16 THE HISTORY OF THE BYZANTINE EMPIRE Leo III issued the first edict against images, which in the Greek Church was directed specifically against the icons. Under Constantine V the struggle became more embittered, and in 765 a fierce persecution was set on foot. In 787 the Empress Irene, an Athenian by birth, succeeded in re- establishing the cult of images, but an Iconoclast reaction under three Emperors of Asiatic origin (8 I3~42^renewed, though with more limited scope, the measures of Leo and Constantine. In 843 the images were finally restored. The Iconoclast movement can be treated neither in isolation from the secular reforms, nor as subordinate to them. In its later stages the attack was directed primarily against the power and influence of the monasteries, as being the strongholds of the cult of images; and the monks reta- liated by boldly challenging the Emperor's constitutional supremacy in Church affairs. But the Isaurians were neither rationalist anti-clericals nor dogmatic innovators. The use of images had not been favoured by the Early Church, and puritan tendencies had appeared sporadically in the fourth and sixth centuries. Asia Minor was their particular centre at this time, and Jewish or Muslim hostility in these parts to a religious use of an art of representation may not have been without effect, as the abusive epithet * Saracen-minded', hurled at Leo III by his opponents, possibly indicates. Christological issues were deeply involved on either side, and it must always be emphasized that for the Byzantines the question was primarily a theological one. Popular feeling and the immense power of tradition were ultimately the deciding factors. The triumph of the icon-defenders was a victory for popular religion and popular ways of thought. The defeat, on the other hand, of the movement towards a separation between the spheres of State and Church reflected no less accurately the Byzantine conviction of the indissolubility of civil and religious government. The reign of Irene, first as regent, and later as Empress after the deposition and blinding of her son, appears at first sight to be merely an interlude between two periods of Iconoclasm. Actually, however, the Second Council of Nicaea (787), which temporarily restored the images, formulated the theory of icon-worship with such success