Memoirs of the Queensland Museum. VOL. X. PART V. Issued 30th OCTOBER, 1935. EDITED BY THE DIRECTOR, HEBER A. LONGMAN, F.L.S., C.M.Z.S. ISSUED BY THE AUTHORITY OF THE CHIEF SECRETARY FOR QUEENSLAND, THE HON. W. FORGAN SMITH, LLD. CONTENTS, New Australian Sawflies (Hymenoptera, Symphyta) - An Abnormal Lorikeet — Plate XXXIII* — — — Notes on Rearing Young Ceratodus — — — - A New Astacopsis from North Queensland — Plate XXXIV* Palaeontological Notes — - Robert Benson, M.A. - H. Greensill Barnard - A. Rudel - — - Kathleen Watson, B.A. - H. A. Longman ~ PAGE. - 211-229 230 - 231-232 - - 233-235 - 236-239 MEMOIRS OF THE QUEENSLAND MUSEUM, Vol. X, Past V. NEW AUSTRALIAN SAWFLIES ( Hymenoptera , Symphyta). By Robert B. Benson, M.A., Department of Entomology, British Museum, London.. I have lately had the opportunity of studying a most interesting collection of sawflies from the Queensland Museum. The collection, of approximately 150 specimens, contained representatives of at least 15 new species ; 13 of these are described in the present paper, while the descriptions of two others will appear later elsewhere. One new genus is describsd Styracotechys, and for this Styracotechyince a new subfamily of the Pergidce, had to be erected. Keys to a new classification of the Pergidce are included so as to show the position of this new subfamily in relation to the other subfamilies, and in the keys a new subfamily Paralypiince is also instituted for the South American Paralypia and allied genera. Mention should also be made here to one further addition to the Australian fauna in Senoclidea ? furvus Konow from Port Darwin. This species was previously known only from New Guinea. The richness of this material, which was collected in the field mostly by Mr. H. Hacker, Entomologist of the Queensland Museum, suggests that there are still many more new species of sawflies yet to be found in Australia. AKGID,E. ANTARGIDIUM Morice. I lately redefined this genus, Benson 1934 (1), and described two new species ; in the present collection there are two more new r ones, both having in the hind wing the recurrent vein nearer to the base of the wing than the cubital, so that the discoidal cell is smaller and shorter than the cubital as it is in Antargidium allucente Benson, fig. 1 of that paper, though the exact position of the recurrent vein is different in three specimens of one of the species. It now appears that the exact position of these veins differs in individuals of the same species, and probably, if long series could be examined, would be found to be of no significance in separating the species. A comparison of the saws of the new species with the three already described shows that the saw figured for A. allucente Benson, fig. 7 in my previous paper, is in some respects not typical for the genus ; in all four other species the hair bands are eplaced by a row of short broad-based spine (see figs. 1 and 2), and the denticulations are of the pattern illustrated in fig. 9 of that paper, although differing in detail. Furthermore the saw of A. allucente Benson was not figured B 212 MEMOIRS OF THE QUEENSLAND MUSEUM. quite accurately, as further examination has revealed that it also has got a few broad-based spines obscured by the hairs in the hair-bands ; these spines are not easily visible and were overlooked before. J rn . m. . Fig. 1. — Antargidium atriceps sp. now, tenth tooth of saw. Fig. 2. — A. rufum sp. now, tenth tooth of saw. ANTARGIDIUM ATRICEPS sp. now $ Colour. — Yellow ; black are the head (except palps and the tips of the mandibles), the meso- and meta-notum, a large spot on the mesopleura, the mesosternum, and the 6 apical segments of the abdomen except the base of the sawsheath in the $ ; apex of hind tibia and all tarsal segments slightly fuscous. Wings infuscate throughout ; stigma and venation black. Length. — § 5*5 to 6-5 mm. ; forewing 5*5 to 6-5 mm. : antenna 1.7 to 2 mm. ; o 6 mm. ; forewing 6 mm. ; antenna 2*5 mm. Puncturation. — Rugulose on face ; rest of insect shining and unpunctured. Head : malar space about two-thirds length of pedicel of antenna ; pedicel about two-thirds as long as broad ; supra-clypeal area with a well-developed carina reaching from clypeus to frontal area where it divides into 2 branches w'hich surround the pear-shaped median fovea ; ocellar basin deep, including front ocellus and surrounded by a pentagonal wall. Legs : with hind basitarsus about as long as three following tarsal segments together. Wings : recurrent vein in hind wings nearer to the base of the wing than the transverse-cubital vein, so much so that the recurrent vein is twice as far from the apex of the cubital vein as the transverse-cubital vein is. Abdomen : hind margin of hypopygium entire; sawsheath as in A. allucente Benson; saw fig. 1. New South Wales, Tamborine, 1 , 1 (holotype and allotype), 21-ii-1927, H. Hacker (Queensland Museum); Queensland. Conondale 1$, 7-M930, H. Hacker (British Museum) ; and a variety in which the transverse cubital vein is only about two-thirds as far from the apex of the cubital vein as the NEW AUSTRALIAN SA W FLIES. 213 recurrent vein (indicating that the positions of these veins are more liable to variation than was at first thought) and in which the black pigment is less widespread, so that the front and sides of mesonotum together with the scutellum and the metanotum are yellow, while the dark mark on the mesopleura is much smaller. Queensland, National Park, 1 §, xi-1920, H. Hacker (Queensland Museum). This species is separated from all others in the genus by the black head in both sexes, the very pale legs, and the carina on the face reaching right to the front of the supraclypeal area ; the sawsheath is most like that of A . apicale W. F. Kirby, but the dorsal tooth is not so sharp when viewed from the side. ANTARGIDIUM RUFUM sp. nov. $ Colour. — Reddish yellow all over, except the tibiae and tarsal segments which are infuscated. Wings, infuscated throughout ; stigma and venation black. Length 6-3 mm. ; forewing 6-3 mm. ; antenna 25 mm. Puncturation obsolete. Head : malar space about half as long as pedicel of antenna ; pedicel about two-thirds as long as broad ; supraclypeal area with the .medial carina present on the hind two-thirds of the area, but not sharp, the front one-third is simply rounded; frons as in A. atriceps sp. nov. Legs with the hind basitarsus distinctly longer than the three following tarsal segments together. Wings : recurrent vein in the hindwing nearer to the base of the wing than the transverse-cubital vein, as in A. allucente Benson (Benson 1934 (1), fig. 1). Abdomen: hind margin of hypopygium entire; sawsheath as in A. dentivalvis Benson, but set up almost erect ; saw fig. 2. New South Wales, Tooloom, 2$$, i-1920, H. Hacker (holotype in Queensland Museum ; paratype in the British Museum). This species is distinguished from all others by the head and body being entirely reddish -yellow, and by the very short almost linear malar space. The sawsheath comes nearest to that of A. dentivalvis Benson, but it is set up almost erect, whereas in that species the sawsheath is not so set. ANTARGIDIUM ? DENTIVALVIS Benson. One female, Queensland, Townsville, Dr. H. Priestly, in bad condition, probably belongs to this species. It differs from the type in being larger (4-5 mm. instead of 4 mm.), in the sawsheath being set up erect at the apex and in the transverse-cubital vein in the hind-wing not being interstitial with the recurrent vein, but received on the cubital nearer the apex of the wing as in A. allucente Benson and typical Arge. 214 MEMOIRS OF THE QUEENSLAND MUSEUM. TENTHREDINIDiE. SENOCLIDEA ? FURVUS Konow. Monophadnus furvus Konow, Wein ent. zt. xvii, 1898 : 232. North Australia, Port Darwin,. 1$, F. P. Dodd. This species, previously known only in New Guinea, is a very interesting addition to the Australian sawfly fauna as it represents the first true Australian Tenthredinid (not counting, of course, Caliroa limazina Retz., which is obviously an alien that has been introduced with fruit trees into a great many parts of the world). PTER YGOPHORI D.E. CLARISSA FLAMMEA, sp. nov. $ Colour reddish -yellow ; head black except for the pale mouth -parts, clypeus, supraclypeal area, and antennae ; a large cream-coloured spot covering the hind half of the outer quarter of abdominal tergum 2. Wings hyaline, slightly infuscate at the apex ; veins at extreme base of wing, including basal half of costa yellow ; rest of veins infuscate ; stigma infuscate, with a pale transparent basal patch. Length 6 mm. ; fore wing 5*5 mm. ; antenna 2 mm. Puncturation. — Head [and thorax dull with fine surface puncturation becoming rugulose on head ; abdomen rugulosely sculptured. Head : mouthparts normal ; malar space about as long as pedicel ; antenna 1 1 -segmented ; subclavate pedicel a little longer than broad ; segment 3 = 4 -f- 5 in length ; only segments 9 and 10 broader than long ; median fovea as a slight rounded depression. Legs : hind-basitarsus about equal to three following tarsal segments. Abdomen : with sawsheath normal ; saw fig. 3. Queensland National Park, 1 $, 25-X-1923, H. Hacker (Queensland Museum). This species is superficially very similar to C. divergens W. F. Kirby, and would run to that species in my key — Benson 1934 (2) p. 469. Externally NEW AUSTRALIAN SAW FLIES. 215 it is only distinguishable on colour characters, the thorax and abdomen being all red, while in C. divergens W. F. Kirby the mesepimeron, mesosternum, and metapleurae together with the last two or three abdominal segments are black, and the costa being yellow at the base instead of all black. The saw is however quite different ; it is more like the saw of C. flavicornis Benson (Benson 1934 (2) fig. 6a). The waved ridges on the side of the saw, however, suggest a link between the saw of C. flavicornis Benson and the spiny-ridged saw of C. ruficollis Benson. Clarissa males. In the collection were males belonging to four different species of this genus, but none of them seemed to belong to any of the described females. I have with some diffidence then described four new species based on the male sex only, which are separated in the key that follows from the already known males of C. atrata G. Turner and C. divergens W. F. Kirby. The male genitalia in this group seem to exhibit but slight and insufficient differences to be relied on yet for separating the species. 1 . Pedicel clearly broader than long ^ 6 Pedicel longer than broad 2 2. Head and thorax metallic greenish -black ; abdomen, except tergum 1, entirely reddish yellow ; hind legs all black. Eyes strongly converge in front so that the malar space is linear diana sp. nov. Head and thorax shining black without metallic tinge ; abdomen, only yellow banded, being black at base and apex ; at least hind tibia mostly yellow. Eyes converge in front, but malar space is still at least half as long as the diameter of an ocellus 3 3. Flagellum of antenna yellow ; the segments each being strongly swollen apically. Puncturation on temples and thorax very fine, almost obsolete. Wings slightly infuscate lucida sp. nov. Antenna all black ; the segments not noticeably swollen out apically. Puncturation on temples very fine and close. Wings hyaline 4 4. Abdomen with whole of segments 2, 3 and often part of 4 yellow divergens W. F. Kirby. Abdomen with only apices of segments 2, 3 and 4 yellow atrata G. Turner. .5. Flagellum of antenna together with a band on abdomen and legs (except coxae, trochanters and tibial spurs) yellow. Inner apical spur to hind tibia, about half as long as hind basitarsus antennata sp. nov. 216 MEMOIRS OF THE QUEENSLAND MUSEUM. Antenna, abdomen and for the most part the hind legs black. Inner apical spur to hind tibia almost as long as hind basitarsus obscura sp. nov. CLARISSA DIANA sp. nov. Colour. — Metallic greenish -black ; yellow are on the fore legs, the front of the femur, the tibia and tarsi, on the middle legs a spot on the apex of the femur, and the whole of the abdomen except tergum 1. Wings slightly infuscate throughout ; veins and stigma black. Length 4*7 mm. ; fore wing 4-5 mm. ; antenna 2-2 mm. Puncturation : head with very fine regular surface puncturation, but shining between the punctures ; the punctures are visible also on the prothorax, but on the mesonotum and mesopleura3 the punctures are apparent only at the sides of the lobes, the middles of the lobes being unpunctured smooth and shining ; abdomen with normal rugulose transverse sculpturing. Head with eyes strongly converging in front so that they almost touch the side of the clypeus and the base of the mandible, leaving but a linear malar space ; length of clypeus and labrum together equals the breadth of the clypeus ; antenna 9-segmented ; distinctly clubbed from the apex of segment 4 ; segment 2 (pedicel) longer than broad ; median fovea as a long groove, joining the frontal furrow and reaching back to the front ocellus behind. Wings as in Clarissa divergens W. F. Kirby, but in the unique type the first transverse cubital nerve is absent in the fore wing. Legs with inner spur on apex of hind tibia about half as long as hind- basitarsus ; hind-basitarsus about equal to 2 following segments in length. Abdomen with apex upturned ; hypopygium entire behind. Queensland, Nanango District, 1 xi-1927, H. Hacker (Queensland Museum). CLARISSA LUCID A sp. nov. <$ Colour black; yellow are the labrum, flagellum of antenna, angles of pronotum behind, legs (except the infuscate coxae, trochanters and tibial spurs) and a band on the abdomen covering terga 2-4 and sterna 2-5. Wings infuscate ; stigma black with a paler base ; costa and venation black basally but brownish apically. Length 6-5 mm. ; forewing 5-5 mm. ; antenna 3*5 mm. Puncturation fine dense and regular on face and frontal region, but obsolete on the temples whose surface is smooth and shining ; obsolete also on NEW AUSTRALIAN SAW FLIES. 217 most of the thorax especially the lobes of the mesonotum where sparse vague punctures are only apparent in the hollows and at the sides ; abdomen transversely rugulose. Head : length of clypeus and labrum together about equals breadth of clypeus ; malar space about as long as two-thirds the diameter of an ocellus ; antenna 12-segmented, not club-shaped, but subserrate, the segments from 4 onwards being of equal breadth apically, but each one being very broad apically and narrow basally, so that the breadth basally is about two -thirds the breadth apically ; median fovea as a deep rounded depression ; vertical furrow clearly marked. Legs with hind basitarsus equal to three following tarsal segments together ; inner hind tibial spur about five-sevenths length of basitarsus ; Abdomen with hypopygium set ujj erect, with the hind margin slightly two-lobed,. being emarginate in the middle. New South Wales, Tooloom, 2^^, i-1926 H. Hacker (holotype in Queensland Museum, paratype in British Museum). CLARISSA ANTENNATA sp. nov. Colour black : yellow are labrum flagellum of antenna hind angles of pronotum, legs (except coxse trochanters and tibial spurs), a band on abdomen covering terga 2-4 and sterna 2-5 and the apex of the hypopygium. Wings almost hyaline, slightly infuscate stigma (except for a pale spot at base), costa and venation dark brown. Length 5-5 mm. ; fore wing 5 mm. ; antenna 4 mm. Puncturation on head fine dense and irregular, though on the clypeus and temples small unpunctured spaces occur between the punctures ; pro- and mesonotum shining betw 3en scattered regular punctures ; meso-pleurse with regular very fine surface punctures ; abdomen transversely rugulose dorsally. Head : length of clypeus and labrum together about equal to breadth of clypeus ; clypeus entire on front margin ; malar space as long as the diameter of an ocellus ; antenna 12-segmented . not club-shaped ; the breadth of segments 4 onwards being the same apically ; pedicel clearly broader than long ; segments from 4 onwards slightly broadened apically ; median fovea as a deep rounded depression ; vertical furrows very distinct. Legs with hind basitarsus equal to 2 following tarsal segments together in length; inner hind-tibial spur about five-sevenths as long as hind basitarsus. Abdomen with hypopygium normal in position, excised in middle on hind margin, and slightly emarginate each side. Queensland, Tamborine Mountain, 1 < J, W. H. Davidson (Queensland Museum). 218 MEMOIRS OF THE QUEENSLAND MUSEUM . CLARISSA OBSCURA sp. nov. Colour black ; pale brown are the labrum, front margin of clypeus, the front legs (except base of femur, trochanters and coxa) the middle and hind tibiae apically as well as the tarsal segments at their joints. Wings slightly infuscate ; stigma and venation black, except basal half of costa which is brownish. Length 6 mm. ; fore wing 5-5 mm. ; antenna 4 mm. though the Brisbane specimen is 8 mm. ; forewing 6-2 mm. ; antenna 4-5 mm. Puncturation on head dense, shallow and irregular, on nota of thorax in the form of regular round scattered punctures with shining smooth spaces between ; the punctures are thicker at the sides of the lobes of the mesonotum and the scutellum, but sparser in the middle of the mesonotal lobes ; on pleurae and sterna the puncturation is irregular and shallow ; on the abdomen the dorsum is transversely rugulose. Head : length of clypeus and labrum together equal to about the breadth of the clypeus ; clypeus emarginate in the middle of the front margin ; malar space in length about two-thirds the diameter of an ocellus ; antennae 14-15 segmented, not club-shaped, increasing in breadth slightly until about segment 6 or 7, and then tapering slightly ; funicle and pedicel very short ; pedicel about twice as broad as long ; segments 4 and onwards being each much broader apically than basally, the basal breadth being about two-thirds of the apical breadth ; median fovea as a deep rounded depression ; vertical furrows clearly defined. Legs with hind basitarsus equal to two following tarsal segments ; inner hind tibial spur in Brisbane specimen equals about five-sixths length of basitarsus, in other specimens almost equals basitarsus in length. Abdomen with hypopygium set up erect, and entire on hind margin. Queensland, 3, Queensland, Brisbane, Mount Cootha, 20-M925, H. Hacker, and 2$$, North Queensland, Ayr, 30-X-1925, S. Bates. Either here we have an extraordinary case of distribution or else, what seems more probable, there was some error in the localities recorded by Westwood. XYLOPERGA PERKINSI sp. nov. ? Colour yellow ; black are the tips of the antennae, an ocellar patch, the shortest middle part of the pronotum, the meso- and meta-notum (except the scutellum and the sides of the mesonotal lobes reaching from front angles of the c 228 Memoirs of the Queensland museum. scutellum to the tegulae), propodeum and front half of abdominal tergum 2, the lower portion of the mesepisternum together with the mesosternum and mesepimeron and the meta-sternum and metepimeron, the middle of hind coxae, and the abdominal sterna. Wings tinged with yellow, especially fore- wings ; hind wings nearly hyaline ; stigma dark brown ; costa and venation brown. Length 17 to 18 mm. ; forewing 14 to 15 mm. ; antenna 3 mm. Pubescence very sparse, almost entirely absent from head (except mouthparts) and thorax, very short and scattered on ventral surface of abdomen. Head. — Face with sparse coarse punctures ; head behind antennae more strongly punctured especially on depressed borders to postocellar area and temples ; malar space about half-length of pedicellum ; antenna 7-segmented ; about as long as shortest distance between eyes ; clavate from apex of 5th segment ; distance between the hind ocelli greater than distance from ocellus to eye ; median fovea and frontal furrow fused to form a pear shape depressed area including the front ocellus ; lateral furrow starts on about a level with the front ocellus and continues back as a slight dejDression to the vertex ; post-ocellar area slightly convex and rounded. Thorax. — Shining all over with coarse scattered rounded punctures becoming dense on the front and side lobes of the mesonotum, especially on the middle of the side lobes, and very sparse on scutellum which is strongly contracted behind, slightly depressed longitudinally in the middle and with very small appendages behind ; mesosternum and metapleurae unpunctured. Abdomen with transverse rugulose sculpture. $ Colour. — Yellow except for the following which are black : tips of mandibles ; the ocellar region and a band stretching backwards to the vertex each side of the post-ocellar area ; pronotum in front and side lobes of mesonotum (except the raised margins of the side lobes stretching from teguke to scutellum) ; depressed parts of metanotum ; dorsal parts of abdominal terga. Wings yellowish hyaline with patch of scales (? androconia) occupying sub-costal, first discoidal and first two cubital cells of forewing ; stigma brown ; tegula and venation yellow in basal half of wings, brown in apical half. Length 13 mm. ; fore wing 11 mm. ; antenna 3 mm. Structure as in female except for wings, genital segments and slightly denser puncturation noticeable on mesopleurse where between punctures the surface is dull and rugulose. West Australia, Cunderdin, 2$$ (including holotype) 3(J