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SMITHSONIAN INSTITUTION
UNITED STATES NATIONAL MUSEUM
Butwetin 181
THE CYCLOPHORID OPERCULATE
LAND MOLLUSKS OF AMERICA
By
CARLOS DE LA TORRE
PAUL BARTSCH
and
JOSEPH P. E. MORRISON
UNITED STATES
GOVERNMENT PRINTING OFFICE
WASHINGTON : 1942
For sale by the Superintendent of Documents, Washington D.1Can a an nei aee tea an Price 65 cents
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ALEXANDER WETMORE,
Assistant Secretary, Smithsonian Institution.
Wasxineron, D. C., June 9, 1942.
II
CONTENTS
Page
Imtroduction and acknowledgments] 2——— =o - oes eee eee oe 1
The cyclophorid mollusks of Cuba, by Cartos pg La Torre and PavuL
RS Bae ee oe eg re ee a ee BE 3
Paraily, Cyclophoridac# 24.22 o_o eae ee SE Se -- =. 3
Subfamily Megalomastominae=_.. =. -2.02teee eee soe) = - 3
GoanustMarcunens 2 soos fe Se SESE 3 2 4
Subgenus. Farcimens:_.- 25.2 Se eo = 5
Subfamily»Aperostominge +22 ts 3) jee a eee eee eee oe SSS 38
Genus Grocidopamane. - a ee ued a Ee 39
Subgenus: Cyclocubana_._.. Sunset. SURE == 39
The eyclophorid mollusks of the West Indies, exclusive of Cuba, by Paun
ARDS CHE aha te ee oe oP Ce ae a RNS ee 43
Family yclophoridaes s.-e- tee tas eee eee 43
Subfamily Megalomastominac louse. ule ye oat ee 43
Genus Harcimen=- 2s = 6 ee ee a ee 43
Subgenus Weopupinias ey aes sae ee 43
Gems Parcrmnoides sake ease BUN oc ee 46
Genus Megalomastoma -2- Oo se Sie BUA 2 ere 48
Subgenus Megalomastomal.b.2 8 22 Sseh ss BLO se 48
Subgenus Megalomastomoides_--_--------------------- 50
Subfamily, Diplommatininae—... 229s 9204s lunes, Ses ee 51
Genus Adelopoma === <== 2 Seek eeen se ee 51
Subfamily. Amphicyclotinaes-._+-2-- =e<= =e oe te 52
(Cremmiss Ci clonatiia ss! hens tte PEN SE ee Soe 2a 52
GenuseAmpnicyeroliiites= = 22 aes Sree ee See eee 54
Subgentis: Ampnzcyclowuiuss= 9 = 99 8 be Be 54
SubsennsiCycloblandia-—- \! = we es 2 oe kt ee Oe 60
Sublamily: Aperostominae.- +95. oo eae fe eee ee oe 62
Genus Crovidnpomarse wee oe aan eee ee 62
SubgentisnGrecidopemaAle les. 22h Le 62
Genirsi@yelogagnapccae < soos hoe te 67
Gents ‘Cyclonendreysia. io. = 12 ee te ee 69
Genie Cyclon isGryikes soe Uae Bee ee a ae oe 71
Subgenus Cyclopusbryag.2- 2 ee eo ee Ye 71
Subgenus Cyclocaymanita__<. 9.223 2s 2 2 2 23 =e 79
GenusiPivenocochiies so ee UNI Si eee 82
Genus: Poleria ing) S222 2 Pe pe ies Se eee ee 105
Subgenusweorert@s 62. 8 Sa ee 106
Subgenus) Cyclobakerza-.— 224 es cee ee eee 115
Genus Aperostemde) 2 en te ees eo eee eee 124
Subgenus! Cycladameraees 2316 / 3022s Se Se ee eb 125
Subgenus Ausirocyclotustc ee = oes eee Sse est 132
Subgenvis Cyelohidalgoa 22. ol | Se ee ee He 136
Pseudogeneric term Incerticyclus___---------------------- 137
IV CONTENTS
Page
The cyclophorid mollusks of the mainland of America, by Pau, Bartscu
and Josprn P. i. MOonnIsON= 22-2220 22 22 ee 142
Famibys@velophoridae--2 22 — ies 2. ee ee ee ee 142
Subfamily Megalomastominae:—- 2.4 =22-22-= == £2 aaa 142
Genus: Lomocyclus ss... - 2S Se eae ees eee ee eee 142
Subfamily: Diplommatininaes:2 2-052") 25-2 Fe ee 148
Genus! Adelonoma 2222 22 one ee a ee eee 148
Subfamily Amphicyclotinaes= 2" 52-5252 2- ae ee 151
Genus Buckleyta_-—. 223. = == PE eee eae ee ee 151
Genus Lagoeyclus- x21 2 eds eh 8 eS ieee 154
Genus Filocyclus:-2. 2.) 5.682 Se oe ae ESS eee eee 157
Genus Calaperostoma:.- =~. 252. — (see ey 159
Genus: Cyrtotoma__.__--- - Sassaeisn nop an Es eee 169
Genus’ Barbacicluss 2220225 <5 3. Ye ee a eee te 175
Genus Calacyclotuss2225. 22. Beni Oe hee eee 178
Genus, Mexcyclotuss.2 2-2 + == Samo he eee ee 179
Genus Megacyclotus. <5 32 Jo 3 eee EE Bae ah eee 181
Genus Amphicyclotus._.. < . 2s be he eee = - 2 183
Subfamily, ;Aperostominae 22-420 26h. ase dees Wee dE abe 186
Genus Apenostoma. -2- 22 ='s- 222512 Se ee 187
Subgenus Jncidostomaz -. - 4.) 3. jt te eke 187
Subgenus: Austrocyclotusco2ae 352 eee ee eae eee 195
Subgenus Neocyclotus.2=-.-22-- 226 MS gy og Be ee 203
Subgenus Cyclopomops.u: 2244-0 sue see See ee 219
Subgenus Aperostoma. 2. - 2h sce ee tesa ee 221
Subgenus’ Cyclohidalgoa___ 2. 4-0 She = ey 268
Genus Dnracyclotués. 2 ooe0 ae el ae oe Ss 274
Pseudogeneric term Incerticyelus.__-/. ~. 22 ==-22--=---==-=- 275
A list. of the mainland cyclophorids.._+ 52-222) 4.ce"'. ate 279
Hixplanation of plates. .2o522. 324222 2 oso 2 So ee ee 283
THE CYCLOPHORID OPERCULATE LAND
MOLLUSKS OF AMERICA
—_———
By Cartos pe LA Torre, Paut Barrscu, and Josepu P. E. Morrison
INTRODUCTION AND ACKNOWLEDGMENTS
Tus contribution is divided into three parts: Parr 1, by Drs. Carlos
de la Torre and Paul Bartsch, deals with the forms inhabiting the
island of Cuba. Parr 2, by Dr. Bartsch, describes the members of
the family known from the West Indies exclusive of Cuba. In Parr 3,
Drs. Bartsch and Joseph P. E. Morrison discuss the mainland fauna.
While Part 3 is by far the largest portion of the paper, it probably
represents only a small fraction of the cyclophorid fauna inhabiting
this extensive territory, the exploration of which has scarcely been
started.
We have not attempted to give complete bibliographic references to
all the species, since this would necessitate an enormous expansion of
the text, but all the citations involving problems of nomenclature have
been noted.
The completeness of this monograph was made possible through
the splendid cooperation of the authorities of the Academy of Natural
Sciences of Philadelphia, particularly Dr. H. A. Pilsbry. We are
indebted also to Prof. H. Burrington Baker, of the University of
Pennsylvania, who unstintingly turned over for our examination all
the material needed in this group, especially from Jamaica. We are
glad, too, to acknowledge our obligation to Dr. d’Alté A. Welch for
his splendid collection of Jamaican cyclophorids, which he placed at
our disposal. The authorities of the Museum of Comparative Zoology
at Cambridge, Mass., through Dr. W. J. Clench, aided materially by
the loan of specimens for study.
The cyclophorid material from Cuba came from several sources,
chief in importance being the extensive collection of Dr. de la Torre,
which furnished the major basis of our understanding of the genus
Farcimen. .Next should be mentioned the large collections made sev-
eral years ago by Dr. Bartsch on several collecting expeditions that
enabled him to comb the island for mollusks. These were made pos-
1
2 THE CYCLOPHORID LAND MOLLUSKS OF AMERICA
sible through the generosity of the late John Brooks Henderson and
through the agency of the Walter Rathbone Bacon Traveling Scholar-
ship of the Smithsonian Institution. Especial thanks are due Prof.
Raul P. Guitart for his helpfulness with specific material needed by
us in the region covered by his explorations, but to mention all the
others who contributed material to our understanding of the fauna of
the Island of Cuba would prolong endlessly these acknowledgments.
We have mentioned under the various species the collectors to whom
we are especially indebted for such contributions,
Our knowledge of the radulas, jaws, and verges discussed in this
paper is based on dissections made by Dr. Morrison.
Part 1.—THE CYCLOPHORID MOLLUSKS OF CUBA
By CARLOS DE LA TORRE AND PAUL BARTSCH
Family CYCLOPHORIDAE Gray
Terrestrial taenioglossate prosobranchiates the mantle cavity of which
has been developed into a thin-walled pulmonary sac. The tentacles
are attenuatedly elongate-conic, bearing the optic papillae at their
outer base. Snout usually short. Foot attenuate posteriorly, entire.
The radula rows consist of seven teeth. The shell shape, size, and
ornamentation are extremely diversified. The operculum, usually
circular, may be a simple chondroid plate or this may bear lamellae
or calcifications of various types.
The family ranges from Mexico south over South America on the
mainland as well as over the West Indian Islands.
In Cuba it is represented by two subfamilies, which the following
key will help to differentiate:
KEY TO THE CUBAN SUBFAMILIES OF THE FAMILY CYCLOPHORIDAE
Sine Cioinez aero) 1iKO Ts ee ee Megalomastominae
Se MMe ce Olek ee ae ee ee Aperostominae
MEGALOMASTOMINAE, new subfamily
Cyclophorid mollusks having an elongate-conic or elongate-turrited
shell. Operculum corneous.
This subfamily embraces the mainland Zomocyclus, which comes
from Guatemala and the adjacent Mexican Chiapas, the Cuban Farci-
men, the Hispaniolan Farcimoides, the Puerto Rican Neopupina,
typical Megalomastoma from the Virgin Islands, and its related sub-
genus Megalomastomoides from eastern Puerto Rico.
Wherever found the animal lives in leaf mulch, or it may even dig
in the loose earth about the base of plants until it is completely
hidden from sight. Bartsch found Farcimen (Neopupina) curtum
(Dall and Simpson) buried completely among the roots of banana
trees on the east slope of El Yunque in Puerto Rico. On days of
protracted rain they may venture forth from their hiding place and
sometimes may climb a foot or so above ground up on the trunk of
a mulch-surrounded tree.
4 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Genus FARCIMEN Troschel
1847. Farcimen TroscHEt, Zeitschr. Malak., vol. 4, p. 44, in part.
1847 (November). Farcimen HERMANNSEN, Indicis generum malacozoorum primor-
dia, vol. 1, p. 489. (Type designated, Cyclostoma tortum Wood.)
Typical Farcimen is known only from Cuba, where it is widely dis-
tributed and presents considerable variation in shell shape and sculp-
ture. In shape it varies from ovate to elongate-conic. The sculpture,
following an initial smooth nuclear portion, may be smooth, axially
lirate, or even ribbed, spiral threads may be present, or the surface
may be pitted as in the Farcimen alutaceum group. The peristome is
heavily reenforced by a thickening and is reflected. The operculum
consists of a simple, thin, multispiral chrondroid plate.
Type: Turbo tortus Wood=Farcimen (Farcimen) tortum (Wood).
The anatomy of /. (F.)/ vinalense scopulorum is described under
that subspecies, p. 34.
All members of the genus Farcimen are mulch-dwellers; they find
their optimum habitat where dead leaves and vegetable detritus be-
come mixed with sand or soil at the base of stone walls or pared6nes or
dead trees. Here usually there is sufficient moisture to furnish ideal
conditions for the development of fungi, upon whose mycelial threads
they seem largely to subsist. Living specimens are therefore rarely
seen on the surface. It is only on wet days or dewy nights that they
appear to come from their hidings and may then climb for some feet
above the level normally occupied by them. Usually they burrow
below the surface and have to be sought by grubbing for them. For
these reasons the shells of most species are partly covered with dirt
when found.
This leaf-mulch dwelling does not tie them to the restricted habitat
that the calciphilous annularids are forced to effect. We therefore
find the various species occupying a much more extended territory than
is the case in the annularids. They are not confined to the moun-
tains but are equally at home in the plains, where their worst enemy
is agriculture, which usually destroys their favorite habitats. Their
greater abundance in the rough hilly country at present merely means
that man’s activities so far have failed to dispossess them.
The absence of strongly developed sculptural characters makes the
genus Farcimen a rather trying group to handle taxonomically.
Words alone seem inadequate to define convincingly our concept of
species and races. Fortunately, by the use of the camera we are able
to portray the characters far better than mere word pictures can accom-
plish. We hope, therefore, that by consulting both descriptions and
figures our contentions will be made clear.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 5
KEY TO THE SUBGENERA OF FARCIMEN
Peristome) thickened/and) retlected2222ee—_ © teas eae lisse ee Farcimen
Peristomemotatnickenedvor) retlecteds2 ses eee Neopupina
Subgenus FARCIMEN Troschel
In the subgenus Farcimen the peristome is thickened and reflected.
The group is peculiar to the island of Cuba.
Type: Turbo tortus Wood=Farcimen (Farcimen) tortum (Wood).
FARCIMEN (FARCIMEN) PSEUDOTORTUM, new species
Megalomastoma tortum of many authors, not of Wood, which equals Farcimen
apertum Poey.
Shell elongate-ovate, the early whorls varying in color from yellow
to pale orange. Nuclear whorls 1.8, well rounded, smooth. Postnu-
clear whorls well rounded, narrowly shouldered at the summit; the
early turns are marked by rather distantly spaced, slender, retractively
curved axial riblets. These riblets on the succeeding turns become
more closely approximated and less regular, and on the last whorl are
less expressed than on the other whorls. In perfect specimens there
is a faint indicaton of obsolete spiral hairlines, at least on the early
turns. Suture rendered conspicuous by the shoulder at the summit.
Base protracted, tapering fairly abruptly toward the open umbilicus.
The umbilicus is bounded by an obsolete carina. Within this the um-
bilical wall appears slightly excavated and is marked by incremental
lines. Aperture almost circular; peristome thickened, heaviest at the
posterior angle and at the junction of the basal lip with the columella,
and narrowest on the parietal wall. Operculum typically farcimenid.
This species is the one that has been considered by many authors
to be F. tortum. We have explained under that species why we do
not agree with this determination. F. (/.) pseudotortum resembles
most nearly forms of F. (F.) wngula (Poey), from which, however,
it is easily distinguished by the fact that the aperture is not protracted
into a clawlike element at the junction of the outer and basal lip.
We are recognizing three subspecies, all occupying parts of Oriente
Province, which the following key will help to differentiate:
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) PSEUDOTORTUM
Umbilicus very narrow.
ShellMarce: hersht more: than) 24min = ee pseudotortum
Shellismalli height less! thant20 mm" 282 ee ee turquinoense
Umbilicusmotivery Napro wes ees lel re CME Eee ee bayamense
6 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) PSEUDOTORTUM PSEUDOTORTUM, new subspecies
PLATE 1, FicurEs 7-9
This race comes from the ancient Cafetal Buena Vista, 18 miles
west-southwest of Bayamo, Oriente Province. The specimen that we
have figured is one collected by Gundlach and labeled UM. tortum. It
is distinguished from the other two by its much larger size and the
ribbing of the early whorls, which is much more conspicuous and a little
more distantly spaced.
The type. U.S.N.M. No. 104424, the specimen referred to above, has
7.5 whorls and measures: Height, 24.9 mm.; greater diameter, 11.8 mm.
FARCIMEN (FARCIMEN) PSEUDOTORTUM TURQUINOENSE, new subspecies
PLATE 1, Ficures 1-3
This race comes from Pico Turquino, Sierra Maestra, Oriente
Province. It is easily distinguished from F. (F.) pseudotortwm
pseudotortum by its small size, and from F. (/.) pseudotortum baya-
mense by its strong ribbing.
The type, U.S.N.M. No. 535874, has 5.3 whorls remaining, the tip
being broken, and measures: Height, 19.3 mm.; greater diameter,
9.9 mm.
FARCIMEN (FARCIMEN) PSEUDOTORTUM BAYAMENSE, new subspecies
PLATE 1, FIGURES 4-6
This race was collected by Victor Rodriguez at the foot of Pan de
Azitcar, Sierra Maestra, southwest of Bayamo. It differs from the
other two forms in being much more rotund and much more openly
umbilicated and in having considerably finer sculpture. The peris-
tome is also more reflected at the junction of the basal lip and columella.
The type, U.S.N.M. No. 535875, has 6.5 whorls and measures:
Height, 18.8 mm.; greater diameter, 10.1 mm.
FARCIMEN (FARCIMEN) YUNQUENSE, new species
PLATE 2, Figures 19-21
Shell very small, pale orange, with the last part of the last whorl
behind the peristome usually red. Nuclear whorls 2, well rounded,
smooth. Postnuclear whorls strongly rounded, narrowly shouldered
at the summit and marked by slender, retractively curved axial riblets,
which are not equal in strength or regularly disposed. Suture strongly
constricted. Periphery well rounded. Base slightly protracted,
rather broadly openly umbilicated, and with an obscure angle at the
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA Z
edge of the umbilicus. Aperture circular; peristome expanded, thick-
ened, and double the width of that of the outer lip on the anterior
columellar wall and the adjacent basal area, and only about one-half
as wide on the posterior columella and parietal side. Operculum
typically farcimenid. The species appears restricted to Yunque de
Baracoa, Oriente Province.
The type, U.S.N.M. No. 535888, was collected by Arango. It has
6.4 whorls and measures: Height, 17.8 mm.; greater diameter, 9.6 mm.
This species was listed as Megalomastoma seminudum by Pfeiffer,
Gundlach, and Arango.
FARCIMEN (FARCIMEN) UNGULA (Poey)
In this species the shell varies immensely in shape and color in the
different subspecies. The form may be elongate-ovate or elongate-
ovate-conic; some individuals are comparatively slender; others, obese.
Most of the specimens, having lost their periostracum, are soiled white
and they range from this through lemon yellow to rose red. The
periostracum is usually thin and translucent. It also varies from
wax color to pale wood brown. The umbilicus varies from open to
closed. Regardless of what the variance mentioned above may be,
the species has in common a thickened peristome, which is protracted
into a clawlike element at the junction of the outer and basal lip.
This species is nearest related to F. (F.) pseudotortum, from which
its ungulate peristome will readily differentiate it.
The species appears confined to Oriente Province, where we are
recognizing nine subspecies, which the following key will help to
differentiate.
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) UNGULA
Umbilicus open.
Shell large, height more than 26 mm.
asthwhorlsoran ge: eas seo ee semivestitum
Masts whorl not oranges. 20-2 See See Fee See holguinense
Shell smaiiers height: 24 mmor less= 232225222 ae perconvexum
Umbilicus closed.
Shell very obese.
Umbilicus bordered with a very conspicuous keel____~__ elephantinum
Umbilicus bordered with an obsolete keel____-__-._------_~- mirandum
Shell not very obese.
Whorls well rounded.
Lip: decidedly, protractenes 2 80 Ts fea eee ee te ungula
ip not decidedly. protracteds=-2 22 ee mayariense
Whorls flattish.
Shelljrose colored = esas Set eee eee ee ee mayense
Shell orange —_--- ele pa ee te ce BAL guantanamense
8 BULLETIN 18 1, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) UNGULA SEMIVESTITUM, new subspecies
PLATE 1, FIGURES 25-27
This race is openly umbilicated. It is much larger than the other
member of that group described here, namely, /. (F.) ungula hol-
guinense. The specimens retaining their periostracum show this to be
wood brown with an olivaceous tinge, streaked with darker axial lines.
When denuded, the last whorl is brilliantly reddish orange. The hair-
like axial threads in this race are stronger and the base of the last
turn is more attenuated than those of &. (7.) ungula holquinense.
The type, U.S.N.M. No. 535884, from Brazo del Pino, Woodfred,
Mayari, Oriente Province, was collected by I. Rivas. It has 6.5 whorls
remaining, having lost the nuclear tip, and measures: Height, 33.0
mm.; greater diameter, 13.8 mm.
FARCIMEN (FARCIMEN) UNGULA HOLGUINENSE, new subspecies
PuaTE 1, Ficures 13-15
The type of this subspecies was collected by Garcia Feria at the
Finca de Garcet, Holguin, Oriente Province. We also have speci-
mens collected by Quesada between Cacocum and San German, and
others obtained by Dr. Aguayo at Baguano, east of Holguin, Oriente
Province.
This race shares with F. (F.) wngula semivestitum the open umbil-
icus. It differs from it in having a much stronger carina limiting the
umbilicus and when denuded in not being brilliant orange on the last
turn, and in having the axial hairlike riblets finer.
The type, U.S.N.M. No. 535877, has 6.8 whorls remaining and mea-
sures: Height, 26.5 mm.; greater diameter, 13.4 mm.
FARCIMEN (FARCIMEN) UNGULA PERCONVEXUM, new subspecies
PLATE 1, Figures 10-12
This is a small race, the type of which was collected by Gundlach at
Yarayabo, Oriente Province. We also have specimens from Central
Palma, Corralillo, northwest of Santiago, from between Baire and
Jiguani, also between Baire and Los Negros, and from San Luis, north
of Santiago. Likewise from Manaca, La Plata. While widely dis-
tributed, it is fairly constant in form, although the coloration varies
materially from flesh colored to roseolate. It is nearest related to F.
(F.) ungula mayariense, from which its much smaller size will readily
differentiate it.
The type, U.S.N.M. No. 535881, has 5.4 whorls remaining and mea-
sures: Height, 24 mm.; greater diameter, 12.5 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 9
FARCIMEN (FARCIMEN) UNGULA ELEPHANTINUM, new subspecies
PLATE 2, FiaurEs 13-15
This very obese race was collected by Dr. S. Molina at Loma Santa
Fe, Guantinamo, Oriente Province. The only member of the ungula
complex resembling this is 7. (F.) ungula mirandum (Pilsbry), from
which its conspicuous tumid umbilical keel distinguishes it. It is also
more inclined toward lemon-yellow, instead of being rosy in coloration,
and the basal lip is more protracted.
The type, U.S.N.M. No. 535887, has 5.0 whorls remaining and mea-
sures: Height, 32.3 mm.; greater diameter, 17.3 mm.
FARCIMEN (FARCIMEN) UNGULA MIRANDUM (Pilsbry)
PLATE 2, Ficurres 16-18
1928. Megalomastoma ungula mirandum Prrspry, Acad. Nat. Sci. Philadelphia
Year Book for 1928, p. 17.
This subspecies was collected by Drs. Pilsbry and d’Alté Welch at
Central Miranda, between Mayari and Santiago, Oriente Province.
It resembles in obeseness F. (F.) ungula elephantinum but is dis-
tinguished by having the shell much more roseolate and the umbilical
bounding keel less conspicuous. Also, the lip is not protracted as in
that subspecies.
The specimen described and figured, U.S.N.M. No. 464468, a topo-
type, has 6.0 whorls remaining and measures: Height, 33.0 mm.;
greater diameter, 15.9 mm.
FARCIMEN (FARCIMEN) UNGULA UNGULA (Poey)
PLATE 1, FicurREs 28-30
1856. Megalomastoma ungula Pory, Memorias sobre la historia natural de la Isla
de Cuba, vol. 2, p. 24, pl. 3, figs. 1-4.
The typical race we have from Santiago, Vista Alegre, San Juan
Hill and the Pedrito Quarry near Santiago, central Oriente, and the
mouth of Magdalena River. The type was collected at Santiago and
given to Poey by Dr. Gutierrez.
The subspecies is of medium size, varying in color from pale lemon
to very faintly roseolate. It has the peristome decidedly protracted
at the junction of the outer and basal lip, giving it a decidedly ungulate
appearance. The umbilicus is practically covered by the reflected peri-
stome, the last whorl being materially attenuated, and terminates in a
fairly conspicuous carina, marking the outer limit of the umbilicus.
The specimen described and figured, U.S.N.M. No. 1044386, is one re-
ceived from Poey collected at Santiago. It has 6.5 whorls remaining
and measures: Height, 29.0 mm.; greatest diameter, 13.2 mm.
10 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
It is distinguished from the other forms with closed umbilicus and
nonobese shape by having the whorls rounded.
FARCIMEN (FARCIMEN) UNGULA MAYARIENSE, new subspecies
PLATE 1, Ficures 19-21
This subspecies was collected by J. B. Henderson at the Farallones
de Nipe, Piedra Gorda, Canapti, near Cayo del Rey, all places near
Mayari. The type comes from Farallones de Nipe.
The race is most nearly related to F. (F.) ungula ungula (Poey),
from which its less protracted aperture and stronger axial sculpture
will readily differentiate it.
The type, U.S.N.M. No. 535883, has 7.2 whorls and measures:
Height, 28.0 mm.; greater diameter, 14.0 mm.
FARCIMEN (FARCIMEN) UNGULA MAYENSE, new subspecies
PLATE 1, FicurEs 16-18
This race was collected by Dr. Ramsden about Alto Songo, La Maya,
Finca Isabelita, and Finca Santa Teresa, Oriente Province. It is a
rose-colored race with closed umbilicus and a strong carina bordering
it. The whorls are marked by rather strong slightly retractively
curved axial riblets, which become weakened on the last turn.
The type, U.S.N.M. No. 535885, from La Maya, has 7.5 whorls re-
maining and measures: Height, 27.9 mm.; greater diameter, 13.5 mm.
The brilliant rose color will differentiate this easily from the other
members with closed umbilicus.
FARCIMEN (FARCIMEN) UNGULA GUANTANAMENSE, new subspecies
PLATE 1, FicurES 22-24
We have this subspecies from a number of stations ranging about
Guantinamo. The individual stations are Vinculo, where the type
was obtained, Saltadero, Sierra des Canasta, and Yateras, Oriente
Province.
This is an orange-colored race with closed umbilicus. It resembles
most nearly /. (F.) ungula mayense, from which its less strong axial
sculpture and paler coloration will differentiate it.
The type, U.S.N.M. No. 535886, has 7.4 whorls and measures:
Height, 18.6 mm.; greater diameter, 13.8 mm.
FARCIMEN (FARCIMEN) CAMAGUEYANUM, new species
Shell of medium size, elongate-ovate, varying in color from uni-
formly wax yellow to bicolor, that is, the early whorls may be pale
and the last darker, frequently dark chestnut-brown. Nuclear whorls
about 1.5, well rounded, smooth. Postnuclear whorls strongly
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 11
rounded, marked by closely spaced, very slender, almost hairlike, re-
tractively curved axial riblets. Suture well marked. Periphery
rounded. Base moderately long, quite openly umbilicated in /. (F.)
camagueyanum mayajiguense, a little less so in F’. (/'.) camagueyanum
camagueyanum, and even less so in F. (F.) camagueyanum florencia-
num, in which the umbilicus is for the greater part concealed. Aper-
ture subcircular; peristome much thickened on the outer, basal, and
inner lip, and reflected; the posterior edge of the reflected peristome
is sharp. The thickened peristome of the inner lip does not quite
reach the preceding turn, but leaves a conspicuous gap. The peristome
of the parietal wall is not thickened. Operculum typically farcimenid.
This species ranges through the north and northwestern portion
of the Sierra Cubitas and extends to the Jatibonico.
We are recognizing three subspecies, which the following key and
descriptions will help to differentiate:
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) CAMAGUEYANUM
IM DUT CUS erat eres Cee eee ee ee ee mayajiguense
Umbilicus not broad.
Shell large, height more than 27 mm-_-__----__----~------- camagueyanum
Shell smaller, height less than 25 mm______--_--------___--- florencianum
FARCIMEN (FARCIMEN) CAMAGUEYANUM MAYAJIGUENSE, new subspecies
PLATE 2, Fiacures 28-30
We have this race from a number of stations, namely, the north
slope of the Sierra de Jatibonico, Veredas de los Broqueles, del
Chorrerén y del Alunado, Sierra de Matahambre, and Sierra de
Canoa, Mayajigua, Santa Clara Province.
This race has a quite regular convex outline with the early whorls
pale olive and the last one much darker, not infrequently purplish.
Its chief distinction, however, lies in the fact that the umbilicus is
more widely open than in any other race and the bounding keel very
conspicuous.
The type, U.S.N.M. No. 535892, comes from the north slope of the
Sierra de Jatibonico at Los Broqueles. Having lost the extreme
tip, the remaining 6.3 whorls measure: Height, 24.5 mm.; greater
diameter, 12.3 mm.
FARCIMEN (FARCIMEN) CAMAGUEYANUM CAMAGUEYANUM, new subspecies
PLATE 2, FIGURES 25-27
This race comes chiefly from the Cubitas Mountains, Camagtiey
Province. Bartsch collected it at Los Corrales de los Cangilones and
at Finca Los Cangilones. Both of these stations are at the foothills
of the Cubitas Mountains. In the mountains it was obtained at
12 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Vereda del Burro in several stations on this path. It was also se-
cured in Paso de la Guanaja, in the Paso de Jas Escaleras, Paso de
los Paredones, and Paso de Lesca. In each of these passes series
of specimens were taken at various stations.
This is the largest race of the group. It is unicolor and has the
umbilicus fairly open, not as wide, however, as in /. (/.) cama-
gueyanum mayajiguense.
The type, U.S.N.M. No. 535898, comes from Los Corrales de los
Cangilones. It has 7.1 whorls and measures: Height, 27.4 mm.;
greater diameter, 14.5 mm.
FARCIMEN (FARCIMEN) CAMAGUEYANUM FLORENCIANUM, new subspecies
PLATE 2, Figures 7-9
This race was collected by Drs. Pilsbry and Welch on the cliffs
near Florencia, Camagiiey Province. The early whorls in this race
are a little darker than in the rest and the last one usually has the
purplish tinge of the two-colored forms, which is also the color
of the interior. The peristome is decidedly expanded, thickened, and
reflected at the junction of the columella and basal lip. It is quite
narrow on the parietal wall and leaves exposed a fairly large por-
tion of the umbilicus, thereby displaying the umbilical carina.
The type, U.S.N.M. No. 535894, has 5.0 whorls remaining and
measures: Height, 21.3 mm.; greater diameter, 11.0 mm.
FARCIMEN (FARCIMEN) WRIGHTI, new species
Shell rather small, varying from elongate-ovate to ovate in out-
line. The color ranges from wax yellow through olive to orange,
and the shells may be unicolor or bicolor, that is, the last whorl may
be chestnut-brown. Nuclear whorls about 1.5, smooth. Postnuclear
whorls well rounded, marked by slender, exceedingly fine, closely
spaced, hairlike axial riblets. Suture strongly constricted. Periph-
ery well rounded. Base narrowly umbilicated, usually almost hid-
den by the reflected peristome. Aperture circular; peristome much
thickened and reflected on the outer, basal, and columellar portion,
much narrower on the parietal wall, where it is somewhat excavated.
The broadly expanded portion of the peristome is always sharp and
knife-edged at the extreme posterior edge of the reflected part. The
peristome of the inner lip touches the preceding turn in Ff. (F.)
wrighti biayaense and F. (F.) wrighti wrighti and almost so in the
other two subspecies.
This species ranges through the southern and eastern complex of
the Sierras de Guaicanamar and Najaza to Sibanicti to the Lomas
de Borje and Santa Cruz, south and east of the Rio Maximo in the
Province of Santa Clara.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 13
We are recognizing four subspecies, as follows:
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) WRIGHTI
Peristome of inner lip almost touching the preceding turn.
Shelli Ova tetas ffi Lbs PEE ee ee Eh oe A ee Tat biayaense
Shellzelongate-ovaite:ssa° 5 ee tise a et Ye Oe ee eee? wrighti
Peristome of inner lip not touching the preceding turn.
Wimbilicaly carina: COnSPICUOUG Soe a= en ee ee eee martianum
Wmibilicalscarinaonot(conspicuouse=s— 2 == eee najazaense
FARCIMEN (FARCIMEN) WRIGHTI BIAYAENSE, new subspecies
PLATE 2, FiguRES 1-3
This subspecies comes from San Martin de Biaya, 15 miles south of
Marti, Camagiiey Province. It is a chubby race in which the early
whorls are pale olivaceous with the last one tending toward reddish.
The interior of the aperture is the color of the exterior of the last whorl.
Its chief distinction from the other members consists of the fact that
the rather broadly expanded inner lip extends up and almost touches
the preceding whorl, leaving a mere line of separation. In this char-
acter it agrees with F. (F.) wrighti wrighti, from which it is readily
distinguished by its gibbose outline. The parietal wall of the peristome
is much narrower than the rest and almost suggests the peculiar lunate
aspect of F. (F.) bituberculatum (Sowerby).
The type, U.S.N.M. No. 535889, has 5.1 whorls remaining, the extreme
tip being lost, and measures: Height, 22.4 mm.; greater diameter,
13.4mm. It was collected by Victor Rodriguez.
FARCIMEN (FARCIMEN) WRIGHTI WRIGHTI, new subspecies
PLATE 2, FIGURES 22-24
Bartsch collected this race in large numbers at Loma Santa Cruz,
near Senado, Camagiiey Province. We have it also from Loma de
Borje and between Puerto Principe and Tunas, where it was collected
by Wright. This subspecies agrees with F. (7.) wrighti biayaense in
the approximation of the expanded lip to the preceding turn. It is,
however, much slenderer than that race, and its color scheme is much
more inclined to bicolor, that is, the early turns wax yellow and the
last one purplish brown.
The type, U.S.N.M. No. 535891, comes from Loma Santa Cruz. It
has 5.6 whorls remaining, having lost the extreme tip, and measures:
Height, 22.7 mm.; greater diameter, 11.8 mm.
Bartsch’s field notes say that the animal of this subspecies is pale
pink, with numerous very small white dots on the back and sides. The
sole of the foot is a little deeper pink than the sides, and the tentacles
are coral-red.
428349—42 2
14 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) WRIGHTI MARTIANUM, new subspecies
PLATE 2, FIcuRES 10-12
We have this race from Finca San Carlos, Marti, which is the type
locality, and also from El Zanjén de Sibanicti, and from San Antonio
Abad, Berrocal, and Guaimaro, Camagiiey Province. This small sub-
species is unicolor orange, with white peristome and white umbilical
region. The peristome is exceedingly expanded and thickened except
at the parietal wall, where it is very narrow and exposes the umbilicus.
In general shape it resembles F. (F.) wrighti biayaense but is readily
distinguished from that by its smaller size and by having the inner lip
decidedly distinct from the preceding turn.
The type, U.S.N.M. No. 535896, which was collected by Victor Rod-
riguez, has 5.8 whorls remaining and measures: Height, 19.3 mm.;
greater diameter, 11.0 mm.
FARCIMEN (FARCIMEN) WRIGHTI NAJAZAENSE, new subspecies
PLATE 2, Figures 4-6
The type of this subspecies comes from El] Cacaotal de Sifontes,
Sierra de Najaza, Camagiiey Province. We have it also from the
Sierra del Cachimbo, which is between the Sierras del Chorrillo and
de Najaza, and from the Sitrra de Guaicanamar, Palomar de San
José, between Guaicanamar and Najaza, and Verada del Telégrafo
in the Sierra del Chorrillo. i.
This race resembles most nearly FP. (F.) wrighti martianum but is
easily distinguished from that by the fact that the peristome is de-
cidedly protracted anteriorly.
The type, U.S.N.M. No. 535897, has 6.9 whorls remaining and meas-
ures: Height, 21.5 mm.; greater diameter, 138.0 mm.
FARCIMEN (FARCIMEN) BITUBERCULATUM (Sowerby)
In this species the shell varies very much in size; some of the races
are of medium stature, while others constitute the largest and most
ponderous shells in the genus. In shape they vary from elongate-
ovate to elongate-conic. In color also there is a great variation;
some races are uniformly pale, others conspicuously bicolor, that is,
the last whorl is purplish plum colored, while the early turns are
usually of a wax tint. In some races both unicolor and bicolor forms
are present. Peristome white with a yellowish tinge. The interior
of the aperture ranges from porcelain white to purplish brown but
is more or less constant for the different races. Nuclear whorls about
1.5, strongly rounded, smooth. The postnuclear whorls, while
rounded, vary materially in convexity in the different races. This is
emphasized by the amount of constriction presented by the suture.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 15
The sculpture consists of retractively curved axial riblets that vary
materially in number and strength in the different subspecies. All
the races show irregularly distributed pitting, which produces an
appearance of drilling by some parasite or of partial decay. This
character shows a tendency toward approaching Farcimen (Farci-
men) alutaceum ([Menke] Pfeiffer), in which it reaches a maximum
development. Periphery well rounded. Base well rounded, or some-
what produced, narrowly umbilicated, with or without a limiting
umbilical carina. The umbilicus in most races is concealed by the
widely expanded and reflected inner peristome. Aperture large, sub-
circular. Peristome much thickened, expanded, and reflected, ex-
cept that of the parietal wall, always wider on the columella than the
rest, and narrow and somewhat excised on the parietal wall. At the
posterior angle of the aperture and at the posterior columellar ex-
pansion, the peristome is thickened into more or less of a knob, which
furnished the specific name bituberculatum. Operculum typically
farcimenid,
The species appears confined to the southern part of the Province
of Santa Clara, where it reaches its maximum differentiation in the
Trinidad Mountain region.
We are recognizing five subspecies:
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) BITUBERCULATUM
Whorls strongly rounded.
SHI rr CO re ee a a ket ee bituberculatum
Sheep COLO rs eee ee tee NU ee ey Se Ee eee See eee minor
Whorls not strongly rounded.
PACH ED NTT) Spe eT SUI ea Ne iva ts GTS CD a giganteum
Axial ribs not irregular or interrupted.
Axial rib Ets) Strong ee BS he eee gutierrezi
FASXAa Te TIDIELS NOt. SUT OW as aes See ee ee ee crassum
FARCIMEN (FARCIMEN) BITUBERCULATUM BITUBERCULATUM (Sowerby)
PLATE 3, FrcurEs 10-12
1850. Cyclostoma bituberculatum SowrErspy, Thesaurus Conchyliorum, vol. 1,
Suppl, p. 164*, pl. 31a, figs. 290-291.
This race was collected somewhere in the mountains of Trinidad,
Santa Clara Province. It differs from all the others in being much
slenderer, unicolor, and much more regularly conic, with the whorls
strongly rounded, in which respect it resembles /. (7.) bitwberculatum
minor (Kobelt).
The specimen figured, U.S.N.M. No. 535901, is the only one we have
that agrees with Sowerby’s figure. It was obtained by Dr. de la Torre
from the ancient collection of D. Tomas Iradi in Cienfuegos. It has
4.9 whorls remaining and measures: Height, 32.7 mm.; greater di-
ameter, 17.3 mm.
16 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) BITUBERCULATUM MINOR (Kobelt)
PLATE 3, FicurEs 16-18
1843. Cyclostoma auriculatum SoweErBy, Thesaurus Conchyliorum, vol. 1, p. 151,
pl. 31, fig. 277 (not Cyclostoma auriculatum Orbigny, 1842).
1902. Megalomastoma (Farcimen) bituberculatum minor Koper, Das Tierreich,
Cyclophoridae, p. 265.
This subspecies was collected by H. N. Lowe at Puriales, Santa Rosa,
Trinidad, Santa Clara Province. It agrees with the typical race, P.
(F.) bituberculatum bituberculatum (Sowerby), in having the whorls
strongly rounded but differs from it in being more ovate and in having
the last whor! plum colored.
The specimen figured, U.S.N.M. No. 535902, has 5.0 whorls remain-
ing and measures: Height, 28.5 mm.; greater diameter, 15.3 mm.
FARCIMEN (FARCIMEN) BITUBERCULATUM GIGANTEUM, new subspecies
PLATE 8, Figures 1-3
We have this subspecies from Sopimpa, a railroad station between
Fomento and Trinidad, and Loma del Caballete, Sierra de Gavilanes
of the Trinidad region, Santa Clara Province. It differs from the
other races of /. (F.) bituberculatum in having the axial ribs of the
later turns irregularly developed; in size it resembles F’. (F'.) betuber-
culatum gutierrezi and F. (F.) bituberculatum crassum, from both of
which the irregularity of its ribs will distinguish it.
The type, U.S.N.M. No. 535904, which comes from Sopimpa, has 5.0
whorls remaining, and measures: Height, 33.7 mm.; greater diameter,
17.8 mm.
FARCIMEN (FARCIMEN) BITUBERCULATUM GUTIERREZI, new subspecies
PLATE 3, Fiaures 4-6
We have this race from a number of stations ranging around Banao,
south of Sancti Spiritus and southeast of Trinidad, Santa Clara
Province; for example, Finca Angelina, La Quinta, El Cacahual, and
Guayacanes; also from Loma de Jarao, Blanquizal, near Guayos.
This is a large pale race, as large as if not larger than 7. (/.)
bituberculatum giganteum, from which it is easily differentiated by
being unicolor. The two races are also zoogeographically distinct.
The type, U.S.N.M. No. 535906, comes from Finca Angelina and
has 5.4 whorls remaining. It measures: Height, 36.8 mm.; greater
diameter, 18.6 mm,
FARCIMEN (FARCIMEN) BITUBERCULATUM CRASSUM, new subspecies
PLATE 3, FicuREs 13-15
This subspecies comes from the southern mountain complex of
Santa Clara Province. We have it from Finca Pitajones, the Cara-
cusey Valley, Trinidad; Loma de Gabino Galvez; Loma Esperanza
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 1y
and Las Damas near Guayos; Finca El Silencio and Cortinas de Veloso,
Canttii; Loma Cariblanca, Fomento, Trinidad; and Loma Caja de
Agua, Tuinict, Sancti Spiritus.
This race is large like /’. (7'.) bituberculatum gutierrezi, which it re-
sembles in many ways but from which it differs by having the peri-
stome very much intensified and in having the axial riblets decidedly
reduced, almost obsolete on the last whorl.
The type, U.S.N.M. No. 535907, was collected by Prof. Pedro Guitart
at Finca Pitajones, Trinidad. It has 4.6 whorls remaining and
measures : Height, 32.8 mm.; greater diameter, 17.8 mm.
FARCIMEN (FARCIMEN) SEMINUDUM (Poey)
Shell varying very greatly in size in the different subspecies. The
shape is equally variable, ranging from elongate-turrited to ovate.
The early whorls are orange-red, or brownish tinted, or even flesh
colored. The last whorl] is dark reddish brown with a purplish tinge.
Nuclear whorls about 1.5, small, well rounded, smooth. The post-
nuclear whorls range from well rounded to inflated. The axial sculp-
ture on the early whorls consists of quite regular and regularly spaced
riblets. On the last turn these may evanesce or become intensified and
irregular. Suture well constricted, rendered conspicuous by the slight
shoulder at the summit of the whorls. Periphery well rounded. Base
moderately openly umbilicated with a carina of varying strength limit-
ing the umbilicus on the outside. Aperture subcircular; peristome
greatly thickened and expanded on the outer, basal, and inner lip,
narrow on the parietal portion, thin and sharp at the posterior edge.
The peristome is somewhat protracted on the anterior columellar por-
tion. Operculum typically farcimenid.
The species appears limited to Santa Clara Province, whence we are
recognizing six subspecies:
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) SEMINUDUM
ast whorl almost Smooth 2s oes 21 es ey Dee ts STs aD magister
Last whorl not smooth.
Axial riblets very fine.
shellilargzesheizhtmore thany2imine sae ee ee collare
Shell.small, height less than, 22,mm- 2-2-2 2) ee poeyi
Axial riblets not very fine.
Axial ribs of last whorl irregular.
SS GTI Vick Gee Ses ee eee FD Oh a a lek seminudum
Shel? elongate-cvatelates fhe eee ee eRe guitarti
Axial ribs of lastswhorl motinregwlars 2525 sn ea ee ern Pe leoni
FARCIMEN (FARCIMEN) SEMINUDUM MAGISTER, new subspecies
PLATE 4, FiIcuRES 19-21
This subspecies comes from the mountains about Trinidad, Santa
Clara Province. We have it from Nacimiento del Rio Caballero
near Letran and from La Vigia.
18 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
It is easily distinguished from the other subspecies by its very
large size and much smoother surface, in which respect it approaches
F. (F.) seminudum poeyi.
The type, U.S.N.M No. 535914, which comes from Nacimiento
del Rio Caballero, has 6.4 whorls remaining and measures: Height,
33.0 mm.; greater diameter, 15.4 mm.
FARCIMEN (FARCIMEN) SEMINUDUM COLLARE (Pcey MSS.), new subspecies
Piate 4, Ficures 7-9
This subspecies is rather widely distributed. It ranges from
Corrallilo eastward along the hills of the north coast to the Sierra
de Meneses, Santa Clara Province. We have it also from a few
scattered places to the south of this range. Through this range we
find considerable variation, not so much in size and general shape
as in the strength of the peristome and the amount of its reflection
over the umbilicus, which is also of variable width, sometimes being
quite open. There is likewise considerable variation in the strength
of the obsolete angle at the outer limitation of the umbilicus. Any
of these characters, if constant, might suggest subspecific distine-
tion, but in the present instance this is precluded because in the
large series of specimens before us we find that the variations re-
ferred to may be present in any of the population groups examined.
Almost 60 lots, most of them representing a considerable series, are
before us. The subspecies is much larger than F. (/.) seminudum
poeyi, with which it shares the fine axial ribbing.
As a rule, in perfect specimens the early whorls are covered by
a thin golden-yellow periostracum; the later whorls are dark, and
by far the largest percentage of specimens are bicolor, though uni-
color individuals are not rare.
The specimen described and figured, U.S.N.M. No. 535909, comes
from La Cantarilla, San Juan de los Yeras, Santa Clara Province.
It has 5.5 whorls remaining and measures: Height, 28.6 mm.; greater
diameter, 14.7 mm.
FARCIMEN (FARCIMEN) SEMINUDUM POEYI, new subspecies
Pirate 4, FIGuRES 25-27
We have this subspecies from Loma de Santa Fé, Camajuani;
San Juan de los Remedios, near Santa Clara; Orillas del Rio Cala-
bazas, Placetas; Los Damas near Guayos; Central Mapos, west of
Sabana, Sancti Spiritus.
This subspecies and F. (F.) s. collare are distinguished from all
the others by having very fine, closely spaced axial riblets, which
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 19
are quite regular on the last turn. It differs from /. (F.) s. collare
by its much smaller size.
The type, U.S.N.M. No. 535917, comes from Loma de Santa Fé.
It has 6.5 whorls and measures: Height, 20.0 mm.; greater diameter,
11.8 mm.
FARCIMEN (FARCIMEN) SEMINUDUM SEMINUDUM (Poey)
PLATE 4, FIGURES 22-24
1854. Megalomastoma seminudum Pory, Memorias sobre la historia natural de la
Isla de Cuba, vol. 1, pp. 405-406.
This subspecies comes from the region of Trinidad. We have it
from La Vigia, Fomento, and Giiinia de Soto, and from Arroyo La
Lima, Finca el Retiro, and Cagiieiras in Sancti Spiritus.
This race is small compared with /. (F.) seminudum magister, from
which it is also readily distinguished by the irregularity of the axial
riblets of the last whorl. It resembles most closely F. (¥.) seminudum
guitarti, from which it differs in having the shell much more inflated.
The specimen figured, U.S.N.M. No. 535915, is one received from
Poey without specific locality. It has 5.0 whorls remaining and meas-
ures: Height, 22.8 mm.; greater diameter, 11.7 mm.
FARCIMEN (FARCIMEN) SEMINUDUM GUITARTI, new subspecies
PLATE 4, FI1GUREs 1-3
This subspecies was collected by Guitart at Finca Caimiabo, Sancti
Spiritus. It is a small race, with the axial riblets on the last turn
irregularly developed. It resembles most nearly the typical /. (F.)
seminudum seminudum (Poey), from which it differs by its smaller
size and much less inflated turns.
The type, U.S.N.M. No. 535916, has 5.2 whorls remaining and meas-
ures: Height, 21.0 mm.; greater diameter, 10.4 mm.
FARCIMEN (FARCIMEN) SEMINUDUM LEONI, new subspecies
PLATE 4, Ficures 16-18
This subspecies was collected by Hermano Leén and Professor Gui-
tart at Hornos de Cal, Vista Alegre, Sancti Spiritus, Santa Clara
Province. It isa little smaller than /. (F.) seminudum magister and
has the axial ribs more strongly developed. It is much larger than
the other three subspecies here listed, differing also from F. (F.) semi-
nudum poey? in having the axial ribs stronger and from the other two
smaller races in having them regular on the last whorl.
The type, U.S.N.M. No. 535908, has 4.5 whorls remaining and meas-
ures: Height, 25.4 mm.; greater diameter, 14.0 mm.
20 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) TORREI (Guitart)
PLATE 6, FIGURES 4-6
1936. Megalomastoma torrei Guirart, Mem. Soc. Cubana Hist. Nat. “Felipe
Poey,” 1936, pp. 105-106, figs. 1, 2.
Shell very small, elongate-ovate. The early whorls are wax yellow
with an orange flush, the last one being darker. The nuclear whorls
are decollated. Postnuclear whorls rather well rounded, marked by
slender, almost hairlike, slightly retractively curved axial riblets which
are about one-third as wide as the spaces that separate them. These
riblets become a little more distantly spaced on the last whorl. Suture
well constricted. Periphery well rounded. Base slightly protracted,
openly, moderately broadly umbilicated, the umbilical angle marked
by an obsolete cord. Aperture circular; peristome thickened, except
on the parietal wall, where it is much thinner. Operculum typically
farcimenid.
The specimens in our collection are paratypes received from Guitart
and were collected on the banks of the Rio Manajanabo. The speci-
men described and figured, U.S.N.M. No. 535918, has 4.5 whorls re-
maining and measures: Height, 16.9 mm.; greater diameter, 8.4 mm.
FARCIMEN (FARCIMEN) OBESUM, new species
Shell varying from medium size to large, pale chestnut-brown, with
the peristome pale yellow. The nucleus consists of a little more than
one turn, which is well rounded and smooth. Postnuclear whorls
rather inflated, moderately well rounded, marked by almost vertical,
gently curved axial riblets which are about as wide as the spaces that
separate them. Suture well impressed. Periphery well rounded.
Base rather openly umbilicated with an obtuse carina marking the
outer edge of the umbilicus. The umbilical wall is marked by the
continuation of the axial riblets. Aperture circular, oblique, decidedly
expanded; the peristome is broadly expanded, reflected, and decidedly
thickened, somewhat auriculated at the posterior angle and also at
the posterior portion of the inner lip, which does not touch the pre-
ceding turn. Operculum typically farcimenid.
This species occupies the northeastern mountains of Santa Clara
Province.
We are recognizing two subspecies, which the following key and
descriptions will help to differentiate :
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) OBESUM
Diameter more than 17, mmo 8.) = ae obesum
Diameter Tess scien ly) WN ee re subobesum
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA mk
FARCIMEN (FARCIMEN) OBESUM OBESUM, new subspecies
PLATE 3, Fiacures 7-9
This race was collected by Bartsch on Loma Platero. It is much
more gibbose than the smaller eastern race, /’. (7.) 0. subobesum.
We have also seen specimens of it from Seibado de Yaguajay and from
El Resbalillo, Cambao, Yaguajay, and Remedios.
The type, U.S.N.M. No. 535908, comes from Loma Platero. It has
4.5 whorls remaining and measures: Height, 31.4 mm.; greater diam-
eter, 18.6 mm.
Bartsch described the animal of specimens collected on Loma Platero
as being of a general pale pink tone with buffish snout. The whole body
except the sole of the foot is covered with small flattened tubercles,
which are marked by numerous fine white dots. Tentacles bright
coral-red. Sole of foot pale pink.
FARCIMEN (FARCIMEN) OBESUM SUBOBESUM, new subspecies
PLATE 3, FIGURES 19-21
This race ranges through the mogotes about Remedios and Yagua-
jay. We have it from the Dos Sierras, Central San Agustin; Caicaje,
Loma de Bueno Vista and the north slope of the Sierra de Meneses
east of Yaguajay, also opposite Jungalito. It can be easily distin-
guished from typical F. (/.) obesum obesum by its much smaller size
and less gibbose form.
The type, U.S.N.M. No. 516710, comes from the paredénes on the
north side of the second mogote west of the gap in the Dos Sierras,
Santa Clara Province. It has 6 whorls remaining and measures:
Height, 28.1 mm.; greater diameter, 14.5 mm.
The specimens collected by Bartsch at the Central San Agustin and
in the Sierra de Jatibonico, opposite Los Broquelles, were described
by him as having the entire animal flesh colored with a rosy flush,
with the tentacles bright coral-red. The papillae of the sides with
numerous white dots.
FARCIMEN (FARCIMEN) ALUTACEUM ([Menke] Pfeiffer)
Shell elongate-ovate, varying very greatly in size in the different
subspecies. The early whorls are wax yellow; this changes to dull
orange on the succeeding turns and finally to dull reddish brown, or
dull plum purple on the last whorl. Peristome white with a yellow
tinge. Interior of the aperture reddish brown. Nuclear whorls
almost 2, strongly rounded, smooth. Postnuclear whorls strongly
rounded, the first marked by more or less regular, retractively slant-
22 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
ing axial riblets which on the succeeding whorls break up into shorter
elements to form a very complex and irregular network of meshes,
between which deep pits are present. The pittings rather than the
meshes that enclose them catch one’s eye. Suture deeply constricted.
Periphery well rounded. Base somewhat produced, openly umbili-
cated, with the edge of the umbilicus more or less carinated, not
pitted. Umbilical wall marked by lines of growth only. Aperture
oval with the long axis oblique to that of the spire. Peristome de-
cidedly expanded, thickened, and reflected, excepting the parietal
portion, widest on the columella, more or less alate at the posterior
part of the columella and outer lip. Operculum typically farcimenid.
This species is confined to the Trinidad Mountain complex of
southern Santa Clara Province.
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) ALUTACEUM
Shell large, height about 30 mm__----___----------------_------- magnificum
Shell medium: sized, height about 25 mm-=__-==_--____ "===" "== =__= alutaceum
Shell small, height about 17 mm__---------------------------------- digitale
FARCIMEN (FARCIMEN) ALUTACEUM MAGNIFICUM, new subspecies
PLATE 4, FicurEs 13-15
This subspecies occurs at Sopimpa, Santa Clara Province. The
shell of this race is very large, agreeing in shape and sculpture with
F. (F.) alutaceum alutaceum ({Menke] Pfeiffer) and F. (F.) alu-
taceum digitale (Gundlach), from which it differs by a little weaker
pitting on the last turn and by its gigantic size.
The type, U.S.N.M. No. 535898, has 5.3 whorls remaining and
measures: Height, 30.8 mm.; greater diameter, 15.4 mm.
FARCIMEN (FARCIMEN) ALUTACEUM ALUTACEUM ([Menke] Pfeiffer)
PLATE 4, FicurEs 10-12
1846. Cyclostoma alutaceum (Menke) Prerrrrr, Zeitschr. Malak., vol. 3, pp.
85-86.
We have this typical race from Quemado Feo, north of Trinidad.
It is intermediate in size between F. (F.) alutaceum, magnificum and
F. (F.) alutaceum digitale (Gundlach). It has the whorls a little less
convex.
The specimen described and figured, U.S.N.M. No. 585899, has 4.5
whorls remaining and measures: Height, 25.1 mm.; greater diameter,
13.3 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 23
FARCIMEN (FARCIMEN) ALUTACEUM DIGITALE (Gundlach)
PLATE 4, FIGuRES 4-6
1857. Megalomastoma digitale GunpLACH, Malak. Blitter, vol. 4, p. 114.
We have this small race from Letrén and Puriales in Trinidad;
Soledad near Cienfuegos; Ternero; and Sierra de San Juan, Santa
Clara Province. The small size of this subspecies will easily differ-
entiate it from the rest.
The specimen described and figured, U.S.N.M. No. 535900, comes
from Letrén and has 5 whorls remaining. It measures: Height,
16.4 mm.; greater diameter, 10.1 mm.
FARCIMEN (FARCIMEN) ROCAI, new species
PLATE 5, Ficures 13-15
Shell of medium size, the early whorls pale yellowish, the last chest-
nut-brown; peristome yellowish white. The nucleus consists of a
little more than one well-rounded smooth turn. The postnuclear
whorls are well rounded and marked by irregular, slightly retractively
curved, somewhat sinuous, poorly developed axial riblets, which
branch and anastomose in an irregular manner. In addition to this
there are fine irregular pittings suggesting those of #. (/.) alutaceum
([Menke] Pfeiffer). Suture well constricted. Periphery well
rounded. Base sloping suddenly toward the obsolete carina that
marks the outer edge of the open umbilicus. The umbilical wall is
marked by the continuation of the incremental lines. Aperture cir-
cular; peristome double, the outer decidedly expanded, thin at the
edge, concave, turned up into conspicuous auricles at the posterior
angle and at the posterior termination of the outer peristome of the
inner lip. It is very narrow on the parietal wall, touching the pre-
ceding whorl only in the middle. Operculum typically farcimenid.
The type, U. S. N. M. No. 535905, was collected by Father Roca at
Cabezadas del Caracusey, which is between Trinidad and Sancti
Spiritus, Santa Clara Province. It has 5.4 whorls remaining and
measures: Height, 17.0 mm.; greater diameter, 15.8 mm.
The irregular sculpture and “worm-eaten” pittings suggest a close
affinity with F. (F.) alutaceum, from which it can readily be dis-
tinguished by the sharp-edged outer peristome, which lacks the heavy
thickening characteristic of /. (F.) alutacewm. Prof. P. Guitart has
also collected specimens of this at Cortinas de Corengue and Veguetas
de Buenos Aires.
24 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) AURICULATUM (Orbigny)
Moderately large shells of very elongate-ovate outline. Early
whorls flesh colored or orange in one race; the last or last two whorls
purplish brown. Nuclear whorls about 1.3, small, well rounded,
smooth. Postnuclear whorls well rounded, appressed at the summit,
marked by fairly regularly developed, slender, retractively curved
axial riblets, which on the last whorl are almost sigmoid; these riblets
are narrower than the spaces that separate them. In addition to this,
the whorls show distantly, irregularly distributed, small, round pits
suggestive of Ff. (f.) alutaceum ({Menke] Pfeiffer). Suture mod-
erately constricted. Periphery well rounded. Base slightly pro-
tracted, moderately openly umbilicated and marked like the spire.
Aperture subcircular; peristome broadly expanded, except that of
the parietal wall, much thickened, with the posterior edge of the
reflection of the outer lip sharp. The peristome of the inner lip is
decidedly produced, which lends to the aperture an earlike appear-
ance, hence the name. It does not approach the preceding whorl but
is separated from it by a large gap. A knoblike thickening may be
present at the posterior termination of the peristome of both the inner
and outer lip. Operculum typically farcimenid.
We are recognizing four subspecies, three of which appear confined
to the southwest side of Santa Clara and southeast of Matanzas
Provinces, while the fourth stretches from the north coast at Cardenas
southward to Coliseo and San Miguel.
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) AURICULATUM
Peristome veryemuch thickened] 2) 22) Sess eee senectum
Peristome not very much thickened.
arly whorls Oranges. 22 =. 2. a See ee eee clenchi
Early whorls pale.
Shell large) height;more than; 29 mms sarees eee ee bicolor
Shell smaller, heightiless: than (26mm 22" See auriculatum
FARCIMEN (FARCIMEN) AURICULATUM SENECTUM, new subspecies
PLATE 5, FicuREs 16-18
Two specimens from Cayo Carenas, near Cienfuegos, collected by
Henderson appear to be semifossil. They differ very greatly from the
other subspecies in having the peristome enormously thickened and
decidedly effused on the columellar side.
The type, U.S.N.M. No. 535911, has 5 whorls remaining and meas-
ures: Height, 26.5 mm.; greater diameter, 14.1 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 20
FARCIMEN (FARCIMEN) AURICULATUM CLENCHI, new subspecies
PLATE 5, Figures 7-9
We have this subspecies from Colonia Guabairo, Central Soledad,
Cienfuegos; and Hatiguanico, Cabo Zapata, Santa Clara Province.
It is much larger than the typical F. (f.) auriculatum auriculatum
(Orbigny) and much darker in coloration, the early whorls usually
being deep orange or the next to the last rosy and the last plum colored.
The type, U.S.N.M. No. 535913, comes from Guabairo. It has 5.3
whorls remaining and measures: Height, 30.0 mm.; greater diameter,
14.7 mm.
FARCIMEN (FARCIMEN) AURICULATUM BICOLOR (Gould)
PLATE 5, Ficures 19-21
1844. Cyclostoma auriculatum bicolor GovuLp, Boston Journ. Nat. Hist., vol. 4,
cover p. 494.
This race centers around Coliseo; we have it also from Los Chivos,
Cimarrones, near Cardenas, Matanzas Province. It resembles most
nearly the typical race in shape, sculpture, and coloration, also in the
shape of the aperture, but it is uniformly larger.
The specimen figured, U.S.N.M. No. 535912, was collected by Bartsch
on the steep paredénes west of the pass at Coliseo. It has 7.8 whorls
remaining and measures: Height, 30.3 mm.; greater diameter, 14.0 mm.
FARCIMEN (FARCIMEN) AURICULATUM AURICULATUM (Orbigny)
PLATE 5, FicurEes 1-3
1842. Cyclostoma auriculatum Orpteny, in Sagra’s Histoire physique, politique et
naturelle de l’Ile de Cuba, vol. 1, p. 257, pl. 22, figs. 1, 2.
1851. Cyclostoma solenatum PorEy, Memorias sobre la historia natural de la Isla
de Cuba, vol. 1, pl. 7, figs. 17, 18.
1852. Oyclostoma idolum (Férussac MS.) Premrer, Monographia pneumono-
pomorum viventium, vol. 1, p. 180.
We have seen this subspecies from the following localities: Zara-
banda and Yaguaramas, Santa Clara Province. It differs from F.
(F.) auriculatum bicolor (Gould), with which it agrees most closely, in
the shape of the aperture and in being much smaller; it is less dark in
color than Ff. (7.) auwriculatum clenchi.
The specimen figured, U.S.N.M. No. 535910, which comes from
Zarabanda, has 6.0 whorls remaining and measures: Height, 25.0 mm. ;
greater diameter, 12.7 mm.
26 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) TORTUM (Wood)
PLATE 5, FiaurEs 10-12, 22-24
1828. Turbo tortus Woop, Index testaceologicus, ed. 2, Suppl., p. 20, fig. 32.
1828. Cyclostoma tortum Woon, ibid., p. 36, pl. 6, fig. 32.
1851. Cyclostoma apertum Pory. Memorias sobre la historia natural de la Isla de
Cuba, vol. 1, pl. 7, figs. 15, 16.
1854. Megalomastoma apertum Poey, ibid., p. 405.
Shell of medium size, elongate-ovate. Early whorls wax yellow,
turning to brown on the last turn. Peristome white. Nuclear whorls
1.7, well rounded, smooth. Postnuclear whorls strongly rounded, the
early ones marked by very regular, very closely spaced, hairlike axial
riblets, which evanesce on the last turn. Faint indications of spiral
lirations are also present. Suture strongly constricted, rendered con-
spicuous by the very narrowly shouldered summit of the whorls.
The last whorl is rather long, bounded anteriorly by a low rounded
carina, which marks the outer edge of the moderately broad open
umbilicus. The umbilical wall is marked by axial threads, which
are stronger here than on the outside of the last turn. Aperture
oblique, subcircular; peristome very much thickened, reflected, nar-
rower on the parietal wall and the columella. Operculum typically
farcimenid.
The two specimens figured, U.S.N.M. No. 516857, came from Los
Palos, Nueva Paz, Habana Province. They were collected by Arango.
One, a complete specimen, has 6.5 whorls; the other has 5.2 whorls
remaining. They measure, respectively: Height, 27.0, 23.5 mm.;
greater diameter, 14.0, 18.9 mm.
There are 38 lots in the collection of the United States National
Museum, all of which fall within the range mentioned above.
Wood’s small figure has caused endless confusion. Most authors
believed that it represented what we are here calling Farcimen pseudo-
tortum, which occupies a region of eastern Cuba that was not explored
in 1828. The aperture as figured by Wood is not quite normal for
F. apertum Poey, a common species in Habana Province, but speci-
mens with the characters here depicted are not infrequent. The
swollen, overhanging middle whorl is also of not infrequent occur-
rence in F. apertum Poey, all of which leads us to believe that F.
tortum is without question Farcimen apertum Poey, which name it
must replace.
In 1851 Poey (loc. cit.) gave two figures of what he called Cyclos-
toma apertum. Evidently through correspondence with Pfeiffer he
was led to believe that his figures represented a variety of F. auricu-
latum, for in 1854 (doc. cit.) he states that the figures were bad. Here
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA Zh
he gives a description of the shell and cites the south coast of Cuba
as the home of the species. He also mentiones the ingenio (sugar
mill) of Don J. Poey near Alacranes as a source of some of his material.
We believe that Poey’s figures are not at all bad, that they repre-
sent quite well the species that occurs abundantly between Habana
and Matanzas. Many specimens distributed by Poey with the name
Megalomastoma apertum belong to this species and add additional
support to our conclusion. Also, Gundlach, in Pfeiffer (Malakozool.
Blatter, vol. 8, p. 122, 1856), describes the animal of M/egalomastoma
apertum and cites therewith Loma de Camoa as the first definite lo-
cality for the species, which we shall consider the restricted type
designation. A translation of Gundlach’s description of the animal
is as follows:
Animal beautiful rose-red with coral-red antennae, small black
eyes on the outer bases of the short antennae. In moving about the
head is projected very slowly about two lines beyond the shell and
the shell is then rapidly drawn after it. The caudal portion does
not project beyond the operculum.
FARCIMEN (FARCIMEN) GUNDLACHI (Pfeiffer)
Medium-sized shells with the early whorls wax color or a little
dusky; the later turns considerably darker, in some cases even plum
colored. Nuclear whorls small, a little more than 2, well rounded,
smooth. Postnuclear whorls inflated, rather strongly rounded,
marked by mere incremental lines or slender riblets, which vanish
on the middle of the turns and are most emphasized on the posterior
portion. There are faint indications of spiral hair lines. These
may be mere color markings. Suture moderately well constricted.
Periphery well rounded. Base protracted, openly umbilicated with a
strongly rounded carina marking the outer limit of the umbilicus.
Aperture almost circular; peristome broadly expanded and reflected
except on the inner lip, which is narrower and adnate to the preced-
ing turn. Operculum typically farcimenid.
The species is confined to the eastern end of Pinar del Rio and the
western part of Habana Provinces where three races are recognizable.
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) GUNDLACHI
Axinerinlets obsolete :.. <2 ke nee Ad eS ae ek ee ee gundlachi
Axial riblets not obsolete.
Shell’-smaill: heizht less'than 20 mm eee gundlachiellum
Shelldarger; height more) than) 22; mma 2) ae ee ee anafense
28 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) GUNDLACHI GUNDLACHI (Pfeiffer)
PLATE 6, FIGURES 22-24
1856. Megalomastoma gundlachi Pretrrer, Malak. Blitter, vol. 3, pp. 48, 121.
1857. Megalomastoma gundlachi Prrirrer, Noy. Conch., vol. 1, p. 96, pl. 26,
figs. 20-22.
Henderson and Bartsch collected this subspecies at various places
in the Cuzco Mountain complex. For example, El Mulo; the upper
reaches of the Rio Hondo; Pefia Blanca (El Huro) ; Lomas de Cuzco;
and Salto de Manantiales. The typical subspecies is remarkably
smooth, with scarcely any indication of riblets at the summit, in
which respect it differs from F. (#.) gundlachi gundlachiellum,
which has slight ribbing on the posterior half of the turns.
The specimen figured, U.S.N.M. No. 535920, comes from Loma del
Cuzco (El Mulo). It has 7.0 whorls and measures: Height, 24.3
mm.; greater diameter, 11.0 mm.
FARCIMEN (FARCIMEN) GUNDLACHI GUNDLACHIELLUM, new subspecies
Puats 6, Ficures 1-3
We have this small race from the mountains north of Candelaria,
such as El Taburete, Cayajabos, and Mogote Soton. Its smaller size
will readily distinguish it from the other two subspecies.
The type, U.S.N.M. No. 535924, coming from El Taburete, Caya-
jabos, has 5.1 whorls remaining and measures: Height, 19.6 mm.;
greater diameter, 10.8 mm.
Bartsch describes the animal of this subspecies, which he collected
at Soton on a hillside west of Taburete, June 11, 1928, as follows:
Animal flesh colored with a pinkish suffusion. The tentacles in some
specimens are a trifle pinker than the general tone of the body; in
others they are coral-red. Eyes black, situated at the outer base of
the tentacles.
FARCIMEN (FARCIMEN) GUNDLACHI ANAFENSE, new subspecies
PLATE 6, Figures 25-27
This subspecies appears to be restricted to the Sierra de Anafe.
We have a lot from Guanajay, Pinar del Rio Province, however,
which agrees in every way with it. It resembles most nearly the
typical race but is easily distinguished from that by having the axial
ribs of the early whorls much finer and more closely spaced. The
last whorl is also smoother.
The type, U.S.N.M. No. 535923, comes from Loma de Anafe. It
has 7.0 whorls and measures: Height, 22.3 mm.; greater diameter,
11.0 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 29
FARCIMEN (FARCIMEN) LEONINUM (Pfeiffer)
Shell varying from small to medium sized, elongate-ovate, usually
wax yellow or with the last whorl much darker, even plum colored.
Nuclear whorls about 2, quite small, well rounded, smooth. Post-
nuclear whorls somewhat inflated, weil rounded, marked by slightly
sinuous axial riblets, which vary materially in spacing and develop-
ment in the two races here recognized. Suture moderately con-
stricted. Periphery well rounded. Base protracted, marked by an
obsolete carina at the junction with the umbilicus, which is moder-
ately open. Aperture subcircular; peristome thickened and reflected,
fairly uniform in size excepting that of the parietal wall, which is
much narrower and adnate to the preceding turn. Operculum typi-
cally farcimenid.
The species is confined to the central part of Pinar del Rio Prov-
ince. We are recognizing two subspecies, which the following key
will help to differentiate:
KEY TO SUBSPECIES OF FARCIMEN (FARCIMEN) LEONINUM
Shelislarce: height more chane24 mm Ses ee eee eee leoninum
Shellysmallsnerent jess) than’ 24) mms ie ee eee leonellum
FARCIMEN (FARCIMEN) LEONINUM LEONINUM (Pfeiffer)
PLATE 5, FIGURES 25-27
1856. Megalomastoma leoninum PFEIFFER, Malak. Blitter, vol. 3, pp. 48, 122.
1857. Megalomastoma leoninum PFEIFFER, Nov. Conch., vol. 1, p. 97, pl. 26,
figs. 23-25.
This medium-sized race comes from the higher reaches about
Rangel. The axial ribs are comparatively strong and rather dis-
tantly spaced, particularly upon the early whorls. Its larger size
will easily differentiate this subspecies from F. (F.) leoninum
leonellum.
The specimen figured, U.S.N.M. No. 535922, comes from the coffee
plantation on the summit of Rangel. It has 5.4 whorls remaining and
measures: Height, 25.0 mm.; greater diameter, 12.5 mm.
FARCIMEN (FARCIMEN) LEONINUM LEONELLUM, new subspecies
PLaTH 5, Fiacurrs 46
We have this subspecies from San José, Rangel; Sierra de Limones,
and also from a station on the road between Rangel and Guajaibon.
Jt differs from the typical race in being considerably smaller and
darker colored and in having the anterior portion of the whorls only
slightly ribbed.
428349—42—_3
30 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The type, U.S.N.M. No. 535921, which comes from San José, Rangel,
has 5.4 whorls remaining and measures: Height, 20.3 mm.; greater
diameter, 10.3 mm.
FARCIMEN (FARCIMEN) MANI (Poey)
PLATE 6, Ficures 16-18
1851. Cyclostoma mani Pory, Memorias sobre la historia natural de la Isla de
Cuba, vol. 1, pl. 7, figs. 19-22.
1854. Megalomastoma mani PorY, ibid., pp. 404-405.
Shell moderately large, elongate-ovate, wax colored throughout,
or with the last whorl brownish or even purplish. Peristome pale
yellow. Nuclear whorls about 1.7, small, well rounded, smooth. Post-
nuclear whorls inflated, strongly rounded, and marked on the early
turns by closely spaced, slender axial riblets, which are as fine as
hairs. On the middle whorl these become quite reduced and on the
later ones obsolete. Suture rather strongly constricted. Periphery
well rounded. Base slightly protracted, narrowly umbilicated, with
an obsolete carina marking the outer edge of the umbilicus. Aperture
subcircular; peristome broadly expanded, particularly so on the inner
lip, where it is decidedly produced, a little narrower on the parietal
wall, where it is adnate to the preceding turn.
The specimen described and figured, a topotype, U.S.N.M. No.
535925, was received from Poey. It has 7.3 whorls remaining and
measures: Height, 26.1 mm.; greater diameter, 13.5 mm.
This species comes from the general region of Rangel and the Santa
Cruz River, and extends northward to Rancho Lucas in the Mount
Guajaibon region.
FARCIMEN (FARCIMEN) VENTRICOSUM (Orbigny)
PLATE 6, Fiaures 13-15
1842. Cyclostoma ventricosa Orpieny, in Sagra’s Histoire physique, politique et
naturelle de l’Ile de Cuba, vol. 1, p. 256, pl. 21, fig. 18.
Shell ovate, decidedly inflated, pale orange. Nuclear whorls 1.5,
small, well rounded, smooth. The postnuclear whorls increase very
rapidly, inflated, strongly rounded and marked on the early whorls by
fine, almost hairlike, retractively curved axial riblets, which on the
first turn are very distantly spaced, while on the succeeding turns
they become decidedly closely approximated, being separated on the
second turn by spaces about as wide as the riblets. On the last three
turns they are obsolete or rather replaced by irregular and irregu-
larly distributed incremental lines. The summit of the whorls is
slightly narrowly shouldered, which renders the suture rather con-
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 31
spicuous. Periphery inflated, strongly rounded. Base short, well
rounded, with a rounded carina marking the outer edge of the open
umbilicus. Aperture almost circular; peristome rather broadly ex-
panded, thickened, and marked by concentric lines of growth of
about equal width, except that of the parietal wall, which is much
narrower. Operculum typically farcimenid.
This species is restricted to Pan de Guajaibén, where it is very
abundant about Hato Sagua. The specimen described and figured,
U.S.N.M. No. 535928, is one of a large series collected by Henderson
and Bartsch. It has 7.8 whorls and measures: Height, 29.3 mm.;
greater diameter, 17.2 mm.
This species is easily distinguished from all other Farcimens by its
stout, short, decidedly inflated form.
FARCIMEN (FARCIMEN) SUBVENTRICOSUM, new species
Shell varying greatly in form in the different subspecies; in some
it is elongate-ovoid; in others, elongate-conic. In color the shells
vary from soiled wax color to pale orange to pale brown; they may
be unicolor or darker on the later turns, and the last may even be
purplish plum colored. Nuclear whorls almost 2, small, well rounded,
smooth. Postnuclear whorls inflated, strongly rounded. In one sub-
species they are marked by rather strong, well-defined axial ribs on
all but the last turn. In others, the axial riblets are in the form of
hair lines on the early turns, evanescing shortly thereafter. Suture
constricted to a varying degree in the different races. Periphery
well rounded. Base moderately produced in all the races, moderately
broadly openly umbilicated with a rounded carina at the outer limit
of the umbilicus. Aperture subcircular; peristome moderately ex-
panded, decidedly thickened and reflected, protracted on the columel-
lar wall and decidedly narrowed on the parietal wall and marked by
concentric lines of growth. Operculum typically farcimenid.
This species ranges from the Rio San Diego north of San Diego de
los Bafos, through the Organos Mountains to Pan de Guajaibén,
through the Sierra Chiquita, west through the Sierra de Galalén and
Sierra de Pico Chico; the Sierra de San Andrés to Vifiales.
We are recognizing three subspecies, which the following key
will help to distinguish:
KEY TO SUBSPECIES OF FARCIMEN (FARCIMEN) SUBVENTRICOSUM
Axial -ribletsirather:strongsdse ssaleien ten wiyoe aA EY oh multistriatinum
Axial riblets not strong.
HEMP STEN Gert: teat WORT aye ened cp iterbeal Cit 2 ge Olga a eae le te si aia! balneorum
SHEMFESCO pea ee NON a CON RE RN is WR RiNL eS ot subventricosum
32 BULLETIN 181, UNITED STATES) NATIONAL MUSEUM
FARCIMEN (FARCIMEN) SUBVENTRICOSUM MULTISTRIATINUM, new subspecies
PLATE 6, Ficures 19-21
This race appears to extend from Los Lagunitas, near the Sierra de
Galalén to San José and to certain of the mogotes of the San Diego
de los Bafos complex. It is easily differentiated from the others by
having rather strong regular axial riblets on all but the last whorl.
The type, U.S.N.M. No. 535934, comes from Los Lagunitas. It has
6.2 whorls remaining and measures: Height, 31.5 mm.; greater diam-
eter, 15.8 mm.
FARCIMEN (FARCIMEN) SUBVENTRICOSUM BALNEORUM, new subspecies
PLATE 6, Ficgures 10-12
This race occupies the mogotes and mountainsides bordering the
Rio San Diego, north of San Diego de los Banos, apparently on both
sides of the river. It is a smaller and slenderer race than /. (/.) sub-
ventricosum multistriatinum. Italso has the ribbing merely indicated.
The type, U.S.N.M. No. 535935, comes from the Cueva de los Indios
on the east bank of the Rio San Diego. It has 8 whorls and measures:
Height, 28.2 mm.; greater diameter, 12.8 mm.
FARCIMEN (FARCIMEN) SUBVENTRICOSUM SUBVENTRICOSUM, new subspecies
PLATE 6, FicurEs 7-9
This subspecies extends from Pan de Guajaibén westward through
the Organos Mountains to Viniales. It is a subventricose race whose
whorls are narrowly shouldered at the summit and whose axial sculp-
ture is reduced to mere lines of growth. It somewhat suggests /. (/.)
ventricosum (Orbigny), but is easily distinguished from that species
by its less inflated form. It differs from the other subspecies by its
more inflated form and less strong sculpture.
The type, U.S.N.M. No. 535933, comes from the northeast end of
Pan de Guajaibén. It has 6.5 whorls remaining and measures: Height,
30.3 mm.; greater diameter, 15.7 mm.
FARCIMEN (FARCIMEN) VINALENSE, new species
Shell elongate-conic, varying materially in outline, unicolor, wax
colored or pale orange or with the last whorl much darker, even plum
colored; peristome white or pale yellow. Nuclear whorls two, well
rounded, smooth. The early postnuclear whorls are marked by slen-
der, rather feebly developed axial threads, which vary materially in
spacing in the two subspecies. The succeeding whorls are inflated,
strongly rounded and marked by less strong threads, which gradually
grow weaker and evanesce on the last whorl. Suture strongly con-
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 33
stricted. Base moderately prolonged, bounded by a rounded carina
at the junction with the open umbilicus. Aperture almost circular;
peristome broadly expanded, thickened, and reflected, broadest on the
columellar wall where it is protracted, narrowest on the parietal wall;
peristome is marked by concentric lines of growth. Operculum typi-
cally farcimenid.
We are recognizing two subspecies. which the following key will
help to differentiate :
KEY TO SUBSPECIES OF FARCIMEN (FARCIMEN) VINALENSE
Shell) elongait Ge 4- Booe. ots. s co rhe tions Mew ate eT ed oes ev ohn te ihe vinalense
SHV UT er 0 6 ge or ee tee i aT a ES Fa er scopulorum
FARCIMEN (FARCIMEN) VINALENSE VINALENSE, new subspecies
PLATE 7, Ficures 4-6
This subspecies ranges about the region of Vifiales. We have it
from the Ensenada de los Bafios; the valley east of Bafos de San
Vicente; Cueva de las Delicias, Sierra de Vifales; the paredén of
the Puerta del Ancén; Sierra de la Chorrera and Hoyo de Jaruco.
It is a decidedly narrow race, of almost cylindric outline, with very
rounded whorls, which readily distinguish it from the more regu-
larly conic F. (F.) vinalense scopulorum.
The type, U.S.N.M. No. 535929, comes from the Ensenada de los
Banos. It has 8 whorls and measures: Height, 28.8 mm.; greater
diameter, 14.1 mm.
FARCIMEN (FARCIMEN) VINALENSE SCOPULORUM, new subspecies
PLATE 7, FIGuRES 7-9
We have this subspecies from the mogotes around Kilometer 14
between Pinar del Rio and Vinales, also from the mogotes adjacent
to the road between Vinales and the Puerta del Ancon, for example:
The Mogotes de la Vega, Capon, Coco Solo, Puertecitas, Trujillo,
Palmarito, Zacarias, Dinamita, Rojas, Cuajani, Rinconada, and Vigil.
This race differs from typical F. (f.) vinalense vinalense in being
smaller, less elongated, more conic, and stouter and in having the
whorls slightly less rounded.
The type, U.S.N.M. No. 535930, comes from the mogote on the
northeast side of the road at Kilometer 14, between Pinar del Rio and
Vinales. It has 8 whorls and measures: Height, 27.3 mm.; greater
diameter 14.4 mm.
Dr. Joseph P. E. Morrison has dissected specimens of this sub-
species and has furnished us with the following description based
upon alcoholic material, No. 6674, collected by Bartsch on Mogote
de la Dinamita.
34 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The foot is short, oval, with undivided sole. The snout is of
medium length, tapering, conical, truncate at the tip. The tentacles
are short, slightly tapering, but blunt at their tips. The eyes are
small, borne on slight prominences at the outer base of the tentacles.
The verge is conspicuous in male individuals. It is attached on the
side of the neck, lateral to the right tentacle, and furnished only
with a seminal groove. The verge is basally somewhat flattened, and
a little swollen in appearance, gradually tapered to a subcylindrical
glandular portion near the tip. The seminal groove begins at the
opening of the vas deferens and extends diagonally across the body
surface to the posterior basal point of attachment of the verge, and
continues on the under or posterior side of the verge, to the narrowly
rounded tip. Ordinarily the terminal part of the verge is looped
or folded upon itself, along the side of the foot.
The jaw is composed of two triangular pieces hinged together
dorsally. The median edge of each jaw is thickened into an obscure
ridge which projects as a distinct cutting point, on the free edge of
the jaw. The surface of the jaw is marked with diagonal striations
(or rows of diamond-shaped scalelike elements) ; running nearly for-
ward at the median line, these extend over the ridge, where they
gradually change direction, to extend diagonally to the cutting edge.
The radula has the characteristic formula of the family Cyclo-
phoridae. The central is tricuspid, much higher than wide. The
lateral is tricuspid, oblique, the central cusp much larger than the
other two. The inner marginal is somewhat oblique, and bears three
cusps, the middle one twice as long as the two smaller ones. The
shorter, outer marginal tooth bears only two subequal cusps on its
cutting edge.
The reduction of cusps to two on the outer marginal tooth appears
to be characteristic of the genus Farcimen.
FARCIMEN (FARCIMEN) SUPERBUM, new species
Shell large, rather stout, very elongate-ovate. Early whorls wax
yellow, gradually turning darker, the major tint being chestnut-
brown; the last whorl may even be purplish. Nuclear whorls about 2,
small, well-rounded, smooth. Postnuclear whorls inflated, strongly
rounded ; the early ones marked by slender, well-elevated axial riblets,
which are rather distantly spaced. These riblets evanesce, and on the
last two turns they are indicated as mere lines of growth. Suture
strongly constricted. Periphery well rounded. Base rather short,
well rounded, moderately broadly openly umbilicated, with a rounded
carina at the outer termination of the umbilical wall. Aperture circu-
lar; peristome decidedly thickened and reflected, produced on the
middle of the columella wall, and much narrower and somewhat sinu-
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA a
ous on the parietal wall, where it is adnate to the preceding turn.
Operculum typically farcimenid.
We are recognizing two subspecies, which the following key will
differentiate :
KEY TO SUBSPECIES OF FARCIMEN (FARCIMEN) SUPERBUM
Shellglarce shershinabpoubl oOsm Mae ae ee re superbum
Shell§smaiiler, height about/Z26.mme2_ = eee itinerarium
FARCIMEN (FARCIMEN) SUPERBUM SUPERBUM, new subspecies
PLATE 7, FIGURES 28-30
This subspecies occurs on the mountains forming the rim of Los
Acostas and Luis Lazo and extends from there eastward on the north
side through Pan de Azticar and Costanera del Abra to the Costanera
de San Vicente. On the south side it extends through Isabel Maria
to the Sierra del Infierno. This race differs from F. (F.) swperbum
itinerarium in being much larger with the whorls more inflated and
more rounded.
The type, U.S.N.M. No. 535931, comes from the Sierra de los Acos-
tas, Luis Lazo, Pinar del Rio. It has 8 whorls and measures: Height,
32.5 mm.; greater diameter, 16.5 mm.
FARCIMEN (FARCIMEN) SUPERBUM ITINERARIUM, new subspecies
PLATE 7, FieuRES 10-12
This subspecies extends from the Potrerito at Luis Lazo, through the
mogotes bordering the road from there to Sumidero, Cabezas and
Tsabel Maria, in Pinar del Rio Province.
It is smaller than the typical race and is more inclined to have the
last whorl] purplish.
The type, U.S.N.M. No. 535932, comes from Potrerito. It has 7.1
whorls and measures: Height, 26.6 mm.; greater diameter, 14.1 mm.
FARCIMEN (FARCIMEN) HENDERSONI, new species
Shell of medium size, elongate-ovate, varying in color on the early
whorls from flesh colored to pale chestnut-brown, while the later
turns are always darker; they may be unicolor, but usually tend
toward purplish plum colored. Nuclear whorls about 2, small, well
rounded, smooth. Postnuclear whorls inflated, strongly rounded,
marked in the one race by rather strong, retractively curved axial
riblets; the other is almost without these. Suture quite strongly
constricted. Periphery well rounded. Base only moderately pro-
duced, openly moderately umbilicated. Aperture circular; peri-
stome very much thickened and reflected, marked by concentric lines
36 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
of growth, produced on the columella and much narrower on the
parietal wall than on the rest. Operculum typically farcimenid.
We are recognizing two subspecies, which the following key will
help to distinguish:
KEY TO SUBSPECIES OF FARCIMEN (FARCIMEN) HENDERSONI
Acetic hare Let ter Orn ee a catalinense
Asal riblets obsolete. 2-2 2s) sane ee eee hendersoni
FARCIMEN (FARCIMEN) HENDERSONI CATALINENSE, new subspecies
PLATE 7, FIGURES 22-24
This race comes from La Catalina north of San Diego de los
Bafios. We also have it from La Cumbre and Hato Caimito west of
Pan de Guajaibén, Pinar del Rio Province. This is smaller than
typical F. (F.) hendersoni and has very pronounced, distantly spaced,
slender riblets, which are present even on the last turn.
The type, U.S.N.M. No. 535937, comes from Cayito de la Catalina.
It has 5.5 whorls remaining and measures: Height, 22.8 mm.; greater
diameter, 11.0 mm.
FARCIMEN (FARCIMEN) HENDERSONI HENDERSONI, new subspecies
Puate 7, Figures 138-15
This subspecies has a rather wide distribution, extending from
Paso Real through Teneria de Guane, through La Murrallia eastward
to Giiira de Luis Lazo; Lagunillas; Mogote del Cerro de Cabras to
Cayo San Felipe; Consolacion del Sur and Entronque Herradura,
Pinar del Rio Province. On the north side it extends through
Mogote Pan de Azticar, the valley east of Bafios de San Vicente
through La Mina and Bella Maria, Pinar del Rio Province. This
race is a little larger than F. (F.) hendersoni catalinense and has
the whorls almost smooth, while catalinense has them strongly ribbed.
The type, U.S.N.M. No. 535936, comes from the Mogote del Cerro
de Cabras. It has 7.1 whorls and measures: Height, 24.0 mm.;
greater diameter, 12.3 mm.
FARCIMEN (FARCIMEN) ARANGOI, new species
PLATE 7, FIacuRES 16-18
Shell very small, ovate. Nuclear whorls decollated. Postnuclear
whorls very strongly inflated, rounded, and marked by slender, re-
tractively curved axial riblets, which grow irregular and less dif-
ferentiated on the last turn. Suture strongly constricted. Base
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA ay
short, inflated, strongly rounded, and marked by an obsolete keel at
the outer rim of the umbilicus. Aperture circular; peristome only
moderately expanded, reflected, and slightly produced on the basal
lip, a little wider on the columellar wall than on the rest, almost as
wide on the parietal wall as on the outer lip. The operculum is not
known.
The type, U.S.N.M. No. 535938, was collected by Arango at La
Caja, Pinar del Rio Province. It has 5.4 whorls remaining and
measures: Height, 16.1 mm.; greater diameter, 8.3 mm.
FARCIMEN (FARCIMEN) GUANENSE, new species
Shell elongate-ovate. Nuclear whorls white, the rest pale orange,
or the last whorl may even be partly or wholly brilliant orange or
purplish. Nuclear whorls 1.5, small, well rounded, smooth. Post-
nuclear whorls well rounded, narrowly shouldered at the summit,
the first three marked by slender, well-raised, rather closely spaced,
and retractively slanting axial riblets, while on the remaining turns
these become obsolete or are merely indicated near the summit.
Suture well constricted. Periphery well rounded. Base protracted,
openly umbilicated, marked by a rounded carina at the umbilical junc-
tion. Aperture circular; peristome rather broadly expanded and re-
flected, protracted on the columella, narrower on the parietal wall
and marked by concentric lines of growth. Operculum typically
farcimenid.
We are recognizing two subspecies, which the following key will
help to differentiate :
KEY TO THE SUBSPECIES OF FARCIMEN (FARCIMEN) GUANENSE
Suenllarse abort 25 Mim Nigh. 2s see er) a bed ERA TO guanense
Shellsmall)aboute23 smn, (hig ee eee ee eee ee eee lagunillense
FARCIMEN (FARCIMEN) GUANENSE GUANENSE, new subspecies
PLATE 7, FIGURES 1-3
This race comes from the region of Guane. We have it from the
caves of the mogotes on the south side of the road opposite the Sierra
de Guane; Los Portales; La Murrallia; and from the mogote at Punta
de la Sierra, Pinar del Rio Province. It is distinguished from F. (F.)
guanense lagunillense by being much larger, more elongate and with
a more protracted base.
The type, U.S.N.M. No. 535926, from Guane, has 7.6 whorls and
measures: Height, 29.2 mm.; greater diameter, 13.4 mm.
\
38 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
FARCIMEN (FARCIMEN) GUANENSE LAGUNILLENSE, new subspecies
PLATE 7, FrcurEs 19-21
This small race was collected by Wright at Lagunillas in the Dis-
trict of San Juan Martinez, Pinar del Rio Province. It differs from
the typical race in being much smaller, having the last whorl pro-
tracted, and having the basal two-thirds of the last whorl orange col-
ored or darker.
The type, U.S.N.M. No. 535927, has 6.4 whorls remaining and meas-
ures: Height, 23.0 mm.; greater diameter, 12.1 mm.
FARCIMEN (FARCIMEN) PROCER (Poey)
PLATE 7, Figures 25-27
1852. Cyclostoma tortum var, Pory, Memorias sobre la historia natural de la Isla
de Cuba, vol. 1, pl. 18, figs. 12-18.
1854. Megalomastoma procer PoEY, ibid., p. 404.
1856. Megalomastoma complanatum PFEIFFER, Proc. Zool. Soc. London, vol. 24,
p. 36.
Shell rather large, pale chestnut-brown with an olivaceous flush;
the early whorls lighter and the last one darkest. Peristome white;
interior of the aperture reflecting the external coloration. Nuclear
whorls almost 2, small, well rounded, smooth. The postnuclear whorls
are well rounded and marked by slightly retractively curved, slender,
hairlike axial riblets on the early turns, which grow less strong on the
succeeding whorls and become obsolete on the last. Suture slightly
constricted. Periphery well rounded. Base somewhat produced,
with an obsolete shoulder at the outer margin of the moderately wide-
open umbilicus. Aperture subcircular; peristome broadly expanded,
widest on the inner lip, thick, reflected. The posterior portion of the
expanded lip almost touches the preceding turn and largely hides the
umbilicus when viewed squarely. The peristome on the parietal wall
is exceedingly narrow. Operculum typically farcimenid.
This species comes from the Isle of Pines, where we collected it in
large numbers on the Sierra de Casas, Sierra de Caballus, Sierra de
Colombo, and the Morrillo del Diablo. In the latter place they appear
to be a trifle smaller.
The specimen described and figured, U.S.N.M. No. 535939, was col-
lected by Bartsch at the northwestern part of the Sierra de Casas. It
has almost 7 whorls remaining and measures: Height, 32.7 mm.;
greater diameter, 15.9 mm.
APEROSTOMINAE, new subfamily
Cyclophorid mollusks having a helicoid shell and an operculum
bearing calcifications on its external surface.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 39
Genus CROCIDOPOMA Shuttleworth
1857. Crocidopoma SHUTTLEWORTH, Journ. Conchyl., vol. 5, pp. 271-272.
1891. Crocidopoma CrossE, Journ. Conchyl., vol. 39, p. 160. (Type, Crocidopoma
jfloccosum (Shuttleworth).)
This genus embraces small aperostomid mollusks whose whorls are
marked by strong spiral cords and whose operculum bears a strongly
elevated and outwardly reflected calcified lamella. The basal
chondroid plate extends beyond the turns of the calcified lamella and
breaks up on the outside into slender, thin, fluted, and striated
fimbriations.
Type: Cyclostoma (Cyclotus) floccosum Shuttleworth=Croci-
dopoma (Crocidopoma) floccosum (Shuttleworth).
The genus appears to be confined to Cuba and Hispaniola. It breaks
up into two subgenera, the typical one of which, Crocidopoma, is con-
fined to Hispaniola, while Cyclocubana is restricted to Cuba.
KEY TO THE SUBGENERA OF CROCIDOPOMA
Fimbriations of chondroid plate extending above edge of calcareous
lamellae Sete Se Meeks ses ee ere ee renee a Crocidopoma
Fimbriations of chondroid plate not extending above edge of cal-
CATEOUS seam el cy ese aaah Bae a Ee Cyclocubana
CYCLOCUBANA, new subgenus
In this subgenus the fimbriations of the outer edge of the chondroid
basal plate extend but little if at all beyond the outer edge of the
calcareous lamella.
Type: Cyclotus perdistinctus Gundlach=Crocidopoma (Cyclocu-
bana) perdistinctum (Gundlach).
KEY TO THE SPECIES OF THE SUBGENUS CYCLOCUBANA
Shebiplanorpoidyse 2tis ithe ON i eA ed Oo UL Oe perdistinctum
Shelividepressedshelicoidy se wet 55 eee ees yest del peter hel sped ysl 5 gundlachi
CROCIDOPOMA (CYCLOCUBANA) PERDISTINCTUM (Gundlach)
Shell planorboid, horn colored. Nuclear whorls about 1.5. Post-
nuclear whorls very strongly rounded, circular in cross-section with
strongly channeled suture, marked by well-raised spiral lirations vary-
ing in number in the different subspecies. The axial sculpture con-
sists of incremental lines. The last whorl may or may not be solute.
Operculum typical of that of the subgenus.
This species differs from Crocidopoma (Cyclocubana) gundlachi
in having the spire very depressed; in the latter, it is depressed-heli-
coid. We are recognizing two subspecies:
40 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
KEY TO THE SUBSPECIES OF CROCIDOPOMA (CYCLOCUBANA) PERDISTINOTUM
Spiral cords ‘at: peristomie 242022) ee ee ee tes toroense
Spirallicords) at; peristome) 322 2a ae ee eee perdistinctum
CROCIDOPOMA (CYCLOCUBANA) PERDISTINCTUM TOROENSE, new subspecies
PLATE 8, Figures 10-12
Shell planorboid. Nuclear whorls 1.5, strongly rounded, smooth.
Postnuclear whorls very much rounded, circular in cross section,
marked by strongly elevated spiral cords, which are of almost equal
strength and are separated by spaces about three times as wide as
the cords. Of these cords 9 occur between the summit and the
periphery and 15 upon the remainder of the shell. The axial sculpture
consists of incremental lines and somewhat rough resting spaces.
Suture very deeply openly channeled. The last whorl is solute for
about one-twentieth of a turn. Base very broadly openly umbilicated.
Operculum as described for the subgenus.
The type, U.S.N.M. No. 355777, was collected by Gundlach at
Monte Toro, Guantanamo, Oriente Province. It has 4 whorls and
measures: Height, 4.2 mm.; greater diameter, 10.2 mm.; lesser diam-
eter, 6.9 mm.
The lesser number of spiral cords will differentiate this from typical
O.(C.) perdistinctum perdistinctum (Gundlach).
CROCIDOPOMA (CYCLOCUBANA) PERDISTINCTUM PERDISTINCTUM (Gundlach)
PLATE 8, FiaurES 13-15
1858. Cyclotus perdistinctus GUNDLACH, Malak, Blitter, vol. 5, pp. 192-193.
Shell small, planorboid, thin, wax colored, with a greenish tinge.
Nuclear whorls 1.5, well rounded, smooth, the last half turn showing
the merest beginning of the spiral sculpture of the succeeding turns,
forming a very slightly elevated apex. Postnuclear whorls circular
in cross section, marked by strongly elevated, slender spiral cords,
which increase in number with the growth of the shell by intercalation.
On the first postnuclear turn 4 of these cords are present on the spire ;
on the second, 7 between the summit and the periphery, while on the
last turn 31 can be counted in the circle of the peristome of the aper-
ture. These spiral cords vary in strength, depending upon whether
they represent intercalated elements or the major cord. They are
never so wide as the spaces that separate them and rarely wider than
one-half of the spaces. In addition to the spiral sculpture the shell
is marked by slender incremental lines, which render the free edge of
the cord slightly roughened. The last twentieth of a turn is solute.
The underside shows a broad open umbilicus more or less funnel
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 41
shaped, which shows all the whorls within. The inside of the oper-
culum is convex, slightly calcified, showing all the turns, and the
outside shows 10 turns of an oblique, outward-turned, calcified
lamella, which is marked by slender striations, the turns of the
lamella overlapping each other somewhat like shingles on a roof,
leaving, however, a narrow space between them.
The specimen described and figured, U.S.N.M. No. 535776, was col-
lected at San Andrés, near Reuter, Oriente Province by Dr. Ramsden,
who has furnished information showing that this falls well within
Gundlach’s concept of Jurisdiccién, Cuba, his type locality. It has 4.1
turns and measures: Height, 4.4 mm; greater diameter, 11.0 mm;
lesser diameter, 7.6 mm.
A translation of Gundlach’s statement relative to the animal of this
species follows: I am using here the generic name C'yclotus because I
consider the species more nearly related to Megalomastoma than Cyclo-
stoma. The animal is very similar to that of Megalomastoma, as well
as its mode of life under decaying leaves in the dark forest at the
sugar plantation Felicitas in the region of Enramada. The animal is
bright rose-red, with the head a little darker within, the antennae are
cinnabar red and the forehead crossed by wrinkles. In crawling the
always very dirty shell is carried almost perpendicular.
CROCIDOPOMA (CYCLOCUBANA) GUNDLACHI, new species
Shell depressed-helicoid, horn colored. Nuclear whorls about 1.5,
vell rounded, smooth; postnuclear whorls strongly rounded, circular
in cross section, with strongly channeled suture. The postnuclear
whorls are marked by narrow, strongly elevated spiral cords, which
vary in number in the different subspecies and which increase in num-
ber by intercalation as the shell increases in size. The spaces between
the spiral cords are wider than the spiral cords and they, as well as the
cords, are crossed by incremental lines. Periphery well rounded; base
broadly, openly umbilicated, and marked like the spire; the same
sculpture extends into the umbilical wall. The last portion of the
last whorl may be adnate to the preceding turn or slightly solute.
The operculum is typically that of the subgenus.
The species occupies the mountain area of Oriente Province. We
are recognizing three subspecies:
KEY TO THE SUBSPECIES OF CROCIDOPOMA (CYCLOCUBANA) GUNDLACHI
Spiral cordsrless than! 255 eesaee yet eee a OE ee ee ignotum
Spiral cords more than 30.
Greater diameter! more than 0 imme se 42 hes eee ee Bee See gundlachi
Greater diameterless) than Oimmeso) ee oS ee oe wrighti
42 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
CROCIDOPOMA (CYCLOCUBANA) GUNDLACHI IGNOTUM, new subspecies
PLATE 8, Figures 7-9
In the Evezard collection of the National Museum there are two
specimens of a subspecies for which we have no specific locality. They
are distinguished from the rest of the material at hand and agree per-
fectly with each other. We are, therefore, reluctantly bestowing a sub-
specific name upon them.
The type, U.S.N.M. No. 585775, has 4.3 whorls and shows at the aper-
ture 10 spiral cords between the summit and the suture, 7 on the base
and 7 on the umbilical wall. This measures: Height, 4.8 mm; greater
diameter, 9.0 mm; lesser diameter, 7.0 mm.
The other specimen, U.S.N.M. No. 316425, has 4.2 whorls and the
same number of spiral cords as the type, and measures: Height, 7.4
mm.; greater diameter, 8.2 mm.; lesser diameter, 6.8 mm.
The extremely small number of spiral cords differentiates it from the
other two races.
CROCIDOPOMA (CYCLOCUBANA) GUNDLACHI GUNDLACHI, new subspecies
PLATE 8, FIGURES 1-3
This subspecies was collected by Gundlach at Monte Toro.
The type, U.S.N.M. No. 535773, has 4 whorls and at the aperture
shows the following number of cords: Between the summit and the
periphery 10, on the base 10, on the umbilicus 16. It yields the follow-
ing measurements: Height, 6.8 mm.; greater diameter, 10.9 mm. ; lesser
diameter, 8.4 mm.
CROCIDOPOMA (CYCLOCUBANA) GUNDLACHI WRIGHTI, new subspecies
PLATE 8, Figures 4-6
In the National Museum we have four specimens collected by Charles
Wright at Potosi, Oriente Province. Potosi, while not on recent maps,
from information at hand we know is situated somewhere north of
Monte Libano and Felicidad. These specimens are differentiated from
the other two members by being a little more closely coiled.
The type, U.S.N.M. No. 535774, has 4.1 whorls and measures: Height,
5.1 mm.; greater diameter, 8.9 mm.; lesser diameter, 6.9 mm. It has 12
spiral cords between the summit and the suture, 11 on the base, and 8
on the umbilical wall, a total of 31 when viewed at the aperture.
The other two specimens are complete and yield, respectively, the
following information: Number of whorls, 4.1 and 3.7; number of
spiral cords between the summit and suture, 12 and 12; spiral cords on
base, 10 and 8; spiral cords on the umbilical wall, 10 and 10. They
measure respectively: Height, 5.3 and 5.0 mm.; greater diameter, 8.3
and 8.2 mm.; lesser diameter, 6.5 and 6.2 mm.
Part 2—THE CYCLOPHORID MOLLUSKS OF THE
WEST INDIES, EXCLUSIVE OF CUBA
By PAUL BARTSCH
Family CYCLOPHORIDAE Gray
For a definition of the family, see p. 3.
KEY TO THE ANTILLEAN SUBFAMILIES OF THE FAMILY CYCLOPHORIDAE
Shell elongate-turrited:=- 1:5) 28105: es 1 ees Se Megalomastominae
Shell not elongate-turrited.
Shelli pmpaidhse se Os a es ee AS Diplommatininae
Shell not pupoid, but planorboid or helicoid.
Operculum chondroid___~---__----------__-----_-_- Amphicyclotinae
Operculum ealcified—-__- = == = ES SS Aperostominae
Subfamily MEGALOMASTOMINAE Torre and Bartsch
Cyclophorid mollusks having an elongate-conic or elongate-turrited
shell. Operculum corneous.
For complete diagnosis, see p. 3.
KEY TO THE WEST INDIAN GENERA OF MEGALOMASTOMINAE
Shell large and robust.
\uWdavanch Cove Coyarerenuhwbon) saris ee Farcimen
Wihorls'omoperculum distinct sae es se ieee Se eee Farcimoides
Shell not large and robust, but small and thin______-----~-_~ Megalomastoma
Genus FARCIMEN Troschel
For diagnosis of the genus, see p. 4.
The subgenus Farcimen is restricted to the island of Cuba and is
therefore not considered in this part of our paper.
Subgenus NEOPUPINA Kobelt
1854. Megaloma Woopwarp, A manual of the Mollusca, p. 177 (not Megaloma
Hall, 1852).
1856. Lomastoma Woopwakrp, ibid., p. xviii (not Rafinesque, 1819).
1902. Neopupina Kosett, Das Tierreich, Cyclophoridae, p. 262.
This subgenus differs from the Cuban Farcimen in having a more
cylindric shell and in having the peristome sharp, i. e., not broadly
thickened and reflected. Operculum a thin chondroid multispiral
plate.
43
44 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Type: Megalomastoma flavula Swainson = Helix crocea Gmelin =
Farcimen (Neopupina) eroceum (Gmelin).
The anatomy of Farcimen (N.) croceum (Gmelin) and F. (W.)
curtum (Dall and Simpson) yields the radula formula 3:3:3:3. The
jaw is provided with a median projection, and the verge is attached
to the side of the neck, gradually tapering without a terminal ap-
pendage. It is traversed by a seminal groove.
KEY TO THE SPECIES OF SUBGENUS NEOPUPINA
J Xo-abebh seo a Os a a curtum
Axial ribs not strong.
Shelliargzesheight.more thane30 mm eee croceum
Shell small height less than’ 20) mms2 222222) Se Se2 es se=2 2 hjalmersoni
FARCIMEN (NEOPUPINA) CURTUM (Dall and Simpson)
PLATE 9, Figures 32-34 (type) ; FiacurEes 35-37
1901. Megalomastoma croceum curtum Dat and Simpson, Bull. U. 8. Fish Comm.
for 1900, vol. 1, p. 484, pl. 53, fig. 2.
Shell small, varying from elongate-ovate to cylindroconic in outline,
varying in color from wax yellow to pale brown. Nuclear whorls
decollated in all our specimens. Postnuclear whorls moderately well
rounded, marked by rather strong, closely approximated axial riblets.
Suture well impressed. Periphery well rounded. Base moderately
produced, very narrowly umbilicated, with a strong carina marking
the outer edge of the umbilicus. Aperture subcircular; peristome not
in one plane, but somewhat sinuous, simple. Operculum typically
farcimenid.
The type, U.S.N.M. No. 159678, comes from Hucares, east Puerto
Rico. It has 5.5 whorls remaining and measures: Height, 21.0 mm. ;
greater diameter, 9.5 mm.
This small species differs from the other two from Puerto Rico in
having the riblets much more strongly developed.
Additional specimens come from various stations on the eastern
slope of Puerto Rico, and since the type is a worn specimen, I am also
figuring a more perfect one, which has almost 7 whorls and measures:
Height, 23.0 mm.; greater diameter, 9.1 mm.
The animals of specimens collected by me about the roots of banana
trees at Mamey in the foothills of the Sierra Luquilla, June 23, 1929,
had the entire dorsal part of the body flesh colored with a grayish
suffusion and a pinkish flush; the sole of the foot was also flesh
colored, while the tentacles were pale orange with a grayish tinge.
There was a reddish spot present under the tentacles.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA AS
FARCIMEN (NEOPUPINA) CROCEUM (Gmelin)
PLATE 9, FIGURES 23-25
1786. Helix cylindracea glabra CHEMNITz, Neues systematisches Conchylien-
Cabinet, vol. 9, p. 166, pl. 35, fig. 1233 (nonbinomial).
1791. Helix crocea GMELIN, Systema naturae, vol. 6, p. 3655.
1798. Cerion truncatum Bouten, Museum Boltenianum, p. 90.
1822. Cyclostoma flavula Lamarck, Histoire naturelle des animaux sans
vertébres, vol. 6, p. 146.
1828. Turbo flavidus (Cyclostoma flavidum) Wood, Index testaceologicus, ed. 2,
Suppl., pp. 19, 36, pl. 6, fig. 31.
1838. Cyvlostoma crocea DesHAYES, in Lamarck’s Histoire naturelle des animaux
sans vertebres, ed. 2, vol. 8, p. 357.
1840. Megalomastoma flavula Swainson, A treatise on malacology, p. 336.
1847. Farcimen flavulum TroscuHett, Zool. Malak., vol. 4, p. 44.
1847. Megalomastoma cylindraceum PreirFER, Zool. Malak., vol. 4, p. 109.
Shell large, cylindroconic, varying in color from soiled wax colored
to bright rose colored. The nucleus is small, consisting of 2 whorls,
which are well rounded and smooth. The postnuclear whorls are
flattened, somewhat constricted a little below the suture, and marked
by closely approximated, low, rounded axial riblets, which become less
conspicuous as the whorls increase. Suture only slightly constricted.
Periphery well rounded. Base slightly protracted, narrowly
umbilicated, and marked by an obsolete fold at the outer edge of the
umbilicus. Aperture subcircular; peristome not in one plane, but
somewhat sinuous, simple, not thickened as in typical Parcimen.
Operculum typically farcimenid.
The specimen figured, U.S.N.M. No. 535945, has 6.5 whorls remain-
ing and measures: Height, 39.1 mm.; greater diameter, 14.7 mm.
This is the common large race distributed all over the north coast
of the Island of Puerto Rico.
I found the adult animal of this to have the forehead, sides, and back
brown with an olivaceous tinge, the foot flesh colored, with a bluish
tinge, tentacles tipped with orange.
The young were flesh colored, with a red spot under the tentacles.
FARCIMEN (NEOPUPINA) HJALMERSONI (Pfeiffer)
PLATE 9, FIGURES 26-28
1875. Megalomastoma hjalmersoni PrrirreR, Malak. Blitter, vol. 22, p. 119.
Shell small, elongate-ovate, varying in color from pale wax yellow
to chestnut-brown on the last turn. Nuclear whorls 2, well rounded,
smooth. The early postnuclear whorls are strongly rounded; the
rest with a constriction below the summit and marked by closely
spaced, slender, retractively slanting axial riblets, which become
obsolete on the last whorl. Suture slightly constricted. Periphery
428349424
46 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
well rounded. Base moderately protracted, narrowly umbilicated,
the umbilicus marked by a strong carina at its outer edge. Aperture
subcircular; peristome somewhat sinuous, not in one plane, simple.
Operculum typically farcimenid.
The specimen described and figured is one of a series, U.S.N.M.
No. 535692, collected by me at Lares, Puerto Rico. It has 7.1 whorls
and measures: Height, 19.7 mm.; greater diameter, 8.9 mm.
This species resembles the large #7’. (W.) erocewm (Gmelin) in the
constriction below the summit and also in the character of the rib-
bing, which is much finer than that of the other small species, F’.
(V.) curtum (Dall and Simpson), from eastern Puerto Rico.
We also have it from a number of additional stations in the west-
central portion of the Island of Puerto Rico.
FARCIMOIDES, new genus
Shells resembling those of the Cuban genus Farcimen in general
outline and thickened peristome, but with a spiral cord marking the
outer limit of the umbilicus. Operculum not simple but with the
inany layers composing it distinct, not fused as in Parcimen.
Type: Oyclostoma orbignyi Pfeifter=Farcimoides orbignyt
(Pfeiffer).
KEY TO THE HISPANIOLAN SPECIES OF FARCIMOIDES
Shell small, height less than 20 mm_--_----__--------------------------- sallei
Shell not small, height more than 25 mm.
Shell sstout-=. 222-0 sh Gees ee tA Os Ae gt domingoense
SHETTETO ty SHOU ee ee orbignyi
FARCIMOIDES SALLEI, new species
PiaTE 9, Figures 29-31
1851. Cyclostoma orbignyi va. y PFEIFFER, Proc. Zool. Soc. London, vol. 19, p. 149.
1853. Cyclostoma orbignyi variety 2 Prrrrrer, Martini-Chemnitz Conchylien
Cabinet, vol. 1, sect. 19, p. 272, pl. 37, figs. 5, 6. ’
Shell small, very elongate-ovate, covered with a wax-colored perios-
tracum. When denuded the shell may be unicolor or milk white,
or the last whorl may be darker, even purplish plum colored. Nu-
clear whorls small, well rounded, smooth. The early postnuclear
whorls inflated, strongly rounded, the last one almost flattened; the
early ones marked by slender, almost hairlike axial riblets which
gradually increase in strength until they become conspicuous, re-
tractively curved, elevated ribs, which are about as wide as or wider
than the spaces that separate them. Suture strongly constricted.
Periphery well rounded. Base rather long, openly umbilicated with
a strong cordlike carina marking the outer limit of the umbilicus.
This has a strong constriction anteriorly and posteriorly. The axial
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 47
riblets extend feebly over the umbilical wall. Aperture circular,
slightly irregular as far as its external plane is concerned; peristome
thickened and marked by concentric lamellae of about the same ex-
pansion all around. Operculum typically farcimoid.
The type, U.S.N.M. No. 535941, was collected by Dr. Abbott at
El Rio, which is between Constancia and Jarabacoa, Dominican
Republic, at an elevation of 4,000 feet. It has 7.5 whorls and
measures: Height, 19.0 mm.; greater diameter, 7.7 mm.
This I consider to be the small form described by Pfeiffer as Cyclo-
stoma orbignyi var. y. It differs markedly from the other two known
Haitian species by its much smaller size, much more inflated whorls,
and much heavier ribbing.
FARCIMOIDES DOMINGOENSE, new species
PLATE 9, Figures 1-3
Shell cylindroconic, stout, varying from wax color to pale brown
on the last turn. Nuclear whorls? The early postnuclear whorls
gain rapidly in size and are fairly well rounded; the later ones are
decidedly flattened. They are all narrowly shouldered at the sum-
mit and marked by slender, sigmoid, retractively curved axial rib-
lets, which are about as wide as the spaces that separate them on the
early turns and wider on the last. These riblets render the summit
of the turns slightly crenulated. Suture well constricted. Periph-
ery well rounded. Base moderately protracted, openly umbilicated,
and marked with a strong cord at the outer junction of the umbilicus,
which is deeply constricted at its umbilical edge and less so dorsally.
Aperture subcircular in a wavy plane; peristome thick, a little nar-
rower on the parietal wall than on the rest. Operculum, typically
farcimoid. ,
The type, U.S.N.M. No. 535940, was collected by Parker in the
Dominican Republic. It has 6.4 whorls remaining and measures:
Height, 28.8 mm.; greater diameter, 12.0 mm.
Two other lots before us are also without specific locality data.
All these are readily distinguished from F. orbignyi (Pfeiffer) by
their much more cylindric form and much stouter outline.
FARCIMOIDES ORBIGNYI (Pfeiffer)
PLATE 9, Figures 4—6
1851. Cyclostoma orbignyi PFEIFFER, Proc. Soc. London, vol. 19, p. 149.
1852. Cyclostoma orbignyi PretrrerR, Martini-Chemnitz Conchylien Cabinet, vol. 1,
sect. 19, p. 272, pl. 37, figs. 3, 4.
Shell almost cylindroconic, varying in color from unicolor to wax
yellow, with the last whorl with a reddish orange band, which is
separated from the summit and base by a paler area. Peristome
48 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
white. Nuclear whorls 2, small, well rounded, smooth. The first
2 postnuclear whorls rather strongly inflated, the rest gradually less
inflated and finally flattened. The postnuclear whorls are marked
by closely spaced, decidedly retractively curved, slender axial riblets,
which are about as wide as the spaces that separate them. Suture
rendered conspicuous by the feeble shoulder of the whorls. Periph-
ery well rounded. Base decidedly protracted, openly umbilicated,
marked by a somewhat obsolete cord at the outer limit of the umbili-
cus. The umbilical wall marked by the continuation of the axial
riblets. Aperture subcircular; peristome thickened and reflected, nar-
rower on the columellar wall than on the rest. Operculum typically
farcimoid.
The specimen figured is one of a large series, U.S.N.M. No. 151351,
from the Sallé collection, probably a topotype. It has 8.4 whorls
and measures: Height, 28.2 mm.; greater diameter, 11.5 mm. The
locality mentioned is Haiti, the designation for all Hispaniola in the
early days. We know Sallé collected only in the Santo Dominican
region, so the species must be referred to the Dominican Republic.
All our other lots are equally deficient in specific locality data.
Genus MEGALOMASTOMA (Guilding) Swainson
1840. Megalomastoma (Guilding) Swarnson, A treatise on malacology, pp. 186,
336, figs. 97 g, h, i.
Shell small, cylindroconic, covered with a persistent periostracum.
Nuclear whorls smooth, the succeeding turns marked with strongly
elevated, retractively curved axial riblets. Base with a strong spiral
keel at the outer limit of the broad open umbilicus. Aperture sub-
circular; peristome double, with a rather deep, somewhat lunate ex-
cision on the parietal wall. Operculum a simple, thin, multispiral,
chondroid plate.
Type: Megalomastoma brunnea (Guilding) Swainson=Adfegalo-
mastoma (Megalomastoma) brunneum (Guilding) Swainson.
This genus ranges from St. Thomas through St. John to Tortola.
The radula of Megalomastoma (Megalomastoma) petiti has the
formula 3:3:3:38. The jaw has a median projection, and the verge
is attached to the side of the neck, gradually tapering without a
terminal appendage. It is traversed by a seminal groove.
KEY TO THE SUBGENERA OF MEGALOMASTOMA
Last: whorl] axially ribbed =t9000.. 22 eee Megalomastoma
Last whorl'axially tuberculatedii tan ee ee ee Megalomastomoides
Subgenus MEGALOMASTOMA (Guilding) Swainson
Megalomastomas having the surface with axial ribs only, and with
the peristome excised on the parietal wall.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 49
Type: Megalomastoma brunnae (Guilding) Swainson=Megalo-
mastoma (Megalomastoma) brunneum (Guilding) Swainson.
KEY TO THE SPECIES OF SUBGENUS MEGALOMASTOMA
Shell elongate-conic.
Axial ribs of early whorls closely spaced__-----_-_-------___- antillarum
Axial ribs of early whorls not closely spaced________---_______ brunneum
Shellgelonzate-0 Va tee ee ee petiti
MEGALOMASTOMA (MEGALOMASTOMA) ANTILLARUM (Sowerby)
PLATE 9, Figures 10-12, 21
1843. Cyclostoma antillarum Sowrrsy, Thesaurus conchyliorum, vol. 1, p. 150, pl.
28, fig. 180.
Shell small, elongate-conic, chestnut-brown with the peristome
white. The nucleus consists of a single smooth turn. The early
postnuclear whorls are inflated and strongly rounded; the later ones
a little less so, marked by moderately elevated, slender, sublamellar,
retractively slanting axial ribs, which are very regular on the first
two turns and rather closely spaced; on the later turns they are much
more distantly spaced and there is a tendency for finer threads to
appear in the broad interspaces. Suture strongly constricted. Per-
iphery well rounded. Base protracted, openly umbilicated, and
marked by a moderately strong, broad, rounded cord at the outer
limit of the umbilicus, which is crossed by the axial ribs. Aperture
subcircular; peristome not on a flat plane, but sinuous, moderately
thickened and reflected, and moderately excised on the parietal wall.
Operculum typically megalomastomid.
The specimen described and figured is one of a lot, U.S.N.M. No.
393653. It was collected by me on Mount Sage, Tortola, British
West Indies. It has 7.5 whorls and measures: Height, 13.6 mm.;
greater diameter, 5.1 mm.
This species recalls closely M@, (M.) brunneum (Guilding) Swain-
son, from which it can be distinguished readily by its smaller size
and more closely spaced axial ribs on the early turns. The umbilical
cord is also less pronounced.
MEGALOMASTOMA (MEGALOMASTOMA) BRUNNEUM (Guilding) Swainson
PLate 9, FIguRES 7-9, 22
1840. Megalomastoma brunnea (Guilding MS.) Swainson, A treatise on ma-
lacology, pp. 186, 336, fig. 97, g, h, i.
Shell elongate-conic, chestnut-brown, with the peristome white.
Nuclear whorls 1.2, well rounded, smooth. The early postnuclear
whorls inflated ; the later ones less so and the last one flattened, marked
by very decidedly retractively curved, slender, sublamellar axial ribs,
50 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
which are rather distantly spaced on all the whorls. On the later
turns axial hairlines are also apparent in the intercostal spaces.
These are irregularly distributed and of irregular numbers. Suture
very strongly constricted. Periphery low, well rounded. Base de-
cidedly produced, flatly umbilicated with a decidely elevated keel at
the outer limit of the umbilicus, which is rendered crenulated by the
axial riblets that extend also over the umbilical wall. Aperture sub-
circular; peristome sinuous, that is not in a flat plane, moderately
thickened and moderately excised on the parietal wall. Operculum
typically megalomastomid.
The specimen described and figured, U.S.N.M. No. 535943, comes
from St. Thomas. It has 8.2 whorls and measures: Height, 15.4 mm. ;
greater diameter, 6.2 mm.
There are 17 additional lots of this species, all from St. Thomas, in
the collection of the United States National Museum.
This species is nearest related to M/. (If.) antillarum (Sowerby),
from which its more elongate form, more distantly spaced ribs, and
more pronounced umbilical keel will readily differentiate it.
MEGALOMASTOMA (MEGALOMASTOMA) PETITI, new species
PLATE 9, FicurEs 16-18, 20
Shell elongate-ovate, sooty chestnut-brown; peristome white.
Nuclear whorls 1.3, well rounded, smooth. The early postnuclear
whorls somewhat inflated, strongly rounded, the last one slightly flat-
tened and marked by slender sublamellar, rather distantly spaced axial
riblets, which are rather irregularly distributed. Suture well con-
stricted. Periphery well rounded. Base rather produced, flatly um-
bilicated, with a moderately strong cord at the outer limit of the
umbilicus. Aperture circular; peristome almost flat, moderately
thickened, slightly reflected and moderately excised on the parietal
wall. Operculum typically megalomastomid.
This species comes from the Island of St. John. The type, U.S.N.M.
No. 535942, was collected by me on Bordeaux Hill, Hurricane Harbor,
St. John. It has 7 whorls and measures: Height, 12.3 mm.; greater
diameter, 5.4 mm.
The United States National Museum contains six additional lots, all
from St. John.
This species is easily distinguished from the other Megalomastomas
by its shorter and much stouter shape.
MEGALOMASTOMOIDES, new subgenus
Shell resembling Megalomastoma in size, shape, and periostracal
covering but differing from it in having the peristome not excised on
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 51
the parietal wall and in having the early postnuclear sculpture as in
typical Megalomastoma, while that of the later turns has the axial
ribs broken up into elongated, somewhat lunate tubercles, which gives
to the surface of these whorls a rasplike appearance. Operculum a
simple, thin, multispiral chondroid plate.
Type: Cyclostoma (Megalomastoma) verruculosum Shuttleworth=
Megalomastoma (Megalomastomoides) verruculosum (Shuttleworth).
Distribution: Puerto Rico.
MEGALOMASTOMA (MEGALOMASTOMOIDES) VERRUCULOSUM (Shuttleworth)
PLATE 9, FiguRES 13-15, 19
1854. Cyclostoma (Megalomastoma) verruculosum SHUTTLEWORTH, Berner Mitth.,
p. 90.
1864. Megalomastoma verruculosum SoweErBy, Thesaurus conchyliorum, vol. 3, pl.
263, fig. 15.
Shell small, elongate-conic. Nuclear whorls decollated in all our
specimens. Postnuclear whorls well rounded, the early ones marked
by slender, retractively curved axial riblets, which are about as wide
as the spaces that separate them. On the last three turns these riblets
break up into segments and give to the surface a rasplike appearance.
Suture strongly constricted. Periphery well rounded. Base pro-
duced, openly flatly umbilicated, and marked by a strong spiral cord
at the outer limit of the umbilicus, which is crossed by the axial rib-
lets, which extend over the umbilical wall. Aperture circular, not m
a single plane, but sinuous; peristome thickened and somewhat re-
flected, of equal width all around, not incised on the parietal wall.
Operculum multispiral, corneous, with central nucleus.
We have this species from Puerto Rico without specific locality.
The specimen described and figured, U.S.N.M. No. 535944, has 7
whorls remaining and measures: Height, 15.1 mm.; greater diameter,
5.7 mm.
This shell is small like typical Megalomastoma but differs from it
markedly by the sculpture and the absence of excision on the parietal
wall.
Subfamily DIPLOMMATININAE Kobelt
This subfamily is defined in the key on p. 48.
Genus ADELOPOMA Doering
For definition of the genus, see p. 148.
52 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
ADELOPOMA OCCIDENTALE (Guppy)
PLATE 40, FIGuRE 6
1872. Diplommatina huttoni occidentalis Guppy, Proc. Assoc. Trinidad, vol. 2, p. 24.
1884. Diplommatina occidentale GopwIN-AUSTEN, Land and Freshwater Mollusca
of India, vol. 1, pl. 45, fig. 8.
1886. Diplommatina occidentalis GoDWIN-AUSTEN, ibid., p. 173.
1893. Diplomatina huttoni occidentale Guppy, Journ. Conch., vol. 7, p. 226.
1898. Palaina (Cylindropalaina) occidentalis KosELT and MOLLENDORFF, Nachrbl.
deutschen malak. Ges., vol. 30, p. 133.
1902. Adelopoma occidentale KozErt, Das Tierreich, vol. 16, p. 480.
Shell minute, sinistral, thin, translucent, alabaster white, except the
nuclear turns, which are buff. Nuclear whorls 1.5, inflated, strongly
rounded, smooth. Post nuclear whorls inflated, strongly rounded,
marked by slender, lamellose, scalariform, decidedly protractively
slanting axial ribs, of which 27 occur on the first and second and 33 on
the last whorl. The spaces separating the axial ribs are four or five
times as wide as the ribs, and are covered by exceedingly fine, micro-
scopic spiral striations. Suture very strongly constricted. Base in-
flated, very strongly rounded, with a narrow umbilical chink, marked
by the continuation of the axial ribs. Aperture irregularly obliquely
oval; peristome double, the inner exserted; the outer narrowly ex-
panded. Operculum typically adelopomid.
The specimen figured is one of nine, U.S.N.M. No. 203654, received
from Bland and comes from Trinidad. It has 6 whorls and meas-
ures: Height, 2.8 mm.; diameter, 1.2 mm.
U.S.N.M. No. 98784 contains six specimens from the same source,
while U.S.N.M. No. 516033 contains seven specimens received from
Dr. George Clapp.
This species differs from A. costaricense in being a little stouter
and having more axial ribs, which also are of uniform expansion
from the summit to the periphery.
Subfamily AMPHICYCLOTINAE Kobelt and Mollendorff
Cyclophorids of helicoid outline with chondroid operculum.
KEY TO THE WEST INDIAN GENERA OF THE SUBFAMILY AMPHICYCLOTINAE
Operculum: with fimbriations=29=— =. — ee Cyclohaitia
Operculum without fimbriations==—= ===". Amphicyclotulus
CYCLOHAITIA, new genus
Small amphicyclotine shells with strong spiral cords on spire and
base. Operculum multispiral, chondroid, with the outer edge of
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 53
the last whorl expanded into fimbriations. In shape and sculpture
the shells of this genus resemble Crocidopoma, but they lack the
subsutural keel and the lirations within the umbilicus; they also re-
semble it in the fimbriations of the operculum, but here we have no
calcareous lamella. The verge likewise is of amphicyclotine pattern
in its attenuation.
Type: Cyclohaitia haitia, new species.
The anatomy of Cyclohaitia shows the radula formula to be
3:3:3:3, and of the jaw is lacking the median points. The verge is
on the back of the neck behind the tentacles and traversed by a vas
deferens. The terminal appendage is long and threadlike, almost as
long as the basal portion.
CYCLOHAITIA HAITIA, new species
Puate 10, FicurEs 12-14
Shell small, helicoid, openly moderately umbilicated, covered with
an olivaceous, wax-colored periostracum. Nuclear whorls 1.5, small,
well rounded, smooth, forming a moderately elevated apex. Post-
nuclear whorls strongly rounded, circular in cross-section, marked
by incremental lines and rather strong spiral cords, of which 5 are
present on the first, 7 on the second, and 9 on the last turn between
summit and periphery in the type. These cords vary somewhat in
strength and spacing. The whorls at the summit slope decidedly
toward the suture and render this very conspicuous. Periphery
strongly rounded. Base strongly rounded, marked by 9 spiral cords
equaling those of the spire, and the continuations of the incremental
lines. The umbilical wall does not have spiral cords, only the con-
tinuation of the lines of growth. Aperture slightly oblique, cir-
cular; peristome simple. Operculum chondroid, multispiral, with
the outer edge of the later whorls much produced to form a more
or less fimbriated, up-turned extension.
The type, U.S.N.M. No. 535855, was collected north of Tiburon,
Haiti, along the road leading to Carcasse, south of the first village,
by C. R. Orcutt. It has 4.4 whorls and measures: Height, 5.2 mm.;
greater diameter, 9.0 mm.; lesser diameter, 6.7 mm. Charles R.
Orcutt also collected 10 additional lots ranging from the type locality
eastward along the south coast of Haiti to the Baie des Flamands.
Some of the dead specimens are much larger than the type. For
example U.S.N.M. No. 403722, from Damassin River has 4.6 whorls
and measures: Height, 8.7 mm.; greater diameter, 12.0 mm.; lesser
diameter, 9.0 mm., but all intermediate sizes are present in lots from
the same locality.
54 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Genus AMPHICYCLOTULUS Kobelt
1912. Amphicyclotulus Koxett, Martini-Chemnitz, Conchylien Cabinet, vol. 1,
sect. 19, pt. 3, p. 918.
Small cyclophorids with chondroid multispiral operculum, the
outer edge of which is not expanded into fimbriations. The spire is
marked by incremental lines and in the typical subgenus bears raised
spiral cords, at least on the early whorls. In this they may also be
present on the base. (In the subgenus Cycloblandia the spiral cords
are absent.) The base is broadly openly umbilicated. The peristome
is not thickened or reflected.
Type: Amphicyclotulus rufescens (Sowerby).
The genus is Antillean.
The anatomy of A. (A.) acutiliratus (Drouet) and A. (A.) mineri
shows the radula to have the formula 3:3:3:3, and the jaw without
median projection. The verge is on the back of the neck behind the
tentacles and is traversed by a vas deferens. The terminal appendage
is long and threadlike, almost as long as the basal portion.
KEY TO THE SUBGENERA OF AMPHICYCLOTULUS
Shell with*spiral lamellae2.0 te eee Amphicyclotulus
Shell) without spiral Jameliaes: Soe 2 3 es ee ee Cycloblandia
Subgenus AMPHICYCLOTULUS Kobelt
Amphicyclotulus in which all or at least the early postnuclear whorls
are marked by raised spiral cords, which may or may not be present
on the base.
Type: Amphicylotulus rufescens (Sowerby).
This subgenus is known from Puerto Rico, Guadeloupe, Dominica,
and Martinique.
KEY TO THE SPECIES OF SUBGENUS AMPHICYCLOTULUS?
Spiral sculpture confined to early postnuclear whorls.
Shell orange-yell ow = soe fe ee ee ee ea mineri
Shell ‘waxtyellowit222 2 ke Use _ Nate ie AE Os ee guadeloupensis
Spiral sculpture not confined to early postnuclear whorls.
Spiral sculpture consisting of strongly raised cords.
Spiral cords on spire strongly crenulated_________ See ee rufescens
Spiral cords on spire not strongly crenulated.
Spiral cord bounding umbilicus stronger than the rest_ acultiliratus
Spiral cord bounding umbilicus not stronger than the rest.
Spiral cords of base distantly spaced_____-__------ schrammi
Spiral cords of base not distantly spaced___-----~- dominicensis
Spiral sculpture not consisting of strongly raised cords.
Spiral sculpture consisting of strongly raised threads.
Spiral’ threads nUMerOuUss at = es eee eo liratus
Spiral ‘threads few.=]-- = 2 == — = Se eee perplexus
14, (A.) portoricensis (Boettger) is not included in this key, as I have not seen speci-
mens of it.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 55
AMPHICYCLOTULUS (AMPHICYCLOTULUS) MINERI, new species
PLATE 10, Figures 15-17
Shell rather large, helicoid, moderately broadly openly umbilicated,
covered by an orange-yellow periostracum. Nuclear whorls 1.6, weil
rounded, smooth, forming a depressed apex. Postnuclear whorls in-
flated, strongly rounded, marked by incremental lines and slender
spiral threads, which cover the anterior two-thirds of the turns, the
posterior third being without them. Of these threads 7 are present on
the second and 11 on the third whorl. On the last half of the last turn
the spiral threads are quite obsolete. Suture well impressed. Pe-
riphery inflated, strongly rounded. Base moderately rounded with
the umbilicus occupying about one-fourth of the diameter of the shell.
The base is marked by incremental lines only. Aperture subcircular,
being protracted to form an acute point at the posterior angle of the
aperture; peristome simple. Operculum typically amphicyclotulid.
The type, U.S.N.M. No. 535856, and a host of specimens come from
Laudat, Dominica. The type has 5.5 whorls and measures: Height,
13.8 mm.; greater diameter, 16.6 mm.; lesser diameter, 13.6 mm.
This species resembles most closely A. (A.) guadeloupensis, from
which its orange coloration and more elevated form readily distin-
guish it.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) GUADELOUPENSIS, new species
PLATE 10, FicurEs 18-20
Shell depressed-helicoid, moderately broadly openly umbilicated,
wax yellow. Nuclear whorls 1.5, well rounded, smooth. The post-
nuclear whorls are well rounded, the first smooth, the rest marked
by weak spiral threads which cover the posterior four-fifths of the
turn, being absent near the summit. They again become obsolete on
the last half turn. In addition to the spiral threads the whorls are
marked by fine incremental lines. Suture moderately well impressed.
Periphery inflated, well rounded. Base short, well rounded, marked
only by the incremental lines. In cross section the last whorl is in-
clined toward being oval. Aperture oblique, subcircular, slightly angu-
lated at the posterior angle; peristome simple. Operculum typically
amphicyclotulid.
The type, U.S.N.M. No. 116462, comes from Guadeloupe. It has 5.1
whorls and measures: Height, 12.8 mm.; greater diameter, 17.6 mm.;
lesser diameter, 14.0 mm.
Additional specimens in the collection of the National Museum are
also labeled merely “Guadeloupe,” without specific locality.
This species resembles most closely A. (A.) méineri from Dominica.
It is distinguished from this by its flatter form and paler coloration.
56 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
AMPHICYCLOTULUS (AMPHICYCLOTULUS) RUFESCENS (Sowerby)
PLATE 10, Ficures 4, 5
1843. Cyclostoma rufescens SowrerBy, Thesaurus conchyliorum, vol. 1, p. 94, pl. 24,
figs. 36, 37.
1843. Cyclostoma rufescens SowERBY, Proc. Zool. Soc. London, vol. 11, p. 60.
“Shell nearly orbicular, generally of a reddish colour, with a short
spire consisting of 4 rounded, spirally ridged and striated volutions;
the ridges crenulated; suture deep; aperture circular, with a thin
sharp peritreme; umbilicus large. From Martinique, communicated
by L. T. Powis, Esq.”
I have not seen specimens that satisfy Sowerby’s description and
figures which I have copied.
Specimens of A. (A.) acutiliratus (Drouet) vary considerably in
tuberculation of their spiral keels, and it is quite possible that
Sowerby’s species represents merely an exaggerated form of Drouet’s
A, (A.) acutiliratus.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) ACUTILIRATUS (Drouet)
PLATE 10, Fraures 1-3
1859. Cyclophorus acutiliratus Drovurt, Essai sur les mollusques terrestres et
fluviatiles de la Guyane francaise, p. 89, pl. 3, figs. 42-44.
Shell moderately large, openly umbilicated, varying in color from
pale yellow to bright red. Nuclear whorls 2.5, well rounded, smooth.
Postnuclear whorls strongly rounded, marked by exceedingly fine,
closely spaced, incremental lines, which give the surface a silky appear-
ance, and strong sublamellar keels, of which 5 are present between the
summit and the suture on the next to the last whorl, and 10 on the last
turn. ‘These keels are not quite regular in strength and distribution.
They average from one-half to one-fourth the width of the spaces that
separate them. Suture channeled. Periphery rounded. Base with
the umbilicus about one-fourth the greater diameter. Base well
rounded, marked by 8 spiral cords. The umbilicus is bounded by a
strong lamella and on the umbilical wall, within the umbilicus, 6 addi-
tional spiral lamellae are present. Aperture oblique, circular; peri-
stome simple, rendered somewhat fluted by the spiral sculpture. Oper-
culum typically amphicyclotulid.
The specimen described and figured, U.S.N.M. No. 585859, a female,
is one of five specimens collected by me at Colson, 14 to 15 km. north of
Fort de France, Martinique. This has 5.4 whorls and measures:
Height, 11.1 mm.; greater diameter, 15.3 mm.; lesser diameter, 12.0 mm.
U.S.N.M. No. 393879 contains seven specimens from the same lo-
cality.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA Bf
U.S.N.M. No. 151358 contains a series of specimens collected by
Sallé on Martinique. These show considerable variation. In some the
spiral cords show slight nodulation, which suggests a trend toward A.
(A.) rufescens (Sowerby) ; in fact, there is a possibility that this may
be an extreme variation of that species. The spiral keels on the um-
bilicus will easily distinguish it from all other known Amphicyclotulus.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) SCHRAMMI (Shuttleworth)
PLATE 10, FicurrEs 6-8
1857. Cyclostoma (Cyclophorus) schrammi SHUTTLEWORTH, Journ. Conchyl., vol.
5, p. 269.
Shell small, depressed-helicoid. Periostracum wax colored. Nu-
clear whorls almost 2, well rounded, smooth. Postnuclear whorls well
rounded, marked by slightly keeled spiral cords, of which 5 are present
on the second, while on the third 2 additional cords appear between the
first near the summit and the suture. At the aperture 10 spiral cords
are apparent. In addition to these spiral cords the shell is marked by
fine incremental lines. Suture moderately constricted. Periphery
well rounded. Base openly umbilicated, the umbilicus forming about
one-fourth of the greater diameter of the shell. The base is sculptured
like the spire, bearing 5 cords, while 3 additional ones are present
on the outer portion of the umbilical wall, the inner part being free
of spiral sculpture. Aperture oblique, almost circular, slightly angu-
lated at the posterior angle; peristome simple. Operculum typically
amphicyclotulid.
U.S.N.M. No. 491754 contains two specimens from Guadeloupe, one
of which we have figured. This has 4.4 whorls and measures: Height,
5.5 mm.; greater diameter, 9.8 mm.; lesser diameter, 7.6 mm.
U.S.N.M. No. 331792 contains two more specimens received from
Petit.
This species is most nearly related to A. (A.) doménicensis, from
which it is easily distinguished by its much more distantly spaced spiral
cords on the base.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) DOMINICENSIS, new species
Pate 10, Frcures 9-11
Shell small, helicoid, openly umbilicated. The nuclear and early
postnuclear turns red, the rest covered by a yellowish periostracum.
Nuclear whorls 2, well rounded, smooth. Postnuclear whorls some-
what inflated, strongly rounded, marked by strongly raised, almost
keellike spiral threads, of which 9 are present between the summit and
the suture on the next to the last whorl and 9 between the summit and
the periphery on the last whorl. The spaces that separate these cords.
58 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
which are quite regular, are a little more than twice as wide. Periphery
rendered almost angulated by a spiral cord. In addition to the spiral
cords the whorls are marked by slender incremental lines. Suture well
impressed. Base rather short, well rounded, marked by the continua-
tion of the incremental lines and 9 spiral cords, which equal those on
the spire in strength, but weaken toward the aperture. The umbilical
wall also shows traces of spiral cords. The umbilicus is about one-
fifth the greater diameter of the last whorl. Aperture oblique; peri-
stome simple. Operculum typically amphicyclotulid.
The type, U.S.N.M. No. 585857, comes from Long Dilton, Dominica.
It has 4.5 whorls and measures: Height, 5.5 mm.; greater diameter,
8.0 mm.; lesser diameter, 7.2 mm. A paratype is in the American
Museum of Natural History.
This species is nearest related to A. (A.) schrammi (Shuttleworth),
from which it can be distinguished readily by its color and the greater
number of spiral cords.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) LIRATUS (Drouet)
PLATE 11, Figures 4-6
1859. Cyclophorus liratus DRovEtT, Essai sur les mollusques terrestres et fluviatiles
de la Guyane francaise, p. 88, pl. 3, figs. 30-82.
Shell decidedly depressed-helicoid, openly umbilicated, covered by a
wax-colored periostracum. Nuclear whorls 1.3, well rounded. Post-
nuclear whorls strongly rounded, marked by rather strong, regular and
regularly spaced spiral cords, of which 14 are present between the
summit and the suture on the last turn. In addition to the spiral
sculpture, the whorls are marked by closely spaced incremental threads.
Suture well impressed. Periphery well rounded. Base moderately in-
flated, well rounded, marked by 18 spiral threads, which equal those
on the spire in strength, and by the continuation of the incremental
lines. The umbilicus is about one-fourth the width of the greater
diameter of the shell, and the umbilical wall bears a few indications of
spiral threads. A cross section of the last whorl would appear broadly
oval. Aperture slightly oblique, circular, with scarcely any angling at
the posterior angle of the aperture; peristome simple. Operculum
typically amphicyclotulid.
The specimen described and figured, U.S.N.M. No. 474037, comes
from Martinique. It has 4.4 whorls and measures: Height, 7.8 mm.;
greater diameter, 14.2 mm.; lesser diameter, 10.3 mm.
This species suggests both A. (A.) méneri and A. (A.) guadeloupen-
sts but differs from them in having the spiral sculpture extend to the
peristome and in being much more depressed.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 59
AMPHICYCLOTULUS (AMPHICYCLOTULUS) PERPLEXUS, new species
PLATE 11, FicurEs 7-9
Shell small, depressed-helicoid, covered by a wax-yellow periostra-
cum, which assumes a rosy tinge toward the apex. Nuclear whorls
almost 2, well rounded, smooth. Postnuclear whorls inflated, strongly
rounded, marked by slender incremental lines and on the next to the
last turn by six low, rounded spiral threads, of which the one at the
summit is weaker than the rest. On the last whorl these threads be-
come obsolete and almost vanish before reaching the peristome. Su-
ture moderately impressed. Periphery well rounded. Base well
rounded, openly umbilicated, the umbilicus being about one-third the
width of the greater diameter. The base is marked by incremental
lines only. Aperture slightly oblique, circular, with a slight angle
at the posterior angle; peristome simple. Operculum typically
amphicyclotulid.
The type, U.S.N.M. No. 535858, comes from Guadeloupe. It has
4.9 whorls and measures: Height, 7.0 mm.; greater diameter, 11.5
mm.; lesser diameter, 8.0 mm.
U.S.N.M No. 43485 contains another specimen from the same lo-
cality.
U.S.N.M. No. 151348a contains two additional specimens collected
by Sallé on Guadeloupe.
The weak sculpture of this species will differentiate it from the
rest of the species.
AMPHICYCLOTULUS (AMPHICYCLOTULUS) PORTORICENSIS (Boettger)
PLATE 11, Ficurres 19-21
1887. Cyclotus portoricensis Borrrcrr, Jahrb. deutschen malak. Ges., vol. 14,
pp. 101-102, pl. 4, fig. 7.
Shell broadly openly umbilicated, umbilicus equaling a fifth part
of the width of the base, depressed-turbinate, solid, olivaceous, a little
lighter on the base, shining, spire little expanded, convex; apex quite
small, somewhat obtuse, after decortication, red. Whorls, 5-5.5, in-
creasing regularly in size, with impressed disjunct suture, convex,
minutely rugulose striate and everywhere densely spirally lirate, lirae
13-15 in the penultimate whorl, last whorl terete, base more convex,
a little expanded at the aperture or slightly descending. Aperture
moderately large, oblique; peristome continuous, straight, acute,
white, with the margin somewhat thickened all around; columella a
little retracted and reflected. Operculum immersed, corneous with
11 turns, its nucleus concave.
60 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Height, 13-13.5 mm.; diameter, 19.5-20.5 mm.; height of aperture,
9.5-10.5 mm.; diameter, 8.5-9.5 mm. Habitat, Rio Blanco, in the
Sierra de Caduros, Puerto Rico.
Boettger says he had before him seven specimens sent to him by
Otto Goldtuss.
I have not seen this species and have freely translated Dr. Boett-
ger’s description and copied his figures.
It is related to A. (A.) guadeloupensis, A. (A.) mineri, and A. (A.)
liratus (Drouet). The strong lirations of the base will distinguish
it from the first two; it is easily differentiated from the last by its
much larger size.
CYCLOBLANDIA, new subgenus
Amphicyclotulus in which the whorls, even the early postnuclear
turns, are without raised spiral cords or threads.
Type: Cyclostoma beauiana Petit= Amphicyclotulus (Cycloblandia)
beauianus (Petit).
We know this subgenus from Guadeloupe, Grande Terre, Dominica,
and Martinique.
The anatomy of A. (Cycloblandia) amethystinus (Guppy) shows
the radula to have the formula 3:3:3:3, and the jaw without median
projection. The verge is on the back of the neck behind the tentacles,
and traversed by a vas deferens. The terminal appendage is long and
threadlike, almost as long as the basal portion.
KEY TO THE SPECIES OF SUBGENUS CYCLOBLANDIA
Shell large, greater diameter more than 15 mm__________________ amethystinus
Shell small, greater diameter less than 12 mm_____-___-______-_- beauianus
AMPHICYCLOTULUS (CYCLOBLANDIA) AMETHYSTINUS (Guppy)
PLATE 11, FIGURES 1-3
1868. Cyclotus amethystinus Guppy, Ann. Mag. Nat. Hist., ser. 4, vol. 1, p. 433.
Shell large, depressed-helicoid, covered by a pale olive periostracum.
Nuclear whorls 1.8, well rounded, smooth. Postnuclear whorls inflated,
strongly rounded, marked by fine incremental lines, which vary some-
what in strength and by inconspicuous hair lines a little darker than
the general coloration of the shell. In cross section the whorls are
rather inclined to oval than circular. Suture well impressed, particu-
larly so on the early turns. Periphery strongly rounded. Base not
inflated, well rounded, marked by incremental lines equaling those on
the spire, which on the last portion of the last whorl become intensi-
fied. Aperture oblique, almost circular; peristome simple. Operculum
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 61
with about 8 whorls, thin, horny, with the outer edge of the turns
slightly upturned and the exposed portion retractively striate.
The specimen described and figured, U.S.N.M. No. 478947, is one of
several hundred which I collected at Danes on the east side of the
watershed east of Portsmouth, Dominica, August 4, 1928. It has 5.0
whorls and measures: Height, 9.8 mm.; greater diameter, 15.2 mm.;
lesser diameter, 12.0 mm. The larger series is registered as U.S.N.M.
No. 473946.
My notes say of the animal: “Entire surface flesh-colored with a
roseate flush, a pale rose-colored space runs across the snout in front
of the tentacles, which are bright orange. The sole of the foot of the
same coloration as the dorsal part. Found on the ground under decay-
ing rubbish.”
AMPHICYCLOTULUS (CYCLOBLANDIA) BEAUIANUS (Petit)
PLATE 11, Ficurres 10-12
1853. Cyclostoma beauiana Petit, Journ. Conchyl., vol. 4, p. 368, pl. 11, figs. 11, 12.
1854. Cyclostoma inornata RepFteLD, Ann. Lyc. Nat. Hist. New York, vol. 6, p. 18,
pl. 1, fig. 7.
Shell small, depressed-helicoid, openly umbilicated, olivaceous horn
colored. Nuclear whorls 1.6, well rounded, smooth. Postnuclear whorls
strongly rounded, marked by fine incremental lines only. The summit
of the whorls is slightly flattened near the suture, which renders this
fairly conspicuous. Periphery well rounded. Base inflated, strongly
rounded, marked like the spire. Aperture oblique, almost circular,
slightly angulated at the posterior angle; peristome simple. Interior
of the aperture reddish orange within, paling toward the edge. Oper-
culum thin, horny, multispiral.
The specimen described and figured, U.S.N.M. No. 535844, is one of
five collected by me at Traicou Chet, Crois Rivier, Guadeloupe, July 31,
1928. It has 4.4 whorls and measures: Height, 7.7 mm.; greater
diameter, 11.0 mm.; lesser diameter, 8.2 mm.
I collected two additional specimens, U.S.N.M. No. 390055, at Habi-
tation La Tapick, Guadeloupe. Two specimens, U.S.N.M. No. 151348,
were collected by Sallé with the locality Guadeloupe. A dead specimen,
U.S.N.M. No. 390085, was obtained at Lepes, Grande Terre, July 30,
1928; and six more, U.S.N.M. No. 393118, were collected at Grande
Ravine, Grande Terre.
These latter specimens are not distinguishable from those taken on
Guadeloupe. Petit’s species was described from Grande Terre and
Redfield’s from Guadeloupe.
428349—42-_5
62 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Subfamily APEROSTOMINAE Torre and Bartsch
This subfamily is defined on p. 38.
KEY TO THE GENERA OF WEST INDIAN APEROSTOMINAE
Basal chondroid plate produced into fimbriations.
Spire with spiral keels,or cords_______._-.—-----______-_--- Crocidopoma
Spire without spiral keels or cords.
Peristome with a breathing notch_---_---___--_____--_-_- Cyclojamaicia
Peristome without a breathing notch__---_---_--__--- Cyclovendreysia
Basal chondroid plate not produced into fimbriations.
Operculum with a strongly elevated calcareous lamella.
Lamelia incurved! throughouts22222 2 Cyclopilsbrya
Lamella not incurved throughout.
Lamella expanded at free edge-------------------- Ptychocochlis
Lamella not expanded at free edge_--__-__--_------------- Poteria
Operculum without a strongly elevated calcareous lamella____ Aperostoma
Genus CROCIDOPOMA Shuttleworth
For definition of Crocidopoma see p. 39.
Our knowledge of the anatomy of Crocidopoma (Crocidopoma)
wortex elevatum is based upon a very poorly preserved specimen of
the animal of this species. It shows a radula having the formula
3: 3: 3: 2. The male reproductive system resembles that of
Amphicyclotulus.
A key to the subgenera of Crocidopoma will be found on p. 39.
Subgenus CRocIpOPOMA Shuttleworth
1857. Crocidopoma SHUTTLEWORTH, Journ. Conchyl., vol. 5, pp. 271-272.
1891. Crocidopoma Crosss, Journ. Conchyl., vol. 39, p. 160.
In this subgenus the flocculations of the operculum extend very
greatly beyond the edge of the calcareous lamella.
Type: Cyclostoma (Cyclotus) flocedsum Shuttleworth=Crocido-
poma (Crocidopoma) floccosum (Shuttleworth).
This subgenus is confined to Hispaniola.
KEY TO THE SPECIES OF THE SUBGENUS CROCIDOPOMA ?
Subsutural ‘Spiral* keel ‘very ‘strong: —2 eS soi ee eee vortex
Subsutural spiral keel not very strong.
Suture very broadly channeled.
Shell planorboid2 22 "2222 eee ee casuelense
Shell’ depressed-helicoid=2_- == 24 ee milleri
Suture not very broadly channeled.
Shell helicoid {2222 eee ee __ orcutti
Shell depressed-helicoid_.._—_.- _- .. =. = SE ee eee ee abbotti
20. (C.) floccosum (Shuttleworth) is not included in this key for lack of material for
differential comparison with material at hand, none of our specimens satisfying the
description thereof.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 63
CROCIDOPOMA (CROCIDOPOMA) VORTEX (Weinland)
Shell small, almost lenticular with a thin yellowish-brown perio-
stracum, which is usually worn on the keels, exposing them as white
lines. The denuded shell is white. Nuclear whorls 1.5, strongly
rounded, smooth. The postnuclear whorls are well rounded and
marked by elevated spiral keels, of which 6 are present between the
summit and the suture on the first turn and 7 on the remaining whorls.
On the last half of the last turn there are usually intercalated threads.
Periphery well rounded. Base well rounded and marked by 5 strong
keels between the periphery and the edge of the umbilicus, 5 more
being present on the umbilical wall. The umbilicus is open and the
whorls can be seen within. Aperture very slightly oblique, circular ;
peristome rendered sinuous by the spiral cords. Operculum typically
crocidopomid.
The species is restricted to the southwestern peninsula.
We are recognizing two subspecies, which the following key will
help to differentiate :
KEY TO THE SUBSPECIES OF CROCIDOPOMA VORTEX
ShellMilenticnilars Wee ae eee ee oe ee er ae ree eee ee are ee vortex
Shell depressed-helicoid 2204. Sh ee) See ee ee eee elevatum
CROCIDOPOMA (CROCIDOPOMA) VORTEX VORTEX (Weinland)
PLATE 11, Figures 13-15
1862. Cyclostoma vortex WEINLAND, Malak. Blitter, vol. 9, p. 90.
1869. Cyclotus vortex PFEIFFER, Noy. Conch., vol. 3, p. 445, pl. 98, figs. 17-20.
Shell small, almost lenticular, with a pale yellowish-brown peri-
ostracum, which when worn from the strong spiral keels leaves the
white shell below exposed. Nuclear whorls 1.5, strongly rounded,
smooth. Postnuclear whorls well rounded, marked by decidedly ele-
vated spiral keels, of which 6 are present between summit and suture
on the first turn, and 7 on the remaining turns. On the last half
of the last turn intercalated finer threads appear between the heavy
keels. In addition to the spiral keels the shell is marked by rather
strong incremental lines. The space between the very strongly ele-
vated keel near the summit and the summit of the whorl makes the
suture a deeply impressed channel. Periphery well rounded. Base
well rounded, marked like the spire, 5 strong keels being present
between the periphery and the edge of the umbilicus and 5 more on
the umbilical wall. The umbilicus is broad and extends to the very
first whorl. The last turn is solute for about one-twentieth of a
turn. Aperture very slightly oblique, circular; peristome ren-
dered slightly sinuous by the spiral cords and somewhat notched by
64 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
the first spiral cord below the suture, which undoubtedly when closed
by the operculum leaves a breathing space. Operculum typically
crocidopomid.
This subspecies has a rather wide range along the coast of the
southern peninsula. We have many specimens, from many stations,
ranging from Port-au-Prince around the end of the island, down
the south coast to near Aquin, to some of the outlying small islands
of the region. Throughout this range the shells are fairly constant,
differing, however, materially in size in the individual colonies and
also in the amount of solution with the last turn. In an exceedingly
large old individual this may extend over considerably more space
than in those not quite so big. Specimens collected by Henderson and
Simpson at La Ferrier, Haiti, U.S.N.M. Nos. 162982 and 490072, and
by W. L. Abbott at Sostia, Dominican Republic, U.S.N.M. Nos.
336766 and 336788, both near the north coast of Hispaniola, are indis-
tinguishable from those of the south peninsula.
The specimen that has served for my description and figure,
U.S.N.M. No. 403073, comes from the type locality, a little west of
Jérémie. It has 4.1 whorls and measures: Height, 6.7 mm.; greater
diameter, 11.8 mm.; lesser diameter, 9.0 mm.
A very large specimen from Anse du Clerc, Dept. du Sud, U.S.N.M.
No. 401741, has 4.6 whorls and measures: Height, 9.9 mm.; greater
diameter, 14.2 mm.; lesser diameter, 9.8 mm.
CROCIDOPOMA (CROCIDOPOMA) VORTEX ELEVATUM, new subspecies
PLATE 11, FicuRes 16-18
This subspecies agrees completely in its sculpture with the typical
form. It differs materially from it, however, in being much more
elevated and in having the keel at the summit much more closely ap-
proximated to the preceding turn, thus making the channeled suture
much narrower. The specimens were collected on top of Mount
Rochelois.
The type, U.S.N.M. No. 535846, was collected by W. J. Eyerdam and
measures: Height, 8.9 mm.; greater diameter, 12.4 mm.; lesser di-
ameter, 9.9 mm.
CROCIDOPOMA (CROCIDOPOMA) FLOCCOSUM (Shuttleworth)
PLATE 12, FIGURE 16
1857. Cyclostoma (Cyclotus) floccosum SHUTTLEWORTH, Journ Conchyl., vol. 5,
pp. 268, 272.
1863. Oyclotus floccosus REEVE, Conchologia iconica, vol. 14, sp. 48.
Shell broadly umbilicated, depressed, thin, with incremental hair
lines and closely spaced, slightly nodulose, spiral keels, which alternate
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 65
in strength, whitish with a thin soiled-yellow periostracum. The apex
does not project materially and is rather obtuse. Suture deeply
channeled. Whorls 4.5, convex, the last descending, keeled above and
solute. Aperture scarcely oblique, circular; peristome simple and
acute. Operculum typical. Height, 7.0 mm.; greater diameter,
12.0 mm.; lesser diameter, 9.0 mm. Aperture, 4 mm. broad and wide.
The above is a translation of Shuttleworth’s description to which
he adds, “Habitat, Haiti Sallé.”
I have mapped Sallé’s travels in the island of Hispaniola. They
did not extend into the range of what I am here calling C. (C.) vortex
(Weinland), which occupies the western portion of the south penin-
sula, and which has been at times confused with the present species.
QC. (C.) floccosum (Shuttleworth) is from Santo Domingo. The
United States National Museum collection contains other species from
Santo Domingo, but it does not have representatives of C. (C.) floc-
cosum. For that reason I am copying Reeve’s figure.
CROCIDOPOMA (CROCIDOPOMA) CASUELENSE Crosse
Puate 41, FiguREs 7-9
1891. Crocidopoma casuelense Crosse, Journ. Conchyl., vol. 39, p. 161.
1902. Crocidopoma casuelense Kozett, Das Tierreich, Cyclophoridae, pp. 259-260,
fig. 54.
Shell planorbid, covered with a thin yellow periostracum (nuclear
whorls decollated). The postnuclear whorls are strongly rounded
and marked by strong spiral keels, of which 7 are present on the first
remaining turn between the summit and the periphery, and 12 on
the same space on the last turn. The first keel at the suture is only
a trifle heavier than the remaining ones. The spaces between these
keels are broader than the keels. The axial sculpture consists of fine
hair lines, and the last whorl is solute for about one-tenth of a turn.
Suture very deeply impressed. Periphery well rounded. Base very
broadly openly umbilicated and marked by 6 spiral keels equaling
in strength those on the spire. There are 7 additional spiral keels
of equal strength and spacing on the umbilical wall. On the base
and umbilical wall the keels are narrower than the spaces that separate
them. Aperture subcircular; peristome slightly reflected, thin.
Operculum typically crocidopomid.
The specimen described and figured, U.S.N.M. No. 331865, is one
of the Redfield collection collected by Cuming. It has 3 postnuclear
whorls remaining and measures: Height, 5.2 mm.; greater diameter,
10.5 mm.; lesser diameter, 7.7 mm. I have been unable to locate the
exact position of Casuela in Hispaniola.
66 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
CROCIDOPOMA (CROCIDOPOMA) MILLERI, new species
PLATE 12, Figures 1-3
Shell helicoid, covered with a pale brown periostracum, or at least
one would be led to believe so by the fragments clinging to the speci-
men at hand. Nuclear whorls about 1.6, well rounded, smooth. Post-
nuclear whorls inflated, very strongly rounded, marked by rather
slender, rather broad, spiral cords, of which 11 are present on the
last two turns between summit and suture. These are separated by
spaces only a little wider than the cords. The one at the summit is
broader than the rest and the space between the summit and this is
deeply inbent to form a wide channel at the suture. Periphery
strongly rounded. Base inflated, strongly rounded, marked by 10
spiral cords which equal those of the spire in strength and spacing.
Umbilicus moderately broad, the umbilical wall of the last whorl
bears 6 spiral cords. Aperture circular; peristome rendered slightly
sinuous by the spiral cords, slightly notched at the heavy cord near
the summit. Operculum?
The type, U.S.N.M. No. 389796, has 3.7 whorls remaining and
measures: Height, 7.0 mm.; greater diameter, 9.9 mm.; lesser diame-
ter, 7.4 mm. It was collected by Gerrit S. Miller on the slope of
Loma de Cielo, Bohoruco Mountains, at an altitude of 3,000 feet,
Polo, Barahona District, Dominican Republic.
CROCIDOPOMA (CROCIDOPOMA) ORCUTTI, new species
PLATE 12, FIGURES 36-38
Shell large for the genus, depressed-helicoid, covered by a pale yel-
lowish periostracum. Nuclear whorls 1.5, well rounded, smooth.
Postnuclear whorls inflated, strongly rounded, marked by moderately
elevated spiral cords, of which 8 are present on the first, 10 on the sec-
ond, and 16 on the last part of the last turn between summit and peri-
phery. These cords increase in number by intercalation and therefore
vary considerably in strength. The first one near the summit, how-
ever, is always stronger than the rest. In addition to this the shell is
marked by rather strong incremental lines, which are more or less
hairlike and give to the spiral cords an almost granular aspect. Suture
profoundly narrowly channeled. Periphery well rounded. Base
strongly rounded, moderately broadly umbilicated, marked by 9 spiral
cords between which finer threads are present at the termination of the
last whorl. The umbilical wall bears 6 spiral cords with an inter-
calated one between the outer heavy keels. Last whorl solute for
about one-twentieth of a turn. Aperture slightly oblique, almost cir-
cular, with a slight notch formed by the subsutural keel; peristome
smooth. Operculum typically crocidopomid.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 67
This species was collected by Charles R. Orcutt on Gimbi Mountain
in the southwestern part of the southern peninsula of Haiti.
The type, U.S.N.M. No. 535847, has 4.5 whorls and measures:
Height, 11 mm.; greater diameter, 14 mm.; lesser diameter, 10.8 mm.
CROCIDOPOMA (CROCIDOPOMA) ABBOTTI, new species
PLATE 12, Figures 13-15
Shell almost lenticular, white (it has lost its periostracum). Nuclear
whorls? Postnuclear whorls strongly rounded, marked by slender,
well elevated, spiral keels, of which 6 are present between summit and
suture on the first, 8 on the second, and 9 on the last whorl. These
keels are only about one-fourth as wide as the spaces that separate
them. The first one near the summit is much stronger and broader
than the rest and is notched at the peristome to furnish a breathing
space. Suture narrowly and profoundly channeled. Periphery well
rounded. Base strongly rounded, marked by 6 slender spiral keels.
The umbilicus is very broad and profound. The wall on the last whorl
bears 5 spiral keels. The last whorl is solute for about one-thirtieth
of aturn. Aperture circular, very slightly oblique; peristome simple,
except that it is rendered wavy by the spiral cords. Operculum?
The type, U.S.N.M. No. 356197, a broken specimen, has lost the
nuclear whorls and part of the last turn. The 2.3 whorls remaining
measure: Height, 6.3 mm.; greater diameter, 10.5 mm.; lesser diameter,
7.4mm. This was collected by Dr. Abbott at “Cave, Savanna, 1 hour
west of Maniel Viejo, Bohoruco Mts.,” southwest Dominican Republic ;
altitude, 2,510 feet.
A young specimen, U.S.N.M. No. 389873, was collected by Gerrit 8S.
Miller at Polo, District Barahona, Bohoruco Mountains, altitude 2,000
feet, Dominican Republic.
While I dislike to base a species on an imperfect specimen, for the
sake of completeness I am here doing so.
CYCLOJAMAICIA, new genus
Medium-sized aperostomids of almost lenticular form, with deeply
channeled suture and a decided narrow notch in the peristome at the
posterior angle of the aperture, which leaves on the preceding parts of
the whorls at this part of the shell a rough raised cord. Operculum
with a multispiral, strongly elevated, obliquely outward curved cal-
careous lamella between the turns of which project thin scalelike fimbri-
ations, the outer extension of the basal chondroid elements; on the in-
ner turns these are worn off to the length of the calcerous lamella,
while on the outer margin in perfect specimens they project to fully
triple the width of the calcareous lamella as partly overlapping, obli-
68 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
quely placed shingles. These fimbriations therefore are not merely a
cutting up of the expanded basal chondroid plate, but they represent in
a measure a decidedly folded and cut feature. These fimbriations on
the outer whorl are also somewhat fluted. They are as thin as tissue
paper and unless very carefully handled might easily be destroyed.
Type: Cyclostoma suturale Sowerby=Cyclojamaicia suturalis
(Sowerby).
The genus is confined to Jamaica.
The radulae of Cyclojamaicia bondi (Vanatta) and C. suturalis
(Sowerby) show the formula 3:3:3:3. The jaw of C. bondi is with-
out median projection. The verge of both is situated on the back of
the neck behind the tentacles and is swollen basally. It is traversed
by a seminal groove only. The terminal appendage is very short and
appears almost chitinized and is shaped like a single turn of the tip
of a wood screw.
KEY TO THE SPECIES OF GENUS OCYCLOJAMAICIA
Suturall channeli wid@ee ee fee ae nace Bene Oe ee ee bondi
Sutural-channel narrows se ae ee suturalis
CYCLOJAMAICIA BONDI (Vanatta)
PLATE 12, Fiaures 7-9
1936. Poteria (Crocidopoma) bondi VAnatTra, Nautilus, vol. 49, pp. 98-99.
Shell lenticular, very broadly umbilicated, covered with a brownish
wax-colored periostracum. Nuclear whorls about 1. Postnuclear whorls
strongly rounded, the first without a keel at the summit, the rest with
the strongly rounded keel at some distance from the preceding turn,
which leaves a broadly channeled suture between that and the keel.
The postnuclear whorls are marked by very closely spaced, hairlike,
sublamellar axial riblets. In crossing the keel at the summit these
become decidedly backward bent. Periphery well rounded. Base well
rounded, marked like the spire. Aperture circular with a notch at the
posterior keel as in Cyclojamaicia suturalis (Sowerby). Operculum
as described in the generic diagnosis.
The specimen figured is one of a series, U.S.N.M. No. 399392, collected
by Charles R. Orcutt at Retreat, 1 mile west of Great House, St. Ann
Parish, Jamaica. It has 4.2 whorls and measures: Height, 6.1 mm.;
greater diameter, 13.0 mm.; lesser diameter, 9.5 mm.
We have 22 lots of this species in the collection of the U. S. National
Museum, which range from Stewart Town in a curve east to Moneague
in St. Ann Parish.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 69
CYCLOJAMAICIA SUTURALIS (Sowerby)
PLATE 12, Figures 10-12
1843. Cyclostoma suturale Sowrrsy, Proc. Zool. Soc. London, vol. 11, p. 29.
1848. Cyclostoma suturale G. B. Sowersy, Thesaurus Conchyliorum, vol. 1,
p. 91, pl. 23, figs. 1-2.
Shell lenticular, very widely umbilicated, covered with an oliva-
ceous, wax-colored periostracum. Nuclear whorls about 1, well
rounded, smooth. Postnuclear whorls strongly rounded, the first one
with a normal suture, the rest with a decidedly elevated keel, sep-
arated from the preceding turn by a very deeply channeled suture.
The keel in fact is a backward reflection of succeeding stages of
the edge of the peristome, near the posterior angle of the aperture,
where there exists a sinus, which forms a notch in the peristome,
which serves as a breathing pore when the operculum closes the
aperture. The postnuclear whorls are marked by very closely spaced,
hairlike, sublamellar axial riblets. These in crossing the keel at the
summit become decidedly backward bent. Periphery well rounded.
Base well rounded, marked like the spire. Aperture circular, except-
ing the sinus at the posterior angle referred to above. Peristome
simple. Operculum as described in the generic diagnosis.
The specimen described and figured, U.S.N.M. No. 535685, is one
of four collected by Dr. H. Burrington Baker at his station J22,
that is, the hill north of Rotten Gut, Manchester Parish, Jamaica.
It has 4.5 whorls and measures: Height, 7.0 mm.; greater diameter,
14.4 mm.; lesser diameter, 10.6 mm.
We have 23 additional lots of this species, all of which come from
the edge of the Cockpit country, from the northeastern part of
St. Elizabeth to the northwestern part of Manchester Parish and
the extreme southern part of Trelawny Parish. All this material
comes from within a radius of 6 miles.
This species is easily distinguished from Cyclojamaicia bondi
(Vanatta) by its much narrower channeled suture between the sum-
mit and the keel. Furthermore, the keel at the summit in sutwralis
is acute, while in C. bondi it is rounded.
Sowerby, in describing this species, said that G. C. Bainbridge,
of Liverpool, had received a few specimens many years ago from
the woods of Demerara. This is undoubtedly a mistake, for nothing
like it has ever been reported from the mainland.
CYCLOVENDREYSIA, new genus
Small, planorboid, widely umbilicated aperostomids, the whorls
of which are circular in cross section, the sculpture consisting of
incremental threads only. Operculum with a multispiral calcareous
lamella, which is outward reflected to touch the succeeding turn at
70 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
the outer edge, leaving a tear-drop-shaped space separating them
basally. This lamella bears strong oblique threads. The dorsal
chondroid plate is projected and upward turned at the outer edge
of the last whorl to the height of the calcareous lamella or slightly
above it. The upturned edge is slightly fimbriated.
Type: Cyclostoma dubiosum C. B. Adams= Cyclovendreysia
dubiosa (C. B. Adams).
We have been unable so far to find a male of this species. The
radula has the formula 3:3:3:3, and the jaw lacks distinct median
projections.
CYCLOVENDREYSIA DUBIOSA (C. B. Adams)
PuatTe 12, Figures 46
1851. Cyclostoma dubiosum C. B. ApAMs, Ann. Lyc. Nat. Hist. New York, vol. 5,
p. 81.
Shell lenticular, very widely openly umbilicated, covered with a
brownish wax-colored periostracum. There is one nuclear turn,
which is rounded and smooth. Postnuclear whorls inflated, strongly
rounded, separated by a profoundly impressed suture and marked
by slender, almost vertical, closely placed, hairlike axial riblets, which
are a little more distantly spaced on the first turn than on the re-
maining turns. Periphery strongly rounded. Base very broadly
openly umbilicated, well rounded, marked like the spire, which is
also the sculpture of the umbilical wall. In cross section the whorls
are circular. Aperture slightly oblique, circular; peristome simple.
Operculum described in our generic diagnosis.
The specimen figured, U.S.N.M. No. 356120, is one of several col-
lected by John B. Henderson at Ipswich, St. Elizabeth Parish,
Jamaica. It has 4.5 whorls and measures: Height, 6.5 mm.; greater
diameter, 14.5 mm.; lesser diameter, 11.0 mm.
The United States National Museum contains 22 lots, and in addi-
tion to this I have seen two lots from the C. B. Adams collection
and four from the Academy of Natural Sciences of Philadelphia.
The material before me enables me to say that the species ranges
from the northwestern corner of St. Elizabeth Parish northward to
Montego Bay, St. James Parish, through Trelawny to Browns
Town, St. Ann Parish. Throughout this range it shows no differen-
tiation into races.
While the shell in general form resembles Cyclojamaicia suturalis
(Sowerby) and @. bondi (Vanatta), its entirely different operculum
and absence of sutural keel remove it at once from that complex.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA Tal
CYCLOPILSBRYA, new genus
Medium-sized aperostomine mollusks having a strongly elevated
spiral lamella on the outside of the whorls of the operculum, which
is concavely curved on the inside.
Type: Cyclostoma jugosum C. B. Adams=Cyclopilsbrya ( Cyclo-
pilsbrya) jugosa (C. B. Adams).
The anatomy of @. (C.) caribaea (Clench and Aguayo) shows the
radula formula 3:3:3:3.
The genus presents two primary modifications, which may be
defined as subgenera, as follows.
KEY TO THE SUBGENERA OF GENUS CYCLOPILSBRYA
Lamella curving uniformly inward to a sharp edge-----------~- Cyclopilsbrya
Lamella not curving uniformly inward to a sharp edge, but with a
broadly expanded fold, which extends from the outside of the free
edge outward and downward almost to meet the next turn___ Cyclocaymania
CYCLOPILSBRYA, new subgenus
In this subgenus the strongly raised spiral lamella is curved inward
and its free outer margin is sharp.
Type: Cyclostoma jugosum C. B. Adams=Cycelopilsbrya (Cyclo-
pilsbrya) jugosa (C. B. Adams).
This subgenus is confined to western Jamaica, where it ranges
through the parishes of Hanover, St. James, Westmoreland, and St.
Elizabeth.
KEY TO THE SPECIES OF SUBGENUS CYCLOPILSBRYA *
Umbilical keel well developed.
Wrinkled sculpture of base strong.
Shell large, greater diameter more than 20 mm-____ westmorelandensis
Shell small, greater diameter less than 15 mm_------~---~ hendersoni
Wrinkled sculpture of base feeble.
Pits outside of umbilical keel profound__-~~---------- glenburniensis
Pits outside of umbilical keel not profound___---_-~-~----- rupisfontis
Umbilical keel obsolete.
Umbilicus very broad.
Retractively slanting axial ribs of base very regular____--___-- striosa
Retractively slanting axial ribs of base very irregular_______~~ jugosa
Umbilicus not very broad.
Shell large, greater diameter more than 24 mm-_----_----__-- caribaea
Shell small, greater diameter less than 18 mm.
Sculpture coarsely wrinkled___-~---------------------- rufilabris
Sculpture not coarsely wrinkled_--~----~~-------~------ asperula
31 have not seen C. (C.) beswicki (Chitty) and therefore have not included it in this key.
72 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
CYCLOPILSBRYA (CYCLOPILSBRYA) WESTMORELANDENSIS (Chitty)
PLATE 12, Ficures 30-82
1857. Cyclotus westmorelandensis Cuirry, Proc. Zool. Soc. London, vol. 25,
. 156.
1898. ease (Plectocyclotus) westmorelandensis KoBELT and MOLLENDORFF,
Nachrbl. deutschen malak. Ges., vol. 29, p. 139, reprint.
1912. Neocyclotus (Plectocyclotus) varians thielei Kosert, Martini-Chemnitz
Conchylien Cabinet, vol. 1, sec. 19, pt. 3, p. 944, pl. 141, figs. 10-12.
Shell large, helicoid, the early whorls red, the last one pale brown
with the appressed summit almost white. The nucleus consists of
1.5 well-rounded smooth turns. The rest of the whorls are marked
by rather strong axial ribs, which are about as wide as the spaces
that separate them. They are very regular on the early turns but
become less so on the last whorl owing to the additional wrinkled
sculpture. Beginning with the fourth turn, rather large, feebly de-
veloped, and irregularly distributed wrinkles, which fade out on the
last part of the last turn, make their appearance. These wrinkles
are best emphasized below the appressed summit. Suture strongly
impressed on all but the last turn where the summit creeps up on
the preceding turn and forms a cord. Periphery well rounded and
marked by the wrinkling characteristic of the upper surface of the
last turn. Base inflated, strongly rounded, moderately broadly,
openly umbilicated and marked by a series of very strong, oblique,
retractively slanting, short cords, which radiate from the outer edge of
the umbilicus, but do not reach the middle of the base. The spaces
separating these cords are not quite as wide as the cords. The rest of
the base is feebly malleated. The umbilical wall is marked by obsolete
ribs, which are the continuation of the strong oblique cords mentioned
above. These, as well as their interspaces, are crossed by incremental
lines. Aperture circular, oblique, decidedly protracted into a clawlike
element at the posterior angle; the outer lip of the peristome is exca-
vated below the summit and protracted on the middle, thin at the edge,
with the inner lip thickened. Operculum typically cyclopilsbryid.
The specimen figured is one of two, U.S.N.M. No. 356099, received
from C. B. Adams. It has 5.5 whorls and measures: Height, 19.2
mm.; greater diameter, 23.8 mm.; lesser diameter, 17.3 mm. It is
without specific locality label.
The species appears confined to the south side of Westmoreland
Parish, Jamaica, east of Savanna la Mar.
CYCLOPILSBRYA (CYCLOPILSBRYA) HENDERSONI, new species
PLATE 12, FiacuREs 21-23
Shell very small, helicoid. Nuclear whorls pale; the succeeding
whorls reddish; the last one yellowish, covered with a thin yellowish
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 73
periostracum. The nucleus consists of 1.5 well-rounded smooth turns.
The postnuclear whorls are inflated, strongly rounded, and marked
by rather irregular wrinkled axial riblets. In addition to this, be-
ginning with the second half of the first postnuclear whorl, wrinkles
irregular in size and distribution make their appearance. These are
rather coarse, particularly so on the last whorl. Suture well im-
pressed, a little less so on the last part of the last whorl. Periphery
well rounded and marked by the continuation of the wrinkles. Base
inflated, strongly rounded, openly umbilicated with a low keel marking
the outer edge of the umbilicus. The entire base is marked by very
strong wrinkles, which almost assume the form of nodules, the spaces
between them being strongly impressed pits. The umbilical wall is
marked by coarse, low, rounded, broad ribs which in turn are crossed
by the incremental lines. Aperture circular, protracted into an angle
at the posterior angle; outer lip of the peristome thin, the inner rather
thickened. Operculum typically cyclopilsbryid.
The type, U.S.N.M. No. 57764, was received from C. B. Adams.
It has 4.9 whorls and measures: Height, 9.9 mm.; greater diameter,
14.2 mm.; lesser diameter, 11.0 mm.
No definite locality accompanies this specimen, but the same species
was obtained by Henderson near Kings, Westmoreland Parish,
Jamaica, which is the only locality we have for it. Kings is on the
south coast of Westmoreland near the boundary of St. Elizabeth
Parish.
CYCLOPILSBRYA (CYCLOPILSBRYA) GLENBURNIENSIS, new species
Puate 12, Ficurres 18-20
Shell large, helicoid, the denuded specimens soiled yellowish white.
The nucleus consists of almost 1.5 well-rounded smooth turns. The
postnuclear whorls are well rounded and marked by retractively
curved, closely approximated, rather broad axial riblets, which are
quite regular on the early whorls but become sinuous and slightly
irregular on the last whorl. Beginning with the middle half of the
penultimate whorl wrinkles make their appearance. These gain in
strength with the increasing whorls, but again fade out on the last
one-tenth of the last turn. These wrinkles are irregular in strength
and distribution and extend in a more or less zigzag pattern from
the summit toward the periphery in the best-developed portion. Su-
ture well impressed except on the last half of the turn, which creeps
up on the preceding whorl. Periphery well rounded. Base strongly
rounded, moderately broadly openly umbilicated, with a very strong
keel marking the outer edge of the umbilicus. The umbilical side
of this keel is limited by a decided excavation. On the outside a
series of rather distantly spaced, protractively curved, short, stout
74 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
riblets radiate from the keel, leaving broad impressed pits between
them. These riblets do not extend over more than one-third of the
base; the rest of the base is marked by incremental lines and slight
pittings. Aperture circular, protracted into a clawlike element at the
posterior angle; outer lip of the peristome thin, the inner somewhat
thickened, particularly so at the umbilical keel. Operculum typically
cyclopilsbryid.
The type, U.S.N.M. No. 535965, was collected by Orcutt on Glen-
burnie Mountains, Westmoreland Parish, Jamaica. It has 5.8 whorls
and measures: Height, 19.2 mm.; greater diameter, 26.5 mm.; lesser
diameter, 19.4 mm.
Four additional lots from the immediate type locality are in the col-
lection of the National Museum.
The profound pits outside of the umbilical keel readily distinguish
this species from C. (C@.) rupisfontis (Chitty), which it most nearly
resembles.
CYCLOPILSBRYA (CYCLOPILSBRYA) RUPISFONTIS (Chitty)
PLATE 12, FicuRES 24-26
1857. Cyclotus rupisfontis Curtrry, Proc. Zool. Soc. London, vol. 25, p. 155.
1898. Neocyclotus (Plectocyclotus) rupisfontis Kopetr and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 139, reprint.
Shell of medium size, elevated helicoid, covered with a dark chest-
nut-brown periostracum except the region immediately below the
summit, which is buff. There are also frequently spiral lines of the
buff coloration on the last part of the last turn. The nucleus con-
sists of 1.3 well rounded, smooth turns. The postnuclear whorls are
inflated, strongly rounded; the first is marked by very regular,
strongly elevated axial riblets, which are separated by spaces about
as wide as the riblets. Beyond this turn the axial sculpture becomes
irregular and the riblets become more or less sinuous on the next
whorl, while on the last whorl they are very sinuous and the indi-
viduals vary materially in strength from the summit to the periphery,
forming in fact a series of jointed slight wrinkles. In addition to
this, there are heavy wrinkles on the last whorl, which are also
irregular and slope in a general, slightly protractive direction. They
are strongest on the middle of the turns and weaken toward the
summit and the periphery and lend to the last whorl an irregularly
nodulose aspect. Suture well impressed except on the last turn,
which creeps very strongly up on the preceding turn, to which it
is appressed, leaving a slight concave area below its summit. Pe-
riphery well rounded. Base inflated, strongly rounded, rather nar-
rowly umbilicated, provided with a strong keel at the outer edge of
the umbilicus. On the umbilical side the keel is strongly inpinched,
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 75
while on the outside there is scarcely any impression. The umbilical
keel itself is rendered rough by the incremental lines and immediately
outside of this there are a few faint obsolete pits. The rest of the
base is almost smooth or at best exceedingly faintly, irregularly,
obsoletely nodulose. Aperture circular, protracted into a decidedly
clawlike element at the posterior angle; outer lip of the peristome
thin, the inner thickened, particularly so at the junction with the
basal keel. Operculum typically cyclopilsbryid.
The specimen figured, Acad. Nat. Sci. Philadelphia, No. 174180,
is one of a series collected by Dr. H. B. Baker on the hills north of
Retreat, Westmoreland Parish, Jamaica. It has 5 whorls and meas-
ures: Height, 19.0 mm.; greater diameter, 24.3 mm.; lesser diameter,
18.0 mm. It resembles most closely @. (C.) glenburniensis, from
which the feeble basal sculpture will readily differentiate it.
The species is found on both sides of the boundary on the south
side of Westmoreland and St. Elizabeth Parishes.
CYCLOPILSBRYA (CYCLOPILSBRYA) STRIOSA (Chitty)
PLATE 12, FicuRES 27-29
1857. Cyclotus jugosus striosus Cuirry, Proc. Zool. Soc. London, vol. 25, p. 156.
Shell very small, decidedly depressed-helicoid; the early whorls
reddish, gradually changing to wax yellow on the last turn. The
nucleus consists of about 1.5 strongly elevated smooth turns; the
succeeding whorls are marked by strongly elevated axial ribs, which
are not quite so wide as the spaces that separate them, and which
on the last turn are most conspicuous on the oblique wrinkles. Be-
ginning with the last whorl, rather rough, decidedly obliquely re-
tractively slanting cords, which are about as wide as or a little wider
than the spaces that separate them, make their appearance. ‘These
cords are well elevated and extend enfeebled to the summit. Near
the summit they assume a little more vertical direction. Suture
strongly constricted, even on the last whorl. Periphery well rounded
and marked by the sculpture of the spire. Base inflated, strongly
rounded, very openly umbilicated, with the merest indication of an
umbilical keel. The strong oblique cords described for the spire ex-
tend over the entire base and pass down on the umbilical wall as less
strong riblets. Aperture circular, slightly angulated at the posterior
angle; outer lip of the peristome thin; the inner is slightly thickened.
Operculum typically cyclopilsbryid.
The specimen figured, U.S.N.M. No. 168515, is one of a lot col-
lected by Henderson and Simpson at Ipswich in the northwestern
part of St. Elizabeth Parish, Jamaica. It has 4.5 whorls and meas-
ures: Height, 9.8 mm.; greater diameter, 14.4 mm.; lesser diameter
10.8 mm.
76 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
We have many additional lots collected by Charles R. Orcutt from
the same general region.
The small size and the depressed-helicoid form will differentiate
this from all the other members of the genus.
CYCLOPILSBRYA (CYCLOPILSBRYA) JUGOSA (C. B. Adams)
PLATE 12, FicurEs 33-35; PLAT 42, Fieures 11-13
1852. Cyclostoma jugosum C. B. ApaMs, Ann. Lyc. Nat. Hist. New York, vol. 5,
. 59.
1852. joi jugosus PFEIFFER, Monographia pneumonopomorum viventium,
vol. 1, p. 27.
1857. Cyclotus jugosus parva CuiTty, Proc. Zool. Soc. London, vol. 25, p. 156.
1898. Neocyclotus (Plectocyclotus) jugosus Koperr and MO.LeNnporFr, Nachrb.
deutschen malak. Ges., vol. 29, p. 188, reprint.
Shell very large, decidedly depressed-helicoid, almost planorbid,
covered with a pale brown periostracum. The nuclear whorls are
flesh colored, the succeeding whorls reddish, gradually paling to flesh
color. The nucleus consists of 1.6 inflated, well-rounded, smooth
whorls. The succeeding turns are marked by strongly elevated, slen-
der, very regular, slightly retractively curved axial riblets, which are
about one-third as wide as the spaces that separate them. On the
last turn and a half strong wrinkles make their appearance, which
are arranged in a peculiar zigzag pattern, the zigzagging being very
broad. These wrinkles are about as wide as the spaces that separate
them. On the last half of the last turn the zigzag arrangement
becomes less conspicuous and the sculpture assumes more the character
of nodulation. Suture well impressed on all the turns. Periphery
well rounded and marked by the continuation of the sculpture of the
spire. Base inflated, strongly rounded, and marked with strong
oblique cords near the umbilicus, which have a protracted slant. The
rest of the base is marked by finer irregular cords, which are irregu-
larly distributed but very strongly developed. Umbilicus broad and
open. The umbilical wall is marked by strong, almost sublamellar
axial riblets and lines of growth. Aperture circular, oblique, slightly
protracted at the posterior angle; outer lip of the peristome thin at
edge, the inner somewhat thickened. Operculum typically cyclo-
pilsbryid.
The specimen figured, U.S.N.M. No. 100685, is one of a large series
collected by Orcutt between Ipswich and Springfield, near the 4-mile
milestone, St. Elizabeth Parish, Jamaica. It has 5.1 whorls and
measures: Height, 16.7 mm.; greater diameter, 27.8 mm.; lesser diam-
eter, 20.4 mm.
This species is readily distinguished from @. (@.) striosa (Chitty)
by its very large size and irregular cording of the base. Chitty
described this as Cyclotus jugosus parva. We find that the size of
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA C6
this species varies very greatly, and that specimens meeting all the
requirements of Chitty’s parva may be found in any of the colonies.
They merely represent individual variation, not zoogeographic races.
This species is distributed through western St. Elizabeth Parish
and the adjacent region of Westmoreland Parish.
CYCLOPILSBRYA (CYCLOPILSBRYA) CARIBAEA (Clench and Aguayo)
Puate 13, Fiaures 48-45
1848. Cyclostoma corrugatum G. B. Sowrrpy, Proc. Zool. Soc, London, vol. 11,
p. 30 (not Cyclostoma corrugatum Menke, Synopsis methodica mol-
luscorum .. . p. 89, 1830).
1935. Poteria caribaea CLENCH and AcuAyo, Nautilus, vol. 49, p. 51, pl. 3, figs. 5, 6.
Shell large, helicoid. The nuclear whorls are pale brown, while the
succeeding turns are chestnut-brown, gradually fading to flesh color.
The nucleus consists of 1.5 well rounded, smooth turns. The post-
nuclear whorls are inflated, strongly rounded, all but the last 1.5 turns
marked by slender, rather weak and moderately distantly spaced,
slightly retractive axial riblets. The last 1.5, in addition to this
sculpture, are marked by strong wrinkles, which are very strong near
the summit and there almost vertical in position; below this there
are oblique wrinkles, which are arranged in a more or less ill-defined
zigzag fashion. Suture strongly impressed except on the last turn,
which is slightly appressed to the preceding turn. Periphery strongly
rounded and marked by the continuation of the axial sculpture. Base
inflated, strongly rounded, with very coarse, oblique, protractively
slanting wrinkled cords, which radiate from the outer edge of the
umbilicus toward the periphery, weakening somewhat toward the
periphery. The base is rather narrowly openly umbilicated and the
umbilical wall is marked by coarse, almost lamellar axial riblets and
fine lines of growth. Aperture circular, slightly protracted into an
angle at the posterior angle; outer lip thin, the inner somewhat
thickened. Operculum typically cyclopilsbryid.
The specimen figured is one of a lot, U.S.N.M. No. 535696, collected
by B. W. Oruda at Spring Mount, St. James Parish, Jamaica, and
received from the Museum of Comparative Zoology. It has 5 whorls
and measures: Height, 16.8 mm.; greater diameter, 24.8 mm.; lesser
diameter, 18.0 mm.
This species was first recognized by G. B. Sowerby, who called it
Cyclostoma corrugatum; that name being preoccupied by Menke’s
corrugatum, the adoption of Clench and Aguayo’s name caribaea
becomes necessary.
The species is rather widely distributed, ranging through Hanover,
Westmoreland, St. James, and St. Elizabeth Parishes.
428349426
78 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
CYCLOPILSBRYA (CYCLOPILSBRYA) RUFILABRIS (Chitty)
PLATE 13, Figures 1-3
1857. Cyclotus jugosus rufilabris Cuirry, Proc. Zool. Soc. London, vol. 25, p. 156.
Shell small, helicoid. Nuclear whorls flesh colored, the succeeding
rose red, the rest brown with a reddish flush; aperture red. The
nucleus consists of a little more than a well-rounded, smooth turn.
The postnuclear whorls are marked by rather closely spaced, poorly
developed, slightly retractively curved axial riblets. Beginning with
the last half of the penultimate whorl irregular wrinkles make their
appearance. These are strongest immediately anterior to the summit,
where they appear as groups of heavy broad nodules. Anterior to
this the finer cords, which are decidedly oblique and irregular, are
present. Suture well impressed on all the whorls except the last part
of the last, in which the summit creeps up on the preceding turn. The
periphery is well rounded and marked by the continuation of the
sculpture of the spire. Base inflated, strongly rounded, with very
strong, protractively curved, oblique wrinkles, which are separated by
spaces as wide as the wrinkles. These do not extend to the periphery,
but fade out toward the middle of the base, the anterior part being
sculptured like the upper surface. The umbilicus is not very broad
and its wall is marked by rather strong axial ribs, which, like the
spaces between them, bear fine incremental lines. Aperture circular,
decidedly oblique, protracted into an angle at the posterior angle;
outer lip of the peristome thin, the inner somewhat thickened.
Operculum typically cyclopilsbryid.
The specimen figured, U.S.N.M. No. 356148, bears the label “Ja-
maica,” without detailed locality. It has 4.5 whorls and measures:
Height, 12.2 mm.; greater diameter, 16.8 mm. ; lesser diameter, 12.5 mm.
Three additional lots are in the collection of the National Museum.
Two of these come from Balaclava, St. Elizabeth Parish, and another
from Pepper, St. Elizabeth Parish.
This species resembles most nearly C. (C.) asperula (Sowerby),
from which it can readily be distinguished by its stronger sculpture.
CYCLOPILSBRYA (CYCLOPILSBRYA) ASPERULA (Sowerby)
PLATE 13, FIGURES 4-6
1843. Cyclostoma asperulum SoweErBy, Thesaurus conchyliorum, vol. 1, p. 91, pl.
23. fis, 3:
1852. Cyclotus asperulus PFEIFFER, Monographia pneumonopomorum viventium,
vole dips 21.
1898. Neocyclotus (Plectocyclotus) asperulus KoseELT and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138, reprint.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 79
Shell small, helicoid, flesh colored, with a brownish flush. The nu-
cleus consists of a little more than a single turn, which is well rounded
and smooth. The postnuclear whorls are inflated, strongly rounded,
and marked by feeble, retractively curved axial riblets, which are
about half as wide as the spaces that separate them. On the last 1.5
turns feeble wrinkles make their appearance, which on the last whorl
are very regularly developed and distributed. They are not very prom-
inent even here. Suture moderately strongly impressed. Periphery
strongly rounded. Base inflated, strongly rounded, very narrowly um-
bilicated, marked by wrinkles, pits, and nodules. The umbilical wall
is marked by slender riblets and lines of growth. Aperture circular,
oblique, protracted into a clawlike element at the posterior angle; outer
lip of the peristome thin, the inner somewhat thickened. Operculum
typically cyclopilsbryid.
The specimen figured, U.S.N.M. No. 474033, bears the label “Ja-
maica,” without definite locality designation. It has 4.6 whorls and
measures: Height, 12.3 mm.; greater diameter, 16.2 mm.; lesser
diameter, 12.5 mm.
This species resembles in size and shape C. (C.) rufilabris (Chitty),
from which it can be easily differentiated by its weak sculpture.
Two additional specimens in our collection are also without locality
designation.
CYCLOPILSBRYA (CYCLOPILSBRYA) BESWICKI (Chitty)
1857. Cyclotus beswicki Cuirry, Proc. Zool. Soc. Lendon, p. 157.
1898. Neocyclotus (Plectocyclotus) beswicki Kospetr and MO6LLENporFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138, reprint.
I have not seen this species, so I quote Chitty’s description:
“Form, subglobose-conic. Colowr, pinkish, with light brown epi-
dermis. Sculpture, very coarse lines of growth, very slight distant
corrugation on the last whorl, coarser below; fine lines of growth within
umbilical keel. Spire, well elevated, with almost straight outlines.
Whorls, 514, moderately rounded, with moderate suture. Aperture,
oblique and much dilated to the right below, depressed above. Peri-
treme, slightly sinuate above, sharp on the right. Umbilicus, moderate,
greatest width 0.25 [6.25 mm.]. Umbilical keel, strongly produced.
Operculum, with lamina of about 9 much incurved close spiral whorls,
which are very slightly concave on the entire margins; end of last a
little incurved to preceding one. Height 0.75 [18.75 mm.]; greatest
breadth 0.9 [22.5 mm.]; least breadth 0.74 [18.50 mm. ].
“Bogue Estate, north-east corner of St. Elizabeth.”
CYCLOCAYMANIA, new subgenus
In this subgenus the spiral lamella is curved inward to the very
edge, but attached to its free edge on the later turns is a broad fold,
80 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
which extends in a curve downward and outward to almost meet
the next whorl.
Type: Poteria caymanensis Pilsbry=Cyclopilsbrya (Cyclocay-
mania) caymanensis (Pilsbry).
KEY TO THE SPECIES OF SUBGENUS CYCLOCAYMANIA
Shell rugose.
Umbilicus: wides2= 34 = ee Bees Sa ee ee caymanensis
Umbilicug: Narrows, (ses ee eee ee eee fonticula
Shell noe srs OS ere ce ee a ee laevitesta
CYCLOPILSBRYA (CYCLOCAYMANIA) CAYMANENSIS (Pilsbry)
PLATE 138, FicurEs 7-9, 10-12 type
1930. Poteria caymanensis PILsBRY, Proc. Acad. Nat. Sei. Philadelphia, vol. 82,
pl. 18, figs. 4, 5.
Shell small, depressed-helicoid, flesh colored when the periostracum
is removed. Nuclear whorls 1.5, well rounded, smooth. Postnuclear
whorls well rounded; the first with axial riblets, the rest with axial
riblets which are less regular and toward the end become obsolete
and decidedly irregular. On the middle of the postnuclear whorls
there are decidedly obliquely retractively slanting rugae, which, com-
bined with the axial sculpture, produce a decidedly rough surface.
On the last part of the last turn the rugae almost disappear and the
whorl, while still rough, is not as conspicuously patterned. Suture
well impressed. Aperture rounded. Base openly umbilicated with a
very strong carina marking the outer limits of the umbilicus. Within
the umbilicus the shell is inpinched above the keel. The surface of
the base has the rugations very strongly marked and here they appear
quite regular and separated by spaces a little narrower than the
raised ridges. The axial threads of growth here also are less pro-
nounced, while on the umbilical wall the axial riblets are more em-
phasized. Aperture decidedly oblique, somewhat peaked at the pos-
terior angle; peristome moderately thickened. The last whorl is
slightly descending and the peristome is therefore rendered free.
Operculum ?
Dr. H. A. Pilsbry collected the type and the paratype of this
species 1 mile south of Georgetown, Grand Cayman. They constitute
Acad. Nat. Sci. Philadelphia No. 150857. The type has 4.5 whorls
and measures: Height, 8.7 mm.; greater diameter, 14.0 mm.; lesser
diameter, 11.0 mm. The paratype has lost the early whorls. It
measures: Height, 10.6 mm.; greater diameter, 16.2 mm.; lesser
diameter, 12.8 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA sl
CYCLOPILSBRYA (CYCLOCAYMANIA) FONTICULA (Preston)
PLATE 18, Figures 13-18; PLATE 42, Fiaures 6-7
1911. Neocyclotus fonticulus PRESTON, Proc. Malac. Soe. London, vol. 9, p. 359, fig.
Shell small, helicoid, covered with a wood-brown periostracum.
Nuclear whorls 1.2, small, well rounded, smooth. Postnuclear whorls
strongly rounded, the first turn marked by slender, retractively curved
axial riblets. Beyond this the axial riblets give way to semilamellar,
more or less closely spaced, irregularly developed, retractively slanting
axial elements. These are rendered rough and more or less wavy by
retractively curved, broad rugae, which on the last part of the last turn
become enfeebled and almost vanish toward the aperture. Suture
strongly impressed. Periphery well rounded. Base moderately well
rounded, limited at the umbilicus by a very heavy keel. The base is
sculptured by the continuation of the fine axial lamellae and by very
strong, retractively curved, quite regular and regularly spaced cords,
which are separated by grooves a little less wide than the cords. The
base is openly umbilicated, the umbilicus being about one-third the
width of the greater diameter of the shell. The umbilical wall appears
excavated above the bounding keel and is marked by the axial lamella-
tion only. In apparently very old, senescent individuals, the last whorl
descends near the aperture. In a majority of specimens, however,
this is not the case. Aperture decidedly oblique, subcircular. In the
specimens where the last whorl is adnate to the preceding turn, the
attachment to the parietal wall terminates in a lunate line and the
peristome is protracted into a decided point at the posterior angle. In
specimens where the last whorl is descending, the aperture becomes
more nearly circular and the posterior angle shows a carina on the
outside. Peristome simple; the outer lip thin; the inner lip, however,
is materially thickened. Operculum typically cyclocaymanid.
Preston’s type shows the senescent state, and I am figuring two speci-
mens, one corresponding to his figure, and another showing the more
prevalent phase. These have 4.9 and 4.8 whorls, respectively, and
measure: Height, 10.0, 9.1 mm.; greater diameter, 14.5, 14.6 mm.; lesser
diameter, 11.5 and 11.0 mm., respectively.
These were selected from a large series of specimens, U.S.N.M. No.
466831, collected by T. Savage English on Grand Cayman Island, with-
out specific locality.
Dr. H. A. Pilsbry obtained this species in the woods southwest of
North Sound, and I found a dead specimen, U.S.N.M. No. 392278, at
the church between St. George and Bowdentown.
82 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
CYCLOPILSBRYA (CYCLOCAYMANIA) LAEVITESTA (Pilsbry)
PLATE 13, FIcuRES 22-24
1930. Poteria laevitesta Pruspry, Proc. Acad. Nat. Sci. Philadelphia, vol. 82, pp.
227-228, pl. 16, figs. 18, 18a, 19, 19a.
Shell small, helicoid, covered with a honey-yellow periostracum.
Nuclear whorls 1.4, small, well rounded, smooth. The postnuclear
whorls are well rounded, the first marked by very regular, retrac-
tively slanting axial riblets, which are not quite so wide as the spaces
that separate them. After this the riblets become less regular and
more inclined to be lamellose and closely approximated, which render
the surface of the spire rather rough but without rugations. Suture
moderately constricted. Periphery almost angulated. Base short,
well rounded, marked like the spire with indications of fine spiral
lirations. A strong keel forms the umbilical angle. The umbilical
wall is excavated above this keel. The base also is devoid of ruga-
tions. The umbilicus is about one-fourth the diameter of the shell.
Aperture decidedly oblique, protracted at the posterior angle into an
almost spurlike element. Peristome simple, outer lip thin; inner lip
decidedly thickened. Operculum typically cyclocaymanid.
The specimen described and figured, Acad. Nat. Sci. Philadelphia
No. 150658, is a paratype collected by Charles B. Taylor on Grand
Cayman. It has 4.6 whorls and measures: Height, 11.5 mm.; greater
diameter, 16.5 mm.; lesser diameter, 13.5 mm.
Genus PTYCHOCOCHLIS Simpson
1895. Ptychocochlis Stmpson, Proc. U. S. Nat. Mus., vol. 17, p. 481.
Simpson replaces Platystoma Klein, several times preoccupied, with
the name Ptychocochlis and cites as type Neocyclotus jamaicensis
Chemnitz, rechristened orthodoxly by Menke in 1880 as Cyclostoma
corrugatum. The type of Ptychocochlis Simpson is Ptychocochlis
corrugata (Menke) = Turbo jamaicensis Chemnitz, Conchylien Cabi-
net, vol. 11, pp. 277-278, pl. 209, figs. 2057, 2058, 1795 (nonbinomial)
rechristened in 1830 as Cyclostoma corrugatum Menke, Synopsis
methodica molluscorum * * *, p. 39.
This genus is differentiated from Poteria by having the elevated
lamellae of the operculum expanded at the summit to resemble the
capital letter T or L inverted, the outer limb of the cross bar being
much longer than the inner and touching that of the succeeding
whorl, thus forming a complete deck.
Type: Cyclostoma corrugatum Menke=Ptychocochlis corrugata
(Menke).
The radulae of Ptychocochlis clappi, P. martensi (Kobelt), and
P. savannensis show the formula 3:3:3:3. The jaw is without a
median projection. The verge of P. vendreysi is situated on the
THB CYCLOPHORID LAND MOLLUSKS OF AMERICA 83
back of the neck behind the tentacles. It is swollen basally and pro-
vided with a seminal groove and a very short simple terminal
appendage.
KEY TO THE SPECIES OF THE GENUS PTYCHOCOCHLIS ‘
Nodulations present on base.
Nodulations on base strong.
Strong nodulations of base extending from umbilical keel to
periphery.
Strong nodulations present on base of all of last whorl.
Upper side of last whorl very coarsely wrinkled__ corrugatior
Upper side of last whorl] not very coarsely wrinkled.
Wrinklestofihbase coaTs@esw. 22 8 gossei
Wrinkles of base fine.
Shell more than 20 mm. in diameter__-______ varians
Shell less than 14 mm. in diameter_-________ gemma
Strong nodulations present on base of early portion of
last whorl only.
Nodulations strong on all the upper surface of last
whorl.
iimhilrcalikeelsstrong= = 42 _ 2 heat Soe ete ees welchi
Umbilical keel obsolete_.__.-__________ manchesterensis
Nodulations not strong on all the upper surface of
last whorl but obsolete on last part.
Nodulose ridges retractively slanting at periphery_ taylori
Nodulose ridges protractively slanting at pe-
riphery.
Nodulations coarse 23 ees ee ee eee ae zigzag
Nodulations finers22 ee ea eee eve shawae
Strong nodulations of base not extending from umbilical
keel to periphery.
Nodulations of upper surface of last whorl few and
CUS LAVET EB yp SJO SL COU een nee eee ee ete ee earn Sense ones Hee) clappi
Nodulations of upper surface of last whorl not few or
distantly spaced.
Shell gigantic, greater diameter more than 32 mm______ senex
Shell not gigantic, greater diameter 27 mm, or less.
Nodulations on base very strong______--______ corrugata
Nodulations on base not very strong___-_________ adamsi
Nodulations on base feeble.
Oblique, protractively curved wrinkles present near umbili-
GalleKGe TS 55 Ee as eh he ba cee Ok See simpsoni
Oblique, protractively curved wrinkles absent near umbili-
cal Keel.
Weak nodules present on upper surface of all of last
whorl.
INodules on wbaseiweakeo2e ssart WA aris Dupe sere) martensi
Nodules on base obsolete.
UmbilicusAwide i) 2 ace. rks Sa eer es hendersoni
Wmbilicusinarro yw wee tse Ne ae eee TER, marianna
4T have not included P. subrugosa (Sowerby) in this key, since I have not seen specimens
of it.
84 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Weak nodules not present on the upper surface of all
of last whorl.
Penultimate whorl with obliquely curved wrinkles.
Wrinkles retractively curved on anterior half
OP WO Ts ee ee ee er lacteofiluvialis
Wrinkles protractively curved on anterior
half of whorl.
Nodulose sculpture strong___-----------_ savannensis
Nodulose sculpture weak__-_---__----------- orcutti
Penultimate whorl without obliquely curved wrin-
kles.
Wrinkles on penultimate whorl arrow-shaped.
Arrow-shaped wrinkles pointing forward____-- minor
Arrow-shaped wrinkles pointing back-
ward.
Shell decidely elevated____________--__-_ magna
Shell not decidedly elevated___---- montegoensis
Wrinkles on penultimate whorl not arrow-
shaped.
Umbilicus: very ‘narrow =22.2-22=-4______ vendreysi
Umbilicus not very narrow_.------------ subglobosa
Nodulations absent, on: base===-— == ae eee campeachyi
PTYCHOCOCHLIS CORRUGATIOR (Chitty)
PLATE 13, FIGURES 37-39
1857. Cyclotus corrugatior Cuirty, Proc. Zool. Soc. London, vol. 25, p. 149.
1859. Neocyclotus (Plectocyclotus) corrugatior KoseLr and MOLLENDoRFF, Nachrb.
deutschen malak. Ges., vol. 31, p. 136.
Shell of medium size, depressed-helicoid. Nuclear whorls flesh
colored, the succeeding turns wood brown, the last one paler, with the
summit of the corrugations lighter than the rest of the shell. Peristome
and interior soiled bluish white. The nucleus consists of about 1
smooth well-rounded turn. The first 114 postnuclear turns are well
rounded and marked by slightly retractively curved axial riblets, which
are about half as wide as the spaces that separate them. The suc-
ceeding turns are also strongly rounded and marked by the continua-
tion of the axial riblets, which here are rendered irregularly disposed
by the strong rugations. Beginning with the third whorl and extend-
ing over the remaining turns, except the last half of the last whorl,
strongly raised, decidedly oblique, protractively slanting ridges make
their appearance. These ridges extend to about one-fourth of the
distance between the summit and the periphery anterior to the summit.
They are rendered slightly nodulose by the axial riblets. The last half
of the last turn is marked by nodules, which are not of uniform
strength or spacing, or the direction of slant. These are also rendered
more finely nodulose by the crossing axial riblets. The suture of all
but the last half turn is deeply impressed. In the last half turn the
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 85
summit of the whorls is appressed to the preceding turn. Periphery
well rounded. Base well rounded, openly umbilicated, and provided
with a strong keel at the outer limit of the umbilicus. The space be-
tween the umbilical keel and the periphery is marked by raised, re-
tractively slanting ridges, which join those on the upper portion of
the whorl, but here have the reverse slant. They are not quite so
strong as those on the upper surface and they weaken in strength
toward the umbilical keel. In addition to this, strong incremental
lines representing the continuation of the finer axial riblets are present.
The umbilical wall is marked by crowded, rather rough axial riblets
and incremental lines. Aperture subcircular, slightly angulated at the
posterior angle where it is also somewhat protracted; the outer per-
istome is strongly evenly curved; the inner slightly thickened at the
umbilical keel. Operculum typically ptychocochlid.
The specimen figured has 5 whorls and measures: Height, 13.5 mm.;
greater diameter, 21.2 mm.; lesser diameter, 15.9 mm. It is one of the
specimens in the Adams collection at Amherst College, and the label
states that it came from Gutters and was donated by Chitty.
The species centers in Manchester Parish, Jamaica, but it also ex-
tends into Clarendon to the east and the eastern part of St. Elizabeth
and to the north into St. Ann Parish.
PTYCHOCOCHLIS GOSSEI, new species
Pate 13, Ficures 34-36
Shell rather large, helicoid. The nucleus and early postnuclear
turns are almost blood-red; the rest unicolor, chestnut-brown, or
banded. In the type there is a white band immediately below the
summit and a broad light zone extending from the periphery to the
middle of the base, and finer paler lines on the anterior portion of the
base, which is also of a paler tint than the upper surface. The nucleus
consists of a single, well-rounded, smooth turn. The postnuclear
whorls are strongly rounded. The first postnuclear whor!] is marked
by slender, retractively curved axial riblets, which are not quite as
wide as the spaces that separate them. These fine riblets continue
over the rest of the whorls, becoming a little heavier and decidedly
irregular in distribution on the last whorl. On the second postnuclear
whorl the beginning of the decidedly oblique, protracted ridges make
their appearance. These become intensified as the shell increases in
size, and on the penultimate whorl] they are about as wide as the spaces
that separate them. They do not extend on the anterior fourth of the
whorls and on the last half of the last whorl they become much en-
feebled. They are never as pronounced as in P. corrugatior (Chitty).
On all the whorls, except the last, the suture is well impressed ; on the
latter the summit is appressed to the preceding turn. Periphery well
86 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
rounded. Base strongly rounded, moderately broadly openly
umbilicated, with a strong broad keel limiting the outer termination
of the umbilicus. The posterior half of the base is decidedly wrinkled,
almost to the very edge of the peristome. On the anterior half the
wrinkling is almost absent. The umbilical keel is rendered somewhat
notched by the axial sculpture. The umbilical wall is marked by fairly
strong axial riblets and lines of growth. Aperture subcircular,
angulated at the posterior angle and slightly effuse at the junction of
the basal and outer lip. The outer lip is thin; the inner, some-
what thickened. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 398348, is one of a series collected by Orcutt
at Hermitage Dam, Jamaica, that is, on the main road between Con-
stant Spring and Stony Hill, St. Andrew. It has 5 whorls and
measures: Height, 18.7 mm.; greater diameter, 25.2 mm.; lesser
diameter, 18.8 mm.
The larger size, less open umbilicus, and more elevated form, as well
as the feebler nodulation, will readily distinguish this species from
P. corrugatior.
PTYCHOCOCHLIS VARIANS (C. B. Adams)
PLATE 13, FIGURES 40-42
1852. Cyclostoma varians C. B. ApAMs, Ann. Lyc. Nat. Hist. New York, vol. 5,
p. 59.
1852. Cyclotus varians PFEIFFER, Monographia pneumonopomorum viventium,
Suppl. 1, p. 27.
1898. Neoayclotus (Plectocyclotus) varians Kosrirr and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 189, reprint.
Shell of medium size, soiled flesh colored, covered by a pale brown
periostracum. The nuclear whorls appear to vary from flesh colored
to reddish. The nucleus consists of a little more than a turn, which
is smooth and well rounded. The postnuclear whorls are strongly
rounded; the first is marked by slender, retractively curved axial rib-
lets, which are not quite so wide as the spaces that separate them.
Beginning with the second postnuclear whorl, strong, oblique, protrac-
tively curved ridges make their appearance. These continue to the
early part of the last whorl. These ridges do not extend quite to the
summit but stop at about one-fourth of the distance between the
summit and periphery, anterior to the summit. The spaces separat-
ing these ridges are about equal to the ridges. They, as well as the
ridges, are crossed by the slender axial riblets. On the last whorl
the oblique ridges gradually disappear and are replaced by irregular
wrinkles that appear to be without definite direction. This sculpture
extends almost to the peristome. The suture is well impressed on all
but the last turn, which is appressed at the summit. Periphery well
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA i7,
rounded. Base strongly rounded, openly moderately broadly umbili-
cated with a strong keel bounding the outer edge of the umbilicus.
The entire surface of the base is marked by irregularly disposed
wrinkles and pits. The umbilical keel is rendered rather rough by
the incremental lines, which extend strongly and roughly into the
umbilicus. Aperture subcircular, rather strongly protracted into an
angle at the posterior angle; outer peristome thin, the inner somewhat
thickened. Operculum typically ptychocochlid.
The specimen figured is one of C. B. Adams’ original lot, No. 6.
This was selected by H. B. Baker ° as the type of Adams’ complex. It
has 5 whorls and measures: Height, 14.6 mm.; greater diameter, 20.8
mm.; lesser diameter, 15.6 mm. No specific locality accompanies this
lot.
The species appears restricted to the north coast of Jamaica in St.
Mary and St. Ann Parishes, that is, the stretch between Port Maria
and Ocho Rios Bay.
PTYCHOCOCHLIS GEMMA (Chitty)
PuaTe 13, FIGuRES 25-27
1857. Cyclotus gemma Currty, Proc. Zool. Soc. London, vol. 25, p. 150.
1898. Neocyclotus (Plectocyclotus) gemma Kosetr and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138, reprint.
Shell small, depressed-helicoid, pale horn colored. The nucleus
consists of a single well-rounded, smooth turn. The postnuclear
whorls are well rounded; the first is marked by slender, retractively
curved axial riblets, which are about as wide as the spaces that sep-
arate them. The second postnuclear whorl shows the beginning of
oblique wrinkles, which are not quite as wide as the spaces that
separate them. On the last whorl these wrinkles assume an irregular
distribution, the spaces between them being a little wider than the
wrinkles. These rugations do not extend quite tothe summit. Suture
well impressed in all the turns. Periphery well rounded. Base
broadly openly umbilicated, marked by very strong, rough, decidedly
oblique, protracted cords, which are wider than the spaces that
separate them and are rendered roughened by the axial riblets. The
umbilical keel is but feebly expressed, and the umbilical wall is
marked by closely spaced axial riblets. Aperture circular, slightly
angulated at the posterior angle, oblique; peristome thin. Operculum
typically ptychocochlid.
The specimen figured is C. B. Adams’ unique type in the Amherst
College collection, received from Chitty, and is said to have come
from Burnt Hill, Westmoreland Parish, Jamaica. It is the only
5 Nautilus, vol. 48, p. 86, 1935.
88 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
specimen that I have seen. It has 4.2 whorls and measures: Height,
7.9 mm.; greater diameter, 12.7 mm.; lesser diameter, 9.0 mm.
PTYCHOCOCHLIS WELCHI, new species
PLatTe 13, Fiaures 28-30
Shell small, depressed-helicoid, the early whorls red, the rest oliva-
ceous with a chestnut-brown ee interior of the arene bluish
white. The nucleus consists of 1.2 well-rounded, smooth turns. The
first postnuclear turn is well rounded and marked by poorly devel-
oped, somewhat irregularly distributed, feeble axial riblets. On the
remaining turns these riblets become more and more irregular owing
to the rugate ridges marking the surface. Beginning with the sec-
ond postnuclear whorl, these ridges gradually develop; they extend
from almost at the summit to the periphery and have a decidedly
protracted slant. They are not quite as wide as the low spaces that
separate them, and are crossed by the slender riblets. Suture strongly
constricted. Periphery well rounded. Base openly umbilicated with
a moderately strong keel marking its outer limit. The base is marked
by weak, retractively curved ridges, which are best developed on the
early pare of the last turn and become obsolete on the later portion.
They are strongest near the periphery. The umbilical wall is marked
by strong, fairly coarse riblets. Aperture circular, protracted into
a decided angle at the posterior angle. Peristome thin. Operculum
typically ptychocochlid.
The type, U.S.N.M. No. 535966, was collected by d’Alté A. Welch
at his station A7, at Hardware Gap, St. Andrew Parish, Jamaica.
Tt has 4.2 whorls and measures: Height, 10.6 mm.; preater diameter,
15.0 mm.; lesser diameter, 11 mm.
Nine topotypes are in Dr. Welch’s collection. This species suggests
P. manchesterensis, from which its coarser sculpture on the upper sur-
face, more pronounced umbilical keel, and feebler sculpture of the base
will readily distinguish it.
PTYCHOCOCHLIS MANCHESTERENSIS, new species
PLATE 13, FicuRES 19-21
Shell of medium size, helicoid, pale chestnut-brown. Nuclear
whorls 1.2, well rounded, smooth. The first postnuclear whorl is
strongly rounded and marked by slender, slightly retractively curved,
rather closely spaced axial riblets. These axial riblets become
stronger and less regular on the succeeding postnuclear turn. On
the last whorl they are decidedly sinuous. Beginning with the sec-
ond postnuclear turn in addition to the axial riblets, the whorls are
marked on the anterior two-thirds by decidedly obliquely slanting,
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 89
protracted ridges, which are less than half the width of the spaces
that separate them. Suture strongly impressed. Periphery of the
last whorl well rounded. Base broadly openly umbilicated, the
outer edge of the umbilicus being marked by an obsolete cord. The
base is marked by rather closely spaced, broad, retractively curved,
moderately elevated ridges, which are best developed on the early
portion of the last turn. On the last portion of the last turn these
ridges become obsolete. The umbilical wall is marked by rough
riblets and incremental lines. Aperture circular, rather effuse at
the junction of the outer and basal lip, and slightly angulated at the
posterior angle; peristome thin. Operculum typically ptychocochlid.
The type is one of six specimens, No. 174135, in the collection of
the Academy of Natural Sciences of Philadelphia. It was collected
by H. Burrington Baker at Belretiro, Manchester Parish, Jamaica.
It has 4.5 whorls and measures: Height, 11 mm.; greater diameter,
17.4 mm.; lesser diameter, 13.0 mm.
PTYCHOCOCHLIS TAYLORI, new species
PLATE 13, Fiecures 31-33
Shell of medium size, varying in shape from helicoid to depressed-
helicoid, covered with a thin olivaceous periostracum. The nucleus
consists of a single smooth, well-rounded turn. The postnuclear
whorls are strongly rounded; the first 1.5 are marked by rather weak,
slender axial riblets which are rather distantly spaced. On the suc-
ceeding turn the axial riblets become irregular, sinuous, and almost
narrowly lamellose. Beginning with the second postnuclear whorl
and extending to the last half of the last turn, very oblique, protac-
tively slanting ridges are present, which are about as wide as the
spaces that separate them. On the last half of the last turn these
ridges become very irregular and have the nodules all arranged in
a retractive position. Suture well impressed. Periphery well
rounded. Base well rounded, openly, rather broadly umbilicated
with a rather strong keel at the outer limit of the umbilicus. The
early part of the base bears quite strong nodulations, which gradu-
ally become weaker toward the end. ‘The umbilical wall is marked
by numerous, closely spaced, slender hairlike riblets. Aperture sub-
circular, oblique, slightly protracted and angulated at the posterior
angle; outer peristome thin, the inner peristome is rather thickened.
The type, U.S.N.M. No. 356032, was collected by C. B. Taylor in
Upper Clarendon Parish, Jamaica. It has 4.4 whorls and measures:
Height, 11.8 mm.; greater diameter, 17.0 mm.; lesser diameter, 13.1
mm.
90 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
An additional specimen from the same locality is considerably
larger. It has 4.8 whorls and measures: Height, 15.2 mm.; greater
diameter, 21.6 mm.; lesser diameter, 16.0 mm.
PTYCHOCOCHLIS ZIGZAG (Chitty)
PLATE 14, Ficures 138-15
1857. Cyclotus zigzag Cu1try, Proc. Zool. Soc. London, vol. 25, p. 150.
1898. Neocyclotus (Plectocyclotus) zigzag KopBetr and MOLLENDoRFF, Nacnro.
deutschen malak. Ges., vol. 29, p. 139, reprint.
Shell of medium size, depressed-helicoid. All our specimens have
lost the epidermis, the remaining shell being soiled white. The nu-
cleus consists of a little more than a single well-rounded, smooth turn.
Postnuclear whorls strongly rounded; the first is marked by feeble,
rather distantly spaced axial riblets. On the succeeding postnuclear
whorls the axial riblets become rather irregular, owing to the nodula-
tions; they also become stronger and appear almost lamellose. Be-
ginning with the second postnuclear whorl, nodular ridges make their
appearance. These ridges are not so wide as the spaces that separate
them and do not extend over the posterior fourth of the turns. They
are decidedly obliquely protractively curved. They become obsolete
on the last fourth of the last whorl. Suture well impressed. Pe-
riphery well rounded. Base openly umbilicated, strongly rounded,
provided with a very strong keel marking the outer limit of the
umbilicus. This keel is decidedly inpinched on the umbilical side,
and a little less so on the outside. The first half of the last turn of
the base is strongly nodulose; on the latter part the nodulation be-
comes enfeebled. The umbilicai wall bears numerous axial riblets and
fine hairlike incremental lines. Aperture circular, oblique, protracted
into a slight angle at the posterior angle. Peristome thin on the
outer lip and thickened on the inner. Operculum typically
ptychocochlid.
The specimen figured, U.S.N.M. No. 399500, is one of a series col-
lected by C. R. Orcutt in Grays Valley, Trelawny Parish, Jamaica. It
has 4.5 whorls and measures: Height, 12.4 mm.; greater diameter,
18.2 mm.; lesser diameter, 13.8 mm.
We have this species from quite a number of localities, all of which
are located on the northern rim of Trelawny and the western portion
of St. Ann Parishes.
PTYCHOCOCHLIS SHAWAE, new species
PLATE 14, FicuREs 10-12
Shell of medium size, helicoid. The nuclear whorls and first post-
nuclear turn pale, the succeeding turns chestnut-brown, gradually
becoming paler on the last whorl. Base a little paler than the spire.
Aperture soiled white with a brownish tinge. The nucleus consists
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 91
of a single well-rounded, smooth turn. Postnuclear whorls well
rounded; the first is marked by slender, decidedly retractively curved,
rather closely spaced axial riblets, which are fairly strong. The
succeeding turns also have axial riblets, which are less regular, and
quite irregular on the last whorl, where they are almost lamellose.
In addition to this, beginning with the second postnuclear whorls,
there are strong, decidedly retractively slanting ridges that do not
extend on the posterior fourth of the turns. These ridges are not
quite so wide as the spaces that separate them, and they become
obsolete on the last third of the last whorl, where they are replaced
by less strong irregular nodulations. Suture strongly impressed.
Periphery well rounded. Base well rounded, openly umbilicated, the
umbilicus bounded by. a strong keel, which is inpinched on both
sides. The base is marked on the early portion of the last turn by
oblique, retractively slanting ridges, which are replaced on the last
half of the turn by irregular wrinkles. The umbilical wall is marked
by riblets and fine lines of growth. Aperture circular, oblique,
slightly protracted into an angle at the posterior angle. Peristome
thin. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 535967, is one of a series of specimens
collected by Orcutt at Balaclava, St. Elizabeth, Jamaica. It has
4.8 whorls and measures: Height, 14.0 mm.; greater diameter, 19.8
mm.; lesser diameter, 15.2 mm.
A host of specimens before me appear to confine this species to
the Cockpit region of northern St. Elizabeth Parish.
PTYCHOCOCHLIS CLAPPI, new species
PLATE 14, Ficures 19-21
Shell very large, helicoid, covered with a chestnut-brown periostra-
cum. The nucleus consists of 1.5 well-rounded smooth turns. The
postnuclear whorls are well rounded, the early ones separated by a
well-marked suture, and the later ones appressed at the summit.
The first 1.5 nuclear whorls are marked by slender, slightly retract-
ively curved axial riblets, which are about as wide as the spaces that
separate them. Beyond this the nodular ridges gradually make their
appearance. These are rather irregular, that is they begin at the
summit as retractive elements, then near the middle of the whorl bend
forward into protractive ridges, thus producing a somewhat arrow-
pointed cone-in-cone effect. This arrangement disappears on the
last half of the last turn, where the nodulations become irregular
and quite obsolete. On the last fourth of the last whorl, spiral
threads make their appearance. The last whorl is strongly con-
stricted below the summit, which causes the posterior portion to
appear decidedly appressed to the preceding turn. Periphery well
92 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
rounded. Base well rounded, openly umbilicated, with a strong keel,
which is not only pinched in on the umbilical side, but appears even
excavated at this place. On the outside of the keel there is also a
depression and here we find strongly developed pits and nodules
which extend over the anterior half of the base. The umbilical
wall is marked by somewhat irregular riblets and lines of growth.
Aperture circular, strongly protracted and angulated at the posterior
angle; outer lip of the peristome thin; the inner somewhat thickened.
The operculum differs from the typical ptychocochlid in having the
outward reflected portion of the whorls not quite touching that of the
expanded inner portion of the succeeding turn, thus leaving a slight
channel between the two in some of the turns.
The type is one of C. B. Adams’ complex of Cyclostoma varians.
It is part of his No. 7 and is labeled Portland, Jamaica, Agnes Hines,
collector.
The type is in the Amherst College collection. It has 5 whorls and
measures: Height, 22.8 mm.; greater diameter, 31.4 mm.; lesser
diameter, 23.0 mm.
PTYCHOCOCHLIS SENEX, new species
PLATE 14, Figures 22-24
This is the largest of the known Jamaican cyclophorids. It has a
helicoid shell. The specimen is without periostracum. The nucleus
consists of a little more than one turn, which is well rounded and
smooth. The postnuclear whorls are well rounded; the first 2.5 are
marked by slender, slightly retractively curved, hairlike axial rib-
lets, which are not quite so wide as the spaces that separate them.
These are followed by a turn in which very strong, retractively
curved ridges are present. On the first half of the last whorl these
ridges assume an arrowpoint arrangement, a cone-in-cone type, in
which the point of the arrow is almost on the middle of the whorl.
On the last half of the last whorl this arrangement becomes obsolete
and the shell is marked by strong incremental lines and nodulations
near the summit, the senescent last portion appearing strongly den-
ticulated. The suture of the early whorls is well impressed, while
on the last turn the summit of the whorls becomes appressed to the
preceding turn, and materially creeps up on this. Periphery well
rounded. Base well rounded, openly umbilicated with a strong,
heavily nodulose keel marking the outer limit of the umbilicus.
From this keel there radiates toward the middle of the base a series
of heavy ridges which are separated by spaces about as wide as the
ridges. These ridges have a protractive slant. They become obso-
lete on the last fifth of the last whorl. The umbilical wall is marked
by coarse ribs, which bear finer hairlike incremental lines. Aper-
THB CYCLOPHORID LAND MOLLUSKS OF AMERICA 93
ture subcircular, oblique, protracted to form an angle at the posterior
angle; peristome moderately thickened all around. Operculum
unknown.
The unique type is in the C. B. Adams collection. It measures:
Height, 22.4 mm.; greater diameter, 33.4 mm.; lesser diameter, 26.2
mm. It was with Adams’ No. 7 of Cyclostoma varians collected by
Agnes Hines in Portland Parish, Jamaica.
PTYCHOCOCHLIS CORRUGATA (Menke)
PuLaAtTe 14, FicuREs 7-9
1795. Turbo jamaicensis CHEMNITZ, Martini-Chemnitz Conchylien Cabinet, vol.
11, p. 277, pl. 209, figs. 2057, 2058 (nonbinomial).
1830. Cyclostoma corrugatum MENKE (=—Turbo jamaicensis Chemnitz), Synopsis
methodica Molluscorum, p. 39.
1857. Cyclotus portlandensis CuirtTy, Proc. Zool. Soc. London, vol. 25, p. 147.
Shell moderately large, helicoid, chestnut-brown, a little paler on
the last whorl, which tends toward buff. The appressed summit of
the whorls is also a little paler than the rest of the shell. There is
a broad dark band at the periphery, while the anterior portion of
the base and the umbilicus are flesh colored with a yellowish tinge,
which is also the color of the interior of the aperture. The nucleus
consists of a little more than a turn, which is well rounded and
smooth. The postnuclear whorls are strongly rounded; the first one
and three-quarters are marked by slender, decidedly elevated, and
rather distantly spaced, slightly protractively slanting axial riblets.
Following this, wrinkles make their appearance. These are arranged
in a somewhat arrow-shaped pattern extending retractively over a
third of the distance between the summit and the periphery, then
bending abruptly protractively toward the periphery. These ridges
are about as wide as the spaces that separate them and fade out at
the summit, which is appressed to the preceding turn as a ridge.
On the last third of the last turn these ridges become decidedly
irregular and gradually fade out. On the last turn there is a strong
depression a little distance below the summit. On all these ridged
whorls the fine axial riblets are well developed, but are rendered
irregular and wavy as they cross the nodules of the ridges. On
the first two turns the suture is strongly impressed ; on the succeeding
turn it is rendered less conspicuous by the appression of the summit
of the whorls to the preceding turn. Periphery well rounded. Base
strongly rounded, openly umbilicated, with a very pronounced keel
marking the outer edge of the umbilicus. This keel is separated
from the umbilical wall by a deeply impressed groove. On the out-
side it is less strongly differentiated. The posterior half of the base
is more or less smooth, while the anterior portion is marked by pro-
428349497
94 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
tractively curved, strong ridges separated by spaces as wide as the
ridges. There are also deep pits at the umbilical keel, which are
more or less regularly disposed. The umbilical keel is rendered
rough by the incremental elements and the umbilical wall is marked
by rough axial riblets and lines of growth. Aperture circular,
oblique, decidedly protracted and angulated at the posterior angle;
outer lip thin, the inner somewhat thickened. Operculum typically
ptychocochlid.
The specimen figured is one of two, U.S.N.M. No. 253665, from
Port Antonio, Portland Parish, Jamaica. It has 4.9 whorls and
measures: Height, 17.7 mm.; greater diameter, 25.2 mm.; lesser
diameter, 18.7 mm.
A large series of specimens before me all come from the north coast
of Portland, ranging from Port Antonio westward.
I have been unable to differentiate Chitty’s Cyclotus portlandensis,
even subspecifically, from this species.
PTYCHOCOCHLIS ADAMSI, new species
PLATE 14, FiaurES 4-6
Shell rather large, helicoid, the upper surface chestnut-brown; the
lower, paler. Interior of aperture soiled white. The nucleus consists
of 1.5 well-rounded smooth whorls. The postnuclear whorls are marked
by rather weak somewhat retractively curved axial riblets, which are
rather distantly spaced. Beginning with the third postnuclear whorl,
weak nodules make their appearance, which extend from the appressed
summit to the periphery in a protractive, decidedly oblique slope. This
type of sculpture extends over only about half a turn, after which the
nodulation becomes irregularly disposed and practically disappears on
the last third of the last turn. The suture of the early whorls is very
strongly impressed. On the last 1.5 turns the summit of the turns be-
comes decidedly appressed and projects over the preceding turn so as
to form a strong ridge, which renders the suture less conspicuous. On
the last whorl there is a decided groove below this appressed portion.
Periphery well rounded. Base strongly rounded, openly umbilicated,
with a very strong keel marking the outer limit of the umbilicus. This
keel is separated by a decidedly impressed groove anteriorly. Posterior
to the keel more or less regularly disposed deep pits are present, while
the spaces between them extend as oblique short cords toward the
periphery; they do not, however, reach the middle of the base. The
umbilical wall is marked by coarse riblets and lines of growth. Aper-
ture circular, oblique, protracted into a decided angle at the posterior
angle, where it joins the preceding turn as a slight shelf; peristome
thin on the outer lip, somewhat thickened on the inner. The oper-
culum is not quite typically ptychocochlid, the outer projection of the
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 95
expanded summit falling short of reaching the succeeding turns in
spots.
The type, from Portland Parish, Jamaica, is one of the C. B. Adams
collection at Amherst College. It has 5 whorls and measures: Height,
19.0 mm. ; greater diameter, 27.0 mm.; lesser diameter, 19.7 mm.
This species resembles quite closely P. corrugata (Menke) but is
easily differentiated by its much less strongly developed sculpture and
the difference in the operculum noted.
PTYCHOCOCHLIS SIMPSONI, new species
PLATE 14, Ficures 16-18
Shell of medium size, helicoid, chestnut-brown with the cord at the
summit of the last whorl buff. The nucleus consists of a little more
than a single low, rounded, smooth whorl. The postnuclear whorls
are inflated and strongly rounded; the first one is marked by very
slender, slightly retractively curved, weak axial riblets. The second
postnuclear whorl shows the beginning of oblique protractive wrinkles
which do not quite extend to the summit. This sculpture continues to
the last half of the last turn, where it becomes irregular. The entire
surface is also marked by incremental lines of more or less irregular
strength. Suture strongly impressed on the early whorls, rendered
less conspicuous on the last turn by the appressed summit. Periphery
well rounded. Base narrowly umbilicated and marked with a mod-
erately strong cord at the outer edge of the umbilicus and a series of
feeble protractively slanting ridges adjacent to the umbilical cord,
which are separated by spaces about as wide as the ridges. The rest
of the base shows an indication of incised spiral lines. The umbilical
wall is marked by rather coarse incremental lines. Aperture circular,
large, protracted into an angle at the posterior angle; outer lip
thin, the inner thickened. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 535968, was collected by Henderson and
Simpson at Bog Walk, St. Catherine Parish, Jamaica. It has 4.8
whorls and measures: Height, 15.3 mm.; greater diameter, 21.3 mm.;
lesser diameter, 15.5 mm.
Its smaller size and feebler sculpture will differentiate this species
from both P. corrugata (Menke) and P. adamsi. The species seems
to be confined to the region of Bog Walk.
PTYCHOCOCHLIS MARTENSI (Kobelt)
Puate 14, Fiaures 1-3; PLATE 42, FicuRres 8-10
1912. Neocyclotus (Plectocyclotus) martensi Kosett, Martini-Chemnitz Conchy-
lien Cabinet, vol. 1, sect. 19, pt. 3, p. 948, pl. 141, figs. 2-4.
Shell large, helicoid, chestnut-brown ; interior of the aperture chest-
nut-brown. The nucleus consists of about one well-rounded smooth
96 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
turn. Postnuclear whorls inflated, strongly rounded; the first is
marked by slender, retractively curved axial riblets, which are about
half as wide as the spaces that separate them. Axial riblets also
cover the rest of the whorls, but beyond the first turn they become
less regular and tend more toward the formation of slender lamellae.
Beginning with the second postnuclear whorl, oblique ridges make
their appearance. These ridges are decidedly protractively curved,
and are a little narrower than the spaces that separate them. They
are strongest on the early turns and gradually fade and disappear on
the middle of the spire, after which they are replaced by irregular,
low wrinklings and nodulations, as well as malleations. Suture
strongly impressed, except on the last turn, which is appressed to the
preceding whorl. Periphery well rounded. Base strongly rounded,
openly umbilicated, with a strongly elevated keel marking the outer
edge of the umbilicus. This keel is decidedly notched and roughened,
and separated from the umbilical wall by a strong groove. On the
outside it is also separated by a groove, but here the groove is weaker
than that on the umbilical side. The rest of the base is marked by
obsolete spiral lines and indications of malleations, as well as strong
incremental lines. The umbilical wall is marked by numerous in-
cremental lines, which tend toward the formation of riblets. Aper-
ture circular, protracted into an angle at the posterior angle; outer
lip of the peristome thin; inner, somewhat thickened. Operculum
typically ptychocochlid.
The specimen figured, U.S.N.M. No. 356092, was collected by John
B. Henderson at Holly Mount, St. Catherine Parish, Jamaica. It has
5 whorls and measures: Height, 19.4 mm.; greater diameter, 28.6
mm.; lesser diameter, 21.0 mm.
The species has a rather wide distribution through the northern
part of theisland. It ranges through Trelawny and St. Ann Parishes
and the northeastern part of St. Catherine Parish.
PTYCHOCOCHLIS HENDERSONI, new species
PLATE 15, FIGuRES 7-9
Shell of medium size, depressed-helicoid. The denuded specimen
white. The nucleus consists of a single well-rounded, smooth turn.
The postnuclear whorls are inflated and strongly rounded; the first
is marked by slender, feeble axial riblets; the succeeding turns in
addition to the axial riblets are marked by protractively curved
ridges, which are about as wide as the spaces that separate them and
which do not extend on the anterior fourth of the turns. These
ridges extend fairly strongly to the very end of the last whorl.
Suture strongly impressed on all but the last turn, which is appressed
to the preceding whorl. Periphery well rounded. Base broadly
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 97
openly umbilicated. The outer edge of the umbilicus is bounded by
a weak keel. ‘The base is marked by rather strong incremental lines
and a series of oblique pits, which are not profound at the outer edge
of the umbilical keel. The umbilical wall is marked by low, rather
broad axial riblets and incremental lines. Aperture circular, oblique,
protracted into an angle at the posterior angle; peristome slightly
thickened. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 535969, was collected between Fort Au-
gusta and Port Henderson in St. Catherine Parish, Jamaica. It has
4.4 whorls and measures: Height, 11.3 mm.; greater diameter, 17.1
mm.; lesser diameter, 13.2 mm.
Two additional specimens from the same locality are a trifle larger.
This species is nearest related to P. martanna, from which it can
readily be separated by its broader umbilicus.
PTYCHOCOCHLIS MARIANNA, new species
PLATE 15, FicuREs 1-3
Shell of medium size, helicoid, chestnut-brown, a trifle paler on
the underside. The nucleus consists of a single well-rounded, smooth
turn. Postnuclear whorls inflated, strongly rounded; the first is
marked by slender, retractively slanting, hairlike axial riblets, which
are as wide as the spaces that separate them. On the succeeding
turns these riblets become decidedly intensified and irregular as they
cross the ridges, nodulations, and pits. Beginning with the second
postnuclear whorl, oblique protractively slanting wrinkles make their
appearance. These ridges are separated by spaces not quite so wide
as the wrinkles. This sculpture continues to the last half of the
last turn, beyond which the wrinkles become irregularly disposed
and also decidedly irregular in size. Toward the end of this whorl
they become almost obsolete. Suture strongly impressed on the
early whorls, adnate on the last whorl. Periphery well rounded.
Base well-rounded, narrowly umblicated, the umbilicus provided with
a strong keel at its outer limit. A deeply impressed groove sepa-
rates the keel from the umbilical wall. The keel is rendered de-
cidedly rough by the incremental elements. The base is marked by
scarcely perceptible indications of nodules. It also gives one the
impression of having obsolete spiral threads. The umbilical wall
is marked by moderately strong incremental lines. Aperture cir-
cular, decidedly oblique, protracted into an angle at the posterior
angle; peristome thin. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 535970, was collected by Henderson at
Roaring River Falls, Ocho Rios, St. Ann Parish, Jamaica. It has 5
whorls and measures: Height, 17.0 mm.; greater diameter, 21.2 mm. ;
lesser diameter, 16.3 mm.
98 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The species appears to be confined to the north coast of St. Ann
and St. Mary Parishes.
PTYCHOCOCHLIS LACTEOFLUVIALIS (Pilsbry and Brown)
PLATE 15, FIGURES 46
1910. Aperostoma (Ptychocochlis) lacteofluviale Prtspry and Brown, Proc. Acad.
Nat. Sci. Philadelphia, 1910, p. 534.
Shell rather small, helicoid, covered with a chestnut-brown perios-
tracum. The nucleus consists of a single well-rounded, smooth turn.
Postnuclear whorls inflated, strongly rounded; the first marked with
mere protractively slanting incremental lines; the rest also show
incremental lines, which gradually increase in size, and form on
the last turn more or less sinuous, almost hairlike riblets. Beginning
with the second postnuclear whorl, weak, protractively slanting axial
wrinkles make their appearance, which do not extend to the pos-
terior third of the whorls and are best emphasized on the middle.
These wrinkles are about as wide as the spaces that separate them.
The wrinkles disappear on the last whorl, where indications of
spiral obsolete threads make their appearance. Suture very strongly
impressed on all but the last half of the last whorl, which is adnate.
Periphery well rounded. Base inflated, strongly rounded, openly
umbilicated, with a weakly developed cord marking the outer limit
of the umbilicus. The base has scarcely any indications of obsolete
nodules, but is marked by feeble spiral cords and incremental lines.
The umbilical wall is marked by rather coarse riblets and incremental
lines. Aperture decidedly oblique, strongly protracted to form an
angle at the posterior angle. Operculum typically ptychocochlid.
The specimen figured, U.S.N.M. No. 356037, is one of a series of
specimens collected by C. B. Taylor at Round Hill, Milk River, Claren-
don Parish, Jamaica. It has 4.7 whorls and measures: Height, 12.5
mm.; greater diameter, 16.5 mm.; lesser diameter, 13.0 mm.
The species seems to be confined to the Milk River region of south-
western Clarendon Parish.
PTYCHOCOCHLIS SAVANNENSIS, new species
PLATE 15, Fieures 13-15
Shell of medium size, depressed-helicoid, covered with a pale
chestnut-brown periostracum. The nucleus consists of a single well-
rounded, smooth whorl. The postnuclear whorls are inflated and
strongly rounded; the first is marked by slender, closely spaced, hair-
like, retractively slanting axial riblets. The second postnuclear
whorl, in addition to slender riblike incremental lines, is marked by
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 99
almost vertical axial ridges, which are separated by spaces about
twice as wide as the ridges. Beginning with the third postnuclear
whorl, the ridges assume a protractively slanting position. Here
they do not reach to the summit but terminate at the anterior third
between summit and suture. These ridges here are not quite so wide
as the spaces that separate them. On the last whorl they again dis-
appear and the whorl is marked by irregular nodules, which become
decidedly enfeebled on the last portion of this turn. Suture strongly
impressed on all but the last turn, which is adnate to the preceding
whorl. Periphery well rounded. Base strongly rounded, openly,
rather narrowly umbilicated and provided with a rather strong keel
at the outer limit of the umbilicus. The base is marked by lines of
growth and scarcely any indications of nodulations. The umbilical
wall is marked by rather rough lines of growth. Aperture circular,
protracted into an angle at the posterior angle. Operculum typically
ptychocochlid.
The type, U.S.N.M. No. 356025, is one of a large series of speci-
mens collected by Henderson at Savanna la Mar, Westmoreland
Parish, Jamaica. It has 4.5 whorls and measures: Height, 13.0 mm. ;
greater diameter, 20.2 mm.; lesser diameter, 14.5 mm.
This species is confined to the region about Savanna la Mar, that
is, the southern side of central Westmoreland Parish.
PTYCHOCOCHLIS ORCUTTI, new species
PLATE 15, FIGuRES 27-29
Shell large, helicoid, covered with a chestnut-brown periostracum,
sometimes with a light zone near the summit and the periphery.
When the periostracum is removed there is a pinkish tinge to the
shell substance of the last whorl. The nucleus consists of 1.5 well-
rounded, smooth whorls. The postnuclear whorls are inflated and
strongly rounded; the first is marked by feeble, slightly retractively
slanting, hairlike riblets, which are not so wide as the spaces that
separate them. Beginning with the second postnuclear whorls, weak,
oblique, low, protractively slanting ridges make their appearance,
which do not reach the anterior fourth between summit and suture.
These disappear shortly after the last whorl is reached, where they
are replaced by feeble, irregular nodules, which become obsolete on
the last part of the last turn. Suture moderately strongly impressed
on the early whorls; the last has the summit appressed, which char-
acteristic renders the sutrure less conspicuous. Periphery well
rounded. Base somewhat inflated, well rounded, openly, moder-
ately broadly umbilicated. The umbilicus is bounded on the out-
side by a cord, which is rendered rough by the incremental lines.
100 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The base is marked by incremental lines and appears free from nodu-
lations. The umbilical wall is marked by coarse, irregular riblets
and lines of growth. Aperture circular, decidedly oblique, with a
strongly protracted angle at the posterior angle; outer lip thin;
the inner, thickened. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 395588, was collected by Orcutt at Arntully
Estate on Stony Valley River, at an elevation of 3,000 feet in St.
Thomas Parish, Jamaica. It has 5.1 whorls and measures: Height,
20.1 mm.; greater diameter, 26.8 mm.; lesser diameter, 20.0 mm.
The species appears to range through the southern slope of the Blue
Mountains in St. Thomas Parish,
PTYCHOCOCHLIS MINOR (Chitty)
PLATE 15, Fiaures 10-12
1857. Cyclotus corrugatior minor Curtry, Proc. Zool. Soc. London, vol, 25, p. 150.
Shell of medium size, depressed-helicoid, covered with a pale brown
periostracum. The nucleus consists of a little more than a smooth
rounded whorl. The postnuclear whorls are inflated and well rounded ;
the first is marked by slender, very poorly developed axial riblets. Be-
ginning with the second, these axial riblets form wrinkles, which do
not extend to the summit of the whorls but are decidedly obliquely
protractively slanting on the posterior half of the whorl, and then
bend at an equal angle, retractively, toward the suture, that is, form-
ing an arrow-shaped angle, producing a cone-in-cone effect. The
spaces that separate these wrinkles are not quite so wide as the wrinkles.
This sculpture disappears on the last whorl, where irregularly de-
veloped and distributed nodules replace it. Suture well impressed on
the early whorls; on the last whorl the summit is appressed and the
suture becomes less conspicuous. Periphery well-rounded. Base well
rounded, rather narrowly umbilicated, with a strong keel marking the
outer edge of the umbilicus, which is decidedly pinched in at its inner
margin. The outside of the umbilical keel is rendered notched by the
irregularly developed incremental lines. The rest of the base is
marked by obsolete spiral threads. The umbilical wall bears rather
coarse, closely spaced riblets. Aperture subcircular, oblique, de-
cidedly protracted into an angle at the posterior angle; outer lip thin,
inner lip somewhat thickened. Operculum typically ptychocochlid.
The specimen figured, U.S.N.M. No. 128016, comes from Mandeville,
Manchester Parish, Jamaica. It has 4.5 whorls and measures:
Height, 12.5mm.; greater diameter, 17.8 mm. ; lesser diameter, 13.3 mm.
This species appears to range through southwestern Manchester
Parish and southeastern St. Elizabeth Parish.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 101
PTYCHOCOCHLIS MAGNA (Chitty)
PLATE 15, Figures 30-32
1857. Cyclotus corrugatior magna CHITTY, Proc. Zool. Soc. London, vol. 25, p. 150.
1912. Neocyclotus (Plectocyclotus) varians knobbei Kosett, Martini-Chemnitz
Conchylien Cabinet, vol. 1, sect. 19, pt. 3, p. 944, pl. 141, figs. 13-14.
Shell rather large, helicoid, pale brown with a rosy flush; interior
of the aperture pale brown. The nucleus consists of a single well-
rounded, smooth turn. The postnuclear whorls are decidedly in-
flated, strongly rounded; the first is marked by slender, hairlike,
slightly retractively curved axial riblets. From there on oblique, re-
tractively slanting ridges make their appearance. These ridges extend
from near the summit to about the middle of the whorl; they then
suddenly change to a decidedly oblique protracted slant, thus produc-
ing a series of more or less regular cone-in-cone shaped areas. On the
last whorl this sculpture is replaced by rather irregular wrinkling on
its first half, while the last half lacks nodulations. Suture strongly
unpressed except on the last whorl, which is appressed to the preceding
turn. Periphery strongly rounded. Base narrowly umbilicated, with
a strong keel marking the outer limit of the umbilicus. The base, as
well as the last part of the upper surface, shows weak spiral threads.
The umbilical wall is marked by rather coarse riblets and incremental
lines. Aperture circular, angulated at the posterior angle; outer lip
thin, the inner lip thicker. Operculum typically ptychocochlid.
The specimen figured, U.S.N.M. No. 535971, is one of three in our
collection labeled “Jamaica,” without specific locality. It has 5 whorls
and measures: Height, 21.0 mm.; greater diameter, 23.9 mm.; lesser
diameter, 18.4 mm.
P. magna is easily distinguished from the other members of the
genus by its extremely elevated outline.
PTYCHOCOCHLIS MONTEGOENSIS, new species
PLATE 15, Ficures 33-35
Shell moderately large, depressed-helicoid, chestnut-brown. Early
whorls with a rosy flush. The nucleus consists of a little more than a
single whorl, which is well rounded and smooth. The postnuclear
whorls are inflated, well rounded; the first half of the first postnuclear
whorl is marked by slender, slightly retractively curved axial riblets;
the second half shows the beginning of decidedly protractively curved
wrinkles, which do not quite extend to the summit, and which are about
as wide as the spaces that separate them. On the penultimate whorl
these wrinkles take a retractive slant on the posterior portion of the
whorl, then bend strongly into a protractive position, thus forming
arrow-points that give to the sculpture of this whorl a cone-in-cone
102 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
shaped effect. On the last whorl the sculpture becomes irregular and
on the last half of the last turn quite obsolete. Suture strongly im-
pressed on the early turns; on the last whorl the summit of the turn is
appressed to the preceding turn and the suture is less conspicuous.
Periphery strongly rounded. Base moderately broadly umbilicated,
with a strong keel marking the outer limit of the umbilicus. This keel
is strongly inpinched on the umbilical margin and less so on the basal
margin. It is rendered rough by the incremental lines. The base does
not have pronounced tubercles but merely here and there an indication
thereof. The umbilical wall is marked with moderately strong riblets
and incremental lines. Aperture circular, the posterior angle pro-
tracted into an angulation; outer lip of the peristome thin, the inner
slightly thickened. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 535972, was collected by Henderson at Mon-
tego Bay, St. James Parish, Jamaica. It has 4.7 whorls and measures:
Height, 15.7 mm.; greater diameter, 21.6 mm. ; lesser diameter, 16.2 mm.
The species seems to be confined to the region about Montego Bay,
where it appears to be rather abundant.
PTYCHOCOCHLIS VENDREYSI, new species
PLATE 15, FicuRES 36-38
Shell rather large, dark chestnut-brown, with a little paler zone
at the suture and a rosy tinge on the early whorls. The interior of the
shell is bright chestnut-brown with a reddish tinge, the edge of the
peristome being a little paler. The nucleus consists of a little more
than a single turn, which is well rounded and smooth. The post-
nuclear whorls are inflated and well rounded; the first turn is marked
by slender, retractively curved axial riblets; the next, in addition to
the axial riblets, which become almost lamellose on the succeeding
turns, is marked by decidedly protractively curved, low, weakly devel-
oped ridges which are separated by equally inconspicuous impressions.
On the remaining turns this sculpture becomes obsolete and the surface
is merely rendered slightly rough by a few pits and weak nodules.
The last portion of the upper surface of the last whorl bears indica-
tions of poorly developed spiral threads. Suture of the early whorls
well impressed; the last whorl has the summit appressed and the
suture thereof not so strongly emphasized. Periphery well rounded.
Base well rounded, openly umbilicated, with a strong keel at the outer
edge of the umbilicus, which is strongly inpinched on its umbilical
margin and feebly impressed on the outer edge. The base, in addi-
tion to incremental lines, has some indications of spiral threads and
an occasional little nodule. The umbilical wall is marked with axial
riblets and incremental lines. Aperture circular, decidedly oblique,
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 103
with the posterior angle produced into a decided angulation. Outer
lip of the peristome thin, the inner lip somewhat thickened. Oper-
culum typically ptychocochlid.
The type, U.S.N.M. No 535978, is one collected by Orcutt at Worthy
Park, 1 mile from Luidas Vale, St. Catherine Parish, Jamaica. It has
4.9 whorls and measures: Height, 18.9 mm.; greater diameter, 25.7
mm.; lesser diameter, 19.1 mm.
The species appears to be confined to St. Catherine Parish, where
it is rather widely distributed.
PTYCHOCOCHLIS SUBGLOBOSA, new species
PLATE 15, FiauREs 16-18
Shell of medium size, helicoid, covered with a dark chestnut-brown
periostracum. There is an indication of a lighter zone at the appressed
summit of the last whorl. The nucleus consists of 1.2 well-rounded,
smooth whorls. The postnuclear whorls are inflated and strongly
rounded; the first is marked with fine, retractively slanting, closely
spaced, hairlike riblets, which become somewhat strengthened on the
succeeding turns and appear as low lamellae on the last whorl. Begin-
ning with the second postnuclear whorl, the upper side of the turns is
marked with irregularly developed and disposed weak low nodulations,
which lend to the surface a somewhat rough and malleated aspect.
Suture strongly impressed except on the last whorl, the summit of
which is appressed to the preceding turn, which causes the suture to be
less pronounced. Periphery well rounded. Base inflated, well
rounded, openly umbilicated with a strong broad rounded keel at the
outer edge of the umbilicus. This keel is decidedly notched and has a
number of not very deep, somewhat protractively, obliquely directed
indentations outside of it. The rest of the base is slightly malleated
and shows low obsolete spiral threads. The umbilical wall is marked
by strong lines of growth. Aperture circular, protracted at the pos-
terior angle into an angulation. Operculum typically ptychocochlid.
The type, U.S.N.M. No. 398910, is one of a series of specimens col-
lected by Orcutt on the east side of Ferry River above Kingston on the
Spanish Town road, St. Andrew Parish, Jamaica. It has 5 whorls
and measures: Height, 15.7 mm.; greater diameter, 19.8 mm.; lesser
diameter, 14.3 mm.
The species ranges through St. Catherine and St. Ann Parishes.
PTYCHOCOCHLIS CAMPEACHYI (H. B. Baker)
Shell small, helicoid, covered with a chestnut-brown periostracum ;
interior of the aperture pale brown. The nucleus consists of about 1.2
well-rounded, smooth whorls. The postnuclear whorls are inflated and
104 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
strongly rounded. In one of the subspecies, P. ¢. petricola, the first
nuclear whorl is marked by very coarse, distantly spaced axial ribs.
In the typical race the axial riblets are fine and less strongly developed.
The succeeding whorls are slightly malleated, almost smooth, and
marked with lines of growth. Suture strongly impressed except on
the last turn, which is appressed to the summit. Periphery well
rounded. Base narrowly umbilicated with a limiting keel at the outer
edge of the umbilicus. The rest of the base is marked like the upper
surface of the last whorl, that is, it shows the lines of growth and
obsolete malleations and indications of spiral threads. The umbilical
wall is marked by incremental lines. Aperture subcircular, decidedly
oblique, decidedly protracted into an angle at the posterior angle;
outer lip of the peristome thin, the inner somewhat thickened. Oper-
culum typically ptychocochlid.
I am recognizing two subspecies :
KEY TO THE SUBSPECIES OF PTYCHOCOCHLIS CAMPEACHYI
Early postnuclear whorls strongly axially ribbed_------------------- petricola
Early postnuclear whorls feebly axially ribbed_----------~----~------~ campeachyi
PTYCHOCOCHLIS CAMPEACHYI PETRICOLA, new subspecies
PLATE 15, Figures 22-24
This subspecies appears confined to the southern part of St. Andrew
Parish, Jamaica. It is differentiated from P. c. campeachyi (H. B.
Baker) by having the first postnuclear whorl provided with strong
axial ribs. It is also larger than that subspecies.
The type, U.S.N.M. No. 535947, comes from Stony Hill, St. Andrew
Parish, Jamaica. It has 4.5 whorls and measures: Height, 12.8 mm.;
greater diameter, 16.6 mm. ; lesser diameter, 12.8 mm.
PTYCHOCOCHLIS CAMPEACHYI CAMPEACHYI (H. B. Baker)
PLATE 15, Figures 19-21
1934. Poteria varians campeachyi H. B. Baxer, Nautilus, vol. 48, pp. 61-62,
figs. 5, 19, 22.
This is the smaller of the two subspecies, which ranges through the
central portion of St. Catherine Parish. It is distinguished from P. c.
petricola by its smaller size and weak axial ribs on the first postnuclear
turn.
The type, Acad. Nat. Sci. Philadelphia No. 189611, was collected by
Dr. H. B. Baker near Campeachy Gully Marsh on the northwest side
of Port Henderson Hill, St. Catherine Parish, Jamaica. It has 4.8
whorls and measures: Height, 12.2 mm.; greater diameter, 15.2 mm. ;
lesser diameter, 11.3 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 105
PTYCHOCOCHLIS SUBRUGOSA (Sowerby)
Priate 15, FicuRES 25, 26
1850. Cyclostoma subrugosum Sowexrsy, 'Thes. conechyliorum, vol. 1, Supplement.
p. 161*, pl. 31B, figs. 308-309.
1852. Cyclotus subrugosus PFEIFFER, Monographia pneumonopomorum viventium,
vol. 1, p. 28.
1898. Neocyclotus (Plectocyclotus) subrugosus Kopecr and MOLLENvorrF, Nachrb.
deutschen malak. Ges., vol. 29, p. 139, reprint.
Since I have not seen this species, I quote Sowerby’s description and
reproduce his figures:
“Shell suborbicular, depressed, slightly corrugated, white; spire
depressed, obtuse, slightly prominent at the apex; volutions four,
rounded, slightly depressed posteriorly; suture distinct, marked with
a nearly obsolete line in front; aperture nearly circular; peritreme
slightly thickened posteriorly, thin on the side of the umbilicus, sepa-
rated from the last volution; umbilicus large, anteriorly keeled.
“This species is related to C. corrugatum.
“Jamaica.”
Genus POTERIA Gray
In 1850 Gray, in his “Nomenclature of Molluscus Animals and Shells
in the Collection of the British Museum,” part 1, Cyclophoridae, page
11, defines the name Poteria, which he had used in his “Synopsis of the
Contents of the British Museum,” in 1840 and 1842, as a nude name, by
stating, “whorls in front of the axis keeled. Peristome straight.”
Here he cites two species without designating a type:
27. CYCLOTUS LINEATUS [Gray].
Cyclostoma jamaicense Sow. Thes. Conch. i. 96. n. 15. t. 23. f. 12, 13., not
Gray.
Aper. jamaicense Pfeiff. 1. c. 104. n. 3.
* * * * * * *
28. CycLOTUS JAMAICENSIS [Gray].
Cyclost. corrugatum Menke, Cat. Malak. 10.; Sow. Proc. Zool. Soc. 1848, 30.,
Thes. Conch. i. 95. t. 23, f. 10, 11.; Pfeiffer, Kuster, Chemn. Conch. Cab.
DTM, Sly AO te ey eel oy Aa
Turbo jamaicensis Chemn. Conch. xi. t. 209. f. 2057, 2058.
T. (Cyclostoma) jamaicensis Gray in Wood’s cat. Supp. t. 6. f. 3.; Lister,
Conch. t. 55. f. 51.; Sloane, Jamaica, ii. t. 240. f. 84.
Aper. corrugatum Pfeiff. 1. c. 104. n. 4.
In other words, he lumps here about everything known from Jamaica
at that time except what had been segregated to form his Cyclotus
lineatus.
Baker, in 1922, Nautilus, vol. 35, pages 14, 15, resurrected the forgot-
ten name Poteria and designated Turbo jamaicensis (Chemnitz) Wood,
1828, as type. This, unfortunately, still leaves some doubt about the
106 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
actual type restriction for the genus, since 7urbo jamaicensis Chemnitz
(nonbinominal), rechristened orthodoxly Cyclostoma corrugatum in
1830 by Menke, is quite distinct from 7'urbo (Cyclostoma) jamaicensis
Gray.
Therefore, I now further restrict the type designation by selecting
Turbo jamaicensis Gray (in Wood, Index testaceologicus, Suppl., p.
18, pl. 6, fig. 3, 1828=Cyclostoma jamaicense, ibid., p. 36, same plate
and figure) as type. The synonymy is as follows:
1850. Poteria Gray, Nomenclature of molluscous animals and shells in the col-
lection of the British Museum, pt. 1, Cyclophoridae, p. 11. (Type: Cyclostoma
jamaicense Gray, in Wood.)
1898. Plectocyclotus Kopett and MOLLENDORFF, Nachrb. deutschen malak. Ges.,
vol. 28, p. 189 (reprint; not in original). Type by subsequent designation
(Pilsbry, Proc. Acad. Nat. Sci. Philadelphia, p. 5383): Cyclostoma jamaicense
Sowerby=Poteria (Poteria) lineata (Gray).
The group is characterized by the possession of a rather heavy
medium-sized shell, which has an umbilical bounding keel that varies
in different species from very strong to obsolete. The shell may be
strongly malleated or less so, or malleations may be absent. The oper-
culum bears a strongly elevated spiral lamella, which is almost at right
angles to the chondroid basal plate, and curves outward. The genus
is confined to Jamaica,
I am recognizing two subgenera, which the following key will help
to differentiate :
KEY TO THE SUBGENERA OF POTERIA
Space between the opercular lamella with retractively curved raised
threadss ee ee Oe ee ee ee ee ee Poteria
Space between the opercular lamella without retractively curved
raised) threads222- 2 Se ee a a eee Cyclobakeria
Subgenus PoTERIA Gray
For synonymy see above.
Poterias in which the broad spaces separating the turns of the la-
mella are reenforced with retractively curved raised threads.
Type: Turbo (Cyclostoma) jamaicensis Gray=Poteria (Poteria)
jamaicensis (Gray).
The radula of Poteria (Poteria) lineata (Gray) has the following
formula 8:3:3:8, and the jaw lacks distinct median projections.
The verge is situated on the back of the neck behind the tentacles.
It is swollen basally and provided with a seminal groove and a very
short simple appendage.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 107
KEY TO THE SPECIES OF SUBGENUS POTERIA
Upper surface of last whorl corrugated.
Shell helicoid.
Penultimate whorl with wrinkles.
Wrinkles and pits COarse2= 22 == ae eee burringtoni
Wrinkles and pits not coarse_—---__--_______--___-______ imitator
Penultimate whorl with axial riblets only__-__----_--_-------- lineata
Shell depressed-helicoid.
Wimbilicalkeelwstnon Ges eee eae corrugatissima
lWimbilical keeltobsoletens =e ee ees ee pallescens
Upper surface of last whorl! not corrugated.
Axialeriblecsealmost lamellae en eee jamaicensis
Axial riblets not almost lamellar.
Whorls' flattened near Summit=—-=-—__ = = plana
Wihortlsnot fattenea: near summit = 2s. een eee crassa
This key does not include Chitty’s Cyclotus nodosus, notatus, and
inutilis, specimens of which I have not seen. I have copied Chitty’s
descriptions and offer suggestions as to their relationship under
each of these species.
POTERIA (POTERIA) BURRINGTONI, new species
Puate 16, Fiaurss 31-33
Shell rather large, helicoid, covered with a chestnut-brown perios-
tracum. The nucleus consists of 1.5 inflated, well-rounded, smooth
turns. The postnuclear whorls are inflated and strongly rounded;
the first is marked by rather strong axial riblets, which are not quite
so wide as the spaces that separate them. On the succeeding turns
these axial riblets become somewhat weakened, irregular in strength
and disposition. In comparison they are here narrower and more
elevated. Beginning with the third whorl, strong axial wrinkles
make their appearance. They are at first rather irregular, but on
the first half of the last whorl they assume a decidedly obliquely
slanting retractive form. These wrinkles render the upper surface
decidedly rough. Suture well impressed, except that of the last turn,
where the summit of the whorls is appressed to the preceding turn.
Periphery well rounded and marked by the continuation of the
oblique wrinkles described for the spire. These wrinkles also extend
over the low rounded base to the umbilical keel. They become weak-
ened toward the aperture. Base openly umbilicated, with a rounded
keel marking the outer limit of the umbilicus. The umbilical wall
bears rather strong axial ribs, which are crossed by incremental lines.
These ribs look as if they might be the continuation of the oblique
wrinkles. Aperture subcircular, oblique, decidedly protracted into
an angle at the posterior angle; peristome somewhat sinuous; that of
the outer lip thin, while that of the inner lip is somewhat thickened.
Operculum typically poterid.
108 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The type of this species, the only specimen known, was collected by
Dr. H. B. Baker at Somerset, Manchester Parish, Jamaica, and bears
the Philadelphia Academy of Natural Sciences catalog number 174145.
It has 5.2 whorls and measures: Height, 17.8 mm.; greater diameter,
27.0 mm.; lesser diameter, 20.0 mm.
This is by far the most roughly corrugated species of the Jamaican
Poterias, having the largest and most distantly spaced wrinkles of all.
POTERIA (POTERIA) IMITATOR, new species
PLATE 16, Ficures 10-12
Shell of medium size, unicolor or more frequently banded with a
broad zone of dark chestnut brown above and below the periphery,
the posterior half, the band at the periphery, and the base being
brownish yellow. The nucleus consists of a single whorl, which is
very small, almost flattened, and moderately well rounded. The first
postnuclear turn is marked by fine, decidedly retractively curved axial
riblets, which are about as wide as the spaces that separate them.
These riblets on the next turn become much increased in size. They
are low, well rounded, and almost as wide as the spaces that separate
them. On the succeeding turn the axial riblets are lost sight of by
their conspicuous wrinkling and pitting. The wrinkles here branch
and anastomose and form an irregular network of meshes enclosing
well-impressed pits, which may be circular or elongate. The summit
of the turns is decidely nodulose on the last whorl. On the early post-
nuclear whorls the suture is well impressed. Periphery well rounded
and marked by the pitting described for the spire. Base well rounded,
broadly openly umbilicated with a decidedly elevated, almost lamellar
keel marking the outer edge of the umbilicus. This keel is rendered
rough by the incremental lines. The anterior third of the base between
the periphery and the umbilical keel, that is the portion usually
covered by a brown band, is smooth; the rest of the base is wrinkled
like the spire, but here the wrinkles are finer and the pits enclosed
between them a little deeper. The umbilical wall bears regular axial
riblets and fine lines of growth. Aperture subcircular, somewhat
oblique, decidedly protracted into an angle at the posterior angle and
thickened at the umbilical keel; the outer lip of the peristome is
thin, and the inner somewhat thickened. The operculum is typically
poterid.
The type, U.S.N.M. No. 535975, was collected by C. R. Orcutt at
Bogue Hill, St. Elizabeth Parish, Jamaica. It has 4.8 whorls and
measures: Height, 16.9 mm.; greater diameter, 23.6 mm.; lesser
diameter, 17.8 mm.
This species occupies the northern part of St. Elizabeth Parish
and the adjacent areas of Manchester Parish,
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 109
At first glance one might consider this to be Aperostoma (Cycla-
damsia) seminudum (C. B. Adams), as far as the general shape and
color scheme are concerned, but it can at once be distinguished by
its poterid operculum.
POTERIA (POTERIA) LINEATA (Gray)
Shell varying from helicoid to decidedly elevated helicoid in out-
line, unicolor or banded; periostracum ranging from brownish buff
to chestnut colored. The nucleus consists of 1.5 decidedly elevated,
inflated, strongly rounded, smooth whorls. The first 2 postnuclear
whorls are marked by slender, very regular, retractively curved axial
riblets; beyond this the axial riblets become less regular and on the
last whorl are mere elevated incremental lines. On the last whorl
wrinkles make their appearance on the upper surface; they are of
irregular size and irregularly disposed. Here, too, spiral threads
are present. The suture of the early whorls is well impressed; on
the last two the summit of the whorls is appressed to the preceding
turn, and the suture becomes less conspicuous. Periphery well
rounded. Base well rounded, openly umbilicated, with a strong keel
marking the outer edge of the umbilicus. The base, also, is marked by
irregular wrinkles, but here they assume a more spiral arrangement.
The deepest pits are next to the umbilical keel. The umbilical wall
is marked by low rounded axial ribs, which, as well as their inter-
spaces, are crossed by incremental lines. Aperture subcircular,
oblique, decidedly protracted into an angulation at the posterior
angle; the outer lip of the peristome is thin, the inner, thickened,
particularly so at the umbilical keel. Operculum typically poterid.
I am recognizing two subspecies, which the following key and
descriptions will help to differentiate :
KEY TO THE SUBSPECIES OF POTERIA (POTERIA) LINEATA
Hermht. more’ than! 20 mmUU ur 202) te see aa lineata
Persntilessithanyt ¢ minister ee Der eee eee es See cycloata
POTERIA (POTERIA) LINEATA LINEATA (Gray)
PLATE 16, Ficures 34-36; PLATE 42, Figures 14, 15
1843. Cyclostoma jamaicense SowrerBy, Thesaurus conchyliorum, vol. 1, p. 96,
No. 15, pl. 23, fig. 12 (not Cyclostoma jamaicense Gray, 1828).
1850. Cyclotus (Poteria) lineatus Gray, Nomenclature of the molluscous animals
and shells in the collection of the British Museum, pt. 1, Cyclophoridae,
Dina:
This subspecies appears to be the most abundant race in Jamaica.
It is rather widely distributed. We have specimens from Westmore-
land, St. James, St. Elizabeth, Manchester, St. Ann, and Clarendon
42834942 8
110 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Parishes. Its larger size and more elevated form will readily dis-
tinguish it from its more western relative.
The specimen figured, U.S.N.M. No. 375939, was collected by Orcutt
at Carisbrook, St. Elizabeth Parish. It has 5.5 whorls and measures:
Height, 21.0 mm.; greater diameter, 26.6 mm.; lesser diameter,
20.3 mm.
POTERIA (POTERIA) LINEATA CYCLOATA (Chitty)
PLATE 16, FiacurEs 7-9
1857. Cyclotus cyclodtus Curtry, Proc. Zool. Soc. London, vol. 25, p. 148.
1898. Neocyclotus (Plectocyclotus) cycloatus Kopett and MOLLENpDoRFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 1388, reprint.
This subspecies appears to be confined to the southwestern part
of Westmoreland Parish. It is distinguished from the typical P.
(P.) l. lineata (Gray) in being smaller and less elevated, with a
weaker umbilical keel, and in having the last whorl usually not so
strongly appressed to the preceding turn.
The specimen figured, U.S.N.M. No. 356017, is one of a large series
collected by Henderson at Mount Pleasant, Westmoreland Parish,
Jamaica. It has 5.2 whorls and measures: Height, 16.0 mm.; greater
diameter, 23.1 mm.; lesser diameter, 17.3 mm.
POTERIA (POTERIA) CORRUGATISSIMA (Chitty)
PLATE 16, Figures 37-39
1857. Cyclotus corrugatissimus Currry, Proc. Zool. Soc. London, vol. 25, p. 155.
1898. Neocyclotus (Plectocyclotus) corrugatissimus KosBeLT and MOLLENDORFF,
Nachrb. deutschen malak. Ges., vol. 29, p. 1388, reprint.
Shell large, very depressed-helicoid, covered by a chestnut-brown
periostracum. The nucleus consists of a little more than one moder-
ately rounded smooth whorl. The postnuclear whorls are strongly
rounded, slightly flattened toward the summit. The first 2.5 are
marked by strong, retractively curved axial riblets, which are sep-
arated by spaces about 1.5 times as wide as the riblets. Beyond this
wrinkles make their appearance. These consist of two series: Strong
wrinkles, which extend from the summit toward the periphery, being
strongest at the summit and fading ventrally, having a protractive
slant; these are crossed by even more oblique, less strong wrinkles,
which are decidedly retractively slanting, the combination forming a
peculiar pattern. Suture strongly impressed on the early whorls,
less conspicuous on the last, where the summit of the whorl is strongly
appressed to the preceding turn. Periphery marked by the combina-
tion of the sculpture described for the spire. Base moderately
rounded, very widely openly umbilicated, with a strong keel marking
THH CYCLOPHORID LAND MOLLUSKS OF AMERTCA 111
the outer edge of the umbilicus. This keel on the first half of the last
whorl consists of oblique, protractive strong short ridges, which on
the last half disappear and give place to rough corrugations. Border-
ing the basal keel, on the outside, is a series of deep pits. The rest
of the base is marked by more or less continuous wrinkles, which have
a slightly oblique trend, being almost spiral in disposition. The um-
bilical wall is covered by strong axial ribs and fine incremental lines.
Aperture subcircular, decidedly oblique, with a conspicuous pro-
tracted angle at the posterior angle; outer lip of the peristome thin,
the inner thickened, particularly so at the basal keel. Operculum
typically poterid.
The specimen figured, U.S.N.M. No. 397856, was collected by Orcutt
5 miles from Black River on the road to Newmarket, St. Elizabeth
Parish, Jamaica. It has 5.25 whorls and measures: Height, 16.6 mm. ;
greater diameter, 26 mm.; lesser diameter, 19.4 mm.
The species appears to be restricted to southwestern St. Elizabeth
Parish, although we have it also from one station in Westmoreland
Parish, just across the border. In shape and in the sculpture of
the upper surface P. (P.) corrugatissima resembles most nearly
P. (P.) pailescens (C. B. Adams), from which it can readily be dis-
tinguished by its much stronger basal keel and much stronger basal
sculpture.
POTERIA (POTERIA) PALLESCENS (C. B. Adams)
PLATE 16, Fieures 28-30
1851. Cyclostoma pallescens C. B. ApAms, Ann. Lyc. Nat. Hist. New York, vol.
5, p. 59.
1852. oti pallescens P¥rIrFER, Monographia pneumonopomorum viventium,
vol. 1, p. 27.
1898. Neocyclotus (Plectocyclotus) pallescens Koper and MOuLenporrr, Nachrb.
deutschen Malak. Ges., vol. 29, p. 138, reprint.
Shell large, decidedly depressed-helicoid, almost planorboid, cov-
ered with a dark chestnut-brown periostracum. The nucleus consists
of about 1.5 moderately elevated, well-rounded, smooth turns. The
first postnuclear whorl is marked by weak, rather elevated axial rib-
lets, which are about as wide as the spaces that separate them. On
the succeeding turn these axial riblets become less regular, and wrin-
kles make their appearance on the upper surface. On the first part
of the last turn these wrinkles have both a decidedly oblique
protractive and retractive arrangement. On the last half of the
last whorl they are more inclined toward a protractive arrange-
ment. These wrinkles do not quite extend to the summit of the whorls
but begin at the anterior termination of the anterior fourth between
the summit and suture, the summit being marked by rough lines of
growth only. Suture strongly impressed, even on the last whorl.
112 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Periphery well rounded. Base very broadly openly umbilicated, with
a very weak cord marking the outer limit of the umbilicus. The sculp-
ture of the base between the periphery and this cord consists of
obsolete wrinkles, which tend toward spiral disposition. The um-
bilical area is marked by coarse ribs and fine incremental lines. Aper-
ture subcircular, somewhat effused at the junction of the outer and
basal lip, protracted into an angle at the posterior angle; outer
lip thin, the inner somewhat thickened. Operculum typically poterid.
The specimen figured, U.S.N.M. No. 356125, was received from C. B.
Adams, bearing the label “Jamaica” without definite locality. It
has 4.9 whorls and measures: Height, 16.7 m..; greater diameter, 27.2
mm.; lesser diameter, 19.8 mm.
The species seems to be confined to northeastern St. Elizabeth Par-
ish, Jamaica. In general shape it recalls P. (P.) corrugatissema
(Chitty), but the coarseness of the wrinkles on the upper surface and
their arrangement in a protractive and retractive series will at once
distinguish it. It can also be differentiated from corrugatissima by
the absence of the strong umbilical keel and in having the basal sculp-
ture also much less strongly developed.
POTERIA (POTERIA) JAMAICENSIS (Gray)
PLATE 16, FicuREs 4-6
1828. Cyclostoma jamaicense GRAY, in Wood, Index testaceologicus, ed. 2, Suppl.,
p. 36, pl. 6, fig. 3 (not Turbo jamaicensis Chemnitz, 1795, nonbinominal ; or
Cyclostoma jamaicense Sowerby, 1843).
1857. Cyclotus novussaltus Cuitty, Proc. Zool. Soc. London, vol. 25, p. 154.
Shell moderately large, helicoid, covered with an elevated brownish
periostracum, which is rendered spotted by wearing of the axial riblets.
The nucleus consists of a little more than a single well-rounded, mod-
erately elevated turn. The postnuclear whorls are marked by a strong
retractively curved axial riblets, which assume the strength of low,
appressed, retractively slanting Jamellae on the last turn. Suture well
impressed on all but the last whorl, which is appressed to the preced-
ing turn. Periphery well rounded, crossed by the axial ribs. Base
somewhat inflated, well rounded, openly umbilicated, with a weak
broad cord at the outer edge of the umbilicus. The base is marked by
the continuation of the axial sculpture. The umbilical wall is marked
by rather strong, distantly spaced axial ribs and fine incremental lines.
Aperture circular, decidedly protracted at the posterior angle; peri-
stome thin all around. Operculum typically poterid.
The specimen figured, U.S.N.M. No. 356077, is one of three received
from C. B. Adams. It has 5.2 whorls and measures: Height, 17.0
mm.; greater diameter, 24.7 mm.; lesser diameter, 19.0 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 113
The species appears confined to the southern coast of St. Elizabeth
Parish, Jamaica. It is readily distinguished from the other non-
corrugated Poterias by its strong lamellalike axial sculpture.
POTERIA (POTERIA) PLANA, new species
PLatTE 16, FIGURES 25-27
Shell small, helicoid, covered with a pale-brown periotracum. The
nucleus consists of a single, moderately elevated, well-rounded, smooth
turn. The succeeding turns are marked by slender, retractively curved
axial riblets, which are inclined to waviness on the last turn, where in-
dications of spiral threads are present. The postnuclear whorls are
flattened on the posterior fourth, anterior to the summit. Suture
strongly impressed, a little less so on the last whorl. Periphery well
rounded and marked by the continuation of the axial sculpture. Base
well rounded, openly umbilicated, with an obsolete cord at the periph-
ery. The base is marked by a continuation of the axial riblets and
indications of obselete spiral threads. The umbilical wall bears
slender riblets. Aperture circular, oblique; peristome thin. Oper-
culum typically poterid.
The type, U.S.N.M. No. 535976, was collected by Henderson at
Montpelier, St. James Parish, Jamaica. It has 4.9 whorls and meas-
ures: Height, 12.5 mm.; greater diameter, 19.2 mm.; lesser diameter,
14.0 mm.
The species appears to be rather widely distributed. The specimens
before me come from Westmoreland, Hanover, St. James, St. Eliza-
beth, and Trelawny Parishes. It resembles most nearly P. (P.) crassa
(C. B. Adams), from which the flattening at the summit will readily
distinguish it.
POTERIA (POTERIA) CRASSA (C. B. Adams)
PLATE 16, FicuREs 16-18
1851. Cyclostoma crassum C. B. Apams, Ann. Lyc. Nat. Hist. New York, vol.
5, p. 59.
1852. Cyclotus crassus Preirrer, Monographia pneumonopomorum viventium,
vol. 1, p. 29.
1855. Cyclotus (Aperostoma) crassus H. and A. ADAMS, The genera of recent
Mollusea, vol. 2, p. 275.
1898. Neocyclotus (Plectocyclotus) crassus KoseLt and MOLLENDorFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138, reprint.
Shell helicoid, small, brownish horn colored. The nucleus con-
sists of a single well-rounded, rather elevated, smooth turn. The
postnuclear whorls are marked by rather strong, retractively curved
axial riblets, which are almost as wide as the spaces that separate
them. These riblets become less strong and less regular on the
114 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
last turn. Suture well impressed on all the whorls except the last
part of the last whorl, which is appressed at the summit. Pe-
riphery well rounded. Base strongly rounded, openly umbilicated,
with an obsolete cord at the outer edge of the umbilicus, marked
by the weak continuation of the axial riblets, which also extend over
the umbilical wall. Aperture subcircular, decidedly protracted at
the posterior angle into an angle; outer lip of the peristome thin,
the inner lip somewhat thickened. Operculum typically poterid.
We have figured Adams’ type, which is in the collection at Am-
herst College. It has 5 whorls and measures: Height, 14.0 mm.;
greater diameter, 19.0 mm.; lesser diameter, 14.9 mm., and is said to
come from the interior of Manchester Parish, Jamaica.
The specimen resembles P. (P.) plana but is easily distinguished
from that by the absence of flattening at the summit.
POTERIA (POTERIA) NODOSA (Chitty)
1857. Cyclotus nodosus Cutty, Proc. Zool. Soc. London, vol. 25, p. 157.
1898. Neocyclotus (Plectocyclotus) nodosus KoBetT and MO.LLENpDoRFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 188, reprint.
I have not seen specimens referable to this species, so quote
Chitty’s description. He says that it came from Maroon Town, St.
James Parish, Jamaica.
“Form, more depressed-conic. Colour, white, with light brown
epidermis, brown at lip and on operculum. Sculpture, lines of
growth, remarkably knotted corrugation on last whorl, pitted deep
about umbilical keel, and coarse lines of growth within. Spire,
depressed with rather straight outlines. Whorls, 54, well rounded,
with deep suture. Aperture, rather oblique to the left below. Peri-
treme, slightly sinuate above. Umbilicus, wide, greatest breadth 0.3
[7.5 mm.], funnel-shaped. Umbilical keel, not wide, but well pro-
duced. Ovperculum, small, strong, sharp-edged lamina of 5 or 6
wide-apart whorls, rising almost equally from the flat plane, end
of last thickened. Height 0.6 [15 mm.], greatest breadth 0.85 [21.25
mm.], least breadth 0.67 [16.75 mm.].”
This appears nearest to P. (P.) corrugatissima (Chitty), which
occupies a range much to the south.
POTERIA (POTERIA) NOTATA (Chitty)
1857. Cyclotus notatus Cuitry, Proc. Zool. Soc. London, vol. 25, p. 148.
1898. Neocyclotus (Plectocyclotus) notatus Kosetr and MOLLENDorFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 188, reprint.
I have not seen specimens referable to this species, so quote Chitty’s
description. He says it came from Trelawny, Jamaica.
“Sculpture, fine and almost smooth, with a slight corrugation on
the upper part of latter half of the last whorl, which is devoid of
THB CYCLOPHORID LAND MOLLUSKS OF AMERICA 115
depression in its upper part. Umbilicus, greatest breadth 0.17 [4.25
mm.], least breadth 0.16 [4.0 mm.]. Umbilical keel, almost obsolete.
Operculum, character of C. corrugatus, more concave than preceding ;
spiral lamina, about 8 whorls, upper margin scarcely reflected.
Height 0.58 [14.5 mm.], greatest breadth 0.64 [16.0 mm.], least breadth
0.56 [14.0 mm.].”
This may be a small subglobular race of P. lineata (Gray),
occupying a range to the northeast of the typical form.
POTERIA (POTERIA) INUTILIS (Chitty)
1857. Cyclotus inutilis Curry, Proc. Zool. Soc. London, vol. 25, p. 155.
1898. Neocyclotus (Plectocyclotus) inutilis Kosetr and MO6tienporrr, Nachrb.
deutschen malak. Ges., vol. 29, p. 188, reprint.
As I have not seen specimens referable to this species, I quote
Chitty’s description :
“Horm, much depressed-conic. Colour, light brown; apex red-
brown. Sculpture, fine lines of growth on the 314 whorls, thence
(for its size) roughly corrugated, except the last quarter of last
whorl, where, above, lines of growth are strong, with slight pitting;
lines of growth fine within the umbilicus. Spzre, much depressed,
with convex outlines. Whorls, 4, well rounded, with very deep
suture. Aperture, oblique from left to right below, dilated above
and much expanded to the right, horizontally elliptical, 0.22 [5.5
mm.], 0.2 [5.0 mm.] vertically. Peritreme, rather sinuate above,
and detached from the penult whorl. Umbilicus, open and large,
greatest breadth 0.14 [3.5 mm.]. Umbilical keel, well defined inside
umbilicus. Operculum, of C. Jamaicensis type, concave exteriorly,
with lamella of 5 whorls, like C. crassus. Height 0.31 [7.75 mm.],
greatest breadth 0.46 [11.5 mm.], least breadth 0.35 [8.75 mm.]”
This species seems to be allied to P. lineata, but because of its
minute size it cannot be that species unless it should prove to be a
dwarf of that form.
CYCLOBAKERIA, new subgenus
Poterias in which the spaces separating the turns of the lamella are
not reenforced by raised threads.
Type: Cyclotus novae-spet Chitty =Poteria (Cyclobakeria) novae-
spet (Chitty).
The distribution of this subgenus is peculiar, being confined to the
two ends of the Island of Jamaica. Slight differences in the oper-
culum of the species from the two regions may indicate that further
division of this subgenus may be necessary. I am leaving the deter-
mination of this to the anatomists who may study the group.
116 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The radula of Poteria (Cyclobakeria) nana has the following for-
mula: 8:3:3:38, and the jaw lacks distinct median projections. The
verge is situated on the back of the neck behind the tentacles. It is
swollen basally and provided with a seminal groove and a very shert
simple terminal appendage.
KEY TO THE SPECIES OF SUBGENUS CYCLOBAKERIA
Zigzag, sculpture: absent 2242) bb ee ee eee ee tryoniana
Zigzag sculpture present.
Zigzag sculpture on last whorl feeble.
Zigzag sculpture on base pronounced___——----—~-~---_---_.-= novaespei
Zigzag sculpture on base not pronounced_--------__--__-_--_- welchi
Zigzag sculpture on last whorl not feeble.
Basal keel strong.
Last part of base outside of keel with protractively
curved cords.
Shell large, greater diameter more than 25 mm-_--___ magister
Shell smaller, greater diameter less than 18 mm_-__--___ nana
Last part of base outside of keel without protractively
curved Cords. 42a Sie Sees eee dentistigmata
Basal keel feeble.
Lamella of operculum very broad, almost touching suc-
ceeding whorlos222 = 2) 2s ee es ie ee chittyi
Lamella of operculum not very broad, not touching suc-
ceeding whorl.
Whorls slightly concave below summit___---__---~-~- notatior
Whorls not slightly concave below summit.
Spire yhelicoide 2223 2 ee eee yallahsensis
Spire depressed-helicoid_____--___--------_---- balnearis
POTERIA (CYCLOBAKERIA) TRYONIANA (Pilsbry and Brown)
PLATE 16, FIGURES 22-24
1910. Aperostoma (Ptychocochlis) tryonianum Pitspry and Brown, Proc. Acad.
Nat. Sci. Philadelphia, 1910, pp. 534-535.
Shell small, helicoid, covered with a golden-brown periostracum.
The nucleus consists of a little more than 1.5 well-rounded, smooth
turns. The postnuclear whorls are marked by slender, retractively
curved, well-raised, rather distantly spaced axial riblets. On the
last turn these riblets become less regular in size and spacing.
Suture strongly constricted in all the whorls. Periphery well
rounded, base strongly rounded, moderately broadly openly umbili-
cated, with a mere suggestion of a limiting keel at the outer edge of
the umbilicus. The base and the umbilical wall are marked by the
continuation of the axial riblets. Aperture circular, decidedly
oblique, very slightly angulated at the posterior angle; peristome thin
all around. Operculum typically cyclobakerid.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 117
The unique type, Acad. Nat. Sci., Philadelphia, No. 101141, is
without specific locality, bearing merely the label “Jamaica.” It has
4.5 whorls and measures: Height, 9.7 mm.; greater diameter, 14.0
mm.; lesser diameter, 10.9 mm.
This species is easily separated from all the other Cyclobakerias as
it has none of the zigzag sculpture characterizing the group.
POTERIA (CYCLOBAKERIA) NOVAESPEI (Chitty)
PLATE 16, Ficures 138-15
1857. Cyclotus novae-spei CuiTry, Proc. Zool. Soc. London, vol. 25, p. 148.
1898. Neocyclotus (Plectocyclotus) novae-spei KoBELT and MOLLENDOoRFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138, reprint.
Shell of medium size, helicoid, covered by a chestnut-brown perio-
stracum. The nucleus consists of a single well-rounded, smooth turn.
The postnuclear whorls are inflated, strongly rounded; the first 2
postnuclear whorls are marked by feeble, hairlike, slightly retrac-
tively curved axial riblets. After this the axial riblets become less
regular and wavy, owing to the additional sculpture. Beginning
with the third postnuclear whorl irregular wrinkles make their
appearance. These are most strongly emphasized near the suture
and weakened toward the middle of the turn. There are also indica-
tions of spiral threads and pits. The suture on all but the last whorl
is strongly impressed; on the last it is rendered less conspicuous by
the fact that the summit of this whorl creeps up materially on the
preceding turn, to which it is appressed. Periphery well rounded.
Base inflated, strongly rounded, openly, moderately broadly umbili-
cated with a very strong lamellar keel at the outer limit of the um-
bilicus. The space immediately below this keel, within the umbilicus,
is strongly excavated, while on the outside there is a strong impres-
sion. The base itself is marked by low zigzag wrinkles, which are
strongest near the umbilical keel and fade toward the periphery. At
the umbilical keel the spaces between these ridges are deeply im-
pressed pits. The umbilical wall is marked by obsolete riblets and
numerous hairlike incremental lines. Aperture circular, protracted
into an angle at the posterior angle; the outer lip of the peris-
tome thin, the inner somewhat thickened. Operculum typically
cyclobakerid.
The specimen figured, Acad. Nat. Sci. Philadelphia, No. 174154, is
one of three collected by H. B. Baker at Negril Hills, southwest of
Retreat, Westmoreland Parish, Jamaica. It has 5 whorls and
measures: Height, 16.2 mm.; greater diameter, 22.1 mm.; lesser
diameter, 16.3 mm.
The species is confined to the southwestern coastal region of
Westmoreland Parish.
118 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
POTERIA (CYCLOBAKERIA) WELCHI, new species
Shell of medium size, helicoid, covered with a chestnut-brown
periostracum. The nucleus consists of a single well-rounded, smooth
turn. The postnuclear whorls are marked by slender, rather closely
approximated, slightly retractively slanting, hairlike riblets, which
become decidedly irregular on the last whorl. Beginning with the
third whorl rough wrinkles make their appearance on the upper
surface of the whorls. These are strongest near the summit and extend
protractively slanting toward the periphery, which they do not quite
reach. The spaces separating these ridges are almost as wide as the
ridges. Suture well impressed on all but the last turn. Here the
summit is appressed and renders the suture less conspicuous. Peri-
phery well rounded. Base moderately broadly umbilicated, well-
rounded, and provided with a strong keel at the outer edge of the
umbilicus. On the umbilical side of this keel the shell is decidedly
excavated. From the umbilical keel protractively curved ridges
radiate toward the periphery. These are very pronounced in the
typical race and feebly expressed in the subspecies P. (C.) welchi
taylor. The posterior portion of the base is smooth, except for indi-
cations of spiral lines. Aperture circular, decidedly oblique, pro-
tracted into an angle at the posterior angle. Operculum typically
cyclobakerid.
I am recognizing two subspecies, which the following key and
descriptions will help to differentiate :
KEY TO THE SUBSPECIES OF POTERIA (CYCLOBAKERIA) WELCHI
Pits outside: of umbilical (keelestrong ee ee eee welchi
Pitsioutside of umbilical keel nottstron ge ess ee eee taylori
POTERIA (CYCLOBAKERIA) WELCHI WELCHI, new subspecies
PLATE 16, Ficures 1-3
The typical race was collected by Welch on Dolphin Mountain,
Jamaica, at an altitude of 1,200-1,450 feet. This mountain is located
in the central western portion of Westmoreland Parish.
This subspecies differs from P. (C.) w. taylori in having the pits
outside of the umbilical keel on the base very deep and the ridges
bordering these very pronounced. Both of these characters are
absent in taylor2.
The type, U.S.N.M. No. 535977, from Dolphin Mountain, Jamaica,
has 5.1 whorls and measures: Height, 20.0 mm.; greater diameter,
26.0 mm.; lesser diameter, 18.9 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 119
POTERIA (CYCLOBAKERIA) WELCHI TAYLORI, new subspecies
PLAtr 17, Ficures 37-39
This race was collected by C. B. Taylor and John B. Henderson
in the Great Valley, Hanover Parish, Jamaica. It is easily differenti-
ated from P. (C.) w. welchi by having the upper surface of the last
whorl obsoletely nodulose on its anterior half, and by having the
pits posterior to the umbilical keel very poorly indicated, and the
radiating ridges bordering the pits almost absent.
The type, U.S.N.M. No. 535978, comes from Great Valley, Hanover
Parish, Jamaica. It has 4.9 whorls and measures: Height, 16.0 mm. ;
greater diameter, 23.9 mm.; lesser diameter, 17.1 mm.
POTERIA (CYCLOBAKERIA) MAGISTER, new name
PLATE 17, Figures 4042
1857. Cyclotus corrugatus Cuirry, Proc. Zool. Soc. London, vol. 25, p. 146 (not
Cyclostoma corrugatum Menke, Synopsis methodica molluscorum, p. 39,
1830).
Shell very large, helicoid, wood brown, a little paler on the early
whorls. The nucleus consists of a single well-rounded, smooth turn.
The postnuclear whorls are inflated, strongly rounded, and marked
by retractively curved axial riblets, which grow stronger as the shell
increases in size and less regular, owing to the nodulations. Begin-
ning with the fourth whorl, oblique, protractively slanting axial
cords make their appearance. These fade out toward the summit
and assume a zigzag course anteriorly ; they become decidedly weaker
on the last one-tenth of the last turn. Suture strongly impressed on
all the whorls, except the last, which is decidedly appressed to the
preceding turn, thus weakening the suture. Periphery well rounded,
crossed by the axial protractive cords. Base inflated, strongly
rounded, openly moderately broadly umblicated, with a strong keel at
the outer edge of the umbilicus. The keel is strongly inpinched on
the umbilical side and less so on the basal side. Radiating from the
umbilical keel a series of strong cords about as wide as the spaces that
separate them extends over the anterior fourth of the base and fades
out toward its center. The posterior portion of the base shows the
feeble continuation of the zigzag cords of the spire. The umbilical
wall is marked by rough, closely spaced incremental threads. Aper-
ture subcircular, protracted at the posterior angle into a decided
point; outer lip thin, the inner thickened. Operculum typically
cyclobakerid.
The type is one of six specimens in the Amherst College collection,
labeled No. 7, collected by Chitty in Portland Parish, Jamaica. It
120 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
has 5.1 whorls and measures: Height, 22.7 mm.; greater diameter, 31.0
mm.; lesser diameter, 23.1 mm.
The large size will readily distinguish this species from all the
other known Cyclobakerias.
POTERIA (CYCLOBAKERIA) NANA, new species
PLATE 16, FiacurES 19-21
Shell quite small, helicoid, covered with a chestnut-brown peri-
ostracum. The nucleus consists of a single well-rounded, smooth
turn. The postnuclear whorls are inflated, strongly rounded, and
marked by slightly retractively slanting, closely spaced axial riblets,
which at first are hairlike, and which on the last whorl become almost
sublamellar. Beginning with the fourth whorl, protractive zigzag
ridges make their appearance, which fade out toward the summit.
The last whorl is decidedly appressed, and the portion that extends
up on the preceding turn has a constriction bounding it which makes
it appear cordlike. Suture strongly impressed, less so on the last
turn on account of the appressed summit. Periphery well rounded
and marked by the continuation of the zigzag cords. Base inflated,
strongly rounded, rather narrowly umbilicated, and marked by strong
protractive cords which are about as wide as the spaces that sepa-
rate them and radiate from the umbilical keel toward the base. The
posterior portion of the base is marked by the feeble continuation
of the sculpture of the upper portion of the whorl. It also shows
indications of spiral threads. The umbilical wall is marked by
feeble riblets and numerous threadlike incremental lines. Aperture
circular, oblique, decidedly protracted into an angle at the posterior
angle; peristome of the outer lip protracted, thin; that of the inner
lip somewhat thickened. Operculum typically cyclobakerid.
The type, U.S.N.M. No. 535979, is one of a series collected by
Orcutt at Cousins Cove, Hanover Parish, Jamaica. It has 4.5 whorls
and measures: Height, 11.8 mm.; greater diameter, 17.0 mm.; lesser
diameter, 12.6 mm. The species appears to be restricted to the north-
west coast of Hanover Parish.
POTERIA (CYCLOBAKERIA) DENTISTIGMATA (Chitty)
PLATE 17, Figures 34-36; PLATE 42, Figures 1-3
1857. Cyclotus dentistigmatus Cuirry, Proc. Zool. Soe. London, vol. 25, p. 154.
1898. Neocyclotus (Plectocyclotus) dentistigmatus, KoBeLtr and MOLLENDORFF,
Nachrb. deutschen malak. Ges., vol. 29, p. 138, reprint.
Shell of medium size, helicoid, covered with a chestnut-brown
periostracum ; nuclear whorls scarlet red. ‘The nucleus consists of 1.5
inflated, well-rounded, smooth turns. The postnuclear whorls are
THB CYCLOPHORID LAND MOLLUSKS OF AMERICA Al
inflated, strongly rounded, and marked by rather coarse axial riblets.
In addition to this, beginning with the second whorl, rugations make
their appearance, which gradually increase in size. These ridges are
less pronounced on the summit and grow stronger toward the periph-
ery. They have a zigzag protractive slant. Suture well impressed, a
little less so on the last turn, which is appressed to the preceding
whorl. Periphery well rounded, marked by the feeble continuation
of the axial sculpture. Base strongly inflated, strongly rounded,
with a strong keel marking the outer edge of the moderately large
umbilicus. The inner edge of this keel is decidedly inpinched, while
the outer is bounded by a series of deep pits which give the shell
the appearance of having been bitten by pointed teeth, hence the
name. The spaces between these pits form cords which are poorly
developed, short, and protractively bent. These cords disappear on
the last portion of the last turn. The posterior portion of the base
is without nodules. The umbilical wall is marked by rather pro-
nounced axial riblets. Aperture subcircular, decidedly oblique, pro-
tracted into an angle at the posterior angle; the outer lip of the peri-
stome is thin, the inner somewhat thickened. Operculum typically
cyclobakerid.
The specimen figured, U.S.N.M. No. 401305, is one of a very large
series collected by C. R. Orcutt 214 miles east of Bath, St. Thomas
Parish, Jamaica. It has 4.9 whorls and measures: Height, 16.2 mm. ;
greater diameter, 25.0 mm.; lesser diameter, 17.9 mm.
The species appears to extend over the entire southeastern portion
of St. Thomas Parish.
POTERIA (CYCLOBAKERIA) CHITTYI, new species
PLATE 17, FiaurEs 4-6
Shell small, helicoid, chestnut-brown; nuclear whorls paler. The
nucleus consists of a single well-rounded, smooth turn. The post-
nuclear whorls are inflated, strongly rounded, and marked by hair-
like, retractively slanting axial riblets, which increase in strength as
the shell progresses in growth. Beginning with the third postnu-
clear whorl feeble axial riblets make their appearance, which increase
in strength up to the last two-tenths of the last whorl, when they
again fade out. These axial cords are arranged in a more or less
zigzag pattern. Below the summit of the last turn, particularly
toward the aperture, there is a slightly concave area. The last part
of the last turn is marked by slender spiral threads. Suture of all
but the last whorl strongly impressed; on this it is rendered less
conspicuous by the appressed summit of the last turn. Periphery
well rounded. Base strongly inflated, well rounded, rather narrowly
openly umbilicated, with a feeble cord marking the outer edge of the
122 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
umbilicus. The base is marked by obsolete nodulations and pittings.
The umbilical wall is marked by numerous, irregular incremental
lines. Aperture subcircular, protracted into an acute angle at the
posterior angle; outer lip of peristome thin, the inner somewhat
thickened. The operculum has the outer edge of the lamella decidedly
expanded, approximating that of the succeeding turn.
The type, U.S.N.M. No. 378167, is one of a large series of almost
all dead shells, collected by Orcutt at the western end of the Essing-
ton Bridge, St. Thomas Parish, Jamaica. It has 4.6 whorls and
measures: Height, 14.0 mm.; greater diameter, 20.0 mm.; lesser diam-
eter, 15.2 mm.
Charles R. Orcutt also collected specimens near Yallahs Bay, St.
Thomas Parish.
POTERIA (CYCLOBAKERIA) NOTATIOR (Chitty)
PLATE 17, FicuREs 13-15
1857. Cyclotus notatior Curry, Proc. Zool. Soc. London, vol. 25, p. 148.
1299, Neocyclotus (Plectocyclotus) notatior Kosetr and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 31, p. 186.
Shell of medium size, rather elevated helicoid, wood brown. The
nucleus consists of a little more than a single low, rounded turn. The
postnuclear whorls are decidedly inflated and marked by retractively
curved, hairlike, closely spaced axial riblets, which become irregu-
larly disposed on the nodulose whorls. Beginning with the fourtb
postnuclear whorl, feeble nodulations make their appearance, which
increase in size as the shell advances in growth. On the last whorl
they assume first a retractive slant on the posterior portion near
the summit, then they make a sudden protractive bend, but owing
to the irregular development of these cords they have a zigzag ap-
pearance. These cords are not quite so wide as the spaces that
separate them. On the last two-tenths of the last whorl they again
fade out. Suture strongly impressed on all but the last whorl, which
is decidedly appressed to the preceding turn. There is an impressed
area below this appressed portion which renders that part of the
shell slightly concave. Periphery well rounded and marked by the
feeble continuation of the axial sculpture. Base inflated, strongly
rounded, very narrowly umbilicated, and marked with a weak keel
at the outer edge of the umbilicus. The base is marked by incre-
mental lines and obsolete spiral striations. The umbilical wall bears
many slender axial riblets: Aperture subcircular, decidedly oblique
and decidedly protracted at the posterior angle. Outer lip thin and
sharp at the edge; inner lip somewhat thicker. Operculum typically
cyclobakerid.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 123
The specimen figured, U.S.N.M. No. 535980, is one of the Adams
collection labeled “jamaicense Sowerby.” It has 4.5 whorls and
measures: Height, 15 mm.; greater diameter, 20.8 mm.; lesser diam-
eter, 16 mm.
Chitty’s notatior came from St. Elizabeth Parish, J amaica. Our
specimen appears to agree with Chitty’s diagnosis.
POTERIA (CYCLOBAKERIA) YALLAHSENSIS, new species
Puate 17, Ficures 16-18
1857. Cyclotus notatior var. a Cuitty, Proc. Zool. Soe. London, vol. 25, p. 148.
Shell very small, elevated helicoid, wood brown, with the nucleus
pale buff, the succeeding turn pale chestnut-brown and the rest
wood brown. ‘The nucleus consists of a single well-rounded, smooth
turn; the succeeding turns are marked by feeble, slightly retractively
curved, closely spaced hairlines, those of the last whorl being a little
stronger. Beginning with the last whorl, feeble nodulose cords make
their appearance. These cords are more or less protractively curved
but irregularly developed so as to produce a zigzag effect. Suture
well impressed, except on the last whorl, which is appressed to the
preceding whorl. Periphery well rounded, marked by the feeble
continuation of the sculpture of the spire. Base inflated, strongly
rounded, narrowly umbilicated, with a weak cord marking the outer
edge of the umbilicus. The base is marked by lines of growth and
a few indications of spiral threads. The umbilical wall is marked
by strong, closely spaced incremental lines. Aperture decidedly
oblique, protracted at the posterior angle to form an angle; peristome
rather thin. Operculum typically cyclobakerid.
The type, U.S.N.M. No. 356093, comes from Yallahs River, St.
Thomas Parish, Jamaica. It has 4.9 whorls and measures: Height,
11.0 mm.; greater diameter, 15.0 mm.; lesser diameter, 11.3 mm. Dr.
Welch also collected a large series at White Horses, St. Andrew
Parish.
This species can readily be distinguished from P. (C.) notatior
(Chitty) by the absence of the depressed area below the summit.
POTERIA (CYCLOBAKERIA) BALNEARIS, new species
PLATE 17, Figures 7-9
Shell small, depressed-helicoid, covered with a dark wood-brown
periostracum. The nucleus consists of a single well-rounded, smooth
turn. The postnuclear whorls are well rounded and marked by
retractively curved hairlike axial riblets. The last whorl also has
feeble zigzag axial riblets, which do not come quite to the summit.
124 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Suture well impressed, less so on the last half of the last turn, which
is appressed to the summit. Periphery strongly rounded, marked by
the feeble continuation of the axial riblets. Base inflated, strongly
rounded, narrowly umbilicated with a weak cord marking the outer
edge of the umbilicus. The base outside of the umbilicus shows a
few weak pits outside of the basal cord and mere indications of obso-
lete nodulations, which fade out toward the end of the last turn.
The umbilical wall is marked by moderately strong incremental lines.
Aperture circular, protracted into a slight angle at the posterior
angle; peristome somewhat thickened all around. Operculum
typically cyclobakerid.
The type, U.S.N.M. No. 535981, was collected by Orcutt near Bath,
St. Thomas Parish, Jamaica. It has lost part of the early turns;
the 3 remaining measure: Height, 12.4 mm.; greater diameter, 16.6
mm.; lesser diameter, 13.2 mm.
The species seems to extend through the Morant Bay region, St.
Thomas Parish.
Genus APEROSTOMA Troschel
1847. Aperostoma TroscHEL, Zeitschr. Malak., vol. 4, p. 44.
1847. Aperostoma PrerrrER, ibid., p. 47. (Type designation, Cyclostoma (Apero-
stoma) blanchetianum Moricand. )
1850. Aperostoma Petit, Journ. Conchyl., vol. 1, p. 88 (C. giganteum Sowerby
designated as type).
1852. Aperostoma HERRMANNSEN, Indicis generum malacozoorum primordia, vol.
2, Suppl., p. 10 (C. mexicanum Menke designated as type).
Troschel (loc. cit.) in 1847 divided the cyclostomids into eight
genera. On page 44 he created the genus Aperostoma and named
three species under it, namely, Cyclostoma volvulus Lamarck, C.
mexicanum Menke, and C. blanchetianum Moricand, without designat-
ing a type. Pfeiffer, in the article following Troschel’s, discussed
his own work on the group and commended Troschel’s endeavors,
which he accepted as sound, but he said that genus 4, Aperostoma,
contains heterogeneous elements, namely, some with calcareous oper-
culum and some with horny. He restricted the name Aperostoma to
shells with a calcareous operculum. He placed those with a horny
operculum in Cyclophorus Montford, citing as type C. volvulus
Miiller.
This restriction of Aperostoma Troschel removes (@. volvulus
Lamarck and C. mexicanum Menke from the genus and leaves only
C. blanchetianwm Moricand, which becomes its type.
KEY TO THE SUBGENERA OF WEST INDIAN APEROSTOMA
Umbilicusibounded by; alikeele se a ee ee Cycladamsia
Umbilicus not bounded by a keel.
SCulpeave .PULOSSH te sete Ee AE ae Se Yee ee eae Austrocyclotus
Sculpture) mot (Tugosee eee ee Cyclohidalgoa
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 125
CYCLADAMSIA, new subgenus
Shells of this subgenus resemble Veocyclotus in shape and general
sculpture of shell but differ in having a keel at the exterior limit of
the umbilicus. Operculum covered on the outside with a thick cal-
careous deposit, which forms a weak ridge on the inner edge of the
whorls and tapers outwardly, its surface being obliquely striated.
Type: Cyclostoma seminudum C. B. Adams = Aperostoma (Cycla-
damsia) seminudum (C. B. Adams).
The subgenus Cycladamsia appears confined to a strip of western
Jamaica lying between longtitude 77°30’ and 78° W. and latitude 18°
and 18°25’ N., except for two crab-carried specimens from Montego
Bay, which is a little farther north.
The radula of Aperostoma (Cycladamsia) seminudum seminudum
(C. B. Adams) and A. (@.) seminudum scabratum have the formula
3:3:38:3, and the jaw without median projection.
KEY TO THE SPECIES OF SUBGENUS CYCLADAMSIA
Whorls strongly pitted or malleated.
Last fourth of base of last whorl with regular oblique ridges.
Oblique ridges extending from umbilical keel to periphery______ ruber
Oblique ridges not extending from umbilical keel to periph-
OV ook sed ee See eR en bairdianum
Last fourth of base of last whorl without regular oblique ridges.
Shell large, greater diameter more than 23 mm___________ seminudum
Shell smaller, greater diameter less than 20 mm_______________ fossile
Whorls not strongly pitted or malleated___________________.. rudisplanusque
APEROSTOMA (CYCLADAMSIA) RUBER (Chitty)
Shell helicoid. Nuclear whorls 1.3, small, well rounded, smooth; the
first half postnuclear whorl with slender, retractively curved axial
riblets, which are separted by spaces about as wide as the riblets.
This stage is followed by a turn in which the axial sculpture becomes
irregular and the ribs obsolete, then by a wrinkled and pitted stage
that continues to the end of the last whorl. In the early portion of
this stage the wrinkles are more or less regular and decidedly retrac-
tively curved. They are not of uniform development, which condi-
tion gives the spaces between the wrinkles a pitted aspect. The
summit of the whorls is appressed; the suture is not impressed,
Periphery well rounded. Base well rounded, openly umbilicated,
the umbilicus marked by a strong cord at its outer termination.
The cord within the umbilicus is pinched in. The sculpture of the
base consists of decidedly obliquely retractively curved cords, which
are wavy and feebly nodulose, enclosing spaces of a little less width
between them, which have a somewhat pitted aspect. The umbilical
428349429
126 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
wall is marked by strong, curved, rugose incremental ridges. Aper-
ture oblique, subcircular and angled at the posterior angle; peristome
slightly thickened on the outer and basal lip and somewhat emarginu-
late on the parietal wall. Operculum typically cycladamsid.
This species is nearest related to A. (C.) batrdianum (Chitty),
from which it can be distinguished by the fact that the oblique
ridges on the last whorl cross the periphery undiminished. In
bairdianum they vanish on the base before reaching the periphery.
I am recognizing two subspecies, which the following ‘key and
descriptions will help to differentiate:
KEY TO THE SUBSPECIES OF APEROSTOMA (CYCLADAMSIA) RUBER
Shell large, greater diameter more than 21 mm___-----~-----~~-~-~------- ruber
Shell smaller, greater diameter less than 18 mm_---_~------------- pretiosum
APEROSTOMA (CYCLADAMSIA) RUBER RUBER (Chitty)
PLATE 17, Ficures 31-33
1857. Cyclotus ruber Curtry, Proc. Zool. Soe. London, vol. 25, p. 153.
This subspecies has the early whorls pale brown; the part of the
periostracum remaining is chestnut-brown. In denuded specimens
the shell is flesh colored, with a pale brownish tinge and usually with
several broad bands of dark brown, the periphery appearing as a
lighter zone.
This subspecies is distinguished from A. (@.) ruber pretiosum
(Chitty) by its much larger size.
Chitty cites Westmoreland as type locality. Westmoreland is the
westernmost as well as one of the largest parishes of Jamaica. An
absence of a definite locality makes it not unlikely that his shells
came from the St. Elizabeth Parish border. The specimen figured,
U.S.N.M. No. 535982, is from the C. B. Adams collection at Amherst
and bears the label C. seminudum Adams var., St. Elizabeths Ja-
maica, E. Chitty donor! This has 5 whorls and measures: Height,
15.3 mm.; greater diameter, 22.0 mm.; lesser diameter, 16.1 mm.
It must be remembered that Adams visited Chitty and obtained
many specimens from him. Chitty did not venture to enter the de-
scriptive field until after Adams’ death. We know that Chitty de-
scribed material from New Hope, Westmoreland, which is near the
southeastern border of Westmoreland. All the material that we
have seen with definite locality comes from the region of Carisbrook,
St. Elizabeth.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 127
APEROSTOMA (CYCLADAMSIA) RUBER PRETIOSUM (Chitty)
PLATE 17, Ficures 25-27
1857. Cyclotus pretiosus Cutty, Proc. Zool. Soc. London, vol. 25, p. 153.
In type of sculpture this subspecies corresponds in every way with
that of typical ruber. It differs from it mainly in its much more
diminutive size.
The specimen figured, U.S.N.M. No. 400998, was collected by C. R.
Orcutt on a tall hill near Maggotty, St. Elizabeth Parish, Ja-
maica. It has 3.4 whorls remaining and measures: Height, 8.5 mm. ;
greater diameter, 15.0 mm.; lesser diameter, 11.2 mm.
In his description Chitty cites New Hope, Westmoreland, as type
locality. We have not seen specimens from New Hope, but we do
have lots from three adjoining parts of St. Elizabeth Parish, namely,
Maggotty, Cheltenham, and a road cut 4 miles north of Black River.
APEROSTOMA (CYCLADAMSIA) BAIRDIANUM (Chitty)
Pate 17, Fiaures 28-30
1857. Cyclotus bairdianus Cuirry, Proc. Zool. Soc. London, vol. 25, p. 153.
Shell helicoid, of medium size, “of rich red brown color, with some.
times a lighter band at the periphery and around the umbilical keel,
yellowish at the suture of the last whorl” (Chitty). The nucleus
consists of about one turn. The first postnuclear whorl is marked
by retractively curved axial riblets, which are a little broader than
the spaces that separate them. The whorl succeeding shows lines of
growth, which are strongest at the summit, and malleations on the
rest of the shell. This type of sculpture continues to the aperture, but
becomes intensified and developed more or less into strong wrinkles.
Suture moderately impressed. Periphery well rounded. Base well
rounded, openly umbilicated and marked with a very strong keel at
the outer edge of the umbilicus and decidedly pinched in below the
keel in the umbilicus. The base is marked by pittings which extend
to the periphery. On the last one-fifth of a turn there are also
strong decidedly retractively curved axial cords, which are separated
by spaces about as wide as the cord. These vanish before reaching
the periphery. The umbilical wall is marked by irregular lines of
growth which render it rather rough. Aperture oblique, circular,
with a feeble angle at the posterior angle, slightly emarginated below
the summit on the outer lip and slightly excavated on the columellar
wall. Operculum typically cycladamsid.
Chitty failed to cite any locality with his description. Three
specimens before me from the Adams collection mixed with “C.
128 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
varians Ads.” No. 14, come from Accompong Town, St. Elizabeth
Parish. Adams credits them to Chitty as donor. Our specimens
were collected by Orcutt at Maggotty, St. Elizabeth Parish, about 5
miles south of Accompong.
The specimen figured, U.S.N.M. No. 535983, comes from Maggotty,
St. Elizabeth Parish. It has 4.5 whorls and measures: Height, 12.0
mm.; greater diameter, 17.0 mm.; lesser diameter, 13.0 mm. These
meet most nearly the measurements cited by Adams. There are, how-
ever, larger individuals obtained at the same locality. One of these
has 5 turns and measures: Height, 14.8 mm.; greater diameter, 21.4
mm.; lesser diameter, 16.2 mm. 4
This species belongs to the group in which the basal sculpture near
the aperture consists of parallel, retractively slanting cords. This
character it shares with A. (@.) ruber (Chitty), from which it differs
in having much feebler sculpture on the latter portion of the spire
as well as the base.
APEROSTOMA (CYCLADAMSIA) SEMINUDUM (C. B. Adams)
Shell varying greatly in shape, which ranges in outline from
depressed-helicoid to subglobular. The color is also decidedly vari-
able, ranging from unicolor to banded, the light area usually being
flesh colored or buff, the darker chestnut-brown. The sculpture is
likewise immensely variable. The nucleus is small, usually consist-
ing of a turn or a turn and a half, which is smooth. The succeeding
postnuclear whorl is marked by finely, retractively curved, rather
closely spaced axial riblets, and from there on the remaining whorls
show rough lines of growth and numerous wrinkles, malleations,
and pittings. The strength of these elements varies very materially.
In some specimens they become decidedly enfeebled toward the pe-
riphery of the last turn; in others, they remain conspicuous. The
suture is moderately impressed. Periphery well rounded, smooth
in some individuals; in other individuals, pitted. Base well rounded,
openly umbilicated, the umbilicus varying materially in size. In
one of the races it is exceedingly broad, while in another it is quite
narrow. A strong keel marks the outer limits of the umbilicus.
The umbilical edge of this keel is deeply impressed. The base varies
materially in sculpture; in some the posterior half is practically
smooth; in others, it is strongly pitted and decidedly rough. The
umbilical wall is marked by strong riblike incremental elements.
The aperture is subcircular, somewhat angulated at the posterior
angle; in some individuals there is a decided sinus on the outer lip
a little below the summit; in others it is absent. The columellar
wall is slightly excavated. Operculum typically cycladamsid, but
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 129
differing materially in the concavity of its outer surface, which ranges
from almost flat to decidedly concave.
This species presents a difficult problem to the taxonomist, for I
was unable to find sharp, clearly cut geographic races, but I did
find intergrading elements in all the characters mentioned; that is,
elevation, width of umbilicus, intensity of sculpture or partial
absence thereof.
I am recognizing four subspecies, which appear to have more or
less of a zoogeographic locus:
KEY TO THE SUBSPECIES OF APEROSTOMA (CYCLADAMSIA) SEMINUDUM
Periphery strongly sculptured.
SpirephelicOid +25 h.22 Siess Oa See ee scabratum
Spirevdepressed:helicoidas= 4s «2 Pe Pe Sa 2 eee eh ee humile
Periphery not strongly sculptured.
Wim Dili@us Natt O Wess ae eee eee eee deburghaeanum
Wmbillcus notmarrowe =e a ee ee ee __ seminudum
APEROSTOMA (CYCLADAMSIA) SEMINUDUM SCABRATUM, new subspecies
PLATE 18, FIcuRES 26-28
This subspecies has a well-elevated, helicoid outline and is exceed-
ingly roughly sculptured, the pitting extending over the periphery.
It is easily distinguished from A. (C.) s. hwmile, which also has the
pitting extending over the periphery, by its elevated form.
The type, U.S.N.M. No. 535984, was collected by Dr. Welch at
The Alps, Trelawny Parish, Jamaica. It has 5 whorls and measures:
Height, 18.7 mm.; greater diameter, 26.0 mm.; lesser diameter,
19.3 mm.
Welch also collected this race at Croydon, east-central Trelawny
Parish. Both of these localities represent the northeastern known
limit of the subspecies.
APEROSTOMA (CYCLADAMSIA) SEMINUDUM HUMILE, new subspecies
PLATE 18, Ficures 29-31
This subspecies has the shell decidedly depressed and finely roughly
sculptured, the pitting and wrinkling extending over the periphery.
It also has an exceedingly wide umbilicus. The depressed form and
large umbilicus and finer sculpture will distinguish this race from
A. (C.) seminudum scabratum.
The type, U.S.N.M. No. 535985, has 5 whorls and measures: Height,
15.2 mm.; greater diameter, 25.6 mm.; lesser diameter, 19.0 mm. It
was collected by Dr. d’Alté Welch at Kyle, St. Elizabeth Parish,
Jamaica.
130 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
This race was collected by C. R. Orcutt at Carisbrook and by
Dr. Welch at Kyle and Bartons. It therefore seems to occupy the
north-central portion of St. Elizabeth Parish.
APEROSTOMA (CYCLADAMSIA) SEMINUDUM DEBURGHAEANUM (Chitty)
Puate 18, Ficurres 35-37
1857. Cyclotus deburghaeanus Cuirry, Proc. Zool. Soc. London, vol. 25, p. 151.
This subspecies was differentiated from typical seminudum by
Chitty chiefly on the basis of its more elevated form, narrower
umbilicus, and flatter operculum, and possibly a little feebler sculpture
of the base, for in both of these subspecies there is a tendency toward
obsoleteness of sculpture. I have found in examining series of
specimens that the relative concavity of the operculum is a variable
element, and the width of the umbilicus is also variable, as is the
elevation, and for this reason I am relegating Chitty’s species to
subspecific rank.
The specimen figured, U.S.N.M. No. 535986, was collected by Dr.
Welch west of Headings, Trelawny Parish, Jamaica. This has 5
whorls and measures: Height, 20.0 mm.; greater diameter, 25.3 mm. ;
lesser diameter, 18.0 mm.
Chitty cites Westmoreland (?) as the type locality. I believe that it
occupies northern Manchester and southern Trelawny Parishes,
extending into the edge of southeastern St. James Parish.
APEROSTGMA (CYCLADAMSIA) SEMINUDUM SEMINUDUM (C. B. Adams)
PLATE 18, FicuRES 32-34; PLATE 42, FIGURES 4, 5
1851. Cyclostoma seminudum C. B. Apams, Ann. Lyc. Nat. Hist. New York, vol.
5, p. 59.
This subspecies is a little less elevated than A. (C@.) seminudum
deburghaeanum (Chitty) and a little more roughly sculptured. It
also has a much broader umbilicus and it, like deburghaeanum, has
a feeble sculpture on the base, where it fails to extend to the periphery.
Adams cites Manchester, Jamaica, as the type locality. Chitty, in
restricting seminudum, says that it comes from the northwest border
of Manchester Parish; he specifically mentions Bogue Hill. I be-
lieve that this subspecies ranges from Somerset, Manchester Parish,
in a northwesterly direction to Phoenix, St. James Parish.
The specimen figured, U.S.N.M. No. 6642, was received from C. B.
Adams. It has 5 whorls and measures: Height, 16.0 mm.; greater
diameter, 23.5 mm.; lesser diameter, 18.1 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 131
APEROSTOMA (CYCLADAMSIA) FOSSILE, new species
PLATE 18, FicurEs 22-24
Shell of medium size, helicoid, yellowish white. The nucleus con-
sists of a little more than one turn; the next two turns are marked by
well-rounded, retractively curved axial riblets, which are a little wider
than the spaces that separate them. The next whorl is marked by
strong rough wrinkles at the summit. The rest of the whorls are
marked by retractively curved cords, which are slightly wavy and
slightly nodulose, producing a somewhat pitted effect on the general
surface. The last half of the last turn is very roughly sculptured.
The wrinkles at the summit are intensified, and the rest loses its
regular retractively curved ribbing, which is replaced with irregular
wrinkles, malleations, and pits. Suture well impressed. Periphery
well rounded. Base well rounded, broadly openly umbilicated with
a moderately strong cord marking the outer limit of the umbilicus.
The base of the early part of the last whorl is marked by the con-
tinuation of the axial cords referred to for the spire, and the same
type of pitting between them. The regularity of the disposition of
these elements disappears on the last half, where a similar type of
pitting and nodulation is present, but no regular distribution of these
elements obtains. The umbilical wall is marked by wavy strong axial
ribs. Aperture very large, somewhat channeled at the posterior
angle and slightly excavated on the columellar wall. Operculum
unknown.
The type, U.S.N.M. No. 535987, and a large series of specimens,
U.S.N.M. No. 375436, were obtained by Mr. Orcutt from fossiliferous
sand sifting in a cave north of One Eye River Sink, near Balaclava,
Manchester Parish, Jamaica. The type has 4.5 whorls and measures:
Height, 11.7 mm.; greater diameter, 18.2 mm.; lesser diameter,
13.9 mm.
An additional lot, U.S.N.M. No. 397355, was obtained on a hill north
of One Eye River Sink below the cave in crevices in the precipice.
This species belongs to the seminudwm group, as far as the type
of sculpture is concerned, but differs from it in the larger aperture
and the much stronger umbilical sculpture and also in having the
sculpture of the spire and base finer. It is evidently a fossil. Or-
cutt failed to find living specimens of it in the region.
APEROSTOMA (CYCLADAMSIA) RUDISPLANUSQUE (Chitty)
PLATE 18, FIGURES 13-18
1857. Cyclotus rudis-planusque Cuitty, Proc. Zool. Soc. London, vol. 25, p. 152.
Shell small or medium sized, helicoid. When perfect, covered with
a wood-brown periostracum, which may be streaked with axial darker
132 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
shades of brown. The nucleus consists of a little more than one
turn, which is small and well rounded. The next turn is marked
by very fine, retractively curved, closely spaced, almost hairlike
riblets, while the rest of the shell bears irregular rugations and
scattered malleations, the latter usually a little below the summit.
The rugations are strongest near the summit of the shell. Suture
well impressed. Periphery well rounded. Base well rounded, mod-
erately broadly umbilicated. The umbilicus is bounded by a strong
keel, which is strongly inpinched on the umbilical side and rendered
somewhat nodulose by the irregular axial sculpture. The umbilical
wall is marked by irregular axial ribs. Aperture large, subcircular,
except for an angulation at the posterior angle, feebly incised below
the summit on the outer lip and somewhat incised on the columellar
wall. Operculum typically cycladamsid.
The two specimens figured, U.S.N.M. No. 356182, are part of four
from Mulgrave, St. Elizabeth Parish, Jamaica. I have selected these
to show the limit of variation in size and also in intensity of sculp-
ture. The larger has 5 whorls, the smaller 4.8. They measure,
respectively: Height, 13.3 and 9.2 mm.; greater diameter, 18.6 and
13.2 mm.; lesser diameter, 14.0 and 10.2 mm.
This species is easily distinguished from the other Cycladamsias
by its extremely reduced sculpture.
Chitty cites Accompong Town, St. Elizabeth Parish, as type
locality. This appears to be the center of its distribution. Some
crab-carried (?) specimens were collected at Montego Bay, St. James
Parish. The living material seen extends from Mocho to Maggoty,
but we also have some dead specimens from 4 miles north of Black
River, St. Elizabeth Parish, and Glen Burnie Mountains, West-
moreland (Parish.
This species presents a puzzle that will be cleared up only when
anatomic material comes to hand for dissection. There are two dis-
tinct sizes, a large and a small element. It is possible that these
may be sexual characteristics. The occurrence of both in the same
locality points toward such a condition.
AUSTROCYCLOTUS, new subgenus
Aperostomine shells of helicoid shape the outside surface of which
is marked by closely placed threads crossing each other in protractive
and retractive series, producing an engine-turned pattern.
Type: Aperostoma (Austrocyclotus) stramineum (Reeve).
Distribution: Panama to Ecuador and Venezuela and the West
Indies.
The radula formula of Aperostoma (Austrocyclotus) grenadense
mesweent is 8:3:3:2. The jaw is without a median projection.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 133
The verge is situated on the back of the neck behind the tentacles.
It is swollen basally and provided with a seminal groove and a very
short simple terminal appendage.
KEY TO THE WEST INDIAN SPECIES OF THE SUBGENUS AUSTROCYCLOTUS
Sculpture very strong and filelike---------------------------------- rugatum
Sculpture not as strong or filelike.
Shell averaging more than 20 mm. in diameter___~---------- vincentinum
Shell averaging less than 15 mm. in diameter__---------------- grenadense
APEROSTOMA (AUSTROCYCLOTUS) RUGATUM (Guppy)
PLATE 17, Ficures 10-12
1864. Cyclotus rugatus Guppy, Ann. Mag. Nat. Hist., ser. 3, vol. 14, p. 246.
Shell of medium size, depressed-helicoid, openly, moderately,
broadly, umbilicated, covered by a golden-brown periostracum. Nu-
clear whorls 1.5, well rounded, smooth, forming a moderately elevated
spire. Postnuclear whorls inflated, strongly rounded, strongly in-
bent at the suture excepting the last fifth of the last turn, which
is marked by rather strong, decidedly retractively curved axial rib-
lets. On the last whorl the riblets are replaced by more or less
zigzagging ridges separated by spaces equaling them. This zig-
zagging produces a filelike resemblance. On the last fifth of a
turn these elements become much disturbed and finally obsolete.
Suture strongly impressed; periphery strongly rounded. Base in-
flated, strongly rounded, and marked by the continuation of the
sculpture at the spire. Umbilical wall marked by closely spaced
axial cords. Aperture oblique, almost circular, slightly angulated
at the posterior angle. Peristome simple. Operculum typically
austrocyclotid.
I ‘collected the specimen described and figured, U.S.N.M. No.
390108, at Brigham Hill, Central Range, Trinidad, on September 2,
1928. It has 4.4 whorls and measures: Height, 12.9 mm.; greater
diameter, 17.5 mm.; lesser diameter, 15.6 mm. I took three additional
specimens, U.S.N.M. No. 394005, on Verdant Vale, about 4 miles north
of Arima, Trinidad, on September 3, 1929.
This species is much more robust and rotund, with the sculpture
much coarser and rougher, than A. (A.) grenadense (Shuttleworth).
APEROSTOMA (AUSTROCYCLOTUS) VINCENTINUM (Pilsbry)
PLATE 17, Figures 1-3
1935. Poteria vincentina Pitspry, Proc. Acad. Nat. Sci. Philadelphia, vol. 87,
p. 4, pl. 1, figs. 2, 2a.
Shell rather large, depressed-helicoid. Nuclear whorls almost 2,
smooth, well rounded. Postnuclear whorls marked by numerous,
134 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
wavy, slender, raised, retractively slanting axial threads, which are
disposed in a more or less zigzag pattern, and lend to the surface
a worn-file effect. Suture well constricted. Periphery somewhat in-
flated, strongly rounded. Base strongly rounded, marked by the
continuation of the sculpture described for the spire. Umbilicus
broad, open. Aperture almost circular, somewhat angulated at the
posterior angle; peristome thin, slightly reflected, a little thicker on
the columellar wall, adnate to the preceding turn at the parietal wall.
Operculum typically austrocyclotid.
The specimen described and figured, U.S.N.M. No. 5385919, was col-
lected by Bartsch on Mount St. Andrews, St. Vincent. It has 4.4
whorls and measures: Height, 13.7 mm.; greater diameter, 22.7 mm. ;
lesser diameter, 17.1 mm.
This species recalls most nearly A. (A.) grenadense (Shuttleworth),
from which its much larger size will readily distinguish it.
APEROSTOMA (AUSTROCYCLOTUS) GRENADENSE (Shuttleworth)
Shell small, depressed-helicoid, openly moderately broadly umbili-
cated. Nuclear whorls about 2, well rounded, smooth, conforming in
increase with the rest of the spire. Postnuclear whorls slightly flat-
tened near the suture, the rest well rounded, marked by incremental
lines and obliquely slanting, low, elongated tubercles, which lend the
surface a wrinkled appearance. This sculpture usually weakens to-
ward the last tenth of a turn, where the incremental lines become
much rougher and irregular. Periphery of last whorl well rounded.
Base strongly rounded, marked like the spire. On the umbilical wall
the tuberculations may weaken. Aperture broadly ovate, somewhat
pointed at the posterior angle. Peristome acute. Operculum of about
six whorls, typically austrocyclotid.
The species appears confined to Grenada. I am recognizing two
subspecies :
KEY TO THE SUBSPECIES OF APEROSTOMA (AUSTROCYCLOTUS) GRENADENSE
Baseystrongly, granulogee ne ee ee ee ee ee grenadense
Base not strongly: granulose: 23 3 = Eee mesweeni
APEROSTOMA (AUSTROCYCLOTUS) GRENADENSE GRENADENSE (Shuttleworth)
PLATE 17, Fiaures 19-21
1857. Cyclostoma (Cyclotus) grenadense SHUTTLEWORTH, Journ. Conchyl., vol. 5,
p. 266.
This subspecies is differentiated from A perostoma (Austrocyclotus)
grenadense mesweent by having the base strongly granulose.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 135
I collected it in large numbers, U.S.N.M No. 473940, under heaps
of leaves and branches in the Cacao plantation on the Annandale Es-
tate, August 25, 1929, and a small number, U.S.N.M. No. 390105, at a
much higher elevation at Granitown House, August 25, 1929. The
latter average a little smaller and stronger sculptured. Both of these
localities are on the western slope of Grenada.
U.S.N.M. No. 57768 contains two specimens received from Bland;
US.N.M. No. 366504 contains eight specimens, and U.S.N.M. No.
366503 two specimens from Vendryes. None of these lots bear specific
locality data, only the label Grenada, but all belong to this race.
Ten specimens from the Annandale lot yield the following
measurements :
Height Greater diameter | Lesser diameter
ae
. 6
9
20
ao
»9
3
. 4
9
9
RrPOOWH RATAT IO OO’
OPOWRN PERRO:
Of the animal of this subspecies from the Annandale estate, my
notes say: “Dorsal parts pale rose color, sides a little paler, sole of
foot rose colored with a buffish tinge; tentacles crimson.”
APEROSTOMA (AUSTROCYCLOTUS) GRENADENSE MCSWEENI, new subspecies
PLATE 17, FIGURES 22-24
Shell agreeing with typical A. (A.) grenadense grenadense
(Shuttleworth) in size and shape but differing from it in having the
sculpture of the spire a little less strong, while that of the base is
very much reduced. I found these characters common to the speci-
mens collected on the east slope of Grenada. My largest series,
U.S.N.M. No. 473942, was obtained in a cacao plantation under the
collected heaps of dead leaves and branches of Baltazar, August 25,
1929. My notes say of the animal, “Coloration the same as that of
the specimens obtained at the Annandale estate.”
I collected the type, U.S.N.M. No. 474088, on the estate of Dr.
Edgar deJ. McSween, one of my medical students now residing in
Grenada, and I gratefully acknowledge his helpfulness during my
sojourn there. The type has 4.3 whorls and its measurements are
136 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
cited below, together with those of a series of specimens obtained at
the same place:
Greater diameter Lesser diameter
Mm.
(type)
ONOCUWOOWO:
1
0
‘4
8
. 6
. 6
6
8
3
@
oak
sue
au
ie
al
.o5
so
. 4
. 0
. 2
Small specimens come from Telescope Hill on the northeast coast,
U.S.N.M. No. 307501. Gerrit S. Miller, Jr., collected representatives
at Mount Pleasant at an altitude of 1,500 feet, U.S.N.M. No. 360782.
Professor Summers collected specimens at an altitude of 700 feet,
without mentioning specific locality, U.S.N.M. No. 356045.
CYCLOHIDALGOA, new subgenus
Aperostomas of helicoid shape with closely crowded axial riblets.
Type: Aperostoma (Cyclohidalgoa) translucidum (Sowerby).
Distribution; Colombia, Venezuela, British Guiana, Trinidad, and
Margarita Island.
Guppy showed the radula formula to be 3: 3: 3: 3.
APEROSTOMA (CYCLOHIDALGOA) TRANSLUCIDUM TRINITENSE (Guppy)
PLATE 18, Figures 7-12
1864. Cyclotus trinitensis Guppy, Ann. Mag. Nat. Hist., ser. 3, vol. 14, p. 245.
1923. Poteria translucida trinitensis H. B. Baxer, Occ. Pap. Mus. Zool. Univ.
Michigan, No, 187, pp. 31-33, pl. 5, figs. B and C.
Baker, in discussing the mainland forms of this species, recognizes
the subspecific distinctness of the Trinidad race, stating, “Shell more
depressed ; suture distinctly impressed; whorls convex.” The group is
an exceedingly difficult one, the proper elucidation of which may be
furnished when enough anatomic data has been adduced.
Guppy describes the Trinidad animal as follows: “The animal is
of a pinkish colour, which is most pronounced about the tentacles.
The eyes are small and black. The mouth is provided with an amber-
coloured, somewhat triangular mandible, divided into two parts by a
median fissure, from which diverge slightly curved rows of minute
denticulations strongly resembling the lingual teeth of some Helicidae.
THE CYCLOPHORID LAND MOLLUSKS OF AMBERICA 137
“The lingual teeth are 3. 1. 3, in arched rows: central broad, triden-
tate; 1st lateral broad, bidentate, with a base much produced out-
wardly; 2nd tridentate; 8rd much hooked and reflexed, tridentate.”
I collected semifossil specimens on the bluffs in a quarry on the
south side of the road between Pampator and Asuncion on Margarita
Island. I failed to find living specimens anywhere on the parts of the
island visited.
INCERTICYCLUS, a pseudogeneric term
Members of the family Cyclophoridae as a rule require a knowledge
of the operculum to enable one to assign them to the proper generic
and subgeneric groups. Since specimens not infrequently drift into
museums without this door, and the rest of the shell characters may
fail to enable one to locate the shell in question, I propose the above
name as a superspecific designation for species that have been
christened but that in our monographic endeavors we are unable to
locate properly for lack of the characters mentioned.
Incerticyclus is not to be considered a genus, nor is it ever to have
a type, but is to remain an open catch-all into which we may place
those species the position of which we are uncertain. When the neces-
sary characters have been adduced these species can be properly
aligned, taken out of Jncerticyclus, and when new material without
opercula is received, it can be placed in this group.
I am placing the following species in Jncerticyclus :
INCERTICYCLUS BAKERI (Simpson)
PLATE 18, FicurEs 1-3
1895. Neocyclotus (Ptychocochlis) bakeri Stmpson, Proc. U. 8S. Nat. Mus., vol. 17,
p. 449, pl. 16, figs. 1, 2.
Shell large, helicoid; nuclear whorls lost. The early postnuclear
whorls are marked by wavy or slightly zigzag, retractively slanting,
slender axial riblets, which are well developed, quite regular and
regularly spaced, and separated by spaces about as wide as the riblets.
On the last half of the last turn, however, there appears another
type of sculpture consisting of strong wrinkles, the first part of
which is retractive, then vertical, then again sloping retractively
to the periphery. These wrinkles vary materially in strength.
Suture moderately well impressed. Periphery well rounded. Base
well rounded, openly umbilicated, and marked by decidedly retrac-
tively curved, very regular ridges, which pass over the umbilical
angle strongly onto the umbilical wall. These ridges are about as
wide as the spaces that separate them. In addition to this the entire
base is marked by fine incremental lines. There is scarcely a sug-
138 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
gestion of the angle at the outer edge of the umbilicus present.
Aperture circular; peristome thin. Operculum?
The type, U.S.N.M. No. 115717, was collected by John B. Hender-
son in the mar] stratum of the Bowden Beds, Miocene, in St. Thomas
Parish, Jamaica. It has 3.4 whorls remaining and measures:
Height, 16.3 mm.; greater diameter, 30.0 mm.; lesser diameter,
20.0 mm.
The peculiar sculpture combined with the absence of the umbilical
keel separates this species from all the other known Jamaican
cyclophorids.
INCERTICYCLUS BOWDENENSIS, new species
PLATE 41, F1cuRES 4-6
Shell small, helicoid, yellowish white. Nuclear whorls decollated.
The early postnuclear whorls somewhat impressed at the summit;
tie last appressed. The early whorls are marked by slender, hairlike
incremental lines. Beginning with the last half of the penultimate
turn, decidedly obliquely protractively slanting axial rugae make their
appearance. They take a sudden retractive slant near the summit
where they are weaker and less regular. Periphery rounded. Base
strongly rounded, openly umbilicated, and marked by lines of growth
only. A rather strong, somewhat notched, moderately broad keel
forms the outer edge of the umbilicus, while the umbilical wall is
marked by riblike gatherings of the incremental lines. Aperture
circular, oblique; peristome simple, thickened to form an auricle at
the posterior angle.
The type, Acad. Nat. Sci. Philadelphia No. 82532a, was collected
by Uselma C. Smith and S. L. Schermo in the Miocene formation
at Bowden, Jamaica. It has 3.5 whorls remaining and measures:
Height, 10.8 mm.; greater diameter, 15.7 mm.; lesser diameter, 12 mm.
Its lesser size and comparatively weak sculpture will readily
distinguish it from Jncerticyclus bakeri (Simpson) and J. schermoz.
INCERTICYCLUS SCHERMOI, new species
PLATE 41, Figures 10-12
Shell of medium size, helicoid, cream yellow. Nuclear whorls 1.5,
small, well rounded, smooth. Postnuclear whorls well rounded; all
but the last turn marked by very strong, slightly retractively curved
axial riblets, which are less than half as wide as the spaces separating
them. The last whorl is very roughly wrinkled. The axial ribs
persist and are even stronger than on the preceding whorls, but they
are rendered irregular by decidedly obliquely protractively slanting
folds. The latter are not quite so broad as the spaces separating
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 139
them. Periphery well rounded. Base strongly rounded, openly um-
bilicated with a very strong keel at the outer edge of the umbilicus.
Deep pits are present at the keel, which extend as impressed grooves
over the anterior third of the base. Anterior to the keel the umbil-
ical wall is also deeply pitted and the impressed areas extending
from the pits over the umbilical wall let the intermediate space appear
as ribs marked by incremental lines. Aperture oblique, circular.
The type, Acad. Nat. Sci. Philadelphia No. 82582, was collected
by Uselma C. Smith and S. L. Schermo in the Miocene beds at
Bowden, Jamaica. It has 4.4 whorls and measures: Height, 13 mm. ;
greater diameter, 19.8 mm.; lesser diameter, 15.8 mm. Thirteen
young and fragmentary specimens are registered under the same
number.
The strong basal sculpture readily differentiates this from /ncer-
ticyclus bowdenensis and the strong spiral keel at the umbilicus sepa-
rates it from Incerticyclus bakeri (Simpson).
INCERTICYCLUS PERPALLIDUS (C. B. Adams)
PLATE 18, FIGURES 4-6
1852. Cyclostoma perpallidum C. B. ApAMs, Ann. Lyc. Nat. Hist. New York,
vol. 5, p. 81.
1852. Cyclotus perpallidus PFreIrFER, Monographia pneumonopomorum viventium,
vol. 1, p. 29.
1857. Cyclostoma perpallidus Currry, Proc. Zool. Soc London, vol. 25, p. 157.
1898. Neocyclotus (Plectocyclotus) perpallidus KosetT and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 9, p. 138, reprint.
Shell of medium size with sides sloping regularly to the spire,
soiled white. The nucleus consists of 1.5 smooth, well-rounded turns.
The postnuclear whorls are moderately well rounded, the first marked
by feeble retractively curved axial riblets. Beginning with the
second, fine wavy axial wrinkles make their appearance. These
wrinkles branch and anastomose, enclosing spaces between them of
varying length. They are a little weaker toward the summit than
on the major portion of the whorls. Suture moderately well im-
pressed on all of the turns. Periphery obscurely angulated and
marked like the spire. Base very moderately rounded, almost flat-
tened, marked by the continuation of the axial wrinkles, which here
are retractively slanting and marked near the umbilicus by strong
coarse ribs, which are wider than the spaces that separate them. The
umbilical wall is marked by the continuation of these ribs and fine
incremental lines. There is only the merest indication of angulation
at, the outer edge of the umbilicus. Aperture subcircular, decidedly
protracted into a claw at the posterior angle, having the outer lip
thin, inner somewhat thickened. Operculum?
140 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Adams’ type, here figured, has 5 whorls and measures: Height, 13.0
mm.; greater diameter, 19.4 mm.; lesser diameter, 15.4 mm.
Adams cites merely Jamaica, but Chitty (doc. cit.) states: “Near
Moore Town, Portland.” He says, “This shell was originally brought
to me by a negro named Shelly, whom I could never, even by money
excite to sufficient energy to collect more. * * * Operculum still
wanting.”
The species has not been rediscovered, and until the operculum has
been found it will be impossible to place it properly. The peculiar
sculpture, the sloping spire, and, above all, the flattish base, differentiate
it from all the other known West Indian cyclophorids.
INCERTICYCLUS PERPLEXUS, new species
PLATE 18, Ficures 19-21
Shell small, depressed-helicoid, covered with a golden-brown perio-
stracum, at least the fragments left on our specimens would indicate
that that is the case. The nucleus consists of 1.4 small, somewhat
inflated, well-rounded, smooth turns. The first postnuclear whorl is
strongly rounded and marked by very regular, rather distantly spaced,
slender, well-raised axial riblets. On the succeeding turns the axial
riblets are more closely approximated and reduced to incremental lines.
In addition to these, the entire surface is marked by strong, very
oblique, decidedly protractively slanting heavy ridges, which are
separated by spaces about as wide as the ridges. These ridges at the
periphery join the ridges on the base, which have a decidedly opposite
slant, the two at their junction forming sharp arrowpoints. Base
broadly, openly umbilicated, inflated, well rounded, and marked by the
heavy rugations mentioned above. The umbilical wall bears coarse,
more or less vertical, riblike incremental lines. Aperture circular;
peristome simple, that of the outer lip thin; the inner lip thickened.
Operculum unknown.
The type, U.S.N.M. No. 535988, was collected by Orcutt at Appleton,
St. Elizabeth, Jamaica. It has 3.3 whorls remaining and measures:
Height, 7.8 mm.; greater diameter, 14.9 mm.; lesser diameter, 11.3 mm.
U.S.N.M. No. 378448 contains two topotypes received from the same
source. These are a little smaller than the type.
INCERTICYCLUS MARTINICENSIS (Shuttleworth)
1857. Cyclostoma (Cyclotus) martinicense SHUTTLEWORTH, Journ. Conchyl., vol.
5, p. 267.
Shell broadly umbilicated, lenticular, thin, finely striated, with dense
fine spiral grooves, pale olive-brown, shining; spire little elevated.
Apex rather obtuse; suture deep; 4-414 convex, rapidly increasing
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 141
whorls, the last scarcely descending ; aperture hardly oblique, circular ;
peristome continous, acute. Operculum calcareous, scarcely convex
on the outside, with ten volutions, depressed and expanded at the edge.
Height, 8 mm.; greater diameter, 14 mm.; lesser diameter, 11.5 mm.
Aperture 6 mm., broad and high.
Shuttleworth states that he had received two specimens of this from
Petit from the Island of Martinique. I have not seen it and am unable
to place it in any of the groups here listed.
INCERTICYCLUS CINEREUS (Drouet)
PuLate 18, FicuRE 25
1859. Cyclophorus cinereus DRovET, Essai sur les mollusques terrestres et fluvia-
tiles de la Guyane francaise, p. 90, pl. 3, figs. 37-38.
The following is a translation of Drouet’s description :
Shell small, umbilicated, subturbinate, thin, scarcely translucent,
finely striate, ashy; whorls 4, convex, apex obtuse; aperture angulatedly
circular, peristome simple, acute. Operculum immersed, concolor, nor-
mal. Height, 4 mm.; greater diameter, 4.5 mm.; diameter of aperture
2mm. * * * This Cyclophorid by its general form, size, and
coloration resembles our Valvata piscinalis Miill. It has no decided
characters except its small size, subturbinate form and ashy color, and
especially the fine regular striations with which its shell is adorned.
The figures emphasizes the axial lirations.
There is no material in the collection of the United States National
Museum that helps us understand this species. The fact that Drouet
describes Cyclophorus liratus and C. acutiliratus immediately before
his @. cinereus and says here that the operculum is normal, leads one
to believe that he was dealing with an amphicyclotid. We know of
none in the West Indies with axial ribs. I therefore include this
species in sedis incertae. It was described from the Island of Marti-
nique.
428349—42—__10
Part 3.—THE CYCLOPHORID MOLLUSKS OF THE
MAINLAND OF AMERICA
By PAuL BARTSCH and JOSEPH P. E. MORRISON
Family CYCLOPHORIDAE Gray
The family Cyclophoridae is defined on p. 3.
The mainland cyclophorids range from Mexico to and over South
America. They belong to four subfamilies, which the following key
will help to differentiate :
KEY TO THE SUBFAMILIES OF CYCLOPHORIDAE ON THE MAINLAND OF AMERICA
Shell elongate-turrited2 22255 a ee Megalomastominae
Shell not elongate-turrited.
FSV a Vey NS oh) 670) (5 Dane ee A ee eee Diplommatininae
Shell not pupoid but planorboid or helicoid.
@perculum’ chondroid= ==. +>_2 422222 "se. eS Amphicyclotinae
Operculum: calcified) ==) 2 eee Aperostominae
Subfamily MEGALOMASTOMINAE Torre and Bartsch
For definition of the subfamily, refer to p. 3.
Genus TOMOCYCLUS Crosse and Fischer
1872. Tomocyclus Crosse and FiscHer, Journ. Conchyl., vol. 20, p. 76.
Shells with elongated-turrited spire. Aperture circular, peristome
double; the inner more or less exserted; the outer broadly flaringly ex-
panded, with a slit terminating in a more or less circular perforation in
the parietal wall below the edge of the peristome. Operculum with
central nucleus, bearing a broadly expanded spiral lamella, which is
reflected to parallel the basal chondroid plate. The outer edge of the
lamella may be appressed to the succeeding turn, or fringes of its outer
ragged edge may be free; in reality the entire lamella is free, and its
appressed position is responsible for its appearing adnate.
Type: Zomocyclus gealei Crosse and Fischer.
T. simulacrum (Morelet) yields the following data: Radula formula :
3:3:3:3. Jaws with central projection. Penis on right side of neck
traversed by a groove only.
142
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 143
The radula of 7. gealei has the formula 3: 3: 3: 3. According to
Crosse and Fischer the jaw has a median projection, the verge is long
and is on the side of the neck behind the tentacles, tapering gradually
and bearing a seminal groove. There is no terminal appendage.
KEY TO THE SPECIES OF TOMOCYCLUS
Axdalusculpiune consisting.o: Slender ribss2s—s === === a eee gealei
Axial sculpture not consisting of slender ribs but of fine incremental
lines.
OuteraperistomMe~naLlTrow.- === =a eae ae ee guatemalensis
Outer peristome not narrow.
Siphonwstronghy, developedss =a se eee ee eee siphonis
Siphon not strongly developed.
Wihorlsestronglyarounded S22.) ee ee constrictus
Whorls not strongly rounded.
Shell large, height more than 85 mm__-_--________ simulacrum
Shell smaller, length less than 27 mm____-___----- copanensis
TOMOCYCLUS GEALEI Crosse and Fischer
PLATE 19, FIGuRE 1
1872. Tomocyclus gealei Crosse and FiscHer, Journ. Conchyl., vol. 20, p. 77.
1886. Tomocyclus gealei FISCHER and Crosse, Mission scientifique au Mexique et
dans l’Amérique centrale, vol. 2, pt. 7, pp. 118-120, pl. 40, figs. 1-3.
1890. Megalomastoma (Tomocyclus) gealei von Martens, Biologia Centrali-
Americana, p. 10.
Shell very large, very elongate-turrited, covered by a pale yellowish-
brown periostracum, which shows a few axial streaks of darker brown.
Peristome bluish white, which is also the color of the interior of the
aperture. The early whorls are decollated in all our specimens; those
remaining are appressed at the summit, well rounded, and marked by
somewhat sinuous, retractively curved axial riblets, which are about
as wide as the spaces that separate them. In addition to this the whorls
are marked by very closely spaced spiral striations. Suture strongly
impressed. Periphery well rounded. Base moderately long with a
strong, heavy, spiral keel bounding the outer edge of the umbilicus,
which is open. Aperture circular; peristome double, the inner slightly
exserted; the outer broadly flaringly expanded, with a slit in the
parietal wall, which at some little distance behind the edge of the
peristome develops into a perforation, the perforation being arched
over by the two sides of the peristome, which almost form a siphon.
Operculum typically tomocyclid.
The specimen described and figured, U.S.N.M. No. 321034, bears the
label “Central America.” It has 7 whorls remaining and measures:
Height, 40.7 mm.; greater diameter, 17.8 mm.; lesser diameter, 11.9
mm. Height of aperture, 14.1 mm. externally, 7.9 mm. internally ;
diameter, 13.2 mm. externally, 7.7 mm. internally.
144 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The United States National Museum collections contain the follow-
ing specimens: U.S.N.M. No. 32069, 3 from Guatemala; U.S.N.M. No.
251126, 1 collected by O. F. Cook at Alta Vera Paz, Guatemala;
U.S.N.M. No. 321081, 2 collected by S. Smith in Guatemala; U.S.N.M.
No. 57333, 3 received from Bland, collected in Guatemala; U.S.N.M.
No. 104444, 1 from Alta Vera Paz, Guatemala; U.S.N.M. No. 316386,
2 from Guatemala; U.S.N.M. No. 321033, 1 collected by Sallé, labeled
“Mexico.”
In size this species resembles 7’. s¢mulacrum, from which it is easily
distinguished by its rather strong axial ribs, which are absent in
simulacrum.
TOMOCYCLUS GUATEMALENSIS (Pfeiffer)
PLATE 19, Ficure 4
1851. Cyclostoma guatemalense PFEIFFER, Proc. Zool. Soc. London, 1851, p. 245.
1852. Megalomastoma guatemalense PFEIFFER, Monographia pneumonopomorum
viventium, vol. 1, p. 132.
1853. Cyclostoma guatemalense PFEIFFER, Martini-Chemnitz Conchylien Cabinet,
vol. 1, sect. 19, p. 267, pl. 36, figs. 18, 14.
1872. Tomocyclus guatemalense Crosse and FiscHer, Journ. Conchyl., vol. 20,
enbGe
1878. nat abun guatemalense REEVE, Conchologia iconica, vol. 20, pl. 8,
fig. 73.
1886. Tompopelie guatemalensis FiscHrER and Crosse, Mission scientifique au
Mexique et dans l’Amérique centrale, vol. 2, pt. 7, p. 124, pl. 40, fig. 11.
We have not seen specimens of this species and copy Pfeiffer’s
figures and translate his description as given in Martini-Chemnitz.
Shell perforate, elongate, rather thick shelled, moderately truncated
above, axially finely striate, white beneath a greenish-brown perios-
tracum. Spire with a somewhat convex outline, gradually tapering.
Whorls 6, slightly arched, the last shorter, descending anteriorly, and
somewhat solute, compressed about the narrow open umbilicus but
not keeled. Aperture parallel with the axis, almost circular. Peri-
stome free, white, double, the inner complete, and only slightly pro-
duced; the outer expanded at right angles to the aperture, excised
over the perforation. Length, 24 mm.; diameter, 8 mm.
Pfeiffer’s figure has 5.8 whorls remaining and measures: Height,
23.2 mm.; greater diameter, 11.6 mm.; lesser diameter, 8.2 mm.
Height of aperture externally, 8.2 mm., internally, 6.1 mm.; diameter
externally, 6.7 mm., internally, 5.1 mm.
Habitat: Alta Vera Paz, Guatemala.
These figures show a finely ribbed shell with a much more narrowly
expanded outer peristome than in any of the other species.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 145
TOMOCYCLUS SIPHONIS, new species
PLATE 19, Fiaure 5
Shell of medium size, olivaceous with a chestnut tinge; peristome
bluish white, which is also the color of the interior. Early whorls
decollated in all our specimens; those remaining well rounded, ap-
pressed at the summit, which is slightly denticulated and marked by
lines of growth, which do not form riblets. There is also an indi-
cation of microscopic spiral striations. Suture strongly constricted.
Periphery well rounded. Base with a weak carina at the outer edge
of the umbilicus, which disappears on the last quarter of the last turn.
Aperture circular; peristome double, the inner moderately exserted ;
the outer broadly flaringly expanded and somewhat wavy with a
strong sinus on the middle of the parietal wall, which connects with
a round opening, over which the two sides of the reflected outer
peristome form a pseudosiphon. Operculum typically tomocyclid.
The type, U.S.N.M. No. 162511, was collected by von Ihering at
Alta Vera Paz, Guatemala. It has 5 whorls remaining and measures:
Height, 31 mm.; greater diameter, 14.6 mm.; lesser diameter, 9.7 mm.
Height of aperture, 12.9 mm. externally, 5.9 mm. internally; diameter,
12.2 mm. externally, 5.8 mm. internally.
This differs from 7. constrictus in having the pseudosiphon much
more strongly developed and the basal carina much weaker; it is also
paler in coloration.
TOMOCYCLUS CONSTRICTUS, new species
PLATE 19, Figure 2
Shell of medium size, elongate-turrited, covered with a chestnut-
brown periostracum. Early whorls decollated in all our specimens;
those remaining strongly rounded, slightly denticulated at the sum-
mit, and marked by feeble obsolete axial riblets corresponding to the
denticulations at the summit; these, however, evanesce on the major
portion of the base. In addition to this, the whorls are marked by
fine spiral striations. Suture strongly constricted. Periphery well
rounded. Base slightly produced, bearing a very strong keel at the
outer limit of the umbilicus. Aperture circular; peristome double,
the inner moderately exserted; the outer broadly, wavily, flaringly
expanded, with a notch in the parietal wall which connects with the
round opening. The two sides of the peristome arch over to partly
cover the pseudosiphon. Operculum typically tomocyclid.
The type, U.S.N.M. No. 162315, was collected by Godman at Coban,
northern Guatemala. It has 5.3 whorls remaining and measures:
Height, 32.8 mm.; greater diameter, 15.2 mm.; lesser diameter, 10.4
mm. Height of aperture, 12.6 mm. externally, 6.5 mm. internally;
diameter, 12.2 mm. externally, 6.3 mm. internally.
146 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The United States National Museum collections contain also the
following specimens:
U.S.N.M. No. 516028, a topotype; U.S.N.M. No. 250693, 4 speci-
mens received from Rolle, collected at Finca de Providencia, Guate-
mala; U.S.N.M. No. 321030, 1 specimen received from Sowerby and
Fulton, labeled “Guatemala.”
This species resembles 7’. stphonis, but differs from it in its weaker
siphonal development and in having the whorls much more strongly
rounded and the basal keel much stronger.
TOMOCYCLUS SIMULACRUM (Morelet)
PLATE 19, FIGURE 6
1849. Cyclostoma simulacrum Morsetet, Testacea novissima insulae Cubanae et
Americae centralis, p. 22.
1852. Megalomastoma simulacrum PFEIFFER, Monographia pneumonopomorum
viventium, vol. 1, p. 181.
1853. Cyclostoma simulacrum PreEtrreR, Martini-Chemnitz Conchylien Cabinet,
vol. 1, sect. 19, p. 267, pl. 36, figs. 11, 12.
1864. Megalomastoma simulacrum Sowersy, Thesaurus conchyliorum, vol. 3,
ul. 263, figs. 3, 4.
1872. Tomocyclus simulacrum Crosser and FiscHer, Journ. Conchyl., vol. 20, p. 76.
1878. Tomocyclus simulacrum KoseEtr, Ilustrirtes Conchylienbuch, p. 195, pl. 61,
fig. 22.
1878. Megalomastoma simulacrum Reeve, Conchologia iconica, vol. 20, p. 8,
fig. 72.
1883. Megalomastoma simulacrum Tryon, Manual Conchology, vol. 2, p. 282, pl.
75, fig. 61.
1886. Tomocyclus simulacrum Crosse and FiscHer, Mission scientifique au
Mexique et dans l’Amérique centrale, vol. 2, pt. 7, pp. 121-124, pl. 40,
figs. 9, 9a; pl. 44, figs. 1, la.
Shell large, covered with a chocolate-brown periostracum, which
grows paler toward the tip. Peristome yellowish bluish white, which
is also the color of the interior of the aperture. Early whorls decol-
lated in all our specimens; those remaining moderately well rounded,
appressed at the summit, and marked by rather irregular incremental
lines. Suture strongly constricted. Periphery well rounded. Base
with a moderately strong keel marking the outer limit of the um-
bilicus. This disappears on the last quarter of the last turn. Aper-
ture circular; peristome double, the inner slightly exserted; the outer
broadly, sinuously flaringly expanded. A slit on the middle of the
parietal wall with the two sides reflected over it partly covers a round
puncture a little behind the edge of the peristome.
The specimen figured, U.S.N.M. No. 203656, was received from
Bland and comes from Guatemala. It has 5.7 whorls remaining
and measures: Height, 88 mm.; greater diameter, 16.6 mm.; lesser
diameter, 11.1 mm. Height of aperture externally, 138.4 mm., in-
ternally, 6.8 mm.; diameter externally, 13.4 mm., internally, 6.7 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 147
The following additional specimens are in the National Museum Col-
lection: No. 321006, from the Redfield Collection; No. 365365, 1 col-
lected by A. A. Hinkley at Chama, Alta Vera Paz, Guatemala; No.
32070, 2 collected by Sarg between Tactic and Tamaju, Alta Vera Paz,
Guatemala.
This species is the largest of the smooth-whorled forms. In size
it resembles 7’. gealei Crosse and Fischer, and in sculpture it agrees
with the other species.
TOMOCYCLUS COPANENSIS (Sowerby)
PLATE 19, Figure 3
1850 Cyclostoma copanense SowerBy, Thesaurus conchyliorum, vol. 1, Suppl.
p. 165*, pl. 31B, figs. 310, 311.
1886. Tomocyclus similacrwm copanensis FISCHER and Crosse, Mission scienti-
fique au Mexique et dans l’Amérique centrale, vol. 2, pt. 7, p. 121, pl.
40, figs. 4, 5, 7, 8, 10.
1886. Tomocyclus simulacrum yar. d FIscHER and Crosse, ibid., p. 121, pl. 40,
fig. 6.
1890. egased (Tomocyclus) simulacrum minus VON MARTENS, Biologia
Centrali-Americana, p. 10.
1890. Meglomastoma (Tomocyclus) simulacrum gracilius VON MARTENS, idem.
Shell small, chestnut-brown. The early whorls decollated in all
our specimens; the latter well rounded, appressed at the summit, and
marked by lines of growth and indistinct spiral striations. Suture
well constricted. Periphery well rounded. Base moderately long,
with a strong keel marking the outer edge of the umbilicus, which
disappears on the last quarter of the last turn. Aperture circular;
peristome double, the inner slightly exserted; the outer flaringly,
somewhat wavily expanded, with a slit on the parietal wall, which
communicates with the circular perforation a little behind the slit.
The two edges of the peristome are bent over to form a siphon.
The specimen described and figured is one of two, U.S.N.M. No.
515763, collected by Godman at Coban, Guatemala. ‘These differ ma-
terially in size, so we are giving the dimensions of both. The speci-
men figured has 5.2 whorls remaining and measures: Height, 26 mm. ;
greater diameter, 12.6 mm.; lesser diameter, 8.7 mm. Height of aper-
ture externally, 9.5 mm., internally, 5.8 mm.; diameter externally,
10 mm., internally, 5.5 mm. The smaller specimen has 5.5 whorls
remaining and measures: Height, 22 mm.; greater diameter, 10.8
mm.; lesser diameter,7 mm. Height of aperture externally, 8.3 mm.,
internally, 5 mm.; diameter externally, 8 mm., internally, 5 mm.
The U. S. National Museum collections contain 3 additional lots: No.
316385, 3 specimens labeled “Guatemala”; No. 515764, 1 specimen
received from Sowerby & Fulton labeled “Guatemala”; No. 321005,
1 specimen from Alta Vera Paz, Guatemala, collected by Cuming.
148 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
There appears to have been considerable misunderstanding about
this species. We have lumped v. Martens’ Megalomastoma (Tomo-
cyclus) simulacrum minus and his M. (T.) simulacrum gracilius
under this name, which we feel justified in doing from the specimens
before us. The small size will readily differentiate this from the
other more or less smooth forms.
Subfamily DIPLOMMATININAE Kobelt
This subfamily embraces a dozen or more genera and some 400
species. Its distribution extends from Siberia south through India,
and in the west Pacific Island complex from Japan to eastern New
Zealand. Only a single genus, Adelopoma, has been reported from
America.
Genus ADELOPOMA Doering
1884. Adelopoma Dorrine, Bol. Acad. Nac. Cienc. Cordoba, vol. 7, p. 457.
1898. Eupalaina KoseLt and MOLLENDoRFF, Nachrb. deutschen malak. Ges., vol.
30, p. 181, in part.
1898. Cylindropalaina KopeLr and MOLLENDOREF, ibid., p. 133, in part.
Small pupiform, sinistral or dextral shells, white, thin. Nuclear
whorls smooth; postnuclear whorls ornamented with scalariform axial
ribs; suture strongly constricted. Aperture circular, with a small
denticle on the columellar wall; peristome double. Operculum thin,
corneous, with 3 or 4 whorls whose outer edge is slightly upturned.
Type: Adelopoma tucma Doering.
Distribution: Costa Rica to Peru.
The radula formula cited by Doering is: 5:4:4:2 for A. twema.
Baker (Occ. Pap. Mus. Zool. Univ. Michigan, No. 137, p. 47, 1922)
gives the radula for A. occidentale Baker from La Fria, Venezuela,
as ‘7:5:5:5. This is our A. baker.
In 1863 Philippi described Pupa lmensis (Malak. Blatter, vol. 14,
p. 75), and this was later referred by Doering and Ancey to the genus
Adelopoma, but Pilsbry has definitely shown that it belongs to the
genus Pupoides in the Pupillidae.
ADELOPOMA TUCMA Doering
PLATE 40, Figure 1
1884. Adelopoma tucma DorrtnG, Bol. Acad. Nac. Ciene. Cérdoba, vol. 7, p. 458,
figs.
1902. Adelopoma tucma Kosett, Das Tierreich, Cyclophoridae, p. 480.
We have not seen this species, but we give a translation of Doering’s
description and copy his figure.
Shell rimate, small, oblong-ovate, pupilliform, thin, white, crystal-
line, with attenuated apex, closely obliquely rib-striate; whorls 6,
convex, the first 2 nuclear, smooth. The postnuclear whorls are ele-
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 149
gantly, closely rib-striately sculptured ; the last exceeds one-fourth the
length of the shell. Suture profound, excavated; aperture circular,
straight; peristome straight, continuous, reflectedly expanded.
Length, 2mm.; diameter, 1mm. Diameter of aperture, 6mm. Oper-
culum small, concave, circular, with concentric and spiral striations,
slightly yellowish.
Radula formula: 5: 4:4: 2.
The locality as given by Doering is Sierra de Tucuman, Argentina.
ADELOPOMA STOLLI (von Martens)
PLatTe 40, Figures 2, 3
1890. Diplommatina stolli von Martens, Biologia Centrali-Americana, p. 20,
pled ig. Lo:
1898. Palaina (Eupalaina) stolli Kosenrt and Métienporrr, Nachrb. Deut.
Malak. Ges., vol. 30, p. 133.
1902. Adelopoma stolli Koprur, Das Tierreich, Cyclophoridae, p. 480.
We have not seen specimens of this species and therefore must
give a translation of v. Martens’ description and copy his figure:
Shell sinistral, imperforate, ovate-conic, with 16 strong ribs on the
penultimate whorl, white; whorls 6, convex, the first 2 smooth, yellow-
ish, the penultimate the largest, the last a little narrower, rounded
and thin; aperture subvertical, circular; peristome continuous, thick-
ened, expanded, white. Length, 2.5 mm.; diameter,1 mm. Aperture
0.75 mm.
Habitat: Northwest Guatemala in the District of Cholhuitz, on
the slope of the Volcano Santa Maria at the plantation Helvetia,
where 2 specimens were found by O. Stoll on the ground in
second growth woods.
Comparing the specimens with Adelopoma occidentale from Trini-
dad, the riblets in A. stolli are very much stronger and less
numerous.
Hinkley (Nautilus, vol. 21, p. 78, 1907), reports this species from
Tampico, where a single specimen was found. We have seen this
specimen, but unfortunately it was so badly fractured that we were
unable to make any comparison. Judging from the distribution,
we rather doubt its identity. It is possible that this represents a
distinct form.
ADELOPOMA BAKERI, new species
Puate 40, Ficure 5
1923. Adelopoma, occidentale H. B. BAKER, Oce. Pap. Mus. Zool. Univ. Michigan,
No. 137, pp. 46-48, fig. 21 (radula).
Shell minute, sinistral, pupoid, white. The nucleus consists of 1.5
inflated mammillated turns, which are smooth. The postnuclear
150 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
whorls are inflated. strongly rounded, and marked by distantly
spaced, slender, scalariform, protractively curved axial ribs, of which
33 are present on the first, 29 on the second, 26 on the third and fourth,
and 12 on the remaining 0.4 of a turn, in the type. Suture very
strongly constricted. Base well rounded, very narrowly umbilicated,
marked by the continuation of the axial ribs. Aperture obliquely
oval; peristome double, the inner expanded, thickened, exserted, and
slightly reflected, adnate to the preceding turn on the parietal wall.
There is a feeble denticle in the middle of the columella. Outer
peristome rather broadly flaringly expanded.
Radula formula: 7:5:5:5.
The type and 27 specimens, No. 140916, are in the collection of the
Philadelphia Academy of Natural Sciences, collected by H. B. Baker
at Quebrada, Venezuela. The type has almost 6 whorls and
measures: Height, 2.3 mm.; diameter, 1.2 mm.
This species is distinguished from A. occidentale in having white
nuclear whorls, and in being stouter, with a much more strongly
developed peristome.
ADELOPOMA COSTARICENSE, new species
PLATE 40, Ficure 4
Shell minute, sinistral, thin, semitranslucent, alabaster white.
Nuclear whorls 1.5, inflated, strongly rounded, smooth. Postnuclear
whorls inflated, strongly rounded, and marked by low, lamellar,
scalariform, decidedly protractively slanting axial ribs, of which 24
are present on the first three whorls and 12 on the last half turn.
The spaces separating the axial ribs are 7 or 8 times as wide as the
ribs, and are covered by exceedingly fine, microscopic, spiral striations.
Suture very strongly constricted. Base inflated, well rounded, with
a narrow umbilical chink, marked by the continuation of the sculp-
ture described for the spire. Aperture broadly obliquely oval; peri-
stome double, the inner strongly exserted and reflected; the outer
moderately broadly expanded, adnate, and appressed to the preceding
turn. Operculum typically adelopomid.
The type, U.S.N.M. No. 516034, comes from Santa Maria, Costa
Rica, at an altitude of 1,550 meters. It has 6 whorls and measures:
Height, 2.8 mm.; diameter, 12 mm. U.S.N.M. No. 516035 contains
9 topotypes.
This species is distinguished from A. occidentale (Guppy) by being
a little slenderer and having fewer axial ribs; these, too, are less
worn at the summit and near the periphery than on the middle, which
is not true of occidentale.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 151
ADELOPOMA OCCIDENTALE (Guppy)
PLATE 40, FIGURE 6
We have described this species from Trinidad (p. 52).
Specimens indistinguishable from it were obtained by the Bond
Venezuelan Expedition of the Academy of Natural Sciences of Phila-
delphia in 1911, at Cariaquita, Venezuela. They are registered as
A.N.S.P. No. 105193 and U.S.N.M. No. 536008.
Cariaquita is not far from the Island of Trinidad, and it is not
surprising that the species should extend to this locality.
Subfamily AMPHICYCLOTINAE Kobelt and Mdllendorff
For the characters of this subfamily, refer to p. 52.
KEY TO THE GENERA OF THE SUBFAMILY AMPHICYCLOTINAE
Spiral sculpture strong.
SHEIUS CISCO 1 eee teeter ek oe eee ee ee Buckleyia
Shell not discoid.
Shell helicoid.
Shell with a notch at base of columella_______---___~ Lagocyclus
Shell without a notch at base of columella.
Shell with two spiral keels on umbilical wall___--~ Filocyclus
Shell without spiral keels on umbilical wall___ Calaperostoma
Spiral sculpture absent or obsolete.
JEieveiSjvopaave TAN oYOUeYS LAR AL ee ee Cyrtotoma
Peristome not reflected.
Posterior angle of aperture with a strong keel_______-__- Barbacyclus
Posterior angle of aperture without a strong keel.
SHELL? (Sra 0 Cae a ee ee Calacyclotus
Shell not smooth but with axial sculpture only.
Axial sculpture not vermiculated.
Axialisculpture vertical = 22222 22s a2 sae Mexcyclotus
Axial sculpture retractives— === 2) == Megacyclotus
Axial sculpture vermiculated________________-_ Amphicyclotus
Genus BUCKLEYIA Higgins
1872. Buckleyia Hiactns, Proc. Zool. Soc. London, 1872, p. 86.
1873. Buckleya Mousson, Malak. Bliitter, vol. 21, p. 17.
Amphicyclotine shells of a discoid shape, having the upper and
lower side almost identical in all features. Operculum with the edge
of the outer turns upturned as flakelike elements.
Type: Aperostoma (Buckleyia) montezwumi Higgins=Buckleyia
martinezi (Hidalgo).
Distribution: Colombia and Ecuador.
KEY TO THE SPECIES OF BUCKLEYIA
SS TTT ISG] Sipe ne aie mec eee ater CE fas EE na ener ea bicincta
Spiral keels 4.
Spirseikeelssbron lh) as Es Se ee ee martinezi
Spiral keelsinweale i). uC reer Ee ee ee ee bifasciata
152 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
BUCKLEYIA BICINCTA, new species
PLATE 19, Fieures 13-15
Shell disk shaped, concave on both the upper and lower surface,
dark olive-green with whitish axial bands, which are rather irregu-
lar in outline and of varying width but of rather regular spacing.
The nucleus consists of almost one turn, which is followed by a
portion having spiral threads, and this in turn is succeeded by a
part which has 2 strong keels at a considerable distance anterior and
posterior to the periphery. The slightly rounded space between these
two keels is marked by 22 slender spiral threads, as well as incremen-
tal lines. On the upper surface there is a moderately strong spiral
thread almost median between the supraperipheral keel and the
suture. There are 20 slender hairlines between this and the supra-
peripheral keel, and 18 between it and the suture on the upper sur-
face. On the lower surface there is a similar arrangement, 22 spiral
threads being present between the strong subperipheral keel and
the median cord and 16 between it and the suture. Suture well im-
pressed, showing the spiral keel on both the upper and lower surface.
Aperture with the upper and lower lip evenly curved; the outer
lip is less arched and rendered eared by the two spiral keels. The
parietal lip corresponds in arching to the outer.
The type, U.S.N.M. No. 316063, comes from Ecuador without
specific locality designation. It has 4.4 whorls and measures: Height,
5.2 mm., greater diameter, 16.5 mm.; lesser diameter, 12.8 mm.
The strong 2-keeled aspect of the shell readily differentiates this
from the other two species.
BUCKLEYIA MARTINEZI (Hidalgo)
Puate 19, Figures 19-21
1866. Cyclophorus martinezi Hmateo, Journ. Conchyl., vol. 14, p. 273, pl. 8, fig. 5.
1872. Aperostoma (Buckleyia) montezumi Hicerns, Proc. Zool. Soc. London,
1872, p. 686, pl. 56, fig. 7.
1873. Buckleya martinezi Mousson, Malakoz. Blatter, vol. 21, p. 17.
Shell disk shaped, concave on both the upper and lower surface,
sides olive-green with paler axial bands; these vary considerably in
width and spacing. The nucleus consists of a fraction of a turn and
is smooth. This is followed by a spirally lirate portion, which in
turn is succeeded by a part bearing 4 strong keels, which are of about
equal strength and spacing. Two of these are on each side of the
periphery and the other two separated from them by a space as wide
as the one that separates these two keels, one above and the other
below on the base. The spaces between these keels are concave and
marked by several fine spiral threads and rather strong threadlike
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 153
incremental lines. Between the suture and the first strong keel, a
little nearer to the latter, there is another spiral thread on the dorsal
surface, which is materially stronger than the fine spiral hairlines cov-
ering the rest of the whorl in this region. Of these fine spiral hair-
lines, 19 are present between the suture and the stronger thread
and 17 between that and the first strong keel, in the specimen which
we are figuring. On the ventral side there is also a median stronger
thread corresponding to one on the upper side and between this
and the suture 19 spiral hairlines are present, while the space be-
tween this and the first strong keel bears 15 spiral hairlines. The
suture is strongly channeled on both the upper and the lower surface,
and shows the spiral keel. Aperture circular; peristome simple,
rendered fluted on the peripheral side. Operculum with central
nucleus, many whorls with the outer edge of the later whorls up-
turned and projecting as flakelike elements.
The specimen described and figured, U.S.N.M. No. 420868, bears
the locality label “Ecuador.” It has 4.5 whorls and measures: Height,
7.5 mm.; greater diameter, 24.7 mm.; lesser diameter, 17.9 mm.
The four strong keels readily differentiate this from the other spe-
cies here discussed.
BUCKLEYIA BIFASCIATA Mousson
PLatge 19, Ficures 16-18
1873. Buckleya bifiasciata Mousson, Malak. Blitter, vol. 21, p. 17.
1875. Buckleya bifasciata PretrrEer, Nov. Conch., vol. 4, p. 132, pl. 129, figs. 15-18.
1876. Cyclophorus bifasciata Pretrrer, Monographia pneumonopomorum vViven-
tium, vol. 4, p. 122.
1897. Buckleyia bifasciata Kosett and M6LLENDorFF, Nachrb. deutschen malak.
Ges., vol. 29, p. 189.
Shell disk shaped, concave on both the upper and lower surface,
of olivaceous ground color, marked with darker spiral zones of vary-
ing width. There are two broad paler bands at some little distance
above and below the periphery, which are edged with darker zones,
which consist of a series of interrupted elongated beadlike elements.
The ground color is not uniform but has a marbled effect. The
nuclear whorls consist of a smooth fraction of a turn followed by a por-
tion bearing spiral hairlines and this in turn is succeeded by whorls that
have 4 low rounded cords (not keels), the space between the two
central ones being wider than that which separates the two adjacent
to these. The central ones are equally distant from the periphery.
Between the central ones 8 spiral threads are present, while between
the two cords on the dorsal surface 5 spiral threads are present.
The rest of the dorsal surface also bears spiral threads, of which
13 are present on the anterior half of the turns, while on the pos-
154 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
terior half they are obsolete. On the ventral surface 7 fine spiral lines
are present between the two cords and 12 on the anterior third of the
rest of the whorls, while posterior to this the spiral sculpture is
obsolete. Suture well impressed, showing the spiral cord. Aperture
almost circular, slightly compressed dorsoventrally ; peristome simple.
The specimen described and figured, U.S.N.M. No. 414156, was
collected by H. Damel at Jerico, Antioquia, Colombia. It has 4.4
whorls and measures: Height, 7.5 mm.; greater diameter, 23 mm.;
lesser diameter, 17 mm.
The weaker keels and the two strong color bands readily distinguish
this species from B. martinezi (Hidalgo).
LAGOCYCLUS, new genus
Amphicyclotine shells of helicoid shape, with spiral striations or
lirations. There is a notch in the peristome at the junction of the
inner and basal lip, which becomes bridged over behind the peristome
to form a conspicuous keel extending over all the whorls.
Type: Lagocyclus crosseanus (Hidalgo).
Distribution: Ecuador and Peru.
KEY TO THE SPECIES OF LAGOCYCLUS
Strong spiral threads present on last whorl.
Subperipheral spiral threads present.
Spiral..cords Strong van reg rl ais se oe eee antoni
Spiral cords weak and irrerular=- =) 22) eee haematommus
Subperipheral spiral threads absent==—===22—=="=s2— 272) = crosseanus
Strong spiral threads absent on last whorl.
Shell larce, jereater diameter s0!Gumm == ees ee eee vasconesi
Shell smaller) greater) diameter 20:0 mm=) 20sec bartletti
LAGOCYCLUS ANTONI (Cousin)
PLATE 20, Figures 6-8
1887. Cyclophorus antonii Cousin, Bull. Soc. Zool. France, 1887, p. 85, pl. 4, fig. 4.
Shell rather small, covered with an olivaceous epidermis; when this
is denuded the shell is soiled white on the last turn, the rest of the
turns being rosy red. The nucleus consists of about one turn, which is
smooth. The postnuclear whorls are marked by spiral cords of which
7 are present on the next to the last whorl and 8 on the last. On
this there is also a finer thread between the heavier cords. In addi-
tion to this, the spire is marked by fine incremental lines. Suture
moderately impressed. Periphery rounded. Base somewhat inflated,
well rounded, and marked with 8 spiral cords equaling those of the
spire immediately below the periphery, and a broad heavy keel mark-
ing the outer edge of the umbilicus. The arched space between this
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 155
keel and the spiral cords is marked by obsolete spiral threads and
coarse incremental lines. The spiral keel marking the outer edge of
the umbilicus terminates in a notch at the peristome. The umbilicus
is moderately broad and open to the tip. Its wall is marked by rib-
like incremental lines. Aperture somewhat oblique, almost circular ;
peristome simple, notched at the umbilical cord, the parietal wall
covered with a moderately thick callus. Operculum unknown.
The specimen described and figured, U.S.N.M. No. 528512, was
received from Sowerby & Fulton with the locality label “Ecuador.”
It has 5 whorls and measures: Height, 11 mm.; greater diameter, 18.7
mm.; lesser diameter, 13.9 mm.
The presence of spiral cords below the periphery readily distin-
guishes this species from L. crosseanus (Hidalgo).
LAGOCYCLUS HAEMATOMMA (Pfeiffer)
PLATE 20, FiauRES 1-3; PLATE 41, FicurEs 1-3
1862. Cyclophorus haematomma P¥etrrerR, Proc. Zool. Soc. London, 1862, p. 276.
1863. Cyclophorus haematomma Prrtrrer, Noy. Conch., vol. 2, p. 217, pl. 57, figs.
12-14.
1897. Amphicyclotus haematomma Kosett and MOLLenpDorFF, Nachrb. deutschen
malak. Ges., vol. 29, p. 139.
1912. Amphicyclotus haematomma Kose.t, Martini-Chemnitz Conchylien Cabinet,
vol. 1, sect. 19, p. 924, pl. 188, figs. 10-12.
Shell helicoid, the early whorls bright brown, gradually fading to
flesh color on the last turn. The nucleus consists of about one turn,
which is well rounded and smooth. The first postnuclear whorl is
also rounded and smooth, but the succeeding turns are well rounded
and marked by spiral threads which vary considerably in size. Of
these 7 are present on the third and fourth turns and 15 between the
summit and the periphery on the last whorl. In addition to this the
upper surface is marked by regular lines of growth which lend it a
somewhat rough aspect. Suture moderately well impressed. Periph-
ery well rounded. Base openly umbilicated, well rounded. The
posterior third is marked by 4 strong spiral cords equaling those on
the spire. Anterior to this there are microscopic spiral lines. There
is a strong axial keel marking the outer edge of the umbilicus and
sightly notching the aperture. The umbilical wall is marked by
coarse lines of growth. Aperture oblique, subcircular, slightly
auriculated at the posterior angle and notched at the basal cord. The
peristome is slightly thickened on the inner lip.
The specimen described and figured, U.S.N.M. No. 536032, is marked
“Ecuador,” without specific locality. It has 5.4 whorls and measures:
Height, 14.6 mm.; greater diameter, 20.5 mm.; lesser diameter, 16.4
mm. Height of aperture, 8.4 mm.; diameter, 8.5 mm.
156 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
L. haematomma is nearest related to ZL. antoni (Cousin), from
which its finer, irregular spiral threads readily distinguish it.
LAGOCYCLUS CROSSEANUS (Hidalgo)
PLATE 20, Ficurres 9-11
1866. Cyclophorus crosseanus Hipareo, Journ. Conchyl., vol. 14, p. 348, pl. 14,
fig. 1.
1897. ERE Ee crosseanus KoBeLt, Nachrb. deutschen malak. Ges., vol. 29,
p. 139.
Shell of helicoid outline, covered with an clive-brown periostracum,
white when this is removed, the basal portion being considerably
darker than the spire. There is also a subperipheral chestnut-brown
band present, which gradually fades toward the base. The nucleus
consists of about one turn and is smooth. The postnuclear whorls are
marked by spiral cords, of which 7 are present on the first and second,
8 on the third, and 12 on the last whorl between the summit and the
periphery. These cords are a little less strong near the summit and
stronger at the periphery. In addition to this the spire is marked
by fine incremental lines. Suture strongly impressed. Periphery
rendered angulated by a spiral cord. Base strongly rounded and
marked by incremental lines only. There is a strong spiral keel at
the outer edge of the umbilicus, which terminates in a notch at the
peristome. Umbilicus moderately broadly openly umbilicated, its
wall marked by almost riblike incremental lines. Aperture slightly
oblique, circular; peristome simple, except the notch referred to above;
parietal wall covered with a thin callus. Operculum corneous.
The specimen described and figured, U.S.N.M. No. 307412, was re-
ceived from Sowerby and Fulton with the locality label “Ecuador.”
It has 5.2 whorls and measures: Height, 14.7 mm.; greater diameter,
22.5 mm.; lesser diameter, 17.0 mm.
This species is readily distinguished from Z. antoni (Cousin) by
lacking the spiral threads on the posterior base.
LAGOCYCLUS VASCONESI (Jousseaume)
PLATE 20, FrcurES 12-14
1897. Cyclophorus vesconesi JOUSSEAUME, Le Naturaliste, vol. 19, p. 250.
1898. Cyclophorus vasconesi JOUSSEAUME, Le Naturaliste, vol. 20, p. 81 (cor-
rection).
1899. Amphicyclotus vesconesi KopeLT and MOLLENDORFF, Nachrb. deutschen
malak. Ges., vol. 31, p. 136.
Shell of helicoid outline, covered with a chestnut-brown periostra-
cum on the later turns; the early turns are reddish. It is a little
paler on the underside than on the spire, and there is a light zone at
the periphery bordered by a moderately broad dark band anterior to
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA kez
this. In addition there are obscure darker spiral lines on the upper
whorls. The nucleus consists of about one smooth turn followed by
whorls which bear 5 moderately strong spiral cords, which are of
equal strength and spacing. The space between the first of these and
the summit is about three times as wide as that separating the cords.
On the last whorl, however, these spiral cords disappear. In addition
to this the shell is marked by incremental lines. Suture moderately
impressed. Periphery well rounded. Base moderately convex, marked
by incremental lines only. sdates setae ecuadorense
Umbilicus not narrow.
Upper surface closely finely ribbed--__ subcingulatum
Upper surface obsoletely finely ribbed_---_- quitense
Axial ribs of umbilical wall not stronger than
those on upper surface.
Last whorl regularly finely ribbed.
Shell pale olive colored__-_-------------- olivaceum
Shell browne eee Be ee eee castaneum
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 223
Upper surface not malleated—Continued.
Posterior angle of aperture not auriculate—Continued.
Shell not decidedly depressed—Continued.
Aperture large—Continued.
Axial ribs of umbilical wall not stronger than
those on upper surface—Continued.
Last whorl not regularly finely ribbed.
Umbilicusivery,, broad42-2 = peruvianum
Umbilicus not very broad.
Summit of last whorl appressed to
preceding: turns=s2 ae esse eae salengoense
Summit of last whorl not appressed
to preceding turn.
Shellphelicoi deers 2 stake ees masvense
Shell depressed-helicoid.
Suture impressed_4------------ agassizi
Suture not impressed__---~-_ nevadense
Aperture not large.
Last whorl deflected below subperipheral brown
band.
Wimpilicis mManrowae eee eee fasciatum
Umbilieus moderately broad: = 2-2" 2 eas ene eet ee pazi
Last whorl not deflected below subperipheral
brown band.
Base with a median brown band.
Peristome with a sinus below basal brown
Deana ee es he Sal Se Does 2 neces caucaense
Peristome without a sinus below basal
brown band.
Greater diameter more than 27 mm_ colombiense
Greater diameter less than 25 mm.
Upper surface regularly finely ribbed_ perezi
Upper surface not regularly finely
ribbed.
Suture strongly impressed____ boliviense
Suture not strongly impressed_ redfieldi
Base without median brown band.
Greater diameter more than 27 mm____-_--___ currani
Greater diameter less than 25 mm.
Aperture deflected below subperiph-
eraliibrowny bande 22 Sen s0 8 wa = nanum
Supraperipheral light zone very
CONSPICUOUS 2S fa ss Re eee eee merrilli
Supraperipheral light zone obso-
Tet Giese haan Ul oes ei pulchellum
Aperture not deflected below subpe-
ripheral brown band.
Upper surface with oblique
scratches His: SoU aed popayanum
Upper surface without oblique
scratches.
Dark band covering half of
DA SC a heteniee Teuneee enna) balsasense
Dark band not covering half
OL DasSexs aera Bue oases cardozi
224 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
THE GROUP OF APEROSTOMA (APEROSTOMA) UMBILICATUM
This group embraces the species having the upper surface of the
last whorl malleated.
APEROSTOMA (APEROSTOMA) UMBILICATUM, new species
PLATE 31, FiauRES 1-3
Shell very depressed-helicoid, very dark brown; the early whorls
paler; interior of aperture bluish white. The nucleus consists of 2
small, well-rounded, smooth turns; the postnuclear whorls are mod-
erately rounded and marked by retractively slanting lines of growth
and numerous, very rough malleations. Suture strongly impressed.
Periphery rendered decidedly angulated by a low keel. Base very
openly umbilicated, somewhat inflated, strongly rounded, and marked
by the continuation of the axial sculpture and strong malleations,
which on the posterior half also produce several impressed spiral
lines. The umbilical wall also is malleated. Aperture irregularly
broadly tear-shaped and drawn out into a rather broad angle at the
posterior angle. There is a groove in the center of this angulation;
peristome simple, very slightly more thickened on the inner lip than
on the outer. The parietal wall is covered by a rather thick callus,
which renders the peritreme complete. Operculum typically aperos-
tomid, of about 8 whorls.
The type, U.S.N.M. No. 307428, comes from Bogota, Colombia.
It has 4.7 whorls and measures: Height, 20.5 mm.; greater diameter,
38.5 mm.; lesser diameter, 27.7 mm. Height of aperture, 14.8 mm.;
diameter, 16.8 mm.
This species is distinguished from all the other members of the
group by its very strong malleations, which lend to the surface a
decidedly rugose aspect.
APEROSTOMA (APEROSTOMA) CUMINGI, new species
PLATE 31, Ficures 19-21
Shell of medium size, depressed-helicoid, covered with a very dark
olivaceous-brown periostracum; the early whorls being denuded of
this appear red; interior of aperture bluish white. The nucleus con-
sists of 2.5 well-rounded smooth turns. Postnuclear whorls well
rounded, the first marked by retractively curved, somewhat irregular,
feeble axial riblets; the last by incremental lines and oblique scratches
and malleations, which render this whorl decidedly rough. Suture
deeply impressed. Periphery rendered angulated by a low keel,
which slopes more abruptly dorsally than ventrally. Base narrowly
umbilicated, inflated, strongly rounded, and marked like the spire,
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 225
except that on the posterior portion the malleations form several
more or less impressed lines. The umbilical wall is rough and shows
less of the malleation. Aperture circular with a slightly obtuse angle
at the posterior angle; peristome simple; that of the outer lip thin,
while the inner is thickened and continues as a heavy callus over the
parietal wall, which renders the peritreme complete.
The type, U.S.N.M. No. 307416, was collected by Cuming in
Colombia. It has 4.3 whorls and measures: Height, 19.3 mm.;
ereater diameter, 30.2 mm.; lesser diameter, 20.8 mm. Height of
aperture, 12.5 mm.; diameter, 14.8 mm.
This species comes nearest to A. (A.) umbélicatum in strength of
the malleations and roughness but is easily distinguished from that
by its slightly more elevated form, weaker sculpture, and much nar-
rower umbilicus.
APEROSTOMA (APEROSTOMA) PAEZENSE, new species
PLATE 31, Ficures 7-9
Shell very depressed-helicoid ; the denuded nuclear whorls reddish ;
the rest of the postnuclear whorls are covered by a rather thick
periostracum, that of the first whorl being olivaceous brown, while
the last whorl is dark chesnut brown. There is a slender narrow
zone of pale olive yellow immediately above the peripheral keel.
There is a very broad almost blackish brown zone occupying the
peripheral keel and about one-fourth of the base anterior to the
keel. A second similar dark narrow zone is separated from the
broad one by a moderately broad space. The rest of the base is paler
than the upper surface; interior of aperture bluish white. The
nucleus consists of a little more than 2 small, well-rounded, smooth
turns. The postnuclear whorls are well rounded, the last one slightly
flattened toward the suture, dropping a trifle below the periphery
on the last fifth of the last turn. The first postnuclear whorl is
marked by feeble retractively slanting lines of growth, which gain
in strength on the next turn, where they are a little heavier than
hair lines and irregular in distribution and development. On the last
turn fine malleations and scratches appear, in addition to the above
sculpture. Suture well impressed. Periphery weakly angulated.
Base broadly openly umbilicated, slightly inflated, well rounded, and
marked by the continuation of the lines of growth which assume the
strength of riblets on the umbilical wall. Aperture irregularly tear-
shaped, decidedly oblique, drawn into an angle at the posterior angle
and somewhat angulated at the junction of the basal and outer lip;
peristome simple; the outer lip thin; the inner thickened and reflected.
The parietal wall is covered by a rather thick callus, which renders
226 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
the peritreme complete. Operculum typically aperostomid, having
about 8 whorls.
The type, U.S.N.M. No. 251169, was collected by H. Pittier in the
valley of the Rio Paez, in the Central Cordilleras, Colombia. It
has 4.5 whorls and measures: Height, 20.9 mm.; greater diameter, 37.3
mm.; lesser diameter, 26.8 mm. Height of aperture, 13.8 mm.;
diameter, 17.7 mm.
In the open umbilication, this species resembles A. (A.) wmbili-
catum, from which it is easily differentiated by its color scheme and
the much larger aperture, as well as the much finer sculpture.
APEROSTOMA (APEROSTOMA) BOGOTENSE (Pfeiffer)
PLATE 31, FIGURES 4-6
1855. Cyclostoma (Cyclotus) bogotense PFEIFFER, Proc. Zool. Soc. London, vol.
2a, nalts
1858. Cyclotus bogotensis PFEIFFER, Monographia pneumonomoporum viventium,
WOL, 2; pi i
1863. Cyclotus bogotensis REEVE, Conchologia iconica, vol. 14, sp. 20.
1897. Neocyclotus (Neocyclotus) bogotensis Kosett and MOLiLEeNporFr, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
1923. Poteria (Neocyclotus) bogotensis, H. B. BAKER, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 187, p. 44.
Shell small, depressed-helicoid, varying from olivaceous to oliva-
ceous-brown, with a dark zone at the periphery and usually a nar-
row light one above it. The base is bicolor, that of the posterior
half being olivaceous, while the anterior half is much paler; interior
of aperture bluish white. The nucleus consists of 2.3 rather large,
well-rounded, smooth turns. The postnuclear whorls are well
rounded, the last one being slightly flattened below the summit and
marked by strong, small malleations, which give the upper surface
of the shell not only a malleated but a somewhat vermiculated aspect.
The last whorl! is slightly decurrent on the last quarter of the turn.
Suture very strongly impressed. Periphery weakly angulated. Base
moderately broadly umbilicated, somewhat inflated, well rounded,
and marked by almost vertical, irregular, incremental lines, which are
a little stronger on the umbilical wall, and feeble malleations. Aper-
ture circular with a slight angulation at the posterior angle; peri-
stome simple, the outer lip thin, the inner slightly thickened posteri-
orly. The parietal wall is covered by a thick callus which renders
the peritreme complete.
The specimen described and figured is one of three, U.S.N.M. No.
816133, which comes from Bogota, Colombia. It has 3.8 whorls and
measures: Height, 13.8 mm.; greater diameter, 20.5 mm.; lesser
diameter, 14.7 mm. Height of aperture, 9.2 mm.; diameter, 9.4 mm.
U.S.N.M. No. 524128 contains 1 specimen without specific locality.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 227
The small size will easily distinguish this from the other members
except A. (A.) inconspicuum (Sowerby), which is even smaller.
APEROSTOMA (APEROSTOMA) SIMILE, new species
PLATE 31, Ficures 13-15
Shell helicoid, small, covered with a horn-colored periostracum,
which on the last whorl becomes decidedly olivaceous. There is a
broad, pale, darker band on the posterior half of the base. The
nucleus consists of about 1.5 smooth turns. The postnuclear whorls
are marked by retractively slanting incremental lines, which become
increasingly closer spaced as the shell advances in age. In addition
to these incremental lines the whorls are marked by rugosities sepa-
rated by more or less strongly impressed scratches. Suture well
impressed. Periphery feebly angulated. Base somewhat inflated,
rather openly umbilicated, well rounded, and marked by the con-
tinuation of the incremental lines, which extend upon the umbilical
wall, and mere indications of rugosities. Aperture subcircular, ob-
lique; peristome simple, the outer thin; the inner somewhat thick-
ened, more so on the parietal wall. Operculum typically aperostomid.
The type, U.S.N.M. No. 535989, was received from Sowerby &
Fulton with the locality “Bogota” (Colombia). It has 4.1 whorls
and measures: Height, 13 mm.; greater diameter, 19.3 mm.; lesser
diameter, 15 mm.
U.S.N.M. No. 307398 contains another specimen from the same
locality, while U.S.N.M. No. 307400 contains a specimen bearing the
label “New Grenada.”
In size and general coloration this species agrees fairly well with
A. (A.) bogotense (Pfeiffer). The less developed rugose sculpture
and more scattered nodulation will readily distinguish it from this.
.
APEROSTOMA (APEROSTOMA) INCONSPICUUM (Sowerby)
PLATE 31, F1acurEs 10-12
1848. Cyclostoma inconspicuum SowrErsBy, Thesaurus Conchyliorum, vol. 1, p.
109, pl. 24, figs. 73-74.
1912. Neocyclotus papayanus fasciata Koxsett (pars), Martini-Chemnitz Con-
chylien Cabinet, vol. 1, sect. 19, pt. 3, p. 901.
1923. Poteria (Neocyclotus) inconspicuum, H. B. BaxKer, Oce. Pap. Mus. Zool.
Univ. Michigan, No. 137, p. 44.
Shell small, helicoid, covered by a pale-brown periostracum. There
ig a narrow pale chestnut zone immediately below the periphery,
and above this a white band of about the same width. The posterior
half of the base below the dark band is olivaceous, gradually paling
toward the umbilicus. The nucleus consists of about 1.5 small, in-
flated, well-rounded, smooth turns. The early postnuclear whorls
228 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
are marked by regular, almost vertical, slender riblets, which are
separated by spaces no wider than the riblets. On the succeeding
turns the axial riblets become reduced to incremental lines, and on
the last one in addition to these oblique, irregularly scattered, im-
pressed scratches are present, which cause the spaces between them
to appear as basal rugosities. Periphery feebly angulated. Base
inflated, strongly rounded, broadly openly umbilicated, and marked
like the spire. The umbilical wall is marked by incremental lines.
Aperture large, subcircular, slightly auriculated at the posterior
angle; peristome simple, thin all around.
The specimen described and figured, U.S.N.M. No. 116667, comes
from the Lea collection and bears no locality label. It has 4.4 whorls
and measures: Height, 12 mm.; greater diameter, 15.9 mm.; lesser
diameter, 12.5 mm.
The small size readily distinguishes this from all the other
Aperostomas.
APEROSTOMA (APEROSTOMA) CINGULATUM (Sowerby)
PLATE 31, FiaurEs 22-24
1843. Cyclostoma cingulatum SowrrsBy, Thesaurus conchyliorum, vol. 1, p. 93,
pl. 29, figs. 213-214.
1852. Cyclotus cingulatus PFEIFFER, Monographia pneumonopomorum viventium,
vol. 1, 18.
1866. Cyclotus pee BuLanp, Amer. Journ. Conch., nor 2, p. 59 (error for
cingulatus; not angulatus von Martens 1874).
1897. Neocyclotus (Neocyclotus) cingulatus Kospetr and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 187.
1923. Poteria (Neocyclotus) cingulatum H. B. BaKker, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 137, p. 42.
Shell depressed-helicoid, covered with a very dark chestnut-brown
periostracum; the early denuded whorls reddish. The nucleus con-
sists of 2 small, well-rounded, smooth turns. The postnuclear whorls
are well rounded, the last slightly flattened below the suture. The
first postnuclear whorl is marked by slender, retractively curved,
somewhat irregular, fine, rather closely spaced riblets, which become
very much enfeebled on the last half of the last turn and toward
the end completely lost. In adition to this, the upper surface of
the last whorl is irregularly slightly malleated. Suture well im-
pressed. Periphery very feebly angulated. Base moderately broadly
openly umbilicated, inflated, and eed by rather irregular wavy
axial riblets, whieh become very much intensified on the umbilical
wall, and rather conspicuous malleations stronger than those on the
upper surface. Aperture oblique, circular, slightly angulated at the
posterior angle; peristome simple, thin on the outer lip and thickened
on the inner, with a very heavy callus on the parietal wall, which
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 229
renders the peritreme complete. The operculum is typically aperos-
tomid of about 8 whorls.
The specimen described and figured, U.N.S.M No. 307422, comes
from Cauca, Colombia. It ‘has 4.4 whorls and measures: Height,
24.7 mm.; greater diameter, 43.8 mm.; lesser diameter, 29.3 mm.
Height of aperture, 17.5 mm.; diameter, 20.5 mm.
There are four additional lots in the National Museum collection :
No. 414158, one specimen from Jerico, Antioquia, Colombia. No.
424727, two specimens collected by H. Daniel at Jerico, Antioquia.
One of these is much larger than the one figured. It has 4.8 whorls
and measures. Height, 31.2 mm.; greater diameter, 52 mm.; lesser
diameter, 35 mm. Height of aperture, 20 mm.; diameter, 26.2 mm.
No. 307419, one specimen labeled “Colombia.” No. 307457, labeled
“South America.”
This species belongs to the group that is only feebly malleated on
the upper surface. The dark color will distinguish it from A. (A.)
veracochanum, which is very much thinner-shelled and much paler.
APEROSTOMA (APEROSTOMA) VERACOCHANUM, new species
PLATE 31, FicuREs 16-18
Shell very depressed-helicoid, covered with a pale olivaceous-brown
periostracum, which on the last part of the last whorl becomes de-
cidedly olivaceous both on the upper and lower surfaces. The early
denuded whorls are reddish; interior of the aperture bluish white.
The nucleus consists of a little more than 2 moderately well-rounded
smooth turns; the post-nuclear whorls are moderately rounded and
marked by weak retractively curved axial riblets, which find their
strongest expression on the first half of the last turn. On the last
half, they are enfeebled. Suture strongly impressed, slightly chan-
neled on the last part of the last whorl. Periphery very weakly
angulated. Base openly umbilicated, strongly rounded, and marked
by the continuation of the axial ribs, which on the umbilical wall
become much strengthened and fused. Aperture very broadly tear-
shaped, with a conspicuously drawn-out angle at the posterior angle,
which has a groove in its middle; peristome simple; the outer lip
thin; the inner slightly thickened. The parietal wall is covered by
a callus which renders the peritreme complete.
The type, U.S.N.M. No. 524045, was collected by the Wilkes Explor-
ing Expedition at Maguas, Peru. It has 4.5 whorls and measures:
Height, 24.4 mm.; greater diameter, 43.2 mm.; lesser diameter, 31.5
mm. Height of aperture, 16.3 mm.; diameter, 18.5 mm.
This species has a much thinner shell and is much paler than
A, (A.) cingulatum (Sowerby).
230 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) ALLANTAYUM, new species
PLATE 32, FicurEs 4-6
Shell of medium size, very depressed-helicoid; the early denuded
whorls are soiled flesh colored; the succeeding turns dark straw
colored with an orange flush, the last reddish chestnut-brown with
a narrow light zone immediately above the narrow dark band at
the periphery; posterior half of the base a little darker than the
upper surface, while the anterior is considerably paler; interior of
aperture bluish white. The nucleus consists of about 2 small, well-
rounded, smooth turns; the postnuclear whorls are moderately well
rounded, all but the last marked by very regular, retractively slanting
axial riblets, which are about as wide as the spaces that separate
them. On the last whorl these become much enfeebled and reduced
to irregular lines of growth. Here, too, we have slight indications
of malleations. The last whorl is slightly decurrent on the last
quarter of a turn. Suture well impressed. Periphery feebly ob-
soletely angulated. Base openly umbilicated, inflated, well rounded
and marked by irregular, wavy, strong incremental lines, which are
developed into irregular ribs on the umbilical wall. Aperture de-
cidedly oblique, very broadly tear-shaped, with a pronounced drawn-
out angulation at the posterior angle, which has a groove in its
middle on the inside; peristome simple; the outer lip thin; the
inner somewhat thickened. The parietal wall is covered by a thick
callus which renders the peritreme complete.
The type, U.S.N.M. No. 524046, comes from the Rich collection
and is labeled “Peru.” It has 4.4 whorls and measures: Height,
17.4 mm.; greater diameter, 30.8 mm.; lesser diameter, 21.2 mm.
Height of aperture, 11.7 mm.; diameter, 13.9 mm.
U.S.N.M. No. 22208 contains another specimen received from Mac-
Andrew labeled “Peru.”
The conspicuous bicolored base readily distinguishes this from the
other feebly malleated members of this group.
THE GROUP OF APEROSTOMA (APEROSTOMA) DUNKERI
Umbilical wall strongly distinctly axially ribbed.
APEROSTOMA (APEROSTOMA) DUNKERI (Pfeiffer)
PLATE 32, Fiaures 1-3
1856. Cyclotus dunkeri Pretrrer, Malak. Blatter, vol. 3, p. 256.
1864. Cyclotus dunkeri Reeve, Conchologia iconica, vol. 14, pl. 2, sp. 9.
1897. Neocyclotus (Neocyclotus) dunkerit KoBett and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 187.
1923. Poteria (Neocyclotus) dunkeri H. B. Baker, Occ. Pap. Mus. Zool. : Univ.
Michigan, No. 1387, pp. 39, 40, 43.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 231
Shell depressed-helicoid, the early whorls soiled flesh white; the
later turns gradually becoming chestnut brown. There is a broad
spiral dark zone covering the posterior half of the base, which grad-
ually becomes paler from the periphery basally. The rest of the base
is much paler, so that there is a bicolor effect on the base; interior of
aperture decidedly bluish white. The nucleus consists of 2 small,
well-rounded, smooth turns. The postnuclear whorls are weakly
rounded, the last one flattened a little below the summit, and marked
by retractively curved, rather regular axial riblets, which are a little
wider than the spaces that separate them. On the last turn, however,
these riblets gradually become enfeebled, and on the last portion of this
whorl they are merely represented by irregular lines of growth. The
last whorl is slightly decurrent on the last fifth of a turn. Suture
strongly impressed. Periphery rounded; the peculiar banding at the
periphery, however, gives it the effect of being angulated, which is an
optical illusion. Base very openly umbilicated, somewhat inflated, well
rounded ; the posterior half is marked by the continuation of the axial
ribs which are stronger than on the upper surface, and on the anterior
half become fused into strong, distinct, broad, rounded ribs, which
cross the sloping umbilical wall. Aperture very broadly tear-shaped
with a decidedly drawn-out auricle at the posterior angle, which has
an impressed groove in its middle; peristome simple, the outer lip
thin; the inner thickened. The parietal wall is covered by a thick
callus, which renders the peritreme complete. Operculum typically
aperostomid, having 10 whorls.
The specimen described and figured is one of two, U.S.N.M. No.
251170, collected by H. Pittier in the Rio Paila Valley, Colombia, at
an altitude of 1,800 meters. It has 4.8 whorls and measures: Height,
20.4 mm.; greater diameter, 38.5 mm.; lesser diameter, 26mm. Height
of aperture, 13 mm.; diameter, 17.5 mm.
The larger size of this species distinguishes it from the other two
members here described as having strong ribs on the umbilical wall.
APEROSTOMA (APEROSTOMA) PAILAENSE, new species
PLatTe 32, Ficures 7-9
Shell quite small, depressed-helicoid. The shreds of periostracum
remaining would indicate that this was straw-colored with a gold
tinge. The denuded surface is uniformly bluish white; interior of
aperture bluish white. The nucleus consists of 2 small, well-rounded,
smooth turns; the postnuclear whorls are moderately well rounded;
the last slightly flattened below the summit, marked by retractively
curved slender axial riblets, which are about as wide as the spaces
that separate them. On the last half of the last whorl these riblets
202 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
become much enfeebled and form rather irregular, broad incremental
elements. Suture strongly impressed. Periphery with a weak
angulation. Base moderately openly umbilicated, inflated, well
rounded, and marked by strong axial ribs which gradually increase
in intensity toward the umbilicus, being strongest on the sloping
umbilical wall. Aperture very broadly oval, with a conspicuous
auricle at the posterior angle, which bears a groove in its middle;
peristome simple; the outer lip thin; the inner thickened. The
parietal wall with a very thick callus rendering the peritreme
complete.
The type, U.S.N.M. No. 251171, was collected by H. Pittier in the
Rio Paila Valley, Colombia, at an altitude of 1,300 meters. It has
4.3 whorls and measures: Height, 16.3 mm.; greater diameter, 27.8
mm.; lesser diameter, 18.9 mm. Height of aperture, 11.7 mm.;
diameter, 13.9 mm.
This species is readily distinguished from the other two with
strong ribs on the umbilical wall by its exceedingly small size.
APEROSTOMA (APEROSTOMA) PAEZICOLUM, new species
PLATE 32, FiauRESs 13-15
Shell depressed-helicoid, the early whorls reddish; the last whorl
olivaceous with axial streaks of brown, which vary in width and
spacing. There is a narrow light zone immediately above the
periphery. Below the periphery the posterior half of the base has
a dark chestnut-brown area; the anterior half is olivaceous brown
and much paler. The axial darker streaks mentioned for the upper
surface also are apparent on the lower. Interior of aperture bluish
white. The nucleus consists of 2.1 small, well-rounded, smooth turns.
The postnuclear whorls are moderately well rounded; the last is
slightly decurrent on the last half whorl and slightly flattened below
the summit. The postnuclear whorls are marked by slender, re-
tractively curved axial riblets, which are a little wider than the
spaces that separate them. On the last whorl these become less
strongly developed and more irregular. There is also a weak indica-
tion of a few scratches, showing the affinity of the species to the mal-
leated form. Suture strongly impressed. Periphery well rounded.
Base openly umbilicated, somewhat inflated, well rounded, and
marked by the continuation of the sculpture characterizing the upper
surface; the riblets, however, become fused on the umbilical wall
to form strong and quite regular rather broad ribs on its sloping
surface. Aperture almost circular with a conspicuous auricle at
the posterior angle, which bears a groove in its middle; peristome
simple; outer lip thin; the inner thickened. The parietal wall is
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 233
covered by a thick callus, which renders the peritreme complete. The
operculum is typically aperostomid, having 10 whor!s.
The type, U.S.N.M. No. 524047, was collected by H. Pittier in the
Rio Paez Valley, Colombia, at an elevation of 2,500 meters. It has
4.7 whorls and measures: Height, 18.7 mm.; greater diameter, 33.1
mm.; lesser diameter, 22.2 mm. Height of aperture, 12.8 mm.;
diameter, 15.6 mm.
Its smaller size will readily differentiate this from A. (A.) dunkera
(Pfeiffer).
GROUP OF APEROSTOMA (APEROSTOMA) CARMIOLI
Umbilical wall closely finely axially ribbed.
APEROSTOMA (APEROSTOMA) CARMIOLI, new species
Puate 32, Ficures 19-21
Shell large, helicoid; the early turns reddish; the later covered on
the upper surface by a golden-yellow periostracum. There is a light
zone immediately above the periphery, while the posterior half of the
base is chestnut brown; the rest is of the same color as the upper
surface; interior of aperture bluish white. The nucleus consists of
2 small, well-rounded, smooth turns; the postnuclear whorls are in-
flated, strongly rounded, and marked on the upper surface by very
weak, retractively curved, flattened riblets, which become evanescent
on the last whorl. Suture strongly impressed. Periphery well
rounded. Base openly umbilicated, inflated, well rounded, and
marked by the continuation of the axial ribs, which gain in intensity
below the middle and become stronger on the umbilical wall, where
they form distinct fine ribs, which are not quite as wide as the spaces
that separate them. The last whorl is slightly decurrent near the
peristome. Aperture large, broadly oval with a conspicuous angle
at the posterior angle, which has a groove in the middle; peristome
simple; the outer lip thin. The basal lip bears 2 notches; the inner
lip is somewhat. thickened, while the parietal wall is covered with a
moderately thick callus, which renders the peritreme complete.
The type, U.S.N.M. No. 25034, was collected by Carmiol in Costa
Rica. It has 4.9 whorls and measures: Height, 27.2 mm.; greater
diameter, 40.4 mm.; lesser diameter, 29.4 mm. Height of aperture,
16.5 mm.; diameter, 19 mm.
U.S.N.M. No. 405227 contains a young specimen collected by M.
Valerio at Chitaria, Costa Rica.
This species differs from the other members of this group by being
considerably larger.
234 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) COSTARICENSE (von Martens)
PLATE 32, Figures 16-18
1876. Cyclotus quitensis costaricensis VON MARTENS, Jahrb. deutschen malak.
Ges., vol. 3, p. 254.
Shell helicoid, covered by a thin golden-yellow periostracum. The
base bears a moderately broad chestnut-colored band, which fades
anteriorly into the basal color, which is a little darker than that of
the upper surface; interior of the aperture bluish white. The nucleus
consists of 2 small, well-rounded, smooth turns; the postnuclear
whorls are inflated, strongly rounded; the last one is slightly flat-
tened below the summit and is marked by obsolete hairlike axial
riblets, which become much enfeebled on the last half of the last turn.
The last fifth of the last turn is slightly decurrent. Suture strongly
impressed. Periphery feebly angulated. Base narrowly umbilicated,
marked by the continuation of the axial ribs, which become fused and
slightly stronger on the umbilical wall. Aperture very oblique, sub-
circular with a decided auriculation at the posterior angle, which
bears a groove in its middle; peristome simple; the basal lip with
2 deep notches, one marking the outer limit of the umbilicus; the
inner lip is slightly thickened. The parietal wall is covered by a
moderately thick callus, which renders the peritreme complete.
U.S.N.M. No. 321045 contains two specimens from the Redfield
collection, apparently received from Gabb. They come from Costa
Rica. The largest of these, which we have figured, has 4.7 whorls
and measures: Height, 23 mm.; greater diameter, 31.7 mm.; lesser
diameter, 24.4mm. Height of aperture, 14.9 mm.; diameter, 15.3 mm.
U.S.N.M. No. 524052 contains another specimen collected by M.
Valerio.
The much smaller size of this species readily differentiates it from
A. (A.) carmioli.
APEROSTOMA (APEROSTOMA) EXIGUUM, new species
PLATE 82, FicurEs 10-12
Shell small, depressed-helicoid; the upper surface covered with a
golden yellow peristracum. There is a broad light zone immediately
above the periphery and a still broader dark zone immediately
below this; the rest of the base is a little paler than the upper sur-
face. Interior of aperture bluish white. The nucleus consists of 2
small, well-rounded, smooth turns. The postnuclear whorls are some-
what inflated, well rounded and marked by slightly wavy, well-de-
veloped axial riblets, which are about as wide as the spaces that
separate them. There is an impressed groove a little below the summit
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA ZO
on the last whorl, which gives the upper edge of this whorl a some-
what flattened aspect. The last whorl also is slightly decurrent near
the peristome. Suture strongly impressed. Periphery weakly angu-
lated. Base openly umbilicated, inflated, well rounded, and marked
by the continuation of the axial riblets, which fuse into heavier riblets
on the umbilical wall. Aperture subcircular, with a conspicuous
auricle at the posterior angle, which has a groove in its center; peri-
stome simple; outer lip thin; the basal lip with 2 deep incisions, the
first of which marks the outer limit of the umbilicus, and the other is
about halfway between this and the periphery; the inner lip is mod-
erately thickened, while the parietal wall is covered by a moderately
strong callus, which renders the peritreme complete.
The type, U.S.N.M. No. 190281, was collected by H. Pittier in the
Zhorquin Valley, Talamanca, Costa Rica. It has 4.5 whorls and
measures: Height, 15.4 mm.; greater diameter, 23.7 mm.; lesser diam-
eter, 16.4mm. Height of aperture, 9.6 mm.; diameter, 11.7 mm.
This species can readily be distinguished from the other members
of the group having two notches on the basal area by its very much
smaller size.
APEROSTOMA (APEROSTOMA) BISINUATUM (von Martens)
PLATE 33, FicureEs 5, 6
1864. Cyclotus bisinuatus voN MARTENS, Malak. Bliitter, vol. 11, p. 118, pl. 3,
figs. 1, 2.
1890. eds (Aperostoma) bisinuatus voN MARTENS, Biologia Centrali-Ameri-
cana, p. 3.
1897. Neocyclotus (Neocylotus) bisinuatus Kopett and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
We have not seen this species and give a translation of von Martens’
description:
Shell umbilicated, turbinate, solid, marked by closely spaced, un-
dulating, confluent riblets, yellowish brown unicolor. Spire conic,
somewhat acute. Whorls 5, quite convex, the last one rounded, flat-
tened at the suture. Umbilicus quite narrow funnel-shaped, delimited
by an obsolete angle. Aperture diagonal, ovate, circular, angulated
above, bluish white within; peristome straight, obtuse, emarginate
above and below. Columellar margin arcuate; major diameter, 38.5
mm.; lesser diameter, 29 mm. Height, 31mm. Aperture, 20 mm.
Discovered in a high plateau of Costa Rica by the late Hoffmann
who sent it to the Berlin Museum.
This species is distinguished from the other members of the group
by the two notches, one at the outer limit of the umbilicus, and the
other diametrically across from it in the outer lip, a little posterior
to the middle.
236 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) IRREGULARE (Pfeiffer)
PLATE 33, FIGURE 4
1855. Cyclostoma (Cyclotus) irregulare PretrrEr, Proc. Zool. Soc. London, vol. 23,
a ee
1858. patie irregularis PrrirFER, Monographia pneumonopomorum viventium,
vol. 2, p. 15.
1864. Cyclotus irregularis Remve, Conchologia iconica, vol. 14, pl. 4, fig. 18.
1890. Cyclotus (Aperostoma) irregularis VON MARTENS, Biologia Centrali-Amer-
icana, Suppl., pp. 3, 596.
1897. Neocyclotus (Neocyclotus) irregularis KopeLtt and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
1923. Poteria (Neocyclotus) irregulare H. B. BAxKer, Occ. pap., Mus. Zool. Univ.
Michigan, No. 187, p. 41.
As we have not seen this species we give a translation of Pfeiffer’s
description :
Shell umbilicated, depressed-turbinate, solid, closely striated, yel-
lowish. The spire is shortly turbinate, acute. Suture profound.
Whorls 5, convex, the last swollen above and broadly canaliculate at
the suture; below marked by a broad median band. The broadly fun-
nel-shaped umbilicus is limited on the outside by a strong angulation.
Aperture oblique, triangularly oval, angulated above; peristome
continuous, slightly adnate to the preceding turn, straight. The right
outer lip dilated; columella thickened moderately arcuate, channeled
basally. Major diameter, 37 mm.; lesser, 30 mm.; altitude, 19 mm.
Habitat : Central Costa Rica.
Reeve’s figure shows a decided notch at the junction of the inner
and basal lip which is responsible for the angulation marking the outer
edge of the umbilicus.
The species appears most nearly related to A. (A.) pittiert (von
Martens) but differs from it in being much larger.
APEROSTOMA (APEROSTOMA) PITTIERI (von Martens)
1900. Cyclotus (Aperostoma) irregularis pittiert VON Martens, Biologia Centrali-
Americana, Suppl., p. 597.
1923. Poteria (Neocyclotus) irregularis pittieri, H. B. BAkrr, Occ. Pap. Mus.
Zool. Univ. Michigan, No. 137, p. 41.
We have not seen this species. Von Martens distinguishes it from
A. (A) trregulare Pfeiffer with the followmg diagnosis, which we
translate :
Distinctly flatter than the typical form [¢rregulare], less vaulted,
29 mm. only in diameter and 16.5 mm. in height. Aperture 12 mm.
Habitat: Salinas Bay, northwestern Costa Rica, collected by H.
Pittier.
The much smaller size distinguishes this from A. (A.) érregulare
(Pfeiffer).
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 237
THE GROUP OF APEROSTOMA (APEROSTOMA) GIGANTEUM
Posterior angle of aperture forming a slight angle, bearing a weakly
impressed median groove.
APEROSTOMA (APEROSTOMA) CONFUSUM (Sykes)
PLATE 33, Ficures 1-3
1846. Cyclostoma gigantewm Preirrer, Martini-Chemnitz Conchylien Cabinet,
vol. 1, sect. 19, p. 11, pl. 1, figs. 11-14 (not OC. giganteum Reeve, Con-
chologia systematica, vol. 2, p. 99, pl. 184, fig. 17, 1842).
1901. Aperostoma confusum SyKeEs, Journ. Malac., vol. 8, p. 106, pl. 10, fig. 2.
1923. Poteria (Neocyclotus) confusum H. B. Baxsr, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 187, p. 42.
Shell very large, helicoid. Our specimen of this species, unfor-
tunately, is sadly denuded. However, it shows that there is a broad
dark chestnut-colored band below the periphery and apparently a
narrow light zone immediately above this; interior of aperture pale
buff. The nucleus consists of about 2.4 small, well-rounded, smooth
turns. The postnuclear whorls are marked by decidedly retractively
slanting, rather irregular, closely approximated axial riblets, which
on the last half of the last turn become confluent incremental ele-
ments. The summit of the whorls is appressed, ‘and there is a slight
concavity on the last turn below the summit. Suture well impressed.
Periphery strongly rounded. Base narrowly umbilicated and marked
by the continuation of the axial sculpture, which does not increase
in differentiation on the umbilical wall. Aperture circular, oblique,
with a small angulation bearing a central groove at the posterior
angle; peristome simple; outer lip thin; the inner thickened and
slightly reflected. The parietal wall is covered by a thick callus,
which renders the peritreme complete. This callus is not in a straight
line with the columella, but forms a little oval sinus.
The specimen that we believe belongs here, U.S.N.M. No. 341766,
was collected by A. E. Heighway at Acandi, a small settlement in the
Gulf of Atrato, on the boundary between Colombia and Panama.
It has 5.3 whorls and measures: Height, 34.2 mm.; greater diameter,
51.5 mm.; lesser diameter, 38.2 mm. Height of aperture, 20.5 mm.;
diameter, 24.8 mm.
This species is differentiated from the rest of the gzgantewm group
by having the spire much more elevated.
APEROSTOMA (APEROSTOMA) GIGANTEUM (Reeve)
PLATE 33, Figures 7-9
1842. Cyclostoma giganteum Reeve, Conchologia systematica, vol. 2, p. 99, pl.
184, fig. 17.
1843. Cyclostoma giganteum SowerBy, Thesaurus conchyliorum, p. 92, pl. 23,
figs. 8, 9.
428349—42——_16
238 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
1843. Cyclostoma giganteum Sowersy, Proc. Zool. Soc. London, 1843, p. 30.
1847. Aperostoma giganteum Prrir¥rer, Zeitsch. Malak., p. 104.
1850. Cyclotus giganteus Gray, Nomenclature of molluscous animals and shells
in the collection of the British Museum, pt. 1, p. 6.
1897. Neocyclotus (Neocyclotus) giganteus Kosrtr and MOLLENDoRF¥, Nachrbl.
deutschen malak. Ges., vol. 29, p. 137.
1901. Aperostoma giganteuwm SyxeEs, Journ. Malac., vol. 8, p. 105, pl. 10, fig. 1.
1923. Poteria (Neocyclotus) gigantea H. B. BAKer, Occ. pap. Mus. Zool. Univ.
Michigan, No. 187, pp. 40, 42.
Shell gigantic, depressed-helicoid. The early whorls pinkish; the
later pale yellow, varying from yellow to pale chestnut, with a light
zone below the summit and a broader yellowish zone immediately
above the periphery. The posterior half of the base is chestnut-
brown, while the anterior half and umbilical wall are yellowish horn
colored; interior of aperture bluish. The nucleus consists of 2 very
small, well-rounded, smooth turns. The postnuclear whorls are
moderately rounded and appressed at the summit with an impressed
groove immediately below the summit. The postnuclear whorls are
marked by regular, retractively curved, closely spaced axial riblets,
which are a little wider than the spaces that separate them. On the
last half of the last whorl they become obsolete and are replaced by
mere lines of growth. Suture strongly impressed. Periphery well
rounded. Base with a very large open umbilicus, inflated, and
marked by the continuation of the axial ribs, which become fused on
the anterior half of the base and quite strong and well differentiated
into strong ribs on the umbilical wall. Aperture oblique, subcircular,
drawn into an obtuse angle at the posterior angle, which has a median
groove; peristome simple; that of the outer lip sharp and slightly
reflected, while the inner lip is thickened. The parietal wall is cov-
ered by a thick callus, which renders the peritreme complete. In the
peristome there is a shallow sinus at the posterior termination of the
columella. Operculum typically aperostomid, having 9 whorls.
The specimen described and figured is one of two, U.S.N.M. No.
951101, collected by H. Pittier on the Cerro de Garagara, Panama, at
20 to 900 meters. It has 3.7 whorls remaining and measures: Height,
32.6 mm.; greater diameter, 56.7 mm.; lesser diameter, 38.3 mm.
Height of aperture, 23.5 mm.; diameter, 27 mm.
The depressed-helicoid form and gigantic dimensions readily dis-
tinguish this from the other members of the gigantewm group.
Reeve, Joc. cit., p. 97, states: “Mr. Sowerby has kindly permitted
us to refer to his figure of Cyclostomata in part 2 of his Species Con-
chyliorum, which we are happy to announce is now nearly ready for
publication.” This work, it would appear, was never published.
Cyclostoma giganteum Sowerby must be considered a manuscript
name, and the name Cyclostoma giganteum must date from Reeve,
1842.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 239
APEROSTOMA (APEROSTOMA) MANABENSE, new species
Puate 34, Fiagures 10-12
Shell of medium size, very depressed-helicoid. The early denuded
whorls pale red, the last covered with a thin, reddish olivaceous, horn-
colored periostracum. There is a narrow light zone immediately
above the periphery, while the posterior half of the base is chestnut-
brown, and the anterior half and umbilical wall are paler than the
upper surface; interior of aperture bluish white with a livid tinge.
The nucleus consists of 2.4 small, well rounded, smooth turns. The
postnuclear whorls are slightly rounded. There is a shallow groove
immediately below the summit. The last whorl is obliquely deflected
for about four-fifths of a turn. The postnuclear whorls are marked
by weak, retractively slanting, very slender, closely spaced axial rib-
lets, which become evanescent on the last half of the last turn, where
they are represented by mere lines of growth. Suture well impressed.
Periphery well rounded. Base broadly, openly umbilicated, inflated,
strongly rounded, marked by the continuation of the axial riblets
which become fused into stronger ribs on the anterior half of the
base and umbilical wall. Aperture decidedly oblique, tear-shaped,
drawn out into a decided angle at the posterior angle; peristome
thin on the outer and basal lip; a little thicker on the inner lip. The
parietal wall is covered by a thick callus, which renders the peri-
treme complete.
The type, U.S.N.M. No. 524066, was collected by O. Haught be-
tween Quevedo and Calcata, Manabi, Ecuador. It has 4.8 whorls
and measures: Height, 23.2 mm.; greater diameter, 48 mm.; lesser
diameter, 29.4mm. Height of aperture, 16.5 mm.; diameter, 20.3 mm.
The wide umbilication suggests A. (A.) gigantewm (Reeve), from
which it is distinguished by its small size.
APEROSTOMA (APEROSTOMA) FISCHERI (Hidalgo)
PLATH 34, FiaurEs 1-3
1867. Cyclotus fischeri Hipateo, Journ. Conchyl., vol. 15, p. 305, pl. 8, fig. 3.
1901. Aperostoma fischeri Syxss, Journ. Malac., vol. 8, p. 105, pl. 10, fig. 3.
1912. Neocyclotus giganteus Kosetr (pars), Martini-Chemnitz Conchylien Cabi-
net, vol. 1, sect. 19, pt. 3, p. 891, pl. 133, fig. 1.
1923. Poteria (Neocyclotus) fischeri H. B. Baker, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 137, p. 42.
Shell moderately large, helicoid. The early whorls rose-red; the
last covered with an olivaceous pale chestnut-colored periostracum.
There is a narrow light zone at the periphery, while the posterior half
of the base is conspicuously bright chestnut-brown; the anterior half
and the umbilical wall are much paler. The interior of the aperture
is bluish white. The nucleus consists of 2 very small, well-rounded
240 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
smooth turns; the postnuclear whorls are moderately rounded,
slightly flattened near the summit, the last one for about one-tenth of
a turn forming a cord at the appressed summit. The postnuclear
whorls are marked by fine, retractively curved axial riblets which are
about as wide as the spaces that separate them but which become ir-
regular and less developed on the last turn, particularly toward the
aperture. Suture well impressed. Periphery feebly angulated.
Base openly umbilicated, inflated, well rounded, and marked by the
continuation of the axial riblets, which extend over the umbilical
wall, where they become somewhat strengthened. Aperture subcir-
cular, decidedly protracted at the posterior angle into an auricle,
which bears a groove in its middle; peristome simple; the outer lip
sharp and somewhat reflected; the inner somewhat thickened. The
parietal wall is covered by a thick callus, which renders the peritreme
complete. There is only a feeble inclination toward an inbending
at the junction of the columella with the parietal wall. Operculum
typically aperostomid, showing 9 whorls.
The specimen described and figured, U.S.N.M. No. 307424, comes
from New Grenada. It has 4.9 whorls and measures: Height, 27.5
mm.; greater diameter, 47 mm.; lesser diameter, 32.4 mm. Height of
aperture, 17 mm.; diameter, 22.8 mm.
U.S.N.M. No. 307421 contains one specimen from Quito, Ecuador.
This species can readily be distinguished from A. (A.) utriaense
by its deflected last whorl.
APEROSTOMA (APEROSTOMA) UTRIAENSE, new species
PLATE 34, FIGURES 4-6
Shell of medium size, depressed-helicoid. The upper surface is
covered with a pale chestnut-brown periostracum, but there is a rather
broad paler area immediately below the summit. There is also a light
zone immediately above the periphery, while a broad chestnut-colored
band extends over the posterior two-fifths of the base, the rest being
pale olivaceous horn colored; interior of aperture bluish white. The
nuclear whorls in the type are lost. The postnuclear whorls are well
rounded and on the last turn there is a groove immediately below the
periphery. The postnuclear whorls are marked by very fine, closely
spaced, retractively slanting axial riblets, which become much weak-
ened on the last whorl. Suture strongly impressed. Periphery well
rounded. Base moderately broadly openly umbilicated, marked by the
continuation of the axial sculpture, but much more roughly so. The
axial ribs of the base become fused into stronger elements on the um-
bilical wall. Aperture subcircular, with a moderately strong angula-
tion at the posterior angle, which bears a median groove; outer lip
thin and somewhat reflected; the inner lip slightly thickened. The
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 241
parietal wall is covered by a thick callus, which renders the peritreme
complete. There is only a weak emargination on the columellar callus.
The type, U.S.N.M. No. 524068, comes from Puerto Utria, Colombia.
It has 3.2 whorls remaining and measures: Height, 27.4 mm.; greater
diameter, 46.7 mm.; lesser diameter, 30.9 mm. Height of aperture,
19.4 mm. ; diameter, 23.3 mm.
This species most nearly resembles A. (A.) brujense, from which it
can readily be differentiated by its less strongly ribbed striate
sculpture.
APEROSTOMA (APEROSTOMA) BRUJENSE, new species
PuLaTe 34, Ficures 13-15
Shell large, depressed-helicoid. The early denuded whorls vary
from rose colored to flesh colored. The last whorl is covered with a
brownish, suffused, horn-colored periostracum. There is a narrow
light zone immediately above the periphery, while the posterior half
of the base is very dark chestnut-brown, and the anterior half and
umbilical wall are a little lighter than the upper surface. The nucleus
consists of 2 small, well-rounded, smooth turns. The postnuclear
whorls are somewhat inflated, well rounded, with an impressed area
immediately below the summit on the last whorl, while the summit
itself appears as an appressed thread. The surface of the postnuclear
whorls is marked by rather coarse, retractively slanting axial rib-
lets, which vary considerably in strength and spacing. Suture well
impressed. Periphery weakly angulated. Base moderately broadly
umbilicated and marked by the continuation of the axial riblets, which
become fused into stronger ribs on the umbilical wall. Aperture sub-
circular with a moderately strong angle at the posterior angle which
bears a median groove; peristome simple; the outer lip expanded and
somewhat reflected; the inner slightly thickened. The parietal wall
is covered by a very thick callus, which renders the peritreme complete.
There is very weak emargination in this callus at the insertion of the
columella.
The type, U.S.N.M. No. 251418, was collected by Maj. E. A. Gold-
man on the Cerro Bruja, Panama, at an elevation of 500 feet. It has
4.7 whorls and measures: Height, 28.6 mm.; greater diameter, 49.4
mm.; lesser diameter, 33mm. Height of aperture, 19.2 mm.; diameter,
24.4 mm.
The following additional specimens are in the National Museum
collections: No. 524069, a topotype from the same source as the type;
No. 228908, a specimen collected by H. Pittier in the mountains around
Gaspasalana, High Mamoni, Panama; No. 251419, five specimens col-
lected by Major Goldman at Gatun, Canal Zone.
This species can be distinguished readily from A. (A.) wtriaense
by its much stronger rib-striate sculpture.
242 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) PORTOBELLENSE, new species
PLATE 34, FIGURES 7-9.
Shell small for the group, depressed-helicoid. The early whorls
red, the last covered on the upper surface by a thin horn-colored perios-
tracum. ‘There is a light supraperipheral zone and a dark area im-
mediately below the periphery on the base. The rest is horn colored;
interior of aperture pale buff. The nucleus consists of about 2 small,
well-rounded turns. Postnuclear whorls well rounded, with a de-
pressed area a little below the summit. The postnuclear whorls are
marked by very slender, retractively curved, closely crowded axial
ribs, which become coarser and more distantly spaced on the last whorl.
Suture well impressed. Periphery slightly angulated. Base openly
umbilicated and marked by the continuation of the axial ribs, which
do not gain materially in strength on the umbilical wall. Aperture
subcircular, drawn into a moderately produced angle at the posterior
angle, which bears a groove in the middle; peristome simple; the outer
lip expanded and slightly reflected; the inner slightly thickened.
The parietal wall is covered by a thick callus, which renders the
peritreme complete. There is only an emargination in the parietal
callus.
The type, U.S.N.M. No. 251484, was collected by E. A. Schwarz
at Porto Bello, Panama. It has 4.7 whorls and measures: Height,
21.5 mm.; greater diameter, 35 mm.; lesser diameter, 23.8 mm. Height
of aperture, 15.2 mm.; diameter, 17.8 mm.
U.S.N.M. No. 251427 was collected by August Busck at Porto Bello,
Panama.
This species can readily be distinguished from all the other members
of the group by its small size.
APEROSTOMA (APEROSTOMA) FULTONI, new species
Piate 35, Figures 14
Shell depressed-helicoid, chestnut brown, with a lighter zone at
the summit and a white zone immediately above the periphery. There
is a broad dark brown band covering two-fifths of the base immediately
below the periphery and a slender narrow band a little posterior to
the middle of the base. The rest of the base is yellow horn colored;
interior of aperture bluish white, showing the banding within. The
nucleus consists of 2 small, well-rounded, smooth turns. The post-
nuclear whorls are strongly rounded with a depressed area imme-
diately below the summit. The postnuclear whorls are marked by
rather regular strong retractively curved ribs, which are about as
wide as the spaces that separate them. Suture strongly impressed.
Periphery obsoletely angulated. Base moderately broadly umbili-
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 243
cated, inflated, strongly rounded and marked by the continuation of
the axial ribs which grow much stronger on the umbilical wall. Aper-
ture very large, circular, with an angulation at the posterior angle;
peristome simple; the outer lip thin, slightly effuse at the junction
of the basal and outer lip; a little thickened on the inner lip. The
parietal wall is covered by a thick callus, which renders the peritreme
complete.
The type, U.S.N.M. No. 307425, was received from Sowerby and
Fulton with the label “Brazil.” It has 4.8 whorls and measures:
Height, 17.8 mm.; greater diameter, 29.9 mm.; lesser diameter, 21.5
mm. Height of aperture, 12 mm.; diameter, 13.4 mm.
This species agrees with A. (A.) amazonense in having the base
strongly axially ribbed but differs from it in having a much larger
aperture.
APEROSTOMA (APEROSTOMA) AMAZONENSE, new species
PLATE 35, FIGURES 4-6
Shell decidedly depressed-helicoid, almost planorbid. The entire
' upper surface is covered by a pale chestnut-brown periostracum.
There is a faint lighter zone immediately above the periphery. The
base is bicolor. The posterior half is a little darker than the upper
surface of the shell, while the anterior half and umbilical wall are
much lighter; interior of aperture bluish white. The nucleus con-
sists of 2 small, well-rounded, smooth turns; the postnuclear whorls
are moderately rounded, the last one defiected for a quarter of a turn.
The postnuclear whorls are marked by retractively curved axial rib-
lets, which are not quite as wide as the spaces that separate them.
These riblets become decidedly enfeebled on the last portion of the
last turn. Suture strongly impressed. Periphery weakly angulated.
Base broadly umbilicated, inflated, strongly rounded and marked by
strong axial ribs, which are more pronounced on the umbilical wall
than on the rest. Aperture subcircular, slightly angulated at the pos-
terior angle; peristome simple; the outer and inner lips both some-
what thickened and slightly reflected. The parietal wall is covered by
a thick callus which renders the peritreme complete. The operculum
is typically aperostomid and shows about 10 whorls.
The type, U.S.N.M. No. 356102, was collected by E. H. Short on
the Amazon River in Brazil. It has 4.5 whorls and measures:
Height, 13 mm.; greater diameter, 24 mm.; lesser diameter, 18.1 mm.
Height of aperture, 9 mm.; diameter, 9.9 mm.
There are two additional lots, both bearing merely the label “Bra-
zil”; U.S.N.M. No. 307425 contains one specimen and U.S.N.M.
No. 316236, three specimens.
244 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
This species agrees with A. (A.) fultoni in having the base rather
strongly axially ribbed, but it differs from it in having a much smaller
aperture.
APEROSTOMA (APEROSTOMA) LAXATUM (Sowerby)
PLATE 35, FIGURE 13
1850. Cyclostoma laxatum Sowersy, Thesaurus Conchyliorum, vol. 1, Suppl.
p. 159*, pl. 314, fig. 302.
1852. Cyclotus laxatus PFEIFFER, Monographia pneumonopomorum viventium,
vol. 1, p. 18.
1897. Neocyclotus (Neocyclotus) laratus Kospetr and MO6OLLEeNnporFr, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
1923. Poteria (Neocyclotus) laratum H. B. BaKer, Occ. Pap., Mus. Zool. Univ.
Michigan, No. 137, p. 41.
Although Cyclostoma laxatum Sowerby has been quoted by many
malacologists, we regret to say that we have none in the National
Museum collection that agree with Sowerby’s description, figure, and
habitat; we must therefore quote Sowerby’s description and copy his
figure. His figure measures: Height, 18.5 mm.; greater diameter, 36.4
mm. Height of aperture, 13.1 mm.; diameter, 16.9 mm.
“Shell of a depressed orbicular form, whitish, covered with a thick
orange-brown epidermis; spire depressed, with a rufous apex; volu-
tions four, rounded, covered not only with distinct lines of growth, but
also with irregular rugulosities, with two narrow zones, the posterior
white, the other fuscous, in front of which the epidermis is more
darkly coloured; aperture not quite circular, but very slightly ellip-
tical, and with a slight angle posteriorly close to the last volution;
peritreme thin, anteriorly very slightly reflected; umbilicus very
large, showing internally the four volutions.
“The shell is somewhat diaphanous, so that the white and dark-
brown zones are distinctly seen within the aperture.
“Columbia. Mus. Cuming.”
THE GROUP OF APEROSTOMA (APEROSTOMA) BLANCHETIANUM
Shell decidedly depressed.
APEROSTOMA (APEROSTOMA) BLANCHETIANUM (Moricand)
PLATE 35, Figures 7-9
1826. Cyclostoma blanehetianum MoricAnpd, Mem. Soe. Genéve, vol. 7, p. 442,
pl. 2, figs. 21-23.
1847. Aperostoma blanchetianum TroscHEL, Zeitsch. Malak., p. 44.
1847. Aperostoma blanchetianum Prertrrer, Zeitsch. Malak., p. 104.
1850. Cyclotus inca (pars) Gray, Nomenclature of molluscous animals and
shells in the collection of the British Museum, pt. 1, p. 6.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 245
1897. Neocyclotus (Neocyclotus) inca (pars) Kosett and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
1923. Poteria (Neocyclotus) blanchetianwm H. B. BaxKeEr, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 187, p. 44.
Shell depressed-helicoid. The early whorls, denuded of the
periostracum, reddish; the later reddish horn colored, the last oliva-
ceous on the upper surface. There is a narrow white zone at the
periphery and a broad dark olivaceous band covering the posterior
two-fifths of the base, the rest being horn colored; interior of aper-
ture bluish white. The nucleus consists of 2 moderately large, well-
rounded whorls. The postnuclear whorls are strongly rounded and
marked by retractively curved, strong, small axial ribs, which are
smaller than the spaces that separate them. On the last half of the
last turn these riblets become much enfeebled and replaced by mere
rough incremental lines. The suture is narrowly deeply channeled.
Periphery very weakly angulated. Base broadly openly umbilicated,
inflated, well rounded and marked by the continuation of the axial
ribs, which grow considerably stronger on the umbilical wall. On
the last half of the last whorl they become fused and more or less
scalelike. Aperture circular, with a slight angulation at the pos-
terior angle; peristome simple; the outer lip thin, slightly reflected ;
the inner slightly thickened and also slightly reflected. The parietal
wall is covered by a rather thick callus, which renders the peritreme
complete. Operculum typically aperostomid, with about 10 whorls.
The specimen described and figured is one of two, U.S.N.M. No.
98110, collected at Bahia, Brazil. It has 4.5 whorls and measures:
Height, 15.2 mm.; greater diameter, 27.4 mm.; lesser diameter, 19.7
mm. Height of aperture, 9.9 mm.; diameter, 11.9 mm.
The following additional specimens are in the National Museum
collections:
No. 104450, five specimens from Bahia received from Moricand;
No. 307417, one specimen labeled “Brazil”; No. 381893, one specimen
from the Amazon Valley. This species closely resembles A. (A.)
peruense but is readily differentiated from it by its deeply incised
narrow suture.
APEROSTOMA (APEROSTOMA) PERUENSE, new species
PLATE 35, FicurEs 10-12
Shell very depressed-helicoid, almost planorbid; the upper surface
dark horn colored. There is a light white band immediately above
the periphery and a broad brilliant chestnut-brown zone immediately
below the periphery. There is also a second, very narrow brown
band separated from the broad band by a space about twice as wide
as the narrow brown band. The rest of the base is horn colored,
246 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The nucleus consists of 2 small, well-rounded, smooth turns; the
postnuclear whorls are strongly rounded and marked by retractively
curved, very regular axial riblets, which are a little narrower than
the spaces that separate them. On the last half of the last turn the
riblets become less pronounced and less regular. There are also some
slight spiral markings on the upper surface. Suture strongly im-
pressed. Periphery strongly rounded. Base broadly, openly um-
bilicated, inflated, and marked by the continuation of the axial ribs,
which become a little intensified on the umbilical wall. Aperture
broadly oval, with a slight auricle at the posterior angle; peristome
simple; the outer lip of the peristome thin; the inner slightly thick-
ened and reflected. The callus on the parietal wall is thick and
renders the peritreme complete.
The type, U.S.N.M. No. 103999, comes from the Rich collection and
bears the label “Peru.” It has 4.2 whorls and measures: Height,
14.3 mm.; greater diameter, 26.8 mm.; lesser diameter, 18.2 mm.
Height of aperture, 10.5 mm.; diameter, 12.3 mm.
This species can readily be differentiated from A. (A.) blanche-
tianum (Moricand) by its lacking the narrowly channeled suture.
APEROSTOMA (APEROSTOMA) LEAI, new species
PLATE 35, Fiaures 17-19
Shell very depressed-helicoid, almost planorbid. The upper sur-
face is orange horn colored, with a little lighter zone at the summit
and a broader white zone immediately above the periphery. The
base bears a broad, bright, chocolate-brown band, which covers about
one-third of the base anterior to the periphery. The rest is horn
colored, with a narrow hydrophanous band on the middle of the base.
Interior of aperture bluish white. The nucleus consists of 2 moder-
ately large, well-rounded smooth turns. The postnuclear whorls are
well rounded, the last half of the last whorl] being slightly flattened
below the suture. The postnuclear whorls are marked by retractively
curved, very regular, rather closely spaced axial riblets, which are
separated by spaces about as wide as the axial riblets. On the last
part of the last turn these riblets become much enfeebled. Suture
deeply impressed. Periphery feebly angulated. On the dark band
below the periphery on the base the axial ribs are almost obsolete;
they grow a little stronger on the rest of the base and are strongest on
the umbilical wall. The umbilicus is broadly openly funnel-shaped.
Aperture broadly oval, with a rather strong angulation at the pos-
terior angle; peristome simple; outer lip thin and slightly outward
reflected; the inner a little thicker. The parietal wall is covered by
a rather thick callus which renders the peritreme complete. Oper-
culum typically aperostomid, showing 6 whorls remaining.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 247
The type, U.S.N.M. No. 104451, comes from Balsas, Peru. It was
in the Lea collection. It has 4.8 whorls and measures: Height, 14.2
mm.; greater diameter, 27.5 mm.; lesser diameter, 19.2 mm. Height
of aperture, 10.7 mm.; diameter, 12.3 mm.
The obsolete ribbing immediately below the periphery distin-
guishes this species from A. (A.) peruense and A. (A). blancheti-
anum (Moricand).
APEROSTOMA (APEROSTOMA) VENEZUELENSE, new species
PLATE 35, FIGURES 20-22
Shell very depressed-helicoid, thin, covered with an olivaceous
periostracum, which is streaked and blotched with darker olive.
There is a faint light zone immediately above the periphery and a
broad weak brown band immediately below it. A second slender
spiral line lies a little posterior to the middle of the base. The rest
of the base is a little paler than the upper surface; interior of
aperture bluish white. The nucleus consists of 2 small, well-rounded,
smooth turns. The postnuclear whorls are strongly rounded, the
last one slightly deflected at its extremity. They are marked by
obsolete axial riblets, which become quite irregular on the last whorl,
where also some slight malleations are present. The last whorl is
slightly flattened a little below the summit and the suture is deeply
narrowly channeled. Periphery obsoletely angulated. Base openly
broadly umbilicated, inflated, and marked by the same type of sculp-
ture as that which characterizes the upper surface of the last whorl,
except that on the umbilical wall the riblets become a little more
distant. Aperture broadly oval, with a slight angulation at the
posterior angle; peristome simple; the outer Jip thin; the inner only
slightly thickened. The parietal wall is covered by a moderately
thick callus, which renders the peritreme complete.
The type, U.S.N.M. No. 307429, was collected by Cuming in Vene-
zuela. It has 4.2 whorls and measures: Height, 14.7 mm.; greater
diameter, 29.7 mm.; lesser diameter, 21 mm. Height of aperture,
11.8 mm.; diameter, 12.8 mm.
This species agrees with A. (A.) depresswm (Da Costa) in having
the axial ribs obsolete; it differs from that, however, in having the
whorls strongly downward flexed near the suture and more rounded.
APEROSTOMA (APEROSTOMA) DEPRESSUM (Da Costa)
PLATE 35, F1icuRES 14-16
1906. Neocyclotus depressus Da Costa, Proc. Malac. Soc. London, vol. 7, p. 9,
pl. 1, figs. 14-16.
1923. Poteria (Neocyclotus) depressus H. B. BAKER, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 187, p. 42.
948 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Shell very depressed-helicoid, almost planorbid, covered by an
olivaceous periostracum, with a moderately broad light zone immedi-
ately above the periphery and the posterior half of the base oliva-
ceous-chestnut colored; the anterior half and the umbilicus are horn
colored; interior of aperture bluish white. The nucleus consists of
2 small, well-rounded, smooth turns. The postnuclear whorls are
slightly rounded with a depression a little below the summit on the
last half of the last turn. The postnuclear whorls are marked by
obsolete, retractively curved axial riblets and faint traces of mallea-
tions in the depressed portion below the summit. Suture very
strongly impressed. Periphery well rounded. Base broadly, openly
umbilicated, inflated, well rounded, and marked by rather regular
axial riblets, which become intensified on the umbilical wall. Aper-
ture circular with an angulation at the posterior angle; peristome
simple; the outer lip thin, the inner thin and slightly reflected. The
parietal wall is covered with a thin callus rendering the peritreme
complete. Operculum typically aperostomid, showing 8 whorls.
U.S.N.M. No. 250742 contains two specimens from Chanchamayo,
Peru. The specimen described and figured has 4.5 whorls and meas-
ures: Height, 18.8 mm.; greater diameter, 26.4 mm.; lesser diameter,
19.5 mm. Height of aperture, 11 mm.; diameter, 12 mm.
This species resembles A. (A.) venezuelense in the obsolete ribbing,
but differs from it in having the whorls much flatter on the upper
surface and not inbent near the suture as in that form.
APEROSTOMA (APEROSTOMA) ECUADORENSE, new species
PLATE 36, Figures 4-6
Shell helicoid, the early whorls reddish horn colored, the last horn
colored. There is a lighter, very pronounced zone immediately above
the periphery, and a darker zone, which gradually fades basally,
occupying the posterior two-fifths of the whorl. The rest of the base
is of about the same shade of horn color as the last whorl; interior
of aperture bluish white, showing the external banding within. The
nucleus consists of 2 moderately large, well-rounded, smooth turns.
The postnuclear whorls are well rounded; the last one is gradually
deflected for half of a turn. The postnuclear whorls are marked by
quite regular, retractively curved, slightly flattened axial riblets,
which are a little broader than the spaces that separate them. On
the last part of the last turn these become somewhat weakened.
Suture well impressed. Periphery rounded. Base narrowly um-
bilicated, inflated, strongly rounded, and marked by strong axial
riblets, which are considerably stronger than those on the upper sur-
face. Aperture oblique, subcircular, with a slight auricle at the
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 249
posterior angle; peristome simple; outer lip thin, the inner thickened
and slightly reflected. A strong callus covers the parietal wall and
renders the peritreme complete. The operculum is typically aper-
ostomid and shows 8 whorls.
The type, U.S.N.M. No. 316105, was collected by Cuming at Quito,
Ecuador. It has 4.6 whorls and measures: Height, 23.3 mm.; greater
diameter, 33.0 mm.; lesser diameter, 25.5 mm. Height of aperture,
14.5 mm.; diameter, 15.7 mm.
U.S.N.M. No. 307407 contains three topotypes, and U.S.N.M. No.
316110 contains another topotype.
This species is differentiated from the others of the group that
have the axial ribs of the umbilical wall stronger than on the upper
surface by its narrow umbilicus.
APEROSTOMA (APEROSTOMA) SUBCINGULATUM (Kobelt)
PLATE 36, Ficures 1-3
1912. Neocyclotus (giganteus?) subcingulatus Kopett, Martini-Chemnitz Con-
chylien Cabinet, vol. 1, sect. 19, pt. 3, p. 892, pl. 133, figs. 4-6.
1923. Poteria (Neocyclotus) subcingulatum H. B. BaxKer, Oce. Pap. Mus. Zool.
Univ. Michigan, No. 137, p. 42.
Shell low, helicoid, covered with a reddish periostracum, which
pales to chestnut-brown on the last turn. There is a very narrow
light zone immediately above the periphery. The base is bicolor; the
posterior portion covering about two-fifths of the base is dark chest-
nut-brown at the periphery and gradually pales anteriorly; the an-
terior portion is bright horn color. The nucleus consists of 2 small,
well-rounded, smooth turns. The postnuclear whorls are well rounded,
the last one having a slight depression immediately below the sum-
mit. All but the last postnuclear whorls are marked by slender,
retractively curved axial riblets, which are not so wide as the spaces
that separate them. On the last whorl, however, these riblets grow
consecutively weaker and become quite evanescent toward the end.
Suture very strongly impressed. Periphery well rounded. Base
broadly umbilicated, inflated, marked by the continuation of the axial
riblets, which are weak at the periphery but grow stronger as they
cross the base, being very strong on the umbilical wall. Aperture
broadly oval, with a slight auricle at the posterior angle; peristome
simple; the outer lip thin; the inner lip slightly thickened and
slightly reflected. The parietal wall is covered by a moderately thick
callus, which renders the peritreme complete.
The specimen described and figured, U.S.N.M. No. 98109, comes
from Quito, Ecuador. It has 4.6 whorls and measures: Height, 19.5
mm.; greater diameter, 36.1 mm.; lesser diameter, 24.6 mm. Height
of aperture, 14.7 mm.; diameter, 16.3 mm.
250 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
U.S.N.M. No. 366106 contains an additional specimen from Quito;
U.S.N.M. No. 126907, one specimen collected by M. B. Kerr is
labeled merely “Ecuador.”
This species, which has the same type locality as A. (A.) ecwador-
ense, is of about the same size as that species but is more depressed,
with a much larger umbilicus.
THE GROUP OF APEROSTOMA (APEROSTOMA) QUITENSE
Medium-sized shells, not decidedly depressed and having a rather
large aperture.
APEROSTOMA (APEROSTOMA) QUITENSE (Pfeiffer)
PLATE 36, Figures 138-15
1852. Oyclostoma (Cyclotus) quitense Prretrrrr, Proc. Zool. Soc. London, vol.
20, p. 61.
1852. Cyclotus quitensis P¥FrIrFrR, Monographia pneumopomorum viventium,
vol. 1, p. 17.
1854. Cyclostoma quitense Prrtrrrr, Martini-Chemnitz Conchylien Cabinet, vol.
1, sect. 19, p. 340, pl. 44, figs. 19-22.
1897. Neocyclotus (Neocyclotus) quitensis KosELT and MOLLENDORFF, Nachrb.
deutschen malak. Ges., vol. 29, p. 138.
1923. Poteria (Neocyclotus) quitensis H. B. BAKeER, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 187, p. 48.
Shell rather large, low, helicoid, covered with a bright chestnut-
brown periostracum. There is a pale, rather broad zone immediately
above the periphery. The base is bicolor, having a broad, very dark
chestnut-brown band immediately below the periphery, which ex-
tends over half of the base. The anterior half of the base, including
the umbilical wall, is bright horn color; interior of aperture decidedly
bluish, showing the external banding within. The nucleus consists
of 2 well-rounded, smooth turns. The postnuclear whorls are well
rounded, the last one is slightly impressed a little below the suture.
They are marked with weak, retractively curved axial riblets. Suture
strongly impressed. Periphery well rounded. The light band above
the periphery produces an optical illusion of angulation. Base mod-
erately broadly umbilicated, inflated, strongly rounded, and marked
by the continuation of the feeble riblets on the posterior half, but from
there on these riblets become much strengthened and are quite strong
on the umbilical wall. Aperture broadly oval, with a rather strong
auricle at the posterior angle; peristome simple; outer lip rather
flaringly expanded and slightly reflected; inner lip somewhat thick-
ened. The parietal wall is covered by a thick callus, which renders
the peritreme complete. The operculum is typically aperostomid,
showing 9 whorls.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 255
The specimen described and figured, U.S.N.M. No. 307446, comes
from Quito, Ecuador. It has 4.6 whorls and measures: Height, 20.8
mm.; greater diameter, 36.9 mm.; lesser diameter, 24.2 mm. Height
of aperture, 14 mm.; diameter, 18.2 mm.
U.S.N.M. No. 307396 contains three topotypes collected by Cum-
ing; U.S.N.M. No. 307431 contains another topotype from the Morelet
collection.
This species is easily differentiated from A. (A.) ecwadorense and
subcingulatum (Kobelt) by its smoother surface and much larger
aperture.
APEROSTOMA (APEROSTOMA) OLIVACEUM, new species
PuLate 36, Ficures 10-12
Shell low, helicoid, the early whorls pinkish, the last olivaceous
with darker axial streaks, which are of irregular width and spacing.
There is a light zone immediately above the periphery and a darker
broad zone covering the posterior two-fifths of the base. The
umbilical wall and the rest of the base are of the same color as the
upper surface of the last turn; interior of the aparture bluish. The
nucleus consists of 2 small, well-rounded, smooth turns. The post-
nuclear whorls are well rounded; however, there is a depressed area
a little below the summit. The postnuclear whorls are marked by
retractively curved axial ribs, which are of about the same width
as the spaces that separate them and which grow consecutively
stronger as the shell increases in size. In addition, there are irregu-
lar, oblique scratches in the depressed area below the summit on
the last turn and feeble indications of spiral markings on the rest
of the last whorl. Suture strongly impressed. The periphery is
well rounded. The light band, however, gives it the optical illusion
of being angulated. Base narrowly umbilicated, inflated, strongly
rounded, and marked by the continuaaion of the axial ribs, which
become slightly stronger on the umbilical wall. Aperture very
large, subcircular; peristome simple; outer lip thin; inner thickened.
The parietal wall is covered by a very heavy callus which is slightly
emarginated, and which renders the peritreme complete.
The type, U.S.N.M. No. 524084, comes from Ecuador. It has 4.6
whorls and measures: Height, 20.8 mm.; greater diameter, 34.2 mm.;
lesser diameter, 23.1 mm. Height of aperture, 13.9 mm.; diameter,
16.6 mm.
U.S.N.M. No. 307423 contains a topotype.
This species in form and sculpture most nearly resembles A. (A.)
castaneum, from which its much paler coloration easily distinguishes it.
252 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) CASTANEUM, new species
PLATE 36, F1guRES 7-9
Shell low, helicoid, bright chestnut-brown on the upper turns, with
a light zone immediately above the periphery and a broad dark
chestnut-brown band immediately below the periphery. The rest
of the base is golden horn colored; interior of aperture bluish white,
the edge buff colored. The nucleus consists of 2 small, well-rounded,
smooth turns. The postnuclear whorls are moderately rounded,
well impressed a little anterior to the suture with a slender cord at
the suture. The last whorl is decurrent for about one-quarter of a
turn. The postnuclear whorls are marked by strong, retractively
curved, very regular axial riblets, which become decidedly weakened
on the last quarter of the last whorl. Suture strongly impressed.
Periphery well rounded. Base moderately broadly, openly umbili-
cated, inflated, strongly rounded, and marked by the continuation
of the axial ribs, which do not become stronger on the umbilical
wall. Aperture very large, very broadly ovate with a slight angu-
lation at the posterior angle; peristome simple; outer lip thin,
decidedly produced on the outer lip and slightly reflected; inner
lip slightly thickened. The parietal wall is covered by a moderately
thick callus, which renders the peritreme complete.
The type, U.S.N.M. No. 524087, comes from Venezuela without
specific locality. It has 4.4 whorls and measures: Height, 18.4
mm.; greater diameter, 32.2 mm.; lesser diameter, 21.4 mm. Height
of aperture, 12.3 mm.; diameter, 15.7 mm.
This species resembles A. (A.) olévacewm but differs from it in
having the axial sculpture much stronger and the last whorl de-
cidedly decurrent and in being of much darker coloration.
APEROSTOMA (APEROSTOMA) PERUVIANUM (Da Costa)
PLATE 36, F1cuREs 16-18
1906. Neocyclotus peruvianus Da Costa, Proc. Malac. Soc. London, vol. 7, p. 98,
pl. 11, figs. 7-9.
1923. Poteria (Neocyclotus) peruvianus H. B. BAxKkrEr, Oce. Pap. Mus. Zool.
Univ. Michigan, No. 187, p. 42.
We have not seen specimens of this species and so give a translation
of Da Costa’s description and copy his figures:
Shell large, openly umbilicated, depressed, grayish brown with a
narrow yellowish band above the periphery. The base bears a broad
dark band. Whorls 4.5, convex, impressed at the suture, obliquely
rib-striated. Aperture obliquely circular, bluish white. Greater
diameter, 38 mm.; lesser diameter, 29 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 253
Type locality : Pozuzo, eastern Peru.
Da Costa further adds: “There is a certain resemblance between
this shell and JW. cingulatus, Sowb., from Colombia, but it is less
openly umbilicated and not so strongly striated. The different geo-
graphical distribution of the two species should be sufficient to dis-
tinguish them. The shelly operculum is thick and consists of 8-9
whorls, which are thickened at the inner edge.”
The very broad umbilicus will distinguish this from the other
members whose last whorl is not finely regularly ribbed.
APEROSTOMA (APEROSTOMA) SALENGOENSE, new species
PLATE 37, FIcuRES 18-20
Shell low, helicoid; the denuded early whorls pinkish, the later
turns soiled white. Where the periostracum remains this is shown
to be golden-yellow on the upper surface. There is a narrow light
zone at the periphery and the base has a broad, dark, colored band
covering two-fifths of its surface. Anterior to this, including the
umbilical wall, the base has a horncolored periostracum; interior of
the aperture bluish white. The nucleus consists of 2 small, well-
rounded, smooth turns, forming a rather pointed apex. The post-
nuclear whorls are well rounded, with a slightly impressed area
a little below the suture on the last turn and a weak cord at the sum-
mit, which is appressed to the preceding turn. The postnuclear
whorls are marked by rather irregular, poorly developed, retrac-
tively slanting axial riblets which become evanescent on the last part
of the last turn. Suture strongly impressed. Periphery feebly an-
gulated. Base moderately openly umbilicated, inflated, well-rounded,
and marked by the same type of sculpture as that which characterizes
the upper surface. Aperture broadly oval with a slight auricle at the
posterior angle; peristome simple; the outer lip produced at its junc-
tion with the basal lip and slightly reflected, thin; the inner slightly
thickened. The parietal wall is covered by a thick callus, which is
separated from the preceding turn by an impressed groove, which
renders the peritreme complete.
The type, U.S.N.M. No. 104432, was collected by Cuming at Salengo
Island, Ecuador. It has 4.6 whorls and measures: Height, 21.5 mm.;
greater diameter, 37.4 mm.; lesser diameter, 25.4 mm. Height of
aperture, 13.5 mm.; diameter, 17.7 mm.
The appressed summit distinguishes this species from the other
members of the group having a moderately broad umbilicus.
428349—42——_17
254 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) MASVENSE, new species
PLATE 37, Figures 1-3
Shell helicoid. The denuded early whorls pinkish, the rest oli-
vaceous with a brownish tinge. There are also irregular axial
streaks of a darker shade. There is a narrow light zone immediately
above the periphery and a broad dark band about one-fifth of the
width of the base anterior to the periphery. The rest of the base
is olivaceous horn colored; interior of the aperture bluish, showing
the dark band within. The nucleus consists of 2 small, well-rounded,
smooth turns. The postnuclear whorls are strongly rounded, the
last one decurrent for some little distance behind the aperture, hav-
ing a strongly impressed groove a little anterior to the summit.
The postnuclear whorls are marked by rather coarse, retractively
curved, and somewhat irregularly developed axial riblets, which be-
come decidedly weakened on the last quarter of the last turn. In
the groove below the summit there are oblique scratches. Sutures
very strongly impressed, almost channeled. Periphery obsoletely
angulated. Base narrowly openly umbilicated, inflated, strongly
rounded, and marked by the continuation of the axial ribs, which
increase but slightly on the umbilical wall. Aperture subcircular
with a conspicuous angulation at the posterior angle; peristome
simple; outer lip thin, slightly expanded; the inner lip somewhat
thickened. The parietal wall is covered by a thick callus, which is
separated from the preceding turn by an impressed line, which
renders the peritreme complete.
The type, U.S.N.M. No. 524048, was collected by Oscar Haught at
Cerro Masve, Guayas Province, Ecuador. It has 4.5 whorls and
measures: Height, 18.9 mm.; greater diameter, 30.2 mm.; lesser di-
ameter, 21.1mm. Height of aperture, 11.8 mm.; diameter, 14.1 mm.
U.S.N.M. No. 524049 contains 15 topotypes from the same source.
The more elevated spire will readily distinguish this species from
A. (A.) agassizi, which also has the axial sculpture comparatively
stronger.
APEROSTOMA (APEROSTOMA) AGASSIZI, new species
Puate 37, Ficures 4-6
Shell depressed-helicoid; the early denuded whorls pink; the last
covered with a chestnut-brown periostracum, which is paler on the
early whorls than on the last. There is a small white zone imme-
diately above the periphery, and a broad dark chestnut brown band,
which covers about a quarter of the base immediately below the
periphery. The rest of the base is yellowish horn colored. The
nucleus consists of 2 rather large, well-rounded, smooth turns. The
THE CYCLOPHORID LAND MOLLUSKS OF AMERTCA B55
postnuclear whorls are moderately well rounded, the last one having
a depressed area a little anterior to the summit. The summit itself,
at least on the last whorl, shows like a cord. The postnuclear whorls
are marked by retractively slanting axial riblets, which on the penul-
timate whorl are about as wide as the spaces that separate them; on
the last half of the last whorl they gradually become weakened and
finally evanescent. Suture very conspicously impressed. Periphery
of the last whorl feebly angulated. Base narrowly umbilicated,
inflated, strongly rounded, and marked by the continuation of the
axial ribs which become irregular and rather poorly emphasized on
the umbilical wall. Aperture large with a moderately conspicuous
angulation at the posterior angle; peristome simple; outer lip rather
produced, thin; the inner slightly thickened. The parietal wall is
covered by a thick callus, which is separated from the preceding turn
by an impressed line, rendering the peritreme complete.
The type, U.S.N.M. No. 57285, was collected by Louis Agassiz in
Brazil, probably in the Amazon River region, since the Agassiz
expedition covered that field. It had 4.4 whorls and measures:
Height, 15.7 mm.; greater diameter, 28.3 mm.; lesser diameter, 19
mm. Height of aperture, 11.1 mm.; diameter, 13.7 mm.
This species is readily distinguished from A. (A.) nevadense by
its deeply impressed suture.
APEROSTOMA (APEROSTOMA) NEVADENSE, new species
PLATE 37, Ficures 15-17
Shell helicoid, the upper surface covered by a brownish olivaceous
periostracum, which on the last half of the last turn becomes olive.
There is a narrow, faint, spiral, lighter zone immediately above the
periphery. The base has a pale olivaceous-brown band covering one-
fifth of the base, anterior to the periphery. The rest is olivaceous
horn colored. The nucleus consists of 2, rather large, well-rounded,
smooth turns. The post-nuclear whorls are well rounded, the last one
is slightly flattened toward the suture. The postnuclear whorls are
marked by very poorly developed, irregular, somewhat wavy axial
riblets. Suture weakly impressed. On the last half of the last turn
the riblets become decidedly irregular and almost obsolete. Periphery
obsoletely angulated. Base narrowly umbilicated, inflated, well
rounded, and marked by irregularly developed and spaced riblets,
which give it a rough surface, even rougher than the spire. Aperture
broadly oval, weakly angulated at the posterier angle; peristome
simple; the outer lip thin; the inner slightly thickened and slightly
reflected. The parietal wall is covered by a moderately thick callus,
which renders the peritreme complete.
256 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
The type, U.S.N.M. No 206494, was collected by Gabaldon and
sons at an altitude of 8,400 ft., in the Sierra Nevada Mountains,
Venezuela. It has 4.3 whorls and measures: Height, 15.8 mm.;
greater diameter, 28.5 mm.; lesser diameter, 19.8 mm. Height of
aperture, 10.9 mm.; diameter, 138 mm.
The rough irregular sculpture and weakly impressed suture will
readily differentiate this species from A. (A.) agassizi.
THE GROUP OF APEROSTOMA (APEROSTOMA) FASCIATUM
Small shells with rather narrow umbilicus and small aperture.
APEROSTOMA (APEROSTOMA) FASCIATUM (Kobelt)
PLATE 37, FiaurEs 7-9
1912. Neocyclotus popayanus fasciata Kopsetr, Martini-Chemnitz Conchylien
Cabinet, vol. 1, sect. 19, pt. 3, p. 900 (pl. 7, figs. 7-10), pl. 189, figs. 1-5.
1923. Poteria (Neocyclotus) popayana fasciata H. B. BAxer, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 137, 40, 44.
1928. Poteria (Neocyclotus) dunkeri var. approaching P. popayana H. B. BAKER,
Occ. Pap. Mus. Zool. Univ. Michigan, No. 137, pp. 40, 45, pl. 5, fig. Ty.
Shell small, helicoid, covered with an olivaceous-brown periostracum,
which fades to olivaceous on the last portion of the last turn. In
addition, there is a pale band immediately above the periphery and in-
dications of additional inconspicuous spiral lighter zones on the upper
surface. There isa dark band of brown about one-fifth width of base
immediately anterior to the periphery and in addition to that lesser
spiral bands of brown on the anterior half of the base; the rest is
olivaceous horn colored; the interior of aperture is bluish white. The
nucleus consists of 2 moderately large, well-rounded, smooth turns.
The postnuclear whorls are well elevated, inflated, and strongly
rounded. ‘The last one is deflected below the periphery for almost a
turn. The postnuclear whorls are marked by feeble, retractively
curved, hairlike riblets, which become much enfeebled on the last
quarter of the last turn. Suture strongly impressed. Periphery well
rounded. Base narrowly umbilicated, inflated, well rounded and
marked by the feeble continuation of the axial ribs on the posterior
half; on the anterior half and the umbilical wall the axial ribs become
very much intensified. Aperture rather small, circular, with a slight
angulation at the posterior angle; peristome simple; the outer lip
slightly flaringly expanded; the inner somewhat thickened. A mod-
erately thick callus covers the parietal wall. It is separated from the
preceding turn by an impressed line and renders the peritreme com-
plete. Operculum typically aperostomid, showing 9 whorls.
The specimen described and figured is one of two, U.S.N.M. No.
57766, which comes from Puerto Cabello, Venezuela. It has 43
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 257
whorls and measures: Height, 16 mm.; greater diameter, 21.8 mm.;
lesser diameter, 16.3 mm. Height of aperture, 9.4 mm.; diameter,
10.3 mm.
U.S.N.M. No. 316273 contains 2 additional specimens labeled
“Venezuela.”
While this species resembles A. (A.) pazi (Crosse) in having the
last whorl deflected, it can be distinguished from it in being much
darker in coloration and in having the umbilicus narrower.
APEROSTOMA (APEROSTOMA) PAZI (Crosse)
PLATE 37, FicurEs 12-14
1866. Cyclotus pazi Crossr, Journ. Conchyl., vol. 14, p. 356, pl. 14, fig. 3.
1897. Neocyclotus (Neocyclotus) pazi KopeLr and MOLLENDoRFF, Nachrb. deutschen
malak. Ges., vol. 29, p. 187.
1923. Poteria (Neocyclotus) pazi H. B. BAKrr, Occ. Pap. Mus. Zool. Univ. Michi-
gan, No. 187, p. 41.
Shell small, helicoid, covered by an olivaceous periostracum, the
early whorls of which have a brownish tinge. There are irregular
streaks of darker axial bands present. There is a narrow pale zone
immediately posterior to the periphery and a weak brownish band
about one-quarter the width of the base anterior to the periphery. The
rest of the base is horn colored; interior of aperture bluish white. The
nucleus consists of two moderately large, well-rounded, smooth turns.
The postnuclear whorls are strongly rounded; the summit of the last
one falls very materially below the periphery of the preceding turn.
The postnuclear whorls are marked by fairly strong, retractively slant-
ing axial riblets, which are about as wide as the spaces that separate
them. On the last half of the last turn these riblets become very
irregular, more or less fused, and quite obsolete on the last portion.
Periphery obsoletely angulated. Base moderately broadly umbili-
cated, inflated, strongly rounded, and marked by the continuation of
the axial sculpture, which becomes a little heavier and more irregular
on the umbilical wall. Aperture almost circular, very slightly angu-
lated at the posterior angle; peristome simple; outer lip thin; inner
lip slightly thickened. The parietal wall is covered by a moderately
thick callus which is separated from the preceding turn by an
impressed line rendering the peritreme complete.
U.S.N.M. No. 307401 contains two specimens, one of which has served
for our figure. They were collected by Cuming on Chimborazo,
Ecuador. The specimen figured has 41/4, whorls and measures: Height,
11 mm.; greater diameter, 22 mm.; lesser diameter, 18.5 mm. Height
of aperture, 9.5 mm.; diameter, 9.9 mm.
This species resembles A. (A.) fasctatum (Kobelt) in having the
last whorl decidedly deflected but differs from it in being much paler
and in having the umbilicus wider.
258 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
APEROSTOMA (APEROSTOMA) CAUCAENSE (Da Costa)
PLATE 37, FicureEs 10, 11
1901. Neocyclotus caucaensis DA Costa, Proc. Malac. Soc. London, vol. 4, p. 240,
pl. 24, fig. 9.
1923. Poteria (Neocyclotus) caucaensis H. B. Baker, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 187, p. 41.
We have not seen specimens of this species and so give a trans-
lation of Da Costa’s description and copy his figure:
Shell moderately profoundly umbilicated, subdiscoid, vivid oliva-
ceous green with the interior of the aperture white and narrowly
banded at and below the periphery. Whorls 4, impressed at the
suture; the last canaliculate below. Aperture circular; peristome
thin, with a cut in the basal lip. Greater diameter, 30 mm.; lesser,
23.5 mm.
Habitat: Province of Cauca, Colombia.
Da Costa then adds: “Of the two specimens received of this singu-
lar shell one was returned by the writer in hopes that others might
be discovered, but without result. In both of the shells sent the
fissure on the lip occurs in precisely the same place, just below the
shallow canal that surrounds the base; a similar fissure is found
in V. giganteus, var. Fischerianus, Hid., from Ecuador, but that
form has no canal on the base.”
This species looks to us as if its peculiarities were due to an in-
jury; however, not having seen it or anything like it, we merely
cite it as described and figured.
APEROSTOMA (APEROSTOMA) COLOMBIENSE (Da Costa)
PLATE 88, FicuRES 22-24
1901. Neocyclotus colombiensis Da Costa, Proc. Mal. Soc. London, vol. 4, p. 240,
pl. 24, fig. 7 (not Cyclostoma colombiensis Férussac MS., Orbigny, 1835).
1923. Poteria (Neocyclotus) colombiensis H. B. BAKER, Occ. Pap. Mus. Zool.
Univ. Michigan, No. 187, p. 48.
Shell helicoid, the upper portion brownish olivaceous. There is
a narrow light band immediately above the periphery and a narrow
dark chestnut-colored band immediately below the periphery. There
is a second dark chestnut-colored band of the same intensity and
width on the middle of the base. Between these two there are
numerous slender, spiral hairlines of dark brown. The rest of the
base is olivaceous horn colored; interior of aperture bluish white.
The nucleus consists of 2 rather large, well-rounded, small turns.
The postnuclear whorls are strongly rounded; the last one drops
considerably below the periphery of the preceding turn and bears
a conspicuous rounded cord at the appressed summit. Below this
cord is an impressed groove. The postnuclear whorls are marked by
HE CYCLOPHORID LAND MOLLUSKS OF AMERICA 259
retractively curved feeble riblets, which are rather irregular in
strength and spacing, and which become weaker on the last turn
and obsolete on the last portion thereof. Suture well impressed.
Periphery weakly angulated. Base narrowly umbilicated, inflated,
strongly rounded, and marked by the continuation of the feeble axial
sculpture, which, however, develops into stronger and broader riblets
on the umbilical wall. Aperture circular with a moderately strong
angulation at the posterior angle; peristome simple; outer lip thin,
slightly expanded and slightly reflected; inner lip moderately thick-
ened. The parietal wall is covered by a heavy callus, which is
strongly separated from the preceding turn and renders the peritreme
complete.
The specimen described and figured, U.S.N.M. No. 307482, was
received from Sowerby and Fulton with the designation “New
Grenada” (Colombia). It has 4.4 whorls and measures: Height,
18.7 mm.; greater diameter, 27.7 mm.; lesser diameter, 20.2 mm.
Height of aperture, 11 mm.; diameter, 12.8 mm.
Da Costa’s figure yields the following measurements: Height, 21.1
mm.; greater diameter, 33.5 mm.; lesser diameter, 24.7 mm. Height
of aperture, 13.4 mm.; diameter, 14.9 mm. It has 5 whorls.
This species recalls A. (A.) caucaense (Da Costa) but lacks the
basal sinus or configuration described therefor. It also suggests
A, (A.) perezi (Hildalgo), but is much larger than that species.
APEROSTOMA (APEROSTOMA) PEREZI (Hidalgo)
PLATE 38, FIGURES 4-6
1866. Cyclotus perezi Hi~paco, Journ. Conchyl., vol. 14, p. 344, pl. 14, fig. 2.
1897. Neocyclotus (Neocyclotus) perezi Kosett and MO6.iLEeNnporFr, Nachrb.
deutschen malak. Ges., vol. 29, p. 137.
1923. Poteria (Neocyclotus) perezi H. B. BaKEr, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 137, p. 43.
Shell small, depressed-helicoid, the upper surface bright reddish
chestnut-brown. There is a very inconspicuous, narrow, slightly
lighter zone immediately above the periphery. The base has a
broad dull chestnut-brown band immediately below the periphery
and another narrow one of the same shade on the middle of the
base. The rest is a little paler than the upper surface. The nucleus
consists of 2 moderately large, well-rounded, smooth turns. The
postnuclear whorls are inflated, strongly rounded, almost flattened
at the summit. The summit of the last one falls slightly below
the periphery of the preceding turn. The postnuclear whorls are
marked by regular fine riblets. Suture strongly constricted. Pe-
riphery well rounded. Base openly umbilicated and marked by
the continuation of the axial riblets, which become a little less
260 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
strong on the umbilical wall. Aperture circular with a slight angu-
lation at the posterior angle; peristome simple; outer lip thin and
the inner slightly thickened. The parietal wall is covered by a
moderately strong callus which renders the peritreme complete.
U.S.N.M. No. 316104 contains two specimens from Quito, Ecuador.
The one figured has 4 whorls and measures: Height, 13.8 mm.;
greater diameter, 24.5 mm.; lesser diameter, 16.8 mm. Height of
aperture, 10.2 mm.; diameter, 11.1 mm.
The small size and more depressed form readily distinguish this
species from A. (A.) colombiense (Da Costa), and the regular rib-
bing of the upper surface differentiates it from A. (A.) redfieldi.
APEROSTOMA (APEROSTOMA) BOLIVIENSE, new species
PLATE 38, Fiaures 7-9
Shell helicoid, covered with a chestnut-brown periostracum, which
is a little paler near the suture than on the major portion of the
turns. There is a narrow, inconspicuous, lighter zone immediately
above the periphery. The base is marked by a broad chestnut-
brown band, which gradually fades from the periphery basally. This
is separated by a narrow band, which is about twice as wide as
the narrow dark chestnut-brown band below it, which occupies
the middle of the base. The general coloration of the base is oliva-
ceous horn colored; interior of aperture bluish white. The nucleus
consists of 2 small, well-rounded, smooth turns. The postnuclear
whorls are strongly rounded; the summit of the last one gradually
falls below the periphery of the preceding turn. The postnuclear
whorls are marked by rather strong, retractively curved axial riblets,
which grow somewhat weaker and irregular on the last part of
the last whorl. Suture very deeply impressed. Periphery obsoletely
angulated. Base narrowly umbilicated, inflated, strongly rounded
and marked by the continuation of the axial ribs, which grow very
slightly stronger on the umbilical wall. Aperture broadly oval, pro-
duced at the junction of the basal and outer lip; peristome simple;
outer lip sharp; inner somewhat thickened. The parietal wall is
covered by a thick callus, which is separated from the preceding
turn by a deeply impressed groove which renders the peritreme
complete.
The type, U.S.N.M. No. 307426, was received from Sowerby and
Fulton with the designation “Veocyclotus inca Orbigny var. Minor
Pfeiffer” and the locality “Bolivia.” It has 4.2 whorls and measures:
Height, 14.5 mm.; greater diameter, 23.9 mm.; lesser diameter, 16.5
mm. Height of aperture, 9.7 mm.; diameter, 10.9 mm.
U.S.N.M. No. 524102 contains one topotype from the same source.
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 261
This species resembles A. (A.) redfieldi. It has the suture much
more profoundly impressed and the median basal dark band more
nearly on the middle of the base.
APEROSTOMA (APEROSTOMA) REDFIELDI, new species
PLATE 38, FicuRES 10-12
Shell small, helicoid. The upper surface is dark chestnut-brown.
There is a broad light band immediately above the periphery, while
below the periphery there is a broad dark chestnut-brown band about
twice as wide as the light zone above, and a second narrow band occu-
pies the base a little posterior to the middle. The rest of the base is
olivaceous, with a brownish tinge; interior of the aperture bluish
white. The nucleus consists of about 2 well-rounded, smooth turns.
The postnuclear whorls are well rounded; the last one is appressed as
a feeble cord at the summit, below which there is an impressed groove.
The postnuclear whorls are marked by feeble, poorly developed axial
riblets which are of irregular size and spacing. On the last part of
the last turn they become quite obsolete. The summit of the last whorl
falls slightly below the periphery of the preceding turn. Periphery
obscurely angulated. Base narrowly umbilicated, inflated, strongly
rounded, and marked by the continuation of the axial ribs, which do
not appear to grow stronger on the umbilical wall. Aperture almost
circular, slightly angulated at the posterior angle; peristome simple;
the outer lip sharp; the inner slightly thickened. The parietal wall is
covered by a thick callus, which is separated from the preceding turn
by a strongly impressed groove, which renders the peritreme complete.
The type, U.S.N.M. No. 307399, comes from Brazil. It has 4.5
whorls and measures: Height, 14.5 mm.; greater diameter, 23.6 mm. ;
lesser diameter, 16.7 mm. Height of aperture, 8.5 mm.; diameter,
10.3 mm.
This species is very closely related to A. (A.) boliviense but has the
height of the aperture proportionately smaller. The color band also
occupies a different position. In A. (A.) boliviense it is on the middle
of the base, while in this species it is more posterior. A. (A.) boli-
viense also has the suture much more strongly impressed.
APEROSTOMA (APEROSTOMA) CURRANI, new species
PLATE 38, Figures 1-3
Shell helicoid, soiled white when denuded. The fragments of the
periostracum remaining in the specimen before us show a golden-
brown tint on the upper surface with a dark subperipheral band on
the base. The nuclear whorls are decollated in our specimen. The
postnuclear whorls are well rounded and marked by retractively
262 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
slanting axial riblets. Suture strongly impressed. Periphery ob-
scurely angulated. Base moderately broadly umbilicated, inflated,
well rounded, with the axial riblets growing a little stronger on the um-
bilical wall. Aperture subcircular ; peristome simple; outer lip slightly
expanded and reflected; inner moderately thickened. The parietal
wall is covered by a rather heavy callus, which is separated from the
preceding turn by an impressed line.
The type, U.S.N.M. No. 322360, was collected by H. M. Curran on
the Rio Gregugy, Bahia, Brazil. It has a little more than 2 whorls
remaining and measures: Height, 16 mm.; greater diameter, 27.5 mm.;
lesser diameter, 10.8 mm. Height of aperture, 12.1 mm.
We dislike to describe imperfect material as new species but are
making an exception in this case because the specimen in question
represents an element completing the range of the group to which
it belongs.
The species can be easily differentiated from A. (A.) merrilli by
its larger size.
APEROSTOMA (APEROSTOMA) NANUM, new species
PLATE 38, Ficures 13-15
Shell small, helicoid; the upper surface horn yellow with a narrow
light zone immediately above the periphery. Below the periphery
there is a narrow blackish-brown zone. The rest of the base is of the
same color as the upper surface. The nucleus consists of 2 well-
rounded, smooth turns. The postnuclear whorls are well rounded; the
last one has an inconspicuous cord at the summit immediately adjacent
to which is an impressed groove. The summit of the last half turn of
the last whorl is gradually deflected until it falls below the dark sub-
peripheral band. The postnuclear whorls are marked by retractively
slanting axial riblets, which are about as wide as the spaces that sep-
arate them. Suture very strongly impressed. Periphery obscurely
angulated. Base narrowly umbilicated, inflated, well rounded, marked
by the continuation of the axial ribs, which become very much enfeebled
on the last half. These axial riblets become stronger on the umbilical
wall. Aperture circular, slightly angulated at the posterior angle;
peristome simple; outer lip thin; inner lip somewhat thickened. The
narrow parietal wall is covered by a thick callus which renders the
peritreme complete.
The type, U.S.N.M. No. 336128, was collected by H. Pittier on the
coastal range somewhere between sea level and 200 meters between
Caracas and Puerto Cabello, Venezuela. It has 4 whorls and meas-
ures: Height, 18.8 mm.; greater diameter, 21.7 mm.; lesser diameter,
15.8 mm. Height of aperture, 8.5 mm.; diameter, 10.1 mm.
THE CYCLOPHORID LAND MOLLUSKS OF AMERTCA 263
The small size and pale coloration readily distinguish this species
from the other members of the group.
APEROSTOMA (APEROSTOMA) MERRILLI, new species
PLATE 38, FieuRES 19-21
Shell low, helicoid. The upper surface olivaceous, with a brownish
tinge. There is a broad light zone immediately above the periphery
and a broad dark zone covering about two-fifths of the base immedi-
ately below the periphery. The rest of the base is a little paler than
the upper surface. The nucleus consists of 2 small, well-rounded,
smooth turns. The postnuclear whorls are strongly rounded. The
last one is gradually deflected until it falls about halfway across the
dark subperipheral band at the aperture. The postnuclear whorls
are marked by retractively curved, slender axial riblets, which are
not quite as wide as the spaces that separate them. These riblets be-
come somewhat confused and irregular on the last half of the last
turn. Suture strongly impressed. Periphery obscurely angulated.
Base openly, rather broadly umbilicated, inflated, strongly rounded,
and marked by the continuation of the axial riblets, which become
obsolete on the umbilical wall. Aperture almost circular with a
slight angulation at the posterior angle; peristome simple; outer lip
thin; inner slightly thickened. The parietal wall is covered by a
moderately thick callus which renders the peritreme complete.
The type, U.S.N.M. No. 58310 comes from Brazil. It has 43
whorls and measures: Height, 14.6 mm.; greater diameter, 23.7 mm,;
lesser diameter, 17 mm. Height of aperature, 9.5 mm.; diameter,
10.9 mm.
U.S.N.M. No. 203708 contains five not quite perfect specimens ob-
tained by H. B. Merrill on the Braganza Railway in the Amazon
Valley; U.S.N.M. No. 57286 contains another specimen obtained by
the Agassiz Expedition to Brazil.
The larger size, darker coloration, and the feeble sculpture on the
umbilical wall readily distinguish this species from A. (A.) nanum.
APEROSTOMA (APEROSTOMA) PULCHELLUM, new species
PLATE 38, Ficures 16-18
Shell small, depressed-helicoid, thin. The early whorls with a
pinkish tinge; the later pale horn colored. There is an inconspicuous
light zone immediately above the periphery and a moderately broad
chestnut band immediately below it. The rest of the base is colored
like the upper surface of the last whorl. The nucleus consists of 2
rather large, well-rounded, smooth turns. The postnuclear whorls
are well rounded, slightly flattened toward the summit, or even some-
264 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
what inbent. The summit of the last turn gradually falls below the
periphery of the preceding turn at the aperture, and is more than
the width of the brown band below the brown band. The postnu-
clear whorls are marked by feeble axial riblets, which are merely
indicated on the last half of the last turn. Suture very deeply im-
pressed. Base broadly openly umbilicated, somewhat inflated, well
rounded and marked by the continuation of the axial ribs, which do
not become intensified on the umbilical wall. Aperture very large;
peristome simple; outer lip expanded and slightly reflected; inner
lip slightly thickened. A very heavy callus covers the parietal wall,
from which it is separated by a strongly impressed groove, which
renders the peritreme complete. Operculum typically aperostomid,
showing 9 whorls.
The type, U.S.N.M. No. 524125, comes from Brazil. It has 4.3
whorls and measures: Height, 12.1 mm.; greater diameter, 22.1 mm. ;
lesser diameter, 15 mm. Height of aperture, 6.9 mm.; diameter,
10.5 mm.
The pale coloration, obsolete supraperipheral light zone, and feeble
axial sculpture on the umbilical wall readily differentiate this species
from A. (A.) merrilli.
APEROSTOMA (APEROSTOMA) POPAYANUM (Lea)
PLATE 39, FiauRES 17-19 (type), 20-22
1839. Cyclostoma popayana LEA, Trans. Amer, Phil. Soc., new ser., vol. 6, p. 94,
pl. 23, fig. 76.
1847. Aperostoma popayanum Pretrrer, Zeitsch. f. Malak., vol. 4, p. 104.
1852. Cyclotus popayanus PretFreR, Monographia pneumonopomorum viventium,
vol. 1, p. 21.
1897. Neocyclotus (Neocyclotus) popayanus Kosetr and MO.tienporrr, Nachrb.
deutschen malak. Ges., vol. 29, p. 187.
1928. Poteria (Neocyclotus) popayana H. B. Baxer, Occ. Pap. Mus. Zool. Univ.
Michigan, No. 187, p. 44.
Shell helicoid, the upper surface olivaceous horn colored with axial
streaks of darker shades, which are of irregular size and spacing.
There is a narrow light zone immediately above the periphery and
a brownish olivaceous band below the periphery, which gradually
fades basally. The rest of the base is horn colored. The nucleus
consists of 2 small, well-rounded, smooth turns, forming a rather
elevated spire. The postnuclear whorls are well rounded; the last
one slightly deflected below the periphery, forming a weak cord at
the appressed summit, below which there is an impressed groove.
The postnuclear whorls are marked by retractively curved slender
axial riblets, which are narrower than the spaces that separate them,
and which on the last turn become rather irregular. Here, too, there
are oblique scratches in the depressed area below the summit and, to
THE CYCLOPHORID LAND MOLLUSKS OF AMERICA 265
a lesser extent, on the rest of the whorl. Periphery obscurely angu-
lated. Base narrowly umbilicated, and marked by the feeble con-
tinuation of the axial ribs, which do not become intensified on the
umbilical wall. Aperture circular with a conspicuous angulation at
the posterior angle; peristome simple; outer lip thin, somewhat ex-
panded and slightly reflected on the basal half; slightly thickened on
the inner lip with the parietal wall covered by a moderately thick
callus which is separated from the preceding turn by an impressed
line, which renders the peritreme complete. Operculum typically
aperostomid, showing 9 whorls.
Lea’s type, U.S.N.M. No. 104453, was collected by Dr. Gibbons near
Popayan, Grenada (that is, southwest Colombia). It has 4.2 whorls
and measures: Height, 14.7 mm.; greater diameter, 22.6 mm.; lesser
diameter, 16 mm. Height of aperture, 9.8 mm.; diameter, 10.7 mm.
A second specimen with the type and bearing the same number
is considerably larger but in every way similar to the dead denuded
type. It measures: Height, 18.4 mm.; greater diameter, 27.5 mm.;
lesser diameter, 19 mm. Height of aperture, 10.8 mm.; diameter,
12.6 mm. It has 4.6 whorls.
We have also figured one of two specimens received from Bland,
U.S.N.M. No. 307406, collected at Marmato, New Grenada (south-
western Colombia). It has 4.4 whorls and measures: Height, 15.4
mm.; greater diameter, 23.5 mm.; lesser diameter, 17 mm. Height
of aperture, 10.4 mm.; diameter, 10.8 mm.
U.S.N.M. No. 356029 contains two young specimens from New
Grenada.
This species appears nearest related to A. (A.) balsasense, from
which it can readily be differentiated by the scratched surface of the
last whorl, which is not present in balsasense.
APEROSTOMA (APEROSTOMA) BALSASENSE, new species
PLATE 39, Fiaures 1-3
Shell depressed-helicoid. The early whorls are flesh colored; the
last two horn colored with a brownish flush. There is a very narrow
light zone immediately above the periphery and a broad dark zone
below the periphery which extends from the periphery to almost the
middle of the base, gradually fading anteriorly. The anterior portion
and umbilical wall are horn colored; the interior of the aperture is
flesh white and shows the external banding. The nucleus consists of
2 small, well-rounded, smooth turns. The postnuclear whorls are well
rounded, appressed at the summit and marked on the penultimate
whorl by very regular and regularly spaced retractively curved axial
ribs. These ribs are about as wide as the spaces that separate them.
On the last whorl the riblets become somewhat irregular and toward
266 BULLETIN 181, UNITED STATES NATIONAL MUSEUM
the end evanescent. The summit of the last whorl drops slightly below
the periphery. Base openly umbilicated, inflated, well rounded, and
marked by the feeble continuation of the axial ribs, which almost
vanish on the umbilical wall. Aperture circular, with a slight angu-
lation at the posterior angle; peristome simple; outer lip very thin;
inner slightly thickened. The parietal wall is covered with a thin
callus which renders the peritreme complete.
The type, U.S.N.M. No. 524126, comes from Balsas, Peru. It has 4
whorls and measures: Height, 13.8 mm.; greater diameter, 23.3 mm.;
lesser diameter, 16.4mm. Height of aperture, 9.3 mm.; diameter, 10.2
mm. .
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 1
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 283.
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 2 3
*
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CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANAT'ON OF PLATE SEE PAGE 283.
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 3
19
18
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 283,
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 4
6
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 283,
U. S NATIONAL MUSEUM BUEEEBRIN 18” iREATE.S
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 284,
U. S. NATIONAL MUSEUM BULLETIN 181
26
99
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CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 284,
PEATE 6)
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 7
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANAT:ON OF PLATE SEE PAGE 284.
U. S. NATIONAL
MUSEUM BULLETIN 181
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 284.
PLATE 8
a
U. S. NATIONAL MUSEUM BUBEERING 81>] PIEATE 9
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGES 284, 285.
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 10
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 285,
U. S. NATIONAL MUSEUM BUELERINGISi) PEATE 11
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 285,
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 12
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 285,
U. S. NATIONAL MUSEUM BUEEE GIN 181 PEATE 13
21 24 eu 30 =
42
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGES 285, 286,
U. S. NATIONAL MUSEUM BULLETIN 181
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 286
PLATE 14
U. S. NATIONAL MUSEUM BUEECERIN 181 PEATE TS
9
oe
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 286,
U. S. NATIONAL MUSEUM BULLETIN 181
36
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 286,
PLATE 16
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 17
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGES 286, 287.
U. S. NATIONAL MUSEUM BULLETIN 181
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 287.
PLATE 18
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 19
wy
15
16
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 287.
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 20
CYCLOPHORID MOLLUSKS OF AMERICA,
FOR EXPLANATION OF PLATE SEE PAGE 287
U. S. NATIONAL MUSEUM BULEEE RING 138i) JPEATKE 21
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGES 287, 288.
U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 22
CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 23
CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 24
10 - 15
CYCLOPHORID MOLLUSKS OF AMERICA.
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CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 26
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CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 27
CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BUEEETIN 181. sPEATE 28
32 |
33
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20
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 33
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U. S. NATIONAL MUSEUM BULLETIN 181 PLATE 34
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U. S. NATIONAL MUSEUM BUEEETIN 181° (PEATE 37
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U. S. NATIONAL MUSEUM BULEEIN 181 PEATE 38)
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CYCLOPHORID MOLLUSKS OF AMERICA.
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U. S. NATIONAL MUSEUM BUEEETIN 181 PEATE Ail
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BULLETIN 181 PLATE 42
CYCLOPHORID MOLLUSKS OF AMERICA.
FOR EXPLANATION OF PLATE SEE PAGE 291,
INDEX
(Principal references are given in boldface type.)
abbotti, Crocidopoma, 62, 67, 285.
acutiliratus, Amphicyclotulus, 54, 56,
285.
Cyclophorus, 56, 141.
adamsi, Ptychocochlis, 83, 94, 95,
Adelopoma, 51, 148.
bakeri, 148, 149, 282, 291.
costaricense, 52, 150, 280, 291.
occidentale, 52, 148, 149, 150,
282, 291.
stolli, 149, 281, 291.
tuema, 148, 279, 291.
affine, Aperostoma 204, 210, 281, 289.
affinis, Cyclotus, 210.
Neocyclotus, 210.
Poteria, 210.
agpesi Aperostoma, 223, 254, 256,
290.
286.
151,
279,
allantayum, Aperostoma, 221, 230, 282,
289.
alutaceum, Cyclostoma, 22.
Farcimen, 4, 15, 21, 22, 23, 24,
amazonense, Aperostoma, 222, 243,
290.
ambiguum, Aperostoma, 203, 205,
281, 289.
ambiguus, Cyclotus, 211.
Neocyclotus, 211.
Poteria, 211.
amethystinus, Amphicyclotulus, 60,
Cyclotus, 60.
Amphicyclotinae, 43, 52, 142, 151.
Amphicyclotulus, 52, 54, 57, 60, 62.
acutiliratus, 54, 56, 285.
amethystinus, 60, 285.
beauianus, 60, 61, 285.
dominicensis, 54, 57, 285.
guadeloupensis, 54, 55, 58, 60,
liratus, 54, 58, 60, 285.
mineri, 54, 55, 58, 60, 285.
perplexus, 54, 59, 285.
portoricensis, 54, 59, 285.
rufescens, 54, 56, 57, 285.
schrammi, 54, 57, 58, 285.
Amphicyclotus, 151, 183.
boucardi, 183, 184, 281, 288.
bourcieri, 163.
cayennensis, 276.
chanchopoyasensis, 162.
cooperi, 180.
cousini, 166.
crosseanus, 156.
cumingii, 167.
delphinulus, 158.
428349—42
283.
279,
211,
285.
285.
20
Amphicyclotus esmeraldensis, 160.
gayi, 278.
goldfussi, 184, 281.
guayaquilensis, 161.
haematomma, 155.
hidalgoi, 167.
lutescens, 181.
maleri, 184, 185, 220, 281, 288.
nigrofasciatus, 164.
olssoni, 178.
orbignyi, 168.
ponderosus, 182.
psilomitus, 274.
rosenbergi, 160.
texturatus, 184, 186, 281, 288.
underwoodi, 176.
vesconesi, 156.
anafense, Farcimen, 27, 28, 284.
angasianum, Aperostoma, 177.
angulatus, Cyclotus, 228.
antillarum, Cyclostoma, 49.
Megalomastoma, 49, 50, 284.
antoni, Lagocyclus, 154, 156, 280, 287.
antonii, Cyclophorus, 154.
Aperostoma, 62, 124, 186, 187, 221, 277.
affine, 204, 210, 281, 289.
agassizi, 223, 254, 556, 279, 290.
allantayum, 221, 230, 282, 289.
amazonense, 222, 243, 279, 290.
ambiguum, 203, 205, 211, 281, 289.
angasianum, 177.
aulari, 195, 196, 199, 200, 282, 288.
aureum, 204, 208, 209, 281, 289.
bairdianum, 125, 126, 127 287.
balsasense, 228, 265, 266, 282, 291.
bartletti, 157.
bejumense, 268, 269, 270, 272, 282,
289,
belli, 268, 269, 279, 289.
berendti, 205, 211, 212, 281, 289.
bisinuatum, 222, 235, 280, 290.
blanchetianum, 187, 221, 222, 244,
246, 247, 279, 290.
bogotense, 221, 226, 227, 279, 289.
boliviense, 223, 260, 261, 279, 290.
brujense, 222, 241, 281, '290.
burringtoni, 196, 202, 282, 289.
carabobense, 196, 201, 282, 289.
cardozi, 221, 223, 266, 282, 291.
carmioli, 221, 233, 234, 280, 290.
castaneum, 222, 251, 252, 282, 290.
caucaense, 223, 258, 259, 280, 290.
chrysacme, 203, 219, 281, 289.
cingulatum, 221, 228, 229, 280, 289.
293
294
Aperostoma colombiense, 223, 258, 280,
290.
confusum, 221, 222, 237, 280, 281,
290.
connivens, 275.
cookei, 205, 215, 279, 281, 289.
cooperi, 180.
corpulentum, 2038, 217, 218, 279, 289.
corrugatum, 105.
costaricense, 221, 234, 280, 290.
cumingi, 221, 224, 279, 289.
cumingii, 167.
currani, 223, 261, 279, 290.
deburghaeanum, 129, 130, 287.
depressum, 222, 247, 282, 290.
duffiianum, 276.
dunkeri, 221, 230, 233, 280, 289.
dunoonense, 269, 2738, 279, 289.
dyeri, 204, 205, 281, 289.
dysoni, 203, 204, 207, 209, 217, 281,
289.
ecuadorense, 222, 248, 250, 251, 280,
290.
exiguum, 221, 234, 280, 289.
fasciatum, 223, 256, 257, 282, 290.
filoliratum, 221, 267, 280, 291.
fischeri, 222, 239, 280, 290.
fossile, 125, 131, 287.
fultoni, 222, 242, 279, 290.
giganteum, 221, 222, 237, 238, 239,
277, 281, 290.
glaucostomum, 196, 199, 282, 288.
goldfussi, 164.
geranulatum, 196, 280, 288.
grenadense, 133, 134, 135, 287.
haughti, 268, 269, 279, 289.
hedui, 187, 191, 279, 288.
hinkleyi, 203, 204, 206, 207, 281, 289.
hitomi, 187, 194, 280, 288.
humile, 129, 287.
inea, 221, 267, 279, 291.
incomptum 187, 192, 282, 283.
inconspicuum, 221, 227, 282, 289.
irregulare, 222, 236, 280, 290.
jamaicense, 105.
kobelti, 187, 199, 279, 288.
kugleri, 196, 201, 282, 289.
lacteofiuviale, 98.
laxatum, 222, 244, 280, 290.
leai, 221, 222, 246, 282, 290.
limellum, 196, 197, 199, 201, 279,
288.
lutescens, 181.
major, 269, 270, 279, 289.
malleatum, 187, 188, 189, 190, 191,
282, 288.
manabense, 222, 239, 280, 290.
masvense, 223, 254, 280, 290.
mcsweeni, 132, 134, 135, 287.
merrilli, 223, 262, 263, 264, 279, 290.
mexicanum, 173.
montezumi, 151, 152.
moricandi, 219, 279, 291.
nanum, 223, 262, 263, 282, 290.
nevadense, 233, 255, 282, 290.
nicaraguense, 205, 214, 216, 281,
289.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Aperostoma nirafe, 187, 193, 195, 279,
288
olivaceum, 222, 251, 252, 280, 290.
paezense, 221, 225, 279, 289.
paezicolum, 221, 232, 280, 290.
pailaense, 221, 231, 280, 289.
palenquense, 183.
panamense, 196, 197, 281, 288.
pazi, 217, 228, 257, 280, 290.
peilei, 196, 200, 279, 288.
perdistinctum, 276.
perezi, 223, 259, 280, 290.
pergrandis, 187, 188, 190, 279, 288.
peruense, 222, 245, 247, 282, 290.
peruvianum, 2238, 252, 282, 290.
pichinchense, 187, 191, 280, 288.
pittieri, 222, 236, 280.
pizarroi, 187, 193, 194, 195, 282, 288.
popayanum, 223, 264, 266, 280, 291.
portobellense, 222, 242, 281, 290.
pretiosum, 126, 127, 287.
princeps, 175.
pulchellum, 228, 263, 279, 299.
pygmaeum, 269, 272, 282, 289.
quitense, 222, 250, 280, 290.
redfieldi, 223, 261, 279, 290.
ruatanense, 204, 207, 281, 289.
ruber, 125, 126, 127, 128, 287.
rudisplanusque, 125, 131, 287.
rugatum, 133, 286.
salengoense, 2238, 253, 280, 290.
salleanum, 170.
sallei, 205, 213, 281, 289.
sanctaemarthae, 203, 217, 218, 279,
289.
santaguitense, 289.
scabratum, 125, 129, 287.
seminudum, 109, 125, 128, 129, 130,
131, 287, 291:
simile, 221, 227, 280, 289.
smithi, 208, 204, 216, 217, 279, 289.
stirlingi, 187, 195, 280, 288.
stramineum, 132, 195, 196, 198, 282,
288.
subcingulatum, 222, 249, 251, 280,
290.
sumichrasti, 204, 209, 211, 281, 289.
translucidum, 136, 179, 268, 269,
272, 279, 289.
trinitense, 136, 268, 269, 271, 282,
287, 289.
tryonianum, 116.
umbilicatum, 221, 224, 225, 226, 279,
289.
utriaense, 222, 240, 241, 280, 290.
valerioi, 205, 213, 215, 280, 289.
venzuelense, 222, 247 248, 282, 290.
veracochanum, 221, 229, 282, 289.
vincentinum, 133, 286.
walkeri, 174.
wetmorei, 203, 279, 291.
Aperostominae, 3, 38, 43, 62, 142, 186.
apertum, Cyclostoma, 26.
Farcimen 5, 26.
Megalomastoma, 26, 27.
arangoi, Farcimen, 36, 284.
asperula, Cyclopilsbrya, 71, 78, 285.
INDEX
asperulum, Cyclostoma, 78.
asperulus, Cyclotus, 78.
Neocyclotus, 78.
atratensis, Calacyclotus 178, 179, 279,
288.
aulari, Aperostoma, 195, 196, 199, 200,
282, 288.
Poteria, 199.
aureum, Aperostoma, 204, 208, 209, 281,
289.
auriculatum, Cyclostoma, 16, 25.
Farcimen, 24, 25, 26, 284.
Austrocyclotus, 124, 132, 187, 195, 197,
202.
aulari, 199.
burringtoni, 202.
carabobense, 201.
glaucostomum, 199.
granulatum, 196.
grenadense, 134.
kugleri, 201. -
limellum, 197.
mesweeni, 135.
panamense, 197.
peilei, 290.
rugatum, 133.
stramineum, 198.
vincentinum, 133.
avus, Cyrtotoma, 169, 170, 281, 288.
bairdianum, Aperostoma, 125, 126, 127,
287.
bairdianus, Cyclotus, 127.
bakeri, Adelopoma, 148, 149, 282, 291.
TIncerticyclus, 137, 188, 139, 287.
Neocyclotus, 137.
balnearis, Poteria, 116, 123, 286.
balneorum, Farcimen, 31, 32, 284.
balsasense, Aperostoma, 223, 265, 266,
282, 291.
Barbacyclus, 151, 175.
boueardi, 175, 177, 280, 288.
princeps, 175, 280, 288.
underwoodi, 175, 176, 178, 280, 288.
bartletti, Aperostoma, 157.
Cyclotus, 157.
Lagocyclus, 154, 157, 282, 287.
Neocyclotus, 157.
Poteria, 157.
bayamense, Farcimen, 5, 6, 283.
beauiana, Cyclostoma, 60, 61.
beauianus, Amphicyclotulus, 60, 61, 285.
bejumense, Aperostoma 268, 269, 270,
202, 282, 289.
bejumensis, Poteria 270.
belli, Aperostoma 268, 269, 279,
Neocyclotus, 201, 268.
berendti, Asperostoma, 205,
281, 289.
Cyclotus, 212.
Neocyclotus, 212.
Poteria, 212.
beswicki, Cyclopilsbrya, 71, 79.
Cyclotus, 79.
Neocyclotus, 79.
biayaense, Farcimen, 12, 13, 14, 283.
bieincta, Buckleyia, 151, 152, 280, 287.
289.
Dlilpeeniee
295
bicolor, Cyclostoma, 25.
Farcimen, 24, 25, 284.
bifasciata, Buckleya, 153.
Buckleyia, 151, 153, 279, 287.
Cyclophorus, 153.
bisinuatum, Aperostoma, 222, 235,
290.
bisinuatus, Cyclotus, 235.
Neocyclotus, 235.
bituberculatum, Cyclostoma, 15.
Farcimen, 13, 14, 15, 16, 283.
blanchetianum, Aperostoma, 187, 221,
222, 244, 246, 247, 279, 290.
Cyclostoma, 124, 244, 267.
Poteria, 245.
bogotense, Aperostoma, 221, 226,
279, 289.
Cyclostoma, 226.
bogotensis, Cyclotus, 226.
Neocyclotus, 226.
Poteria, 226.
boliviense, Aperostoma, 223, 260,
279, 290.
bondi, Cyclojamaicia, 68, 69, 70, 285.
Poteria, 68.
poucargs Amphicyclotus, 183, 184,
288.
Barbacyclus, 175, 177, 280, 288.
Cyclophorus, 184.
Cyclostoma, 184.
Cyclotus, 175, 177.
Neocyclotus, 177.
bourcieri, Amphicyclotus, 163.
Calaperostoma, 159, 163, 164,
287.
Cyclophorus, 163.
Cyclostoma, 163.
bowdenensis, Incerticyclus,
291.
brasiliense, Cyclostoma, 277.
braziliensis, Incerticyclus, 277, 279, 291.
brujense, Aperostoma, 222, 241, 281, 290.
brunnea, Megalomastoma, 48, 49.
brunneum, Megalomastoma, 48, 49, 284,
285.
Buckleya, 151.
bifasciata, 153.
martinezi, 152.
Buckleyia, 151.
bicincta, 151, 152, 280, 237.
bifasciata, 151, 153, 279, 287.
martinezi, 151, 152, 154, 280, 287.
burringtoni, Aperostoma, 196, 202, 282,
28
Poteria, 107, 286.
Calacyclotus, 151, 178.
atratensis, 178, 179, 279, 288.
olssoni, 178, 179, 279, 281, 288.
Calaperostoma, 151, 159.
bourcieri, 159, 163, 164, 280, 287.
chanchapoyasense, 159, 162, 164,
So uaO Ke
cousini, 159, 166, 167, 280, 288.
cumingi, 159, 167, 169, 279, 288.
esmeraldense, 159, 160, 161, 280, 287.
guayaquilense, 159, 161, 280, 287.
hidalgoi, 159, 166, 167, 279, 287.
280,
261,
281,
280,
138, 139,
296
Calaperostoma leai, 159, 165, 279, 288.
nigrofasciatum, 159, 164, 166, 280,
288.
orbignyi, 159, 168, 279.
pittieri, 159, 164, 281, 287.
purum, 159, 162, 280, 287.
rosenbergi, 159, 160, 280, 287.
camagueyanum, Farcimen, 10, 11, 283.
campeachyi, Poteria, 104.
Ptychocochlis, 84, 103, 104, 286.
carabobense, Aperostoma, 196, 201, 282,
289.
eardozi, Aperostoma, 221, 223, 266, 282,
291.
Poteria, 266.
caribaea, Cyclopilsbrya, 71, 77, 286.
Poteria, 77.
carmioli, Aperostoma, 221, 233, 234, 280,
290.
castaneum, Aperostoma, 222, 251, 252,
282, 290.
casuelense, Crocidopoma, 62, 65, 291.
catalinense, Farcimen, 36, 284.
caucaense, Aperostoma, 223, 258, 259,
280, 290.
eaucaensis, Neocyclotus, 258.
Poteria, 258. r
cayennense, Cyclostoma, 276.
cayennensis, Amphicyclotus, 276.
Cyclophorus, 276.
Incerticyclus, 276, 281.
caymanensis, Cyclopilsbrya, 80, 285.
Poteria, 80.
Cerion truncatum, 45.
chanchapoyasense, Calaperostoma, 159,
162, 164, 282, 287.
chanchapoyasensis, Amphicyclotus, 162
chittyi, Poteria, 116, 121, 286.
chrysacme, Aperostoma, 203, 219, 281,
289.
Neocyclotus, 219.
cinereus, Cyclophorus, 141.
Ineerticyelus, 141, 287.
cingulatum, Aperostoma 221, 228, 229,
280, 289.
Cyclostoma, 228.
Poteria, 228.
cingulatus, Cyclotus, 228.
Neocyclotus, 228, 253.
elappi, Ptychocochlis, 82, 83, 91, 286.
clenchi, Farcimen, 24, 25, 284.
collare, Farcimen, 17, 18, 19, 283.
colombiense, Aperostoma, 223, 258, 280,
290.
colombiensis, Neocyclotus, 258.
Poteria, 258.
complanatum, Megalomastoma, 38.
confusum, Aperostoma, 221, 222, 237,
280, 281, 290.
Poteria, 237.
connivens, Aperostoma, 275.
Cyclotus, 275.
Incerticyclus, 275, 282, 291.
Neocyclotus, 275.
constrictus, Tomocyclus, 143, 145, 281,
287.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
cookei, Aperostoma 205, 215, 279, 281,
289.
cooperi, Amphicyclotus, 180.
Aperostoma, 180.
Cyclotus, 180.
Cyrtotoma, 180.
Mexcyclotus, 180, 181, 281, 288.
copanense, Cyclostoma, 147.
copanensis, Tomocyclus, 148, 147, 281,
287.
corpulentum, Aperostoma, 203, 217, 218,
279, 289.
corpulentus, Cyclotus, 217.
Neocyclotus, 217.
Poteria, 217.
corrugata, Ptychocochlis, 82, 83, 93, 95,
286.
corrugatior, Cyclotus, 84.
Neocyclotus, 84.
Ptychocochlis, 83, 84, 85, 86, 286.
corrugatissima, Poteria, 107. 110, 112,
114, 286.
corrugatissimus, Cyclotus, 110.
Neocyclotus, 110.
corrugatum, Aperostoma, 105.
Cyclostoma, 77, 82, 938, 105, 106,
119.
corrugatus, Cyclotus, 115, 119.
costaricense, Adelopoma, 52, 150, 280,
291.
Aperostoma, 221, 234, 280, 290.
costaricensis, Cyclotus, 234.
cousini, Amphicyclotus, 166.
Coen 159, 166, 167, 280,
288.
Cyclophorus, 166.
crassa, Poteria, 107, 113, 286.
‘| erassum, Cyclostoma, 113.
Farcimen, 15, 16, 283.
crassus, Cyclotus, 113, 115.
Neocyclotus, 113.
crocea, Cyclostoma, 45.
Helix, 44, 45.
croceum, Farcimen, 44, 45, 46, 285.
Crocidopoma, 39, 53, 62.
abbotti, 62, 67, 285.
casuelense, 62, 65, 291.
elevatum, 62, 63, 64, 285.
floccosum, 39, 62, 64, 65, 285.
gundlachi, 39, 41, 42, 284.
ignotum, 41, 42, 284.
milleri, 62, 66, 285.
orcutti, 62, 66, 285.
perdistinctum, 39, 40, 284.
toroense, 40, 284.
vortex, 62, 63, 65, 285.
wrighti, 41, 42, 284.
crosseanus, Amphicyclotus, 156.
Cyclophorus, 156.
Lagocyclus, 154, 155, 156, 280, 287.
cumingi, Aperostoma, 221, 224, 279, 289.
See ae 159, 167, 169, 279,
288.
Cyclophorus, 158.
cumingii, Amphicyclotus, 167.
Aperostoma, 167.
Cyclophorus, 167.
currani, Aperostoma, 2238, 261, 279, 290.
curtum, Farcimen, 3, 44, 46, 285.
Megalomastoma, 44
Cycladamsia, 124, 125.
bairdianum, 127.
deburghaeanum, 130.
fossile, 131.
humile, 129.
pretiosum, 127.
ruber, 126.
rudisplanusque, 131.
seabratum, 129.
seminudum, 130.
cycloata, Poteria, 109, 110, 286.
cycloatus, Cyclotus, 110.
Neocyclotus, 110.
Cyclobakeria, 106, 115
balnearis, 123.
ehittyi, 121.
dentistigmata, 120.
magister, 119.
nana, 120.
notatior, 122.
novaespei, 117.
taylori, 119.
tryoniana, 116.
welchi, 118.
yallahsensis, 123.
Cycloblandia, 54, 60.
amethystinus, 60.
beauianus, 61.
Cyclocaymania, 71, 79.
caymanensis, 80.
fonticula, 81.
laevitesta, 82.
Cyclocubana, 39.
gundlachi, 42.
ignotum, 42.
perdistinctum, 39.
toroense, 40.
wrighti, 42.
Cyclohaitia, 52, 53.
haitia, 53, 285.
Cyclohidalgoa, 124, 136, 187, 268.
bejumense, 270.
belli, 268.
dunoonense, 273.
haughti, 269.
major, 270.
pygmaeum, 272.
translucidum, 272.
trinitense, 136, 271.
Cyclojamaicia, 62, 67.
bondi, 68, 69, 70, 285.
suturalis, 68, 69, 70, 285.
Cyclophoridae, 3, 43, 187, 142.
Cyclophorus, 124.
acutiliratus, 56, 141.
antonii, 154.
bifasciata, 153.
boucardi, 184.
bourcieri, 163.
cayennensis, 276.
cinereus, 141.
cousini, 166.
erosseanus, 156.
cumingi, 158.
cumingii, 167.
INDEX
Cyclophorus delphinulus, 158.
disjunctus, 219.
dysoni, 207.
esmeraldensis, 160.
gayi, 278.
goldfussi, 184.
guayaquilensis, 161.
haematomma, 155.
hidalgoi, 167.
liratus, 58, 141.
lutescens, 180, 181.
maleri, 185.
martinezi, 152.
mexicanus, 173.
nigrofasciatus, 164.
orbignyi, 168.
ponderosus, 182.
psilomitus, 274.
rosenbergi, 160.
salleanus, 170.
texturatus, 186.
underwoodi, 176.
vasconesi, 156.
vesconesi, 156, 160.
volvulus, 124.
Cyclopilsbrya, 62, 71.
asperula, 71, 78, 285.
beswicki, 71, 79.
caribaea, 71, 77, 286.
caymanensis, 80, 285.
fonticula, 80, 81, 285, 291.
glenburniensis, 71, 73, 75, 285.
hendersoni, 71, 72, 285.
jugosa, 71, 76, 285, 291.
laevitesta, 80, 82, 285.
rufilabris, 71, 78, 79, 285.
rupisfontis, 71, 74, 285.
striosa, 71, 75, 76, 285.
westmorelandensis, 71, 72, 295.
Cyclopoma, 219.
disjunctum, 219.
Cyclopomops, 187, 219, 278.
moricandi, 219.
Cyclostoma, 41.
alutaceum,, 22.
antillarum, 49.
apertum, 26.
asperulum, 78.
auriculatum, 16, 25.
beauiana, 60, 61.
bicolor, 25.
bituberculatum, 15.
blanchetianum, 124, 244, 267.
bogotense, 226.
boucardi, 184.
bourcieri, 163.
brasiliense, 277.
cayennense, 276.
cingulatum, 228.
copanense, 147.
97
corrugatum, 77, 82, 93, 105, 106,
119.
crassum, 113.
erocea, 45.
cumingii, 167.
298
Cyclostoma disjunctum, 219.
distinetum, 275.
dubiosum, 70.
duffianum, 276.
dysoni, 208, 207.
flavidum, 45.
flavula, 45.
floccosum, 39, 62, 64.
gayi, 278.
giganteum, 124, 237, 238.
glaucostomum, 199.
grenadense, 134.
guatemalense, 144.
guayaquilense, 161.
idolum, 25.
inca, 267.
incomptum, 191, 192, 276.
inconspicuum, 227.
inornata, 61.
irregulare, 236.
jamaicense, 105, 106, 109, 112.
jugosum, 71, 76.
laxatum, 244.
lutescens, 181.
mani, 30.
martinicense, 140.
mexicanum, 124, 173.
moricandi, 220.
orbignyi, 46, 47.
pallescens, 111.
perpallidum, 139.
ponderosum, 182, 276.
popayana, 264.
prominula, 277.
psilomitum, 274.
purum, 162.
quitense, 250.
rufescens, 56.
schrammi, 57.
seminudum, 125, 126, 130.
simulacrum, 146.
solenatum, 25.
stramineum, 198, 276.
striata, 165.
striatum, 165.
subrugosum, 105.
suturale, 68, 69.
texturatum, 186, 276.
tortum, 4, 26, 38.
translucidum, 272.
varians, 86, 92, 93, 127, 128.
ventricosa, 30.
verruculosum, 51.
volvulus, 124.
vortex, 63.
Cyclotus, 41.
affinis, 210.
ambiguus, 211.
amethystinus, 60.
angulatus, 228.
asperulus, 78.
bairdianus, 127.
bartletti, 157.
berendti, 212.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Cyclotus beswicki, 79.
bisinuatus, 235.
bogotensis, 226.
boucardi, 175, 177.
cingulatus, 228.
connivens, 275.
cooperi, 180.
corpulentus, 217.
corrugatior, 84.
corrugatissimus, 110.
corrugatus, 115, 119.
costaricensis, 234.
erassus, 113, 115.
eycloatus, 110.
deburghaeanus, 130.
dentistigmatus, 120.
duflianus, 277.
dunkeri, 230.
dysoni, 208.
filo-liratus, 267.
fischeri, 239.
floccosus, 64.
gemma, 87.
giganteus, 238.
glaucostoma, 200.
glaucostomus, 199.
granulatus, 196.
inca, 244, 267.
incomptus, 189, 191.
inutilis, 107, 115.
irregularis, 236.
jamaicense, 123.
jamaicensis, 115.
jugosus, 76.
quitensis, 250.
laxatus, 244.
lineatus, 105, 109.
magna, 101.
mexicanus, 173.
minor, 100, 213.
multilineatus, 212.
nodosus, 107, 114.
notatior, 122, 123.
notatus, 107, 114.
novae-spei, 115, 117.
novussaltus, 112.
pallescens, 111.
parva, 76, 77.
pazi, 257.
perdistinctus, 39, 40.
perezi, 259.
perpallidus, 139.
pittieri, 236.
popayanus, 264.
portlandensis, 93, 94.
portoricensis, 59.
pretiosus, 127.
ruber, 126.
rudis-planusque, 131.
rufilabris, 78.
rugatus, 133.
rupisfontis, 74.
stramineus, 198.
striosus, 75.
INDEX
Cyclotus subrugosus, 105.
translucidus, 271, 272.
trinitensis, 136, 271.
varians, 86.
vortex, 63.
westmorelandensis, 72.
zigzag, 90.
Cyclovendreysia, 62, 69.
dubiosa, 70, 285.
eylindraceum, Megalomastoma, 45.
Cylindropalaina, 148.
Cyrtotoma, 151, 169.
avus, 169, 170, 281, 288.
cooperi, 180.
fischeri, 169, 170, 281, 288.
goldmani, 169, 173, 174, 281, 288.
ignotum, 169, 170, 281, 288.
mexicanum, 169, 173, 174, 281,
palmeri, 169, 172, 281, 288.
salleanum, 169, 170, 281, 288.
walkeri, 169, 174, 281, 288.
deburghaeanum, Aperostoma, 129,
287.
deburghaeanus, Cyclotus, 130.
delphinulus, Amphicyclotus, 158.
Cyclophorus, 158.
Filocyclus, 158, 282, 287.
dentistigmata, Poteria, 116,
291.
dentistigmatus, Cyclotus, 120.
Neocyclotus, 120.
depressum, Aperostoma, 222,
290.
depressus, Neocyclotus, 247.
Poteria, 247.
digitale, Farcimen, 22, 23, 283.
Megalomastoma, 23.
Diplommatina occidentale, 52.
occidentalis, 52
stolli, 149.
Diplommatininae, 43, 51, 142, 148.
disjunctum, Cyclopoma, 219.
Cyclostoma, 219.
disjunctus, Cyclophorus, 219.
distinctum, Cyclostoma, 275.
distinctus, Incerticyclus, 275, 291.
Neocyclotus, 275.
domingoense, Farcimoides, 46, 47, 284.
Te ti Amphicyclotulus, 54, 57,
285.
dubiosa, Cyclovendreysia, 70, 285.
dubiosum, Cyclostoma, 70.
duffianum, Aperostoma, 276.
Cyclostoma, 276.
duffianus, Cyclotus, 277.
Incerticyclus, 276, 282.
Neocyclotus, 277.
288.
130,
120,
247,
dunkeri, Aperostoma, 221, 230, 233, 280,
289. 3
Cyclotus, 230.
Neocyclotus, 230.
Poteria, 230, 256.
269, 273, 279,
dunoonense, Aperostoma,
289
dyeri, Aperstoma, 204, 205, 281, 289.
299
dysoni, Aperstoma, 203, 204, 207, 209,
217, 281, 289.
Cyclophorus, 207.
Cyclostoma, 203, 207.
Cyclotus, 208.
Neoeyclotus, 208, 209.
Pliatystoma, 208.
Poteria, 208.
ecuadorense, Aperostoma, 222, 248, 250,
251, 280, 290.
elephantinum, Farcimen, 7, 9, 283.
elevatum, Crocidopoma, 62, 63, 64, 285.
esmeraldense, Calaperostoma, 159, 160,
161, 286, 287.
esmeraldensis, Amphicyclotus, 160.
Cyclophorus, 160.
Eupalaina, 148.
exiguum, Aperostoma, 221, 234, 280, 289.
Farcimen, 1, 3, 4, 5, 34, 48, 45, 46.
alutaceum, 4, 15, 21, 22, 23, 24, 283.
anafense, 27, 28, 284.
apertum, 5, 26.
arangoi, 36, 284.
auriculatum, 24, 25, 26, 284.
balneorum, 31, 32, 284.
bayamense, 5, 6, 283.
biayaense, 12, 13, 14, 283.
bicolor, 24, 25, 284.
bituberculatum, 13, 14, 15, 16, 283.
camagueyanum, 10, 11, 283.
catalinense, 36, 284.
clenchi, 24, 25, 284.
collare, 17, 18, 19, 283.
crassum, 15, 16, 283.
croceum, 44, 45, 46, 285.
curtum, 3, 44, 46, 285.
digitale, 22, 23, 283.
elephantinum, 7, 9, 283.
flavulum, 45.
fiorencianum, 11, 12, 283.
giganteum, 15, 16, 283.
guanense, 37, 284.
guantanamense, 7, 19, 283.
guitarti, 17, 19, 283.
gundlachi, 27, 28, 284.
gundlachiellum, 27, 28, 284.
gutierrezi, 15, 16, 17, 283.
hendersoni, 35, 36, 284.
hjalmersoni, 44, 45, 285.
holguinense, 7, 8, 283.
itinerarium, 35, 284.
lagunillense, 37, 38, 284.
leonellum, 29, 284.
leoni, 17, 19, 283.
leoninum, 29, 284.
magister, 17, 19, 288.
magnificum, 22, 283.
mani, 30, 284.
martianum, 13, 14, 283.
mayajiguense, 11, 12, 283.
mayariense, 7, 8, 10, 283.
mayense, 7, 10, 283.
minor, 15, 16, 283.
mirandum, 7, 9, 288.
multistriatinum, 31, 32.
300
Farcimen najazaense, 13, 14, 283.
obesum, 20, 21, 283.
perconvexum, 7, 8, 283.
poeyi, 17, 18, 19, 283.
procer, 38, 284.
pseudotortum, 5, 6, 7, 26, 283.
rocai, 23 284.
scopulorum, 4, 33, 284.
seminudum, 17, 19, 283.
semivestitum, 7, 8, 283.
senectum, 24, 284.
Subobesum, 20, 21, 283.
subventricosum, 31, 32, 284.
superbum, 34, 35, 284.
torrei, 20, 284.
tortum, 4, 5, 26, 284.
turquinoense, 5, 6, 283.
ungula, 5, 7, 9, 10, 283.
ventricosum, 30, 32, 284.
vinalense, 32, 33, 284.
wrighti, 12, 13, 283.
yunquense, 6, 283.
Farcimoides, 3, 43, 46.
domingoense, 46, 47, 284.
orbignyi, 46, 47, 284.
Sallei, 46, 285.
fusciata, Neocyclotus, 227, 256.
Poteria, 256.
fasciatum, Aperostoma, 223, 256, 257,
282, 290.
Filocyeclus, 151, 157.
delphinulus, 158, 282, 287.
filoliratum, Aperostoma, 221, 267, 280,
291.
filo-liratus, Cyclotus, 267.
Poteria, 267.
fischeri, Aperostoma, 222, 239, 280, 290.
Cyclotus, 239.
Cyrtotoma, 169, 170, 281, 288.
Poteria, 239.
fischerianus, Neocyclotus, 258.
flavidum, Cyclostoma, 45.
flavidus, Turbo, 45.
flavula, Cyclostoma, 45.
Megalomastoma, 44, 45.
flavulum, Farcimen, 45.
floccosum, Crocidopoma, 39, 62, 64, 65
282.
?
’
Cyclostoma, 39, 62, 64.
floccosus, Cyclotus, 64.
florencianum, Farcimen, 11, 12, 283.
fonticula, Cyclopilsbrya, 80, 81, 285, 291.
fonticulus, Neocyclotus, 81.
fossile, Aperostoma, 125, 131, 287.
fultoni, Aperostoma, 222, 242, 279, 290.
gayi, Amphicyclotus, 278.
Cyclophorus, 278.
Cyclostoma, 278.
Incerticyclus, 278, 279.
gealei, Megalomastoma, 143.
Tomocyclus, 142, 143, 147, 281, 287.
gemma, Cyclotus, 87.
Neocyclotus, 87.
Ptychocochlis, 88, 87, 285.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
gigantea, Poteria, 238.
giganteum, Aperostoma, 221, 222, 237,
238, 239, 277, 281, 290.
Cyclostoma, 124, 237, 238.
Farcimen, 15, 16, 283.
giganteus, Cyclotus, 238.
Neocyclotus, 2388, 239.
glabra, Helix, 45.
glaucostoma, Cyclotus, 200.
Poteria, 200.
glaucostomum, Aperostoma, 196,
282, 288.
Cyclostoma, 199.
glaucostomus, Cyclotus, 199.
Neocyclotus, 200.
glenburniensis, Cyclopilsbrya, 71, 73, 75.
285
199,
goldfussi, Amphicyclotus, 184, 281.
Aperostoma, 184. i
Cyclophorus, 184.
goldmani, Cyrtotoma, 169, 173, 174, 281,
288.
gossei, Ptychocochlis, 83, 85, 286.
gracilius, Megalomastoma, 147, 148.
granulatum, Aperostoma, 196, 280, 288.
granulatus, Cyclotus, 196.
Neocyclotus, 196.
Poteria, 196.
grenadense, Aperostoma, 133, 134, 135,
287.
Cyclostoma, 134.
guadeloupensis, Amphicyclotulus, 54, 55,
58, 60, 285.
guanense, Farcimen, 37, 284.
guantanamense, Farcimen, 7, 10, 283.
guatemalense, Cyclostoma, 144.
Megalomastoma, 144.
guatemalensis, Tomocyclus, 143, 144,
281, 287.
guayaquilense, Calaperostoma, 159, 161,
280, 287.
Cyclostoma, 161.
guayaquilensis, Amphicyclotus, 161.
Cyclophorus, 161.
guitarti, Farcimen, 17, 19, 283.
gundlachi, Crocidopoma, 39, 41, 42, 284.
Farcimen, 27, 28, 284.
Megalomastoma, 28.
gundlachiellum, Farcimen, 27, 28, 284.
gutierrezi, Farcimen, 15, 16, 17, 283.
Habropoma lutescens, 181.
mexicanum, 171, 173.
salleanum, 170.
haematomma, Amphicyclotus, 155.
Cyclophorus, 155.
Lagocyclus, 154, 155, 156, 280, 287,
291.
haitia, Cyclohaitia, 53, 285.
haughti, Aperostoma, 268, 269, 279, 289.
hedui, Aperostoma, 187, 191, 279, 288.
Helix crocea, 44, 45.
glabra, 45.
hendersoni, Cyclopilsbrya, 71, 72, 285.
Farcimen, 35, 36, 284.
Ptychocochlis, 83, 96, 286.
INDEX
hidalgoi, Amphicyclotus, 167.
Calaperostoma, 159, 166, 167, 279,
287.
Cyclophorus, 167.
hinkleyi, Aperostoma, 203, 204, 206, 207,
281, 289.
hitomi, Aperostoma, 187, 194, 280, 288.
hjalmersoni, Farcimen, 44, 45, 285.
Megalomastoma, 45.
bolguinense, Farcimen, 7, 8, 283.
humile, Aperostoma, 129, 287.
idolum, Cyclostoma, 25.
ignotum, Crocidopoma, 41, 42, 284.
Cyrtotoma, 169, 170, 281, 288.
{mitator, Poteria, 107, 108, 286.
{nea, Aperostoma, 221, 267, 279, 291.
Cyclostoma, 267.
Cyclotus, 244, 267.
Neocyclotus, 245, 267.
Poteria, 267.
Incerticyclus, 137, 275.
bakeri, 137 188, 139, 287.
bowdenensis, 138, 139, 291.
braziliensis, 277, 279, 291.
cayennensis, 276, 281.
cinereus, 141, 287.
connivens, 275, 282, 291.
distinctus, 275, 291.
duffianus, 276, 282.
gayi, 278, 279.
martinicensis, 140.
perpallidus, 139, 287.
perplexus, 140, 287.
prominulus, 277, 279, 291.
schermoi, 138, 291.
Incidostoma, 187.
hedui, 191.
hitomi, 194.
incomptum, 192.
kobelti, 190.
malleatum, 188.
nirafe, 193.
pergrandis, 188.
pichinchense, 191.
pizarroi, 193.
stirlingi, 195.
incomptum, Aperostoma, 187, 192, 282,
288
Cyclostoma, 191, 192, 276.
incomptus, Cyclotus, 191.
Neocyclotus, 192.
Poteria, 192.
Pterocyclos, 192.
inconspicuum, Aperostoma, 221, 227, 282,
- 289,
Cyclostoma, 227.
Poteria, 227.
inornata, Cyclostoma, 61.
inutilis, Cyclotus, 107, 115.
Neocyclotus, 115.
Poteria, 115.
irregulare, Aperostoma, 222, 236, 280,
290.
Cyclostoma, 236.
Poteria, 236.
428349—42—_20
301
irregularis, Cyclotus, 236.
Neocyclotus, 236.
itinerarium, Farcimen, 35, 284.
jamaicense, Aperostoma, 105.
Cyclostoma, 105, 106, 109, 112.
Cyclotus, 123.
jamaicensis, Cyclotus, 115.
Neocyclotus, 82.
Poteria, 106, 107, 112, 286.
Turbo, 82, 93, 105, 106, 112.
jugosa, Cyclopilsbrya, 71, 76, 285, 291.
jugosum, Cyclostoma, 71, 76.
jugosus, Cyclotus, 76.
Neocyclotus, 76.
knobbei, Neocyclotus, 101.
kobelti, Aperostoma, 187, 190, 279, 288.
kugleri, Aperostoma, 196, 201, 282, 289.
lacteofluviale, Aperostoma, 98.
lacteofluvialis, Ptychocochlis, 84, 98, 286.
laevitesta, Cyclopilsbrya, 80, 82, 285.
Poteria, 82.
Lagocyclus, 151, 154, 159.
antoni, 154, 156, 280, 287.
bartletti, 154, 157, 282, 287.
erosseanus, 154, 155, 156, 280, 287.
haematomma, 154, 155, 156, 280, 287,
291.
vasconesi, 154, 156, 280, 287.
lagunillense, Farcimen, 37, 38, 284.
laxatum, Aperostoma, 222, 244, 280, 290.
Cyclostoma, 244.
Poteria, 244.
laxatus, Cyclotus, 244.
Neocyclotus, 244.
leai, Aperostoma, 221, 222, 246, 282, 290.
Calaperostoma, 159, 165, 279, 288.
leonellum, Farcimen, 29, 284.
leoni, Farcimen, 17, 19, 283.
leoninum, Farcimen, 29, 284.
Megalomastoma, 29.
limellum, Aperostoma, 196, 197, 199, 201,
279, 288.
limensis, Pupa, 148.
lineata, Poteria, 106, 107, 109, 110, 115,
286, 291.
lineatus, Cyclotus, 105, 109.
Liracyclotus, 186, 274.
psilomitus, 274, 282, 287.
liratus, Amphicyclotulus, 54, 58, 60, 285.
Cyclophorus, 58, 141.
Lomastoma, 48.
lutescens, Amphicyclotus, 181.
Aperostoma, 181.
Cyclophorus, 180, 181.
Cyclostoma, 181.
Habropoma, 181.
Mexcyclotus, 179, 180, 181, 288.
magister, Farcimen, 17, 19, 283.
Poteria, 116, 119, 287.
magna, Cyclotus, 101.
Ptychocochlis, 84, 101, 286.
magnificum, Farcimen, 22, 283.
major, Aperostoma, 269, 270, 279, 289.
Poteria, 270.
302
maleri, Amphicyclotus, 184, 185, 220,
281, 288.
Cyclophorus, 185.
malleatum, Aperostoma, 187, 188, 189,
190, 191, 282, 288.
manabense, Aperostoma, 222, 239, 280,
290.
manchesterensis, Ptychocochlis, 83, 88,
285.
mani, Cyclostoma, 30.
Fareimen, 30, 284.
Megalomastoma, 30.
marianna, Ptychocochlis, 83, 97, 286.
martensi, Neocyclotus, 95.
Ptychocochlis, 82, 83, 95, 286, 291.
martianum, Farcimen, 13, 14, 283.
martinezi, Buckleya, 152.
Buckleyia, 151, 152, 154, 280, 287.
Cyclophorus, 152.
martinicense, Cyclostoma, 140.
martinicensis, Incerticyelus, 140.
masvense, Aperostoma, 223, 254, 280,
290.
mayajiguense, Farcimen, 11, 12, 283.
mayariense, Farcimen, 7, 8, 10, 283.
mayense, Farcimen, 7, 10, 283.
mcsweeni, Aperostoma, 132, 134,
287.
Megacyclotus, 151, 181.
palenquensis, 182, 183, 281, 288.
ponderosus, 182, 183, 281, 288.
Megaloma, 48.
Megalomastoma, 3, 41, 48, 48, 50, 51.
antillarum, 49, 50, 284.
apertum, 26, 27.
brunnea, 48, 49.
brunneum, 48, 49, 284, 285.
complanatum, 88.
curtum, 44,
eyclindraceum, 45.
digitale, 23.
flavula, 44, 45.
gealei, 143.
gracilius, 147, 148.
guatemalense, 144.
gundlachi, 28.
hjalmersoni, 45.
leoninum, 28.
mani, 30.
minor, 16.
minus, 147, 148.
mirandum, 9.
petiti, 48, 49, 50, 284.
procer, 38.
seminudum, 7, 19.
simulacrum, 146.
torrei, 20.
tortum, 5, 6.
ungula, 9.
verruculosum, 51, 284.
Megalomastominae, 3, 43, 142.
Megalomastomoides, 3, 48. 50.
verruculosum, 51.
merrilli. Aperostoma, 223, 262, 263, 264,
279, 290.
135,
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Mexcyclotus, 151, 179.
cooperi, 180, 181, 281, 288.
lutescens, 179, 180, 181, 288.
mexicanum, Aperostoma, 173.
Cyclostoma, 124, 173.
Cyrtotoma, 169, 173, 174, 281, 288.
Habropoma, 171, 173.
mexicanus, Cyclophorus, 173.
Cyclotus, 173.
milleri, Crocidopoma, 62, 66, 285.
mineri, Amphicyclotulus, 54, 55, 58, 60.
285.
minor, Cyclotus, 100, 213.
Farcimen, 15, 16, 283.
Megalomastoma, 16.
Neocyclotus, 260.
Poteria, 213.
Ptychocochlis, 84, 100, 286.
minus, Megalomastoma, 147, 148.
mirandum, Farcimen, 7, 9, 283.
Megalomastoma, 9.
montegoensis, Ptychocochlis,
286.
montezumi, Aperostoma, 151, 152.
moricandi, Amphicyclotus, 220.
Aperostoma, 219, 279, 291.
Cyclostoma, 220.
multilineatus, Cyelotus, 212.
multistriatinum, Farcimen, 31, 32, 284.
najazaense, Farcimen, 13, 14, 283.
nana, Poteria, 116, 120, 286.
nanum, Aperostoma, 223, 262, 263, 282,
290
84, 101,
Neocyclotus, 125, 187, 203.
affinis, 210.
ambiguus, 211.
asperulus, 78.
bakeri, 137.
bartletti, 157.
belli, 201, 268.
berenadti, 212.
beswicki, 79.
bisinuatus, 235.
bogotensis, 226.
boueardi, 177.
caucaensis, 258.
chrysacme, 219.
cingulatus, 228, 253.
colombiensis, 258.
connivens, 275.
corpulentus, 217.
corrugatior, 84.
corrugatissimus, 110.
crassus, 113.
eycloatus, 110.
dentistigmatus, 120.
depressus, 247,
distinctus, 275.
duffianus, 277.
dunkeri, 230.
dysoni, 208, 209.
fasciata, 227, 256.
fischerianus, 2:
fonticulus, 81.
gemma, 8&7.
INDEX
Neocyclotus giganteus, 238, 239.
glaucostomus, 200.
granulatus, 196.
inea, 245, 267.
incomptus, 192.
inutilis, 115.
irregularis, 236.
jamaicensis, 82.
jugosus, 76.
knobbei, 101.
laxatus, 244.
martensi, 95.
minor, 260.
nodosus, 114.
notatior, 122.
notatus, 114.
novae-spei, 117.
pallescens, 111.
panamensis, 197.
pazi, 257.
peilei, 200.
perezi, 259.
pergrandis, 188, 190.
perpallidus, 139.
peruvianus, 252.
popayanus, 264.
quitensis, 250.
rupisfontis, 74.
stramineus, 198.
subcingulatus, 249.
subrugosus, 105.
thielei, 72.
translucidus, 272.
trinitensis, 271.
varians, 8&6.
westmorelandensis, 72.
zigzag, 90.
Neopupina, 3, 5, 43.
eroceum, 45.
eurtum, 44.
hjalmersoni, 45.
nevadense, Aperostoma, 223, 255, 282,
290.
nicaraguense, Aperostoma, 205, 214, 216,
281, 289.
nigrofasciatum, Calaperostoma,
164, 166, 280, 288.
nigrofasciatus, Amphicyclotus, 164.
Cyclophorus, 164.
nirafe, Aperostoma, 187, 193, 195, 279,
288.
nodosa, Poteria, i14.
nodosus, Cyclotus, 107, 114.
Neocyclotus, 114.
notata, Poteria, 114.
notatior, Cyclotus, 122, 123.
Neocyclotus, 122.
Poteria, 116, 122, 123, 286.
notatus, Cyclotus, 107, 114.
Neocyclotus, 114.
novae-spei, Cyclotus, 115, 117.
Neocyclotus, 117.
Poteria, 115, 116, 117, 2&6.
novussaltus, Cyclotus, 112.
obesum, Farcimen, 20, 21, 283.
159,
303
occidentale, Adelopoma, 52, 148, 149,
150, 151, 282, 291.
Diplommatina, 52.
occidentalis, Diplommatina, 52.
Palaina, 52.
olivaceum, Aperostoma, 222, 251, 252,
280, 290.
olssoni, Amphicyclotus, 178.
Calacyclotus, 178, 179, 279, 281, 288.
orbignyi, Amphicyclotus, 168.
Calaperostoma, 159, 168, 279.
Cyclophorus, 168.
Cyclostoma, 46, 47.
Farcimoides, 46, 47, 284.
orcutti, Crocidopoma, 62, 66, 285.
Ptychocochlis, 84, 99, 286.
paezense, Aprostoma, 221, 225, 279, 289.
ae Aperostoma, 221, 232, 280,
90.
pailaense, Aperostoma, 221, 231, 280,
Palaina occidentalis, 52.
stolli, 149.
palenquense, Aperostoma, 183.
palenquensis, Megacyclotus,
281, 288.
pallescens, Cyclostoma, 111.
Cyclotus, 111.
Neoecyclotus, 111.
Poteria, 107, 111, 286.
palmeri, Cyrtotoma, 169, 172, 281, 288.
panamense, Aperostoma, 196, 197, 281,
288.
panamensis, Neocyclotus, 197.
Poteria, 197.
parva, Cyclotus, 76, 77.
pazi, Aperostoma, 217, 257, 280, 290.
Cyclotus, 257.
Neocyelotus, 257.
Poteria, 257.
peilei, Aperostoma, 196, 200, 279, 288.
Neocyelotus, 200.
perconvexum, Farcimen, 78, 283.
perdistinctum, Aperostoma, 276.
Crocidopoma, 39, 40, 284.
perdistinctus, Cyclotus, 59, 40.
perezi, Aperostoma, 223, 259, 280, 290.
Cyclotus, 259.
Neocyclotus, 259.
Poteria, 259.
pergrandis, Aperostoma, 187, 188, 190,
279, 288.
Neoeyclotus, 188, 190.
Poteria, 188.
perpallidum, Cyclostoma, 139.
perpallidus, Cyclostoma, 139.
Cvclotus, 1389.
Incerticyclus, 139, 287.
Neocyclotus, 139.
perplexus, Amphicyclotulus, 54, 59, 285.
Incerticyclus, 140, 287.
peruense, Aperostoma, 222, 245, 247, 282,
290
182, 183,
peruvianum, Aperostoma, 223, 252, 282,
290.
304
peruvianus, Neocyclotus, 252.
Poteria, 252.
petiti, Megalomastoma, 48, 49, 50, 284.
petricola, Ptychocochlis, 104, 286.
pichinchense, Aperostoma, 187, 191, 280,
288
pisecinalis, Valvata, 141.
pittieri, Aperostoma, 222, 236, 280.
Calaperostoma, 159, 164, 281, 287.
Cyclotus, 236.
Poteria, 236.
pizarroi, Aperostoma, 187, 193, 194, 195,
282, 288.
plana, Poteria, 107, 113, 114, 286.
Platyraphe, 220.
Platystoma, 82.
dysoni, 208.
Plectocyclotus, 106.
poeyi, Farcimen, 17, 18, 19, 283.
ponderosum, Cyclostoma, 182, 276.
ponderosus, Amphicyclotus, 182.
Cyclophorus, 182.
Megacyclotus, 182, 183, 281, 288.
popayana, Cyclostoma, 264.
Poteria, 256, 264.
popayanum, Aperostoma, 223, 264, 266,
280, 291.
popayanus, Cyclotus, 264.
Neocyclotus, 264.
portlandensis, Cyclotus, 93, 94.
portobellense, Aperostoma, 222, 242, 281,
290.
portoricensis, Amphicyclotulus, 54, 59,
285.
Cyclotus, 59.
Poteria, 62, 82, 105, 106.
affinis, 210.
ambiguus, 211.
aulari, 199.
balnearis, 116, 123, 286.
bartletti, 157.
bejumensis, 270.
berendti, 212.
blanchetianum, 245.
bogotensis, 226.
bondi, 68.
burringtoni, 107, 286.
campeachyi, 104.
cardozi, 266.
caribaea, 77.
caucaensis, 258.
caymanensis, 80.
chittyi, 116, 121, 286.
cingulatum, 228.
colombiensis, 258.
confusum, 237.
corpulentus, 217.
corrugatissima, 107, 110, 111, 112,
114, 286.
crassa, 107, 1138, 286.
cycloata, 109, 110, 286.
dentistigmata, 116, 120, 287, 291.
depressus, 247.
dunkeri, 230, 256.
dysoni, 208.
fasciata, 256.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
Poteria filo-liratus, 267.
fischeri, 239.
gigantea, 238.
glaucostoma, 200.
granulatus, 196.
imitator, 107, 108, 286.
inca, 267.
incomptus, 192.
inconspicuum, 227.
inutilis, 115.
irregulare, 236.
jamaicensis, 106, 107, 112, 286.
laevitesta, 82.
laxatum, 244.
lineata, 106, 107, 109, 110, 115, 286,
291.
magister, 116, 119, 287.
major, 270.
minor, 213.
nana, 116, 120, 286.
nodosa, 114,
notata, 114.
notatior, 116, 122, 123, 286.
novaespei, 115, 116, 117, 286.
pallescens, 107, 111, 286.
panamensis, 197.
pazi, 257.
perezi, 259.
pergrandis, 188.
peruvianus, 252.
pittieri, 236.
plana, 107, 113, 114, 286.
popayana, 256, 264.
quitensis, 250.
rugata, 202.
sanctaemarthae, 218.
santaguitensis, 272, 2738.
smithi, 216.
stramineus, 198.
subcingulatum, 249.
taylori, 118, 119, 287.
translucida, 272.
trinitensis, 186, 271.
tryoniana, 116, 286.
vincentiana, 133.
welchi, 116, 118, 119, 286
yallahsensis, 116, 123, 287.
pretiosum, Aperostoma, 126, 127, 287.
pretiosus, Cyclotus, 127.
princeps, Aperostoma, 175.
Barbacyclus, 175, 280, 288.
procer, Farcimen, 38, 284.
Megalomastoma, 38.
prominula, Cyclostoma,. 277.
prominulus, Incerticyclus, 277, 279, 291.
pseudotortum, Farcimen, 5, 6, 7, 26, 283.
psilomitum, Cyclostoma, 274.
psilomitus, Amphicyclotus, 274.
Cyclophorus, 274.
Liracyclotus, 274, 282, 287.
Pterocycios, 161.
incomptus, 192.
Ptychocochlis, 62, 82.
adamsi, 83, 94, 95, 286.
campeachyi, 84, 103, 104, 286.
clappi, 82, 83, 91, 286.
corrugata, 82, 83, 93, 95, 286.
INDEX
305
Ptychocochlis corrugatior, 83, 84, 85, 86, | salleanum, Aperostoma, 170.
286
gemma, 83, 87, 285.
gossei, 83, 85, 286.
hendersoni, 83, 96, 286.
lacteofluvialis, 84, 98, 286.
magna, 84, 101, 286.
manchesterensis, 83, 88, 285.
marianna, 838, 97, 286.
martensi, 82, 83, 95, 286, 291.
minor, 84, 100, 286.
montegoensis, 84, 101, 286.
orcutti, 84, 99, 286.
petricola, 104, 286.
savannensis, 82, 84, 98, 286.
senex, 83, 92, 286.
shawae, 83, 90. 286.
simpsoni, 83, 95, 286.
subglobosa, 84, 103, 286.
subrugosa, 83, 105, 286.
taylori, 83, 89, 286.
varians, 83, 86, 286.
vendreysi, 82, 84, 102, 286.
welchi, 83, 88, 285.
zigzag, 83, 90, 286.
pulchellum, Aperostoma, 223, 263,
290.
Pupa limensis, 148.
Pupoides, 148.
purum, Calaperostoma, 159, 162,
279,
280,
Cyclostoma, 162.
pygmaeum, Aperostoma, 269, 272,
289.
quitense, Aperostoma, 222, 250, 280, 290.
Cyclostoma, 250.
quitensis, Cyclotus, 250.
Neocyclotus, 250.
Poteria, 250.
redfieldi, Aperostoma, 223, 261, 279, 290.
rocai, Farcimen, 23, 284.
rosenbergi, Amphicyclotus, 160.
282,
Calaperostoma, 159, 160, 280, 287.
Cyclophorus, 160.
ruatanense, Aperostoma, 204, 207, 281,
289.
ruber, Aperostoma, 125, 126, 127, 128,
287.
Cyclotus, 126.
rudisplanusque, Aperostoma, 125, 131,
287.
radis-planusque, Cyclotus, 131.
eee, Amphicyclotulus, 54, 56, 57,
285.
Cyclostoma, 56.
rufilabris, Cyclopilsbrya, 71, 78, 79, 285.
Cyclotus, 78.
rugata, Poteria, 202.
rugatum, Aperostoma, 133, 286.
rugatus, Cyclotus, 133.
rupisfontis, Cyclopilsbrya, 71, 74, 285.
Cyclotus, 74.
Neocyclotus, 74.
salengoense, Aperostoma, 223, 253, 280,
290.
Cyrtotoma, 169, 170, 281, 288.
Habropoma, 170.
salleanus, Cyclophorus, 170.
sallei, Aperostoma, 205, 213, 281, 289.
Farcimoides, 46, 285.
sanctaemarthae, Aperostoma, 203, 217,
218, 279, 289.
Poteria, 218.
santaguitense, Aperostoma, 289.
santaguitensis, Poteria, 272, 273.
See Ptychocoechlis, 82, 84, 98,
286.
scabratum, Aperostoma, 125, 129, 287.
schermoi, Incerticyclus, 138, 291.
een Amphicyclotulus, 54, 57, 58,
Cyclostoma, 57.
scopulorum, Farcimen, 4, 33, 284.
seminudum, Aperostoma, 109, 125, 128,
129, 130, 131, 287, 291.
Cyclostoma, 125, 126, 130.
Farcimen, 17, 19, 283.
Megalomastoma, 7, 19.
seminudus, Cyclostoma, 1380.
semivestitum, Farcimen, 7, 8, 283.
senectum, Farcimen, 24, 284.
senex, Ptychocochlis, 83, 92, 286.
shawae, Ptychocochlis, 83, 90, 286.
simile, Aperostoma, 221, 227, 280, 289.
simpsoni, Ptychocochlis, 83, 95, 286.
simulacrum, Cyclostoma, 146.
Megalomastoma, 146.
Tomocyeclus, 142, 148, 144, 146, 147,
281, 287.
siphonis, Tomocyclus, 143, 145, 146, 281.
smithi, Aperostoma, 203, 204, 216, 217,
279, 289.
Poteria, 216.
solenatum, Cyclostoma, 25.
stirlingi, Aperostoma, 187, 195, 280, 288.
stolli, Adelopoma, 149, 281, 291.
Diplommatina, 149.
Palaina, 149.
stramineum, Aperostoma, 132, 195, 196,
198, 282, 288.
Cyclostoma, 198, 276.
stramineus, Cyclotus, 198.
Neocyclotus, 198.
Poteria, 198.
striata, Cyclostoma, 165.
striatum, Cyclostoma, 165.
striosa, Cyclopilsbrya, 71, 75, 76, 285.
striosus, Cyclotus, 75.
subcingulatum, Aperostoma, 222, 249,
251, 280, 290.
Poteria, 249.
subcingulatus, Neocyclotus, 249.
subglobosa, Ptychocochlis, 84, 103, 286.
subobesum, Farcimen, 20, 21, 283.
subrugosa, Ptychocochlis, 83, 105, 286.
subrugosum, Cyclostoma, 105.
subrugosus, Cyclotus, 105.
Neocyclotus, 105.
subventricosum, Farcimen, 31, 32, 284.
306
sumichrasti, Aperostoma, 204, 209, 211,
281, 289.
superbum, Farcimen, 34, 35, 284.
suturale, Cyclostoma, 68, 69.
suturalis, Cyclojamaicia, 68, 69, 70, 285.
taylori, Poteria, 118, 119, 287.
Ptychocochlis, 83, 89, 286.
texturatum, Cyclostoma, 186, 276.
texturatus, Amphicyclotus, 184, 186, 281,
288
Cyclophorus, 186.
thielei, Neocyclotus, 72.
Tomocyclus, 8, 142.
constrictus, 143, 145, 281, 287.
copanensis, 148, 147, 281, 287.
gealei, 142, 143, 147, 281, 287.
guatemalensis, 148, 144, 281, 287.
simulacrum, 142, 148, 144, 146, 147,
281, 287.
siphonis, 143, 145, 146, 281.
toroense, Crocidopoma, 40, 284.
torrei, Farcimen, 20, 284.
Megalomastoma, 20.
tortum, Cyclostoma, 4, 26, 38.
Farcimen, 4, 5, 26, 284.
Megalomastoma, 5, 6.
tortus, Turbo, 4, 5, 26.
translucida, Poteria, 272.
translucidum, Aperostoma, 136, 179, 268,
269, 272, 279, 289.
Cyclostoma, 272.
translucidus, Cyclotus, 271, 272.
Neocyclotus, 272.
trinitense, Aperostoma, 136, 268, 269,
271, 287, 289.
trinitensis, Cyclotus, 186, 271.
Neocyclotus, 271.
Poteria, 136, 271.
truncatum, Cerion, 45.
tryoniana, Poteria, 116, 286.
tryonianum, Aperostoma, 116.
tuema, Adelopoma, 148, 279, 291.
Turbo flavidus, 45.
jamaicensis, 82, 93, 105, 106, 112.
tortus, 4, 5, 26.
turquinoense, Farcimen, 5, 6, 283.
umbilicatum, Aperostoma, 221, 224, 225,
226, 279, 289.
Underwoodi, Amphicyclotus, 176.
Barbacyclus, 175, 176, 178, 280, 288.
BULLETIN 181, UNITED STATES NATIONAL MUSEUM
underwoodi, Cyclophorus, 176.
ungula, Farcimen, 5, 7, 9, 10, 283.
Megalomastoma, 9.
utriaense, Aperostoma, 222, 240, 241,
280, 290.
valerioi, Aperostoma, 205, 213, 215, 280,
289.
Valvata piscinalis, 141.
varians, Cyclostoma, 86, 92, 93, 127, 128.
Cyclotus, 86.
Neocyclotus, 86.
Ptychocochlis, 83, 86, 286.
vasconesi, Cyclophorus, 156.
Lagocyclus, 154, 156, 280, 287.
vendreysi, Ptychocochlis, 82, 84, 102, 286.
venezuelense, Aperostoma 222, 247, 248
282, 290.
ventricosa, Cyclostoma, 30.
ventricosum, Farcimen, 30, 32, 284.
veracochanum, Aperostoma, 221, 229,
282, 289.
verruculosum, Cyclostoma, 51.
Megalomastoma, 51, 284.
vesconesi, Amphicyclotus, 156.
Cyclophorus, 156, 160.
vinalense, Farcimen, 32, 33, 284.
vineentina, Poteria, 133.
vinecentinum, Aperostoma, 133, 286.
volvulus, Cyclophorus, 124.
Cyclostoma, 124.
vortex, Crocidopoma, 62, 63, 65, 285.
Cyclostoma, 63.
Cyclotus, 63.
walkeri, Aperostoma, 174.
Cyrtotoma, 169, 174, 281, 288.
welchi, Poteria, 116, 118, 119, 286.
Ptychocochlis, 83, 88, 285.
westmorelandensis, Cyclopilsbrya, 71,
72, 295.
Cyclotus, 72.
Neocyclotus, 72.
wetmorei, Aperostoma, 203, 279, 291.
wrighti, Crocidopoma, 41, 42, 284.
Farcimen, 12, 13, 283.
yallahsensis, Poteria, 116, 123, 287.
yunquense, Farcimen, 6, 283.
zigzag, Cyclotus, 90.
Neocyclotus, 90.
Ptychocochlis, 88, 90, 286.
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