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ADDITIONS AND CORRECTIONS
to
An Annotated List of Generie Names of

the Seale Insects (Homoptera: Coccoidea)

By Louise M. Russeuu

Systematic Entomology Laboratory

SUPPLEMENT

Miscellaneous Publication No. 1015

Agricultural Research Service

UNITED STATES DEPARTMENT OF AGRICULTURE

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PREFACE

This publication contains additions and corrections to “An Anno-
tated List of Generic Names of the Scale Insects (Homoptera: Coc-
coidea” by Harold Morrison and Emily R. Morrison, published in
October 1966. It extends through December 31, 1963, as did the Mor-
rison and Morrison publication, and in addition includes citations to
a 1965 paper by De Lotto, which has a crucial bearing on certain
coccid generic names, and the 1966 diaspidid catalog of Borchsenius.

Morrison and Morrison treated names proposed by Amyot, 1847:
488-503, as uninominals and hence as invalid names without explain-
ing their action. Neave listed the Amyot names without comment.
Although inclusion in Neave’s list does not imply validity of the names,
their presence there has led some workers to question the correctness of
the Morrison and Morrison conclusions.

It therefore seems relevant to note the following statement by
Amyot, 1845: 369: “Ce qui doit le plus frapper au premier abord, dans
cet ouvrage, est le mode de nomenclature que nous y avons adopté,
sous le titre de Méthode mononymique, et qui consiste dans l’applica-
tion d’un nom unique, donné a chaque espéce, au lieu de deux, le nom
générique et le nom spécifique, comme cela se pratique dans la nomen-
clature en usage depuis Linné.”

This system obviously does not meet the requirements of Article
11(c) of the International Code of Zoological Nomenclature. The
matter was settled unequivocally by the International Commission in
Opinion 686, 1963: 423, when Amyot’s Méthode mononymique was
placed on the Official Index of Rejected and Invalid Works in Zoologi-
cal Nomenclature.

Tt is evident from their comments on Amyot names that Morrison
and Morrison were not aware of Opinion 686.

A few Amyot coccid names were omitted by Morrison and Morrison.
They are included in the present list for the sake of completeness, and
notes or corrections are added under a few Amyot names included
by Morrison and Morrison.

Morrison and Morrison usually cited the name of a type-species
in its original combination but sometimes cited a later combination if
one had been used by the describer of the genus. In the latter instances
that I have recognized, I have added or substituted the original name

combination of the type-species.

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~

Entries in the present list are arranged by the page number of the
original publication and are followed by the generic name under which
the addition or correction appears. Generic names omitted from the
original list are here included in alphabetical sequence following the
page number and are marked “(new entry).”

Except for the references given at the end of this list, publications
cited may be found in the coccid bibliographies of Morrison and
Renk, 1957, and Morrison and Morrison, 1965.

I extend my appreciation to all persons who have informed me of
omissions and errors in the 1966 publication and especially to Giovanni
De Lotto, Plant Protection Research Institute, Pretoria, South Africa.
I am also deeply indebted to Curtis W. Sabrosky, Systematic Ento-
mology Laboratory, U.S. Department of Agriculture, for advice on
nomenclatorial problems.

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ADDITIONS AND CORRECTIONS
Acontonidia: Change “triangularia” to “triangularis”
Acreagris: Insert “123,” before “pl.”

Albastaspis: Change “295” to “275”

Anamefiorinia: Change “caswariniae” to “casuarinae”

Anaparaputo: Change “1961” to “1962”

Anisococcus: Insert “21,” before “26.”

Anophococcus: Add after discussion “This synonymy was
indicated by Hoy, 1962: 23, and 1963: 132-133. See discussion
under Anothococcus Ferris.”

Anoplaspis Leonardi, 1898: After “(207)” add “nomen nudum;
Leonardi, 1898, in Berlese and Leonardi, Riv. di Patol. Veg.
(1897) 6: 131 [289].”

Antakaspis: Change “terminalis” to “terminaliae”

Antecerococcus: Change “punctiferus Green, 1900,” to “puncti-
ferus Green, 1901,”

Aonidia: Change “43” to “42”
Aonidomytilus: Change “102.” to “102-103.”
Aspidaspis: Change “45 ;” to “45, nomen nudum ;”

Aulacaspis: Change “1834” to “1833” and insert “C. L.” before
“Scott”

Austrolichtensia: Change“ Lecaniodiaspis” to “Lecanodiaspis”
and add after discussion of type-species “Fuller, 1897, first
used the name ‘Lecanodiaspis (4?) hakearum’ for the species

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that he later, 1899: 457, called ‘Zichstensia hakearwm sp. n.’ In
1899 : 458, he stated, ‘In my preliminary list this species is men-
tioned asa Lecaniodiaspis (?);....”

Balachowskiella: Change “1956” to “1955”
Boisduvalia: Change “1822” to “1882”

Brachyscelis: Insert “Type-species: Brachyscelis pileata
Schrader, 1863, by subsequent designation by Lindinger, 1937:
179, through substitution of Apiomorpha Ribsaamen for pre-
occupied Brachyscelis.”

Caelostoma: Change “Cockrell” to “Cockerell”

Cephalaspis: Add after discussion “Borchsenius, 1963: 1162,
accepted the concept of Cephalaspis MacGillivray and pro-
posed Parainsulaspis to replace the preoccupied name. See dis-
cussion under Parainsulaspis.”

Cerecococcus: Insert “H.” before “Scott”

Ceroplastes: Change “chilensis Gray, 1827,” to “janetrensis
Gray, 1828,”

Chentraspis: Change “284, 286;” to “284-286;” and “6:111”
to “6: 110-111”

Chionaspis: Insert as first two references “Targioni-Tozzetti,
1868, (separate) Soc. Ital. di Sci. Nat. Atti 11:5; 1869, 11: 698,
nomen nudum ;”

Chnaurococcus: Insert “22,” before “40”

Chortinaspis: Change “46,” to “45-46,”

Coccinella: Insert after second sentence of discussion “Gmelin
did not include a species name under Coccinella.”

Coccinilla (new entry) Amyot, 1847, Soc. Ent. de France Ann.
(ser. 2) 5: 493.

Amyot wrote “Coccus ou Coccinilla’ and included the species name

“cacti Linn.”

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LIST OF GENERIC NAMES OF THE SCALE INSECTS 3 :

P

4. Cocconidia: Change “1758” to “1767” and add after discussion
“Under Cocconidia Amyot included the species name ‘adoni-
dum Linn.’ The latter name stood for the mealybug that has
been known erroneously as Pseudococcus adonidum (Linnaeus,
1758 or 1767). De Lotto, 1965: 226-228, has shown that there
is no Coceus adonidum Linnaeus, 1758, that Coccus adonidum
Linnaeus, 1767, is not a scale insect, and that Pseudococcus
adonidum auct. is Pseudococcus longispinus (Targioni-
Tozzetti, 1867).”

Coccopoa: Change “uninomial” to “uninominal” and “1761” to
“1758” Insert after first sentence of discussion “In 1758
phalaridis was validated by bibliographic reference to the 1746
Fauna Suecica where the species was described as completely
as in 1761.”

Coccopsidia (new entry) Amyot, 1847, Soc. Ent. de France Ann.
(ser. 2) 5: 496.
Amyot proposed this name for Coccus psidii Chavannes, thus appar-

ently publishing the name before the description of psidii appeared in
the 1848 Chavannes publication.

43 Cockerellella: Change “p. 71.” to “pp. 71, 75.”

45 Constockiella (new entry) MacGillivray, 1921, The Coccidae,
p. 423.

A lapsus for Comstockiella Cockerell.

46 Cornimytilus (new entry) Borchsenius, 1963, Zool. Zhur. 42:
1165.

TYPE-SPECIES : Lepidosaphes afganensis Borchsenius, 1962, by original
designation and monotypy.

The author indicated a close relationship between this genus and
Lepidosaphes Shimer and Eucornuaspis Borchsenius.

Cornoculus: Insert “78,” before “81” and change type-species
to read “Cornoculus cornutus Ferris, 1955, by monotypy;
Cornoculus oculatus Ferris, 1955, by original designation,
nomen nudum.” Insert before discussion “Ferris designated
the type-species of this genus as ‘Cornoculus oculatus, new
species; the only included species.’ However, he described the
only included species under the name C’. cornutus, and did not
mention oculatus again.”

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Crisicoccus: Insert “22,” before “45”

Cryptophyllaspis: Insert as first reference “1897, in Leonardi,
Riv. di Patol. Veg. 5: 375 ;”

Cyphoma: Change “Cyphoma characias” to “Orthezia chara-
cias” and delete brackets.

Cystococcus: Insert as second reference “ (1897, Separate, Notes
on Coccidae, p. 11).” and change “19022” to “1902h”

Cystococcus (new entry) Lindinger, 1937, Ent. Jahrb. 46: 182.

An alternate spelling for Cisticoccus A. Costa.

Dactylopius: Change “1758” to “1767” and add after discussion
“De Lotto, 1965 : 226-228, has shown that there is no Coccus
adonidum Linnaeus, 1758, that Coccus adonidum Linnaeus,
1767, is not a scale insect, and that Pseudococcus adonidum
auct. is Pseudococcus longispinus (Targioni-Tozzetti, 1867).
Since it is recognized that Coccus adonidum Linnaeus, 1767,
the possible type-species of Dactylopius, was misidentified,
it is most urgent that this case be submitted to the International
Commission.”

Delococeus: Change “Formicoccus” to “Formicococcus”

Diaprosteci: Change “1758” to “1767” and add after discussion
“De Lotto, 1965 : 226-228, has shown that there is no Coccus
adonidum Linnaeus, 1758, that Coccus adonidum Linnaeus,
1767, is not a scale insect, and that Pseudococcus adonidum
auct. is Pseudococcus longispinus (Targioni-Tozzetti, 1867).
Since the type-species was misidentified, the case should be
referred to the International Commission.”

Diaspidistus: Change to “Diaspidistis”

Diaspis: Change “echinocactus” to “echimocacti”

Dorthesia: Change type-species to read “Orthezia characias
Bose d’Antic, 1784, by inonotypy, = Aphis urticae Linnaeus,

1767.” and change “Coccus urticae Linnaeus.” to “Orthezia
urticae (Linnaeus).”

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LIST OF GENERIC NAMES OF THE SCALE INSECTS 9)

Dorthezia (new entry) Signoret, 1869, Soc. Ent. de France Ann.
(ser.4) 8: 833.

Apparently a lapsus for Dorthesia since it appeared twice against seven
spellings of Dorthesia in Signoret, 1868-77. See discussion under Dorthesia.

Dynaspidiotus: Change “131,” to “131-152,”

Emmereziaspis: Change “a//audi de Charmoy” to “allwaudi
Grandpré and Charmoy”

Eucornuaspis (new entry) Borchsenius, 1963, Zool. Zhur. 42:
1168.

TYPE-SPECIES: ( Mytilaspis machili Maskell, 1898) =Aspidiotus pinnae-
formis Bouché, 1851, currently Hucornuaspis pinnaefornis
(Bouché), by original designation.

Borchseninus, 1966: 58, synonymized machili with pinnaecformis. He
indicated a close relationship between this genus and Cornuaspis
Borchseninus.

Eulepidosaphes: Change “384” to “364”

Euparlatoria: Change “Parlatoria oleae” to “Diaspis oleae”

Fernaldiella: Change first sentence of discussion to read
“Leonardi presented this spelling in his generic key (p. 4)

and in a list of species (p. 112); other (3) references in lis
paper were to Ferna/della.”

Ferrisiana: Insert “Type-species: Dactylopius virgatus Cock-
erell, 1893, by substitution of Ferrisiana for Ferrisia Full-

away.”

Froggatiella (new entry) Morrison and Renk, 1957, U.S. Dept.
Agr. Mise. Pub. 734: 129.

A lapsus for Froggattiella Leonardi.
Fulbrightia: Insert “1950,” after “gallicola Ferris,”
Genistaspis: Change “zelihiae” to “zelihae”

Globulicoccus: Change type-species to read “Coccus fuscus
Gmelin, 1789, ‘Z.[ecanium] . . . Dougl.’ of Lindinger.”

Gossyparia: Change “Coccus spurius Modeer, 1778,” to “Gos-
syparia spuria (Modeer, 1778),”

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Greenacoccus: Insert “Type-species: Afonophlebus stebbingii
Stebbing, 1902, by substitution of Greenacoccus tor Greeniclla
MacGillivray.”

Greeniella (non Cockerell, 1897) : Change ‘75, 474.” to “459.”
Insert after first sentence of discussion “Greeniella appeared
on pp. 70 and 459. The spelling Grcenella appeared on p. 75,
where the type was designated, and on p. 474, where the name
was changed to Greenacoccus.”

Greenoripersia: Change “1921” to “1929”
Guerinia: Change “39” to “31”

Guerinococcus: Insert “Type-species: Guerinia tinctoria Tar-
gioni-Tozzetti, 1868, believed to be the same as Coccus ser-
ratulae Fabricius, 1775, by substitution of Guerinococcus for
Gucerinia Vargion-Tozzett).”

Haliaspis (new entry) Takagi, 1963, Insecta Matsumurana 26:
118.

TYPE-SPECIES: Chionaspis spartinae Comstock, 1883, by original
designation.

This genus was established to receive three North American species
formerly placed in Duplachionaspis MacGillivray.

Hemiaspidis: Change “/7emaspidus” to “Hemaspidis”
Hemiberlesiella: Delete brackets.

Hemilecanium: Insert after first sentence of discussion “Green,
1916: 6-7, indicated that theobromae was a synonym of
7mOIMCANS.”

Incisaspis: Delete “311,” and add after discussion “The spelling
[ne‘aspis appeared on p. 3811 where the type-species was
designated.”

Kermesoides: Insert “1869,” after “7argionia nigra Signoret,”

Lakshadia: Change “7'achardia lacca (Kerr), by substitution.”
to “Coccus lacca Kerr, currently Laccifer lacca (Kerr) .”

Lattaspidiotus: Insert “394,” before “457-458” and change ‘“‘de
Charmoy,” to “Grandpré and Charmoy,”

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LIST OF GENERIC NAMES OF THE SCALE INSECTS 7

Lecanodiopsis (new entry) Lichtenstein and Signoret, 1886,
Soc. Ent. de France Ann. (Bul. Ent.) (ser. 6) 6: xxxix,

A lapsus for Lecanodiaspis Targioni-Tozzetti. The species involved, L.
dugesii, is currently placed in Ceroplastodes Cockerell.

Liucoccus: Change “Liuococcus ehrhorniodes” to “Liucoccus

ehrhornioides”
Longisomus: Insert “nomen nudum,” before “by monotypy.”
Lophococcus: Change “1902” to “1901”
Luzulaspis: Insert *?” after “Aspidiotus”

Lyraspis: Change “46; to “46, nomen nudum ;” and add after
discussion “Lepage. 1939: 313, stated that in 1939 Ferris (per-
sonal communication) informed him that Lyrasp/s was a syn-
onym of Viveaspis MacGillivray.”

Macracanthopyga: Change “1957, by 87,” to “1957b: 66, 1957c:

ol,
Mallophora: Change “1896,” to “1897,”

Marchaliella: Change “J/onophlebus hellenicus Gennadius,
1883,” to “Chionuspis lepineyi Balachowsky, 1928,”

Marlattaspis: Change “monotypy.” to “monotypy,=Asp/diotus
lataniae Signoret, 1869.” Change “Signoret.” to “(Signoret ).”

Maskellia: Insert as second and fourth references * (1897, Misc.
Pub. 177: 1-2); (1897, Separate, Notes on Coccidae, p. 5).”

bo

Maskelliella (new entry) Ferris, 1937, Microentomology 2: 4.

A lapsus or emendation for Maskeliella Leonardi. See discussion under
Maskeiiella.
Mixorthezia: Change “1926” to “1925”

Moluscococecus: Change to “J/o/luscococcus”

Murataspis: Change “megaporus Balachowsky” to “megapora
Balachowsky, 1928,”

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Mycococcus: Change “copernicae” to “coperniciae”
Mytilaspis: Change “1798” to “1789”

Mytilella: Insert “4,” before “20”

Najococcus: Change “Lindinger” to “Green” -
Natalaspis: Insert “unnamed” before “var.”

Neogreenia: Insert before discussion “The name Paragreenia
appeared on p. 78 where the type-species was designated.”

Neopulvinaria: Change “¢mertina” to “imeretina”
Nuculaspis: Insert “75,” after “45,”

Odonaspis: Change “284” to “284-286”
Onceropyga: Insert “78,” before “208”
Ourococeus: Insert “Separate,” before “Notes”
Parafairmairia: Change “1876” to “1872”

Parainsulaspis (new entry) Borchsenius, 1963, Zool. Zhur. 42:
1162.

TYPE-SPECIES: Mytilaspis cocculi Green, 1896, through substitution of
Parainsulaspis for Cephalaspis MacGillivray.

Borchsenius erroneously designated Lepidosaphes laterochitinosa Green,
1925, as the type-species. The describer proposed this name as a replace-
ment for Cephalaspis MacGillivray, 1921, a name preoccupied in Pisces.
He included Mytilaspis cocculi Green, 1896, the type-species of Cephalaspis
MacGillivray, in Parainsulaspis but designated Lepidosaphes laterochiti-
nosa as the type of his genus, a species not included in Cephalaspis. Under
the provisions of Article 67(i) of the International Code of Zoological
Nomenclature, cocculi Green becomes the type-species of Parainsulaspis.
Borchsenius indicated a close relationship between this genus and Para-
lepidosaphes Borchsenius and Jnsulaspis Mamet.

Paraonidea: Change “394” to “454”

Parasaissetia: Change type-species to read “Lecantum nigrum
Nietner, 1861, currently Saissetia nigra (Nietner).”

Paraselenaspidus: Change “1953” to “1954”

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LIST OF GENERIC NAMES OF THE SCALE INSECTS 9

Pelomphala: Change “292” to “392”

Phaulaspis: Change “284” to “284-286”

Phenacaspis: Change “38” to “48”

Pinaspis (new entry) Lindinger, 1942, in Titschack Beit. z.
Fauna Perus 2: 126.
A lapsus or emendation for Pinnaspis Cockerell.

Pinomytilus (new entry) Borchsenius, 1963, Zool. Zhur. 42:
1173.

TYPE-SPECIES : Lepidosaphes pseudotsugae Takahashi, 1957, by original

designation and monotypy.
The describer indicated a close relationship between this genus and
Eucornuaspis Borchsenius and Cornimytilus Borchsenius.
Pistaciaspis (new entry) Borchsenius, 1963, Zool. Zhur. 42:
1165.

TYPE-SPECIES: Lepidosaphes pistaceae Archangeiskaya, 1930, by orig-
inal designation.

The deseriber indicated a close relationship between this genus and
Lepidosaphes Shimer.

Planococcoides: Change “Laing, 1956” to “Laing, 1929”

Planococcus: Change type-species to read “Dorthesia citri
Risso, 1813, as interpreted by Ferris, 1950, by original
designation.”

“s

Porphyrophora: Change “1883” to “1833”
Protargionia (new entry) Ferris, 1937, Microentomology 2: 106.

A lapsus for Partargionia MacGillivray.

Pseudaonidea (new entry) MacGillivray, 1921, The Coccidae,
p- 453.

A lapsus for Pseudaonidia Cockerell.

Pseudaulacaspis: Change “Aulacaspis” to “Diaspis”

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Pseudococcus: Change type-species to read “Dactylopius long-
ispinus 'Targioni-Tozzetti, 1867, by subsequent designation of
Fernald, 1903b: 96.” Add after discussion “De Lotto, 1965:
226-228, has shown that Coccus adonidum Linnaeus, 1767, is
not a scale insect, and that Pseudococcus adonidum auct. is
Pseudococcus longispinus (Targioni-Tozzetti, 1867).”

Pterolecanium: Change “48” to “47-48” and insert “(Luleca-
niwm)” before “pulchrum”

Pulvinariella: Change “1830” to “1829”

Pulvinella: Insert “(Pulvinella)” before “pulchella” and “J.
P.” before “da Fonseca”

Radiaspis: Insert as first reference “1938, Microentomology 3:
45,75, nomen nudum;”

Rhodococcus: Change “rosaelutea” to “rosaeluteae”’
Ripersiella: Insert “7m Cockerell,” before “1899”
Saccharicoccus: Change “1893” to “1895”

Sakaramyaspis: Change “Mamet, 1953,” to “Mamet, 1954,”
Sasakiaspis: Change “1885” to “1886”

Schizentaspidus: Change “Schizentapidus” to “Schizenta-
spidus”

Schizochlamys: Change “1900” to “1899”
Selenaspidus: Delete brackets.

Signoretia: Change “Stal, 1869” to “Targioni-Tozzetti, 1868”
and add “by monotypy” after “1864”

Signorettia: Change reference to read “1868, (separate) Soc.
Ital. di Sci. Nat. Atti 11: 25; 1869, 11: 699.”

Solenophora: Change “Maskell, 1899,” to “Maskell, 1890,”

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LIST OF GENERIC NAMES OF THE SCALE INSECTS Let

Spaerolecanium (new entry) Leonardi, 1908, (non Stile, 1908),
Portici R. Scuola Super. di Agr. Lab. Zool. Gen. e Agr. Bol. 3:
180.

Apparently a lapsus for Sphaerolecanium. This spelling appeared on
p. 180. Sphaerolecanium appeared on p. 181, in Leonardi, 1918: 213, and
in Silvestri, 1920 : 312.

Sphaerococcopsis: Change “Sphaerococcopsis inflatipes” to
“Sphaerococcus inflatipes”

Sphaerolecanium: Change “180-181.” to “181.” Add after dis-
cussion “This spelling appeared on p. 181, in Leonardi, 1918:
213, and in Silvestri, 1920: 312. The spelling Spaerolecanium
appeared on p. 180.”

Suturaspis: Change “268” to “262-263”

Symeria: Change “Lepidosaphes epiphytidis (Maskell)” to
“Mytilaspis epiphytidis Maskell”

Tachardia: Insert “R.” before “Blanchard”

Talla: Change type-species to read “Lecaniwm quercus Réaumur
(‘gallinsecte en forme de rein,’ Réaumur, 1738: 79) of von
Heyden, 1860=Coccus quercus Linnaeus, 1758, currently
Kermes quercus (Linnaeus), by monotypy.” Insert before dis-
cussion “This insect was named quercus by Linnaeus in 1758
when he referred to Réaumur’s figures of the ‘gallinsecte’ on
oak.”

Tecaspis: Change “wmtali” to “umtalii”
Tozzetia: Change to “Tozzettia”
Trachycoccus: Insert “78,” before “215”

Trechocorys: Change “monotypy.” to “monotypy,=Dactylo-
pius longispinus Targioni-Tozzetti, 1867, through misiden-
tification of Coccus adonidum Linnaeus, 1767.” Add after
discussion “De Lotto, 1965: 226-228, has shown that Coccus
adonidum Linnaeus, 1767, is not a scale insect, and that Pseudo-
coccus adonidum auct. is Pseudococcus longispinus (Targioni-
Tozzetti, 1867). Since the type-species was misidentified, the-
case should be referred to the International Commission.”

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