CANOTIA

QK

188

.C44

2012

Volume 8, issue 1
Contents

Checklist of Vascular Plants of Organ Pipe Cactus National Monument, Cabeza Prieta
National Wildlife Refuge, and Tinjas Altas, Arizona

Richard Stephen Felger, Susan Rutman, Thomas R. Van Devender, and

Steven M. Buckley 1

April 2012

Vascular Plant Herbarium
School of Life Sciences, Arizona State University

CANOTIA

(2R

i m*»

XdU

Editor: Leslie R. Landrum
P. 0. Box 874501
School of Life Sciences
Arizona State University
Tempe, AZ 85287-4501
(les.landrum@asu.edu)

Associate Editor: Orbelia R. Robinson
Botany Department
California Academy of Sciences
875 Howard Street
San Francisco, CA 94103-3009
(orobinson@calacademy.org)

Production Editor: Scott T. Bates
University of Colorado at Boulder
Rm. 318-CIRES Bldg.
Boulder, CO 80309
(scott.thomas.bates@gmail.com)

Canotia publishes botanical and mycological papers related to Arizona. These may include contributions to the
Vascular Plants of Arizona project, checklists, local floras, new records for Arizona and ecological studies. All
manuscripts are peer-reviewed by specialists. Acceptance for publication will be at the discretion of the editor. At
least 30 printed copies of each issue are distributed to libraries in the United States, Europe, and Latin America.
Anyone may download copies free of charge at http://www.canotia.org.

Canotia is named for Canotia holacantha Torr. (Celastraceae), a spiny shrub or small tree nearly endemic to Arizona.

ISSN 1931-3616

MERTZ LHSRAR t
NEW YORK
tsOYAMlOAL

GARDEN

CHECKLIST OF VASCULAR PLANTS OF ORGAN PIPE CACTUS NATIONAL
MONUMENT, CABEZA PRUT A NATIONAL WILDLIFE REFUGE, AND
TINAJAS ALTAS, ARIZONA

Richard Stephen Felger1,2, Susan Rutman3, Thomas R. Van Devender1,2, and

Steven M. Buckley4,5

’Herbarium, University of Arizona, P.O. Box 210036, Tucson, AZ 85721
2 Sky Island Alliance, P.O. Box 41165, Tucson, AZ 85717

3Organ Pipe Cactus National Monument, 10 Organ Pipe Drive, Ajo, AZ 85321

4National Park Service, Sonoran Desert Network, 7660 E. Broadway Blvd.,

Ste. 303, Tucson, AZ 85710

^School of Natural Resources and the Environment, University of Arizona,

Tucson, AZ 85721

ABSTRACT

The contiguous Organ Pipe Cactus National Monument, Cabeza Prieta National
Wildlife Refuge, and the Tinajas Altas region within the Sonoran Desert in southwestern
Arizona have a vascular plant flora of 736 taxa (species, subspecies, varieties, and hybrids)
in 420 genera and 94 families. Elevation and ecological diversity decrease from east (Organ
Pipe) to west (Tinajas Altas) while aridity increases from east to west, all correlating with
decreasing botanical diversity. Organ Pipe Cactus National Monument, which includes an
ecologically isolated Sky Island of dwarfed woodland rising above actual desert, has a flora
of 657 taxa in 395 genera and 93 families, of which 1 1 percent (72 species) are not native.
Cabeza Prieta National Wildlife Refuge has a documented flora of 426 taxa in 266 genera
and 63 families, of which 8.8 percent (37 species) are not native. The Tinajas Altas region
has a flora of 227 taxa in 164 genera and 47 families, of which 5.3 pecent (12 species) are
not native. Non-native species make up 10.7 percent (78 species) of the total flora of the
three areas, but among these only 7 percent (52 species) are established in the flora area as
reproducing populations. Six non-natives have become invasive species that pose serious
threats to the native ecosystems: Sahara Mustard ( Brassica tournefortii). Red Brome
(. Bromus madritensis subsp. rubens), Buffelgrass ( Cenchrus ciliaris), Arabian and
Mediterranean Grass (Schismus arabicus and S. barbatus), and Tamarisk ( Tarnarix
chinensis).

Key words: Sonoran Desert, flora, Arizona, Organ Pipe Cactus National Monument,
Cabeza Prieta National Wildlife Refuge, Tinajas Altas, non-native species, invasive species

INTRODUCTION

In this publication we provide a checklist of the vascular plants for the
contiguous protected areas of Organ Pipe Cactus National Monument, Cabeza Prieta
National Wildlife Refuge, and the Tinajas Altas region (the flora area) of
southwestern Arizona (Fig. 1). Detailed background information on these regions,

Checklist of Vascular Plants of Organ Pipe Cactus National Monument, Cabez Prieta National
Wildlife Refuge, and Tinajas Altas Region, Arizona. CANOTIA 8: 1-53, 2012.

©2012 R. S. Felger. Non-commercial, personal, educational, or scientific use is freely
granted without asking permission; you are requested to cite this work.

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CANOTIA Vol. 8

2012

including specific geographic information in a regional gazetteer and their biota, can
be found in Broyles et al. (2007), Felger and Broyles (1997, 2007), and Felger et al.
(2007a).

A specimen-based listing for the flora area and surrounding regions is provided
by Felger et al. (1997, 2007b). Flowever, in view of significant new findings,
taxonomic revisions, and corrections we offer a current and concise checklist for the
flora area — Organ Pipe to the Tinajas Altas region. This checklist is specimen-based
with 736 taxa (species, subspecies, varieties, and hybrids) in 420 genera and 94
families (Table 1). Elevation and ecological diversity decreases from east (Organ
Pipe) to west (Tinajas Altas) while aridity increases from east to west, all correlating
with decreasing botanical diversity. This listing is preliminary to our larger work on
the flora and vegetation of the flora area in preparation. Although there has been
extensive documentation of plants from this region, diligent botanists should be able
to discover some additional species.

The present flora area is part of five major, contiguous protected areas and a
sixth de facto area that make up the heart of the Sonoran Desert, here called the “Six
Dry Borders Areas”: 1) Organ Pipe Cactus National Monument, 2) Cabeza Prieta
National Wildlife Refuge, and 3) Sonoran Desert National Monument in Arizona; 4)
Reserva de la Biosfera El Pinacate y Gran Desierto de Altar in Sonora and 5)
Reserva de la Biosfera Alto Golfo de California y Delta del Rio Colorado in Sonora
and Baja California; and the 6) Barry M. Goldwater Range. The Goldwater Range in
Arizona is not a protected area per se but generally is managed as one by the U.S Air
Force and U.S. Marine Corps. Spanning 210 miles (338 km) from San Felipe, Baja
California, to just southwest of Phoenix, Arizona, these six areas cover 7,515,221
acres (3,041,302 ha), or 1 1.7 percent of the Sonoran Desert, making them the largest
zone of contiguous protected desert anywhere in the Americas. This bio-network
supports a flora of at least 865 vascular plant taxa (species, subspecies, and varieties)
including 91 non-native species in 448 genera and 107 families (Felger et al. 2007a
and b, and new records including Cuscuta spp., Mentzelia longiloba var.
pinacatensis J. J. Schenk & L. Hufford, Psorothamnus fremontii, and Salvinia
molesta D. S. Mitchell in Salviniaceae). This flora includes more than one-third of
the total flora of the entire Sonoran Desert, a region covering about 100,000 mi2
(310,000 km2) in five states in Mexico and the United States (Shreve and Wiggins
1964).

Table 1 compares the flora of the three protected areas of Arizona, and the
combined Six Dry Borders Areas. The three areas of the present floristic listing have
a collective size of 1,270,699 acres (514,234 ha). The Six Dry Borders Areas, about
six times larger than the included three areas of the present floristic listing, have a
total flora of 860 species, an increase of only about 15 percent above that of the
present flora listing.

ORGAN PIPE CACTUS NATIONAL MONUMENT

The Monument covers 330,689 acres (133,825 ha) and ranges from 980 to
4808 ft (299 to 1465 m) elevation. The majority of the Monument falls within
Shreve’ s (1951) Arizona Uplands subdivision of the Sonoran Desert. The higher

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

3

elevations of the Ajo Mountains, along the eastern margin of the Monument, support
an ecologically isolated Sky Island rising above actual desert vegetation. At these
higher elevations one finds a dwarfed woodland that includes Juniper ( Juniperus
arizonica ), Scrub Oak ( Quercus turbinella), and Rosewood ( Vauquelinia
calif ornica). The southwestern comer of the Monument transitions into the Lower
Colorado Valley subdivision of the Sonoran Desert (Felger 2000; Felger et al. 1992).
Works on the vegetation of the Monument include those by Brown and Warren
(1986), Lowe et al. (1992), Mulroy (1971), Steenbergh and Warren 1977, Warren
and Anderson (1987), and Warren et al. (1981). The National Park Service is in the
process of updating existing maps through its Inventory and Monitoring program.

The first checklist of the Monument plants was completed by McDougall (ca.
1940). That list and one prepared by Phillips (1967) were based on field experience
and specimens. Subsequent checklists compiled by Adams (1971) and Jordan (1975)
contained undocumented and questionable listings. Janice Bowers’ (1980) flora was
the first critical, specimen-based flora of the Monument and serves as the basis for
all future botanical work in the Monument. The non-native flora was discussed by
Felger (1990) and a detailed flora of Quitobaquito with ethnobotanical information
was prepared by Felger et al. (1992). Additions to the flora were published by
Pinkava et al. (1992) and Felger et al. (1997, 2007b) provided a comprehensive
annotated flora.

The total present flora of the Monument includes 657 taxa (species,
subspecies, varieties, and hybrids) in 395 genera and 93 families. Non-native species
account for 1 1 percent of the flora (72 species), although many are not established as
reproducing populations.

CABEZA PRIETA NATIONAL WILDLIFE REFUGE

The Refuge covers 860,010 acres (348,034 ha) and ranges from 680 to 3293 ft
(207 to 1004 m) in elevation. The entire Refuge is within Shreve’s (1951) Lower
Colorado Valley subdivision of the Sonoran Desert. A preliminary, specimen-based
flora of the Refuge by Simmons (1965, 1966) was based on first-hand observations
and specimens. Phelps (1981) produced a checklist of the Refuge, Tunnicliff et al.
(1986) compiled a list for the Barry M. Goldwater Military Range that included the
Refuge, and Felger et al. (1997, 2007b) provided substantial additions for Goldwater
Range. While the flora is well documented, many remote areas, especially the
rugged mountains are expected to contain unrecorded range extensions, such as the
discovery of Scallop-leaf Sage {Salvia vaseyi) new for Arizona by Cain et al. (2010).

Detailed vegetation studies were conducted by Malusa (2003). We list a
documented flora of 426 taxa in 266 genera and 63 families. Non-native plants (37
species) make up 8.8 percent of the flora, although some are not established as
reproducing populations.

TINAJAS ALI AS REGION

This is the most arid portion of the flora area and adjoins the western margin of
the Cabeza Prieta National Wildlife Refuge. The core area is the 80,000 acres
(32,375 hectares) of the Tinajas Altas region encompassing the Tinajas Altas

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Mountains from near the Cipriano Pass to the Mexican border at Frontera Canyon
and the south end of the Lechuguilla Valley drained northward by Coyote Wash and
southward by La Jolla Wash. Elevation ranges from ca. 950 to 2764 ft (ca. 290 to
842 m). The region is within Shreve’s (1951) Lower Colorado Valley subdivision of
the Sonoran Desert.

Felger and Van Devender (2012) provided an abridged checklist of the flora in
a work on the Tinajas Altas region by Broyles et al. (2012), and that information is
included here with some revisions and new discoveries. Felger et al. (forthcoming)
provide a comprehensive treatment of the modem and deep history flora of the
Tinajas Altas region. Annotated checklists for the Goldwater Range included Tinajas
Altas plants (Felger et al. 1997, 2007b). We document a flora of 227 taxa (including
2 hybrids) in 164 genera and 47 families. There are 12 non-native species in the
flora, which is only 5.3 percent of the total flora. We include some plants from
immediately adjacent localities that are expected to occur in the core area.

The boundaries for this area are those of the Tinajas Altas addition to Cabeza
Prieta National Wildlife Refuge as proposed by U.S. Fish and Wildlife Service
(1970a & b) and as incorporated within the Cabeza Prieta Refuge for a few months
in 1975 by the Department of the Interior (Tunnicliff et al. 1986: 3.14). Those
boundaries would have extended the northern refuge boundary westward to a point
about one mile west of Cipriano Pass and then southward to the U.S.-Mexico
border. That area encompassed land in townships 12, 13, and 14S in range 17W and
townships 12 and 13S in range 18W, totaling approximately 80,000 acres (U.S. Fish
& Wildlife Service 1970b) and included Raven Butte, the Tinajas Altas Mountains,
and portions of the Lechuguilla Desert and Davis Plain, but the land was not
congressionally included within the refuge. In turn, because of its special biological,
historical, archaeological, geological, and scenic values, the Bureau of Land
Management in coordination with the Department of Defense subsequently
designated much of the former addition as an Area of Critical Environmental
Concern (ACEC) of irregular configuration comprising about 60,500 acres (U.S.
Departments of the Air Force, Navy, and Interior 2006), and later as a Special
Natural Interest Area of comparable though not congruent boundaries (Range
Management Department 2007).

NON-NATIVE SPECIES

Approximately 10.7 percent of the total flora, or 78 species, are not native to
the region (see Felger 1990, 2000; Felger et al. 2007b; Van Devender et al. 2009;
Wilson et al. 2002). Among the total non-native flora, only 7 percent (52 species)
are established as reproducing populations and a number of these occur only in
disturbed habitats. Twenty-seven species do not have reproducing populations in the
flora area: these include “strays” such as Shepherd’s Purse ( Capsella bursa-
pastoris), Safflower ( Carthamus tinctorius), and Wheat ( Triticum aestivum), as well
as others that have persisted from cultivation, such as the Figs ( Ficus carica) and
Pomegranates ( Punica granatum ) at Quitobaquito (e.g., Felger et al. 1992), and/or
plants extirpated from the flora area such as Watercress (. Nasturtium officinale).
Among the fully established non-natives, a few have become invasive species that

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

5

pose serious threats to the native ecosystems, e.g., Sahara mustard ( Brassica
tournefortii). Red Brome ( Bromus madritensis subsp. rubens ), Buffelgrass
( Cenchrus ciliaris ), Arabian Grass ( Schismus arabicus), Mediterranean Grass
( Schismus barbatus ), and Tamarisk ( Tamarix chinensis) (Felger 1990, 2000; Felger
et al. 2007b; Van Devender et al. 2009).

THE CHECKLIST

Plants in this checklist are grouped as Pteridophytes (ferns and lycopods or
“ferns and fern relatives”), Gymnosperms (cone-bearing plants), and Angiosperms:
Magnoliidae (magnoliids), Eudicotyledoneae (eudicots), and Monocotyledoneae
(monocots). Within these categories the plants are listed alphabetically by family,
genus, species, infraspecific taxa (subspecies or varieties), and hybrids. The
accepted scientific names are in bold.

Nomenclature in the checklist utilizes our best interpretation of current
systematic knowledge. We generally follow or interpret the taxonomy of the Flora of
North America, Vascular Plants of Arizona (Vascular Plants of Arizona Editorial
Committee, 1992+), and the Jepson Flora Project (2012), and utilize Tropicos (201 1)
as reference for authors of scientific names and indications of “accepted name”
when provided (e.g., Poaceae). References supporting taxonomic interpretations
generally not included in those more general works or of special interest are cited in
the checklist with the particular taxa.

Selected synonyms of scientific names are italicized within brackets [--].
Vernacular, or common, names follow the scientific names or synonyms and when
known, or deemed worthwhile, are given in English, Spanish, and the Fiia C-ed
O’odham dialect, respectively. English language common names largely follow
those provided by the Jepson Flora Project and USDA Plants Database (201 1). The
Spanish-language names are italicized. The O’odham names are from Felger et al.
(2007b). These indigenous names are based largely on Gary Nabhan’s work in
Felger et al. (1992, 2007b). Plant family designations follow the APG III system
(Angiosperm Phylogeny Group; Stevens 2011).

A voucher specimen is cited for each of the three areas in which the taxon is
present, with the exception of some reliable sight records or published references
when a specimen was not available. Specimens cited are at the University of
Arizona Herbarium (ARIZ), unless otherwise indicated by the standardized
abbreviations for herbaria (Index Herbariorum 2009), or at the herbaria of Organ
Pipe Cactus National Monument (identified as ORPI) or Cabeza Prieta National
Wildlife Refuge (identified as CAB). All herbarium specimens cited have been seen
by Felger, although some have been seen as images. The name of the collector and
collection number is given. In cases where more than one collector is listed on a
label, generally only the first collector’s name is given. If no collection number is
provided on the herbarium label, then the specimen is identified by the date of
collection, for example, Harbison 6 Mar 1937. Additional specimens are cited in
Felger et al. (2007b) as well as the Southwest Environmental Information Network
(SEINet 2011), and are found in the regional herbaria including ARIZ, ASU, DES,
SD, and the herbaria at Organ Pipe Cactus National Monument (ORPI) and Cabeza

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Prieta National Wildlife Refuge (CAB). Richard Felger’s collections are identified
as RSF, Susan Rutman’s as SR, and Tom Van Devender’s as TRV.

Area designations are: CP = Cabeza Prieta National Wildlife Refuge, OP =
Organ Pipe Cactus National Wildlife Refuge, TA = Tinajas Alias region. Non-native
plants established as reproducing populations in the flora area are indicated with an
asterisk (*) and non-native species not established as reproducing populations are
marked with two asterisks (**).

PTERIDOPHYTES - FERNS AND LYCOPHYTES

MARSILEACEAE - WATER-CLOVER FAMILY

Marsilea vestita Elooker & Greville subsp. vestita. Hairy Water-clover, Hooked Pepperwort
CP: RSF 01-571.

PTERIDACEAE - MAIDENHAIR FERN FAMILY

Argyrochosma limitanea (Maxon) Windham subsp. limitanea [Notholaena limitanea Maxon var.
limitanea ]. Southwestern False Cloak-fern
OP: Nichol 23 Feb 1939.

Astrolepis cochisensis (Goodding) D. M. Benham & Windham subsp. cochisensis [Notholaena
cochisensis Goodding. N. sinuata var. cochisensis (Goodding) Weatherby. Cheilanthes cochisensis
(Goodding) Mickel]. Scaly Cloak-fern, Jimmy-fern

CP: RSF 93-29. OP: Goodding 455-41. TA: TRV 10 Mar 1980.

Astrolepis sinuata (Lagasca ex Swartz) D. M. Benham & Windham subsp. sinuata [ Notholaena
sinuata (Lagasca ex Swartz) Kaulfus. Cheilanthes sinuata (Lagasca ex Swartz) Domin.]. Wavy Star-
scaled Cloak-fern

OP: RSF 90-568.

Bommeria hispida (Mettenius ex Kuhn) Underwood. Hairy Bommeria
OP: 17 Dec 1939, Harbison 26317 [ SD).

Cheilanthes lindheimeri Hooker. Fairy Swords
OP: RSF 90-559.

Cheilanthes parryi (D. C. Eaton) Domin [ Notholaena parryi D. C. Eaton]. Parry’s Lip-fern
CP: Simmons 21 Nov 1964. OP: Wirt 5 Dec 1990. TA: Harrison 6575.

Cheilanthes villosa Davenport ex Maxon. Hairy Lip-fern
OP: RSF 05-267.

Cheilanthes wrightii Hooker. Wright’s Lip-fern
OP: Bowers 1003.

Cheilanthes yavapensis T. Reeves ex Windham. Yavapai Lip-fern
OP: Darrow 3863.

Notholaena californica D.C. Eaton subsp. ealifornica [ Cheilanthes deserti Mickel]. California
Cloak-fern

CP: Wiggins 26 Mar 1932. OP: RSF 88-1 35 (ORPI). TA: Reeves 5353 (ASU).

Notholaena standleyi Maxon [ Cheilanthes standleyi (Maxon) Mickel], Star Cloak-fern
CP: RSF 92-575. OP: Yatskievych 81-92. '

Pellaea truncata Goodding. Spiny Cliff-brake
OP: Bowers 1701.

Pentagramma triangularis (Kaulfuss) Yatskievych, Windham & E. Wollenweber subsp. maxonii
(Weatherby) Yatskievych, Windham & E. Wollenweber [Pityrogramma triangularis (Kaulfuss)
Maxon var. maxonii Weatherby], Goldback Fern
OP: Bowers 930.

SELAGINELLACEAE - SPIKE-MOSS FAMILY
Selaginella arizonica Maxon. Arizona Spike-moss; flor de piedra
CP: RSF 92-578. OP: Bezy 25 Oct 1964.

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Selaginella eremophila Maxon. Desert Spike-moss

CP: RSF 92-585. OP: RSF 88-1 1 3. TA: Reichenbacher 28 Mar 1981.

GYMNOSPERMS - CONE-BEARING PLANTS
CUPRESSACEAE - CYPRESS FAMILY

Juniperus arizonica (R. P. Adams) R. P. Adams [J. coahuilensis (Martinez) Gaussen ex R. P.
Adams var. arizonica R. P. Adams. J. erythrocarpa Cory, in part: New Mexico, Arizona], Arizona
Juniper. See Adams ef al. (2006).

OP: RSF 90-508.

EPHEDRACEAE - JOINT-FIR FAMILY

Ephedra aspera Engelmann ex S. Watson [E. nevadensis S. Watson var. aspera (Engelmann ex S.
Watson) L.D. Benson], Boundary Ephedra; canutillo ; ku'ukpalk

CP: Yeatts 3653. OP: Bowers 1266. TA: Hodgson 2100 (DES).

ANGIOSPERMS - FLOWERING PLANTS

MAGNOLIIDS - MAGNOLIIDEA (majority of basal dicots)
ARISTOLOCHIACEAE - BIRTHWORT FAMILY

Aristolochia watsonii Wooton & Standley [A. brevipes Bentham var. acuminata S. Watson, not zl.
acuminata Lamaire, A. porphyrophylla FI. Pfeifer], Indian-root; hierba del indio, zapatito
CP: RSF 92-700. OP: RSF 86-275.

SAURURACEAE - LIZARD-TAIL FAMILY

Anemopsis californica (Nuttall) Hooker & Arnott. Yerba mansa; hierba del manso ; va:vis
OP: Mearns 2786 (US).

EUDICOTS - EUDICOTYLEDONEAE (more advanced dicots)

ACANTHACEAE - ACANTHUS FAMILY
Anisacanthus thurberi (Torrey) A. Gray. Desert Honeysuckle
OP: Bowers 1240.

Carlowrightia arizonica A. Gray. Arizona Wrightwort; lemilla
CP: Lehto L23542 (ASU). OP: RSF 88-266.

Dicliptera resupinata (Vahl) de Jussieu. Fold-wing; alfalfdla
OP: Lockwood 163.

Justicia californica (Bentham) D. N. Gibson [Beloperone californica Bentham]. Desert
Hummingbird-bush; chuparrosa\ wipisimal

CP: Benson 10787. OP: Peebles 10825. TA: Shreve 5940.

Justicia candicans (Nees) L. D. Benson [Jacobinia ovata A. Gray], Chuparrosa
OP: Nichol 23 Feb 1939.

Justicia longii Hilsenbeck [Siphonoglossa longiflora (Torrey) A. Gray] . Longflower Tube-tongue
OP: Daniel 2590 (ASU).

Ruellia nudiflora (Engelmann & A. Gray) Urban [/?. midiflora var. runyonii (Tharp & F. A.
Barkley) B. L. Turner], Violet Ruellia
OP: Lockwood 163.

ADOXACEAE - ADOXA FAMILY

**Sambucus cerulea Rafinesque [S. nigra Linnaeus subsp. cerulea (Rafinesque) Bolli. S. mexicana
C. Presl ex de Candolle. S. cerulea var. mexicana (C. Presl ex de Candolle) L. D. Benson. S.
neomexicana Wooton. S. cerulea var. neomexicana (Wooton) Rehder]. Blue Elderberry; tdpiro. See
Eriksson and Donoghue (1997).

OP: Tinkharn Apr 1942.

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AIZOACEAE - AIZOON FAMILY

*Mesembryanthemum erystallinum Linnaeus. Crystal Iceplant; hielitos
OP: SR 1 1 May 1995.

*Mesenibryanthemum nodiflorum Linnaeus. Slender-leaf Iceplant; hielitos
CP: SR 1 1 May 1995. OP: SR 1 1 May 1995.

Trianthema portulacastrum Linnaeus. Horse-purslane; verdolaga de cochi ; kasvan
CP: Adobe Well (Simmons 1966). OP: RSF 88-430.

AMARANTHACEAE- AMARANTH FAMILY (includes Chenopodiaceae)

* Amaranthus albus Linnaeus [A. graecizans of authors, not A. graecizans Linnaeus], Tumbleweed
Amaranth

OP: Harbison 13 Dec 1939 (SD).

Amaranthus crassipes Schlechtendahl var. crassipes. Spreading Amaranth
CP: RSF 92-713. OP: Bowers 1531 (ORPI). TA: RSF 04-24.

Amaranthus fimbriatus (Torrey) Bentham ex S. Watson. Fringed Amaranth; bledo, quelitillo;
cuhukkia i:vakl

CP: RSF 92-663. OP: Bowers 1520. TA: RSF 04-25.

Amaranthus palmeri S. Watson. Careless Weed, Pigweed; quelite de las aguas; cuhugia
CP: RSF 92-665. OP: Bowers 1521.

Amaranthus xtucsonensis Henrickson. Tucson Amaranth
OP: Harbison 26249.

Atriplex canescens (Pursh) Nuttall. Four-wing Saltbush; chamizo cenizo ; ’onk ’i:vaki
CP: RSF 93-61. OP: SR 20031031-1 (ORPI).

Atriplex elegans (Moquin-Tandon) D. Dietrich [A. elegans subsp . fasciculata (S. Watson) H. M.

Hall & Clements], Wheelscale Orach; chamizo cenizo; ’onk ’i:vakl
CP: RSF 92-581. OP: Bowers 912.

Atriplex lentiformis (Torrey) S. Watson subsp. lentiformis. Quail Bush, Lens Scale, Big Saltbush;

chamizo grande

OP: Nichol 28 Apr 1939.

Atriplex linearis S. Watson [A. canescens (Pursh) Nuttall subsp. linearis (S. Watson) H. M. Hall &
Clements]. Narrowleaf Saltbush. See Felger (2000) and Felger et al. (2007b).

CP: Las Playas, 31 Jan 1992, RSF, observation. OP: RSF 90-488.

Atriplex pacifica A. Nelson. Pacific Orach, South Coast Saltscale
CP: Eiber 7. OP: Bowers 1225.

Atriplex polyearpa (Torrey) S. Watson. Desert Saltbush, Allscale Saltbush; chamizo cenizo ; ’onk
’i:vakl

CP: TRY 9 Mar 1980. OP: RSF 87-268. TA: RSF 02-04.

**Bassia hyssopifolia (Pallas) Kuntze. Smother Weed
OP: SR 15 Apr 2005 (ORPI).

*Chenopodium murale Linnaeus. Netleaf Goosefoot; chual, choal ; ’onk ’i:vaki, kaupdam
CP: RSF 93-366. OP: Bowers 1015. TA: RSF 98-149.

Chenopodium watsonii A. Nelson. Watson’s Goosefoot
OP: Bowers 1019 (ORPI).

Com ph rena sonorae Torrey. Sonoran Globe Amaranth
OP: Wirt 13 Nov 1989 (ORPI).

Monolepis nuttalliana (Schultes) Greene. Poverty Weed; ’opon
CP: RSF 93-77. OP: Bowers 1160.

Nitrophila occidentalis (Moquin-Tandon) S. Watson. Alkali Weed
OP: Bowers 1333.

*Salsola tragus Linnaeus [S', australis R. Brown. S. iberica (Sennen & Pau) Botschantzev ex
Czerepanov. S’, kali of authors, not S. kali Linnaeus. S. kali var. tenuifolia Tausch. S. pestifer Nelson].
Russian Thistle, Tumbleweed; chamizo volador; hejel ’e’esadam, vo:po’odam sa’i
CP: RSF 92-644. OP: Bowers 1720.

Suaeda nigra (Rafinesque) J. F. Macbride [S’, moquinii (Torrey) Greene. S. torreyana S. Watson].
Bush Seepweed; quelite salado; s-cuk onk

CP: RSF 01-544. OP: TRY 30 Aug 1978.

2012

CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

9

Tidestromia lanuginosa (Nuttall) Standley subsp. eliassoniana Sanchez-del Pino & Flores Olvera
[T. eliassoniana (Sanchez-del Pino & Flores Olvera) Sanchez-del Pino]. Woolly Honeysweet; hierba
ceniza, hierba lanuda

CP: RSF 92-666. OP: RSF 88-408. TA: RSF 04-64.

ANACARDIACEAE - CASHEW FAMILY

Rhus aromatica Aiton var. trilobata (Nuttall) A. Gray [/?. trilobata Nuttall], Skunk Bush; limita
OP: Harbison 25670.

Rhus kearneyi F. A. Barkley subsp. kearneyi [ Schmaltzia kearneyi (F. A. Barkley) F. A. Barkley],
Desert Sumac; limita del desierto

CP: Simmons 27 Sep 1964. TA: Harrison & Kearney 6573 (isotype).

APIACEAE (UMBELLIFERAE) - CARROT OR PARSLEY FAMILY
Bowlesia incana Ruiz & Pavon. Hoary Bowlesia
CP: Elias 10202. OP: Bowers 1132.

Daucus pusillus Michaux. Wild Carrot; zanahoria silvestre
CP: RSF 93-343. OP: Nichol 26 Mar 1939.

Eryngium nasturtiifolium de Jussieu ex F. Delaroche. Hierba del sapo
CP. Lehto L22460.

Lomatium nevadense (S. Watson) J. M. Coulter & Rose. Biscuit-root
OP: SR 2003-333 (ORPI).

Spermolepis echinata (Nuttall ex de Candolle) A. Heller. Bristly Scale-seed

CP: RSF 93-360. OP: McDougall 69.

Yabea microcarpa (Hooker & Arnott) Koso-Poljanski [Caucalis microcarpa Hooker & Arnott],
False Carrot

OP. Wirt 1 Apr 1990.

ARISTOLOCHIACEAE, see MAGNOLIIDS

APOCYNACEAE - DOGBANE FAMILY (includes Asclepiadaceae )

Asclepias albicans S. Watson. White-stem Milkweed; jumate, candelilla
CP: Engard 912 (ASU). TA: Shreve 5941.

Asclepias erosa Torrey. Giant Sand-milkweed, Desert Milkweed; hierba del cuervo
CP: RSF 92-622. TA: Simmons 21 May 1965.

Asclepias linaria Cavanilles. Narrow-leaf Milkweed
OP: Darrow 3856.

Asclepias nyctaginifolia A. Gray. Mojave Milkweed
OP: Wirt 10 Aug 1990.

Asclepias subulata Decaisne. Desert Reed-stem Milkweed, Rush Milkweed \jumete, mata candelilla
CP: Harlan 447. OP: Bowers 1538 (ORPI). TA: RSF 04-03.

Funastrum hartwegii (Vail) Schlechter [F. cynanchoides (Decaisne) Schlechter var. hartwegii
(Vail) Krings. Sarcostemma cynanchoides Decaisne subsp. hartwegii (Vail) R. W. Holm. Funastrum
heterophyllum (Engelmann ex Torrey) Standley]. Climbing Milkweed; giiirote

CP: Duncan 28 Mar 1970. (CAB). OP: Parker 7982. TA: TRV9-10 Mar 1980.
Haplophyton cimicidum A. de Candolle [H. cimicidum var. crooksii L. D. Benson. H. crooksii (L.
D. Benson) L. D. Benson], Cockroach Plant; hierba de la cucaracha
OP: Supernaugh 432.

Matelea cordifolia (A. Gray) Woodson. Heart-leaf Milkweed Vine
OP: Supernaugh 440 (ORPI).

Matelea parvifolia (Torrey) Woodson. Spearleaf
CP: RSF 94-29. OP: Mulroy 2 May 1970.

Metastelma arizonicum A. Gray [Cynanchum arizonicum (A. Gray) Shinners]

OP: Darrow 2436.

ASCLEPIADACEAE, see APOCYNACEAE

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CANOTIA Vol. 8

2012

ASTERACEAE (COMPOSITAE) - ASTER OR DAISY FAMILY

Acamptopappus sphaerocephalus (Harvey & A. Gray) A. Gray var. sphaerocephalus. Goldenhead
OP: Tinkham 18 Apr 1942.

Acourtia nana (A. Gray) Reveal & R.M. King [Perez i a nana A. Gray]. Desert Holly
OP: SR 20061007-1.

Acourtia wrightii (A. Gray) Reveal & R. M. King [Perez ia wrightii A. Gray], Brown-foot
CP: RSF 93-306. OP: Bowers 1702.

Adenophyllum porophylloides (A. Gray) Strother [Dvssodia porophylloides A. Gray], Dogweed
CP: RSF 92-735. OP: TRY 85-109.

Ageratina paupercula (A. Gray) R. M. King & H. Robinson [Eupatorium pauperculum A. Gray],
Santa Rita Snakeroot

OP: Darrow & Gould 4686.

Ambrosia ambrosioides (Cavanilles) W. W. Payne [Franseria ambrosioides Cavanilles]. Canyon
Ragweed; chicura\ nunui jej

CP: Vorhies 11 Apr 1924. OP: Mearns 2736 (US). TA: TRV 9-10 Mar 1980.

Ambrosia confertiflora de Candolle. Slim-leaf Bur-sage; estafiate\ mo'ostalk
CP: RSF 92-538. OP: Bowers 915.

Ambrosia cordifolia (A. Gray) W. W. Payne [Franseria cordifolia A. Gray]. Heart-leaf Bur-sage
OP: Shreve 6198.

Ambrosia deltoidea (Torrey) W. W. Payne [Franseria deltoidea Torrey]. Triangle-leaf Bur-sage;
chamizo forrajero; tadsad, va:gita

CP: RSF 92-680. OPT RSF 90-401. TA: TRV 26 Mar 1983.

Ambrosia dumosa (A. Gray) W. W. Payne [Franseria dumosa A. Gray], White Bur-sage; chamizo ;
tadsad

CP: Shreve 5918. OP: Bowers 991. TA: RSF 04-07.

Ambrosia dumosa x A. ilicifolia (putative hybrid)

TA: RSF 98-136.

Ambrosia ilicifolia (A. Gray) W. W. Payne [Franseria ilicifolia A. Gray], Holly-leaf Bur-sage
CP: Goodding 30 Nov 1938. TA: Vorhies 16 Apr 1924.

Ambrosia monogvra (Torrey & A. Gray) Strother & B. G. Baldwin [Hymenoclea monogvra Torrey
& A. Gray], Slender Burrobrush; jecota; 'kvadhod
CP: RSF 87-261. OP: RSF 87-261.

Ambrosia salsola (Torrey & A. Gray) Strother & B. G. Baldwin var. pentalepis (Rydberg) Strother
& B. G. Baldwin [Hymenoclea salsola Torrey & A. Gray wax. pentalepis (Rydberg) L. D. Benson].
White Burrobrush, Cheesebush; 'i:wadhod

CP: RSF 93-166. OP: Mearns 2768 (US). TA: Webster 24245.

Arida arizonica (R. C. Jackson & R. R. Johnson) D. R. Morgan & R. L. Hartman [ Machaeranthera
arizonica R. C. Jackson & R. R. Johnson. M. arida B. L. Turner & D. B. Home. M. coulteri (A.

Gray) B. L. Turner & D. B. Home var. arida (B. L. Turner & D. B. Home) B. L. Turner. M.
ammophila Reveal. Psilactis coulteri A. Gray], Arid Tansy-aster

CP: RSF 93-435. OP: Jackson 3043-2 (isotype of M. arizonica).

Arida carnosa (A. Gray) D. R. Morgan & R. L. Hartman [Aster carnosus (A. Gray) A. Gray ex
Hemsley, not A. carnosus Gilbert. A. intricatus (A. Gray) S. F. Blake. Leucosyris carnosa (A. Gray)
Greene. Machaeranthera carnosa (A. Gray) G. L. Nesom var. intricata (A. Gray) G. L. Nesom],
Alkali Aster

OP: Stimson 233.

Artemisia dracunculus Linnaeus [A. dracunculoides Pursh], Tarragon; estragon
OP: Goodding 484-45.

Artemisia iudoviciana Nuttall subsp. albula (Wooton) D. D. Keck. Western Mugweed, White Sage,
Silver Wormwood; estafiate
OP: Harbison 26269.

Baccharis brachyphylla A. Gray. Short-leaf Baccharis

CP: RSF 92-533. OP: Warren 10 Nov 1983. TA: Goodding 1186.

Baccharis salicifolia (Ruiz & Pa von) Persoon subsp. salicifolia [B. glutinosa Persoon]. Seep
Willow; batamote; susk

CP: RSF 92-576. OP: Nichol 28 Apr 1939.

2012

CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

Baccharis sarothroides A. Gray. Desert Broom; escoba amarga, romerillo\ susk kuagig
CP: RSF 92-706. OP: Mearns 2775 (US). TA: RSF 04-26. 04-27.

Bahiopsis parishii (Greene) E. E. Schilling & Panero [Viguiera parishii Greene. V. deltoideci A.
Gray var. parishii (Greene) Vasey & Rose]. Parish’s Goldeneye; arioso
CP: RSF 95-295. OP: Harbison 26168. TA: RSF 04-84.

Baileya multiradiata Harvey & A. Gray. Many-flowered Desert-marigold; gi:ko
CP: RSF 92-747. OP: Nichol 17 Apr 1939.

Baileya pleniradiata Harvey & A. Gray. Woolly Desert-marigold
CP: Benson 10782. OP: Bowers 1171. TA: RSF 05-48.

Bebbia juncea (Bentham) Greene var. aspera Greene. Sweetbush, Chuckwalla Delight; hauk ’u’us
CP: Simmons 2 Oct 1963. OP: Benson 9907. TA: TRY 5 Mar 1983.

Brickellia atractyloides A. Gray var. atractyloides. Spiny-leaf Brickellbush
CP: RSF 92-583. TA: Vorhies 16 Apr 1924.

Brickellia californica (Torrey & A. Gray) A. Gray. California Brickellbush
OP: Bowers 929 (ORPI).

Brickellia coulteri A. Gray var. coulteri. Triangle-leaf Brickellbush; pachaba
CP: RSF 93-271. OP: Harbison 26143.

Brickellia frutescens A. Gray. Shrubby Brickellbush
OP: Darrow 3841 .

Calycoseris wrightii A. Gray. White Tack-stem

CP: RSF 93-340. OP: Parker 7926. TA: RSF 05-145.

**Carthamus tinctorius Linnaeus. Safflower; cdrtamo
OP: Bowers 1717 (ORPI).

*Centaurea melitensis Linnaeus. Malta Star Thistle
CP: RSF 92-716. OP: RSF 89-251.

Chaenactis carphoclinia A. Gray var. carphoclinia. Pebble Pincushion
CP: RSF 93-403. OP: RSF 03-352. TA: TRY 26 Mar 1983.

Chaenactis stevioides Hooker & Amott. Desert Pincushion

CP: RSF 93-369. OP: Nichol 14 Mar 1939. TA: Lindquist 26 Mar 1983.

Cirsium neomexicanum A Gray. New Mexico Thistle; cardo\ gewel
OP: Gould 31 Mar 1948.

Diaperia verna (Rafmesque) Morefield var. verna [Evax multicaulis de Candolle. E. verna
Rafinesque var. verna]. Spring Pygmy Cudweed, Rabbit Tobacco
CP: Harlan 167. OP: SR 2003-391.

Dieteria asteroides Torrey var. glandulosa (B. L. Turner) D. R. Morgan & R. L. Hartman
[Machaeranthera asteroides (Torrey) Greene var. glandulosa B. L. Turner], Fall Tansy-aster
OP: Goodding 485-45.

Eclipta prostrata (Linnaeus) Linnaeus [E. alba (Linnaeus) Hasskarl. E. erecta Linnaeus]. False
Daisy; chile de agua, hierba del tajo
OP: Niles 724.

Encelia farinosa A. Gray ex Torrey. Brittlebush; incienso , hierba del bazo, rama blanca\ tohaves
CP: RSF 92-54. OP: McDougall 55. TA: Goodding 2204.

Encelia farinosa x E. frutescens

OP: Bowers 1131. TA: RSF 01-585.

Encelia frutescens (A. Gray) A. Gray. Button Encelia

CP: Lehto L23586 (ASU). OP: Bowers 1130. TA: RSF 04-65.

Ericameria cuneata (A. Gray) McClatchie var. spathulata (A. Gray) H. M. Hall [Haplopappus
cuneatus A. Gray var. spathulata (A. Gray) S. F. Blake], Wedgeleaf Goldenbush
OP: RSF 90-513.

Ericameria laricifolia (A. Gray) Shinners [Haplopappus laricifolius A. Gray], Turpentine Bush
OP: Bowers 969.

*Erigeron canadensis Linnaeus [Convza canadensis (Linnaeus) Cronquist]. Horseweed; cola de
caballo

CP: RSF 92-568. OP: RSF 87-322.

Erigeron lobatus A. Nelson. Desert Fleabane

CP: RSF 93-325. OP: RSF 86-182. TA: RSF 98-118.

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CAN0T1A Vol. 8

2012

Eriophvllum lanosum (A. Gray) A. Gray 1 Antheropeas lanosum (A. Gray) Rydberg], Woolly Daisy
' CP: Whipple 3944. OP: Harlan 03-6.

Gaillardia arizonica A. Gray [ G . arizonica var. pringlei (Rydberg) S. F. Blake], Arizona Blanket-
flower

CP: Malusa 18 Apr 2001. OP: McDougall 63.

Gamochaeta stagnalis (I. M. Johnston) Anderberg [ Gnaphalium stagnale I. M. Johnston], Desert
Cudweed, Rosy Everlasting
OP: SR 20050318-9.

Geraea canescens Torrey & A. Gray. Desert Sunflower, Desert Gold
CP: Shreve 5927. OP: McDougall 8. TA: TRV27 Mar 1983.

Gutierrezia arizonica (A. Gray) M. A. Lane [ Greenella arizonica A. Gray], Arizona Snakeweed
CP: RSF 93-367. OP: Bowers 1154.

Gutierrezia sarothrae (Pursh) Britton & Rusby. Broom Snakeweed, Ruby Matchweed; hierba de la
vibora ; siw tadsagi

CP: RSF 92-584. OP: Bowers 935.

Gymnosperma glutinosum (Sprengel) Lessing [Selloa glutinosa Sprengel], Gumhead
CP: Benson 10802. OP: Bowers 988. TA: McLaughlin 1969.

**Helianthus annuus Linnaeus. Sunflower; mirasol
OP: SR 21 Jul 1998 (ORP1).

Helianthus niveus (Bentham) Brandegee subsp. tephrodes (A. Gray) Heiser. Dune Sunflower;
mirasol de las dunas ; hi:wai
CP: RSF 92-775.

Hymenothrix wislizeni A. Gray. Thimblehead
CP: RSF 92-536. OP: Harbison 26196.

Hy menoxys odorata de Candolle. Bitterweed
CP: RSF 93-382.

Isocoma acradenia (Greene) Greene var. acradenia [ Haplopappus acradenius (Greene) S. F.
Blake], Alkali Goldenbush

CP: Tule Well (Simmons 1965, 1966). OP: Simmons 11 Oct 1963.

Koanophvllon palmeri (A. Gray) R. M. King & H. Robinson [Eupatorium palmeri A. Gray (a
segregate off', solidaginifolium A. Gray)]. Umbrella Thoroughwort
CP: Tule Well (Simmons 1965, 1966). OP: RSF 90-528.

*Lactuca serriola Linnaeus. Prickly Lettuce, Compass Plant
CP: RSF 93-375. OP: Bowers 1756. TA: RSF 10-162.

Laennecia coulteri (A. Gray) G. L. Nesom [Conyza coulteri A. Gray]. Coulter’s Horseweed
CP: Harlan 269 (CAB). OP: RSF 87-291. TA: RSF 02-14.

Logfia arizonica (A. Gray) Holub [Filago arizonica A. Gray], Arizona Fluffweed
CP: RSF 92-561. OP: RSF 88-274. TA: RSF 98-114.

Logfia depressa (A. Gray) Holub [Filago depressa A. Gray], Dwarf Fluffweed

CP: RSF 93-400. OP: Lehto 6-15440-b (ASU, mixed sheet with Sp’locline gnaphaloides) .
Logfia filaginoides (Hooker & Amott) Morefield [L. californica (Nuttall) Holub. Filago californica
Nuttall], California Fluffweed

CP: RSF 92-573. OP: Bowers 1248. TA: RSF 98-144.

Machaeranthera tagetina Greene [Aster tagetinus (Greene) S. F. Blake]. Mesa Tansyaster
OP: RSF 05-219.

Malacothrix fendleri A. Gray. Fendler’s Desert-dandelion
CP: Cuttler 17 Mar 1995 (CAB). OP: RSF 03-259.

Malacothrix glabrata (A. Gray ex D. C. Eaton) A. Gray [M. californica de Candolle var. glabrata
A. Gray ex D. C. Eaton]. Desert Dandelion

CP: McLaughlin 2983. OP: SR 2003-399.

Malacothrix sonorae W. S. Davis & P. H. Raven. Sonoran Desert Dandelion
CP: Harlan 220. OP: Niles 554.

Monoptilon bellioides (A. Gray) H. M. Hall. Mojave Desert Star

CP: SR & Tibbitts 18 Feb 2002. OP: Benson 10625. TA: Halse 31 Mar 1973.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

13

*Oncosiphon piluliferum (Linnaeus f.) Kallersjo [Matricaria glob if era (Thunberg) Frezi ex
Harvey], Pineapple- weed. False Chamomile; manzanilla
OP: SR 15 Apr 2005. CP: Harlan 165.

Packera quereetorum (Greene) C. Jeffrey [Senecio quercetorum Greene], Oak Creek Ragwort
OP: Bowers 1284.

Palafoxia arida B. L. Turner & M. I. Morris var. arida. Spanish Needles
CP: Benson 10787. OP: RSF 88-406. TA: TRY 27 Mar 1983.

Parthenice mollis A. Gray. Annual Monsterwort
OP: SR 20031025-13.

Pectis cyiindrica (Fernald) Rydberg. Sonoran Chinchweed
CP: RSF 01-566.

Pectis linifolia Linnaeus var. linifolia [P. linifolia var. marginal is Fernald. P. punctata Jacquin],
Romero macho

OP: Wirt 13 Aug 1990.

Pectis papposa Harvey & A. Gray var. papposa. Desert Chinchweed; manzanilla del coyote ; ban
manzanuya

CP: RSF 92-557. OP: Bowers 1535. TA: RSF 04-56.

Perityle ajoensis Todsen. Ajo Rock Daisy

OP: Todsen 2292 (isotypes ARIZ, ORPI).

Perityle ernoryi Torrey. Desert Rock Daisy

CP: Whipple 3912. OP: McDougall 50. TA: RSF 04-57.

Peucephyllum schottii A. Gray. Pygmy-cedar; romero del desierto
CP: Hodgson 2752 (DES). TA: TRY 5 Mar 1983.

Pleurocoronis laphamioides (Rose) R. M. King & H. Robinson [Hofmeisteria laphamioides Rose]
OP .Gould 3019.

Pleurocoronis pluriseta (A, Gray) R.M. King & H. Robinson [ Hofmeisteria pluriseta A. Gray].
Arrow-leaf; canutillo

CP: Benson 10804. TA: Goodding 2103.

Pluchea odorata (Linnaeus) Cassini var. odorata [P. purpurascens (Swartz) de Candolle],
Saltmarsh-fleabane, Alkali Camphor-weed; jara
OP -.RSF 86-204.

Pluchea sericea (Nuttall) Coville [Tessaria sericea (Nuttall) Shinners], Arrow-weed; cachanilla;
kornagl 'u'us, 'u'us kokomadk
OP: Bowers 1315.

Porophyllum gracile Bentham. Slender Poreleaf; odora, hierba del venado
CP: SR 2003-19. OP: RSF 88-118 (ORPI). TA: TRY 5 Mar 1983.

Prenanthella exigua (A. Gray) Rydberg [Lygodesmia exigua (A. Gray) A. Gray], Brightwhite
CP: Bowers 2996. OP: Gould 2990. TA: RSF 98-132.

Psathyrotes ramosissima (Torrey) A. Gray. Desert Velvet, Turtleback
CP: Yeatts 3246 (CAB). TA: Halse 31 Mar 1973.

Pseudognaphalium canescens (de Candolle) Anderberg [Gnaphalium canescens de Candolle. G.
wrightii A. Gray], Wright’s Cudweed
OP: Bowers 1927.

Psilostrophe cooperi (A. Gray) Greene. Paper-daisy, Paper- flower
CP: RSF 93-302. OP: McDougal 22 Mar 1941.

Rafinesquia californica Nuttall. California Chicory'

CP: RSF 93-299. OP: McDougall 29.

Rafinesquia neomexicana A. Gray. Desert Chicory

CP: Harbison 26156. OP: Shreve 6206. TA: McLaughlin 1974.

Senecio flaccidus Lessing var. monoensis (Greene) B. L. Turner & T. M. Barkley [S’, monoensis
Greene], Sandwash Groundsel, Threadleaf Ragwort
OP: Bowers 1600.

Senecio lemmonii A. Gray. Lemmon Groundsel

CP: Simmons 24 Jan 1965. OP: Kearney 10849.

Senecio mohavensis A. Gray. Mojave Groundsel, Mohave Ragwort
CP: RSF 93-91. OP: McDougall 76. TA: TRY 26 Mar 1983.

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CANOTIA Vol. 8

2012

*Sonchus asper (Linnaeus) Hill subsp. asper. Prickly Sow Thistle; chinita\ ho’idkam, ’i:vakl
CP: RSF 93-98. OP: Nichol 4 May 1939. TA: RSF 02-10.

*Sonchus oleraceus Linnaeus. Common Sow Thistle; chinita\ hauvi, hehewo

OP: Bowers 1594. TA: Frontera Canyon, 18 Mar 1998, RSF, observation.

Stephanomeria exigua Nuttall subsp. exigua. Wire-lettuce
OP: SR 2003-457.

Stephanomeria pauciflora (Torrey) A. Nelson. Desert-straw, Desert Wire-lettuce
CP: Cain 15 Nov 2003. OP: RSF 86-293. TA: TRV 86-144.

Stephanomeria schottii (A. Gray) A. Gray. Schott’s Wire-lettuce
CP: Reeves 6780.

Stylocline gnaphaloides Nuttall [5. arizonica Coville], Everlasting Neststraw
OP: Lehto 6-15440-b (ASU, mixed sheet with Logfia depressa).

Stylocline micropoides A. Gray. Desert Neststraw

CP: Whipple 3926. OP: Niles 553. TA: RSF 98-143.

Thymophylla concinna (A. Gray) Strother [Dyssodia concinna (A. Gray) B. L. Robinson],

Dogweed; manzanilla de coyote\ ban mansani:ya
CP: Niles 341. OP: RSF 88-119.

Thymophylla pentachaeta (de Candolle) Small var. belenidium (de Candolle) Strother [Dyssodia

pentachaeta (de Candolle) B. L. Robinson var. belenidium (de Candolle) Strother], Five-needle

Prickly-leaf

CP: RSF 92-636A. OP: Beal 10 Mar 1987 (ORPI).

Townsendia annua Beaman. Annual Townsend Daisy
OP: SR 2003-287.

Trichoptilium incisum (A. Gray) A. Gray. Yellow-head

CP: RSF 92-776. OP: SR 2003-316 (ORPI). TA: TRV 10 Mar 1986.

Trixis californica Kellogg var. californica. California Threefold

CP: Reichenbacher 470. OP: Lockwood 162. TA: Vorhies 16 Apr 1924.

Uropappus lindleyi (de Candolle) Nuttall [Microseris linearifolia (de Candolle) Schultz Bipontinus.

M. lindleyi (de Candolle) A. Gray], Silverpuffs
CP: RSF 93-352. OP: McDougall 98.

**Verbesina encelioides (Cavanilles) Bentham & Hooker f. ex A. Gray subsp. exauriculata (B. L.

Robinson & Greenman) J. R. Coleman. Golden Crownbeard
OP: Harbison 26271 (SD).

Xanthisma gracile (Nuttall) D. R. Morgan & R. L. Hartman [Haplopappus gracilis (Nuttall) A.

Gray. Machaeranthera gracilis (Nuttall) Shinners]. Slender Goldenweed
OP: Bowers 1698.

Xanthisma spinulosum (Pursh) D. R. Morgan & R. L. Hartman var. gooddingii (A. Nelson) D. R.

Morgan & R. L. Hartman [ Haplopappus spinulosus (Pursh) de Candolle subsp. gooddingii (A.

Nelson) H. M. Hall. Machaeranthera pinnatifida (Hooker) Shinners var. gooddingii (A. Nelson) B.

L. Turner & Hartman]. Spiny Goldenweed

CP: Whipple 3932. OP: Cooper 554. TA: Kurtz 1169.

Zinnia acerosa (de Candolle) A. Gray. Desert Zinnia; zinia del desierto
OP: Bowers 1692.

BERBERIDACEAE - BARBERRY FAMILY

Berberis haematocarpa Wooton [Mahonia haematocarpa (Wooton) Fedde]. Red Barberry
OP: Niles 536.

Berberis harrisoniana Kearney & Peebles. Kofa Mountain Barberry
OP: Phillips 76-2.

BIGNONIACEAE - BIGNONIA FAMILY

Chilopsis linearis (Cavanilles) Sweet subsp. arcuata (Fosberg) Henrickson. Desert-willow; mimbre
CP: RSF 92-547. OP: SR 20061008-13.

2012

CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

15

BORAGINACEAE - BORAGE FAMILY (includes Hydrophyllaceae)

Amsinckia intermedia Fischer & C. A. Meyer. Devil’s Lettuce, Fiddleneck; cetkom
CP: RSF 93-58. OP: RSF 88-305. TA: TRV 5 Mar 1983.

Amsinckia tessellata A. Gray var. tessellata. Desert Fiddleneck; cetkom
CP: RSF 93-181. OP: Cooper 727. TA: RSF 10-161.

Cryptantha angustifolia (Torrey) Greene. Narrow-leaf Cryptantha, Desert Cryptantha
CP: Shreve 6219. OP: Bowers 1593. TA: TRV 5 Mar 1983.

Cryptantha barbigera (A. Gray) Greene. Bearded Cryptantha
CP: Kearney 1088. OP: RSF 87-260. TA: RSF 98-137.

Cryptantha ganderi I. M. Johnston. Dune Cryptantha
CP: RSF 93-388.

Cryptantha holoptera (A. Gray) J. F. Macbride. Winged Cryptantha
CP: Benson 10791 .

Cryptantha maritima (Greene) Greene [includes var. maritima and var . pilosa I. M. Johnston].
White-haired Cryptantha

CP: Benson 10758. OP: Gould 3005. TA: Kearney 10905.

Cryptantha micrantha (Torrey) I. M. Johnston subsp. micrantha. Dwarf Cryptantha

CP: Shreve 6217. TA: Butterwick 6265.

Cryptantha pterocarya (Torrey) Greene var. cycloptera (Greene) J. F. Macbride. Wing-nut
Cryptantha

CP: Kearney 10893. OP: McDougall 19. TA: TRV 10 Mar 1980.

Cryptantha racemosa (S. Watson ex A. Gray) Greene. Bushy Cryptantha
TA: TRV 5 Mar 1983.

Eucrypta chrysanthemifolia (Bentham) Greene var. bipinnatifida (Torrey) Constance. Spotted
Hideseed

CP: Daniel 2656 (ASU). OP: Benson 10661.

Eucrypta micrantha (Torrey) A. Heller. Desert Hideseed; peluda

CP: SR 2003-40. OP: McDougall 43. TA: TRV 5 Mar 1983.

Harpagoneila palmeri A. Gray. Palmer’s Grappling-hook
OP: RSF 05-218.

Heliotropium curassavicum Linnaeus [H curassavicum var. oculatum (A. Heller) I. M. Johnston ex
Tidestrom]. Alkali Heliotrope; hierba del sapo; babad 'i:vakl, kakaicu 'i:vaki
CP: RSF 92-577. OP: McDougall 90.

Lappula redowskii (Hornemann) Greene var. cupulata (A. Gray) M. E. Jones [L. occidentalis (S.
Watson) Greene]. Crowned Stickseed

CP: Kearney 11029. OP: Parker 7990.

Narna demissum A. Gray var. demissum. Purple Mat
CP; RSF 93-442. TA: RSF 98-112.

Nama hispidum A. Gray var. spathulatum (Torrey) C. L. Hitchcock [N. coulter i A. Gray]. Bristly
Nama

CP: SR 18 Feb 2002. OP: Fay 744. TA: RSF 04-52.

Pectocarya heterocarpa (I. M. Johnston) I. M. Johnston. Mixed-nut Comb-bur
CP: Parker 7936. OP: Parker 7936. TA: TRV 5 Mar 1983.

Pectocarya platycarpa (Munz & I. M. Johnston) Munz & I. M. Johnston. Broad-winged Comb-bur
CP: Whipple 3949. OP: Benson 10664. TA: TRV 5 Mar 1983.

Pectocarya recurvata I. M. Johnston. Arched Comb-bur
CP: RSF 93-362. OP: Parker 7936 A.

Phacelia a f finis A. Gray. Limestone Phacelia
OP: Bowers 1069 (ORPI).

Phacelia coerulea Greene. Sky-blue Phacelia

CP: Lehto L 23535 (ASU). OP: Kearney 10824.

Phacelia crenulata Torrey ex S. Watson var. ambigua (M. E. Jones) J. F. Macbride [R. ambigua M.
E. Jones. P. minutiflora J. W. Voss ex Munz], Desert Heliotrope

CP: Reeves 6766 (ASU). OP: Bowers 1028. TA: TRV 5 Mar 1983.

Phacelia distans Bentham. Caterpillar Phacelia, Femleaf Phacelia
CP: McManus 661. OP: Kearney 10853.

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Phacelia neglecta M. E. Jones. Alkali Phacelia

CP: Lehto L23540 (ASU). OP: TRV85-S. TA: RSF 05-62.

Phacelia pedicellata A. Gray. Pedicellate Phacelia

CP: Irwin 93 (ASU). OP: Niles 552. TA: RSF 98-145.

Phacelia ramosissima Douglas ex Lehmann [P. ramosissima var. latifolia (Torrey) Cronquist].
Branching Phacelia

OP: Harbison 26261 .

Phacelia rotundifolia Torrey ex S. Watson. Roundleaf Phacelia
OP: Christy 1263 (ASU).

Pholistoma auritum (Lindley) Lilja ex Lindblom var. arizonicum (M. E. Jones) Constance. Blue
Fiesta-flower

CP: RSF 93-62. OP: Kearney 10817.

Plagiobothrys arizonicus (A. Gray) Greene ex A. Gray. Arizona Popcorn-flower
OP: RSF 03-271.

Plagiobothrys jonesii A. Gray. Mojave Popcorn-flower
OP: TRY 19 Feb 1984.

Tiquilia canescens (A. de Candolle) A. T. Richardson var. puichella (I. M. Johnston) A. T.
Richardson. Woody Crinklemat

CP: TRV 9 Mar 1980. OP: SR 2003-211.

Tiquilia palmeri (A. Gray) A.T. Richardson. Palmer’s Crinklemat
CP: RSF 93-410. TA: TRV 27 Mar 1983.

BRASSICACEAE (CRUCIFERAE) - MUSTARD FAMILY

Boechera perennans (S. Watson) W. A. Weber [Arabis perennans S. Watson], Perennial Rockcress
OP: Nichol 14 Mar 1939.

**Brassica nigra (Linnaeus) W. D. J. Koch. Black Mustard; mostaza negra
OP: SR 2003-449.

*Brassica tournefortii Gouan. Sahara Mustard; mostaza

CP: RSF 93-418. OP: Bowers 1030. TA: RSF 04-33.

**Capsella bursa-pastoris (Linnaeus) Medikus. Shepherd’s Purse; hierba del pastor
CP: RSF 93-201.

Caulanthus lasiophyllus (Hooker & Arnott) Payson [Guillenia lasiophylla (Hooker & Arnott)
Greene. Thelypodium lasiophyllum (Hooker & Arnott) Greene], California Mustard
CP: Reeves 6822 (ASU). OP: RSF 88-133. TA: Reeves 5404 (ASU).

Descurainia pinnata (Walter) Britton subsp. ochroleuca (Wooton) Detling. Western Tansy
Mustard; pamita\ su'uvad

CP: RSF 93-277. OP: Nichol 14 Mar 1939. TA: RSF 05-152.

Dimorphocarpa pinnatifida Rollins. Dune Spectacle-pod
CP: RSF 92-27. TA: RSF 98-138.

Dithyrea californica Harvey. California Spectacle-pod

CP: RSF 92-28. T A: TRV 27 Mar 1 983.

Draba cuneifolia Nuttall ex Torrey & A. Gray \D cuneifolia var. integrifolia S. Watson. D.
cuneifolia var. sonorae (Greene) Parish], Wedgeleaf Draba
CP: RSF 93-62. OP: Bowers 1006. TA: RSF 05-36.

*Eruca vesicaria (Linnaeus) Cavanilles subsp. sativa (Miller) Thellung [ E . sativa Miller], Garden-
rocket, Salad-rocket, Arugala

OP: Conner 16 Feb 2005.

Erysimum capitatum (Douglas ex Hooker) Greene [perhaps E. capitatum var. purshii (Durand)
Rollins]. Western Wallflower

OP: Tinkham 15 Apr 1942.

Hesperidanthus linearifolius (A. Gray) Rydberg [ Schoenocrambe linearifolia (A. Gray) Rollins.
Sisymbrium linearifolium (A. Gray) Payson. Thelypodiopsis linearifolia (A. Gray) Al-Shehbaz],
Purple Rock-mustard

OP: Bowers 1090.

Lepidium densiflorum Schrader. Common Peppercress, Coommon Pepper-weed
OP: Bowers 1063.

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17

Lepidium lasiocarpum Nuttall subsp. lasiocarpum. Sand Pepper-weed; lentejilla
CP: RSF 93-426 . OP: Shreve 6203. TA: TRY 10 Mar 1980.

Lepidium thurberi Wooton. Arizona Pepper-weed
OP: McDougall 32.

Lyrocarpa coulteri Hooker & Harvey. Lyre-pod; ban censanig

CP: Goodding 1554. OP: Parker 7914. TA: Harrison 6563.

**Matthiola longipetala (Ventenat) de Candolle [M. bicornis (Sibthorp & Smith) de Candolle],
Night-scented Stock

OP: SR 27 Apr 2005.

**Nasturtium officinale W. T. Aiton [ Rorippa nasturtium-aquaticum (Linnaeus) Hayek],

Watercress; berro

OP: Williams Spring (extirpated, Felger et al. 1992).

Physaria tenella (A. Nelson) O’Kane & Al-Shehbaz [Lesquerella tenella A. Nelson]. Desert
Bladderpod

CP: RSF 92-60. OP: McDougall 15. TA: TRY 5 Mar 1983.

Sibara angelorum (S. Watson) Greene. Angled Rockcress
CP: Lehto L23523 (ASU).

Sibara virginica (Linnaeus) Rollins. Winged Rockcress
CP: Harlan 495.

**Sisymbrium altissimum Linnaeus. Tumble Mustard, Jim Hill Mustard
OP: SR 15 Mar 2005 (ORPI).

♦Sisymbrium irio Linnaeus. London Rocket; pamiton; ban cinsanig
CP: RSF 92-64. OP: Kearney 10830.

♦Sisymbrium orientale Linnaeus. Indian hedge-mustard
CP: RSF 05-158. OP: SR 20050315-1.

Streptantheila longirostris (S. Watson) Rydberg. Long-beak Twist-flower
CP: RSF 92-20. OP: Baker 7701 (ORPI).

Streptanthus carinatus C. Wright ex A. Gray subsp. arizonicus (S. Watson) Kruckenberg, Rodman,
& Worthington [5. arizonicus var. luteus Kearney & Peebles], Arizona Jewel Flower
OP: Nichol 26 Mar 1939.

Thelypodium wrightii A. Gray \ Stanleyella wrightii (A. Gray) Rydberg]

OP: Hesselberg 16 Oct 1966.

**Thlaspi arvense Linnaeus. Fanweed, Roadside Pennycress
OP: Pinkava 9993 (ASU).

Thysanocarpus curvipes Hooker [T. curvipes var. elegans (Fischer & C. A. Meyer) B. L.

Robinson]. Lacepod

CP: RSF 93-93. OP: Kearney 10816.

BURSERACEAE - TORCHWOOD FAMILY

Bursera microphylla A. Gray. Elephant Tree; torote; ‘usabkam

CP: Elias 10308. OP: Bowers 1095. TA: Vorhies 16 Apr 1924.

CACTACEAE - CACTUS FAMILY

Carnegiea gigantea (Engelmann) Britton & Rose [Cereus giganteus Engelmann], Saguaro, Sahuaro;
saguaro ; ha:san

CP: Rebman 2884 (DES). OP: RSF 90-70. TA: Tinajas Altas, 19 Mar 1998, RSF,
observation.

Cylindropuntia acanthocarpa (Engelmann & J. M. Bigelow) F. M. Knuth [Opuntia acanthocarpa
Engelmann & J. M. Bigelow], Buckhorn Cholla; choya; ciolim (also hanami, the general term for
cholla)

Cylindropuntia acanthocarpa var. coloradensis (L. D. Benson) Pinkava [Opuntia acanthocarpa
var. coloradensis L. D. Benson]

CP: Rebman 2883 (ASU). OP: Baker 7578 (ASU). TA: Reeves R5395 (ASU).

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Cylindropuntia acanthocarpa var. major (Engelmann & J. M. Bigelow) Pinkava [Opuntia
echinocarpa Engelmann var. major Engelmann. O. acanthocarpa var. major (Engelmann & J. M.
Bigelow) L. D. Benson]

CP: 7 Harbison 17004 & 17005 (SD). OP: Steenbergh 5-2662-1 (neotype, RSA/POM).

Cylindropuntia acanthocarpa x C. spinosior

OP: Baker 8366 (ASU, SD).

Cylindropuntia arbuscula (Engelmann) F. M. Knuth [ Opuntia arbuscula Engelmann]. Pencil
Cholla; siviri, tasajo', vipinoi

OP: Baker 7573 (ASU).

Cylindropuntia bigelovii (Engelmann) F. M. Knuth [ Opuntia bigelovii Engelmann], Teddybear
Cholla; choya guera\ hadsadkam (also hanamT, the general term for cholla)

CP: Benson 10766. OP: Baker 7588 (ASU, ORPI). TA: Mearns 348 (US).

Cylindropuntia echinocarpa (Engelmann & J. M. Bigelow) F. M. Knuth [ Opuntia echinocarpa
Engelmann & J. M. Bigelow. O. wigginsii L. D. Benson], Silver Cholla

CP: Rebman 2881 (SD). TA: SE ofTinajas Altas Mts, 3 Feb 1990, RSF , observation.
Cylindropuntia fulgida (Engelmann) F. M. Knuth [ Opuntia fulgida Engelmann], Chain-fruit Cholla;
choya\ hanaml

Cylindropuntia fulgida var. fulgida [Opuntia fulgida var. fulgida]

CP: Rebman 2888 (DES). OP: Harbison 26170.

Cylindropuntia fulgida var. mamillata (Schott ex Engelmann) Backeberg [ Opuntia fulgida var.
mamillata (Schott ex Engelmann) J. M. Coulter]

OP: Near Kuakatch, 10 Apr 2000, SR (photo).

Cylindropuntia leptocaulis (de Candolle) F.M. Knuth [ Opuntia leptocaulis de Candolle]. Desert
Christmas Cholla; tasajillo', 'ajl vipinoi, ce'ecem vipinoi
CP: Rebman 2886 (SD). OP: RSF 88-470.

Cylindropuntia ramosissima (Engelmann) F. M. Knuth [ Opuntia ramosissima Engelmann],
Diamond cholla

CP: Harbison 15 Nov 1937 (SD). OP: Baker 7776 (ASU, ORPI). TA: Reeves 5394 (ASU).
Cylindropuntia spinosior (Engelmann) F. M. Knuth [ Opuntia spinosior (Engelmann) Tourney].
Cane Cholla

CP: RSF 92-546. OP: Harbison 26188 (ARIZ, SD).

Cylindropuntia versicolor (Engelmann ex J. M. Coulter) F. M. Knuth [ Opuntia versicolor
Engelmann]. Staghorn Cholla; siviri
OP. Baker 7752 (ASU).

Echinocactus polycephalus Engelmann & J. M. Bigelow var. polycephalus. Many-headed Barrel
Cactus, Cottontop Cactus, Cannonball Cactus; biznaga
CP: RSF 92-596. TA: Reeves 5426 (ASU).

Echinocereus engelmannii (Parry ex Engelmann) Lemaire. Strawberry Hedgehog Cactus; pitayita\
'isvigl

Echinocereus engelmannii var. acicularis L. D. Benson

OP: Peebles 14555.

Echinocereus engelmannii var. chrysocentrus (Engelmann & J. M. Bigelow) Rumpler
CP: Sheep Mt, 31 Jan 1992, RSF, observation. TA: Blackwell 709.

Echinocereus nicholii (L. D. Benson) B. D. Parfitt [ E . engelmannii var. nicholii L. D. Benson].
Golden Hedgehog Cactus

CP: RSF 93-280. OP: Benson 9901.

Echinocereus santaritensis W. Blum & Rutow. Claret-cup Cactus
OP: Baker 14050 (ASU).

Echinomastus erectocentrus (J. M. Coulter) Britton & Rose var. acunensis (W. T. Marshall) Bravo
[E. acunensis W. T. Marshall. Neolloydia erectocentra (J.M. Coulter) L. D. Benson var. acunensis
(W. T. Marshall) L. D. Benson], Acuna Cactus

OP: Supernaugh 2 Jan 1951 (lectotype, DES; isolectotype, ORPI).

Ferocactus cylindraceus (Engelmann) Orcutt [F. acanthodes (Lemaire) Britton & Rose. F.
acanthodes var. eastwoodiae L. D. Benson. F. acanthodes var. lecontei (Engelmann) G. E. Lindsay].
Mountain Barrel Cactus, Desert Barrel Cactus; biznaga', jiavul
CP: Benson 10809. OP: RSF 88-456. TA: RSF 90-5.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

19

Ferocactus emoryi (Engelmann) Orcutt [F. covillei Britton & Rose], Emory’s Barrel Cactus;
biznaga; jiavul

CP: Har bison 16849 (SD). OP: Pinkava 14371 (ASU).

Ferocactus wislizeni (Engelmann) Britton & Rose. Compass Barrel Cactus, Fishhook Barrel Cactus;
biznaga ; jiavul

CP: Harbison 16847 (SD). OP: Benson 9895.

Grusonia kunzei (Rose) Pinkava [Opuntia kunzei Rose. O. stanlyi Engelmann ex B. D. Jackson var.
kunzei (Rose) L. D. Benson. Corynopuntia kunzei (Rose) M. P. Griffiths], Desert Club-cholla, Devil
Cholla

CP: Hodgson 18704 (DES). OP: Mearns 2735 (CAS). TA: Harbison 16999 & 17000 (SD).
Grusonia parishii (Orcutt) Pinkava [ Opuntia parishii Orcutt. O. stanlyi Engelmann ex B.D. Jackson
var. parishii (Orcutt) L.D. Benson. O. stanlyi var. peeblesiana L.D. Benson. Corynopuntia parishii
(Orcutt) F.M. Knuth], Parish Club-cholla
OP: RSF88-110.

Lophocereus schottii (Engelmann) Britton & Rose var. schottii [Cereus schottii Engelmann.
Pachycerus schottii (Engelmann) D. R. Hunt], Senita; sinita ; ce:mT
CP: Harbison 17061 (SD). OP: Blakley 323 (DES).

Mammillaria grahamii Engelmann subsp. grahamii [Among the many synonyms are: M.
microcarpa Engelmann. M. grahamii var. arizonica Quehl. M. grahamii var. oliviae (Orcutt) L. D.
Benson. M. milleri (Britton & Rose) Bodekerj. Arizona Fishhook Cactus; cabeza de viejo; ban 'isvig,
ban cekida

CP: E side Pinacate flow, 31 Jan 1992, RSF, observation. OP: Benson 9945. TA: Reeves
5391 (ASU).

Mammillaria tetrancistra Engelmann. Corkseed Fishhook Cactus; cabeza de viejo
CP: McManus 688. OP: Steenbergh 1-662-1 (ORPI). TA: RSF 02-6.

Mammillaria thornberi Orcutt [M. fasciculata of authors, not Engelmann]. Thornber Fishhook
Cactus; cabeza de viejo ; ban mauppa
OP: Harbison 26183 (SD).

Opuntia basilaris Engelmann & J. M. Bigelow var. basilaris. Beavertail Cactus

CP: RSF 92-621. TA: Tinajas Altas Canyon, 28 Apr 2008, RSF, observation.

Opuntia chlorotica Engelmann & J. M. Bigelow. Pancake Prickly-pear; nopal rastrero
CP: Harbison 16 Mar 1937 (SD). OP: Nichol 28 Mar 1939 (ORPI).

Opuntia engelmannii Sairn-Dyck ex Engelmann. Prickly-pear; nopal

Opuntia engelmannii var. engelmannii [O. phaeacantha Engelmann var. discata (Griffiths) L. D.
Benson & Walkington], Desert Prickly-pear; naw, ’i:bhai

CP: Agua Dulce Pass, 12 Jun 1992, RSF, observation. OP: RSF 90-430.

Opuntia engelmannii var. flavispina (L. D. Benson) B. D. Parfitt & Pinkava [O. phaeacantha var.
flavispina L. D. Benson], Yellow-spine Prickly-pear; nopal

CP: RSF 92-737. OP: Nichol 27 Apr 19 39 (isotypes: ARIZ, ORPI).

**Opuntia engelmannii var. linguiformis (Griffiths) B. D. Parfitt & Pinkava [O. linguiformis
Griffiths], Cow- tongue Prickly-pear

OP: 1 mi E of Lukeville, 3 Nov 2003, SR, observation.

Opuntia phaeacantha Engelmann. Brown-spine Prickly-pear; nopal
OP: SR 2003-535 (ARIZ, ORPI).

**Opuntia santa-rita (Griffiths & Hare) Rose. Purple Prickly-pear
OP: Nichol 27 Apr 1939 (ORPI, extirpated).

Peniocereus greggii (Engelmann) Britton & Rose var. transmontanus (Engelmann) Backeberg
[Cereus greggii Engelmann var. transmontanus Engelmann], Desert Night-blooming Cereus; reina
de la noche, sarramatraca ; ho'ok wa'o

CP: Rebman 2890 (ASU). OP: RSF 90-574. TA: Mearns 2811 (CAS/DS, US).

Peniocereus striatus (Brandegee) Buxbaurn [Cereus striatus Brandegee. Neoevansia striata
(Brandegee) Sanchez-Mejorada. Wilcoxia striata (Brandegee) Britton & Rose. Cereus diguetii F. A.
C. Weber. Wilcoxia diguetii (F. A. C. Weber) Diguet & Guillaumin], Sacamatraca, sarramatraca ;
’i:kul!

OP: Nichol 20 Apr 1939.

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Stenocereus thurberi (Engelmann) Buxbaum [Cereus thurberi Engelmann. Lemaireocereus thurberi
(Engelmann) Britton & Rose]. Organ Pipe; pitaya, pitaya dulce\ cucuvis

CP: Agua Dulce Mts, photo, 2 June 1963 (Simmons 1965). OP: Hodgson 14 (ASU).

CAMPANULACEAE - BELLFLOWER FAMILY

Nemacladus orientalis (McVaugh) Morin [A. glanduliferus Jepson var. orientalis McVaugh], Redtip
Thread Plant

CP: RSF 93-329. OP: Benson 10663. TA: RSF 05-147.

Triodanis biflora (Ruiz & Pavon) Greene. Small Venus Looking-glass
OP: Bowers 1285.

CANNABACEAE - HEMP FAMILY (includes Ulmaceae, in part)

Celtis pallida Torrey subsp. pallida [C. tala Gillies ex Planchon var. pallida (Torrey) Planchon].
Desert Hackberry; garambullo\ kuavull

CP: RSF 94-25. OP: Nichol 23 Feb 1939.

Celtis reticulata Torrey. Netleaf Hackberry; ciimero
OP: Goodding 303-41 .

CAPPARACEAE - CAPER FAMILY

Atamisquea emarginata Miers ex Hooker & Arnott [Capparis atamisquea Kuntze]. Palo hediondo
OP: Harbison 26181 .

CAPRIFOLIACEAE, see ADOXACEAE

CARYOPHYLLACEAE - PINK FAMILY
Achyronychia eooperi Torrey & A. Gray. Sand-mat, Frost-mat
CP: Eiber 23. OP: Parker 7984. TA: TRY 26 Mar 1983.

Cerastium texanum Britton. Mouse-ear Chickweed
OP: RSF 05-270.

Drymaria viscosa S. Watson. Sticky Drymary
CP: Hodgson 2080 (DES).

**Herniaria hirsuta Linnaeus var. cinerea (de Candolle) Loret & Barrandon. Burstwort
OP: RSF 03-245.

Loeflingia squarrosa Nuttall [ L . squarrosa subsp. cactorum Barneby & Twisselmann], Spreading
Pygmy- leaf

CP: SR 2003-375. OP: RSF 03-184. TA: RSF 92-608.

Silene antirrhina Linnaeus. Sleepy Catchfly

CP: RSF 93-363. OP: SR 2003-445 (ORPI).

CLEOMACEAE-CLEOME FAMILY

Polanisia dodecandra (Linnaeus) de Candolle subsp. traehysperma (Torrey & A. Gray) H. H. litis
[P. traehysperma Torrey & A. Gray]. Western Clammy weed
OP: SR 23 Nov 1994 (ORPI).

Wislizenia refracta Engelmann subsp. refracta. Jackass-clover
CP: Corner Well (Simmons 1966). OP: Peebles 14558.

CONVOLVULACEAE - MORNING GLORY FAMILY
Cuscuta, see Costea (2007) and Costea and Stefanovic (2010).

Cuscuta californica Hooker & Arnott var. californica. Chaparral Dodder, California Dodder \fideo',
vepegi vasai

OP: McDougall 97.

Cuscuta legitima Costea & Stefanovic \C. umbel lata var. reflexa (J. M. Coulter) Yuncker]

CP: RSF 92-740.

Cuscuta salina Engelmann. Salt Dodder \fideo\ vepegi vasai
OP \RSF 89-241.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

21

Cuscuta tuberculata Brandegee. Desert Dodder; fideo\ vepegi vasai
OP: RSF 10532.

Cuscuta umbellata Kunth var. umbellata. Dodder; Jideo; vepegi vasai
CP: RSF 92-687-B. OP: RSF 88-414.

Evolvulus alsinoides (Linnaeus) Linnaeus [E. alsinoides var. acapulcensis (Willdenow) van

Ooststroom. E. alsinoides var. angustifolia Torrey]. Mouse Ears; oreja de raton
OP: Nichol 26 Mar 1938.

Ipomoea costellata Torrey. Crest-rib Morning-glory
OP: SR 20031026-17 (ORPI).

Ipomoea cristulata Hallier f. [/. coccinea of authors, not Linnaeus. Quamoclit gracilis Hallier f.].

Scarlet Morning-glory'

OP: Wirt 13 Oct 1990.

Ipomoea hederacea Jacquin. Ivy-leaf Morning-glory'

CP: RSF 92-746. OP: Wilson 189.

Jacquemontia pringlei A. Gray
OP: Goodding 227 .

CRASSULACEAE - STONECROP FAMILY

Crassula connata (Ruiz & Pavon) A. Berger [C. connata var. erernica (Jepson) M. Bywater &

Wickens. Tillaea erecta Hooker & Arnott], Pygmy Stonecrop
CP: RSF 93-162. OP: RSF 03-356. TA: RSF 05-26.

Dudleya arizonica Rose [D. pulverulenta (Nuttall) Britton & Rose subsp. arizonica (Rose) Moran.

Echeveria arizonica (Rose) Kearney & Peebles], Arizona Liveforever

CP: Simmons 27 Oct 1962. OP: Bowers 1268 (ORPI). TA: Kurtz 1170.

Graptopetalum rusbyi (Greene) Rose
OP: Darrow 3863.

CROSSOSOMATACEAE - CROSSOSOMA FAMILY

Crossosoma bigelovii S. Watson. Ragged Rock-flower

CP: RSF 93-86. OP: Parker 7961. TA: Webster 24254.

CUCURBITACEAE - GOURD FAMILY

Brandegea bigelovii (S. Watson) Cogniaux. Desert Star-vine

CP: Nichol 13 Nov 1937. OP: Benson 9932. TA: Goodding 2092.

Cucurbita digitata A. Gray. Finger-leaf Gourd, Coyote Gourd; calabacilla, chichi coyota ; 'adavl, 'ad
CP: RSF 92-671. OP: Simmons 1 Oct 1963. TA: Goodding 2090.

Cucurbita palmata S. Watson, Desert Coyote Gourd
TA: RSF 01-584.

Echinopepon wrightii (A. Gray) S. Watson. Wild Balsam-apple
OP: Wirt 13 Oct 1990.

Marah gilensis (Greene) Greene. Big Root
OP: Harbison 26258.

Tumamoca macdougalii Rose. Tumamoc Globe-berry
OP: Baker 7638 (ASU).

EUPHORBIACEAE - SPURGE FAMILY

Acalypha californica Bentham [A. pringlei S. Watson], California Copperleaf; hierba del cancer
CP: RSF 92-649 & Turner. OP: Van Devender 84-450. TA: Goodding 5 Dec 1935.

Chamaesyce, see Euphorbia

Croton sonorae Torrey. Sonora Croton; rama blanca
CP: RSF 92-566. OP, Bowers 944.

Croton wigginsii L. C. Wheeler. Dune Croton
CP: RSF 93-386.

Ditaxis adenophora (A. Gray) Pax & K. Hoffmann [ Argythamnia adenophora A. Gray], Glandular

Silverbush

OP: Van Devender 85-107.

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Ditaxis brandegeei (Millspaugh) Rose & Standley var. intonsa I.M. Johnston [Argythamnia
brandegeei Millspaugh var. intonsa (I. M. Johnston) J. W. Ingram]. Sonoran Silverbush
CP: Goodding 4358.

Ditaxis lanceolata (Bentham) Pax & K. Hoffmann [Argythamnia lanceolata (Bentham) Muller
Argoviensis]. Narrowleaf Silverbush

CP: Whipple 3935. OP: RSF 88-458. TA: RSF 04-72.

Ditaxis neomexicana (Muller Argoviensis) A. Heller [Argythamnia neomexicana Muller
Argoviensis]. New Mexico Silverbush

CP: RSF 92-661. OP: RSF 86-29 1. TA: RSF 04-10.

Ditaxis serrata (Torrey) A. Heller var. serrata. [Argythamnia serrata (Torrey) Muller Argoviensis],
Sand Silverbush. See Steinmann and Felger (1997).

CP: RSF 92-783. TA: Frontera Canyon, 18 Mar 1998, RSF, observation.

Euphorbia. For placement of Chamaesyce in Euphorbia, see Steinmann and Porter (2002).
Euphorbia abramsiana L. C. Wheeler [Chamaesyce abramsiana (L. C. Wheeler) Koutnik], Desert
Sandmat, Abrams’ Spurge; golondrina

CP: RSF 01-577. OP: RSF 86-277. TA: RSF 04-44.

Euphorbia albomarginata Torrey & A. Gray [Chamaesyce albomarginata (Torrey & A. Gray)
Small]. Rattlesnake 'Weed', golondrina

CP: RSF 92-562. OP: Bowers 893.

Euphorbia arizonica Engelmann [Chamaesyce arizonica (Engelmann) Arthur], Arizona Spurge;
golondrina

OP: McLaughlin 1934.

Euphorbia capitellata Engelmann [Chamaesyce capitellata (Engelmann) Millspaugh]

OP: McDougal 14 Apr 1941 .

Euphorbia eriantha Bentham. Beetle Spurge

CP: RSF 93-424. OP: Benson 9928. TA: RSF 04-12.

Euphorbia florida Engelmann [Chamaesyce florida (Engelmann) Millspaugh]

CP: RSF 92-660 (CAB). OP: Baker 7884 (ASU).

Euphorbia heterophylla Linnaeus
OP: SR 20031026-40.

Euphorbia hyssopifolia Linnaeus [Chamaesyce hyssopifolia (Linnaeus) Small], Hyssop Spurge
CP: Harlan 309. OP: Wirt 27 Jul 1990.

Euphorbia melanadenia Torrey [Chamaesyce melanadenia (Torrey) Millspaugh]. Golondrina
OP: Bezy 25 Oct 1964.

Euphorbia micromera Boissier ex Engelmann [Chamaesyce micromera (Boissier ex Engelmann)
Wooton & Standley], Golondrina

CP: Harlan 300. OP: RSF 86-294.

Euphorbia pediculifera Engelmann var. pediculifera [Chamaesyce pediculifera (Engelmann) Rose
& Standley var. pediculifera]. Louse Spurge; golondrina
CP: McLaughlin 1959. OP: Mearns 2746 (US).

Euphorbia platysperma Engelmann ex S. Watson [Chamaesyce platysperma (Engelmann ex S.
Watson) Shinners], Dune Spurge. Flat-seed Spurge
TA: Douglas 876 (ASU).

Euphorbia polycarpa Bentham [Chamaesyce polycarpa (Bentham) Millspaugh. E. polycarpa var.
hirtella Boissier], Desert Spurge; golondrina', vi'ibam

CP: Shreve 6214. OP: Parker 7934. TA: RSF 04-13.

*Euphorbia prostrata Aiton [Chamaesyce prostrata (Aiton) Small]

OP: Wirt 20 Jul 1990 (ORPI).

Euphorbia setiloba Engelmann ex Torrey [Chamaesyce setiloba (Engelmann ex Torrey) Millspaugh
ex Parish], Fringed Spurge; golondrina

Tropicos and others recognize Chamaesyce setiloba Norton, Contributions from the United
States National Herbarium 25: 345 (1925). However, Kelly Allred (personal communication 23
December 2011) provided the following opinion: Parish (1925, Cat. PI. Salton Sink 6, 1913, reprint
of Carnegie Institution Washington Publication 193, 1914) reported, ‘‘‘’Chamaesyce setiloba Millsp.
ined. Euphorbia setiloba Engelm. In sandy or loose soil in which it is often partly buried. Mounds at

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

23

the old beach line east of Holtville (8087). Sand wash in the desert southwest of Brawley (8301).”
The confusing part of this is the “ined.” after Millspaugh, leading one to believe that Parish did not
intend to effect the combination. It would seem to mean, rather, that the combination had not hitherto
been published (Millspaugh proposed it, but it was ined.) and that Parish was deliberately bringing
together the genus and the basionym as suggested by Millspaugh. Hence Millspaugh ex Parish is the
correct citation, being sufficient to make the combination effectively published.

CP: RSF 92-535. OP: LehtoL 18317 ( ASU). TA: RSF 04-14.

Euphorbia spathulata Lamarck. Warty Spurge
CP: RSF 01-569.

Euphorbia trachysperma Engelmann [Chamaesyce trachysperma (Engelmann) Millspaugh]

CP: RSF 01-568.

Jatropha cardiophylla (Torrey) Muller Argoviensis. Limberbush; sangrengado
OP: Bowers 1393.

Jatropha cinerea (Ortega) Muller Argoviensis. Ashy Limberbush; sangrengado ; komagi va:s
OP: Bowers 1392.

Jatropha cuneata Wiggins & Rollins. Desert Limberbush; sangrengado ; va:s
CP: Peebles 10862. OP: Parker 7965. TA: Harrison 6567.

Sebastiania bilocularis S. Watson [Sapium biloculare (S. Watson) Pax], Arizona Jumping Bean;
hierba de la flecha; 'ina hita

CP: RSF 94-31. OP: Mearns 2753 (US).

Stillingia linearifoiia S. Watson

CP: SR 18 Feb 2002. OP: Baker 8787 A (ORPI).

Tragia nepetifolia Cavanilles. Noseburn; ortiga
OP: Van Devender 84-453.

FABACEAE (LEGUMINOSAE) - LEGUME FAMILY
Acacia, see Acaciella, Senegalia, and Vachellia

Acaciella angustissima (Miller) Britton & Rose [ Acacia angustissima (Miller) Kuntze]. White-ball
Acacia

OP: Parker 7741.

Acmispon, see Brouillet (2008) and Jepson Flora Project (2012).

Acmispon brachycarpus (Bentham) D. D. Sokoloff [Lotus humistratus Greene]. Hill Locust
OP: McDougall 14.

Acmispon maritimus (Nuttall) D. D. Sokoloff var. brevivexillus (Ottley) Brouillet [Lotus
salsuginosus Greene var. brevivexillus Ottley. L. humilis Greene], Bird-foot Trefoil
CP: Peebles 10863. OP: Bowers 1105.

Acmispon rigidus (Bentham) Brouillet [Lotus rigidus (Bentham) Greene]. Desert Rock-pea
CP: Shreve 6228. OP: Van Devender 76-1 . TA: Harrison 3610.

Acmispon strigosus (Nuttall) Brouillet [Lotus strigosus (Nuttall) Greene var. tomentellus (Greene)
Isely. L. tomentellus Greene]. Hairy Lotus

CP: Shreve 5923. OP: Parker 7939. TA: TRV 9 Mar 1980.

Astragalus didymocarpus Hooker & Amott var. dispermus (A. Gray) Jepson. Dwarf White
Milkvetch

OP: RSF 03-260.

Astragalus insularis Kellogg var. harwoodii Munz & McBurney. Sand Locoweed
CP: Eiber 33.

Astragalus lentiginosus Douglas ex Hooker. Freckled Milkvetch
Astragalus lentiginosus var. australis Bameby
OP: Walden 19 Apr 1964 (ASU).

Astragalus lentiginosus var. yuccanus M. E. Jones
OP: Tinkham 18 Apr 1942.

Astragalus nuttallianus de Candolle var. imperfectus (Rydberg) Barneby. Small-flowered
Milkvetch

CP: RSF 93-270. OP: Nichol 24 Feb 1939.

Calliandra eriophylla Bentham var. eriophylla. Fairy Duster; huajillo
CP: RSF 93-48. OP: Bowers 1048.

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CANOTIA Vol. 8

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Coursetia glandulosa A. Gray [C. microphylla A. Gray], Sdmota
OP: Parker 7919.

Dalea mollis Bentham. Silky Dalea

CP: Goodding 1701. OP: Harbison 26218. TA: RSF 05-67.

Dalea pogonathera A. Gray var. pogonathera. Bearded Prairie-clover
OP: Parker 7922.

Dalea pringlei A. Gray var. pringlei. Pringle’s Prairie-clover
OP: Nichol 16 May 1937.

Desmodium procumbens (Miller) Hitchcock [ D . procumbens var. exiguum (A. Gray) B. G.
Schubert. D. procumbens var. transversum (B. L. Robinson & Greenman) B. G. Schubert. D.
wigginsii B. G. Schubert]. Western Tickfoil
OP: SR 20060916-2.

Galactia wrightii A. Gray. Wright’s Milkpea
OP: Bowers 1543.

Hoffmannseggia glauca (Ortega) Eifort [Caesalpinia glauca (Ortega) Kuntze], Hog Potato; camote
de raton; ’i:kovi

CP: RSF 01-556.

Lupinus arizonicus (S. Watson) S. Watson [ L . arizonicus subsp. sonorensis J. A. Christian & D. B.
Dunn], Arizona Lupine; lupino\ tas ha:hag

CP: Peebles 10891. OP: RSF 88-284. TA: TRY 25 Mar 1983.

Lupinus concinnus J. Agardh. Elegant Lupine
CP: RSF 93-349. OP: Bowers 1601.

Lupinus sparsiflorus Bentham [ L . sparsiflorus subsp. mohavensis Dziekanowski & D. B. Dunn. L.
sparsiflorus var. mohavensis (Dziekanowski & D. B. Dunn) Welsh]. Coulter’s Lupine
CP: RSF 93-327. OP: Bowers 1096.

Marina parryi (Torrey & A. Gray) Barneby [Dalea parrvi Torrey & A. Gray], Parry Dalea, Parry’s
False Prairie-clover

CP: Elias 10219. OP: Bowers 986.

**Medicago polymorpha Linnaeus [M. hispida Gaertner]. Burclover
OP: RSF 88-1 42 B (ORPI).

*Melilotus indicus (Linnaeus) Allioni. Yellow Sourclover; trebol agrio
OP: Bowers 1607.

Mimosa distachya Cavanilles var. laxiflora (Bentham) Barneby [M. laxiflora Bentham], Arizona
Mimosa; garabatillo

OP: Bowers 1517.

Nissolia schottii (Torrey) A. Gray. Schott’s Yellowhood
OP: Bowers 939.

OIneya tesota A. Gray. Ironwood; palo fierro\ ho’idkam

CP: Little Tule Well, 12 Jun 1992, RSF, observation. OP: Cooper 577. TA: RSF 08-171.
Parkinsonia, see Felger et al. (2001), Haston et al. (2005), Hawkins (1996), and Hawkins et al.
(1999).

**Parkinsonia aculeata Linnaeus. Mexican Palo Verde; bagote, retama, guacapora
OP: SR 2003-524.

Parkinsonia florida (Bentham ex A. Gray) S. Watson [Cercidium floridum Bentham ex A. Gray].
Blue Palo Verde; palo verde', ko'okomadk, kalisp

CP: RSF 93-415. OP: Nichol 26 Apr 1939. TA: Van Devender 86-11.

Parkinsonia microphylla Torrey [Cercidium microphyllum (Torrey) Rose & I. M. Johnston],
Foothill Palo Verde; palo verde-, kek cehedagT

CP: Agua Dulce Mts, 12 Jun 1992, RSF, observations. OP: RSF 90-404. TA: RSF 08-177.
Phaseolus acutifolius A. Gray var. acutifolius. Tepary; tepari del monte', bavi
OP: Wirt 13 Oct 1990.

Phaseolus filiformis A. Gray [P. wrightii A. Gray], Desert Bean; ban bavi, cepulin bavi
CP: RSF 92-780. OP: Peebles 14554 A.

Prosopis glandulosa Torrey var. torreyana (L. D. Benson) M. C. Johnston [P.juliflora (Swartz) de
Candolle var. torreyana L. D. Benson]. Western Honey Mesquite; mezquite\ kui
CP: Harlan 213 (CAB). TA: Harrison 6581 .

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

25

Prosopis pubescens Bentham. Screw Bean; tornillo; kujel
OP: RSF 86-327.

Prosopis velutina Wooton \P . juliflora (Swartz) de Candolle var. velutina (Wooton) Sargent], Velvet
Mesquite; mezquite ; kui

CP: Lehto 22487 (ASU). OP: RSF 87-270.

Psorothamnus emoryi (A. Gray) Rydberg var. emoryi [ Dalea emoryi A. Gray], Emory’s
Srnokebush

CP: RSF 92-757.

Psorothamnus fremontii (Torrey ex A. Gray) Barneby [Dalea fremontii Torrey ex A. Gray],
Fremont’s Dalea

TA: Sundt 2 Apr 2011.

Psorothamnus spinosus (A. Gray) Barneby [Dalea spinosa A. Gray], Smoke Tree; palo cenizo
CP: SR 17 Feb 2002. OP: Nichol 3 Mar 1939.

Rhynchosia texana Torrey & A. Gray [R. senna Gillies ex Hooker var. texana (Torrey & A. Gray)
M. C. Johnston. R. senna var. angustifolia (A. Gray) Grear], Texas Snout-bean. See Turner (201 1).
OP: Baker 7552 (ORPI),

Senegalia greggii (A. Gray) Britton & Rose [Acacia greggii A, Gray; A. greggii var. arizonica
Isely], Catclaw; uha de goto, gatuho\ ’u:pad. See Seigler et al. (2006)

CP: RSF 92-537 (CAB). OP: Cooper 564. TA: RSF 04-21.

Senna eovesii (A. Gray) H. S. Irwin & Barneby [Cassia covesii A. Gray]. Desert Senna; hojasen,
daisillo; ko'ov! ta.taml

CP: RSF 92-764. OP: Bowers 885.

Tephrosia vicioides Schlechtendal [T. tenella A. Gray], Red Hoary-pea
OP: SR 5 Sep 1999.

Trifolium wormskioldii Lehmann [T. wormskioldii var. arizonicum (Greene) Barneby. T. lacerum
Greene. T. willdenovii Sprengel], Cow Clover
OP: Bowers 1205 (ORPI).

Vachellia constrieta (Bentham) Seigler & Ebinger [Acacia constricta Bentham], White-thorn
Acacia; mezquitillo ; gidag. See Seigler and Ebinger (2005).

CP: RSF 94-23. OP: Bowers 928.

Vicia ludoviciana Nuttall ex Torrey & A. Gray subsp. ludoviciana [V. exigua Nuttall], Deerpea
Vetch

OP: Cooper 578.

FAGACEAE - BEECH FAMILY

Quercus turbinella Greene [Q. ajoensis C. H. Muller. Q. turbinella subsp. ajoensis (C. H. Muller)
Felger & C. H. Lowe], Scrub Oak; encinillo

OP: Muller 9519 & Tucker (isotypes of Q. ajoensis, ARIZ, ORPI).

FOUQUIERIACEAE - OCOTILLO FAMILY

Fouquieria splendens Engelmann subsp. splendens. Ocotillo; ocotillo; melhog

CP: Agua Dulce Mt, 12 Jun 1992, RSF, observations. OP: RSF 88-132. TA: RSF 04-15.

GENTIANACEAE - GENTIAN FAMILY

Eustoma exaitatum (Linnaeus) Salisbury ex G. Don subsp. exaltaturn. Catchfly Gentian
OP: Mearns 1 Feb 1 894 (US).

Zeltnera arizonica (A. Gray) G. Mansion [Centaurium arizonicum (A. Gray) A. Heller. C.
calycosum (Buckley) Fernald var. arizonicum (A. Gray) Tidestrom], Arizona Centaury
OP: RSF 86-212.

GERANIACEAE - GERANIUM FAMILY

*Erodium cicutarium (Linnaeus) L’Heritier ex Aiton. Filaree, Heron’s Bill, Redstem Stork’s Bill;
alfilerillo ; hoho'ibad

CP: Whipple 3917. OP: RSF 88-279. TA: RSF 05-143.

Erodium texanum A. Gray. Texas Filaree, Desert Stork’s Bill
CP: RSF 93-398. OP: RSF 03-335. TA: RSF 05-34.

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CANOTIA Vol. 8

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Geranium carolinianum Linnaeus. Crane’s Bill
OP: Bowers 1711.

GROSSULARIACEAE - GOOSEBERRY FAMILY
Ribes quercetorum Greene. Oak Gooseberry
OP: SR 20061022-5.

KRAMERIACEAE - RAT ANY FAMILY

Krameria bicolor S. Watson, 1886 [K. gray i Rose & Painter, 1906]. White Rhatany; cosahui
CP: Johnson 26 Mar I960. OP: Bowers 1386. TA: RSF 04-16.

Krameria erecta Willdenow ex Schultes [K. parvifolia Bentham]. Little-leaf Rhatany, Range
Rhatany; cosahui

CP: RSF 92-766. OP: Crooks 31 Mar 1937. TA: RSF 08-191.

LAMIACEAE (LABIATAE) - MINT FAMILY

Hedeoma nana (Torrey) Briquet subsp. macrocalyx W. S. Stewart. False Pennyroyal; oregano
CP: RSF 94-11 . OP: Kearney 10852.

Hyptis albida Kunth [H. emotyi Torrey]. Desert Lavender; salvia. See Felger et al. (2007) and
Fishbein and Harlan (1998).

CP: 93-440. OP: McDougall 61. TA: Kurtz 1161.

Monardella arizonica Epling. Arizona Monardella
OP: Baker 7640 (ORPI).

Salvia columbariae Bentham. Desert Chia; chia\ da:pk
CP: Simmons (1966). OP: RSF 88-281.

Salvia pinguifolia (Fernald) Wooton & Standley. Rock Sage
OP: Darrow 3850.

Salvia vaseyi (Porter) Parish [Audibertia vaseyi Porter]. Scallop-leaf Sage
CP: Cain & Jansen 15 Nov 2003.

Scutellaria mexicana (Torrey) A.J. Paton [ Salazaria mexicana Torrey], Bladder-Sage. See Patton
(1990).

CP: RSF 92-51. OP: Wilson 9 Apr 1993 (ORPI). TA: Gentry 3524.

Teucriuni cubense Jacquin subsp. depressum (Small) E. M. McClintock & Epling. Dwarf
Germander

CP: RSF 93-99. OP: Wirt 12 Apr 1989.

Teucrium glandulosum Kellogg. Desert Germander

CP: Simmons 26 Oct 1962. OP: SR Mar 31 2005. TA: RSF 02-14.

LINACEAE - FLAX FAMILY

Linum lewisii Pursh [ L . perenne L. var. lewisii (Pursh) Eaton & Wright]. Prairie Flax
OP: McDougall 9 Apr 1941 .

LOASACEAE - STICKLEAF FAMILY

Eucnide rupestris (Baillon) H. J. Thompson & W. R. Ernst [Sympetaleia rupestris (Baillon) A. Gray
ex S. Watson]. Rock-nettle, Velcro Plant

CP: Monson 1 . OP: Wirt 5 Dec 1990.

Mentzelia affinis Greene. Triangle-seed Blazing Star; pegapega
CP: McLaughlin 2976. OP: RSF 88-287.

Mentzelia albicaulis (Douglas ex Hooker) Douglas ex Torrey & A. Gray (this species complex
includes M. desertorum (Davidson) H. J. Thompson & J. E. Roberts). White-stem Blazing Star
CP : McLaughlin 2986. TA: TRY 6 Mar 1983.

Mentzelia involucrata S. Watson [M. involucrata var. megalantha I. M. Johnston], Silver Blazing
Star; pegapega

CP: Kearney & Peebles 11012. OP: Parker 7970.

Mentzelia isolata Gentry

OP: Wirt 13 Aug 1990.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

27

Mentzelia lortgiloba J, Darlington var. yavapaiensis J. J. Schenk & L. Hufford. Blazing Star;
pegapega. See Schenk and Hufford (2010).

CP; Yatskievych 79-242. OP: Har bison 26217. TA: Frontera Canyon, 18 Mar 1998, RSF,
observation.

Mentzelia puberula J. Darlington. Argus Blazing Star

CP; Cain 15 Nov 2003. TA; Reichenbacher 480.

Petalonyx linearis Greene. Narrowleaf Sandpaper Plant
OP; Bowers 1259. TA; Goodding 5335.

Petalonyx thurberi A. Gray subsp. thurberi. Thurber Sandpaper Plant; hadsadkam
CP: Simmons 19 May 1963 (CAB). OP: Bowers 1723.

LYTHRACEAE - LOOSESTRIFE FAMILY (includes Punicaceae)

**Punica granatum Linnaeus. Pomegranate; granada; galna:yu
OP: TRY 31 Aug 1978.

MALPIGHIACEAE - MALPIGHIA FAMILY

Cottsia gracilis (A. Gray) W. R. Anderson & C. Davis [Janusia gracilis A. Gray], Fermina. See
Anderson and Davis (2007).

CP: RSF 93-287. OP: RSF 88-125.

MALVACEAE - MALLOW FAMILY (includes Sterculiaceae)

Abutilon abutiloides (Jacquin) Garke ex Britton & P. Wilson. Pintapan

OP: Bowers 990.

Abutilon incanum (Link) Sweet [A. pringlei Hochreutiner] . Pelotazo
CP: RSF 92-728. OP: Parker 7906.

Abutilon malacurn S. Watson. Yellow Indian Mallow
CP: RSF 93-8. OP: Wirt 11 Nov 1988.

Abutilon palmeri A. Gray. Palmer’s Indian Mallow
CP: Lehto L23521 (ASU). OP: Benson 9923.

Anoda abutiloides A. Gray. Indian Anoda
OP: SR 8 Nov 1995 (ORPI).

Anoda pentaschista A. Gray. Field Anoda
OP: SR 24 Sep 2006.

Ayenia filiformis S. Watson [A. pusilla of various authors, not A pusilla Linnaeus]. Dwarf Ayenia
CP: RSF 92-736. OP: Bowers 1020.

Ayenia microphylla A. Gray. Shrub Ayenia

OP: Fonts 441.

Eremalche exilis (A. Gray) Greene. White Mallow

CP: Reeves 6787 (ASU). OP: McDougall 70.

Herissantia erispa (Linnaeus) Brizicky. Bladder Mallow
CP: RSF 92-770. OP: Parker 7949.

Hermannia pauciflora S. Watson. Dwarf Burstwort
OP: Fallon 30 Mar 2011.

Hibiscus biseptus S. Watson. Arizona Rose-mallow

OP: Baker 7557 (ASU, mixed collection with H. coulteri).

Hibiscus coulteri Harvey ex A. Gray. Desert Rose-mallow
CP: RSF 93-286. OP: Harbison 26248.

Hibiscus denudatus Bentham var. denudatus. Rock Hibiscus

CP: Shreve 5924. OP: McDougall 59. TA: Reeves R5431 (ASU).

Horsfordia alata (S. Watson) A. Gray. Pink Velvet-mallow

CP: Shreve 5932. OP: Bowers 1526. TA: Kurtz 1158.

Horsfordia newberryi (S. Watson) A. Gray. Orange Velvet-mallow

CP: RSF 92-571 . OP: Darrow 2411. TA: Borrego Canyon, 16 Jun 1992, RSF, observation.
*Malva parviflora Linnaeus. Cheeseweed; malva, quesitos; tas ha:hag, ha:hag cu’igkam
OP: Bowers 1563.

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CANOTIA Vol. 8

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Malvastrum bicuspidatum (S. Watson) Rose subsp. bicuspidatum. Shrubby False Mallow
OP: RSF 90-553.

Malvella leprosa (Ortega) Krapovickas [Sida leprosa (Ortega) K. Schumann], Alkali Mallow
CP: RSF 01-563 .

Malvella sagittifolia (A. Gray) Fryxell [Sida lepidota A. Gray var. sagittifolia A. Gray], Arrow-leaf
Alkali Mallow

CP: Simmons 4 Oct 1964.

Rhynchosida physocalyx (A. Gray) Fryxell [SWa physocalyx A. Gray]

OP: Warren 10 Nov 1983.

Sida abutifolia Miller [Sidafdicaulis Torrey & A. Gray. S. procumbens Swartz]

OP: RSF 03-423.

Sphaeralcea ambigua A. Gray subsp. ambigua. Desert Globemallow; mal de ojo
CP: Shreve 6223. OP: Bowers 1145. TA: RSF 04-78.

Sphaeralcea coulteri (S. Watson) A. Gray [5. orcuttii Rose], Annual Globemallow; mal de ojo\
hadam tatk, niatum. See Felger (2000) and Felger et al (2007b).

CP: Kearney 10900. OP: Parker 7943. TA: RSF 05-141.

Sphaeralcea emoryi Torrey ex A. Gray. Emory Globemallow; mal de ojo
CP: RSF 92-15. OP: Benson 9921. TA: RSF 10-167.

Sphaeralcea laxa Wooton & Standley. Caliche Globemallow
OP: Goodding 297-41 .

MARTYNIACEAE - DEVIL’S CLAW FAMILY

Proboscidea altheifolia (Bentham) Decaisne. Desert Devil’s Claw, Desert Unicorn-plant; cuernitos,
una de gato\ ban 'ihugga

CP: RSF 92-767. OP: Gould 3217. TA: RSF 04-18.

Proboscidea parviflora (Wooton) Wooton & Standley subsp. parviflora. Devil’s Claw; cuernitos,
una de gato\ 'ihug

" OP: Wirt 13 Aug 1990.

MOLLUGINACEAE — CARPET- WEED FAMILY
*Glinus lotoides Linnaeus
CP: RSF 92-565.

*Mollugo cerviana (Linnaeus) Seringe. Thread-stem Carpet-weed, Indian Chickweed
CP: RSF 92-715. OP: RSF 88-419. TA: RSF 04-50.

MONTIACEAE — MINER’S LETTUCE FAMILY

Calandrinia ciliata (Ruiz & Pavon) de Candolle [C. ciliata var. menziesii (Hooker) J. F. Macbride],
Fringed Redmaids

OP: RSF 03-264.

Cistanthe monandra (Nuttall) Hershkovitz [Calyptridium monandrum Nuttall], Sand-cress
OP: Nichol 14 Mar 1939.

Claytonia perfoliata Donn ex Willdenow subsp. mexicana (Rydberg) John M. Miller & K. L.
Chambers [Montia perfoliata (Donn ex Willdenow) Howell. M. mexicana (Rydberg) Standley &
Steyermark], Miner’s Lettuce

OP: Nichol 14 Mar 1939.

Phemeranthus aurantiacus (Engelmann) Kiger [Talinum aurantiacum Engelmann], Orange Flame-
flower

OP: Bowers 1798.

MORACEAE - MULBERRY FAMILY
**Ficus carica Linnaeus. Fig; higuera; su:na
OP: RSF 88-452 (extirpated).

Morus microphylla Buckley. Littleleaf Mulberry; mora cimarrona\ gohi
OP: Bowers 1291 .

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

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NYCTAGINACEAE - FOUR-O’CLOCK FAMILY
Abronia villosa S. Watson var. villosa. Sand-verbena; verbana de la arena
CP: Benson 10783. TA: TRV 27 Mar 1983.

Acleisanthes longiflora A. Gray. Ange! Trumpets
CP: RSF 92-564.

AlHonia incarnata Linnaeus [A. incarnata var. nudata (Standley) Munz. A. incarnata var. villosa
(Standley) Munz], Windmills, Trailing Four-o’clock

CP: RSF 92-678. OP: Nichol 14 Mar 1939. TA: RSF 04-23.

*Boerhavia coccinea Miller [5. caribaea Jacquin], Scarlet Spiderling
CP: RSF 92-63 7. OP: TR V 85-114. TA: RSF 92-609.

Boerhavia coulteri (Hooker f.) S. Watson var. palmeri (S. Watson) Spellenberg [B. spicata Choisy
var. palmeri S. Watson]. Spiderling; juantilipln

CP: RSF 92-676. OP: Wirt 25 Jul 1990. TA: RSF 04-29.

Boerhavia erecta Linnaeus. Spiderling; juantilipln-, makkurm ha-jeved
CP: RSF 92-692. OP Gould 3214. TA: RSF 04-28.

Boerhavia rnegaptera Standley. Large-winged Spiderling
OP: Bowers 954.

Boerhavia pterocarpa S. Watson. Wing-fruit Spiderling
OP: Bowers 1533 (ORPI).

Boerhavia triquetra S. Watson [B. intermedia M. E. Jones. B. triquetra var. intermedia (M. E.
Jones) Spellenberg. B. maculata Standley]. Spiderling; juanamipili, juantilipin. See Spellenberg
(2007).

CP: RSF 01-572. OP: Warren 1 1 Aug 1975.

Boerhavia wrightii A. Gray. Large-bract Spiderling; juantilipln
CP: RSF 92-677. OP: Bowers 1530. TA: RSF 04-08.

Commicarpus scandens (Linnaeus) Standley. Climbing Wart-club

CP: E margin of Cabeza Prieta, 23 Feb 2003, SR, observation. OP: RSF 91-136.

Mirabilis laevis (Bentham) Curran var. villosa (Kellogg) Spellenberg [M. bigelovii A. Gray], Desert
Four O’clock

CP: Shreve 5931. OP: Benson 10678. TA: Goodding 1517.

Mirabilis multiflora (Torrey) A. Gray var. multiflora. Colorado Four O’clock
OP: SR 20031026-43.

Mirabilis tenuiloba S. Watson. Long-lobed Four O’clock
TA: RSF 92-613.

QLEACEAE - OLIVE FAMILY

Forestiera phillyreoides (Bentham) Torrey [F. shrevei Standley]. Desert Olive
OP: Shreve 6201 (isotype of F. shrevei).

Fraxinus velutina Torrey. Velvet Ash; fresno

OP: Benson 10686 (native? extirpated?).

Menodora scabra A. Gray [M. scoparia Engel mann ex A. Gray]. Twinberry
C'P: Harlan 392. OP: Darrow 2429.

ONAGRACEAE - EVENING-PRIMROSE FAMILY

For revisions in Onagraceae see Wagner et al. (2007).

Chylismia arenaria A. Nelson [Camissonia arenaria (A, Nelson) P. H. Raven]. Fortuna Suncup
CP: TRV 9 Mar 1980. TA: Harrison 3608.

Chylismia claviformis (Torrey & Fremont) A. Heller subsp. peeblesii (Munz) W. L. Wagner &
Hoch [Camissonia claviformis (Torrey & Fremont) P. H. Raven subsp. peeblesii (Munz) P. H.
Raven]. Browneyes

CP: Furlow 14 Mar 1979. OP: Bowers 1104. TA: TRV 5 Mar 1983.

Chylismia claviformis subsp. rubescens (P. H. Raven) W. L. Wagner & Hoch [Camissonia
claviformis subsp. rubescens (P. H. Raven) P. H. Raven], Browneyes
CP: Shreve 6211. OP: Bowers 1138.

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Epilobium canum (Greene) P. H. Raven subsp. latifolium (Hooker) P. H. Raven [Zauschneria
californica C. Presl subsp. latifolia (Hooker) D. D. Keck]. California Fuchsia, Hummingbird
Trumpet

OP: Nichol 14 Mar 1939.

Eremothera boothii (Douglas) W. L. Wagner & Hoch subsp. condensata (Munz) W. L. Wagner &
Hoch [Camissonia boothii (Douglas) P. H. Raven subsp. condensata (Munz) P. H. Raven]. Woody
Bottle-washer

CP: Niles 356. OP: Bowers 1097.

Eremothera chamaenerioides (A. Gray) W. L. Wagner & Hoch [Camissonia chamaenerioides (A.
Gray) P.H. Raven]. Willow-herb Evening-primrose

CP: RSF 93-121. OP: Bowers 1106. TA: TRV 5 Mar 1983.

Eulobus californicus Nuttall ex Torrey & A. Gray [ Camissonia californica (Nuttall ex Torrey & A.
Gray) P.H. Raven. Oenothera leptocarpa Greene]. California Suncup
CP: RSF 93-41 7. OP: McDougall 45. TA: TRV 5 Mar 1983.

Oenothera arizonica (Munz) W. L. Wagner [O. deltoides Torrey & Fremont var. arizonica Munz.
O. californica (S. Watson) S. Watson subsp. arizonica (Munz) W.M. Klein. O. avita (W. M. Klein)
W. M. Klein subsp. arizonica (Munz) W. M Klein], Arizona Evening-primrose
CP: Shreve 6222. OP: Kearney 10856.

Oenothera curtiflora W. L. Wagner & Hoch [Gaura parviflora Douglas ex Lehmann], Lizard-tail,
Velvet-leaf Gaura

CP: Benson 10748. OP: RSF86-174A.

Oenothera deltoides Torrey & Fremont subsp. deltoides. Dune Evening-primrose, Devil’s Lantern
CP: RSF 92-37.

Oenothera primiveris A. Gray. Yellow Desert Evening-primrose

CP: Shreve 6222. OP: Nichol 24 Feb 1939. TA: RSF 04-54.

OROBANCHACEAE - BROOMRAPE FAMILY

Castilleja exserta (A. Heller) T. 1. Chuang & Heckard subsp. exserta [ Orthocarpus exsertus A.
Heller. O. purpurascens Bentham], Purple Owl’s-clover

CP: Cutler 17 Mar 1995 (CAB). OP: Benson 10656.

Castilleja lanata A. Gray. Indian Paintbrush
OP: Nichol 4 May 1939.

Orobanche cooperi (A. Gray) A. Heller. Desert Broomrape; mo'otadk
CP: SR 18 Feb 2002. OP: RSF 88-300. TA: TRV 6 Mar 1983.

Orobanche fasciculata Nuttall. Yellow Broomrape
OP: SR 2003-497 { ORPI).

OXALIDACEAE - OXALIS FAMILY
Oxalis albicans Kunth. Wood sorrel
OP: Darrow 3854.

PAPAVERACEAE - POPPY FAMILY

Argemone gracilenta Greene. Prickly Poppy, Cowboy’s Fried Eggs; cardo
CP: RSF 92-11. OP: Cooper 611.

*Argemone ochroleuca Sweet subsp. ochroleuca. Mexican Prickly Poppy; cardo
OP: Johnson 25 Apr 1992 (ASU).

Eschscholzia californica Chamisso subsp. mexicana (Greene) C. Clark. Mexican Gold-poppy;
amapolita del campo; ho:hi ’e’es
OP: Niles 556.

Eschscholzia minutiflora S. Watson. Little Gold-poppy

CP: SR 2003-381. OP: RSF 88-290. TA: TRV 5 Mar 1983.

PHRYMACEAE - MONKEY-FLOWER FAMILY
Mimulus guttatus Fischer ex de Candolle. Seep Monkey-flower
OP: Bowers 1206.

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31

Mimulus rubellus A. Gray. Little Monkey-flower
OP: Shreve 6202.

PHYTOLACCACEAE - POKE WEED FAMILY
Rivina humilis Linnaeus. Rouge Plant, Pigeon Berry
OP: Goodding 294-41.

PLANT AGINACEAE - PLANTAIN FAMILY (includes Scrophulariaceae, in part)

Antirrhinum. Vargus et al. (2004) recognize Neogaerrhinum and Sairocarpus; also see Oyama and
Baum (2004). We are retaining Antirrhinum for these genera following the Jepson Flora Project
(2012).

Antirrhinum filipes A. Gray [Asarina filipes (A. Gray) Pennell. Neogaerrhinum filipes (A. Gray)
Rothmaler], Yellow Twining Snapdragon

CP: SR 2003-27. OP: McDougall 84.

Antirrhinum nuttallianum Bentham subsp. subsessile (A. Gray) D. M. Thompson [ Sairocarpus
pusillus (Brandegee) D. A. Sutton], Lesser Snapdragon
OP: Thompson 264.

Antirrhinum watsonii Vasey & Rose [A. kingii S. Watson var. watsonii (Vasey & Rose) Munz.
Sairocarpus watsonii (Vasey & Rose) D. A. Sutton], Watson’s Snapdragon
OP: Baker 7707 (ASU, ORPI).

Keckiella antirrhinoides (Bentham) Straw subsp. microphylla (A. Gray) Straw \Penstemon
microphyihis A. Gray]. Bush Penstemon
OP: Bowers 1281 .

Maurandya antirrhiniflora Humboldt & Bonpland ex Willdenow [Asarina antirrhiniflora
(Humboldt & Bonpland ex Willdenow) Pennell. Maurandella antirrhiniflora (Humboldt & Bonpland
ex Willdenow) Rothmaler]. Blue Snapdragon-vine, Blue Twining Snapdragon
CP: RSF 92-544. OP: Bowers 933.

Nuttallanthus texanus (Scheele) D.A. Sutton [L inaria texana Scheele. L. canadensis (Linnaeus)
Dumont de Courset var. texana (Scheele) Pennell]. Blue Toadflax, Texas Toadflax
OP: TRV 10 Mar 1978.

Penstemon parryi (A. Gray) A. Gray. Desert Penstemon; jaritos, varita de San Jose; hevel 'e'es
CP: Childs Mt, rare, Mar 2004, Curtis McCasland, observation. OP: Gould 3006. TA:
Tinajas Altas, rare, 10 Jan 2002, RSF, observation.

Penstemon pseudospectabilis M. E. Jones subsp. pseudospectabilis. Mojave Beard-tongue
CP: RSF 93-160. OP: Lockwood 161. TA: Hodgson 2723 (DES).

Plantago ovata Forsskal var. fastigiata (E. Morris) S. C. Meyers & Liston [P. insularis Eastwood.
Not P. insularis Nyman ex Briquet. P. fastigiata E. Morris. P. insularis var. fastigiata (E. Morris)
Jepson], Woolly Plantain, Indian Wheat; pastora; mumsa

CP: RSF 93-320. OP: Parker 7908. TA: RSF 05-123.

Plantago patagonica Jacquin [P. purshii Roemer & Schultes], Pastora
CP: RSF 92-560. OP: SR 2003-454.

Pseudorontium cyathiferum (Bentham) Rothmaler [Antirrhinum cyathiferum Bentham], Desert
Snapdragon. See Oyama and Baum (2004) and Vargus et al. (2004).

CP: Benson 10792. OP: Baker 7715 (ASU). TA: Frontera Canyon, 18 Mar 1998, RSF,
observation.

Stemodia durantifolia (Linnaeus) Swartz. Purple Stemodia

OP: Dakan 27 Feb 1972 (ORPI). Van Devender 87-5.

Veronica peregrina Linnaeus subsp. xalapensis (Kunth) Pennell. Purslane Speedwell, Necklace-
weed

CP: RSF 92-563. OP: Gould 2987.

PLUMBAGINACEAE - LEADWORT FAMILY

Plumbago zeylanica Linnaeus [P. scandens Linnaeus]. Doctor-bush; estrenina
OP: Parker 7744.

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POLEMONIACEAE - PHLOX FAMILY

Aliciella Intifolia (S. Watson) J. M. Porter subsp. latifolia [Gilia latifolia S. Watson]. Broad-leaf
Gilia

CP: Benson 10795. TA: Halse 31 Mar 1973.

Eriastrum diffusum (A. Gray) H. Mason. Woolly-star

CP: Kearney 10870. OP: RSF 88-293. TA: TRV 6 Mar 1983.

Eriastrum eremieum (Jepson) H. Mason subsp. yageri (M. E. Jones) H. Mason. Desert Woolly-star
OP: Tinkham Apr 1942.

Gilia flavocincta A. Nelson subsp. australis (A. D. & V. E. Grant) A. G. Day & V. E. Grant [G.
ophthalmoides Brand subsp. australis (A. Nelson) A. D. & V. E. Grant], Lesser Yellowthroat Gilia
OP: McDougall 44.

Gilia stellata A. Heller. Star Gilia

CP: Monson 20 Mar 1958. OP: Kearney 10811. TA: McLaughlin 1972.

Ipomopsis multiflora (Nuttall) V.E. Grant [Gilia multiflora Nuttall], Many-flowered Gilia
OP: Baker 7641 (ASU).

Langloisia setosissima (Torrey & A. Gray) Greene subsp. setosissima. Bristly Langloisia
CP: SR 2003-34. OP: Bowers 1119. TA: RSF 98-102.

Leptosiphon aureus (Nuttall) J. M. Porter & L. A. Johnson subsp. aureus [Linanthus aureus
(Nuttall) Greene var. aureus ]. Golden Desert-trumpets
OP: McDougall 39.

Linanthus bigelovii (A. Gray) Greene. Bigelow’s Desert-trumpet
CP: RSF 92-39.

Linanthus cf. bigelovii (large-flowered form with short simple leaves)

CP: RSF 93-427.

Linanthus cf. bigelovii x L. jonesii (possible hybrid)

CP: Bowers 1228. OP: Whipple 3925.

Linanthus demissus (A. Gray) Greene. Desert Snow
OP: McDougall 68.

Linanthus jonesii (A. Gray) Greene. Jones’ Desert-trumpet

CP: Benson 10762A (RSA). OP: Gould 2994. TA: TR 26 Mar 1983.

Linanthus cf. jonesii (large-flowered form, perhaps an unnamed taxon)

CP: Mason 3601.

Loeseliastrum schottii (Torrey) Timbrook \ Langloisia schottii (Torrey) Greene]. Schott’s Calico
TA: TRV 27 Mar 1983.

Phlox tenuifolia E.E. Nelson. Mountain Phlox

OP: Bowers 932.

POLYGALACEAE - MILKWORT FAMILY
Polygala macradenia A. Gray. Gland-leaf Milkwort
OP: Darrow 3857.

POLYGONACEAE - BUCKWHEAT FAMILY

Chorizanthe brevicornu Torrey subsp. brevieornu. Brittle Spineflower, Short-horn Spineflower
CP: Niles 344. OP: Benson 10675. TA: TRV 5 Mar 1983.

Chorizanthe corrugata (Torrey) Torrey & A. Gray. Wrinkled Spineflower
CP: RSF 93-394. OP: SR 6 Apr 1998 (ORPI).

Chorizanthe rigida (Torrey) Torrey & A. Gray. Devil’s Spineflower; tapacola
CP: RSF 92-1. OP: Parker 7923. TA: RSF 05-33.

Eriogonum abertianum Torrey. Abert’s Wild Buckwheat
OP: Kearney 10833.

Eriogonum deflexum Torrey var. deflexum [E. deflexum var. turbinatum (Small) Reveal]. Skeleton
Weed

CP: SR 22 Mar 2003. OP: Parker 7979.

Eriogonum fasciculatum Bentham var. polifolium (Bentham) Torrey & A. Gray. Flat-top
Buckwheat

CP: RSF 92-52. OP: Parker 8009. TA: Shreve 5942.

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Eriogonum inflaturn Torrey & Fremont. Desert Trumpet, Bladder Stem
CP: RSF 93-339. OP: RSF 88-460. TA: Reeves 5441 (ASU).

Eriogonum maculatum A. Heller. Spotted Wild Buckwheat
OP: Walden 18 Apr 1964 (ASU).

Eriogonum thomasii Torrey. Thomas’ Wild Buckwheat

CP: Shreve 6.213. OP: Parker 7968. TA: TRV 5 Mar 1983.

Eriogonum thurberi Torrey. Thurber’s Wild Buckwheat

CP: RSF 93-438. OP: Lehto 15443-f TA: RSF 05-93.

Eriogonum trichopes Torrey. Little Desert Trumpet

CP: RSF 92-776. OP: Bowers 1253. TA: RSF 05-28.

Eriogonum wrightii Torrey ex Bentham var. nodosum (Small) Reveal [E. wrightii var. pringlei (J.
M. Coulter & Fisher) Reveal]. Bastard-sage

CP: McLaughlin 1965. OP: RSF 90-562. TA: Goodding 1449.

Nemacaulis denudata Nuttall var. gracilis Goodman & L. D. Benson. Woolly Heads
CP: Eiber 27. TA: TRV 27 Mar 1983.

*Polygonum argyrocoleon Steude! ex Kunze. Silver-sheath Knotweed
CP: RSF 92-562 (CAB). OP: RSF 05-161.

Pterostegia drymarioides Fischer & C. A. Meyer. Woodland Threadstem
OP: Kearney 10815.

Rurnex hymenosepalus Torrey. Sand Dock, Wild-rhubarb; cahaigre; wakondam
OP: McDougall 82.

PORTULACACEAE - PURSLANE FAMILY (in part see MONTIACEAE and TALINACEAE)
Portulaca halimoides Linnaeus [P. parvula A. Gray]. Silk-cotton Purslane
CP Harlan 305. OP: RSF 87-281. TA: RSF 04-61.

*Portulaca oleracea Linnaeus. Purslane; verdolaga ; ku'ukpalk
CP: RSF 01-554. OP: RSF 87-318.

Portulaca suffrutescens Engelmann
OP: SR 20060924-5.

Portulaca umbra ticola Kunth subsp. lanceolata (Engelmann) J. F. Matthews & Ketron [P.
lanceolata Engelmann]. Winged Purslane
OP: Wilson 193.

PRIMULACEAE - PRIMROSE FAMILY
**Anaga!lis arvensis Linnaeus subsp. arvensis. Scarlet Pimpernel
OP: RSF 88-138.

Androsace occidentalis Pursh [A. occidentals var. arizonicus (A. Gray) H. St. John]. Western Rock-
jasmine

OP: Kearney 10851 .

RANUNCULACEAE - RANUNCULUS FAMILY
Anemone tuberosa Rydberg. Desert Windflower

OP: RSF 05-266 .

Clematis drummondii Torrey & A. Gray. Texas Virgin-bower; barba de chivato
CP: RSF 92- 722. OP: Harbison 25672.

Delphinium scaposurn Greene. Bare-stern Larkspur; espulita cimarrona
CP: RSF 93-309. OP: Parker 7960.

Myosurus cupulatus S. Watson. Mousetail
OP: RSF 05-199.

Myosurus minimus Linnaeus. Dwarf Mousetail
OP: Gould 2986 (extirpated)

RESEDACEAE - MIGNONETTE FAMILY
Oligomeris linifolia (Vahl) J. F. Macbride. Desert Cambess

CP: Benson 10776. OP: Nichol 28 Apr 1939. TA: RSF 04-53.

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RHAMNACEAE - BUCKTHORN FAMILY

Condalia globosa l.M. Johnston var. pubescens I.M. Johnston. Crucillo; kawk kuavuli
CP: RSF 94-21. OP: Peebles 14329.

Frangula betulifolia (Greene) Grubov subsp. betulifolia [ Rhammis betulifolia Greene]. Birchleaf
Buckthorn; salicieso

OP: Nichol. 23 Feb 1939.

Rhamnus erocea Nuttall. Holly-leaf Buckthorn, Holly-leaf Redberry. (We are unable to distinguish
R. crocea in Organ Pipe from R. ilicifolia Kellogg; Wolf (1938) considered them to be conspecific,
e.g., R. crocea subsp. ilicifolia (Kellogg) C. B. Wolf.)

OP: Parker 8001 .

Ziziphus obtusifoiia (Hooker ex Torrey & A. Gray) A. Gray var. canescens (A. Gray) M. C.
Johnston [ Condalia lycioides (A. Gray) Weberbauer var. canescens (A. Gray) Trelease]. White
Crucillo, Graythorn; abrojo

CP: Harlan & Steinmann 283 (CAB). OP: RSF 89-233. TA: RSF 08-170.

ROSACEAE - ROSE FAMILY

Vauquelinia ealifornica (Torrey) Sargent subsp. sonorensis W. J. Hess & Henrickson. Sonoran
Rosewood

OP: Nichol 23 Feb 1939.

RUBIACEAE - MADDER FAMILY

Galium aparine Linnaeus. Stickywilly, Goose-grass Bedstraw
OP: Kearney & Peebles 10845.

Galium microphyllum A. Gray. Bract-leaf Bedstraw
OP: RSF 03-415.

Galium stellatum Kellogg [ G . stellatuni var. eremicnm Hilend & J. T. Howell], Starry Bedstraw
CP: Shreve 5937. OP: Bowers 1251 . TA: Harrison & Kearney 6574.

RUTACEAE - RUE OR CITRUS FAMILY
Ptelea trifoliata Linnaeus. Hop Tree

OP: Harbison 13 Dec 1939 (SD).

Thamnosma montana Torrey & Fremont. Turpentine-broom
CP: Engard 914 (DES). TA: Harrison & Belden 3612.

SALICACEAE - WILLOW FAMILY

Populus fremontii S. Watson subsp. fremontii. Fremont Cottonwood; alamo; 'auppa
OP: Peebles 14563.

Salix gooddingii C. R. Ball. Goodding Willow; sauce, sauz; ce'ul
OP: Peebles 14562.

SANTALACEAE - SANDALWOOD FAMILY (includes Viscaceae)

Phoradendron ealifornicum Nuttall. Desert Mistletoe; /p/7; ha:kvad
CP: RSF 92-9. OP: Mearns 2742 (US). TA: RSF 04-59.

SAPINDACEAE - SOAPBERRY FAMILY

Dodonaea viscosa Jacquin var. angustifolia (Linnaeus f.) Bentham. Hop-bush; tarachico
OP: Lockwood 159.

Sapindus drummondii Hooker & Arnott [S. saponaria Linnaeus var. drummondii (Hooker &
Amott) L. D. Benson], Soapberry; jaboncillo, amolillo
OP: Bowers 1340.

SAURURACEAE, see MAGNOLIIDS

SCROPHULARIACEAE, see OROBANCHACEAE, PHRYMACEAE, and
PLANTAGINACEAE

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35

SIMAROUBACEAE - QUASSIA FAMILY

Castela emoryi (A. Gray) Moran & Felger [Holaccmtha emoryi A. Gray], Crucifixion-thorn; corona
de cristo

CP: RSF 92-545. OP: Gould 3012.

SIMMONDSIACEAE - JOJOBA FAMILY

Simmondsia chinensis (Link) C. K. Schneider. Jojoba, Goat-nut; jojoba; hohovai
OP: Tibbitts 6 Mar 2003.

SOLANACEAE - NIGHTSHADE or POTATO FAMILY

Capsicum annuum Linnaeus var. glabriusculum (Dunal) Heiser & Pickersgill [ C . indicum
Dierbach var. aviculare Dierbach. C. annuum var. aviculare (Dierbach) D'Arcy & Eshbaugh].
Chiltepin; chiltepin; a'al ko'okol
OP: Wilson 207.

Chamaesaracha coronopus (Dunal) A. Gray. False Nightshade
CP: Hardy [Hardies] 3 Dec 1935. TA: RSF 04-63.

Datura discolor Bernhardi. Desert Datura; toloache

CP RSF 92-778. OP: Gould 3216. TA: RSF 04-64.

Lycium andersonii A. Gray var. andersonii [L. andersonii var. deserticola (C. L. Hitchcock)
Jepson]. Desert Wolfberry; salicieso; s-toa kuavulT

CP: SR 2003-31. OP: RSF 90-47. TA: RSF 08-196.

Lycium berlandieri Dunal var. longistylum C. L. Hitchcock. Bachata, salicieso
CP: TRV 9 Mar 1980. OP: Bowers 908.

Lycium brevipes Bentham var. brevipes [ L . richii A. Gray]. Salicieso
OP: SR 2003-107 (ORPI).

Lycium ealifornicum Nuttall ex A. Gray var. californicum [L. californicum var. arizonicum A.
Gray]. California Desert-thorn

CP: SR 20050115-8 (specimen not located). OP: RSF 90-42.

Lycium exsertum A. Gray. Arizona Desert-thorn; salicieso
OP: Bowers 1564.

Lycium fremontii A. Gray var. fremontii. Fremont Wolfberry; salicieso; kuavulT
CP: RSF 92-42. OP: Bowers 1011. TA: McLaughlin 1967.

Lycium macrodon A. Gray. S-cuk kuavulT

CP: RSF 93-168. OP: Mearns 2740 (CAS). TA: Harrison 6571.

Lycium parishii A. Gray. Parish Wolfberry; salicieso

CP: Harrow 14 Apr 1941. OP: Shreve 6208. TA: TRV 9 Mar 1980.

Nicotiana clevelandii A. Gray. Desert Tobacco; tabaquillo del coyote; ban vivga
CP: Benson 10810. OP: Bowers 1044 (ORPI). TA: Kearney 10913.

**Nicotiana glauca Graham var. glauca. Tree Tobacco

OP: Hwy 85, near Visitor Center, 5 Sep 2005, SR, observation.

Nicotiana obtusifolia M. Martens & Galeotti [N. trigonophylla Dunal], Coyote Tobacco, Desert
Tobacco; tobaco de coyote; O'odham ha-vivga

CP: Cain 15 Nov 2003. OP: Mearns 2744 (CAS/DS). TA: Harrison 6568.

Petunia parviflora de Jussieu [Calibrachoa parviflora (de Jussieu) D'Arcy], Small-flowered Petunia
OP: RSF 88-317.

Physalis acutifolia (Miers) Sandwith [ P . wrightii A. Gray]. Sharpleaf Ground-cherry

CP: Reported from Las Playas (Simmons 1966) but no specimen located; it occurs in
adjacent Sonora (Felger 2000). OP: Warren 17 Nov 1979.

Physalis lobata Torrey [Quincula lobata (Torrey) Rafinesque], Hairy Ground-cherry
CP: RSF 93-1 01. OP: McDougall 62.

Physalis versicolor Rydberg [P. crassifolia Bentham var. versicolor (Rydberg) Waterfall], Desert
Ground-cherry; tomatillo del desierto

CP: Shreve 5930. OP: Parker 7731. TA: Goodding 4902.

*Solanum americanum Miller [S. nodiflorum Jacquin]. Black Nightshade; chichiquelite; cuvi vupui

OP: Bowers 1329.

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Solanum douglasii Dunal. Douglas Nightshade
OP: Supemaugh 443.

Solanum hindsianum Bentham. Mala mujer, tomatillo espinoso
OP: Lechner 28 Jan 1986.

Solanum xanti A. Gray. Purple Nightshade
OP: Tinkham I Apr 1942.

STERCULIACEAE, see MALVACEAE

TALINACEAE - TALINUM FAMILY

Talinum paniculatum (Jacquin) Gaertner. Pink Baby’s-breath, Jewels of Opar
OP .SR 20061022-14.

TAMARICACEAE - TAMARISK FAMILY

**Tamarix aphylla (Linnaeus) H. Karsten. Athel, Saltcedar; pino salado
CP: RSF 02-15. OP: SR 20 Apr 2001 (ORPI).

*Tamarix chinensis Loureiro [T. ramosissima Ledebour]. Saltcedar, Tamarisk; pino salado', ’onk
’u’us

CP: Simmons 28 Oct 1962 (CAB). OP: Bowers 1391.

URTICACEAE - NETTLE FAMILY

Parietaria hespera B. D. Hinton var. hespera. Desert Pellitory
CP: RSF 93-90. OP: RSF 88-298. TA: RSF 04-55.

ULMACEAE, see CANNABACEAE

VERBENACEAE - VERBENA FAMILY
Aloysia wrightii (A. Gray) A. Heller ex Abrams. Oreganillo
CP: RSF 93-39. OP: Nichol 16 May 1937.

Glandularia bipinnatifida (Nuttall) Nuttall var. bipinnatifida [Verbena ciliata Bentham. V.
bipinnatifida var. ciliata (Bentham) B. L. Turner]

OP: Tinkham 1 Apr 1942.

Glandularia gooddingii (Briquet) Solbrig [Verbena gooddingii Briquet]. Desert Verbena
CP: RSF 93-65. OP: Kearney 10842.

**Lantana camara Linnaeus. Lantana; confiturilla negra
OP: SR 24 Mar 2003.

Tetraclea coulteri A. Gray. Coulter’s Wrinklefruit

CP: RSF 92-741. OP: SR 24 Nov 1997 (ORPI).

Verbena bracteata Lagasca & Rodriguez. Big-bract Verbena
CP: RSF 92-555.

Verbena menthifolia Bentham. Mint Vervain

Attributions of V. halei Small to Arizona, New Mexico, and northwestern Mexico are based
on misidentifications of V. menthifolia (Nesom 2010).

CP: RSF 92-556. OP: RSF 88-454.

Verbena xylopoda (L. M. Perry) G. L. Nesom [V. neomexicana (A. Gray) Small var. xylopoda L. M.
Perry], Arizona Vervain. See Nesom (2010).

OP: Kearney 10846.

VISCACEAE - see SANTALACEAE

ZYGOPHYLLACEAE - CALTROP FAMILY
Fagonia, see Felger (2000).

Fagonia californica Bentham subsp. californica [F. laevis Standley. F. californica subsp. laevis
(Standley) Wiggins]

CP: RSF 92-53. TA: RSF 93-195.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

37

Fagonia californica subsp. longipes (Standley) Felger & C. H. Lowe [F. longipes Standley]

CP: RSF 92-572. OP: McDougall 51.

Fagonia pachyacantha Rydberg [F. californica var. glutinosa Pringle ex Vail]

CP: RSF 93-169. TA: Lindquist 25 Mar 1983.

Kailstroemia californica (S. Watson) Vail. Baihurm, mal de ojo
CP: RSF 92-674. OP: Bowers 1529. TA: RSF 04-47.

Kailstroemia grandi flora Torrey ex A. Gray. Summer poppy; baiburin, mal de ojo
CP: RSF 92-686. OP: Bowers 1528.

Larrea tridentata (de Candolle) Coville [L. divaricata Cavanilles subsp. tridentata (de Candolle)
Felger & C. H. Lowe]. Creosotebush; hediondilla, gobernadora; segai, segoi
CP: Yeatts 3657. OP: Nichol 26 Mar 1939. TA: RSF 04-1 7.

*Tribulus terrestris Linnaeus. Puncture Vine, Caltrop, Goathead; torito, toboso
CP: Simmons 7.

MONOCOTS-MONOCOYLEDONEAE

AGAVACEAE, see ASPARAGACEAE

ALLIACEAE, see AMARYLLIDACEAE

AMARYLLIDACEAE - AMARYLLIS FAMILY
Allium macropetalum Rydberg. Desert Onion; cebollin

CP: RSF 93-117. OP: Phillips 83-87 (ASU, ORPI).

Zephyranthes longifolia Hemsley. Plains Rain-lily

OP: Bowers 1802.

ARECACEAE - PALM FAMILY

**Washingtonia fllifera (Linden ex Andre) H. Wendland ex de Bary. California Fan Palm
OP: RSF 88-258 (ORPI, removed).

ASPARAGACEAE - ASPARAGUS FAMILY (includes Agavaceae, Nolinaceae or Ruscaceae, and
Themidaceae)

Agave x ajoenesis W. C. Hodgson. Ajo Mountain Agave
OP: Hodgson 4478 (holotype, DES).

Agave deserti Engelmann subsp. simplex Gentry. Desert Agave; lechuguilla, mezcaf 'a'ud
CP: Gentry 20590 . OP: Hodgson 5766 (ASU, ORPI). TA: RSF 90-19.

Agave schottii Engelmann var. schottii. Shin-dagger; amolillo; utko je:j
OP: Baker 7758 (ASU).

Dichelostemma capitatum (Bentham) Alfred Wood subsp. pauciflorum (Torrey) Keator [D.
pulchellum (Salisbury) A. Heller var. pauciflorum (Torrey) Hoover. Brodiaea capitata Bentham var.
pauc (flora Torrey]. Blue Dicks, Wild Hyacinth; coberia, cobena, ajo bianco-, ha:d
CP: Crawford 3 (ASC). OP: RSF 05-254.

Hesperocallis undulata A. Gray. Ajo Lily, Desert Lily; ajo silvestre; a:sos
CP: Reeves 6783 (RSA). OP: McDougall 37. TA: RSF 05-119.

Nolina bigelovii (Torrey) S. Watson. Desert Tree-beargrass
CP: RSF 92-601. TA: Shreve 6233.

Nolina microcarpa S. Watson. Beargrass; sacahuiste\ moho
OP: Bowers 1296 (ORPI).

Triteleiopsis palmeri (S. Watson) Hoover [ Brodiaea palmeri S. Watson], Blue Sand-lily
CP: RSF 93-384. T A : TRY 27 Mar 1983.

Yucca baccata Torrey var. brevifolia L. D. Benson [Y. arizonica McKelvey], Banana Yucca; ddtil;
howij

OP: RSF 05-278.

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CANOTIA Vol. 8

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COMMELINACEAE - SPIDERWORT FAMILY
Commelina erecta Linnaeus. White-mouth Dayflower
OP: Wilson 191.

CYPERACEAE - SEDGE FAMILY

Cyperus esculentus Linnaeus var. leptostachyus Boeckeler. Yellow Nutgrass; coquillo amarillo,
cebollin\ vasai su:vl

CP: RSF 01-575.

Cyperus laevigatus Linnaeus. Flat Sedge
OP: McDougall 34.

Cyperus mutisii (Kunth) Andersson [Mariscus mutisii Kunth. C. asper (Liebmann) O’Neill]

OP: Goodding 306-41 .

Cyperus squarrosus Linnaeus [C. aristatus Rottboll. Mariscus squarrosus (Linnaeus) C. B. Clarke].
Dwarf Sedge

CP: Monson 9. OP: Clark 10990 (ORPI). TA: RSF 04-41.

Eleoeharis coloradoensis (Britton) Gilly. Dwarf Spikerush
OP: Peebles 14565.

Eleoeharis flavescens (Poiret) Urban var. flavescens. Yellow Spikerush
OP: Darrow 2403.

Eleoeharis rostellata (Torrey) Torrey. Traveling Spikerush
OP: McDougall 35.

Schoenoplectus americanus (Persoon) Volkart ex Schinz & R. Keller [Scirpus americanus Persoon.
S. olneyi A. Gray]. Bulrush; tule \ va:k
OP: Peebles 14564.

HYDROCHARITACEAE - WATERWEED FAMILY (includes Najadaceae)

Najas marina Linnaeus. Holly-leaf Water-nymph; sargazo
OP: Pinkava 2363 (ASU).

IRIDACEAE - IRIS FAMILY
Sisyrinchium demissum Greene. Blue-eyed Grass
OP: SR 23 Apr 1999 (ORPI).

JUNCACEAE - RUSH FAMILY
Juneus bufonius Linnaeus. Toad Rush

OP: Clark 11501 (ORPI, extirpated).

Juneus cooperi Engelmann. Cooper’s Rush
OP: Bowers 1309.

Juneus mexicanus Willdenow ex Roemer & Schultes f. [J. articus Willdenow var. mexicartus
(Willdenow ex Roemer & Schultes f.) Balsev]. Mexican Rush, Wire Rush
OP: Reichhardt 69.

LILIACEAE - LILY FAMILY

Calochortus kennedyi Porter var. kennedyi. Desert Mariposa Lily; ajo amarillo. cobena amarilla\
ha:dkos

OP: RSF 05-176.

NOLINACEAE, see ASPARAGACEAE
POACEAE (GRAMINEAE) - GRASS FAMILY

Aristida adseensionis Linnaeus. Six-weeks Three-awn; zacate tres barbas
CP: RSF 92-681. OP: Bowers 1034. TA: RSF 04-69.

Aristida californica Thurber var. californica. California Three-awn; tres barbas de California
CP: Reeder 6835.

Aristida californica var. glabrata Vasey. Smooth California Three-awn; tres barbas de California
OP: Reeder 8218.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

39

Aristida purpurea Nuttall var. nealieyi (Vasey) Allred [A. glauca (Nees) Walpers]. Purple Three-

awn; tres barbas

CP: Reeder 7591. OP: RSF 90-49. TA: Vorhies 16 Apr 1924.

Aristida purpurea var. parishii (Hitchcock) Allred [A. parishii Hitchcock]. Parish Three-awn
CP: Reeder 6830. OP: RSF 92-102.

Aristida ternipes Cavanilles var. gentilis (Henrard) Allred [A. gentilis Henrard. A. hamulosa
Henrard. A. ternipes var. hamulosa (Henrard) Trent]. Poverty Three-awn; zacate araha de tres
barbas

CP: RSF 92-710. OP: SR 2003-306 (ORPI).

Aristida ternipes var. ternipes. Spider Grass; zacate araha
CP: RSF 9 2-5 79b. OP: Bowers 917.

*Avena fatua Linnaeus. Wild Oats; avena silvestre, avena cimarrona, avena loca\ 'agspi mudat-kam,
ko:ksam

OP: SR 14 Apr 1998.

Bothriochloa harbinodis (Lagasca) Herter [ Andropogon barbinodis Lagasca]. Cane Bluestem;
zacate popotillo

CP: Goodding 127 . OP: Bowers 911. TA: Tinajas Altas Canyon, 19 Mar 1998, RSF,
observation.

Bouteloua aristidoides (Kunth) Grisebach [6. aristidoides var. arizonica M. E. Jones]. Six-weeks
Needle Grama; aceitilla, navajita aguja

CP: RSF 92-667. OP: RSF 86-280. TA: RSF 04-31.

Bouteloua barbata Lagasca var. barbata. Six-weeks Grama; navajita, zacate liebrero ; s-cuk
mudadt-kam

CP: RSF 92-683. OP: RSF 86-279. TA: RSF 04-32.

Bouteloua barbata var. rothroekii (Vasey) Gould [B. rothrockii Vasey]. Rothrock Grama; zacate
liebrero

OP: Bowers 1708.

Bouteloua curtipendula (Michaux) Torrey. Side-oats Grama; banderilla ; da:pk vasai
OP: Nichol 4 May 1939.

Bouteloua repens (Kunth) Scribner & Merrill [B. fdiformis (E. Fournier) Griffiths], Slender Grama;
navajita delgada

OP: Van Devender 85-113.

Bouteloua trifida Thurber. Red Grama; navajita china
CP: RSF 92-648. OP: SR 20050320-46.

Bromus carinatus Hooker & Arnott [£. arizonicus (Shear) Stebbins. B. carinatus var. arizonicus
Shear. B. marginatus Nees ex Steudel]. California Brome
CP: RSF 92-2. OP: Bowers 1209.

*Bromus madritensis Linnaeus subsp. rubens (Linnaeus) Husnot [B. rubens Linnaeus], Red Brome;
bromo rojo

CP: RSF 93-324. OP: SR 12 May 2003. TA: RSF 02-8.

**Bromus tectorum Linnaeus. Cheat Grass, Downy Chess
OP: RSF 86-133.

Cenchrus, see Chemisquy et al. (2010) and Columbus and Smith (2010).

*Cenchrus ciliaris Linnaeus [Pennisetum ciliare (Linnaeus) Link]. Buffelgrass; zacate buffel

CP: RSF 92-727. OP: RSF 86-448. TA: Frontera Canyon, 18 Mar 1998, RSF, observation.

* Cenchrus echinatus Linnaeus. Southern Sandbur; guachapori
OP: RSF 87-319.

*Cenchrus setaceus (Forsskal) Morrone [Pennisetum setaceum (Forsskal) Chiovenda. P. ruppelii
Steudel], Fountain Grass

CP: RSF 92-646. OP: RSF 87-333.

*Cenchrus spinifex Cavanilles, 1799 [C. incertus M. A. Curtis, 1835. C. pauciflorus Bentham,

1844], Common Sandbur; huizapori, guachapori
CP: Reeder 6836.

Chloris virgata Swartz. Feather Fingergrass; zacate lagunero
CP: RSF 92-550. OP: RSF 87-315.

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CANOTIA Vol. 8

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Cottea pappophoroides Kunth. Cotta Grass; zacate papo
CP: RSF 02-38. OP: RSF 02-73.

*Cynodon dactylon (Linnaeus) Persoon var. dactylon. Bermuda Grass; zacate bermuda, zacate
ingles ; ki: weco vasai, ’a’ai hihimdam vasai

CP: Eagle Tank, 13 Jun 1992, RSF, observation. OP: RSF 87-320.

*Dactyloctenium aegyptium (Linnaeus) Willdenow. Crowfoot Grass; zacate de cuervo
OP: RSF 87-289.

Dasyochloa pulchella (Kunth) Willdenow ex Rydberg [Triodia pulchella Kunth. Tridens pulchellus
(Kunth) Hitchcock. Erioneuron pulchellum (Kunth) Tateoka]. Fluff Grass; zacate borreguero
CP: RSF 92-641. OP: Bowers 921. TA: RSF 04-11.

Digitaria californica (Bentham) Henrard var. californica [Trichachne californica (Bentham)

Chase], Arizona Cottontop; zacate punta blanca

CP: Goodding 924G. OP: Bowers 923. TA: RSF 10-185.

Distichlis spicata (Linnaeus) Greene [ D . stricta (Torrey) Rydberg], Salt Grass; zacate salado\ ’onk,
vasai

OP: Nichol 28 Apr 1939.

*Echinochioa colona (Linnaeus) Link. Jungle Rice, Leopard Grass; zacate pinto, zacate tigre, zacate
rayado

CP: RSF 92-717. OP: TRY 9 Sep 1984.

*Echinochloa crus-galli (Linnaeus) P. Beauvois. Barnyard Grass; zacate de agua, zacate de corral
CP: RSF 92-718 (ASU).

Elymus elymoides (Rafinesque) Swezey [Sitanion hvstrix (Nuttall) J. G. Smith], Bottlebrush
Squirreltail

OP: Bowers 1302.

*Enneapogon cenchroides (Lichtenstein ex Roemer & Schultes) C. E. Hubbard. Soft-feather Pappus
Grass

OP: SR 20061022-1.

Enneapogon desvauxii P. Beauvois. Spike Pappus Grass, Nine-awn Pappus Grass; zacate lobero
CP: Monson 24 May 1957. OP: SR 20060927-1.

**Eragrostis barrelieri Daveau. Mediterranean Love Grass

OP: RSF 87-276.

*Eragrostis cilianensis (Allioni) Vignolo-Lutati ex Janchen. Stink Grass; zacate apestoso

Perry and McNeill (1986) confirm that the Allioni is given credit for the basionym and that
Janchen is the publishing author, thus supporting the authorship shown here for this species (John
Reeder, personal communication 2004 and Kelly Allred, personal communication 23 December
2011).

CP: RSF 92-567. OP: Van Devender 84-447. TA: RSF 04-21.

Eragrostis intermedia Hitchcock. Plains Love Grass
OP: Nichol 4 May 1939.

*Eragrostis lehmanniana Nees. Lehmann Love Grass
CP: RSF 92-647. OP: SR 9 May 1996.

Eragrostis peetinacea (Michaux) Nees [ E . diffusa Buckley. E. tephrosanthes Schultes. E. arida
Hitchcock. E. peetinacea var. miserrima (E. Fournier) Reeder], Carolina Love Grass
CP: RSF 01-560.

*Eriochloa acuminata (J. Presl) Kunth var. acuminata [ E . lemmonii Vasey & Scribner var. gracilis
(E. Fournier) Gould]. Southwestern Cup Grass; zacate taza
OP: RSF 86-266.

Eriochloa aristata Vasey var. aristata. Bearded Cup Grass; zacate taza
CP: RSF 92-719. OP: RSF 02-73a (ASU).

Festuca microstachys Nuttall [Vulpia microstachys (Nuttall) Munro]. Small Fescue. For the two
Festuca, see Columbus and Smith (2010) and Jepson Flora Project (2012).

OP: RSF 03-426.

Festuca octoflora Walter [ Vulpia octoflora (Walter) Rydberg], Sixweeks Fescue, Eight-flowered
Fescue

CP: RSF 93-89. OP: Nichol 14 Mar 1939. TA: TRY 10 Mar 1980.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

41

Heteropogon contortus (Linnaeus) P. Beauvois ex Roemer & Schultes [Andropogon contortus
Linnaeus]. Tanglehead; zacate Colorado

CP: RSF 92-651. OP: Reeder 8911. TA: TRY 25 Mar 1983.

Hilaria belangeri (Steudel) Nash [H. belangeri var. longifolia (Vasey) Hitchcock]. Curly Mesquite;
zacate chino

OP: RSF 05-179.

Hilaria rigida (Thurber) Bentham ex Scribner [Pleuraphis rigida Thurber], Big Galleta; tobosa
CP: RSF 93-405. OP: TR V 87-8. TA: RSF 04-46.

Hopia obtusa (Kunth) Zuloaga & Morrone [ Panicum obtusion Kunth], Vine Mesquite; zacate de
gma

OP: Bowers 1796.

*Hordeum rnurinum Linnaeus subsp. giaucum (Steudel) Tzvelev [R stebbinsii Covas. H.
leporinum Link subsp. giaucum (Steudel) T. A. Booth & A, J. Richards]. Wild Barley; cebadilla
silvestre

CP: RSF 93-72. OP: Nichol 28 Apr 1939.

Leptochloa dubia (Kunth) Nees [ Diplachne dubia (Kunth) Scribner], Green Sprangletop
OP: Gould 4690.

Leptochloa fusca (Linnaeus) Kunth subsp. uninervia (J. Presl) N. W. Snow [L. uninervia (J. Presl)
Hitchcock & Chase. Diplachne uninervia (J. Presl) Parodi], Mexican Sprangletop
OP: SR 2003-442.

Leptochloa panicea (Retzius) Ohwi subsp. brachiata (Steudel) N. W. Snow [L. brachiata Steudel.
L.filiformis (Persoon) P. Beauvois], Red Sprangletop; deparramo rojo
CP: RSF 92-564. OP: Bowers 1518.

Leptochloa viscida (Scribner) Beal. Sticky' Sprangletop
CP: Mom on 7.

Melica frutescens Scribner. Woody Melic
OP: Goodding 302-41 .

Muhlenbergia alopecuroides (Grisebach) P. M. Peterson & Columbus [ Lycurus alopecuroides
Grisebach. L. setosus (Nuttall) C. Reeder], Bristly Wolftail. For placement of Lycurus in
Muhlenbergia , see Peterson et al. (2010).

OP: Wirt 29 Oct 1989 (ORPI).

Muhlenbergia appressa C. O. Goodding. Devil’s Canyon Muhly
OP: SR 2003-339.

Muhlenbergia dumosa Scribner ex Vasey. Bamboo Muhly; carricillo, otatillo
OP: Reeder 8909.

Muhlenbergia emersleyi Vasey [M gooddingii Soderstrom], Bullgrass, Bull Muhly; zacate de toro
OP: Goodding M-3 73.

Muhlenbergia fragilis Swallen. Delicate Muhly

OP: SR 20061022-2.

Muhlenbergia microsperma (de Candolle) Kunth. Little-seed Muhly; liendrilla
CP: RSF 92-569. OP: Goodding 480-45. TA: RSF 04-51 .

Muhlenbergia porteri Scribner ex Beal. Bush Muhly; zacate aparejo; ku:kpadag
CP: RSF 93-45. RSF. OP: Mulroy 6 Oct 1969.

Muhlenbergia rigens (Bentham) Hitchcock. Deer Grass; zacate del venado
OP: Goodding 479-45.

Muhlenbergia tenuifolia (Kunth) Kunth [M monticola Buckley], Slender Muhly
OP: Reeder 8910.

Panicum alatum Zuloaga & Morrone var. alatum
CP: RSF 01-555.

*Panicum antidotale Retzius. Giant Panic Grass, Blue Panic Grass; panizo azul
OP: RSF 87-329.

Panicum hallii Vasey var. hallii [P. lepidulum Hitchcock & Chase], Hall’s Witch Grass
OP: Goodding 3 1 2-4 1 .

Panicum hirticaule I. Presl var. hirticaule [P. capillare Linnaeus var. hirticaule (J. Presl) Gould].
Mexican Panic Grass

CP: RSF 92-657. OP: RSF 88-428.

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CANOTIA Vol. 8

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Pappophorum vaginatum Buckley. Pappus Grass
OP: Albee Apr 1937.

*Phalaris caroliniana Walter. Carolina Canary Grass; alpistillo
CP: Harlan 494.

*Phalaris minor Retzius. Little-seed Canary Grass; alpistillo ; ba:bkam
CP: RSF 92-569. OP: Bowers 1 722.

Phragmites australis (Cavanilles) Trinius ex Steudel subsp. berlandieri (E. Fournier) Saltonstall &
Hauber [P. communis Trinius]. Common Reed, Reed Grass; carrizo; va:pk
OP: RSF 86-214.

**Poa annua Linnaeus. Annual Blue Grass, Winter Grass; pastito de invierno
OP: Darrow 2405.

Poa bigelovii Vasey & Scribner. Bigelow Blue Grass; zacate azul anual
CP: RSF 93-54. OP: Bowers 1250.

*Polypogon monspeliensis (Linnaeus) Desfontaines. Rabbitfoot Grass, Annual Beard Grass; zacate
cola de zorra

CP: Henry 05/08/1996 (CAB). OP: McDougall 91.

*PoIypogon viridis (Gouan) BreistrofTer [Agrostis semiverticillata (Forsskal) C. Christensen.
Polypogon semiverticillata (Forsskal) Hylander], Water Bent Grass, Water Beard Grass
OP: Bowers 1311 .

*Schismus arabicus Nees [5. barbatus subsp. arabicits (Nees) Marie & Weiller], Arabian Grass
CP: RSF 93-35. OP: Bowers 1109. TA: TRV 5 Mar 1983.

*Schismus barbatus (Linnaeus) Thellung. Mediterranean Grass
OP: RSF 88-301 .

Setaria grisebachii E. Fournier. Grisebach’s Bristle Grass; zacate cola de zorra
OP: Goodding 474-45.

Setaria liebmannii E. Fournier. Liebmann’s Bristle Grass; zacate cola de zorra
OP: Goodding 25 Sep 1943.

Setaria macrostachya Kunth (this species complex includes S. leucopila (Scribner & Merrill) K.
Schumann). Plains Bristle Grass; zacate tempranero
CP: RSF 92-570. OP: Bowers 1294.

*Sorghum halepense (Linnaeus) Persoon. Johnson Grass; zacate johnson
CP: Simmons 30 Oct 1962 (CAB). OP: RSF 87-325.

Sporobolus airoides (Torrey) Torrey. Alkali Sacaton; zacaton alcalino
CP: RSF 92-604. OP: Bowers 1805.

Sporobolus cryptandrus (Torrey) A. Gray. Sand Dropseed; zacate arenero
CP: RSF 92-64 3d. OP: RSF 90-474. TA: Goodding 599.

Sporobolus flexuosus (Thurber ex Vasey) Rydberg [5. ctyptandrus var. flexuosus (Thurber ex
Vasey) Thurber]. Mesa Dropseed
OP \Wirt 8 Sep 1990.

Sporobolus pyramidatus (Lamarck) Hitchcock [5. patens Swallen. S. pulvinatus Swallen], Whorled
Dropseed

OP : RSF 88-420.

Stipa speciosa Trinius & Ruprecht var. speciosa [Achnatherum speciosum (Trinius & Ruprecht)
Barkworth. Jarava speciosa (Trinius & Ruprecht) Penailillo], Desert Needle Grass. See Columbus
and Smith (2010) and Jepson Flora Project (2012).

OP: RSF 90-529. TA: TRV 26 Mar 1983.

Trichloris crinita (Lagasca) Parodi [71 mendocina (Philippe) Kurtz. Chloris crinita Lagasca].
Feather Finger Grass; zacate escoba
OP: RSF 87-280.

Tridens eragrostoides (Vasey & Scribner) Nash. Lovegrass Tridens
OP: Goodding 295-41 .

Tridens muticus (Torrey) Nash var. muticus. Slim Tridens

CP: Whipple 3955. OP: Bowers 1559. TA: RSF 04-75.

**Triticum aestivum Linnaeus. Wheat; trigo; pilkan
OP: RSF 05-161.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

43

Urochloa arizonica (Scribner & Merrill) Morrone & Zuloaga [Panicum arizonicum Scribner &
Merrill. Brachiaria arizonica (Scribner & Merrill) S. T. Blake]. Arizona Signal Grass; piojillo de
arizona

CP: Bowers 1532. OP: Bowers 1519.

POTAMOGETONACEAE - POND WEED FAMILY

Stuckenia pectinata (Linnaeus) Borner [ Potamogeton pectinatus Linnaeus. Coleogeton pectinatus
(Linnaeus) Les & R. R. Haynes]. Sago Pondweed
OP: RSF 86-270.

RUPPIACEAE - DITCH-GRASS FAMILY
Ruppia cirrhosa (Petagna) Grande. Ditch-grass
OP: RSF 86-222.

RUSCACEAE, see ASPARAGACEAE
TPIEMIDACEAE, see ASPARAGACEAE
TYPHACEAE - CATTAIL FAMILY

Typha domingensis Persoon. Southern Cattail; tule; 'uduvhag

CP: Cabeza Prieta Tanks, 12 Jun 1992, RSF, observation (also see Simmons 1966). OP:
RSF 89-235. TA: Wilder 08-383.

ZANNICHELLIACEAE - HORNED-PONDWEED FAMILY
Zannichellia palustris Linnaeus. Homed-pondweed
CP: Harlan 471. OP: Peebles 14566.

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CANOTIA Vol. 8

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Table 1. Numbers of total taxa, non-native species, genera, families, and areas for Organ Pipe Cactus
National Monument, Cabeza Prieta National Wildlife Refuge, Tinajas Altas Region, and the six
protected areas of the total Dry Borders area. Taxa numbers represent species plus subspecies or
varieties, and hybrids.

OP, CP, TA
combined

Organ

Pipe

(OP)

Cabeza

Prieta

(CP)

Tinajas

Altas

(TA)

Six Dry
Borders

Areas

Taxa

732

655

423

228

862

Non-native species

78

72

37

12

91

Genera

420

395

266

164

448

Families

94

93

63

47

107

Acres

1,270,699

330,689

860,010

80,000

7,515,221

Hectares

514,234

133,825

348,034

32,375

3,041,302

Checklist of Plants of Southwestern Arizona. Figure 1. Organ Pipe Cactus National Monument,
Cabeza Prieta National Wildlife Refuge, Tinajas Altas Region (the flora area), and adjacent areas.
Drafted by H. Jeff Boyd.

2012

CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

45

Checklist of Plants of Southwestern Arizona. Figure 2. Cabeza Prieta National Wildlife Refuge.
Photographer David Taylor, looking northward above O’Neill’s Grave, 26 Mar 2010. Xeroriparian
vegetation along small arroyos including Hilaria rigida, Parkinsonia floridci ,
and Prosopis glandulosa. Photo by RSF.

Checklist of Plants of Southwestern Arizona. Figure 3. Cabeza Prieta National Wildlife Refuge.
Pinta Sands, 27 Mar 2010. Spring annuals including Abronia vil/osa , Brassica tournefortii in
foreground, Cryptantha angustifolia , and Oenothera deltoides. Photo by Joan Scott.

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CANOTIA Vol. 8

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Checklist of Plants of Southwestern Arizona. Figure 4. Organ Pipe Cactus National Monument.
Southeast- facing slope in the southern Diablo Mountains with saguaros ( Carnegiea gigantea) and
organ pipes (, Stenocereus thurberi). 12 Apr 2006. Photo by SR.

Checklist of Plants of Southwestern Arizona. Figure 5. Organ Pipe Cactus National Monument.
Quitobaquito pond including Baccharis salicifolia in foreground, and Populus fremontii,
Schoenoplectus americanus, Typha domingensis in background. 29 Sep 2011. Photo by SMB.

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CHECKLIST OF PLANTS OF SOUTHWESTERN ARIZONA

47

Checklist of Plants of Southwestern Arizona. Figure 6. Tinajas Altas Mountains. Desert bighorn
ram and including Bursera microphylla and Hyptis cilbida. 16 Apr 2011. Photo by Jim Malusa.

Checklist of Plants of Southwestern Arizona. Figure 7. Tinajas Altas Mountains, Surveyors Tank
Canyon, looking eastward. Canyon bottom vegetation including Encelia farinosa , Olneya
tesota , Pros o p is glandulosa, and Senegalia greggii. 29 Mar 2010. Photo by RSF.

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ACKNOWLEDGEMENTS

Over the years many different friends and colleagues have provided information and
assistance, and/or accompanied us in the field. In this regard we thank Kelly W. Allred, Daniel F.
Austin, Tina J. Ayers, Marc A. Baker, Anne Christine Barber, Brad Boyle, Bill Broyles, J. Travis
Columbus, Mark A. Dimmitt, Ellen Dorn, Luke Taylor Evans, Mark Fishbein, Judy Ann Gibson,
Edward Erik Gilbert, W. Eugene Hall, Stephen F. Hale, Gayle Harrison Hartmann, James L. Heard,
Ann Lane Hedlund, James Henrickson, Wendy Caye Hodgson, Sarah Hunkins, Philip D. Jenkins,
Richard Burton Laugharn, the late Cynthia A. Lindquist, Elizabeth Makings, Jim Malusa, Curtis
McCasland, Michelle McMahon, Guy Nesom, J. Mark Porter, Adrianne G. Rankin, Jon P. Rebman,
the late Charlotte Reeder and John Reeder, Frank W. Reichenbacher, Karen Reichhardt, Glenn Rink,
Rusty Russell, Andrew Salywon, Andrew C. Sanders, Silke Schneider, Joan Evelyn Scott, Richard
Spellenberg, Victor Steinmann, David Jonathan Taylor, Timothy Tibbitts, Larry Toolin, Gordon C.
Tucker, Raymond M. Turner, Sula Vanderplank, Benjamin T. Wilder, Michael F. Wilson, and
George Yatskievych. We thank H. Jeff Boyd for drafting the map. The authors thank Lacrecia
Johnson of the United States Fish and Wildlife Service for generous assistance in funding for the
publication of this manuscript. We thank two anonymous reviews whose promptings greatly
improved the manuscript. Our special thanks go to Leslie R. Landrum for substantive suggestions and
editing and to Scott T. Bates for expert production editing. Richard Felger writes: I especially
appreciate Bill Broyles introducing me to this great place and for good times in the field. Much of my
work has been supported by the Wallace Research Foundation.

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