3 coli’ SO Sne ; ot a) NO ISS ZAR A. A Ar NNO ON RRS oa s Oia Ass ass AN, \) RRL XX Ving . \\ \ eh \\ ; i in a Wat: ake } been act UR eat m4 mies WAT) die Aen ae me Pat. ind é 7 a : Ve a Bae | hak oo ry ¥ me Bitty my ey ' had Ay im in 4 ie SMITHSONIAN INSTITUTION UNITED STATES NATIONAL MUSEUM BuL.teETin 182 MONOGRAPH OF THE WEST INDIAN BEETLES OF THE FAMILY STAPHYLINIDAE BY RICHARD E. BLACKWELDER UNITED STATES GOVERNMENT PRINTING OFFICE WASHINGTON : 1943 For sale by the Superintendent of Documents, Washington, D.C. - - - - - - - -« = - = Price $1.00 ADVERTISEMENT The scientific publications of the National Museum include two series, known, respectively, as Proceedings and Bulletin. The Proceedings series, begun in 1878, is intended primarily as a medium for the publication of original papers, based on the collec- tions of the National Museum, that set forth newly acquired facts in biology, anthropology, and geology, with descriptions of new forms and revisions of limited groups. Copies of each paper, in pamphlet form, are distributed as published to libraries and scien- tific organizations and to specialists and others interested in the different subjects. The dates at which these separate papers are published are recorded in the table of contents of each of the volumes. The series of Bulletins, the first of which was issued in 1875, con- tains separate publications comprising monographs of large zoologi- cal groups and other general systematic treatises (occasionally in several volumes), faunal works, reports of expeditions, catalogs of type specimens, special collections, and other material of similar nature. The majority of the volumes are octavo in size, but a quarto size has been adopted in a few instances in which large plates were regarded as indispensable. In the Bulletin series appear volumes under the heading Contributions from the United States National Herbarium, in octavo form, published by the National Museum since 1902, which contain papers relating to the botanical collections of the Museum. The present work forms No. 182 of the Bulletin series. ALEXANDER WETMORE, Assistant Secretary, Smithsonian Institution. CONTENTS Page RCDOCUCTIONM a Soe ae See te re a se ators Sen este tek UC petssteay) 1 IA CEN OW LEG PVE Mt Sea eras eee AEE Ste Af Bet, PS 2 GINO T A Taye ee ee ee i ee ef ns ST AN Sherry 4 Bunpematic irertment...o.. 802555 oh oS Sau eee eee ek a) 30 Method of treatment ce sees Se esigues allel fee ost ee. | 30 Haunalirelationships= <2] 262 Ste SS Le me Bee late h erceal) 36 BOULCEs OL TECOTOR. ae as eae aes 5 Sos A et ref 38 Mermimnolapy = 22 5 Steer A het eal ote aye Ded Pel seek 2 = 39 Arrangement of subfamilies and genera_______.._----_------------ 40 Key to the subfamilies of West Indian Staphylinidae_____________- 40 Subfamily Pins Tin aes ee See ee eee easter eee kl etree Ed | 41 Key to West Indian genera of Piestinae____.._....___...-_----- 41 Genus Hy potelus Pirichson S22 es ssh a eit: bee 8) ee 41 Gens Prestus Gravenhorst..2...-<-....ctasisoouet gle als 43 Key to West Indian species of Piestus._.........-..--.---- 44 Bubpramily Oseerron are Syl pe. eaters, ehh) de Wt see 5I Key to West Indian genera of Omaliinae___-_-._--------.----- 51 Gentiselhiocono0 mus WCCl 3 25 ma et RE) oe ee. 51 Genus: Onalium- Gravenhorst.s!s: #2 ").2) 4s2 8 28-ee =.=. 53 Key to West Indian species of Omaliwm____..------------ 54 Buplamily/ Ox VTe bias 8 a eee SP pn eet. 57 Key to West Indian genera of Oxytelinae_______.._..______--- 57 Genus: Carneizmis samonelle=.- «2. s3t4 aot Joe oe ee KS 58 Key to West Indian species of Carpelimus__._------------ 59 Genuz Lorrentomus Bierig.sh.28 52. genase eed. 220k ek. - 87 Key to West Indian species of Torrentomus__...---------- 87 Genus-A pocelliisEirichsonan tes ao ere dy ke Tk 88 Key to West Indian species of Apocellus_._._...._..__-.----- 89 Genus Oxiytelue Gravenhorst’< = =s242--2)521~h2279- 42-32 3 - = 5 = 91 Key to West Indian species of Oxytelus__._._.....--------- 92 Geutisiw aross Siar pete tn oy ee. ok ee a 103 Key to West Indian species of Parosus__......----------- 103 Genus Hiimovruedweson wetter sob. cette oh oe t= 105 Key to West Indian species of Thinobius........-..------ 105 Genus Platystethus Mannerheimes = ..22212. 1 2088/48 ae 2 109 CeO DNIS Dale CUaH DO aMMOUC) Gece oY PE a a bald og ABE ae i 112 Key to West Indian species of Bledius._.___._....--------- 113 MULARINEO OOMORIINA eee Bee ee ee Poe Ba ae th FS 119 Key to West Indian genera of Osoriinae___....-.-.-.--------- 119 Genus: Psetdolispinodes Bernhauer! 2. 2.22 n eee ee 120 Key to West Indian species of Pseudolispinodes___-_-.----- 121 Genus ErapinsanWriCUBOlis 52 20 ees lie ot eg hE do et 129 Key to West Indian species of Lispinus._.._...------------ 130 IV BULLETIN 182, UNITED STATES NATIONAL MUSEUM Systematic treatment—Continued. Page Subfamily Osor11naE—Continued. Genus Inopeplus Smith... 222225 2 ese eee e cate senses 138 Key to West Indian species of Inopeplus_-_..------_---_-- 139 GenusiGlyptoma, ErichSone sss. e ae eee ee 141 Key to West Indian species of Glyptoma____________------ 142 Genus Jspeson Schautucs 2.22" 822 ok es cee ee aes 144 Key to West Indian species of Espeson___---------------- 144 Genus}; 2horacophonus Motschulsky--_2__-_ = = 222 ee 148 Key to West Indian species of Thoracophorus__.____._----- 149 Genussileusis laporte..22 .86oo. 3 ees sense. ees ee 155 Genus. Paralispinus Bernhauers_-- 3. Se ee 156 Key to West Indian species of Paralispinus__.._...------- 157 Genus Lepiochirus Germar.. 22.2252 .-22..0 24sec es 162 WNeotrochus: New: PEWS! = = a a So a ea AY ee 164 Key to West Indian species of Neotrochus__.._-_--.-------- 165 GenuszHoloirochus richson 222 22 16h ss Se eee ae 169 Key to West Indian species of Holotrochus___-_.--_------- 170 GGenuseMamogonussbauviel == 22245252 oo See ee ee a 172 Genus Osorus. Latreillessse 2 i ee ee Sie ee ee 174 Key to West Indian species of Osortus__..._-.------------ 175 Sublamily SrvnOropINAwe : 2s ok OE ee ee ee 202 Genus: Siylopodits Beniek..4 2-23 eet eee Pes ee 202 Key to West Indian species of Stylopodus____________- 203 Subfamily; SueNinaws 2.95340. eee eae: ls OR eee te eS 208 GenuseStenusWatreilles. 2 Se, Ee ee ee 208 Key to West Indian species of Stenus___.-.._._---____- 209 Subfamily. HUAMSTHETINADY! <3. 38 Sees Saeko eee eee eee 226 Genus Hvoctavius Bierioe 22 se ee ee 297 Genus. 2 amotius- Schautusse= === 2) 28 tse ee ee 2907 Genus. Stencesthetus Sharps 2225-28 eee eee 228 Subfamily PampoRINAmcGL 18 tee eu: ee ee ee ee 228 Key to West Indian genera of Paederinae________--------_--- 229 Genus “Ehinochars Wraatz=. 2) See ee ee eee a Se 231 Key to West Indian species of Thinocharis_____.---------- 232 Genus Lithocharis Boisduval and Lacordaire________________-_- 239 Key to West Indian species of Lithocharis__.__...--__-___ 240 Genus.Aderocharis Sharply 32 220020 eee: ere ee 250 Key to West Indian species of Aderocharis___.___.-___-__. _- 251 Genus: Stilomedon: Sharp. <8. 2008 Sens Wee ee ee ee 255 Key to West Indian species of Stilomedon___---------.__-- 256 Genus Sunius Stephens! 2. 2 2220s Bets ee Oe eee ee 259 Key to West Indian species of Sunitus_....._....___.._--__- 261 Genus Iledon: Stephensge.c 22st ee ee 270 Key to West Indian species of Medon__------------------ 270 Genus Orus' Casey oo: cece eee ee ee ee Did Key to West Indian species of Orus______-__.-_--2._2--_- 277 Genus Scopacie Prichsons<> 2200" Ta See ee eee 279 Key to West Indian species of Scopaeus________..--__---- 280 Genus-Monisia Sharps 2325 3s oe ee ee ee eee 296 Key to West Indian species of Monista___._....-..._____- 296 Genus Rugilus Samouelle_________________________---_ eee 299 Key to West Indian species of Rugilus._...........------. 300 CONTENTS V Systematic treatment—Continued. Page Subfamily PanpertnaE—Continued. Genus Scopobium : Blackwelder_- =... a 306 Genus: Lathrobium Gravenhorst:.... =< —.—e e 307 Key to West Indian species of Lathrobium__._.____-_______ 308 Genus Lobrathium Mulsant and Rey__--_-----------_---_-_--_ 311 Key to West Indian species of Lobrathium__.___...______- 312 Genur:PrredemiscPaprieissc 2 te ee i Mt 321 Key to West Indian species of Paederus..-__--___-_---_-- 322 Genus, Homocotarsus Hochhuthy 3 36 et eo ee 325 Key to West Indian species of Homoeotarsus___-__-_--_- ~~ 326 Genus. Ochthephtlum. Stephens... ..--2<-2b. oo $:<< set on no 9. 331 Key to West Indian species of Ochthephilum___------_---- 332 Gann: Bigerypla Casey: 4428s soo ae eth ee ne 5 335 Key to West Indian species of Biocrypta__.._._..._-----_-- 336 Gens nites TiC bSony Oe ery Pee 8 ee AE Ek 343 mUmsOBnacte Mow i PChMss 2 <= 2 at eked eae Ged eee Se 345 Key to West Indian species of Suniophacis__..._____------ 345 REPO SOURS SNOW ODS EE Se nok eae ee Eagle VATS 348 Key to West Indian species of Stilosaurus__..-_-.__------ 348 Genusssrannoderis Sharp. | 22. =. aetna Ae See h 351 Key to West Indian species of Stamnoderus____.______---- 351 CemisvSiniphaers: Bierig8 0 he. 2 te Dope Guth g 356 Key to West Indian species of Stiliphacis___._..___...------ 356 GenussSunrosaunusibieriy ss a eee ae ee tO eek 361 GeRGsSUntOCRaris Sharples -~-isees seeibelpe o2 ae Fs ee 362 Roneiie New. fens. 5. 222.25 c nnn Ree eee elests 364 GenyisjAstenus StephenszeetaeF te Snacee cette te oes ees 365 Key to West Indian species of Astenus___---------------- 366 Genus, Echiaster Brichson_....< 44+. --teet aacsaeeeeds canes. 369 Key to West Indian species of Echiaster_____-_----------- 369 Genus \Sphaeronum Sharp... .e4elisletol escent 374 Genus: Pinophilus, Gravenborst... ..- 2492! sseecatsh ees. 376 Key to West Indian species of Pinophilus__-.-.-__-------- 377 Genus Aracocerus, Nordmann.__........ 2r“iem score lt yiietele 386 Genus, Palommus Wrichson.....5.2.2...520 tL ewieeesth a0. 388 Suplamily STAPHYERININAB 2.2.0. ..5. WA inenaous te: glimatd 393 Key. to tribes. of Staphylininae_._._. . ssed std cages ane. 394 Key to West Indian genera of Staphylinini____------_--_----- 394 Genus: Pasderamimus. Sharp... sie) s2uweseet hve. 395 Key to West Indian species of Paederomimus___-.-----~---- 395 Genus: Philonthus Stephens... duaM sucoseolS oe. . 398 Key to West Indian species of Philonthus____..-_-_------- 400 Genus. Belonuchus Nordmann.......--22292J.eberevl! suuL..- 420 Key to West Indian species of Belonuchus_-_---.--.------ 421 Gena Cafus, Stephens... sole SEAS Sates oe) 435 Key to West Indian species of Cafius............--------- 436 Genus. Frichsontus Fauyel_... 2325S LS Le 440 Genus: Stanhylinus Linnacus.........02016 wu ousaenisouiel.. 443 Genus) Creaphiius Samouelle...1sbubouie lf on2nigeinne uuLe_. 447 Genus: Xanthopygus Krastz.. 2911 tun doses ayweet) ulre.. 449 Genus) Phelothalpus Kraate in. ene 2 RU) eo uat AL... 451 Key to West Indian species of Philothalpus__.....-----. 452 VI BULLETIN 182, UNITED STATES NATIONAL MUSEUM Systematic treatment—Continued. Page Subfamiily SrapHyLinrnazE—Continued. Genus Drochus sHrichsone2= 2202s. seen se see oe ece sae soe a 455 Genus: Cephaloxynum-Bernhauer "2272 eee eee 458 Key to West Indian species of Cephalorynum__-.---------- 459 Genustiolsustirichsontes ee ee aes, Soe eee eee eee 460 Key to West Indian species’ of Holsws so 2-24. 2--.=-== 461 Genus-Heterothops Stephens=s0s 25. eee eee ee ee 464 Key to West Indian species of Heterothops_.__-.---------- 464 GenussAlcylophorus Nordmann 222222 Vs bas Tee ee eee 466 Key to West Indian species of Acylophorus__.------------ 467 Genus Atanygnathius:JacobsONe=2 2522822 — = eee oe eee ee eee 471 Key to West Indian species of Atanygnathus_....--------- 471 Key to West Indian genera of Xantholinini._.._.____---_------ 473 GeénuseXaentholinus*Dejeans 1022 Be eg ee EL 473 Key to West Indian species of Xantholinus__--.---------- 475 Genus DThyneocephalus Guérinss ssssss20 bees 228 eee 490 Genus-PlochionocerustSharps.- 202 35 Jes A ee eee 491 Genus-Leptacinus-Erichsoms:. soe ee ee ee 493 Key to West Indian species of Leptactnus___..___----_---- 493 Genus: Lithocharodes Sharp. <. << 225. 20ve os on Be ese 496 Key to West Indian species of Lithocharodes_.._-.---_-_-- 496 Genus-Iicrolinus:-Casey.22.022 5233532 ea i ae. 502 Genus: Stenolunus*Bierig © 322k Sue see De Ot ye ae 503 Genus. Oligolinius Casey =. o2sssscssact ea eee ee 504 Key to West Indian species of Oligolinus..........--_---- 505 Subfamily (ACHYPORINAM 3554 52454as¢ oe22 so Oe ee ee 509 Key to West Indian genera of Tachyporinae______._________-- 509 Genus: Leucoparyphus Kraata2 bo 555s 2% 0 Le oe da ce 510 Genus: Coproporus: Kraatzs 22.53 bss, Serene) Sie 512 Key to West Indian species of Coproporus........-------- 512 Genus Conosomus:-Motschulsky 2225228) 3 eee 524 Genus Bryoporus. Kraatzs<... 522 ee ee lad eked Fee 526 Key to West Indian species of Bryoporus_..---.---------- 527 Subfamily. HyPocyPHTINnAb 442222 sheet ei ee 53 Genus-Andey pis Hommess- 2 5c eee 3 eee 532 Subfamily ABOCHARINAB so co.ceosee se ee oe 533 Genus. Deinopsis Matthews. aaah eS Sele Beko ae en 534 Genus: Myllaena Brichsonztsee. 1s saasot oe ee ek eee 534 Genus; Pronomaea.Trichson.. ...- cased 2l eee see See 535 Genus: Giigota, Mannerheim_ ba eso 2 ace led_ et ae 535 Genus Thyreonenus. Manns... 2. ee eee cee 538 Genus EburniolasMannt) )- oeoee cet tee od eo - Woe 538 Genus: Thazteria Fenyes...... .. eon cee e_ 539 Genus: Perinthus ‘Casey aiai o> 22sscs ethol el ot ger Le 539 Genus. Barychara: Sharp...-..202+ 2. 2edsete i pee 539 Genus (Gnypetosoma, Cameron: 22 scene3t 0 set et ee 539 Genus, Alisalia Casey. 2.2... >. JeesleG avtenels ct ae 540 Genus Brachychara Sharp... ts o--el Fouad ol pee 540 Genus Gyrophaena.Mannerheim. 20) 4sseu seissss ene. 540 Genus; Diesiota Mulsant and Rey- i 2tee=2)..seece Sect ae 544 Genus. Coenonica, Kraatz... 22.5. ete a ee ee pee 544 Genus Placusa Erichson Genus Sclusa Prichsons. 52 ssee ono ee eee eee 545 CONTENTS Systematic treatment—Continued. Subfamily ALEocHARINAE—Continued. Genuseanecturellas Camerones = 2222 2 ee once eas ses scke Genus: Bolttochara; Mannerheim==-=..-...-22-2s2.-t-s--5---<- Genusextenovtota bicrig# Aes _ ee? toate ee A eee se Genus: Huvindg SURED .s eu beet Se SORE nn oS ees secs Genus falagre® Samoucles 222 5. 6 28k Sy Oe es Genus cronerg: OUST pss 52 Se eae ee See eee GeHUSUINEOLGTG DUS 2. Rie te oo ne eee Genus Pseudognypeta Cameron®—-— 2-2 -- 2 noe ee sees GenusiGnypeta- Thomson s=as2sese2sre 2 sa cela te Be Genus-Aitheia sbhomson-==2-<2sece=s22-— St Pw eee Genus. Alevonoia Phomson=.225225 52 2-222 2S Bees Gems: Guajira Dienes. 2-2 foes Soh e ee eet oe ce ee Genus Mimacrotona Cameron! ===. .....~--------.--=---+.--- Genus Pseudothamiaraea Cameron....----------------------- Gems yras Htephensa-. fo. Sie Mn ee ek ee Genus, “enotoma, Camenrone os. 2550 estes te eters Genus anlandsia WKranty see ee ee Genus: Phlocopora;brichsone =22. 225 22555 See ae ee (Couns icalea, eriChsO0 = 2 5-- 2 eek wee oe eS Genus ermiogaster Casey 21). x2 hee el ee Sat ae. Genus -Termiophya Wasmann=. 22 325..52254202. 2 es GenusehuinorecisOlerens os ofee sess eo eee ee Se ee Geniis Ocjusa WIRRtt i Se et ee a Genus Aleochara Gravenhorst. =. 2. 2252. - 655 sse5e Stee RP aCOMIs bie Cavallo eee tee ee ee oe poe ca ee ee eee Pelleriineiscabiane:- £25. 2)sc we ees Ul see pete ols eae Pee iO onan as tek 2 ee Oe ess ee eee eae eee son. See PATCLGLGNN GUT ht er a Ee eS = ewes VII Page 546 546 546 546 547 548 548 548 549 549 555 555 555 555 556 556 557 557 558 558 558 559 559 559 562 585 603 624 625 ad — SOMNONRWNHE ll OWMDNANTEWHe Hispaniola.o<2. 522.26 Puerto Ricoz-_.--2-2= » Virgin Islands. -2...=- Ot: Martin.and St. Kitts jeroups. <<. 2.22 2045.2 Sse err Be } Antiguaw=- 2.22822. m Montserfate.2.2225=- »» Guadeloupe.........- & Dominica ss sss55 2 ek y Martinique: 2.22.22 SoS be UCla =m ae ff Barbadoss. 222 42-522. sols-VINCentes— = uae HiGrenadinesi2s222 42-22 PEGTreN AG ae oso = arr Vill ILLUSTRATIONS FIGURES MONOGRAPH OF THE WEST INDIAN BEETLES OF THE FAMILY STAPHYLINIDAE By Ricwarp E. BuackweELDrr INTRODUCTION For many years the West Indian Islands have been a place of unusual interest to American naturalists and those of many other countries as well. The list of entomologists who have studied the insects of the West Indies is long, and several of our large museums have sponsored such studies. The Smithsonian Institution, through the Walter Rathbone Bacon Scholarship and other means, has con- ducted several investigations of the fauna of the islands, but until the present study was undertaken none had been concerned with the insects of that region. The Walter Rathbone Bacon Scholarship is administered by the Smithsonian Institution for the study of the fauna of the regions outside of the United States, and in 1935 it was awarded to me for the purpose of studying the Staphylinidae of the West Indies. The program of study that was followed involved the collecting of Staphylinidae on each of the islands from Cuba to Trinidad; the examination of all local collections; the study of types in the British Museum and the collection of Dr. Malcolm Cameron; the mounting, labeling, and sorting of the collections assembled; the identification of the species; and the preparation of a report, to in- clude a systematic revision of the family as it occurs in the West Indies. In pursuance of this plan, I spent 21 months in the islands collecting specimens and data, a month in London studying types, and 14 months at the United States National Museum preparing and studying the collections. In addition, some time was spent on the manuscript during 1938-40, and after my return to the Na- tional Museum in July 1940, this paper occupied most of my time for 12 months. The number of specimens of Staphylinidae collected was between 45,000 and 50,000. In addition to these, approximately 8,800 speci- mens from the West Indies were examined in the British Museum, over 400 in Dr. Cameron’s collection, several hundred in the Ameri- can Museum of Natural History, over 1,600 from the Museum of 1 2 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Comparative Zoology, and about 1,500 in the United States Na- tional Museum. Several smaller collections were also studied, yield- ing valuable additions to the series. These are listed in a later para- graph. For comparisons and the drawing up of keys and diagnoses many specimens from other parts of the New World were studied. These were principally in the United States National Museum but were partly from the collections mentioned above. The study has included 91 genera (6 of them new) and 468 spe- cies (exclusive of the Aleocharinae). Of these, 829 species were rep- resented by type material in some of the collections examined. New species (which were also included in the above 329) numbered 212. We collected at least 179 species in the West Indies, including nearly half of the 212 new species. Only 34 species were not represented in material examined by me, and about half of these were in genera that were not studied in detail (Palaminus in particular). ACKNOWLEDGMENTS This study was made possible by the Smithsonian Institution through the Walter Rathbone Bacon Scholarship and the facilities of the United States National Museum. It was carried out with the assistance of Dr. Edward A. Chapin, curator of the division of insects, without whose constant advice and encouragement it could never have been completed in the time available. It would have been impossible to make positive identifications of many species, or to have included many others at all, without the cooperation of the British Museum (Natural History). G. J. Ar- row, curator of Coleoptera, kindly made it possible for me to study the numerous types in the collections of the Museum and was of spe- cial help in arranging for the loan of a considerable number of speci- mens. This enabled me to compare them directly with the series at the United States National Museum and to draw up descriptions for which there was not time during the brief stay in England. Dr. W.R. Thompson, of the Imperial Institute of Entomology, was also of much help to me during my studies at the British Museum. Many species of West Indian Staphylinidae have been described by Dr. Malcolm Cameron, who lives in a suburb of London. , Yo S. thy voy) e oy OV0770S » SODINAINFNI lee, Qe SONY 7S! SYWYHYa SINI/d 4O 77S YNYAVH 13 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 96 SOE PbE O0e see vee £86 200 166 OLE OLE B96 L9E 196 O96 tee DO 2 OP 927 260 068 606 yo" PEC LEE O08 yw Ne G OCF 627 IGE 2p a S96 . e HLYE TY INOIHINYN " PIE Sw WN OHINID (A . “ao, ® 996 2279 NOLSONIM O/NOLNY LU0d £9¢ MOLITLSYD oot AVE OLIONNY VAVNWET ree IN aN 2 enogYyvH G70 Sip Sr rer s LL NHOL HSINVES nae 0 vip 8 oe H or Z 696 ul SLE qm bY INOW INOW OLE ow sty ele 12 ele Ley. Ava SNNY LS clr ONOd YOLYIITIV ‘OlP Vip eer Oo JITVUAIONVH L6€ 6 86 /07 oor HLOOW TS YAY MIVTG 120 cra 0ePr a or S/ FAWNOD v7 wayN LS VN WS ‘ ve bor ‘ ODVILNYS ® ge al SZAIUNO OTS “0 ( OP st \ MONOGRAPH OF WEST INDIAN STAPHYLINIDAE L7 os ANGULLA ‘ ST MARTIN oe? 0 ST. BARTHOLOMEW ST MARTIN & ST KITTS GROUPS S ST. EUSTATIUS BASSE TERRE ST. KITTS ‘ NEVIS Map No. 7. BULLETIN 182, UNITED STATES NATIONAL MUSEUM 18 °8 “ON dvIT 262 eNogerH : HS/TONF ° NMOL GvOw O70 HLNOW TVS v2 SLNIVS 77 a ce WrHe Yd aie SONVISI FAl¥ 9 $L2 av 82 v6Z snnor ss $62, £62 Ts 262. = ate VIDLLNY | a} MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 19 cane oat MONTSERRAT WOOOLANDS s TRANTS BAY 263 PLYMOUTH > 271 270 262 O GARRAS @ Map No. 9. BULLETIN 182, UNITED STATES NATIONAL MUSEUM 20 ‘OT=ONeshN Ose SILNIVS S77 SLANT 79 Fern SIOFAE S104L FPYFLISSVG 239 tl & 09 PL py ere FVYFIL FLILIFd <> o> JYYFL ISSVG 06 Id-¥- FLNIOD FYION FLNIOS Fu PLS se tiast SIOINVES LS FYYFZL JONVYO MILNIAVT JINOW 37 JOvHISIO & SIND] L40d FIMOTIGMNI MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 21 DOMINICA 260 ) PORTSMOUTH LONDONDERRY M. DIABLOTIN BASSINVILLE a ST. JOSEPH 259 240 TROIS PITONS GRAND BAY Map No. ll. 22 BULLETIN 182, UNITED STATES NATIONAL MUSEUM MARTINIQUE M PELEE RST PIERRE LA TRIN/TE ty Go v LAMENTIN FORT DE FRANCE ° ve oe © FRANCOIS a A e MARIN °° Ge bf et Map No. 12. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 23 ST LUCIA °¢ GROS /I§LET 2/3 2// ANSE LA RAYE £97 208 eM 2809 2/5 " 2/7 CANARIES 2/6 2/8 2/9 M. GIMIE SOUFRIERE ANSE CHOISEUL LABORIE VIEUX FORT Map No. 13. 24 BULLETIN 182, UNITED STATES NATIONAL MUSEUM BARBADOS e 191 SPRING HALL e ST. Lucrs 19 ST. ANOREWS CHURCH a 196 SPEIGHTS TOWN 203 200 202 CANE GARDEN a HOLETOWN F 187 BATHSHEBA 194 BRIDGETOWN sag 190 CHRIST CHURCH s Mar No. 14. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 20 ST VINCENT lé4 LA SOUFRIERE 183 182 164 ORY w# CHATEAU BLLA/R “EORGETOWN i. 173 BUCCAMENT R 166 176 I77 169 W71 165 ‘a r 179 YAMBU R 70 /67 KINGSTOWN I72 /8/ (63 185 CALLIAQUA ee °9 Map No. 15. 449008—42——-3 26 BULLETIN 182, UNITED STATES NATIONAL MUSEUM ST VINCENT GRENADINES BEQUIA ; oa BATTOWIA Q Pe ae te QUATRE ISLE BALLICEAUX MUS TIQUE if Ss oO L/TTLE MARTINIQUE CARRIACOU DIAMOND ROCK de ISLE RONDE GRENADA Map No. 16. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Dl SAUTEUVAS 197 Lanroine 16l /60 GOUYAVE 135 GRENVILLE 451 GRAND ETANG o ST GEORGES Mar No. 17. BULLETIN 182, UNITED STATES NATIONAL MUSEUM “28 (SORT ANIA A ENE NS ER TS TIE A FOISATAS SLON AYN, yi NYUNY? Bg / 121 ar ee 62/ HONOSOEYHIS ee? ed HLNOWATA 6i/ HINOSOBXOY 29 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE '6TSON SAVIN [ae eee RR VONYOW OY VAS ai a el Wt vIva v7 $01, OONYNYI 4 NYS O87 Oe # LVUYISLNON lg O&A OGHEHS FAIYN GYGINEL ald VITNVYZNYN JONVYD FUONKS a SNVHS INDY? ai ee 2Z/ 4 921 InILSNONY aS ee ® Of $21 eel a i Sie Sit PB Hd ISOF LS oe ol NivdS 40 LYOd /0/ AVG YYONY Ira 80/ 42N0S SNYS a 8 SYSTEMATIC TREATMENT METHOD OF TREATMENT It has been my endeavor throughout this monograph to present in uniform style all the information known for each species. Thus I have included keys to genera and species, complete synonymy, a standardized description, the exact type locality, the disposition of all types, all known records, the number and locations of all speci- mens I have examined, notes regarding the relationships and other items of interest, and what is known of the habits of the species. What my intention has been in each of these phases is indicated in the following paragraphs. In the case of genera it has not been pos- sible to give as exhaustive a treatment. The synonymy with original citations, the genotypes of all names, a diagnosis of the important characters, and notes on relationships are given in each case. All keys given are original and are frequently based on characters that have not been commonly recorded in the past. Many of these are structural characters described in a previous study of the mor- phology of the Staphylinidae (Blackwelder, 1936). No attempt whatsoever has been made to make the keys natural or to make them present a regular classification. They are considered to be purely artificial aids to the identification of the species, although in some cases the resulting arrangement may coincide with a natural group- ing. For my conception of the arrangement of the groups one should consult the Systematic Catalog (p. 562). In the generic synonymy all names are listed that have been used as genera or subgenera and are now considered synonyms or sub- genera. The latter are treated as synonyms except for the indi- cation of their usual status. Wherever possible, misspellings and nomina nuda are included since there is always some chance that further information will change their status. The generic synon- ymy is based principally on two sources: The Junk Coleopterorum Catalogus (Bernhauer, Schubert, and Scheerpeltz, 1910 to 1934) and new synonyms discovered during the present study. The new synonymy is not indicated as such in the list but is usually discussed in the “Remarks” at the end of the diagnosis. Exceptions to the above are the subfamily Paederinae and the tribe Lispinini. In these all the synonymy is based on new studies of the genotypes, undertaken as necessary sidelines of the present study and recently 30 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE ol published as generic revisions of the groups (Blackwelder, 1939a and 1942). It is obviously impossible to make identifications of species unless the genus is definitely known. The genus depends largely upon its genotype for its definitive characters, and it is therefore very im- portant to have designated genotypes for all the generic names. Very few designations have been made in this family, even by stu- dents who have made careful studies of genera and subgenera. Such studies may lose much of their value if their writers’ conception of the genus differed from the conception of the original describer of the genus. Our rules of procedure compel us to go back to the origi- nal conception, and this should be established as the cornerstone of the foundation of each generic revision. I have, therefore, listed genotypes for all names, although this has involved the designation of a great many. The International Rules of Zoological Nomen- clature have been scrupulously adhered to, and all doubtful cases are discussed in footnotes. It is possible that previous designations have been overlooked in some cases, but careful search has been made and every original description examined. An indication is always given as to the source of the designation, whether it is a monobasic genus or designation was made originally, subsequently, or in the present work. The generic diagnoses form, in my estimation, one of the impor- tant contributions of this study. It is very surprising to find that few of our several thousand named genera of Staphylinidae have ever been carefully described. Original publications far too often base a genus simply upon a certain species, or upon a single differ- ence between it and a related genus. In trying to remedy this I have assembled a diagnosis containing the characters that appear to be of generic value in that subfamily, giving them in a uniform man- ner for comparison. These were almost always drawn from actual specimens, wherever possible from the genotype. Some characters used in these descriptions have not been commonly used in the past. Explanations of them will be found under “Terminology” or in my study of the morphology (Blackwelder, 1936). It is believed that these characters will be found to hold for most of the species of the genus in any part of the world. Frequently several species have been examined, but it is, of course, impossible to be certain without exam- ining all the species of each genus. They will hold, I believe, for all the American species. The keys to species are often based in part upon species known to me only by descriptions. Although this has forced me to rely in part on the characters (frequently superficial) that are to be found o2 BULLETIN 182, UNITED STATES NATIONAL MUSEUM in the descriptions, it seems worth while in enabling one to have a complete picture of the genus without having to consider a series of “species inquirendae.” If these keys be found to be faulty, it will seldom affect the species I had before me but only the doubtful ones. Before each specific name in the key is a number indicating the position of that species in the sequence of the text. Very careful attention has been given to the specific synonymies. Although the Junk catalog has again been used as a starting point, a large amount of synonymy has been found that is not recorded in that already voluminous work. In addition, each species is re- corded under each generic name with which it has ever been coupled. In general, I have tried to cite all references to each species, but in the case of cosmopolitan species this is obviously impracticable. In such cases, only the original reference is given. The date and page refer to the detailed citations in the bibliography. Throughout the work the descriptions have been arranged on a uniform pattern. The color and markings are given first; then the details of the head, pronotum, elytra, and abdomen, with their ap- pendages, in that order; the special characters of the male and the female; the special characters of the legs; and finally the length. Throughout each genus I have tried to mention the same characters in the same order and manner to facilitate comparisons. Certain terms and usages are explained in a later paragraph. There is a strong and highly commendable tendency among mod- ern entomologists to give exact dimensions in the descriptions of insects. Doubtless these are of great value in many groups. How- ever, in the Staphylinidae there is almost always a considerable range of size due to at least two factors: (1) The actual difference in size of individuals; and (2) the apparent difference caused by greater or less retraction of the flexible abdomen. A very exact measurement of total length is of little use if the specific range is great. Since in the present work direct measurements have not been found to be of great value, the length is given in millimeters and major fractions thereof, and all minor measurements are given as proportions of one part to another. These are expressed as frac- tions rather than as ratios because they have thus been found to be more readily understandable and comparable. Several factors have prevented the use of primary genitalic charac- ters in the descriptions. The first of these was the lack of time to develop techniques and make the numerous dissections and prepara- tions. Another was the fact recently demonstrated in other studies that some genera do not present usable specific characters on the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 30 genitalia. And so little has been done on this phase of the study of the Staphylinidae that there is not a proper foundation on which to start such a study. This is not to be construed as a declaration that these structures may not in many cases prove to be of great value in classification. The sexual characters of the seventh, eighth, and ninth segments have been recorded wherever possible. They are fre- quently of high value in specific distinctions, and they generally enable one to distinguish the sexes. The type or original locality of a species is frequently of interest in cases in which two closely related species are confused and in other situations. It has been found, however, that there are several conceptions of the meaning of the term “type locality.” As used herein the type locality is the actual place at which the holotype was taken, whether this exact spot is now known or not. However, spe- cific type localities have seldom been given for West Indian species, and it has been necessary to content oneself in many cases with merely the name of the island. In every case, however, the locality is given with as much exactness as possible. In certain species it has been found that the published locality does not agree with that on the type specimen. These cases are individually discussed in the text. The present location of type specimens and material from the type series is of considerable interest to persons desiring to make an intensive study of a species or genus. Our accepted procedure nowadays demands the recording of the type depository, but in the past such record was more often omitted than not. It has there- fore been thought advisable to record the disposition of types wher- ever possible. In the case of all new species the types have been carefully designated and labeled, and catalog numbers have been assigned to those in the United States National Museum. The types in the British Museum (principally of Sharp’s and Cameron’s spe- cies) are in part holotypes and in part undesignated lectotypes (syntypes), but they have all been treated uniformly as holotypes. The same is true of the types in Dr. Cameron’s collection. Many of the species, however, were described long ago by Erichson, Fauvel, or others without designation of or record of disposition of types. In these cases I have indicated the probable location of the type, based on the location of the collection as recorded by W. Horn in his work “Uber entomologische Sammlungen” (Berlin-Dahlem, 1935-37). In order to give a complete outline of the distribution of each species, I have listed all the records I have been able to find, whether 34 BULLETIN 182, UNITED STATES NATIONAL MUSEUM previously published or represented by specimens in collections or new ones from my own collecting. These are arranged geographi- cally in the following order: (1) The West Indies from the Bahamas Islands through the Greater Antilles and the Lesser Antilles to Trinidad (and Curacao); (2) South, Central, and North America; (3) Europe, Africa, and the Orient. In the case of large islands, such as Cuba, Jamaica, Hispaniola, Puerto Rico, and Trinidad, spe- cific localities are listed when known; but with the smaller islands this has seemed unnecessary. In all cases of my own records, how- ever, reference is given to the collecting stations listed hereinafter (p. 584), where detailed localities can be obtained. For continental American and for Old World records only the country (or State in the United States) is given, with the source of the record. The rec- ords for Guadeloupe have generally been divided between Basse Terre and Grande Terre. (See map No. 10). These are virtually separate islands and are very different in topography and vegeta- tion. Records from Hispaniola have been separated as from Haiti and the Dominican Republic. This is simply a matter of con- venience. There frequently arise situations when another student is attempt- ing to use a work of this sort, in which he needs to know what specimens and how many were before the describer or reviser. He will also want to know if the types have been examined. I have therefore given for each species the number and location of the speci- mens that I have examined, with specific mention of the types if they were studied. Other data of interest not included in the previous sections are mentioned under “Remarks.” Comparisons with similar species are often given, and mention of the subgenus is made if it has been determined. And what is known of the habits or occurrence of the species is given in a separate paragraph. Certain exceptions to this general plan should be noted. The sub- families Euaesthetinae and Aleocharinae and the genera Palaminus and Stenolinus are not treated in the monographic manner that was used for the others. The reasons for these omissions vary as outlined below. The subfamily Aleocharinae contains a large and varied assortment of insects but is rather homogeneous in structure. Since the time of Erichson its subdivisions have been based in the primary divisions upon the number of the tarsal and palpal segments. The usual minute size of these species, and the occasionally great specializa- tion, make it very difficult to count these segments under ordinary MONOGRAPH OF WEST INDIAN STAPHYLINIDAE oD conditions of examination. Indeed, it is absolutely necessary to know the amount of magnification used by the previous workers in order that one should not see more than they were able to see and therefore get a different result. One recent worker has gone so far as to suggest that possibly all Aleocharinae have the tarsi 5-segmented in reality, although they have been described as having them 5-5-5, 5-5-4, 5-4-4, 4-44, 3-3-3, and even more reduced. This is certainly not the case, but examples are numerous to show that this group is a plastic or changing one in regard to these matters of segmentation (and perhaps other characters). It should be ob- vious, therefore, that a reliable classification cannot be built primarily upon the tarsal and palpal segmentation, and inasmuch as our only classifications now do depend on those characters there is a high de- gree of probability that there are duplications and involved synony- mies to be discovered. In such a situation it is very hard to see what is gained by describing new species or by attempting to make identi- fications. If one may judge from the flood of new species of Aleo- charinae that have been described in recent years, there are students of this family who will not agree with this point of view. Never- theless, until some sort of practicable order be brought into what appears now to be a hopeless chaos (from the point of view of all save a few specialists), I believe it far better not to add to the con- fusion by publishing new species or even new records, except in a few cases where the species are readily recognizable in spite of the generic confusion. For this subfamily, therefore, I have not at- tempted to prepare diagnoses, keys, or descriptions, although many of the West Indian species are probably recognizable when isolated from all foreign species. It has been thought worth while to list the species that have been described and give the West Indian rec- ords. It appears that this is all that can be done at the present time without adding to the confusion that already exists. The subfamily Euaesthetinae is a small and relatively unstudied one. Virtually none of the genera have been described at all, and generic assignments can be made only with the greatest difficulty. The time was not available to make a generic revision of the group, so I have merely listed the described species and the known records. The genus Palaminus contains several species described from the West Indies. The other American species are very imperfectly known and the limits of specific variation have not been ascertained. Some material in this genus was available to me, but it seemed insuf- ficient to permit decisions as to specific limits. I have, therefore, listed the species without keys, given the known records, and left the remaining specimens to be studied at some later time. 36 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Certain other genera have not been available to me except from the original descriptions. ‘These include chiefly genera recently de- scribed by Bierig. Without in any way intending to question the validity of these genera, I have been unable to include them in proper form for lack of specimens to examine. In some cases translations of the original descriptions are given, and in some only a brief sum- mary is included. These genera will have to be restudied before they can be fitted into the classification presented here. The genus Cephaloplectus has been found in Trinidad (species godmani) and would normally have been included in this study as the subfamily Cephaloplectinae. However, Dr. C. H. Seevers, of Chicago, has recently studied this group and believes that it defi- nitely does not belong in the Staphylinidae. His results wil! be pub- lished soon, but the genus is omitted from this paper on the basis of his opinion. FAUNAL RELATIONSHIPS Although our records are not complete enough to enable us to draw final conclusions as to the distribution (either present or his- torical) of the family Staphylinidae in the West Indies and sur- rounding regions, it is possible to get some indications of the general situation. The facts of distribution so far as known are presented in the following tabulation, in which the second group indicates the number that do not show any interisland distribution, and the succeeding groups deal with those that do yield evidence on this subject. Total number of species— Recorded from ‘West: Indiese 2) a2. = te se oe eee 468 Represented by type material_._________________---- 117 (plus 212 new) = 829 Mesceribedijas newest) 2h ee oe. es es ee a ee Se ee ee 212 Not herein jrecognized 2222 22-28 4-23 2S a ee 34 SPO OMby: Dl, TS) apy ch ea A ee Se ee ee eee ee eee 270 rom (Greater }Antillesonly®.i2) 20s a see eee 182 Krom) Lesser, "Antilles oily ope ko ib eit Stl pile Dive Sener es ae ee 34 From ‘Trinidad’ only. 2 ng ee eee 54 Showing distribution’ in West Indies) 2222 eo 2 se a te ee 198 Krom ‘most tof islands 2 (e008 PO) Sete ey ee eae Eee eee 49 Krom: Greater Antilles (only22s--2 2 eo ee ee eee 48 From. Lesser Antilles only si. 222). cts | es ee St ee ee 22 From 1-or more in each group.2o- 2 ee 38 Showing distribution outside West Indies__-__-__________--__---------._- 97 From West Indies and America, only_222:l-2u) 220) Sate se 79 From West Indies and eastern hemisphere only__------------------ pe MONOGRAPH OF WEST INDIAN STAPHYLINIDAE an Hroniavwese, Ludies/ and North’ America ronly2ve22 soon eee ee 27 From Greater Antilles only and North America only________-__--__- 2 From Greater Antilles only and Trinidad and North America only____ 1 From Lesser Antilles only and North America only____-__--__-_____ 0 From both groups and North America only__---_------~-- epee eee eer ee 5 From Trinidad and North America only Wrom. West Indies and Central, America :onlystes222222. 1-2 ee 46 From Greater Antilles only and Central America only__--__-_------~- 4 From Lesser Antilles only and Central America only___------------- it Hromipotk ercoups and ‘Central, America onlys_4222222-_ =: Bees os 3 From Trinidad only and Central America only___--__------__-__--__- aL From West Indies and South America only___._____--________1__-_____- 56 From Greater Antilles only and South America only__---------_----- 0 From Lesser Antilles only and South America only__--------------- 7 Hrom both groups*and ‘South’ Ameriea “only 22 2 2s_ 25. 2 8 From Trinidad only and South America only_-_--_----_--__---------~- 6 From West Indies and North America and Central America_________-____ 2 From West Indies and North America and South America_____---_______ 3 From West Indies and Central America and South America________-______ 13 From Trinidad and Central America and South America________________- 6 From West Indies and North America and Central America and South DETTE ee aa Rie popes bs eke SNS a oa ee) Oe ag 15 From Trinidad and North America and Central America and South America. 1 From these tabulations it appears that 8 percent of the species not restricted to the islands occur also in North America only, 9 percent in Central America only, 21 percent in South America only; 2 per- cent in both North and Central America, 8 percent in both North and South America, 13 percent in both Central and South America, and 15 percent in all three parts of the mainland. Furthermore, combining the above, 27 percent occur in the islands and in North America (at least), 46 percent in the islands and in Central America (at least), and 56 percent in the islands and in South America (at least). Of these same 97 species that occur in the islands and on the mainland, 16 percent occur also in the Eastern Hemisphere. And 2 species in the West Indies occur also in the Eastern Hemi- sphere but not on the American mainland at all. Some entomologists have considered Trinidad (and Tobago) to be South American and not related to the West Indies faunally. In support of this the present study includes 54 species that occur in Trinidad alone and 18 species that occur in Trinidad and South America but not in the true West Indies. It should be noted that the staphylinid fauna of Venezuela is only slightly known, and it is likely that many more of the Trinidad species will be found also on the mainland. The evidence that bas led me to include Trinidad 38 BULLETIN 182, UNITED STATES NATIONAL MUSEUM in this study of the West Indies is principally the number of species occurring in the islands and also in Trinidad. ‘This number is 50, and only half of these are yet known from South America. Six genera occur in Trinidad and South America but not in the true West Indies. These are Leptochirus, Ophites, Suniocharis, Ro- netus, Philothalpus, and Plochionocerus. They support the conten- tion that Trinidad is faunally South American. These statements are all based on the data now available and will undoubtedly be revised as our knowledge of the West Indian Staphy- linidae increases. ‘The conclusions are in general accord with those derived from studies of other groups of animals and of plants, except as to the inclusions of Trinidad and Tobago in the West Indies. Although it is not practicable to include here lists of the species recorded from each island, it is of interest to note the total number of species that have been taken on each one. The totals include the species collected by me, those unpublished records I have found in various collections, and all the previously published records: Ber- muda, 2; Bahamas Islands, 1; Cuba, 172; Isle of Pines, 4; Cayman Islands, 3; Jamaica, 128; Hispaniola, 119; Mona, 1; Puerto Rico, 103; Vieques, 3; Culebra, 1; St. Thomas, 19; St. John, 11; Tortola, 1; St. Croix, 37; St. Bartholomew, 1; St. Kitts, 11; Antigua, 32; Mont- serrat, 832; Guadeloupe, 64; Les Saintes, 1; Dominica, 32; Martinique, 5; St. Lucia, 50; Barbados, 17; St. Vincent, 76; Mustique, 9; Carria- cou, 7; Grenada, 84; Tobago, 11; Trinidad, 122; and Curagao, 1. Together these records include 468 species. Of these 59 are known to occur also in South America, 51 in Central America, and 28 in North America. SOURCES OF RECORDS Several sources of records have been mentioned in previous para- graphs. The largest one, of course, has been the identifications made by me of the specimens available in several collections. Many of these identifications amount to rediscovery of species unreported since their original proposal 50 to 100 years ago. A considerable number of species have been found already identi- fied in the collections of the United States National Museum, the British Museum, Dr. M. Cameron, and myself. These determina- tions have been checked in all cases and, if verified, have been in- cluded. A number of these records are believed to be errors in identification or to represent synonyms. These appear in the syn- onymy or are discussed in the sections headed “Remarks.” The entire literature on the Staphylinidae, so far as time and facilities would permit, has been searched for references to the West Indies. Records that appear to be based on correct identifications and locality records have been included; others are discussed in the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 39 “Remarks” or included in brackets as doubtful records. One source of these records has been found somewhat unreliable. Many records of “Siidamerika” in the Junk Catalog should be changed to St. Thomas, St. John, or Puerto Rico. These are principally Erichson’s species, and the error was in the translating of his localities. The principal sources of West Indian records in the literature (be- sides the Junk Catalogs) are the works of Erichson, DuVal, Fauvel, Leng and Mutchler, Cameron, Wolcott, and Bierig. Dr. Cameron and Sr. Bierig have contributed the largest number of new species all within the past 30 years. TERMINOLOGY Some terms have been employed in this paper that have not com- monly been used before in the Staphylinidae. A number of these are used and defined in a previous paper on the morphology of the Staphylinidae (Blackwelder, 1936). Others, which may not be en- tirely self-explanatory, include the following: The segments of the antennae have usually been referred to in the literature as “joints”; this is herein used only for the articulation between the segments, as “the outer segments are pubescent beyond the third joint,” that is, the fourth to eleventh segments. The base and basal angles of the head are described in relation to a line tangent to the middle of the base; if the base touches this line at two points, it is said to be emar- ginate; the distance of the eyes from the base is measured to this line. Margining of the abdomen implies the presence of parater- gites, either singly or in pairs, but these may be present in unmar- gined species as well. The words punctation, tuberculation, and sculpture are used as entirely separate and distinct. Sculpture is restricted to the minute reticulations, striations, aciculations, and unevennesses of the suriace seldom visible at all with magnifications less than X30 (on specimens of 5 mm. length or less). Tuberculi may be structurally related to punctures (or may bear punctures) but are always elevated above the surface. Umbilicate punctures have an obvious flat bottom to the depression with a smaller punctule at the middle (often setiferous). The terms spine, horn, and tooth are applied only to immovable processes of the bodywall, whereas seta, hair, setula, fimbria, calcar, and spur are used for movable processes—all morphologically setae. The word spinule as used in Osorius refers to processes that grade from large setae into tubercles bearing setae and into ordinary spines. As used in this bulletin, the words dung, manure, and excrement refer to cow dung, horse manure, and human excrement, respectively. 40 BULLETIN 182, UNITED STATES NATIONAL MUSEUM ARRANGEMENT OF SUBFAMILIES AND GENERA The arrangement of subfamilies used here is partly the result of studies carried out on the genera included herein and differs from that used in the Junk Catalog and other recent works in the follow- ing points: (1) The subfamily Oxytelinae is restricted to what is called the tribe Oxytelini in the Junk Catalog, and the Piestini, Oma- lini, and Osoriini are elevated to the rank of subfamilies; (2) the Staphylininae and Xantholininae are retained as tribes, but the Xanthopygini and Quediini are considered to be subordinate to the former; (3) the Cephaloplectinae are omitted as not belonging to the Staphylinidae; and (4) the Hypocyphtini are elevated to the rank of subfamily. In certain groups the arrangement of genera is based upon studies of the genotypes and represents new conclusions as to the relation- ships. For example, the subfamily Piestinae is restricted to the sub- tribes Piesti and Trigonuri through the removal of all the other subtribes to the new subfamily Osoriinae. The latter is rearranged into three tribes and seven subtribes, all of which occur in the West Indies. The subfamily Paederinae has been completely rearranged into a system based on the relative specialization of its members, but no attempt has been made to arrange categories between the sub- family and the genus, although such groups do certainly exist. The position of the tribes Xantholinini and Staphylinini have been re- versed because of the obviously greater specialization of the former, but no comprehensive study of the groups has yet been made. Some rearrangement of genera in the subfamily is proposed. KEY TO SUBFAMILIES OF WEST INDIAN STAPHYLINIDAE 1. Antennae inserted into vertex between eyes________--____-__-_--_-_ 2 Antenna inserted at front or side margin of head_____-______-__________ 3 2 Losterioricoxae: conieal;: not ‘expanded sa eee aie ek ees STENINAE Posterior coxae expanded laterally under the femora______ ALEOCHARINAE 3. Abdomen not margined above at sides, and fourth segment of maxillary palpus notjenlargsedvorSecurito rm el ee Se ae eee OSORIINAE Abdomen margined at sides above, or fourth segment of maxillary palpus enlargedand more or' less securiform= 222% 2) — eee 4 @. WerGb eGwO sOCOUIT = ee Oa tn es ee ees OMALIINAE Without paired ocellis sere Ase) phi ae Seraph Nee cet eee 5 5. Antennae “lO:segmented] tsk 4 ia ie eee HYPocYPHTINAE Antenna: 1-segmenteds: 2: <8 22) wees oe soe Se oe ion Ge 6 6: Anterior coxae small, elobose-.2-- = 4 cert PIESTINAE ANCTEXIO“ COXAG! CONICAL eXSenie dias sa terse 2 ne ee ee ee eee A iC “. \With elongate labral processesi22 2) Se Se STYLOPODINAE ‘Without’ elongate: labral processes:_- — 52s Ue, Pe ee ee 8 Svan tennae: distinctly, clubbed 2222 oe we Peet pe ees EQUAESTHETINAE Antennae, notdistinctly (clubbed == 22a) sa ee eee 9 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 4] 9. Second abdominal sternite present--__-______________--___--- OXYTELINAE Second andominalistemmite: absentee 32 aa se Set St ek Be 10 10. Antennae inserted along anterior margin of head__----------_--~-_--~- afl Antenna inserted at sides of head near eyes_____---_-_---~-- TTACHYPORINAE 11. Intersegmented membranes with pattern of rectangular areas. PAEDERINAE Intersegmental membranes with pattern of longitudinal rows of variously SHAPCOAVEAS ARH See prea ere et Se eee ae Ge Eee STAPHYLININAE Subfamily PIESTINAE As previously constituted this was a moderate-sized subfamily of varied structure and appearance, which has been found to be dif- ficult to define. Difficulties encountered during the study of the West Indian species led to a paper recently published? in which several of the subtribes of the Piestinae (Piestini of Bernhauer and Schubert ?) were transferred to the new subfamily Osoriinae. The genus Cephaloxynum is here transferred to the subfamily Staphy- lininae. These changes leave the Piestinae with only two West Indian genera, Piestus and Hypotelus, although other genera not found there will probably also remain in the subfamily. These lat- ter include 7rigonurus, Siagonium, Trigites (Pseudeleusis, Triga), and perhaps others. This subfamily is thus greatly reduced in size, while being made much more homogeneous through the removal of extraneous elements. KEY TO WEST INDIAN GENERA OF PIESTINAE 1. Head with regular impressed line between antennae; elytra BUTTA Teer ore Ei reeet a = i ree ey. hese th oe Peds 8 ed 9 eh te oe Ij. Piestus Head without impressed lines; elytra not striate___-__-__--_~- I. Hypotelus I. Genus HYPOTELUS Erichson Hypotelus Ertcuson, 1840, p. 840. Genotype.—H. pusillus Erichson (designated by Duponchel, 1841). Diagnosis —Body slender, subdepressed, glabrous; head rounded in front, not narrowed posteriorly; eyes small but prominent; an- tennae longer than head and pronotum, basal segment (of male) with a prominent tooth; mandibles small, unarmed; fourth segment of the maxillary palpus at least twice as long as third; labium sub- semicircular; labial palpi short, stout; pronotum quadrate, some- what narrowed posteriorly; prosternum not greatly elongate; elytra not extending much farther posteriorly than the metasternum, punc- tate; abdomen margined; sternites of first and second segments not present. 1Proc. U. S. Nat. Mus., vol. 92, pp. 75-90, 1942. 2 Coleopterorum catalogus, pars 19, vol. 5, p. 4, 1910. 449008-——42——-4 42 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—This is a small tropical American genus of which the species are very uniform in appearance. After repeated attempts I am unable to separate the West Indian specimens into more than one species. Since one of Bierig’s names is available, it is adopted for the specimens I have examined. I have examined 140 specimens of our species of this genus. Of these 28 are in the British Museum, 14 are in the collections of the United States National Museum, and 98 were collected by me during 1935-87, The specimens in the British Museum were labeled H. praecow Erichson and ZH. hostilis Fauvel. The former is a misplaced speci- men of a Lispinus, and the latter (28 examples) are a misidentifica- tion of H. insulanus Bierig as described herein. 1. HYPOTELUS INSULANUS Bierig Hypotelus insulanus Bierta, 1934e, pp. 348, 344, figs. 2, 3. Description.—Piceous to rufotestaceous, elytra testaceous. Head more finely and irregularly punctate than pronotum, feebly and con- fusedly but moderately densely strigulose; basal segment of antenna in male more or less tuberculate, the tubercle being merely a carinate prominence on the upper inner edge. Pronotum three-tenths wider than long, sides rounded and gradually narrowed posteriorly from the middle or apical two-fifths, just visibly emarginate before the posterior angles; midline slightly elevated, smooth; sometimes with a pair of irregular foveae before base; punctures moderately coarse, separated by one to three times their diameter; strigulae obsolescent in places. EHlytra scarcely longer than wide, hind angles broadly rounded; punctation shallow but otherwise similar to that of pro- notum, occasionally with a trace of serial arrangement; not at all strigulose. Length, 2 mm. Type locality—Cuba, Aspiro, at base of the Sierra del Rosario. Types.—Described from five examples collected March 28, 1934. Types are not mentioned in the description but are presumably in the collection of Alexander Bierig, Habana, Cuba. (One “cotype” is in the United States National Museum, No. 52699.) Records—The following are the records known to me: Cuba: Aspiro (Bierig, 1984), Cayamas (Schwarz, in U.S.N.M.). Jamaica: Bath in St. Thomas (Blackwelder station 392B), Morant Bay (Black- welder station 886A), Trinityville (Blackwelder station 428), Santa Cruz (Blackwelder station 421). Hispaniola: DoMInicAN Repusiic, Jarabacoa (Darlington, in M.C.Z%. and U.S.N.M.). St. Vincent: (British Museum). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 43 Specimens examined.—I have seen the paratype and 13 other exam- ples in the United States National Museum, 28 in the British Museum under the name #, hostilis Fauvel, three from the Museum of Com- parative Zoology, and 98 collected by Chapin and Blackwelder in 1935-37. Remarks—I1 am unable to segregate these specimens into more than one species. There is considerable variation in my large series in the punctation and sculpture, in the pronotal foveae, and in other characters. This species can be distinguished from the Central American ZZ. hostilis Fauvel by the presence of ground sculpture on the pronotum, the absence of basal depressions on the pronotum, and the evenly spaced pronotal punctures. Sr. Bierig records this species from under the bark of a “gum” tree (tropic birch ?). Our Cuban specimens were collected “on ceiba”; that is, on a silk cotton tree, presumably under the bark. The Jamaica examples were taken from under bark of the silk cot- ton tree and the tropic birch, as well as flying at dusk. II. Genus PIESTUS Gfavenhorst Piestus GRAVENHORST, 1806, p. 228. Trichocoryne GRAY, 1832, p. 306. Tricoryna LAporTE, 1834, p. 126 (misspelling). Trichocoryna BRULLE, 1837, p. 91 (misspelling). Subgenus Zirophorus DALMAN, 1821, p. 372. Zyrophorus LATREILLE, 1832, p. T7 (misspelling). Irenaeus LEACH, seq. Latreille, 1832, p. 87 (as synonym of Zyrophorus). Traeneus LAcorDATRE, 1854, p. 127 (misspelling). Subgenus Antropiestus BERNHAUER, 1917a, p. 45. Genotype.—P. sulcatus Gravenhorst (monobasic). Of Z'richoco- ryne, Zirophorus penicillatus Dalman=Trichocoryne penicillatus (Dalman) (monobasic) ; of Z7rophorus and Trenacus, Z. fronticornis Dalman=/. fronticornis (Dalman) (designated here) ; of Antropies- tus, A. andinus Bernhauer (as Piestus (Antropiestus) andinus Bern- hauer) (monobasic). Diagnosis —Body broad, depressed, glabrous, shining; head with impressions on vertex; eyes well developed but not prominent, an- tennae elongate, segments 4 to 10 pubescent; mandibles dentate; last segment of maxillary palpus longer than the third, labial palpi short, filiform; gula narrow, elongate; pronotum quadrate; prosternum not exceedingly elongate; anterior coxae globose or transverse, not pro- jecting; anterior tibiae dentate externally; elytra generally striate (always in our species), not extending beyond metasternum poste- 44. BULLETIN 182, UNITED STATES NATIONAL MUSEUM riorly; alate; posterior coxae contiguous, “transverse” (in the usage of previous writers, but “not expanded laterally and caudally” in the usage of Blackwelder, 1936, p. 76) ; abdomen margined; sternites of first and second segments not present. Remarks — With the exception of P. capricornis Laporte all the West Indian species belong to the subgenus Piestus (s. str.), which is characterized by its unarmed head, smooth pronotal disk, and stri- ate elytra. P. capricornis belongs to the subgenus Zirophorus Dal- man, characterized by the two horns on the front of the head. It is doubtfully West Indian. Another species of Piestus is recorded by Leng and Mutchler (1914, p. 403) from Haiti; it is P. meaicanus Castelnau. The type in the British Museum is very distinct from all West Indian examples of the genus I have seen and leads me to assume that the record is based on a misidentification. Specimens of a Piestus from Cuba, Haiti, and Trinidad in the British Museum were identified by Dr. Cameron as P. minutus Erichson. They dif- fer, however, from Central and South American examples of minutus in the same collection by the sculpture of the head and the puncta- tion of the pronotum. They are herein assigned to P. penicillatus (Dalman). I collected 268 examples of our species of this genus during 1935- 37; 62 examples were already in the collections of the United States National Museum; and 198 examples were studied in the collections of the British Museum. In addition to these I have seen 25 examples from the Museum of Comparative Zoology and 3 from the American Museum of Natural History. KEY TO WEST INDIAN SPECIES OF PIESTUS 1.. Front, of head prolonged as two. horns_—-_—-—+____-- === 5. capricornis Hront.of head notprolonged = —- =e eee 2 2. Mivtha nwith aS ECS Gn ae es as ee ie Se eee 1. sulcatus Blytra “with sixth ’stria’ obsolescent’2. + S52 ie ke ee en Pe eee 3 8. Sulci of pronotum one-third length of pronotum or less_____-_-__________- 4 Sulci about one-half length of pronotum___------__-___-__-___ 4. pygmaeus 4. Pronotal sculpture very fine and sparse; large discal area SIMOOUN VR eke ee Ss oe 8. penicillatus Pronotal sculpture comparatively coarse and very dense; only a small smooth ares se 2 eee eee) ee Nak eee Oe Ee ee ee ee 2. fulvipes 1. PIESTUS SULCATUS Gravenhorst Piestus sulcatus GRAVENHORST, 1806, p. 224.—LEPELETIER and SERVILLE, 1825, p. 128.—EnricuHson, 1840, p. 835.—Lacorpairg, 1854, p. 128.—SHaArp, 1876, p. 407.—BERNHAUER and ScHUBERT, 1910, p. 7. (Not Laporte, 1834.) Description.—Piceous to rufotestaceous. Head with supra-anten- nal tumescences very feeble, united behind by a ridge feebly set off MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 45 from vertex, and separated from the feebly convex central clypeal area only by two shallow foveae; clypeus not separated from transverse ridge; sparsely, moderately coarsely punctate throughout and very distinctly strigulose; gular sutures impressed, leaving gula convex. Pronotum truncate before and behind, sides rounded; width to length as 3 to 2; middle moderately impressed; lateral sulci coarse, extending to middle of length; sparsely not finely punctate; densely and very distinctly strigulose. /Hlytra with striae rather coarsely punctate, intervals punctured as pronotum; sixth stria complete, wide, coarsely punctate, usually briefly double near humerus; a vague seventh stria on deflexed portion, which is rather coarsely sculptured. Length, 314 to 44% mm. Type locality —Brazil. Types.—Presumably in Zoologisches Universitét Museum, Breslau. Records —The following are the records known to me: Cuba: (Laporte, 1835). Dominica: (Blackwelder stations 287, 246B). St. Lucia: (Blackwelder stations 220D, 220G, 220H, 226, 227, 231). St. Vincent: (British Museum). Grenada: (Blackwelder station 182). Trinidad: St. Augustine (Weber, in M.C.Z.). South America: Braz (Gravenhorst, 1806; LePeletier and Serville, 1825; Fauvel, 1865; Sharp, 1876; British Museum; U.S.N.M.). Specimens examined.—I have examined 33 examples in the British Museum, 1 in the United States National Museum, 1 in the Museum of Comparative Zoology, and 104 collected by me in 1935-87. Remarks.—This species also appears to follow the cocoa cultiva- tion. Besides from the decaying cocoa pods, I have taken this species under dry dung and under fermenting chips on breadfruit tree stumps. 2. PIESTUS FULVIPES Erichson Piestus fulvipes ERIcuHSoNn, 1840, p. 833.—LAcORDAIRE, 1854, p. 128—FAvuvEL, 1865, pp. 22, 27.—FLEvuTIAUXx and SAtifé, 1889, p. 382.—BrERNHAUER and SCHUBERT, 1910, p. 7.—Lrena and Murcurrr, 1914, p. 403. Description.—Rufopiceous. Head with supra-antennal tumes- cences very large and prominent, joined to the posterior ridge which is much lower and divided at the midline by a deep notch, and sepa- rated from the triangular upper part of the clypeus by an irregular V-shaped depression, which continues to the sides where it separates the lateral clypeal prominences from the supra-antennal; clypeus ab- ruptly declivous in front, very irregularly and somewhat coriaceously sculptured; vertex strigulose (prominences obsoletely so); no punc- tures except behind the posterior ridge; gular sutures scarcely at all impressed, gula nearly even with postoccipital areas. Pronotum 46 BULLETIN 182, UNITED STATES NATIONAL MUSEUM quadrate; width to length as 3 to 214; midline finely and evenly im- pressed, impunctate; lateral sulci feeble, not exceeding one-third the length of pronotum; sparsely, finely, and very irregularly punctate; confusedly strigulose, more regularly at sides. EHlytra with striae moderately punctate; intervals finely irregularly punctate; sixth stria marked by a short impressed line at basal third and scattered punc- tures posteriorly. Length, 7 mm. Type locality—Guadeloupe, Pointe-a-Pitre. Types.—One example in the British Museum from the Chevrolat collection. Collected by F. de L’Herminier. Records.—The following are the records known to me: [Hispaniola: Hartt (Leng and Mutchler, 1914).] Guadeloupe: (Hrichson, 1840; Fauvel, 1865; Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914; British Museum). Specimens examined.—Three examples of this species were studied in the British Museum, and one of these was afterward borrowed for further study. Remarks.—This species has apparently been collected only once, probably at the same time as ?. capricornis. Tf this is so, then the remarks under capricornis apply equally here, and this also must be regarded as a doubtful West Indian record. The record of Haiti by Leng and Mutchler is unsupported by any specimens or data known to me and is distinctly to be questioned. Nothing has been recorded concerning the habits of this species. 3. PIESTUS PENICILLATUS (Dalman) Zirophorus penicillatus DALMAN, 1821, pp. 375, 376; 1823, pp. 24, 25.—LEPELE- TIER and SERVILLE, 1828, p. 818.—Gray, 1832, p. 306—HEricHson, 1840, p. 834.— FAUVEL, 1865, p. 27.—SHArRP, 1887, p. 711. Trichocoryne striata GRAY, 1832, p. 306.—Ericuson, 1840, p. 884.—FAuven, 1865, p. 29.—BERNHAUVER and SCHUBERT, 1910, p. 7. Trichocoryne penicillatus (Dalman) Gray, 1832, p. 306.—EriIcHson, 1840, p. 834.—FAUVEL, 1865, p. 29. Piestus paenicillatus Laprortr, 1834, p. 129 (misspelling). Piestus striatus (Gray) Laporte, 1834, pp. 126, 128—Hricuson, 1840, p. 834.— BeERNHAUER and SCHUBERT, 1910, p. 7. (Not Guérin, 1837, p. 32.) Piestus penicillatus (Dalman) LaAportr, 1934, p. 127.—Ericuson, 1840, p. 834.— LAcorpDAIRE, 1854, p. 128—Scrrpa, 1855, p. 302.—Favuver, 1865, pp. 28, 27.— SHarp, 1887, p. 711.—BeRNHAUER and ScHuBERT, 1910, p. 7. Piestus erythropus Ericson, 1840, p. 884.—LacorpatrE, 1854, p. 128.—DuVat, 1857, p. 46.—FAvvEL, 1863, p. 444; 1865, pp. 23, 28, 29, 30.—SHarp, 1887, p. 713.—BrERNHAUER and ScHuBERT, 1910, p. 7.—Lenc and MurcHter, 1914, p. 403.—Wotcortt, 1924, p. 77.—Gowpey, 1926, p. 11.—Wotcort, 1936, p. 196. Description.—Piceous to rufopiceous, pronotum sometimes paler. Head with large and prominent supraantennal tumescences, connect- ed behind by a complete transverse ridge, and separated from the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 47 moderately convex central clypeal area by an even V-shaped de- pressed line which does not reach the margins; with a few sparse and rather fine punctures; irregularly and obsoletely sculptured be- fore the impressed line, obsoletely strigulose behind it; gula de- pressed, sutures not separately impressed except at base. Pronotum quadrate; width to length as 8 to 2; midline smoothly shallowly impressed; lateral sulci feeble, not more than one-third the length of pronotum; finely sparsely punctate and confusedly strigulose, more regularly and densely at the sides. Hlytra with striae finely punctate, intervals each with an irregular row of fine punctures; sixth stria represented by a vague impression near humerus or apically, or with a few impressed punctures at basal third. Length, 314 to 7 mm. Type locality —‘Guadeloupe.” Of striata Gray, “West Indies”; of erythropus, “Cuba.” Types.—Presumably in the Naturhistoriska Riksmuseum, Stock- holm. Of erythropus, presumabiy in the Zoologische Museum, Ber- lin; of striata Gray, unknown. Records —The following are the records known to me: Cuba: (Erichson, 1840; Fauvel, 1863, 1865; DuVal, 1857; Sharp, 1887, as erythropus; Leng and Mutchler, 1914, as erythropus), Guantanamo (Rams- den, in A. M. N. H.), Soledad (Darlington, in M. C. Z. and U. S. N. M.), Buenos Aires, Trinidad Mountains (Darlington, in M. C. Z.), Upper Ovando River, Oriente (Darlington, in M. C. Z and U. S. N. M.). Jamaica: (Gowdey, 1926, as erythropus), Bath in St. Thomas (Blackwelder stations 392B, 426B), Liguanea Plain (Brues, in M. C. Z.), Whitfield Hall, Blue Mountains (Darlington, in M. C. Z.). Hispaniola: (Fauvel, 1863, 1865) ; Harri, Kenskoff (Darlington, in M.C.Z. and U.S.N.M.), Morne La Hotte foothills (Darlington, in M.C.Z. and U.S.N.M.) ; DoMINICAN REPUBLIC, Jarabacoa (Darlington, in M.C.Z.). Puerto Rico: (Fauvel, 1865; Leng and Mutchler, 1914, as erythropus; Wolcott, 1924, 1936), Utuado (Busck and Richmond, in U.S.N.M.), El Yunque (Black- welder station 851A). {St. Bartholomew: (Erichson, 1840; Fauvel, 1865). Apparently an error for Guadeloupe (see “Remarks.”) ] Guadeloupe: (Dalman, 1821, 1823). St. Lucia: (Blackwelder stations 207A, 207B, 211C, 211D, 444G). Grenada: (Blackwelder stations 132, 134, 137, 153B; British Museum). Tobago: (Blackwelder station 116B). Trinidad: (Busck, in U.S.N.M.), Sans Souci (Blackwelder station 98B), Man- zanilla (Blackwelder stations 102, 103A, 104A), Moruga (Blackwelder sta- tion 111), Caura Valley (Adamson, as Blackwelder station 444A), St. Augus- tine (Weber, in M.C.Z. and U.S.N.M.). [South America: VenezueLaA (Seriba, 1855), Brazm (Bernhauer and Schubert, 1910).] [Central America: Mexico (Bernhauer and Schubert, 1910).] Specimens examined—One hundred twenty-one examples have been examined: 10 in the British Museum, 31 in the United States 48 BULLETIN 182, UNITED STATES NATIONAL MUSEUM National Museum, 1 in the American Museum of Natural History, 23 from the Museum of Comparative Zoology, and 56 collected by me in 1935-387. Remarks.—There has been much confusion concerning this species. The following summary is presented in the hope of correcting this: In 1821 Dalman described the species from “Guadeloupe”; in 1823 he redescribed it from the same locality; and in 1828 LePeletier and Serville described it from the same place. In 1832 Gray described a supposedly new genus, which he called 7richocoryne and which he based on a species “from the West Indies” of which he remarked, “It appears to be the same as... Zirophorus penicillatus” but which he figured on plate 82 as “T’richocoryne striata, G. R. Gray” (this is clearly a case of a stillborn synonym, but confusion arose because he also figures on plate 52 a species he calls Zirophorus striatus Leach, which is quite a different insect. In 1834 Castelnau cited striata Gray as synonym of Piestus bicornis Olivier (this was a mistake for the striata of Leach but gave rise erroneously to the record of Colombia for striata Gray); Castelnau further cited pens- cillatus erroneously as a synonym of P. sulcatus Gravenhorst, thereby involving the sulcatus records of Cuba, Cayenne, and Brazil, which of course do not apply to the true penicillatus Dalman. In 1840 Krichson cited penicillatus from Puerto Rico and St. Bartholomew, giving Dalman as the source of the latter (this is presumably an error for Guadeloupe) ; he further described a new species he called erythropus from Cuba and to which he attached as a synonym striata Gray (since striata Gray was above proved to be a synonym of penicillatus, then erythropus must likewise be a synonym of it). In 1857 DuVal and in 1863 Fauvel considered erythropus from Cuba as a valid species. In 1865 Fauvel cited penicillatus from the same islands as Erichson (merely copying the error of St. Bartholomew) and cited erythropus from “Santo-Domingo; Cuba; Opelousas, Mex- ique,” with striata Gray as synonym. In 1887 Sharp threw doubt on the Mexican record but left the species valid; and finally in 1910 Bernhauer and Schubert accepted erythropus (with synonym striata Gray) from “Antillen, Mexiko,” and penicillatus from “Mexiko, Brasilien” (this last record must be either an error or based upon new material, and the type locality is omitted). The uniting of erythropus with penicillatus in the present study leaves us with un- questioned records for the one species from Guadeloupe, Puerto Rico, Cuba, and Hispaniola, and my collections extend these throughout the islands to Trinidad. This species is somewhat variable in the extent of the traces of the sixth stria. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 49 Schwarz recorded this species (on specimens in the United States National Museum) “on ceiba” (silk cotton tree). I collected it from under bark of logs of the same tree, in rotten coconut husks, in old banana stalks, and in rotten cocoa pods. Specimens from the Museum of Comparative Zoology are labeled “fermenting palm” and “bark of log.” 4, PIESTUS PYGMAEUS Laporte Piestus pygmaeus Laporte, 1834, pp. 127, 180—Hricuson, 1840, p. 885.—Lacor- DAIRE, 1854, p. 128.— FAUVEL, 1865, pp. 23, 33, 34—SHarp, 1876, p. 407; 1887, pp. 713, 721.—FLEuTIAUx and SArLL#&, 1889, p. 882.—BreRNHAUER and SCHUBERT, 1910, p. 7.—Lene and Muroutrr, 1914, p. 403.—Bovine and CrarigHEap, 1931, p. 114.—ScHEERPELTZ, 1933, p. 994. Piestus rufipennis CHEevrotat, MS.—FLEUTIAUX and SALLf, 1889, p. 382. Description.—Piceous to rufopiceous, elytra rufotestaceous. Head flat, with supraantennal tumescences very feeble, merging posteriorly into the flat surface of the vertex, separated from the feebly convex central clypeal area by a well-impressed U-shaped line which curves outward at the sides and is continued as a black line to the side mar- gins; sparsely and rather finely punctate, and distinctly strigulose throughout (more finely and densely on the clypeus) ; gular sutures impressed, leaving gula convex through most of its length. Prono- tum quadrate, sides feebly rounded; width to length as 3 to 2; mid- line feebly impressed and irregularly punctate; lateral sulci coarse, extending nearly to middle of length; sparsely and finely punctate with scattered larger punctures especially near the depressions, obso- letely strigulose at the sides and base only. /lytra with striae finely punctate, intervals each with an irregular row of moderately fine punctures; sixth stria represented by an irregular band of punctures. Length, 3 to 384 mm. Type locality—Colombia. Types.—Two examples on one card in the British Museum. Records.—The following are the records known to me: [Hispaniola: Hartr (Leng and Mutchler, 1914).] Guadeloupe: (Fauvel, 1865; Sharp, 1887; Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914), Gourbeyre (A.M.N.H.). Dominica: (Blackwelder station 237). St. Lucia: (Blackwelder stations 220K, 220G, 220H, 221, 226, 230). St. Vincent: (British Museum). Grenada: (British Museum). Trinidad: (British Museum), Caura Valley (Adamson, as Blackwelder station 444A), Mt. Tucuche (Darlington, in M.C.Z.). South America: Brazm (Erichson, 1840; Fauvel, 1865; Sharp, 1876, 1887; Béving and Craighead, 1931; British Museum), ArGENTINA (British Mu- seum), CotompraA (Castelnau, 1834; Erichson, 1840; Fauvel, 1865; British Museum). 50 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Central America: PANAMA (Sharp, 1887; British Museum; U.S.N.M.), Nica- RAGUA (Sharp, 1887; British Museum), GUATEMALA (Fauvel, 1865; Sharp, 1887; British Museum; U.S.N.M.), BrirrisH HonpurAs (Sharp, 1887; Brit- ish Museum; U.S.N.M.), Mexico (Fauvel, 1865; Sharp, 1887; British Museum; U.S.N.M.). Specimens examined.—I have examined 151 examples in the British Museum, 1 borrowed from the Museum of Comparative Zoology, 2 in the American Museum of Natural History, 30 in the United States National Museum, and 108 collected by me in 1935-37. Remarks.—This widespread species apparently inhabits the West Indies as far north as cocoa 1s grown in abundance. The northern islands grow cocoa in small quantities, but I was not able to collect there at the proper season to determine the complete range of this insect. Besides the cocoa pods from which I took it frequently, I found it under chips on fresh stumps of the breadfruit tree. The record of this species from Haiti by Leng and Mutchler is at least possibly a misidentification. 5. PIESTUS CAPRICORNIS Laporte Piestus capricornis LAPorTE, 1834, pp. 127, 129.—HEricuson, 1840, p. 883.—La- CoRDAIRE, 1854, p. 128.—FavveL, 1865, pp. 22, 26, 27.—SHarp, 1876, p. 406; 1887, p. 711.—FLEUTIAUXx and SALLE, 1889, p. 382.—BrRNHAUER and ScuHv- BERT, 1910, p. 6.—LeENG and MutcHter, 1914, p. 403. Zirophorus capricornis (Laporte) SHARP, 1887, p. 711.—BERNHAUER and ScHu- BERT, 1910, p. 6. Piestus muticus FAuveEL, 1902, p. 22 (as variety).—BrERNHAUER and ScHUBERT, 1910, p. 6 (as variety). Description—Rufopiceous. Head with two feeble horns, short, thick, and directed forward; deeply foveate between the eyes, trian- gular area between depression elevated and smooth; very sparsely and rather finely punctate. Pronotum nearly twice as wide as long, sides straight; smooth, scarcely punctate at sides but with a few moderately large and shallow punctures at center of disk; minutely strigulose at sides; midline finely impressed. /lytra with strong punctate striae, sixth abbreviated at middle, sometimes impressed at apex. Length, 114 mm. Type locality—Guadeloupe, Pointe-a-Pitre. (Castelnau cited this species from “Cayenne,” but the type is labeled “Piestus capricornis Lap. Type ex coll. Chevrolat. Guadeloupe. L’Herminier” and “Pointe-a-Pitre.”) Types.—One example from the Chevrolat collection labeled type in the British Museum. Records.—The following are the records known to me: Guadeloupe: (Laporte, 1834; Hrichson, 1840; Fauvel, 1865; Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). South America: CAYENNE (Laporte, 1834; Fauvel, 1865), Brazm (Fauvel, 1902, as muticus; Bernhauer and Schubert, 1910, as muticus). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 51 Specimens examined.—The type is the only example seen. Remarks.—This species belongs to the subgenus Zirophorus as de- fined by Sharp (1887, p. 711). There is at least considerable doubt of the West Indian records, and I expect that further collecting will show the species to be strictly South American. Nothing has been recorded concerning the habits of this species. Subfamily OMALIINAE This subfamily is poorly represented in most tropical regions but may be immediately recognized by the pair of ocelli on the vertex of the head. Ocelli occur in no other West Indian staphylinids, but a single ocellus does occur in species of at least one other group in other parts of the world. These beetles commonly occur on flowers but with us are often taken on decaying fruit. Two genera are here recorded from the West Indies. KEY TO WEST INDIAN GENERA OF OMALIINAE 1. Mesosternum usually more or less carinate; coarsely punctate ELT) ONO ip ane a re ee ae ae Bee DA Eee ee ee. = IV. Omalium Mesosternum not carinate; not coarsely punctate above__ III. Phloeonomus III. Genus PHLOEONOMUS Heer Phloconomus HEER, 1838a, p. 184. Distemmus LECONTE, 1861, p. 69. Phlaeonomus FAvveEL, 1868a, p. 48 (misspelling). Subgenus Phloeostiba THomsSoN, 1861, p. 208. Subgenus Xylostiba GANGLBAUER, 1895, p. 731. Subgenus Xanthonomus BERNHAUER, 1926, p. 312. Genotypes.—Omalium pusillum Gravenhorst=Phloeonomus pusil- lus (Gravenhorst) (designated here) ; of Distemmus, Trogophloeus argus LeConte=Distemmus argus (LeConte) (monobasic) ; of Phloe- ostiba, Staphylinus planus Paykull=Omalium planum (Paykull) of Gyllenhal=Phlocostiba plana (Paykull) of Thomson (designated here); of Xylostiba, Omalium monilicorne Gylenhal=Phloconomus (Xylostiba) monilicornis (Gyllenhal) (monobasic); of Xanthono- mus, Phloeonomus (Xanthonomus) toxopeanus Bernhauer (mono- basic). Diagnosis—Body depressed; head subtriangular; with two ocelli posteriorly; gula wide, narrowed anteriorly; antennae moderately long, five apical segments enlarged and densely pubescent; labrum transverse, truncate or somewhat rounded; mandible short; third segment of maxillary palpus enlarged, fourth more slender and nearly as long as first three; pronotum transverse; anterior coxal cavities confluent, widely open behind; anterior coxae transverse, not 52 BULLETIN 182, UNITED STATES NATIONAL MUSEUM prominent; mesosternum not carinate; elytra extending beyond metasternum posteriorly; posterior coxae contiguous, transverse, scarcely expanded under the femora; abdomen widely margined; sternites of first and second segments not present; femora unarmed, tibiae spinose, fifth tarsomere longer than first four. Remarks.—This is a boreal genus but is represented in the Ameri- can Tropics by more species than Omalium. It is distinguished from Omalium principally by not having the mesosternum carinate. One species only is known from the West Indies. 1. PHLOEONOMUS PEDICULARIUS (Erichson) Omalium pedicularium Exicuson, 1840, p. 887.—Fauvet, 1866, p. 14, 16—LrENne and Mutcuter, 1914, p. 403.—Worcort, 1924, p. 77; 1936, p. 196. Homalium pedicularium (Erichson) GEMMINGER and Haron, 1868, p. 667. Homalium lacrymale FrrutiIaux and SALy#, 1889, p. 381. Phloeonomus pedicularius (Erichson) BERNHAUER and SCHUBERT, 1910, p. 59. Omalium lacrymale (Fleutiaux and Sallé) BrernHAveR and Scuuserr, 1910, p. 54, Omalium lachrymale Lene and MutcHreEr, 1914, p. 403. Description——Piceous to piceocastaneous, pronotum and base of abdomen castaneous, elytra testaceous with scutellar area, outer mar- gin, and wide apical band piceocastaneous (sometimes testaceous with head and elytral markings darker). Head with vague depressions above base of antennae and in front of antennae; not punctate but with distinct scaly ground sculpture. Pronotum nearly one-half wider than long, evenly rounded at sides, anterior angles rounded, posterior angles prominent; sides of disk scarcely explanate; with two vague longitudinal impressions on the disk; the sparse and feeble punctation obscured by the dense scaly sculpture. Mesoster- num not at all carinate. Hlytra fully one-fourth wider than pro- notum, widest posteriorly, just broader than long; punctures mod- erate, separated by a little more than their diameter but obscured by the dense but rather feeble scaly ground sculpture. Abdomen very minutely punctured; sculpture obsolescent. Length, 134 to 2 mm. Type locality—Puerto Rico. Of lacrymale, Gourbeyre, Guade- loupe. Types.—In either the Hope Museum, Oxford, or the Zoologische Museum, Berlin; of Zaerymale, possibly in the Musée National d’His- toire Naturelle, Paris. Records.—The following are the records known to me: Cuba: San Juan Mountains (M.C.Z.). Jamaica: Santa Cruz (Blackwelder station 421), Maggoty (Blackwelder station 402), Trinityville (Blackwelder station 428), Moneague (Blackwelder sta- tion 869B), Troy (Blackwelder station 409), Montego Bay (A.M.N.H.), Cinchona (A.M.N.H.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 53 Hispaniola: DomMInIcAN REPUBLIC, Santiago (U.S.N.M.). Puerto Rico: (Hrichson, 1840; Fauvel, 1866; Leng and Mutchler, 1914; Wolcott, 1924, 1936). Guadeloupe: (Fleutiaux and Sallé, 1889, as lacrymale; Leng and Mutchler, 1914, as lachrymale) ; Gourbeyre (A.M.N.H.). Grenada: (Blackwelder stations 132, 137, 153A). Trinidad: Manzanilla (Blackwelder station 103A). Specimens examined.—I have examined 2 specimens in the United States National Museum, 5 in the American Museum of Natural History, 1 in the Museum of Comparative Zoology, 6 collected by Chapin and Blackwelder in Jamaica, and 300 collected by me in 1935-37. Remarks—tIn the Bernhauer and Schubert catalog pedicularius is placed in the genus Phloeonomus and lacrymale in the true Omal- zum. Iam unable to separate them in the material at hand and find no records of either species having been taken since the original description. Whether the species is found also in South America is not known to me, but it seems distinct from the two described species from that region, as well as from the Central American ones. I have collected this species in rotting cocoa pods, under ferment- ing chips on a fresh stump, flying at dusk, and in a mixed lot from fungus and rotten grapefruit. IV. Genus OMALIUM Gravenhorst Omalium GRAVENHORST, 1802, p. 111. Ochtherenus MorscHULsKyY, 1860a, p. 546. Homaliwm GEMMINGER and HAROLD, 1868, p. 665 (emendation). Ornalium WotcoTt, 1924, p. 77 (misspelling). Subgenus Stenomalium BERNHAUER, 1939, p. 194. Genotypes.—Staphylinus rivularis Paykull=Omalium rivulare (Paykull) (indicated by Duméril, 1806; designated by Latreille, 1810). (The designation of depressum by Samouelle, 1819, and of planum Paykull by des Gozis, 1886, therefore cannot stand.) Of Ochthexenus, O. clavicornis Motschulsky (monobasic); of Steno- malium, Omalium (Stenomalium) rufiventre Bernhauer (designated here). Diagnosis.—Body depressed; head broadly triangular; with two ocelli posteriorly; gula wide, narrowed anteriorly; antennae moder- ately long, five apical segments enlarged and densely pubescent; labrum transverse, feebly emarginate; mandibles short; third seg- ment of maxillary palpus enlarged, fourth nearly as long as first three; pronotum transverse; anterior coxal cavities confluent, widely open behind; anterior coxae transverse, not prominent, mesosternum more or less carinate; elytra extending beyond the metasternum posteriorly ; posterior coxae contiguous, transverse, scarcely expanded 54 BULLETIN 182, UNITED STATES NATIONAL MUSEUM under the femora; abdomen widely margined; sternites of first and second segments not present; femora unarmed, tibiae spinose, fifth segment of hind tarsi longer than first four together. Remarks.—This is a boreal genus that is very poorly represented in the American Tropics. At least several of the species described by Sharp from Central America will be found to belong to the genus Phloeonomus rather than to Omalium. I have seen 34 specimens, which I assign to 4 species. These are found only in the Greater Antilles. KEY TO WEST INDIAN SPECIES OF OMALIUM 1. Side margins of pronotum distinctly curved up; lateral depression large-and deepens: ase tas = Se oi a ee teense 4. laborium Side margins of pronotum explanate but not curved up; lateral depressions; feeble ;orsmodersiles2=- 2a = a a ee ae ee ee 2 2, Smooth preocellar ridges very distinct_-_._-_-____-.----__ 1. jamaicensis Preoceliarridgesmecbles22a2 = <5 1 Sa ee ee ee ee es ee S 3. Intervals of pronotum distinct, shining _-_____-__.________. 2. darlingtoni Intervals-of pronotum almost invisible=—2.-—_ 2225222 -- = 3. cubanum 1. OMALIUM JAMAICENSIS, new species Description—Bright rufous throughout. Head broadly elevated along middle; feebly margined above and behind the eyes; feebly nar- rowed behind eyes to margin, thence abruptly ; ocelli prominent at base, not set off by smooth areas; punctures moderate but obscured by ground sculpture. Pronotum one-third wider than long; sides rounded in front, without lateral angle, straight from middle to prominent basal angles; strongly explanate at the sides but not up-curved; very feebly foveate at middle of sides; disk very vaguely depressed on either side of midline posteriorly; moderately coarsely and rather densely punc- tate, the intervals narrow and with traces of ground sculpture. Elytra nearly one-third wider than pronotum, one-fourth longer than wide; very feebly explanate at the sides; with coarse punctures, not very dense, but obscured by irregular ground sculpture. Abdomen above with moderately coarse but not dense punctures obscured by dense ground sculpture. Length, 134 mm. Type locality —Jamaica, Yallahs Valley, Blue Mountains. Types.—Holotype, American Museum of Natural History, collected by J. A. Grosbeck on February 27, 1911. Records.—The following is the only record known to me: Jamaica: Yallahs Valley (Grosbeck, in A.M.N.H.). Specimens examined.—I have seen only the holotype. Remarks.—This species is distinct in numerous characters, such as the shape of the pronotum and the elytra and the punctation and sculpture of the head, pronotum, elytra, and abdomen. The type was collected during general sweeping. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 55 2. OMALIUM DARLINGTONI, new species Description—Piceous to rufopiceous, the margins rufous. Head broadly raised along midline; margined above and behind the eyes; feebly narrowed behind eyes to margin, then abruptly; ocelli promi- nent at base, separated from punctate area by smooth diagonal promi- nences; punctures moderately large but irregular, intervals sometimes large and shining; in front with ground sculpture only. Pronotum one-third wider than long; sides rounded in front, feebly angled at middle, then straight to prominent posterior angles; strongly ex- planate at sides but not upcurved; feebly foveate at middle of sides; disk with very feeble U-shaped impression; strongly and coarsely punctate, with narrow and convex intervals; with traces of ground sculpture on the intervals. Hlytra two-sevenths wider than pronotum, just longer than wide; feebly explanate at sides; densely and moder- ately coarsely but irregularly punctate with traces of ground sculp- ture on the intervals. Abdomen above sparsely and shallowly but not finely punctate; with feeble scaly ground sculpture. Length 21% to 3 mm. Type locality —Jamaica, main range of the Blue Mountains, eleva- tions between 5,000 and 7,388 feet. Types.—Holotype and 16 paratypes, Museum of Comparative Zool- ogy; nine paratypes in the United States National Museum (No. 52481) ; collected by Dr. P. J. Darlington on August 17-19, 1934. Records.—The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 26 types. Remarks.—This series shows very little variation and is therefore my principal reason for keeping separate the other three species. The characters used are not striking, yet they seem to be constant and to be sufficient to separate the species if used with care. I have received no record of the habits of this species. 3. OMALIUM CUBANUM, new species Description —Rufopiceous, margins somewhat paler. Head broadly raised along midline; margined above and behind the eyes; scarcely narrowed behind eyes to margin; then abruptly ; ocelli promi- nent at base, separated from punctate area by smooth diagonal prominences; punctures moderately large but very irregular, the inter- vals often large and shining; in front with ground sculpture only. Pronotum nearly one-half wider than long; sides feebly arcuate in front, distinctly angled at center, thence straight or slightly emarginate to the distinct basal angles; broadly explanate at sides but not up- curved; very feebly foveate at middle of sides; disk with a distinct U-shaped impression ; strongly and coarsely but irregularly punctate, 56 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with broad shining convex intervals; without ground sculpture on the disk. lytra about one-fourth wider than pronotum, a little longer than wide; feebly explanate at the sides; coarsely and moder- ately densely punctate, with narrow and convex intervals; with traces of ground sculpture on the intervals. Abdomen above impunctate except for a pair of discal punctures on each segment; with dense ground sculpture. Length, 2 mm. Type locality—Cuba, Pico Turquino, summit at 6,000 feet. Types.—Holotype, Museum of Comparative Zoology, collected by Dr. P. J. Darlington on June 16-21, 1936. Records.—The following is the only record known to me: Cuba: Pico Turquino (Darlington, M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species differs from darlingtoni and jamaicensis also by the punctation of the abdomen. I have little hesitation in describing it from a unique specimen since the series of darlingtoni indicates that there is little variation. I have received no record of its habits. 4. OMALIUM LABORIUM, new species Description.—Clear rufopiceous throughout. JZead broadly raised along midline, margined above and behind the eyes; feebly narrowed behind the eyes to margin, then abruptly; ocelli very prominent at base; with very large coarse punctures except in front, separated by irregular convex prominences; in front with distinct almost scaly ground sculpture; submentum and mentum with very coarse punc- tures; third segment of maxillary palpus not enlarged, fourth elon- gate, as long as first three together but not more slender than third. Pronotum nearly three-fourths wider than long, sides rounded in front but feebly angulate at middle, thence straight to posterior angles, which are prominent; disk strongly explanate at the sides and foveate at middle of sides; with two feeble longitudinal impres- sions near midline; very strongly and coarsely punctate, with narrow and convex intervals; no ground sculpture. lytra one-twelfth wider than pronotum, just longer than wide; feebly explanate at sides and apex; punctures very coarse and deep, narrowly separated by convex ridges, and sometimes vaguely serial. J/esosternum carinate at base and apex only, with distinct ground sculpture. Abdomen impunc- tate except for a pair of large punctures on the disk of each sternite, but with dense scaly sculpture. Length, 214 to 3 mm. Type locality.— Haiti, east slope of Morne La Selle at elevation of about 6,000 feet. Types.—Holotype and two paratypes, females, U.S.N.M. No. 52367, collected by me on August 8, 1935. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 57 Records.—The following are the records known to me: Hispaniola: Harr1, Morne La Selle (Blackwelder station 22C), La Visité (Dar- lington, in M.C.Z. and U.S.N.M.); DominicAn ReEPuBLIC, Loma Rucilla (Darlington, in M.C.Z.). Specimens examined.—I have seen only the three types and three specimens from the Museum of Comparative Zoology. Remarks.—This species seems to be quite distinct from the described American ones. It was found at a high elevation (for the West Indies), similarly to the Central American species recorded by Sharp. The specimens were taken by beating plants along the edge of the jungle, where it was replaced by pine forest. Subfamily OXYTELINAE This name has been used for groups of various ranks in recent years, but in the Junk Catalog ® it is treated as a subfamily, which includes the Piestinae, the Omaliinae, the true Oxytelinae, and the Osoriinae. In this condition it is exceedingly difficult to define the subfamily or to arrange characters so that it may be placed in a key. The Osoriinae, the Omaliinae, and the true Oxytelinae may each be defined readily by a single morphological character of considerable importance, and there seems to be no reason whatsoever for grouping them under one head. Indeed, recent studies indicate that these four groups are not at all closely related, and each of them is there- fore treated here as a subfamily. The Oxytelinae may be recognized among all staphylinids by the presence of a complete sclerite in the second ventral abdominal seg- ment. This gives them an additional “visible ventral segment.” Eight genera have been recorded from the West Indies. KEY TO WEST INDIAN GENERA OF OXYTELINAE 1. Elytra dehiscent or inner apical angles rounded___---__---_--___---__~- 2 Elytra not dehiscent; inner apical angles not rounded___------__--_--~-~- aa 2. Front tibia with one or two external rows of spines__----__-----+--__-- 3 Front tibia without external rows of spines__-_------------- X. Thinobius BOLO’ DiOlas WiItNLWOLLOWSs OL spines... -_= = XII. Bledius rout tibia, withone-row of spines22=) 2 eee XI. Platystethus 4. Labrum emarginate and dentate at sides________ EACETEE NYS _ 95) IX. Parosus Gabrum) transverse) but, not. emarginates=a2- 5 4-5 tet dee nd tes 5 5. Front tibia with an external row of. spines-_-_--__-__-_-_~_ VIII. Oxytelus Pron tipiar Wwiatnout.external stow..Of Spinesss22—-—-. ee 6 Gee E0dy SIMOOLAT ANG Shen tT yah eee es es VII. Apocellus Body with more or less dense punctures or sculpture__-___-__-_------- if 7. Head very strongly constricted behind____._-_--_---__- VI. Torrentomus Head not very strongly constricted_ ou. 222-+0--i2.-L_---- V. Carpelimus ® Bernhauer and Schubert, 1910. 449008—42—_5 - 58 BULLETIN 182, UNITED STATES NATIONAL MUSEUM V. Genus CARPELIMUS Samouelle Carpelimus SAMOUELLE, 1819, p. 174. Trogophloeus MANNERHEIM, 1830, p. 49. Taenosoma MANNERHEIM, 1830, p. 50. Carpalimus STEPHENS, 1832, p. 324 (misspelling). Tragophloeus Laporte, 1834, p. 125 (misspelling). Carpalinus BruLif, 18387, p. 92 (misspelling). Trogophlaeus Westwoop, 1839, p. 172 (misspelling). Boopinus KiiMa, 1904, p. 56. Trogoploeus EICHELBAUM, 1909, p. 123, (misspelling). Subgenus Teropalpus Sorter, 1849, p. 330. Trogolinus SHARP, 1900, p. 231. Subgenus Thinodromus Kraatz, 1858, p. 866. Subgenus T'roginus Mutsant and Rey, 1879, p. 316. Subgenus Amisammus DES GoziIs, 1886, p. 5. Amisanimus BrRNHAUER and ScHUBERT, 1910, p. 95 (misspelling). Genotypes—C. bilineatus Stephens (designated by des Gozis, 1886). Of Trogophloeus, Oxytelus corticinus Gravenhorst=T'rogo- phloeus corticinus (Gravenhorst) (monobasic); of Z'aenosoma, T. gracile Mannerheim (designated here); of Boopinus, Trogophloeus memnonius Erichson=Boopinus memnonius (Erichson) (designated by Tottenham, 1939); of Teropalpus, T. suturalis Solier (designa- tion compulsory, since the other two originally included species were doubtfully assigned to the genus) ; of 7rogolinus, Trogophloeus uni- color Sharp=Trogolinus unicolor (Sharp) (designated by Totten- ham, 1939); of TAénodromus, Trogophloeus dilatatus Krichson= Thinodromus dilatatus (Erichson) (monobasic); of 7’roginus, T'ro- gophloeus exiguus Erichson=Troginus exiguus (Erichson) (desig- nated by Sharp, 1887, and by Tottenham, 1989); of Amzsammus, Carpalimus arcuatus Stephens=Amisammus arcuatus (Stephens) (monobasic and original designation). Diagnosis —Head cylindrical, eyes generally prominent, without ocelli; antennae 11-segmented; labrum large, transverse; maxillary palpus subulate; gular sutures united; pronotum not sulcate; hypo- mera with a lobe partly behind the coxae; anterior coxal cavi- ties open behind and confluent; front coxae large, exserted; scutellum generally not visible; middle coxal cavities narrowly separated ; posterior coxae contiguous, “transverse”; abdomen margined, first sternite absent; tibiae not spinose; tarsi 3-segmented. Remarks.—This is a very difficult genus. It has not been properly studied in any part of the New World, at least. Careful comparison with types and paratypes has made me reasonably sure of the identi- fication of all but four of the species recorded from the West Indies. A key is presented that includes only the species known to me by specimens. It is considered only as a preliminary attempt and will not be found adequate for all material. Indeed, I have numerous MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 59 specimens I am unable to place in the key, and I am leaving them aside for the time being. The descriptions are brief but contain the characters upon which my conclusions have been based. The record of Trogophloeus corticinus (Gravenhorst) from Guade- loupe by Leng and Mutchler cannot be verified in the material avail- able to me. It may be the same as the 7’. fulvipennis Fauvel, but I am unable to prove the identity of these two species. I have omitted 7’. corticinus as a doubtful record. The name Z7’rogophloeus has long been used for this genus in spite of the existence of the older name Carpelimus. In pointing out this error Tottenham (1939, p. 227) uses the spelling Carpalimus, ap- parently first used by Stephens in 1832. If this name was validated by Samouelle, the original spelling must be accepted, in the absence of proof that it was anerror. The arrangement of subgenera is based on the assignment of the genotypes in the Bernhauer and Schubert catalog. I have examined over 1,500 examples of this genus from the West Indies; 906 are in the British Museum, 63 in Dr. Cameron’s collec- tion, 131 in the Museum of Comparative Zoology, 27 in the American Museum of Natural History, 142 in the United States National Museum, and 367 were collected by me during 1935-37. These I assign to 37 species, of which 18 appear to be new. KEY TO WEST INDIAN SPECIES OF CARPELIMUS 4 1. Elytra and abdominal tergites with abrupt discal protuberances Ole CATLIN AC See are eee ee eee eee eee Sere ee ee 7. cornucopius Hivtravand:tergites not-modified-as-abover= 2) te a 2 2. Anterior angles of pronotum prominent, tuberculate, pronotum Gistiichiye hex apr on ales eares eee ee NEE he RAE Petts cles nd ee 3 Anterior angles of pronotum not tuberculate; pronotum not dis- HNChlyshexAs Ona ese seer ea eeeet Ae eee See Cee en Sere IE eee 8 3. Pronotum with a complete transverse arcuate groove near DAS Cpe oo atin Sal nr) aie 2 Beem on rts ree LB SRNR, LAIN ES 1. croceipes Pronotum without basal groove or interrupted at middle_______-________ 4 =-Eronotum: very: densely sculptured, opaque 22) 8 Ne 5 Pronotum not very densely sculptured, shining_____________-__________ iv Comturyure: Lely tin Chat Gi aie eee Se nie ee EE 4. correctus Hiyira’ cogmmely “punctate. ve.) Se ee ee Bot ee 5. darlingtoni 6. Apical margin of elytra distinctly paler______________._________ 6. beattyi VER ars UNL COLOR OUR eee le 2 SB es eh wre MVR TEN OR UU BOY t 7. Pronotum very feebly punctate or sculptured__.____--_________ 3. sericeus Pronotum rather densely and distinctly sculptured____________ 2. fulvipes 8. Pronotum with a complete transverse arcuate groove near base_________ 9 Pronotum without basal groove or with it interrupted at middle_______ 14 9. Pronotum distinctly punctate, generally coarsely so_________-________ 10 Pronotum. noyidistinetly punetates 252k wi seo i eg a *The following species have not been recognized in the material available to me and are not included in the key; their original descriptions are placed after the other species: C. fulvipennis (Fauvel), testaceipennis (Cameron), sordidus (Cameron), and demmeli (Bierig). 60 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 10. Coarse punctures of pronotum distinct-_-__-__----------------- 8. smithi Punctures. of pronotum not discrete, obscured by ground Sculpture: hs = a e 9. weberi 11. Pronotum with distinct longitudinal (or diagonal) impressions________ 12 “Pronotum without distinct longitudinal impressions______------------ 13 12. Longitudinal depressions forming a deep transverse median STOO VE ee ee ee es ee ee ee eee 138. borinquensis Longitudinal depressions feeble and not forming a deep trans- verse median. groove=—--=. 2. =f 8s Ro 12. obsolescens 13. Elytra moderately coarsely punctate; pronotum without trace Ol # DUN CCU CS== = = =a ne ea ee ee ee 11. impunctus Blytra very coarsely punctate; pronotum with fairly distinct shallow punctures: near bases i Sekt Ne te see 10. hispaniolus 14. Pronotum with longitudinal depressions (sometimes obsolescent) -------- 15 Pronotum entirely without trace of longitudinal depressions (sometimes midline vaguely elevated) ~----_-___-________________.___ 26 15. Wlytra “coarsely punctate.- £2 -~ = las = ee ae ee ee eee 16 Hiytra moderately or finely punctate=2=22= =e eee 18 16. Punctures of pronotum not distinct, obscured by sculpture__ 14. danforthi Punctures of pronotum distinct, not obscured by sculpture____----_--- 17 17. Pronotum distinctly over one-fourth wider than long_--__----~~ 18. aridus Pronotum less than one-fourth wider than long_------__-----_- 20. phaios 18. Pronotum with distinct individual punctures separated by smooth= intervals! =~ s2-2 222 eae Se ee eee 21. scrobiger Pronotum with very dense punctures or more or less distinct sculpturé 2... 22 S62 Soe eh Ba ee Se en ee 8 Se 19 19. Punctures of pronotum very fine and dense____..__+_.__._=_-_._=.-=-= 20 Punctures of pronotum not very fine or very dense (or densely sculptured). o22 2 2) 2 2222 ae ee ee en ee 22 20 ehlytra black.or ruftopiceus= 2322522 eee ee ae 22. petomus Hilytra rufous: or rufotestaceous2us=2 = 22 ee ees 21 21. Very slender: .eyes mot, prominent. 22 = es _ 25. subtilior Moderately stout; eyes very prominent_____-______~____-_-__-_ 19. aeolus 22.°Deep black. throughout__2 2222S Se 15. cubensis At least in part rufescent, rufous, or testaceouS______--_--_-—__+ -4=--2 23 23. Dlytra distinctly paler. iat ‘apexi.2255_ 2 st ee eee 16. imitator Hlytramot abruptly paler at apexise see 2 te wees Se Sees a 24 24, Pronotum shining, not very, densely, sculptured-=-—_.._=+——_S2—=———4-== 25 Pronotum not very shining, densely sculptured_----------- 17. varicornis ah-Body very. Slender... Mf. 28 2 ee ee 25. prolixus Body.. moderately Stout--=<> =. 8 2 ee eee 23. flavipes 26. Black, elytra marked with yellow from humerus to apex and alongsapiealsmargine <2 8 3s. hw, ee 28. discipennis Blytra not:so marked 24 2 kt a ee ee ee 27 27. Pronotum with a distinct narrow smooth midline_--_-_---___ o2. aequalis Pronotum without smooth; midline... -22- —2.:8 6 3 2 bee te 28 28. Pronotum very densely sculptured, not at all shining___-_____-_____-____- 29 Pronotum not very densely sculptured, more or less shining__----____- 30 29., Black or piceous, througho G20) 8 2 fee a a 26. haplomus Black, pronotum rufescent, elytra piceorufous with apical third indefinitely, .piceotestaceous.=2—...--- 8 27. tunapunae MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 61 80. Sides of pronotum arcuate or straight posteriorly______.___-_-_---_---__- 81 Sides of pronotum«distinctly emarginate posteriorly_----__-~~- 33. dissonus 31. Pronotum with distinct punctures among sculpture__-----___ 31. conformis Pronotum without distinct punctures among sculpture_-------------__ 32 32. Elytra moderately coarsely punctate________-___--__--.------ 29. chapini Hivsravvery minutely punctuates 22 2) 30. rulomus 1. CARPELIMUS CROCEIPES (Fauvel) Trogophloeus arcuatus FAuvEt, 1863, p. 489; 1868, p. 379 —FLeuTIAUX and SaLLé, 1889, p. 881.—BERNHAUER and ScHusert, 1911, p. 95.—LENe and MUTCHLER, 1914, p. 403. (Not Stephens, 1835.) Trogophloeus croceipes WAUVEL, 1866b, p. 879.—SHaRP, 1887, p. 702.—FLEUTIAUX and SALx&, 1889, p. 381.—BrRNHAUER and ScHUBERT, 1911, p. 95.—CAMERON, 1918, p. 324.—Lrne and Mutcuter, 1914, p. 403. (Not Bierig, 1935.) Description.—Black throughout. Head rather deeply bi-impressed between the antennal ridges; eyes at nearly half their width from base. Pronotum two-fifths wider than long; the sides bituberculate, the anterior angles prominent and tuberculate; disk with two arcuate transverse depressions scarcely interrupted at middle; with large and distinct punctures separated by shining convex intervals; without ground sculpture. Hlytra with moderately coarse punctures obscured by the irregular surface and traces of ground sculpture. Length, 214 mm. Type locality —Cuba (for both names). Types.—A unique in the British Museum, labeled “Cuba Poey” (for both names). Records.—Vhe following are the records known to me: Cuba: (Fauvel, 1863; Sharp, 1887; British Museum; Bernhauer and Schubert, 1911; Leng and Mutchler, 1914), Sierra del Rosario (Bierig, in U.S.N.M., as arcuatus), Hongolosongo, Oriente (Darlington, in M.C.Z.), Daiquiri, Oriente (Darlington, in M.C.Z. and U.S.N.M.), San Blas, Trinidad Mountains (Darlington, in M.C.Z.). Jamaica: (Hubbard, in U.S.N.M.) Balaclava (Blackwelder station 399A). Hispaniola: DomInicAN Repusric, Santiago (Darlington, in M.C.Z.), Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.). Montserrat: (Hubbard, in U.S.N.M.). Guadeloupe: (Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). Specimens examined.—I have seen 2 examples in the United States National Museum and 15 in the Museum of Comparative Zoology, in addition to the type in the British Museum. Remarks.—This is a very distinct species. It was renamed by Fauvel in 1868 because the earlier name areuatus was preoccupied. The name croceipes has been erroneously applied by Bierig to a very differ- ent insect, which is herein identified as smithi Bernhauer. The Jamaica examples were found on sticks and leaves more or less submerged in the center of a small stream. 62 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. CARPELIMUS FULVIPES (Erichson) Trogophlocus fulvipes Ericason, 1840, p. 804.—Duvvat, 1857, p. 43.—FAvvEL, 1863, pp. 439, 440.—LeEConrTE, 1877, p. 247.—Scuwanrz, 1878, p. 442.—SHarp, 1887, p. 697.—FAvvEL, 1901, p. 71.—VivapaA, 1901, p. 83.— BERNHAUER, 1908a, p. 230.—BERNHAUER and ScHUBERT, 1911, p. 99.—LENG and MuTcHIeER, 1914, p. 403.—NorMaAN, 1920, p. 694.—WotcorT, 1924, p. 77—Breric, 1935a, pp. 9, 11.—Wotcortt, 1936, p. 196. Trogophloeus rubripennis FAUVEL, 1863, p. 440.—BERNHAUER and SCHUBERT, 1911, p. 97 (as synonym of aridus DuvAL).—LeneG and Mutcuter, 1914, p. 403.— Bieri, 1935a, p. 9. Trogophloeus senilis SHarp, 1880, p. 51—Favuver, 1901, p. 71.—Suarp, 1908, p. 543.—BERNHAUER and SCHUBERT, 1911, p. 104. Trogophloeus texranus CASEY, 1889b, pp. 332, 834, 385.—Favuver, 1901, p. 71.— BERNHAUER and ScHUBERT, 1911, p. 99.—NotTmAn, 1920, p. 694. Trogophloeus aequalis GuNDLACH, 1893, p. 300.—WotcorT, 1924, p. 77. (Not Duval.) Description.—Head black, pronotum piceous or rufopiceous, elytra rufocastaneous, abdomen piceous. Head with two rather feeble de- pressions between the antennal ridges; eyes separated from base by about one-third their length. Pronotum nearly one-third wider than long, irregularly hexagonal, sides without tubercles, anterior angles distinctly tuberculate; disk with two distinct longitudinal depres- sions; with not very fine but moderately dense punctures, scarcely at all obscured by ground sculpture. H/ytra moderately finely and densely but distinctly punctate, with very little trace of ground sculpture. Length, 2 to 3 mm. Type locality.—Puerto Rico. Of senzlis, island of Oahu, Hawaiian Islands; of tevanus, Galveston, Tex.; of rubripennis, Cuba. Types.—Either in the Hope Museum, Oxford, or the. Zoologische Museum, Berlin. Of sendlis, in British Museum; of teaanus, holo- type and three paratypes, U.S.N.M. No. 37207, in the Casey collec- tion in the United States National Museum; of rubripennis, presumably in the collection of Blanche Rancin, Caén, France. Records.—The following are the records known to me: Cuba: (Fauvel, 1901; DuVal, 1857; Fauvel, 1863, as fulvipes and rubripennis; Leng and Mutchler, 1914, as both; Bierig, 1935, as fulvipes and rubripennis), Cayamas (Schwarz, in U.S.N.M.), Baragua (Christenson, in U.S.N.M.; C. F. Stahl, in M.C.Z. and U.S.N.M.), Rio Almendares (Bierig, in U.S.N.M.), Cayajabos (Bierig, in U.S.N.M, as rubripennis), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Cauto El Cristo, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Trinidad Mountains (Darlington, in M.C.Z.). Jamaica: (Fauvel, 1901), Milk River (Blackwelder station 415), Kingston (Black- welder stations 3, 379), Spanish Town (Blackwelder station 377), May Pen (Blackwelder station 424A), Black River (Blackwelder stations 416, 419), Santa Cruz (Blackwelder station 421), Ocho Rios (Darlington, in M.C.Z.). Hispaniola: (Fauvel, 1901) ; Hartz, Bayeux (Hoffman, in U.S.N.M.), Port-au- Prince (Darlington, in M.C.Z.) ; Dominican Repuslic, San Francisco Moun- tains (Busck, in U.S.N.M.), Barahona (Darlington, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 63 Puerto Rico: (Hrichson, 1840; DuVal, 1857; Fauvel, 1863; LeConte, 1877; Sharp, 1887; Fauvel, 1901; Leng and Mutchler, 1914; Notman, 1920; Wolcott, 1924, 1986), Isabela (Hoffman, in U.S.N.M.), Jayuya (Oakley, in U.S.N.M.), Bayamon (Mills, in U.S.N.M.), Guanica (Blackwelder station 855A; Darling- ton, in M.C.Z. and U.S.N.M.), Lake Cartagena (Blackwelder station 46A), Maricao (Blackwelder station 47A). St. Croix: (Fauvel, 1901; Blackwelder stations 333, 341, 344). Guadeloupe: (Fauvel, 1901). Martinique: (Fauvel, 1901). St. Vincent: (Fauvel, 1901). Grenada: (Fauvel, 1901). South America: BrAziL, ARGENTINA (Fauvel, 1901). Central America: GUATEMALA, NICARAGUA, PANAMA (Sharp, 1887); Mexico, GUATEMALA, NICARAGUA, PANAMA (Fauvel, 1091); Mexico Villada, 1901). North America: Fiorina, Lourstana (LeConte, 1877; Sharp, 1887) ; CaLirornira, Fioripa, LOUISIANA, TEXAS (Fauvel, 1901) ; Texas (Casey, 1889b, as teranus) ; FLoripA (Schwarz, 1878). Hawaii: (Fauvel, 1901; Sharp, 1880, as senilis). Specimens exwamined.—I have seen 34 specimens in the United States National Museum, 35 from the Museum of Comparative Zoology, and 73 collected by me during 1935-37. Remarks.—I believe this species to be properly identified although specimens labeled fu/vipes in the British Museum really belong to correctus or one of the other species. A specimen from Sr. Bierig shows that his identification of this species is the same as mine. My specimens were taken from along the sandy banks of streams, on the muddy banks of ponds, from cow dung, and flying at dusk. 3. CARPELIMUS SERICEUS (Cameron) Trogophloeus (Boopinus) sericeus CAMERON, 1923, p. 392.—SCHEERPELTz, 1933, p. 1088. Trogophloeus (s. str.) sericews Cameron, SCHEERPEL?TZ, 1933, p. 1088. Trogophloeus sextagonalis Breric, 1935, p. 10, fig. 1. Description.—Black, pronotum and elytra sometimes feebly rufes- cent. Zead with two feeble depressions between the antennal ridges; eyes separated from base by only a ridge. Pronotwm over one-fourth wider than long, irregularly hexagonal, sides without tubercles, the anterior angles very prominent and coarsely tuberculate; disk with two moderate longitudinal depressions; with rather fine and irregular punctures feebly obscured by ground sculpture. Z7ytra with mod- erate punctures of irregular shape and obscured by indefinite ground sculpture. Length 214 to 3 mm. Type locality —Jamaica. Of sextagonalis, Cuba. Types.—Three examples in Dr. Cameron’s collection. Of sextago- nalis, in Bierig collection; and one cotype in the United States Na- tional Museum (No. 52710). 64 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following are the records known to me: Cuba: Caimito del Guayabal (Bierig, 1935; Bierig, in U.S.N.M.), Laguna La Canao (Bierig, 1935), Cayajabos (Bierig, 1985), Rio Almendares (Bierig, 1935.) (All these as sextagonalis.) Jamaica: (Cameron, 1923; Scheerpeltz, 1933), Santa Cruz (Blackwelder station 421), Black River (Blackwelder station 416), May Pen (Blackwelder station 425B). St. Croix: (Blackwelder station 332). Specimens examined.—I have seen three examples in Dr. Cameron’s collection, one in the United States National Museum, and nine collected by me during 1935-87. Remarks.—Although I have not compared directly any Cuban ex- amples of sextagonalis with the type of sericeus, I have no doubt of the identity of these two species, since a “cotype” of sextagonalis received from Sr. Bierig is exactly similar to specimens I had pre- viously compared with the type of serzceus. This species is quite similar to beattyi and fulvipes but seems to be readily distinguishable from them by color and the details of punctation. My specimens were taken from the muddy banks of ponds, from bat guano, and flying at dusk. 4. CARPELIMUS CORRECTUS, new species Description.—Head black, pronotum piceorufous, elytra rufous, abdomen piceocastaneous. Head with two moderate depressions be- tween the antennal ridges; eyes at about half their length from base. Pronotum over one-third wider than long, irregularly hexagonal, sides not tuberculate, the anterior angles prominent and tuberculate; disk with two feeble longitudinal depressions along the median pos- terior ridge; with very dense granular ground sculpture. Hlytra with rather fine punctures very irregular and obscured by the irregular ground sculpture. Length, 214 to 214 mm. Type locality.—St. Lucia, the Roseau River about 6 miles south of Castries, elevation 250 feet. Types.—Holotype and 14 paratypes, U.S.N.M. No. 52431, collected by me on March 29, 1936. Records.—The following are the records known to me: Jamaica: (Hubbard, in U.S.N.M.; Cameron collection), Trinityville (Black- welder station 428), Milk River (Blackwelder station 415), Spanish Town (Blackwelder station 3877), Troy (Blackwelder station 409), Kingston (Darlington, in M.C.Z.). Hispaniola: Harrr (Cameron collection) ; DoMINICAN REPUBLIC, Sanchez (Dar- lington, in M.C.Z.). Puerto Rico: Maricao (Blackwelder station 47A). St, Croix: (Blackwelder station 341). Guadeloupe: Gourbeyre (A.M.N.H.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 65 St. Lucia: (Blackwelder stations 209A, 211A; Cameron collection). Barbados: (Blackwelder stations 193A, 200, 202A). St. Vincent: (Blackwelder station 170; British Museum). Grenada: (British Museum). Trinidad: St. Augustine (Withycombe, in British Museum; Darlington, in M.C.Z.). Specimens examined.—I have seen 33 examples in the British Mu- seum, 14 in the collection of Dr. Cameron, 2 in the American Museum of Natural History, 3 in the Museum of Comparative Zoology, 1 in the United States National Museum, and 34 collected by me during 1935-37. Remarks.—This species has stood under the name fulvipes in col- lections, but that seems to be more properly applied to another, which is distinguished by having the pronotum not opaque. This species probably will be found on all the islands. My specimens were found under stones and sand along streams, in dung, and flying at dusk. 5. CARPELIMUS DARLINGTONI, new species Description.—Piceous, pronotum and elytra somewhat rufescent, abdomen castaneous. Mead with two feeble depressions between the antennal ridges; eyes at about half their length from base. Pronotum three-sevenths wider than long, not distinctly hexagonal because of the curvature of the base and sides; lateral and anterior angles dis- tinctly tuberculate, basal angles very prominent; disk with two dis- tinct long longitudinal impressions and vague depressions laterally ; with very dense granular ground sculpture. Hlytra with punctures not very fine but very indistinct and mixed with irregular ground sculpture. Length, 314 mm. Type locality—Hispaniola, San José de las Matas, Dominican Republic, elevation 1,000 to 2,000 feet. Types —Holotype, Museum of Comparative Zoology, collected by Dr. P. J. Darlington in June 1938. Records.—The following is the only record known to me: Hispaniola: Dominican Repusric, San José de las Matas (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks—This species differs from correctus in having the sides of the pronotum tuberculate as well as in the coarser punctation of the elytra. I have received no record of its habits. 6. CARPELIMUS BEATTYI, new species Description—Head black, pronotum piceous to rufous, elytra piceocastaneous or rufocastaneous, with apical margin distinctly 66 BULLETIN 182, UNITED STATES NATIONAL MUSEUM testaceous, abdomen piceocastaneous. ead with two moderate de- pressions between the antennal ridges; eyes at about one-fourth their length from base. Pronotwm over one-fourth wider than long, ir- regularly hexagonal, sides with a single very blunt tubercle within each posterolateral emargination, the anterior angles prominent and bluntly tuberculate; disk with two very feeble longitudinal impres- sions; with rather fine punctures obscured by indefinite ground sculp- ture. Llytra with rather fine punctures of irregular shape and obscured by indefinite ground sculpture. Length, 214 to 3 mm. Type locality.—St. Croix, Wheel of Fortune Estate, one-half mile east of Frederiksted. Types.—Holotype and 13 paratypes, U.S.N.M. No. 52467, collected by me on November 17, 1986. Records—The following are the records known to me: Hispaniola: Harrr, Diquini (Mann, in M.C.Z.).; DominicANn Repusuic, Rio San Juan (Miller, in U.S.N.M.). St. Croix: (Blackwelder station 332). Guadeloupe: Basse Terre (Blackwelder station 79C). St. Lucia: (Blackwelder stations 209A, 211A). Specimens examined.—I have seen 4 specimens in the United States National Museum in addition to the 14 types, and 1 in the Museum of Comparative Zoology. Remarks.—This species can be distinguished at once by the testace- ous apical margin of the elytra. It is somewhat variable in the strength of the pronotal ridges and depressions. My specimens were taken from bat guano, from beneath rotting fruit, and from under stones and sand along streams. 7. CARPELIMUS CORNUCOPIUS, new species Description —Rufocastaneous, head piceous. Head with two feeble depressions between the abrupt antennal ridges; the eyes at very nearly their length from base; antennae very slender, extending to apex of elytra. Pronotum one-fifth broader than long, distinctly hexagonal; anterior angles scarcely tuberculate, lateral angles rounded, basal angles rather abrupt; disk with two distinct longi- tudinal impressions united near base, and with vague depressions laterally; with dense but not fine punctures distinct; without ground sculpture. Hlytra behind humerus with a series of tubercles and gradually elevated to apical fourth into a spatulate posteriorly di- rected horn; with punctures as on pronotum but a little less dense; without sculpture. Abdomen with tergites 7 to 8 each with a pair of short abrupt longitudinal carinae. Length, 324 mm. Type locality —Grenada, 6 miles southeast of Gouyave on road to Grenville. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 67 Types —Holotype in the United States National Museum, No. 52356, collected on January 24, 1936. Records.—The following is the only record known to me: Grenada: (Blackwelder station 155). Specimens examined.—I have seen only the holotype. Remarks.—This species may be recognized at once by the unique elevations of the elytra and tergites, as well as by the shape of the pronotum and the form of the antennae. I have some doubts that it is a true Carpelimus but can find no obvious distinguishing feature and know of no other genus to which it might be referred. The type was found along the edge of a stream. 8. CARPELIMUS SMITHI (Bernhauer) Trogophloeus smithi BERNHAUER, 1908, p. 229—BeRNHAUER and ScHusert, 1911, p. 104. Trogophloeus (Carpalimus) smithi CAMERON, 1918b, p. 323 (as new species).— Lene and MurTcH er, 1914, p. 403.—SCHEERPELTZ, 1933, p. 1082. Trogophlocus (Carpalimus) smithianus SCHEERPELTZ, 1933, p. 1082 (as new name for smithi Cameron). Trogophloeus croceipes Birra, 1935a, p. 9 (not Fauvel). Description —Black, pronotum and elytra feebly rufescent. Head with two feeble depressions between the antennal ridges; eyes at nearly half their length from base. Pronotwm about one-fourth wider than long, anterior angles rounded; disk with a strong arcuate transverse basal depression and two rather indefinite longitudinal ones; punc- tures coarse and moderately dense; without distinct ground sculpture except in the basal groove. Llytra a little more coarsely punctate than pronotum and somewhat obscured by indefinite ground sculpture. Length, 214 to 344 mm. Type locality —“Grenada (leg. Smith), Bolivien, Argentinien (Buenos Aires, Resistencia).” Of smthi Cameron, Grenada, Mount Gay Estate, Leeward side. Types-—Presumably in the collection of Dr. Max Bernhauer. Of smithi Cameron, in the British Museum, and one “cotype” in the United States National Museum (No. 50879). Records—The following are the records known to me: Cuba: Aspiro (Bierig, in U.S.N.M., as croceipes), Guanabo (Bierig, in U.S.N.M., as croceipes), Hongolosongo, Oriente (Darlington, in M.C.Z.), Soledad (Darlington, in M.C.Z.). Hispaniola: Hart, Port-au-Prince (Darlington, in M.C.Z. and U.S.N.M.). Grenada: (Bernhauer, 1908; Cameron, 1918; Scheerpeltz, 1933; British Museum ; U.S.N.M.). Trinidad: St. Augustine (Darlington in M.C.Z. and U.S.N.M.). South America: BoriviA, ARGENTINA (Bernhauer, 1908). 68 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have examined 78 examples (including the type) in the British Museum, 5 in the United States National Museum, and 5 from the Museum of Comparative Zoology. Remarks.—This species may be distinguished by its complete basal pronotal depression and the punctation of the pronotum and elytra. Although it is impossible for me to be certain, it seems highly prob- able that the Bernhauer and the Cameron Grenada species are identi- cal, being from the same lot. If the holotype be one of the South American specimens, then our species may be different and would be called smithianus Scheerpeltz. Specimens of this species were sent by Sr. Bierig under the name croceipes. 'They are certainly not the croceipes of Fauvel, and I am unable to distinguish them from smtht of Cameron or the examples from Haiti and Trinidad. I find no record of the habits of this species. 9. CARPELIMUS WEBER], new species Description.—Piceous, pronotum and base of elytra rufescent, ab- domen castaneous. Head with two moderate depressions between the antennal ridges; eyes at about half their length from base. Pronotum one-third wider than long; anterior angles rounded; disk with a strong arcuate transverse basal depression, the two very short longi- tudinal depressions uniting with each other but separated from the basal one; punctures moderately fine and dense and somewhat ob- secured by indefinite ground sculpture. Hlytra much more coarsely punctate than pronotum, and with traces of ground sculpture. Length, 2 mm. Type locality —Trinidad, base of Galeota Point. Types.—H olotype, Museum of Comparative Zoology, and one para- type, United States National Museum No. 52482, collected by N. A. Weber, in September (type) and March 1935. Records.—The following are the records known to me: Trinidad: Galeota Point (Weber, in M.C.Z.), Maracas Falls (Weber, in U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species is very similar to smthi except for the sculpture of the pronotum. I have received no record of its habits. 10. CARPELIMUS HISPANIOLUS, new species Description—Piceous. Head with two moderate depressions be- tween the antennal ridges; eyes separated from base by about one- third their length. Pronckn, one-fourth wider than long; sides and angles not tuberculate; sides distinctly emarginate in posterior half; disk with a large arcuate transverse basal depression and vague traces MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 69 of two short longitudinal ones; with obsolescent punctures, a little more distinct basally, but very shallow ; without distinct ground sculp- ture. lytra with very coarse punctures separated by one-half their diameter or less by fairly flat intervals; without distinct ground sculp- ture. Length, 3 to 314 mm. Type locality —Haiti, northeastern foothills of Morne La Hotte, elevation 3,000 to 4,000 feet. Types.—Holotype and one paratype in Museum of Comparative Zoology, one paratype in the United States National Museum (No. 52483), collected by Dr. P. J. Darlington on October 12, 1984. Records.—The following is the only record known to me: Hispaniola: Haiti, Morne La Hotte (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This species is readily distinguished from tmpunctus and obsolescens by the size of the elytral punctures. It is also dis- tinct from obsolescens by its obsolete pronotal depressions, and from ampuncius by the proportions of the pronotum and its size. I have received no record of the habits of this species. 11. CARPELIMUS IMPUNCTUS, new species Description.—Black. Head with two moderate depressions be- tween the antennal ridges; eyes separated from base by about half their length. Pronotwm one-fifth wider than long, sides and anterior angles not tuberculate; disk with a large arcuate transverse basal depression and two very feeble longitudinal ones united with it; with- out distinct punctures or sculpture but minutely and irregularly punctulate. Hlytra moderately coarsely and rather densely punctate; without distinct ground sculpture. Length, 214 to 234 mm. Type locality —Jamaica. Types—Holotype and four paratypes, U.S.N.M. No. 52468, col- lected by H. G. Hubbard. Records.—The following is the only record known to me: Jamaica: (Hubbard, in U.S.N.M.), Specimens examined—Ii have seen only the five types. Remarks.—This species can be distinguished from all other West Indian species of Carpelimus by its virtually impunctate pronotum and feeble longitudinal impressions. I have no record of its habits, 12. CARPELIMUS OBSOLESCENS, new species Description.—Black, pronotum and elytra more or less rufescent. Head with two very feeble depressions between the antennal ridges, the latter very small; eyes separated from base by less than half their length. Pronotum one-eighth wider than long; sides and angles not 70 BULLETIN 182, UNITED STATES NATIONAL MUSEUM tuberculate; disk with a large transverse arcuate basal impression and two feeble longitudinal ones united with it and at middle with each other; with moderate punctures only slightly obscured by vague ground sculpture. lytra with moderately coarse punctures some- what obscured by the irregular ground sculpture. Length, 214 to 314 mm. Type locality.—Hispaniola, Ennery, Haiti, near an elevation of 1,000 feet. Types.—Holotype and two paratypes in the Museum of Compara- tive Zoology; three paratypes in the United States National Museum (No. 52484); all collected by Dr. P. J. Darlington on September 6-11, 1934. Records.—TYhe following are the records known to me: Hispaniola: Harr1, Ennery (Darlington, in M.C.Z and U.S.N.M.), Port-au- Prince (Darlington, in M.C.Z.). Specimens examined.—\ have seen only the six types. Remarks.—This species differs considerably from ipunctus in the punctation of the pronotum and elytra as well as by its very much reduced antennal ridges. T have no record of its habits. 13. CARPELIMUS BORINQUENSIS, new species Description.—Piceous, pronotum and elytra more or less rufescent, abdomen castaneous. ead with two feeble depressions between the antennal ridges; eyes separated from base by about one-half their length. Pronotwm one-fourth wider than long; sides and angles not tuberculate; sides behind middle distinctly emarginate; disk with a large arcuate transverse basal impression, sometimes partially inter- rupted at middle, and two short longitudinal depressions deeply united to form a transverse median groove; without distinct punctures (except traces at base) but with minute punctulae throughout; without ground sculpture. LZlytra with moderately coarse punctures, but no distinct ground sculpture. Length, 214 to 244 mm. Type locality—Puerto Rico, Maricao Forest, elevation 2,000 to 3,000 feet. Ty pes.—Holotype and five paratypes in the Museum of Comparative Zoology; four paratypes in the United States National Museum (No. 52485); collected by Dr. P. J. Darlington in May 1938. Records.—The following are the records known to me: Puerto Rico: Maricao (Darlington, in M.C.Z. and U.S.N.M.), El Yunque, Dar- lington, in M.C.Z. and U. S. N. M.). Specimens examined.—tI have seen only the 10 types. Remarks.—This species is distinct from all others in this group by the deep median groove, which is as strong as the basal one. Ihave received no record of its habits. “I — MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 14. CARPELIMUS DANFORTHI, new species Description.—Black. Head with two very feeble depressions be- tween the antennal ridges; eyes not distinctly separated from base. Pronotum one-sixth wider than long, all sides and angles rounded ; disk with feeble traces of two longitudinal depressions; with moderate punctures not very dense but almost completely obscured by dense and distinct ground sculpture. Llytra coarsely punctate, the intervals vaguely rugose; with rather irregular ground sculpture. Length, 2 to 234 mm. Type locality —Puerto Rico, south side of Guanica Lake. Types.—Holotype and one paratype, U.S.N.M. No. 52469, collected by me on January 5, 1937. Records.—The following are the records known to me: Jamaica: Kingston (U.S.N.M.). Puerto Rico: Guanica (Blackwelder station 355A), Ensenada (A.M.N.H.). Specimens examined.—I have seen the two types, three other ex- amples in the United States National Museum, and two specimens in the American Museum of Natural History. Remarks —This species may be distinguished among those with coarsely punctate elytra by the strong ground sculpture of the pro- notum which obscures the punctures. The paratype is quite pale in color but does not differ structurally from the type. It is apparently immature in regard to the color. The Jamaica examples have the pronotal sculpture somewhat less distinct but do not seem to differ otherwise. The types were taken from under rubbish on the muddy bank of the lake; the Jamaican examples were found abundantly in green- houses. 15. CARPELIMUS CUBENSIS (Bierig) Trogophloeus cubensis Breria, 1935a, p. 15, 16, fig. 8. Description.—Black. Head with two moderate depressions between the antennal ridges; eyes at about two-thirds their length from base. Pronotum one-fourth wider than long, angles rounded; disk with two feeble longitudinal depressions joining the basal foveae; without very distinct punctation but granularly sculptured. lytra with moderate punctures very slightly obscured by traces of ground sculpture. Length, 114 to 134 mm. Type locality —Cuba. Types.—tIn collection of Alexander Bierig. One “cotype” in the United States National Museum (No. 52705). Records.—The following are the records known to me: Cuba: Rio Almendares (Bierig, 1935), Playa de Jaimanitas (Bierig, 1935), Valle del Yumuri (Bierig, 1935), Rangel (Bierig, 1985), Sierra Bonilla (Bierig, in U.S.N.M.), Sierra del Rosario (Bierig, in U.S.N.M.). Jamaica: Spanish Town (Blackwelder station 377). 12 | BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen two examples in the United States National Museum and one collected by Chapin and Blackwelder in February 1937. Remarks.—This species is most readily distinguished by its deep black color. The Jamaican example does not agree in every detail but presents no characters sufficient to separate it. The Jamaican example was taken flying at dusk. 16. CARPELIMUS IMITATOR (Bierig) Trogophloeus imitator Breric, 1935a, p. 12, 18, 14, 15, fig. 6. Description.—Black, pronotum sometimes rufescent, elytra piceo- castaneous with apex narrowly paler. Head with two feeble impres- sions between the antennal ridges; eyes separated from base by nearly half their length. Pronotwm over one-fourth wider than long, the angles rounded; disk with two rather feeble longitudinal impres- sions posteriorly; punctures not very fine but somewhat obscured by indistinct ground sculpture. “lytra with moderately large punc- tures more or less obscured by indefinite ground sculpture. Length, 14% to 134 mm. Type locality —Cuba, “Aspiro (al pie de la Sierra del Rosario).” Types.—tIn collection of Alexander Bierig. Two “cotypes” in the United States National Museum (No. 52708). Records—The following are the records known to me: Cuba: Aspiro (Bierig, 1935), Rio Almendares (Bierig, in U.S.N.M.), Cayajabos (Bierig, in U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.). Jamaica: Santa Cruz (Blackwelder station 421), Spanish Town (Blackwelder station 377), Black River (Blackwelder station 416), Trinityville (Black- welder station 428), Gordon Town (Blackwelder station 868B), Kingston (Blackwelder station 885), Moneague (Blackwelder station 875). Puerto Rico: Maricao (Blackwelder stations 47A, 47B), Mayagiiez (Blackwelder station 50A), Jayuya (Oakley, in U.S.N.M.), Utuado (Busck, in U.S.N.M.), Guaniea (Darlington, in M.C.Z.). St. Croix: (Blackwelder stations 333, 341, 344). Guadeloupe: Basse Terre (Blackwelder stations 75, 79B; A.M.N.H.). Specimens examined.—I have seen 2 examples from the Bierig col- lection and 11 others in the United States National Museum, 17 in the American Museum of National History, 1 in the Museum of Com- parative Zoology, and 53 collected by Chapin and Blackwelder dur- ing 1935-37. Remarks.—This species is rather feebly set off by the pale elytral apex. The above description was taken from one of the “cotypes” in the United States National Museum. My specimens were taken from under dung, rotten mangoes, rot- ten guavas, and fresh chips, from the muddy banks of ponds and streams, and flying at dusk. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE (3 17. CARPELIMUS VARICORNIS (Bernhauer) Trogophloeus varicornis BERNHAUER, 1904a, p. 24.—BrrNHAUER and ScHUBERT, 1911, p. 105.— LENG and Murcuter, 1914, p. 403. Trogophtoeus (Taenosoma) laetipennis CAMERON, 1923, p. 395.—ScHEERPELTz, 1938, p. 1085. Trogophioeus (s. str.) laetipennis Cameron, SCHEERPELTZ, 1933, p. 1085. Description.—Piceous, pronotum rufopiceous, elytra testaceous. Head with two moderate depressions between the antennal ridges; eyes separated by about half their length from base. Pronotum one- fourth wider than long, the angles rounded; disk with two feeble lon- gitudinal depressions; punctures moderate but obscured by rather indefinite ground sculpture. lytra moderately coarsely punctate, without very distinct ground sculpture. Length, 114 mm. Type locality—Grenada, Vendome Estate, Leeward side. Of laetipennis, Grenada, Mount Gay Estate, Leeward side. Types.—Probably in the British Museum, although Bernhauer made no mention of them and I did not find them there. Of Jaeti- pennis, in the British Museum. Records.—The following are the records known to me: Cuba: Cauto El Cristo, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Savana La Mar, Oriente (Darling- ton, in M.C.Z. and U.S.N.M.), Maisi, Oriente (Darlington, in M.C.Z.). Hispaniola: Harri, Port-au-Prince (Darlington, in M.C.Z.). Grenada: (Bernhauer, 1904; Leng and Mutchler, 1914; Cameron, 1923, as laetipennis). Trinidad: Galeota Point (Weber, in M.C.Z.), St. Augustine (Weber, in M.C.Z.), Nariva Swamp (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen four examples (including the type of daetipennis) in the British Museum. One of these was bor- rowed for further study. Also 21 examples from the Museum of Comparative Zoology. Remarks.—This is one of the species that is very difficult to place ina key. The characters used above appear to be sufficient to set it apart. There seems to be little doubt that the types of varicornis and daetipennis came from the same material, although not from ex- actly the same locality. One of the other examples of laetipennis came from Vendome Estate where the type of varicornis was found. I find no record of the habits of this species. 18. CARPELIMUS ARIDUS (DuVal) Trogophloeus aridus DuVAt, 1857, p. 43.—Fauven, 1863, p. 439.—Suarp, 1887, p. 698.—GUNDLACH, 1893, p. 800—BrrNnHaver, 1910, p. 856.—BrRNHAUER and ScHusert, 1911, p. 97.—Lene and Murcuter, 1914, p. 403; 1917, p. 198. — Wo corr, 1924, p. 177; 1936, p. 198.—Brrrie, 1935a, p. 9. Description.—Black, pronotum and elytra sometimes more or less rufescent. Head with two rather feeble depressions between the 449008—42——-6 74 BULLETIN 182, UNITED STATES NATIONAL MUSEUM antennal ridges; eyes separated from base by not over one-sixth of their length. Pronotum almost one-half wider than long, sides and angles rounded; disk with two distinct longitudinal depressions unit- ing with the two halves of the transbasal depression; with moderately large punctures not or scarcely obscured by ground sculpture. Elytra more coarsely punctate than the head, the intervals vaguely rugose, without distinct ground sculpture. Length, 2 to 214 mm. Type locality—Cuba. Types.—The type should be in the Sharp collection in the British Museum, but I did not find it. Records —The following are the records known to me: Cuba: (DuVal, 1857; Fauvel, 1863; Sharp, 1887; Gundlach, 1893; Bernhauer and Schubert, 1911; Leng and Mutchler, 1914; Bierig, 1985; Cameron col- lection), La Coronella (Bierig, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Cameron collection), Kingston (Blackwelder station 379), Spanish Town (Blackwelder station 377), Balaclava (Blackwelder station 399A), Trinityville (Blackwelder station 428), May Pen (Blackwelder station 425B), Ocho Rios (Darlington, in M.C.Z.). Hispaniola: Hartt (Cameron collection), Bayeux (Hoffman, in U.S.N.M.), Port- au-Prince (Darlington, in M.C.Z.), Ennery (Darlington, in M.C.Z. and U.S.N.M.); DominicAN Repustic, Rio San Juan (Miller, in U.S.N.M.), Sanchez (A.M.N.H.; Darlington, in M.C.Z. and U.S.N.M.), Barahona (Dar- lington, in M.C.Z. and U.S.N.M.). Puerto Rico: (Gundlach, 1893; Leng and Mutchler, 1914; Wolcott, 1924, 1936). Guanica (Blackwelder station 3554; Darlington, in M.C.Z. and U.S.N.M.). St. Croix: (Blackwelder station 341). Barbados: (British Museum; Blackwelder station 198B). St. Vincent: (Blackwelder station 170; British Museum, as memnonius). Grenada: (British Museum, as memnonius). Trinidad: St. Augustine (Darlington, in M.C.Z.). Specimens examined.—\I have seen 45 examples in the British Museum, 8 in Dr. Cameron’s collection, 14 from the Museum of Com- parative Zoology, 4 in the United States National Museum, 1 in the American Museum of Natural History, and 43 collected by me during 1935-37. Remarks.—Part of the specimens in the British Museum were placed under the name 7'rogophloeus memnonius Erichson. This species is otherwise known only from the Mediterranean area and appears to me to be distinct in details of punctation. My specimens were taken from under stones and on mud banks along streams and ponds, from partly submerged debris in a stream, and flying at dusk. 19. CARPELIMUS AEOLUS, new species Description.—Head black, pronotum rufopiceous, elytra light rufocastaneous, abdomen piceous. ZZead with two moderate depres- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 15 sions between the antennal ridges; eyes scarcely at all separated from base. Pronotwm about one-fifth wider than long, the angles rounded; disk with two feeble longitudinal depressions posteriorly ; with punctures rather fine and dense, not distinct but not definitely obscured by ground sculpture. Zlytra with rather fine but irregular punctures; without distinct ground sculpture. Length, 1144 mm. Type locality—Jamaica, between Spanish Town and Kingston. Types.—Holotype and two paratypes, U.S.N.M. No. 52450, col- lected by Chapin and Blackwelder on February 2, 1937. Records.—The following are the records known to me: Jamaica: Spanish Town (Blackwelder station 377), Santa Cruz (Blackwelder station 421), Kingston (Darlington, in M.C.Z. and U.S.N.M.). Specimens exwamined.—\ have seen only the three types and two specimens from the Museum of Comparative Zoology. Remarks.—This species may be distinguished from swbtilior by its normally stout form and prominent eyes. The types were taken flying at dusk. 20. CARPELIMUS PHAIOS, new species Description.—Head black, pronotum and elytra rufopiceous, abdo- men piceous. Head with two rather feeble depressions between the antennal ridges; eyes separated from base by nearly two-thirds their length. Pronotum about one-fifth wider than long, sides and angles rounded; disk with two feeble longitudinal depressions united with basal foveae; punctures moderate, very distinct, with shining inter- vals. LHlytra very coarsely punctate; with traces of ground sculp- ture. Length, 2 mm. Type locality —Jamaica, between Bath in St. Thomas and Lower White Horses by way of Sunning Hill, Trinityville, and Cedar Hill. Types.—Holotype and one paratype, U.S.N.M. No. 52448, collected by Chapin and Blackwelder on February 28, 1937. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Vinales (A.M.N.H.). Jamaica: Trinityville (Blackwelder station 428), Moneague (Blackwelder sta- tion 435). Specimens examined.—\ have seen the two types and one other specimen in the United States National Museum, and two in the American Museum of Natural History. Remarks.—This species may be distinguished from its neighbors by the proportions of the pronotum and the details of punctation as given in the key. The single example from Cuba is paler than the types but is otherwise the same. Our specimens were taken flying at dusk and by sweeping grass in a meadow, 76 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 21. CARPELIMUS SCROBIGER (Cameron) Trogophloeus (Boopinus) scrobiger CAMERON, 1923, p. 393.—ScHEERPELTz, 1933, p. 1088. Trogophloeus (s. str.) serobiger Cameron, SCHEERPELTZ, 1933, p. 1088. Description—Black. Head with two moderate depressions between the antennal ridges; eyes scarcely separated from base. Pronotwm nearly one-half wider than long, angles rounded; disk with two feeble longitudinal depressions joining the separated halves of the arcuate basal depression; with fine punctures separated by fairly smooth shining intervals. /lytra with punctures a little larger than on pronotum but rather angular, intervals with vague ground sculpture. Length, 1144 to 2 mm. Type locality —Grenada, St. Johns River, Leeward side. Types.—In the British Museum. Records—The following are the records known to me: Jamaica: Bath in St. Thomas (Blackwelder station 392B), Trinityville (Black- welder station 428). Puerto Rico: Aguadilla (Busck, in U.S.N.M.), Utuado (Busck, in U.S.N.M.). Vieques Island: (Busck, in U.S.N.M.). St. Vincent: (Blackwelder station 176A). Grenada: (Cameron, 1923; Scheerpeltz, 1933; British Museum; Cameron col- lection; U.S.N.M.). Specimens examined.—I have seen 15 examples in the British Museum, 4 in Dr. Cameron’s collection, 4 in the United States Na- tional Museum, and 18 collected by me during 1935-37. Remarks.—The description was drawn from a St. Vincent example. This is one of the smallest of our species having the punctures of the pronotum discrete and not obscured by ground sculpture. My examples were taken from under stones along the edge of streams and flying at dusk. 22. CARPELIMUS PETOMUS, new species Description—Head black, pronotum and elytra piceorufous, abdo- men black with apex rufescent. Head with two feeble depressions between the antennal ridges; eyes scarcely separated from base. Pro- notum one-fifth wider than long, angles rounded; disk with two feeble rounded depressions along middle; with rather fine punctures not distinctly separate yet not obscured by ground sculpture. L’lytra finely and irregularly punctate, and with indefinite ground sculpture. Leneth, 1144 to 2 mm. Type locality —St. Thomas, Charlotte Amalie. Types.—Holotype and seven paratypes, U.S.N.M. No. 52470, col- lected by me on October 21, 1935. One paratype has been deposited in the British Museum and one in the collection of Dr. Cameron. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE CE Records.—The following are the records known to me: Jamaica: Milk River (Blackwelder station 415), Spanish Town (Blackwelder station 377). Puerto Rico: Guanica (Blackwelder station 357A). Vieques Island: (Leonard, in U.S.N.M.). St. Thomas: (Blackwelder station 68). Antigua: (Blackwelder station 295). Specimens examined.—I have seen the 11 types and 8 other speci- mens in the United States National Museum. Remarks.—This species is one of the group that differ in details of sculpture and in arrangement of colors. ‘These are separated as shown in the key. My specimens were taken from under seaweed on the beach or flying at dusk or to light. 23. CARPELIMUS FLAVIPES (Erichson) Trogophloeus flavipes ErtcHson, 1840, p. 808.—DvuVat, 1857, p. 44.—FAvvEL, 1863, p. 441.—BrRNHAUER and SCHUBERT, 1911, p. 99.—LENG and MUTCHLER, 1914, p. 403. (Not Motschulsky, 1861.) Trogophloeus hiemalis Bieria, 1935a, p. 18, 14, 15, fig. 3. Trogophioeus flavibasis Bieria, 1935a, p. 18, fig. 4. Description.—Black, pronotum rufopiceous, elytra rufocastaneous. Head with two feeble depressions between the antennal ridges; eyes separated from base by less than one-fourth their length. Pronotwm about one-fourth wider than long, anterior angles rounded; disk with two feeble short depressions not united with basal foveae: with rather fine punctures a little indistinct but without distinct ground sculpture. Llytra not very finely but rather irregularly punctate, the punctures angular and somewhat obscured by indefinite ground sculpture. Length, 114 to 2 mm. Type locality —*Americae meridionalis insulis St. Johannis et St. Thomae.” Of hiemalis, Cuba; of flavibasis, Cuba, Sierra del Rosario, vicinity of Cayajabos. Types.—HKither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of Azemalis and flavibasis, in collection of Alex- ander Bierig (one “cotype” of each in the United States National Museum, Nos. 52701 and 52704). Records.—The following are the records known to me: Bahamas Islands: Nassau (Knab, in U.S.N.M.). Cuba: (Fauvel, 1863; DuVal, 1857; Leng and Mutchler, 1914), Playa de Mari- anao (Bierig, 1935), Aspiro (Bierig, 1935; U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Cayajabos (Bierig, 1935, as flavibasis; U.S.N.M.). Jamaica: (Hubbard, in U.S.N.M.; Cameron collection), Santa Cruz (Black- welder station 421), Spanish Town (Blackwelder station 377), Bath in St. Thomas (Blackwelder station 392H). 78 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Hispaniola: Hartt (Cameron collection). Puerto Rico: Mayagiiez (Blackwelder station 50D), Guanica (Blackwelder station 355A). St. Thomas: (Erichson, 1840; DuVal, 1857; Fauvel, 1863; Leng and Mutchler, 1914). St. A (Erichson, 1840; DuVal, 1857; Fauvel, 1863; Leng and Mutchler, 1914). St. Croix: (Blackwelder station 344). Guadeloupe: (A.M.N.H.). St. Lucia: (Cameron collection). Barbados: (Blackwelder station 200). St. Vincent: (Blackwelder stations 166, 176A; British Museum). Grenada: (H. H. Smith, in British Museum and U.S.N.M.). Central America: Mexico (Bernhauer and Schubert, 1911). Specimens examined.—I have seen 670 examples in the British Museum, 14 in Dr. Cameron’s collection, 17 in the United States National Museum (including “cotypes” of hiemalis and flavibasis), 2 in the American Museum of Natural History, and 14 collected by me during 1935-87. Remarks.—This widespread species is somewhat variable in small details but seems to be distinguishable by the characters used in the key. I am not able to separate the two Cuban species described by Sr. Bierig. My specimens were taken from under stones and trash, along streams and ponds, and flying at dusk. 24. CARPELIMUS SUBTILIOR (Cameron) Trogophloeus (Taenosoma) subtilior CAMERON, 1923, p. 396.—SCHEERPELTZ, 1933, p. 1088. Trogophloeus (s. str.) subtilior Cameron, SCHEERPELTZ, 1933, p. 1088. Description—Head black or piceous, pronotum and elytra light rufocastaneous, abdomen somewhat picescent. Head with two feeble depressions between the antennal ridges, and with a feeble fovea on the vertex; eyes separated from base by about their length. Pronotum scarcely wider than long, the angles rounded; disk with two feeble longitudinal depressions uniting with the basal foveae; punctures not distinct, but with rather dense coarse sculpture. Elytra finely punctate; without distinct ground sculpture. Length, iY, to 134 mm. Type locality—Grenada, Balthazar. Ll'ypes.—Unique type in the British Museum. Records.—The following are the records known to me: Jamaica: Spanish Town (Blackwelder station 377), Fern Gully (Blackweider station 374). Grenada: (Cameron, 1923). Specimens examined.—I have seen the type in the British Museum and three examples collected by Chapin and Blackwelder in 1937. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 79 Remarks.—This species may be recognized by its very slender form and not prominent eyes. The above description was drawn from a Fern Gully example. My specimens were taken flying at dusk. 25. CARPELIMUS PROLIXUS (Bierig) Trogophloeus prolizus Briere, 1935a, pp. 12, 18, fig. 2. Description.—Black, elytra rufocastaneous. JZead with two feeble depressions between the antennal ridges; eyes scarcely separated from base. Pronotwm about one-fourth wider than long, the angles rounded; disk with two rather feeble longitudinal depressions unit- ing with the basal foveae; punctures rather fine but obscured by indefinite ground sculpture. lytra with punctures moderately fine and somewhat angular; with indefinite ground sculpture. Length, 134 mm. Type locality—Cuba. Types.—tin collection of Alexander Bierig. One “cotype” in the United States National Museum (No. 52707). Records.—The following are the records known to me: Cuba: Rio Almendares (Bierig, 1985; Bierig, in U.S.N.M.), Laguna La Canoa (Bierig, 1985), Playa de Marianao (Bierig, 1935), Aspiro (Bierig, 1935). Specimens examined.—I have seen only the “cotype” in the United States National museum. Remarks.—This species can be distinguished from those near it in the key by its slender form. I find no record of its habits. 26. CARPELIMUS HAPLOMUS, new species Trogophloeus hatuey Brieric, MS. Description—Black, the elytra very slightly rufescent. Head with two very feeble depressions between the antennal ridges; base not strongly marked but eyes not much separated from it. Prono- tum over one-third wider than long, angles rounded; disk without trace of longitudinal depressions but with feeble basal foveae; sculp- ture very fine and very dense, surface entirely opaque. lytra with rather coarse punctures separated by irregular intervals; with very feeble and indefinite ground sculpture. Length, 1144 mm. Type locality—St. Croix, Frederiksted. Types —Holotype, U. S. N. M. No. 52451, collected by me during November 1936. Records.—The following are the records known to me: Cuba: Rio Almendares (Bierig, in U.S.N.M., as hatucy). St, Croix: (Blackwelder station 838). 80 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen only the type and one example from Cuba labeled 7. hatuey Bierig in the United States National Museum. Remarks.—This species is rather distinct according to the char- acters employed in the key. The Cuban example, which is a “co- type” of hatwey, is identical with the type except for the presence of an indistinct testaceous spot on one elytron. The type was caught flying to light at night. 27. CARPELIMUS TUNAPUNAE, new species Description.—Black, the pronotum rufescent, the elytra piceorufous with the apical third indefinitely testaceous. Head with scarcely a trace of depressions between the antennal ridges; eyes separated from base by one-fourth their Jength, constriction abrupt but feeble. Pronotum one-fourth wider than long, the angles rounded; disk without trace of longitudinal depressions or basal foveae; sculpture very fine and very dense, granular, surface opaque. lytra with moderate punctures obscured by irregular intervals and traces of ground sculpture which is more evident basally. Length, 114 to 124 mm. Type locality —Trinidad, foothills north of Tunapuna. Types.—Holotype and two paratypes in the Museum of Compara- tive Zoology; two paratypes in the United States National Museum (No. 52482) ; collected on May 25, 1935, by Dr. N. A. Weber. Ona paratype collected 12 miles north of Arima. Fecords.—The following are the records known to me: Trinidad: Tunapuna (Weber, in M.C.Z. and U.S.N.M.), Arima (Weber, in M.C.Z.). Specimens examined.—I have seen only the five types. Remarks.—This species is similar to haplomus in most characters. It differs in the shape of the head behind the eyes and in the coloration. I have received no record of its habits. 28. CARPELIMUS DISCIPENNIS (Bierig) Trogophloeus discipennis Bieric, 1935a, p. 16, fig. 7. Description.—Black, elytra striped with testaceous from humerus to apex and along apical margin. Head with two very feeble depres- sions between the antennal ridges; separated from base by about their length or a little more. Pronotwm one-fourth wider than long, angles rounded; disk without distinct depressions; very densely granulosely sculptured and opaque. lytra with punctures scarcely evident because of the irregularity of the intervals; with some ground sculpture especially near base. Length, 114 to 134 mm. CO b= MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Type locality—Cuba. Types.—tIn the collection of Alexander Bierig. One “cotype” in the United States National Museum (No. 52702). Records.—The following are the records known to me: Cuba: Rio Almendares (Bierig, 1985; U.S.N.M.), Playa de Marianao (Bierig, 1985), Cayajabos (Bierig, 1935). Jamaica: Fern Gully (Blackwelder station 874), Trinityville (Blackwelder station 428), Moneague (Blackwelder stations 375, 485), Santa Cruz (Black- welder station 421), Milk River (Blackwelder station 415), Gordon Town (Blackwelder station 384), Ocho Rios (Blackwelder station 373). Specimens examined—I have seen the one “cotype” and 41 examples collected by Chapin and Blackwelder in February 1937. Remarks.—This is a species definitely distinguished by its dense sculpture and elytral coloring. I find no difference between the Jamaican and the Cuban examples. Our specimens were taken flying at dusk or by sweeping in a grassy meadow. 29. CARPELIMUS CHAPINI, new species Description.—Black, pronotum and elytra more or Jess rufescent. Head with two very feeble depressions between the antennal ridges; eyes scarcely separated from the feebly defined base. Pronotwm one-fourth wider than long, angles rounded; disk without trace of longitudinal depressions; without distinct punctures but densely and granularly sculptured. Zlytra with moderately coarse . punctures, the intervals with vague ground sculpture. Length, 114 to 144 mm. Type locality—Jamaica, between Spanish Town and Kingston. Types.—Holotype and five paratypes, U.S.N.M. No. 52429, collected by Chapin and Blackwelder on February 2, 1937.. One paratype has been deposited in the British Museum. Records.—The following is the only record known to me: Jamaica: Spanish Town (Blackwelder station 3877). Specimens examined.—\ have seen only the seven types. Remarks.—This species appears to be distinguishable by details of sculpture. It is closest to conformis. The types were taken flying at dusk. 30. CARPELIMUS RULOMUS, new species Description—Piceous. Head with two feeble impressions be- tween the antennal ridges; eyes scarcely separated from feebly de- fined base. Pronotum nearly one-fourth wider than long; angles rounded; disk without trace of longitudinal depressions, but mid- line vaguely elevated at base; with dense and fine punctures appear- ing very much like ground sculpture; surface shining. Llytra 82 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with very fine but irregular punctures; without distinct ground sculpture. Length, 2 mm. Type locality—Dominican Republic, Barahona. Types.—Holotype, Museum of Comparative Zoology, collected by Dr. P. J. Darlington in September 1938. FRecords.—The following is the only record known to me: Hispaniola: DoMINICAN Repuslic, Barahona (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species differs from chapini principally in the very different punctation of the elytra. I have no record of its habits. 31. CARPELIMUS CONFORMIS, new species Trogophloeus conformis Brierta, MS. Description.—Black, the elytra feebly rufescent. Head with two feeble depressions between the antennal ridges; eyes separated from base by less than one-third their length. Pronotum about one-fifth wider than long, angles rounded; disk without trace of longitudinal depressions; with a vague trace of smooth midline; punctures distinct but not dense, intervals densely but not coarsely sculptured. Hlytra with rather coarse punctures, the intervals with very feeble ground sculpture. Length, 1144 mm. Type locality—Cuba, El] Cano, Habana Province. Types—Holotype in the United States National Museum (No. 52706), collected in April 1934 by Sr. Bierig. Records.—The following is the only record known to me: Cuba: El Cano (Bierig, in U.S.N.M.). Specimens examined.—I have seen only the holotype in the United States National Museum. Remarks—This species may be distinguished by the distinct pro- notal punctures among the ground sculpture. I find no record of its habits. 32. CARPELIMUS AEQUALIS (DuVal) Trogophloeus aequalis DUVAL, 1857, p. 44.—FAvVEL, 1868, p. 441.—BERNHAUER and ScHusert, 1911, p. 97.—Lene and Mutcutrr, 1914, p. 403.—Wotcort, 1924, p. 77; 1936, p. 196.—Brerta, 1935a, pp. 9,17. (Not Gundlach, 1893.) Trogophloeus (Taenosoma) funestus CAMERON, 1923, p. 393.—SCHEERPELTZ, 1933, p. 1086. Trogophloeus (s. str.) funestus Cameron, SCHEERPELTZ, 1933, p. 1086. Trogophloeus (Troginus) amplicollis Brrrie, 1935a, p. 17, fig. 9. Description —Black, pronotum and elytra feebly rufescent. Head with two feeble depressions between the antennal ridges; eyes not distinctly separated from base. Pronotum about one-fifth wider than MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 83 long, the angles narrowly rounded; disk without trace of longitudinal impressions but with a narrow smooth midline; with moderate punc- tures rather dense and obscured by indefinite ground sculpture. Elytra with moderately coarse punctures separated by irregular inter- vals which are vaguely sculptured. Length, 2 to 2144 mm. Type locality —Cuba. Of amplicollis, Cuba, Playa de Marianao; of funestus, Jamaica. Types.—Presumably in the collection of Blanche Rancin, Caén, France. Of amplicollis, in collection of Alexander Bierig; of funestus, in collection of Dr. Cameron. One “cotype” of amplicollis is in the United States National Museum (No. 52700). Records.—The following are the records known to me: Bermuda: (Weber, in M.C.Z. and U.S.N.M.). Bahamas Islands: Andros Islands, Mangrove Cay (Mann, in A.M.N.H.). Cuba: (DuVal, 1857; Fauvel, 1863; Leng and Mutchler, 1914), Playa de Marianao (Bierig, 1935, and in U.S.N.M., as amplicollis). Jamaica: (Cameron, 1923, as funestus), Santa Cruz (Blackwelder station 421), Spanish Town (Blackwelder station 3877). Specimens examined.—\ have seen 7 examples (including the type of funestus) in Dr. Cameron’s collection, 2 examples (including a “cotype” of amplicollis) in the United States National Museum, 1 in the American Museum of National History, 2 from the Museum of Comparative Zoology, and 11 collected by Chapin and Blackwelder in February 1937. Remarks.—This is a distinct species, which may be recognized by the smooth midline of the pronotum. It is a rather robust species. Our examples were taken flying at dusk. 33. CARPELIMUS DISSONUS (Bierig) Trogophloeus dissonus BrERIG, 1935a, pp. 11, 12, fig. 5. Description—Black. Head with two moderate depressions between the antennal ridges; eyes scarcely separated from base. Pronotwm about one-fifth wider than long, angles rounded; disk without longi- tudinal depressions but with basal foveae ; not distinctly punctate with rather indefinite sculpture leaving surface shining but not smooth. Elytra with punctures obsolescent, giving an appearance of feeble sculpture. Length, 21/4 mm. Type locality.—Cuba. Types.—In collection of Alexander Bierig. One “cotype” in United States National Museum (No, 52703). Fecords.—The following are the records known to me: Cuba: Habana (Bierig, 1935), Batabano (Bierig, 1935), Playa de Marianao (Bierig, in U.S.N.M.). Specimens examined.—I have seen only the one “eotype” in the United States National Museum. 84 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks—This species may be distinguished from those close to it by its obsolescent punctation, especially that of the elytra. I find no record of its habits. 34. CARPELIMUS TESTACEIPENNIS (Cameron) Trogophloeus (Taenosoma) testaceipennis CAMERON, 1923, p, 895.—SCHEERPELTZ, 1933, p. 1089. Trogophloeus (s. str.) testaceipennis Cameron, SCHEERPELT2, 1933, 1089. Original description—Black, greasy-lustrous, the elytra rufo- testaceous, the base and scutellar region narrowly infuscate. Thorax with discal impressions ill-defined. Abdomen black, shining. An- tennae with the first three joints brownish testaceous. Legs testaceous. “Head densely coriaceous as in 7’. sordidus, mihi; the eyes large and temples short. Antennae with the third joint distinctly shorter than the second, the fourth small, scarcely longer than broad, the fifth larger, a little longer than broad, the sixth to the tenth transverse, gradually increasing in breadth, the penultimate only slightly trans- verse. Thorax rather strongly transverse, the sides gently rounded in front and pretty strongly contracted towards the base; disk with four obsolete impressions; sculpture dense and coriaceous, without distinct puncturation. Elytra a little longer and broader than the thorax, slightly transverse, closely and by no means finely punctured, shining. Abdomen black, shining, very finely and closely punctured in front, more sparingly on the last three segments, pubescence fine, not sericeous, moderately close. Length, 2 mm.” Type locality. —Haiti. Types.—Unique type in collection of Dr. Cameron. Records.—The following is the only record known to me: Hispaniola: Harri (Cameron, 1923; Cameron collection). Specimens examined.—I have seen only the unique type, which I examined very briefly in Dr. Cameron’s collection. Remarks.— “This species is of almost identical build with 7. nigrellus Shp., but differs in the sculpture and colour” (from original description). I find no record of its habits. 35. CARPELIMUS SORDIDUS (Cameron) Trogophloeus (Taenosoma) sordidus CAMERON, 1923, p. 894.—SCHEERPELTZ, 1933, p. 1088. Trogophloeus (s. str.) sordidus Cameron, SCHEERPELTZ, 1933, p. 1088. Original description — ‘Black, moderately shining; the elytra dark brown. Head densely punctured; thorax with the impression on the disk very obsolete. Antennae with the first two joints brownish tes- taceous. Legs testaceous. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 85 “Head without impression on the vertex, closely and densely sculp- tured, appearing to be coriaceous rather than punctured; eyes large, temples short. Antennae with the third point distinctly shorter than the second, the fourth scarcely longer than broad, the fifth as long as broad, larger than the fourth and sixth, sixth to the tenth transverse, gradually increasing in breadth, the penultimate about half as broad again as long. Thorax transverse, widest about the middle, the sides in front gently rounded and narrowed, rather strongly contracted in a nearly straight line behind; disk with four very obsolete impressions; densely coriaceous and with moderate close but obsolete puncturation. Elytra nearly half as long again as, and distinctly broader than the thorax, about as long as broad; rather coarsely and pretty closely punc- tured, finely and sparingly pubescent. Abdomen black, shining, very finely and moderately closely punctured and pubescent, rather more sparingly on the last two or three segments; finely coriaceous. Length. 1.75 mm.” Type locality.—Grand Cayman, Cayman Islands. Types.—Unique type in collection of Dr. Cameron. Records.—The following is the only record known to me: Cayman Islands: Grand Cayman (Cameron, 1923; Cameron collection). Specimens examined.—I have seen no examples of this species. Remarks.—‘Closely allied to 7’. misellus, Shp., but the thorax is more transverse and more strongly contracted behind, the sculpture more coriaceous, the puncturation appearing obsolete, the elytra rather more coarsely punctured, the antennae longer, the penultimate joints less transverse” (from original description). I find no record of its habits. 36. CARPELIMUS DEMMELI (Bierig) Trogophloeus demmeli Brera, 1935a, p. 16. Original description— “Appears undoubtedly most closely related to the previous species (discipennis). However, it differs in the concolorous brownish-black eltyra, the obscuro-ferrugineous legs with the lighter knees and pallid tarsi. The sides of the elytra are more parallel; on the disk of the pronotum, in certain lights, one can see vestiges of the prebasal fossae; and the apical antennal enlarge- ment appears more gradual. For the rest, one does not find differ- ences of sufficient clarity to describe, they being of the same size” (translated). Type locality.—Cuba. Types.—In collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: Habana (Bierig, 1935), Caimito del Guayabal (Bierig, 1935). 86 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen no examples of this species. Remarks.—‘In proof of my gratitude I dedicate this rarity to my estimable friend and colleague, the painter Albert Demmel, who gave it to me, having himself caught it at night on the tablecloth. The second example was found in the mud of a pond” (translated from original description). This species appears to me to be a synonym of discipennis, but in the absence of specimens I retain it as a species not known to me. 87. CARPELIMUS FULVIPENNIS (Fauvel) Trogophloeus fulvipennis FAUVEL, 1863, p. 440.—SHARP, 1887, p. 699.—HAMILTON, 1889, p. 118.—CocKERELL, 1893, p. 8326.—BERNHAUER and SCHUBERT, 1911, p. 98 (as variety of corticinus Gravenhorst).—Lrene and MutTcHtsr, 1914, p. 4038 (as variety).—GowpbrEy, 1926, p. 11.—ScHEERPELTZ, 1933, p. 1083 (as aberra- tion of corticinus Gravenhorst).—Breria, 1985, p. 15. Original description—Not very elongate, broad, subdepressed, black, base of antennae and feet testaceous; pronotum piceous, disk with four longitudinal impressions, sides rotund; elytra fulvorubrous, apex scarcely dilute, base somewhat infuscate, shining, one-third longer than wide; tip of abdomen concolorous. Length, 124 mm. “Of the form of 77. pusillus Er., less elongate, wider; facies and coloration of the preceding (7. rubripennis Fauvel), to which it is probably closely related, but differs in reality in the following points: the size is smaller by half, the body is much more thick-set, wider, not linear, more brilliant, the palpi are testaceous, the antennae are paler, with the two basal segments testaceous. The pronotum is of the size of the head, a little elongated, scarcely one-fourth wider than long; the sides are regularly arcuate, not angulate, with the greatest width at about the center; upon the disk, with four small fossulae united to form a longitudinal impression. The elytra are paler especially at the apex, a little longer. The abdomen is a little broader, unicolorous, the outer margin of the penultimate segment and all of the last appear a little less pale under a certain light. The punctation is finer and denser on the head, pronotum, and abdo- men, stronger and more visible on the elytra. The feet are pale testaceous.” (Translated.) Type locality—Cuba. Types—Presumably in the collection of Blanche Rancin, Caén, France. Records.—The following are the only records known to me: Cuba: (Fauvel, 1863; Sharp, 1887; Cockerell, 1893; Leng and Mutchler, 1914). Jamaica: (Cockerell, 1898; Gowdey, 1926). Specimens examined.—I have seen no examples of this species. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 87 Remarks.—It appears to me unlikely that this species is a variety of T. corticinus (Gravenhorst), but Iam unable to add to the information outlined above. I find no record of its habits, VI. Genus TORRENTOMUS Bierig Torrentomus Breria, 1934e, p. 213. Genotype.—T. torrei Bierig (original designation and monobasic). Diagnosis —Parallel and feebly convex; body with rather dense sculpture; without ocelli: labrum transverse, arcuate; head very strongly constricted behind; pronotum not transversely sulcate; inner apical angles of elytra not broadly rounded; tibiae pubescent. Remarks.—The above characters were taken from Sr. Bierig’s de- scription. In addition to these the new species here described shows the following features: Antennae 11-segmented, filiform; gular sutures united in anterior half; fourth segment of maxillary palpus subulate; anterior coxal cavities confluent and open behind; front coxae conical, prominent; posterior coxae contiguous, transverse ; sternite of first abdominal segment absent, of second complete; tarsi 3-segmented, This genus is very similar to both Oxytelus and Carpelimus, although it is more likely to be confused with the former. I have seen examples only of the new species tenebrosus. KEY TO WEST INDIAN SPECIES OF TORRENTOMUS t Pronotumewigesteat. middle:.s22- 2 ose ae eee aan 1. torrei Pronotum widest) near anterior angles=2>- 2-22) = - == eae 2. tenebrosus 1. TORRENTOMUS TORREI Bierig Torrentomus torrei Brertc, 1934e, p. 214, pl. 11, figs. 1, 3. Description —Ferrugineous, pronotum testaceous, elytra brownish posteriorly, apex of abdominal segments testaceous. ead slightly wider behind eyes, angles rotund; supra-antennal prominences mode- rate; sculpture densely granulose and regular; antennae scarcely thickened externally, second segment articulated with diagonally truncate end of first, labrum nearly semicircular. Pronotum one-half wider than long, as wide as head, sides and base arcuate; widest at middle where sides are feebly angled; sculpture densely granulose but less fine than on head and more coriaceous. Llytra scarcely wider than pronotum. Abdomen with very fine and scarcely per- ceptible sculpture. Length, 114 to 14% mm. (From Bierig.) Type locality—Cuba, Sierra del Grillo, province of Habana. Types.—One example in collection of Alexander Bierig. 88 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following is the only record known to me: Cuba: Sierra del Grillo (Bierig, 1934). Specimens examined.—I have seen no examples of this species. Remarks.—This species is known to me only from the original description. The type was taken by sifting fallen leaves. 2. TORRENTOMUS TENEBROSUS, new species Description.—Castaneous. Head slightly wider behind eyes, angles rotund; constricted part completely set off from rest by a circular sulcus around the neck; clypeal area quadrate, set off by a feebly im- pressed black angulate line; supraantennal prominences rather feeble; with very fine and very dense scaly sculpture throughout ; antennae not at all thickened externally, second segment articulated with diagonally truncate end of first segment; labrum nearly semi- circular but almost emarginate in front; maxillary palpus subulate, third segment large, fourth acicular; gular sutures converging on constricted part, united and broadly Y-shaped in front. Pronotum one-third wider than long, one-twelfth wider than head; sides and base arcuate, hind angles scarcely evident, widest at front; with ir- regular punctate depressions at posterior third of sides, and two large widely separated foveae near the base of disk; with very dense fine scaly sculpture. Hlytra one-fourth wider than pronotum, one- ninth wider than long; sides nearly straight, apical angles scarcely rounded; with dense and fine scaly sculpture. Abdomen feebly sub- muricately sculptured. Length, 214 mm. Type locality—Jamaica, vicinity of Trinityville, parish of St. ‘Thomas. Types.—Holotype and two paratypes in the United States National Museum (No. 52471), collected on February 28, 1937, by Chapin and Blackwelder (one paratype from Troy, parish of Trelawney, February 16, 1937). Records —The following are the records known to me: Jamaica: Trinityville (Blackwelder station 428), Troy (Biackwelder station 409). Specimens examined.—I have seen only the three types. Remarks.—This species is doubtfully referred to Zorrentomus, since that genus is not otherwise represented before me. These specimens were caught flying at dusk. Vil. Genus APOCELLUS Erichson Apocellus EricHson, 1840, p. 812. Ocaleomorpha FLEISCGER, 1921, p. 114. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 89 Genotype —Lathrobium sphaericollis Say=Apocellus sphaericollis (Say) (designated by Duponchel, 1841). Diagnosis.—Body glabrous, robust; integuments smooth; no ocelli; but with distinct pits of dorsal arm of tentorium; gular sutures united anteriorly; antennae 11-segmented ; labrum quadrate; palpi subulate ; anterior coxal cavities confluent, entirely open behind; anterior coxae prominent; posterior coxae contiguous, not expanded laterally under the femora; abdomen strongly margined, first sternite not present; sternite of second segment normal, not carinate; in male, seventh sternite more or less modified; posterior trochanters small; tibiae without rows of spines; tarsi 3-segmented; posterior ones larger than half of the tibia. Remarks.—This is a genus readily distinguished by its facies as well as by structural characters. I have seen 33 examples of this genus from the West Indies. KEY TO WEST INDIAN SPECIES OF APOCELLUS 1. Pronotum and elytra longitudinally strigulose________-_-_-_--_--- 2, planus Pronotum and: elytra, not striguloses22_2=5 = — 2 oe ease se 1. ustulatus 1. APOCELLUS USTULATUS Erichson Apocellus ustulatus EricHson, 1840, p. 818.—LacorpalrgE, 1854, p. 120.—KRAATzZ, 1858a, p. 807.— BERNHAUER and ScHusbertT, 1911, p. 108.—LENe and MuTcH- ER, 1914, p. 404. Description —F lavotestaceous to rufotestaceous, elytra piceotesta- ceous with apex nigropiceous, apex of abdomen nigropiceous. Head one-eighth wider than pronotum, suborbicular, smooth, feebly im- pressed longitudinally on each side in front; supraantennal tubercles large and prominent; antennae longer than head and pronotum, incrassate; labrum rather large, twice as wide as long, rather deeply emarginate, with membranous lobes at anterior angles; gular sutures united from near constriction, Y-shaped anteriorly; third segment of maxillary palpi large, fourth acicular. Pronotum small, one-fifth wider than long; apex one-third wider than base, half as wide as elytra; convex, smooth. Llytra two-fifths longer than pronotum; smooth, shining. Abdomen smooth. AJfale, posterior margin of sev- enth sternite interrupted by an oval membranous cushionlike area; eighth sternite broadly concave, rather deeply triangularly emargi- nate posteriorly. Length, 2 to 22 22 mm. Type locality.—St. ae Types.—Probably either in the Hope Museum, Oxford, or the Zo- ologische Museum, Berlin. Records.—The following are the records known to me: St. John: (Erichson, 1840; Lacordaire, 1854). St. Croix: (Blackwelder stations 331, 344). 449008—42——7 90 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Montserrat: (Blackwelder station 269). Guadeloupe: Basse Terre (Blackwelder station 75; A.M.N.H.). Dominica: (Blackwelder station 242). Grenada: (British Museum). Specimens examined.—l have examined 2 examples in the British Museum, 3 in the American Museum of Natural History, and 21 collected by me in 1935-37. Remarks.—The specimens in the British Museum were placed under A. gracilis Sharp from Central America. I was unable in a hurried examination to separate these or my specimens from the type of gracilis. However, A. ustulatus was described much earlier from St. John (Virgin Islands of the United States), and I cannot separate it from the specimens under consideration. I therefore conclude that the West Indian examples represent the ustulatus of Erichson and that the grac?/is Sharp should not be in our lists unless it is a synonym of ustulatus, which T am unable to prove. IT have collected this species from under cow dung, from under seaweed on the beach, and flying at dusk. 2. APOCELLUS PLANUS Sharp Apocellus planus SHARP, 1876, p. 401; 1877, p. 147.—BrERNHAUER and SCHUBERT, 1911, p. 107.—ScHEERPELTz, 1933, p. 1091. Description.—Testaceous, head pronotum, elytra, and tip of abdo- men in part infuscate. MWead scarcely wider than pronotum, oval, very feebly impressed at sides; supraantennal prominences large and abrupt; impunctate but densely confusedly strigulose; antennae slen- der, as long as head, pronotum, and elytra together; labrum large, twice as wide as long, deeply emarginate; gular sutures united from near constriction, Y-shaped anteriorly; third segment of maxillary palpi large, fourth acicular. Pronotum about one-half as wide as elytra, scarcely wider than long, apex one-fifth wider than base; very densely and rather coarsely longitudinally strigulose. //ytra seven-ninths wider than long, nearly one-half longer than pronotum ; with dense strigulae as on pronotum. Abdomen smooth. Afale, seventh sternite almost completely divided by a large hemispherical cavity in the thickened posterior margin, cavity one-third as wide as segment; eighth sternite broadly semicircularly emarginate. Length, 214 mm. Type locality—Brazil, Ega, Amazons. Types.—In the Sharp collection in the British Museum. Records.—The following are the records known to me: Trinidad: St. Augustine (C. L. Withycombe, in British Museum). South America: Brazin (Sharp, 1876, 1877), ArnGENTINA (Scheerpeltz, 1933). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE OI Specimens examined.—I have examined seven specimens of this species in the British Museum. Four are from Trinidad. One of these was borrowed for further study. Remarks.—This species is easily distinguished by the strigulose surface. I find no record of its capture since the original description and no indication of its habits. VIII. Genus OXYTELUS Gravenhorst Oxrytelus GRAVENHORS?, 1802, p. 101. Caccoporus THOMSON, 1861, p. 127. Styloxys DES Gozis, 1886, p. 15. Styloris EICHELBAUM, 1909, p. 119. Oyzrtelus BERNHAUER and ScHUBERT, 1911, p. 109. Subgenus Hpomotylis THomson, 1861, p. 128. Subgenus Tanycraerus THOMSON, 1861, p. 129. Subgenus Anotylus THomson, 1861, p. 1380. Anotyhus BERNHAUER, 1988, p. 22. Subgenus Emopotylus BERNHAUER, 1910, p. 359. Genotype—Staphylinus piceus Fabricius = Ozxytelus piceus (Fabricius) (indicated by Duméril, 1806; designated by Latreille, 1810). (The designation of “O. depressum Grav. (= tetracarinatus Block)” by des Gozis in 1886 cannot stand.) Of Anotylus, Oxy- telus nitidulus Gravenhorst (designated here), of Caccoporus, Staphylinus piceus Linnaeus = Owytelus piceus (Linnaeus) (mono- basic) ; of Epomotylus, Qaytelus sculptus Gravenhorst (monobasic) ; of Tanycracrus, Oxytelus lutetpennis Erichson (monobasic) ; of Emopotylus, Oxytelus cuernavacanus Bernhauer (monobasic). Diagnosis —Body moderately depressed; without ocelli; antennae 11-segmented, more or less thickened apically; labrum transverse, emarginate; gular sutures united in anterior two-thirds; fourth seg- ment of maxillary palpi more or Jess subulate; pronotum longitudi- nally sulcate; anterior coxal cavities confluent, entirely open behind; front coxae conical, prominent; middle coxal cavities separated by union of mesosternal and metasternal processes; elytra not dehiscent ; posterior coxae contiguous, “transverse,” not much expanded under the femora; abdomen margined; sternum of first segment absent, second complete, not carinate; posterior trochanters small; front tibiae with a single row of spines; tarsi 3-segmented. Remarks.—This large genus shows a considerable variation in many characters. Exceptions can be found to many of those in the above diagnosis, but the members of the genus are readily dis- tinguishable at a glance by their peculiar facies. I have examined over 1,900 mounted specimens of this genus but have not had time to sort the rest of my West Indian series, which amount to over 10,000 additional examples. 92 BULLETIN 182, UNITED STATES NATIONAL MUSEUM KEY TO WEST INDIAN SPECIES OF OXYTELUS 1. Head, pronotum, and elytra finely sculptured and Opaque s2---— 6. scorpio Head, pronotum, and elytra not both finely sculptured and opaque__---_-- 2 2. Elytra!' muricately -seulptured2- 225212 So) ae ee eee 3 Blytra notimuricately sculptured :» 25. laeviceps Elytra irregularly and indistinctly punctate_______________ gob toetes WE 31 31. Punctures of pronotum elongate, impressed, sparse___._____ 26. jamaicensis Punctures of pronotum round, shallow, moderately dense_--__ 31. exiguus 32. Sculpture of head distinct and definite, more or less clearly longitudinal, especially at ‘sides: of disk=~— 2.2 22 eee 29. weberi Sculpture of head vague and indefinite, not clearly longitudinal even at SidG6.25. 2 os ee eee pees eh Re Le end Bae, 30. trinitatis 1. OSORIUS REGULUS Darlington Osorius regulus DARLINGTON, 1937, pp. 286, 288. Description. — Castaneous to rufotestaceous. Head about nine- tenths as wide as pronotum; eyes small, not at all prominent; an- terior margin of clypeus slightly rounded, irregularly crenate, an- gles not very prominent; labrum distinctly emarginate, alutaceous except on declivity; antennal segments not transverse; surface very finely alutaceous and with tiny punctulae, also a few large punc- tures on vertex. Pronotum strongly narrowed behind, one-fourth wider than long, base three-fourths as wide as apex, sides rather sinuate; side margins very strong near base; with narrow shining midline; with fine punctures almost completely obscured by the dense alutaceous ground sculpture, and a very few larger setigerous punctures. “//ytra one-tenth narrower than pronotum, nearly one- fifth wider than long, widened apically; humeral margins strongly elevated; with a few setigerous punctures, coriaceous throughout. Abdomen feebly alutaceous except at apex, impunctate. Anterior face of front tibia with numerous scattered setigerous punctures. Length, 14 to 15 mm. Type locality—sSouthwestern Haiti, Morne La Hotte, 7,000 feet to summit (about 7,800 feet). Types.—Holotype (M.C.Z. No. 20233) and four paratypes in the Museum of Comparative Zoology; one paratype in the United States National Museum (No. 51249). Records—The following is the only record known to me: Hispaniola: Harir1, Morne La Hotte (Darlington, 1987; U. S. N. M.; A.M.N.H.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 177 Specimens examined.—I have studied only the paratype in the United States National Museum and another from the Museum of Comparative Zoology. Remarks.—This is the largest osoriine known from the West In- dies and among the largest of the genus. Darlington records that the wings are so reduced as to render the insect flightless. The types were taken “in thick damp leaf mold, rotting logs, and under other cover in wet cloud forest.” 2. OSORIUS DARLINGTONI, new species Description.—Piceous, frequently rufescent in part. Head about nine-elevenths as wide as pronotum; eyes small, not at all prominent; anterior margin of clypeus slightly emarginate, strongly crenate, the angles not prolonged but tuberculate; labrum truncate, alutaceous except on declivity; antennal segments not transverse; surface very distinctly alutaceous, with rather coarse sparse punctulae and a few larger punctures. Pronotum strongly narrowed behind, one- tenth broader than long, base four-fifths as wide as apex; sides scarcely sinuate; side margins somewhat expanded just at base; the basal angles prominent; without shining midline; sculptured and punctured as the head. Ziytra one-tenth narrower than pronotum, about one-eighth wider than long, widest at apical third; humeral margins distinctly elevated; strongly coriaceous and with a very few large setigerous punctures. Abdomen distinctly alutaceous throughout. Anterior face of front tibia with scattered setigerous punctures. Length, 9 to 10 mm. Type locality—Cuba, south side of Pico Turquino, elevation 3,000 to 5,000 feet. Types.—Holotype and 7 paratypes in the Museum of Comparative Zoology, 6 paratypes in the United States National Museum (No. 52487) ; collected by Dr. P. J. Darlington in June and July 1936 at the localities listed below. Records.—The following is the only record known to me: Cuba: Pico Turquino (Darlington, in M.C.Z. and U.S.N.M.), Upper Ovando River, Oriente (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 14 types and 2 imma- ture specimens. Remarks.—This species is most similar to regulus but differs in its smaller size, in the emargination of the clypeus, the narrower pro- notum, and so on. It represents the extreme development of clypeal crenation in which the crenulae are almost tubercles. I have received no record of the habits of this species. 178 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 3. OSORIUS CRENULIFRONS Notman Osorius crenulifrons Norman, 1925, pp. 9, 15, 18—ScHEERPELTz, 1933, p. 1133.— Dar.ineTon, 1937, pp. 286, 289, 291, 292. Description —Rufopiceous. Head about five-sixths as wide as pronotum; eyes relatively very prominent; clypeus truncate, very unevenly crenate, angles indefinite; labrum appearing emarginate though straight below the declivity, alutaceous; antennae slender; with sparse fine punctures obscured by the strong dense alutaceous ground sculpture. Pronotum about one-fifth wider than long, feebly narrowed posteriorly, base nearly as wide as apex; sides feebly arcuate and sinuate; lateral margin scarcely widened posteriorly ; with a vague impression at midline in part; with moderately fine punctures separated by three to four times their diameter, obsoletely alutaceous. lytra barely wider than pronotum, a little longer than wide, sides rather strongly arcuate; coriaceous; impunctate. Abdo- men alutaceous, scarcely punctate. Anterior face of front tibia with scattered setigerous punctures. Length, 9 mm. Type locality—Cuba, San Diego (de los Banos, Pinar del Rio Province). Types.—Holotype, U.S.N.M. No. 26344. Records.—The following is the only record known to me: Cuba: San Diego de los Banos (Notman, 1925; Darlington, 1937; U.S.N.M.). Specimens examined.—I have examined the type, the only speci- men known. Remarks.—This species is very similar to salutator of Darlington but may be distinguished by characters noted under that species. I find no record of its habits. 4. OSORIUS FRATELLUS Darlington Osorius fratellus DARLINGTON, 1937, pp. 286, 291. Description.—Piceous, pronotum, and elytra rufescent. Head seven-eighths to nine-tenths as wide as pronotum; eyes scarcely at all prominent; clypeus truncate, coarsely crenate, angles not very promi- nent; labrum very slightly emarginate, alutaceous; antennal segments not transverse; with a few coarse punctures and sparse finer ones ob- scured by dense alutaceous ground sculpture. Pronotwm widest at anterior third, moderately narrowed behind, one-fourth wider than long, base nearly as wide as apex; margin moderately expanded near base; with scarcely a trace of midline; with very sparse fine punctures and dense alutaceous ground sculpture. Z7Jytra not narrower than pronotum, about as long as wide; narrowed to base and to apex from apical third; humeral margin feeble; with a few large setigerous punctures; coriaceous. Abdomen feebly alutaceous; above with only MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 179 a few tuberculate setigerous punctures. Anterior face of front tibia with numerous scattered setigerous punctures. Length, 9 to 10 mm. Type locality —Southeastern Haiti, Furcy, La Selle Range, elevation about 5,000 feet. Types.—Holotype (M.C.Z. No. 20235) and one paratype in the Museum of Comparative Zoology. Records——The following are the records known to me: Hispaniola: Harti, Furcy (Darlington, 1937), Tardieu (Darlington, 1937). Specimens examined—tI have seen only the paratype from the Museum of Comparative Zoology. Remarks.—This species is only feebly distinguished from salutator, but I am unable to form an opinion as to its true status for lack of material. I find no record of its habits. 5. OSORIUS SALUTATOR Darlington Osorius salutator DARLINGTON, 1937, pp. 286, 290, 291, 292. Description—Black. Head four-fifths as wide as pronotum; eyes slightly prominent; clypeus truncate, coarsely crenate, angles not very prominent; labrum very slightly emarginate, alutaceous; antennal segments not transverse; with a few coarse punctures and sparse finer ones obscured by dense alutaceous ground sculpture. Pronotum widest at anterior third, moderately narrowed behind, one-fourth wider than long, base nearly as wide as apex; margin not greatly expanded near base; with scarcely a trace of midline; with very sparse fine punctures and dense alutaceous ground sculpture. LHlytra only slightly narrower than pronotum or shorter than wide, narrowed to base and to apex from apical third; humeral margins feeble; with a few large setigerous punctures; coriaceous. Abdomen feebly aluta- ceous except at apex, above with only a few tuberculate setigerous punctures. Anterior face of front tibia with numerous scattered setigerous punctures. Length, 10 to 1214 mm. Type locality —Southeastern Haiti, La Visité and vicinity, La Selle Range, 5,000 to 7,000 feet elevation. Types.—Holotype (M.C.Z. No. 20234) and four paratypes in the Museum of Comparative Zoology; one paratype in the United States National Museum (No. 51247). Records.—The following are the records known to me: Hispaniola: Hariri, La Visité (Darlington, 1987; M.C.Z.; U.S.N.M.), Mount Basil (Darlington, 19387). Specimens examined.—I have seen one paratype in the United States National Museum and one paratype and two other specimens from the Museum of Comparative Zoology. 180 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—The specimens from Mount Basil in northern Haiti are reported by Darlington to be somewhat different from the types. The sinuation of the sides of the pronotum is not sufficient to make a clear distinction between this and crenulifrons, but in the latter the sides at base are less explanate, the hind angles not prominent externally, and the base is not feebly emarginate. The types were taken “in thick leaf mold in wet cloud forest.” 6. OSORIUS HIRTILABRIS Darlington Osorius hirtilabris DARLINGTON, 19387, pp. 284, 286, 292. Description.—Piceous. ead about nine-tenths as wide as pro- notum; eyes very prominent; clypeus truncate, anterior margin crenate (serrately); labrum long, truncate, with rounded angles, alutaceous basally, swollen and shining anteromedially; antennae slender; surface minutely punctulate and finely alutaceous. Pro- notum one-third wider than long, strongly narrowed behind, base barely three-fourths as wide as apex; sides arcuate anteriorly, sinu- ate posteriorly; lateral margin much broader basally; without dis- tinct median line; sparsely punctate; with only a trace of ground sculpture. lytra as wide as pronotum, scarcely longer than wide, not much enlarged posteriorly; humeral margin feeble; with a very few large setigerous punctures; coriaceous. Abdomen moderately densely punctate, not distinctly alutaceous except at base. Anterior face of front tibia with setigerous punctures scattered. Length, 9 to 11 mm. Type locality —Southwestern Haiti, “northeastern foothills of the Massif de la Hotte, 2,000-4,000 feet.” Types.——Holotype (M.C.Z. No. 20236) and 16 paratypes in the Museum of Comparative Zoology; 2 paratypes in the United States National Museum (No. 51250). Records.—The following are the records known to me: Hispaniola: Hartt, Morne La Hotte (Darlington, 19387; M.C.Z.; U.S.N.M.), Miragoane (Darlington, 1987), Kenskoff (Darlington, 1937). Specimens examined.—I have seen only the two paratypes in the United States National Museum and one from the Museum of Comparative Zoology. Remarks.—Very distinct by the unusual characters of the labrum. The types were “taken struggling in a pond of flood water in scrubby (formerly rain-forest) country.” 7. OSORIUS DAEMON Darlington Osorius daemon DARLINGTON, 1937, pp. 286, 293. Description —Piceous. Head scarcely narrower than pronotum; eyes not prominent; clypeus narrow, distinctly emarginate at middle, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Si angles not prominent, margin coarsely and irregularly crenate; labrum emarginate, unusually large, alutaceous; outer antennal seg- ments as broad as long; surface strongly alutaceous, with minute punctures and a few coarse ones. Pronotum moderately narrowed behind, one-fifth wider than long, base four-fifths as wide as apex; sides feebly sinuate; lateral margin very little broader basally; with- out trace of midline; distinctly alutaceous and finely punctulate. Elytra about as wide as pronotum, nearly as long as wide; sides feebly arcuate; surface shining but coriaceous, sparsely, irregularly, and rather finely punctate; with a few large setigerous punctures. Abdomen feebly alutaceous; moderately closely but not densely punc- tate, seventh and eighth tergites more closely. Anterior face of front tibia with setigerous punctures scattered. Length, 10 mm. Type locality —Southwestern Haiti, northwestern foothills of the Massif de la Hotte, elevation 2,000 to 4,000 feet. Types.—Holotype (M.C.Z. No. 20237) and one paratype in the Museum of Comparative Zoology. Records.—The following are the records known to me: Hispaniola: Haiti, Massif de la Hotte (Darlington, 1937), Tardieu (Darlington, 1937). Specimens examined.—I have seen only the paratype from the Museum of Comparative Zoology. Remarks.—The clypeus in this species is rather asymmetrical but is quite distinctive. The description is based upon the paratype. I find no record of the habits of this species. 8. OSORIUS CUBENSIS, new species Description.—Rufopiceous, pronotum and elytra almost rufous. Head four-fifths as wide as pronotum, eyes not very prominent; clypeus feebly rounded, angles evident but not prominent, coarsely crenate; labrum large, emarginate above declivity only, alutaceous; apical antennal segments nearly as wide as long; with sparse mod- erate punctures and a few large ones obscured by strong alutaceous ground sculpture. Pronotuwm one-sixth wider than long, rather strongly narrowed posteriorly, base eight-ninths as wide as apex; sides arcuate in front, nearly straight behind; lateral margin scarcely widened posteriorly ; with a trace of smooth midline; with moderately fine punctures separated by two to four times their diameter; alu- taceous only at base. “/ytra about one-tenth wider than pronotum, one-tenth wider than long, sides feebly rounded; coarsely coriaceous, impunctate. Abdomen alutaceous, with moderately dense large sub- muricate punctures. Anterior face of front tibia with a few scat- tered setigerous punctures. Length, 7 mm. Type locality—Cuba, San Antonio de los Banos. 182 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Types.—Holotype, United States National Museum No. 52380, col- lected by José H. Pazos. (This specimen was the paratype of O. buscki recorded by Notman.) Also 10 paratypes from localities listed below, which are deposited in the United States National Museum and the Museum of Comparative Zoology. Records.—The following are the records known to me: Cuba: San Antonio de los Banos (Pazos, in U.S.N.M.), Buenos Aires, Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.), Rangel Mountains (Darling- ton, in M.C.Z and U.S.N.M.), Imias (Darlington, in M.C.Z.), Soledad (Darlington, in M.C.Z.). Specimens examined.—\ have seen only the 11 types. Remarks.—This species seems to be quite distinct from buscki by its rounded clypeus, its less alutaceous pronotum, and its scattered punctures on the anterior tibiae. It was recognized as probably distinct in the National Museum collection by Dr. Chapin. One additional specimen from Rangel Mountains differs from the others in having the pronotal punctures distinctly coarser. This difference appears to me to be much too slight to warrant separating it specifically, since only the size of the punctures is involved, not arrangement or density or shape. I have no record of its habits. 9. OSORIUS STRICTUS, new species Description —Rufopiceous. Head five-sixths as wide as prono- tum; the eyes moderately prominent; the clypeus feebly rounded, the angles not prominent, somewhat indistinctly crenate; labrum large, feebly emarginate both above and below the declivity, alutaceous; apical antennal segments nearly as wide as long; with sparse moderate punctures and a few large ones obscured by strong alutaceous ground sculpture. Pronotum one-eighth wider than long, rather strongly narrowed behind middle, base seven-eighths as wide as apex; sides distinctly emarginate before basal angles; lateral margin distinctly enlarged and recurved at basal angle; without trace of midline; with fine and shallow punctures separated by three to five times their diameter; feebly alutaceous throughout. Hlytra very nearly as wide as pronotum, as long as wide; coarsely coriaceous and im- punctate except for a few large punctures. Abdomen with aluta- ceous sculpture on each segment but obsolescent at apex of each; with moderately coarse but sparse setigerous punctures, denser apically. Anterior face of front tibia with setigerous punctures scattered. Length, 8 mm. Type locality.—Cuba, mountains north of Imias, in eastern Oriente Province, elevation 3,000 to 4,000 feet. Types—Holotype, Museum of Comparative Zoology, collected by Dr. P. J. Darlington on July 25-28, 1936. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 183 Records—The following is the only record known to me: Cuba: Imias (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks—This specimen differs from cubensis in numerous small details and especially in the shape of the pronotum. I have received no record of its habits. 10. OSORIUS SOCORS Darlington Osorius socors DARLINGTON, 1937, pp. 286, 294. Description.—Piceous. Head about four-fifths as wide as prono- tum; eyes slightly prominent; clypeus truncate, margin finely crenu- late; labrum scarcely emarginate, alutaceous; antennae slender; faintly or not alutaceous except basally, distinctly punctulate; with a few larger punctures at sides. Pronotum one-tenth wider than long, slightly narrowed behind, base nine-tenths as wide as apex; sides feebly arcuate; apical angles denticulate; lateral margin only slightly broader basally, with conspicuous smooth midline; not alutaceous but strongly punctulate. lytra as wide as pronotum and as long as wide; sides nearly parallel; with a few large setigerous punc- tures; coriaceous. Abdomen not distinctly alutaceous, finely not densely punctate. Anterior face of front tibia with setigerous punc- tures scattered. Length, 744 mm. (From Darlington.) Type locality—Central Cuba, Central Soledad, near Cienfuegos, Santa Clara Province. Types.—Holotype in the Museum of Comparative Zoology (No. 91238). Records.—The following is the only record known to me: Cuba: Central Soledad (Darlington, 1937). Specimens examined.—I have seen no examples of this species. Remarks.—Though separated in the key from manni, this species is very similar to it. In fact I think it very unlikely that it can be retained as distinct. The considerable difference in size appears to me to be the only positive character distinguishing them. Since both are represented by unique types, it will be necessary to obtain addi- tional material before a solution can be found, unless a study of the genitalia of this genus reveals usable specific characters. (The type of manné is a female.) “Taken with a Berlese funnel near Harvard House.” 11. OSORIUS MANNI Notman Osorius manni NoTMAN, 1925, pp. 9, 19, 25—ScHrEERPELTZ, 1933, p. 1184.—Dar- LINGTON, 1937, pp. 286, 294, 295, 296. Description.—Rufopiceous. Head nearly six-sevenths as wide as pronotum; eyes relatively prominent; clypeus wide, truncate and very 184 BULLETIN 182, UNITED STATES NATIONAL MUSEUM feebly emarginate, rather finely crenulate; labrum truncate, aluta- ceous; antenna slender; alutaceous and finely punctulate. Pronotum one-fourth wider than long; moderately narrowed posteriorly, base about as wide as apex; sides feebly evenly arcuate except near base; lateral margin scarcely widened posteriorly; without distinct mid- line; alutaceous and sparsely finely punctate. Llytra very slightly wider than pronotum, nearly one-seventh wider than long; coriaceous, with a few large setigerous punctures. Abdomen feebly alutaceous except on apical segments; with sparse muricate setigerous punctures especially apically on each segment. Front tibiae not known (lost from type). Length, 11 mm. Type locality—Cuba, Pinares in Oriente. Types.—Unique holotype in the United States National Museum (No. 26345). Records.—The following are the records known to me: Cuba: Pinares (Notman, 1925; Darlington, 1937; U.S.N.M.), Yunque de Baracoa, Oriente (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype and one exam- ple in the Museum of Comparative Zoology. Remarks.—This species is very similar to cautis and socors of Dar- lington. It has been taken only twice. I find no record of its habits except that Dr. Darlington’s specimen was collected at an elevation of 1,000 to 1,800 feet. 12. OSORIUS CAUTIS Darlington Osorius cautis DARLINGTON, 1937, pp. 286, 295. Description.—Piceous. Head eight-ninths as wide as pronotum; eyes only very slightly prominent; clypeus truncate, anterior margin crenulate, angles slightly prominent; labrum slightly emarginate, alutaceous; antennae slender; alutaceous and finely punctulate, with a few large setigerous punctures. Pronotwm one-seventh wider than long, moderately narrowed behind; base four-fifths to six-sevenths as wide as apex; sides nearly straight, feebly sinuate; lateral margin broader posteriorly; without distinct midline; finely alutaceous and punctulate. HJytra as wide as or slightly wider than pronotum, as long as wide, nearly parallel; surface shining, sparsely punctulate, with a few large setigerous punctures. Abdomen not distinctly alu- taceous; sparsely punctate above. Anterior face of front tibia with a single series of setigerous punctures. Length, 11 to 1114 mm. Type locality—Southwestern Haiti, summit of ridge just above Roche Croix, on a northern spur of Morne La Hotte, elevation 5,000 feet. Types.—Holotype (M.C.Z. No. 20239) and seven paratypes in the Museum of Comparative Zoology; one paratype in the United States MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 185 National Museum (No. 51248); and one paratype in the American Museum of Natural History. Records.—The following are the records known to me: Hispaniola: Harri, Roche Croix (Darlington, 1987; M.C.Z.; U.S.N.M.), Morne La Hotte (Darlington, 1937). Specimens examined.—l\ have seen only the paratype in the United States National Museum and one from the Museum of Comparative Zoology. Remarks.—This species differs from manni in having the clypeus much narrower, the angles slightly produced, and in having the labrum distinctly emarginate. I am unable to see any distinct difference in the sides of the pronotum, and the differences in the size of the sub- marginal channels is very slight. “Taken under various ground cover in wet cloud forest.” 13. OSORIUS MEDIOCRIS Darlington Osorius mediocris DARLINGTON, 1937, pp. 286, 296, 297. Description.—Piceous to rufopiceous. Head six-sevenths as wide as pronotum; eyes rather prominent; clypeus truncate, strongly cre- nate; labrum not emarginate, alutaceous; antennae with outer seg- ments as wide as long; strongly alutaceous, finely sparsely punctulate, with a few large punctures. Pronotum one-tenth to one-seventh wider than long, feebly narrowed behind, base about seven-ninths as wide as apex; sides feebly arcuate; lateral margin somewhat broader basally; without trace of midline; alutaceous, with rather fine punc- tures. Llytra not quite so wide as pronotum, as long as wide, not much expanded; coriaceous, with a few large setigerous punctures. Abdomen scarcely visibly alutaceous except basally below; rather finely and closely submuricately punctate. Anterior face of front tibia with setigerous punctures in a single series. Length, 814 mm. Type locality—Southeastern Haiti, La Visité and vicinity, La Selle Range, 5,000-7,000 feet elevation. Types.—Holotype (M.C.Z. No. 20240) and one paratype in the Museum of Comparative Zoology. Records.—The following is the only record known to me: Hispaniola: Hartt, La Visité (Darlington, 1937). Specimens examined.—\I have seen only the paratype from the Museum of Comparative Zoology. Remarks.—Since the removal of the paratype of buschi as a new species, the size difference between mediocris and buschki becomes very great. They differ also in the relative size of the head and the form of the labrum. JT find no record of the habits of this species. 449008—42 13 186 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 14. OSORIUS BIARMATUS Darlington Osorius biarmatus DARLINGTON, 1937, pp. 285, 287. Description —Black. Head as wide as pronotum; eyes not at all prominent; front margin of clypeus feebly crenate, angles conspicu- ously, acutely, and somewhat divergently produced, margin evenly emarginate between; labrum emarginate, alutaceous; antennae faintly incrassate from the seventh segment; surface coriaceous in front, alutaceous at rear, and with minute punctulae; moderately punctate at sides of vertex and clypeus. Pronotum much narrower basally, one-fifth to one-sixth wider than long; apex one-half wider than base; sides distinctly sinuate; margins not enlarged basally; coria- ceous and with minute punctulae, but not alutaceous; moderately coarsely, not closely punctate; median stripe impunctate, not well defined. Hlytra as wide as pronotum, one-twentieth to one-tenth longer than wide, nearly parallel; punctured as pronotum; not aluta- ceous, but coriaceous. Abdomen moderately closely and coarsely punctate; eighth tergite faintly alutaceous. Anterior face of front tebia with single series of setigerous punctures. Length, 714 to 734 mm. Type locality —Southwestern Haiti, northeastern foothills of the Massif de la Hotte, 2,000 to 4,000 feet elevation. Types—Holotype (M.C.Z. No. 20232) and one paratype in the Museum of Comparative Zoology. Records.—The following is the only record known to me: Hispaniola: Harri, Massif de la Hotte (Darlington, 1937). Specimens examined.—I have seen only the paratype from the Museum of Comparative Zoology. Remarks.—Darlington records that this species combines charac- ters of the large species with crenate clypeus and the small species with smooth clypeal margin. I find no mention of its occurrence. 15. OSORIUS BUSCKI Notman Osorius buscki NoTMAN, 1925, pp. 9, 19, 24——ScHEERPELTz, 1933, p. 11382.—Dar- LINGTON, 1987, p. 286, 297. Description—Piceorufous. Head just over three-fourths as wide as pronotum; eyes moderately prominent; clypeus rounded, some- times very feebly biemarginate at the sides, anterior margin strongly crenulate; labrum emarginate, alutaceous; outer antennal segments about as wide as long; alutaceous and sparsely punctulate, with a few larger punctures. Pronotum one-seventh wider than long, feebly narrowed behind, base nearly six-sevenths as wide as apex; sides evenly arcuate; lateral margins somewhat wider posteriorly; a vague trace of midline; alutaceous, and with moderate punctures separated -— MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 187 by three times their diameter on the disk. Zlytra about as wide as pronotum, as long as wide, feebly enlarged; coriaceous, impunctate. Abdomen alutaceous; with moderate muricate punctures. Anterior face of front tibia with a single series of setigerous punctures. Length, 5 mm. Type locality— ‘Santo Domingo” (=Dominican Republic). Types.—Holotype in the United States National Museum (No. 26346), collected by August Busck. (The recorded paratype is now the type of O. cubensis Blackwelder.) Records.—The following are the records known to me: Hispaniola: DOMINICAN Repuetic (Notman, 1925; U.S.N.M.); Harri, La Visité (Darlington, 1987; A.M.N.H.), Kenskoff (Darlington, 1937). Specimens exumined.—I have seen only the type and one of the specimens collected by Dr. Darlington at La Visité. Remarks.—The paratype that Notman designated was from Cuba and is believed to be a distinct species. It is herein described as cubensis new species. Darlington has collected nine additional spec- imens of this species in Haiti. I find no record of its habits. 16. OSORIUS ORIENTE, new species Description—Rufopiceous. Head thirteen-fourteenths as wide as pronotum; eyes prominent; clypeal margin scarcely arcuate except when viewed from behind, with traces of emarginations near the angles, margin feebly crenate; labrum very feebly emarginate, aluta- ceous; antennal segments 7 to 10 about as broad as long; with a very few large punctures, minute punctulae obscured by strong alu- taceous sculpture. Pronotum one-sixth wider than long, moderately narrowed behind, base eleven-fourteenths as wide as apex; sides evenly arcuate, without trace of basal emargination; lateral margins scarcely at all widened posteriorly; without trace of midline; feebly alutaceous, with sparse and fine punctulae. lytra as wide as pro- notum, one-seventh longer than wide; coriaceous, impunctate. Abdo- men alutaceous, more densely on eighth tergite; with moderate sub- muricate punctures not dense. Anterior face of front tibia with a single series of setigerous punctures. Length, 514 to 7 mm. Type locality—Cuba, mountains north of Imias, in eastern Ori- ente Province, elevation 3,000 to 4,000 feet. Types.—Holotype and one paratype, Museum of Comparative Zoology, and one paratype in the United States National Museum (No. 52488), collected by Dr. P. J. Darlington on July 25-28, 1936. Records.—The following is the only record known to me: Cuba: Imias (Darlington, in M.C.Z. and U.S.N.M.). 188 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 3 Specimens examined.—I\ have seen only the three types. Remarks —This species differs from buscki by its fine pronotal punctures, the lack of emargination of the sides basally, the long elytra, and its greater size. It differs from twrquinus in the shape of the pronotum. I have received no record of its habits. 17. OSORIUS TURQUINUS, new species Description.—Piceous to rufopiceous. Head eight-ninths as wide as pronotum; eyes not very prominent; clypeus feebly rounded, very feebly emarginate near the angles, which are prominent but not produced, anterior margin rather feebly crenate; labrum very feebly emarginate, alutaceous; antennal segments all longer than wide; with sparse but irregular punctures obscured by strong alu- taceous ground sculpture. Pronotum one-eighth wider than long, strongly narrowed behind, widest at front, base just two-thirds as wide as apex; sides nearly straight; lateral margins feebly explanate at basal angles; without trace of midline; with rather fine sparse punctures not obscured by feeble alutaceous sculpture. Elytra one- sixteenth narrower than pronotum, as long as wide; sides evenly arcuate; with a few rather large but very shallow punctures, scat- tered; coriaceous. Abdomen alutaceous, much more densely on eighth tergite; with rather small but submuricate setigerous punc- tures, moderately dense. Anterior face of front tibia with a single series of setigerous punctures. Length, 814 to 9 mm. Type locality—Cuba, south side of Pico Turquino, elevation 3,000 to 5,000 feet. Types.—Holotype, Museum of Comparative Zoology, and para- type, United States National Museum (No. 52489), collected by Dr. P. J. Darlington in June 1936. Records.—The following is the only record known to me: Cuba: Pico Turquino (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—lI have seen only the two types. Remarks.—This species is very distinct from buscki by the shape of the pronotum. I have received no record of its habits. 18. OSORIUS LATIPES (Gravenhorst) Oxrytelus latipes GRAVENHORST, 1806, pp. 183, 184, 196, 198.—Ortvier, 1811, p. 616.— Say, 1834, p. 462.—LrConte, 1859, p. 578; 1877, p. 216. Molosoma latipes (Gravenhorst) Say, 1834, p. 462.—Ericuson, 1840, p. 757.— LECOoNTE, 1859, p. 578; 1877, p. 216. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 189 Osorius latipes (Gravenhorst) ErtcHson, 1840, p. 757.—LAcorpAtre, 1854, p. 113.—Scripa, 1855, p. 801.—LEConTr, 1859, pp. 37, 462.—Summers, 1874, p. 52.—SuHarp, 1876, p. 386, 432.—LeConre, 1877, p. 216—ScHwarz, 1878, p. 442.—Dury, 1879, p. 165.—SHarp, 1887, p. 682.—HAmILTOoN, 1895, pp. 327, 357.—Dury, 1902, p. 125.—BLAtTcHLEY, 1910, p. 473.—BERNHAUER and Scuusert, 1911, p. 144.—NoTMAN, 1920, p. 699; 1925, pp. 3, 8, 23, 25.— SCHEERPELTZ, 1933, p. 1184—JonrEs, 1935, p. 151.—Dariineron, 1937, pp. 287, 299. Description.—Piceous to rufopiceous, pronotum and elytra gen- erally more rufous. Head about six-sevenths as wide as pronotum; eyes scarcely at all prominent; clypeus feebly broadly emarginate, angles somewhat prominent; labrum very short, a little emarginate above declivity, alutaceous; outer antennal segments about as wide as long; with rather coarse punctures especially at sides; obsoletely alutaceous. Pronotum one-twelfth wider than long, rather strongly narrowed behind, base three-fourths as wide as apex; lateral margin not at all widened posteriorly; with an impressed midline set off by irregular rows of punctures; with very coarse punctures sometimes united longitudinally, separated transversely by less than twice their width; alutaceous ground sculpture almost completely obsolete. Elytra about as wide as pronotum, a little longer than wide, parallel ; finely coriaceous; with very coarse and irregular punctures, more or less serial; not alutaceous. Abdomen not alutaceous, except vaguely apically; very coarsely but not muricately punctate. An- terior face of front tibia with a few very fine scattered setigerous punc- tures. Length, 4 to 6 mm. Type locality —‘*America septentrionali.” Types.—Presumably in the Zoologische Museum, Berlin. Records.—The following are the records known to me: Cuba: Soledad (Darlington, 1937). North America: FtormA, ALABAMA, VIRGINIA, DisTRICT OF COLUMBIA, MARYLAND, PENNSYLVANIA, OHIO, KENTUCKY, INDIANA, IowA, NEBRASKA, SOUTH DAKOTA, KANSAS, Missouri, ARKANSAS, TExAS (Gravenhorst, 1806; Say, 1834; Notman, 1925; Darlington, 1937; ete.). Specimens examined —I have examined many examples from North America, but from the West Indies I have seen only the example collected by Dr. Darlington. Remarks.—The only West Indian specimen known was collected by Dr. Darlington in Cuba. He records that it has the head more strongly alutaceous than is usual in this species. The above de- scription was taken principally from specimens collected near Washington, D. C. It has been collected in the United States from South Dakota to Texas and from Florida to New England. 190 BULLETIN 182, UNITED STATES NATIONAL MUSEUM This species has been recorded as “burrowing in the ground beneath stones.” 19. OSORIUS MINOR Notman Osorius minor NoTMAN, 1925, pp. 7, 14, 25.—ScHEERPELTZ, 1933, p. 1134. Description.—Dark rufopiceous. Head nine-tenths as wide as pro- notum; eyes moderately prominent; clypeus feebly emarginate in front, angles feebly tuberculate; labrum feebly emarginate above de- clivity, alutaceous; outer segments of antennae as wide as long; with numerous large punctures; densely strongly alutaceous. Pro- notum one-ninth wider than long, strongly narrowed posteriorly; base nearly five-sixths as wide as apex; sides feebly arcuate and very slightly sinuate, posterior angles rounded; lateral margin not wider posteriorly; with an impressed middle band set off by serial punctures; with very coarse frequently elongate punctures, gener- ally separated transversely by less than half their width; very obso- letely alutaceous. E/ytra scarcely wider than pronotum, a little wider than long, not much enlarged posteriorly; with moderately large punctures, frequently serial and somewhat united; coriaceous but not alutaceous. Abdomen very vaguely alutaceous; punctures rather large, often submuricate. Anterior face of front tibia with a very few scattered setigerous punctures. Length, 4 mm. Type locality—Trinidad, ward of Montserrat in the county of Caroni. Types.—Holotype in the United States National Museum (No. 27330). fecords.—The following are the records known to me: St. Vincent: (British Museum). Grenada: (British Museum; Cameron collection). Trinidad: Montserrat (Notman, 1925, U.S.N.M.). Specimens examined.—lI have seen the unique type in the United States National Museum, 18 examples in the British Museum, and 2 in Dr. Cameron’s collection. Remarks.—This species is similar to latipes but is distinct by its strongly alutaceous head. It is possible that this will eventuaily be found to fall within the ranges of variation and distribution of latipes, but at present it is best retained as distinct. The specimens in the British Museum and in Dr. Cameron’s col- lection were labeled Osorius parcus Sharp. O. minor differs from the Central American specimens in its smaller size, less distinct elytral punctation, its obsolescent abdominal sculpture, and the less distinctly bidenticulate angles of the clypeus. I find no record of the habits of this species. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 191 20. OSORIUS SCHWARZI Notman Osorius schwarzi NoTMAN, 1925, p. 7, 18, 26.—ScHEERPELTz, 1933, p. 1136.—Dar- LINGTON, 1937, pp. 287, 299, 300. Description—Black, elytra rufopiceous mediobasally. Head about seven-eighths as wide as pronotum; eyes a little prominent; clypeus broadly emarginate in front, angles prominent; labrum scarcely emarginate, even above declivity, alutaceous; antennae slen- der; with moderately large but not very sparse punctures obscured by very strongly and densely alutaceous ground sculpture. Prono- tum about one-seventh wider than long, moderately narrowed pos- teriorly, base three-fourths as wide as apex; sides scarcely but evenly arcuate; hind angles feebly rounded; lateral margins not enlarged pos- teriorly; without midline except as impunctate band; with very coarse punctures, generally separated by less than twice their diam- eter; alutaceous. Zlytra as wide as pronotum, a little longer than wide; sides nearly straight; coriaceous, but with large punctures more or less serial, and moderately distinct alutaceous ground sculp: ture. Abdomen alutaceous; with submuricate punctures. Anterior face of front tibia with a few scattered setigerous punctures. Length, 5 mm. Type locality—Cuba, Cayamas, about 15 miles west-northwest of Cienfuegos, province of Santa Clara. Types.—Holotype and two paratypes in the United States Na- tional Museum (No. 26334); one paratype in the collection of How- ard Notman; one paratype in the Museum of Comparative Zoology. Records.—The following is the only record known to me: Cuba: Cayamas (Notman, 1925; Darlington, 1987; U.S.N.M.). Specimens examined.—\ have examined the three types in the United States National Museum. Remarks.—This species has not been reported since its description. It is rather feebly differentiated from haitiellus and minor. I find no record of its habits. 21. OSORIUS HAITIELLUS Darlington Osorius haitiellus DARLINGTON, 1937, pp. 287, 299, 300. Description —Black, elytra in part rufopiceous. Head nine- tenths as wide as pronotum; eyes scarcely prominent; clypeus trun- cate, not crenulate, angles denticulate; labrum not emarginate, decliv- ous in front, alutaceous only at sides; apical antennal segments about as wide as long; impunctate; strongly alutaceous; with setigerous tuberculi at sides. Pronotum about as wide as long, moderately narrowed behind, basal angles narrowly rounded, base nearly five- sixths as wide as apex; sides rather sinuate; lateral margin not wider posteriorly; without definite midline; with coarse punctures 192 BULLETIN 182, UNITED STATES NATIONAL MUSEUM sometimes united longitudinally, transversely separated by twice their width; alutaceous. /7ytra about one-twentieth wider than pronotum, one-twentieth longer than wide, sides nearly parallel; rather coarsely and indefinitely but serially punctate; coriaceous. Abdomen not alutaceous except feebly at apex; rather coarsely submuricately punc- tured. Anterior face of front tibia with setigerous punctures in a short series. Length, 414 to 5144 mm. Type locality—Southwestern Haiti, “northeastern foothills of the Massif de la Hotte, 2,000-4,000 feet.” Types.—Holotype (M.C.Z. No. 20242) and three paratypes in the Museum of Comparative Zoology; one paratype in the United States National Museum (No. 51246). Records.—The following are the records known to me: Cuba: Mountains north of Imias, Oriente (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Hartt, Massif de la Hotte (Darlington, 1987; M.C.Z.; U.S.N.M.)., Tardieu (Darlington, 1987), La Visité (Darlington, 1937), Kenskoff (Darling- ton, 1937). Specimens examined.—I have seen only the one paratype in the United States National Museum, one paratype and two other speci- mens from the Museum of Comparative Zoology. Remarks.—This species differs from schwarzi especially in the denticulate clypeal angles and the single series of punctures on the tibiae. No record has been made of its habits. 22. OSORIUS INTERMEDIUS Erichson Osorius intermedius ErtcHson, 1840, p. 754.—Lacorpamr, 1854, p. 113.— CANDEZE, 1861, p. 331.—So.tsky, 1869, p. 265,—ViILLapa, 1901, p. 33.—EICHEL- BAUM, 1909, p. 126.—BERNHAUER and SCHUBERT, 1911, p. 144.—BERNHAUER, 1920b, p. 188.—NotTMAN, 1925, pp. 11, 24, 25—ScHEERPELTz, 1933, p. 1134. (Not Cameron, 1928.) Description —Black. Head eight-ninths as wide as pronotum; clypeus broad, with angles forming rounded obtuse prominences and margin between sinuate; labrum short, alutaceous; antennae slender ; with moderately fine punctures throughout; densely alutaceous; with a few tuberculi at sides of clypeus. Pronotwm two-sevenths wider than long, rather strongly narowed behind, base two-sevenths as wide as apex, sides straight in front, posterior angles obtuse but slightly prominent; lateral margin feebly widened posteriorly; with only a trace of impressed midline; with moderately fine abrupt punctures separated by three to five times their diameter, alutaceous, and with an occasional punctiform depression. Llytra scarcely wider than pronotum, a little wider than long, rather broadly rounded at sides; coarsely coriaceous, but with distinct punctiform depressions. Abdo- men distinctly alutaceous only on apical segments; with a very few MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 193 large tuberculate punctures. Anterior face of front tibia with an irregular series of setigerous punctures. Length, 12-14 mm. Type locality—Columbia interiore.” Types.—Possibly in the Zoologische Museum, Berlin. Records —The following are the records known to me: Trinidad: (British Museum); Manzanilla (Blackwelder station 104B), Mount Tucuche (Darlington, in M.C.Z. and U.S.N.M.), Mount St. Benedict (Weber, in M.C.Z.). South America: VENEZUELA (British Museum), Cotompt1a (Erichson, 1840; Lacor- daire, 1854; Notman, 1925; British Museum). [Central America: Costa Rica (Notman, 1925), Mexico (Candéze, 1861).] Specimens examined—tI have examined four specimens in the British Museum, three from the Museum of Comparative Zoology, and eight collected by me in 1935-387. Remarks.—TYhe Costa Rican examples recorded by Notman have been examined in the United States National Museum. They differ rather strongly from my specimens in their coarse clypeal puncta- tion, and I believe them to be distinct. My examples seem to agree closely with ones in the British Museum from Colombia and Venezuela. The larval example recorded from Mexico by Candeéze is certainly very doubtfully referable to this species. I collected these specimens from a very rotten log in a damp situation. 23. OSORIUS HUBBARDI Notman Osorius hubbardi NoTMAN, 1925, pp. 6, 11, 25.—ScHEERPELTzZ, 1933, p. 1184.—Dar- LINGTON, 1937, pp. 287, 297. Description.—Piceorufous. Head six-sevenths as wide as prono- tum; eyes moderately prominent; clypeus broad, angles prolonged in acute processes nearly as long as distance between them, anterior margin not crenulate; labrum emarginate, obsoletely alutaceous; an- tennae slender; with scattered moderate punctures posteriorly and a few small tubercles anteriorly ; strongly and densely alutaceous. Pro- notwm one-sixth wider than long, moderately narrowed posteriorly, base five-sixths as wide as apex; sides rounded but sinuate; lateral margin scarcely widened posteriorly; without distinct midline; with moderate but impressed sparse punctures; alutaceous. Hlytra one- seventh wider than pronotum, one-tenth wider than long; strongly arcuate at sides; coarsely coriaceous, but with indefinite large punc- tures. Abdomen alutaceous, with large sparse punctures. Anterior face of front tibia with a single series of setigerous punctures, but with three or four additional punctures apically. Length, 914 mm. Type locality.—Jamaica. Types.—Holotype in United States National Museum (No. 26331). 194 BULLETIN 182, UNITED STATES NATIONAL MUSEUM FPecords.—The following are the records known to me: Jamaica: (Notman, 1925; British Museum; U.S.N.M.). Specimens examined. —I have seen only the unique type and the specimen mentioned below. Remarks.—This species has not been reported since its description in 1925. It is distinct by the horned clypeus from all but notmani, from which it differs in size and abdominal punctation. The specimen in the British Museum labeled “Osorius hubbardi Nordm.” was borrowed and compared directly with the type. It seems to be identical. It was received by the British Museum in 1845 but bears no other data. I find no record of the habits of this species. 24. OSORIUS NOTMANI Darlington Osorius notmani DARLINGTON, 1937, pp. 287, 297. Description.—Piceous or rufous. Head about five-sixths as wide as pronotum; eyes not at all prominent; clypeus with angles produced in conspicuous slender teeth about one-fourth as long as the distance between them, each tooth with a denticle at base on inner side, margin not crenulate; labrum scarcely emarginate, alutaceous only at sides, unusually large; apical antennal segments about as wide as long; strongly alutaceous anteriorly, more feebly on vertex; with rather coarse and dense punctures except at front and center. Pronotum less than one-tenth wider than long, much narrowed behind, base two-thirds or three-fourths as wide as apex; sides broadly rounded throughout; basal angles broadly rounded; lateral margins not at all enlarged posteriorly; with trace of impressed midline in part; not very distinctly alutaceous; moderately coarsely but not closely punc- tate; midline impunctate, margined by an irregular series of punc- tures. LHlytra about one-tenth narrower than pronotum, as long as wide, slightly expanded ; coriaceous and punctured a little more finely and closely than pronotum (but very shallowly). Abdomen very feebly and irregularly alutaceous, moderately closely and shallowly punctate except at apex. Anterior face of front tibia with setigerous punctures in a single slightly irregular series. Length, 6 mm. Type locality —Jamaica, near Blue Mountain Peak, 5,000-7,388 feet elevation. Types.—Holotype (M.C.Z. No. 20241) and one paratype in the Museum of Comparative Zoology. Records.—The following is the only record known to me: Jamaica: Blue Mountain Peak (Darlington, 1937). Specimens examined.—I have seen only the paratype from the Museum of Comparative Zoology. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 195 Remarks.—This species is also very distinct. It is similar only to hubbardi in the structure of the clypeus, but differs from that and other Osorius by the rounded basal angles of the pronotum. I find no record of its habits. 25. OSORIUS LAEVICEPS Notman Osorius laeviceps NorMANn, 1925, p. 7, 15, 25.—ScHEERPELTZ, 1933, p. 1184.—Dar- LINGTON, 1937, pp. 287, 301. Description.—Piceous, elytra frequently rufescent. Head ten- elevenths as wide as pronotum; eyes relatively prominent; clypeus truncate, angles not prominent; labrum truncate, alutaceous; outer antennal segments about as wide as long; very coarsely and densely punctate except at middle; without trace of ground sculpture. Pro- notum one-tenth wider than long, strongly narrowed posteriorly, base three-fourths as wide as apex; sides feebly sinuate; basal angles rounded; lateral margin not wider posteriorly; with impressed mid- line outlined by serial punctures; with coarse impressed punctures; generally separated by about their diameter; without trace of ground sculpture. lytra a little wider than pronotum, one-sixth longer than wide, sides moderately arcuate; with coarse shallow punctures generally separated by about their diameter; slightly coriaceous but not alutaceous. Abdomen with large shallow dense punctures, not alutaceous. Anterior face of front tibia with setigerous punctures in single series. Length, 314 to 334 mm. Type locality.—Puerto Rico, San Juan. Types.—Holotype in American Museum of Natural History. FRecords—The following are the records known to me: Hispaniola: DoMINICAN REPUBLIC, SAnchez (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: San Juan (Notman, 1925; Darlington, 1987), Adjuntas (Black- welder station 49), Maricao (Blackwelder station 47A), Mayagiiez (Black- welder station 50B, 50C), El Yunque (Darlington, in M.C.Z. and U.S.N.M.) Specimens examined.—I have examined the type in the American Museum of Natural History, 28 examples from the Museum of Com- parative Zoology, and 40 collected by me in 1935-87. Remarks.—My specimens have not been directly compared with the type but I believe them to be the same. The above description was taken principally from a specimen from Mayagiiez. Notman records that the type was taken by sifting. My speci- mens were collected from dung, from rotting cocoa pods, and in quantity from the rotting “fruit” of an imported palmlike tree. 26. OSORIUS JAMAICENSIS, new species Description—Piceous, pronotum and elytra rufopiceous. Head not distinctly narrower than pronotum; eyes relatively prominent; clypeus truncate, angles not prominent; labrum feebly emarginate 196 BULLETIN 182, UNITED STATES NATIONAL MUSEUM above declivity, smooth except at sides; outer antennal segments transverse; with very coarse irregular punctures except on median area; with traces of alutaceous sculpture among the punctures. Pro- notum scarcely wider than long, moderately narrowed posteriorly, base two-thirds as wide as apex; sides rather evenly rounded, basal angles narrowly rounded; median smooth stripe indefinite; with very large punctures, more or less elongate; feebly alutaceous at sides. Elytra as wide as pronotum, about one-sixth longer than wide; not much enlarged posteriorly; with large and more or less serial punc- tures obscured by strongly coriaceous surface and traces of aluta- ceous ground sculpture. Abdomen only very vaguely alutaceous; moderately and sparsely punctate. Anterior face of front tibia with a single series of setigerous punctures. Length, 314 mm. Type locality—Jamaica, Roaring River Falls, parish of St. Anns. Types.—Holotype, U.S.N.M. No. 52472, collected on September 13, 1917, by Harold Morrison. Records.—The following is the only record known to me: Jamaica: Roaring River Falls (Morrison, in the U.S.N.M.). Specimens examined.—I have seen only the unique type. Remarks.—Like others of the small Osorius this is not distin- guished by striking characters. When large series of all these species are available, it should be possible to evaluate the feeble characters of punctation and sculpture and decide the status of these segregates that we now believe to be species. I have no record of its habits. 27. OSORIUS EGGERSI Bernhauer Osorius eggersi BERNHAUER, 1904, p. 19.—BERNHAUER and ScHusBert, 1911, p. 144.— CAMERON, 1913, p. 8326.—Lena and Mutcuter, 1914, p. 404—NotMAn, 1925, pp. 6, 22, 25.—ScHeEerPEttz, 1933, p. 1133.—Dartineton, 1937, pp. 287, 298, 299. Description.—Piceous, pronotum and elytra rufous. Head scarcely narrower than pronotum; eyes a little prominent; clypeus truncate, angles not prominent; labrum truncate, smooth except at extreme sides; outer segments of antennae wider than long; with sparse coarse punctures especially at sides; obsoletely alutaceous. ‘Pronotum a trifle longer than wide, rather strongly narrowed to base, base two- thirds as wide as apex; sides straight in front, arcuate behind, angles broadly rounded; lateral margin not wider basally; disk with median impunctate stripe bordered by series of large punctures; with coarse punctures, sometimes elongate or uniting longitudinally, separated transversely by about their diameter; with only a trace of alutaceous ground sculpture. Zlytra nearly one-tenth wider than pronotum. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 197 nearly one-sixth longer than wide; expanded but sides straight in front; with coarse shallow punctures, more or less serial; not aluta- ceous, but coriaceous. Abdomen not alutaceous except feebly at apex; moderately coarsely and densely punctate, punctures not muricate. Anterior face of front tibia with a single series of small setigerous punctures. Length, 3 to 3144 mm. Type locality —‘Insula St. Thomas.” Types.—In collection of Dr. Bernhauer. Records —The following are the records known to me: Cuba: Santiago (Notman, 1925; Darlington, 19387; A.M.N.H.). Jamaica: Montego Bay (Notman, 1925; Darlington, 1937; A.M.N.H.), Trinity- ville (Blackwelder station 428), Moneague (Blackwelder station 375). Hispaniola: Harri, Kenskoff (Darlington, 1937). Puerto Rico: (U.S.N.M.). St. Thomas: (Bernhauer, 1904; Leng and Mutchler, 1914; Notman, 1925; Dar- lington, 1937). Central America: PANAMA (Notman, 1925; Darlington, 1987; U.S.N.M.). Specimens examined.—I have seen 10 examples in the United States National Museum and 8 collected by me in 1935-37. Remarks.—This description was drawn from a specimen from Trin- ityville, Jamaica. I am unable to distinguish it from examples of eggersi from Panama. My examples were taken flying at dusk. 28. OSOQRIUS LEWISI, new species Description.—Piceous, pronotum and elytra more or less pubescent. Head one-eighth narrower than pronotum; eyes somewhat prominent; clypeus feebly evenly emarginate, angles tuberculate; labrum very short, emarginate above declivity; outer segments of antennae trans- verse; with scattered rather coarse shallow punctures obscured by ground sculpture which becomes very strong in front. Pronotwm one-twelfth broader than long, rather strongly narrowed to base, base three-fifths as wide as apex; sides evenly arcuate, basal angles broadly rounded; lateral margin not wider basally; disk with median im- punctate stripe bordered by fairly definite series of punctures; with irregularly spaced and somewhat elongate moderately coarse punc- tures, separated transversely by more than their diameters; with very dense and fine ground sculpture. Hlytra one twenty-fourth wider than pronotum, one twenty-fifth longer than wide, feebly expanded with the sides evenly arcuate; with very large shallow punc- tures more or less serial obscured by coriaceous sculpture. Abdomen without distinct alutaceous sculpture except basally, punctures muri- eate or tuberculate. Anterior face of front tibia with a single series of setigerous punctures. Length, 334 mm. Type locality —Grand Cayman, west end of Georgetown. 198 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Types.—Holotype, Oxford University, collected on April 25, 1938, by C. B. Lewis and G. H. Thompson. Records.—The following is the only record known to me: Cayman Islands: Granp CAYMAN (Lewis, in Oxford University collection). Specimens examined.—I have seen only the holotype. Remarks.—This species is very similar to eggerst. It differs in the very evident sculpture of the front of the head and the nature of the abdominal punctures. The type was taken in a light-trap. 29. OSCRIUS WEBERI, new species Description.—Rufocastaneous, head somewhat picescent. Head one-seventh narrower than pronotum; eyes a little prominent; cly- peus evenly and rather deeply emarginate, the angles prominent; labrum truncate, slightly emarginate above the declivity, smooth except at sides; outer segments of antennae scarcely as long as wide; without distinct punctures but with sparse irregular tubercles basally and behind the antennal ridges, densely and rather coarsely aluta- ceous. Pronotum about one-seventh wider than long, moderately narrowed to base with the sides feebly sinuate throughout; apex one- fourth wider than base; the hind angles moderately rounded; lateral margins not abrupt but distinctly wider basally; disk with median impunctate stripe irregularly outlined and somewhat elevated in basal half; with rather large, usually elongate, but very shallow and irregular punctures, more or less united along median band; with ground sculpture feeble except inside punctures. Llytra about as wide as pronotum, one-fifth longer than wide, widest just behind middle but sides straight before and behind; surface very uneven, without distinct punctures but with a few tubercles laterally; with traces of ground sculpture on the coriaceous surface. Abdomen with feeble ground sculpture and sparse rather indistinct large punctures. Anterior face of front tibia with an indistinct series of setigerous punctures. Length, 4 mm. Type locality —Trinidad, 12 miles north of Arima. Types.—Holotype, Museum of Comparative Zoology, and one para- type, United States National Museum (No. 52490), collected by N. A. Weber on June 23, 1935. Records —The following is the only record known to me: Trinidad: Arima (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species is readily distinguished from eggersi by its shallow pronotal punctures, its larger size, and the more deeply emarginate clypeus. I have received no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 199 30. OSORIUS TRINITATIS, new species Description—Castaneous, head, disk of pronotum, and apical third of elytra picescent. Head scarcely narrower than pronotum; eyes a little prominent; clypeus evenly but only moderately deeply emargi- nate, the angles somewhat prominent but not at all tuberculate; labrum truncate, slightly emarginate above the declivity, with traces of ground sculpture throughout; outer segments of antennae much wider than long; with rather indefinite coarse shallow punctures obscured by very indefinite ground sculpture. Pronotwm one-six- teenth wider than long, moderately narrowed to base with the sides nearly even and only slightly arcuate; apex about one-third wider than base; the hind angles broadly rounded; margin not distinctly wider basally; disk with median impunctate stripe irregularly out- lined, elevated only near base; with moderate and round but irregu- lar and very shallow punctures, not coalescing but somewhat obscured by traces of ground sculpture. Z/ytra one-sixteenth wider than pronotum, one-tenth longer than wide, widest at apical fourth, sides feebly arcuate; surface very uneven, without distinct punc- tures but with indistinct tubercles throughout, with traces of minute ground sculpture on the coriaceous surface. Abdomen with distinct moderate punctures separated by about twice their diameter and somewhat obscured by the indistinct ground sculpture. Anterior face of front tibia with a regular series of setigerous punctures placed a little laterally. Length, 3 mm. Type locality —Trinidad, 10 miles north of Arima, elevation 1,800 feet. Types.—Holotype, in Museum of Comparative Zoology, and one paratype, United States National Museum (No. 52501), collected on March 28, 1935, by Dr. N. A. Weber. Records. —The following is the only record known to me: Trinidad: Arima (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species differs from weberi very greatly in the sculpture of the head, as well as in size and the shape of the antennal segments. I have received no record of its habits. 31. OSORIUS EXIGUUS Notman OsOrius exiguus NoTMAN, 1925, pp. 7, 16, 25.—ScHEERPELTz, 1933, p. 1183.—Dar- LINGTON, 19387, pp. 287, 300. Osorius micros NOTMAN, 1925, pp. 7, 22, 25—Dartinaton, 1937, pp. 287, 301. (Not Sharp, 1887.) Description.—Rufopiceous to rufotestaceous. Head nine-tenths as wide as pronotum; eyes scarcely at all prominent; clypeus truncate, 200 BULLETIN 182, UNITED STATES NATIONAL MUSEUM angles not prominent; labrum short, coarsely alutaceous; outer antennal segments transverse; rather coarsely and densely but very shallowly punctate; not alutaceous. Pronotum one-fourteenth wider than long, rather strongly narrowed posteriorly, base five-sevenths as wide as apex, basal angles rounded; with narrow and _ indefinite median impunctate stripe; with coarse, shallow punctures, generally separated by less than their diameter; not alutaceous. H’lytra as wide as pronotum, one-fifteenth longer than wide, sides nearly parallel; with very indefinite shallow punctures obscured by coriaceous texture ; not distinctly alutaceous. Abdomen with moderate but dense punc- tures; coriaceous but not alutaceous. Anterior face of front tibia with an indefinite series of setigerous punctures, Length, 2 mm. Type locality—Cuba, Cayamas, about 15 miles west-northwest of Cienfuegos, province of Santa Clara. Types.—Holotype and one paratype in the United States National Museum (No. 263837). Records.—The following is the only record known to me: Cuba: Cayamas (Notman, 1925; U.S.N.M.). Specimens examined.—I have seen only the two types and possibly a third specimen. Remarks.—With the types of this species I have associated the specimen that Notman recorded as O. micros Sharp. I can find nothing to distinguish it except its length of 3mm. I have compared this specimen (micros of Notman) directly with the type of micros Sharp in the British Museum and made the following notes at that time: “It (Notman’s specimen) differs in lacking the two discal rows of very strong punctures, and in lacking the dense ground sculpture of the head. The elytra are not so coarsely and irregularly punctured. Pronotum also lacks ground sculpture.” The size of the type of micros Sharp is 324 mm. QO. micros Sharp should therefore be removed from our list, and the O. micros of Notman should be placed as a synonym of exiguus Notman. I find no record of the habits of this species. 32. OSORIUS EXALATUS, new species Description —Rufocastaneous. Head as wide as pronotum; eyes very small, scarcely prominent though abrupt; clypeus produced into a lobe, which is broadly semicircular except for a prominent median angulation; labrum large and semicircular, with sparse and feeble scaly ground sculpture; outer segments of antennae wider than long; with fine but scattered punctures and with traces of ground sculpture. Pronotum one-fourth wider than long, rather strongly narrowed to base, base two-thirds as wide as apex; sides straight in front, arcuate MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 201 or sinuate behind, the hind angles rather broadly rounded; lateral margin not abrupt but much wider posteriorly; disk slightly flat- tened medially in front but midline not smooth; with minute irregular punctures, and with distinct ground sculpture medially. Hlytra closely appressed to body, almost as wide as pronotum, nearly one- fourth wider than long, scarcely expanded posteriorly; without dis- tinct punctures or sculpture but surface both coarsely and finely coriaceous. Abdomen with sparse and rather fine punctures and a few larger muricate punctures; with alutaceous lustre. Anterior face of front tibia without visible setigerous punctures. Length, 2144 mm. Type locality —Trinidad, Aripo Valley, elevation 2,000 feet. Types.—Holotype, Museum of Comparative Zoology, collected by N. A. Weber on April 19, 1935. Records.—The following is the only record known to me: Trinidad: Aripo Valley (Weber, in M.C.Z.). Specimens examined.—l\ have seen only the holotype. Remarks.—This is another species like webevi, which has the elytra small and closely applied to the body. It has the aspect of a subter- ranean beetle. It is distinct by its labrum and impunctate tibiae from all save webert and micropterus and from these by the absence of the medium pronotal groove. It is distinct from micropterus further by the shape of the clypeus. The type was taken in a cave. 33. OSORIUS MICROPTERUS, new species Description.—Testaceocastaneous. Head as wide as pronotum; eyes small, not very prominent though abrupt; clypeus a broad smooth semicircular lobe; labrum broadly rounded, indefinitely sculp- tured; outer antennal segments broader than long; with sparse and irregular but distinct moderately coarse punctures; with ground sculpture behind the eyes but only very vaguely on the disk. Pro- notum one-tenth wider than long, strongly narrowed posteriorly, apex nearly one-half wider than base; sides very feebly but regularly arcuate; basal angles scarcely evident; disk rather deeply impressed along midline almost to base, without very distinct punctures but with faint traces of ground sculpture which appears to be in part composed of minute punctulae. Llytra closely appressed to body, just wider than the pronotum, not quite as long as wide; feebly en- larged near middle; surface coarsely uneven but without distinct punctures or tubercles on the disk; finely but irregularly coriaceous with traces of ground sculpture. Abdomen with isolated shallow punctures; shining along midline but laterally with faint coriaceous ground sculpture. Anterior face of front tibia without visible seti- gerous punctures. Length, 1°, mm. 449008—42——14 202 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Type locality —Trinidad, Ortocre River, 5 miles southeast of Rio Claro. Types—Holotype, Museum of Comparative Zoology, collected by N. A. Weber on January 3, 1935. Records.—The following is the only record known to me: Trinidad: Rio Claro (Weber, in M.C.Z.). Specimens examined,—I have seen only the holotype. Remarks.—This is the smallest Osorius I have scen. It is un- usually distinct because of its rounded clypeus and the smooth shin- ing appearance of the integuments under low magnification. I have received no record of its habits. Subfamily STYLOPODINAE This subfamily has been known under the names Megalopinae and Megalopsidiinae, with the type genera Megalops and Megalopsidia, respectively. The present change of name of the subfamily to agree with that of the type genus (as explained under Stylopodus) is not required by the International Rules of Zoological Nomenclature but is in keeping with the interpretation followed by many American taxonomists. Only one genus has been described in this subfamily, and it is represented in our region by at least six species. It may be readily recognized by its very large and prominent eyes, transversely sulcate pronotum, and shining integuments. XXVI. Genus STYLOPODUS Benick Stylopodus Brnick, 1917, p. 190 (as subgenus of Megalops). Subgenus Megalopsidia Lene, 1918, p. 204 (as new name for Megalops). Megalops Erxtcuson, 1840, p. 751. (Not Lacepéde, 1803.) Aulacotrachelus BENICK, 1921, p. 1 (as new name for Megalops). Subgenus Gata, new name. Perostylus BENIcK, 1917, p. 190. (Not Pilsbry, 1894.) Subgenus Megalopsidiella BERNHAUER, 1933, p. 333. Genotypes.—Megalops cephalotes Erichson=Stylopodus cephalotes (Erichson) (original designation). Of Megalops, Oxyporus caelatus Gravenhorst= Megalops caelata (Gravenhorst) (designated by Ben- ick, 1917) ; of Megalopsidia and Aulacotrachelus, same as of Megalops, under section II, f, of Rule 30 of the International Rules of Zoo- logical Nomenclature; of Perostylus, Megalops praedita Sharp= Stylopodus (Perostylus) praeditus (Sharp) (original designation) ; of Gata, same as of Perostylus, under same rule as above; of Afega- lopsidiella, Megalopsidia ogloblini Bernhauer=Stylopodus (Megalop- sidiella) ogloblini (Bernhauer) (monobasic and original designation). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 203 Diagnosis —Body in part subcylindrical, glabrous; eyes very large and prominent, antennae 11-segmented, clavate; clypeus feebly sepa- rated from vertex, anterior angles toothed; labrum visible only as two slender processes; maxillary palpus filiform, first segment short; mentum oblong; gular sutures strongly converging at base but anterior to pits are more widely separate and parallel; prothorax subcylindri- cal; pronotum transversely sulcate; prosternum with a transverse row of punctures in front of the coxae; anterior coxal cavities separated by a slender process of the prosternum, half open behind; anterior coxae very small but prominent, without sulcus on anterior face; meso- sternum carinate; middle coxal cavities separated by union of meso- sternal and metasternal processes; posterior coxae contiguous, small, transverse, not expanded under femora; abdomen margined; first and second sternite absent; third sternite carinate at middle basally ; sexes not greatly differing; tibiae unarmed, tarsi 5-segmented. Remarks.—This is a very distinct genus in regard to a great many characters. I know of no important summary or study of its struc- ture, but the following can be mentioned as being of unusual structure or development: The bicornute labrum, the toothed clypeus, the cla- vate antennae, the gular sutures, the prosternum and coxal cavities, the mesosternal and metasternal processes, and the fourth tarsomere. This genus was described by Erichson under the name AMegalops and was known by that name for 90 years. In 1918 Leng found it to be preoccupied and renamed it Megalopsidia. In 1921 Benick came to the same conclusion and offered the new name Avwlacotrachelus, in apparent ignorance of Leng’s proposal. The genus now stands in all recent works as Megalopsidia Leng. However, in 1917 Benick pro- posed simultaneously two new subgenera, Sty/opodus and Perostylus. Nomenclaturally, these must be considered as available names with priority over Megalopsidia and all later names. It is therefore necessary to choose one of these two for the name of the genus and leave the other (with Megalopsidia and the still later AMegalopisidi- ella) as subgenera. Inasmuch as Perostylus is itself a homonym, Stylopodus is the first available name and must be accepted for this genus. The only name that escapes unchanged from this reorgani- zation is that of the subgenus Megalopsidiella. I know of 6 species from the West Indies and have examined 36 examples from the islands beside a considerable series from South and Central America. KEY TO WEST INDIAN SPECIES OF STYLOPODUS te Whvirasentirelysimpunctaics= ste sere ee ee ee ee See 2 a es ee 2 Hivinaawithysome: punctines 222s ees Sean ea ee ee 3 2. Elytra with large testaceous discal area__.________-_____-__-_______- 1. smithi Hiytra*unicolorous; piceocastanéous22— 22 2 hava) ee 2. impressus 204 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 3. Elytra testaceous, at most with margins black___--____________ 3. humeralis Elytra black, at most with several testaceous spots______-_________--___--- 4 #4. JRunctureszof, elytra inva; 2roup,0n: disk. <2 eee ee ee eee 5 Punctures' of elytra scattered. ys 6. rambouseki 5. Elytral punctures arranged in two distinct series_______________ 5. punctatus Elytral punctures not arranged in two distinct series________ 4. laeviventris 1. STYLOPODUS SMITHI (Cameron) Megalops smithi CAMERON, 1913b, p. 3828.—LENe and MurcH er, 1917, p. 198. Megalopsidia smithi (Cameron) SCHEERPELTZ, 1933, p. 1143. Description.—Black, elytra yellow with humeral callus, base, suture, sides, and apical margins narrowly rufopiceous. Head with eyes broader than elytra; ninth and tenth segments of antennae transverse; vertex in front with a circular impression enclosing a smooth round central elevation; with another elevation above each eye posteriorly ; impressions of vertex not very distinctly punctate; clypeus obtusely produced at center, with prominent projection at lateral angles. Pronotum subeylindrical, sides bituberculate in front; as wide as long; with four transverse punctate grooves, the first very feeble along the margin, the second scarcely interrupted at middle but finely punc- tate, third interrupted at middle by a ridge and more coarsely punc- tate, fourth along posterior margin not interrupted. //ytra one-third wider than pronotum, one-third wider than long; humeral callus prominent but sutural callus absent; sutural stria distinct but disk entirely without impressions or punctures. Abdomen impunctate, shining. Length, 3144 mm. Type locality.—St. Vincent. Types.—One example labeled type in the British Museum. Eleven other specimens in the same series have the importance if not the actual status of paratypes. Four additional specimens were found in another series in the same collection. Records.—The following is the only record known to me: St. Vincent: (Cameron, 1913; British Museum). Specimens examined —\ have seen only the 16 examples in the British Museum. Remarks.—This species seems distinct by its entirely smooth and even elytra. The example I borrowed from the British Museum is labeled “Lot 14, Aprl. 5. 400 ft. Under bark. Shady glen.” 2. STYLOPODUS IMPRESSUS (Sharp) Megalops impressus SHARP, 1876, p. 380; 1887, p. 676.—BeRNHAUER and ScHu- BERT, 1911, p. 151. Description —Black to rufopiceous throughout. Head with eyes one-tenth wider than elytra; vertex in front with a circular impres- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 205 sion posteriorly and enclosing a smooth rounded elevation; with another elevation above each eye posteriorly; impressions not very distinctly punctate; ninth and tenth antennal segments transverse ; clypeus produced at center into an angle which is nearly acute, lat- eral angles strongly produced. Pronotum subcylindrical, one-twelfth wider than long, sides scarcely tuberculate but with prominences; with four strong and deeply punctate transverse grooves, all but the fourth interrupted narrowly at middle; intervals very convex, shin- ing. Elytra one-third wider than pronotum, one-fourth wider than long; entirely impunctate, but with a distinct longitudinal impres- sion at middle of disk, abrupt internally, slightly diagonal; sutural stria impressed ; humeral callus prominent, but without sutural callus. Abdomen impunctate, shining. Length, 314 to 5 mm. Type locality —‘Villa Nova,” Amazon Valley, Brazil. Types.—One female in the British Museum. Records—The following are the records known to me: Trinidad: (C. E. Bryant, in British Museum), La Brea (Morrison, in U.S.N.M.). South America: Braziu (Sharp, 1876, 1887), Botrvra (W. M. Mann, in U.S.N.M.). Central America: PANAMA (Sharp, 1887). Specimens examined.—I have not examined the unique type or Sharp’s example from Panama. Specimens before me include one borrowed from the British Museum and two in the United States National Museum. Remarks.—There is very little difference between the Trinidad and the Bolivia examples. The species is obviously widespread and will undoubtedly be found throughout northern South America. Sharp records the type from under chips. 3. STYLOPODUS HUMERALIS (Cameron) Megalops humeralis CAMERON, 1913b, p. 327.—Lene and Murcuter, 1917, p. 198. Megalopsidia humeralis (Cameron) SCHEERPELTz, 1933, p. 1142. Description—Black, elytra yellow with humeral callus, base, and apex narrowly black. Head and antennae much as in laeviventris. Pronotum a little longer than broad, a little wider in the middle, equally narrowed in front and behind; with sides evenly rounded; narrower than head and elytra; with four coarsely punctate trans- verse impressions on each side, the first are separated from each other by narrow, smooth space on middle of disk; the second are separated by a broader space bearing a large puncture, the third by a wider im- punctate space, the fourth follow the posterior margin and are not interrupted. #lytra about as long as pronotum, scarcely transverse ; sutural callus, sutural stria, and short punctate stria on disk well marked. Abdomen as in Jlaeviventris. Length, 4 mm. (From Cameron. ) 206 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Type locality—St. Vincent. Types.—One specimen labeled type in the British Museum. Records.—The following is the only record known to me: St. Vincent: (Cameron, 1913). Specimens examined.—I have seen only the unique type. Remarks.—When I examined the type I made a note that the punc- tate stria on the elytron has three punctures but is entirely lacking on one elytron. This species is similar in coloration to smzthi, and I regard it as a doubtful species. The length of the “clypeal spines” is of no value whatever, and there appears to be some variability in the impressed punctures of the head and pronotum. I find no record of its habits. 4. STYLOPODUS LAEVIVENTRIS (Cameron) Megalops laeviventris CAMERON, 1913b, p. 326.—LENG and Mutcutrr, 1917, p. 198. Megalopsidia laeviventris (Cameron) SCHEERPELTZ, 1933, p. 1142. Description —Black, shining. ead with eyes nearly as wide as elytra; vertex in front with a feeble broad semicircular impression enclosing the anterior end of a large elevation; elevation with two or three punctures impressed in transverse row; with distinct punc- tures; antennal segments 9 and 10 transverse; clypeus truncate, angles acutely but only moderately produced. Pronotwm subcylin- drical, as long as wide; sides not tuberculate but with prominence at sides anteriorly; with four transverse grooves, first feeble and impunctate, second obsolete except for three or four small depres- sions on each side, third moderately deep with punctiform depres- sions laterally, interrupted and deflected at middle, fourth basal, moderate, punctate; intervals moderately convex, shining. lytra nearly one-half wider than pronotum, over one-fifth wider than long; humeral callus moderate; sutural stria impressed; discal group of punctures separated from suture by distinct elevation; group con- sisting of three or four punctures. Abdomen impunctate except feebly on apical segments. Length, 314 to 4 mm. Type locality—St. Vincent. (Cameron records the locality as Grenada but the labeled type is from St. Vincent.) Types.—One example labeled type in the British Museum. Ten other specimens in the same series have the importance if not the actual rank of paratypes. Three additional specimens are in the same collection. Records.—The following are the records known to me: 4 As later pointed out by Dr. Cameron (1930, p. 315) the so-called clypeal spines are in reality elongate processes of the sides of the labrum. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 207 St. Vincent: (British Museum). Grenada: (Cameron, 1913; British Museum). Specimens examined. —I have seen only the 14 examples in the British Museum, of which one was borrowed for further study. Remarks.—This species seems amply distinct from the other West Indian species. The pronotal grooves are feebler than in any other of our species. I find no record of its habits. 5. STYLOPODUS PUNCTATUS (Erichson) Megalops punctatus ErtcHson, 1840, p. 752.—Lacorpaire, 1854, p. 111, pl. 15, fig. 4 —Favvet, 1901, p. 74—BrrnyHaver and ScHusert, 1911, p, 151—Cam- ERON, 1913, p. 328.—BeENIcK, 1917, pp. 191, 192.—ScHEERPELTz, 1933, p. 1143. Megalopsidia (s. str.) punctata (Erichson) SCHEERPELTZ, 1983, p. 1143. Description—Piceous, elytra fuscous, sometimes with humeral spot and apical sutural spot testaceous. Head with eyes a little nar- rower than elytra; clypeus truncate but with angles shortly acutely produced and deflected; vertex without linear impressions but with numerous deep coarse punctures; no individual elevations except over base of antennae; ninth and tenth antennal segments transverse, eighth as broad as long. Pronotum subcylindrical, sides not tubercu- late but with prominences anteriorly; scarcely wider than long; with four feeble transverse punctate impressions, first and third inter- rupted at middle, second and third deflected posteriorly on disk, punctures coarse, intervals convex, shining. “7/ytra one-third wider than pronotum, nearly three-tenths wider than long; humeral callus feeble; sutural stria impressed; with two rows of four large punc- tures at center of disk. Abdomen impunctate, shining. Length 3144 mm. Type locality —Colombia. Types —In either the Hope Museum, Oxford, cr the Zoologische Museum, Berlin. Records.——The following are the records known to me: Trinidad: Moruga (Blackwelder station 111). Curacao: (Fauvel, 1901). South America: Brazin (Fauvel, 1901), Coromsra (Erichson, 1840; Lacordaire, 1854; Fauvel, 1901). Central America: Mexico (Fauvel, 1901). Specimens examined—I have seen only two specimens from Trinidad. Remarks.—I have little doubt of the identification of this species because of the exact agreement with Erichson’s description and the ease of separating it in the key. My specimens were found in decaying cocoa pods. 208 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 6. STYLOPODUS RAMBOUSEKI (Bierig) Megalopsidia rambouseki Brera, 1934e, p. 220, pl. 11, figs. 7, 8. Megalopsidia bierigi Rambousek MS., Breric, 1934e, p. 220. Description —Black, variegated with transparent rufous on most of the elevated parts, elytra with sutural region yellawish and with large ivory-white spot at humeral callus, with same color on posterior half of seventh and following abdominal segments. /Zead with eyes only a little narrower than elytra; vertex with two semielliptical tubercles, oblique and touching or confluent behind; intervals sepa- rately and coarsely punctate. Pronotum subhexagonal, widest at apical third; sides with blunt denticles and protuberances; of the four bulky elevations, the two approximate anterior ones are clearly hemispherical, the two basal, separated by a protuberance, more en- larged, are obliquely oval, with exterior face subvertical, and almost contiguous to the marginal postlateral protuberances; intervals feebly punctate. Hlytra a little wider than pronotum; with two oblique postscutellar callosities, the suture simply elevated, flanked by a stria and with a posterior prelateral callus, semicircular from the interior. Length,4mm. (From Bierig.) Type locality—Cuba. Types.—tIn collection of Alexander Bierig. Fecords.—The following are the records known to me: Cuba: Rio Hondo (Bierig, 1934), Sierra Bonilla (Bierig, 1934), Sierra del Rosario (Bierig, 1934). Specimens examined.—I have seen no examples of this species. Remarks.—This species is known to me only from Sr. Bierig’s description and figures. It seems to me amply distinct from the other West Indian species. It was placed by Bierig in the subgenus Megalopsidia. It was recorded from fungus on tree trunks. Subfamily STENINAE Two genera have been described in this subfamily but only one is so far known from the West Indies. Like the preceding it is readily recognized by its large and prominent eyes but differs in having the integuments more densely punctate or rugose, in having the pronotum not transversely sulcate, and in other characters given in the diagnoses. XXVIII. Genus STENUS Latreille Stenus LATREILLE, 1796, p. 77. Nestus Rey, 1883, p. 183. Subgenus Hemistenus Morscuutsky, 1860, p. 557. Mesostenus Rey, 1883, p. 183 (not Gravenhorst, 1829). Parastenus Hrypren, 1905, p. 262 (new name for Mesostenus Rey). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 209 Subgenus Hypostenus Rey, 1883, p. 183. Astenus LyYNcH, 1884, p. 341. Stenosidotus LyNcu, 1884, p. 338. Areus Casry, 1884, p. 150. Subgenus J'esnus Rey, 1883, p. 183. Mutinus Casey, 1884, p. 146. Genotypes.—Stenus 2-guttatus Fabricius (designated by Latreille, 1810). Of Nestus, Stenus (Nestus) boops Liungh (designated by Tottenham, 1940) ; of Hypostenus, Stenus (Hypostenus) kiesenwetteri Rosenhauer (designated by Tottenham, 1940); of Astenus, Stenus (Astenus) speculifrons Fauvel (monobasic) ; of Stenosidotus, Stenus (Stenosidotus) aenescens Lynch (monobasic) ; of Hemistenus, Stenus (Hemistenus) pallitarsis Stephens (designated by Tottenham, 1939) ; of Areus, Stenus flavicornis Erichson = Areus flavicornis (Erich- son) (designated here); of Mesostenus, Stenus (Mesostenus) im- pressus Germar (designated by Tottenham, 1939); of Parastenus, Stenus (Parastenus) impressus Germar (by synonymy with Meso- stenus) ; of Tesnus, Stenus (Tesnus) opticus Gravenhorst (designated by Tottenham, 1940); of Mutinus, Stenus (Mutinus) dispar Casey (designated here). Diagnosis.—Body elongate, cylindrical; labrum transverse, rounded in front; antennae 11-segmented, elongate and slender, more or less clubbed; first segment of maxillary palpus elongate, third large, fourth minute; last segment of labial palpus normal; mentum sub- quadrate, strongly carinate at middle; gula broad, sutures conver- gent at middle; anterior coxal cavities confluent, open behind; an- terior coxae small but conical and rather prominent; middle coxal cavities narrowly separated by union of mesosternal and metasternal processes; posterior coxae “conical,” not expanded laterally because of the proximity of the pleurites; first and second abdominal sternites not sclerotized; third carinate; males with apical segments modified ; tarsi 5-segmented. Remarks—This very large and widespread genus is readily recog- nizable by its appearance. The large eyes and the insertion of the antennae into the upper surface of the head near the eyes will dis- tinguish it from all other staphylinids. The genus has been divided into several subgenera, but there is no complete agreement as yet on the characters to be used for this purpose. I have examined 113 West Indian examples belonging to 20 species of which 15 appear to be new. These belong to at least three of the subgeneric groups listed above. KEY TO WEST INDIAN SPECIES OF STENUS ie Abdomenistronelyenianeined = st wen bo the Bl eo eS 2 Abdomenmotdistinctlyamareined = 229-2 4 es se se 4 210 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. es) 10. ti. 12. 13. 14. 15. 16. 1%. 18. 19. Punctures of pronotum and elytra strongly and distinctly united in channels, with intervening ridges very distinct-__.______________________ 2. chapini Punctures of pronotum and elytra not strongly united in channels, ridges not strong and distinet-s <2 2225. As ee Se ee 3 . Densely punctate throughout, subopaque_______-_______________ 3. rulomus Not densely punctate, shining===.. = _ 3s ee eee eee 4. odius . Fourth segment of anterior tarsus strongly bilobed____________________ 5 Mourthyseementsnots pilObe da 2s ees ee ee 1. trinitatis . Head with median::smooth? spaceon: vertex2ue 2-2 ee 6 Head without median smooth space on vertex___--______--_____--___~ 10 . Punctures of pronotum and elytra coarse and deep, similar to those Of Hheads2se 224 Wg 2 Be a, ee ai Punctures of pronotum and elytra fine and scattered, much less coarse than those om, thes ese ee ee ee ee ee See 17. lucens . Abdomen punctured as forebody but less coarsely_____________---______ 8 Abdomen with relatively fine shallow punctures and strong ground sculp- TUT 2: SHEN OE SEE aN ee Ee ead ae 18. dyeri ~whronotum tuberculate at+sides, of bases. 222 -2-=- = 20. antillensis Pronotum not, tuberculatesatssides Of base. 2222" sss = ts eee eee 9 . Head with large and well-defined additional smooth area both at each posterior upper margin of eye and behind each antennal fossa________ 14. cubanus Head without large and well-defined additional smooth area_-____--__ 17 Subopaque: length) 2 mm. 2. 28 2 a ee eee 5. bakeri Shining sength:3) to; b7mm=- bas at oP ee ee 11 Punctures of elytra generally separated by nearly their diameter. 6. jamaicensis Punctures of elytra generally separated by half or less of their diameter. 12 Punctures of pronotum somewhat serial along midline behind center___ 13 Punetures of pronotum not at all serial along midline behind center____ 14 Highth tergite with dense scaly ground sculpture____--______ 11. morrisoni Highth tergite without distinct scaly ground sculpture__-____--_ 12. bryanti Pronotum .as broad 8S) lone sense. oe ee ee eee 10. mutchleri Pronotum’ distinctly longer thane broad -—-- =e ee ee 15 Outline of pronotum from above with a distinct postaplen! protuberance Ont Gach! side ss 252s: i. ees ahs ged ee ais ae aA wees 8. augustinus Outline of pronotum from above not interrupted near apex__-----_---~ 16 Pronotum much more densely punctate than elytra_-_--_-_____- 7. jugalis Pronotum not much more densely punctate than elytra__---_-~ 9. haitiensis Median area of head narrow but very distinct and extending throughout length’! of ‘vertex+.220). 22 OE) a 4) eee 16. darlingtoni Median area not extending throughout length of vertex__-._-_----_~-- 18 Median area very broad, well-defined___________-________ 18. hispaniolus Median. area feeble; ill-defined +" = ="= 2 see ee ee eee 19 Vertex with a small but distinct depression on each side of midline near GOTtOR, Meee 7 eae Ee PR ARE dee a eee 15. adamsoni Vertex scarcely at all’ depressed—+ 23.4)! 220 et ees 19. cubensis 1. STENUS TRINITATIS, new species Description—Shining black. Head rather deeply concave above between the eyes, with a narrow acute median longitudinal ridge; labrum broadly rounded; with moderately fine irregularly shaped and spaced punctures; without ground sculpture or smooth areas except MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Qt for the ridge. Pronotum with punctures indistinct because of fre- quent uniting transversely, giving almost a coarsely wrinkled appear- ance; without ground sculpture; scarcely longer than wide, widest near middle, arcuately narrowed in front with angles obsolete, feebly emarginate behind with the angles obtusely rounded. Llytra scarcely longer than broad, nearly one-third wider than pronotum; with humeral callus feeble; punctures larger than those on pronotum, irregularly shaped, less uniting except at the sides; without ground sculpture. Abdomen not margined; with shallow punctures finer than on elytra and becoming still finer apically; with indistinct ground sculpture. Z’arsus with fourth segment not bilobed; posterior tarsus long, first segment longer than fifth. dale, all sternites flattened, especially apically, eighth with a narrow excision one-third as wide as apex and twice as long as wide, with apex very narrowly rounded. Female, sternites unmodified. Length, 4 mm. Type locality—Trinidad, St. Augustine. Types——Holotype, male, and two paratypes in the Museum of Comparative Zoology; three paratypes in the United States National Museum (No. 52538); all collected in April 1929 by Dr. P. J. Dar- lington. Records.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the six types. Remarks.—This species is very distinct by the concavity of the vertex and the nature of the punctation. It is :solated from all other West, Indian species by having simple tarsi in conjunction with an unmargined abdomen. I have received no record of its habits. 2. STENUS CHAPINI, new species Description.—Shining black. Head rather feebly longitudinally impressed between the eyes; labrum broadly rounded; with very coarse and deep punctures separated by less than half their diameter by narrowly convex intervals, no central smooth space; vertex con- stricted behind; without ground sculpture. Pronotum very coarsely feebly punctured, separated by less than their diameter, with occa- sional intervals lower so that the punctures are somewhat united irregularly; one-sixteenth longer than wide; widest at middle, scarcely narrowed to apex, moderately narrowed to base; without ground sculpture. Llytra nearly one-half wider than pronotum, one-tenth wider than long; with humeral callus feeble and a slight impression mesad; punctures as coarse as on pronotum but more fre- quently and distinctly united in irregular rows; with sparse pu- bescence. Abdomen strongly margined throughout; with moder- 212 BULLETIN 182, UNITED STATES NATIONAL MUSEUM ately large punctures separated by about their diameter by flat inter- vals; pubescence moderate. TZarst with fourth segment not at all bilobed; posterior tarsi moderately long, first segment about as long as fifth. Male, eighth sternite very feebly broadly emarginate. Length, 3 to 384 mm. Type locality —Jamaica, about 5 miles west of Black River (11% miles southeast of Scotts Cove). Ty pes.—Holotype, female, and four paratypes, U.S.N.M. No. 52388, collected by Chapin and Blackwelder on February 24, 1937; and three paratypes, same number, collected by H. G. Hubbard. Records.—The following are the records known to me: Jamaica: (Hubbard, in U.S.N.M.), Black River (Blackwelder station 416), Castle Daly in St. Anns (Darlington, in M.C.Z. and U.S.N.M.), Cinchona (Darlington, in M.C.Z. and U.S.N.M.), Whitfield Hall in Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the eight types in the United States National Museum and seven specimens from the Museum of Comparative Zoology. Remarks.—This species is distinguished from all others in our region by its dense punctation and rugose appearance on both the elytra and the pronotum as well as by its filiform tarsi. It was found by Dr. Chapin and me on the damp muddy edge of a small pond. 3. STENUS RULOMUS, new species Description —Shining black; unusually flattened above. Head feebly longitudinally biimpressed; labrum broadly rounded; with moderately coarse deep punctures irregularly spaced but separated by abrupt intervals which have traces of ground sculpture; without smooth areas. Pronotum punctured as the head but intervals more irregular with punctures occasionally partly united with traces of ground sculpture on intervals; as long as wide, widest at middle, feebly narrowed to apex, feebly emarginate to base. lytra one- ninth broader than long, one-fourth wider than pronotum; with humeral callus indistinct ; punctured and sculptured similarly to pro- notum. Abdomen margined throughout; with moderately small punctures separated by about their diameter, the flat intervals with obsolescent sculpture. 7Z’arsus with fourth segment not bilobed ; pos- terior tarsus rather short, first segment about as long as fifth. Male, sixth sternite just visibly emarginate; seventh with a shallow arcuate emargination in median third, surface depressed before emargination; eighth sternite with an abrupt triangular emargina- tion twice as wide as deep, apex not rounded, sides straight. Fe- male unknown. Length, 3 mm. Type locality—Bahama Islands, Nassau. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 23 Types-—Holotype, male, in Museum of Comparative Zoology ; col- lector and date unknown. Records.—The following is the only record known to me: Bahamas: Nassau (M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species has a very distinct appearance which is due in part to its flattened dorsum. It differs from odius by the male characters and from chapini by the less tortuous elytral punctures. T have received no record of its habits. 4. STENUS ODIUS, new species Description—Shining black. Head rather feebly longitudinally biimpressed between the eyes; labrum rather truncate in front; with very coarse and deep but irregular punctures generally separated by one-half their diameter or less; without distinct smooth spaces; without ground sculpture. Pronotum with deep but irregular punc- tures frequently elongate transversely, the intervals abrupt and ir- regular; one-eighth longer than wide; widest at middle, moderately narrowed to apex, rather abruptly emarginate behind; without trace of ground sculpture. “lytra as wide as long, one-half wider than pronotum; humeral callus feeble; punctures as on pronotum but a little coarser and with intervals somewhat flattened; pubescence rather long. Abdomen strongly margined throughout; with rather small punctures separated by their diameter or more by flat inter- vals; pubescence moderate. Yarsus with fourth segment not at all bilobed; posterior tarsus moderately long; first segment nearly as long as fifth. JMfale, seventh tergite feebly emarginate, eighth with a very broad but shallow and completely rounded emargination. Female, unknown. Length, 334 mm. Type locality—Cuba, Soledad near Cienfuegos. Types.—Holotype, male, in Museum of Comparative Zoology. FRecords—The following is the only record known to me: Cuba: Soledad (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species differs from chapini in having the pro- notal and elytral punctures less uniting and tortuous, and in being much more strongly shining. I have received no record of its habits. 5. STENUS BAKERI Bernhauer Stenus bakeri BeRNHAUvER, 1910, p. 364—BERNHAUER and Scuupert, 1911, p. 172. (Not Bernhauer, 1914.) Hypostenus bakeri (Bernhauer) Lene and MurTcHtIEr, 1914, p. 404. 214 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Description—Similar to alumnus Sharp but differing as follows: Black, subopaque. Head as wide as elytra; more strongly and densely punctate; without metallic lustre; antennae not clavate. Pronotum shorter, scarcely longer than wide; more coarsely and densely punctate. Zlytra much shorter, together somewhat wider than long; more coarsely punctate and much more densely and con- fusedly channeled. Abdomen cylindrical; somewhat more densely punctate. J/ale, seventh sternite somewhat flattened; eighth sternite with a broad triangular emargination, not large and extending not far in the surface of the sternite. Length, 2 mm. (From Bern- hauer.) Type locality.—Cuba, Habana. Types.—One male, presumably in the collection of Max Bernhauer. Records.—The following is the only record known to me: Cuba: (Leng and Mutchler, 1914), Habana (Bernhauer, 1910). Specimens examined.—I have seen no examples of this species. Remarks.—This species was rather briefly described without men- tion of subgeneric characters. These have been assumed from its position in the catalog (Bernhauer and Schubert, 1910, p. 172). If the species is really subopaque, it is quite distinct from all those from the West Indies before me by that character. I find no record of its habits. 6. STENUS JAMAICENSIS, new species Description.—Black, pronotum and elytra generally violet-black. Head not distinctly biimpressed between the eyes; not abruptly ele- vated at edge of eyes; labrum broadly rounded; with very coarse and deep punctures separated generally by less than half their diameter, intervals flat; without distinct impressed areas; without ground sculpture. Pronotum one-third longer than wide, widest just be- hind middle, not greatly narrowed either way; with very coarse and deep punctures, separated by less than half their diameter by evenly convex intervals, without ground sculpture. lytra about one-half wider than pronotum, one-fifteenth longer than wide; with humeral callus scarcely at all elevated and without impressions; with punc- tures as on pronotum but generally separated by their diameter or more; without ground sculpture or pubescence. Abdomen not mar- gined; with basal segments punctured as elytra but with punctures less coarse, becoming less impressed apically; eighth tergite only with distinct scaly ground sculpture; with sparse pubescence only at tip; eighth sternite bordered with a row of minute spinules; basal part of ninth tergite broadly emarginate and bordered with fine spinules of much greater length than on the eighth sternite. Zarsus with fourth segment bilobed. Length, 414 to 5144 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 25 Type locality —Jamaica, between Mocho and Catadupa, in south- ern St. James Parish. Types——Holotype and two paratypes, U.S.N.M. No. 52887, col- lected by Chapin and Blackwelder in 1935-37; two paratypes col- lected by H. G. Hubbard; and one paratype collected by Harold Morrison on September 12, 1917. Records—The following are the records known to me: Jamaica: (Hubbard, in U.S.N.M.), Sharps Grove near Chapelton (Morrison, in U.S.N.M.), Mocho, in St. James Parish (Blackwelder stations 16, 406), Man- chioneal (Blackwelder station 364). Specimens examined.—I have seen only the six types. Remarks—I am unable to distinguish sexes among these six speci- mens. The type and one other have the ninth sternite narrowly excised at apex, but all the others have this segment so far retracted that it is invisible. The type and two others collected by me were all taken by sweep- ing low herbage along the road. On each occasion further collecting failed to produce more specimens. 7. STENUS JUGALIS Erichsen Stenus iugalis EricHson, 1840, p. 736. Stenus jugalis Erichson, FAvuvEL, 1901, p. 74 (as synonym of cupreus Laporte).— BERNHAUER and SCHUBERT, 1911, p. 173 (as synonym of cupreus).—BENICcK, 1917, pp. 300, 301.—ScHErRPELTz, 1933, pp. 1176, 1178.—Benicx, 1988, p. 148. Description—Piceous. Head not distinctly biimpressed but with midline feebly raised; without smooth areas; labrum broadly rounded; punctures rather coarse and deep, regular, generally sepa- rated by about half their diameter; intervals moderately convex, with a faint alutaceous luster; pubescence distinct. Pronotum one- third longer than wide, widest at middle, scarcely narrowed in front, slightly but emarginately narrowed behind; punctures coarse and deep but very dense, separated by less than half their diameter by convex intervals; without ground sculpture or pubescence. Llytra about one-half wider than pronotum, nearly one-sixth longer than wide; humeral callus not prominent; punctation similar to that of pronotum but a little coarser, sparser, and less abrupt, intervals not so convex; without ground sculpture or pubescence. Abdomen not margined ; with same type of punctures as elytra but less coarse and less dense; with distinct ground sculpture on seventh and eighth tergites; pubescence very short. Tarsi with fourth segment strongly bilobed. Male, sixth sternite slightly flattened at middle posteriorly and with an indefinite area of minute sculpture and denser pubes- cence; seventh feebly emarginate at middle, surface flattened and with a longitudinal whitish spongy area; eighth sternite with a nar- 216 BULLETIN 182, UNITED STATES NATIONAL MUSEUM row deep excision, all angles rounded, sides straight. Female, apical sternites not modified. Length, 4144 to 5 mm. Type locality —Colombia. Types.—Either in Hope Museum, Oxford, or Zoologische Museum. Berlin. Records —The following are the records known to me: Trinidad: Montserrat (Busck, in U.S.N.M.), San Rafael (Adamson, as Black- welder station 444F). South America: (British Museum), VENEZUELA (British Museum), CoLoMpBra (Erichson, 1840). Specimens examined.—I have examined six examples in the British Museum, two in the United States National Museum, and one collected by Dr. A. M. Adamson on January 4, 1936. Remarks.—This species appears to be properly associated with Erichson’s name jugalis. I have compared our specimens with the South American examples in the British Museum. I do not have enough material to determine its variability, but I believe it to be fairly constant in the characters used in the key. I find no record of its habits. 8. STENUS AUGUSTINUS, new species Description—Piceous black. Head not impressed between the eyes but vaguely elevated in front; without smooth areas; labrum broadly rounded; punctures moderately coarse and deep, regular, separated by much less than their diameter, intervals rather flat but not sculptured. Pronotum one-fourth longer than wide, widest just behind middle, moderately evenly narrowed in front, broadly emar- ginate behind; punctures coarse and deep but very dense, separated by less than one-half diameter by irregular convex intervals; without ground sculpture. /ytra three-eighths wider than pronotum, one- eleventh longer than wide; humeral callus not at all prominent; punctation similar to that of pronotum but a little coarser and sparser; without ground sculpture. Abdomen not margined; with same type of punctures as elytra but much less coarse and less dense; with traces of ground sculpture on the eighth tergite; pubescence short. Zarsus with fourth segment strongly bilobed. fale, seventh sternite very slightly emarginate; eighth sternite with a large tri- angular notch about as deep as broad, angles narrowly rounded. Female, seventh and eighth sternites not modified. Length, 4 to 414 mm. Type locality —Trinidad, St. Augustine. Ty pes.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology, and one paratype, male, in the United States National Museum (No. 52492); collected in April 1929 by Dr. P. J. Darlington. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE QI Records.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—\ have seen only the three types. Remarks.—The slight pronotal protuberance is not equally strong in the three specimens, but this species can be distinguished from jugalis by its much narrower head, by the absence of the whitish pubescent area on the seventh sternite of the male, and by the wider emargination of the eighth sternite of the male. I have received no record of its habits. 9. STENUS HAITIENSIS, new species Description—Shiny black to uniformly picescent. Mead very slightly depressed between the eyes but without raised midline; with- out smooth areas; labrum rounded but feebly truncate in front; punctures moderate but irregular, sometimes separated by nearly their diameter, intervals flat and shiny. Pronotum three-eighths longer than wide, widest just before middle, feebly narrowed before and behind, sides feebly emarginate behind; punctures moderately coarse and deep, irregularly separated by one-half their diameters by feebly convex intervals; without ground sculpture. Hlytra one- half wider than pronotum, one-twelfth longer than wide; humeral callus very feeble; punctation similar to that of pronotum but coarser, not at all serial; without ground sculpture. Abdomen not margined; with same type of punctures as pronotum but less coarse and less dense; with distinct ground sculpture on seventh and eighth tergites; pubescence very short. 7Z’ars? with fourth segment strongly bilobed. Male, sixth and seventh sternites densely and finely punc- tate along middle, seventh very feebly emarginate, eighth with an abrupt triangular emargination more than twice as deep as wide. Female, sixth and seventh sternites punctate as in male, eighth truncate. Length, 4 to 5 mm. Type locality—Haiti, northeastern foothills of the Massif de La Hotte; elevation 2,000 to 4,000 feet. Types.—Holotype and three paratypes in Museum of Comparative Zoology, collected on October 10-24, 1934, by Dr. P. J. Darlington. Two paratypes with same data in the United States National Mu- seum (No. 52493). Records—The following are the records known to me: Hispaniola: Hart, Massif de La Hotte (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—lI have seen only the six types. Remarks.—This species differs from jugalis principally as indi- eated in the key, but also in the more abrupt and narrower emargina- tion of the eighth sternite of the male and the feeble development of the spongy area on the seventh sternite of the male. I have received no record of its habits, 449008—42-—15 218 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 10. STENUS MUTCHLERI, new species Description.—Shiny piceous black. Head scarcely at all impressed between the eyes; labrum broadly rounded; with very coarse and deep punctures separated by much less than their diameter by feebly convex intervals; without distinct smooth areas; without ground sculpture. Pronotum with coarse and deep punctures, very narrowly separated, the intervals almost rugose and with traces of ground sculpture; as long as broad, widest at the middle, arcuately nar- rowed to apex, emarginately narrowed to base. “lytra one-half wider than pronotum, one-tenth wider than long; humeral callus feeble but with anterior face impunctate; punctures coarser than on pronotum, more distinct and sometimes separated by one-half their diameter by shining intervals; without trace of serial arrangement; pubescence rather long. Abdomen not margined; with moderately large but feeble punctures rather dense; with traces of ground sculpture apically. Zarsus with fourth segment strongly bilobed. Male, seventh sternite very feebly emarginate; eighth with a small triangular emargination twice as wide as deep. Female, seventh sternite very feebly emarginate; eighth sternite truncate. Length, 21/, to 234 mm. Type locality —Cuba, 7 kilometers north of Vinales. Types.—Holotype and 10 paratypes in the American Museum of Natural History, collected on September 16-22, 1913. Four para- types in the United States National Museum (No. 52494) and two in the Museum of Comparative Zoology. Records—The following are the records known to me: Cuba: Vinales (A.M.N.H., U.S.N.M., M.C.Z.), Candelaria (M.C.Z.), Soledad (M.C.Z. and U.S.N.M.). Specimens examined.—I have seen the 17 types and 3 other speci- mens. Remarks. This species is similar to jugalis but differs by its short pronotum, broader emargination of the eighth sternite of the male, and the absence of the spongy area of the seventh sternite. It also differs markedly in size and appearance. I have received no record of its habits. 11. STENUS MORRISONI, new species Description.—Black, elytra and abdomen piceous. Head not bi- impressed between the eyes; not abruptly elevated at edge of eyes; labrum broadly rounded, feebly flattened in front; with coarse and deep punctures generally separated by less than half their diameter, interstices rather flat, without impunctate areas; without ground sculpture; pubescence distinct. Pvonotum one-third longer than wide, widest near middle, sides arcuate in front, almost emarginate MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 219 behind; with coarse and deep punctures generally separated by less than half their diameter by convex intervals; without ground sculp- ture; pubescence very short but distinct. H7/ytra one-half wider than pronotum, one-seventh longer than wide; with humeral callus scarcely elevated and without impressions; punctures less abrupt than on pronotum, appearing a little more separate, intervals less convex; without ground sculpture; pubescence short but distinct. Abdomen not margined; with punctures similar to elytra but smaller, sparser apically, very much finer on eighth tergite which has distinct scaly ground sculpture. Zarsus with fourth segment bilobed. ale, sixth sternite with posterior half at center flattened and with a whitish spongy surface, seventh sternite with same on apical two-thirds, feebly emarginate at center; eighth sternite with a narrow acuminate excision extending through three-fourths of its length, rounded at sides. Length, 5 mm. Type locality—tTrinidad, Department of Agriculture Grounds, Port of Spain. Types.—Holotype, male, U.S.N.M. No. 52386, collected by Harold Morrison on November 23, 1918. Records.—The following is the only record known to me: Trinidad: Port of Spain (Morrison, in U.S.N.M.). Specimens ecamined.—I have seen only the unique type. Remarks.—This species is very similar to jugalis Erichson as I have determined that species. This is distinguished, however, besides the character given in the key, by the much deeper and narrower excision of the eighth sternite in the males. I have no record of its habits. 12. STENUS BRYANTI, new species Description.—Piceous. Head with two longitudinal impressions scarcely distinct; without smooth areas; labrum broadly rounded; punctures moderately coarse, somewhat irregular; without ground sculpture; pubescence evident. Pronotwm one-fourth lo ger than wide, widest about middle, sides in front rounded, nearly straight behind; punctures coarse, deep, dense, separated by less than half their diameter; without ground sculpture; pubescence short. Hlytra three-eighths wider than pronotum, one-eleventh longer than broad; humeral callus scarcely evident; punctures similar to pronotal but intervals a little less convex; no ground sculpture; pubescence a little longer. Abdomen not margined; punctured as the elytra but less coarsely or closely, intervals flat; without ground sculpture except a trace on the eighth tergite; pubescence rather long and dense. Male, seventh sternite with pubescence more dense at middle apically; eighth deeply narrowly triangularly emarginate, apex rather 220 BULLETIN 182, UNITED STATES NATIONAL MUSEUM rounded, sides arcuate apically, emargination extending through half the length of the segment. Memale, apical sternites not modified. Tarsus with fourth segment strongly bilobed. Length, 314 to 4 mm. Type locality.—Trinidad, cocoa estate about 6 miles from La Brea, county of St. Patrick, ward of La Brea. Types.—Holotype, male, and 1 paratype, U.S.N.M. No. 52383, col- lected by Harold Morrison on October 18, 1918; 1 paratype from Aripo Savanna, H. Morrison, October 26, 1918; 10 paratypes in the British Museum labeled “Trinidad B.W.I. 1908. C. E. Bryant.” Records.—The following are the records known to me: Trinidad: (Bryant, in British Museum), La Brea (Morrison, in U.S.N.M.), Aripo (Morrison, in U.S.N.M.). Specimens examined.—I have seen 3 specimens in the United States National Museum and 10 borrowed from the British Museum. Remarks.—This species is very similar to morvrtsoni but differs by its smaller size, less coarse and more irregular punctures of vertex, and more definitely serial pronotal punctures, as well as the sculpture of the eighth tergite. I have no record of its habits. 13. STENUS DYERI, new species Description —Piceous. Head rather distinctly biimpressed on vertex between eyes; elevated at middle and smooth, shining; with additional smooth areas at posterior angle of eye and behind anten- nal ridge, but all areas rather small and indefinite; punctures mod- erately coarse or less, irregularly spaced, intervals flat; vague ground sculpture near antennae; pubescence long and very evident. Pro- notum one-third longer than wide, widest just behind middle; sides arcuate in front, feebly emarginate behind; punctures rather coarse but irregular in size and spacing, intervals nearly flat, not in series along midline; without ground sculpture; pubescence moderate. Elytra one-half wider than pronotum, one-thirteenth longer than wide; humeral callus scarcely evident, with a feeble elevation be- tween it and suture; punctation similar to that of pronotum but a little coarser and sparser; with distinct but rather indefinite ground sculpture; pubescence long and sparse. Abdomen not margined; punctures abrupt and shallow, not dense at base and more sparse apically, diameter less than half that of elytral punctures. fale, un- known. Female, apical segments more densely punctate at middle; eighth sternite slightly emarginate. Zarsus with fourth segment bilobed. Length, 484 mm. Type locality —Cuba. Types.—Holotype, female, U.S.N.M. No. 52382, “F. J. Dyer Coll.,” September 29, 1916. Pecords.—The following is the only record known to me: MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Bok Cuba: (Dyer, in the U.S.N.M.). Specimens examined.—I have seen only the unique type. Remarks.—This species is distinguished from adamsoni by the rela- tive length of the pronotum and by the flatter intervals between the pronotal punctures. It differs from cubanus in its size, in having the abdominal segments not very abruptly constricted, and in having the smooth areas of the head poorly defined. I have no record of its habits. 14. STENUS CUBANUS, new species Description.—Piceous. Head distinctly biimpressed between the eyes; with an elevated central smooth area, another at upper posterior angles of eye, and one behind the antennal ridge; labrum broadly rounded; punctures moderately coarse but confused and crowded in depressions; without ground sculpture; pubescence short but dis- tinct. Pronotwm one-eighth longer than wide, widest near middle, sides feebly arcuate in front, distinctly emarginate behind; with coarse, irregular punctures, with an irregular smooth area at middle and others at sides; intervals moderately convex; without ground sculpture; pubescence short. Hlytra three-fifths wider than pro- notum, scarcely wider than long; humeral callus shghtly prominent, with another feeble prominence between humerus and suture; punc- tures coarser than on pronotum, irregular in size and density; with- out ground sculpture; pubescence short. Abdomen not margined; with punctures very much finer than on elytra, rather sparse; no ground sculpture except on apical segments; pubescence moderately long. TYarsus with fourth segment bilobed. Male, sixth sternite flattened apically and with denser pubescence, seventh same through- out its length; eighth with an even triangular emargination, slightly wider than deep, apex scarcely rounded, outer angles gradually rounded; eighth tergite alone with ground sculpture. /emale, apical sternites not modified; seventh and eighth tergites and sixth, seventh, and eighth sternites with ground sculpture. Length, 3 to 314 mm. Type locality—Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, Santa Clara Province. Types.—Holotype, male and two paratypes, U.S.N.M. No. 52384, collected by E. A. Schwarz on February 28 in 1902, 1908, or 1904. Records—The following is the only record known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This species is distinct from all others in our region by the large impunctate shining areas on the head. The male ab- dominal characters are likewise distinctive. I have no record of its habits. 222 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 15. STENUS ADAMSONI, new species Description—Piceous. Head with two distinct short longitudinal impressions between the eyes; middle elevation with indefinite smooth area, also one at upper apical corner of eye, and one behind antennal ridge; labrum broadly rounded; punctures moderate but deep and irregular, intervals convex, shining; without ground sculpture; pu- bescence distinct. Pronotum about as long as wide, widest near mid- dle; sides feebly arcuate in front, feebly emarginate behind; punctures rather coarse but very irregular, sometimes partly united and some- times leaving wide smooth spaces between; without ground sculpture; pubescence short but distinct. /Vytra three-fifths wider than pro- notum, one-seventh wider than long; humeral callus slightly prom- inent; punctured similarly to pronotum but a little more coarsely, sparsely, and regularly; intervals convex; without ground sculpture; pubescence short but distinct. Abdomen not margined; punctured similarly to elytra but less densely, less coarsely especially posteriorly, and less deeply; without ground sculpture; pubescence short but distinct. Zarsus with fourth segment bilobed. Jfale, unknown. Female, ventral abdominal segments not modified except for denser punctures at middle of apical segments. Length, 4 mm. Type locality—Trinidad, Department of Agriculture Grounds, Port of Spain. Types —Holotype, female, U.S.N.M. No. 52385, collected October 24,1918, by Harold Morrison. Records.—The following is the only record known to me: Trinidad: Port of Spain (Morrison, in U.S.N.M.). Specimens examined.—I have seen only the unique type. Remarks.—This species is not strongly separated from cubanus but it seems best at present to separate it even though the male is unknown. It may prove to be a South American species, but I have been unable to find any to which it can be ascribed. I have no record of the habits of this species. 16. STENUS DARLINGTONI, new species Description—Piceous. Head feebly but broadly biimpressed be- tween the eyes; median elevation with a definite and distinct smooth area extending throughout the vertex but much broader posteriorly ; also indefinite areas along eye and behind antennal ridge; labrum broadly rounded; punctures rather fine but irregular and deep, the intervals very convex and shining; without ground sculpture; pu- bescence short but distinct. Pronotwm one-fifth longer than wide, widest near middle; sides almost straight from middle to front angles, feebly emarginate behind; punctures coarse and very irregular, some- times more or less united in grooves, usually with an irregular smooth MONOGRAPH OF WEST INDIAN STAPHYLINIDAE noe area behind middle of disk; without ground sculpture; pubescence very short. lytra two-thirds wider than pronotum, one-fourteenth wider than Jong; humeral callus rather prominent; punctured simi- larly to pronotum but more coarsely and a little more sparsely and regularly; the intervals unevenly convex; without ground sculpture; pubescence distinct. Abdomen not margined; punctures smaller than on pronotum, a little shallower, intervals flat ; without distinct ground sculpture; pubescence distinct. Zarsus with fourth segment bilobed. Male, seventh and eighth sternites with denser and finer pubescence, eighth with a broad semicircular emargination; ninth flattened trans- versely but somewhat concave longitudinally, narrowed posteriorly with a small arcuate emargination between the two spiniform apices; seventh, eighth, and ninth sternites with very indefinite ground sculp- ture. Female, apical segments without special modifications, sculp- ture asinmale. Length, 44% to 5 mm. Type locality —Trinidad, St. Augustine. Types—Holotype, male, and three paratypes in the Museum of Comparative Zoology; three paratypes in the United States National Museum (No. 52424) ; collected in April 1929 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the seven types. Remarks.—This species is distinguished by the size and length of the median smooth area of the head, as well as by the spiniform apices of the ninth sternite of the male. I have received no record of its habits. 17. STENUS LUCENS Cameron Stenus (Mesostenus) lucens CAMERON, 1913b, p. 329.—LEna and MutcH ter, 1917, p. 198.—ScHEERPELTZ, 1933, p. 1192. (Not Benick, 1917.) Description—Black. Head with eyes broader than elytra; with rather broad, raised, almost impunctate central space, rest of surface with moderately large not very close punctures; central space and interstices smooth and polished without trace of ground sculpture; glabrous; antennae long and slender. Pronotwm widest at middle, scarcely longer than wide, equally narrowed in front and behind, impressed on either side at the widest part; punctures much finer and more sparing than on the head, interstices smooth and _ polished; glabrous. “lytra shorter than pronotum, transverse; with distinct humeral callus and impression internal to latter; disk impressed; punctures fine and scattered as on pronotum, interstices smooth, polished; glabrous. Abdomen margined, gradually narrowed to apex, with a few scattered punctures at base of segments, rest of surface almost impunctate; pubescence scanty. Tarsus with fourth segment 224 BULLETIN 182, UNITED STATES NATIONAL MUSEUM bilobed. Male, eighth tergite with horseshoe-shaped emargination; sixth and seventh sternites impressed at middle, eighth with a narrow deep triangular excision. Length, 34% mm. (From Cameron.) Type locality— Grenada, Balthazar. Types.—One specimen labeled type in the British Museum. Records.—The following is the only record known to me: Grenada: (Cameron, 1913). Specimens examined.—I have seen only the unique type. Remarks.—This species is distinct by its nearly impunctate elytra. I find no record of its habits. 18. STENUS HISPANIOLUS, new species Description—Shining black. Head not at all impressed between the eyes but midline slightly elevated, elevation broadly oval, smooth, not extended anteriorly; with antennal prominence smooth but with- out definite smooth area above eye; labrum very broadly rounded; punctures irregular in size and spacing but often rather dense and moderately coarse; without distinct ground sculpture. Pronotum slightly longer than wide, widest near middle; sides not arcuate in front but angulate halfway between middle and apex, anterior angles scarcely evident, feebly emarginate behind; punctures considerably coarser than on head, irregularly spaced but rather dense especially along midline, occasionally with irregular smooth areas; without ground sculpture. Zlytra one-half wider than pronotum, about as wide as long; humeral callus moderate; with punctures about as coarse as on pronotum but sparser on the disk, separated by much less than their diameter laterally, the intervals rather flat; without ground sculpture. Abdomen not margined; punctures rather shallow and sparse, somewhat indefinite apically; with ground sculpture on two apical tergites. Zarsus with fourth segment bilobed. Jale, sixth, seventh, eighth, and ninth sternites flattened, eighth with a small triangular notch three-fourths as wide as the apex, nearly as deep as wide, with apex narrowly rounded; ninth emarginately truncate with prominent spiniform apices. Female, sternites unmodified. Length, 3 to 4 mm. Type locality—Haiti, tang Lachaux, on the southwest peninsula. Types—Holotype and 12 paratypes, Museum of Comparative Zoology; 9 paratypes in the United States National Museum (No. 52496). Records.—The following are the records known to me: Cuba: Soledad (Darlington, in M.C.Z. and U.S.N.M.), Buenos Aires, Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.), Hongolosongo, Oriente (Darlington, in M.C.Z.). Hispaniola: Harri, Stang Lachaux (Darlington, in M.C.Z. and U.S.N.M.) ; Domi1n- ICAN REPUBLIC, Sanchez (Darlington, in M.C.Z.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 225 Specimens examined.—I have seen only the 22 types. Remarks.—This species is not entirely satisfactory in the key. The smooth areas of the head are often distinct but are not as regular as in the species in the first half of couplet 9. It can be distinguished from adamsoni and cubensis by the size of the median area of the vertex, and from most other species by the unimpressed head. I have received no record of its habits. 19. STENUS CUBENSIS Bernhauer Stenus cubensis BERNHAUER, 1910, p. 364. Stenus (Hypostenus) cubensis Bernhauer, BerNHAUER and ScHusert, 1911, eli. H re cubensis (Bernhauer) Lena and Murcuter, 1914, p. 404. Description—Black with slight aeneous luster. Head not im- pressed between the eyes but feebly elevated along midline, the elevation narrowly and irregularly smooth and shining; without other smooth areas; labrum broadly rounded; punctures coarse but irregu- lar and obscured by short hairs; without distinct ground sculpture. Pronotum scarcely at all longer than wide, widest near middle; sides nearly straight in front and only very feebly emarginate behind; punctures a little coarser than on the head but irregular, sometimes with suggestion of sculpture on the intervals. /lytra one-half wider than the pronotum, somewhat wider than long; humeral callus moderate; punctures a little coarser and sparser than on pronotum, of irregular shape; without sculpture on the intervals. Abdomen with punctures similar to those on head, sparser apically; with distinct ground sculpture on the two apical tergites. Male, eighth sternite with a triangular emargination one-half deeper than wide; seventh sternite scarcely emarginate posteriorly, flattened and with somewhat dense and long pubescence. Length, 214 mm. Type locality—Cuba, Habana. Types.—One male, presumably in the collection of Max Bernhauer. fecords.—The following are the records known to me: Cuba: (Leng and Mutchler, 1914), Habana (Bernhauer, 1910), Soledad (Darling- ton, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen three examples of this species from the Museum of Comparative Zoology (of which one has been deposited in the United States National Museum). Remarks.—This species was originally placed in the subgenus Hypostenus. It is not strikingly different from adamsoné and others. It appears to be distinguishable by the feeble development of the smooth areas of the head and the distribution of the punctation. Ten other specimens which agree closely with this species (from Soledad, Aguadores in Oriente, and Buenos Aires in the Trinidad Mountains) differ principally in the size and shape of the emargina- 226 BULLETIN 182, UNITED STATES NATIONAL MUSEUM tion of the eighth sternite in the male. In these it is smaller, may be as wide or twice as wide as deep, and there is less obvious modifi- cation of the punctation of the seventh and eighth sternites. I find no record of its habits. 20. STENUS ANTILLENSIS Benick Stenus (Hypostenus) antillensis BeNIcK, 1917, p. 301, 302.—ScHEERPELTZ, 1933, Dp. Lvs: Description—Black. Head moderately wide, somewhat wider than the pronotum and somewhat smaller than the elytra, vertex feebly impressed; with median broadly elevated area, another behind the antennal ridge, and another at posterior margin of eye, smooth and shining. Pronotum scarcely longer than wide; sides expanded, strongly rounded at middle, arcuate before and behind; coarse punc- tures separated at middle to form midline; behind on each side with a distinct shining tubercle. Hlytra as long as pronotum, scarcely widened behind humerus; together somewhat deeply emarginate behind; the convex disk is papresed 4 in front at suture, and on each side with two wide transverse impressions which are feeble and do not reach entirely to the suture; coarsely punctate, more coarsely and a little more sparsely than pronotum. Abdomen not margined; in front coarsely and densely punctate, behind much finer and some- what more sparsely punctate. Zarsi with fourth segment bilobed, first as long as two following together. Afale, seventh sternite finely punctate and pubescent at middle, eighth with a broad shallow rounded emargination, surface at middle finely and densely punctate and densely pubescent. Jnteguments without ground sculpture. Length, 444 mm. (From Benick.) Type locality—Guadeloupe. Types.—One male in collection of Ludwig Benick. Records.—The following is the only record known to me: Guadeloupe: (Benick, 1917). Specimens examined.—I have seen no examples of this species. Remarks.—This species is apparently distinct by its tuberculate pronotum as well as by the male abdominal characters. I find no record of its habits. Subfamily EHUAESTHETINAE This subfamily has been very imperfectly characterized, and most of its genera are known chiefly from the included species, there being no adequate descriptions or diagnoses. I know of no character or group of characters that will distinguish the subfamily from the ie Pselaphidae. The flexibility of the abdomen is a relative condition at best, but some of these and many MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 224 other staphylinids have the abdomen almost completely rigid. Very few staphylinids have eight visible ventral abdominal segments, the Euaesthetinae not being among those few. The number of tarsal subsegments varies so greatly in the Staphy- linidae that the presence of more segments in the Euaesthetinae than is characteristic of the Pselaphidae can scarcely be taken as a charac- ter of importance. There are quite enough staphylinids with 3- segmented tarsi to make this character useless in separating the Staphylinidae and Pselaphidae. Three species, placed in three genera, have been recorded from the West Indies. I have seen examples of two of these but am un- able to verify either the generic or specific assignment. In the meager literature on this group I have been unable to find characters for a generic key or for generic diagnoses. I have therefore merely listed the published records and synonymy. XXVIII. Genus EXOCTAVIUS Bierig Exoctavius Breric, 1984e, p. 221. Genotype. —E.. bermudezi Bierig (monobasic and original designa- tion). Remarks.—No information is available to me other than that in the original description referred to above. Only one species is known. 1, EXOCTAVIUS BERMUDEZI Bierig Ezxoctavius bermudezi Breria, 1934e, p. 222, pl. 11, fig. 6. Type locality—Cuba, Sierra del Rosario, vicinity of the source of the Rio Taco-Taco. Types—Holotype and allotype in the collection of Alexander Bierig. (An example sent to the United States National Museum labeled cotype was collected at Aspiro. It may not have been in the original series, since that locality was not mentioned in the origi- nal description. ) Records.—The following are the records known to me: Cuba: Sierra del Rosario (Bierig, 1934), Aspiro (Bierig, in U.S.N.M.). Remarks.—The two types (and one other example) are the only ones known, but the description may be adequate when the other American Euaesthetinae are sufficiently known to permit comparisons. XXIX. Genus TAMOTUS Schaufuss Tamotus ScHAUFUSS, 1872, p. 248 (nomen nudum) ; 1877, p. 289. Genotype.—T. femoratus Schaufuss (monobasic). Remarks.—i can find no diagnosis of this genus or even a dis- cussion of a few characters. Careful study will be required to de- termine its characters and their limits. Only one species is known. 228 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 1. TAMOTUS FEMORATUS Schaufuss Tamotus femoratus SCHAUFUSS, 1874, p. 289; 1883, p. 166, 170.—FAuveL, 1902a, p. 88.—EIcHELBAUM, 1909, p. 132.—BERNHAUER and ScHUsBERT, 1911, p. 188.— Lene and MutcuHterr, 1914, p. 404. Type locality—Cuba. Types.—Possibly in the Zoologische Museum, Berlin. Records.—The following is the only record known to me: Cuba: (Schaufuss, 1874; Leng and Mutchler, 1914). Remarks—This species has apparently not been reported since the original description. The information available is insufficient to place it in its proper relation to the rest of the subfamily. XXX. Genus STENAESTHETUS Sharp Stienaesthetus SHARP, 1874, p. 79. Genotype.—S. sunioides Sharp (monobasic). Remarks.—I can find no adequate diagnosis of this genus. It has apparently not been mentioned since its original proposal. Four species have been assigned to it. 1. STENAESTHETUS IMMARGINATUS (Erichson) Huaesthetus immarginatus Ericuson, 1840, p. 748. Type locality —‘Valle Araguensi Columbiae.” Types.—In either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: St. Vincent: (British Museum). South America: CotomprA (Hrichson, 1840). Remarks.—One example from St. Vincent is in the British Museum. It is the sole authority for the transfer of this species from Huaesthe- tus to Stenaesthetus, as well as for the West Indian record. Subfamily PAEDERINAE This large and important subfamily presents more difficulties than most of the others. A revisionary treatment of the genera was attempted by me in 1939 as part of this study, but some changes have been made in the prsent manuscript. Several groups of genera are in need of furthur revision. Among these may be mentioned the Pinophilini, the A/yrmecosaurus-group, the Medon-Lithocharis com- plex, and the Stilicopsis-group. With many of the smaller species segregation to genus can be made only after examination of the prosternal process under the front coxae. This usually necessitates the relaxing and lifting of one front coxa. In the West Indian fauna the genera can later MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 229 be recognized by other characters, such as the sculpture of the head and the armature of the labrum. In most classifications of the Staphylinidae the Paederinae and Staphylininae are separated by the character of the exposure or concealment of the mesothoracic spiracles (sometimes called pro- thoracic) or by the shape of the posterior coxae. In the case of the first of these characters the Paederinae contains every degree from completely free and visible spiracle to one completely hidden by one of several means. The character is simply not usable for this separation, but it is not easy to find a substitute. The shape of the posterior coxae is not a usable character in this instance. It is im- possible to key out certain lathrobiids and philonthids without refer- ence at this point to other characters. I have attempted to show previously (Blackwelder, 1936) that these two subfamilies have posterior coxae that are structurally similar. Studies of the larvae (Béving, 1931) confirm the opinion that these subfamilies are closely related. Careful study has so far shown only one fundamental structural character that distinguishes the adults—the pattern of the pigmented areas on the intersegmental membranes of the abdomen. Illustrations of these are given in the paper mentioned above (Blackwelder, 1936). They can sometimes be seen with a binocular microscope of fairly high power, or may be examined readily in slide preparations under the compound micro- scope. In the Paederinae the sclerotized or pigmented areas are rec- tangular and are arranged in transverse rows. In the Staphylininae they are triangular or irregular and are arranged in definite longi- tudinal bands. KEY TO WEST INDIAN GENERA OF PAEDERINAE 4 1. Maxillary palpus with the last segment very small, generally subulate__ 4 Maxillary palpus with the last segment as long as the preceding, obliquely INAECHE ESI AC Casa sa eee a eR SD 2 2, Abdomen- strongly-margined, -punctates.=%2 00) Sts ers DOO re oe S Abdomen not margined, coarsely and regularly sculptured. LXI, Palaminus 3. Mandibles edentate or with a single internal tooth near the base. LX. Araeocerus Inner edge of mandibles near the middle with a parallel slender tooth, which is oblique, and emarginate at apex___________________ LIX. Pinophilus 4, Prosternum not dilated under the front coxae as far as hypomera________ 5 Prosternum expanded laterally and caudally, either connate with hypomera or very larrowly separated’ from thenye: 22 Voults ies Fae ee 22, 5. Antennae anteriorly flexile and strongly geniculate at first joint, basal seg- THENUPVET Yeh CLOT AA Cress Se See aE ES REGS Ue See = Sad. SER OLE” 6 Antennae posteriorly flexile, not strongly geniculate, basal segment not very CLON ZA Ce= weer eter unas stra tem SeeRey Rae ner NN Phe eas Ao zie et ALLEY EU ALT) ak 9 4 This key, with the exception of the first three couplets, has been adapted from the key in my recent revision of the Paederini (Blackwelder, 1939). 230 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 10. at. Neck less than one-fourth as wide as head__-_____________ XLVIIL. Ophites Neck more than one-fourth as wide asheadl_22.0 42-0 eet a 7 . Elytra with a pleural fold near side margin___________ XLV. Homoeotarsus Hlytra,withont trace. ofa, pleural folds +... 2. 3. et 8 uabbume mot Centaie. 22 5h. oie ee XLVII. Biocrypta Labrum bidentate (the denticles sometimes obtusely rounded). XLVI. Ochthephilum . Fourth segment of maxillary palpus net strongly compressed nor very short, flabrouse inde lsc ns seers. et we Secepl eee Sal eli ater = eyes eyed sp Legeib oe 10 Fourth segment of maxillary palpus compressed, truncate, and pubescent. XLIV. Paederus Neck never less than one-fourth as wide as head; apex of posterior tibia With. a.‘distinct ctenidium: on both sides: 4 Se eee 11 Neck variable; apex of posterior tibia with a distinct ctenidium only on inner .edge fsa 3-y asl bo oretaed ol melts wk asipee 25 Adee als} Fourth segment of maxillary palpus longer than greatest width of third; labrum semicircularly emarginate; punctation dense and umbilicate. . XLI. Scopobium Fourth segment of maxillary palpus shorter than greatest width of third; labrum bilobed or triangularly emarginate; punectation generally not um- bilichteente dt. Alt. watey soda Loar teiprte TP ovreupeg beeriel ater os 12 12. Elytra with a pleural fold near side margin___________ XLITI. Lobrathium Elytra without trace of a pleural fold____-_____________ XLII. Lathrobium 13. Neck (usually) one-fourth as wide as head, or more_____-______________ 14 Neck one-fifth to one-eighth as wide as head_________-_________-____ 20 14. Antennae with first two segments larger, rest verticillate, slender, of equal thicknessithroughoutes sect aetees te epee obs 5S ee XXXI. Thinocharis Antennae normal. oon et eA es a Ee 15 16. iT. 18. 19. 21. . Pronotum distinctly longer than wide; seventh abdominal sternite in male generally distinctly modified; punctures of head not distinctly umbilicate though not "very nels 2s sees ERO a ee ee ee ee eee XXXVII. Orus Pronotum not or scarcely longer than wide; seventh abdominal sternite of male rarely modified; punctures of head umbilicate or fine and dense. 16 Head and pronotum densely and very finely punctate or sculptured, without any umbilicate punctures except at margins_______-_ XXXII. Lithocharis Head with few or many distinct umbilicate punctures, surface sometimes densely punctulate...—.... = =} ee ee MG Labrum without median tooth_._<+s+--2. —8.3s-.—- = fe een eee 19 Labrum with median tooth or prominence___----_.-~-_-------------_+-+ 18 Umbilicate punctures of head sparse and not strong--_ XXXIITI. Aderocharis Head densely and strongly umbilicately punctured___ XXXIV. Stilomedon Gular sutures distinctly diverging posteriorly from before middle. XXXY. Sunius Gular sutures most approximate along middle or at base. XXXVI. Medon . Gular sutures always united, at least basally________-_-__-_-_ XL. Rugilus Gular sutures never united in any part (although often very closely approxi- IMAteG), 2% oe ee re ee eS Ss 21 Labrum not dentate, feebly emarginate; vertex sometimes carinate in males. XXXIX. Monista Labrum not denticulate, truncate and narrowly emarginate. XXXVIII. Scopaeus 13 further examination shows that this character is rather misleading. The presence of threo or four spinules in a row well below the apex is not sufficient to place the specimen in this half of the couplet; the row must be definite and extend almost to the apex. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 231 22. Prosternum connate with edges of hypomera___-----------_~----------~ 23 Prosternum not connate with edges of hypomera_____-_--------------- 26 23. Body covered with dense coarse verrucose sculpture__-_-_~~-~ LY. Ronetus Body not covered with dense coarse verrucose sculpture__------------- 24 24. Prothorax very elongate; head grooved behind eyes____ LVIII. Sphaeronum Prothorax not very elongate; head not grooved behind eyes_--------~-- 25 oF iiabrmm QUAGrIGenta elses fee Oi Aaa See Ene LVII. Echiaster abrumebigentatesst7-23 hs + seo a. te oln Sete Ph ee LVI. Astenus %GLaprumudenticulate.or: broadly. lobed in, front -.=__.-_.__--_--—~--___ 27 Labrum not denticulate, truncate and narrowly emarginate. XLIX. Suniophacis 27. Prothorax narrowly prolonged at middle in front-_-_--_- LI. Stamnoderus Prothorax, not distinctly prolonged in-front.=-* —--- —_==22--!) a 28 28. Head coarsely umbilicately punctate throughout______~-~_- LII. Stiliphacis Head not distinctly umbilicately punctate throughout__-___-___-__---_-- 29 29; Headsand: pronotum’ densely, sculptured=_222--2==—--- _*- SS eas 30 Head and pronotum not densely sculptured____________---- L. Stilosaurus 380. Labrum with two long and very slender hornlike denticles. LIII. Suniosaurus Labrum with denticles not slender and hornlike_____--_--_ LIV. Suniocharis XXXI. Genus THINOCHARIS Kraatz Thinocharis KRaatz, 1859b, p. 142. Subgenus Sciocharis Lyncu, 1884, p. 260. Subgenus Sciocharella Casry, 1905, pp. 151, 158. Genotypes—Thinocharis pygmaca Kraatz (designated by Black- welder, 1939). Of Sctocharis, S. castanoptera Lynch (designated by Blackwelder, 1939); of Sciocharella, S. delicatula Casey (mono- basic). Diagnosis —Body slender, moderately convex; antennae posteriorly flexile, not strongly geniculate, basal segment not very elongate, first and second segments larger than the remainder, which are of uniform slenderness, somewhat verticillate; labrum without median tooth, with or without paired denticles; fourth segment of maxillary palpus very small, subulate; gular sutures not at all united, variable; pro- sternum carinate posteriorly, short, not dilated laterally under the coxae ; hypomera prolonged in a short lobe partially behind the coxae; anterior coxal cavities open behind, confluent; front coxae large, ex- serted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, third carinate at middle; seventh sternite generally modified in the male; tarsi 5- segmented, anterior tarsus either slender or dilated. Remarks.—This genus will undoubtedly be a large one with several more subgenera. Its developments somewhat parallel those found in the genera related to Lithocharis and Medon, but I am at present unable to believe that the peculiar form of the antennae is not a char- acter of generic importance. 232 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Medonella Casey is not at all similar to Thinocharis, but some of the South American species of 7’hinocharis will probably require new subgenera. All our species belong to the subgenera Sctocharis and Sciocharella, which differ from Thinocharis (s. str.) in having the labrum bidentate. Sczocharis has the gular sutures most approxi- mate along the middle or posteriorly, whereas Sciocharella has them most approximate anteriorly. I have examined 182 specimens of this genus from the West Indies, including the types of all but one of the valid species. Of these 130 are in the British Museum, 2 are in the Casey collection and 2 in the general collection of the United States National Museum, and 40 were collected by me in 1935-387. These belong to six species, of which two appear to be new. KEY TO WEST INDIAN SPECIES OF THINOCHARIS 1. Gular sutures most approximate anteriorly “__-___-_-___-_________-__-____ 2 Gular sutures most approximate posteriorly or along middle “_-___-______ 3 2. Punctures of head moderate:.and distinct=—=—*—-- *2222"" SS 5. smithi Punctures of head: minutezand wery dense_-=. "+ _ +s es- _-__ =. 6. exilis 8. Labrum with a median denticle and two laterals; seventh sternite of male not smodified—= =.=" -_ == sees a ee ee eee 1. poundi Labrum without median denticle; seventh sternite of male modified___--~- + 4. Labrum with very short minute denticles; emargination of seventh sternite in male not bounded laterally in any way_--_~_----~---.+---_--- 2. bakeri Labrum with distinctly triangular and prominent denticles; emargination of seventh sternite in male bounded laterally by denticles________-____--___ 5 5. Elytra with discrete tuberculi; emargination of seventh sternite in male bounded laterally by a minute denticle-___-_______________-___-- 3. chapini Elytra without discrete tuberculi; emargination of seventh sternite in male bounded slaterallys bya strong, tooth=—2—2 ee eee 4, fuscina 1. THINOCHARIS POUNDI, new species Description —Rufopiceous, elytra testaceous with large sutural in- fuscation, abdomen rufotestaceous. /7Zead nearly one-fourth wider than long; eyes large, at much less than their diameter from base; sides behind eyes nearly parallel, base feebly arcuate but angles mod- erately prominent; base slightly emarginate at middle; neck nearly one-half as wide as head; gular sutures moderately separated along middle, feebly diverging at both ends; without distinct punctures but with dense and rather uneven sculpture; with very fine pubescence and the usual longer hairs; labrum with two small acute denticles and a smaller one between them. Pronotum one-sixth wider than long, seven-eighths as wide as head; widest at anterior angles, moderately narrowed posteriorly, angles all rounded; with a rather indefinite 4The West Indian species can be separated also on the sculpture of the pronotum. In Bciocharella (2) the surface is finely or coarsely punctate; in Sciocharis (3) it is tubercu- late but not at all punctate. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 233 smooth midline which is grooved posteriorly; impunctate but with moderately dense tubercles throughout; without ground sculpture; pubescence fine. Llytra as wide as long, one-seventh wider than pro- notum; anteriorly with regular and even punctation or sculpture, but with ground sculpture along suture basally, tuberculi apically, and some indefinite punctures laterally. Abdomen very finely densely and indistinctly punctate; with dense fine pubescence. Jfale, seventh sternite unmodified; eighth sternite with a triangular emargination four times as wide as deep. Female, sternites unmodified. Length, 216 to 234 mm. Type locality—tTrinidad, 5 miles west-southwest of Manzanilla junction on the Plum Road. Types.——Holotype, male, and two paratypes, females, U.S.N.M. No. 52394, collected by me on December 18, 1935. Records.—The following is the only record known to me: Trinidad: Manzanilla (Blackwelder station 104A). Specimens examined.—I have seen only the three types. Remarks.—This species is distinguished by the simple male charac- ters and the structure of the labrum. It belongs in the subgenus Sciocharis as herein defined. The types were collected from rotting cocoa pods. 2. THINOCHARIS BAKERI (Casey) Sciocharis bakeri Casry, 1910, p. 188.—LeNe and MurtcuHtrer, 1914, p. 405; 1947, p. 199. BLACKWELDER, 1939, p. 102. Thinocharis bakeri (Casey) BERNHAUER and SCHUBERT, 1912, p. 228.—BLAcK- WELDER, 1939, p. 102. Description—Head rufopiceous, pronotum rufous, elytra castane- ous, abdomen rufotestaceous. Head scarcely wider than long; eyes rather large, at about their length from base; sides behind eyes nearly straight and parallel, base nearly straight at sides, slightly emarginate at middle, posterior angles rather prominent; neck one- half as wide as head; gular sutures moderately separated along middle, diverging feebly at both ends; with sparse setigerous punc- tures obscured by dense ground sculpture; labrum not distinctly emarginate but with two small but prominent median denticles. Pronotum one-seventh wider than long, scarcely narrower than head; widest at anterior angles, feebly narrowed posteriorly, angles all rounded ; with dense but discreet tuberculi throughout, generally with a minute puncture visible on each tubercle; no other punctures or definite sculpture; with very fine and inconspicuous pubescence; with distinct midline, feebly grooved at base. Llytra as long as wide, nearly one-fourth wider than pronotum; not distinctly punctate, but with dense tuberculate sculpture; pubescence fine and dense. Ad- 449008—42-16 934 BULLETIN 182, UNITED STATES NATIONAL MUSEUM domen not enlarged; very finely, densely, and indistinctly punctured and pubescent. Male, seventh sternite with two rounded emargina- tions separated by a blunt tooth, the emarginations not separated laterally from the margin of the segment, with unusually large and abundant hairs posteriorly; eighth sternite with a deep and broadly rounded emargination bordered with larger hairs. /emale unknown. Length, 24% mm. Type locality —Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types—Unique holotype in Casey collection in United States National Museum (No. 38216). Records.—The following is the only record known to me: Cuba: (Leng and Mutchler, 1914), Cayamas (Casey, 1910). Specimens examined.—I have seen only the unique type. Remarks.—This species is similar to fuscina except for the very distinct male characters. It has not been reported since the original description and is not represented in the abundant material collected at Cayamas by E. A. Schwarz. It belongs in the subgenus Sciocharis as herein defined. I find no record of its habits. 3. THINOCHARIS CHAPINI, new species Description —Head dark rufopiceous, pronotum rufous, elytra and abdomen testaceous, elytra broadly infuscate along suture. Head scarcely wider than long; eyes large, at a little less than their length from base; sides behind eyes nearly parallel, base feebly arcuate, angles moderately prominent; base slightly emarginate at middle; neck not quite one-half as wide as head; gular sutures moderately separated along middle, feebly diverging at both ends; with moderate punctures especially at sides obscured by dense ground sculpture; labrum not distinctly emarginate but with two rather large and blunt denticles at middle. Pronotwm one-sixth wider than long, scarcely narrower than head; widest at anterior angles, moderately narrowed posteriorly, angles rounded; with rather indefinite smooth midline, which is feebly canaliculate posteriorly; without punctures but with moderately dense tuberculi throughout, each with a minute puncture on its top; without ground sculpture; pubescence exceed- ingly short and fine. ZJytra as long as wide, one-fourth wider than pronotum; with some indistinct ground sculpture basally along su- ture but with tuberculi as on pronotum but sometimes not as distinct. Abdomen very finely, densely, and indistinctly punctulate; with dense fine pubescence. ale, seventh sternite with two rounded emargina- tions separated by a blunt triangular tooth and feebly separated from MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 235 edge of segment by a minute denticle; eighth sternite with a deep tri- angular emargination. /emale, sternites not modified, 244 to 3 mm. Type locality Jamaica, between Luana and Kensworth by way of Lacovia, Santa Cruz, and Spur Tree, parishes of St. Elizabeth and Manchester. Types.—Holotype, male, and two paratypes, female, U.S.N.M. No. 52393, collected by Chapin and Blackwelder in February 1937. The paratypes were taken near Milk River. Records—The following are the records known to me: Jamaica: Santa Cruz (Blackwelder station 421), Milk River (Blackwelder sta- tion 415). Hispaniola: DominicAN Repusiic, Villa Altagracia (Darlington, in M.C.Z.), Jarabacoa (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the three types and three examples collected by Darlington. Remarks.—This species is rather similar to bakeri in the male characters but differs in details of these and in the sculpture of the elytra. It belongs to the subgenus Sctocharis as herein defined. The types were caught flying at dusk. 4. THINOCHARIS FUSCINA Cameron Thinocharis fuscina CAMERON, 1913b, p. 342—LeNeG and MurcHtrr, 1917, p. 199.—ScHEERPELTZ, 1933, p. 1246.—BLACKWELDER, 1939a, p. 102. Description.—Head dark rufous, pronotum rufous, elytra and ab- domen rufotestaceous, the elytra sometimes infuscate on the disk. Head one-tenth wider than long; eyes moderately large, separated by a little less than their length from base; sides behind eyes straight and nearly parallel, base feebly arcuate but angles rather prominent; somewhat emarginate at middle; neck nearly one-half as wide as head; gular sutures moderately approximate along middle, diverging feebly at both ends; with numerous moderate punctures obscured by the dense ground sculpture; with dense short pubescence and longer hairs at sides and back; labrum not distinctly emarginate but with two rather large triangular denticles at middle. Pronotwm one- seventh wider than long, scarcely as wide as head; widest at anterior angles, feebly narrowed posteriorly, angles all rounded but the pos- terior more broadly; with rather dense but discreet tuberculi through- out, sometimes appearing to have a minute puncture on top of each tubercle; without other punctures or sculpture; pubescence short and dense; midline rather vaguely smooth and sometimes feebly canal- iculate posteriorly. “lytra scarcely longer than wide; about one- sixth wider than pronotum; not distinctly punctate, with tuberculate sculpture not very dense or fine but without distinct tuberculi; pubescence fine and dense. Abdomen very finely densely and indis- 236 BULLETIN 182, UNITED STATES NATIONAL MUSEUM tinctly punctate; with dense fine pubescence. Male, seventh sternite strongly produced at middle, with two semicircular emarginations separated by a broad triangular tooth and bounded laterally by acute processes directed posteromedially; eighth sternite with a deep broadly rounded emargination. Female, with sternites unmodified. Length, 214 to 2384 mm. Type locality —Grenada, Chantilly Estate. Types—In the British Museum. (One specimen in the United States National Museum, No. 50876, is designated cotype—or paratype). Records.—The following are the records known to me: Grenada: (Cameron, 1913; British Museum; U.S.N.M.). Trinidad: Tunapuna (Weber, in M.C.Z.). Specimens examined.—I have seen 115 specimens in the British Museum, 1 in the Museum of Comparative Zoology, and 7 in the United States National Museum. The last were part of the original series, received in exchange. Remarks.—I am unable to separate the females of this species from those of chapini, but the male characters are very distinct from those of any other of our species. It belongs in the subgenus Sciocharis as herein defined. I find no record of its habits. 5. THINOCHARIS SMITHI Cameron Thinocharis smithi CAMERON, 1913b, p. 343.—LENe and MurcHter, 1917, p. 199.— SCHEERPELTz, 1933, p. 1247.— BLACK WELDER, 1939a, p. 102. Description —Piceous. Head nearly one-half wider than long; eyes very large, occupying almost the entire side of the head; emar- ginately truncate behind; neck set into the emargination; neck about one-third as wide as head; gular sutures moderately separated, feebly converging anteriorly; punctures moderate and rather irregular, gen- erally separated by less than their diameter; without ground sculp- ture; with numerous long hairs, especially at sides and back; labrum with two small but acute and prominent denticles near middle, not emarginate. Pronotwm one-sixth wider than long, seven-eighths as wide as head; widest at anterior angles, feebly narrowed posteriorly with hind angles and base rounded; with large punctures, generally distinctly umbilicate, separated by less than half their diameter; with a trace of smooth area along midline behind middle. /lytra slightly wider than long, not quite as wide as pronotum; with moderately Jarge distinct punctures, generally separated by once to twice their diameter; without ground sculpture. Abdomen somewhat wider than elyta in part; very indistinctly punctate but with numerous MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 237 long hairs. fale, seventh sternite very slightly and broadly emargi- nate; eighth sternite with a triangular emargination twice as wide as deep with the apical angles rounded. Female, sternites not emargi- nate. Length, 2 to 2144 mm. Type locality—Grenada, Belleview Estate. Types.—in the British Museum. Records.—The following are the records known to me: St. Vincent: (British Museum). Grenada: (Cameron, 1918). Specimens examined.—I have seen 12 examples (including the type) in the British Museum and 7 in the United States National Museum. The latter were from the original series and were received in exchange. Remarks.—This species is distinguished from all others in our region by its large distinct punctures and lack of ground sculpture. It belongs in the subgenus Sciocharella as herein defined. I find no record of its habits. 6. THINOCHARIS EXILIS (Erichson) Lithocharis exilis BEricuson, 1840, p. 627.—Fauvet, 1888, p. 25.—HICHELBAUM, 1909, p. 146.—BLACKWELDER, 1939a, p. 102. Sciocharis atratula Lyncu, 1884, p. 265.—FAUVEL, 1888, p. 25.—H1cHELBAUM, 1909, p. 146. Sciocharis fragilis SHarp, 1886, p. 574.—Favuver, 1888, p. 25.—BLACKWELDER, 1939a, p. 102. Sciocharis minuta SHARP, 1886, p. 574. Sciocharella delicatula CASry, 1905, p. 159.—Dury, 1906, p. 258.—HICHELBAUM, 1909, p. 146.—CaskEy, 1910, p. 188.—BLaATCHLEY, 1910, p. 431. Sciocharella pertenuis Casey, 1910, p. 188.—Lrene and MurcuHter, 1914, p. 405; 1917, p. 199. Thinocharis evilis (Erichson) BeERNHAUER and ScHUBERT, 1914, p. 229.—BERN- HAUER, 1921, p. 36.—SCHEERPELTZ, 1933, p. 1246.—BLACKWELDER, 1939a, p. 102. Thinocharis atratula (Lynch) BERNHAUER and SCHUBERT, 1914, p. 229 (as Synonym of evilis). Thinocharis pertenuis (Casey) BERNHAUER and ScHUBERT, 1914, p. 229.—LENe and MutcHtirr, 1914, p. 405.—BLACKWELDER, 1939a, p. 102. Thinocharis fragilis (Sharp) BERNHAUER and SCHUBERT, 1914, p. 229.—BLACcK- WELDER, 1939a, p. 102. Thinocharis minuta (Sharp) BERNHAUER and SCHUBERT, 1914, p. 229. (Not Casey, 1905. ) Thinocharis delicatula (Casey) BrERNHAUER and ScHuBERT, 1914, p. 229.—BLAcK- WELDER, 1939, p. 102. Thinocharis (Sciocharis) exilis (Hrichson) ScHEERPELTZ, 1983, p. 1246. Description.—Castaneous to ferrugineous, elytra testaceous with sight infuscation along suture, abdomen rufotestaceous. Head scarcely wider than long; eyes rather small, at more than their length from base; posterior angles rounded, base not emarginate; neck one- 238 BULLETIN 182, UNITED STATES NATIONAL MUSEUM half as wide as head; gular sutures rather widely separated, moderately diverging behind; not distinctly punctate, but densely and very finely sculptured throughout; with very fine and inconspicuous pubescence; labrum not emarginate but with two fine denticles above margin at middle. Pronotum one-sixth wider than long, as wide as head; sides nearly parallel, posterior angles more broadly rounded; with dense ground sculpture as on head; no trace on midline. “lytra one-third wider than pronotum, a trifle wider than long; very finely and rather densely punctate and vaguely tuberculose, but not densely sculptured as head and pronotum; pubescence fine and dense. Abdomen rather fusiform, not wider than elytra; very finely and indistinctly punctate and pubescent. J/ale, seventh sternite feebly divided into three lobes by shallow angular emarginations, before which are clusters of hairs; eighth sternite with a shallow emargination, rounded at apex and with apical angles rounded. Female, seventh and eighth sternites not emarginate. Length, 134 to 2 mm. Type locality—Colombia. Of atratula, province of Buenos Aires, Argentina; of pertenuis, Cayamas, Cuba; of fragilis, Guatemala; of minuta, San Miguel, Pearl Islands, Panama; of delicatula, Selma, Alabama. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of atratula, in the Museo Argentino Ciencias Naturales, Bernard Rivadavia, Buenos Aires; of pertenuis and deli- catula, in the Casey collection in the United States National Museum ; of fragilis and minuta, in the British Museum. Records.—TYhe following are the records known to me: Cuba: (Leng and Mutchler, 1914, as pertenuis), Cayamas (Casey, 1910, as per- tenuis; Schwarz, in U.S.N.M.). Antigua: (Blackwelder station 275B). Dominica: (Blackwelder stations 237, 252). St. Lucia: (Blackwelder stations 218, 220D, 226). Trinidad: (British Museum), St. Augustine (Weber, in M.C.Z.), Tunapuna (Weber, in M.C.Z.). South America: Braziwt (British Museum), ARGENTINA (Lynch, 1884, as atratula), CoLoMBIA (Erichson, 1840). Central America: PANAMA (Sharp, 1886, as minuta), GUATEMALA (Sharp, 1886, as fragilis). North America: ALABAMA (Casey, 1905, as delicatula), FLrorips (Casey collection in U.S.N.M.), InprIAna (Blatchley, 1910). Specimens examined.—I have examined 8 specimens in the British Museum, 2 in the Museum of Comparative Zoology, 5 in the Casey collection and 2 others in the United States National Museum, and 18 collected by me in 1935-87. As the above synonymy was not known to me when I visited the British Museum, I did not examine the types of Sharp’s two species from Central America. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 239 Remarks.—This is an unusually small and obscure species, which shows some variation in certain characters. Its lengthy synonymy is due more to the confusion in generic names than to anything else, although most of the synonyms have been recorded only once each. It is distinguished from our other species by the modification of the seventh sternite in the male and by the dense sculpture. It belongs in the subgenus Sctocharella as herein defined. The name atratula is listed as a synonym on the authority of Fau- vel. The two Casey species are placed as synonyms after a careful examination of the types. Sharp’s two species are included on the basis of the original descriptions and a note by Fauvel. My specimens were found in dung, in decaying cocoa pods, in a pile of dead grass, and flying at dusk. XXXII. Genus LITHOCHARIS Beisduval and Lacordaire Lithocharis BorspuvaAt and LACoRDAIRE, 1885, pp. 420, 481. Lithocharius Kraatz, 1857a, p. 664 (misspelling). Litocharis HAGENS, 1865, p. 110 (misspelling). Lithocaris Houper, 1882, pp. 78, 97 (misspelling). Metaxryodonta Castry, 1886a, p. 29. Lithocarus KNaAus, 1904, p. 154 (misspelling). Arthocharis CAMERON, 1921, p. 372. Subgenus Pseudomedon MuULsAnt and Rey, 1878, p. 122. Ramona Casrty, 1886b, p. 213. Subgenus Ophiomedon SHARP, 1886, p. 567, Subgenus Stilocharis SHARP, 1886, p. 576. Genotypes.—Paederus ochraceus Gravenhorst=Lithocharis ochra- cea (Gravenhorst) (designated by Thomson, 1861). Of Arthocharis, Paederus ochraceus Gravenhorst=Arthocharis ochracea (Graven- horst) (designated by Blackwelder, 1939) ; of Metaxyodonta, M. tes- tacea Casey (monobasic) ; of Pseudomedon, Lathrobium obsoletwm Nordmann=Pseudomedon obsoletum (Nordmann) (designated by Blackwelder, 1939); of Ramona, R. capitulum Casey (monobasic) 3 of Ophiomedon, O. stipes Sharp (designated by Blackwelder, 1939) ; of Stilocharis, S. longula Sharp (monobasic). Diagnosis—Body moderately slender, parallel, moderately con- vex; head generally finely and irregularly but densely punctate, with or without ground sculpture; eyes generally at less than their length from base; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, first two not much larger than rest; labrum with a median tooth which is prominent on the upper sur- face or with paired denticles; fourth segment of maxillary palpus very small, subulate; gular sutures variable, prosternum carinate in part, sometimes prolonged under the coxae but not dilated laterally ; 240 BULLETIN 182, UNITED STATES NATIONAL MUSEUM hypomera sometimes feebly lobed behind the coxae; anterior coxal cavities entirely open behind, confluent; front coxae large, exserted ; middle coxal cavities confluent; posterior coxae contiguous, coni- cal; first and second abdominal sternites absent; seventh sternite of male sometimes slightly emarginate and with a ctenidium along pos- terior margin; eighth sternite of male emarginate. Figure 1.—Apical abdominal sternites of males of Lithocharis: A, L. dorsalis Erichson; B, L. secunda, new species; C, L. ochracea (Gravenhorst); D, L. sororcula Kraatz. Remarks.—This genus has been a source of difficulty for many years and has been given very different status by various writers. Casey separated it as a subtribe distinct from J/edon and most other writers have considered it to be a subgenus of Afedon. In my revi- sion of the Paederini (Blackwelder, 1939) I endeavored to separate these two as valid genera and assigned several subgenera to each. The genotype of Lithocharis is nearly cosmopolitan, but the generic char- acters are not pronounced or obvious. I assign nine West Indian species to this genus, three of them new. Of these, five belong in the typical subgenus, three in the sub- genus Stilocharis, and one in the subgenus Ophiomedon. The sub- genus Pseudomedon is apparently not represented in the West Indies. I have examined 653 examples of this genus from the West Indies. Of these, 270 are in the British Museum, 35 in Dr. Cameron’s collec- tion, 40 in the United States National Museum, and over 800 col- lected by me in 1935-37. KEY TO WEST INDIAN SPECIES OF LITHOCHARIS & 1, Gular- sutures converging posteriorly{o<2) 2. L-=- ees te 2 Gular.suturesparallelfalonge middle=—s= 8.22 fe see 9. heres 2. Labrum with a median tooth or prominence_______________-____&______ 3 Labrumy with two denticles near ‘center: 22-222 -- a ee 6 3. Seventh sternite of male with a ctenidium on posterior margin____-_-_- 4 Seventh sternite of male unmodified__-___________-_____________- 4. volans 18 The species LL. mendacia Cameron is unknown to me, and I am unable to include it in the key since the necessary characters are not mentioned in Dr. Cameron’s description. It is placed as No. 5 in the text. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 241 4. BHighth sternite with two groups of long golden hairs posteriorly____-_____ 5 Eighth sternite without groups of long golden hairs posteriorly__ 1. sororcula 5. Ctenidium of equal spinules, occupying less than one-third of width of seg- YOU VEYA Bak gt ee gh ae NR A ag le acetate Oe Ek SEER 3. secunda Ctenidium with spinules smaller at ends, occupying more than one-third of width of segment when viewed from below__---------------- 2. ochracea GssAnteriorstarsindilated=..225 25 8s 5 2 ee tk 8. posticata IATICCEIOIe taESIe NOU SOT UCC es se a ee ee ae 7 7. Pronotum with midline shining, indistinctly sculptured_-_-_-__-- a 6. limbata Pronotum with midline sculptured about as densely as rest-_----- 7. dorsalis 1. LITHOCHARIS SORORCULA Kraatz Lithocharis sororcula KRraatz, 1859b, p. 140.—BrERNHAUER and ScHUBERT, 1912, p. 243.—CAmERON, 1928, pp. 245, 247; 1931, pp. 156, 162; 1983, p. 40.— BLACKWELDER, 1939a, p. 102. Medon sororculum (Kraatz) BERNHAUER and ScHuBrERT, 1912, p. 243.—ScHEER- PELTZ, 1933, p. 1262.—BLACKWELDER, 1939a, p. 102. Description —Agreeing in all respects with the description of ochracea except for the male characters. J/ale, seventh sternite with a slight emargination posteriorly throughout the middle half of the margin and occupied by a ctenidium of black fimbriae which are shorter in the middle and at the ends, also with a few unusually large setae projecting posteriorly over the ctenidium from the pos- terior surface of the segment; eighth sternite with a rather deep rounded emargination which is narrowly bordered at apex, but with- out trace of the large clusters of golden hair found in ochracea. Length, 3 to 4 mm. T'ype locality.—Ceylon. Types.—Presumably in the Deutsche Entomologische-Institut, Ber- lin-Dahlem. Records.—The following are the records known to me: Jamaica: Santa Cruz (Blackwelder station 421), Milk River (Blackwelder sta- tion 415), Spanish Town (Blackwelder station 377). Puerto Rico: Salinas (Blackwelder station 60), San Juan (Blackwelder stations 345, 353B), Lajas and Bokueron (Dozier, in U.S.N.M.). St. Croix: (Blackwelder stations 317, 325, 330, 331, 341). St. Kitts: (Blackwelder stations 297, 299, 300, 302, 305, 306, 312). Antigua: (Blackwelder stations 275A, 275B, 280B, 283, 284, 285B, 287, 290). Montserrat: (Blackwelder stations 265B, 267B). Guadeloupe: Basse Terre (Blackwelder stations 70, 75), Grande Terre (Black- welder station 86). St. Lucia: (Blackwelder station 225). Barbados: (Blackwelder stations 188, 194, 196). St. Vincent: (Blackwelder stations 180, 185). Grenada: (Blackwelder stations 131, 150, 151). Orient: CeYLon, INprA, SINGAPORE (Kraatz, 1859; Cameron, 1928, 1931). Specimens examined.—I have examined 5 examples in Dr. Cam- eron’s collection, 4 in the United States National Museum, and 90 collected by me in 1935-87. 242 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks —Although very similar to ochracea in all other char- acters I have observed, this species is very distinct by the male struc- ture of the seventh abdominal sternite. The West Indian examples seem to be exactly similar to the oriental ones as figured by Dr. Cam- eron (1931). It has not been possible to associate definitely with these males any of the females of the ochracea complex. ‘This species belongs in the subgenus Lithocharis s. str. I have collected this species from horse manure and cattle dung, in rotting grass, and flying at dusk. 2. LITHOCHARIS OCHRACEA (Gravenhorst) Paederus ochraceus GRAVENHORST, 1802, pp. 58, 60. Paederus rubricollis GRAvENHOorsT, 1806, p. 188. Sunius rubricollis (Gravenhorst) STEPHENS, 1829, p. 287. Paederus obscurus Kirpy, MS.—StTEPHENS, 1829, p. 287. Sunius ochraceus (Gravenhorst) STEPHENS, 1829, p. 282. Rugilus rubricollis (Gravenhorst) MANNERHEIM, 1830, p. 40. Lithocharis ochracea (Gravenhorst) BotspuvAL and LAcORDAIRE, 1835, pp. 432, 433. Sunius brunniceps FAtRMAIRE, 1849, p. 290. Lithocharis fastidiosa FairMAIRE and GERMAIN, 1861, p. 488. Tjithocharis ochrea SUMMERS, 18738, p. 192 (misspelling). Lithocharis brunniceps (Fairmaire) FAuveL, 1878, p. 66. Medon ochracea (Gravenhorst) CHyzkEr, 1885, pp. 10, 19. Metaxyodonta alutacea CasrEy, 1886a, p. 380. Metaxyodonta quadricollis CAsry, 1886a, pp. 31, 32. Lithocharis alutacea (Casey) Casey, 1886b, p. 282. Tithocharis quadricollis (Casey) CASEY, 1886b, p. 232. Lithocharis rubricollis (Gravenhorst) SerpLirz, 1891, p. 373. Medon (Lithocharis) ochraceus (Gravenhorst) DrEvitL£, 1907, p. 141, 149. Medon (Lithocharis) alutacea (Casey) BERNHAUER and ScHusert, 1912, p. 242. Medon (Lithocharis) brunniceps (Fairmaire) BerRNHAUER and SCHUBERT, 1912, p. 242. Medon (Lithocharis) fastidiosus (Fairmaire and Germain) BERNHAUER and ScHvupert, 1912, p. 242. Medon (Lithocharis) quadricollis (Casey) BERNHAUER and ScHusert, 1912, p. 243. Medon (Lithocharis) rubricollis (Gravenhorst) BrERNHAUER and SCHUBERT, 1912, p. 248. Medon (Arthocharis) ochracea (Grayenhorst) CAMERON, 1921, p. 372. Description—Color variable but generally as follows: Head black, pronotum, elytra, and abdomen rufous, elytra and abdomen fre- quently clouded with castaneous. ead scarcely wider than long; eyes moderate, separated from base by about their length or a little less; posterior angles very broadly rounded, sides and base not dis- tinct; neck about two-fifths as wide as head; gular sutures moder- ately separated, straight, converging from apex to base; labrum not distinctly emarginate, with a very small and blunt tooth at middle not projecting beyond the margin but being the end of a feeble ridge MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 243 on the upper surface; with a few scattered punctures obscured by very irregular ground sculpture, which is chiefly composed of minute punctulae. Pronotum as broad as long, slightly wider than head; sides nearly parallel, angles rounded; without punctures but with rather flat tuberculi throughout; without ground sculpture. Llytra as long as wide, one-fourth wider than pronotum; with fine and mod- erately dense tuberculate sculpture but not with distinct tuberculi separated by smooth surface as on pronotum. Abdomen very finely and densely punctured and pubescent. Male, seventh sternite at middle of posterior margin with a comb of short black setae occupy- ing about the middle half of the segment and directed strongly in- wardly to give the appearance of a narrow striated band on the pos- terior border of the segment, the setae diminishing in size at the sides and bounded by a small group of hairs; eighth sternite with a rather broadly rounded notch, and laterally with two thick brushes of very long golden hairs. Female, sternites not modified. Length, 3 to 4 mm. Type locality—Germany, duchy of Brunswick. Of rudbricollis, not recorded but presumably Germany; of brunniceps, “Taiti”; of fastidiosa, Santiago, Chile; of alutacea, Santa Clara County, Califor- nia; of guadricollis, Lake County, California. Types.—Presumably in the Zoologische Museum, Berlin. Of rub- ricollis, same; of brunniceps, possibly in the Muséum Royal d’His- toire Naturelle de Belgique, Brussels; of fastidiosa, same; of alutacea, holotype, male, U.S.N.M. No. 38207, in the Casey collection in the United States National Museum; of quadricollis, holotype, male U.S.N.M. No. 38208, in the Casey collection in the United States National Museum. fecords.—The following are the records known to me: Jamaica: (Cameron, 1931), Santa Cruz (Blackwelder station 421), Spanish Town (Blackwelder station 377), May Pen (Blackwelder station 425C). Puerto Rico: Salinas (Blackwelder station 60), San Juan (Blackwelder station 345), Lajas, Guanica, Ponce (Dozier, in U.S.N.M.). St. John: (Erichson, 1840). St. Kitts: (Blackwelder stations 302, 306). Antigua: (Blackwelder stations 281A, 287). Guadeloupe: Basse Terre (Blackwelder station 75), Grande Terre (Blackwelder station 86). Dominica: (Blackwelder stations 237, 240, 252). Barbados: (Blackwelder stations 194, 195). Grenada: (Blackwelder station 150; British Museum; U.S.N.M.). South America: Brazit (Hrichson, 1840), Cotne (Fauvel, 1868; Fairmaire, 1861, as fastidiosa). Central America: GUATEMALA (Sharp, 1886). North America: CatrrorNIA (Casey, 1886, as alutacea and quadricollis). Europe: (Hrichson, 1839a, 1840; Deville, 1907 ; Fowler, 1888). 244 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Africa: (Gridelli, 1930), ZANzrBarR, Mauritius (Cameron, 1931), MADEIRA, CANARIES (Wollaston, 1854-65). Orient: New Gurnea, TAHITI, CeLEBES, CreyLon, InpIA, CHina (Fauvel, 1878), Inp1a (Cameron, 1931). Specimens examined.—Although I have seen numerous specimens labeled ochracea in all collections, and many of these are undoubtedly properly identified, the only specimens of which I have examined the male characters are seven in the United States National Museum and 27 collected by me in 1935-37. Remarks.—Only males have been included in the records of this and related species because of the difficulty of separating the females. The male characters agree closely with those illustrated by Cameron (1931) and are readily distinguished from all others I have seen. This species is almost cosmopolitan, having been recorded from nearly every part of the world except Australia and the northern parts of Europe and Asia. Its extensive synonymy is in large part due to this distribution, but it should be noted that not all the synonymy has been specifically checked in respect to the male characters. I have examined the types of alutacea and quadricollis in the Casey collection and find them to be identical with ochracea in this respect. This species is, of course, the genotype of the subgenus Litfhocharis s. str. I have collected this species from horse manure and cattle dung, in cocoa pods, and flying at dusk. 3. LITHOCHARIS SECUNDA, new species Description—Agreeing in all respects with the description of ochracea except for the male characters. J/ale, seventh sternite with a slight abrupt emargination of the posterior margin occupying the center fourth and filled with a ctenidium of short equal black fimbriae, strongly depressed entad and abruptly limited laterally, with a few abnormally large setae along margin at sides of the ctenidium; eighth sternite with a rather broad rounded emargination, much wider than deep, and with a large brush of long golden hairs on each side. Length, 21% to 4144 mm. Type locality —Jamaica, between Luana and Newport by way of Lacovia, Santa Cruz, and Spur Tree, parishes of St. Elizabeth and Manchester. Types.—Holotype and 32 paratypes, males, U.S.N.M. No. 52396, collected by Chapin and Blackwelder on February 24, 1937 (holotype), and during 1935-87. The paratypes are also from the localities listed below. Records. Jamaica: Milk River (Blackwelder station 415), Spanish Town (Blackwelder station 877), May Pen (Blackwelder station 425C), Black River (Blackwelder station 420), Newport (Blackwelder station 412A), Santa Cruz (Blackwelder station 421). The following are the records known to me: MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 245 Hispaniola: DoMINICcAN REPUBLIO: San José de las Matas (Blackwelder station 38). Be Rico: Salinas (Blackwelder station 60). St. Croix: (Blackwelder stations 330, 339). Antigua: (Blackwelder stations 275A, 280B, 285B, 290). Montserrat: (Blackwelder station 270). Guadeloupe: Basse Terre (Blackwelder station 70). St. Lucia: (Blackwelder station 213). Specimens examined.—tI have seen only the 33 types. Remarks.—This species is rather close to ochracea in its male char- acters as well as all other characters observed. It differs constantly, however, in the form and size of the ctenidium of the seventh sternite and can be distinguished at a glance even with low magnification. As with the other species of this complex no females can be definitely associated with these males. It belongs in the subgenus Lithocharis s. str. I have collected this species from cattle dung and horse manure and flying at dusk. 4. LITHOCHARIS VOLANS, new species Description.—A greeing in all respects with the description of ochra- cea except for the male characters. fale, seventh sternite with a very slight but broad emargination; eighth sternite with a shallow but angular emargination with the lateral angles rounded. Length, 3 to 344 mm. Type locality —Jamaica, Hope Gardens, Kingston. Types.—Holotype, male, U.S.N.M. No. 52397, collected by me on July 4, 1935, and one paratype collected by Chapin and Blackwelder on February 24, 1937, near Santa Cruz, parish of St. Elizabeth. Fecords.—The following are the records known to me: Jamaica: Kingston (Blackwelder station 1C), Santa Cruz (Blackwelder station 421), Milk River (Blackwelder station 415). {Puerto Rico: Salinas (Blackwelder station 60).] Specimens examined—I\ have seen only the two types and three other specimens mentioned below. Remarks.—This species differs from all the others of this group known to me in lacking any ctenidial modification of the seventh sternite in the male, but the presence of the single median tooth appears to be sufficient to establish it as a Lithocharis, at least at present. A second specimen from Santa Cruz differs from the type in having the emargination of the eighth sternite broader, deeper, and rounded at apex. The specimen from Puerto Rico also differs in the same character, having the emargination much more abrupt, scarcely wider than deep, and with all the angles narrowly rounded. These specimens were collected from dung and manure, and flying at dusk. 246 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 5. LITHOCHARIS MENDACIA Cameron Lithocharis mendacius CAMERON, 1923, p. 399. Medon (Lithocharis) mendacius (Cameron) SCHEERPELTZ, 1933, p. 1261. Description — Black, greasy-lustrous, the thorax pitchy-red, the elytra shining castaneous-red, largely infuscate posteriorly. Abdomen brown. Antennae and legs reddish testaceous. Length 4.4 mm. Closely allied to L. mendax Shp., and differing only in the following points: The antennae are a little shorter and the penultimate joints more transverse, the eyes are smaller, the head is densely coriaceous, with a few scarcely visible obsolete punctures, the thorax is sculptured similarly to the head, but the fine obsolete punctures are more numer- ous; the elytra are differently coloured, the abdomen is reddish brown, é unknown.” (From Cameron.) Type locality —Jamaica. Types.—Unique female in the collection of Dr. Cameron. Records.—The following is the only record known to me: Jamaica: (Cameron, 1923). Specimens examined.—I have seen only the unique type, which I examined briefly in Dr. Cameron’s collection. Remarks.—I am unable to place this species either in my key or among my specimens because of the absence of the male characters. The descriptions seems to indicate that it is a Lithocharis in the strict sense. I find no record of its habits. 6. LITHOCHARIS LIMBATA Erichson Lithocharis limbata EricHson, 1840, p. 621.—SHArpP, 1886, pp. 550, 551—Lrne and MutTcHter, 1914, p. 405. Medon limbatus (Hrichson) BERNHAUVER and ScHuUBERT, 1912, p. 242.—BLAcK- WELDER, 1939a, p. 102. Lithocharis (Stilocharis) limbata Erichson, BLACKWELDER, 1939a, p. 102. Description.—Rufopiceous, elytra castaneous to rufocastaneous with sides and apex testaceous, abdomen castaneous to rufocastaneous. Head subquadrate, base somewhat diagonal on each side, angles nar- rowly rounded; eyes large, separated from base by about half their length; labrum with a broad shallow emargination with two moderate denticles; gular sutures moderately separated in front, very approxli- mate behind; with punctures almost completely obscured by dense but fine ground sculpture. Pronotum with irregular and somewhat in- distinct punctures separated by shining intervals, midline shining, not distinetly sculptured. ytra rather densely and tuberculately punc- tured. Anterior tarsus not dilated. Male, eighth sternite truncate. Female, eighth sternite prominently rounded. Length 314 to 414 mm. Type locality—Colombia. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 247 Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Hispaniola: Harri, Dessalines (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: San Juan (Blackwelder stations 45, 345, 353B), Mayagiiez (Black- welder station 50B), Villalba (Blackwelder station 62). St. Croix: (Blackwelder station 330). Guadeloupe: Basse Terre (Blackwelder stations 75, 77B), Grande Terre ( Black- welder station 86). Dominica: (Blackwelder stations 235B, 286, 237, 245, 248A, 246B; Busck, in. U.S.N.M.). Martinique: (British Museum). St. Lucia: (Blackwelder stations 207C, 220C, 220D, 221, 225, 226). Barbados: (Blackwelder stations 198B, 194, 195, 188, 201A). St. Vincent: (British Museum; Cameron collection ; Blackwelder stations 165, 168, 180, 185). Grenada: (British Museum; Cameron collection; Blackwelder stations 137, 139). Trinidad: St. Joseph (Blackwelder station 180), Manzanilla (Blackwelder station 108A). South America: Colombia (Erichson, 1840; Sharp, 1886; British Museum). Central America: GUATEMALA (Sharp, 1886; British Museum), PANAMA (Sharp, 1886; British Museum; Bierig, in the Cameron collection). Specimens examined.—I have examined 239 specimens from the West. Indies in the British Museum, 7 in the collection of Dr. Cameron, 1 in the Museum of Comparative Zoology, 8 in the United States National Museum, and 100 collected by me in 1985-37; also a few specimens from Central America. Remarks.—The identification of these West Indian specimens with the Colombian species has not been verified by me, although I have examined the latter briefly without noting any differences. The species is somewhat variable in color but can be separated by the characters in the key. It belongs in the subgenus S/élocharis. I have collected it from dung, manure, and excrement, and from rotten fruit, decaying cocoa pods, and fermenting cocoanut husks. 7. LITHOCHARIS DORSALIS Erichson Lithocharis dorsalis Er1tcHson, 1840, p. 616.—FLEUTIAUX and SALxLs, 1889, p. 380.— LENG and Murcutkrr, 1914, p. 405—Wotcorr, 1924, p. 78; 1936, p. 197. Medon (Lithocharis) dorsalis (Erichson) BERNHAUER and SCHUBERT, 1912, p. 242, Lithocharis obfuscata CAMERON, 1928, p. 399. Medon (Lithocharis) obfuscatus (Cameron) SCHEERPELTZ, 1933, p. 1261. Description—Head rufopiceous, pronotum rufous, elytra castaneous to testaceous with sides and apex paler, abdomen castaneous. Head subquadrate with posterior angles rounded and often robust; eyes at their length or more from base; labrum broadly emarginate, with two blunt denticles within the emargination ; gular sutures widely separated at base and apex but most approximate behind middle; with discrete 248 BULLETIN 182, UNITED STATES NATIONAL MUSEUM though somewhat indefinite fine punctures, with vague ground sculp- ture on shining intervals. Pronotwm punctured similarly to head ; mid- line not distinctly shining. LZlytra feebly tuberculately punctured but not densely tuberculately sculptured. Anterior tarsus not dilated. Male, eighth sternite with a rounded emargination about five times or more as wide as deep. Female, eighth sternite not modified. Length, 214 to 234 mm. Type locality—Puerto Rico. Of obfuscatus, Jamaica. Types.—Khither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of obfuscatus, type and 11 others in the collection of Dr. Cameron, two cotypes and one other in the British Museum. Pecords.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.). Jamaica: (Cameron, 1923; British Museum; Cameron collection), Spanish Town (Blackwelder station 377), Newport (Blackwelder station 412A), Balaclava (Blackwelder station 397B), Trinityville (Blackwelder station 428), Milk River (Blackwelder station 415), Moneague (Blackwelder station 3875), May Pen (Blackwelder station 425C), Whitfield Hall in Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Hart1, Ennery (Darlington, in M.C.Z.) ; DomMIntcAN ReEpusriio, Con- stanza (Darlington, in M.C.Z.), Mount Quita (Darlington, in M.C.Z.). Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914; Wolcott, 1924, 1986), Adjuntas (Blackwelder station 48C), Maricao (Blackwelder station 47A), Villalba (Blackwelder station 62), San German (Dozier, in U.S.N.M.). St. Croix: (Blackwelder station 325). Antigua: (Blackwelder station 292). Montserrat: (Hubbard, in U.S.N.M.). Guadeloupe: (Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). St. Lucia: (Blackwelder stations 207B, 220D, 226, 230). St. Vincent: (British Museum; Cameron collection). Grenada: (Blackwelder station 132; British Museum; Cameron collection). Trinidad: (Fry collection, in British Museum), Manzanilla (Blackwelder stations 102, 104A), Moruga (Blackwelder station 111). Specimens examined.—tI have examined 24 specimens in the British Museum, 22 in the collection of Dr. Cameron, 6 from the Museum of Comparative Zoology, 14 in the United States National Museum, and 51 collected by me in 1935-37. Remarks.—This species has not been reported under Erichson’s name since the original description. The description seems to apply well to the species named obfuscatus by Cameron, except for the statement “tarsis . . . anticis leviter dilatatus.” This is perhaps an error, since the tarsus is rather stout and flattened beneath even in these “undilated” forms. It belongs in the subgenus Stilocharis. This is a variable species, but I have not been able to find any basis for dividing it. All specimens from the West Indies that I have seen labeled infuscatus Erichson belong to this species, and if they are actually MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 249 infuscatus then obfuscatus becomes a synonym. I do not have South American examples of ¢nfuscatus for comparison. My specimens were collected from decaying cocoa pods, from rotting banana stalks, from dung, and flying at dusk. 8. LITHOCHARIS POSTICATA Erichson Lithocharis posticata ErtcHson, 1840, p. 619.—LeNa and MurTcuHter, 1914, p. 405.—WotcottT, 1924, p. 78; 1936, p. 197. Medon (Lithocharis) posticatus (Hrichson) BERNHAUER and ScHuBeErT, 1912, p. 248. Description.—Piceous, elytra testaceous with apical third blackish, apex of abdomen testaceous. //ead a little broader than pronotum ; posterior angles somewhat rounded; labrum bidenticulate; densely punctate, smooth in front at middle, finely sparsely pubescent, shin- ing. Pronotum a little narrower than elytra, slightly wider than long, moderately narrowed posteriorly; densely but not very finely punc- tate, with smooth midline; finely and sparsely pubescent, shining. Hlytra one-third longer than pronotum; densely finely punctate, finely pubescent. Abdomen very finely punctate, with longer pubes- cence. Anterior tarsus feebly dilated. Jale, eighth sternite tri- angularly excised. Female, unknown. Length, 2 mm. (From Erichson. ) Type locality—Puerto Rico. x Types—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. fecords.—The following is the only record known to me: Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914; Wolcott, 1924, 1936). Specimens examined.—lI have seen no examples of this species. Remarks.—Apparently this species has never been seen since it was described by Erichson. I am not able to refer to it definitely any of the specimens before me and therefore include it as an unknown species. It appears to belong to the subgenus Stilocharis. I find no record of its habits. 9. LITHOCHARIS HERES, new species Description.—Rufotestaceous. Head one-ninth wider than long, not emarginate at base, sides evenly rounded into base; eyes moderate, separated by more than their length from base; labrum with two denticles within a broad emargination; gular sutures moderately close, parallel along middle, diverging at both ends; with sparse punctures (possibly somewhat umbilicate) completely obscured by very dense scaly ground sculpture. Pronotum as wide as head, nearly as long as wide; sides feebly converging with all angles strongly rounded; disk without impressions or midline; with punc- tures and sculpture as on head. “lytra not distinctly punctate but 44900S—42——17 250 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with tiny elevated setigerous tuberculi, rather dense and even; pub- escence moderately long. Anterior tarsus feebly dilated. Abdomen finely asperately punctate. Male, seventh sternite not modified; eighth with a small rounded emargination. Female, sternites not modified. Length, 314 to 3144 mm. Type locality—Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, male, and five paratypes, U.S.N.M. No. 52409, collected by E. A. Schwarz on January 138 (also January 6, March 2, and December 19) of 1902, 1903, or 1904. One paratype has been deposited in the collection of Dr. Cameron and one in my own collection. Records.—The following is the only record known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Specimens examined.—I have seen only the eight types in the United States National Museum and one other example in the Museum of Comparative Zoology. Remarks.—The position of this species is not altogether certain. The dense ground sculpture obscures the punctures to a large extent, but it is possible that it belongs closer to Medon. It is chiefly dis- tinguished by the form of the gular sutures. It belongs in the sub- genus Ophiomedon. I have no record of its habits. XXXIII. Genus ADEROCHARIS Sharp Aderocharis SHARP, 1886, p. 552. Subgenus Panscopaecus SHARP, 1889, p. 262. Subgenus Dorocharis BLACKWELDER, 1939a, p. 99. Genotypes.—Paederus corticinus Gravenhorst= Aderocharis corti- cinus (Gravenhorst) (designated by Blackwelder, 19392). Of Doro- charis, Aderocharis (Dorocharis) chapini Blackwelder (monobasic and original designation) ; of Panscopacus, P. lithocharoides Sharp (monobasic). Diagnosis.—Body rather stout, moderately convex, head generally emarginate at base, with angles prominent, punctures distinctly um- bilicate (in the genotype) or dense and of indefinite shape, with or without ground sculpture; eyes at more than their length from base; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, first two not much larger than rest; labrum with median tooth but without paired denticles; fourth segment of maxil- lary palpus very small, subulate; gular sutures not at all united, rather widely separated in front but very approximate posteriorly; punctures of pronotum as on head or tuberculate; prosternum cari- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Zo nate in part, not greatly prolonged between coxae; hypomera pro- longed in a short lobe partially behind the coxae; anterior coxal cavities open behind, confluent; front coxae very large, exserted; middle coxal cavities confluent; posterior coxae contiguous, conical ; first and second abdominal sternites absent, seventh sternite more or less modified in the male, usually biemarginate, eighth emarginate in male only; anterior tarsus dilated. Remarks.—This genus is not strongly distinguished from Lithocharis and may be only a subgenus of it. I am led to keep it separate because of the ease with which it can be separated at a glance by its much Ficure 2.—Apical abdominal sternites of males of Aderocharis: A, A. cubensis, new species; B, A. suturalis, new species; C, A. dubitans, new species; D, 4. conifer Cameron. greater size. The male characters are also quite different and, as in Lithocharis, are the only satisfactory means of separating many of the species. I have examined 59 examples from the West Indies. Of these 19 are in the British Museum, 11 in the collection of Dr. Cameron, 5 in the Museum of Comparative Zoology, 8 in the United States National Museum, and 16 were collected by me in 1935-87. KEY TO WEST INDIAN SPECIES OF ADEROCHARIS 1. Seventh abdominal sternite of male emarginate__________________________ 2 Seventh abdominal sternite not at all emarginate______-________ 1. cubensis 2. Seventh abdominal sternite of male biemarginate___.____________________ 3 Seventh abdominal sternite of male with three emarginations__ 4. suturalis 38. Emarginations separated by a large process_____________________ 3. conifer Emarginations feebly separated by a flat are___--_-__-_________ 2. dubitans 1. ADEROCHARIS CUBENSIS, new species Description.—Rufocastaneous. Head not distinctly emarginate at base nor with very prominent angles but eyes small and separated from base by one-half more than their length; gular sutures convergent toward base but separated throughout in the male; antennal segments 252 BULLETIN 182, UNITED STATES NATIONAL MUSEUM becoming distinctly broader apically; with very fine and dense but distinctly umbilicate punctures, and with a few larger scattered um- bilicate punctures; with irregular ground sculpture at sides posteriorly. Pronotum with a poorly defined smooth midline; with umbilicate punctures and sculpture at sides as on head. Llytra not distinctly punctate, but with rather dense fine tuberculi, sometimes arranged in irregular transverse rows, and bearing fine short hairs. Abdomen rather densely and finely but distinctly muricately punctured; with dense short pubescence. J/a/e, seventh sternite slightly sinuate pos- teriorly but not at all emarginate; eighth sternite with a very large triangular emargination. Jemale, sternites not modified. Length, 414 mm. Type locality —Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, male, U.S.N.M. no. 52898, collected by KE. A. Schwarz on February 15 of 1902, 1903, or 1904; two paratypes, female, in the Museum of Comparative Zoology, one in the United States Na- tional Museum, from Soledad. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U:S:N.M:): Specimens exvamined.—I have seen only the four types. Remarks.—This species is unique in its feeble male characters as well as in the shape of the head. The holotype is slightly immature, being of paler color and having the punctation a little less distinct and deep, though of the same type and density. The examination of the species related to Lithocharis by use of the male genitalia may throw addi- tional light on their proper position, but at present this species ap- pears to be best placed in this genus. I have no record of its habits, although one specimen was taken at light. 2. ADEROCHARIS DUBITANS, new species Description.—Head black, pronotum and abdomen piceous, elytra and apex of abdominal segments rufopiceous. Head emarginate posteriorly, with angles moderately prominent; eyes small, separated from base by less than one-half more than their length; gular sutures in male converging posteriorly, never very distant; densely and mod- erately finely punctate, punctures not definite in shape, with large setig- erous punctures at sides, pubescence fine and dense; without distinct ground sculpture. Pronotum with distinct narrow smooth midline; with dense tuberculi unusually flat, appearing almost like small um- bilicate punctures; with short pubescence but no ground sculpture on disk. Hlytra with tuberculi very dense, scarcely distinguishable, some- times forming transverse rows; apparently with additional ground MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 258 sculpture; pubescence dense and distinct. Abdomen very minutely and densely punctulate and pubescent; with indistinct ground sculpture. Male, seventh sternite with two emarginations separated by a broad feebly rounded lobe and bounded laterally by two broadly triangular processes; eighth with a deep rounded emargination nearly twice as wide as deep. Yemale, unknown. Length, 6144 mm. Type locality—Trinidad, 5 miles west-southwest of Manzanilla junction on the Plum Road. Types.—Holotype, male, U.S.N.M. No. 52399, collected by me on December 18, 1935. Records.—The following are the records known to me: Trinidad: Manzanilla (Blackwelder station 104A). Specimens examined —I have seen only the unique type. Remarks.—This species is distinguished principally by the male characters. Three specimens in the British Museum under the name Medon latro Sharp are probably this species, although I have not examined the male characters. They were taken at Caparo, Trinidad, by Klages in June 1914. Tt is possible that this species or the following one is the same as A. latro Sharp, but I am not able to form an opinion without seeing the male characters of the type. The latter is in the British Museum and is mounted under glass. The type was collected from decaying cocoa pods. 3. ADEROCHARIS CONIFER Cameron Aderocharis conifer CAMERON, 1918b, p. 8345.—LrNe and MurcHteER, 1917, p. 199.— BLACKWELDER, 1939a, p. 102. Aderocnaris obscurior CAMERON, 1913b, p. 346—LeENG and MurTrcHiEr, 1917, p. 199.— BLACKWELDER, 1939a, p. 102. Medon conifer (Cameron) SCHEERPELTZ, 1933, p. 1268—BLACKWELDER, 1939a, . 102. eae obscurior (Cameron) SCHEERPELTz, 1933, p. 1263.—BLAcKWELDER, 1939a, p. 102, Description —Piceous, pronotum rufopiceous, elytra and abdomen rufotestaceous with disk in each case more or less infuscate. Head emarginate posteriorly but with angles not very prominent; eyes small, separated from base by nearly twice their length; gular sutures converging posteriorly and never very distant; not differing between the sexes; very irregularly but distinctly punctate with dense pubes- cence, posteriorly becoming somewhat tuberculate, more dense, and with some ground sculpture. Pronotvm with distinct smooth mid- line, somewhat elevated; with dense but discrete tuberculi bearing dense pubescence, and mixed with ground sculpture at sides posteri- orly. Hlytra tuberculate as pronotum but a little more coarsely, without distinct group sculpture. Abdomen very finely and densely 254 BULLETIN 182, UNITED STATES NATIONAL MUSEUM punctulate and pubescent. Afale, seventh sternite with two feeble rounded emarginations separated by a large triangular lobe, each emargination interrupted by a protruding angulate process, the median lobe fimbriate posteriorly; eighth sternite deeply triangularly emarginate. Female, sternites unmodified. Length, 6 to 7 mm. Type locality.—St. Vincent, Leeward side. Of obscurior, Grenada, Chantilly Estate, Windward side. Types.—In the British Museum (for both names). Records.—The following are the records known to me: St. Vincent: (Cameron, 1913; Cameron collection; United States National Museum). Grenada: (Cameron, 1913, as obscurior ; Cameron collection). Specimens examined.—I have seen 19 examples (including the two types) in the British Museum, 11 in the collection of Dr. Cameron, and 6 (including one cotype) in the United States National Museum. The last were received from the original series by exchange. Remarks.—I was unable to distinguish the types of obseurior and conifer and believe them to be the same. The male characters are much more complex than in any other of our species. I am unable to verify the statement made by Dr. Cameron that the triangular lobe of the seventh abdominal sternite of the male is emarginate at the apex. The surface of the lobe is slightly concave throughout but the apical margin is not distinctly emarginate. I find no record of its habits. 4, ADEROCHARIS SUTURALIS, new species Description.—Piceous to rufous, pronotum, elytra, and abdomen a little more rufous, elytra with margin generally rufescent or rufo- testaceous. Head feebly emarginate at base, angles prominent; eyes small, at about one-half more than their length from base; gular sutures different in the sexes; not normally punctured but with mod- erately dense tuberculi with a minute puncture and fine hair at top of each; with irregular ground sculpture on base only. Pronotwm with distinct but indefinitely smooth midline; with tuberculi as on head but stronger; without ground sculpture. Hlytra with tuberculi as on pronotum but with pubescence more evident; without ground sculpture. Abdomen very minutely and very densely punctulate and pubescent. Jfale, gular sutures united throughout from small tri- angular submentum; seventh abdominal sternite with four small triangular processes separated by three rounded emarginations, the middle one a little wider than deep, the outer pair as deep as wide; eighth sternite very broadly, shallowly, and evenly emarginate. Female, gular sutures not united, moderately converging from sub- mentum to base; abdominal sternites not modified. Length, 5 to 6144 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 255 Type locality —Puerto Rico, 2 miles north-northwest of Villalba. Types.—Holotype, male, and 14 paratypes, 5 males and 9 females, U.S.N.M. No. 52400, collected in 1935-387 by Chapin and Blackwelder. The paratypes are from Villalba and the localities mentioned below. Also three paratypes in the Museum of Comparative Zoology, from Cuba, Jamaica, and Haiti. Records.—The following are the records known to me: Cuba: Soledad (Darlington, in M.C.Z.). Jamaica: (Wight, in M.C.Z.), May Pen (Blackwelder station 425C), Milk River (Blackwelder station 415), Santa Cruz (Blackwelder station 421), Newport (Blackwelder station 412A). Hispaniola: Haiti, Dessalines (Darlington, in M.C.Z.). Puerto Rico: Villalba (Blackwelder station 62), Adjuntas (Blackwelder station 49), Ensenada (Dozier, in U.S.N.M.). Specimens examined.—I have seen only the 18 types. Remarks.—This species is very distinct by the male characters as well as by the sculpture of the head. My specimens were collected from horse manure and flying at dusk. XXXIV. Genus STILOMEDON Sharp Stilomedon SHarp, 1886, p. 565. Subgenus Polymedon Casey, 1905, pp, 151, 155. Genotype.—Lithocharis connexa Sharp=Stilomedon connexum (Sharp) (designated by Blackwelder, 1939). Of Polymedon, Lith- ocharis tabacina Casey=Polymedon tabacinum (Casey) (original designation and monobasic). Diagnosis —Body elongate, moderately convex; punctures of head densely umbilicate, with or without ground sculpture; antennae pos- teriorly flexile, not strongly geniculate, basal segment not much elongate, first two not much larger than rest; labrum with median tooth and one or two pairs of denticles; fourth segment of maxillary palpus very small, subulate; gular sutures not at all united but narrowly separated, feebly diverging in front; punctures of pronotum normal, not umbilicate or tuberculate, without dense ground sculpture; prosternum not strongly carinate, prolonged between coxae as far as mesosternum but not much dilated under coxae; hypomera pro- longed in a short lobe partially behind the coxae; anterior coxal cavities open behind, confluent; front coxae very large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, third carinate at base; seventh sternite not modified in either sex, eighth feebly emarginate in male only. Remarks.—The comparison of the genotypes of Stilomedon and Polymedon shows them to be very similar in structure though differ- ing moderately in facies, principally in the shape of the pronotum. 256 BULLETIN 182, UNITED STATES NATIONAL MUSEUM I have seen from the West Indies 118 specimens representing three species. KEY TO WEST INDIAN SPECIES OF STILOMEDON 1. Head without ground sculpture; umbilicate punctures crowded_------~--- Z Head in part with ground sculpture; umbilicate punctures not crowded. 1. insularum o. Pronotum densely 1ugosely punctale=— = 22 eee 3. audanti Pronotum rather sparsely discretely punctate____---__----~-- 2. connexum 1. STILOMEDON INSULARUM (Cameron) Medon (Neomedon) insularum CAMERON, 1923, p. 899.—SCHEERPELTzZ, 1933, p. 1250.—BLACKWELDER, 1939a, p. 108. (Not insulare Casey, 1905.) Stilomedon insularum (Cameron) BLACKWEUDER, 1939a, p. 103. Description ——Piceous to rufous, pronotum generally a little more rufescent with lighter spots at the anterior angles, elytra generally paler than pronotum, tip of abdomen paler. Head quite distinctly emarginate at center of base with a short vertical groove in base of vertex, eyes small, at about twice their length from base; posterior angles moderately rounded; labrum with median tooth large, very prominent above, additional denticles very small and close to median; gular sutures rather narrowly separated but converging posteriorly ; surface with moderate umbilicate punctures, not very dense, intervals flat and with dense ground sculpture especially anteriorly. Pronotum with sides moderately converging posteriorly ; not or feebly impressed on disk; midline feebly channeled posteriorly; with moderately fine punctures which appear to be obsoletely umbilicate; with fine indis- tinct ground sculpture. lytra not distinctly punctate, rather densely set with posteriorly directed tuberculi which bear the fine but long pubescence; without ground sculpture except along suture. Abdomen very finely and densely asperately punctulate; pubescence rather long. Male, seventh sternite unmodified; eighth sternite with a very broad and very shallow emargination, but this bordered by long hairs directed to the sides. Female, sternites not modified. Length, 314 to 4 mm. Type locality —Jamaica. Types.—Unique female type in the collection of Dr. Cameron. Records—The following are the records known to me: Cuba: Santiago de las Vegas (Dimmock, in U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.). Jamaica: (Cameron, 1923), Trinityville (Blackwelder station 428). Hispaniola: Haiti, Port-au-Prince (Audant and Ducasse, in U.S.N.M.) ; DoMINI- OAN Repusiic, San Francisco Mountains (Busck, in U.S.N.M.), Mount Quita (Darlington, in M.C.Z.). Specimens examined.—I have seen the type in Dr. Cameron’s col- lection, six examples in the United States National Museum, two that were borrowed from the Service Technique (and later presented to the United States National Museum), one in the Museum of Com- parative Zoology, and one collected by me in 1935-37. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE PASY Remarks.—This species is readily distinguished by its dense ground sculpture. Some variation has been observed in the impressions and channeling of the pronotum. It belongs in the subgenus Stilomedon s. str. The examples from Cuba were taken “in ceiba” (silk cotton tree) and the one from Jamaica flying at dusk. 2. STILOMEDON CONNEXUM (Sharp) Lithocharis connexa SHARP, 1876, p. 254; 1886, p. 566—BLACKWELDER, 1939a, pp. 108, 122. Lithocharis convexa SHARP, 1876, p. 262 (typographical error) ; 1886, p. 566. Stilomedon connera (Sharp) SHarp, 1886, pp. 566, 567.—BeERNHAUER and ScHU- BERT, 1912, p. 230..-LeN@ and MutTcuter, 1914, p. 405.—Birnrtie, 1933, p. 489.— BLACKWELDER, 1939a, pp. 103, 122. Stilomedon convera (Sharp) SHarp, 1886, p. 566—BERNHAUER and SCHUBERT, 1912, p. 230. Description Rufous, elytra and abdomen sometimes rufotesta- ceous. Head not emarginate behind, hind angles broadly rounded; eyes small, at nearly twice their length from base; labrum not de- cidedly emarginate, with median denticle not very distinct, additional pair rather widely separated, large at base but acute and rather long; gular sutures very close but feebly converging posteriorly; surface with dense umbilicate punctures, the intervals frequently united in rows, without ground sculpture. Pronotwm with sides rather strongly converging posteriorly; with two feeble longitudinal impressions posteriorly on disk along middle; midline not distinctly channeled; punctures not umbilicate but moderately coarse, usually separated by less than their diameter; without ground sculpture ex- cept at posterior angles. Zlytra with indistinct punctation, which is evidently the result of the flattening of small tuberculi with the punc- tule on top; without distinct ground sculpture. Abdomen rather densely and finely but asperately punctured; with rather long and dense pubescence. Male, seventh sternite unmodified, eighth with a very broad rounded emargination. Female, sternites unmodified. Length, 314 to 4 mm. Type locality “Amazons” (=Amazon Valley, Brazil). Types—“Two males in the British Museum” (according to the criginal description, but a note in the collection of the British Museum says, “Type not found.” I found no South American exam- ples in that collection). Records.—The following are the records known to me: Cuba: (Leng and Mutchler, 1914; United States National Museum). Jamaica: Trinityville (Blackwelder station 428), Milk River (Blackwelder sta- tion 415), Santa Cruz (Blackwelder station 421), Bath in St. Thomas (Blackwelder station 892H). Hispaniola: Harri, Miragoane (Darlington, in M.C.Z.). Guadeloupe: (Leng and Mutchler, 1914). 258 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Dominica: (Blackwelder stations 235A, 287, 251). St, Lucia: (Blackwelder stations 221, 231). St. Vincent: (British Museum). Grenada: (Blackwelder station 1382; British Museum). Trinidad: Manzanilla (Blackwelder station 103A), Caura Valley (Weber, in M.C.Z.) South America: Braziz (Sharp, 1876, 1886), VeENEzUELA (Bernhauer and Schubert, 1912). Central America: PANAMA (Sharp, 1886; United States National Museum), Mexico (Bernhauer and Schubert, 1912). Specimens examined.—Beside a considerable number of specimens from Panama, I have seen 12 examples from the West Indies in the British Museum, 5 in the United States National Museum, and 69 collected by Chapin and Blackwelder in 1935-87. Remarks.—Even in a small series this species shows some variation in details of sculpture. Although I believe the West Indian exam- ples to be the same as the Panama ones (four of which were received from the Biologia material), I do not have males from Panama to compare with males from the islands. The above description was drawn from a St. Lucia example. This species belongs in the sub- genus Stilomedon s. str. We have collected it in rotting cocoa pods, flying at dusk, and flying to light at night. 3. STILOMEDON AUDANTI, new species Description.—Rufous, elytra generally rufotestaceous. Head fee- bly emarginate behind, posterior angles very broadly rounded; eyes small, at about twice their length from base; labrum with median tooth rather small, lateral teeth rather widely separated, moderately large but stout; gular sutures narrowly separately but converging poste- riorly; surface with rather large and dense umbilicate punctures, the intervals not much united in rows, flat, sometimes half as wide as punctures; without ground sculpture on disk. Pronotum with sides moderately converging posteriorly; scarcely impressed on disk; mid- line feebly channeled posteriorly; punctures normal, moderately coarse, dense, more or less confluent, intervals somewhat rugose and somewhat obscured by indefinite coarse ground sculpture. lytra not punctured ; with rounded tuberculi frequently separated by more than their diameter, and in part obscured by coarse ground sculpture; with three indistinct rows of larger setigerous punctures longitudinally on each elytron. Abdomen very finely, densely, and indistinctly punc- tulate; pubescence moderately long. Male, seventh sternite unmodi- fied; eighth very feebly triangularly emarginate. Female, sternites unmodified. Length, 3 to 4mm. Type locality —Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, Province of Santa Clara. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 259 Types.—Holotype, male, and seven paratypes, U.S.N.M. No. 52406, collected by E. A. Schwarz on January 5 of 1902, 1903, or 1904. Records.—The following are the records known to me: Cuba: (U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Baragua (Christenson, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Harti, Port-au-Prince (Audant, in U.S.N.M.). Puerto Rico: (U.S.N.M.), Ponce (Dozier, in U.S.N.M.). St. Croix: (Blackwelder stations 333, 341). Specimens examined.—Beside the types I have examined four ex- amples in the United States National Museum, two that were lent from the collection of the Service Technique in Haiti, two from the Museum of Comparative Zoology, and two collected by me in 1935-37. Remarks. —This species is rather similar to connexum but differs in the punctation and sculpture of the head, pronotum, and elytra, in the emargination of the base of the head, and in the shape of the labral denticles. It belongs in the subgenus Stilomedon s, stv. The types were taken “on palmetto”; one other specimen from Cuba was found “in soil of sugarcane fields”; one specimen from Puerto Rico and one from Cuba were found in trash on banana boats at Philadelphia; my specimens were caught flying at dusk. XXXV. Genus SUNIUS Stephens Sunius STEPHENS, 1829, p. 287. Subgenus Hypomedon Mutsant and Rey, 1878, p. 122. Chloécharis LyNcH, 1884, p. 257. Chloeocharis EICHELBAUM, 1909, p. 148 (misspelling). Cloecharis Biertc, 1934f, p. 328 (misspelling). Hemimedon Casty, 1905, pp. 152, 160. Lena Casey, 1805, p. 189. Euastenus Fiori, 1915, p. 10. Subgenus Sunius s. str. Oligopterus Casry, 1886a, p. 12. Medonella Casty, 1905, pp. 154, 180. Micromedon Casry, 1905, pp. 153, 155. Xenocharis Breria, 1934f, p. 328. Subgenus Caloderma Casry, 1886a, p. 5. Subgenus Trachysectus Casey, 1886a, pp. 32, 388. Genotypes.—Paederus melanocephalus Fabricius=Sunius melano- cephalus (Fabricius) (designated by Westwood, 1888). Of Chloe- charis, C. rufula Lynch (monobasic); of Oligopterus, O. cuneicollis Casey (monobasic) ; of Micromedon, Medon seminigrum Fairmaire= Micromedon seminigrum (Fairmaire) (monobasic); of Medonella, M. minuta Casey (monobasic and original designation) ; of H'wastenus, FE, pallidus Fiori (monobasic) ; of Xenocharis, X. occipitalis Bierig (monobasic and original designation) ; of Hypomedon, Lithocharis 260 BULLETIN 182, UNITED STATES NATIONAL MUSEUM debilicornis Wollaston=Hypomedon debdilicornis (Wollaston) (des- ignated by Blackwelder, 1939a); of Hemimedon, H. rujfipes Casey (designated by Blackwelder, 19392) ; of Lena, L. testacea Casey (mon- obasic) ; of Caloderma, C. rugosa Casey (designated by Blackwelder, 1939); of Trachysectus, Lathrobium confluentum Say=Trachysectus confluentus (Say) (monobasic and original designation). Diagnosis —Body elongate, moderately convex; punctures of head umbilicate, with or without dense ground sculpture; antennae pos- teriorly flexile, not strongly geniculate, basal segment not much elongate, first two not much larger than the rest; labrum without median denticle, with or without a pair of denticles near middle; fourth segment of maxillary palpus very small, subulate; gular su- tures not at all united, moderately distant, most approximate anteri- orly; punctures of pronotum similar to head in general; prosternum not carinate on disk, prolonged between coxae as far as mesosternum but not much dilated under coxae; hypomera prolonged in a short lobe partially behind coxae; anterior coxal cavities open behind, con- fluent; front coxae very large, exserted; middle coxal cavities conflu- ent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, third carinate at base, seventh and eighth not modified in either sex (unless males are unknown); anterior tarsi slender. Remarks.—Although the distinguishing characters of this genus are not very pronounced, it seems desirable and justifiable to recognize the variations within the old genus Medon by forming several genera, each of which has groups of species that can with propriety be made subgenera. The name /Zypomedon was at first adopted for this genus. When it was found that the name Sunius of Stephens was the valid name of one of the subgenera (previously cited under the name Oligop- terus), it became necessary to use the older name for the genus. The subgenera Hypomedon and Caloderma are the only ones besides the typical subgenus which have been found in the West Indies. The subgenus Sunius s. str. differs from both of these in having the head and pronotum sparsely punctate and shining. The subgenus Caloderma differs from Hypomedon in having the umbilicate punc- tures dense and in the absence of ground sculpture. The species Medonella minuta was described by Casey as possessing four denticles on the labrum. I am unable to see more than two on either of the types and therefore place the name Medonella as a synonym of Sunius gs. str. A “cotype” of Xenocharis occipitalis Bierig in the United States National Museum leaves no doubt that the genus Xenocharis is a synonym of Sunius s. str. Before receipt of this specimen I had MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 261 confused the species with Medonella ferrugenea (Sunius ferrugincus) , which is somewhat similar in both structure and appearance. Sr. Bierig’s figure of the two species shows them to be distinct but does not prevent the uniting of Xenocharis with Sunius s. str. I have seen 240 examples of this genus from the West Indies. They are assigned to nine species, of which four appear to be new. KEY TO WEST INDIAN SPECIES OF SUNIUS 1. Head and pronotum with very dense ground sculpture____-___-___-_------- 2 Head and pronotum without dense ground sculpture, shining__----_--~- 3 2. Elytra normally, moderately coarsely punctate; less than 3 mm. long. 8. debilicornis Elytra impunctate, with rather dense rounded tuberculi; 4 mm. long. 9. terresi S- Pronotum™= more or” less distinctly: punctatelee eer se wie | Ses 4 Pronotum finely? tuberculatest*2_ si2 eae Ute Sees ee aes ed oe 7. weberi 4. Eyes large, at less than twice their length from base__-_----__--________ 5 Byes small, at more than twice their length from base___-____-_________ 6 5. Pronotum with median smooth stripe not canaliculate_________ 6. curtulus Pronotum without median smooth stripe, but finely canaliculate posteriorly. 5. oblitus 6. Punctures of head usually separated by less than their diameter__________ 7 Punctures of head usually separated by more than their diameter________ 8 7. Sides of pronotum distinctly converging behind; length about 4 mm. 4. darlingtoni Sides of pronotum nearly parallel; length 2 to 244 mm __-____-_ 8. occipitalis 8. Head with moderate, somewhat umbilicate punctures; elytra not breader TSEL ELTA EL ee SRS REA SS SETS NE es By a ee ey 1. oakleyi Head with fine punctures; elytra broader than long--___-_-- 2. ferrugineus 1. SUNIUS OAKLEYI, new species Description —Rufopiceous, basal two-thirds of elytra indefinitely testaceous, abdomen castaneous with apex testaceous. Head one- thirteenth wider than long; base emarginate only above the neck; eyes very small, separated from base by over three times their length; sides slightly arcuate, basal angles rounded; labrum with a small triangular emargination at middle but without trace of denticle; gular sutures moderately separated, rather feebly converging apically; with large but not very distinctly umbilicate punctures generally separated by their diameter or more, with a smooth midline; without ground scu!p- ture. Pronotum one-twelfth wider than long, as wide as head; sides feebly converging posteriorly ; with punctures similar to those of head but denser, middle stripe indefinite, very vaguely irregularly chan- neled posteriorly; without ground sculpture. E/ytra one-eighth longer than wide, a trifle wider than pronotum; surface uneven, punc- tures scarcely evident; without ground sculpture. Abdomen finely but sparsely tuberculately punctate. Male, apex of seventh sternite trun- 262 BULLETIN 182, UNITED STATES NATIONAL MUSEUM cate or very vaguely emarginate; eighth with a broad rounded emargination less than one-third as deep as wide. Female, unknown. Length, 244 mm. Type locality —Puerto Rico, 4 miles southeast of Adjuntas, elevation about 3,000 feet. Types.—Holotype, male, U.S.N.M. No. 52402, collected by me on October 4, 1935; two paratypes from the Museum of Comparative Zoology, from El Yunque and Maricao, ecords.—The following are the records known to me: Puerto Rico: Adjuntas (Blackwelder station 48A), Maricao Forest (Darlington, in M.C.Z.), El Yunque (Darlington, in U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This species is named for R. G. Oakley, who was of great assistance to me in the collecting of this and numerous other species. This species is quite distinct from ferrugineus, but except for the denticles on the labrum it seems to be a normal Swnius s. str. The type was found in a very rotten stump. 2. SUNIUS FERRUGINEUS (Bierig) Medonella ferruginea BrERIc, 1934f, pp. 326, 329, fig. 3. Description —F¥errugineous. Head quadrangular, one-fourth longer than wide (from the figure, about as wide as long), basal angles rather broadly rounded, base not truncate; eyes separated from base by more than twice their length; with fine punctures separated by less than their diameter except on the vertex; without dense ground sculpture. Pronotwm as wide in front as head, as broad as long, sides feebly converging behind and rounded into base; punctures homogeneous, more variable in size, lacking in narrow irregular midline; without dense ground sculpture. Flytra reduced, one- seventh wider than long; much more coarsely and simply punctate, the intervals elevated and very finely irregularly coriaceous. Abdo- men with very uneven granulose sculpture. J/ale, eighth sternite with a large triangular incision. Length, 214 mm. (From Bierig.) Type locality——Cuba, Playa de Marianao. Types.—In the collection of Alexander Bierig. Records.—The following is the only record known to me: Cuba: Playa de Marianao (Bierig, 1934). Specimens examined.—I have seen no examples of this species. Remarks.—The characters given in the original description appear to be sufficient to place this species in the key, although its reference , to this genus is perhaps doubtful. It appears to belong to the sub- genus Sunius s. str. Bierig records this species from sand under dry seaweed. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 263 3. SUNIUS OCCIPITALIS (Bierig) Xenocharis occipitalis Brertc, 1934f, p. 329, figs. 1, 4—BLACK WELDER, 1939a, p. 122. Description—Rufotestaceous throughout. ead as long as broad, base rather strongly emarginate, triangular above the neck; eyes very small, separated from base by more than three times their length; sides slightly arcuate, basal angles narrowly rounded; labrum with two distinct denticles near the center; gular sutures moderately separate but very feebly converging apically; surface with rather small umbilicate punctures, separated by their diameter or less, absent along middle; without ground sculpture. Pronotum as long as broad, scarcely narrower than head; sides feebly converging posteriorly; with punctures similar to those on head but sparser, generally sep- arated by more than half their diameter and frequently more than their diameter; middle stripe smooth, slightly elevated posteriorly ; without ground sculpture. lytra barely longer than wide, as wide as pronotum; punctured as pronotum but less distinctly umbilicately ; somewhat coriaceous but without ground sculpture. Abdomen very minutely but not very densely asperately punctate. fale, seventh sternite flattened at middle and becoming feebly concave posteriorly, produced posteriorly in center half, the lobe with a rounded emargi- nation about half as deep as wide, eighth sternite with an emargina- tion of breadth equal to that of seventh but as deep as wide, set off by a minute inwardly directed tubercle at the inner edge of the pos- terior angles. Female, sternites not modified. Length, 2 mm. Type locality —Cuba. Types—In the collection of Alexander Bierig. (One “cotype” is in the United States National Museum (No. 52729) through the generosity of Sr. Bierig.) Records.—The following are the records known to me: Cuba: San Vicente, Sierra de los Organos (Bierig, 1984), Rangel, Sierra del Rosario (Bierig, 1934; Bierig, in U.S.N.M.), Aspiro (Bierig, 19384), Bejucol (Bierig, 1934), Cruz de Piedra, province of Habana (Bierig, 1934), Cayamas (Schwarz, in U.S.N.M.). Specimens examined.—I have seen one male and one badly broken female in the United States National Museum, in addition to the “cotype.” Remarks.—I am unable to separate the Cayamas specimens from the “cotype” of occipitalis. The description has been taken from the Cayamas examples. The punctation of the head and pronotum indicates that this species belongs in the subgenus Sunius s. str. It is very similar to Medonella minuta Casey (=Sunius minuta) from Florida from which it is dis- tinguished by the very small eyes and long temples as well as by de- tails of punctation. The types were found under fallen leaves, 264 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 4. SUNIUS DARLINGTONI, new species Description—Piceous, elytra rufocastaneous or rufous. Head trans- verse, not emarginate behind, with angles moderately rounded; eyes very small, at over three times their length from base; labrum with a small median emargination bordered by a pair of small blunt tubercles, and with a still smaller pair more laterally; gular sutures very closely approximated near base; with dense and fine but dis- tinctly umbilicate punctures generally separated by less than half their diameter; without distinct ground sculpture except in depres- sion between eyes and also behind eyes laterally. Pronotum nearly as wide as head, about one-fourth wider than long, the sides rather strongly converging and rounded into base; disk with a trace of im- pression on either side of midline near base; punctation very similar to that of head but obsolescent; without a distinct smooth midline; without distinct ground sculpture. Hlytra with rather indistinct submuricate punctures separated by a little more than their diam- eters; without definite ground sculpture. Abdomen sparsely minutely submuricately punctate, with indistinct ground sculpture. Afale, eighth sternite with a very small semicircular emargination. Female, sternites not modified. Length, 4 to 444 mm. Type locality—Dominican Republic, Loma Vieja, south of Con- stanza, elevation about 6,000 feet. Types.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology, collected in August 1938 by Dr. P. J. Darling- ton; two paratypes in the United States National Museum (No. 52510). (All three paratypes collected at Loma Rucilla and the moun- tains to the north in June 1938.) Records —The following are the records known to me: Hispaniola: DoMINICAN Repustic, Loma Vieja (Darlington, in M.C.Z.), Loma Rucilla (Darlington, in M.C.Z. and U.S.N.M.). Specimens cramined.—I have seen only the four types. Remarks.—This species is distinguished by the obsolescent pronotal punctures, the small eyes, and the denticles of the labrum, in addition to the characters in the key. It is an unusually shiny species and probably belongs in the subgenus Swnius s. str. I have received no record of its habits. 5. SUNIUS OBLITUS (Erichson) Lithocharis oblita ErtcuHson, 1840, p. 618.—BLACKWELDER, 1939a, p. 104. Medon (Lithocharis) oblita (Hrichson) BeERNHAUER and SCHUBERT, 1912, p. 242.—BLACKWELDER, 1939a, p. 104. Description.—Black or piceous, elytra rufotestaceous, abdomen cas- taneous or picescent. Head transverse, not emarginate behind, with angles moderately rounded; eyes large, at about their length from MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 265 base; labrum with a small median emargination but entirely without denticles; gular sutures moderately close, most approximate along middle, diverging at both ends; with rather coarse umbilicate punc- tures, very uneven in size, separated generally by less than their width by flat intervals, a little more sparse at center of vertex; with- out distinct ground sculpture. Pronotum as wide as head, nearly one- fifth wider than long, sides feebly converging and evenly rounded into base; punctation very similar to that of head; without a smooth mid- line but finely channeled along it posteriorly; without ground sculp- ture. LHlytra not distinctly punctate, surface very uneven and with sparse fine setigerous tubercles, very indistinct; without definite ground sculpture. Abdomen minutely punctate and pubescent, with indistinct ground sculpture. fale, seventh sternite unmodified, eighth with an abrupt rounded emargination. Female, sternites not modi- fied. Length, 234 to4 mm. Types locality —Colombia. Types.—Kither in the Hope Museum, Oxford, or Zoologische Mu- seum, Berlin. Records.—The following are the records known to me: St. Lucia: (Blackwelder station 220D). St. Vincent: (British Museum). Grenada: (Cameron collection; British Museum; United States National Museum). South America: Cotompia (Erichson, 1840; Bernhauer and Schubert, 1912). Specimens examined.—I have seen 158 examples in the British Mu- seum, 6 in the collection of Dr. Cameron, and 1 collected by me in April 1936. (These are all from the West Indies.) Remarks.—This species also is known to me only from the West Indies but seems to be properly associated with the Erichson name. It is distinguished by its smooth shining integuments and channeled pronotum. It belongs in the subgenus Sunius s. str. My specimen was found in rotting cocoa pods. 6. SUNIUS CURTULUS (Erichson) Lithocharis curiula Extcuson, 1840, p. 618.—Lene and MurcHtsr, 1914, p. 405.— BLACKWELDER, 1939a, p. 104. Medon (Lithocharis) curtulus (Erichson) BerNHAvER and SCHUBERT, 1912, p. 241.—BLACKWELDER, 1939a, p. 104. Description—Head black to rufopiceous, pronotum rufopiceous to rufous, elytra rufopiceous to rufous with apex sometimes paler, abdo- men castaneous. /Zead transverse, rather strongly emarginate behind but with angles broadly rounded; eyes large, at a trifle less than their length from base; labrum with a small triangular emargination at middle, entirely without teeth; gular sutures moderately close, most approximate along middle, diverging at both ends; with rather coarse 449008—42 18 266 BULLETIN 182, UNITED STATES NATIONAL MUSEUM umbilicate punctures, intervals flat, nearly absent on middle of vertex; without ground sculpture. Pronotwm scarcely narrower than head, one-ninth wider than long, sides feebly converging posteriorly; less coarsely umbilicately punctured than head, with an impunctate me- dian band in basal half but not channeled; without ground sculpture. Elytra not distinctly punctate, surface uneven and with sparse feeble setigerous tubercles; without ground sculpture. Abdomen exceed- ingly finely but not densely punctate, with fine long pubescence; with indistinct transverse ground sculpture. Male, seventh sternite not modified; eighth with a moderately deep rounded emargination. Female, sternites not modified. Length, 3 to 314 mm. Type locality—Colombia. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Grenada: (Cameron collection, British Museum; United States National Museum). Trinidad: (Bernhauer and Schubert, 1912; Leng and Mutchler, 1914), Tacarigua (Blackwelder station 107C). South America: VENEZUELA (Bernhauer and Schubert, 1912), Corompra (Erich- sou, 1840; Bernhauer and Schubert, 1912). Specimens examined.—I have seen 27 examples in the British Mu- seum, 7 in the collection of Dr. Cameron, and 1 collected by me in December 1935. (AJl these are from the West Indies.) Remarks.—Although I have seen no specimens from Colombia or Venezuela, I believe this identification to be correct. It is distin- guished by the absence of ground sculpture and the absence of a median pronotal channel. It belongs in the subgenus Sunius s. str. My specimen was taken from under moss and earth on a rock. 7. SUNIUS WEBER], new species Description.—Piceous, elytra piceocastaneous with apical sixth tes- taceous. /Zead a trifle broader than long (exclusive of the eyes), base distinctly emarginate only above the neck; eyes very large, separated from base by about half their length; sides rounded into base; labrum with a small median emargination but no denticles; gular sutures moderately separated, most approximate before middle, thence feebly diverging; with coarse, distinct, and crowded umbilicate punctures, the intervals convex and without sculpture. Pronotum about one- seventh wider than long, the sides moderately converging posteriorly and rounded into base; without any punctures or distinct ground sculp- ture but with fine rounded tuberculi separated by one to two times their diameter; middle stripe irregular but smooth, not elevated. Elytra about as long as wide, about one-eighth wider than pronotum 5 surface very uneven, without distinct punctures but with somewhat MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 267 indistinct tuberculi as on pronotum; without distinct ground sculp- ture. Abdomen with minute punctures obscured by irregular seulp- ture. Male, unknown. Female, sternites not modified. Length, 3 mm. Type locality —Trinidad, 11 miles north of Arima, elevation 1,800 feet. Types.—Holotype, female, in the Museum of Comparative Zoology, collected on March 28, 1935, by N. A. Weber. Records.—The following is the only record known to me: Trinidad: Arima (Weber, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species is unique in the coloring of the elytra but is immediately distinguished from all other West Indian species by possessing tuberculi on the pronotum and elytra instead of umbilicate punctures. It belongs in the subgenus Caloderma, of which it is our only respesentative. I have received no record of its habits. 8. SUNIUS DEBILICORNIS (Wollaston) Lithocharis debilicornis WouuLAston, 1857, p. 194. Lithocharis brevicornis ALLARD, 1857, p. 747, pl. 14, fig. 2. Lithocharis aegyptiaca MorTscHULSKY, 1858a, p. 644. Lithocharis pallida MoTscHULSKY, 1858a, p. 644. Lithocharis occulta WATERHOUSE, 1876, p. 108. Medon (Hypomedon) debdilicornis (Wollaston) Mursant and Rey, 1878, p, 164. Chloecharis rufula LYNCH, 1884, p. 259. Medon debdilicornis (Wollaston) FAUVEL, 1888, p. 25. Medon brevicornis (Allard) GANGLBAUER, 1895, p. 528. Medon aegyptiacus (Motschulsky) GANGLBAUER, 1895, p. 523. Medon (Chloecharis) debilicornis (Wollaston) DEvILLE, 1907, pp. 145, 148. Medon (Hypomedon) occultus (Waterhouse) BERNHAUER and ScHusEr?, 1912, p. 238. Medon (Hypomedon) aegyptiacus (Motschulsky) BERNHAUER and SCHUBERT, 1912, p. 238. Medon (Hypomedon) brevicornis (Allard) BERNHAUER and ScuuBert, 1912, p. 238. Medon (Hypomedon) rufulus (Lynch) BERNHAUVER and ScHuBeRt, 1912, p. 238. Chloecharis debilicornis (Wollaston) BERNHAUER and ScHuBERT, 1912, p. 238. Medon (Lithocharis) pallida (Motschulsky) BeRNHAUER and ScHUBERT, 1912, p. 248. Hypomedon debilicornis (Wollaston) LeENa and Mutcuter, 1914, p. 405. Huastenus pallidus Fiori, 1915, p. 10. Lithocharis rufulus (Lynch) CAMERON, 19381, p. 154. Medon (Hypomedon) pallidus (Fiori) ScHrrrPELrz, 1933, p. 1257. (Not Motschulsky, 1858.) Medon pallidum (Motschulsky) CAMERON, 1934, p. 79. Description.—Rutfotestaceous or fuscotestaceous, elytra testaceous. Head scarcely wider than long, feebly emarginate at middle of base, 268 BULLETIN 182, UNITED STATES NATIONAL MUSEUM sides arcuate but angles not completely rounded; eyes small, at nearly twice their length from base; labrum with a small emargination at middle, but without denticles; gular sutures moderately close in front, widely separated posteriorly ; with moderate umbilicate punctures, sep- arated by their diameter or less and obscured by dense ground sculp- ture. Pronotum scarcely wider than long, almost as wide as head; sides moderately convergent posteriorly, angles rounded; surface not impressed; midline impunctate but sculptured; with punctation and sculpture as on head. lytra normally, moderately coarsely punc- tured, without ground sculpture. Abdomen with fine asperate punc- tures, not dense; with distinct ground sculpture. fale, seventh and eighth sternites not modified. /’emale, not externally distinguishable from male. Length, 1384 to 244 mm. Type locality—Madeira, near Funchal. Of aegyptiacus, Egypt; of rufulus, Argentina, Chacabuco, Buenos Aires Province; of occul- tus, Rodriguez Island, Indian Ocean; of pallidus Fiori, “Itaha meridionale,” “nord di Cotrone (prov. di Catanzaro)”; of brevicornis, France, “aux environs de Rozoy-en-Brie”; of pallidus Motschulsky, “Indes orientales.” Types.—Unique type in the British Museum. Of aegyptiacus, presumably in the Zoologische Universitat Museum, Moscow; of rufulus, in the Museo Argentino Ciencias Naturales, Bernar Riva- davia, Buenos Aires; of occultus, in the British Museum; of pallidus Fiori, presumably in collection of A. Fiori; of brevicornis, disposition unknown; of pallidus Motschulsky, with aegyptiacus. Records.—The following are the records known to me: Cuba: (U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Jamaica: Milk River (Blackwelder station 415), Santa Cruz (Blackwelder sta- tion 421), Trinityville (Blackwelder station 428). St. Croix: (Blackwelder station 341). Antigua: (H. E. Box, as Blackwelder station 444C). Guadeloupe: (Fauvel, 1888; Leng and Mutchler, 1914). St. Lucia: (H. E. Box, as Blackwelder station 444B). St. Vincent: (Cameron collection). Grenada: (British Museum; Cameron collection). Trinidad: St. Augustine (British Museum). South America: (Gridelli, 1930), Aracrentrna (Lynch, 1884, as rufulus). Central America: Mexico (Gridelli, 1930). North America: SourH CaroLINA (Fauvel, 1878, 1888), Texas (Gridelli, 1980), Texas, Frorma (Casey collection, United States National Museum, as testacea). Europe: FrANCE (Fauvel, 1872, 1878, 1888; Allard, 1857; Mulsant and Rey, 1877; Portevin, 1929; Deville, 1907), Corsica, IrAty, Siciry (Fiori, 1915; Deville, 1926). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 269 Africa: (Gridelli, 1930), Maprrra (Wollaston, 1857; Fauvel, 1872, 1877, 1888), Capp VERDES (Fauvel, 1872, 1878, 1888; Ganglbauer, 1895), Eeypr (Fauvel, 1872, 1878, 1888; Ganglbauer, 1895), Tunis, Ropriguez ISLAND, CANARIES, Sr. HELENA, ALGERIA, AZORES (Waterhouse, 1876, 1879). Asia and Orient: Persta (Fauvel, 1878, 1888; Ganglbauer, 1895), Sram, Srnea- PORE, INDIA (Fauvel, 1872, 1888; Ganglbauer, 1895; Cameron, 1921, 1931), JAPAN (Fauvel, 1878, 1888; Ganglbauer, 1895). Australia: (Fauvel, 1878, 1888). Cceania: New CALEDONIA (Fauvel, 1888), HAwaAr (Sharp, 1908; Gridelli, 1930). Specimens examined.—l have seen from the West Indies, beside many from the Atlantic islands and the Indo-Oriental region and North America, 7 examples in the British Museum, 3 in Dr. Cameron’s collection, 2 in the Museum of Comparative Zoology, 4 in the United States National Museum, and 12 collected by me in 1935-87. Remarks.—The lengthy synonymy is due to failure to recognize that this is virtually a cosmopolitan species. Although so widespread, I know of no instance in which more than a few specimens have been taken at one time. It is the type of the subgenus Hypomedon. My specimens were taken in trash heaps in sugarcane fields and flying at dusk. 9. SUNIUS TERRESI, new specics Description.—Rufous, abdomen picescent. Head one-sixth wider than long, rather indefinitely emarginate behind, with angles mod- erately rounded; eyes large, at a little more than their length from base; labrum with two broad rounded lobes separated by a small emargination, without denticles; gular sutures moderately close, most approximate along middle, diverging at both ends; with mod- erately dense and small umbilicate punctures, not much sparser on disk, somewhat obscured by dense ground sculpture. Pronotum one- sixth wider than long, as wide as head exclusive of eyes; sides feebly converging and evenly rounded into base; without midline or chan- nels, but with a trace of midline formed of linearly arranged inter- vals; punctures similar to those of head but almost completely ob- scured by the dense strong ground sculpture. Hlytra with rather dense rounded tuberculi, impunctate and without ground sculpture. Abdomen minutely and densely punctate and pubescent. d/ale, un- known. Female, sternites not modified. Length, 4 mm. Type locality—Haiti, Diquini, on coastal plain between Port-au- Prince and Leogane. Types.—Holotype, female, U.S.N.M. No. 52405, collected by J. B. Terres. Records.—The following is the only record known to me: Hispaniola: Hair, Diquini (Terres, in United States National Museum). 270 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens exwamined.—lI have seen only the unique type. Remarks.—This species is distinguished by its dense ground sculp- ture and rather dense umbilicate punctures of head and pronotum and strongly tuberculate sculpture of elytra. It belongs in the subgenus Hypomedon. I have no record of its habits. XXXVI. Genus MEDON Stephens Medon STEPHENS, 1832, p. 273. Oxymedon Casry, 1905, p. 177. Subgenus Platymedon CAsry, 1889a, p. 184. Paramedon Casey, 1905, p. 166. Subgenus Medonodonta CaAsry, 1905, p. 176. Subgenus Tetramedon Casey, 1905, p. 178. Genotype.—Medon ruddii Stephens (monobasic). Of Oxymedon, O. rubrum Casey (monobasic) ; of Afedonodonta, M. alutacea Casey (monobasic); of Paramedon, P. arizonicum Casey (designated by Blackwelder, 1939) ; of Platymedon, P. laticollis Casey (monobasic) ; of Tetramedon, T. rufipenne Casey (monobasic). Diagnosis—Body elongate, moderately convex; punctures of head umbilicate; antennae posteriorly flexible, not strongly geniculate, basal segment not much elongate, first two not much larger than rest; labrum without median denticle, with one or two pairs of denticles near middle; fourth segment of maxillary palpus very small, subulate; gular sutures not at all united, never very distant but most approximate along middle or posteriorly; punctures of pronotum more or less tuberculate; prosternum longitudinally carinate along disk, prolonged posteriorly between coxae but not much dilated beneath them; hypomera prolonged in a short lobe partially behind the coxae; anterior coxal cavities open behind, confluent; front coxae very large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, third carinate at base, seventh unmodified in both sexes, eighth ster- nite not or feebly modified in male only. Remarks.—This name is used here in a restricted sense probably about equivalent to the subgenus Medon of Bernhauer and Schubert. It is not very abruptly distinguished from its close neighbors, but in this status permits a more careful grouping of its species. I have seen 89 examples of this genus, belonging to seven new species. KEY TO WEST INDIAN SPECIES OF MEDON i Head with denseveround sculptures 22.2 a ee ee ee ee 2 Head-without dense croundsculpturels== 2-3 =e eae 3 2; Head’ distinctly longer: than: broad— 2:22) !22 es eh ee ae 7. cordatum Head distinctly broader than, long=—._=+—-=-- 25+ 822 4 6. schwarzi 8. Midline of pronotum elevated, narrowly impunctate_-___----_------------ 4 Midline of pronotum not elevated ===. 2222-32 2oS Sen eee 5 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Diet 4. Head with many larger umbilicate punctures scattered among dense small (OT CS ea a eee ee 2. matasana Head without larger umbilicate punctures___________-_____________ 1. johni 5. Pronotum distinctly punctate; midline not deeply canaliculate____________ 6 Pronotum not distinctly punctate; midline deeply canaliculate__ 5. cubanum 6. Pronotum punctured as head, not very densely____-__________ 4, trinitatum Pronotum very densely punctate, more densely than head___ 3. dominicanum 1. MEDON JOHNI, new species Deseription.—Piceous, pronotum generally rufescent, elytra and base of abdomen castaneous, apex of abdomen testaceous. /Zead one- ninth wider than long, base not emarginate, sides rounded into base from eyes; eyes large, at a little less than their length from base; labrum with two distinct denticles near middle in a rather wide emargination; gular sutures moderately separated in front, becoming very close posteriorly; with dense but not large umbilicate punctures, separated by narrow convex intervals; without distinct ground sculp- ture. Pronotum as long as wide, one-tenth narrower than head; sides feebly converging posteriorly and rounded into base; disk impressed on either side along midline posteriorly; midline narrowly smooth and shining; with dense but feebly developed tuberculi, each bearing a minute punctule and hair at top, separated by less than their diameter by smooth shining intervals. Hlytra with moderate punctures ob- secured by submuricate or tuberculate rugosities; with long pubescence. Abdomen densely and finely punctate, more sparsely apically. Male, seventh and eighth sternites not modified. Female, externally indis- tinguishable from male. Length, 4 mm. Type locality—sSt. Lucia, one-half mile northwest of Duboulay Estate on road to Soufriére, district of St. Remy. Types.—Holotype and 32 paratypes, U.S.N.M. No. 52408, collected by me in April and May 1936; two paratypes from Dominica. Two paratypes have’ been deposited in the British Museum, two in Dr. Cameron’s collection, and two in my own collection. Records.—The following are the records known to me: Dominica: (Blackwelder station 237). St. Lucia: (Blackwelder station 207B, 220D, 221, 224, 226, 227, 280, 231). Specimens examined.—l have seen only the 40 types. Remarks.—I find considerable variation in the labrum of this species. In several there are two hairs arising between the denticles which seem to mark two minute additional denticles; in some the edges of the emargination are angular and almost denticlelike; and in some the two teeth are so reduced as to be rather indistinct. This species belongs in the subgenus J/edon s. str. I have collected this species from cut banana stalks and from rotting cocoa pods. 272 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. MEDON MATASANA, new species Description.—Piceocastaneous, humeri and sides of abdomen some- what paler. Head exclusive of eyes about one-sixth wider than long, base slightly emarginate over the neck, sides very feebly converging behind, the angles narrowly rounded; eyes large, at about their length from base; labrum with a small but abrupt emargination deeper than wide, without denticles; gular sutures fairly close and nearly parallel from near the front; with dense but rather small umbilicate punctures interspersed with larger ones; the intervals flat and without distinct ground sculpture. Pronotum about one-sixth wider than long, about one-tenth narrower than head; sides moderately converging and rounded into base; disk feebly impressed along midline posteriorly ; midline narrowly and indefinitely smooth and shining, vaguely canaliculate posteriorly; with very dense and irregular umbilicate punctures, the intervals not wide enough to be flattened; without dis- tinct ground sculpture. H/ytra with rather dense submuricate punc- tures obscured by surface rugosity; with rather long pubescence. Abdomen finely and moderately densely punctate; with indistinct ground sculpture. A/ale, eighth sternite with a large triangular emargination, twice as broad as deep, with all angles rounded. Female, sternites unmodified. Length, 4 to 414 mm. Type locality—Dominican Republic, San José de las Matas, elevation 1,000 to 2,000 feet. Types.—Holotype and seven paratypes in the Museum of Com- parative Zoology; five paratypes in the United States National Mu- seum (No. 52511) ; collected in June 1938 by Dr. P. J. Darlington. fecords.—The following are the records known to me: Hispaniola: DoMINICAN REPUBLIC, San José de las Matas (Darlington, in M.C.Z. and U.S.N.M.), Mount Quita-Espuela (Darlington, in M.C.Z. and U.S.N.M.), Santiago de Jos Caballeros (Darlington, in M.C.Z.),. Villa Altagracia (Darlington, in M.C.Z.), Sanchez (Darlington, in M.C.Z.). Specimens ewamined.—I have seen only the 13 types. Remarks.—This species is distinct by its labral emargination, the punctation of the pronotum and elytra, and the male characters. It is larger than any of our species except dominicanum. It apparently belongs in the subgenus Afedon s. str. I have received no record of its habits. 3. MEDON DOMINICANUM, new species Description—Head black, pronotum piceous, elytra and abdomen castaneous with humeral areas of elytra indefinitely paler. Head about one-seventh wider than long, base somewhat emarginate espe- cially above the neck, sides parallel but moderately rounded into base ; eyes large, at a little more than their length from base; labrum with MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 218 a small and abrupt emargination, deeper than wide, the apex rectangu- lar, without denticles; gular sutures narrowly separated, feebly con- verging posteriorly; with fine and dense but very distinct umbilicate punctures interspersed with a few much larger ones, the intervals narrow but flat and with ground sculpture at sides basally. Pronotum about one-sixth wider than long, only a trifle narrower than head; sides rather strongly converging and rounded into base; disk not impressed along midline; midline not smooth or canaliculate; with dense umbilicate punctures of two sizes as on head; without distinct ground sculpture. //lytra with rather dense submuricate punctures obscured by coarse rugosity of the surface; with long pubescence. Abdomen very finely and densely punctulate, with indistinct ground sculpture. Male, sixth sternite with anterior plica extended at middle to make a narrow elevated median ridge which projects as a blunt process at posterior margin; seventh sternite transversely impressed and broadly emarginate; eighth with a large excision, deeper than wide, wider just inside the apical angles. Female, sternites unmodified. Length, 5 to 6 mm. Type locality —Dominican Republic, Sanchez. Types.—Holotype, male, and five paratypes in the Museum of Com- parative Zoology; five paratypes in the United States National Mu- seum (No. 52512), collected in July 1938 by Dr. P. J. Darlington. Records.—The following are the records known to me: Hispaniola: Hartt, Ntang Lachaux (Darlington, in M.C.Z. and U.S.N.M.) ; DoMINICAN Repusiic, Sanchez (Darlington in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 11 types. Remarks.—This is the largest Medon known to me. It is very dis- tinct by the male characters, in which it differs greatly from J. matasana, which is its closest neighbor geographically as well as zoologically. It probably belongs in the subgenus Medon s. str. I have received no record of its habits. 4, MEDON TRINITATUM, new species Description.—Piceous, elytra and abdomen piceocastaneous, ely- tron with a triangular pale spot at outer apical corner extending to suture and to middle of side. Mead one-fifth wider than long, base emarginate only above the neck, sides parallel and rather nar- rowly rounded into base; eyes large, at about their length from base; labrum with a small abrupt emargination, scarcely as deep as wide, without denticles; gular sutures obsolescent, feebly divergent from near front; with moderately coarse umbilicate punctures sepa- rated by one-half to once their diameter; without ground sculpture. Pronotum one-eighth wider than long, as wide as head; sides moder- ately converging behind and rounded into base; disk not impressed 274 BULLETIN 182, UNITED STATES NATIONAL MUSEUM along midline; with only a trace of smooth midline; with punctures as on head but sometimes a little denser; without ground sculpture. Elytra with punctures irregular, feebly submuricate, surface very uneven though not actually rugulose; with long pubescence. Ab- domen with minute and rather indistinct punctures and rather vague ground sculpture. A/ale, eighth sternite with a broad shallow emar- gination. Female, sternites not modified. Length, 3 to 334 mm. Type locality —Trinidad, Tunapuna. Types.—Holotype, male, and seven paratypes in the Museum of Comparative Zoology; four paratypes in the United States National Museum (No. 52513) ; collected in June to September 1935 by N. A. Weber. (Paratypes partly from the localities listed below.) Records.—The following are the records known to me: Trinidad: Tunapuna (Weber, in M.C.Z. and U.S.N.M.), St. Augustine (Weber, in M.C.Z. and U.S.N.M.), Nariva Swamp (Weber, in M.C.Z.), Galeota Point (Weber, in M.C.Z.), Macqueripe Bay (Weber, in M.C.Z.), Balandra Bay (M.C.Z.). Specimens examined.—lI have seen only the 12 types. Remarks.—This species is distinct from the other West Indian species by the color pattern of the elytra. It probably belongs in the subgenus Medon s. str. I have received no record of its habits. 5. MEDON CUBANUM, new species Description—Rufous, elytra faintly picescent. Head a trifle broader than long, the base not at all emarginate, the sides slightly expanded behind the eyes and moderately rounded into base; eyes small, at almost three times their length from base; labrum with a small median emargination about as wide as deep, without denticles; gular sutures rather close, diverging at both ends; with rather small but dense and distinctly umbilicate punctures usually separated by half or less of their diameter; without ground sculpture. Pronotum as wide as long, a trifle narrower than head; sides moderately con- verging and rounded into base; rather coarsely canaliculate along midline throughout; with some depression along midline posteriorly ; setigerous punctures apparently umbilicate but almost completely obsolete, not dense; without ground sculpture. //ytra with moder- ately fine and dense punctures obsolescent, separated indefinitely by less than their diameter; with long pubescence. Abdomen with strongly umbilicate punctures fairly dense, without distinct ground sculpture. Afale, eighth sternite with an abrupt rounded emargina- tion from one-half deeper than wide to about as wide as deep. Fe- male, unknown. Length, 314 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 210 Type locality —Cuba, Loma del Gato, Cobre Range, Oriente Prov- ince, elevation about 3,000 feet. Types.—Holotype and one paratype, males, in the Museum of Comparative Zoology; one paratype, male, in the United States National Museum (No. 52514); collected on July 3-7, 1936, by Dr. P. J. Darlington. Records.—The following are the records known to me: Cuba: Loma del Gato (Darlington, in M.C.Z. and U.S.N.M.), Pico Turauino (Darlington, in M.C.Z.). Specimens examined.—I have seen only the three types. Remarks.—This species is distinguished by its small eyes, by the obsolete punctuation of the pronotum, and by the male characters. It is also unique in the enlargement of the head behind the eyes. It probably belongs in the subgenus Medon s. str. I have received no record of its habits. 6. MEDON SCHWARZI, new species Scioporus caribaeus Bireria, MS. Description —Rufotestaceous. Head one-seventh wider than long, rather indefinitely emarginate behind, with angles moderately rounded; eyes small, at about twice their length from base; labrum with a small median emargination and a pair of more feeble emargi- nations between this and the sides, without denticles; gular sutures moderately close, most approximate along middle, diverging at both ends; with very sparse umbilicate punctures separated by one to three times their diameter, with dense scaly ground sculpture. Pronotum almost as wide as head, as long as wide; sides feebly con- verging and evenly rounded into base; with an impunctate but sculptured median area, not channelled; with punctures and sculp- ture as on head. Llytra not distinctly punctate, surface uneven and with indistinct setigerous tubercles moderately dense; without ground sculpture. Abdomen very minutely punctured and _ pubescent; ground sculpture indistinct. J/ale, eighth sternite with a small rounded emargination about three times as wide as long. Female, sternites not modified. Length, 4144 mm. Type locality —Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, female, U.S.N.M. No. 52404, collected by E. A. Schwarz on February 26 of 1902, 1903, or 1904; one paratype in the United States National Museum and two in the Museum of Comparative Zoology, taken in August 1932 and 1934 at Soledad, near Cienfuegos, Cuba, by B. B. Leavitt and P. J. Darlington. (Two “cotypes” of S. caribaeus are in the United States National Museum, through the kindness of Sr. Bierig.) 276 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records —The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Rangel (Bierig, in U.S.N.M. as caribacus). Specimens examined.—I have seen the four types as above, the two “cotypes” of caribaeus, and three other specimens from the Museum of Comparative Zoology. Remarks.—This species is distinguished by its dense ground sculp- ture, its sparse umbilicate punctures, and other minor characters. It belongs in the subgenus Platymedon. One of Dr. Darlington’s specimens was found under a stone in company with ants of the species Odontomachus haematoda (Linnaeus). 7. MEDON CORDATUM, new species Description —Rufocastaneous. Head one-sixth longer than wide, not emarginate behind; sides parallel, nearly straight, moderately broadly rounded into base; eyes small, about three and a half times their length from base; labrum with a triangular emargination nearly twice as broad as deep with the outer angles rounded, without denticles; gular sutures approximate and parallel from near base to near apex; with somewhat irregular but moderate umbilicate punc- tures separated by one to two times their diameter; with rather feeble scaly ground sculpture. Pronotwm as wide as head and as long as broad; sides rather strongly converging behind from anterior quarter and rounded into base; midline feebly canaliculate along middle; punctured and sculptured as head. #lytra with punctures about as coarse as on pronotum but a little denser; surface uneven but not distinctly sculptured. Abdomen with fine punctures not dense and with a few coarse punctures on each segment; without distinct ground sculpture. Afale, eighth sternite with a broad tri- angular emargination, three times as wide as deep. Female, un- known. Length, 4144 mm. Type locality—Cuba, Buenos Aires, Trinidad Mountains, eleva- tion 2,500 to 3,500 feet. Types.—Holotype, male, in the Museum of Comparative Zoology, collected on May 8-14, 1936, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Cuba: Buenos Aires (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species is very distinct because of the elongate parallel head and the ground sculpture of the head and pronotum. It belongs in the subgenus Platymedon. I have received no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 214 XXXVII. Genus ORUS Casey Orus CASEY, 1884, p. 136. Subgenus Leucorus Casry, 1905, pp. 191, 192. Subgenus Pycnorus Casey, 1905, p. 194. Genotypes—Orus punctatus Casey (designated by Blackwelder, 1939). Of Leucorus, L. rubens Casey (designated by Blackwelder, 1939) ; of Pyenorus, Scopaeus dentiger LeConte=Pycnorus dentiger (LeConte) (designated by Blackwelder, 1939). Diagnosis —Body slender, strongly constricted; punctures of head not umbilicate; eyes normal; antennae posteriorly flexile, strongly geniculate, basal segment not very elongate, first and second segments not larger than rest; labrum not very deeply emarginate; fourth seg- ment of maxillary palpus subulate; gular sutures moderately separate, not strongly converging at either end; neck more than one-fourth as wide as head; pronotum distinctly longer than wide; prosternum not expanded under coxae laterally; hypomera feebly lobed behind the coxae; anterior coxal cavities open behind; front coxae large, exserted ; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, fourth and fifth sometimes with a transverse fold, seventh modified, eighth emarginate; basal half of front tibia with a concavity lined with diagonal ctenidia; apex of posterior tibia with a ctenidium only on inner edge. Remarks.—This genus is very similar to Scopaeus, from which it is distinguished principally by the width of the neck. The wide separa- tion of these two genera in the key (see Blackwelder, 1939) is mis- leading and is corrected in the generic arrangement. I have examined 51 specimens of this genus and describe two new species to receive them. The Scopaeus umbra of Sharp is distinct from either of these and is not known to me from the West Indies. KEY TO WEST INDIAN SPECIES OF ORUS 1. Elytra with distinct punctures, without ground sculpture____-____ 1. volans Elytra without distinct punctures, with dense ground sculpture. 2. cameroni 1. ORUS VOLANS, new species Description —Testaceous. Head distinctly emarginate behind, the angles rounded ; the eyes separated from base by more than three times their length; labrum bidenticulate, with a minute prominence at each side; gular sutures moderately separate, almost parallel; without distinct punctation but with dense but feeble ground sculpture. Pvo- notum one-fourth longer than wide, angles rounded, sides not at all emarginate in front; midline slightly elevated posteriorly ; with rather distinct punctures but obscured by fine ground sculpture. /lytra somewhat coriaceous but with distinct and rather large punctures; without distinct ground sculpture. Male unknown. Female, eighth 278 BULLETIN 182, UNITED STATES NATIONAL MUSEUM sternite prominently rounded; eighth tergite rounded. Length, 134 to 214 mm. Type locality —Jamaica, between Toll Gate and Newport by way of Milk River and Figurary Gully. Types.—Holotype and three paratypes, females, U.S.N.M. No. 52422, collected by Chapin and Blackwelder on February 25, 1937. Records—The following are the records known to me: Jamaica: Milk River (Blackwelder station 415), Spanish Town (Blackwelder station 377). Specimens ewamined.—I have seen only the three types and one other example. Remarks.—The sculpture of the elytra is the chief character sep- arating this from cameroni. It belongs in the subgenus Leucorus. The types were taken flying at dusk. 2. ORUS CAMERONI, new species Description —Castaneous. Head distinctly emarginate behind, the angles rounded; eyes separated from base by more than twice their length; labrum minutely bidenticulate; gular sutures moderately separated, closest along middle; very finely and densely punctulate, the punctulae not very distinct from the ground sculpture. Prono- tum nearly one-fourth longer than wide, angles rounded; sides not emarginate in front; midline scarcely evident in any part; with very minute punctulae obscured by feeble ground sculpture. L/ytra not distinctly punctate, with dense and fine subasperately tuberculate sculpture. Male, fourth and fifth sternites with a transverse fold; seventh with a longitudinal impression with a few rounded spicules scattered along it; eighth sternite with a narrow parallel incision, four times as deep as wide, the angles rounded. Female, eighth stern- ite prominently rounded; eighth tergite rounded. Length, 114 to 2mm. Type locality—Jamaica, between Bath and Lower White Horses by way of Sunning Hill, Trinityville, and Cedar Hill. Types.—Holotype, male, and 18 paratypes, U.S.N.M. No. 52420, col- lected by Chapin and Blackwelder on February 28, 1937. One para- type has been deposited in the collection of Dr. Cameron, and one in the British Museum. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in United States National Museum), Baragua (Chris- tenson, in United States National Museum). Jamaica: (Cameron collection), Trinityville (Blackwelder station 428), Montego 3ay (Blackwelder station 407), Spanish Town (Blackwelder station 377), Milk River (Blackwelder station 415), Moneague (Blackwelder station 375), Santa Cruz (Blackwelder station 421), Fern Gully (Blackwelder station 374). Hispaniola: Hattr (Cameron collection). Grenada: (British Museum; Cameron collection). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 279 Specimens examined.—I have seen 12 examples in the British Mu- seum and 13 in Dr. Cameron’s collection under the name wmbra Sharp. Six more are in the United States National Museum, and 16 were collected by Chapin and Blackwelder in February 1937. Remarks.—This species is apparently widely distributed although imperfectly collected. It belongs in the subgenus Leucorus. The types were collected from under seaweed and flying at dusk. XXXVIII. Genus SCOPAEUS Erichson Scopaeus ErtcHson, 1840, p. 604. Polyodontus Sotrer, 1849, p. 310 . Scoponeus MorscHuLsky, 1858a, p. 641. Leptorus Casry, 1886b, pp. 217, 220. Scoponaeus HICHELBAUM, 1909, p. 150 (emendation). Pseudorus Casey, 1910, p. 190. Scopoeoneus CAMERON, 1934, p. 79 (misspelling). Scopaeoneus CAMERON, 1940, p. 89 (misspelling). Subgenus Scopaeodera CAsrEy, 1886b, p. 217. Subgenus Scopaeopsis CAsry, 1905, p. 191. Subgenus Scopacoma Casey, 1905, p. 191. Genotypes.—Paederus laevigatus Gyllenhal=Scopaecus laevigatus (Gyllenhal) (designated by Duponchel, 1841; designation of 8. didy- mus Krichson by Blackwelder, 1939, in error). Of Polyodontus, P. angustatus Solier (monobasic) ; of Scoponeus, S. testaceus Motschul- sky (designated by Blackwelder, 1939) ; of Leptorus, Scopaeus exiguus Erichson=Leptorus exiguus (Erichson) (designated by Blackwelder, 1939; the designation of texanus Casey by Tottenham, 1939 was subsequent to the above and therefore invalid); of Pseudorus, P. prolixipennis Casey (designated by Blackwelder, 1939); of Sco- pacodera, E'chiaster nitidus LeConte = Scopaeodera nitida (LeConte) (designated by Blackwelder, 1939) ; of Scopacopsis, Echiaster opacus LeConte = Scopacopsis opaca (LeConte) (designated by Blackwelder, 1939) ; of Scopaecoma, Scopaeus rotundiceps Casey = Scopaeoma ro- tundiceps (Casey) (designated by Blackwelder, 1939). Diagnosis —Body generally slender and strongly constricted; head finely punctate; vertex not carinate in males; antennae posteriorly flexile, not strongly geniculate, basal segment not very elongate, first and second not much larger than rest; labrum bidentate or quadri- dentate, not very deeply emarginate; fourth segment of maxillary palpus subulate; gular sutures always separate; neck one-fifth to one- eighth as wide as head; prosternum somewhat dilated under the coxae but widely separated from hypomera; hypomera somewhat lobed behind the coxae; front coxal cavities separate, open behind; front coxae large, exserted ; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; fourth and fifth sternites of male sometimes with a transverse fold or 280 BULLETIN 182, UNITED STATES NATIONAL MUSEUM foveae; eighth sternite of male emarginate; apex of posterior tibiae with a ctenidium on inner edge only; tarsi 5-segmented; fourth seg- ment not dilated. Remarks.—This genus is unusually well represented in our fauna, and about half of the species are new. The classification of the sub- genera is still in an unsatisfactory state, although I have attempted an arrangement in a previous paper. The records of Scopaeus mollis Sharp from Jamaica in Gowdey, 1926, and of Scopaeus illustris Fauvel from Cuba by Leng and Mutch- ler, 1914, are probably misidentifications. Until the specimens can be reexamined, they must be regarded as doubtful records. Of this genus from the West Indies I have seen 179 examples in the British Museum, 53 in the collection of Dr. Cameron, 32 in the United States National Museum, 41 from the Museum of Comparative Zoology, and 116 collected by me in 1935-37. These belong to 17 species of which 9 are new. One other species is known to me only from the original description. KEY TO WEST INDIAN SPECIES OF SCOPAEUS 1. Head, pronotum, and elytra shining, scarcely punctate or sculptured____-~ 2 Head, pronotum, and elytra more or less densely punctate or sculptured__ 3 2. Head and elytra with obsolete sculpture_--___-____._____________ 18. chapini Head and elytra smooth and shining, as pronotum____--~__ 17. pulchellus oe, Head*truncate or emarsinate’ behind==—t =" --- 2 eee 4 Head “distinctly: rounded behind2 22512 a ee eee 17 4. Punctures of head “close, rugulose, umbilicate’____-_________- 14. auripilis Punctures of head fine and dense, not umbilicate, or with dense fine ground sculpture only_------ pe Ses ei a oR Oe ts a ee ee 5 5. Disk of pronotum with distinct individual round tuberculi__-------__--~_ 6 Disk of pronotum with normal fine punctures___-___-_-_____-___-_____-_- 7 6.) Apical! margin ‘of ‘elytra paler: -2=2h2 822 ee Sees eee 12. arena Elytra unicolorous <= 223-5222 2 se eee elias 13. darlingtoni 7. Pronotum usually parallel, front angles broadly rounded, sides in front not distinctly: .emarginate.2s— 222+ =. ee a 8 Pronotum more or less narrowed behind, front angles distinct, sides in front distinetly. emarginates #27!) #28 Set te Se ae oe ee ee 15 8. Pronotum very much depressed; color entirely testaceocastaneous or rufotestaceouss 2% {= 2Sh.s et eee a eee 11. angusticollis Pronotum not very much depressed; color piceocastaneous to testaceous__ 9 9. Fourth (and fifth) sternites of male with a transverse fold_---_----__~- 10 Mourth and fitth: sternites without fold222. 22-22 == 2-2 ee 13 10. Eyes at about twice their length from: base_.----___-___~---_-__--+_-_- re Eyes at about three times their length from base___----_---_--~- 5. rulomus 11.,, Fold. of fourth’ sternite complete across disk22+-1222) 2. s)=ee_ ee ee 12 Fold of fourth sternite divided at middle__-_-________-_____ 4. fasciatellus 12. Seventh sternite of male broadly and feebly emarginate, eighth deeply CINATPIN ATO ee ne ee ee 3. potamus Seventh sternite of male not at all emarginate, eighth feebly emarginate. 2. rivularis MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 281 1sourchisterniteof male without carinals fo Sees ee see ee eae 14 Fourth sternite of male with a small posterior carina______~ 1. dominicanus 14. Pronotum densely and distinctly punctate_________-____-___~_ 8. pygmaeus Pronotum sparsely and rather indistinctly punctate__--.___----_- 9. filum 15. Punctures of pronotum very fine and dense, without evident individual CLTATE LO re are ae Pe eA De Ea ES ee ee ee et 16 Punctures of pronotum not very fine though dense, diameter of each clearly VAS IID) Cesare es BE oe we Bh ee ie ee a Se ee eS 10. cupiens IGA pexcOLelybranacrowlypalers- oe fe 7. marginatus Kiytra’ wnicolorouss -2=—2- 25s Ue ee a ee ee 6. antennalis Meellytrartinely;puUnChates seas ee Sse ee ee eo ee 16. boxi Hiviravdistinetly, tuberculates222 5 os 2s ess ee Santas 15. rambouseki 1. SCOPAEUS DOMINICANUS, new species Description —Rufocastaneous. Head distinctly emarginate behind, the angles rather narrowly rounded; the eyes separated from base by about three times their length; labrum with two slender denticles near middle and a pair of small lateral ones; gular sutures narrowly sepa- vated in front and feebly diverging posteriorly; with very fine and dense setigerous punctures but without distinct ground sculpture. Pronotum about one-fourth longer than wide, anterior angles fairly broadly rounded; sides in front scarcely emarginate, nearly straight to rounded basal angles; without trace of midline except as posterior ele- vation; punctures and sculpture as on head but a little less distinct. Elytra very finely and densely tuberculate or rugulose-sculptured. Male, fourth and fifth sternites without fold; fourth with a small but abrupt keel at middle of posterior edge; eighth truncate and with a small triangular incision about as deep as wide. Yemale, eighth sternite prominently rounded; eighth tergite rounded. Length, 214 mm. Type locality—Dominican Republic, Sanchez. Types.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology; one paratype, female, in the United States National Museum (No. 52415); collected in July 1938 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Hispaniola: DoMINIcAN Repusuio, Sanchez (Darlington, in M.C.%, and U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—TVhis species is distinct from all the other West Indian Scopaeus by the carinate fourth sternite of the male. It belongs in the subgenus Scopaeus s. str. I have received no record of its habits. 2. SCOPAEUS RIVULARIS, new species Description.—Castaneous, frequently somewhat picescent in part, apical borders of elytra always paler than rest. Head very feebly 449008—42 19 282 BULLETIN 182, UNITED STATES NATIONAL MUSEUM emarginate behind, the angles rather narrowly rounded; eyes sepa- rated from base by a little over twice their length; labrum with two small median denticles; gular sutures rather narrowly separated in front, feebly diverging posteriorly; finely and densely punctured, the punctures obscured by indistinct ground sculpture. Pronotum one-fourth longer than wide, anterior angles very broadly rounded ; sides in front not emarginate, straight to rounded posterior angles; without trace of midline; punctures and sculpture similar to that of head. Flytra with sculpture similar to that of head but deeper and a little less dense. Male, fourth and fifth sternite with a transverse fold, seventh not emarginate; eighth sternite with a very feeble rounded emargination, not abrupt. Female, eighth sternite promi- nently rounded; eighth tergite with a slightly arcuate prominence. Length, 2 to 214 mm. Type locality —Jamaica, 2 miles northeast of Gordon Town on road to Mavis Bank. T'ypes.—Holotype, male, and four paratypes, U.S.N.M. No, 52419, collected by Chapin and Blackwelder in February 1937. Records—The following are the records known to me: Jamaica: Gordon Town (Blackwelder station 368B), Trinityville (Blackwelder station 428). Specimens examined.—I have seen only the five types. Remarks.—This species is superficially rather similar to several of its neighbors, but is distinguished by the feeble male characters. It belongs in the subgenus Scopaeus s. str. 3. SCOPAEUS POTAMUS, new species Description —Rufopiceous, the head generaliy hghter than the pro- notum, elytra paler along apex. Head distinctly emarginate behind, the angles rather narrowly rounded; the eyes separated from the base by about twice their length; labrum with median denticles rather enlarged at base, with an indistinct outer pair; gular sutures mod- erately narrowly separated in front of middle and feebly diverging posteriorly; without distinct punctures but with dense fine ground sculpture. Pronotwm one-fourth longer than wide; anterior angles strongly rounded, sides scarcely emarginate in front, feebly arcuate to posterior angles; with trace of impressed midline; with ground sculpture as on head but less distinct. H/ytra not distinctly punc- tate but densely subasperately or tuberculately sculptured. ale, fourth and fifth sternites with a transverse fold, seventh broadly feebly emarginate; eighth sternite with a rather large and broad emargination, one-third as deep as wide. Female, eighth sternite rounded; eighth tergite rounded. Length, 2 to 214 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 283 Type locality —St. Lucia, Roseau River, at elevation of about 200 feet, about 6 miles above its mouth. Types Holotype, male, and three paratypes, U.S.N.M. No. 52417, collected by me on March 29, 1936. Records.—The following is the only record known to me: €t. Lucia: (Blackwelder station 209A). Specimens examined —I have seen only the four types. Remarks.—This species is readily distinguished by the male charac- ters. It belongs in the subgenus Scopaeus s. str. The types were collected in the sand at the edge of a stream in the forest. 4. SCOPAEUS FASCIATELLUS Erichson Scopaeus fasciatellus ExicHson, 1840, p. 609.—BERNHAUER and ScHusert, 1912, p. 242.—LreNe and Murcnuter, 1914, p. 405.—Wotcort, 1924, p. 78; 1936, p. 197.— BLACK WELDER, 1989a, p. 106. Description.—Head and pronotum rufotestaceous, elytra and abdo- men cinnamomeous. /fead feebly emarginate behind, the angles mod- erately rounded; eyes separated from base by over twice their length ; labrum with two median denticles and two lateral ones; gular su- tures narrowly separated and virtually parallel throughout; very finely and densely punctate but with smooth shining intervals. Pronotum one-fifth longer than wide, anterior angles rather broadly rounded; sides in front not emarginate, scarcely arcuate to base; with narrow smooth midline basally; rather indistinctly punctate with surface uneven but shining. Z7ytra not distinctly punctate, surface almost asperately sculptured, pubescent, without other dense ground sculpture. Male, fourth and fifth sternite with a transverse fold widely divided at middle to form two foveae on each segment; seventh sternite flattened at middle, eighth very feebly broadly emar- ginate. Female (from Erichson), eighth sternite feebly triangularly produced. Length. 214 mm. Type locality —Puerto Rico. Types.—Kither in Hope Museum, Oxford, or the Zoologische Mu- seum, Berlin. Records.—The following are the records known to me: Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914; Wolcott, 1924, 1936), Ciales (Blackwelder station 63). Specimens examined. —I\ have seen only one example collected by me on October 12, 1935. Remarks.—The specimen on which this description is based agrees with the original description of Erichson in all essential characters. I have little doubt that the identification is correct. The shape of 284 BULLETIN 182, UNITED STATES NATIONAL MUSEUM the head and the male characters indicate that this species belongs in the subgenus Scopacus s. str. My specimen was taken from under pebbles or flying at the bank of a river. 5. SCOPAEUS RULOMUS, new species Description.—Testaceous with abdomen somewhat darker. Head feebly emarginate behind, the angles moderately rounded; eyes very small, separated from base by nearly three times their length; labrum with two small denticles at middle, and with a very indistinct smaller denticle at each side, the median pair enlarged at base; gular sutures moderately separated in front and feebly diverging posteriorly; punctures fine and irregular, separated by their diameter by flat shining intervals. Pronotwm about one-third longer than wide, an- terior angles moderately narrowly rounded; sides in front feebly emarginate, nearly straight to the rounded posterior angles; midline very vague; punctures a little sparser and less distinct than on head; surface similarly shining. /lytra not distinctly punctate but with very irregular and almost asperate sculpture, though surface is shin- ing. Afale, fourth and fifth sternites with a transverse fold widely interrupted at middle to form two separated pairs of foveae; eighth sternite truncate and with a broad and shallow acuminate emargina- tion. Female, eighth sternite rounded; eighth tergite unmodified. Length, 134 mm. Type locality —Jamaica, between Bath and Lower White Horses by way of Sunning Hill, Trinityville, and Cedar Hill. Types.—Holotype, male, and one paratype, female, U.S.N.M. No. 52491, ‘collected by Chapin and Blackwelder on February 28, 1937, and February 2, 1937. Records.—TVhe following are the records known to me: Jamaica: Trinityville (Blackwelder station 428), Fern Gully (Blackwelder sta- tion 374). Specimens examined.—t\ have seen only the two types. Remarks.—TVhis is a very small and fragile species which belongs in the subgenus Scopaeus s. str. It is chiefly remarkable for the creat distance between the eyes and the base of the head. The types were both taken flying at dusk. 6. SCOPAEUS ANTENNALIS Cameron Scopacus antennalis CAMERON, 1913b, p. 349.—LenG and MutcHuer, 1917, p. 199. Scopaeus antennatus SCHEERPELTZ, 1938, p. 1265 (misspelling). Scopaeus bicoloricornis Brertc, MS. Description.—Piceous to piceocastaneous or castaneous. Head slightly emarginate behind, angles moderately rounded; labrum with two denticles separated by a small rounded emargination, and with ~- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 285 two additional minute denticles laterally; the median denticles are somewhat enlarged at base; gular sutures moderately separated, fee- bly diverging posteriorly; with minute punctures so dense as to appear like ground sculpture, but surface still shining. Pronotum one-ninth longer than wide, anterior angles narrowly rounded, sides feebly emarginate in front, moderately arcuate posteriorly; with scarcely a trace of midline; not very distinctly punctate but with sculpture forming irregular punctiform depressions, without addi- tional ground sculpture. /lytra not punctured but with distinct and strong tuberculi separated by less than their diameter; without ground sculpture except basally along suture. A/a/e, fourth and fifth sternites with a transverse fold, seventh feebly broadly emarginate; eighth sternite with a deep triangular emargination, apex narrowly rounded, about two-thirds as deep as wide, and with outer angles rounded. Female, eighth sternite rounded; eighth tergite rounded. Length, 214 to 3 mm. Type locality —Haiti. Types.—In the collection of Dr. Cameron. Records.—The following are the records known to me: Cuba: (Bierig and Cameron collection, as bicoloricornis), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Aguadores in Oriente (Darlington, in M.C.Z. and U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Baragua (Christenson, in U.S.N.M.). Hispaniola: Hairr (Cameron, 1913; Leng and Mutchler, 1917; Scheerpeltz, 1933), Desbarriere (Darlington, in M.C.Z.) ; Dominican REPUBLIC, Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.), Constanza (Darlington, in M.C.Z.), SAnchez (Darlington, in M.C.Z. and U.S.N.M.), San José de las Matas (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: Jayuya (Oakley, in U.S.N.M.). St. Croix: (Blackwelder station 341). Guadeloupe: Basse Terre (Blackwelder station 79B). St. Lucia: (Biackwelder station 209A). Specimens examined.—\ have seen the unique type in Dr. Cam- eron’s collection, 16 examples in the United States National Museum, 19 from the Museum of Comparative Zoology, and 12 collected by me in 1935-37. ‘Two additional specimens are in Dr. Cameron’s col- lection under this name but labeled also as cotypes of bicoloricornis Bierig (MS. name). Remarks. —I did not examine the two “cotypes” from Cuba. The above description was taken entirely from the specimens in the United States National Museum. Dr. Cameron did not mention the folds of the fourth and fifth sternites, and I failed to examine the type for this character. If they are not present, then my specimens may be a distinct species. The shape of the head and the male char- acters place this species in the subgenus Scopaeus s. str. 286 BULLETIN 182, UNITED STATES NATIONAL MUSEUM My specimens were taken “on cucumber,” under rotting mango fragments, under sand and gravel along a stream, and flying at dusk. The Baragua specimen was taken “in soil of sugarcane fields.” 7. SCOPAEUS MARGINATUS Cameron Scopaeus marginaits CAMERON, 1913b, p. 3849.—LeENG and MUTCHLER, 1917, p. 199.—SCHEERPELTZ, 1933, p. 1267.—Brackwevpre, 1939a, p. 106. Description.—Piceous, the apex of the elytra narrowly rufotesta- ceous. Zead slightly emarginate behind, the angles rounded; labrum with two moderate median denticles separated by a rounded notch, outer angles prominently angled, almost denticulate; gular sutures moderately separated in front, divergent posteriorly; very finely but discretely punctate, without distinct ground sculpture. Pronotwn about one-sixth longer than wide; anterior angles narrowly rounded ; sides scarcely emarginate in front, moderately arcuate to base; rather indistinctly and confusedly punctate, not distinctly sculptured but rather coriaceous. ///ytra not punctate but with tuberculate sculp- ture, the tuberculi rather feeble and rounded, surface uneven. JZale, fourth and fifth sternites with a transverse fold, seventh slightly emarginate and feebly impressed, eighth with a deep narrow excision, the outer angles broadly rounded. /emadle, eighth sternite rounded. Length, 214 to 3 mm. Type locality. — Jamaica. Types.—tIn collection of Dr. Cameron. Records—The following are the records known to me: Cuba: Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Cameron, 1913; Leng and Mutchler, 1917; Scheerpeltz, 1983 ; Hubbard, in U.S.N.M.), Trinityville (Blackwelder staticn 428), Montego Bay (Black- welder station 407), Santa Cruz (Blackwelder station 421), Gordon Town (Blackwelder stations 868B, 382, 383), Blue Mountains (Darlington, in M.C.Z.). Specimens examined.—I have seen the type and 8 other specimens in Dr. Cameron’s collection, 1 in the Museum of Comparative Zoology, 2 females in the United States National Museum, and 19 examples collected by Chapin and Blackwelder in February 1937. Remarks.—There is some variation in the density of the punctation of the head and pronotum and likewise in the appearance of vague ground sculpture. The shape of the head and the characters of the male indicate that this species belongs in the subgenus Scepaeus s. str., although the classification of the subgenera is not yet in a very settled condition. Our specimens were collected under stones along the edge of streams and flying at dusk, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 287 8. SCOPAEUS PYGMAEUS Erichson Scopaeus pygmaeus ErtcHson, 1840, p. 608.—BERNHAUER and ScHuseRt, 1912, p. 251.—Lena and Mutcuier, 1914, p. 405—Wotcorr, 1924, p. 78; 1936, p. 197.—BLACKWELDER, 1939a, p. 106. Scopaeus simplicicollis CAMERON, 1913b, p. 348.—LENG and MuTcHier, 1917, p. 199.—ScCHEERPELTZ, 1933, p. 1269.—BLAcCKWELDER, 1939a, p. 106. Description.—Cinnamomeous, sometimes picescent or paler in part, apical half of each elytron indefinitely bordered with testaceous. Head very feebly emarginate behind, the angles narrowly rounded; eyes separated from base by two and one-half times their length; labrum with two very slender median denticles and with a distinct pair of outer denticles; gular sutures rather narrowly separated anteriorly, feebly divergent posteriorly; finely and densely punctate but with intervals shining; without ground sculpture. Pronotum one- fifth longer than wide; anterior angles moderately narrowly rounded, sides in front scarcely emarginate, feebly arcuate to base; midline feeble, slightly depressed; not distinctly punctate but with dense though not coarse ground sculpture. lytra not very distinctly punctate but pubescent and minutely and indefinitely broken up, with ground sculpture along suture at base. Male, fourth and fifth sternites without trace of transverse fold, seventh very slightly emarginate; eighth sternite with a small abrupt rounded emargination. Female, eighth sternite slightly produced at middle; eighth tergite rounded. Length, 2 to 244 mm. Type locality—Puerto Rico. Of simplicicollis, Balthazar, Gre- nada. Vypes.—Kither in Hope Museum, Oxford, or the Zoologische Mu- seum, Berlin. Of stmplicicollis, in the British Museum. Records.—The following are the records known to me: Jamaica: Spanish Town (Blackwelder station 377), Santa Cruz (Blackwelder station 421), Black River (Blackwelder station 416), May Pen (Blackwelder station 425B), Milk River (Blackwelder station 415). Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914; Wolcott, 1924, 1936), Guanica (Blackwelder station 355A), Jayuya (Oakley, in United States National Museum). St. Croix: (Blackwelder stations 341, 344; Beatty, in United States National Museum). St. Lucia: (Cameron collection). Barbados: (British Museum). St. Vincent: (Cameron, 1913, as simplicicollis; British Museum; Cameron col- lection; Scheerpeltz, 1933; Blackwelder stations 170, 174, 176A). Grenadines: Mustique (British Museum; Cameron collection). Grenada: (Cameron, 1913, as simplicicollis; British Museum; Cameron collec- tion; Scheerpeltz, 1933). Specimens exwamined.—I have examined 122 examples in the British Museum (including the type of simplicicollis) , 29 examples in the col- 288 BULLETIN 182, UNITED STATES NATIONAL MUSEUM lection of Dr. Cameron, 1 in the United States National Museum, and 57 collected by Chapin and Blackwelder in 1935-87. Remarks.—This is a variable species in respect to the sculpture of the pronotum and elytra, the length of the temples, and the abrupt- ness of the emargination of the eighth sternite of the male. The de- scription is drawn principally from a specimen from St. Vincent. This species belongs in the subgenus Scopaeus s. str. My specimens were collected from rubbish on the muddy bank of Jagoons and ponds, from under seaweed on the beach, from under stones along streams, and flying at dusk. 9. SCOPAEUS FILUM Sharp Scopaeus fllum SHarp, 1886, p. 542.—Casry, 1886, p. 220—BrRNH“AUER and ScHu- BERT, 1912, p. 247.—-BLACKWELDER, 1939a, p. 106. Description.—Castaneous or ferrugineous to testaceous. Head trun- cate behind, not distinctly emarginate, angles moderately rounded; labrum with two small denticles at middle; gular sutures rather nar- rowly separated in front of middle, feebly divergent posteriorly; very finely and rather densely punctate but with shining intervals; without ground sculpture. Pronotwm one-fifth longer than wide, anterior angles rather broadly rounded, sides not distinctly emargi- nate in front, straight behind and broadly rounded into base; midline sometimes feebly channeled at base or apex; pronotum without dis- tinct punctures except minute punctulae; surface slightly uneven but without ground sculpture. Zlytra not punctate but with a dense in- distinct sculpture which seems to be obsoletely tuberculate. Male, fourth and fifth sternites without transverse folds, seventh not dis- tinctly emarginate; eighth sternite with a small shallow rounded abrupt emargination, less than one-half as deep as wide. Female, eighth sternite prominently rounded; eighth tergite rounded. Length, 2 mm. Type locality.—Guatemala, Paso Antonio. Types—tIn the British Museum. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Baragua (Christensen, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Cameron collection), Milk River (Blackwelder station 415). Hispaniola: Hartt (Cameron collection), Dominican Repusiic, Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.), Sanchez (Darlington, in M.C.Z.), Barahona (Darlington, in M.C.Z. and U.S.N.M.), Villa Altagracia (Darling- ton, in M.C.Z.). Central America: GUATEMALA (Sharp, 1886). Specimens examined.—I have seen 8 examples in the United States National Museum, 16 from the Museum of Comparative Zoology, 9 in the collection of Dr. Cameron, and 2 collected by Chapin and Black- welder in February 1937. I have not examined the types. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 289 Remarks.—Since I have not compared the West Indian specimens with the type of fidwm I am not able to say definitely that they are the same. The description of that species agrees entirely with our speci- mens except that in the latter the elytra are not visibly punctate. I believe it probable that they are the same. My specimens were collected at light or flying at dusk; the Soledad examples were taken in a Berlese funnel. 10. SCOPAEUS CUPIENS, new species Description.—Piceous to rufopiceous, elytra and abdomen castane- ous. Head slightly emarginate behind, angles moderately rounded ; labrum with two small denticles at middle, each somewhat enlarged at base; gular sutures straight, narrowly separated in front, divergent posteriorly; punctures distinct and dense but surrounded by fine in- distinct ground sculpture. Pronotwm nearly one-sixth longer than wide; anterior angles rather narrowly rounded; sides in front dis- tinctly emarginate, moderately arcuate posteriorly; with indistinct smooth midline; not distinctly punctured but sculptured to resemble obsolete umbilicate punctures, more distinctly punctiform along mid- dle; without other ground sculpture. 2lytra not punctate but with small tuberculi separated by less than their diameter, and with addi- tional ground sculpture along suture basally. fale, unknown. Female, eighth sternite broadly rounded; eighth tergite rounded at apex. Length, 214 mm. Type locality —Jamaica. Types.—Holotype, female, and one paratype, U.S.N.M. No. 52416, collected by H. G. Hubbard. Records.—The following are the records known to me: Jamaica: (Hubbard, in United States National Museum), Santa Cruz (Black- welder station 421). Specimens ecamined.—I have seen only the two types. Remarks.—The paratype is imperfect, having lost the abdomen. The species is distinguished from the others principally by details of sculpture. The shape and sculpture of the head place it in the sub- genus Scopaeus s. str. The paratype was caught flying at dusk. 11. SCOPAEUS ANGUSTICOLLIS Cameron Scopaeus angusticollis CAMERON, 1913b, p. 346.—LeNG and Murcuner, 1917, p. 199.—ScHEERPELTZ, 1933, p. 1265. Description.—Reddish brown, abdomen lighter with posterior mar- gin of tergites and apex testaceous. Head emarginate posteriorly, angles rounded; closely and finely punctate. Pronotum. one-fourth longer than wide, anterior angles rounded but traceable; feebly nar- rowed posteriorly; with a fine median line posteriorly and an obsolete 290 BULLETIN 182, UNITED STATES NATIONAL MUSEUM impression on each side near base; punctures fine and close, much finer than on head. £lytra finely and closely but less distinctly punctured than pronotum. Male, unknown. Female (presumably with ster- nites unmodified). Length, 344 mm. (From Cameron.) Type locality —Grenada, Balthazar. Types.—Unique female in the British Museum. Records.—The following are the records, known to me: Cuba: Aguadores, in Oriente (Darlington, in M.C.Z.). Grenada: (Cameron, 1918; Leng and Mutchler, 1917; Scheerpeltz, 1983; British Museum). Specimens examined.—I have seen only the unique type and one specimen from the Museum of Comparative Zoology, as explained below. ; Remarks.—Although the original description does not mention the presence or absence of ground sculpture on the head, I assume that it is not present. This species belongs in the subgenus Scopaeus s. str. Since the above description was written I have received one ex- ample from Cuba that agrees very closely with it. The specimen is a female and shows the following additional characters: Head with- out ground sculpture; eyes separated from base of head by two and a half times their length; pronotum unusually rounded but much de- pressed, without posterior depressions, although the disk is flattened ; length, 3 mm. I find no record of its habits. 12. SCOPAEUS ARENA, new species Description.—Head rufous, pronotum rufous or rufopiceous, elytra dark rufopiceous with apex narrowly testaceous, abdomen castaneous. Head truncate behind, angles moderately broadly rounded; labrum with two large prominent teeth more or less modified or denticulate at base, and a pair of minute outer denticles; gular sutures narrowly separated, parallel from in front of middle; without punctation but with very fine dense ground sculpture. Pronotum two-sevenths longer than wide; anterior angles rounded continuously with sides, not distinctly emarginate in front, feebly arcuate posteriorly; mid- line vaguely elevated posteriorly; sculptured similarly to head or a trifle more distinctly. Hlytra not punctate but with rather dense rounded tuberculi, with additional tuberculate ground sculpture along suture at base. Male unknown. Female, eighth sternite rounded. Length, 344 to 3144 mm. Type locality—St. Lucia, Millette River, at junction with the Roseau River, 5 miles south of Castries. Tyves.—Holotype and one paratype, females, United States Na- tional Museum No, 52418, collected by me on April 3, 1936. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 291 Reeords.—The following is the only record known to me: St. Lucia: (Blackwelder station 211A). Specimens examined.—I have seen only the two types. Remarks.—The shape and sculpture of the head place this species in the subgenus Scopaeus s. str. The types were taken under stones and sand at the edge of a stream in the dense forest. 13. SCOPAEUS DARLINGTONI, new species Description.—Piceocastaneous. Head feebly emarginate behind, the angles rather broadly rounded; labrum with two small median denticles and possibly a lateral pair; gular sutures parallel, narrowly separated; with dense round tuberculi somewhat obscured on disk: without distinct ground sculpture. Pronotum one-ninth longer than wide, the anterior angles rather broadly rounded, sides scarcely emar- ginate in front, evenly curved into base behind; disk depressed on either side of midline near base, midline very fine; with round tuber- culi more distinct than on head; without distinct ground sculpture. Elytra not punctate but with tuberculi as on pronotum, a little coarser but somewhat confused by indistinctly rugulose surface. Male, fourth and fifth sternites with a complete transverse fold on each at middle, seventh broadly feebly emarginate; eighth with a broad feeble emargination opening into a triangular excision about as deep as wide. Female, unknown. Length, 314 mm. Type locality —Cuba, Buenos Aires in Trinidad Mountains, eleva- tion 2,500 to 3,500 feet. 7Types.—Holotype, male, in the Museum of Comparative Zoology, collected on May 8-14, 1936, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Cuba: Buenos Aires, in Trinidad Mountains (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species is at once distinguishable by the tuberculi on the head, pronotum, and elytra. Those on the head can be ob- served only with a magnification of about 60 times. S. arena has the tuberculi on the elytra but lacks them on the pronotum and head. The svecies belongs in the subgenus Scopaeus s. str. I have received no record of its habits. 14. SCOPAEUS AURIPILIS Cameron Scopacus auripilis Cameron, 1913b, p. 347. —Lene and Murcurer, 1917, p. 199.— SCHEERPELTZ, 1933, p. 1265. Description—Head and pronotum ferrugineous or black, elytra black with apical margin sharply testaceous, abdomen black with apex of eighth and ninth tergites testaceous. Head emarginate behind, 292 BULLETIN 182, UNITED STATES NATIONAL MUSEUM posterior angles right, only blunted at the extreme apex; punctation close, rugulose, umbilicate. Pronotwm scarcely longer than broad, widest at anterior angles which are obtuse, gradually narrowed behind to rounded posterior angles, more abruptly in front; with a very fine shining median carinula at base but without trace of basal foveae; punctation as on head. “lytra with punctures close and rugulose, rather finer than that of pronotum, especially behind. Abdomen closely and finely punctured. Pubescence long and yellow throughout. Male, unknown. Female, without special modifications. Length, 3 mm. (From Cameron.) Type locality —Grenada, Black Forest Estate, Windward side. Types.—Unique female in the British Museum. Records —The following are the records known to me: St. Vincent: (Cameron, 1918; Leng aud Mutchler, 1917; Cameron collection ; Scheerpeltz, 1933). Grenada: (Cameron, 1913; Leng and Mutehler, 1917; British Museum). Specimens examimed.—I have seen only the type in the British Museum and one example in Dr. Cameron’s collection. Remarks.—My examination of the type confirms the relative dimen- sions given by Dr. Cameron. One specimen from each island is all that are now known of this species. The shape and sculpture of the head indicate that it belongs in the subgenus Scopacus s. str. I find no record of its habits. 15. SCOPAEUS RAMBOUSEKI (Bierig) Scopaeopsis rambouseki Birric, 1984a, p. 24, 26, 27, figs. 1-10. Description.—Piceous, anterior margin of pronotum rufopiceous. Head truncate or feebly rounded behind, angles rounded, sides feebly converging from eyes posteriorly; labrum with two large slender teeth somewhat enlarged and modified at base, and with two small denticles outside the others; gular sutures parallel, very narrowly separated throughout; very finely and indistinctly punctate but with smooth intervals; without ground sculpture. Pronotum about one-fifth longer than wide; anterior angles broadly rounded, sides straight in front and moderately arcuate behind; with indistinct midline and feeble basal impressions; rather indistinctly and irregularly punctate but with flat intervals. lytra not punctate, but with feeble and irregular tuberculi, often separated by their diameter, intervals with traces of ground sculpture. fale, unkown. Female, seventh tergite with a small abrupt shallow emargination at middle, eighth tergite biemar- ginate to form a small triangular median lobe; eighth sternite not modified. Length, 314 mm. Type locality.—Cuba, “Morro (Habana) ; Rio Ariguanabo; Caimito del Guayabal.” MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 293 Types—In the collection of Sr. Bierig. One paratype is in the collection of Dr. Cameron and one is in the United States National Museum (No. 52725). Records.—The following are the records known to me: Cuba: (Cameron collection), Habana (Bierig, 1934), Rio Ariguanabo (Bierig, 1934), Caimito del Guayabal (Bierig, 1984), Baragua (Christenson, in U.S.N.M.), Soledad (Weber, in M.C.Z.). Specimens examined.—I did not see the “cotype” supposed to be in Dr. Cameron’s collection, but I have studied one in the United States National Museum; also two examples in the National Museum aud one specimen from the Museum of Comparative Zoology. Remarks.—The characters of the tergites of the female distinguish this species readily. The specimen from Soledad does not show the emargination of the seventh tergite. ‘There is some difference in the shape of the head between the two Baragua specimens, in one being truneate, in the other distinctly rounded. The Soledad specimen agrees with the latter in this character. This species was described in the genus Scopaeopsis, which is here considered to be a subgenus of Scopaeus. ‘The shape of the head and the characters of the male, how- ever, indicate that it belongs in the subgenus Scopaeus s. str. Bierig records the species as follows: “Vive en orillas humedas y parec ser raro”; the other specimens were taken at light or by sweeping. 16. SCOPAEUS BOX], new species Description—Head and pronotum rufous or piceorufous, elytra testaceous to castaneous, indefinitely paler at apex, abdomen castane- ous. Head truncate behind, almost rounded, the angles moderately rounded; eyes separated from base by less than twice their length; labrum with two denticles at middle and a small triangular additional denticle at each side; gular sutures moderately separated, most ap- proximate a little before middle, very feebly diverging posteriorly ; finely and densely punctured, but irregularly and with vague ground sculpture. Pronotum one-fourth longer than wide; anterior angles almost completely rounded from neck to sides, which are nearly straight; with trace of smooth midline; punctures fine and dense; without ground sculpture. Hlytra with punctures indistinct but dense, the surface appearing subasperate. J/ale, fourth and fifth sternites with a transverse fold, seventh not emarginate, eighth sinuate at mid- dle but not distinctly emarginate. Memale, eighth sternite rounded ; eighth tergite a little prominent at middle. Length, 2 to 244 mm. Type locality —St. Lucia, Roseau River, at elevation of 200 feet, about 6 miles above its mouth. Types.—Holotype, male, and three paratypes, U.S. N. M. No, 52418, collected by me on March 29, 1936. 294 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Pecords.—The following is the only record known to me: St. Lucia: (Blackwelder station 209A), Specimens examined.—I have seen only the four types. Remarks.—This species is very similar to rivularis from Jamaica but differs constantly in numerous small characters. The elytra are not distinctly bordered in apical half; the head is not at all emarginate behind; the temples are shorter; the labrum has an extra pair of denticles; the gular sutures are closest less anteriorly; the ground sculpture of the head is less distinct; with trace of smooth midline; and eighth sternite of male even more feebly emarginate. Until we find specimens on the intervening islands, it is best to keep the two separate. The species appears to belong in the subgenus Scopaeus s. str. The types were found in sand along the stream. 17. SCOPAEUS PULCHELLUS Erichson Scopacus pulchellus ErtcuHson, 1840, p. 609.—SuHarp, 1876, p. 248; 1886, p. 547.— CasEy, 1886b, p. 220; 1905, p. 217.—BERNHAUVER and ScHuneErT, 1912, p. 250.— BLACKWELDER, 1939a, p. 106. Scopaeus apicipennis SHARP, 1886, pp. 547, 548.—Bernuaver and Scuusert, 1912. p. 250. Scopaeodera pulchella (BHrichson) Casey, 1856b, p. 220; 1905, p. 217. Description—Dark rufous, outer apical angles of elytra luteous, abdomen castaneous, picescent at apex. Head rounded behind but emarginately interrupted at neck; labrum with two large teeth sep- arated by a rather large medium notch, and two small outer denticles: gular sutures moderately separated, most approximate before mid- dle; with very sparse obsolete and fine punctures, surface very smooth and strongly shining. Pronotwn one-third longer than wide, anterior angles very broadly rounded; sides in front feebly emarginate, pos- terior evenly arcuate to base; smooth, shining, almost entirely im- punctate; without midline of any sort. #A7y/ra not distinctly punctate or sculptured, surface slightly uneven but shining. J/c/e, fourth and fifth sternites without trace of transverse fold; seventh not emarginate, but with a narrow longitudinal groove apically; eighth sternite miss- ing from my specimen. /ema/e, eighth sternite prominently rounded. Length, 224 mm. Type locality —‘V alle Araguensi Columbiae” (= Venezuela). Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Grenada: (British Museum: Cameron collection; Blackwelder station 155). Trinidad: Tacarigua (Blackwelder station 107A). South America: VENEZUELA (Erichson, 1840), CotomBta (Casey, 1886, 1905; Sharp, 1876, 1886). Central America: Mrextco (Bernhauer and Schubert, 1912), Guaremata (Sharp, 1886, as apicipennis), PANAMA (Sharp, 1886, as apicipennis). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 295 Specimens examined.—I have seen seven examples in the British Museum, two in Dr. Cameron’s collection, and three collected by me in 1935-37. Remarks.—Although I have seen no examples from South America, it seems likely that these specimens from the West Indies are Erich- son’s species. They were so identified in the British Museum and Dr. Cameron’s collection. The above description was taken from my Trini- dad examples. The specimen I have referred to as a male has the abdomen muti- lated. The sixth, seventh, eighth, and ninth tergites and the eighth and ninth sternites are missing. The groove on the seventh sternite is the only evidence that it isa male. The smooth integuments of this species place it in the subgenus Scopaeodera. The types and my specimens were found under stones at the edge of the river. 18. SCOPAEUS CHAPINI, new species Description.—Piceous, pronotum rufopiceous. Head completely and evenly rounded behind from eye to eye; labrum rather strongly bidentate, with a moderate notch between the teeth; gular sutures most approximate along middle, never very close; with dense punctation or sculpture nearly obsolete, surface strongly shining. Pronotwm one- third longer than wide, anterior angles rounded continuously with sides, sides in front feebly emarginate, posteriorly evenly arcuate to base; sculptured still more obsoletely than head, surface strongly shining, without definite midline; with two rather strong impressions at base leaving a narrow cariniform median ridge. Hlytra coriaceous, not punctate but with sparse obsolescent tuberculi; shining. dale, fourth and fifth sternites without trace of a transverse fold; seventh sternite very feebly and widely emarginate; eighth sternite with a broad triangular emargination, one-half as deep as wide, with outer angles completely rounded into sides of sternite. Hemale, unknown. Length, 3 mm. Type locality—Jamaica, between Luana and Kensworth via La- covia, Santa Cruz, and Spur Tree, parishes of St. Elizabeth and Manchester. Types—Holotype, male, U.S.N.M. No. 52412, collected by Chapin and Blackwelder on February 24, 1937. Records —The following is the only record known to me: Jamaica: Santa Cruz (Blackwelder station 421). Specimens examined.—\ have seen only the unique type. Remarks.—This species does not fall clearly into any subgenus in my key (Blackwelder, 19392). However, if the obsolescent sculpture be considered to be in the process of disappearing, phylogenetically, 296 BULLETIN 182, UNITED STATES NATIONAL MUSEUM then the species keys out to Scopacoma, in which it seems to belong by the combination of other characters. The type was caught flying at dusk. XXXIX. Genus MONISTA Sharp Monista SHARP, 1876, p. 271. Genotype.—M onista typica. Sharp (original designation; Interna- tional Rules, Article 30, I, d). Diagnosis.—Head finely punctate, sometimes biimpressed to form a median ridge; eyes small; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, first and second not enlarged; labrum rather large, feebly emarginate, without denticles; gular sutures separated but parallel and very approximate; fourth segment of maxillary palpus subulate; neck one-fifth as wide as head or less; prothorax not prolonged in front; prosternum rather widely dilated under the front coxae but widely separated from the hypomera; hypomera feebly lobed behind the coxae; front coxal cavities narrowly open behind; anterior coxae large, exserted ; middle coxal cavities con- fluent ; posterior coxae contiguous, “conical”; first and second abdomi- nal sternites absent, eighth emarginate in male; apex of posterior tibia with a distinct ctenidium only on the inner edge; tarsi 5-segmented, fourth segment lobed beneath. Remarks—The West Indian species of this genus are at once distinguished from all Staphylinidae known to me by the peculiar carinate structure of the vertex of the male. This is apparently not present on many of the species described from South and Central America. The genus has been placed near Astenus and E'chiaster but is more similar in structure to Scopaeus and Stilicus. I have seen from the West Indies 34 specimens belonging to four species of which three are new. KEY TO WEST INDIAN SPECIES OF MONISTA 1. Pronotum with fine ground sculpture ; cephalic carina close to neck. 1. personata Pronotum without distinct ground sculpture; cephalic carina distant from TGC HK 8) Beat ee te MR I hn Se ea ee 2 2. Head minutely tuberculate.=-- +--+ 5 oe eee 4. vola Head more or less distinctly punctate, not at all tubereulate____________ 3 3. Punectures of head and pronotum obsolescent__-------__------- 2. trinitatis Punctures of head and pronotum distinct-__---___-___--__- 3. darlingtoni 1. MONISTA PERSONATA Cameron Monista personata CAMERON, 1913b, p. 339.—LeNG and MutTcHter, 1917, p. 198..— SCHEERPELTZ, 1933, p. 1240.—BLACKWELDER, 1939a, p. 106. Description —Rufotestaceous, apex of elytra paler. Head orbicu- lar; eyes small, not prominent; vertex modified in male; impunctate, but with a few tiny tuberculi especially above the eyes; with distinct MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 297 ground sculpture above and behind but merely coriaceous in front; labrum broadly rounded but very feebly emarginate at middle. Pronotum as broad as long, evenly rounded at sides but more nar- rowed in front; with feebly umbilicate punctures separated by less than their diameter and obscured by fine ground sculpture. Hlytra rather finely and indistinctly submuricately punctate, surface some- what uneven but punctures net abrupt; without ground sculpture. Abdomen finely sparsely and minutely submuricately punctured; without ground sculpture. dfale, vertex behind the eyes with a cir- cular depression divided longitudinally by a strong carina; eighth abdominal sternite broadly and feebly emarginate. Memale, vertex unmodified; eighth sternite not emarginate. Length, 214 to 3 mm. Type locality —Grenada. Types.——Twelve specimens (one labeled type) in the British Museum. Records.—The following is the only record known to me: Grenada: (Cameron, 1913; Leng and Mutchler, 1917; British Museum). Specimens examined.—I\ have seen the 12 examples in the original series in the British Museum and 7 in the collection of Dr. Cameron, One example was borrowed from the British Museum for further study. Remarks—The male of this species is easily recognizable at a glance by the unusual carinate head and the posterior position of the carina. [ find no record of its habits. 2. MONISTA TRINITATIS, new species Description—Rufous or rufotestaceous, with apex of elytra broadly paler. Head subhexagonal; eyes small but prominent, at over twice their length from base; vertex not modified (in female) ; labrum broadly rounded but feebly emarginate at middle; with rather: coarse punctures nearly obsolete throughout, nearly tuberculose basally; with irregular ground sculpture on the clypeus. Pronotum one-twelfth longer than wide, widest at anterior two-fifths, thence. strongly narrowed to neck, moderately narrowed to base; with punc- tures as on head, but without distinct ground sculpture. //ytra with punctures less distinct than pronotum, the intervals smooth and flat; without distinct ground sculpture. Abdomen finely sparsely and minutely submuricately punctate; without ground sculpture. fale, unknown. Female, sternites unmodified. Length, 21 mm. Type locality—Trinidad, foothills north of Tunapuna. Types.—Holotype in the Museum of Comparative Zoology; one paratype in the United States National Museum (No. 52357) ; col-. lected on June 29, 1935, and May 25, 1935, by N. A. Weber. 449008—42 20 298 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following is the only record known to me: Trinidad: Tunapuna (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species is distinguished by its obsolescent puncta- tion and lack of pronotal ground sculpture. I have received no record of its habits. 3. MONISTA DARLINGTONI, new species Description.—Head and pronotum piceous, elytra rufopiceous with apical fourth testaceous, abdomen rufous. Head suborbicular; eyes small but prominent, at over twice their length from base; vertex not modified (in female); labrum broadly rounded and_ feebly emarginate at middle; punctures coarse and distinct, usually separated by less than half their diameter; without distinct ground sculpture. Pronotum as broad as long, widest at anterior two-thirds, thence strongly narrowed to neck, and rather strongly to base; with punctures as on head but a little less distinct; without ground sculpture. Z?ytra with umbilicate punctures almost entirely obso- lete having merely the central setigerous punctule: without distinct ground sculpture. Abdomen finely, sparsely, and minutely submu- ricately punctate; without ground sculpture. Male, unknown, Female, sternites unmodified. Length, 314 mm. Type locality —Trinidad, St. Augustine. Types.—Holotype in the Museum of Comparative Zoology; one paratype in the United States National Museum (No, 52358) ; col- lected in April 1929 by Dr. P. J. Darlington. Records—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens ewamined.—I\ have seen only the two types. Remarks.—This species is distinguished by the distinct punctures of the head and pronotum and the absence of sculpture on the clypeus. Like the preceding species the lack of males leaves us in doubt as to the condition of the vertex in that sex. I have received no record of its habits. 4. MONISTA VOLA, new species Description —Rufous, apex of elytra and all of abdomen rufotes- taceous. /Zead subquadrate; eyes small but somewhat prominent; vertex modified in male; impunctate but with rather dense setigerous tubereuli especially posteriorly, and with minute ground sculpture on clypeus; labrum broadly rounded but feebly emarginate at middle. Pronotum one-sixth longer than wide, widest at anterior two-fifths, thence strongly narrowed to neck, feebly narrowed to base; with umbilicate punctures almost completely obsolete, making surface MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 299 slightly uneven; without ground sculpture. lytra sparsely, finely, and irregularly submuricately punctate, surface between smooth and flat; without ground sculpture. Abdomen finely, sparsely, and minutely submuricately punctate; without ground sculpture. Male, vertex between the eyes with a circular depression divided by a longitudinal carina which ends posteriorly in a smooth excrescence and which bears several long hairs; eighth sternite feebly broadly emarginate. Female, unknown. Length, 2144 mm. Type locality—Dominica, one-fourth mile north of the Boery River on coastal plain. Types.—Holotype, male, U.S.N.M. No. 52407, collected by me on June 26, 1936. Records—The following is the only record known to me: Dominica: (Blackwelder station 252). Specimens examined—I have seen only the unique type. Remarks.—Like personata this is distinguished by its carinate head but differs from it in having the carina distant from the neck, between the eyes, The type was caught flying at dusk. XL. Genus RUGILUS Samouelle Rugilus SAMOVELLE, 1819, p. 173. Stilicus LEPELETIER and SeRvILLe, 1825, p. 495. Stilicosoma Casey, 1905, p. 219. Genotypes.—Paederis orbicwlatus Fabricius=Rugilus orbiculatus (Fabricius) (original designation; also designated by Curtis, 1828). Of Stilicus, Paederus orbiculatus Fabricius=Stilicus orbiculatus (Fabricius) (designated by Thomson, 1861); of Stilicosoma, Stili- cus rufipes Germar=Stilicosoma rufipes (Germar) (monobasic). Diegnosis—Head not emarginate posteriorly, with dense umbilicate punctures but without ground sculpture; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, first and second not large with rest small; labrum large and broad, not much emarginate, bidenticulate at middle, with outer angles more or less toothed; fourth segment of maxillary palpus subulate; gular sutures united; neck jess than one-fifth as wide as head; pronotum strongly narrowed in front but not prolonged; prosternum carinate between coxae but not much dilated under them; anterior coxae large, ex- serted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; seventh ster- nite sometimes and eighth always emarginate in male; apex of posterior tibia with a ctenidium only on inner sides. Remarks.—In Casey’s key to the Paederini the statement that this and related genera have the prosternum “greatly dilated laterally 300 BULLETIN 182, UNITED STATES NATIONAL MUSEUM under the coxae as far as the inflexed sides of the pronotum” is mis- leading. Even in views from directly above or behind these lateral expansions do not reach laterally nearly as far as the hypomera, in fact are scarcely at all dilated. In my key (Blackwelder, 1939a) these genera are grouped with those not expanded as far as the hypomera. The name Rugilus was validated in 1819 by Samouelle and is there- fore older than Stilicus, which appears to have been used first in 1825 by Lepeletier and Serviile. The record of Stilicus cribratus Sharp from Jamaica by Gowdey, 1926, is probably a misidentification. Until the specimens can be reexamined, it must be regarded as a doubtful record. This is a world-wide genus of considerable size. It is wel! repre- sented in America, both temperate and tropical. I have examined 17 specimens from the West Indies in the British Museum, 11 in Dr. Cameron’s collection, 25 from the Museum of Comparative Zoology, 12 in the United States National Museum, and 9 collected by Chapin and Blackwelder in 1935-87. These I assign to seven species, of which two seem to be new. KEY TO WEST INDIAN SPECIES OF RUGILUS 1. Pronotum distinctly punctate only along the midline______---_ 7. coprophilus Pronotum coarsely and strongly umbilicately punctate throughout, except FOO MIdMN eee 235 Oe ee ea a a ee ee 2 2: eaytrarcoarsely punctate se = ee ee eae os See ee ee 3 EKlytra finely or indistinctly, not densely punectate____------__-- 6. insularis 3. Elytra with coarse separate punctures_________._____-.--__-__-- 1. buscki Elytra with dense coarse rugulose sculpture_________-.__-__--__-_------- 4 4. Seventh sternite of male emarginate or tuberculate_______-____-_--_-_-_- 5 Seventh sternite of: male unmodified_..--.-- 2242... eet 2. jucundus 5. Seventh sternite of male with simple emargination_____--__________-___~- 6 Seventh sternite of male with additional modifications__________ 5. chthonus 6. Seventh sternite of male with rounded emargination one-third as deep as wide; anglessroundedio2 22. 20S 2 ae Ae ee 3. agnatus Seventh sternite of male with semicircular emargination two-thirds as deep aS wide, angles not rounded___--__-_____________-____ 4. cupreus 1. RUGILUS BUSCKI, new species Description Black, shining. ead scarcely wider than long, rounded behind except for a slight prominence above neck, angles entirely obliterated; eyes large, at less than their length from base ; labrum with two very minute denticles at middle, lateral angles indistinctly dentate; umbilicate punctures moderately large, fre- quently elongate, rather dense throughout; without smooth area on vertex; without distinct ground sculpture. Pronotwm one-sixth longer than wide, about three-fourths as wide as head; widest at anterolateral angles; sides scarcely narrowed in front, feebly arcuate posteriorly to base; with a distinct but narrow smooth median band; - MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 301 umbilicate punctures a little smaller than on head, more regular, dense; without smooth areas; without ground sculpture. /ytra as wide as long, as wide as head; conjointly emarginate behind; punc- tures very coarse and deep, separated by convex intervals; without ground sculpture. Abdomen finely and moderately densely sub- muricately punctate. Male, unknown. Female, sternites not modi- fied. Length, 4144 mm. Type locality —Trinidad, ward of Montserrat. Types.—Holotype, female, U.S.N.M. No. 52411, collected on June 29, 1903, by August Busck. Records.—The following is the only record known to me: Trinidad: Montserrat (Busck, in United States National Museum). Specimens examined —I have seen only the unique type. Remarks.—This species is sufficiently distinguished from the other West Indian species, but I am unable to assert that it is positively distinct from the South American ones. 2. jugalis Erichson is the only species recorded from Venezuela, but I have not seen examples of it. I have no record of its habits. 2. RUGILUS JUCUNDUS (Cameron) Stilicus jucundus CAMERON, 1918b, p. 340.—LEne and MuTcHLER, 1917, p. 198.— SCHEERPELTZ, 1933, p. 1242.—BLacKWELpDER, 1939a, p. 107. Description.—Piceous, slightly rufescent, apex of elytra narrowly testaceous, more widely at sides. Mead scarcely wider than long, rounded behind except for a slight prominence above neck, angles entirely obliterated; eyes large, at less than their length from base; labrum with two rather large denticles at middle, outer angles also produced in small teeth; umbilicate punctures moderate, seldom elongate, not very dense; without distinct smooth area on vertex; without distinct ground sculpture. Pronotwm two-ninths longer than wide, three-fourths as wide as head; widest at anterolateral angles; sides very feebly emarginate in front, feebly arcuate posteriorly to base; with a distinct elevated smooth median band; umbilicate punc- tures a little smaller than on head, dense, without distinct smooth areas; without ground sculpture. H/ytra as wide as long, one- twelfth wider than head; conjointly emarginate behind; punctures coarse and deep, separated by broad convex intervals; without ground sculpture. Abdomen very finely and densely submuricately punc- tate. Male, seventh sternite not modified; eighth sternite with a triangular excision deeper than wide, the angles narrowly rounded. Female, sternites unmodified. Length, 4 to 444 mm. Type locality —St. Vincent, Leeward side. Types.—In the British Museum. 302 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following are the records known to me: St. Vincent: (Cameron, 1913; Leng and Mutchler, 1917; British Museum; Cam- eron collection). Grenada: (Cameron collection). Specimens examined.—I have seen 12 examples (including the type) in the British Museum and 7 in Dr. Cameron’s collection. One of the former was borrowed for further study. Remarks.—Distinguished from agnatus, besides the characters men- tioned in the key, by the seventh sternite. The above description was drawn entirely from a male borrowed from the British Museum. I find no record of its habits. 3. RUGILUS AGNATUS (Cameron) Stilicus agnatus CAMERON, 1913b, p. 340.-—LENG and MurcHirr. 1917, p. 198.— SCHEERPELTZ, 1933, p. 1240.—BLACKWELDER, 1939a, p. 107. Description.—Black, shining, apex of elytra narrowly testaceous, more broadly at sides. /Zead scarcely wider than long, rounded behind except for a slight prominence above neck, angles broadly rounded ; eyes large, at less than their length from base; labrum finely bidentic- ulate at middle, lateral angles dentate; umbilicate punctures rather large especially on vertex, not often elongate; with an indefinite smooth area at center of vertex; without ground sculpture. Pronotwm one- sixth longer than wide, three-fourths as wide as head; widest at an- terolateral angles; sides in front feebly emarginate, posteriorly feebly arcuate to base; with a distinct elevated smooth middle band; umbili- cate punctures large, on the disk often separated by smooth intervals but without a distinct smooth area; without ground sculpture. /lytra scarcely longer than wide, as wide as head; conjointly emarginate be- hind; punctures coarse and deep, not dense, with strong convex in- tervals; without ground sculpture. Abdomen very finely and indis- tinctly punctured. Afale, seventh sternite with a rounded emargina- tion, one-third as deep as wide, lateral angles broadly rounded; eighth sternite with an abrupt triangular excision about as deep as wide, angles very narrowly rounded. Female, sternites not modified. Length, 314 to 4mm. Type locality —Grenada, Balthazar. Types.—In the British Museum. Records.—The following is the only record known to me: Grenada: (Cameron, 1918; Leng and Mutchler, 1917; British Museum; Cameron collection). Specimens examined.—I have seen five examples, including the type, in the British Museum and two in the collection of Dr. Cameron. One of the former was borrowed for further study. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 303 Remarks.—The above description of the male characters differs from Dr. Cameron’s description in the structure of the seventh sternite. It was taken from one of the original series borrowed from the British Museum. The species is apparently even more distinct from jucundus than was previously thought. I find no record of its habits. 4. RUGILUS CUPREUS (Cameron) Stilicus cupreus CAMERON, 1913b, p. 341.—LENe and MurcHtier, 1917, p. 198.— SCHEERPELTZ, 1933, p. 1241. Description.—Bronze-black with coppery lustre, elytra brown, shin- ing. Head transverse, posterior angles rounded, as wide as elytra; punctation close, umbilicate, with a small smooth space on vertex. Pronotum. trapezoidal, scarcely longer than width at anterior angles; gradually narrowed in a straight line to base, more strongly narrowed in front; disk with smooth median line; punctation ason head. Hlytra distinetly longer than pronotum, longer than broad; with moderately coarse and rugulose punctation. Abdomen closely and finely punc- tured. Male, unknown. Female, sternites not modified. Length, 41% mm. (From Cameron.) Type locality —Jamaica, Newcastle district, elevation 3,000 feet. Types.—Unique type (female) in the collection of Dr. Cameron. Records —Yhe following are the records known to me: Jamaica: (Leng and Mutchler, 1917), Newcastle (Cameron, 1913), Blue Mountains (Darlington, in M.C.Z). Specimens examined.—I have seen only the type, which I examined briefly in Dr. Cameron’s collection, and the specimen mentioned below. Remarks.—An example taken by Dr. Darlington in 1934 close to the type locality has not been compared with the type but appears to be this species. It shows the following additional characters: Outer apical corner of elytra indefinitely paler; head scarcely measurably broader than Jong; eyes at about their length from base; labrum with two small median denticles, none lateral; pronotum one-tenth longer than wide, seven-eighths as wide as head; with indistinct shiny area on each side; elytra as long as broad, two-fifths wider than pronotum and two-sevenths longer; punctures indistinct and mixed with flat tuberculi, all obscured by rugulose surface; male, seventh sternite with a semicircular emargination almost as deep as wide, the angles right; eighth with a more open emargination of same size, one-half wider than deep. Length, 4 mm. The key characters were taken from this specimen since the type is not available for further study. It is distinct by its elytral sculpture as well as by the male characters. JT find no record of its habits. 304 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 5. RUGILUS CHTHONUS, new species Description.—Piceocastaneous, shining. Head one-ninth wider than long, not rounded behind although hind angles are broadly rounded; eyes large, at about their length from base; labrum with two large slender but blunt denticles, lateral angles slightly dentate; um- bilicate punctures rather large, often elongate; vertex with an indefi- nite median smooth area; without ground sculpture except vaguely in bottom of punctures. Pronotum one-eighth longer than wide, three- fourths as wide as head; widest at anterior angles, sides distinctly emarginate in front, distinctly arcuate to base; with distinct elevated smooth midline, moderately large and strongly umbilicate punctures dense on either side but leaving an indefinite smooth area on each side behind middle; without ground sculpture. /Jytra as long as wide, scarcely wider than head; conjointly emarginate behind; punctures rather coarse and deep, separated by convex intervals; without ground sculpture. Abdomen with dense fine punctures much obscured by dense and rather indistinct ground sculpture. Male, unknown. Female, eighth sternite prominently rounded. Length 544 mm. Type locality —Puerto Rico, 5 miles north of Villalba. Types.—Holotype, female, U.S.N.M. No. 52410, collected by me on October 12, 1935. Records.—The following are the records known to me: Hispaniola: Hairs, Furey (Mann, in M.C.Z.), LaVisité (Darlington, in M.C.Z. and U.S.N.M.), Desbarriere (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: Villalba (Blackwelder station 64). Specimens examined.—I have seen only the type and five examples from the Museum of Comparative Zoology. Remarks.—This species is distinct from all others now known from the West Indies by the irregular smooth spaces of the pronotum and by its size. As the specimens from Hispaniola differ slightly from the type, they are not made paratypes. They have the punctures of the head less united and not appearing to be arranged in ares; they do not have the pronotal punctures near the midline posteriorly quite so crowded ; and they have the lateral smooth areas smaller, scarcely more than enlarged intervals. The type was collected by sifting dead leaves and trash in a damp gully. All examples were taken at elevations of 3,000 to 7,000 feet. 6. RUGILUS INSULARIS (Cameron) Stilicus insularis CAMERON, 1913b, p. 341.—LeNeG and MutcHter, 1917, p. 198.— SCHEERPELTZ, 1933, p. 1242.—BLACKWELDER, 1939a, p. 107., Description.—Black, shining, elytra sometimes narrowly and indis- tinctly paler along apical margin. Head one-seventh wider than long, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 305 not rounded behind although posterior angles are rounded; eyes large, at about their length from base; labrum minutely bidenticulate, indis- tinctly dentate at outer angles; umbilicate punctures moderate, some- times elongate in front; vertex with a smooth median area; without ground sculpture. Pronotum as wide as long, five-sixths as wide as head; widest at anterior angles, sides distinctly emarginate in front, distinctly arcuate posteriorly; with a distinctly elevated smooth middle band, umbilicate punctures dense, with intervals wider and shiny just behind middle on each side but not forming a distinct smooth area; without ground sculpture. Hlytra scarcely wider than long, a trifle wider than head; conjointly emarginate behind; punctures not coarse but abrupt and elongate, the intervals rather coriaceous. Abdomen moderately sparsely but very finely and submuricately punctured. Male, seventh sternite flattened at center between two small tubercle- like prominences; eighth with a broadly rounded emargination half as deep as wide but very abrupt. Female, eighth sternite rounded. Length, 4 to 4144 mm. Type locality —Jamaica, Chester Vale district. Types.—Unigue type in the collection of Dr. Cameron. Records.—The following are the records known to me: Jamaica: (Leng and Mutchler, 1917), Chester Vale (Cameron, 1913), Spanish Town (Blackwelder station 377), Santa Cruz (Blackwelder station 421), Milk River (Blackwelder station 415), Trinityville (Blackwelder station 428), Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.), Kingston (Darling- ton, in M.C.Z.). Specimens examined.—I have seen the unique type in Dr. Cameron’s collection, 11 specimens from the Museum of Comparative Zoology, and 8 examples collected by Chapin and Blackwelder in February 1937. Remarks.—This species is described from specimens I compared with the type. It is chiefly distinguished by its elytral punctation and male characters. My specimens were caught fiying at dusk. 7. RUGILUS COPROPHILUS (Bierig) Stilicus coprophilus Brmric, 1936, p. 138, pl. 6, figs. 3, 4. Description Rufopiceous, apex of elytra narrowly testaceous, more widely at sides than at suture. Head one-tenth wider than long, rounded behind; eyes large, at less than their length from base; labrum scarcely emarginate, bidenticulate, with a prominent median ridge on upper surface, lateral angles slightly dentate; with rather dense and moderately coarse umbilicate punctures; vertex with an indefinite median smooth area; without distinct ground sculpture. Pronotwm one-eighth longer than wide, two-thirds as wide as head; widest at anterior angles, sides not distinctly emarginate in front, distinctly arcuate posteriorly; with obsolescent umbilicate punctures distinct 306 BULLETIN 182, UNITED STATES NATIONAL MUSEUM along middle smooth space but scarcely discernible toward sides; with- out ground sculpture. //ytra as long as wide, as wide as head, con- jointly emarginate behind; punctures rather feeble and sparse, some- times somewhat muricate. Abdomen sparsely and very finely sub- muricately punctured. Jfale, seventh sternite slightly flattened apically at middle; eighth with a rounded emargination half as deep as wide. Female, eighth sternite rather strongly and prominently rounded. Length, 3 to 314 mm. Lype locality —*Cuba ; probablemente en toda la isla y durante todo elano. Isla de Pinos.” Types.—In the collection of Alexander Bierig. One “cotype” is in the United States National Museum (No. 52724). Records.—The following are the records known to me: Cuba: (Bierig, 1936), Rangel, Sierra del Rosario (Bierig, in U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Isle of Pines: (Bierig, 1936). Specimens examined.—In addition to the “cotype” I have seen 11 examples in the United States National Museum and 8 from the Mu- seum of Comparative Zoology. Remarks.—The above description is taken entirely from the speci- mens in the United States National Museum. The species is distin- guished from all others in the West Indies by its obsolete pronotal punctation. J find no record of its habits. XLI. Genus SCOPOBIUM Blackwelder Scopobium BLACKWELDER, 1939a, pp. 87, 107, 121. Genotype.—O phiomedon anthracinum Cameron=Scopobium an- thracinum (Cameron) (original designation). Diagnosis—Punctures dense and umbilicate; antennae normal; lab- rum broadly semicircularly emarginate; fourth segment of maxillary palpus acicular, longer than greatest width of third; gular sutures very approximate throughout their length; neck about one-half as wide as head; prosternum not dilated under coxae; hypomera pro- longed in a lobe partly behind the coxae; anterior coxal cavities en- tirely open behind; front coxae very large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and sec- ond abdominal sternites absent, third strongly carinate at middle of base; basal half of front tibia with a concavity lined with diagonal ctenidia; front tarsi broadly expanded; apex of posterior tibia with a ctenidium on each side. Remarks.—This is a very distinct genus, which combines certain features of Aderocharis and the Domene-Lathrobium series. It is at present represented by only one species. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 307 1. SCOPOBIUM ANTHRACINUM (Cameron) Ophiomedon anthracinum Cameron, 1913b, p. 344.—Lene and MurcHLer, 1917; p. 198.—ScHEERPELTZ, 1938, p. 1247—BrackWeLber, 1939a, pp. 107, 121. Medon anthracinum (Cameron) BLACKWELDER, 1939a, p. 107. Scopobium anthracinum (Cameron) - BLACK WELDER, 1939a, pp. OT 121s Description.—Piceorufous. Head quadrate, not narrowed behind; eyes at nearly twice their length from base; with very dense umbilicate punctures; labrum with a large semicircular emargination. Pronotwm as broad as long, scarcely narrowed posteriorly, the basal angles some- what rounded; with a distinct impressed midline posteriorly; with dense umbilicate punctures a little smaller than those on head. //ytra two-sevenths wider than long, longer than pronotum; with moderately coarse punctures separated by less than their diameter; intervals with very vague traces of ground sculpture. Abdomen with each segment strongly impressed at base; with fine but feeble submuricate punctures and traces of ground sculpture. Male, seventh sternite with a semi- circular emargination bordered by two ridges which form curved spatulate processes projecting posteriorly ; eighth sternite with a deep emargination parallel-sided near apex and about as broad posteriorly as long. Female, seventh sternite with a median sinuation; eighth sternite rounded. Length, 6 mm. Type locality —Grenada, Chantilly Estate. Types.—In the British Museum. Records.—The following are the records known to me: Bt. Lucia: (Blackwelder station 209A). St. Vincent: (Cameron, 1913; Leng and Mutchler, 1917; British Museum). Grenada: (British Museum). Specimens examined.—I have seen four examples in the British Museum and one collected by me in March 1936. Remarks.—This is a very distinct species, which I made the type of a new genus in a previous paper on the Paederini. It can be recog- nized readily by the large semicircular emargination of the labrum. My example was taken from under stones along a stream. XLII. Genus LATHROBIUM Gravenhorst Lathrobium GRAVENHOorST, 1802, p. 51. Latrobium Dat, 1828, p. 16 (misspelling). Lathobium LxConte, 1859, p. 572 (misspelling). Centrocnemis JOSEPH, 1868, p. 865. Subgenus Tetartopeus C2watina, 1888, p. 349. Subgenus Lethrobium s. str. Litolathra Casry, 1905, p. 71. Subgenus Lathrolepta CAsry, 1905, p. 72. Subgenus Deratopeus Casry, 1905, p. 73. Subgenus Abletobium Casey, 1905, p. 70. Subgenus Apteralium Casry, 1905, p. 70. Subgenus Lathrobiopsis Casey, 1905, p. 72. Subgenus Lathrobioma Casry, 1905, p. 72. 308 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Genotypes.—Staphylinus elongatus Linnaeus= Lathrobium elonga- twm (Linnaeus) (designated by Latreille, 1810; since this is a valid designation, that of des Gozis in 1886 of multipunctum is invalid). Of Centrocnemis, Lathrobium (Centrocnemis) krniense Joseph (monobasic) ; of Litolathra, L. suspecta Casey (designated by Black- welder, 1939) ; of Lathrolepta, Lathrobiwm debilis LeConte=Lathro- lepta debilis (LeConte) (monobasic) ; of Deratopeus, D. parvipennis Casey (designated by Blackwelder, 1939); of Tetartopeus, Lathro- bium terminatum Gravenhorst=Lathrobium (Tetartopeus) termina- tum Gravenhorst (designated by Blackwelder, 1939) ; of Abletobium, A. pallescens Casey (monobasic) ; of Apteralium, Lathrobium brevi- penne LeConte=A pteralium brevipenne (LeConte) (designated by Blackwelder, 1939) ; of Lathrobiopsis, L. texana Casey (monobasic) ; of Lathrobioma, Lathrobium tenue LeConte=Lathrobioma tenuis (LeConte) (designated by Blackwelder, 1939). Diagnosis —Head with punctures not very dense, not coarsely umbilicate; eyes normal; antennae posteriorly flexile, not strongly geniculate, basal segment not very elongate, outer segments normal; labrum bilobed or triangularly emarginate; fourth segment of maxil- lary palpus small, acute, glabrous, not compressed, shorter than greatest width of third; neck over one-fourth as wide as head; pros- ternum not dilated under the front coxae as far as the hypomera; anterior coxal cavities entirely open behind; elytra without a fold above side margin; first and second abdominal sternites absent; apex of posterior tibia with a distinct ctenidium on both sides. emarks.—This genus is distinguished from Lobrathium by the lack of a laterai fold on the elytron. It is a large genus but is repre- sented in the West Indies by only four species, of which three appear to be new. I have seen only five examples. KEY TO WEST INDIAN SPECIES OF LATHROBIUM 1. Punctures on disk of pronotum three times as large as those on head. 2. canoaensis Punctures of pronotum not much larger than those on head__-__-_.____._. 2 2. Hlytra shorter thangpronotimil] 22 es Sees Pee ee Elytra longer than pronotum__ ee ee ee ee ee, CPI tatu 3. Basal angles of head distinet_-___. ft ean ae . 4, subterraneum Basal angles of head broadly rounded____--__-_--__------ 3. dominicanum 1. LATHROBIUM TRINITATUM, new species Description—Testaceous, head and pronotum more or less rufes- cent. Head truncate behind, with hind angles narrowly rounded; with rather feeble punctures not very dense except at base; without ground sculpture except near base; gular sutures distinctly separated along middle and equally divergent to base and apex. Pronotum with irregular median smooth area not outlined by a distinct series MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 309 of punctures; the moderate punctures evenly scattered over the rest of the surface, usually separated by less than their diameter; without ground seulpture. “lytra conjointly longer than wide, wider than pronotum; punctured similarly to the pronotum, a iittle more sparsely on the disk near base; with a vague suggestion of ground sculpture. Hind tarsus with first segment scarcely longer than second. Male, unknown. Female, apical sternites not modified. Length, 644 mm. Type locality—Trinidad, Cedros, on southwest peninsula. Types.—Holotype, female, in Museum of Comparative Zoology, collected on May 4, 1929, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Trinidad: Cedros (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species belongs in the subgenus Lathrobium s.str. It is the only species known from the southern islands and is a normal appearing Lathrobium as distinguished from the subter- ranean L. dominicanum and L,. subterrancum. I have received no record of its habits. 2, LATHROBIUM CANOAENSIS (Bierig) Tetartopeus canvaensis Brerie, 1984f, p. 325, fig. 2. Description.—Black, elytral suture and humers! edge obscure cas- taneous, outer apical angles irregularly luteous. Head as wide as long, sides completely rounded into base; eyes moderate, separated from base by about one-half more than their length; punctures vary- ing from very small to very large, usually umbilicate; with distinct ground sculpture only basally. Pronotum nearly one-fifth longer than wide; sides nearly straight and feebly converging posteriorly, the angles broadly rounded; with very narrow and irregular smooth midline, feebly channelled posteriorly ; punctures on the disk umbili- cate and three times as large as many of those on head, not arranged in series, usually separated by less than their diameter. //ytra nor- mal, with punctures half as large as those on pronotum, without distinet ground sculpture. Hind tarsus with first segment shorter than second. Male, fifth, sixth, and seventh sternites depressed, eighth with a narrow rounded emargination one-half deeper than wide. Female, sternites not specially modified. Length, 714 mm. Type locality —Cuba, Laguna La Canoa, province of Pinar del Rio. Types.—Unique holotype, female, in collection of Alexander Bierig. Records—The following are the records known to me: Cuba: Laguna La Canoa ( Bierig, 1934). North America: Frorima (Bierig, in U.S.N.M.). Specimens cvramined—\ have seen only the example mentioned below. a0 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—One male specimen from Florida was sent to the Na- tional Museum as a “cotype” of canoaensis (U.S.N.M. No. 52726). It seems to agree with the description in most points but cannot possibly be any kind of primary type because the original descrip- tion specifically states that only a single specimen was known. I have not been able to identify this specimen (or species) with any of the described North American species. The above description was drawn from the Florida specimen. The type was found in a crevice in mud. 3. LATHROBIUM DOMINICANUM, new species Description—Piceous, often rufescent at borders of sclerites. Head with hind angles broadly rounded; with moderate punctures sparse on the disk and seldom separated by less than their diameter laterally; without ground sculpture; gular sutures well separated, diverging from near the front. Pronotwm with median smooth area outlined by two regular series of about fourteen punctures, the series sometimes interrupted near base by duplication of punctures; lat- erally with an arcuate series and a few scattered punctures; without ground sculpture. Hlytra very short, appressed; with indistinct and irregular submuricate punctures not regularly serial; without dis- tinct ground sculpture. Hind tarsus with first segment scarcely longer than second. J/ale, seventh sternite unmodified; eighth with a small triangular emargination, twice as wide as deep, with the angles rounded and apex acute. Female, unknown. Length, 6 mm. Type locality—Dominican Republic, Loma de la Pena, northwest of Constanza. Types.—Holotype, male, Museum of Comparative Zoology, and paratype, male, United States National Museum (No. 52497) ; col- lected in August 1938 by Dr. P. J. Darlington at elevation of 5,000 feet. Records.—The following is the only record known to me: Hispaniola: Dominican Repusic, Loma de la Pefia (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This is one of the subterranean type of paederid so distinct as a group and among themselves as species. It is readily distinguishable by the characters given in the key. It belongs in the subgenus Apteralium, although differing con- siderably in facies from the type species. I have received no records of its habits except that it was taken at about 5,000 feet elevation, probably in ground cover. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 3li 4. LATHROBIUM SUBTERRANEUM, new species Description.—Piceorufous. Head with hind angles narrowly rounded; with moderately coarse punctures sparse on the disk and often separated by less than their diameter laterally; without ground sculpture; gular sutures distinctly separated in front, widely divergent behind. Pronotuwm with median area outlined by two regular series of about 18 very fine punctures; laterally with an arcuate series and a few scattered punctures; without ground sculpture. Hlytra very short, appressed ; with indistinct and irregular submuricate punctures not serial; without distinct ground sculpture though the punctures are somewhat obscured by the uneven surface. Hind tarsus with the first segment distinctly longer than the second. Jfale, unknown. Female, apical sternites not modified. Length, 6 mm. Type locality —Puerto Rico, El Yunque, 3,000 feet elevation. Types.—Holotype, female, in the Museum of Comparative Zoology, collected in May 19388 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Puerto Rico: KH] Yunque (Darlington, in M.C.Z.). Specimens examined.—l\ have seen only the holotype. Remarks.—This is a very unusually slender species of the sub- terranean type. It is very distinct by the characters given in the key and belongs in the subgenus Apteralium, where it is similar to dominicanum but different from the type species, brevipennis. It is the only specimen of this genus so far found in Puerto Rico. It was found at 3,000 feet elevation, probably under ground cover. XLUI. Genus LOBRATHIUM Mulsant and Rey Lobrathium MuLsant and Rey, 1878, p. 78. Bathrolium Gozis, 1886, p. 14. Lathrobiella Casry, 1905, pp. 75, 133. Lathretaxis Casty, 1995, pp. 74, 122. Subgenus Platydomene GANGLBAUER, 1895, pp. 504, 507. Subgenus Hulaihrobium Casey, 1905, pp. 73, 115. Lathrotropis Casey, 1905, pp. 74, 115. Subgenus Pseudolathra Casey, 1905, pp. 74, 129. Linolathra Casry, 1905, pp. 75, 131. Microlathra Casey, 1905, pp. 75, 142. Paralathra Casty, 1905, pp. 75, 130. Genotypes.—Lathrobium multipunctum Gravenhorst=Lobrathium. multipunctum (Gravenhorst) (designated by Blackwelder, 1939). Of Lathrobiella, Lathrobium collare Erichson—Lathrobiella collaris (Erichson) (designated by Blackwelder, 1939); of Bathrolium, Staphylinus punctatus Yourcroy=Lathrobium punctatum (Four- croy)=Bathrolium punctatum (Fourcroy) (implied by Gozis, 1886) ; B12 BULLETIN 182, UNITED STATES NATIONAL MUSEUM of Lathrotaxis, Lathrobium longiuscula Gravenhorst=Lathrotaxis longiuscula (Gravenhorst) (designated by Blackwelder, 1939); of Platydomene, Lathrobium bicolor Erichson=Platydomene bicolor (Erichson) (designated by Blackwelder, 1939); of Hulathrobiwm., Lathrobium grande WLeConte=Fulathrobium grande (UeConte) (monobasic); of Lathrotropis, Lathrobium jacobinum LeConte= Lathrotropis jacobina (LeConte) (designated by Blackwelder, 1939) ; of Pseudolathra, Lathrobium anale LeConte=Pseudolathra analis (LeConte) (designated by Blackwelder, 1939) ; of Linolathra, L. fili- tarsis Casey (designated by Blackwelder, 1939); of Micro/uthra, Lathrobium pallidula YWeConte=Microlathra pallidula (“LeConte) (designated by Blackwelder, 1939) ; of Paralathra, P. filicornis Casey (monobasic). Diagnosis.—Body elongate, parallel, feebly constricted ; integuments moderately sparsely punctate, not wmnbilicate; eyes present and nor- mal; antennae posteriorly flexile, not strongly geniculate, basal seg- ment not very elongate, first and second not much larger than rest; labrum bilobed; fourth segment of maxillary palpus acicular, shorter than the greatest width of the third segment; gular sutures separate, variable in position; neck fully one-half as broad as head; prosternum not dilated under coxae; hypomera feebly lobed behind the coxae; anterior coxal cavities open behind; front coxae large, exserted; elytra with a longitudinal fold above side margin; middle coxal cavities con- fluent; posterior coxae contiguous, “conical”; first and second abdomi- nal sternites absent; seventh sternite of male sometimes modified, eighth always emarginate; basal half of anterior tibia with a con- cavity lined with diagonal ctenidia; apex of posterior tibia with a distinct etenidium along both inner and outer edges. Remarks.—This genus has generally been considered a subgenus of Lathrobiwm. Tt was raised to the rank of genus, distinguished from Lathrobivm by the presence of the elytral fold and containing several subgenera (see Blackwelder, 1939). It includes most of the previously described members of the Lathrobia from the West Indies. I have seen 87 examples of this genus from the West Indies. These are assigned to ten species, of which five are new. KEY TO WEST INDIAN SPECIES OF LOBRATHIUM 1. E\lytra:. normal: not}shorter,than) pronotwme=_ 2223-2. eee 2 Elytra reduced, distinctly shorter than pronotum___----------------____ 8 2. Median space of pronotum outlined by two definite series of punctures (sometimes sirrecular) M3 ee ee a A ee 4 Median space outlined at most by bands of punctures not arranged in BING 16, SOTIOS S22 22.0 Mog es 2A sgh aig Dea 5 ek aie oe es ee eel 3 5. Seventh sternite of male longitudinally suleate; piceous, with apex of elytra Purescembte! 174) Pyar atv eps s 3 EE SS See eee 1. discolor Seventh sternite of male not suleate; black, with elytra rufous. 2. pectorale MONOGRAPH OF WEST INDIAN STAPHYLINIDAE B13 4. Gular sutures most approximate along middle__________---_______ 7. odium Gularssutures diverging) from) before! middles<2 2 4. ets ek ee 5 §. Elytra with two or three discal series of punctures_____________________ é Elytra with four or five discal series of punctures______________ 3. rubidum 6. Seventh sternite of male with angles of emargination acutely toothed; outer apicalsthird: or elytra paleris 22s sis) eet esee) ae ee See 6. darlingtoni Seventh sternite} of male moti toothediss 4s sek ee Sk 7 7. Piceous or rufopiceous, elytral apex indefinitely paler______ 5. margipallens Rufous, elytra picescent with apical half or two-fifths testaceous. 4. nitidum §. Pronotum with two regular series of punctures____.______________-______ 9 Pronotum without distinct single series______________-_____ 10. pluviosum oF Dorsal series: with 6) punctures eee ss ee eee eel Se Pees 9. borinquense Donsalaseries: withyaboutel2 punctures. = 8. reductum 1. LOBRATHIUM DISCOLOR (Erichson) Lathrobium discolor Ertcuson, 1840, p. 598.—BreRNHAUER and ScHusert, 1912, p. 258. Description.—Nigropiceous to rufous (immature), apical two- thirds of elytra generally rufopiceous (always more rufous than remainder). Head with sparse moderate punctures separated by one to three times their diameter; without ground sculpture; gular su- tures moderately widely separated, parallel throughout. Pronotum with sparse moderate punctures; median smooth stripe irregular, not bordered by a regular series of punctures; lateral punctures not more than partly in series; without ground sculpture. //ytra with mod- erately coarse but rather shallow and indefinite punctures in irregular series, one sutural, about four discal, and two posthumeral; without distinct ground sculpture. Hind tarsus with first segment slightly shorter than second. Afale, seventh sternite with a longitudinal con- cavity in apical two-thirds, impunctate except at sides, the edges abrupt and elevated, the segment vaguely emarginate at center; eighth sternite with an ogival emargination two and one-half times as long as wide, the apex narrowly acute, the angles not rounded; sixth and even fifth sternites sometimes with a trace of median im- pression. Female, sternites not modified. Length, 7 to 8 mm. Type locality — America meridionalis ins. St. Thomae.” Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Cuba: Soledad (Darlington, in M.C.Z. and U.S.N.M.), Aguadores, in Oriente (Darlington, in M.C.Z.). Hispaniola: Harri, Mtang Lachaux (Darlington, in M.C.Z.); Dominican Re- PUBLIC, Barahona (Darlington, in M.C.Z.), Sanchez (Darlington, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z.). St. Thomas: (Erichson, 1840). {South America: (Bernhauer and Schubert, 1912, by error).] 449008—42——_21 314 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen 23 examples collected by Dr. Darlington, of which 8 are now deposited in the United States Na- tional Museum. Remarks.—The male genitalia leave little doubt of the identity of this species, only odiwm being at all similar. That species is dis- tinguished only by its single series of punctures bordering the prono- tal median space, whereas in discolor the space is irregular and marked by bands of punctures not in a single series. The two species occur in the same part of Cuba, but I find no intergrades in either series. They both belong in the subgenus Lobrathium s. str. I find no record of the habits of this species. 2. LOBRATHIUM PECTORALE (Erichson) Lathrobium pectorale ERIcHSoN, 1840, p. 596.—BERNHAUER and SCHUBERT, 1912, p. 265.—LeEnG and MutTcHtER, 1914, p. 405. Description.—Nigropiceous, elytra rufous. Head more densely punctate at sides, smooth in front, punctures notably sparse. Prono- tum densely punctate at sides, median longitudinal space smooth. Elytra more densely but less strongly punctate, subserially in front. Male, seventh sternite broadly feebly emarginate; eighth sternite deeply triangularly excised. Female, wnknown. Length, 6 mm. (From Erichson.) Type locality —‘Americae meridionalis ins. St. Thomae.” Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following is the only record known to me: St. Thomas: (Erichson, 1840; Bernhauer and Schubert, 1912; Leng and Mutchler, 1914). Specimens examined.—I have seen no example of this species. Remarks.—The characters given above seem to be sufficient to dis- tinguish this species from any known to me. The male characters distinguish it from odiwm. It is not possible to determine from the original description to which subgenus it belongs. I find no record of its habits. 3. LOBRATHIUM RUBIDUM (Fauvel) Tithocharis rubida Fauver, 1863, p. 486.—BrackweELpEr, 1939a, p. 110. Lathrobium rubidum (Fauvel) Virrapa, 1901, p. 31.—BrrnHAvER and Scuu- BERT, 1912, p. 266.—LENG and MurcHLer, 1914, p. 405.—BLAcKWELDER, 1939a, p. 110. (Not Casey, 1905.) Lobrathium rubidum (Fauvel) BLAcKWELDER, 1939a, p. 110. Description.—Rufous to rufotestaceous, abdomen generally darker, elytra sometimes infuscate basally. JZead with large and smaller punctures separated by one to three times their diameter; without ground sculpture; gular sutures rather widely separated, most ap- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 315 proximate before middle, rather widely diverging posteriorly. Pro- notum with median smooth space outlined by two irregular series of about 12 punctures, generally separated by about their diameter, between these and margin are scattered punctures not arranged in series but extending throughout the length; without ground sculpture. Elytra with rather feeble punctures in series, one sutural, four discal, and several indefinite ones posthumeral; without ground sculpture. Hind. tarsus with first segment very little longer than second. Male, seventh sternite unmodified, eighth with a triangular emargination one-half wider than deep, with apex narrowly rounded and lateral angles broadly rounded. Female, eighth sternite shghtly prominent at middle. Length, 4 to 414 mm. Type locality.—Cuba. Types.—Presumably in collection of Blanche Rancin, Caén, France. Records.—The following are the records known to me: Cuba: (Fauvel, 1863: Bernhauer and Schubert, 1912; Leng and Mutchler, 1914), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Jamaica: Kingston (Darlington, in M.C.Z.). Hispaniola: DoMINICAN REPUBLIC, Sanchez (Darlington, in M.C.Z. and U.S.N.M.), Barahona (Darlington, in M.C.Z.). Specimens examined.—I have seen four examples in the United States National Museum and five collected by Dr. Darlington. Remarks.—This species is distinguished by its large number of discal series of punctures on the elytra, by the arrangement of the punctures on the pronotum, and by the male characters. The de- scription was drawn from the examples in the United States National Museum. It probably belongs in the subgenus Pseudolathra. I find no record of its habits. 4. LOBRATHIUM NITIDUM (Erichson) Lathrobium nitidum Ertcnson, 1840, p. 599.—Lyncuw, 1884, p. 239.—BrRNHAUER and ScuHupert, 1912, p. 264——ScCHEERPELTZ, 1933, p. 1280.—BLACKWEDCDER, 1939a, p. 110. Lobrathium nitidum (Erichson) BLacKWELDER, 1939a, p. 110. Description—Rufous or rufopiceous, elytra piceous with apical third testaceous. Head with moderate punctures separated by one- half to two times their diameter; without ground sculpture; gular sutures rather widely separated, most approximate in front, rather widely diverging posteriorly. Pronotwm with median smooth space outlined by two rather straight series of about ten punctures, not evenly spaced; laterally with an arcuate series of about seven punc- tures and a few between this and margin, but none between the two series or posteriorly; without ground sculpture. FH/ytra with very feeble punctures in series, one sutural, two discal, and one post- humeral; without ground sculpture. Hind tarsus with first segment 316 BULLETIN 182, UNITED STATES NATIONAL MUSEUM a little longer than second, Male, seventh sternite prominently rounded but truncated in the middle third by a broad rounded emargination, the angles rounded; eighth sternite with a triangular emargination four times as wide as deep and preceded by a triangular depression. Female, apical sternites feebly rounded. Length, 314 mm. Type. locality — Columbia.” Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Cuba: Aguadores in Oriente (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Harri, Port-au-Prince (Darlington, in M.C.Z.), Ennery (Darlington, in M.C.Z.) ; DominicaN Repustic, Villa Altagracia (Darlington, in M.C.Z.), Puerto Plata (Darlington, in U.S.N.M.). St. Croix: (Blackwelder station 344). St. Lucia: (H. FE. Box, as Blackwelder station 444G). Specimens exramined.—I have seen the two specimens collected by me in 1935-37 and nine collected by Dr. Darlington. Remarks.—The description is drawn from the two specimens now in the United States National Museum. These agree with the original description of nitidwm but are distinguished from other species by the male characters. The color arrangement of the elytra will at once distinguish it among the West Indian species. It belongs in the subgenus Pseudolathra. My specimens were caught flying at dusk. 5. LOBRATHIUM MARGIPALLENS (DuVal) Lathrobium margipallens DuVAL, 1857, p. 41.—BeRNHAUER, and ScHupert, 1912, p. 263.—LeNe and MuTcHLER, 1914, p. 405.—BLACKWELDER, 1939a, p. 110. Lathrobium marginipallens CHEVROLAT and FAUVEL, 1868, p. 485 (misspelling). Lathrobium insulare CAMERON, 1922, p. 113.—ScHEERPELTZ, 1933, p. 1277. Lobrathium margipallens (DuVal) BLACKWELDER, 1939a, p. 110. Description.—Piceous, more or less rufescent especially in front, elytra with apex narrowly and indefinitely paler. Mead with sparse moderate punctures separated by one to three times their diameter; without ground sculpture; gular sutures moderately separated, most approximate before middle, rather widely divergent posteriorly. Pronotum very sparsely punctate; middle smooth space outlined by two regular series of punctures, varying from 7 to 10 punctures in each series; with a sigmoid lateral series of 7 to 9 punctures, and a few between this and margin; without ground sculpture. Zlytra with rather feeble punctures in series, one sutural, two discal, and one post- humeral; without ground sculpture. Hind tarsus with first segment longer than second. Male, seventh sternite slightly produced in median three-fifths, the process truncate and with a feeble rounded MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 317 emargination at middle; eighth sternite with a rounded emargination three times as wide as deep, the outer angles rounded and somewhat produced, the surface of the segment behind the emargination forming a triangular concavity, which is impunctate but with the usual ground sculpture. Female, seventh sternite not modified; eighth somewhat truncate. Length, 314 to 4 mm. Type locality —Cuba. Of insulare, Jamaica. Types.—The type should be in the British Museum (from Guerin- Méneville via the Sharp collection), but I did not find it there. Of insulare, in the collection of Dr. M. Cameron. Records.—The following are the records known to me: Cuba: (Duval, 1857; Bernhauer and Schubert, 1912; Leng and Mutchler, 1914; British Museum), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Maisi in Oriente (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Cameron, 1922; Cameron collection, as insulare), Black River (Black- welder station 416), Santa Cruz (Blackwelder station 421). Snecimens examined—I have seen one specimen in the British Museum, two in the United States National Museum, seven collected by Dr. P. J. Darlington, and three collected by me in 1935-87. PRemarks.—I am unable to distinguish the Jamaican from the Cuban examples. The description was drawn from a Cuban example in the United States National Museum. This species belongs in the subgenus Pseudolathra, My specimens were taken flying at dusk. 6. LOBRATHIUM DARLINGTONI, new species Description—Rufous, head rufopiceous or picescent, elytra piceous with outer apical third rufotestaceous. MHWead with sparse moderate punctures separated by one to three times their diameter; without ground sculpture; gular sutures moderately separated, most approxi- mate near apex. Pronotwm very sparsely punctate, median smooth space outlined by two regular series of 7 to 10 punctures; lateral] punc- tures not in series; without ground sculpture. Hlytra with moderately coarse punctures in series, 1 sutural, 2 discal, and 1 posthumeral; with- out ground sculpture. Hind tarsus with first segment longer than second. J/ale, seventh sternite with a semicircular emargination in median third, the angles produced into short but acute processes; eighth sternite with a rounded emargination three times as wide as deep, the outer angles rounded and somewhat produced, the surface of the segment behind the emargination forming a triangular concavity which is impunctate but with the usual ground sculpture. Female, sternites not emarginate. Length, 414 mm. Type locality —Trinidad, St. Augustine. Types.—Holotype, male, and two paratypes, male and female, in the Museum of Comparative Zoology ; three paratypes, male, in the United 318 BULLETIN 182, UNITED STATES NATIONAL MUSEUM States National Museum (No. 52502); collected in April 1929 by Dr. P. J. Darlington (and on May 9, 1935, by Dr. N. A. Weber). Records.—The following are the records known to me: Trinidad: St. Augustine (Darlington and Weber, in M.C.Z. and U.S.N.M.). Specimens ecamined.—I have seen only the six types. Remarks.—This species is very distinct by the characters of the sev- enth sternite of the male. It can usually be distinguished also by its coloration. It belongs in the subgenus Pseudolathra, One specimen was collected at light. 7. LOBRATHIUM ODIUM, new species Description.—Piceous or rufopiceous, with elytra rufous at least in apical third. Head with rather large punctures separated by one to three times their diameter; without ground sculpture except in region, about the neck; gular sutures moderately widely separated, nearly parallel, diverging only near base. Pronotum with median smooth space outlined by two rather irregular series of about 12 punctures, the series interrupted near base and apex by duplication of punctures; laterally with scattered punctures of various sizes; without ground sculpture. LJytra with feeble punctures in impressed longitudinal series, 1 sutural, 4 discal, and an indefinite band posthumeral; without ground sculpture. Wind tarsus with first segment a little longer than second. Male, seventh sternite with an abrupt and deep longitudinal concavity in apical two-thirds, the bottom impunctate, the apical edge slightly emarginate; eighth sternite with an abrupt emargination twice as deep as wide, the sides feebly concave, the angle acute. Female, seventh sternite unmodified, eighth rounded. Length, 6 mm. Tupe locality—Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, male, and seven paratypes, U.S.N.M. No. 52423, collected by EK. A. Schwarz. One paratype has been deposited in the British Museum. Lecords.—The following is the only record known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.). Specimens excamined.—I have seen only the nine types. Remarks.—This species is distinct from all others from the West Indies except discolor by the male characters. It is very similar to discolor but seems to differ constantly in the arrangement of the color of the elytra. This is not an important character, but it seems best to keep these separate until examples of discolor from the Virgin Islands can be compared directly with it. Z. odiwm belongs in the subgenus Lobrathium s. str. T have no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 319 8. LOBRATHIUM REDUCTUM, new species Description —Piceous, the head generally black, base of elytra nar- rowly rufescent. Head with rather coarse and sparse punctures; sides behind eyes arcuate and rounded into base without angles; without ground sculpture; gular sutures straight, moderately separated in front and diverging to base. Pronotwm with median smooth space out- lined by two regular series of 12 to 14 punctures, the series sometimes interrupted by a misplaced or doubled puncture; with a few scattered punctures laterally; without ground sculpture. “lytra very small, appressed; with fine and somewhat indistinct submuricate punctures not in series; surface uneven but without distinct ground sculpture. Hind tarsus with first segment not noticeably longer than second. Male, seventh sternite unmodified ; eighth with a very small triangular incision, the angles rounded, the apex acute. Female, sternites unmodi- fied. Length, 6 to 7 mm. Type locality—Dominican Republic, Valle Nuevo, southeast of Constanza, elevation about 7,000 feet. Types.—Holotype, male, and five paratypes, in Museum of Com- parative Zoology; four paratypes in the United States National Museum (No. 52498) ; collected in August 1938 by Dr. P. J. Darlington. PRecords.—The following is the only record known to me: Hispaniola: DomINIcAN ReEpustic, Valle Nuevo (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—t\ have seen only the 10 types. Remarks.—This species is one of a group of three collected by Dr. Darlington that are very distinct by the subterranean habitus. It differs greatly from boringuense and pluviosum in the punctation of the pronotum, but probably belongs, with those two species, to an undescribed subgenus. It was found at elevations of 6,000 to 7,000 feet in cloud-forest, prob- ably in ground cover. 9. LOBRATHIUM BORINQUENSE, new species Description—Rufous, with abdomen more or less piceous. Head above entirely impunctate except for a large setigerous puncture over each eye, a pair between the eyes, and a row of four above the neck; without trace of ground sculpture; gular sutures widely separated, nearly parallel throughout. Pronotum nearly twice as long as head or elytra and twice as long as broad; with median space outlined by two even rows of six punctures; with 7 to 10 scattered punctures laterally ; without trace of ground sculpture. lytra very small, appressed; each with 4 longitudinal series of large punctures, the inner two rows of about 4 punctures, the outer 2 of 6 to 8; without trace of ground sculp- ture. Hind tarsus with first segment about as long as second. Abdo- 320 BULLETIN 182, UNITED STATES NATIONAL MUSEUM men with all visible ventral segments deeply pitted on each side near front, the pit abrupt and flaringly excavated posteriorly, the excava- tion bordered by a distinct cariniform edge which extends half the length of the segment. Male, eighth sternite truncate with posterior border thickened and raised at both sides of center giving appearance of a narrow median groove, apical median portion of segment flattened and smooth. /emale, eighth sternite truncate but otherwise unmodi- fied. Length, 614 to 744 mm. Type locality —Puerto Rico, El Yunque, elevation about 3,000 feet. Types.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology; one paratype, male, in the United States National Museum (No. 52499); collected in May 1938, by Dr. P. J. Darlington, FRecords.—The following is the only record known to me: Puerto Rico: El Yunque (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—lI have seen only the three types. Remarks.—This is a very striking species with a distinctly sub- terranean appearance. The excavations on the abdominal sternites are unique among the staphylinids known to me. It appears to belong in an undescribed subgenus. I have received no record of its habits, but there is a strong pre- sumption that it occurs in ground cover. 10. LOBRATHIUM PLUViIOSUM, new species Description—Rufous, abdomen slightly picescent. Head with moderately large but very sparse setigerous punctures; without trace of ground sculpture; gular sutures rather widely separated and parallel. Pronotum one-half longer than head or elytra, one-fourth longer than wide; with an ee median smooth space not out- lined by series of ae es, punctures sparse and moderate; without trace of ground sculpture. El ytra very small, appressed ; in sparse and rather indefinite punctures not arr aged serially ; rere some- what uneven but without ground sculpture. Hind tarsus with first segment about as long as second. Abdomen without excavations at the sides of the sternites. Afale, eighth sternite with a small rounded emargination and a narrow smooth stripe along the entire midline. Female, eighth segment narrowly rounded and without smooth stripe. Length, 5 to 5144 mm, Type eae i Mominicen Republic, vicinity of Valle Nuevo, at elevation of about 6,000 feet. Types.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology; one paratype, male, from Loma de la Pefia, northwest of Constanza, in the United States National Museum (No. 52500) ; all collected in August 1938 by Dr. P. J. Darlington. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Sok Records.—The following are the records known to me: Hispaniola: DomInicAN Repupiic, Valle Nuevo (Darlington, in M.C.Z.), Con- stanza (Darlington, in U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This is a distinctly subterranean species in appearance. It is-readily distinguished from reductum and borinquense by the absence of serial punctures on the pronotum and elytra as well as by the male characters, but it belongs in the same undescribed subgenus. The types in the Museum of Comparative Zoology were found in cloud forest, probably under ground cover. XLIV. Genus PAEDERUS Fabricius Paederus Fasnrictus, 1775, p. 268. Poederus Lucas, 1857, p. 51. (misspelling). Poederomorphus GAUTIER, 1862, p. 75. Paederomorphus Marsrvt, 1866, p. VI (misspelling). Paederidus Mutsant and Rey, 1877, p. 245. Leucopaederus Casey, 1905, pp. 59, 67. Paederillus Casny, 1905, pp. 59, 62. Subgenus Pseudopaederus BERNHAUER, 1915b, p. 187, Subgenus Gnathopaederus CHapin, 1927, p. 75. Subgenus Paederognathus WENDELER, 1928a, p. 37. Gnathopaederus WENDELER, 1927, p. 1 (not Chapin, 1927). Subgenus Neopacderus BLACKWELDER, 1939a, p. 97. Genotypes.—Staphylinus riparius Linnaeus=Paederus riparius (Linnaeus) (designated in 1810 by Latreille). Of Poederomorphus, P. pedoncularius Gautier (monobasic) ; of Leucopaederus, Paederus ustus LeConte=Leucopaederus ustus (LeConte) (monobasic); of Paederillus, Paederus littorarius Gravenhorst=Paederillus littora- rius (Gravenhorst) (designated by Blackwelder, 1939); of Paederi- dus, Paederus ruficollis Fabricius=Paederus (Paederidus) ruficollis Fabricius (designated by Blackwelder, 1939); of Gnathopaederus Chapin, G. szechuanus Chapin (original designation; monobasic) 5 of Gnathopaederus Wendeler, G. turrialbanus Wendeler (monobasic) ; of Paederognathus, Gnathopaederus turrialbanus Wendeler=Pae- derognathus turrialbanus (Wendeler) (monobasic) ; of Pseudopae- derus, Paederus (Pseudopaederus) nigerrimus Bernhauer (desig- nated by Blackwelder, 1939); of Meopaederus, Paederus morio Mannerheim=Paederus (Neopacderus) morio Mannerheim (origi- nal designation), Diagnosis.—Body elongate, moderately convex; head exserted, con- stricted behind; antennae inserted under sides of front of head, not geniculate but posteriorly flexile, 11-segmented, filiform; labrum short, transverse, emarginate at middle; fourth segment of maxillary palpus short, compressed, pubescent; mentum transverse; submentum trans- O22 BULLETIN 182, UNITED STATES NATIONAL MUSEUM verse; gular sutures converging toward line of constriction from both ends; prosternum short, not carinate, abbreviated between and under coxae forming an acute point which does not attain the mesosternum:; anterior coxal cavities narrowly open behind; front coxae very large, conical, prominent; middle coxal cavities confluent; posterior coxae large, conical, contiguous; abdomen margined; first and second ster- nites absent; eighth sternite excised in male, lobed in female; tarsi 5-segmented, fourth segment bilobed. Remarks.—I have made no changes in this genus except in the char- acters used to distinguish it and its parts. It is one of the most easily recognized of the Staphylinidae. Fifty-four examples from the West Indies are herein assigned to four species, of which one is new. KEY TO WEST INDIAN SPECIES OF PAEDERUS 1. Color uniform rufotestaceous throughout______--__-___---- 3. jamaicensis Color*in' part blue.or black2a2 =a a ee 2 2. Body black throughout=-.<2 2-22. = SS ee ee 4. morio Body. in:part rufous.or testaceous— 222222 es ee ee ee 3 S$: Abdomen: entirely black..8¢ 222. 2223-2 oe es 2. homonymus Base of abdomen testaccouSi=-_==-s—2"ss- 22 2 a 1. mutans 1. PAEDERUS MUTANS Sharp Paederus mutans SHARP, 1876, p. 286.—BERNHAUER and ScHUBERT, 1912, p. 209.— PLAVILSTSCHIKOV, 1929, p. 29.—BLACKWELDeER, 1939a, p. 112. Description.—Rufotestaceous, elytra cyanescent, seventh and eighth abdominal segments black. /Zead with eyes a little wider than long, with moderate but very shallow punctures separated by one to three times their width, each puncture with a long black hair; labrum with a small triangular notch at middle, rather truncate in front with mar- gin indistinctly crenulate. Pvonotum one-sixth longer than wide, wid- est at anterior fourth, distinctly narrowed behind; punctures finer and sparser than on head but with similar hairs. ytva as wide as pro- notum, over one-sixth longer than wide; punctures coarse and deep, separated by less than their diameter by feebly convex intervals; su- ture feebly raised throughout. Abdomen with sparse and fine punc- tures more or less transversely serial, each with a long black hair, submuricate especially on apical segments; with feeble ground sculp- ture. Male, eighth sternite with “the usual excision.” Female, eighth sternite deeply semicircularly emarginate on each side at apex, leaving prominent angles and an acuminate triangular median lobe. Length, 8to 9mm. Type locality —“Tapajos and Para,” Amazon Valley, (Brazil). Types.—In the British Museum. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE a2e Records.—The following are the records known to me: Trinidad: (Bernhauer determination, in British Museum). South America: Braziu (Sharp, 1876), Bottvra, Peru (Plavilstschikov, 1929). Specimens examined.—I have seen the types and 16 other examples from South America and 1 from Trinidad in the British Museum, the latter borrowed for further study. Remarks.—The above description is drawn from the female from Trinidad but does not conflict with the original. I find no record of the habits of this species. 2. PAEDERUS HOMONYMUS, new name Paederus tricolor ERicHson, 1840, p. 663.—SHarpP, 1886, p. 6138.—BERNHAUER and ScHUuBERT, 1912, p, 211.—LENG and MurTcHLErR, 1914, p. 404; 1917, p. 199.— BLACKWELDER, 1939a, p. 112. (Not Fabricius, 1787.) Paederus thoracicus MARSHALL, 1878, p. xxx (not Erichson; not Stephens).— Lene and Mutcuter, 1917, p. 198. Description —Black, prothorax and mesosternum rufotestaceous, elytra aeneocyanescent. Head with eyes and exclusive of neck as wide as long; with moderate and shallow punctures separated by two to three times their diameter; with ground sculpture feeble in front, absent at center of vertex, and strong posteriorly; punctures with moderately long hairs; labrum truncate in front, coarsely crenulate, and with a small quadrate notch at middle. Pronotwm one-fifteenth longer than wide, widest at anterior fourth, distinctly narrowed be- hind; punctures scarcely evident but separated by three to four times their diameter, much denser at sides, only marginal ones bearing hairs; disk with a slight median prominence near base. Elytra as wide as pronotum, one-fourth longer than wide; punctures coarse and deep, with intervals feebly convex. Abdomen with very sparse submuri- cate setigerous punctures and distinct ground sculpture. Male, eighth sternite with a deep parallel rounded excision. Length, 9 to 914 mm. Type locality—*. .. Ins. St. Vincentii Americae ...” (=St. Vincent). Types.—In either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Martinique: (Marshall, 1878; Leng and Mutchler, 1917, as thoracicus). St. Vincent: (Erichson, 1840; British Museum). (Guatemala: (Bernhauer determination, in British Museum; Bernhauer and Schubert, 1912).) Specimens examined.—I have seen only the 12 specimens in the British Museum; 1 of these I borrowed for further study. Remarks.—The one example in the British Museum from Guate- mala was determined by Bernhauer as this species, but my brief exam- 324 BULLETIN 182, UNITED STATES NATIONAL MUSEUM ination of it revealed the following differences: The Guatemalan ex- ample has no ground sculpture on the back of the head, has the elytra more irregular, almost sculptured, and the elytral punctures in irreg- ular grooves but scarcely evident. This specimen was probably the basis for the Guatemalan record in the Bernhauer and Schubert catalog. The specimens recorded from Martinique by Marshall as thoracicus Erichson are undoubtedly not that species but the present one. I find no record of the habits of this species. 38. PAEDERUS JAMAICENSIS, new species Paecderus jamaicensis BERNHAUER, MS. Description—Rufotestaceous throughout. /Zead with eyes a little broader than long; without punctures except for a few large setiger- ous ones; with rather indistinct ground sculpture; labrum with a moderate triangular notch at center, edge somewhat crenate. Prono- tum about one-sixth longer than wide, widest at anterior third, feebly narrowed behind; with a few setigerous punctures especially at the sides. Hlytra at apex as wide as pronotum, closely appressed to body and much narrower at base; with about 10 large setigerous punctures; the surface uneven but not distinctly sculptured. Abdo- men with about 10 large setigerous punctures on each segment; with very vague transverse ground sculpture. A/ale, unknown. Yemale, eighth sternite prolonged into an acutely triangular process. Length, 6 mm. . Type locality —Jamaica. Types.—Holotype, female, in the British Museum. Records.—The following is the only record known to me: Jamaica: (British Museum ace. 45.110). Specimens examined.—I have seen only the unique type. Remarks.—This specimen seems so very distinct from all other species of Paederus known from the West Indies that I describe it in spite of the chance that it is immature. It certainly does not belong to any of the other species listed here. It belongs in the subgenus Neopaederus. I find no record of its habits. 4, PAEDERUS MORIO Mannerheim Pacderus morio MANNERHEIM, 1830, p. 89.—BeERNHAUER and ScHusBERT, 1912, p. 208.—Lene and Murcuter, 1914, p. 404—BLacKWELpeR, 19359a, pp. 112, 120. Description.—Black, elytra aeneous. Head over one-fourth wider than long; with moderately fine but very irregular punctures rather obscured by the dense fine ground sculpture; punctures at sides with short hairs; labrum truncate in front, with a small quadrate notch at middle, sides slightly arcuate and feebly crenulate. Pronotum MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 325 one-eleventh longer than wide, widest at apical fourth, distinctly nar- rowed behind; with scattered punctures similar to head but sparser; ground sculpture distinct but very fine. Zlytra slightly wider than pronotum, three-eighths wider than long; widest at apex, suture barely half as long as total; not distinctly punctate but with a few fine hairs; surface coarsely coriaceous, very uneven. Abdomen with sparse but somewhat transversely serial submuricate punctures; with very fine ground sculpture. Male, seventh sternite longitudinally depressed at middle, very feebly emarginate posteriorly; eighth sternite with a very deep parallel excision rounded at base and with outer angles rounded. Female, eighth sternite with a rounded emar- gination at each side to form prominent outer angles and a rounded median lobe. Length, 7 to 9 mm, Type locality— Insula St. Domingo .. .” (=Hispaniola). Types.—Presumably in Zoologisk Museum, Helsingfors. Records—The following are the records known to me: Hispaniola: (Mannerheim, 1830, as island of “St. Domingo”; Leng and Mutchier, 1914, as island of ‘‘Haiti’) ; Harr1, Morne La Selle (Blackwelder station 22C), Kenskoff (Blackwelder station 23B; Darlington, in M.C.Z. and U.S.N.M.), Furey (Mann, in M.C.Z.) Specimens examined.—I have seen 22 specimens collected by me on Morne La Selle and 1 at Kenskoff, as well as 5 from the Museum of Comparative Zoology. Remarks.—There can be little doubt that these specimens belong to Mannerheim’s species. My specimens were taken at considerable ele- vations, 6,000 feet on Morne La Selle and close to 4,000 feet at Kenskoff. All the previous published records refer to the island as a whole (under the name of Haiti or Santo Domingo), and it is probable that the specimens came from Haiti. If the species is found else- where in the island it will probably be in the north-central moun- tains of Haiti or the higher central range of the Dominican Republic. My examples were all taken while beating trees and shrubs at the edge of the jungle. XLV. Genus HOMOEOTARSUS Hochhuth Homoeotarsus HocHHvuTH, 1851, p. 34. Homaeotarsus LACOoRDAIRE, 1854, p. 89 (misspelling). Spirosoma MOTscHULSKyY, 1858b, p. 206. Subgenus Hesperobiwm Casky, 1886a, p. 33. Subgenus Lucryptina Casry, 1905, pp. 24, 28. Subgenus Gastrolobium Casry, 1905, p. 31. Subgenus Homoeobium BLACKWELDER, 1939a, p. 96, 114, 118. Subgenus Nemoeotus BLACKWELDER, 1939a, p. 96, 114, 120. Genotypes—H. chaudoirt Hochhuth (monobasic). Of Spirosoma, S. fulvescens Motschulsky (monobasic) ; of Lucryptina, Cryptobium 326 BULLETIN 182, UNITED STATES NATIONAL MUSEUM opacum Sharp=Hucryptina opaca (Sharp) (monobasic and original designation); of Gastrolobium, Lathrobium bicolor Gravenhorst= Gastrolobium bicolor (Gravenhorst) (designated by Blackwelder, 1939) ; of Hesperobium, Cryptobium tumidum LeConte= Hesperobium tumidum (LeConte) (original designation) ; of Homocobium, Homo- eotarsus (Homoecobium) bakerianum Blackwelder (monobasic and original designation) ; of Nemocotus, Cryptobium rubiginosum Bern- hauer=Homocotarsus (Nemoeotus) rubiginosus (Bernhauer) (mono- basic and original designation). Diagnosis —Body very elongate, parallel; integuments moderately densely punctate; antennae anteriorly flexile, strongly geniculate, basal segment much elongate; labrum bilobed; fourth segment of maxillary palpus subulate; gular sutures separate; neck more than one-half as wide as head, abruptly constricted across dorsal surface; prosternum not dilated under the coxae; hypomera feebly lobed behind coxae; anterior coxal cavities open behind; front coxae large, exserted; elytron with a longitudinal fold above side margin; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; third longitudinally carinate at base; fourth and fifth in male sometimes with a median fovea, fifth sometimes lobed (males sometimes dimorphic) ; basal half of anterior tibia with a concavity lined with diagonal ctenidia, with a correspond- ing prominence on the femur; apex of posterior tibia with a ctenidium on each side. Remarks.—This genus is distinguished by the structure of the an- tennae and the presence of the pleural elytral fold. It is rather variable in the secondary male characters, and the subgenus G'astrolo- bium, to which all our species belong, is highly developed in this respect. At least some of the species have dimorphic males. The name given above to this genus has not previously been used in connection with New World faunas (except temporarily by Casey, 1889a, p. 182). The division of the genus Cryptobium on the char- acter of the elytral fold leaves us with four subgeneric names, of which one must be the generic name as well. The oldest name is Homoeotarsus and must be accepted on that basis. I have seen 187 specimens from the West Indies. These I assign to 4 species, of which 2 are new. KEY TO WEST INDIAN SPECIES OF HOMOEOTARSUS 1. Head with ground sculpture throughout___--___-____--_____-_ sete ees Vertex between eyes without ground sculpture__-_-__-_--_--_--__- 1, solus 2. Pronotum with dense ground sculpture___.._____.___-__-_-__ 4. testaceipes Pronotum* without: ground Sculpture: 2 te ee ees ae ee 33 5. Fovea of fifth sternite obsolescent, very shallow__------__-- 3. obsolescens Foven of fifth sternite distinct, deep, usually oval and strongly concave. 2. albipes MONOGRAPH OF WEST INDIAN STAPHYLINIDAE oot 1. HOMOEOTARSUS SOLUS, new species Description—Uniform black to rufopiceous throughout, the pro- notum sometimes with a blue-green tinge. Mead with coarse punc- tures scarcely distinctly separated near base, each puncture well defined posteriorly but gradually excavated in front, almost absent near clypeus, very scattered between eyes; ground sculpture visible within the punctures basally and laterally, but area between and in front of eyes entirely smooth, shining; eyes in front of middle, separated by more than twice their length from base; basal antennal segment not quite as long as distance between antennae. Pronotwm with narrow impunctate median stripe, rest with coarse punctures usually separated by less than half their diameter (more irregular at sides) ; without ground sculpture. “lytra with coarse punctures separated by about half their diameter by convex intervals, with no trace of serial arrangement; without ground sculpture. J/ale, fourth sternite with a distinct transverse setigerous fovea; fifth with a small circular fovea and usually with a posterior lobe one-fourth as broad as segment extending past middle of sixth sternite and with its apex truncate; sixth and seventh sternites impunctate along middle. Female, without modifications of the sternites. Length, 6 mm. Type locality—Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, female, U.S.N.M. No. 52589, collected on Janu- ary 18 by E. A. Schwarz. Also 24 paratypes in the United States National Museum and the Museum of Comparative Zoology. Records—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Pico Turquino (Darlington, in M.C.Z. and U.S.N.M.) Gran Piedra Range, Oriente (Darlington, in M.C.Z. and U.S.N.M.), mountains north of Imias, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Loma del Gato, Cobre Range, Oriente (Darlington, in M.C.Z and U.S.N.M.). Jamaica: Cinchona (Darlington, in M.C.Z.). Hispaniola: Hattr, northeast foothills of the Massif de La Hottie (Darlington, in M.C.Z. and U.S.N.M.); Dominican REPUBLIC, Constanza (Darlington, in M.C.Z. and U.S.N.M.), foothills south of Santiago (Darlington, in M.C.Z.). Specimens examined.—I have seen only the 25 types. Remarks.—This is a very distinct species because of its uniform color and the unsculptured interocular space of the head. It belongs in the subgenus Gastrolobium. I have no record of its habits except that it is frequently found at elevations up to 5,000 feet. 328 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. HOMOEOTARSUS ALBIPES (Erichson) Cryptobium albipes EricHson, 1840, p. 566.—BERNHAUER and ScHUBERT, 1912, p. 279.—LeNe and MuTCHLER, 1914, p. 405.—WotcortT, 1924, p. 78; 1936, p. 197.— BLACK WELDER, 1939a, p. 113. Cryptobium marginellum BERNHAUER, 1908b, p. 322.—BrERNHAUER and SCHUBERT, 1912, p. 284.—Lene@ and MutcuHrer, 1914, p. 405. Homocotarsus albipes (Erichson) BLACKWELDER, 1989a, p. 113. Description —Pitchy black with apex of elytra and abdominal ter- gites narrowly testaceous, varying to castaneous with head piceous and pronotum rufous. Zead with moderately coarse and dense punc- tures generally separated by less than their diameter (basally and laterally by less than half their diameter); punctures absent near clypeus but scattered between eyes; with dense and distinct ground sculpture throughout; eyes in front of midline, at twice their length from base; basal segment of antennae not quite as long as distance between antennae. Pronotwm with narrow impunctate median stripe, rest with coarse punctures separated by half their diameter or less (a little more at sides); without ground sculpture. Llytra with coarse punctures separated by about one-fourth their diameter by convex intervals, with no trace of serial arrangement; without ground sculpture. Jfale dimorphic, always with a well-developed transverse fovea on fourth and fifth sternites, fifth sternite with or without a broad rounded posterior lobe extending to the seventh sternite. Female without foveae or other modifications of the sternites. Length, 7 to 8 mm. Type locality—Puerto Rico. Of marginellum, St. Vincent. Types.—In either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of marginellum, “In der Sammlung des k. k. natur- historischen Hofmuseums in Wien und in meiner eigenen” (Bern- hauer). Records.—The following are the records known to me: Cuba: Cayamas (Schwarz in U.S.N.M.), Cauto El Cristo, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Soledad (Darlington, in M.C.Z and U.S.N.M.), Imias, Oriente (Darlington, in M.C.Z.), Baragua (Scaramuzza, in M.C.Z.), Aguadores, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Marcata, Rio Bayamo (Darlington, in M.C.Z.). Jamaica: Kingston (Blackwelder station 379; Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Harrt, Ennery (Darlington, in M.C.Z. and U.S.N.M.), Plaine de l’Artibonite (Darlington, in M.C.Z. and U.S.N.M.); DominicAn REPUBLIC, Barahona (Darlington, in M.C.Z.). Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914, as both; Wolcott, 1924, 1936), Bayamon (Lesesnes, in U.S.N.M.). St. Vincent: (Bernhauer, 1908, as marginellum; Leng and Mutchler, 1917, as marginellum; Blackwelder stations 166, 170, 176A). Grenada: (British Museum). South America: VENEZUELA, COLOMBIA (Bernhauer and Schubert, 1912). Central America: GUATEMALA, Mexico (Bernhauer and Schubert, 1912). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 329 Specimens examined.—I have seen 23 examples in the British Mu- seum, 31 from the Museum of Comparative Zoology, 3 in the United States National Museum, and 59 I collected during 1935-37. Remarks—TVhis is a variable species both in color and puncta- tion. The latter varies in arrangement and somewhat in density, but very little in the size of the punctures. The dimorphism of the male is similar to that observed in North American species of Gas- trolobium, to which subgenus this species belongs. I have collected this species from under stones at the edge of streams. 3. HOMOEOTARSUS OBSOLESCENS, new species Description—Black, anterior and posterior margin of pronotum and elytra more or less rufescent. Mead with moderate punctures, each more abrupt posteriorly, narrowly separated basally but by about their diameter on vertex and still more sparsely in front; with very dense and very strong ground sculpture throughout; eyes in front of midline, at over twice their length from base; basal segment of antennae not quite so long as distance between antennae. Pronotum with irregular median smooth stripe, rest with coarse irregular punc- tures generally very dense but sparser at sides; without ground sculpture. Llytra with coarse deep punctures separated by convex intervals, with no trace of serial arrangement; without ground sculp- ture. Jfale dimorphic, fourth sternite with a distinct fovea, fifth with only a trace of a setigerous fovea, not lobed, or with a broad rounded triangular lobe reaching to the seventh sternite. Female, sternites not modified. Length, 7 to 814 mm. Type locality Trinidad, 3 miles northeast of St. Augustine on the Tacarigua River, county of St. George, ward of Tacarigua. Types.—Holotype, male, and one paratype, female, U.S.N.M. No. 52425, collected by me on December 22, 1935; two paratypes in the British Museum, one from Caparo and one labeled merely Trinidad; two paratypes from Tobago in the United States National Museum, and five paratypes from the Museum of Comparative Zoology. Records.—The following are the records known to me: Tobago: (Blackwelder station 119). Trinidad: (Birch, in B.M.), Tacarigua (Blackwelder station 107A), Caparo (Klages, in B.M.), St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen two specimens in the British Museum, five from the Museum of Comparative Zoology, and four collected by me in 1935-87, Remarks.—The examples in the British Museum were labeled Cryptobium marginellum Bnhr. and Cryptobiwm assimile Er., the former apparently in the handwriting of Dr. Bernhauer. They ap- pear to me to differ from the descriptions of those species and to 449008—42 22 330 BULLETIN 182, UNITED STATES NATIONAL MUSEUM agree with my Trinidad specimens. The obsolete fovea of the fifth sternite of the male is quite distinctive, regardless of the presence or absence of the lobe. This species belongs in the subgenus Gastrolobium. My specimens were found beneath stones at the edge of streams. 4. HOMOEOTARSUS TESTACEIPES (Bierig) Gastrolobium testaceipes Birrie, 1935b, p. 39, pl. 4, figs. 5, 6. Gastrolobiwm congener BieRrIc, 1935b, p. 40. Description —Black, sometimes more or less rufescent. Head with moderate punctures, each more abrupt posteriorly, narrowly sepa- rated basally but by about their diameter on vertex and still more sparsely in front; with very dense and strong ground sculpture throughout; eyes in front of midline, at over twice their length from base; basal segment of antennae not quite so long as distance between antennae. Pronotwm with irregular impunctate median stripe, rest with moderate umbilicate punctures, usually separated by more than their diameter and somewhat obscured by the coarse dense ground sculpture. lytra with moderately coarse deep punctures, abrupt only in front, the intervals flat, without serial arrangement; surface exceedingly minutely coriaceous. J/ale dimorphic, fourth sternite with a transverse fovea; fifth with a small circular fovea, with or without a narrow acuminate lobe, which extends to the seventh ster- nite. Memale, sternites not modified. Length, 8 to 10 mm. Type locality—Cuba, “Cercania de Guanabo; Caimito del Guaya- bal.” Of congener, Cuba, “Sierra del Rosario, Rio Taco-Taco.” Types.—In collection of Alexander Bierig (for both). (Three “co- types” of congener are in the United States National Museum, No. 52727.) Records.—The following are the records known to me: Cuba: Guanabo (Bierig, 1935), Caimito del Guayabale (Bierig, 1935), Sierra del Rosario (Bierig, 1935, as congener), Aspiro (Bierig, in U.S.N.M., as congener), Buenos Aires, Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.), Maisi, Oriente (Darlington, in M.C.Z. and U.S.N.M.), Rangel Mountains, Pinar del Rio (Darlington, in M.C.Z.), Soledad (J. G. Myers and Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Harri, EHonery (Darlington, in M.C.Z. and U.S.N.M.), mountains near Port-au-Prince (Darlington, in M.C.Z. and U.S.N.M.), northeast foothills of the Massif de La Hotte (Darlington, in M.C.Z. and U.S.N.M.), Camp Perrin (Darlington, in M.C.Z.) ; DominicAN REPUBLIC, Villa Altagracia (Darlington, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.), Constanza (Darlington, in M.C.Z.). Specimens examined.—Beside the paratypes of congener, I have seen 33 examples from the Museum of Comparative Zoology. Remarks.—Although IT have not seen examples identified by Bierig as testaceipes, the three examples of congener, in conjunction with the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 331 original descriptions and a series from various parts of Cuba, lead me to believe that the two species cannot be distinguished. The relative widths of the pronotum and elytra, as recorded, do not apply to these “cotypes,” and the remaining characters appear to me to be of very much less than specific importance when we take into consideration the variation found throughout this genus. Tn addition to the male characters this species differs from albipes in the presence of ground sculpture on the pronotum and in other details. The species is recorded from the edge of water, beneath stones and decaying plant debris. XLVI. Genus CCHTHEPHILUM Stephens Ochihephilum STEPHENS, 1829, p. 287. (Not Ochthephilus Mulsant and Rey, 1856.) Cryptobium MANNERHEIM, 1830, p. 38. Ochtephilum STEPHENS, 1832, p. 200 (misspelling). Octhephilum LaAcorparme, 1854, p. 89 (misspelling). Octephilum BERNHAUVER and ScHUBERT, 1911, p. 92 (misspelling). Subgenus Ababactus SHARP, 1885, p. 533. Subgenus Cryptobiella CAsEy, 1905, p. 29. Subgenus Asienobium BERNHAUER, 1911, p. 411. Subgenus Neobactus BLACKWELDER, 1989a, p. 96. Genotypes.—Paederus fracticornis Paykull=Ochthephilum fracti- corne (Paykull) (monobasic). Of Cryptobium, Paederus fracticornis Paykull=Cryptobium. fracticorne (Paykull) (monobasic) ; of Aba- bactus, A. depressus Sharp (designated by Blackwelder, 1939); of CUryptobiella, C. colonica Casey (original designation; the designation of C. rostratum Sharp by Bierig, 1935, is not valid); of Astenobiwm, A. excellens Bernhauer (monobasic) ; of Veobactus, Cryptobium (Neo- bactus) nunenmacheri Blackwelder (monobasic and original desig- nation). Diagnosis.—Body moderately elongate, not distinctly parallel; in- teguments strongly punctate, often without ground sculpture; antennae anteriorly flexile, strongly geniculate, basal segment much elongate ; labrum bidenticulate; fourth segment of maxillary palpus subulate; gular sutures separate; neck more than one-fourth as wide as head; prosternum not dilated beneath the coxae; hypomera feebly lobed be- hind the coxae; anterior coxal cavities open behind; front coxae large, exserted ; elytra without pleural fold; middle coxal cavities confluent ; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, third carinate at base; fourth and fifth sternites of male not modified; basal half of anterior tibiae without diagonal ctenidia; apex of posterior tibia with a ctenidium on each side. Remarks.—This genus was restricted by Casey to the small European species. It is apparent from a study of the genotypes of this and Casey’s new genera that they are not all of the same value. Cryptobium 332 BULLETIN 182, UNITED STATES NATIONAL MUSEUM is therefore restored to the American lists with three subgenera, and the remaining names are grouped under Homocotarsus and Biocrypta. Cryptobium centrale Sharp of Central and South America is listed by both Bernhauer and Schubert and Leng and Mutchler as occurring in Guadeloupe. It is at least likely that this is a misidentification, but I am not able to say to which of our species (or genera) it really pertains. Inasmuch as the name Ochthephilum has priority over Cryptobium (and the two are isogenotypic), the former must be accepted as the valid name for this genus. J have seen only three examples of this genus from the West Indies. They belong to two new species, and one other species has been recorded from Cuba and one from Trinidad. KEY TO WEST INDIAN SPECIES OF OCHTHEPHILUM 1. Eyes at not over twice their length from base; basal segment of antenna longer than: interantennal «distances +_-=—=— ee Be ee i 15, Eyes at four times their length from base; basal segment of antenna as long aginterantennalldistance 22 a ee ee 3. darlingtoni 2. Basal segment of antennae twice as long as interantennal distance; head very strongly narrowed) infrontsc == - ee eee 2. trinitatum Basal segment of antennae one-half longer than interantennal distance; head feebly narrowed «ine fronts 22. SS eee eee see 1. atlanticum 1. OCHTHEPHILUM ATLANTICUM (Bierig) Cryptobiella atlantica Birerie, 19385b, p. 41, pl. 4, figs. 8, 9, 10. Description.—Piceous, anterior and posterior margin of pronotum rufescent, apex of elytra testaceous. Head not greatly narrowed in front of eyes; punctures moderately large, umbilicate, clearly im- pressed on disk, somewhat sparse on clypeal area, intervals on vertex narrower than punctures; with dense ground sculpture; with a feeble epicranial depression with clustered punctures; eyes at or a little be- hind the middle; basal seoment of antennae about four-fifths as long as head but only one-half longer than distance between antennae; an- tennal grooves distinct; labrum scarcely emarginate and with two — minute denticles; neck one-half as wide as head. Pronotwm with punc- tures denser than on head, somewhat vague and subserial; with a broad smooth median band, canaliculate posteriorly; with ground sculpture. Male, seventh sternite feebly emarginate and with disk impressed; eighth with a narrow excision nearly three times as deep as width at apex. Length,5to6mm. (From Bierig.) Type locality—Cuba, “Embocadura del Rio Cojimar: Playa de Marianao.” Types.—In collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: Rio Cojimar, Playa de Marianao (Bierig, 1935). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 330 Specimens eramined.—ti have seen no examples of this species. Remarks-—The differences between this and trinitatwm are listed under the latter species. Certain of the characters in the above descrip- tion were taken from the figures given by Bierig. This species belongs in the subgenus Cryptobiella as stated by Bierig, although his genotype designation for it is not valid (see Blackwelder, 1939a). Recorded as found in accumulated seaweed. 2. OCHTHEPHILUM TRINITATUM, new species Description.—Rufous, elytra piceous with apical margin testaceous. Head narrowed in front of the eyes; eyes situated at middle of length; coarsely and densely, almost umbilicately punctate, sparser in front; with rather indistinct ground sculpture throughout; basal segment of antennae three-fourths as long as head, over twice as long as distance between antennae; antennal grooves distinct; labrum with distinct denticle at sides of a rounded median emargination; neck half as wide as head. Pronotum with narrow smooth and slightly elevated median band, not canaliculate at base; punctures rather large, umbilicate, leaving smooth areas, without ground sculp- ture. Hlytra with coarse punctures, narrowly separated by convex intervals, without trace of serial arrangement; intervals with very indistinct ground sculpture. Afale, seventh sternite with a very shal- low but abrupt median emargination and the surface depressed pos- teriorly before the emargination; eighth with an abrupt and deep narrow emargination twice as deep as wide and gradually rounded to apex. Female, eighth sternite truncate. Length, 5 mm. Type locality —Trinidad, 8 miles northeast of St. Augustine on the Tacarigua River, county of St. Georges, ward of Tacarigua. Types.—Holotype, female, and four paratypes, male, U.S.N.M. No. 52540, collected by me on December 22, 1935; four paratypes in the Museum of Comparative Zoology; one paratype in the British Museum. Fecords.—The following are the records known to me: Trinidad: Tacarigua (Blackwelder station 107C), St. Augustine (Withycombe, in the British Museum; Darlington, in M.C.Z. and U.S.N.M.), Cedros (Darlington, in M.C.Z.). Specimens examined.—\ have seen only the ten types. Remarks.—This species differs from atlanticwm in the characters used in the key, as well as the presence of denticles and an emargina- tion on the labrum and the canaliculate median pronotal band. It belongs in the subgenus Cryptobiella. The type was found under moss on a rock. 334 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 3. OCHTHEPHILUM DARLINGTONI, new species Description.—Testaceocastaneous, a little dark in front (probably somewhat immature). Head somewhat expanded behind the eyes, nearly circular; eyes very small, separated from base by nearly four times their diameter, from apex by one and one-half times; basal segment of antenna barely half as long as head, about as long as distance between antennae; antennal grooves moderate; labrum with a narrow emargination between two blunt teeth; neck half as broad as head; punctures moderate, not distinctly umbilicate though sloping anteriorly and sculptured within, often separated by less than their diameter, very sparse in front; with very fine scaly ground sculp- ture throughout. Pronotum with an impunctate median band not outlined by regular series of punctures; punctures less coarse than on head but more irregularly spaced; feebly depressed medially be- fore base; with ground sculpture as on head. /lytra shorter than pronotum, narrower than abdomen; with punctures irregularly unit- ing, giving a subrugose appearance, the intervals not flattened ; with- ‘ out ground sculpture but with a somewhat coriaceous texture. Male, unknown. /emale, sternites not modified. Length, 7 mm. Type locality —Cuba, south side of Pico Turquino, elevation 3,000 to 5,000 feet. Types.—Holotype in the Museum of Comparative Zoology, col- lected in June 1936 by Dr. P. J. Darlington. Records—The following is the only record known to me: Cuba: Pico Turquino (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This specimen differs so greatly from the description of atlanticum Bierig that I cannot believe that it is the same. The eyes are small and placed well before the middle, the basal segment of the antenna is very much shorter, the punctures are not umbilicate, and the length is greater. The small elytra with coriaceous texture give it the appearance of a subterranean species. I have received no record of its habits. 4. OCHTHEPHILUM TRINIDADENSE (Bernhauer) Cryptobium trinidadense BERNHAUER, 1934, p. 161. Original description—‘Dem C. rostratum, Sharp, in der Gestalt ausserordentlich ahnlich, durch die Fiarbung und viel feinere und dichtere Punktierung das ganzen Korpers auf den ersten Blick zu unterscheiden. “Briunlichrot, der Halsschild rétlichbraun, der Kopf und die Fliigeldecken pechschwarz, der schmale Hinterrand der Fliigeldecken, der vorderste Teil des Kopfes, die Taster und Beine rétlichgelb, die MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 339 Fiihler braunlich, die Wurzel und Spitze geiblich. Der Kopf ist etwas schmiler als der Halsschild, viel langer als breit, hinter den in der Mitte der Seiten gelegenen Augen unmerklich erweitert, fein und sehr dicht, zwischen und vor den Augen stiirker und viel weit- laufiger punktiert, daselbst g@linzend, hinten matt, vor dem Vorder- rande dusserst zart chagriniert. Die Fiihler sind gestreckt, die vor- letzten Glieder viel linger als breit. Der Halsschild ist viel schmiler als die Fliigeldecken, um ein Drittel linger als breit, vor der Mitte schwach erweitert, ausserhalb der breiten, erhabenen spiegelgliinzen- den Mittelzone ziemlich kraftig und dicht, etwas ungleichmiissig punk- tiert glainzend. Die Fliigeldecken sind betrichtlich linger als der Halsschild, miissig stark und sehr dicht punktiert, wenig glinzend. neben dem Seitenrande ohne Kiellinie. Der Hinterleib ist aiisserst fein und dicht, fast chagrinartig punktiert. Linge: 5.5mm. Beim g ist das sechste Sternit in der Mitte des Hinterrandes breit dreieckig ausgerandet.” Type locality—Trinidad, St. Augustine. Types.—In either the collection of Dr. Bernhauer or in the British Museum; collected on April 15, 1926, by C. L. Withycombe. Records.—The following is the only record known to me: Trinidad: St. Augustine (Bernhauer, 1934). Specimens examined.—I have seen no example of this species. Remarks—The original description seems to be sufficient to place this species in the genus Ochthephilum (Cryptobium of authors) but will not enable me to place it in the key. It seems to differ from the other species in the feeble male characters. I find no record of its habits. XLVII. Genus BIOCRYPTA Casey Biocrypta CAsry, 1905, pp. 26, 51. Genotype—Cryptobium prospiciens LeConte=Biocrypta prospi- ciens (LeConte) (monobasic and original designation). Diagnosis—Integuments moderately punctate, sometimes with ground sculpture; antennae anteriorly flexile, strongly geniculate, basal segment elongate: labrum not denticulate; fourth segment of maxillary palpus subulate; gular sutures separate; neck more than one-fourth as wide as head; prosternum not transversely impressed before coxae, not dilated beneath the coxae; hypomera feebly lobed behind coxae; anterior coxal cavities open behind; front coxae large, exserted; elytra without trace of pleural fold; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second ab- dominal sternites absent; third carinate at base; fifth sternite of male with a median fovea and a posterior lobe; basal half of anterior tibia 336 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with a concavity lined with diagonal ctenidia; apex of posterior tibia with a ctenidium on each side. Remarks.—This genus is very similar to Homocotarsus but it is distinguished by the absence of the pleural fold. From Ochthe- philum it differs by lacking denticles on the labrum. From the West Indies I have seen 98 specimens, representing nine species, of which eight appear to be new. KEY TO WEST INDIAN SPECIES OF BIOCRYPTA 1. Eyes moderately small, separated from base by not over three times their diameters, clytta; normal==225.0 =. tf RE een ee ee, 2, Eyes very smal], separated from base by at least four times their diameter; LYLE QOST CCT CO CS ae aaa eee Se ee ee 5 . Vertex with’ dense’ ground “sculpture. 2.122 ius eee 1. darlingtoni Vertex between the eyes at most with traces of ground sculpture____-___ 3 38. Head longer than broad; sternites of female unmodified; fifth sternite of male with a very large circular fovea______.____.._____-____ 4, dominicana Head as broad as or broader than long; fifth sternite of female generally modified ; fifth sternite of male with at most a moderate fovea_-___--_ 4 4. Head impunctate along center of vertex; punctures seldom separated by less than theircdiameters 232.2 22) fk ee Ae oe ee 3. trinitatis Head more or less punctate throughout; punctures usually separated by less Lhanet hein Ciamelers 22 = = es en ee 2. fulvipes 5. Eyes minute, separated from base by about eight times their diameter. 9. hispaniola Eyes very small but separated from base by only about four times their to Gigmete rn. 21-38 vos ees tee is ee ad 2 ee Be eee = a6 6. Disk of pronotum finely coriaceous but without distinct ground sculpture. 7 Disk of pronotum with distinct ground sculpture, not at all coriaceous____ 8 %.. Head distinctly swollen behind eyes22 28! 2 ne eee 5. haitia Head scarcely at all swollen behind eyes____---__.---____-_ 8. jamaicensis 8. Head between the eyes with dense ground sculpture____-____-__--_ 7. densa Head between eyes without ground sculpture________________-- 6. cubensis 1. BIOCRYPTA DARLINGTONI, new species Description.—Rufopiceous, the pronotum black, posterior margin of elytra rufotestaceous. Head with eyes at twice their length from base, at their length from apex; basal segment of antennae nearly two-thirds as long as head, as long as distance between them; an- tennal grooves distinct; with moderate punctures and a few very coarse ones between the eyes; with very fine and dense ground sculp- ture throughout. Pronoiwm with distinct and slightly elevated smooth median band set off by fairly regular series of punctures; rather coarsely and fairly densely punctured; without ground sculp- ture. Hlytra much larger than pronotum; with dense moderate punctures separated by convex intervals, not distinctly serial; without ground sculpture. Male, fourth sternite with a slightly elevated transverse fovea occupying middle seventh of the width; fifth sternite MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 330 with an obsolescent fovea of about the same size; eighth sternite rounded. Female, unknown. Length, 74% mm. Type locality —Trinidad, St. Augustine. Types.—Holotype, male, in Museum of Comparative Zoology; one paratype, male, in United States National Museum (No. 52503) ; col- lected in Apri! 1929 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species is readily distinguished by the dense ground sculpture of the vertex as well as by the feebly developed male characters. It is a typical member of the genus. I have received no record of its habits. 2. BIOCRYPTA FULVIPES (Erichson) Cryptobium fulvipes Exicuson, 1840, p. 566—Horn, 1885, p. $98 —BrrNHAUER and Scuvuserr, 1912, p. 283.—Lene and MurcHtrrr, 1914, p. 405.—Wotocort, 1924, p. 78; 1936, p. 197.—BLAcKWELDER, 1939a, p. 114. Biocrypta fulvipes (Erichson) BLACKWELpDER, 1939a, p. p. 114. Description—Rufous or rufocastaneous with head picescent. Head with punctures of very irregular sizes, rather scattered; with feeble ground sculpture except between the eyes; eyes situated at middle of length of head; basal segment of antennae scarcely over one-half as long as head, about as long as distance between them; antennal grooves distinct. Pronotum with irregular median band not elevated or canaliculate; punctures moderate, rather sparse but irregularly spaced, not at all serial; without ground sculpture. /ytra much larger than pronotum; with dense moderate punctures separated by flattened inter- vals, not distinctly serial; intervals with indistinct ground sculpture. Male, fourth sternite unmodified, fifth with a transverse fovea at middle and a posterior lobe about one-third as broad at base as seoment and extending to middle of sixth sternite or middle of seventh, with apex truncate; eighth sternite with a small abrupt rounded emargination. Female, all sternites unmodified except for fifth, which has a central smooth area rising to an abrupt semicircular posterior rim. Length, 714 mm. Type locality—Puerto Rico. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records—The following are the records known to me: Puerto Rico: (Erichson, 1840; Horn, 1885; Bernhauer and Schubert, 1912; Leng and Mutchler, 1914; Wolcott, 1924, 1986), Villalba (Blackwelder station 64), Maricao Forest (Darlington, in M.C.Z.), El Yunque (Darlington, in M.C.Z, and U.S.N.M). Antigua: (H. E. Box, as Blackwelder stations 4441 and 444M). 308 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Guadeloupe: (Bernhauer and Schubert, 1912; Leng and Mutchler, 1914). Grenada: (Bernhauer and Schubert, 1912; Leng and Mutchler, 1914; British Museum). South America: COLOMBIA, VENEZUELA, Ecuapor (Bernhauer and Schubert, 1912). Specimens examined.—l have seen 18 examples in the British Museum, two collected by Dr. Harold E. Box, three from the Museum ot Comparative Zoology, and two collected by me on October 12, 1935. Remarks.—This species differs from the genotype in the following particulars (among others): Much more distinct punctures on head and elytra and much less distinct ground sculpture, shorter basal segment of antennae, coarser punctation of pronotum, and the much larger fovea of the fifth sternite of the male and the smaller emar- gination of the eighth. There is at least some doubt of the South American records. I have seen specimens only from Puerto Rico, Antigua, and Grenada and have no direct evidence that it is as widespread a species as is indicated in the Junk Catalog. My specimens were taken while sifting leaves in a damp gully. 3. BLIOCRYPTA TRINITATIS, new species Description.—Rufopiceous. Head with eyes separated from base by about twice their length; basal segment of antennae about two- thirds as long as head, as long as distance between them; antennal grooves distintt; with punctures of various sizes rather sparse; with ground sculpture only at base and apex. Pronotum with median smooth area not elevated but set off by fairly regular series of mod- erate punctures; sparsely and only moderately coarsely punctate; without ground sculpture. #/ytra larger than pronotum; with dense moderately coarse punctures separated by flattened intervals, not at all serial; without distinct ground sculpture. Male, unknown. Female, all sternites unmodified except for fifth which has a central smooth area rising to an abrupt semicircular posterior rim. Length, 8 mm. Type locality —Trinidad, St. Augustine. Types.—Holotype, female, in Museum of Comparative Zoology, col- lected in April 1929 by Dr. P. J. Darlington. Pecords.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z.). Specimens ewamined.—I have seen only the unique type. Remarks.—This specimen had been confused with fulvipes but seems to be very distinct by the sparse punctation. Even an isolated female seems worthy of description in this genus, of which only one species is so far known from South America (Argentina). T have received no record of the habits of this species. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 339 4. BIOCRYPTA DOMINICANA, new species Description —Black, sometimes picescent in part. Head with eyes about at middle of length; basal segment of antennae about half as long as head, about as long as distance between them; antennal grooves distinct; with moderate but uneven punctures fairly dense basally but sparse in front; with rather indistinct ground sculpture basally. Pronotum with slightly elevated smooth median band set off by punc- tures only partly in series; with moderate punctures frequently sepa- rated by about their diameter; without distinct ground sculpture. Elytra much larger than pronotum; with dense but rather small punc- tures separated by somewhat flattened intervals, not at all serial; without distinct ground sculpture. fale, fourth sternite with a large circular fovea; fifth with a still larger circular fovea occupying nearly the entire middle third of the segment; seventh with a narrow oval excision one-third as long as segment, with angles somewhat pro- longed, the excision bordered by a thin bead which extends backward us two short spiniform processes between the extended angles; eighth very deeply narrowly emarginate. Female, sternites not modified. Length 614 to 7144 mm. Type locality—Dominican Republic, Sanchez. Types.—Holotype, male, and two paratypes, female, in the Museum of Comparative Zoology; two paratypes, male and female, in the United States National Museum (No. 52504); collected in July 1938 by Dr. P. J. Darlington. fecords.—The following are the records known to me: Hispaniola: DoMINICAN Repusric, Sanchez (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—\ have seen only the five types. Remarks.—Vhis species is very distinct by the exceptional develop- ment of the male characters. The details of the emargination of the eighth sternite are unique among the Cryptobia, and the foveae of the fourth and fifth sternites reach their greatest development in this species. It is, however, a typical member of the genus as herein defined. I have received no record of its habits. 5. BIOCRYPTA HAITIA, new species Description.—Rufopiceous, head and pronotum somewhat darker. Head with eyes small, separated from base by about four times their diameter, by one and one-half times from apex; basal segment of antennae a little over half as long as head, about as long as distance between them; antennal grooves feeble; with moderately coarse punc- tures, distinctly umbilicate, dense at base, sparse in front where they are obscured bv the dense but irregular ground sculpture. Pronotum with irregular impunctate median band not marked by series of punc- 340 BULLETIN 182, UNITED STATES NATIONAL MUSEUM tures; punctures about as on head but less dense and more irregularly placed; without distinct ground sculpture, but with surface through- out minutely uneven. Hlytra shorter and scarcely wider than pro- notum, narrower than the abdomen, closely appressed ; with punctures about as coarse as on head but denser and less regularly shaped, not at all serial; ground sculpture as on pronotum. fale, with a small circular pale area (scarcely a fovea) at center of posterior margin of fifth sternite; seventh with punctation and vestiture finer and denser along middle posteriorly; eighth with a narrow attenuated emargina- tion, a little deeper than wide at apex, at least three times as deep as wide at middle. Female, sternites unmodified. Length, 714 mm. Type locality —Haiti, Morne La Hotte, 5,000 to 7,800 feet. Types.—Holotype, male, in Museum of Comparative Zoology ; para- type, female, in United States National Museum (No. 52506) ; collected on October 16-17, 1934 by Dr. P. J. Darlington. fecords.—The following is the only record known to me: Hispaniola: Harri, Morne La Hotte (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks—This species has the aspect of a subterranean insect. It may be distinguished by the peculiar male characters which are unique in the Cryptobia of the West Indies. It was taken at a high altitude (for the West Indies), probably under ground cover. 6. BICCRYPTA CUBENSIS, new species Description.—Rufopiceous, the head and pronotum sometimes less rufescent. Head with eyes small, separated from base by over four times their diameter, by two times from apex; basal segment of an- tenna three-fifths as long as head, a little longer than distance between them; antennal grooves feeble; with moderately coarse but sparse and sometimes umbilicate punctures interspersed with minute punctures; with dense ground sculpture throughout (obsolescent in small area of vertex between the eyes). Proenotwm with irregular impunctaie median band marked by punctures that are only partly serial; punc- tures a little less coarse than on head, sparse and seldom serial; with fine but distinct ground sculpture throughout. LZ/ytra shorter and scarcely wider than pronotum, narrower than the abdomen, closely appressed ; with punctures about as coarse as on head but denser and less regularly shaped, not at all serial; without distinct ground sculp- ture, but surface strongly coriaceous. ale, fifth sternite with a small median oval fovea, posterior fifth of margin prolonged in a narrow abrupt but tapering lobe which extends to middle of seventh; eighth not distinctly emarginate. Fema/e, fifth sternite with a central smooth area rising to an abrupt semicircular posterior rim. Length 814 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 341 Type locality —Cuba, south side of Pico Turquino, in province of Oriente, elevation 3,000 to 5,000 feet. Types.—Holotype, male, and paratype, female, in Museum of Com- parative Zoology; two paratypes, female, in United States National Museum (No. 52509) ; collected in June 1936 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Cuba: Pico Turquino (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the four types. Remarks.—This is one of the subterranean type, which may be distinguished readily by the sculpture as well as by the male charac- ters. In the latter it is most like jamaicensis, which has the sternal lobe similar but the eighth sternite emarginate. (The only male of cubensis I have seen has the apex of the eighth sternite somewhat chipped away. Enough remains to lead me to believe that it was not emarginate as in jamaicensis.) These specimens were taken at considerable elevations, probably under ground cover. 1. BIOCRYPTA DENSA, new species Description —Rufocastaneous, the head and pronotum somewhat picescent. Head with eyes small, at about four times their length from base, and about two times from apex; basal segment of antennae about three-fifths as long as head, scarcely longer than distance between them; antennal grooves feeble; with uneven but not very coarse punc- tures, each declivous or excavated in front, not distinctly umbilicate, sparse in front, posteriorly separated by one to two times their diam- eter; with fine but dense ground sculpture throughout. Pronotwm with irregular impunctate median band not marked by definite series, vaguely foveate near base; punctures much less coarse than on head but umbilicate and seldom at all serial; with fine but dense ground sculpture throughout. Z’lytra shorter and scarcely wider than pro- notum, narrower than the abdomen, closely appressed ; punctures about as coarse as on head but often indistinct and sometime coalescent, sur- face both coarsely and finely coriaceous but without other ground sculpture. fale, fifth sternite with a small oval median anterior fovea, the middle third of posterior margin abruptly prolonged in a rectangular lobe, which extends over base of seventh; eighth with a small rounded notch preceded by an indefinite triangular smooth area (gutter). Female, unknown. Length, 714 mm. Type locality—Haiti, La Visité and vicinity in the La Selle Range, 5,000 to 7,000 feet elevation. Types.—Holotype, male, in Museum of Comparative Zoology, col- lected on September 16-23, 1934, by Dr. P. J. Darlington. 342 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following is the only record known to me: Hispaniola: Harti, La Visité (Darlington, in M.C.Z.). Specimens examined.—\ have seen only the unique type. Remarks.—This species is very distinct by its even fine ground sculpture. It is similar to jamaicensis in the male characters except that the lcbe is broader and not tapering to the truncate apex. I have received no record of its habits except that it was taken at a considerable altitude, probably under ground cover. 8. BIOCRYPTA JAMAICENSIS, new species Description.—Rufocastaneous, forepart sometimes picescent. Head with eyes small, separated from base by nearly four times their diame- ter, by two times from apex; basal segment of antenna nearly three- fifths as long as head, distinctly longer than distance between them; the antennal grooves feeble; very irregularly punctured, some punc- tures fairly coarse, dense posteriorly but only a few isolated in front ; without distinct ground sculpture but vaguely coriaceous, more strongly in front. Pronotwn with irregular impunctate median band marked by very irregular series of punctures; punctures much as on head; without distinct ground sculpture but surface irregularly coriaceous. 7ytra shorter and scarcely wider than pronotum, nar- rower than the abdomen, closely appressed; with punctures a little coarser than on head but sometimes united irregularly and separated by irregular convex intervals; without distinct ground sculpture. Male, without trace of fovea on fourth sternite; with a moderate transversely oval fovea on fifth, and with a narrow triangular lobe extending to apex of sixth sternite; sixth and seventh without vesti- ture along middle; eighth with a small rounded triangular emargina- tion, about as deep as wide. Female, fifth sternite with a subbasal median smooth area rising to an abrupt semicircular posterior rim. Length, 8 to 9 mm. Type locality—Jamaica, main range of Blue Mountains at an elevation of 5,000 to 7,000 feet. Types.—Holotype, male, and paratype, female, in the Museum of Comparative Zoology; two paratypes, male, in the United States Na- tional Museum (No. 52507) ; all collected between August 17 and 19, 1934, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the four types. Remarks.—This is likewise a subterranean form in appearance. It is readily distinguished by the characters in the key as well as by the male characters. This is the only species so far taken on Jamaica MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 343 but is similar in many respects to the mountain species of Cuba and Hispaniola. I have received no record of its habits, 9. BIOCRYPTA HISPANIOLA, new species Description.—Rufocastaneous, somewhat picescent in front. Head somewhat enlarged behind the eyes; eyes minute, separated from base by eight times their diameter, by pearly three times from the apex; basal segment of antenna unusuaily slender and curved, about half as Jong as head, and as long as distance between them; antennal grooves very feeble; with moderately fine anteriorly directed punctures sepa- rated by one to two times their diameter; with dense but very fine and rather indistinct ground sculpture. Pronotum with vague impunc- tate median band not outlined by series of punctures; punctures a little finer than on head, usually separated by about twice their diame- ter; surface smooth, without ground sculpture. Elytra slightly shorter and scarcely wider than pronotum, narrower than abdomen, closely appressed ; conjointly deeply emarginate behind; punctures obsoles- cent, sometimes vaguely submuricate ; with dense but indistinct ground sculpture. fale, with a circular fovea in middle sixth along apical margin of fourth and fifth sternite, sixth and seventh transversely impressed and smooth before base, eighth elongate but apex deeply incised, the slit preceded by a narrow groove. Female, sternites not modified. Length, 7 to 10 mm. Type locality—Hispaniola, Loma Rucilla and vicinity in the Dominican Republic; elevation 5,000 to 8,000 feet. Types.—Holotype and 138 paratypes, male, and 21 paratypes, female, in the Museum of Comparative Zoology; 27 paratypes, male and female, in the United States National Museum (No. 52505) ; collected in June 1938 by Dr. P. J. Darlington. Records.—The following are the records known to me: Hispaniola: DoMINICAN RepuBLic, Loma Rucilla (Darlington, in M.C.Z. and U.S.N.M.), Valle Nuevo (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 62 types. Remarks.—This is a very distinct species because of the size of the eye, the shape of the head and basal antennal segment, the sculpture, and the male characters. Very little variation is exhibited by the fine series examined. The specimens were taken in cloud forest presumably under ground cover. XLVIII. Genus OPHITES Erichson Ophites EricHson, 1840, p. 627. Genotype.—O. versatilis Erichson (designated by Duponchel, 1841, not Blackwelder, 1939a). 344 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Diagnosis —At least part of integuments punctate, sometimes with ground sculpture; antennae anteriorly flexile, strongly geniculate, basal segment much elongate; labrum bidenticulate; maxillary pal- pus elongate, fourth segment globose and acuminate; gular sutures united; neck less than one-fourth as wide as head, nearly half as long as head; prosternum longitudinally impressed before each coxa, not dilated beneath them; hypomera broadly lobed over base of coxae; anterior coxal cavities open behind; front coxae large, ex- serted; elytra without distinct pleural fold; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; subbasal sternites of male unmodified ; basal half of anterior tibia without concavity but with area of diag- onal ctenidia; apex of posterior tibia without distinct ctenidium on either side. Remarks.—This is a very distinct genus because of the remarkable shape of the head. It is unusual also in the arrangement of the sculpture, which is almost restricted to the head, while only the elytra are densely punctate. The genus is neotropical. Only one species has been found in our region and that in Trinidad. 1. OPHITES BUGNIONI Fauvel Ophites bugnioni Fauvet, 1901, p. 80.—BrERNIZAUFR and ScHUBERT, 1912, p. 277. Description.—Black, apex of elytra and abdomen and appendages rufotestaceous. Head abruptly and evenly narrowed from eyes to neck; with a few scattered punctures obscured by the dense sculp- ture, which is composed of minute setiferous punctures. Pronotum nearly twice as long as broad, strongly narrowed in front, sides a little emarginate behind middle; with a few moderate umbilicate punctures about basal angles, two discal series and others obsoles- cent; smooth, shining. Z/ytra rectangular, one-half wider than pronotum, the humeral angles tuberculate; with coarse submuricate punctures more or less serial especially near the suture; without ground sculpture. Abdomen very densely and minutely submuri- cately punctate (appearing like sculpture). Afale, eighth sternite with a U-shaped excision nearly half as deep as visible length of seoment, two-thirds as wide as deep, the apices rounded. Female, eighth sternite broadly and very feebly emarginate. Length, 10 mm. Type locality—*Colombie: Palomino.” Types——Presumably in the collection of Blanche Rancin, Caén, France. fRecords.—The following are the records known to me: Trinidad: St. Augustine (Darlington, in M.C.Z. and U.S.N.M.), Four Roads (Thaxter, in M.C.Z.). South America: CoLompria (Fauvel, 1901). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 345 Specimens examined.—l have seen 24 examples from the Museum of Comparative Zoology, of which some will be deposited in the United States National Museum. Remarks.—This is one of the most easily recognized staphylinids in our region. It is, of course, a member of the South American fauna and is believed to be Fauvel’s species, although I have not been able to compare it with other specimens of that species. It differs widely in punctation and sculpture from O. versatilis Erichson. Dr. Darlington collected this series from deep grass along a pond. XLIX. SUNIOPHACIS, new genus Genotype.—S. concolor Blackwelder (designated here). Diagnosis—Body rather short and stout; antennae moderately filiform, not geniculate; labrum without teeth, truncate, narrowly emarginate at center; maxillary palpi long, fourth segment minute, pubescent; gular sutures united; prothorax nearly circular, not much prolonged anteriorly at middle; prosternum expanded under coxae nearly to hypomera but not connate with the latter; anterior coxal cavities closed behind (by the sternum beneath the coxae rather than by the hypomera); front coxae conical, prominent; middle coxal cavities confluent; posterior coxae conical, approximate; abdomen margined, somewhat contracted at base; first and second sternites absent; in general, eighth sternite feebly emarginate in males, feebly prolonged in females; fourth segment of tarsus lobed beneath. Remarks.—This genus appears to be distinguishable from Stilo- saurus, Suniocharis, Stamnoderus, Suniosaurus, and Stiliphacis by the characters mentioned in the key. The genera in this complex are not yet sufficiently understood to make the generic limits certain, but so far as the West Indian species are concerned the genera are readily recognized. Of this genus I have seen 1 example in the Museum of Compara- tive Zoology, 1 in Dr. Cameron’s collection, and 12 in the United States National Museum. These I divide into three rather feebly differenti- ated species, of which two are described as new. KEY TO WEST INDIAN SPECIES OF SUNIOPHACIS 1, Pronotum with umbilicate punctures united in distinct longitudinal grooves. 3. hubbardi Pronotum with umbilicate punctures irregular, not in even rows________ 2 DP eLy tra “entirely testaCeOU ses saw aa ee REE a SC hk 1. concolor Hlytrainipart piceons oxiinfuseates 2 -— ea et See 2. thoracicus 1. SUNIOPHACIS CONCOLOR, new specics Description.—Rufopiceous, elytra testaceous. Head moderately large, suborbicular, almost as wide as elytra; eyes moderate, not very 449008—42 25 346 BULLETIN 182, UNITED STATES NATIONAL MUSEUM prominent; sides behind eyes a little elongate but evenly rounded to base; punctures dense and umbilicate but feeble and shallow, with scaly sculpture on intervals and more generally in front; labrum rounded at sides and in front on either side of a shallow emargination. Pronotum one-eleventh longer than wide, over five-eighths as wide as elytra, widest at anterior third, strongly narrowed in front to narrow apex, moderately behind to rounded angles; sides finely tuberculate ; more strongly umbilicately punctate than head but with punctures generally united in irregular longitudinal rows; without ground sculp- ture except vaguely on the intervals. Prosternum not at all longi- tudinally carinate on disk. “7Vytra almost as wide as long, about two- fifths longer than pronotum; with punctures rather fine but depressed so as to appear larger, surface uneven. Abdomen rather finely but not densely punctate; with rather long pubescence. Lighth sternite broadly and feebly lobed at apex. Length, 2 mm. Type locality —Jamaica. Types.—Holotype and one paratype, U.S.N.M. No. 52389, collected by H. G. Hubbard. Records.—The following is the only record known to me: Jamaica: (Hubbard, in U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—These specimens seem to be distinct from thoracicus though rather similar to that species. Again I can find no evidence of sex in the two specimens before me. T have no record of the habits of this species. 2. SUNIOPHACIS THORACICUS (Cameron) Stilicopsis thoracicus CAMERON, 1923, p. 3898.—SCHEERPELTZ, 1933, p. 1237.—BLacK- WELDER, 1939, p. 115. Monista antillana Birrte, MS. Description.—Black ; elytra rufotestaceous, each with a black median band from lateral margin nearly to suture; apex of abdomen obscurely testaceous. Head large, subquadrate, rounded behind eyes which are small; outer antennal segments transverse; punctures moderately close, superficial and umbilicate ; intervals distinctly coriaceous. Pro- notwm searcely longer than broad, widest before middle, anterior angles rounded, sides obliquely contracted in front to narrow neck, posteriorly less strongly contracted to rounded posterior angles; shin- ing at middle, with trace of midline; sculpture distinctly coriaceous; with a few obsolete umbilicate punctures sparingly distributed and indistinct. Prosternum with only a trace of a longitudinal carina on disk. Flytra a little longer and distinctly wider than pronotum, almost as wide as head, longer than broad; shining, moderately finely and moderately closely punctate; finely pubescent. Abdomen nar- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 347 rowed at base and apex, extremely finely and moderately closely punc- tate throughout and very finely pubescent. Male, unknown. Length, 2mm. (From Cameron.) Type locality —Haiti. L'ypes—Unique type in collection of Dr. Cameron, (A “cotype” of antillana is in the United States National Museum.) Reeords—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Habana (Bierig, in U.S.N.M. as antil- lana). Hispaniola: Haitr (Cameron, 1923). Specimens examined —I have seen the unique type, which I ex- amined briefly in Dr. Cameron’s collection, and nine examples in the United States National Museum. Remarks.—The examples from Cuba have not been compared with the type but differ from the original description only in the paler color of each part. This might well be due to differences in lighting or to differences in personal judgment. In my specimens the labrum is rounded at the sides and feebly so in front on each side, with a very small acuminate notch at center. I do not find any evidence of sex in my examples. I have no record of the habits of this species. 3. SUNIOPHACIS HUBBARDI, new species Description—Piceous, apical third of elytra testaceous. Head suborbicular, eight-ninths as wide as elytra; eyes moderately large, not very prominent, sides behind the eyes somewhat elongate, evenly rounded; labrum rounded at sides and in front on either side of a small shallow notch. Pronofwm scarcely as wide as long, over three-fourths as wide as head, widest at anterior third; strongly nar- rowed in front to smal! neck, moderately narrowed posteriorly to rounded basal angles; sides finely tuberculate; without midline; with distinct umbilicate punctures almost always united in long grooves; intervals with distinct ground sculpture. Prosternum not at all longi- tudinally carinate on disk. Hlytra scarcely longer than wide, two- fifths wider than pronotum; punctures moderately fine, feebly sub- muricate. Abdomen very finely muricately punctate, without ground sculpture but with rather long pubescence. Highth sternite unmodi- fied. Length, 234 mm. Type locality— Jamaica. Types.—Holotype, U.S.N.M. No. 52392, collected by H. G. Hubbard. Records.—The following are the records known to me: Jamaica: (Hubbard, in U.S.N.M.), Kingston (Darlington, in M.C.Z.). Antigua: (Weber, in M.C.Z.). Specimens examined.—t have seen only the holotype and two ex- amples in the Museum of Comparative Zoology. 348 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—The description of the elytral coloring given above should be modified as follows: A median black band from margin nearly to suture is black, the apical third is testaceous, the remaining part at base and along suture is so darkly piceous as to be scarcely distinguish- able from the black band. It appears to me to be a coloration very similar to that described by Cameron for thoracicus, but there seems to be ample distinction in the sculpture of the pronotum. The two specimens in the Museum of Comparative Zoology differ from the type in having a much lighter color with the elytral spot much clearer. They do not seem to differ otherwise. I have no record of the habits of this species. L. STILOSAURUS, new genus Genotype—S. rulomus Blackwelder (designated here). Diagnosis —Body moderately slender, strongly constricted; an- tennae filiform, not geniculate; labrum enlarged at middle, bidentate with additional angulations; maxillary palpus long, third segment enlarged, fourth minute; gular sutures united; prothorax slightly oval, not much prolonged at middle in front; prosternum expanded under coxae as far as but not connate with the hypomera; anterior coxal cavities closed behind (by the sternum beneath the coxae rather than by the hypomera) ; front coxae conical, prominent; middle coxal cav- ities confluent; posterior coxae conical, approximate; abdomen mar- gined; somewhat contracted toward base; first and second sternites absent; in general, eighth sternite feebly emarginate in males, feebly prolonged or rounded in females; fourth segment of tarsus lobed beneath. Remarks.—This genus is readily recognizable by the absence both of umbilicate punctures and of dense coarse sculpture. It usually has fine ground sculpture and sometimes shows traces of umbilicate punc- tures, as if they were obsolescent. I have seen 11 examples belonging to three species, all of which are herein described as new. KEY TO WEST INDIAN SPECIES OF STILOSAURUS i. Head with epistomal suture and anterior part of coronal suture distinct. 8. coronalis Head without raised line marking epistomal and anterior part of coronal SUUUTCS 22 2s ee ee ne ae ee 2 2. Anterior part of pronotal midline marked by a raised line in a depression. 2. lineatus Anterior part of pronotal midline depressed but without ralsed line, 1. rulomus 1. STILOSAURUS RULOMUS, new species Description.—Rufopiceous to castaneous. Head moderately large, semicircular behind, quadrate in front, about as wide as elytra; eyes MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 349 moderately large, prominent; labrum with two acutely triangular median teeth adjacent at base but without other denticles; impunc- tate but with dense fine ground sculpture and sparse rather indistinct round tubercles; without elevated lines marking sutures on vertex. Pronotum slightly longer than wide, as wide as head, widest just before middle, sides evenly rounded posteriorly, base not distinct; with three large setae along each side; midline feebly depressed and without raised line; with sculpture and tuberculi more distinct than on head. lytra one-fourth wider than long, four-fifths as long as pronotum; with very coarse punctures somewhat obscured by the uneven surface. Abdomen with punctures indistinct; without dis- tinct ground sculpture. fale, seventh sternite feebly emarginate throughout; eighth with a broad emargination at least four times as wide as deep. Female, sternites not modified. Length, 214 mm. Type locality—Dominican Republic, cloudforest in the vicinity of Valle Nuevo; elevation about 6,000 feet Types.—Holotype, male, and two paratypes in the Museum of Com- parative Zoology; two paratypes in the United States National Museum (No. 52520); collected in August 1988 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Hispaniola: DoMINICAN ReEPuBLic, Valle Nuevo (Darlington, in M.C.Z. and U.S.N.M.) Specimens examined.—I have seen only the five types. Remarks.—This species may be distinguished by its lack of raised lines on the head and pronotum. It is also unique in the distinct- ness of the tuberculi. I have received no record of its habits except that it was taken in cloudforest. 2. STILOSAURUS LINEATUS, new species Description —Castaneous or testaceocastaneous. Head moderately large, semicircular behind, quadrate in front, about as wide as elytra; eyes moderately large, prominent; labrum with two acutely triangu- lar median teeth adjacent at base and flanked by small angulations but no other denticles; without punctures or distinct ground sculp- ture but with indistinct tuberculi basally and laterally only, surface rather uneven; without elevated lines marking sutures on vertex. Pronotum one- cool longer than wide, not quite as wide as head, widest near middle, sides rather feebly arcuate and rounded into base; with two to four large setae along each side; midline feebly impressed in apical half and marked by a fine smooth raised line; with dense ground sculpture, and with tuberculi a little more distinct than on head. lytra two-fifths wider than long, five-sevenths as long as pronotum; with coarse punctures almost completely obscured 300 BULLETIN 182, UNITED STATES NATIONAL MUSEUM by unevenness of the surface. Abdomen without distinct punctures but with irregular pitted sculpture. Male, seventh sternite very broadly evenly and feebly emarginate; eighth same as seventh. Female, sternites unmodified. Length, 214 mm. Type locality—Puerto Rico, El Yunque Peak; elevation about 3,000 feet. Types.—Holotype, male, and paratype, female, in the Museum of Comparative Zoology; paratype, female, in the United States National Museum (No. 52521) ; collected in May 1938 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Puerto Rico: El Yunque (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—t have seen only the three types. Remarks.—This species may be known by its reduced elytra and the raised line on the pronotum. I have received no record of its habits. 3. STILOSAURUS CORONALIS, new species Description —Testaceous. Head moderately large, suborbicular, almost as wide as elytra; eyes moderately large, prominent; evenly rounded behind eyes; without distinct punctures or sculpture but with indistinct tuberculi basally and laterally; with a fine raised line marking the position of the epistomal suture between the front of the eyes, aid another marking the anterior fifth of the coronal suture; labrum with median lobes forming two very blunt triangular teeth flanked by a pair of obtuse angulations. Pronotum suborbicular, scarcely wider than long, five-sixths as wide as head, widest near middle, evenly rounded Soneeienty but sides becoming straight near Apex; piled with about three large setae; with a distinct raised midline through most of the length; tuberculi larger and more con- spicuous than on the head but obscured by the dense fine scaly ground sculpture. /lytra three-fourths as long as wide, nine-tenths as long as pronotum; with moderate punctures obscured by the uneven surface. Abdomen exceedingly minutely punctate or sculptured ; with rather dense pubescence. Male, unknown. Female, sternites rounded or broadly lobed. Length, 124 mm. Type locality. =enhaat Sayamas, 15 miles west-northwest of Cien- fuegos, Province of Santa Clara. Types.—Holotype and one paratype, females, U.S.N.M. No, 52390, collected by KE. A. Schwarz. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Specimens examined.—I{ have seen only the two types and one example from the Museum of Comparative Zoology. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 35k Remarks.—TVhis species is unique in the genus by having fine ele- vated lines marking the epistomal and coronal sutures. This con- dition is known to occur among other Staphylinidae only in the sub- family Tachyporinae and one or two other Paederinae. T have no record of the habits of this species. LI. Genus STAMNODERUS Sharp Stamnoderus SHARP, 1886, p. 607. Genotype.—S. godmani Sharp (designated by Blackwelder, 1939). Diagnosis —Body moderately slender, strongly constricted; anten- nae posteriorly flexile, not strongly geniculate, basal segment not very much elongate, segments slender, swollen anteriorly toward apex; eyes prominent; labrum large, produced at middle, without denticies; fourth segment of maxillary palpus very small, subulate; gular sutures united; neck one-fifth as wide as head or less; prothorax narrowly prolonged in front; prosternum carinate along middle throughout, prolonged posteriorly between coxae and laterally be- neath them, connate with edges of hypomera; hypomera not pro- longed behind anterior coxal cavities which are entirely open behind above but entirely closed by prosternal processes; front coxae very large, exserted; middle coxal cavities confluent; posterior coxae con- tiguous, “conical”; first and second abdominal sternites absent, third carinate at base, seventh sometimes emarginate in male, eighth emar- ginate in male; tarsi 5-segmented, fourth segment lobed beneath. Remarks.—This is one of the easiest of all our Paederini genera to recognize because of its prolonged prothorax. It is known only from temperate and tropical America. The record of Stamnoderus sallaci Sharp from Jamaica by Gowdey, 1926, is probably a misidentification. Until the specimens can be reexamined, it must be regarded as a doubtful record. I have seen 151 examples of this genus and assign them to four species, of which one is described as new. KEY TO WEST INDIAN SPECIES OF STAMNODERUS 1, Clypeus smooth or at most with obsolete punctation__-_________-- 2. labeo Clypeus with distinct punctures or ground sculpture__-_______--__-____-~- 2 Ze eunctation or, headeopsolete posteriorly=“2-=_ = = Se eee 3 Punctation of headidistincet throughout: 22. =- 225-2 1. varians oy coronal suture, marked by.a raised line. a 3. bernhaueri Coronalvsutureyentirely- unmarked 22 on ee ee 4. cameroni 1. STAMNODERUS VARIANS Cameron Stamnoderus varians CAMERON, 1918, p. 335.—LENe@ and MutcHter, 1917, p. 198.— SCHEERPELTZ, 1933, p. 1237.—BLACKWELDER, 1939a, p. 115. Description.—Piceous, apex of elytra and abdomen transparent rufous. Head without trace of coronal or epistomal sutures; clypeus 352 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with strong umbilicate punctures as on vertex; labrum transverse with a broad median anterior lobe. Pronotuwm with punctures obsolescent, not distinctly umbilicate; with only a feeble flat midline; no ground sculpture; lateral impressions moderate but distinct. 7ytra slightly shorter than long; sides evenly arcuate, not expanded posteriorly; with moderately coarse and distinct punctures, generally separated by less than their diameter by rather flat intervals. Abdomen very finely and not distinctly submuricately punctate. fale, eighth sternite with a shallow emargination. Female, sternites not modified. Length, 21% to 31% mm. Type locality —Grenada, Balthazar. Types.—Twelve examples (one labeled type) in the British Mu- seum). Records.—The following are the records known to me: St. Vincent: (Cameron, 1913; Leng and Mutchler, 1917). Grenada: (Cameron, 1913; Leng and Mutchler, 1917). Specimens examined.—I have seen 56 examples (including the type) in the British Museum and 6 in the collection of Dr. Cameron. One example was borrowed from the British Museum for further study. Remarks.—Vhis species seems to be distinct from the others of our region by the strength of the umbilicate punctures of the head. It is also distinct in coloration. I find no record of its habits. 2. STAMNODERUS LABEO (Erichson) Sunius labeo ErtcHson, 1840, p. 648.—SHarp, 1886, p. 607.—BrackwEvper, 1939a, p. 115. Stamnoderus labco (Hrichson) SHaAxrp, 1886, p. 607 —BERNHAUER and ScHuBERT, 1912, p. 220.—CAMERON, 1913p, p. 334.—Lene and Murcntrr, 1914, p. 404.— BLACKWELDER, 1939a, p. 115. Stamnoderus delauneyi FLeutTiAux and SALif, 1889, p. 380, pl. 7, fig. 7.—BrRn- 1912, p. 220.—CAMERON, 1913b, p. 8834—-LENG and MurcHter, 1914, p. 404— BLACKWELDER, 1939a, p. 115. Stamnoderus apicalis CAMERON, 1913b, p. 335.—LreNe and Murcuiksr, 1917, p. 198.—SCHEERPELTZ, 1933, p. 1237—BLACKWELDrER, 1939a, p. 115. Stamnoderus incarenatus Breric, 1936, pp. 140, 142, 148, pl. 6, figs. 8-10. Stamnoderus caribacus Breric, 1936, pp. 141, 142, pl. 6, figs. 5-7. Stamnoderus truncatus Brarie, 1936, p. 142, pl. 6, figs. 11-15. Description.—Rufotestaceous, elytra and abdomen testaceous. Head without trace of coronal or epistomal sutures; clypeus without distinct punctures but sparsely pubescent; vertex with obsolete punc- tures vaguely umbilicate; labrum transverse with a large broad median anterior lobe. Pronotum with only a trace of midline; punctures obsolete, vaguely umbilicate; sparsely pubescent; lateral impressions well defined. Zlytra as wide as long, enlarged to apical fourth; with MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 353 punctures rather dense but indefinite, intervals irregularly convex. Abdomen very finely, more or less submuricately punctate. Male, eighth sternite very broadly and feebly emarginate; ninth tergite sub- rectangularly emarginate, declivous, and with four black hairs at each side. Female, eighth sternite arcuate posteriorly; ninth tergite deeply triangularly emarginate, with three or four black hairs at each side. Length, 214 to 314 mm. Type locality—“Americae ins. St. Johannis” (=St. John in the Virgin Islands of the U.S. A.). Of delauneyi, Guadeloupe, Camp- Jacob and Trois Rivieres; of apicalis, Jamaica, Hope River district; of tncarenatus, caribaeus, and truncatus, Cuba. Types.—tiIn either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of delauneyi, possibly in the Musée d’Histoire Naturelle National, Paris; of apicalis, in collection of Dr. Cameron; of incarenatus, caribacus, and truncatus, in collection of Alexander Bierig, and “cotypes” in the United States National Museum (Nos. 52721, 52722, 52720) or the collection of R. E. Blackwelder. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in United States National Museum), Sierra Bonilla (Bierig, 1936, as incarenatus), Sitio Perdido (Bicrig, 1936, as incarenatus), Sierra del Rosario (Bierig, 1936, as incarenatus, truncatus, and caribaeus), Sierra del Grillo (Bierig, 1936, as caribaeus), Aspiro, El Sabicu, and El Rangel (Bierig, 1936, as truncatus), Buenos Aires in Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.), Soledad (Darlington and Leavitt, in M.C.Z. and U.S.N.M.), Baracoa (Darlington, in M.C.Z.). Isie of Pines: Sierra de las Casas (Bierig, 1936, as caribaeus). Jamaica: (Leng and Mutchler, 1917, as apicalis), Hope River (Cameron, 1913, as apicalis), Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Haiti, Port-au-Prince (Darlington, in M.C.Z.), Ennery (Darlington, in M.C.Z. and U.S.N.M.); DomiIniIcAN ReEpuBlic, Sanchez (Darlingten, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: Villalba (Blackwelder station 64), El Yunque (Darlington, in M.C.Z.) St. John: (Erichson, 1840). Guadeloupe: Basse Terre (Fleutiaux and Sallé, 1889, as delauneyi; Leng and Mutchler, 1914; Blackwelder station 81). Dorainica: (Blackwelder station 244A). Specimens examined—I have examined 8 examples (including the type of apicalis) in the collection of Dr. Cameron, 18 in the British Museum, 28 from the Museum of Comparative Zoology, 9 in the United States National Museum, and 15 collected by me in 1935-87. Remarks.—Although I have not compared any of my specimens directly with the type of apicalis, they differ in no way from the original description except to show a slight variability and indicate a wide range. The examples from Dominica and Guadeloupe seem to fit perfectly the description of delawneyi but also are identical 354 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with Puerto Rican examples identified as /abeo Erichson. I have therefore united the three species under the oldest name. Sr. Bierig has kindly sent “cotypes” of his three Cuban species. I find it possible to distinguish the three specimens when considered alone, but the differences are duplicated in several of my series of labeo from other islands and from Cayamas, Cuba. I am forced to conclude that these are merely extremes of a somewhat variable species. These conclusions might be changed by the examination of large series of these staphylinids, but at present I find insufficient evidence to justify keeping them separate. I have collected this species only under Jeaves, either mn rain-forest or in sheltered gullies. 3. STAMNODERUS BERNHAUERI Cameron Stamnoderus bernhaueri CAMERON, 1918b, pp. 383, 334.—Lene and MurcHums, 1917, p. 198.—ScHEERPELTZ, 1933, p. 1237.—Breric, 1936, p. 140.—BLAcK- WELDER, 1939a, p. 115. Stamnoderus dissimilis CAMERON, 1918b, p. 334.—LeENG and MutcHtigr, 1917, p. 198.—ScCHEERPELTZ, 1933, p. 1237.—Burrrie, 1936, p. 140.-—BLACKWELDER, 19389a, p. 115. Description —Rufotestaceous, elytra and abdomen generally testa- ceous. Head with a distinct raised line along the coronal (or epi- cranial) suture; epistomal sutures not marked except by irregular depressions ; clypeus with dense umbilicate punctures, but vertex nearly smooth; labrum transverse with a large broad median anterior lobe. Pronotum with distinct finely raised midline, except at base ; punctures exceedingly fine and situated on small rounded tuberculi, not very dis- tinct; without ground sculpture; lateral impressions rather feeble; sometimes with a more or less well-defined depression at middle of base. “lytra as wide as long, enlarged to apical third; with rather coarse and distinct punctures separated by about their diameter. Abdo- men finely and more or less submuricately punctate. J/ale, seventh sternite shallowly but rather abruptly emarginate at middle; eighth similarly but twice as deeply emarginate; ninth tergite truncate with angles feebly prolonged. Female, eighth sternite broadly expanded posteriorly. Length, 314 mm. Type locality —Jamaica, Hope River district (for both names). Types.—One type and one other specimen in the collection of Dr. Cameron. The unique type of dissimiélis is in the same collection. Records.—The following are the records known to me: Jamaica: (Leng and Mutchler, 1917; Bierig, 1986; Hubbard, in U.S.N.M.), Hope River (Cameron, 1913, for both), Kingston (Blackwelder station 381), Moneague (Darlington, in M.C.Z.), Blue Mountains (Darlington, in M.C.Z and U.S.N.M.). Specimens examined.—I have seen three examples in Dr. Cameron’s collection (including both the types), four from the Museum of Com- MONOGRAPH OF WHST INDIAN STAPHYLINIDAE 300 parative Zoology, one in the United States National Museum, and three collected by me during 1935-87. Remarks.—I have been unable to separate these two species even by the differential characters given by Dr. Cameron. These characters all refer to structures or conditions that vary even among the four examples now before me. The fact that the types were taken at the same place and apparently represent opposite sexes when considered with the variation shown by my small series appears to indicate un- avoidably that the two names represent just one variable species. It is interesting to note here that the other species of this genus from our region are also variable and cannot be separated by color or any charac- ter that shows variation within a single series. My specimens were taken flying at dusk or found incidentally in the net while taking fireflies at night in tall brush. 4. STAMNOBERUS CAMERONI, new species Deseription.—Rufotestaceous, the head and pronotum sometimes a little darker than the elytra. Head without raised lines along coronal suture; epistomal suture not marked except by irregular depressions ; clypeus with rather feeble punctures, not distinctly umbilicate, becom- ing obsolete on the vertex ; labrum transverse with a large broad median anterior lobe. Pronotwm with only a trace of midline; punctured as clypeus; without ground sculpture; lateral impressions feeble; with an irregular depression at middle of base. Hlytra one-ninth longer than wide, nearly parallel; with rather coarse and distinct punctures separated by nearly their diameter. Abdomen finely and densely sub- muricately punctate. Jfale, seventh sternite broadly feebly emargin- ate; eighth with a broadly triangular emargination, about eight times as wide asdeep. Female, sternites not modified. Length, 3 to 4mm. Type locality—Dominican Republic, San José de las Matas; elevation 1,000 to 2,000 feet. Types.—Holotype, male, and five paratypes in the Museum of Com- parative Zoology; four paratypes in the United States National Museum (No. 52518) ; collected in July 1938 by Dr. P. J. Darlington. Records.—Yhe following are the records known to me: Hispaniola: Harri, Port-au-Prince (Darlington, in M.C.Z.); DoMINICAN RE- PUBLIC, Villa Altagracia (Darlington, in M.C.Z.), Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.), San José de las Matas (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—lI have seen only the 10 types. Remarks.—This species is very similar to bernhaueri but differs in the absence of the raised coronal line and the less distinctly umbilicate punctures. I have received no record of its habits. 356 BULLETIN 182, UNITED STATES NATIONAL MUSEUM LIT. Genus STILIPHACIS Bierig Stiliphacis Breric, 1938b, p. 142. Genotype.—S. occipitalis Bierig (original designation and mono- basic). Diagnosis—Body moderately stout; antennae moderately slender, not geniculate; labrum with two large median teeth and sometimes two minute additional ones; maxillary palpus long, third segment enlarged, fourth minute, pubescent; gular sutures united; prothorax nearly circular, not much prolonged anteriorly at middle; proster- num expanded under coxae nearly to hypomera but not connate with the latter; anterior coxal cavities closed behind (by the sternum be- neath the coxae rather than by the hypomera); front coxae large, prominent; middle coxal cavities confluent; posterior coxae “conical,” approximate; abdomen margined, somewhat contracted at base; first and second sternite absent; in general, eighth sternite emarginate in males, rounded in females; fourth segment of tarsi lobed beneath. Remarks.—This genus is known to me from an example sent to me by Sr. Bierig several years ago and four species which appear to be new. The above diagnosis was prepared before the appearance of Bierig’s description. As herein defined this genus differs from Stilicopsis (paradoxa) in having labral denticles and in having the pronotal punctation similar to that of the head, instead of having the punctures replaced by tub- erculi. The genus Dibelonetes is known to me only from an unde- termined species from Nicaragua, but this differs from S#liphacis in the suborbicular shape of the head. Unless additional characters are found, it is likely that these two will be united. I have studied 78 examples from the West Indies. They belong to five species, of which three appear to be new. KEY TO WEST INDIAN SPECIES OF STILIPHACIS 1 labrum: denticulate.at centerso === es ee = Set Se ee eee 2 abrum’ jnot-.dentichilate. 5 2-4 2a Le et See Se exieun 2. Pronotum nearly as wide as long_-----~- gE See Tih hee eT ete aes 3 Pronotum one-third longer than wide__--.-------------.... 5. darlingtoni §..Elytra;reduced, widestiat apex _<-2e) = oc a 4. occipitalis EBlytra, not reduced, narrowed: at apex... so 22222} es eee 4 4. Head completely rounded behind; 4 mm. long__-------------- 2. trinitatis Head truncate behind; 244 mm. long_.-+--------=-_--- ria saat 3. dentatus 1. STILIPHACIS EXIGUA (Erichson) Sunius exiguus EricHson, 1840, p. 647.—KrAatz, 1857a, p. 666.—Brerte, 1938, p. 142, Stilicopsis exigua (Erichson) BerNHAvER and ScHusenrt, 1912, p. 220.—Lene and MutTcHtrr, 1914, p. 404.—Wotcort, 1924, p. 78; 1936, p. 197. Stilicopsis circum/lexrus CAMERON, 1913b, p. 332.—LEN@ and Murcnutirr, 1917, p. 198.—SCHEERPELTZ, 1933, p. 12837.—BLAcKweEtner, 1939a, p. 115. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 304 Stilicopsis auripilis CAMERON, 1913b, p. 3833—Lrena and Murcutrr, 1917, p. 198.—SCHEERPELTZ, 1933, p. 1237.—BLACKWELDER, 1939, p. 115. Stiliphacis exigua (BErichson) Breric, 1938a, p. 142. Description.—Rufopiceous, elytra and abdomen testaceous, elytra usually with a black lateral spot. Head large, subquadrate, as wide as elytra; eyes large, prominent; sides behind eyes straight for less than length of eyes, then abruptly constricted; closely but coarsely umbilicately punctured ; antennal segments transverse from the third; labrum feebly broadly lobed at center, on each side of lobe with a minute tooth. Pronotum scarcely wider than long, seven-eighths as wide as head and elytra, widest at anterior fourth, strongly narrowed and rounded in front, feebly narrowed posteriorly; sides finely tuber- culate; punctation similar to that of head. Prostermwn rather indis- tinctly longitudinally carinate on the disk. //ytra nearly as long as broad, a little Jonger than pronotum; moderately but densely punctate. Abdomen shining, indistinctly punctate, pubescent. fale, seventh sternite with a broad shallow emargination; eighth broadly and deeply emarginate. Length, 214 to 3 mm. Type locality—Puerto Rico. Of circumslexus, “Mustique I., Grena- dines, W. I.”; of auripilis, Mount Gay Estate, Grenada. Types.—tIn either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. The types of cirewnflewus and auripilis are in the British Museum. Records.—The following are the records known to me: Puerto Rico: (Erichson, 1840; Leng and Mutchler, 1914; Wolcott, 1924), Jayuya (Wotleoctt, 19386). Grenadines: Mustique (Cameron, 1913, ag circumflerus; Leng and Mutchler, 1917, as circumflerus). Grenada: (Cameron, 1913, as auripilis; Leng and Mutchler, 1917, as auripilis). Trinidad: St. Angustine (Weber, in M.C.Z.). Specimens examined.—I have examined 48 specimens in the British Museum, 5 in the collection of Dr. Cameron, and 1 from the Museum of Comparative Zoology. Remarks.—I am unable to accept as of specific importance the char- acters mentioned by Dr. Cameron to separate his specimens from Grenada and Mustique. A specimen of each series that I borrowed from the British Museum for further study differ in degree from the descriptions in various points and seem to be inseparable by any char- acter other than depth of color and extent of infuscation of the elytra. These differ in no way from the description of S. exigua (Krichson) except those with the elytra infuscate laterally. I found no specimens labelled exigua in either the British Museum or Dr. Cameron’s collec- tion and therefore conclude that he overlooked this name. T find no record of the habits of this species. 358 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. STILIPHACIS TRINITATIS, new species Description.—Rufopiceous, apical spot of elytron and penultimate sternite testaceous. Head ovate, hind angles completely rounded, base emarginate above neck; eyes separated from base by a little more than their length; labrum with two slender denticles, longer than distance between them, these flanked by a pair of smaller denticles; umbilicate punctures coarse and dense, rarely uniting, often irregular in shape; without ground sculpture on the intervals. Pronotum about one- seventh longer than wide, not prolonged in front, widest at anterior third, sides rather strongly converging to narrowly rounded basal angles; sides rather coarsely tuberculate; umbilicate punctures a little smaller than those on head, occasionally uniting but not definitely in longitudinal rows; without ground sculpture. Zlytra a little wider than long, one-fourth longer than pronotum; with rather coarse punc- tures separated by about one-third their diameter by feebly convex intervals; without distinct ground sculpture. Abdomen finely and obsoletely submuricately punctate; surface uneven. Male, unknown. Female, sternites unmodified. Length, 324 mm. Type locality —Trinidad, Mount Tucuche. Types.—Holotype, female, in the Museum of Comparative Zoology, collected in April 1929 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Trinidad: Mount Tucuche (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—In addition to its unusual size, this species may be dis- tinguished by its labrum and its rounded head. I have received no record of its habits. 3. STILIPHACIS DENTATUS, new species Description —Rufotestaceous with elytra a little paler to piceous with elytra testaceous and marked with black on the disk and apically along the suture. ead moderately large, suborbicular, one-fifteenth wider than elytra; eyes moderate, not very prominent, at about their length from base; sides behind eyes rather broadly rounded into base ; umbilicate punctures distinct and dense, frequently irregular in shape, without ground sculpture on the intervals; labrum at middle with two acute teeth much longer than distance between them, and flanked by a pair of smaller ones. Pronotwm about as long as wide, not prolonged in front, about three-fourths as wide as elytra, widest at anterior third, strongly narrowed in front, moderately narrowed behind to rounded angles; sides coarsely tuberculate; umbilicate punctures very irregular but distinet; without ground sculpture. /lytra about as wide as long, nearly one-fourth longer than pronotum; with moderate slightly elongate punctures separated by their diameter or less; 1n- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 359 tervais convex; without ground sculpture. Abdomen with fine but obsolescent submuricate punctures. Male, eighth sternite broadly and feebly emarginate, the apex of the emargination interrupted by a feeble broad lobe of the sclerite. Female, unknown. Length, 3 mm. Type locality —Grenada, 1 mile south of Sauteurs, at the northern end of the island. Types.—Holotype, male, in the United States National Museum, No. 52391, collected by me on January 13, 1936. Twelve paratypes are in the British Museum. Records.—The following are the records known to me: St. Vincent: (British Museum). Grenada: (Blackwelder station 137; British Museum). Specimens examined.—I have seen only the 13 types. Remarks.—The type specimen differs from extqua (auripilis and circumpexus of Cameron) in the strongly dentate labrum and rounded basal angles of head. In labral structure it is close to the prevailing conception of Dibelonetes. The paratypes were placed in the British Museum under the name Didelonetes stigma Erichson. A specimen of the latter from Brazil appears to be larger, with coarse punctation, and has a much larger elytral black spot than our species. The type was found in decaying cocoa pods. 4. STILIPHACIS OCCIPITALIS Bierig Stiliphacis occipitalis Breric, 1938b, pp. 141, 142, figs. 3-5. pp. 115, 122. Stiliphacis truncata Birria, MS. 3LACK WELDER, 1939a, Description—Testaceous. MHcad moderately large, subhexagonal, including the eyes as wide as elytra; eyes rather prominent, separated from base by about their length; sides behind eyes feebly converging, angles rather broadly rounded ; umbilicate punctures large and dense, frequently irregular in shape, without ground sculpture on the in- tervals; labrum transverse, at middle with two acute teeth about two- thirds as long as labrum, longer than distance between them, and a pair of minute denticles outside them. Pronotum a little longer than wide, not prolonged in front, five-sixths as wide as elytra, widest at anterior third, strongly narrowed in front, moderately narrowed behind to rounded angles; sides indistinctly tuberculate; umbilicate punctures as on head, irregular and often uniting in short grooves; without ground sculpture. //ytra one-fifth wider than long, as long as pro- notum; with rather coarse impressed punctures separated by feebly convex intervals; without ground sculpture. Abdomen sparsely and rather indistinctly submuricately punctate. Mo/e,unknown. Female, eighth sternite rounded. Length, 2 mm. Type locality—Cuba, Sierra del Rosario. Types.—tIn the collection of Alexander Bierig. 360 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records—The following is the only record known to me: Cuba: Sierra del Rosario (Bierig, 19388; Bierig, in Blackwelder collection), Specimens examined.—I have seen only one example in my own col- lection, received from Sr. Bierig some years ago under the name truncata. Remarks.—This species is very similar to dentatus but seems to be distinct. It can apparently be recognized by its small size and pale color as well as by the reduced elytra. I find no record of its habits. 5. STILIPHACIS DARLINGTONI, new species Description —Castaneous, somewhat picescent especially in front, elytra paler. ead ovate, wider than the elytra; eyes prominent, at one-half more than their length from base; sides completely rounded into base which is emarginate above the neck; labrum with two slender teeth a little longer than distance between them, flanked by a pair of small denticles; with moderately large aud dense umbilicate punctures, frequently united in short irregular channels; without ground sculp- ture on the intervals. Pronotum ene-third longer than wide, widest at middle, sides behind nearly straight, basal angles moderately rounded ; sides indistinctly tuberculate ; punctures as on head but not as flat-bottomed, only occasionally united; without ground sculpture. Elytra barely wider than long, seven-eighths as long as pronotum, nar- rower than abdomen or head; with rather coarse impressed punctures separated by feebly convex intervals; without distinct ground sculp- ture. Abdomen with coarse punctures basally on each segment, be- coming finer toward apex. Jale, eighth sternite truncate. Female, eighth sternite prominently rounded. Length, 2% to 3 mm. Type locality —Jamaica, main range of the Blue Mountains, eleva- tion 5,000 to 7,388 feet. Types.—Holotype, male, and seven paratypes in the Museum of Comparative Zoology; six paratypes in the United States National Museum (No. 52519) ; collected on August 17-19, 1934, by Dr. P. J. Darlington. Records—The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 14 types. Remarks.—This species is distinct by the punctures of the pronotum differing in appearance from those of the head, as well as by the coarse punctures of the abdomen, the reduced elytra, and the feeble male characters. I have received no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 361 LIiI. Genus SUNIOSAURUS Bierig Suniosaurus BiExie, 1988b, p. 139. Genotype.—S. cuadriceps Bierig (original designation and mono- basic). Diagnosis.—Body slender, elongate; antennae filiform, not verticil- late, posteriorly flexile at first joint; labrum with two long, slender, hornlike processes within a slight emargination; third segment of maxillary palpus much enlarged, fourth minute; gular sutures appar- ently united behind from near middle; neck about two-fifths as wide as head; pronotum quadrate; hypomera divided by a diagonal line descending from base to middle of side; prosternum longitudinally carinate at middle, dilated under coxae nearly as far as the hypo- mera; anterior coxae large, prominent; mesosternum not carinate ; mid- dle coxal cavities confluent; abdomen strongly margined; sternites of first and second segments not present; front femora dilated; tarsi 5-segmented, slender, fourth tarsomere simple. Remarks.—These characters are taken from the specimen collected by N. A. Weber at Soledad. Sr. Bierig’s diagnosis was very brief but does not conflict with the above except in regard to the gular sutures, which he described as obsolete. They are very clear in the specimen examined by me. There is some question as to whether this genus will key out prop- erly in my key to the Paederini (Blackwelder, 1939), since the pros- ternal process is expanded under the coxae only about halfway to the hypomeral edge. It seems clearly to belong in the present series, however, rather than in the Medon-group. This genus was said to be intermediate between Suniocharis and Suniotrichus, but my study of these two genera shows them to be very widely separated in the subfamily, the former being close to Dibel- onetes and lM chiaster, the latter being close to Thinocharis. My exam- ination indicates that it probably is more nearly related to Suniocharis and will stand next to that genus in the key. It may be distinguished readily by the size and shape of the Jabral denticles. I have seen only one specimen, which seems to belong to the geno- type species. 1. SUNIOSAURUS CUADRICEPS Bierig Suniosaurus cuadriceps Brerie, 1938b, pp. 140, 148.—BLacKWELDER, 1939a, p. 122. Description.—Rufocastaneous. Head subquadrate; eyes at about twice their length from base; antennae long and slender, no seg: ments transverse; with moderate indistinctly umbilicate punctures almost completely obscured by dense scaly ground sculpture. Pro- notum as wide as head, as broad as long, sides rounded into base; without impunctate midline; without punctures but sculptured as 449008—42-_—-24 362 BULLETIN 182, UNITED STATES NATIONAL MUSEUM head but more densely and finely; with short pubescence. Hlytra nearly as broad as head and pronotum, nearly one-fourth longer than broad; with rather dense rounded setiferous tuberculi, with rather indistinct but granulose ground sculpture between. Abdomen with fine and dense but obsolescent submuricate punctures, without definite ground sculpture. J/a/e, unknown. VYemale, apical tergites and sternites unmodified. Length, 4 mm. Type locality—Cuba. Types.—tin the collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: La Lisa (Bierig, 1958}, El Mariel (Bierig, 1938), Sierra de Anafe {Bierig, 1938), Aspiro (Bierig, 1988), Soledad (Weber, in M.C.Z.). Specimens examined —I\ have seen only the one specimen, from Soledad in the Museum of Comparative Zoology. Remarks.—This specimen is assigned to this species without much hesitation. The description given above is that of my specimen and has not been compared with the type or changed to fit Bierig’s description. I have some doubt as to the sex of the specimen I have seen, al- though it has no apparent modifications of the abdomen. Sr. Bierig did not record the sex of his specimens. The type was found under fallen leaves, and Mr. Weber’s example was collected in a Berlese funnel. LIV. Genus SUNIOCHARIS Sharp Suniocharis SHARP, 1886, p. 586. Genotype.—S. modesta Sharp (designated by Blackwelder, 1939). Diagnosis.—Body slender; antennae short, flexile at first joint; labrum indistinctly bidenticulate; mandibles short, acutely triden- tate; third segment of maxillary palpus much enlarged, fourth minute; gular sutures narrowly separated, feebly converging pos- teriorly at least in front; neck very slender; hypomera divided by a diagonal line descending from base to apex; prosternum longitu- dinally carinate at middle, dilated posteriorly under coxae as far as the hypomera but not connate with it; anterior coxae large, promi- nent; mesosternum not carinate; middle coxal cavities confluent ; abdomen strongly margined; sternites of first and second segments not present; sternite of third segment longitudinally carinate at base; front femora dilated; tarsi 5-segmented, very slender, fourth tarsomere simple. Remarks—Bernhauer has described a subgenus of this as Para- suniocharis (1933, pp. 519, 520), with genotype Suniocharis (Para- suniochaiis) boxi Bernhauer (monobasic and designated). I have MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 303 seen no examples and am unable to state whether it belongs here or not. Sharp’s original description of the tarsus is as follows: “The terminal joint is, however, inserted above the fourth joint very near the base of the third; as the fourth joint is placed underneath the terminal one, and is not wider than it, the foot appears to be 4-jointed.” I have not been able to verify this on my single speci- men. The tarsi there are very distinctly slender, the segments de- creasing In size from the first to the fourth, and with the apex of each only a little diagonally truncate. I have seen only three specimens of a species that appears to be distinct from the previously known species. 1. SUNIOCHARIS VOLANS, new species Description.—Rufopiceous, pronotum rufous, elytra indefinitely testaceous at apex. Head subquadrate, truncate behind, the angles right; eyes small, at much more than their length from base; an- tennae short and stout, segments 7 to 10 transverse, 11 longer than 9 and 10 together; punctate but with fine and rather dense tuber- culi somewhat obscured by indefinite ground sculpture; clypeus trun- cate in front. Pronotwm as wide as head, one-eighth wider than long, obtrapezoidal, widest at front, feebly narrowed behind, angles moderately narrowly rounded ; with feebly depressed midline basally ; with tuberculi as on head or a little coarser; with short pubescence. Elytra about as wide as long, one-sixth longer than pronotum; sur- face densely covered with tiny elongate tuberculi, arranged longitu- dinally more or less in series, shining surface visible between; with rather long pubescence. Abdamen rather densely, finely, and trans- versely serially submuricately punctate, less distinct and dense apically. fale, unknown. Yemale, apical tergites and_ sternites unmodified. Length, 3 mm. Type locality —Trinidad, between Sangre Grande and Sans Souci by way of Toco, parishes of St. Andrew and St. David. Types.—Holotype, female, U.S.N.M. No. 52395, collected by me on December 15, 1935; two paratypes, female, in Museum of Compara- tive Zoology and United States National Museum, collected in 1935 by N. A. Weber. Records.—The following are the records known to me: Trinidad: Sangre Grande (Blackwelder station 100B), Maracao Valley (Weber, in U.S.N.M.), San Rafael (Weber, in M.C.Z.). Specimens examined.—t\ have seen only the three types. Remarls.—This species differs from the Central American species described by Sharp in its much smaller size, stouter antennae, and probably other characters not mentioned in the descriptions. The type was caught flying at dusk. 364 BULLETIN 182, UNITED STATES NATIONAL MUSEUM LV. RONETUS, new genus Genotype —R. ortinolus Blackwelder (designated here). Diagnosis—Slender and elongate, moderately constricted; anten- nae posteriorly flexile, not strongly geniculate, basal segment not much elongate, some apical segments transverse; eyes prominent; head not grooved behind eyes; labrum short and broad, with four large equal triangular teeth; gular sutures united throughout; third segment of maxillary palpus enlarged, elongate, fourth minute; neck less than one-fourth as wide as head; prothorax elongate but not greatly prolonged in front; prosternum expanded under the coxze and connate with the hypomera; hypomera not much lobed behind the coxae; anterior coxal cavities open above but closed below by the sternal processes; anterior coxae large, exserted; middle coxal cavi- ties confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; tarsi 5-segmented, slender, fourth seg- ment not lobed beneath; integuments all densely coarsely tuberculate in appearance. Remarks.—This is one of our most distinct genera because of the coarse sculpture. The tuberculi seem to bear one hair and to be covered with a white precipitate, but I have not been able to wash this off by any means. It would key out to or near A/yrmecosaurus in my key to the Paederini (Blackwelder, 1939) but is distinguished at once from that peculiar genus in lacking the ridges at the sides of the head, pronotum, and elytra. In most other respects these two genera are rather similar. I have seen only one example, the genoholotype. 1. RONETUS ORTINOLUS, new species Description.—Castaneous or piceous, color obscured by dense cover- ing of whitish secretion or precipitate. Mead semicircular behind, nearly one-half longer than wide, unusually convex; eyes prominent, at one-half more than their length from base; antennae filiform to fifth joint, five penultimate segments moniliform; surface through- cut with indistinct but dense and coarse verrucose sculpture, each prominence bearing an anteriorly-projecting seta and covered with a flaky deposit. Pronotwm one-half longer than wide, three-fourths as wide as head, widest at middle, sides straight in front and be- hind, basal angles narrowly rounded; with sculpture as on head. Elytra about one-sixth longer than wide, one-tenth longer than pro- potum; sculptured as head but without the obvious setae. Abdomen sculptured as elytra. Sexes not distinguished. Length, 4 mm. Type locality —Trinidad, La Ortinola Estate, Maracas Valley. Types.—Holotype, sex unknown, in the Museum of Comparative Zoology, collected on September 18, 1935, by N. A. Weber. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 365 Records.—The following is the only record known to me: Trinidad: Maracas Valley (Weber, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—I know of no other staphylinid that can be confused with this. It differs from Myrmecosaurus by the shape of the head and the absence of ridges and from all others by the verrucose sculpture. I have received no record of its habits. LVI. Genus ASTENUS Stephens Astenus STEPHENS, 1832, p. 275. Sunius ERIcHSON, 1839a, p. 528 (not Stephens, 1829). Subgenus Mecognathus Wo LLaAsToNn, 1854, p. 595. Suniogaster Reirrer, 1909, p. 151. Subgenus Neognathus Ssarp, 1874, p. 69. Subgenus Astenognathus RettTeR, 1909, p. 150. Subgenus Hurysunius Retrrer, 1909, p. 149. Genotypes.—Staphylinus angustatus Fabricius=Astenus angus- tatus (Fabricius) (designated by Westwood, 1838); the designation of brunneus Stephens by des Gozis, 1886, cannot stand; the designa- tion of Blackwelder, 1939, was made in ignorance of that of West- wood and in oversight of that of des Gozis. Of Sunius, Staphylinus angustatus Fabricius=Sunius angustatus (Fabricius) (designated by Blackwelder, 1939); of Astenognathus, Sunius bimaculatus Erichson=Astenus (Astenognathus) bimaculatus (Erichson) (desig- nated by Blackwelder, 1939); of Hurysunius, Sunius paradowus Eppelsheim= Astenus (Furysunius) paradoxus (Eppelsheim) (desig- nated by Blackwelder, 1939); of Mecognathus, M. chimaera Wollaston (monobasic); of Swniogaster, Sunius ampliventris Reitter=Astenus (Suniogaster) ampliventris (Reitter) (monobasic) ; of Neognathus, N. angulatus Sharp (monobasic). Diagnosis—Very slender and elongate, moderately convex; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, slender; eyes prominent; head not grooved be- hind the eyes; labrum short and very wide, bidentate; gular sutures united throughout; third segment of maxillary palpus enlarged but not subspherical, fourth subulate; neck not very slender; prothorax not prolonged in front; prosternum prolonged posteriorly between and under coxae and laterally as far as the hypomera with which it is connate; hypomera somewhat prolonged in a lobe behind the coxae; anterior coxal cavities open behind above but closed beneath by the sternal processes; anterior coxae large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first 366 BULLETIN 182, UNITED STATES NATIONAL MUSEUM and second abdominal sternites absent, seventh generally not modi- fied, eighth emarginate in male; tarsi 5-segmented, fourth segment lobed beneath. Remarks.—This large genus is well represented in all parts of America. It is easily recognizable by its slender form and apically enlarged abdomen. The subgenera that have been named are not sufficiently well known to me to permit me to place the West Indian species in their proper positions. I have seen from the West Indies 57 specimens, representing 3 species, KEY TO WEST INDIAN SPECIES OF ASTENUS 1. Supraantennal ridge joining upper margin of eye as a distinct smooth line. 3. juvus Supraantennal ridge not reaching to wpper margin of eye_-------------__ 2 2 Miytramacniatevor Dicoloreds 2-2.) esas ae oa ee eee 2. cubensis Elytra not maculate, unicolorous________________- 8 nee 1. cinctiventris 1. ASTENUS CINCTIVENTRIS (Sharp) Bunius cinctiventris SHarp, 1886, p. 601. Astenus cinctiveniris (Sharp) BeERNHAUER and SCHUBFR?T, 1912, p. 215.—Bieric, 1933, p. 517. Description.—Rufotestaceous to testaceous, usually with indefinite piceous or cloudy areas on head and pronotum, and seventh abdominal tergite piceous except at apex. Head slightly longer than broad; eyes at about their length from the neck; sides evenly and strongly rounded to base; with dense umbilicate punctures indistinct, obscured in front by ground sculpture; a band of ground sculpture separating the punctures from the front edge of clypeus; supraantennal ridge disappearing before the eye; beneath with rather coarse abrupt punc- tures; ground sculpture strong in front but vague posteriorly; united gular suture scarcely at all depressed at base. Pronotum two-sevenths longer than wide, about seven-eighths as wide as head, widest at an- terior three-eighths, sides nearly straight in front, feebly arcuate and moderately converging behind; with umbilicate punctures dense but indistinct and frequently united longitudinally. Hlytra widest at middle, narrowed to both ends; with moderate asperate punctures, moderately dense but not serial. Abdomen moderately and densely somewhat serially muricately punctate. Male, eighth sternite with a small triangular emargination; ninth tergite with a deep narrowly triangular notch between short processes. Female, unknown. Length, 3 to 4 mm. Type locality —Guatemala. T'ypes.—A series in the British Museum (one labeled type). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 367 Records.—The following are the records known to me: Hispaniola: DOMINICAN REPUBLIC, San José de las Matas (Darlington, in M.C.Z.), Sanchez (Darlington, in U.S.N.M.). St. Lucia:(H. E. Box, as Blackwelder station 444B). St. Vincent: (British Museum). Grenadines: Mustique (British Museum). Grenada: (British Museum). Central America: GUATEMALA, Mrxico, PANAMA (Sharp, 1886), Costa Rica (Bierig, 1933). Specimens examined.—I have seen 38 examples from the West Indies in the British Museum, 2 from the Museum of Comparative Zoology, and 4 collected by Dr. H. E. Box in St. Lucia. Remarks.—1 was unable to distinguish the West Indian examples from the Guatemalan type of this species in the British Museum. It is probably a more widespread species than our records yet indicate. Dr. Box collected his specimens from decomposing trash in sugar- cane fields. 2. ASTENUS CUBENSIS, new species Description.—Rufous or rufotestaceous, sometimes with abdominal segments piceous, disk of head and pronotum infuscate, and elytra with a lateral black spot of variable size. Head a little longer than wide; eyes at about their length from neck; sides evenly and strongly rounded to base; with dense umbilicate punctures above extending to smooth margin of clypeus in front; with indistinct ground sculpture on ridges between punctures, especially in front; supraantennal ridge disappearing in front of eyes; beneath with rather coarse punctures, each one ill-defined anteriorly; ground sculpture confined to area near mouth parts; united gular suture rather depressed at base. Pronotum one-fourth longer than wide, seven-eighths as wide as head, widest at anterior third, sides straight and strongly converging in front, feebly arcuate and moderately converging behind; with dense umbili- cate punctures, seldom united or much elongate; without ground sculpture. Zlytra narrowed posteriorly, strongly convex; with asper- ate punctation, moderately dense, sometimes appearing to be in transverse rows. Abdomen rather strongly and densely, somewhat serially, muricately punctate. Male, eighth sternite with a moderate triangular notch at apex; ninth tergite with a rounded emargina- tion somewhat deeper than wide between two acute processes. [e- mile, tergites and sternites unmodified. Length, 3 to 314 mm. Type locality —Cuba, Cayamas, about 15 miles west-northwest of Cienfuegos, southern Santa Clara Province. Types.—Holotype, male, and three paratypes, U.S.N.M. No. 52480, collected in January by E. A. Schwarz. 368 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records —The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Jatibanico (Christenson, in U.S.N.M.), Limones (W. M. Mann, in U.S.N.M.), Soledad (Weber, in M.C.Z.). Jamaica: Blue Mountains (Darlington, in M.C.Z.). Specimens examined.—I have seen four examples beside the types in the United States National Museum, and two in the Museum of Comparative Zoology. Remarks.—This species is normally one of the ones with maculate elytra, but even in our eight specimens there is great variation, the dark spot being almost completely absent in two cases and covering the entire basal two-thirds in another. I have no record of the habits of this species except that the Soledad example was collected in a Berlese funnel. 8. ASTENUS JUVUS, new species Description —Rufotestaceous, apex of abdomen feebly picescent. Head about one-seventh longer than broad; eyes at a little more than their length from base; scarcely narrowed behind eyes, basal angles moderately rounded; with very dense umbilicate punctures above ob- scured by ground sculpture in front; supraantennal ridge clearly uniting with upper margin of eye; beneath with rather coarse abrupt punctures; ground sculpture visible in certain lights on all of the ventral aspect; united gular sutures somewhat impressed near base. Pronotum nearly one-fifth longer than wide, about as wide as head, widest on anterior third, sides feebly arcuate and strongly converging anteriorly, nearly straight and feebly converging posteriorly; with dense umbilicate punctures; with indistinct ground sculpture on the ridges. Hlytra widest at middle, narrowed to both ends; with coarse asperate punctation, moderately dense. Abdomen finely and moder- ately sparsely, somewhat serially, submuricately punctate. fale, un- known. Female, eighth sternite rounded. Length, 4 mm. Type locality —Trinidad, Montserrat, in county of Caroni. Types.—Holotype, female, U.S.N.M. No. 52427, collected by August Busck. fecords.—The following is the only record known to me: Trinidad: Montserrat (Busck, in U.S.N.M.). Specimens ewamined.—I have seen only the unique type. Remarks—This species seems to be distinct from the true West Indian species by the character of the supraantennal ridge. I am not able to determine whether it is distinct from all the South American species, however. I find no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 369 LVII. Genus ECHIASTER Erichson Echiaster ExtcHson, 1840, p. 636. Subgenus Leptogenius Casry, 1886b, p. 214. Genotypes.—E’. longicollis Erichson (designated by Duponchel, 1841). Of Leptogenius, L. brevicornis Casey (monobasic). Diagnosis —Body moderately elongate and strongly constricted, with very dense sculpture; head not grooved behind eyes; vertex not carinate; eyes large, prominent; antennae posteriorly flexile, not strongly geniculate, basal segment not much elongate, first and second not enlarged with rest slender; labrum moderately large, with four triangular denticles; third segment of maxillary palpus enlarged but not subspherical, fourth segment subulate; gular sutures united; neck slender; prothorax not elongate or prolonged in front; prosternum dilated under front coxae and connate with hypomera ; hypomera feebly lobed behind the coxae; anterior coxal cavities open behind above but closed beneath by the prosternal processes; anterior coxae large, ex- serted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent, eighth sternite sometimes broadly emarginate in male; tarsi 5-segmented, fourth seg- ment not strongly lobed beneath. Remarks.—This genus is confined to temperate and_ tropical America. It is distinct by its dense sculpture and quadridentate labrum. I am unable to distinguish Leptogenius Casey as more than a subgenus. The record of E'chiaster solitarius Sharp from Jamaica by Gowdey, 1926, is probably a misidentification. Until the specimens can be reexamined, it must be regarded as a doubtful record. I have examined 63 specimens from the West Indies, which I assign te 6 species of which 1 appears to be new. KEY TO WEST INDIAN SPECIES OF ECHIASTER 1. Elytra with intervals between punctures smooth and shining. 6. impressicollis Elytra densely punctate, intervals not shining or distinct__.._--_--______ 2 2 wyes large, at less’ than their length from base .--_ == 3 Eyes small or moderate, at their length or more from base____-_________ es 8. Elytra black, obscurely lighter at the shoulders; head densely granularly BCUUPEUT EG seta LSE LU Bee hers OL a DES ee ee ee 4. sharpi Hliytra testaceous with apical third black; head densely and finely umbili- CATO PUNCHING Cet aE oe at ee ae ee ee 5. buphthalmus 4. Byes moderate, at about their length from base____-__-__________ Ses 5 Eyes very small, at twice their length from base___._.___----__- 1. microps 5. “Dull pitechy black; elytra obscurely lighter at the shoulders”__ 2. distinctus PELriUSiMOUSH MOG AS a DOV. es ee ae eS ee es) 3. waterhousei 370 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 1. ECHIASTER MICROPS, new species Description—Rutous to rufotestaceous, head, pronotum, and abdo- men sometimes picescent in part, elytra with apical half black. ead moderately large, subquadrate; eyes very small, at nearly twice their length from base; no punctures visible, but sculpture very dense and granulose and tuberculose; antennae short, segments 7 to 10 trans- verse. Pronotum nearly four-fifths as wide as head, as broad as long; widest at anterior third, thence abruptly narrowed to neck, moder- ately narrowed to base; disk longitudinally impressed on each side of midline, leaving a distinct elevated convex ridge; sculpture similar to that of head. F/ytra as broad as long, one-third longer than pro- notum; sculpture similar to that of head and pronotum but with very fine pubescence. Abdomen near base wider than elytra, narrower posteriorly; with moderately fine but not dense submuricate punctures; pubescence rather long apically. fale, eighth sternite feebly broadly emarginate. Female, eighth sternite broadly rounded. Length 114 to 2144 mm. Type locality —Montserrat. Types.—Holotype, female, and one paratype, female, U.S.N.M. No. 52408, collected in February and March by H. G. Hubbard. Records.—The following are the records known to me: [Cuba: Cayamas (Schwarz, in U.S.N.M.).] Montserrat: (Hubbard, in U.S.N.M.). Specimens examined.—\ have seen the two types and one other specimen, Remarks.—The Cuban example is placed with this species with some doubt. It is considerably larger but seems to differ otherwise only in the less distinct pronotal impressions and slightly more rounded pronotum. The sculpture of the forebody has been examined with a binocular microscope at a magnification of over 200 diameters. At this and all lower magnifications it appears to be densely granulose and tuber- culose. I find no record of the habits of this species. 2. ECHIASTER DISTINCTUS Cameron Echiaster distinctus CAMERON, 1913b, p. 388.—LENG and MuTcHLER, 1917, p. 198.— SCHEERPELTZ, 1933, p. 1238. Description——Dull pitchy black, elytra obscurely reddish testaceous at the shoulders, abdomen ferruginous at apex. Head large, subquad- rate, longer than broad, posterior angles rounded; eyes rather small; temples large, longer than diameter of the eyes; punctation close, rugose, and umbilicate; antennae short, segments 6 to 10 transverse. Pronotum narrower than head and elytra, broadest at anterior angles, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 371 which are rounded, posterior angles obtuse; scarcely longer than wide, carinate posteriorly on midline; sculptured as head. “lytra as long as broad, closely and rugosely punctured. Abdomen closely punctate and shortly pubescent. Length,244 mm. (From Cameron.) Type locality— Jamaica, Wag Water River. Ty pes.—Unique type in the collection of Dr. Cameron. Records.—The following is the only record known to me: Jamaica: (Leng and Mutchler, 1917), Wag Water River (Cameron, 1913). Specimens examined. —I have seen only the type, which I examined briefly in Dr. Cameron’s collection. Remarks.—A though I cannot say from direct examination, it seems possible that an error was made in the original description of the sculpture of the head and pronotum. In the description of this species and of waterhousei the sculpture of the head and pronotum is described as dense, rugose, and umbilicate. An examination of the latter species (cotypes sent froma the British Museum) shows that this sculpture is not at all umbilicate (as defined by Cameron, 1930, p. x) but is coarsely granulose or tuberculose. I find this appearance of dense granulation under all magnifications from 21 to 204 times. From this I conclude that the sculpture of déstinctus is probably similarly granulose instead of umbilicate as described. Furthermore, on waterhousei there is no sign of rugosity, at least in the sense of ridges or wrinkles. I find no record of the habits of this species. 8. ECHIASTER WATERHOUSE! Cameron Echiasier waterhouset CAMERON, 1913b, p. 8836—LeNa and MUTCHLER, 1917, p. 198.—ScHEERPELTZ, 1933, p. 1289.—BLACKWELDER, 1939a, p. 116. Description —Rutotestaceous, often more testaceous behind, apical alf of elytra black or infuscate, abdomen infuscate before the apex. Head moderate, subquadrate ; eyes moderate, at about their length from base; impunctate but densely granulose and tuberculose; antennae short, segments 5 to 10 transverse. Pronotum nearly five-sixths as wide as head, as long as broad; widest at anterior two-fifths, strongly nar- rowed and slightly produced in front, moderately narrowed behind; disk longitudinally impressed basally along the midline; sculpture similar to that of head. Hlytra as broad as long, two-fifths longer than pronotum; sculptured similarly to pronotum but a little less densely and sometimes vaguely in longitudinal series. Abdomen some- what fusiform; with fine and moderately dense submuricate punctures on the four basal tergites, apical tergites with coarse muricate punc- tures bearing long black hairs. Length, 2 to 214. Type locality —Grenada, Chantilly Estate. Types.—Fourteen specimens (one labeled type) in the British Museum; five in the collection of Dr. Cameron (one labeled cotype) ; 372 BULLETIN 182, UNITED STATES NATIONAL MUSEUM six in the United States National Museum (one labeled cotype and paratype, No. 50874). Records.—The following are the records known to me: [Cuba: Marianao (Cameron collection, as minutus Sharp.).] Jamaica: (Cameron collection), Trinityville (Blackwelder station 428), Troy (Blackwelder station 409). Hispaniola: DoMInIcAN Repusiic, SAinchez (Darlington, in M. ©. Z.). St. Vincent: (Cameron, 1913). Grenada: (Cameron, 1913; Leng and Mutchler, 1917; Blackwelder station 137). Specimens examined—I have seen 14 examples in the British Museum, 6 in the collection of Dr. Cameron, 1 from the Museum of Comparative Zoology, 6 in the United States National Museum, and 5 collected by me in 1935-37. Remarks.—The single specimen labeled minutus in the Cameron collection is very likely this species. T was not able to compare it with the type of ménutus in the British Museum but find that the descrip- tion of minutus is insufficient to separate it from waterhouset. The discontinuous distribution indicated above from the original publication is verified by my own captures. Further collecting will undoubtedly bring many new records. My specimens were collected from decaying cocoa pods. 4. ECHIASTER SHARPI Cameren Hchiaster sharpi CAMERON, 1913b, p. 337, 8338.—Lrna and Murcrtier, 1917, p. 198.— SCHEERPELTZ, 1983, p. 1239. Description—Head and pronotum ferruginous, elytra black, ob- scurely lighter at the shoulders, abdomen black, apex obscurely lighter. Head large, orbicular; eyes large, their diameter much greater than the length of the temples; sculpture close, rugose, umbilicate; antennae short, segments 6 to 10 transverse. Pronotwm narrower than head, dis- tinctly longer than broad, oval-oblong, carinate at center (with a raised central line), anterior angles rounded, posteriorly obtuse; sculp- tureason head. #7/ytra distinctly longer than broad, and little longer than the pronotum; closely and rugosety punctured. Abdomen nar- rowed at apex; closely punctate and shortly pubescent. Length, 3 mm. (From Cameron.) Type locality —Jamaica, Hope River district. Types.—Unique type in the collection of Dr. Cameron. Records —The following is the only record known to me: Jamaica: (Leng and Mutchler, 1917), Hope River (Cameron, 1913). Specimens examined.—I have seen only the unique type, which I examined briefly in Dr. Cameron’s collection. Remarks.—The remarks concerning punctation under distinctus apply equally to this species. I find no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Sie 5. ECHIASTER BUPHTHALMUS Cameron Echiaster buphthalmus CAMERON, 1913b, p. 337.—LeENe and MurcHter, 1917, p. 198.—ScHEERPELTz, 1933, p. 1238.—BLacKWELDER, 1939a, p. 116. Description —Rufotestaceous, elytra paler with apical third black, apex of abdomen piceous. Head moderately large, suborbicular; eyes large, at about half their length from base; very densely and very finely umbilicately punctured throughout; antennae short, segments 7 to 10 transverse. Pronotwm about five-sevenths as wide as head, one-fifth longer than wide; widest at anterior third, strongly narrowed in front, moderately narrowed behind; with just a trace of umbilicate sculp- ture as on head but surface appearing (with magnification of 85 times) densely granulose and tuberculose **; with a longitudinal impression on each side of midline, more separated than usual. Hlytra as broad as long, one-sixth longer than pronotum; densely granulose and tuber- culose, with trace of longitudinal series but not rugose. Abdomen nar- rowed posteriorly; at base densely granulose, becoming less distinct apically ; seventh to ninth tergites with dense black hairs arising from small submuricate punctures. Length, 214 to 3 mm. Type locality.—St. Vincent. Types—F our specimens (one labeled type) in the collection of Dr. Cameron; a series of 12 examples in the British Museum also contains a specimen labeled type, from Mustique. Records—The following are the records known to me: Antigua: (Blackwelder station 275A). St. Vincent: (Cameron, 1918; Cameron collection). Grenadines: Mustique (Cameron, 1913; Leng and Mutchler, 1917; British Museum). Grenada: (British Museum). Trinidad: St. Augustine (Weber, in M.C.Z. and U.S.N.M.), Tunapuna foothills (Weber, in M.C.Z.). Specimens examined.—I have examined 12 specimens in the British Museum, 4 in the collection of Dr. Cameron, 8 from the Museum of Comparative Zoology, and 1 collected by me in 1935-87. The above description was taken from a specimen borrowed from the British Museum and the one taken by me. Remarks.—This species is distinct from all the others examined by me in having the head actually umbilicately punctured. The specimen labeled type in the Cameron collection must be ac- cepted as the holotype, inasmuch as Dr. Cameron recorded that the type was in his collection. ‘The examples in the British Museum un- doubtedly have the value and usefulness of paratypes, though not entitled to such status under the International Code. My specimen was found in dung. On certain specimens that have been dirty the remnants of the dirt in the sculpture depressions give a distinct impresslon of umbilicate punctures, 374 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 6. ECHIASTER IMPRESSICOLLIS Cameron Echiaster impressicollis CAMERON, 1913b, p. 858.—LeNe and Murcnu er, 1917, p. 198.—ScHEERPELTZ, 1933, p. 1238. Description—Dirty brownish testaceous, abdomen black at apex. Head oval, longer than broad, gradually rounded and contracted be- hind the eyes, without trace of posterior angles; coarsely and rugosely punctured; antennae as long as head, segments 7 to 10 transverse. Pronotum twice as long as broad, widest at middle, gradually nar- rowed in front and behind with all angles obliterated; closely and rugosely sculptured. Zlytra not longer than pronotum, very coarsely but not very closely punctate, with intervals smooth and shining; with coarse pubescence. Abdomen with tergites 3 to 6 very coarsely punc- tured and pubescent with intervals smooth and shining; last two segments very finely punctured and pubescent, shining. Length, 4mm. (From Cameron.) ality.— Grenada 1azar, Type locality —Grenada, Balthazai Zypes.—Unique type in the British Museum. y Records.—The following is the only record known to me: Grenada: (Cameron, 1913; Leng and Mutchler, 1917; British Museum). Specimens examined.—I have seen only the unique type in the British Museum. Remarks —From my study of the type I can record the following to supplement or correct the above description which was rearranged from the original publication: Head exactly one-third longer than wide (length from nuchal constriction to apex of clypeus; width not including eyes); labrum divided into four short triangular lobes which are twice as long as wide at base; sculpture of head not rugose but composed of closely spaced hemispherical tuberculi instead of punctures or depressions; pronotum almost exactly two-thirds as wide as long, sculptured as head; elytra exactly as long as pronotum, one-ninth narrower than long; hind tarsus with first four segments decreasing gradually in length, first longer than the fifth, filiform ; pubescence not at all visible to me. At the time I studied the type I made a note that it was probably not congeneric with the other West Indian species of chiaster, but I am unable now to cite any character that will permit it to be separated. I therefore retain it here until it can be restudied. I find no record of its habits. LVIII. Genus SPHAERONUM Sharp Sphaeronwm SHArp, 1876, p. 225. Sphacrinum Suarp, 1876, p. 36. Sphaerinium Casey, 1905, p. 55 (misspelling). Sphaeronium Casey, 1905, p. 55 (misspelling). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Ste Genotype —sS. pallidum Sharp (designated by Casey, 1905). Diagnosis —Body very narrow and elongate; sides of head strongly grooved behind the eyes; vertex not carinate; eyes small, not prominent; antennae posteriorly flexile, rather strongly geniculate, basal segement not much elongate, first and second not enlarged with rest slender; labrum broadly feebly emarginate, not denticulate; third segment of maxillary palpus subspherical, fourth minute, retractile; gular sutures united throughout; neck very slen- der; prothorax elongate but not greatly prolonged in front; pro- sternum dilated under the front coxae and connate with the hypomera; hypomera not lobed behind the coxae; anterior coxal cavities open behind but closed beneath by the prosternal expansions; anterior coxae large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; first and second abdominal sternites absent; tarsi filiform, five-segmented, fourth segment not lobed beneath. Remarks.—This is the first record of this genus north of the Amazon Valley in Brazil. There can scarcely be any question of the identification because of the very distinct features of this genus. I have examples of the genotype for comparison. One specimen from Cuba is all that I have seen from the West Indies. It seems to be a new species, although I am not able to compare it with all the Amazon species. 1. SEFHAERONUM BERBERUM, new species Description.—Rufocastaneous. ead with sides evenly rounded into base, eyes at about four times their length from base; with a distinct groove from below eye to neck on each side; with rather small but distinctly umbilicate punctures except along front margin, with dis- tinct but not strong scaly ground sculpture; neck one-ninth as wide as head. Pronotum about twice as long as broad; widest at anterior two-fifths, moderately but evenly narrowed behind, more abruptly and nearly emarginately in front; with punctures about half as large as on head, less regular and more obscured by the irregular ground sculpture; narrow midline impunctate, somewhat elevated behind. Elytra a little shorter than pronotum, one-fifth longer than broad; with shallow and somewhat indistinct punctures usually separated by their diameter or more; without distinct ground sculpture. Abdomen without distinct punctures but surface very uneven, not distinctly sculptured. MZale,unknown. Female, sternites not modified. Length, 5 mm. Type locality—Cuba, Soledad near Cienfuegos. Types.—Holotype, female, in the Museum of Comparative Zoology, collected on October 27, 1926, by Dr. P. J. Darlington. 376 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Lecords.—The following is the only record known to me: Cuba: Soledad (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species is very distinct among West Indian sta- phylinids. It differs from S. pallidum in its dense punctation of the head and pronotum. I have received no record of its habits. LIX. Genus PINOPHILUS Gravenhorst Pinophiius GRAVENHORST, 1802, p. 201. Lycidus Laporte, 1834, p. 121. Lyeidius LEACH, in Dejean, 1836, p. 73 (misspelling). Pityophilus BRULLE, 18387, p. 75. Lycidius LACoRDAIRE, 1854, p. 102 (misspelling). Phinopnilus REDTENBACHER, 1874, p. 75 (misspelling). Subgenus Heteroleucus SHarp, 1886, p. 629. Subgenus Pinophilinus HICHELBAUM, 1908, p. 85. Subgenus Metapinophilus GRIDELLI, 1927, pp. 114, 117. Genotypes.—P. latipes Gravenhorst (monobasic). Of Pityophilus, same as Pinophilus (Rules, Article 30, I, f); of Lycidus, L. latipes Gravenhorst (stillborn synonym); of Metapinophilus, Pinophilus reticulatus Eppelsheim=Pinophilus (Metapinophilus) reticulatus Eppelsheim (designated here) ; of Zeteroleucus, H. marginatus Sharp (designated here); of Pinophilinus, Pinophilus fauveli Schubert= Pinophilus (Pinophilinus) fauveli Schubert (designated here). ay) D Ficure 3.—Paramere of genitalia of males of Pinophilus: A, P. aguayot Bierig; B, P. flavipes Erichson; C, P. hispaniolus, new species; D, P. bierigi, new species; E, P. darlingtoni, new species; F, P. schwarzi, new species; G, P. subterraneus, new species; H, P. trinitatis, new species; I, P. danfortht, new species. Diagnosis —Head margined beneath the eyes; antennae inserted at sides of head close to eyes; labrum very short, more or less bilobed ; mandibles with a median inner tooth; fourth segment of maxillary palpus obliquely hatchet-shaped; gular sutures separate; hypomeron prolonged in a lobe partly behind the coxa; sternite expanded under MONOGRAPH OF WEST INDIAN STAPHYLINIDAE oid the coxae and united with the hypomera; front coxae very large, ex- serted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; abdomen strongly margined, first and second sternites ab- sent, third longitudinally carinate at base; tarsi 5-segmented, middle tibia not spinose on outer side. Remarks.—This genus contains some of the largest staphylinids known from the West Indies. It is readily distinguished from all other West Indian genera by the enormously expanded anterior tarsi. Many of the species are very difficult to distinguish by external characters, but the median lobe of the male aedeagus provides satis- factory characters. ‘These may often be observed in undissected speci- mens, but the removal of the aedeagus and the mounting of it sep- arately are very easy and much to be recommended. I have examined 86 examples of this genus and assign them to 13 species, of which 9 are new. One other species described from the West Indies is not available to me. KEY TO WEST INDIAN SPECIES OF PINOPHILUS i eronotum»with Gense vround sculptures 22 2a ee a ee eee 2 Pronotum jshining, without ground sculptures 22232.) ssa oe Eee es 7 2. AGLOUNGESCulpture, of: minutespunctulacs 2: st see ie Sel ee ee 3 Ground:sculpture of minute reticulations 2-2. tos ke 6 8. Head with three distinct sizes of punctures between the eyes______--___ ok Head with only very large and very small punctures between the VCS Mee ees Set Se eee Saab ee ae ea es oe See 8. bierigi 4° mMichthysternitesof, malesbroadly «truncate. 2222s ee 5 Highth sternite of male somewhat produced and then narrowly truncate. 11. aguayoi 5. Punctures along midline of pronotum often separated by nearly their dia- STATES Tyee nee ee ee RE ne ee ee ee eS 9. flavipes Punctures never separated by more than half their diameter. 10. hispaniolus 6. Pronotum impunctate along midline_______-_______-___ 12. insigniventris Pronotum without impunctate median band--_~-_-----_____ 13. schwarzi feetivira, red): length 6:b.mm 232 — = 2 at ee ee 1, vermiformis Bivirasbiack lengths 12 alOwlon mms 2 tt elle ees ot Sy ee ae, 8 8. Pronotum as broad as or broader than long, coarsely punctate___________ 9 Bronosumwonger: chan. broad, finely, punctate... == S222 en ee At 9. Elytral intervals flat, only slightly rugose_____.___-_____-~ 5. subterraneus Hlytral intervals convex, strongly rugose.22 22-22 et Se 10 10. Intervals near basal angles of pronotum with numerous or dense punctulae, 7. jamaicensis Intervals at most with scattered punctulae____-____________ 6. darlingtoni PIPE SIeSHOn Pronobuni, Parallela =. 222 oe 4, trinitatis PronorulMecistinectly broader invfronte sa See ee et ee 12 12. Pronotum very sparsely punctate except near midline___________ 3. cubanus Pronotum moderately densely punctate, a little more sparsely at sides. 2. danforthi 449008—42——_25 278 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 1. PINOPHILUS VERMIFORMIS Cameron Pinophilus vermiformis CAMERON, 1913b, p. 330.—LENe and MutTcHtLerR, 1917, p. 198.—ScHEERPELTZ, 1938, p. 1210. Description —Head and elytra reddish brown; thorax and abdomen pitchy brown, the posterior margins of the segments and apex of abdomen reddish. Head with about five large punctures on each side between and in front of the eyes; without “visible” ground sculpture. Pronotum with close, moderately coarse umbilicate punctures except on smooth midline. //ytra coarsely and closely punctate, more so than thorax. Abdomen pretty closely and moderately coarsely punc- tate, more finely but less closely posteriorly. Length,6.5mm. (From Cameron. ) Type locality —Mustique, Grenadines. Types.—In the British Museum. Records.—The following are the records known to me: Grenadines: Mustique (Cameron, 1913; Leng and Mutchler, 1917; Scheerpeltz, 1933; British Museum). Specimens examined. —I have examined the type briefly. Remarks.—This species differs from danforthi in the large punc- tures of the head, as well as its very small size and reddish elytra. Only the single type is known. I find no record of its habits. 2. PINOPHILUS DANFORTHI, new species Description.—Piceous, tip of abdomen rufous. Head almost with- out the very large punctures between and in front of the eyes, with small punctures and minute punctulae, the former somewhat grouped ; surface shining. Pronotwm not coarsely but rather densely punctate, the intervals smooth and shining, and with occasional punctulae; with a narrow and irregular impunctate midline. /lytra with rather strong punctures separated by about their diameter or less by flat intervals; without ground sculpture. Abdomen with rather coarse and dense punctures strongly excavated behind; with very fine in- distinct transverse strigulae. J/a/e, seventh sternite with a median longitudinal concavity which forms posteriorly a semicircular emar- gination, the corners being slightly prolonged; eighth sternite with a very large rectangular emargination preceded by a narrow smooth area, the base of the emargination prolonged as a triangular or rounded lobe less than one-third as long as the notch is deep, with a group of many short acute setae before the smooth area. Female, sternites unmodified. Length, 15 to 19 mm. Type locality —Puerto Rico, Humacao. Ty pes.—Holotype, male, U.S.N.M. No. 52426, collected by Miguel A. Diaz on November 18, 1930; seven paratypes from Haiti in the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 319 United States National Museum and seven in the Museum of Com- parative Zoology. Records—The following are the records known to me: Hispaniola: Haiti, Kenskoff (Dozier, in U.S.N.M.), swamps north of Dessalines (Darlington, in M.C.Z. and U.S.N.M.), Miragoane (Darlington, in M.C.Z. and U.S.N.M.), Etang Lachaux (Darlington, in M.C.Z. and U.S.N.M.), Ennery (Darlington, in M.C.Z.). Puerto Rico: Humacao (Diaz, in U.S.N.M.). Specimens examined.—lI have seen only the 15 types. Remarks.—This is a very distinct species because of its shining integuments and large size. It is distinguished from all other species by its male characters; from all but favipes and darlingtoni by its size. P. flavipes has the integuments sculptured; darlingtoni has the elytra reduced and appressed. T have no record of the habits of this species. 3. PINOPHILUS CUBANUS, new species Description.—Piceous, tip of abdomen rufous. Head without very large punctures, with rather small punctures sparse and leaving smooth areas, and with sparse minute punctulae; surface shining. Pronotum longer than broad; punctures as on head, separated by at least half their diameters along disk, very sparse at sides, the intervals smooth and shining, and with occasional punctulae; with irregular median stripe impunctate but with some punctulae. LH lytra with rather strong punctures usually separated by less than their diameter by flat intervals; without ground sculpture. Abdomen with rather coarse and dense punctures often excavated behind; surface somewhat alutaceous but without distinct sculpture. J/a7e, unknown. Female, sternites unmodified. Length 12 mm. Type locality—Cuba, Cayamas. Types.—Holotype, female, U.S.N.M. No. 52515, collected by E. A. Schwarz; one paratype, female, in the Museum of Comparative Zoology. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Specimens examined.—lI have seen only the two types. Remarks.—This species differs from danforthi in its smaller size and sparse punctation of the pronotum. It probably also differs in the character of the male, at present unknown. I have received no record of its habits. 4, PINOPHILUS TRINITATIS, new species Description—Piceous, apex of abdomen rufous. Head without very large punctures, with small punctures rather sparse and a few 380 BULLETIN 182, UNITED STATES NATIONAL MUSEUM minute punctulae; surface shining. Pronotwm longer than broad, parallel; punctured and sculptured as head, more densely on disk; with irregular impunctate midline. /ytra with rather strong punc- tures usually separated by nearly their diameter by flat intervals; without distinct ground sculpture. Addomen with rather fine and shallow, sometimes submuricate, excavated punctures rather dense; without distinct ground sculpture. Jfale, eighth sternite with an acute triangular emargination about as deep as wide, with the an- gles broadly rounded, surrounded by a narrow gutter. Female, un- known. Length, 12 mm. Type locality —Trinidad, St .Augustine. Types.—Holotype, male, in the Museum of Comparative Zoology, collected in April 1929 by Dr. P. J. Darlington. Records—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z.). Specimens examined,—t have seen only the unique type. Remarks.—This species is readily distinguished from all others in our fauna by the parallel pronotum. It is the only species so far known from Trinidad. I have received no record of its habits. 5. PINOPHILUS SUBTERRANEUS, new species Description—Rufous to rufopiceous. Head with sparse, coarse punctures, mostly in groups, and with numerous fine punctulae; with- out ground sculpture, surface shining. Pronotwm a little broader than long, arcuately narrowed into base; with moderately coarse punctures generally separated by one-fourth their diameter; with- out smooth midline; without ground sculpture but with an occa- sional punctule. //ytra shorter than pronotum, narrower than ab- domen; with punctures about as on pronotum, the intervals a little more convex; without ground sculpture. Abdomen a little less coarsely but more densely punctate than elytra; without distinct sculpture on the intervals. Jfale, eighth sternite with a shallow rounded emargination about one-fifth as deep as wide. Female, sternites not modified. Length, 16 to 19 mm. Type locality—Cuba, Buenos Aires in the Trinidad Mountains, elevation 2,500 to 8,500 feet. Types.—Holotype, male, and one paratype, female, in Museum of Comparative Zoology; one paratype, female, in the United States National Museum (No. 52516); collected on May 8-14, 1936, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Cuba: Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 381 Specimens examined.—I have seen only the three types. Remarks.—This species has a peculiar subterranean habitus that is distinct from all others except darlingtoni. From that it is dis- tinguished by the flat elytral intervals, as well as by the male characters. I have received no record of its habits. 6. PINOPHILUS DARLINGTONI, new species Description.—Piceous, apex of abdomen rufous. Head with scat- tered coarse punctures, usually in groups, and with rather dense coarse punctulae separated by one to three times their diameter; without ground sculpture, shining. Pronotum a little broader than long, arcuately narrowed into base; with moderately coarse punc- tures generally separated by less than half their diameter; without smooth midline; without ground sculpture, but with an occasional punctule. “lytra shorter than pronotum, narrower than abdomen; punctures not as coarse as on pronotum but denser, the intervals dis- tinctly rugose but without ground sculpture. Abdomen with punc- tures rather fine, very shallow, and widely excavated behind; with strong ground sculpture in the punctures and indistinctly on the intervals. Male, eighth sternite with a triangular emargination half as dcep as wide with all angles rounded. Female, eighth sternite feebly broadly lobed behind. Length, 17 to 18 mm. Type locality—Jamaica, main range of the Blue Mountains, ele- vation 5,000 to 7,388 feet. Types.—Holotype, male, and one paratype, female, in the Museum of Comparative Zoology; one paratype, female (No. 52517), in the United States National Museum; collected on August 17-19, 1934, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I\ have seen only the three types. Remarks.—This is our largest species and one of two so far found in Jamaica. It is distinguished by its subterranean habitus and its rugose elytral intervals. I have received no record of its habits. 7. PINOPHILUS JAMAICENSIS, new species Description—Piceous. Head with scattered large punctures usu- ally in groups, sparsely interspersed with rather coarse punctulae}; without ground sculpture. Pronotum a trifle broader than long, arcu- ately narrowed to rounded basal angles; with very coarse punctures usually separated by half their diameter or less; without smooth mid- 382 BULLETIN 182, UNITED STATES NATIONAL MUSEUM line; with occasional punctulae on the intervals hecoming dense near basal angles. “lytra shorter than pronotum, narrower than abdo- men; punctures less coarse or distinct than on pronotum but denser, the intervals somewhat convex and rugose, and with traces of ground sculpture. Abdomen with dense punctures excavated and submuri- cate, almost indistinguishable; without distinct ground sculpture. Male, unknown. Female, sternite not modified. Length, 1214 mm. Type locality Jamaica, main range of Blue Mountains, elevation 5,000 to 7,388 feet. Types.—Holotype, female, in the Museum of Comparative Zoology, collected on August 17-19, 1934, by Dr. P. J. Darlington. Records —The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This is also a species of subterranean habitus. It dif- fers greatly from darlingtoni in size as well as the distribution of punctulae on the pronotum. I have not hesitated to describe it from a single female because of its distinctness and the scarcity of material from the high mountain regions. I have received no record of its habits. 8. PINOPHILUS BIERIGI, new species Description.—Black, front margin of head and apex of abdomen rufescent. J7Zead with a few very large punctures between and in front of the eyes, and with rather dense punctulae throughout. Pronotum with moderately large and not sparse punctures separated by flat intervals, the latter densely punctulate, the punctures densely sculptured within. /7ytra with strong punctures separated by their diameter or less by moderately convex intervals; surface strongly coriaceous. Abdomen with moderately coarse but very shallow punctures, each densely sculptured within and some feebly sub- muricate. Afale, eighth sternite strongly produced into a rounded lobe. Female, unknown. Length, 11 mm. Type locality—Puerto Rico, Mayagiiez. Types.—Holotype, male, U.S.N.M. No. 52473, collected by A. Suro on September 15, 1930. Records.—The following is the only record known to me: Puerto Rico: Mayagiiez (Suro, in U.S.N.M.). Specimens examined.—I have seen only the unique type. Remarks.—This species may be distinguished by the dense punc- tulae of the head, as outlined in the key. It belongs in the same group as aguayot and flavipes, of which the status is not certain. It MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 383 was kindly presented to the National Museum by Dr. §. T. Danforth, of Mayagiiez. I have no record of its habits. 9. PINOPHILUS FLAVIPES Erichson Pinophiius flavipes Ericuson, 1840, p. 674—Morscuutsky, 1857, p. 516.— SHARP, 1886, p. 621.—BrERNHAUER and ScHUBERT, 1912, p. 193.—LeNG and MUuTCHLER, 1914, p. 404.—Wotocorr, 1924, p. 78; 1936, p. 196. Description —Black, posterior border of abdominal segments rufes- cent. Head with about five very large punctures on each side between and in front of the eyes, and with numerous rather small punctures interspersed with minute punctulae, the latter more sparse in front; without ground sculpture. Pronotum with large and moderately dense punctures, the intervals flat and rather densely minutely punctulate; the large punctures also very densely sculp- tured inside; with a narrow irregular impunctate midline. Elytra with strong punctures separated by their diameter or less by strongly convex intervals; without distinct ground sculpture. Abdomen very coarsely but not sparsely punctate, the. punctures either excavated behind or submuricate; with rather indistinct transverse strigulae. Male, eighth sternite feebly truncate. Female, eighth sternite with a moderately large rounded lobate expansion. Length, 12 to 16 mm. Type locality—Puerto Rico. T ypes.—Kither in the Hope Museum, Oxford, or in the Zoologische Museum, Berlin. Records —The following are the records known to me: Bahamas: Cat Island (Darlington, in M.C.Z. and U.S.N.M.). Cuba: Cayamas (Schwarz, in U.S.N.M.), Baragua (Stahl and Christenson, in U.S.N.M.; Searamuzza and Stahl, in M.C.Z. and U.S.N.M.), Laguna La Canoa (Bierig collection and U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Central Jaronu (Scaramuzza, in M.C.Z.). Caymans: GRAND CaYMAN (Lewis, in Oxford University collection and U.S.N.M.). Hispaniola: DomMiInIcAN RepuBLic, Barahona (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: (Erichson, 1840; Sharp, 1886; Bernhauer and Schubert, 1913 ; Leng and Mutchler, 1914; Wolcott, 1924, 1936). Specimens examined.—I have seen 10 examples in the United States National Museum, 11 from the Museum of Comparative Zoology, and 5 from Oxford University. Remarks.—This species is somewhat variable but may be recognized by the presence of the three distinct sizes of punctures on the head. The related species lack the middle size entirely though having the minute punctulae more variable. Most specimens were taken at light, but I have seen examples labeled “from mud along river.” 384 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 10. PINOPHILUS HISPANICOLUS, new species Description—Black or piceous. Head with scattered large punc- tures usually in groups, interspersed with small punctures and minute punctulae; without ground sculpture. Pronotum with rather large and dense punctures usually separated by less than half their diameter, the intervals flat and rather sparsely minutely punctulate; the large punctures densely sculptured within; without distinctly impunctate midline. /7ytra with punctures as large as on pronotum, dense and separated by convex rugose intervals; without distinct ground sculpture. Addomen with punctures rather fine and dense, shallow and excavated behind, sometimes submuricate, always sculp- tured within; without distinct ground sculpture. fale, eighth sternite broadly truncate. Female, sternites not modified. Length, 11 to 13 mm. Type locality—Dominican Republic, Sanchez. Types.—Holotype, male, in Museum of Comparative Zoology; one paratype, female, in the United States National Museum (No. 52541) ; collected in July 1938 by Dr. P. J. Darlington. Another paratype in the Museum of Comparative Zoology from Etang Lachaux in Haiti. Records.—The following are the records known to me: Hispaniola: Harri, Etang Lachaux (Darlington, in M.C.Z.); DomMINican Re- PUBLIC, Sanchez (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This species may be distinguished from flavipes by its dense punctation and by the very different aedeagus of the male. From aguayoi it is distinguished by the aedeagus as well as a slight difference in the shape of the eighth sternite. I have received no record of its habits. 11. PINOPHILUS AGUAYOI Bierig Pinophilus aguayoi Brrerta, 1935b, p. 29, pl. 3, fig. 1. Description—Black. Head with a few large punctures _be- tween and before the eyes interspersed with small punctures and punctulae; without ground sculpture. Pronotum with very large and rather dense punctures usually separated by half their diameter or less and completely filled with dense ground sculpture; the inter- vals rather densely irregularly punctulate. lytra with rather coarse punctures separated by their diameter or less by strongly convex intervals, somewhat obscured; without distinct ground sculpture. Abdomen moderately coarsely but not sparsely punctate, some- times submuricately; with indistinct transverse strigulae. Made, eighth sternite somewhat produced and narrowly truncate. Female, sternites not modified. Length, 814 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 385 Type locality Cuba. Types—In collection of Alexander Bierig. One “cotype” is in the United States National Museum (No. 52716). Pecords.—The following are the records known to me: Cuba: Habana (Bierig, 1985; U.S.N.M.); Cumanayagua (Bierig, 1935), El Rangel (Bierig, 1935), Aspiro (Bierig, 1935), Soledad (Darlington, in M.C.Z. and U.S.N.M.), Aguadores, in Oriente (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined.—i have seen only the paratype in the United States National Museum and five examples collected by Dr. Dar- lington. Remarks —This species is distinguished by its sparse punctulae on the head and the very dense sculpture of the pronotum, but more particularly by the aedeagus of the male. It is one of a group of species occurring throughout the New World which can be dis- tinguished with certainty only by the male characters. This group includes the West Indian béerigi, aguayot, flavipes, and hispaniolus, the North American latipes and parcus, the Central American albipes, erythropterus, sequnctus, panamensis, guatemalenus, iners, and linea- ticollis, and the South American swu/ffusus. I find no record of the habits of this species. 12. PINOPHILUS INSIGNIVENTRIS Bernhauer Pinophilus insigniventris BERNHAUER, 1918, p. 65.—ScHEERPELTZ, 1933, p. 1208. Description—Black. Head extremely finely and moderately densely punctate and with a number of coarse large punctures; with fine and dense ground sculpture. Pronotum rather strongly and densely punc- tate except along midline; very distinctly and extremely densely re- ticulately sculptured. lytra with deep dense punctures, the inter- vals more or less united into ridges, but shining. Abdomen finely and somewhat densely submuricately punctate; with ground sculpture. Sewves not distinguished. Length,8 mm. (From Bernhauer.) Type locality—Guadeloupe. Types.—in the collection of Dr. Max Bernhauer. Records.—The following is the only record known to me: Guadeloupe: (Bernhauer, 1918). Specimens examined. —ti have not seen examples of this species. Remarks.—This species is very similar to sehwarzi from Cuba. It differs as described principally in having an impunctate midline on the pronotum but will probably be distinguished by the male aedeagus when there is ample material available. I find no record of its habits. 386 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 13. PINOPHILUS SCHWARZI, new species Description.—Black or piceous. Head with large, scattered, some- what umbilicate punctures and a few minute punctulae; with dense reticulate ground sculpture except in front. Pronotwm with punctures a little smaller than on head, somewhat obscured by very dense reticu- late ground sculpture but often separated by their diameter; without distinct midline. Z/ytra with deep and strong punctures as large as those on head separated by their diameter or less by feebly convex intervals; without distinct ground sculpture. Abdomen with mod- erately coarse excavated or submuricate punctures; ground sculpture indistinct except in punctures. Jfale, eighth sternite somewhat trian- gularly produced. Female, eighth sternite rounded. Length, 9 to 11 mm. Type locality—Cuba, Cayamas, 15 miles west-northwest of Cien- fuegos, province of Santa Clara. Types.—Holotype, female, U.S.N.M. No. 52474, collected by E. A. Schwarz; one paratype from Soledad, Cuba, in the Museum of Com- parative Zoology ; 16 paratypes from the Cayman Islands in the Oxford University collection and 7 in the United States National Musevm, collected by Dr. C. B. Lewis. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Caymans: GRAND CayMAN (Lewis, in Oxford University and U.S.N.M.). Specimens examined.—I have seen only the 25 types. Remarks.—This species is similar only to insigniventris which is not available for comparisons. If differs from all other West Indian Pinophilus in the dense reticulate ground sculpture. I have no record of its habits. LX. Genus ARAEOCERUS Nordmann Aracocerus NORDMANN, 1836, p. 7. Aracocerus NORDMANN, 1886, p. 157 (misspelling). Subgenus Scotocerus BERNHAUER, 1918, p. 67. Araeocrus FALL, 1932, p. 58 (misspelling). Areocerus FALL, 1932, p. 58 (misspelling). Genotypes —A. niger Nordmann (monobasic). Of Scotocerus, Araeocerus (Scotocerus) curtipennis Bernhauer (designated here). Diagnosis —Head margined beneath the eyes; antennae inserted at sides of head close to eyes; labrum very short, almost bilobed; man- dibles very slender, without teeth; fourth segment of maxillary palpus large and obliquely hatchet-shaped; gular sutures separate but ap- proximate from posterior fifth to middle; hypomera prolonged in a lobe partly behind the coxae; prosternum expanded laterally beneath coxae and united with hypomera; front coxae very large, exserted; MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 387 posterior coxae approximate, “conical”; abdomen margined at sides, first and second sternites absent, third longitudinally carinate at base ; tarsi 5-segmented., Remarks.—This diagnosis was taken from the specimen of parvi- pennis in my own collection. Other species were not available for comparison. The genus is distinguished from Pinophilus only by the absence of the large median tooth on the mandible. This seems to be scarcely sufficient justification for the separation, but 1t will be fol- lowed here until further studies are made. Only one species has so far been found in the West Indies. 1. ARAEOCERUS PARVIPENNIS Bierig Aracocerus parvipennis Brzric, 1935b, p. 30, pl. 3, figs. 2, 3. Deseription.—Black to rufocastaneous. Head quadrate; eyes sepa- rated from base by scarcely their Jength; labrum transverse, with a deep rounded notch at middle; with a few scattered very large and deep punctures and with very fine and indistinct ground sculpture. Pronotum with sides feebly curved, somewhat narrowed posteriorly, with an indistinct smooth median line; punctures relatively fine, irreg- ular, the intervals usually about half their diameter; with ground sculpture as on head. Elytra shorter than pronotum, narrower than abdomen; with punctures a little larger than on pronotum and sepa- rated usually by less than half their diameter; without ground sculp- ture. Abdomen with moderately coarse and shallow excavated punc- tures, the intervals with vague ground sculpture. J/a/e, seventh ster- nite broadly and feebly emarginate; eighth with a rounded emargina- tion over four times as wide as deep. Female, sternites not modified. Length, 9 to 12 mm. Type locality—Cuba. Types.—In the collection of Alexander Bierig. (One “cotype” 1s in the collection of the writer.) Records.—The following are the records known to me: Cuba: Playa de Marianao (Bierig, 1935), El Rangel (Bierig, 1935), La Lisa (Bierig, 1935), Sierra Bonilla (Bierig, 1935), Aspiro (Bierig, in Blackwelder collection), Rangel Mountains in Pinar del Rio (Darlington, in M.C.Z.), Gran Piedra Range in Oriente (Darlington, in M.C.Z.), Loma del Gato in Oriente (Darlington, in U.S.N.M.), Buenos Aires, Trinidad Mountains (Dar- lington, in U.S.N.M). Specimens examined—I have seen only one example in my own col- lection, received some years ago from Sr. Bierig, and four collected by Dr. Darlington. Remarks.—This species is very distinct from the other West Indian Pinophilini. It has a distinct subterranean habitus and an unusually distinct ground sculpture. It was originally captured by sifting fallen leaves. 388 BULLETIN 182, UNITED STATES NATIONAL MUSEUM LXI. Genus PALAMINUS Erichson Palaminus Ertcyson, 1840, p. 681. Genotype.—P. pilosus Erichson (designated by Duponchel, 1841). Diagnosis —Antennae inserted at sides of head close to eyes; la- brum very short; fourth segment of maxillary palpus obliquely hatchet-shaped; gular sutures separate; hypomera feebly lobed be- hind the coxae; prosternum expanded beneath the coxae and broadly united with the hypomera; front coxae large, exserted; middle coxal cavities confluent; posterior coxae contiguous, “conical”; abdomen not margined, first and second sternites absent, third longitudinally carinate at base; middle tibia not spinose on outer side; tarsi 5-seg- mented, Remarks.—This extensive genus is well represented in the West Indies. Although many species have been described, there has been almost no attempt to determine the extent and limits of specific varia- tion. This is true for all the other parts of the Western Hemisphere as well. Although the present collection contains about 100 ex- amples, and J have seen about 80 in other collections, I feel unable to determine the variation in characters which have been used as of primary importance. Since only five species have been definitely recognized by comparison with types, it seems premature to attempt to give a key to all the known species. Therefore, in the present study I am presenting a summary of the records and data so far pub- lished with additional records of which there is reasonable certainty, and I am omitting keys, descriptions, and new species. 1. PALAMINUS ANACORETA Bierig Palaminus anacoreta Breria, 1935b, p. 34, pl. 3, fig. 7. Type locality—Cuba, Portugalete, province of Habana. Types.—Unique type in collection of Alexander Bierig. fecords—The following is the only record known to me: Cuba: Portugalete (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 2. PALAMINUS APTERUS Bernhauer Palaminus apterus BERNHAUER, 1918, p. 82—NoTMAN, 1929, pp. 9, 17.—BuErIa@, 1935b, p. 88.—SCHEERPELTZ, 1933, p. 1213. Type locality.— Guadeloupe. Types.—In collection of Dr. Max Bernhauer. Records.—The following are the records known to me: Guadeloupe: (Bernhauer, 1918 ; Notman, 1929, A.M.N.H.). Specimens examined.—I have seen no examples of this species. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 389 3. PALAMINUS BERMUDEZI Bierig Palaminus bermudezi Brrria, 1935b, p. 35, pl. 4, figs. 2, 3. Type locality —Cuba, Sierra del Rosario, vicinity of El Rangel. Types.—In collection of Alexander Bierig. Records—The following is the only record known to me: Cuba: Sierra del Rosario (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 4. PALAMINUS BIFIBUS Notman Palaminus bifidus Norman, 1929, pp. 9, 14.—ScHEERPELTZ, 1933, p. 1213.—Brerte, 1935b, p. 86.—Wotcort, 1936, p. 196. Type locality—Puerto Rico, KE! Yunque. Types.—Holotype, male, allotype and four other paratypes, in the American Museum of Natural History. Records—The following are the records known to me: Puerto Rico: El Yungue (Notman, 1929; Wolcott, 1986), Aibonito (Notman, 1929; Wolcott, 1936), Adjuntas (Notman, 1929; Woicott, 1936). Specimens exwamined—I have examined the six types and two other examples in the American Museum of Natural History. 5. PALAMINUS BRUCHIANUS Bierig Palaminus bruchianus BreRia, 1935b, pp. 36, 387, 38, pl. 3, figs. 8, 9. Type locality.—Cuba, Playa de Marianao. Types.—In collection of Alexander Bierig. Records.—The following is the only record known to me: Cuba: Playa de Marianao (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 6. PALAMINUS CORIACEUS Cameron Palaminus coriaceus CAMERON, 1913b, p. 331.—Lene and MuTCHLER, 1917, p. 198.— Norman, 1929, pp. 9, 17.—ScHeEERPELTz, 1933, p. 1214.—Bieria, 1935b, p. 36. Type locality —Jamaica. Types.—In the collection of Dr. Cameron. (This species was originally recorded from “Jamaica, Haiti.” I found three speci- mens in the Cameron collection, one from Jamaica and one from Haiti, both being labeled type. I hereby designate the Jamaican type to be lectotype of this species.) Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in United States National Museum). Jamaica: (Cameron, 1913; Leng and Mutchler, 1914; Notman, 1929; Cameron collection). Hispaniola: Hartt (Cameron, 1913; Leng and Mutchler, 1917; Cameron collec- tion). 390 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—lI have seen the type and two other examples in Dr. Cameron’s collection, and two in the United States National Museum. 7. PALAMINUS DUBIUS Notman Palaminus dubius Norman, 1929, pp. 11, 16.—ScuHrerPettz, 1933, p. 1214. Type locality —Guadeloupe, Gourbeyre. Types.—Unique holotype in American Museum of Natural History. Records.—The following is the only record known to me: Guadeloupe: Gourbeyre (Notman, 1929). Specimens examined.—I have seen only the unique type. 8. PALAMINUS GRANDICOLLIS Notman Palaminus grandicollis NorMAN, 1929, pp. 9, 16.—ScHIERPELTZ, 1933, p. 1215.— WotcoTt, 1936, p. 197. Type locality—Puerto Rico, Aibonito. Types.—Holotype (male), allotype, and one other paratype, in American Museum of Natural History. Records.—The following are the records known to me: Puerto Rico: Aibonito (Notman, 1929; Wolcott, 1936), Adjuntas (Notman, 1929; Wolcott, 1936). Specimens examined.—I have seen the three types and one other specimen in the American Museum of Natural History. 9. PALAMINUS HYLAEUS Bierig Palaminus hylaeus Bierta, 1935b, pp. 32, 33, 34, pl. 3, fig. 10. Type locality.—Cuba. Types—In collection of Alexander Bierig. (A cotype is in the United States National Museum, No. 52717.) Records.—The following are the records known to me: Cuba: Playa de Marianao (Bierig, 1935), Sierra Bonilla (Bierig, 1935), Sierra del Rosario, Cayajabos (Bierig, 1935), Aspiro (Bierig, 1935), Corralillo (Santa Clara) (Bierig, 1935), Habana (Bierig, in United States National Museum). Specimens examined. —I have seen only one example in the United States National Museum. 10. PALAMINUS INSULARIS Cameron Palaminus insularis CAMERON, 1913b, pp. 331, 332.—LENG and MuTcH Ler, 1917, p. 198.—NorMaN, 1929, p. 12, 17.—ScHEERPELTz, 1933, p. 1215.—Wo corr, 1936, p. 197. Type locality —Jamaica, Hope Gardens district. Types.—Unique holotype in collection of Dr. Cameron. Records.—The following are the records known to me: Jamaica: (Leng and Mutchler, 1917; Notman, 1929), Hope Gardens (Cameron, 1913; Cameron collection). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 391 Puerto Rico: Aibonito (Notman, 1929; Wolcott, 1936). Guadeloupe: Gourbeyre (Notman, 1929). Specimens examined.—I have seen only the unique type. 11. PALAMINUS LENGE Notman Palaminus lengi NoTMAN, 1929, pp. 8, 14—ScHEERPELTZ, 1933, p. 1215.—Wotcort, 1936, p. 196. Type locality—Puerto Rico, Adjuntas. Types——Holotype, female, in the American Museum of Natural History. Records.—The following are the records known to me: Puerto Rico: Adjuntas (Notman, 1929; Wolcott, 1986; A.M.N.H.; Oakley, in U.S.N.M.), Mayagtiez (Dozier, in U.S.N.M.). Specimens examined.—I have seen the six types in the American Museum of Natural History and two examples in the United States National Museum. 12. PALAMINUS PARVIPENNIS Notman Palaminus parvipennis NotMAn, 1929, pp. 9, 14.—ScHEERPELTZ, 1933, p. 1215.— Wotcorrt, 1936, p. 196. Type locality.—Puerto Rico, El Yunque. Types——Holotype (male), allotype and four other paratypes in the American Museum of Natural History. Records.—The following are the records known to me: Puerto Rico: El Yunque (Notman, 1929; Wolcott, 1936), Loiza (Notman, 1929; Wotcort, 1936), Villalba (Blackwelder station 64), Mayagiiez (Dozier, in U.S.N.M.). Specimens examined.—I have seen the six types in the American Museum of Natural History, two examples in the United States Na- tional Museum, and one collected by me in 1935-37. 13. PALAMINUS PERALUTACEUS Bierig Palaninus peralutaceus BiERiG, 1935b, p. 37, pl. 3, figs. 11, 12, pl. 4, figs. 1. Type locality—Cuba, Sierra del Rosario, vicinity of El Rangel. Types.—tIn the collection of Alexander Bierig. Records.—The following is the only record known to me: Cuba: Sierra del Rosario (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 14 PALAMINUS PILUM Bierig Palaminus pilum BinriGc, 1935b, pp. 38, 34, 36, pl. 3, figs. 4-6. Type locality.—Cuba. Types.—tIn the collection of Alexander Bierig. 392 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following are the records known to me: Cuba: Portugalete (Bierig, 1985), Sierra de Anafe (Bierig, 1935), Sierra del Rosario, Aspiro (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 15. PALAMINUS PROCERUS Notman Palaminus procerus NoTMAN, 1929, pp. 18, 16.—ScHEERPELTZ, 1933, p. 1216.—Wot- corT, 1936, p. 197. Type locality——Puerto Rico, Aibonito. Types.—Holotype and one paratype, males in the American Museum of Natural History. Records.—The following are the records known to me: Puerto Rico: Aibonito (Notman, 1929; Wolcott, 1936), Maricao (Morrison, in U.S.N.M.). Specimens examined.—I have seen only the two fragmentary types in the American Museum of Natural History and one example in the United States National Museum. 16. PALAMINUS PUSILLUS Notman Palaminus pusillus NoTMAN, 1929, pp. 9, 15.—ScHEERPELTz, 1933, p. 1216.—Wot- coTT, 1986, p. 197. Palaminus pusilus Bierta, 1935, p. 36. Type locality —Puerto Rico, El Yunque. Types——Unique holotype in the American Museum of Natural History. Records.—The following is the only record known to me: Puerto Rico: El Yunque (Notman, 1929; Wolcott, 19386). Specimens examined.—lI have seen only the unique type. 17. PALAMINUS ROSARIENSIS Bierig Palaminus rosariensis Bierta, 1935b, p. 88, pl. 4, fig. 4. Type locality—Cuba. Types.—In the collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: Cayajabos (Bierig, 1935), Rio Taco-Taco (Bierig, 1935). Specimens examined.—I have seen no examples of this species. 18. PALAMINUS SCITULUS Notman Palaminus scitulus NoTMAN, 1929, pp. 9, 15.—ScHEERPELTz, 1933, p. 1216.—BreErIe, 1935, p. 36.—WoLcotTT, 1936, p. 197. Type locality—Puerto Rico, Aibonito. Types.—Holotype, male, and allotype in the American Museum of Natural History. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 393 Records.—The following is the only record known to me: Puerto Rico: Aibonito (Notman, 1929; Wolcott 1936). Specimens exanined.—I have seen only the two types. 19. PALAMINUS VARIABILIS Erichson Palaminus variabilis ERicHson, 1840, p. 683, pl. 1, fig. 8—LacorpatrgE, 1854, p. 104.—SHArpP, 1876, p. 340.—FLEUTIAUX and SALL&, 1889, p. 381.—BERNHAUER and SCHUBERT, 1912, p. 201.—CameEron, 1918b, p. 331.—LeNe@ and MurcHLEr, 1914, p. 404.—Gowpey, 1926, p. 12—Norman, 1929, p. 12.—ScHEERPELYz, 1933, DP Qi: Type locality—‘In Columbia et in Americae meridionalis ins. Puerto-Rico et St. Thomae.” Types—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. fecords.—The following are the records known to me: Jamaica: (Gowdey, 1926). Puerto Rico: (Erichson, 1840). St. Thomas: (Hrichson, 1840). Guadeloupe: (Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). St. Vincent: (British Museum; Cameron collection). Grenada: (British Museum; Cameron collection). Trinidad: San Fernando Hill (Morrison, in U.8.N.M.), Palo Seco (Morrison, in U.S.N.M.), D’Abadie (Morrison, in U.S.N.M.), Port of Spain (Morrison, in U.S.N.M.), Montserrat (Busck, in U.S.N.M.). South America: CoLomBIA (Erichson, 1840; Lacordaire, 1854), Venezueta (Bern- hauer and Schubert, 1912). Central America: (Bernhauer and Schubert, 1912). Specimens examined.—I have seen 33 examples in the British Museum, 5 in Dr. Cameron’s collection, and 21 in the United States National Museum. Subfamily STAPHYLININAE This large subfamily contains most of the large species of Staphy- linidae. It is readily distinguished by the characters used in the key from all except the Paederinae, and from these it is known by characters discussed in the introductory remarks to that group. Four rather distinct groups of genera occur in this subfamily. In recent works they have usually been treated as tribes, viz: Xantholinini, Staphylinini, Xanthopygini, and Quediini. As pointed out previously (Blackwelder, 1940) these four are not of equal rank, since the Xantholinini differ more from all the others than they do among themselves. Relationships are therefore better shown by using two tribes, the Xantholinini and the Staphylinini, and then dividing the latter into three subtribes, the Staphylinina, 4490084226 394 BULLETIN 182, UNITED STATES NATIONAL MUSEUM the Kanthopygina, and the Quediina. For convenience the genera of the three last subtribes are arranged in a single key to the tribe. KEY TO TRIBES OF STAPHYLININAE 1. Bases of antennae approximate; prosternum with a pair of sclerites in front Of the sternite: sass a ae ee eee XANTHOLININI Bases of antennae distant; prosternum without anterior sclerites. STAPHYLININI Certain genera that have been placed in the Xantholinini are herein transferred to the Staphylinini. It is possible that a new subtribe will have to be erected for them there, but they certainly have nothing to do with the Xantholinini. These are Diochus and Holisus (both of them having been previously transferred by certain writers). To these I now add Cephaloxynwm, proposed by Bern- hauer in the Piestini but surely a member of this group closely related to Holisus. Both of the characters used above will distin- guish these and ail other Staphylinini from all Xantholinini, so far -as known to me. KEY TO WEST INDIAN GENERA OF STAPHYLININI 1. Lateral marcin-of pronotun doubles) 222 222 ee ee ee iD Lateral’margin -of pronotum’ simple_22-22 2! 2.22 fee eee ss ee eee 13 -2. Fourth segment of maxillary palpus much shorter and more slender than? thirds. 22-3 ha el ee ee eA SS se A eee 3 Fourth segment of maxillary palpus nearly equal to or longer than thirds.) 222 22 een nn bee eee ko soe SR Sees Be eee 5 3. Labrum large, prominent at middless222 222 es LXXI. Diochus MAabrum PDO De Ce ees a a Se ee ee a ee ee 4 4. Side of pronotum with two marginal lines not uniting anteriorly. LXXII. Cephaloxynum Side of pronotum with three marginal lines uniting before anterior angle, LXXIII. Holisus 5h Mareinal lines of pronotum Separate: — =) *e se es ee ee 6 Marginal lines of pronotum united near apex_______-_____-_--_--_--._- 8 6°\Head: margined beneath the eyesi 244-24. es eet ee ee ee 7 Head not margined beneath the eyes_-___________---- LXVIII. Creophilus 7. Upper lateral line of pronotum extended around front angles and then united TOS IO WET Sees 22a es sot eee ee ee LXX. Philothalpus Upper lateral line of pronotum not extended around front angles. LXIX. Xanthopygus 8. Posterior femora with two rows of spines beneath; gular sutures united in firontseaht.~ _ Peers ree ee eR ea LXIV. Belonuchus Posterior femora at most with one row of spines; gular sutures generally unitedionly, basally. =e ees hs oa ee ee ee ee 9 9. Side piece of pronotum inflexed (or head margined beneath eyes) —---—~ 10 Side piece of pronotum not inflexed; head not at all margined beneath Cy.es is kt Shr RE USO, GUAR Se LXII. Paederomimus 10. Head and pronotum with dense umbilicate punctures__ LX VII. Staphylinus Head and pronotum not densely umbilicately punctate_______----_--__- ey MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 395 11. Fourth segment of maxillary palpus large, fusiform ____-.______---_---- cn, Fourth segment of maxillary palpus large near base but becoming very slender DL Call ly eee ere ee ee ee Be ee eS LXVI. Hrichsonius 12. Pronotum with impunctate median area and dense punctures at sides. LXV. Cafius Pronotum virtually impunctate except for distinct series of punctures. LXIII. Philonthus 13..Antennae stronsly-geniculates=—— = aaa ea eases LXXV. Acylophorus ATHLCNNAC TN Ole Pen CUlALC= So 225 eh Se 2 ee a eee 14 SNA PLA SM AA om oe i ee rs eee Sb Se a LXXVI. Atanygnathus EASIEST Cy ym a re en er a ea Se LXXIV. Heterothops LXII. Genus PAEDEROMIMUS Sharp Paederomimus SHARP, 1885, p. 439. Genotype.—P. difformiceps Sharp (at least implied by Sharp, and, if necessary, fixed here). Diagnosis—Head without dense umbilicate punctures, not margined beneath the eyes; antennae widely separated at front of head, outer segments densely pubescent from the fourth joint; labrum bilobed; palpi elongate and slender, fourth segment of maxillary longer than third, labial similar to maxillary; gular sutures united behind middle; jateral margin of pronotum double, lines united near the coxae; an- terior coxal cavities entirely open behind; front coxae large, exserted ; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent; intersegmental membranes of abdomen with pattern of irregular rounded sclerotized areas arranged in longitudinal bands; femora unarmed. Remarks.—This genus was established by Sharp for species formerly of the genus PAilonthus but having the prosternum transversely di- vided into two regions of different texture and having the side pieces of the pronotum not inflexed. It is a large genus confined to tropical or subtropical America. I have seen 33 examples of this genus from the West Indies. I assign these to four species, of which one appears to be new. KEY TO WEST INDIAN SPECIES OF PAEDEROMIMUS 1. Abdomen rufotestaceous, segments six to seven piceous____-~- 2. cribricollis Abdomen at most with eighth segment testaceous________________________ 2 2. Eighth tergite and sometimes posterior border of ninth rufotestaceous. 3. interjectus Abdomensentirel ya piceOussOns Dla Ck == = a ee ee a ee Se 3 See VOLSAl pPronotal SCLies Of Six. PUNCLUECS=-— == =—- aoe == ao 4. lustralis Dorsal pronotal series of four punctures________--______________ 1, tetricus 1. PAEDEROMIMUS TETRICUS, new species Description —Rufopiceous. Head quadrate with angles rounded; eyes separated from base by one-third more than their length; in front 395 BULLETIN 182, UNITED STATES NATIONAL MUSEUM with a feeble median longitudinal impression nearly half as long as head; with large and somewhat impressed punctures scattered at sides; with very fine strigulose ground sculpture. Pronotum one-eighth longer than wide, widest at anterior fourth with anterior angles broadly rounded, feebly emarginately narrowed to rounded basal angles, base sinuate ; dorsal series of four punctures, the first and fourth widely separated; and with about eight smaller lateral punctures; with ground sculpture as on head. “lytra with fine and dense punc- tures not very distinct but forming a coarsely irregular surface; with- out ground sculpture. Abdomen with fine but not dense submuricate punctures; without distinct ground sculpture. Mfale, unknown. Fe- male, eighth sternite rounded. Length, 414 mm. Type locality —Trinidad, Port of Spain. Types.—Holotype, female, U. 8. N. M. No. 52466, collected on “11.6” by August Busck. Records.—The following is the only record known to me: Trinidad: Port of Spain (Busck, in U.S.N.M.). Specimens examined.—I have seen only the unique type. Remarks.—This species differs from lustra/is in having only four punctures in the pronotal series, and from the other West Indian species in its piceous color and the shape of the pronotum. I have no record of its habits. 2. PAEDEROMIMUS CRIBRICOLLIS (Erichson) Philonthus cribricollis Ertcuson, 1840, p. 487.—SHarp, 1885, p. 448. Paederomimus cribricollis (Brichson) SHARP, 1885, pp. 448, 450.—BrRNHAUER and ScHUBERT, 1914, p. 368. Paederomimus trinidadensis BERNHAUER, 1915¢, p. 296.—SCHEERPELTZ, 1933, p. 1376. Description—Head piceous, pronotum rufopiceous, elytra piceous with humeral third rufotestaceous, abdomen rufotestaceous with sixth and seventh segments picescent. ead suborbicular, eyes at less than their length from base; in front with a longitudinal impression nearly half as long as head; with large impressed punctures scattered at sides ; surface smooth but with vague trace of strigulose ground sculpture. Pronotum nearly one-half longer than wide, widest in anterior third, emarginately narrowed to rounded posterior angles; dorsal series of four to six very large impressed punctures (five is apparently normal), and with about seven lateral punctures; surface smooth but with vague strigulae as on head. Zlytra with large punctures separated by con- vex intervals of less than their diameter; without ground sculpture. Abdomen very sparsely and finely submuricately punctate; without trace of ground sculpture. Male, unknown. Female, eighth sternite rounded, Length, 7 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 397 Type locality — “Columbia ..., Mexico...” Of trinidadensis, Caparo, Trinidad. Types—tIn either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of trinidadensis, presumably in coliection of Dr, Bernhauer. Records.—The following are the records known to me: Grenada: (British Museum). Trinidad: Caparo (Bernhauer, 1915, as trinidadensis; Scheerpeltz, 1933, as same; British Museum), Manzanilla (Blackwelder station 103A), South America: Cotompra (Hrichson, 1840; Sharp, 1885; Bernhauer and Schu- bert, 1914), VenrzurLa (Bernhauer and Schubert, 1914). Central America: Mexico (Hrichson, 1840; Sharp, 1885, with doubt; Bernhauer and Schubert, 1914). Specimens examined.—I have seen 11 examples from the West Indies in the British Museum and 2 collected by me on December 18, 19385. Remarks.—The examples in the British Museum from Grenada were labeled ertbricollis and the one from Trinidad trinidadensis. I am unable to find any difference between these or between the original descriptions, Brazilian examples of cribricollis also seem to be the same. This species is readily distinguished from the other West Indian species by its coloring. My examples were found in rotting cocoa pods. 8. PAEDEROMIMUS INTERJECTUS Bernhauer Paederomimus interjectus BERNHAUER, 1908b, p. 8333.—BERNHAUER and SCHUBERT, 1914, p. 368.—LeENG and Murcutrr, 1914, p. 405. Paederomimus smithi CAMERON, 1922, p. 121—ScHEERPELTz, 1933. p. 1876. Description.—Piceous, elytral humerus and apex of abdominal seg- ments rufescent, eighth segment rufotestaceous. Head suborbicular; eyes at one-third more than their length from base; in front with a median longitudinal impression, scarcely half as long as head; with very large impressed punctures scattered at sides; without distinct ground sculpture. Pronotwm one-sixth longer than wide, widest at anterior angles, evenly narrowed to completely rounded base; dorsal series of six punctures, with a lateral series of five and several others; without distinct ground sculpture. Zlytra with large punctures sepa- rated by convex intervals of about their diameter; without ground sculpture. Abdomen with sparse and moderately fine excavated or submuricate punctures; without ground sculpture. fale, unknown. Female, eighth sternite rounded. Length, 5 to 5144 mm. Type locality —St. Vincent, Windward side. Of smithi, St. Vin- cent, Windward side. Types—Presumably in collection of Dr. M. Bernhauer. Of srithi, in the British Museum. 398 BULLETIN 182, UNITED STATES NATIONAL MUSEUM fecords.—The following are the records known to me: St. Vincent: (Bernhauer, 1908; Leng and Mutchler, 1914; Cameron, 1922, as smithi; British Museum, as smithi; U.S.N.M.). Specimens examined.—I have seen 12 examples in the British Mu- seum and 7 in the United States National Museum. Remarks.—These two species were undoubtedly described from ma- terial from the same lot collected by H. H. Smith. The examples in the United States National Museum are from the original series of smithi, received in exchange from the British Museum. I find no record of the habits of this species. 4. PAEDEROMIMUS LUSTRALIS (Erichson) Philonthus lustralis ERIcHSON, 1840, p. 489.—Suarp, 1885, p. 438. Paederomimus lustvalis (Hrichson) SHARP, 1885, p. 488.—BERNHAUER and ScHU- BERT, 1914, p. 8368.—LENeé and Mutcnter, 1914, p. 405.—Wotcort, 1924, p. 78; 1936, p. 197. Paederomimus insularis BERNHAUER, 1908, p. 3383.—BERNHAUER and ScHUBERT, 1914, p. 368.-—Lene and MUTCHLER, 1914, p. 405. Description.—Black, more or less aeneous, elytra fuscoaeneous. Head subovate; with large impressed punctures scattered at sides. Pronotum one-half longer than wide; sides straight, parallel; base and posterior angles rounded; dorsal series of six punctures, and with five lateral punctures. /ytra densely and profoundly punctate. Abdomen densely and finely punctate. Male, eighth sternite pro- foundly bisinuate. Female, eighth sternite rounded. Length, 5 mm. (From Erichson.) Type locality—Puerto Rico. Of insularis, St. Thomas. Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of tmsularis, presumably in the collection of Dr. Max Bernhauer. Records.—The following are the records known to me: Puerto Rico: (Hrichson, 1840; Leng and Mutchler, 1914, as both; Wolcott, 1924, 1936). St. Thomas: (Bernhauer, 1908, as insularis; Bernhauer and Schubert, 1914). Specimens examined.—I have seen no examples of this species. Remarks.—Although the description of insularis is very brief and is almost entirely a comparison with interjectus, I believe it is possible to be reasonably certain that this is the species actually occurring in St. Thomas. It is distinguished by its black color and the six punc- tures in the pronotal series. In all but the latter character it is close to tetricus. I find no record of its habits, LXIII. Genus PHILONTHUS Stephens Philonthus STEPHENS, 1829, p. 279. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 399 Subgenus Gabrius STEPHENS, 1829, p. 279. Subgenus Bisnius STEPHENS, 1829, p. 283. Gefyrobius THomson, 1860, p. 166. Gefyribus JOHANSEN, 1914, p. 888 (misspelling). Cheilocolpus SotiEr, 1849, p. 320. Subgenus Rabigus MULSANT and Rey, 1875, p. 523. Subgenus Hecoptolonthus BERNHAUER, 1912, p. 206. Subgenus Pseudophiionthus BERNHAUER, 1915a, p. 302. Subgenus Onychophilonthus NERESHEIMER and WAGNER, 1924, p. 156. Subgenus Nephronthus BERNHAUER, 1982, p. 147. Nephrontus SCHEERPELTZ, 1984, p. 1782 (misspelling). Genoty pes.—Staphylinus laminatus Creutz=Philonthus laminatus (Creutz) (designated by Lacordaire, 1854). Of Eccoptolonthus, Philonthus (Eccoptolonthus) conradti Bernhauer (monobasic) ; of Cheilocolpus, Staphylinus (Cheilocolpus) pyrostoma Solier (desig- nated here); of Gabrius, G. suaveolens Stephens (designated by Lacordaire, 1854); of Gefyrobius, Staphylinus nitidulus Graven- horst = Gefyrobius nitidulus (Gravenhorst) (monobasic) ; of Bisnis, Staphylinus cephalotes Gravenhorst=Bisnius cephalotes (Graven- horst) (designated by Lacordaire, 1854); of Rabigus, Staphylinus —pullus Nordmann=Rabigus pullus (Nordmann) (designated by Tottenham, 1939); of Mephronthus, Philonthus (Nephronthus) grandis Bernhauer (monobasic) ; of Onychophilonthus, Staphylinus marginatus Fabricius=Philonthus (Onychophilonthus) marginatus (Fabricius) (monobasic) ; of Pseudophilonthus, Philonthus (Pseudo- philonthus) bicoloripennis Bernhauer (monobasic). Diagnosis —Head and pronotum without dense umbilicate punc- tures, sometimes feebly margined beneath the eyes; antennae widely separated at front of head, outer segments pubescent from the fourth joint; labrum bilobed; palpi moderately slender, fourth segment of maxillary longer than third, labial similar to maxillary; gular sutures not united or only posteriorly; pronotum virtually impunctate except for longitudinal series, lateral margin double, the lines not united or only in front; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular,” sometimes with a small expansion over the base of the trochanter ; first and second abdominal sternites absent ; intersegmental membranes of abdomen with a pattern of irregular rounded sclerotized areas arranged in longitudinal rows; femora unarmed. Remarks.—This genus is not distinguished from Belonuchus by characters of great importance or definiteness. The combination of femoral armature and united gular sutures will distinguish the West Indian species of Belonuchus, while in general they can be recognized by a somewhat different appearance. A species was recently described by Bierig (1940) in this genus under the name caribaeus. A drawing of the head and pronotum was given, 400 BULLETIN 182, UNITED STATES NATIONAL MUSEUM and in his remarks Bierig states that the species was identified by Rambousek as Paederomimus sp. Bierig believes. however, that it is a true Philonthus (s. str.) and describes it as such. Although most generic characters are not mentioned in the description, the figure raises doubts as to the generic assignment. It appears possible that the species really belongs in either Paederomimus or Belonuchus, but further examination of the types is necessary for a decision. Since I have no other source of information I list this as a doubtful Philonthus and give the original description in full. The characters given seem to be sufficient to carry it through the key to stlvaticus. Since that species is hkewise not now before me, I am unable to point out distinc- tions, if any do exist. Fourteen species are recognized from the West Indies, but one of these is left undescribed for lack of material. I have examined over 650 examples from the West Indies. KEY TO WEST INDIAN SPECIES OF PHILONTHUS 1, Pronotal series with three punctures._..-.—.2.--.— oe Pronotal series with more than three punctures_____-__________________ 3 2. Mlytra very distinetly and ‘coarsely punctates 2-2 ses Bsa oe as Elytra rather finely and somewhat indistinctly punctate__-__ 1. hepaticus 3: Pronotal series withtfour punctures: £2 oa. fe ee 4 Pronotal series with more than four punctures_______-_____________-_. ateh 4. Head with row of four punctures between eyes____--______-_-________ AO Head without four punctures in row between eyes_______---___-_-__-_- 10 5, Apex of elytra testaceous#2222 lsh eee Se ee 9. flavolimbatus Apex of elytra at most very narrowly testaceous or indistinctly rufescent__ 6 6. Pronotum rufous or at least rufescent, not much narrowed in front_______-_ ac *ronotum black (when mature), distinctly narrowed in front--_____-_--_ 9 7. Head elongate and distinctly broader posteriorly__________- 2. thermarum Head not elongate and not broader posteriorly____________________-____- 8 8. Elytra densely and moderately finely punctate, apical margin and suture testaceOUsH BGs wb oath es sar. cee is torn tt ere beer bee 5. discoideus Elytra not densely and rather coarsely punctate, margins not testaceous. 3. ventralis 9. Punctures of abdomen very fine and dense; elytra unicolorous. 7. longicornis Punctures of abdomen not fine or dense; elytra generally marked with PUfOuUSs oc3 Ferrera Ps Renewal ee est eee § eel eee eae | een 8. varians 10. Punctures of elytra indistinct; abdomen sparsely punctate throughout. 4. audanti Punctures of elytra small but distinct; abdomen more densely punctate POSsterior]y 2220 Be a ee eee eee ees 5a 6. figulus 11. Sides of pronotum parallel or narrowed in front------_ 14. havaniensis Pronotum: narrowed) posteriorly2sc_ sei toe. bitte fie segs i ey 12 12..iMlytra, shining bronze-ereen.=. =. af on Se ree 13 Hlytra rufous or rufopiceous... = 2 ee ee 14 174 species known only from quarantine interceptions keys to this point but is left undescribed because of uncertainty as to its true habitat. It is believed to have originated in Cuba. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 401 33. Wromy@ubae2 =o EE ta ee los ea 10. caribaeus AEST TUM eR VNC ee ae es 11. silvaticus 14. Pronotal punctures very coarse and deep_~___----_--__--~.-- 13. trinitatis Pronotal punctures; moderates= soos 25 Se ae ee 12. waterhousei 1. PHILONTHUS HEPATICUS Erickson Philonthus hepaticus ER1cHson, 1840, p. 451, Philonthus vilis Ertcuson, 1840, p. 451. Philonthus orphanus EricHson, 1840, p. 452. Philonthus nanus MELSHEIMER, 1846, p. 36. (Not Gravenhorst, 1806.) Philonthus cinctutus MELSHEIMER, 1846, p. 37. Philonthus palleolus MELSHEIMER, 1846, p. 37. Staphylinus rufipennis Sorrer, 1849, p. 317. (Not Fabricius, 1801; not Graven- horst, 1802.) Philonthus varicolor BOHEMAN, 1858, p. 29. (Not variticolor Nordmann, 1837.) Philonthus pyropterus Kraatz, 1859a, p. 12 (as new name for rufipennis Solier, not Gravenhorst). Philonthus rufipennis (Solier) FAIRMAIRE and GERMAIN, 1861, p. 451. (Not Gravenhorst, 1802.) Philonthus pauwvillus SousKy, 1868, p. 133. Piilonthus cinctulus Horn, 1884, p. 223. (Error; not Gravenhorst, 1802.) Philonthus parvimanus SHARP, 1885, p. 406. Description.—Head black, pronotum rufopiceous, elytra and abdo- men piceocastaneous with apical borders narrowly paler. Head suborbicular, posterior angles entirely rounded; eyes at about their length from base; disk impunctate except for two large punctures forming (with two supraocular) a row between the eyes; with very fine punctulae and strigulose ground sculpture. Pronotwm one- twelfth longer than wide, distinctly narrowed in front with sides straight, with base completely rounded into sides; with two discal series of three punctures, and with about four lateral punctures; ground sculpture as on head. Elytra with moderate but rather indefinite punctures generally separated by their diameter or less and sometimes indistinctly submuricate; without ground sculpture. Abdomen finely and irregularly but not densely submuricately punc- tate; with only a trace of transverse ground sculpture. Male, eighth sternite with an abrupt triangular emargination twice as wide as deep ; seventh sternite not at all emarginate. Female, eighth sternite rounded. Length, 4 to 7 mm. Type locality —‘America septentrionali.” Of vilis, St. Thomas; of orphanus, “Caracas Columbiae”; of cénctutus, nanus, and palleo- lus, Pennsylvania; of parvimanus, (Central America) ; of pauwillus, Mexico; of rufipennis and pyropterus, Chile; of varicolor, California. Types—Presumably in the Zoologische Museum, Berlin. Of vilis and orphanus, either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin; of cinctutus, nanus, and palleolus, in the Museum of Comparative Zoology, Cambridge, Mass.; of par- 402 BULLETIN 182, UNITED STATES NATIONAL MUSEUM vimanus, presumably in the Zoologische Museum, Leningrad; of rufipennis and pyropterus, not known; of varicolor, in Naturhisto- rische Riksmuseum, Stockholm; of pauwwillus, in Zoologische Museum, Leningrad. Records.—The following are the records known to me: Bahamas: Nassau (Busck, in U.S.N.M.). Cuba: (Duval, 1857; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; as vilis), Cayamas (Schwarz, in U.S.N.M.). Jamaica: Santa Cruz (Blackwelder station 421), Spanish Town (Blackwelder station 877), Kingston (Blackwelder station 1C), Trinityville (Blackwelder station 428), Milk River (Blackwelder sfation 415), May Pen (Black- welder station 425C), Gordon Town (Blackwelder stations 368B, 382, 383), Newport (Blackwelder station 412A), Troy (Blackwelder station 409). Hispaniola: (Leng and Mutchler, 1914, as vilis) ; Harri, Trouin (Blackwelder station 24B), St. Mare (Mann, in M.C.Z. and U.S.N.M.); Dominican RE- PUBLIC, Boca Chica (Blackwelder station 41), San José de las Matas (Black- welder station 38; Darlington, in M.C.Z.), Mount Diego de Ocampo (Darling- ton, in M.C.Z. and U.S.N.M.), Sanchez (Darlington, in M.C.Z.). Puerto Rico: Hormigueros and Lajas (Dozier, in U.S.N.M.), Maricao (Black- welder station 47A), San Juan (Blackwelder stations 483A, 45, 345, 353B), Adjuntas (Blackwelder station 48C), Guanica (Smyth, in collection of Rio Piedras Experiment Station). St. Thomas: (WHrichson, 1840; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; as vilis). St. Croix: (Blackwelder stations 317, 325, 330). St. Kitts: (Blackwelder stations 299, 302, 304, 306, 312). Antigua: (Blackwelder stations 275A, 279B, 281A, 285B; H. E. Box, as Black- welder stations 444K, 4440). Montserrat: (Blackwelder stations 262, 265B, 270). Guadeloupe: (Leng and Mutchler, 1914, as vilis), Basse Terre (Blackwelder stations 75, T8B). Les Saintes: (Fleutiaux and Sallé, 1889, as vilis). Dominica: (Blackwelder stations 252, 254B). Barbados: (Blackwelder stations 193B, 194, 195, 201A, 203). St. Vincent: (Blackwelder stations 168, 175). Grenadines: Carriacou (Blackwelder station 144B). Tobago: (Blackwelder stations 117A, 123). Trinidad: Port of Spain (Blackwelder stations 115, 130). South America: VENEZUELA (Fauvel, 1891), ArGeNTINA (Lynch, 1884), CuILe (Solier, 1849; Fauvel, 1864; Fairmaire and Germain, 1861), CotomsBra (Erichson, 1840, as orphanus). Central America: MExIco, GUATEMALA, NICARAGUA, PANAMA (Sharp, 1885, as hepaticus and vilis), Mexico (Villada, 1901), North America: (Hrichson, 1840), Carrrornta, NevapA, ARIZONA, PENNSYLVANIA, Kansas (Boheman, 1858; Horn, 1884; Melsheimer, 1846). Australia: (Fauvel, 1878). Specimens examined.—I have seen from the West Indies 2 exam- ples in the British Museum, 6 from the Museum of Comparative Zoology, 6 in the United States National Museum, and 148 collected by me during 1935-87. In addition series are available from North, Central, and South America. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE AQ3 Remarks.—This species occurs throughout the Americas. It is somewhat variable in coloring and in the distinctness of elytral and abdominal punctation. The description was drawn from a Jamaican series. With one possible exception, this is our only species with only three punctures in the pronotal series. My specimens were collected from dung, manure, and excrement, from under seaweed on the beach, from rubbish along streams, from among decaying forest debris, and flying at dusk. 2, PHILONTHUS THERMARUM Aubé Philonthus thermarum AuBk&, 1850, p. 316. Philonthus exilis KRAAtz, 1851, p. 298. Phitonthus angustatus KRaatz, 1859b, p. 92. Philonthus pygmaeus KRraAatz, 1859b, p. 98. (Not Snellen.) Gabrius fuscolaterus MOTSCHULSKY, 1859, p. 76. Xantholinus inclinans WALKER, 1859, p. 51. Gabrius thermarum (Aubé) MULSANT and Rey, 187Ta, p. 546. Philonthus fuscolaterus (Motschulsky) Fauvet, 1903b, p. 271. Piiilonthus inclinans (Walker) FAUVEL, 19035b, p. 271. Philonthus (Gabrius) thermarum Aubé, REITTER, 1909, p. 131. Description —Head black or piceous, pronotum piceous to rufous, elytra and abdomen castaneous, paler at sides and sometimes pices- cent. Head ovate, slightly wider behind the eyes, basal angles mod- rately rounded; eyes at nearly twice their length from base; disk impunctate except for two large punctures forming (with the two supraocular) a row between the eyes, and a similar pair posterior to these; with very fine and indistinct ground sculpture. Pronotum nearly one-sixth longer than wide, sides arcuate but not distinctly narrowed in front, basal angles completely rounded; with two discal series of four punctures, the anterior more distant, and with about five lateral punctures; ground sculpture as on head. Llytra with moderate punctures separated by about their diameter by slightly irregular but shining intervals. Abdomen very finely and moder- ately sparsely submuricately punctured; with very indistinct ground sculpture. J/ale, eighth sternite with a triangular emargination about twice as wide as deep but almost completely filled by a trans- lucent membrane. Female, eighth sternite rounded. Length, 3 to 4 mm. Type locality —Paris. Of angustatus, “insula Ceylan et in India ovientali”; of pygmaeus, same; of fuscolaterus, “les montagnes de Noura-Ellia 4 Ceylan”; of eatlis, Brandenburg; of inclinans, Ceylon. Types.—Presumably in the Musée National d’Histoire Naturelle, Paris. Of angustatus, ewilis, and pygmaeus, presumably in the Deutsche Entomologische Institut, Berlin; of fwscolaterus, presum- ably in the Zoological University Museum, Moscow; of zénclinans, probably in the British Museum. 404 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records —The following are the records known to me: Jamaica: (Gowdey, 1926), Troy (Blackwelder station 409). Hispaniola: Harirr, Cul-de-Sac Plaine (Blackwelder station 80). Puerto Rico: Lajas, Guanica, Sabana Grande, and Boqueron (Dozier, in U.S- N.M.), Maricao (Blackwelder station 47A). St. Croix: (Blackwelder station 321). Antigua: (Blackwelder stations 275B, 280B, 285B). Guadeloupe: Basse Terre (Blackwelder station 75). Grenada: (H. H. Smith, in British Museum and Cameron collection). South America: FrReNcH GUIANA (Fauvel, 1872). North America: (Fauvel, 1872), Missourr, Districr or CotuMBIA (Horn, 1884). Europe: (Aubé, 1850; Kraatz, 1851, 1857; Fauvel, 1872; Fairmaire and Laboul- bene, 1856; Gangibauer, 1895; Everts, 1898, 1922; Johansen, 1914; Portevin, 1929). Asia and Orient: CHiNA, JAPAN, SUMATRA, CEYLON, JAVA, New CALEDONIA (Fauvel, 1872; Ganglbauer, 1895; Kraatz, 1859; Motschulsky, 1859; Walker, 1859; Sharp, 1889; Fauvel, 1903; Cameron, 1932). Africa: Maperra (Fauvel, 1872), Mapverra and East Arrica (Gangibauer, 1895). Specimens examined.—I have seen 6 examples from the West In- dies in the British Museum, 1 in Dr. Cameron’s collection, 5 in the United States National Museum, and 11 collected by me during 1935-37. Remarks.—This species is very widely distributed in the temperate parts of the Northern Hemisphere. It is distinguished from other West Indian Philonthus by its small size and its elongate oval head. The description was drawn principally from an example from Antigua. AXantholinus breviventer Sperk has been included in the synonymy of this species. Since it was published before thermarum (1885), it would have priority over that name, but since it seems likely that Sperk’s species really was a Scopaeus, I believe it better to drop the name until it can be proved that it belongs here. My specimens were found in dung and manure and in rotting grass. 8. PHILONTHUS VENTRALIS (Gravenhorst) Staphylinus ventralis GRAVENHORST, 1802, p. 174. (Not Gené, 1830.) Staphylinus anthravzy GRAvENHORST, 1802, p. 176. Staphylinus immundus GRAVENHORST, 1806, p. 66. Staphylinus celer GRAVENHORST, 1806, p. 66. Staphylinus rotundiceps Kirpy, MS.—STEPHENS, 1829, p. 283. Philonthus ventralis (Gravenhorst) STEPHENS 1832, p. 232. Philonthus picicollis STEPHENS, 1832, p. 224. Quedius fulvipes STEPHENS, 1832, p. 229. (Not Fabricius, 1792.) Bisnius rotundiceps StrrHENsS, 1832, p. 248. Philonthus anthrav (Gravenhorst), NFRDMANN, 1837, p. 473. Philonthus proximus WOLLASTON, 1857, p. 189. (Not Kraatz, 1859.) Philonthus celer (Gravenhorst) Kraarz, 1857a, p. 596. (Not Heer, 1888.) Philonthus fortunatus WoLLAstTon, 1865, p. 493. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 405 Philonthus rotundiceps (Stephens) MULSANT and Rey, 1875, p. 483. Philonthus immundus (Gravenhorst) FLEUTIAUX and SALLE, 1889, p. 380. Philonithus fulvipes (Stephens) FLeuTraux and Sarit, 1889, p. 380. (Not Fabricius, 1792.) Description—Head black, pronotum rufopiceous, elytra and ab- domen piceocastaneous, margins of elytra sometimes narrowly tes- taceous. Head suborbicular or subquadrate with angles rounded, not narrowed behind the eyes; eyes at about their length from base; sometimes with an irregular depression in front; disk with two large punctures forming (with two supraocular) a line between the eyes, the supraocular punctures being placed a little more anteriorly, and with two large punctures posteriorly at sides of vertex; with very Sne strigulose ground sculpture. Pronotum scarcely longer than wide, with base completely rounded, but not distinctly narrowed in front; with two discal series of four punctures, the posterior more distant, with another puncture not in line but not clearly part of the anterior marginal series, and with about four lateral punctures; ground sculpture more distinct than on head. /lytra with moderate punctures distinct and regular, generally separated by about their diameter; intervals smooth. Abdomen with very fine and moder- vtely sparse submuricate punctures; with a vague trace of strigulose ground sculpture, and with iridescent lustre. JJ/ale, eighth sternite with a very feeble emarginate sinuation and a triangular area in front of it marked by the absence of punctures. Female, eighth sternite rounded. Length, 5 to 7 mm. Type locality —‘Lusitania.” Of immundus, Paris; of celer, Paris; of anthrax, “America septentr.”; of picicollis, England; of fulvipes, England; of fortwnatus, Teneriffe; of proximus, Madeira; of ro- tundiceps, England. Types.—Probably in the Zoologische Museum, Berlin. Of anthrax, immundus, and celer, probably in the Zoologische Museum, Berlin; of fulvipes, in the British Museum; of fortwnatus, in the British Mu- seum; of proximus, in the British Museum; of picicollis, in the British Museum; of rotundiceps, in the British Museum. Records.—The following are the records known to me: Cuba: Cayamas (Casey collection, in U.S.N.M.), Soledad (Darlington, in M.C.Z.). Jamaica: (Cameron collection; Cameron in Blackwelder collection; Gowdey, 1928), Santa Cruz (Blackwelder station 421), Milk River (Blackwelder sta- tion 415), Spanish Town (Blackwelder station 377), Moneague (Blackwelder station 875), Kingston (Blackwelder station 1C), Trinityville (Blackwelder station 428), May Pen (Blackwelder station 425C), Black River (Black- welder station 420), Newport (Blackwelder station 412A). Hispaniola: DoMInNICAN Repusiic, Santiago (Blackwelder station 33A), San José de las Matas (Blackwelder station 38). 406 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Puerto Rico: Lajas, Juana Diaz, Ensenada (Dozier, in U.S.N.M. and in Wolcott collection), Mameyes (Blackwelder station 57), Guanica (Blackwelder sta- tion 357B), San Juan (Blackwelder stations 43B, 51, 845, 358, 45), Lake Cartagena (Blackwelder station 46A), Adjuntas (Blackwelder station 48C), Humacao (Blackwelder station 54). Culebra Island: (Busck, in U.S.N.M.). St. Thomas: (Blackwelder stations 66, 67). St. Croix: (Blackwelder stations 317, 325, 327, 328, 330, 331, 339, 344). St. Kitts: (Blackwelder stations 296, 297, 300, 802, 304, 305, 306, 312). Antigua: (Blackwelder stations 273, 275A, 275B, 280B, 281A, 284, 285B, 287, 290, 292, 294, 444K). Montserrat: (Blackwelder stations 265B, 267B). Guadeloupe: Basse Terre (Blackwelder stations 70, 75, 77B), Grande Terre (Blackwelder stations 86, 89, 90B). Dominica: (Blackwelder stations 286, 240, 257A, 257B, 258). St. Lucia: (Blackwelder stations 210, 212, 218). Barbados: (Blackwelder stations 189, 194, 195, 198A, 199, 201A, 203). St. Vincent: (Erichson, 1840; Fauvel, 1872; Blackwelder stations 165, 172, 185). Grenada: (British Museum; Blackwelder stations 1381, 189, 151, 161). Tobago: (Blackwelder station 123). Trinidad: Port of Spain (Blackwelder stations 110, 130). South America: CAYENNE (Fauvel, 1872). Worth America: (Erichson, 1840; Fauvel, 1872), IowA, VirGINIA, CALIFORNIA, Texas, NEw York (Casey collection, in U.S.N.M.), NEw Mexico (Cameron collection), MicHIGAN, Missouri, NEvADA (Horn, 1884). Europe: GERMANY, SWEDEN, FRANCE, CYPRUS, ENGLAND, FINLAND (Erichson, 1840; Kraatz, 1857; Thomson, 1860; Fauvel, 1872; Fowler, 1888). Asia: Asta MINoR, CAucASUS, TRANSCAUCASUS, SrBEeRTA, InpIA (Fauvel, 1872; Cameron, 1932). Africa: ALGERIA, MADEIRA, CANARIES, CAPE VERDES (Fauvel, 1872). Specimens examined.—I have seen 6 examples from the West Indies in the British Museum, 7 in Dr. Cameron’s collection, 1 in my own col- lection, 4 in the collection of Dr. G. N. Wolcott, 1 in the Museum of Comparative Zoology, 11 in the United States National Museum, and 363 collected by me during 1935-37. Remarks.—This is our most common and widespread Philonthus. It undoubtedly occurs on all the islands, though it has not been recorded from Martinique and some of the small islands. It is readily dis- tinguished from most of our species by the characters given in the key, but from discoideus it is sometimes separated with difficulty. The latter species has the humerus and borders of elytra testaceous and has slight differences in the male characters. My specimens were collected from dung and manure, from a pile of rotting grass, from the refuse of lime and grapefruit canneries, and flying at dusk. 4. PHILONTHUS AUDANTI, new species Description—Black or piceous, pronotum distinctly rufescent, ab- domen iridescent. ead suborbicular but with posterior angles vis- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 407 ible; eyes at about one-third their length from base; without depres- sion in front; vertex without punctures except for one above each eye but not forming a line of four with the supraocular punctures with two of which each of the discals forms a small triangle; without distinct ground sculpture but with some luster. Pronotwm as wide as long, base and angles completely rounded, feebly narrowed in front; with two discal series of four punctures, and with about four lateral punctures; with faint luster as on head. //ytra with punctures rather indistinct and indefinite, appearing as a cluster of minute punctures; surface fairly even and without ground sculpture. Abdomen very sparsely and moderately finely submuricately punctate; without ground sculpture but with strong iridescence. Jfa/e, eighth sternite with a small rounded emargination, three times as wide as deep, and with a rather wide smooth border. Female, unknown. Length, ol4 mm. Type locality—Haiti, 5 miles south of St. Mare. Types.—Holotype, male, U.S.N.M. No. 52465, collected by me on August 15, 1935. Records.—The following is the only record known to me: Hispaniola: Hariri, St. Mare (Blackwelder station 26). Specimens examined.—I have seen only the unique type. Remarks.—This species differs from all others in the West Indies in the position of the discal pair of punctures on the head and in the elytral punctation. It is distinguished from figulus by its much smaller size as well as the characters mentioned in the key. The type was found under dung. 5. PHILONTHUS DISCOIDEUS (Gravenhorst) Staphylinus discoideus GRAVENHORST, 1802, p. 38. Staphylinus suturalis MARSHAM, 1802. p. 509. (Not Panzer, 1794.) Staphylinus testaceus GRAVEN HORST, 1806, p. 60. Staphylinus lepidulus Krrpy, MS.—STEPHENS, 1829, p. 278. Quedius suturalis (Marsham) STEPHENS, 1829, p. 278. (Not Kiesenwetter, 1845; not Thomson, 1867.) Quedius lepidulus STEPHENS, 1882, p. 223. Staphylinus conformis BoIspUVAL and LACoRDAIRE, 1835, p. 398. Philonthus discoideus (Gravenhorst) NorDMANN, 1836, p. 97. Philonthus suturalis (Marsham) NorpMANN, 1836, p. 91. Phiionthus ruficornis MELSHEIMER, 1846, p. 38. (Not Motschulsky, 1860; not Hochhuth, 1860; not Broun, 1880.) Philonthus conformis (Boisduval and Lacordaire) GANGLBAUER, 1895, p. 455. (Not Sharp, 1876.) Philonthus rufipennis GerHAxrpT, 1910, p. 555 (as aberration). (Not Gravenhorst, 1802; not Solier, 1849.) Philonthus lepidulus (Stephens) BerNHAUER and ScHUBERT, 1914, p. 336. (Not LeConte, 1863.) Philonthus testaceus (Gravenhorst) BERNHAUER and ScHusert, 1914, p. 336. 408 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Philonthus discoidens CAMERON, 1933, p. 42 (misspelling). Philonthus gerhardtianus SCHEERPELTZ, 1933, p. 13840 (new name for rufipennis). Description.—Head black, pronotum rufopiceous to rufous, elytra and abdomen castaneous, sometimes picescent, elytral suture and posterior and lateral margins narrowly testaceous. Head subquad- rate with posterior angles moderately rounded; eyes at about two- thirds their length from base; without distinct impression in front; disk with two punctures forming (with two supraocular) a row of four between the eyes, the supraoculars being placed a trifle more anteriorly, and with a pair of lateral discal punctures just back of the eyes; with minute punctulae and very fine strigulose ground sculpture. Pronotum as broad as long, scarcely narrowed in front, base com- pletely rounded; with two discal series of four punctures, with another puncture not in line but not clearly part of the anterior marginal series, and with about five lateral punctures; with ground sculpture as on head. lytra with moderate punctures of indefinite shape, generally not separated by their diameter; without distinct ground sculpture. Abdomen with fine and rather dense submuricate punc- tures; with indistinct transverse strigulose ground sculpture. Male, eighth sternite with a small triangular or rounded emargination over four times as wide as deep and preceded by a triangular smooth area. Female, eighth sternite rounded. Length, 5 to 6144 mm. Type locality—(Not recorded.) Of suturalis, not recorded; of ruficornis, Pennsylvania; of lepidulus, England ; of testaceus, Sweden; of conformis, Paris; of gerhardtianus and rufipennis, Liegnitz. Types.—Probably in the Zoologische Museum, Berlin. Of ruficornis, probably in the Museum of Comparative Zoology, Cambridge, Mass.; of lepidulus, in the British Museum; of swturalis, possibly in the British Museum; of testaceus, in Naturhistorisk Riksmuseum, Stockholm; of conformis, unknown; of rufipennis, presumably in collection at Hed- wigsschule, Liegnitz. Records.—The following are the records known to me: Cuba:(Erichson, 1840; Fauvel, 1872; British Museum). Jamaica: (Cameron collection), Spanish Town (Blackwelder station 377). Hispaniola: Hair, Kenskoff (Blackwelder station 23B). Puerto Rico: San Juan (Blackwelder station 43B). St. Croix: (Blackwelder station 344). St. Kitts: (Blackwelder stations 297, 299, 302, 305). Antigua: (Blackwelder stations 275B, 290). Montserrat: (Blackwelder station 267B). Guadeloupe: Grande Terre (Blackwelder station 89). Grenada: (British Museum). North America: (Iauvel, 1872), ArLaNTic States to Nevapa and Arizona (Horn, 1884), NrvApA, ArIzoNA, VirGINIA, New York, Texas (Casey collection, in U.S. N. M.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 409 Europe: GeRMANY, FRANCE, SWEDEN, Russia, ENGLAND, FINLAND, DENMARK, Cyprus (Erichson, 1839, 1940; Fauvel, 1872, Mulsant and Rey, 1875; Nord- mann, 1836; ete.) Africa: BARBARIE, MADEIRA, CANARIES, CAPE VERDES (Fauvel, 1872). Asia and Orient: Srperta, Asta Minor, INpIA, HAwarr (Nordmann, 1886; Fauvel, 1872; Cameron, 1932; Sharp, 1908). Specimens exwamined.—I have seen 7 examples from the West Indies in the British Museum, 1 in Dr. Cameron’s collection, and 14 collected by me during 1935-37. Remarks. —This species is very similar to ventralis, but apparently it can be distinguished by the testaceous margins (especially the sutural) of the elytra. The emargination of the eighth sternite in the male is generally deeper and more angular than in ventralis, where it is merely a slight sinuation of the margin. I usually collected this in company with much more numerous examples of ventralis, and I am not certain that there are really two species involved. My specimens were found in dung, in a pile of rotting grass, and flying at dusk. 6. PHILONTHUS FIGULUS Erichson Philonthus figulus ErtcHson, 1840, p. 464—SAHLBERG, 1844, p. 790.—CHEVROLAT and FAvvEL, 1863, p. 484.—SHarp, 1885, p. 415.—FAUvVEL, 1891, p. 115.—BErn- HAUER and ScuHusBeErT, 1914, p. 888.—LENG and MUTCHLER, 1914, p. 406.— PLAVILSTSCHIKOv, 1929, p. 80.—SCHEERPELTZ, 1983, p. 1342. Philonthus amnazonicus SHARP, 1876, p. 165; 1885, pp. 415, 416—Favver, 1891, p. 115.—BrrRNHAUER and SCHUBERT, 1914, p. 338.—LENG and MuTcHLer, 1914, p. 406. Description —Black. Head suborbicular; eyes at less than half their length from base; without impression in front; disk impunctate except for two punctures between the eyes but close to the supraocular series and not forming a line of four with any pair of the latter; with- out distinct ground sculpture. Pronotum scarcely longer than wide, posterior angles completely rounded; feebly narrowed in front; with two discal series of four punctures, the posterior more distant, and with about four lateral punctures; with minute punctulae but without ground sculpture. /ytra with rather fine, posteriorly-directed, some- times submuricate punctures generally separated by about their di- ameter transversely; without ground scultpure. Abdomen with mod- erately fine but not dense submuricate punctures (a little denser pos- teriorly); iridescent but without distinct ground sculpture. JMale, eighth sternite with triangular emargination about one-third as deep as wide and with the outer angles broadly rounded. Female, eighth sternite with a feeble rounded emargination not over one-fifth as deep as wide. Length, 7 to 9 mm. Type locality —Brazil. Of amazonicus, “Para, Ega, Amazons Val- ley,” Brazil. 449008—42——27 410 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Types——Presumably in the Zoologische Museum, Berlin. Of amazonicus, in the British Museum. Records.—The following are the records known to me: Cuba: (Leng and Mutchler, 1914, as amazonicus), Habana (Fauvel, 1863, 1891; Bierig, in Cameron collection). Puerto Rico: Maricao (Blackwelder station 47A). St. Vincent: (H. H. Smith, in British Museum). Trinidad: St. Augustine (Darlington, in M.C.Z.). South America: Braziu (Erichson, 1840; Sahlberg, 1844; Sharp, 1876, as amazoni- cus ; 1885, as figulus), BraziL, CAYENNE (Fauvel, 1863), VENEZUELA (Fauvel, 1891; British Museum), ARGENTINA (Scheerpeltz, 1983), Peru (Plavilstschi- kov, 1929). Central America: Mrexico, GUATEMALA, NICARAGUA, CosTA Rica, PANAMA (Fauvel, 1891; Sharp, 1885), GuATEMALA (Cameron collection). Specimens examined.—I have seen from the West Indies one ex- ample in the British Museum, one in Dr. Cameron’s collection, one from the Museum of Comparative Zoology, and nine collected by me during 1935-37. Lemarks.—This species may be distinguished by its black color, the interocular punctures not forming a straight row of four, and the definite though small elytral punctures. It is the only West Indian species known to me to have the eighth sternite of the female emar- ginate and differing from that of the male only in degree. My specimens were found under excrement. 7. PHILONTHUS LONGICORNIS Stephens Philonthus longicornis STEPHENS, 1882, p. 237. Philonthus scybalarius NoRDMANN, 1836, p. 94. Philonthus fuscicornis NorpMANN, 1886, p. 96. Philonthus feralis Ertcuson, 1840, p. 469. Philonthus promtus ErtcHson, 1840, p. 929. Philonthus varians FATRMAIRE, 1849, p. 290 (not Paykull). Philonthus algiricus MoTtscHULSKy, 1858a, p. 668 . Philonthus pedestris WALKER, 1859, p. 51. Philonthus asemus KRAATz, 1859b, p. 86. Philonthus perplerus FAIRMAIRE and GERMAIN, 1861, p. 481. Philonthus fumosus SousxKy, 1868, p. 134. Philonthus promptus Pettit, 1870, p. 65 (misspelling). Cafius laetabilis OLLIFF, 1887, p. 501. Philonthus laetabilis (Oliff) Lea, 1899, p. 540. Philonthus linkei BERNHAUER, 1908c, p. 34 (as variety). Philonthus rubromaculatus BERNHAUER, 1915e, p. 9. (Not Blanchard, 1843.) Philonthus piceicornis GRIDELLI, 1920, pp. 18, 19. Philonthus lactabilis Everts, 1922, p. 124 (misspelling). Description.—Black, elytra piceocastaneous. Head orbicular; eyes at less than their length from base; not impressed in front; disk im- punctate except for two large punctures forming (with two supra- ocular) a line between the eyes; with indistinct strigulose ground MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 411 sculpture. Pronotum scarcely longer than wide, with base broadly rounded but angles discernible, sides arcuately but distinctly converg- ing in front; with two discal series of four punctures, the posterior more distant, and with about five lateral punctures; ground sculpture as on head. Hlytra with surface irregular and punctures directed posteriorly, rather dense, and often appearing submuricate; without ground sculpture. Abdomen with very fine and not very dense punc- tures; iridescent but without distinct ground sculpture. A/ale, eighth sternite with an emargination about three times as wide as deep but preceded on the segment by a triangular smooth area about as deep as wide. Female, eighth sternite truncate. Length, 7 to 8 mm. Type locality—England. Of algiricus, Algeria; of fumosus, Mex- ico; of fuscicornis, Cape of Good Hope; of Jaetabilis, “Port Lincoln, South Australia; Hobart, Tasmania”; of perplexus, Santiago, Chile; of promtus, Pennsylvania; of scybalarius, Berlin; of asemus, “India boreali”; of feralis, Colombia; of pedestris, Ceylon; of piceicornis, not recorded; of linkei, Germany; of rubromaculatus, southern Italy. Types.—In British Museum. Of promtus, probably in the Zoo- logische Museum, Berlin; of feralis, either in the Hope Museum, Ox- ford, or the Zoologische Museum, Berlin; of perplexus, probably in either the Musée Royal d’ Histoire Naturelle Belgique, Brussels, or the Museo Nacional, Santiago, Chile; of pedestris, probably in the British Museum; of Jaetabilis, probably in the Australian Museum, Sydney; of piceicornis, in the Museo Entomologicae “Pietro Rossi,” Duino; of fuscicornis, “Mus. Reg. Berol.” ; of scybalarius, unknown; of algiricus, probably in the Zoological Museum, Moscow; of asemus, in the Deutsche Entomologische Institut, Berlin-Dahlem; of fwmosus, in the Zoological Museum, Leningrad; of /inkei and rubromaculatus, presumably in the collection of Max Bernhauer. Records.—The following are the records known to me: Hispaniola: Harri, Kenskoff (Blackwelder station 23E). Puerto Rico: Lajas (Dozier, in U.S.N.M.). St. Kitts: (Blackwelder station 305). Antigua: (Blackwelder station 280B). Montserrat: (Hubbard, in U.S.N.M.). Grenada: (British Museum). South America: CorompBia (Erichson, 1840, as feralis), ARGENTINA, CHILE (British Museum). Central America: Mexico (Solsky, 1868, as fumosus). North America: “INverywhere” (Horn, 1884; Erichson, 1840). Europe: (Stephens, 1832; Fauvel, 1872; Ganglbauer, 1895; Nordmann, 1836; Thomson, 1860; Reitter, 1909; Johansen, 1914). Africa: Maprrras, CANARIES, AZORES, CAPE VERDES, ASCENSION, CAPE OF GooD Hope, MADAGASCAR (Fairmaire, 1849; Wollaston, 1871; Fauvel, 1872). Asia and Orient: Smperra, JAPAN, CEYLON, TAHITI, AUSTRALIA, NEw CALEDONIA, TASMANIA, Hona Kona, SUMATRA (Fairmaire, 1849; Fauvel, 1872; Olliff, 1887; Cameron, 1932; British Museum). 412 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I\ have seen five examples from the West Indies in the British Museum, two in the United States National Museum, and three collected by me during 1935-87, Lemarks.—This species is widespread in both hemispheres but is generally not common. It is one of the few all black species in our region and can be distinguished by the combination of characters given in the key. My specimens were all taken from dung, but the Hubbard example from Montserrat was taken “in refuse of arrowroot.” 8. PHILONTHUS VARIANS (Paykull) Staphylinus varians PAYKULL, 1789, p. 45. Staphylinus nitens GRAVEN HORST, 1802, p. 26. (Not Schrank, 1781.) Staphylinus opacus GRAVENHCRST, 1802, p. 26. (Not Block, 1799 ; not Roth, 1851.) Staphylinus bipustulatus GRAVENHORST, 1802, p. 87. (Not Linnaeus, 1761.) Staphylinus aterrimus MARSHAM, 1802, p. 513. (Not Gravenhorst, 1802.) Staphylinus bimaculatus MARSHAM, 1802, p. 525. (Not Schrank; not Graven- horst, 1802.) Staphylinus nigrans DEJEAN, 1821, p. 22.—DAHL, 1823, p. 16. Staphylinus unicolor Kirpy, MS.—STEPHENS, 1829, p. 278 (nomen nudum). (Not Herbst, 1784.) Philonthus varians (Paykull) STEPHENS, 1829, p. 280. Philonthus opacus (Gravenhorst) STEPHENS, 1829, p. 281. (Not LeConte, 1863.) Philonthus aterrimus (Marsham) STEPHENS, 1829, p. 281. Philonthus bipustulatus (Fabricius) STEPHENS, 1829, p. 281. Staphylinus intaminatus Kirpy, MS.—STEPHENS, 1829, p. 281. Staphylinus lituratus (Samouelle) STEPHENS, 1829, p. 281 (nomen nudum). Staphylinus punctiventris Kirpy, MS.—STEPHENS, 1829, p. 281. Quedius unicolor STEPHENS, 1852, p. 224. (Not Kiesenwetter, 1847.) Philonthus punctiventris STEPHENS, 1832, p. 255. (Not Kraatz, 1857.) Philonthus intaminatus STEPHENS, 1882, p. 235. Philonthus nitens (Gravenhorst) STEPHENS, 1882, p. 236. Philonthus lituratus STEPHENS, 1882, p. 238. Staphylinus nigricans ZETTERSTEDT, 1840, p. 65. Philonthus costatus Baupt, 1848, p. 29. Quedius incompletus HocHHuTH, 1849, p. 153. Philonthus alpigradus MuLsant and Rey, 1875, p. 481. Philonthus scutatus EprpeLSHEIM, 1895, p. 127. (Not Erichson, 1840.) Philonthus unicolor (Stephens) Rerrrer, 1909, p. 180. (Not Kiesenwetter, 1847.) Philonthus incompletus (Hochhuth) BERNHAUER, 1915e, p.9. (Not Roubal, 1911.) Philonthus bimaculatus (Marsham) BERNHAUER and SCHUBERT, 1914, p. 359.) (Not Gravenhorst, 1802.) Philonthus piceicoris GRIDELLI, 1920, pp. 18, 19. (Not Eppelsheim, 1895.) Philonthus proteus Everts, 1922, p. 124. Philonthus fuscicoris SCHEERPELTZ, 1933, p. 1866 (as aberration). Description.—Black, elytra with indefinite reddish subsutural spot in apical two-thirds. Head scarcely wider than long, subquadrate ; eyes at about their length from base; not impressed in front; disk impunctate except for two large punctures forming (with two supra- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 413 ocular) a line between the eyes; with rather indefinite strigulose ground sculpture. Pronotwm as wide as long, base rounded with angles scarcely discernible, distinctly narrowed in front; with two discal series of four punctures, the posterior more distant, and with about four lateral punctures; ground sculpture as on head. Llytra without distinct punctures but with posteriorly directed setae arising from very small submuricate punctures on surface irregularities. Abdomen sparsely and moderately finely submuricately punctured ; with scarcely a trace of ground sculpture. fale, eighth sternite with a large triangular emargination a little wider than deep and pre- ceded by a smooth triangular depression. Female, eighth sternite rounded. Length, 7 mm. Type locality—Sweden. Of alpigradus, Auvergne, France; of nitens, Brunswick, Germany; of punctiventris, Suffolk, England; of unicolor, England; of aterrimus (Great Britain) ; of piceicoxis, not recorded; of intaminatus and lituratus, England; of opacus, Bruns- wick, Germany; of fuscicovis, north Italy; of incompletus, Tiflis, Caucasus; of costatus, Sardinia; of scutatus, Gabon, West Africa; of bipustulatus, Brunswick, Germany; of bémaculatus, England. Types—In Naturhistisk Riksmuseum, Stockholm. Of piceicomis, in Museo Entomologicae “Pietro Rossi,” Duino; of intaminatus, lituratus, unicolor, and punctiventris, in British Museum, London; of aterrimus, probably in the British Museum; of nitens and bipustu- latus, in Zoological University Museum, Breslau; of opacus, prob- ably in Zoologische Museum, Berlin; of alpigradus, possibly in Musée d'Histoire Naturelle, Lyon; of incompletus, presumably in Zoologische Kabinett dem Polytechnic Institut, Kiev; of fuscicowis, probably in collection of Otto Scheerpeltz; of costatus, in Zoological University Museum, Turin; of scwtatus, in Naturhistorische Museum, Wien; of bimaculatus, unknown, Records.—The following are the records known to me: Cuba: (Erichson, 1840). St. John: (Erichson, 1840). St. Vincent: (Erichson, 1840). North America: NorruH ATLANTIC StaTEs to WASHINGTON and CALIFORNIA (Horn, 1884). Europe: Cyprus, Syrta, Asta MINoR, Caucasus, ENGLAND (Erichson, 1840; Kraatz, 1857; Fauvel, 1872; Mulsant and Rey, 1875; Ganglbauer, 1895; Stephens, 1882). Africa: Care or Goop Hopr, BarBARIE, ABYSSINIA, TENERIFFE, CAFFRARIA (Erich- son, 1840; Fauvel, 1872; Boheman, 1848). Asia: SIBERIA, JAPAN, BurMA (Fauvel, 1872; Cameron, 1932). Specimens examined.—I have seen no examples from the West Indies. The description was drawn from a North American example in the Casey collection. 414 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—This species is distinguished from longicornis by the marking of the elytra and the sparse punctures of the abdomen. It also differs in the male characters. I find no record of its habits. 9. PHILONTHUS FLAVOLIMBATUS Erichson Philonthus flavolimbatus Ericuson, 1840, pp. 471, 929.—LeConTe, 1866, p. 348.— Horn, 1884, pp. 190, 191, 192, 247.—Suarp, 1885, p. 416.—CockKERELL, 1898, p. 143.—GrirrirH, 1900, p. 566.—FAxL, 1901, pp. 14, 69.— Knaus, 1905, pp. 349; 1906, p. 380.—F ALL and CockrrELL, 1907, p. 166—Lene and Murcuter, 1914, p. 401.—BernHaAver and Scuusert, 1914, p. 8338.—Norman, 1920, p. 706.—Tor TENHAM, 1939, p. 238. Philonthus apicipennis Lyncu, 1884, p. 155.—FAavvet, 1887, p. 232 (as synonym) .— VILLADA, 1901, p. 30.—BrerNHAUER and ScHupeRrtT, 1914 p. 338.—TorreNHAM, 1939, p. 233. (Not Cameron, 1932.) Description—Head black, pronotum rufopiceous, elytra piceous with apical border testaceous (much more widely near suture), abdo- men piceous with posterior margins rufous. Head suborbicular, pos- terior angles entirely rounded; eyes at about their length from base; disk impunctate except for two large punctures forming (with two supraocular) a row between the eyes; with very fine punctulae and very fine strigulose ground sculpture. Pronotum one twenty-fourth longer than wide, distinctly narrowed in front but with sides feebly arcuate ; base broadly rounded but posterior angles traceable; with two discal series of four punctures, fairly evenly spaced, and with about four lateral punctures; ground sculpture more distinct than on head. Elytra with indefinite posteriorly-directed submuricate punctures, not very dense; without ground sculpture. fale, eighth sternite with a very feeble triangular emargination at least four times as wide as deep, posterior angles rounded. Female, eighth sternite truncate. Length, 414 to 5144 mm. Type locality—Colombia. Of apicipennis, “America meridionali et boreali.” Types.—Lither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of apicipennis, in the Museo Argentina Ciencias Natural Bernard Rivadavia, Buenos Aires. Records.—The following are the records known to me: Martinique: (Bernhauer and Schubert, 1914). Grenadines: Carriacou (Blackwelder stations 143B, 144B). South America: Cotomsta (Erichson, 1840); Braz (Sharp, 1885), ARGENTINA (Lynch, 1884), New GreNADA, BrAziIL, ARGENTINA (Bernhauer and Schubert, 1914). Central America: Mrxico, GUATEMALA, NicaRaGua, PANAMA (Sharp, 1885). North America: CArotina (Erichson, 1840), GutF STATes and Arizona (Horn, 1884), CaLIFoRNIA, FtLortpa (Leng, 1920), Texas, Arizona (U.S.N.M.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE Ath Specimens examined.—I have seen only six examples from the West Indies, collected by me in January 1936. Numerous examples from Southern United States are in the United States National Museum. Remarks.—This species is apparently widely distributed in tropical and subtropical America. Its present known distribution in the West Indies probably indicates that it has been introduced there recently. It is distinguished from other West Indian species by the testaceous border of the elytra, which is very similar to that found in some species of Baryodma. My specimens were all found in cattle dung. 10. PHILONTHUS CARIBAEUS Bierig Philontus (s.str.) caribacus Brrrie, 1940, p. 141, fig. 19, 1. Original description —Longitud como 8 mm.; cabeza 0,9 mm. de ancho, protérax 0,9 por 1.8, élitros 1.4 por 1.5 mm., de ancho por largo, maxima anchura abdominal (3.er segmento) 1.4 mm. Lustroso. De un negro de pez, repliegues laterales y margenes posterio-tergitales del abdomen obscuro-castaiios, élitros aéneos, bucales, escapo antenal y base de los articulos 2 y 3 de un testaceo subido, miembros testaceos, tarsos y extremo antenal mas palidosatin. Cabeza allanada en el plano epicraneal, recta a la base y los temporales algo divergentes, de ancho redondeado en los angulos occipitales, los ojos de poca convexidad; de 2 puntos gemelos superoculares el exterior el mas grosero, al borde occipital mas numerosos puntos que a la gena posterior. Antenas 2.4 mm., engrosadas de modo sucesivo y doblando, finalmente, casi 2 veces el grosor del escapo; éste poco mas largo que el articulo 3, 2.° como de ¥% mas corto y de la longitud del 4.°, siguientes decreciendo paulati- namente, tiltimos algo aplastados y mas cdénicos, 10° tan ancho como largo, terminal inflado, oblicuo-truncado en el dpice. Torax paralelo, costados subsinuados en la mayor mitad antical, mirgenes proximal y distal arqueadas, angulos redondeados, los anteriores desaparecidos por completo (visto desde arriba); disco achagrinado, microscépica- mente, en sentido transverso; puntos bastante obsoletos, los laterales en numero de 4 y en figura de rombo. Elitros de htmeros con- vexos, de lados subrecto-divergentes, la sutura ligeramente abultada dentro de una hondonada leve, integral; punctuacién medio- cremente fina, neta, oblicuo-impresa desde atrds, transversalmente subcontigua; pubescencia de suma sutileza, de color grisaceo. Escudete rugoso-coriiceo y con confusos poros de pelitos acostados, negros. Abdomen cdénico desde el tercer segmento, los puntos tergitales distintamente mas groseros que los elitrales, algo menos apretados, oblongos, piliferos, esparciéndose en las dreas apicales. Estiletes fuscos, sdlo sutilmente pubescentes, espinas tibiales débiles, obscuras. 416 BULLETIN 182, UNITED STATES NATIONAL MUSEUM “En el ¢, solo esta’ cortamente recto-recortado el apice del 6.° esternito abdominal, mientras en la hembra se nota una punta obtusa. Los protarsos estan algo dilatados hacia la base.” Type locality —Cuba. Types.—In the collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: Province of Habana (Bierig, 1940), province of Pinar del Rio (Bierig, 1940). Specimens examined.—I\ have seen no examples of this species. Remarks.— ‘Hace anos, este coleédptero fué determinado por Ram- bousek como Paederomimus sp., y, en efecto, clerta semejanza es inne- gable. Pero un examen concienzudo de los caracteres primordiales de este género y de Philonthus aclaré que ha de afiadirse al tiltimo; sin embargo bajo un subgénero aun acrear. Es de talle esbelto y muy gracil, a la armadura tibial escasea y remeda la del ruso brunicollis Hochh., en el cual, ademas, veo un parentesco estrecho, general. Se sehala luego por el t6rax subcilindrico, un ligero irisar superficial, excepto los élitros aéneos, y la palidez del extremo antenal. En con- cepto erichsoniano entra en el grupo de las especies de 6 puntos pronotales.” See Remarks under genus Philonthus. “Se encuentra durante todo el afio en Ja hojarasca, en los Noviembres principalmente (Bierig) .” 11. PHILONTHUS SILVATICUS Cameron Philonthus silvaticus CAMERON, 1922, p. 119.—-SCHEERPELTZ, 1933, p. 1861. Description —Black, elytra bronze-green. Head broader in front, sides gradually convergent behind the eyes, posterior angles broadly rounded; eyes at less than their length from base; with a pair of widely separated discal punctures between the eyes; without strong ground sculpture. Pronotum nearly cylindrical, scarcely narrowed behind; with two discal series of five punctures, and with about five lateral punctures. “/7ytra moderately finely and moderately closely punctate. Abdomen with the bases of the three basal tergites closely and mod- erately coarsely punctate, the rest much more sparingly and more finely punctate. Zale, eighth sternite with a small rounded emargination. Female, (eighth sternite rounded). Length, 634 to 7 mm. (From Cameron.) Type locality —Jamaica. Types.—In Dr. Cameron’s collection. Records—The following is the only record known to me: Jamaica: (Cameron, 1922). Specimens examined.—I have seen nine examples of this species, including the type, in Dr. Cameron’s collection. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 417 Remarks.—This species is quite similar to what I recognize as havaniensis but is apparently distinguishable by the shape of the pronotum. If, is distinguished from waterhouset and most other species by the greenish color of the elytra. Dr. Cameron described this species as “very black.” Examples of waterhousei that he de- scribes as “black” are generally slightly castaneous or rufescent. It is assumed that this species is actually black and not picescent. I find no record of its habits. 12. PHILONTHUS WATERHOUSEI Cameron Philonthus waterhousei CAMERON, 1922, p. 120.—ScHEERPELTZ, 1933, p. 1569. Description.—Piceous, generally feebly rufescent. Head transverse, somewhat narrowed behind eyes, the latter at distinctly more than their length from base; basal angles rounded; with a slight depression in front; disk with two punctures forming (with two supraocular) a line of four between the eyes, and with scattered punctures at sides and base leaving only the center of vertex impunctate; without dis- tinct ground sculpture. Pronotum about one-third longer than wide, widest at anterior third, feebly emarginately narrowed to basal third, with basal angles broadly rounded; with two discal series of five punc- tures, fairly evenly spaced, and with about seven lateral punctures; with vague ground sculpture. “/ytra with moderate punctures sep- arated by about their diameter; the surface even and without ground sculpture. Abdomen rather irregularly but finely and submuricately punctured; with traces of ground sculpture. J/a/e, eighth sternite with posterior margin sinuate but not distinctly emarginate, bordered at center with a narrow smooth rim. VYemale, eighth sternite rounded. Length, 4 to 5 mm. Type locality.—St. Vincent, Leeward side. Types.—In the British Museum; one “cotype” in Dr. Cameron’s collection, six “cotypes” in the United States National Museum (No. 50878). Records.—The following are the records known to me: St. Vincent: (Cameron, 1922; British Museum; Cameron collection ; U.S.N.M.). Grenada: (British Museum; Cameron collection; Cameron, 1922; U.S.N.M.). Specimens examined.—I have seen 12 examples in the British Mu- seum, 7 in Dr. Cameron’s collection, and 6 in the United States National Museum (each including the types mentioned above). Remarks.—This species is distinguished by the characters given in the key and by the very feeble characters of the male. The descrip- tion is drawn from a St. Vincent example in the United States National Museum. I find no record of its habits. 418 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 13. PHILONTHUS TRINITATIS, new species Description—Head piceous, pronotum piceorufous, elytra rufous but paler at base, abdomen rufous with fourth and fifth visible seg- ments piceous. Head (including eyes) one-sixth wider than long, suborbicular, hind angles scarcely traceable; eyes at a little less than their length from base; with a strong median longitudinal depression in anterior half; disk with two punctures forming (with two supra- ocular) a nearly straight row of four between the eyes, and with scat- tered very coarse punctures at sides and base; without ground sculp- ture. Pronotwm one-third longer than wide, widest near front angles, feebly emarginately narrowed to basal angles, which are moderately rounded, base truncate; with discal series of five or six punctures (and another anterior pair which is not truly marginal) very deep and coarse, nearly umbilicate, and with five or six lateral punctures; with- out ground sculpture. //ytra with rather coarse punctures separated by less than their diameter; the surface somewhat rugose but without ground sculpture. Abdomen irregularly but finely and submuricately punctate; without ground sculpture. Male, unknown. Female, eighth sternite rounded. Length, 6 mm. Type locality—Trinidad, St. Augustine. Types.—Holotype in the Museum of Comparative Zoology. Records.—The following is the only record known to me: Trinidad: St. Augustine (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species is not very similar to P. waterhouse?. The coloring, the strength of the pronotal punctures, the shape of the head and pronotum, the coarseness of the elytral punctures, and the com- plete lack of any trace of ground sculpture are among the differences. I would hesitate to describe a single Philonthus as new, especially from Trinidad, if it were not so unusual in appearance and structure. None of the South or Central American species known to me are at all similar. ‘The general facies is that of a Belonuchus, but the femora are without spines (except for one row on the anterior) and the gular sutures are united only behind the middle. I have received no record of its habits. 14. PHILONTHUS HAVANIENSIS (Laporte) Staphylinus havaniensis LAports, 1834, p. 116. Philonthus havaniensis (Laporte) Laporte, 1834, p. 116.—Favvet, 1891, p. 115.— BERNHAUER and SCHUBERT, 1914, p. 340.—LeNe and MurcuHter, 1914, p. 406.— Wotcort, 1924, p. 78; 1936, p. 197. Philonthus trepidus ErtcHson, 1840, p. 489.—FAvuver, 1891, p. 115 (as synonym of havaniensis).—BERNHAUER and SoHvusBert, 1914, p. 340. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 419 Philonthus obscurus FAUVEL, 18638, p. 434; 1891, p. 115 (as synonym of havanien- sis) —BERNHAUER and ScHupeRT, 1914, p. 340.—Lene and Mutcutrr, 1914, p. 406. (Not Gravenhorst, 1802.) Philonthus cubensis Breric, MS. Description —Black (frequently more or less rufescent, especially the pronotum) with brownish pubescence on elytra and abdomen, Head subquadrate, very feebly narrowed behind the eyes to rounded basal angles; eyes at almost their length from base; sometimes flattened or impressed in front; with a pair of punctures forming (with two supraocular) a line of four punctures between the eyes; impunctate area of disk smaller than usual; with very fine strigulose ground sculpture, and sometimes greenish lustre. Pronotum one-fourth longer than wide, sides parallel with basal angles broadly rounded; with two discal series of six punctures, the anterior less distinctly part of the series but distinctly not marginal, and with about four lateral punc- tures; with ground sculpture as on head, lustre more coppery when present. /lytra with surface sometimes subrugose so that punctures are not readily visible; they are directed posteriorly and more or less submuricate; shining or with some trace of tuberculate ground sculp- ture. Abdomen with moderately fine and dense punctures either strongly excavated behind or submuricate; with more or less distinct ground sculpture. Male, eighth sternite with a rounded or triangular emargination about one-fifth to one-half as deep as wide, with a narrow smooth border, frequently the border emarginately depressed. Female, eighth sternite rounded or truncate. Length, 4 to 614 mm. Type locality—Cuba. Of trepidus, “Americae ins. St. Johannis et Puerto Rico”; of obscurus, Cuba. Types.—Probably in the Muséum National d’Histoire Naturelle, Paris. Of trepidus, either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin; of obscurus, presumably in the collection of Blanche Rancin, Caén, France. Fecords.—The following are the records known to me: Cuba: (Laporte, 1834; Fauvel, 1863; Leng and Mutchler, 1914), Cayamas (Schwarz, in U.S.N.M.), Matanzas Province (Bierig, in Blackwelder collec- tion), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: Spanish Town (Blackwelder station 377), Black River (Blackwelder station 416), Kingston (Blackwelder station 379). Hispaniola: Hartt, Bayeux (Hoffman, in U.S.N.M.), Port-au-Prince (Audant collection). Puerto Rico: (Erichson, 1840, as trepidus; Leng and Mutchler, 1914; Wolcott, 1924, 1936), Maricao (Blackwelder station 47A), Guanica (Blackwelder station 356), Mayagiiez (Blackwelder station 50B). St. John: (Hrichson, 1840, as trepidus; Leng and Mutchler, 1914). St. Croix: (Blackwelder station 341; H. A. Beatty, in U.S.N.M.). Antigua: (Blackwelder station 287; H. E. Box, as Blackwelder stations 444M, 4440). 420 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen 14 examples in the United States National Museum, 4 from the Museum of Comparative Zoology, 1 in the collection of the Service Technique in Haiti, 1 in my own collec- tion, 10 collected by Dr. H. E. Box in 1987, 4 collected by H. A. Beatty in 1938, and 18 collected by me during 1935-37. Remarks.—This species is very variable in the details of sculpture and punctation as well as in the shape and extent of the male char- acters. It may be distinguished by the number of punctures in the pronotal series and the shape of the pronotum. My specimens were collected from dung, the muddy banks of ponds, the rotting “fruit” of an imported palm, and flying at dusk. The examples from Dr, Box were taken “among decaying forest debris in Inga laurina forest in mesophytic (voleanic) district.” LXIV. Genus BELONUCHUS Nordmann Belonuchus NORDMANN, 1836, p. 129. Trapeziderus MoTSCHULSKY, 1859, p. 77. Musicoderus SHARP, 1885, p. 455. Genotypes.—Staphylinus haemorrhoidalis Fabricius = Belonuchus haemorrhoidalis (Fabricius) (monobasic). (Misspelled haemoroi- dalis by Nordmann.) Of Trapeziderus, T. bicolor Motschulsky monobasic) ; of Afusicoderus, M. cephalotes Sharp (designated here). Diagnosis—Head without dense umbilicate punctures, not mar- gined beneath the eyes; antennae widely separated at front of head, outer segments pubescent from fourth segment; labrum bilobed; palpi elongate and slender, fourth segment of maxillary longer than third, labial similar to maxillary; gular sutures united from before middle, at least in the males; lateral margin of pronotum double, the lines united near coxal articulation; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent; interseg- mental membranes of abdomen with a pattern of irregular rounded sclerotized areas arranged in longitudinal rows; front femora gen- erally finely, and posterior generally coarsely spinose. Remarks.—This genus has always been separated from Philonthus in an unsatisfactory manner. The spinosity of the posterior femore is more constant than that of the anterior femora but yet does not hold throughout. The gular sutures have here been found to be fairly satisfactory for this purpose, since in Philonthus (at least in the West Indies) these sutures are either separate throughout or united well behind the middle. In counting the punctures in the discal pronotal series I have not tried to follow a set rule but have included any marginal puncture that is in the proper position to fall within the series. If the marginal MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 421 puncture is clearly out of line it is not counted. This method was taken into consideration in the key in the case of the species not repre- sented by specimens at hand. All references to this character are corrected to this method. Musicoderus apparently cannot be retained even as a subgenus, at least as far as our one species is concerned. Dr. Cameron expressed to me his opinion that it is a synonym of Belonuchus and I concur in this opinion. The record of Alusicoderus cephalotes Sharp from Jamaica by Gowdey, 1926, is probably a misidentification. Until the specimens can be reexamined, it must be regarded as a doubtful record. Of West Indian specimens I have examined 256, which I assign to 16 species of which 7 are believed to be new. KEY TO WEST INDIAN SPECIES OF BELONUCHUS 1. Elytra bright rufous or testaceous with head and pronotum black ____—~ 2D Elytra black, bluish, or body rufescent throughout____________________ | ZA ADUOMeH ADleOlOred st eLss AIT C VE ss TP ga 2 et EES re ee 3 AD GOMeNT UNTCOLOROUS# 2 2 ieee ST eens feed ee Ee a A 11. mundus 3. Abdomen rufotestaceous, apex black______________________ 14. rufipennis Abdomensplack= apex, rufotestaceousua. 226.2 «seis ase tb 12. satyrus 2 VADeXZOLApdomen abruptly; paler—-2-. 2-2 ea A Oe Te 5 AD AOMENEUNTCOLOLOUS fae ee Ek Ne ney. RANT BATONS STN YMA Teen 10 5. Dorsal series of pronotum with five punctures________________-____-__ 6 Dorsal series of pronotum with less than five punctures_______________ 9 OG: Apical. segmentralone paler es= eS ANAL een eee 5. amplus (iwonorimorerapicalsézmentsy paler2 2:22 227 oe Oi eee ee 7 i. Leadvandepronotumubluishiws 3.455. sven) T__ View bape 6. coelestinus EHeadvandapronotum: black.orjpiceouse.* 2-3. ee ees 8 Srpnen oN aT OT CSS ee eee eng oe ees en ae 9. bugnioni en sche apowbeal Oem se ee en See ee ens See 8. agilis 9. Pronotum with single pair of dorsal punctures__________-____ 7. cognatus Pronotum with dorsal series of three punctures________________._+--__- 18 10; Hront of head: with,a, longitudinal, depression———._-_-—-._.- 2-22. 2 Front of head without longitudinal depression_____________---________ 11 lh roninof head wwilhipbroddsconcavitye—.=—- = ft) — eye 8. trinitatis Front of head entirely without depressions_____________.__ 10. danforthi 12. Dorsal series of pronotum with four punctures______________ 2. oakleyi Dorsal’ series ‘of pronotum! with’ five ‘punctures 222) Vl Ves! theese See 13 13. Abdomen (and sometimes other parts) rufous_______-_-_____-______- 14 Abdomen black, or) piceocastaneouss- 222501 ss _ be Se Me ees eel iee re 15 14. Pronotum distinctly narrowed posteriorly_________-____-______ 16. antiguae Pronotum not distinctly narrowed posteriorly___--__-__-__ 13. dominicus 15. Punctures of elytra generally separated by about their diameter or more. 16 Punctures of elytra separated by much less than their diameter. 1. hispaniolus 148 One specimen in the present collection runs out at this point in the key. It was taken by the Plant Quarantine Service at New York from boats that had come last from Cuba. Inasmuch as there is considerable chance of error in assuming that Cuba is the actual home of the species, it is not named or described here. 422 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 16. Much depressed; elytral interyals with indistinct ground sculpture; length GSD oe a ree ee 15. cacao Not depressed ; elytral intervals smooth, shining; length 6 to 9 mm. 4, gagates 1. BELONUCHUS HISPANIOLUS, new species Description—Piceous throughout. Head narrowed behind the eyes with sides completely rounded into base; eyes at barely their length from base; in front with a median longitudinal impression nearly one-half as long as head; with large and impressed punctures sparsely scattered at sides; with fine strigulose ground sculpture. Pronotum about one-eighth longer than wide, widest near front, feebly emarginately narrowed posteriorly to rounded basal angles; discal series with five punctures, and with about five lateral punc- tures; ground sculpture as on head. lytra with punctures rather small and dense, generally separated by distinctly less than their diameter, the bottom of each appearing granulate; surface even, without ground sculpture. Abdomen with very sparse and fine, often submuricate punctures; without distinct ground sculpture but with iridescent lustre. Afale, unknown. Female, eighth sternite rounded. Length, 7 mm. Type locality —Dominican Republic, San Francisco Mountains. Types.—Holotype and one paratype, females, U.S.N.M. No. 52468, collected on “6.9.05” by August Busck. Records.—The following is the only record to me: Hispaniola: DoMINIcAN Repustic (Busck, in U.S.N.M.). Specimens examined.—I have seen only the two types. Remarks.—This species is distinguished by its dense granulate elytral punctures and by its uniform black color. I find no record of its habits. 2. BELONUCHUS OAKLEYI, new species Desecription—Piceous, head, pronotum, and abdomen with irides- cent luster. Head with sides evenly rounded from eyes to neck; eyes at less than their length from base; in front with a longitudinal im- pression nearly one-half as long as head; with a few moderately large scattered punctures; with fine strigulose ground sculpture. Pronotum one-sixth longer than wide, widest at anterior angles, feebly and sinu- ately narrowed posteriorly, basal angles broadly rounded; discal series of four punctures, with about six lateral punctures; with ground sculp- ture as on head. Elytra with moderate punctures generally separated by about their diameter; intervals flat but surface somewhat ridged and finely coriaceous. Abdomen with moderately sparse and fine sub- muricate or excavated punctures; with only a vague trace of ground sculpture. Male, unknown. Female, eighth sternite rounded. Length, 6 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 423 Type locality.—Puerto Rico, Mayagiiez. Types.—Holotype, U. S. N. M. No. 52460, collected by me on Octo- ber 5, 1935. Records.—The following is the only record known to me: Puerto Rico: Mayagiiez (Blackwelder station 50B). Specimens examined.—I have seen only the unique type. Remarks.—This species is distinguished by having only four punc- tures in the pronotal series. It is also an unusually small and slender species, with a violet luster on head and pronotum. The type was found in a mass of rotting “fruit” at the base of an imported palmnlike tree. 8. BELONUCHUS TRINITATIS, new species Description.—Black, pronotum and abdomen more or less piceous. Head quadrate, sides arcuate but not distinctly narrowed behind eyes, basal angles rounded but marked by ridges; eyes separated from base by about their length; with a concavity in front but without longitu- dinal impressed line; punctures large, impressed, umbilicate, but very sparse; with very fine strigulose ground sculpture. Pronotum scarcely longer than wide, widest at anterior angles, sides straight and feebly narrowed to base which is strongly rounded; dorsal series of five punc- tures, and with about five lateral punctures; ground sculpture as on head. “lytra with moderately coarse punctures generally separated by more than their diameter by somewhat uneven intervals; surface indistinctly coriaceous. Abdomen sparsely and rather finely submuri- cately punctate; with only a trace of transverse ground sculpture. Male, eighth sternite flattened or concave and with a rounded emar- gination about three times as wide as deep. Yemale, with eighth sternite truncate. Length, 8 to 9 mm. Type locality—Trinidad, Department of Agriculture Stock Farm at St. Joseph, 6 miles east of Port of Spain. Types.—Holotype, male, and four paratypes, U.S.N.M. No. 52458, collected by me on January 5, 1936. Records.—The following is the only record known to me: Trinidad: Port of Spain (Blackwelder station 130). Specimens examined.—I have seen only the five types. Remarks.—This species is distinguishable by the lack of median longitudinal impression at the front of the head. The shape of the concavity differs between the sexes. The females have a smaller de- pression, it being more nearly longitudinal but still distinguishable from the linear impression of the other species. The types were found in cattle dung. 424 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 4. BELONUCHUS GAGATES Erichson Belonuchus gagates ErtcHson, 1840, p. 424.—Duvat, 1857, p. 38.—CHEVROLAT and FAuUVEL, 1863, p. 4838.—BERNHAUER and ScHUBERT, 1914, p. 370.—LENG and MutcHter, 1914, p. 406; 1917, p. 199.—Wotcorr, 1924, p. 78.—Gowpey, 1926, p. 12.—WotcortT, 1936, p. 197. Belonuchus minar ErtcHson, 1840, p. 424.—BrrNHAUER and ScHUBERT, 1914, p. 371.—LENG and MuUTCHLER, 1914, p. 406. Musicoderus nigrocaeruleus CAMERON, 1913a, p. 175. Musicoderus nigrocoeruleus LENG and MuTCHLER, 1917, p. 198 (misspelling). Belonuchus gowdeyi BERNHAUER, MS.—Gowpey, 1928, p. 2.—BERNHAUER, 1935, p. 91. Belonuchus nigrocaeruleus (Cameron) SCHEERPELTzZ, 1933, p. 1379. Description.—Black to rufopiceous, often with metallic lustre except on elytra. Head slightly narrowed behind the eyes, and sides rounded into base; eyes at about their length from base; in front with a longi- tudinal impression of varying length; punctures large and umbilicate but very sparse; with fine strigulose ground sculpture. Pronotwm not or scarcely longer than wide, widest in front with front angles rounded, sides nearly straight, moderately narrowed posteriorly to completely rounded base; dorsal series of five punctures, and with five or six lat- eral punctures; with ground sculpture as on head. 7ytra with punc- tures separated by once or twice their diameter by flat even intervals; not distinctly coriaceous. Abdomen very sparsely and moderately finely submuricately or excavately punctate; without distinct ground sculpture but generally somewhat iridescent. fale, eighth sternite with a rounded emargination about three times as wide as deep. Female, eighth sternite rounded. Length, 61% to 9 mm. Type locality— ‘Puerto Rico ..., Cuba...” Of gowdeyi Cin- chona, Jamaica; of minav, St. Thomas and St. John; of nigrocaeru- leus, Newcastle district, Jamaica. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of gowdey?, in the British Museum; of minax, with gagates,; of nigrocaeruleus, in Dr. Cameron’s collection. Records.—The following are the records known to me: Bahamas Island: Nassau (Busck, in U.S.N.M.). Cuba: (Erichson, 1840; DuVal, 1857; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; British Museum), Marianao (Rambousek, in Cameron col- lection), Habana (Rambousek, in Cameron collection; Baker, in Casey col- lection, U.S.N.M.), Cayamas (Schwarz, in U.S.N.M.), Santiago de las Vegas (Dimmock, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Department of Agriculture collection, Jamaica; Gowdey, 1926, 1928; Leng and Mutchler, 1917, as gagates and nigrocaeruleus; Bernhauer, 1935, as gowdeyi; British Museum; Schaus and Hubbard, in U.S.N.M.), Cinchona (British Museum, as gowdeyi), Neweastle (Cameron, 1913, as nigrocueru- leus), Gordon Town (Blackwelder station 383). Hispaniola: (Leng and Mutchler, 1914). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 425 Puerto Rico: (Erichson, 1840; DuVal, 1857; Leng and Mutchler, 1914; Wolcott, 1924, 1936), Bayamon (Busck, in U.S.N.M.), Juana Diaz (Dozier, in U.S.N.M.), Ensenada (Dozier, in U.S.N.M.), Maricao (Morrison, in U.S.N.M.; Black- welder station 47A), Mayagiiez (Blackwelder station 50B; Dozier, in U.S.N.M.), San Juan (Blackwelder station 51), El Yunque (Blackwelder sta- tion 851A), Ponce (Wolcott, 1924). St. Thomas: (Erichson, 1840; Leng and Mutcher, 1914; as mina). St. John: (Erichson, 1840; Leng and Mutchler, 1914; as minar). Montserrat: (Hubbard, in U.S.N.M.). Specimens examined—I have seen 11 examples in the British Museum (including the type of gowdey?), 14 in the collection of Dr. Cameron (including the type of nigrocaeruleus), 2 from the Museum of Comparative Zoology, 66 in the United States National Museum (including 4 in the Casey collection), and 12 collected by me in 1935-87. Remarks.—This species is rather variable in several characters. It seems to be our only widespread all-black species of this genus, but it has been found in numbers only under carrion. Examination of the types of nigrocaeruleus and gowdeyi has shown them to be the same as gagates. This species has not been definitely recorded from new specimens since its description. This species has been collected from dead-meat traps, from “crowns of Tillandsia,” from beneath dung and fresh chips, from the rotting “fruit” at the base of an imported palm, in rotten grapefruit, and from under chips and rubbish on stumps. 5. BELONUCHUS AMPLUS, new species Description.—Piceous, apical segment of abdomen rufotestaceous. Head with sides arcuately narrowed behind eyes, basal angles broadly rounded; eyes separated from base by one-third more than their length; in front with a feeble longitudinal impression extending nearly to middle; with very large and somewhat umbilicate punctures rather impressed, scattered except along middle and in front, dense be- hind eyes; with very fine strigulose ground sculpture. Pronotum nearly one-fourth longer than wide, widest at anterior fifth, feebly sinuately narrowed to rounded basal angles; discal series of five im- pressed punctures, with about six lateral punctures; ground sculpture even finer than on head. //ytra with moderately large but indefinite punctures, more or less united transversely and sometimes submuricate ; without ground sculpture. Abdomen with moderately coarse punc- tures, sometimes excavated behind or submuricate. JMJale, unknown. Female, eighth sternite rounded. Length, 9 mm. Type locality —St. Lucia, one-half mile northwest of St. Remy on road to Soufriére (214 miles southeast of Soufriére). Types.—Holotype and one paratype, females, U.S.N.M. No. 52457, collected by me on April 22, 1936. 449008—42—_28 426 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following is the only record known to me: St. Lucia: (Blackwelder station 221). Specimens examined.—I have seen only the two types. Remarks.—This species is very similar to coelesténus Bernhauer but is distinguished by its greater size and the absence of any bluish color on the head, pronotum, and elytra. The types were collected from rotting cocoa pods. 6. BELONUCHUS COELESTINUS Bernhauer Belonuchus coelestinus BERNHAUER, 1908b, p. 330.—BrRNHAUVER .and SCHUBERT, 1914, p. 8369.—LENG and MutcHrkEr, 1914, p. 406. Belonuchus coeruleus CAMERON, 1922, p. 122.—ScHEERPELTz, 1933, p. 1377. Belonuchus smithi FavveL, MS.—CAMERON, 1922, p. 122. Description.—Head and pronotum shining bluish, elytra cyaneous, abdomen nigropiceous with two apical abdominal segments testaceous. Head quadrate, as long as wide; sides parallel with rounded posterior angles; in front with a median longitudinal impression; with four punctures between the eyes in a rectangle, remainder, except along middle, with large isolated punctures. Pronotum somewhat longer than wide, somewhat narrowed posteriorly; with dorsal series of five strong punctures, and at the sides with a few scattered punctures. Elytra strongly and densely punctate. Abdomen strongly, but not deeply, and somewhat densely punctate, posteriorly more scattered and fine. Male, unknown. Female, (sternites not modified). Length, 6to7mm. (From Bernhauer and Cameron.) Type locality.—St. Vincent (for both). Types—tIn the collections of the K. K. naturhistorischen Hof- museums in Wien and Dr. Bernhauer. The type of coeruleus is in the British Museum. Records.—The following are the records known to me: St. Vincent: (Bernhauer, 1908; Leng and Mutchler, 1914; Cameron, 1922, as coeruleus). Specimens examined.—I have seen only the two examples in the British Museum. Remarks.—This species is known to me only from the two descrip- tions and the two examples in the British Museum. In addition to the correspondence of Dr. Bernhauer’s and Dr, Cameron’s descriptions, the type specimens almost certainly came from the same lot, since both were taken by H. H. Smith on St. Vincent. From my brief examination of the type of coeruleus I can add the following dimensions: Head one-ninth wider than long, pronotum one-eighth longer than wide, elytra exactly as long as wide. ‘This species will be readily distinguishable by its color. I find no record of its habits. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 427 7. BELONUCHUS COGNATUS Sharp Belonuchus cognatus SHARP, 1885, p. 429.—BERNHAUER and ScHusErt, 1914, p. 369.—SCHEERPELTZ, 1933, p. 1877. (Not Fauvel, 1891.) Description.—Piceous black, abdomen piceocastaneous with two apical segments rufotestaceous. Head with sides arcuately expanded behind eyes and rounded into base, but with posterior angles marked by a ridge; eyes separated from base by fully twice their length; in front with a rather feeble impression about one-third as long as head ; punctures large, umbilicate, and impressed but very sparse even at sides; with fine strigulose ground sculpture. Pronotum about one- tenth longer than wide, widest at anterior angles, sinuately narrowed to rounded basal angles; with each discal series reduced to a single puncture, and with only three or four lateral punctures; with ground sculpture a little less distinct than on head. LHlytra with moderate punctures somewhat impressed, generally separated by their diameter or more by uneven intervals; surface indistinctly coriaceous. Abdo- men with moderately sparse submuricate and excavated punctures; with trace of ground sculpture. Male, eighth sternite rounded but very feebly emarginate at apex. Female, eighth sternite rounded and not at all emarginate. Length, 9 to 10 mm. Type locality —Panama, Bugaba. Types.—In the British Museum. Records—The following are the records known to me: St. Vincent: (British Museum). Trinidad: (Bryant, in Cameron collection), ward of Montserrat (Busck, in U.S.N.M.). South America: VENEZUELA (Bernhauer and Schubert, 1914; Cameron collection), ARGENTINA (Scheerpeltz, 1933) ; CoLomBra, VENEZUELA (British Museum). Central America: British HoNDURAS, GUATEMALA, NicARAGUA, PANAMA (Sharp 1885; Bernhauer and Schubert, 1914; British Museum) ; GuATeMALA (Cam- eron collection). Specimens examined.—I have seen three examples from St. Vincent in the British Museum, two from Trinidad in Dr. Cameron’s collec- tion, and two from Trinidad in the United States National Museum. Remarks.—This species is at once distinguished from all others in our fauna by the single puncture in the dorsal “series” of the pronotum. It differs from haemorrhoidalis of South America in having the entire two apical segments pale. I find no record of its habits. 8. BELONUCHUS AGILIS Erichson Belonuchus agilis Ericuson, 1840, p. 423.—DvuvVAt, 1857, p. 37.—CHEvROLAT and Favvet, 1863, pp. 432, 433.—BrrnHAveR and Scuusert, 1914, p. 369.—LEeNa and Murcutrr, 1914, p. 406—BrrnHAuER, 1921, p. 170.—Gownbey, 1926, p. 12. Belonuchus chevrolati FAUVEL, 1863, p. 432.—BERNHAUER and ScHusert, 1914, p. 369.—Lene abd MuTCHIER, 1914, p. 406. 428 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Description.—Piceocastaneous, elytra darker, two apical abdominal segments testaceous. /7ead quadrate, sides arcuate but not narrowed, basal angles rounded but also marked by a ridge; eyes at nearly twice their length from base; very feebly impressed in front, punctures moderately large, impressed, scattered sparsely at sides; with fine strigulose ground sculpture. Pronotuma very little longer than wide, widest at anterior fifth, very strongly narrowed to rounded basal angles; discal series of five punctures, and with about six lateral punc- tures; ground sculpture as on head. 7ytra with moderate punctures generally separated by more than their diameter by flat intervals; surface very finely and indistinctly coriaceous. Abdomen with very sparse punctures excavated behind; with very indistinct ground sculp- ture. J/ale, unknown. Female, eighth sternite rounded. Length, 914 mm. Type locality —Cuba. Of chevrolati, Habana, Cuba. Types.—Presumably in the Zoologische Museum, Berlin. Of chevrolati, presumably in the collection of Blanche Rancin, Caén, France. Records —The following are the records known to me: Cuba: (Erichson, 1840; DuVal, 1857; Fauvel, 1863; Leng and Mutchler, 1914; British Museum), Habana (Fauvel, 18638, as chevrolati), Cayamas (Schwarz, in U.S.N.M.). Jamaica: (Bernhauer and Schubert, 1914; Leng and Mutchler, 1914; Gowdey, 1926; British Museum). [Hispaniola: (Bernhauer and Schubert, 1914.) ] [South America: CoLtomBiA (Bernhauer and Schubert, 1914).] Specimens examined.—I have seen six examples in the British Museum and one in the United States National Museum. Remarks.—This species is distinguished by its coloration, the feeble impressions of the head, and the very sparse punctures of the abdomen. Its most striking feature, however, is the shape of the pronotum, which is so strongly narrowed posteriorly. The specimen before me was taken “on ceiba” (silk-cotton tree). The record of South America by Bernhauer and Schubert is prob- ably an error, since they do not give the type locality. 9. BELONUCHUS BUGNIONI Fauvel Belonuchus bugnioni Fauvet, 1901, p. 86, pl. 2, fig. 183—BrrNHAUER and ScHUBERT, 1914, p. 869.—LEeNG and MuTcHier, 1914, p. 406. Belonuchus exrimius BERNHAUER, 1917, p. 105.—ScHEERPELTzZ, 1933, p. 1378. Description.—Black, elytra bluish, last two abdominal segments and margin of third rufotestaceous. //ead quadrate, coarsely and sparsely punctured at sides; in front with a median longitudinal im- pression. Pronotum one-third longer than wide, posteriorly sinuately narrowed; dorsal series of five punctures, with a few lateral punc- MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 429 tures. H/ytra densely and rather finely punctate. Abdomen sparsely punctate, especially on eighth tergite. Male, unknown. Female, (sternites not modified). Length, 6 to 7 mm. (From Fauvel and Bernhauer.) Type locality —Martinique. Of eximius, Guadeloupe. Types.—Presumably in the collection of Blanche Rancin, Caén, France. Of eviméus, presumably in the collection of Dr. Bernhauer. Records.—The following are the records known to me: Guadeloupe: (Bernhauer, 1917, as eximius). Martinique: (Fauvel, 1901; Leng and Mutchler, 1914). Specimens examined.—I have seen no examples of this species. Remarks.—There seems to be little doubt that these two names represent a single species. It is distinguished by the blue color of the elytra alone and by the extent of the pale area at the apex of the abdomen. I find no record of its habits. 10. BELONUCHUS DANFORTHI, new species Description—Head black, pronotum and abdomen _ picescent, elytra distinctly rufescent. Head rather strongly narrowed behind the eyes, the basal angles obliterated but densely and coarsely pubes- cent; eyes at one-half more than their length from base; without any depressions on front of vertex; with moderate punctures very sparse, absent on center of vertex; without ground sculpture. Pvro- notum over one-fifth longer than wide, widest at anterior third, feebly sinuately narrowed posteriorly, with all angles rounded; dorsal series of five punctures, and with about seven lateral punc- tures; with very fine strigulose ground sculpture. H7/ytra with mod- erately coarse punctures separated by less than their diameter by feebly convex intervals; with only a trace of ground sculpture. Abdomen rather finely and sparsely submuricately punctate; with rather indistinct transversely strigulose ground sculpture. Male, eighth sternite with an abrupt rounded median posterior lobe nearly as long as wide and about one-fifth as wide as apex of segment. Female, unknown. Length, 514% mm. Type locality —Puerto Rico, about 13 miles east of Maricao and north of Yauco, elevation about 3,000 feet. Types.—Holotype, male, U.S.N.M. No. 52459, collected by me on October 3, 1935. Records.—The following is the only record known to me: Puerto Rico: Maricao (Blackwelder station 47B). Specimens examined.—I have seen only the unique type. Remarks.—This is one of those species which is near the dividing line between Belonuchus and Philonthus. It has the united gular 43 BULLETIN 182, UNITED STATES NATIONAL MUSEUM sutures of the former but the posterior femora are unarmed. The anterior femora are spinose beneath. It keys to ¢trinitatis but can be immediately distinguished by its smaller size. The unimpressed front of the head and the male characters serve to separate it from all other West Indian species of either Belonuchus or Philonthus. The type was found in rotting guava fruit. 11. BELONUCHUS MUNDUS Erichson Belonuchus mundus Ericson, 1840, p. 425.—Sotsxky, 1869, p. 263.—BERNHAUEB and ScHUBERT, 1914, p. 371. Description.—Black, elytra rufotestaceous, abdomen sometimes pi- cescent. Head slightly narrowed behind eyes, which are about their length from base; anterior half with a median longitudinal impressed line; punctures coarse and impressed, absent on vertex and sparse even behind eyes; very finely and indistinctly strigulose. Pronotwm about one-eighth longer than wide, very slightly wider in front, basal angles completely rounded; dorsal series of five (or sometimes six) punctures, and with about six lateral punctures; ground sculpture es on head. Hlytra with moderately coarse but uneven punctures separated by their diameter or less; intervals smooth and flat. Ad- domen with moderately coarse and dense punctures, sometimes sub- muricate; surface iridescent but without visible ground sculpture. Male, eighth sternite with a rounded triangular emargination twice as wide as deep. Female, eighth sternite rounded. Length, 6 to 7 mm. Type locality—Americae meridionalis ins. St. Vincenti. .”. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: St. Lucia: (Blackwelder stations 221, 226, 227, 230, 231). St. Vincent: (Erichson, 1840; British Museum; U.S.N.M.). Grenada: (British Museum; Blackwelder stations 132, 153A). [South America: (Bernhauer and Schubert, 1914).] Specimens examined.—I have seen 12 examples in the British Museum, 59 collected by me during 1935-37, and 3 in the United States National Museum. The last were from the original series, received in exchange. Remarks.—This species is distinguished from all other West In- dian species by having the elytra alone red. The Bernhauer and Schubert record of South America is almost certainly an error, since they do not mention the type locality. My specimens were all taken in freshly cut or rotting cocoa pods. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 431 12. BELONUCHUS SATYRUS Erichson Belonuchus satyrus ERicHson, 1840, p. 421.—BrerRNHAUER and ScHuBERT, 1914, p. 371. Description.—Iridescent black, elytra and apex of abdomen rufo- testaceous. Head straight at sides, with basal angles broadly rounded; eyes at about their length from base; in front with a feeble longitudinal impression extending nearly to middle; with only the usual scattered large punctures; with very fine strigulose ground sculpture. Pronotwm scarcely wider than long, widest at anterior angles, feebly narrowed posteriorly, basal angles completely rounded ; discal series of five punctures, and with about five lateral punctures; ground sculpture as on head. Eytra with moderate punctures, fre- quently separated by less than their diameter and sometimes indis- tinctly transversely serial; without ground sculpture. Abdomen with moderately sparse submuricate punctures; without distinct ground sculpture although vaguely alutaceous. J/ale, supraantennal prominences strongly elevated; eighth sternite broadly and very feebly emarginate. Female, supraantennal prominences not ele- vated; eighth sternite rounded. Length, 10 to 12 mm. Type locality—Colombia. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. fecords.—The following are the records known to me: Trinidad: (Birch and Bryant, in British Museum; Busck, in U.S.N.M.), Man- zanilla (Blackwelder stations 103A, 104A), Moruga (Blackwelder station 111), Sangre Grande (Blackwelder station 100A). South America: CotomsBra (Erichson, 1840; British Museum), CoLompra, VENE- ZUELA, NEw GRANADA (Bernhauer and Schubert, 1914). Specimens examined.—I have seen 2 examples in the British Mu- seum, 1 in the United States National Museum, and 10 collected by me in 1935-87. Remarks.—This species is readily distinguished by its size and color, as well as the development of the antennal ridges in the male. My specimens were all taken from cocoa pods (both freshly cut and well decayed). 13. BELONUCHUS DOMINICUS, new species Description.—Head piceous, pronotum piceocastaneous, elytra rufo- castaneous, abdomen rufous. Head feebly narrowed behind the eyes, sides rounded into base; eyes at a little more than their length from base; in front with a longitudinal impression half as long as head; with rather coarse and impressed punctures scattered at sides; with fine strigulose ground sculpture. Pronotum about one-tenth longer than 432 BULLETIN 182, UNITED STATES NATIONAL MUSEUM wide, with sides nearly parallel, base broadly rounded; dorsal series of five punctures, and with about seven lateral punctures; ground sculp- ture ason head. /ytra with rather coarse and dense punctures, some- times not sharply outlined and in transverse rows; without ground sculpture. Abdomen with sparse and moderately coarse punctures, generally excavated behind; without distinct ground sculpture, but somewhat iridescent. J/a/e, eighth sternite with an abrupt rounded emargination twice as wide as deep. Female, eighth sternite rounded. Length, 8 to 9 mm. Type locality—Dominica, Hillsboro Estate, at mouth of the Layou River, Leeward side. Types.—Holotype, male, and four paratypes, U.S.N.M. No. 52462, collected by me on June 2, 1936. One paratype in my own collection. Records.—The following is the only record known to me: Dominica: (Blackwelder station 237). Specimens examined,—I have seen only the six types. Remarks.—This species is distinct by its rufescent color, by its male characters, and by the parallel pronotum. The six specimens are very uniform in these respects and seem to preclude the possibility of the color being due to immaturity. The types were found in rotting cocoa pods. 14. BELONUCHUS RUFIPENNIS (Fabricius) Staphylinus rufipennis Fasrictus, 1801, p. 597.—EricHson, 1840, p. 426.— SHARP, 1885, p. 431. (Not Stephens, 1829; not Solier, 1849.) Staphylinus formosus GRAVENHORST, 1806, p. 72.—ErRIcHSON, 1840, p. 426.— SHARP, 1876, p. 160; 1885, p. 431. Cafius apicalis DEJEAN, 1838, p. 68.—ERICHSON, 1840, p. 426. Belonuchus formosus (Gravenhorst) ErrcHson, 1840, p. 426.—GrEMMINGER and Harotp, 1868, p. 598 (as synonym of rufipennis)—SHarP, 1885, p. 481.— BLATCHLEY, 1910, p. 379.—BERNHAUER and SCHUBERT, 1914, p. 370.—LENe@, 1920, p. 108.—ScHEERPELTZ, 1933, p. 1878. Philonthus rufipennis (Fabricius) ErtcHson, 1840, p. 426. (Not Grayenhorst, 1802; not Solier, 1849.) Belonuchus pallipes MreLSHEIMER, 1846, p. 35.—LAcorDAIRE, 1854, p. 80.—GrEM- MINGER and HAROLD, 1868, p. 598.—BERNHAUER and SCHUBERT, 1914, p. 870,— LENG, 1920, p. 108. Belonuchus rufipennis (Fabricius) GrMMINGER and Harorp, 1868, p. 598 (as valid). —SuHarp, 1885, p. 431.—BrerNHAUER and ScHUBERT, 1914, p. 370. Belonuchus apicalis (Dejean) GEMMINGER and HARoLD, 1868, p. 598. Description.—Black, elytra rufotestaceous, four basal tergites of abdomen rufous. /Zead rounded posteriorly from the eyes into base; eyes separated from base by about their length; in front with a longi- tudinal impression about one-third as long as head; with only a few scattered punctures, none along middle; with very fine strigulose ground sculpture. Pronotum scarcely longer than wide, scarcely MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 433 at all narrowed posteriorly, basal angles completely rounded; with ground sculpture as on head. “lytra with moderately coarse punc- tures separated by their diameter or more, laterally with less flat intervals; without ground sculpture. Abdomen with sparse and moderately fine submuricate punctures, without distinct ground sculpture but vaguely alutaceous. Male, eighth sternite with a feeble triangular emargination three times as wide as deep. Female, eighth sternite rounded. Length, 64% to 74% mm. Type locality—*Americae meridionali.” Of formosus, Carolina; of apicalis, Cayenne; of pallipes, Pennsylvania. Types.—Disposition not known to me. Of formosus, presumably in the Zoological University Museum, Breslau; of apicalis, not known to me; of pallipes, presumably in the Museum of Compara- tive Zoology, Cambridge, Mass. Records—The following are the records known to me: Tobago: (Blackwelder station 116B). Trinidad: (Bryant, in Cameron collection), Moruga (Blackwelder station 111), Manzanilla (Blackwelder station 103A), Port of Spain (airchild, in M.C.Z.) Mount Tucuche (Darlington, in M.C.Z. and U.S.N.M.). South American: (Fabricius, 1801), CotomMpra, CAYENNE (FErichson, 1840), CotomBIA, Brazit (Sharp, 1876, 1885), ARGENTINA (Scheerpeltz, 1933). Central America: Mexico, BRITISH HONDURAS, GUATEMALA, NICARAGUA, PANAMA (Sharp, 1885). North America: FLoripa, Georgia, CAROLINA, NEw JERSEY, NEW YORK, PENNSYL- VANIA, OHIO, QUEBEC, INDIANA, NEw Mexico, Arizona (Gravenhorst, 1806; Erichson, 1840; Melsheimer, 1846; Sharp, 1885; Leng, 1920; Blatchley, 1910; ete.). Specimens examined—From the West Indies I have seen 3 examples in Dr. Cameron’s collection, 8 from the Museum of Com- parative Zoology, and 14 collected by me in December 1935. Numer- ous examples from North and South America were also examined. Remarks.—\ regret the necessity for changing an old and well- established name but can find no other tenable procedure. The Gem- minger and Harold catalog puts the names formosus and rufipennis in their proper relationship, but subsequent writers have not followed that work in this respect. The Bernhauer and Schubert catalog, although recognizing that the two are equal, places the older name as a synonym of the younger. Unless formosus be held to be a dis- tinct species, it cannot be retained unless ruficollis is a homonym. Such a previous use of the name I am not able to find, and I there- fore use ruficollis as the valid name. This species is distinguished by its red abdomen with black apex, which is a combination unique among West Indian representatives of this genus. My specimens were all taken from freshly cut or rotten cocoa pods, 434 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 15. BELONUCHUS CACAO, new species Description —Head rufopiceous, pronotum rufous, elytra and abdo- men rufocastaneous, the tip of latter rufescent but not strongly set off from rest. Head quadrate, posterior angles marked by ridges, eyes at one-half more than their length from base; in front with a strong median longitudinal impression, about half as long as head; with large impressed punctures scattered not very sparsely at sides; with fine strigulose ground sculpture. Pronotwm about one-sixth longer than wide, widest at anterior angles and arcuately narrowed to rounded basal angles; discal series with five punctures, and with about seven lateral punctures. Hlytra with moderate punctures sepa- rated by about their diameter but often indistinct and with intervals forming indefinite transverse ridges; surface finely coriaceous. Abdomen with fine and not very sparse punctures, generally sub- muricate; with very fine strigulose ground sculpture. Male, un- known. Yemale, eighth sternite truncate. Length, 544 mm. Type locality —Trinidad, one-half mile south-southwest of Man- zanilla junction on the Plum Road. Types.—Holotype and paratype, females, U.S.N.M. No. 52464, col- lected by me on December 18, 1935. The paratype was taken at Sans Souci on December 15, 1935. Records.—The following are the records known to me: Trinidad: Manzanilla (Blackwelder station 102), Sans Souci (Blackwelder station 98B). Specimens examined.—l\ have seen only the two types. Remarks.—This species is distinguished by its very much de- pressed form, by its quadrate head, and by the arcuate sides of the pronotum. The types were taken in rotting cocoa pods. 16. BELONUCHUS ANTIGUAE, new species Description —Rufotestaceous throughout. Head not narrowed be- hind eyes but broadly rounded from eyes to neck; eyes separated from base by a little more than their length; in front with a longitudinal median impression about one-third as long as head; with rather large impressed punctures scattered sparsely at sides; with very fine strigu- lose ground sculpture. Pronotwm about as wide as long, widest in front, sinuately narrowed posteriorly to rounded basal angles; dorsal series with five punctures, and with about six lateral puctures; with ground sculpture as on head. //ytra with moderate punctures fre- quently individually indistinct because of uneven surface, often ar- ranged in transverse rows; without ground sculpture. Abdomen moderately sparsely and finely punctate, submuricately or excavately ; without distinct ground sculpture but somewhat iridescent. Male, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 435 eighth sternite with a feeble triangular emargination three times as wide asdeep. /emale, eighth sternite rounded. Length, 6 to 714 mm. Type locality— Antigua, near summit of Boggy Peak, parish of St. Mary, elevation 1,300 feet. Types.—Holotype, male, and five paratypes, U.S.N.M. No. 52461, collected by Dr. H. E. Box on May 30, 1937. One paratype has been deposited in the British Museum and one in Dr. Cameron’s collection. Records.—The following is the only record known to me: Antigua: (Box, as Blackwelder station 444Q). Specimens examined.—I have seen only the nine types. Remarks.—This species is very distinct by its coloring, and it is of such uniform color throughout these nine examples that I cannot ascribe it to immaturity. The types were collected “from inside of scarlet bracts of a large unknown feliconia sp. (one of the rarest plants of Antigua) in dense mesophytic woodlands.” LXV. Genus CAFIUS Stephens Cafius STEPHENS, 1829, p. 288. (Not Boisduval and Lacordaire, 1835.) Subgenus Remus HowMe, 1837, p. 64. (Not Thomson, 1860; not Mulsant and Rey, 1875.) Pseudidus MULSANT and Rey, 1875, p. 574. Subgenus Bryonomus CASEY, 1885, p. 313. Subgenus Huremus Brera, 1934a, p. 68. Subgenus Philonthopsis Kocu, 1936, p. 178. Subgenus Pseudoremus Kocu, 1936, p. 179. Genotypes.—Staphylinus fucicola Samouelle = Cafius fucicola (Samouelle) (designated by Lacordaire, 1854). Of Remus, R. seric- eus Holme (monobasic); of Pseudidus, Remus sericeus Holme= Pseudidus sericeus (Holme) (designated by Tottenham, 1939); of Bryonomus, Cafius canescens Miklin= bryonomus canescens (Miiklin) (designated here) ; of Philonthopsis, Cafius sabulosus Fauvel=Cafius (Philonthopsis) sabulosus Fauvel (designated here) ; of Pseudoremus, Cafius lithocharinus LeConte=Cafius (Pseudoremus) lithocharinus LeConte (designated here) ; of Euremus, Philonthus bistriatus Erich- son=Cafius (Huremus) bistriatus (Erichson) (designated here). Diagnosis—Head and pronotum without dense umbilicate punc- tures, generally margined beneath the eyes; antennae widely separated at front of head, outer segments densely pubescent from the fourth joint; labrum bilobed; palpi moderately stout, fourth segment of maxillary much longer than third, fusiform, labial palpus similar to maxillary; gular sutures united basally ; pronotum with lateral margin double, the lines united near the coxal articulation; anterior coxal 436 BULLETIN 182, UNITED STATES NATIONAL MUSEUM cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular,” sometimes with a small expansion over the base of femur; first and second abdominal sternites absent ; intersegmental membranes of abdomen marked with a pattern of irregular rounded sclerotized areas arranged in longitudinal bands; femora unarmed. Remarks.—As far as I can determine this genus has always been ascribed to Stephens (1832) who was the first to give a diagnosis of it. It was, however, validated by Mannerheim in 1830 and by Stephens in 1829 by the listing of several valid species, and may have been used even earlier. The genus is readily recognized by its habitus as well as by its littoral habits. Koch (1936) has partially revised the genus, although a false im- pression of completeness is given by his paper, which entirely omits mention of over 15 described species or names. It should be pointed out also that there is apparently an error in the key at couplet 25 and 30. The subgenus Remus apparently can be distinguished from only part of the subgenus Pseudoremus. I have examined 171 examples from the West Indies and assign them to three species, all previously described. KEY TO WEST INDIAN SPECIES OF CAFIUS 1. Head with deeply impressed median line anteriorly_-~-----~- 2. caribeanus Head without impressed median line anteriorly_------------------------- os 2. Pronotum with series of punctures along sides of median space. 3. bistriatus Pronotum without single series of punctures along sides of median space. 1. subtilis 1. CAFIUS SUBTILIS Cameron Cafius subtilis CAMERON, 1922, p. 121.—ScHEERPELTZ, 1933, p. 1370.—Brrrie, 1934a, pp. 65, 66. Cafius sericeus var, subtilis Cameron, Kocu, 1936, p. 192. Description—Black, pronotum sometimes picescent, elytra and abdomen castaneous. Head quadrate, not enlarged behind eyes but with basal angles moderately rounded; eyes separated from base by about one-half more than their length; without any longitudinal im- pressions on vertex; with moderately coarse and somewhat umbilicate punctures (not impressed) scattered rather densely except along middle; with fine and dense scaly ground sculpture. Pronotwm over one-fifth longer than wide, widest in anterior fifth, thence scarcely at all narrowed to narrowly rounded posterior angles; disk with median. band impunctate but not outlined by punctures that are distinctly serial; punctures at sides moderately densely and evenly scattered with scaly ground sculpture more distinct than on head. #7ytra with- out punctures but with dense tuberculate sculpture with minute and LS — °°} MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 437 indistinct ground sculpture intermixed. Abdomen finely and densely submuricately punctate; with dense but not very distinct ground sculp- ture. Male, eighth sternite with triangular emargination as deep as wide, with inner edges beveled and outer angles rounded. Female, eighth sternite rounded. Length, 314 to 4144 mm. Type locality —Jamaica, Port Royal. Types.—In Dr. Cameron’s collection. Records.—The following are the records known to me: Cuba: (Koch, 1936), Playa de Marianao (Bierig, 19384; U.S.N.M.). Jamaica: (Bierig, 1984; Koch, 1936), Port Royal (Cameron, 1922). Puerto Rico: San Juan (Blackwelder station 438A). St. Croix: (Blackwelder stations 326, 329, 344). St. Kitts: (Blackwelder station 307). Antigua: (Blackwelder station 276). Montserrat: (Blackwelder station 269). Guadeloupe: Basse Terre (Blackwelder station 77A). Dominica: (Blackwelder station 2401%B). Specimens examined.—\ have seen 13 examples in Dr, Cameron’s collection and 22 collected by me during 1935-37. Remarks.—This species is very similar to sericeus Holme, aqguayot Bierig, and pruinosus Erichson. Whether these represent species or varieties (subspecies) I am not in a position to decide. The length of the antennae, the punctation of the head, and the anterior angles of the pronotum are so variable in both the West Indian examples and in sericeus from the Eastern Hemisphere as well as aguayoi from eastern North America that they will not serve to distinguish them. For the present it is best to retain subtilis as a valid species, at least until adequate series of all these species can be studied. This species be- longs in the subgenus Remus Holme. I have found it under seaweed and drift on sandy beaches. 2. CAFIUS CARIBEANUS Bierig Oaflus corallicola var. caribeanus Birrta, 1934a, pp. 67, 68, fig. 2.—KocH, 1936, p. 175, 176, 186. Description.—Black, the abdomen more or less castaneous. Head straight or slightly rotund behind the eyes with the angles rounded ; the eyes separated from base by about their length; with a fine im- pressed median line from between the eyes to front margin; with coarse, somewhat impressed and umbilicate punctures scattered rather densely except along middle and in front; with distinct strigulose ground sculpture. Pronotwm about one-fifth longer than wide, widest at anterior third, feebly narrowed to basal third and thence straight to prominent basal angles; with smooth midline marked by two con- fused rows of moderate punctures, and with rather densely but ir- regularly scattered punctures laterally with more or less smooth band 438 BULLETIN 182, UNITED STATES NATIONAL MUSEUM next to the series; with strigulose ground sculpture more distinct than on head. lytra without distinct punctures but densely sculptured, apparently with two sizes of asperities. Abdomen with fine and mod- erately dense submuricate punctures; with indistinct ground sculpture. Male, eighth sternite with a triangular emargination about as deep as wide with outer angles rounded. Female, eighth sternite rounded. Length, 5 to 6 mm. Type locality.—Cuba, Playa de Marianao. Types.—In collection of Alexander Bierig. One “cotype” (para- type) is deposited in the United States National Museum (No. 52745), Records——The following are the records known to me: Cuba: Playa de Marianao (Bierig, 1934; United States National Museum). Jamaica: Bluefields Bay (Blackwelder station 418). Puerto Rico: San Juan (Blackwelder station 483A), Humacao (Blackwelder stations 55, 56). St. Croix: (Blackwelder stations 326, 336). Antigua: (Blackwelder station 272). Guadeloupe: Basse Terre (Blackwelder station 77A). Dominica: (Blackwelder station 2401%4B). Grenadines: Carriacou (Blackwelder station 143A). Grenada: (Blackwelder station 163). Specimens examined.—I have seen a cotype received from Sr. Bierig and 20 examples collected by me during 1935-37. Remarks.—I have compared this species with examples of coralli- cola from Singapore (identified by Dr. Cameron) and find it to be very similar, if not indistinguishable. However, since I have only one Cuban example and two Singapore ones I do not feel in a posi- tion to decide the variability of the characters that present the slight differences. Until the species in this group can be studied with ade- quate material (and such has not been available to anyone heretofore, as far as West Indian examples are concerned), it seems best to re- tain caribeanus as a valid species. I have found it under seaweed and drift on sandy beaches. 3. CAFIUS BISTRIATUS (Erichson) Philonthus bistriatus Ericuson, 1840, p. 502.—LEContTE, 1863, pp. 39, 40.— ScHWARZ, 1878, p. 441. Philonthus bilineatus ErtcHson, 1840, p. 503.—BrrNHAUER and ScHUBERT, 1914, p. 380.—Lrena@ and MuTcHLIER, 1914, p. 406.—ScHEERPELTz, 1933, p. 13869. Cafius bistriatus (Erichson) Horn, 1884, p. 237—WickHAM, 1895, pp. 293, 294.—SCHEERPELTZ, 1933, p. 1869.—Kocn, 1936, pp. 175, 187. Cafius vistriatus HAMILTON, 1894, p. 13 (misspelling). Cafius bilineatus (Erichson) SCHEERPELTZ, 1933, p. 1869.—KocHu, 1936, pp. 176, 187. Cafius rufifrons Birra, 1934b, pp. 67, 68.—Kocu, 1986, pp. 176, 187. Cafius rufrifrons Bierta, 1934b, p. 67, fig. 3 (misspelling). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 439 Description—Head black, rest rufocastaneous with pronotum sometimes feebly picescent. Head not enlarged behind eyes but with basal angles moderately rounded; eyes separated from base by one-third more than their length; without any longitudinal im- pression on the vertex; with moderately coarse umbilicate punctures (not impressed) scattered not very densely but absent along center and in front; with fine but very distinct and dense scaly ground sculpture. Pronotum scarcely at all longer than wide, widest at anterior angles, sinuately narrowed to posterior angles, which are rounded but discernible; with impunctate midline marked by two fairly regular series of large and somewhat umbilicate punctures, with rather dense punctures laterally separated by an irregular smooth space from the series; with ground sculpture less distinct than on head. Llytra with dense punctures obscured by irregular and indefinite ground sculpture. Abdomen with fine but not very dense submuricate punctures; with indistinct ground sculpture espe- clally apically. Afale, eighth sternite with a moderate rounded emargination wider than deep and bordered with a translucent membrane. Female, eighth sternite rounded. Length, 5 to 7 mm. Type locality—New York, Long Island. Of bélineatus, St. John; of rufifrons, Cuba, Playa de Marianao. Types.—Presumably in the Zoologische Museum, Berlin. Of bilineatus, either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin; of rufifrons, holotype and allotype in Bierig col- lection, and a “cotype” in the United States National Museum (No. 52746). ftecords.—The following are the records known to me: Bahamas: Water Cay, Egg Island (Wickham, 1895), Egg Island, Eleuthera Island, Harbor Island (Wickham, in U.S.N.M..) Cuba: Playa de Marianao (Bierig, 1934, as rufifrons). Jamaica: Port Royal (Cameron collection; Koch, 1986), Montego Bay (Black- welder station 407), Bluefields Bay (Blackwelder station 418). Mona Island: (Hoffman, in U.S.N.M.). Puerto Rico: San Juan (Blackwelder station 438A). St. Thomas: (Koch, 1936). St. John: (Hrichson, 1840, as bilineatus). St. Croix: (Blackwelder station 326). St. Kitts: (Blackwelder station 307). Antigua: (Koch, 1986; Blackwelder station 276). Montserrat: (Blackwelder station 268). Guadeloupe: (Blackwelder station 77A). Dominica: (Blackwelder station 24014B). St. Lucia: (Blackwelder station 232). Barbados: (Blackwelder stations 190, 192, 197). Grenadines: Carriacou (Blackwelder station 143A). Grenada: (Blackwelder station 163). Tobago: (Blackwelder station 120). 440 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Trinidad: Balandra Bay (Blackwelder station 97). South America: (Koch, 1986; Bernhauer and Schubert, 1914, as bilineatus). North America: (Hrichon, 1840; Horn, 1884; Koch, 1936), FLoriwa, VIRGINIA, MASSACHUSETTS, NEW JERSEY, NEw YORK, RHODE ISLAND, QUEBEC (U.S.N.M.; ete.). Specimens examined.—I have seen 7 examples in Dr. Cameron’s col- lection, 12 in the United States National Museum, and 96 collected by me during 1935-37. Remarks.—This species evidently ranges from the Massachusetts coast to South America by way of the West Indies but has not been recorded from the Gulf of Mexico or from Central America. West Indian examples seem to be identical with North American ones. This is the only species of Staphylinidae so far recorded from Mona Island. It belongs in the subgenus Huremus Bierig. The descrip- tion was drawn from a series from Puerto Rico. The example in the United States National Museum from Miami, Florida (from Bierig collection, as rufifrons), differs decidedly from the series herein identified as bistriatus in the extent of the ground sculpture of the head and pronotum. This specimen is apparently immature, but if it is really the rufifrons of Bierig, then that name may represent a valid species. On the basis of the West Indian ex- amples and the original description I believe that rufifrons is the same as bistriatus. I have found it under seaweed and drift on sandy beaches. LXVI. Genus ERICHSONIUS Fauvel Bisnius THomMson, 1860, p. 168. (Not Stephens, 1829.) Drichsonius Fauver, 1872, p. 427. (Not Schubert, 1911.) Actobius FAUVEL, 1876, p. 72. Neobisnius GANGLBAUER, 1895, p. 464 (new name for Hrichsonius). Genotypes.—Staphylinus cinerascens Gravenhorst= F’richsoniv. cinerascens (Gravenhorst) (designated by Tottenham, 1939). Of Actobius, Staphylinus cinerascens Gravenhorst=Actobius cineras- cens (Gravenhorst) (Rules, Article 30, II, f) ; of Veobisnius, Staphy- linus cinerascens Gravenhorst=Neobisnius cinerascens (Graven- horst) (Rules, Article 30, IT, /). Diagnosis —Head and pronotum without dense umbilicate pune- tures, not margined beneath the eyes; antennae widely separated at the front of head, outer segments densely pubescent from the fourth joint; labrum bilobed; palpi moderately stout, fourth segment of maxillary longer than third, subulate, labial similar to maxillary; gular sutures very approximate posteriorly but not united; lateral margin of pronotum double, the lines united near the coxal articula- tion; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular”; first and second MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 44] abdominal sternites absent; intersegmental membranes of abdomen with a pattern of irregular rounded sclerotized areas arranged in longitudinal bands; femora unarmed. Remarks.—The name Neobisnius was proposed by Ganglbauer ap- parently under the impression that H7ichsonius was preoccupied. However, the two previous uses (Dana, 1849, and Westwood, 1849) both use the spelling Z’vichsonia, which does not preoccupy F’rich- sonius under our present code of nomenclatural procedure. It there- fore becomes necessary to restore H’richsonius as the valid name of this genus. Seven species of this genus have been described from the West Indies, and I have made every effort to recognize them. I have seen. type material of five of these species and have material in con- siderable series from Cuba to Grenada. These I am unable to separate into more than one species. This one may very likely be found on some part of the mainland of America as well, but I have not been able to recognize it among the species available to me. 1. ERICHSONIUS HUMILIS (Erichson) Philonthus humilis ErtcHson, 1840, p. 512.—CHeEvrotat and FAvvEL, 1863, p. 435,—Lrene and MuTcHLER, 1914, p. 406.—WotcoTt, 1924, p. 78; 1936, p. 197. Neobisnius humilis (Hrichson) BERNHAUER and SCHUBERT, 1914, p. 323.—LENG and Murcuter, 1914, p. 406.—Breria, 1933, pp. 50, 54. Neobisnius margipallens BERNHAUER and ScHUBERT, 1914, p. 323. (Not Lath- robium margipallens DUVAL, 1857, p. 41.) Neobisnius limbatus CAMERON, 1922, p. 118.—ScHrERPELTZ, 1933, p. 1327. Neobisnius funerulus CAMERON, 1922, p. 119.—ScHEERPELTZ, 1933, p. 1827. Neobisnius nigrocoeruleus CAMERON, 1922, p. 118. Neobisnius nigrocaeruleus SCHEERPELTZ, 1938, p. 13827 (misspelling). Neobisnius carbonarius Breric, 1933, p. 49. Neobisnius demmeli Brieric, 1933, p. 54. Description.—Piceous, the pronotum sometimes rufescent, narrow apex of elytra and abdominal segments generally pale. Head sub- quadrate with angles broadly rounded, seldom visibly narrowed be- hind eyes; eyes separated from base by a little more than their length; with moderate punctures of even size, generally separated by less than their diameter but rather indistinct posteriorly; sometimes with very indistinct ground sculpture. Pronotwm over one-fifth longer than wide, widest in anterior fourth and thence emarginately narrowed to rounded base; smooth midline narrow, not outlined by punctures in series; punctures similar to those on head but generally denser, with- out large smooth areas but punctures indistinct at sides; sometimes with very indistinct ground sculpture. /lytra with fine and rather dense submuricate punctures, not distinct if viewed from behind. Abdomen very finely and moderately densely submuricately punctate, with indistinct ground sculpture. Male, eighth sternite with a small 449008—42—__99 442 BULLETIN 182, UNITED STATES NATIONAL MUSEUM semicircular emargination. J emale, eighth sternite broadly rounded. Length, 3 to 5 mm. Type locality.— Puerto Rico. Of limbatus, Haiti; of nigro- coeruleus, Jamaica (although Dr. Cameron recorded this species only from Haiti, the example labeled type is from Jamaica) ; of funerulus, St. Vincent; of demmeli, Cuba, province of Pinar del Rio, Sierra del Rosario, vicinity of Cayajabos; of carbonarius, Cuba, province of Pinar del Rio, Sierra del Rosario, Cayajabos. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of limbatus and nigrocoeruleus, in Cameron col- lection; of funeru/us, in the British Museum; of carbonarius and demmeli, in collection of Alexander Bierig (one “cotype” of the latter in the United States National Museum (No. 52744). Records.—The following are the records known to me: Cuba: (Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; Cameron collec- tion, as nigrocoeruleus and hwmilis), Cayajabos (Bierig, 1933, as dem- meli and carbvbonarius; U.S.N.M.), Almendares (Bierig, in U.S.N.M.), Cay- amas (Schwarz, in U.S.N.M.), Aspiro (Bierig, in U.S.N.M., as carbonarius), Sierra del Rosario (Bierig, 1933, as carbonarius and demmeli), Soledad (Darlington, in M. C. Z. and U.S.N.M.), Cauto El Cristo, Oriente (Darling- ton, in M.C.Z. and U.S.N.M.), Rangel Mountains, Pinar del Rio (Dartling- ton, in M.C.Z. and U.S.N.M.). Jamaica: (Cameron, 1922, as nigrocoeruleus; Hubbard, in U.S.N.M.), Black River (Blackwelder station 416), May Pen (Blackwelder station 424A), Milk River (Blackwelder station 415), Spanish Town (Blackwelder sta- tion 877), Santa Cruz (Blackwelder station 421), Moneague (Darlington, in M.C.Z.), Kingston (Darlington, in M.C.Z.; Blackwelder station 379). Hispaniola: Harrr (Cameron, 1922, as limbatus and nigrocoeruleus; British Museum; Cameron collection) ; DOMINICAN REPUBLIC, Barahona (Darling- ton, in M.C.Z. and U.S.N.M.), Sanchez (Darlington, in M.C.Z.), Constanza (Darlington, in M.C.Z. and U.S.N.M.), San José de las Matas (Darling- ton, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z.). Puerto Rico: (Erichson, 1840; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; Wolcott, 1924, 1936), Jayuya (Oakley, in U.S.N.M.). Virgin Islands: Tortola (Cameron collection). St. Croix: (Chevrolat and Fauvel, 18638; Leng and Mutchler, 1914; Blackwelder station 341). St. Lucia: (Blackwelder stations 209A, 211A; Cameron collection). Barbados: (Blackwelder station 198B). St. Vincent: (British Museum and Cameron collection, as arduus Sharp; Cameron, 1922, as funerulus ; Blackwelder stations 166, 170, 176A). Grenada: (H. H. Smith, in British Museum and Cameron collection, as arduus; U.S.N.M.). Specimens ewamined.—I have seen 14 examples in the British Mu- seum, 18 in Dr. Cameron’s collection, 31 from the Museum of Com- parative Zoology, 8 in the United States National Museum, and 2 others received from Sr. Bierig, and 126 collected by me during 1935-37. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 443 Remarks.—This is a variable species in size, color, and details of sculpture. I was unable to separate the specimens segregated under the above names in the British Museum and Dr. Cameron’s collec- tion, and cannot now separate my specimens satisfactorily. It seems fairly certain that only one species is represented, occurring from Cuba to Grenada, at least. It differs from arduus Sharp of Central America in size, the unimpressed front of the head, and many de- tails of sculpture. One example from the Dominican Republic and one from Trinidad differ from the others in the punctation of the pronotum and elytra. They do not fall in the range of variation observed in the large series, but I am unwilling to describe them as new from unique speci- mens. They are now in the Museum of Comparative Zoology. IT have collected this species from dung and manure, from the muddy banks of ponds, from beneath stones along streams, and flying at dusk. LXVII. Genus STAPHYLINUS Linnaeus Staphylinus LINNAEUS, 1758, p. 421. Staphilynus ScHAEFFER, 1779, index (misspelling). Staphilinus Ovivier, 1791, p. 378 (misspelling). Slaphylinus FAIRMAIRE and GERMAIN, 1861, p. 413 (misspelling). Ouchemus Drs Gozis, 1886, p. 14. Subgenus Platydracus THOMSON, 1858, p. 29. Bemasus MULsAnt and REy, 1875, p. 257. Subgenus Dinothenarus THOMSON, 1858, p. 29. Subgenus Abemus MuLtsant and REy, 1875, p. 242. Subgenus Parabemus REI’TER, 1909, p. 118. Subgenus Chitocompsus BERNHAUER, 1913, p. 232. Subgenus Huwryolinus BERNHAUER, 1915, p. 297. Subgenus Nesiolinus BERNHAUER, 1915, p. 122. Subgenus Plesiolinus BERNHAUER, 1916, p. 93. Subgenus Chaetodracus MUtiEr, 1926, p. 27. Subgenus Apostenolinus BERNHAUER, 1934, p. 9. Genotypes.—S. mawillosus Linnaeus (designated by Latreille, 1810; later designations include S. erythrocephalus Fabricius, by Samou- elle, 1819; S. murinus Linnaeus, by Curtis, 1839; 8. olens Miiller, by Blanchard, 1845; and S. hévtus Linnaeus, by des Gozis, 1886). Of Ouchemus, Staphylinus erythropterus Linnaeus=Ouchemus ery- thropterus (Linnaeus) (original designation); of Platydracus, Staphylinus stercorarius Olivier=Platydracus stercorarius (Olivier) (monobasic) ; of Bemasus, Staphylinus lutarius Gravenhorst= Be- masus lutarius (Gravenhorst) (designated here); of Dinothenarus, Staphylinus pubescens DeGeer=Dinothenarus pubescens (DeGeer) (monobasic) ; of Abemus, Staphylinus chloropterus Panzer =A bemus chloropterus (Panzer) (designated here) ; of Parabemus, Staphylinus 444 BULLETIN 182, UNITED STATES NATIONAL MUSEUM fossor Scopoli=Parabemus fossar (Scopoli) (monobasic) ; of Chito- compsus, Staphylinus polyphemus Bernhauer=Chitocompsus poly- phemus (Bernhauer) (monobasic) ; of Huryolinus, Staphylinus sem- cyaneus Bernhauer=Furyolinus semicyaneus (Bernhauer) (mono- basic) ; of Vestolinus, Staphylinus bakeri Bernhauer=WNesiolinus ba- kert (Bernhauer) (designated here) ; of Plesiolinus, Staphylinus cos- taricensis Bernhauer=Plesiolinus costaricensis (Bernhauer) (desig- nated here); of Chaetodracus, Staphylinus patricius Bernhauer= Chaetodracus patricius (Bernhauer) (monobasic) ; of A postenolinus, Staphylinus (Apostenolinus) cariniceps Bernhauer (monobasic). Diagnosis —Head and pronotum with dense umbilicate punctures, usually feebly margined beneath the eyes; antennae widely sepa- rated at front of head, outer segments densely pubescent from the fourth joint; labrum strongly bilobed; palpi moderately elongate, fourth segment of maxillary much longer than third, labial similar to maxillary; gular sutures separate throughout; lateral margin of pronotum double, the lines united, side piece inflexed; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent, except possibly for a remnant of the second at sides; inter- segmental membranes of abdomen with a pattern of irregular rounded sclerotized areas arranged in longitudinal bands; femora unarmed. Remarks.—In the Bernhauer and Schubert Catalog the genera Staphylinus and Ocypus are more or less united, with eack group containing several subgenera. The characters that distinguish the two appear to be of generic value, and I am therefore omitting Ocypus and its subgenera from Staphylinus. Inasmuch as these names are somewhat mixed in the Catalog, I list below the names that have been coupled with Staphylinus but that are not herein placed in that genus. Genus Ocypus SAMOUELLE, 1819, p. 172. Anodus NorDMANN, 1836, p. 11. (Not Spix, 1829.) Ocys Curtis, 1889, p. 758 (misspelling). Acypus ERICHSON, 1839a, p. 444 (misspelling). Alapsodus ToTtTENHAM, 1939, p. 226 (as new name for Anodus). Subgenus Goérius STEPHENS, 1829, p. 275. Rayacheila MorscHutsky, 1845, p. 40 (typographical error). Ragacheila MANNERHEIM, 1846, p. 226 (misspelling). Rhagacheila MANNERHEIM, 1846, p. 226 (misspelling). Rhagochila MoTscHuLsky, 1858, p. 668. Matidus MorscHutsky, 1860a, p. 569. Subgenus TJ'asgius STEPHENS, 1829, p. 276. Subgenus Pseudocypus MULSANT and Rey, 1875, p. 291. Subgenus Parocypus BerNHAveR, 1915f, p. 52. Subgenus Protocypus MULtErR, 1923, p. 136. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 445 Subgenus Aulacocypus MULLER, 1925, p. 40. Subgenus Neotasgius MULLER, 1925, p. 41. Subgenus Xanthocypus MULLER, 1925, p. 41. This rearrangement is of course based on the genotypes. I have depended on the Bernhauer, Schubert, and Scheerpeltz catalogs for the relationships of the species designated. Of the above the geno- types are as follows: Of Ocypus, Staphylinus cyaneus Paykull= Ocy- pus cyaneus (Paykull) (original designation) ; of Anodus, A. falcifer Nordmann (designated by Tottenham, 1939); of Alapsodus, Anodus faleifer Nordmann=Alapsodus falcifer (Nordmann) (original designation) ; of Goérius, Staphylinus olens Miller = Goérius olens (Miiller) (designated here); of Rhagochila, R. inderiensis Mot- schulsky (monobasic); of Matidus, M. forficularius Motschulsky (monobasic); of Zasgius, Staphylinus ruftpes Latreille = Tasgius rupfipes (Latreille) (monobasic); of Pseudocypus, Staphylinus mus Brullé=Pseudocypus mus (Brullé) (designated by ‘Tottenham, 1939) ; of Parocypus, Staphylinus (Parocypus) dehradunensis Bern- hauer (monobasic); of Protocypus, Staphylinus (Protocypus) fulwotomentosus Eppelsheim (monobasic); of Aulacocypus, Ocypus gloriosus Sharp = Aulacocypus gloriosus (Sharp) (designated here) ; of Neotasgius, Ocypus brevicornis Weise = Neotasgius brevicornis (Weise) (monobasic) ; of Xanthocypus, Ocypus weisei Harold = Xan- thocypus weiset (Harold) (monobasic). In addition to these the name 7'richoderma Stephens has been listed as a subgenus of Staphylinus. Its type is Staphylinus murinus Lin- naeus = Trichoderma murinus (Linnaeus) (designated by Westwood, 1838). (The designation of nebulosus Fabricius by des Gozis, 1886, is therefore invalid.) Since murtnus is likewise the type of Ontholestes Ganglbauer, 1895 (designated by Tottenham, 1940), these two are isogenotypic. Zvrichoderma, being the oldest name, is to be used for the genus containing murinus. It will be noted that the true genotype of Staphylinus is also the type of Creophilus. This means that the name Staphylinus should be applied only to mawillosus and its allies (now known as Creophilus) and that the genus including erythropterus, etc., must take a later name. Several names are available for this, but none later than Ouchemus des Gozis need be considered. However, to most students of this family and to many other coleop- terists, this transfer of the name Staphylinus to another genus, the suppression of Creophilus, and the introduction of a newer name for the large genus now considered the type of the family will seem to be an unnecessary change of established names. Rather than make this decision in the present paper, I am using Creophilus and Staphy- linus in the sense of the Junk Catalog and leaving a final decision for a more propitious moment. 446 BULLETIN 182, UNITED STATES NATIONAL MUSEUM The only West Indian species of this genus is a widespread Central and South American species, which has apparently entered our region from both ends. It has been collected several times in Trinidad but has been reported from Cuba only once (Fauvel’s record is merely a copy of DuVal’s). The genus in its broad sense is readily recognized among our species by its size and dense pubescence and the dense punctation of the head and pronotum. 1. STAPHYLINUS ANTIQUUS Nordmann Staphylinus antiquus NORDMANN, 1886, p. 35.—ERICHSON, 1839b, p. 373.—SAHL- BERG, 1844, p. 788.—Lucas, 1857, p. 50.—SHArp, 1876, pp. 150, 154, 155, 156; 1884 pp. 381, 382, 383, 385; 1887, p. 786.—FAvvEL, 1901, p. 85.—VimLapa, 1901, p. 29.—BERNHAUER and SCHUBERT, 1914, p. 375,—BERNHAUER, 1933, p. 44. Ocypus cubae DUVAL, 1857, p. 36.—FAvveL, 1863, p. 432.—LENG and MUTCHLER, 1914, p. 406. Staphylinus cubae (DuVal) BERNHAUER and ScHUBERT, 1914, p. 377.—Lene and MourcuHtikgr, 1914, p. 406.) Description—Black with fuscous pubescence, abdomen opaque ru: fopiceous with apex rufocastaneous. ead with mandibles obtrian- gular, basal angles narrowly rounded ; eyes at less than half their length from base; with moderately dense large umbilicate punctures, the in- tervals narrow but generally visible; with indistinct ground sculpture on the intervals. Pronotwm as wide as long, quadrate, base rounded but with angles distinct; with punctation similar to that of head but without ground sculpture and leaving an irregular smooth midline posteriorly. LHlytra very densely and finely punctate, the punctures not separately visible but forming a dense sculpture; very densely pubescent. Abdomen very finely and densely punctate and pubescent, with a transverse row on each segment of very large posteriorly directed setigerous punctures. A/ale, eighth sternite with a very fee- ble emargination six times as wide as deep, and with surface smooth around emargination. Female, eighth sternite rounded. Length, 10-17 mm. Type locality—Para (Brazil). Of cubae, Cuba. Types.—Presumably in the Zoologische Museum, Berlin. Of cubae, presumably in the British Museum. Records.—The following are the records known to me: Cuba: (DuVal, 1857; Fauvel, 1863; Leng and Mutchler, 1914; as cubae). Trinidad: (Birch and Bryant, in British Museum), Caparo (Blackwelder collec- tion). South asad Brazit (Nordmann, 1837; Sahlberg, 1844; Lucas, 1857), Cotom- BIA, Brazin (Hrichson, 18389b; Sharp, 1876), VrENEzuUELA, Brazit, PERU (Fauvel, 1891), Cotompra, Peru, Braz (Fauvel, 1901). Central America: NicArAagua (Sharp, 1876), Mexico, GUATEMALA, HONDURAS, NIcARAGUA, PANAMA (Sharp, 1884), Mexico (Fauvel, 1891, 1901; Villada, 1901). MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 447 Specimens examined.—l have seen four examples in the British Museum and one in my own collection. Remarks.—I have been unable to find any differences between these West Indian specimens and examples of antiquus from Mexico and South America. Since DuVal’s description of Ocypus cubae fits this species exactly, it seems probable that it also belongs here. I find no record of the habits of this species. LXVIII. Genus CREOPHILUS Samouelle Creophilus SAMOUELLE, 1819, p. 172. Creophulus Voris, 1934, p. 240. Genotype—Staphylinus mawillosus Linnaeus=Creophilus maail- losus (Linnaeus) (original designation). Diagnosis—Head and pronotum without dense umbilicate punc- tures; head not margined beneath eyes; antennae widely separated at front of head, outer segments densely pubescent from seventh joint; labrum strongly emarginate at middle; fourth segment of maxillary palpus slightly shorter than third; labial palpus similar to maxillary; gular sutures separate, most approximate along middle; lateral margin of pronotum doubled, the marginal lines separate; anterior coxal cavities open behind except for a sclerotization about the spiracle (peritreme) ; front coxae large, exserted; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent; intersegmental membranes of abdomen with a pattern of irregular rounded sclerotized areas arranged in longitudinal rows; femora un- armed beneath. Remarks—This is a small genus but contains the largest species of Staphylinidae found in the West Indies. It occurs throughout the world, but none of its species are truly cosmopolitan. Only one species is recorded from the West Indies. 1. CREOPHILUS MAXILLOSUS (Linnaeus) Staphylinus mavillosus LINNAEUS, 1758, p. 421. Staphylinus anonymus Suuzer, 1761, p. 17. Staphylinus tertius SCHAEFFER, 1766, pl. 20, fig. 1. Staphylinus balteatus Drarer, 1774, p. 18 (as synonym). Staphylinus fasciatus Furssty, 1775, p. 21. Staphylinus nebulosus Fourcroy, 1785, p. 165. Staphylinus villosus GravenHorst, 1802, p. 160. (Not DeGeer, 1774.) Creophilus mazillosus (Linnaeus) SAMOUELLE, 1819, p. 172. Emus nebulosus MANNERHEIM, 1830, p. 20. Creophilus ciliaris SrEPHENS, 1882, p. 202. Creophilus fasciatus LAportTE, 1834, p. 111 (as new species). Creophilus villosus (Gravenhorst) LAporTE, 1834, p. 111. Emus mazillosus (Linnaeus) BotspuvAL and Lacorparre, 1835, p. 358. Staphylinus arcticus ERIcHson, 1839b, p. 348. 448 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Staphylinus cinerarius Ertcuson, 1839b, p. 350. Staphylinus ciliaris (Stephens) Ericuson, 1842, p. 211. Creophilus bicinctus MANNERHEIM, 18438, p. 229. Staphylinus bicinctus MANNERHEIM, 1848, p. 229. Emus villosus (Gravenhorst) MANNERHEIM, 1843, p. 229. Staphylinus orientalis MoTscHULSKY, 1857p, p. 67. (Not Bernhauer and Schubert, 1914. ) Creophilus orientalis (Motschulsky) MorscHULSKY, 1860b, p. 120. Creophilus fulvago MorscHutsky, 1860b, p. 120. Creophilus arcticus (Erichson) MorscHutsky, 1860b, p. 120. Oreophilus cinerarius (Erichson) Kraatz, 1874, p. 291. Creophilus imbecillus SHarp, 1874, p. 28.—Kraatz, 1874, p. 291. Creophilus medialis SHARP, 1874, p. 28.—Kraatz, 1874, p. 291. Creophilus subfasciatus SHarP, 1874, p. 28.—Kraatz, 1874, p. 291. Emus cinerarius (Erichson) FAuvEt, 1886, p. 50. Creophilus balteatus (Degeer) HAmILTon, 1894, p. 19. Creophilus pulchellus Meter, 1899, p. 99. Creophilus canariensis BERNHAUER, 1908b, p. 334. Oreophilus fasciatus (Fuessly) BERNHAUER and ScHUBERT, 1914, p. 398. Creophilus nebulosus (Fourcroy) BERNHAUER and ScHuBERT, 1914, p. 398. Oreophilus anonymus (Sulzer) BERNHAUER and ScHuBERT, 1914, p. 398. Creophilus ciliaroides Hatcu, 1988, p. 149 (as aberration). Description *.—Black, pubescence frequently whitish and arranged in a definite pattern, forming a white band across the elytra and irregular areas at the sides of the fourth and fifth abdominal tergites. Head about three-eighths wider than long, sides evenly rounded from front angles into base, eyes not interrupting side outlines; ver- tex finely but deeply punctate, with minute irregular punctures be- tween; without definite ground sculpture, but with a few large setigerous punctures behind the eyes and along margins. Pronotwm about one-fifth wider than long, truncate in front, nearly semi- circular behind, with posterior angles traceable; disk impunctate except for the minute punctulae as on head; with numerous setigerous punctures at anterior angles and along edges laterally. “lytra densely and rather finely punctate and densely pubescent; with a few much larger setigerous punctures arranged in a longitudinal discal series and a few submarginal individuals; surface coriaceous. Abdomen densely punctate and pubescent, with a few larger setiger- ous punctures; with indistinct transverse ground sculpture. Male, seventh sternite very feebly but not very widely emarginate; eighth with a rounded triangular emargination nearly twice as wide as deep; ninth sternite distinct. Female, seventh and eighth sternite rounded; ninth divided longitudinally into two sclerites. Length, 12 to 23 mm. 1° For a complete description of the structural characters, see Blackwelder, 1936. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 449 Type locality.“ Europe.” Ty pes.—Presumably in the Museum Uppsala. Records.—The following are the records known to me: Cuba: (Leng and Mutchler, 1914; Erichson, 1839b; Sharp, 1884). Isle of Pines: (Holland, 1917). Jamaica: Montego Bay (Blackwelder station 17), Kingston (Blackwelder station 434). go acts GUATEMALA (Sharp, 1884), Mexico (Erichson, 1839b, as vil- losus; Sharp, 1884; Fauvel, 1875). North America: Throughout the UNITrep STATES, CANADA, LABRADOR, NEWFOUND- LAND, GREENLAND, and ALASKA. Eurasia: KAMCHATKA, SIBERIA, EUROPE, ENGLAND, INDIA, JAPAN, SYRIA, PERSIA, Caucasus, AMUR, RUSSIA. Africa: Egypt, MADEIRAS, CANARIES, BARBARY COAST. Specimens examined.—I have seen seven examples collected by me on July 20, 1935, and six collected by Chapin and Blackwelder gn March 4,1937, Remarks.—This species is immediately recognizable among all West Indian staphylinids, or, indeed, among staphylinids of the entire world, by its characteristic size, form, and coloring. It is reported to be “almost cosmopolitan” but seems to be entirely absent from South America, tropical Africa, Australia, and Oceania; in fact re- stricted to the regions north of latitude 15° N. In a previous paper (Blackwelder, 1936) I treated the American representative under the name vé/osus without examining the validity of that name, which is better known to American entomologists. I now believe villosus to be a synonym of mawillosus in company with numerous other names, some of which have been considered to be va- rieties (subspecies). Although specimens have so far been reported only from Cuba and Jamaica, I believe it likely that the species will be found also in His- paniola and possibly in Puerto Rico. During my own collecting, car- rion was not available for examination on those islands, and it is only in this situation that the species occurs normally. Our 13 specimens were taken from beneath a dead hawk and a dead dog. LXIX. Genus XANTHOPYGUS Kraatz Xanthopygus Kraatz, 1857, p. 539. Lampropygus SHARP, 1884, p. 346. Heteropygus BERNHAUER, 1906, p. 195. Genotypes.—Staphylinus wanthopygus Nordmann=X anthopygus gvanthopygus (Nordmann) (by absolute tautonymy). Of Lampro- pygus, Staphylinus wxanthopygus Nordmann=Lampropygus wan- 450 BULLETIN 182, UNITED STATES NATIONAL MUSEUM thopygus (Nordmann) (designated here) ; of Heteropygus, Lampro- pygus (Heteropygus) giganteus Bernhauer (monobasic).?° Diagnosis —Head not densely umbilicately punctate, margined be- neath the eyes; antennae widely separated at anterior margin of head, outer segments densely pubescent from the fourth joint; labrum bi- lobed; palpi slender, fourth segment of maxillary longer than third, labial similar to maxillary; gular sutures becoming very approximate posteriorly; pronotum with lateral margin double, upper line not extending around anterior angle and never united with lower; ante- rior coxal cavities open behind, except for a large peritreme; front coxae very large, exserted; posterior coxae contiguous, “triangular” ; first and second abdominal sternites absent ; intersegmental membranes of abdomen marked with a pattern of irregular rounded sclerotized areas arranged in longitudinal rows; femora not strongly spinose beneath. Remarks.—This genus is to a certain extent given a new meaning by the return to it of XY. zanthopygus which is its true genotype. That species is not before me at present, but peaus agrees with the published conceptions of the genus and has been used (with XY. cognatus Sharp) in writing the diagnosis. This genus is distinguished from Philothalpus (and Xenopygus) by the disappearance of the upper marginal pronotal line at the anterior angles. This group of genera is badly in need of a thorough revision, and when that is made the number of genera will probably be further reduced. Nine West Indian examples of the genus have been seen. They belong to the following single species: 1. XANTHOPYGUS PEXUS (Motschulsky) Belonuchus perus MorscHuLsky, 1858, p. 667—GErMMINGER and HArozp, 1868, p. 596. Lampropygus pexus (Motschulsky) BrerNHAvER, 1906, p. 196.—BrERNHAUER and SCHUBERT, 1914, p. 407.—Lrna and MurcHter, 1914, p. 405. Description.—Black, apex of abdomen rufotestaceous. /Zead nearly one-third wider than long; eyes not prominent, occupying not over one-half the sides; the posterior angles marked by a ridge; with very scattered punctures except along middle, more dense along base, feebly umbilicate; with very fine and irregular strigulose ground sculpture. Pronotum scarcely wider than long, widest at anterior 2” The above designation of L. wanthopygus as genotype of Lampropygus may seem to be arbitrary but is based on what I believe to be sound reasoning. In proposing the name Lampropygus, Sharp was in error because he included the previously established type of Xanthopygus. His name will therefore always be at least in part synonymous with Xanthopygus. Less confusion will result if the name Lampropygus is definitely and irre- voeably rejected in this manner, and, if the other species included does prove to be distinct generically, a new name can be proposed for it that will have a more definite application and will not involve a change in the concept of an established genus. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 451 third, almost emarginate to front angles, sinuately narrowed to un- evenly rounded base; punctures similar to those of head at sides but with two irregular discal series of about 10 punctures; ground sculp- ture as on head. Llytra with moderately coarse and rather dense punctures separated by about their diameter; surface without ground sculpture, but uneven. Abdomen with moderate punctures strongly excavated behind, sometimes submuricate; surface alutaceous but with- out distinct ground sculpture. J/ale, eighth sternite feebly and broadly emarginate. Female, eighth sternite truncate. Length, 14 to 18 mm. Type locality —Colombia. Types—Presumably in the Zoological University Museum, Moscow. Records.—The following are the records known to me: St. Vincent: (H. H. Smith, in British Museum). Trinidad: (Bryant and Birch, in British Museum; Busck, in U.S.N.M.), Port of Spain (Marshall, in British Museum), Caparo (Blackwelder collection), Ariapito (Blackwelder collection), St. Augustine (Darlington, in M.C.Z. and U.S.N.M.), Mount Tucuche (Darlington, in M.C.Z.). South America: Cotompra (Motschulsky, 1858), CoLomBiA, VENEZUELA, GUIANA, Braziu (Bernhauer and Schubert, 1914), CotomBia, Brazin (British Museum). Specimens examined.—I have seen six examples in the British Museum, four from the Museum of Comparative Zoology, one in the United States National Museum, and two in my own collection. Specimens from South America also were examined in the British Museum. Remarks.—The single example from St. Vincent is perhaps not suffi- cient evidence that this species has extended its range north of Trini- dad, but I have no definite reason for doubting the record. This species is distinguished from cognatus of Central America by its sparser and less coarse punctures on the head and pronotum, by its coarser elytral punctation, and by the pale abdominal apex extend- ing only to the middle of the seventh tergite. I find no record of its habits. LXX. Genus PHILOTHALFPUS Kraatz Philothalpus KRAATZ, 1857, p. 540. Philotalpus GEMMINGER and HAROLD, 1876, index (misspelling). Subgenus Xenopygus BERNHAUER, 1906, p. 196. Subgenus Oligotergus Breria, 1937, p. 204. Genotypes—Staphylinus anceps Erichson=Philothalpus anceps (Erichson) (designated here). Of Xenopygus, Philonthus analis Krichson=Xenopygus analis (Krichson) (designated here); of Oligotergus, Philothalpus (Oligotergus) oculatus Bierig (monobasic). Diagnosis —Head without dense umbilicate punctures, margined beneath the eyes; antennae widely separated at front of head, outer 452 BULLETIN 182, UNITED STATES NATIONAL MUSEUM segments pubescent from fourth or fifth joint; labrum bilobed; palpi moderately long and slender, fourth segment of maxillary much longer than third, third segment of labial elongate, truncate, with apex sometimes compressed; gular sutures very approximate from middle; lateral margin of pronotum doubled, lines separate until around anterior angles; anterior coxal cavities entirely open behind; front coxae very large, exserted; posterior coxae contiguous, “tri- angular”; first and second abdominal sternites absent; intersegmental membranes of abdomen with pattern of irregular rounded sclerotized areas arranged in longitudinal rows; femora unarmed beneath. Remarks—This is a tropical American genus that apparently does not occur in the true West Indies. Xenopygus has always been placed some distance from this genus and apparently has never been compared with it. I am unable to find any character of generic importance to distinguish them, and I therefore reduce Yenopygus to the status of subgenus, although I am not certain that it can be retained even to that extent. It is distinguished by the complete lack of fine pubescence on the fourth antennal segment, by the more prominent subocular ridge, and by the uncompressed tip of the fourth segment of the maxillary palpus. I have seen 20 examples from the West Indies. These belong to two species which belong to two subgenera as defined above. KEY TO WEST INDIAN SPECIES OF PHILOTHALPUS 1. Pronotum with punctures evenly scattered except along midline. 1. fasciatus Pronotum with two distinct dorsal series outlining midline__---- 2. analis 1. PHILOTHALPUS FASCIATUS (Nordmann) Philonthus fasciatus NorDMANN, 1836, p. 112.—Ericuson, 1839b, p. 8397.—SHarp, 1884, p. 355. Staphylinus anceps ERIcHSon, 1839b, p. 396.—LACORDAIRE, 1854, p. 78. Staphylinus segmentarius ErIcHSON, 1839b, p. 397.—SHArpP, 1876, pp. 134, 136; 1884, p. 355. Staphylinus fasciatus (Nordmann) ErtcHson, 1839b, p. 897.—SHarp, 1884, p. 355. Philothalpus fasciatus (Nordmann) Kraatz, 1857a, p. 540.—SHaArp, 1884, p. 355.— BERNHAUER and SCHUBERT, 1914, p. 402 (as variety). Philothalpus segmentarius (Erichson) Kraarz, 1857a, p. 540.—SHarp, 1884, pp. 355, 356, 357.— BerNHAUER and SCHUBERT, 1914, p. 401 (as variety). Philothalpus luteipes SHArp, 1876, pp. 133, 134.—BrerNHAUVER and SCHUBERT, 1914, p. 402 (as synonym of segmentarius). Philothalpus anceps (Erichson) BreRNHAUVER and ScHuBeERT, 1914, p. 402.— PLAVILSTSCHIKOV, 1929, p. 80. Description—Head and pronotum black, elytra rufopiceous to testaceous, abdomen piceous with apex of segments paler and an apical band formed by testaceous apex of seventh tergite and testa- ceous base of eighth. Head (including the eyes) nearly one-fourth wider than long; eyes prominent, occupying three-fourths of the MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 453 sides; anterior part between eyes flattened; with scattered, coarse, indistinct umbilicate punctures except at center of vertex, dense behind eyes, sparse in front; surface with strong scaly ground sculp- ture throughout; pubescence long and conspicuous. Pronotwm one- fifth longer than wide, widest at anterior fifth, scarcely narrowed to prominent anterior angles, sinuately narrowed to strongly rounded base; punctation, sculpture, and vestiture similar to those of head. Elytra with outer posterior angles completely rounded; densely and rather coarsely punctate, but without distinct ground sculpture. Abdomen with third to fifth sternites with an arcuate basal eleva- tion; punctures irregular but generally fine and often submuricate; with very vague transverse ground sculpture. MA/fale, eighth sternite with a small, rounded, triangular emargination. Female, sternites rounded. Length, 8 to 9 mm. Type locality—Mexico. Of anceps, Colombia; of segmentarius, Colombia. Lypes.—Disposition not known. Of anceps and segmentarius, in either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Trinidad: (Bryant, in British Museum), Caparo (Klages, in British Museum), Manzanilla (Blackwelder station 108A), Port of Spain (Blackwelder station 115). South America: COLOMBIA (Erichson, 1889b, as anceps and segmentarius ; Sharp, , 1884; British Museum), VENEZUELA (British Museum), Brazim (Sharp, 1876, as luteipes; British Museum), Peru (Bernhauer and Schubert, 1914; Plavilst- schikov, 1929), British GUIANA (British Museum). Central America: PANAMA (British Museum), PANAMA and GUATEMALA (Sharp, 1884, as segmentarius), Mexico (Nordmann, 1836; Erichson, 1839b, as fasciatus; Sharp, 1884, as fasciatus). Specimens examined—From the West Indies I have seen nine examples in the British Museum and two collected by me in December 1935. Remarks.—The examples in the British Museum were separated under the names anceps and segmentarius, but I was unable to find any difference except color. A specimen of the dark form was borrowed and appears to be inseparable from the light form which I collected. This species belongs in the subgenus Philothalpus s.str. Inasmuch as fasciatus is the older name, it cannot be used as a variety of anceps. I have, therefore, suppressed anceps, which may be con- sidered to be a subspecies or variety by some workers. However, fasciatus must be the valid species. My specimens were found in freshly cut cocoa pods and flying at dusk. 2. PHILOTHALPUS ANALIS (Erichson) Philonthus analis Ertcuson, 1840, p. 495.—Lucas, 1857, p. 50.—SHARP, 1876, pp. 132, 186; 1884, p. 348. (Not Heer, 1838-42; not Fauvel, 1907.) 454 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Aanthopygus analis (Erichson) Kraarz, 1857a, p. 540.—SuHarp, 1876, p. 132,— BERNHAUER, 1906, p. 196. Lampropygus analis (Krichson) SHArp, 1884, pp. 347, 348.—Brrnuauer, 1906, p. 196. Xenopygus analis (Hrichson) BERNHAUER, 1910, p. 377.—BERNHAUER and ScHU- BERT, 1914, p. 405.—SaLt, 1929, pp. 450, 464.—ScHEEFRPELTz, 1933, p. 1416.— BALDUF, 1935, p. 82. Description—Head greenish, pronotum black with greenish tinge, elytra piceous, abdomen piceous with last two segments rufotesta- ceous. //ead (including eyes) one-fourth wider than long; eyes only slightly prominent, occupying two-thirds of the sides; anterior part not distinctly flattened; with scattered punctures except on vertex, punctures indistinctly umbilicate; with fine but distinct scaly ground sculpture interspersed with fine punctulae. Pronotum scarcely longer than wide, widest at anterior third, moderately narrowed to anterior angles, sinuately narrowed to strongly rounded base; with two in- definite series of large punctures along midline, of eight to nine punctures, with a lateral series of seven or eight and scattered addi- tional punctures; ground sculpture finer than on head; vestiture nearly absent. Hlytra conjointly widely emarginate, outer angles not evenly rounded; densely and coarsely punctate, but without distinct ground sculpture. Abdomen with sternites 3 to 5 with an angulate basal ele- vation; punctures sparse but fine and submuricate; with very vague transverse ground sculpture. d/ale, seventh sternite with a median longitudinal pubescent oval fovea, posteriorly very slightly and broadly emarginate; eighth sternite with a triangular notch a little wider than deep. Fema/le, sternites rounded. Length, 11 to 14mm. Type locality—‘Cayenne . . . ; Colombia.” Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Fecords—The following are the records known to me: Trinidad: Caparo (Blackwelder collection), Ariapita Valley (Blackwelder collec- tion), Arima (Kisliuk and Cooley, in U.S.N.M.), Manzanilla (Blackwelder station 103A), Mount Tucuche (Darlington, in M.C.Z. and U.S.N.M.). South America: (Sharp, 1884), Corompra, FrencuH GuIANA (BHrichson, 1840), Brazit (Sharp, 1876; British Museum), Brazit, Peru, GUIANA, VENEZUELA (Bernhauer and Schubert, 1914), ARGENTINA, PERU (Scheerpeltz, 1933), Boxrtvia (Costella, Lopez, and Mann, in U.S.N.M.). Central America: Mrxico, GUATEMALA, PANAMA (Sharp, 1884; British Museum ; Bernhauer and Schubert, 1914), Mexico, PANAMA (Blackwelder collection). Specimens examined.—l have seen from the West Indies one ex- ample in the United States National Museum, three from the Museum of Comparative Zoology, two in my own collection, and two collected by me in 1935-87. In addition I have seen numerous specimens from Central and South America. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 455 Remarks.—The above description was taken from a Trinidad ex- ample collected in 1935. Several series from South and Central Amer- ica indicate that the species is much more variable than the Trinidad examples indicate. The elytra are frequently greenish, and the entire green surface may be somewhat metallic. The punctures in the pro- notal series are very variable, numbering from 7 to 10. Immature specimens (from Bolivia) are dark rufous throughout except for the greenish head and the testaceous abdominal apex. This species be- longs in the subgenus Xenopygus as defined above. My examples were collected from freshly cut cocoa pods. LXXI. Genus DIOCHUS Erichson Diochus HRIcHson, 1889b, p. 300. Rhegmatocerus MOTSCHULSKY, 1858, p. 657. Genotype-—Diochus nanus Erichson (monobasic). Of Rhegmato- cerus, R. punctipennis Motschulsky (designated here). Diagnosis.—Head and pronotum without dense umbilicate puncta- tion; head not margined beneath the eyes; antennae moderately well separated at anterior margin of head, outer segments densely pubescent; labrum large, suborbicular; third segment of maxillary palpus large, acicular; gular sutures separate, converging toward base; pronotum with side margin double, the lines separate through- out; anterior coxal cavities open behind; front coxae large, exserted ; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent; intersegmental membranes of abdomen with a pat- tern of irregular rounded sclerotized areas arranged in longitudinal rows; femora unarmed beneath. Remarks.—This genus has generally been placed in the Xantholinini, but I am unable to reconcile many of its characters with those of the true xantholinines. The front of the head is entirely different from AXantholinus, the gular sutures are not united, the elytral suture is not beveled; the pattern of the intersegmental membranes of the abdo- men is different, and the males are clearly different from the females by the usual modifications of the seventh and eighth sternites. Six species have been recorded from the West Indies, but I am un- able to distinguish them. I have seen 244 examples from the West Indies, including the types of four of the names. I believe all are one species. 1. DIOCHUS NANUS Erichson Diochus nanus ERICHSON, 1839b, p. 801.—DUPONCHEL, 1841, p. 57.—-LACORDAIRE, 1854, p. 65.—CHEVROLAT and FAUVEL, 1863, p. 430.—LyNcH, 1884, p. 172.— SHARP, 1885, pp. 466, 467.—F AvVEL, 1891, p. 106.—HIcHELBAUM, 19()9, p. 161.— BLATCHLEY, 1910, p. 400.—BERNHAUER and SCHUBERT, 1914, p. 319.-—LeNne and MourcHtrr, 1914, p. 406.—Cammron, 1922, pp. 116, 117—BERNuAUER, 1929, p. 193. 456 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Diochus schaumi Kratz, 1860, p. 27.—SHarp, 1885, p. 467.—CAsey, 1906, pp. 431, 432, 483. Diochus parvulus KrAatz, 1860, p. 27. Diochus longicornis SHARP, 1876, pp. 184, 185; 1887, p. 791. Diochus vicinus SHARP, 1876, p. 185. Diochus tarsalis SHARP, 1876, p. 185. Diochus flavicans SHARP, 1876, p. 185. Diochus inornatus SHARP, 1885, p. 466 Diochus vilig SHARP, 1885, p. 467. Diochus maculicollis FAUVEL, 1891, p. 106. Diochus brevipennis Casey, 1906, pp. 481, 482. Diochus thoracicus CASEY, 1906, p. 482. Diochus pallidiceps CAsEy, 1906, p. 482. Rhegmatocerus conicicollis E1cHELBAUM, 1909, p. 161. Diochus perpletus CAMERON, 1922, p. 116. Diochus apicipennis CAMERON, 1922, p. 116. Diochus antennalis CAMERON, 1922, p. 117.—BERNHAUER, 1929, p. 193. Diochus pumilio BERNHAUER, 1929, p. 193. Description—Piceous to rufopiceous, margins sometimes paler, abdomen sometimes piceocastaneous. Head very smail, one-sixth longer than wide, sides arcuate behind eyes and evenly rounded into base; clypeus a little produced between base of antennae but only as a broad flat surface; with a few scattered punctures except along middle posteriorly; with very indefinite and indistinct ground sculp- ture. Pronotum one-sixth wider than head, two-sevenths longer than wide, oval, without trace of angles; with two discal pairs of punc- tures behind middle, a discal pair more widely separated in anterior half, and an irregular lateral series of three or four on each side; without ground sculpture. lytra three-sevenths wider than pro- notum; not distinctly punctate except for two discal rows, one sutural and one post-humeral, of four to seven submuricate punc- tures; surface uneven, feebly radially grooved near scutellum. Ab- domen densely and very finely punctate, with a few larger submu- ricate punctures and rather indefinite ground sculpture. dale, seventh sternite feebly emarginate in median fourth, eighth with a narrow rounded but very shallow emargination more distinct from behind than from beneath. Female, seventh sternite not emarginate, eighth broadly rounded. Length, 214 to 314 mm. Type locality —“In variis Americae meridionalis regionibus, e Co- lumbia.. .” Of schaumi, “Louisiana... Georgia”; of parvulus, Bahia, (Brazil) ; of longicornis, Tapajos, Brazil; of vicinus, Tapajos, Brazil; of tarsalis, Tapajos, Brazil; of flavicans, Tapajos, Brazil; of inornatus, “Nicaragua ..., Panama... ”3 of vélis, “Guatemala, near the city”; of maculicollis, “Caracas, Colonie Tovar,” Venezuela ; of brevipennis, St. Louis, Mo.; of thoracicus, Long Island, N. Y.; of pallidiceps, Austin, Tex.; of perplexus, Mount Gay Estate, Grenada; of apicipennis, St. Vincent; of antennalis, Grenada: of pumilio, Vera- cruz, Mexico. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 457 Types.—Hither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of schawmi and parvulus in the Deutsche Ento- mologische Institut; of longicornis, vicinus, tarsalis, flavicans, inor- natus, vilis, perplexus, and apicipennis, in the British Museum; of maculicollis, in the collection of Blanche Rancin, Caén, France; of brevipennis, thoracicus, and pallidiceps, in the Casey collection in the United States National Museum; of pumilio, in collection of Max Bernhauer; of antennalis, not recorded but in either the collection of Malcolm Cameron or the British Museum. FRecords.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Aspiro (Bierig, in Blackwelder collec- tion), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: (Hubbard, in U.S.N.M.; Cameron collection), Spanish Town (Black- welder station 877), Milk River (Blackwelder station 415), Santa Cruz (Blackwelder station 421), Trinityville (Blackwelder station 428), Kingston (Darlington, in M.C.Z.). : Hispaniola: DoMINICAN REPUBLIC, Sanchez (Darlington, in M.C.Z. and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z.), Villa Altagracia (Darlington, in M.C.Z.). Puerto Rico: Maricao (Blackwelder station 47A). St. Croix: (Blackwelder stations 341, 344). Guadeloupe: Basse Terre (Blackwelder station 79C). St. Lucia: (Blackwelder stations 207C, 209A, 211A, 444B). Barbados: (Blackwelder stations 193A, 202A). St. Vincent: (Cameron, 1922a; British Museum). Grenadines: Mustique (British Museum; Cameron collection). Grenada: (Cameron, 1922a; British Museum; Cameron collection). Trinidad: Tacarigua (Blackwelder station 107C), foothills north of Tunapuna (Weber, in M.C.Z. and U.S.N.M.), St. Augustine (Weber, in M.C.Z. and U.S.N.M.), base of Galeota Point (Weber, in M.C.% and U.S.N.M.), 11 miles north of Arima (Weber, in M.C.Z. and U.S.N.M.). South America: (Erichson, 1839), Braz (Sharp, 1876, as longicornis, vicinus, tarsalis, and flavicans; Kraatz, 1860), Cotompra (Erichson, 1839b; Sharp, 1885), ARGENTINA (Lynch, 1884), VENEZUELA (Fauvel, 1891). Central America: GuATEMALA, PANAMA, Nicaracua (Sharp, 1885, as nanus, inornatus, schaum, and vilis), Mexico (Bernhauer, 1929). North America: LOUISIANA (Kraatz, 1860; Sharp, 1885), Ruope IsLaAnp, Vir- GINIA, NorRTH CAROLINA, NEw YorkK, Missourt, TExas (Casey, 1906, as palli- diceps), MICHIGAN, FLortpA, OHIO, MASSACHUSETTS, MISSISSIPPI (U.S.N.M.). Specimens examined—tI have seen 165 examples in the British Museum (including types of Dr. Cameron’s species), 27 in Dr. Cameron’s collection, 28 from the Museum of Comparative Zool- ogy, 4 in the United States National Museum, 1 from Sr. Bierig in my own collection, and 49 collected by me during 1935-37. In addi- tion to these I have examined the types of Casey’s species and good series of North American specimens. Remarks.—My examination of examples of nanus, schaumi, per- plexus, apicipennis, and antennalis in the British Museum and in Dr. Cameron’s collection left me unable to distinguish any of the 449008—42—30 458 BULLETIN 182, UNITED STATES NATIONAL MUSEUM five or to separate my own specimens from any of them. A similar experience was had with the three Casey species. [Examination of the characters used to separate these has shown that they are variable characters and apparently not of specific value. I can only con- clude that the names are synonymous. In addition to these species of which I have seen examples, a careful examination of the other described American species (except formicetorum Bernhauer) shows that the same characters have been used in every case, and no varia- tions are recorded which are not included in the series at hand. I am forced to conclude that they too are synonyms, and I feel that the evidence is strong enough to justify placing the names in the formal synonymy. My specimens were collected from among decomposing trash in canefields, from under dung, rotting fruit, moss, and stones along edge of streams, and flying at dusk. LXXII. Genus CEPHALOXYNUM Bernhauer Cephaloxynum BERNHAUER, 1907, p. 282. Subgenus Ponticulus Brerta, 1931, p. 424. Genotype-—C. gestroi Bernhauer (monobasic). Of Ponticulus, Cephaloxynum (Ponticulus) rambouseki Bierig (original designa- tion and monobasic). Diagnosis —Form slender, depressed; head and pronotum without dense umbilicate punctures; head not margined beneath or behind the eyes; antennae widely separated at anterior margin of head, outer segments pubescent from the fourth joint; labrum bilobed (though separated from the clypeus by a smooth area that might be mistaken for the labrum itself) ; fourth segment of maxillary palpus shorter and much more slender than the third; labial palpus similar to maxillary; gular sutures united, at least basally; lateral margin of pronotum doubled, the lines not clearly united anteriorly; ante- rior coxal cavities open behind; front coxae very large, exserted; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent; femora unarmed beneath; tarsi 5-segmented. Remarks.—This genus was described in the Piestinae, but I can see no reason for regarding it as anything but a true staphylinine, especially since the redefinition of the Piestinae. Ihave assumed that Sr. Bierig’s species from Cuba is actually a Cephalowynum, since the genus is not otherwise available to me in identified material. I con- sider that Ponticulus cannot be maintained even as a subgenus on the basis of the characters given by Bierig, and his fgures and descrip- tion leave little doubt that rambouseki is congeneric with tricolor. If tricolor is properly assigned to Cephalorynum, then this transfer to the Staphylininae is valid. If not, then the genus here treated would take the name Ponticulus. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 459 I have seen four specimens from the West Indies representing two species of which one is new. KEY TO WEST INDIAN SPECIES OF CEPHALOXYNUM 1. Head with two deep foveae on vertex_._--__~----_=+~~-- 2. rambouseki Heads withoultovede: OnuVellexa=2 2225S a 2 See 1. chapini 1. CEPHALOXYNUM CHAPINI, new species Description.—Head rufopiceous, pronotum and abdomen rufotes- taceous, elytra fuscotestaceous. //ead quadrate, one-tenth longer than wide; with coarse and distinctly umbilicate punctures scattered evenly except along middle, generally separated by more than their diameter ; surface with fine but distinct scaly ground sculpture throughout. Pronotum one-ninth longer than wide, front angles broadly rounded, sides straight posteriorly and moderately narrowed, hind angles dis- tinct; more finely punctate than head, but evenly except for smooth median space; with scaly ground sculpture finer than on head. Elytra moderately densely but feebly punctured; surface shining but slightly uneven. Abdomen sparsely and finely submuricately punc- tured; with indistinct transverse strigulae. Length, 2144 mm. Type locality —Jamaica, between Fern Gully and Ewarton in vicin- ity of Moneague. Types—Holotype, in the United States National Museum (No. 52456) collected by Chapin and Blackwelder on February 2, 1937. Records.—The following is the only record known to me: Jamaica: Moneague (Blackwelder station 375). Specimens examined.—\ have seen only the unique type. Remarks.—This species is very distinct by its even head without ridges or depressions. The shape of the pronotum is also distinctive, as well as the even arrangement of the pronotal punctures, The type was taken flying at dusk. 2. CEPHALOXYNUM RAMBOUSEKI Bierig Cephalorynum rambouseki Brertc, 1931, p. 424.—ScHEERPELTzZ, 1933, p. 995.— Brieria, 1937, p. 278. Description.—Testaceous, head picescent especially about the eyes, elytra piceous except indefinitely at base, fourth and fifth visible ab- dominal segments piceous. /7cad quadrate, as broad as long; with a deep circular fovea on each side of midline at center of vertex, smooth midline feebly but broadly elevated toward front; with coarse punc- tures separated by about half their diameters; with very vague stri- gulose ground sculpture. Pronotum as broad as long, front angles not rounded, sides gradually curved into rounded base; without dis- tinct punctures; ground sculpture as on head. /7ytra with punctures 460 BULLETIN 182, UNITED STATES NATIONAL MUSEUM rather dense but obsolescent, very shallow; with scarcely a trace of ground sculpture. Abdomen with sparse and fine indistinct sub- muricate punctures; without distinct ground sculpture. Length, 2 to 254 mm. Type locality --Cuba, Tapaste, Sierra Bonilla. Types.—Type, February 29, 1930, and one paratype from Rangel, Sierra del Rosario, Cuba, February 9, 1930, in the collection of Alex- ander Bierig, Habana, Cuba. [In 1937 Bierig recorded also one “hypotype” in his collection. An example sent to the United States National Museum in 1938 (No. 52698) is labelled “cotype” but appar- ently is not more than a hypotype. ] Records.—The following are the records known to me: Cuba: Tapaste in the Sierra Bonilla (Bierig, 1931), Rangel in the Sierra del Rosario (Bierig, 1981), Camoa in the province of Habana (Bierig, in U.S.N.M.), Buenos Aires in the Trinidad Mountains (Darlington, in M.C.Z. and U.S.N.M.). Specimens examined—I have seen only the one “cotype” in the United States National Museum and two specimens from the Museum of Comparative Zoology. Remarks.—This species is very distinct because of the foveate vertex as well as by the impunctate pronotum. The color pattern is likewise unique in our fauna. The types were found under bark of fallen logs. LXXIII. Genus HOLISUS Erichson Holisus Er1cHson, 1839b, p. 298. Piestomorphus MorscHULSKY, 1858, p. 666. Hyptioma CAsEy, 1906, p. 359, 362. Genotype.—H. analis Erichson (designated by Duponchel, 1841). Of Piestomorphus, P. ater Motschulsky (monobasic) ; of Hyptioma, H. cubensis Casey (monobasic). Diagnosis —Form very much depressed; head and pronotum with- out dense umbilicate punctures; head not margined beneath or behind the eyes; antennae widely separated at anterior margin of head, outer segments pubescent from the fourth joint; labrum bilobed; fourth segment of maxillary palpus shorter and much more slender than the third; labial palpus similar to maxillary; gular sutures united; lateral margin of pronotum tripled, the lines united anteriorly ; anterior coxal cavities partly closed by a pair of sclerites behind the spiracles; front coxae very large, exserted; posterior coxae contiguous, “triangular”; first and second abdominal sternites absent ; intersegmental membranes with a pattern of irregular rounded sclerotized areas arranged in longitudinal rows; femora unarmed beneath; tarsi 5-segmented. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 461 Remarks.—This genus is restricted to the Western Hemisphere, where it is represented by a considerable number of species, each being generally very rare in collections. It was described as a xantholinid but seems to be a normal member of the true Stayphy- linini, The two West Indian species described by Erichson have been cataloged as occurring only in South America and have therefore been overlooked by students of the West Indies fauna. They were described from St. John and St. Vincent without mention of any South American localities. I believe I have recognized them both in the present series. I have seen 26 examples of this genus from the West Indies. ‘These belong to three species of which one is new. KEY TO WEST INDIAN SPECIES OF HOLISUS SPOUT CAT aCe eo eee Ee eee 2 Mahiumenocrcalinatel- = ee EO ee 1. debilis 2eBlackathroushout, except tor wuster=.2. === -<- 2-22 = ee 3. laevis Inmeereat spart rufous ‘or ‘testaceousi ofS. Sis ea eee tse 2. guildingii 1. HOLISUS DEBILIS Erichson Holisus debilis ErtcHson, 1839b, p. 300.—Lacorpaire, 1854, p. 65.—BERNHAUER and ScHvueErt, 1914, p. 321. Hyptioma cubensis Casry, 1906, p. 862.—HicHEeLBAum, 1209, p. 162.—Casey, 1910, p. 106.—BeRNHAUER and SCHUBERT, 1914, p. 320.—ScHEERPELTZ, 1933, p. 1326. Holisus cubensis (Casey) CasEy, 1910, p. 106—Lrne and Murcuter, 1914, p. 406.—ScCHEERPELTZ, 1933, p. 1324. Holisus rufoniger CAMERON, 1922, p. 117.—ScHEERPELTZ, 1933, p. 1326. Holisus erythroderus Fauve, MS—CaMEron, 1922, p. 117. Description —Head piceous or rufopiceous, pronotum, elytra, and abdomen rufous or rufotestaceous, elytra more or less infuscate es- pecially apically. Head quadrate, as wide as long; with coarse and indistinct umbilicate punctures evenly scattered and generally sepa- rated by about their diameter, in front arranged in denser series to outline a Y-shaped elevation; surface with distinct longitudinal strigulae. Pronotum suborbiculate, the sides nearly straight but hind angles not at all prominent; with coarse punctures scattered over sur- face except along middle, very little denser posteriorly; surface with distinct diagonal strigulae throughout. Zlytra rather finely and densely punctate, the surface shining but feebly coriaceous. Adbdo- men sparsely and moderately finely submuricately punctate; with in- distinct transverse strigulae. Length, 3 to 44% mm. Type locality—St. John. Of rufoniger, Haiti; of cubensis, Habana, Cuba. Types—In either the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of rufoniger, in collection of Dr. Cameron; of cubensis, in Casey collection, U.S.N.M. No. 48241. 462 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Records.—The following are the records known to me: Bahamas: Watling Island (Greenway, in M.C.Z.). Cuba: (Casey, 1906; Leng and Mutchler, 1914), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Hispaniola: Hartt (Cameron, 1922). Puerto Rico: El Yunque (Blackwelder station 351A). St. John: (Erichson, 1839b; Lacordaire, 1854). St. Lucia: (Blackwelder stations 220F, 227). St. Vincent: (Cameron, 1922; British Museum). (South America: (Bernhauer and Schubert, 1914).) Specimens examined—I have seen eight examples collected by me in 1935-37, three from the Museum of Comparative Zoology, and one in the British Museum, beside the three types of cubensis in the United States National Museum. I have not examined the type in Dr. Cameron’s collection. Remarks.—This is a rather variable species in details of color and punctation, but I am not able to find characters that will divide it at all satisfactorily. The Erichson species was certainly unknown to Col. Casey when he described cubensis, and it was probably unknown to Dr. Cameron when he described rufoniger. I have directly compared the type of cubensis with specimens from the other islands, but of rufoniger I have made only a brief examination. I believe there is little doubt that they are all the same species. The record of South America by Bernhauer and Schubert, 1914, is almost certainly an error, since they make no mention of the type locality. My specimens were collected from decaying cocoa pods, from under chips and frass on stumps, and flying at dusk. 2. HOLISUS GUILDINGII Erichson Holisus guildingiti ErecHson, 1839b, p.800.—LAcorDAIRE, 1854, p. 65.—BrrN- HAUER and ScHUBERT, 1914, p. 322. Description —Head black to rufopiceous, pronotum and abdomen rufous or rufotestaceous, elytra testaceous; pronotum with aeneous luster. Head quadrate, as wide as long, sides arcuate; with distinctly elongate umbilicate punctures fairly evenly scattered and not very distinctly marking ridges in front, leaving a smooth midline; surface with fine but. dense longitudinal strigulae. Pronotwm suborbiculate but with basal angles slightly prominent; with coarse punctures more or less serially limiting the smooth midline and scattered laterally, more sparsely in front; with strigulae generally diagonal. Llytra moderately densely and feebly punctate; surface shining but slightly uneven. Abdomen sparsely and moderately finely submuricately punctate; with indistinct transverse strigulae. Length, 314 to 5144 mm, MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 463 Type locality. —St. Vincent. Types.—Presumably in the Hope Museum, Oxford. Records —The following are the records known to me: Puerto Rico: El Yunque (Blackwelder 351A), Mayagiiez (Blackwelder station 358B). Antigua: (H. E. Box, as Blackwelder station 444Q). St. Lucia: (Blackwelder station 227). St. Vincent: (Hrichson, 1839b) ; Lacordaire, 1854). [South America: (Bernhauer and Schubert, 1914).] Specimens examined.—I have seen only the nine examples col- lected by me in 1935-37. Remarks.—This species also is variable in details of the arrange- ment of punctures and ridges on the head but seems to be distinct from debilis by the elongation of the punctures of the head, by the sparse and coarse pronotal punctures, and by the prominence of the hind angles of the pronotum. The record of South America by Bernhauer and Schubert is almost certainly an error since they do not cite the type locality. These specimens were collected from rotting cocoa pods, from under chips and frass on stumps, from under bark, and “from inside of scarlet bracts of a large unknown species of Heliconia” (H. E. Box). 38. HOLISUS LAEVIS, new species Description —Black, with aeneous luster. Head quadrate, as wide as long; with coarse punctures arranged in an indefinite semicircle back of the bases of the antennae and two confused bands converging toward the neck, with two punctures on the frontal region and scat- tered punctures at the sides behind the eyes; surface finely but dis- tinctly longitudinally strigulate throughout. Pronotum suborbi- cular, anterior angles broadly rounded, sides not emarginate pos- teriorly but posterior angles traceable; with a median smooth space, and scattered punctures laterally, more or less in two series in front; surface strigulose, longitudinally behind but diagonally in front. Elytra moderately coarsely and densely punctate, the punctures obscured by fine radiating wrinkles, but surface shining. Abdomen with sparse punctures greatly excavated behind; surface transversely strigulate. Length, 6 mm, Type locality —Jamaica, 1 mile east of Bath in parish of St. Thomas. Thomas. Types.—Holotype, in the United States National Museum (No. 52455), collected by Chapin and Blackwelder on February 8, 1937. Records.—The following are the records known to me: Cuba: Mountains north of Imias, Oriente (Darlington, in M.C.Z.). Jamaica: Bath in St. Thomas (Blackwelder station 392B). 464 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—I have seen only the type and one example from the Museum of Comparative Zoology. Remarks.—This species is distinguished by its quadrate head, its coarse and sparse pronotal punctures, and its uniform black color. I am unable to distinguish the Cuban example from the type. It was taken at an elevation of 3,000 to 4,000 feet. A species very similar to this occurs in Central and South America. It may differ in details of sculpture or may be the same species. I am unable to determine the range of variation of the West Indian species so cannot state definitely whether they are the same or not. The type was found under bark of a silk cotton tree. LXXIV. Genus HETEROTHOPS Stephens Heterothops STEPHENS, 1829, p. 284. Trichopygus NORDMANN, 1836, p. 187. Heterotops NoRDMANN, 1836, p. 187 (misspelling). Genotypes.—Staphylinus binotatus Gravenhorst=Heterothops bi- notatus (Gravenhorst) (designated here). Of Z'richopygus, Tachy- porus dissimilis Gravenhorst=Trichopygus dissimilis (Gravenhorst) (designated by Tottenham, 1939). Diagnosis——Head and pronotum without dense umbilicate punc- tures, more or less margined beneath the eyes; antennae widely sepa- rated at front of head, not geniculate nor serrate, outer segments densely pubescent from third or fourth segment; labrum transverse}; mouthparts short; palpi subulate, fourth segment of maxillary shorter than third, third segment of labial slender; gular sutures separate; lateral margin of pronotum simple; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, with a small expansion over the base of femur; first and second abdominal sternites absent; femora unarmed; tarsi all 5-segmented. Remarks.—This is a moderate-sized genus occurring throughout the world but generally with rather limited specific ranges. It is distinguished principally by the structure of the mouthparts, but in the West Indies may be known among Quediinae by its 5-seg- mented tarsi or the short palpi. Two new species are all that are known to me from the West Indies. KEY TO WEST INDIAN SPECIES OF HETEROTHOPS 1. Eyes at more than their length from base_______-__________ 1. rambouseki Eyes at much less than their length from base___--____________ 2. ocularis MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 465 1. HETEROTHOPS RAMBOUSEKI, new species Hetcrothops liliputanus RAMBOUSEK, MS. Description.—Head and tip of abdomen piceous, rest testaceous or rufescent. Head elongate, widest posteriorly; eyes at nearly three times their length from base; not punctate except at the sides; with very indistinct ground sculpture. Pronotum scarcely wider than long, broadly rounded behind, somewhat arcuately narrowed in front; with a single pair of discal punctures just before middle; with ground sculpture as on head. Zlytra with rather fine submuricate punctures separated by two or three times their diameter; with rather distinct transverse strigulose ground sculpture. Abdomen with punctation finer than on elytra and denser posteriorly; ground sculpture in- distinct. Length, 114 to 2 mm. Type locality —Cuba, district of Jamaica, province of Habana. Types.—Holotype in collection of Dr. Cameron, collected on Oc- tober 5, 1928, by Dr. Rambousek; and two paratypes in United States National Museum (No. 52476). (The type was labeled paratype of H. liliputanus by Dr. Rambousek, in whose collection there are undoubtedly additional examples.) Records—The following are the records known to me: Cuba: Habana (Rambousek, in Cameron and Rambousek collections), Cayamas (Schwarz, in U.S.N.M.). Jamaica: Bath in St. Thomas (Blackwelder station 387A). Specimens examined.—I have seen only three specimens of which the two in the United States National Museum are in very poor condition. The example in the Cameron collection was borrowed for study and is herein designated as the holotype. Remarks.—This species is readily recognized by the size of the eyes, as well as by the length of the body which is distinctly less than that of ocularis. My specimen was taken from fungus (a species of Hydnum). 2. HETEROTHOPS OCULARIS, new species Description—Head and pronotum piceous, elytra and abdomen castaneous (sometimes picescent). Mead suborbicular, as broad as long; eyes very large, occupying more than half the length; with marginal punctures only; with vague traces of ground sculpture. Pronotum one-seventh wider than long, broadly rounded behind, some- what arcuately narrowed in front; with a single pair of discal punc- tures at apical third; with traces of ground sculpture as on head. EHlytra with fine submuricate punctures separated by one to two times their diameter; without distinct ground sculpture. Abdomen with punctures as on elytra but more distinctly muricate; at most with 466 BULLETIN 182, UNITED STATES NATIONAL MUSEUM traces of ground sculpture. Male, eighth sternite with a small emar- gination three times as wide as deep. Female, eighth sternite rounded. Length, 21% to 31% mm. Type locality —Trinidad, mouth of cave in Aripo Valley; elevation 9,600 feet. T'ypes—Holotype and one paratype (females) in the Museum of Comparative Zoology; one paratype (male) in the United States Na- tional Museum (No. 52530) ; collected April 19, 1935, by N. A. Weber. Records.—The following is the only record known to me: Trinidad: Aripo Valley (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—l have seen only the three types. Remarks.—This species is at once distinguished from H. rambouseki by the very large eyes which occupy most of the sides of the head. This development also forces the infraocular ridge onto the ventral surface of the head. In details of sculpture and in appearance the two species differ little. The types were found at the mouth of a cave, but I have received no other record of their habits. LXXY. Genus ACYLOPHORUS Nordmann Acylophorus NORDMANN, 1836, p. 127. Rhygmacera MOTSCHULSKY, 1845, p. 40. Rhegmacera Fauve, 1895, p. 275 (misspelling). Genotype.—A. ahrensii Nordmann (designated here) ; the citation as type of A. glaberrimus (Herbst) by Bierig in 1938 can be recognized only through the synonymy of ahrensii and glaberrimus. Of Rhyg- macera, R. nitida Motschulsky (monobasic). Diagnosis —Head and pronotum without dense and umbilicate punctures, margined beneath the eyes; antennae widely separated at front of head, strongly geniculate at the first joint, segments densely pubescent from the third joint; labrum transverse but rounded, with a few long setae; mouth-parts not very long or very slender, fourth seoment of maxillary palpus enlarged and either truncate or acumi- nate, labial palpus with second and third segments much enlarged and flattened, forming a distinct club; gular sutures separate but approximate near base; lateral margin of pronotum single; anterior coxal cavities open behind; front coxae large, exserted ; posterior coxae contiguous, “triangular,” with a small expansion over the base of the femur; first and second abdominal sternites absent, except for a rem- nant of the second at the side; femora unarmed; all tarsi 5-segmented ; the sexes usually not externally distinguished. Remarks.—This rather small genus is found in most parts of the world but the species are usually not widespread. It is distinguished from Heterothops principally by the structure of the mouthparts. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 467 Two species described from Cuba are not available to me. Four others that appear to be new are described below. I have seen 20 examples from the West Indies. KEY TO WEST INDIAN SPECIES OF AOYLOPHORUS i Species described by Bierig trom Cubase. 2228 ee 2 Species described here from Hispaniola and Trinidad____________________ 3 2; Abdomen withsdistinct..aeneous. luster.__=-—._.-—=.~___._-..-.. 4. hatuey Abdomen without distinct aeneous luster_____________-_______ 2. insulanus 3. Scutellum densely and finely punctate throughout; punctures of abdomen coarse, tuberculate; abdomen with strong aeneous luster____ 5. metallicus Scutellum with a group of rather coarse punctures surrounded by smooth areas; punctures of abdomen rather fine; abdomen without strong aeneous [USCC Tee eee ERS SE ae IAT SRR el ey Same Meena eee APART be BAAR SEONG, 4 4. Fourth segment of maxillary palpus abruptly diagonally truncate, about twice aS5 lONS 4aASy DIOR se cnet ee reese aN er ote ae Nae ees 6. darlingtoni Fourth segment of maxillary palpus elongate, attenuated, about three times ASMLON SaaS); DEOAC st. x Peete a. Pee eee As | ee eek. ee 5 5. Abdomen with slight aeneous luster___.__-_____-__...._--_____ 3. aenescens Abdomen without trace of aeneous luster___.___-______-_______ 1. trinitatis 1. ACYLOPHORUS TRINITATIS, new species Description.—Black, pronotum rufopiceous, margins of abdominal segments rufescent. Mead as broad as long, subcircular, eyes about half as long as head; maxillary palpus with third and fourth segments somewhat enlarged, fourth acuminate; with a few large setiferous punctures around sides of disk and one above each eye, and with a few minute and indefinite punctulae; without ground sculpture. Pro- notum one-sixth wider than long, sides and base completely rounded from front angles; with a pair of large discal punctures at middle and another at apical sixth, and one near side at apical third; with scat- tered punctulae as on head and without ground sculpture. Scutellum with a few punctures as on base of elytra surrounded by smooth margin. “lytra with moderately coarse but distinctly muricate punc- tures separated by one to two times their diameter; without ground sculpture. Abdomen with rather coarsely submuricate punctures bearing very strong setae and separated by one to two times their diameter; without ground sculpture. Length, 5 to 5144 mm. Type locality —Trinidad, foothills north of Tunapuna. Types.—Holotype and one paratype in the Museum of Compara- tive Zoology; two paratypes in the United States National Museum (No. 52581) ; collected on May 25, 1935, by N. A. Weber. (Paratypes partly from the other localities listed below.) Records —The following are the records known to me: Trinidad: Tunapuna (Weber, in M.C.Z. and U.S.N.M.), Cedros (Darlington, in M.C.Z.), St. Augustine (Darlington, in U.S.N.M.) Specimens examined.—I have seen only the four types. 468 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Remarks.—This species is not greatly different in any obvious char- acter from the others described here but seems to be distinct as outlined in the key. It is much smaller than darlingtoni, which is the only other species known from Trinidad. I have received no record of its habits. 2. ACYLOPHORUS INSULANUS Bierig Acylophorus insulanus Brirerie, 1938, pp. 125, 128, 129, figs. 16, 28. Description.—Intense black, border of seventh tergite and apex of abdomen rufous. Head with postgena densely punctate and pubes- cent; occipital furrow distinct; fourth segment of maxillary palpus slightly enlarged and acuminate; finely and densely punctured behind the eyes. Pronotwm one-twelfth wider than long. Llytra one-fourth broader than long. Scutellwm with punctures as on base of elytra. Abdomen with punctures crowded at the base of each segment, so as to elevate the narrow intervals. Length, 614 to 7144 mm. (From Bierig.) Type locality.—Cuba. Lypes.—tIn the collection of Alexander Bierig. fecords.—The following are the records known to me: Cuba: El Mariel (Bierig, 1988), Laguna La Canoa (Bierig, 1938). Isle of Pines: (Bierig, 1938). Specimens examined.-—I have seen no examples of this species. Remarks.—This species is placed in the subgenus Acylophorus s. str. by Bierig. It had previously been identified as A. pronus Erichson, from which he distinguishes it. The types were found on the banks of streams and ponds, about the base of plants. 3. ACYLOPHORUS AENESCENS, new species Deseription.—Black, abdomen with faint bluish luster. Head sub- orbicular, eyes more than half as long as head; maxillary palpus with third and fourth segments somewhat enlarged, fourth acuminate; with a few large setiferous punctures around sides of disk and one above each eye, and with scattered punctulae almost obsolete; without ground sculpture. Pronotum slightly wider than long, with a pair of large discal punctures just before middle and another subapical, and one near side at apical fifth; with obsolete punctulae as on head and without ground sculpture. Scutellum with a few punctures as on base of elytra surrounded by a broad smooth margin. lytra with moderately coarse but distinctly muricate punctures separated by one to two times their diameter; without ground sculpture. Abdomen with moderate submuricate punctures bearing strong setae and sep- arated by two to four times their diameter; without ground sculpture. Length, 5144 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 469 Type locality—Dominican Republic, Sanchez. Types.—Holotype in the Museum of Comparative Zoology, col- lected in July 1938 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Hispaniola: DoMINICAN Repusuic, Sanchez (Darlington, in M.C.Z.). Specimens examined.—I have seen only the unique type. Remarks.—This species differs from metallicus in the sparse coarse scutellar punctures, in the nontuberculate punctures of the abdomen, and in the weakness of the luster. I have received no record of its habits. 4. ACYLOPHORUS HATUEY Bierig Acylophorus (s. str.) hatuwey Breria, 1938a, pp. 120, 126, 127, 128, figs. 2, 29. Description.—Black, apical border of abdominal tergites rufous, with distinct aeneous luster. Head short oval; postgena punctured and pubescent; occipital furrow well-marked; maxillary palpus with third and fourth segments somewhat enlarged, fourth acuminate but shorter than in insu/anus. Sceutellum finely punctate, as base of elytra. Elytra scarcely longer than pronotum, more crowded and more asperate than in insulanus. Abdomen with punctures moderately dense on the basal tergites. Length, 514 to 7mm. (From Bierig.) Type locality.—Cuba. Types.—In the collection of Alexander Bierig. Records.—The following are the records known to me: Cuba: El Rangel, Sierra del Rosario (Bierig, 1988), Laguna del Somorrostro (Bierig, 1938), Aspiro (Bierig, 1938). Specimens examined.—I have seen no examples of this species. Remarks.—This species also is placed by Bierig in the subgenus Acylophorus s. str. It is described as being closely related to A. inswl- anus and appears to differ principally in the aeneous luster. I find no record of its habits. 5. ACYLOPHORUS METALLICUS, new species Description —Black, pronotum rufopiceous, abdomen with strong aeneous luster. Head suborbicular; eyes about half as long as head; maxillary palpus with third and fourth segments somewhat enlarged, fourth acuminate; with a few large setiferous punctures around sides of disk and one above each eye, and with scattered punctulae almost obsolete; without ground sculpture. Pronotum slightly wider than long; with a pair of large discal punctures near middle and another pair at apical three-fourths, and a diagonal pair on each side near front angle; with obsolete punctulae as on head and without ground sculpture. Scutellwm punctured as base of elytra with only a very narrow smooth border. Hlytra with moderately coarse but submuri- 470 BULLETIN 182, UNITED STATES NATIONAL MUSEUM cate punctures often touching transversely and separated by twice their diameter or more longitudinally; without ground sculpture. Abdomen with setiferous punctures appearing to be coarsely tuber- culate, moderately dense; with strong luster. Length, 7 mm. Type locality—Dominican Republic, Sanchez. Types.—Holotype in the Museum of Comparative Zoology; one paratype in the United States National Museum (No. 52532); col- lected in July 1938 by Dr. P. J. Darlington. Records.—The following is the only record known to me: Hispaniola: DoMINICAN REpPuBLIc (Darlington, in M.C.Z. and U.S.N.M.). Specimens ecamined.—I have seen only the two types. Remarks.—This species is readily distinguishable by its strong luster, by the tuberculose abdominal punctures, and by the punctures of the pronotum. I have received no record of its habits. 6. ACYLOPHORUS DARLINGTONI, new species Acylophorus longicornis var. thaxteri BERNHAUER, MS. Description —Black, margin of apical segments rufous; pubescence: castaneous. Head nearly as broad as long, suborbicular; eyes a little less than half as long as head; fourth segment of maxillary palpus en- larged, fusiform or diagonally truncate; with a few large setiferous punctures around sides of disk and one above each eye; without dis- tinct punctulae or ground sculpture. Pronotom about one-ninth broader than long; with a pair of large discal punctures just before middle and a pair submarginal in front, and with one or two near the anterior angle; without punctulae or ground sculpture. Scutellum punctured as base of elytra with a distinct smooth border. “lytra with moderately coarse and strongly submuricate punctures, some- times uniting transversely, usually separated by twice their diameter longitudinally; without ground sculpture. Abdomen with setiferous punctures dense at base and sparse at apex of each segment, rather strongly submuricate; without ground sculpture. Length, 8 to 9 mm. Type locality —Trinidad, Cedros. Types.—Holotype and six paratypes in the Museum of Comparative Z oology; five paratypes in the United States National Museum (No. 52533) ; collected on May 4, 1929 by Dr. P. J. Darlington. Records.—The following are the records known to me: Trinidad: Cedros (Darlington, in M.C.Z.), St. Augustine (Darlington, in M.C.Z.), Four Roads (Thaxter, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen one example from the Museum of Comparative Zoology in addition to the types. Remarks.—In addition to its size this species may be recognized by the structure of the maxillary palpus. Ten of the paratypes were taken in “ditches in Brown’s cacao.” MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 471 LXXVI. Genus ATANYGNATHUS Jacobson Tanygnathus ERICHSON, 1888a, p. 417 (not Wagler). Tanygnatus EICHELBAUM, 1909, p. 159 (misspelling). Atanygnathus JACOBSON, 1909, p. 521. Tanygnathinus Re1TTER, 1909, p. 105. Genotypes—Tanygnathus terminalis Erichson (monobasic) (for all three names). Diagnosis —Head and pronotum without dense umbilicate punc- tures, margined beneath the eyes; antennae widely separated at front of head, not strongly geniculate, segments pubescent from the second joint; labrum transverse and densely hairy in front; mouth parts much elongate, palpi very long and slender, fourth segment of maxil- lary longer than the third, the labial similar to the maxillary; gular sutures separate; lateral margin of pronotum single; anterior coxal cavities entirely open behind; front coxae large, exserted; posterior coxae contiguous, “triangular,” with a small expansion over the base of femur; first and second abdominal sternites absent; femora un- armed except sometimes for a strong comb of setae at the base of the anterior ones; anterior tarsi 5-segmented, middle and _ posterior 4-segmented. Remarks.—This is the only American genus of the Quediini (or even of the Staphylininae) that does not have all the tarsi 5-segmented. It isa small genus but occurs all over the world. I have seen 82 examples from the West Indies. I assign these to two species, both previously described but one heretofore known only from the original description. KEY TO WEST INDIAN SPECIES OF ATANYGNATHUS 1. Elytral punctures exceedingly fine, somewhat indistinect________ 2. laticollis Hlytral punctures not’ very fine; distinet-=_ =o 1. heterocerus 1. ATANYGNATHUS HETEROCERUS Cameron Atanygnathus heterocerus CAMERON, 1922, p. 123 (as variety of antennalis Sharp) .—ScHEERPELTZ, 1933, p. 1468. Description —Head piceous, pronotum rufopiceous to rufous, elytra and abdomen castaneous to rufocastaneous. Head more or less strongly narrowed behind the eyes; not distinctly punctate except at sides, but with rather indistinct scaly ground sculpture. Pro- notum over one-fifth wider than long, completely rounded posteriorly and arcuately narrowed in front; with a single pair of discal punc- tures in front of middle; with ground sculpture as on head but less distinct. /ytra with moderate punctures separated by less than their diameter, bluntly and indistinctly submuricate. Abdomen with fine posteriorly excavated punctures, moderately sparse; without distinct ground sculpture. Length, 4 to 5 mm. 472 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Type locality.—Grenada. Types.—In the British Museum. Records.—The following are the records known to me: St. Vincent: (Cameron, 1922; British Museum; Cameron collection; U.S.N.M.). Grenada: (Cameron, 1922; British Museum; Cameron collection; U.S.N.M.; Blackwelder station 155.) Specimens examined.—I have seen from the West Indies 30 ex- amples in the British Museum, 6 in Dr. Cameron’s collection, 7 in the United States National Museum, and 1 collected by me in January 1936. (All but the last of these are from the original series.) Remarks.—This species appears to be distinct from laticollis by the punctation of the elytra. It was described by Dr. Cameron as a variety of antennalis Sharp but I have not been able to verify this. For the present I believe it best to consider it as a distinct species, although the characters mentioned by Dr. Cameron are quite insuffi- cient to distinguish it from antennalis. My specimen was found under stones along a small stream, 2. ATANYGNATHUS LATICOLLIS (Erichson) Tanygnathus laticollis FRIcHSON, 1839b, p. 289.—DLAcoRDAIRE, 1854, p. 60.— Lene and MutcHtrr, 1914, p. 405. Atanygnathus laticollis (Hrichson) BERNHAUER and SCHUBERT, 1916, p. 440. Description.—Piceous to rufopiceous, pronotum rufous, elytra and abdomen piceocastaneous or rufocastaneous. Head not narrowed behind the eyes but strongly triangular in front; without punctures except for one or two at sides, but with very indistinct ground sculp- ture. Pronotwm over one-fifth wider than long, completely rounded posteriorly and arcuately narrowed in front; with a single pair of discal punctures and a corresponding pair of anterior submarginal ones; with ground sculpture as on head. lyé¢ra rather finely and not very distinctly punctate; surface with irregular and indistinct ground sculpture. Abdomen finely and moderately densely sub- muricately punctate; with distinct but irregular ground sculpture. Length, 4 to 5mm. Type locality —“Amer. mer. ins. St. Vincentii.” Types.—Probably in the Hope Museum, Oxford. Records.—The following are the records known to me: Cuba: Soledad (Darlington, in M.C.Z. and U.S.N.M.), Rangel Mountains, Pinar del Rio (Darlington, in M.C.Z.), Hanabanillo Falls, Trinidad Mountains (Darlington, in M.C.Z.) Hispaniola: DoMInIcAN Repusiico, San José de las Matas (Darlington, in M.C.Z.), Sanchez (Darlington, in M.C.Z and U.S.N.M.), Puerto Plata (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: Mayaguez (Blackwelder station 50B). Antigua: (H. E. Box, as Blackwelder stations 4448, 444U). St. Vincent: (Hrichson, 1839b; Lacordaire, 1854; Blackwelder station 166). Trinidad: Maracas Falls (Weber, in M.C.Z. and U.S.N.M.). MONOGRAPH OF WEST INDIAN STAPHYLINIDAB 473 Specimens examined.—lI have seen 31 specimens from the Museum of Comparative Zoology and seven collected by me during 1935-37. Remarks.—Although I found no examples identified as this species in the British Museum or in Dr. Cameron’s collection, there is little doubt of the identity of the species. One example labeled wilis Sharp in Dr. Cameron’s collection seems to belong to this species, but I cannot state whether daticollis and vil7s may be the same. My examples were taken among the rotting “fruit” of an imported palm and under stones and sand along streams. KEY TO WEST INDIAN GENERA OF XANTHOLININI 1. Head and pronotum densely umbilicately punctate. LXXIX. Plochionocerus Head and pronotum at most sparsely punctate_-___________________--_ 2 2. Fourth segment of maxillary palpus much shorter than third, more or less ROTC UNG Saas OSes Eee aiid oA BEUTAEE a Ae Eh ee Ste 5 Fourth segment of aeuiaey palpus not or scarcely shorter than third, not TCL CU ale eee ee = Ree Se a Ee 8 i 3 3. Upper marginal line of pronotum joining lower line. LXXVIII. Thyreocephalus Upper marginal line of pronotum extending to or toward anterior angles with- OuL-UNItINe With LOWEN Tine hese so ee SE es sae ee 4 4. Fourth segment of maxillary palpus conical, slightly shorter than third. LXXX. Leptacinus Fourth segment of maxillary palpus distinctly longer than third. LXXVII. Xantholinus Da Ocnlarerrooves:onevande distinet===-2 =~ 22.20 =e ce eee 6 Ocular-eroovesmabsent: Of Inaitstin Coa as ae Be eer ee See 7 6 Gularsutures visible ithroughoutsit 22-224 4h eas LXXXITI. Stenolinus Gular sutures obsolete except in front___-_________-- LXXXIV. Oligolinus fepANLerioracansi, strong) y,,dilateds. 2s. ah oe ee LXXXII. Microlinus Anterior tarsi not much dilated. 2 2—~-=---22— LXXXI. Lithocharodes LXXVII. Genus XANTHOLINUS Dejean Xantholinus DEJEAN, 1821, p. 28. Subgenus Megalinus MULSANT and Rey, 1877), p. 45. Metacyclinus Retrrer, 1909, p. 137. Subgenus Wotolinus CAsry, 1906, p. 371. Subgenus /diolinus Casey, 1906, p. 375. Subgenus Notolinopsis Casry, 1906, p. 377. Subgenus Hypnogyra Casry, 1906, p. 394. Subgenus Calontholinus Retrrer, 1908, p. 114. Subgenus Milichilinus Reirrer, 1909, p. 137. Subgenus Typhlolinus Rerrrer, 1909, p. 188. Xantholinim WAwERKA, 1928, p. 33. Genotypes.—Paederus fulgidus Fabricius=Xantholinus fulgidus (Fabricius) (designated here). Of Metacyclinus, Staphylinus gla- bratus Gravenhorst=Xantholinus (Metacyclinus) glabratus (Gyra- venhorst) (designated here); of Milichilinus, Xantholinus decorus Erichson= Xantholinus (Milichilinus) decorus (Erichson) (mono- 449008—42 31 474 BULLETIN 182, UNITED STATES NATIONAL MUSEUM basic); of Typhlolinus, Xantholinus (Typhlolinus) hungaricus Reitter (monobasic); of Hypnogyra, Xantholinus gularis We- Conte=Xantholinus (Hypnogyra) gularis (LeConte) (designated here); of MNotolinopsis, N. capensis Casey (designated here); of Notolinus, N. fumipennis Casey (designated here); of Jdiolinus, Xantholinus crassicornis Hochhuth=J/diolinus crassicornis (Woch- huth) (monobasic); of Megalinus, Staphylinus glabratus Graven- horst=Xantholinus (Megalinus) glabratus (Gravenhorst) (mono- basic); of Calontholinus, Xantholinus (Calontholinus) fasciatus Hochhuth (monobasic). Diagnosis —Head and pronotum not densely umbilicately punc- tate; antennal grooves distinct, ocular grooves absent; antennae inserted close together at anterior margin of head, basal segment generally less than half as long as head, outer segments densely pu- bescent from the fourth joint; labrum small, feebly emarginate; fourth segment of maxillary palpus distinctly longer than third: last segment of labial palpus similar in shape to maxillary; gular sutures united or closely approximate in part; neck generally more than one-third as wide as head; pronotum with side margin double, the lines always separate; prosternum with a pair of large sclerites in front of the sternite; elytra with suture broadly beveled; middle coxae widely separated; posterior coxae contiguous, “triangular”: first and second abdominal sternites absent or represented only by tiny sclerites at the sides; intersegmental membranes of abdomen marked with a pattern of angular areas in longitudinal bands; anterior tarsi not much dilated. Remarks.—This genus is distinguished by the structure of the palpi and the separate marginal lines of the pronotum. It is a world- wide genus and as herein defined is more abundantly represented in the Western Hemisphere than was thought by Col. Casey. The authorship of this genus has apparently been erroneously cited for many years. In the Encyclopédie Méthodique (1825, p. 475) Serville gives a key to the groups of Staphylininae and obviously in- tends to propose the genus Xantholinus. However, he fails to give a Latin form to the name and also fails to cite any species. It is therefore impossible to accept his proposal of the genus, although this is the reference cited by all recent writers. The question of ac- cepting this use of the name is, however, not important since it was used at least twice at previous dates. In 1823 Dahl used the name at the head of a list of 8 species, of which 3 or 4 were valid names, and in 1821 Dejean used the name over a list of 21 species of which 9 were valid. Although there is no question that new specific names proposed in these lists are nomina nuda and not valid, there seems to MONOGRAPH OF WEST INDIAN STAPHYLINIDAE A75 be no reason for not accepting new generic names that were pro- posed for valid species. Accordingly Dejean becomes the author of Xantholinus wnless and until a previous validation of the name is found. The names 7'yphlodes Sharp and Vulda DuVal have been cited as subgenera of Yantholinus. The original descriptions of these ap- pear to me to contain characters of sufficient importance to separate them generically. I have examined 655 examples of this genus from the West Indies. These I assign to 16 species of which 11 appear to be new. One other species recorded from the West Indies is not recognized in the present collections. KEY TO WEST INDIAN SPECIES OF XANTHOLINUS 1. Head broader than long, broader than elytra (at least in male). 17. hydrocephalu Head longer than broad, not broader than elytra_____._______-___.____ 2 2. Pronotum entirely without discal punctures___-___--_-------__ 16. illucens Pronotum with few or many discal punctures_______-___----__________- 3 3. Head with additional fine postantennal grooves_____-____-----_-__-___ a Head without additional postantennal grooves_________________-_____ 10 4. Head and pronotum with distinct ground sculpture of lines____________ 5 Head and pronotum without’ ground seulpture.:.i.-__________________ 6 5. Center of vertex sparsely punctate; pronotum with 12 or 13 punctures in GISCAIRSE Ti Come 2 a he es eae LB ete De 14, jamaicensis Center of vertex almost impunctate; pronotum with 9 to 10 punctures in GOLSaTE SCLiCC tee a ae She EE a ee oe 15. impunctus 6. Center of vertex sparsely punctate or eyes separated from base of head by barely threestimes. theim Vength 22 2 2 a ee 7 Center of vertex impunctate and eyes separated from base by well over three EM ESRGhe live] Cm gees ee ee Pe ee ee ae eee 13. solitarius (oetilytraswithishumeri (distinctly «palerse) = lok ees eee 12. beattyi Hiv traywwithouc paleysnumertss 2s sa at ee es Se 8 or Heads) one-fourth) longer jthan)| wides_ 9220 = = ee 8 11. haitius Head. about. one-eighth longer than wide or less___________-.__________ 9 9. Pronotal series irregular but with numerous punctures_____-_-_ 9. audanti Pronotal series with not over seven punctures____-_------~- 10. morrisoni 10. Coarse punctures of head interspersed with distinct and dense DUNG Cua Case ee = ae Mee ee Se ee i ee 11 Coarse punctures of head not interspersed with punctulae____--__---~ 13 iieeedrd: wath distinet ground) sculptures2-2_—=----.2 = 12 Head “without distinct cround* sculptures 222 2S 6. cubensis Ely tral withidensersroundisculptwures = 2 eh eee 8. rulomus Elytra uneven but without distinct ground sculpture____-__~_ 7. darlingtoni Seyi Ae LO COLOL OM Si ears ota pee OS a ee ee 14 ES) GEARLOG Teer es eee See RA in) PR) ere ete 15 i eMost ofnelytralspunctures im series_ 2 == eee 1. oakleyi Ontyrouterspuncturestin seriessa2== See Sasa ee ee 2. insulatus 476 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 15. Elytra paler near apex (or outer apical angle) _--_-__________--___-__ 16 Hlytra,,paler near, base. J =. 62-2 ne Sees a ee ee et 5. humeralis 16; Antennal. grooves. punCltate. 2. _* 2-222 cea eee 17 Antennal Prooves impunctate 222 = _ 2 Sn eee 3. fuscipennis 17. Outer apical diagonal half of elytra testaceous____________________- a Apex: of-ely tra’ indefinitely paleni2 +220 See oes eee eee 4. attenuatus 1. XANTHOLINUS OAKLEYI, new species Description—Head black, pronotum and abdomen rufopiceous to rufocastaneous, elytra uniform castaneous or piceocastaneous. Head over one-sixth longer than wide, distinctly but narrowly cordate in- cluding mandible; antennal grooves strong, impunctate except for a single apical puncture; without additional post-antennal grooves or ocular grooves; with scattered umbilicate punctures of uneven size leaving a smooth median area; without ground sculpture. Pronotum three-eighths longer than wide, widest in anterior third, evenly nar- rowed posteriorly to rounded base; with median smooth area outlined by two series of nine punctures, with lateral series of six or seven punctures but almost no others; without ground sculpture. Elytra not very coarsely or densely punctate, nearly all the punctures ar- ranged in series; the intervals flat and indistinctly coriaceous. Ab- domen sparsely but not very finely submuricately punctate, with indefinite transverse strigulae. Length, 314 to 444 mm. Type locality—Puerto Rico, 18 miles east of Maricao. Types.—Holotype and paratype, U.S.N.M. No. 52434, collected by me on October 3, 1935; and one paratype from Villalba. Fecords.—The following are the records known to me: Hispaniola: DoMINIcAN RrepuBiic, Mount Diego de Ocampo (Darlington, in M.C.Z.). Puerto Rico: Maricao (Blackwelder station 47A), Villalba (Blackwelder sta- tion 62). Specimens examined.—I have seen only the three types and one example from the Museum of Comparative Zoology. Remarks.—This species is distinguished by the serial arrangement of the elytral punctures and the lack of extraserial punctures on the pronotum. The types were found in dung and manure. 2. XANTHOLINUS INSULATUS Cameron Xantholinus insulatus CAMERON, 1922, p. 115. Xantholinus insularis FAUVEL, MS.—CAmeERoNn, 1922, p. 115. p. 1131 (misspelling). (Not Bernhauer, 1904.) SCHEERPELTZ, 1933, Description —Rufopiceous, head black. Head one-eighth longer than wide, not enlarged behind the eyes, basal angles broadly rounded; antennal grooves rather feeble though not shortened; ocular grooves 71 One specimen taken in quarantine at New York City from cargo that originated in Haiti seems very distinct. The absence of reliable data prevents its description here. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 477 absent; with large and somewhat elongate umbilicate punctures at sides; without minute punctulae or ground sculpture. Pronotwm nearly one-fifth longer than wide, widest in anterior third, moderately narrowed posteriorly with sides emarginate at basal third ; with smooth midline marked by two series of about eight punctures, and with an irregular lateral series and a few scattered punctures; without ground sculpture. Hlytra with moderately coarse but generally not serial punctures, intervals flat. Abdomen sparsely and finely submuricately punctate, and with indistinct transverse strigulae. Length, 5 mm. Type locality —Grenada, Mount Gay Estate. Types —tIn the British Museum. Records.—The following are the records known to me: Grenadines: Mustique (Cameron, 1922; British Museum). Grenada: (Cameron, 1922; British Museum; Scheerpeltz, 1933). Specimens examined.—I have examined the type in the British Museum and one other example that was borrowed for further study. Remarks—This species has not been collected since its discovery. It is not strikingly distinct from related species but seems to be valid as set out in the key. I find no record of its habits. 3. XANTHOLINUS FUSCIPENNIS Cameron Xantholinus fuscipennis CAMERON, 1922, p. 115 (as variety of vilis Sharp).— SCHEERPELTZ, 1938, p. 1314 (as variety of pusillus Sachse). Description—Head black, pronotum and abdomen piceous, elytra piceous with apical third indefinitely testaceous. Head about one- sixth longer than wide, sides arcuate behind eyes and rounded into base; antennal grooves strong and straight, not distinctly punctate; ocular grooves absent; with large umbilicate punctures at sides and base, but vertex impunctate; without punctulate or distinct ground sculpture. Pronotum about three-eighths longer than wide, widest in front, sides narrowed to the rounded base but slightly emarginate behind middle; with smooth midline marked by two series of eight punctures, and with an irregular series of four or five laterally ; with- out ground sculpture. “lytra with moderately coarse and not dense punctures almost entirely in series; intervals rather flat. Abdomen moderately sparsely and rather finely submuricately punctate, and with indistinct transverse strigulae. Length, 4 to 5 mm. Type locality —Jamaica. Types.—tIn collection of Dr. Cameron. Records —The following are the records known to me: Jamaica: (Cameron, 1922; Scheerpeltz, 1933; Cameron collection), Santa Cruz (Blackwelder station 421), May Pen (Blackwelder station 425C), Bala- claya (Blackwelder station 397B), Trinityville (Blackwelder station 428), Black River (Blackwelder station 420), Kingston (Blackwelder station 1C), Milk River (Blackwelder station 415), Moneague (Blackwelder station 375). 478 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Specimens examined.—lI have seen 12 examples in Dr. Cameron’s collection and 15 collected by Chapin and Blackwelder in February 1937. Remarks.—This species was described as a variety of X. vilis Sharp. The West Indian specimens identified as v2/is in the British Museum are quite distinct, but I have not compared specimens of fuscipennis directly with the type of vilis. I believe, however, that it will prove to be a distinct species. Among West Indian species it is distinguished by its elytral coloration and the impunctate antennal grooves, Our examples were collected from dung and manure and flying at dusk, 4. XANTHOLINUS ATTENUATUS Erichson Xantholinus attenuatus ErR1cHson, 1839b p. 330.—Scrima, 1855, p. 299.—Duvat, 1857, p. 34—CHeEvRoLAT and FAuvet, 1863, p. 430.—Fauver, 1866, p. 350.— SHaArp, 1876, p. 203; 1885, p. 484—BrerkNHAUER and ScHusert, 1914, p. 301.— LENG and MuTcuHter, 1914, p. 406—Wotcort, 1924, p. 78; 1936, p. 197. Description.—Head black, pronotum and abdomen rufocastaneous, elytra rufescent with outer apical corner indefinitely testaceous. Head about one-fifth longer than wide, sides behind eyes feebly arcuate but head not widened posteriorly, basal angles marked by ridges at sides; antennal grooves strong, with one or more distinct punctures, especially near apex; ocular grooves absent; with large umbilicate punctures at sides, vertex almost impunctate except be- hind; with minute punctulae throughout but without ground sculpture. Pronotum over one-fifth longer than wide, widest at anterior angles, thence feebly and evenly narrowed to rounded base; with smooth midline marked by two series of six or seven punctures, and with an irregular lateral series and a few other punctures; without ground sculpture. //ytra with coarse but not dense punc- tures more or less serial, but irregular in size and spacing. Abdo- men sparsely and not coarsely submuricately punctate, and with indistinct transverse strigulae. Length, 4144 to 7 mm. Type locality—*Americae insulis St. Thomae et Puerto Rico... St. Vincentil,...-_ 2 eas ee ee 2 Hlytra:. at most-withymargin recuryvedi=— 2-2 ss) oe Aaa eee eee me 2. Pronotum relatively coarsely punctate_______--_--_--__---__-- 11. buscki Pronotum not;or minutely, punctate:= 2-23)" eee 3 3: Dlytra .distinetly punctates = 22 - ess See ee ee eee 10. rutilus Klytra not. or: minutely, punctave— 22a ee ee eee a 4. Apex of elytra and sides of pronotum paler________-___________-__-___ 5 Notas abovess: 22-5 2 Se See Se ee ee 6 5. Apical third of elytra and posterior margin of pronotum abruptly pale. 9. pulchellus Narrow margin of elytra indefinitely paler; base of pronotum not paler. 8. ebonus 6.-Pronotum: testaceous, strigulose2-se=-+-<- 3 = 2 eee 7. flavicollis Pronotum.piceow™ mot istrigulose. 222s: = ee eek eee 6. cacao MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 513 7. Elytra distinctly (sometimes irregularly) punctate____.____-__-_--_-___- 8 Hivirasonlyaveryeaminutely;punctulate. = ess" 2 2 ee ee eck 10 Sr CLONOLUMe a Stine] ye Sibi eulOsei a=. ee et ee ee ee 9 PronotumEnoe siniculosess== 2S sss ee ee 2. apicalis OF Dlyitawmstriou] oses eases oie ee eee ee 3. hepaticus i ytraenoeysinigulOses = ase SS Ne a 1. sharpi iO -=Pronotum-and? elytra; strigulose. 22. s=. = eee Ee 5. weberi Pronotumand ‘elytra; not: at «all istrigulose1 +22 *=s-+ = = 4, trinitatis 1. COPROPORUS SHARPI Cameron Coproporus sharpi CAMERON, 1922, p. 123—ScHEERPELTz, 1934, p. 1518. Description.—Head black, pronotum testaceous, sometimes clouded on the disk, elytra testaceous more or less clouded apically, abdomen testaceous generally clouded at base. Head not distinctly punctate but with minute punctulae and transverse strigulae. Pronotum not distinctly punctate but irregularly strigulose and with minute punc- tulae. Hlytra without concavity at sides except for a submarginal groove; distinctly punctate but the punctures irregular and more or less composite; without distinct strigulae. Abdomen with punctures obscured by irregular strigulae. Male, eighth sternite with a large triangular emargination about one-half wider than deep; eighth tergite with four blunt teeth separated by rounded notches. Female, eighth tergite with four slender but triangular processes, the middle a little longer and separated to base. Length, 114 to 214 mm. Type locality—St. Vincent. Types.—In the British Museum. Records.—The following are the records known to me: Antigua: (H. E. Box, as Blackwelder station 444Q). Montserrat: (Hubbard, in U.S.N.M.). St. Lucia: (Blackwelder station 226). St. Vincent: (Cameron, 1922; British Museum). Specimens examined.—I have seen two examples (including the fype) in the British Museum, eight in the United States National Museum, six collected by Dr. H. E. Box, and one collected by me in April 1936. Remarks.—This species is the smallest of those having the sides of the elytra unimpressed. It can also be distinguished by the irregular elytral punctation. I have seen one specimen from the Museum of Comparative Zoology, collected by Dr. W. M. Mann in Haiti, that keys out to this species. It is 8 mm. long but differs in no other respect I can find except color. I believe it to be a distinct species but will leave it till more specimens are available. My specimens were taken in cocoa pods and “from inside of scarlet bracts of a large Heliconia.” 514 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. COPROPORUS APICALIS (Erichson) Tachinus apicalis Er1cHson, 1889b, p. 250.—LacorparRE, 1854, p. 55.—DuVat, 1857, p. 33. Coproporus apicalis (Erichson) BERNHAUER and ScHUBERT, 1916, p. 489. Erchomus apicalis (Erichson) Lene and MurcuHtier, 1914, p. 407.—Wotcort, 1924, p. 79; 1936, p. 198. Description.—Piceous with margins pale to rufopiceous with pro- notum and elytra rufous. Head not distinctly punctate or strigulose but with very minute punctulae. Pronotwm not distinctly punctate or strigulose but with minute punctulae. Hlytra not flattened or con- cave at sides, except for a narrow stria along the upturned margin; with very distinct but rather irregular punctures separated by about twice their diameter; without strigulae. Abdomen not very distinctly punctate but irregularly strigulose. fale, eighth sternite with a large rounded triangular notch about one-half wider than deep; eighth tergite quadridentate, the middle pair of teeth acute and not widely separated. Female, eighth tergite with six slender lobes, the middle four much longer, the two middle deeply separated. Length, 134 to 214, mm. Type locality —Puerto Rico. Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records —The following are the records known to me: Cuba: Soledad (Leavitt, in M.C.Z. and U.S.N.M.), Loma del Gato, Oriente (Darlington, in M.C.Z.), Pico Turquino, (Darlington, in U.S.N.M.). Jamaica: Trinityville (Blackwelder station 428), Kingston (Blackwelder sta- tion 1D), Troy (Blackwelder station 409), Balaclava (Blackwelder station 403). Hispaniola: Haiti, northeast foothills of La Hotte Range (Darlington, in M.C.Z.), Ennery (Darlington, in M.C.Z. and U.S.N.M.), Camp Perrin (Dar- lington, in M.C.Z.), Etang Lachaux (Darlington, in M.C.Z.) ; Dominican Repusiic, Sinchez (Darlington, in M.C.Z. and U.S.N.M.), foothills south of Santiago (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: (EHrichson, 1839b, DuVal, 1857). Specimens examined.—I have seen 16 examples from the Museum of Comparative Zoology and 6 specimens collected by me during 1935-37, Remarks.—This species is distinguishable by its punctate elytra, smooth pronotum, and unimpressed elytral margin. My examples were taken on rotting mango fragments and flying at dusk. 3. COPROPORUS HEPATICUS (Erichson) Tachinus hepaticus Er1cHson, 1839b, p. 249.—LAcorDATRE, 1854, p. 55. Tachinus converus ERICHSON, 1839b, pp. 248, 249.—LAcorDAIRE, 1854, p. 55.— FAUVEL, 1863, p. 429. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 515 Coproporus converus (Erichson) FAuveL, 1863, p. 429.—LENa and MUTCHIER, 1914, p. 407.—BERNHAUER and ScHUBERT, 1916, pp. 489, 490.—SCHEERPELTZ, 1933, p. 1516.—BLACK WELDER, 1938, p. 8. Coproporus ignavus SHARP, 1876, p. 87; 1883, p. 305.—BERNHAUER and SCHUBERT, 1916, p. 490.—BLACKWELDER, 1938, p. 8. Erchomus inflatus Horn, 1877, p. 107.— HENSHAW, 1898, p. xliii—FaLL and COCKERELL, 1907, p. 153.—LENe, 1920, p. 111 (as synonym of converus).— BLACKWELDER, 1988, p. 7. Erchomus ignavus (Sharp) SHarp, 1883, p. 305. Erchomus converus (Hrichson) SHarp, 1883, p. 8306.—HENSHAwW, 1898, p. xliii— HUBBARD, 1899, p. 2 (suppl.).—LENe, 1920, p. 111. Erchomus hepaticus (Hrichson) Viiiapa, 1901, p. 28. Ooproporus inflatus (Horn) BERNHAUER and ScHUBERT, 1916, p. 490.—SCHEER- PELTZ, 1933, p. 1516.—BLACK WELDER, 1938, p. 7. Coproporus hepaticus (Erichson) SCHEERPELTz, 1934, p. 1516. Description.—Black with abdomen somewhat rufescent (frequently largely rufescent, especially on the elytra). Head not distinctly punc- tate but with minute punctulae and distinct strigulae. Pronotum not distinctly punctate but distinctly strigulose and with minute punctulae. Hlytra vaguely flattened but not concave at sides, except for a narrow stria along the upturned margin; with fine and irregular but distinct punctures; transversely strigulose. Abdomen with coarse but rather indefinite punctures somewhat obscured by irregular strigulae. Male, eighth sternite with a very large triangular emargi- nation nearly as deep as wide, the sides sinuate and the apical angle acuminate; eighth tergite quadridentate, the teeth longer than usual and more slender, the middle pair nearly twice as long as outer. Female, eighth tergite with four slender processes, the middle pair a little longer and deeply separated. Length, 214 to 314 mm. Type locality —Colombia. Ofignavus, Anana, Amazons, Brazil; of convexus, Brazil; of inflatus, Camp Grant, Ariz. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of convewus, either in the Zoologische Museum, Berlin, or the Natural History Museum, Geneva; of inflatus, in the Academy of Sciences of Philadelphia; of ignavus, in the British Museum. Records.—The following are the records known to me: Cuba: (Fauvel, 1863, as converus), Cayamas (Schwarz, in U.S.N.M.), San Antonio de los Banos (Pazos, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: Cayey (Cotton, in U.S.N.M.), Bayamon (Lesesne, in U.S.N.M.), San Juan (Blackwelder station 44). Trinidad: Manzanilla (Blackwelder station 104A). South America: Cotompra (Hrichson, 1839b), Braz (Sharp, 1883, as ignavus; Erichson, 1839b, as converus; Fauvel, 1863, as converus; British Museum), ARGENTINA, PARAGUAY (Scheerpeltz, 1933), ARGENTINA (British Museum). Central America: Mexico, GUATEMALA, PANAMA (Sharp, 1883, as ignavus), MExico (Fauvel, 1863, as converus ; U.S.N.M.), MExico (Villada, 1901). 516 BULLETIN 182, UNITED STATES NATIONAL MUSEUM North America: ArizoNA (Horn, 1877, as inflatus), ARIZONA, TEXAS, CALIFORNIA (Blackwelder, 1938, as inflatus; U.S.N.M.), Arizona (Hubbard, 1899, as convenus). Specimens examined.—From the West Indies I have seen 10 ex- amples in the United States National Museum, 6 from the Museum of Comparative Zoology, and 8 collected by me in 1935-1937. In addi- tion I have seen numerous examples from North, Central, and South America, including the types of ignavus and inflatus. Remarks.—This species can be distinguished by its size and the un- impressed sides of the elytra. The northern specimens (Cuba, Puerto Rico, Mexico, North America) seem to be uniformly paler than the Trinidad or South American ones, though this may be due in part to the age of the specimens. I can find no other characters to separate them. The Puerto Rican examples were collected “under bark of Bucare tree,” the Cuban ones “on ceiba” (silk cotton tree), and the Trinidad ones in very old cocoa pods. 4. COPROPORUS TRINITATIS, new species Description.—Rufous, sometimes very feebly picescent in part. Head not distinctly punctate but with minute punctulae; without trace of ground sculpture. Pronotwm not distinctly punctate but with very minute punctulae; without trace of strigulae. /lytra rather flattened at the sides but not concave, except for a narrow stria along the upturned margin; not distinctly punctate or strigulose but with minute punctulae. Abdomen distinctly but rather irregularly punc- tate, with traces of strigulae. Male, eighth sternite with a large some- what rounded triangular emargination about one-third wider than deep; eighth tergite quadridentate, the middle pair separated only half way to base. Female, eighth tergite with six slender lobes, the middle four nearly equal in length but the two middle more triangular. Length, 134 to 3 mm. Type locality.—Trinidad, the Tacarigua River, 2 miles north of Tacarigua or 4 miles northeast of St. Augustine. T'ypes.—Holotype, male, and four paratypes, U.S.N.M., No. 52454, collected by me on December 22, 1935. Records.—The following is the only record known to me: Trinidad: Tacarigua (Blackwelder station 107A). Specimens examined.—I have seen only the five types. Remarks.—This species can be distinguished by its entirely impunc- tate and smooth head, pronotum, and elytra. It resembles rvtilus but differs in lacking the elytral concavity. The types were found under stones at the river’s edge. Pe MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 517 5. COPROPORUS WEBERI, new species Description—Dark rufopiceous, head black, apex of abdomen rufous; sometimes rufous throughout. Head not punctate, but with a few minute punctulae obscured by the distinct transverse strigulae. Pronotum with punctulae and sculpture as on head, sometimes obso- lescent. Hlytra rather flattened at sides but not distinctly concave, except for a narrow stria along the margin; with very minute punc- tulae and irregular transverse strigulae. Abdomen with small punc- tures much obscured by irregular strigulose sculpture. dale, eighth sternite with an equilateral triangular emargination, the angles pro- longed as parallel ridges, bearing coarse setae at the apices; eighth tergite quadridentate, the processes of same shape, size, and separa- tion, the middle pair more posterior. Female, eighth tergite with four slender lobes, equally separated though middle pair not so deeply, middle pair a little more slender and longer. Length, 2144 mm. Type locality.—Trinidad, St. Augustine. Types——Holotype, male, and seven paratypes, in the Museum of Comparative Zoology; six paratypes in the United States National Museum (No. 52542) ; collected in July 1935 by N. A. Weber. Records.—The following are the records known to me: Trinidad: Maracas Falls (Weber, in M.C.Z.), St. Augustine (Weber, in M.C.Z. and U.S.N.M.), foothills north of Tunapuna (Weber, in M.C.Z. and U.S.N.M.). Specimens examined.—I have seen only the 14 types. Remarks.—TVhis species is very distinct by the strong strigulae of all parts of the dorsal surface. AJ] others of our species having these strigulae throughout have the elytra concave at the sides. The strigulae are sometimes fainter on the pronotum but those on the elytra are enough to distinguish the species. The flattening at the sides of the elytra may at times seem to attain convexity. In these cases the specimens would key out to flavicollis and cacao. From the former it is distinguished by its dark pronotum; from the later by its size and the sparse strigulae of the pronotum, I have received no record of the habits of this species. 6. COPROPORUS CACAO, new species Description.—Piceous, sometimes slightly rufescent (or even testa- ceous). Head not punctate but exceedingly minutely punctulate and with scarcely a trace of strigulae. Pronotwm not distinctly punctate but minutely punctulate, without strigulae (although iridescent). Elytra with a lateral concavity formed principally by the broadly upturned margin; not distinctly punctate but with punctulae dis- tinct; sometimes at lateral third and along apical margin with fine 518 BULLETIN 182, UNITED STATES NATIONAL MUSEUM short diagonal grooves; with rather feeble transverse strigulae. Abdomen with punctures rather obscured by irregular strigulae. Male, eighth sternite with a narrow triangular emargination scarcely as wide as deep; eighth tergite quadridentate, the middle pair much longer and not widely separated. Female, eighth tergite with four slender processes, the middle pair much longer. Length, 1 to 144 mm. Type locality —St. Lucia, DuBoulay Estate, district of St. Remy, 3 miles southeast of Soufriére. Types.—Holotype, female, U.S.N.M. No. 52453, collected by me on April 23, 1936. Records.—The following are the records known to me: St. Lucia: (Blackwelder stations 220B, 220G). Trinidad: Montserrat (Busck, in U.S.N.M.). Specimens examined.—Besides the type I have seen one specimen in the United States National Museum and one collected by me in May 1936. Remarks.—This species can be distinguished by the combination of characters given in the key. My second example from St. Lucia is entirely testaceous but is otherwise similar to the type. The Trini- dad example lacks the diagonal grooves of the elytra, but this does not appear to be of great importance. My specimens were found in rotting grapefruit and among fer- menting chips on a fresh stump. 7. COPROPORUS FLAVICOLLIS Scriba Tachinus nitidulus FERIcHSON, 18389b, p. 247.—LAcorDAIRE, 1854, p. 55. (Not Fabricius, 1781.) Coproporus flavicollis ScrrBa, 1855, p. 297.—BERNHAUER and ScHUBERT, 1916, p. 491. Erchomus strigosus SHARP, 1883, p. 299. Erchomus nitidulus (Erichson) Lena and MutTcHter, 1914, p. 407—Wotcort, 1924, p. 79; 1936, p. 198. Coproporus nitidulus (Erichson) BeRNHAUER and ScHuBeRT, 1916, p. 491. Coproporus strigosus (Sharp) BERNHAUER and ScHUBERT, 1916, p. 492. Description—Head and pronotum testaceous, elytra piceous to piceocastaneous with apex paler, abdomen piceous with apex piceo- testaceous. ead without punctures or distinct punctulae; with dis- tinct but irregular strigulae. Pronotum not distinctly punctate but with minute punctulae and distinct transverse strigulae. /7ytra with a distinct lateral concavity formed principally by the broadly up- turned margin; not distinctly punctate but with rather indistinct punctulae; with distinct transverse strigulae. Abdomen not clearly punctate but with very irregular strigulae. Male, eighth sternite with a narrow triangular emargination scarcely as wide as deep. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 519 Female, eighth tergite with four processes, the middle pair longer and more slender. Length, 1 to 1144 mm. Type locality —Venezuela. Of nitidulus, Puerto Rico; of strigosus, Guatemala. Types.—Presumably in the Senckenburg Museum, Frankfurt-am- Main. Of nitidulus, either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin; of s¢vigosus, in the British Museum. Records.—The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: Spanish Town (Blackwelder station 377). Puerto Rico: (Erichson, 1839b, as nitidulus). South America: VENEZUELA (Scriba, 1855). Central America: MExico, GUATEMALA (Sharp, 1883, as strigosus), Mrxico (Fenyes, in U.S.N.M.), Panama (Schwarz, in U.S.N.M.). Specimens examined.—I have seen one specimen from the West Indies in the British Museum, four from the Museum of Comparative Zoology, one in the United States National Museum, and two col- lected by me during 1935-87. Four examples from Central America were found in the United States National Museum, and the type of strigosus was examined in the British Museum. Remarks.—This species may be distinguished by its testaceous head and pronotum. Central American examples which I compared with the type of strigosus are indistinguishable from my West Indian specimens. The above description was drawn from my Jamaican examples. My specimens were caught flying at dusk. 8. COPROPORUS EBONUS, new name Tachinus piceus WrRicHson, 1889b, pp. 246, 250.—LAcorDAIRE, 1854, p. 55. (Not Stephens, 1829.) Coproporus piceus (Erichson) BERNHAUER and ScHUBERT, 1916, p. 491.—SAtrt, 1929, pp. 485, 450, 464.— BERNHAUER, 1934, p. 217. Erchomus piceus (Erichson) Lena and MurcHter, 1914, p. 407.—Wotcorr, 1924, p. 79; 1936, p. 198. Description.—Piceous, pronotum, and abdomen more or less rufes- cent, sides of pronotum and apex of elytra narrowly and indefinitely paler. Head very minutely punctulate and with traces of strigulae, but shining. Pronotum not distinctly punctate but with minute punctulae; with traces of strigulae as on head. Llytra with a dis- tinct lateral concavity formed principally by the broadly upturned margin; punctulae a little more distinct than on the head; strigulae more distinct than on the pronotum. Abdomen with distinct but very irregular strigulae obscuring any punctures. Jale, eighth ster- nite with an abrupt triangular emargination as deep as wide; eighth 520 BULLETIN 182, UNITED STATES NATIONAL MUSEUM tergite quadridentate, the middle teeth longer and narrower. Fe- male, eighth tergite with four slender processes, the middle pair longer. Length, 1144 to 2 mm. Type locality.—Puerto Rico. Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Jamaica: Moneague (Blackwelder station 370), Porus (Blackwelder station 423). Puerto Rico: (Erichson, 1839b; Wolcott, 1924, 19386), Adjuntas (Blackwelder station 48C). St. Lucia: (Blackwelder stations 220K, 224, 226, 227). Grenada: (Blackwelder station 153A). Specimens examined.—I have seen 5 examples in the British Mu- seum and 18 collected by me during 1935-87. Remarks.—This species is very similar to pulchellus but apparently can be distinguished by the smaller size of the pale areas as well as by the male characters. Both these species vary somewhat in the former character and might be considered to be a single species ex- cept for the distinct difference in shape of the emargination of the eighth sternite. This description was drawn from an example from St. Lucia. My specimens were found in rotting cocoa pods, under-chips on fresh stumps, in dung, and under bark. 9. COPROPORUS PULCHELLUS (Erichson) Tachinus pulchellus Er1cHson, 1839b, p. 247.—LAcorDAIRE, 1854, p. 55. Coproporus cumanensis Scripa, 1855, p. 297.—BeRNHAUVER and SCHUBERT, 1916, p. 492. Tachinus infimus DuVAL, 1857, p. 33.—CHEvRoLAT and FAvUvEL, 18638, p. 480.— BLACKWELDER, 1938, p. 9. Coproporus infimus (DuVal) CHEyROLAT and FAuvvEL, 18638, p. 430..—BERNHAUER and ScHUBERT, 1916, p. 490.—BLACKWELDFR, 1938, pp. 2, 9. Coproporus distans SHARP, 1876, p. 92; 1888, p. 299.—BrRNHAUER and ScHU- BERT, 1916, p. 492. Erchomus distans (Sharp) SHarp, 1883, p. 299, 3800 Erchomus infimus (DuVal) Lene and Murcuter, 1914, p. 407.—LeEna, 1920, p. 111.— BLACKWELDER, 1938, p. 9. Ooproporus pulchellus (EHrichson) BERNHAUER and SCHUBERT, 1916, p. 492.— CAMERON, 1922, p. 123. Description—Piceous, pronotum generally somewhat rufescent with margins testaceous, elytra rufocastaneous with apical third testaceous. Head not distinctly punctate but with irregular and indistinct strigulae and minute punctulae. Pronotwm not distinctly punctate but with minute punctulae and traces of strigulae. lytra with a distinct lateral concavity formed in part by the upturned mar- gin; not very distinctly punctate but with minute punctulae and MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 521 traces of transverse strigulae. Abdomen with distinct but very ir- regular strigulae obscuring any punctures. fale, eighth sternite with an abrupt triangular emargination over twice as wide as deep; eighth tergite bluntly quadridentate, the middle teeth rather widely sep- arated. Female, eighth tergite with four slender processes, the inner pair longer and more slender than the outer. Length 134 to 2 mm. Type locality—*In Puerto Rico et in Colombia.” Of infimus, Cuba; of distans, Rio Purus, Amazons, Brazil; of cwmanensis, Vene- zuela. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of cumanensis, probably in the Senckenburg Mu- seum, Frankfort am Main; of déstans, in the British Museum; of infimus, either in the Muséum National d’Histoire Naturelle, Paris, or the British Museum, London. Records—The following are the records known to me: Cuba: (Fauvel, 1863), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.), mountains north of Imias, Oriente (Darlington, in M.C.Z.). Jamaica: (Hubbard, in U.S.N.M.), Troy (Blackwelder station 409), Manchioneal (Blackwelder station 9B), Spanish Town (Blackwelder station 377), Trin- ityville (Blackwelder station 428), May Pen (Blackwelder station 425C), Milk River (Blackwelder station 415), Santa Cruz (Blackwelder station 421), Whitfield Hall (Darlington, in M.C.Z, and U.S.N.M.). Hispaniola: Harri, Port-au-Prince (Audant, in U.S.N.M.) ; DOMINICAN REPUBLIC, Puerto Plata, (Darlington, in M.C.Z.), Mount Quita—Espuela (Darlington, in U.S.N.M.). Puerto Rico: (Erichson, 1839b; U.S.N.M.). Antigua: (Blackwelder station 277A; H. E. Box, as Blackwelder station 4440). Dominica: (Blackwelder stations 285B, 237, 248A, 251, 252, 257C). St. Lucia: (Blackwelder stations 220B, 220C, 221, 224, 226, 227, 230, 2381; H. E. Box, as Blackwelder station 444B). St. Vincent: (U.S.N.M.; British Museum). Grenada: (U.S.N.M.; British Museum), Blackwelder stations 182, 137, 153A). Trinidad: Sangre Grande (Blackwelder station 100A). South America: Cotompra (Hrichson, 1839b), Brazin (Sharp, 1883), VENEZUELA (Seriba, 1855, as cumanensis). Central America: NICARAGUA, GUATEMALA (Sharp, 1883), Mexico (British Museum). North America: FrLoripa (Leng, 1920; Blackwelder, 1938). Specimens examined—¥rom the West Indies I have seen 295 examples in the British Museum, 8 from the Museum of Compara- tive Zoology, 22 in the United States National Museum, and 1907 collected by me during 1935-37. Remarks.—This is the commonest West Indian Coproporus and is equally abundant in Central America. It can generally be dis- tinguished by its small size and its coloring, but from ebonus it is 449008—42——34 522 BULLETIN 182, UNITED STATES NATIONAL MUSEUM separated by rather unsatisfactory characters as mentioned under that species. My specimens were taken from rotting citrus fruit, from decaying cocoa pods, in freshly cut chips of coconut husks, under chips on fresh stumps, from fungus, from manure, in decomposing trash in sugarcane fields, among decomposing forest debris, and flying at dusk. 10. COPROPORUS RUTILUS (Erichson) Tachinus rutilus ERICHSON, 1839b, p. 253.—SHArRP, 1883, p. 804.—BLACKWELDER, 1938, pp. 2, 4. Tachinus terminalis ErtcHson, 1889b, p. 250.—LAcorDATRE, 1854, p. 55.—CHEVRO- LAT and FAUVEL, 1863, p. 430.—DvuVAaL, 1857, p. 32. Tachyporus brevis Scrrpa, 1855, p. 296. —BLACKWELDER, 1938, pp. 4, 5. Coproporus brevis (Scriba) BERNHAUER and ScHusertT, 1916, p. 492. (Not Sharp, 1876.) Coproporus terminalis (Hrichson) CHEvROLAT and FauvatL, 1863, p, 429.—LENG and MutTcHter, 1914, p. 407.—BrrNHAUER and ScHusert, 1916, p. 493.— Woxcort, 1924, p. 79; 1936, p. 198. Erchomus rutilus (Erichson) SHARP, 1883, p. 804.—WotcorT, 1936, p. 198.— BLACKWELDER, 1988, p. 4. Cilea rutilus (Erichson) Lene and MourcuHter, 1914, p. 406—Wotcorr, 1924, p. 79; 1936, p. 198 —BLACKWELDER, 1938, p. 4. Coproporus rutilus (Erichson) BrrNaavEerR and ScHupert, 1916, p. 492.— BreRNHAUER, 1918, p. 91.—SCHEERPELTZ, 1933, p. 1518—BrERNHAUER, 1934, p. 218.—Wotcort, 1936, p. 198.—BLACKWELDER, 1938, pp. 2, 3, 4. Coproporus obscurior BERNHAUER, 1918, p. 92 (as aberration).—ScHEERPELTZ, 1934, p. 1518. Coproporus piceorufus BERNHAUER, 1918, p. 92 (as aberration).—ScHEERPELTZ, 1934, p. 1518. Description —Clear rufous to rufopiceous. Mead minutely punc- tulate but not otherwise punctate or sculptured. Pronotum not dis- tinctly punctured or sculptured, but sometimes with one or two rather large punctiform depressions at the sides of the disk. Hlytra with a large and distinct depression at the sides but slightly separated from the margin; rather finely but very distinctly punctured, the intervals smooth and shining. Abdomen punctured similarly to the elytra but a little more finely and with irregular strigulae among the punctures. JM/ale, eighth sternite with a large abrupt triangular emargination about one-third wider than deep; eighth tergite quadridentate, the median teeth longer and more acute, the median notch rather broad. Female, eighth tergite divided into four pro- cesses, the two middle more slender and a little longer than the outer. Length, 144 to 2 mm. Type locality —“*Americae ins. Puerto Rico et St. Thomae, in Colum- bia.” of brevis, Venezuela; of obscurior, Panama; of piceorufus, Argentina and Brazil; of terminalis, Puerto Rico. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 523 Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of brevis, probably in the Senckenburg Museum, Frankfurt-am-Main; of obseurior and piceorufus, presumably in the collection of Dr. Max Bernhauer; of terminalis, with rutilus. Records —The following are the records known to me: Cuba: (Fauvel, 1863; DuVal, 1857; as terminalis; Wright, in M.C.Z.), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S.N.M.). Jamaica: Trinityville (Blackwelder station 428). Hispaniola: Hariri, Sources Puantes (Darlington, in M.C.Z.), Port-au-Prince (Darlington, in U.S.N.M.); Dominican Repusuic, Sanchez (Darlington, in M.C.Z.). Puerto Rico: (Erichson, 1839b, as both; DuVal, 1857, as terminalis; Fauvel, 1863, as terminalis; Sharp, 1883), Ponce (R. G. Oakley, in U.S.N.M.). St. Thomas: (Erichson, 1839b; Sharp, 1883). St. Croix: (Blackwelder stations 341, 344). Montserrat: (Hubbard, in U.S.N.M.; Blackwelder station 269). Dominica: (Blackwelder stations 251, 252). St. Vincent: (British Museum). Grenadines: Mustique (U.S.N.M.; British Museum). Grenada: (U.S.N.M.; British Museum). Trinidad: Port of Spain (Blackwelder station 115), St. Augustine (Weber, in M.C.Z. and U.S.N.M.), foothills north of Tunapuna (Weber, in M.C.Z. and U.S.N.M.). South America: Cotompia (Erichson, 1839b; Sharp, 1883), Brazin (Sharp, 1883), ARGENTINA, Braz, (Bernhauer, 1918, as piceorufus), WENEZUELA (Seriba, 1855, as brevis). Central America: Mexico, BriTIsH HONDURAS, GUATEMALA, NicARAGUA (Sharp, 1883), PANAMA (Bernhauer, 1917, as obscurior). North America: Texas (Blackwelder, 1938). Specimens examined.—F rom the West Indies I have seen 124 exam- ples in the British Museum, 24 from the Museum of Comparative Zoology, 18 in the United States National Museum, and 11 collected by me during 1935-37. Remarks.—In general this species is very distinct by its bright red color from all except trinitatis. From this it differs in having the definite lateral impression of the elytra. However, the specimens I have identified as terminalis are darker, although the pronotum is still lighter than the rest of the body. I can find no way to separate these specimens or the original descriptions. The above description was taken from specimens from Dominica. My specimens were found under seaweed on the beach and, more commonly, flying at dusk. 11. COPROPORUS BUSCKI, new species Description.—Piceous, elytra somewhat rufescent at base. Head rather irregularly but distinctly punctate; surface minutely uneven but not distinctly strigulose. Pronotuwm distinctly but somewhat irregularly punctate; surface irregularly and indistinctly strigulose. 524 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Elytra with a large and deep depression along the lateral margin; a little more coarsely punctate than pronotum; surface uneven but not strigulose. Abdomen not very distinctly punctate but with irregular strigulae. Male, eighth sternite with a large triangular emargina- tion about one-half wider than deep, the angles narrowly rounded; eighth tergite quadridentate, the median pair of teeth longer and separated by a notch of same size and shape as the teeth. Female, eighth tergite with four slender processes, the inner pair a little longer than the outer. Length, 224 mm. Type locality —Trinidad, ward of Montserrat. Types.—Holotype, male, and two paratypes, female, U.S.N.M. No. 52452, collected on July 4, 1905, by August Busck. Records.—The following is the only record known to me: Trinidad: Montserrat (Busck, in U.S.N.M.). Specimens examined.—I have seen only the three types. Remarks.—This species is distinguished by the relatively coarse punctures of the pronotum. Its robust size will also serve to sep- arate it from the species with impressed elytra. I have no record of its habits. LXXXVII. Genus CONOSOMUS Motschulsky Conurus STEPHENS, 1829, p. 272. (Not Kuhl, 1820.) Conosomus MoTscHuULsky, 1857b, p. 54. Conosoma KRAatz, 1857a, p. 431. Canurus KRAATZ, 1874, p. 292 (misspelling). Genoty pes.—Staphylinus littoreus Linnaeus= Conosomus littoreus (Linnaeus) (designated here). Of Conurus, Tachyporus pubescens Gravenhorst=Conurus pubescens (Gravenhorst) (designated here); of Conosoma, Staphylinus littorevs Linnaeus=Conosoma littorea (Linnaeus) (designated here). Diagnosis —Head inclined, not margined at the sides beneath; hypostomal suture and anterior part of coronal suture distinct ; antennae 11-segmented, inserted at sides of head; labrum trans- verse; maxillary palpus subulate; gular sutures separate; anterior coxal cavities closed behind by the union of the prosternum and hypomera beneath the coxae; front coxae large, exserted; posterior coxae contiguous, “transverse”; abdomen not margined, first and second sternite absent; tibiae fimbriate at tip with equal spinules; tarsi 5-segmented, first segment of posterior tarsus short. Remarks.—The two names Conosomus and Conosoma were pub- lished in the same year (1857) apparently entirely independently as new names for the preoccupied Conurus Stephens. Conosoma has subsequently been used almost exclusively, with Conosomus only MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 525 rarely cited even as a synonym. However, in 1860 (1860a, p. 575) Motschulsky wrote that his name was older than that of Kraatz and that Kraatz had improperly credited the name to himself. In view of this it appears that Motschulsky’s name is the older and is to be used for the genus, and the Kraatz name may reasonably be considered an emendation or misspelling. This genus is world-wide in distribution and is well represented in North and South America. It is readily recognized by its con- vex tapering form and dense vestiture. Five species have been described from the West Indies but I am unable to distinguish them. I have therefore considered them to be only one species. 1. CONOSOMUS INTERRUPTUS (Erichson) Conurus interruptus ERIcHSon, 1889p, p. 225. Conosoma interruptum (Erichson), BERNHAUER and SCHUBERT, 1916, p. 468, Conosoma vitraci BERNHAUER, 1918, p. 50.—ScHEERPELTZ, 1933, p. 1500. Conosoma caribeanum Birrta, 1937, p. 276, figs. 4, 5. Conosoma cubanum Bteric, 1937, p. 277, fig. 6. Conosoma conforme Brrria, 1937, p. 278, fig. 7. Description —Piceocastaneous, basal spot on elytra and apex of abdomen more or less rufescent. Head transverse, inserted into prothorax almost to the eyes; frontal area testaceous; antennae slender at base, apical segment variable in shape; with rather fine and irregular punctures between the eyes. Pronotum strongly con- vex, nearly one-third wider than long, sides evenly rounded; with very fine and vaguely submuricate punctures throughout; surface shining. /lytra with punctures similar to those of pronotum but somewhat obscured by irregular ground sculpture. Abdomen with very fine punctures rather obscured by fine and irregular ground sculpture, the punctures more distinctly submuricate posteriorly. Vestiture very conspicuous throughout. Length, 114 to 3 mm. Type locality—‘*Valle Araguensi Columbiae.” Of vitraci, Guade- loupe; of caribeanum, cubanum, and conforme, Cuba. Types.—Kither in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Of witraci, in the Bernhauer collection; of cari- beanum, cubanum, and conforme, in the Bierig collection. One “cotype” of conforme is in the United States National Museum. Records.—The following are the records known to me: Cuba: Sierra del Rosario, El Rangel (Bierig, 1937), Sierra de Bonilla, Cueva del Indio (Bierig, 1937), Pico Somorrostro (Bierig, 1937), Aspiro (Bierig, 1937), Cayamas (Schwarz, in U.S.N.M.), Soledad (Darlington, in M.C.Z. and U.S. N. M.), Pico Turquino (Darlington, in M.C.Z. and U. S. N. M). Jamaica: Bath in St. Thomas (Blackwelder station 387A). 526 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Hispaniola: Harti, Roche Croix (Darlington, in M.C.Z.) ; DOMINICAN REPUBLIC, Valle Nuevo (Darlington, in M.C.Z and U.S.N.M.), Constanza (Darlington, in M.C.Z. and U.S.N.M.). Puerto Rico: El Yunque (Darlington, in M.C.Z. and U.S.N.M.). Montserrat: (Hubbard, in U.S.N.M.). Guadeloupe: (Bernhauer, 1918, as vitraci; Busck, in U.S.N.M.). St. Vincent: (British Museum). Trinidad: Tacarigua (Blackwelder station 107C), Guepo Bay (Darlington, in M.C.Z.). South America: Cotomsra (EHrichson, 1839b). Specimens examined.—I have seen 2 examples in the British Museum, 18 from the Museum of Comparative Zoology, 15 in the United States National Museum, and 5 collected by me during 1935-37. Remarks.—\ am unable to distinguish the species recently named by Bierig from Cuba or that of Bernhauer from Guadeloupe. The characters used seem to me to be variable and all extremes are to be found in the series before me. I have hesitated to reduce to synonymy the two species I have not seen (cubanus and caribeanus) but find no other procedure tenable on the information at hand. My examples were taken from under moss and dirt on a rock beside a stream and from fungus (Hydnum sp.). LXXXVIII. Genus BRYOPORUS Kraatz Bryoporus KRraatz, 1857a, p. 452. Subgenus Bryophacis Retrrer, 1909, p. 102. Genotypes.—Tachinus cernuus Gravenhorst = Bryoporus cernuus (Gravenhorst) (designated here). Of Bryophacis, Bolitobius rufus Erichson = Bryoporus (Bryophacis) rufus (Erichson) (designated by Tottenham, 1939.) Diagnosis —Head inclined, strongly margined beneath the eyes; antennae inserted at sides in front of eyes, 11-segmented, outer seg- ments pubescent from the fourth joint; hypostomal suture present, coronal suture absent; labrum rather large, subquadrate; palpi stout, fourth segment of maxillary about as long as third, conical; gular sutures widely separated; anterior coxal cavities entirely open behind; front coxae very large, exserted; posterior coxae contiguous, “trans- verse,” expanded laterally and caudally under the femora; first and second abdominal sternites absent; tarsi 5-segmented, first segment of posterior moderate or short. Remarks.—This small genus is structurally very close to Bolitobius but can be distinguished by its equal spinules at the apex of the tibiae and by the conical fourth segment of the maxillary palpus. I have seen 21 examples from the West Indies representing 7 species, of which 6 appear to be new. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 520 KEY TO WEST INDIAN SPECIES OF BRYOPORUS 1. Elytral punctures very sparse, not forming distinct series______-_______-_-_ 2 Hiytralspunctures) in. mumerous: series-=. === = - = a a ee 3 2. Elytra broader than pronotum, punctures very irregular in size__ 6. cubanus Elytra not broader than pronotum, punctures sparse but of same size. 7. subterraneus 8. Pronotum with sparse transverse scratches____-__--___---~- 5. aciculatus Pronotum with punctures and punctulae only_____--__-_______--____-__-~ a 4. Black or piceous throughout, except for edges_______-_--___-__-_______-_- 5 Head and pronotum: brightirufousee 2-2 seme ee 3. bicolor 5. Head, pronotum, and elytra with at least traces of ground sculpture. 4. obscurus Waithoutminace: Or s2round) Sculptures. 222 2a aS A 6 Gehiytral’series;recular and, Gistincto222 =~ 38 2s a eee 1. regulus Elytral series in part irregular and confused____--_-__----_-_-- 2. similis 1. BRYOPORUS REGULUS, new species Description.—Black or piceous, edges sometimes rufescent. Head suborbicular, deflected; without distinct punctures on the disk but with irregular minute punctulae sometimes in clusters; without ground sculpture. Pronotum two-sevenths wider than long, widest at basal fourth; basal angles obtuse but distinct; strongly convex; with a single pair of large discal punctures at apical third and scat- tered rather coarse punctulae; without ground sculpture. Llytra with very large but very shallow punctures, excavated behind, in eight longitudinal series, the fifth, sixth, and seventh not perfectly regular but readily traceable; without ground sculpture. Abdomen with punctures finer than on elytra and often submuricate; without ground sculpture. Length, 314 to 6 mm. Type locality—Dominican Republic, cloudforest in the vicinity of Valle Nuevo; elevation about 6,000 feet. Types.—Holotype and three paratypes in the Museum of Compara- tive Zoology; four paratypes in the United States National Museum (No. 52543); collected in August 1938 by Dr. P. J. Darlington. (Paratypes partly from localities listed below.) Records.—The following are the records known to me: Hispaniola: Hartt, La Visité (Darlington, in M.C.Z.); DominicAN REPUBLIO, Valle Nuevo (Darlington, in M.C.Z. and U.S.N.M.), foothills south of San- tiago (Darlington, in M.C.Z. and U.S.N.M.); Sfnchez (Darlington, in M.C.Z. and U.S.N.M.), San José de las Matas (Darlington, in M.C.Z.), Constanza (Darlington, in M.C.Z.). Specimens examined.—I have seen the eight types and two other specimens from the Museum of Comparative Zoology. Remarks.—This species is distinguished as outlined in the key. It attains the largest size of any so far taken in the West Indies. The type was taken in cloudforest, but I have received no other record of the habits of this species. 528 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. BRYOPORUS SIMILIS, new species Description.—Piceous, abdominal segments rufescent behind. Head suborbicular, deflected; without distinct punctures on the disk but with irregular scattered punctulae; without ground sculpture. Pro- notum about one-fifth wider than long, widest at basal third; basal angles obtuse but distinct; strongly convex; with a pair of moderate discal punctures at apical fourth (and possibly another behind mid- dle) ; with strong but irregular punctulae but no ground sculpture. EHlytra with moderately large but very shallow punctures excavated behind and in eight longitudinal series; the sixth and seventh and apex of fifth confused; without ground sculpture. Abdomen with rather fine and not dense submuricate punctures; without trace of ground sculpture. Length, 414 mm. Type locality—Trinidad, base of Galeota Point. Types.—Holotype in the Museum of Comparative Zoology, collected on September 20, 1935, by N. A. Weber. Records.—The following is the only record known to me: Trinidad: Galeota Point (Weber, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species is very similar to regulus but has the elytral series more confused, the discal pronotal pair more anterior, and the pronotum proportionately less wide. T have received no record of its habits. 3. BRYOPORUS BICOLOR, new species Description—Head and pronotum rufous, elytra rufopiceous, ab- domen rufocastaneous. Head oval, the eyes much shorter than their distance from base; without discal punctures but with irregular punc- tulae; without ground sculpture. Pronotum one-seventh broader than long, widest at basal third; basal angles narrowly rounded; with a pair of discal punctures near apical third and another pair near basal fourth ; with punctulae more distinct and regular than on head; with- out distinct ground sculpture. FH7Jytra with large but shallow and posteriorly excavated punctures, the longitudinal series scarcely trace- able except at suture and sides, the punctures usually separated by their diameter or less; without ground sculpture. Abdomen with punctures similar to those of elytra but much finer, submuricate ; with- out distinct ground sculpture. Length, 414 to 5 mm. Type locality —Dominican Republic; cloudforest in the vicinity of Valle Nuevo; elevation about 6,000 feet. Types.—Holotype and two paratypes in the Museum of Comparative Zoology; two paratypes in the United States National Museum (No. 52544) ; collected in August 1938 by Dr. P. J. Darlington. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 529 Records.—The following are the records known to me: Hispaniola: Dominican Repusric, Valle Nuevo (Darlington, in M.C.Z. and U.S.N.M.), Loma Rucilla (Darlington, in M.C.Z.). Specimens examined.—I lave seen only the five types. Remarks. —This species is distinct by the bright rufous color of the head and pronotum, as well as by the shape of the head and the size of the eyes, and the basal pair of pronotal punctures. The types were taken in cloudforest, but I have received no other record of the habits of this species. 4. BRYOPORUS OBSCURUS (Erichson) Boiitobius obscurus ERICHSON, 18389b, p. 272.—Lene and MutTcHter, 1914, p. 407.— BERNHAUER and SCHUBERT, 1916, p. 460.—Wotcort, 1924, p. 79; 1936, p. 198. Description.—Piceous, edges of pronotum, elytra, and abdominal segments more or less rufescent. Head suborbicular, deflected; eyes at more than their length from base; without punctures on the disk; with indistinct ground sculpture. Pronotwm about one-fourth wider than long, widest at basal third; basal angles obtuse but distinct; rather strongly convex; with a single pair of discal punctures at an- terior fourth and another pair near base; surface indistinctly pitted and with traces of ground sculpture. lytra with moderately large but shallow and irregular punctures in longitudinal series, the sutural impressed, the rest not very distinct; with traces of ground sculpture between. Abdomen with punctures a little finer and less distinct than on elytra; without distinct ground sculpture. Length, 314 to 4 mm. Type locality —Puerto Rico. Types.—Either in the Hope Museum, Oxford, or the Zoologische Museum, Berlin. Records.—The following are the records known to me: Puerto Rico: (Erichson, 1839b). Grenada: (British Museum). Specimens examined.—I have seen only one example in the British Museum. It was borrowed for further study. Remarks.—The specimen on which the above description was based was placed in the British Museum under the name Bryoporus seri- atus Erichson from Brazil. My examination showed that the two were not the same, and I cannot now find any character to distinguish it from obscurus, which was described from the West Indies and ap- parently has not been reported since. I find no record of its habits. 5. BRYOPORUS ACICULATUS, new species Description.—Dark rufous, feebly picescent in part. Head sub- orbicular, eyes occupying less than half the length; without discal 530 BULLETIN 182, UNITED STATES NATIONAL MUSEUM punctures but with distinct minute punctulae; without ground sculp- ture. Pronotum about one-fourth wider than long, widest near basal third, basal angles obtuse but distinct; with a pair of discal punc- tures at apical third and another pair at basal fifth; with fairly coarse punctulae and irregular transverse aciculations or scratches; without ground sculpture. Hlytra with fairly coarse shallow punc- tures, excavated behind, in eight longitudinal series, somewhat im- pressed and not clearly defined at sides of disk; without ground sculpture. Abdomen with fine submuricate punctures rather dense; without distinct ground sculpture except on apical segments, Length, 4 mm. Type locality—Cuba, mountains north of Imias, eastern Oriente province; elevation 3,000 to 4,000 feet. Types.—Holotype in the Museum of Comparative Zoology, col- lected on July 25-28, 1936, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Cuba: Imias in Oriente (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species is very distinct because of the scratches on the pronotum. It also is unusual in having the rows of elytral punctures somewhat impressed. I have received no record of its habits. 6. BRYOPORUS CUBANUS, new species Description.—Piceorufous, the elytra rufotestaceous. Head sub- orbicular, a little broader than long, eyes nearly half as long as head; without discal punctures, but with scattered minute punctulae; with- out ground sculpture. Pronotwm about one-eighth broader than long, widest at basal third, basal angles somewhat rounded; with a pair of discal punctures at apical fourth and another pair at base; with minute punctulae but no ground sculpture. lytra with punc- tures indistinct except in sutural series and in an area at outer apical third and toward humerus, where they are coarse but sparse; without ground sculpture. Abdomen with moderate punctures broadly exca- vated behind and somewhat obscuring each other; without ground sculpture. Length, 214 to 3 mm. Type locality Cuba, Soledad, near Ciefuegos. Types.—Holotype in the Museum of Comparative Zoology; para- type in the United States National Museum (No. 52545); collected on January 15 and October 15, 1926, respectively. Records.—The following is the only record known to me: Cuba: Soledad (Darlington, in M.C.Z. and U.S.N.M.). Specimens ewamined.—I have seen only the two types. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 531 Remarks.—This species is readily distinguished by the testaceous elytra as well as by the feeble elytral punctation. The latter appears to be variable, but the large punctures are never regularly serial. Ihave received no record of its habits. 7. BRYOPORUS SUBTERRANEUS, new species Description.—Testaceous, the elytra and abdomen somewhat ru- fescent. Head almost as broad as long but somewhat narrowed in front, eyes at at least three times their length from base; with scat- tered irregular small punctures, but without ground sculpture. Pro- notum scarcely wider than long, widest just behind middle, only slightly narrower in front than at hind angles; with a pair of large discal punctures at apical fifth and another pair at basal fourth, and with large punctulae throughout; without ground sculpture. lytra shorter and narrower than pronotum; with coarse but irregular punc- tures throughout, serial only along suture; without distinct ground sculpture. Abdomen with coarse but very shallow punctures, feebly submuricate; with traces of strigulose ground sculpture, especially apically. Length, 3 mm. Type locality—Jamaica, main range of the Blue Mountains; ele- vation 5,000 to 7,388 feet. Types.—Holotype in the Museum of Comparative Zoology, col- lected on August 17-19, 1934, by Dr. P. J. Darlington. Records.—The following is the only record known to me: Jamaica: Blue Mountains (Darlington, in M.C.Z.). Specimens examined.—I have seen only the holotype. Remarks.—This species is very distinct because of its subterranean facies. The reduced elytra, the punctation of the head and elytra, and the shape of the pronotum will also serve to distinguish it. I have received no record of its habits. Subfamily HYPOCYPHTINAE This group has generally been classified as a tribe in the subfamily Tachyporinae, but it differs from the other members of that sub- family in so many points that I see no other course at present than to raise the tribe to a subfamily. These characters include the absence of the frontal sutures, the elongate lacinia of the maxilla, the com- plete submental sutures, the large mesosternal intercoxal process, the 4-seomented tarsi, the absence of paratergites on the second abdom- inal segment, and the extreme development of the male genitalia. Only one genus has been found in the West Indies. 5932 BULLETIN 182, UNITED STATES NATIONAL MUSEUM LXXXIX. Genus ANACYPTUS Horn Anacyptus Horn, 1877, p. 87. Microcyptus Horn, 1882, p. i. Micrecyptus EICHELBAUM, 1909, p. 196 (misspelling). Genotype.—Hypocyphtus testaceus LeConte= Anacyptus testaceus (LeConte) (monobasic). Of Microcyptus, Hypocyphtus testaceus LeConte= Microcyptus testaceus (LeConte) (Rules, Article 30, IT, f). Diagnosis —Head short and strongly deflected, not distinctly mar- gined beneath the eyes; hypostomal and coronal sutures not visible; antennae 10-segmented with the two basal and three apical segments enlarged, inserted at sides of head close to eyes; labrum semicircular ; maxillary palpus subulate, third segment large; gular sutures widely separated; pronotum and elytra extended beneath the body; anterior coxal cavities not at all concave but open behind; front coxae very large, exserted; middle coxae narrowly separated; posterior coxae very large, contiguous, broadly ‘expanded laterally and caudally beneath the femur and with an expansion over the base of the femur; abdomen narrowly margined basally, first and second sternites absent ; tarsi 4-segmented. Remarks—This genus contains the smallest species of Staphy- linidae known. It is distinguishable by its size, by the shape of the antennae, and by the structure of the posterior coxae. The name Aficrocyptus was proposed by Horn to replace Ana- cyptus, which he believed to be preoccupied. However, under our present code of nomenclature, Anacypta Illiger does not preoccupy Anacyptus, and the latter must be accepted. Only one species is known from the West Indies. 1. ANACYPTUS TESTACEUS (LeConte) Hypocyphtus testaceus LeContr, 1863, p. 30.—Horn, 1877, p. 125—HENsHAw, 1881, p. 219.—E1cHELBAUM, 1909, p. 196. Anacyptus testaceus (LeConte) Horn, 1877, p. 87, 125.—Scuwarz, 1878, p. 440.— HENSHAW, 1881, p. 219. Microcyptus testaceus (LeConte), ScHwarz, 1889, p. 160.—Dury, 1902, p. 124.— WASMANN, 1903, p. 98.—ErcHELBAUM, 1909, p. 196.—BLAtcHLEY, 1910, p. 442.—BERNHAUER and ScHuserT, 1916, p. 495. Description.—Rufotestaceous. Head sparsely minutely punctulate, without ground sculpture. Pronotum minutely punctulate and pubescent, the tiny setulae arranged to appear like striae; without ground sculpture. //ytra minutely punctulate and pubescent, the tiny setulae forming longitudinal and sometimes diagonal lines; with- out distinct ground sculpture. Abdomen generally very much retracted, with rather dense setae at the sides. Length, 2/5 mm. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 533 Type locality.—Athens, Ga. Types.—In the LeConte collection in the Museum of Comparative Zoology, Cambridge, Mass. Records —The following are the records known to me: Cuba: Cayamas (Schwarz, in U.S.N.M.). Montserrat: (Hubbard, in U.S.N.M.). North America: ArIzoNA, TEXAS, FLormpA (U.S.N.M.), Grorara (LeConte, 1863), GerorGIA, ARIZONA (Horn, 1877), LAKE SuperRiIon (Casey collection). Specimens examined.—I have seen five examples from the West Indies in the United States National Museum and numerous North American examples. Remarks.—This minute species appears to have a wide range although it is seldom collected. A. goeldii Wasmann from Brazil is very similar, if one may judge from the description, and may possibly be the same. It was found in termite nests. This species was originally reported from under pine bark, but I find no record concerning the West Indian examples. Subfamily ALEOCHARINAE Although this is by far the largest subfamily of staphylinids, it is a very heterogeneous one and one that is very poorly known from the standpoint of classification and morphology. Many thousands of species have been described, but comparatively few generic revisions have been published, and no adequate key to the genera has been proposed. The principal classifications that have been proposed have been based on the segmentation of the tarsi and palpi. It is known that these characters are not stable in some groups, but no allowance has been made for this fact. In addition, the structure of the tarsus is such that very different counts can be arrived at with different magnifications. Under these conditions it is exceedingly difficult, as well as uncertain, to make generic identifications, although species can sometimes be readily determined by comparison. Under these conditions it appears useless to add to the number of species requiring future reclassification, especially as no particular purpose is served by describing these obscure forms at this time. For this reason, I have compiled the records that have been published from the West Indies, but I have not described the new species that un- doubtedly do exist in our collections, and I have not attempted to identify the known species in this group. There have been recorded from the West Indies 156 species belong- ing to 41 genera. 534 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Genus DEINOPSIS Matthews Deinopsis MATTHEWS, 1888, p. 193. Dinopsis KRaatz, 1857b, p. 37. 1. DEINOPSIS GRACILIS Cameron Deinopsis gracilis CAMERON, 1922, p. 123.—BERNHAUFR and SCHEERPELTZ, 1926, p. 503. Records.—The following are the records known to me: St. Lucia: (Cameron, 1922; Cameron collection). Genus MYLLAENA Erichson Myllaena ERicHson, 18387, p. 382. Centroglossa MATTHEWS, 1838, p. 194. Myllana DUPONCHEL, 1841, p. 57 (misspelling). Millaena FENYES, 1918, pp. 3, 10 (misspelling). Myllanea FENYES, 1918, p. 15 (misspelling). 1. MYLLAENA CELERRIMA Cameron Myllaena celerrima CAMERON, 1922b, p. 650.—BERNHAUER and SCHEERPELTZ, 1926, p. 505. Records.—The following are the records known to me: St. Vincent: (Cameron, 1922b; British Museum). Grenada: (Cameron, 1922b; British Museum). 2. MYLLAENA CURTICORNIS Cameron Myllaena curticornis CAMERON, 1922b, p. 651.— BERNHAUER and SCHEERPELTZ, 1926, p. 505. FRecords.—The following are the records known to me: Grenada: (Cameron, 1922b; British Museum). 3. MYLLAENA DIFFICILIS Cameron Myllaena difficilis CAMERON, 1922b, p. 651.—-BERNHAUER and SCHEERPELTZ, 1926, p. 505. Records.—The following are the records known to me: Jamaica: (Cameron, 1922b; Cameron collection). 4. MYLLAENA DIVERSICORNIS Cameron Myllaena diversicornis CAMERON, 1922b, p. 650.—BERNHAUER and SCHEERPELTZ, 1926, p. 505. Records.—The following are the records known to me: St. Vincent: (Cameron, 1922b; British Museum; Cameron collection). Grenada: (Cameron, 1922b; British Museum.) 5. MYLLAENA FRAGILIS Sharp Myllaena fragilis SHARP, 1883, p. 286.—FENYES, 1920, p. 188.—BERNHAUER and SCHEERPELTZ, 1926, p. 506. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 535 Records.—The following are the records known to me: St. Lucia: (Cameron collection). Central America: GUATEMALA (Sharp, 1883). 6. MYLLAENA GRANULATA Cameron Myllaena granulata CAMERON, 1922b, p. 652.—BERNHAUVER and SCHEERPELTz, 1926, p. 507. Records.—The following are the records known to me: Jamaica: (Cameron, 1922b; British Museum; Cameron collection). 7. MYLLAENA INDEFATIGABILIS Cameron Myllaena indefatigabilis CAMERON, 1922b, p. 651.—BERNHAUER and SCHEERPELTZ, 1926, p. 507. Records.—The following are the records known to me: St. Lucia: (Cameron, 1922b; Cameron collection). 8. MYLLAENA OBSCURA Cameron Myllaena obscura CAMERON, 1922b, p. 651.—BERNHAUER and SCHEERPELTZ, 1926, p. 508. Records —The following are the records known to me: Hispaniola: Hartt (Cameron, 1922b; British Museum; Cameron collection). Genus PRONOMAEA Erichson Pronomaea ERIcHSON, 1837, p. 378. Pronomea REDTENBACHER, 1874, p. 73 (misspelling). 1. PRONOMAEA DEBILIS Cameron Pronomaea debilis CAMERON, 1922b, p. 652.—BERNHAUER and SCHEERPELTZ, 1926, p. 510. Records.—The following are the records known to me: [Jamaica: (Cameron, 1922b; Bexnhauer and Scheerpeltz, 1926) .] Tortola: (Cameron collection). Remarks.—The Jamaican record is probably an error since no specimens in the Cameron collection are labeled Jamaica. Tortola is in the Virgin Islands. Genus OLIGOTA Mannerheim Oligota MANNERHEIM, 1830, p. 72, 1. OLIGOTA ALBIDICORNIS Bernhauer Oligota albidicornis BERNHAUER, 1923b, p. 1438.—BrERNHAUER and SCHEERPELTZ, 1926, p. 512. Records.—The following are the records known to me: Guadeloupe: (Bernhauer, 1923). 536 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. OLIGOTA CADAVERINA Bierig Oligota cadaverina Brerta, 1934d, p. 113, fig. 1. Records —The following are the records known to me: Cuba: (Bierig, 1934). 3. OLIGOTA CENTRALIS Sharp Oligota centralis SHanp, 1883, p. 293.—FENyYEs, 1918, p. 56.—Cameron, 1922, p. 124.—BrerRNHAUER and SCHEERPELTZ, 1926, p. 512.—Breria, 1934, p. 114, fig. 2. Records.—The following are the records known to me: Cuba: (Bierig, 19384). Central America: GUATEMALA (Sharp, 1883; Bierig, 1984). 4. OLIGOTA CHRYSOPYGA Kraatz Oligota chrysopyga Kraatz, 1859, p. 45.—Frnyes, 1918, p. 56.—BERNHAUER and ScHEERPELTZ, 1926, p. 512.—Breric, 1934, pp. 113, 116. Oligota ventralis Fauvet, 1889, p. 171.—Fenyss, 1918, p. 56.—BERNHAUER and SCHEERPELTZ, 1926, p. 512. Records.—The following are the records known to me: Grenada: (British Museum; Cameron collection). Africa: (Bernhauer and Scheerpeltz, 1926; etc.). Orient: (Bernhauer and Scheerpeltz, 1926; etc.). 5. OLIGOTA HYPOCYPTINA Bernhauer Oligota hypocyptina BrERNHAUER, 1923, p. 143.—BreRNHAUER and SCHEERPELTZ, 1926, p. 5138. Records.—The following are the records known to me: Guadeloupe: (Bernhauer, 1923). 6. OLIGOTA LAXATA Cameron Oligota lawata CAMERON, 1922, p. 124.—BrRNHAUER and SCHEERPELTZ, 1926, p. 513. Records—The following are the records known to me: Jamaica: (Cameron collection). Grenada: (Cameron, 1922; British Museum). 7. OLIGOTA LUTEICORNIS Bierig Oligota luteicornis Breric, 1934, p. 119, figs. 5, 10. Records.—The following are the records known to me: Cuba: (Bierig, 1934). 8. OLIGOTA MACULICORNIS Cameron Oligota maculicornis CAMERON, 1922, p. 124.—BERNHAUER and SCHEERPELTZ, 1926, p. 514.—Brerie, 19384, p. 117. Records.—The following are the records known to me: Cuba: (Bierig, 1934). Hispaniola: Hartt (Cameron, 1922; Cameron collection). | MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 537 9. OLIGOTA MINUTA Cameron Oligota minuta CAMERON, 1931, p. 82.—SCHEERPELTZ, 1934, p. 1532. Records.—The following are the records known to me: Grenada: (Cameron collection). Trinidad: (Cameron, 1931; British Museum; Cameron collection). 10. OLIGOTA MINUTISSIMA Bernhauer Oligota minutissima BERNHAUER, 1923, p. 144.—BERNHAUER and SCHEERPELTZ, 1926, p. 5138. (Not Stephens, 1832; not Heer, 1841.) fecords.—The following are the records known to me: Guadeloupe: (Bernhauer, 1923). 11. OLIGOTA PARVA Kraatz Oligota pygmaea KRAATZ, 1858b, p. 3852. (Not Solier, 1849.) Cligota parva KRAAtTz, 1862, p. 300. Oligota contempta WoLLAstTon, 1867, p. 231. Gligota aliena MULSANT and REy, 1873, p. 134. Oligota californica Casry, 1911, p. 230. Oligota congruens Casry, 1911, p. 230. Oligota esmeraldae CASEY, 1911, p. 231. Records.—The following are the records known to me: Jamaica: (Cameron collection). Tortola: (Cameron collection). St. Vincent: (British Museum; Cameron collection). Grenada:(British Museum; Cameron collection). South America: ARGENTINA (Bernhauer and Scheerpeltz, 1926). Central America: MExico (Cameron collection). North America: NEVADA, CALIFoRNIA (Bernhauer and Scheerpeltz, 1926; etc.). Europe: (Woliaston, etc.). Orient: CHina (Cameron collection). 12. OLIGOTA RHOPALOCERA Bernhauer Oligota rhopalocera BERNHAUER, 1923, p. 145—BreRNHAUER and SCHEERPELTZ, 1926, p. 516.—Birria, 1934, p. 116. Records—The following are the records known to me: Cuba: (Bierig, 1934). St. Thomas: (Bernhauer, 19238). 13. OLIGOTA RUFA Cameron Oligota rufa CAMERON, 1922, p. 125.—BERNHAUER and SCHEERPELTZ, 1926, p. 516. Records.—The following are the records known to me: Jamaica: (Cameron collection). St. Lucia: (Cameron, 1922). 449008—42 35 5388 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 14. OLIGOTA SMITHI Cameron Oligota smithi CAMERON, 1922, p. 124—BERNHAUER and SCHEERPELTZ, 1926, p. 513. Records.—The following are the records known to me: Grenada: (Cameron, 1922; British Museum). 15. OLIGOTA TESTACEORUFA Bernhauer Oligota testaceorufa BreRNHAUER, 1923, p. 144.—BERNHAUER and SCHEERPELTZ, 1926, p. 513. Records—The following are the records known to me: St. Thomas: (Bernhauer, 1923). 16. OLIGOTA TRICOLOR Bierig Oligota tricolor Birric, 1934, p. 116. Records.—The following are the records known to me: Cuba: (Bierig, 1934). 17. OLIGOTA ZONATA Bierig Oligota zonata Bieric, 1934, p. 115. Records.—The following are the records known to me: Cuba: (Bierig, 1934). 18. OLIGOTA, species indeterminate Hispaniola: Hartr (Cameron collection). Tortola: (Cameron collection). Genus THYREOXENUS Mann Thyreorenus MANN, 1923, p. 329. 1. THYREOXENUS PARVICEPS Mann Thyreozrenus parviceps MANN, 1923, p. 330, 331.—BrRNHAUER and SCHEERPELTZ, 1926, p. 519.—IEi Merson, 1935, pp. 371, 372. Records.—The following are the records known to me: Trinidad: (Mann, 1923). South America: BririsH GUIANA (Mann, 1923). 2. THYREOXENUS, species indeterminate Trinidad: (Seevers, 1937). Genus EBURNIOLA Mann Eburniola MANN, 1923, p. 33. 1. EBURNIOLA LEUCOGASTER Mann Eburniola leucogaster MANN, 1923, p. 33, fig. 31—BrERNHAUER and SCHEERPELTZ, 1926, p. 519.—Emerson, 1935, pp. 369, 370, 371, 373, 381, 387, 389, fig. 2. MONOGRAPH OF WEST INDIAN STAPHYLINIDAB 539 Records.—The following are the records known to me: Trinidad: (Mann, 1923; Emerson, 1935). South America: BriTIsH GUIANA (Mann, 1923; Emerson, 1935). Genus THAXTERIA Fenyes Thazrteria FENYES, 1921, p. 17. 1. THAXTERIA INSULARIS Fenyes Tharteria insularis FENYeS, 1921, p. 17.—BerNHAUER and ScHUBERT, 1926, p. 521. Records.—The following are the records known to me: Grenada: (Fenyes, 1921). Genus PERINTHUS Casey Perinthus Casey, 1889¢, p. 192. 1. PERINTHUS XENOCOSTALIS Seevers Perinthus xenocostalis SEEVERS, 1937, p. 9, pl. 3, fig. 18. Records.—The following are the records known to me: Trinidad: (Seevers, 1937). South America: BririsH GUIANA (Seevers, 19387.) 2. PERINTHUS DUDLEYANUS Casey Perinthus dudleyanus Casey, 1889c, pp. 194, 196.—SrEvrRs, 1937, pp. 9, 10, 11, pl. 8, fig. 19. Records —The following are the records known to me: Trinidad: (Seevers, 1937). South America: BririsH GUIANA, Ecuapor (Seevers, 1937). Central America: PANAMA (Casey, 1890; Seevers, 1937). Genus BARYCHARA Sharp Barychara SHARP, 1883, p. 292. 1. BARYCHARA FLAVIPENNIS Cameron Barychara flavipennis CAMERON, 1922, p. 125.—BERNHAUER and SCHEERPELTZ, 1926, p. 521. Records.—The following are the records known to me: Hispaniola: Hair1 (Cameron, 1922; Cameron collection). Genus GNYPETOSOMA Cameron Gnypetosoma CAMERON, 1922, p. 127. 1. GNYPETOSOMA CALOCERA Cameron Gnypetosoma calocera CAMERON, 1922, p. 127.—BERNHAUER and SCHUBERT, 1926, p. 523. Schistoglossa calocera FAuvEL, MS.—CAMERON, 1922, p. 127. Records —The following are the records known to me: St. Vincent: (Cameron, 1922; British Museum). 540 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. GNYPETOSOMA FARREA Cameron Gnypetosoma farrea CAMERON, 1922, p. 128.—BrrRNHAUER and ScHEERPELTZ, 1926, p. 523. Schistoglossa farrea FauvEL MS.—CAMErOoN, 1922, p. 128. Records —The following are the records known to me: St. Vincent: (Cameron, 1922; British Museum). Genus ALISALIA Casey Alisalia CASEY, 1911, p. 219. Alysalia FENYES, 1918, p. 67 (misspelling). 1. ALISALIA BRUNNEA Cameron Alisalia brunnea CAMERON, 1922, p. 126.—BrrRNHAUER and SCHEERPELTZ, 1926, p. 524. Records.—The following are the records known to me: Hispaniola: Hartt (Cameron, 1922; Cameron collection). 2. ALISALIA PICEA Cameron Alisalia picea CAMERON, 1922, p. 126.—BERNHAUER and SCHEERPELTZ, 1926, p. 524. Records—The following are the records known to me: Hispaniola: Hartr (Cameron, 1922; Cameron collection). Genus BRACHYCHARA Sharp Brachychara SHARP, 1883, p. 267. 1. BRACHYCHARA ATERRIMA Cameron Brachychara aterrima CAMERON, 1922b, p. 637.—BeRNHAUsR and SCHEERPELTZ, 1926, p. 526. Records—The following are the records known to me: St. Vincent: (Cameron, 1922b; British Museum). Genus GYROPHAENA Mannerheim Gyrophaecna MANNERHEIM, 1880, p. 74. 1. GYROPHAENA AENEICOLLIS Cameron Gyrophaena aencicollis CAMERON, 1922b, pp. 639, 642.—BreRNHAUER and SCHEER- PELTZ, 1926, p. 528. Records—The following are the records known to me: St. Vincent: (Cameron, 1922b; British Museum). Grenada: (Cameron, 1922b; British Museum; Cameron collection). 2. GYROPHAENA ATOMARIA Cameron Gyrophaena atomaria CAMERON, 1922b, pp. 637, 647.—BERNHAUER and SCHEEB- PELTZ, 1926, p. 529. Cuba: (Cameron collection). Hispaniola: Haitr (Cameron, 1923; Cameron collection). 2. PHLOEOPORA PRODUCTA Cameron Phloeopora producta CAMERON, 1923, p. 388.—BERNHAUER and SCHEEFRPELTZ, 1926, p. 720. Records—The following are the records known to me: Hispaniola: Harrr (Cameron, 1923; Cameron collection). 3. PHLOEOPORA RELIGATA Erichson Phloeopora religata EricHson, 18389b, p. 79.—BrERNHAUER and SCHEERPELTZ, 1926, p. 720. Records.—The following are the records known to me: Puerto Rico: (Erichson, 1839b). Genus GCCALEA Erichson Ocalea EricHson, 1837, p. 298. 1. GCALEA, species indeterminate West Indies: (British Museum). Genus TERMITOGASTER Casey Termitogaster CASEY, 1889, p. 63. 1. TERMITOGASTER BREVIS Mann Termitogaster brevis MANN, 1923, p. 344.—BrERNHAUER and SCHEERPELTZ, 1926, p. 734. Records—The following are the records known to me: Trinidad: St. Joseph (Mann, 19238). South America: BriTIsH GUIANA (Mann, 1923). 2. TERMITOGASTER EMERSONI Mann Termitogaster emersoni MANN, 1923, p. 842.—BERNHAUER and SCHEERPELTZ, 1926, p. 734. Records. The following are the records known to me: eo Trinidad: St. Joseph (Mann, 1923). South America: BririsH GUIANA (Mann, 1923). Genus TERMITOPHYA Wasmann Termitophya WASMANN, 1903, p. 95. 1. TERMITOPHYA FLAVIVENTRIS Mann Termitophya flaviventris MANN, 1923, p. 352.—BrrNHAUER and SCHEERPELTZ, 1926, p. 736. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 559 Records.—The following are the records known to me: Trinidad: St. Joseph (Mann, 1923). South America: -BriTISH GUIANA (Mann, 1936). Genus EUTHORAX Solier Buthorawv Souter, 1849, p. 345. Myrmecochara KRAATz, 1857b, p. 40. Campoporus LYNCH, 1884, p. 64. Camponotus FAuVEL, 1887, p. 230 (error). Eurynotida Casey, 1906, p. 343. Dinusina BERNHAUER, 1908a, p. 249. 1. EUTHORAX PICTIPENNIS Kraatz Kuthorax pictipennis Kraatz, 1857, p. 41.—SHaArp, 1883, p. 163.—BsrNHAUER and ScCHEERPELTZ, 1926, p. 738. Records——The following are the records known to me: St. Vincent: (British Museum). Central America: Mexico (Sharp, 1883). North America: LouISIANA (Bernhauer and Scheerpeltz, 1926). Genus CCYUSA Kraatz Ocyusa KRAAtTz, 1856, p. 156. 1. OCYUSA GRANELLA Cameron Ocyusa granella CAMERON, 1923, p. 387.—BERNHAUER and SCHEERPELTZ, 1926, p. 743. Schistoglossa granella FAUVEL, MS.—CAMERON, 1923, p. 387. Records.—The following are the records known to me: Grenada: (Cameron, 1923; British Museum; Cameron collection). Genus ALEOCHARA Gravenhorst Aleochara GRAVENHORST, 1802, p. 67. Aleohcara SAHLBERG, 1834, p. 351 (misspelling). Aleochora FENYES, 1918, p. 9 (misspelling). 1. ALEOCHARA BIMACULATA Gravenhorst Aleochara bimaculata GRAVENHORST, 1802, p. 187.—LENG and MutTcHtier, 1914, p. 407.—BERNHAUER and SCHEERPELTZ, 1926, p. 792. Aleochara duplicata Er1icHson, 1889b, p. 167. Aleochara alticoia SHarp, 1883, p. 148. Records.—The following are the records known to me: Hispaniola: Harrt (Leng and Mutchler, 1914). South America: Cotompia (Bernhauer and Scheerpeltz, 1926). Central America: Mexico, GUATEMALA (Bernhauer and Scheerpeltz, 1926; etce.). North America: (Gravenhorst, 1802; etc.). 560 BULLETIN 182, UNITED STATES NATIONAL MUSEUM 2. ALEOCHARA CAMERONI Bernhauer and Scheerpeltz Aleochara bugnioni CAMERON, 1923, p. 889. (Not Fauvel, 1901.) Aleochara cameroni BERNHAUER and SCHEERPELTZ, 1926, p. 726. Records.—The following are the records known to me: St. Vincent: (Cameron, 1923; British Museum; Cameron collection). Grenada: (Cameron, 1923; British Museum). 3. ALEOCHARA LATERALIS Erichson Aleochara lateralis ErtcHson, 1839b, p. 161.—IFAvUVEL, 1863, p. 428.—BERNHAUER and SCHEERPELTZ, 1926, p. 778. Aleochara bonariensis LYNCH, 1884, p. 70.—BERNHAUER and SCHEERPELTZ, 1926, p. 778. FRecords.—The following are the records known to me: Cuba: (Fauvel, 1863). South America: Braz, CoLomBiA (Fauvel, 1863). 4. ALEOCHARA NIGROCOERULEA Cameron Aleochara nigrocoerulea CAMERON, 1923, p. 389. Aleochara nigrocaerulea BERNHAUER and SCHEERPELTZ, 1926, p. 778 (misspelling). Fecords.—The following are the records known to me: Jamaica: (Cameron, 1923; Cameron collection). 5. ALEOCHARA NOTULA Erichson Aleochara notula ErtcHson, 18389b, p. 167.—DuVAt, 1857, p. 32.—CHEvroraT and FAUVEL, 1868, p. 429.—SHaArp, 1888, pp. 148, 149.—FLeuTIAux and Satrf, 1889, p. 379.—ViLiapa, 1901, p. 27.—FeEnyes, 1908, p. 65.—LENG and Mutcn- LER, 1914, p. 407.—F enyes, 1920, p. 413.—BrrRNHAUER and SCHEERPELTZ, 1926, p. 794. Aleochara duplicata Lyncu, 1884, p. 78.—Frnyes, 1920, p. 413. (Not Erichson, 1839.) Aleochara signaticollis FAmRMATRE and GERMAIN, 1861, p. 413.—Favuvet, 1866, pp. 285, 288.—FAvvEL, 1868, p. 63.—SHARP, 1883, p. 148.—FrnyEs, 1920, p. 413. Records. Cuba: (DuVal, 1857; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914). Jamaica: (Cameron collection). St. Thomas: (Hrichson, 1839b; DuVal, 1857; Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; Fenyes, 1920.) Guadeloupe: (Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). St. Vincent: (British Museum). Grenadines: Mustique (British Museum). South America: (British Museum; Cameron collection), CH1ne (Fairmaire and Germain, 1861, as signaticollis; Fauvel, 1866, 1868; Fenyes, 1920; Sharp, 1883), ARGENTINA (Fenyes, 1920), Central America: Mexico, GUATEMALA, NICARAGUA (Sharp, 1883), Mexico (Vil- lada, 1901). North America: (Fenyes, 1920). | The following are the records known to me: MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 561 6. ALEOCHARA PUBERULA Klug Aleochara puberula Kuiue, 1832, p. 189. Aleochara deserta EricHson, 1839b, p. 173. Aleochara vaga EricHson, 18389b, p. 172. Aleochara decorata AuBE, 1850, p. 311. Aleochara armitagei WoLLASTON, 1854, p. 559. Aleochara brunnescens MorscHULsKY, 1858, p. 243. Aleochara sanguinolenta MoTSCHULSKY, 1858, p. 241. Aleochara dubia FAUVEL, 1868, p. 428. Aleochara analis MacLeEay, 1878, p. 135. Aleochara major EICHELBAUM, 1911, p. 176, Records.—The following are the records known to me: Cuba: (Fauvel, 1863, as dubia). Cosmopolitan: (Bernhauer and Scheerpeltz, 1926, ete.). 7. ALEOCHARA TAENIATA Erichson Aleochara taeniata ERIcHSON, 1889b, p. 165—Suarp, 1883, pp. 150, 151.— FLEUTIAUX and SALLf, 1889, p. 379.—Lene and Murcutrr, 1914, p. 407.— Fenyes, 1920, p. 404.—PLavinstscHikov, 1929, p. 80.—BERNHAUER and SCHEERPELTZ, 1926, p. 788. (Not Solksy.) Aleochara lapidicola SAHLBERG, 1844, p. 517.—FrnyeEs, 1920, p. 404. Aleochara orypodia Suarp, 1883, pp. 150, 151; 1887, p. 776.—Frnyrs, 1920, p. 404. Aleochara simulairic SuHarp, 1883, p. 151; 1887, p. 776—Fxrnyes, 1920, p. 404.— BERNHAUER and SCHEERPELTZ, 1926, p. 783. Records.—The following are the records known to me: Jamaica: (Cameron collection). St. John: (Erichson, 1839b; Sharp, 1883; Fenyes, 1920). Guadeloupe: (Fleutiaux and Sallé, 1889; Leng and Mutchler, 1914). St. Vincent: (British Museum). Grenada: (Cameron collection; British Museum, as simulatriz). South America: Brazin (Sahlberg, 1844, as lapidicola; British Museum; Fenyes, 1920) ; CAYENNE, Peru (Sharp, 1883), Peru (Plavitschikov, 1929). Central America: GUATEMALA (Sharp, 1883, as simulatriz ; Fenyes, 1920; British Museum), GUATEMALA, PANAMA (Sharp, 18838, as ozypodia), PANAMA (Sharp, 1887, as similatriz). North America: ArRIzoNA (Fenyes, 1920). 8. ALEOCHARA VERBERANS Erichson Aleochara verberans ErRIcHson, 1839b, p. 164.—CHEvRoLAT and FAUVEL, 1863, p. 428.— SHaArp, 1883, p. 149.— Lene and MutrcHier, 1914, p. 407.—Fernyes, 1920, p. 410.—BERNHAUER and SCHEERPELTZ, 1926, p. 788. Records.—The following are the records known to me: Cuba: (Chevrolat and Fauvel, 1863; Leng and Mutchler, 1914; British Museum). South America: Brazin, CoLtomBIA (Hrichson, 1839b; Chevrolat and Fauvel, 1863; Sharp, 1888), Brazin (Fenyes, 1920; British Museum), ARGENTINA (Bern- hauer and Scheerpeltz, 1926). Central America: Mmxico (Sharp, 1883; British Museum). SYSTEMATIC CATALOG This check-list contains the names of all species of Staphylinidae known to occur in the West Indies, arranged in the same order as in the descriptive section. The numbers after the generic names and before the specific names will aid in referring back to the descriptions, which are similarly numbered. Generic synonymy is omitted except where necessary to link the present names to ones previously in use. Specific synonymy is restricted to the separate specific names that have been proposed. Synonyms are printed in italics. The abbreviation “etc.” is used to indicate that the species is known from parts of the world other than the West Indies and continental America. The word “to” between the names of two regions indicates that the species occurs in one or several of the intervening islands as well as on those actually mentioned. For detailed distribution (and synonymy) see the fore- going descriptive section. PIESTINAE Hypotelus Er. (1) Tadnsulanus Brot. 22a ee eee eee Cuba, Jamaica, St. Vincent Piestus Grav. (IT) 14 sulcatus Gravee = eee Dominica to Trinidad, South America QED OS TH eee Oa eG Ee ec) he eee Guadeloupe sepenicillatys: (Dalim. ss ee | eee ee ee eee Cuba to Trinidad striata (Gray) erythropus Hr. 4, pygmaeus ape=--=s2 2-2 o ene Guadeloupe to Trinidad, South America, Central America rufipennis (FF. & 8.) Dn CADLICOINIS Wea Dees n. ee eee eee Guadeloupe, South America muticus Fy. OMALIINAE Phloeonomus Heer (IIT) f'.-pedicularius (br)\o2 25 22 eee 2 eee Cuba to Trinidad lacrymale (F. & 8.) Omalium Grav. (IV) tee jamaicensis: BUkKwr saan ee Jamaica 2° darlingtoni (BK yr ee Jamaica 3. cubanumy Blew st 225 ee Se ee we ee ee Cuba AY SV UOT L UII Uy a ee Hispaniola OXYTELINAE Carpelimus Sam. (V) Trogophloeus Mannh, il; eroceipes: (PVis) a2 See A Ae pee ee ee Cuba to Guadeloupe arcuatus (Pvyl.) 562 MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 563 Carpelimus Sam. (V)—Continued. 2. ONnrort 10. tt: 12: 13. 14. 15. 16. 17. 18. 19. 20. 21. 22. 24. 25. 26. 27. 28. 29. 30. 31. 32. 33. 34. 35. 36. 37. ~ fihvipes *QMES) 222s) 3s— 2 aes Cuba to Grenada, South America, Central America, ete. rubripennis (F v1.) senilis (Shp.) texanus (Csy.) aequalis (Gundl.) Mm SELICSUS U@alils) tease eee Ee Cuba, Jamaica, St. Croix sextagonalis (Brg.) Mm COPRCCIIS) Db IKWrse sees na ea eee Jamaica to Trinidad OPN O TOMAR URW = a ee Hispaniola MDCACHY Bk Witees- eos se oS ee Hispaniola to St. Lucia + CORNUCOPIUIS DKW. se sea e Grenada MASmMithinGB nn ry=+~ 22 See a ee Cuba to Trinidad smithi (Cam.) smithianus (Scheerp.) croceipes (Brg.) Ve De Lie blkWit-= 2s. eae wee eee See 2.) Se ee Se Trinidad hispaniolus: BikiwP 222252 = oes eee ss ee Hispaniola TM PUNCtUSpBNGWR == — a se ee ee Jamaica ODSoleScenSNBiK wrest te a ek oe ee ee BD Hispaniola DOTINGUCHSISHBIK wits- =~ Sa ee Puerto Rico GantorthicBlkwits =.= 2252 se oo sea tS Jamaica, Puerto Rico CUDENSISHGB oy) =e Sn i ie ee Cuba, Jamaica TMLcAtOLw GSTS. sola Cuba to Guadeloupe SViSAITET CO TENT Soa (OES TMT 5) pea wr Cuba to Trinidad lactipennis (Cam.) ATIGUSS (DIVA eae ee a ee eee Cuba to Trinidad BeOS PAB oyaeie & ee ee Ee Be Jamaica DHA OS WB WW = cae a ee Cuba, Jamaica BCLObI Sere (Cams) aaa = ee Ss ee Jamaica to Grenada PetOMUSNBU wre Ae ke SIS ee ee Jamaica to Antigua al avIDES CHE) 22 aoe a Bahamas to Grenada, Central America hiemalis (Brg.) flavibasis (Brg.) Sub imiorn (Cams) pao ee ee Jamaica, Grenada DOM Ss CBiSs) eas eo ee Cuba ADL OMT Sees lkiyy reese eo eee Cuba, St. Croix GUNADILN eM wrt 20 Sa ee a ee Trinidad Giscipennigu(Bye:) 2 ss. = Cuba, Jamaica chapinis: Bllkwr2 so. sis 2-5 2a 2 ee Jamaica UL OTIS aes ayy Tae ae eR DS a Hispaniola contormis Blkwr-2>ss- 2" Jo 2 eee Cuba L2equalis | (DuVialjija= 22s = = Bermuda, Bahamas, Cuba, Jamaica funestus (Cam.) amplicollis (Brg.) GISSONUS ss, (BEE) a ce eee esl | ee ee Cuba LESEAGEI PEI Ses (CAIN ee seas ee a ee Hispaniola SORGIOMS (Cams) eae i ee ee ee Caymans CLE rEUMETC Lhe (GET) es se ees De Cuba Uva era Sta GH Wile) ae etre ee a ye eee Cuba, Jamaica 564 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Torrentomus Brg. (VI) Aa torrei: Brets:e! te Se Be te Cuba 2: tenebrosus) wBlkwr. 522. es Jamaica Apocellus Er. (VII) J. ustulatus r= 2s ane ee eee ee ee St. John to Grenada 2. planus: Shps222 2-233 Ss Se eee ee eee Trinidad, South America Oxytelus Grav. (VIII) 1. insignitus Grav_-___ Cuba to Trinidad, South America, Central America, North America, ete. americanus Mannh. pumilio Boh. 2. pluvius :Blkwtst. <2. =82 3s se A eee Trinidad POON CER OTIS MH Vernet ee ee St. Vincent, South America 4° incisus, Mots = 22222_2— Bermuda, Cuba to Trinidad, South America, Central America, ete. ferrugineus Kr. laevior Shp. bledioides Blackb. laxipennis Fairm. FyshOTOMIS a Wawel a 2 ae ee ee Puerto Rico GiaSCOTpios HW seen ea eee ee Jamaica, Hispaniola, Puerto Rico Wao loreOSus WOlc 222 2- so os eee Jamaica to Grenada, ete Sschapinine blow rl s-2 25 2 2 os ee ee Se rr ee ee Jamaica Oo jamaicensis Blkwre 2 22.22 se oe ee ee Be ee Eee ee Jamaica Parosus Shp. (IX) fevskahitgZioyl (nee se eee eee St. Vincent, Grenada 2 antillamim. Wend] 22-22) aa eee ee eee Guadeloupe Thinobius Kies. (X) davormmatus, Cames=2-- aoe eee ae eee Cuba, Jamaica. Hispaniola tenebrophilus Brg. flavifer Brg. 2. sopaculusiGames-2 -..8 25s oe Cuba, Jamaica, Hispaniola. Dominica morio Brg. nebulosus Brg. cubensis Brg. SMITI CONS OAM a 8 ee ee ee eee eee St. Vincent, Grenada 4, Jamaicensis Blikwri.2--.-2-- 22. Jamaica 5. exasperatus Blkwr__-----__-__----___- Jamaica, Hispaniola, Montserrat Ganitidulus *Bnhr 22222 Soe ee ee Cuba, Jamaica, Puerto Rico piceus Cam. Platystethus Mannh. (XI) 1. Spiculus:Er-_-2. = # Cuba to Trinidad, South America, Central America exiguus DuVal Bledius Sam. (XII) 1. ecaribbesnus Blkwr_ 2-22 ee ee Hispaniola to Tobago 2; esposus Blkwr2ss-- 3-2. he einai oe Antigua eo. withyeombel), Bohr. -. =e ee ere ele Trinidad 4..eubensis Blkwr_.=—--_ 3-4 a ee re eee Cuba 5: :trinidadensis Blkwr ===." =e ee ae Trinidad 6. jamaicensis Blkywr- =. 2555 eee Cuba, Jamaica abesttyin bik Wr 2 ee ee eo ee Vieques, St. Croix §) ceratus bllowr 2s eee Cuba, Hispaniola MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 565 OSORIINAE Pseudolispinodes Bnhr. (XIII) IeMOnU Pde MB IKWiHe 2 £2 2 Fe ee a Se Trinidad Deore VCLSUS@PEs Wiles sss js es ee Lo eee ea Hispaniola So MUM PALAU aI) ae ee on eS Se ee ew Cuba to Trinidad claviger (Cam.) Aeirregularis rp licwr 2 fel. 2s ee Cuba, Jamaica, Hispaniola DaGantorthineblikw r= 22 one ee Puerto Rico, Guadeloupe G. stenellus@Hir)) 2. -- Cuba to Grenada, South America, Central America, North America fauveli (Shp.) 1887 flavipennis (E'vl1.) tenuis (LeC.) that OM COUUIS ds Uy Bas oe mae ee ee pa ee Montserrat Sisnigrifrons: (ivi. 22.26.8920 er Se Cuba to St. Lucia sparsepunctatus (Cam.) OMenaAdeloupser blk wites 22 k= Lee ee eS aa ee Guadeloupe Lispinus Er. (XIV) Pa COMI Sees ees en eee ee ee Cuba, Puerto Rico, Central America QA ATCINICUS ERK WIs =o oe eee es ae ea Trinidad SCatenn, SNP Ais ans ah tt Trinidad, South America, Central America AMANO UINUS Tee Lee Jamaica, Hispaniola HsAtELenUatus Pre ates sen Pe Puerto Rico, Guadeloupe, Trinidad GrstriolavWrs= see Cuba, Trinidad, South America, Central America Me gnSubamigs Huygeese! 2 ke Se Cuba to Grenada, Central America granadensis Evi. Sap OXa MES owes oS 0 ee a eee St. Lucia Inopeplus Smith. (XV) HSSRWOLCO CLigy ES Mccoy re hc oi le Sere Ce os ae eS ee Hispaniola Zremutchieni iBilowies 25 2 2 a Guadeloupe omassistans. Bikwr ss) ots 2 Ad on ee eee St. Lucia A BSGRIae ulus, Bika 22 oo ek ee Guadeloupe Glyptoma Er. (XVI) Calocerus Fyl. ieee dagensis: UBikwr. 22 eee Trinidad BEreaGelupensis! «CWend.) _...2- 2-3 a wi Guadeloupe Hspeson Schauf. (XVII) Me CTASSULUS ME Vilar a2 Sto ee oy Montserrat to St. Vincent SeCMIMECEOIGCS yD 2 8 ee Guadeloupe to Trinidad PemIDOTAULI Se OCHA te = 2 Lk 2 oo ee ne Cuba to Trinidad AAPPRLLLGESTA SB Vil meee ee a ee Guadeloupe, South America Thoracophorus Mots. (XVIII) eRe “WENO nF Puerto Rico to Grenada PereCrane lust lows 2 22 ee a a fo Re eal Trinidad 3. brevicristatus (Horn)_--._ Cuba to Guadeloupe, Central America, North America See Page BON SIS OOM. 852222 eo Soe Cuba to Trinidad aeRO Ne) gos 2 Saat ee ar St. Thomas eperehil beams Eevee ee ee Trinidad euiGeTi tbe Lia (Etec 18 F So eet ee Puerto Rico Hleusis Lap. (XIX) PICU DED RCW ie 2 8 Sa ed ge Pen Tale Cuba 566 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Paralispinus Bnhr. (XX) 1; politus «(Shp.)2222==-2222— Guadeloupe to St. Vincent, Central America 2 OXI oul s CHES) eee Cuba to Grenada, South America, Central America, North America, ete. laevigatus (Kr.) rufescens (LeC.) rufus (Iv1.) fauveli (Shp.) 1876 aruensis (F vl.) pallescens (Blackb.) obscuripennis Bnhr. 3) rulomuS+BIKWii2s2. 2 Se oe eee ee eee Jamaica 4 ChepUSCULUS bl KW a= sea a eee ee eee ~ Cuba to Trinidad 5; punctatus; Blkwre 2s. 2 — a oe eee Puerto Rico Leptochirus Germ. (XXI) 1; maxillosus (Wab:)=]-=2===22=-— Trinidad, South America, Central America bicolor Lucas bolivianus Heller collaris Fv1. anthracinus Fvyl. Neotrochus Blkwr. (XXII) 1 minor (y))ess2=— = Cuba, Jamaica, Grenada, Trinidad, South America 2. -trinita tis (Bliiwrt=s=2242 =e se a eases See Trinidad 32 cylindrus (ir) 222222. Puerto Rico, Grenada, Trinidad 4 smithi-€Cam:) 22222 228 We ea ee eee St. Vineent, Trinidad Holotrochus Er. (XXIII) i poundi. Bliktwr- 25.22 2523S en 52 ee ee eee Trinidad 2) volvulus Hrsa<-<==--<=ssessse2eseeeosee 24 - Senssescaate2 Puerto Rico Mimogonus Fvl. (XXIV) i etumator: (ivls))22s=2-—=—-5=—— Cuba to Guadeloupe, Central America, ete. fauveli (Cam.) Osorius Latr. (XXV) iireoulus Darl 2.5. =. 2s asce eens SSeS ee Hispaniola 2, darling toniJBlktwti 22+ 2222 5. 2S eae Cuba 3. erenulifirons: INOUM —-=222- 2532252 =. Sees eee eee Cuba 4; fratellus Darliu.2-2=255.22-. oS se Seen san see ee Hispaniola 5. Salutator.Darl--- ==... ----2222--=2 ee tee ee Hispaniola 6; hirtilabris, .Darl<- = 2-2-2 22 se a eee Hispaniola “;, (daemon, Warli-28. = 2 sa oe eee eee eee ee Hispaniola Si icubensis) BIKWr. 22a en en eae ea eee eee Cuba 9. ‘Strictiis BIK Wr 22-2222 osese se ase asa eeesa= 4 Le Cuba TO socors DarlbSie sto.) non a re n e e e e Cuba 11. manni Notm:.2- ==. == 9-2 sa ee eee eee Cuba 12; cautis Darl: 2225. ES Hispaniola 15. mediocris, Danl.. 2-4 ee ee Hispaniola i4. biarmatus Darl. ite rosie tee ee eee eee Hispaniola OE DUSCKE SNOW = 225 a ee Hispaniola 16:: Oriente Bik wr.-2= + 22525262 slhseo5o-e5-- ee ee eee Cuba 1%. turquinus Blk Wr s.2 25+ 2222 a Se Cuba TSswatipes, (Glave) a= 5 oe a eee as Cuba, North America LOMminor- NOM os- 222 ee eee St. Vineent, Grenada, Trinidad 20.. schwaral Notm-_-_... = ..5-- 2 a Cuba 21.. haitiellus..Dgp)..-=2 23 ee ee eee Hispaniola 22. intermedius Hits22. 2222 Soot Soe ee ee ee Trinidad, South America MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 567 Osorius Latr. (XXV)—Continued. Comat D ATO leeIN O GMa ae eee een ee ee ee Jamaica Ae OO GINN AD ) SANT | em ee ee ee ee SE ee ee ee Jamaica eA CV COD SIN © UE eee ae ae ee ee eee a Puerto Rico 2 Game UTI COM SIS ial ayy sen ee ee we ee ee eek Jamaica smeSCeLSie Wee sees eae oe ae Cuba to St. Thomas, Central America SoC VusES Lee Ey | ayy Te rene i ee eet ee ee ee ee ee Caymans AO EWG DC Iulem Eph Ke Wyile eas ear ee e 2 S oe 2 oe ee Pe Trinidad SOMULITLALISRIbIK Wire tae te ee eee ee ee ee Trinidad CMO RTS UILISMON O Cea hee sees et eet PP ee eee et ek Cuba micros Notm. SON EVES baileys tees en ee EE aes ee eee Trinidad Som CLOPLCDUS MS UW 2s es a a Trinidad STYLOPODINAE Stylopodus Benick (XXVI) SRS ry aT CMT (aM) a oe ee pt ee een ra ree ale St. Vincent Zeimpressus| (Shp) == 22-22222 Trinidad, South America, Central America SAM ENUINVE ETL (OTN) es A ae En al a ee St. Vineent 4 Aeviventris: (Cam) 22 22a ee es ee St. Vincent, Grenada 5. punctatus (Hr.)--- Trinidad, Curacao, South America, Central America GerampousekiM@@bTre)\. 2225 2 2 See SE ee ee Cuba bierigi (Brg.) STENINAE Stenus Latr. (XXVII) A emCLATUG Se SW ae So ee ee ee Se a en Trinidad Zam CH Us key aa, ee ee SL Re Ie eee Jamaica eSemeIG ULL ONIN SES UWI ens ers ae ee Se ah at ee Bahamas EO OL UULS Reeds Ul Kayser a tr es Ln ee Nee eee Ae Cuba Drs DAK Gem ts Meee eres NER A ha oe ee eee Cuba Gre | AMMA CENSiS ails ley ee oe ee Se eee Jamaica smAI ULL S Tw ee we ee s St. Vincent, South America SSMU SUIS GLTNUES, SSL Me see ee ee Trinidad SM AILICN SIS ES LW it ease ie oe Ae ee Hispaniola LOM nIUtChleni eb kw o- = ee s ese 2 bay ee ee ie ea ae Cuba MIST OT TISO Mite KW ee ee Trinidad LAN, «BVA AEH) he 8 Ea ee cee gn ECR ee eG SB ee Trinidad ide yemm LY, CSTE Le > Rc Vy glia seer ee lala eB tel Cuba EES CUD ATM Saris Wwe ee a ee ee ee Cuba aL Oe hed TIT SOT > Wales oe ae ee et ee at a Trinidad NG mC ane OMe awash oa 5 Se ee Ee Eee ee Trinidad LEC: LNT ECSTANS| ox CFE 0 ee ee i a a a ee ee ee ener ee oy tly eee Grenada A SoS DAML ONS a UW Dee eer re a ee Cuba, Hispaniola iS CUUCUSIS MME sae ans 0 a ee ee eee Re Ue SA ee Cuba PU MAN LUNCH SIS py Clee a= 2 kee ee eee Guadeloupe EUAESTHETINAE Exoctavius Brg. (XXVIII) RR DELMUCe Zit Peek swe a oe eee ee Cuba Tamotus Schauf. (XXIX) NERLCINOLALIGES CHAULS ere a-: Sat eos eR OF oe ee ee se es Cuba Stenaesthetus Shp. (XXX) i. iMiMmMAn eis buUsE (Ds) 2-2 ane saa eee St. Vincent, South America 568 BULLETIN 182, UNITED STATES NATIONAL MUSEUM PAEDERINAE Thinocharis Kr. (XXXT) i; pound: BlKkwiresn-se2+ soe. eae ea ee ees Trinidad 2. balkeri. i(Csys)is= = se a ae re es Cuba 3; Chapini” Blkwos) 22) 22a ae oo eee Jamaica, Hispaniola 4. fascina Came. 222 ee a ee ee ee Grenada, Trinidad Bicsmithiv Camis tt. ee eee eda oh ie a een ne ee St. Vincent, Grenada 6. exilis (Hr.)------ Cuba to Trinidad, South America, Central America, North America atratula (lynch) fragilis (Shp.) minuta (Shp.) delicatula (Csy.) pertenuis (Csy.) Lithocharis B. & L. (XXXII) al Se SOLROECU] ae Kites wee oe ee a ae ee ee Jamaica to Grenada, ete. 2. ochraceah( Grays). == 2222) eke Jamaica to Grenada, South America, Central America, North Ameriea, ete. rubricollis (Gray.) obscurus (Steph. ) brunniceps (Fairm.) fastidiosa F. & G. alutacea (Csy.) quadricollis (Csy.) iS. SCCUNCA Dllkwit 25-2 ee ee eee Jamaica to St. Lucia AP SV OLAS. 2) bs Whe ae ees ee ee ne Jamaica ye MOM OA CIA, COS ae ee ee eee Jamaica 6. limbata Hr____ Hispaniola to Trinidad, South America, Central America Te CLOTS ALD SE To i Cuba to Trinidad obfuscata Cam. ST POSt Cate: CHa oe ee Se ee Se ee Puerto Rico THOT Gg! TBS Nyy sees ae a aa ee er re eee Cuba Aderocharis Shp. (XX XIII) OS CUDETISTS BU voit ss a ae ee re re ee Cuba DCU LEDS. ES Wyatcea crac cs ce es ea en ee Trinidad Fa Paek 610) 6) 5 oak (O60 PSR ee PUR aS INS oS eR WS BN St. Vincent, Grenada obscurior Cam. 4 OSUGURALS isl wes 8 ese aa ee ee Cuba to Puerto Rico Stilomedon Shp. (XXXIV) I insularam (Cani.) ois a = ee Cuba, Jamaica, Hispaniola 2. connexum (Shp.)_--. Cuba to Trinidad, South America, Central America S$. 2UGantl Bl Kyi ee ee ee ee ee Cuba to St. Croix Sunius Steph. (XX XV) Hypomedon sensu Blkwr., 1939 OSU CVA RLS Wy Tee ee re ca ee es ee Puerto Rico 2. ferrugineus: \((Br¢.) 2222-6 ee oe sy ne Cuba oy OCCIDITRUS~ (Bre: ) S20 2 Se ee eee Cuba 4. aarlingtoni, Bikwt 2222c2 Bl kyr a ee eee Hispaniola, Puerto Rico $.. cubanus: Blkwit2 a2] a22s52se2seo coon ae eee ee Cuba 4. trinitatis (Blkwrostsecn 22 Soa ee eS eee Trinidad 5. ‘subterraneus, Blkwr2 2 2-22 2 ae ea ee a ae Cuba G6. darlingtoni’ Blkwr- = 23-3 ees Jamaica ac Aamsicensis: (Blkiwitiiis<=-2..5 S22 e5 se ee Jamaica 82 biericil@ Bik wire a ee ee Puerto Rico Osiivines hil ==. ae ee Bahamas, Cuba to Puerto Rico 10 ghispaniolus! (Blkwr- ss. .ac—— 225 -- eeee e Hispaniola Ti. aeudyol Bres.o = Wo ee eee eee Cuba 12. “insigniventris, Bohr. -=2222. eee Guadeloupe 13. Schwarz (Blkiwt2s=. ssa S55 as oo oe ee eee Cuba Araeocerus Nord. (LX) j. parvipennis Bret 5-22-2522 S222 22 aoe See eee Cuba Palaminus Er. (LXI) isanacoreta: Bre. se. ie See ea Cuba 2. /anterus#bnhrs_ = = 222 Sees 2 ee eee Guadeloupe S$. permudeziv Bros. 2 ee See eee eee Cuba 4 :peralutaceusi Bre 3-2. = = = ee eee Cuba 140 pilont (Birger a ee ee eee Cuba 15S procerusz NOtM== = 2 se er eee Puerto Rico 16) pusillusMNOtm: 222 <2 22-2 a= ee ee eee Pue-to Rico af rosariensis' Bro?o=- 22-52 == 3 a ss eee eee eee Cuba 1S8"scitulus® Note 4-5 22s 2s see eee a eee Puerto Rico 19isvariabilis ‘Hrs W-- = -==—-=--==— Jamaica to Trinidad, South America STAPHYLININAE Paederomimus Shp. (LXITI) phar WCETET CUS cS LI I a a Trinidad 2. eribricollis (Er.)_-. Grenada, Trinidad, South America, Central America trinidadensis Bnhr. So. anterjechus Snhti. 2. Sloe ee oe ee St. Vincent smithi Cam. 4, LUStTaLIS GH) eee Puerto Rico, St. Thomas insularis Bnhr. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 573 Philonthus Steph. (LXIII) fe hepaticus: Hrs + —- oes oe Ka Bahamas, Cuba to Trinidad, South America, Central America, North America, ete. vilis Er. orphanus Er. nanus Melsh. cinctutus Melsh. palleolus Melsh. rufipennis (Sol.) varicolor Boh. pyropterus Kr. paucillus Sols. parvimanus Shp. DethermarumeAUbG=2=-2-—ses2 58 oe Se ee Jamaica to Grenada, South America, North America, ete. exilis Kr. angustatus Kr. pygmaeus Kr. fuscolaterus (Mots.) inclinans (Walk.) Si ventralisn(Gravy.) 22-5225 2. ee ee LS Cuba to Trinidad South America, North America, ete. anthraxv (Grav.) immundus (Gray.) celer (Grav.) picicollis Steph. fulvipes (Steph.) rotundiceps (Steph.) fortunatus Woll. prozimus Woll. Ae audantioBlkwrte-=* == S2. Sa a ee Hispaniola Ha aiscoideus: (Grave) sse2=-=--s= Cuba to Grenada, North America, éte. suturalis (Marsh.) testaceus (Grav.) lepidulus (Steph.) conformis (B. & L.) ruficornis Melsh. rufipennis Gerh. gerhardtianus Scheerp. 6: figulus)Hr=+-ses"= Cuba to Trinidad, South America, Central America amazonicus Shp. (lONnPicorMMise Steph! - 2 = se Hispaniola to Grenada, South America, Central America, North America, ete. scybalarius Nordm. fuscicornis Nordm. feralis Er. promtus Er. varians Fairm. algiricus Mots. pedestris Walk. asemus Kr. perplecus I. & G. fumosus Sols. 574 Philon BULLETIN 182, UNITED STATES NATIONAL MUSEUM thus Steph. (LXIII)—Continued. 7. longicornis Steph.—Continued. laetabilis (Oliff) linkei Bnhr. rubromaculatus Bnhr. piceicornis Grid. 8. varians (Payk.)---._ Cuba, St. John, St. Vincent, North America, etc. bipustulatus (Grav.) aterrimus (Marsh.) nitens (Gray.) opacus (Grav.) intaminatus (Steph.) lituratus Steph. nigrans (Dej.) punctiventris Steph. unicolor (Steph.) bimaculatus (Marsh.) costatus Baudi incompletus (Hochh.) scutatus Epp. alpigradus M. & R. proteus Everts piceicoxis Grid. fuscicovis Scheerp. 9. flavolimbatus Er_____-~ Martinique, Carriacou, South America, Central America, North America apicipennis Lynch 10, caribacus “Bre. a= 22222522 = ek ead ee ee ee ee —- Cuba 145. Silvaticus Cam 22 ohs22252. Se. eee Jamaica 72, waterhousei Cam 232) 2h 25-3 ae St. Vincent, Grenada AS atrinitatis Bl kw. e222 — eS a eee ee Trinidad 14.havaniensis (Maps) t see ashes oa eee ee Cuba to Antigua trepidus Er. obscurus Fvl. cubensis Brg. MS. Belonuchus Nordm. (LXIV) 2 2 3. 4 hispaniolus'* Blkwl=2.222 22-2 22=--- liliputanus Ramb., MS. 2. OGUlEnIS=Blikwree. 2 eee Acylophorus Nordm. (LXXY) ark wnhnd eH tharteri Bnhr., MS. Atanygnathus Jacobs. (LXXVI) i. heterocerus Cam----.22-~ - 2. laticollisi( Mrs) =~ = 2 trinitatis@Bllnwre 225 22222 ee f AnSUlAGUS One ee ee . aenescens Blkwr_-_-------------- , natuey *Biee ee ee ee eee , metallicus: Bikwr-----—-- 222 . darlingtoni Blkwr__----_------~~- Ss ee a St. Vincent, Trinidad Trinidad, South America, Central America Trinidad, South America, Central America Cuba to Trinidad, South America, Central America, North America ce si aA oo eh eo ee Jamaica notes sose = Shee ee es Cuba Bahamas, Cuba to St. Vincent ee eee ore Puerto Rico to St. Vincent opr de ea Ww Cuba, Jamaica er ee ee Cuba, Jamaica 2 ee ae 2 ee Trinidad aoe = wie, Jee een Trinidad 22h Wace oe ae eR OR Cuba, Isle of Pines 2 cance SNe! ees Hispaniola bes ee Se ee Se Cuba hdd ee ee Ze => Hispaniola 2 ee oaee en ee ute Prinidad ae oa ee St. Vincent, Grenada ee _._._ Cuba to Trinidad MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 577 Xantholinus Dej. (LXXVII) RmOgkie vip lewis eee a8 ts a Hispaniola, Puerto Rico SAMOS UIVCUSM@ Ams 2h eS ee el ae Se Mustique, Grenada Seruscipennisn @amet: Hi tyier Lei UE Bete Ee) Jamaica 4. attenuatus Hr______ Cuba to Trinidad, South America, Central America OHUMET AIGA ee ee et I Cuba to St. Vincent puncticeps DuVal punctiger G. & H. Givcubensisveblicwy si. 225 3. S520 Se 5 ep eet | ee ee Cuba (Coy Garline tomnie Blinwi tees oe oe See ee Hispaniola SapculomuSt bs kewirse eee oe SP soe ee Hispaniola Svaudanitiy (Blow ee «28 ieee Hispaniola OR IMOLMISON web Twila see ete Jee 2 ee Pes, a Trinidad dG Ss ewes ee eer ee et ie ee Hispaniola OAD CA UL YAl eis Livy Ae ee ek Cuba, Hispanola, St. Croix Soe S OMG ATU Sia Es kya Seas een eee eee ee See ee Guadeloupe Bayes) PIN N COT ST Sp BS Meayy ste so ere ne Se ae Jamaica By GRUROUEIGUUS Ey OW Uns ener ee oe SO ie ee a Cuba GS NRICONS i pee eee ee eee Puerto Rico to Trinidad, South America ij. hydrocephalus) Wylie ane a Trinidad, South America Thyreocepkalus Guer. (LXXVIII) fe SUDENS_Shps2-< eee St. Vincent, South America, Central America Plochionocerus Shp. (LXXIX) iebrachyplelus Shp! a= .- se ane eo eeee Trinidad, Central America Leptacinus Er. (LXXX) CSPOT Ene CAN ees ee ee ee ee es Se we ae St. Thomas apicipennis Bnhr. 2. parumpunetatus (Gyll.)-------_ Cuba to Grenada, North America, ete. longicollis (Steph. ) ampliventris DuVal subimpressus (Dej.) radiosus (Peyr.) pallidipennis Mots. tricolor Kr. flavipennis Kr. amissus (EF. & C.) breviceps Waterh. papuensis Fy]. sardous Fiori rubricollis Reitt. fauveli Cam. Lithocharodes Shp. (LXXXI) iemclaviscapas (@amisyee soe Pao kee Be Grenada, St. Vincent claviger Cam., MS. CULT LOTiil mE Keypress rae eee SA RS ea i ae ee Cuba Sms etie ula tates Uw == 2 Gk oe 2 Se 2 et ee Hispaniola APA COLI ES cpu ere ee a ee ee a ee Se Trinidad HslamMbpousektvblkwir, (BTSs MS.) cee eee oo een ae a ee Cuba Sr ETUC ANE Sys) ayy. Tee ae i ee ie ee eee Trinidad (epUMColLomo (Sams )\ee one ee eee see RE ee ee Jamaica Microlinus Csy. (UXXXII) TE DUSIOMUILeOs) rea eee eae Jamaica, Trinidad, North America Stenolinus Brg. (UXXXIII) ae FN VEO WEL CENT Ss ie eee er cena i Le Rg ee Cuba 578 BULLETIN 182, UNITED STATES NATIONAL MUSEUM Oligoninus Csy. (LXXXIV) i. hubbandi¢Blkwrte=..—. =. -ss25-522-22—sa senses Se Montserrat 2.darlinetonif@Blkwr—--- = --s.-—. 2-5 ee ee Hispaniola Syettlarius. (r.) 2. St. Vincent, South America, Central America 4;,testaceipennis!«(Mwls) 2232 62522525 st ones Ee tee Cuba pallidipennis (F v1.) 5. variabilis; Blkwr=— = 25 225 Sao 2- Sees = a EE Cuba, Hispaniola TACHYPORINAE Leucoparyphus Kr. (LXXXV) dee sil phoides™ (iy) 2s=2=s== === Puerto Rico to Trinidad, North America, etc. suturalis (Panz.) marginalis (Grav.) limbatus (Grav.) pictus (Er.) geminatus (Rand. ) marginicollis Kr. limbifer Mots. koreanus Buhr. Coproporus Kr. (LX XXVI) VA Shar pie @ aie sec ee srk es ies eee Se Antigua to St. Vincent 2 sapicalish(hrs)=*-.cs2522-2 oul eee Cuba to Puerto Rico eo: hepaticus (ir) 22s 122s Cuba, Puerto Rico, Trinidad, South America, Central America, North America converus (Er.) ignavus Shp. inflatus (Horn) 4. trinitatis VBlikwrts ooo. 2. soe eee ee ee Trinidad Be webert SB likwirl ee een ee te ee ee ee Trinidad 6; "cacao Blikwi 2-22 Soa eS ee St. Lucia, Trinidad (oatlavicollis Scribas==..—5 se s—=== =a eee Cuba, Jamaica, Puerto Rico, South America, Central America nitidulus (Er.) strigosus (Shp.) 8. (ebonus®Blikwr! 22) see oe ee Jamaica to Grenada piceus (Er.) 9) pulchellus: (hr) 2222 —— 2a Cuba to Trinidad, South America, Central America, North America cumanensis Seriba infimus (DuVal) distans Shp. 10.-Cubensisy 1(@s ya) Sa Oe ee ee eee ee ee Cuba (CwGensatarC amiss 22262 ae Se a St. Vincent 8; TauveliG@ am 2222 Be an eee eee ee ee ee St. Vincent flavicornis Cam. 9: ‘granifera. Cam 2-2 252) ao ee ee eee Jamaica 10., hydrocephalasCam-2==—- 2-222 255 2 es eas St. Vincent, Grenada 21. jamaicensiss Cam’ —--— 222-_ 2-- 2 ee ee ee Jamacia 12; laxatae Came: t 22 Oke ee ee St. Vincent, Grenada 134 Oblitay Shpies==ss Soe es See ee St. Lucia, Central America 14. obseuripes' Cam2-.= 22-2. ee Jamaica THe persimilish@ am == sets= een Se eee St. Vincent 16, piceicollisyiCam=es— 2-2 = = eee St. Vincent, Grenada (on DUD CLAM ORM See 2 a es ee ae ee ee ea Hispaniola 18) Quassa Shp 46222 — 82 en ee eee Grenada, South America 19: nufula) Cam 2 eee A a St. Vincent, Grenada 20; ‘smith ‘Cam 222322. oe ee St. Vincent, Grenada 21, sparsepunctata® CaMie-=--5. 22a. ane eee aaa Jamaica 22. aplendidula: Mrs2: Se St. Thomas, South America Diestota M. & R. TPCADICRISUE Oe iN es ee St. Vincent, Grenada puncticeps Cam. 2sperata, Shp-s 222 - + Cuba to Grenada, South America, Central America 3. laticornis Shp_------- St. Vincent, Mustique, Grenada, Central America Coenonica Kr. if. puncticolliSektts=< ==) --22 232 eas eee St. Vincent, Grenada, ete. indica Mots. cribriceps Shp. MONOGRAPH OF WEST INDIAN STAPHYLINIDAE 581 Placusa Er. AT panglignCam sates keto Ee es ee eel St. Vincent DemneLenorastern@am <8 oo 2s ae ee ete St. Vincent Ser COMECLOMIBE akOGw N= = — Ao oe cet bop ic St. Vincent insularis Cam. 4s luetuosae@ame ss = a os oe ee St. Vincent Silusa Er. deeGiversicollise Came == 52 oso oe ee St. Vincent, Grenada ome Lene lag Camis se = sie Ss oe Sea ee St. Vincent Theeturella Cam. Min sidiosau Camas <> 2 52 35 a te St. Vincent, Grenada Bolitochara Mannh. jescincticastra (Bie <2 23a ee i ee ee Cuba Xenobiota Brg. lphernhaveri (Bry ss 4 set a a oe a nt Cuba Euvira Shp. CUD AT AEST eee a ae ee Be SN eee A Cuba Deena VINCS Dies > 3 22 Se oo a ee eee Cuba Swinsnlanrisy Cam... — ooo sen ek ee oe a ah tae St. Vincent Falagria Sam. HePDaASiventrisn (Cam, )i--22 22-25 ee ee Jamaica 2ceepnalotes $CaMmet:< 49... 5 ho se ee a ed Grenada 8. concinna Er__ St, Vincent, Grenada, South America, Central America, ete. parae Shp. fovea Shp. longipes Woll. currag Shp. Anise Shp 22 eo 8 oe a ee Jamaica to Grenada, Central America Meronera Shp. i Valbicincta: ((br;)i22-.-.-2 === Cuba, Jamaica, Grenada, South America Central America Neolara Shp. sevalboruibatad Clin) 228 = 2 as Grenada, South America Peg GUY OEE NOEEY OPEN a Cuba Pseudognypeta Cam. Ug joel. (OF hn Jamaica Gnypeta Thoms. ie DASALISMO@ ams meoNe oan Soe eee ee Jamaica, St. Lucia maxbasiventris, Cam:.2. 22... 2. ee les Grenada Seehra cls aS hpess 2 = sae er) See ee Jamaica, Central America 4-sSanctae-luciae, Cam:=== 0-222 ssa eo ee St. Lucia Atheta Thoms. eA CCOC CHS OAT esate ee SS oe eee Mustique, Grenada Dal pinennign Cama 2 2e 8 ol oe Oa eee Mustique, Grenada SSA LOTTA ta GH) es Hispaniola to Grenada AeA OMA De CAM os ape ker RS 2s i ee Jamaica, Hispaniola Peanciilarum i 96% see I> ice Bee Sir Uhe 8 18, ON, 2 : Pie ran oe « aw te ea 74 Yt 2? : mit - SS CS - = : ITHSONIAN INSTITUTION Ss UTIO 9088 01421 ” 3 i 2666