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THE ATOR: :
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23 AUG 2002
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Botany Series “=

S)2

NATURAL
HISTORY
MUSEUM

VOLUME 32 NUMBER 1 27 JUNE 2002

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World list abbreviation: Bull. nat. Hist. Mus. Lond. (Bot.)

ISSN 0968-0446

The Natural History Museum Botany Series
Cromwell Road Vol. 32, No. 1, pp. 1-59
London SW7 SBD Issued 27 June 2002

Typeset by Ann Buchan (Typesetters), Middlesex
Printed in Great Britain by Henry Ling Ltd, at the Dorset Press, Dorchester, Dorset

AX CBG LEYS) 2

Bull. nat. Hist. Mus. Lond. (Bot.) 32(1): 1—5 Issued 27 June 2002

Taxonomic notes on some African species i
the family Calymperaceae (Musci)

Ditiinsitien te a

THE NATURAL
HISTORY MUSEU!

H 90n
25AGE2802
PRESENTED

GENERAL UBRAR

i a

LEN T. ELLIS
Department of Botany, The Natural History Museum, Cromwell Road, London SW7 5BD

SYNOPSIS. Syrrhopodon usambaricus Broth. ex S. Orban is placed in synonymy with Syrrhopodon asper Mitt., and
Syrrhopodon lisowskii S. Orban in synonymy with Syrrhopodon gardneri (Hook.) Schwagr. The distinctive features of
Syrrhopodon stuhlmannii Broth. are discussed, and the only records of Calymperes moluccense Schwagr. from Africa are

redetermined as Calymperes palisotii Schwagr.

The research for this paper was largely undertaken in response to
difficulties encountered in identifying the specimens of Syrrhopodon
collected during the British Bryological Society Expedition to
Mulanje Mountain, Malawi, 1991. Determination of this material
would have been considerably more difficult without the important
primary accounts of the African species of Syrrhopodon by Orban
(1981) and Orban & Reese (1986).

Syrrhopodon asper Mitt. in J. Linn. Soc. Bot. 7: 151: 1863. Type:
Tanzania, Kilimanjaro, Hannington s.n. (NY!-holotype, BM!-
isotype).

Fis. 4.

Syrrhopodon usambaricus Broth. ex S. Orban in Acta Bot. Hung. 24:
113 (1978), syn. nov. Type: Tanzania, Usambara, Lutindi, 1902,
Liebusch s.n. (H-BR!-holotype).

DISCUSSION. The leaves in Syrrhopodon asper Mitt. consist of a
linear-lanceolate chlorophyllose limb extending from a subelliptical
hyaline base. They possess a prominent marginal rib and are spinulose
to various degrees. As in many species of Calymperes and
Syrrhopodon, the leaves in different specimens can vary widely in
their relative dimensions (Fig. la—c), and range from a stubby 4 mm
to aslender 7.5 mm long. The marginal ribs, viewed in cross-section,
are well differentiated. Commonly, a superficial layer of
chlorophyllose cells encloses small dorsal and ventral groups of
stereids that are separated by a median row of guide cells (a costa-
like arrangement of tissues). This arrangement of cells in the marginal
rib is plainly developed in the leaves of some specimens and less
well developed in others (Fig. le—h). Towards the base in all leaves,
the marginal rib becomes a flattened, undifferentiated, often
unistratose band of linear cells. The region in the leaf base in which
the margin transforms from a differentiated rib to an undifferentiated
band also varies between specimens.

The type specimen of Syrrhopodon usambaricus Broth. ex S.
Orban (Liebusch s.n., H-BR) represents a form of S. asper Mitt.
with tall, slender shoots. The leaves are relatively fine and nar-
row, and hardly curl when dry. The marginal ribs are well
differentiated, and in the leaf base, the transition from polystratose
rib to undifferentiated band occurs well below the apex of the
hyaline lamina (Fig. li). In the isotype material of S. asper
(Hannington s.n., BM) the shoots are small and have shorter,
stubbier leaves. These curl when dry, are notably spinulose, and
have prominent, well-differentiated marginal ribs. The region
along the leaf at which the margin transforms from differentiated
polystratose rib to unistratose/bistratose band tends to be adjacent
to the apex of the hyaline lamina (Fig Ik).

© The Natural History Museum, 2002

Intermediate expressions of these contrasting features of
Hannington s.n. and Liebusch s.n. are apparent in the great range of
material now available for examination, and indicate that these
superficially distinct type specimens represent extreme forms of the
same species. For example, Wood 1726 (Tanzania, BM, BM-K) has
leaves proportioned like those of the type of Syrrhopodon asper that
become curled when dry, but some cross-sections through the distal
hyaline leaf base show a degree of differentiation closer to that in the
type of S. usambaricus (Fig. 1)).

SPECIMENS EXAMINED. Malawi. Mulanje Mountain, June 1991:
Hodgetts 2041a, 2047c, 2220b, 2532a, 2669a (RNG!); Kathumba
MS5915a (RNG!), M5916 (BM!); Kungu M3123a (RNG!); Longton
M8054a, M8058a, 8425b (RNG!); Magombo M4041b, 4042b
(RNG!); Porley 35a, 278a (RNG!); Russell M6055b, M6063a,
M6068a (RNG!); Wigginton M1034a, M1682a (RNG!), M1201a
(BM!). Uganda. Kadese, Ruwenzori Mts, above Miniba camp,
2700 m, 22 January 1962, Loveridge JPL397 (BM!). Kenya. Mutha
Hill, August 1938, Boy Joana 7519 (BM!, BM-K!). Tanzania.
Usambara, Lutindi, 1911, Liebusch s.n. (H-BR!); Usambara Ouset,
créte Matundsi-Mashindei, SE of Ambangudu Tea Estate, 1300 m, 5
February 1985, Pécs 8533/R (BM!); Kilimanjaro: above Marangu,
2000 m, 13 July 1948, Hedberg 1144e (BM-K!); on path between
Marangu and Bismark Hut, 2400 m, 24 February 1953, Wood 1726
(BM!, BM-K!). Morogoro District: Nguru Mts, ridge behind
Dikurura Valley, 1700-1900 m, 6°02'S 37°32'E, Pécs 89119/W
(BM!); Nguru ya Ndege Hill NNW of Morogoro town, summit,
1200-1350 m, 6° 42’S 37° 36’E, Pocs & Knox 88252/H (BM!).
Mozambique. Namiuli, Makua Country, 1887, Last s.n. (BM-K!).

Syrrhopodon stuhlmannii Broth. Bot. Jahrb. Syst. 24: 240 (1897).
Type. Tanzania, Uluguru, Bergwald, 1600 m, Stuhlmann 8809
(BM!-isotype).

Fig. 2a—d.

DISCUSSION. In Orban & Reese (1986) Syrrhopodon usambaricus
Broth. ex S. Orban [=Syrrhopodon asper Mitt.] is keyed out beside
S. stuhlmannii Broth. Large specimens of S. asper are superficially
similar to those of S. stuhlmannii. Both species possess leaves with
marginal ribs that have a costa-like structure (viewed in cross-
section). However, the species are easily distinguished. In leaves of
S. stuhlmannii the marginal ribs are mostly smooth; the cells of the
chlorophyllose lamina are ventrally roundly protuberant, and dorsally
flat to barely protuberant (Fig. 2a—d). The rib at the margin of the
distal hyaline lamina is very well developed and strongly differenti-
ated, with a median row of guide cells often more than nine cells
wide (Fig. 2c, d). In contrast, most superficial cells of the marginal

2 L.T. ELLIS

Fig. 1 a—l. Syrrhopodon asper Mitt. a—c: leaves; d-I: cross-sections of leaf through (d: chlorophyllose lamina, e—h: margin of chlorophyllose lamina, i-l:
margin around distal hyaline lamina. a Drawn from Pécs 8533R (BM). b Drawn from Pécs & Knox 88252/H (BM), c Drawn from Boy Joana 7519
(BM). d, g, i, Drawn from Liebusch s.n. (H-BR, holotype of Syrrhopodon usambaricus). e, ; Drawn from Wood 1726 (BM). f, k Drawn from Hannington
s.n. (BM, isotype of Syrrhopodon asper). h, 1 Drawn from Pécs 89119/W (BM).

TAXONOMIC NOTES ON CALYMPERACEAE

sae OR a

a-d, 1, J

Ra vedelelenan

J )

50 um

0.5 mm

Fig. 2 a—d. Syrrhopodon stuhlmannii Broth. a—d: cross-sections through leaf margins (a, b: in chlorophyllose limb, c, d: in distal hyaline base). e, f:
Syrrhopodon gardneri (Hook.) Schwagr. e, f: gemmae on leaves (e: in cluster at apex (ventral surface), f: in groups in mid-leaf). g—j. Calymperes
palisotii Schwagr. g, h: apices of gemmiferous leaves (g: in ventral view showing gemmae-producing region (gemmae lost), h: in dorsal view); 1, j: cross-
sections of chlorophyllose limb (i: showing half of costa, and j: lamina). a, c Drawn from Pécs & Knox 89053/AK (BM). b, d Drawn from Stuhlmann
8809 (BM, isotype of Syrrhopodon stuhlmannii). e Drawn from Kathumba M5073b (RNG). f Drawn from Longton M8375a (RNG). g, h, i, } Drawn

from Lisowski 50255 (EGR).

ribs and the cells of the chlorophyllose lamina in S. asper are replete
with acute projections (Fig 1d—l). If apparent at all, the median row
of guide cells in the rib at the margin of the distal hyaline lamina is
usually less than six cells wide (Fig. 1i-l).

Syrrhopodon stuhlmanni Broth. remains a distinct species,
apparently endemic to Tanzania.

SPECIMENS EXAMINED. ‘Tanzania. Uluguru, Bergwald, 1600 m,
Stuhlmann 8809 (BM!-isotype of S. stuhlmannii). Uluguru Mountains,
30 July 1941, Eccles AH8648 (BM!), AH8652 (BM!). Morogoro
District. Nguru Mts, ridge above ‘Spirit Lake’ at the north source of

Chazi River above Chazi Falls, 2000—2100 m, 6°00’S 37°30'E, 4 Feb-
ruary 1989, Pécs & Knox 89053/AK (BM!); Lushoto District, West
Usambara Mts, 5 km east of Mgwashi village on west slope of Gonja
Hill, 1600-1700 m, 4° 47'S 38° 33 'E, Pécs & Krog 88205/R (BM!).

Syrrhopodon gardneri (Hook.) Schwagr., Sp. musc. frond. suppl. 2
(1): 110 (1824).
Fig. 2e, £,.3,

Calymperes gardneri Hook. Musci Exot. 2: 146 (1819). Type:
Nepal, Gardner [1205] (BM!-holotype, BM!-isotype).

<SecaNOO8. POE. 1; Beh
seectna L aaladoss «| Shara
: , » SER

a, b c-f

L.T. ELLIS

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1mm

Fig. 3

Syrrhopodon lisowskii S. Orban in Egri Ho Si Minh Tandrképz6
Fois. Fiizetei 18: 81 (1987), syn. nov. Type: Zaire, Haut Shaba,
Env. de Kasumbalesa, Colline Kibwe I., 1400 m, 20 March 1971,
Lisowski s.n. (EGR!-holotype).

DISCUSSION. The holotype of Syrrhopodon lisowskii S. Orban
possesses narrow, gemma-bearing leaves that have strongly incurved
margins. Orban & Reese (1986) use these features to distinguish S.
lisowskii from S. gardneri (Hook.) Schwagr. However, cross-sec-
tions of the leaves from the types of S. gardneri and S. lisowskii are
indistinguishable (Fig. 3-1), and an examination of a large number of
specimens has shown that leaves with incurved margins and/or
bearing gemmae fit comfortably within the range of features
characteristic of S. gardneri.

In the latter species, young leaves often tend to possess strongly
incurved margins and can have blunt, almost cucullate apices. The
isotype material of Syrrhopodon gardneri (BM) has many leaves
with a lamina approaching the degree of incurvature apparent in S.
lisowskii (Fig. 3e, f). Gemmae occur in many specimens of S.
gardneri and their presence is not correlated with the relative
breadth of the leaf or incurvature of the leaf margins. They may be

0.5 mm

50 um

aI. Syrrhopodon gardneri (Hook.) Schwigr. a—b: leaves; c-f: details of leaf apex, (c, d: in lateral view, e, f: in ventral view); g—I: cross-sections
through leaf margin (g, h: in chlorophyllose limb, i, j: around apex of hyaline base, k, 1: in distal hyaline base). a, c, f, g, i, kK Drawn from Gardner H1205
(BM, isotype of Syrrhopodon gardneri). b, d, e, h, j, 1 Drawn from Lisowski s.n.CEGR, holotype of Syrrhopodon lisowskii).

produced on the ventral and sometimes dorsal surfaces of the costa,
often in a bunch at the leaf apex (Fig. 2e). In a collection from
Malawi (Longton M8375a) gemmae occur in small groups at inter-
vals along the costa, associated with the loosely transverse rows of
prominent costal teeth. The leaves of this specimen are not espe-
cially narrow nor are the margins of the chlorophyllose lamina
particularly incurved (Fig. 2f). In all respects it is identifiable as S.
gardneri.

SPECIMENS EXAMINED. Malawi. Mulanje Mountain, June 1991:
N. Hodgetts M2008e, M2026d, M2048b, M2066a, M2067b, 2383a,
2555a, M2638a, 2661a, 2662c (RNG!); Kathumba M5058(BM!),
M5073b(RNG!); Kungu M3241 (BM!), M3267c, M3268a, M3033
(RNG!); Longton M8231a, M8232a, M8375a, M8648a (RNG!);
Magombo M4238a (RNG!); O’Shea 7107b, 7108d, 7110a, 7111b,
7138a, 240a, 7240b pro parte, 7241b, M7404c, M7406b, M7508a,
M7509b (RNG!), 7318a, 7389a (BM!); Porley 98a, 232a, 317a,
329a (RNG!); Wigginton M1009b pro parte, M1024b pro parte,
M1217a, M1219a, M1225a, M1243a, M1247a, M1417a, M1576a
(RNG!). Zomba Plateau, forestry campsite opposite Chawe School,
1470 m, 8 August 1993, Stevenson s.n. (BM!). Uganda. Ishasha

TAXONOMIC NOTES ON CALYMPERACEAE

Gorge, Kanunga, 7500 ft, 6 June 1952, Lind 14 (BM!); Bwindi
National Park, forest 1 km east of the Zaire border near Rukubira,
1680 m, 0°59’53"S 29°35’50"E, 3 February 1996, Matcham U1118a
(BM!). Democratic Republic of Congo. Lushiji, October 1923,
Overlaet 347 (BM!). Angola. Huilla District, near Humpata, 3800—
5500 ft, May 1860, Welwitsch 6 (BM!- isotype of Calymperes
welwitschii Dub.). Madagascar. Ankafana, 1880, Deans Cowan
s.n. (BM!).

Calymperes palisotii Schwagr. Sp. musc. frond. suppl. 1 (2): 334
(1816). Type: Nigeria, ‘Oware’ [Warri], Palisot de Beauvois s.n.
(BM!- lectotype).

Fig. 2g-j.

DISCUSSION. Orban (1995) identified three specimens from conti-
nental Africa as Calymperes palisotii ssp. moluccense (Schwagt.)
M. Menzel in M. Menzel & Schultze-Motel [=Calymperes
moluccense Schwagr.]. These specimens, collected by Lisowski
from Zaire and Guinea, would have been the first African records for
C. moluccense. Unfortunately, all three collections are Calymperes
palisotii Schwagr. (non C. moluccense). The cells of the
chlorophyllose lamina are ventrally roundly protuberant (Fig. 21, j)
and the lamina at the apex of gemmiferous leaves forms a broad
point (Fig. 2g, h). These features are typical of C. palisotii, and are
especially well represented in collections from Africa. As explained
by Ellis (1987) and Ellis & Tan (1999), C. moluccense (syn. C.
palisotii ssp. moluccense) is distinguishable from C. palisotii by its

R)

possession of laminal chlorocysts that are ventrally drawn out as
acute, often coronate-papillose projections, and gemmiferous leaves
with a rounded rather than broadly pointed apex.

Calymperes moluccense Schwagr. has yet to be found in conti-
nental Africa or the adjacent islands of the Indian Ocean, and must
still be regarded as an Indo-Pacific species.

SPECIMENS EXAMINED. Guinea. Conakry, Campagnie Miniére,
23 December 1961, Lisowski 918 (EGR!). Democratic Republic of
Congo. Haut-Zaire, Kisangani: prés de la Porte, 11 December 1977,
Lisowski 50255 (EGR!); centre de la ville, 22 December 1977,
Lisowski 50379 (EGR!).

ACKNOWLEDGEMENTS. | am grateful to the directors and staff of EGR
(especially Dr S. Orban), NY and RNG for the loan of specimens.

REFERENCES

Ellis, L.T. 1987. Taxonomic notes on Calymperes. Journal of Bryology 14: 681-690.
—— & Tan, B.C. 1999. The moss family Calymperaceae (Musci) in the Philippines.
Bulletin of The Natural History Museum, London (Botany Series) 29 (1): 146.
Orban, S. 1981. Studies on African Calymperaceae, III. Conspectus of the African
species of Syrrhopodon Schwaegr. Acta Botanica Hungarica 27 (1-2): 169-177.
1995. Studies on African Calymperaceae, VI. New data to continental Africa and

Madagascar. Acta Botanica Hungarica 39 (3-4): 227-234.
& Reese W.D. 1986. Notes on the taxonomy of African Syrrhopodon. Abstracta
Botanica 10: 349-255.

PRETNG Sen e

Bull. nat. Hist. Mus. Lond. (Bot.) 32(1): 7-11

6 ge

Issued 27 June 2002

Typification of Ranunculus bullatus L.

(Ranunculaceae)

RICCARDO M. BALDINI

Museo Botanico, Universita degli Studi di Firenze, via G. La Pira 4, I-50121 Firenze, Italy; rbaldo@ unifi.it

CHARLES E. JARVIS

Department of Botany, The Natural History Museum, Cromwell Road, London SW7 SBD; c.jarvis@nhm.ac.uk

SYNOPSIS. The typification of Ranunculus bullatus L. is discussed and a lectotype and epitype chosen which maintain usage
of the name in the sense of subsp. bullatus. Ranunculus bullatus subsp. cytheraeus (Halacsy) Vierh. is recognized at species rank,
and the combination R. cytheraeus (Halacsy) Baldini is made to accommodate it. Both taxa are described and illustrated. The
names R. bullatus L. var. cuneifolius Coustur. & Gand., R. bullatus L. var. cyrenaicus Pamp., R. bullatus L. var. ovatus Freyn, R.
bullatus L. var. prolifer Gennari, R. semicalvus Jord. & Fourr. and Jonosmanthus plantagineus Jord. & Fourr. are also typified.

INTRODUCTION

Ranunculus bullatus L., s. l., is a polymorphic taxon distributed in
the Mediterranean area (Greuter et al., 1989; Jalas & Suominen,
1989) by scattered populations from Portugal to Greece and North-
ern Africa (Maire, 1964). It is well-known that there are many
difficulties in distinguishing the several infraspecific taxa described
within this species (Gandoger, 1889; Vierhapper & Reichinger fil.,
1935; Davis, 1965) because many diagnostic characters overlap. As
a part of a wider study of this group of taxa in Europe (Baldini, in
prep.), it is necessary to establish the correct application of the
Linnaean name.

LECTOTYPIFICATION OF RANUNCULUS
BULLATUS L.

Linnaeus (1753: 550) used a diagnosis ‘Ranunculus foliis ovatis
serratis, scapo nudo unifloro’ cited from his earlier Hortus
Cliffortianus (Linnaeus, 1738) along with three synonyms from,
respectively, Bauhin (1623), Dodoens (1583) and Clusius (1601).
He also recognized one unnamed variety (var. 8) based on a series of
polynomials of Morison (1680) ‘8. Ranunculus latifolius bullatus
autumnalis, flore pleno & prolifero. Moris. hist. 2. p. 447. s. 4. t. 31.
f. 49. 50. 51.’. Finally, Linnaeus added ‘Habitat in Lusitania, Creta’ ,
and a symbol to indicate a perennial life form (Fig. 1).

Although there are two sheets referred to this taxon in Linnaeus’
Herbarium at LINN (numbering follows Savage, 1945), 715.10
lacks the relevant Species Plantarum number (i.e. ‘8”) and is there-
fore a post-1753 addition to the herbarium (see Turland & Jarvis,
1997: 458-459), and 715.11 is entirely unannotated. We therefore
do not accept either sheet as original material for this name, although
both are identified (e.g. by Davis, 1965: 179) as belonging to the
typical subspecies. There is no relevant material in any of the other
‘general’ Linnaean collections (e.g. H, MW, S, SBT, UPS), nor is
there any in the Burser Herbarium (UPS) which can be associated
with the Bauhin polynomial cited by Linnaeus. However, there is a
specimen in the Clifford Herbarium (Herb. Clifford: 229, Ranuncu-
lus 7, BM! (viewable at http://www.nhm.ac.uk/botany/databases/
clifford)) which can be associated with both var. B of Hortus
Cliffortianus (Linnaeus, 1738) and var. B of Species Plantarum

© The Natural History Museum, 2002

(Linnaeus, 1753) and is a double-flowered form (‘flore pleno &
prolifero’) of R.. bullatus. It is similar to the plant portrayed by
Cornut (1635: t. 95; cited by Linnaeus as a synonym in 1738 but
omitted in 1753) and Morison (1680: s. 4, t. 31, f. 50). The Clifford
specimen is original material for the name.

The other original elements for the name are drawn from the cited
and uncited illustrations. The woodcut of Dodoens’ (1583: 426)
‘Ranunculus Lusitanicus’ was copied by Morison (1680: s. 4, t. 31,
f.51,as ‘Ranunculus latifolius autumnalis caule hirsuto flore minimo,
nobis’) via the uncited Clusius (1601: 238, as ‘Ranuncul. grumosa
rad. I. species IT) and represents a plant from Crete (notwithstand-
ing Dodoens’ belief that the plant came from Portugal) identifiable
as R. bullatus subsp. cytheraeus (Hal.) Vierh. (see Greuter et al.,
1989: 428).

Clusius’ (1601: 238) ‘Ranunculus grumosa radice I.’ , also copied
by Morison (1680: s. 4, t. 31, f. 49, as ‘Ranunculus latifolius
autumnalis fl. simplici, nobis’), represents a plant from the Iberian
Peninsula, identifiable as R. bullatus L. subsp. bullatus.

Lastly, for his unnamed variety B, Linnaeus cited Morison’s
(1680: s. 4, t. 31, f. 49, 50 and 51) along with a single, composite
polynomial that Linnaeus derived from those used by Morison for
his three illustrations (for the identities of these, see above).

There are therefore six possible elements from which a lectotype
could be chosen: the Clifford material, the element represented by
Dodoens’ illustration, that represented by Clusius’ figure, and the
three Morison figures associated with variety B. Clifford’s material
and Morison’s f. 50 have petaloid staminodia (‘. . . flore pleno &
prolifero’), are almost certainly of cultivated origin and would not
effectively restrict the application of subsp. bullatus. Dodoens’
illustration and Morison’s f. 51 belong to subsp. cytheraeus so either
would make an unfortunate and disruptive type choice. However,
Clusius’ illustration (copied by Morison as his f. 49) belongs to
subsp. bullatus and ‘Ranunculus grumosa radice I.’ in Clusius,
Rariorum plantarum historia: 238 (1601) is therefore designated
here as the lectotype of Ranunculus bullatus L. (Fig. 2). Given its
somewhat schematic nature, it seems wise to designate an epitype to
establish its application beyond any possible doubt, which we do
here (see Taxonomic Summary).

Study of the variation in Ranunculus bullatus L. s.l. by the senior
author (Baldini, in prep.), leads to the conclusion that R. bullatus L.
subsp. cytheraeus (Halacsy) Vierh. is best recognized at species
rank. This is based on several distinct features, including flower size

8

balletus, 8. RANUNCULUS foliis ovatis ferratis, {capo nudo
unifioro. Hort. cliff. 129: ae
stat latifolius bullatus, afphodeli radice. Bawh.
pia. LOT.
Rasunculus lufitanicus. Dod. pempt. 429.
Ranunculus grumofa radice 1. Cluf. bift. 1. it 238.
B. Ranunculus latifolius bullatus autumnalis, flore pleno
& prolitero. Morif. bifk. 2. p-447.S. 4-8 31-S+ 49: $0
ers):
Habitat in Lufitania, Crota, ¥
Fig. 1 The protologue of Ranunculus bullatus Linnaeus, Species

Plantarum: 550 (1753).

Ranuncul. grumofaradice 1.

Fig. 2 ‘Ranunculus grumosa radice I.’ in Clusius, Rariorum plantarum
historia: 238 (1601), the lectotype of Ranunculus bullatus L.

and arrangement, leaf morphology and habit, and their correlation
with geographical distribution. A new combination, relevant syn-
onymy, key and species descriptions and illustrations are given
below.

R.M. BALDINI AND C.E. JARVIS

TAXONOMIC SUMMARY

Key to Ranunculus bullatus and R. cytheraeus

l Leaf lamina ovate, crenate, never mucronate, more or less bullate and
pubescent, sometimes glabrous or hispid above or beneath; scape to 20—
25 cm long, never branched, more or less pubescent, 1-flowered;
achenes with a short, curved beak .............cccccsseccesseeeees 1. R. bullatus

— Leaf lamina obovate to elliptical, crenate-denticulate, mucronate, upper
surface villose especially at the margin; scape to 8 cm long, branched
with small bracteole at the base of each branch, with spreading hairs, 1—
2-flowered; achenes with a short, uncinate beak ...... 2. R. cytheraeus

1. Ranunculus bullatus L., Sp. pl. 1: 550 (1753). Type: “Ranuncu-
lus grumosa radice I.’ in Clusius, Rar. pl. hist.: 238 (1601),
lectotype, designated here. Epitype: ‘Flora Lusitanica. Prov.
Estremadura, ad pagum, 1848, leg. Dr. Welwitsch 198°’ (BM!),
designated here.

Fig. 3.

Ranunculus supranudus Jord. & Fourr., Brev. Pl. Nov. 1: 1 (1866).
Type. ‘Algeria: Daroussa prope Bone, ex dom. A. Joannon’. Not
located.

R. semicalvus Jord. & Fourr., Brev. Pl. Nov. 1: 2 (1866). Type:
‘Ajaccio, 15 octobre 1866, col. De Marsilly 101 bis’ (P!, lectotype,
designated here; BM!, K!-isotypes).

R. bullatus L. var. prolifer Gennari, Spec. e var. Fl. Sard.: 29 (1866),
non Cavara & Grande in Boll. Soc. Bot. Ital.: 101 (1925). Type:
‘Ranunculus bullatus scapo diviso seu prolifero, Cagliari ai
Cappuccini, 9bre. 1863, Gennari s. n.’ (TO!- lectotype, desig-
nated here).

Tonosmanthus semicalvus (Jord. & Fourr.) Jord. & Fourr., Icon. Fl.
Eur. 2: 14 (1869).

I. plantagineus Jord. & Fourr., Icon. Fl. Eur. 2: 14 (1869). Type:
‘Algeria, Costantine, tab. CCXLIV. — 326’ in Jordan & Fourreau,
Icon. Fl. Eur. 2. (1869), lectotype, designated here.

R. bullatus L. var. & ovatus Freyn in Willk. & Lange, Fl. Hisp. Prodr.
3: 919 (1880). Type: ‘Malaga, Willkomm exsicc. 1844, n. 418’,
(BM!-lectotype, designated here; G!-isotype).

R. bullatus L. var. B bellidifolius Freyn in Willk. & Lange, Fl. Hisp.
Prodr. 3: 919 (1880). Type: ‘Malaga, Kalisch’. Not located.

R. bullatus L. subsp. supranudus (Jord. & Fourr.) Vierh. in Osterr.
Bot. Z. 84: 129 (1935).

R. bullatus L. subsp. bullatus var. plantagineus (Jord. & Fourr.)
Maire in Jahand. & Maire, Cat. Pl. Maroc: 994 (1940).

R. bullatus L. subsp. bullatus var. supranudus (Jord. & Fourr.) Maire
in Jahand. & Maire, Cat. Pl. Maroc: 994 (1941).

R. bullatus L. subsp. eu-bullatus Maire, Fl. Afrique N. 11: 139
(1964).

Perennial herb with fibrous roots and fusiform tubers; leaves (2—)3—
5(-7) in basal rosette; petiole 2—5(—7) cm long, with silky hairs;
lamina ovate, crenate, sometimes crenate-denticulate, never mucro-
nate, more or less bullate and pubescent, sometimes glabrous or
hispid above or beneath, (1—)2—8 x 1-5(—6) cm; scapes erect, never
branched, up to 20-25 cm long, more or less pubescent, sometimes
glabrous, 1—flowered; flowers 1—-2(—3) cm in diameter, with 5 green
sepals 0.3-0.6 cm long and 5—10(—12) yellow and obovate petals
0.5-1.5 cm long; receptacle glabrous, not elongated; achenes gla-
brous, ovate-inflated with a short and curved beak.

DISTRIBUTION
EUROPE. Tutin (1964: 234); Greuter et al. (1989: 428); Jalas & Suominen
(1989: map no. 1839); Tutin & Akeroyd (1993: 281). Portugal: Pereira

TYPIFICATION OF RANUNCULUS BULLATUS L.

Fig.3 = Ranunculus bullatus L. From the epitype specimen, Welwitsch 198 (BM), A. Maury delineavit. A. whole plant x 1; B. petal x 4; C. achene x 20;
the additional leaves (x 1) show two of the more frequent foliar morphotypes found in this variable species. The upper leaf (D) is from a collection from the
Balearic Islands (Bianor 2518, BM!); the lower leaf (E) is from a collection from Sicily (Ross 103, G!).

R.M. BALDINI AND C.E. JARVIS

Fig.4 Ranunculus cytheraeus (Halacsy) Baldini. From material from Cyprus (Agia Napa distr., Famagusta, 15 Dec. 1940), Davis 2052 (BM), A. Maury

delineavit. A. whole plant x 2; B. petal x 5; C. achene x 20.

Coutinho (1974: 279). Spain: L6pez Gonzalez (1986). Balearic Islands:
Smythies (1984: 448). Corsica: Paradis et al. (2000). Italy (incl. Sardinia
and Sicily): Fiori (1898: 506; 1924: 666); Zangheri (1976: 161); Pignatti
(1982: 318). Crete: Turland et al. (1993: 132 sub R. bullatus L. p.p.).
Turkey: Davis (1965: 179). Cyprus: Meikle (1977: 48).

NORTH AFRICA. Maire (1964: 137 excl. Libya where only R. cytheraeus
(Halacsy) Baldini occurs). Morocco: Battandier (1888: 10); Jahandiez &
Maire (1932: 250); Négre (1961: 241). Algeria: Battandier & Trabut (1904:
12); Quézel & Santa (1962: 370). Tunisia: Battandier & Trabut (1904: 12);
Pottier-Alapetite (1979: 156).

2. Ranunculus cytheraeus (Halacsy) Baldini, stat. nov.

Ranunculus bullatus L. var. cytheraeus Halacsy in Vierh., K. K.
Zool.-Bot. Ges. Wien 54: 483 (1904). Type: ‘Ranunculus bullatus
L. v. cytheraeus. Foliis grosse inciso-dentatis, sparse pilosis
subglabrisve, petalis 5, obovatis vel obovato-cuneatis. Ins. Cythera:
in dumosis siccis reg. inf., 25.X1.1903, Chr. Leonis 5° (LY!-
lectotype, designated here; LY!-isotypes).

Fig. 4.

R. bullatus L. var. cuneifolius Coustur. & Gand. in Bull. Soc. Duffour:

TYPIFICATION OF RANUNCULUS BULLATUS L.

36 (1914). Type: ‘Ranunculus bullatus L., Spec., 774. V.
cuneifolius Cousturier et Gandoger. Var. nov.! Cat. Cam. 37 — var.
Créte: Candie. Dans les collines au nord d’ Hagios-Déka. Décembre
1913, P. Cousturier 1479 (LY !-lectotype, designated here; MPU!-
isotype).

R. bullatus L. forma cuneifolius (Coustur. & Gand.) Coustur. &
Gand. in Bull. Soc. bot. France 63: 13 (1916).

R. bullatus L. var. cyrenaicus Pamp. in Nuovo Giorn. Bot. Ital., n. s.,
26: 212 (1919). Type: ‘Ranunculus bullatus L. var. cyrenaicus
Pamp., 5 Gennaio 1917: in fioritura, depressione del terreno fra le
due Palme e la linea ferroviaria di Benina in direzione del Giok
(Kebir). Arabo = Ain el noogia = occhio di pecora. Gli arabi ne
raccolgono i fiori e li uniscono alla mastice ricavata dalla pianta
Hellek (Periploca laevigata) detta Elek, per darle il colorito
giallo. Zanon 24’ (FI!-lectotype, designated here).

R. bullatus L. var. prolifer sensu Cavara & Grande in Bull. Soc. Bot.
Ttal.: 101 (1925), non Gennari, Spec. e var. Fl. Sard.: 29 (1866).

R. bullatus L. subsp. cytheraeus (Halacsy) Vierh. in Osterr. Bot. Z.
84: 129 (1935).

R. bullatus L. subsp. cyrenaicus (Pamp.) Maire, Fl. Afrique N. 11:
139 (1964).

Perennial herb with fibrous roots and fusiform tubers; leaves 3—5(—
9) in basal rosette, with silky hairs; pedicels 0.5—1.5(—2.5) cm long;
lamina from obovate to elliptical, crenate-denticulate, mucronate,
upper surface villose especially at the margin and somewhat pubes-
cent beneath; scapes erect, 3—7(—8) cm long, with spreading hairs,
branched with small bracteole at the base of branch; flowers 1-2,
(0.5—)1—1.5(—2) cm in diameter, with 5 green sepals 0.3—0.5(-0.7)
cm long and 5(-6) yellow and obovate petals, (0.5—)0.8—1.2(—1.5)
cm long; receptacle glabrous and elongated; achenes glabrous, 0.1—
0.3 cm long, inflated with a short and uncinate beak.

DISTRIBUTION

GREECE. Cythera: Halacsy (1904 sub R. bullatus L. var. cytheraeus
Halacsy); Davis (1965: 179 sub R. bullatus subsp. cytheraeus (Halacsy)
Vierh.). Cyprus: Davis (1965: 179 sub R. bullatus subsp. cytheraeus (Halacsy)
Vierh.); Meikle (1977: 48 sub R. bullatus subsp. cytheraeus (Halacsy)
Vierh.). Crete: Rechinger (1943: 188 sub R. bullatus L. var. cuneifolius
Coustur. & Gand. p.p. & var. cytheraeus Halacsy in Vierh. p.p.); Barclay
(1986: 82 sub R. bullatus L. p.p.); Turland et al. (1993: 132 sub R. bullatus L.
p.p.).

NORTH AFRICA. Libya: Béguinot & Vaccari (1912a, 1912b sub R.
bullatus sensu Bég. & Vaccari, non L. (1753); Pampanini & Zanon (1919:
212 sub R. bullatus L. var. cyrenaicus Pamp.); Pampanini (1931: 203 sub R.
bullatus L. var. cyrenaicus Pamp.); Cavara & Grande (1925 sub R. bullatus
var. prolifer sensu Cavara & Grande, non Gennari (1866); Maire (1964: 139
sub R. bullatus L. subsp. cyrenaicus (Pamp.) Maire; Davis (1965: 179 sub R.
bullatus L. subsp. cytheraeus (Halacsy) Vierh.); Qaiser (1984: 35 sub R.
bullatus L. subsp. cytheraeus (Halacsy) Vierh.).

ACKNOWLEDGEMENTS. We are grateful to the curators of the herbaria
cited above for permission to study the collections in their care, and in
particular to Prof. P. Schafer (MPU). We are also grateful to Giorgio
Padovani (Museo Botanico dell’ Universita degli Studi di Firenze) and
Laura Settesoldi (Erbario Tropicale di Firenze) for their kind assistance in
transferring images to CD. CEJ acknowledges generous grant support to
the Linnaean Plant Name Typification Project by the Linnean Society of
London.

11

REFERENCES

Barclay, C. 1986. Crete. Checklist of the vascular plants. Englera 6: 1-138.

Battandier, J.-A. 1888. Dicotylédones I. Thalamiflores. Jn J.-A. Battandier & L.
Trabut (Eds), Flore de l’Algérie. Alger.

& Trabut, L. 1904. Flore analytique et synoptique de |’Algérie et de la Tunisie.
Alger.

Bauhin, C. 1623. Prodromus theatri botanici. Basel.

Béguinot, A. & Vaccari, A. 1912a. Contributo alla flora della Libia. Monografie e
Rapporti Coloniali 16: 1-70. 10 plates. Roma.

& 1912b. Specie nuove o rare per la Flora della Libia. Padova.

Cavara, F. & Grande, L. 1925. Altre piante rare o nuove della Cirenaica. Bollettino
della Societa Botanica Italiana: 100-105.

Clusius, C. 1601. Rariorum plantarum historia. Antverpiae.

Cornut, J.-P. 1635. Canadensium plantarum. Paris.

Davis, P.H. 1965. Ranunculus. In P.H. Davis (Ed.), Flora of Turkey and the East
Aegean Islands 1: 146-195. Edinburgh.

Dodoens, R. 1583. Stirpium historiae pemptades sex, sive libri xxx. Antverpiae.

Fiori, A. 1898. Ranunculus (Tourn.) L. Jn A. Fiori & G. Paoletti (Eds), Flora Analitica
d'Italia 1: 500-516. Padova.

— 1924. Raninculus L. In A. Fiori, Nuova Flora Analitica d'Italia 1: 659-678.
Firenze.

Gandoger, M. 1889. Ranunculus. In Florae Europae 1: 167-169. Parisiis.

Gennari, P. 1866. Specie e varieta pitt rimarchevoli e nuove da aggiungere alla Flora
Sarda: 29. Cagliari.

Greuter, W., Burdet, H.M. & Long, G. 1989. Med-Checklist 4. Dicotyledones
(Lauraceae-Rhamnaceae). Genéve.

Halacsy E. von, 1904. Bericht der Sektion fiir Botanik. Versammlung am 29. April
1904. Verhandlungen der kaiserlich-koniglichen zoologisch-botanischen Gesellschaft
in Wien 54: 483-484.

Jahandiez, E. & Maire, R. 1932. Catalogue des plantes du Maroc 2. Alger.

Jalas, J. & Suominen, J. 1989. Atlas Florae Europaeae 8. Nymphaeaceae to
Ranunculaceae. Helsinki.

Linnaeus, C. 1738. Hortus Cliffortianus. Amsterdam.

— 1753. Species plantarum. Holmiae [Stockholm].

L6épez Gonzalez, G. 1986. Ranunculus L. Sect. 6. Physophyllum Freyn. In S. Castroviejo
et al. (Eds), Flora Iberica 1: 325-326. Madrid.

Maire, R. 1964. Ranunculus. In R. Maire, (Ed.), Flore de l’Afrique du Nord 11: 112-
204. Paris.

Meikle, R. D. 1977. Flora of Cyprus 1. Kew.

Morison, R. 1680. Plantarum historiae universalis Oxoniensis 2. Oxonii.

Négre, R. 1961. Petite flore des régions arides du Maroc Occidental 1. Paris.

Pampanini, R. 1931. Prodromo della Flora Cirenaica. Forli.

& Zanon, V. 1919. Nuovi contributi alla conoscenza della flora della Cirenaica.
Nuovo Giorn. Bot. Ital., n.s., 26: 205-220.

Paradis G., Pozzo di Borgo, M.-L. & Jeanmonod, D. 2000. Ranunculus bullatus L.
In D. Jeanmonod (Ed.), Notes et contributions a la flore de Corse, X VI. Candollea
55: 59 (fig. 2), 62.

Pereira Coutinho, A.X. 1974. Flora de Portugal, 2™ ed. (Reprint). Leutershausen.

Pignatti, S. 1982. Flora d’Italia 1. Bologna.

Pottier-Alapetite, G. 1979. Flore de la Tunisie. Angiospermes-Dicotyledones. Apetales-
Dialypetales 1. Paris.

Qaiser, M. 1984. Ranunculaceae. Jn S.M.H. Jafri & A. El-Gadi (Eds), Flora of Libya
108: 1-54. Tripoli.

Quézel, P. & Santa, S. 1962. Nouvelle Flore de l’Algérie et des régions désertiques
meéridionales 1. Paris.

Rechinger K.H. fil. 1943. Flora Graeca. Wien.

Savage, S. 1945. A catalogue of the Linnaean herbarium. London.

Smythies, B.E. 1984. Flora of Spain and the Balearic Islands. Checklist of vascular
plants. II. Cruciferae-Rutaceae. Englera 3: 213-486.

Turland, N.J., Chilton, L. & Press, J.R. 1993. Flora of the Cretan area. Annotated
checklist & atlas. London.

& Jarvis, C.E. (Eds) 1997. Typification of Linnaean specific and varietal names
in the Leguminosae (Fabaceae). Taxon 46: 457-485.

Tutin, T.G. 1964. Ranunculus L. In T.G. Tutin et al. (Eds), Flora Europaea 1: 223-237.
Cambridge.

— & Akeroyd, J.R. 1993. Ranunculus L. In T.G. Tutin et al. (Eds), Flora Europaea,
2™ ed. 1: 269-285. Cambridge.

Vierhapper, F. & Rechinger, K.H. fil. 1935. Bearbeitung der von Ignaz Dorfler im
Jahre 1904 auf Kreta gesammelten Bliiten- und Farnpflanzen. Osterreichische
Botanische Zeitschrift 84: 123-197.

Zangheri, P. 1976. Flora Italica 1. Padova.

bee oe

Bull. nat. Hist. Mus. Lond. (Bot.) 32(1): 13-59

4
7,

Issued 27 June 2002

Catalogue of the lichens of Thailand

PATRICIA A. WOLSELEY

Department of Botany, The Natural History Museum, Cromwell Road, London SW7 5BD

BEGONA AGUIRRE-HUDSON

58 Willowhayne Drive, Walton-on-Thames, Surrey KT12 2NR

PATRICK M. McCARTHY

Australian Biological Resources Study, GPO Box 787, Canberra, A.C.T. 2601, Australia

Synopsis. A catalogue of the lichens of Thailand is provided based on published reports. It includes publication details of 554
species and infra-specific taxa together with information on distribution in Thailand, occurrence in other south and south-east
Asian countries, literature references and the locations of named specimens.

INTRODUCTION

Before the 1990s, knowledge of the diversity and distribution of
the lichen flora of Thailand was based almost entirely on collec-
tions made by a handful of foreign visitors in the first half of the
twentieth century (Wolseley & Aguirre-Hudson, 1991). During a
research project on lichens as environmental indicators based at
the Natural History Museum, London, a comprehensive survey of
literature concerning lichens in south-east Asia was made and a
data base constructed, which forms the basis of this checklist
(Aguirre-Hudson & Wolseley, 1994). It was not possible during
this time to trace all types described in the literature, or to confirm
the accuracy of many earlier identifications. Since then there has
been a rapid and productive expansion of lichenological research
by Thai practitioners, especially at the Lichen Research Unit,
Ramkhamhaeng University, Bangkok, further expanding the list
and including many newly described species (Boonpragob et al.,
1998; Pooprang et al., 1999; Vongshewarat et al., 1999; Hom-
chantara & Coppins, 2002). The present checklist is conceived as
a working document for all researchers in south-east Asia to
provide both the literature sources and herbaria in which speci-
mens from Thailand may be seen.

SCOPE AND ARRANGEMENT

This catalogue documents all 554 currently accepted taxa that have
been reported in the literature from Thailand, as well as synonyms
previously applied to Thai specimens. Genera and species are
arranged in alphabetical order, and generic and family concepts
follow Hawksworth et al. (1995). Where new combinations have not
been made, taxa have been placed in the current genus within square
brackets []. Accepted names are shown in bold and synonyms in
italics. Distribution within Thailand is given where possible by
province including new provinces and their former names (Fig. 1).
Literature pertaining to distribution in Thailand is cited and refer-
ences provided. Location of specimens in herbaria is indicated by
abbreviations following Index Herbariorum (Holmgren et al., 1990)
as shown below:

AAU: | Herbarium Jutlandicum, Botanical Institute, University of
Aarhus, Denmark.

© The Natural History Museum, 2002

BKF: Forest Herbarium, Royal Forest Department, Bangkok,
Thailand.

BM: Herbarium, Botany Department, The Natural History
Museum, London, UK.

Cc Herbarium, Botanical Museum, University of Copenhagen,
Denmark.

CANB: Australian National Herbarium, CSIRO, Canberra,
Australia.

CMU: Herbarium, Biology Department, Chiang Mai University,
Thailand.

COLO: Herbarium, Colorado University, Colorado, USA.

DUKE: Hebarium, Botany Department, Duke University, Dur-
ham, North Carolina, USA.

E Herbarium, Royal Botanic Garden, Edinburgh, UK.

FH: Farlow Refence Library and Herbarium of Cryptogamic
Botany, Harvard University, Cambridge, Massachusetts,

USA.
GB: Herbarium, Botanical Museum, Gothenburg, Sweden.
H: Herbarium, Botanical Museum, University of Helsinki,
Finland.
KF: Herbarium, Forest Research Institute, Seoul, Korea.
KYO: Herbarium, Botany Department, Kyoto University, Japan.
| Be Rijksherbarium, Leiden, Netherlands.
LG: Herbarium, Botany Department, Liége University, Belgium.
OSA: — Herbarium, Osaka Museum of Natural History, Japan.
PSU: | Herbarium, Department of Biology, Prince of Songkla

University, Haad Yai, Thailand.

RAMK: Herbarium, Biology Department, Ramkhamhaeng Uni-
versity, Bangkok, Thailand.

TNS: Herbarium, Department of Botany, National Science Mu-
seum, Tokyo, Japan.

TUR-V: Vainio herbarium, Biology Department, University of
Turku, Finland.

UPS: — Botanical Museum, Uppsala University, Sweden.

US: United States National Herbarium, Botany Department,
Smithsonian Institution, Washington DC, USA.

Distribution of taxa in in other countries in adjacent areas of south
and south-east Asia is indicated under the subheading ‘ASIA’. We
have avoided making taxonomic judgements on specimens we have
not examined. Consequently, a number of highly dubious identifica-
tions remain, as do species which will almost certainly be transferred
to other genera.

Fig. 1

Although this Catalogue is based exclusively on published
reports and Homchantara’s unpublished thesis, we have not
included data from an anonymous checklist on the web-site of the
Lichen Research Unit at Ramkhamhaeng University, Bangkok
(www.ru.ac.th/lichen). This checklist includes the names (without
additional information) of 404 species based on specimens held at
RAMK. However, although many names can be attributed to
literature sources and authoritative determinations, a significant
proportion are unauthenticated and are previously unpublished
taxa which must be considered possibly doubtful. We are aware
that this Catalogue documents a rather small portion of what is
almost certainly a very diverse lichen flora; indeed, the actual
number of species is probably in excess of 2000. However, we
hope that it will provide a basis for future taxonomic research on
Thai lichens, and that the research programs currently underway
in Thailand will ensure its rapid obsolescence.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Key to provinces of Thailand

Northern region
Mae Hong Son
Chiang Mai
Chiang Rai
Phayao

Nan

Lamphun
Lampang

Phrae

Uttardit

10. Tak

11. Sukhotai

12. Phitsanulok

13. Kamphaeng Phet
14. Phichit

15. Nakhon Sawan
Northeastern region
16. Petchabun

7 T0e1

18. Udon Thani

19. Nong Khai

20. Sakon Nakhon
21. Nakhon Phanom
22. Kalasin

23. Maha Sarakham
24. Khon Kaen
Eastern region

25. Chaiyaphum

26. Nakhon Ratchasima
27. Buri Ram

28. Surin

29. Roi Et

30. Yasothon

31. Si Sa Ket

32. Ubon Ratchathani
Southwestern region
33. Uthai Thani

34. Kanchanaburi

35. Ratchaburi

36. Phetchaburi

37. Prachuap Khiri Khan
Central region

38. Chai Nat

SOOT SON) Na

39. Sing Buri

40. Lop Buri

41. Suphan Buri

42. Ang Thong

43. Phra Nakhon Si Ayutthaya

44. Saraburi

45. Nakhon Nayok

46. Nakhon Pathom

47. Pathum Thani

48. Nonthaburi

49. Bangkok

50. Samut Prakan

51. Samut Songkram

52. Samut Sakhon

Southeastern region

53. Prachin Buri

54. Chachoengsao

55. Chon Buri

56. Rayong

57. Chanthaburi

58. Trat

Peninsular region

59. Chumphon

60. Ranong

61. Surat Thani

62. Phangnga

63. Phuket

64. Krabi

65. Nakhon Si Thammarat

66. Phatthalung

67. Trang

68. Satun

69. Songkhla

70. Pattani

71. Yala

72. Narathiwat

New provinces

73. Nong Bua Lamphu formerly in Udon
Thani

74. Mukdahan formerly in Nakhon Phanom

75. Amnat Charoen formerly in Ubon
Ratchathani

76. Sra Kaeo formerly in Prachin Buri

Map of Thailand showing location of provinces. Reproduced with permission from Hamalainen & Pinratana (2001).

CATALOGUE

Anisomeridium (Miill. Arg.) M. Choisy;

Monoblastiaceae

[Anisomeridium|] Microthelia asiaticum Vain., Hedwigia

46: 180 (1907).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 149), Hawksworth

(1985: 135).
SPECIMEN.

ASIA. _ India.

TUR-V (holotype).

Anisomeridium foliicola R. Sant. & Tibell, Austrobaileya

2: 531 (1988).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).

CATALOGUE OF THE LICHENS OF THAILAND

SPECIMENS. CANB.

ASIA. India, Papua New Guinea, Philippines, Sri Lanka.

Anthracothecium Hampe ex A. Massal.;
Pyrenulaceae

Anthracothecium macrosporum Miill. Arg., Linnaea 63:
44 (1888).
Anthracothecium pyrenuloides auct. non (Mont.) Miill. Arg.

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99, as
Anthracothecium pyrenuloides).

SPECIMENS. BM.

ASIA. India, Indonesia.

Anthracothecium parameroides (Vain.) Zahlbr., Cat. Lich.
Univ. 1: 466 (1922).
Bottaria parameroides Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 179; 1909: 143,
both as Bottaria parameroides).

SPECIMEN. TUR-V (holotype).

ASIA. Indonesia.

Anthracothecium prasinum (Eschw.) R.C. Harris, in Egan,
Bryologist 90: 163 (1987).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-
Hudson (1997a: 333), Boonpragob et al. (1998: 216).

SPECIMENS. BM, RAMK.

ASIA. Papua New Guinea.

Anthracothecium roseum (Vain.) Zahlbr., Cat. Lich. Univ.
1: 468 (1922).

Bottaria rosea Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 179; 1909: 143,
both as Bottaria rosea).

SPECIMEN. TUR-V (holotype).
ASIA. —

Arthonia Ach.; Arthoniaceae

Arthonia complanata Fée, Essai Crypt. Ecore. Offic.: 54
(1824).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 139).
SPECIMEN. TUR-V.

ASIA. China, Indonesia, Papua New Guinea, Philippines.
Arthonia aff. mira R. Sant., Symb. Bot. Upsal. 12(1): 77
(1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. —

15

Arthonia obscuratula Zahlbr., Cat. Lich. Univ. 2: 71
(1922).

Arthonia stenographella subsp. obscurata Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 140, as Arthonia
stenographella subsp. obscurata).

SPECIMEN. TUR-V (holotype).

ASIA. —

Arthonia ochrodes Nyl., in Willey, Syn. Gen. Arthonia: 10
(1890).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 142).
SPECIMEN. TUR-V.

ASIA. —

Arthonia perpallens Ny\1., Ann. Sci. Nat., Bot., sér. 4, 20:
231 (1863).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 141).
SPECIMENS. TUR-V.
ASIA. —

Arthonia rhizophorae Vain., Bot. Tidsskr. 29: 141 (1909).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 141).
SPECIMEN. TUR-V (holotype).

ASIA. —

Arthonia varia Ny. var. stenographella (Ny1.) Vain., Ann.
Acad. Sci. Fenn., ser. A, 15(6): 393 (1921).

Arthonia stenographella Ny.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 140, as A.
stenographella).

SPECIMEN. TUR-V (holotype).

ASIA. Philippines, Sri Lanka.

Arthopyrenia A. Massal.; Arthopyreniaceae
Arthopyrenia siamensis Vain., Suom. Elain-ja Kasvit.
Seuran Van. Julk. 1(3): 55 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 55).
SPECIMEN. TUR-V (holotype).

ASIA. —

Arthothelium A. Massal.; Arthoniaceae
Arthothelium monosporum (Vain.) Zahlbr., Cat. Lich.
Univ. 2: 129 (1922).

Arthonia monospora Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 54, as

Arthonia monospora).
SPECIMEN. TUR-V (holotype).

ASIA. —

16

Arthothelium ruanum (A. Massal.) Korb., Parerg. Lich.:
263 (1861).

Arthonia ruana A. Massal.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 139, as Arthonia
ruana).

SPECIMEN. TUR-V.

ASIA. —

Arthothelium ruanum var. phaeosporum (Vain.) Zahlbr.,
Cat. Lich. Univ. 2: 132 (1922).

Arthonia ruana var. phaeospora Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 139, as Arthonia
ruana var. phaeospora).

SPECIMEN. TUR-V (holotype).

ASIA. —

Arthothelium spectabile Flotow ex A. Massal., Ric. Auton.
Lich. Crost.: 54 (1852).

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99).
SPECIMEN. BM.

ASIA. China, India, Malaysia, Sri Lanka.

Bacidia De Not.; Bacidiaceae
Bacidia affinis Vain., Acta Soc. Fauna Fl. Fenn. 53(1):
146 & 154 (1922).

Bacidia atrosanguinea Th. Fr.

THAILAND. Chumphon (Koh TaoI.). Paulson (1930: 99, as Bacidia
atrosanguinea).

SPECIMEN. BM (not located).

ASIA. —

Bacidia manilensis var. siamensis Vain., Suoml. Elain-ja
Kasvit. Seuran Van. Julk. 1(3): 46 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 46).
SPECIMEN. TUR-V (holotype).
ASIA. —

Bacidia umbrina var. turgida Th. Fr., Lichenogr. Scand. 1:
365 (1874).

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99).
SPECIMEN. BM (not located).
ASIA. —

Bathelium Ach.; Trypetheliaceae

Bathelium madreporiforme (Eschw.) Trevis., Spighe e
Paglie: 20 (1853).

Laurera madreporiformis (Eschw.) Riddle.

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217, as Laurera madreporiformis).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

SPECIMENS. CANB.

ASIA. India, Myanmar.

Brigantiaea Trevis.; Brigantiaeaceae

Brigantiaea leucoxantha (Spreng.) R. Sant & Hafellner, in
Hafellner & Bellemére, Nova Hedwigia 35: 246 (1981).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary). Wolseley & Aguirre-Hudson (1997a: 333), Boonpragob et al.

(1998: 216).
SPECIMENS. BM, RAMK.

ASIA.
Taiwan.

India, Malaysia, Papua New Guinea, Philippines, Sri Lanka,

Brigantiaea tricolor (Mont.) Trevis., Spighe e Paglie: 9
(1853).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.),
Lampang (Doi Khun Tan N.P.). Wolseley & Aguirre-Hudson (1997a:

333):
SPECIMENS. BM, CMU.

ASIA.
Taiwan.

India, Indonesia, Malaysia, Papua New Guinea, Sri Lanka,

Buellia De Not.; Physciaceae

Buellia blastenioides Vain., Hedwigia 46: 171 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 114).
SPECIMEN. 'TUR-V (holotype).

ASIA. —

Buellia pleiophoroides var. meiospermoides Vain., Bot.
Tidsskr. 29: 113 (1909).

THAILAND. Chiang Mai (Doi Suthep N.P.), Trat (Koh Chang 1.),
Uthai Thani (Huay Kha Khaeng N.P.). Vainio (1909: 113), Wolseley

& Aguirre-Hudson (1997a: 339).
SPECIMENS. BM, TUR-V (holotype).

ASIA. —

Buellia stramineoatra Vain., Hedwigia 46: 172 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 115).
SPECIMENS. TUR-V (syntypes).

ASIA. —

Buellia subdives Vain., Hedwigia 46:172 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1907: 172; 1909: 114).
SPECIMENS. 'TUR-V (syntypes).

ASIA. —
Bulbothrix Hale; Parmeliaceae

Bulbothrix goebelii (Zenker) Hale, Smithsonian Contr. Bot.
32: 14 (1976).

CATALOGUE OF THE LICHENS OF THAILAND
Parmelia addenda Vain.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Queen Sirikit Botanical Garden), Trat (Koh Chang I.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Vainio (1909: 107, as
Parmelia addenda), Hale (1976a: 14), Mongkonsuk et al. (1996:
16), Wolseley & Aguirre-Hudson (1997a: 333).

SPECIMENS. C (lectotype), TUR-V (isolectotype), BKF, BM, CMU,
US, Queen Sirikit Botanical Garden Herbarium.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Taiwan.

Bulbothrix hypochraea (Vain.) Hale, Phytologia 28: 480
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Pooprang et al. (1999: 118).

SPECIMENS. RAMK.

ASIA. —

Bulbothrix isidiza (Ny1.) Hale, Phytologia 28: 480 (1974).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P., Maetang), Lampang (Doi Pae Luang), Loei (Ban
Sui, Phu Kradung), Phitsanulok (Phu Hin Rongkla, N.P.), Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-
Hudson (1995: 318; 1997a: 333; 1997b: 355-356), Moon et al.

(2000a: 99).
SPECIMENS. BKF, BM, CMU.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea,
Philippines, Sri Lanka, Taiwan.

Bulbothrix laevigatula (Ny1.) Hale, Phytologia 28: 480
(1974).

Parmelia hookeri Taylor.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 37, as
Parmelia hookeri).

SPECIMEN. TUR-V.

ASIA. —

Bulbothrix meizospora (Ny1.) Hale, Phytologia 28: 480
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Loei
(Phu Rhoa N.P.), Nakhon Nayok (Khao Yai N.P.). Boonpragob et al.

(1998: 216), Pooprang et al. (1999: 118).
SPECIMENS. BM, RAMK.

ASIA. India, Indonesia, Nepal, Sri Lanka.

Bulbothrix pigmentacea (Hale) Hale, Phytologia 28: 480
(1974).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &

Aguirre-Hudson (1995: 318; 1997a: 333; 1997b: 355-56).
SPECIMENS. BKF, BM, CMU.

ASIA. Malaysia, Philippines.

Bulbothrix queenslandica (Elix & G.N. Stevens) Elix,
Mycotaxon 47: 126 (1993).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Loei
(Nahaew N.P.), Phitsanulok (Phu Hin Rongkla N.P.). Pooprang et al.
(1999: 119).

SPECIMENS. RAMK.

ASIA. —

Bulbothrix sensibilis (J. Steiner & Zahlbr.) Hale,
Phytologia 28: 481 (1974).

THAILAND. Loei (Nahaew N.P.). Pooprang et al. ( 1999: 119).
SPECIMENS. RAMK.

ASIA. India, Sri Lanka.

Bulbothrix setschawensis (Zahlbr.) Hale, Phytologia 28:
481 (1974).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Lampang, Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-
Hudson (1997a: 333).

BKF, BM, CMU.
ASIA. China, India, Nepal.

SPECIMENS.

Byssoloma Trevis.; Pilocarpaceae
Byssoloma chlorinum (Vain.) Zahlbr., Cat. Lich. Univ. 8:
233 (1932).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Papua New Guinea.

Byssoloma leucoblepharum (Ny\.) Vain., Dansk Bot. Ark.
4: 23 (1926).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Wolseley &
Aguirre-Hudson (1997a: 333), Boonpragob et al. (1998: 216).

SPECIMENS. BM, RAMK.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Singapore, Vietnam.

Byssoloma tricholomum (Mont.) R. Sant., Symb. Bot.
Upsal. 12(1): 480 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. China, Philippines, Singapore.
Calenia Miill. Arg. emend. R. Sant.; Gomphillaceae
Calenia graphidea Vain., Ann. Acad. Sci. Fenn., ser. A, 15:

163 (1921).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 216).

SPECIMENS. RAMK.

18

ASIA. Indonesia, Malaysia, Papua New Guinea, Philippines.

Calenia aff. phyllogena (Miill. Arg.) R. Sant., Symb. Bot.
Upsal. 12(1): 349 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. India, Papua New Guinea, Philippines, Sri Lanka.

Calopadia Veézda; Ectolechiaceae
Calopadia fusca (Miill. Arg.) Veézda, Folia Geobot.
Phytotax. 21: 215 (1986).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. CANB.

ASIA. Indonesia, Papua New Guinea, Philippines, Sri Lanka.

Caloplaca Th. Fr.; Teloschistaceae

Caloplaca aequata (Hue) Zahlbr., Cat. Lich. Univ. 7: 211
(1931).

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99).
SPECIMEN. BM (not located).

ASIA. —

Caloplaca flavorubescens (Huds.) Laundon, Lichenologist
8: 147 (1976).

Caloplaca aurantiaca (Lightf.) Th. Fr.

THAILAND. Chiang Mai (Doi Inthanon N.P.), Chumphon (Koh
Tao I.). Paulson (1930: 99, as Caloplaca aurantiaca).

SPECIMENS. BM.

ASIA. Sri Lanka, Taiwan.

Caloplaca holocarpa (Hoffm.) A.E. Wade, Lichenologist
3: 11 (1965).
Caloplaca pyracea (Ach.) Th. Fr.

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99, as
Caloplaca pyracea).

SPECIMEN. BM (not located).

ASIA. China, India.

[Caloplaca] Placodium testaceorufum Vain., Hedwigia 46:
170 (1907).

THAILAND. ‘Trat (Lem Ngob). Vainio (1909: 111, as Placodium
testaceorufum).

SPECIMEN. TUR-V (holotype).

ASIA. —
Canomaculina Elix & Hale; Parmeliaceae

Canomaculina subsumpta (Nyl.) Elix, Mycotaxon 65: 477
(1997).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Rimeliella subsumpta (Nyl.) Kurok.

THAILAND. Chiang Mai (Maetang). Kurokawa (1991: 9, as

Rimeliella subsumpta); Moon et al. (2000a: 99).
SPECIMEN. TNS.

ASIA. India, Papua New Guinea.

Canomaculina subtinctoria (Zahlbr.) Elix, Mycotaxon 65:
477 (1997).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Loei
(Nahaew N.P.). Mongkonsuk et al. (1996: 17), Pooprang et al.
(1999: 119).

SPECIMENS. RAMK.

ASIA. China, India, Indonesia, Nepal.

Canoparmelia Elix & Hale; Parmeliaceae

Canoparmelia adspersa (Vain.) Elix & Hale, in Elix,
Johnston & Verdon, Mycotaxon 27: 278 (1986).

Parmelia adspersa Vain.

THAILAND. ‘Trat (Lem Ngob). Vainio (1907: 168, as Parmelia
adspersa).

SPECIMENS. C (isolectotype), TUR-V (lectotype and syntype).

ASIA. Papua New Guinea, Philippines.

Canoparmelia concrescens (Vain.) Elix & Hale,
Mycotaxon 27: 278 (1986).

THAILAND. Phitsanulok (Phu Hin Rongkla, N.P.). Moon et al.
(2000a: 100).

SPECIMENS. TNS.

ASIA. —

Canoparmelia ecaperata (Miill. Arg.) Elix & Hale, in Elix,
Johnston & Verdon, Mycotaxon 27: 278 (1986).

Parmelia malaccensis var. laeteflavens Vain.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P., Maetang), Loei (Phu Kradung N.P.), Nakhon
Ratchasima (Khao Yai N.P.), Phitsanulok (Phu Hin Rongkla, N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Vainio (1921:
38, as Parmelia malaccensis var laeteflavens), Wolseley & Aguirre-
Hudson (1995: 318; 1997a: 333; 1997b: 355-56), Boonpragob et al.
(1998: 216), Moon et al. (2000a: 100).

SPECIMENS. BM, CMU, RAMK, TNS, TUR-V (lectotype and
syntype of P. malaccensis var laeteflavens), US.

ASIA. India, Nepal.

Canoparmelia owariensis (Asahina) Elix, Mycotaxon 47:
127 (1993).

Pseudoparmelia owariensis (Asahina) Hale.

THAILAND. Chiang Mai (Doi Saget), Loei (Ban Sui, Phu Kradung),
Phitsanulok (Phu Hin Rongkla, N.P.). Hale (1976b: 39, as
Pseudoparmelia owariensis), Moon et al. (2000a: 100).

TNS.
ASIA. China (Hong Kong).

SPECIMENS.

CATALOGUE OF THE LICHENS OF THAILAND

Canoparmelia pustulescens (Kurok.) Elix, Mycotaxon 47:
127 (1993).

THAILAND. Chiang Mai (Doi Saget), Loei (Phu Kradung). Moon

et al. (2000a: 101).
SPECIMENS. TNS.

ASIA. — India.

Canoparmelia salacinifera (Hale) Elix & Hale, in Elix,
Johnston & Verdon, Mycotaxon 27: 279 (1986).
Pseudoparmelia salacinifera (Hale) Hale.

THAILAND. Chiang Mai (Doi Saget, Doi Suthep N.P.), Loei (Phu
Kradung N.P.). Hale (1976a: 46, as Pseudoparmelia salacinifera),
Wolseley & Aguirre-Hudson (1995: 318), Moon et al. (2000a: 101).

SPECIMENS. BKF, BM, CMU, TNS, US.

ASIA. —

Canoparmelia texana (Tuck.) Elix & Hale, in Elix,
Johnston & Verdon, Mycotaxon 27: 279 (1986).

THAILAND. Chiang Mai (Chiang Dao N.P., Queen Sirikit Botani-
cal Garden), Lampang (Doi Khun Tan N.P.), Loei (Phu Kradung

N.P.). Hale (1976a: 53), Moon et al. (2000a: 101).
SPECIMENS. BKF, BM, CMU, TNS.

ASIA. India, Malaysia, Sri Lanka.

Catillaria A. Massal.; Catillariaceae

Catillaria testaceolivens (Vain.) Zahlbr., Cat. Lich. Univ. 4:
79 (1927).

Lecidea testaceolivens Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 173; 1909: 117,
both as Lecidea testaceolivens).

SPECIMEN. TUR-V (holotype).

ASIA. —

Catillaria unicolor (Vain.) Zahlbr., Cat. Lich. Univ. 4: 83
(1927).

Lecidea unicolor Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 173; 1909: 118,
both as Lecidea unicolor).

SPECIMEN. TUR-V (holotype).

ASIA. —

Cetrariopsis Kurok.; Parmeliaceae

Cetrariopsis wallichiana (Taylor) Kurok., Mem. Nat. Sci.
Mus., Tokyo 13: 140 (1980).

Cetraria wallichiana (Taylor) Miill. Arg.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 36, as Ce-
traria wallichiana), Kurokawa (1980: 140), Wolseley &
Aguirre-Hudson (1995: 318; 1997a: 333).

SPECIMENS. BM, PSU, TNS.

ASIA. China, Indonesia, Nepal, Philippines, Taiwan.

Cetrelia W.L. Culb.& C.F. Culb.; Parmeliaceae

Cetrelia braunsiana (Mill. Arg.) W.L. Culb. & C.F. Culb.,
Contr. U.S. Natl. Herb. 34: 493 (1968).

Cetraria collata f. isidiata Asahina.

THAILAND. Chiang Mai (Doi Inthanon N.P.). Sato (1962: 3, as
Cetraria collata f. isidiata), Wolseley & Aguirre-Hudson (1997a:

353:93')):
SPECIMENS. BM, TNS.

ASIA.
Taiwan.

China, India, Malaysia, Papua New Guinea, Philippines,

Cetrelia japonica (Zahlbr.) W.L. Culb. & C.F. Culb., Contr.
U.S. Natl. Herb. 34: 511 (1968).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Nayok (Khao
Yai N.P.). Wolseley & Aguirre-Hudson (1997a: 333), Boonpragob

et al. (1998: 216).
SPECIMENS. BM, RAMK.

ASIA. Indonesia, Malaysia, Taiwan.

Cetrelia nuda (Hue) W.L. Culb. & C.F. Culb., Contr. U.S.
Natl. Herb. 34: 513 (1968).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1997a: 337, 339).

SPECIMEN. BM.

ASIA. China, Taiwan.

Cetrelia olivetorum (Nyl.) W.L. Culb. & C.F. Culb, Contr.
U.S. Natl. Herb. 34: 515 (1968).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1997a: 337, 339).

SPECIMEN. BM.

ASIA. China, India, Papua New Guinea, Taiwan.

Chaenotheca (Th. Fr.) Th. Fr.; Coniocybaceae
Chaenotheca trichialis (Ach.) Th. Fr., Nova Acta Regiae
Soc. Sci. Upsal., ser. 3, 3: 251 (1860).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 216).

SPECIMEN. RAMK.

ASIA. — India.

Chiodecton Ach.; Roccellaceae
Chiodecton congestulum Nyl., Bull. Soc. Linn.
Normandie, sér. 2, 2: 106 (1868).

THAILAND. Trat (Koh Chang I.). Thor (1990: 41).
SPECIMENS. C, TUR-V.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Taiwan.

20

Chiodecton subsphaerale Nyl., Bull. Soc. Linn.
Normandie, sér. 2, 2: 106 (1868).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 138).
SPECIMENS. TUR-V.
ASIA. Malaysia.

Chroodiscus Miill. Arg.; Thelotremataceae
Chroodiscus mirificus (Kremp.) R. Sant., Symb. Bot.
Upsal. 12(1): 311 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka, Vietnam.

Chrysothrix Mont.; Chrysothricaceae

Chrysothrix candelaris (L.) Laundon, Lichenologist 13:
110 (1981)

Lepraria xanthina Vain.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Trat (Koh Chang I.), Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Vainio (1909: 151, as Lepraria xanthina), Wolseley &

Aguirre-Hudson (1997b: 355).
SPECIMENS. BM, TUR-V.

ASIA. China (Hong Kong), India, Philippines, Sri Lanka.

Cladia Ny1; Cladoniaceae
Cladia aggregata (Sw.) Nyl., Compt. Rend. Hebd. Séances
Acad. Sci. 83: 8 (1876).

THAILAND. Loei (Phu Miang, Phu Kradung N.P.). Yoshimura
(1978: 37).

SPECIMENS. AAU, C.

ASIA. China (Hong Kong), India, Indonesia, Malaysia, Nepal,
Papua New Guinea, Philippines, Taiwan.

Cladonia P. Browne; Cladoniaceae

Cladonia calyciformis Nuno, J. Jap. Bot. 47: 161 (1972).

THAILAND. Loei ( Phu Kradung N.P.). Yoshimura (1978: 37).
SPECIMENS. C, TNS (holotype).
ASIA. —

Cladonia ciliata var. tenuis (Florke) Ahti, Ann. Bot. Fenn.
15: 8 (1978).

Cladonia tenuis (Florke) Harm.

THAILAND. Phu Mieng. Yoshimura (1978: 37, as Cladonia tenuis).
AAU, C.

China, India.

SPECIMENS.

ASIA.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Cladonia didyma var. vulcanica (Zoll. & Moritzi) Vain.,
Acta Soc. Fauna Fl. Fenn. 4: 145 (1887).

Cladonia vulcanica Zoll.

THAILAND. Chiang Mai (Doi Inthanon N.P.), Loei (Phu Kradung
N.P.). des Abbayes (1956: 259, as Cladonia vulcanica), Yoshimura

(1978: 38, as Cladonia vulcanica).
SPECIMENS. TNS.

ASIA.
Taiwan.

India, Indonesia, Papua New Guinea, Philippines, Sri Lanka,

Cladonia furcata (Huds.) Schrad., Spicil. Fl. Germ.: 107
(1794).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. China, India, Indonesia, Taiwan.

Cladonia gymnopoda Vain., Acta Soc. Fauna Fl. Fenn. 10:
172 (1894).

THAILAND. Loei (Phu Miang). Yoshimura (1978: 38).
SPECIMENS. AAU, C.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Taiwan.

Cladonia modesta Ahti & Krog, Ann. Bot. Fenn. 24: 88
(1987).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 216).
SPECIMENS. RAMK.

ASIA. —

Cladonia ramulosa (With.) Laundon, Lichenologist 6: 225
(1984).

Cladonia pityrea (Florke) Fr.

THAILAND. Chiang Mai (Doi Suthep N.P.), Khu Kawng, Loei
(Phu Luang N.P.). Vainio (1921: 46, as C. pityrea), Yoshimura
(1978: 38, as C. pityrea).

SPECIMENS. C.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Philippines, Sri Lanka.

Cladonia rangiferina (L.) F.H. Wigg., Primit. Fl. Holsat.
90: 29 (1780).

THAILAND. Loei (Phu Kradung N.P.). Yoshimura (1978: 37),
Kurokawa & Kashiwadani (1988: Lich. Rar. Crit. Exs. No. 60).

C;: TNS:

ASIA. China, India, Taiwan.

SPECIMENS.

Cladonia rappii A. Evans, Trans. Connecticut Acad. Arts
38: 297 (1952).

Cladonia calycantha Delise ex Nyl.

CATALOGUE OF THE LICHENS OF THAILAND

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Phu Kradung
N.P.). Yoshimura (1978: 37, as C. calycantha).

SPECIMENS. AAU, BM, C.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea,
Philippines, Taiwan.

Cladonia siamea des Abbayes, Kew Bull. 2: 262 (1956).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Loei (Phu Kradung N.P.). Yoshimura (1978: 37), Huovinen & Ahti
(1986: 182), Stenroos (1986: 164), Kurokawa & Kashiwadani (1988:
Lich. Rar. Crit. Exs. Nos. 61, 62).

SPECIMENS. BM (holotype), C, H (isotype), RAMK, TNS.

ASIA. Indonesia, Papua New Guinea.

Cladonia siamea f. evoluta des Abbayes, Kew Bull. 2: 262
(1956).

THAILAND. Loei (Phu Kradung N.P.). des Abbayes (1956: 262).
SPECIMENS. BKE, BM (holotype).

ASIA. —

Cladonia siamea f. pulvinata des Abbayes, Kew Bull. 2:
263 (1956).

THAILAND. Loei (Phu Kradung N.P). des Abbayes (1956: 263).
SPECIMENS. BKF, BM (holotype).

ASIA, —

raat squamosa (Scop.) Hoffm., Deutschl. Fl. 2: 125

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 216).

SPECIMENS. RAMK.

ASIA. China, India, Indonesia, Papua New Guinea, Philippines,
Taiwan.

Cladonia subpityrea Sandst., in Zahlbruckner, Ann.
Naturhist. Mus. Wien 42: 61 (1928).

THAILAND. Loei (Dan Sai). Yoshimura (1978: 38).

SPECIMENS. C.

ASIA. Papua New Guinea, Philippines, Taiwan.

Cladonia transindica Ahti, Ann. Bot. Soc. Zool-Bot. Fenn.
‘Vanamo’ 32(1): 72 (1961).

THAILAND. locality not known. des Abbayes (1964), Ahti (1984:
44), Stenroos (1988: 212).

SPECIMEN. ?

ASIA. China, Indonesia, Malaysia, Vietnam.

Coccocarpia Pers.; Coccocarpiaceae

Coccocarpia dissecta Swinscow & Krog, Norweg. J. Bot.
23: 254 (1976).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden, Doi

21

Suthep N.P., Doi Inthanon N.P.), Khiri Khan, Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Mongkonsuk et al. (1996: 16).

SPECIMENS. BM, Queen Sirikit Botanical Garden Herbarium.

ASIA. Malaysia, Papua New Guinea, Philippines.

Coccocarpia erythroxyli (Spreng.) Swinscow & Krog,
Norweg. J. Bot. 23: 256 (1976).

Coccocarpia molybdaea Pers.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Nakhon Ratchasima (Khao Yai N.P.), Trat (Koh Chang I.), Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Paulson (1930: 99,
as C. molybdaea), Arvidsson (1982: 62), Boonpragob et al. (1998:

216).
SPECIMENS. BM, GB, L, RAMK, TNS.

ASIA. China (incl. Hong Kong), India, Malaysia, Myanmar, Papua
New Guinea, Philippines, Singapore, Sri Lanka, Taiwan.

Coccocarpia glaucina Kremp., Nuovo Giorn. Bot. Ital. 7:
14 (1875).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Nayok (Khao
Yai N.P.). Boonpragob et al. (1998: 216).

SPECIMENS. BM, RAMK.

ASIA. Malaysia, Papua New Guinea, Philippines.

Coccocarpia palmicola (Spreng.) Arv. & D.J. Galloway,
Bot. Not. 132: 242 (1979).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden, Doi
Suthep N.P., Doi Inthanon N.P., Chiang Dao N.P.), Nakhon Nayok
(Khao Yai N.P.), Tak (Ton Krabakyai N.P.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Arvidsson (1982: 76), Wolseley &
Aguirre-Hudson (1997a: 333; 1997b: 355), Mongkonsuk et al.
(1996), Boonpragob et al. (1998: 216).

SPECIMENS. BM, L, RAMK, TNS, Queen Sirikit Botanical Gar-
den Herbarium.

ASIA. China (incl. Hong Kong), India, Malaysia, Myanmar, Papua
New Guinea, Philippines, Singapore, Sri Lanka, Taiwan.

Coccocarpia pellita (Ach.) Mill. Arg. emend. R. Sant.,
Symb. Bot. Upsal. 12(1): 420 (1952).

THAILAND. Nakhon Nayok (Khao Yai N.P.), Uthai Thani (Huay

Kha Khaeng Wildlife Sanctuary). Boonpragob et al. (1998: 216).
SPECIMENS. BM, RAMK.

ASIA. China (Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Philippines, Sri Lanka.

Collema F.H. Wigg.; Collemataceae

Collema coccophyllum Nyl., Syn. Meth. Lich. 1: 112
(1858).

THAILAND. Chiang Mai (Doi Suthep). Degelius (1974: 69).
SPECIMEN. TNS.

ASIA. India.

22

Collema coilocarpum (Miill. Arg.) Zahlbr., Cat. Lich.
Univ. 3: 34 (1924).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary). Wolseley & Aguirre-Hudson (1997a: 333), Boonpragob et al.

(1998: 217).
SPECIMENS. BM, RAMK.

ASIA. India, Papua New Guinea, Philippines, Sri Lanka.

Conotrema Tuck.; Stictidaceae
Conotrema lumbricoides Sipman, in Aptroot et al.,

Biblioth. Lichenol. 64: 52 (1997).

THAILAND. Nakhon Sri Thammarat (Khao Luang N.P.).

Vongshewarat et al. (1999: 227)
SPECIMENS. RAMK.

ASIA. Papua New Guinea.

Cresponea Egea & Torrente; Roccellaceae
Cresponea chloroconia (Tuck.) Egea & Torrente,

Mycotaxon 48: 310 (1993).

Lecanactis deminuens (Nyl.) Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 121, as Lecanactis

deminuens).
SPECIMEN. TUR-V.

ASIA. —

Crocynia A. Massal.; Lecideaceae

Crocynia pyxinoides Nyl., Sert. Lich. Trop. Labuan
Singapore: 37 (1891).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson (1995:

320; 1997a: 333, 339; 1997b: 356), Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. Malaysia, Singapore.

Cryptothecia Stirt.; Arthoniaceae

Cryptothecia candida (Kremp.) R. Sant., Symb. Bot. Upsal.
121); 63: (1952):

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMEN. RAMK.

ASIA. India, Indonesia, Malaysia, Sri Lanka.

Cryptothecia obtecta Patw. & Sethy, Biovigyanam 13(2):
46 (1987).

THAILAND. Northern Thailand. Wolseley (1991: 240).
SPECIMEN. BM.

ASIA. — India.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Dichosporidium Pat.; Chiodectonaceae

Dichosporidium boschianum (Mont.) Thor, Opera Bot.
103: 64 (1990).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. RAMK.

ASIA. China (Hong Kong), India, Malaysia, Philippines, Singa-
pore, Sri Lanka.

Dictyonema C. Agardh ex Kunth; Meruliaceae

Dictyonema sericeum (Sw.) Berk., J. Bot. (London) 2: 639
(1843).

Dictyonema sericeum f. thelephora (Spreng.) Parm.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 151), Parmasto
(1978: 110, 111).
SPECIMEN. TUR-V.

ASIA. China (incl. Hong Kong), Indonesia, Papua New Guinea,
Philippines, Taiwan.

Dirinaria (Tuck.) Clem.; Physciaceae
Dirinaria aegialita (Ach.) B. Moore, Bryologist 71: 248
(1968).

THAILAND. Chiang Mai ( Doi Inthanon N.P., Chiang Dao N.P.),
Lampang (Doi Khun Tan N.P., Doi Pae Luang N.P.). Wolseley &
Aguirre-Hudson (1997a: 333).

SPECIMENS. BM, CMU.

AsIA. China (Hong Kong), India, Indonesia, Malaysia, Philip-
pines, Sri Lanka, Taiwan, Vietnam.

Dirinaria applanata (Fée) D.D. Awasthi, in Awasthi &
Agarwal, J. Indian Bot. Soc. 49: 135 (1970).

THAILAND. Khiri Khan, Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Wolseley & Aguirre-Hudson (1997a: 333).

SPECIMENS. BM.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Philippines, Sri Lanka, Taiwan, Vietnam.

Dirinaria confusa D.D. Awasthi, Biblioth. Lichenol. 2: 56
(1975).

THAILAND. Fang (northern Thailand). Yoshimura (1978: 35).
SPECIMEN. C.

ASIA. China, Papua New Guinea, Taiwan.

Dirinaria consimilis (Stirt.) D.D. Awasthi, in Awasthi &
Agarwal, J. Indian Bot. Soc. 49: 135 (1970).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Sara Buri, Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Awasthi (1975: 93), Wolseley & Aguirre-
Hudson (1995: 321; 1997a: 333; 1997b: 355).

BM, CMU.

India, Indonesia, Malaysia, Myanmar, Vietnam.

SPECIMENS.

ASIA.

CATALOGUE OF THE LICHENS OF THAILAND

Dirinaria picta (Sw.) Clem. & Shear, Gen. Fung.: 323
(1931).

Physcia picta Nyl.

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Khiri
Khan, Trat (Koh Chang I.). Vainio (1909: 112, as Physcia picta),
Paulson (1930: 100, as P. picta), Mongkonsuk et al. (1996), Wolseley
& Aguirre-Hudson (1995: 321).

SPECIMENS. BM, TUR-V, Herbarium Queen Sirikit Botanical

Garden.

ASIA. China (Hong Kong), India, Indonesia, Malaysia, Myanmar,
Papua New Guinea, Philippines, Singapore, Sri Lanka, Taiwan,
Vietnam.

Echinoplaca Fée; Gomphillaceae

Echinoplaca epiphylla Fée, Essai Crypt. Ecorc. Offic.: 93
(1824).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. RAMK.

ASIA. China (Hong Kong), India, Indonesia, Papua New Guinea,
Philippines, Vietnam.

Echinoplaca pellicula (Miill. Arg.) R. Sant., Symb. Bot.
Upsal. 12(1): 367 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. RAMK.

ASIA. China, Indonesia, Papua New Guinea, Philippines, Sri Lanka,
Vietnam.

Endocarpon Hedwig; Verrucariaceae

Endocarpon adscendens Miill. Arg., Bull. Trav. Soc.
Murith. Valais 10: 58 (1881).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. CANB.
ASIA. —

Enterographa Fée; Chiodectonaceae

[Enterographa] Chiodecton pallidellum var. olivaceoalba
Vain., Bot. Tidsskr. 29: 138 (1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 138), Thor (1990:
86).

SPECIMEN. 'TUR-V (not located).
ASIA. —

[Enterographa\ Chiodecton pallidellum vac.
olivaceostraminea Vain., Bot. Tidsskr. 29: 137 (1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 137), Thor (1990:
86).

SPECIMEN. TUR-V (holotype).

ASIA. —

23

Eremothecella Syd.; Arthoniaceae

Eremothecella macrocephala (R. Sant.) Thor, Liicking &
Matsumoto, Symb. Bot. Upsal. 32(3): 39 (2000).

Stirtonia macrocephala R. Sant.

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217, as Stirtonia macrocephala).

SPECIMENS. RAMK.

ASIA. Indonesia, Malaysia, Papua New Guinea, Philippines.

Erioderma Fée; Pannariaceae

Erioderma sorediatum D.J. Galloway & P.M. Jorg.,
Lichenologist 7: 139 (1975).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Nayok (Khao
Yai N.P.). Wolseley & Aguirre-Hudson (1995: 321; 1997a: 339),
Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK.

ASIA. Malaysia, Papua New Guinea, Sri Lanka.
Eschatogonia Trevis., ?Acarosporaceae

Eschatogonia prolifera (Mont.) R. Sant., in Swinscow &
Krog, Macrolichens East Africa: 86 (1988).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Lampang (Wiang
Kosai N.P.), Nakhon Ratchasima (Khao Yai N.P.), Satun (Thaleban
N.P.), Uthai Thani (Huay Kha Kaeng Wildlife Sanctuary).
Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK.

ASIA. —
Everniastrum Hale ex Sipman; Parmeliaceae

Everniastrum cirrhatum (Fr.) Hale ex Sipman, Mycotaxon
26: 237 (1986).

Parmelia cirrhata Fr.
Cetrariastrum cirrhatum (Fr.) W.L. Culb. & C.F. Culb.
Parmelia vulgaris Vain.

THAILAND. Chiang Mai (Doi Suthep, Doi Inthanon, Chiang Dao
N.P.), Loei (Phu Luang). Vainio (1921: 38, as Parmelia vulgaris),
Sato (1962: 3, as P. cirrhata), Moon et al. (2000a: 101, as
Cetrariastrum cirrhatum).

SPECIMENS. BM, CMU, TNS, TUR-V.

ASIA. China, India, Indonesia, Nepal, Sri Lanka, Taiwan.

Everniastrum nepalense (Taylor) Hale ex Sipman,
Mycotaxon 26: 239 (1986).

Cetrariastrum nepalense (Taylor) W.L. Culb. & C.F. Culb.
Parmelia americana (Meyen & Flotow) Mont.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P., Maetang), Yala (Banglang N.P.). Vainio (1921: 38,
as Parmelia americana), Sato (1962: 6, as P. americana), Wolseley
& Aguirre-Hudson (1995: 321; 1997a: 333), Moon et al. (2000a:
101, as Cetrariastrum nepalense).

24

SPECIMENS. BM, CMU, PSU, RAMK, TNS, Kurokawa, Lich.
Rar. Crit. Exs. No. 78.

ASIA. China, India, Indonesia, Nepal.

Everniastrum scabridum Elix & Pooprang, in Pooprang et
al., Mycotaxon 71: 112 (1999).

THAILAND. Chiang Mai (Doi Suthep N.P.). Pooprang et al. (1999:

i A
SPECIMENS. BM, CMU, RAMK (holotype).

ASIA. —

Everniastrum sorocheilum (Vain.) Hale ex Sipman,
Mycotaxon 26: 242 (1986).

Parmelia sorocheila Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Sato (1962: 4, as

Parmelia sorocheila Vain.).
SPECIMEN. OSA.

ASIA. China, Indonesia, Philippines, Taiwan.

Everniastrum vexans (Zahlbr. ex W.L. Culb. & C.F. Culb.)
Hale ex Sipman, Mycotaxon 26: 242 (1986).

Cetrariastrum vexans Zahlbr. ex W.L. Culb. & C.F. Culb.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Loei (Phu Luang N.P.). Wolseley & Aguirre-Hudson (1995: 318;

1997a: 333), Moon et al. (2000a: 101, as Cetrariastrum vexans).
SPECIMENS. BM, TNS.

ASIA. China, India, Indonesia, Papua New Guinea, Philippines,
Taiwan.

Fuscopannaria P.M. Jg@rg.; Pannariaceae

Fuscopannaria siamensis P.M. Jérg. & Wolseley, in
Jegrgensen, J. Hattori Bot. Lab. 89: 256 (2000).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Jorgensen (2000:

256).
SPECIMENS. BM (holotype).

ASIA. —

Glyphis Ach.; Graphidaceae

Glyphis cicatricosa var. confluens (Zenker) Zahlbr.,
Denkschr. Kaiserl. Akad. Wiss., Wien Math.-Naturwiss. Cl.
83: 112 (1909).

Graphis cicatricosa var. confluens (Zenker) Vain.

THAILAND. Trat (Koh Chang I.), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Vainio (1909: 136, as Graphis cicatricosa var.

confluens).
SPECIMENS. BM, TUR-V.

ASIA. Vietnam.
Graphidastra (Redinger) Thor; Chiodectonaceae

Graphidastra byssiseda (Miill. Arg.) Thor, Opera Bot. 103:
80 (1990).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Wolseley & Aguirre-Hudson (1997: 333).

SPECIMENS. BM.

ASIA. India, Indonesia, Nepal.

Graphina Miill. Arg.; Graphidaceae

Graphina boschiana Mill. Arg., Flora 65: 385 (1882).
Graphis concolor Ny\.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 128, as Graphis

concolor).
SPECIMENS. TUR-V.

ASIA. India, Indonesia, Sri Lanka.

Graphina columbiana (Ny1.) Zahlbr., Cat. Lich. Univ. 2:
402 (1924).
Graphis obtecta var. columbiana Nyl.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 51, as
Graphis obtecta var. columbiana).

SPECIMEN. TUR-V.

ASIA. India.

Graphina consimilis (Vain.) Zahlbr., Cat. Lich. Univ. 2:
402 (1923).
Graphis consimilis Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 177; 1909: 127,
both as Graphis consimilis).

SPECIMEN. TUR-V (holotype).

ASIA. Philippines.

Graphina fissurinoidea (Vain.) Zahlbr., Cat. Lich. Univ. 2:
406 (1923).

Graphis fissurinoidea (Ny1.) Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 128, as Graphis
fissurinoidea).

SPECIMEN. TUR-V.

ASIA. —

Graphina glaucorufa (Vain.) Zahlbr., Cat. Lich. Univ. 2:
407 (1923).

Graphis glaucorufa Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 51, as
Graphis glaucorufa).

SPECIMEN. TUR-V (holotype).

ASIA. —

Graphina intermedians (Vain.) Zahlbr., Cat. Lich. Univ. 2:
411 (1923).

Graphis intermedians Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 50, as
Graphis intermedians).

CATALOGUE OF THE LICHENS OF THAILAND

SPECIMEN. TUR-V (holotype).

ASIA. —

Graphina intricata (Eschw.) Miill. Arg., Flora 71: 510
(1888).

Graphis subintricata Kremp.
Graphis extricata Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 126, as Graphis
subintricata and G. extricata).

SPECIMENS. TUR-V.

ASIA. Sri Lanka.

Graphina ruiziana var. gracilior Miill. Arg., Flora 65: 385
(1882).

Graphis ruiziana var. gracilior Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 125, as Graphis

ruiziana vat. gracilior).
SPECIMENS. TUR-V.

ASIA. Indonesia.

Graphina simplex (Vain.) Zahlbr., Cat. Lich. Univ. 2: 425
(1923).
Graphis simplex Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1907: 127 as Graphis
simplex).

SPECIMEN. TUR-V (holotype).

ASIA. —

Graphina streblocarpa var. pauperior Vain., Suom. Elain-
Ja Kasvit. Seuran Van. Julk. 1(3): 50 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 50).
SPECIMEN. TUR-V (holotype).

ASIA. —

Graphina subobtecta var. feracior (Vain.) Zahlbr., Cat.
Lich. Univ. 2: 427 (1923).

Graphis subobtecta var. feracior Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 51, as
Graphis subobtecta var. feracior).

SPECIMEN. TUR-V (holotype).

ASIA. —
Graphis Adans.; Graphidaceae

Graphis afzelii Ach., Syn. Meth. Lich.: 85 (1814).

THAILAND. Chiang Mai (Baw Luang), Nakhon Ratchasima (Khao
Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).

Boonpragob et al. (1998: 217).
SPECIMENS. RAMK, BM.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Sin-
gapore, Sri Lanka, Vietnam.

Za
Graphis albonitens Miill. Arg., Hedwigia 30: 53 (1891)
THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99).
SPECIMEN. BM.

ASIA. —

Graphis glaucocinerea Vain., Hedwigia 46: 178 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1907: 178; 1909: 135).
SPECIMENS. TUR-V, FH (both syntypes).

ASIA. —

Graphis hossei Vain., Suom. Elain-ja Kasvit. Seuran Van.
Julk. 1(3): 54 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 54),
Patwardhan & Kulkarni (1976: 127).

SPECIMEN. TUR-V (holotype).

ASIA. — India.

Graphis leptospora Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 53 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 53).
SPECIMENS. TUR-V (holotype).

ASIA. —

Graphis lineola Ach., Lichenogr. Universalis: 264 (1810).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 134).

SPECIMEN. TUR-V.

ASIA. China (incl. Hong Kong), India, Indonesia, Papua New
Guinea, Philippines.

Graphis ochrocheila Vain., Hedwigia 46: 178 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1907: 178; 1909: 135)
SPECIMEN. TUR-V (holotype).

ASIA. —

Graphis pyrrhocheiloides Zahlbr., Cat. Lich. Univ. 2: 321
(1923).

Graphis pyrrhocheila Vain.

THAILAND. Trat (Lem Ngob). Vainio (1907: 179; 1909: 136, both

as Graphis pyrrhocheila).
SPECIMEN. TUR-V (holotype).

ASIA. India, Indonesia.

Graphis sauroidea Leight., Trans. Linn. Soc. London 25:
452 (1866) .

THAILAND. Trat (Koh Chang I.). Vainio (1909: 133).
SPECIMENS. TUR-V.
ASIA. —

Graphis siamensis Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 52 (1921).

26

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 52).
SPECIMENS. TUR-V (holotype).
ASIA. —

Graphis striatula (Ach.) Spreng., Syst. Veg., 16th ed, 4:

250 (1827).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 134).
SPECIMENS. TUR-V.

ASIA. China, India, Indonesia, Malaysia, Sri Lanka.

Graphis subdisserpens Nyl., Bull. Soc. Linn. Normandie,
sér. 2, 7: 16 (1874).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 132).
SPECIMENS. TUR-V.
ASIA. — India.

Graphis subdisserpens f. arbusculaeformis Vain., Bot.
Tidsskr. 29: 132 (1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 132).
SPECIMENS. TUR-V (holotype).

ASIA. —

Graphis tenella Ach., Syn. Meth. Lich.: 81 (1814).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1909: 133;
1921: 54).
SPECIMENS. TUR-V.

ASIA. China, India, Indonesia, Sri Lanka.

Graphis tenella var. epiphaea Vain., Bot. Tidsskr. 29: 133
(1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 133).
SPECIMENS. TUR-V (holotype).

ASIA. —

Graphis tenella var. leptocarpoides Vain., Bot. Tidsskr. 29:
133 (1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 133).
SPECIMENS. TUR-V (syntypes).
ASIA. —

Graphis tenuis Vain., Suom. Elain-ja Kasvit. Seuran Van.
sulk. 1(3):53:(1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 53).
SPECIMENS. TUR-V (syntypes).
ASIA. —

Graphis trichospora Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 52 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 52).
SPECIMEN. TUR-V (holotype).

ASIA. —

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Gyalideopsis Veézda; Gomphillaceae
Gyalideopsis rubescens Veezda, Folia Geobot. Phytotax.
14: 67 (1979).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Wolseley & Aguirre-Hudson (1997a: 333),

Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. —

Gymnoderma Ny|I.; Cladoniaceae

Gymnoderma coccocarpum Ny|l., Syn. Meth. Lich. 2: 26
(1863).

THAILAND. Nakhon Nayok. Yoshimura & Sharp (1968: 635-
640), Hawksworth & Yoshimura (1973: 503), Yoshimura (1982:
84), Kashiwadani & Gradstein (1982: 80).

SPECIMENS. US.

ASIA. China, India, Indonesia, Malaysia, Taiwan.
Gyrostomum Fr.; Thelotremataceae

Gyrostomum scyphuliferum (Ach.) Nyl., Ann. Sci. Nat.
Bot., sér. 4, 16: 96 (1862).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Trat (Koh Chang I.). Vainio (1909: 121), Wolseley
& Aguirre-Hudson (1997a: 333), Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK, TUR-V.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Phil-
ippines, Taiwan, Vietnam.

Haematomma A. Massal; Haematommataceae

Haematomma collatum (Stirt.) C.W. Dodge, Beih. Nova
Hedwigia 38: 41 (1971).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Lampang (Doi Khun Tan N.P.), Uthai Thani (Khao Nang Ram
Wildlife Sanctuary). Staiger & Kalb (1995: 89).

SPECIMENS. BKF, BM, CMU, COLO.

ASIA. India, Indonesia, Papua New Guinea, Sri Lanka.

Haematomma puniceum (Ach.) A. Massal., Atti Reale Ist.
Veneto, Sci. Lett. Arti, ser. 3, 5: 253 (1860).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Uthai Thani (Khao Nang Ram Wildlife Sanctuary). Vainio (1921:
43), Wolseley & Aguirre-Hudson (1997a: 339).

SPECIMENS. BKF, BM, CMU, TUR-V.

ASIA. China, India, Indonesia, Singapore, Sri Lanka, Taiwan,
Vietnam.

Haematomma wattii (Stirt.) Zahlbr., Cat. Lich. Univ. 5:
776 (1928).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.).
Staiger & Kalb (1995: 89), Wolseley & Aguirre-Hudson (1997a:
3535: 309):

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. BM, CMU.

ASIA.
pines.

India, Indonesia, Malaysia, Papua New Guinea, Philip-

Hafellia Kalb, H. Mayrhofer & Scheidegger;
Physciaceae

Hafellia disciformis (Fr.) Marbach & H. Mayrhofer, in
Marbach, Biblioth. Lichenol. 74: 259 (2000).
Buellia disciformis (Fr.) Mudd

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 45, as

Buellia disciformis).
SPECIMENS. TUR-V.

ASIA. China, India, Indonesia, Vietnam.
Heterodermia Trevis.; Physciaceae
Heterodermia albicans (Pers.) Swinscow & Krog,
Lichenologist 8: 113 (1976).

Physcia crispa var. mollescens (Nyl.) Vain.

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Trat
(Koh Chang I.). Vainio (1909: 111, as Physcia crispa var. mollescens),
Mongkonsuk et al. (1996: 17).

SPECIMENS. TUR-V, Queen Sirikit Botanical Garden Herbarium.
ASIA. China, Malaysia, Philippines.

Heterodermia dendritica (Pers.) Poelt, Nova Hedwigia 9:
31 (1965).

Anaptychia dendritica Pers.

THAILAND. Hase Plantation, probably Laos. Kurokawa (1962: 54;
1973: 602, both as Anaptychia dendritica).

SPECIMEN. TNS.

ASIA. China, India, Indonesia, Papua New Guinea, Philippines,
Taiwan.

Heterodermia diademata (Taylor) D.D. Awasthi,
Geophytology 3: 113 (1973).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.,
Chiang Dao N.P.). Wolseley & Aguirre-Hudson (1997b: 356).

SPECIMENS. BM.

ASIA. India, China, Malaysia, Papua New Guinea, Sri Lanka.

Heterodermia esorediata (Vain.) Du Rietz & Lynge, in
Lynge, Videnskaps. I. Mat.-Naturv. Kl. 16: 14 (1924).

Anaptychia esorediata Vain.

THAILAND. Locality not given. Kurokawa (1959: 179, as
Anaptychia esorediata).

SPECIMEN. TNS (not seen).

ASIA. India, Indonesia, Philippines.

Heterodermia fragilissima (Kurok.) J.-C. Wei & Y.-M.
Jiang, Lich. Xizang: 111 (1986).

zy

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1997a: 333).

SPECIMENS. BM.

ASIA. China.

Heterodermia hypocaesia (Yasuda) D.D. Awasthi,
Geophytology 3: 113 (1973).

Anaptychia hypocaesia Yasuda

THAILAND. Chiang Mai (Doi Suthep N.P., Maetang District), Loei
(Phu Kradung N.P.). Kurokawa (1973: 603, as Anaptychia

hypocaesia).
SPECIMENS. TNS.

ASIA. China, Indonesia, Papua New Guinea, Philippines, Taiwan.

Heterodermia hypoleuca (Ach.) Trevis., Atti Soc. Ital. Sci.
Nat. 11: 615 (1868).

Anaptychia hypoleuca (Ach.) A. Massal.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Queen Sirikit Botanical Garden), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Vainio (1921: 45, as Anaptychia hypoleuca),
Mongkonsuk et al. (1996: 17).

SPECIMENS. BM, TUR-V, Queen Sirikit Botanical Garden Her-
barium.
ASIA. China, India, Malaysia, Papua New Guinea, Sri Lanka,

Taiwan.

Heterodermia incana (Stirt.) D.D. Awasthi, Geophytology
2 Wai 73),

Anaptychia incana (Stirt.) Zahlbr.

THAILAND. Hase Plantation, probably Laos. Kurokawa (1962: 92,
as Anaptychia incana).

SPECIMEN. TNS.

ASIA. China, India, Indonesia, Taiwan.

Heterodermia leucomelos (L.) Poelt, Nova Hedwigia 9: 31
(1965).

Anaptychia leucomelaena (L.) A. Massal.
Anaptychia neoleucomelaena Kurok.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Trat (Koh Chang I., Lem Dan). Vainio (1921: 44, as Anaptychia
leucomelaena), Kurokawa (1961: 51, as A. neoleucomelaena),
Wolseley & Aguirre-Hudson (1997a: 333).

SPECIMENS. BM, TUR-V.

ASIA. China, India, Indonesia, Malaysia, Myanmar, Papua New
Guinea, Philippines, Sri Lanka, Taiwan.

Heterodermia microphylla (Kurok.) Skorepa, Bryologist
75: 490 (1972).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Nayok (Khao

Yai N.P.). Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. China, India, Papua New Guinea.

28

Heterodermia pandurata (Kurok.) J.-C. Wei, Enum. Lich.
China: 112 (1991).

Anaptychia pandurata Kurok.

THAILAND.
95-96).

Hase Plantation, probably Laos. Kurokawa (1962:

SPECIMENS. TNS.

ASIA. Taiwan.

Heterodermia speciosa (Wulfen) Trevis., Atti Soc. Ital. Sci.
Nat. 11: 614 (1868).
Anaptychia speciosa (Wulfen) A. Massal.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.).

Vainio (1921: 45, as Anaptychia speciosa).
SPECIMENS. BM, TUR-V.

ASIA.
Lanka.

China (incl. Hong Kong), India, Indonesia, Malaysia, Sri

Hypogymnia (Nyl.) Nyl.; Hypogymniaceae
[Hypogymnia] Parmelia coilocarpa Vain., Suom. Elain-ja
Kasvit. Seuran Van. Julk. 1(3): 40 (1921), auct. non Stirt.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 40, as
Parmelia coilocarpa).

SPECIMENS. BM, TUR-V (syntype).

ASIA. —

Hypogymnia pseudobitteriana (D.D. Awasthi) D.D.
Awasthi, in Awasthi & Singh, Geophytology 1: 101 (1972).

THAILAND. Chiang Mai (Doi Suthep Pui N.P., Doi Inthanon N.P.).
Wolseley & Aguirre-Hudson (1995: 318).

SPECIMENS. BM.

ASIA. India, Malaysia, Papua New Guinea, Taiwan.

Hypogymnia subphysodes (Kremp.) Filson, Victorian Nat.
87: 327 (1970)
Parmelia subphysodes Kremp.

THAILAND. Chiang Mai (Doi Suthep, N.P.). Vainio (1921: 40, as
Parmelia subphysodes), Wolseley & Aguirre-Hudson (1997a: 333).

SPECIMENS. BM, TUR-V.

ASIA. —

Hypogymnia vittata (Ach.) Gas., Acta Soc. Linn. Bordeaux
53: 66 (1898).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1997a: 333, 337).

SPECIMENS. BM.

ASIA. China, India, Malaysia, Nepal, Taiwan.

Hypotrachyna ( Vain.) Hale; Parmeliaceae

Hypotrachyna adducta (Ny1.) Hale, Phytologia 28: 341
(1974).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

THAILAND. Chiang Mai (Doi Inthanon N.P.). Pooprang et al.

(1999: 119), Moon et al. (2000a: 101).
SPECIMENS. RAMK, TNS.

ASIA. China, India, Malaysia, Nepal, Philippines, Taiwan.

Hypotrachyna adjuncta (Hale) Hale, Phytologia 28: 341
(1974).

Parmelia exsecta Taylor.
Hypotrachyna exsecta (Taylor) Hale.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Loei (Phu Luang N.P), Phitsanulok. Sato (1962: 3, as Parmelia
exsecta) Kurokawa (1986: 265, as Hypotrachyna exsecta), Pooprang

et al. (1999: 120), Moon et al. (2000a: 101, and as H. exsecta).
SPECIMENS. BKF, BM, CMU, RAMK, TNS.

ASIA. India, Malaysia, Nepal, Philippines, Sri Lanka, Taiwan.

Hypotrachyna bahiana (Ny1.) Hale, Smithsonian Contr.
Bot. 25: 23 (1975):

Parmelia bahiana Nyl.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 38, as

Parmelia bahiana).
SPECIMEN. TUR-V.

ASIA. —

Hypotrachyna chlorobarbatica Elix & Pooprang, in
Pooprang, Boonpragob & Elix, Mycotaxon 72: 113 (1999).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Pooprang et al.

(19992 113).
SPECIMENS. CANB (isotype), RAMK (holotype).

ASIA. —

Hypotrachyna coorgiana Patw. & Prabhu, Bryologist 80:
348 (1997).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Phitsanulok

(Phu Hin Rongkla N.P.). Pooprang et al. (1999: 120).
SPECIMENS. RAMK.

ASIA. India.

Hypotrachyna crenata (Kurok.) Hale, Phytologia 28: 341
(1974).
Parmelia crenata Kurok.

THAILAND. Chiang Mai (Maetang District). Kurokawa (1988:

167-168, as Parmelia crenata), Moon et al. (2000a: 101).
SPECIMENS. TNS.

ASIA. India, Indonesia, Taiwan.

Hypotrachyna dactylifera (Vain.) Hale, Smithsonian Contr.
Bot. 25: 30 (1975).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).

Moon et al. (2000a: 101).
SPECIMENS. TNS.

ASIA. —

CATALOGUE OF THE LICHENS OF THAILAND

Hypotrachyna ducalis (Jatta) Hale, Smithsonian Contr.
Bot. 25: 33 (1975).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Wolseley & Aguirre-Hudson (1997a: 335),

Boonpragob et al. (1998: 217).
SPECIMENS. BKF, BM, CMU, RAMK.

ASIA. —

Hypotrachyna granulans K.H. Moon, Kurok. & Kashiw.,
Bull. Natn. Sci. Mus., Tokyo, ser. B, 26: 135 (2000).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Moon et al. (2000b:
135-137).

SPECIMENS. TNS (holotype), RAMK (isotype).

ASIA. —

Hypotrachyna imbricatula (Zahlbr.) Hale, Smithsonian
Contr. Bot. 25: 41 (1975).

Parmelia imbricatula Zahlbr.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Pacho, Maetang),
Loei (Ban Sui, Phu Kradung). Hale (1971: 16, as Parmelia

imbricatula), Moon et al. (2000a: 101).
SPECIMENS. TNS.

ASIA. India, Malaysia, Philippines, Taiwan.

Hypotrachyna immaculata (Kurok.) Hale, Smithsonian
Contr. Bot. 25: 41 (1975).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.). Pooprang et al.
(1999: 120).

SPECIMEN. RAMK.

ASIA. _ India.

Hypotrachyna infirma (Kurok.) Hale, Phytologia 28: 341
(1974).

Parmelia infirma Kurok.

THAILAND. Loei (Ban Sui, Phu Kradung). Hale & Kurokawa
(1964: 179, as Parmelia infirma), Awasthi (1977: 177), Moon et al.
(2000a: 101).

SPECIMEN. TNS.

ASIA. India, Nepal, Taiwan.

Hypotrachyna kingii (Hale) Hale, J. Jap. Bot. 43: 324
(1968).
Parmelia kingii Hale.

THAILAND. Nakhon Nayok (Khao Yai N.P.). Hale (1968: 324, as

Parmelia kingii), Moon et al. (2000a: 101).
SPECIMENS. TNS (isotype), US (holotype).

ASIA. Indonesia.

Hypotrachyna massartii (Hue) Hale, Phytologia 28: 341
(1974).

THAILAND. Loei (Phu Kradung). Moon et al. (2000a: 102).

29

SPECIMENS. TNS.

ASIA. —

Hypotrachyna masonhalei Patw. & Prabhu, Bryologist 80:
348 (1997).

THAILAND. Chiang Mai (Doi Suthep N.P.). Pooprang et al. (1999:
121).

SPECIMEN. RAMK.

ASIA. — India.
Hypotrachyna orientalis (Hale) Hale, Phytologia 28: 341
(1974).

Parmelia orientalis Hale.

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Nayok (Khao
Yai N.P.). Hale (1971: 435, as Parmelia orientalis Hale), Culberson
& Hale (1973: 166, as P. orientalis), Wolseley & Aguirre-Hudson
(1995: 318), Boonpragob et al. (1998: 217), Moon et al. (2000a:
102).

SPECIMENS.
(holotype).

BM, CMU, KF, TNS (isotype), UPS (isotype), US

ASIA. India, Malaysia, Papua New Guinea, Philippines.
Hypotrachyna osseoalba (Vain.) Y.S. Park & Hale, Taxon
38: 88 (1989).

Parmelia osseoalba Vain.
Hypotrachyna formosana auct. non (Zahlbr.) Hale.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Maetang), Loei (Ban Sui, Phu Kradung N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Vainio (1921: 39, as Parmelia osseoalba), Wolseley
& Aguirre-Hudson (1995: 318, as Hypotrachyna formosana),
Boonpragob et al. (1998: 217, as H. formosana), Moon et al. (2000a:
102).

SPECIMENS. BKF, BM, CMU, RAMK, TNS, TUR-V (holotype).
ASIA. China, India, Indonesia, Malaysia, Philippines, Taiwan.
Hypotrachyna physcioides (Ny1.) Hale, Smithsonian Contr.
Bot. 25: 54 (1975).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Pacho, Maetang),
Loei (Nahaew N.P., Phu Kradung). Pooprang et al. (1999: 120),

Moon et al. (2000a: 102).
SPECIMENS. RAMK, TNS.

ASIA. India, Malaysia, Papua New Guinea.

Hypotrachyna ramkhamhaengiana Elix & Pooprang, in
Pooprang, Boonpragob & Elix, Mycotaxon 72: 113 (1999).

THAILAND. Sakon Nakhon (Phu Pan N.P.). Pooprang et al. (1999:
11S):

SPECIMENS. RAMK (holotype).

ASIA. —

Hypotrachyna scytodes (Kurok.) Hale, Phytologia 28: 341
(1974).

Parmelia scytodes Kurok.

30

THAILAND. Chiang Mai (Maetang District). Culberson & Hale

(1973: 166, as Parmelia scytodes), Moon et al. (2000a: 102).
SPECIMENS. TNS.

ASIA. India, Malaysia.

Imshaugia S.L.F. Mey.; Parmeliaceae

Imshaugia placorodia (Ach.) S.L.F. Mey., Mycotaxon 35:
189 (1989).

Parmeliopsis placorodia (Ach.) Nyl.

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).

Mongkonsuk et al. (1996: 17, as Parmeliopsis placorodia).
SPECIMENS. Queen Sirikit Botanical Garden Herbarium.

ASIA. —

Ionaspis Th. Fr.; Hymeneliaceae

Tonaspis tropica Aptroot, in Aptroot et al., Biblioth.
Lichenol. 64: 73 (1997).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.

ASIA. Papua New Guinea.

Lasioloma R. Sant.; Ectolechiaceae

Lasioloma arachnoideum (Kremp.) R. Sant., Symb. Bot.
Upsal. 12(1): 547 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Vietnam.

Laurera Rchb.; Trypetheliaceae

Laurera benguelensis (Miill. Arg.) Zahlbr., Cat. Lich.
Univ. 1: 503 (1992).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Nakhon
Nayok (Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Mongkonsuk et al. (1996: 17), Wolseley & Aguirre-
Hudson (1997a: 339, 1997b: 356), Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK, Queen Sirikit Botanical Garden Her-

barium.

ASIA. — India.

Laurera keralensis D.K. Upreti & A. Singh, Bull. Jard.
Bot. Belg. 57: 374 (1987).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Sakon Nakhon

(Ratchapat Sakon). Vongshewarat et al. (1999: 227).
SPECIMENS. RAMK.

ASIA. — India.

Laurera megasperma (Mont.) Riddle, Bull. Torrey Bot.
Club 44: 323 (1917).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Vongshewarat

et al. (1999: 228).
SPECIMENS. RAMK.

ASIA. India, Papua New Guinea, Sri Lanka.

Laurera meristospora (Mont. & Bosch) Zahlbr., Cat. Lich.
Univ. 1: 505 (1922).

Thelenella interrupta Vain.
Clathroporina interrupta (Vain.) Zahlbr.

THAILAND. Trat (Koh Chang I.), Nakhon Ratchasima (Khao Yai
N.P.). Vainio (1907: 180, 1909: 149, both as Thelenella interrupta),
McCarthy (1995: 347, as Laurera aff. meristospora), Boonpragob et

al. (1998: 217).
SPECIMENS. RAMK, TUR-V (holotype of T: interrupta).

ASIA. Indonesia, Papua New Guinea.

Laurera meristosporoides P.M. McCarthy &
Vongshewarat, in Vongshewarat et al., Mycotaxon 71: 228
(1999).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.). Vongshewarat et

al. (1999: 228).
SPECIMENS. CANB (holotype).

ASIA. —

Laurera phaeomelodes (Miill. Arg.) Zahlbr., in Engler &
Prantl, Nat. Pflanzenfam. 1, 1: 71 (1903).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.). Vongshewarat et

al. (1999: 230).
SPECIMENS. RAMK.

ASIA. _ India.

Laurera subdiscreta (Nyl.) Zahlbr., Cat. Lich. Univ. 1: 506
(1922).

THAILAND. Buri Ram (Khao Kadong), Sakon Nakhon (Phu Pan

N.P.). Vongshewarat et al. (1999: 230).
SPECIMENS. RAMK.

ASIA. India.

Lecanactis K6rb.; Roccellaceae

Lecanactis flavisedella (Ny1.) Tehler, Willdenowia 22: 206
(1992).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson
(1997a: 333, 339).

SPECIMENS. BM.

ASIA. India.

Lecanora Ach.; Lecanoraceae

Lecanora achroella Nyl., J. Bot. 14: 263 (1876).
Lecanora monodorae Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 109, as Lecanora
monodorae).

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. TUR-V.

ASIA. —

Lecanora cinereocarnea (Eschw.) Vain., Etud. Class. Lich.
Brésil 1: 80 (1891).

THAILAND. Chiang Mai (Doi Suthep N.P.), Trat (Koh Chang I[.,
Lem Dan). Vainio (1909: 108; 1921: 41).

SPECIMENS. TUR-V.

ASIA. Philippines, Vietnam.

Lecanora fumigata Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 42 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 42).
SPECIMEN. TUR-V (holotype).

ASIA. —

Lecanora lividocarnea Vain., Philipp. J. Sci., sect. C, 8:
100 (1913).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 41).
SPECIMENS. TUR-V.

ASIA. Philippines.

Lecanora lividoglauca Vain., Suom. Elain-ja Kasvit.
Seuran Van. Julk. 1(3): 41 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 41).
SPECIMEN. TUR-V (holotype).

ASIA. —

Lecanora phaeocardia Vain., Suom. Elain-ja Kasvit.
Seuran Van. Julk. 1(3): 41 (1921).

Lecanora phaeocardia var. lividofuscescens Vain.
Lecanora phaeocardia var. subcrenulans Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.), Nong Boa, Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Vainio (1921: 41,
42), Lumbsch et al. (1996: 283-285).

SPECIMEN. TUR-V (holotype of Lecanora phaeocardia var.
lividofuscescens).

ASIA. India, Indonesia.

Lecanora pulicaris (Pers.) Ach., Syn. Meth. Lich.: 336
(1814).
Lecanora subfusca var. chlarona Ach.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 108, as Lecanora
subfusca var. chlarona).

SPECIMENS. TUR-V.

ASIA. China, India, Papua New Guinea, Philippines.
Lecanora subfusca Ach., Lichenogr. Universalis: 393
(1810).

THAILAND. Chiang Mai (Doi Suthep N.P.), Chumphon (Koh Tao
I.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Paulson
(1930: 99), Wolseley & Aguirre-Hudson (1997b: 356).

31
SPECIMENS. BM.

ASIA. China, Sri Lanka.

Lecanora subgranulata Nyl., Lich. Nov.-Zel.: 62 (1888).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 108).
SPECIMENS. TUR-V.

ASIA. —

Lecidea Ach.; Lecideaceae

Lecidea inolescens Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 48 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 48).
SPECIMENS. TUR-V (type not located).

ASIA. —

Lecidopyrenopsis Vain.; Lichinaceae
Lecidopyrenopsis corticola Vain., Hedwigia 46: 172
(1907).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 117), Zahlbruckner

(1926: 172), Henssen (1979: 485; 1989: 102).
SPECIMENS. C (isolectotype), TUR-V (lectotype).

ASIA. —

Leprocaulon Nyl.; Fungi Imperfecti

Leprocaulon arbuscula (Nyl.) Nyl., Lich. Ins. Guineensis:
8 (1889).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Nakhon Ratchasima (Khao Yai N.P.). Wolseley & Aguirre-Hudson

(1997a: 333), Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. China, India, Taiwan.
Leptogium (Ach.) S.F. Gray; Collemataceae

Leptogium asiaticum P.M. Jgrg., Herzogia 2: 466 (1973).

THAILAND. Chiang Mai (Doi Inthanon N.P., Chiang Dao N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &
Aguirre-Hudson (1997a: 333).

SPECIMENS. BM, CMU.

ASIA. China, India, Malaysia, Papua New Guinea, Sri Lanka.

Leptogium azureum (Sw. ex Ach.) Mont., in Webb &
Berthelot, Hist. Nat. Iles Canar. 3(2): 129 (1840).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.),
Lampang (Doi Khun Tan N.P.), Nakhon Nayok (Khao Yai N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary), Yala (Banglang

Reservoir). Boonpragob et al. (1998: 217).
SPECIMENS. BM, CMU.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea, Sri
Lanka, Vietnam.

32
Leptogium burgessii (L.) Mont., in Webb & Berthelot,
Hist. Nat. Iles Canar. 3(2): 129 (1840).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).

Mongkonsuk et al. (1996: 17).
SPECIMEN. Queen Sirikit Botanical Garden Herbarium.

ASIA. India, Indonesia.

Leptogium caesium (Ach.) Vain., Acta Soc. Fauna Fl.
Fenn. 7: 225 (1890).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 116).
SPECIMEN. TUR-V.

ASIA. India, Indonesia, Papua New Guinea.

Leptogium chloromelum (Sw.) Nyl., Syn. Meth. Lich. 1:
128 (1858).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).

Mongkonsuk et al. (1996: 17).
SPECIMEN. Queen Sirikit Botanical Garden Herbarium.

ASIA. India, Malaysia, Sri Lanka, Taiwan.

Leptogium cochleatum (Dicks.) P.M. Jorg. & P. James,
Lichenologist 15: 113 (1983).

Leptogium tremelloides auct.

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99, as
Leptogium tremelloides).

SPECIMEN. BM.

ASIA. — India.

Leptogium coralloideum (Meyen & Flot.) Vain., Ann.
Acad. Sci. Fenn., ser. A, 6(7): 110 (1915).

Leptogium chloromelum auct. non. (Sw.) Nyl.

THAILAND. Chiang Mai (Doi Inthanon N.P.), Chumphon (Koh
Tao I.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Paulson
(1930: 99, as Leptogium chloromelum auct. non. (Sw.) Nyl.).

BKF, BM, CMU.

Indonesia, Papua New Guinea, Philippines.

SPECIMENS.
ASIA.

Leptogium cyanescens (Rabenh.) K6rb., Syst. Lich. Germ.:
420 (1855).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Lampang (Doi Khun Tan N.P.), Nakhon Nayok
(Khao Yai N.P.), Tak (Ton Krabakyai N.P.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Wolseley (1991: 236), Wolseley &
Aguirre-Hudson (1997a: 333, 1997b: 355), Boonpragob et al. (1998:
par

SPECIMENS. BKF, BM, CMU.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Sri Lanka, Taiwan.

Leptogium denticulatum Nyl., Ann. Sci. Nat. Bot., sér. 5, 7:
302 (1867).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Nakhon Nayok (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Hudson (1997a: 333; 1997b: 355), Boonpragob et al. (1998: 217).
BKF, BM, CMU.

ASIA. China (incl. Hong Kong), India, Malaysia.

SPECIMENS.

Leptogium javanicum Mont., Syll. Gen. Sp. Crypt.: 379
(1856).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &
Aguirre-Hudson (1997a: 333).

SPECIMENS. BKF, BM, CMU.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philippines.
Leptogium marginellum (Sw.) S.F. Gray, Nat. Arr. Brit. Pl.
1: 401 (1821).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Queen Sirikit Botanical Garden), Tak (Ton Krabakyai N.P.), Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley (1991:
236), Mongkonsuk et al. (1996: 17).

SPECIMENS. BKF, BM, CMU, Queen Sirikit Botanical Garden
Herbarium.
ASIA. India, Indonesia, Papua New Guinea, Philippines.

Leptogium moluccanum (Pers.) Vain., Etude Class. Lich.
Brésil 1: 223 (1890).

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 46), Sato
(1962: 2).
SPECIMENS. TUR-V, TNS.

ASIA. China, India, Indonesia, Papua New Guinea, Taiwan.
Leptogium resupinans Nyl., Ann. Sci. Nat. Bot., sér. 4, 15:
368 (1861).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Jorgensen (1997:
128).

SPECIMENS. BM.

ASIA. India.

Leptogium trichophorum Miill. Arg., Flora 72: 505
(1889).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.).
Vainio (1921: 46).

SPECIMENS. BM, CMU, TUR-V.

ASIA. China, India, Indonesia, Philippines, Taiwan.

Letrouitia Hafellner & Bellem.; Letrouitiaceae

Letrouitia domingensis (Pers.) Hafellner & Bellem., Nova
Hedwigia 35: 281 (1982).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Lampang (Doi Khun Tan N.P.), Nakhon
Ratchasima (Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary), Yala (Banglang N.P.). Wolseley et al. (1994:
122), Boonpragob et al. (1998: 217).

SPECIMENS. BM, CMU, RAMK.

CATALOGUE OF THE LICHENS OF THAILAND

ASIA. India, Papua New Guinea, Philippines, Sri Lanka, Taiwan,
Vietnam.

Letrouitia flavocrocea (Ny1.) Hafellner & Bellem., Nova
Hedwigia 35: 281 (1982).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary),
Yala (La Or Rung). Wolseley et al. (1994: 122).

SPECIMENS. BM, CMU.

ASIA. India, Philippines.
Letrouitia leprolyta (Ny1.) Hafellner, Nova Hedwigia 35:
693 (1981).

THAILAND. Chumphon (N. of Pranburi), Lampang (Doi Luang),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Hafellner
(1981: 694), Wolseley et al. (1994: 122), Wolseley & Aguirre-
Hudson (1997a: 333, 339; 1997b; 355).

SPECIMENS. BM.

ASIA. India, Indonesia, Papua New Guinea, Sri Lanka.

Letrouitia subvulpina (Ny1.) Hafellner, Nova Hedwigia 35:
705 (1982).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Wolseley et al. (1994:
122), Wolseley & Aguirre-Hudson (1997a: 333; 1997b: 355),

Boonpragob et al. (1998; 217).
SPECIMENS. BM, RAMK.

ASIA. India, Indonesia, Papua New Guinea, Sri Lanka.

Letrouitia transgressa (Malme) Hafellner & Bellem., Nova
Hedwigia 35: 710 (1982).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Lampang (Doi
Luang), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).

Wolseley et al. (1994: 122).
SPECIMENS. BM, CMU.

ASIA. India, Indonesia, Taiwan.

Letrouitia vulpina (Tuck.) Hafellner & Bellem., Nova
Hedwigia 35: 281 (1982).

THAILAND. Satun(Thaleban N.P.), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Wolseley et al. (1994: 122), Wolseley & Aguirre-
Hudson (1997a: 333, 339; 1997b: 355).

SPECIMENS. BM.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka.

Lobaria (Schreb.) Hoffm.; Lobariaceae
Lobaria discolor (Bory) Hue, Nouv. Arch. Mus. Paris, sér.
4, 3: 23 (1901).

THAILAND. Chiang Mai (Chieng Dao N.P., Doi Inthanon N.P., Doi
Suthep N.P.). Yoshimura (1978: 39), Wolseley (1991: 234).

BM, C.

ASIA. China, India, Indonesia, Philippines, Sri Lanka, Taiwan.

SPECIMENS.

a3

Lobaria isidiosa (Miill. Arg.) Vain., Philipp. J. Sci., sect. C,
8: 129 (1913).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Sato (1962: 3), Yoshimura (1971: 295).

SPECIMENS. TNS.

ASIA. China, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka, Taiwan, Vietnam.

Lobaria pseudopulmonaria Gyeln., Acta Fauna Fl. Univ.,
ser. 2, 1(5/6): 6 (1933).

THAILAND. Chiang Mai (Chieng Dao N.P., Doi Suthep N.P.), Loei
(Phu Luang N.P). Yoshimura (1971: 291; 1978: 39).

SPECIMENS. C, KYO, TNS.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea,
Philippines, Taiwan.

Lobaria retigera (Bory) Trevis. Schedae Lich. Veneta No.
75 (1869).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Wolseley & Aguirre-Hudson (1997a: 334).

SPECIMENS. BM.
ASIA. China, India, Indonesia, Philippines, Sri Lanka, Taiwan,
Vietnam.

Maronea A. Massal; Fuscideaceae

Maronea constans (Ny1.) Hepp, Flecht. Eur.: 771 (1860).

THAILAND. Chiang Mai (Doi Suthep N.P.). Wolseley & Aguirre-
Hudson (1997a: 334).

SPECIMENS. BM.

ASIA. India, Philippines.

Mazosia A. Massal.; Opegraphaceae
Mazosia melanophthalma (Mill. Arg.) R. Sant., Symb.
Bot. Upsal. 12(1): 117 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. RAMK.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Philippines, Sri Lanka, Vietnam.

Megalospora Meyen; Megalosporaceae

Megalospora bicolorata (Vain.) Zahlbr., Cat. Lich. Univ. 4:
87 (1927).

THAILAND. Chiang Mai (Doi Sutep N.P.). Vainio (1921: 48).
SPECIMEN. 'TUR-V (not located).

ASIA. —

Megalospora sulphurata Meyen, in Meyen & Flotow, Nov.

Actorum Acad. Caes. Leop.-Carol. Nat. Cur. 19, suppl.:
228 (1843).

34

THAILAND. Chiang Mai (Doi Suthep N.P.). Wolseley & Aguirre-
Hudson (1997a: 339).

SPECIMENS. BM.

ASIA. India, Indonesia, Papua New Guinea.

Megalospora tuberculosa (Fée) Sipman, Biblioth.
Lichenol. 18: 156 (1983).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Sipman (1983: 238), Wolseley & Aguirre-Hudson

(1997a: 339), Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK, US.

ASIA. China, India, Taiwan.

Menegazzia A. Massal.; Hypogymniaceae

Menegazzia terebrata (Hoffm.) A. Massal., Neag. Lich.: 3
(1854).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1995: 323; 1997a: 339).

SPECIMEN. BM.

ASIA. China, India, Malaysia, Taiwan.

Microtheliopsis Miill. Arg.; Microtheliopsidaceae

Microtheliopsis uleana Miill. Arg., Flora 73: 195 (1890).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB.

ASIA.
Lanka.

Indonesia, Malaysia, Papua New Guinea, Philippines, Sri

Myelochroa (Asahina) Elix & Hale; Parmeliaceae
Myelochroa aurulenta (Tuck.) Elix & Hale, Mycotaxon
29: 240 (1987).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Phitsanulok (Phu

Hin Rongkla N.P.). Moon et al. (2000a: 102).
SPECIMENS. TNS.

ASIA. —

Myelochroa entotheiochroa (Hue) Elix & Hale,
Mycotaxon 29: 240 (1987).

THAILAND. Chiang Mai (Maetang). Moon et al. (2000a: 102).
SPECIMENS. TNS.

ASIA. —

Myelochroa irrugens (Hue) Elix & Hale, Mycotaxon 29:
241(1987).

THAILAND. Chiang Mai (Maetang). Moon et al. (2000a: 102).
SPECIMENS. TNS.

ASIA. —

Myelochroa perisidians (Ny1.) Elix & Hale, Mycotaxon 29:
241 (1974).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Parmelina perisidians (Ny]1.) Hale.

THAILAND. Loei (Ban Sui). Hale (1976c: 39, as Parmelina
perisidians), Moon et al. (2000a: 102).

TNS, US.
ASIA. Philippines, Sri Lanka.

SPECIMENS.

Myelochroa siamea Kurok., J. Jap. Bot. 73: 13 (1998).

THAILAND. Chiang Mai (Maetang District). Kurokawa (1998:

13), Moon et al. (2000a: 102).
SPECIMENS. BM, TNS.

ASIA. —
Myelochroa xantholepis (Mont. & Bosch) Elix & Hale,
Mycotaxon 29: 241 (1987).

Parmelina xantholepis Mont. & Bosch.

THAILAND. Chiang Mai (Maetang, Doi Suthep N.P., Chiang Dao
N.P.), Phitsanulok (Phu Hin Rongkla, N.P.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Hale (1976c: 53, as Parmelina

xantholepis), Moon et al. (2000a: 102).
SPECIMENS. BM, CMU, TNS, US.

ASIA. India, Indonesia, Philippines.

Myeloconis P.M. McCarthy & Elix; Myeloconaceae
Myeloconis erumpens P.M. McCarthy & Elix,
Lichenologist 28: 406 (1996).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB, RAMK.

ASIA. Papua New Guinea.

Myriotrema Fée; Thelotremataceae
Myriotrema albocinctum Hale, Bull. Brit. Mus. (Nat.
Hist.), Bot. 8(3): 273(1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. Malaysia, Sri Lanka.
Myriotrema bahianum (Ach.) Hale, Mycotaxon 11: 132
(1980).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).

Homchantara (1999: 151-153).
SPECIMENS. BM, RAMK.

ASIA. Malaysia, Papua New Guinea, Philippines.

Myriotrema clandestinum (Fée) Hale, Mycotaxon 11: 133
(1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.
ASIA. — Sri Lanka.

CATALOGUE OF THE LICHENS OF THAILAND

Myriotrema compunctum (Ach.) Hale, Mycotaxon 11: 133
(1980).

THAILAND. Lampang (Teak Improvement Centre), Rayong

(Klaeng District). Homchantara (1999: 153).
SPECIMENS. BM, RAMK.

ASIA. Indonesia, Philippines, Sri Lanka, Vietnam.

Myriotrema costaricense (Miill. Arg.) Hale, Mycotaxon 11:
133 (1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Boonpragob et al. (1998:

217), Homchantara (1999: 156-158).
SPECIMENS. BM, RAMK.

ASIA. India, Malaysia.

Myriotrema decorticum Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 277 (1981).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Homchantara (1999: 159-161).

SPECIMENS. BM.

ASIA. Sri Lanka.

Myriotrema desquamans (Miill. Arg.) Hale, Mycotaxon
11: 133 (1980).

THAILAND. Chantaburi (Khitchakut N.P., Khao Soi Dao Wildlife
Sanctuary), Nakhon Sri Thammerat (Khao Luang N.P.), Sukhothai
(Ramkhamhaeng N.P.), Tak (Ton Krabak Yai N.P.). Homchanatara

(1999: 161-163).
SPECIMENS. BM, RAMK.

ASIA. India, Malaysia, Papua New Guinea, Philippines, Sri Lanka.

Myriotrema fissurinum Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 279 (1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Phetchabun (Phu
Hin Rongkla N.P.). Homchantara (1999: 159).

SPECIMENS. RAMK.

ASIA. India, Sri Lanka.

Myriotrema glaucescens (Ny1.) Hale, Mycotaxon 11: 133
(1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Homchantara (1999: 168).

SPECIMENS. BM, RAMK.

ASIA. India, Sri Lanka.

Myriotrema grandisporum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara

(1999: 224-225, as Myriotrema ‘unidentified 10’).
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Be

Myriotrema khaoyaianum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara
(1999: 209-211, as Myriotrema ‘unidentified 3’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Myriotrema khuntanense Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Lampang (Doi Khuntan N.P.). Homchantara (1999:
226-227, as Myriotrema ‘unidentified 11’).

SPECIMENS. BM (holotype), RAMK (isotype).

ASIA. —

Myriotrema masonhalei (Patw. & C.R. Kulk.) Hale,
Mycotaxon 11: 134 (1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. RAMK.

ASIA. India.

Myriotrema microporum (Mont.) Hale, Mycotaxon 11: 134
(1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Phetchabun (Phu Hin Rongkla N.P.). Boonpragob

et al. (1998: 217), Homchantara (1999: 175).
SPECIMENS. BM, RAMK.

ASIA. India, Indonesia, Malaysia, Philippines, Sri Lanka, Taiwan.

Myriotrema microstomum (Mill. Arg.) Hale, Mycotaxon
11: 134 (1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Phetchabun (Phu Hin
RongklaN.P.), Sukhotai (Ramkhamhaeng N.P.). Homchantara (1999:

Wi ap
SPECIMENS. BM, RAMK.

ASIA. Sri Lanka, Taiwan.

Myriotrema minutulum (Hale) Hale, Mycotaxon 11: 134
(1980).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Homchantara (1999: 180).

SPECIMENS. BM.

ASIA. Sri Lanka.

Myriotrema minutum (Hale) Hale, Mycotaxon 11: 134
(1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara
(1999: 182).

SPECIMEN. RAMK.
ASIA. Malaysia, Sri Lanka.

Myriotrema olivaceum Fée, Essai Crypt. Ecorc. Offic.: 103
(1824).

36

THAILAND. Loei (Na Haeo N.P., Phuteen Suansai forest, Ban
Bomoeng Noi). Homchantara (1999: 185).

SPECIMENS. RAMK.
ASIA. Sri Lanka.

Myriotrema phaeosporum (Ny1.) Hale, Mycotaxon 11: 134
(1980).

Leptotrema phaeosporum (Nyl.) Miill. Arg.

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99, as
Leptotrema phaeosporum).

SPECIMEN. BM (not located).

ASIA. —

Myriotrema polytretum Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 291 (1981).

THAILAND. Chiang Mai (Doi Suthep N.P.), Lampang (Doi Pi
Luang Ditrict), Loei (Phu Ruea N.P.), Uthai Thani (Huay Kha
Khaeng N.P.). Homchantara (1999: 187).

SPECIMENS. BM, RAMK.

ASIA. Malaysia, Sri Lanka.

Myriotrema rugiferum (Harm.) Hale, Mycotaxon 11: 135
(1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Phetchabun (Phu Hin
Rongkla N.P.). Homchantara (1999: 189).

SPECIMENS. BM, RAMK.

ASIA. India, Papua New Guinea, Philippines, Sri Lanka.

Myriotrema subconforme (Ny1.) Hale, Mycotaxon 11: 135
(1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. RAMK.

ASIA. India.

Myriotrema subgranulosum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Loei (Na Po District).

Homchantara (1999: 205-207, as Myriotrema ‘unidentified 1”).
SPECIMENS. BM (holotype), RAMK (isotype).

ASIA. —

Myriotrema subminutum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yao N.P.), Phetchabun
(Phu Hin Rongkla N.P.), Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Homchantara (1999: 215-217, as Myriotrema ‘uniden-
tified 6’).
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Myriotrema terebratulum (Ny1.) Hale, Mycotaxon 11: 135
(1980).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Loei (Na Po District),

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et al. (1998:

217), Homchantara (1999: 196).
SPECIMENS. BM, RAMK.

ASIA. — India.

Myriotrema viride Nagarkar & Hale, Mycotaxon 35: 440
(1989).

THAILAND. Sukhothai (Ramkhamhaeng N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Boonpragob et al. (1998: 217),

Homchantara (1999: 199).
SPECIMENS. BM, RAMK.

ASIA. Malaysia.

Myriotrema wightii (Taylor) Hale, Mycotaxon 11: 135
(1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Kanchanaburi (Erawan
N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).

Homchantara (1999: 201).
SPECIMENS. BM, RAMK.

ASIA. India, Malaysia, Philippines, Sri Lanka.

Nephroma Ach.; Nephromataceae

Nephroma helveticum Ach., Lichenogr. Universalis: 523
(1810).

Nephroma tropicum (Mill. Arg.) Zahlbr.

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Wolseley & Aguirre-Hudson (1995: 324; 1997a: 334, 339, as Neph-

roma tropicum).
SPECIMENS. BM, CMU.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea.
Nephromopsis Miill. Arg.; Parmeliaceae
Nephromopsis stracheyi (Church. Bab.) Miill. Arg., Flora
74: 374 (1891).

Nephromopsis ornata auct. non (Mill. Arg.) Hue

THAILAND. Chiang Mai (Doi Suthep N.P), Loei (Phu Rua N.P.).
Wolseley & Aguirre-Hudson (1995: 324, as Nephromopsis ornata

auct. non (Miill. Arg.) Hue).
SPECIMENS. BKF, BM, CMU.

ASIA. China, India, Malaysia, Taiwan.

Normandina (Nyl.) Vain.; incert. sed.

Normandina pulchella (Borrer) Nyl., Ann. Sci. Nat., Bot.,
sér. 4, 15: 382 (1861).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Wolseley & Aguirre-Hudson (1997a: 334),

Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. China, Malaysia, Papua New Guinea.

CATALOGUE OF THE LICHENS OF THAILAND

Ocellularia Meyen; Thelotremataceae

Ocellularia albomaculata Hale, Bull. Brit. Mus. (Nat.
Hist.) Bot. 8(3): 300 (1981).

THAILAND. Satun (Thalae Ban N.P.). Homchantara (1999: 237).

SPECIMENS. BM.

ASIA. Sri Lanka.

Ocellularia allosporoides (Ny\.) Patw. & C.R. Kulk.,
Kavaka 5: 5 (1977).

THAILAND. Lampang (Wiang Kosai N.P.), Chantaburi (Khitchakut
N.P.), Kanchanaburi (Khao Lem N.P.), Nakhon Ratchasima (Khao
Yai N.P.). Boonpragob et al. (1998: 217), Homchantara (1999: 239).

SPECIMENS. BM, RAMK.

ASIA. _ India.
Ocellularia arecae (Vain.) Hale, Mycotaxon 11: 136
(1980).

Thelotrema arecae Vain.
Thelotrema interpositum Nyl.

THAILAND. Chantaburi (Khitchakut N.P.), Loei (Na Haeo N.P.),
Nakhon Ratchasima (Khao Yai N.P.), Phetchabun (Phu Hin Rongkla
N.P.), Trat (Koh Chang I.). Vainio (1909: 119, as Thelotrema
arecae), Salisbury (1971: 277, as T. interpositum), Hale (1981: 305),
Boonpragob et al. (1998: 217), Homchantara (1999: 242).

SPECIMENS. RAMK, TUR-V (holotype).

ASIA. India, Malaysia, Philippines.

Ocellularia ascidioidea Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 300 (1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Phetchabun (Phu
Hin Rongkla N.P.), Sukhotai (Ramkhamhaeng N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Boonpragob et al. (1998:
217), Homchantara (1999: 245).

SPECIMENS. BM, RAMK.

ASIA. Malaysia, Sri Lanka.

Ocellularia asiatica (Vain.) Hale, Hedwigia 46: 175
(1907).

Thelotrema asiaticum Vain.

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Trat (Koh Chang
I.). Vainio (1909: 120, as Thelotrema asiaticum), Patwardan &
Kulkarni (1977a: 10, as T. asiaticum), Boonpragob et al. (1998:
pany

SPECIMENS.
India.

RAMK, TUR-V (holotype).
ASIA.
Ocellularia chonestoma (Leight.) Zahlbr., Cat. Lich. Univ.
2: 586 (1923).

THAILAND. Kanchanaburi (Khao Lem N.P.). Homchantara (1999:
248).

SPECIMENS. RAMK.

37

ASIA. Malaysia, Philippines, Sri Lanka.

Ocellularia concolor Meyen & Flot., Nov. Actorum Acad.
Caes. Leop.-Carol. Nat. Cur. 19, suppl.: 230 (1843).

THAILAND. Chantaburi (Khitchakut N.P.), Chiang Mai (Doi Suthep
N.P.), Nakhon Ratchasima (Khao Yai N.P.), Phetchabun (Phu Hin
Rongkla N.P.), Sukhothai ( Ramkhamhaeng N.P.). Boonpragob et

al. (1998: 217), Homchantara (1999: 250).
SPECIMENS. BM, RAMK.

ASIA. India, Malaysia, Philippines.

Ocellularia conformis (Fée) Hale, Mycotaxon 11: 136
(1980).

THAILAND.
(1999: 250).

SPECIMENS.

India.

Phetchabun (Phu Hin Rongkla N.P), Homchantara

RAMK.

ASIA.

Ocellularia crassa (Miill. Arg.) Hale, Mycotaxon 11: 136
(1980).

THAILAND. Phetchabun (Phu Hin Rongkla N.P). Homchantara
(1999: 250).

SPECIMENS.
India.

RAMK.

ASIA.

Ocellularia diacida Hale, Mycotaxon 7: 378 (1978).

THAILAND. Chantaburi (Kitchakut N.P., Namtok Phliu N.P., Khao
Soi Dao Wildlife Sanctuary), Chiang Mai (Doi Inthanon N.P., Doi
Suthep N.P.), Lampang (Doi Pae Luang), Loei (Na Haeo N.P., Phu
Riea N.P.), Nakhon Ratchasima (Khae Yai N.P.), Phetchabun (Phu
Hin Rongkla N.P), Sakon Nakhon (Phu Phan N.P.), Sukhothai
(Ramkhamhaeng N.P.), Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Boonpragob et al. (1998: 217), Homchantara (1999:
250).

SPECIMENS. BM, RAMK.

ASIA. India, Sri Lanka.

Ocellularia diospyrosis Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Tak (Ton Krabak Yai N.P.). Homchantara (1999: 314—

315, as Ocellularia ‘unidentified 1’).
SPECIMENS. BM (holotype).

ASIA. —

Ocellularia dolichotata (Ny1.) Zahlbr., Cat. Lich. Univ. 2:
589 (1923).

Thelotrema siamense Vain.

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Trat (Koh Chang
I.). Vainio (1907: 174; 1909: 120, as Thelotrema siamense), Hale
(1981: 303), Sethy et al. (1987: 193), Boonpragob et al. (1998: 217),

Homchantara (1999: 264).
SPECIMENS. BM, C (isolectotype), RAMK, TUR-V (lectotype).

ASIA. India, Malaysia, Papua New Guinea, Philippines, Singapore.

38

Ocellularia emersa (Kremp.) Miill. Arg., Flora 69: 310
(1886).

THAILAND. Chanthaburi (Khitchakut N.P.), Loei (Phu Ruea N.P.),
Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et al. (1998:
217), Homchantara (1999: 267).

SPECIMENS. RAMK.

ASIA. Sri Lanka.

Ocellularia epitrypa (Ny1.) Hale, Mycotaxon 11: 136
(1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Boonpragob et al. (1998:

217), Homchantara (1999: 269).
SPECIMENS. BM, RAMK.

ASIA. India, Philippines, Sri Lanka.

Ocellularia eumorpha (Stirt.) Hale, Mycotaxon 11: 136
(1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Na Haeo N.P.).
Homchantara (1999: 271).

SPECIMENS. BM, RAMK.

ASIA. India, Sri Lanka.

Ocellularia fissa (Ny1.) Hale, Mycotaxon 11: 136 (1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. RAMK.

ASIA. — India.

Ocellularia flavescens Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Homchantara (1999:

339-341, as Ocellularia ‘unidentified 13’).
SPECIMENS. BM (holotype), RAMK (isotype).

ASIA. —

Ocellularia groenhartii Hale, Mycotaxon 3: 173 (1975).

THAILAND. Chantaburi (Khao Soi Dao N.P., Kitchakut N.P.,
Namtok Phliu N.P.), Kanchanaburi (Khao Laem N.P.), Lampang
(Wiang Kosai N.P.), Nakhon Ratchasima (Khao Yai N.P.).

Boonpragob et al. (1998: 217), Homchantara (1999: 272).
SPECIMENS. BM, RAMK.

ASIA. India, Indonesia.

Ocellularia guptei (Nagarkar, Sethy & Patw.) D.D.
Awasthi, Mycotaxon 23: 79 (1986).

THAILAND. Chantaburi (Khitchakut N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Boonpragob et al. (1998: 217), Homchantara
(1999: 276).

SPECIMENS.
India.

RAMK.
ASIA.

Ocellularia inthanonensis Homchantara & Coppins,
Lichenologist 34: (in press).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

THAILAND. Chiang Mai (Doi Inthanon N.P.). Homchantara (1999:
333-334, as Ocellularia ‘unidentified 10’).

SPECIMENS. BM (holotype), RAMK (isotype).

ASIA. —

Ocellularia kansriae Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Pechabun (Phu
Hin Rongkla N.P.), Sukhothai (Ramkhamhaeng N.P.). Homchantara
(1999: 335-337, as Ocellularia ‘unidentified 11’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Ocellularia kratingensis Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chantaburi (Khitchakut N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Homchantara (1999: 316-318, as Ocellularia
‘unidentified 2’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. Malaysia, Philippines, Sri Lanka.

Ocellularia leucina (Miill. Arg.) Hale, Mycotaxon 11: 137
(1980).

THAILAND. Chantaburi (Namtok Phliu N.P.), Kanchanaburi
(Kroengkawia), Nakhon Ratchasima (Khao Yai N.P.). Boonpragob
et al. (1998: 217), Homchantara (1999: 278).

SPECIMENS. RAMK.

ASIA. India.
Ocellularia marivelensis (Vain.) Hale, Mycotaxon 11: 137
(1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217), Homchantara (1999: 281).

SPECIMENS. RAMK.

ASIA. India, Malaysia, Philippines.
Ocellularia massalongoi (Mont.) Hale, Mycotaxon 11: 137
(1980).

THAILAND. Chantaburi (Namtok Phliu N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary), Yala (Banlang N.P.). Boonpragob et al. (1998: 21h);
Homchantara (1999: 283).

SPECIMENS. BM, RAMK.

AsIA. India, Malaysia, Papua New Guinea, Philippines.
Ocellularia meiosperma (Ny1.) Hale, Mycotaxon 11: 137
(1980).

THAILAND. Chantaburi (Namtok Phliu N.P.), Chiang Mai (Doi
Suthep N.P.), Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217), Homchantara (1999: 281).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Malaysia, Philippines, Sri Lanka.

CATALOGUE OF THE LICHENS OF THAILAND

Ocellularia melanopthalma Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Phetchabun (Phu Hin Rongkla N.P.). Homchantara
(1999: 324-326, as Ocellularia ‘unidentified 6’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Ocellularia monosporoides (Ny1.) Hale, Mycotaxon 11:
137 (1980).

THAILAND. Chantaburi (Khitchakut N.P.), Phetchabun (Phu Hin
Rongkla N.P.). Homchantara (1999: 288-290).

SPECIMENS. RAMK.

ASIA. Sri Lanka.

Ocellularia microascidium (Vain.) Zahlbr., Cat. Lich.
Univ. 2: 595 (1923).

Thelotrema microascidium Vain.

THAILAND. Trat (Koh Kong I.). Vainio (1907: 176, as Thelotrema
microascidium).

SPECIMEN. ‘TUR-V (holotype).

ASIA. —

Ocellularia neoleucina Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chantaburi (Namtok Phliu N.P.). Homchantara (1999:
333-334, as Ocellularia ‘unidentified 15’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Ocellularia neopertusariiformis Hale, Bull. Brit. Mus.
(Nat. Hist.), Bot. 8(3): 315 (1981).

THAILAND. Loei (Phu Ruea N.P.), Nakhon Ratchasima (Khao Yai
N.P.), Phetchabun (Phu Hin Rongkla N.P.). Homchantara (1999:

290-293).
SPECIMENS. RAMK.

ASIA. Malaysia, Sri Lanka.

Ocellularia nylanderiana Hale, Phytologia 26: 414 (1973).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. RAMK.

ASIA. — India.

Ocellularia paleoamplior Aptroot & Sipman, J. Hattori
Bot. Lab. 91: 333 (2001).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara
(1999: 326-328, as Ocellularia ‘unidentified 7’).

SPECIMENS. RAMK (holotype).
ASIA. —

Ocellularia papillata (Leight.) Zahlbr., Cat. Lich. Univ. 2:
597 (1923).

39

THAILAND. Chantaburi (Namtok Phliu N.P., Khitchakut N.P.),
Lampang (Doi Mae Luang), Nakhon Ratchasima (Khao Yai N.P.),
Sakhon Nakhon (Phu Phan N.P.), Sukhothai (Ramkhamhaeng N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Boonpragob et
al. (1998: 217), Homchantara (1999: 295).

SPECIMENS. BM, RAMK.

ASIA. India, Philippines, Sri Lanka.

Ocellularia peremergens Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Loei (Na Haeo N.P.), Nakhon Ratchasima (Khao Yai N.P.).
Homchantara (1999: 331-332, 337, 349-350, as Ocellularia ‘uni-
dentified 9’, ‘12’, 18’).

SPECIMENS. BM (holotype), RAMK (isotype).

ASIA. —

Ocellularia perforata (Leight.) Miill. Arg., Hedwigia 31:
284 (1892).

THAILAND. Chantaburi (Khao Soi Dao Wildlife Sanctuary, Namtok
Phliu N.P., Kitchakut N.P.), Nakhon Ratchasima (Khao Yai N.P.),
Nakhon Sri Thammerat (Khao Luang N.P.), Petchabun (Phu Hin
Rongkla N.P.), Sukhothai (Ramkhamhaeng N.P.). Boonpragob et al.
(1998: 217), Homchantara (1999: 297).

SPECIMENS. RAMK.

ASIA. China (Hong Kong), India, Malaysia, Papua New Guinea.
Ocellularia pertusariformis (Leight.) Zahlbr., Cat. Lich.
Univ. 2: 598 (1923).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.

ASIA. India.

Ocellularia pluripora Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 319 (1981).

THAILAND. Chanthburi (Khitchakut N.P., Namtok Phliu N.P.),
Loei (Phu Ruea N.P., Na Haeo N.P.), Nakhon Ratchasima (Khao Yai

N.P.). Boonpragob et al. (1998: 217), Homchantara (1999: 304).
SPECIMENS. RAMK.

ASIA. India, Malaysia.

Ocellularia pluripoides Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chiang Mai (Doi Suthep N.P.). Homchantara (1999:

318-319, as Ocellularia ‘unidentified 3’).
SPECIMENS. BM (holotype).

ASIA. —
Ocellularia punctulata (Leight.) Zahlbr., Cat. Lich. Univ.
2°-599 (1923).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.

ASIA. India, Malaysia.

40

Ocellularia rhicnoporoides Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chantaburi (Namtok Phliu N.P.). Homchantara (1999:
329-330, as Ocellularia ‘unidentified 8’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Ocellularia sticticans Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 323 (1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. RAMK.

ASIA. — Sri Lanka.

Ocellularia subleucina Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chantaburi (Khitchakut N.P.). Homchantara (1999:

341-343, as Ocellularia ‘unidentified 14’).
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Ocellularia wolsleyanum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara

(1999: 347-349, as Ocellularia ‘unidentified 17’).
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Opegrapha Ach.; Opegraphaceae

Opegrapha aff. lambinonii Sérus., Lejeunia 90: 2 (1978).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Sérusiaux (1985:

23):
SPECIMEN. LG.

ASIA. India, Papua New Guinea, Philippines, Sri Lanka.

Opegrapha prosodea Ach., Methodus: 22 (1803).
THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 99).
SPECIMEN. BM.

ASIA. China, India, Malaysia, Papua New Guinea, Taiwan.

Opegrapha robusta Vain., Bot. Tidsskr. 29: 137 (1909).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 137).
SPECIMENS. TUR-V (syntypes).

ASIA. Indonesia, Papua New Guinea.

Opegrapha subvulgata Ny1., Flora 52: 71 (1869).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 136).
SPECIMEN. TUR-V.

ASIA. — India.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Pannaria Delise ex Bory; Pannariaceae
Pannaria dispartita (Ny1) Vain., Suomal. Elain-ja Kasvit.
Seuran Vanamon Julk. 1(3): 45 (1921).
THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 45).
SPECIMEN. TUR-V.

ASIA. —

Paraparmelia Elix & J. Johnst.; Parmeliaceae

Paraparmelia rodriguesiana (Hue) Elix & J. Johnst.,
Mycotaxon 27: 280 (1986).

THAILAND. Phitsanulok (Phu Hin Rongkla, N.P.). Moon et al.
(2000a: 102).

SPECIMEN. TNS.

ASIA. —

Parmelaria D.D. Awasthi; Parmeliaceae

Parmelaria thomsonii (Stirt.) D.D. Awasthi, J. Hattori Bot.
Lab. 63: 368 (1987).

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Phu Rua N.P.).

Wolseley & Aguirre-Hudson (1995: 325).
SPECIMENS. BM, DUKE, TNS.

ASIA. India, Nepal.

Parmelia Ach.; Parmeliaceae

Parmelia saxatilis (L.) Ach., Methodus: 204 (1803).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden),

Mongkonsuk et al. (1996: 17).
SPECIMEN. Queen Sirikit Botanical Garden Herbarium.

ASIA. China, India.

Parmeliella Miill. Arg.; Pannariaceae

Parmeliella brisbanensis (C. Knight) P.M. Jgrg. & D.J.
Galloway, Fl. Australia 54: 316 (1992).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary). Wolseley & Aguirre-Hudson (1995: 325; 1997a: 334, 1997b:

356), Boonpragob et al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. —

Parmeliella papillata P.M. Jorg., Biblioth. Lichenol. 78:
133 (2001).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Jgrgensen (2001: 134).

SPECIMENS. BM.

ASIA. Papua New Guinea.

CATALOGUE OF THE LICHENS OF THAILAND

Parmelina Elix; Parmeliaceae

Parmelina quercina (Willd.) Hale, Phytologia 28: 483
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Mongkonsuk et al. (1996: 17).

SPECIMEN.
ASIA. China, India.

Queen Sirikit Botanical Garden Herbarium.

Parmelina usambarensis (J. Steiner & Zahlbr.) Hale,
Phytologia 28: 483 (1974).

Parmelia usambarensis J. Steiner & Zahlbr.

THAILAND. Loei (near Phu Kradung). Hale (1976c: 51, as Parmelia
usambarensis), Singh (1979: 222, as P. usambarensis), Kurokawa &
Kashiwadani (1988: 7, as P. usambarensis), Moon et al. (2000a:
102).

SPECIMENS.

India.

TNS.
ASIA.
Parmelinella Elix & Hale; Parmeliaceae
Parmelinella chozoubae (Singh & Sinha) Elix &
Pooprang, in Pooprang et al., Mycotaxon 71: 121 (1999).

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Nahaew N.P.),
Nakhon Ratchasima (Khao Yai N.P.), Phitsanulok (Phu Hin Rongkla

N.P.). Pooprang et al. (1999: 121).
SPECIMENS. RAMK.

ASIA. _ India.

Parmelinella simplicior (Hale) Elix & Hale, Mycotaxon
29: 242 (1987).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.). Moon et al.
(2000a: 102).

SPECIMENS.

India.

TNS.

ASIA.

Parmelinella wallichiana (Taylor) Elix & Hale, Mycotaxon
29: 242 (1987).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P., Doi Pacho, Maetang), Loei (Ban Sui, Phu
Kradung), Phitsanulok (Phu Hin Rongkla, N.P.), Tak (Ton Krabakyai
N.P.), Uthai Thani ( Huay Kha Khaeng Wildlife Sanctuary). Wolseley
& Aguirre-Hudson (1995: 326; 1997a: 334; 1997b: 356), Moon et
al. (2000a: 103).

SPECIMENS. BKF, BM, CMU, TNS.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Phil-
ippines, Sri Lanka, Taiwan.

Parmelinopsis Elix & Hale; Parmeliaceae
Parmelinopsis expallida (Kurok.) Elix & Hale, Mycotaxon
29: 242 (1987).

Parmelina expallida (Kurok.) Elix & Hale.

THAILAND. Chiang Mai (Maetang District). Hale (1976c: 30 as
Parmelina expallida), Patwardhan & Prabhu (1977: 111, as

4]

P. expallida), Wolseley & Aguirre-Hudson (1995: 325), Moon et al.

(2000a: 103).
SPECIMENS. TNS, US.

ASIA. India, Taiwan.

Parmelinopsis horrescens (Taylor) Elix & Hale,
Mycotaxon 29: 242 (1987).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.). Pooprang et al.
(1999; 121).

SPECIMENS. RAMK.

ASIA. India, Sri Lanka.

Parmelinopsis microlobulata (D.D. Awasthi) Elix & Hale,
Mycotaxon 29: 242 (1987).

THAILAND. Nakhon Sri Thammarat (Khao Luang N.P.),
Phitsanulok (Phu Hin Rongkla N.P.). Pooprang et al. (1999: 121).
SPECIMENS. RAMK.

ASIA. India.

Parmelinopsis minarum (Vain.) Elix & Hale, Mycotaxon
29: 243 (1987).

THAILAND. Chiang Mai (Doi Suthep N.P.). Pooprang et al. (1999:

£22);
SPECIMENS. RAMK.

ASIA. —
Parmeliopsis (Nyl. ex Stizenb.) Nyl.; Parmeliaceae

Parmeliopsis ambigua (Wulfen) Nyl., Lich. Lapp. Orient.:
121 (1866).

THAILAND. Lampang (Khuntan N.P.). Pooprang et al. (1999: 122).
RAMK.

ASIA. Nepal.

SPECIMENS.

Parmotrema A. Massal.; Parmeliaceae

Parmotrema abessinicum (Kremp.) Hale, Phytologia 28:
334 (1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).

Pooprang et al. (1999: 122).
SPECIMENS. RAMK.

ASIA. India, Malaysia.

Parmotrema amaniense (J. Steiner & Zahlbr.) Krog &
Swinscow, Lichenologist 15: 129 (1983).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).

Wolseley & Aguirre-Hudson (1997a: 334).
SPECIMENS. BKF, BM.

ASIA. India.

Parmotrema andinum (Miill. Arg.) Hale, Phytologia 28:
334 (1974).

Parmelia andina Miill. Arg.

42

THAILAND. Chiang Mai (Doi Suthep, Chiang Dao N.P.). Hale
(196Sa: 236, as Parmelia andina).

SPECIMENS. BM, CMU, TNS.

ASIA. India, Malaysia.

Parmotrema austrosinense (Zahlbr.) Hale, Phytologia 28:
335 (1974).

THAILAND. Phitsanulok (Lan Hin Taek, Phu Hin Rongkla N.P.).
Moon et al. (2000a: 103).

SPECIMENS. TNS.

ASIA. China.

Parmotrema chinense (Osbeck) Hale & Ahti, Taxon 35:
133 (1986).

Parmelia perlata Ach.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Trat (Koh Chang I.). Vainio (1909: 104, as

Parmelia perlata Ach.).
SPECIMENS. BM, CMU.

ASIA.
Taiwan.

China, Malaysia, Papua New Guinea, Philippines, Sri Lanka,

Parmotrema corniculans (Ny1.) Hale, Phytologia 28: 335
(1974).

THAILAND. Chiang Mai (Doi Inthanon N.P.). Pooprang et al.

(1999: 122).
SPECIMEN. RAMK.

ASIA.
pines.

Indonesia, Laos, Malaysia, Papua New Guinea, Philip-

Parmotrema crinitum (Ach.) M. Choisy, Bull. Mens. Soc.
Linn. Soc. Bot. Lyon 21: 175 (1952).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Mongkonsuk et al. (1996: 17).

SPECIMEN. Queen Sirikit Botanical Garden Herbarium.

ASIA. China, India, Papua New Guinea, Sri Lanka, Taiwan.

Parmotrema cristiferum (Taylor) Hale, Phytologia 28: 335
(1974).

Parmelia andina Taylor.

THAILAND. Chiang Mai (Chiang Dao N.P., Doi Suthep N.P.),
Nakhon Nayok (Khao Yai N.P.), Udaun, Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Hale (1965b: 241, as Parmelia andina),
Boonpragob et al. (1999: 217), Moon et al. (2000a: 103).

SPECIMENS. BM, TNS, US.

ASIA. China, India, Indonesia, Malaysia, Myanmar, Papua New
Guinea, Philippines, Sri Lanka, Taiwan.

Parmotrema dilatatum (Vain.) Hale, Phytologia 28: 335
(1974).

THAILAND. Chiang Mai (Doi Suthep N.P., Queen Sirikit Botanical
Garden), Nakhon Ratchasima (Khao Yai N.P). Mongkonsuk et al.
(1996: 17), Boonpragob et al. (1999: 217), Moon et al. (2000a: 103).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

SPECIMENS. RAMK, TNS, Queen Sirikit Botanical Garden
Herbarium.
ASIA. India, Indonesia, Malaysia, Papua New Guinea, Sri Lanka.

Parmotrema dominicanum (Vain.) Hale, Phytologia 28:
335 (1974).

THAILAND. Loei (Phu Kradung). Moon et al. (2000a: 103).
SPECIMENS. TNS.

ASIA. —

Parmotrema elacinatulum (Kurok.) Streimann, Biblioth.
Lichenol. 22: 94 (1986).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Pooprang et al.

(1999: 123).
SPECIMEN. RAMK.

ASIA. Papua New Guinea.

Parmotrema eunetum (Stirt.) Hale, Phytologia 28: 336
(1974).

THAILAND. Chiang Mai (Doi Suthep N.P., Maetang). Moon et al.

(2000a: 103).
SPECIMENS. TNS.

ASIA. Sri Lanka.

Parmotrema eurysacum (Hue) Hale, Phytologia 28: 336
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).

Mongkonsuk et al. (1996: 17).
SPECIMEN. Queen Sirikit Botanical Garden Herbarium.

ASIA. —

Parmotrema explanatum (Hale) Hale, Phytologia 28: 336
(1974).

Parmelia explanata Hale.

THAILAND. Loei (Phu Kradung). Hale (1965b: 199, as Parmelia

explanata), Moon et al. (2000a: 103).
SPECIMENS. TNS (holotype), US (isotype).

ASIA. —

Parmotrema gardneri (C.W. Dodge) Sérus., Bryologist 87:
5 (1984).

Parmelia gardneri C.W. Dodge.

THAILAND. Chiang Mai (Doi Inthanon N.P.), Trat (Koh Chang I.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Vainio (1909:
105, as Parmelia gardneri), Wolseley & Aguirre-Hudson (1997a:
334; 1997b: 355).

SPECIMENS. BM, TUR-V.
ASIA. Papua New Guinea.

Parmotrema hababianum (Gyeln.) Hale, Phytologia 28:
336 (1974).

CATALOGUE OF THE LICHENS OF THAILAND

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Pooprang et al. (1999: 125).

SPECIMEN. RAMK.

ASIA. China, India, Sri Lanka.

Parmotrema hypotropum (Ny1.) Hale, Phytologia 28: 337
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Mongkonsuk et al. (1996: 17).

SPECIMEN. RAMK.

ASIA. —

Parmotrema incrassatum Hale ex DePriest & B.W. Hale,
Mycotaxon 67: 207 (1998).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Pooprang et al. (1999: 123).

SPECIMENS. Queen Sirikit Botanical Garden Herbarium.

ASIA. China (incl. Hong Kong).

Parmotrema lobulascens (J. Steiner) Hale, Phytologia 28:
337 (1974).

THAILAND. Chiang Mai (Doi Inthanon N.P.), Nakhon Nayok (Khao
Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Boonpragob et al. (1999: 217).

SPECIMEN. BM, RAMK.

ASIA. —

Parmotrema maclayanum (Miill. Arg.) Hale, Phytologia
28: 337 (1974).

Parmelia breviciliata Hale.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Queen Sirikit Botanical Garden), Loei (Ban Sui, Phu Kradung, Phu
Luang), Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Kurokawa & Kashiwadani (1988: 4, as Parmelia breviciliata),
Moon et al. (2000a: 103).

BM, TNS.

ASIA. Papua New Guinea.

SPECIMENS.

Parmotrema mellissii (C.W. Dodge) Hale, Phytologia 28:
337 (1974).

THAILAND. Loei (Phu Kradung). Moon et al. (2000a: 103).
SPECIMENS. TNS.

ASIA. China, India, Papua New Guinea, Taiwan.

Parmotrema nanfongense (Kurok.) DePriest & B.W. Hale,
Mycotaxon 67: 204 (1998).

THAILAND. Chiang Mai (Doi Suthep N.P.). Moon et al. (2000a:
104).

SPECIMEN. TNS.

ASIA. Papua New Guinea, Taiwan.

Parmotrema nilgherrense (Ny1.) Hale, Phytologia 28: 338
(1974).

43

Parmelia nilgherrensis Ny.
Parmelia siamensis Vain.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Lampang (Doi Khun Tan N.P.), Nakhon Nayok (Khao Yai N.P.), Tak
(Ton Krabakyai N.P.), Yala (Banglang Reservoir). Vainio (1921: 37,
as Parmelia siamensis), Hale (1965a: 333, as P. nilgherrensis),

Wolseley & Aguirre-Hudson (1997a: 334).
SPECIMENS. BM, TUR-V (holotype of Parmelia siamensis).

ASIA. China, India, Papua New Guinea, Philippines, Sri Lanka.

Parmotrema overeemii (Zahlbr.) Elix, Australas. Lichenol.
42: 23 (1998).

THAILAND. Chiang Mai, Lampang (Doi Khun Tan N.P.), Loei
(Phu Kradung N.P.), Nakhon Ratchasima (Khao Yai N.P.), Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Pooprang et al.

(1999: 123).
SPECIMENS. BKF, BM, CMU, RAMK, TNS, TUR-V.

ASIA. Indonesia, Papua New Guinea.

Parmotrema pancheri (Hue) Hale, Phytologia 28: 338
(1974).
Parmelia pancheri Hue.

THAILAND. Chiang Mai (Doi Suthep). Hale (1965b: 236, as
Parmelia pancheri), Moon et al. (2000a: 104).

SPECIMENS. TNS, US.

ASIA. —

Parmotrema permutatum (Stirt.) Hale, Phytologia 28: 338
(1974).

Parmelia permutata Stirt.

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden). Hale
(1965b: 302, as Parmelia permutata), Kashiwadani (1975: 81),

Moon et al. (2000a: 104).
SPECIMEN. TNS.

ASIA. India, Indonesia, Malaysia, Papua New Guinea.

Parmotrema planatilobatum (Hale) Hale, Phytologia 28:
338 (1974).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Pooprang et al.

(1999: 124).
SPECIMEN. RAMK.

ASIA. India, Indonesia, Malaysia, Sri Lanka.

Parmotrema platyphyllinum (Vain.) Elix, Australas.
Lichenol. 42: 23 (1998).

Parmelia platyphyllina Vain.
Parmelia zollingeri auct. non Hepp.

THAILAND. Phitsanulok (Lan Hin Taek), Trat (Koh Chang I.).
Vainio (1909: 106, as Parmelia platyphyllina), Hale (1965b: 267, as

P. zollingeri), Moon et al. (2000a: 104).
SPECIMENS. TNS, TUR-V.

ASIA. —

44

Parmotrema poolii (C.W. Dodge) Krog & Swinscow,
Lichenologist 15: 130 (1983).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden),
Phitsanulok (Lan Hin Taek). Moon et al. (2000a: 104).

SPECIMENS. TNS.

ASIA. Papua New Guinea.

Parmotrema praesorediosum (Ny1.) Hale, Phytologia 28:
338 (1974).

THAILAND. Chiang Mai (Doi Saget, Maesa, Doi Suthep N.P.,
Queen Sirikit Botanical Garden), Songkla. David et al. (1990:
1299), Mongkonsuk et al. (1996: 17), Moon et al. (2000a: 104)

SPECIMENS. CANB, TNS, US, Queen Sirikit Botanical Garden

Herbarium.

ASIA. China (incl. Hong Kong), India, Malaysia, Papua New
Guinea, Singapore, Sri Lanka, Taiwan.

Parmotrema procerum (J. Steiner & Zahlbr.) Hale,
Phytologia 28: 338 (1974).

Parmelia procera J. Steiner & Zahlbr.

THAILAND. Chiang Mai (Doi Suthep N.P.). Hale (1965b: 303, as
Parmelia procera).

SPECIMENS. TNS, US.

ASIA. —

Parmotrema pseudocrinitum (des Abbayes) Hale,
Phytologia 28: 338 (1974).

THAILAND. Loei (near Phu Kradung). Kurokawa & Kashiwadani
(1988: 6), Moon et al. (2000a: 104).

SPECIMEN. TNS.

ASIA. —

Parmotrema pseudonilgherrense (Asahina) Hale,
Mycotaxon 5: 441 (1977).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Pooprang et al. (1999: 124).

SPECIMEN. RAMK.

ASIA. China, India, Nepal.

Parmotrema rampoddense (Ny1.) Hale, Phytologia 28: 338
(1974).

THAILAND. Chiang Mai (Doi Suthep N.P., Queen Sirikit Botanical
Garden, Maetang), Loei (Phu Kradung N.P.), Phitsanulok (Lan Hin
Taek, Phu Hin Rongkla, N.P.), Kurokawa & Kashiwadani (1988: 6),

Mongkonsuk et al. (1996: 17), Moon et al. (2000a: 104).
SPECIMENS. BM, TNS.

ASIA.
Taiwan.

China, India, Indonesia, Papua New Guinea, Sri Lanka,

Parmotrema robustum (Degel.) Hale, Phytologia 28: 338
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden),

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Phitsanulok (Lan Hin Taek, Phu Hin Rongkla, N.P.). Mongkonsuk et

al. (1996: 17), Moon et al. (2000a: 104).
SPECIMENS. TNS, Queen Sirikit Botanical Garden Herbarium.

ASIA. India, Papua New Guinea.

Parmotrema rubromarginatum Elix & Pooprang, in
Pooprang et al., Mycotaxon 72: 115 (1999).

THAILAND. Loei, Nakhon Ratchasima. Pooprang et al. (1999:

115).
SPECIMENS. CANB (isotype), RAMK (incl. holotype).

ASIA. —

Parmotrema saccatilobum (Taylor) Hale, Phytologia 28:
339 (1974).

THAILAND. Phuket (Ban Pae). Moon et al. (2000a: 104).
SPECIMEN. TNS.

ASIA. China, India, Papua New Guinea, Singapore, Taiwan.

Parmotrema sancti-angeli (Lynge) Hale, Phytologia 28:
339 (1974).

Parmelia sancti-angeli Lynge.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P., Doi
Chieng Dao N.P., Maetang), Loei (near Phu Kradung), Phitsanulok
(Lan Hin Taek, Phu Hin Rongkla, N.P.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Hale (1965b: 307, as Parmelia sancti-

angeli), Moon et al. (2000a: 104).
SPECIMENS. BM, TNS, US.

ASIA. China, India, Indonesia, Laos, Papua New Guinea, Sri
Lanka, Taiwan.

Parmotrema stuppeum (Taylor) Hale, Phytologia 28: 339
(1974).

Parmelia perforata f. claudelii (Harm.) Vain.
Parmelia maxima Hue.

THAILAND. Chiang Mai (Doi Suthep N.P.), Phuket (Ban Pae), Trat
(Koh Chang I.). Vainio (1909: 105, as Parmelia perforata f. claudelii),

des Abbayes (1964, as P. maxima), Moon et al. (2000a: 104).
SPECIMENS. TNS, TUR-V.

ASIA. India.

Parmotrema subarnoldii (des Abbayes) Hale, Phytologia
28: 339 (1974).

THAILAND. Loei, Nakhon Ratchasima. Pooprang et al. (1999:

124)
SPECIMENS. RAMK.

ASIA. India, Indonesia, Papua New Guinea.

Parmotrema submarginale (Michx.) DePriest & B.W.
Hale, Mycotaxon 67: 208 (1998).

Parmotrema michauxianum (Zahlbr.) Hale.

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden).
Mongkonsuk et al. (1996: 17).

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. Queen Sirikit Botanical Garden Herbarium.

ASIA. —

Parmotrema sulphuratum (Nees & Flot.) Hale, Phytologia
28: 339 (1974).

THAILAND. Loei (Na Haew N.P.), Nakhon Sri Thammarat ( Khao

Luang). Pooprang et al. (1999: 124).
SPECIMENS. BM, RAMK.

ASIA. Indonesia, Malaysia, Papua New Guinea.

Parmotrema thailandicum Elix & Pooprang, in Pooprang
et al., Mycotaxon 72: 117 (1999).

THAILAND. Phitsanulok (Phu Hin Rongkla N.P.).

SPECIMENS. CANB (isotype), RAMK (holotype).

ASIA. —

tse tinctorum (Nyl.) Hale, Phytologia 28: 339

Parmelia tinctorum Ny.
Parmelia coralloidea (Meyen & Flot.) Vain.

THAILAND. Chumphon (Koh Tao I.), Chiang Mai (Doi Suthep
N.P., Doi Inthanon N.P., Doi Pacho, Chiang Dao N.P., Queen Sirikit
Botanical Garden, Maetang), Chiang Rai, Loei (Ban Sui, Pha Nok
Kao), Phitsanulok (Lan Hin Taek, Phu Hin Rongkla, N.P.), Udon-
Thani. Vainio (1921: 37, as Parmelia coralloidea), Sato (1962: 4, as
P. tinctorum), Mongkonsuk et al. (1996: 17), Wolseley & Aguirre-
Hudson (1997a: 334; 1997b: 355), Moon et al. (2000a: 104).

SPECIMENS. BKF,BM,CMU,RAMK, TNS, TUR-V, Queen Sirikit
Botanical Garden Herbarium.

ASIA. China (incl. Hong Kong), Indonesia, India, Laos, Malaysia,
Papua New Guinea, Philippines, Singapore, Sri Lanka, Taiwan.
Parmotrema Zollingeri (Hepp) Hale, Phytologia 28: 339
(1974).

THAILAND. Chiang Mai (Queen Sirikit Botanical Garden), Loei

(Ban Su). Moon et al. (2000a: 105).
SPECIMENS. TNS.

ASIA. China, India, Indonesia, Philippines, Taiwan.
Pertusaria DC.; Pertusariaceae

Pertusaria asiana Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 44 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 44);
Oshio (1976: 116).

SPECIMEN. TUR-V (holotype).

ASIA. —

Pertusaria bengalensis Vain., Hedwigia 46: 169 (1907).

THAILAND. Trat (Koh Chang I.). Vainio (1907: 169; 1909: 110),

Oshio (1976: 116).
SPECIMEN. TUR-V (holotype).

ASIA. —

45

Pertusaria pustulata (Ach.) Duby, Bot. Gall. 2(2): 673
(1830).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 109), Archer (1997:

1:35):
SPECIMENS. TUR-V.

ASIA. China, India.

Pertusaria sphaerulifera Vain., Hedwigia 46: 169 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1907: 169; 1909: 109).
SPECIMEN. 'TUR-V (holotype).

ASIA. —

Pertusaria subnegans Vain., Hedwigia 46: 170 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1907: 170; 1909: 110).

SPECIMEN. TUR-V (holotype).

ASIA. —

Phaeographina Mill. Arg.; Graphidaceae

Phaeographina caesiopruinosa (Fée) Miill. Arg., Mém.
Soc. Phys. Hist. Nat. Genéve 29(8): 49 (1887).

Graphis caesiopruinosa Fée.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 49, as
Graphis caesiopruinosa).

SPECIMEN. TUR-V.

ASIA. India, Malaysia, Sri Lanka.

Phaeographina chlorocarpoides (Ny}1.) Zahlbr., Cat. Lich.
Univ. 2: 321 (1924).

Graphis chlorocarpoides Ny].

THAILAND. Chaing Mai (Doi Suthep), Trat (Koh Chang I.). Vainio
(1909: 124; 1921: 49, both as Graphis chlorocarpoides),

Kashiwadani (1977: 45).
SPECIMENS. TUR-V, TNS.

ASIA. China, Indonesia, Philippines, Taiwan.

Phaeographina chrysenteron (Mont.) Mill. Arg.
Hedwigia 30: 52 (1891).

THAILAND. Chumphon (Koh Tao I.). Vainio (1921: 49), Paulson
(1930: 100).

SPECIMEN. TUR-V, BM.

ASIA. China, India, Indonesia, Sri Lanka.

Phaeographina heterocarpoides (Ny1.) Zahlbr., Cat. Lich.
Univ. 2: 439 (1924).

Graphis heterocarpoides Ny).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 123, as Graphis
heterocarpoides).

46
SPECIMEN. TUR-V.

ASIA. —

Phaeographina persimilis (Vain.) Zahlbr., Cat. Lich. Univ.
2: 442 (1923).

Graphis persimilis Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 123, as Graphis
persimilis).

SPECIMENS. TUR-V.

ASIA. —

Phaeographina scalpturata (Ach.) Mill. Arg., Flora 65:
399 (1882).

Graphis scalpturata Ach.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 49, as Graphis

scalpturata).
SPECIMENS. TUR-V.

ASIA. China (Hong Kong), India, Indonesia.

Phaeographina schmidtii (Vain.) Zahlbr., Cat. Lich. Univ.
2: 446 (1923).
Graphis schmidtii Vain.

THAILAND. ‘Trat (Koh Chang I.). Vainio (1907: 176; 1909: 124,
both as Graphis schmidtii).

SPECIMEN. TUR-V.

ASIA. —

Phaeographina subintricata (Kremp.) Mill. Arg., Bull.
Herb. Boissier 3: 320 (1895).

Graphis subintricata Vain.

THAILAND. Trat (Koh Kong I.). Vainio (1909: 126, as Graphis
subintricata).

SPECIMEN. TUR-V.

ASIA. China, India.

Phaeographina subrigida (Ny1.) Zahlbr., Cat. Lich. Univ.
2: 446 (1923).

Graphis subrigida Ny].

THAILAND. Chiang Mai (Doi Suthep). Vainio (1909: 122, as
Graphis subrigida).

SPECIMEN. TUR-V.

ASIA. —

Phaeographis Miill. Arg.; Graphidaceae
Phaeographis exaltata (Mont. & Bosch) Miill. Arg., Flora
65: 336 (1882).

Graphis diversa Nyl.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 129, as Graphis
diversa).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
SPECIMEN. TUR-V.

ASIA. Indonesia, Philippines.

Phaeographis nylanderi (Vain.) Zahlbr., Cat. Lich. Univ. 2:
382 (1923).

Graphis nylanderi Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 130, as Graphis
nylanderi).

SPECIMEN. TUR-V.

ASIA. —

Phaeographis rugulosa (Vain.) Zahlbr., Cat. Lich Univ. 2:
444 (1923).

Graphis rugulosa Vain.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 49, as
Graphis rugulosa).

SPECIMEN. ‘TUR-V (holotype).

ASIA. —

Phaeographis sericea (Eschw.) Miill. Arg., Flora 71: 523
(1888).

Graphis sericea (Eschw.) Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 130, as Graphis
sericea).

SPECIMEN. TUR-V.

ASIA. China (Hong Kong).

Phaeographis subtigrina (Vain.) Zahlbr., Cat. Lich. Univ.
2: 387 (1923),

Graphis subtigrina Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 131, as Graphis
subtigrina), Patwardhan & Kulkarni (1977b: 721), Wirth & Hale
(1978: 29).

SPECIMEN. TUR-V.

ASIA. India.

Phaeophyscia Moberg; Physciaceae

Phaeophyscia hispidula (Ach.) Essl., Mycotaxon 7: 301
(1978).

Physcia setosa (Ach.) Nyl.

THAILAND. Chiang Mai (Doi Inthanon N.P.), Chumphon (Koh
Tao I.). Vainio (1909: 130), Paulson (1930: 100, as Physcia setosa).

SPECIMENS. BM.

ASIA. China, India, Indonesia, Malaysia.

Phlyctis Wallr.; Phlyctidaceae

[Phlyctis| Phlyctella himalayensis Nyl., Lich. Nov.-Zel.: 73
(1888).

Phlyctis polyphragmia (Nyl.) Vain.

CATALOGUE OF THE LICHENS OF THAILAND

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 43, as
Phlyctis polyphragmia).

SPECIMEN. TUR-V.

ASIA. India, Nepal.

[Phlyctis| Phlyctella neozelandica Ny|., Flora 52: 121
(1869).

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 100, as
Phlyctella neozelandica).

SPECIMEN. BM.

ASIA. —
Phyllopsora Miill. Arg.; Phyllopsoraceae
Phyllopsora furfuracea (Pers.) Zahlbr., in Engler & Prantl,

Nat. Pflanzenfam. 1(1): 138 (1905).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. BM, RAMK.

ASIA. Papua New Guinea.
Physcia (Schreb.) Michx.; Physciaceae
Physcia sorediosa (Vain.) Lynge, Natur. Skrift. I. Math.-

Naturvid. Kl. 16: 27 (1924).

THAILAND. Chumphon (Koh Tao I.), Uthai Thani (Huay Kha
Khaeng Wildlife Sanctuary). Paulson (1930: 100).

SPECIMEN. BM (not located).

ASIA. Malaysia, Papua New Guinea.

Physcidia Tuck.; Phyllopsoraceae

Physcidia cylindrophora (Taylor) Hue, Bull. Soc. Linn.
Normandie 6(1): 97 (1908).

THAILAND. Chiang Mai (Doi Suthep), Nakhon Ratchasima (Khao
Yai N.P.). Kalb & Elix (1995: 286), Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK.

ASIA. India, Malaysia, Papua New Guinea, Philippines.

Physcidia wrightii (Tuck.) Tuck., Proc. Amer. Acad. Arts 5:
400 (1862).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Loei (Phu Kradung N.P.), Nakhon Ratchasima (Khao Yai N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Kurokawa &
Kashiwadani (1988: 15), Kalb & Elix (1995: 291), Wolseley &
Aguirre-Hudson (1997a: 334), Boonpragob et al. (1998: 217).

BM, RAMK, TNS.
ASIA. Malaysia.

SPECIMENS.

Physma A. Massal.; Collemataceae

Physma byrsaeum (Ach.) Tuck., Syn. N. Amer. Lich. 1: 115
(1882).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay

47

Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson
(1995: 327).

SPECIMEN. BM.

ASIA. China (Hong Kong), India, Malaysia, Papua New Guinea.

Physma plicatum (Pers.) Hue, Bull. Soc. Linn. Normandie,
sér. 5, 9: 136 (1906).
Trat (Koh Chang I.). Vainio (1909: 116).

C, TUR-V.

THAILAND.
SPECIMENS.

ASIA. Indonesia.

Platismatia W.L. Culb & C.F. Culb.; Parmeliaceae
Platismatia erosa W.L. Culb & C.F. Culb., Contr. U.S.
Natl. Herb. 34: 526 (1968).

THAILAND. Chiang Mai Province (Doi Inthanon N.P.). Wolseley
& Aguirre-Hudson (1997a: 337).

SPECIMEN. BM.

ASIA. China, India, Indonesia, Nepal, Philippines, Taiwan, Viet-
nam.

Polymeridium (Miill. Arg.) R.C. Harris;
Trypetheliaceae

Polymeridium albidum (Miill. Arg.) R.C. Harris, Acta
Amazonica 14 (Suppl.): 69 (1984) [‘1986’].
Arthopyrenia siamensis Vain.

THAILAND. Chiang Mai (Doi Suthep). Vainio (1921: 55, as
Arthopyrenia siamensis), Harris (1993: 624).

SPECIMEN. TUR-V (holotype of Arthopyrenia siamensis).
ASIA. Malaysia, Papua New Guinea, Philippines.
Polymeridium catapastum (Nyl.) R.C. Harris, Acta
Amazonica 14 (Suppl.): 70 (1984) [‘1986’].

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Vongshewarat
et al. (1999: 231).

SPECIMENS. CANB, RAMK.

ASIA. Malaysia, Papua New Guinea.

Polymeridium quinqueseptatum (Nyl.) R.C. Harris, in
Tucker & Harris, Bryologist 83: 12 (1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Vongshewarat

et al; (19992231).
SPECIMENS. CANB, RAMK.

ASIA. China (Hong Kong), India, Malaysia, Philippines.

Porina Ach.; Trichotheliaceae

Porina aff. albicera (Kremp.) Overeem, Bull. Jard. Bot.
Buitenzorg, sér. 3, 4: 112 (1922).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

48
SPECIMENS. CANB.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka, Vietnam.

Porina americana Fée, Essai Crypt. Ecorc. Offic.: 83
(1824).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB.

ASIA. —

Porina bellendenica Miill. Arg., Hedwigia 30: 56 (1891).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. CANB.

ASIA. —

Porina conica R. Sant., Symb. Bot. Upsal. 12(1): 232
(1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. CANB.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka, Vietnam.

Porina corruscans (Rehm) R. Sant., Symb. Bot. Upsal.
12(1);:223'G952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB.

ASIA. India, Indonesia, Papua New Guinea, Philippines.

Porina eminentior (Ny1.) P.M. McCarthy, Lichenologist
32: 42 (2000).
Clathroporina eminentior (Nyl.) Mill. Arg.

THAILAND. Chiang Mai (Doi Inthanon), Uthai Thani (Khao Nang
Rum, Khlong Plou). McCarthy (1995: 329, as Clathroporina
eminentior).

SPECIMENS. BM.

ASIA. Papua New Guinea.

Porina epiphylla (Fée) Fée, Essai Crypt. Ecorc., Suppl. 1:
76 (1837).
Micropeltis schmidtiana Rostrup.

THAILAND. Nakhon Nayok (Khao Yai N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Trat (Koh Chang I.). Rostrup (1902: 211); Santesson

(1952: 234), Boonpragob et al. (1998: 217).
SPECIMENS. CANB.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea,
Philippines, Sri Lanka, Vietnam.

Porina erawanensis P.M. McCarthy & Vongshewarat, in
Vongshewarat et al., Mycotaxon 71: 231 (1999).

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

THAILAND. Kanchanaburi (Erawan N.P.). Vongshewarat et al.
(1999: 231).

SPECIMENS. CANB (holotype), RAMK (isotype).

ASIA. —

Porina internigrans (Nyl1.) Miill. Arg., Rep. Australas.
Assoc. Advancem. Sci. 1895: 452 (1895).

THAILAND. Sukothai (Ramkhamhaeng N.P.). Vongshewarat et al.
(1999: 233).
SPECIMEN. RAMK.

ASIA. India, Papua New Guinea.

Porina kansriae P.M. McCarthy, Lichenologist 31: 239
(1999).

THAILAND. NakhonRatchasima(Khao YaiN.P.). Boonpragobetal.

(1998: 217, as Porina aff. ulceratula Zahlbr.), McCarthy (1999: 239).
SPECIMENS. CANB (incl. holotype).

ASIA. —

Porina lucida R. Sant., Symb. Bot. Upsal. 12(1): 240
(1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB.

ASIA.
Lanka.

Indonesia, Malaysia, Papua New Guinea, Philippines, Sri

Porina mastoidea (Ach.) Miill. Arg., Bot. Jahrb. Syst. 6:
399 (1885).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 217).
SPECIMENS. CANB.

ASIA. India, Malaysia, Papua New Guinea, Philippines.

Porina aff. octomera (Miill. Arg.) F. Schill., Hedwigia 67:
274 (1927).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

CANB.

ASIA. Vietnam.

SPECIMENS.

Porina rufula (Kremp.) Vain., Acta Soc. Fauna Fl. Fenn.
7(2): 227 (1890).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. CANB.

AsIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka.

Porina siamensis P.M. McCarthy, Lichenologist 31: 242
(1999).

THAILAND. Nakhon Nayok (Khao Yai N.P.), Nakhon Ratchasima
(Khao Yai N.P.). McCarthy (1999: 242).

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. CANB (incl. holotype).

ASIA. —

Porina tetracerae (Ach.) Mill. Arg., Bot. Jahrb. Syst. 6:
401 (1885).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMENS. CANB.

ASIA. China, India, Malaysia, Papua New Guinea, Sri Lanka.
Porina aff. vezdae Liicking, Nova Hedwigia 52: 285
(1991).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 212).
SPECIMENS. CANB.

ASIA. —

Porina virescens (Kremp.) Miill. Arg., Flora 66: 331
(1883).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 212).
SPECIMENS. CANB.

ASIA. India, Indonesia, Malaysia, Papua New Guinea, Philip-
pines, Sri Lanka, Vietnam.

Porina wolseleyae P.M. McCarthy, Lichenologist 31: 244
(1999).

THAILAND. Nakhon Nayok (Khao Yai N.P.). McCarthy (1999:

244).
SPECIMENS. CANB (holotype).

ASIA. —

Pseudocyphellaria Vain.; Lobariaceae

Pseudocyphellaria argyracea (Delise) Vain., Hedwigia 37:
35 (1898).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 217).

SPECIMEN. RAMK.
ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua

New Guinea, Philippines, Sri Lanka, Taiwan.

Pseudocyphellaria aurata (Sm.) Vain., Acta Soc. Fauna
FI. Fenn. 7: 183 (1890).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &
Aguirre-Hudson (1997a: 334).

SPECIMENS. BKF.
ASIA. China, India, Indonesia, Malaysia, Philippines, Taiwan.

Pseudocyphellaria beccarii (Kremp.) D.J. Galloway, Bull.
nat. Hist. Mus. London (Bot.) 24: 119 (1994).

49

THAILAND. Chiang Mai (Doi Inthanon N.P.). Wolseley & Aguirre-
Hudson (1997a: 334).

SPECIMEN. BM.

ASIA. Malaysia.

Pseudopyrenula Miill. Arg.; Trypetheliaceae
Pseudopyrenula diluta var. degenerans Vain., Bot. Tidsskr.
29: 148 (1909).

Pseudopyrenula endoxanthoides Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 148), Harris (1998:
138).

SPECIMEN. TUR-V (holotype).

ASIA. —

Pyrenula Ach.; Pyrenulaceae

Pyrenula anomala (Ach.) Vain., Ann. Acad. Sci. Fenn., set.
A, 6(7): 189 (1915).

THAILAND. Buri Ram (Khao Kadong), Chanthaburi (Numtokphliu
N.P.), Chiang Mai (Queen Sirikit Botanical Garden), Lumpang
(Khuntan), Sakon Nakhon (Phu Pan N.P.). Vongshewarat et al.

(1999: 234).
SPECIMENS. RAMK.

ASIA. India, Papua New Guinea, Philippines, Vietnam.

Pyrenula approximata Vain., Bot. Tidsskr. 29: 145 (1909).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 145).
SPECIMEN. TUR-V (holotype).

ASIA. —

Pyrenula aspistea Ach., Ges. Naturf. Freunde Berlin Mag.
Neuesten Entdeck. Gesammten Naturk. 6: 17 (1814).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 147).
SPECIMENS. RAMK, TUR-V.

ASIA. _ India, Indonesia, Papua New Guinea, Philippines, Sri Lanka.

Pyrenula confinis (Ny1.) R.C. Harris, More Florida
Lichens: 109 (1995).
Bottaria confinis (Nyl.) Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 143, as Bottaria

confinis).
SPECIMENS. RAMK, TUR-V.

ASIA. India, Papua New Guinea, Philippines.

Pyrenula feracissima Vain., Bot. Tidsskr. 29: 145 (1909).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 145).
SPECIMENS. TUR-V (holotype).

ASIA. —

Pyrenula glabriuscula (Ny}.) Vain., Bot. Tidsskr. 29: 146
(1909).

50
THAILAND. Trat (Koh Chang I.). Vainio (1909: 146).
SPECIMENS. TUR-V (holotype).

ASIA. China, Philippines.

Pyrenula leucostoma Ach., Syn. Meth. Lich.: 124 (1914).
Bottaria libricola (Feé) Vain.

THAILAND. Trat (Lem Ngob). Vainio (1909: 142, as Bottaria

libricola).
SPECIMENS. TUR-V.

ASIA. Indonesia, Papua New Guinea.

Pyrenula macularis (Zahlbr.) R.C. Harris, Mem. New York
Bot. Gard. 49: 94 (1989).
Rayong. Vongshewarat et al. (1999: 234).

RAMK.

THAILAND.
SPECIMENS.

ASIA. Papua New Guinea.

Pyrenula nitida (Weig.) Ach., Ges. Naturf. Freunde Berlin
Mag. Neuesten Entdeck. Gesammten Naturk. 6: 21 (1814).
Chumphon (Koh Tao I.). Paulson (1930: 100).

BM (not located).

THAILAND.
SPECIMEN.
ASIA. China, India, Indonesia, Sri Lanka.

Pyrenula parvinuclea (Meyen & Flot.) Aptroot, in Aptroot
et al., Biblioth. Lichenol. 64: 165 (1997).

Bottaria denudata (Nyl.) Vain.

THAILAND. Chachesoa, Samut Prakan (Muang Boran), Trat (Koh
Chang. I.). Vainio (1909: 143, as Bottaria denudata), Vongshewarat

et al. (1999: 234).
SPECIMENS. CANB, RAMK, TUR-V.

ASIA. India, Indonesia, Malaysia, Papua New Guinea.

Pyrenula subaggregata Miill. Arg., Bot. Jahrb. Syst. 6:
410 (1885).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 144).
SPECIMEN. TUR-V.

ASIA. — India.

Pyrenula subferruginea (Malme) R.C. Harris, Mem. New
York Bot. Gard. 49: 101 (1989).

THAILAND.
(1999: 235).

SPECIMEN. RAMK.

Kanchanaburi (Erawan N.P.). Vongshewarat et al.

ASIA. Papua New Guinea.

Pyrenula submarginata Vain., Bot. Tidsskr. 29: 146
(1909).

THAILAND. Trat (Koh Chang I.). Vainio (1909: 146).
SPECIMEN. TUR-V (holotype).

ASIA. —

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY
Pyrenula velatior Miill. Arg., Flora 68: 20 (1885).
THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 100).
SPECIMEN. BM.

ASIA. —
Pyrgillus Nyl.; Pyrenulaceae

Pyrgillus indicus (Kremp.) Aptroot, Biblioth. Lichenol. 44:
84 (1991).

Pyrgillocarpon indicum (Kremp.) Nadv.

THAILAND. Chiang Mai (Doi Inthanon, N.P., Doi Suthep N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &
Aguirre-Hudson (1997a: 334, 339, as Pyrgillocarpon indicum).

SPECIMENS. BM.

ASIA. India, Malaysia.
Pyrgillus javanicus (Mont. & Bosch) Nyl., Mém. Soc. Sci.
Nat. Cherbourg 5: 334 (1858).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-
Hudson (1997a: 334, 339), Boonpragob et al. (1998: 217).

SPECIMENS. BM, RAMK.

ASIA. Indonesia, Malaysia, Papua New Guinea, Philippines.

Pyrrhospora K6rb.; Lecanoraceae

Pyrrhospora russula (Ach.) Hafellner, in Kalb &
Hafellner, Herzogia 9: 86 (1992).

Lecidea russula Ach.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Chiang Dao N.P.), Nakhon Ratchasima (Khao Yai N.P.), Uthai
Thani (Huay Kha Khaeng Wildlife Sanctuary). Vainio (1921: 48, as
Lecidea russula), Sato (1962: 3, as L. russula), Hafellner (1993:
737), Wolseley & Aguirre-Hudson (1997a: 333, 339; 1997b: 356),

Boonpragob et al. (1998: 217).
SPECIMENS. BM, CMU, TNS, TUR-V.

ASIA. China (Hong Kong).

Pyxine Fr.; Physciaceae

Pyxine coccifera (Fée) Nyl., Mém. Soc. Imp. Sci. Nat.
Cherbourg 5: 108 (1857).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.),
Lampang (Doi Khun Tan N.P.), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Vainio (1921: 45), Wolseley & Aguirre-Hud-

son (1991: 172, 173; 1997a: 334; 1997b: 355).
SPECIMENS. BKF, BM, CMU, RAMK.

ASIA. India, Indonesia, Malaysia, Sri Lanka, Vietnam.

Pyxine consocians Vain., Philipp. J. Sci., Sect. C, 8: 109
(1913).

THAILAND. Chiang Mai, Uthai Thani. Wolseley & Aguirre-Hudson
(1997a: 334; 1997b: 355, 356).

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. BKF, BM, CMU.

ASIA. India, Malaysia, Papua New Guinea, Philippines.

Pyxine coralligera Malme, Bih. Kongl. Svenska Vetensk-
Akad. Handl. 23(3): 40(1897).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.),
Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary). Wolseley &

Aguirre-Hudson (1997a: 334; 1997b: 356).
SPECIMENS. BKF, BM, CMU.

ASIA. India, Papua New Guinea.

Pyxine retirugella Ny1., Ann. Sci. Nat. Bot., sér. 4, 11: 240
(1859).

Pyxine asiatica Vain.
Pyxine retirugella var. laevior Vain.
Pyxine copelandii Vain.

THAILAND. Trat (Koh Chang I.), Uthai Thani (Huay Kha Khaeng
Wildlife Sanctuary). Vainio (1909: 113, as Pyxine asiatica, P.

retirugella var. laevior and P. copelandii). Rogers (1986: 149).
SPECIMENS. BM, TUR-V.

ASIA. India, Malaysia, Papua New Guinea, Philippines.

Pyxine schmidtii Vain., Hedwigia 46: 170 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 112).
SPECIMEN. TUR-V (holotype).

ASIA. —

Ramalina Ach.; Ramalinaceae
Ramalina calicaris (L.) Rohl, Deutshl. Fl. 3(2): 139
(1813).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.). Sato
(1962: 6), Yoshimura (1978: 37).

SPECIMENS. C.

ASIA. China, India, Indonesia, Taiwan.

Ramalina conduplicans Vain., Suom. Elain-ja Kasvit.
Seuran Van. Julk. 1(3): 35 (1921).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).

Vainio (1921: 35), Awasthi (1960: 177), Sato (1962: 6).
SPECIMENS. TUR-V (syntypes).

ASIA. —

Ramalina farinacea (L.) Ach., Methodus: 263 (1803).

THAILAND. Chiang Mai (Chiang Dao N.P.), Chumphon (Koh Tao
I.), Trat (Koh Chang I.). Vainio (1909: 104), Paulson (1930: 100),
Yoshimura (1978: 37).

BM, C, TUR-V.

ASIA. China, Indonesia, Sri Lanka, Vietnam.

SPECIMENS.

Ramalina hossei Vain., Suom. Elain-ja Kasvit. Seuran Van.
Julk. 1(3): 36 (1921).

51

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 36),

Kashiwadani (1988: 129-133).
SPECIMENS. TNS, TUR-V (holotype).

ASIA. Taiwan.

Relicina (Hale & Kurok.) Hale; Parmeliaceae

Relicina abstrusa (Vain.) Hale, Phytologia 28: 484 (1974).

THAILAND. Nakhon Ratchasima (Khao Yai N.P), Loei (Phu
Kradung N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary). Culberson et al. (1981: 25), Boonpragob et al. (1999: 218),
Moon et al. (2000a: 105).

SPECIMENS. BM, RAMK, TNS.

ASIA. China, India, Indonesia, Malaysia, Philippines, Taiwan.
Relicina circumnodata (Ny1.) Hale, Phytologia 28: 484
(1974).

THAILAND. Phitsanulok (Lan Hin Taek, Phu Hin Rongkla, N.P.),
Phuket (Ton Sai). Moon et al. (2000a: 105).

SPECIMENS. TNS.

ASIA. Malaysia, Philippines.

Relicina subabstrusa (Gyeln.) Hale, Phytologia 28: 485
(1974).

THAILAND. Nakhon Ratchasima (Khao Yai N.P). Boonpragob et

al. (1999: 218).
SPECIMENS. RAMK.

ASIA. Malaysia, Philippines, Taiwan.

Relicina sublanea (Kurok.) Hale, Phytologia 28: 485
(1975).

THAILAND. Yala (Wang Pra, Bang-Lang N.P.). Elix (1996: 120).
SPECIMENS. PSU.

ASIA. Indonesia, Malaysia, Philippines, Sri Lanka.

Relicina sublimbata (Ny1.) Hale, Phytologia 28: 485
(1974).

THAILAND. Loei (Phu Kradung). Hale (1975: 31), Wolseley &

Aguirre-Hudson (1995: 328), Moon et al. (2000a: 103).
SPECIMENS. BM, TNS, US.

ASIA. Myanmar.
Relicinopsis Elix & Verdon; Parmeliaceae

Relicinopsis intertexta (Mont. & Bosch) Elix & Verdon,
Mycotaxon 27: 281 (1986).

Parmelia ecoronata Ny\.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 107, as Parmelia

ecoronata).
SPECIMENS. BM, TUR-V.

ASIA. India, Indonesia, Philippines, Malaysia, Papua New Guinea.

52

Relicinopsis rahengensis (Vain.) Elix & Verdon,
Mycotaxon 27: 282 (1986).

Parmelia rahengensis Vain.
Pseudoparmelia rahengensis (Vain.) Hale.

THAILAND. Nong Boa near Raheng, Chiang Mai (Doi Saget, Doi
Suthep N.P., Doi Inthanon N.P., Chieng Dao N.P., Queen Sirikit
Botanical Garden, Maetang), Lampang (Doi Khun Tan N.P.),
Phitsanulok (Lan Hin Taek, Phu Hin Rongkla, N.P.), Uthai Thani
(Huay Kha Khaeng Wildlife Sanctuary). Vainio (1921: 39, as
Parmelia rahengensis), Hale (1976a: 42, as Pseudoparmelia
rahengensis), Wolseley & Aguirre-Hudson (1995: 329; 1997a: 334;
1997b: 335, 366), Moon et al. (2000a: 105).

SPECIMENS. BKF, BM, CMU, TNS, US, TUR-V (lectotype).
ASIA. —

Relicinopsis malaccensis (Ny1.) Elix & Verdon, Mycotaxon
27: 282 (1986).

THAILAND. Loei (Nahaew N.P.), Phitsanulok (Phu Hin Rongkla

N.P.). Pooprang et al. (1999: 125).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Malaysia, Philippines, Sri Lanka.

Rimelia Hale & A. Fletcher; Parmeliaceae
Rimelia reticulata (Taylor) Hale & A. Fletcher, Bryologist
93: 28 (1990).

Parmelia cetrata f. soredifera Vain.
Rimelia clavulifera (Rasénen) Kurok.

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.,
Maetang), Chiang Rai (Ban Lang), Loei (Phu Kradung), Phitsanulok
(Lan Hin Taek, Phu Hin Rongkla, N.P.). Vainio (1921: 37, as
Parmelia cetrata f. soredifera), Sato (1962: 3), Wolseley &
Aguirre-Hudson (1995: 329), Moon et al. (2000a: 105, as Rimelia
clavulifera).

SPECIMENS. BKF, BM, CMU, TNS, TUR-V.

ASIA. China, India, Indonesia, Malaysia, Papua New Guinea, Sri
Lanka, Taiwan.

Roccella DC.; Roccellaceae

Roccella phycopsis Ach., Lichenogr. Universalis: 440
(1803).

THAILAND. Chumphon (Koh Tao I.). Paulson (1930: 100).
SPECIMENS. BM.

ASIA. =

Sarcographa Fée; Graphidaceae

Sarcographa labyrinthica Miill. Arg., Mém. Soc. Phys.
Hist. Nat. Genéve 29(8): 46 (1887).

Graphis labyrinthica (Ach.) Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 131, as Graphis
labyrinthica).
SPECIMENS. BM, TUR-V.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

ASIA. Indonesia, Malaysia, Sri Lanka.

Sarcographa medusulina (Nyl.) Miill. Arg., Flora 70: 77
(1887).

THAILAND. Chiang Mai (Doi Suthep N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife Sanctu-
ary). Vainio (1909: 131), Wolseley & Aguirre-Hudson (1997a: 334;

1997b: 355), Boonpragob et al. (1998: 218).
SPECIMENS. BM, RAMK.

ASIA. —

Spilonema Bornet; Coccocarpiaceae

Spilonema schmidtii (Vain.) Henssen, Symb. Bot. Upsal.
18: 97 (1963).

Leptogidium mooreii (Hepp) Nyl.
Leptodendriscum schmiadtii Vain.

THAILAND. Trat (Koh Chang I). Vainio (1909: 115; 1921: 34, as

Leptodendriscum schmidtii), Henssen (1963: 97).
SPECIMENS. 'TUR-V (lectotype, syntype).

ASIA. —

Sporopodium Mont.; Ectolechiaceae

Sporopodium hossei Vain., Suom. Elain-ja Kasvit. Seuran
Van. Julk. 1(3): 47 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 47).
SPECIMEN. TUR-V (holotype).

ASIA. —

Sporopodium leucoxanthum (Spreng.) Vain., Suom. Elain-
Ja Kasvit. Seuran Van. Julk. 1(3): 47 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 47).
SPECIMEN. TUR-V (holotype).

ASIA. —
Sticta (Schreb.) Ach.; Lobariaceae
Sticta boschiana Mont. & Bosch., in Junghuhn, Pl.

Junghuhn. 4: 436 (1855).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Wolseley (1991: 233).

SPECIMENS. BM.

ASIA. Indonesia, Philippines.

Sticta fuliginosa (Dicks.) Ach., Methodus: 280 (1803).

THAILAND. Chiang Mai (Doi Inthanon N.P., Doi Suthep N.P.).
Wolseley (1991: 233), Wolseley & Aguirre-Hudson (1997a: 334).

SPECIMENS. BM.

ASIA. China, India, Indonesia, Philippines.

Sticta nylanderiana Zahlbr., Cat. Lich. Univ. 3: 356
(1925).

CATALOGUE OF THE LICHENS OF THAILAND

THAILAND. Chiang Mai (Chiang Dao N.P., Doi Inthanon N.P., Doi
Suthep N.P., Doi Saget). Yoshimura (1978: 38), Wolseley (1991:

233), Wolseley & Aguirre-Hudson (1997a: 334).
SPECIMENS. AAU, BM, C.

ASIA. China, India, Taiwan.

Sticta platyphylloides Ny|., in Hue, Bull. Soc. Bot. France
34: 22 (1887).

THAILAND. Chiang Mai (Doi Suthep N.P.). Yoshimura (1978: 38).
SPECIMEN. AAU.

ASIA. China, India.

Sticta weigelii Isert, in Acharius, Lichenogr. Universalis:
446 (1810).

THAILAND. Nakhon Nayok (Khao Yai N.P.). Yoshimura (1978:
39).

SPECIMEN. AAU.

ASIA. China (incl. Hong Kong), India, Indonesia.

Sticta wrightii Tuck., Amer. J. Arts, ser. 2, 28: 204 (1859).
THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 45).
SPECIMENS. TUR-V.

ASIA. China, Taiwan.

Strigula Fr.; Strigulaceae
Strigula maculata (Cooke & Massee) R. Sant., Symb. Bot.
Upsal. 12(1): 186 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218).
SPECIMENS. CANB.

ASIA. India, Indonesia, Philippines, Sri Lanka, Vietnam.

Strigula melanobapha (Kremp.) R. Sant., Symb. Bot.
Upsal. 12(1): 188 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

SPECIMENS. CANB.

ASIA. China, India, Malaysia, Papua New Guinea, Philippines, Sri
Lanka.

Strigula smaragdula Fr., Linnaea 5: 550 (1830).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

SPECIMENS. CANB.

ASIA. China, India, Indonesia, Malaysia, Nepal, Papua New
Guinea, Philippines, Sri Lanka, Taiwan, Vietnam.

Strigula subtilissima (Fée) Miill. Arg., Flora 66: 346
(1863).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

53
SPECIMENS. CANB.
ASIA. China, Indonesia, Papua New Guinea, Philippines, Sri Lanka,
Taiwan, Vietnam.
Syncesia Taylor; Roccellaceae
Syncesia albiseda (Ny1.) Tehler, Fl. Neotropica 78: 37
(1996).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson
(1997a: 339).

SPECIMENS. BM.

ASIA. | Sri Lanka.

Teloschistes Norman; Teloschistaceae
Teloschistes flavicans (Sw.) Norman, Nytt Mag.
Naturvidensk. 7: 229 (1853).

Teloschistes flavicans f. glaber Vain.

THAILAND. Trat (Koh Chang I.). Vainio (1909: 111, as Teloschistes

flavicans f. glaber).
SPECIMENS. TUR-V.

ASIA. China, India, Indonesia, Sri Lanka.

Thelotrema Ach.; Thelotremataceae

Thelotrema alboannuliforme Nagarkar, Sethy & Patw.,
Kavaka 12(2): 57 (1985).

THAILAND. Loei (Na Haeo N.P.), Nakhon Ratchasima (Khao Yai

N.P.). Boonpragob et al. (1998: 218), Homchantara (1999: 64-66).
SPECIMENS. RAMK.

ASIA. | India.

Thelotrema albosorellum (Ny1.) Tuck., Gen. Lich.: 139
(1872).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218), Homchantara (1999: 66-67).
SPECIMENS. RAMK.

ASIA. India, Malaysia, Sri Lanka.

Thelotrema astroideum (Berk. & Broome) Hale,
Mycotaxon 11: 131 (1980).

THAILAND. Chantaburi (Namtok Phliu N.P.), Nakhon Ratchasima
(Khao Yai N.P.). Boonpragob et al. (1998: 218), Homchantara

(1999: 69-71).
SPECIMENS. RAMK.

ASIA. India, Malaysia, Sri Lanka.

Thelotrema calathiforme Vain., Hedwigia 46: 174 (1907).
THAILAND. Trat (Koh Chang I.). Vainio (1909: 119).
SPECIMENS. FH , TUR-V (both syntypes).

ASIA. — India.

54

Thelotrema canarense Patw. & C.R. Kulk., Norweg. J.
Bot. 24: 128 (1977).

THAILAND. Chiang Mai( Doi Suthep N.P.), Kanchanaburi (Erawan
N.P.), Sakhon Nakhon (Phu Phan N.P.). Homchantara (1999: 71—

73).
SPECIMENS. BM, RAMK.

ASIA. — India.

Thelotrema colobicum Nyl., Bull. Soc. Linn. Normandie,
sér. 2, 7: 169 (1873).

THAILAND. Chantaburi (Kitchakut N.P.), Chiang Mai (Doi Suthep
N.P.), Loei ( Na Haeo N.P., Phu Ruea N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Phetchabun (Phu Hin Rongkla N.P.), Sukhothai
(Ramkhamhaeng N.P.), Uthai Thani (Huay Kha Khaeng Wildlife
Sanctuary). Boonpragob et al. (1998: 218), Homchantara (1999:
73-76).

SPECIMENS. BM, RAMK.

ASIA. India, Sri Lanka.

Thelotrema dilatatum (Miill. Arg.) Hale, Mycotaxon 11: 13
(1980).

THAILAND. Chantaburi (Namtok Phliu N.P.). Homchantara (1999:

76-78).
SPECIMENS. RAMK.

ASIA. India, Malaysia, Sri Lanka.

Thelotrema expallescens Ny1., Flora 59: 560 (1876).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

SPECIMENS. RAMK.

ASIA. —

Thelotrema glaucopallens Ny1., Ann. Sci. Nat. Bot., sér. 4,
2: 327 (1863).

THAILAND. Chantaburi (Khao Soi Dao N.P., Kitchakut N.P.).

Homchantara (1999: 78-81).
SPECIMENS. RAMK.

ASIA. India, Malaysia.

Thelotrema imperfectum Hale, Bull. Brit. Mus. (Nat.
Hist.), Bot. 8(3): 255 (1981).

THAILAND. Uthai Thani (Huay Kha Khaeng Wildlife Sanctuary).
Homchantara (1999: 81-83).

SPECIMENS. BM.

ASIA. Sri Lanka.

Thelotrema laceratulum Miill. Arg., Flora 70: 399 (1887).

THAILAND. Lampang (Wiang Kosai N.P.), Nakhon Ratchasima
(Khao Yai N.P.), Uthai Thani (Huay Kha Khaeng Wildlife

Sanctuary). Homchantara (1999: 84-86).
SPECIMENS. BM, RAMK.

ASIA. India.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Thelotrema lacteum Kremp., in Nylander, Flora 47: 269
(1864).

THAILAND. Sakonnakhon (Phu Pham N.P.). Homchantara (1999:

86-89).
SPECIMENS. RAMK.

ASIA. India, Papua New Guinea, Sri Lanka.

Thelotrema laemense Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Kanchanaburi (Khao Laem N.P.). Homchantara (1999:
134-136, as Thelotrema ‘unidentified 3’).

SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Thelotrema lepademersum Nagarkar, Sethy & Patw.,
Kavaka 12(2): 59 (1985).

THAILAND. Chantaburi (Namtok Phliu N.P.), Nakhon Ratchasima

(Khao Yai N.P.). Homchantara (1999: 89-91).
SPECIMENS. RAMK.

ASIA. | India.

Thelotrema leprieurii (Mont.) Hale, Mycotaxon 11: 131
(1980).

THAILAND. Chantaburi (Namtok Phliu N.P.). Boonpragob et al.

(1998: 218), Homchantara (1999: 91-93).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Sri Lanka.

Thelotrema leprocarpoides Hale, Bull. Brit. Mus. (Nat.
Hist.), Bot. 8(3): 259 (1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218).
SPECIMENS. RAMK.

ASIA. India.

Thelotrema leprocarpum (Ny1.) Tuck., Gen. Lich.: 139
(1872).

THAILAND. Loei (Na Haeo N.P.), Sukhothai (Ramkhamhaeng

N.P.). Homchantara (1999: 94-96).
SPECIMENS. RAMK.

ASIA. India, Indonesia, Sri Lanka.

Thelotrema monosporum Ny\l., Ann. Sci. Nat. Bot., sér. 4,
15: 46 (1860).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Homchantara (1999: 96-100).

SPECIMENS. BM.

ASIA. India, Sri Lanka.

Thelotrema mongkolsukii Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Sukhothai (Ramkhamhaeng N.P.). Homchantara (1999:
129-131, as Thelotrema ‘unidentified 1’).

CATALOGUE OF THE LICHENS OF THAILAND
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Thelotrema nureliyum Hale, Bull. Brit. Mus. (Nat. Hist.),
Bot. 8(3): 259 (1981).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218), Homchantara (1999: 100-102).

SPECIMENS. RAMK.

ASIA. Sri Lanka.

Thelotrema patens Ny\., Acta. Soc. Sci. Fenn. 26: 17
(1900).

THAILAND. Chiang Mai (Doi Suthep), Phetchabaun (Phu Hin
Rongkla N.P.). Homchantara (1999: 102-104).

SPECIMENS. BM, RAMK.

ASIA. Malaysia, Sri Lanka.

Thelotrema phliuense Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Chantaburi (Namtok Phliu N.P.). Homchantara (1999:
138-140, as Thelotrema ‘unidentified 5’).

SPECIMENS. BM (holotype).

ASIA. —

Thelotrema phlyctidioides (Miill. Arg.) Hale, Mycotaxon
11: 132 (1980).

THAILAND. Chantaburi (Namtok Phliu N.P.), Chiang Mai ( Doi
Suthep N.P.), Nakhon Ratchasima (Khao Yai N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Homchantara (1999: 104-117).

SPECIMENS. BM, RAMK.

ASIA. Sri Lanka.

Thelotrema piluliferum Tuck., Proc. Amer. Acad. Arts 7:
227 (1868).

THAILAND. Chiang Mai (Doi Suthep N.P.). Homchantara: 1999:
107-108.

SPECIMENS. BM.

ASIA. Malaysia, Sri Lanka.

Thelotrema platysporum Harm., Bull. Séances Soc. Sci.
Nancy, sér. 3, 13: 41 (1912).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara

(1999: 110-112).
SPECIMENS. BM, RAMK.

ASIA. India, Malaysia, Papua New Guinea, Sri Lanka.

Thelotrema poeltii Patw. & C.R. Kulk., Norweg. J. Bot. 24:
130 (1977).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218).
SPECIMENS. RAMK.

ASIA. — India.

55

Thelotrema porinoides Mont. & Bosch, in Junghuhn,
Enum. Pl. Insul. Java Sumatra: 151 (1855).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218), Homchantara (1999: 114-117).
SPECIMENS. RAMK.

ASIA.
pines.

India, Indonesia, Malaysia, Papua New Guinea, Philip-

Thelotrema pseudoexanthismocarpum (Patw. & C.R.
Kulk.) Hale, Mycotaxon 11: 132 (1980).

THAILAND. Chiang Mai (Doi Suthep N.P.), Satun (Thalae Ban
N.P.), Sukhothai (Ramkhamhaeng N.P.), Uthai Thani (Huay Kha

Khaeng Wildlife Sanctuary). Homchantara (1999: 117-120).
SPECIMENS. RAMK.

ASIA. — India.

Thelotrema pycnophragmium Nyl., Sert. Lich. Trop.
Labuan Singapore: 5 (1891).

THAILAND. Chantaburi(Namtok Phliu N.P.). Homchantara (1999:
120-122).

SPECIMENS. RAMK.

ASIA. India, Malaysia.

Thelotrema rhododiscum Homchantara & Coppins,
Lichenologist 34: (in press).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Homchantara

(1999: 136-138, as Thelotrema ‘unidentified 4’).
SPECIMENS. E (isotype), RAMK (holotype).

ASIA. —

Thelotrema rugatulum Ny1., Bull. Soc. Linn. Normandie,
sér. 2, 7: 168 (1873).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218).
SPECIMENS. RAMK.

ASIA. India

Thelotrema subhiatum Patw. & Nagarkar, Biovigyanam 6:
9 (1980).

THAILAND. Chantaburi (Kitchakut N.P.), Phetchabun (Phu Hin
Rongkla N.P.), Sakonnakhon (Phu Phan N.P.). Homchantara (1999:
120-122).

SPECIMENS.

India.

RAMK.

ASIA.

Thelotrema turgidulum (Miill. Arg.) Hale, Mycotaxon 11:
132 (1980).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et

al. (1998: 218), Homchantara (1999: 125-129).
SPECIMENS. RAMK.

ASIA. India.

56
Thelotrema weberi Hale, Phytologia. 27: 497 (1974).

THAILAND. Chiang Mai ( Doi Suthep N.P.). Homchantara (1999:
107-108).

SPECIMENS. BM.

ASIA. Malaysia, Papua New Guinea, Sri Lanka.

Tricharia Fée emend. R. Sant.; Gomphillaceae.

Tricharia aff. vainioi R. Sant., Symb. Bot. Upsal. 12(1):
382 (1952).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

SPECIMEN. RAMK.

ASIA. China, Indonesia, Papua New Guinea, Philippines.

Trypethelium Spreng.; Trypetheliaceae

Trypethelium eluteriae Spreng., Anleit. Kenntn. Gew. 3:
351 (1804).

THAILAND. Chumphon (Koh Tao I.), Nakhon Ratchasima (Khao
Yai N.P.). Paulson (1930: 100), Boonpragob et al. (1998: 218).

SPECIMENS. BM, CANB, RAMK.

ASIA. China (incl. Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Taiwan, Vietnam.

Trypethelium nigroporum Makhija & Patw., J. Hattori
Bot. Lab. 73: 202 (1993).

THAILAND. Loei (Naheaw), Sakonnakhon (Phu Pan N.P.).
Vongshewarat et al. (1999: 235).

SPECIMENS. RAMK.

ASIA. — India.

Trypethelium tropicum (Ach.) Miill. Arg., Bot. Jahrb. Syst.
6: 393 (1885).

THAILAND. Nakhon Ratchasima (Khao Yai N.P.). Boonpragob et
al. (1998: 218).

SPECIMENS. CANB, RAMK.

ASIA. China (Hong Kong), India, Indonesia, Malaysia, Papua
New Guinea, Singapore, Vietnam.

Trypethelium variolosum (Ach.) Miill. Arg., Flora 68: 255
(1885).

Pseudopyrenula ochroleuca (Eschw.) Vain.
Pseudopyrenula ochroleuca var. subdissocians Vain.
Trypethelium ochroleucum var. subdissocians (Vain.) Zahlbr.

THAILAND. Chiang Mai (Doi Suthep N.P.), Trat (Koh Chang I.).
Vainio (1909: 147; 1921: 55, as Pseudopyrenula ochracea), Harris
(1998: 145).

SPECIMEN. TUR-V (holotype of Pseudopyrenula ochroleuca vat.
subdissocians).

ASIA. India, Indonesia, Malaysia, Papua New Guinea.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

Tylophoron Nyl. ex Stizenb.; Caliciales

Tylophoron moderatum Ny|., Bot. Zeitung 20: 279 (1862).

THAILAND. Chiang Mai (Doi Suthep N.P.), Uthai Thani (Huay
Kha Khaeng Wildlife Sanctuary). Wolseley & Aguirre-Hudson
(1997a: 334, 339).

SPECIMENS. BM.

ASIA. China, India, Taiwan.

Usnea Dill. ex Adans.; Usneaceae

Usnea australis Fr., Syst. Orb. Veg.: 282 (1825).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.).
Vainio (1921: 33), Sato (1962: 4), Santisuk (1988: 99),

SPECIMEN. TUR-V.

ASIA. —

Usnea baileyi (Stirt.) Zahlbr., Denkschr. Kaiserl. Akad.
Wiss., Wien Math.-Naturwiss. KI. 83: 182, 191 (1909).

Eumitria perrubescens Vain.
Usnea implicita (Stirt.) Zahlbr.
Usnea implicita subsp. perrubescens Stitt.

THAILAND. Loei (Phu Luang, Chiang Mai (Doi Suthep N.P.).
Vainio (1921: 35, as Eumitria perrubescens), Motyka (1936: 62, as
Usnea implicita subsp. perrubescens), Rogers & Stevens (1988:
358), Yoshimura (1978: 35).

AAU, C, TUR-V (holotype of Eumitria perrubescens).

ASIA. China (incl. Hong Kong), Indonesia, Malaysia, Myanmar,
Papua New Guinea, Philippines, Taiwan.

SPECIMENS.

Usnea baileyi subsp. chrysopora (Stein) Asahina, J. Jap.
Bot. 42: 7 (1967).

THAILAND. Chiang Mai (Doi Suthep N.P., Chiang Dao N.P.).
Yoshimura (1978: 36).

SPECIMENS. C.

ASIA. Philippines.

Usnea baileyi subsp. septentrionalis Asahina, in Hara, FI.
Eastern Himalaya: 598 (1966).

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Thung Salaeng,

Phu Luang N.P.). Yoshimura (1978: 35).
SPECIMENS. AAU, C.

ASIA.

Usnea dasycera (Ny1.) Motyka, Usnea: 552 (1936-38).
Usnea perplexans f. dasycera (Nyl.) Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 33, as
Usnea perplexans f. dasycera).

AAU, C.

India, Nepal, Taiwan.

SPECIMENS.

ASIA. —

Usnea endochroa var. papillata (Vain.) Zahlbr., Cat. Lich.
Univ. 6: 564 (1930).

Eumitria endochroa var. papillata Vain.

CATALOGUE OF THE LICHENS OF THAILAND

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 34, as
Eumitria endochroa var. papillata).

SPECIMEN. TUR-V.

ASIA. Philippines.

Usnea endorhodina (Vain.) Zahlbr., Cat. Lich. Univ. 6: 564
(1930).

Eumitria endorhodina Vain.

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 34, as
Eumitria endorhodina).

SPECIMEN. TUR-V.

ASIA. Philippines.

Usnea florida var. subcomosa Vain., Philipp. J. Sci. Sect.
C, 4: 652 (1909).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 33).
SPECIMEN. TUR-V.

ASIA. China, Philippines.

Usnea himantodes Stirt., Scott. Naturalist (Perth) 7: 75
(1883).

Usnea neoguineensis var. gracilior Asahina.
Usnea neoguineensis var. neoguineensis Asahina.

THAILAND. Loei (Phu Luang). Asahina (1968: 497, as Usnea
neoguineensis vat. gracilior), Yoshimura (1978: 36, as U. neoguine-
ensis and U. neoguineensis var. gracilior), Stevens (1990: 410).

SPECIMENS. AAU, TNS (holotypes of Usnea neoguineensis var.
gracilior and Usnea neoguineensis var. neoguineensis).

ASIA. China, Indonesia, Papua New Guinea, Philippines, Taiwan.
Usnea hossei Vain., Suom. Elain-ja Kasvit. Seuran Van.
Julk. 1(3): 34 (1921).

THAILAND. Chiang Mai (Doi Suthep N.P., Doi Inthanon N.P.).
Vainio (1921: 34), Zahlbruckner (1933: 581), Sato (1938: 4; 1962:

4), Herre (1963: 65), Santisuk (1988: 99).
SPECIMENS. BKF, TNS, TUR-V (syntypes).

ASIA. China, Malaysia, Taiwan.

Usnea leucospilodea Nyl., J. Linn. Soc. London (Bot.) 20:
30 (1883).

THAILAND. Chiang Mai (Doi Suthep N.P.). Asahina, (1969: 37),
Yoshimura (1978: 36).

SPECIMEN. AAU, C.

ASIA. India, Papua New Guinea, Laos, Malaysia, Taiwan.

Usnea luridorufa Stirt., Scott. Naturalist (Perth) 7: 104
(1881).

THAILAND. Doi Suthep. Motyka (1937-38: 523), Awasthi (1986:
374).

SPECIMEN. AAU, GC:
ASIA. China.

a7

Usnea pectinata Taylor, in Hooker, J. Bot. (London) 6: 191
(1847).

THAILAND. Chiang Mai (Doi Suthep N.P.), Loei (Phu Kradung
N.P., Phu Luang N.P.). Awasthi (1986: 362), Yoshimura (1978: 36),

Kurokawa & Kashiwadani (1988: 51).
SPECIMENS. Kurokawa, Lich. Rar. Crit. Exs. No. 347, C, AAU.

ASIA. China, India, Malaysia.

Usnea perplexans Stirt. Scott. Naturalist (Perth) 6: 103
(1881).

THAILAND. Chiang Mai (Doi Suthep N.P.). Vainio (1921: 33).
SPECIMEN. AAU.

ASIA. India, Philippines.

Usnea rubicunda Stirt., Scott. Naturaliust (Perth) 6: 102
(1881).

THAILAND. Chiang Mai (Doi Suthep N.P), Phu Miang. Yoshimura
(1978: 36).
SPECIMENS. AAU.

ASIA. China, India, Indonesia, Taiwan, Vietnam.

Xanthoparmelia (Vain.) Hale; Parmeliaceae

Xanthoparmelia congensis (Stein) Hale, Phytologia 28:
486 (1974).

THAILAND. Chiang Mai (Doi Saget), Loei (Ban Sui, Phu Kradung),
Lumpang (Khuntan N.P.), Phitsanulok (Lan Hin Taek, Phu Hin
Rongkla, N.P.), Sakonnakhon (Phu Pan N.P.). Pooprang et al. (1999:
125), Moon et al. (2000a: 105).

SPECIMENS. RAMK, TNS.

ASIA. China (incl. Hong Kong).

Xanthoparmelia mougeotina (Ny1.) D.J. Galloway, New
Zealand J. Bot. 18: 538 (1981).

THAILAND. Lumpang (Khuntan N.P.). Pooprang et al. (1999: 125).
SPECIMEN. RAMK.

ASIA. Papua New Guinea.

ACKNOWLEDGEMENTS. We are grateful to those organisations and
individuals who have assisted this project over many years: the Leverhulme
Trust who provided the initial funding to Pat Wolseley and Begofia Aguirre-
Hudson for this research in Thailand; The Natural History Museum for
supporting this research during and since initial funding; Prof. Teuvo Ahti
(Helsinki), Prof. Jack Elix (Canberra), Dr Seppo Huhtinen (Turku), Dr
Hiroyuki Kashiwadani (Tokyo), Laszlo L6kés (Budapest), Dr Simone
Louwhoff (London), the Royal Forestry Department, Thailand; Prof. Kansri
Boonpragob, Dr Natsurang Homchantara and the staff and students of the
Lichen Research Group, Ramkhamhaeng University, Bangkok; Dr Wanaruk
Saipunkaew (Chiang Mai University), Dr Oy Kanjanavanit and M.R.
Smansnid Svasti.

This catalogue is dedicated to Thai lichenologists and especially to Prof.
Kansri Boonpragob and Dr Natsurang Homchantara under whose direction
lichenology now flourishes in Thailand.

58

REFERENCES

Abbayes, H. des 1956. Quelques Cladonia (lichens) des regions inter-tropicales,
nouveaux ou peu connus, conservés dans I’ herbier de Kew. Kew Bulletin 2: 259-266.

1964 [‘1963"]. Lichens nouveaux ou interessants du Vietnam. Revue Bryologique
et Lichénologique 32: 216-222.

Aguirre-Hudson, B. & Wolseley, P.A. 1994. Bibliography of southeast Asian
lichenology, 1810-1992. Journal of the Hattori Botanical Laboratory 76: 313-350.

Ahti, T. 1984. The status of Cladina as a segregated from Cladonia. Beitréige zur Nova
Hedwigia 79: 25-61.

Archer, A.W. 1997. The lichen genus Pertusaria in Australia. Bibliotheca Lichenologica
69: 1-249.

Arvidsson, L. 1982. A monograph of the lichen genus Coccocarpia. Opera Botanica
67: 1-99.

Asahina, Y. 1968. Lichenologische Notizen (211-212). Journal of Japanese Botany
43: 495-499.

—— 1969. Lichenologische Notizen (215-216). Journal of Japanese Botany 44: 33-
39,

Awasthi, D.D. 1960. On a collection of macrolichens by the Indian expedition to Cho-
oyu, east Nepal. Proceedings of the Indian Academy of Sciences 51: 169-180.

1975. A monograph of the lichen genus Dirinaria. Bibliotheca Lichenologica 2:
1-108.

— 1977. [‘1976’] Lichen genus Parmelia in India I — subgenera Parmelia and
Amphigymnia. Biological Memoirs, Lich. I 1(1-2): 155-229.

—— 1986. Lichen genus Usnea in India. Journal of the Hattori Botanical Laboratory
61: 333-421.

Boonpragob, K., Homchantara, N., Coppins, B.J., McCarthy, P.M. & Wolseley,
P.A. 1998. An introduction to the lichen flora of Khao Yai National Park, Thailand.
Botanical Journal of Scotland 50(2): 209-219.

Culberson, C.F., Culberson, W.L. & Johnson, A. 1981. A standarized TLC analysis
of B-orcinol depsidones. The Bryologist 84: 16-29.

& Hale, M.E. 1973. Chemical and morphological evolution in Parmelia sect.
Hypotrachyna: product of ancient hybridization? Brittonia 25: 162-173.

David, F., Elix, J.A. & Samsudin, M.W. 1990. Two new aliphatic acids from the lichen
Parmotrema praesorediosum. Australian Journal of Chemistry 43: 1297-1300.
Degelius, G. 1974. The lichen genus Collema with special reference to the extra-

European species. Symbolae Botanicae Upsalienses 20(2): 1-215.

Elix, J.-A. 1996. A revision of the lichen genus Relicina. Bibliotheca Lichenologica
62:1-150.

Hafellner, J. 1981. Monographie der Flechtengattung Letrouitia (Lecanorales,
Teloschistinae). Nova Hedwigia 35: 645-729.

1993. Die Gattung Pyrrhospora in Europa. Eine erste Ubersicht mit einem
Bestimmungsschliissel der Arten nebst Bemerkungen zu einigen aussereuropiischen
Taxa (lichenisierte Ascomycotina, Lecanorales). Herzogia 9: 725-747.

Hale, M.E. 1965a. A monograph of Parmelia subgenus Amphigymnia. Contributions
of the United States National Herbarium 36(5): 193-358.

1965b. Six new species of Parmelia from southeast Asia. Journal of Japanese
Botany 40: 199-205.

—— 1968. New Parmeliaceae from southeast Asia. Journal of Japanese Botany 43:
324-328.

1971. Morden-Smithsonian expedition to Dominica: the lichens (Parmeliaceae).

Smithsonian Contributions to Botany 4: 1-25

1975. A monograph of the lichen genus Relicina (Parmeliaceae). Smithsonian

Contributions to Botany 26: 1-32.

1976a. A monograph of the lichen genus Bulbothrix Hale (Parmeliaceae).
Smithsonian Contributions to Botany 32: 1-29.

—— 1976b. A monograph of the lichen genus Pseudoparmelia Lynge (Parmeliaceae).
Smithsonian Contributions to Botany 31: 1-62.

1976c. A monograph of the lichen genus Parmelina Hale (Parmeliaceae).
Smithsonian Contributions to Botany 33: 1-60.

—— 1981. A revision of the lichen family Thelotremataceae in Sri Lanka. Bulletin of
the British Museum (Natural History), Botany 8: 227-332.

& Kurokawa, S. 1964. Studies on Parmelia subgenus Parmelia. Contributions
from the U.S. National Herbarium 36(4): 121-191.

Hamalainen M. & Pinratana A. 2001. Atlas of the dragonflies of Thailand. Bangkok.

Harris, R.C. 1993 [*1991"]. A revision of Polymeridium (Miill. Arg.) R.C. Harris
(Trypetheliaceae). Boletim do Museu Paraense Emilio Goeldi, Série Botanica 7:
619-644.

—— 1998. A preliminary revision of Pseudopyrenula Mill. Arg. (lichenized Asco-
mycetes, Trypetheliaceae) with a redisposition of the names previously assigned to
the genus. /n M.G. Glenn, R.C. Harris, R. Dirig & M.S. Cole (Eds) Lichenographia
Thomsoniana: North American lichenology in honour of John W. Thomson, pp. 133-
148. New York.

Hawksworth, D.L. 1985. A redisposition of the species referred to the ascomycete genus
Microthelia. Bulletin of the British Museum (Natural History), Botany 14: 43-181
—— Kirk, P.M., Sutton, B.C. & Pegler, D.N. 1995. Ainsworth & Bisby’s Dictionary

of the Fungi. 8th ed. Wallingford.

P.A. WOLSELEY, B. AGUIRRE-HUDSON AND P.M. McCARTHY

—— & Yoshimura, I. 1973. 358. Proposal to conserve the generic name Gymnoderma
Nyl. 1863. [Lichenes] against Gymnoderma Humb ex Steud. 1824. [Fungi]. Taxon
22: 503.

Henssen, A. 1963. Eine revision der Flechtenfamilien Lichinaceae und Ephebaceae.
Symbolae Botanicae Upsalienses 18: 1-123.

1979. Problematik der Gattungsbegrenzung bei den Lichinaceen. Berichte der

Deutschen Botanischen Gesellschaft 92: 483-506.

1989. Metamelanea and Stromatella, new genera of Lichinaceae. Lichenologist
21: 101-118.

Herre, A.W.T.C. 1963. The lichen genus Usnea and its species at present known from
the Philippines. Philippine Journal of Science 92(1): 41-76.

Holmgren, P.K., Holmgren, N.H. & Barnett, L.C. 1990. Index Herbariorum Part 1:
The Herbaria of the World. New York.

Homchantara, N. 1999. The taxonomic and ecological aspects of the Thelotremataceae
in southeast Asia. Unpublished Ph.D. thesis. John Moores University Liverpool.
& Coppins, B.J. (2002). New species of the lichen family Thelotremataceae in SE

Asia. Lichenologist 34: (in press).

Huovinen, K. & Ahti, T. 1986. The composition and contents of aromatic lichen
substances in Cladonia, section Unciales. Annales Botanici Fennici 23: 173-188.

Jorgensen, P.M. 1997. Further notes on hairy Leptogium species. Symbolae Botanicae
Upsalienses 32(1): 113-130.

2000. Notes on some east-Asian species of the lichen genus Fuscopannaria.

Journal of the Hattori Botanical Laboratory 89: 247-259.

2001. New species and records of the lichen family Pannariaceae from Australia.
Bibliotheca Lichenologica 78: 109-139

Kalb, K. & Elix, J.A. 1995. The lichen genus Physcidia. Bibliotheca Lichenologica
57: 265-296.

Kashiwadani, H. 1975. Ennumeration of Anaptychiae and Parmeliae of Papua New
Guinea. Reports of the Cryptogams in Papua New Guinea: 75-85. Tokyo.

1977. Rare lichens collected in the Izu Islands. Memoirs of the National Science

Museum, Tokyo, Botany 10: 43-48.

1988. Ramalina hossei Vainio (lichen) found in Buthan and Formosa. Bulletin of

the National Science Museum, Tokyo, Botany 14: 129-133.

& Gradstein,S.R. 1982. Notes on Baeomyces sanguineus Asahinaand Gymnoderma
coccocarpum Nyl. Miscellanea Bryologica et Lichenologica 9: 79-81.

Kurokawa, S. 1959. Anaptychiae (lichens) and their allies in Japan (2). Journal of
Japanese Botany 34: 174-184.

1961. Anaptychiae (lichens) and their allies of Japan (6). Journal of Japanese

Botany 36: 51-56.

1962. A monograph of the genus Anaptychia. Beihefte zur Nova Hedwigia 6: 1—

155.

1973. Supplementary notes on the genus Anaptychia. Journal of the Hattori

Botanical Laboratory 37: 563-607.

1980. Cetrariopsis, a new genus in the Parmeliaceae, and its distribution.

Memoirs of the National Science Museum, Tokyo 13: 139-142.

1988. Materials for the distribution of lichens in Japan (10). Journal of Japanese

Botany 63: 167-168.

1991. Rimeliella, a new lichen genus related to Rimelia of the Parmeliaceae.

Annals of the Tsukuba Botanical Garden 10: 1-14.

1998. A new record and a new species in Myelochroa (Parmeliaceae). Journal of

Japanese Botany 73: 113.

& Kashiwadani, H. 1988. Indices to taxa distributed under Lichens rariores et
critici exsiccati Fascicules I-XIV (Nos. 1-700). Tokyo.

Lumbsch, H.T., Guderley, R. & Elix, J.A. 1996. A revision of some species in
Lecanora sensu stricto with a dark hypothecium (Lecanorales, Ascomycotina). The
Bryologist 99: 269-291.

McCarthy, P.M. 1995. A reappraisal of Clathroporina Miill. Arg. (Trichotheliaceae).
Lichenologist 27: 321-350.

— 1999. Three new species of Porina (Trichotheliaceae) from Thailand. Lichenologist
31: 239- 246.

Moon, K.H., Kurokawa, S. & Kashiwadani, H. 2000a. A list of Thailand species of
Parmelia (sens. lat.) Preserved in the National Science Museum. Jn K. Matsura (Ed.)
Proceedings of the First and Second Symposia on Collection Building and Natural
History Studies in Asia. National Science Museum Monographs 18. Tokyo: 97-106.

——. & 2000b. A new species of Hypotrachyna (lichenised Ascomycetes,
Parmeliaceae) from Thailand. Bulletin of the National Science Museum, Tokyo, ser.
B, 26: 135-138.

Mongkonsuk, P., Boonpragob, K., Homchantara, N. & Boonchai, W. 1996. Diver-
sity of the lichens at Queen Sirikit Botanic Garden, Mae Rim, Chiang Mai. Botanical
Garden Organization Newsletter 4: 13-22.

Motyka, J. 1936. Lichenum generis Usnea studium monographicum, Pars systematica
1: 1-304. Lublin.

— 1937-1938. Lichenes generis Usnea studium mongraphicum, Pars systematica 2:
305-651. Lublin.

Oshio, M. 1976. Pertusaria variolosa (Kremp.) Vain. and allied species. Miscellanea
Bryologica et Lichenologica 7(6): 114-116.

Parmasto, E. 1978. The genus Dyctionema (Thelophorolichenes). Nova Hedwigia 29:
99-144.

CATALOGUE OF THE LICHENS OF THAILAND

Patwardhan, P.G. & Kulkarni, C.R. 1976. Some additions to the lichen flora of India
IV: Graphis and Graphina (Family Graphidaceae). Biovigyanam 2: 123-132.

— 1977a. A contribution to our knowledge of the lichen flora of India I: Family
Thelotremataceae. Kavaka 5: 1-17.

—— 1977b. Some additions to the lichen flora of India V: Genera Phaeographis and
Phaeographina (Family Graphidaceae). Current Science 46: 720-721.

— & Prabhu, A.V. 1977. Some additions to the lichen flora of India II: Parmelina
Hale and Parmotrema Mass. Biovigyanam 3: 111-113.

Paulson, R. 1930. Lichens from Kaw Tao, an island in the Gulf of Siam. The Journal
of the Siam Society, Natural History Supplement 8(2): 99-101.

Pooprang, T., Boonpragob, K. & Elix, J.A. 1999. New species and new records in the
lichen family Parmeliaceae (Ascomycotina) from Thailand. Mycotaxon 71: 111-
127.

Rogers, R.W. 1986. The genus Pyxine (Physciaceae, lichenized Ascomycetes) in
Australia. Australian Journal of Botany 34: 121-154.

—— & Stevens, G.N. 1988. Usnea baileyi complex (Parmeliaceae, Lichenised Asco-
mycetes) in Australia. Australian Systematic Botany 1: 355-361.

Rostrup, F.G.E. 1902. Fungi. /n J. Schmidt (Ed.), Flora of Koh Chang; Contributions
to the knowledge of the vegetation in the Gulf of Siam. Botanisk Tidsskrift Kjobenhavn
24: 355-367.

Salisbury, G. 1971. The Thelotremata of Angola and Mocambique. Revista de Biologia
7: 271-280.

Santesson, R. 1952. Foliicolous lichens I: A revision of the taxonomy of the obligately
foliicolous lichenized fungi. Symbolae Botanicae Upsaliensis 12(1): 1-590.

Santisuk, T. 1988. An account of the vegetation of northern Thailand. Geoecological
Research 5: 1-101.

Sato, M. 1938. Enumeratio Lichenum Ins. Formosae (IV). Journal of Japanese Botany
14: 463-469.

— 1962. Lichenes of Doi Inthanon and Doi Suthep northwestern Thailand. /n T. Kira
& T. Umeaso (Eds), Nature and life in Southeast Asia Wl: \-4.

Sérusiaux, E. 1985. Goniocysts, goniocystangia and Opegrapha lambinonii and
related species. Lichenologist 17: 1-25.

Sethy, P.K., Nagarkar, M.B. & Patwardhan, P.G. 1987. Materials for a lichen flora
of the Andaman Islands — III. Mycotaxon 28: 191-198.

Singh, K.P. 1979. Some new records of lichens from India. Bulletin of the Botanical
Survey of India 21: 221-223.

Sipman, H.J.M. 1983. A monograph of the lichen family Megalosporaceae. Bibliotheca
lichenologica 18: 1-241.

Staiger, B. & Kalb, K. 1995. Haematomma-studien. I. Die Flechtengattung
Haematomma. - Bibliotheca Lichenologica, 59, Berlin.

Stenroos, S. 1986. The family Cladoniaceae in Melanesia |. Cladonia sect. Uncialis.
Annales Botanici Fennici 23: 161-164.

— 1988. The family Cladoniaceae in Melanesia 4. The genera Cladia, Cladina,
Calathaspis and Thysanothecium. Annales Botanici Fennici 25: 207-218.

59

Stevens, G.N. 1990. Usnea himantodes Stirton and its synonyms. Lichenologist 22:
409-412.

Thor, G. 1990. The lichen genus Chiodecton and five allied genera. Opera Botanica
103: 1-92.

Vainio, E.A. 1907. Lichenes novi rarioresque, 4. Hedwigia 46: 168-181.

— 1909. Lichenes. /n J. Schmidt (Ed.), Flora of Koh Chang; Contributions to the
knowledge of the vegetation in the Gulf of Siam. Botanisk Tidsskrift Kjobenhavn 29:
104-151.

—— 1921. Lichenes in summo monte Doi Sutep (circ. 1675 m.s.m.) in Siam boreali
anno 1904 AD: re C.C. Hosseus collect. Suomalise Elain-ja Kasvitieteelisen Seuran
Vanamon Julkaisuja 1(3): 33-55.

Vongshewarat, K., McCarthy, P. M., Mongkolsuk, P. & Boonpragob, K. 1999.
Additions to the lichen flora of Thailand. Mycotaxon 70: 227-236.

Wirth, M. & Hale, M.E. 1978. Morden-Smithsonian expedition to Dominica: The
lichens (Graphidaceae). Smithsonian Contributions to Botany 40: |-64.

Wolseley, P. 1991. Observations on the composition and distribution of the ‘Lobarion’
in forests of south east Asia. In D.J. Galloway (Ed.), Tropical lichens: their
systematics, conservation, and ecology. Systematics Association Special Volume 43:
217-243. Oxford.

— & Aguirre-Hudson, B. 1991. Lichens as indicators of environmental change in
the tropical forests of Thailand. Global Ecology and Biogeography Letters 1: 170-
ikgfeys

& 1995. Key to lichen genera in Thailand with special reference to epiphytic

taxa, Part 1: Macrolichens. Natural History Bulletin of the Siam Society 43: 303-

335;

& — 1997a. The ecology and distribution of lichens in tropical deciduous and

evergreen forests in Thailand. Journal of Biogeography 24: 327-343.

& 1997b. Fire in tropical dry forests: corticolous lichens as indicators of

recent ecological changes in Thailand. Journal of Biogeography 24: 345-362.

Moncrieff, C. & Aguirre-Hudson, B. 1994. Lichens as indicators of environ-
mental stability and change in the tropical forests of northern Thailand. Global
Ecology and Biogeography Letters 4: 116-123.

Yoshimura, I. 1971. The genus Lobaria of eastern Asia. Journal of the Hattori
Botanical Laboratory 34: 231-364.

1978. Some lichens of Thailand collected by the Danish botanists, 1958-68.

Bulletin of the Kochi Gakuen Junior College 9: 35-40.

1982. Distribution of Gymnoderma coccocarpum Ny. and G. insulare Yoshim. &

Sharp. Bulletin of the Kochi Gakuen Junior College 13: 83-86.

& Sharp, A.J. 1968. A revision of the genus Gymnoderma. American Journal of
Botany 55: 635-640.

Zahlbruckner, A. 1926. Lichenes (Flechten). Jn A. Engler & K. Prantl (Eds), Die
Naturlichen Pflanzenfamilien 8: 1-270. Leipzig.

1933. Flechten der Insel Formosa II. Feddes Repertorium Specierum Novarum

Regni Vegetabilis 33: 22-68.

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