eis a ene,
Mi
a Perna, WaRORT OF Phe
Beeman oF AEGIS OF
a
as Bh? FTG NOLAN
i TP Bhd CPO
2 em BAG, BARR AE A
Pricrpe errs tal foie busy Tea
Renee gies Ae ks TTY SOR
i ® Toate a my if ing I P nih
wv a s 7 aah age a, v, : ; » - ; 5 7
ater Dek aro WOULD Haat in
c y 7 : iw. ia at
; Nv 7 LY iw aan : aa) j a tytn i V
, ra i) ; aD, r 7a
| Ne ie Hrs
2
ey ee
fi ch
ANNUAL REPORT OF THE
BOARD OF REGENTS OF
THE SMITHSONIAN
INSTITUTION
SHOWING THE
OPERATIONS, EXPENDITURES, AND
CONDITION OF THE INSTITUTION
FOR THE YEAR ENDED JUNE 30
L949
’ (Publication 3996)
UNITED STATES
GOVERNMENT PRINTING OFFICE
WASHINGTON 1950
For sale by the Superintendent of Documents, U. S. Government Printing Office
Washington 25, D. C. Price $2.75 (Buckram)
a oT Bk Ook AR ASNGS Vin Ag,
SMITHSONIAN INSTITUTION,
Washington, December 9, 1949.
To the Congress of the United States:
In accordance with section 5593 of the Revised Statutes of the
United States, I have the honor, in behalf of the Board of Regents,
to submit to Congress the annual report of the operations, expendi-
tures, and condition of the Smithsonian Institution for the year ended
June 30, 1949. I have the honor to be,
Respectfully,
A. Wrermore, Secretary.
BI
SEP ~ 11950
a Sehee Lienntl
CONTENTS
Page
Mii GeO bro LC ras tym nent Wings 4 Slvr els SE Aer hl en Sl cA ofS v
Generalstatement see = eeunke so Se eet Bo 3 Be ee Lt) 1
Presentation of the Wright Brothers’ Aeroplane of 1903 to the United
Statese Nationale Muse wit 2) 52 toe 2 oe eae eee ee ee Be ee 3
The establishment. ____..__-- ie kD Bay aioe ahd Leer ee aie alee ae SS a ao 5
Fiber BOasrdkotpecentcaa== pe tnt a5 as aie eee Bee a ee ey eee eee 6
PSSITY ATR COS eee ss AS Lh ay SR A 6 teen eoiiens = n a s ne e U
IND PLOPMAt ONS = aes execs Soe Pee ae ee eee ee 7
WASILODS tet eps. aps BL win a hal ue Peres SH hen Re nt UO ET OE BA 8
Sixteenth James Arthur annual lecture on the sun___.___.___________-- 8
Summary of the year’s activities of the branches of the Institution_______ 9
Publications. = 222s 252s=22< fel 8a README eo AOE TY Li elt OL ICRRMS Shee Oe | SLES 14
ALS Altay see ol lan rs ein 1s Rs apres a ae OE aks 2 DAA septs SRE Lhd 15
Appendix 1. Report on the United States National Museum____________ 16
2. Report on the National Gallery of Arti! (22 20) 2S 2 25
3. Report on the National Collection of Fine Arts___________- 41
4 Reportontthesreem Gallery of Artes = sees ee ee eee 47
5. Report on the Bureau of American Ethnology____________- 55
6. Report on the International Exchange Service___________-_- 89
7. Report on the National Zoological Park_-_-______-______-- 97
8. Report on the Astrophysical Observatory ---____________-- 109
9: Report on-the National Air Museumis22-2 2222-25252 -_ === 114
10. Report on the Canal Zone Biological Area__--____________- 126
ileReport, onyiheslibrarys=22 == 720 shoe ea eee eae 132
25 Repongion pPublcabiong==2 263.993 2225s. - ose ode eee 136
Report of the executive committee of the Board of Regents________-____- 143
GENERAL APPENDIX
hestormavionsolestarsbyaliyInanyoDILZCl dae ae eee 153
Aloe Cream OI ToS Capel, loyy ANoveriao leRVE ee eee enneeeoe 161
The 200-inch Hale telescope and some problems it may solve, by Edwin
Pole eas ee eee eye oe ee ee ee eye Ae ake 3 175
The determination of precise time, by Sir Harold Spencer Jones________- 189
The elementary particles of physics, by Carl D. Anderson_____-_____--- 203
Recent advances in virus research, by Wendell M. Stanley___-__-___--_-- 213
Ground-water investigations in the United States, by A. N. Sayre____--- 219
Modern soil science, by Charles E. Kellogg_.____._._-...---------------- 227
Aime in-ovolution bys EeZeUnere cejsee= oss se- Sao ao cal ee cess 247
More about animal behavior, by Ernest P. Walker___-__________------- 261
IV CONTENTS
Page
The breeding habits of the weaverbirds: a study in the biology of behavior
patterns, by dlerbert Priedmann--..= 222252522 -s2 oo See eee ee 293
New Zealand, a botanist’s paradise, by Egbert H. Walker______________ 317
The archeological importance of Guatemala, by A. V. Kidder__________- 349
Excavations at the prehistoric rock-shelter of La Colombiére, by Hallam
MGSO WIS Bisa 5, olan eo Ds ee ee ee ee pee 359
Ronne Antarctic research expedition, 1946-1948, by Commander Finn
Ronne aU Ore Neg hes. oe a te 2 Se ee ee ee ee pe ee 369
whe staterot science, by Karl. ©, Compton=c- 22- = ee eee 395
LIST OF PLATES
Secretary 6. report: Lu atOs two: ess sk ee 2s Se a eee ee 54
ihe formation of stars (Spitzer blates) jae 2) fe see ee ee one ee 160
Whe origin of theyearth.@Page):, Plates Iai! ss. 2ei eee Se 166
The 200-inch Hale telescope (Hubble): Plates 1-10________________-_-- 182
Modern soil science (Kellogg): Plates 1,2. 225-5. 8 ee 246
Animal behavior (Ernest Walker): Plates 1-16._..._..........._.._--- 278
The breeding habits of the weaverbirds (Friedmann): Plates 1-8_______- 310
New Zealand, a botanist’s paradise (Egbert Walker): Plates 1-10______- 326
The archeological importance of Guatemala (Kidder): Plates 1-6__-_____- 358
Excavations at La Colombiére (Movius): Plates 1-7_____.._--_________- 366
Ronne Antarctic research expedition (Ronne): Plates 1-8_----_____--_--- 374
THE SMITHSONIAN INSTITUTION
June 30, 1949
Presiding Officer ex officto— Harry S. Truman, President of the United States.
Chancellor.—Frep M. Vinson, Chief Justice of the United States.
Members of the Institution:
Harry 8S. Truman, President of the United States.
ALBEN W. BaRKLEY, Vice President of the United States.
Frep M. Vinson, Chief Justice of the United States.
Grorecs C. MARSHALL, Secretary of State.
Joun W. Snyper, Secretary of the Treasury.
Louis JoHNSON, Secretary of Defense.
Tom C. Cuarx, Attorney General.
JessE M. Donatpson, Postmaster General.
Jutius A. Krua, Secretary of the Interior.
CHARLES F, BRANNON, Secretary of Agriculture.
CHARLES SAWYER, Secretary of Commerce.
Maurice Tosin, Secretary of Labor.
Regents of the Institution:
Frep M. Vinson, Chief Justice of the United States, Chancellor.
ALBEN W. BARKLEY, Vice President of the United States.
Cuinton P. ANDERSON, Member of the Senate.
LEVERETT SALTONSTALL, Member of the Senate.
Wat7TEerR F. Grorce, Member of the Senate.
CLARENCE Cannon, Member of the House of Representatives.
E. E. Cox, Member of the House of Representatives.
Joun M. Vorys, Member of the House of Representatives.
Harvey N. Davis, citizen of New Jersey.
Arraur H. Compton, citizen of Missouri.
VANNEVAR Busu, citizen of Washington, D. C.
Rosert V. Fueming, citizen of Washington, D. C.
JEROME C. HuNSsAKER, citizen of Massachusetts.
Executive Commitiee—Ropert V. FLEMING, chairman, VANNEVAR
CLARENCE CANNON.
Secretary — ALEXANDER WETMORE,
Assistant Secretary. JOHN E. Grar.
Assistant Secretary.—J. L. Keppy.
Administrative assistant to the Secretary.—LouisE M, PEARSON.
Treasurer.—J. D. Howarp.
Chief, editorial division. WeEBsTER P. TRUE.
Librarian.—Leiua F, Cuarx.
Administrative accountant.—TuHomas F, Cuark.
Superintendent of buildings and labor.—L. L. OLIVER.
Personnel officer —B. T. CARWITHEN.
Chief, division of publications.—L. E. COMMERFORD.
Property, supply, and purchasing officer—ANTHONY W, WILDING.
Photographer.—F. B. Kustner.
Busi,
VI ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
UNITED STATES NATIONAL MUSEUM
Director.—A. REMINGTON KELLOGG.
Chief, office of correspondence and records—HrLeNA M. WEIss.
Editor.—Pauu H. OEHSER.
Assistant librarian.— ELIsABETH H. Gazin.
SCIENTIFIC STAFF
DEPARTMENT OF ANTHROPOLOGY:
Frank M. Setzler, head curator; A. J. Andrews, chief preparator.
Collaborator in anthropology: W. W. Taylor, Jr.
Division of Archeology: Neil M. Judd, curator; Waldo R. Wedel, associate
curator; M. C. Blaker, scientific aid; J. Townsend Russell, honorary
assistant curator of Old World archeology.
Division of Ethnology: H. W. Krieger, curator; J. C. Ewers, associate curator;
C. M. Watkins, associate curator; R. A. Elder, Jr., assistant curator.
Division of Physical Anthropology: T. Dale Stewart, curator; M. T. Newman,
associate curator.
DEPARTMENT OF ZOOLOGY:
Waldo L. Schmitt, head curator; W. L. Brown, chief taxidermist; Aime
M. Awl, scientific illustrator.
Associates in Zoology; T. 8S. Palmer, W. B. Marshall, A. G. Béving, C. R.
Shoemaker, W. K. Fisher.
Collaborator in Zoology: R. 8. Clark.
Collaborator in Biology: D. C. Graham.
Division of Mammals: D. H. Johnson, associate curator; H. W. Setzer, asso-
ciate curator; N. M. Miller, museum aid; A. Brazier Howell, collaborator;
Gerrit S. Miller, Jr., associate.
Division of Birds: Herbert Friedmann, curator; H. G. Deignan, associate
curator; Alexander Wetmore, custodian of alcoholic and skeleton collec-
tions; Arthur C. Bent, collaborator.
Division of Reptiles and Amphibians: Doris M. Cochran, associate curator.
Division of Fishes: Leonard P. Schultz, curator; E. A. Lachner, associate
curator; L. P. Woods, associate curator; D. 8. Erdman, scientific aid; W.
T. Leapley, museum aid.
Division of Insects: L. O. Howard, honorary curator; Edward A. Chapin,
curator; R. E. Blackwelder, associate curator; W. D. Field, associate
curator; O. L. Cartwright, associate curator; Grace E. Glance, associate
curator; W. L. Jellison, collaborator.
Section of Hymenoptera: S. A. Rohwer, custodian; W. M. Mann, assist-
ant custodian; Robert A. Cushman, assistant custodian.
Section of Diptera: Charles T. Greene, assistant custodian.
Section of Coleoptera: L. L. Buchanan, specialist for Casey collection.
Section of Lepidoptera: J. 'T. Barnes, collaborator.
Division of Marine Invertebrates: F. A. Chace, Jr., curator; P. L. Illg, asso-
ciate curator; Frederick M. Bayer, assistant curator; L. W. Peterson,
G. S. Cain, museum aids; Mrs. Harriet Richardson Searle, collaborator;
Max M. Ellis, collaborator; J. Percy Moore, collaborator; Mrs. M. 8.
Wilson, collaborator in copepod Crustacea.
Division of Mollusks: Harald A. Rehder, curator; Joseph P. E. Morrison,
associate curator; R. Tucker Abbott, assistant curator; W. J. Byas, museum
aid; P. Bartsch, associate.
Section of Helminthological Collections: Benjamin Schwartz, collabo-
rator,
Division of Echinoderms: Austin H. Clark, curator.
SECRETARY’S REPORT VII
DEPARTMENT OF Borany (Nationant HERBARIUM):
EK. P. Killip, head curator; Henri Pittier, associate in botany.
Division of Phanerogams: A. C. Smith, curator; E. C. Leonard, associate
curator; E. H. Walker, associate curator; Lyman B. Smith, associate
curator; V. E. Rudd, assistant curator.
Division of Ferns: C. V. Morton, curator.
Division of Grasses: Jason R. Swallen, curator; Agnes Chase, research asso-
ciate; F. A. McClure, research associate.
Division of Cryptogams: E. P. Killip, acting curator; Paul S. Conger, asso-
ciate curator; G. A. Llano, associate curator; John A. Stevenson, custodian
of C. G. Lloyd mycological collections; W. T. Swingle, custodian of Higher
Algae; David Fairchild, custodian of Lower Fungi.
DEPARTMENT OF GEOLOGY:
W. F. Foshag, head curator; J. H. Benn, exhibits preparator; Jessie G.
Beach, aid.
Division of Mineralogy and Petrology: W. F. Foshag, acting curator; E. P.
Henderson, associate curator; G. S. Switzer, associate curator; F. E.
Holden, exhibits preparator; Frank L. Hess, custodian of rare metals and
rare earths.
Division of Invertebrate Paleontology and Paleobotany: Gustav A. Cooper,
curator; A. R. Loeblich, Jr., associate curator; David Nicol, associate
curator; W. T. Allen, L. Pendleton, museum aids; J. Brookes Knight,
research associate in Paleontology.
Section of Invertebrate Paleontology: T. W. Stanton, custodian of
Mesozoic collection; J. B. Reeside, Jr., custodian of Mesozoic collection.
Division of Vertebrate Paleontology: C. L. Gazin, curator; D. H. Dunkle, asso-
ciate curator; Norman H. Boss, chief exhibits preparator; W. D. Crockett,
scientific illustrator; A. C. Murray, F. L. Pearce, preparators.
Associates in Mineralogy: W. T. Schaller, S. H. Perry, J. P. Marble.
Associates in Paleontology: T. W. Vaughan, R. S. Bassler.
DEPARTMENT OF ENGINEERING AND INDUSTRIES:
Frank A. Taylor, head curator.
Division of Engineering: Frank A. Taylor, acting curator.
Section of Civil and Mechanical Engineering: Frank A. Taylor, in charge.
Section of Marine Transportation: Frank A. Taylor, in charge.
Section of Electricity: K. M. Perry, associate curator.
Section of Physical Sciencesand Measurement: Frank A. Taylor, in charge.
Section of Land Transportation: S. H. Oliver, associate curator.
Division of Crafts and Industries: W. N. Watkins, curator; F. C. Reed,
associate curator; E. A. Avery, museum aid; F. L. Lewton, research
associate.
Section of Textiles: G. L. Rogers, assistant curator.
Section of Wood Technology: William N. Watkins, in charge.
Section of Manufactures: F. C. Reed, in charge.
Section of Agricultural Industries: F. C. Reed, in charge.
Division of Medicine and Public Health: G. S. Thomas, associate curator.
Division of Graphic Arts: J. Kainen, curator; E. J. Fite, museum aid.
Section of Photography: A. J. Wedderburn, Jr., associate curator.
DEPARTMENT OF History:
Charles Carey, acting head curator; T. T. Belote, Museum historian.
Divisions of Military History and Naval History: M. L. Peterson, associate
curator; J. R. Sirlouis, assistant curator.
Division of Civil History: M. W. Brown, assistant curator.
Division of Numismatics: 8S. M. Mosher, associate curator.
Division of Philately: C. L. Manning, assistant curator.
VIII ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
NATIONAL GALLERY OF ART
Trustees:
Frep M. Vinson, Chief Justice of the United States, Chairman.
Grorcse C. MarsHatt, Secretary of State.
Joun W. Snyper, Secretary of the Treasury.
ALEXANDER WETMORE, Secretary of the Smithsonian Institution.
SaMvuEL H. Kress.
FERDINAND LAMMOT BELIN.
DuNCAN PHILLIPS.
CHESTER DALE.
Paut MELLon.
President.—SAMUEL H. Kress.
Vice President—FERDINAND LAMMOT BELIN.
Secretary-Treasurer.—HUNTINGTON CAIRNS.
Director.—Davip E. FINLEY.
Administrator.—Harry A. McBrive.
General Counsel. HUNTINGTON CAIRNS.
Chief Curator—JoHN WALKER.
Assistant Director.—MacGILL JAMES.
NATIONAL COLLECTION OF FINE ARTS
Director.—Tuomas M. Braces.
Curator of ceramics.—P. V. GARDNER.
Exhibits preparator.—G. J. Martin.
Assistant librarian.— ANNA M. LINK.
FREER GALLERY OF ART
Director.—A. G. WENLEY.
Assistant Director.—JoHN A. PoPpE.
Associate in Near Eastern art.—RicHARD ETTINGHAUSEN.
Associate in Far Eastern art.—W. R. B. AcKER.
Research associate—Gracr DUNHAM GUEST.
BUREAU OF AMERICAN ETHNOLOGY
Director.—MatTTHEWw W. STIRLING.
Associate Director —F RANK H. H. Roserts, Jr.
Senior ethnologists—H. B. Couuins, Jr., Joan P. Harrineton, W. N. FENTON.
Senior anthropologists —G. R. Wituny, P. Drucker.
Collaborators —FRANcES Densmore, JoHn R. Swanton, A. J. WaRina, Jr.
Editor—M. HELEN PALMER.
Assistant librarian.—Miriam B. KetcHum.
Scientific illustrators —Epwin G. Cassnpy, E. G. SCHUMACHER.
Archives assistant.—M. W. Tucker.
INsTITU1E OF SocraL ANTHROPOLOGY.—G. M. Fostrmr, Jr., Director.
River Basin SurvEyS.—F RANK H. H. Rosmrts, Jr., Director.
INTERNATIONAL EXCHANGE SERVICE
Chief.—D. G. WILLIAMs.
SECRETARY’S REPORT Ix
NATIONAL ZOOLOGICAL PARK
Director.—Wiiu1aAM M. Mann.
Assistant Director.—Ernest P. WALKER.
Head Keeper.—F Rank O. Lowe.
ASTROPHYSICAL OBSERVATORY
Director.—Loyat B. ALDRICH.
Assistant librarian.— Marjorie KuNzE.
Division oF ASTROPHYSICAL RESEARCH:
Chief.—Wi.ii1amM H. Hoover.
Instrument makers.—ANDREW KRAMER, D. G. Tatsert, J. H. Harrison,
Research associate-—CHARLES G, ABBOT.
DIvIsION OF RADIATION AND ORGANISMS: ~
Chief —R. B. WirHrow.
Plant physiologist (physicochemical). —LrONARD PRICE.
Biological aid (botany).—Y. B. Ensrap.
NATIONAL AIR MUSEUM
Advisory Board:
ALEXANDER WETMORE, Chairman.
Mas. Gen. GRANDISON GARDNER, U.S. Air Force.
Rear Ap. A. M. Prinz, U. S. Navy.
Grover LOENING.
WixuiaM B. Srovut.
Assistant to the Secretary for the National Air Musewm.—Caru W. Mirman,
Curator.—P. E. GARBER.
Associate curators.—S. L. Brrers, R. C. Srrospet, W. M. Mates.
Exhibits preparator.—S. L. Porter.
CANAL ZONE BIOLOGICAL AREA
Resident Manager.—Jamus ZETEK.
ty o Lien 7
ow
REPORT OF THE SECRETARY OF THE SMITH-
SONIAN INSTITUTION
ALEXANDER WETMORE
FOR THE YEAR ENDED JUNE 30, 1949
To the Board of Regents of the Smithsonian Institution:
GENTLEMEN: I have the honor to submit herewith my report show-
ing the activities and condition of the Smithsonian Institution and
its bureaus during the fiscal year ended June 30, 1949.
GENERAL STATEMENT
The Institution continued vigorously to pursue its program of
activities in ‘‘the increase and diffusion of knowledge” as stipulated
by its founder, James Smithson. The increase of knowledge is fos-
tered by original scientific researches and explorations in the fields
of anthropology, biology, geology, and astrophysics; the diffusion of
knowledge, by publications in a number of series that are distributed
free to libraries and educational institutions throughout the world,
by extensive museum and art gallery exhibits, by the International
Exchange Service for the world-wide interchange of scientific and
governmental publications, and by a large correspondence, both
national and international.
I present first certain general features of the year’s activities,
together with a summary of the work of the several bureaus of the
Institution, to afford a concise picture of the events of the year.
Next follow appendixes containing more detailed reports on each
bureau, and finally there appears the financial statement of the Execu-
tive Committee of the Board of Regents. The appendixes contain
reports on the United States National Museum, the National Gallery
of Art, the National Collection of Fine Arts, the Freer Gallery of
Art, the Bureau of American Ethnology, the International Exchange
Service, the National Zoological Park, the Astrophysical Observatory,
the National Air Museum, the Canai Zone Biological Area, the Smith-
sonian library, and the publications of the Institution.
1
2 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
When the Smithsonian Institution began its operations more than
one hundred years ago, it carried on its research programs largely by
subsidizing the work of scientists not on its own staff, and by publish-
ing the results of their work. As these pioneer researches expanded
in scope and became somewhat stabilized, bureaus gradually grew up
around the Institution, each with its own staff specializing in the
work of that particular field. The value of the various activities
gradually became known to the Nation, and eventually one by one
they were recognized as public necessities by the Congress. Most of
them are now supported largely by Government funds although re-
maining under Smithsonian direction. At present, nearly all the
research and exploration of the Institution is done through these
bureaus, notably the United States National Museum, the Bureau
of American Ethnology, and the Astrophysical Observatory.
As stated in last year’s report, the Institution has for many years
operated under the handicap of shortages of personnel and of ade-
quate housing space. I reported that the Smithsonian Institution
has today the same amount of space that it had in 1911 in which to
accommodate four times as many visitors and four times as many
museum specimens. Much the same condition still prevails. Some
slight gain was apparent in personnel in a few of the scientific divi-
sions, but not sufficient for the prompt execution of essential cura-
torial work and adequate research on the National collections. The
crowded condition, particularly in the buildings of the National
Museum, remained unalleviated. In the report of the Director of
the Museum it will be noted that there is a considerable decrease in
number of specimens accessioned during the year, a decrease which,
he says, “may be attributed in part to the inadequacy of available
storage facilities for the preservation of such materials.”’ More ade-
quate building space is one of our major needs.
Though hampered by space conditions it should be brought to
attention that the Smithsonian Institution continues to grow and to
expand its usefulness year by year. In the 5 years during which I
have served as Secretary, three additional activities have been added
to its responsibilities—the Canal Zone Biological Area, the National
Air Museum, and the River Basin Surveys, the latter a unit of the
Bureau of American Ethnology. The work of these new activities
has notably augmented Smithsonian efforts toward the increase and
diffusion of knowledge in widely diversified fields, as will be seen in
reading the detailed reports appended hereto. The purpose in calling
attention to deficiencies is to emphasize the obvious fact that a growing
SECRETARY’S REPORT 3
institution such as the Smithsonian, of so vital interest and importance
to the American people, must receive increased financial support if
it is to continue to meet its full obligations and to further the high
ideals of its founder, James Smithson, who left his entire fortune in
trust to the United States of America for the benefit of all mankind;
PRESENTATION OF THE WRIGHT BROTHERS’ AEROPLANE OF 1903
TO THE UNITED STATES NATIONAL MUSEUM
On December 17, 1948, the forty-fifth anniversary of the first flight
by Wilbur and Orville Wright at Kitty Hawk, N. C., the original
aeroplane that made that historic flight became the property of the
American people. At a formal ceremony in the Museum attended
by many high civil and military officials the plane was presented to
the United States National Museum by Milton Wright on behalf of
the estate of Orville Wright.
The story of the plane goes back to December 17, 1903, when the
Wright Brothers were ready after several years of research and
experiment to test out their gasoline-engine-powered biplane at
Kitty Hawk on the coast of North Carolina. With Orville at the
controls, the machine was released, and after a 40-foot run on the
launching track, it lifted into the air in full flight. In Orville Wright’s
own words:
“The flight lasted only 12 seconds, but it was nevertheless the first
in the history of the world in which a machine carrying a man had
raised itself by its own power into the air in full flight, had sailed
forward without reduction of speed, and had finally landed at a point
as high as that from which it started.”
Three more flights were made the same day, but after the last
flight a strong gust of wind turned the plane over, damaging it so
badly that no more trials were made that year. The damaged
machine and engine were sent back to the Wrights’ workshop in
Dayton, and 13 years later were restored, using all the original parts
available. The aeroplane was displayed at the Massachusetts
Institute of Technology and later at several aeronautical exhibitions.
In 1928 Orville Wright had it sent as a loan to the Science Museum
at South Kensington, London, England, where it remained on exhibi-
tion until World War II. Owing to the danger of damage by bomb-
ing, the plane was removed to a safe place for the duration of the war,
When Orville Wright died on January 30, 1948, it was learned
from papers in his files that he wished the Kitty Hawk aeroplane to
be returned to the United States and placed in the National Museum.
The executors of his estate conferred with officials of the Science
4 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Museum and of the Smithsonian Institution, and with the generous
cooperation of the British Government the actual transfer of the
plane took place in November 1948. It was brought across the
Atlantic to Halifax on the Mauretania, from there to Bayonne,
N.J., on the Navy carrier Palau, and to Washington by Navy truck.
At the formal presentation on December 17, 1948, the ceremonies
were opened by the Secretary of the Smithsonian Institution. After
the invocation by Maj. Gen. Luther D. Miller, Chief of Chaplains,
Department of the Army, and greetings by the Presiding Officer,
Chief Justice Fred M. Vinson, Chancellor of the Smithsonian Institu-
tion, a message from the President of the United States was read by
Col. Robert B. Landry, Air Force Aide to the President. His Britan-
nic Majesty’s Ambassador, Sir Oliver Franks, K. C. B., C. B. E.,
then spoke on “Britain and the Wright Brothers,” after which the
presentation of the aeroplane was made by Milton Wright, of Dayton,
Ohio, on behalf of the estate of Orville Wright. Mr. Wright told of
his boyhood recollections of his uncles’ bicycle shop where the Kitty
Hawk plane was fabricated, and concluded thus:
“The aeroplane means many things to many people. To some it
may be a vehicle for romantic adventure or simply quick transporta-
tion. To others it may be a military weapon or a means of relieving
suffering. To me it represents the fabric, the glue, the spruce, the
sheet metal, and the wire which, put together under commonplace
circumstances but with knowledge and skill, gave substance to
dreams and fulfillment to hopes.”
The aeroplane was accepted on behalf of the Smithsonian Institu-
tion by Chief Justice Fred M. Vinson, Chancellor of the Institution,
and the address of acceptance was given by Vice President-Elect
Alben W. Barkley, a regent of the Institution. In the course of his
address Mr. Barkley expressed one thought that doubtless was in the
minds of all participants in the ceremony:
“Tt is a matter of deep regret to all of us that Orville Wright could
not have been here today to see this wide public recognition of achieve-
ment, and receive in person the fitting acclaim to his brother, to
himself, and to their Kitty Hawk plane. We are grateful to all of
those who have made it possible to bring the plane back to its native
soil, and especially to the heirs of the estate of Orville Wright, for
depositing the Kitty Hawk machine here where all America will have
an opportunity to see it, and where all may do it fitting honor.”
The Kitty Hawk aeroplane now hangs suspended from the ceiling
of the north hall of the National Museum’s Aris and Industries
Building, where the presentation ceremony was held. Directly back
of the main entrance, the plane is the first object to meet the eyes of
SECRETARY’S REPORT 5
the thousands of visitors who throng the Museum daily. As thus
displayed it bears the following label:
The Original
WRIGHT BROTHERS’ AEROPLANE
The world’s first
power-driven heavier-than-air machine in which man
made free, controlled, and sustained flight
Invented and built by Wilbur and Orville Wright
Flown by them at Kitty Hawk, North Carolina
December 17, 1903
By original scientific research the Wright Brothers
discovered the principles of human flight
As inventors, builders, and flyers
they further developed the aeroplane, taught man to fly
and opened the era of aviation
Deposited by the Estate of Orville Wright
°
“The first flight lasted only twelve seconds, a flight very modest compared
with that of birds, but it was nevertheless the first in the history of the world in
which a machine carrying a man had raised itself by its own power into the air
in free flight, had sailed forward on a level course without reduction of speed, and
had finally landed without being wrecked. The second and third flights were a
little longer, and the fourth lasted 59 seconds covering a distance of 852 feet over
the ground against a 20 mile wind.’”—W1.LzBur and OrvILLE WriGcaHt.
(From Century Magazine, vol. 76, September 1908, p. 649.)
This is not the final resting place of the plane, however—it is
destined eventually to occupy the place of honer in the National Air
Museum, the most recent bureau of the Smithsonian Institution.
Preliminary plans for the Air Museum envision a special centrally
located exhibit area for the Wright aeroplane of 1903, to serve as a
memorial to the birth of aviation.
THE ESTABLISHMENT
The Smithsonian Institution was created by act of Congress in
1846, according to the terms of the will of James Smithson, of England,
who in 1826 bequeathed his property to the United States of America
“to found at Washington, under the name of the Smithsonian Insti-
tution, an establishment for the increase and diffusion of knowledge
among men.’ In receiving the property and accepting the trust,
Congress determined that the Federal Government was without
authority to administer the trust directly, and, therefore, constituted
an ‘establishment’? whose statutory members are “the President,
the Vice President, the Chief Justice, and the heads of the executive
departments.”
6 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
THE BOARD OF REGENTS
The following changes occurred during the year in the personnel of
the Board of Regents:
On January 20, 1949, Vice President Alben W. Barkley (formerly a
regent by appointment from the Senate) became ‘ex officio a member
of the Board.
On February 14, 1949, the following regents were appointed from
the House of Representatives: Clarence Cannon of Missouri; John M.
Vorys of Ohio; and E. E. Cox of Georgia to succeed Samuel K.
McConnell of Pennsylvania.
On March 8, 1949, Senators Leverett Saltonstall and Clinton P.
Anderson were appointed to succeed Vice President Alben W. Barkley
who became an ex officio member of the Board, and Senator Wallace
H. White of Maine, retired.
On March 10, 1949, Dr. Jerome C. Hunsaker was appointed a citizen
regent from Massachusetts for the statutory term of 6 years, to succeed
Frederic C. Walcott, retired.
The roll of regents at the close of the fiscal year, June 30, 1949,
was as follows:
Chief Justice Fred M. Vinson, Chancellor; Vice President Alben W.
Barkley; members from the Senate: Walter F. George, Clinton P.
Anderson, Leverett Saltonstall; members from the House of Repre-
sentatives: Clarence Cannon, John M. Vorys, E. E. Cox; citizen mem-
bers: Harvey N. Davis, Arthur H. Compton, Vannevar Bush, Robert
V. Fleming, and Jerome C. Hunsaker.
Proceedings.—The Board of Regents held its annual meeting on
January 14,1949. Present: Chief Justice Fred M. Vinson, Chancellor;
Representative Clarence Cannon, Representative John M. Vorys,
Dr. Arthur H. Compton, Dr. Harvey N. Davis, Dr. Robert V. Flem-
ing, Secretary Alexander Wetmore, and Assistant Secretary John E.
Graf.
The Secretary presented his annual report covering the activities
of the Institution and its bureaus, including the financial report of the
Executive Committee, for the fiscal year ended June 30, 1948, which
was accepted by the Board. The usual resolution authorizing the
expenditure by the Secretary of the income of the Institution for the
fiscal year ending June 30, 1950, was adopted by the Board.
It was announced that in support of the work of the Astrophysical
Observatory John A. Roebling had made a further generous gift
which is of major importance in carrying on these scientific investi-
gations.
The annual report of the Smithonsian Art Commission was pre-
sented by the Secretary and accepted by the Board. A resolution was
adopted to reelect the following members for 4-year terms: Archibald
SECRETARY’S REPORT he
G. Wenley, David E. Finley, Eugene E. Speicher, Paul Manship.
The following officers were reelected for the ensuing year: Chairman,
Paul Manship; vice chairman, Robert Woods Bliss; secretary,
Alexander Wetmore.
The Board was advised that in an attempt to recover the Gellatly
art collection from the Secretary in his status of a private individual,
though acting as custodian under the Smithsonian Institution, Mrs.
Charlayne Gellatly’s attorneys had filed action in the District Court
of the United States for the District of Columbia on June 18, 1947.
Under date of June 17, 1948, Judge J. McGuire rendered a decision
that, in the opinion of the Court, there was no merit in Mrs. Gellatly’s
claims, since it was found that there was a valid gift to the United
States by the deceased, John Gellatly, before his death and before his
marriage. On July 19, 1948, the attorney for Mrs. Gellatly filed notice
of appeal before the United States Court of Appeals for the District
of Columbia. Marvin C. Taylor, special attorney, Department of
Justice, represented the Institution.
On the evening of March 1, 1949, an informal meeting of the Board
was held at dinner in the Main Hall of the Smithsonian Building, with
the Chancellor, Chief Justice Fred M. Vinson, presiding. At this
meeting heads of the various activities under the Institution pre-
sented statements relative to their work. These statements, with the
ensuing discussion, provided a general view of the existing operations
of the Smithsonian, particularly in the research field.
FINANCES
A statement on finances, dealing particularly with Smithsonian
private funds, will be found in the report of the Executive Committee
of the Board of Regents, page 143.
APPROPRIATIONS
Funds appropriated to the Institution for the fiscal year ended June
30, 1949, totaled $2,259,000, allotted as follows:
@eneraleadministra tions sae see ee ee ee ee $46, 794
INationale@iuse ini fae ee ae eee ee ee eg 712, 560
Buresawvof American hthuolopy ss 222 22225 2 eee ee 71, 996
AstrophysicalObsernvatorye. sesso etek ot sete - caer 101, 590
INationalaCollectionsoteliner Arts == eee ee ee ee eee 32, 543
International Exchange Service=* 22 v. oul sas See be ee ole 68, 938
Maintenance and-OperaviOn. 2.052) nee eee eo eo 764, 626
Dervicerdivisions=seses see ee ee ee ee 274, 448
INationaleAirsMiniscn mee ee ei ee ee ee ee eee 180, 285
CanalkZoneybiolocicalwArca to 3 ins so = ee oe ee eee 4, 760
Wnallottedses: -tee See Seek! Wha BEM NE esa A ee ae a Oe 460
DLT C come 2 a ce ld Se pA ee rt eA Ne PE TR ea 2, 259, 000
866591—50——-2
8 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
In addition $1,073,500 was appropriated to the National Gallery of
Art, a bureau of the Institution but administered by a separate board
of trustees; and $528,848 was provided in the District of Columbia
appropriation act for the operation of the National Zoological Park.
Besides these direct appropriations, the Institution received funds
by transfer from other Federal agencies, as follows:
From the State Department, from the appropriation Cooperation
with the American Republics, 1949, a total of $97,900 for the operation
of the Institute of Social Anthropology, including the issuance of
publications resulting from its work.
From the National Park Service, Interior Department, $118,500
for archeological projects in connection with River Basin Surveys.
VISITORS
The number of visitors to the Smithsonian buildings for the year
was 2,606,104, an all-time record of attendance. This was an increase
of 212,605 over the previous year’s attendance. April 1949 was the
month of largest attendance with 371,871 visitors; August 1948, the
second largest with 313,364. Records for the five buildings show the
following number of visitors: Smithsonian, 494,880; Arts and In-
dustries, 1,148,303; Natural History, 689,233; Aircraft, 198,648;
Freer, 75,040.
A summary of attendance records is given in table 1:
TABLE 1.—Visttors to the Smithsonian buildings during the year ended June 80, 1949
Smith- Arts and Natural Adrorate Freer
Year and month sonian Industries | History Buildin Gallery Total
Building | Building | Building 8 of Art
1948
Duly a he oS ee eee 61, 529 128, 635 74, 243 24, 557 9, 510 298, 474
USUE ADC) a Say ee pe ee 65, 412 136, 704 75, 026 26, 672 9, 550 313, 364
September.a22 08-35 22 ee 45, 178 90, 321 61, 839 18, 460 7, 269 218, 067
Octobere-2)- 228s: fe Fast es 34, 460 66, 329 47, 962 13, 670 5, 460 167, 881
INovemberis= 2221 -iee ee 27, 380 50, 700 39, 829 11, 833 4,415 134, 157
December- 22s 222-22 22222-522 18, 242 42, 191 23, 419 8, 512 3, 153 95, 517
1949
JANUAIY See ee ee 26, 748 59, 837 37, 212 11, 085 4,124 139, 006
Mebruary 2 Seer aes See 22, 949 54, 470 35, 220 10, 842 4, 032 127, 513
Miarchste S220 eee ee 25, 650 66, 814 41, 452 12, 499 5, 092 151, 507
April see 32 ee eee 64, 804 177, 144 97,135 23, 532 9, 256 371, 871
VT yates oe Bate Mee non SOee ee 47, 718 142, 007 88, 029 19, 653 6, 172 303, 579
Jules. 5252222 s ses ean es 54, 810 133, 151 77, 867 17, 333 7, 007 290, 168
Total: === 442-5 3ee 494, 880 1, 148, 303 1689, 233 198, 648 75, 040 2, 606, 104
' Not including 31,249 persons attending meetings after 4:30 p. m.
SIXTEENTH JAMES ARTHUR ANNUAL LECTURE ON THE SUN
In 1931 the Institution received a bequest from James Arthur, of
New York, a part of the income from which was to be used for an
annual lecture on some aspect of the study of the sun.
SECRETARY’S REPORT 9
The sixteenth Arthur lecture was given in the auditorium of the
National Museum on April 14, 1949, by Sir Harold Spencer Jones,
Astronomer Royal of Great Britain, the arrangements being made
through Dr. S. A. Mitchell, of the Leander McCormick Observ-
atory, University of Virginia. The title of Sir Harold’s lecture was,
“The Determination of Precise Time,” a subject on which he is a
world authority. His lecture will be published in full in the Annual
Report of the Board of Regents of the Smithsonian Institution for 1949.
SUMMARY OF THE YEAR’S?ACTIVITIES OF THE BRANCHES OF
THE INSTITUTION
National Museum.—Approximately 446,000 specimens were added
to the collections, for the most part as gifts or as transfers from Gov-
ernment agencies, bringing the total number of catalog entries to
31,679,046. Outstanding accessions for the year included: In an-
thropology, an important collection of 51 artifacts representing the
work of American Indians, Eskimo of Alaska, and natives of Pacific
islands, given by Georgetown University; 17 gold-embossed silver
vessels given by the Government of Tibet to President Truman and
in turn presented by him to the Smithsonian Institution; and valu-
able skeletal remains recovered in northern Australia by Frank M.
Setzler, a member of the Commonwealth of Australia-National Geo-
graphic Society-Smithsonian Institution Expedition to Arnhem
Land; in zoology, maramal specimens from many distant parts of the
world including Northern Territory of Australia, Nepal, Malay Pen-
insula, Korea, Okinawa, Philippine Islands, and New Guinea, 778
birds from Arnhem Land, Australia, and 1,164 from India and Nepal,
14,000 fishes from the Solomon Islands and the East Indies, and
5,000 from the Persian Gulf and the Red Sea; in botany, 2,382 plants
of Fiji, 5,854 plants of Colombia, and 2,157 plants of China; in geology,
20 kinds of minerals hitherto unrepresented in the National collec-
tions, a 42-carat brazilianite gemstone, the largest ever found in
Brazil, an 8,750-gram stony meteorite that fell at Girgenti, Italy, and
many thousands of fossil specimens collected by staff members in
various parts of the United States; in engineering and industries,
the original Wright Brothers’ aeroplane of 1903, a collection of elec-
trical measuring instruments, early lamps, and electronic tubes, some
of them made in the 1880’s, and an exhibit showing the development
of electric hearing aids; in history, a group of relics bequeathed by
Gen. John J. Pershing, including uniforms, flags, and medals, a note-
worthy collection of European gold and silver coins from the four-
teenth to the twentieth century presented by Paul A. Straub, of
New York City, and a complete set of Allied military currency pre-
sented by the Department of the Army.
10 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Field work was conducted in Arnhem Land in northern Australia,
India and Nepal, the Persian Gulf and the Red Sea, New Zealand,
the Canadian Arctic, nine different countries in South and Central
America, and many parts of the United States. The Museum pub-
lished its Annual Report, 3 Bulletins, 25 Proceedings papers, and 2
papers in the Contributions from the United States National Herbar-
ium. The division of history was elevated to the status of a full
department of the Museum, with five divisions—military history,
naval history, civil history, numismatics, and philately.
National Gallery of Art——During the year there were 1,529,568
visitors to the Gallery, an average daily attendance of 4,225. Acces-
sions as gifts, loans, or deposits numbered 1,174, including 10 paint-
ings and 50 prints and drawings from the estate of the late R. Horace
Gallatin, and 891 prints and drawings from Lessing J. Rosenwald.
Eleven special exhibitions were held at the Gallery, and two traveling
exhibitions were circulated to art galleries, museums, and other organ-
izations throughout the country. In response to inquiries received
by the Gallery, nearly 1,000 research problems requiring reports were
investigated, and advice was given regarding 233 works of art brought
to the Gallery for opinion. Numerous books and articles on art sub-
jects were published by staff members. New publications continued
to be added to the literature available at the Gallery for purchase by
the public. Some 15,000 persons attended the special tours of the
Gallery, 20,000 the “Picture of the Week” talks, and 18,000 the lec-
tures in the auditorium. The Gallery’s collections of art works has
grown so fast that all available exhibition space was in use during
the year. To provide for expansion, contracts have been let for the
completion of 12 more galleries in unfinished areas of the Gallery
building. ‘Some 50,000 persons attended the 46 Sunday evening
concerts given in the Gallery’s East Garden Court.
National Collection of Fine Arts.—At the annual meeting of the
Smithsonian Art Commission of December 7, 1948, a number of
paintings were accepted for the National Collection. The Commis-
sion passed a resolution calling attention to the inadequacy of the
present art exhibition facilities in the National Museum and recom-
mending that the Secretary of the Smithsonian Institution take action
to provide proper space for the preservation and exhibition to the
public of the National Collection of Fine Arts. Two miniatures were
acquired through the Catherine Walden Myer fund. Under the
provisions of the Ranger bequest, seven paintings temporarily as-
signed to various art institutions were recalled for final consideration
by the Smithsonian Art Commission. Two of these paintings were
accepted for the National Collection, and the others were returned
to the institutions to which they were originally assigned. A large
SECRETARY'S REPORT 11
amount of information on art subjects was furnished to visitors in
person, as well as by mail and phone. Members of the staff lectured
on art topics to several organizations, and six special art exhibitions
were held during the year, for most of which catalogs were furnished
by the organizations sponsoring the exhibitions.
Freer Gallery of Art.—Additions to the collections included Chinese
bronze, jade, lacquer, marble, and painting; Syrian glass; Syrian or
Egyptian gold; Arabic manuscript; Persian manuscript, painting, and
stone sculpture; Indian painting; and Turkish painting. The work
of the professional staff was devoted to the study of new accessions
and to research within the collection of Chinese, Japanese, Iranian,
Arabic, and Indian materials. Reports were made upon 2,563 objects
and 372 photographs of objects submitted to the Gallery for examina-
tion, and 369 Oriental language inscriptions were translated. The
repair and restoration of the walls of Whistler’s Peacock Room were
completed early in the year, and work was begun on the ceiling.
Visitors to the Gallery numbered 74,846 for the year, and 1,724 came
to the Gallery offices for special purposes. Sixteen groups were given
instruction in the exhibition galleries by staff members, and 13 lec-
tures were given in art galleries and museums, before clubs, and to
various associations.
Bureau of American Ethnology.—Dr. M. W. Stirling, Director of
the Bureau, devoted 4 months to a continuation of his archeological
work in Panamé in cooperation with the National Geographic Society.
Heretofore undescribed ceramic cultures were found at Utivé and
Barriles, and much new information was obtained on the classic
Chiriqui and Veraguas cultures. Dr. Frank H. H. Roberts, Jr.,
contmued to direct from Washington the very extensive operations
of the River Basin Surveys, a unit of the Bureau created to rescue
important archeological sites threatened by the construction of dams
and the creation of river basin reservoirs. ‘The work was done in
cooperation with the National Park Service, the Bureau of Reclama-
tion, the Army Corps of Engineers, and local organizations. Surveys
of threatened sites covered 69 reservoir areas in 21 States. Since
the program started, 2,107 archeological sites have been located and
recorded, and of these, 456 have been recommended for excavation
or testing before they are destroyed by construction work. Dr.
John P. Harrington continued his revision of the Maya grammar.
Toward the end of the year he went to Old Town, Maine, to pursue
ethnological and linguistic studies on the Abnaki Indians. Dr. Henry
B. Collins, Jr., conducted archeological excavations at Frobisher Bay
on Baffin Island in the Canadian Arctic. Ruins were found of old
Eskimo semisubterranean houses made of stones, whale bones, and
turf, the evidence showing that the site has been occupied succes-
12 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
sively by Eskimos of both the prehistoric Dorset and Thule cultures.
Dr. William N. Fenton continued his field work and library research
on the Iroquois Jndians, obtaining the life history of an aged Seneca
and recording Seneca rituals, prayers, and legends. Dr. Gordon R.
Willey devoted the year to studying and writing up the results of
previous field work. His monographic work, “Archeology of the
Florida Gulf Coast,”’ was completed and at the close of the year was
in process of being published by the Smithsonian Institution.
The Institute of Social Anthropology, an autonomous unit of the
Bureau, is financed by State Department funds to carry out coopera-
tive training in anthropological teaching and research with the other
American republics. Institute staff members, under the directorship
of Dr. George M. Foster, Jr., continued to give courses in anthropology
and to conduct cooperative research and field work in Brazil, Colom-
bia, México, and Pert.
The Bureau published its Annual Report and two Publications of
the Institute of Social Anthropology. The last two volumes of the
Handbook of South American Indians, volumes 5 and 6, were in
press at the close of the year.
International Hachanges.—The Smithsonian International Exchange
Service is the official United States agency for the interchange of gov-
ernmental and scientific publications between this country and the
other nations of the earth. ‘The Exchange Service handled during the
year a total of 840,125 packages of publications, weighing 796,700
pounds. These figures represent an increase over the previous year
of 80,006 packages, but a decrease of 15,489 pounds in weight, indicat-
ing by the lighter weight per package that most institut'ons have about
completed shipment of material held up during the war. Shipments
are now made to all countries except Rumania, and efforts to resume
exchanges with that country are being continued. The number of
sets of United States official publications sent abroad in exchange for
similar publications of other countries is now 96—58 full and 38
partial sets. There are also sent abroad through the Exchange
Service 81 copies of the Federal Register and 75 copies of the Con-
gressional Record.
National Zoological Park.—The collection was improved during the
year by the addition of a number of rare animals. At the close of the
fiscal year there were 3,724 specimens in the collection, an increase
of 927 over the previous year. These represented 755 different species,
an increase of 65. Among the rare or unusual animals received by
gift, exchange, or purchase were the rare Meller’s chameleon, a
spectacled bear, a pair of pigmy marmosets—smallest of all monkeys,
an African two-horned rhinoceros, a pair of wombats, a pigmy ant-
eater, orang-utans, and chimpanzees. The total number of creatures
SECRETARY'S REPORT 1433
born or hatched at the Zoo was 157—56 mammals, 62 birds, and 39
reptiles. Personne! recruitment and training for the organization
progressed satisfactorily, and the most needed repairs and minor im-
provements to buildings and grounds were carried out. The year’s
total of visitors to the Zoo was the largest ever recorded—3,346,050,
an increase of more than 300,000 over the previous year. Groups
from schools, some as far away as Maine, Florida, Texas, and Cali-
fornia, numbered 1,844, aggregating 93,632 individuals.
Astrophysical Observatory —Year-long tests at the three most
promising sites for a new high-altitude solar observing station indicate
that the best skies prevail at the Clark Mountain, Calif., site, the
second-best site being Pohakuola, Hawaii. Estimates of the cost of
establishing a field station on Clark Mountain, however, proved to be
in excess of available funds, forcing postponement of building opera-
tions. Data and tables were prepared which simplify computations
at the field observing stations by eliminating the tedious curve-plotting
process heretofore used in obtaining the air mass. Daily observations
of the solar constant of radiation were continued at the Montezuma,
Chile, and Table Mountain, Calif., stations. Intercomparisons
between the substandard silver-disk pyrheliometer S. I. No. 5 and the
instruments in use at Miami, Montezuma, and Table Mountain show
no material changes in constants, confirming the adopted scale of
pyrheliometry. Special radiation measurements started in 1945 at
Camp Lee, Va., under contract with the Office of the Quartermaster
General, were continued there, half of the year by the Observatory
and half by the Quartermaster Board; similar measurements were also
made at Miami, Fla., and at Montezuma, Chile. The work of the
Division of Radiation and Organisms has been concerned chiefly with
reorganizing and reequipping the laboratories. Besides new office
space which has been established, five rooms are being converted into
constant-condition rooms for biological experimentation, and four
chemistry laboratories will be available. In addition, a photographic
laboratory, an X-ray room, a cytology laboratory, an electronics
laboratory, and two general laboratories are being arranged.
National Air Museum.—The Air Museum was given the responsi-
bility of receiving, bringing to Washington, and preparing for exhibi-
tion the original Wright Brothers aeroplane of 1903, presented to the
National Museum in December 1948. A storage depot to be used by
the Air Museum until it has a building of its own was acquired in
November 1948 in the former Douglas Aircraft plant at Park Ridge,
Ill. There a field organization was installed, and the Air Museum
assumed custody of the storage facility itself and the large collection
of planes and other aeronautical material stored there by the U. S.
Air Force for the Museum. The Advisory Board held three meetings
14 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
during the year which were devoted mainly to advancing the acquisi-
tion of a building site and a suitable museum building in the Wash-
ington area. The Museum expects during the coming year to sub-
mit to Congress a report regarding sites and a building, the prelim-
inary study of which has been prepared in cooperation with the Public
Buildings Administration. Among outstanding accessions of the
year were the Swoose, the historic B~17—D bomber that served through-
out World War II from Bataan to the defeat of Japan, presented by
the city of Los Angeles; Maj. Alford Williams’ renowned Gulfhawk-2
presented by the Gulf Oil Co.; a Japanese Baka Bomb, or “suicide
plane,” transferred by the Department of the Navy; and 10 scale
models of recent types of Naval aircraft received from the manufac-
turers who produced the original planes. New accessions totaled
122 objects from 40 different sources.
Canal Zone Biological Area.—A new building for woodworking and
carpenter shops and for living quarters for the warden-caretaker was
completed during the year, the old quarters being converted into a
two-room laboratory unit. Work on the new 14,000-gallon water
tank was halted by heavy rains but can be completed with 2 or 3
weeks of dry weather. The most urgent needs are the fireproofing of
existing buildings and the construction of a new six-room laboratory
and storage building. Twenty-nine scientists representing many
different organizations worked at the laboratory during the year, and
their contributions have added materially to our knowledge of tropical
life. Among the interesting researches were the work of Drs. Scho-
lander and Walters of the Arctic Research Laboratory at Point Barrow,
Alaska, on the metabolic reactions to temperature in various animals
and plants in order to obtain a tropical counterpart for similar work
on Arctic forms in Alaska; the studies of Drs. Clark and Soper of the
Research Laboratory of Eastman Kodak on the effects of tropical
conditions on photographic equipment and materials, including color
film; and the Resident Manager’s own special studies, particularly the
long-term termite-resistance tests.
PUBLICATIONS
In carrying out the diffusion of knowledge, the Institution issues
eight regular series of publications and six others that appear less fre-
quently. All these series, embodying the results of Smithsonian
researches, are distributed free to more than a thousand libraries,
both here and abroad, as well as to a large list of educational and
scientific organizations. The findings of Smithsonian scientists,
chiefly in the fields of anthropology, biology, geology, and astrophysics,
are therefore made readily available to all through this wide free
distribution.
SECRETARY'S REPORT 15
A total of 71 separate volumes and pamphlets were issued during the
past year. Among the outstanding publications to appear were Dr.
Henry Field’s compilation in the Smithsonian Miscellaneous Collec-
tions entitled ‘Contributions to the Anthropology of the Soviet
Union,”’ which presents, for the first time in English, accounts of
recent findings in this little-known area; a revised edition of the popu-
lar handbook of the National Aircraft Collection, which is in effect a
brief history of aeronautics from the mythical flying horses of antiquity
to the supersonic jet planes of today; two more volumes in the famous
series of Life Histories of North American Birds, prepared by A. C.
Bent, bringing to 17 the number so far issued in the series; and a
paper by Jason R. Swallen on new grasses from several countries of
South and Central America, in the Contributions from the United
States National Herbarium.
The total number of copies of publications in all series distributed
during the year was 267,491. A complete list of the year’s publica-
tions will be found in the report of the Chief of the Editorial Division,
Appendix 12.
LIBRARY
Of the 57,671 publications received by the Smithsonian library
during the year, 7,287 came as gifts from many different donors.
Another 17,713 were periodicals mostly received in exchange for
Smithsonian publications from research institutions and other scien-
tific and educational organizations throughout the world. Containing
the record of progress in science and technology, these periodicals are
indispensable in the prosecution of the Institution’s own work.
Increasingly heavy demands upon reading and reference services
of the library were noted during the year, the interlibrary loans total-
ing 2,619 publications to 89 different libraries. The new position of
assistant librarian in charge of the Astrophysical Observatory library
was filled by the promotion of an acquisitions assistant.
New exchanges arranged during the year numbered 338; 6,884
volumes and pamphlets were cataloged, and 31,184 cards were added
to catalogs and shelflists; 1,060 volumes were sent to the bindery,
and 1,026 were repaired in the Museum.
At the close of the year, the library’s holdings totaled 921,206
volumes, more than half of which are housed in the Library of Congress
as the Smithsonian Deposit.
Respectfully submitted.
ALEXANDER WETMORE, Secretary.
APPENDIX 1
REPORT ON THE UNITED STATES NATIONAL MUSEUM
Srr: I have the honor to submit the following report on the condi-
tion and operations of the United States National Museum for the
fiscal year ended June 30, 1949.
COLLECTIONS
Approximately 446,000 specimens (88,000 less than last year)
were incorporated into the National collections during the year and
were distributed among the six departments as follows: Anthropology,
4,099; zoology, 279,621; botany, 38,708; geology, 109,499; engineering
and industries, 2,610; and history, 11,104. The decrease in the number
of specimens accepted for the Museum’s collections may be attributed
in part to the inadequacy of available storage facilities for the preser-
vation of such materials; consequently, a finer screening of collections
from prospective donors is now mandatory. Most of the accessions
were acquired as gifts from individuals or as transfers from Govern-
ment departments and agencies. The complete report on the Museum,
published as a separate document, includes a detailed list of the year’s
acquisitions, of which the more important are summarized below.
Catalog entries in all departments now total 31,679,046.!
Anthropology.—The most noteworthy additions to the archeological
collections were as follows: A black-figured Attic lecythus of the fifth
century, B. C., presented to President Harry S. Truman as a token of
gratitude from the people of Greece and lent by the President; 11
gold-plated ornaments from Veraguas, Panam4, and 2 gold fishhooks
from Colombia, a gift from Karl P. Curtis; and 47 prehistoric earthen-
ware vessels from the Valley of Nasca, Peri, presented to the late
Gen. John J. Pershing by former Peruvian President Augusto B.
Leguia and donated by General John J. Pershing.
Handicrafts and material culture of many of the world’s peoples
were represented in the additions to the ethnological collections. An
unusually important collection of 51 specimens representing the work
of American tribes of the Great Plains and the Great Lakes, of Arizona
and New Mexico, as well as of the Eskimo of Alaska, of the Igorot of
the Philippine Islands, and of the Marquesans and Maori of the
1 The revised tabulation of the National collection of insects during the past year, in addition to the
normal increment, has increased last year’s total by more than 4,400,000 specimens.
16
SECRETARY'S REPORT 17
Southeast Pacific, assembled over a period of more than a century,
was presented by Georgetown University. President Harry 8S. Tru-
man presented to the Smithsonian Institution 17 gold-embossed silver
vessels received at the White House as a gift from the Government
of Tibet in appreciation of an American gift of wireless receiving and
transmitting sets made during World War II. Included are two
butter lamps and stands, four teacup stands and covers, two bowls
for grain offerings, one teapot, and two beer mugs, all decorated in
gold-embossed designs derived from Chinese-Tibetan folklore and
Buddhist religious art. A collection of 287 folk, costume, and his-
torical portrait dolls, representing the native dress of peoples of many
lands, was received as a bequest from the late Mrs. Frank Brett Noyes.
The Don Diego Columbus mahogany table, traditionally known as
the writing desk of Diego Columbus, was conditionally bequeathed
by Mrs. Edith Keyes Benton. This table had been preserved for
centuries in the cathedral of Santo Domingo City and was presented
by Archbishop Nouel to Commander Frederick L. Benton, U.S.N., in
recognition of his work in Santo Domingo during the influenza epidemic
of 1918. One of the rarest of musical instruments, a musical gong,
kyung, carved from white marble, was presented by Ju Whan Lee,
director of the Korean Court Music Conservatory at Seoul, Korea.
The largest accession received by the division of physical anthro-
pology consisted of the skeletal remains recovered in northern Australia
by Frank M. Setzler, a member of the Commonwealth of Australia-
National Geographic Society-Smithsonian Institution Expedition to
Arnhem Land. Australian skeletal material available for study in
the United States is rather limited. Four casts of African fossil
primates, which illustrate certain characteristics of antecedent special-
ization, were also acquired during the year.
Zoology.—The collections made by the Museum staff detailed to the
Arnhem Land field expedition, under the joint sponsorship of the
Commonwealth of Australia, National Geographic Society, and the
Smithsonian Institution, have added many previously unrepresented
forms of animal life to the National collections. These collections
included not only vertebrates but invertebrates as well.
Accessions that enhanced the usefulness of the mammalian collec-
tion came from the Northern Territory of Australia, Nepal, Malay
Peninsula, Korea, Okinawa, Philippine Islands, New Guinea, and New
Hampshire. Field work financed wholly or in part by the W. L.
Abbott fund resulted in the addition of birds not hitherto represented
in the National collection. Included among these accessions were
2,815 skins and 388 eggs of Colombian birds; 900 skins, 24 skeletons, and
2 sets of eggs of Panamanian birds; 778 bird skins, many of which were
not represented in the collection, as well as 51 skeletons and 2 eggs
18 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
from Arnhem Land, Australia; and 1,164 bird skins procured by the
joint National Geographic Society-Yale University-Smithsonian
Institution Expedition to India and Nepal. Other accessions com-
prised 611 bird skins from Nyasaland: 177 birds and 1 egg from
northeastern Venezuela: 171 bird skins from Pacific War areas; and
125 bird skins from Korea.
Snakes, lizards, and frogs from Arnhem Land, amphibians from
Perit, reptiles and amphibians from Honduras, and a general collection
from Virginia and North Carolina constituted the most important
additions to the herpetological collection.
The most noteworthy accessions received by the division of fishes
were nearly 14,000 specimens from the Solomon Islands and the East
Indies, which were presented by Dr. Wilbert M. Chapman; 14,300
from Arnhem Land; and approximately 5,000 from the Persian Gulf
and the Red Sea, resulting from a survey sponsored by the Arabian-
American Oil Co. Other important collections of fishes came from
Puerto Rico, Panam4, British Columbia, and Florida.
Approximately 25,000 miscellaneous insects from South Pacific
Islands came to the Museum by transfer from the U. S. Commercial
Co. Among other large lots were approximately 12,000 flies; 3,500
chalcidoid wasps; 500 beetles; and some 53,000 insects transferred
from the United States Bureau of Entomology and Plant Quarantine.
During the year considerable significant material was added to the
marine invertebrate collection, of which the most important accessions
were 11,765 miscellaneous invertebrates from the Department of
Zoology, University of California; 70 lots of paratypes, hypotypes,
and topotypes of hydroids from the Allan Hancock Foundation,
University of Southern California; 760 marine invertebrates from
California and Mexice; 709 specimens from Bahama Islands; 1,781
from Pacific Islands and California; 452 from the Persian Gulf and
the Red Sea; and 859 from Arnhem Land. By transfer from the
Office of Naval Research, the Museum acquired 3,668 invertebrates
from Point Barrow, Alaska. The United States Geological Survey
transferred 568 specimens from the Marianas Islands.
A rare deep-water Pleurotomaria, dredged at a depth of 160 fathoms
off Natal, South Africa, and presented by Dr. Cecil von Bonde, con-
stituted the most notable accession received by the division of
mollusks. From other sources the division received 250 Peruvian
terrestrial and fresh-water mollusks and 540 marine mollusks from
Canton Island, and 150 Japanese land mollusks. Exchanges brought
to the Museum approximately 1,080 shells from Spain and lesser
numbers from South Africa, Italy, and Cuba. By transfer the Museum
received about 1,200 mollusks obtained in the Caroline Islands from
the United States Geological Survey; approximately 30,600 specimens
SECRETARY’S REPORT 19
from the Naval Medical Research Institute; and 600 marine and land
shells of the Solomon Islands from the Naval Medical School. Mem-
bers of the staff obtained about 1,200 mollusks in Arnhem Land and
some 1,500 in the region of the Persian Gulf and the Red Sea.
Botany.—As exchanges, the National Herbarium received 2,382
plants, comprising a collection made in Fiji by Dr. A. C. Smith, from
the Arnold Arboretum of Harvard University; 5,854 plants of Colom-
bia from the Facultad de Agronomfa, Universidad Nacional, Medellin;
and 2,157 Chinese plants from the National Szechwan University.
The Division of Rubber Plant Investigations, United States Depart-
ment of Agriculture, transferred 865 plants from eastern Colombia.
The Oficina de Estudios Especiales, Mexico City, presented 394
Mexican grasses. = nc ne ae ne Landscape with Boatman.
A painting by Murillo, “The Return of the Prodigal Son,” given
by the Avalon Foundation, was accepted by the Board of Trustees
on December 10, 1948. At the same time the Board accepted the
portrait of Daniel Boardman, by Ralph Earl, from Mrs. W. Murray
Crane; “Interior of a Church,” by Pieter Neeffs, from Senator Theo-
dore Francis Green of Rhode Island; and two paintings, ‘‘Repose,”’
by John Singer Sargent, and “Head of a Girl,” by James Abbott
McNeill Whistler, from Curt H. Reisinger. On December 22, 1948,
the Board of Trustees accepted from Dr. G. H. A. Clowes a painting,
“Allegory,” Venetian School about 1500, and from Vladimir Horo-
witz a painting, ““Head of a Young Girl,” by Renoir. The Board
of Trustees accepted from Miss Georgia O’Keeffe on March 8, 1949,
a gift of the following three paintings:
Artist Title
Marsden Hartley...) =) 322 al 2 es Se Landscape No. 5.
Ar Chun GAM ONG 55-285 ee Sane Seeue ess Moth Dance.
Georgia. Osieetiens. rere oe Ae ee To be selected later.
28 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
During the fiscal year, the portrait of Captain Patrick Miller by
Raeburn, previously on loan, was given to the Gallery by Mrs.
Dwight Davis.
SCULPTURE
On December 10, 1948, the Board of Trustees accepted from
Stanley Mortimer, Jr., a ‘“‘Portrait Bust of a Member of the Order of
San Iago” attributed to Leone Leoni, which had previously been on
loan to the Gallery. At the same time the Board accepted from
Miss Mildred Howells a portrait medallion of her father, William
Dean Howells, and herself, by Augustus Saint-Gaudens, to be held
for a National Portrait Gallery.
PRINTS AND DRAWINGS
A gift from Lessing J. Rosenwald of 309 additional prints and
drawings was accepted on December 10, 1948, to be added to the
Lessing J. Rosenwald Collection. At the same time, two volumes of
“The Georgics’”’ of Virgil with 119 illustrations by Andre Dunoyer de
Segonzac were accepted as a gift from the artist. This gift was
inspired by an earlier gift to the Gallery of a collection of Segonzac’s
prints and drawings made in memory of the late Frank Crowninshield.
The Board of Trustees, during the fiscal year, received 50 prints and
drawings from the collection of the late R. Horace Gallatin. On
March 8, 1949, the Board accepted from Miss Georgia O’ Keeffe three
water colors by John Marin entitled “Movement, Boat and Sea,
Deer Isle, Maine,’ “White Mountain Country, Summer,” and
“Storm over Taos, New Mexico.”’ The Board of Trustees accepted
from Mr. Rosenwald on May 3, 1949, 582 additional prints and draw-
ings. Received during the fiscal year from George Matthew Adams
were 20 etchings by Alphonse Legros.
PHOTOGRAPHS
The Board of Trustees on March 8, 1949, accepted from Miss
Georgia O’Keeffe a key set of photographs, consisting of about 1,500
prints, by Alfred Stieglitz.
EXCHANGE OF WORKS OF ART
During the fiscal year 1949 the Board accepted the offer of Chester
Dale to exchange the portrait of Ralph Waldo Emerson by Sully,
which was being held for the National Portrait Gallery, for the por-
trait of the Sicard David Children by Sully, which was then on loan
to the Gallery. The Board also accepted the offer of Lessing J.
Rosenwald to exchange the prints ‘Sacrifice to Priapus,”’ by Jacopo
de Barbari, ‘Conversion of St. Paul,’’ by Lucas van Leyden, and
SECRETARY’S REPORT 29
“Solomon Worshipping Idols,’ by the Master M. Z., for superior
impressions of like prints now included in the Rosenwald Collection
at the National Gallery of Art.
WORKS OF ART ON LOAN
During the fiscal year 1949 the following works of art were received
on loan by the National Gallery of Art:
From Artist
Chester Dale, New York, N. Y.:
1 BCR (0) oes ae de Ag ee ire oN ee Pee Domergue.
Mrs bhilipyliydics 8-2 ss 2 seo seee eee S Zuloaga.
SanvSepulvedaase= = sss te ee eee Zuloaga.
iarRubiaydelAbanicoss222-- 2-2 sse— ee osa2 Zuloaga.
Mrs. Brooks Goddard, Paris, France (via the
National Collection of Fine Arts):
Mrursical eins pina tio epee eae rere Romaine Brooks Goddard.
PEHewBaleory 55 22 5 UM A sere es Se eg Romaine Brooks Goddard.
Sketcne fete ee ssf oe ee ey, Bele eee Romaine Brooks Goddard.
SelfsPortral tees eee ee eee ee eee Romaine Brooks Goddard.
Alfred Stieglitz Collection:
(Miss Georgia O’ Keeffe, New York, N.
Y.)
Chimneys and Water Tower-------------- Demuth.
AGCowssiskulliwithpRediers= eee O’ Keeffe.
Ihinerand=Curviess=s28s8s cose ee eee eee O’ Keeffe.
Chauncey Stillman, New York, N. Y.:
AVHalberdier 24 tee ee ss ee ae asl Pontormo.
George Matthew Adams, New York, N. Y.:
Sretchings an 2 ole as sue ee ee awe Alphonse Legros.
C. S. Gulbenkian, Lisbon, Portugal:
28 pieces of Egyptian sculpture.
3 pieces of eighteenth-century French fur-
niture.
1 fourteenth-century Arabian bottle.
1 sixteenth-century Persian rug.
7 eighteenth-century French books.
The Italian Government:
AVmarblerscatueOi Davide sane sees ae ee Michelangelo.
Robert Woods Bliss, Washington, D. C.:
32 objects of Pre-Columbian art.
LOANED WORKS OF ART RETURNED
The following works of art on loan were returned during the fiscal
year 1949:
To Artist
Chester Dale, New York, N. Y.:
OnitherBeachee a ane ee eee eae ees Winslow Homer.
Mme. Charlotte Fuerstenberg, at New York,
INceyY -:
Sea abr Pstaquers. sess o2— oneness Cezanne.
30 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Anonymous loan:
Paradise, Valleyi43t 2-442 42-63-20 ee en John La Farge.
Robert Woods Bliss, Washington, D. C.:
16 objects of Pre-Columbian art.
Peabody Museum, Harvard University, Cam-
bridge, Mass.:
70 objects of Pre-Columbian art.
WORKS OF ART LOANED
During the fiscal year 1949, the Gallery loaned the following works
of art for exhibition purposes:
To Artist
Albright Art Gallery, Buffalo, N. Y.:
Joseph; Widener. -.-.---2 See +s. sseesS Augustus John.
Art Institute of Chicago, Chicago, IL:
Alexander: Hannltonassse oo eo Trumbull.
William) Dhormtone -enAs) 22-2 ee ee Stuart.
Columbus Gallery of Fine Arts, Columbus, Ohio:
iAbrahsimeltineoinivres 62 2S. sano ee Healy.
Corcoran Gallery of Art, Washington, D. C.:
‘The White Girl. 2-2 sa.=0* = ess eee Whistler.
Dallas Museum of Fine Arts, Dallas, Tex.:
George Washington (Vaughan-Sinclair) ----- Stuart.
Dayton Art Institute, Dayton, Ohio:
ackawanna Valleys sof ses- 22. 2 eee Inness.
Fort Worth Art Association, Forth Worth, Tex.:
Breezing Wp. eee aA a ee Winslow Homer.
Metropolitan Museum of Art, New York, N. Y.:
Captain Charles Stewart. -.--.-.--------- Sully.
Pack Memorial Library, Asheville, N. C.:
Tomas *Vawsone 26 sa 2se ese = = ee Mather Brown.
Henry, Laurense... 2 = 23st atest est Copley.
Andre wa acCkSON= =a = esac arene eee eee Earl.
WiilltaminatVloores ae ene ee = = ae Feke.
General) William: Moultries === 22-- — = === Charles Willson Peale.
JohnmiC: Calhoune ses. 226-2. ae Rembrandt Peale.
JOUnNeBaptista Ashe nas sas ee ee eee Stuart.
Matilda Caroline; @ruger. = .=..-----=-—-=- Stuart.
Hrancis, Hopkinson... 25 S22_2- = 22 ee Sully.
Annebiddle Hopkinson] —5-—. 222-4... 2 355-2 Sully.
JosiasrAllston ease oe ae eee eee Theus.
Walliam Rogerst=.4 ies ease es ee eee Trumbull.
Portraits, Inc., New York, N. Y.:
Mrs#Chester Dale: si) 210 0 sees fat or Bellows.
Mr Chester Dale. 25. == 252-2. GASES Bellows.
Scott and Fowles, New York, N. Y.:
Joseph Widener... 2.522.225. 5.css5558 Augustus John.
SECRETARY'S REPORT oil
EXHIBITIONS
During the fiscal year 1949 the following exhibitions were held at
the National Gallery of Art:
American Paintings from the Collection of the National Gallery of Art. Exhi-
bition of American paintings, featuring a group of portraits from Pocahontas to
General Eisenhower. Continued from previous fiscal year, through July 11, 1948.
American Folk Art. Exhibition consisting of 104 water-color renderings from
the Index of American Design. July 18 to September 7, 1948.
American Graphic Art from the Eighteenth Century to the Present Day.
Selection from the collections of the Library of Congress, the Smithsonian Institu-
tion, and the National Gallery of Art. September 19 to November 14, 1948.
Paris the Favorable Climate. Exhibition of prints and drawings by Bonnard,
Vuillard, Maurice Denis, Andre Dunoyer de Segonzac, and Matisse, arranged in
memory of Frank Crowninshield. November 21, 1948, to January 11, 1949.
Michelangelo’s ‘‘David.’’ Lent to the National Gallery of Art by the Italian
Government. January 24 to June 28, 1949.
Gulbenkian Collection of Egyptian Sculpture. Lent for an indefinite period to
the National Gallery of Art by C. 8. Gulbenkian. Opened January 30, 1949.
Studies of Medieval Cathedrals. Exhibition of photographic studies lent to
the National Gallery of Art by Clarence Ward, head of the Department of Fine
Arts, Oberlin College. January 30 to February 13, 1949.
Gulbenkian Collection of Eighteenth Century French Objects. Additions to
earlier loan by C. S. Gulbenkian, on exhibition at the National Gallery of Art
for an indefinite period. Opened February 20, 1949.
American Paintings from the Collection of the National Gallery of Art. Feb-
ruary 20 to April 10, 1949.
Early Italian Engraving. Exhibition of early Halian engravings, lent to the
National Gallery of Art by various museums and anonymous lenders. April 17
to June 19, 1949.
R. Horace Gallatin Collection. Exhibition of prints bequeathed to the National
Gallery of Art by Mr. Gallatin. Opened June 26, 1949.
The following exhibitions were displayed in the cafeteria corridor
of the National Gallery of Art during the fiscal year 1949:
Whistler Prints. Rosenwald Collection; one gift of Myron A. Hofer. Con-
tinued from previous fiscal year through July 18, 1948.
Audubon Prints. Mrs. Walter B. James Collection. July 20 to December 12,
1948.
Index of American Design. Water-color renderings of early American toys.
December 13, 1948, to February 15, 1949.
Index of American Design. Water-color renderings of early American furniture
and textiles. February 16 to March 28, 1949.
Legros Prints. George Matthew Adams Collection. March 29 to May 15,
1949.
Seymour Haden Prints. Rosenwald Collection and gift of Miss Elisabeth
Achelis. May 16 to June 12, 1949.
Ostade Prints. Rosenwald Collection and gift of Mrs. Addie Burr Clark.
Opened June 13, 1949.
a2 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
TRAVELING EXHIBITIONS
Rosenwald Collection.—Special exhibitions of prints from the Rosen-
wald Collection were circulated to the following places during the
fiscal year 1949:
Kenneth Taylor Galleries, Nantucket, Mass.:
26 French prints.
July 26 to August 23, 1948.
Watkins Gallery, American University, Washington, D. C.:
26 French prints.
October 13 to 30, 1948.
Los Angeles County Museum, Los Angeles, Calif.:
20 Blake prints.
October 1948.
Wyncote Woman’s Club, Wyncote, Pa.:
11 prints.
October 17 to 23, 1948.
Rutgers College, New Brunswick, N. J.:
9 Italian prints.
October 1948.
Fogg Museum of Art, Harvard University, Cambridge, Mass.:
1 Rembrandt drawing.
November 1948.
Museum of Modern Art, New York, N. Y.:
1 Munch print.
November 1948 to January 1949.
Walters Art Gallery, Baltimore, Md.:
6 Gavarni drawings.
January 22 to March 6, 1949.
Walters Art Gallery, Baltimore, Md.:
5 miniatures.
January 27 to March 138, 1949.
City Art Museum, St. Louis, Mo.:
17 prints.
March 1949.
Institute of Contemporary Arts, Washington, D. C.:
11 Klee prints.
March 21 to April 22, 1949.
Philadelphia Museum of Art, Philadelphia, Pa.:
3 Lehmbruck prints.
May 1949.
Art Gallery of Toronto, Toronto, Canada:
67 prints.
May 1949.
SECRETARY'S REPORT
33
Index of American Design.—During the fiscal year 1949 exhibitions
from this collection were shown at the following places:
Library of Congress, Washington, D. C.
Western Reserve Historical Society,
Cleveland, Ohio.
Shaker Village Work Camp, Pittsfield,
Mass.
New York State Historical Association,
Cooperstown, N. Y.
Damariscotta Information Bureau,
Damariscotta, Maine.
University of ‘Tennessee,
Tenn.
Wustum Museum of Fine Arts, Racine,
Wis.
City Art Museum, St. Louis, Mo.
William Rockhill Nelson Gallery, Kan-
sas City, Mo.
Munson- Williams-Proctor Institute,
Utica, N. Y.
Toledo Museum of Art, Toledo, Ohio.
Mint Museum, Charlotte, N. C.
Museum of Fine Arts, Montgomery, Ala.
Knoxville,
Schenectady Museum, Schenectady,
Na ¥2
University of Oklahoma, Norman,
Okla.
University of Michigan, Ann Arbor,
Mich.
North Carolina College, Durham, N. C.
Art Institute, Zanesville, Ohio.
Atlanta University, Atlanta, Ga.
Currier Gallery of Art, Manchester,
N. H.
Stephens College, Columbia, Mo.
Brown University, Providence, R. I.
Fort Valley State College, Fort Valley,
Ga.
Washington College, Chestertown, Md.
Everhart Museum, Scranton, Pa.
Art Gallery, Grand Rapids, Mich.
Florida Agricultural and Mechanical
College, Tallahassee, Fla.
Farnsworth Museum, Rockland, Maine.
Tuskegee Institute, Tuskegee, Ala.
Young Playways, Inc., Washington,
Dr CE:
Smith College, Northampton, Mass.
Prairie View University, Prairie View,
Tex.
University of North Dakota, Grand
Forks, N. Dak.
American University, Washington, D. C.
Rockford Art Association, Rockford, II.
Sweet Briar College, Sweet Briar, Va.
Arkansas Agricultural, Mechanical and
Normal College, Pine Bluff, Ark.
Alfred University, Alfred, N. Y.
Fisk University, Nashville, Tenn.
St. Paul Public Library, St. Paul, Minn.
Spelman College, Atlanta, Ga.
Arnot Art Gallery, Elmira, N. Y.
Kenneth Taylor Galleries, Nantucket,
Mass.
CURATORIAL ACTIVITIES
The Curatorial Department accessioned 1,118 new gifts to the
Gallery during the fiscal year. Advice was given in the case of 233
works of art brought to the Gallery for opinion, and 58 visits were
made by members of the staff in connection with proffered works of art.
Almost 1,000 research problems requiring reports were investigated
in response to inquiries received by the Gallery. During the year,
16 individual lectures were given by members of the curatorial staff,
both at the Gallery and elsewhere. In addition Miss Elizabeth
Mongan gave a seminar at Alverthorpe, Jenkintown, Pa., for Swarth-
34 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
more College honor students; Charles Seymour, Jr., gave a course
at Johns Hopkins University on Renaissance Art; and Charles M.
Richards gave a survey course on art history under the auspices of
the Department of Agriculture. Miss Mongan also made the
arrangements for Arthur M. Hind’s American lecture tour, in connec-
tion with the publication of Part II of his ‘‘Harly Italian Engraving,”
under Gallery auspices. Mr. Seymour served on three and Miss
Mongan on two art juries.
Special installations were prepared for: the Michelangelo “David”
lent to the National Gallery of Art through the courtesy of the
Italian Government; 28 pieces of Egyptian sculpture lent to the
Gallery by C. S. Gulbenkian placed on exhibition in January 1949;
and eighteenth-century furniture and books also lent by Mr. Gulben-
kian. The cataloging and filing of photographs in the George Martin
Richter Archive continued to make progress, with the gradual
enlargement of the collection.
Further activities of the department are indicated under the
heading of “‘ Publications.”
RESTORATION AND REPAIR OF WORKS OF ART
Necessary restoration and repair of works of art in the Gallery’s
collections were made by Stephen S. Pichetto, Consultant Restorer
to the Gallery, until his death in January 1949. No successor to
Mr. Pichetto has as yet been appointed, but necessary minor repairs
on the works of art have been continued under the care of Mr.
Pichetto’s residual staff. All work was completed in the Restorer’s
studio in the Gallery, with the exception of the restoration of two
paintings, work on which is being completed in the New York studio
of S. S. Pichetto, Inc.
PUBLICATIONS
During the year Mr. Cairns published two books, ‘‘The Limits
of Art,’’? Pantheon Books, Inc., and ‘‘Legal Philosophy from Plato
to Hegel,” Johns Hopkins Press. He also edited a volume entitled
“Lectures in Criticism,’”’ Pantheon Books, Inc., and contributed an
introduction to “Epicurus, My Master,’’ by Max Radin, University
of North Carolina Press. He also contributed articles and reviews
to the Columbia Law Review, Human Events, Saturday Review
of Literature, New York Herald Tribune, Baltimore Evening Sun,
Law and Contemporary Problems, The Scientific Monthly, and to
the volume El Actual Pensamiento Juridico de los Estados Unidos,
Buenos Aires.
A series of 12 articles on masterpieces in the Gallery, prefaced
by one entitled ‘‘New Friends for Old Masters,” is being published
by John Walker in the Ladies Home Journal. An article by Mr.
SECRETARY’S REPORT 35
Walker, “The Art of Duplicating Great Art,” appeared in Vogue
on August 15, 1948, and another, ‘‘American Masters in the National
Gallery,” in the National Geographic Magazine in September 1948.
Mr. Walker also contributed two book reviews, the first reviewing
Bernard Berenson’s ‘‘ Aesthetics and History in the Visual Arts”
to the October 1948 Gazette des Beaux-Arts, and the second, entitled
‘The Philosophy of a Connoisseur,’ a review of Mr. Berenson’s
‘Sketch for a Self-Portrait,’ to the New York Times for April 24,
1949. Charles Seymour, Jr., published two articles, ““Note on the
Relationship between an Illustration by Travies de Villers and
Daumier’s ‘Le Fardeau’,” in the Journal of the Walters Gallery for
1948, and in the Summer Bulletin of the Columbus Gallery of Fine
Arts the text of the address given by him for the inauguration of a group
of sculpture by Georg Ehrlich in the Columbus Gallery of Fine Arts.
Printing of ‘‘Masterpieces of Sculpture from the National Gallery
of Art,’ a volume prepared by Mr. Seymour, was begun during the
summer of 1949. Mrs. Fern R. Shapley has written two book reviews,
a review of Bernard Berenson’s ‘‘ Aesthetics and History’ which is
to be published in the next number of the College Art Journal, and
one on Evelyn Sandberg-Vavala’s ‘‘ Uffizi Studies” published in the
January 1949 Gazette des Beaux-Arts. Miss Elizabeth Mongan
contributed six articles for the volume honoring Paul Sachs; an article
for the Color Print Society on ‘Rockport,’ a colored lithograph
by Stella Drabkin; descriptions of 27 illuminated miniatures to Pro-
fessor Faye for the second edition of Seymour de Ricci’s “Census
of Manuscripts in America.”’ An article on Rowlandson by David
Keppel was published in the winter, 1949, number of The Art Quarterly.
An article by James W. Lane entitled ‘Religious Art Exhibit’
appeared in the Interracial Review, and one on ‘‘Contemporary
Religious Sculpture Exhibition” in the Catholic University Bulletin;
he contributed two book reviews on ‘‘Van Eyck’s the Holy Lamb,”
by Leo Van Puyvelde, and ‘‘Robert Louis Stevenson,’”’ by David
Daiches, to the Catholic World, and one on “American Landscape
Painting,” by Wolfgang Born, to the Magazine of Art. Charles M.
Richards wrote a report on a code for intermuseum loans for the
American Association of Museums.
An illustrated catalog of the Gulbenkian Egyptian sculpture was
issued for the opening of the exhibition, and Mr. Seymour prepared
a pamphlet on the Michelangelo ‘‘ David,” which was placed on sale
during its exhibition. The book of illustrations of the Mellon Collec-
tion went to press in the late spring of 1949; work on the new National
Gallery of Art catalog is at an advanced stage.
The Publications Fund during the past fiscal year has continued to
add new subjects to the supply of inexpensive color reproductions
36 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
offered to the public, including 11’’ x 14’’ color prints and color
post cards. Five large collotype reproductions supplemented the
already long list of subjects available. A silk-screen print of an
anonymous fifteenth-century colored woodcut from the Rosenwald
Collection was also published.
The Gallery is continuing to meet the demand for illustrated
catalogs of its various collections. The Mellon catalog is in process
of publication, a third printing of the Kress catalog ordered, and a
fifth edition of the Chester Dale catalog was published during the year.
Two new publications were issued this year: an ‘Arts and Crafts
Bibliography,” by Erwin O. Christensen, and a catalog of the ‘‘Egyp-
tian Sculpture from the Gulbenkian Collection.” A group of engraved
Christmas cards was added to the usual series of color and Rosenwald
subjects.
Final negotiations have been made for the printing in gravure of
the book, ‘‘Masterpieces of Sculpture from the National Gallery of
Art,” and it will be available by October 1949; the publisher now
has the final manuscript for ‘‘ Made in America,”’ by Mr. Christensen;
the Gallery received a stock of ‘Popular Art in the United States,”’
also by Mr. Christensen, which will go on sale on July 4, 1949; and
“Pictures from America,”’ by John Walker, will shortly be published.
EDUCATIONAL PROGRAM
During the year approximately 15,000 persons attended the Gen-
eral, Congressional, and Special Topic Tours, while over 20,000
attended the Picture of the Week. More than 18,000 came to hear
the lectures and other programs in the auditorium. At least two-
thirds of this lecture audience were regular attendants at these
Sunday afternoon lectures. Many of them brought out-of-town
visitors, and stated that this lecture series was becoming one of the
Capital’s chief Sunday attractions. The motion picture, ‘The
National Gallery of Art,” continues to be popular with clubs, educa-
tional organizations, and similar groups. During the past 12 months,
19 persons borrowed this film.
The publication of the monthly Calendar of Events, announcing
Gallery activities, including notices of exhibitions, lectures, Gallery
talks, tours, and concerts was continued during the year by the
Educational Department. About 3,900 of the Calendar of Events
are mailed each month.
LIBRARY
A total of 283 books, 221 pamphlets, and 31 periodicals were given
to the Gallery; 494 books, 18 pamphlets, and 282 periodicals were
purchased, and 40 subscriptions to periodicals were purchased.
Exchanges with other institutions included 47 books, 114 pamphlets,
SECRETARY'S REPORT 37
13 periodicals, and 420 bulletins. Of the 1,762 books borrowed and
returned during the year, the Library of Congress lent 1,676 books to
the Gallery on the usual interlibrary loan basis, and the remaining
86 books were borrowed from 25 public and university libraries.
INDEX OF AMERICAN DESIGN
During the year the Index of American Design continued to expand
as the result of gifts and exchanges. Three hundred and thirty-six
persons studied Index material at the Gallery; of this number, 301
were new users and 25 revisited the collection for study purposes.
The use of photographs of Index drawings was increased by about
40 percent, with 1,796 photographs being sent out on loan, exchange,
or purchase. Fifty exhibitions of original water-color renderings
were circulated in 25 States.
PRESIDENT TRUMAN’S INAUGURAL RECEPTION
On January 20, 1949, the President’s Inaugural Reception was
held in the National Gallery of Art. The Seventh Street ground
floor and main floor lobbies were especially furnished and decorated
for the occasion; the rotunda and the two garden courts were appro-
priately decorated with flowers; under arrangements made by the
White House staff, a platform was built in the West Sculpture Hall
where the President addressed the guests who could not be received
personally in the West Garden Court. Three sections of the Marine
Band Orchestra played during the reception. The total number of
guests was approximately 8,000.
CUSTODY OF GERMAN PAINTINGS
On April 6, 1949, the Gallery accepted custody of the 97 paintings
from Berlin museums which had been on an exhibition tour of the
United States, part of the group of 202 German paintings stored in
the Gallery building by the Department of the Army from December
1945 to March 1948. After the last exhibition of this collection of
paintings in Toledo, Ohio, the collection was brought to Washington
and stored in the Gallery for about 2 weeks pending final shipping
arrangements. On April 20, 1949, the collection was delivered to
the Army for return to the American Zone in Germany.
The exhibition of the Berlin paintings in 13 museums throughout
the United States resulted in the collection of $303,605.35 through ad-
mission fees and voluntary contributions for the relief of German
children in the American Zone in Germany. ‘These funds were de-
posited with the Gallery and were later disbursed in accordance with
instructions received from the Department of the Army. During the
tour 1,307,001 persons viewed the paintings, in addition to 964,970
38 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
who saw them during the time the paintings were on exhibition at the
National Gallery of Art in Washington.
CUSTODY OF GERMAN SILVER
On January 7, 1949, the Gallery returned to the Department of the
Army for transport to Germany the 44 sealed cases containing silver-
ware and glassware and belonging to the Hohenzollern family. The
cases had been stored in the Gallery since April 11, 1947.
CUSTODY OF WHITE HOUSE FURNITURE
On November 24, 1948, the Gallery accepted custody of certain
items of paintings, sculpture, and furniture belonging to the White
House for storage in the building until the repairs to the White House
are completed.
Shipments of these items started on December 3, 1948, and con-
tinued for several days thereafter. At the present time there are 76
works of art—paintings and sculpture—stored in the Gallery’s storage
rooms and 25 vanloads of furniture stored in the packing space on the
main floor.
The necessary arrangements for fire prevention, inspection, and
fumigation have been established and are being carried out.
NEW CONSTRUCTION
During the past fiscal year, the Committee on the Building approved
the construction in the southwest moat of a small workroom for the
use of the gardening staff in maintaining and growing certain plants
for the garden courts and landscaping. Later, when funds become
available, it is planned to construct two small greenhouses adjacent
to this workroom.
The growth of the Gallery’s collections of works of art has been so
rapid that all available exhibition space is now being utilized. As a
matter of fact there are already several paintings which cannot be ex-
hibited because there is no space in the present galleries. For this
reason the Committee on the Building recommended that, to take care
of the most urgent needs, the unfinished spaces 61-66 and 68-70, on
the main floor, be completed as soon as funds are available. These
galleries will be used for new acquisitions of paintings in the American
and British schools and will also make possible some rearrangement in
galleries already finished so as to make available additional space
therein.
The Committee on the Building also recommended that the so-called
copyists’ room be finished to furnish office space for the Educational
Department, which is now operating in rather cramped quarters.
Funds have been generously made available from private sources to
complete this work, and contracts have been entered into with Eggers
SECRETARY'S REPORT 39
and Higgins, Architects, and Vermilya-Brown Company, General
Contractors, for the completion of 12 galleries in these unfinished
areas. The floor plan has been approved, and bids are now being taken
from subcontractors. It is anticipated that actual construction will
begin in August 1949 and that the work will be completed by May 1950.
CARE AND MAINTENANCE OF THE BUILDING
The usual routine work in connection with the care and maintenance
of the building andits mechanical equipment was carried on throughout
the year.
The three older refrigeration compressors were completely dis-
mantled and overhauled, including the purge compressors. Three
chilled-water pumps, including the electric motors, were completely
overhauled and realigned by the mechanical staff. Twelve supply
fans were cleaned and repainted to protect them against corrosion,
The structural steel base for the large 400-horsepower motor driving
No. 2 Worthington refrigeration machine was strengthened in order
that this large motor would remain in alignment. ‘To correct serious
leaks in two of these machines, the technical staff successfully made
and installed the necessary parts.
The cornice metal lining at the top of the exterior wall of the
building developed leaks, and approximately 50 percent of the joints
in the metal lining were cleaned and soldered.
In connection with the Inaugural Reception, the technical staff
installed floodlights on three sides of the building, assisted the person-
nel of the U.S. Army Signal Corps in the installation of a loud-speaker
system on the main floor, and installed extra electric lines and water
lines for the use of the caterer. The maintenance staff erected exten-
sive checking facilities for the proper care of wraps.
Twelve new display cases were constructed by the staff for the
Gulbenkian Exhibition.
Care and improvement of the Gallery grounds and other miscella-
neous work progressed satisfactorily. Potted plants, totaling 2,366,
which were used for decoration in the two garden courts, were grown in
the southwest moat. In addition, over 350 large pots of chrysanthe-
mums were also grown in this moat area, and these plants provided
the decoration for the two garden courts during the months of October
and November.
COMMITTEE OF EXPERT EXAMINERS
During the year the United States Civil Service Commission’s
Committee of Expert Examiners, composed of staff members of the
Gallery, aided in the drafting of standards for Civil Service positions in
which a knowledge of the history of art is a basic requirement. The
866591—50-——4
40 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Committee also performed preliminary work in the preparation
of the examination announcement for art positions which was dis-
tributed by the Civil Service Commission with a closing date of
April 19, 1949. From this examination registers of eligibles will be
established for appointment to art positions in the Gallery and
elsewhere in the Government. This will give the present incumbents,
most of whom are serving indefinite war-time appointments, an
opportunity to attain permanent status, and will also make available
a greater number of qualified candidates.
OTHER ACTIVITIES
Forty-six Sunday evening concerts were given during the fiscal
year, all concerts being held in the East Garden Court. A Mozart
Festival of six concerts was given in the autumn with the highest
attendance rate for the season. The five Sunday evenings in May
were devoted to the Gallery’s annual American Music Festival. An
estimated 50,000 persons attended these concerts.
During the year the photographic laboratory of the Gallery made
17,709 prints, 1,342 black-and-white slides, 1,005 color slides, 3,873
negatives, in addition to infrared photographs, ultraviolet photographs,
X-rays, and color separation negatives.
A total of 3,500 copies of press releases, 128 special permits to copy
paintings in the National Gallery of Art, and 117 special permits to
photograph in the Gallery were issued during the year.
OTHER GIFTS
Gifts of books on art and related material were made to the Gallery
library during the year by Paul Mellon and others. Gifts of money
during the fiscal year 1949 were made by the Avalon Foundation and
The A. W. Mellon Educational and Charitable Trust, and a cash
bequest was received from the Estate of the late William Nelson
Cromwell.
AUDIT OF PRIVATE FUNDS OF THE GALLERY
An audit of the private funds of the Gallery has been made for the
fiscal year ended June 30, 1949, by Price Waterhouse & Co., public
accountants, and the certificate of that company on its examination
of the accounting records maintained for such funds will be forwarded
to the Gallery.
Respectfully submitted.
Huntineton Cairns, Secretary.
THe SECRETARY,
Smithsonian Institution.
APPENDIX 3
REPORT ON THE NATIONAL COLLECTION OF FINE ARTS
Sir: I have the honor to submit the following report on the activi-
ties of the National Collection of Fine Arts for the fiscal year ended
June 30, 1949.
THE SMITHSONIAN ART COMMISSION
The twenty-sixth annual meeting of the Smithsonian Art Commis-
sion was held in the Regents’ Room of the Smithsonian Building, on
Tuesday, December 7, 1948.
The members present were: Paul Manship, chairman; Alexander
Wetmore, secretary (member, ex officio); George Hewitt Myers;
George H. Edgell; Lloyd Goodrich; John Taylor Arms; Archibald G.
Wenley, Gifford Beal, and Robert Woods Bliss. Thomas M. Beggs,
Director of the National Collection of Fine Arts, and John E. Graf,
Assistant Secretary of the Smithsonian Institution, were also present.
The Commission recommended the reelection of Archibald G. Wen-
ley, David E. Finley, Eugene E. Speicher, and Paul Manship for the
usual 4-year period.
The following officers were reelected for the ensuing year: Paul Man-
ship, chairman; Robert Woods Bliss, vice chairman; and Dr. Alexander
Wetmore, secretary.
The following were reelected members of the executive committee
for the ensuing year: David E. Finley, chairman, Robert Woods Bliss,
and Gilmore D. Clarke. Paul Manship, as chairman of the Commis-
sion, and Dr. Alexander Wetmore, as secretary of the Commission, are
ex officio members of the executive committee.
The Secretary summarized the status of exhibition and storage of
the art objects of the National Collection of Fine Arts which at present
are housed in space intended for the natural history collections in the
Natural History Building. 5----2 2: Qi Caulormige -2cecceson sae 5.8 8
The cars that made up the remaining 10.30 percent came from
every one of the remaining States, as well as from Alaska, Bahamas,
Canada, Canal Zone, Chile, Cuba, Guam, Hawaii, Honduras, Italy,
Japan, Mexico, Netherlands, Newfoundland, Poland, Puerto Rico,
Sweden, Trieste, Trinidad, and Virgin Islands.
It is well known that District of Columbia, Maryland, and Virginia
cars bring to the Zoo many people from other parts of the United
States and of the world, but no figures are available on which to base
percentages.
FINANCES
The regular appropriation provided in the District of Columbia
appropriation act was $492,600, and there was a supplemental appro-
priation in the second deficiency bill of $36,248 to provide for the
increased salaries of $330 per annum authorized by Congress. Of
the total of $528,848 which was available, about $11,474 will remain
unexpended, subject to minor changes in final bills. This saving
was mainly from salaries because of the impossibility of filling posi-
tions promptly.
The stone restaurant building, which was constructed in the park
in 1940 under an allotment of $90,000, is under a 3-year lease obtained
by competitive bidding at $10,212 per annum. This money is de-
posited in the general fund of the United States Treasury. The
concessionaire serves meals and light refreshments, and sells novelties.
NEEDS OF THE ZOO
The chief need of the Zoo is for the replacement of antiquated
structures that have long since ceased to be suitable for the purpose.
The more urgently needed buildings are: (1) A new administration
building to replace the 144-year-old historic landmark now in use for
an office building for the Zoo, but which is neither suitably located nor
well adapted for the purpose. This building is in an excellent loca-
tion for a public recreational structure, and could probably be rehabili-
tated and used for recreational purposes, perhaps as a children’s
108 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
museum, and thus maintained as a historic building. The new
office building should be better located both from the standpoint of
accessibility to the public and convenience for the administration of
the Zoo. (2) A new building to house antelopes and other medium-
size hoofed animals that require a heated building.
STATUS OF THE COLLECTION
Class Species individ Class Species jadivid:
Mammialse2-= 2) <2 os2s5 22252 235 786; || M@rustaceanstass saan 1 2
Birds 32. 2 ee ea ee a 346 1,911 |} Arachnids____ 2 3
Reptiles sae eee ae ees 121 509 || Insects_._--.. 1 100
Amphibians = sees a ree 22 1644S lollusks 22 seen eee eee 3 23
Bishee | Sede): A. aeRO 24 226
Total. 53. #2225 335_-5-23 755 3, 724
SUMMARY
Animals Onenan a dlily ly, M948. a. ole SR oe ee ee 2, 797
‘Accessionsduringythe yearn. 22555220 seo] es ee eee ee Tao
Total number of animals in collection during the year___._______-_ 4, 548
Removals for various reasons such as death, exchanges, return of animals
onydeposit: vetese StL as. See se eee pee to ice rrn crate ih Tage Oe 824
Im-collection on June.o0; 9408 2. 0. ae es eee ee eee 3, 724
Respectfully submitted.
W. M. Mann, Director.
Dr. A. WrTmore,
Secretary, Smithsonian Institution.
APPENDIX 8
REPORT ON THE ASTROPHYSICAL OBSERVATORY
Srr: I have the honor to submit the following report on the opera-
tions of the Astrophysical Observatory for the fiscal year ended
June 30, 1949:
The Observatory includes two research divisions: (1) the Division
of Astrophysical Research, concerned chiefly with solar radiation
problems, and (2) the Division of Radiation and Organisms, con-
cerned with the biological effects of radiation.
During the year a new room adjoining the Director’s office was
built for the administrative assistant and for the files of the Obeerva-
tory. The resulting consolidation of needed information nsar at
hand has materially improved the efficiency of operation.
Considerable progress can be reported concerning the new revised
editions of the Smithsonian Meteorological Tables and the Smith-
sonian Physical Tables, mentioned in last year’s report. R. J. List,
editor of the Meteorological Tables revision, had practically completed
his manuscript at the end of the fiscal year. The difficult task of
revising the Physical Tables, the last revision of which had been issued
in 1934, was begun in September 1948 under the direction of Dr.
W.E. Forsythe. An office in Cleveland, Ohio, and an assistant were
furnished to Dr. Forsythe. At the close of the fiscal year he reports
that approximately one-half of the tables for the new edition have
been completed.
(1) DIVISION OF ASTROPHYSICAL RESEARCH
Previous to 1946 the Observatory had for many years maintained
three high-altitude field stations for solar-constant observations. In
1946 the Tyrone station, which for 7 years had been operated on
Burro Mountain (altitude 8,000 ft.) in southwestern New Mexico,
was abandoned because skies there had progressively deteriorated,
mainly the result of increasing mining and smelting operations in that
general region. As a temporary measure, to aid in certain studies
referred to below under contract with the Quartermaster Corps, the
Tyrone station equipment was transferred to and installed at Miami,
Fla. Since then much effort has been spent to find the most suitable
location for a third high-altitude field station to replace the aban-
doned Tyrone site. In last year’s report we mentioned that after
109
110 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
careful investigation three promising sites had been chosen for further
study and that in May 1948 a recording Eppley pyrheliometer was
installed at each of these sites, namely, (1) Torreén, Coahuila, Méx-
ico; (2) Mountain Pass, near Clark Mountain, California; and (3)
Pohakuola, Hawaii. The three pyrheliometers were operated for a
period of 1 year, ending June 1949. The resulting records indicate
the uniformity and the quality of the sky for each day during the
period. It is clear from the records that the best skies prevailed
at the Clark Mountain location. The second-best site was Poha-
kuola. This spot, 6,500 feet above sea level on the Island of Hawaii,
yielded some records of unusually clear and uniform skies, but such
skies were not the rule. At Clark Mountain, during the period
June 8 to March 31, there were 171 days with skies sufficiently good
for satisfactory observations, while at Table Mountain, Calif., during
the same period actual observations were made on 135 days. From
studies of these records and other sources, it appears that the Clark
Mountain region is in general considerably drier and more free of
haze and clouds than any other high-altitude location at present
known in the northern hemisphere.
In view of this, estimates were obtained of the cost of establishing
a field station at an altitude of 6,500 feet on the south slope of Clark
Mountain. Owing to the prevailing high prices for building materials
and labor, the estimates proved to be in excess of available funds.
It is hoped that sufficient funds may become available, but pending
this the Observatory plans immediately to enlarge its facilities at
Table Mountain sufficiently so that it will be possible to proceed
without delay with the special experimental problems mentioned in
last year’s report.
Work at Washington.—W. H. Hoover, Chief of the Division, in
addition to supervision of the work in progress, prepared data and
tables which will help to simplify the computations in the field. In
the past, to obtain the air mass (or length of path of the solar beam
in the atmosphere) it has been necessary to plot carefully a series
of theodolite readings against time, to read off desired altitude values,
and finally to enter an air-mass-altitude table. With the aid of
Hoover’s data, the observer, by reading the theodolite at specified
intervals, may enter the tables directly to determine the air mass.
This eliminates the tedious curve-plotting process.
A new instrument, designed by Dr. John W. Evans, of the High
Altitude Observatory of Harvard University, and described by him
in the Journal of the Optical Society of America, December 1948, was
kindly lent to the Astrophysical Observatory by Dr. Menzel of Har-
vard University to test and to determine its adaptability to Smith-
sonian work. The instrument is a photometer especially designed
SECRETARY'S REPORT 111
for determining the brightness of the sky immediately surrounding
the sun. Excellent results have been obtained with it at the Harvard
Station at Climax, Colo. It is of considerable interest to compare its
readings with simultaneous readings of the Smithsonian pyranometer
which also measures the brightness of the sky in a zone around the
sun. In preparation for comparison tests a rigid mounting has been
prepared for the instrument with slow-motion adjustment in altitude
and azimuth.
During the fiscal year, two silver-disk pyrheliometers, Nos. 80 and
81, were built, calibrated and sold at cost, one to the Hebrew Institute
of Technology, Haifa, Palestine, and the other to the Dublin Institute
for Advanced Learning. Inadditiontwomodified Angstrom pyrheliom-
eters and one special instrument for the spectroscopic determination
of atmospheric water vapor have been prepared for the Belgium
Meteorological Institute. These were nearly completed at the end
of the year.
Dr. C. G. Abbot, research associate of the Observatory, continued
his studies of the dependence of weather upon solar changes. This
work has been published in Smithsonian Miscellaneous Collections,
vol. 111, Nos. 5,6, and 7. Dr. Arctowski’s studies of solar and terres-
trial atmospheres were retarded by illness, but his work was resumed
before the close of the year.
Work win the field——Daily observations of the solar constant were
in progress throughout the year, as far as skies permitted, both at
Montezuma, Chile, and at Table Mountain, Calif. The skies during
the year were apparently normal at Table Mountain, but at Monte-
zuma, the observers noted an unusual number of days with light cirrus
clouds.
Early in the year Mr. Hoover carried the Observatory’s substandard
silver-disk pyrheliometer S. I. No. 5 to Miami for direct comparisons
with the pyrheliometers at that station. In February 1949, in the
course of changing the personnel at Montezuma, Chile, substandard
S. I. No. 5 was carried to Montezuma by the new Montezuma ob-
server, and brought back in April by the retiring observer, after inter-
comparisons had been made in Chile. The previous year S. I. No. 5
had been carried to Table Mountain by the director for similar inter-
comparisons. Thus there are now very recent direct comparisons
between all field pyrheliometers and substandard S. I. No. 5, which
in turn was carefully compared in 1947 with the absolute water-flow
standard. These many intercomparisons show no material changes
in constants. They satisfactorily confirm the adopted scale of pyr-
heliometry. A revision of Dr. Abbot’s paper of 1922 on “The Silver-
disk Pyrheliometer”’ is in preparation, summarizing the constants of
all silver-disk pyrheliometers, and describing certain changes which
112 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
have been adopted in recent years, both in the instrument itself and
in the method of use.
In June 1945 special radiation measurements were started at Camp
Lee, Va., under contract with the Office of the Quartermaster General,
in connection with their long-range study of the causes for the deteri-
oration of tents and tent materials. This contract has been renewed
each year since then, and the work has now extended to include similar
radiation measurements at Miami, Fla., a wet, sea-level station, and
at Montezuma, Chile, a dry, high-altitude station. The Observatory
completed the Camp Lee measurements January 1, 1948, and since
then they have been continued by the Quartermaster Board at Camp
Lee, with the Observatory acting in an advisory capacity, and giving
assistance when difficulties arise. The measurements at Miami,
begun in December 1947, have continued throughout the present
fiscal year. Similar radiation measurements and textile exposures
were begun at Montezuma, Chile, in December 1948, and will continue
approximately 2 years. Five reports to the Office of the’ Quarter-
master General were made during the year summarizing,the data
obtained at Miami and at Montezuma.
In January 1949, the Director visited the Miami field station, to
inspect the work in progress. While there he obtained special bolo-
graphs showing the absorption effects of known quantities of water
vapor in the atmosphere. Measurements of these bolographs con-
firmed the correctness of precipitable water curves which Mr. Fowle
had determined in earlier work at Washington and which have since
been used many times in our solar-constant program. This work is
discussed in Smithsonian Miscellaneous Collections, vol. 111, No. 12,
soon to be issued.
(2) DIVISION OF RADIATION AND ORGANISMS
(Report prepared by Dr. R. B. Withrow)
The work of the Division for the past year has been concerned
chiefly with reorganizing and reequipping the laboratories. New
office space has been established in conjunction with the basement
laboratories. These offices have been furnished with desks and cases
and will accommodate a maximum of nine individuals.
Most of the laboratories have been repainted and are being re-
equipped with modern lighting facilities. The laboratory furniture
has been reconditioned and new metal furniture ordered to supplement
that already available.
Five rooms are being converted into constant-condition rooms for
biological experimentation with equipment for controlling the tem-
perature, humidity, radiation, and nutritional environment.
SECRETARY'S REPORT is
Four chemistry laboratories will be available, including a general
laboratory, a balance room, a dark room held at room temperature
for pigment analyses, and an insulated and air-conditioned dark room
controlled at 0° C. or above for protein and enzyme analyses.
In addition, a photographic laboratory, a room for X-ray facilities,
a cytology laboratory, an electronics laboratory, and two general
laboratories are being set up, all of which are being designed with
new plumbing to supply gas, compressed air, and water, and with
new electrical power outlets.
The Research Corporation has very generously made a grant to the
Division for reequipping the laboratories with modern experimental
facilities and for work on the mechanism of visible radiation on growth
processes in plants, The Division also has been assigned a contract
by the Chemical Corps, Department of the Army, providing funds for
personnel and equipment for research on the effect of growth regula-
tors on metabolic activities of plant tissues.
Respectfully submitted.
L. B. Aupricn, Director.
Dr. A. WETMORE,
Secretary, Smithsonian Institution.
APPENDIX 9
REPORT ON THE NATIONAL AIR MUSEUM
Sir: I have the honor to submit the following report on the opera-
tions of the National Air Museum for the fiscal year ended June 30,
1949.
INTRODUCTION
This year, the first full year of the National Air Museum as a
bureau of the Smithsonian Institution, was one of many activities.
In addition to normal museum operations, the bureau was concerned
especially with the return from England of the Wright Brothers’
renowned aeroplane, the Kitty Hawk; with the acquisition and man-
agement of a field storage facility; with the accession of the U.S. Air
Force aircraft collection; and with the basic study and planning for
a site and building for the aeronautical collections.
Karly in 1948 the Institution was informed that the late Dr. Orville
Wright had expressed the desire to present the Kitty Kawk to the
United States National Museum and that the executors of Dr. Wright’s
estate would institute the necessary legal action to bring this about.
Prior to the receipt of this news the Smithsonian had effected the
administrative transfer of all aeronautical museum activities and ex-
perienced personnel from the National Museum to the newly estab-
lished National Air Museum. Therefore, in order to have the expert
assistance of the Air Museum staff, the Secretary, through the
Director of the National Museum, delegated to the National Air
Museum the responsibility for the reception, exhibition, and preserva-
tion of the Wright plane. The details are indicated under Curatorial
Activities presented later in this report.
Negotiations begun last year with the U.S. Air Force to acquire a
storage depot for the Air Museum were successfully consummated on
November 1, 1948. On that date the bureau was granted occupancy
of 267,475 square feet of floor space within building T-6 of the former
Douglas Aircraft plant at Chicago Orchard Airport, Park Ridge, IIL.,
and installed a field organization to operate the facility. On the same
date the Museum assumed tentative custody (pending inventory) of
the large aircraft collection stored in this building by the Air Force
and on May 1, 1949, upon completion of the inventory, assumed full
responsibility for its preservation.
114
SECRETARY'S REPORT 115
Along with these several extra activities the bureau continued, with
the Office of Design and Construction of the Public Buildings Ad-
ministration, the further study of sites and a building for the Museum.
This was done in accordance with the recommendation of the Advisory
Board. The result of the study is recorded later in this report.
No changes were made during the year in the departmental organi-
zation of the bureau except the occasional employment of temporary
clerical help. This was especially necessary in connection with the
reception and exhibition of the Wright Brothers’ aeroplane. The
bureau found it difficult, on the other hand, to fill several positions
available at its field storage facility because of the higher wage scale
prevailing in the Chicago area for comparable work. As a result,
the work program planned for this field organization was not fully
carried out. In all other respects the bureau completed the year in
good condition.
ADVISORY BOARD
In April of this year the Board experienced a change in membership
as a result of the retirement of its U. S. Air Force representative,
Maj. Gen. E. M. Powers. General Powers had served on the Board
since its inception late in 1946, having been designated to the office
by General Spaatz. His wise counsel during the formative days of
the establishment of the National Air Museum was most helpful. To
succeed him on the Board, Gen. H. H. Vandenberg, Chief of Staff,
Department of the Air Force, designated Maj. Gen. Grandison
Gardner who met with the Board for the first time at its sixth meeting
in June 1949.
During the year three meetings of the Advisory Board were held
in Washington, on August 26, 1948, December 20, 1948, and June 29,
1949. Deliberations in these meetings were directed principally to-
ward the advancement of the Air Museum’s major projects, namely,
the acquisition of a building site and a suitable museum building in
the Washington area.
As directed by the Board at its August 1948 meeting, the study of
a suitable museum building was continued this year in cooperation
with the Public Buildings Administration.
STORAGE OF MUSEUM MATERIAL
In accordance with a resolution adopted by the Advisory Board
last year, the bureau completed negotiations on November 1, 1948,
to take over the storage operations of that portion of one of the former
Douglas Aircraft buildings (T-6) at the Chicago Orchard Airport,
Park Ridge, Ill., containing the collection of aeronautical museum
material stored there by the United States Air Force for the National
116 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Air Museum. Some additional floor space within this building, ad-
jacent to that containing the Air Force collection, together with two
two-floor structures erected therein and suitable for office use, was
acquired at the same time.
Immediately following this transaction the bureau installed a field
organization to operate the facility, consisting of an associate curator
in charge, an aircraft technician, and a guard force to patrol the area
24 hoursa day. Until February 1, 1949, the military personnel of the
Air Materiel Command of the U. 8S. Air Force, which had been
detailed to care for the Air Force collection, remained on duty to assist
the bureau’s organization in readying itself to assume its responsibil-
ities. With this accomplished, three programs of work were initiated:
(1) The rearrangement by classes of the aeronautical materials packed
in boxes and crates; (2) the cleaning and sealing of all openings of
assembled aircraft and rust-proofing of component parts; and (3)
the inspection and inventory of all items composing the collection
preparatory to its transfer from the Air Force to the Air Museum.
On May 1, 1949, the inventory was completed and the transfer was
effected of 1,366 aeronautical objects including 97 aircraft to the Air
Museum. The preservation work was in progress at the end of the
year.
While these activities were in progress the bureau took steps to
provide the maximum protection of the materials in storage. In
addition to the acquisition of hand fire extinguishers installed in fixed
positions over the area and of larger extinguishers mounted on hand
trucks, an intercommunicating system was selected and a contract
let for its installation at 12 stations distributed strategically over the
Museum’s storage area. This will enable a guard on patrol to com-
municate quickly with the administration office in any emergency.
The bureau also designed and contracted for the construction and
erection of a high wire fence to enclose the major part of the area.
These projects were in progress at the end of the year.
PLANNING
MUSEUM SITE AND BUILDING
During the year the bureau continued the investigation of sites and
a building for the Air Museum. For this purpose it had the valuable
cooperation of the Federal Works Agency, Public Buildings Adminis-
tration, Office of Design and Construction, through an arrangement
involving the transfer of funds.
Planning and designing a museum building for aircraft and aviation
collections involves factors not usually encountered in museum struc-
tures. For example, although the history of practical aviation spans
a comparatively short period of years, the steps in its development
SECRETARY'S REPORT 117
are many. Therefore, there must be imposed limitations of selection
of materials of both historical and technological significance not only
to avoid incomprehensive public displays but also impractical housing
requirements. The aeronautical collections will include material
both of small and uncommonly great dimensions and weight. Ex-
perience indicates that approximately 30 percent of the total available
floor area of a technical museum structure is required for its mainte-
nance and operations services and that for the safety of the visiting
public ample passageways must be established in all exhibition areas.
It can be readily understood that these requirements necessitate a
compromise between the ideal and a realistic aviation museum buiding.
Both the Advisory Board and the bureau’s staff gave careful atten-
tion during the year to factors such as these which brought about a
number of changes in the plans originally developed last year.
CURATORIAL ACTIVITIES
The curator, Paul E. Garber, reports on the year’s work as follows:
At the beginning of the fiscal year the staff was moved into new and
improved quarters which provided more facilities for the expanding
personnel. Office and shop equipment were acquired, an efficient
procedure for handling correspondence was adopted, and added space
was allocated to the library, the reference files, and the photographic
files. The constant efforts of the staff in the maintenance of the
exhibits are reflected in the improvement of individual displays, but
the extreme overcrowding in the present Aircraft Building and the
Aeronautical Hall assigned to the bureau in the Arts and Industries
Building has approached the danger point to both visitors and speci-
mens. Asaresult, the addition of large exhibits has been brought to a
standstill. Happily, the facility at Park Ridge, Ill., provides for the
storage of material which might otherwise be lost to the Museum.
Only by the acquisition of a permanent building for the Museum in
the Washington area can this situation be corrected.
EXHIBITION
The outstanding accomplishment of the year was the receipt of the
Wright Brothers’ aeroplane of 1903 and its exhibition and preparation
for the presentation ceremony on December 17, 1948, the forty-fifth
anniversary of its historic flight. The curator was assigned the
pleasant duty of representing the Smithsonian in meeting Dr. Herman
Shaw, Director of the Science Museum, London, and of accepting
from him at Halifax, Nova Scotia, custody of the aeroplane, This
was effected on November 12, and following the transfer of the aero-
plane from the S. S. Mauretania to the Navy Carrier U.S. S. Palau,
the curator accompanied the plane to Bayonne, N. J., saw to its re-
loading on a Navy truck and accompanied the truck convoy to the
118 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
National Museum. There the Spirit of St. Louis had been moved so
that the two noted aircraft could share the same North Hall of the
Arts and Industries Building and, with the assistance of photographs
and drawings provided by M. J. B. Davy of the Science Museum, the
Kitty Hawk was assembled. Some details of the engine and trans-
mission were installed from sketches and photographs furnished by
Charles Taylor, the mechanic who assisted the Wrights in the original
construction of the plane and engine. The aeroplane was suspended
in the front of the hall with cables and splices donated by the Jacoel
Equipment Co. At the ceremony of presentation, the “Karly Birds,”
the association of pioneer pilots, many of whom had been trained by
the Wrights, were among the honored guests. A number of them have
since augmented the Kitty Hawk exhibit by donations of their own
records and relics. Acknowledgments are also made to the Air Force
Technical Museum and to S. Dunham, Dayton, Ohio, for photographs
and drawings for addition to the Museum’s reference and exhibition
material on the Wrights. At the close of the fiscal year progress is
being made on an auxiliary exhibition case to be placed under the
Kitty Hawk in which the story of the Wright Brothers will be told in
detail.
As the National Air Museum progresses, its purposes and services
have become better known, and very helpful cooperation has been
received in the matter of accessions. The following examples are
outstanding. The presentation of Alford Williams’ renowned Gulf-
hawk-2 October 11, followed an impressive flight-demonstration of the
remarkable aerobatic combination of pilot and plane. The Gulf Oil
Company formally presented the airplane at the National Airport and
soon after, Major Williams’ technician, Frank Tye, who had main-
tained the plane in splendid condition throughout its 12 years of stren-
uous flying, assembled it in the Aeronautical Hall. The Department
of the Navy, Bureau of Aeronautics, repaired and transferred a Jap-
anese Baka Bomb to the Museum and provided, as auxiliary material,
examples of both jet and rocket engines used in these ‘‘suicide planes.”
The collection of scale models which reviews the evolution of aircraft
used in Naval service was improved by 10 recent types received from
manufacturers who produced the original planes for the Navy. The
Department of the Navy assisted also in the special anniversary cele-
bration held in the Aircraft Building to commemorate the thirtieth
anniversary of the first trans-Atlantic aircraft flight made by the
NC-4. An illustrated description of this Curtiss-built flying boat,
the hull of which is in the Museum, was prepared by the staff and
printed by courtesy of the Curtiss-Wright Corporation. Speakers
included Vice Adm. John D. Price, U. S. N., and Capt. Holden C.
Richardson, U.S. N., Ret.
SECRETARY’S REPORT 119
The first Roadable Autogyro, transferred from the Civil Aeronautics
Administration, was reconditioned by them for museum purposes.
In the previous report, the services of the Air Force were acknowledged
in moving the Army Curtiss Racer from the Aircraft Building to the
Aeronautical Hall—a move made necessary by restricted space. This
year, the plane was equipped with the original floats with which Lt.
(now General) James Doolittle won the Schneider Trophy Race in
1925. The bracing wires for this restoration were kindly provided
by the MacWhyte Company, Kenosha, Wis. Valuable assistance
was received this year from the Air Force in unloading and mounting
four large engines donated by the Wright Aeronautical Corporation,
in covering with Plexiglas the sides of the DeHavilland—4 and Gen.
William Mitchell’s Spad—16, and in replacing the windows in the first
nonstop transcontinental airplane, the 7-2.
The exhibit which illustrates the accomplishments of John Joseph
Montgomery of California, a renowned pioneer of gliding whose first
glides were made in 18838, received additions through the cooperation
of the Montgomery family, the San Diego Junior Chamber of Com-
merce, and the biographer, Winsor Josselyn. ‘Through the generosity
of the Firestone Tire and Rubber Company, the wheels on the Voisin
bomber of World War I were equipped with tires. In the auxiliary
exhibit which accompanies the Flagplane of the First World Flight,
the group of portrait sculptures were renovated by its sculptor, Joseph
A. Atchison, and the stereopticon story of this flight was reactivated.
A special exhibition of model aircraft as flown by hobby enthusiasts
was prepared in August 1948 during the period of the national show
and contest. Improvements were made in the display of Col. Charles
Lindbergh’s accessories and flight clothing. Extensive cleaning,
rearranging, labeling, and repairing have brought the exhibits to a
condition believed to be as presentable as the crowded conditions and
work program permit.
Special exhibitions arranged by the staff and involving the use of
Museum material away from the bureau included a group of cases
set up by Bolling Field for the Air Force anniversary on September 18,
containing engines, models, and relics of the military air arm; and the
loan of the original Liberty engine and models of historic Air Force
planes for the technical exhibit and air show held at Andrews Field,
Maryland, February 15.
STORAGE
Among the numerous aircraft installed this year in the Museum’s
storage facility at Park Ridge, IIl., was the Swoose, flown there under
its own power. This historic B-17—D bomber had served throughout
World War II from Bataan to the defeat of Japan. Completing its
military career as the command plane of Gen. George H. Brett, the
866591—50 9
120 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Swoose was acquired by the city of Los Angeles as a war memorial.
In 1948 the city, through Mayor Fletcher Bowron, presented the plane
to the National Air Museum, and with the cooperation of Grover
Loening and Maj. Gen. E. M. Powers of the Advisory Board, arrange-
ments were made for the reconditioning of the plane by the Air Force.
In due time this was accomplished under the direction of the Swoose’s
wartime flight engineer, Captain Boone, and in April 1949, with its
wartime pilot, Col. Frank Kurtz, at the controls, the plane was flown
to the Chicago Orchard Airport and delivered there to the Museum’s
storage facility. The Air Force cooperated not only in this spectacular
delivery but also in the tremendous project of transferring its huge
collection of trophy aircraft and accessories to Museum custody.
The screening, cataloging, and arrangement of this stored collection
at Park Ridge was under way as the year closed.
The historic and trophy aircraft and engines which are being
assembled for the Museum by the Department of the Navy are
stored at Norfolk, Va. During the year the curator inspected this
facility, checking the condition of the NC—4’s wings and other parts,
the Japanese ‘‘Emily,’”’ the German Dornier 335 which had recently
been moved there, and the service types which are in ‘‘canned”’ con-
tainers. All were in good condition.
Acknowledgments are made to Eastern Air Lines for earmarking
one of its first DC-3’s for the collection, to the Civil Aeronautics
Administration for reserving its famous Boeing 247—D for the Mu-
seum, and to the Martin Aircraft Company for the gift of a half-scale
flight prototype of the PBM ‘“Mariner.”” These will be stored tem-
porarily by the donors.
INFORMATIONAL SERVICES
Interest in the Museum is widespread, and its services to the
industry, Government departments, students, research workers, his-
torians, authors, craftsmen, and the air fraternity in general are daily
becoming more in demand as reflected in the numbers of inquiries
and requests received by letter, personal visit, and telephone. Radio
programs in which the Museum participated included, Information
Please, We The People, and the Air Force Hour. The curator told
the story of the Swoose over the radio both in Los Angeles and Omaha,
and television programs illustrated the Kitty Hawk, Gulfhawk-2, the
NC-4, and the Museum’s model collection.
The Bureau of Ordnance, Department of the Navy, borrowed a
number of the Museum’s scale models to be used as patterns for re-
search problems; the Interior Department was assisted with aero-
nautical details in some of its museum dioramas; the Public Schools
of the District of Columbia received help in conducting their aero-
SECRETARY’S REPORT 121
nautical courses; and Pan American Airways was loaned photographs
of Santos-Dumont’s airships for use in its publicity displays on Brazil.
The historic sections of the Aircraft Year Book were compiled with
help of the Museum staff; Bettman Archive received identification on
a group of unlabeled photographs of airplanes; the Prewitt Aircraft
Company used the Museum’s reference files during their search for
details of rotary aircraft; and the United States Chamber of Com-
merce and the Vallejo, Calif., museum, received assistance in display-
ing exhibits.
Another edition of the Handbook of the National Aircraft Collection
was issued, embodying changes which bring it up to date. This is
the eighth printing of 10,000 since the first issue in 1928. During the
year the curator lectured on technical and historical aspects of flight
and the progress of the National Air Museum to the Aero Club of
Washington, the Air Transport Association, the “99ers” association
of women flyers, the Washington Association of Building Superin-
tendents, the Civitan Club, several local fraternal and church groups,
and served as judge of scale-model craftsmanship at the National
Capital Air Show, and at a kite contest held by local units of the
Boy Scouts.
In conducting its informational services the staff acknowledges the
help given by members of the “Early Birds,” collectors of aeronautical
photographs and clipping scrapbooks, pilots, manufacturers, airmen,
and many others who donated reference material to the Museum’s data
files and library. These helpful source data are assembled and readily
available for serious study.
SURVEY
The survey over the Nation of aeronautical materials of technical
and historical significance was continued during the year. Much of
the work was conducted by staff correspondence. Frequently, how-
ever, it became necessary to undertake direct investigation and study
of suggested material and consultations with those acquainted with
the material. It was in this connection, primarily, that the following
visits away from Washington were made by the staff:
Middletown, Pa., Olmsted Air Force Base, July 23, by associate curator Robert
C. Strobell, to examine a group of Japanese trophy airplanes which had been
evaluated and tested.
Buffalo, N. Y., Airport, August 31, by associate curator Stephen L. Beers, to in-
spect two Curtiss engines used in early Naval aircraft and two French engines
of World War I.
Roosevelt Field, Long Island, N. Y., and East Orange, N. J., October 11-16, by
the curator, to examine a group of historic American, English, and French
airplanes and to determine the availability of a Benoist airplane of 1912.
Halifax, Nova Scotia, November 9-22, by the curater, to obtain the Wright
Brothers’ aeroplane of 1903.
122 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Los Angeles, Calif., Inglewood Field, and March Field, January 19-29, by the
curator, to receive, recondition, and test hop the Boeing B-17~D bomber
Swoose.
Aberdeen and Baltimore, Md., March 15, by Mr. Beers, to inspect aircraft and
obtain a scale model of the Martin bomber, type MB-1 of 1918.
Jackson Center, Ohio, May 28, by Mr. Strobell, to arrange receipt and shipment
of Benoist airplane.
Langley Field, Va., May 23-25, by the curator and Mr. Beers, to attend the
National Advisory Committee for Aeronautics conference and inspect
material of museum significance.
Miami, Fla., June 15-18, by Mr. Beers, to attend Eastern Airlines conference and
inspect equipment and aircraft made available to the Museum.
Numerous trips were made by staff members between the base and
field units in connection with management of the storage area and
procurement and placement of accessions.
ACCESSIONS
New accessions totaled 122 objects from 40 sources. The majority
of these were solicited by the Museum and involved considerable prior
research by the staff to determine the significance and need of each
object in the over-all picture of the history and development of aero-
nautics. In addition, each accession required staff attention in a
variable amount in arranging for the procurement and shipment of the
object and in incorporating it into the aeronautical collection.
Except where otherwise indicated the accessions received this year
and listed below were entered in the Museum’s records as gifts or
transfers:
NATIONAL AIR MUSEUM ACCESSIONS DURING THE FISCAL YEAR ENDED
JUNE 80, 1949
ABEL, A. H. (See under Port of Oakland, Board of Port Commissioners.)
ABRAMS, TALBERT (See under Abrams Instrument Corp.).
ABRAMS INstRUMENT Corp., Lansing, Mich: (Through Talbert Abrams) The
Explorer, single pusher monoplane with empennage extended on twin booms;
believed to be the first American aircraft designed primarily for aerial mapping
and survey work (N. A. M. 629, loan).
AERONCA ArrcRAFT Corp., Middletown, Ohio: (Through John A. Lawler) First
production Aeronca sport plane, 1929 (N. A. M. 647, loan).
Arropropucts Division, GeNERAL Motors Corp., Dayton, Ohio: (Through
W. F. Stover) Four Aeromatic propeller assembly displays illustrating types,
mechanisms, and production steps (N. A. M. 651).
ALIHAN, Dr. Mixa (See under Kollsman Instrument Division, Square D Co.).
ALLEN, Witu1aM B., Jr. (See under U. 8S. Post Office Department.)
Bastow, J. G., Oakland, Calif.: American flag insignia of the First Aero Squadron,
World War I, cut from the fuselage fabric of a Salmson airplane (N. A. M. 625).
Brcx, Toomas H. (See under Crowell-Collier Publishing Co.)
BisicuKkow, WIiLu1AM. (See under Comet Model Airplane and Supply Co.)
SECRETARY’S REPORT 123
Bravpiey, R. F., San Francisco, Calif.: A framed black-and-white photograph of
an early air meet and a souvenir log book of the first round-trip, trans-Pacific
Pan American Airways passenger flight, 1936 (N. A. M. 636).
BriskIN, Irvine. (See under Columbia Pictures Corp.)
Brooxityn Pouyrecunic Instirure, Brooklyn, N. Y.: (Through Dr. D. G.
Lockward) A 160-hp. Curtiss V-X aircraft engine used to power a Curtiss R-2
Army reconnaissance plane cf 1916 (N. A. M. 631).
CuHauuinor, G. R. (See under Kansas City, Mo., Chamber of Commerce.)
CuHancr Voucur Arrcrart, Div. or Unirep Arrcrart Corp., Stratford, Conn.:
(Through John J. Hospers) Two 1:16-scale airplane models, U. S. Navy,
World War II: F4U-—4 and OS2U-1 (N. A. M. 630).
Con, Don, Williamsville, N. Y.: Two aircraft engines, rotary, French, World
War I: a Clerget 9B and a Gnome ‘‘Monosoupape”’ (N. A. M. 655).
Conus, C. 8. (See under Pan American-Grace Airways, Inc.)
CoutumBiA Pictures Corp., Hollywood, Calif.: (Through Irving Briskin) Two
scale medels of Prof. John J. Montgomery’s gliders reproduced for the motion
picture “Gallant Journey.’”’ These show the Santa Clara of 1905 and the
Evergreen of 1911 (N. A. M. 626).
Comer Moprt AIRPLANE AND SuppLty Co., Chicago, Ill.: (Through William
Bibichkow) Wind tunnel stated by donor to be the first practical mass-produced
type; for personal and classroom use; throat 14” x 22”; embodies the principles
of full-scale designs; accessories are included (N. A. M. 643).
CROWELL-COLLIER PuBLIsHING Co., New York, N. Y.: (Through Thomas H.
Beck) Original water-color painting by Melbourne Brindle depicting the first
flight of the Wright Brothers’ aeroplane at Kitty Hawk, N. C., December 17,
1903 (N. A. M. 654).
DeHart, Dana C., San Francisco, Calif.: A major portion of a cabane strut from
the Curtiss ‘‘R’’ which was one of the first two planes to carry scheduled air
mail from New York to Chicago via Cleveland, September 5 to 17, 1918
(N. A. M. 628).
Dintz, Goutp (deceased), Omaha, Nebr.: Two wooden propellers: a ‘‘Paragon’”’
1911, used on the Army’s first airship, and a “Flottorp” of early 1920 type
inscribed with famous autographs (N. A. M. 644).
DoouiTtLE, Gun. J. H., New York, N. Y.: A jagged fragment, found in China,
from the right-engine nacelle of a North American B-25 “Mitchell”? bomber
which took part in the Tokyo raid of April 18, 1942 (N. A. M. 604).
Dovua.as Arrcrart Co., Inc., Santa Monica, Calif.: A 1:16-size scale model of
the Douglas SBD Navy carrier-based dive bomber which saw extensive service
in World War II (N. A. M. 620).
Frank, JouHNn P. (See under North Carolina Granite Corp.)
Frrprericx, D. 8. (See under Rohm and Haas Co.)
Friss, LEonarpD, London, England: A large display poster printed from the
original painted by the donor, advertising the International Aviation Tourna-
ment held at Belmont Park, L. J., October 22 to 30, 1910 (N. A. M. 648).
Gitcnrist, Mrs. Guy, Dutch Flat, Calif.: The starter’s flag used in the Oakland-
Honolulu Dole Race, 1927. Given in memory of her brother, Maj. Edward
Howard (N. A. M. 653).
Goopwin, Cuiaire V. (See under Port of Oakland, Board of Port Commissioners.)
Grumman, L. R. (See under Grumman Aircraft Engineering Co.)
GRUMMAN AIRCRAFT ENGINEERING Co., Bethpage, L. I., N. Y.: (Through L. R.
Grumman) Four 1:16-size scale models of Grumman aircraft, U. S. Navy,
World War II: an F3F biplane fighter, an F4F ‘‘Wildcat,” a TBM ‘Avenger’
torpedo bomber, and an F7F “Tigercat” (N. A. M. 634).
124 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Gur Orn Corp., Pittsburgh, Pa.: The Gulfhawk—2 airplane which was flown by
Maj. Alford Williams for 12 years. It illustrates the last of the Navy’s biplane
fighters, an F8F (N. A. M. 652).
HEFFERNAN, Carr. J. B. (See under National Military Establishment, Depart-
ment of the Navy.)
Hosprrs, Joun J. (See under Chance Vought Aircraft, Div. of United Aircraft
Corp.)
Hussewi, Cuarues H., Cleveland, Ohio: Twelve color prints of current-design
private aircraft as painted by the donor for the 1949 calendar of Thompson
Products, Inc. (N. A. M. 642).
Kansas Crry, Mo., CaHamBer or Commerce: (Through G. R. Challinor) A
pressed-coal briquette, carried on Berlin Airlift, 1949; part of the one-millionth
ton. It was mined in the Ruhr; flown from Frankfurt to Berlin and thence to
Kansas City, Mo., as a feature of a ceremony acclaiming the Airlift (N. A. M.
645).
Kartveut, A. (See under Republic Aviation Corp.)
KoLusMAN INSTRUMENT Diviston, SeuarE D Co., Elmhurst, N. Y.: (Through
Dr. Milla Alihan) A machmeter; an instrument used to record speed in Mach
Number, which expresses the speed of the aircraft in relation to the speed of
sound (N. A. M. 649).
LAWLER, JOHN A. (See under Aeronca Aircraft Corp.)
LocxwarpD, Dr. D. G. (See under Brooklyn Polytechnic Institute.)
Menruor, Kenneto ©. (See under Wright Aeronautical Corp.)
NatrionaL Miiitary EsTaABLISHMENT:
Department of the Air Force. From Air Force Technical Museum, Wright-
Patterson A. F. Base, Dayton, Ohio: Five 1: 72-size, black plastic World War
II aircraft recognition models: a C—47 two-engine transport, a PB2Y four-
engine fiying boat, a PBM-8 two-engine flying boat, an SB2U single-engine
Navy dive bomber, and a TBD single-engine torpedo bomber (N. A. M. 638).
Department of the Navy, Washington, D. C. From Office of the Deputy Chief
of Naval Operations (Air): (Through Vice Adm. John D. Price) A plaque,
replica of one installed at Lisbon, Portugal, in 1946, commemorating the first
trans-Atlantic flight by the NC-4, May 1919 (N. A. M. 646). From Office
of Public Information, Bureau of Aeronautics: A Navy summer-weight flying
suit and helmet, World War II type, worn by Col. Marion E. Carl, U.S. M. C.,
when he flew the Douglas D-558 Skystreak to a world’s speed record of
650 m. p. h. on August 25, 1947 (N. A. M. 637). From Office of Public Rela-
tions: (Through Capt. J. B. Heffernan, U. S. N., Ret., Curator for Navy
Department) Scale models of three planes used by the Navy in World War II:
XPB2Y experimental version of Consolidated ‘‘Coronado”’ used for bombing
and transport; XPBM experimental version of Martin ‘“‘Mariner’” used for
patrol-bombing and antisubmarine service; and TBD Douglas ‘Devastator’
carrier-based torpedo plane (N. A. M. 635, loan).
Nevin, Rosert §., Baltimore, Md.: A 1:24-size scale model of the Martin MB-1
twin-engined Army bomber of 1918, made by lender (N. A. M. 621, loan).
NortH CaroLInaA GRANITE Corp., Mount Airy, N. C.: (Through John P.
Frank) Architects’ model of the Wright Brothers Memorial on the summit of
Kill Devil Hill, Kitty Hawk, N. C. (N. A. M. 6388).
Pan AMERICAN-GRACE ArRways, INc., New York, N. Y.: (Through C. §. Collins)
The Fairchild FC—2 five-passenger cabin monoplane with which the scheduled
commercial operations of Panagra System in South America were inaugurated
September 1928 (N. A. M. 650).
SECRETARY’S REPORT 125
Perry, Kenneta M., Washington, D. C.: A German Air Force garrison cap of
the World War II period (N. A. M. 622).
Port OF OAKLAND, Boarp or Porr ComMISSIONERS, Oakland, Calif.: (Through
A. H. Abel) The “‘Diamond”’ airplane, identified by donors as the first airplane
constructed in California (1910), and the Kemp engine used to power it; a
“Wiseman-Cooke” airplane constructed by Fred Wiseman and flown by him in
what was probably the first cross-country air-mail flight in America, Petaluma
to Santa Rosa, Calif., February 17, 1911. This plane was also flown by Weldon
Cooke (N. A. M. 639). (Through Claire VY. Goodwin) A pilots’ control wheel
from the cockpit of Sir Charles Kingsford-Smith’s Southern Cross airplane, first
to fly from the United States to Australia, May 31—June 9, 1928; and a facsimile
of the log kept by the copilot, Charles Ulm, on that flight (N. A. M. 640).
Pricn, Vich Apm. Joun D. (See under National Military Establishment, De-
partment of the Navy.)
Rerusiic Aviation Corp., Farmingdale, L. I., N. Y.: (Through A. Kartveli)
three 1:16 scale models: Republic P—47—-N ‘“‘Thunderbolt”’ Air Force single-seat
fighter, Republic F-—84 “Thunderjet’”’ Air Force single-seat fighter, Republic
RC-3 “Seabee’’ all-metal, four-place amphibian personal plane; and a 1:48
scale model: Republic XF-12 “Rainbow” Air Force four-engine, long-range,
high-altitude photo-reconnaissance plane (N. A. M. 641).
Roum anD Haas Co., Philadelphia, Pa.: (Through D. §. Frederick) Four ex-
amples of Plexiglas forms used in construction of aircraft: a P—47 bubble canopy,
an astradome, a Crocker-Wheeler nose section for B—26 aircraft, and a Sikorsky
Helicopter nose (N. A. M. 627).
Spratt, GeorGE, Deep River, Conn.: A Curtiss V—8 air-cooled aircraft engine
of about 1907, used by the donor’s father in experiments with movable-wing
aircraft (N. A. M. 624).
Stover, W. F. (See under Aeroproducts Division, General Motors Corp.)
U.S. Post Orrick DepartTMENT, Washington, D.C.: (Through William B. Allen,
Jr.) The pouch used to carry air mail on a flight commemorating the thirtieth
anniversary of air mail; a flight was made between New York and Washington,
D. C., in an Air Force P-80 jet aircraft (N. A. M. 619).
Viuas, Jack, Chicago, Ill.: Hull of the Curtiss Flying Boat in which the donor
made the first flight across Lake Michigan on July 1, 1913 (N. A. M. 632).
Waker, Mas. Tuomas L., Glen Echo, Md.: Japanese equipment, World War
II: an Hitachi aircraft engine from a ‘‘Cypress” biplane primary trainer, a
cutaway supercharger and fuel metering device produced by Mitsubishi, and
a group of five instruments also manufactured by Mitsubishi (N. A. M. 600).
Weems, Capt. P. V. H., Annapolis, Md.: A collection of plotters illustrating
many forms used to solve navigation problems involving position, direction,
and distance (N. A. M. 656).
Wricut, Orvititr, Estate or, Dayton, Ohio: The original Wright Brothers’
aeroplane of 1903 (in custody for the U. S. National Museum) (U. 8S. N. M.
181390).
Wricut AmronauticaL Corp., Wood-Ridge, N. J.: (Through Kenneth C.
Mebrhof) Four radial aircraft engines: Curtiss ‘‘Challenger,”’ Wright ‘‘Cyclone’”’
18BA, Wright “Cyclone” 14A, and Wright “‘Cyclone”’ 9GB (N. A. M. 623).
Respectfully submitted. Cart W. Mirman,
Assistant to the Secretary for the National Air Museum.
Dr. A. WeTmorg,
Secretary, Smithsonian Institution.
APPENDIX 10
REPORT ON THE CANAL ZONE BIOLOGICAL AREA
Sir: It gives me pleasure to present herewith the annual report of
the Canal Zone Biological Area for the fiscal year ended June 30, 1949.
IMPROVEMENTS MADE
A new building, halfway up to the laboratory level and near the
generators, was completed. The ground floor will be used largely
for the woodworking machinery and carpenter shop, the upper part
as living quarters for the warden-caretaker. ‘The old cottage just
below the large laboratory building, formerly occupied by the warden-
caretaker, has been converted into a very desirable two-room lab-
oratory unit. The forest is close by, making the building exception-
ally suitable for observation and study of mammals and other forms.
Work on the 14,000-gallon concrete gravity-flow water tank was
halted by heavy rains which made it impossible to use the truck and
large concrete mixer. However, the excavation is made and the
gravel and reinforcement iron are at the site; 2 or 3 weeks of dry
weather will permit completion of the tank.
During the year, from allocated funds, the launch Snook was
purchased from The Panama Canal. This is a very sturdy, well-
built boat 40 feet long, with an 11-foot beam—large enough to ac-
commodate 40 passengers. It is in very good condition and will
serve even for towing.
New generators were installed during the year, providing a depend-
able source of electricity for continuous use.
Eight steel herbarium cases, needed for many years, have been
received and the herbarium specimens transferred to them. These
specimens are now in excellent condition. Eight steel storage cabinets
were also received, providing dustproof storage for much of the
laboratory equipment.
SCIENTISTS AND THEIR STUDIES
Twenty-nine scientists came to the laboratory during the year.
Although many of them stayed only a short time, their acquaintance
with the island and its facilities will no doubt bring others to the
laboratory in the near future. The contribution of these scientists
has added materially to our knowledge of life under tropical conditions.
126
SECRETARY’S REPORT 127
Dr. Franz Schrader and Dr. Sally Hughes Schrader returned to
continue their cytological studies.
Dr. Frank A. Hartman and Robert Albertin, of the Department
of Physiology of the Ohio State University, spent some time on the
island, and used the laboratory as their base for excursions up the
Chagres River Basin and into the Volcan region of Chiriqui Province.
They studied in great detail the anatomy of the adrenals in sloths
and the coati-mundi while on the island. The adrenals of more
than 600 vertebrates were collected during expeditions in the Republic
of Panama. Field studies were made of selected cases, and the rest
of the material was taken to the island for further treatment in prep-
aration for cytological examination later at their University labora-
tory. The skins of the birds obtained by them were donated to the
United States National Museum. The hearts of a number of the
vertebrates were sent to Dr. Struthers, of the University of Syracuse,
for anatomical study of the coronaries and other blood vessels.
Dr. Hartman plans to return for a much more extensive survey,
especially in relation to the effects of the male hormone, particularly
in the sloth. The use of the island as a base for excursions to other
nearby regions emphasizes one of the unique features of the Canal
Zone Biological Area.
Dr. Per F. Scholander and Dr. Vladimir Walters, of the Arctic
Research Laboratory at Point Barrow, Alaska, spent considerable
time on the island studying the metabolic reactions to temperature
in various animals and plants in order to obtain a tropical counterpart
for the work done on Arctic forms in Alaska. A deep-freeze was
used in some of their work, and the analysis of the tropical mammals
and birds in cold gave just the information needed to interpret
properly the findings in the Arctic. The work on the island with
the deep-freeze proved of basic importance for formulating a theory
on the relation between insulation and metabolism.
R. Joseph Kowal, in charge of the laboratory of the Bureau of
Entomology and Plant Quarantine at Gulfport, Miss., returned in
order to reexamine and evaluate the special series of termite-resistance
tests initiated by him 5 years ago, including the very valuable series
of soil poisons. He was assisted by Russell EK. Fontaine, in charge
of the insect- and pest-control work of the United States Army in
the Caribbean area.
Dr. Walter Clark, in charge of the Research Laboratory of Eastman
Kodak, who had come to inaugurate the large air-conditioned labora-
tory in the outskirts of Panama City, accompanied by Dr. Cleve C.
Soper, in charge of Eastman Kodak’s tropical research work here,
spent several weeks on the island in connection with their corrosion
and deterioration studies. Asin past years, Dr. Clark took thousands
128 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
of feet of splendid motion pictures of the animal life. The work of
the Eastman Kodak Company on the island, and in Panama City,
is yielding most valuable data on corrosion and deterioration, as
well as a better understanding of the influence of the Tropics on
color film.
EK. P. Killip, head curator of the department of botany, United
States National Museum, spent a short time on the island, collecting
material for the herbarium. He went over most of the laboratory’s
herbarium specimens and later will send such additional sheets as
may be needed to augment or supplement the collection.
Dr. Charles C. Adams, pioneer ecologist of the Americas, was
another visitor who came to the Tropics for first-hand knowledge.
He stressed the value of the island for ecologists, both plant and
animal, in providing an intimate acquaintance with jungle life.
Dr. and Mrs. H. N. Moldenke, of the New York Botanical Garden,
visited the island after their return from the Second Pan American
Botanical Congress held in Tucum4n, Argentina. Dr. Moldenke’s
chief interest was the Verbenaceae, and he very kindly rechecked the
laboratory’s material in this family.
Dr. Marshall Stone, of the University of Chicago, again revisited
the island for a short time.
Phil W. Longenecker, student at Colorado College, spent the month
of July on the island. He made a list of 98 species of birds that he
positively identified, with copious notes on their habits. His
studies also included observations on a number of the mammals.
In his report he states that he did not find it necessary to go far into
the forest, as there was so much to see within a mile of the laboratory.
Dr. Eugene Eisenmann, of New York City, visited the island again
this year to continue his ornithological studies. Dr. Eisenmann is
an authority on the birds of this region. He will prepare a list,
brought up to date, of the birds observed on the island.
Oliver E. Mosser, of Smithtown Branch, N. Y., came for a few
days, specifically to make certain important observations of army
ants for Dr. Schneirla. In addition he studied birds and mammals.
Frank W. Hunnewell and his sister Louise revisited the island and
stayed several weeks, following up his botanical studies. They
showed the same deep interest in the island that they had displayed
ever since Dr. Barbo:ir was actively connected with its direction.
lt is a pleasure to record again a short visit by Dr. and Mrs.
Matthew W. Stirling and Richard Stewart, who were in Panama on
archeological reconnaissance on behalf of the National Geographic
Society and the Smithsonian Institution. Motion pictures were
taken by Mr. Stewart to complete the reel covering the island.
Dr. Wetmore, Secretary of the Smithsonian, revisited the island,
SECRETARY’S REPORT 129
at which time conferences were held with the writer on island matters,
plans for the future, improvements that would be desirable. W. M.
Perrygo, of the National Museum, accompanied him as assistant.
John E. Graf, Assistant Secretary of the Smithsonian Institution,
spent 2 weeks on the island in June and July, examining the laboratory
facilities and discussing its operations.
A. C. Langlois, of the Bahamas, whose deep interest in palms and
horticulture in general are well known, was a welcome visitor for a
few days, during which brief time he was able to observe the palms as
they grow in their natural habitat.
W. E. Lundy, secretary-treasurer of The Panama Canal Natural
History Society, a keen student of nature, spent a week on the island,
and as on earlier visits prepared a detailed report of the mammals,
birds, reptiles, and other forms that he saw. These yearly lists from
Mr. Lundy form a very valuable record.
Miss KE. Thomas and Miss Marie Weir, local naturalists of note,
again revisited the island for a few profitable days.
The writer continued his special research problems, particularly
the long-term termite-resistance tests, and fruit-fly populations. The
large Berlese funnel has been kept in operation, and it is of interest
to note the great number of species of mites, pseudoscorpions, and
ants, particularly some of the very rare genera, that have been
collected in this manner.
An interesting development from work done at Barro Colorado
Island that should be mentioned here, since it has not previously
been noted, is the availability of a phonograph record of jungle sounds
by day and by night, familiar and friendly to those who know them,
mysterious and sometimes fearful to the uninitiated. The work is
that of Dr. Arthur A. Allen and Dr. Paul Kellogg of Cornell University
who, during the war, made a long series of recordings in the jungle
for training use with American troops assigned to outpost duty. A
considerable part of the work was done on Barro Colorado Island,
though it is only recently that the material has been released and
prepared in form available to the public. The voices of howler
monkeys, birds, and amphibians, reproduced faithfully by painstaking
techniques, carry fully the ordinary sounds heard during the 24 hours
about the Island.
MORE URGENT NEEDS
One of the most urgent needs is the fireproofing of certain structures
by the use of concrete posts and concrete blocks, which could be
accomplished gradually. These buildings are the Barbour and Chap-
man houses, the kitchen and its adjacent storerooms, and the main
laboratory building.
130 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Even more urgent is a new laboratory and storage building, which
will require a small addition to the clearing back of our present
laboratory site. This structure should be built with concrete posts
and sills, and concrete-block sides, and should have at least six rooms.
Provision should be made for the periodical use of heat in order to
reduce the growth of molds. ‘This building would provide separate
rooms for the library, for scientific records, for storage of cameras
and other delicate apparatus, for the herbarium, for laboratory glass-
ware, and for chemicals. The entire building should be well ventilated,
but in addition glass windows should be provided so that when
necessary these can be tightly closed and heat used. High humidity,
the subsequent rapid growth of fungus, and the need of protection
from termites are problems of the first order in the Tropics.
Other urgent needs are steel storage cabinets, metal bookcases
with closing glass fronts, metal card-index cabinets for the species
index and the library index, and sectional steel letter files.
TaBLe 1.—Annual rainfall, Barro Colorado Island, Canal Zone
Total Stalion Total Station
Year inches average Year inches average
192522223 eee (042g ie Sea se 119 footer 124. 13 110. 12
192622. ees ree! 118. 22 PUSH GW OSSs ts: Sa See = 117. 09 110. 62
Cys eee 116. 36 ANA 68: OS Oh Se 115. 47 110. 94
LO2Sase 2 ee ee 101. 52 Ps So O40 Ss ae ee ee 86. 51 109. 43
D2 Qe ts 2 eel 87. 84 HOGT DG OL2S aes ae eee 91. 82 108. 41
NOS ORS s eee ee 76. 57 1 ACO 1 ty 2 a SO 108. 55
LOSI OS ea eee 1238. 30 04769) 1943222. See 120. 29 109. 20
LOS2 etres chon ees 113. 52 LOSS 7 GN NOLES 22 ogee Ss 111. 96 109. 30
NOG Sees oe er LOTS LOS O21 O45 2> 22 se Se ae Se 120. 42 109. 84.
LOS 4S eee ee 122. 42 1OGO4 N94 Ge ee oe eee 87. 38 108. 81
19S es Seu eet ee 148. 42 1UNOVS DOA ee ree bee ae 71. 92 107. 49
LOSG 4252 2 pee 93. 88 LOS: 98iO4S =e a ee 83. 16 106. 43
TABLE 2.—Comparison of 1947 and 1948 rainfall, Barro Colorado Island, Canal
Zone (inches)
Total Accumu-
Station | Years of | Excessor | lated ex-
Month average | record j|deficiency| cess or de-
ficiency
DANUSLY See ee ee ee 1,84 2M ee ne eee | ae eee
Webruarys ee See as fea ee 1, 22 23 —1.03 —1.03
Marchese: 5 eae ener eney, Sa mye ee 1.36 23 —1.19 —2, 22
Aprile sche 2 RS oe ee aes ea ee 2.81 24 +0. 11 —2.11
ia y eee ee ae ee eae a eee em ee 10. 85 24 —0.05 —2.16
JN eee es ee ee 11.10 24 —4, 78 —6. 94
BIL (eee ge ee pe ee ee ee ee 11. 61 24 —0.16 —7.10
IANIQUB tH eae ee ee ee ee 12. 44 24 —1.98 —9.08
September. a a eee 10. 27 24 —3.5 —12. 63
Ostober2223 so ae eae eee 13. 07 24 —2. 33 —14. 96
November! 25-220 ose oe ie ot eee eS 18. 84 24 +1. 49 —13. 47
December ss sess aes ee ee 11.02 24 —9. 80 —23. 27
TVCAt! See Si aeen oe ee ee 1OG2437 baw =. 855 2 eee —23. 27
(Dye aee as =e ae re ee TQRIESE ES S| ees —5. 36
NG) eee ee See Se ee ee ee 99520-22282. 25.255. 35-3 —17. 91
SECRETARY’S REPORT aul
FISCAL REPORT
During the fiscal year 1949, $12,256.05 in trust funds was available.
Of this amount $11,118.90 was spent, leaving on hand only $1,137.15
with which to face the new fiscal year. In addition to this, $1,122.30
is still on deposit, representing local collections.
During the year only $1,243.00 was collected as fees from scientists,
as compared to $1,907.75 last year. This decline is very largely due
to the high cost of transportation to the Isthmus, which keeps many
from coming. Despite the higher cost of food and other items, the
laboratory has not increased its per diem charge to scientists. Those
from institutions that sustain table subscriptions still receive a discount
of 25 percent.
The following institutions continued their support to the laboratory
through the payment of table subscriptions:
Smithsonianvinstitution ss.) boii a ye Ache alps 1 Sc lt Aa $500. 00
American’ Museum of Natural History =: 5.2222. +225L4-22221- 300. 00
Bastmany Wodak: Companys 2. teed 22 Se Peet ee yO 1, 000. 00
News VOrk ZOOlOpICAl NOCIGLY. = 2022 Ne eee ee 300. 00
University oe Cnicagos-22 2-6 iss. 54 52s steno Neat eke ae 300. 00
OhioistaterUniversivy® -s-2- 3 {ee rer ees Lee ee eee 300. 00
The Forest Products Laboratory, United States Department of
Agriculture, contributed $25.00 a month as service fees for facilities
furnished.
It is most gratifying to record donations from Dr. Eugene Eisen-
mann, Dr. Oliver P. Pearson, Mrs. Dorothy Edgerton, Miss Louise
Hunnewell, Mr. Frank W. Hunnewell, and the Botanical Society of
Washington.
The sum of $5,000 was made available by the Smithsonian Institu-
tion from appropriated funds, and of this amount $4,997.53 was used
for permanent improvements. The Institution also contributed $3,000
from its private funds, in addition to its table fees.
Respectfully submitted.
JAMES ZeTEK, Resident Manager.
Dr. ALEXANDER WETMORE,
Secretary, Smithsonian Institution.
APPENDIX 11
REPORT ON THE LIBRARY
Str: I have the honor to submit the following report on the activities
of the Smithsonian library for the fiscal year ended June 30, 1949:
All the continents and most of the countries of the world were
represented among the 57,671 publications received by the library
during the year. These books, pamphlets, and serials were pre-
dominantly scientific and technical in character and touched all the
special and related fields of interests of the Smithsonian Institution
and its branches. The International Exchange Service transmitted
5,082 of them, and the rest came by mail or by other means of delivery.
Acquisitions by purchase included 1,792 volumes, three collections
of pamphlets on special subjects, and subscriptions for 279 periodicals.
Gifts of 7,287 publications came from many different donors.
The library owes a lasting debt of gratitude to these friends of the
Institution, at home and abroad, for their generous contributions
of books and papers, many of which the library would not otherwise
have been able to acquire. Not yet statistically recorded in the
library is the important gift of the large Ferdinand Perret Research
Library of the Arts and their Affiliated Sciences, presented by Mr.
Perret to the National Collection of Fine Arts.
The library’s principal strength and the backbone of its usefulness
lies in the large collections of publications, chiefly serials, issued by
the research institutions, scientific societies, universities, academies,
museums, and observatories all over the world, which the Smith-
sonian Institution receives in exchange for its own publications.
These are the primary sources of the records of progress in science
and technology, in the arts and industries. Ready access to them is
indispensable to the work of the Institution. The larger number
of the 17,713 periodical entries recorded during the year were these
exchange publications, and many monographic works received in
exchange were separately cataloged. There were 338 new exchanges
arranged, and 7,008 volumes and parts needed to fill gaps in serial
sets, or for special purposes, were obtained in response to 726 special
requests made to the issuing agencies.
132
SECRETARY’S REPORT se
A grand total of 17,771 publications were sent to the Library of
Congress. Of these, 1,978 volumes and 4,582 periodical parts were
recorded as permanent additions to the great Smithsonian Deposit
there. The others were foreign and domestic dissertations, docu-
ments, and miscellaneous publications of little immediate importance
to the work of the Institution.
The Army Medical Library selected for transfer 859 publications
no longer needed here in the sectional library of the division of
medicine. Also sent to the Army Medical Library were 1,068
currently received medical dissertations and 1,927 other publications
on medical subjects. 33, 441. 68
——————— 6, 092, 775. 69
otalsinvestInentsts 42 s—= ese aaeeee 2 ae eee 9, 141, 069. 65
146 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
CASH BALANCES, RECEIPTS, AND DISBURSEMENTS DURING FISCAL
YEAR 19491
Cash balance/onphand! June’ 30) 194823225) 3 es See $563, 847. 37
Receipts other than Freer endowment:
Income frompnvestMmentsas sae aes ee eee $156, 219. 10
Giftsyandtcontnbutions#s= 452s ose eee 48, 143. 71
Salesvor pubheatians= 2! ooe = (eee) es ewe ewes 33, 281. 09
IMS Cel Ane OUS) eRe ER Pe eek Cis | 11, 566. 10
Total receipts other than Freer endowment-_-__-___------- 249, 210. 00
Receipts from Freer endowment:
Income fromanvestments== = 25a ae ee $282, 265. 48
Total receipts from Freer endowment-_-_-_-_-__--_--------- 282, 265. 48
TE GER eh es oe ee (Ee hie) Brie SR oa Oe 1, 095, 322. 85
Disbursements other than Freer endowment:
AGministrations= 22 22 oe e eee eo eae ees $43, 422. 75
PST ORGLORIS eee ee I a erent Ee 45, 618. 12
Bio) 75 2 PRE eS Se. dee ee Se ee ees 3, 977. 10
Buildings—care, repairs, alterations__._______- 136. 00
@ustodiantieesNetos: Gees 2 28 ee 3, 293. 15
Miscellaneous 4222s. ano ees eta Sas 3, S22. 07
Mesearchesse. 22. Soe Une eee ae eee 127, 412. 84
Smithsonian Retirement System_____________- 3, 608. 28
Purchases of securities (net) _.__..-.-__-_____- 4, 508. 63
Total disbursements other than Freer endowment_-_-_-_-_-__- 235, 799. 64
Disbursements from Freer endowment:
Salaries = 2 Areeet ee es eee ee $83, 480. 37
iRurchasestoncollectionsss.2 = — eee 125, 050. 00
Custodian fees etohaisa 2 2-2 eee 10, 858. 00
Miscellaneoud@oe ss soe ee cee oe a nee eee 26, 594. 80
Purchases of securities (net) __._._.._..________- 80, 631. 18
Total disbursements from Freer endowment__....__.-_--- 326, 614. 35
Investment of current funds in U. 8. Bonds________-__-__-.___- 2, 578. 13
fhotaldisbursements= = So o= 3226-2 sea ee eee 564, 992. 12
Cashibalante June30, 1049incteeabso cuss. sees ee eee 530, 330. 73
TOtalG: Ses oa55 he eee ee EE Fa en ee 1, 095, 322. 85
1 This statement does not include Government appropriations under the administrative charge of the
Institution.
SECRETARY’S REPORT
ASSETS
Cash:
United States Treasury cur-
Tent Account ae = see ae $360, 201. 95
In banks and on hand_----- 170, 128. 78
530, 330. 73
Less uninvested endowment
fundshe 2s ea we eee 89, 826. 17
ED OE CE
Travel and other advances___-_----------- 11, 585. 42
Cash invested (U.S. Treasury notes) ------ 502, 815. 37
Investments—at book value:
Endowment funds:
Freer Gallery of Art:
Stocks and bonds__ $6, 059, 334. 01
Uninvested capital_ 33, 441. 68
——_—_—__————_ 6, 092, 775. 69
Investments at book value other than Freer:
Stocks and bonds_-__--- $1, 929, 118. 64
Real estate and mort-
gage notes=—— 2 62, 790. 83
Uninvested capital____-_ 56, 384 49
Special deposit in U. S.
Treasury. Interest at
6G; percent==ss====—"— 1, 000, 000. 00
—— 3, 048, 293. 96
147
$954, 905. 35
9, 141, 069. 65
10, 095, 975. 00
UNEXPENDED FUNDS AND ENDOWMENTS
Unexpended funds:
Income from Freer Gallery of Art endowment-_------------
Income from other endowments:
Restricted == sees oa es ae ee $187, 425. 28
Goneral os besa ts ee Ok ao ee ee 85, 599. 74
Giltstindvpranite: 2. 5eie Sees io es ls ee
Endowment funds:
RreersGallery Of Att. o22s-. 2526s ee aon $6, 092, 775. 69
Other:
Restrictedes sss) ee —— $1, 345, 776. 78
General 2 oe eek ee 1, 702, 517. 18
3, 048, 293. 96
393, 411. 62
273, 025. 02
288, 468. 71
954, 905. 35
9, 141, 069. 65
10, 095, 975. 00
148 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The practice of maintaining savings accounts in several of the
Washington banks and trust companies has been continued during
the past year, and interest on these deposits amounted to $824.74.
In many instances, deposits are made in banks for convenience in
collection of checks, etc., and later such funds are withdrawn and
deposited in the United States Treasury.
Disbursement of funds is made by check signed by the Secretary
of the Institution and drawn on the United States Treasury.
The foregoing report relates only to the private funds of the
Institution.
The Institution gratefully acknowledges gifts from the following:
American Philosophical Society, for Iroquois research.
W. W. Corcoran, for B. F. Starr.
Eickemeyer Estate, for preservation, etc., of Rudolph Eickemeyer photographic
collection.
KE. P. Killip, for use of Department of Botany.
National Academy of Sciences, for study of flora of Okinawa.
National Geographic Society, expedition to western Panamé,
Research Corporation.
John A. Roebling, additional contribution for researches in radiation.
The following appropriations were made by Congress for the
Government bureaus under the administrative charge of the Smith-
sonian Institution for the fiscal year 1949:
Salariesiandvexpensessueseree i se eee oe Sao ae eee eee eae $2, 259, 000. 00
INStIOnalY SOOO sICO Ean K Shae sie eee SE Fs See 528, 848. 00
In addition, funds were transferred from other Departments of the
Government for expenditure under direction of the Smithsonian
Institution as follows:
Cooperation with the American Republics (transfer from the State
Weparvment). 28 es eee ewe nee eee ee meee eee ee eee $97, 960. 00
Working Fund, transferred from the National Park Service,
Interior Department, for archeological investigations in River
Basins throughout the United States_.._.........--.-.------- $118, 500. 00
The Institution also administers a trust fund for partial support
of the Canal Zone Biological Area, located on Barro Colorado Island
in the Canal Zone.
The report of the audit of the Smithsonian private funds follows:
SEPTEMBER 14, 1949.
To THE Boarp or REGENTs,
SMITHSONIAN INSTITUTION,
Washington 25, D. C.
We have examined the accounts of the Smithsonian Institution relative to its
private endowment funds and gifts (but excluding the National Gallery of Art
and other departments, bureaus, or operations administered by the Institution
under Federal appropriations) for the year ended June 30, 1949. Our examina-
tion was made in accordance with generally accepted auditing standards, and
SECRETARY’S REPORT 149
accordingly included such tests of the accounting records and such other auditing
procedures as we considered necessary in the circumstances.
The Institution maintains its accounts on a cash basis and does not accrue
income and expenses. Land, buildings, furniture, equipment, works of art,
living and other specimens and certain sundry property are not included in the
accounts of the Institution.
In our opinion, the accompanying financial statements present fairly the
position of the private funds and the cash and investments thereof of the Smith-
sonian Institution at June 30, 1949 (excluding the National Gallery of Art and
other departments, bureaus or operations administered by the Institution under
Federal appropriations) and the cash receipts and disbursements for the year
then ended, in conformity with generally accepted accounting principles applied
on a basis consistent with that of the preceding year.
Prat, Marwick, MircHe.y & Co.
Respectfully submitted.
Rozert V. FLEMING,
VANNEVAR Bus,
CLARENCE CANNON,
Executive Commiitee.
ae oxi oe ah rf rs be ay on me webrope sy a
: - yah noes & Fa i" eater ee eT
be a “ose Yon aor Twit 7 ans Se Oe | nd: eitedes " t ay ufstieet rt
=e ei 9) alSerds ede ytups ater tical Firstly: ae
WAY at idea: i Witgo easteiax nic més, Gwe lesieviloe ga seth ut
= fas hea eiled S pene ous aul 9 ae i atc =)
a
=
sad
52
| ll
5
I
=
e<
Bei
= ath VOUS droit) aluontaineg Lejanert,, gies tachment sill sipping 3 u
- ory es ah ) >, 7 =f
. 13} uo) To Toes itd dy MVE2 aS i r hile ifs 4 ty. Ds Ur Pitul of# ving 602 ce: ‘cn ni
7 Tee ow te ; J foros Ath, gnivnitss 9) Ube t e ny saint peliad iedliaaial aig
wahAy (eR oaY i f Perkigiuiethe Siciliano i o> oo eae
x z U: 8 ¢c o
ee) mee: <—- er Pi) e > —
Ficure 3.—Upper, our view of some spiral nebulae. The arrows indicate veloc-
ities. Note that spiral B, which is twice as far from us as spiral A, is receding
twice as fast. OC, three times as far, is receding three times as fast, and so on.
Lower, the view of an observer on spiral B, considering himself to be at rest.
It is the principle of relativity that he has just as much right as we do to con-
sider himself at rest. He gets the same view as we do; all the spirals are
receding from B with velocity proportional] to distance.
Tracing the motions back in time (there is no evidence that the
spirals are accelerating or decelerating) shows that all the spiral
nebulae would have been near our galaxy between 2 and 3 billion
years ago. The coincidence of this with the age of the earth and the
age of the meteorites is too marked to need further comment—the
whole universe seems to have started with a bang about 3 billion years
ago!
‘ THE BEGINNING OF TIME
This curious evidence that the spiral nebulae were all close to—if
not entangled with—our galaxy 8 billion years ago, means that the
formation of the solar system at that time probably took place under
conditions somewhat different from those of today. To be sure of the
reasoning, we must examine the conditions of 3 billion years ago more
carefully; it was this reexamination which led, in 1945, to the most
bizarre suggestion of all in this field already rich in speculation. It
was put forward by the English biologist, J. B. S. Haldane, and is
based on a new theory—or philosophy—of relativity proposed in 1932
by the English mathematician, E. A. Milne. First we shall speak of
Milne and his brand of relativity.
To make the reasoning clear we must start with Einstein’s earlier
relativity theory which links space and time in such a way that if
one observer is moving at constant velocity past another his
measurements of distances and time intervals will differ from those
172 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
of the first observer, although the relation of time and distance is
such that they both observe the same laws of physics. LHinstein
formulated his relativity on the philosophy that it is simply impos-
sible to tell which observer is ‘‘at rest.’”? Complicated as it sounds,
this scheme has been developed to form a logically complete theory
in terms of mathematical transformations. Milne extended the
established principles of relativity in his “cosmological principle,”’
which is, in effect, an assumption that the view of the whole universe
from one spiral nebula must be the same as the view from any other.
Moreover, he has redefined distance measurements in terms of the
travel time of light signals, as in radar ranging, thus reducing both
time and distance measurements to readings of clocks, in principle.
¢ e e ‘ ¢ . ’ ¢ eo e ¢ e oe
° e e e e o e ¢ 7 e a e e
e o . . ¢ e e a e ¢ eo ¢ e
e ‘ ¢ ¢ o ° e ’ ° eo eo e ¢
Us
o e e ° 2 , e ¢ ‘ eo 6 oe eo
Figure 4.—Milne’s picture of the universe. If all measurements are made in
atomic time, on Milne’s theory, the universe started expanding from a point
3 billion atomic years ago. As we see it now the spiral nebulae shown in the
left diagram are all moving away from us and (if we could see far enough)
would be much more numerous near the “edge.”” At this edge the velocity of
recession is equal to the velocity of light, so we can never hope to see the edge
itself. On the other hand, if clock time is used for all our measurements, the
universe is static and the spiral nebulae, as shown on the right above, are
uniformly distributed on to infinity. The more distant nebulae are redder be-
cause we see them as they were many years ago with “‘slow” atoms. The
“edge’’ of this picture comes when this reddening gets so extreme that galaxies
are no longer visible.
Milne then raises the disturbing question: How are we sure that
our clocks are reading constant intervals of time? In fact, the slow-
ing down of the earth’s rotation (which is normally our ‘master
clock’’) has been measured as one-thousandth of a second per century
by comparison with the planets, and we have no philosophically
sound assurance that the planets keep “‘perfect time.”
The cosmological principle leads mathematically to two kinds of
time, one of which is speeding up relative to the other. Milne has
shown that pendulum clocks, the earth, and the planets keep ‘‘dy-
ORIGIN OF THE EARTH—PAGE tzZ3
namic’”’ or clock time, while vibrating atoms and radioactive decay
have constant period only in “kinematic”’ or atomic time. There is
no philosophical reason for choosing one kind as the “correct” time;
if we used a pendulum clock to time atoms we would find, after a very
long interval, that the atoms are gradually speeding up in their
vibration, if we used an atomic clock we would similarly find that the
planets are slowing down in their orbits.
If this is correct—and no one has yet proved it otherwise—the
age of the earth is 3 billion atomic years as determined from radioactive
decay, but it is many more clock years, since in the past the clock
year was shorter than the atomic year. (They are equal at present—
by definition.)
The coincidence between the age of the earth and the time of reces-
sion of the spiral nebulae Milne explains as a result of the difference
in these two kinds of time. Since the light we observe from a spiral
100 million light-years away left there 100 million years ago, we are
seeing the atoms there ticking off the units of atomic time in use 100
million years ago. Compared to our present atoms, these early
atoms ran slow; as a result, the light they emitted is redder than the
light emitted now by similar atoms on the earth.
From this effect and his cosmological principle, Milne calculates
that in the past infinite number of clock years there were 3 billion
atomic years. The origin of the earth, and the time when all the
spirals were close to our galaxy, both of them 3 billion atomic years
ago, therefore occurred at the beginning of time (since one could hardly
expect more than infinite time on the clock scale).
Now for Haldane’s suggestion, which he calls ‘A Quantum Theory
of the Origin of the Solar System.” It is based, as its name implies,
on the well-established quantum theory of radiation, and on a mathe-
matical result of Milne’s theory: that the universe, as measured in
atomic time, has expanded with the velocity of light, starting from a
point of zero radius 3 billion atomic years ago.
Since the universe started from zero radius, Haldane was able to
pick an early enough instant, just a fraction of a second after the start
of atomic time, when the whole universe was but a fraction of an inch
in diameter—much smaller than the wave length of visible light—
smaller, by far, than the wave length of X-rays or gamma rays.
(These fractions are too small to write out easily; the first requires
72 zeros after the decimal point, the second, 62!) ‘The wave lengths
of radiation in existence in this small universe could scarcely have
been bigger than the universe itself, Haldane reasoned, therefore the
only radiation in existence was of these incredibly short wave lengths.
But the basic principle of the quantum theory is that radiant energy
comes only in packets, or ‘quanta,’ inversely proportional to the
wave length in size. So, at this early instant all radiation was in
174 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
giant quanta of very small waves. And the energy of one of these
giant quanta can easily be calculated as sufficient to knock one or
more planets out of the sun. The even smaller waves at a somewhat
earlier instant would have been in quanta with sufficient energy to
tear apart stars, and even earlier, to tear apart the galaxies from some
primordial globe of matter.
The details of this remarkable suggestion have been carried no
farther, but Haldane’s investigation points up one important general
fact: whether or not Milne’s new relativity is accepted, conditions
at the time of the origin of the solar system were probably consider-
ably different from those today. If Milne’s cosmology is accepted,
the relationship between radiation and matter was most radically
different. It may seem that this iast and most fantastic speculation—
which can neither be completely explained nor fully evaluated here—
contradicts our former conclusion that the solar system was formed
from a rotating nebula of gas and dust. However, the condensation
of the planets and the distribution of angular momentum (which
have been so difficult to explain in all previous theories) may follow
from further mathematical investigation of the first second of atomic
time. In fact, if the details can be worked out rigorously, Haldane’s
suggestion may lead to confirmation of Milne’s cosmology, which is
as yet lacking.
In an echo of the introductory remarks it scarcely needs to be
emphasized that we have no complete theory of the origin of the
earth. The reader may be impressed with the diverse investigations
involved and with the promise of the latest speculations; or he may
notice the infinite regression implicit in any question of origins: if
the planets were formed from dust or planetesimals, whence came the
dust or planetesimals? if the dust and planetesimals came from a
primordial nebula, whence came the primordial nebula? if the primor-
dial nebula was formed by the absorption of a giant quantum by a
fragment of matter, whence came the original matter and radiation
in the universe? and so on, ad infinitum (clock time).
THE 200-INCH HALE TELESCOPE AND SOME
PROBLEMS IT MAY SOLVE!
By Epwin Hussite
Mount Wilson Observatory
[With 10 plates]
In 1609 Galileo turned his telescopes toward the sky. His favor-
ite—it was the fifth, finished within 6 months of the first trial—was
about 5 feet long and had a lens about 2 inches in diameter. It
magnified nearly 30 times and its light-gathering power was equal to
about 80 human eyes. He called it ‘Old Discoverer,” and with it he
saw mountains on the moon, phases of Venus, four moons of Jupiter,
and stars innumerable beyond the limit of the unaided eye.
It was then that the explorations of space began—the explorations
that have swept outward in wave after wave as telescopes developed,
until in our time we study a region of space so vast that it may be a
fair sample of the universe itself. Today there is nearing completion
a new telescope, far more powerful than any previously made, and it
is proper to consider its significance both as an engineering achieve-
ment, and as an instrument for further explorations. With this end
in view, I propose to discuss briefly the development of telescopes in
general, the 200-inch in particular, and some of the problems it may
help us to solve.
Galileo’s optic tubes with single-lens objectives grew rapidly into
telescopes from 20 to 25 feet long with lenses 2 to 3 inches in diameter.
There the development stopped, for practical purposes, because of
the engineering difficulties with still longer tubes.
The longer focal lengths were considered desirable in order to over-
come color difficulties. With a single lens, each different color was
brought to a focus at a different distance from the lens. Hence the
image, when focused for any particular color, was blurred by the out-
of-focus images in other colors. The long telescopes represented an
attempt to spread out the images of different colors over so long a
distance that one color could be focused with minimum interference
1 Alexander F. Morrison lecture, delivered in Pasadena, Calif., April 8, 1947. Reprinted by permission,
with slight alterations, from Publications of the Astronomical Society of the Pacific, vol. 59, No. 349, August
1947.
175
176 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
from the others. Telescopes 100, 150, and even 200 feet long were
actually constructed, with lenses from 3 to 6 inches in diameter.
These monstrous instruments, however, were too unwieldy for use,
and the real work during the first century and a half after Galileo’s
time was done with the smaller telescopes.
Finally, in the middle of the eighteenth century, the color problem
was solved by replacing the single-lens objective with a compound
objective, each of whose separate components, made of different kinds
of glass, canceled out most of the color effects of the other. These
color-free (achromatic) lenses gave much better images and permitted
the use of relatively short tubes for a lens of a given diameter. Tele-
scopes immediately entered a new period of growth which culminated
in the 40-inch lens, with a focal length of 63 feet, at the Yerkes
Observatory in Wisconsin. The 40-inch was finished in 1892, and
since that time developments have concerned lenses for special pur-
poses rather than for greater light-gathering power. For technical
reasons, it seems unlikely that larger lenses will be made in the
foreseeable future.
This greatest of all lenses had been ordered originally by a group of
enthusiasts here in southern California in connection with a plan for
a ‘University System.” The project did not fully materialize, and
the unfinished telescope was bought and completed by the University
of Chicago.
The very large telescopes of recent times, in which light-gathering
power is the most important consideration, are all reflectors, not
refractors. The light is funneled to a focus, not by refraction through
a convex lens but by reflection from a concave mirror. ‘These tele-
scopes are free from color effects because all colors are reflected in
the same way.
The first reflector was made by Isaac Newton in 1672, in a deliberate
effort to avoid the color troubles of single-lens refractors. His first
model had a burnished metal mirror, about an inch in diameter,
figured to a concave spherical surface, and mounted at the bottom
of a tube about 6 inches long. The image, which would lie in the
middle of the upper end of the tube, was thrown to the side by a small
plane mirror set at 45° to the axis, just below the focus. Newton
presented the toy to the Royal Society, where it may still be seen,
sitting on a volume of his famous Principia.
Although Newton’s reflector avoided the color problem, it suffered
another defect, known as spherical aberration, arising from the
spherical surface of the main mirror. It was not until 50 years later
when Hadley, in 1722, found a method of parabolizing concave mirrors,
that the development of reflectors finally got under way. About 90
years ago metal mirrors were replaced by glass, silvered on the front
surfaces. In our time aluminum has been substituted for silver,
200-INCH HALE TELESCOPE—HUBBLE ZAC
low-expansion glasses have been developed, methods of parabolizing
have been perfected, and engineering problems of constructing
telescopes have been solved as they arose.
The 40-inch refractor was installed at Yerkes under the direction
of George E. Hale. He clearly saw that, regardless of the success
of this telescope, the quest for still greater light-gathering power
depended upon mirrors rather than lenses. Refractors were pref-
erable for certain types of work (including, for instance, visual
resolution of double stars, precise measurement of position, wide-
angle photography, ectc.), but for light-gathering power, with all that
it implies, the future lay with the reflector. Because the reflections
are from the front surfaces, transparency and absolute homogeneity
of the glass are not demanded; the mirror may be supported from the
back and sides, instead of from the rims alone as in the case of lenses,
and, of course, there are no color effects.
Hale took the lead in America in encouraging the development of
large reflectors. A 24-inch of unusual perfection was made by G. W.
Ritchey and installed at Yerkes. It proved so successful that plans
for a 60-inch were immediately set in motion. When Hale left
Yerkes to establish the Mount Wilson Observatory, he was able to
transfer Ritchey and the unfinished 60-inch mirror to Pasadena, where
the telescope was completed in 1908. The work of the 60-inch on
Mount Wilson so fully justified the faith in larger reflectors that plans
for a new one were immediately made, this time for a 100-inch mirror.
This reflector, completed during the first World War, marked an
important epoch in the history of astronomy. It is still the greatest
telescope in operation. Four large reflectors with mirrors from 60
to 84 inches have since been completed (two in Canada and two in the
United States), and others, including a 120-inch for Lick Observatory,
are in process of planning or construction.
The 100-inch opened up new fields of investigation of the very first
importance, and furnished glimpses of even richer fields beyond. If
more light-gathering power were available, these more distant fields
could be explored. In the face of this challenge the possibility of
larger telescopes was the favorite topic of conversation among astron-
omers at Mount Wilson, and presumably at other places as well.
We talked of 200 inches, or 300, and even dreamed of still more light.
One of the group, F. G. Pease, drew tentative designs for a 300-inch,
and demonstrated that the engineering features were not impossible.
Again Hale took the lead. Through his efforts funds were secured
in 1928 in the form of a gift from the International Education Board
to the California Institute of Technology for the establishment of an
astronomical observatory and laboratory. An Observatory Council,
with Hale as chairman, and with the greatest experts in the country
as advisers, administered the details of the project. When Hale
178 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
dropped out, Max Mason took over the chairmanship. It was decided
that a 200-inch reflector was as bold a step beyond the 100-inch as
could be justified in view of the unknown problems, both optical and
engineering, that might be encountered. Laboratories and shops
were erected on the California Institute campus, a site for the ob
servatory was found on Mount Palomar, a disk of pyrex glass was
achieved by the Corning Glass Company, and the project proceeded
steadily until it was interrupted by the war. Work was resumed soon
after VJ-day, and the telescope has since been completed.
The proper fields for the 200-inch are determined primarily by its
immense light-gathering power. Because adequate consideration of
all the possible applications would require more time than is now
available, I propose to limit the following discussion to three typical
problems. These problems are, first, the existence of canals on Mars;
second, the relative abundance of the chemical elements in stars; and,
third, the large-scale structure of the universe. Each problem
represents a particular aspect of light-gathering power, namely,
resolution, dispersion, and depth penetration.
As a brief introduction, let me comment on the telescope itself.
The mirror intercepts a beam of light 200 inches or 17% feet in
diameter—in other words, it gathers as much light as a million human
eyes, or four 100-inch reflectors. It funnels this light to an image at
the primary focus, 55% feet in front of the mirror. There an image
of the sky is formed such as you may see on the ground glass of a
camera. This image may be examined visually with a microscope,
recorded on a photographic plate, analyzed with a spectrograph, or
studied by other techniques. Actually, most of the work will consist
of direct photography or spectrum analysis. By using long time
exposures, it is possible to photograph stars or nebulae several times
fainter than can be seen in the eyepiece. For this reason, the 200-
inch is best described as a huge camera.
Now let us consider some typical problems for the 200-inch. I shall
start with a problem concerning a member of the solar system. The
telescope will not be turned on the sun because of temperature effects—
in some ways it would act as a burning glass. Nor does it offer any
unique advantages for the study of the moon. In that field it will
serve merely to improve data of a kind that can be got nearly as well
with several other telescopes. In the field of planetary photography,
however, the opportunities are unique because, for the first time, it
may be possible to photograph all that the eye can see with a telescope
of moderate size. An immediate application is to the highly contro-
versial question of canals on Mars.
The canals are described as very fine, dark lines running along great
circles, sometimes doubled, and often converging or crossing at spots
called “‘oases.”’ Such fine, hairlike patterns, superposed on the back-
200-INCH HALE TELESCOPE—HUBBLE 179
eround of large, well-known markings, have been recorded by various
trained visual observers, using telescopes of all sizes from 6 inches up-
ward, during the whole of the past 70 years since Schiaparelli first
reported them. ‘The canal systems, if real, would almost necessarily
imply the existence now or in the past of intelligent beings on Mars.
But other trained observers, again using telescopes of all sizes,
report no trace of canals. KE. E. Barnard, perhaps the greatest of the
American visual observers, studied the planet over many years with
the then largest telescopes in the world, including the 36-inch refractor
at Lick, the 40-inch at Yerkes, and the 60-inch reflector at Mount
Wilson, and, although he saw an immense amount of detail, he found
no canals.
The two groups of observers flatly contradict each other, and since
the observations are personal impressions neither group can demon-
strate the validity of its assertions. Evidently the controversy must
be resolved by photography. Once photographs are available on which
the canals should appear if they are real features of the planet, the
question will be settled beyond reasonable doubt. The test has not
been possible as yet because existing equipment, although closely ap-
proaching the required standards, does not fulfill them. The 200-
inch, however, should meet all the necessary conditions and settle the
question.
The problem is as follows: Mars is a small object. The image at
the primary focus of the 100-inch is less than 5 inch at the most
favorable oppositions, and less than % inch at the long Cassegrain
focus. In order to get an image large enough to serve as a critical
test of fine markings (i. e., to make the resolution of the photographic
plate comparable with the optical resolution of the telescope) it is
necessary to use an enlarging lens. Thus the total light collected by
the telescope is spread over a much larger, and correspondingly fainter,
image. Furthermore, because of the atmosphere on Mars, it is neces-
sary to photograph the surface markings through deep orange or red
filters, still further reducing the effective brightness of the image.
The reduction is so great that photographs with existing telescopes
require time exposures instead of snapshots. The exposures may be
only a second (or even a fraction of a second, with the 100-inch) but
they are long enough to permit the dancing or shimmering of the
image to smear out the finest detail.
You doubtless realize that a telescope magnifies the twinkling of
stars along with everything else. Critical observations are restricted
to periods of maximum steadiness (good seeing, as it is called), and
even then, the shimmer is appreciable under high magnification. The
eye can “hold”? an image under these conditions, but photography
with time exposures is helpless. The shimmer smears out the fine
details. It is for this reason that, in the case of fine markings such
180 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
as the canals on Mars are said to be, the eye can see more than the
photographic plate can record.
However, the 200-inch will collect so much light that, for the first
time, it will be possible to photograph enlarged images through filters
with snapshots. These exposures will be short enough to catch a
dancing image at the end of a flicker—when it is momentarily at rest
as it reverses direction. If many exposures are snapped on a movie
film, a certain percentage of them may be expected to record what the
eye can see (at least with telescopes of moderate sizes).
There is much more to the story, but it is too technical for the
present discussion. But it can be said with some confidence that the
200-inch may settle the long-standing controversy concerning the
canals on Mars.
The second problem I have selected for discussion involves spectrum
analysis. You know, of course, that light reaches us as a jumble of
waves of all different wave lengths, each representing a different color.
It is possible, with prisms or gratings, to spread the colors out into
an ordered sequence or spectrum, running from the long waves of thered
to the short waves of the violet, and beyond in either direction. Such
spectra of stars and nebulae show phenomena of profound significance
at certain particular wave lengths. Spectrum analysis involves the
isolation and study of these particular regions.
Your radio offers an analogy. With the tuning dial you run along
the spectrum of radio waves and isolate a particular wave length in
order to hear a particular station which is broadcasting on that wave
length.
If there were no tuning device, and you heard all programs at once
with a nonselective receiver, the result would be bedlam. The step
from such a nightmare to the clear reception of messages from indi-
vidual stations suggests the nature of the step in astronomy from the
study of integrated light to spectrum analysis.
Light from stars and nebulae originates in atoms. There are as
many kinds of atoms as there are chemical elements, and the atoms
may have various stable states. Each stable state of each kind of
atom represents a set of broadcasting stations, sending messages
concerning the nature of the atoms and the physical conditions under
which they exist. By tuning in and reading these messages, it is pos-
sible to identify chemical elements, to determine temperatures, pres-
sures, and other physical attributes, and even to measure motion in
the line of sight (radial velocity).
But in order to read the messages clearly it is necessary to achieve
precise tuning—that is, to spread out the spectrum on the maximum
possible scale. It is here that the great light-gathering power of the
200-inch offers new possibilities.
200-INCH HALE TELESCOPE—HUBBLE 181
The length over which a spectrum can be spread, and still remain
bright enough to be photographed, depends upon the brightness of
the object. The sun has been spread out over a spectrum about 50
feet long from red to violet; the brightest stars, over about 3 feet,
and the faintest naked-eye stars over about 1 foot. The shortest
spectra giving useful information are about one-tenth of an inch long,
and have been obtained from stars and nebulae about a hundred
thousand times fainter than the faintest naked-eye stars. With the
200-inch, all the stellar and nebular spectra can be lengthened about
four times, and consequently the analysis can be carried out much
more precisely than was hitherto possible.
One new field, now faintly glimpsed, can be explored rather fully.
The important data are the relative abundances of the different chem-
ical elements in different kinds of stars. These data are derived from
the comparative study of the different stations (or lines) due to differ-
ent chemical elements in a spectrum, and require the longest practical
spectra (the highest possible dispersion) for adequate analysis.
There is reason to believe that more than 99 percent of the atoms
in the universe are hydrogen. Even by weight, hydrogen, with the
simplest and lightest of all atoms, probably contributes a large frac-
tion of the total matter in the universe. There are insistent sugges-
tions that the relative abundance of hydrogen varies considerably
from star to star. There is also some reason to suppose that the rela-
tive abundance of other elements does not vary widely in the stars,
although the physical conditions of the stars do vary widely (from
giants to dwarfs, from hot blue stars to cool red stars). The supposi-
tion rests mainly on negative evidence and requires further study
with powerful instruments.
It is believed that the 200-inch alone can adequately explore this
field, now dimly outlined with existing telescopes. What is now sug-
gested by analysis of three or four of the very brightest stars can be
critically tested in these objects, and the study can be extended in a
comparable way over a large sample collection of stars in general.
We cannot predict the final results of the exploration. They may
represent the next major chapter in the development of our knowledge
of the universe, or they may prove to be relatively trivial. But the
unexplored field looms as a challenge, and the challenge will be met.
The data are immensely important because they bear directly on
two very fundamental problems, namely, the source of stellar energy
and the origin of chemical elements.
Geologists, studying the history of the earth’s surface, assure us
that the sun has been pouring out energy at a fairly constant rate
over the last several hundred million years at least. Possible sources
for the unfailing supply were not only unknown but were unimagined,
182 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
until the modern science of nuclear physics was developed. Now ex-
planations may be sought over a wide range of nuclear reactions giv-
ing various lifetimes to stars up to the limit set by Einstein’s famous
formula,
energy =mass X (velocity of light)?
For instance, if the whole of the sun’s mass were transformed directly
into energy, the sun could radiate at the present rate for a million
million years. But, if nuclear reactions supplied the energy, the
possible lifetime (with the present rate of radiation) would be reduced
according to the particular reaction involved.
The nuclear reactions, in general, produce the transmutation of
elements—the old dream of the alchemists. The most plausible of
the current theories concerning the source of the sun’s energy, pro-
posed by H. Bethe, is based on the carbon cycle in which, because of
the presence of carbon at temperatures found in the sun, hydrogen
nuclei may combine to form helium, releasing energy in the process.
One test of the theory is furnished by a comparison of the relative
abundance of the different isotopes of carbon actually observed in the
sun with the relative abundances involved in the carbon cycle.
In a vaguely analogous way, it is possible to speculate on the build-
ing up of all elements from the primitive hydrogen atoms, and these
speculations may be guided by the observed relative abundances of
elements in stars of widely different physical characteristics.
Thus the data on abundances, derived from large-scale spectra,
bear directly on all theories concerning the source of stellar energy,
the origin of chemical elements, the past history of the universe, and
its future evolution.
The third unique field of investigation for the 200-inch is cos-
mology—the structure and behavior of the universe as a whole. As-
tronomers hope that the observable region of space—the region that
can be observed with telescopes—is a fair sample of the universe, and
they attempt to infer the nature of the universe from the observed
characteristics of the sample. The 200-inch, because of its great
light-gathering power, should penetrate into space about twice as far
as the 100-inch, and consequently will permit us to explore a volume
of space about eight times that now available. The probability that
the observable region may be a fair sample of the universe will thus
be greatly increased.
It was the 100-inch that opened this new field and prepared the
way for the new telescope. The picture developed rather suddenly
during the 1920’s. The sun with its family of planets seems isolated
and lonely in space, but we know that it is merely one of the stars—
one of several thousand million stars which, together, form the stellar
system. This system is a swarm of stars which drifts through space
Smithsonian Report, 1949.—Hubble PLATE 1
DOME OF THE 200-INCH TELESCOPE ON PALOMAR MOUNTAIN
Smithsonian Report, 1949.—Hubble PEATE 2
200-INCH HALE TELESCOPE
Meeting of the American Astronomical Society and the Astronomical Society
of the Pacific on July 1, 1948.
Smithsonian Report, 1949.—Hubble PLATE 3
MARS
Upper: Photograph with 100-inch reflector, September 2, 1924.
Lower: Drawing by Pettit with 20-inch reflector, July 12, 1939. These typical
pictures illustrate the fact that as vet photography does not furnish an objective
test of the existence of “canals” on Mars.
Smithsonian Report, 1949.—Hubble
EXTRAGALACTIC NEBULAE
These nebulae are examples of the stellar systems which serve as landmarks in
the exploration of the universe. The group above (NGC 3185, 3187, 3190,
3193) is at a distance of about 8 million light-years and appears to be receding
from us at the rate of about 850 miles per second.
Smithsonian Report, 1949.—Hubble PLATE 5
NGC 2261
The first photograph made with the 200-inch Hale telescope.
ZG VWaHYV GSLOATAS NI NOIS3Y
9 ALVI1d 2[999H— 6r6| ‘340dey UeluOsy{IUIG
Smithsonian Rerort, | PLATE 7
MESSIER 87 (NGC 4486)
Smithsonian Report, 1949.—Hubble PLATE 8
NGC 5204
Smithsonian Report, 1949.—Hubble PLATE 9
NGC 3359
Smithsonian Report, 1949.—Hubble PEATE 0
MESSIER 3 (NGC 5272)
200-INCH HALE TELESCOPE—HUBBLE 183
as a swarm of bees drifts through the air. From our position some-
where within the system, we look out through the swarm of stars,
past the boundaries, into the universe beyond.
Those outer regions are empty for the most part—vast stretches
of empty space. But here and there, scattered at immense intervals,
we find other stellar systems comparable with our own. They are so
remote that individual stars can be seen only in a few of the nearest
systems. In general they appear as faint patches of light, resembling
tiny clouds, and have long been called by the Latin word for clouds—
that is, ‘‘nebulae.”’
We now know that these nebulae are huge stellar systems averaging
about a hundred million times as bright as the sun. They are the
true inhabitants of space—vast beacons that serve as landmarks for
the exploration of the universe. We see a few that appear large and
bright. These are the nearer nebulae. Then we find them smaller
and fainter in constantly increasing numbers, and we know we are
reaching out into space farther and ever farther, until, with the faintest
nebulae that can be detected with the largest telescope, we have
reached the frontiers of the observable region.
This region has been explored with the 100-inch out to distances so
remote that light, speeding at 186,000 miles per second, requires 500
million years to make the journey. Thus the observable region at
present is a sphere, centered on the observer, with a radius of about
500 million light-years. Throughout this sphere about a hundred
million nebulae are scattered, each a stellar system comparable to
our own system of the Milky Way.
The study of this observable region as a sample of the universe has
led to the recognition of two large-scale features. The first feature
is homogeneity. The nebulae are scattered singly, in groups, and
even in great clusters, but when very large volumes of space are com-
pared, their contents are found to be quite similar. On the grand
scale, the observable region appears to be very much the same, in all
directions and at all distances.
The second characteristic is the fact that light waves from distant
nebulae seem to grow longer in proportion to the distance they have
traveled. It is as though the stations on your radio dial were all
shifted toward the longer wave lengths in proportion to the distances
of the stations. In the nebular spectra the stations (or lines) are
shifted toward the red, and these red-shifts vary directly with dis-
tance—an approximately linear relation.
The red-shifts are most easily interpreted as evidence of motion in
the line of sight away from the earth—as evidence that the nebulae
in all directions are rushing away from us, and that the farther away
they are, the faster they are receding. This interpretation lends itself
directly to theories of an expanding universe. The interpretation is
866591—50——13
184 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
not universally accepted, but even the most cautious of us admit that
red-shifts are evidence either of an expanding universe or of some
hitherto unknown principle of nature.
The two observed characteristics of the observable region, namely,
the approximately uniform distribution and the approximately linear
law of red-shifts, must be satisfied by any theory of the universe.
They are the only observational results on the grand scale that can
be used as tests. They serve to eliminate many theories formerly
developed on insufficient data, but several modern theories survive
the tests. These latter theories all permit the observed features in
a limited region near the observer but they predict that departures
from the simple approximate laws of distribution and of red-shifts
will be found when the measures are extended to greater distances.
These departures differ from theory to theory, and, if the measures
can be extended to the necessary distance, will distinguish the correct
theory from the false.
Thus the most important observational problems in cosmology may
be described as the small, second-order effects of great distances.
The nebulae appear to be distributed in a roughly uniform manner
and the red-shifts appear to be roughly proportional to distance, out
to the limits of the 100-inch. The next step is to determine these
features more precisely over the limited range of the 100-inch and
approximately out to far greater distances.
Attempts have been made to attain the necessary precision with
the 100-inch, and the results appear to be significant. If they are
valid, it seems likely that red-shifts may not be due to an expanding
universe, and much of the current speculation on the structure of the
universe may require re-examination. The significant data, however,
were necessarily obtained at the very limit of a single instrument,
and there were no possible means of checking the results by inde-
pendent evidence. Therefore the results must be accepted for the
present as suggestive rather than definitive.
The problem is essentially one for the 200-inch. This new telescope
will penetrate into space out to a thousand million light-years, and
the second-order effects of great distance will be so conspicuous that
they cannot be missed.
As a particular and final example, let me mention the effects of
increasing red-shifts on apparent brightness. It is well known that
a rapidly receding light appears fainter than a similar, but stationary,
light at the same momentary distance. The reason is that the stream
of light-quanta from the moving light is thinned out by the recession
so that fewer quanta per second reach the observer. Since brightness
is measured by the rate of arrival of quanta, the receding light appears
abnormally faint.
200-INCH HALE TELESCOPE—HUBBLE 185
Actually the dimming factor (the reduction of apparent brightness)
is a simple fraction represented by velocity of recession divided by
the velocity of light. Recession at one one-hundredth the velocity
of light reduces the apparent brightness by 1 percent; at one-tenth
the velocity of light, by 10 percent, and so on. Thus the effects of
recession would be negligible until velocities of several hundred miles
per second were reached. The effects would be appreciable at a
few thousand miles per second, and conspicuous at several tens of
thousands of miles per second.
If red-shifts are evidence of actual recession, the dimming factors
should become appreciable near the limits of measurement with the
100-inch and should be conspicuous near the limit of the 200-inch.
At the very limits of direct photographs with the 200-inch, the factor
should approach the order of 40 to 50 percent, and should be unmis-
takable.
We may predict with confidence that the 200-inch will tell us
whether the red-shifts must be accepted as evidence of a rapidly
expanding universe, or attributed to some new principle of nature.
Whatever the answer may be, the result will be welcomed as another
major contribution to the exploration of the universe.
I have mentioned the three specific problems of canals on Mars,
relative abundance of chemical elements in stars, and the nature of
the red-shift, because they illustrate the unique powers of the 200-
inch telescope in three aspects, namely, resolution, dispersion, and
space penetration.
Because these problems are of first importance, and can be solved,
they, together with others of a similar kind, will be included in the
initial research programs. The solutions of these problems alone
will fully justify the construction of the telescope.
But such a program is merely a logical beginning—the first carefully
considered stage in the exploration of vast unknown regions of the
universe. As the darkness is pushed back, greater problems will
doubtless emerge which we cannot now foresee.
FIRST PHOTOGRAPHS WITH THE 200-INCH HALE TELESCOPE?
The first photographs with the 200-inch Hale reflector on Palomar,
made under normal observing conditions, confirm the most optimistic
predictions of its designers. Such a statement, as usual, requires some
explanation. The photographs were made as routine tests to record
progress in the tedious program of adjustments. Seeing was never
better than “‘average,”’ the aluminum coat was dusty and grimy, and
the mirror showed a turned-up edge. These handicaps, of course,
2 A later article by Dr. Hubble from the Publications of the Astronomical Society of the Pacific, vol. 61,
No. 360, June 1949,
186 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
will be eliminated or avoided in time, but during the tests they caused
some loss of light and appreciable loss of definition. Nevertheless,
the test plates record stars and nebulae fully 1.5 magnitudes fainter
than the extreme limit of the 100-inch reflector on Mount Wilson.
The faintest star images, on the better plates, were, however, a little
more than 1 inch in diameter, and, at the threshold, it was sometimes
difficult to distinguish with certainty between stars and nebulae.
Thus the 200-inch has registered already the full gain in light-
gathering power corresponding to the size of the main mirror. The
slight additional gain that may be expected with a clean, sensibly
perfect mirror surface will be accounted for by the absence of a
Newtonian flat and by the very transparent sky over Palomar.
The greatest improvement in the future will be in definition, as
indicated by the size of faint star images. Definition is very sensitive
to the seeing, and, while the test plates approached the definition
to be expected under average conditions, they indicated that the
mirror is not yet in shape to operate at maximum efficiency on the
rare nights of fine seeing. The trouble arises from the turned-up
edge and can be eliminated by the retouching now in progress. The
improved definition will be significant, particularly for distinguishing
nebulae from stars at the threshold of long exposures. In the higher
latitudes, the telescope records many more nebulae than stars.
The turned-up edge was well known from Hartman tests, and its
effects could be predicted with some confidence. The photographs
were made primarily to confirm and record these effects. However,
the first plates were so impressive that a set of full exposures was
made to serve as a record of performance before the mirror was
removed for retouching. About 60 photographs were assembled over
the 3 months from January 26 to April 28, as opportunities arose
during the normal program of adjustments. Of this number, perhaps
half a dozen represent full exposures under average seeing conditions,
and a like number show good performance with reduced apertures or
with a Ross correcting lens for enlarging the usable field. Selections
from the files are illustrated in plates 5 to 10, and comments on them
are given below. The full aperture (200-inch) and Eastman 103a—O
emulsions were used in all exposures except those to which special
references are made in the comments. The scale of the original
negatives is about 1 mm=12’’.
Plate 5, NGC2261; R.A.=6"36"6, Decl. =+8°47’ (1950); Jan. 26,
1949; 15 min. exposure, poor seeing; enlarged 34x.
This plate shows the first of the photographs with the Hale tele-
scope. It is recorded as PH—-1-H (i. e., Palomar, Hale, No. 1, followed
by the initial of the observer). It was made under poor conditions
as a preliminary test of the mechanical operations and procedures
involved in direct photography at the prime focus. The trial was
200-INCH HALE TELESCOPE—HUBBLE 187
successful, except for the large size of the star images produced by
the poor seeing. The exposure was made on January 26, 1949, about
10 p. m., after waiting more than a week for a break in the weather.
The object, NGC2261, is a well-known, variable galactic nebula—a,
comet-shaped mass with the variable star, R Monocerotis, at the apex.
Plate 6, S.A. 57; R.A.=13"6"3, Decl.=-+29°37’ (1950); Jan. 27,
1949; 60 min. exposure, seeing average; enlarged 7% ; center is 2/5 N.
and 3/4 W. of BD+30°2371. This selected area contains one of the
most reliable magnitude sequences available for faint stars extending
to the 21st magnitude. Exposures of 1 minute registered stars to
about 19.7, and of 3 minutes, to about 20.7. Exposures of 5 or 6
minutes reached the extreme limits of the 100-inch, beyond the end
of the sequence. From these data it is estimated that the 60-minute
exposures permitted by the dark sky above Palomar reached at least
1.5 magnitude beyond the 100-inch.
The threshold of the plate is dominated by nebulae rather than by
stars, and this fact emphasizes the tremendous range of the telescope.
Some of the faintest nebulae recorded are presumably at about twice
the distance reached with the 100-inch or, in round numbers, at about
1,000 million light-years. This figure, of course, refers to average
nebulae. Individual images on the photograph may represent dwarf
nebulae at lesser distances or giant nebulae, even more remote.
Plate 7, Messier 87 (NGC4486); R.A.=12°2873, Decl.=-+12°42’
(1950); Apr. 27, 1949; exposure 45 min., seeing average; enlarged 8X.
The object is one of the brightest members of the Virgo cluster of
nebulae, whose distance is of the order of 7.5 million light-years. It
is classified as a peculiar elliptical nebula (KOp). The photograph
shows the nebula, presumably a globular mass of type II stars (1. e.,
stars similar to those in globular star clusters), surrounded by an
extensive, tenuous atmosphere of supergiant stars. ‘This phenomenon
was suggested by the best photographs made with the 100-inch on
Mount Wilson, but is conspicuous on the 200-inch plate.
Plate 8, NGC5204; R.A.=13"28"0, Decl.=+ 58°38’ (1950); Jan.
31, 1949; exposure 30 min., seeing average; enlarged 4X.
The object is a dwarf, late-type spiral in Ursa Major, at an esti-
mated distance of less than 3 million light-years. The plate is in-
cluded to show the ability of the telescope to resolve the neighboring
stellar systems so that the brighter stars can be studied individually.
Photographs of several of the larger spirals, such as M 81, NGC2403,
etc., have been made, but the coma-free field of the telescope at full
aperture is so small (about 5 minutes of are in diameter) that the
plates are not suitable for reproduction on a scale sufficient to show
the resolution to advantage.
Plate 9, NGC3359; R.A.=10"43"4, Decl.=+63°20’ (1950); Apr.
27, 1949; exposure 45 min., seeing average; enlarged 3X.
188 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
This late-type barrel spiral is an isolated stellar system or, possibly,
an outrider of the Ursa Major cloud of bright nebulae, and its distance
is of the order of 5 million light-years. The plate is included to illus-
trate the resolution of fairly distant nebulae, and the opportunities
now available for the study of the very brightest stars in stellar
systems.
Plate 10, Messier 3 (NGC5272); R.A.=13539"5, Decl.=+28°38’
(1950); Apr. 21, 1949; exposure 3 min., made with an aperture of 160
inches, and a Ross correcting lens; seeing average; enlarged 8X.
This plate, of a well-known globular star cluster, is included to
illustrate the use of a Ross correcting lens, placed a few inches in
front of the plate, to enlarge the coma-free field of the telescope.
The lens performed well with the 160-inch aperture but, with the
full 200-inch, it exaggerated the effects of the turned-up edge of the
main mirror. The provisional mounting of the lens did not permit
the use of a guiding eyepiece, so the plate shows the successful per-
formance of the tracking mechanism of the telescope during an
unguided exposure of 3 minutes. The usable field with this correcting
lens is more than 15 minutes of are in diameter.
THE DETERMINATION OF PRECISE TIME!
By Str Haroup SPENCER JONES
Astronomer Royal of Great Britain
Of the three fundamental physical units, there is an essential
distinction between the unit of time and the units of mass and length.
The units of mass and length are represented by material standards
to which any mass or length can be related, either directly or indirectly.
But the unit of time cannot be represented by any material standard.
For practical purposes time can be thought of in the Newtonian sense
as something which flows uniformly. The passage of time can be
marked by a clock, and any simple natural phenomenon which obeys
one definite law without perturbation might be used to mark off equal
intervals of time and therefore to serve as a clock. The rotation of
the earth provides us with a natural clock. We shall see later that it
is not a perfect clock, but that it is sufficiently uniform for almost all
practical purposes; it has, moreover, the great advantage of never
stopping.
We can therefore define the unit of time as the period of rotation
of the earth. Some reference object must be selected against which
to measure the rotation. For the purposes of everyday life, time
must be related to the sun, whose rising and setting gives the alterna-
tion of daytime and nighttime. The day defined by the rotation of
the earth with respect to the sun is called the true solar day; it is the
interval between two consecutive transits of the sun across the
meridian of any place. With this unit, true solar time is obtained
by dividing the true solar day into 24 hours and calling the instant of
meridian passage of the sun 12 hours. The time given by a sundial
is true solar time. For practical purposes, however, true solar time
is not convenient; because the motion of the sun across the heavens
is not uniform, the length of the solar day varies in length throughout
the year. For civil purposes, therefore, a mean solar day is used,
whose length is equal to the average length of the true solar days
throughout the year. The time based on the mean solar day as unit
is called mean solar time. The relationship between mean solar time
and true solar time at some particular instant is defined by means of
1 Sixteenth Arthur lecture, given under the auspices of the Smithsonian Institution April 14, 1949,
189
190 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
a convention, into the details of which I need not enter. The extreme
differences between mean and true solar times range from 16 minutes
about November 3, when true noon precedes mean moon, to 14%
minutes about February 12, when true noon follows mean moon.
Astronomers, however, find it more convenient to determine time
by the observation of the stars. There are many stars but only one
sun and, moreover, the time of transit of a star can be determined more
accurately than the time of transit of the sun. The sidereal day is
defined by the rotation of the earth relative to the stars. It is about
4 minutes shorter than the solar day. If we imagine the sun and a
star to be on the meridian of some particular place at the same instant,
then after the lapse of one sidereal day the star will again be on the
meridian; but, because of the orbital motion of the earth round the
sun, the earth will have to turn a little more in order to bring the sun
onto the meridian. In the course of a year the earth completes its
orbit around the sun and there must consequently be exactly one more
sidereal day in the year than mean solar days.
If the relative positions of a number of stars in the equatorial region
of the sky have been accurately determined, we can think of them as
equivalent to the graduations on the face of a clock. As the earth
rotates, a telescope, fixed so as to be able to move only in the meridian,
will sweep across these stars in turn, each at a definite specific instant
of sidereal time. By observing the transit of stars whose positions
are known, the sidereal times at the instants of meridian transit are
therefore determined. The beginning of the sidereal day or, in other
words, Oh. of sidereal time, is defined by the transit of the point in
the sky at which the ecliptic crosses the equator from south to north;
this point is called the vernal equinox or the First Point of Aries.
By defining the commencement of the sidereal day in this manner,
we are provided with a means for converting from sidereal time to
mean solar time, which is required for the purposes of everyday life.
But it has one inconvenience. The First Point of Aries is not fixed
relative to the stars. It has a slow retrograde motion, due to the
precessional motion of the earth’s axis, and superposed on this uniform
motion is a slow to-and-fro drift, caused by the nutational or nodding
motion of the axis. The nutation depends upon the relative positions
and distances of the sun and moon from the earth. The principal
term in the nutation has a period of about 18 years and a semi-
amplitude of about 1 second of time. There is also a 6-monthly term
amounting to 0.08 second and a number of short-period terms
amounting to 0.020 second, of which the principal term has a period
of 2 weeks. The precision of modern clocks is such that these small
terms cannot be neglected. The true sidereal day, measured relative
to the true position of the First Point of Aries, is therefore not abso-
lutely uniform in length, and it is necessary to introduce the con-
DETERMINATION OF PRECISE TIME—-SPENCER JONES 191
ception of mean sidereal time, measured relative to the mean position
of the First Point of Aries. Actual observation of the stars provides
the astronomer with true sidereal time, which he then has to correct
for the nutation to obtain mean or uniform sidereal time.
The determinations of time by astronomical observations are used
to control the performance of a standard clock, determining its error
at a specific instant and the rate of increase or decrease of that error,
the clock then being used to obtain the time at other instants. This
usually involves extrapolation to some time subsequent to the latest
observation. For such extrapolation to be accurate, the time de-
terminations must not be affected by serious errors and the standard
clock must be of high precision. The determination of precise time
therefore involves two problems, the determination with high ac-
curacy of the time at specific instants and the development of time-
keepers of very high precision.
The sidereal time of the transit of a star across the meridian is
equal to the right ascension of the star. Sidereal time can therefore
be determined by observing the times of meridian transit of stars of
known right ascension. The conventional method of making the
observations has been to use a transit instrument. This consists of a
telescope, mounted on an axis at each end of which is a cylindrical
pivot. The pivots rest in fixed bearings, adjusted so that the common
axis of the pivots is as nearly as possible horizontal and pointing in an
east-west direction. If the axis of the pivots were exactly horizontal
and in the east-west direction and if the optical and mechanical axes
of the telescope coincided, the axis of the telescope would be in the
meridian plane, whatever direction the telescope was pointing to.
This ideal condition is never achieved and there are always small
errors of level, of azimuth, and of collimation. These adjustments
are liable to continual change; there are slow seasonal changes, as-
sociated with changes of temperature and possibly also with sub-
surface moisture; there are also more rapid changes, which are cor-
related with changes of circumambient temperature and with the
direction of the wind. To control these changes frequent observa-
tions of level, of azimuth, and of collimation are essential, which take
up a disproportionate amount of the observing time. The error of
collimation can, however, be eliminated if the telescope is reversed
in its bearings in the middle of each transit, half the transit being
observed before reversal and the other half after reversal. It is not
possible to reverse large transit instruments sufficiently quickly and
it has accordingly become customary to use small transit instruments,
which can be rapidly reversed, for the determination of time; as it
is the brighter stars which are observed, a large aperture is not needed.
There are other factors which have also to be taken into considera-
tion. The pivots will never be absolutely cylindrical; their figures
192 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
have to be determined with great accuracy and appropriate corrections
made to the observations. Flexure of the axis can cause troublesome
systematic errors. If the horizontal axis is not equally stiff in all
directions, its flexure will vary according to the direction in which the
telescope is pointed. If the two halves are not equally stiff, the
telescope will be twisted from the meridian by a variable amount.
Personal equations between different observers are somewhat trouble-
some, though they do not exceed a few hundredths of a second when
the so-called impersonal micrometer is used. Before its introduction,
the method of observing was for the observer to press a hand-tapper
at the instant the star crossed each of a number of vertical spider
wires in the focal plane of the telescope; by so doing, he closed an
electric circuit which sent a current to a recording chronograph, which
recorded not only the signals from the telescope but also time signals,
every second or alternate seconds, from the clock. The instants
of the star crossing the wires could then be read off at leisure after the
observations had been completed. With this method of observing,
the times determined by different observers could differ by as much as
halfasecond. The reason is easy to see; one observer might wait until
he saw the star actually bisected by the wire before he pressed the
tapper, with the result that, because of the time required for the mes-
sage to travel from his brain to his eye and to be converted into
muscular action, his signal would inevitably be late; another observer
would, as it were, shoot the flying bird, gauging the rate of motion of
the star so that his tap is made as nearly as possible at the instant at
which the star is actually bisected. The personal equations can be
determined by what are called personal equation machines; the
transit of an artificial star is observed, the times at which the star is at
certain positions during the transit beng compared with the observed
times. Although an observer will unconsciously form a fixed habit
in observing so that his personal equation remains substantially
constant, small variations, depending upon the physical condition of
the observer, do occur.
The method of observing now almost universally employed is to
have a single movable wire in the micrometer eyepiece instead of
a number of fixed wires. The wire can be traveled along by the
observer, who adjusts its speed so as to keep the star continually
bisected by the wire. As the wire moves along, contacts are auto-
matically made in certain positions, sending signals which are re-
corded on the chronograph. In order to relieve the observer of some
of the strain of maintaining a uniform motion of the wire, it is now
common to drive the wire mechanically at the speed appropriate to the
motion of the star, using an electric motor with some form of con-
tinuously variable gearing. With this method of observing, the
personal equations of different observers are very small, usually not
DETERMINATION OF PRECISE TIME—SPENCER JONES 193
more than two or three hundredths of a second; it is for this reason
that this form of micrometer is called the “impersonal”? micrometer.
Small though these residual personal equations are, they remain
remarkably constant and can be determined by personal equation
machines. They seem to arise from two causes: there is “bisection
error,” an observer systematically bisecting an image to the right or
to the left of its center; this error changes sign at the zenith with
instruments in which the observer changes the direction in which he
faces, according to whether he is observing a north or a south star;
there is also “‘following error,” an observer systematically setting the
wire in front of or behind the center of a moving image. This error
does not change sign at the zenith.
If the pivots are not exactly cylindrical, the telescope will be
twisted out of the meridian by an amount varying with its position.
The figures of the pivots must therefore be determined with great
accuracy and appropriate corrections applied to the observed times of
transit. The figures of the pivots must be determined at intervals,
as they may change slowly in the course of use through wear. Other
variable errors can be introduced through slight mechanical imperfec-
tions in the telescope; if there is the slightest play in the eyepiece
micrometer or in the objective, errors will be introduced which will
vary with the position of the telescope.
When all the possible sources of error which can affect observations
with a transit instrument are borne in mind, it is rather surprising
that the observations are as accurate as they are. The probable error
of a single time determination is usually about two-hundredths of a
second. This was quite accurate enough before the era of clocks of
high precision and before there were any practical requirements for
very precise time. The scatter of the observations is, however, incon-
veniently large for the adequate control of the performance of the
modern quartz-crystal clock. For these reasons the conventional
transit instrument is likely to be gradually replaced for the purpose
of time determination by some other type of instrument. Several
modifications of the transit instrument have been considered which
eliminate or minimize some of its disadvantages. The most accurate
results are given, however, by an entirely different instrument known
as the photographic zenith tube. It consists of a fixed vertical telescope
pointing to the zenith, which has a mercury horizon at the bottom of
its tube, whose purpose is to reflect the light from a star to a focus in
the plane of the second principal point of the objective. The funda-
mental principle of the instrument was due to Sir George Airy, who
first used it for the reflex zenith tube at Greenwich: when the light is
brought to a focus accurately in the plane of the second principal
point of the objective, the results are unaffected by tilt of the telescope.
The troublesome error of level is therefore immaterial, while any error
194. ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
of azimuth does not affect observations made in the zenith. The
telescope is constructed so that the objective and the plate holder can
be rotated through 180°, the observations being made photographically
in order to eliminate personal equations and to give greater accuracy.
Suppose two exposures are given on a star at times which are sym-
metrical about the time of meridian transit, the objective and the
photographic plate being rotated through 180° between them. The
two images will lie on a line exactly parallel to the meridian. If,
however, the two times of exposure are not exactly symmetrical, the
images will be slightly staggered; by measuring the staggering and
knowing the clock times of the two exposures, the clock time of
meridian transit can be inferred.
In practice an exposure of finite length is required to give a
measurable image on the plate. During this exposure the plate holder
is traveled along at the speed appropriate to the motion of the star,
signals being sent to the chronograph at certain definite positions of
the plate holder. After reversal the plate carriage retraces its path,
and signals are sent during the course of the second exposure at the
same positions.
With this design of instrument, collimation error does not enter,
there are no pivot errors to be considered, and the various sources of
error inherent in a movable instrument are avoided. At the Naval
Observatory, Washington, a photographic zenith tube, designed and
used by F. E. Ross originally for the determination of the variations
of latitude, has been used for some years for the determination of time.
An instrument on the same general principle, but differing materially
in details of design, is in an advanced stage of construction for the
Royal Greenwich Observatory. The errors of time determination
should not exceed 2 or 3 milliseconds, which will permit a tight control
of the performance of the observatory clocks.
For the purpose of time determination it is necessary to assume
positions for the stars which are observed. These positions will have
random errors, whose effects can be reduced by observing sufficient
stars. But they may also be affected by systematic errors; if, for
instance, the errors vary with right ascension they will introduce a
spurious systematic variation in the derived clock error through the
year. For the purpose of time determinations and in order that the
times determined at different observatories can be directly compared,
there is an international agreement to use the positions of the stars
given in the fundamental star catalog known as the FK3. These
are bright stars, whereas with the photographic zenith tube, inasmuch
as observations are restricted to a narrow belt at the zenith, it is
necessary to use fainter stars. Their positions must therefore be
determined by transit circle observations and tied on to the FK3
system, The photographic observations will in course of time provide
DETERMINATION OF PRECISE TIME—SPENCER JONES 195
some measure of control over the periodic errors in right ascension of
the FK3 system itself.
Until about 25 years ago, pendulum clocks of the regulator type
were used as the standard clocks in observatories. A considerable
improvement in precision was brought about by the invention of the
free-pendulum clock. In an ordinary pendulum clock the timekeeping
is impaired by the variable friction involved in driving a train of
wheels to move the hands and record the actual time on a dial. An
appreciably higher accuracy is to be expected if the pendulum is
allowed to swing freely, except when it receives periodically impulses
to maintain its swing, and is thereby relieved from all extraneous
work. To achieve this had been the aim of horologists for many
years, but although many attempts were made it was not really
successfully accomplished until the invention by W. H. Shortt of his
free-pendulum clock. The master pendulum is enclosed in an airtight
case, in which the air pressure is reduced to about 1 inch of mercury
and which is maintained at constant temperature and swings freely,
except for small impulses, given at half-minute intervals, to main-
tain the amplitude at a nearly constant value. The slave clock
is a normal synchronome electric clock, which is adjusted when swing-
ing as an independent clock to lose about 6 seconds a day. The
synchronizing action required from the master free pendulum is there-
fore a one-way action—always an accelerating action. The slave
pendulum itself releases electrically the impulsing lever of the free
pendulum, which falls when the free pendulum is at the midpoint of
its swing. The impulse arm falls on the top of a small pivoted wheel,
mounted on the free pendulum; this being a dead point, and the
impulse not commencing to be given until the pendulum swings
outward from the central position, the amount of the impulse does
not depend upon any slight variation in the synchronization between
the two pendulums which may occur.
The synchronization of the slave pendulum is effected by means of a
light flexible spring carried on it. The impulse arm of the free
pendulum, after it has fallen clear of the pendulum, actuates a device
which closes an electric circuit and sends a current through a small
electromagnet adjacent to the slave pendulum. If the slave clock has
dropped sufficiently behind the master, the armature of this electro-
magnet will, when the electromagnet is excited, engage the bent end of
the light spring on the slave pendulum. The end of the spring is then
held fixed and, as the pendulum swings, the spring is flexed and the
pendulum is accelerated; if, on the other hand, the slave pendulum is
closely in phase with the master, the end of the spring passes under
the armature before the electromagnet is excited, and nothing happens.
The length and strength of the spring are so adjusted that when the
synchronizing action occurs the slave pendulum is accelerated by
196 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
%4o second. As the natural losing rate of the slave clock, 6 seconds
a day, is equivalent to go second per minute, the synchronizer, which
is actuated each half minute, should hit and miss alternately. For
this reason it is called the ‘‘hit-and-miss” synchronizer.
The first experimental Shortt free-pendulum clock was installed at
the Edinburgh Observatory in 1921. It at once proved to be such an
improvement upon previous pendulum clocks that two were installed
at the Greenwich Observatory in 1923 and others in subsequent years.
It was the excellent performance given by these clocks that made it
necessary for astronomers for the first time to introduce the conception
of mean sidereal time. Previously true sidereal time had been uni-
versally used, as clocks were not good enough to be able to show up the
small effects due to the short-period terms in nutation. The free-
pendulum type of clock is capable of an accuracy of about one-
hundredth of a second a day. Detailed investigation of their
performance has shown, however, that such clocks are liable to
frequent small erratic changes of rate, of the order of about 3 milli-
seconds a day. Small though such changes are, they cause, by
integration, an irregular wandering of the clock. For sending out
time signals, it is always necessary to extrapolate beyond the latest
time determination; these erratic changes of rate restrict the accuracy
with which the error of the clock can be extrapolated. It can, on
occasion, happen that 2 weeks or more may elapse without any check
on the performance of the clock being possible and the transmitted
time signals may consequently be appreciably in error. Moreover,
because of the errors of observation, there is a natural scatter in the
derived errors of the clock. In interpolating between the observed
errors there is no means of distinguishing between scatter due to errors
of observation and scatter due to the irregular wandering of the clock.
It is possible, of course, to attempt to reduce the effects of the
wandering by using the mean of several clocks. Nevertheless, very
high accuracy cannot be obtained, because residual effects due to the
irregularities are always present.
A new standard of accuracy has been provided in recent years by
the use of an oscillating quartz crystal, developed originally to serve
as a precision standard of frequency. The quartz clock is based upon
the piezoelectric property of quartz. If a plate of quartz is com-
pressed, the two opposite faces become electrically charged, one
positively and the other negatively. Conversely, if two opposite
faces are given positive and negative charges respectively, the piece
of quartz experiences a mechanical contraction or expansion. By
rapidly alternating the electric charges, the quartz can be maintained
in mechanical vibration. In the quartz clock an oscillating electrical
circuit is used, the dimensions of the crystal being adjusted so that its
DETERMINATION OF PRECISE TIME—SPENCER JONES 197
natural resonance frequency is equal to the frequency of the oscillating
circuit. Under these conditions a strong vibration is set up and the
quartz crystal takes control and locks the frequency of the oscillating
electrical circuit to its own resonance frequency. Quartz is a very
stable substance and, provided it is maintained at a very uniform
temperature and the drive circuit is properly designed, the frequency
remains constant to a high degree of accuracy. It is usual for the
crystal to be cut to give a frequency of 100,000 cycles a mean time
second, the dimensions of the quartz then being conveniently small.
This frequency is divided down in steps electronically, either by the
use of multivibrators or by frequency subdivision until an output
with a frequency of 1,000 cycles a second is obtained. The output
of this frequency is used to drive a phonic motor, from which time
signals can be obtained at any desired intervals.
Such clocks have many advantages over pendulum clocks. They
have proved to have very high short-period stability. Their erratic
changes of rate are less than half a millisecond a day, and the clocks
themselves can be relied upon to about 1 millisecond a day. For
extrapolating between scattered time determinations they are there-
fore much superior to pendulum clocks. They have, moreover, the
advantage of the great flexibility inherent in dealing with 100,000
vibrations a second instead of only a single one. Electronic methods
can be used for quickly and accurately determining the relative errors
and rates of the clocks. For such purposes at Greenwich, decimal
counter chronometers are used. This device consists of a scale-of-ten
counter, and is actuated by the 100,000-cycle output per second from
one of the quartz crystals. When it is switched on, it will start
counting these vibrations, recording the count on five decade dials,
reading, respectively, tenths, hundredths, thousandths, ten-
thousandths, and hundred-thousandths of a second. To compare
two quartz clocks, a seconds signal from the phonic motor driven by
the one clock is used to start the count and a signal from the second
clock to stop it. The time difference between the two clocks, accurate
to a hundred-thousandth of a second, is thus obtained in a fraction of a
second. As a check, the second clock can be used to start the count
and the first clock to stop it. The difference in frequency of the two
clocks is obtained by feeding the 100,000 ¢. p. s. outputs from the two
clocks into a comparator, so that they beat against one another, and
timing the beats. It is possible to obtain an accuracy of one part in
10”° in the measurement of the frequency difference.
At Greenwich, the clocks are used in groups of three, one phonic
motor being provided for each group of three clocks. One of the
clocks is selected to drive the phonic motor, but regular comparisons
are made between each pair of clocks in the group. Automatic beat
198 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
counters record the integrated time difference between each pair,
A-B, B-C, C-A, the third comparison providing a check on the
other two.
A further convenience of the quartz clocks is that it is not necessary
to maintain separate mean-time and sidereal-time clocks as it is
with pendulum clocks. By means of suitable gearing, it is possible
to take sidereal seconds direct from the phonic motor which gives
also mean time seconds. The ratio of the mean time second to the
sidereal time second is 1.002 737 909 293. This ratio can be closely
represented by a gearing of 119/114 multipled by 317/330, which is
only 4 parts in 10° small. These sidereal second signals are used for
recording on the chronograph during the time determinations. When
the rate of the clock relative to these signals has been derived it is a
simple matter to infer its rate relative to true mean time seconds.
The small error in the conversion from mean time to sidereal time is,
of course, eliminated.
For short-period prediction quartz clocks leave little to be desired.
They have not as yet, however, reached the stage at which long-period
prediction has the accuracy that is desirable. The difficulty arises
from a slow drift in frequency to which they are all liable. The crystal,
after cutting, appears to go through a slow ageing process; the drift
in frequency is rather rapid at first, but progressively diminishes
though it sems never to cease altogether. Iffor any reason the crystal
should stop, through a tube or resistor giving out, it will not, when
restarted, follow along its previous ageing curve; a new ageing cycle
sets in. Any small disturbance, such as a slight temperature change,
can alter the frequency drift somewhat. The effect of the frequency
drift on the error of the clock increases with the square of the time
so that, even though the drift may be quite small, its effects will
become important with lapse of time. With the present scatter in
the actual time determinations, several months’ observations are
needed to give a sufficiently accurate derivation of the frequency
drift, but there is always the uncertainty whether during this period
some small disturbance may not have caused the rate of drift to change
slightly.
Moreover there are extraneous effects which can complicate the
determination. During a period of several months, there will be a
wide range in the right ascensions of the stars which are used for the
time determinations. If there are periodic errors in the fundamental
system of star places, a spurious factor will have entered into the
determination of the frequency drift. The motions of the earth’s
poles cause further complications. The poles have an irregular mo-
tion, which is roughly circular, but with a variable radius. The
extreme departures of the true poles from their mean positions are
about 30 feet. ‘The movement of the pole along the meridian causes
DETERMINATION OF PRECISE TIME—SPENCER JONES 199
a variation in latitude, which can be observed with a zenith tele-
scope. The movement in the perpendicular direction causes a dis-
placement of the meridian. The motion of the pole has two main
components, with periods of a year and of about 14 months respec-
tively. As a consequence of this motion, it would be found that if we
had a perfect clock, with no rate at all, and observations which were
entirely free from error, the clock would appear to have a slightly
variable rate. This apparent variation of rate will affect the deter-
mination of the frequency drift and give a spurious value.
It is not possible at an observatory to measure the component of
the polar motion at right angles to the meridian. At Greenwich
an approximate compensation for the motion is made through the
cooperation of the Naval Observatory, Washington, which sends
regularly to Greenwich the observed movement of the pole along the
meridian of Washington. If Washington were 90° in longitude west
of Greenwich, the displacement along the meridian of Washington
would also be the displacement at right angles to the meridian of
Greenwich. But the longitude of Washington is only 77° west of
Greenwich. However, the use of the Washington latitude-variation
data does enable the greater part of the polar-motion effect to be elim-
inated from the Greenwich clock curves and it has been noticeable
that the inferred performance of the clocks has thereby been improved.
The development of an atomic or molecular clock, in which the
frequency of some selected atomic or molecular vibration will be
subdivided to give a frequency closely equal to that of an oscillating
quartz crystal and used to lock the vibrations of the crystal, is
already foreshadowed by the work in progress at the National Bureau
of Standards, Washington, in the development of an ammonia clock,
in which the frequency of one particular mode of vibration of the
ammonia molecule is used as the control. This work is as yet in its
early stages and has not gone beyond the point of showing that the
control of a quartz crystal in the way suggested is practicable. When
the clock has been developed to the stage at which the accurate con-
trol of a precision quartz clock becomes possible, the crystal will be
prevented from drifting in frequency. ‘The clock error curve over a
long period of time should then be represented by a straight line.
Departures from a straight line could be attributed to periodic errors
in the star places, to the polar motion, or to irregularities in the
rate of rotation of the earth itself. Much more accurate long-term
prediction would become possible, with a considerable gain in the
accuracy of timekeeping.
It has been well established that the length of the day is subject
to small fluctuations. It has long been known that there are discord-
ances between the observed and the tabular positions of the moon
which are not attributable to imperfections in the theory of the
866591—50-——14
200 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
motion of the moon. In the development of the theory, the gravita-
tional effects which have been neglected are far too small to amount
to anything like the discordances which are observed. In more
recent years it has been proved that there are similar fluctuations in
the motions of Mercury, Venus, and the sun; but for these bodies
the effects are much smaller than for the moon because their mean
motions are much less rapid. It was the comparative smallness of
the effects for these bodies which made their detection difficult. So
there are, in effect, four clocks which agree together and one clock,
our earth, which differs from the other four. The natural conclusion
is that it is the earth which is at fault and that the length of the day,
which has been adopted as the unit of time and assumed to be invar-
iable, is actually subject to small variations.
The changes in the length of the day are found, from the analysis
of the observational data, to be of two different kinds. There is a
slow progressive increase in length, of the order of 1 millisecond in
the length of the day in the course of a century. This progressive
increase is caused by tidal friction, more particularly in the shallow
sea; it acts as a brake on the earth. Though so small in amount, the
effect on the mean longitudes of the moon and the planets increases
with the square of the time and is large enough to make the position
of the moon 20 centuries ago, if computed from its present motion in
longitude, very considerably in error. The effect was actually first
detected in 1679 by Hailey from the early observations of eclipses.
Superposed on the progressive increase of length there are also irregular
changes, the day sometimes increasing in length and sometimes de-
creasing; these changes cannot be attributed to tidal friction, because
frictional effects can cause only a slowing down and never a speeding
up in the earth’s rotation. These changes are due to changes in the
earth’s moment of inertia and could be accounted for quantitatively
if the earth expanded or contracted slightly by 4 or 5 inches.
There is one essential difference between the two phenomena.
A change in the moment of inertia of the earth is something that con-
cerns the earth alone. The apparent displacements of all the other
bodies are strictly proportional to their mean motions. But tidal
friction is something that concerns the earth and the moon jointly;
the total angular momentum of the earth-moon system is conserved,
but there is interaction between the earth and the moon. The appar-
ent displacements of Mercury, Venus, and the sun will again be
proportional to their mean motions but the same will not hold for
the moon; its displacement will not have the same ratio to its mean
motion. It is this difference in the case of the moon which makes it
possible to separate the two effects of tidal friction and of change
of the moment of inertia of the earth.
DETERMINATION OF PRECISE TIME—SPENCER JONES 2(1
Though the changes in the length of the day have been fully estab-
lished by these observations, the data are not sufficiently accurate to
decide whether the changes occur suddenly or whether they are spread
over a few days, a few weeks, a few months, or even over a year or two.
If they occur rather suddenly, they could be detected with ease by
quartz clocks in their present stage of development; if spread over a
few months, the larger changes could be detected, but changes of
smaller amount would be likely to escape detection. Since a few
years ago, when quartz clocks were adopted at Greenwich as the
basis for the time service, a close watch has been kept for any evidence
of a change in the earth’s rotation. Once or twice small changes
have been suspected but there has always been some factor which
has made a definite conclusion impossible—perhaps one of the clocks
has changed its rate or has stopped at the crucial time, or there has
been some uncertainty in the determination of frequency drift. The
evidence provided by the observations of occultations of stars by
the moon is that there has been no major change in the earth’s rate
of rotation since about 1918. There may possibly have been small
changes, but no definite conclusions are as yet possible.
It is not inconceivable that there may be small annual variations
in the rate of rotation of the earth. There are seasonal displacements
of matter over the earth’s surface; there is, for instance, a high-pressure
region over Siberia at one season of the year and a low-pressure region
at another season, entailing the displacement of large atmospheric
masses, with corresponding change in the moment of inertia. Such
effects would be tangled up with effects due to periodic errors in star
places and with the effects of the polar motion. Much more is likely
to be learned about these matters when the atomic clock has reached
a further stage of development, so that the frequency drift of the
quartz crystal can be eliminated. Observations with photographic
zenith telescopes should gradually smooth out any residual periodic
errors in star places, while the information they provide about the
variation of latitude will furnish basic data which can be used subse-
quently to separate polar motion effects from small variations in the
earth’s rotation. It may prove, however, that the earth itself is
rather like a pendulum clock in its behavior and that its rate of rota-
tion is liable to frequent and small irregular changes, so that we can
at present merely observe their integrated effect.
The question may arise in the near future how the unit of time
should be defined. Clocks are now at a stage when their stability
for short periods is of a higher accuracy than the earth’s rotation
itself. The earth, however, has the advantage over any clock that
it has no liability to a stoppage. It may be possible to develop
atomic clocks to a stage at which they can be run for several years
202 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
without stopping and to maintain accurate time; with a battery of
such clocks, all controlled by the same atomic vibration, it would be
possible to bridge over the stoppage of any single clock and thereby
to maintain an accurate standard of time more or less indefinitely.
There will be definite objections to using as the fundamental unit of
time a unit that is known to be variable. A new unit should be
absolutely invariable. A clock based fundamentally upon a length
which is controlled by an atomic wave length, and upon the velocity
of light, for instance, seems theoretically ideal.
Note ‘ADDED IN PROOF.—Since the lecture was delivered, investi-
gations at the Greenwich Observatory have established the existence
of a fairly regular annual variation in the rate of rotation of the
earth. Relative to uniform time the earth gets behind by about 60
milliseconds in May-June and ahead by a similar amount in Novem-
ber. The corresponding variations in the length of the day amount
to somewhat more than 1 millisecond a day on either side of the
mean value. H.S. J.
THE ELEMENTARY PARTICLES OF PHYSICS!
By Cart D. ANDERSON
California Institute of Technology, Pasadena
The idea of elementary particles of matter, of small, discrete, in-
divisible particles out of which all matter in the universe is consti-
tuted, is as old as recorded history. The Greeks in their philosophical
speculations discussed at length the question of the ultimate nature of
matter. They realized that there were only two possible choices open
to them; either matter must be thought capable of being divided into
smaller and smaller units without end, or else it must consist of small
units which are themselves wholly indivisible. Many of the Greek
philosophers experienced a philosophical difficulty in trying to conceive
of infinite divisibility, whereas others found it equally difficult to think
of a particle as being truly indivisible. The difficulty is closely akin to
that which one experiences when contemplating the limits of the uni-
verse, and trying to decide in his own mind whether it pleases him
more to think of the universe as unbounded and extending to infinity,
or to imagine a finite universe with definite bounds beyond which there
is nothing, not even space. The idea of the existence of indivisible ma-
terial particles, however, seems to have had more appeal to the Greeks,
and the atomic hypothesis was expounded and developed in the fifth
century B. C., chiefly by Thales, Leucippus, and his distinguished
pupil, Democritus, until in many respects it resembled the views which
are held today.
The views of Democritus were prominent for 500 years but began
to wane after the beginning of the Christian Era and by about A.D.
200 had almost wholly disappeared from European philosophical
thought. The idea of material atoms did not really appear again in
Europe until about the middle of the seventeenth century, a time
marking the beginning of the great era of scientific experimentation
which has continued with an ever increasing tempo up to the present.
During this period, through scientific research based on experimen-
tation, the atomic theory of matter slowly developed. Highlights in
1 Based on material presented in the Sigma Xi Annual Address, A. A. A. S. Centennial, Washington,
September 1948. Reprinted from American Scientist, vol. 17, No. 2, April 1949, by permission from The
Society of the Sigma Xi.
203
204 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
this development were the Laws of Chemical Proportion as discovered
and enunciated by Dalton near the beginning of the nineteenth cen-
tury, and later the successes of the Kinetic Theory of Gases. By the
beginning of the twentieth century, the concept of the chemical atom
had received general acceptance as a theory based on scientific experi-
mentation. The idea of atoms had thus been removed from the realm
of philosophical speculation and had become a proved scientific fact.
According to this picture all matter depending upon its nature consists
of a mixture of varying numbers of the ninety-odd different chemical
atoms. The size and the mass and other properties of most of the
chemical atoms had been determined although not with great preci-
sion.
DISCOVERY OF FIRST ELEMENTARY PARTICLES
During the time when the chemical atom was being firmly estab-
lished as a scientific fact, other scientific investigations were succeed-
ing in proving the existence of at least one particle of matter which
was more elementary in character than the chemical atoms. In the
decade from 1890 to 1900 the discovery of X-rays and radioactivity,
and studies of the phenomena associated with the discharge of elec-
tricity through gases, soon proved the existence of the electron and
showed that the atoms of chemistry must all be considered as complex
structures, structures which are themselves built up of particles of a
more elementary character.
The electron was distinguished from the other particles previously
studied by physicists and chemists in one very important respect. It
was established as a unique particle in the sense that all electrons
were found to be identical with one another, no matter from what form
of matter they were derived. For the first time then the presence of
a particle truly elementary in character was revealed to science. It
was found always to carry a negative electric charge and to have a
mass about 2,000 times less than the hydrogen atom, the simplest
and least massive of all the chemical atoms. The electron immedi-
ately took its place as one of the elementary particles common to all
forms of matter.
The following 30 years, from 1900 to 1930, were extremely fruitful
in furthering our knowledge of the properties of the chemical atoms.
The work of Moseley showed that chemical atoms were members of
a family, all of them being related to one another in a perfectly definite
and simple way. In 1911 the experimental genius of Rutherford in
Cambridge, England, proved the existence of the atomic nucleus, and
in 1919 he succeeded for the first time in producing an atom of oxygen
from the disruption of the nucleus of an atom of nitrogen. Thus in
1919 the will of man for the first time was able to cause the disinte-
gration of an ordinarily stable element, with the accompanying release
ELEMENTARY PARTICLES OF PHYSICS—ANDERSON 205
of nuclear energy. These and other investigations all combined to
prove that the proton, the nucleus of the simplest of all the chemical
atoms, hydrogen, is a constituent of all other chemical atoms, and
hence is in fact one of the elementary particles of matter.
In 1930, then, the physicist had at his disposal two elementary ma-
terial particles, the electron and the protron, in terms of which to try
to understand the structure of all matter. In this undertaking the
physicist realized many great successes, but in many instances his
efforts resulted in sharp failures. Apparently the world was not to
be understood in terms as simple as these.
In general the physicist was successful in understanding those phe-
nomena which we may classify, for want of a better term, as extra-
nuclear phenomena, and he was unsuccessful in understanding those
phenomena which we may classify as nuclear phenomena. By extra-
nuclear phenomena we mean those processes in which the electrons
which form the outer shells of the atom are the active participating
agents; in this type of phenomena the central core of the atom, or the
nucleus, is present but remains undisturbed and does not participate
actively. Nuclear phenomena, on the other hand, are those in which
the nucleus is the active participant.
Extranuclear phenomena and nuclear phenomena have a great
many distinguishing characteristics. One of the most interesting and
important of these distinguishing characteristics is concerned with the
level of energies involved. Extranuclear phenomena involve very low
energies as compared with nuclear phenomena. The physicist uses the
term electron volt as a measure of energy. The energies of extranuclear
phenomena are found usually to range from a fraction of one electron
volt to several electron volts, whereas nuclear phenomena are found
usually to correspond to several millions of electron volts.
In our environment almost every phenomenon in nature represents
an extranuclear phenomenon: for example, the burning of coal, the
growth of plants, the generation of electric power by conventional
means, the fermentation of wine, the explosion of dynamite, and others
in uncountable numbers. Nuclear phenomena are not so common-
place, but a few examples may be mentioned: for example, the gen-
eration of the sun’s heat, the decay of radium, the manufacture of plu-
tonium, the absorption of cosmic rays in the earth’s atmosphere, the
explosion of an atom bomb.
The concept of energy has been introduced here because of the great
importance that this concept has in the discussion of any physical phe-
nomenon. I have stated that extranuclear phenomena represent low-
energy phenomena and nuclear phenomena represent high-energy
phenomena. To be more accurate I should have said that in extra-
nuclear phenomena we find low concentrations of energy; that is, the
energy changes that one associates with a single elementary particle
206 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
are low in extranuclear phenomena and high in the case of nuclear
phenomena. Moreover, physicists for the past several years have been
studying certain phenomena which represent energy concentrations
many thousands of times greater than those represented even by
nuclear phenomena. This range of energies has been called the range
of ludicrously high energies. So far the only opportunity the physicist
has had to study phenomena in the range of ludicrously high energies
is in connection with observations associated with cosmic rays, and we
shall see in a moment that important knowledge of the elementary
particles of matter has come from studies of phenomena in the range
of ludicrously high energies.
As stated previously, by 1930 two elementary particles of matter
were known to the physicist, the electron which always occurred with
a negative electric charge and the proton which always occurred with
a positive charge. When considered in a manner consistent with the
theoretical concepts as they had been developed up to that time in
terms of the quantum mechanics, the negative electron and the positive
proton served quite successfully as building blocks in terms of which
to understand the structure of atoms so far as the extranuclear phe-
nomena were concerned. But when attempts were made to picture the
structure of the nuclei of the various chemical atoms, or to understand
nuclear phenomena, the attempts usually ended in failure.
Then suddenly in 1932 two new elementary particles were dis-
covered: the neutron and the positive electron, or positron. The known
elementary particles were therefore doubled in number, increasing from
two to four, and providing the physicist with more material with which
to work.
The discovery of the neutron, which came as a result of experiments
performed in Germany, in France, and in England, was immediately
welcomed, for now neutrons together with protons could serve as the
building stones for the various types of atomic nuclei. One very grave
and fundamental problem which formerly had been present was now
removed immediately, for it was no longer necessary to assume the
existence of electrons inside the nucleus, a concept which always had
been accompanied by serious theoretical difficulties.
The discovery of the positive electron, or positron, came during a
series of experiments being performed for the purpose of measuring
the energies of the particles produced by cosmic rays. The discovery
of the positron was an unexpected discovery. This statement is true
even though, about 2 years before, a British physicist, Dirac, had an-
nounced a new theory which actually predicted the existence of posi-
trons. This new feature of physical theory was not welcomed by
physicists, however; it was on the contrary considered to be an un-
fortunate defect in the theory, and many attempts, by Dirac himself
ELEMENTARY PARTICLES OF PHYSICS—ANDERSON 207
and others, were made to remove it, although all were unsuccessful.
If even one physicist in the world had taken the theory of Dirac
seriously, he would have had an admirable guide leading directly to
the discovery of the positron. Had this happened, the positron
would almost certainly have been discovered by 1930 rather than in
1932. However, after the positron was shown actually to exist, then
it was a very short time indeed until many of its properties were
understood in terms of the Dirac theory.
ELEMENTARY PARTICLES AND RADIATION
The discovery of the positron represented the first instance in which
it was recognized that an elementary particle of matter may have only
a transitory existence. In ordinary matter, for example, the average
life span of a positron is only a few billionths of a second, for when
a positron and a negative electron come close to one another they
mutually annihilate one another—the two particles disappear and in
their place one finds only radiation. The whole of the material sub-
stance constituting the particles is spontaneously transformed into
radiant energy. Measurements show that this process is quantita-
tively in accord with the now famous Einstein equation H=mc?, which
relates mass and energy. ‘The process which is the inverse of the
annihilation of material particles also occurs, namely, the production
of particles out of radiation. If radiation of sufficiently high energy
is passed through matter, electrons and positrons are generated. In
this process the material substance of the two particles is actually
created out of the energy represented by the radiation, and again in
conformity with the Einstein equation H=me’.
In the light of these happenings one must change basically his con-
cept of the elementary particles of matter; these particles are no longer
to be thought of as permanent objects which always preserve their
identity, and which serve only as building blocks of matter by joining
together in groups to form the more complex chemical atoms. One
must recognize instead the possibility of the creation of material
particles out of radiation, and the annihilation of material particles
through the production of radiation. Such a possibility as this, of
course, was completely inconceivable to the Greeks in their long
philosophical discussion on the indivisibility of matter versus the
divisibility of matter.
A further step toward a realization of the great complexity inherent
in the relationships among the elementary particles of matter came in
1935 with the discovery of the positive and negative mesotrons, or
positive and negative mesons as they are now often called. This dis-
covery was also made in investigations of the high-energy phenomena
occurring when cosmic rays are absorbed in their passage through
matter.
208 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The mesotron is a particle some 200 times as massive as an electron,
and therefore about one-tenth as massive as either a proton or neutron.
It occurs with both positive and negative electric charge. The dis-
covery of the mesotron did not come quickly and accidentally as was
the case with the positron and the neutron. It came only after the
completion of a sustained series of observations, covering a period of
4 years, which were designed to remove certain inconsistencies always
present when we attempted to understand certain ‘cosmic-ray phe-
nomena in terms of the elementary particles then known. These
inconsistencies were removed in terms of the existence of the mesotron,
whose discovery was publicly announced in 1936.
Unlike the neutron, the mesotron was not a particle to be imme-
diately welcomed by the physicist. The physicist makes his advances
by simplifying his understanding of nature; hence, a physical world
which could be explained in terms of only one or two distinct elemen-
tary particles would be most to his liking. The discovery of the
mesotron did not introduce a simplification; rather it complicated the
situation for it increased the number of material elementary particles
from four to six. Apparently the Creator does not favor a world of
too great simplicity.
Before the discovery of the mesotron a Japanese physicist, Yukawa,
had postulated on theoretical grounds the possible existence of parti-
cles of a mass intermediate between a proton and an electron. His
theory, however, was not generally known to physicists at that time,
and did not have any part at all in the discovery of the mesotron.
Had this theory been generally known it is still doubtful if it would
have affected the course of cosmic-ray research, since, unlike the
Dirac theory of the positron, it would not have served as so useful a
guide in pointing out the most fruitful directions for the research to
follow.
Like the positron the mesotron has a very short life expectancy.
In free space, both positive and negative mesotrons have a normal life
span of just over two-millionths of a second, after which time they
spontaneously disintegrate. Very recent observations have shown
that in all probability the spontaneous disintegration of a mesotron
results in the simultaneous production of an electron and two neutrinos.
Neutrinos are the interesting elementary particles which had pre-
viously been invented in order to balance energy and momentum in
the process in which an electron is produced when a radioactive nucleus
decays. A similar situation exists in the case of the decay of a
mesotron except that here, because the mesotron disappears entirely,
it is necessary to postulate the emission of two neutrinos in order to
balance energy and momentum.
In free space mesotrons spontaneously decay after about two-
millionths of a second. In the presence of matter, a mesotron of
ELEMENTARY PARTICLES OF PHYSICS—ANDERSON 209
negative charge may terminate its existence in an even shorter time.
It does this by entering an atomic nucleus or, in the language of the
physicist, by undergoing nuclear capture.
The mesotrons observed in cosmic rays are produced by the very
high energy particles of the primary cosmic-ray beam as it comes into
the earth from outer space and plunges through the earth’s atmos-
phere. In a manner somewhat analogous to the creation of positrons
and electrons, the mesotrons are born out of the tremendous energies
carried by the primary cosmic-ray beam.
There are many interesting phenomena involved in the birth and
death of mesotrons and in the violent nuclear processes which accom-
pany these phenomena, but it will not be possible to discuss them here.
However, I should like to mention in this connection two important
advances which have been made within the last 2 years.
RECENT ADVANCES IN NUCLEAR RESEARCH
One of these is the work under way in Bristol, England, by Powell
and his coworkers, which has consisted of a detailed analysis of the
tracks produced by mesotrons in the emulsions of photographic plates.
These investigators have discovered a mesotron of a new type which is
heavier than the ordinary mesotron. It is about 285 times as massive
as an electron, whereas the ordinary mesotron is about 215 times as
heavy. The heavy mesotron has only a very short life; it lives only
about one one-hundredth as long as the light mesotron, after which
time it disintegrates and produces a light-weight mesotron and another
particle which is probably a neutrino. The negatively charged heavy-
weight mesotron may also directly enter an atomic nucleus and give
rise to a violent nuclear disruption.
Although both the newly discovered heavy mesotrons and the light
mesotrons discovered in 1936 have some properties in common—e. g.,
both types of particles occur with positive and negative charges, both
have short lives, and both are found in cosmic rays—nevertheless in
some very fundamental respects they are entirely different types of
elementary particles. The heavy mesotron interacts very strongly
with atomic nuclei, but the light mesotron interacts only very weakly
with atomic nuclei. Another difference lies in the respective values of
that important property known as the spin or angular momentum;
recent researches indicate that the heavy mesotron has an integral
spin, whereas the light mesotron has a half-integral spin.
In all probability it is the heavy mesotron and not the light mesotron
which is to be identified with the particle first postulated on theoretical
grounds by Yukawa in 1934. The theory of Yukawa even in its pres-
ent state today is very primitive. However, this theory still provides
the best basic concept in terms of which to understand processes in-
volving mesotrons, and after further development in the future the
210 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Yukawa theory may possibly provide an understanding in terms of
mesotron exchange forces of that all-important problem as to the
nature of the forces acting between the particles inside a nucleus. So
far no satisfactory theory has been developed in terms of which to
understand many of even the simplest phenomena involving the
nucleus. To acquire a quantitative understanding of the interactions
of the elementary particles of matter and of the fundamental nuclear
processes is one of the great tasks of theoretical physics today.
To complete our list of elementary particles we should also include
the photon. This particle, together with the neutrino as noted above,
is, however, in a somewhat different category from the other types
of particles. The photon is not a material particle, in the sense that
it cannot be identified with any particle which can exist at rest and
have associated with it a finite amount of ponderable material sub-
stance. Photons are to be identified only with radiation or radiant
energy. ‘The neutrino must also be placed in a special category, since
it cannot have associated with it an appreciable amount of ponderable
material substance if any at all, and since it has never been directly
observed.
Tn all, then, the physicist at the present time recognizes at least 10
distinct elementary particles of matter. Whether this list is complete
or not no one can say with certainty. The indications are that the
list is not complete, for evidence seems to be rapidly accumulating for
the existence of at least one additional elementary particle. This
particle is found in cosmic rays and appears to have a mass some 1,000
times the mass of the electron. But what its properties are and how
it is related to the light and heavy mesotrons and to the other elemen-
tary particles of matter is a subject which must await the results of
further observations.
The thought of probable further additions to the list of elementary
particles of matter suggests a question which is quite apart from
physics and has to do simply with the naming of new particles. We
have here actually an interesting example of the great difficulties that
physicists sometimes have merely in assigning labels or names to the
various concepts which their experience or their theories have brought
forth. It is usually necessary to choose some sort of name for these
concepts, whether they be elementary particles of matter or something
else, at a time before all the facts regarding them are known. In 1937
the term mesotron was suggested to designate the new particle of
intermediate mass discovered in the cosmic rays in 1936. Since then
this term has often been contracted to meson and has been so em-
ployed. Since the discovery of the new particle whose mass is greater
than the mass of the original cosmic-ray mesotron, the term mesotron
or meson has been employed to designate both types of particles and
the Greek-letter prefixes 7 and yp used to differentiate between them.
ELEMENTARY PARTICLES OF PHYSICS—ANDERSON 911
Thus the term r-mesotron or r-meson designates the heavier particle
and y-mesotron or p-meson designates the lighter particle. This
nomenclature seemed satisfactory for a time until continued experi-
mentation began to show more and more clearly the important basic
differences between the two types of particles. It is beginning to be
quite apparent now that the properties of these two types of particles
are such that they will not naturally fall into the same classification.
Thus the use of a common generic term, such as mesotron or meson,
to designate both these types of particles may in the future prove to
be quite inconvenient and illogical. Just what should be done with
respect to nomenclature at this time is not clear, but it is a matter
which should receive very serious consideration, especially in view
of the apparent entry of still another new elementary particle into
the fold.
Another important advance that I want to mention is the recent
success in producing mesotrons in the large cyclotron on the University
of California campus at Berkeley. This represents the first time that
it has been possible by artificial or laboratory methods to imbue a
single particle of matter with an energy sufficiently high to make pos-
sible the creation of mesotrons. This they have succeeded in doing
in Berkeley with their beam of a-particles, or helium nuclei, which
have been accelerated to an energy of 400 million electron volts.
They observed the production of both the heavy and light mesotrons,
and all indications are that the mesotrons thus produced are identical
with those previously observed among the particles produced by
the cosmic rays.
Now in the design stage are other particle-accelerating machines
which will yield particle energies several times the 400 million electron
volts so far achieved in the Berkeley cyclotron. When these machines
are in operation, working at energies up to 6 or 7 billion electron volts,
we can expect to learn much more about mesotrons and the other ele-
mentary particles of matter. Moreover, we must expect that a con-
tinuation of research in cosmic rays will also extend our knowledge
in this field, since in the cosmic rays particles are available for study
whose energies are even 10 to 100,000 times greater than those to be
expected from any of the accelerators that are being planned.
In conclusion I should like to indicate the possible significance of
these new discoveries to science and to the world at large.
In this discussion I have classified physical phenomena, according
to the energy associated with them, into three categories: (1) low-
energy or extranuclear phenomena, (2) high-energy or nuclear phe-
nomena, and (3) extremely high-energy or what we might call, for
want of a better name, elementary-particle phenomena. Knowledge
of the first of these, low-energy or extranuclear phenomena, has already
profoundly affected the life of nearly every human being on earth.
2A ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The industrial revolution, our mechanized civilization, the shrink-
ing of the world through advances in communication and transporta-
tion have all come as a direct application of our knowledge of low-
energy or extranuclear phenomena. Indirectly it has been responsible
for the political and economic organization of the whole earth. Our
present age might well be classified as an extranuclear age.
Since the explosion of the atomic bomb, and the achievement of the
release of nuclear energy on a large scale, it seems rather clear that
we are now entering a new period in which nuclear phenomena are
destined to have an important part in shaping the world, at least
politically if not economically, in the very near future. Just how
great will be the influence on the world of our knowledge of nuclear
phenomena no one can say.
It is only 50 years since our direct knowledge of the electron was
not much more than a faint green glow in a glass tube—and now no
one would deny that our knowledge of the properties of the electron
has had an effect of profound importance in shaping our civilization.
It is also only about 50 years since the world’s knowledge of nuclear
phenomena consisted of nothing more than the thoughts passing
through the mind of Becquerel as he pondered a darkened area on a
photographic plate. At present our knowledge of all these fields is
incomplete, but particularly is this true of nuclear phenomena, and
most particularly true of high-energy phenomena or the phenomena
of the elementary particles.
So far, the world’s knowledge of the phenomena of high energies
or the interactions between the elementary particles is represented by
nothing more than a few printed pages in the scientific journals, by
discussions among physicists, or perhaps by an occasional lecture.
But we can look forward with anticipation and even excitement to
the new discoveries which are surely to come as studies are carried
forward of elementary particles and very high-energy processes. New
phenomena of great beauty, extreme complexity, and novelty are
certain to be revealed and finally to be understood. Whether our
knowledge of these new phenomena will then exert a great or a small
influence on the world as a whole no one can say. I believe it would
be most unwise, however, in the light of the history of scientific
development, to expect this influence to be small.
RECENT ADVANCES IN VIRUS RESEARCH !
By WENDELL M. STANLEY
Professor of Biochemistry and Director of the Virus Laboratory
University of California
Viruses are small infectious agents that can cause disease in man,
other animals, plants and bacteria. They range in size from about
10 my, a size slightly smaller than that of certain protein molecules, in
an almost continuous spectrum of sizes up to about 300 muy, a size
slightly larger than that of certain accepted living organisms. A given
virus can multiply and cause disease only when within the cells of
certain specific living organisms. No virus has been found to repro-
duce in the absence of living cells. During multiplication viruses
occasionally change or mutate to form a new strain which in turn
causes a new disease. Viruses were not discovered until 1892 when
Iwanowski demonstrated that the causative agent of the mosaic disease
of tobacco would pass through a filter that retained all known living
organisms. Six years later Beijerinck proved that this agent was
not an ordinary living organism and recognized it as a new type of
infectious disease-producing agent—namely, a virus. The same
year Loeffler and Frosch demonstrated that foot-and-mouth disease
of cattle was caused by a virus. The discovery of the first virus
disease of man, that of yellow fever, was made in 1901 by Reed and
coworkers.
Since the original discovery of the infectious, disease-producing
agent known as tobacco mosaic virus, well over 300 different viruses
capable of causing disease in man, animals, and plants have been
discovered. Among the virus-induced diseases of man are smallpox,
yellow fever, dengue fever, poliomyelitis, certain types of encephalitis,
measles, mumps, influenza, virus pneumonia, and the common cold.
Virus diseases of animals include hog cholera, cattle plague, foot-and-
mouth disease of cattle, swamp fever of horses, equine encephalitis,
rabies, fowl pox, Newcastle disease of chickens, fowl paralysis, and
certain benign as well as malignant tumors of rabbits and mice. Plant
virus diseases include tobacco mosaic, peach yellows, aster yellows,
1 Talk presented at the Medal Day Meeting at The Franklin Institute, October 20, 1948. Reprinted
by permission from Journal of the Franklin Institute, vol. 246, No. 6, December 1948.
213
214 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
potato yellow dwarf, alfalfa mosaic, curly top of sugar beets, tomato
spotted wilt, tomato bushy stunt, corn mosaic, cucumber mosaic, and
sugarcane yellow stripe. Bacteriophages, which are agents capable
of causing the lysis of bacteria, are now regarded as viruses.
The viruses have been separated as a special group of infectious,
disease-producing agents by means of several general properties, no
one of which is, however, exclusively characteristic of viruses. Never-
theless, no great amount of difficulty has been encountered in the
segregation of the virus group. Viruses are characterized by their
small size, by their ability to reproduce or multiply when within the
living cells of a given host, by their ability to change or mutate during
multiplication, and by their inability to reproduce or grow on artificial
media or in the absence of specific living cells. The sole means of
recognizing the existence of a virus is provided by the multiplication
of the virus which is, of course, usually accompanied by manifesta-
tions of disease. Viruses spread from diseased to normal susceptible
hosts by different methods. Some are transferred by direct contact,
as when a diseased leaf is caused to rub against a healthy leaf by a
gust of wind, or when a normal person or animal comes into direct
contact with a diseased person or animal. Such viruses can usually
be spread by indirect contact through the medium of nonspecific
animate or inanimate objects. Some viruses cannot be transferred
by direct contact, but require an intermediate host such as a mosquito,
louse, or leafhopper. Im some cases a highly specific intermediate
host is necessary, and a more or less definite period of incubation
within this host may be required before the virus can be transmitted.
Because properties such as reproduction and mutation have long
been considered characteristic of living entities, viruses were, for
many years, regarded as living organisms somewhat smaller than
ordinary bacteria. However, the isolation in 1935 of tobacco mosaic
virus in the form of a crystalline nucleoprotein of unusually high
molecular weight and the subsequent isolation of still other viruses
in the form of high molecular weight nucleoproteins, some of which
were also crystallizable, cast doubt upon the validity of classifying all
viruses as organisms. With the exception of virus activity, the
properties of some of the smaller viruses are quite similar to the
properties of ordinary protein molecules, whereas at the other extreme
with respect to size, the properties of the viruses are more nearly like
those of accepted living organisms. The viruses, therefore, serve as a
bridge between the molecules of the chemist and the organisms of
the bacteriologist, and provide us with new reasons for considering
that life, as we know it, owes its existence to structure, to a specific
state of matter, and that the vital phenomenon does not occur spon-
taneously, but is possessed in varying degrees by all matter. It is
ADVANCES IN VIRUS RESEARCH—STANLEY 915
obvious that a sharp line dividing living from nonliving things can-
not be drawn and this fact serves to add fuel for discussion of the
age-old question “What is life?”
Attempts to learn something about the nature of viruses through
studies on their general properties began with Beijerinck’s work in
1898 and were continued in different laboratories for over 30 years
without too much success. Although Beijerinck and Allard made
important contributions, perhaps the most significant work was that
of Vinson and Petre during the years from 1927 to 1931 when they
showed that tobacco mosaic virus could be subjected to several kinds
of chemical manipulations without loss of virus activity. Never-
theless, in 1932 the true nature of viruses was a complete mystery. It
was not known whether they were inorganic, carbohydrate, hydro-
carbon, lipid, protein, or organismal in nature. It became necessary,
therefore, to conduct experiments which would yield information of
a definite nature. Tobacco mosaic virus was selected for these initial
experiments because it appeared to provide several unusual ad-
vantages. Large amounts of highly infectious starting material
were readily available and the virus was known to be unusually
stable. Furthermore, it was possible to titrate or measure the amount
of this virus in a preparation with ease and rapidity and with great
accuracy. During the course of a wide variety of early exploratory
experiments, it was found that the enzyme pepsin inactivated tobacco
mosaic virus only under conditions under which pepsin is active as a
proteolytic agent. It was concluded that tobacco mosaic virus was
a protein or very closely associated with a protein which could be
hydrolyzed by pepsin. With this as a lead, efforts were made to
concentrate and purify tobacco mosaic virus by means of the methods
previously employed in work with proteins. Soon, by means of a
combination of procedures involving salting-out, isoelectric precipita-
tion and adsorption on and elution from an inert material, a crystal-
line material was obtained which possessed the properties of tobacco
mosaic virus. This crystalline material was found to be a nucleo-
protein with rod-shaped molecules or particles about 280 by 15my in size
and with a molecular weight of about 40,000,000. Early skepticism
that a virus could exist in the form of a crystallizable nucleoprotein
has largely disappeared, chiefly because the results of a vast amount
of experimental work have indicated that the virus activity is a
specific property of the rod-shaped nucleoprotein.
Tobacco mosaic virus exists in the form of many strains which
appear to have arisen by a process similar to that of mutation in
higher organisms. Several of these strains have been obtained in
purified form by means of differential centrifugation. Purified prep-
arations obtained from plants diseased with different strains of
866591—50-——15
216 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
tobacco mosaic virus were found to possess properties quite similar to,
yet in every case distinctive from, those of purified preparations of the
ordinary strain. Spectacular progress has been made in the establish-
ment of the nature of the chemical changes which accompany the
mutation of tobacco mosaic virus. The amino acid composition of
purified preparations of eight strains of tobacco mosaic virus and of
two types of influenza virus has been determined. The results ob-
tained with the strains of tobacco mosaic virus indicate that the
mutation of a virus can be accompanied by the elimination of one or
more amino acids from the virus structure, by the introduction of one
or more new amino acids into the virus structure, or by a change in the
concentration of one or more amino acids present in the virus structure.
This work has great significance for it has provided the first informa-
tion regarding the nature of the structural changes which accompany
mutation. Extension of this work may reveal the exact nature of the
chemical differences between virulent and avirulent virus strains, and
provide important information regarding the mutation process in
higher organisms.
Attempts have been made to change the structure of tobacco mosaic
virus by means of known chemical reactions in vitro in an effort to se-
cure chemically modified active virus. Although several types of chem-
ical derivatives of this virus were produced and were found to possess
full virus activity, the inoculation of such virus derivatives to normal
Turkish tobacco plants always resulted in the production of ordinary
tobacco mosaic virus. ‘The results indicated that the chemical deriva-
tives were converted to ordinary virus following their introduction into
the cells of the plant, or, more probably, that the infecting molecules
may not necessarily function as exact patterns for reproduction. De-
spite these results it still appears that it may be possible to make
changes in vitro similar to those which occur in nature, and thus secure
a heritable chemical modification. Obviously this is a field in which
important new results can be anticipated.
Following the isolation of tobacco mosaic virus in the form of a
crystalline nucleoprotein having individual molecules or particles about
15 by 280 my in size, studies were undertaken in several laboratories to
determine if other viruses could be obtained in purified form, mainly by
techniques involving high-speed centrifugation. Some of these puri-
fied viruses are crystallizable nucleoproteins having either rodlike or
spherical particles. Some are nucleoproteins which have, as yet, not
been crystallized. Others are large particles consisting of nucleo-
protein, lipid, and carbohydrate, and possessing, in some cases, a
degree of morphological differentiation which resembles that of organ-
isms. Still other viruses have, as yet, defied isolation and purification,
possibly, in some cases, because of extreme instability. The viruses
ADVANCES IN VIRUS RESEARCH—STANLEY Pail
which have been purified possess varied shapes and form an almost
continuous spectrum of sizes. The smaller rod or spherically shaped
viruses appear to be simple nucleoproteins, some of which can be
obtained in crystalline form. These appear to have chemical and
physical properties which, neglecting virus activity, would tend to
place them in the molecular world. The larger viruses have a com-
position and properties which are characteristic, not of molecules, but
of organisms. The viruses have certainly provided a link between the
molecules of the chemist and the organisms of the biologist. Yet
there is no place at which a line can be drawn dividing the molecules
from the organisms.
The viruses appear to form a continuous series with respect to
structure, ranging from the smaller viruses, which are simple nucleo-
proteins with many properties similar to those of ordinary molecules,
on through viruses with a gradually increasing complexity of structure,
to the larger viruses, which, with respect to structure and properties,
are similar in many respects to organisms. However, it must be re-
membered that the properties of only a relatively few purified viruses
have been determined. In view of the possibility that these represent
the more stable and more easily purified viruses, one cannot be certain
that a true picture of the chemical and physical properties of viruses as
a whole has been obtained as yet. Information regarding the mode of
reproduction of viruses is needed most urgently. At present it is not
known whether viruses reproduce by fission or by means of some new
process. The solution of this puzzle would certainly represent a most
important and significant advance, for the basic reactions character-
istic of virus reproduction may well represent the fundamental process
which characterizes all living things.
GROUND-WATER INVESTIGATIONS IN THE
UNITED STATES!
By A. N. Sayre
Geologist in Charge, Ground Water Branch
Water Resources Division, U. S. Geological Survey
Before discussing ground-water investigations in the United States
I should like to outline briefly the broad problems of water supply,
water control, and conservation, in the solution of which ground-
water investigations play an important part.
For mere existence, a man requires only about 2 quarts of water
per day. However, even in simple pastoral or agricultural settings,
the biological necessity represents only a part of the total needs for
water, and in our own complex industrial and agricultural economy
great quantities of water are needed for a multitude of purposes.
Water is needed for sanitation, for washing clothes, for facilitating
sewage disposal, for scrubbing floors, and for processing foods. It is
needed for fire protection, for generating power, and for industrial
processes, for irrigation, for air conditioning, and even for producing
atomic energy. The task of providing water at the right time and
place is a serious problem which fully occupies the attention of
thousands of engineers and chemists and a smaller number of geolo-
gists. Intimately associated with the water-supply problem is the
problem of controlling floods to minimize the erosion of our soils and to
conserve floodwater for beneficial uses. For obvious reasons, many
of our agricultural, industrial, and urban developments have taken
place along waterways where they are vulnerable to the ravages
of floods which appear to become more costly almost year by year.
There is good reason to believe that the demand for water supply will
continue to increase and that the demand for control and conservation
of floodwaters will also increase. Projects for accomplishing these
ends will become more expensive and their planning and design will
require greater knowledge of our water resources and of the basic
geologic and hydrologic factors affecting them.
1 Presented at the joint meeting of the Society of Economic Geologists and Geological Society of America,
Ottawa, Canada, December 1947. Reprinted by permission from Economic Geology, vol. 43, No.7, Novem-
ber 1948.
219
220 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Especially during the past century, the use of water has increased
at an amazing rate. In 1850 only 83 cities in the United States had
public water supplies, and only a small proportion of the homes in
these cities had water piped directly from the city mains. By 1939
there were 12,760 ? municipal waterworks and thousands of industries
had private supplies from wells or surface-water sources. As the
number of waterworks increased, the uses of water and the per-
capita consumption also increased. The quantities of water used
for a few purposes are given below. Flush toilets, bathing and
laundry, street cleaning, and fire protection require an average of
about 40 to 75 gallons per day per capita.* Processing a ton of steel
in highly finished form requires about 65,000 gallons* of water;
making a gallon of gasoline takes 7 to 10 gallons® of water. Vast
quantities of water are used for air conditioning and for making paper,
explosives, coke, textiles, and a host of other products. Thus, a large
city, such as Chicago, with numerous industries may have a per-capita
water consumption as high as 250 to 300 gallons per day.
Only a few thousand acres in the West was irrigated in 1850, but
21 million acres ® was irrigated in 1939, and many additional irrigation
projects are under construction. An acre of cotton uses about 2.57
acre-feet, or 800,000 gallons, of water during the growing season; an
acre of alfalfa requires about 4 acre-feet of water; irrigation of truck
gardens, fruits, sugarcane, rice, and other crops also requires large
amounts of water. In eastern United States, supplemental irrigation
is increasing because the application of a relatively small amount of
water when it is needed by crops may double or triple the yield of
the land.
The greatly increased use of water has, in many places, almost fully
utilized the readily available water supplies, drawn ground-water
levels dangerously low, caused sea water to enter streams and ground-
water reservoirs in coastal areas, and permitted oil-well brines or
factory wastes to pollute many of our ground-water reservoirs and
streams. Thus, the development of additional water supplies for
new projects or industries has become increasingly difficult and costly.
Nevertheless, it would be a mistake to infer that our water supplies
are approaching exhaustion. Actually, much can be done to conserve
and thereby increase the total amount of water available for beneficial
use. For example, in many places spacing pumped wells over wider
2 Engineering News-Record, vol. 123, p. 414, 1939.
3 Turneaure, F. E., and Russell, H. L., Public water supplies, 4th ed., p. 19. John Wiley & Sons, New
York, 1940.
«Lloyd, Kenneth M., Industry and water supply in Ohio. Ohio State Univ. Exper. Stat. News, p. 31,
April 1946.
‘ Jordan, Harry E., Industrial requirements of water. Amer. Water Works Assoc. Journ., vol. 38, pp.
65-68, 1946.
6 Census Bureau.
’ National Resources Committee, Regional Planning, pt. 6, p. 91, 1938.
GROUND-WATER INVESTIGATIONS—-SAYRE 221
areas would prevent excessive lowering of the ground-water levels.
Artificial recharge of ground-water reservoirs by spreading flood-
waters and by other means has been successful in several areas.
Abatement of pollution in streams and ground-water reservoirs,
retention of floodwaters in reservoirs for later use, control of reservoir
stages by forecasting normal and flood flows of streams, control of
silt and sedimentation, and other measures are being carried out to
increase the supply available for perennial beneficial use. The
continued growth and prosperity of the Nation will depend to a large
degree upon the success with which these problems are attacked and
solved.
Unlike most mineral resources, water is not exhaustible, in the strict
sense, because it is replenished from time to time by precipitation. A
surface reservoir may be dangerously low and be refilled in the nick
of time by heavy rains. Heavy pumping may cause ground-water
levels to decline progressively until pumping is no longer economically
feasible, but when pumping is temporarily or permanently reduced
the water levels usually recover. Likewise, a period of heavy rainfall
following a drought period may induce recharge sufficient to restore
water levels essentially to predrought levels.
Only a part of the water taken from streams or pumped from wells
is actually consumed. In many manufacturing processes water is
merely a washing agent and is essentially unreduced in volume by
its use. Water used in boilers or in quenching hot metal is partly
evaporated, but the remainder is discharged or re-used. Water used
in irrigation is partly evaporated and partly transpired by plants, but
there is always an excess which is discharged and which carries away
undesirable salts. The excess water from all these uses returns to
the stream or to the ground altered by the concentration of minerals
contained in it, or by the addition of dissolved constituents, or of
color, or sediment, or simply by the addition of heat. It may be
re-used for the same or other purposes with or without the addition
of new water. For example, the water of the Pecos River, in Texas
and New Mexico, is used and re-used for irrigation and domestic
supply seven or eight times between its source and Girvin, Tex. Al-
though water is added from tributary areas along its course, each
time the water from the river is used the mineral concentration
increases, and a few miles above Girvin it is so highly mineralized
that even the most resistant crops are unable to survive its application.
Even so, it may still have potential use, because water also possesses
the energy of position and in its journey from the mountains to the
sea it may be used many times over for generating hydroelectric
power.
Another characteristic peculiar to water results largely from the
vagaries of precipitation. Many places are faced successively with
Dae, ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
water shortages and destructive floods. Although enormous sums
have been spent on flood control, complete protection from floods is
difficult and often prohibitively costly, and in only a few places has
it been accomplished.
Whatever conclusions are reached with regard to the economics of
flood control by storage reservoirs, from the standpoint of conservation
of water supply it is advantageous to salvage as much of the floodwater
as possible. In certain areas where water supply is now inadequate,
as it is in many places west of the 100th meridian, much of the water
from precipitation escapes to the sea during floods. In Los Angeles
County, Calif., much of the floodwater is retained in surface reservoirs
from which it is later discharged into specially prepared recharge
basins and seeps into the underground aquifers. Some of the flood-
water is caught in seepage reservoirs designed to promote infiltration
into the ground-water basins, and some of it escapes to the sea, but
essentially all the floodwater in the Los Angeles area will be put to
beneficial use when the projects now under consideration are com-
pleted. Plans for similar flood control in other parts of the country
are in various stages of execution.
Ground water and surface water are so intimately related that for
proper solution of the over-all problems of water supply, control, and
conservation it is now necessary to have all the facts regarding both.
Precipitation, which is the source of both, is partly lost, largely through
evaporation. Especially during the growing season, a large part
enters the ground and is transpired by plants. The remainder per-
colates downward below the plant roots to become ground water, or
runs off directly as surface flow. The ground water, returning to the
surface as seeps or springs, provides the base flow of the streams which
prevails through periods of low precipitation, On the other hand,
especially in the West, many streams lose water by seepage in certain
stretches and thus recharge the ground-water reservoirs.
Many of the basic ground-water investigations in the United States
are carried on cooperatively by the United States Geological Survey
and State or local agencies, including State geological surveys, State
engineers, counties, and municipalities. In Illinois ground-water
investigations are made by the State Geological Survey and the State
Water Survey; and in Missouri investigations are made by the State
Geological Survey. In California a large staff of engineers in the
State Division of Water Resources for many years has been investi-
gating overdraft of ground-water supplies. Recently arrangements
were made whereby the United States Geological Survey, in addition
to its investigations in Los Angeles, Orange, and Santa Barbara
Counties, will assist the Division of Water Resources in the geological
phases of a State-wide inventory of the water resources of California.
GROUND-WATER INVESTIGATIONS—SAYRE 223
Various other Federal and State agencies are obtaining some data on
ground water in connection with special phases of their work.
The ground-water investigations of the United States Geological
Survey began more than 50 years ago. At that time ground-water
supplies were little developed. Consequently, most of the early field
investigations were of the exploratory type. Laboratory and field
studies by King, Slichter, and later by Meinzer outlined the broad
principles of ground-water occurrence and movement. However,
before the deep-well turbine pump was developed in the early part of
this century, use of ground water in large quantities was limited to
areas of springs or artesian flow, or to areas where the water level was
within reach of suction pumps. After the turbine pump was intro-
duced, it became possible to pump economically even where water
levels are deep; power costs also decreased, and well drilling and finish-
ing methods were improved to increase the efficiency of wells. Because
of these advances, ground water came to be used in ever-increasing
quantities, first in areas where surface water was not readily available,
and later in areas where ground-water supplies were more economical
because they obviated the long pipe lines, collection works, and costly
treating plants needed for surface-water supplies. ‘The first great
expansion of ground-water supplies was made largely without technical
guidance. Because of their immense storage capacity, the ground-
water reservoirs were regarded as inexhaustible, and in many places
development was well advanced before progressively declining water
levels brought about the realization that ground-water reservoirs may
be depleted. As a result, demands for detailed ground-water studies
steadily increased. These studies are thorough, systematic investiga-
tions which include areal geologic mapping; subsurface studies occa-
sionally augmented by geophysical surveys and test drilling to deter-
mine the structure, thickness, and the sequence of water-bearing and
non-water-bearing beds; collecting data on fluctuations of water levels
in relation to precipitation and to pumpage; determining the recharge
and perennial yield; inventories of ground-water withdrawal; test
pumping to determine coefficients of permeability, transmissibility,
and storage; determining the quality and temperature relationships,
and the relations between surface and ground water. The work does
not include supervision, construction, or control of water supplies.
The United States Geological Survey is now making cooperative
ground-water investigations in 42 States and in Alaska, Hawaii, the
Virgin Islands, and Puerto Rico. In general, the State or local
authorities are most familiar with the needs in their States, and they
are largely responsible for designating the areas in which investigations
are to be made. Most of the ground-water staff of the United States
Geological Survey now have headquarters in field offices, of which
there are about 40.
B24. ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
In addition to the cooperative investigations with States and local
agencies, certain investigations primarily of Federal interest are being
carried out with use of Federal funds only. For example, an extensive
program of ground-water investigations is being conducted in the
Missouri River Basin to provide basic information needed for planning
and constructing the various water projects that are authorized or
planned. This information will include determination of existing
ground-water conditions and probable effect of irrigation on ground-
water levels, especially with reference to waterlogging and drainage,
canal locations with reference to seepage, areas where irrigation with
ground water is feasible, the location of potential sources for farmstead
and municipal water supplies, and so on. Other investigations, such
as that recently started in the Central Valley of California, will provide
basic information needed in the project to use excess floodwaters for
recharging the heavily pumped ground-water reservoirs in the San
Joaquin portion of the valley. Several projects also are under way
in connection with defense plans.
A large program of measurement of ground-water levels in observa-
tion wells has long been an integral part of the cooperative program.
Recently a small Federal fund was provided for extending this pro-
eram, analyzing the data, and determining the current status of our
ground-water supplies on a Nation-wide scale. The program includes
the investigation of the possibilities of forecasting low-stage stream
flows from fluctuations of ground-water levels, enabling more efficient
control of reservoir and river stages in the operation of hydroelectric
plants and permitting considerable economies in the generation of
power. Conversely, base flow is an index of ground-water storage.
As the ability of the ground-water reservoirs to absorb water ma-
terially affects the runoff to be expected from rainfall, the analysis
of data on ground-water levels should aid in flood forecasting.
In addition to and as part of the above program, several research
projects are under way, including the study of movement of water
through soils and water-bearing materials; infiltration from streams
and its effect on the temperature and quality of the ground water;
the occurrence of ground water in fractures and solutional openings
in impermeable rocks, such as limestone, tightly cemented sandstone,
granite, etc., and the improvement of our techniques for locating
producing wells in such rocks; earth subsidence resulting from the
withdrawal of ground water, and related problems connected with
elasticity and compressibility of artesian aquifers; methods of analyz-
ing results of pumping tests to determine coefficients of transmis-
sibility and storage; and a number of other projects.
With respect to the probable future of ground-water investigations,
it should be pointed out that ground-water development was expand-
GROUND-WATER INVESTIGATIONS—SAYRE 225
ing rapidly in the middle thirties. In 1935 the total use of ground
water in the United States amounted to about 10 billion gallons a day.
The development was greatly accelerated by the needs of the war,
so that by 1945 the total pumpage had nearly doubled.2 Although
the war ended more than 2 years ago there has been no sign of a
decrease in the use of ground water. In fact, both surface water and
ground water are now being used in greater quantities than ever
before. As the use of water approaches ever more closely the limits
of the available supply, it is believed that water-resources investiga-
tions will be needed with ever-increasing urgency because water
constitutes the prime factor in the continued development of the
Nation’s industrial and agricultural economy.
§ Guyton, W. F., Industrial use of ground water in the United States. Abstract, Journ. Washington
Acad. Sci., vol. 39, No. 3, pp. 105-106, Mar. 15, 1949. (To be published in Proc. Geol. Soc. Amer.)
es ‘ gations ban tat ipiecepyiieaseeis dur ai
: once ange. ‘ aaly Ampust ra yey &
ioe Gey efida-letiie Py aia oa:
Aes #1) dl orang heh hae, |
aa ane ON, ue
fra Bia neva, Tih yt
10% 0, ast) ssa,
ai
a + hosts
n Sousa a
har wiih, af te
PY,
A of nid thy guys $ rhs
ia oldiol ohh to
tbonit,od, leg exit)
9 SFE finger thrton ay, oe ‘ies sank ARG, out “olga,
pO CO lark sthen a has eo tay at)
Sif wi j 4 “| i
ev iccuvans AeecT Oper) ay, i
iy yibitan? eifal Ai nent uy oat aah iy Hitt pF mide Haid i) ni Fat yhhgabh lt tae = 45
Vols CMTC. Cars bog) Jet ADSL od cad ED RAD rh Male, OE ae nit Bj deh hepa.
- J wie in u
i ad ’ ‘¥ v4
; |
i { 7 Cd tae
14
»
. 7 5 iw Ay ; Ta ; i ¢ et me yah, -
Rnisuly off Vodeanl PRC Kimi protein) | + Resa geet
A t , . 5 oy :
: . . ut Vi 4G CT I
; ‘iaT) uel ages, o Hy
LF f bp » Me i vs cf f LA ley , ae cine ;
ye
mn fh) F ju mind ike me { rar ie GSA Bal | ; ute ice’ i ;
me ~ AGA thie? pla in Die 61 yl news a
, j
pes Made : ; ref a tReeT yA a Gite (pee whi. wt, ray
950 pap its wiv, taee: {10 : pPerviady are helena q
As fine! rd a le 4 y yHL’s ede At vy WL } i ‘ an a
eT ai Lay Pye # ig fs fon fA ? a) ae aealivuaty ; ,
-~ A P : j f
as : - :
s , ag ‘i « ?
ik ey) Cheer WAY, Gens Gone may! fh, ary
i
a
= ——
eet
dell As eee? an Ute heb ete
os |
m
2
>
=
_
re NOCCS
sSitesasd von ) i es
ei,
iy = pasoe|as.——
uebosin—
_ 4)
ttl
= areas ese etis sense NWoenad fetete tanetes spear amma ee ll
: z Te uoljos0doaz
uolsosa jowsou : ee tile i
puo jjo-uny eeoeds
f — 0; usnjas sfosauiw | v ¥ y V
Puo Jayjow aiu0bsD wroy
|
Jajjow diuDbs0 | |
ezisayjuds
SJUDId
@DIxo|p uoqso9 uol{osidsuosy uegboj4in
230 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
materials, soil science is not easily classified as a science in the tradi-
tional patterns. Often it is included with the physical sciences, like
chemistry and physics. Yet it is also grouped as correctly with the
biological sciences, along with plant physiology and bacteriology.
Then again, soil science is appropriately grouped with geology and
geography as an earth science. Actually soil science uses the prin-
ciples and methods of all three of these groups, and a soil scientist
who uses the principles of only one or two of the groups, to the exclu-
sion of the others, can only have small principles that have little
prediction value and soon bog down in contradiction. In addition
to those already mentioned, there are important principles and
methods peculiar to soil science itself that do not belong to any other
science. In application, the principles of soil science must be inti-
mately related to those of the social sciences.
It needs to be emphasized that the experimental method and the
method of scientific correlation are essential in both fundamental
soil science and in the application of its principles to practical prob-
lems in farming, gardening, and forestry.
Soils must be studied in relation to one another and to the whole
environment, both natural and cultural, to understand their forma-
tion and the influence of the individual factors of climate, vegetation,
parent rock, relief, and time. How any one of these factors operates
depends upon the others. The significance of any one soil charac-
teristic depends upon the others. Any soil is a combination of
characteristics, produced by a combination of factors, each of which
influences the functioning of the others.
Experiments are needed, both natural and artificial, to learn how
individual soils behave and how they respond to treatment. These
must be specifically related to individual kinds of soil, however, if the
results are to be used in developing principles or as the bases for
practical predictions. Thus the experimental methods and the
methods of scientific correlation are intimately interwoven in produc-
tive research in soil science.
SOIL AND LANDSCAPE
Let us look briefly at the implications of this concept of soils. They
are natural bodies, each with its own unique morphology; they are
dynamic bodies, developing with the natural landscape itself; they ac-
curately reflect, at any moment, the combined or synthetic influence of
the living matter and climate, acting upon the parent rock through
processes conditioned by relief, over a period of time; they are dis-
tributed over the earth according to orderly discoverable and definable
geographic principles.
The geological process of mountain building, rock formation, and
landscape evolution from which the parent materials of soils originate,
MODERN SOIL SCIENCE—KELLOGG 25K
are still going on along with soil formation. The natural erosion of the
uplands gradually removes a little of the surface bit by bit while the
soil film settles down, and fresh minerals are added to the soil from
beneath. With the warping of the landscape these processes are
accelerated or retarded.
At several of the Soil Conservation Experiment Stations of the
United States Department of Agriculture, rates of erosion were deter-
mined under permanent vegetation (15).2 Under the natural forest
Exchangeable Cations (M.E,/100gms.) Exchangeable Cations (M.E./100gms.)
pH 10 20 30 40 pH
5.1
4.8
49
4.4
Si
6.2
$.6
B. From granite
a ed,
Percentage of Clay
A. From diabase
Fiaurs 2.—A comparison of the clay content and exchangeable bases from two
soils developed in similar environments, except for the difference in parent
rock. The fundamentally important differences in clay content and in ex-
changeable bases are obvious between the soils and among horizons within
each soil (9).
cover of the Cecil soil of the Piedmont, erosion proceeds at a rate of
about 1 foot in 10,000 years. This value was determined on a 10-
percent slope. Yet on a 14-percent slope of the Muskingum silt loam
of Ohio, considerably over 200,000 years would be required to remove
1 foot by erosion under the forest cover.
Under a well-established grass cover, normal erosion proceeds
slowly on the dark-colored soils developed under tall prairie grasses.
On the Marshall silt loam near the Nebraska-Iowa line, with 9-per-
cent slope, nearly 14,000 years would be required to remove 1 foot
under bluegrass. On a black soil of east Texas, Austin clay with a
slope of 4 percent, the figure is nearly 900,000 years.
3Numbers in parentheses refer to literature cited at the end of this article.
866591—50-——_16
Zaz ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
These values probably do not represent the real extremes, but rather
relate to normal erosion. Some soils erode much more rapidly under
clean cultivation with bad effects upon soil productivity. Through
proper cropping systems and soil management practices, erosion of
soil under use should be kept somewhere near the normal rate.
Organic Organic
Motter(Z) pH SOW ue e100 0) 50 100. ~— PH Matter()
eth lie 3 posse TMT | Depth Pe l ae 55 6.2
Serere, iO ¥ 1
oso SEMIN ETM ss
> (]
ZOmennS'5 seeeene yA the ise a
on a
< Qj } Ie
os Femme aN
O
20" — Rey 6 4
lo 82 SN eee
Tl, 50:7.
nt 64 se : Sees
ieee Gee
7.8 08
HO. 7:6
cle?
oO res :
RRO ! A. 40”
PGR CERI Rt iat is)
A. Miami Stit Loam-
Undulating to Rolling
B. Bethel Silt Loam- Flat
Ficur& 3.—A comparison of the mechanical composition of two soils developed
from similar parent materials and in similar environments, except for relief.
On soils of undulating to gently rolling relief there is continually a small
amount of erosion in the natural landscape. As the surface is thus slowly
eroded, each soil horizon works down into the one beneath. Over a period of
several thousand years the whole soil profile, while remaining at the same
length, may sink into the landscape a foot or more. Thus the soil is kept
constantly renewed with new minerals. This is illustrated by the Miami silt
loam at the left. The soil at the right is developed from similar material on
flat upland where there is little or no erosion in the natural landscape. The
leached material accumulates at the surface, and clay formation and the accu-
mulation of clay are accentuated in the middle portion of the profile, with the
development of a claypan. Whereas the Miami silt loam is a well-drained soil,
pervious to roots and water, the Bethel is an imperfectly drained soil. During
wet periods excess water is held up by the claypan which is also impenetrahle
to mast roots. Kinds of crops and soil-management practices for optimum
production are quite different indeed, even though the soils lie side by side on
the same farm (2).
Percolating water gradually dissolves the minerals in the soil and the
rock beneath. Clarke has said that this process alone reduces the
surface of the United States on an average of about a foot in 30,000
years (6). From a study of streams, Dale and Stabler (7) estimated
MODERN SOIL SCIENCE—KELLOGG 233
several years ago that the average rate of denudation for the United
States as a whole was about 1 foot in 8,760 years, with suspended
matter accounting for about 65 percent and dissolved matter 35 per-
cent. Solution progresses much more rapidly than this in areas of
soft limestone and high rainfall, and, of course, the process is almost
infinitely slow on the hard rocks of steep slopes.
Other landscapes receive part of the erosion products and part of
the solution products from the upland. It is probable that a third of
the population of the world get their major food supply from alluvial
souls recently rejuvenated by additions of fresh rock minerals to their
surface. The Nile is a famous example. In flood stage the water of
the Nile contains over 1,000 parts per million of suspended matter
relatively rich in phosphorus, potassium, nitrogen, and other plant
nutrients. Part of this covers the soil in the flood plain and part of it
moves out into the sea. Barrell (1) estimated that the Nile Delta
alone contains the equivalent of nearly 12,000 cubic miles of rock, to
say nothing of the soluble material contributed to the sea water.
According to Barrell’s figures the rock material in the delta of the
Niger River is equivalent to a wedge-shaped mountain range some
18 miles wide at the base, 3 miles high at the top, and 1,000 miles long.
This gives some idea of the enormous movement of surface soil
material as a natural process. In addition, in the Tropics especially,
volcanoes often shower the landscape with fresh rock or ash. mates
‘ LE
pee Af: Le
Z Ses
ELE
row
ae SA
The areas of each great soil group shown
on the map include areas of other groups too
small to be shown separately Especially are
there small areas of the azonal and intrazonal
§roups included in the areas of zonal groups
AZONAL
Soils without well-developed soi! characteristics. (Many areas
of these soils are included with other groups on the map.)
LITHOSOLS AND SHALLOW SOILS
oe ll
Fo O46 F O))
a Shallow soils consisting largely of an imperfectly
weathered mass of rock fragments, largely but
ee | not exclusively on steep slopes.
Resear.
RON BF
SANDS (ORY)
Very sandy soils.
ALLUVIAL SOILS
Soils developing from recently deposited alluvium
that have hadlittle or no modification by pro =
cesses of soil formation.
EK
866591—50 (Face page 236)
Thad.
VTE
Yiya®
'
“ate a? Hind “Too5
_
S
a Bypoe runtesid
to We Hind - #100)
abbateenry td eaten etree
ale
Ss: paca eal Sean kee:
<7 a. etn
AR ecmeiieaall
oa lie LEG
»
ontardignns. shay tanita 'o equrg tev)
Fenpfeipen Fh alae Ti YO sone tes prvlenemah act yetael
‘ey byes are Nheere eur gem ett ro peo 24) >
(dabviow ne ele
pew ate to shoe dares! beroind yi.t
—_— soto
39 m 7 2102 Anime» HeIOo aR
5 oe dey r Dun 2 hae Memo Geo ber gra
‘ tea uve bo baw eneiey unsinse bamud e604 matteo \e chee berioeg! award
eee ieGupers a cb cath: abr te ef - aipay “oooge balesrot N
Wros 691. cy al on t ;
oe . ‘ ’ pa trem wedt f pees WO tt = pg BOR. A) 205908, "wos
le eR sienqnere # Ret nto iG ES ai wena hee ahion bed2g-ab ndings > aS
= SESE Fue ve pred | eo Sng ‘betewwt = :
Obwo ahs =—~ ¥ ~~ : mn Peg : cence poi MALLOW §O Peat A
“oe carers " % whe Meine as s2gi2:
we > Dart cout davteheniwbaeo oe A brains Sait =f seman mot eee te “1 il
' ya ‘ yes sel :
aTRy iow
p08 nWwORE eee
“via {8 Bice Mwond- Aaibsy tijd 10 owentl
orb bron OMe, epee Di MONTIEL: tae toa, hq ngamet
veasge ry ae ews POD, ae wt tan oakue
| peed year lateats ral THREES RO me Daze”
; ‘ho ert andigey Dea etereemsl ot looo to aioe Sow fs
. Qu rere 4 date Pao te ce + AChR Oem Gunde 1h ww D
é Ss. +
OPTS agg Cus Nah
+ dati iden tegavor'l
-
——
—
z 7 pe Ke Sa fibmdt - ub) a A we apa Gant tigu
¥ S wwgete bm
) ,enngsy Bie lon ©
*
5 Mey wae
- ira ii o
*
fart waste 7] bbe gry Hear hy
,
cork Wag na
ay a aly p digts.
ee to aloe eword J16d
ebrwieakyy bend (nin
ao seh6 ren “BMW Vo bice owned de
t, har get
i
boar»
abnaieeerg bint ris!
Les
ign +t pte
ts ” seta
pa
44
MODERN SOIL SCIENCE—KELLOGG De Th
causes of soil characteristics, because of failure to see the common
characteristics of unlike soils found in the same ecological region.
In a broader study one might reach altogether different conclusions.
Suppose, for example, that one studies in detail the upland soils on
smooth slopes from limestone and sandstone in West Virginia in con-
trast to those on smooth slopes from limestone and sandstone in
western North Dakota and in southern Arizona. Here again, striking
differences are apparent, but the most important of these relate to
differences in climate and vegetation; geology would seem relatively
unimportant.
Thus by grouping the local soil types into higher categories, the
broad soil groups that dominate the landscapes of great regions are
compiled. When one considers individual practices in the garden, on
the farm, or in the forest, it is the local landscape and the local soil
types that are important. When one considers the great movements
of population, the potentialities of nations, and the historical trends
of peoples, the significant factors are the great soil groups. Thus
the gardener sees soil characteristics with a different emphasis than
does the geographer, but soil science has much to contribute to both.
PRIMITIVE SOCIETIES ON THE SOIL
The soil supports plants, animals, and man himself. Primitive man
must have been as much the helpless product of his environment as
were the wild animals. He lived close to the soil and was a food
gatherer. He took the plants and animals, including fish, that were
available in his own landscape. Most of these were eaten with little
change by cooking, storage, or refining. No doubt it was a risky busi-
ness. Families might fail to get food because of drought, deep snows,
wars, or other calamities, and starve. However, when things went
well, although they might live on a few foods for a time, during the
year there was usually a variety. Then too, primitive man ate vigor-
ously. He ate whole foods—skins, hulls, seeds, and other parts that
are now thrown away. It is only recently that modern man has
attempted to create a balanced diet. Early man received or failed to
receive his proteins, minerals, and vitamins, unconsciously. Of course,
no scientist was available to study our savage ancestors at the time,
but many studies have been made of relatively primitive peoples and
of the physical degeneration caused by the substitution for native
whole foods, like cereal grains, milk, cheese, fruits, and meat, of refined
sugar, white flour, and similar products of more ‘‘advanced”’ societies
(12). Thus native peoples may become degenerated, or even extinct,
after contact with Western Europeans. ‘There are reasons to suggest
that people, like plants and animals, over a long period come to an
adjustment with their food supply (11).
938 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Marett held that people having food deficient in some essential ele-
ment, say calcium, phosphorus, or iodine, gradually develop the ability
to conserve this element. That is, in the evolutionary process the
ability to get along satisfactorily with only a little of some element
would have survival value. Such people would be most likely to have
children able to thrive and grow strong. But in the process, size,
skin color, and other features, even the psychological characteristics
and social traits, are altered. Marett believed that differences in food
composition, most of which were closely related to the local environ-
ment before the modern period, had a great deal to do with the origin
of races, with the physical and social differences among the different
peoples of the earth.
Marett argued, for example, that in regions with acid soils deficient
in calcium, people of small size would be favored. The physiological
strain of lactation upon females would be much greater in humid
regions where soils are generally leached and acid and the foods de-
ficient in calcium and phosphorus, than in arid regions where soils
usually contain abundant lime (calcium). Thus on acid soils where
food is deficient in calcium the bodily strain would lead to adjustments
for economizing lime through decreased size, especially of bones. He
suggested that the operation of these forces may have been important
in the development of fine bones among modern people as contrasted
to our more coarse-boned ancestors. Such changes are very gradual,
and are not marked until after long-living in a particular landscape.
MAN AS A CULTIVATOR
As civilization developed, man became a cultivator. He began to
direct the course of nature toward his own ends, and ceased to be
simply a food gatherer dependent only upon the natural bounty of
the landscape. He ceased to be primarily a thief, and became a
grower, 2 homemaker, a planner, and a conservationist in the only
sense the term has any social meaning. As he gained in experience
he learned to satisfy himself more easily; in fact, some people in the
society could cease to be food gatherers. Social structures rose with
the evolution of trades and professions. As the efficiency of food
production increased, more and more people could be released from
food gathering to develop the arts and sciences, to make the other
things man needed for his health and comfort, and, unfortunately,
to make war.
From the dawn of history to the rise of modern science the accumu-
lation of learning about agriculture was a terribly slow process.
Experience, which was passed down from father to son over the genera-
tions, was the only guide. Only a few departures were made, because
there was no substitute for such experience. Further, there was little
MODERN SOIL SCIENCE—KELLOGG 239
realization that experience on one soil, in one landscape, could not be
relied upon where another soil was involved. Migrations were often
disastrous for this reason. Then, too, world history records changes
in the landscape under the very feet of the farmer, like the slow spread
of the Sahara Desert as it gradually expands to its former position,
following the moist period of glacial times (8).
Unfortunately, the early scientists of Greece and Rome reached
little into the problems of agriculture. With a few conspicuous excep-
tions, the philosophers of that day accepted farming as the job of
slaves, beneath the dignity of trained scholarship.
THE GREAT DISCOVERIES
The tempo of man’s struggle with his environment completely
changed with two great forces: the rise of modern science and the great
discoveries. The most important fact of Western culture was the
opening of new land in the world. The forces leading to the pessi-
mism of Malthus were already destroyed before his famous essay on
population had been printed. Science began to increase productive
efficiency. Western Europeans found new homes in the landscapes
of the Americas.
Europe had been bound by an aristocracy based upon land. Al-
though many came to the new world to seek gold and adventure,
most people came to find land and to build homes on the only security
they knew. Gradually the east coasts of the new world filled up. In
the beginning people were confined to land near the sea and to navi-
gable waters, as they had been in the centuries before. But modern
science came to the aid of discovery. The European colonists pushed
into the interior, especially in North America. Railroads had made
possible the exploitation of interiors of continents, of the great areas
of black soils. Except for a few isolated spots, these soils were scarcely
used by civilized folks at the time of the Treaty of Westphalia when
modern nationalism had its birth. During the nineteenth century
the black soils (Chernozem and Prairie soils) and the brown soils
(Chestnut and Brown soils) in North America were occupied. The
frustrated, the persecuted, the seekers of new opportunity had a place
to go, and it was a good place with good soil. For over 200 years a
man and woman could carve themselves out a farm home on the colo-
nial frontier, in the Ohio Valley, on the great prairies of Illinois and
Iowa, and finally on the Great Plains to the west. Grassland needed
only cultivation, once transportation was established.
In addition to the fine soils, there were other free resources, the
forests and minerals. There was land in North America, Central
America, South America, New Zealand, and Australia. Following
the great discoveries Europeans found opportunities throughout the
240 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
world. This progress is credited by many people to western civiliza-
tion, but it would have been a poor civilization indeed that could not
have succeeded with these riches. With such abundant resources for
its citizens, it is hard to imagine how any American government could
possibly have failed.
At the present time, however, people must make a go of things where
they are, or else move into areas where a great deal of careful planning
is necessary for successful agriculture or industry. There is no more
Chemical Composition (%) Chemical Composition (%)
CaO MgO oO 50 100 100 CaO MgO
0 50
Ol 02 Rosie _| Depth RQIYGY By 04 2.0
O
te NW ae
RA XO QOS oo .0
OS 15.4
0.03 35.5
—AAACDSL
Xe oe
O25 0:
B. Conomingo Silt Loam
of Maryland
0.2 Tr 100
a ET AP TE I TF LAD LT LD A
7 Eee ==
FIO OKD KOO COCO O Oe,
RK KRG ROKER KK OL
i Nes SKS SERRE SPRERS
‘ “ Nas OPS
RRR SRKRKKS OKI K RK KR K
POS Nee
ROKK LKR KR SKLAR
eataces xO eatetstctes ate =
25 PARAHIPPUS
PARAHIPPUS .
MIOHIPPUS
30 t 5
wy MIOHIPPUS MESOHIPPUS
O
3544 O ‘
2 16
re) MESOHIPPUS
40 |
45 | 9
a aii OROHIPPUS
50
I
HYRACOTHERIUM
55 HYRACOTHERIUM
Ficure 1.—The evolution of the horse in North America correlated with the
absolute time scale. (Based on the work of E. D. Cope, W. D. Matthew, W. B.
Scott, G. G. Simpson, R. A. Stirton, and others.)
250 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
nor even in successive species of ages as much apart as 1 million years,
to believe that the minute differences present had any value as a
determining factor in survival.
Returning to the duration of a genus, it will be interesting to
consult other groups, to see whether the figure of 5% million years
found in the horses is more generally applicable. Similar values are
found in other groups of Mammalia, such as the land carnivores.
For these Simpson constructed a ‘‘survivorship curve’ showing that
the mean duration of a genus of carnivores is 6% million years. But
for the bivalve Mollusca, he obtained a mean as high as 78 million
years.
Just to add a few more figures: Swinnerton found 20 million years
as the average lifetime of a genus of Triassic ammonites. Lingula, on
the other hand, a brachiopod, has been known for its persistence since
the Cambrian. This genus has lasted over 400 million years. From
this evidence the terrestrial Mammalia appear to have genera of a
short duration, but even this is a matter of a few million years.
One might be inclined to interpret these different rates of generic
evolution in terms of species steps, assuming that each lineage would
pass through several evolutionary changes, each constituting a new
species, before the accumulated differences justify one in calling the
descendants a new genus. It is certain that this process underlies
the evolution of the horse, but it is conceivable that in other groups
genera may have originated without the interposition of a series of
species steps. The time rate of species formation, therefore, must
now be considered.
Selecting again terrestrial forms of life, mainly mammals, as our
first examples, because more detailed evidence is available, we find
that since the end of the last glaciation, some 10,000 to 20,000 years
ago, only minor subspecies have appeared. The differences are
confined to body size, color, slight differences in body proportions,
in the development of appendages, etc. The British race of the red
deer, for instance, has evolved since Britain became separated from
the Continent some 7,500 years ago. This is shown by fossil evidence,
since the early postglacial specimens found in the Thames belong to
the Continental race. Moreover the characters of Cervus elaphus
scoticus, as the British race is called, are probably not fixed genetically,
since, when the breed was transferred to a favorable environment,
in New Zealand, it reverted in many respects to the Continental type.
Other examples could be given of the insignificant character of
Postglacial differentiation in species. ;
Greater differences are observed in some, but by no means the
majority, of Upper Pleistocene species, of about 50,000 to 100,000
years ago. For the marmot of that time, for instance, Wehrli found
that the shape of the temporal ridges has since become stabilized.
TIME IN EVOLUTION—ZEUNER 251
These ridges run into the upper posterior edge of the processus
postorbitalis in a certain number of fossil specimens, while in recent
Marmota marmota they have moved to the upper side of the processus.
That this character has become practically fixed since the Upper
Pleistocene, is shown by the following figures:
TaBLeE I
Marmota Inter- Primitive
tyz.e mediate type
(Percent) (Percent) (Percent)
Wppersbleisvocenes 32. 4- =) = oe a IS 33 53 13
FES COT Gee ae a a res ee a ae Byer eye I a 98 LM, 2
Forms of this degree of variation would, in the recent fauna, be re-
garded as subspecies.
Going back to the Middle Pleistocene, about 250,000 years ago,
the differences become more conspicuous, but they are still treated
as subspecific by most taxonomists. It is only in the Lower Pleisto-
cene, about 500,000 years ago, that we encounter ancestral forms on
which the taxonomists agree that they must be classified as distinct
species. This applies, for instance, to the European elephants,
Elephas antiquus, and the mammoth, which Soergel has shown to
have evolved from an early Pleistocene common ancestor, Hlephas
meridionalis. If this view is correct, nearly half a million years
were required to produce a new species by way of a gradual change.
If it is not correct, the point of divergence lies farther back in the
past and the rate of species change is longer. In the lineages of
rhinoceroses, bears, and deer, similar evidence is available and there
are cases in the Recent fauna which have been explained as the results
of geographical isolation during the glaciations (the common crow
and the hooded crow, for instance). But again there are species
which have changed much less since the Lower Pleistocene, which
are regarded as no more than subspecifically distinct from their
Recent descendants and, therefore, must have a slower rate of species
formation. In the Cromer Forest Bed (about 500,000 years ago),
14 percent of the species are regarded as “Recent,” though it is quite
likely that subspecific differences will be detected by future revisers.
Five hundred thousand years, then, appears to be a fast rate of
species evolution in terrestrial mammals and no faster rate has, to
my knowledge, yet been found in other groups of the animal king-
dom. Let us now consider the question, how long have species re-
mained stable or unchanged? To answer it we have to resort to ma-
rine groups. Lower Miocene Mollusca of Java, for instance, are
regarded as conspecific with Recent forms. They have been stable
for 30 million years. Surveying fossils in general, this has to be
regarded as a high figure, but it need not be a maximum.
252 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
It is conceivable that plant species behave differently. Small
has analyzed several groups of plants from the chronological point of
view, notably the diatoms. He came to the conclusion that the mean
duration of species is measured in millions of years. Furthermore,
he distinguished in the diatoms long-lived species (up to 100 million
years) and short-lived species (one million years or less). In other
groups he encountered shorter rates than this. These time rates are
of much the same order as in the animal kingdom. But there is one
interesting difference: while in animals the evidence suggests gradual
change of specific characters, Small regards the changes in the diatoms
(and perhaps other plants) as sudden.
Summarizing, we may say that half a million years is a short life-
time for a species and, so far as our admittedly incomplete evidence
goes, something ike a minimum. The lifetime of genera appears to
be rarely less than 5 million years. Short rates are observed in ter-
restrial groups like most Mammalia, while long rates are frequent in
mainly marine groups like the bivalve shells and the diatoms. There
appears to be a certain amount of correlation between the rates of
species evolution and the degree of changeability of the environment,
slow rates being frequent in environments like the sea in which living
conditions are exceedingly stable.
Hitherto we have been discussing changes in single lines of descent.
Let us now consider the phenomenon of phyletic splitting. Branch-
ing is a frequent event in phylogenesis. An ancestral species may
evolve into two divergent descendant species by the disappearance of
intermediates, or the ancestral species may remain unaltered and a
new type emerge as aside branch. In fact, few groups would survive
without splitting, since owing to the action of internal (genetic) and
external (environmental) factors, many species become extinct.
If the rate of splitting equals the rate of extinction, the number of
species in a systematic category, like a genus or a family, remains
constant. But if the rate of splitting is greater than the rate of ex-
tinction, the number of species in the group under consideration will
rise increasingly steeply along an exponential curve.
There are many different ways in which the rate of splitting can be
plotted in relation to time. I have selected two simple methods of
plotting which can be applied readily to any group from which suffi-
cient fossil material has been adequately studied by a taxonomist.
One is to plot the number of species existing in any period or sub-
period, or smaller stratigraphical division, on the absolute time scale.
The other consists of doing the same for the newly appearing species
only. In the first, the surviving species modify the picture. The
second is a truer presentation of the rate of splitting, but the material
is often not complete enough for its application.
TIME IN EVOLUTION—ZEUNER 253
Now let us consider a few examples. Figure 2 shows the genus
Salenia. These sea urchins start in the Lower Cretaceous. The
number of species rises rapidly to a climax in the Upper Cretaceous
(“increase phase’”’). Cases like this one, of a rapid, almost sudden,
blossoming-out of a group were called ‘explosive evolution” by Schin-
dewolf. We may borrow the term and call the phase of rapid increase
the explosive phase. In Salenia it is followed by a catastrophic drop
followed by a slow ‘‘decline phase” leading to almost complete
extinction. Of the cause of this sudden drop we are completely igno-
rant; it is an unusual feature, as will be seen when other diagrams are
considered.
As regards the increase phase, we learn that in Salenia it lasted for
about 40 million years.
Figure 3 shows the Brachiopod genus Lingula. Again it reveals a
rapid rise in the number of species at the beginning, from the Cam-
brian to the Ordovician, of the order of 40-50 million years. But
between the increase and the decline phases, a more or less “‘station-
ary’’ phase is intercalated, during which the number of species did
not vary greatly. The decline phase of Lingula set in with the Car-
boniferous. It is a long-drawn-out phase and continues into the
present.
Another interesting case is that of the coelacanths, a group of the
fringe-finned fishes or Crossopterygii (fig. 4). This group never had
a truly explosive phase. Nevertheless, there is an increase phase
from the end of the Devonian to the Triassic, lasting about 100
million years.
One more example may be given, the molluscan genus Poiretia
(fig. 5). Here, the explosive phase in the evolution of the genus
is 80-40 million years. Other examples confirm that the length of
the explosive phase is of the order of a few 10-million years, the
extremes so far found being 30 and 100 million years and the mean
around 50 million years.
As an example from the plant kingdom, illustrating at the same
time a group with stable, geometrical characters living in an environ-
ment which changes but little, diatoms may be shown. Small in-
vestigated the chronology of their evolution, and the diagram of
Hemiaulus shown (fig. 6) is a translation into our method of plotting
of one of his diagrams. It shows a steepening rise to a climax in the
Cretaceous. The increase phase lasted about 80 million years, no
longer than in groups living in less stable environments, but there was
a pronounced initial lag phase.
Now it is interesting to consider some higher systematic categories.
One might expect that the explosive phase in the evolution of higher
systematic units, like families, orders, classes, is longer than in
genera. But this is not so.
254 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
J
eke)
See
He 50 fe)
million years
CRETACEOUS PALAEOGENE NEOGENE
FIGURE 2,—Diagram of the numbers of species in the sea-urchin genus Salenia
from the Cretaceous to the present day. (Surface area proportional to the
number of species.)
TATA 200 Tele) i
400
million years
U.CAMB.ORD. SILUR.DEV. CARB. PERM, TRIAS.JURAS.CRET. CAINOZOIC
Ficure 3.—Number of species in the brachipod genus Lingula, from the Upper
Cambrian to the present day. (Surface area proportional to the number of
species.)
TIME IN EVOLUTION—ZEUNER 255
COELACANTHIDAE
SRT RES TE
250 200 150 100 50 1@}
Diplocercididae
_ 300
million years
DEV, CARBON. PERM. TRIAS. JURASS.CRET. CAINOZOIC
Ficure 4.—Diagram showing frequency of species of coelacanth fishes in relation
to time. Inthe Upper Devonian the family Diplocercididae flourished. In the
Carboniferous it was replaced by the Coelacanthidae, which reached their climax
in the Triassic. The small number of coelacanths recorded from the Permian
may not represent their true frequency owing to the fact that relatively few
marine deposits are known from the Permian. Possibly, therefore, the rise to
the climax in the Triassic was not interrupted in the Permian, as is shown in
the diagram, but was continuous from the Lower Carboniferous; if so, the rise
would have taken about 100 million years and the rate of evolution would then
be one of the slowest known. The coelacanths are regarded as a very con-
servative group which changed but little in the course of time.
60 50 40 30 86.20 lO oO
million years
PALAEOGENE NEOGENE.
Figure 5.—Diagram showing the number of new species of the molluscan genus
Poiretia appearing in the different subdivisions of the Tertiary. The maximum
number of new species was produced in the late Palaegene, within 30 to 40
million years of the appearance of the genus.
256 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The first example of this kind is that of the rotaliid Foraminifera
(fig. 7), showing the number of existing genera in the superfamily.
Again there is a phase of steepening increase. It lasted from the
early Cretaceous into the Oligocene, for about 70 million years.
Another case is that of the genera in the brachipod family, Spiriferi-
dae, with an explosive phase of about 80 million years. A third
example shows the orders in the class of the true fishes. This diagram
0
HEMIAULUS
(DIATOMACEA)
TOTAL OF SPECIES LIVINGIN
ANY PERIOD
NO. OF NEW SPECIES
PRODYCEDIN ANY PERIOD
Triassic ———> |Jurassic—o|L.Cret. |UpCretac|Eocene OL |Miocene|Prioc. | R.
200 MILLION [100 MILLION YEARS
Figure 6.—The diatom genus Hemiaulus. The number of species living in any
period (blocks) and the number of new species appearing in any period (curve)
from the Permian to the present day. (Numerical material from Small, 1948.
Ordinate scale linear.)
150 100 © 50 fo)
million.years
TRIASSIC JURASSIC CRETACEOUS PALAEOGENE NEOGENE
Figure 7.—Diagram showing the number of genera in the superfamily Rotalioidea
(Foraminifera) present in various geological epochs. (Surface area of blocks
proportional to the number of species.)
(fig. 8), which is based on Romer’s work, is not strictly to scale, but
there is no doubt that orders like the sturgeons and gar pikes had
explosive phases of some 20-60 million years.
Finally, the number of orders within the class of the winged insects
(Pterygota) must be mentioned. Since about a hundred orders have
been distinguished, it is possible to plot them in much the same way
as species in a genus. Once more, an explosive phase emerges, an
exceptionally large number of orders appearing during the Upper
Carboniferous and the Permian, over a period of about 60 million
years.
TIME IN EVOLUTION—ZEUNER 257
TABLE 2.—Insect orders
Existing New
1 DYERICO LOVE OLE es as Toes Aes popes SEG we gs yee Adis Mallee erent yen aes 0 0
ower! Carboniferous ee sac te ete ae en eit a eee Pe 2 2
Wpper Carboniferous! =. a2. es 2S 25s oe nee cee aes ee 18 16
Perm is nes is eee eae eee Pe ne ees Bg 37 30
IMlESOZO1C =e Sree 2 ee Mee at gs tei iE E Ey Ti ib oem pe Ros Ye pe eS eee ep 31 22
CAS oy RN Mey een al RE te ape ree OnE ee et eet oe ey ge 38 13
BB eY 1s} 0) ee ey Se As ISR eS Vege ae aR eee EN Pore Se A PE aero SE CI Sea ee eC
million
ears
| |
aoe > : CAINOZOIC
i i
sul CRETACE-
i y OUS
$ 2
= eo
7 NG JURASSIC
3 of
°.
8 :
on | jee
4 PERMIAN
B,
: 5 CARBON
: : ‘ -IFEROUS
DEVONIAN
SILURIAN
FiacurE 8.—The evolution of the true fishes correlated with the geological time
scale. Based on a diagram by A. S. Romer.
To summarize, increase phases (often of the ‘‘explosive” type) in
the evolution of genera, families, and orders lasted, so far as evidence
goes, something like 15-100 million years, with a mean of about 50 or
60 million years.
In other words, the increase phase of evolution is of the same length
[of the order of 50-60 million years (+30)], irrespective of the sys-
tematic unit investigated. It is difficult to resist the temptation to
speculate on the significance of this result. What it appears to mean
is that the number of species steps involved in the evolution of new
systematic categories is the same irrespective of the rank of the cate-
gory. It takes no longer for a new order to evolve than for a new
genus to appear.
258 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
If this is accepted as a valid inference from the material studied,
then the difference between genera, families, orders, and classes must
lie in the quality of the unit steps involved in their evolution. This
is not a new idea,? though we have arrived at it from a new and un-
expected direction.
What is meant by the statement that the quality of the steps de-
cides the systematic category which is evolving, is easy to illustrate
but difficult to formulate. A good example is the evolution of the
jaws of the fishes from gill arches. This is an extraordinarily interest-
ing case of change of function of an organ. Its real significance,
however, is that this change provided the fishes with a decided ad-
vantage over their environment. It opened up new and better food
supplies and, as Sewertzoff expresses it, “increased the general life-
intensity of these animals.”’ A biting-mouth skeleton is important
also as an offensive and defensive weapon. It is, I think, sufficiently
evident that this particular change of function was full of evolutionary
potentialities. Applying Sewertzoff’s terminology, such changes may
be called aromorphs.
On the other hand, if one considers the extreme adaptation of a leaf
insect (Phyllium), one finds that it has—to use Sewertzoff’s phrase-
ology—led to a decrease in the life-intensity of the animal. No new
food supplies have been made available by the process of adaptation,
and its evolutionary potentialities are nil. This type of adaptation is
typical of the lower systematic categories.
There is another interesting feature that. emerges from the chrono-
logical treatment of evolution. It is that there appears to be no
correlation between fast rates of species evolution with groups having
a rapid succession of generations. Elephants are among the most
slowly breeding animals known and yet their rate of evolution was
the fastest so far found. On the other hand, Drosophila, which,
because of its rapid succession of generations, is so much used in
experimental biology, is a genus 50 million years old. It appears
therefore that a rapid succession of generations must not be taken as
a, substitute for long periods of time. It is indeed surprising to find
that the number of generations is not the only factor ruling the rate
of change in evolution and that this change is in a vague way correlated
with absolute time. Does this perhaps mean that external, environ-
mental factors are influencing evolution over very long periods of
time? I am unable to answer this question, or to offer any other
explanation.
I hope I have been able to show that a study of the chronology of
evolution is well worth the effort. The suggestions made here must
?This has been put forward in different forms, for instance, by G. G. Simpson and R. Goldschmidt.
TIME IN EVOLUTION—ZEUNER 259
not be taken as final results. For the time being the material is
still too scanty to make generalizations sufficiently safe. Some of
the time rates, however, have been found by several workers inde-
pendently, and some of the rules—if one be allowed to use this term—
have been deduced by more than one worker.
BIBLIOGRAPHY
GoLpscHMIpT, R.
1940. The material basis of evolution. 436 pp. New Haven.
ScHINDEWOLF, O. H.
1947. Fragen der Abstammungslehre. Aufs. Reden Senckenb. Nat. Ges.,
Frankfurt a M., vol. 1. 23 pp.
Simpson, G. G.
1944. Tempo and mode in evolution. 237 pp. New York.
SMALL, J.
1946. Quantitative evolution—VIII. Proc. Roy. Irish Acad., ser. B, vol.
51, No. 4, pp. 53-80.
1948a. Quantitative evolution—IX—XIII. Proc. Roy. Irish Acad., ser. B,
vol. 51, Nos. 17-21, pp. 261-346.
1948b. Some laws of organic evolution. 15 pp. Privately printed.
ZHUNER, F. E.
1946a. Biologicai evolution and time. Ch. XII in Dating the Past. London.
1946b. Time and the biologist. Discovery, vol. 7, No. 8, pp. 242-249, 256.
(Some diagrams shown in the discourse are figured in this article,
and acknowledgments are due to the editor of Discovery for per-
mitting their reproduction.)
he sabi Bry)
“SS
ay -_ iL ° fo
— :
a han Fhe ae BS Aydihiet pond Win! : idee organ
4 anibe tne ENE “spe pie rie
rh eta)
iD ;
pone i 9n0 rn anon Noe cgens vd 4 wl
# Th ’ ; ; s i
ee : ie oe a
+3 “=F crits Myf 17 f
i Vee
\ ¥ _ ‘ ii srry ‘9 ve a
‘ 3 mr - = _ ,
pormliwat wih) .cubiwloys Wn siead Wsquieit alt O45
Z re : : , 5 Z| :
‘ ‘ : +t us s Kf% era a
eat) pan deine ce ack ailotegati iets Sal eg TE
: A . ty On eh HO? 995 a mbt —— hs _
S t a ; B
7 i ee “= Pada) CRA a”
; ‘ | ¥ yas any BY wks btee ds Bh. oheadiz ii er "i — fs
| | ; . ppt ee:
sy A A600 Dah botal. tL, anier 1 e74 f—aohuior ave ii avid ae a :
eae he " e-88 AS * an 0 —_— L
are ie el fa A heel’ ; (Or er TELS A edt if "5 by aaalstta nity? dope
eh ibe 5: iar’ OG ae vega TS Df dial Soe <
; Sue. « Shade 3 op iat 1) f f fio i Add CASAS . rel adres oe be ir. ine
: ' - il ni : ‘. : ; : ca A tons : ,
EAS. hee a ps hs - r tk , oe
rive: Menten TLE Le Lee ir } with) hia neil toys uotyclotl. evel
| a : ag a Sf « ip a a o,f \ Par) he i Ni
sta, At : Ls rte 4 | : one al 7 ‘a hl Riri | bad bia at Us
ie ania ‘edity pill te Sig a beats rw riyais Sine
ne ‘ets sai ae ih TOR see sada Cr Ye ow Vw ii wel ; writs! tte i gaks a)
= r j i» (ape ¥ ee 43) bia i sada! 12 399%
MORE ABOUT ANIMAL BEHAVIOR !
By Ernest P. WALKER
Assistant Director, National Zoological Park
[With 16 plates]
INTRODUCTION
The scope of the subject ‘‘animal behavior’ is almost unlimited, for
every species has its own particular pattern of behavior and the indi-
viduals of the species show many variations even from that pattern, so
that the subject could really be made an exhaustive study for each
and every species. Although psychologists have made extensive
studies of human behavior, actually they have merely made a good
beginning. Glimpses of animal behavior and some conception of how
behavior patterns may have developed frequently help in explaining
human behavior and human reactions, for our reactions are the result
of a long series of experiences of trial and error and elimination by
natural causes just as has been the case with those of other animals.
Similar causes have brought about similar reactions in many
instances, and different causes and environment have developed
different reactions; there is, therefore, a remarkable diversity of
behavior pattern among the different kinds of animals that live under
many different conditions. Those that have not been able to adjust
themselves to their environment have become extinct and are usually
known only by their fossil remains; those that could adapt themselves
and multiply are the forms that have survived to the present day.
The activities in an animal’s life are essentially the same as the basic
activities of humans. Individuals must survive, and to do so they
must be able to avoid enemies, to obtain food and shelter; and for the
species to survive, the individuals must reproduce and the young must
be given an opportunity to start their own struggles for survival.
No doubt everyone who has observed animals closely has seen them
do many things that could not be explained in the light of our ex-
periences and practices. However, we must judge and interpret the
1 An earlier article by the same author, under the title ‘‘Animal] Behavior,’ appeared in the Annual
Report of the Smithsonian Institution for 1940, pp. 271-312, 18 pls. As the first paper has been out of print
for several years, some of the same ideas and examples contained in it are incorporated in the present paper,
although not as exact quotations.
261
262 § ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
actions of animals in the light of the lives they normally lead in the
wild. If we are well enough informed as to the conditions under
which they live and their mode of life, we can often understand actions
that would otherwise be incomprehensible or aimless to us. I do not
believe that animals do any more aimless things than humans. On
the contrary, I am often impressed by the direct and efficient
manner in which they carry on the work of their lives.
The habits of animals are so intimately associated with and gov-
erned by their structure that one who is somewhat familiar with either
the habits or the structure can often make rather close deductions
from one regarding the other.
Every habit or type of behavior that has been developed because it
helps the animal in its life is, of course, a special trait. Some are so
little known that attention might be focused upon them and con-
sideration given to the manner in which they benefit the animal.
To understand the animals best, we must realize that merely because
we cannot perform a given act does not mean that an animal cannot
do it. For example, the sense of smell in most humans is so unde-
veloped that we cannot trail another animal by scent, as we know many
animals do. Also, our hearing is so dull we cannot detect sounds that
we know many other animals hear. Likewise, other senses of animals
are probably so much more keenly developed than ours that they re-
celve impressions or information of which we know little or nothing.
For example, some animals give off vibrations that other animals are
able to detect; the presence of a warm live body can be detected by
certain snakes at distances of several feet even though their eyes and
nostrils are not functioning; and apparently impressions are received
by insects through their antennae and by fish through their lateral
lines.
How individual animals acquire their behavior patterns is a fas-
cinating field for study. Some actions are apparently taught to the
young by the parents; some are learned by the young by observing
others of their kind; some are learned by the trial and error system, or
as we know it, “by bitter experience”’; and some come to the animal
by instinct; that is, the animal reacts in a certain way (usually the
right way) under certain circumstances without previously having
had an opportunity to learn consciously to act in that manner under
those circumstances. How such reactions develop is explained by
various theories and is a separate study.
ECOLOGY, STRUCTURE, AND HABITS
Ecologists refer to an animal as occupying an ecological niche, that
is, the main activities of the animal take place within a certain type
of habitat. It may live entirely within the water, on or under the
ground, or almost entirely on trees, or combinations of any of these,
ANIMAL BEHAVIOR—ERNEST P. WALKER ”263
and its food is definitely limited by the products that it can obtain in
the habitat which it occupies. Animals that do not use the same food
or are not antagonistic, because one does not prey upon the other,
can occupy the same areas or overlapping areas, so that ecological
niches do not have definite, clearly defined boundaries, but are rather
areas or space spheres of activity having also a time component. The
structure of the animal to a large extent governs the type of ecological
niche that it may occupy, since through evolution the animal’s form
has been modified to adapt it to the type of ecological niche that has
been available to its long line of ancestors down to itself. Thus we
can to some extent explain and understand the great variety of forms
of animal life.
We might consider two little creatures of approximately the same
size which occupy different ecological niches and are very different
in structure, and see how they fit into their respective spheres. These
are golden hamsters (Mesocricetus auratus) and “‘flying’’ squirrels
(Glaucomys volans).
The golden hamster of Syria averages about 4 ounces in weight,
with a maximum of about 8 ounces. It is a stout-bodied, short-necked,
short-armed, and short-legged little mammal with short fingers and
toes and a tail only about half an inch in length. It has very large
cheek pouches that open inside the lips and extend far back of the
shoulders, in which it can carry surprising loads of food or nest mate-
rial. It is rich golden-reddish brown above and white beneath, with
white hands and feet. The little creature is an inhabitant of an arid,
rocky region where it lives mainly among rocks or in burrows in the
ground around the rocks. Syria’s climatic conditions result in a
scarcity of food material for such little creatures for long periods of
time; therefore, the storage of food is necessary and the cheek pouches
are of great value in carrying food to its burrow or den. It works
industriously at this, apparently the only limit to the amount of food
it carries being determined by the amount that is available.
They are relatively slow-moving, clumsy little creatures in com-
parison with many other animals that we know, but are obviously
well adapted to leading their lives in an efficient manner in their
habitat. Their movements remind one of military tanks in that they
are slow, ponderous, and persistent. There is nothing sprightly or
agile about them, but if they want to go in a certain direction or up a
certain surface they keep at it so persistently that frequently they
succeed, and they can wedge themselves through surprisingly small
crevices. They display remarkable persistence and determination to
get into crevices that would seem to be so small and uncomfortable as
not to be attractive to them.
The mouths of rodents are definitely on the underpart of the head
and back of the nose, making it difficult for them to cut upward when
866591—50-—18
264 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
in their normal position. The animals solve this problem very
readily, however, by throwing themselves on the back or side and
cutting upward with their teeth. A picture of such an effort is shown
in which my “Shehammy” has thrown herself on her back and is
trying to cut upward on the lower edge of a door so that she can
squeeze through the crack between it and the floor (pl. 1, fig. 1).
This aperture was only seven-eighths of an inch high and, when she
tried to force her way through it standing on her feet, she was much
too thick and her body was much too tense, but in the inverted posi-
tion she extended her arms forward so that she was able to get a grasp
on the lower edge of the door and pull herself through the crevice,
which she did repeatedly. This ‘““chammy,”’ one of my pets, did it
regularly, and I have seen other rodents do it occasionally.
Frequently one of my golden hamsters, when permitted to run
about my study in the evening, would clamber up in the crack between
a bookcase and the wall a foot or more above the floor and stay there
for a considerable time. She usually rested by bracing her feet
against the wall and her back against the bookcase and apparently
was content to stay in that position. This suggests to me that these
animals adopt such a position as a means of being comfortable in
fissures between the rocks among which they live.
When hamsters want to get down from an elevation that they have
reached, they apparently do not think of jumping down even a few
inches, but lower themselves as far as possible with the hind feet,
sustaining themselves by the top of the hind feet, which are bent at the
ankle. Next they release one foot and then the other and fall head
downward, usually landing on their noses, but sometimes on their
backs (pl. 2). They then sit up, sneeze, rub their noses, and go about
their business. It will be noted that they do not utilize their hind
claws or toes which are too small to be effective. This method,
while probably a relatively inefficient action, has persisted because
it is not seriously harmful in the circumstances under which they
live, and is the best they can do with their short fingers and toes
which are not of much use to them for such descents.
At night, when foraging for food, hamsters apparently spend very
little time eating, preferring to fill their pouches, take the load of food
home, and return for more (pl. 1, fig. 2). In this way they gather
food when it is available and eat at leisure in their nests without
being exposed to danger. During the daytime, their sleeping period,
they frequently wake up and eat, probably consuming much more
food at this time than during the night when they are actively foraging,
grooming, exploring, or visiting their neighbors.
When “‘hammies” are away from their homes or other shelter and
there is a sudden alarm, they dash for shelter with surprising speed for
little folk with short legs but if they are touched or attacked before
ANIMAL BEHAVIOR—ERNEST P. WALKER 265
they can get away, they throw themselves on their backs incredibly
quickly and are prepared to fight savagely with their sharp, fairly
strong teeth and tiny claws. If they are in their nests, they pay little
attention to outside disturbances, at most sniffing from just inside the
doorway.
Baby ‘‘hammies” arrive in litters of as many as 17, tiny, blind,
naked, pink, helpless little fellows that lie on their backs and nurse
while mother “‘hammy” hovers over them to keep them warm in the
snug cup-shaped nest of fibers she has constructed in a remote portion
of the burrow or den. The little ones grow very rapidly, and by the
thirteenth day they have a good coat of fur, are eating solid food, and
begin wandering about in the burrow, den, or cage. Even though
their eyes are not yet open, they find their way back to the nest if
mother has not detected their absence and carried them home. By
the fifteenth day their eyes are open, and by the seventy-third day they
produce babies of their own. ‘They bear large families in rapid suc-
cession but cease to breed regularly when about 1 year old, and the
life span is short (probably a maximum of about 3 years). There is
a high mortality rate, probably throughout life.
The structure and lives of the flying squirrels (Glaucomys volans) of
North America are in marked contrast to those of the hamster.
Flying squirrels weigh about 3 ounces, have slender bodies, long,
slender, strong arms, fingers, legs, and toes with fairly long, curved
claws, and a rather long tail. Theskin of the body is much larger than
is necessary to enclose the delicate little form and is extended outward
along the side as hair-covered membrane indistinguishable from the
skin of the body. This skin reaches down to the wrists and ankles,
so their outline is almost square when the little creatures put their
arms forward and out and their legs back and out as they do while
gliding through the air. This adaptation more than doubles the area
of the upper and lower surfaces as it appears when the animals are at
rest. In keeping with this form, the hairs of the sides of the tail are
long, and closely set, and project at almost right angles, whereas those
of the upper and lower surfaces are very short and lie close to the bone
of the tail, so that the tail is, in cross section, almost like a feather.
All these are modifications for life in an entirely different habitat from
that of the hamster.
Flying squirrels live among the trees, generally making their nests
in hollow limbs, old woodpecker holes, or other shelters high above
the ground. They are strictly nocturnal, having very large eyes
adapted to admitting the maximum light available. They are not
limited in their movements to climbing up and down the trees or to
mere leaping between objects, for their remarkable form gives them
power to glide long distances. ‘To do this they seek an elevated point,
leap out into space and spread their arms and legs so that they take
266 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
on the form of miniature gliders (pl. 4, fig. 1). When approaching a
landing, which they generally try to make on a tree or other object
not on the ground, they swing slightly upward to check the glide and
bring their long arms and legs forward as shock absorbers (pl. 4,
fig. 2). Instantly upon alighting they dart around to the other side
of the tree or limb, which is probably a provision to escape any
enemy that might be following them.
Before flying squirrels take off on a glide they almost invariably
sway the head and body as far as possible first to one side then the
other, repeated several times, and often raise as high as they can
stand and crouch as low as possible. 'Thisis probably a sort of range
finding by triangulation.
They feed very largely on nuts and acorns which they gather and
store in their nests and probably in almost any location in the forest
in which they can find a cavity that is large enough to hold a nut
securely. I assume this from the fact that my pets regularly place
nuts at many places about the house other than in their nest box.
They particularly selected the top of the window, the medicine cabinet
in the bathroom, my hand, my pockets, inside my collar, and even
the depression between my arm and body when I have my arm close
to my body. When placing nuts, they have an interesting habit of
trying to make them secure wherever they are leaving them. They
force them into the crevice or onto the surface where they are leaving
them, then tap them three or four times with their front teeth. This
suggests that they probably similarly try to wedge the nuts in cracks
in the bark or slight depressions on the tops of tree limbs in the wild.
In addition to nuts and seeds, they eat some fruits, berries, and
insects. My pets eat mealworms, grasshoppers, and small bits of
meat in moderation.
“Glauckies” (short for their scientific name) are very hesitant
about going down to the ground, feeling much more at home leaping
about in the trees, gliding from place to place; however, they will go
down on occasion. When they do so, they are plainly not at home
and run with the arms and legs extended so as to hold the body as
high above the ground as possible, the gliding membrane being
pulled upward close to the body so that a very peculiar effect is
produced.
Instead of sleeping soundly and ignoring disturbances in their
neighborhood, as do the hammies and many other burrowing creatures
who know they are safest in their dens, the flying squirrels sleep
lightly and at the least disturbance in their neighborhood they peer
out or dash away.
“Glauckies’”’ are born in families of two to six pink, helpless little
ones, with the gliding membranes plainly evident. They develop
ANIMAL BEHAVIOR—ERNEST P. WALKER 267
slowly, are not well clothed in fur until about the twenty-eighth day,
and continue nursing until about the fifty-sixth day. In recognition
of the dangers of an active life in trees, they, like young tree squirrels
(Sciurus), start out exploring near the nest with great caution, and
venture farther and take more chances only as they gain strength,
experience, and confidence. Their gestation period is about 60 days
and young are born when the parents are about 1 year old.
These brief descriptions of the widely different hamsters and flying
squirrels, which inhabit very different ecological niches, give a glimpse
of the specialization among animals which enables them to survive
and to lead their lives under diverse conditions.
Giraffes (Giraffa) have developed such a high degree of specializa-
tion that they are able to fill an ecological niche without competition
with other mammals except in those portions of their range which
overlap that of the elephants. Much of the food of the giraffe is
taken from the upper portions of shrubs or from the lower branches
of trees far above the reach of most mammals. In addition to its long
legs and very long neck, the giraffe has elongate, mobile lips and can
extend its tongue several inches to help it gather in leaves and twigs.
The moles (Scalopus and other genera), which burrow in the earth,
are blind, have exceedingly short forelimbs, with nose and tail very
sensitive to touch, large, powerful hands armed with long, strong
claws, and very strong muscles in the fore part of the body, particularly
those associated with the arms. The fur is very soft and short and
will lie in any direction; thus it does not impede the mole when it
desires to run backward in its burrow, which it often does. Moles
feed on earthworms, grubs, and other small animal life they catch in
the earth. (See pl. 5, fig. 2.)
In marked contrast are the gibbons (Hylobates and Symphalangus),
highly specialized members of the primate or monkey group. Their
arms are exceedingly long and powerful, with elongate hands and four
strong fingers, the thumb being small and situated far back on the
hand. The legs are rather long, and the feet are better fitted for
grasping than are the hands. The body is small. These adaptations
enable gibbons to be the most expert of all mammals in swinging by
their arms through the forest from branch to branch and even from
tree to tree. In this process the hands are used as hooks, the thumb
taking no part in grasping. Since the hands are usually occupied
when traveling through the trees, gibbons carry objects such as food
or branches by grasping them with the feet. The same method of
carrying objects is used by chimpanzees (Pan) and orangutangs
(Pongo).
Whereas the gibbons, chimpanzees, gorillas (Gorilla), and orang-
utangs lack tails, the spider monkeys (Ateles) of Central and South
268 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
America possess long, prehensile tails which they use in many
ways, especially to hang by or to use as a supplementary hand
in steadying themselves. In the picture (pl. 11) it will be seen that
one of the animals is using its tail to hold itself while it leans far
forward. On one occasion I saw seven spider monkeys in this group
leaning forward, held by their tails, watching for the door to open so
they could go into the house for their afternoon meal.
Some rodents have large, powerful incisor teeth that protrude
much more than those of the majority of rodents, and one naturally
surmises that this trait is associated with some habit peculiar to them,
which indeed it is. The animals possessing such teeth use them in
burrowing; that is, they cut away the earth, remove small stones,
and cut roots, thus performing most of the functions that the powerful
hands of the mole perform and, in addition, do the cutting work
which the mole’s hands cannot do. Some other rodents, and many
of the insectivores also, have projecting incisor teeth, but these are
relatively small and slender and are obiously not suitable for such
heavy work as digging, moving small stones, or cutting. These are
used as forceps for the picking up of food, which consists mainly of
small fish, insects, or worms.
Such animals as the kangaroos (Macropus), the kangaroo rats
(Dipodomys), the African spring hare or spring hass (Pedetes), the
jerboas (Alactaga, Scarturus, Salpingotus) and others have large,
long, powerful hind legs, relatively small, weak, short forelegs, and
long tails which are either tufted with long hairs near the tip or are
rather heavy, often very thick near the base (pl. 5, fig. 1). These
animals are leapers and use the hind legs almost exclusively in propul-
sion. The tail is a balancing member; it also acts as a tripod leg
in some forms.
The types of leaps of different animals vary greatly. One interest-
ing difference is the contrast between the leap of the flying squirrel,
an animal that leaps directly toward its objective with a very flat
trajectory, often planning to land below its target and then climb
up to it, and the leap of the tarsier (Tarsisus) which has a very high
trajectory and lands the leaper at its objective. After becoming
accustomed to photographing the leaps of flying squirrels, I tried
photographing tarsiers leaping and aimed my camera as I would
for the squirrels. With the pronounced leap upward of the tarsier,
I at first caught only the tail and hind portion of the animal in the
upper portion of the picture. After a few such experiences I learned
the difference in their leaps and corrected my aim accordingly.
The list of such highly specialized forms could be expanded indefi-
nitely to include representatives of all groups of animal life, for
every animal is specialized to lead a certain type of life.
ANIMAL BEHAVIOR—ERNEST P. WALKER 269
CLEANLINESS
Almost all animals are fastidiously cleanly about their persons and
their homes. Because of the fact that many animals live on or in
the ground or on trees or other surfaces that frequently are dirty
according to our standards, we often erroneously consider the animals
to be uncleanly. As a matter of fact, practically every animal takes
great pains to avoid soiling its coat, and when it does unavoidably
become soiled, the animal at the first opportunity painstakingly cleans
itself by shaking, rolling in the sand, washing in water, or licking itself.
Even those pests, the common Norway rat, roof rat, and house mouse,
which have found that they can obtain food and shelter in man’s
filthy surroundings, endeavor to keep themselves clean. If they are
observed when not alarmed, it will be seen that they carefully pick
their way through their surroundings in an effort to avoid soiling
themselves, and after they have finished their exploration they will
invariably be seen to groom carefully. If one will part the fur of
small mammals it will be seen that the skin is generally immaculately
clean.
A star-nosed mole (Condylura cristata) that I am keeping as a pet
comes out of the ground, goes into a dish of water and at once begins
grooming. In a few minutes it has washed thoroughly.
Animals have developed many ways for keeping themselves and
their nests clean. Apparently flying squirrels rarely void excreta
during their entire sleeping period, which may be as much as 16 hours.
Obviously they wish to keep the inside of the tree or other nest loca-
tion clean, so wait until they go out at night when they can see
clearly and there is a minimum of danger from outside enemies.
Many of the burrowing mammals have little toilet rooms that open
off of the main burrow where they deposit their excrement, thus keep-
ing the main burrow clean. Young North American opossums
(Didelphis virginiana) of an age to be in the pouch or still be clinging
to their mother apparently do not release their excretion unless they
are stroked on the lower abdominal region. This is probably a pro-
vision for keeping the pouch clean until such time as the mother is
ready to clean the little ones.
POSTURES
The new-born babies of many of the small mammals naturally lie on
their backs. This leaves them in the proper position for nursing
when the mother hovers over them, protecting them, hiding them,
and keeping them warm.
Most hoofed animals do not voluntarily sit on their haunches. A
horse assumes this position briefly when getting up, but this does not
constitute true sitting. The exceptions to the rule are the tapirs,
270 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
which frequently sit on their haunches with the forefeet supporting
the front part of the body (pl. 14, fig. 2).
A position that would appear to us to be uncomfortable is regularly
adopted by flying squirrels, which normally live in hollow trees. The
body hangs in a vertical position, either head up or head down, sus-
pended by the fingers or the toes, most of the load actually being on
the fingernails or the toenails (pl. 3, fig. 1). They sleep in such poses
at least a portion of the time. The hanging position is probably
assumed as the result of having accommodated themselves to situations
where there is no satisfactory ledge or bottom in the den, so that a
tree cavity that would otherwise be unsuitable can be utilized by
clinging to its rough surfaces. However, they now take this pose
freely even when comfortable nests or rests are available. Of course,
they also take other positions such as curled up in a ball or lying on
the side or the back.
The inverted position so often used by sloths (Choloepus, Bradypus,
and Scaeophus), wherein they hang beneath a limb, is well known and
is entirely suitable for their purposes. Bats generally hang head
downward.
HOMING INSTINCT
When an animal has become established at a given location, it
almost invariably develops a strong attachment for its home even
though such home does not provide entirely satisfactory quarters.
Almost every animal has a rather definite area in which it lives its
entire life, and it rarely leaves this range except when forced to do so
by very unusual circumstances. This is commonly shown when
captive animals escape, for they usually remain nearby for some time
and frequently go back into the cages if they are able todo so. People
who are familiar with animals regularly take advantage of this trait
by leaving the cage door open, refraining from frightening the animal,
and placing food in the cage or nearby until the animal is recaptured.
On one occasion I was keeping a specimen of the least short-tailed
shrew (Cryptotis parva) in a jardiniere. I had observed it trying to
leap out, but it is not adapted to leaping and was apparently unable
to reach the top. After a few days I discovered that it had escaped
during the night probably by superior jumping gained by persistent
exercise. It was so tiny that it was useless to search for it in the
house, so I waited until evening and then placed the jardiniere on the
floor almost directly below the place at which it had been standing on
my desk. JI crumpled around it an old blanket so that it formed a
ramp or runway from the floor up to the top of the jardiniere. I then
mashed a mealworm and dragged particles of it in a trail on the
blanket from where it touched the floor to the top of the jardiniere.
Within a couple of hours the little fellow was back in his home.
ANIMAL BEHAVIOR—ERNEST P. WALKER 201
Plainly he had come to recognize the jardiniere as home and was glad
to get back to it when it was made possible for him to do so.
Remarkable developments of the trait of returning to the home
range are shown by carrier pigeons (Columba), and birds, mammals,
and fish that migrate.
SELF-PRESERVATION
Humans ordinarily give little thought to the matter of self-preserva-
tion, although it becomes a vital subject during wartime. Among
other animals it is ever foremost. In its broadest aspects self-preserva-
tion involves not only active and passive resistance to enemies, but
also the ability to obtain food and shelter. Humans have recently
had brought to their attention the fact that they, like other animals,
must either be prepared to defend themselves or run away when an
ageressor attacks or threatens. If there is no place to which they can
escape, or if they cannot successfully fight off their enemy, they have
only the alternatives of being subjugated and made slaves, as some-
times happens among humans, or of being destroyed, which in the
case of animals may mean being devoured.
There are very few animals that are aggressive to the extent of
trying to drive away or subjugate their neighbors merely for the
pleasure of the victory. Ordinarily, they will start a fight only when
it is necessary to obtain food, shelter, or a mate.
The methods used by animals to avoid, escape, or defeat an enemy
are almost as numerous as the species of animals, for the individuals
of every species employ one or more methods. All are interesting
and some are very remarkable.
The seeking of shelter is the most common of passive defensive
measures and is practiced to a greater or less degree by all animals.
Some go into burrows in the ground, crevices in rocks, or holes in
trees, while some of the larger ones retreat into dense jungles or
forests.
“Freezing,” the act of remaining motionless, is extensively employed,
as at the first intimation of danger almost every animal ceases prac-
tically all movement. Since an object which is very conspicuous
when moving is readily overlooked when stationary, “‘freezing’’ is a
highly successful means of avoiding detection by enemies. Even
those that depend mainly on their ability to scent their prey may
have difficulty locating it if they cannot see it.
Rabbits and hares (Leporidae), quail (Colinus), and grouse (Tetra-
onidae), when perfectly still in their native haunts, are very difficult
to see, and often will not move unless one almost steps on them.
This method of escape is applicable in almost any type of surrounding,
even on seemingly barren plains or expanses of snow. It is most
effective, of course, if the color pattern of the animal is such that it
272 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
harmonizes and blends with the surroundings or if the outline of the
animal is obliterated by some type of camouflage. Most animals are
so colored that they blend perfectly with their surroundings, some,
such as the ptarmigan (Zagopus), certain hares (Lepus), and weasels
or ermine (Mustela), even changing to white in winter to blend with
the snow.
Among insects, camouflage and mimicry have been developed to a
high degree of perfection in many species. Some resemble their
natural surroundings so closely that they are difficult to distingusih
even though one is looking directly at them. Resemblance to dead
or green leaves or stems is the most common camouflage, the pattern
and form of the insect being modified to a strikingly perfect imitation
of the color and form of those parts of the plant on which the insect
usually lives. In the type of specialization known as mimicry, one
species of insect very closely resembles and usually behaves like
another which is distasteful or dangerous to animals. By this
resemblance the imitator is often let alone by an anima) that mistakes
it for the undesirable insect which it resembles.
When it is not possible to avoid an enemy, some animals use @ more
or less passive method which is well illustrated by the armadillos
(Dasypus, Tolypeutes, and related genera) which roll themselves into
a ball, completely protecting the feet, tail, and head within the armor
plate of the body (pl. 6, figs. 1 and 2). This method effectively
baffles many animals that would ordinarily devour them, and when
the would-be attacker becomes discouraged and leaves, the armadillo
unrolls itself and goes about its business. Another method is the
feigning of death, which is well illustrated by the American opossum
which, when alarmed, falls on its side with the mouth partially opened
and appears so limp and inert that it is often left for dead by animals
that would vigorously attack if they surmised the animal to be alive.
Apparently opossums are unpalatable to many animals, so that this
means of protection is very effective. When danger ceases to threaten,
the opossum gradually resumes activity, but if the attacker is merely
waiting nearby and makes a movement, the opossum will usually
again go into its death-feigning act. It is supposed that this behavior
is a type of fainting induced by fright and is perhaps not actually
under the control of the animal. When the hog-nosed snake of the
eastern United States, frequently called spreading adder (Heterodon),
feels itself in danger, it feigns death, throws itself on its back, and
assumes various grotesque poses if it is seriously aggravated. It does
not, however, do this unless the annoyance is great and continued.
Many insects feign death when a disturbance occurs in their vicinity.
This is illustrated by the many beetles that inhabit low shrubbery,
which drop from their elevated locations to the ground and remain
quiet or quickly take refuge in a more protected location.
ANIMAL BEHAVIOR—ERNEST P. WALKER 273
One of the most remarkable modifications of feigning death of
which I have heard was witnessed and described to me as follows by
John N. Hamlet, of the United States Fish and Wildlife Service:
Three of us recently saw a Cooper’s hawk (Accipiter cooperi) chase a spotted sand-
piper (Actitis macularia). The piper dropped into the water and stayed under for
several seconds. The Cooper’s lit on the stream edge a few yards down the
stream. The sandpiper came to the surface and floated down the stream with
its wings open flat on the water and its neck stretched out. It passed within 3
feet of the hawk who gave it no more than a casual glance. The piper floated
down stream about 20 yards and took off and disappeared.
The live sandpiper drifted close by the hawk but was not recognized
because it was not moving in its usual pose. This is a choice example
of the effectiveness of ‘freezing’? and assuming an unusual pose.
Mr. Hamlet has witnessed similar action by sandpipers on two other
occasions.
Bluff is another effective defense that is employed by many animals.
Its best form is for the animal to face its enemy in a pose not usually
assumed by it and that makes it appear as large as possible, ferocious,
and threatening. Many, if not all, of the owls (Strigiformes) bluff
by crouching low, spreading their wings at almost right angles to the
body and ruffling the body feathers until the bird appears several
times larger than it really is. The bittern does likewise, and most
mammals bluff to some degree. A good example is the dwarf weasel
(Mustela rixosa campestris), only about the size of a cigar, who stands
his ground, opens his mouth wide, barks, and even attacks if need be,
although its teeth and jaws are so small it can scarcely break the
skin of one’s hand. The domestic cat’s (Felis cattus) high-arched
back, bushed-out tail, and wide-open, snarling mouth present a good
example, and many of us have witnessed the hestitation of a dog
suddenly confronted by such an attitude. Often this hesitation
gives the cat an opportunity to escape without having to fight.
Fighting for mates is definitely beneficial to the species, for by
nature’s law of the survival of the fittest, which must prevail through
all species, only those survive that are best able to take care of them-
selves. This, taken in its broadest sense, includes not only physical
strength but mental alertness and adaptability to varying and new
conditions. Males that are physical weaklings and would not father
vigorous offspring are ordinarily vanquished in encounters for mates
and therefore leave no progeny. By this process nature has consist-
ently eliminated the unfit and has improved each of the species.
The occasional maiming of individuals in their conflicts is not in
itself injurious to the species, as such mutilations are not inherited
and, if the parent was vigorous, the progeny stand excellent chances
of being vigorous even though the parent may have been injured in
some of its conflicts.
274 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Among animals that live in colonies or in herds, conflicts are gen-
erally rare except among males that are fighting to determine who
shall be master of the herd, as in the case of bison. However, on one
occasion I witnessed a definite fight in a prairie dog (Cynomys) colony.
In this case the aggressor was outside of the burrow and trying to
keep another one in the burrow and, at the same time, to fill the
burrow with dirt to bury him. Several others were within a few
feet watching the proceedings but taking no part in it although it
was obvious that they sympathized with the “‘underdog.’”’ Whenever
the one that was in the burrow would attempt to come out, the one
that was outside would try to bite and scratch him, and, when he had
him forced back into the burrow, would scratch and push earth into
the entrance. Finally, after perhaps 20 minutes, he lost interest and
left. The bystanders almost immediately went to the burrow, greeted
and apparently sympathized with the one that had been attacked,
and in a few minutes the normal life of the colony had been resumed.
This method of fighting by trying to imprison the victim is also used
by prairie dogs in closing the entrance to the burrow which a snake
has entered.
ADAPTABILITY
The readiness with which animals accept approaches or friendliness
of man varies greatly. Some seem to be so thoroughly imbued with
caution and suspicion of man’s intentions that they can be tamed
only with the greatest of effort. Others respond almost immediately
to handling and friendly treatment. Examples of the latter are the
gray foxes of the United States (Urocyon) which tame very readily
and in marked contrast to the general wariness and slow taming of
the red foxes (Vulpes). The beaver (Castor) is another animal that
tames easily, sometimes merely a few hours of kind treatment being
sufficient to win its confidence. Young hair seals (Phoca) also tame
almost immediately when captured. Usually they seem to have no
fear when picked up, looking to their captors as friends and becoming
affectionate pets, sometimes swimming after a boat that is leaving
them behind after its occupants have picked them up and petted
them briefly.
Penguins are fearless and very curious as to visitors on land or on
the ice or snow where they normally have few or no enemies, but in
the water they are cautious as there they are accustomed to watching
for enemies such as killer whales (Orca), sea leopards (Hydrurga), and
large fish.
Another phase of the adaptability of animals is the degree to which
wild animals can survive where man has established himself. Most
of them appear to have no fear of man’s presence and his activities
so long as they are not actually molested and their haunts and food
ANIMAL BEHAVIOR—ERNEST P. WALKER 275
supply are not destroyed. We can see numerous examples of animals
that have gone ahead fairly successfully with their ives when man
has not interfered too much with conditions essential for them to live.
Another group consists of animals that thrive in close proximity to
man and either become a part of man’s household and receive his
direct aid in their existence, or adapt themselves to the conditions
that man provides and obtain food and shelter in spite of his utmost
efforts to control them. In this group are the Old World rats (Rattus)
and mice (Mus) that have become established almost throughout the
world wherever man has made settlements. Other examples are the
European house sparrow (Passer domesticus), commonly called English
sparrow in the United States, and the European starling (Sturnus
vulgaris) both of which have become firmly and widely established in
North America. These two birds, like the rats and mice, have be-
come pests in some regions where they have adapted themselves to a
remarkable degree and have become very plentiful. They have found
sufficient food and shelter around man’s activities so that they have
thrived where other less adaptable forms have not been able to sur-
vive. Indeed, in many cases they fight the native birds and take
over the ecological niches normally occupied by the local birds that
are less adaptable and aggressive and have not been successful in
defending their territory.
It is sometimes said that there is a third group of animals comprising
those that cannot tolerate proximity to man but, if we study the
problem, we usually find that man’s activities so changed conditions
essential to them for food or shelter that they could not survive, or
else that man intentionally killed them off.
Most animals, if given to understand that man will not harm them,
will become tame, and if man will feed them it is surprising how
friendly even very wild kinds will become. I have a pet big brown
bat (Eptesicus fuscus) that in a few days became so tame it would
return to me after each of its flights about the room (pl. 13, fig. 2).
Even when I awaken her she makes some effort to overcome her
stupor and come into my hand to be warmed preliminary to taking
her evening flights and receiving mealworms, of which she is very
fond?
Chipmunks (Zamias) that have been tame during the summer or
fall, then go into partial or complete hibernation and are inactive
for a period of from 5 to 6 months, come out in the springtime and
resume their friendships and friendliness practically as though they
have not been interrupted. One individual that came in the window
onto my desk and was so tame that I took a picture of it while it was
on my hand obtaining food, returned the following spring and re-
7 fa the Saturday Evening Post for February 4, 1950, is an account of further results of my studies of
ats.
276 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
sumed our relationship where it had broken off with the beginning
of bad weather the previous fall.
Another example of the confidence of wild creatures is the case of
two flying squirrels that are greatly loved guests in our home. They
have such fondness for both my wife and myself that they want to
be with us at every opportunity, even to sleeping in our pockets.
They leap to us as a means of quick locomotion and for the pleasure
of the leap and glide (pl. 4, fig. 2). Their behaviour has been treated
at some length in the National Geographic Magazine for May 1947,
“Flying Squirrels, Nature’s Gliders.”
OBTAINING FOOD
The obtaining of food is an ever-present problem for all animals.
With some it is apparently a very simple process; with others it brings
into play the use of the special structures with which nature has pro-
vided them. The food-getting habits of practically every animal
would make an interesting study, but only a few of the widely differ-
ent methods can be mentioned here.
The peculiarly shaped bill of the flamingo (Phoenicopterus), which
looks as though it had been broken in the middle and the tip bent
downward, is used in an interesting manner. As will be seen from
the picture (pl. 7, fig. 1), the bird extends its neck almost straight
downward, which brings the upper portion of the bill closest to the
ground. The edge of the bill then scrapes along the ground in very
shallow water, and the lower mandible opens and closes rapidly.
The water thus scooped into the mandible is strained between the fine
laminations of the upper and lower mandibles and then discharged
after the minute crustaceans on which the flamingo feeds have been
strained out.
Relatives of the flamingo are the swans and other water fowl
(Anatidae), many of which obtain their food in fairly shallow water.
They eat a wide variety of plant and animal material such as seeds,
green leaves, tubers, fleshy roots, mollusks, and crustaceans, and a few
of them eat some fish. When the food is not too far below the surface
of the water, they “‘tip up’ so that they can extend the neck and head
to the maximum distance below the surface (pl. 7, fig. 2). Others dive
for their food.
Herons (Ardeidae) feed mainly on small fish, frogs, and other
animal life, which they obtain by standing practically motionless in
the water and waiting for the unwary victim to come within reach.
They suddenly extend their long necks to a surprising distance and
catch the prey before it has a chance to dart away. They may stand
in such a position for hours at a time waiting for their victims to
approach.
ANIMAL BEHAVIOR—ERNEST P. WALKER 207
A very different method of obtaining food is employed by the birds
of prey (Falconiformes), which catch their victims by a quick swoop.
An outstanding example of a bird that employs this method is the
falcon (Falco), which while soaring or flying almost out of sight will
detect a bird in flight, or a bird or small mammal on the ground, dive
from thousands of feet at speeds of as much as 280 miles an hour with
the wings almost folded but extended just enough to enable it to steer
itself, and strike and carry off its victim in its talons. In these dives
the speed of the bird is so great that a whistling sound is produced.
On one occasion I heard the whistling and an instant later a half-grown
chicken was struck within about 8 feet of where I was standing, the
falcon having come down at about a 45° angle over my head. It
scarcely seems possible that a bird can fly with such speed and accuracy
as to overtake and capture a startled duck in the air, but the duck
hawk (Falco peregrinus anatum) does this regularly, providing a
thrilling demonstration of skill and dexterity. Hunting with falcons
was a royal sport in the Old World for many centuries and then was
almost discontinued in Europe. It continued in Asia and is now
popular again in Europe and the United States, where many different
kinds of birds of prey are being trained to capture game or lures and
bring them to the trainer. Among the birds that have been so trained
are the beautiful, dainty little sparrow hawk (Falco sparverious),
Cooper’s hawk (Accipiter cooperi), duck hawk, the prairie falcon
(Falco mexicanus), golden eagle (Aquila chrysaetos), and even the
barred owl (Strix varia).
Another and little-known method of obtaining food.is practiced by
the skimmers (Rynchops nigra), in which the lower mandible is con-
siderably longer than the upper. Their regular practice in feeding is
to fly so close over the water that the tip of the lower mandible is in
the water, and when they come upon fish they merely lower them-
selves enough so that they can scoop up the fish.
Practically all woodpeckers (Picidae) feed on insects, most of
which they capture in their burrows in wood. ‘These birds may often
be seen going up or down trees, keeping the body often in a vertical
position by grasping the trunk with the feet and leaning back on
the stiff tail. Through their keen hearing they detect an insect under
the bark or in the wood, and then proceed to cut away the bark
or wood with their beaks until they approach close enough to the in-
sect or worm so that they can draw it out of its burrow with their
long-barbed tongues. The work of the woodpeckers in removing
insects from the wood is far more beneficial than harmful, for if the
insect is allowed to remain there it may do considerable damage,
whereas the woodpecker ordinarily works only to excavate wood that
has already been damaged by the insect, and by removing it prevents
278 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
further damage. The flickers (Colaptes), members of the woodpecker
family, have found a less laborious method of obtaining a considerable
portion of their food—they have learned to eat ants on the ground
and have taken to feeding extensively on these and other insects in
addition to what they get from the trees. The California wood-
pecker (Balanosphyra formicivora baird2) and its relatives feed to some
extent on acorns, which are very plentiful in many portions of their
range, and they have adopted the practice of making holes in the
bark of trees or in posts in which they place acorns for future use.
Sometimes a trunk or limb of a tree or post will be studded ‘with
hundreds of acorns, each in its own separate setting which holds it
securely with the end of the nut projecting slightly from the hole.
The night hawks (Chordeiles), whippoorwills (Antrostomus), swifts
(Chaetura), swallows (Hirundo) and their relatives capture practically
all their food in their very wide capacious mouths while flying. Some
birds such as the tyrant flycatchers (Tyrannidae) and a few others
have the habit of sitting on a perch that gives a good view of sur-
rounding territory and watching for insects. As these come within
range, they are captured in the air in a short flight by the bird, which
then returns to its perch to watch for more. The hummingbirds
(Trochilidae), as is well known, have a very long beak and tongue
which they extend into flowers to feed on the nectar and insects that
they find there. Other small birds known as honey creepers (Co-
erebidae and Melithreptidae) have a long, sickle-shaped beak which
they use in obtaining insects, nectar, and fruit pulp and juices.
Feeding habits among the mammals also vary widely. The langur
and colobus monkeys (Presbytis and Colobus) feed mainly on the
leaves of certain trees. Many of the bats, particularly the small ones
(Microchiroptera), feed almost exclusively on insects that they cap-
ture in flight. Some forms are carnivorous, killing other bats and
small birds and perhaps small mammals. Vampire and false vampire
bats (Desmodus and Diaemus) lap blood that they obtain from warm-
blooded animals by cutting off a very thin layer of the skin with their
razorlike incisor teeth. Other very large bats with a wingspread of
as much as 4 feet, which inhabit the Tropics, are known as fruit bats
(Pteropus and related genera) because they feed largely on fruit.
These bats frequently travel in considerable flocks and move to
various regions from time to time to find food.
Most rodents eat mainly plant food such as seeds or nuts, leaves,
stems, or roots of plants; many, however, also eat some insects and
small amounts of meat. A few have developed highly specialized
feeding habits together with specialized teeth and shape of jaws.
These include the fish-eating rat of South America (Icthyomys) and
an insectivorous type (Rhynchomys) from Luzon Island in the Philip-
pines. Nothing has been recorded regarding the habits of this animal,
Smithsonian Report, 1949.—Ernest P. Walker PLATE 1
1. GOLDEN HAMSTER (MESOCRICETUS AURATUS)
Lying on her back, she is trying to cut upward on the lower edge of a door. In
this position she pulled herself through the %-inch crack under the door.
2. GOLDEN HAMSTER (MESOCRICETUS AURATUS)
The cheek pouches are filled with seeds. The pouch is outlined by a white dotted
line.
‘doip puv 00} 1940 oy} 9Ssvolor 07 ApBaI
puv posBofar JOO} auO “VY SsII :19Y SUIP[OY o1¥ Jooj Jo soovjans szoddn ‘sso] Jo YYSuI| [[NJ 0} posaMo] ‘199 U90 ‘aspa ey} IOAO J[asioYy BursaMo] “WJay
YOOTH SHL SACS SSAHON] OF ATSHS V WOY4 NMO”d ONILLAD (SNLVYENVY SNLADIMDOSAW) YSLSWVH N3G105
c 3a1lvid JI¥Je A “d IS2UIA—"Gp6| ‘Oday uetuosyztuISg
Smithsonian Report, 1949.—Ernest P. Walker PLATE 3
1. EASTERN FLYING SQUIRREL (GLAUCOMYS VOLANS)
Hanging head downward is one of its favorite positions while eating.
2. EASTERN FLYING SQUIRREL (GLAUCOMYS VOLANS)
The flattened tail, long, strong fingers and toes, large eyes, and edge of the gliding
membrane are clearly shown.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 4
1. EASTERN FLYING SQUIRREL (GLAUCOMYS VOLANS)
A glide as seen from beneath and to one side.
.
2. EASTERN FLYING SQUIRREL (GLAUCOMYS VOLANS)
Preparing to alight from a glide.
The two pictures on this plate were taken with the electronic flash which operates at about 1/5000 of a second.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 5
1. EGYPTIAN JERBOA (JACULUS JACULUS)
A characteristic pose, with the very small arms and hands held close to the throat.
The hairy pads beneath the feet are protection against hot sand and the abrasive
action of rough surfaces.
2. TOWNSEND'S MOLE (SCAPANUS TOWNSENDII) (MUSEUM SKIN)
Showing the broad, powerful forefeet with large, straight claws for digging; the
lene, sensitive, mobile snout; short, sensitive tail; and dense, soft, short, plush-
ike fur.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 6
1. THREE-BANDED ARMADILLO (TOLYPEUTES TRICINCTUS)
Rolled up as a protection against enemies. The top of the head is the large inset
portion in the front of the shell; the tail is lying along side of the head. (Photo-
graph by Joao Moojen.)
2. THE BACK OF THE SAME ANIMAL
The hinged arrangement of the plates of the back permit it to roll up completely.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 7
1. FLAMINGOES (PHOENICOPTERUS)
The birds are shown feeding in shallow water, grooming, and resting.
2. WHISTLING SWAN (CYGNUS COLUMBIANUS)
“Tipping up” to obtain food from the bottom. In this position these birds can
reach to a depth of about 45 inches.
(uslooyy ovor
kq ydeisojoyg) ‘“xuey Joy uo sey puly yuo
S}l puv ‘lapynoys Joy uo We 4YS syr ‘uIyO Joy
Jopun pay SI YIIM JoyJOUL vy} OF Sursuryo st Aqeq W
(SOLVNOYMOL
SNHdOaVOS) HLOIS GANVYW 3uvuy SHL
é
‘att
8 ALV1d
£
a
é
‘1oYYBJ 9yQ JO Yowuq oy} UO a1B SoIqed OMT,
(SNHOOVF
XIMHLIITNIVD) LESOWNYVYW GSELANL-ALIHM °1
oF
JOHeA\ “q 1S2UIY—"GpG| ‘Oday ueruosy wg
Smithsonian Report, 1949.—Ernest P. Walker PLATE 9
1. FEMALE SOUTH AMERICAN OPOSSUM (METACHIRUS NUDICAUDATUS)
This animal lacks the marsupial pouch. Some of her babies clinging to her
nipples are to be seen between her hind foot and her tail. (Photograph by
Joao Moojen.)
2. FEMALE SOUTH AMERICAN OPOSSUM (METACHIRUS NUDICAUDATUS)
Hight baby opossumsare clinging to the nipples between the hind legs of their mother.
(Photograph by Joao Moojen.)
"YOIS 94} JO pus oY} UO 4ooSUTI
uv YO}Bd 0} YSnOUs Ivy pusejxo p[_nod snsuo0y ST
{le} o[IsSuesyoid oy} puv qooy Suidsvis oy} 910N *ainyord SI Ul UMOYS [JOM ST JOO}
“BLOUIBD OY} SuIyoyVM st ado YJo_ SYP =“ APQUepued oy} jo Aqrypiqe Suidseis o[qevyIvulel oy, “qySusy ut
492} 8 INOGB QUIT] 991} B SUTAIIVO ST 41 JOOF YU SII SAT UT
-apul y1OM soo VSOYM piBzZIT poziperoeds ATYSIy VW
(WIIWNd VHYNVSONDSIAN) NOAISWVHD “Z (iqa8v OSONOd) NVLNONVYO NVYLYWNS “1
Ol ALVId JAVA “d S2UIF—'Gp6] ‘qaoday ueiuosy qu
‘pavoq B 0JUO SUIPOY SI [rey S,4eyyou oy} Jo dry oy,
‘]I@] Joy JO oseq oy} punowe [1e} sit poddeam sey YOIM ‘Yyouq Joy UO SuIplA Aqvq B sBYy JouI0D jo] Joddn ayy Ul o[RUIEJ OY YT,
(SNSOYSWNAA SAIALY) SABMNOW YaCIdS
PM SBE Ace! JOWEA “d I89UIY— G6] ‘WOdoy ueruosyzrus
Smithsonian Report, 1949.—Ernest P. Walker PLATE 12
aad te eae shoo Tere gree man
tiara et
PLAINS LEAST WEASEL (MUSTELA RIXOSA CAMPESTRIS)
Upper and middle, in summer coat; lower, in winter coat (this form apparently
does not become entirely white). This is a very rare form of which practically
nothing has been recorded, and these may be the only pictures ever taken of
it. About three-quarters natural size.
“qSLIM
8 1OYING oy} UO SUTPFIS SI YOIYM (SUDI0a ShwwoznD)y))
Jolainbs sutAy udtojsvo uv IOAO SuIAy st yeq oy, re} opIsuoyoud A]5u0I4s St YIM JloszT SurApvoys st 4]
(SnOsna SNOISA1ldy) LVG NMOYG Olg “Z (SNAW14 SOLOd) NOFYYNIM *1
€l 3LV1d JIA[EA\ ‘qq 1S9U14—"6} 6] ‘J40dey ueruosyAI US
Smithsonian Report, 1949.—Ernest P. Walker PLATE 14
1. TWO-TOED SLOTH (CHLOEPUS DIDACSYLUS)
Hanging beneath a limb is its usual pose. The forefeet have two toes, the hind
feet, three toes.
2. YOUNG MALAYAN TAPIR (ACROCODIA INDICA)
The striped baby coat is being replaced by the solid gray and black of the adult
coat. Age 3 months.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 15
1. MALE BLACK AUSTRALIAN SWAN (CHENOPIS ATRATA)
The bird is attacking a man in defense of the half-grown young in the foreground.
The female was just to the left outside the picture.
A AT UI A aii
Lh PAAR RRO SIE iirc roe, ment
es : «ee ae 1 wer ee : 7 Fe eo ;
BEN he \ gn Js ol eee ion Sena F soe ease ee Be ae |
2. MALAYAN PORCUPINES (ACANTHION BRACHYURUM)
The two animals alternate head to tail, in which position they regularly sleep.
Smithsonian Report, 1949.—Ernest P. Walker PLATE 16
1. SIX-BANDED ARMADILLO (DASYPUS SEXCINCTUS)
The animal is on its side asleep. They tremble almost constantly when sleeping.
2. THIRTEEN-LINED GROUND SQUIRRELS (CITELLUS TRIDECEMLINEATUS) ASLEEP
Three rest on the top of their heads, one is lying on its back, and one in the back-
ground is mainly right side up.
ANIMAL BEHAVIOR—ERNEST P. WALKER 279
but we can judge from its structure that it probably eats insects or
other soft animal food. The food of carnivores, the flesh eaters, is
rather uniformly flesh or fish, but carnivorous animals frequently pass
over the principal meaty portions of a carcass to drink the blood
and eat parts of the viscera, thereby obtaining valuable vitamins.
Almost invariably the glands or other parts which they eat are known
to scientific workers as being excellent sources of vitamins. In one
experiment that was conducted under my direction, we separated
chicken viscera into its various component parts and then offered
these to small carnivores. Almost all immediately chose the pancreas,
which suggests that it is probably a valuable food. Most people
have witnessed a dog burying a bone and later digging it up and chew-
ing on it. It is possible that the bacteria of the soil act on the bone
to make it more palatable and more digestible and possibly to elimin-
ate any danger of ptomaine poisoning. A few carnivores, such as the
binturong (Arctictis) and the palm civets (Paradozxurus and related
genera), have taken to a diet consisting largely of fruit.
An ingenious method was once demonstrated to me by the West
African marsh civet or marsh mongoose (Atiliz paludinosus) in the
National Zoological Park. This animal, about the size of a large cat,
had a remarkable method of breaking bones. He was commonly
given two pieces of horse ribs 4 inches in length and, when he had
eaten most of the meat off of them, he would take a piece of bone
between his forepaws, raise himself up on his hind feet with the hind
legs well extended and with his forepaws well above the level of his
head, and then quickly throw the bone down on the cement floor of the
cage from a height of 2% to 3 feet. If he was not satisfied with the
results of the first throw, he would repeat the process. The pro-
cedure described suggests that these animals probably use the same
methods in breaking the shells of mollusks and land crabs on which
they feed in their native haunts of West Africa.
The shortest mammal in the Americas and almost the smallest in
the world is the lesser short-tailed shrew (Cryptotis parva) which
inhabits the eastern United States. It weighs as much as two dimes,
less than one-fourth of an ounce, and naturally such a tiny creature
cannot cope with large antagonists in the usual manner. It normally
lives in loose soil and leafmold where it feeds on earthworms, insects,
and a wide variety of small animal life including frogs. Even a
vigorous earthworm is a difficult creature for the tiny shrew to subdue
in the usual manner, but earthworms lose their activity within a few
seconds after the shrew gives them a few light bites; sometimes a
single nip will suffice. Apparently the saliva carried into the very
minor injury made by the shrew’s teeth is poisonous to the earthworms
and takes effect very quickly. When the earthworm has become
quiet the shrew proceeds to devour it, and it may be that this special-
866591—50-—_19
280 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
ized saliva accelerates digestion. Little is known regarding the
poisonous bite of these little creatures, but it is apparently similar
to that of a slightly larger close relative, the short-tailed shrew
(Blarina) whose saliva is definitely known to be poisonous to many
small creatures and whose bite on a man’s finger is sufficiently
poisonous to produce pain extending halfway up the arm with resultant
irritation lasting as much as a week. Such bites are somewhat com-
parable to the bites of snakes in which a secretion from specialized
glands is introduced under the skin of the victim, although these
shrews do not have specialized teeth or glands that are primarily
poison producers.
Among the reptiles, the chamaeleon (Chamaeleonidae) has the
interesting habit of obtaining the insects on which it feeds almost
exclusively by suddenly extending its tongue to a distance almost as
great as the length of its head and body and catching the unwary
insect with it. The tongue is attached near the front of the mouth
and folds back into the mouth like that of the frogs and toads, which
also capture their prey in a similar manner (pl. 10, fig. 2).
Emil Liers’ study of otters (Zutra) in North America have shown
that these remarkably intelligent and playful animals feed largely on
crayfish and numerous small invertebrates instead of fish, which they
have generally been supposed to eat. It is strange that this was not
discovered long ago, emphasizing the fact that there is still a fertile
field for research on animal life.
Through the long period of development of each species, animals
have learned that in general each individual or pair requires a cer-
tain amount of territory in which to sustain itself. Other species
that do not conflict with it will be tolerated in this territory, while
those that would be competitors may be driven out. Forms of life
that constitute the food supply are rarely devoured to the point of
extermination, and animals can therefore ordinarily obtain sufficient
food within a rather definite territory. Some require but a small
range for this purpose; others must have an extensive territory and
move to various parts of it at frequent intervals to obtain the necessary
food. For example, the wolf (Canis nubilus and related forms), an
animal that preys on other animal life, usually has a range about 20
miles in diameter and, except when the female is living at the den
caring for the young, it traverses this range, generally in large circuits,
returning to any given portion of the range about once every 2 weeks.
Seasonal fluctuation in the food supply, such as the migration of
fishes or the dying off of insects, results in the migration of forms that
prey upon them to locations in which food can be found. For example,
insect-eating birds have an abundant supply of food in the Northern
Hemisphere during the summer but, with the approach of winter,
ANIMAL BEHAVIOR—ERNEST P. WALKER 281
insects and fruits become scarce and many of the birds cannot survive.
By migrating to regions farther south, however, they are able to
obtain an adequate supply of such food. Some forms that do not
migrate change their feeding habits during the year. During the
summer when fruit and insects and other animal life are abundant and
easily obtained, they feed extensively on these elements, and in the
winter they eat seeds and plant life that has matured during the
summer. Good examples of this are the birds of the sparrow group
(Fringillidae), in which the young are fed almost exclusively on insects
and fruit and the parents eat such food extensively when it is available,
but during the winter when few insects or fruits are to be had, they
feed mainly on seeds. Others feed only on certain types of food and
apparently cannot survive unless they can obtain these particular
foods. Notable among these are the cross-bills (Loxia), birds of the
Northern Hemisphere about the size of sparrows with usually some
bronze, red, or purple markings. They feed on seeds of the spruce,
hemlock, or pine which they obtain directly from the cones by perch-
ing on them and reaching under the scales of the cones with their
peculiar crossed mandibles and extracting the seeds. Such beaks
are probably well adapted for this purpose but are a serious handicap
to eating the type of food that most birds consume; therefore these
birds are erratic in their occurrence in a given range. If there is a
failure of the seed crop of the spruce, hemlock,-or pine, they must
move to regions in which there is a good crop of seeds on these trees.
Such sturdy, hardy animals as the American bison (Bison bison)
and caribou (Rangifer) have extensive ranges that embrace both
summer and winter forage grounds to which the animals migrate
seasonally, some making a round trip of several hundred miles each
year, measured in straight lines.
We are prone to think of our food selection and handling as being
superior to that of animals, but after observing the care with which
animals select the choice morsels and pass over food that they detect
as being contaminated or not palatable to them, I believe that the
wild ones probably are far better fed and are better judges of food
than we are. Studies of the feeding habits of antelopes on the native
range disclose that in a 14-day period they eat 24 to 30 different kinds
of vegetation. Possibly some of the material was eaten as many of
us have taken food during war time, merely for the purpose of survival,
but obviously they picked and selected foods that they preferred and
thought were best for them. If one will watch a wild rabbit or almost
any wild animal, he will see that it constantly and carefully selects
or rejects food. On many occasions I have endeavored to induce
animals, either wild or in captivity, to eat food that I had seen them
pass over. Rarely could I get them to do so, and on one occasion,
282 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
the third time I offered a certain nut to a very friendly wild squirrel,
it took the nut, opened it, and dropped it, perhaps to show me that
the nut was not fit to eat.
TRANSPORTATION OF OBJECTS
The need of transporting food, nest materials, or young, and occa-
sionally objects that apparently are taken for ornamental purposes
or merely because the animal likes them, have been solved in various
ways. Most animals carry objects in their mouths to some extent,
just as a dog carries a stick or ball, but there are a number of ways
in addition to this: for example, some of the Old World monkeys
(Macaca and others) have cheek pouches that open inside the lips,
but outside the jaws, in which they are able to store food, which must
be a great convenience to them when they are trying to obtain their
share in the presence of other greedy monkeys. They then grab what-
ever food they can, put it in their cheek pouches, and eat it at their
leisure. Monkeys with food in their pouches sometimes look as though
they had mumps on one or both sides. Internal cheek pouches are
found in a number of other animals such as the chipmunks (Tamas
and Hutamias) and hamsters, which use them as shopping bags or
baskets into which to place food to carry it home to the den where it
is stored for future use (pl. 1, fig. 2). Another type of cheek pouch is
the external one which is fur-lined. This is present only in a few
North American forms such as the pocket gophers (Geomys, Thomomys,
and related genera), the pocket mice (Perognathus) and kangaroo rats
(Dipodomys and Microdipodops). Their pockets open outside the
mouth, the skin of the face being folded inward to form the pockets.
The pockets can be turned inside out to clean them, then pulled back
into place by a special muscle. The owners of these pockets carry
food, nest material, and even earth in them.
We are so accustomed to seeing mother cats carry their kittens
by grasping the skin at the back of the neck that we commonly think
of this as being the principal way of carrying young; however, there
are many other methods. The mother squirrel grasps the skin of
the baby’s abdomen in her lips or teeth so that the little one hangs
in an inverted position beneath her head, grasping her neck with its
hands and feet and curling its tail around her neck, thus aiding in
the carrying and leaving no dangling appendages to interfere with
mother’s hands and feet in her travels through the trees. Baby
monkeys cling tightly to the mother in most cases, but the white-
tufted marmoset (Callithrix jacchus) mothers generally carry the
babies only when nursing them; the rest of the time the fathers carry
them (pl. 8, fig. 1). They cling to his long fur and ride on his back or
under surface and hold on so securely that he can make leaps through
the trees without dislodging them. Baby gibbons cling to the mother
ANIMAL BEHAVIOR—ERNEST P. WALKER 283
on almost any part of her body and frequently take a position around
her body almost like a belt.
The Brazilian mammalogist, Joio Moojen, of the Museu Nacional,
Brazil, has informed me that the baby of the rare sloth (Scaeophus
torquatus) clings tightly to the mother and is almost completely hidden
in her long loose hair. If no danger threatens, it may cling to her
back, which is frequently the underside; however, if danger threatens,
it is brought onto her chest or abdomen which is usually her upper
side or is between her and a tree trunk. In this position it is well pro-
tected between the mother and the trunk or branch to which she is
clinging and is so well hidden in the fur that it can scarcely be seen
(pl. 8, fig. 2).
We commonly think of the marsupials as always carrying the
young in the mother’s pouch. This is true for many of the forms,
including the common opossums (Didelphis) of North and South
America, but there are some marsupials that lack the pouch on the
abdomen of the mother. In some of these species the young are
carried or dragged about by the mother as they cling to her nipples.
Good examples of this type are the small South American mouse
opossums (Marmosa and allied forms) which range in size from
about that of a house mouse to that of a common rat. The young
hang suspended from the mother’s nipples which are in a cluster
between her hind legs, and in this location they are well protected.
When the young become larger, they are dragged on their backs,
as the mother walks along the ground. If one becomes detached from
the nipple, it is lost, for apparently they cannot again attach them-
selves to the nipples, and the mother appears to make no effort to
rescue them. The Metachirus pictured with the little ones hanging
from her nipples had eight babies (pl. 9, fig. 2).
Rodent babies of a number of different forms in various parts of the
world also cling to their mother’s nipples until they are of good size,
and when she travels about on the ground or through the trees, they
dangle or are dragged about on their backs. The North American
wood rat or pack rat (Neotoma) has this habit, and as many as four
young, which individually may weigh almost a fourth as much as
their mother, will be dragged about by her.
In the Tasmanian “‘wolf”’ (Thylacinus), a marsupial, the pouch is
a fold or shelf of skin on the abdomen between the hind legs which
opens backward—a sort of rumble seat—and the young are carried
in this. The best pouch of all is possessed by the kangaroos; they
carry the babies in a large, deep, baglike fold of the skin of the mother’s
abdomen until they may weigh almost one-fourth as much as the
mother. When there are no young in this pocket, or when they are
very little, it is shrunken and drawn up until it is rather small, but
as the young grows the pouch stretches and can accommodate one
284 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
young until it is well able to take care of itself. There is a muscle
around the entrance of this pouch which acts like a draw string and
constricts the opening until it is very small, preventing foreign objects
from getting in and the young from tumbling out.
The very beautiful, graceful kangaroo rat mothers of North America
lift their babies by grasping them by the back of the neck with their
lips or teeth. They steady the little ones by holding them with the
forearms, then hop along on their hind legs. This method of carrying
the young was observed and photographed by that great naturalist,
the late Vernon Bailey.
The North American opossum is said to carry leaves and straw for
its nest by wrapping the tip of its tail around the material. Some
doubt exists, however, that this is an established habit, as only a few
instances have been reported.
As previously mentioned, gibbons, orangutangs and chimpanzees
frequently carry objects grasped in their hind feet as well as in their
hands. On one occasion I saw a chimpanzee pick up a piece of banana
in one hand, a piece of bread in the other hand, and a head of lettuce
with one foot. The ability to carry objects in the foot is particularly
useful to animals that regularly traverse the forest by swinging by
their arms from limb to limb. An orangutang carried a limb about
2% inches in diameter and 8 feet in length from an outside cage to an
inside cage in the National Zoological Park, handling it much of the
time by grasping it with the hind foot (pl. 10, fig. 1).
A number of the animals that burrow, particularly those that make
extensive tunnels such as the pocket gophers of North America, the
mole-rats (Cryptomys.and Bathyergus) of Africa, and the bamboo-rats
(Rhizomys and related genera) of Asia, push the earth before them,
placing the hands close’to the sides of the head and against the earth
and supplying the motive power for moving the body forward by the
hind legs. Others scrape earth rearward with the forefeet and then
send it farther rearward by strokes of the hind feet. This is the most
common method of those that do not make continuous tunnels. The
prairie dogs (Cynomys) of North America move a great deal of the
earth that they use in building mounds by this method, and they also
push earth before them.
Beavers carry at least part of the earth or mud they use by holding
it against the breast with the hands.
Elephants (Elephas and Loxodontia) pick up objects by encircling
them with the trunk.
Apparently both whales (Cetacea) and seals (Pinnipedia) some-
times hold their young to their sides by means of the flipper. This
procedure has been observed so rarely and for such brief glimpses that
little is known of it. Whalers say that a mother whale sometimes
ANIMAL BEHAVIOR—ERNEST P. WALKER 285
uses this method to take her baby under water with her out of danger.
Seals may do it for the same reason.
Recently my wife and I witnessed a female black individual of the
gray squirrel of the eastern United States collecting bark fiber for her
nest. She went onto a dead limb of a tulip tree about 8 feet from our
window and began loosening a strip of bark about a foot in length.
Some was torn loose entirely but left hanging by a small strand, other
parts were not hanging down but were well loosened from the branch.
I thought at first she was not satisfied with the quality of the bark,
and I was about ready to offer her advice on how to collect bark
without wasting so much of it. Finally she began picking up the
hanging fibers in her mouth, placing them crosswise and tucking the
ends in so they would not drag and be in her way. When she came
to the strands that still kept the pieces of bark attached to the limb
she cut them off. In a few instances she backed up and pulled them
loose by tugging. I then realized that she did not need any advice
on how to gather bark with her equipment. She had loosened all the
bark she needed but left it attached so it would not fall to the ground,
then when she had enough loosened she began gathering it up. Of
course she could not have had much loose bark in her mouth and still
cut more bark loose from the limb with her teeth. As usual she had
done her work in the most efficient way.
Many times I have offered my pet “flying” squirrels a tidbit that
they preferred to the one they were eating. ‘This confronted them
with a real problem. Their experience through the ages has been not
to drop food, for it falls to the ground and is ordinarily lost. They
usually solve the problem by looking for a place in which to cache the
food they are eating, then come back to me to take what I am offering.
Likewise I have many times put the same problem or a similar one up
to the gray squirrels (Sciwrus carolinensis) of the eastern United
States. Most of them try to solve it by attempting to take the second
morsel without dropping the first. Usually they cannot do this but
make several attempts and finally finish by sitting nearby to eat the
first piece and then come for the second, or sometimes they give up
the attempt and take the first piece home or at some distance to eat,
then return for the second. When I am giving them peanuts in the
shells there is often an amusing struggle to get two together in the
mouth so that they can both be held. Sometimes a squirrel will hug
the nuts in its arms next to its neck and take a few short hops to get
away and work on them at its leisure. I saw one old lady squirrel
develop an ingenious method of solving the problem. After working
with the nuts for a few seconds on several occasions she sat there and
shelled one of the nuts, put the kernels in her mouth, then picked up
the other nut quite easily. Later another learned this method and
286 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
both squirrels now do it regularly. Having learned the method, they
usually hesitate after I have given them one nut to see if I will give
them a second. Why make a trip home with only one nut when they
can just as well carry two? The second squirrel to learn this is
younger than the first one and may have observed the older one per-
form the feat, for as pointed out elsewhere I believe animals learn
much from observing others of their kind. However I saw the entire
process of learning to do the act the first time with both of them, and
thereafter it was a regular procedure to use the new accomplishment.
Captive love birds (Agapornis) sometimes place straws under the upper
tail coverts to transport them to the nesting site.
My wife recently witnessed a novel method of transportation
adopted by a yellowjacket (Vespula maculifrons). The bees were
feeding on and carrying away pieces of raisins put on our fourth-floor
window ledge for the birds. Generally they cut off a small piece of a
raisin and never carried away a whole one, but she saw one bee roll a
raisin with its head to the edge of the ledge and push it over. When
the raisin started to fall, the bee followed it a short distance, then came
back and repeated the process until it had dropped four raisins. My
wife does not think the raisin was accidentally pushed over, as the
bee’s movements seemed aimed at pushing it to the edge. Perhaps
the nest was near the building and this bee had discovered an easy
way of getting raisins to the nest.
CARING FOR AND TEACHING THE YOUNG
The type and amount of care and teaching that animal parents
give the young varies from nothing to a very good education. Any
consideration of this subject at once raises the question of how much
the animal does by instinct and how much it learns from its parents
or others. Of course, no conclusive or complete answer can be given
to this, but there are many fragments of information that we can piece
together to give us some light on the subject. Some animal mothers
never see their young and give them no attention whatever. Among
these are such animals as most snakes and lizards, which lay eggs that
are hatched by the heat from the soil or from decaying vegetation,
and the parent takes no part in their incubation. Exceptions are the
pythons (Pythonidae) and skinks (Scincidae), which incubate their
eggs, and the female alligator which stays near the nest to keep away
intruders that might harm the eggs. However, the mothers take no
part whatever in caring for or instructing the young. The mound-
building birds of Australia and New Guinea (Megapodidae) do not
incubate the eggs or care for the young; the mother lays the eggs in a
mass of vegetation which she, together with other birds of the same
species, scrape together, and in which all of them lay their eggs as in a
sort of communal incubator.
ANIMAL BEHAVIOR—WALKER 287
On the other hand many animals, particularly mammals and most
birds, give a great deal of care to the young and obviously give them
definite instructions. The bears (Ursus and Huarctos) are well-known
examples. When mother bear begins leaving the den in the spring, the
young are left inside and are apparently told to remain there. They
do not begin coming out for some time—until the mother feels that
they have developed enough to need a slightly larger world. She
first permits them to play close to the entrance to the den while she
stands guard, and later, when they are stronger, she takes them with
her on foraging expeditions, at which time she tears open decayed
stumps and logs to expose ants, grubs, mice, and other delicacies.
Likewise, she turns over stones for animal food that can be found under
them, digs up roots, and leads the babies where acorns can be found.
Mother bear is a strict disciplinarian and does not permit the young to
stray faraway. If danger threatens and she feels that for any reason
she cannot take the young to the den, she often sends them up a tree
while she stays on the ground not far away. She is usually successful
in her instructions to them to keep silent, although occasionally a baby
will become so frightened that it will ery, which often results in its
being soundly spanked and cuffed for its infraction of her rules. If
she desires not to send them up a tree but decides to run away, she
keeps them close to her, and if they are disposed to lag or become
tired, she will sometimes cuff them along ahead of her, sometimes
tumbling them end over end so that they will have an incentive to
keep up with her. Throughout this entire time, when the young are
with the mother, observers have seen that it is definitely a training
period in which the mother shows the young where and how to obtain
food, what sounds and smells to avoid, and what are apparently safe
in their haunts.
Apparently about the same procedure is followed among the foxes
(Vulpes), wolves and wild dogs (Canis), and wildcats (Felis), for the
mother leaves the little ones in the den until they are able to begin
playing about the entrance, where she finally permits them to sun
themselves and romp and engage in tussles with brothers and sisters,
gaining strength and agility. She brings them food, over which they
struggle, and finally she brings live food so that they have the actual
experience of handling live prey. Among the foxes and wolves, the
father often participates in bringing food, and both mother and father
stand guard near the entrance to give the alarm for the young to take
refuge inside when danger threatens. Usually a short bark or two is
sufficient to warn the little ones to take shelter. When they are old
enough and strong enough to venture farther away from the den, the
mother takes them on hunting expeditions on which they learn to
catch small prey that is within their strength. On these expeditions
288 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
they learn to be alert, to beware of danger, to be on the watch for
prey and how to catch it, and what to avoid.
Mother deer (Odocoileus), antelope (Antilocapra), and moose (Alces)
hide their newly born young and leave them hidden for some days.
The baby antelopes may be left in plain sight on a grass-clad or very
sparsely vegetated plain. Baby deer are hidden in the grass or sparse
shrubbery, and baby moose may be hidden in such places or in slightly
more dense vegetation. The mother then goes her way to get her
food and rest, and returns to the little one only to nurse it at intervals
of several hours. Thus the young are not exposed to the hazards of
following the mother for the first few days until they have gained
strength and are able to travel with such speed and endurance that
they stand a good chance of survival by escaping with her. The mother
cottontail rabbit (Sylvilagus) scratches out or selects a slight depression
in the soil and lines it with fur that she plucks from the underside of
her body. The depression is too small for her to be in it with the
babies, but she crouches over it, and when the little ones nurse, they
reach upward or climb up through the soft downy nest to reach her
nipples. The cottontail nurses her little ones only at rather long in-
tervals—apparently not at all during the daytime—and as long as 30
hours are known to elapse between feedings in some instances. When
danger threatens she dashes away and the enemy usually follows her.
Mother sea otters (Enhydra), which spend a great deal of time in
the ocean, lie on their backs much of the time and the babies rest on
the mother’s ventral surface. When she dives for food she leaves the
little one floating on the surface while she goes to the bottom and
picks up sea urchins and other food which she brings to the surface
and eats while lying on her back.
A mother flying squirrel that raised her family in my home, has
given me many glimpses of how she cares for her babies. Flying
squirrels are, of course, strictly nocturnal and there would be many
hazards for them in the daytime; therefore I was not surprised to find
that ‘‘Mother Glaucky” carries her babies back into the nest when
she finds that they have strayed out during the daytime. Perhaps
she and the tree squirrels teach their babies by demonstration and by
voice, but I have not been able to detect much evidence of this. The
young play among themselves about the nest, gradually gaining
strength and agility and venturing farther from the nest. The
length of time that baby flying squirrels and baby tree squirrels are
weak and uncertain in their movements and are dependent on the
mother is much greater than is generally supposed. Baby flying
squirrels do not venture out of the nest until they are about 60 days
of age and then only to explore in the immediate vicinity of the nest.
By the seventieth day they are venturing somewhat farther, but their
muscles are still soft and they have not gained agility or confidence.
ANIMAL BEHAVIOR—ERNEST P. WALKER 289
In the wild they would probably not go farther than a few feet in
their own nest tree. It is not until they are about 90 days old that
they are ready to assume full activity. Golden hamsters, on the
other hand, develop very rapidly, and by the thirty-fifth day, although
not full size, they are apparently on their own in all respects.
Swans (Cygnus, Chenopis, and Olor) are remarkably good parents
and keep close watch that their young are protected from enemies.
The female generally stays in the background and keeps the young
with her while the male goes out to meet the intruder. If the danger
is imminent he will approach with a direct rush and make vicious bites
with his beak and will strike severe blows with his wings. If the
danger is not imminent but he still feels an intruder might do harm
he frequently approaches indirectly, that is, comes up to one side of
the enemy apparently as though to catch it off guard. On one
occasion while I was sitting very quietly on a rock in the swan yard
trying to get a picture, the male swan persistently worked around to
one side of me and when I did not move he finally grabbed my arm
with his beak and tried to strike me with his wings. As soon as I
started to move away he was satisfied. In this instance I had been
trusting to my lack of motion to allay his suspicions, but my efforts
to “‘freeze’”’ were not successful. One of the poses of the male black
Australian swan (Chenopis atrata) threatening an intruder is shown
in plate 15, figure 1.
SLEEP
Generally we think of sleep as a very simple state of inactivity
which is similar in all animals, but actually the attitudes and types of
sleep of various animals differ considerably. I think it likely that
most, if not all, of the mammals that live in burrows well beneath the
surface of the ground sleep very soundly, as they are comparatively
free from danger while in their dens. Naturally, my observations of
this fact have been limited and to my knowledge it has not been
carefully studied; however, the few creatures of the burrowing type
that I have been able to study all seem to sleep very soundly. A
golden hamster can be picked up and handled gently within 30 seconds
after it has ceased activity and has thrown itself down to go to sleep.
I have similarly handled pocket mice when they were asleep in their
nests and have found that they were quite difficult to arouse, which is
in marked contrast to the great alertness of many other forms.
Animals that live above ground are, of course, subject to a wide
variety of hazards when they are asleep and must therefore, as it were,
“sleep with one eye open.” This is particularly true of rabbits, most
birds that sleep in the open and, no doubt, most other creatures that
are in similar exposed locations. My pet flying squirrels selected a
laundry bag hanging on a bathroom door for their nest and appear
290 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
not to be disturbed by the swinging of the bag, which perhaps simu-
lates the swaying of the trees in which they would normally live.
They are, however, distinctly disturbed by vibration produced by
rubbing the door or the rod on which the bag hangs, which would
probably remind them of the disturbance made by an enemy climbing
to their nest in a tree. When thus awakened, they usually react in
one of two different ways. If sharply startled they are very likely to
dash out of the nest ready to take off in a glide or run to another, safer
location; or if the alarm is not so sudden they may merely quickly
and quietly go to the entrance of the nest and look out to see the
cause of the disturbance. This behavior is, of course, associated with
the rather exposed location in which they live, where danger may
arrive from almost any direction and their best chance for survival
may be to flee.
I am inclined to think that the African elephant shrews (Macrosce-
lides rufescens) sleep with both eyes open instead of only one. I
have kept some in my den to study them at all hours of the day and
night, and I have yet to see them with their eyes closed. They are
invariably sitting upright with their eyes wide open, or at most only
slightly closed. This trait suggests that they probably sit above
ground in more or less exposed locations and are perpetually alert
for danger.
A wide variety of poses are assumed by animals in sleeping. In
addition to the well-known attitudes of lying on the ventral surface,
the back, or the side, a great many curl up in a very compact ball.
The squirrels and others with bushy tails tuck the head and feet well
inside and wrap the tail around them so that it actually affords some
protection and warmth for the back of their necks and their backs.
In this curled-up position they may lie on the side, but more frequently
the head and feet are on the underside with the top of the head actually
resting on the surfaces on which they are sleeping. This is a common
position among a great many of the rodents. The giant anteater lies
on its side, curls its head and feet together, and covers itself with
its very long-haired tail which serves as a blanket and, perhaps in
the wild, to some extent as camouflage. The sloths sleep hanging
beneath a limb with the head thrown upward and forward so that it
rests on the chest, or they may be partially sitting in the fork of a
tree with the head forward between the upper arms, the tree trunk,
and the chest. Some animals sleep standing up. Horses commonly
do this, and some elephants do much of their sleeping standing.
Most of the bats sleep hanging head downward, being suspended by
the nails of their hind feet. The red bat (Nycteris), which sleeps
hanging on a twig in a tree, has an extensive membrane between the
hind legs which it draws downward so that it serves as a protection
to the ventral parts of the body. When hanging head downward,
ANIMAL BEHAVIOR—ERNEST P. WALKER 291
bats are in a good position to take off for flight, for they are generally
at an elevated location and have merely to let go with their toes and
spread their wings to be in full flight.
I have noticed that the nine-banded, the six-banded, and the hairy
armadillos (Dasypus and Huphractus) all tremble almost continuously
in their sleep, particularly when lying on their backs or sides, as they
often do. This is unique among mammals with which I am ac-
quainted, but I have no theory to explain it.
Malayan porcupines (Acanthion brachyurum) like to sleep side by
side and have an interesting method of avoiding the spines of another
that has already lain down. Each succeeding one merely faces in
the opposite direction from the last one in the row. I have seen as
many as five lying asleep in such alternating head and tail positions,
but when I tried to take their pictures in this arrangement I usually
disturbed some of them, so that I have never been able to photograph
more than two together (pl. 15, fig. 2).
The tiny bat parakeets (Loriculus) sleep hanging head downward,
clinging to the perch by their feet.
Of all the small mammals I have observed the females are much
more particular regarding the nest than the males. The females will
move it about, cut on the nest box, assemble nest material and keep
it well shaped into a snug nest, whereas most of the males are far less
particular, usually working on the nest only enough for it to be
passably comfortable. Of course, my wife noticed this before I did
and pointed out that females have stronger instincts for home main-
tenance than males.
Most mammals, when they have the opportunity to awaken natur-
ally, like to sit and “think,” or perhaps just sit, like many people
who cannot start off ‘in high gear.’”’ Those that I have observed,
after about half an hour start normal activity.
Hibernation was discussed briefly in my previous paper on Animal
Behavior and has been extensively treated in other literature so will
not be further mentioned herein.
CONCLUSION
The better our knowledge of animal habits and behavior, the better
we are prepared to cope with the problems in connection with animal
life and the administration of resources in which animals play a part
of greater or less importance. Biologists are constantly being con-
fronted with problems of how to control, circumvent, keep away, or
increase wild life. The problems may be simple or very complex,
but are always interesting.
Wildlife administration has become an important branch of national,
State, and local government work for we have come to realize that
many of the forms are highly beneficial and should be protected and
292 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
built up to the maximum possible numbers, and that a few are injurious
and should be controlled.
As the activities of mankind are extended, the importance of wildlife
protection increases correspondingly, for we could not live without
animal life and the extirmination of any form is a serious loss. Better
understanding of animals leads to recognition of their value and there-
fore to more interest in their protection, and the study of animal life
as a profession, as a hobby, or merely through casual observation yields
much pleasure.
Norr.—All photographs are by Ernest P. Walker unless otherwise listed.
THE BREEDING HABITS OF THE WEAVERBIRDS
A STUDY IN THE BIOLOGY OF BEHAVIOR PATTERNS
By HERBERT FRIEDMANN
Curator, Division of Birds,
U. S. National Museum
[With 8 plates]
The weaverbirds, as their popular name implies, are, by and large
birds noted for the elaborate nests they build—in many cases actually
weave—out of grasses, straws, rootlets, and other similar materials.
Included in this family (Ploceidae) are some of the finest, most expert,
and most famous of all avian architects. In no other single bird
group of similar status has the habit of nest building been carried to
greater heights, indeed their only rivals for excellence in this particular
are some of the hang-nests or troupials of the New World.
My interest in the weaverbirds began some 30 years ago when I had
the opportunity of studying the actual weaving methods in captivity
of one species, the red-billed weaver, Quelea quelea. Not long after-
ward when I began my studies of parasitic birds I learned that at
least two species of weavers, and possibly several others, not only
built no nests at all, but laid their eggs in the occupied nests of other
kinds of birds to whose care the eggs and the subsequent young were
left. A few years later, over a year’s field work in South and East
Africa gave me ample opportunity to become familiar with the surpris-
ing range of aspects the nest-building habit exhibits in this family of
birds. In this paper it is not my purpose to attempt to describe each
and every one of these aspects, but to discuss them from the point of
view of the biological implications they present. ‘The family is a
large and varied one and contains a great many species (about 275),
many of which have highly divergent nesting habits, which offer very
suggestive data to the naturalist concerned with the evolution of
habits in birds.
In order to appreciate more adequately the significance of the
various aspects of the nest-building habit in these birds, and to
evaluate them more properly in connection with other parts of the
life histories of birds involved, it is necessary to make a few preliminary
293
294 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
general remarks. Birds, as a rule (there are, of course, exceptions),
go through a fairly definite cycle of behavior patterns during the year,
and repeat this cycle every year of their adult lives. Very briefly
stated, the cycle consists, in typical fashion, of the following parts:
Migration or, in the case of nonmigratory birds such as the weavers,
the fragmentation, or breaking-up, of “winter,” nonbreeding flocks
into pairs of birds; the establishment of individual breeding territories
(the extent and definiteness of which vary in different species) ; court-
ship and mating; nest building; egg laying; incubation; care of the
young; and, finally, migration or the return of the individual birds or
pairs to the “winter” flocks. Each of these parts of the whole cycle
is subject to great variation, and each may be influenced in its devel-
opment and expression by its antecedent stages, and each may exert a
similar influence on its succeeding stages.
Like any other sequence of events or stages, the undue development
of any one part may tend to throw out of line one or more of the
remaining parts. When everything runs smoothly according to
what may be looked upon as a normal pattern, it is very difficult
to observe the sequential effects of each part on its successor and it is
rarely possible to get even vague glimpses or hints of how the per-
fected whole cycle came to be developed through the ages. It is
only when something deviates from this normal pattern that we have
much chance of learning anything of the factors that control or
influence it. With these general thoughts in mind, it is instructive
to examine in some detail what has transpired in one family of birds
with respect to one part of the life cycle, the nest-building habit,
and, in connection with it, to related portions of the cycle.
The weaverbirds have been divided into a number of subfamilies
(the exact number differing in different classifications), which in a
general way are characterized by different nest-building habits.
The arboreal, so-called typical weavers (Ploceinae) construct a
suspended type of woven nest, usually shaped like a ball or a closed
oval with a lateral or a downward-extending entrance tube or
“vestibule.” Most of the species of this group are colonial, some-
times as many as 50 or more nests being built in the same tree, and
often with no others on any of the surrounding trees for considerable
distances. On the other hand, some species are quite solitary, like
Reichenow’s weaver (Ploceus reichenowi), of which usually only one,
and apparently never more than two pairs nest on the same tree, and
in those cases where there are two, only one seems to be breeding at
atime. The instinct to build is, however, very strong in most members
of this group as farasknown. Thus, Bates (1930, p. 484) noted of the
hooded weaver (Ploceus cucullatus) in Cameroon that ‘. . . it seems
to be a necessity of the birds’ nature to be always busy with their
nest; they will occupy themselves in their spare time with tearing down
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 295
unused nests, strewing the debris on the ground under the tree . . .”
Of the spectacled weaver of South Africa (Ploceus ocularius) Roberts
(1940, pp. 337-338) records that the nest has a protruding entrance
tube which is usually from 3 to 6 inches in length, but that in several
instances the nest-building activities of the birds were so great that
the entrance tubes became extended to a length of 6 feet! One
nest in the Albany Museum at Grahamstown, recorded by Stark,
has an entrance ‘‘upwards of 8 feet long.” It is a well-known occur-
rence in captivity, such as in zoologicalparks, for different species
of weavers, when given quantities of raffia or straw, to weave a vast
quantity of rather formless masses of compact, densely woven mate-
rial over the branches in the cage and even over the wire mesh of the
cage itself. The birds do so equally avidly whether they are in
breeding or in nonbreeding plumage; in other words, the urge to
build, which in most birds is seasonal and is part of the cyclical
sequence of behavior patterns, is here extended far beyond its normal
limits. Furthermore, in at least some of the species of typical weavers
(many are still very poorly known as far as details of habits are
concerned), the bulk, if not all, of the actual nest construction is
done by the males and not by the females. (In most ordinary
birds the female does most of the nest building.) Thus, in the
masked weaver (Ploceus velatus mariquensis) Taylor (1946, pp.
145-155) found that the males did all the nest building, except for
some of the lining which was put in by the females. When a nest
is completed the male that built it immediately starts to make another,
and in one colony a single male wove no fewer than 11 completed nests.
In another species, the Baya weaver of India (Ploceus philippinus),
Ali (1931) found that the males were polygamous and that the number
of mates each was able to get depended on the number of completed
nests he was able to build for them, the actual courtship and mating
behavior taking place in or around the newly finished nest. Ali
writes that in his experience with this species—
... in the initial stages of an adult nesting colony, no hens are as a rule in evidence,
and I have been unable to discover their whereabouts during the first few days.
It would appear that the instinct to breed asserts itself earlier in the adult cocks
than in the hens, for it is not until the time when the nests have progressed to a
stage where the egg-chamber is finished or nearly so, that some of the females
first become physiologically “‘ripe.”” They now visit the colony quite obviously
with the sole object of “‘prospecting”’ for laying sites, i. e., to discover if there are
any nests that are ready for their occupation. ... Two hens often fight for the
possession of an acceptable nest. The successful hen henceforward boldly enters
the nest and busies herself with finishing off and making the interior comfortable.
In no case have I been able to observe the cooperation between male and female
so often described. The lion’s share of the building—in fact all of it—is undoubt-
edly done by the cock alone. Her contribution is only the “‘interior decoration”’.. .
The “building mania,’ as it has been called, that comes over the adult cock at
this season is a sure indication of his readiness to breed...
866591—50——20
296 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
While my own field notes are less detailed for any one species of
weaver than are Ali’s on the Baya, I found males of a dozen or more
related African species actively engaged in nest building. In the
cases where my notes are fullest (the Cape weaver, Ploceus capensis
olivaceus, the black-headed weaver, Ploceus nigriceps nigriceps, and
the Kenya vitelline weaver, Ploceus witellinus uluensis) my observa-
tions indicate that the male does most of the nest weaving, if not all
of it. In some other species (Jackson’s weaver, Ploceus jacksoni, and
the spotted-backed weaver, Ploceus spilonotus) there seemed to be
more activity among the females in this regard but still the bulk of the
building was carried on by the males. (It may be that some of the
“females” were really males in nonbreeding plumage, a point that
could have been determined only by more collecting at the time.
That this is not unlikely may be adduced from the fact that in my
field notes on the masked weaver, Ploceus velatus arundinarius, I
wrote that the females take part in the nest-building activities, but
Taylor’s careful study of a slightly different subspecies of this bird,
referred to above, convinced him that the males did all the actual
construction and that the females merely added or rearranged some
lining materials. It may well be that the birds I recorded as females
were males that had not yet come into nuptial plumage.) Chestnut
weavers (Ploceus rubiginosus), watched in captivity, showed more
nest-building activity among males than females. Of Speke’s weaver
(Ploceus spekei) van Someren (1916, p. 409) noted that, ‘‘dozens of
nests are built by the male, but only one is occupied; thus there are
always plenty of old nests in all stages of completion.”
In one of the forest-dwelling, relatively solitary, or at least not highly
colonial, typical weavers, Malimbus cassini, of West Africa, Bates (p.
478) found that both sexes take care of the young. He shot a male
and a female at their nest, which had, ‘‘. .. a woven entrance tube 2 or
3 feet long, so thin that its walls were transparent, and the birds could
be seen entering and leaving, feeding young.’’ In the Cape weaver,
Ploceus olivaceus capensis, previously mentioned, Skead found that
both parents fed the young, and it appears that this behavior is fairly
widespread in the entire group.
Another section of this subfamily contains the so-called bishop
birds (Huplectes) and the whydahs (Coliuspasser and its close allies).
These birds are far more terrestrial than the members of the genus
Ploceus and their habits are somewhat different. Lack (1935, pp. 817
ff.) studied the fire-crowned bishop (Huplectes hordacea hordacea) in
Tanganyika Territory and found that the males are polygamous and
do most of the nest building, each female merely finishing or lining the
one it occupies, and each hen continuing to add to the nest throughout
the period of incubation, eventually making it so thick that the ob-
server can no longer see through it, and adding a small saclike ledge
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 297
above the entrance. One male was found to have three mates at the
same time, and may have had still others. The males have very
definite territories and the hens apparently seek out the established
cock birds. Actual mating takes place when the nest is in process of
being built by the male, but the female has, in all probability, already
settled in his territory for some time before this. Incubation of the
eggs and feeding of the young are solely the business of the hens,
which incubate chiefly at night, the warmth from the sun presumably
being sufficient for the eggs during the daylight hours.
Less complete data on the whydahs (Coliuspasser ardens and C.
jackson) suggest that these birds are monogamous. Thus, in writing
of the latter species Jackson (1938, pp. 1469-1470) states that he re-
gards the evidence against the supposed pologamy of this whydah as
conclusive. Near Nairobi, Kenya Colony, he had extremely favor-
able conditions for watching this species within a fenced-in enclosure
with open grass, outside of which enclosure the grass had been burned.
In such a small area it was easy enough to count and mark with a stick each
dancing ground. This done, the whole area was hurriedly quartered with the aid
of two boys to move and frighten away all the birds present; we then retired a
short distance, sat down and waited for them to return. The cocks very soon
appeared; the females were much more wary, but returned in due course. Some
of them settled in the grass and remained there, evidently on their nests, while
others were occupied in going to and fro with fine grass in their bills; these latter
rarely remained hidden for more than a minute at atime. Next day we returned,
and by quartering every yard of the area we discovered four nests with three
eggs, three with two eggs, one with a single egg and three not yet completed.
Each nest when found was marked by tying a knot in a wisp of tall grass close by.
At the end of a week we again returned; but no more nests were found, and on no
occasion did the females equal the number of cocks, but I accounted for this
through my failing to detect one or two of them as they sneaked back to their
nests containing incubated eggs .. .
At least it seems from this account that there were not more hens
than cocks as would have to be the case in a polygamous species.
Unlike many of the weavers whose courtship is performed at the
newly constructed nest, a group of males of Jackson’s whydah makes
a roughly circular dancing area by breaking or snipping off the tall
grass to make a clearing of from 2 to 6 feet in diameter. In the
middle of this clearing are left standing a sizable number of grasses
forming a dense tuft into which the males partly excavate little re-
cesses. Then a number of males go through a leaping peformance,
generally with no hens around to watch them. Jackson describes the
position assumed by the cock as follows:
The head is thrown back like that of a proud Turkey-cock, the beak being held
horizontally; the feet hang downward, the tail is held straight up till it touches
the ruff at the base of the head and neck, the ends of the feathers falling in a curve
downward, with the exception of two tail-feathers which are held straight out
and downward.
298 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
While actually rising in the air the half-open wings are worked with a very
quick shivering motion, and the feet are also moved up and down very rapidly,
beating the air.
The bird springs straight up in the air, sometimes for a few inches and sometimes
to the height of two or more feet, and then drops.
The whole of the plumage is much puffed out throughout the performance,
which is repeated five or six times, with a short interval for rest.
The game would appear to be somewhat fatiguing, as the bird rarely makes
more than five or six jumps at a time without a short rest . . . They very often
assume their curious jumping attitude some little distance before they arrive at
their playground. . .
Besides the data quoted above it may be added that apparently
but one male may make use of one dancing area, and that often at the
end of the jumping dance it appears to try to burrow into the shallow
recesses of the central tuft of grasses (as though there were nests
there).
Jackson’s whydah shows the courtship behavior pattern developed
to a greater degree of display and ostensible rivalry between males,
and to a greater specialized areal usage (courtship, or dancing, ground
as contrasted with nesting site or even breeding territory) than others
of its relatives, but the difference seems to be more one of degree than
of kind. As typified by its habits, this section of the terrestrial Plo-
ceinae may be said to be characterized by elaboration of courtship
behavior from individual displays near the nest to a complex display
at a dancing ground, and by what seems to be monogamy. (More
detailed information is needed on this point, however. In the case of
the black whydah, Coliuspasser concolor, and of the red-collared why-
dah, Coliuspasser ardens, there are observational data supporting a
monogamous state; in the long-tailed wydah, Coliuspasser procne,
similar but less extensive data suggest polygamy.) Recently V. D.
van Someren (1945, pp. 131-141) has concluded that Jackson’s why-
dah is polygamous, but his own presentation of the case is not too pos-
itive. Thus, he writes that—
. polygamy appears to be general, and seems to arise because of the imper-
fect correlation between the maturation of the males and females. Some males
mature early, others late, and the early males may cease dancing and start moult-
ing while later males are just beginning to assume breeding plumage and dance.
This irregular maturation of the males may be spread over several months, while
by contrast, the females mature almost simultaneously, and all nests are found
at the same stage of building or incubation within a few days. Since the sex-
ratio of the mixed flock is almost 50:50, late maturing males are thus able to
mate with several females, because the mature females probably now outnumber
the mature males. Males may commence dancing some four months before the
first nest is found, but these early males are probably unsuccessful at mating be-
cause of the unready state of the females. Males may start dancing while still
in non-breeding plumage, but the behaviour pattern of these immature males is
undeveloped in several respects.
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 299
It may be noted that the assumption that late-maturing males are
able to have several mates is based on the thought that more females
are mature in the later than in the earlier part of the season, but, only
a few lines above, van Someren informs us that all the females mature
simultaneously, which would imply that the females mature later
than the early-maturing males. It would seem, from this, if van
Someren’s assumption be correct, that the early-maturing males might
be no longer in breeding condition late in the season when mates are
available, or else they would compete with the later-maturing cocks
for the hens to a degree sufficient to diminish the likelihood of the
late-maturing mates having more than a single mate apiece.
The care of the eggs and the young appears to be left wholly to the
females, and, as far as the incomplete evidence goes, the actual nest
building is also done by the hens of the various species of whydahs.
We have but little reliable data as to the territorial aspects of the
lives of most of these birds, except for Jackson’s whydah. In his
study of this bird van Someren found that—
. . . true territorial behavior becomes evident early in the sexual break-up of the
flock. The males, isolating themselves on rings (7. e., dancing areas) establish a
well-defined territory of small extent, of which the ring itself is the focal point;
the territory extends all round the ring at a radius of 6 to 10 feet from the central
tuft.
A female alighting anywhere within this territory may be solicited by courtship
behavior by the male on the ring, even though she may not alight on the ring
itself. Another full-plumaged male alighting on this territory is treated in one of
two ways, depending on the attitude of his tail as he alights. If he alights with
his tail arched and the two outer plumes drooping as in the dancing attitude, he
is attacked with pursuit flight if the owner is present in the territory. If however,
the intruding male alights with his tail folded in the normal flight attitude he is
usually solicited and displayed to be the owner as if he was a female. It is very
noticeable that when a male returns to his territory from outside it, the tail is
arched and the two outer plumes drooped the moment he crosses the boundary; the
bird alights in the dancing attitude, and thus shows his ownership by his
appearance...
Where two or more rings are found within a few inches of one another...
they are all formed by the one male, who may use them alternately while dancing,
and keep them all in good order . . . rings occupied by two separate males are
not found closer than about 12 feet. These boundaries are accepted by the other
members of the flock early in the break-up, hence territorial squabbles are seldom
seen late in the season . . . This territory is related purely to sexual functions
and has no food significance; feeding is carried out in a mixed flock even in the
height of the dancing season, on neutral ground where sexual rivalry is notably
absent.
Furthermore, this territory appears of no significance to the females apart from
the fact that they are attracted to the rings; they are unaware of the boundaries
of the male territories. At nesting time, the males cease dancing vigorously and
the main dancing area may become completely deserted; the females nest in a
different area which is usually some distance away from the dancing grounds. The
300 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
nests tend to be grouped together, and are usually about 20 to 30 feet away from
the nearest ring if males have been dancing previously in the neighborhood, i.e.
well outside the territory boundaries.
In connection with the development of courtship posturings, it may
be pointed out that, unlike the members of the arboreal Ploceinae,
the whydahs, and, to a somewhat lesser extent, the bishop weavers,
have very marked sexual dimorphism in the breeding plumage of the
adults and many species have elongated or otherwise specialized
plumes in the nuptial dress of the males. In the nonbreeding season
the sexes generally look alike.
Before going on to the next subfamily, we may briefly summarize
the situation in the typical weavers (Ploceinae). Many of the species
are colonial (which means, in effect, that in most of them the individual
nesting territories are nonexistent), and in those species that have been
most fully observed the nests are built largely or wholly by the males.
Furthermore, in some forms it seems that the males are regularly
polygamous and that the number of mates each one acquires depends
on the number of nests he has been able to provide for his mates.
From the standpoint of our hypothetical “standard” picture of the
cycle of breeding behavior patterns this implies that the first stage—
migration or the fragmentation of flocks into individuals or pairs—is
omitted, that territoriality is likewise skipped, and that the usual
sequence of courtship and nest building is reversed. As we have noted,
the actual courtship and mating takes place in and about the newly
completed nests the males have constructed. The Ploceinae take 2
years to acquire adult plumage and to come into breeding condition.
This summary is correct as far as it goes, but there is still one more
variation in the reproductive behavior pattern exhibited in this sub-
family. One species, sometimes called Rendall’s seed-eater, some-
times (and more properly) the cuckoo finch, Anomalospiza wmberbis,
a bird with no very close relatives, but apparently nearer to the bishop
weavers than to any other assemblage, is wholly devoid of any nest-
building or incubating or rearing instincts, and is, in short, parasitic.
It is still very imperfectly known, and all that may be said with any
certainty is that it is parasitic on small ground-nesting (or near the
ground nesting) warblers of the genera Cisticola and Prinia. (Six
species of the former and one, possibly two, of the latter genus are
known as hosts of the cuckoo finch.) As Delacour (19438, p. 71) has
recently pointed out, the fact that Anomalospiza agrees with the
Viduinae in being parasitic does not necessarily imply close relation-
ship to the members of the latter group. In most respects it seems
best placed, taxonomically, with the Ploceinae, but, it must be ad-
mitted, is an aberrant member of that group. It is aslightly gregari-
ous bird, living, at least in the nonbreeding season, in loose flocks.
Nothing is on record concerning its courtship, sexual relations, or
territorial habits.
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 301
The next subfamily is a much smaller assemblage—the buffalo
weavers (Bubalornithinae)—containing only two species, each with
several races. Unlike the typical weavers these birds do not construct
nests of fine weaving and elaborate structure, but make rather bulky,
massive nests of twigs and thorny branches, rough and untidy in
general appearance, and not suspended from, but placed on top of,
the branches of large trees (in my experience often baobabs). The
birds are colonial and the nests are often placed so close together that
they are actually in direct contact with one another. Jackson (p.
1380) records that he saw where branches had given way under the
combined weight of too many of these nests on several occasions.
Brehm found as many as 18 nests in 1 mimosa tree in northeastern
Africa. The nests are often 2 or 3 feet across and are used and
repaired and added to year after year. Each nest of the black buffalo
weaver (Bubalornis albirostris) contains two or more chambers, lined
with grass and straw, each with an entrance facing away from the
other. The only nests I ever saw of the other species, the white-
headed buffalo weaver (Dinemellia dinemelli), had but a single cham-
ber. There are descriptions in the literature of nests of Bubalornis
containing more than two entrances. Priest (vol. 4, p. 220) writes
of one that “‘. . . there were numerous entrance holes, and it looked
as if about a dozen birds lived in each of these communal nests . . .”
That the urge to build is extended in these weavers beyond the usual
small part of the annual cycle of most birds, as it is in the Ploceinae,
is indicated by some observations made in Darfur, in the Sudan, by
Lynes, who noted that “‘. . . at all seasons we found these Textors
(=Bubalornis) hanging about their everlasting great nest-clusters,
into which, even in midwinter, birds with quite inactive sexual organs
would sometimes carry twigs as if nesting . . .”
Many years ago, in southwestern Africa, Andersson described the
nests as follows:
The collective nests consist externally of an immense mass of dry twigs and
sticks, in which are to be found from four to six separate nests or holes of an oval
form, composed of grass only, but united to each other by intricate masses of
sticks, defying the ingress of any intruder except a small snake. In each of these
separate holes are laid three or four eggs . . . I obtained no less than forty of
these eggs . . . and on the following day the birds were busy in repairing one of
the collective nests which had been injured during the collection of the eggs .. .
I believe these nests are annually added to, for, so far as I have been able to see,
the same nest is retained for several consecutive seasons.
We do not have nearly as complete information on the buffalo
weavers as on the typical ones, but what data are available indicate
that the birds are colonial, that there is little or no observable evidence
of any prenuptial fragmentation of wintering flocks, and that the
males do at least part of the nest building. Whether they do it all
or not is still unknown.
302 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The evidence does suggest that the nest-building habit is indulged in
out of the breeding season and by nonbreeding birds; not too different
from the typical weavers in this respect.
The next subfamily, the sparrow weavers and social weavers (Ploce-
passerinae), shows a great indulgence in nest building, in and out of
the breeding time, which culminates in the truly gigantic communal
structures of the social weaver (Philetairus socius). Thus, of one
South African sparrow weaver, Plocepasser mahali, Roberts (1940,
p. 332) writes that—
. a single pair of birds will construct as many as a dozen nests of whitish grass
stems on the projecting branches, these being arched over the top with two
entrances below on opposite sides, so that there is no cavity for the eggs and
evidently made for amusement only; the nests in which the eggs are laid have
only one entrance and are warmly lined with feathery grass tops .. .
A somewhat different description is given by Stark (1900, pp. 84-85)
who informs us that the species is—
. of social habits, it remains in flock all the year round and breeds in company,
several nests being generally built in a single tree. Rarely have I met with more
pugnacious birds; the males in spring are constantly fighting, and so desperate
are their quarrels that the combatants frequently lie exhausted, side by side, on
the ground, incapable for further movement ... The nests are large, roughly
built, kidney-shaped structures, usually placed near the ends of the branches of a
mimosa or other thorny tree. They are constructed of long grass-stems, the
blades and flowering tops being woven together, the stiff stalks projecting in all
directions. During the winter each nest has two entrances from below, separated
in the interior by a narrow bridge of grass, on which the birds roost. At the
beginning of the breeding season one entrance is stopped up with leaves and grass,
a shallow cavity being left in which the female deposits two or three eggs...
As soon as the young are on the wing, the second entrance is unstopped, and the
nest is again used, both by the old and young birds, as a roosting place. The
nests are annually repaired and last for many years.
A somewhat similar account holds for another species, the gray-
headed social weaver of East Africa, Pseudonigrita arnaudi. Jackson
(p. 1384) comments on the remarkable nest of this bird—
. . . it is very large and quite exceptionally compact, and has two entrances
pointing downwards. During the period between breeding seasons these nests
are used for roosting, the birds resting on the ridge between the two entrances.
In the breeding season one of the holes is stopped up and the eggs are deposited
in a depression beyond the ridge on the material used for stopping up the second
entrance. The nest is firmly woven to several twigs or branches . . . in small
clumps of five to eight nests together.
Of a slightly different race of the same species Jackson found (p. 1385)
the nests ‘‘. . . were packed together so closely as to form almost one
compound nest.”
Probably the most remarkable of all weaverbirds’ nests is that of
the famous sociable weaver of the western arid portions of South
Africa, Philetairus socius. The enormous communal nests built by
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 303
these little sparrowlike birds attain truly great proportions—as much
as 25 feet long and 15 feet wide at the base and from 5 to 10 feet in
height! While each nest is the product of not a lone individual, or
even a pair, but of a whole flock of as many as 75 or 80 pairs, still the
sheer bulk of the nesting material gathered and placed by the birds
is a striking testimony to the tremendous year-round urge of the nest-
building instinct in this species. The time I spent studying this bird
in the western Transvaal in 1925 was one of the most fascinating
experiences an ornithologist could have, and I cannot refrain from
including here part of my notes, at the expense of repeating some items
already published in an earlier paper (1930).
As the common name of the bird implies, Philetfairus is very social
in its habits; in fact it is probably as social as any bird could possibly
be. It is always found in flocks, feeds in flocks, and breeds in large,
many-apartmented compound nests. The smallest flocks that I saw
contained about 20 birds; the largest one at least 150. The flocks
seem to stay pretty much in the same general vicinity all the year
round, and the birds use their huge, massive nests as roosting places
during the nonbreeding season. With this extreme sociability and
sedentary habit of life the territorial relations of the species have-been
modified in a way that is quite remarkable, perhaps unique among
birds. Instead of each pair of birds having its own breeding territory,
each flock seems to have a definite territory, and as the individual
flocks are usually far enough apart not to compete with each other,
the boundaries of these territories are seldom crossed by individuals
of other flocks and other territories. However, in a few cases in my
own experience two flocks were fairly close together (i. e., two nests
were on trees not very far apart), and the birds mingled more or less
while feeding, but in these cases far more fighting and quarreling was
observed than in all the others together. In an area approximately
100 miles long and 10 miles wide, or 1,000 square miles in all, I found
only 26 nests of the social weaver, so it can be seen that the flocks
ordinarily do not live in very close juxtaposition to each other. (The
nests are so large, and so conspicuous at great distances, and the
country so open and easy to examine, the trees being so relatively few
in number, that I am quite certain I found practically every nest in
this area.)
The nests observed varied in size as did the flocks. The smallest
nest found measured about 3 feet in diameter at the base and was
about 3 feet high and had about 10 entrances on the under surface,
indicating that it contained that number of individual nests. The
largest one found was incomplete, i. e., a piece of it had broken off,
breaking its supporting branches by its weight, but the remaining
part was a large, flat, horizontal mass of straw, more or less repaired
at its broken edge, and measured about 25 by 15 feet at the base and
304. ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
was about 5 feet high. The part that had broken off must have been
about 5 feet in diameter each way. This nest contained about 95
nests within it.
In a locality where these birds occur it is impossible to remain long
unaware of their presence. ‘Trees are not so numerous but that each
one becomes an object of more or less importance in the landscape.
Needless to say a tree on which there is a social weaver’s nest is a very
conspicuous object, visible for a great distance and widely proclaims
the presence of the builders. But the birds themselves soon intrude
upon one’s consciousness with their noisy, harsh, chattering notes as
they fly by in flocks or feed in scattered bunches upon the seeds of the
small, stunted shrubs and plants that wrest an existence from the
inhospitable soil. While feeding they keep up an incessant chatter
much like a flock of house sparrows, and, like them, frequently quarrel
over bits of food. In flight they all act in unison with a precision
quite remarkable for birds of their type, the whole flock turning,
rising, falling, wheeling, and stopping more or less together.
Although the birds live in compound “‘apartment-house’’ nests,
feed and fly in flocks, and are at all times exceedingly gregarious, they
seem to establish fairly strong mating relations as far as my field
observations indicate. If they were haphazardly promiscuous they
would be forever in each other’s way getting in and out of the entrance
holes of the individual nests in the large communal structures. As a
matter of fact, the harmony of life within each colony, the lack of
what may be likened to traffic congestion, i. e., the coming and going
of birds in the task of providing food for the young, the fact that out of
numbers of individual nests examined by various observers none were
found with unusual numbers of eggs or young, all argue for individu-
ality in nest occupancy. Whether each male has only one or several
mates is, however, unknown.
There have been several attempts to explain the structure of the
large, composite nests of this species, some writers claiming that each
pair of birds builds an individual nest, all of them close together, and
then the flock builds the common roof over all the nests, while other
writers have recorded that the flock builds a large structure and then
each pair builds its individual nest into this structure. J never saw
the actual beginning of a nest, and the smallest nests I found were, as
mentioned above, complete structures with numbers of nests within
them. However, Roberts (1940, p. 333) describes the construction
of the communal nest as follows:
. . . first a roof is constructed of coarse straws in the strong branches of a large...
tree, and under this a great number of nest-chambers are made by nipping off
the straws to form a tunnel upwards with a chamber at one side of the top of the
tunnel; each pair of birds has its own nest-chamber, and scores of pairs may
occupy the same colony.
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 305
To this I may add that regardless of how the first start is made, it is
true that all through the nonbreeding season, the entire flock does a
large amount of roofing and general enlarging of the whole affair so
that it is true that subsequent individual nests are built into the large
structure. The nests are added to year after year, and frequently
become so large and heavy that they break the branches upon which
they rest, and crash to the ground. All the birds seem to work
together equally, apparently the males as well as the female(?)s, and
even during the breeding season, when they have eggs or young in
the nest, the male birds may be seen carrying straw to the roof or
other parts of the common structure, not necessarily close to their
own respective individual nests. The huge, massive affairs are
composed wholly of dried grasses of a rather coarse, tough sort that
grows commonly in southwestern Africa, and the seeds of which
enter into the diet of the weavers very largely. The material is not
really woven or even plaited on the surface of the nest, but is rather
roughly put together in about the same way that hay is put into a well-
made hay rack, but with a fairly definite thatching arrangement,
causing the rain to run off and not soak through. The under side
of the nest presents the rough, hard ends of the coarse straws and forms
a very uneven surface.
In the sparrow weavers and social weavers (subfamily Plocepas-
serinae) we find, then, as far as our incomplete data permit us to
generalize, an annual behavior cycle characterized by lack of migration
or winter flock fragmentation, a substitution of a communal flock
territory for individual ones as far as nesting is concerned, and a very
marked development, both in seasonal duration and in individual
activity, of the nest-building habit. The published observational
data indicate that both sexes participate in nest building, but these
data are open to question because of the similarity in plumage of the
males and females; whether one or the other does most of the con-
struction is not known. Nothing appears to be on record concerning
the courtship habits, so it is not possible to ascertain whether this
part of the cycle comes before or is associated with already completed
nests as it is in some of the typical weavers (Ploceinae).
Turning now to the next group, the weaver finches (subfamily
Passerinae), which group includes the ubiquitous house sparrow
(Passer domesticus) and its relatives, we find a different range of nest
types. Some, like the house sparrow, build fairly bulky, formless,
untidy nests in trees, on ledges, cornices of buildings, even in holes
in trees, and other elevated sites (never on the ground). When
built in the branches of a tree the nest usually is domed with an en-
trance on one side, and fairly abundantly lined with feathers and other
soft materials; when built in a hole the lining is much reduced as is
306 § ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
the rest of the nest structure. The birds are multiple-brooded; both
sexes take part in nest building and in caring for the young.
Although the nests are not such as would, in and of themselves,
suggest that their makers were overly involved in building activities,
there is evidence that in a closely related species, the Cape sparrow
of South Africa, Passer melanurus, the birds use the nests throughout
the year as sleeping places “ . . . especially in winter, when nests with
more warm material are often specially built for the purpose” (Roberts,
1940, p. 334). In other words, in this species we find some indication
of nest-building activity outside of the breeding season. Whether
this is true for other forms of the genus is not known.
To return to the house sparrow, the reproductive behavior cycle,
as reported by Jourdain and Tucker (1938, pp. 157-158), is quite
peculiar and is still in need of further study before it can be properly
interpreted. The—
. . . prominent feature of breeding-season is noisy display, in which sometimes
one, but commonly several males hop with loud chirpings, round female with
elevated bill and tail and drooping wings, but merely elicit pecks from irritated
hen . . . Whole performance commonly ends with sudden dispersal of participants
and appears unconnected with coition or even pairing. Gengler relates latter to
rough-and-tumble scrimmages between several males without display, female
commonly becoming involved as well, though selection of mates as result of these
tussles seems not very clearly demonstrated. Coition is normally solicited by
female with drooping wings and twittering note, without display by male, and
may be repeated as many as a dozen or fifteen times in succession. Same observer
states that both mated and unmated birds of both sexes are involved in displays,
but that mated males display only to other females, never their own. He inter-
prets display as relict of former genuine courtship, now functionless except as
outlet for persistent display instinct . . . exceptionally coition may be preceded
by typical display of male without usual solicitation of female.
There is evidence, as well, that the species has a polyandrous or promis-
cuous tendency, and Thompson (quoted by Jourdain and Tucker)
considers the noisy displays are explained partly by this tendency,
and partly by the males coming into breeding condition before the
females.
Other members of the subfamily, such as the yellow-throated spar-
rows of the genus Petronia, the rock sparrows of the genus Gymnoris,
and the gray-headed sparrow, Passer griseus, appear to nest chiefly
if not wholly in holes in trees, in old woodpecker or barbet holes, or
even in suitable natural holes of not too large a size. They generally
line these nesting holes with fibers and feathers. The gray-headed
sparrow has adapted itself to human habitations and frequently
nests under the eaves of buildings. The chestnut sparrow, Sorella
eminibey, not infrequently makes use of old nests of other weaverbirds
although it does at other times build for itself.
The absence of adequate data on the members of this group, other
than the house sparrow, makes it impossible to generalize on any broad
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 307
scale or sure foundation. It is safer, then, merely to summarize the
picture in the one relatively well-known member. The picture of
courtship display is markedly altered from what we are in the habit
of considering normal for most passerine birds—males displaying to
any females but their own mates; females apparently soliciting rather
than permitting coition, a precarious monogamy with a tendency
toward polyandry and promiscuity.
The scaly weavers of the genus Sporopipes form a subfamily by
themselves, the Sporopipinae. They are not too well known, but I
have found them in very loose flocks or small assemblages in the dry
thornbush veldt of the Transvaal, where they feed on the ground
like the Passerinae. The South African species (Sporopipes squami-
frons) breeds during the southern winter as a rule, but at times during
the summer as well, suggesting a not too well delimited nesting time.
These birds are not colonial breeders, but build their roughly globular
nests of grass stems and fine twigs, with a fairly pointed lateral en-
trance, in the middle of the dense thorny branches of shrubs and low
trees. Two nests that I found were less globular than published
descriptions indicate is usual. They were somewhat similar to the
untidy structures of the house sparrow, but smaller, slightly more
compact, and less irregular in shape. In my field notes I described
them as horizontal cylinders rather poorly closed at one end, and made
of grasses, fine twigs, straws, etc. One nest containing three eggs
was being very timidly guarded by two of the birds, presumably a
pair (the sexes look alike). The birds would not stay near the nest
while I was close to it, but returned to it as I walked away. Nothing
seems to be on record concerning courtship, mating, or territorial
behavior in any of the scaly weavers.
We now come to the subfamily Viduinae, the indigo birds and the
widow birds, containing a dozen species, three of which are definitely
known to be parasitic and the others are suspected of having similar
breeding habits. This group is somewhat intermediate between the
Ploceinae and the next subfamily, the Estrildinae. Like the mem-
bers of the Ploceinae, the Viduinae take 2 years to acquire adult
plumage, and do not breed until then (the Estrildinae breed when 1
year old, as do the majority of small passerine birds). The adult
males have a breeding plumage in which they are very different from
the brown, streaky hens and year-old birds, the former of which they
resemble in the nonbreeding plumage. (The Estrildinae do not
show any seasonal plumage change as arule.) The best known of the
Viduinae is the pin-tailed widow bird, Vidua macroura, and the fol-
lowing description of its habits is taken largely from my field notes
coupled with pertinent data in the literature.
Vidua macroura is a gregarious bird and is usually seen in flocks of
from 5 to 50 birds, depending on the season. In the breeding season
308 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
in South Africa, where seasons are definite, the flocks tend to break
up and the birds pair off more or less. Yet it is not uncommon to
see small flocks all through the breeding season. Such flocks usually
contain but one full-plumaged male and the rest. of the birds are in the
brown hen type of plumage. In some cases I shot into the flocks and
found that the brown birds were year-old males, but in two cases the
birds proved to be females with fairly enlarged ovaries. It seems,
therefore, that this bird is somewhat polygamous, although I should
judge from most of the cases I have observed (and they are many)
that it is frequently, if not usually, more or less monogamous. In
equatorial Africa all the individuals of the species in any one locality
do not breed at the same time and these flocks usually contain a
breeding pair and either year-old birds or nonbreeding adults. The
lack of definite seasons complicates things superficially to the extent
that the apparent state of affairs has no real relation to the actual
conditions.
This widow bird is largely terrestrial in habit and gathers most, if
not all, of its food on or near the ground. However, in Natal, at
least, during the southern winter the birds go about in large flocks and
spend much time in the trees, where they act and sound not unlike
small finches such as the North American redpoll, Acanthis linaria.
They are by no means confined to trees and are found in tall grass
and in reeds along stream banks. During the breeding season the
males often use isolated trees as perches from which to sing and to
watch over their territories, but the birds spend by far the greatest
part of the time on the ground.
On November 24, at Woodbush, Transvaal, I saw an adult male
in full breeding plumage. It was perched on a bush in an open
grassy field, and as I approached it flew off to a nearby bush and then
to another not far off as I came close again. It made a small circling
flight and came back to the original bush. On and off during the
rest of the day I found it there each time I visited the spot and found
by repeated trials that it could not be induced to leave it. It had
definitely established its territory there, and apparently the bush in
which it was first found was its singing perch. The next day I spent
a couple of hours watching it and tried to make it fly off, but it would
not go more than a hundred feet and then circle back gradually.
There was a single hen bird in the immediate vicinity. I shot the
male and found the testes were much enlarged. The plumage was
still very fresh; in fact the long central rectrices still retained a little
of their sheaths and one of them was so loose that it came out when I
skinned the bird.
In the same region I watched two other males that were established
in their individual territories. One of them was watched for 3
successive days and was apparently without a mate as yet. It had a
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 309
territory about 400 yards in diameter, considerably larger than that of
the first male, but more open, less bushy, and probably contained
possibilities of no greater number of nests to parasitize than the other.
The third and last male had a smaller breeding area and was usually
accompanied by three or four brownish henlike birds. I shot one of
these birds and found it to be a male—a year-old bird in first nuptial
plumage.
The courtship displayed was first observed at Woodbush, Transvaal,
on December 1. The male flew up from the ground and hovered about
2 feet in the air directly over a female, with his body feathers shghtly
ruffled and his wings beating rapidly. With each wing beat the four
long rectrices were violently jerked and made to stream boisterously
over the female, much after the cascade type of tail display of Coli-
uspasser ardens and Coliuspasser procne.
On another occasion, in equatorial East Africa, I saw a male display
to a female that was perched in a thorn tree. The display was similar
to the one already described; the male danced in a stationary posi-
tion as though suspended in midair a couple of feet above the female.
On still other occasions I watched males courting when there were
several of the brownish hen-feathered birds present. In all such
cases I noticed definitely that the male tended to confine his atten-
tions to one particular bird. It seemed as though there was but one
female and that the other brown birds were year-old males. In one
case I shot the whole band (five brown birds) and found that one was a
female in breeding condition and the rest were young males.
Inasmuch as this widow bird is parasitic in its breeding habits it
is interesting to compare it with the cowbirds (Méolothrus) of the
Americas. The chief difference seems to be in their sexual relations.
Both are more or less monogamous but the Vidua tends toward
polygamy while the Molothrus tends toward polyandry.
The vocal efforts of this species are not remarkable. The usual
call notes are weak, high, but sharp tsips, something like the weaker
notes of the redpoll (Acanthis linaria). When a band of birds calls
simultaneously and rapidly they produce a light twittering chorus.
The song is a rapid but modulated repetition of the call note and
usually consists of from 5 to 10 syllables and occasionally more.
It is given in flight as well as when at rest. Curiously enough, I
never heard a male sing while going through his display dance before
a female.
As is well known, this species is parasitic in its breeding habits;
i. e., it lays its eggs in the nests of other birds and leaves them to their
care. Vidua macroura is not the only ploceid exhibiting this habit—
V. regia and V. paradisea and, as we have already noted, Anoma-
lospiza imberbis are also parasitic, and probably the other species
of Vidua will in time be found to be parasitic as well. Vidua
310 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
macroura is parasitic chiefly on waxbills and generally lays but one
egg ina nest. I have seen sets containing two, three, and even, four
of the widow bird’s eggs along with those of the victims, but such
sets are not usual. The eggs are pure white and differ from those of
the common fosterers only in size.
The following birds have been found to be parasitized by Vidua
macroura:
Lonchura scutatus Estrilda rhodopyga
Estrilda astrild Estrilda delamerei
Estrilda subflava Lagonosticta senegala
Estrilda melpoda Lagonostica rubricata
Estrilda massaica Amauresthes fringilloides
Estrilda melanotis Coliuspasser ardens
The incubation period is 12 days.
The breeding season in South Africa is late in the southern summer—
January, February, and early March, sometimes earlier. In Kenya
Colony the species breeds during both the short and the long rainy
seasons. The short rains come in November, December, and Jan-
uary; the long rains in April, May, June, and July. As one goes
northward the rains shift to later in the calendar year; thus in the
southern Sudan the long rains extend into September and start
correspondingly later than in Kenya Colony.
The young V2dua does not always crowd out or starve out its nest
mates (at least in the few cases I watched) as do the young cuckoos
and cowbirds in so many cases, but all grow up together. Fully
fledged young Vidua macroura are often found in flocks of young
waxbills after leaving the nest but they do not remain long in these
assemblages. Before they get ready to molt (postjuvenal molt)
they form flocks of their own. I have seen as many as 15 or 20
young pin-tailed widow birds together. Frequently one or two adult
birds, often males in breeding plumage, are found in these flocks.
My observation on Vidua regia, the shaft-tailed widow bird, Vidua
jischeri, the straw-tailed widow bird, and Vidua orientalis, one of the
indigo finches, while much less complete than those on Vidua macroura,
also indicate that the superficially apparent polygamy is actually not
real, that while one male in adult breeding plumage may be accom-
panied by a small flock of brown henlike individuals, most of the
latter are immature birds of both sexes and only one in a group may
be an adult female. In the case of the straw-tailed widow bird,
Vidua fischeri, I once observed what seemed to be a territorial fight
between two males in full breeding plumage.
To summarize the behavior-pattern cycle in the Viduinae, we may
characterize it as follows: apparently monogamous and solitary(?), but
solitary only with respect to its own age group (adults), not solitary
CLIGI ‘ZG “1OA ISI] “YBN ‘snyy ‘sowy *[INg ‘urdeyg wo17)
SNWIMYADIN SNADO1d AO ANONOD ‘Z SNLVYIINOND SNADO1d AO ANONOD *}
\ SH abe uuPWpallJ—"6p6| ‘ode UeluOsy WIG
SHONVY LNA
ISAN IVAGIAIGN| MOHS OL LSAN 4O ACIS YSGNN ‘Zz SNIDOS SNYIVLATIHd 3AO LSAN ‘1
¢ ALV 1d uuewipatl.J—6p6| ‘Oday uPIUOsYyzIUIG
LSAN TIWNOWWOD SIL
-NVSIDS S3HL AO YANHOD V YSGNN SHAGTING AHL AO ANO “2 SNIDOS SNYIVLATIHd AO LSAN TIVWWS ATYIVSA GNV MAN ‘1
© saLvad uuewipalJ—¢6p6| ‘Woday weruosyytWIG
Smithsonian Report, 1949.—Friedmann PLATE 4
1. A VERY LARGE OLD NEST OF PHILETAIRUS SOCIUS, PARTS OF WHICH HAD
FALLEN DOWN BY THEIR OWN WEIGHT
oe sss a
2. THE SOCIAL WEAVERBIRD, PHILETAIRUS SOCIUS
Smithsonian Report, 1949.—Friedmann PLATE 5
foe
—
>
j
Pd
1. NEST OF PLOCEUS OCULARIUS
(From Chapin, Bull. Amer. Mus. Nat. Hist., vol. 37, 1917.)
2. NEST OF EUPLECTES FLAMMICEPS
(From Chapin, Bull. Amer. Mus. Nat. Hist., vol. 37, 1917.)
Smithsonian Report, 1949.—Friedmann PLATE 6
Ry
Upper, dancing ground of Coliuspasser jacksoni; middle, male Coliuspasser
jacksoni on its dancing ground; lower, male Coliuspasser displaying to female
on dancing ground.
(All photographs on this plate from Van Someren, Journ. East Africa Nat. Hist. Soc., vol. 18, 1945.)
Smithsonian Report, 1949.—Friedmann PLATE 7
1. CUCKOO FINCH, ANOMALOSPIZA IMBERBIS
(From Shelley, Birds of Africa, vol. 4, 1905.)
’
2. YOUNG ANOMALOSPIZA BEING FED BY PRINIA FALVICANS
(From Roberts, Ann. Transvaal Mus., vol. 5, 1917.)
VISSaY VNGIA AO ANO AGNV NMO
Sl] 340 S999 SSaYHHL HLIM SNVSOIAV 14 VINIYd AO LSAN °Z SNLONOTMIdS SNADO1d AO ANONOD °|
8 31V1d uueulpaliJ—'6p6| ‘j40dayy ueluosyyIUIg
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN oll
as far as immature “hangers-on” are concerned; courtship display well
developed in all species; nest-building, incubation, and rearing instincts
completely lacking in three members (V. macroura, paradisea, and
regia) and probably in the others as well. The young of the three known
parasitic species do not seem to evict or to starve out their nest mates
of the host species, but may grow up in apparent amity with them.
Roberts (1939, pp. 106-107) finds that while this is so, the female
parasite usually destroys an egg of the host when depositing its own
in the nest, but no such observations have been published. Usually
there is but a single egg of the parasite in any one nest, but Roberts
has found one instance where “‘five eggs of the common waxbill were
all replaced by eggs of the Pin-tailed Widow-Bird.” Delacour and
other writers have implied that the Viduinae are parasitic chiefly on
waxbils, and even go so far as to suggest that each of the Viduinae has
its particular Estrildinae host species, but this is by no means definitely
established. Thus, the pin-tailed widow bird is known to parasitize
at least nine species of Hstrilda and Lagonosticta, and two ploceine
weavers, Coliuspasser ardens and Amauresthes fringilloides, while there
is some evidence that Vidua regia lays its egg in the nest of a warbler,
Prinia flavicans.
The last subfamily of weaverbirds, comprising the waxbills, grass
finches, and mannikins, is the Estrildinae. Delacour (1943, pp. 71-72)
has recently summarized the characteristics of this group as follows:
Small weaver-finches of highly specialized color pattern, never showing a primi-
tive streaked sparrow-like brown plumage and horn-colored bill; sexes alike or
different; immature always different from adult females. No eclipse plumage in
males, with one exception. Nestlings always showing brightly colored, swollen
spots, lobes or bands at the gape, and an ornamentation of the tongue or palate,
consisting of spots or lines. Eggs numerous and always white; nests globular
with a side entrance, but not woven. Young birds become adult within a year
of their birth and are then able to breed, while it takes two years for young
Viduinae and Ploceinae to mature. Peculiar song and courtship variable but
consistent, in a general way, in large groups of genera. Ten primaries in the wing,
the first being very short and falcate, with the exception of two genera (Clytospiza
and Spermophaga) where it is moderately long, not parasitic.
This large group is composed of three natural subdivisions: the wax-
bills, chiefly found in Africa, but with one genus in Asia; the grass
finches, found in Australia and some of the islands of the south Pacific;
and the mannikins, found in Africa, Asia, and Australia. The
Estrildinae never weave elaborate nests like the Ploceinae but con-
struct roughly globular nests of grasses and leaves, with the entrance
on one side, and which are usually built near or on the ground, in the
grass, oF in bushes and low trees. The nests are very large for the
size of their builders. A number of species frequently use old nests
of other weavers, but usually do a certain amount of work on the
866591—50-——21
312 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
nests themselves, such as adding to or rearranging the lining. Thus,
the bronze mannikin, Lonchura cucullatus, sometimes builds its own
nest, but often breeds in old nests of other weavers, particularly of
species of Ploceus. At Kaimosi, in western Kenya Colony, I found a
rather untidy, loosely constructed nest of dried grasses and plant
fibers, lined with grass seed-heads and feathers; it contained four white
eggs and was evidently the nest of a pair of these mannikins, which
were constantly seen on or about it. On the other hand, a day later I
saw a Lonchura enter an old Ploceus nest, and, wondering what the
bird might be doing there, I cut down the nest and found in it two
eggs exactly like those found in the other nest the day before. The
bird acted in a very excited manner as I examined the nest. Jackson
(p. 1473) also records that this species breeds in old nests of Ploceus
reichenowi, which it lines with grasses. Other Estrildinae known to
use old nests of other species not infrequently are the silverbill,
Lonchura cantans, the cut-throat finch, Amadina fasciata, the red-
headed finch, Amadina erythrocephala, the common waxbill, Estrilda
astrild, the zebra waxbill, Estrilda subflava, the lavender waxbill,
Estrilda perreini, and the cordon-bleu, Uraeginthus bengalus.
Aside from the fact that nest building in many of the Estrildinae
is not so fixed in its pattern but that the birds may either build new
nests for themselves or make use of old nests of other species (often
very different in design from those their own species would construct),
it is worth noting that in a good number of species the males take part
in the task of incubating the eggs. Thus, in writing of the zebra
waxbill, Jackson (p. 1517) goes so far as to say that “. . . as is gen-
erally the case with Waxbills, the males assist in incubation.” Infor-
mation on the courtship habits and sexual relations of the Estrildinae
is still rather scanty, at least as far as significant and reliably
worked-out details are concerned, but what data there are indicate
nothing unusual in either respect. The birds appear to be monog-
amous, and, as is so frequently the case with species in which the
sexes look alike, the courtship antics do not show any peculiar or
marked developments.
In review, then, the great family of weaverbirds exhibits an aston-
ishing range of diversity and variety in the mode of expression of the
different parts of the reproductive behavior cycle. In the beginning
of the breeding time we find everything from marked fragmentation
of wintering flocks into pairs, to year-round gregariousness, and in
courtship from a pattern that comes prior to nest building to one that
follows the completion of the nests by the males, from solitary antics
to elaborate display on special dancing grounds, and, on the other
hand, to almost none at all, or, as in the case of the house sparrow,
to barren but promiscuous displaying by mated males, seemingly
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 313
devoid of direct reproductive function, coupled with apparently
monagamous coition-inviting display by mated females. Nest build-
ing may vary from solitary to highly communal, and to none at all,
and even to parasitism, from slovenly put together masses of material
to amazingly fine and intricate weaving, or huge, communal super-
structures, or may be reduced to merely relining a disused nest of
another species or to lining a hole in a tree. Nest construction may
be done entirely by the male, by both sexes, or largely by the female,
or may be omitted entirely. Sexual relations vary from solitary
monogamy or social monogamy to polygamy, polyandry, and to ap-
parent promiscuity. Incubation in some species, or groups of species,
is performed solely by the hens, while in others the cocks share the
task with their mates, or, in the case of still others, neither sex takes
any care of the eggs, but are parasitic. 'The members of the subfam-
ilies Ploceinae and Viduinae do not come into breeding condition or
acquire adult breeding plumage until they are 2 years old; the mem-
bers of the other groups breed when 1 year old; this in itself is a pro-
found difference. In some forms of the Viduinae and Ploceinae it
permits a type of breeding-season gregariousness, although only a
single adult male and female are usually involved in each little flock.
Few, if any, families of birds offer such a bewildering array of
variations of the parts of the annual cycle, and I cannot help but
wonder if some of these variations may not have been due originally
to the extremes to which, in previously established variations, some of
the stages had been carried. At least the situations created by some
of these extreme developments seem to have been propitious for
further and even quite contrary subsequent changes.
Paradoxical as it sounds, it is possible that the excessive develop-
ment of the nest-building habit may actually have been a contributing
factor in the origin of the complete absence of nest building and egg
care that we know as brood parasitism. In cases of extreme indul-
gence in nest construction such as we find in the social weaverbird
(Philetairus socius) and some of its relatives (Plocepasser etc.), the
huge bulky structures are added to, chiefly by the males, all through
the nonbreeding season. By the time the birds are ready to make
their own individual nest tunnels in the already existing superstruc-
ture they are not acting very differently from birds that make use of
old nests of other species which they then repair. In the case of the
numerous species of typical weavers (Ploceinae) in which the not yet
breeding males construct many nests, the subsequently mated females
are again in a not dissimilar position of taking over nests which they
themselves have not built, and relining them and breeding in them. It
seems that Ali (1931) must have had some such thought in mind
when he noted that Baya weavers occasionally laid eggs in disused
314 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
nests of others of their own kind instead of making new ones for
themselves, as this prompted him to raise the following argument:
If the bird laid in disused nests it would only succeed in avoiding the labour
of building, but would still have to incubate the eggs itself. If on the other hand
it was successful in slipping into an unguarded Baya nest whence the brooding
hen had gone (as actually happened on September 18) and in laying its eggs there,
it would be, quite involuntarily, but with good effect all the same, compelled to
retire on the return of the legitimate occupant, leaving its egg to be hatched by
the Baya. Would such a process not tend, in course of time, to develope into,
and establish, a habit of systematic and voluntary parasitism as has been observed
in some African weavers?
In this connection it may be recalled that Lynes (1924, p. 661) found
that in nesting colonies of several species of African weavers related
to the Baya, studied by him in the Darfur Province of the Sudan,
many nests contained one or two extra eggs of the same species as the
host, but recognizably distinct by virtue of different color or state of
incubation, in other words, eggs that probably were laid by other
individuals of the same kind. It seems then, both in Asia and in
Africa, that not infrequently female weavers, ordinarily using nests
they have not built themselves, may lay an occasional egg in a nearby
nest of their own species.
The Viduinae are, as stated earlier in this paper, intermediate be-
tween the typical weavers (Ploceinae) and the waxbills (Hstrildinae).
In many species (perhaps the majority) of the former group, and also
in a good number of forms of the latter group, the hens breed in nests,
the actual construction of which has been foreign to their experience
and their efforts; in many forms of the latter group, and at least some
members of the former subfamily, the care of the eggs is taken over,
at least in part, by the cocks.
The parasitic mode of reproduction occurs, as far as known, in five
widely separated and quite unrelated families of birds—the ducks, the
cuckoos, the honey-guides, the weaverbirds, and the hang-nests (cow-
birds). There can be little doubt that the development of brood
parasitism has taken place independently in each of these five groups,
and it is not without significance, or at least suggestive value, that
this highly aberrant reproduction pattern has developed among the
small passerine birds (generally considered to be the most highly
evolved of all the birds) in those two families some of whose members
have carried the habit of nest building to its highest and most complex
development. It is all the more noteworthy that in the weaverbirds,
a larger group than the hang-nests and one with greater diversity of
behavior patterns, the parasitic habit has developed in two sub-
families, apparently {independently—the cuckoo finch, Anomalospiza
imberbis, in the Ploceinae, and in the members of the Viduinae, three
of which are definitely known to be parasitic, and the rest of which
BREEDING HABITS OF WEAVERBIRDS—FRIEDMANN 315
are strongly suspected of having the same habit. Many more details
have still to be learned of the annual cycle of behavior patterns in
these birds before it may be possible to attempt to determine the
precise causes and the subsequent evolutionary paths that twice in
the history of the weaverbirds have lead from nesting and incubation
and caring for the young to a state of brood parasitism.
REFERENCES
Aut, Sauim A.
1931. The nesting habits of the Baya (Ploceus philippinus). Journ. Bombay
Nat. Hist. Soc., vol. 34, No. 4, pp. 947-964.
Bates, Grorce L.
1930. Handbook of the birds of West Africa.
BELCHER, CHARLES F,
1930. The birds of Nyasaland.
CuaPiNn, JAMES P.
1917. The classification of the weaver birds. Bull. Amer. Mus. Nat. Hist.,
vol. 37, art. 9, pp. 243-280.
DELACOUR, JEAN.
1943. A revision of the subfamily Estrildinae of the family Ploceidae,
Zoologica, vol. 28, pt. 2, pp. 69-86.
Deacour, JEAN, and Epmunp-Buanc, F.
1933-1934. Monographie des veuves. Oiseau, n. s., vol. 3, pp. 687-726; vol.
4, pp. 52-110.
FRIEDMANN, HERBERT.
1922. The weaving of the red-billed weaver bird in captivity. Zoologica,
vol. 11, No. 16, pp. 355-372.
1929. The cowbirds. A study in the biology of social parasitism.
1930. The sociable weaver bird of South Africa. Nat. Hist., vol. 30, No. 2,
pp. 205-212.
1935. Bird societies. Jn A Handbook of Social Psychology, chap. 5 (edited
by Carl Murchison).
JACKSON, FREDERICK J.
1938. The birds of Kenya Colony and the Uganda Protectorate. 3 vols.
Jourpain, F. C. R., and Tucknr, B. W.
1938. In Witherby, Jourdain, Ticehurst, and Tucker, Handbook of British
Birds, vol. 1, pp. 157-158.
Lack, Davin.
1935. Territory and polygamy in a bishop-bird, Euplectes hordacea hordacea
(Linn.). Ibis, ser. 18, vol. 5, pp. 817-836.
Lynes, HuBERT.,
1924. The birds of north and central Darfur. Ibis, ser. 11, vol. 6, pp. 661-
678.
Prisst, Cecit D.
1933-36. The birds of Southern Rhodesia. 4 vols.
Roserts, AUSTIN.
1917. Parasitism amongst finches. Ann. Transvaal Mus., vol. 5, No. 4,
pp. 259-262.
1939. Notes on the eggs of parasitic birds in South Africa. Ostrich, vol. 10,
pp. 1-20, 100-117.
1940. The birds of South Africa.
316 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
SHELLEY, G. E.
1905. The birds of Africa, vol. 4, pt. 1, pl. 31 (opposite p. 108).
Srark, ARTHUR C.
1900. The birds of South Africa, vol. 1.
TAYLOR, J. SNEYD.
1946. Notes on the masked weaver. Ostrich, vol. 17, No. 3, pp. 145-155,
Van Someren, V. D.
1945. The dancing display and courtship of Jackson’s whydah. Journ.
East Africa Nat. Hist. Soc., vol. 18, Nos. 3-4, pp. 1381-141.
Van SomEREN, V. G. L.
1916. List of birds collected in Uganda and British East Africa, with notes
on their nesting and other habits. Ibis, ser. 10, vol. 4, pp. 373-472.
NEW ZEALAND, A BOTANIST’S PARADISE
By Eaprert H. WALKER
Associate Curator, Department of Botany, U. S. National Museum
[With 10 plates]
INTRODUCTION
The wisest traveler learns as much as possible before a trip, sees
all he can during his journey, and corrects and enlarges his knowledge
by further reading and inquiry after returning home. This article is
the outgrowth of the author’s short but full visit in New Zealand,
which was ideal in nearly every respect except for lack of advance
knowledge of the country, especially of its botany. It suggests what
the writer would have liked to know in advance but had to learn on
the trip and after it. The suggestions given here for further reading
may be of interest, not only to the fortunate few who will visit New
Zealand in person, but the greater number who may do so vicariously
by reading and by listening to those who have gone.
The plants of New Zealand can hardly fail to gain the attention of
the visitor, and the student of New Zealand will find abundant refer-
ence to them in his reading. People in an industrial country may
ignore the plant life, but those in an agricultural land like New
Zealand cannot escape the imprint of the vegetation on their
lives. New Zealand is a land of enthusiastic and competent amateur
naturalists, and its professional botanists are outstanding. The visitor
will find a local naturalist in nearly every town or center, who is eager
to share his specialties with the stranger and to show him the offerings
of the field. ‘The traveler to New Zealand will probably first meet
the introduced flora which dominates the landscape in the inhabited
parts. Only when he visits the more remote and undisturbed areas
will he see many of the native plants. If he has an economic or
agricultural bias, probably the grasslands, the backbone of New
Zealand’s economy, will impress him most. (See pl. 9.) If he is
conservation-minded, the sight of the vast area of shrubland and
fernland (Pterrdium aquilinum var. esculentum) will make him pain-
fully aware of man’s destruction of the native forest. But the soul
of the pure botanist, undisturbed by problems of economics and con-
317
318 | ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
servation, will be stirred most as he enters the forest or “bush” where
abounds the native New Zealand flora, so rich in plants found in no
other country—that is, the endemic species. (See pls. 2 and 3.)
Besides the grassland, scrub, and bush he will see other plant forma-
tions. The extensive plantations of trees, all planted in rows of uni-
form age, are very impressive. The species so grown are all exotics,
that is, not native to New Zealand, the principal one being the Monte-
rey pine (Pinus radiata), a useless tree in its native California, but
here by far the most economically important tree to be found. Not
only does it occur in plantations, but it is to be found almost every-
where as a hedge tree or windbreak (pl. 9, fig. 1) and even as a
naturally planted weed invading wasteland.
It is essential in understanding the peculiarities of the flora of New
Zealand to know its location and climate as contrasted with that of
more familiar areas. The vegetation or the major plant formations
will then be discussed, after which the flora or the elements which
compose the vegetation will be taken up. Finally some consideration
will be given to the past and present study of botany in the country.
LOCATION AND CLIMATE
New Zealand extends from about the 34th to the 47th parallel south
latitude, a distance of about 900 miles. (Fig. 1, and fig. 2, p. 333.)
It consists essentially of three islands, North, South, and Stewart
Islands, with a few small nearby islands or islets. There are several
outlying island groups, politically and biologically part of New
Zealand, of much interest, but they are not included in this discussion.
For vivid geographical comparison, suppose the three main islands of
New Zealand were inverted and superimposed on North America at a
corresponding latitude, with the North Cape of North Island at Cape
Lookout about the center of the Atlantic coast of North Carolina.
Then the South Cape on Stewart Island would lie north of Quebec in
Canada. The East Cape at the end of the Grisborne Peninsula of
North Island would be in southern West Virginia, and Mount Egmont
at Harpers Ferry, W. Va. Wellington, at the southern end of North
Island, would be in west-central Pennsylvania, and Christchurch on
the east side of South Island would be on the shore at the east end of
Lake Ontario. Both areas are in temperate zones of the earth’s
surface, but their climates are in striking contrast. Eastern North
America has a continental climate with extremes of temperature and
a moderate, irregular precipitation. New Zealand, however, has a
strong oceanic climate with far milder temperatures throughout and
a much smaller difference in temperature between the northern and
southern ends and between winter and summer. The precipitation is
fairly evenly distributed throughout the year, although it varies
from place to place. New Zealand owes its climate to the unifying
NEW ZEALAND—EGBERT H. WALKER 319
LS
SO arr
—
3 <5 --t-Fe-
apricorrg "| 7
b; Db
- a
“7
N\ \ SS
> =Chatharn &
aN Na
SER S|!
Tee |
SS
1Bocirity ves
IES —
Anti vOWeSN|
eee aN
°° Acwkleind [5.22
: wot a
ace ! 4 |
sas PH eg bos y —-
7GILUG 2
oF) A |
Du lacguarrie ss
!
Se eee
|
eZrneraldl [4
Ficure 1—Chart of Tasman Sea showing deflection of the cold Antarctic drift
by the warm East Australian current from the Tropics. (From W. C. Davies
(12).)
influence of the warm Tasman Sea on the prevailing westerly winds
which blow over it from Australia, over a thousand miles away. In
its oceanic climate lies the explanation of many of its vegetational
contrasts with other countries.
The vegetation in northern New Zealand is far more tropical in
appearance than its geographical counterpart in North Carolina.
320 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
This ‘subtropical’ aspect of the forests can be seen even in northern
South Island and grades into the characteristic features of the dense
temperate rain forests of the west coast of South Island. The south-
ern-beech forests of Nothofagus in the south are clearly Temperate
Zone forests. (See pl. 1, fig. 2.)
The mountains in New Zealand cause more changes in the climate
from place to place than does the latitude. The mountain ranges
and plateaus of North Island le mostly east of the center. They
are largely volcanic and influence the vegetation not only through
their effect on the winds and moisture but also on the soil. The
volcanic ash and pumice readily absorb more moisture, much of which
seeps away beyond the reach of the plants growing on the surface.
Mount Egmont is a majestic isolated volcanic cone on the west coast
with vegetation in characteristic altitudinal zones from sea level to
the perpetually snow-covered summit.
In South Island the rugged Southern Alps parallel the west coast
and thrust their peaks far into the zone of permanent snow. (See
pl. 7, fig. 1.) Their highest peak is Mount Cook, its summit 12,349
feet above the Tasman seashore less than 24 miles to the west. They
are formed by erosion of uplifted land rather than volcanic activity
and are composed largely of friable greywacke rock. (See pl. 7, fig. 2.)
These ranges drain the prevailing westerly winds of most of their
moisture. Thus the west coast has a heavy rainfall of around 200
inches a year, while on the plains of the eastern leeward side there
may be as little as 20 inches. The highest annual rainfall yet re-
corded is 228 inches at Puysegur Point on the west coast, and the
lowest, 13 inches, in Central Otago only 150 miles away. Dense rain
forests cover the steep western slopes and the narrow coastal plain
below, whereas on the east are the natural grasslands and the broad
cultivated plains. In Central Otago is a semidesert area. The
transition fromheavy rain and dense forest to sunshine and almost
barren eroding slopes may be made in a surprisingly short time in
driving over the divide. Other ranges and hills, especially the Kaikora
Ranges in Marlboro and those in the rugged Banks Peninsula, inter-
rupt this picture and diversify the ecological conditions and vegeta-
tion. Thus New Zealand has a great variety of distinctive, plant
formations in a remarkably small space, a fact which makes bota-
nizing a most interesting and relatively easy occupation. The climate
of New Zealand has been presented by Kidson (16, 17).!
THE GRASSLANDS
The grasslands of New Zealand are the foundation of its agricul-
tural economy, and one is sure to be impressed by their extent and
1 Numbers in parentheses refer to the bibliography.
NEW ZEALAND—EGBERT H. WALKER 321
.
variation. They are of two kinds: first, the original tussock grass-
lands of native species (pl. 9, fig. 2), and second, the pastures with a
sward formed of introduced species (pl. 9, fig. 1). The history of
New Zealand is largely the story of man’s replacement of the native
bush with pastures and the exploitation of the native grasslands in
feeding his flocks of sheep. In the ashes of the bush he planted grasses
and began the process of adapting the sward-forming techniques, so
well developed in his native England, to the conditions of this new and
promising land. He fought a continual battle with the few native
plants with weedlike tendencies and the more numerous and generally
more vigorous exotics, which he intentionally or unintentionally
brought from the far parts of the earth. From early blunderings he
has now developed the technique of sward growing to a very advanced
degree, and agricultural progress or deterioration in large parts of
North Island and certain regions of South Island depends on the com-
position and condition of this pasture sward. Much land formerly
covered with fern, scrub, or blackberry is now grass-covered, with a
high sheep-carrying capacity. It was a most enlightening experience
to see the work of the Animal Research Station at Ruakura near
Hamilton in Auckland Province, in breeding and mixing strains of
grasses, on which, with proper rotation, an amazing number of sheep
can graze throughout the year without additional feed. The various
types of these artificial grasslands of North Island have been carefully
mapped and analyzed in a publication by Madden (19). Something
of the history and significance of these grasslands can be gleaned from
the account written by an English‘agronomist, Stapledon, who visited
Australia and New Zealand in 1926 (25). The planted pastures of
South Island, developed by essentially the same means, are well
described in Hilgendorf’s ecological survey of the grasslands (13),
which supplements Madden’s.
The natural grasslands are very different from the man-made
pastures. They occur most extensively in South Island, but smaller
areas are to be found in North Island, especially in the central plateau.
A relatively low rainfall with a cooler and more even temperature are
among the principal factors governing the development of grasslands
rather than shrublands or “bush.’”’ The plant composition of this
formation varies considerably according to local conditions. Pas-
toralists generally recognize five tussock grasses. ‘Two of them, snow
grass (Danthonia raoulii var. flavescens) ? and red tussock (D. raoulu
var. rubra) are the tall tussocks, 3 to 6 feet high, while the short
tussocks, 1 to 2 feet high, are the silver tussock (Poa caespitosa) and
the hard or fescue tussock (Festuca novae-zealandiae). The blue
tussock (Poa colensoi) is only 6 to 9 inches high.
2 The names, both common and scientific, in general use in discussing these grasses vary considerably.
This makes research on the grasslands rather difficult. Zotov (28) recognizes only four tussock grasses.
322 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The history of the tussock lands is an almost continuous story of
progressive deterioration due to overgrazing, burning, rabbit infesta-
tion, and increased wind erosion, land slip, soil creep, and water
erosion. The present deserts in northern Canterbury, Marlboro,
and Central Otago Provinces were grasslands when white settlement
began, and the carrying capacity of most sheep runs is today far less
than it was in the beginning. Everywhere one can see erosion that
is of recent origin. A most definite sign of overgrazing in South
Island is the excessive development of the scabweed, Raoulia lutescens.
(See pl. 8, fig. 2.) Usually the rabbit population increases as the
tussocks diminish and more open spaces are formed. This only
accelerates the destructive process. It is important to keep in mind
that the tussock grasses themselves are rarely grazed, except the new
growth which springs up after they are burned over, which is tradi-
tionally done annually. The role of the tussocks is to furnish pro-
tection to the smaller grasses and other plants which grow among
them and furnish most of the feed.
The tussock grasslands of South Island have been dealt with rather
fully by Zotov (28) and recommendations presented for restoring these
areas to production. First, annual burning must be eliminated or, if
absolutely necessary to eliminate shrubby invaders, replaced with
carefully controlled burning. The number of grazing animals must
be reduced to the carrying capacity of the land and rotational grazing
introduced in order to restore the fertility. When necessary, the tus-
sock grass must be replanted with selected unpalatable strains or jor-
danons, and, when a protective covering is thus established, highly
palatable strains of native species must be sown between the tussocks.
Hardly any of these measures are now used by the sheepmen. It was
most gratifying to have a glimpse of the Government’s research work in
tussock-grass restoration at its field station in Hutt Valley near
Wellington. The work done there is preliminary to research and
experimentation in the tussock country itself and will surely some
day result in restoration of much depleted land. Probably some
areas have gone almost beyond reclamation and will remain, as have
so many other parts of the world, monuments to man’s lack of fore-
sight and self-control in seizing all the produce of the land rather than
just its surplus.
SHRUBLAND AND FERNLAND
No traveler in New Zealand can fail to be impressed, and at the
same time generally depressed, by the vast extent of land covered by
shrubs and ferns. Unlike the grasslands, they bring no sense of well-
being to man, and, compared with the forest, they at first seem
botanically unattractive. But neither impression is wholly correct.
NEW ZEALAND—EGBERT H. WALKER 323
Shrubland is any plant community in which tall trees are wanting
and shrubs dominate. The fern, which comprises the fernland, is the
native variant of the world-wide bracken, Pteridium aquilinum var.
esculentum, or P. esculentum of many authors. Fernland is here
linked with shrubland because the fern reaches shrub size and the
formation is as dense and impenetrable as the densest thicket of
woody shrubs. Furthermore, this fern community is closely related
ecologically to the other most extensive shrub community, that
dominated by manuka? (Leptospermum scoparium or L. ericoides—
Myrtaceae) (pl. 5, fig. 2). Together these two cover more area than
do the other shrub formations, of which there are many in very
different habitats and of diverse composition, form, and origin. As
in other lands, these shrub formations develop in response to certain
natural conditions. These conditions may develop over a long
period of time. When sudden changes occur favorable to the growth
of a shrub community the formation is called an induced formation.
Shrub formations follow certain volcanic eruptions and sometimes
floods or places of excessive erosion. But more significantly they are
man-induced, coming along after the forest has been destroyed with
ax and fire, and grass has been sown on the ashes, or where man’s fires
and his overabundant greedy sheep have destroyed the natural
grass cover. (See pl. 4, fig. 1.)
The manuka (Leptospermum scoparium—Myrtaceae) is a shrub or
small tree with an amazing adaptability and persistence. It usually
forms a community without the bracken or it may be variously mixed
with this fern. It seems able to grow anywhere, wet or dry, in good
soil or bad, and in heat or reasonable cold, but not in alpine con-
ditions or deep forest shade. Its outstanding ability to thrive on
poor soil makes it rush in where man has done his best to destroy the
land. It is especially prominent on the gumlands of North Auckland,
dug over and the fertility dissipated in the search for fossil kauri gum,
desired as an ingredient in high-grade varnish. Manuka is extremely
plastic in its response to its environment. Within this community
there are some 81 other species of plants, many of them of great
interest to the botanist, not the least being the bulbous-rooted New
Zealand orchids. Manuka is an important source of fuel for man,
3 To the foreigner the widespread use in New Zealand of native Maori names for trees and other plants is
somewhat disconcerting. Very often there is no other name, as for example, manuka for Leptospermum
scoparium or kauri for Agathis australis. These names have often been made into the specific scientific
names, as taraire in Beilschmeidia taraire and tawa in B. tawa. Another common practice of many New
Zealanders is to use in speech a specific scientific name for a common name, as macrocarpa for Cupressus
macrocarpa, lawsoniana for Cupressus lawsoniana, and radiata for Pinus radiata, the Monterey pine.
Another disconcerting practice is the use of some common English name, such as pine, for something which
a foreigner at least would hardly recognize as such. Native pine in New Zealand refers to species of Podo-
carpus. ‘Thus the traveler is in a new nomenclatorial atmosphere. For popular names of New Zealand
plants see Andersen (2) and Cheeseman (4).
324 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
and in its shade grow seedlings of many forest trees, which eventually
rise up and wipe out this ‘‘nurse” species by overshading it. So the
manuka shrubland fills a varied and not altogether harmful place in
New Zealand’s plant economy.
One cannot feel quite so resigned to the fernland, though one
quickly does resign from the job of trying to penetrate it. Its adapta-
bility and prolificness closely matches the manuka, but within its
dense growth there are few if any other plants. Like the manuka, it
cannot endure much shade or cold, and so is not a denizen of the
forest and alpine or subalpine slopes. Man may burn its tangled
fronds, but new ones rise quickly from the unharmed underground
stems. However, overstocking with cattle which eat the tender
young fronds catches the bracken in its “tendon of Achilles,” and if
the practice is persisted in, this scourge can eventually be conquered.
A third shrubland community of much prominence is not only the
result of man-made conditions, but of man’s introduction of plants.
The English gorse, Ulex europaeus, was first brought, no doubt, to
relieve man’s nostalgic longing for the lovely English countryside and
to lend color to the generally colorless New Zealand vegetation. But
this was an imprudent act. From the hedgerows it spread easily to
adjacent fields, dry, gravelly river beds, formerly forested hillsides, and
pastures. It is quite indifferent to the quality of the soil. Large open
spaces soon became impenetrable thickets. Man constantly burns it
off, but fire seems only to improve the viability of its seeds and to
impoverish the soil, which harms the accompanying fodder plants
more than it does the gorse. Control over large areas by grubbing it
out of the ground is hopeless in this land of limited labor and large
demands on human resources. Handcuffed with this gorgeous yellow
culprit are the broom (Cystisus scoparius—Leguminosae), the rose, and
the blackberry, and several adventive shrubs from adjacent Australia,
especially hakea (Hakea acicularis—Proteaceae). One American
shrub of this category is the tree-lupine of California, Lupinus arboreus,
brought as a sand binder and now spreading beyond its first plantings
to other sandy and gravelly spots, not always according to man’s
wishes.
The term “scrub” is often applied to any shrub formation. In
Australia it is erroneously applied to certain forest formations, but
Cockayne (7) applies the term in a more restrictive sense to any
community of divaricating, stiff, shaggy, and often spiny shrubs.
Such shrubs have numerous extremely wiry or rigid, much interlaced
branches and twigs which zigzag at a wide angle in every direction.
The thickets they form are close, unyielding, and often cushionlike
masses. ‘To push ones way through this scrub is impossible and to
travel over it is often a hazardous undertaking. The scrub is usually
subalpine and is composed of various species of Coprosma (Rubiaceae),
NEW ZEALAND—EGBERT H, WALKER 325
Cassinia and tree-daisy (Olearia—Compositae), wild Irishman (Dis-
caria toumatou—Rhamnaceae—pl. 6, fig. 1), and Myrtus (Myrtaceae),
though altogether there are about 55 species in 18 families which have
this divaricating habit.
The plant collector who makes pressed specimens almost meets his
Waterloo when he tries to make a herbarium specimen to represent
adequately such a plant. Some divaricating shrubs further thwart
the collector by dropping their leaves, flowers, and fruits almost at
the first gentle touch. The divaricating branches of pohuehue
(Muehlenbeckia astoni—Polygonaceae) are pliant enough, but when a
representative specimen has been warped into a plant press there is
rarely a leaf or fruit left in situ, and a vivid supplementary description
is needed to bring to the observer’s mind any adequate concept of the
original habit of the plant.
One will find various shrub formations in a wide range of habitats.
Besides the extensive hillside formations of manuka and fern, and the
subalpine scrub, this type of vegetation is often found on sea coasts
(both rocky and sandy), wet lands, mineral lands, areas of volcanic
ash and pumice, and wind-swept shores and mountain slopes. Its
component species, growth forms, and adaptations to environmental
conditions are of much interest. In some places associations of trees,
dwarfed to shrub size by wind, salt spray, or soil influences, resemble
and merge into shrub formations.
FORESTS
The principal natural resource in New Zealand when the pakeha or
white man first came was its trees; at the present time it is its grass.
But the white man could live only secondarily on the forest, so the
trees had to go in order that he might provide for his primary need—
food. Hence, this natural resource, which formerly covered almost
the whole of North Island and much of South Island, was sacrificed
at a rate hardly equaled anywhere else in the world. It took Europe
four centuries to exploit its forests and America two centuries, but
New Zealand accomplished this in one century. Although the exten-
sive natural forests of the past are gone, the remaining fragments are
sufficient to tell us a great deal about New Zealand’s botanical history.
According to Cockayne (7), 385 species of plants are characteristic of
the forests. Of these, 99 are trees, 63 shrubs, 51 herbs, 26 grasslike
plants, 88 ferns, 26 climbing plants, 15 epiphytes, and 13 parasites.
Ninety percent are endemics. So it is in the New Zealand forest that
a visiting botanist will find his greatest delight in becoming acquainted
with the true New Zealand flora. If he arrives in North Island, as is
most likely (and much regretted by the people of South Island), his
first acquaintance will almost surely be with the rain forest, which is
composed of many species of trees and shrubs of many genera and
326 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
families. (See pl. 3, fig. 1.) Later he will meet the strikingly dif-
ferent southern-beech forest, composed almost entirely of one or two
species of Nothofagus and few shrubs. (See pl. 1, fig. 2.) There, if he
is from the Temperate Zone of North America or Europe, the traveler
will feel much more at home, although all the plants will be new to him.
The rain forest in New Zealand is clearly tropical in its origin and
affinities; indeed, it is often designated as subtropical rain forest,
although it lies entirely in the Temperate Zone. Its character is the
result of the oceanic climate with its mild, rather uniform temperatures,
and its abundant, evenly distributed rainfall, which assures a high
atmospheric humidity throughout the year. The southern-beech
forest of Nothofagus, however, is clearly temperate in origin, with its
nearest affinity the Nothofagus forests in Patagonia, Tierra del Fuego,
and southern Chile, on one side, and in Tasmania and adjacent
Australia, on the other. This forest occurs in New Zealand in cooler
and less humid regions than most of the rain forests. It depends
more on ground water and thrives in a less humid atmosphere. All
the forests in New Zealand are evergreen. There are a few deciduous
trees but none of them are forest dominants, so there are no deciduous
forests in New Zealand.
The rain forest is complex, the beech forest relatively simple. These
two forests may be contrasted in part, as follows:
Rain forest Southern-beech forest
Tropical in appearance. Temperate in appearance.
Composed of many tree species in many Composed of one or two tree species in
genera. one genus.
Dense within. Open within.
With several plant strata.‘ With only an open layer of shrubs be-
tween canopy and ground.
Bases of trunks often with plank but- Trunks without buttresses.
tresses.
With many vines or lianas. With very few vines.
Loaded with epiphytes. With only a few parasites.
With many ferns. With few ferns.
Each forest formation has various forest associations within it, these
being quite complex in the rain forests and relatively simple in the
southern-beech forest. The forest associations are named according
to the dominant species within them, the principal associations in the
rain forests being: (1) kauri (Agathis australis—Pinaceae), (2) mixed
dicotylous-taxad,®? and (3) “white pine” or kahikatea (Podocarpus
dacrydiovdes).
4 “Forest is piled upon forest.””—Humboldt.
5 Cockayne’s term for this forest association varies. It is called a mixed taxad forest (6) and dicotylous-
podocarp forest (7). Species of New Zealand “pine,’’ Podocarpus or Dacrydium, members of the yew family
Taxaceae, are dominant or characteristic. There are also many trees of various families of dicotyledons,
whose seeds have two cotyledons in contrast with the conifers which have several and the monocotyledons
with only one cotyledon. The use of the term dicotylous rather than dicotyledonous is in conformity with
Cockayne’s usage (7).
Smithsonian Report, 1949.—Egbert H. Walker PLATE 1
]
1. LAKE FERGUS NEAR THE HOMER TUNNEL, WESTERN OTAGO, SOUTH ISLAND
Southern-beech (Nothofagus) forests clothe the mountains high up toward the snowy peaks of the Southern
Alps.
2. SOUTHERN-BEECH FOREST (NOTHOFAGUS FUSCA), PARADISE, LAKE WAKATIPU,
OTAGO PROVINCE, SOUTH ISLAND
Without looking at the foliage one would think himself to be in a northern-beech (Fagus grandifolia) forest
in temperate North America.
Cseraed O° M Aq ydeiz0j04q) ‘a[sunf[ esuap siqq 9z119}0BIBY CS|R
soyAydide juespunge pue (jJa[) snuepUed SUIGUII[D 9q,J, *901} ISaI0} (‘UOSTON ‘AIN{YSUT U9IQYMBD Asoeqin>+o ‘seraeqd “OM AQ
esny 8 8q 0] SMOIS puB “4SOY S}I Se[suRIys pus SdojaAus pue ‘punois qdeis0j0yd) 480} LOT UM0ID 94 JO pwaids 9Y} PUR J9eJ G9T SI IYSIaY [eI0}
94} 0} UMOP sayover ‘ayAydide ue sv ast SuIseq Al[[ensn BIBI BYAL aL ‘SeyouBsq IsIY ay} 0} YSiY JoaJ QO PUB JOJBUIBIP UL Joa] 44 fT SL 80 Uy,
LSSYO4 AVXVL-SNOTALOSIC GNV1IS| HLYON ‘GNVY1TIHONY HLYON ‘LSSYHOS
GaXIW SHLNI (VLSNEOY SONSCISOYLAW) VLVY V “Zz S3LVLSVNOdIVM AHLNISSYLIYNVy Vv... ALNHYWANVL,, ‘1
P SINAN
J9¥]2A\ H 42934 —6h6| “HOdey weruosyztUIg
ae
¢ 3LV1d
Cprey sf 4q ydeisojoyg) * (44811)
HOOT [BIIOA B SULIAAOD (Digns DYNnODY) Caeys as[qejeseA Useds
pues -(punoise10} pue Jeyued) “ds pisniydng ‘(4Jap) s271qvzoeds visiw7ag
GNV1S| HLYON
“AONIAOYd NOLONITISM ‘SONVY WOVE _L
SINV1d HOOY NIVLNNOW SAO ASTGAW V °Z
€ ALV1d
(sarang "DO °M Aq ydeis
-0JOUq) *4S910j YOIISIYI JO BIBIYS [RIBAS BY BSIIdUICD SIIqBY asIaATp JO
se1oeds gel] Auvu pue ‘soyAydide ‘seuely ‘sued neyiu 04} ‘SLLI8} BOL,
1SSHYO4 AVXVL
“SNOTALODSIG GSXIW AHL NI HLMOYUDS SSN3Q AHL ‘ff
JONIeM “LI 1100G39—*4b¢4)] ‘qu0dayy UPIUOsSU]IWIC
Smithsonian Report, 1949.—Egbert H. Walker PLATE 4
1. EDGE OF WAIPOUA STATE FOREST, NORTH AUCKLAND, NORTH ISLAND
The untouched primeval kauri forest (right) formerly extended over the now cut-over stump, bracken,
grass, and exotic shrub covered sheep-grazing land (left). (Photograph by E. H. Walker.)
2. COASTAL DYSOXYLUM SPECTABILE FOREST, STEPHENS ISLAND, COOK STRAIT
The forest floor is free of undergrowth, probably partially damaged by cattle. The Dysorylum has
characteristically twisted trunks and above-ground roots. The slender trunks are Piper eacelswm.
(Photograph by L. Cockayne, courtesy New York Botanical Garden.)
Smithsonian Report, 1949.—Egbert H. Walker PLATE 5
1. TREE FERN, SHORE OF LAKE ROTO-ITI, NORTH ISLAND
Large tree ferns of many kinds are denizens of the forests and cut-over lands, and often of the roadsides
i je toe Bye ae ees os 0 eer i 7 Pe
2. MANUKA (LEPTOSPERMUM SCOPARIUM), EAST COAST SOUTH ISLAND
This abundant association clothes vast open lands from sea coast and swamps to mountain sides with an
almost impenetrable thicket. (Photograph by L. Cockayne, courtesy New York Botanical Garden.)
Smithsonian Report, 1949.—Egbert H. Walker PLATE 6
1. WILD IRISHMAN (DISCARIA TOUMATOU), A DIVARICATING SHRUB, DRY EAST
COAST OF SOUTH ISLAND
This characteristic growth habit is found in many New Zealand shrubs. See also plate 7, figure 1.
(Photograph by L. Cockayne, courtesy New York Botanical Garden.)
2. COLLECTING DONATIA NOVAE-ZELANDIAE IN AN ALPINE BOG, MAUNGATUA
RANGE, OTAGO PROVINCE, SOUTH ISLAND
This dense flat mat is so solid that footprints hardly show. Other species of this genus occur only in
Tasmania and southern South America. (Photograph by E. H. Walker.)
Smithsonian Report, 1949.—Egbert H. Walker PLATE 7
1. HOOKER VALLEY BELOW THE HERMITAGE, EAST SIDE OF SOUTHERN ALPS,
CANTERBURY PROVINCE, SOUTH ISLAND
A debris-choked glacial valley with braided streams thick with glacial grindings is gradually invaded by
wild Irishman shrubs (Discaria towmatou) and alluvial fans of weathered rock. (Photograph by E. G.
Holt, U. S. Soil Conservation Service.)
CS '¢ r -
£ aS >”
SOUTH ISLAND
Rain clouds, rising against the range, drench the rain forests on the lower slopes and cover the higher
peaks with deep snow, whence flow the glaciers. (Photograph by E. G. Holt, U. 8. Soil Conservation
Service.)
Smithsonian Report, 1949.—Egbert H. Walker PLATE 8
1. EROSION AND ITS CAUSE NEAR WAIHO DOWNS, SOUTH OF TIMARU, CANTERBURY
PROVINCE, SOUTH ISLAND
When sheep overgraze the grass and cut the turf, erosion occurs. Fencing out the sheep (left) allows the
vegetation to help control erosion. (Photograph by E. G. Holt, U.S. Soil Conservation Service.)
2. AN OVERGRAZED HILL IN TUSSOCK-GRASS COUNTRY, LINDA PASS, OTAGO
PROVINCE, SOUTH ISLAND
Soil, bared by overgrazing between the tussocks, is covered with pale green scabweed (Raoulia lutescens)
which in turn nurses tussock-grass seedlings. (Photograph by E. H. Walker.)
Smithsonian Report, 1949.—Egbert H. Walker PLATE 9
reat
Ee
1. PASTURE LAND, HERETAUNGA PLAIN, HAWKES BAY, NORTH ISLAND
Rich pastures of introduced grasses are often separated by hedges and windbreaks of lombardy poplars
and Monterey pine or cypress. Turnips or swedes are grown for winter feed. (Photograph by E. G.
Holt, U.S. Soil Conservation Service.)
2. BREAST HILL STATION IN THE LOW TUSSOCK LAND OF CANTERBURY PROVINCE,
SOUTH !JSLAND
Sheep stations (ranches) are protected by windbreaks of planted pine. The roadside vegetation is little
grazed and more luxuriant. (Photograph by E. G. Holt, U.S. Soil Conservation Service.)
Smithsonian Report, 1949.—Egbert H. Walker PEATE LO
CANTERBURY PROVINCE, SOUTH ISLAND
”
The ‘‘pimples’”’ on the near end are flowers. This weird plant is characteristic of open shingle slopes in
the dry area. (Photograph by L. Cockayne, courtesy New York Botanical Garden.)
a
ac 7 E ee mes - ea i
2. BULL KELP (DURVILLEA ANTARCTICA) ON ROCKS AT LOW TIDE, DOG ISLAND,
FOVEAUX STRAIT
This flat-bladed leathery kelp grows abundantly on wave-lashed recky shores. (Photograph by L.
Cockayne, courtesy New York Botanical Garden.)
NEW ZEALAND—EGBERT H. WALKER S20
The kauri forest is the best known of the rain-forest associations,
this being due to the noble as well as the highly commercial attributes
of its prominent component, the kauri tree itself. (See pl. 2, fig. 1.)
Originally almost all of North Island north of the 38th parallel, which
crosses the island about the base of the Bay of Plenty, was a vast
kauri forest. The title of a recent booklet, “The Waipoua Forest:
the Last Virgin Kauri Forest of New Zealand” (21), shows its present
state. The Waipoua State Forest or the Waipoua Forest Reserve in
North Auckland contains about 40,000 acres, of which about 27,600
acres is actually forested. Perhaps the Trounson Kauri Park with
only about 975 acres is too small to consider, but nevertheless it con-
tains a primeval kauri stand of limited extent, donated by its lumber-
man namesake to the State for a preserve. Because of its small size
and the natural degeneration from its exposed margin, its longevity
as a primeval forest may be limited. The kauri forest formation is
composed of many other tree species besides Agathis australis, as well
as characteristic shrubs and ferns, including huge tree ferns, and a few
herbaceous plants. Among the many climbers is the kie-kie or climb-
ing pandanus (Freycinetia banksit) (pl. 2, fig. 2), the only New Zealand
member of this tropical family, Pandanaceae. This forest varies
according to the presence or absence of the actually dominant trees,
especially of Beilschmeidia taraire in the northern and B. tawa in the
southern part of its range. Various studies have been made of this
forest from different points of view, but probably Cockayne’s (5) and
McGregor’s (21) are of greatest interest to the botanist.
Impermanence seems to be woven into the fabric as well as the
history of the kauri forest. Cockayne (6) has stated that the kauri
is always in a state of progression or retrogression. Seedling kauris
cannot normally reach maturity within the kauri forest itself, but
must grow up where there is more light, for this forest is predomi-
nantly dark, gloomy, dense, and almost impenetrable. Seedlings
grow well, however, in the shade of the manuka or in accidental
clearings and along roadsides cut through the forest. By some it is
thought that the kauri forest will in time cease to exist unless man
takes a hand to perpetuate it. But the Maori people first came only
800 years ago and the white man less than 150, and there were kauri
forests many centuries earlier. There is in New Zealand now a lively
controversy over the preservation of the Waipoua State Forest.
Shall it remain as it now stands, or be reduced in size? Shall it be
left wholly untouched, or is it to be altered within by forest manage-
ment? Shall it furnish timber to commercial exploiters or forest
managers, or be a recreation ground and memorial of the past to the
6 Actually the kauri tree is not dominant, as it grows singly or in clusters. Kauri forests are so named
because of the prominence and commercial value of the kauri, The broad-leaved dicotylous species are
actually dominant.
866591—50——22
328 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
public, or an untouched natural research laboratory to the scientist?
Able protagonists have arisen to fight for its preservation from com-
mercial exploitation (21).
The many complexities and variations in the mixed dicotylous-
taxad forest can only be suggested here. Such a forest may be a rimu
forest with the ‘‘red pine” (Dacrydium cupressimum) dominant, a
totara forest with Podocarpus totara dominant, or a tawa forest with
the principal trees Beilschmeidia taraire or B. tawa of the dicotyledon-
ous family Lauraceae, though usually with Podocarpus or Dacrydium
also present. These forest types intergrade and vary extensively, so
it is often difficult to determine just what kind of tropical forest one
finds himself in.
The parallels of 38° and 42° south latitude are significant as the
southern limits of quite a number of species and the northern limits
of others. As noted above, the kauri ceases at about 38° south.
Some of the 100 or so others which drop out at this point are the toru
(Persoonia toru—Proteaceae), the white-flowered tawari (Izerba breri-
oides—Saxifragaceae), the climbing fern (Lygodium articulatum), and
the trailing fuchsia (Fuchsia procumbens—Onagraceae). The 42d
parallel, where many other species drop out, cuts off the northern end
of South Island. The fact that this unnatural boundary does not co-
incide with Cook Strait between North and South Islands suggests
the geologically recent separation of the islands.
In addition, certain plant forms drop out at various points. For
example, the numerous epiphytes of the mixed rain forests of the west
coast of the South Island are conspicuously lacking in the rain forests
of Banks Peninsula, the vicinity of Dunedin in Otago, Southland, and
Stewart Island. The composition of these mixed rain forests changes
also with variations in the atmospheric humidity, the soil moisture
and composition, the altitude, and perhaps the geological and biological
history. The dicotylous-taxad association of the better-drained land
gives way to a much more pure association of ‘‘ white pine,” kahikatea
(Podocarpus dacrydioides) in poorly drained or swampy lands, as
formerly existed in some prominence in Canterbury on the dry side
of South Island. Here the deleterious effect of the drier air, inimical
to the best growth of most rain-forest trees, was compensated for by
the greater and more steadily available ground water. The swamp-
land forests of Stewart Island are dominated by another tree, Dacry-
dium intermedia. Altitude, or the changed environmental factors
that go along with changes in altitude, also cause changes in this
mixed association. As it rises higher on the mountains it is gradually
replaced by southern-beech forests. On Mount Egmont, however,
they give way to another altitudinal forest association composed of
kamahi (Weinmannia racemosa—Cunoniaceae) and the New Zealand
NEW ZEALAND—EGBERT H. WALKER 329
cedar (Libocedrus bidwillii—Pinaceae). ‘These numerous changes are
dealt with by Cockayne (7) and in many special accounts and reports
of the vegetation of certain localities.
The southern-beech forest extends intermittently, essentially from
the central plateau of North Island to Southland in South Island (pl.1,
fig. 1), but is lacking in a stretch of Westland and does not occur on
Stewart Island. In former times it was often called birch, but this
erroneous designation is seldom used today. These forests are usually
composed of just one species of Nothofagus, or at most two. The spe-
cies change rather strikingly with changes in altitude. The forester
is not content with the taxonomist’s five species of this genus, but is
able to recognize in this complex a great many more entities or varia-
tions, most of them more or less significant to him in dealing with this
economically important forest association. It is now the most im-
portant native timber tree, since the kauri is practically gone as a source
of commercial timber. Hence, the foresters are working out ways of
conserving, extending, and using the beech forests of South Island.
Soil conservationists are interested in Nothofagus forests, for these
can prevent erosion on mountain slopes, if man would only let them
alone or aid Nature to restore them. But man’s ax and fire have laid
waste large tracts of mountain slopes. And now another danger
threatens the beech forests. ‘The deer which man introduced for
sport, or their progeny, are busy browsing and trampling the young
beech seedling so that in places the undergrowth of young seedlings
has disappeared. The solution for the regeneration of these forests
seems to be the extermination of the deer, or at least the keeping of
the population under control at a much lower level than it is today.
Indeed, the problems of future forests involve many factors in this
fascinating country.
THE FLORA, OR THE PLANT SPECIES COMPRISING THE
VEGETATION
Many species of New Zealand plants are possibly unimportant in
considering the structure, origin, and distribution of the vegetation or
plant associations. They are, however, extremely important in rela-
tion to the flora as a whole and the understanding of its origins,
affinities, and distribution. There are too many interesting species to
deal with them at all thoroughly here, but some aspects of the flora
can be reviewed. ‘The marine and fresh-water algae (pl. 10, fig. 2),
and the other lower cryptogams must be omitted, but with full recog-
nition of their great interest and importance. Although New Zealand
is famous as a pteridologist’s paradise, the ferns, too, must be largely
ignored. (See pl, 5, fig. 1.)
One’s first impulse in looking at the New Zealand flora is to divide
it into two categories, the native plants and those introduced by man.
330 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Each seems a separate unit and the visitor with limited time tends to
pass over the introductions. There are, however, so many interesting
and important botanical features in both floras that neither should
be omitted.
THE NATIVE FLORA
The indigenous flora of New Zealand is among the most distinctive
of the world’s floras. It is not especially rich in number of species,
Cheeseman’s Manual (4) listing only 1,763 species, but it contains a
wealth of fascinating members, mostly belonging to genera and families
unfamiliar to the traveler from northern lands. About 78 percent of
the indigenous species of ferns and seed plants are endemics, that is,
are not found anywhere else in the world. This becomes 88 percent
if the ferns and monocotyledonous plants are excluded, and even
higher if only the forest species are considered. Forty genera are
found only in New Zealand. Among the more conspicuous or signifi-
cant are the lacewood (Hoheria—Malvaceae), one of the few natives
which has conspicuous and ornamental flowers and is deciduous, and
two composite genera, Haastia and Raoulia, which form the ‘‘ vegetable
sheep” (pl. 3, fig. 2, and pl. 10, fig. 1), for which New Zealand is noted.
The reasons for the occurrence of these species and genera only in
New Zealand are mostly obscure, and leave much yet to learn. They
may have evolved here or they may have had a widespread distribu-
tion and have died out elsewhere. It cannot readily be determined
that these represent a flora which developed in New Zealand, rather
than one that came from some other part of the world in remote
geological times, but the evidence is that such flora did evolve here.
It is called the palaeozelandic flora and certain genera are tentatively
assumed to belong to it. They include the three strictly endemic
genera just mentioned, besides others now found elsewhere but which
probably originated here. Among them are the distinctive New
Zealand pine (Dacrydium) in the Taxaceae, the New Zealand broom
(Carmichaelia—Leguminosae), the widespread shrub Coprosma (Rubi-
aceae) of distinctive divaricating habit and many species, and the
woody genus Hebe (Scrophulariaceae), which is often combined with
the widespread genus Veronica with mostly herbaceous species.
The majority of the nonendemic species and genera are found also in
Australia. One might suppose this indicated a relation between the
floras of the two regions, but too much emphasis should not be given
to this numerical superiority of the Australian element. It is just as
important in considering this relationship that certain prominent
Australian groups do not occur in New Zealand. Thus in these
islands there are no native eucalypts (Hucalyptus—Myrtaceae), bottle-
brushes (Callistemon—Myrtaceae), Melaleuca (Myrtaceae), wattles
(Acacia—Leguminosae), and other significant genera, especially
NEW ZEALAND—EGBERT H. WALKER 331
legumes, and only two genera, Persoonia and Knightia, of the con-
spicuously Australian family Proteaceae. Few of the Australian ele-
ments are familiar to travelers from the North Temperate Zone. One
that a visitor to the New Zealand forest will soon meet, however, is the
supplejack (Rhipogonium scandens—Liliaceae), a conspicuous vine
which hangs from the tops of the trees. Another is the genus Celmisia
(Compositae), whose many New Zealand species are among the chief
ornamentals in the montane and alpine vegetation (pl. 3, fig. 2).
Besides the Australian element there are indigenous species and
genera, found also in the Malay Archipelago and the Pacific region,
but they are fewer than the Australian ones. The kauri is of this
group, for certain other species of Agathis grow in Australia, New
Caledonia, Fiji, and elsewhere. The climbing pandanus (Freycinetia)
(pl. 2, fig. 2) and the only New Zealand palm (Rhopalostylis) (pl. 8,
fig. 1) belong to definitely Malayan-Pacific groups.
The third group of indigenous but not endemic elements is the sub-
antarctic, most conspicuously represented by the southern-beech
(Nothofagus), already mentioned. There are a good many genera and
some species in this group with distribution around the southern
Pacific, some with extensions into more northern regions. They are
of much interest to plant geographers in understanding the past geo-
logical history of this whole southern region.
Finally, there are New Zealand plants with a world-wide distribution
constituting the cosmopolitan element. An example is the widespread
bracken, Pteridium aquilinum, the New Zealand variety of which, var.
esculentum, grows to such great size, as already noted. Most of these
cosmopolitan species are seashore or littoral plants.
These floristic elements and their origin have been discussed by
various authors, including Wallace (27) and Cockayne (7). The sub-
antarctic flora has been most thoroughly dealt with by Hooker in his
introduction to the Flora Antarctica (14) and by Skottsberg (24).
BOTANICAL DISTRICTS
More significant to the traveler than the origin of the indigenous
elements of the flora, is the distribution of the species within the islands.
This has already been mentioned in connection with the various plant
formations and their occurrence, but consideration of the species
rather than the formations brings out more clearly the botanical
divisions of New Zealand. Through a lifetime of study of the vegeta-
tion and plants of this area, Cockayne (7) has divided the country,
exclusive of the outlying islands, into 16 more or less distinct botanical
districts. There is, of course, some disagreement with these divisions
and other workers will alter them when new studies and interpre-
tations are made, as Cockayne himself predicted. The districts are
332 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
shown on the accompanying map, figure 2.” The following char-
acterizations are largely derived from Cockayne’s publications and
the writer’s personal observations. Only a few of the many outstand-
ing species are here mentioned.
Northwest of North Cape lies the seldom-visited Three Kings Dis-
trict composed of a small group of islands with this name. Their
flora includes 10 species of plants not found elsewhere in New Zea-
land, 6 of which are obviously related to New Zealand species, 3 have
relatives only toward the north, and 1, a species of Chloris (Gramin-
eae), is of almost cosmopolitan affinity, although not found elsewhere
in New Zealand. The impoverishment of this flora by introduced
grazing animals, especially goats, and its recovery after these were
removed is an interesting study with significant implications for other
likewise devastated areas. These Islands have recently been treated
botanically by Oliver (23) and Baylis (8).
In the North and South Auckland Districts there are more than 100
species of plants which do not occur farther south, or extend only a
short distance beyond the southern limit at approximately the 38th
parallel. The kauri tree has already been mentioned in this connec-
tion. The taraire (Beilschmeidia taraire), usually the dominant tree
in the kauri forest, does not go farther south as does the other impor-
tant kauri associate, the tawa (B. tawa). Several ferns are found
only here, as well as the parasite Cassytha paniculata (Lauraceae),
which resembles, but is no relative of, the dodder (Cuscuta) of wide
distribution. This parasite genus occurs in the Pacific area, Australia,
and the gumlands of northern North Auckland. It is a hazard to
walking wherever it grows as it binds together the manuka and other
shrubs by dull green resistant cords. An interesting phenomenon is
the occurrence on the small islands adjacent to these districts of
species or varieties similar to those on the main island, but with larger
leaves, flowers, or fruits, among other differing characters. These
islands are difficult to reach, but one can often find these distinct
plants growing in private gardens, nurseries, and parks where enthusi-
astic New Zealand botanists grow them. Several species have their
northern limit in the South Auckland district, one being the silver
beech (Nothofagus menziesii). Although there are no natural grass-
lands in this area, there are extensive man-made pastures with certain
distinct plant species, mostly grasses. The mild climate makes it
possible to grow certain citrus fruits and other subtropical crops with
success.
¥ After this map was prepared Cockayne and Allan (8) proposed an additional district, the Sounds-Nelson
Botanical District, comprising the South Island portion of the Ruahine-Cook District, which formerly
straddled the Cook Strait. This district was recognized in Cockayne’s larger work (7), which is followed
in this survey, but which does not have a map suitable for reproduction here.
NEW ZEALAND—EGBERT H. WALKER 333
THREE KiWGS Is 5° 1
North Cape
C. Maria v. Dieman \
Doubtless Bay
Mangonui®
R
Hokianga Hr. at
ae
2 Qonegr BARRIER I.
| EXPLANATION. C. Colville
2
BOTANICAL (DISTRICT.
AUCKLAND : Bay of Plenty
r foe eWHITE 1
Three Kings ....
North Auckland
South Auckland d } East Cape
(2) Waikato Subdistrict ay
4, :-(b) Thames Subdistrict NORTH : UA
Volcanic Plateau ag a \ Fo
East Cape... oobr A: K
Berea epesaui es ae \ Me paremoany
| Rushine-Cook .... oi @Gisborne
ISLAND Y
(a) Wellington Subdistrict
(b) Marlborough Sounds Subdistrict
North-eastern South Island ....
North-western South Island ...
Eastern South Island
Western South Island
North Otago .... eee a!
South Otago .... =< an
Fiord nee = 2
Stewart :
Se B.
ri
Sere}
2457 HANMER
oN pba
wt ae
Greymouth
Taramakau R.
“runui R.
2
= AWaimakariri R.
a @LCHRISTCHURCH
Banks Penin.
Jackson Hd.
ez Tt,
xe ingBiigy OR
6 ES en ‘ta R,
Big Bay. Sout § u
. j, S —
Milford Sd.¢ ‘/%*. = et aitaki R. Map of
aru
™ NEW ZEALAND
L. Te(A nau we rg Aira i showing
Sy sy 2a? // DUNEDIN * Tetra ets
A 4 Ry AT 42,, Proposed Botanical Districts
oy i C teri. R.
; J rs - Tach 00
L t iF, A Invercargill fy, {Ma R. 100 75 50 25 O 1
feces canter or am “BBet Pt, =
Pane =, English Miles
a ° of Ta R. *
STEWART istAND Ey
Ss ge 5 L. Cockayna Des.
=
=n
Figure 2.—The botanical districts of New Zealand, according to L. Cockayne (6).
304 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
The first natural or tussock grasslands as one progresses southward
are found in the Volcanic Plateau District. Here one begins to see
certain characteristic subalpine shrub formations, especially of
monoao (Dracophyllum subulatum—Epacridaceae). The volcanic
mountains, three of which are still active, raise their summits far above
timber line and support a fascinating alpine flora.
The northern tip of the East Cape Botanical District is within the
northern plant zone marked off by the 38th parallel, so contains some
northern plants. The southern part in the Hawkes Bay region is
drier, hence it has certain agricultural possibilities not found else-
where. Maize or Indian corn (Zea mays—Gramineae) is grown for
its grain, but elsewhere mostly for fodder.
To the botanist Mount Egmont is probably the first attraction of
the Egmont-Wanganui District in the southwest portion of North
Island. However, its high mountain flora is less rich than is that in
the mountains of the central plateau, which are visible in good
weather from Egmont’s higher slopes. The zones of vegetation on
this isolated volcanic cone, however, are very vivid. There are few
plants endemic to this district and the southern-beech forests are
absent. rvyya] aly eyez % -o2
au 2 SANYC JST] AHALWWIS - J
W209: SLNMY M3d LUA HLM STRAW -EG
eae ee 24
Ww Cp: ( 311Y31S) SQNVD ON] -BanA}-79
‘W2 CP: NvOUNDIaNLY Baddf>7-2q IV)
‘W2O] » 3or
WW JC; NEPUNIIAN Badd) -'
WOO" : asAv] sNsaLo —-D
WD OC: (Ci6! -2\6\ NI 5nq} NYINSId9eL | St
( 299) NI NiTaw 48 9Nq) WHITOIN - b/
Snag oa
|
|
ee |
SNIAO[AI— "6p 6] “40dayy URIUOSYATUIG
Smithsonian Report, 1949.—Movius
1. THE SITE LOOKING EAST
(See explanation of plates, p. 367.)
2. TRENCH DUG THROUGH RIVER-LAID TERRACE DEPOSITS AT LA COLOMBIERE
(See explanation of plates, p. 367.)
Smithsonian Report, 1949.—Movius PLATE 4
1. MAIN TRENCH NEAR REAR WALL OF ROCK-SHELTER
(See explanation of plates, p. 367.)
2. SECTION NEAR WESTERN END OF LA COLOMBIERE
(See explanation of plates, p. 367.)
Smithsonian Report, 1949.—Movius PRATE 5
1. BROKEN MAGDALENIAN BONE OBJECT FOUND AT LA COLOMBIERE
(See explanation of plates, p. 367.)
2. ENGRAVED PEBBLE FOUND AT LA COLOMBIERE
(See explanation of plates, p. 367.)
Smithsonian Report, 1949.—Movius
OBVERSE AND REVERSE SURFACES OF ENGRAVED PEBBLE FOUND AT LA
COLOMBIERE
(See explanation of plates, p. 368.)
Smithsonian Report, 1949.—Movius PLATE 7
) = Ru
SS \ gout
\\ iy + \ Ml
SN \
5)
OUTLINE DRAWINGS OF ANIMALS ENGRAVED ON PEBBLE FOUND AT LA COLOMBIERE
(See explanation of plates, p. 368.)
ROCK-SHELTER OF LA COLOMBIERE—MOVIUS 367
ACKNOWLEDGMENTS
On behalf of the expedition, it is my pleasure to take this oppor-
tunity of thanking Professor Marcel Thoral, Director of the Labora-
toire de Géologie, Faculté des Sciences de Université de Lyon, and
Dr. Jean Viret, Director of the Muséum des Sciences Naturelles,
Lyon, for their generous assistance and kind cooperation in connec-
tion with our investigations at La Colombiére. It is also my privi-
lege to thank Messrs. Joe E. Cason and Raymond H. Thompson,
graduate students in the Department of Anthropology, Harvard
University, for their expert help in deciphering the outlines of the ani-
mals engraved on the La Colombiére pebble and reproduced on plates
6 and 7 of this article.
EXPLANATION OF PLATES
PuiatTe 1
General view of the rock-shelter of La Colombiére showing the Late Pleistocene
terraces of the Ain Valley in the vicinity of Poncin. The site, which is 46 meters
long and some 12 meters wide, faces due south and directly overlooks the Ain
River.
PLATE 2
La Colombiére. Photograph taken by Dr. Mayet and M. Pissot in April
1913, showing the stratification revealed in a trench dug in the western end of the
site.
PLATE 3
1. The site looking east, showing the undisturbed surface of the deposits after
clearing. In the left foreground what appeared to be the top of a filled-in lower
cave may be noted.
2. A deep trench, the outer or southern half of which was 2 meters wide, was
dug through the river-laid terrace deposits at La Colombiére to determine whether
or not the site had been occupied prior to its invasion by the river in Late Pleisto-
cene times. Note the rails and trucks, used for removing the excavated material,
in the foreground.
PLATE 4
1. Near the rear wall of the rock-shelter the width of the main trench was
expanded to 4 meters. Bedrock was finally reached below a total thickness of
11.85 meters of river-laid sands, silts, and gravels, of which the so-called La
Colombiére terrace was formed. There was no evidence of an early occupation
of the site.
2. Section near the western end of La Colombiére showing the intimate and
direct association of the gravelly occupation layers with the sands laid down when
the Ain River invaded the site during the late stages of the formation of the 20-
to 23-meter terrace.
PLATE 5
1. Broken Magdalenian bone object perforated at one end and decorated with
chevron pattern crossed on one side by parallel lines.
2. The interesting engraved pebble described in the accompanying text was
found in the main (Upper Aurignacian/Gravettian) occupation layer at La
368 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
Colombiére in direct association with an extensive hearth (diameter: 2 m.; thick-
ness: 40 em.), on the surface of which the meter scale is resting, as shown in the
photograph.
PLATE 6
The beautifully engraved pebble discovered during the 1948 season in the
Upper Aurignacian occupation layer at La Colombiére, near Poncin (Ain).
Upper: The obverse surface of the pebble showing a very realistic horse, which
is by far the most easily recognizable of any of the engravings on either face.
In addition, there is an extremely finely drawn male reindeer (with shed antlers),
an ibex, and two carnivores (possibly bear). The heads of the two former animals
appear upside down on the right of the photograph. At least two other as yet
unidentified ungulates may also be distinguished.
Lower: The reverse surface of the pebble on which a second horse and an out-
standingly fine woolly rhinoceros are depicted. There are also the heads of two
other partially completed rhinoceroses, and the outline of the body and legs of
what appears to be a cervid of some type.
PLATE 7
Outline drawings of some of the most easily distinguished animals engraved
on the interesting pebble found at the La Colombiére rock-shelter in an Upper
Aurignacian context. Nos. 1-4, upper surface; Nos. 5 and 6, lower surface.
On the basis of the exceedingly skillful and very realistic portrayal of the forms
represented, it is at once apparent that the drawings reproduced come from the
hands of men who knew their models intimately at first hand. In particular, the
engravings of the horse (No. 1) and the woolly rhinoceros (No. 6) are so remarkably
alive that there is no mistaking the subjects. Note that all four legs are shown
in the case of Nos. 1, 2, 3, and 6, and that the hair on the neck, forelegs, and
shoulders of the rhinoceros has been cleverly arranged so as to suggest shading.
Figure 1
Prehistoric archeology deals with the immense span of time between the first
appearance of man and the beginnings of written record—a period of perhaps some
1,000,000 years’ duration. As indicated on this chart, which shows the relative
duration of prehistoric time, during approximately 49/50ths of this period man
was in the Old Stone Age, or Paleolithic and Mesolithic stages, of cultural
development. These stages cover the entire Pleistocene or Glacial Epoch, as well
as much of Early Post-Glacial or Recent times. During the Pleistocene the
northern regions and mountainous areas of the globe were subjected four times
to the advances and retreats of the ice sheets (those of the Alps are known as
Giinz, Mindel, Riss, and Wiirm), river valleys and terraces were being formed,
and profound changes were being induced in the fauna and flora of the Earth.
Throughout the entire span of the Old Stone Age (including both the Paleo-
lithic and the Mesolithic periods) man was a food-gatherer depending for his sub-
sistence on hunting wild animals and birds, fishing, and collecting wild fruits,
nuts, and berries. On the basis of the levidence obtained: to. date, particularly
that from western Europe, it is possible to recognize three main groups of funda-
mental traditions employed by our Stone Age ancestors in manufacturing their
stone implements. These subdivisions are as follows: (a) core tool traditions,
(b) flake tool traditions, and (c) blade tool traditions. The industry found in
1948 at the rock-shelter of La Colombiére by the Peabody Museum of Harvard
University’s expedition to eastern France belongs in the blade tool category; typo-
logically it appears to have close affinities with what is known as the Gravettian
stage in the Upper Paleolithic sequence of western Europe.
RONNE ANTARCTIC RESEARCH EXPEDITION, 1946-1948 !
By CoMMANDER FINN Ronne, U.S. N. R.
[With 8 plates]
My interest in polar exploration dates from 1909, when my father,
Martin Ronne, was selected to accompany Capt. Roald Amundsen
to the Antarctic, on the memorable expedition on which the South
Pole was first reached. That association continued until the great
explorer’s untimely death, after which my father went with Admiral
Byrd on his First Antarctic Expedition, 1928-30. During all this
time I had closely followed the detailed work involved in the planning
and successful execution of polar expeditions. Martin Ronne died
suddenly in Norway in 1932, in his seventy-first year. On Byrd’s
Second Antarctic Expedition, 1933-35, I had the good fortune to
follow in my father’s steps in the capacity of ski expert.
Upon my return to the United States in 1935, I began to plan for a
small Antarctic expedition of my own, on which we would sledge and
map the coast line from the Palmer Peninsula to the Ross Sea area.
I planned to be set ashore, with four men and sufficient dog power,
in the Charcot Island area by a Norwegian whaler and to be picked
up by the whaler several months later on the Ross Sea side. However,
an independent expedition did not prove possible at the time, and my
modest plans eventually were merged into the United States Antarctic
Service Expedition,? on which I acted as second-in-command of the
east base, on Stonington Island, in Marguerite Bay, Palmer Land.
EXPEDITION PLANS
In May 1941, I obtained a commission in the United States Navy,
in which I served until the end of hostilities. Off and on in my spare
time I began formulating plans for an expedition to the old base of
the United States Antarctic Service Expedition on Stonington Island,
a location I considered well suited to geographical exploration, since
within flying range lay the unexplored Weddell Sea coast line and
perhaps the termination of the mountain axis of the Palmer Peninsula.
1 Reprinted by permission from The Geographical Review, vol. 38, No. 3, 1948.
2 English, R. A. J., Preliminary account of the United States Antarctic Expedition, 1939-41, Geogr. Rev.,
vol, 31, pp. 466-478, 1941. The operations from the east base are described on pp. 469-473.
369
370 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
I also planned to carry on an extensive scientific program of at least
a year’s duration with a small group of competent men. To eliminate
the necessity of sending the expedition ship back to civilization, I de-
cided to let it freeze in at a small cove close to the base; the expedition
members themselves would man it. Through Congressional action
I was able to obtain from the Navy Department, on a loan basis, a
sturdily constructed ocean-going wooden tug (pl. 1, fig. 1). From
the Army Air Forces, Office of Research and Development, many
articles of equipment were obtained for testing, including three air-
planes, two snowmobiles (‘‘weasels’’), camping equipment, and numer-
ous types of clothing.
Only those who have had the experience of planning and organizing
an expedition can fully appreciate the enormous amount of work in-
volved, especially in the matter of financial backing. I contacted
many scientific organizations and foundations interested in Antarctic
research, but with few concrete results. Under the auspices extended
by the American Geographical Society of New York, and by selling
the exclusive news rights of the expedition to the North American
Newspaper Alliance, Inc., and with a few subscriptions from interested
friends and a contract with the Office of Naval Research for the
scientific results to be obtained, I was finally able to get the expedition
under way. It was not before December 8, 1946, however, that I
was definitely assured of the minimum required financial support.
Through continuous hard work day and night, and spurred on by a
strong determination to sail, we were ultimately able to assemble the
thousands of needed articles of equipment essential for a polar
expedition.
UNDER WAY
On the afternoon of January 25, 1947, we threw the mooring lines
off our ship, christened The Port of Beaumont, Texas. 'The road had
been long and rough, and many obstacles lay ahead, but we were on
our way at last. Brief stop-overs were made at Balboa, C. Z., and
Valparaiso and Punta Arenas, Chile. To avoid the dangerous roaring
forties with our topside weight, which included 8 airplanes, 112 drums
of gasoline and lubricating oil, and 43 northern sledge dogs, we sailed
in the sheltered waters of southern Chile’s inland passage. On board
ship much work was done to the three airplanes, particularly the
Beechcraft C—45 exploratory plane, in which a complete electrically
operated trimetrogon camera unit was installed, and also a radio
altimeter and extra transmitters and receivers for long-range commun-
ication. Two additional gas tanks were placed in the fuselage, so
that the plane now had a maximum cruising time of 9 hours.
Our passage between Cape Horn and Marguerite Bay was fortu-
nately very smooth, and we encountered only a relatively small amount
ANTARCTIC RESEARCH EXPEDITION—RONNE 371
80° 7s° 70° eke 60° perien? a 65° o
RONNE ANTARCTIC RESEARCH EXPEDITION =. st¥ LK"
— 1947-1948 a OE ——
PROVISIONAL SKETCH MAP 2 tne
PREPARED BY THE AMERIGAN GEOGRAPHICAL SOCIETY See pees LQ
Statute miles
100
100 ° 290
os 64
Features named by the Expedition. ......4¢. Tuve SS
Rlightilinessenctcrcm-cteneerte tc.
New coastline ....c.-c.ccccccceseeores Re
A Miceisont a
are
ca
ee" 662
6"
=" =
40° 702
oe
J“? aA
Lowell Thomas If tes ,
Mt.Horne $ ital
Hauberg Wee ZKe s
ne
2 Tie peataze
4°
is, ” 4.Haed
er fi 000
xX ge wie! Skene 79
° plateau
a ie
~v
~ COATS
& ly LAND
as)
mT
x
CSR ae aera
Figure 1.—General map illustrating the principal explorations of the Ronne
Antarctic Research Expedition.
312 ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
of pack ice and bergs. We anchored off our main base in Stonington
Island on March 12, 1947 (pl. 1, fig. 1). Shortly before my depar-
ture from the United States, I had learned through the Department of
State that, 2 years before, the British Government had established
a permanent base on the island in continuance of a program begun
in 1943 under wartime secrecy. It had also established and was
maintaining other bases on the Palmer Peninsula. Great Britain,
through the Falkland Islands, has long laid claim to this sector of
Antarctica, and these five bases were under the administration of the
Falkland Islands Dependencies Survey. However, it has been the
policy of the United States not to recognize the claim of any govern-
ment in the Antarctic, nor has the United States Government made
any claims of its own.
I had knowledge that in the 6 years since the departure of the United
States Antarctic Service Expedition in 1941 ships from several other
countries had visited the American camp site. In 1943 the Argentine
gunboat Primero de Mayo had visited the American base, and an
Argentine ship and two Chilean vessels had been there shortly before
we arrived in 1947. Upon our arrival we were greeted by the British
leader, Maj. K. S. Pierce Butler, commander of the Falkland Islands
Dependencies Survey, 1947-48, and later we became acquainted
with the other 10 men,, who were occupying their own quarters
constructed about 200 yards from the American camp site. As I
investigated, I was appalled at the amount of wanton damage that
had been done to the three large and three small buildings constitut-
ing the American base. After much hard work the base was made
livable again and occupied by our expedition (pl. 1, fig. 2).
GEORGE VI SOUND TRIP
I decided to attempt to establish an operational base at the south-
east corner of George VI Sound, 300 miles to the south, before we
anchored the ship in its final position for the winter freeze-in. I
hoped to be able to set up a cache of gasoline, stores, and one of our
two weasels at this halfway point and thus facilitate the transporta-
tion of such heavy equipment into the field at a later date. The
attempt did not prove successful, and we later abandoned this loca-
tion in favor of Cape Keeler, on the Weddell Coast. However, the
journey did reveal some new features.
At 5 a.m. on March 23, after riding out a number of strong south-
easterly gales with velocities of as much as 60 miles an hour, we hoisted
anchor and steamed south along the Falliéres Coast, past Cape
Berteaux, to the entrance of George VI Sound. A group of islands,
which I named the Bugge Islands, were found in approximately
69°10’ S., 68°55’ W. Three large islands, the largest about a mile
ANTARCTIC RESEARCH EXPEDITION—RONNE 373
and a half long, stretched for some 5 miles in a northeasterly direction.
Numerous small islands were 15 to 20 feet high, and most of them
were bare. ‘The larger islands, however, were covered with snowcaps
more than a hundred feet high, though bedrock was exposed at the
water’s edge.
Thanks to the skillful piloting of our skipper, Commander Isaac
Schlossbach, U. S. N. (ret.), who was also second-in-command of the
expedition, the vessel moved steadily among the huge shelf ice and
glacier-formed bergs that blocked the entrance to the sound. At
4 o’clock in the afternoon we reached 69°20’ S., our farthest point,
a new record for ships navigating in this region of the Antarctic.
To make a landing anywhere was virtually impossible. I was unable
to see the 150-foot ice wall which marks the entrance to the sound and
over which I had traveled in 1940, but it was obvious that these
tabular bergs had recently broken off from the shelf ice of the sound
itself.2 (Our plane flights several months later revealed that the
shelf edge was discharging bergs such as those among which we were
now sailing and that the face of the shelf had moved back 35 miles
in 7 years.) Not only did the conditions ahead offer an immediate
danger to our only means of transportation back to the civilized world,
but had we continued to search for a suitable landing place farther
south, a sudden change in the weather so late in the season might
have blown these huge bergs in upon us and blocked exit for another
year, The risk was too great, so I gave orders to return to the open
Marguerite Bay.
Our ship was moored in Back Bay, a cove a third of a mile from the
base. As temperatures fell during the first week of May, it became
safely frozen in the bay ice, and it remained so until the summer thaw
of the following year partly released it from the icy grip (pl. 2, fig. 2).
WINTER PREPARATION AND TRAIL PLANS
The winter passed rapidly, and an immense amount of preparatory
work had been accomplished by the time we were ready to start the
field program. On July 15, [ led a sledge party up to the plateau,
6,000 feet high, 17 miles east of our base to establish a meteorological
station. This station was manned and operated during the entire
flying season and, in conjunction with a station later established at
Cape Keeler, 125 miles to the south of the Weddell Sea side, made
it possible for H. C. Peterson, our meteorologist, to forecast the highly
variable weather with good accuracy.
By August all three planes had been unloaded from the ship,
assembled, and made ready. I intended to use the single-engine,
8 Ronne, Finn, The main southern sledge journey from East Base, Palmer Land, Antarctica, in Reports
on Scientific Results of the United States Antarctic Service Expedition, 1939-1941, Proc. Amer. Philos.
Soc., vol. 89, No. 1, pp. 18-22, 1945. This journey confirmed the insularity of Alexander I Island.
374 | ANNUAL REPORT SMITHSONIAN INSTITUTION, 1949
650-horsepower Norseman plane, which had been especially designed
for cold-weather work, for flying gasoline caches to various points
along the Weddell Coast. Weeks of continuous overcast, however,
prevented us from completing this program. In November, by the
time the weather had improved sufficiently for the southern explora-
tory flights, we had deposited 28 drums of high-octane gasoline at the
Cape Keeler Advance Base.
During the winter Major Butler and I had decided to cooperate in a
surface field program. A joint British-American Weddell Coast sledge
party consisting of four men, Major Butler and surveyor Douglas
Mason of the F. I. D. S., and two members of my expedition, Walter
Smith, navigator, and Arthur Owen, dog driver, were to cross the
plateau by dog team to the eastern side of the Palmer Peninsula.
They would sledge south along the Weddell Coast to Cape Knowles,
beyond which the territory was virgin so far as surface travel was
concerned, though the United States Antarctic Service Expedition had
made an exploratory flight as far south as Mount Tricorn. When the
surface party reached Mount Tricorn, the two Americans, now forming
the Ronne Weddell Coast party, would continue southward into the
unknown as far as supplies would permit, in order to establish ground
control points for our aerial mapping. So long a sledge trip without
the aid of supporting dog-team parties would be possible because our
Norseman plane was to deposit several caches of man and dog food
along the route of travel (fig. 2).
RESCUE OF BRITISH AIRMEN
As a first step the small British Auster plane took off for Cape
Keeler on September 15, followed by the larger Norseman plane loaded
with 3,000 pounds of trail supplies. The smaller plane was to make
the initial landing in the field and pick out a suitable landing area for
the heavily loaded Norseman. The Auster did not have adequate
radio communication and in flight unfortunately became separated
from the other plane. When the Norseman did not sight the Auster
at the Cape Keeler rendezvous, a search was made, but darkness and
bad weather were approaching, and Capt. James W. Lassiter had to
turn back. By 10 o’clock that evening a storm had set in, and the
British plane was still missing and unreported. Accordingly, I made
all the facilities of my expedition available to Major Butler for his use
in searching for the missing plane. Captain Lassiter and Lt. Charles
J. Adams made numerous unsuccessful searching flights in the over-
cast weather during the next 8 days. On the ninth day, when hope
was dwindling rapidly, Captain Lassiter located the three lost British-
ers walking back on the sea ice 40 miles south of our base. We learned
that they had actually landed at Cape Keeler on the day of the out-
ward flight and, when they were not sighted by the Norseman, had
Smithsonian Report, 1949.—Ronne
1. THE *“‘PORT OF BEAUMONT, TEXAS,’’ AT ANCHOR OFF THE MAIN BASE, STON-
INGTON ISLAND, MARCH 12, 1947
2. THE MAIN BASE ON STONINGTON ISLAND
Neny Island on the right.
Smithsonian Report, 1949.—Ronne PLATE 2
—
wif
1. WEASEL HAULING THE BEECHCRAFT INTO POSITION FOR FLIGHT AT THE MAIN
BASE
2. THE ICE-LOCKED VESSEL SEEN FROM THE GLACIER NEXT TO THE BASE
Red Rock Ridge in the background.
Smithsonian Report, 1949.—Ronne PLATE 3
1. THE MAIN BASE (LEFT FOREGROUND), STONINGTON ISLAND
Ship at anchor near ice cliff.
—
i .
2. DOG TEAM NEAR THE TERMINAL CLIFFS OF ONE OF THE BIG GLACIERS NEAR
hie BASE
Smithsonian Report, 1949.—Ronne PLATE 4
1. AT THE ENTRANCE TO NENY TROUGH
Heavily crevassed glacier front.
2. COMMANDER RONNE, W. R. LATADY, AND CAPT. LASSITER BESIDE THE C-45
AFTER THE LONG SOUTHERN FLIGHTS
Smithsonian Report, 1949.—Ronne PEATE 5
VIEW OF BASE AND
*
VIEW OF BASE AND NENY FJORD FROM 10,000 FEET
Smithsonian Report, 1949.—Ronne PLATE 6
1. NEW BEDFORD INLET
K-17 photograph taken late in the evening on the first flight south. Altitude of
plane, 10,000 feet, height of mountain in center of picture: 4,100 feet (taken
from ground survey).
2. CAPE KEELER, ON THE WEDDELL. SEA COAST
Advanced base used as an intermediate stop on flights into the unknown south.
Smithsonian Report, 1949.—Ronne PEATE 7
1. WRIGHT INLET, EAST COAST OF PALMER PENINSULA
At the head is Mount Tricorn (5,450 feet).
2. HEAVY CLOUDS NEAR STEELE ISLAND
The cause of our spending a night in the field and the end of our first long flight
south.
Smithsonian Report, 1949.—Ronne PLATE 8
(nn we
ee ate
1. ALONG THE SOUTHEAST COAST OF PALMER PENINSULA
Scaife Mountains in the center.
2. LASSITER SHELF ICE, SEPARATED BY OPEN WATER FROM LOOSE PACK ICE IN
THE WEDDELL SEA
ANTARCTIC RESEARCH EXPEDITION—RONNE
375
i XC. Eielson - ne
\ SCALE 1:3,000,000
mee {" Doileman I. Statute miles
Rey! ow 2 30
7 N\\c Sha
S/he Shartpnnesy] Route of Sledge Party ----»---
\ Steele I.
fe MEHIe Ce Bh Features named by
é wee Sys the Expedition........C Adams ~7!
, eet Elevations in feet
\ Hons inveev...............s600
Cruse in
Mt. Andrew Jackson > = Lamplugh Bay C7 Veesy Edge of shelf ice
13760 amy
Odom Bay Cc mepe id
C:Knowles
Mt Russell Owen & ee 5 Wn
pon OMt, Ward ae we ae ington
i Gruening Gis
72°
126
Is ureato fi AmMerica nner. ee ee ee ee eee 87
BreensGalleryrolArt 2 soe eee pera ee ee Oeier ere 47
ilprany eee n= he een Sasori k & Sates BA ete Seo SSS a ee 132
NationaleAirs iuscum=—eeee see wee ae aa ees eee Pe 122
Nationale C@olle ction fe time Atr (cee eee ee eee ee eee 42
iNanonalkGallerviolsArtx sess =2* san ee Ieee ae ean arene vee ee 27
Nea tonale\Guseuineee ssa Scie ss noe wr eae et SA ae Sek 2 eee 16
iNationaleZoolosicalabarks= == ssn 56 sa= sone snes eae AS eee 98
NCE NGA eee he oe iene ee ae eI AORN Ram HS IIA. Soe eee viii, 52
Adams, Chere ley Cs meee Bo at oe ne crete west Sle ot en de ea eee E 128
Administrative accountant of the Institution (ihomasy he Clark)a ses. == v
Administrative assistant to the Secretary (Louise M. Pearson) -_-_------ Vv
MDCT ULE OUCE Eta a= ann eke SaaS! ReS sees ree hes eee ee eee 127
Aldrich, Loyal B., Director, Astrophysical Observatory--...-..-------- ix, 113
PAU eTIMey Aur t hii Tet Ngeees See ewe Bee A epee Beka DE se HS th Seen sito Beye oy = 129
A Grit VWallliania pee me seers ee eee tfc ie 2 BL ewe ee eats vii, 23
Amencan Mistoriesl Association, Reportss. 62-22-2022 222 oo] eee 141
Anderson, Car] D. (The elementary particles of physics) _--_------------ 203
Anderson», Clinton=—P—(regent of the Institution) 2222522242222 2-2 2 222 v, 6
Animal behavi ior) Morevabout (Emest PS Walker)= 3292225322" " 2-2-2 261
Antaretie research expedition, Ronne, 1946-1948 (Commander Finn
ENO TIIAC) eee ea ae Sere a ey te ee ob aree 100s heer SA yn es 369
OPLOME A IOUSE Nae Aeon nes ae ete ee 7, 26, 59, 107, 131, 142, 148
CanalyZoneBiolociesloAred 222 92282. S52 22 SS sre sees eae 131
Iishitubkeromoocial Anthropology te a. oa one a ee ee ene eee a 59
INE OnaAlmGallenycOmAnr Gee = eee en oe ea oe eee ees 26
NationalZoolosicalebarika sania 92s As Nee oe eee es eee eee 107
TEVETUSW BA AYER (eipaeG | OVUGKG Ley ade ete et pn ted lp ly eM et ek pe 142
River Basin SURVEYSe erat cee thon Stes een Reh ee eee 59
STAM Se TAT 1 Te ae lar ne Ne ers wy hr oS er A Se eS a 7, 148
Arab rmanenineri Cane Corfe een ee eee se ee ee ee 22
Anchicvimstitute OLoNOrth Americas. 2a Sess oe ee eee Tere 23
PAC COWS Kal ered @niiay, Kwan 6 eae See Say See es RR ee ee 111
Srinser Ohimpelsaylonrks ose wie hae ee on ee Oe SE ee nee os 41
Ani Murleckure torxteenth: = occa eee es see no a een eee eee 8
Assistant Secretary of the Institution (John E. Graf)__-__--------- Wi On 4d ae
Assistant Secretary of the Institution (J. L. Keddy) _------------------
INSETO Diy sical’ OOSCEVALOLY oe oes ae ee eee eee a seca cs ee rg, 1c}; 109
Astrophysical Research, Division Off. =) o> 5922 s2s—n=—seeo == ix, 109
Radiation(and Organisms, Division Of- 229-2222 os 22 =. = bey 1}, Wi
4 BSCS) 00 cr ge ae a in a ot aa ag Sg eg ln th ee ee Seat 109
SIR Gi Sh So OD, a Ree Sieae ek ncaa (hela Mic ge Ae emesis Se ee oe AO a ix
INECHISOM OSC DN OAT ae Se ene ae aan ee e ee ee eaeia aeeoe ae 119
Attorney General of the United States (Tom C. Clark, member of the
MS GUC WETOM oe ae ae ene eee ee oe pete ote ere oat es Vv
ANISH OV ee ce cee ee a ee Sr eats eee aa ie ore 79
Australia-National Geographic Society-Smithsonian Institution Expedition. 21
ING CTV BeAr ea eee ere 2 een ee sie one es eee vii
Acyl SAGTIAG GN pect eee mee eee a et ete ee nes eee vi
B
ASA Cl wams GOnG OU tps ese 2 a el ae ee DS ee ee ae 75
Barbeaul ©, Mess.s02 = BPs tt ey ee Be eet ee 57
412 INDEX
Page
Barkley, Alben W., Vice President of the United States (member of the
ln SiG Uti On) Me ee os oe ee eee ieee oes ee eee ee ee ee eee v, 4, 6, 20
IS ATES diy che 2 eps regen vk EL EAS i oes Be Rr a cl vi
IBA TESOL OR meets As LM bie en eg ges Mer ee de ee Se vi
Bassler: shai s ca seh ria eae Ree AUR aes AN eo Oe ee ee vii
Bauxars JA JOSep hick = tee TS Se a en a ies es ae | ce 74
Bavier shred erick lees 20 eee ers et od i eee en a gc eer ara vi
BY GFEN GS alpsei eyStS EA eat ae nl Aa i oe a apres eae ea IMS EG De Metin ns 14 aA ad ahd hse vii
Beal Git ond el. See ere ts te A gee AE a tS ene es ee 41
Beaulac: Willardiiaed Sos. ey. Se Bee ee La ee ee 60
Beers, Stephen tue. 2225 Sas bere SOE. hapa ape ces Sa 5 pee a ee a ix, 121) 122
Beggs, Thomas M., Director, National Collection of Fine Arts_.__-__--- ah
41,
Belin, Ferdinand Lammot, Vice President, National Gallery of Art §'viii, 25, 26
Bell aRiobenty tess Sar, ees Ss ey nee Ne ge AOE ee tale eee e ee 63, 80, 81
1 XSI Ohi s pa Dg el egies ae ag Sle eRe eae aimee A Nira wae S Beye! reas ewes oe © Wie Pye ees vii
dB Yon al oie [ts] Ree aes perenne hy tie eee ne ery eT AY eS Sorts el Ee ere vii
BembtmeAM Ginn Coe eee oe es te ere eee ek, pe ee vi
STAC Ewell Creel eee is teres ee ate eas Nee Re eee ye oe ea er ee vi
BlakersVini@s ted 26 a lis? Ce Se Se ce ae: eye eee ae vi
Bliss Robert AWO0dSH = be sen sok ho eee ee ee ee ee 7,41
IBIS ESAS UC aot es Bes sg aa ee a a ee 74, 75
HEY GY 0} OVS) pal ©)2 0) reac See | ae amp tT ee RE OR on eg es ree ere NP J ve Se § 120
BOSS WING renin bra eT Se eee fe rcs ee ee Os ee ee ea ee vii
BB Givin oe WAS EGE Uae OEE ee Se Ee i TNR ch a I alee vi
IBOWO Meee GI CTs see eee ae el Rage pe YR Sareea et ea 102
IBOWSlTer., WAe Mie een og ae es ke eget eee el cha a A See es et 23
Brannon, Charles F., Secretary of Agriculture (member of the Institution) - Vv
| BD eh ine) ie © pk ial cena AREA ine peel ayia a Se Ee eC EUR AS ek RN oS ks 75
STO wira Ae Va eS ee Se ee Ge eee ee ee a ge eee Be oe vil
TOs Wikeedins see eee cat re, See tet lee eee Sy Nt oy eS vi
Buchanan plies ee eee rae Se eee 8 reset Ae noms ied Vpted BFS) Tea aS BT vi
lekoial, \yiallbeyan Wife ee ee ae SNE ene A” RERUN IOT SS RENE |b ermine ee! 70
BEES Tags Ear Lo TG Ui as sas ct a yc kl 67
Bush, Vannevar i(tegent) of thedlnstitution)2 2 = == = 2-2 en ee v, 6, 149
1 BSAA PR i wa Sele eh Rr eg mleeieyac ey le reagan NIN * See, ele me ge Py nie OW A vi
C
(CEH Gaels tape pete terreno in pean name 6 A npeign te A Lae sen Pe + Ayes bes vi
Cairns, Huntington, Secretary-Treasurer and General Counsel, National
CallenvzotpArte tare chs Sone a ore oo Ee eae viii, 25, 34, 40
alice Ose 1 lS ae see ee ra ye a 63, 64, 81
Canale Zone B1ologi call Ate seems ey ee eee ix, 14, 126
ISGalare ports fe seme el ere es eo ee ee ee 131
JIMpPLOVEMentsuNAd ey c= 2 te eee a re ee See eee 126
Nigrewurgent NC6dS soe een ee ee 129
Raintalistablessewee eee ea Ie OT re ee eee 130
R4(S) oY 0) ol ray ge en ag anagem eek eetiete Pome ete Te ee ony rt eh eyes Poles x indie ae 126
SIGISINAGES Aware WoSie Gi HOChese esas en Se ee oe See eee eee 126
Cannon? (Clarence (regentror the Institution )o-- 25-2222 ee v, 6, 149
@sreys Charles ere ee Re Sia te et ees oe oe ee vii, 24
@anrilcers Vie scl Miri a tae re eee Fe ee ee eee 22
Cartwrcht Ona ses Sean ss meee ee ee eke keno ee eee vi
Canwithen.By i personneltoticen== "9 eee ee ee eee v
Casse diy fe clive sae a ete ee Tas ee ee ee ee ae viii
Chace he Ae aT re es Aa ee ee ce Ray ee ee ee vi
Ghamperiohmelisraes ss ota cece ee ee ae eee eee pn ner 85
Chancellor of the Institution (Fred M. Vinson, Chief Justice of the United
FSET eso) i eccne ee ape rte eerie igo Se agiee Meee yan ee Se ar a ed oe v, 4, 7, 20
hve pind war ee ee eee eee ne tee ee eee eee vi
Chase vAgnageh + ss She = se Me ar ee Fane Se Se ren Se ae vii
Chief Justice of the United States (Fred M. Vinson, Chancellor of the
AVery G16 Gn 61.0 ra) see ee ee v, viii, 4, 7, 20, 25
W@oriste serve etree Oe a ste a 36
@lark vAustingbl. 5.05355. e = le ae ores vi
Clark, Leila KF, brarian.of the Institution. 22. -2-2=2-522-222 ee v, 135
@larkaR Sete otek oe ile eo eye Ie gle ie tee erg een eee ee vi
INDEX 413
Page
Clark, Thomas F., administrative accountant of the Institution________-
Clark, Tom C., Attorney General of the United States (member of the Bn-
SUILUtION) eS os eer = ee ee See S See ale Soe ee wie es ee ve Vv
Clarke a Walter oe etn A eee ee edt SERED A Meare et See 14, 127
@ lair Ked G ila re eI) ene ae ey re oe eS ee ee Fee SE 41
CoalenG correwlhs = saa 4 SLB aks 518 soe ae see ae ee Dae ae ee Fe 69, 70
CochransD oriswMp essa see e ee eens eee ee lel Ee dE vi
Collins Henry Dear ta i OI ee ee nod ee viii, 11, 56, 37
Collins ei lOy Git s=5 94 54-5 29-2 esac aa eh ee bee a ne ve ee Be ER 70
Commerford, L. E., chief, division of publications_--—-----..----------- Vv
Compton Arthurshen(recentiofathemnstitution aa 2 sess] senna v, 6
Compton Karine @hhestate ols cience) masa eee ee ee 395
Cone era lt S a ae ea cg re A hee op fo rt eh a aa ee he vii, 23
Coopers Gustav Ao oS) epee yo eke RE Se Oe eS eee Vil, 2a, oc
Coopereaul licen. eas ase eee TRE eee ea ee ae ee UP, 73, Uy, CU.
Crore 155 18, GReygainie OH Wag line MAUI RON) ee v, 6
Craigias IR aad(oy aio a ee eel ae ee eee See es een eS 60, 61
(Carey eine, Aiice 1D Ses Sea Re Spc neo, Nes es te ee ee ee ee ee ee ee vii
Crmixent wi OsG ete on ee ee tee ne eee ae ea ae eee eee 59
Cunning who Derby si.) Gs ee eae naa ee we eae ee eee eee ae 75
Curator, National Air Museum (Paul E. Garber)__________- IN NAO, UA, ee
Cushman, 1R(0}] 0 Sf et rik peek wee lle oh mre sere repel ance vane Ae nal pe fet et pee Eee
D
TB eK, (Cae Sa piesa kx eth pce i Pag RUN et ects I PF SURRR yg A t Vili, 25, 26
DauchertyppicwandeD) = "ae Seas shee oe to Be See see eee eee 69, 70
Daughters of the American Revolution, National Society of, Report- - --- 141
Davis: ‘Harvey, N. (egentrof the Institution)i22-_—..---2 = --2-2----=--5 v, 6
TOES ich Jeg lB) ete eae a apn a che oer eee ee aN can ee A 118
Deron Gaye oe ae See 2 ee ee enn Se er See 5 See eee Vino
Densmore, IRIE TCC Gere een es ee a ye Soe ea Viii, 58, 88
Director of the National Museum (CA@RemimeatoneWellogg) maaan vi, 24
Donaldson, Jesse M., Postmaster General of the United States (member
of the Institution) _ nS a Se Ge eR eE mee e RD espe e Aa DC RMP SL AS Vv
DON ONO MO lie © Meet see ee este es ea Ne SS ee ee ee he ee eee 79
Dri CKkerae i nilifp ees eee tee ee ee ee re ee ee See ee viii, 67, 68,69
Die eho ee ss" =e kea ees Se eee ee Sheet eas ec see es cal
Ay Talc] Ce) Me ieeete oe mee ee ten ee ee ee a Pe ae ek Se eee vii, 23
Cu CKG GOMEZ ih isha ere eae as Es Se a Se ES Se
E
Partie helompin on ae. hnorntony hace) 2 2222524 2- 40222 2 See eee 161
iastmanss Donal di 225 S87 2 See Se ea ep aaah ere ek reals 65
dgellit George ries ssh Bie eo Ske a ee ee Al
Haditonialadivasiony © hietr@Webster im lGUe) asa a ee v, 142
Beversvangehigeiis.. 52 us aoe heen oe = eee 38
iscnim ann wc oeCn Ces =o es a ee eee Sse ee ee eee 128
Elder, R. A. Jr. Be reser, Uae ee os ole es bes eo he Lae oe eee vi
Elementary ‘particles of physies) Lhes(C@arleD. Anderson) =- 225545222 203
Poise Viaxn Mie a enact Sha Bea” ee ue oe So ee es eee vi
Elstad, NES BS er aoe a aN as ee Nl. ye aa Oh NS RH yn yy EM ee 5 2 ix
Gn siMeers eC OLDS Olen sae set es ten ee ae ne es Nees Sy a ee 61, 62
ramen onaldiSins2s8 2. we ota See eee 4 eS ae ee oe ee vi; 22
Mstablishiment se hhest. 22s) eee. eee a 2 Oe Coe eee 5
Hthnology., bureausof-Americans=225282=— 952 s=a25s 5552 >seenen ee viii, 11, 55
PNUO OU cist eee ey Sen eS ee es ee nee ae Se ee Ae a ee 87
Collectionsss392 2 = aa eee ee ae ae ee ee ee a ee eee 87
Hditonalaworkeandenublications a= eee 12, 86, 141
ihustrationsa: 36 2oae 2222-7 ee eee ee Eee ee eee 86
Institutesot,cocialsAnthropologye = 322242 e225 2222 ee ee 12, 58
ibratyeeeae se See es ee ee ee ee eee Le eee eS 86
Miscellanecousse2- 2222 eure tee oes a ee Ola See ee ee 88
Reporte he See nest He eee Ce ee kee Sees ee eee oe eeaee 55
Rivers basinv Surveys os 26 2 ese eee ee eee | aaa. eee 11, 61
Special-researches!../ See = 323 Sac Seah ee es ae Re ee See 58
Stati. 5. See ee ck Oe ea te viii
414 INDEX
Page
Bttinghausen; Richard... 2253222525 2.55 2 ee ease eee See S viii, 52, 53
Bvans, JGR W222 2a ecm ae en gt Bao ars has apg be: Rens 110
Biwers® Jie C2 ae ie ee pclae ere tat rte a vi
Executive Committee of the Board of Regents___.--_-_._------------- v, 149
Membersec22 = seu eat euieew ce ee cee ee eee oe kee es 2 SS ce ee v, 149
1 =) O10) Pe ee ee ees 143
GANT OTOU © OTL ER GL ODS cet es ney ee 148
aNCSTS{ Ol i ee ee eee Oe ee ee ee ee eee ce eee 147
Y\0T0 bl Renee ane ene eS ee ei rar nee aS 148
Cash balances, receipts, and disbursements during fiscal year 1949_ 146
Classificationvof investments =a. = aa ee ee 145
HreerGallerycorArt fundistet 222 spe as a ee ee 145
GBS cos ee ae ee a etn rade a | 148
MATE MSOMLATAMEIAG Oyyy TUNE Math ULL eee ed 143
SUMMary: OMmendowMents so = ee a 145
Unexpended funds and endowments.___------------==-------- 147
F
Haine hile Davida see ce ae ee eae eee er ace e re eee eee vii
Bie rh GO Tt Val tsa rene NY een ere cae een ee eee eee Vill, 12, 57, 58
client Se ee a8 ee ee Se ee ee ee ee vi
1 EILANG 1 YG1 21S) el lee ec ste lh aye pear emia each 2 eel ey 7, 144
PASTS OPO ELV ET OTS ee ae ee age ee pe eee ere ee U
inimiayaon, Tolan AL, ayevel 1eenel Mls = ee ee ee eee se soeaee 50
Finley, David E., Director, National Gallery of Art___.___--_- viii, 7, 25, 26, 41
RISE T: VV EE Seete cet, Stes mt St Sa ee nn ae eee ats pa ee eee vi
GET e CS °] ap cd fll mts arose nme n Oape e ren an a o p aps Nyel npa d Bos vii
Kleming, Robert’ V. (regent of the Institution) —_ 2-3. --- 25-25 ee v, 6, 149
ontaine SV USselI oes? * Soe Ae ee ee eee Senate eee 127
1 ENOp esByra KS Ah, 9 Meet i pate Ng gt A RC pro RN WRN DUT CS I SEES ey 109
TDCI] ci chp, le {es aie eae a RC My aR ey, Se AT she Na ye aoe EY vii
Foster, G. M., Jr., Director, Institute of Social Anthropology--_---- viii, 12, 59
henalids) ise (Olbnygirs = = = see Se Se: sh rere oe ne AP” OMS BE Ro 4
TAS STS UD) OLO UU ge te we ye tees eee ee ae toe a ae ee 79
HredericksonwD avidehs == eae er te Sas eos ae ee Oe eee eee 64
Hreery Gralll ery: Ot AiG manerg eee re ee ene ee ees viii, 11, 47
PATE GO TA GL ERIN CE epee eae eee re aa ee ee ee ee 150
Chanvestiny exhibitions. ses 2 one en = Se ee ee 50
CWOleChionseeeres Base ss ee a estes ee oe ele eae ae ee a 47
Docent SeLvVICeand Ober sta hee bLvbL eS pee eet 51
FLOM Ona VRCUIGTCS otter ney trea a ee a ee 54
1ST ray ae mers ee, 2a A OR TE I ae ee ek ee ae 50
Publications =e. See 8 oe 3, eee. Se ey eter 7 gh Shey ees 50, 141
Repairs toune;collections 22m s.. oe ee eee eee 49
REDOrE Maes FARR A Se ee ee ee ee 47
Reproductionss!et= 253 osa5a5 [oe SUS eee eda See 50
SpEeclalvisitOrssae 2 S552 en See eee eee ee eee 51
ASIST: dpe enh mcs ee See Ae opel Seed enemy Aye DONG OE EE viii
Briedimann wien bernie esi ten en yen es ee oe fe ay vi
(The breeding habits of the weaverbirds: a study in the biology of
behavior patterns) ance ee ee ae ee ee ee ee 293
G
Crs oss cam ee ahs Dn ee ee. en eae. eS ixey A,
Gardner: Maj) Gen. Grandisom. 3.2.2.2 224.5 525. -6o= 5 oe ee ee rbsey, JENS
Gardnenees Vee: oo ee oe Sey oe ee 8 se eee eel A ee eee aye ee vili, 45
Gazi Mid eB eS Sh a) ta ieee heed Se eee vii, 23
Gazin, srchisabetily hast. ee pe eh Os he Ae eee vi
Gellatiyecollection~2——< =< 25-228 see en Se Bee es Eee u
George, Walter) (regent, of the Institution)—— 22 == 2.22 ae v, 6
Glance: (Grace who. 25 22 ao we ee ee oe eel ee ee oe ee ee ee vi
Goodrichsilloy dace. Vaio aie 2 ee ee eee 41
Graf, John E., Assistant Secretary of the Institution___--_------- v, 6, 41, 129
(Greer yore DS Oe sais ee a ape eee eer eee Ase o vi
Greene aC@harlés Te. 2.88222 ee Se ee oe ee ne ae vi
Ground-water investigations in the United States (A. N. Sayre) --------- 219
Guatemala, The archeological importance of (A. V. Kidder) ____-_-------- 349
Guest, Grace Dunham. - o-oo 22 ee ee koe 2 eee viii
H
Page
HigleoMiar- Genk sWillish) a 92ers Bape oo oe eS ee 55
Hale telescope, The 200-inch, and some problems it may solve (Edwin
1 (RUT 09 5) (23) eee ae at CO ne ln ee, Ee Pee ene er ene ee ee, 175
HET TI BEVO Derby ees ee a ae Soe eA ae) eS eat ey ee 79
Han dley2Charlesi@s ihre 52 ypc pee elo 8 gel (EL pees i Bs Ee 22
aT rine DOme ys) Olah 21 sa ae se 9 Nemes i eee viii, 11, 56
LE (Vert hag] Res al Gade ee A ee ee AN cee Ns Re ete Pyle ON a fe al le 8
TE (Sia Stolt Ao 2 eee A en Rn aye Ae een nee aml mee ere Fa te gt a ix
Elar manera mike At = mee = pwr lant aa eae arene Pot ols A Hekate Aes Ee ee D7
LE [SrA Sip) B90) OTs) a al in ee ee nt eee i ene ei eg See ee ee 69
ElendersOn eer ttt he we See eek es A oN Ee oe ee vii
SiS eal Ores nat cal Dias eae eleaien S clae Oe ee ye SR Re SON ON ee Ee ee Be ape be vii
LE pts Gc) a6 ota Wik s/s eee ge RO pe a Se ee eee eee: 85
TEL GIIG ine gl Ol Deas eR ana Cee ee tee Ce Se eee vii
SE LoU Ke Yes Sand 2 Fiet5] 0) ea a a Pd a ee ee Sees, 85
elo lim berg Alba me tree ta Sete ey toe ee et ee Ee Oe re 60
ETO OVER, Walia tiny ids: soccer oe eat Pe eae eae Se ix, 110
ROUSE kare Ghigo yLOTI Woe a oe tet oh Ce ry eee oh ce vey et 67
ESTO UGE yyste nA ete ae a eg ac enc ee gt ct a ah ear 82
Howard: J.-D). treasurer of the Insmtutione 22292522522 5-2 4 see Vv
LOW aicleg lO) ate ome eRe Pena ern Se a heey ad eee ns IS Se vi
IS LON ell LOR BY VALS) pe ee en ee een ee Tas ee eee vi
Hubble, Edwin (The 200-inch Hale telescope and some problems it may
SO live) ae ee eee ec ee La er ey oe I ce ya Se 175
iaohestaclk vise nt ote Se ha ee eye oe 2 a Phy Le CRUD
eure wel ae E sinks Wars ees oar eo ek ae a a en Se eh th Acre RS 128
Elunsaker, Jerome ©, (regent of the Institution) _--.-..----=--=-s=s=24- v, 6
Els a © Millian ews eae Ne Se Clr es ee tas oe os BS pees 66
I
TEU care? 221 Dias eel SE neh i ee Re A Oe ee ae ee Seems ee. Se oes vi
BEsraal easyer eh pn Ven ee ae 2 ee yee oe Pen ane hth eee BAS 23
Imstiimberom SOclaIvAnthrOpoOlopye. 2-42 == 5 22 a eee eee viii, 12, 58
International Bxchange Services. 2. 9-2. ja. ee viii, 12, 89
Foreign depositories of governmental documents___..-------------- 90
Horerpnuexchange agenciess. =. 2o5 2252 a seco en So ea 95
Interparliamentary exchange of the official journal_____------------ 93
Rackaces received and sente se eau as fossa es ee eae ease 89
RE DORt a: oe 2 tk ana Reet a Re 8D) sane Vv nS SSeS ee 89
SCOPE N Ge ile e steers eee as Me on FE Sa ahd Ge = one A eye meee 79
J
James, Macgill, Assistant Director, National Gallery of Art------------ viii, 25
VellisoneeWianite == awe See se ee we en ne oa sae vi
RFK Tas Coma ag) aa yri lia Leer pets IE PL ETERNAL P S eae So tee vi, 21
Babnconmeliredenickwoate<- (ie 2 ewe het week eer eee ee eS See 75
Johnson, Louis, Secretary of Defense (member of the Institution) - ~~ ----- Vv
rice Nelle ie eee ee 2k ees ee Se ee a Se oe ee vi
K
TROY OYE 0) Jae pel tle GR CN ys Se SAA ep RR Re Ie el Ue i RE vii
KG ya eelcOyee ee Seer een tee Ser es etic Rae oe ee ee ee 79
Keddy: J. is ‘Assistant Secretary of the Institution. 5222_--==-=-2==---- Vv
Kellogg, A. Remington, Director of the National Museum-_------------- vi, 24
Kelloge ‘Charles ES (Modern soil science) 2225_ 2252 52 See 27 Sees 227
TRCTIIOared Bt 0h ape eg el SN EN eae REE Se eC 129
Gell n eI Sae le See eee eth ee 2 erm ar Semen as aug ch en 60
Ge Pel A VAG eer ts ae ee ne ee Wee ais een ae ee 2s eee 35
iestner sabe photographer - eae se Soe eee eo es Meee sere = ee Vv
econ uri Vural bs ee ne Oona ee see Sey ok a Ree eee yan eee ae vili
Keyes Cc harlesmiy- tse tee ec ee ae ae ne mn a a en te See Sem 71
Kidder, A. V. (The archeological importance of Guatemala) _---_-------- 349
RG ip tee eee eee Pe fee te es Ae ee leas Be ee ye vii, 22, 128
5 ERLE AUN Ceca Oe) CI sg eh, a oA IRN PE NE ERNE TSE Ry ES 72, (6, (7
Rerigiit ee Brookes: = ines re ns ke ME aR SL ee eee eho vii
416 INDEX
Page
Kowal Jose phim ba ases = 1 sen ae ele ee SE ae ee ee 127
Kram i 6 Adv re Wy svat oe Sea ee ae gt ee ly in ee I ix
Kress, Samuel H., President, National Gallery of Art____---------- viii, 25, 26
iMriegen gH Wists Se Lee en ot eee a ee ee ee ye eee ee eee vi
ikroeberpAvis cavee=ceewecenheccoen es se hok esha hele sees Se Ree em 58
Krug, Julius A., Secretary of the Interior (member of the Institution) ---- Vv
Kubler (Georses soe Sky se So Soe ea ee eee ee eee 60
Kemi e Var] Orie sea eae ete ae ed a ee ee ix
1a ee O10) al Die: 0) Pee eee ee eee Le nee ee Se Se 120
L
Wtehner wiley ke == seen ae eae Saas = ee mn or meee ote ee ee vi
La Colombiére, Excavations at the prehistoric rock-shelter of (Hallam L.
Movius, Jr.). aA a eRe Snes Se one ee Ree ee oe Sac sal oes eee 359
Landry, (CGN EY Sel oe DE ee ema ORT SLC ea 4
HANG AMC Wane ene a Ae eee La ae ee eee eS See SS eee eee 35
Wane] Os yAte Cire =a ame ts aren = on wk SAS Rs > eS Se ee eA 129
MeamleyeaWi bit Heme tee aaa e eet tenn fe Se oy eer ee ee eee 2S eee vi
Weonard hia Cas «enemas oa: Se senate see RRC eS Pe wees vii
Lewton, PHL acprk dwac we eien de ane ane eh oae aa tha ee 5y cre Ne gee vii
Librarian of the Institution (lellask=Clark) es 2 es eee eee v, 135
ibracy aes neme ek kes ee oe ae ee eh a ee Seer 15, 36, 45, 50, 86, 132
Bureau) of American @ithnolory -----~— 952-2252 alu. 5 Se ee eee 86
BreeriGalleryaor Aarts. occwas s Seah AL. Sete. Oy i Se ee 50
National Collection-of FinevArtse: 2 ==> s-<2s2-5=2522"=5s=2=e ase ee 45
Wational Gallery of-Art;9 222 225= 2=* sie 568 ane SS eee 36
FRC On tia a2 ais iL Mosinee leet Spe agile A en ee ee eee eee eee 132
Smithsonian tes & atte ee te ETA ee A er eee ee See eee 15, 132
iin keAma aes a alse 2 ste A eae ee en ee eee eee viii
AS baphtee pee eee Ane 8 ht See oe ye IN Re oe ee eee 109
hlamoy George Avot ou Seg h ae aE ks Ce eee eee vii, 23
iihoeblich-rAGMto JT 232 22 nes {oS aainin mPa So lter ae iw bee ke SOS Se vii, 23, 82
ihocning* Grover.) .-2s & = seen sod se eh eek ee ees eee ee eee ee EE ee ix, 120
Longnecker, BST aN Vie te a a sty 2 oo ed vie = Nee ee 128
Lowe, Frank O., Head Keeper, National Zoological Park__.----------- ix
Lundelius, Ernest gc irr aa ALSO G AY Ale SRE EE Oe. So) a ee 79
Prindell CC slit — ae eee eee KE Sk oe ok ek ee ee ee Se 23
iran y i Wiss Hieyy 2 se 2 a IU DRO LY SS ee ee no ee eee 129
M
INGA WASNT A Sia! S228" ao Benes 2S Te soli eco oe eee eneere aaa ix
Mann, William N., Director, National Zoological Park___.--------- vi, ix, 108
Manning, CR ESS Rae ee ea ees ee ee ee eee vii
IManshipwrPaul: <2. 24 3 eee ee oe eee ee 7,41
JW We) 6) Pec fee] Bae PR a SI ees SS ee ee a ee ee eon Sona vil
Marshall, George C., Secretary of State (member of the Institution) - - ~~ - V, Vili
Marshall, Wire ene feria Ge as vi
Martin, Grit eee sn eens ie aoe eee ee ee Vili
Mayer-Oakes, Walliams! 20 ein ies nee fA oe SA ee ee 80
McBride, Harry A., Administrator, National Gallery of Art_--.-------- viii, 25
McClure, 1 UR eae ie a el Oe ae. 2 neat ad er AE See i Vii, 23
McConnell, SST Uh Rs ap ee eS een ee en ke eg ny 6
IMeGresor*J chin @sscoseccere eae aeos tae ee eee eee (el
MeiKenzie Gordon PO S22 2eesecned 1 ac lat eee Se eee eR ES eee 79
McQOuiren.* Mr222 20 cercon 3 lL Se Soe Oe ee ee ee 79
Mielion™ Paulie a) 2522 20. ire te. Sy ae bee ee Deo eee ae ee ees Vili, 25, ep
Membersiof the Institutiona222 2242 ee oe ee eee
Menzel) 0 r2< tts 20.20 as sets 2 ee nee ARS SS te Ee eee ee 110
Metcalf (Georve: =. 625.4 -2r coe sata eee: PE Ee Ree ee ee ee (2,40
Willer Caner 2hecs224424 2208s ent ae ae se ee ee ee eee 63
Miller *Gerrit(S = drecrcn et eeeetctreet ea case ee eee eee vi
Miller Mass Gene iuther Di teee2ee wees Sete een eee ote sete el eee eee us
Miller sN; -Miei<4% 22244250 05 actrees ot aae eee ee Se See eee vi
Miller. “Robertititesi8 ius ve SU Ae Fe eer as Oe See eee ae 21
NhitchellMS Ack: tus oes Uae bin doe eee ee ook es dee ee eee eee 9
————_
INDEX 417
Page
LOT G OS AA Chg tel en. ee ee ee ae eee os eee ix, 125
IVI HA AUG eG ca te peaks cee 9 cath Neenah res eG OR Ae eA praeck aA IR Re 64
ING Genie s SELENE sees he at cet Rea ne Ns De pea Le een 128
Migiden keg Mirgs ore Net sere rane ere ee Ot ah eR mala 128
Mion ante blizalbeth epee © ote ot nye tales edna Eee ha 1c Une ait Aen ation Sovo4vco
1 oroy ene [1 622) och cs ee ee PO ee ee ee eee AON me Pee esac ated © 2/0 vi
IMomrisom sy) OSep leks Bp Sie ak es ere re a UE aes i ee ke ee vi
ISU ove wey oecl CaN eG 5 Zit Sle a So cea et ree ee ee Ae eee REE Ee OI SS eT? ¢ vii
Jp OSV GXSs ANTS isl. [2 A A Se ae rel yee eee ee Meee ee ee vii
INOS Sere OL Vir AE peg ee ape eagle Le ag pe ig eg 128
IN (OREN A ONeS Ek Ol ov ha tek) bee ee a a Ss eae ene ee 21
Movius, Hallam L., Jr. (Excavations at the prehistoric rock-shelter of La
Coloma ere ye eres Ree Ee es Ny a Fe ET Sn eM al gi 359
IVI aay Atel eee pe dcen Sea Spe aan pe rte ter al SIE ey sda a Bere patel ae vii, 23
MyersCatherine,. Walden. funds 2200 oo he Se 2a eh po petted ted 10
MiversGeorgetnewitts= 1 22 ee ton a segan ee ee ee ee ee ee 41
N
NiationaleAin Marse tin.) eyt ios 28 See ia ee ae 2 eS ix, 13, 114
IAIC CESSTOINS Mert Sia rae ee tS oe ee kee er ee 122
Acivisony board so6 22" see oe tes see ee ee es See aa ix, 115
Gurston alvactiviticse ss cee ere ee a rere ee eee Leen eure iy renee 117
Jibs gtl] OF BO oT ken pet ie een ae eee ee po en Rel arpa ee ee a Meme = eh pay LBL NZ
Inform atlonsaSenviCes ea yee ee a ae ee eae Be es 120
Museumrsiterand bulldinge= === 2255.0 hoe eS ee 116
NEA a ou Naa ES eR Ce Nice eae Sane eae Ae ep Neo aes me PET our ELL a 116
VESEY CY ONE a ete eee ee dO a ae ote a AE 114
PS IEEEN gee Ses 2 es rpg mts SR Sop I ne sas mao eee meer Ue Send Ly de Nee oy BES 1X
Storage eceean = te kee sent ee eee ae eS Nee tia SEG
SURVeV eee eee ee ee on ayan os eee a eal ee th See 121
National: Collectionmot Mine wArtsi = 2 oes). eee eae easy vile LOD Ast
DEepositsG- see LU SS eee Se Se eee eet eee iter ty ive ye ae 42
Imtormationsenvi Cems = a eee ea eee ee eae 45
Ibilonary Ane ote nate Ras Oe Rone ee oe ee ee re es ee 45
MEOH NHAC CE LCC eee eee eee ee pe Pes ee oe 43
oans;vorothers museums andloreaniza tions mse eee 43
Mier a@atherines Waldens fund]*2 5222) 2. = Jel Ao = eae eee ae eee 42
IBReSCLVa lO nets Bete ee er ees eee ee ee AS eee ee ee 45
Rangeriblenrya Ward tung sss. fen sen es eee eee eee 44
BERT Oe eee ee ei era rly Un (OPIN as SNL, pp 41
SUM MS OMA ATS © OATS STO 1 eee eee ee 41
Specialvexhibitionsesea2 sane ae= sees oe eee s eee Ps eee 46
SU Fame Benet i he RR me MRD ee a ree ae ERR A CF AY a ce 2 VN © aN viii
Wathdrawals iby Ownerss4 2a" 98 eens. St Soe Sor See et Bee 43
NationalaGallenyaofeATt= ao ssa ose == se See ee ee vill, 10; 25
INCCESSIONS eee ae Ree rn ee ae ee SS ee ere oan eee 27
IAC QUMISIETONS re OMIT t; Gee meen ey eres ee ye ee eee ne 26
JN OYOL AO) OV IEE AUON OVS oe a ss ABS ree hg uae Seger 26
INTHE TOYO U2 NOW cle Yo) ceo win et sprees oe papal ee tobe Rela Stier See el See TS TNL? i 10, 27
Audit o1 private tunds.of the: Gallery=.22 222-022 s8e oes =e 40
@arevandimaintenancerorst ier touil cit pee eee 39
(Climmmnnninies O? Gosia GxnnNinenes Ee seed eee esen 39
Gua tonial ea cc ivaGles sete eee ace ea tered eee Rn anyday een ea eee 33
I DfohoKee atop ae yl fob oved tons Ae eee ee es ee ee 36
Exchangesvol wOrks,O1artes so6 5 = eee sea see ea eee eee eee ce 28
HXecubiviercoMmmitteel= == == a Uae aes ee een eee ee ee 25
TEST Nada T ETE © 1S ps eae eee lt oe a meg eek ee ce ep 31
Hin anNcerCOMIMItlee as Seas sas ee ee aa A ea ee eee 36
German paintings. «custody:0f==<1 2.) er 2s eee ee eee 37
Germantsilver,custod yiole ase 22s ee ee ee eee eee
index: ofeAmericanyDesi¢nixna=.-5=5<5 5255555552045 2 se eases lt 8 33, 37
GID TAT Vegan 2 mn a ee i a ee eee ed Recie e mee A ee 3
FoanedaworksoPart returned =. sso ee oeet Oe ie ee sees 29
Newiconstructionis===<22.52=2 25-55 s2-455 5852 2 Jee: eS 38
OCS Stee 5 ee apa a a ae Be Ree es ee ae ale ok Lee AS viii, 25
418 INDEX
National Gallery of Art—Continued Page
Organizations «<.=-5. 255 5225s 522s 5sesrnesssesese sa eee eee 25
Othervactivities.=+—-.-223. Send ae net sae en eee Ae oe 40
Othenciltss* . Varta a a eee BS a ee oe ae a Del ee ee 40
(Psin Gin Gs Sa 2 aa by in a a i oo el at cn A en A aA A PATE
Bhotographs'a a ase Aes a he eR oe me eee ee ae 28
President eiruman sina weurel recep to lessee a ee ee ee 37
Print svanesadrawints:es4 as-s2s2s5eee52822eeenrscSoss eso eee 28
Publicationgac 22 oss ss 4224424 oeeee dob en tas nee snes see AN ee 34
ReDOrhs Sees 32 aa 5 at ale a a le ea ss oo A a Mie ee Rh 25
Restorationyand repair OfawOLks Of/artea== ee ae ee eee 34
Rosenwaldvcollection==22 5523592 h-.2: 522552232222 5) See eee 32
Sculpturckesne suet Stee Ay oS ie ao ee A ee eee 28
ALLAVelIng.OxMIDIGlONS == ==o a aan—r Qos wa ne Aaeee S le eh SS tele ee 32
WMRUStCESSe k= eee ee eee eee eet ee eee cede een Os See ee viii, 25
Wihiteseiouse furniture custo cya of ss ee a eee
Wiorks:oMartdoanedios = 2225 222 Slee Sosa eek Oe SEN Peewee 30
Wiorksiof-anton loantoese. 52a a ak So ween ete ee ee 29
National cosrap bi cis OCle tyme ee eee 9, 11, 21, 55
National Geographic Society-Yale University-Smithsonian Institution
I XpeditiOn® 252.8222 24k once sunes hee ease a eee ase 22
National sViiseume 22 s.. 5 Stee ee ae Soe ie ae eee vi, 9, 16
Changes anOrganization: 25-52 2S n eae ee
WOME CHO TIS ee ee ee a me ee ea eer 16
Bxploration/and: fieldwork: 22" 9a e aos eee ee Oe 21
Publications ee. | se 25 eos ote soo. eee Oe eee eee 23, 138
Re DOR 2 2a ee ae eae es oe one oe ne ae eee eae ee ee 16
SHEE HG ee pis Fe See alee yap Ay SRI IN Ne RENE ERIS AAA eT vi
ING Gro rn ea Pearl S era Oasys es eg ee ee a ay 61, 62
NationaleZoolopical Park! 2.0.02. 20 bee 2 sae ee ae LED. OF
Ha CSN G1 TO ae Rig RL A ae IE RS imine ronda MeN Gps Rk 98
Birthsjand ~hnatehings.—. os 222 22 eee ee 103
Depositors and, donors and, their, gilts. = 22252225505 5— 4 se ee 99
Exhibits). 25s 0s .ostaets soa seg aa ee 97
Meedine theyanimals-\., 5 s2esosons aoe See eae 105
IMATNCGS a5, oe en eo as res = as ns a oe or ee 107
Maintenance) 1d sina O Velie ribs eee ea ee 105
Needs ofthe; 70022252255 2225s a sees oe ee eee ee ee een 107
TERS TOO at ee es cs cece esc a a on 97
Dbalie se oe Sas oe ee ee Sa re at ae ix
Statuspofithescollection-2=5= =.= =. eo. ee eee eee 108
WASItOTS Scene Sh See So ooo oa ee EE eee 106
Ne wan an eV aes So. Re es Sa be oa 2 en tS See ee vi
Newman stanley2.2 2 so oe See ee oe ee a eal ns ead eh es 60
New Zealand, a botanist’s paradise (Egbert H. Walker) ______-_-------- 317
INicolMiDavides2 5 scac= se Soe ae aoe ae es ee See ee vii
O
Obere Ke lery Oe = eee ee ee ee Ae SS eee eee ee 59
Ochser ah aulehistses = Se aes ea SS ee See eee oy a ey ee eee vi
Offictalsyotethemins tit ic ries eee Se ete ee ene ee ee Vv
Oliver, L. L., superintendent of buildings and labor________------------ v
Oliver Ori weaas fees was vole os Te Se SE ene etre noe fy nee ae ore vii
Orr-ikKennethi(G= oe =!s tet ameane ass Soe ae eee Ce ee ee ae 71
Osborne. Homer Douglassa= > 5 =A ers See eS eee Re ee ee 68, 69, 70
Ee
Pacific Science Congress jpevienthe= =~ = 52. ee ee ea ae 22
inagew Lhornton (ihe origin of the earth) -2 5-82 yes. eee eee 161
Ralmer, Medieleno =. 2222-22 o-oo eee eek oe Seeks eee viii
Ralimers NS SS. 2232 38 ke 2 eee Se Se ee oe eee vi
H e1ST Tu c/otea) Th | Leo 8 aa poe ep epee eth SRSA Re Tifa ty 9 Eis ee vii, 23
Pearson, Louise M., administrative assistant to the Secretary_---------- v
Peat. Marwick; Mitchell & Cos... 2-5-2 Scone 2 e522. eae es 149
Rendicton sles sic 2 Seam hae ee Mee on yn i eee ee ee ee vii
IPOER VAG wasn a2 See eee eye ees cw rt pe a ee ee vii
3 es bs t . }
INDEX 419
Page
ECT ero Oe Ete ee ren en care Sa rn a ae oe Res Re ES Be et vii
HEC TSIay OAV eV bea ee tele S9k eats DL ee eee ee bee ye ee 22, 129
ersounel olicer (Bb. --. Canwithen)/ = 22.225 on. sole nee Lee Vv
RECETSOU SR = te a eee ne nes ae age) te ee es ae ae dpe vi
ISLETS OU ey lem yc ett) es eae le ol re Rene RTC Ln 2 palomino ie ee vii
mnilips sy Oincane fee ee a ee ler eae Roe hon AM Dee ae vili, 25, 26
photographer. (lb. ACeStnen) ono: = scm ee oe oe el eee eee a Vv
ICHeLLO ms OLepHen Oss Nee nana: See a ee ee De aes arg 34
PCTSCOD ys LD) ONAL seaate we ote in Gules wen AO ne Ee 59
EMC Cle Les EUG INdiana ec ee ne Lay Sed ete tee api Oe Cheb dete eee gts vii
lnwoods-rlastics: Corp an se ee ele ee Sie ie en ee 22
Pope, John A., Assistant Director, Freer Gallery of Art__ viii, 51, 52, 53, 54
Postmaster General of the United States (Jesse M. Donaldson, member of
hem NS LUGUMGIOU) mers, eee eee Serre) cee eee ds ee ety of pan eee eee Vv
| GLE ehot yal Slee at ees ee er pat ae el PE epee TARE at eM! NLL Fy ix
Peon Reurepe ANd EY ol atoll Oy Rea ee ee eS Sere ie epee Sees 115, 120
Precise time, The determination of (Sir Harold Spencer Jones) __________ 189
President of the United States (Harry 8S. Truman, Presiding Officer ex
OLtCrOro hat ne Blin SU UG UG 1O rn) ae ey me 4
Presiding Officer ex officio of the Institution (Harry S. Truman, President
oOLzthesUmitedistates)/sssr ee ee a ns ee ee eet che ee epte Es 2 Vv
Ce nmiCe w AGL ONG) ten hee Me ee ees ee ee Ee eee 118
EL Conm CeO Mares ketene Se Saco Sere rete Lene 2k pe eee can al ee im ix
BnICe MW oLernousesds COs 20.2 Soeie eae nee oe ee ee ee Le 40
rice wmLveate AGM Ato Vite. 2a. 2 5, hee weats ete lS es eee eee ix
Property, supply, and purchasing officer (Anthony W. Wilding)________ 4
Publications ®2 = = - =" eee eos eee aera Nf eae 14, 23, 34, 50, 61, 86, 136
Nppropriation tor printing and binding=2"_2- 22 2 2 (pe ees 142
Americans ristoricalwASsociatione 2. = ss oan ae ee 141
Bunesucor- American Huanolooye.- eee oo eee ee 86, 140
AONU AIRE DOM eee oe ee ne ein PEL je olan BE onthe 86, 140
Instituteof socialyAnthropologye == =p 22s an hee 61, 86, 140
Daughters of the American Revolution, National Society___________ 141
BAS Elo it ora apeg ears eps See EE ep tp Sa eee apna NE perp tg Nien 136
reer Gallery, Arta 2.0 = SecA es Oh Man Saye PW pa h pert Sey bi pls Be dee. 50, 141
Occasional yRapers se —4 so eek el eee aS oat ae 50, 141
INationalvGallenyioh Att = 25 89- = «sate ee pe oes BAN eal TS eds, 34
Nationa lense Wists fae pie oe lye Ot ee edt Be 23, 138
PACT VESRT IESG POT GAs eee ot ay — sees I ny TE ree oa a ae ey 139
BS ULE Gin epeees eet fe en eee Soe RAD oe Le Leas el ae mS 140
Contributions from the United States National Herbarium__-___ 140
RTO CCE CER GS Sep ds Oe Fe ye eee Oe 0 ee i ee ee ee 139
BERGE POIs tre see See ae NS Sap I oS ee a a eee Oh pe 136
SME RSOMIa nw ames se eee ser eae oe! a eae ho epi re 14, 136
PATE LCD ORU = ses anon ee Se EE Sea pa ee ame oe 137
Miscellancous*C@ollections®s-- =o. 200. 2 ln ae Cae en 136
RS ONE 621] Ux Te A Oper ep RO ca a) NTN 138
Publications. division of, chict (i,k. Commerford)_2 2-22) 2 22a) ae Vv
Q
Quartermaster General; Oflice of. the. 22 25-252 5 ily 1p
18
Radiation and: Organisms. DivisiontOles <5 2" =e ee eee eee IB} IIe
FERVETIA STO CUS bm tees econ ie apace ei en ae ee eee 10
PVECIOMIA CIO MMs UREA EO fe ereiey eae hn rea Boe ae Ann Se eds ea eee 61, 62
Re eC RH a CRE ere as ie A eS 2 eee Oke eee eee 2 eee vii
TRCEXENSI KS C5 i ey J ee ee Pe a SP Rg Uh ew Ae Bet vii
TRRSVeN SV aU TSI af BXOLEN WW.) Eee a yh ee a a v, 6
Xe CUGIVEy CO TIMI CC lee i eee ee eee ee een v, 149
TEU © YO Ob teenie ae oe Arcee oe bei tee od A Ft Senne, SN ae Se eee 143
INTe Tia RG) ane oo cA A as an pe hn ce ee ee ee eye 2 Se v, 6
J EATON CYS) SOU BY Of i ce a ey al A
RC IiGl eee eV Tet CURA ae ie eS re re eo edn DOSE I eee ee ees vi
RichardsCharlesh Misa nse >a se Oe eee 22 wo eee 34, 35
420 INDEX
Page
Richardson Capt; HoldensCs2ece. 4 =. sects = eee een ee on em eee ree 118
RiddellyicranciswAse. jose ee eeten sen eee ees ee een eee eee 64, 69, 70
RiddelibiartysS: Jt 224 s=sS==52 seed e eee en eee ee aa see eee ee , 20
Ripley tose wOillome: «<2 2k 222 eee ected ee see ee eee ed awe ee eee ee 22
IRiVerbasin: SURVEYS -ace=ceee ooee ea en aks ae ee Renee ak eee ee ee viii, 11, 61
Roberts, Frank H. H., Jr., Associate Director, Bureau of American Eth-
nology, and Director, River Basin Surveys_.-.-.----.------- vili, 11, 55, 63
Robinsoniieljdbis~ eee wee ete ees eee ee Sates bees eee ee eee 76
ReOeHline wd OHNWAL i. oo ee oe ee ee eee eee eee 6
OS OTS by Gre a eee ete ee eed eel erbss wk ese eet en ne eee: aE vii
INO Wely NS wAPE == tee ee eee ne eee eee ecko oes Be Eee eee vi
Ronne, Commander Finn (Ronne Antarctic research expedition, 1946-48)_ 369
ROWE) .OlT MEO. 2 LRyROLs oe GENT RSS eee) ub a Ae ee 60
WR Gas (ble eee ee eee ee ee See ee LS vii
ARTTSSE ll cae WO WTASGIN Cle as lp ne ee ee vi
S
Saltonstall, Leverett (regent of the Institution)_.____..___________-____ v, 6
Sawyer, Charles, Secretary of Commerce (member of the Institution) ____ Vv
Sayre, A. N. (Ground- -water investigations in the United States)_________ 219
Schallen Wily oeeestee eee cecece ete tee eee UE ee vii
Schmitt, Wisidosbisn ssa eee eee ae ee eee vi
Scholander, eta gel eel oe ee ee Oe ee ee ene Oe re Eee 14, 127
Schraden Brany. je Sek en eee ee 127
pebraderssalhyMuches*. 3. LOR lat. Pe a eee ee. Cee ee 127
Schultz plieconard Wee ae ee eee ee eee ee ee ee ee vi
Schumacher wh aGecec ose see eee eee ies er) sere vill
SchwarcZ Poenjamins aL eels ee le a a es ee ee See vi
Sciences hestate ot (Karl Cl. Compton) 2.4540 2222) Geese eae 395
Searles iVirs pElancietymicharasOn. os. 2 onton oer ee Re vi
Secretary of Agriculture (Charles F. Brannon, member of the Institution) __ Vv
Secretary of Commerce (Charles' Sawyer, member of the Institution) _____ Mu
Secretary of Defense (Louis Johnson, member of the Institution) it Gees
Secretary of the Institution (Alexander Wetmore) -_-___- NW walt, he (oy, IL, P45}. 128
Secretary of the Interior (Julius A. Krug, member of the Institution) ____ yi
Secretary of Labor (Maurice Tobin, member of the Institution)________-
Secretary of State (George C. Marshall, member of the Institution)_-_ v, viii, 25
Secretary of the Treasury (John W. Snyder, member of the Institution) --
viil, 25, 26
Setzler phrankOMaiy er! Jee. SOTO Tee eed MOL Bie eee vi, 21
Setzerghlenhys Woe acoso eee er re er ee vi, 22
SeyMour Charles. Rsv. see te eee EEE EOE ae ee ee 34, 35
Shalkop, Roberts... -.u cee eeee tee Beet ee eee Set eee 80
Shapley Meri Roce eee ee REESE ee A eS 35
NS eaves MEG Tern aS pepe eee, A pn oe 117
Shepard. Donald WD. 2 eee eee ee ee ee ee ee ee 25
Shih-hsiano-: Wangs aqcuco. == ee ee a ee ee eee 24
Shiner! oe liye Ae 8 Sa ee Sn Se oe eee
Shippeerde Wie kee er ee ee Se SB Rhea Tie ra ae eS Doe ee eee 42, 03; uy
Shoemaker (C) Re 22s 2s ei ee sal ee a eee eee vi
Sirlouis, J. a A a Lae sa Lea tal COIS keira Shee vii
Smith, IN Oia thew rae ban are ee BE De ay Ue ERO Re Coe vii
Smith-"Carlyle'S2s2 225/252 58ers = fates Oe eee ee eee 84
Smith, Danae oe a BE apes Sie el a a ea Oa NI ae ee See vii, 22
Smithsonian Art: Commission... ae 6, 10, 41
Snyder, John W., Secretary of the Treasury (member of the Institution)_ v, viii, 26
Soil science, Modern (CharlesshKelloce) ana se ass saa ae ee 227
Solecki, Ralph PS Fe eR, NP 8 IR PE A SIR 15) 9 De Se eet ane ec ncn Pee Se See 63, 64
Soper, OV CR Tati Cs ona iether Diente inept iocredninier tN Beilin sig 7 > 14, 127
South American Botanical Congress, Second_..........----------=---- 22
Speicher lucene By. 24 6c ce ec eee ee oe eee eee ee ee 7, 41
Spencer Jones) iri arold Lao oe aoe ee Le ee Se te 9
(The determination of precise time) sae Soe ee eee eee 189
Spitzer, uyman, Jr. (The formation of, stars).-2..22-525 2-52 —s4seeeeee= 153
INDEX 421
Page
HED Ry ee ne a SNe ae een meg 7 ne Ce ae eae 2S Gy ae Vv, vi, viii, ix
AStropiny sl cals ODSCTVatO Tye sacs ee ee ee te ix
Bureaujof American) Ethnology 22225285 2) ee ee eet viii
@anal’ZonerBiological Areas] Sos 22 ee ee ee ae a ix
BireermiGallenyromenrG 251 sale A sue also eta Bl el rae CBr ae viii
Intemational Exchange Servicel 222252222222) ee woe ix
Nationale Aine Museums. loess ee ee Foe oe ee ae. Ss ee ix
National Collectionzof Kine: Arts: 2225 2) 5 225. - sae es eee viii
NationaliGallerycot: Artis oe 2. 2 sins wees bike © eg a pb te See Rae vili
Nationale Viusetimiss aoe 2 eek. me Meme eee a pee ee eee vi
National ZoolosicaloPark ase. 2. 20m eu dso eae ee as eee ix
RSKES CST GT OT eben IO Sa Se ea eta Rng te eA ee vege SS en ey Pi rs vi
(SUD OYSY OY 00 E79 shay ea ae tenia Ural nce oh te ee ern Renae Gey a er A Oe | Vv
Stanley, Wendell M. (Recent advances in virus research)_____________- 213
DGATGOM glen Wis OES! ee eens Siete Coy Vale Sethe a ie dake meee eed ty ak me vii
POLAT KO EC OWC Tob rete she aM Sips An 9d ONG ee yeah he Dp pps ee 82
Stars, ane formation of Gayman Spitzer) dirs) =a ee eee ee 153
SLelansson ss Valllnj alma eee rey rape i eee eal pees Seen ee. Seles 56
Prephenson. IODSri Mime = ss eel oe oe ee 82, 83
SEC VENSOMe JONMVAw wn 2 ees = ks Se oe ee a ee ee vii
PUG WW ALTU SECC MAT Cys os ae 2 cue es ee es a Satin a ae 128
Diewath nM DAle eee. foo ee ME ee ee ete en vi, 22
Stirling, Matthew W., Director, Bureau of American Ethnology__________ viii,
11, 55, 88, 128
SrTlingvincssVintbaeweW = sb seer = ees ss Sak eee hs Senet ws ee ee 128
Shouew ViaTsnelley > 2256 ae oe 6 ie eo re ed Ge ee 128
SHOU SMIVALUUTeNTIN ES) ise ea ee oe ee ie oi a ae ix
SbEOWC MeLVOWCTUS Cm oe ee eee ee eee Seen eee Ea ix, 121, 122
SGUDDS UTM SwAtse fe ese ei Se Soe See lea oe eee. ee oe 53
Summary of the year’s activities of the branches of the Institution________ 9
Superintendent of buildings and labor (L. L. Oliver) ___-______________- Vv
RON He rae AS OMe heen oe Me ae eS eee 2 ep ee vii, 23
SM AULOMMOOUMEL eRe ea tani ns eee ea = ee ee Sal viii, 88
SSN WVEUGUIEAN ES AY 6 ee 2 a et eg ee ee ee ey 20 vii
SAS eC RAISE ae ee ea ee en een eee eee See ie 8 vii
T
lea yer tg) ein Cree ar pe ee Ea ee Oot eh Pr ae ix
“ACI Ne Wee 1 yee £7 pt le ae Va i Pe erent Tie UE af Nyro Mee, SO Me ee vii
MRA OT MMV US TVA Oo) ey tet ae saa ee ae Ne, AS 2 7
MER VIOP WE ol k= Soe eee eee Se ee eo es ei Ee vi
ghhackernC OliN= soe cetes. oe aol ee ek ee Bee nie os oe eee 56
PN OMAS MISS Bhi sao as RE oes hoes aaa cane a eee Se 5 lee 129
PRO MIR S Cre ON Se eee a keer eet ie St ee Ceres oS a vii
Time, The determination of precise (Sir Harold Spencer Jones)_________- 189
“Mime ain Gxrollvypvon (Ch, 1d, VCUIMER) eee ea cence oc ee enece secede 247
Tobin, Maurice, Secretary of Labor (member of the Institution)_________ Vv
MOUuTLCLOUPEIATTY GAs 22 Soe sao hee oe aes eR Sa tse apr ae a oe ee @
(MRS UR Ot Wa lbaALATAKOIN jy IDL Jako ewe) 2 v
shrucmWebstert.. Chich editorial, Divisions = ===s=2-56 42 = - = 2 eae v, 142
Truman, Harry 8., President of the United States (Presiding Officer ex
Oi c1OROfes chem ln sStiGutlon) i ar ee a aerle ce nee eee eee Vv
“MaKe fepes INI Ne ies eo ee eS oS Se eS ae ee ae eee ea SeS viil
200-inch Hale telescope, The, and some problems it may solve (Edwin
TB ADNG) 6) Le) ley eu pa PO AAS SY roars mane co oti aR MT ie Pe on SRE Cry Pts em 175
PTGS MERC AINE = ape wince ct ah aoe a hts Ae SI I sd ee ena eee 118
V
NYEvayo Kes ov ayes fee yee © (ey aleg dc laps cs Dem A aa es ra a a ee ne ees eee 115
NUE TNES OV 0d IN (oe oe aan NE ei a ga pe on eS eee eed ae Sees eka a vii
Vice President of the United States (Alben W. Barkley, member of the
AD TES 6:5 GAL GY ny) ee a ee ee are te ote ners ee v, 4, 6, 20
Viking Fund
422 INDEX
Page
Vinson, Fred M., Chief Justice of the United States (Chancellor of the
MMS GG WGI OT) ee os ee ee re ee Vv, vill, 4,7, 20) 25
Virus research, Recent advances in (Wendell M. Stanley) _-_--_-----_--_- 213
MicHOISS. oe ips on eae eee ee eee eee EE ee See eee 8, 11, 27, 50, 51, 106
Breer Gallery, of Artss a. 52 ote ct pecan scek ene eae le eee 1S 50, 51
National iGallery: of Arte. 222 oc SN ope i Sa i eRe ea ee viii, 12, 58, 59
Williams, D. G., Chief, International Exchange Service__-_---------_-- viii, 96
Walsontaliesterstn vs 2st e ee ee een eee eee eee ee 82
Vii S Oa MeV hn NAG ASS rs Ale fe ae 2 ne Se st ek eo I Re ee ie ee eer vi
WaitierstpAmnol divi = =sta= aan — ee we dee eh ee ee ea ee ee ee eee ee 66
RVIGhTOW eM iee ieee nk Ss Aes ele A ee ee ak ee ae Se eee ix, 112
NWMitiiyAaV Vane nel, = ee eee a eee 1 ee eee ee ee ee 19
VOT yi OTN Cae aes eee ee am ee oe ee ree ee ee 51
VV. Oc ire Meal cree ete SRE eee ren er tee ere ere ee 67
WiOOdShsltvi ate Ee ee een ne oe ee a ea ee Tee SS eens tee ee mee vi
LNVerbed og IN AD COS ON vette ye Gk lM SR a NSN DE ES I al op Ly pis al AN ed Be 3, 4, 20
Wiriehtesrothers;seroplanciof 9038 =2—e2== = ==2 eee ees nen ee 3, 20, 114) 117
Presentation of, to the United States National Museum__-__----_~-- 3, 20, 118
a6
RA1eNUNIVeTsiti ae eee ck eee ee ek ee ee ee ee ee eee 22,
BYU Cl Gan Creuse Lie eee ene. ME Ee ee eet ee ee ee ee 65
Z
AAC OWE TO LOR Ae She AS ne tt er eee cs A at eg ne 70
Zetek, James, Resident Manager, Canal Zone Biological Area____------- reg, ssl
ETE Tee pap Hse (CHET TAO VO LUT OT) ees eee eee ee 247
, PP ai, Pema’: , (eles iN rcs Rigas
Det te anata PM se leh He’ vce alec tN PRR Mh
So ruchrindiay: lio: ™ a) Cee ae BO wee WINE Were!
Pitti ait Ce | Anan i my dpe a OTL
i im
‘piv de onion Wi Leslie Wis dikecd lise athens RGR AMANO WAN: ee hit) ta tn epee
Thien i | ; ; ; i ie £ ely tae yc om i)
, 6 dee Sil Ain piel " ‘eelatiog: CRE a
Oh BAH PAR i hs hie, ue Gia AS ceverie, Mada tts ah’ ae phe oe RI ue
Mie ere Vie ba et cw a ee net her Be cae Leer em n Ty ye)
ne MIT Y hai h Mint aay ig ) aD Mae Daa’ Ae a j ey ee eh.
Core wig. ae Ld eS i by Pe Ae ; nea ie eee ah:
ran 7“ ev tre Ne ae Lay van ae CARD en
Bee « “t Peon |
iP seoontam id Ty bay COA ee een, ea Aten Tae a wien ae
| ; ye me eid Rr
aa ee pias ae Fhe BO ae
ihe ree Om ae oe mae a ae a both ah
, te hig SN ert, Mae Ba WS. ih,
oe Arana pean ihe, Hee. a oe (gala) Apt oi
hte Pebehansh ae ere | be foi
Aad eens by. WAASE IY td ey Lil, vane Weyl) i” }
Weis Lats | cae iad idle aac
a Hi eres a anes ee ie to ay a an + ari r ir ‘ : % a |
Nise Ae fa ty re A re Lh ibaa Ph dies: saga de ce
Lae ih i nob » Paina tt! re Hist!” (EGO Bl + | ; o& A :
at ea i ¢
Ch eee f |
POLE if ee
bat a 3 ie
He, bs i
pith, <, 5
; am
ii nl ae eae ea ete | rae Te ie
Lis Biase seth waite Mave yen | eh:
edie ent et Wr Leet Paha TR y ih we er pind pen (te
re lsat i Ys my uit . ihe, P ‘A "9 se oe x q.. a Var nae hendle +. re) ot
knits ail Nilay EN ethiseeah hegen Ue allan hak WD sie wah a Cc eg a -