CATALOGUE TERTIARY MOLLUSCA IV THE DEPARTMENT OF GEOLOGY. PART I. AUSTRALASIA. ■^^' CATALOGUE TERTIARY MOLLUSCA IN THE DEPAETMENT OF GEOLOGY BRITISH MUSEUM (NATURAL HISTOHY). PART I. THE AUSTEALASIAxX TERTIAEY MOLLUSCA. GEOEGE F. HARRIS, F.G.S., MEMB. SOC. GEOL. PKANCE ; MEMB. SOC. BELGE GEOL. TKEAS. MALACOL. SOC. LOND. PRINTED BY ORDER OF THE TRUSTEES. SOLD BY LONGMANS AND CO., 39, PATERNOSTER ROW. B. QUARITCH, 15, PICCADILLY. DULAD AND CO., 37, SOHO SQUARE, W. KEGAN PAUL AND CO., CHARING CROSS ROAD, W.C. AND AT THE BRITISH MUSEUM (NATURAL HISTORY), CROMWELL ROAD, S.W. 1897. {AN rights reserved.) HERTFORD : PRINTED BY STEPHEN AUSTIN AND SONS. PEEFACE. The Catalogue of the Cephalopodous Mollusca by Dr. A. H. Foord, commenced in 1888 (two parts of which are issued, the third part being now in the press), together with the Systematic List of British Oligocene and Eocene Mollusca by Mr. E,. Bullen Newton, in 1891, are the only volumes already published on the fossil Mollusca in this Department. The present work by Mr. Gr. F. Harris, on Tertiary Mollusca, commences a new Catalogue ; the first part being devoted to descriptions and figures of the shells of Australasia (exclusive of the Cephalopoda), to be followed by other geographical series in the collection, of equal interest. Although thirteen volumes on Fossil Yertebrata, eight volumes on Fossil Invertebrata, and three upon Fossil Plants, have already been published by the Trustees, the larger part of the Mollusca and Bryozoa, and the whole of the Brachiopoda, Annelida, Arthropoda, Echinoderma, and Ccelentera, remain to be recorded, as well as the greater portion of the fossil plants, so that probably not less than thirty volumes will be needed to contain. even a brief account of these very varied and extensive collections. HENRY WOODWARD. British Museum (Natural History), Dki'artment op Geology. March 25, 1897. INTRODUCTION. The Tertiary MoUusca of Australasia present many features of special interest, both to geologists and zoologists. The remarkable abundance and perfect state of preservation of the Gastekopoda, especially of Victoria, enable us to shed much light on certain questions relating to phylogeny, which cannot fail to assist the zoologist as indicating the oiigin of many of the principal groups of that division of the Mollusca, and as imparting some idea of the past history of the modern molluscan fauna of Australasia generally. The larval shells, especially, have proved to be of great interest, and I venture to think that many of the details concerning them as recorded in this Catalogue will prove serviceable, not only from the phylogenetic standpoint, and in regard to the early history of the individual, but as a contributii)u to the systematic classification of the Gasteropoda. The phenomena of growth in this division of the mollusca have never been studied, except in regard to restricted groups; and I have occasionally criticized modern methods that give undue importance (as it would seem) to the number and disposition of plications on the columella, the absence or presence of liriE within the outer margin of the aperture, the relative length of the anterior canal when present, and modifications in detail of external sculpture. These and kindred phenomena aie frequently utilized by the systematist for purposes of specific and even of generic classification, whereas in many instances a careful study of ontogeny would show that they were mere peculiarities either of the individual or as characterizing a distinct stage of growth. I do not intend to imply that the features of growth just mentioned are never of importance for purposes of classification ; but until each case is studied on its own merits, in other words until the ontogeny of each has been carefully worked out, it is impossible to say INTRODUCTION. ■whether any particular phenomenon observed is of value to the systematist, or not. A phenomenon may be of the highest importance in the classification of certain genera, and be of little value from that point of view in reference to others. The study of the development of the Gasteropoda would show better than anything else what value is to be ascribed to any particular structural feature in the shell, in that sense ; but little has hitherto been attempted in that direction. An exhaustive study of auxology cannot, obviously, be chronicled in a Catalogue like the present; but I have seized this opportunity of selecting the most promising species and genera of Australasian Tertiary Gasteropoda from' the material in the Museum, and, as far as is consistent with the plan of the book, have detailed certain structures of general interest which may hereafter prove of service to workers in elaborating the genesis of the modern Gasteropod fauna of the antipodes. To the geologist, also, particulars concerning the growth of these Tertiary shells are not without their significance. It is very difficult for paleontologists, by the methods now commonly adopted in classifying the Mollusca, to say how much of a variation from a normal type shall constitute a species : the personal equation is always with us. And seeing that Tertiary beds are commonly classified according to the relative percentage of extinct species of mollusca found in them, and that for the reason just given it is almost impossible to arrive satisfactorily at such percentages, it seems to me that, rightly understood, the phylogeny of that division of the animal kingdom is a broader and surer basis for classification of the various horizons, and might be made to run pari passu with the Lyellian method. Its most useful application, no doubt, would be in reference to homotaxis. This suggestion is to a large extent prompted by the great difficulty which geologists experience in working out the synchrony of the Tertiary beds of Australia, to this day a source of much controvei'sy between them. As instances of the application of phylogeny in matters of this kind let me refer to pp. 17, 18, where in reference to the so-called Pteropoda it will be noted that the assemblage of genera found in certain rocks classified by competent authorities as Eocene is not such as might have been expected in rocks of that age, when the known history and origin of the Pteropoda are taken into INTRODUCTION. IX consideration. As a further illustration T may refer to p. 9G, "where, after giving some account of the larval shells of the VoLUTiDJK (p. 92 et sqq.) and the light they throw on the origin and evolution of that family, I state that tlie development of these larval Volutes in the majority of instances in specimens found at the localities mentioned .... is much more advanced than one would have expected to find in beds as old as the Eocene — that is, on comparison with what is found in the true Eocene in other parts of the world. And it would not be difficult to give other evidence of the same nature. No doubt some families make more rapid strides in one area than in another ; but it is surprising to find in the Gasteropoda how uniform over large areas the rate of progression has been in regard to many of the widely distributed families. There is, for example, the genus that has survived from early Mesozoic times, with hut little modification in the later stages of its journey ; it has had its day, and has settled down as a more or less fixed form. Such a genus is of little use for homotaxial purposes, though eminently interesting from a purely phylogenetic point of view. What we want to study specially for the purposes of Tertiary homotaxis are those families, quite numerous enough, the chief members of which, having parted from the main stock in late Jurassic or Cretaceous times, arrive in the Eocene, with strong tendencies to variation, and, at the same time, are rapid travellers and become ubic|uitous, as is the case with many prominent genera of the Australasian Tertiary Gasteropoda. But this is not the place to enlarge on the value of phylogeny to the geologist, and the subject is introduced here only as in some measure the outcome of a study of the Tertiary MoUusca from the Antipodes now catalogued. In the foregoing observations special stress has been laid on the study of Gasteeopoda, but it must not be understood that the fossil Lamellibeanchiata are one whit inferior in point of interest from the phylogenetic or the geological stand- point. The want of sufficient suitable material in the Museum for the study of the auxology of the Australasian Tertiary LAMELLiBEAXcniATA, is the sole cause for the omission. It should, however, be added that in many specimens the prodissoconch is excellently preserved. The n.imes employed to designate stages of growth of the individual, and the sense in which they are applied in this work, X INTRODUCTION. are, as nearly as may be, those of Professor Alpheus Hyatt, ^ as amended by Messrs. S. S. Euckman and F. A. Uather.'' The terminology alluded to is as follows : — 1. Embryonic. "This stage includes all individual history from the ovum up to the time when the organism can be referred definitely to its class." In the fossil state we can only have left for investigation the hard part of the organism, which in the case of the Gasteropoda is here called the Protoconch, and in the Lamellibranchiata the Pyo(iissoeo7i'ch. 2. Brephic. This stage immediately succeeds the Embryonic, and it is generally undei-stood that during it no specific characters can be distinguished ; but this point will be discussed presently. 3. Neanic. "Luring this stage specific characters and all other morphological features present in the adult, appear and undergo development." 4. Eplielic. That stage denoting "the period of full develop- ment of the individual, when all specific characters are clearly recognizable." 5. Gerontie. "During this stage changes take place which are due to gradual failure of powers." It is subdivided into : " a, Catahatic stage, in which the individual loses its ephebic characters ; and h, Hypodrophic stage, in which the continued loss of characters causes pronounced reversion." In view of the fact that so little is known concerning the phylogeny of the Gasteropoda, I have found it somewhat difficult in certain cases to apply this terminology satisfactorily. Por instance, in the case of shells that are quite smooth it is hard to say where the protoconch ends and the brephic stage begins ; but in very many instances a characteristic varix is present, denoting the termination of the former. The brephic stage, as a rule, in reference to these fossils, was a short one, the growth having been very vigorous immediately on the animal's escape from the capsule, and conchological specific characters consequently make their appearance at a very early stage. Some of the more prominent results obtained from this study of Australasian shells may now be refisrred to. Proc. Boston Soc. Nat. Hist. vol. xxiii. 1888, p. 396. Zoologiscliea Auzeiger, Nos. 405-6, 1892, pp. 420, 429. INTKODUCTIOX. XI It ■will be noticed (p. 7) that I have retained Adaon in its usual place, amongst the OpismoBRANcniAXA ; but it must be confessed that this is done with some misgiving. It is generally concluded,' judging from the structure of its shell and its persistent operculum, that that genus should be placed amongst the lower Opisthobranchs; but it should be observed that the earlier portion of the protoconch is distinctly sinistral, and has all the essential characteristics of the Pyramidellid^. On that ground alone one might feel inclined to classify Aetceon, and possibly some of its associated genera, amongst the Peosobranchiata. We should be strongly fortified in so doing by the important discovery by Bouvier,- in a careful anatomical investigation, that Aetceon has a twist in its visceral nerve-loop, and is, consequently, streptoneurous. In view of what is known of the general organization of the genus, even in spite of the evidence afforded by the protoconch and the study of the nervous system, we are hardly warranted, however, in removing it from the Opisthobranchs ; and for the present it will be safe to conclude that we are here dealing with inherited streptoneury.' The value of the elevation of the spire in the classification of the C0NID.E is discounted by what we find in the ontogeny of Conus cuspidatus (p. 32), where the amount of elevation is shown to be merely a peculiarity of the individual. The impossibility of satisfactorily diiferentiating Stcrcula from Pleurotoma when dealing with fossil specimens, is insisted upon (p. 37) ; and, incidentally, the position of the sinus as a prime element in the classification of the Pleurotomid^ is criticized. Although the majority of protoconchs in species of the Gastero- poda appear to be smooth and without sculpture, many exceptions are noted. In particular, Pleurotoma icangamiieiisis (p. 46) is remarkable in that respect. The initial turn of the protoconch is smooth and obsoletely striated, and this is followed by an obtuse angulation, which rapidly develops into a bold, acute, median spiral carina, with accompanying lineation. See also Columharium acan- thoslephes (p. 52), Telcoclulus gracillimuin (p. 65), Harpa ahhreviata ' Howes, Proc. Make. Soc. London, vol. ii. 1896, p. 68. * Bull. Soc. Pliilom. Taris, t. v. 1893, p. 64. XU INTRODUCTION'. (p. 81), Latlriis tatei (p. 147), Leucozonia staminea (p. 151), Pseudovaricia mirabilis (p. 160), and Murex otwayensis (p. 177). Althougli Tve occasionally find that the main features of the sculpture of the adult are foreshadowed even in the protoconch, that is by no means a general rule. The protoconch of Pleurotonia alta (p. 45), for instance, is perfectly smooth, yet in the brephic stage a most pronounced peripheral carina makes its appearance ; on the other hand, in P. wanganuiensis (p. 46) the strong spiral ornament obtains from the protoconch to the ephebic stage, though profuse additional ornament was suddenly produced in the brephic stage. On this evidence, and seeing how closely allied the two species are in some other respects, one would assume that the latter species most probably descended from an earlier stock of the former. By way of contrast we find in Bathytoma angustifrons (p. 49) and Morio gradatu (p. 201) that but few of the main features of the ornament in the adult were foreshadowed even in the brephic stage, and that ornament is not of a permanent character, but was modified at the caprice of the individual. That, and similar evidence given in this Catalogue, points pretty conclusively to a general rule that may be established (at least, so far as these Australasian fossils are concerned, and it is possibly of much wider application), namely : that when the main features of the ornament are foreshadowed in the early brephic stage, and especially when they obtain even in the protoconch, that ornament may be regarded as of value in the determination of species ; but when, on the contrary, the ornament does not make its appearance until the late neanic stage, and is not, even in an elementary sense, completed until what may be regarded, by analogy, as the early ephebic, that ornament merely characterizes the individual, and is only of negative use for the purposes of classification. The size of the protoconch is very variable (e.g. Cancellaria wannonensis, p. 66), as is well known, even in the offspring of a single individual ; that difference being commonly attributed to carnivorous proclivities on the part of the larger specimens when in the embryonic stage. In this connection it is noteworthy that the size of the protoconch does not seem to have much influence in determining the size of the shell in the adult. The larger protoconch is not ver}' frequently, it would seem, accompanied by the production of a larger adult shell than comes from a much smaller protoconch, that is, in the same species. There are, INTRODUCTION. however, exceptions to tliis, and correlatively it may be noted that the shape of the protoconch occasionally determines the general shape of the shell. In Bathytoma angustifrons (p. 49), for instance, the Museum specimens show that the elevation or depression (as the case may be) of the turns of the protoconch to a large extent determines the relative elongation of the growth of the whole shell, in that particular species. Evidence is given (pp. 95, 96, and description of the species of Scaphella) that some of the Australasian Tertiary Volutid^ possessed horny protoconehs ; and the classification of that family as deduced from a study of the larval shells, together with some account of the origin and development of the groups they represent, will be found (p. 92 et sqq.). Cf. Fusus hezagonalis (p. 131) in this connection. The ontogeny of Mitra multisiilcata (p. 120) forms an interesting study. As a result of the details of gi'owth, it is shown that the number of columellar plications is not necessarily of value in classification in such a genus as Mitra ; for in the brephic stage there were only two plaits, and the suspicion of a third, whilst in the ephebic no fewer than five are observable. A very remarkable mode of growth is recorded in reference to Tylospira coronata (p. 222) ; whilst Cerithium apheles (p. 224) is full of interest. The shell in the latter exhibits longitudinal costfe in the brephic stage, which are lost in the neanic, but reappear strongly in the ephebic, only to die out again in the gerontic stage. There is thus a high possibility of this species having descended from a well-marked longitudinally costate stock, with a tendency for its descendants to assume a much plainer type of ornament. I have stated in an earlier part of this Introduction that, although it is generally understood that during the brephic stage of growth no specific characters can be distinguished, it was desirable to discuss the matter. Some of the observations just made prove that the shell is often highly sculptured in that stage, and many other examples are recorded in the text. It is by no means rare to find that the chief ornament relied upon for the classification of the species (in regard to individuals) suddenly makes its appearance in the brepliic stage, and, not infrequently, starts abruptly from the termination of the protoconch. A glance through the plates accompanying this work will, perhaps, help to make this observation INTRODUCTION. clearer. Dr. Jackson's investigations' led him to the con- clusion that characters almost inseparable from those of the adult frequently make their appearance in the brephic stage of the moUusca. In view of these facts it seems desirable to modify the general statement that during the brephic stage no specific characters can be distinguished. As a rule, no doubt, that statement is perfectly accurate, and the brephic stage is unquestionably well defined in the Gasteropoda as a whole ; but a little qualification seems necessary. I have found that the greatest difficulty in defining the brephic stage is in those cases where a strong varix has been thrown up at the conclusion of the embryonic stage, and it seems right that that should be so. Por the varix certainly indicates a pause in the growth of the shell, and it is reasonable to assume that during that pause the animal was passing through the brephic stage, but did not continue to make the normal shell of that period, except partially, it may be, in some instances. I have even doubted whether in certain instances the varix alluded to was not, in fact, the only manifestation of the growth of the shell during the brephic stage. Turning now to some other points brought out in this Catalogue, it may be noted that it introduces frequent changes in nomen- clature, especially in regard to genera and subgenera. Many well-known names, I am sorry to say, have disappeared as the result of bibliographic research. But I am not proud of having made these changes — they were inevitable if the rule of priority must be applied. In particular, I have been at some pains to discover the names of the type species of both genera and subgenera, in consequence of which a new interpretation has, occasionally, had to be placed on a well-known group. Some new genera and species are proposed; and species hitherto described have been considerably revised in regard to their generic position. In the description of species my general plan has been to give only the salient features, usually in the nature of a short diagnosis, where the species had formerly been fully treated of ; but where the material in the Museum collection permitted, and the species had hitherto received but scanty notice, it has been dealt with at greater length, ' "Phylogeny of the Pelecypoda " : Mem. Boston Soc. Nat. Hist. vol. iv. No. 8, 1890, p. 290. INTEODDCTION. XY Some difficulty has occasionally been experienced in assigning species to their correct geological position in the Tertiary. In. regard to the Australian "Eocene" and Miocene I have, where possible, followed the classification of Professor Tate in his various works quoted, although, as will already have been noticed, I have considerable doubt whether the "Eocene" of that continent is really homotaxial with that of North-Western Europe. Again, in certain localities where two formations occur, as at Muddy Creek, Trclissick, etc., it has sometimes been doubtful from which horizon the particular specimens being catalogued came. In such cases I have been much assisted by the useful list of Muddy Creek fossils drawn up in Mr. John Dennant's memoir^ on the beds in that locality, which has been quoted from time to time. Again, Sir James Hector's list^ of New Zealand localities with their geological horizons have proved valuable in the same connection ; whilst Professor Button's Catalogue ' has of necessity been i-eferred to on many occasions. Nevertheless, there are a few specimens from that colony, the exact provenance of which I have not been able to fix, as will be observed ; such specimens have, for the most part, been in the Museum for many years, and the name of the locality, which they bear, may possibly represent a place where a camp was temporarily made. The collection from Parimoa appears to come from two horizons, the Miocene and Pliocene, and in the case of a new record for that locality it has been found impossible to give the geological age with certainty. At Wanganui, again, both Pliocene and Post-Pliocene beds are found ; in general it has not beea difficult to assign a particular species from this locality to its correct horizon, but there are exceptions. These, and similar points, I must leave to Colonial geologists, who alone, by study on the spot, can satisfactorily clear them up. The locality of Limestone Creek, Glenelg river, Victoria, is here called Post-Pliocene, instead of Newer Pliocene ; I believe it will ultimately be found to be younger than the true Pliocene. It should be explained that by " Australasia " throughout this work is meant, the Australian continent, Tasmania, New Zealand, and the Chatham Islands. 1 Trans. Roy. Soc. South Aust. Tol. xi. 1889, p. 30. - Kep. Geol. Exp. 1890-91, p. 121 (and other Reports in the same Series). » Catalogue Tert. Moll, and Echin. X.Z. 1873. XVI INTRODUCTION. This ■work will not embrace the description of the Tertiary Cephalopoda in the British Museum, as they are being catalogued by Dr. A. H. Foord and Mr. G. C. Crick'; but it may be mentioned that so far as Australasia is concerned only two Tertiary species, viz., Aturia atari, Basterot, var. australis, M'Coy, and Nautilus geelongensis, Poord, are found in the collection, and they have already been dealt with.- Finally, I must express my indebtedness to several friends who have assisted me, from time to time, during the preparation of this Catalogue. Foremost stands Mr. Edgar A. Smith, whose courtesy in permitting me frequently to consult the modern moUusca in the Zoological Department of the Museum, and for much advice in regard to matters of nomenclature, I most thankfully acknowledge ; Mr. R. Bullen I^ewton has also rendered me assistance in regard to nomenclature, for which I tender him my best thanks ; Mr. B. B. Woodward has on several occasions given me the advantage of his extensive knowledge of bibliography, which has been of signal service, and for which I am greatly indebted to him ; Mr. C. D. Sherborn has kindly permitted me to refer to the MS. of his Index Animalium. Professor Tate, of the University of Adelaide, kindly sent me a collection of specimens and copies of his publi- cations on the Tertiary moUusca of Australia, which have been of much use ; whilst I am indebted to my friend Mr. H. W. Burrows for his ready assistance in reference to the classification of mollusca. To Miss G. M. Woodward I must also tender my thanks for the preparation of the beautiful drawings which adorn this Catalogue, GEO. F. HARRIS. 2hth March, 1897. ' Catalogue of Fossil Cephalopoda, British Museum. 2 Id. Part II. 1891, pp. 332, 354. TABLE FAMILIES, GENEEA, AND SUBGENERA. » Class GASTEROPODA. Order Puliionata. Family Helicid^e: page Helix 1 Liparus ........ 2 Family Bdlimulid^ : Placostylus ....... 4 Family Limn^id.e : Limnrea . 4 Family Amphibolid.e : Amphibola 5 Oi'der Opisthobranchiata. Family Act^onid^e : Actseon 7 Solidula 8 Tornatellsea ....... 9 Triploca 9 Adelactreon 10 Family Scaphandrid^. : Scaphander 12 Family Bullid^e : Bulla 13 Bullinella 13 Family RixGiccLiDiE : Ringicula 15 b XVlll TABLE OF FAMILIES, GENERA, AND SUBGENEIU. PAGE Family LuiACiNiDiE : .^g Limaciua .•••■■' Family Cavoliniid^ : ^g Clio Chelyconus , Cythara Clathurella . Narona . Admetula Sveltia . An cilia Glabella Cryptospira 20 Styliola ■••■■■' 21 Vaginella .•••■'■' Family UiiBRACULiDiE : 22 Umbraculum .••■■■' Order Prosobranchiata. Family Terebrid^ : 2^ Terebra .•••■''' Family Conid^ : 27 Conus .••••■■■ 27 Leptocouus . . ■ • • ■ Lithoconus •••■•■ 34 Family Pleurotomid^ : gg Pleurotoma ••■■'' 43 Bathytoma ■'■"'' 51 Columbarium ' " " ' ' 55 Drillia • • ' g^ Mausilia .■■•■■■' 58 .59 • 60 Daphuella .■•••" . Teleochilus " ' ' Family Cancellariid.e : ^^ Caucellaria •■■■■"■„_ 68 . 69 Family Olivid^ : ^^ Olivella 72 Family Harpid/e : ^g Harpa •■■■"" 78 Eocithara ••••'■ Family Marginellid^ : g2 Marginella 90 lAULE OF FAMILIKS, GENERA, AND SOBGENEllA. xix Family Volutid.*; : page Volutilithea 97 Lyria 98 Voluta •■•..... 99 Pterospira 100 Aulica 101 Volutocouus ..... 107 Amoria 108 Scapliella HO Eopsei^hia 115 Family Mitrid.e : Mitra 119 Caucilla . . . . . .123 Uromitra 124 Conomitra . . . . . .129 Family Fasciolariid.b : Fusus 131 Latirofusus . . . . . . . .1.15 Streptochetus 1,37 Fasciolaria ... .... 138 Latirus ]42 Leucozonia . . . . . . _ .148 Latirolageua . . . . . 150 Family Buccinid.-e : Tritonofusus .152 Siplionalia . . . . . . 154 Pseuflovaricia . . . . . .159 Cautharus Igj Eiithria . . . . . . . Iii2 Donuautia . . . . . 162 Phos 1()4 Loxotaphnis . . . . . 165 Zemira .167 Family Nassid.e : Nassa .168 Family Ml'ricid.b : Typhis 170 Murc.x . . .172 Tiiplox 172 Chicoieus 178 Muricopsis 179 Muricidea . . . .181 TABLE OF FAMILIES, GENEKA, AND STJBGENEKA. Family Muricid^ {continued) : page Sistrum 184 Family Lotoriid^ : Lotorium 185 Colubraria 194 Apollo 195 Family Cassidid^ : Cassis 197 Semicassis 198 Casmaria 200 Morio 201 Family Ctpr^id^ : Cyprsea 203 Bernayia 204 Luponia 205 Erosaria 208 Umbilia 209 Trivia 212 Erato 214 Family Strombid^ : Strombus 216 Seraphs 218 Family Strdthiolariid.e : Struthiolaria 218 Tylospira 222 Family CERiTHiiDiE : Cerithium 223 Colina 227 Batillaria 228 Newtoniella 229 Triforis 231 Diastema 231 Family Trichotropid^ : Trichotropis 233 Family Vermetid^ : Tenagodes 233 Family Turritellid^e : Turritclla 234 Family Solariid^ : Solarium ........ 243 Heliacus 245 TABLE OF FAMILIES, GENERA, AND SUBGENERA. Family Hipponycid^ : Hipponyx Family Capulid^ : Crepidula . lauacus Family CALYPTRiEiD^ : Calyptrrea . Calyptropsis Family Xenophorid^ : Xenojihora Tugurium Family NATiciDiE : Natica Lunatia Stigmaulax . Sigaretopsis Euspira Ampiillospira SEQUENZIIDiE : Seqiienzia . SCALID^ : Cirsotrema Crosseia Eglisia EULIMIDiE : Eulima Niso TURBINIDiE ; Turbo Phasianella Collonia Family Family Family Family Family Trochid^ : Trochus . Authora Coelotroclius Claiiculus . Thalotia . Bankivia Liopyrga Ethalia Solariella PAGE 246 246 248 249 251 253 254 255 260 262 263 264 265 266 267 269 270 271 272 272 275 276 276 277 278 279 279 280 281 282 283 TABLE OF FAMILIES, GENEKA, AND SUBGENERA. Family Delphinulid.e : page Liotia 283 Family HALiOTiDiE : Haliotis 284 Family Pleurotomariid^ : Pleurotomaria . 286 Family Fissdrellid^ : Fissurellidea . . . . . . . 2 "^7 Emarginula ....... 288 Subemarginula ...... 289 Tugalia 289 Class SCAPHOPODA. Family Dentaliid^ : Dentalium 290 Cadulus 297 Class LAMELLIBRANCHIATA. Order Tetrabranchia. Family OsTREiDiE : Ostrea 298 Gryphgea 3U2 Family Anomiid^e : Placimanomia ....... 303 Family Dimyid^ : Dimya 305 Family SpoNDTLiDiE : Spondylus 307 Family Limid^e : Lima . 309 Limatula . . . . . . 311 Limea 312 Family Pectinid^e : Peoten 313 Pseudamussium ...... 322 Amussium ....... 324 Family Aviculid^ : Margaritifera 325 Vulsella 326 Pinna . 326 TAliLE OF I'AMILIES, GENERA, AND SOBGIONEUA. XXiU Family MYTiLiDiE : page Mytilus 327 Septifer 328 Modiola 329 Creiiella 329 Family ARCiDiE : Area 330 Barbatia 331 Plagiarca ...... 335 Fossularca ..... 335 Cucullsea 336 Pectimculus 339 Limarca ... .... 343 Limopsis 344 Family NuculiDjE : Nucula 347 Family Nucolanid.e : Nuculana 348 Family Trigoniid^e : Trigonia ........ 352 Family Unionid^ : Uoio 355 Anodonta 356 Family Carditid^ : Cardita 357 Carditella 362 Family Crassatellitid.E : Crassatellites 363 Family Cardiid.i: : Cardium 366 Protocardium 368 Family Chamidje : Chama 369 Family VEXERiDiE ; Meretrix 370 Sunctta 372 Chione 372 Family Ungulinid.'e : Diplodonta 375 Family Donacid^'e : Doiiax 376 7 TABLE OF FAMILIES, GENERA, AND SOBGENEKA. Family Gariidj. : page Gari 377 Family Mesodesmatid^e : Mesodesma ....... 378 Family Mactrid^ : Mactra 379 Zeoatiopsis ....... 380 Family Myid^ : Corbula 381 Family SAXiCAViD.a; : Capistrocardia 383 Family PnoLADiDiE : Barnea 384 Order Dibranchia. Family LuciNlD^ : Lucina 385 Family Tellinid^ : Tellina 386 Family Scrobiculariid^ : Semele 388 Family Cuspidariid.5; : Cuspidaria 389 Family Pandorid^ : Myadora 390 LIST OF COLLECTIONS. The following are the principal sources whence the Collections catalogued in this work were obtained : — 1. A series of mollusca from the Miocene and Pliocene of New Zealand, chiefly from Parimoa and Onekakara. Presented by the Hon. W. B. B. Mantell, about the year 1860. 2. From various horizons of the Tertiary in South Australia. Presented by the Governors of Adelaide ITuseum, in 1861. 3. A few specimens from the Tertiaiy of Adelaide, South Australia. Presented by M. Marshall, Esq., in 1861. 4. From the Tertiary of the River Murray, South Australia. Presented by Commodore Goodenough, R.N., in 1876. 5. From the Pliocene and Post-Pliocene of New Zealand. Pre- sented by Lieut.- Col. L. W. Wilmer, late 12th Regiment, in 1882. 6. Several specimens from the Tertiary of Bairnsdale, Victoria. Presented by W. H. Grigson, Esq., in 1882. 7. A large and important series from the Eocene and Miocene of Muddy Creek, Hamilton, Victoria. Presented by John Bennant, Esq., F.G.S., in 1886. 8. A large series from various horizons of the Tertiary of South Australia, and a few specimens from Western Australia. Presented by H. Y. L. Brown, Esq., F.G.S., in 1887. 9. Many specimens from the Tertiary of Gippsland, Victoria. Presented by the liev. J. S. M. Rogce, in 1887. 10. From the River Murray cliffs. South Australia. Presented by Wm. Evans, Esq., in 1887. c XXVI LIST OF COLLECTIONS. 1 1 . Several specimetis from the Eocene and Miocene of Muddy- Creek, Victoria. Presented by Professor T. Rupert Jones, F.Ii.S., in 1887. 12. From the Tertiary of Border Town, South Australia. Pre- sented by E. Litton, Esq., in 1888. 13. From a raised beach at Opua, near Russell, Jfew Zealand. Presented by F. A. Bather, Esq., M.A., F.G.S., in 1894. In addition to the above donations : — 14. A very large and valuable collection from various horizons of the Tertiary of New Zealand, obtained in exchange from Sir James Sector, K.C.M.G., F.R.S., in 1875. 15. Many specimens from the later Tertiary of New Zealand. Transferred from the Museum of Practical Geology, in 1880. 16. By purchase on several occasions, chiefly in 1876 {Strzelecki Collection), 1879, 1881, 1890, and 1896. Class GASTEROPODA. Order PULMONATA. Family HELICIDJE. Genus HELIX, Linnajus {em. Miiller). [Linn. Syst. Nat. ed. 10, 1758, p. 768; Miiller, Verm. Terr. Fluy. vol. ii. 1774, p. 12.] Several recent writers have referred to H. pomatia, Linnaeus, as the type of Helix ; there seems to be no reason, however, for departing from the emendation of Miiller, as recognized by authors such as Pfeiffer,' who made an exhaustive study of the genus. Type. — Helix lampas, Miiller. Helix tasmaniensis, 6. B. Sowerby. 1845. Helix tasmaniensis, G. B. Sowerby, in Strzelecki, Phys. Desc. X.S.W. p. 298, pi. six. fig. 5. 1878. Helix tasmaniensis, K. Etheridge, jun.. Cat. Aust. Foss. p. 178. 1880. Eelix tasmaniensis, Johnston, Pap. Roy. Soc. Tas. 1879, p. 88. 1888. Helix tasmaniensis, Johnston, Geol. Tasmania, pi. xxxiv. fig. 9. The description by Sowerby being based on very imperfect materials, the following amplification may be quoted: — "Shell thin, globosely conical, umbilicated ; whorls 4h, rapidly increasing, oniamented with fine obliquely-arcuate striaj, crossed by fine spiral lines . . . , flatly convex above, convex below, last whorl tumid and slightly descending in front ; suture somewhat im- pressed, spire depressly conical ; umbilicus deep, narrow, scarcely one-sixth of the diameter ; aperture oblique, ovately lunate ; ' Monog. Helic. Viv. 1«18, p. xv. peristome simple ; margins slightly approximating, basal margin slightly dilated and reAexed."— Johnston, 1880. Only one fragmentary specimen of the numerous examples in the Museum bears any trace of shell structure ; the remainder are internal casts. The ornamentation of the shell fully bears out Mr. Johnston's description, except that no spiral lines are to be seen on the fragment alluded to. The casts present consider- able variation in the elevation of the spire, occasionally, indeed, being somewhat depressed, when they closely resemble the living ffelix confusa, Pfr., of Queensland, North Australia, etc. In the last-mentioned shell, however, certain adult forms attain to con- siderable spiral elevation, especially in the specimens in the Museum from North Australia. Judging from the sculpturing and general aspect of the casts, it is highly probable that IT. tasnianiensis should be classed in the subgenus Rhytida, but the materials are not perfect enough to enable one to be quite certain on this point. Mr. Johnston {op. cit. 1880, p. 89) describes two new species of Helix from the same locality as the Museum specimens come from : one of these, H. huxleyana, is very minute ; and the other, S. geilstonensis, is described as orbicularly depressed, the whorls being markedly convex below ; a variety of H. sinclairi is also created which has, inter alia, a narrow umbilicus. Neither of these is represented in the British Museum Collection. Dimensions (of the type specimen). — Height 22i mm. ; breadth 34 mm. Form, and Loc. — Yellow Limestone (Travertine), probably of Pliocene age : all from the neighbourhood of Hobart Town, Tasmania. 96906. The type specimen figured by Sowerby. Purchased. 46592-3. Two specimens. Presented hj Norton Allport, Esq. 98217. Two casts ; from Geilston Quarry. Purchased. G. 71. Several casts of adult specimens. Purchased. Genus LIPARITS, Albers. [Die Heliceen, 1850, p. 172.] Test ovately conical, with an obtusely conical spire, whorls to the number of 6 or 7, aperture ovate, peristome simple, acute, I columella twisted so as to form an obscure but somewhat callous plication, the slight callosity almost covering the very minutely perforated umbilicus, and extending more or less over the body- whorl towards the posterior portion of the aperture. Type. — Bulimus atomatus, Gray. Liparus gunni, G. B. Sby. (sp.). 1845. Bulbms gumiii, G. B. Sowerby, in Strzelecki, Phys. Desc. N.S.W., etc., p. 298, pi. xix. fig. 6. 1878. Bulimus gunnii, E. Etheridge, jun., Cat. Aust. Foss. p. 177. 1880. Bulimtis gunnii, Johnston, Pap. Roy. Soc. Tas. 1879, p. 90. 1888. Bulimus gunnii, Johnston, Geol. Tasmania, pi. xxiv. fig. 7. This species is represented by one example only in the British Museum — the type figured by Mr. Sowerby. It is a cast of the interior, and no vestige of the shell itself appears thereon. Mr. Johnston was only acquainted with fragments of the cast, l^ever- theless the present writer is of opinion that the structure of the shell may, to some extent, be inferred from a careful examiuation of the type specimen. From the deepness and general aspect of the sutures it is certain that the test must have been very thin, and, as in certain other representatives of the genus Liparus, the lines of growth no doubt existed both interiorly and exteriorly. These latter are strongly reflected in the cast under consideration, from which it is legitimate to conclude that the shell was by no means smooth, although Mr. Sowerby says " leeviffatis?" Unfortunately, a large part of the cast is imbedded in the hard calcareous matrix in such a manner as to mask the phenomena of the aperture, so that the reference of the species to the genus Liparus is not altogether without doubt. When the specimen is compared with the living Liparus leeuivinensis, E. A. Smith,' of N.W. Australia, however, the doubt almost decreases to the vanishing point, for, as far as can be judged from an internal cast, the two must be of the same genus, and, indeed, they are closely allied specifically. The chief specific difference is the more conical and tumid shape of the fossil. On comparing the latter with the living L. kingi, Gray, from King George's Sound, which ' Proc. Malac. Soc. Loud. vol. i. 1894, p. 94, pi. vii. fig. 27. 4 PLACOSTTLTJS. it also resembles in some respects, it is found not to be so elongate, and the lines of growth are more oblique. Dimensions. — Height 26 mm. ; breadth 14^ mm. Form, and Loc. — Yellow Limestone (Travertine), probably of Pliocene age : Hobart Town, Tasmania. 96907. The type specimen figured by Sowerby. Pureliased. Genus PLACOSTYLUS, Albers. [Die Heliceen, 1850, p. 147-] Flacosffhts, Eeck (MS.), Index MoUuscorum, 1837, p. 57 (list name only). Test imperforate, oblong, conical, rugosely striated; whorls to the number of 7 or 8 ; columella twisted, plicated ; aperture irregular but mostly ovate, or oblong ; peristome thick, having an expanded and reflected callous margin, tuberculated neai' the anterior extremity of the aperture. T>/pe. — Bulimus fihratus, Martyn. Placostylus, sp. An imperfect specimen in the Museum collection presents the generic characters of Placostylus, but its condition precludes the possibility of referring it accurately to any known species. It nearest resembles the living P. hovinus, Brug., of N^ew Zealand, and its close ally the variable P. fibratus, Martyn. Form, and Loc. — Post-Pliocene: King's Creek, Darling Downs, Queensland. 75373. One specimen. Presented by Richard Daintree, Esq. Family UMJSmiDM. Genus LIMN^A, Lamarck (em. Rang). [Lamlc, Mem. Soc. Hist. Nat. Paris, 1799, p. 75; Sander Rang, Man. Hist. Nat. Moll. 1829, p. 176.] liimnaea, sp. Two indeterminate specimens of this genus exist in the collec- tion from South Australia ; one an imperfect internal cast from Tertiary beds in Fowler's Bay district, the other a monstrosity from Willunga. S. Y. L. Brown Coll. Family AMPHIBOLIDiE. Genus AMPHIBOLA, Schumacher. [Essai Nouv. Syst. Habit. Testaces, 1817, pp. 58, 190.] Ampullacera, Quoy and Gaimard, Voy. de 1' Astrolabe, Zool. t. ii. 1832, p. 196, pi. xv. figs. 1-9. Thallicera, Swainson, Malacology, 1840, p. 196. The test of this genus -is not unlike an Ampullaria, thoiigh it differs in many essential particulars. It is thick, subglobose, and rugose ; spire somewhat depressed ; outer margin of the aperture with a deep sinus posteriorly ; inner margin callous and expanded ; columella broad and reflected at the base ; umbilicus large. Type. — Amphilola australis, Schumacher. Amphibola avellana, Bruguiere (sp.). 1775. Spengler, Beschaftigungeii d. Natur. Freunde, Berlin, Bd. i. p. 395, pi. ix. figs. 4, 5. 1781. Nerita nnx avellana merito, etc., Chemnitz, Conch. Cab. Bd. v. p. 262, pi. clxxsviii. figs. 1919-20. 1784. Limax crenata, Martyn, Univ. Conch, pi. Ixix. 1789. Bulimus avellana, Bruguiere, Ency. Meth. [Vers] t. i. p. 297. 1790. Helix avellana, Gmelin, Linn. Syst. Nat. ed. 13, p. 3640. 1790. Helix crenata, Gmelin, Linn. Syst. Nat. ed. 13, p. 3623. 1802. Helix avellana, Bosc. Hist. Nat. des Coq. Paris, t. iv. an. s. (1801-2) p. 38. 1802. Bulimus avellana, Bosc. Hist. Nat. des Coq. Paris, t. iv. an. x. (1801-2) p. 92. 1817. Amphibola australis, Schumacher, Essai Nouv. Syst. Habit. Testaces, p. 190. 1832. Ampullacera avellana, Quoy and Gaimard, Voy. de 1' Astrolabe, Zool. t. ii. p. 196, pi. XV. figs. 1-9. Ampullaria avellana | Deshayes and Milne Edwards, Lamk. Hist. Ampullacera avellana) Nat. Anim. sans Vert. 1838, pp. 537-41. 1840. Thallicera avellana, Swainson, Malacology, p. 196. 6 A31PHIB0LA. 1873. Amphibola avellana, Von Martens, List Moll. N.Z. p. 40. 1873. Amphibola avellana, Hutton, Cat. Marine Moll. N.Z. p. 58. 1873. Amphibola avellana, Hutton, Cat. Tert. Moll. N.Z. p. 17. 1879. Amphibola avellana, Hutton, Ann. Mag. Nat. Hist. vol. iii. 5th ser. p. 181, pi. xxii. 1880. Amphibola avellana, Hutton, Man. N.Z. Moll. p. 35. 1893. Amphibola avellana, Hutton, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 36. The inner margin of the aperture, in certain individuals, is extremely callous and folded into a rough plication in the vicinity of the umbilicus ; the latter is broad and keeled outwardly. The species is frequently quoted as A. nux-avellana, Chemnitz; but as that author's designation of it is not binomial it cannot be accepted, and the description as a whole must be ignored. Martyn in 1784 figured the species under the name of Limax crenata, without description. The first author to regularly describe it was Bruguiere, and he included it under Bulimus, a circumstance which Schumacher overlooked, apparently, in creating the generic name. Deshayes and Milne Edwards express their willingness to acquiesce in the genus AmpuUacera proposed by Quoy and Gaimard, but, as in other cases, adhere to the description given by Lamarck. It inhabits salt marshes near the sea, or lives in brackish water. Professor Hutton, in 1879, published particulars concerning its anatomy, amplifying those previously given by Quoy and Gaimard, in which he remarks that " The animal lives between tide-marks in salt or brackish water on mud flats in sheltered bays. When found at all, it is always found in large quantities. It is very sluggish in its habits, and feeds on the vegetable matter contained in the mud, passing large quantities through its alimentary canal. Although air-breathing, it will live for a week or ten days in fi-esh water, and for more than a fortnight in salt water, without being exposed to the air." Dimensions. — Height 23 mm. ; breadth 2ih mm. Form, and Loc. — Post-Pliocene : Wanganui ; Opua, near Russell, I^ew Zealand. G. 466. Two specimens. Presented by Lieut.- Col. W. Wilmer. G. 8281. Several specimens from a raised beach at Opua, 60 ft. to 100 ft. above sea-level. Presented ly F. A. Bather, Fsq., M.A., F.G.S. Order OPISTHOBRANCHIATA. Family ACTiEOMD^. Genus ACTION, Montfort. [Conch. Syst. t. ii. 1810, p. 315.] "Tornatelle," Lamarck, Cours Zool. 1812, p. 117. Tornatella, Lamarck, Hist. Nat. Anim. sans Vert. t. vi. pt. 2, 1822, p. 219. Speo, Risso, Hist. Nat. Eur. Merid. t. iv. 1826, p. 235. Myosota, Gray, Proc. Zool. Soc. Lond. 1847, p. 160. Characterized by a not very prominent protoconch with sinistral nucleus; thick columella with one strong spiral, slightly oblique plication. Type. — Valuta tornatilis, Linn. Actaeon scrobiculatus, Tenison-Woods. [Plate I. Figs, la-c] 1877. Actceon scrobiculatus, Temson-Woods, Pap. Roy. Soc. Tas. 1876, p. 102. 1889. Tornatella scrohiculata, Dennant, Trans. Eoy. Soc. South Aust. vol. xi. p. 48. 1893. Tornatella scrohiculata, Tate and Dennant, Trans. Roy. Soc. South Aust. vol. xvii. p. 223. This species is described as follows by Mr. Tenison- Woods : — " Shell oblong, ovate, solid, apex acute, smooth only at the extreme summit; whorls seven, cancellate with very distinct spiral liraj, much finer longitudinal striae ; interstices rounded or punctate, lii-aj on the last whorl broad and subdivided by a fine groove, longitudinal striae subdistant (so that the interstices are transversely oblong) and passing occasionally over the lirse, so as to make them subgranular, especially at the anterior margin ; aperture subauriform, posteriorly acutely attenuate ; peristome anteriorly everted and recurved, plait conspicuous, solid, obtuse." The species is compared with A. pinguis, D'Orbigny, of the European iliocene, from which it may be distinguished in being narrower, and in the disposition of the columellar plication. The 8 size of the plication, its situation, and the excavated columella behind it, recall the genus Solidula to some extent, though it materially differs in only possessing one plication, which is not bifurcate. Mr. Tenison- Woods does not give a figure of this species, but from his description the identification appears certain ; it is now figured. Dimensions (of the figured specimen). — Length 9|^ mm. ; breadth 4^^ mm. ; length of aperture 6 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G; 4296. One specimen. Presented hy John Dennant, Esq. Genus SOLIDULA, Fischer de Waldheim. [Museum Demidoff (Moscow), t. iii. 1807, p. 226.] Dactylus, Schumacher, Essai Nouv. Syst. Habit. Testaces, 1817, pp. 70, 234. Buccimdus [Plancus), H. and A. Adams, Gen. Rec. Moll. vol. ii. (1854) p. 5. The columella of this genus carries two plications, the anterior prominent and bifid, the posterior comparatively inconspicuous when the shell is entire ; between them the columella is spirally excavated. These, amongst other characters, distinguish Solidula from Adeeon. Type. — Voluta solidula, Linn. Solidula alba, Hutton (sp.). 1873. Buccimdus albus, Hutton, Cat. Marine Moll. N.Z. p. 51. 1873. Buccinulus albus, Hutton, Cat. Tert. Moll. N.Z. p. 16. 1880. Buccinulus albus, Hutton, Man. N.Z. Moll. p. 119. 1893. Tornatella alba, Hutton, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 37, pi. vi. fig. 2. This is a typical example of the genus. It is spirally and closely sulcated, and longitudinal strise are distinctly seen in the grooves. On one of the British Museum specimens several flat ridges between the sulci bear a minor sulcation. The sutures are not deep ; the outer margin of the aperture is extremely thin, and is not sinuous anteriorly. TORNATELLiEA. 9 Dimensions. — Length 14 mm. ; breadth 7 mm.; length of aperture 9 mm. Form, and Loc. — Pliocene : " Lower beds " at Wanganui, N.Z. G. 464. Two specimens. Presented ly Lieut.- Col. W. Wiliner. Genus TORNATELL^A, Conrad. [Journ. Acad. Nat. Sci. Philad. (n.s.) vol. iv. p. 294.] Several authors have confounded Tornatellma with Solidula, but it may be readily distinguished from the latter by the marked depression on the anterior portion of the aperture ; by the difference in disposition of the columellar plications, which are brought more prominently forward, the foremost anterior plication being situated relatively nearer the anterior extremity of the shell ; and by the greater thickness of the shell in the neighbourhood of the outer border of the aperture, which is frequently crenulated. These characters suffice, also, to justify separation from Actceon, sensu stricto. It may be remarked, that the fossil usually known as Actceon simulatus, from the Barton beds, is undoubtedly a re- presentative of this American genus in England. Tornatellcea, moreover, has sometimes been confused with Tornatella, Lamarck, which is a synonym of Actceon. Tijpe. — Turnatellcea hella, Conrad. Subgenus TRIPLOCA, Tate. [Journ. Roy. Soc. N.S.W. vol. xxvii. 1894, p. 186.] The principal difference between this and Tornatellcea, sensu stricto, consists in the former having three columellar plications instead of two, the two anterior of which are situated close together ; the spire is longer, and the outer border of the aperture is not crenulated. Type. — Triploca ligata, Tate. Tornatellsea (Triploca) ligata, Tate (sp.). 1894. Triploca ligata, Tate, Journ. Roy. Soc. N.S.W. vol. sxvii. p. 186, pi. xi. fig. 7. 1895. Tornatclhea {Triploca) ligata, Cossmann, Ess. Paleoconch. Comp. p. 50. 1 0 ADELACT^ON. Shell thick, spirally linear sulcate ; sulci not punctated, close together and varying in prominence, sometimes obliterated. M. Cossmann states that this species possesses minute punctations in the spiral sulci, but neither of the specimens in the Museum exhibit that peculiarity, though when seen under the microscope slight irregularities exist where the faint lines of growth intersect the sulci. The shells are slightly more tumid than that repre- sented by the figure given by Professor Tate; the other specific characters, however, are identical in the main, and the present writer hesitates to make a specific distinction on such unimportant diiferences. The species is evidently very variable; even the deep sulcation bordering the suture (so prominent in the speci- mens selected for description by the last- mentioned author) is occasionally reduced to such dimensions that it barely surpasses in size the ordinary spiral sulcations. The latter, too, are by no means always persistent over the whole of the whorls, one of the Museum specimens showing the body-whorl sulcated over two- thirds of the surface only. It would seem that when ornamented in the manner lastly described, the sulcation bordering the suture is stronger, as though the missing sulci had gathered together at that point. On the other hand, when sulcated throughout, the sutural sulcus is relatively smaller — a phenomenon observable also in certain European forms of Act^onid^, where it seems to be merely an individual characteristic. Q'he visible portion of the protoconch (Plate I. Figs. 2a-l) consists of about H turns, and is remarkably small, hiding the sinistral nucleus — at least in the Museum specimens. Dimensions. — Length 5-5 mm. ; breadth 3 mm. ; length of aperture 3 mm. Form, and Loc. — Eocene : Adelaide. G. 9305. Three specimens. Purchased. Genus ADELACT..ffiON, Cossmann. [Ess. Paleoconcli. Comp. 1" liv. 1895, p. 54.] Mijonia, A. Adams, Ann. Mag. Nat. Hist. 3rd ser. vol. v. 1860, p. 406 {non Dana, 1847). Protoconch with sinistral nucleus ; not very large. Sutures of the shell canaliculated, or impressed ; whorls decussated by fine ADELACTiEON. 11 stria3, lines of growth inconspicuous; columella slightly excavated and carrying a small but well-marked plication ; columellar border expanded and frequently covering the small umbilicus. This genus resembles Actaon in its general characters, but may be readily distinguished when the earlier portion of the protoconch can be satisfactorily determined ; the protoconch, as a whole, is not so well developed as in Adceon. In proposing this new name for Myonia, M. Cossmann remarks that the fossil forms slightly differ from the living, and he bases a new description and diagnosis on a Miocene species, Acteeon papyraceus, Basterot, from the neighbourhood of Bordeaux, quoting that species as the type of the genus. But, in substituting a generic name for one preoccupied, it is desirable to conform to usage by quoting as the type of the genus that appertaining to the genus displaced ; therefore the type species proposed by the French author alluded to should be superseded by that referring to Myonia. Type. — Myonia japonica, A. Adams. Adelactaeon olivellaeformis, Tate (sp.). 1894. ActcBopyramis oUvellaformis, Tate, Journ. Roy. Soc. N.S.W. vol. xxvii. p. 181, pi. xi. fig. 2. 1895. Adelactieon oUvellaformis, Cossmann, Annuaire Geol. Univ. t. x. p. 756. Protoconch (Plate I. Figs. 3a-b) small, earlier portion hidden by later turns in many specimens, but comparatively free in the one now figured. It is smooth and polished ; more elevated than in A. papyraceus, and is implanted obliquely on succeeding whorls. Shell as a whole elongate ; aperture less than half the total length of the shell ; whorls slightly turriculate, smooth, with minute spiral strias, which become more prominent on the anterior portion of the body-whorl. The classification of this species, as remarked by M. Cossmann {op. cit. supra), is a matter of doubt ; the embracing character of the last whorl is distinctly that of the Act^onid^, but the earlier part of the protoconch is different to that of Adelactaon, which, sensu stricto, is obtuse, and is implanted in such a manner as to render its sinistral character hardly perceptible. 12 SCAPHANDER. Dimensions. — Lengtli 10 mm.; breadth 3 mm.; length of aperture 4 mm. Form, and Log. — Eocene : Muddy Creek, Victoria. G. 9307. Two specimens. Purchased. Family SCAPHANDRID^. Genus SCAPHANDER, Montfort. [Conch. Syst. t. ii. 1810, p. 335.] This genus may be readily distinguished from Bulla and Bullinella by the enormous capacity of the body-whorl, which is much dilated anteriorly ; there exists no columella properly so-called, the whorls not being in contact with each other. Type. — Bulla lignaria, Linn. Scaphander tenuis, sp. nov. [Plate I. Figs. 4(j-(;.] Shell with small but deep apical umbilicus ; spirally striated, the striae frequently being regular and equidistant over part of the surface and closing in antero-posteriorly, farthest apart in the median portion of the whorl, closest together towards the apex ; lines of growth conspicuous on the smooth exterior and in the thin spiral grooves, causing the latter to assume a minutely punctated appearance ; aperture extending the whole length of the shell, constricted posteriorly, expanded, curved, and produced anteriorly; inner margin furnished with a conspicuous and slightly reflected smooth band ; in consequence of the thinness of the shell, both striae and lines of growth are seen in the polished interior of the aperture. Mr. John Dennant, in giving ' a list of species of mollusca found in the Muddy Creek beds, alludes to a MS. species of Professor Tate as Scaphander fragilis. As that has, apparently, neither been figured nor described, the present writer is unable to state in what way it may be related to this species. 1 Trans. Eoy. Soc. South Aust. vol. xi. 1889, p. 48. 13 Dimensions. — The largest specimen is not perfect, but attained at least to the following dimensions : length 21 mm. ; breadth 12 mai. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4171. Tour specimens. Presented lij John Demiant, Esq. Family BTJLLID^. Genus BULLA, Linnseus. [Syst. Nat. ed. 10, vol. i. 1758, p. 725.] Bulla australis, Gray. 1825. Bulla austrnlis, Gray, Ann. Phil. vol. xxv. p. 408. 1850. Bulla oblonga, A. Adams, Thes. Conch, vol. ii. p. 577, pi. cxxiii. fig. 74. Two fragments, and an internal cast, referable apparently to this species, are in the Museum collection. One is rather more cylindrical than the typical form of the species, but is too imperfect for description. Dimensions. — Length 53 mm. ; breadth 32 mm. Form, and Loc. — Post-Pliocene (?) : South and "West Australia. 97721. Two specimens, from Logues, Champion Bay, "Western Australia. Presented hij R. Ether idge, E'^q., jun., and R. Y. L. Brown, Esq. 42691. An internal compressed cast, from "Tertiary beds," South Australia. Presented ly the Governors of Adelaide Museum. Genus BTJLLINELLA, U. B. Newton. [Syst. List Edwards Coll. B.M. 1891, p. 265.] BulUna, Risso, Hist. Nat. Eur. Merid. t. iv. 1826, p. 51 [non Ferussac, 1822). Cylichna, Loren, Ofv. K. Vet. Akad. Forhandl. 1847, p. 142 {tion Burmeister, 1844). Form cylindrical ; spire deeply perforated at the summit ; last whorl covering all the others ; surface generally ornamented by spiral striaj especially marked at the anterior portion of the whorl ; 14 BULLINELLA. outer marprin longer than the axis of the shell ; columella having near its anterior extremity a small plication ; often umbilicated. This genus may be distinguished from Bulla, with which it is often confounded, by being more cylindrical in form and by the possession of the columellar plication alluded to. Type. — Bulla cylindracea, Pennant. Bullinella exigua, Tenison-Woods (sp.). 1880. Cylichna exigua, Tenison-Woods, Proc. Linn. Soc. N.S."W. vol. iv. p. 19, pi. ii. fig. 6. 1893. Cylichna exigua, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. xvii. p. 223. This species appears to play the same rdle in the Australian Eocene as does B. uniplicata, J. de C. Sby. {—B. hruguierei, Deshayes), in the Eocene of North-western Europe, and B. galba, Conrad, and B. saint-hilairei, Lea, of homotaxial beds at Claiborne, Alabama, in the "United States. It is a long, cylindrical shell ; the aperture is dilated anteriorly ; columellar border flattened in the vicinity of the rather deep umbilicus and carrying a well-marked, twisted plication ; spiral striaj on the body-whorl most conspicuous towards the anterior portion, and on and in the deep apical de- pression ; rugosities of growth strongly developed. Dimensions. — Length 17 mm. ; breadth 7 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4297. Several specimens. Presented ly John Dennant, Fsq. G. 4748. One specimen. Pres. hy Prof. T. Rupert Jones, F.R.S. Bullinella arachis, Quoy and Gaimard (pp.). 1832. Bulla arachis, Quoy and Gaimard, Voy. de 1' Astrolabe, Zool. t. ii. p. 361, Atlas, pi. xxid. figs. 28-30. 1850. Bulla {Cylichna) arachis, A. Adams, Thes. Conch, vol. ii. p. 590, pi. exxv. figs. 133-4. 1873. Cylichna striata, Hutton, Cat. Marine Moll. N.Z. p. .52. 1873. Cylichna striata, Hutton, Cat. Tert. Moll. N.Z. p. 16. 1880. Cylichna striata, Hutton, Man. N.Z. Moll. p. 120. 1887. Cylichna arachis, Dennant, Trans. Roy. Soc. Vict. vol. xxiii. p. 240. 1893. Cylichna arachis. Button, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 37. non Cylichna arachis, Tenison-Woods, Pap. Eoy. Soc. Tas. 1876, p. 102 (=Cylichna ivoodsi, Tate). KINGICULA. 15 The specimens in the Museum are larger than the average of the living B. arachis ; also, they are relatively broader in pro- portion to their length and are not quite as cylindrical. In all other characters, however, they are essentially similar, and the differences indicated merely constitute a local variation of the species. Dimensions. — Length 22 mm. ; breadth 10 mm. Form, and Loc. — Post-Pliocene : Limestone Creek, Glenelg river, Victoria. G. 5579- Two specimens. Purchased. Bullinella, sp. A mutilated cast of what is evidently a very large example of this genus, probably allied specifically to B. arachis, forms part of the Museum collection. It is cylindrical, the sides being remarkably parallel to each other ; the apical perforation is relatively small, and there is a trace of a prominent denticle on the anterior portion of the columella. Ditiioisions. — Length 30 mm.; breadth 15 mm. Form, and Loc. — "Tertiary" : Null arbor Plains, South Australia. G. 9303. Internal cast, mutilated. H. Y. L. Brown Coll. Family RITs^GICULIDiE. Genus RINGICULA, Deshayes. [Hist. Nat. Anim. sans Vert. t. viii. ed. 2, 1838, p. 342.] Shell small, mamillated protoconch ; aperture canaliculated posteriorly, excavated anteriorly ; outer margin usually very thick, occasionally denticulated within ; columellar border thick and callous, the callus extending from the anterior extremity to the posterior canaliculation ; columella arched and furnished with from two to four plications. M. Morlet has grouped ' the species ' Journ. de Conchyl. 3^ ser. t. xviii. 1878, p. 113 et sqq. ; id. 3^ ser. t. xx. 1880, p. 150 ; id. 3^ ser. t. xxii. 1882, p. 2U0. 16 EINGICTTLA.. according to the presence or absence of denticulations on the outer margin of the aperture, and with reference to the number and disposition of the columellar plications. Type. — Auricula ringens, Lamarck. Ringicula lactea, Johnston. [Plate I. Figs, ba-d.'] 1880. Ringicula lactea, Johnston, Pap. Roy. Soc. Tas. 1879, p. 34. 1888. Ringicula lactea, Johnston, Geol. Tasmania, p. 239. 1893. Ringicula lactea, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. xvii. p. 223. Mr. Johnston describes this species as follows : — " Shell white, shining, minute, conical, sub-turreted, of about 6 rapidly increasing ■whorls; whorls convex, ornamented by regular, distant, punctate, spiral sulci, about 12 on last whorl ; under lens broad, distant, obsolete, transverse lirae may be traced ; suture deeply impressed ; columella abbreviate, with two thick, tortuous folds, the lower being the larger ; aperture roundly notched posteriorly and anteriorly ; outer margin thick and reflexed, forming a broad varix continuous with the last tortuous fold of columella." He does not give a figure of the shell, but the description agrees generally with specimens in the Museum. It may be added that beyond the two columellar folds alluded to, a third one exists posteriorly, the direction of which is approximately at right angles to the others, and, as seen in perfect specimens, it resembles a " tooth." Some specimens from Muddy Creek are but veiy faintly striated, the columellar callosity being thick and partly spread over the body-whorl, from which it is sharply defined. The protoconch of those from the locality mentioned is very small, composed of a turn and a half, smooth and somewhat obtuse. R. lactea may be distinguished from the living R. denticulata, Gould, of Port Jackson and elsewhere in Australia, in that the latter is more tumid, the anterior canal is more deeply cut, the columellar callosity being much drawn out, whilst the spiral lineations are closer together. Dimensions. — Length 3^ mm. ; breadth 2 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9304. Three specimens. Ftirchased. LIMACINID^. 17 Family LIMACINID^. The anatomy of the soft parts of Pteeopoda has led Dr. Paul Pelseueer to the conclusion ^ that pteropods are not entitled to be regarded as a class of mollusca of the same value as the Cephalopoda, Gasteeopoba, Scaphopoda, and LAMELLiBEAifCHiATA. Pteropods, he believes, are not primitive mollusca, but are a derived and recent group, and he completely abstracts certain Primary fossils usually referred to the Pteeopoda from that group, as represented by living forms. Boas had arrived- at a similar con- clusion, though not on such abundant material. Dr. Pelseneer discusses the matter at some length {op. cit. p. 90), and expresses the opinion that real pteropods had no existence as such until the commencement of the Tertiary epoch. He bases his arguments on comparative anatomy, embryology, and palaeontology, in that section dealing with the origin and phylogeny of the whole group. He finds that the pteropods are Gasteeopoda in which the adap- tation to pelagic life has so modified their external characters as to give them an apparent symmetry ; that they do not constitute a distinct subclass, nor even an order. They belong to the Tectibranchiate Opisthobranchs, and they differ less from Tecti- branchs than these differ from the other Opisthobranchs. And, he is of opinion that the Thecosomata to which the Australian fossils presently to be described may be referred, have descended from the Bdlloidea ( = Cephalaspidea). The general classification of the Pteeopoda amongst Opistho- branchiate mollusca is here adopted, but without prejudice as to the disposal of pre-Tertiary fossils usually relegated to the group, the consideration of which does not fall within the scope of the present work. It is not a little remarkable as a phase in the evolution of the Pteeopoda, that immediately the Thecosomata as a group was well established, members of it became extremely numerous in the Lower Tertiaries of Australia. But the assemblage of genera is not such as might have been expected in rocks as old as the Eocene, and it is highly probable that, regarded ' "Challenger" Reports — Zoology, vol. xxiii.: Pteropoda, part 3: Anatomj-, p. 96. ^ Spolia Atlantica, pp. 94, 95. 1! from the evolutionary standpoint of the Pteropoda alone, the Australian " Eocene," as exemplified by the Lower beds of Muddy Creek, Schnapper Point, etc., is younger than the Eocene of typical districts like the London, Hampshire, and Paris basins. A very common genus, for instance, in the Australian beds mentioned is Vaginella, which, as will be seen hereafter, has a distinct Miocene facies, and it is found in typical beds of that age in Europe and America. Stijliola, again (when rightly interpreted), is rather a Miocene than an Eocene subgenus ; as for Spirialis, its range in time, from the Middle Eocene to the present duy, imparts no evidence of homotaxial value. It will be observed that Styliola is here included as a subgenus of Clio ; neither its embryonic nor its structural differences entitle it to rank as a genus. Its structural features go to show that Styliola is much more nearly related to Clio (^sensu stricto) than to Creseis ; the forms included under the latter designation have retained some of the more archaic characters of the LiMACiNiDiE. Genus LIMACINA, Lamarck. [Anim. sans Vert. t. vi. pars 1, 1819, p. 291.] Seterofusus, Fleming, Mem. "Wernerian Soc. Edin. vol. iv. 1822, p. 498. 8piratella, Blainville, Diet. Sci. Nat. t. xxxii. 1824, p. 284. Heliconoides, D'Orbigny, Voy. Amerique Meridionale, t. v. 1836, p. 174. Spirialis, Eydoux and Souleyet, Eevue Zoologique, t. iii. 1840, p. 235. Helicophora, Gray, Synopsis of the Contents of the Brit. Mus. 1842, p. 59. Sca:a, Philippi, Fauna Moll. Sicilias, 1844, p. 164. Protomedea, G. 0. Costa, Microdoride Mediterranea, 1861, p. 73. JEmholus, Jeffreys, British Conchology, vol. v. 1869, p. 114. " Shell umbilicate, with turns gradually increasing ; with a fairly large aperture ; and with a columella not prolonged into a rostrum ; surface smooth or striated. The height of the spire, the form of the surface and that of the aperture, and the size of the umbilicus, vary according to the species." — Pelseneer. Limacina is universally regarded as a genus established by 19 Cuvier, but on reference to the French savant's work ' it will be noticed that he refers merely to " Les Limacines"; the author who really established the genus was Lamarck, as above quoted. The last-mentioned author, however, did not give the name of the species correctly. Tijpe. — Limacina helicialis (= Clio heliciyia, Phipps). Limacina tertiaria, Tate (sp.). 1887. Spirnli.i tertiaria, Tate, Traas. Eoy. Soc. South Aust. vol. ix. p. 196, pi. XX. figs. I2a-e. 1889. Spiralis tertiaria, Dennant, id. vol. xi. p. 48. Shell umbilicated, thin, and shining ; spire depressed to the level of the body- whorl, which is greatly expanded; the last whorl rapidly enlarges, is regularly curved, except near the suture, where it is rapidly descending ; umbilicus small, but deep ; the peristome is slightly reflected in its vicinity ; surface of shell ornamented with fine lines. Australian authors include this species in a genus Spiralis ; it is presumed that Spirialis is intended. Dr. Pelseneer divides - Spirialis, Eydoux and Souleyet, into two groups, according to the presence or absence of an umbilicus, and other characters. The species above described, following that classification, falls into the genus Limacina. Dimensions. — Diameter about 1 mm. Form. a}i(l Loc. — Eocene : Muddy Creek, Victoria. G. 9308. Three specimens. Purchased. Family CAVOLINIID^. Genus CLIO, Linnaeus. [Syst. Nat. eil. 12, 1767, vol. i. pt. 2, p. 109-1 ; tion 0. F. Miiller, 1776.] Cleodora, Pi'ron and Lesueur, Ann. Mus. Hist. Nat. Paris, t. xv. 1810, p. 66. Balantium, Eenson, Journ. Asiat. Soc. Bengal, vol. vi. 1837, p. 151. ' Eegne Animal, t. ii. 1817, p. 380. - Op. cit. pp. 15, 32. 20 Shell some'n'hat angular, compressed dorso-ventrally, with lateral keels. An anterior transverse section is thus always angular laterally, A crest or rib generally extends longitudinally along the back, and usually projects. The embryonic shell is variable in form, but may always be distinctly separated. Type. — Clio pyramidata, Linn. Subgenus STYLIOLA, Lesueur. \Ih Blainville, Manuel de Malacologie, 1825, p. 655.] Shell conical, straight, elongated ; surface smooth or faintly striated ; dorsal groove not parallel to axis of shell, but slightly oblique, with only the anterior extremity (which ends in a rostrum) in the median line ; embryonic portion ends in a pointed apex. Type. — Styliola recta, Lesueur (MS.) = Cleodora suhula, Quoy and Gaimard. Clio (Styliola) rangiana, Tate (sp.). 1887. Styliola rangiana, Tate, Trans. Eoy. Soc. South Aust. vol. ix. p. 194, pi. XX. fig. 2. 18S9. Styliola rangiana, Dennant, id. vol. xi. p. 48. Its rounded sides and dorsal groove serve to distinguish this species from C. licarinata, Tate, found in the same beds. Near the summit of the shell is a conspicuous inflation, whilst the apex is much swollen and abruptly tapers to a fine point. Pro- fessor Tate's figure barely does justice to this structure, and it is interesting enough to demand another figure, which is now given (Plate I. Figs. 6«-5) as seen under a microscope with a one-inch objective. The phenomena in the neighbourhood of the apex recall the genus JEucMlotheca, Fischer, to some extent, but many fundamental differences exist. In particular, C. rangiana has no reversed or collareted border round the aperture ; whilst its pronounced longitudinal sulcus, extending from the aperture almost to the apex, is very distinctive. The whole shell is somewhat compressed dorso-ventrally, and its almost smooth and polished surface is ornamented with oblique transverse striations. Professor Tate remarks that, compared with shells of the living VAGTNELLA. 21 Cleodora siihcla, this species is distinguished by its elliptical aperture, proportionately greater breadth, and by its abruptly tapering apex. Dimensions. — Length 5 mm.; diameter of aperture about Imiu. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9306. Three specimens. Purchased. Genus VAGINELLA, Daudin. [Bull, des Sciences (Soc. Philom.) Paris, t. ii. No. 43, 1800, p. 145, pi. xi. fig. I.] Shell long, ventricose, depressed ; apex sharp-pointed, con- stricted in front ; aperture slightly canaliculated and compressed laterally. The occurrence of this genus in Australia is very remarkable, the more so that it is found in beds of Eocene age, whereas in Europe it is characteristic of the Upper Tertiaries, especially tlie Langhian division of the Miocene. Type. — Vaginella depressa, Daudin. Vaginella eligmostoma, Tate. 1887. Vaginella eligmostoma, Tate, Trans. Roy. Soc. South Aust. vol. ix. p. 195, pi. xs. fig. 7. 1889. Vaginella eligmostoma, Dennant, id. vol. xi. p. 48. 1893. Vaginella eligmostoma, Tate and Dennant, id. vol. xvii. p. 223. This species, which is very common in beds near the base of the section at Muddy Creek, has been aptly compared by Professor Tate to V. depressa, Basterot. The present writer, who has found large numbers of the latter species at Moulin de Lagus, the Coquilliere at Leognan, and other localities near Saucats, south of Bordeaux, is enabled to state from comparison of abundant material of both the Australian and European species mentioned, that V. eligmostoma is narrower, longer, more depressed in pro- portion to its size, and more prominently constricted beneath the expansions of the aperture. The representation of V. depressa as having simple margins meeting at an angle on either side of the aperture is not borne out by the facts, though the margins are not sinuated to the same extent as in the Australian species. The 22 TJMBEACULtTM. author of the latter remarks that the shell is angulated at the sides and sculptured by very fine arched striae descending to the lateral angulations. "The aperture is narrowly oval-oblong, with trun- catedly rounded ends ; each lip has a median curvature bounded on each side by a broad insinuation." The apex is in nearly all cases well preserved. The shell is very variable in length, some specimens being much narrower in proportion than others ; such differences may be adjusted, however, in reviewing a large number, and are not of specific value. Dimensions. — Length 7 mm.; breadth about Hmm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9310. Several specimens. Purchased. Family TJMBRACULIDJE. Genus UMBRACULUM, Schumacher. [EssaiNouv. Syst. Habit. Testaces, 1817, p. 177.] " Ombrelle," Lamarck, Cours Zool. 1812, p. 114. Umhrella, Lamarck, Anim. sans Vert. t. vi. 1819, p. 339. Gastroplax, Blainville, Diet. Sci. Nat. t. xviii. 1820, p. 176. Omhrella, Blainville, Diet. Sci. Nat. t. xxxii. 1824, p. 267. Umbella, D'Orbigny, Moll. Cuba, t. i. 1841, p. 115. Operculatum, H. and A. Adams, Gen. Bee. Moll. vol. ii. 1854, p. 41. Shell orbicular, broad, patelliform, apex very small ; ornamented exteriorly with concentric lines of growth ; internal surface with concentrically undulating striae radiating from the centre to the circumference of the shell, but more prominent towards the centre. The genus is generally called Umhrella, but, as the synonymic references above indicate, Umhraculum has priority. The designa- tion "Ombrelle," if it were not a mere list name, could not stand, in consequence of its being French, and not Latin. The term Operculatum created by Linnaeus ^ and brought on by the brothers Adams cannot be accepted, first because it was not made in a binomial sense, and secondly as it was proposed prior to the date 1 Mus. Tess. 1753, p. 116. TJMBEACULUir. 23 of the tenth edition of Linnaeus' " Systema Naturae." Acardo,^ commonly quoted as being synonymous with Umhraculum, is said by its author to be a bivalve ; from the figure given it might be the epiphysis of a whale's vertebra. M. Cossmann^ states that Vmbrella has priority over Umhraculum, as it was proposed by Chemnitz in 1788, whereas the latter dates from 1817. On referring to the joint work of Martini and Chemnitz that observation is hardly borne out. Martini^ alludes to "Umbella," but in his description^ he is clearly not binomial, and did not intend to name a genus Umhella ; he rather calls it Lepas, as " Lepas vertice et limbo,'" etc. And a few pages farther on' the same kind of thing is described as ^^ Lepas Indiea vertice,''^ etc. Subsequently Chemnitz^ refers to " Umhella Chinensis, testa In- tegra," etc. ; though in the explanation of the plates the term is abbreviated to " Umhella Chinensis." As these authors were not binomial, their work must be ignored; and Umhraculum should be adopted. Type. — Umhella chinensis, Schumacher {ex Martini). Umbraculum australe, sp. nov. [Plate I. Figs, la-c.'] ? Umbrella austrina, Tate, MS. (list name). Shell broad, moderately elevated ; small but prominent protoconch, which is slightly oblique ; lines of growth well marked on both the outer and inner surfaces ; two inconspicuous sulcations diverge from the protoconch anteriorly at an angle of about 35° ; inner surface of the shell smooth and shining, radially striated, orbicular muscle marks large and rather depressed, causing corrugations ; margin thin and rough. * Commer9on MS., Bniguiere, Ency. Meth. (Vers) t. i. 17S9, p. 1 ; cf. also Lamarck, Mem. Soc. Hist. Nat. Paris, 1798, p. 81 ; id. Syst. Anim. sans Vert. 1801, p. 130. 2 Ess. Paleoconeh. Comp. 1895, p. 131. 5 Conch. Cab. Bd. i. 1769, p. 90, pi. vii. fig. 49 ; also pi. vi. fig. 41. * Op. cit. p. 103. « Op. cit. p. 106. 6 Op. cit. Bd. s. 1788, p. 341, pi. clxix. figs. 1645-6. 24 It is interesting to find this genus in the fossil state. Compared •with the living U. mediterranea, Lamarck, the Australian shell is rather more elevated, is not so elliptical, and the inner radiating striae are not as well pronounced in the neighbourhood of the muscular impressions. Dimensions. — Height 10 mm.; maximum breadth 33 mm.; minimum breadth 29 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4183. One specimen. Presented hij John Bennant, Esq. Order PROSOBRANOHIATA. Family TEREBRID^. Genus TEREBRA, Lamarck. [Mem. Soc. Hist. Nat. Paris, 1799, p. 71.] Shell subulate, many-whorled, with a short anterior canal and smooth columella. This genus has been divided into many sections, but the writer agrees with Tryon^ that it cannot be advantageously dealt with in that manner, though an exception may be permitted in the case of the subgenus JEuryta. Type. — Buccinum suhulalum, Linnaeus. Terebra catenifera, Tate. 1886. Terehra catenifera, Tate, Southern Science Record, January, p. 5. 1889. Terehra catenifera, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 160, pi. viii. fig. 14. Shell pyramidal, whorls flattened convexly ; " slightly over- lapping ; double-banded and nodulose in front of the suture, the posterior band rather the broader, and separated by a shallow sulcus, in the centre of which winds a subangular ridge defined by linear grooves; about 20 pairs of nodulations on the penultimate whorl ; anterior half of each whorl distantly and superficially ' Manual of Conchology, vol. vii. 1885, p. 7. 25 spirally ridged ; the Tvhole surface arcuately striated by lines of growth; base spirally ridged and transversely wrinkled." — Tate. It may be compared with the living T. mariesi, E. A. Smith, of Japan. Dimemiom. — Length 43 mm.; breadth 7 mm. Form, and Loc. — Miocene : Muddy Creek, Victoria. G. 4229. Five specimens. Prefiented hj John Dennant, Esq. Terebra platyspira, Tate. 1886. Terebra platyspira, Tate, Southern Science Eecord, January, p. 6. 1889. Terebra platijspira, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 159, pi. viii. fig. 12. 1893. Terebra platyspira, Tate and Dennant, id. vol. x\\i. pt. 1, p. 221. Neither of the specimens in the Museum presents a perfect protoconch, but Professor Tate remarks that it consists of " a relatively large papillary pullus of two turns ; the first turn of the pullus subglobose, with the tip reverted and immersed, much broader than the next two or three whorls." The sutures of the shell are slightly canaliculated and distantly bordered by a linear sulcation ; the whorls are ornamented with transverse curvilinear ridges and a few indistinct spiral striae most conspicuously de- veloped in adult specimens. This species has considerable analogy with certain European Miocene forms of Terehra — in particular T. perhisa, Lamarck, from the Langhian beds south of Bordeaux. It is narrower than that species, however, and the whorls are flatter and more turri- culate. It is allied also to the living T. cingulifera, Lamarck, from the Philippines, China, etc. Dimetmons. — Length 40 mm. ; breadth 5 '5 mm. Professor Tate states that the largest examples attain to 60 mm. in length. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4230. Two specimens. Presented hj John Dennant, Esq. Terebra additoides, Tenison-Woods. 1877. Terebra additoides, Tenison- Woods, Pap. Roy. Soc. Tas. 1876, p. 95. 1889. Terebra additoides, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 163. 1893. Tcreha additoides, Tate and Dennant, id. vol. xvii. pt. 1, p. 221. 26 The specimen in the Museum is too mutilated for minute description. The posterior portion of the later whorls have a narrow ante-sutural band ; the plications are narrow, slightly bent, and subnodose, and between them are very fine and close reticulations. Dimensions. — Length about 9 mm. ; breadth 3 mm. Form, and Loc. — Eocene : Table Cape, Tasmania. 83991. One specimen. Purchased. Terebra ustulata, Deshayes. 1857. Terehra ustulata, Deshayes, Journ. Concliyl. p. 97, pi. iii. fig. 12. 1859. Terehra ustulata, Deshayes, Proc. Zool. Soc. p. 294. The sutural band is not so deeply impressed as in typical specimens in the Museum from Tasmania, as found living, whilst the body-whorl is more convex ; these differences may constitute a variety from which the living forms may have descended, but they are not worthy of specific rank. Dimensions. — Length 33 mm. ; breadth 9 mm. Form, and Loc. — Post- Pliocene : Limestone Creek, Glenelg river, Victoria. G. 5555. Three specimens. Purchased. Terebra geniculata, Tate. 1886. Terebra geniculata, Tate, Southern Science Record, January, p. 6. 1889. Terebra geniculata, Tate, Trans. Roy. Soc. South Aust. vol xi. p. 161, pi. IX. fig. 8. The protoconch (Plate II. Figs. \a-l) of this species is subacute and composed of three convex turns. The whorls of the shell are much constricted posteriorly, and between the constriction and the suture there is a row of tubercles ; the remainder of the surface of the whorls has distant rugose ribs which are angulate, and the whole presents rather the aspect of certain forms of Drillia than of Terehra. That it ought not to be classified with the Pleueoxomid.^, however, is clear from the general phenomena of the aperture. It diff'ers from T. mutica, Tate, from the same locality but of 27 Eocene age, in the whorls being more angulate, the anterior canal being more excavated, and in other minor features. Dimensions. — Length 9 mm ; breadth 3 mm. Form, and Loc. — Miocene : Muddy Creek, Victoria. G. 9309. Two specimens. Family CONID..S. Genus CONUS, Linnaeus. [Syst. Nat. ed. 10, 1758, p. 712.] Shell convolute, turbinate; aperture narrow and long, edentulous ; columella smooth. An attempt to divide the Eocene fossils of this genus into subgeneric groups based on observations made on living examples of Conus, must always, the writer believes, be attended with great uncertainty: for the genus itself had only just been established; and although in a very unstable condition, it had not yet had full opportunity of modifying, with development in diiierent directions. No doubt, to a large extent, certain features of modern groups of Cones were conceived in Eocene times, but they were not sufficiently well differentiated to enable the species to be allocated clearly to the subgenera as established on living species. Conus was pregnant with evolutionary possibilities in the early Eocene. Nevertheless, the writer has essayed a subdivision in spite of the difficulties and uncertainties existing. Ti/pe. — Conus marmoreus, Linnaeus. Subgenus LEPTOCONUS, Swainson. [Malacology, 1840, p. 312.] The author describes Leptoconus as follows : — " Shell light, conic, sometimes striated ; spire elevated, acute, concave ; the basal whorl carinated, detached, and sinuated above, and con- tracted near the suture." Other characters which seem to be fairly persistent in this group are the deeply-cut canal bordering the suture and the more or less staged convolutions. Type. — Conus grandis, G. B. Sowerby. 28 Conus (Leptoconus) ligatus, Tate. 1890. Comes ligatus, Tate, Trans. Eoy. Soc. South Aust. vol. xiii. pt. 2, p. 196, pi. ■vii. figs. 4, \a-b; pi. viii. fig. 9. 1893. Conus ligatus, Tate and Dennant, id. vol. xvii. pt. 1, p. 221. Protoconch (Plate II. Pigs. 2a-l) composed of two smooth volations, the earlier of ■which is somewhat oblique and inflated. The spire of the shell proper is of variable height, in some specimens being comparatively depressed and in others so elongated as to be one-third the length of the shell. Spire-whorls with an obtuse, thick shoulder and ornamented with closely-set, spiral striae, which persist over the whole surface of the body-whorl also, being especially accentuated on the anterior half. The body-whorl varies in shape from pyriform to elongate ovate, the peripheral keel defined anteriorly by a linear constriction, posterior to which there are one or two threads on the peripheral band. The aperture is narrow, with a comparatively shallow anal sinus. The range of variation in this species is so great that innumerable "varieties" could be established. One of them, as pointed out by Professor Tate, closely resembles C. deperditus, Bruguiere, of the Eocene of North-western Europe. This species partakes of the characters of both Leptoconus and Lilhoconus ; it has been assigned to the former subgenus on account of its elevated and staged spire and striated surface. Dimensions. — Length 40 mm. ; breadth 2r5 mm. Form, and Loc. — Eocene : Victoria. 48052. Several specimens, in various stages of growth ; from Schnapper Point. Purchased. 70424. Pour specimens ; from Meribee Plains. Purchased. G. 4242. Two specimens ; from Muddy Creek. Presented hxj John Dennant., Esq. G. 5490. One specimen ; from Muddy Creek. Purchased. Conus (Leptoconus) heterospira, Tate. 1890. Conus heterospira, Tate, Trans. Eoy. Soc. South Aust. vol. xiii. pt. 2, p. 197, pi. vii. figs. 6, 5a. 1893. Conus heterospira, Tate and Dennant, id. vol. xvii. pt. 1, p. 221. 29 This species differs from C. ligatus in being more elongate, ■whilst it is minus the rim-like keel of that species, and the lineatious on the whorls of the spire are not conspicuous. The staging of the whorls is a peculiar feature ; when in the brephic and neanic periods of growth there is little or no attempt at staging, but in the adult, and especially in senile forms, the anterior whorls drop very sharply. It is highly probable that this catabatic phenomenon is the precursor of the establishment of an elevated spire in its descendants ; but that is a point which cannot be definitely settled until the phylogeny of the Gasteropoda has been worked out. Dimensions.— \iGii^th 31 mm. ; breadth 15 mm. Form, and Loc. — Eocene : Victoria. 48052. A series of ten specimens ranging from the neauic to the catabatic stages ; fi'om Schnapper Point. Furchased. 70423. Four specimens ; from Meribee Plains. Furchased. Conus (Leptoconus) ne^wtoiii, sp. nov. [Plate II. Figs. ?,(i-d.'] Protoconch small, composed of two and a half smooth turns, indistinctly striated with growth -lines anteriorly ; posterior portion slightly oblique with reference to the axis of the shell. Spire elevated, acute, convex, about one- sixth the total length of the shell ; spire-whorls seven in number, staged, excavated, spiral lineatious bold and continuous, growth-lines prominent, sinuated. Keel sharp and plain, not being crenulated even in the brephic stage. Body-whorl elongate, ornamented throughout its length by distant spiral sulcations which are punctated where crossed by the lines of growth. Aperture long and narrow ; outer margin thin and arched ; posterior sinus very deep ; anteriorly the aperture is somewhat dilated and the columella is slightly twisted. The elongation and narrowness of the body-whorl are dis- tinctive features. Its nearest ally in the Australian Tertiaries is C. extenuatus, Tate, which, however, is broader, whilst its body-whorl is not so long, its periphery is not so sharply keeled, the earlier whorls are crenulated, and the ornamentation of the spire differs. C. newtoni closely resembles C. sieholdi, Reeve, 30 living in the seas round the coasts of China and Japan, but in the modern form the whorls are not so much staged and the spiral lineations are not so well marked ; the principal point of difference, however, is the shape and shallowness of the posterior sinus. Dimensions. — Length 62 mm.; breadth 26 mm.; length of aperture 52 mm. Form, and Loc. — Eocene (?) : Muddy Creek, Victoria. G. 4239. One example. Prese7ited hj John Dennant, Esq. G. 5494. One example. Purchased. Conus (Leptoconus) murravianus, Tate. 1890. Conns mnrravianiis, Tate, Trans. Eoy. Soc. South Aust. vol. xiii. pt. 2, p. 200, pi. vii. fig. 2. Protoconch (Plate II. Pigs. 4a^b) composed of three smooth turns implanted obliquely with reference to the axis of succeeding whorls; the initial portion is exceedingly small in comparison with the anterior turns. These latter are depressed and deeply canaliculated. The shell proper has an elevated spire with six whorls, which are somewhat staged, slightly excavated ; and spiral lineations are not prominent, except towards the anterior half of the body-whorl. It is noteworthy that in the brephic stage the periphery of the body-whorl was obscurely crenulated, a character which disappeared, however, as the neanic period was reached. Aperture narrow, slightly expanded in front ; outer margin thin ; with a shallow posterior sinus. This species presents characters of no less than three subgenera : by its very elevated spire and the obscure crenulations it recalls Conospirus, which does not appear to stand on a very firm basis ; its protoconch presents the features of Lithooonus as represented in the Australian Tertiary by C. dennanti and C. piillidescens ; but the general contour of the shell and the staged whorls reflect Leptoconus, to which subgenus the species is here referred. Specifically it has a near ally in C. hrocchii, Bronn, of the Italian Pliocene. Dimensions. — Length 41mm.; breadth 16 mm. Professor Tate records a length of 61mm. Form, and Loc. — Eocene : Eiver Murray clifPs, G. 9136. One specimen. Presented ly William Evans, Esq^. 31 Conus (Leptoconus) convexus, sp. nov. [Plate II. Figs, oa-d.'] Protoconcli composed of two smooth turns, the initial portion of ■which is lateral and immersed ; much elevated above the remainder of the shell. The whorls of the spire, five in number, are slightly convex, and are thus not excavated as in other species mentioned ; they are ornamented by five or six deeply-cut sulci, somewhat irregular in places owing to the intersection of growth marks ; these latter, however, do not approach tessellation. The whorls are not staged, but the suture is distinctly canaliculated, especially as the growth approaches the ephebic stage. The body-whorl has an elevated band at its periphery, which is rugosely lineated ; its whole surface is covered by small, undulating, irregular sulci, which broaden anteriorly and are interrupted by sinuous growth- lines. Aperture long, narrow, its two margins parallel with each other ; outer margin thin, curved, and distinctly crenulated within ; anal sinus broad and comparatively shallow, inner margin straight. Its protoconch is very different to that of C. pulhdescens ; and the convexity of the whorls, deep sulci, canaliculate suture, and crenulated margin of the aperture, are highly distinctive. On comparison with C. complicatus, Tate, in addition to the characters just mentioned, it will be observed that C. convexus is broader, and the spire less elevated, whilst the ornament is peculiar to it ; moreover, the whorls are not staged, neither is the sinus deeply, arcuately notched. Dimensions (of the type specimen). — Length 17 mm.; breadth 9 mm. Form, and Loc. — Eocene : Meribee Plains, Victoria. 70424. Two specimens. Purchased. Subgenus LITHOCOXUS, Morch. [Cat. Yoldi, Fasc. i. 1832, p. 66.] Shell conical, spire more or less elevated, but often depressed, not coronate nor staged ; body-whorl sharply carinate ; aperture dilated in front, with a rather deep posterior sinus. Type. — Conus milkpunctatus, Linnxus. 32 Conus (Lithoconus) cuspidatus, Tate. 1890. Conus cuspidatus, Tate, Trans. Eoy. Soc. Soutti Aust. vol. siii. pt. 2, p. 194, pi. vii. fig. 1. 1893. Conus cuspidatus, Tate and Deunant, id. vol. svii. pt. 1, p. 221. Protoconch (Plate IT. Figs. 6a-b) conspicuously elevated, rising to a sharp point, composed of four to five slowly and regularly increasing turns, forming a very distinctive feature. Spire of the shell proper with seven to eight volutions, slightly concave and deeply lineated spirally. In the brephic stage the spire is much elevated in all the specimens examined, and this character may he perpetuated to the gerontic stage, or the whorls may become flatter and less elevated during the neanic, and absolutely depressed during the later stages of growth. Such a wide range in elevation of the spire is not often observable in Conus ; the amount of elevation is merely an individual peculiarity. This may be noted by those who seize upon this unstable property as a subgeneric character and often allow it to influence them in defining a species. Body-whorl either obtusely or acutely angulate at the periphery, depending on the elevation of the spire ; lines of growth well marked, spiral striae plainly developed anteriorly. Aperture narrow, dilated in front, channelled behind ; columella twisted, bordered by an elevated ridge in young specimens, and very pro- nounced in the catabatic stage where the anterior portion is usually much fractured, owing apparently to difficulties of growth. The columellar twist leads to a slight constriction of the body-whorl. Dimensions. — Length 49 mm.; breadth 25 mm. Form, and Loc. — Eocene : Muddy Creek, Yictoria. G. 5489. Three specimens. Purchased, Conus (Lithoconus) pullulescens, Tenison-Woods. 1880. Conus pullulescens, Tenison-Woods, Proc. Linn. Soc. N.S.W. vol. iv. p. 3, pi. i. tig. 4 {non fig. 3). 1890. Conus pullulescens, Tate, Trans. Eoy. Soc. South Aust. vol. xiii. pt. 2, p. 190, pi. xi. fig. 9. 1893. Conus pullulescens, Tate and Dennant, id. vol. xvii. pt. 1, p. 221. This species may be distinguished from C. cuspidatus, amongst other things by its extremely characteristic and prominent proto- conch (Plate 11. Pigs. la-h). This is very large, composed of 33 three turns, somewhat turbinate, with deeply canaliculate sutures. The protoconch is oblique with reference to the axis of the shell. The spire of the shell pi'oper is not much elevated, its whorls are narrow, separated by a linear suture, ornamented with a few spiral threads and closely-set sinuous lines of growth. Body- whorl sharply keeled at the periphery, and covered with fiat spiral threads which become obsolete with age, except anteriorly. The figure accompanying Mr. Tenison-Wood's memoir above cited refers to a young shell, and is practically unrecognizable. The interpretation placed upon it by Professor Tate is here adopted. Dimensions. — Length 39 mm.; breadth 21mm. Form, and Loc. — Eocene : Victoria. 70423. Two specimens ; from Meribee Plains. Purchased. 73220. Three specimens ; from Schnapper Point. Purchased. Or. 4240. One specimen ; from Muddy Creek. Presented hy John Dennant, Esq. G. 5491. One specimen ; from Muddy Creek. Purchased. G. 4745. Two specimens ; from Muddy Creek. Presented hy Prof. T. Rupert Jones, F.R.S. Conus (Lithoconus) dennanti, Tate. 1890. Conus dennanti, Tate, Trans. Eoy. Soc. South Aust. vol. siii. pt. 2, pi. xi. fig. 7 (plate issued July, 1892). 1893. Conus dennanti, Tate and Dennant, id. vol. xvii. pt. 1, p. 221 (list name) . This species does not appear to be described, except in the explanation of the plate above cited, where the following words occur: " Conus dennanti, sp. nov. {C. piillulescens, var., Tate)." The protoconch (Plate II. Figs. 8«-5) differs from that of C. piillulescens in not being angulate, or turbinate ; it is com- posed of three smooth, convex turns, imbedded in the crown of the shell, being in fact the only part whicli stands above the flat surface produced by the convolution of the shell-whorls proper. This upper surface is much depressed, has linear sutures, and is boldly, spirally striated. The periphery of the body-whorl is more sharply keeled than any other Australian Tertiary Conus; the surface of the whorl is covered by rather closely-set spiral linea- tions and sinuous growth-lines. Aperture very narrow, outer 34 border parallel with the inner, but slightly dilated anteriorly ; columella twisted in front. Dimensions. — Length 33 mm. ; breadth 20 mm. Form, and Loc. — Eocene : Victoria. 70423. One specimen ; from Meribee Plains. Purchased. G. 5493. Two specimens ; from Muddy Creek. Purchased. Conus (Litliocoiius) ptychodermis, Tate. 1890. Conus ptychodermis, Tate, Trans. Roy. Soc. South Aust. vol. xiii. pt. 2, p. 196, pi. vli. fig. 3. This species may be readily distinguished from all other Australian Tertiary Conus, by the peculiar, rugged aspect of the body-whorl caused by the inter-crossing of growth folds and the irregular character of the distant spiral lineations and ridges. The spire is much depressed, with canaliculated sutures; the periphery of the body- whorl is sharply keeled ; the anterior portion of the columella is twisted and prominently lineated. The protoconch is not well preserved in the Museum specimen, but it evidently had the general features of that of C. pullulescens, though more Dimensions. — Length 20 mm. ; breadth 9 mm. ; length of aperture 17mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 5489. One specimen. Purchased. Subgenus CHELYCONUS, Morch. [Cat. Yoldi, Fasc. i. 1852, p. 69.] Spire elevated, last whorl convex near suture, rounded at the shoulder, posterior sinus not very deep. Type. — Conus testudinarius, Gmelin {ex Martini). Conus (Chelyconus) ralphi, Tenison- Woods. 1879. Comis ralphii, Tenison- Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 228, pi. xxi. fig. 14. 1890. Conus ralphii, Tate, Trans. Eoy. Soc. South Aust. vol. xiii. pt. 2, p. 198, pi. vli. fig. 6. 35 Protoconch (Plate IT. Figs. 9a-h) conical, moderately elevated and composed of two smooth turns, somewhat similar to that of C. heterospira. In the brephic stage, and, possibly, to the commencement of the neanic, the whorls of the spire are coronate, but as the adult form is reached these disappear and the shoulders become rounded. Mr. Tenison-Woods was only acquainted, ap- parently, with young forms of the species, and the figure in his memoir cited above does not convey a correct idea of the shell, as each whorl is represented as being carinate and nodosely crenulated. Professor Tate has traced the young forms to the adult and firmly established the species. The spire is elevated, composed of eight whorls striated spirally above. The body-whorl is also spirally striated, the striae being most conspicuous on the anterior half. Aperture narrow, posterior sinus shallow and following growth- lines ; columella slightly twisted anteriorly. Dimensions — Length 43 mm. ; breadth 23 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 5492. One specimen. Purchased. G. 4241. One specimen. Presented hj John Dennant, Esq. Conus (Chelyconus) catus, Hwass. 1792. Conus catiis, Hwass, Eney. Meth. (Vers) t. i. pt. 2, p. 707. Shell bulbous, spire convex, striated above ; body-whorl finely striated on the posterior half, and having deep spiral sulcations anteriorly. Aperture narrow, but widening in front and some- what dilated ; posterior sinus conspicuous ; columella twisted anteriorly where a well-marked ridge or fold is produced. The species differs from the earlier C. raJphi, with which it is closely related, in being much broader anteriorly and by the prominent coluraellar fold. The specimens in the Museum are not well enough preserved to show the protoconch or the condition of the shell in the brephic stage. This species has living representatives over a large geographical area, including the Red Sea, Mauritius, Java, China, Xew Cale- donia, and Polynesia generally, and it is interesting to find it as a fossil. Dimensions. — Length 44 mm. ; breadth 24 ram. 36 CONUS. Form, and Loc. — Pliocene : obtained from a well-cTigging 10 feet in depth, Chatliani Islands. G. 9502. Two specimens. Transferred from the Museum of Practical Geology. Conus, sp. The following specimens, for the most part casts, are indeter- minable specifically, but are interesting to record as establishing the marine character of Tertiary deposits in certain parts of Australia, and are worthy of notice also on other grounds : — G. 9499. Five casts of a species allied to C. pullulescens, but larger, the body-whorl being more convex; the columella is distinctly twisted. From NuUarbor Plains, South Australia. R. Y. L. Brown Coll. G. 9500. Cast of a species which may be specifically identical ■with the preceding. From Fowler's Bay District, South Australia. S. Y. L. Brown Coll. G. 9152. Cast imbedded in a mass of fragments of Brtozoa. From near Border Town, South Australia. Presented ly R. Littotv, Esq., F.G.8. G. 353. Cast of a species allied to C. murravianus in having an elongated spire ; but it is relatively broad, and the inner border of the aperture is curved. From Bairnsdale, Victoria. Presented hy W. H. Grigson, Esq. G. 9501. Cast imbedded in ferruginous sandstone. From Flemington, near Melbourne. Transferred from the Museum of Practical Geology. G. 9493. Two casts, from the Miocene " Cow?«-beds" of Mokihinui river, New Zealand. Sir James Hector Coll. G. 9494. Fragment of a species composed of six elevated ■whorls, having a protoconch composed of three smooth turns implanted obliquely on the spire. It is closely allied to and may be specifically identical -with C. trailU, Hutton. From Awamoa, i^ew Zealand. Sir James Hector Coll. G. 4237. A new species of Conus, too imperfect for detailed description. It is 60 mm. in length, with a rather elevated spire and elongate body-whorl ; anterior of the columella twisted ; PLEUEOTOJIID.E. 37 aperture dilated in front. It may doubtless be referred to the subgenus Leptoconns. From the Tertiary of Muddy Creek, Victoria. Fresented btj John Bennant, Enq, Family PLEUROTOMID^. In assigning the species here described to their systematic positions in the Pleueotomid^, it may be remarked that in view of the present unsatisfactory condition of the classification of many of the so-called genera and subgenera in the family, some of the con- clusions anived at must, of necessity, be of a tentative chai'acter. The group has been divided into subfamilies according to the presence or absence of an operculum, and the nature of that when present. The systematic position of many of the species is dependent on the situation of the sinus, its depth, and so forth. The length of the anterior canal, details of ornament, peculiarities of the radula, and the like, have also been permitted to rank high — even as generic characters. The family as a whole is not of very great antiquity, its main features having been carved out in the Eocene ; whilst many of its broad characteristics were not evolved until the Miocene, or early Pliocene. Possibly, no group of the moUusca has been so prolific in yielding so many diversified forms in such a short space of geological time. The variation of individuals in the living as well as in the fossil state is often so wide as to render it impossible to satisfactorily define the limits of range permissible in species. That difficulty, combined with the latitude of the personal equation, has led to the establishment of innumerable so-called species of the Pleurotomid^, which appear to the writer to be founded merely on ontogenetic characters. The problem, already difficult with the living forms, is intensified when the fossils are considered. Here we have no operculum to assist, and many other points utilized in the classification of modern Plkukotomid^ are missing. Shorn of the characters of their opercula, the chief difference between Pleurotoma and Surcula, for example, lies in the form and position of the sinus ; in the former genus it is deeper, and is placed farther away from the suture than in the latter. If these characters were in any way constant they might be rendered useful ; but, as it is, we are 38 PLEUEOTOMi. left without any material conchological differences between the two genera mentioned, and the assignment of any but typical si^ecies to either of them must remain doubtful. It is highly probable, however, that a special study of the protoconchs, and the mode of growth of the shells as found in the fossil condition, will assist in clearing up many difficulties. Very few of the Australian Tertiary species of PLEUEOTOiiin^ have been described ; of these some have never been figured, and the descriptions of the remainder are often so imperfect as to render identification impossible. Several forms new to science are now recorded. Genus PLEUROTOMA, Lamarck. [Mem. Soc. Hist. Nat. Paris, 1799, p. 73.] Shell turriculated, spire long ; collumellar margin smooth ; outer margin of the aperture with a narrow, deep sinus, situated some distance from the suture ; canal long and straight. Type. — Murex habylonius, Linnseus. Pleurotoma murndaliana, Tenison-Woods. 1879. Fleurotoma murndaliana, Tenison-'Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 226, pi. xx. fig. 5. 1889. Pleurotoma murndaliana, Deimant, Trans. Eoy. Soc. South Aust. vol. xi. p. 44. 1893. Pleurotoma murndaliana, Tate and Dennant, id. vol. x\ii. pt. 1, p. 221. Protoconch composed of three elongate turns, the initial portion being slightly inflated, whilst the anterior turns are obtusely carinate. Shell narrow, elongate ; whorls ten in number, slightly convex, and having several bold, irregular, spiral threads or ridges, rather rugose where crossed by growth-lines, and some- what granulated in the neighbourhood of the sinus. Canal long, slender, and twisted ; outer margin serrate, sinus large and deep, and situated some distance from the suture. Dimensions. — Length 47 mm.; breadth 13 5 mm.; length of aperture 20 mm. Form, and Loc. — Eocene : Yictoria. 73232. A variation in which the whorls of the spire carry three acute spiral ridges, or keels, the posterior of which has developed a median sulcus, and is granulose. From Brighton. Purchasid. PLEUEOTOIIA. 89 G. 4232. Two specimens ; from Muddy Creek. Presented hy Johi Dennant, Esq. G. 5502. A very elongate example, from Muddy Creek. Furchased. Pleurotoma septemlirata, sp. nov. [Plate II. Figs. lOa-i.] Shell large, fusiform, tumid ; protoconch small, smooth, devoid of ornament, terminating abruptly at its junction with the shell in the brephic stage ; whorls eleven in number, suture canaliculate and bimarginate, rugosely spirally Urate ; there are seven principal spirally striated lirae on the penultimate whorl in the adult, but at younger stages of growth they are fewer in number, decreasing gradually to four ; the body-whorl is irregularly Urate throughout. Between the large compound liration on the margin of the suture anteriorly, and the elevated ridge denoting the position of the sinus, is a broad, deep sulcation, which, like the sulci between the lirce, is finely spirally striated and interrupted by lines of growth. Aperture large, broad, ovate, contracted anteriorly ; outer margin thin, sulcate within, crenulated or serrated, arcuate ; sinus broad, deep, situated distantly from the suture ; columellar margin smooth, curved, slightly callous in senile individuals ; canal very short and broad. Typically this species has some analogy with P. murndaliana, but is larger, more tumid, has a shorter canal, and is not sub- granose, whilst the deep, broad sulcation in front of the suture is characteristic. Its great range of variation is remarkable, and might at first sight lead to the establishment of several new species. The type specimens, as above indicated, are strongly Urate, but with sufficient material these may be traced almost to the vanishing point, some individuals, indeed, being only very faintly striated, and to all intents and purposes smooth. Pari passu the shell is shorter and more tumid, presenting the aspect of Pusionella, though it could not be classified with that genus, on account of its deep sinus, amongst other things. Dimensions. — Length 60 mm.; breadth 22 mm.; length of aperture 25"5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. 40 PLEUKOTOMA. G. 4231. Typical adult and senile specimens, and an example in which the lirse are not so bold, and the shell is relatively more tumid. Presented hy John Dennant, Esq. G. 4233. Four specimens, in which the ornamentation is much reduced in importance, and numbers of additional spiral lineations are introduced. Presented hy John Pennant, Esq. G. 5501. Two specimens, strongly lirate, but more elongate than typical examples. Purchased. G. 5499. A smooth specimen, in which the lirse have dis- appeared, or are only represented by obsolete lineations, whilst the growth-lines are more prominent. Purchased. Pleurotoma trilirata, sp. nov. [Plate III. Figs. l«-rf.] Shell small, elongate. Protoconch large, composed of two and a half smooth turns ; the earlier portion is much depressed, and even in the initial part the suture is canaliculate and marginate ; at a later stage the protoconch is longitudinally costate and subnodose at the periphery, and this continues up to the brephic stage, where it gives way to the bold spiral lirse which form the sole ornamentation of the whorls in subsequent stages of growth. Whorls six, flat, suture canaliculate and marginate ; spiral lirae three in number on the penultimate whorl, with a smaller lineation between the posterior two, which becomes more accentuated on the body-whorl. The latter is boldly lirated throughout ; lines of growth inconspicuous. Aperture small, pyriform ; outer margin arcuate ; sinus shallow, situated not far from the suture ; columellar margin smooth, not callous, slightly twisted in front ; anterior canal very short. The most interesting feature of this species is the highly sculptured protoconch, which differs from that of any other Australian form here described. The longitudinal costse, how- ever, precede the spiral lirse as in some other species of Pleurotoma. Dimensions. — Length 10-5 mm. ; breadth 4 mm.; length of aperture 3-5 mm. Form, and Loc. — Eocene : Hobson's Bay, Victoria. 48052. The type specimen. Purchased. I PLKUROTOMA. 41 Pleurotoma consutilis, Teiiison-"WoocTs. 1880. Pleurotoma consutilis, Tenison-Woods, Proc. Liun. Soc. X.S.W. vol. iv. p. 9, pi. ii. fig. 5. 1889. Pleurotoma coiisuiilis, Dennant, Trans. Roy. Soc. South Aust. vol. xi. p. 44. 1893. Pleurotoma consutilis, Tate and Dennant, id. vol. x\ii. pt. 1, p. 221. Protoconch composed of one and a half smooth, shining whorls, the anterior portion of which is obtusely keeled medially ; this keel develops into a strong feature in the brephic and neanic stages, but on approaching the adult is not so clearly marked. Shell small, fusiform ; whorls obliquely cancellate, the transverse and longitudinal lines being regularly spaced; aperture broad, outer margin much produced towards the middle, sinus broad, deep, and some distance from the suture ; columella distinctly twisted ; anterior canal short. Dimensions. — Length 1 2 mm. ; breadth 5 mm. ; length of aperture 5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9328. Three specimens. Purchased. Pleurotoma subconcava, sp. nov. [Plate III. Figs. 2a-J.] Shell large, elongated; from 11 to 12 whorls; the later portion of the protoconch (the only part preserved in the type specimen) is distinctly longitudinally ribbed ; beyond the sinuous fracture denoting the commencement of the brephic stage these costae suddenly cease and give way to spiral lineations, with a prominent median sulcation indicating the position of the sinus. The whorls throughout the brephic and neanic stages are flat, with a deeply impressed suture, but in the adult stage they become convex, and have a postero-median subangulation at the periphery. Between this and the suture is a well-marked concave depression. The ornament consists of closely-set spiral lineations, which are nearest together in the depression alluded to ; the suture is margined by one or two lineations bolder than the rest. The lines of growth are fairly conspicuous, producing rough, irregular corrugations here and there ; these are most noticeable in the neighbourhood of the 42 PLEUROTOMA. peripheral subangulation, and on the anterior portion of the body- whorl. Aperture small, being about one-third the length of the whole shell, pyriform and produced in front ; outer lip thin, sinus broad, arcuate, situated on the periphery; columella covered by a thin deposit of shell enamel, not callous posteriorly, slightly twisted in the middle ; anterior canal rather long and deep. Dimensions. — Length 58 mm. ; breadth 16 mm. ; length of aperture with canal 23 mm. Form, and Loc. — Eocene : Meribee river, Victoria. 70421. The type specimen. Purchased. Pleurotoma salebrosa, sp. nov. [Plate III. Figs. Za-d.'] Shell fusiform ; whorls eight in number, in the young growth flat ; in the neanic and ephebic stages subangulately convex, rapidly increasing in size ; protoeonch two smooth turns, globose, the later turn being wider than the succeeding whorl. Ornament consisting of from 13 to 14 deep, undulating sulcations on each whorl of the spire, the body-whorl being sulcated throughout in a similar manner ; lines of growth not very conspicuous, but the surface of the shell is peculiarly, distantly corrugated, the corru- gations being five in number on the penultimate whorl, and rising into indistinct, broad, oblique ribs. Aperture pyriform, much contracted in front ; outer lip thin, slightly sulcated within ; sinus situated about half-way between the peripheral subangulation and the suture, broad and shallow ; columella smooth, striated vertically, but not callous, twisted in front ; anterior canal long, broad at its extremity. This shell differs from P. stihconcava in being relatively more tumid, having a larger aperture and fewer whorls ; by the oblique broad, corrugated costse ; and by the position of the sinus, which in this case is indicative of Surcula of authors. The protoeonch, also, is widely different, as will have been noticed. Pimensions. — Length 33'5 mm. ; breadth 12 mm.; length of aperture 17 mm. Form, and Loc. — Eocene : Hobson's Bay, Victoria. 48052. The type specimen. Purchased. [ PLEUEOTOMA. 43 Pleurotoma clarse, Tenison-Woods. 1880. Pleurotoma clarce, Teuisou- Woods, Proc. Liun. Soc. N-S-W. vol. iv. p. 11, pi. iii. fig. 11. 1893. Pleurotoma clarce, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. XTO. pt. 1, p. 221. 1895. Pleurotoma (Surcula) clarce, Tate and Dennant, id. vol. xix. pt. 1, p. 111. Shell elongate, fusiform, and thin ; posterior part of the spire ribbed, anteriorly the ribs become obsolete ; shoulder of last whorl obtusely angulate ; entire external surface of whorls covered with small, closely-set spiral liraa ; aperture broad, the broad sinus being situated near the suture. From the position of the sinus this species might be classed with Surcula ; but in view of the difficulties of making use of that variable characteristic (which in the case of the present species ranges between the shoulder and the suture, but is nearest to the latter), as already stated, no attempt is here made to separate Surcula from Pleurotoma. Dimensions. — Length 17"5mm. ; breadth 6 mm.; length of aperture 9 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4235 and G. 4236. Several specimens, showing range of variation. Presented by John Letma'it, Esc[. Pleurotoma samueli, Tenison- Woods. [Plate III. Figs. 5a-J.] 1879. Pleurotoma samueli, Tenison-Woods, Proc. Linn. Soc. N.S.W. [ vol. iii. p. 226, pi. xx. fig. 3. 1893. Pleurotoma samueli, Tate and Dennant, Trans. Eoy. Soc. South. Aust. vol. xvii. pt. 1, p. 221. Shell fusiform, elongate ; protoconch turbinate, composed of three smooth turns, the suture of which is marginate. In the brephic stage the margination is much accentuated, and at a later period of growth gives way to a row of small, distant granules. In the same manner, the median, tuberculated carina of the adult commences in the brephic stage by large obtuse uodulations. The shell is thus described by Mr. Tenison- Woods : — "It is a polished shell, with whorls angular in the middle, supporting 44 PLEUEOTOMA. a single, somewhat distant series of coarse, blunt, somewhat square tubercles. These are exactly oa the line of the sinus, and at each side the lines of growth curve away from it. The sinus itself is deep, broad, and somewhat quadrate. The aperture is long and round, rather square posteriorly, and the canal is long and only slightly curved." The marginate character of the protoconch is noteworthy ; from its metamorphosis in later stages of growth, it would appear that the species has descended from a stock in which the suture was accompanied by a large flat keel, though no shell of that character has been recorded as occurring in the Australian Tertiaries. Another point of interest in this species is its compound character : it possesses the tubercular sutural coronation and produced spire characteristic of Clavatula, and shows what value is to be placed on that feature ; the special nature of the peripheral carina and the arcuate, long canal somewhat resemble Surcula; whilst the position of the suture and other general features of the shell recall Pleurotoma. Clearly it was pregnant with evolutionary possibilities. Dimensions. — Length 12 mm. ; breadth 4 mm. ; length of aperture 7 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9329. Three specimens. Purchased. Pleurotoma optata, sp. nov. [Plate III. Figs. 4«-A.] Shell fusiform ; aperture (with anterior canal) more than one- half the length of the whole shell ; whorls seven in number, slightly convex, keeled above and below the suture, which is near the centre of the whorls, the anterior keel being the most prominent ; the other ornament consists of several closely-set small spiral lineations, crossed by rather conspicuous growth-lines, and accompanied by a peculiar damascened structure. Protoconch composed of two smooth turns, the anterior being subangulate and vertically striated ; terminating abruptly against the shell of the brephic stage, which is longitudinally, obliquely costated and marginate. Aperture elongately pyriform, much constricted i PLEtTKOTOMA. 45 anteriorly; outermargin thin, undulating; sinus broad and shallow; columella covered by a thin deposit of vertically striated callus, twisted anteriorly ; canal long, curved, broad in front. This shell agrees with others here described in being longi- tudinally costated in the earlier stages of growth, which costse disappear in the adult. It has the aspect of what is generally included under Surcula, but the sinus is far removed from the suture, as in typical Pleiirotoma. Dimensions. — Length 16'5mm. ; breadth 6 mm.; length of aperture 9 5 mm. Form, and Loc. — Eocene : Hobson's Bay, Victoria. 48052. The type specimen. Purchased. Pleurotoma alta, nom. mut. 1873. Pleurotoma pagoda, Hutton, Cat. Tert. Moll. N.Z. p. 5 {non Reeve). 1893. Pleurotoma pagoda, Hutton, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 48, pi. vi. fig. 21. Protoconch composed of two elevated smooth turns ; whorls six in number, smooth, or only marked with growth-lines, very strongly and acutely keeled at the periphery, and with one or two inconspicuous spiral lineations, as an individual characteristic, above or below the keel ; body- whorl subangulate in front, denoting the position of growth as the suture became developed ; aperture ovate, contracted anteriorly, and terminating in a rather long canal; sinus deep, situated between the keel and the suture, but nearer the former ; columella twisted, covered with a smooth, thin deposit of shelly enamel. On conchological grounds this species might be relegated to Surcula, but, as previously remarked, any such reference must always be exceedingly doubtful with fossil specimens. An interesting point is that, whilst many strongly keeled species of PLEUEOTOMiii.a; bear traces of angulation in the protoconch which lead up to the keel in subsequent stages of growth, P. alta is without this in spite of the fact that the carina is most strongly developed in the brephic stage. This species has considerable analogy with Surctila jani, Bellardi, from the Upper Miocene of the Piedmont basin. The specific name alta is here proposed in substitution for pngoda, preoccupied, as iudicatud. 46 PLEUBOTOMA. Dimensions. — Length 18 mm. ; breadth 7-5 mm. ; length of aperture 10mm. Form, and Loc. — Miocene : New Zealand. G. 9498. Several specimens, from Parimoa, Middle Island. W. B. D. Mantell Coll. G. 9495. A tumid variety, from Awamoa. Sir James Hector Coll. Pleurotoma •wanganuiensis, Hutton. 1873. Pleurotoma wanganuiensis, Hutton, Cat. Tert. Moll. N.Z. p. 4. 1893. Pleurotoma wanganuiensis, Hutton, Linn. See. N.S.W., Macleay Mem. Vol. p. 50, pi. vi. fig. 28. Professor Hutton describes this as follows: — "Shell fusiform, the spire longer than the body-whorl. Whorls strongly spirally ribbed, and obliquely longitudinally ribbed in the centre. From 13 to 16 longitudinal ribs in a whorl. Suture very finely obliquely striated, and with a spiral rib. Aperture oval, canal short and recurved." Specimens in the British Museum agree very well with this description, but the figure given in the memoir above cited represents a more tumid variety. The protoconch (Plate II. Pigs. Wa-h) is most remarkable, and merits detailed description. Of the four specimens in the collection, two exhibit the characteristic fracture or undulation at the termination of growth of the protoconch. As thus defined, the latter is composed of from two and a half to three turns; the initial turn is smooth and obsoletely striated, and this is followed by an obtuse angulation which rapidly develops into a bold, acute, median spiral carina, with a conspicuous lineation both above and below it, and another bordering the suture. Up to this point, no longitudinal costse or other ornamentation are developed, and if the growth had con- tinued in the same manner a shell somewhat resembling P. alia would have resulted. Instead of that, however, the brephic stage is characterized from its commencement by the profuse ornamen- tation, suddenly introduced, which obtains in the adult. This should prove of material value in working out the genetic relation- ships of the group. PLEUEOTOMA. 47 By the short canal and longitudinal costae this shell presents some analogy with Drillia, but it differs in the position of the sinus, which is distant from the suture, and in the absence of a posterior callosity on the columellar border. The length of the anterior canal is not of much value in systematics, and in some species of Fleurotoma may only indicate sexual differences. The ribs and position of the sinus are characteristics recalling Surcula. Dimensions. — Length 14 mm. ; breadth 5 mm. ; length of aperture 6 mm. Form, and Loc. — Pliocene : New Zealand. G. 1618. Two adult examples, from Wanganui. G. 9497. An exceptional form, from Shakespeare Cliff. Siir James Sector Coll. Pleurotoma buchanani, Hutton. 1873. Pleurotoma buchanani, Hutton, Cat. Tert. Moll. N.Z. p. 4. 1893. Pleurotoma buchanani, Hutton, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 50, pi. vi. fig. 26. " Shell elongate -fusiform. Whorls obliquely longitudinally plaited, and with fine spiral ribs below the sinus area ; above the sinus area smooth, concave, with a slight ridge at the suture. Between 11 and 15 longitudinal plications on a whorl. Aperture oval, canal produced, rather bent." — Hutton. Protoconch composed of two oblique smooth turns, the anterior portion having four deep spiral sulci, cutting through the small longitudinal costte of the brephic stage, into which the protoconch imperceptibly passes. This species might be classified with Sxtrcula ; the sinus is situated half-way between the suture and the periphery, and is not deep ; the development of longitudinal costae removes it from Pleurotoma, sensu stricto. Dimensions. — Length 20 mm. ; breadth 7'5 mm. ; length of aperture 10" 5 mm. Form, and Loc. — Pliocene : New Zealand. G. 1617. Adult and juvenile specimens, from Wanganui. G. 9496. Three specimens, from Shakespeare Cliff. Sir James Hector Coll. 48 BATHYTOMA. Pleurotoma, sp. indet. Impression, in compact "white limestone, of a large species, probably one of the Pleueotomid^. It is furnished with large equidistant tubercles at the periphery of the whorls, and has a marginate suture, but the specimen is not well enough preserved to warrant specific description. Prom the Tertiaiy of Nullarbor Plains, South Australia. H. Y. L. Brown Coll. Genus BATHYTOMA, Harris and Burrows. [Eoc. and Oligo. Paris Basin, 1891, p. 113.] Bolichotoyna, Bellardi, Bull. Malac. Ital. t. i. 1875, p. 21 {non Hope, 1839). Tryon,* in reviewing the genera of Pleueotomid^, follows Pischer'' in including Dolichotoma as a subgenus of Genotia, and remarks that in common with Pseudotoma, Oligotoma {=Astheno- tomd), Roumiltia, and Cryptoconus its characters are not important. The American author alluded to does not monograph the fossil species, nor does he explain the matter further, so that it is impossible to arrive at his precise meaning. Professor Tate^ remarks that he was not satisfied that Dolichotoma is clearly definable from Genotia, and therefore describes the Australian species which follow as Genotia. I^ow the type of Genotia is Murex mitriformis, Wood, which is an elongate shell with oblique suture, narrow aperture, straight columella, and wide and shallow sinus situated some distance in front of the suture. The type of Bathrjtoma {olim Bolichotoma) is more solid and turbinate, it has a wider and relatively shorter aperture, a very deep sinus situated near the shoulder of the whorl, its outer margin is highly arcuate, and its columella is distinctly twisted. These are differences enabling the two genera to be distinguished at sight. M. Cossmann,* whilst drawing attention to the fact that the Australian Tertiary species are, perhaps, examples of Bathytoma, ' Manual of Conch, vol. vi. 1884, p. 154. - Manuel de Conchyl. 1883, p. 589. =* Journ. Koy. Soc. N.S.W. vol. xxvii. 1894, p. 174. '' Ann. Geol. Universe!, t. x. 1895, p. 756. BATHTTOMA. 49 states that they have not a twisted columella. As the possession of this is a typical generic factor, it may be observed that the French author's statement is not correct; and though, it is true, the figures accompanying Professor Tate's memoir ^ do not exhibit a twist or plication, that author expressly states- that the specimens have a slightly twisted columella. Pleurotoma atradoides, Watson, dredged by the "Challenger" off the Philippines in 375 fathoms of water, should be relegated to this genus. Type. — Murex cata2)hr actus, Brocchi. Bathytoma angustifrons, Tate (sp.). 1893. Dolichotoma angustifrons, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. xvii. pt. 1, p. 221 (Hst name). 189-1. Genotia angustifrons, Tate, Joum. Eoy. Soc. N.S.W. p. 175, pi. x. figs. 7, 7a-i {non 7c). Protoconch composed of two turns, somewhat oblique and slightly longitudinally striated anteriorly. The elevation or depression of these turns to a large extent determiues the relative elongation in the growth of the whole shell. Where it is depressed, the whorls tend to become broad and strongly keeled; where, on the other hand, it is much elevated, the carinations are not so sharp and the shell becomes very long. It is interesting to observe also that but few of the main features of the ornament were foreshadowed in the brcphic stage, and the extremely diversified character of that ornament as the animal became adult is mei'ely an individual characteristic, of no value for systematic purposes. If the shells of a number of very young specimens be compared, no one would have any difficulty in relegating them to a single species; but as they get larger the ornament tends to become so variable that many malacologists would feel inclined to admit the extreme types of variation as of specific rank. Professor Tate's description and figure refer to a commonly prevailing form. The sinus in this species is unusually deep, though not more so than in some forms of the type species from the Pliocene of • Journ. Roy. Soc. X.S.W. vol. xsvii. 1894, pi. x. fig. 7, etc. ' Op. cit. p. 174. 60 BATHTTOMA. the Mediterranean area. Comparing the shell with that of B. catapJiracta of the Upper Tertiary of Italy, the latter will be found to be relatively more elongate and not so rich in ornament, whilst the body-whorl is, in the majority of specimens, more tumid. The aperture of the Australian shell is somewhat narrower. But it is unquestionably a near ally of the European species mentioned ; and bearing in mind the great range of varia- tion in both shells, it would even be desirable on many grounds to unite the two species. The living B. atractoides, Watson, of the Philippines is, perhaps, synonymous in its general characters, though in that case the name proposed by Mr. "Watson would take precedence. The present writer hesitates to deal with the matter, however, as only one specimen of B. atractoides is at his disposal, and that has a much broader, though shallower sinus than any of the fossil examples here quoted. This species may also be compared with B. viabrunnea, Dall (sp.), living in the West Indies. Dimensions. — Length 37 mm.; breadth 16 mm.; length of aperture 19 mm. Form, and Loc. — Eocene : Victoria and South Australia. 48052. Several specimens, from Hobson's Bay. Purchased. 70422. Several specimens, from Meribee river. Purchased. 73224. Four examples, from Brighton. Purchased. G. 4234. Several specimens, from Muddy Creek. Presented hy John Pennant, Esq^. G. 5500 and G. 5503. Two specimens, from Muddy Creek. Purchased. G. 9134. One specimen, from Murray river, near Adelaide. Presented hj William Evans, Esq. G. 9330. Three examples, from Mornington. Purchased. Bathytoma decomposita, Tate (sp.). 1893. Bolichoioma decomposita, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. svii. pt. 1, p. 221 (list name). 1894. Genotia decomposita, Tate, Journ. Eoy. Soc. N.S.W. vol. xxvii. p. 175, pi. X. fig. 7c. Protoconch similar to that of the last-named species. The transverse ornament is not of such a rugged character as in I COLrilBAKITJM. 51 B. angustifrons, but the essential points of difference are the more turbinate character of its spire, and its relatively longer aperture. The body-whorl in typical specimens is rather inflated, and the transverse fold on the columella is very conspicuous. It is closely allied to B. engonia, Watson. Dimensions. — Length 25 mm. ; breadth 1 3 mm. ; length of aperture 16'5 mm. Form, and Loc. — Eocene : Gellibrand river, Victoria. G. 9326. Two specimens. Purchased. Bathytoma fontinalis, Tate (sp.). 1893. Dolichotoma fontinalis, Tate and Dennant, Trans. Roy. Soc. South Aust. vol. xvii. pt. 1, p. 221 (list name). 1894. Genotia fontinalis, Tate, Journ. Roy. Soc. N.S.W. vol. xxvii. p. 175, pi. X. fig. 4. Protoconch slightly more oblique, but is, otherwise, similar to that of the other two species mentioned. The shell may be readily distinguished by its more fusiform character ; the spire- whorls carry a rough angulation, between which and the suture posteriorly is an excavated area with a number of spiral lineations. The lines of growth being rather prominent, a species of oblique reticulation is set up, and where these pass over the peripheral angulation, rudimentary compressed tubercles result. The spiral lineations of the body-whorl are prominent, and occasionally granulose. The sinus is not quite as deeply cut as in the other species mentioned ; aperture pyriform, constricted posteriorly ; columella erect and bearing a well-marked fold. Dimensions. — Length 20 mm. ; breadth 18-5 mm. ; length of aperture 11 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9327. Two specimens. Purchased. Genus COLUMBARIUM, E. von Martens. [Conch. Mittheil. Bd. ii. 1881, p. 105.] Fiisus, in part, of authors. Protoconch bulbous; shell with a comparatively short body- whorl, and long, straight, narrow, nearly closed canal, similar 52 COLUMBAKITJM. to that of the genus Fusus ; margin of the shoulder slightly curved, but there is no sinus, properly so-called. Tryon,' although including Columbarium amongst the Pleuko- TOJirn^, remarks that he feels " pretty well satisfied " that he was correct in placing the typical species of the genus in Fusus. It may be remarked, however, that regarding Murex colus, Linn., as the type of Fusus, the type of the present genus does not, even conch ologically, resemble it very closely. The inclusion of this genus in the Pleukotomidje rests almost exclusively on the character of the radula, which is distinctly toxoglossate. The passage can be understood, conchologically, if Ancistrosyrinx, Dall, be studied in that connection. If we exclude Pleurotoma cedonulli, Reeve, which, as Professor Dall shows,^ should be classed in the genus last mentioned, then the living examples of Columbarium would appear to belong exclusively to the seas of Australia and Japan. Tt/pe. — Pleurotoma spinicinda, E. von Martens. Columbarium acanthostephes, Tate (sp.). 1888. Fusus acanthostephes, Tate, Traus. Eoy. Soc. South Aust. vol. x. p. 133, pi. vii. fig. 7. 1889. Fusus acanthostephes, Dennant, id. vol. xi. p. 41. 1893. Fusus acanthostephes, Tate aud Dennant, id. vol. xvii. pt. 1, p. 219. 1895. Fusus acanthostephes, Tate aud Dennant, id. vol. xix. pt. 1, p. 111. This is a very typical example of the genus. Its protoconch (Plate III. Pigs. 6a-J) is composed of two smooth turns, the earlier of which is bulbous and the later carinate medially. This carina- tion in the brephic stage is developed to an abnormal extent, and is even spinose, as in adult individuals. The spire is turreted ; the whorls, six in number, are " ex- cessively angulated, and coronated at the keel with large erect or radially extended, compressed, hollow spikes. Last and penultimate whorls with three rounded serrated ridges on the posterior slope, reduced to one on the middle whorls, and entirely absent on the earlier ones. Base of last whorl very contracted and precipitous, ornamented with four or five angular keels, the ' Manual of Concbology, vol. yi. 1884, p. 175. - Bull. Museum Comp. Zool. Harvard, vol. xviii. 1889, p. 78. I i COLUMBAEirM. 53 third from the coronal fringe serrated, the others carrying short, crowded, recurved prickles. Beak encircled with prickly ridges." — Tate. Inner margin of the aperture covered with a leaf-like expansive callosity, extending into the anterior canal, which is very long. This species has considerable analogy with the living C. pagoda var. spinicincta, Von Martens, of West Australian seas, and it is highly probable that C. acanthostephes is merely a synonym of that ; in the absence of an example of a shell of the living species alluded to, the present writer hesitates to further deal with the matter, however. Dimensions. — Length 38 mm.; breadth 13 mm.; length of aperture 7 mm. ; length of anterior canal 21 mm. Form, and Loc. — Eocene : Victoria. 70416. Three specimens, from Port Jackson. Purchased. 73229. Several specimens, from Schnapper Point. Purchased. G. 4274. One example, from Muddy Creek. Presented hy John Pennant, Esq. Columbarium foliaceum, Tate (sp.). 1888. Fusus foUaeeus, Tate, Trans. Eoy. Soc. South Aust. vol. x. p. 132, pi. vii. fig. 10. 1889. Fusus foliaceus, Dennant, id. vol. xi. p. 41. 1893. Fusus foliaceus, Tate and Dennant, id. vol. xvii. pt. 1, p. 219. Protoconch of the same general character as in C. acanthostephes, but not so bulbous and relatively much smaller. A remarkable feature is, that in the brephic stage, and immediately after the animal emerged from the capsule, bold longitudinal ribs make their appearance ; but as the neanic stage is approached these disappear, and are not resuscitated in the adult. These costae, which seem to be connected with longitudinal striae on the protoconch (seen when the latter is well preserved), are nodosely angulate medially. The growth of the shell from the neanic stage onwards exhibits no variation worthy of record. The shell differs from C. acanthostephes in having what Professor Tate calls the " coronal fringe " well developed by the absence of "prickles," and the plain character of the area from the peripheral angulation to the suture. The shell as a whole is 54 COLUMBARIUM. not so ornate ; and the protoconch differs, as already described. It is closely allied to C. pagodoides, "Watson, obtained by the "Challenger" expedition off Sydney in 410 fathoms of water; from which it differs in having a much smaller protoconch and a non-spinose anterior canal. Fusus craspedotus, Tate,^ is stated by its author to belong to the same group as his, F. foliaceus (= Columharium foliaceum). Several examples of that species are in the Museum collection ; and the present writer, from a careful examination of that and other material, believes that the differences in the respective proto- coachs, and the general phenomena of the aperture and ornament, are sufficient to warrant generic separation. At the same time it must be remembered that as the status of this genus largely depends on the character of the radula, and as that part of the animal is not found in fossil specimens, the classification of fossil species which conchologically resemble the recent forms, must always remain doubtful. Dimensions. — Length 45 mm. ; breadth 13'5 mm. ; length of aperture 9 mm. ; length of anterior canal 23 mm. Form, and Loc. — Eocene : Victoria. 73228. A series exhibiting stages of growth from the very young to the adult, from Schnapper Point. Purchased. G. 4287. One specimen, from Muddy Creek. Presented ly John Bennant, Esq. Columbarium spiniferum., Tate (sp.). 1888. Fusiis spiniferus, Tate, Trans. Eoy. Soc. South Aust. vol. s. p. 124, pi. vii. fig. 1. Similar to C. acanthostephes, but with shorter spire ; the posterior slope of the whorls has no spiral ornament ; angular ridge spinose, the spines being irregularly developed in regard to size ; in front of this, and coinciding with the posterior angle of the aperture, is a slender, crenulated, spiral carina, and minor spiral threads are developed anteriorly; aperture subquadrate, contracted in front; outer margin dentate ; anterior canal long and spinose. 1 Trans. Roy. Soc. South Aust. vol. x. 1888, p. 134. 55 Dimensions. — Length 30 mm.; breadth 10 mm.; length of aperture 7 mm. ; length of anterior canal 17 mm. Form, and Loc. — Eocene : Hiver Murray cliffs, near Morgan. G. 9334. Three specimens. Purchased. Genus DRILLIA, Gray. [Ann. Nat. Hist. vol. i. 1838, p. 28.] This differs from Pleurotoma in having longitudinal ribs ; the last whorl is, typically, short; and the columella is furnished with a posterior callosity. The outer lip is thick, flexuous, and a deeply-cut sinus is situated near the suture ; whilst the columella is curved and frequently umbilicate. Type. — Drillia umbilicaia, Gray. Drillia trevori, Tenison "Woods. 1879. Drillia trevori, Tenison -Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 227, pi. XX. fig. 4. 1889. Drillia trevori, Dennant, Trans. Eoy. Soc. South Aust. vol. xi. p. 44. Protoconch composed of two tumid, shining turns, the posterior portion being laterally immersed. The longitudinal costae, which become so well developed at later stages of growth, commence in the brephic by prominent tubercles. Shell regularly and spirally grooved ; suture hidden by an elevated prolongation of the posterior part of the whorls, between which and the ribs referred to is a rather broad and flat space. Aperture very small ; outer margin greatly curved, sinus deep and near the suture ; columella covered by a leaf-like expansion and having a tuberculose callus posteriorly ; anterior canal short and somewhat twisted. This species is said to occur both in the Eocene and Miocene beds of Muddy Creek. Mr. Tenison- Woods states (o/;. cit. p. 228) that it considerably resembles D. beraudii of the south-east coast of Tasmania. Dimensions. — Length 13 mm.; breadth 6 mm.; length of aperture 5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4220. Three young and one adult specimens. Presented by John Dennant, Fsq. 56 Drillia oblongula, sp. nov. [Plate III. Figs. 7fl-i.] Shell small, turreted, fusiform ; protoconch witli two smooth subangulate turns, closely resembling that of D. trevori. Whorls six in number, longitudinally obliquely costated in the brephic and neanic stages, closely spirally lineate, the lineations being irregularly granular at the points of intersection of the growth- lines; typically the costated whorls are deeply sulcated spirally, and in later stages of growth a lineation is developed in each sulcus. On arrival at the adult stage this lineation becomes more conspicuous, but never attains to the same proportions as the originally-formed lineations. These latter persist over the whole body-whorl down to the tip of the anterior canal. Suture marginate and accompanied, anteriorly, by a broad band, medially sulcated, denoting the former positions of the sinus. Aperture narrow, but inflated towards the middle ; outer margin arcuate, the deep, broad sinus being situated near the suture ; inner margin covered by a thin coating of enamel ; columella slightly twisted ; anterior canal very short. This differs from D. integra, Tenison- Woods, in not being con- spicuously ribbed, and by the absence of costae in the adult, though the shell is somewhat corrugated longitudinally at that stage. It differs also in not being broadly sulcate above, nor regularly spirally lirate ; whilst the outer margin is not thickened to an appreciable extent. A form from the same locality has smoother whorls. Dimensions. — Length 13'5 mm. ; breadth 5 mm.; length of aperture 6 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4219. Adult and young examples. Presented hy John Bennajit, Esq. Drillia vixumbilicata, sp. nov. [Plate III. Figs. 8a-4.] Protoconch composed of two smooth elongnted turns, the anterior of which is faintly spirally striated. Shell lanceolate, having seven subangulate convex whorls ; the earlier of these are distinctly MANGILIA. 57 costate ; the whole surface of the whorls is spirally striated, but not so conspicuously between the angulation and the suture ; the latter is marginate. Aperture small and narrow ; inflated towards the middle ; outer margin produced, arcuate ; deep, broad sinus in the proximity of the suture; inner margin curved, enamelled, with a posterior callosity ; umbilicus very small ; anterior canal small, almost truncate. Dimensions. — Length 15-5 mm. ; breadth 4"5 mm. ; length of aperture 5'5 mm. Form, and Loc. — Eocene : Muddy Creek, Yictoria. G. 4221. An adult specimen. Presented hy John Dennant, Eaq. Genus MANGILIA (Leach MS.), Eisso {em.). I [Hist. Nat. Eur. Merid. t. iv. 1826, p. 219.] Bda (Leach MS.), Gray, Proc. Zool. Soc. 1847, p. 13-1. Shell fusiform, not perforated ; aperture commonly narrow, with a short truncated canal ; inner margin varicose ; sinus near the suture. Typically the shell is longitudinally costate. It is usual to include species with a well-marked, deep sinus ; the type of the genus, however, has a mere curvature as a sinus. In regard to Bela, see p. 60. Ti/pe. — Mangelia costulata, Eisso. Mangilia obsoleta, sp. nov. [PI. Ill, Figs. 9«-Zi.] Shell small ; protoconch composed of two smooth turns, deeply umbilicated ; whorls four and a half to five in number, spire-whorls short, subangulate not far from the suture, oblique costae not very prominent and becoming obsolete on the anterior portion of the body-whorl ; the latter is very large in proportion to the size of the shell ; spiral strife not prominent, equidistant, more numerous between the peripheral subangulation and the suture. Aperture elongate ; outer margin curved, slightly varicose, with an elevated deep sinus, also varicose, near the suture; columellar margin covered with a thin deposit of smooth callus; canal short. This is by no means a typical Mangilia ; but for the absence 58 MANGILIA. of denticulations -within the outer margin of the aperture and the comparatively inconspicuous longitudinal costse, it might have been included in the subgenus Cythara. Dimensions. — Length 10 mm.; breadth 4-5 mm. ; length of aperture 6 mm. Form, and Loc. — Eocene (?) : Muddy Creek, Victoria. G. 4222. One specimen. Presented by John Demiant, Esq. Subgenus CYTHARA, Schumacher. [EssaiNouv. Syst. Habit. Testaces, 1817, p. 245.] Shell elongate ; longitudinally costate ; aperture narrow ; outer margin more or less denticulated within ; columellar border striated. Typically the shell is columbelliform. Type. — Cythara striata, Schumacher. Mangilia (Cythara) glabra, sp. nov. [Plate III. Figs. \Qa-b.] Shell solid, narrow, elongate ; whorls four in number, obliquely costellated, the costae being large, rounded, curved (especially near the suture), equidistant, and extending over the suture, which is much hidden in consequence ; between the ribs are several spiral lineations. Aperture narrow, widening somewhat in the middle ; outer margin thick, callous, obscurely denticulated within; sinus deep, oblique, near the suture and bordered by a thick callosity; columellar border covered, the callus being thick posteriorly; columella excavated ; anterior canal short and truncated. It is closely allied to M. hicohr, Reeve, living in the seas of IS'orthern Australia and the Philippines, but is narrower, whilst tlie columella is excavated instead of being convex or sinuate. More distantly it is related also to If. capillacea, Reeve, of the same geographical regions. Dimensions. — Length ITS mm.; breadth 5 mm.; length of aperture 6 mm. Form, and Loc. — Miocene (?) : Muddy Creek, Victoria. G. 4201. Slightly rolled specimens. Presented hy John Dennant, Fsq. CLATHCEELLA. 59 Genus CLATHURELLA, Carpenter. [Mazatlan Cat. 1857, p. 399.] Differs principally from Mangilia in its whorls being more rounded and cancellate ; sinus varicose and sutural ; inner margin of the aperture denticulate posteriorly. It is closely allied to that genus, however. Type. — Pleurotoma purpurea, Montagu. Clathurella bidens, Tenison-Woods (sp.). 1879. Mangelia bidens, Tenison-'Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 227, pi. sx. fig. 2. 1895. Mangilia bidens, Tate and Deunant, Trans. Roy. Soc. South Aust. vol. xix. pt. 1, p. 111. Protoconch composed of two smooth, angulate, elevated turns. Shell fusiform, tumid ; whorls six in number, obliquely costate and distinctly Urate ; between the subangular periphery and the deeply-cut suture the lirse are smaller and more numerous. Aperture rather broad ; outer margin varicose, produced, and thin at the edge, dentate within ; sinus large, broad, and situated near the suture ; inner margin curved, with a callous denticle posteriorly ; columella slightly twisted ; canal broad and short. Dimensions. — Length 15 mm.; breadth 7 mm. ; length of aperture 8 mm. Form. a?id Loc. — Eocene : Muddy Creek, Victoria. G. 4202. Four examples. Presented by John Bennant, Esq. Clathurella obdita, sp. nov. [Plato III. Figs. lU-J.] Shell fusiform, elongate, solid ; protoconch composed of one and a half smooth, convex turns terminating abruptly against the shell proper, which, even in the brephic stage, is distinctly angulate, and possesses the general characteristics of the ornamentation of the adult. Whorls live in number, spirally lirate throughout, the lirte being larger and farther apart in front of the peripheral sub- angulation. The growth-lines develop into equidistant, oblique, inconspicuous costae, best seen in the early stages of growth ; and 60 DAPHXELLA, 'where they cross the larger spiral lirae, and especially at the periphery, teeth-like tubercles result. Aperture narrow ; outer margin thick, dentate within, varicose, produced, the anterior portion bent and partly encircling the short canal; sinus sutural, very conspicuous, deep and bordered by the posterior extension of the outer margin, which covers the suture in that region, and is reflected towards the thin enamel of the inner margin ; a prominent denticle is placed on the latter near the sinus ; columella somewhat excavated, sharply twisted anteriorly. This species differs from M. lidens in being more elongate, the spire is much longer in proportion to the aperture, is narrower, the columella is more sharply twisted ; the longitudinal costse are not so conspicuous, the subangulation at the periphery is more towards the centre of the whorls, and the surface of the shell between that and the suture is more sloping. The shell as a whole is not so rugged. Dimensions. — Length 9 mm. ; breadth 4 mm. ; length of aperture 4-5 mm. Foi'm. and Loc. — Eocene (?) : Muddy Creek, Victoria. G. 4204. Two specimens. Presented ly John Bennant, Esq. Genus DAPHNEL.LA, Hinds. [In Belcher, Voy. " Sulphur," Mollusca, vol. ii. 1844, p. 25.] Shell thin, fusiform ; whorls convex ; body -whorl usually elongate, surface finely cancellated; outer margin slightly curved, broad shallow sinus near the suture ; columella excavated ; canal short, truncate. None of the species here described are typical examples of the genus ; they appear to be intermediate between Baphnella {seiisu strido) and its subgenus Pleuroiotnella, Yerrill. They are not included in the latter, by reason of the sinus not being deep enough, and from the absence of the "smooth concave surface" between the shoulder of the whorls and the suture. By some authors they would possibly be classified with Bela ; that genus, however, appears to be much misunderstood. It does not seem to be recognized that the type of Bela (Leach MS.), Gray,' is Murex ' Proc. Zool. Soc. 1847, p. 134. DAPHNELLA. 61 neiula, Montagu, which is practically synonymous with Mangelia costidata, Risso, the type of the genus Mangilia {em.). Jfoue of the Australian Tertiary forms now figured and described bear the slightest resemblance to " Murex nebula," which Tryon ' classifies with Raphitoma, Bellardi, a subgenus of JDaplmeUa. Type. — Pleiirotoma lymnmiformis, Kiener. Daphnella sculptilis, Tate (sp). [Plate lY. Figs. l«-5.] 1888. Bela sculptilis, Tate, Trans. Eoy. Soc. South Aust. vol. x. p. 173, pi. iv. fig. 1. 1889. Bela sculptilis, Demiant, id. vol. xi. p. 45 (list name). Protoconch composed of one and a half smooth, depressed, obtuse turns, the earlier portion of which is laterally disposed, and the later longitudinally striated. Whorls tumid, six in number, subangulate at the periphery and somewhat excavated between that and the suture. The latter is canaliculate and margined. Surface covered by closely -set spiral lineations, remarkably regular from the brephic stage onwards, and crossed by conspicuous growth - lines, which occasionally produce minute granulations at their points of intersection, but are for the most part more clearly defined in the sulci between the lineations referred to. Aperture nearly one-half the length of the shell, narrow, obliquely terminated posteriorly and curved anteriorly where a short broad canal exists ; outer margin thin, moderately curved, sinus broad but shallow, and occupying the whole of the sutural depression ; inner margin covered by a very thin deposit of enamel, best developed anteriorly ; columella twisted. This species, although figured, has not, apparently, been previously described. Dimensions. — Length 47 mm.; breadth 18 mm.; length of aperture 21 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4194. Three specimens. Presented hy John Bennant, Esq. 1 Manual of Concliology, vol. vi. 1881, p. 307. 62 DAPHNELLA. Daphnella crassilirata, Tate (sp.). [Plate IV. Figs. 2«-S.] 1888. Bela crassilirata, Tate, Trans. Roy. Soc. South Aust. vol. x. p. 173, pi. iv. fig. 7. 1889. Bela crassilirata, Dennant, id. vol. xi. p. 45 (list name). Protoconcli similar to that of B. sculptilis, but somewhat canali- culate, and the anterior portion is more strongly striated spirally. Shell tumid ; whorls five to six in number, angulate at the periphery, between which and the suture is a deep depression ; suture canaliculate and marginate ; surface of the whorls uniformly and deeply sulcated spirally, and crossed by regular growth-lines which produce a cancellation especially well developed in the neighbourhood of the suture. Aperture rather broad, pyriform, obtuse posteriorly ; outer margin very thin, sinus broad, but shallow between the suture and the periphery ; inner margin covered by thin deposit of callus, through which the spiral sulcations are visible posteriorly ; columella excavated, twisted anteriorly. This species, though figured, does not appear to have been hitherto described. It differs essentially from D. sculptilis in being shorter, having the peripheral angulation placed nearer the suture, in being much more deeply excavated between ; the surface of the shell is more deeply sulcated, and the aperture is wider. Dimensions. — Length 25 mm.; breadth ll'Smm. ; length of aperture 13'5 mm. Form, and Zoc— Eocene : Muddy Creek, Victoria. G. 4193. Two specimens. Fresented, hij John Bennant, Fsq. G. 5496. One example. Furchased. Daphnella pulchra, Tate (sp.). 1888. Bela pulchra, Tate, Trans. Roy. Soc. South Aust. vol. x. p. 173, pi. iv. fig. 2. 1889. Bela pulchra, Dennant, id. vol. xi. p. 45 (list name). Protoconch composed of one and a half smooth, polished turns, turbinate, depressed, and canaliculate. Shell as a whole short and DAPnNELLA. 63 tumid ; whorls five in number, convex ; suture slightly canaliculate and bordered by a shallow excavation. Surface covered by spiral striaj, the penultimate whorl carrying six prominent striations, between each of which are three or four minor ones. Growth-lines regular, producing a species of cancellation, the points of inter- section of the larger with the revolving striae being crowned by a small tubercle. These groAvth-lines, also, are of two orders, three or four minor ones being situated between the more prominent of them. Aperture broad, rounded posteriorly and truncated anteriorly ; outer margin thin, curved, the sinus being marked by a mere sinuosity in the neighbourhood of the suture; columella excavated. This species does not appear to have been described before, though figured by Professor Tate, as above indicated. It may be readily distinguished from the two species of Laplmella already described, by the granulated aspect of the whorls, as seen under a lens, by the comparatively shallow excavation bordering the suture, and by the special character of the sinus. Dimensions. — Length 18 mm.; breadth 9 mm.; length of aperture 9 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4157. Two examples. Presented htj John Bennant, Esq. Daphnella striata, Hutton. 1873. Bela striata, Hutton, Cat. Tert. Moll. N.Z. p. 5. 1893. Daphnella striata, Hutton, Linn. See. N.S.W., Macleay Mem. Vol. p. 52, pi. vii. fig. 33. Shell cominelliform, body-whorl longer than the spire ; whorls convex, six in number, finely spirally striated ; aperture ovato- elongate, wide anteriorly; outer margin sinuous, but there is no sinus properly so-called, reflected ; columellar margin covered by a leaf-like expansion of smooth shell enamel, which is continuous with the outer margin, near the suture is a small callosity ; columella twisted ; canal short, recurved. Dimensions. — Length 20 mm. ; breadth 9 mm. ; length of aperture 10 mm. Form, and Loc. — Pliocene : Shakespeare Cliff, Wanganui, New Zealand. G. 9509. Two specimens. Sir James Hector Coll. 64 TELEOCHILUS. TELEOCHILUS/ nov. gen. Protoconch (Plate III. Figs. I2c-d) large, from one to one and a half depressed turns, spirally and longitudinally striated from the initial portion onwards, the spiral striae developing into well- defined sulcations as the shell passes from the brephic to the neanic stages of growth. Shell as a whole elongate, the aperture being much longer relatively than, the spire ; whorls slightly convex and spirally sulcated throughout, lines of growth well marked ; suture typically canaliculate ; aperture with thin, erect, outer margin, constricted posteriorly, and broad towards the anterior ; columella plain, with a thin deposit of callus through which the sulci appear. It is difficult to satisfactorily classify this genus. By its general' configuration it has considerable analogy with Daphnella ; but may be easily differentiated by the unique characteristics of the protoconch, and the absence of a posterior sinus. The last- mentioned circumstance might be taken advantage of to remove Teleochilus from the Pleueotomii)^, did we not know that the possession of a sinus is not an absolutely necessary qualification for inclusion in that family. In some respects Teleochilus recalls certain forms of Mitra, but it has no columellar plications, nor even the semblance of a fold ; whilst its protoconch is distinctive. At the same time, it approaches the genus Dilaphus, Philippi, which, although deprived of folds on the pillar, is classified with the Mitrid^, chiefly on anatomical grounds. Dihnphus differs only from Mitroidea, Pease, by the absence of plaits. The soft parts of the animal were first described by Mr. A. Garrett,^ who was unable to detect any difference between them and those of Cylmdra, Schumacher. Unfortunately, it is not possible to ascertain the relationships of Teleochilus in the same manner, as its soft parts are denied to us; on conchological considerations alone it is included in Plettrotomid^, and may be assigned a position in the neighbourhood of Daphnella. Type. — Daphnella gracillima, Tenison- Woods. 1 t6'a.eios ' complete ' ; x*"^"^ ' hp-' = Proc. Zool. Soc. 1872, p. 843. CANCELLARIA. 65 TeleocMlus gracillimum, Tenison- Woods (sp.). 1876. Daphnella graciUima, Tenison-Woods, Pap. Roy. Soc. Tas. 1876, p. 106. 1879. Daphnella graciUima, Tenison-Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 226, pi. xx. fig. 10. Mr. Tenison -Wood's description of this species is as follows : " Shell fusiform, thin, fragile, shining, whorls sis, gracefully sloping, last longer than the spire, finely striate lengthwise with irregular ridges of growth which become fine, close, and rounded at the lip ; regularly and somewhat distantly grooved with rather broad, flat, shallow, conspicuously striate grooves, one of which is much broader just below the suture, which is distinctly canaliculate ; apex obtuse and Natica-Wke ; mouth narrow, finely rounded at the suture ; outer lip thin ; canal short, base of the pillar finely and obliquely decussate behind." The general characteristics of the protoconch have been alluded to in the generic description. The figure of the species given by its author is a representation of a much worn specimen ; it is deemed advisable, therefore, to refigure it (Plate III. Figs, \2a-d). Dimensions. — Length 25 mm. ; breadth 10 mm. ; length of aperture 16mm. Form, and Loc. — Eocene : Muddy Creek, Yictoria. G. 4161. Four examples, shell of the adult. Presented hy John Dennant, Esq. G. 4162. Two examples, exhibiting the neanic stage of growth. Presented hy John Dennant, Esq. Family CANCELLAEIID^. Genus CANCELLARIA, Lamarck. [Mem. Soc. Hist. Xat. Paris, 1799, p. 71.] Shell oval, fusiform ; aperture slightly canaliculated ; columella stnmgly plicated ; outer margin not reflected, crenulated. Type. — Voltda reticulata, Linnasus. 66 CANCELLAEIA. Cancellaria wannonensis, Tate. 1889. Cancellaria wannonensis, Tate, Trans. Roy. Soc. South Aust. vol. si. p. 156, pi. viii. fig. 11. Shell solid, ventricose ; protoconch elevated, more vigorously developed in some individuals than others, typically composed of two and a half turns, but occasionally two and sometimes three turns, faintly longitudinally ridged anteriorly ; whorls strongly ribbed and spirally lirate ; aperture wide, outer border arcuate, inner border callous, especially in the adult stage ; columella imperforate, having three prominent plications, the posterior of which is the largest, frequently with a smaller ridge between the central and the posterior plications, and some rugosities on the flat portion of the callus in the umbilical region. It has considerable affinities with the living C. semperiana, Crosse, from New Caledonia, but the aperture of the latter is much more dilated, the longitudinal costse are bolder, and the shell as a whole not so solid. Dimensions. — Length 32 mm.; breadth 21mm.; length of aperture 21 mm. Form, and Loc. — Miocene : Muddy Creek, Victoria. G. 4259. A senile example. Presented hy John Bennant, Esq. G. 5524. One specimen. Purchased. G. 9374. A series exhibiting stages of growth. Purchased. Cancellaria gradata, Tate. 1889. Cancellaria gradata, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 155, pi. X. fig. 12. 1889. Bivetia gradata, Cossmann, Ann. Geol. Univ. t. v. p. 1091. This species is very different to any other Cancellaria from Australasia in the British Museum. The shell is thin, cancellate, sharply carinate, with a broad spiral platform at the suture ; the spire in consequence is staged ; the protoconch is composed of one and a half turns, the early portion of which is implanted obliquely and is somewhat inflated, the remainder being decidedly convex with deeply-cut suture ; aperture quadrately oval, outer border crenulated, lirate within ; columella slightly callous, furnished with three prominent plications. CANCELLAEIA. 67 This is by no means a typical Cancellaria. M. Cossmann remarks {op. cit. supra) that it should be classified with Bivetia ; but the type of that subgenus has a posterior canal, varices, and other characteristics absent in C. gradata. For the present it may remain with Cancellaria proper, the present writer having no inclination to increase the number of subgenera with which this group is burdened. Dimensions. — Length 12 mm.; breadth 6-5 mm. ; length of aperture 6 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4262. One specimen. Presented ly John Dennant, Esq. G. 9372. An adult example with well-preserved protoconch. Purchased. Cancellaria, sp. G. 5680. Two imperfect, rolled specimens ; from the post- Pliocene beds of Limestone Creek, Glenelg river, Victoria. Purchased. Subgenus NARONA, H. and A. Adams. [Genera Recent Moll. vol. i. 1854, p. 277.] Shell not umbilicated, oblong, costulated, or cancellated ; whorls depressed near the suture ; columellar border with two oblique plications. Type. — Cancellaria clavatula, Sowerby. Cancellaria (Narona) capillata, Tate. 1889. Cancellaria capillata, Tate, Trans. Eoy. Soc. Soutli Aust. vol. xi. p. 158, pi. X. %. 10. 1889. Karona capillata, Cossmann, Ann. Geol. Univ. t. v. p. 1091. 1893. Cancellaria capillata, Tate and Dennant, Trans. Roy. Soc. South Aust. vol. -xvii. pt. 1, p. 221. Shell small, solid ; protoconch smooth, except anteriorly, where it is longitudinally costated, composed of one and a half convex turns, the earlier portion being somewhat oblique with reference to the axis of the shell; whorls convex, lirate, and slenderly 68 CANCELLAEIA. costated ; the spiral liise produce small nodose crenulations on the costaj. The external ornament is much interrupted by inflated variciform ridges on several individuals in the Museum. Aperture oblong, outer border lirate within ; columella with two small transverse plications ; umbilicus partly covered over by the columellar border. Dimensions. — Length 5 mm. ; breadth 2-75 mm. ; length of aperture 2'5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9369. Four adult examples. Purchased. Cancellaria (Narona) etheridgei, Johnston. 1880. Cancellaria etheridgei, Johnston, Proc. Roy. Soc. Tas. 1879, p. 32. 1889. Cancellaria etheridgei, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 157, pi. is. fig. 6. 1889. Karona etheridgei, Cossmann, Ann. Geol. Univ. t. v. p. 1091. 1893. Cancellaria etheridgei, Tate and Dennant, Trans. Eoy. Soc. South Aust. vol. xvii. pt, 1, p. 221. This species is closely allied to C. capillata, but the longitudinal costae are relatively larger and oblique ; columella with two plications situated posteriorly. The specimen in the Museum is not sufficiently well preserved to enable the protoconch, or minute ornament, to be described. The general character of the embryonic shell was, evidently, similar to the last- mentioned species, however. I)imension,s. — Length 5 mm. ; breadth 3 mm. ; length of aperture 2"5 mm. Form, and Loc. — Eocene : Spring Creek, Yictoria. G. 9373. Two worn specimens. Furchased. Subgenus ADMETULA, Cossmann. [Ann. Soc. Eoy. Malac. Belg. t. xxiv. 1888, p. 22-1.] This subgenus differs from Admete, MoUer, in the possession of varices and by the thickness of the test. Ty2}e. — Buccinum evidsum, Solander. CANCELLAEIA. 69 Cancellaria (Admetula) varicifera, Ten.-Woods. 1S79. Cancellaria varicifera. Ten. -Woods, Proc. Linn. See. X.S.W. vol. iii. p. 231, pi. xxi. fig. 12. 1889. Cancellaria varicifera, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 15.5. 1893. Cancellaria varicifera, Tate and Dennant, Trans. Roy. Soc. South Aust. vol. xvii. pt. 1, p. 221. Professor Tate remarks that this is the antipodean analogue of C. erulsa, Solander, from Barton, and that it differs "only by its much finer ornament, and by the lirte within the outer lip." No doubt there is a close I'esemblance between the two species, but a study of the protoconch is sufficient to separate them immediately. The Australian shell commences with a relatively large protoconch, the earlier portion of which is obtuse and even slightly inflated, and is composed of only one and a half turns ; the Bartonian .shell, on the other hand, has a somewhat pyramidal protoconch com- posed of from two and a half to three turns, the earlier portion is not oblique, and the whole is regularly coiled. There must be some error respecting the lirae alluded to : both the Australian and European shells possess these within the outer lip, typical examples of C, evulsa from Barton having them especially prominent opposite the varices. The Australian shell is thinner. Bimensmis. — Length 14 mm. ; breadth 8 mm. ; length of aperture 5'5 mm. Form, and Loc. — Eocene : Victoria. 48052. Specimens much more tumid than the type ; this has considerable analogy with C. evulsa ; from Schnapper Point. Purchased. G. 4260. An elongate specimen ; from Muddy Creek. Presented hy John Dennant, Esq. G. 9371. A series exhibiting stages of growth. Purchased. Subgenus SVELTIA, Jousseaume. [Le Xaturaliste, t. i. ser. 2, 1887, p. 214.] Shell not umbilicated, long, turriculated ; suture deep ; aperture ovate ; posterior canal hardly perceptible ; anterior canal large and cutting into the peristome ; columella with two unequal, rather salient plications. Type. — Cancellaria varicosa, Brocchi. 70 Cancellaria (Sveltia) epidromiformis, Tate. 1889. Cancellaria epidromiformis, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 154, pi. viii. fig. 9. 1889. Sveltia epidromiformis, Cossmann, Ann. Geol. Univ. t. v. p. 1091. Shell long, thick with sub-turreted spire ; protoconch obtuse, composed of one and a half turns, the earlier portion being planted obliquely with reference to the axis of the shell, smooth, except anteriorly, where frequent, microscopic, longitudinal lineations appear ; whorls deeply excavated at the suture, where an inward- sloping spiral platform occurs ; spirally and longitudinally ribbed, granulated at the points of intersection, the former being the more prominent ; interrupted here and there by varices ; aperture relatively small, ovate ; outer border plicate within ; columella imperforate, with three plications ; anterior canal truncate. This is a typical example of the subgenus Sveltia, except that it has three columellar plications instead of two. In the young these are even more marked than in the adult. Dimensions. — Length 20 mm.; breadth 10 mm.; length of aperture 9 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4260. An adult example. Presented hy John Bennant, Esq. G. 9370. Two specimens, one of which is a well-preserved example of the neanic stage of growth. Furcliased. Family OLIVID^. Genus OLIVELLiA, Swainson. [Malacology, 1840, p. 34.] Olivina, D'Orbigny, Voy. Amerique Meridionale, t. v. 1841, p. 417 [non Morch). A comparison of the soft parts of Olivella and Oliva exhibits many features of interest, fully sustaining the generic separation of the two. The head of Oliva, for instance, is furnished with slender tentacles and eyes, the median tooth of its radula is OLIVELIA. Yl tricuspidate; on the other hand, a typical OUvella has neither tentacles nor eyes, ami the median tooth of the radula carries a great number of small denticles, in addition to which the radula has accessory lateral plates. In spite of these important anatomical differences, the shells of certain species of both genera closely resemble each other, and it is, therefore, almost impossible to classify them with any degree of accuracy from an examination of the test alone. Fortunately, however, the more extreme forms of the shells are not difficult to determine generically, and the Australasian Tertiary fossils about to be described are allocated to the genus OUvella without much doubt. In general, the shells may externally be distinguished from those of OUva by the columella not being plicate (or lirate) throughout its entire length, whilst the columellar border is more obliquely inclined with reference to the axis, and is sometimes slightly excavated (as in 0. nymphalis, Tate) ; the spire is more elevated, though this last character is by no means constant. Ti/pe.— Oliva Uplicata, G. B. Sowerby. Olivella angustata, Tate (sp.). 1889. OUva angmtata, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 144, pi. \m. figs. 7a-*. 1889. OUva angustata, Dennant, id. vol. si. p. 43. The spire in this species is long, in adult specimens being more than one-third the entire length of the shell ; suture deep ; aper- ture narrow, wider where the columella is slightly excavated; plications variable in number, situated on the anterior half of the' columellar border only. Compared with 0. nitidula, Desh., of the Parisian Eocene, it is found to be longer, its aperture is narrower and straighter, and Its columella is less excavated ; in other respects it presents con- siderable analogy with the shell of that species. Dimensions. — Length 26 mm.; breadth 8-5 mm. ; length of aperture 15-5 mm. Form, and Zoc— Eocene : ]\:uddy Creek, Victoria. G. 4170. Three specimens. Presented hy John Dennant, Esq. G. 9367. Examples showing stages of growth. Purchased. 72 Olivella nymphalis, Tate (sp.). 1889. Oliva nymphalis, Tate, Tians. Roy. Soc. South Aust. vol. -xi. p. 145, pi. vii. fig. 7. 1889. Oliva iii/mphalis, Dennant, id. vol. xi. p. 43. 1889. Olivella >i!/mphalis, Cossmann, Ann. Geol. Univ. t. v. p. 1090. This is a more stunted form than 0. angustata, whilst the plications are only two or three in number, being confined to the anterior portion of the columella. The whorls have a tendency to become invaginate, and the spiral sulcation near the middle of the body-whorl is prominent. A close European ally is 0. nritreola, Lamarck, a more elongate shell, from the Eocene of the Cotentin and Paris Basin. Dimensions. — Length 8"5 mm. ; breadth 4 mm. ; length of aperture 4 mm. Form, and Loc. — Miocene : Muddy Creek, Victoria. G. 9368. Two specimens. Purchased. Olivella, sp. G. 9511-2. Two pieces of argillaceous limestone containing casts of several shells, including a species of Olivella closely allied to 0. angustata; from the Lower Miocene (Mangapakeha beds) of Akuakua, east coast of Auckland, N.Z. Sir James Hector Coll. Genus ANCILLA, Lamarck. [Mem. Soc. Hist. Nat. Paris, 1799, p. 70; also Syst. des Anim. sans Vert. 1801, p. 73.] Ancillaria, Lamarck, Ann. Mus. 1810, t. xvi. p. 302. Sparella, Gray, Guide Moll. Brit. Mus. pt. 1, 1857, p. 26. Shell oblong, occasionally acuminate ; suture sometimes canali- culate, but more frequently covered over by callus ; columella excavated, twisted, and carrying several plications anteriorly, which are not hidden by callous growth, whereas the posterior portion of the columella is thick, callous, or gibbose. A depression, or groove, usually separates the callosity of the spire from what is commonly termed the " smooth zone," though this zone is often not as smooth as the callous spire, especially when strongly exhibiting the lines of growth as in certain fossil species. The 73 relative size of this zone is of specific walue taken in conjunction with cognate characters ; aperture grooved posteriorly, and truncate and deeply sinuous in front. Ancilla may bo distinguished from Olivancillaria, with which it is frequently confounded, in that the latter, sensu stricto, has the columella callous thi-oughout its entire length, the spire is usually much more acuminate, and the sutures are deeply canaliculate. The columella of the section Agaronia, however, is not callous, except to a feeble extent posteriorly. Ancilla is readily dis- tinguished from that subgenus, as the latter has such well- developed sutures, and the comparative absence of callosity on the spire and body-whoii is well marked. The subgenus Sparella, almost universally adopted by modern authors, must disappear in synonymy, for the following reasons : — The type of Ancilla is A. cinnamomea, Lamarck, and the type of Sparella is A. alhisuleala, Sby. ; but these two species are synonymous with each other, the only distinguishing feature between them being that the impressed groove of the latter is white — a character without constancy. It is generally stated that a more or less prominent denticle on the outer margin of the aperture characterizes Sparella, but that is found also in the type species of Ancilla ; the spread of the callosity over the spire is usually regarded as a special feature, but what value that may possess from a morphological point of view does not in any way affect the synonymity of the type species mentioned. Ti/pe. — Ancilla cinnamomea, Lamarck. Ancilla pseudaustralis, Tate (sp.). 1889. AnciUaria pseudaustralis, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 148, pi. vi. fig. 13; pi. vii. fig. 1. 1893. Ancillaria pseudaustralis, Tate and Dennant, id. vol. svii. pt. 1, p. 220. This is the largest Australian Ancilla in the Museum. Professor Tate compares it with the European Eocene A. luccinoides, Lamk., but it differs markedly from the latter : the broad band trespassing anteriorly on the smooth zone of the body- whorl, terminating at the inner margin of the aperture by a pronounced denticle forming a slight postei'ior prolongation of the band, or of the groove which borders it posteriorly, at once distinguishes this species from 74 A. huccmoides. The granulations on the callus, the manner in which a portion of the latter is spread over the whorls of the spire from the columellar border nearly to the protoconch, and the relatively smaller size of the twisted anterior of the columella, are also distinctive features. Dimensions. — Length 46 mm. ; breadth 20 mm. ; length of aperture 26 '5 mm. Form., and Loc— Eocene : Muddy Creek, Victoria. G. 4197. Two adult specimens. Presented hj John Dennant, Esq. G. 5507. Two younger examples. Purchased. G. 9331. A series exhibiting stages of growth. Purchased. Ancilla semilsevis, Tenison-Woods (sp.). 1879. AncUlaria semilievis, T. "Woods, Proc. Linn. Soc. N.S.W. vol. iii. p. 229, pi. XX. fig. 7. 1889. AncUlaria semilcevis, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 148. 1893. AncUlaria semilcevis, Tate and Dennant, id. vol. xvii. pt. 1, p. 220. This shell has an acuminate spire, the callus is not strongly developed, and the sutures are bordered by a slight angulation producing a spiral ridge on the surface. The smooth zone is rather narrow, and the spiral band and denticle are well pronounced. It presents some analogy with A . arenaria, Cossmann, from the Eocene of the Paris basin ; the latter, however, is a smaller species, whilst the aperture of the shell is not so large relatively, and its columella is not excavated so deeply. Dimensions. — Length 18 mm.; breadth 7"5 mm. ; length of aperture 9 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4224. Three specimens. Presented hy John Dennant, Esq. G. 9333. Several examples showing stages of growth. Purchased. Ancilla lanceolata, Tate (sp.). 1889. AncUlaria lanceolata, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 147, pi. vii. fig. 2. The callus of this species presents some analogy with that of A. hehera, but the spire is contracted slightly at the apex; the shell is not so cylindrical, and is much more elongated. None of the specimens in the Museum exhibit the denticle ; its existence, 75 however, may be inferred from the feeble groove running round the anterior portion of the body-whorl. The columella is exca- vated, and for the size of the shell its plicated anterior extremity is not very large. Professor Tate remarks that this species is intermediate in character between A. hehera and Apseudaustralis. Dimensions. — Length 32 mm. ; breadth 10 mm. ; length of aperture 17-5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4199. An example of the senile stage of growth. Presented by John Bennant, Esq. G. 5509. One specimen, with columella abnormally excavated. Purchased. G. 9332. Two forms in which the spire is very callous. Purcliased. Ancilla papillata, Tate (sp.). 1889. Ancillaria papillata, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 146, pi. vii. fig. 4. This may be readily distinguished from all other Australian Tertiary Ancillte by the peculiar aspect of the callous growth around the spire, which is sculptured with narrow, sharply rounded, spiral ridges of unequal size, the larger with two or three smaller ones interposed. The spiral band, though deep, is very narrow, and terminates with a sulcated denticle. The shell is almost cylindrical. Dimensions. — Length 29 mm. ; breadth 12*5 mm. ; length of aperture 17 mm. Form, and Loc. — Miocene : Muddy Creek, Victoria. G. 4194. Two specimens. Presented hij John Dennant, Esq. G. 5508. Two adult examples. Purchased. G. 9324. A series exhibiting stages of growth. Purchased. Ancilla ligata, Tate (sp.). 1889. AncUlaria ligata, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. U7, pi. vii. fig. 6. This small shell resembles A. semihcvis, but it has a more obtuse apex, and is strongly angulate at the suture ; callus com- paratively thin, denticle not prominent. 76 Dimensions. — Length 1 7 mm. ; breadth nearly 6 mm. ; length of aperture 8 mm. Form, and Log. — Eocene: Aldinga, South Australia. G. 9325. A series exhibiting stages of growth. Purchased. Ancilla hebera, Hutton (sp.). 1873. Ancillaria hebera, Hutton, Cat. Tert. Moll. N.Z. p. 6. 1880. Ancillaria mucronata, Johnston, Geol. Tasmania, pi. xxxi. fig. 12 {non Sowerby). 1889. Ancillaria hebera, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 147, pi. vii. fig. 5. Although differing in several important particulars, this species is not unlike A. papillata. It lacks the solidity of the latter, however, whilst the spire is not sculptured in the same manner, the shell is not so cylindrical in shape, and the aperture is wider and proportionately larger. Professor Tate compares it with A. olivula, Lamarck, of the Paris basin, which it closely resembles, though, as he remarks, the callosity is larger, more gibbous and granulated in the Australasian fossil. The twisted anterior portion of the columella is relatively much smaller than in the European form. Dimensions. — Length 38 mm. ; breadth 13"5 mm. ; length of aperture 22 mm. Form, and Loc. — Eocene : Tasmania and Victoria. 83979. A tumid example, from Table Cape. Purchased, G. 4198. An elongate form, from Muddy Creek. Presented hy John Dennant, Esq. G. 9376. Several examples, showing stages of growth ; from Piver Murray cliffs. Purchased. Ancilla australis, G. P. Sby. (sp.). 1830. Ancillaria australis, G. B. Sowerby, Spec. Concliyl. p. 7, figs. 44-6. 1834. Ancillaria albisulcata, Quoy and Gaimard, Voy. de I'Astrolabe, Zool. t. iii. p. 19, pi. xlix. figs. .5-12 [non Sowerby). 1834. Ancillaria australis, Quoy and Gaimard, id. p. 20, pi. xlix. figs. 13-17. 1844. Ancillaria australis, Deshayes, in Lamarck, Hist. Nat. Anim. sans Vert. ed. 2, t. x. p. 592. 77 1S50. Ancillaria albisidcata, Gray, Fig. Moll. vol. iv. p. 4, pi. xix. figs. 1-3 («o« Sowerby). 1850. Ancillaria australis, Gray, id. vol. iv. p. 4, pi. xix. fig. 5. 1859. Ancillaria australis, Sowerby, Thes. Conch, vol. iii. p. 63, pi. ccxi. figs. 1, 2. 1864. Ancillaria australis, Reeve, Conch. Icon. vol. xv. pi. iii. figs. la-h. 1873. Ancillaria australis, Von Martens, Crit. List. Moll. N.Z. p. 23. 1873. Ancillaria [Anaulax) australis, Hutton, Cat. Marine Moll. N.Z. j). 17. 1873. Ancillaria australis, Hutton, Cat. Tert. Moll. N.Z. p. 6. 1880. Ancillaria australis, Hutton, Man. N.Z. Moll. p. 58. 1893. Ancillaria australis, Hutton, Linn. Soc. N.S.W., Macleay Mem. Vol. p. 44. The variable proportion of the callus on the spire, even in specimens from the same locality, causes this shell to assume a variety of forms. Sometimes it renders it mucronate, the callus being finely granular and extending over the posterior half of the columella, in which case it is impossible to observe the actual shape and number of the whorls. In other specimens, owing to the comparative absence of callus, the spire is almost acuminate, the suture being distinctly cut in, but in any case the protoconch is entirely covered up. The examples from Parimoa have the greatest thickness of callus on the spire above the body-whorl, and it is so disposed as to render the shell somewhat cylindrical with an angulate ridge above the anterior suture. In the two specimens from Glenelg river the callus is closely corrugated, leading to the formation of a number of spiral ridges ; an accidental mutilation of the spire of one of them reveals the protoconch, which, as far as can be seen, is composed of two turns, the earlier portion being very minute and the later somewhat inflated. The denticle on the outer margin of the aperture is practically a prolongation of a deeply-cut sulcus ; the anterior canal is broad and deep ; columella excavated and carrying several small plications anteriorly. In well-preserved specimens with but little callus, what is commonly termed the " varnished zone " of the body-whorl is light-brown in colour, in front of which is a band of the same tint, but darker. The whorl is finely striated, both longitudinally and spirally. Compared with the older A. pseudaustralis, the latter is more elongate, and its spire is more acuminate. Living examples of the species are found in Australia, New Zealand, and Tasmania. Some of the specimens of A. australis in the Museum very closely 78 resemble A. mucronata, Sowerby, and it is possible that the latter is a local variation of the former. Dimensions. — Length 41mm.; breadth 19'5 mm. ; length of aperture 25 mm. Form, and Loc. — Pliocene and post-Pliocene : New Zealand and Victoria. G. 5563. Two specimens of stunted growth, with callus spirally corrugated ; from Limestone Creek, Glenelg river. Purchased. G. 9521. Several fragments of the spire, exhibiting subangulate mucionation ; from Parimoa, Middle Island. W. B. D. Mantell Coll. G. 9514. A number of specimens showing wide range of variation, the spires of some being acuminate, of others mucronate ; from Wanganui. Sir James Hector Coll. G. 9520. An example of the adult, from Onekakara. Transferred from the Museum of Practical Geology. Family HARPID^. Genus HARPA, Lamarck. [Mem. Soc. Hist. Nat. Paris, 1799, p. 71.] Shell ovate, ventricose ; last whorl very large, ornamented with regularly spaced longitudinal ribs, prolonged over the suture and joined to preceding whorl; spire short; aperture wide, channelled anteriorly, columellar border callous, the callosity extending over part of the whorls, occasionally up to the protoconch ; columella twisted. Type. — Puccinum liar pa, Linnaeus. Subgenus EOCITHARA, Fischer. [Man. Conchy!. 1883, p. 601.] Columellar callosity more sharply defined and elevated than in Harpa [sensu stricto) ; posterior prolongations of the transverse ribs frequently flattened against and practically covering the suture. This, together with the subgenus Silia, Mayer, includes almost all fossil forms of the genus. Type. — Harpa mutica, Lamarck. 79 Harpa (Eocithara) lamellifera, Tate. . 1889. Harpa lamellifera, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 149, pi. vi. fig. 2. The protoconch of this species (Plate lY. Figs. 3«-J) is very different to that of either S. siilcosa or H. tenuis, descriptions of ■which follow. It is more depressed than that of the type of the genus, from which it differs also in being striated. It recalls the protoconch of the Pliocene and living Valuta musica, Linn., though it is not quite as large. Compared with the protoconch of specimens of H. mutiea, Lamk. (the subgeneric type), of the Parisian Eocene, it is relatively much larger and is more depressed, though of the same general character. The ornamentation of the whorls consists of thin, elevated, curved lamellae close together, the anterior portions of which are twisted so as to obscure the suture ; between each of these are three or four fine longitudinal lineations and many spiral stri« closely set, the latter being continued up the posterior side of the lamellae. The columellar border is covered with callus ; the anterior portion, however, is comparatively free therefrom, being sharply defined and elevated. The successive stages of growth of the anterior canal have left a well-marked, broad, rounded ridge, between which and the elevated columellar border is a deep rugose sulcus. Dimensions. — Length 34-5 mm.; breadth 23 mm.; length of aperture 27 mm. Form, and Loc. — Eocene : IMuddy Creek, Victoria. G. 4215. Two examples of the adult. Presented hy John Bennant, Esq. G. 9336. Two specimens of the neanic stage of growth. Purchased. Harpa (Eocithara) sulcosa, Tate. 1889. Harpa sulcosa, Tate, Trans. Roy. Soc. South Aust. vol. xi. p. 150, pi. vi. fig. 10. It is noteworthy that the subgeneric distinction previously observed should be emphasized also in the protoconchs of this 80 and its congener H. tenuis. The embryo of S. sulcosa has only two and a half turns visible, the nucleus appears to be covered over, and is slightly oblique, as exemplified by weathered speci- mens ; the last turn is longitudinally lineated, the earlier portion being smooth and shining. This species is readily distinguished from any other Harpa of the Australian Eocene by the flat area bordering the suture ; the transverse lamellae are very close together, and are exceedingly thin over the area alluded to. The columellar callosity is thin, but sharply defined, with the formation of a deep sulcation and perforation anteriorly. Dimensions. — Length 2.5 mm.; breadth 16 mm.; length of aperture 17 mm. Form., and Loc. — Eocene : Muddy Creek, Victoria. G. 4218. Specimen having protoconch abnormally developed. Presented hy John Dennant, Esq^. G. 9337. Two examples. Purchased. Harpa (Eocithara) tenuis, Tate. 1889. Sarpa tenuis, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 151, pi. vi. fig. 1. The anterior of the protoconch (Plate IV. Eigs. Aa-l) is slightly oblique, and almost entirely hidden by subsequent turns; there are two and a half of these latter, and the whole are micro- scopically granulated, the later turns being also transversely and longitudinally lineated. This is a larger species than S. lamellifera, and it has fewer longitudinal lamellae ; the interlamellar striae, both longitudinal and transverse, are not so prominent, except towards the anterior portion of the body-whorl, where a species of corrugation results ; the shoulder of the last whorl is roughly angulate, and this is accentuated by scaly projections from the lamellfe in that region. There seems to be a great deal in common between this and H. puUigera, from the blue clays at Schnapper Point ; judging from Professor Tate's figure {op. cit. pi. vi. fig. 9) it may, perhaps, be regarded as an extreme form of this species. Dimensions. — Length 36 mm. ; breadth 22 ram. ; length of aperture 28-5 mm. 81 Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 5511. An adult specimen, longitudinal costiB distantly separated. Purchased. G. 9335. Two examples, one of whicli illustrates the senile stage. Purchased. Harpa (Eocithara) abbreviata, Tate. 1889. Sarpa abbreviata, Tate, Trans. Eoy. Soc. South Aust. vol. xi. p. 150, pi. vi. fig. 7. Protoconeh (Plate IV. Figs. 5a-h) composed of two turns ; it has the general features of that of H. tenuis, but differs in that the initial portion is much inflated and is sharply defined from the remainder. The anterior portion of the turns may be divided into two parts, the earlier of which (like the first turn) is smooth, and the later is obscurely ribbed. The last-mentioned character is of much value in working out the phylogeny of the Gasteropoda ; the more pronounced generic characters were imparted to the shell whilst yet in the embryonic stage, prior to the escape of the animal from the capsule. In the brephic stage the incised suture is initially accompanied by an elevated border, which becomes modified as the animal approaches the neanic stage, and altogether disappears in the adult. The shell in the brephic stage also has some rather bold spiral elevations on the whorl, which, in conjunction with the closely-set longitudinal ribs, cause cancella- tion, but this is wanting in the adult form. A cursory examination might lead one to think that this species was merely a variation of H. tenuis, with which it could not possibly be confounded, however, when the protoconeh and shell in the brephic stage are compared. The ordinary spire-whorls are subangulated medially, and ornamented by rather distant, slightly elevated lamellae. Dimensions. — Length 24 mm.; breadth 14-5 mm. ; length of aperture 18-5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4216. Two specimens. Presented hy John Pennant, Esq. 82 MAEGIiVELLA. Family MARGlNELLIDiE. Genus MARGINELLA, Lamarck. [Mem. Soc. Hist. Nat. Paris, 1799, p. 70.] Shell oval or jblong; smooth; spire short, sometimes incon- spicuous ; outer margin frequently thickened and dentate ; columella plicate ; aperture slightly canaliculate anteriorly. Type. — Valuta glabella, Linnaeus. Marginella wentworthi, Tenison-Woods. 1877. Marginella weniworthii, Tenison-Woods, Pap. Roy. Soc. Tas. 1876, p. 109. 1878. Marginella wentworthii, E. Etheridge, jun., Cat. Aiist. Foss. p. 165. 1878. Marginella wentworthi, Tate, Trans. Phil. Soc. Adelaide, 1877-8, p. 92. 1888. Marginella wentworthii, Johnston, Geol. Tasmania, pi. xxxi. figs. 5, 5a. 1889. Marginella wentworthi, Dennant, Trans. Eoy. Soc. South Aust. vol. xi. p. 43. 1895. Marginella wentworthi, Tate and Dennant, id. vol. xix. pt. 1, p. 111. Protoconch composed of a globose, obtuse, smooth turn, barely distinguishable from the succeeding whorl. Shell small', elongate, and polished ; spire moderately elevated ; whorls five in number in adult specimens, convex, body-whorl obtusely angulate at the periphery, surface longitudinally striated by lines of growth, and obscurely spirally lineated, suture accompanied by an anterior spiral depression ; aperture elongate, somewhat inflated towards the middle ; outer margin thickened, furnished with a number of small crenulations terminated posteriorly by a rather large denticle, or tubercle, behind which is a broad, slightly canaliculate depres- sion ; inner margin with four prominent plications ; anterior portion of the aperture widely channelled. This species resembles M. contahulata, Deshayes, from the Middle Eocene of the Paris Basin ; but the latter is more lanceo- late, is minus crenulations on the inner side of the outer margin, and has bifid and divergent plications. Dimensions. — Length 7 mm. ; breadth 4 mm. ; length of aper- ture 4 mm. MABGINELLA. 88 Form, and Loc. — Eocene : Muddy Creek, Victoria. 6. 4213. A series exhibiting stages of growth. Presented hy John Dennant, Esq. G. 9341. Several specimens of the adult stage. Purchased. Marginella muscarioides, Tate. 1878. Marginella muscarioides, Tate, Trans. Phil. Soc. Adelaide, 1877-8, p. 91. 1889. Marf/inella muscar aides, Dennant, Trans. Roy. Soc. South Aiist. vol. xi. p. 43. This species is characterized by its broad aperture, which opens out anteriorly and is deeply channelled ; the body-whorl is gibbose, and well-preserved specimens exhibit inconspicuous, narrow, longi- tudinal bands of brownish tint. It differs from M. wentworthi in the characters above noted, as well as by the absence of crenulations on the outer margin. The latter in M. muscarioides is much thickened, especially towards the middle, where an obscure denticle is observable in some specimens ; the inner margin is somewhat callous, and carries lour conspicuous plications. Professor Tate remarks that in a broad sense this species is a diminutive M. muscaria, Lamarck, from which, however, it differs, amongst other things, in that the callous outer margin of the latter species is extended "on to the penultimate whorl, thence spreading over the base of the shell, and giving rise to a flattened appearance varicosely margined on the left side." The sutural region in M. muscarioides is somewhat depressed. Mr. Dennant quotes this species as occurring in the younger beds at Muddy Creek. Pimensions. — Length 7"5 mm. ; breadth -i'S mm. ; length of aperture 5 mm. Form, and Loc. — Miocene: Muddy Creek, Victoria. G. 4212. Several specimens. Presented hj John Pennant, Esq. G. 9338. Examples illustrating stages of growth. Purchased. 84 MAEGINELLA. Marginella winteri, Tate. 1878. Marginella tvinteri, Tate, Trans. Phil. Soc. Adelaide, 1877-8, p. 94. 1880. Mitra coarctata, Tenison-Woods, Proc. Linn. Soc. N.S.W. vol. iv. p. 8, pi. ii. fig. 10. 1889. 2[nr(jinella winteri, Dennant, Trans. Eoy. Soc. South Aust. vol. xi. p. 43. Shell elongate, subcylindrical, smooth ; protoconch obtuse, com- posed of two smooth turns, the anterior being disproportionately large ; spire elevated, three-whorled, body-whorl comprehensive, ornamented by longitudinal curved growth-lines, suture con- spicuous ; aperture elongate, tapering posteriorly, the sinus being very small, broad anteriorly, and channelled ; outer margin moderately thickened, but variable in individuals, closely crenu- lated within ; columella callous, especially towards the anterior, where five plications occur in the adult : these latter are close together, and only four are well developed, the fifth one, posteriorly, being very small, and altogether absent in young specimens. This is by no means a typical Marginella, but it does not fall conveniently into any of the recognized sections of the genus. The writer does not, however, feel inclined to add to these latter, already too numerous for systematic purposes. It differs from Marginella {sensu stricto) in its cylindrical shape and exsert spire, and by the possession in the adult of five columellar plaits. It has certain affinities with Volvarina, Hinds, ^ which, however, has only four plications on the columella, whilst its outer margin is not crenulated, at least in the type species, M. avena, Valenciennes. Specifically it closely resembles the living M. gracilis, C. B. Adams {=z M. hibalteata. Reeve), from the "West Indies, which is much smaller, more lanceolate, and is only quadriplicate. Dimensions. — Length 15'5mm. ; breadth 7 mm.; length of aperture 10-5 ram. Form, and Loc. — Eocene : Muddy Creek, Victoria. 1 Proc. Zool. Soc. 1844, p. 75. MAEGINELLA. 85 G. 4211. A large series of specimens showing stages of growth. Presented hij John Dennant, £sq. G. 9340. Four examples of the adult. Purchased. Marginella propinqua, Tate. 1878. Margiiiella propinqua, Tate, Trans. Phil. Soc. Adelaide, 1877-8, p. 94. 1832. Marginella propinqua, Kirk, Trans. N.Z. Inst. vol. xiv. p. 409. 1889. Marginella propinqua, Dennant, Trans. Roy. Soq. South Aust. vol. xi. p. 43. 189-5. Marginella propinqua, Tate and Dennant, id. vol. xix. pt. 1, p. 111. This species differs from M. winteri in being broader, and in having a much shorter spire. In studying the stages of growth, which are illustrated by an excellent series of specimens in the Museum, the following particulars concerning the columellar plications may be noted. In the neanic stage of some individuals four equal-sized, well-developed plaits occur, whilst in others the two posterior plaits are much smaller than the anterior. As the ephebic stage is attained a fifth, posterior plication appears, which is placed farther within the aperture of the shell, and is very small. In many individuals, however, only four plaits occur, even towards the gerontic stage, and one specimen in the collection possesses three large anterior plaits and one very small posterior. Another point of interest is the development of the crenulations on the outer margin. In the majority of specimens these are found from the neanic stage onwards ; but it is clear from others that crenulations were not developed, even up to the early portion of the ephebic stage, although the thickening of the outer margin took place under both conditions. The foregoing observations, in the opinion of the writer, are calculated to shed some light on the systematic value of the colu- mellar plications and the crenulations on the outer margin, in this group. It is customary with authors, dealing with Marginella, to lay especial stress on both these characters, in assigning species to subordinate divisions of the genus. Yet in M. propinqua these features are very unstable, and certainly ought not to rank higher than ontogenetic characters. Some authors might be disposed to create another specific name for the non-creuulate forms; but in carefully examining a series of the shells, it is 86 MAEGINELLA. impossible not to recognize the phenomena alluded to as mere variations of individuals of one species. The protoconch, the shape of the shell, form of aperture, the suture — in fact, every other characteristic except the number of plications and the occurrence of marginal crenulations — are identical in the forms here retained in M. propinqua. Biinensiotis. — Length 12 mm.; breadth 5"5mm.; length of aperture 9' 5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 9339. A series exhibiting stages of growth. Purcliased. Marginella inermis, Tate. 1878. Marginella inermis, Tate, Trans. Phil. Soc. Adelaide, 1877-8, p. 93. 1889. Marginella inermis, Dennant, Trans. Roy. Soc. South Aust. vol. xi. p. 43. 18'J5. Marginella inermis, Tate and Dennant, id. vol. xix. pt. 1, p. 111. Shell pyriform, attenuating anteriorly, smooth ; spire very short, composed of from two to three whorls, body-whorl comprehensive, extending almost the whole length of the shell, longitudinal growth- lines conspicuous, suture well defined, polished ; aperture elongate, broadly rounded posteriorly and slightly produced anteriorly where it is widely channelled ; outer margin thickened and crenulated, though comparatively smooth towards the posterior extremity ; columellar margin furnished with four plications in the majority of individuals, though one adult specimen in the Museum has only three, and another three with the trace of a fourth. Its shape, the elongation of the aperture, and its rounded form in the neighbourhood of the posterior sinus, serve to readily distinguish 21. inermis from all other Australasian Marginella!, Except for the dentated outer margin, this species might perhaps be included in the subgenus Prunum. Dimensions. — Length 12 mm.; breadth 6*5 mm. ; length of aperture 10'5 mm. Form, and Loc. — Eocene : Muddy Creek, Victoria. G. 4207. Several examples of the adult. Presented hy John Pennant, Fsq. MAEGINELLA. 87 Subgenus GLABELLA, Swainson. [Malacology, 1840, p. 324.] The principal distinguishing features of this subgenus are the more or less conical form of the spire, which is well developed, and the longitudinal plaits on the peripheral subangulation of the body-whorl present in the majority of the species. The first species mentioned by Swainson is quoted as "P. ruffina, Sw." ; as it is not clear what he intended by that specific name, M. hifasciata, Lamarck (one of the typical species mentioned by Swainson), is here adopted as the type of this subgenus. In reference to M. ovata and M. cornea, two new species here described from the Tertiary of New Zealand, it may be noted tliat certain kinds of Marginella have been recognized in the newer beds of that colony ; but for the most part these have neither been figured nor described, unless, indeed, two or three lines of remarks can be called a description. The present writer has not been able to identify either of the species above mentioned with the meagre descriptions alluded to, and is of opinion that these latter ought to be suppressed as being of no real value to systematists. Type. — Marginella lifasciata, Lamarck. Marginella (Glabella) turbinata, G. B. Sowerby. 1846. MarijincUa turbinata, G. B. Sowerby, Thes. Conch, vol. i. p. 3S.5, pi. Ixxv. figs. 70-1. 1865. Marghuila turbinata, Reeve, Conch. Icon. vol. xv. pi. sxii. fig. 122. 1867. Marginella turbinata, Angas, Proc. Zool. Soc. Lond. p. 196. 1878. Marginella turbinata, Tate, Traus. Phil. Soc. Adelaide, 1877-8, p. 86. 1879. Marginella turbinata, Weinkauff, Conch. Cab. (ed. Kiister), p. 86, sp. 115, pi. xvi. figs. 9, 12. 1883. Marginella [Glabella) turbinata, Tryon, Manual of Conchology, vol. v. p. 23, pi. vii. figs. 4, 5. 1886. Marginella {Glabella)