CHAPTER XI AS REGARDS HEREDITY Problems of Heredity—Physiological Units—A Digression — The Germ-Cells — Transmission of Acquired Characters—Inconceivability—A Priori Argument— Practical Conclusion HEREDITY is the relation of genetic continuity which links generation to generation. An inheritance is all that the organism is or has to start with on its life- journey in virtue of the hereditary relation to parents and ancestors. In all ordinary cases, the inheritance has its initial material basis in the egg-cell and the sperm-cell which unite in fertilisation at the beginning of a new life, and these two kinds of germ-cells, which bear the maternal and the paternal contributions, have their peculiar virtue of reproducing like from like, just because they are the unchanged or very slightly changed cell-descendants of the fertilised ova from which the parents arose. A bud or a cutting separated off from a living creature—tiger-lily or potato, polyp or worm—reproduces an entire organism like the parent, if the appropriate nurture-conditions are available 5 and it can do so because it is a fair sample of the parental organisation. Similarly a germ- cell or two germ-cells in conjunction can develop into a creature like the parent or parents, in virtue of being the condensed essence of the parental organisation. And the germ-cell is this because of its direct continuity 154 habiting another medium, are at the same