FLORA OF WINNESHIEK COUNTY BY B, SHIMEK FROM ilUwaA GEOLOGICAL SuRVEY, VOL. XVI. ANNUAL REPORT, 1905 yy 147-211. DES MOINES 1906 ALBERT R. MANN LIBRARY New YorK STATE COLLEGES OF AGRICULTURE AND HomE ECONOMICS AT CORNELL UNIVERSITY ak 160.863" University Library WMA 3 1924 001 280 5 22 mann Cornell University The original of this book is in the Cornell University Library. There are no known copyright restrictions in the United States on the use of the text. http://www.archive.org/details/cu31924001280522 THE PLANTS OF WINNESHIEK COUNTY. 147 ” THE PLANIS OF WINNESHIEK COUNTY. ; BY B. SHIMEK. Winneshiek county presents a flora of unusual richness. Both because of its geographic position and its varied surface features it offers conditions which have made possible the development of a variety of plants scarcely equalled in any other county of the state. Its northerly position and its rough topography, especially along the Upper Iowa or Oneota river have brought a northerly flora, such as belongs to the heavily wooded regions of Minnesota and Wisconsin; its wooded knobs and ridges along the Turkey and the head waters of the Yellow rivers remind one of the rough wooded areas of southern Iowa; while the prairies are but a continuation of the greater prairies of the west. Each of these territories presents a variety of conditions. The roughest includes the driftless area and the drift border, with their deep gorges with narrow alluvial bottoms, their exposures of both sandstones and limestones, their shaded mossy banks and wooded slopes with not infrequent small bogs, and their drier wooded ridges with occasional treeless barren summits ; the more southerly timbered ridges present somewhat similar con- ditions, but without the prominent rock-exposures, and with floral areas less sharply defined than those which characterize a region cut by deep gorges; and the treeless areas include both the rich level prairie and the prairie bogs of the Iowan drift, and the more rolling and drier prairie of the Kansan. The soils are derived chiefly from loess and drift (which are discussed. else- where in this report), and therefore vary comparatively little in chemical ‘composition. Occasionally coarser, sandy material appears on tke ridges, but on the whole a finer soil prevails, 314288 148 FLORA OF WINNESHIEK’ COUNTY, , its fertility, however, varying with topography and plant-cover- ing, and with the consequent possibility of retaining the humus- laden surface soil. Alluvial tracts are not, for the most part, of great extent in the county, being limited by the narrow valleys. In the rougher parts sandy and rocky tracts are frequent. The investigations upon which this report is chiefly based were carried on largely during the spring, summer and fall of 1903. Every township in the county was visited, and certain typical localities were more thoroughly studied. Thus Moneek and Ft. Atkinson were selected for the study of the flora of the rougher territory of the two principal river systems in the southern part of the county; the vicinity of Calmar for the southerly prairie, and Orleans township and Bluffton for that in the western part of the county; for the rougher wooded region the entire gorge of the Oneota was traversed a number of times across the county, Decorah, Bluffton and Kendallville being the main centers of investigation, while the hilly country bordering Canoe creek and Bear creek received some attention; and the vicinity of Hesper, with its woods and prairies, and unique exposures of St. Peter sandstone proved the richest and most interesting single locality in the county. The flora of this county had received some previous attention. Arthur’s ‘‘Flora of Iowa’’* contains numerous references to plants from this county, which are based, at least in large part, on material furnished by Mrs. M. C. Carter of Hesper, and Mr. K. W. D. Holway, formerly of Decorah; individual references are also made to Winneshiek county plants by a number of other authors who did not secure the material themselves, but received it from others—chiefly from Mrs. Carter, and Messrs. E. W. D. Holway, Herbert E. Goddard, Thos. E. Savage, Alois Kovarik, and EK. Orr.+ *Contributions to the Flora of Iowa, J. C. Arthur, 1876, with continuation, Nos II VI in the Proceedings of the Davenport Academy of Natural Sciences, Vols. III and IV {Such are the following: Ae ae Cerastium arvense var. oblongifoli i : i i To e ‘Bot Cin, Vor SEV, vi ifo umn, var., Hollick and Britton, Bull. . otes on the uatic —R. I. 7 5 . Hi State Univ ot pe aauati ai epeeama of Iowa,—R. I. Cratty, Bull Lab. Nat. Hist., An illustrated Flora, etc,, Britton and Brown. 8 Vols. , 1896-1898. Vol he Uyperaces of Towa, R.1. Cratty, Bull. Lab. Nat Hist , State Univ. of Towa, The lowa Pteridophyta, B. Shimek. ibid., Vol. V,, 1901 i Pgh a ryptogams of Iowa, — . Ta. - Bei. % Ix, 02 also reprint, as Cont. No, 20, Bot. Dap, Ta. B. Col. of A. and MA a ‘Bos Wal otany of the No. U. 8.,— Asa Gray, several e DB. Manual of the Flora of the U. S.,—N. L. Britten, ist ed., 1901, : and ed., 1905, THE PLANTS OF WINNESHIEK COUNTY. 149 Aside from the resident students of botany the most extensive collectors have been the following: Mr. T. J. Fitzpatrick, who made a trip down the Upper Iowa in company with Dr. Paul Bartsch in the summer of 1895. He has published, in part jointly with Mrs. Fitzpatrick, a number of reports on the plants collected,* including, however, a large number of plants obtained from other collectors. Prof. Thos. E. Savage, who visited the county in the spring of 1899. A small part of his collection was noted in some of the references cited, but the greater part is here reported for the first time. The present writer, who made a number of trips to the county in 1902 and 1903. His collections and those of Prof. Savage are now in the Herbarium of the State University. They include a large number of lower cryptogams, especially fungi, lichens and mosses, a report upon which cannot be here ineluded. Two short reports treating of the plants of the county have been published by the writer,* and in the present paper it is purposed to report upon all the plants obtained in the field, and also to include such as have been definitely reported from the county by others. All the papers treating of Winneshiek county plants which have been heretofore published, have been little more than mere annotated lists. Very little attention has been paid to the eco- nomic value of the native plants, though this is often note- worthy, and the public has treated them rather as an undesirable encumbrance which must be removed as soon as possible to make way for the plants of the field and pasture. Aside from the use which we may make of individual plants or their products, the native flora has a vastly more important function whose influ- ence extends far beyond the limits of any particular tract of land upon which it is developed, especially in a territory with a much broken surface such as is found in this county. It develops a better soil, conserves moisture, and prevents erosion and the *Notes on the Flora of Northeastern Iowa,—Proc. Ia. Acad. Sci. «» Vol. V, 1898. Manual of the Flowering Plants of Iowa, 1899. : The Orchidaceze of lowa,—Proc. Ia. Acad Sci., Vol. VII, 1900. The Scrophulariacez of Ilowa,—Proc. Ia Acad. Sci., Vol. X, 1903. Hi ae Ploragh of the St. eter Sandstone in Winneshiek County, Iowa, Bull. Lab. Nat. is ¥ . Notes on Some: Towa Plants, —Proc. Dav. Acad Sci., Vol. X, 1904. ‘150 FLORA OF WINNESHIEK COUNTY. consequent clogging of our streams with sand and mud. It is superior to any aggregation of cultivated or pasture plants for the reason that it is not removed to leave the soil bare during a large part of the year, as is the case with cultivated crops, and it is not eaten or tramped upon by cattle as in the | case of pastures. Moreover, through long adjustment. to existing con- ditions the native plants are perfectly adapted to the various surfaces upon which they occur, and they have become more pliant under the varying moods of our uncertain climate. They are, therefore, more certain of persisting, and thus ‘continuing their benificent influence. In a rough country the dangers from erosion and desiccation are great, not only to the land itself, but to the streams and water-courses. The steeper slopes should never be cleared of their covering of native vegetation, either by cultivation or by pasturing, for not only will they not be profit- able, but they will be a menace to better lands and to valuable springs and streams. But aside from these uses to which all our native plants lend themselves more or less readily, many of them more directly, and therefore more appreciably, affect the interests of man. Among the questions to which this usefulness of plants gives rise may be mentioned the problems of the forests, weeds, ornamental plants, medicinal plants, and consideration of rusts, smuts and other fungi which attack plants. The last subject has received much attention locally from Mr. Holway and will not be dis- cussed here. I. The Forest Problem. Originally not less than one-fourth of the surface of Winne- shiek county was covered with forest. This was sometimes ‘secant, as upon the rocky slopes and drier hill-tops, or consisted of trees of but little value, as upon the narrow lower bottom lands. Here, as elsewhere, the forest was developed chiefly upon poorer soils. The sandy alluvial bottom lands, the rocky slopes, the gravelly or clayey hills—these formed the favorite habitat of trees. Even where a veneer of rich soil and leaf-mould appeared it was the effect rather than the cause of the forest. The forest prevented erosion; it retained moisture which made easier the disintegration of both organic and inorganic mater- THE FOREST PROBLEM. ~’ 152 ials; it annually contributed its leaves to the accumulating soil; it harbored worms and other burrowing animals which brought fine soil-materials to the surface; and in its shelter the burden of dust-laden winds was deposited. So man thought that he saw alluring promise in the richness of the forest soil, and this coupled with the prospect of immediate gain from its products, led him to remove the forest. But an awakening has already come, and men realize that with the removal of the riches of the forest they also lose the richness of the soil, for the rains and melting snows quickly strip it from the hillsides. The land is then practically worthless, for it will make neither field nor pasture—it is fit only for growing trees, as it has grown trees in the past. Few counties in the state have suffered more than Winneshiek in this respect. The principal forest areas were in the roughest territory, unsuited to the ordinary purposes of agriculture. Man’s greed and thoughtlessness combined in many cases to strip the best, if not all, of the forest from these hill- sides, but this was not the gravest error, for if left to its own resources the forest would renew itself. But an attempt was made in many cases to cultivate or pasture the stripped areas, and this was done on the steepest slopes with uniformly dis- asterous results. More acres were cultivated that still. other acres might be secured, under the pretext that the children of the land-holder must not be left without inheritance. The desire for immediate gain was, however, responsible for this, for men had not yet learned that a growing forest is one of the most splendid legacies which they may leave to their children. The best of the legacy which we ourselves received has been dis- sipated, in Winneshiek county as elsewhere, but conditions for the renewal of the forest are here very favorable. There are three distinct forest areas in the county, drained respectively by the Upper Towa, the Turkey and the Yellow rivers. Of these the first is much the largest within the county, but all agree in having a relatively large amount of adjacent rough land, as has been noted. Much of this land is worthless, or at least of little value, for agricultural purposes, and it would entail but little present loss, and would vastly benefit posterity, if given over to the cultivation of timber. This would not require a large amount 152 ‘ FLORA. OF: WINNESHIBK COUNTY. of expensive nursery stock, nor great: labor im the: cultivation of the soil, nor assiduous attention to the welfare of the trees. The territory is so well adapted to the growth of trees that it is: necessary only to strew seed on the ground in suitable places, and to keep out stock and fires, and the forest will take care of itself. For this purpose seed should: be gathered: in considerable quantity, preferably from nearby trees, and should not be per- mitted to become too dry. Seed maturing in spring or early summer may be scattered at once if the land is not devoid of other vegetation, or it may be covered with a thin layer of sand or soil in a shady place and left until fall. Seeds. and fruits which mature in autumn should be scattered just before the leaves begin to, fall. They will thus be better protected both against the: severity of winter, and against squirrels and chip- munks., Less than one-half the: seed. sown. in this manner will germinate, and: for that. reason it should: be. scattered. liberally. With a little care it may be secured in abundance from native or acelimated. trees, practically without expense, in connection with a, pleasant summer or autumn outing. If a little trouble is taken, each year to: re-seed where: the stand is. thin, to remove. rotting wood in order that it may not spread its infection by: scattering countless. spores of fungi, to trim out excessive under- brush, and to protect the trees against domestic animals and. fires, the forest, will be a vast improvement upon. those which the first: settlers; found. Native trees. will be found most satisfactory for reforestation, for as yet no introduced forest tree has demonstrated: its superiority over the native species. Moreover, in a county as well timbered as this both quantity and variety may be secured: with comparative ease. In order that some conception of the range of choice: may be. given, a list of the native trees. and shrubs: is here presented, the species being grouped. according to habitat. In selecting:trees for any tract it is well that heed be: given to the character of its surface in order that an appropriate choice may be made from species occupying similar stations elsewhere. The following list contains the native trees and shrubs: of the county. Their distribution, abundance, full scientific names, TREES AND SHRUBS. 153 ete., may be determined from the systematic list which makes up the closing part of this paper. TREES AND SHRUBS. 1. Species belonging to bogs and wet places.—The species are shrubs, the last two, only, becoming small trees. They are: the meadow-sweet (Spiraea), red-osier dogwood (Cornus stoloni- fera), button-bush (Cephalanthus), shiny willow (Salia lucida) and pussy. willow (8. discolor). 2. Species of the low alluvial bottoms.—The shrubs are: the false indigo (Amorpha fruticosa), sandbar willow (Salia interior) and heart-leaved willow (9. cordata), the willows some- times becoming small trees. The soft maple (Acer saccharinum), red birch (Betula nigra), almond-leaved willow (Salia amygdaloides) and black willow (8. nigra) are trees. 3. Species of higher rich alluvial bottoms.—The wild grape (Vitis vulpina) and poison ivy (Rhus radicans) are woody vines, the latter sometimes a low shrub; the elder (Sambucus can- adensis), flowering currant (Ribes floridum) and the wahoo (Euonymus) are shrubs, the last sometimes a small tree; all the species of haws (Crataegus) and the sheep-berry (Viburnum lentago) are small trees, the latter sometimes a shrub; and the following species are trees: the box-elder (Acer negundo), hack- berry( Celtis), the ashes (Fraxinus, except F’. pennsylvamica), honey locust (Gleditsia), the butternut and walnut (Juglans), cottonwood (Populus deltoides) and white elm (Ulmus americana). 4. Species of upland thickets—-The Missouri gooseberry (Ribes missouriensis) and the hazel (Corylus americanus) are shrubs, and the haws (Crataegus) and wild crab (Malus) are small trees. 5. Species of wooded rocky banks and open slopes.—The fol- lowing are woody vines: the purple virgin’s-bower (Atragene), climbing bittersweet (Celastrus) and the two honeysuckles (Lonicera). The shrubs are: the American yew (Taxus), shad- bush (Amelanchier botryapium), the dogwoods (Cornus alterni- folia, amomum, aspcrifolia and circinata), shrubby cinquefoil (Dasiphora), dwarf cherry (Prunus pumila), leather-wood (Dirca), prairie nine-bark (Opulaster), wild gooseberry (Ribes 154 FLORA OF WINNESHIEK COUNTY. cynosbati), the wild roses (Rosa blanda and humilis), wild black raspberry (Rubus occidentalis), dewberry (Rubus procumbens), wild red raspberry (R. strigosus) red-berried elder (Sambucus pubens), bladder-nut (Staphylea), high bush-cranberry (Vibur- num opulus) and arrow wood (Viburnum dentatum). The fol- lowing species are often shrubs,’ but also become small trees: hoary alder (Alnus), alder-leaved. June-berry (Amelanchier alnifolia), round leaved June-berry (A. rotundifolia), panicled dogwood (Cornus candidissima), choke cherry (Prunus virgin- jana), staghorn sumach (Rhus hirta), prickly ash (Xanthoxy- lum) and black haw (Viburnum lentago). The June-berry (Amelanchier canadensis) and blue beech (Carpinus) are small trees. The following species are forest trees: the balsam (Abies), red cedar (Juniperus viginiana), white pine (Pinus), hard maple (Acer saccharum), cherry birch (Betula lenta), canoe birch (Betula papyrifera), red ash (Fraxinus pennsylva- mica), balm of Gilead (Populus candicans), American aspen (Populus tremuloides) and rock elm (Ulmus racemosa). _, 6. Species of upland, mostly rather open woods.—The iron wood (Ostrya) is a small tree, and the following are forest trees: the hickories (Hicoria), large-toothed aspen (Populus grandidentata), all the oaks (Quercus), the white oak (Q. alba) preferring deep woods, and red elm (Ulmus fulva). 7. Species of deep upland woods——The hispid greenbrier (Smilax hispida) and the Virginia creeper (Parthenocissus) are woody vines; the following are trees: red oak (Quercus rubra), white oak (Q. alba), black cherry (Prunus serotina) and basswood (Tilia). All the species of the preceding group may also be found in deeper woods. 8. Species of prairie .and forest borders——Most of the species of this group are shrubs, as follows: juniper (Juniperus communis), lead plant (Amorpha canescens), New Jersey tea (Ceanothus americana), red root (Ceanothus ovatus pubescens), bush honeysuckle (Diervilla), prairie rose (Rosa arkansana), wild rose (R. humilis), prairie willow (Salix humilis), and wolf-berry (Symphoricarpos occidentalis). The blackberry (Rubus nigrobaccus) is also found along borders, but extends to deep woods, and the smooth sumach (Rhus glabra) is found along borders, in open woods, on dry slopes, etc. But one small “TREES AND SHRUBS. 155 tree, the wild red cherry (Prunus pennsylvanica) , properly belongs to this group. ' In addition to the foregoing species, which are native, the red currant (Ribes rubrum) and the black locust (Robinia pseud- acacia) may be found in waste places or openings, having escaped from cultivation. The red currant appears to be native. Not ali the species here listed are of like value, nor are all ‘usable for the same purposes. The most useful lumber trees are the ashes, butternut and walnut, red and rock elms, cherry ’ birch, black cherry, the hickories, the oaks, red cedar and white pine. For ties and posts the white oak, bur oak, honey locust and black locust are most servicable. The most valuable native shade trees are the hard maple, hackberry, ashes, American ‘elm, basswood, canoe birch and white pine. The most useful species for windbreaks and shelter belts are the soft maple, ashes, boxelder, cottonwood, balm of Gilead and red cedar. All of these are used more or less for fuel. Among the native ornamental trees and shrubs the hard maple, Virginia creeper and the sumachs are remarkable for the splendor of their autumn foliage; the bittersweet for its bright fruit; the June-berry, crab-apple, hawthorns and black locust for their flowers; the false indigo, honeysuckles, wild roses, elderberries, atragene, bladdernut, dogwoods, black haw, sheep-berry, New Jersey tea and nine-bark for flowers and foli- age; and the white pine, balsam fir, shining willow and red- osier dogwood for the beauty of the foliage and crown. As already noted these native species are eminently suitable for all purposes for which trees and shrubs. are used, but never- theless, numerous species have been introduced in cultivation for ornamental purposes. Among the introduced conifers are the Scotch and Austrian pines, Norway spruce, European larch, arbor-vite and hemlock, and among the deciduous trees the silver poplar, Lombardy poplar, buckeye, catalpa, mountain ash and white willow. In this connection a report on a tree-census of a portion of the city of Decorah, including several blocks and streets in the residence part, may be of interest. The number 156 FLORA OF WINNESHIEK COUNTY. to the right in each case indicates the number of trees found within the territory selected: Hard maple ......... ... 97 Red cedar ..........-..- 11 Olt Maple ..esscwivess 86 Balsam fir ...........-- 7 Box elder .............. 64 Walttt sco; cence acess 5 American elm ........... 62 Red lM .a.ce