ALBERT R. MANN LIBRARY AT CORNELL UNIVERSITY CORNELL UNIVERSITY LIBRARY \\ Cornell University Library The original of this book is in the Cornell University Library. There are no known copyright restrictions in the United States on the use of the text. http://www.archive.org/details/cu31924089569143 SELECTED WESTERN FLORA MANITOBA, SASKATCHEWAN, ALBERTA MACMILLAN’S CANADIAN SCHOOL SERIES SELECTED WESTERN FLORA MANITOBA, SASKATCHEWAN, ALBERTA BY B. J. HALES, B.A. PRINCIPAL, NORMAL SCHOOL, BRANDON AUTHORIZED BY THE ADVISORY BOARD OF THE DEPARTMENT or Epucation For Usr 1x Hicu ScHoois anp | CoLLeGiIatE INSTITUTES IN MANITOBA Toronto THE MACMILLAN COMPANY OF CANADA, LIMITED ; 1919 CopvricutT, CaNaDA, 1915 By THE MACMILLAN COMPANY OF CANADA, Limtro REPRINTED, 1916 PREFACE Tus Flora contains 81 families, 291 genera, and more than 650 species. -While making no claim to being a complete flora of the territory covered, it is believed that it contains all the species likely to. be studied by students in the High Schools and Collegiate Institutes of the Prairie Provinces. The arrangement of families follows the order of Engler & Prantl which has now been almost universally adopted, and the nomenclature has been made to conform as closely as possible to the Vienna rules. The capitalization is that of the Index Kewensis, which is now being largely followed, and differs from that of many of the earlier floras in that adjectives derived from names of places are spelled without the capital. : In- the descriptions the aim has been to introduce only such character- istics as can be observed by the beginner, and as far as possible to bring out the points of contrast between species found in the same locality. While the descriptions have mostly been written with the plant in hand, yet full reference has been made to all the well-known floras covering the area. The author wishes especially to acknowledge the help derived from the following: ‘‘ Gray’s New Manual of Botany,’ revised by Rob- inson and Fernald ; ‘‘Illustrated Flora,’’ Britton & Brown, both first and second editions; Gray’s ‘‘Synoptical Flora of North America”? ; Piper’s ‘‘ Flora of Washington’’ ; and ‘‘ A New Manual of Rocky. Moun- tain Botany,” Coulter & Nelson. .. Thanks are also due to the following persons for help: Miss M. E. Moodie and Mr. G. S. Johnson for checking the lists for Alberta and Saskatchewan, respectively; Mr. D. W. Yuill for trying out and check- ing the key ; Professor B. L. Robinson, Harvard University, for confirm- ing a list of names; Mr. J. M. Macoun, Curator of the Herbarium of the Geological Survey, Ottawa, for constant and invaluable assistance in fur- nishing descriptions and confirming the determination of species; and to many ex-students of my own who have so kindly sent specimens for identification from all parts of the Canadian West. Special mention should also be made of the assistance received from Professor John Macoun’s ** Catalogue of Canadian Plants.’’ B. J. H. BranNpDon, August 1, 1915. \ ANALYTIC KEY TO THE FAMILIES OF THE COMMON PLANTS OF MANITOBA, SAS- KATCHEWAN, AND ALBERTA ‘(Carried out to genera where the genus differs in characteristics from the family.) DIVISION I. PTERIDOPHYTA Fern-like or rust-like plants without true flowers and reproducing by means of spores. a. Rush-like plants with jointed stems, the joints surrounded by con- spicuous toothed sheaths ; the stems terminating in conical spikes. Equisetacee, p. 4 Aa. Fern-like plants with leaf-like fronds. Fronds from a creeping rootstock . . . . . Polypodiacee, p.1 Fronds from fleshy roots . . . . . . «. Ophioglossacee, p. 2 DIVISION II. SPERMATOPHYTA Plants with true flowers and reproducing by means of seeds. SUBDIVISION I. GYMNOSPERMA Trees or shrubs with needle-shaped leaves and ovules not enclosed in an ovary. a. Flowers in catkin-like spikes which become conesin fruit . Pinaceae, p. 4 SUBDIVISION II. ANGIOSPERM Plants having the ovules borne in a closed ovary which matures into a fruit. , Cuass 1. MONOCOTYLEDONEZ Stems, when present, without central pith or annual rings; seeds with only one cotyledon; leaves usually straight-veined. vii \ Vili ANALYTIC KEY TO THE FAMILIES a. Small, elliptical or flask shaped, aquatic plants without’ true stems or leaves, and swimming freely on the water . Lemnacee, p. AA. Plants with true stems and leaves, the latter sometimes reduced to scales. B. Perianth free from the ovary or wanting.. u. Perianth wanting or scale-like. Dp. Flowers enclosed in husk-like scales (glumes) ; grass-like plants with jointed stems, sheathing leaves and 1-seeded fruit. : Stems hollow; leaf-sheaths split . . Gramineae, p. Stems generally solid, often triangular; leaf-sheaths not split... . ¥ 4) @) Cyperacee, p. pp. Flowers not enclosed in planned, Rush-like marsh plants with an ovary of 3 carpels Juncacee, p. Leafy immersed plants with perfect flowers Najadacea, p. Flag-like marsh plants with flowers in dense cylin- drical spikes & oslo a a 4 Typhacee, p. cc. Perianth always present, either colored or herbaceous but never scale-like. aw. Pistils numerous in ahead orring . Alismacea, p. EE. Pistil one; compound. Stamens 3; marsh herbs with rush-like leaves and flowers in spikes . . . . Juncaginacea, p. Stamens 4 SA ud: yee ta “see Ae MIANTHEMUM, p. Stamens 6; Divisions of the perianth alike. Rush-like plants . . . . . . Juncacee, p. Notrush-like . . .. . . . Liliaceae, p. Divisions of the perianth not alike; leaves net- Weinéd 5 « ¢ » & w= G TRILLIUM, Dp. BB. Perianth adhering to the ovary. Immersed aquatic herbs . . . . . . Hydrocharitacea, p. Terrestrial herbs. Stamens 2; flowersirregular . . . . . Orchidacee, p Stamens 3 coe ew ew ew ew ww...) Tridacee, p. Stamens6. ...... . . . . Amaryllidacee, p Cuass 2. DICOTYLEDONEZ Stems with a central pith; if woody, the wood arranged in annual rings; leaves net-veined; embryo with two cotyledons. A. Corolla absent. B. Flowers moncecious or dioecious; one or both kinds in catkins or catkin-like clusters. \ 14 “. ANALYTIC KEY TO THE FAMILIES . x Fertile flowers, only, in catkins . . . . . .'Urticacea, p. 32 Sterile flowers, only, in catkins ; ; the fertile iu a guplket in- volucre . . . woe . Fagacee, p. 31 Both sterile and fertile Bowers i in ontistas: Ovary many-ovuled . . . . . « Salicacee, p25 Ovary 1-2-seeded ; cell eavulad: Calyx regular, succulent in fruit. . . . Urticacee, p. 32 Calyx wanting; two or three fertile flowers in the axil of each scale . . : as tases . Betulacee, p. 28 BB. Flowers not in catkins. v. Ovary or its cells 1—4-ovuled. p. Pistils more than one, distinct. Stamens on the calyx; leaves with stipules . Rosacea, p. 60 Stamens on the receptacle; petioles spreading at the base Ranunculacee, p. 44 pp. Pistil one, either simple or compound. B. Ovary superior. ‘ Stipules sheathing the stem . « « Polygonacee, p. 35 Stipules not sheathing the stem. F. Herbs. Style or stigma 1; flowers unisexual Urticacea, p. 32 Style 2-3-branched ; inflorescence not a spike Euphorbiacee, p. 81 Flowers small in dense heads or spikes accompanied by scarious or colored bracts Amaranthacee, p. 39 Flowers small; densely clustered; no scarious bracts . . . . . . . Chenopodiaceae, p. 37 ¥. Shrubs or trees. Leaves opposite. , Fruit 3-celled » oo. « « « « Rhamnacee, p. 85 Fruit a double samara . . . . Acerace@, p. 83 Fruit a single samara . . . . . .Oleacea, p. 107 Leaves alternate ; Styles2 . . . « . .« Urticacee, p. 32 EBB. Ovary inferior or apparently so. Aquatic herbs . . ... . Haloragidacee, p. 96 Terrestrial. Herbs with colored calyx. 2 Leaves opposite, simple . . .Nyctaginacee, p. 40 Leaves alternate, simple . . . COMMANDRA, p. 34 Shrubs. Leaves silvery or brownish scurvy Eleagnacee, p. 93 cc. Ovary or its cells many-ovuled. bp. Ovary superior; calyx present. Ovaries 2ormore . .. .. .. . . Ranunculacee, p. 44 Ovary single, of 1 or 2 cells. xX ANALYTIC KEY TO THE FAMILIES Calyx of separate sepals . . . . . Caryophyllacee, p, 40 Calyx 5-toothed . . . . . .. . . .).) 60@LUX, p. 105 aa. Calyx and corolla both present. B. Corolla of separate parts. c. Stamens more than twice as many as the sepals. pb. Calyx free from the ovary. EB. Pistils several or many ; distinct. Aquatic herbs with peltate leaves . Nymphacee, p. 44 Terrestrial plants. Filaments united in a tube . . .« Malvaceae, p. 89 Filaments not united, inserted on the calyx Rosacea, p. 60 BB. Pistils forming 1 ovary but may have several styles or stigmas. Ovary 1-celled, simple. - Ovules2 . ... . . =. . . . £Rosacee, p. 60 . Ovulesmany .. .. . . Ranunculaceae, p. 44 Ovary compound. Ovary 1-celled. Sepals 2; succulent plants with watery juice _and free central placenta . Portulacacee, p. 43 Sepals 3-5; placenta parietal . . Cistacee, p. 89 Ovary several-celled. Terrestrial herbs; filaments united forming a | tube. . . . . . « . Malvacee, p. 89 Aquatic or marsh adh: Leaves pitcher-shaped . Sarraceniacee, p. 57 Leaves peltate . . . . . Nymphacee, p. 44 Trees with 2-celled anthers , . . Tiliacee, p. 89 pp. Calyx adherent to the compound ovary. ‘Ovary 1-—5-celled. Fleshy-stemmed plants without true foliage Cactacee, p. 92 Leaf-bearing plants. Calyx 2-parted . . . . . . . Portulacacee, p. 43 Calyx of more than 2 parts. Leaves opposite, without stipules Sazifragacee, p. 58 Leaves alternate, with stipules. . Rosaceae, p. 60 Leaves opposite, with Liisa rough-leaved herbs. . . ‘4 . . Doasacee, p. 92 cc. Stamens not more than twice as many as the petals. p. Stamens of the same number as the petals and opposite to them. Ovary one. “Ovary 2-5-celled. 238 Calyx minute, petals valvate in bud . Vitacee, p. 86 Calyx 4~5-cleft, foldedinwardin bud Rhamnacee, p. 85 ANALYTIC KEY TO THE FAMILIES Ovary 1-celled. Anthers opening by lids . . . . Berberidacea, p. Anthers not opening by lids. Style 1, unbranched, stigma 1 Primulacee, p. pp. Stamens not of the same number as the petals, or if of the _ same number alternate with them. Bu. Calyx free from the ovary. F, Ovaries 2 or more, separate or slightly united. Stamens united, with a large common stigma Stamens free from each other. Stamens on the receptacle. Leaves fleshy . . “ . - Crassulacee, p. Leaves not fleshy. . Ovary 5-lobed with 1 style Primulacee, p. Ovaries distinct, styles and stigmas sepa- rate. . . . . . Ranunculacee, p. Stamens on the calyx. With stipules . . . . . . Rosacea, p. Without stipules . . . . Saxifragacee, p. FF. Ovary 1. : Ovary simple with 1 parietal placenta. Leguminose, p. Ovary compound. a. Ovary 1-celled. Corolla irregular. Petals 4, stamens6 . . . Fumariacee, p. Petals and stamens5 . . . Violacea, p. Corolla regular. Ovule solitary. Shrubs ©. . . . . Anacardiacee, p. Herbs... . . . . « Crucifere, p. Ovules not solitary. | Ovules at the bottom of the cell Caryophyllacee, p. Ovules attached to the sides of the cell. Leaves glandular hairy Drosoracee, p. Leaves not glandular hairy. Petals 4. Stamens 6, equal, pod on a stipe Capparidacee, p. Stamens 6, 4 long and 2 short, pod sessile . . Crucifere, p. Petals 3 or 5. Calyx 5-lobed, lobes equal Sazxifragacea, p. Asclepiadacee, p. 104 68 90 51 58 xii ANALYTIC KEY TO THE FAMILIES Calyx of 3 equal or 5 unequal lobes aa. Ovary 2-several-celled. Flowers irregular. Anthers opening by a pore at the top _ Polygalacee, Anthers opening down the side Balsomimaten, p- Flowers regular. Stamens just as many or twice as many as petals. Ovules 1 or 2 in a cell. Herbs. a Flowers unisexual . Euphorbiacee, p. Flowers perfeet. Cells of the ovary the same num- ber as thesepals Geraniacee, p. Cells of the ovary twice as many as the sepals . Linacea, p. Shrubs or trees. Leaves compound or palmately lobed . . . . . Aceracee, p. Leaves simple, alternate; « climb- ing shrub . . CELASTRUS, p. Ovules several or many in a cell. Leaves compound, of three obcordate leaflets . . . . Ovwalidacea, p. Leaves simple. Style 1, stamens free from calyx. Uy Ericacee, p. Styles 2 or 5; leaves opposite from swollen joints Caryophyllacee, p. Stamens not just as many or twice as many as the petals. Trees or shrubs. Stamens fewer than the petals Oleacee, p. Stamens more numerous than the petals. . . . . Aceracee, p. EE. Calyx attached to the ovary for at least part of its length. Tendril-bearing, herbaceous vines Cucurbitacee, p. Not tendril-bearing. 3 Ovules more than 1 to a celk. Ovary 1-celled; sepals2 . . Portulacacee, Cistacee, p. 89 p. 80 84 the 79 82 78 100 40 107 83 132 p. 43 ANALYTIC KEY TO THE FAMILIES xiii Ovary 2—many-celled; stamens on the calyx. Style 1; stamens 4-8 . . . Onagracee, p. Styles 2 or3; stamens5or10 Sazifragacee, p. Ovules only 1 in a cell. Stamens 5 or 10. Trees or shrubs . . . . . CRATEGUS, p. Herbs. Fruit dry, styles2 . . . Umbellifere, p. Fruit berry-like; styles 2-5 Araliacee, p. Stamens 2, 4 or 8. Style 1, stigma 1, fruit a drupe Cornacee, p. Style 1, stigma 2-4-lobed; fruit dry Onagracee, p. BB. Parts of the corolla more or less united. c. Stamens more numerous than lobes of corolla. Ovary 1-celled Placenta 1, parietal . . . . . . . Leguminose, p. Placente 2;parietal . . . . . % . Fumariacee, p. Ovary 2-celled; cells l-ovuled . . . . Polygalacee, p. Ovary 5-many-celled. Stamens free from the corolla. Style 1; leavessimple . . . . . . Ericacee, p. Styles 5; leaves 3-foliate . . . . Oxalidacea, p. Stamens attached to the corolla. Trees or shrubs; style 1 . . « . Ericacee,'p. Herbs; filaments united ina tube . . Malvaceae, p. cc. Stamens not more numerous than the corolla lobes. p. Stamens of the same number as the corolla lobes and opposite to them; style] . . . . Primulacee, p. pp. Stamens alternate with the corolla lobes or fewer. B. Ovary free from the calyx tube. F. Corolla regular. Gc. Stamens as many as the corolla lobes. H. Ovaries more than 1, or if 1, deeply lobed. Ovaries 2, or if 1, then 2-horned. Stamens united . Asclepiadacee, Dp. Stamens distinct . . Apocynacee, p. Ovary 1, deeply 4-lobed. Leaves alternate . Boraginacee, p. Leaves opposite . . . Labiate, p. aH. Ovary 1, not deeply lobed. I. Ovary 1-celled. Seed 1, corolla dry Plantaginacee, p. Seeds several—many. Leaves entire and opposite. Gentianacee, p. 93 58 67 97 97 100 93 68 50 80 100 78 100 89 104 111 110 114 117 127 108 xiv ANALYTIC KEY TO THE FAMILIES Leaves more or less divided, or compound. Corolla white, bearded within, leaves compound of 3 leaflets MENYNANTHES, p. Corolla white or purplish, not bearded within, leaves toothed, lobed, or divided - 1m. Ovary 2-10-celled. Leafless twining parasite cUSCUTA, p. Leaves alternate. Stamens free from the corolla; style Ug nade he PS Ericacee, p. Stamens on the corolla tube. Stamens 4. Leafy stemmed, leaves opposite Verbenacee, p. Stemless, corolla dry and mem- braneous Plantaginacee, p. Stamens 5. Fruit 2 or 4 seed-like nutlets Boraginacee, p. Fruit a pod or berry; few- many-seeded. Style or stigma 3-branched Polemoniacee, p. Style2-branched ; pod4-seeded. » Convoloulacee, p. Style and stigma single, fruit a berry . . . Solanaceae, p. eq. Stamens fewer than the corolla lobes. Stamens, with anthers, 2 or 3. Ovary 4-lobed . . . . . . Lycopus, p. Ovary 2-celled. Stemless herbs ... . Plantaginacee, p. Leafy stemmed herbs with capsule flat- tened and notched at the apex VERONICA, p. Trees... . . . . « Oleacea, p. Fr. Corolla irregular. Stamens, with anthers, either 2 or 4. Ovary 4-lobed, splitting, at maturity, ‘into 4 nutlets. . . . . . . . Labiate, p. Ovary 1-celled. ; Stamens 2; aquatic, often free swimming et ydrophyllacee, p. 109 121 112 100 117 127 114 113 112 120 120 127 125 107 117 Lentibulariacee, p. 125 ANALYTIC KEY TO THE FAMILIES Stamens 4; root parasite without green foliage . . . . . Orobanchacee, p. Ovary 2-celled; seeds numerous . Scrophulariacee, p. ER. Ovary adhering to the calyx tube. ; Tendril-bearing, trailing or climbing herbs Cucurbitacee, p. - Tendrils absent. Stamens separate. Stamens as many as the lobes of the corolla. Stamens free from the corolla Campanulacee, p. Stamens inserted on the corolla -tube. ‘ Leaves opposite or perfoliate Caprifoliacee, p. Leaves opposite with stipules or whorled without stipules . . . Rubiaceae, p. Stamens 1 less than the lobes of the corolla Linnae, p. Stamens united by their anthers. Flowers separate, corolla irregular and split down one side . . . . . Lobeliacee, p. Flowers united in heads on a common recep- tacle and surrounded by an involucre Composite, p. xv 133 134 SELECTED WESTERN FLORA MANITOBA, SASKATCHEWAN, AND ALBERTA DIVISION I. PTERIDOPHYTA. (VASCULAR CRYPTOGRAMS.) _ Plants without true flowers, and reproducing with an alternation of generations. The sexual generation, the prothallus, is incon- spicuous, while the asexual consists of conspicuous fronds, some of which bear the reproductive bodies or spores. I. POLYPODIACEZ (Frrn Famity). Leaf-like herbaceous fronds springing from a creeping rootstock; spores enclosed in sporangia (spore-cases) collected in groups called sori or fruit dots, each of which is covered by a Seale called an indusium ; sori arranged in lines or clusters on the back or margin of the frond division. 1. ASPIDIUM. Woop Fern. Fronds pinnate; fruit dots round, borne on the back of the fertile . fronds which scarcely differ from the sterile; stipes not jointed. 1. A. cristatum, (L.) Sw. Fic. 1.— Prothallus of a Fern. Fronds lanceolate to oblong, the pinne oblong to triangular, deeply cut into 6 to 10 pairs of sharply serrate segments; fruit dots about half way between the margin and the midvein, the covering almost rounded and smooth; stipe and rootstock covered with light brown scales. Moist cold woods, Man. and westward. B 1 2 SELECTED WESTERN FLORA 2. CYSTOPTERIS. Buapper Fern. | Fruit dots on the back of the frond, the covering inflated. Deli- cate ferns with 2-3-pinnate fronds. 1. C. fragilis, (L.) Berna. Fronds lanceolate, 2 or 3-pinnate, the pinnules cut-toothed. Coolies, Alta. 4 3. PTERIS. Bracken. Spore case a continuous line on the back of the frond around the margin, and covered by the edge of the frond folding backwards over it. Fronds 1-3-pinnate. \ Fic. 2, — Fruiting Frond of ‘Fern. ; Sori at a. One sorus with its Fig. 3. — Aspidium cristatum. indusium at b. e eee 1. P. aquilina, L. Brake. Frond stout, 1-2 ft. high, 3-forked at the top, each branch 2-pinnate;- rootstock black, widely spreading. Rocky Mts. and E. Man. II. OPHIOGLOSSACE. More or less fleshy or succulent: plants, consisting of a frond-like stem and leaf growing from a, short rootstock or fleshy.root, . The frond consists of two parts, the one fern-like and sterile, the other a spike or panicle of rather large sporangia, which open at maturity OPHIOGLOSSACE 3. by a transverse slit. The spores are developed from the main tissue of the fruiting stem, and the prothallus is underground. Fie. 4. — Bo- trychium Lunaria. 1. BOTRYCHIUM. Moonworr. Fleshy plants from a short, erect rootstock. The sterile portion ternately or pinnately compound; the fertile portion pinnately divided, with sessile sepa- rate sporangia in rows on the sides of the branches; spores yellow. 1. B. virginianum, (L.) Sw. RATTLESNAKE FERN. Sterile portion sessile, attached above the middle of the plant, the main divisions on short’ stalks, 1 or 2- pinnate, and the leaflets pinnatifid; fertile part 2 or 3- pinnate. Cool rich woods, Man.—Alta. 2. B. Lunaria, (L.) Sw. The sterile part of the frond almost sessile, 1=pinnate, the divisions entire and somewhat fan-shaped, very succulent. Foothills of Rocky Mts. Fie. 5. — Equisetum arvense. et, sterile shoot; f, fertile shoot showing the spike at a; 6, sporophyll, with spo- rangia; $, spore. 4 SELECTED WESTERN FLORA III. EQUISETACEZ (Horserain FamILy). Rush-like plants from a perennial creeping rootstock; stems hollow, jointed, and sheathed at the joints. 1. EQUISETUM. Horseratt. Stems simple or branched, the nodes solid and surrounded by a toothed sheath; fruit borne in a terminal cone-shaped organ formed by a number of shield-like bodies, closely fitted together, and bearing the sporangia on their inner surfaces. J. E. arvénse, L. Common Horseratu. Fertile stems 2-10 in. high, simple or with very few branches, destitute of chlorophyll; sterile stems green and profusely branched, the branches springing in whorls from the nodes. Wet places, common. 2. E. fluviatile, L. Prexs. Stems 3 in.—4 ft. high, sparingly branched, all green and bearing on the top the fruit, which soon falls off, leaving a withered tip. Deep, damp soil, common. DIVISION II. SPERMATOPHYTA. (PHANEROGAMS, OR FLOWERING PLANTS.) Plants bearing flowers with stamens or pistils or hotly and reproducing by seeds. SUBDIVISION I. GYMNOSPERMA. Seeds naked at the base of a scale. ; IV. PINACEZ (Pine Famity). Trees or shrubs with a resinous juice, and mostly entire needle- shaped leaves; flowers borne in a sealy catkin which at maturity becomes a cone, or else berry-like by the scales becoming fleshy; seeds naked, at the base of each scale; mostly evergreen. 1. PINUS. Pine. Cones formed of imbricated woody scales maturing the second year, and spreading when ripe, allowing the seed to fall; leaves never springing singly from the branch, when in 2’s the pair forming a cylinder, when in more than 2, each triangular. PINACEA - 5 1. P. Banksiana, Lamb. Jack Pine. Leaves in 2’s, stout, about 1 in. long, divergent along the branches; cones usually curved, the scales blunt or ending with a minute prickle; a low tree. Sandy soil, E. and N. Man., N. Sask., and Alta. 2. P. strébus, L. Wuuire Prine. Leaves in 5’s, long and slender, the fascicles clustered in large tassels at the ends of the branches; cones 3-6 in. long, cylindrical, nodding, often slightly curved. A lofty tree with white wood, the most valuable timber tree of all the pines, but now scarce. . 8S. E. Man. 3. P. sylvéstris, L. Scorcu Pre. Leaves in 2’s about 2 in. long; fascicles given off all along the branch, but more clustered at the end; cones 13-3 in. long, the scales thick and rigid with a tubercle on the centre; bark gray.