Cambridge Biological Series FLOWERING PLANTS AND FERNS o? ii m m i-q a a a m a CAMBRIDGE BIOLOGICAL SERIES FLOWERING PLANTS AND FERNS CAMBRIDGE UNIVERSITY PRESS C. F. CLAY, MANAGER LONDON : FETTER LANE, E. C. 4 LONDON : H. K. LEWIS, 136, GOWER ST, W.C. i NEW YORK : G. P. PUTNAM'S SONS BOMBAY ) CALCUTTA V MACMILLAN AND CO., LTD. MADRAS j TORONTO : J. M. DENT AND SONS, LTD. TOKYO : MARUZEN-KABUSHIKI-KAISHA ALL RIGHTS RESERVED A DICTIONARY OF THE FLOWERING PLANTS AND FERNS BY J. C. WILLIS M.A., Sc.D., Hon. Sc.D. (Harvard), F.R.S. EUROPEAN CORRESPONDENT, LATE DIRECTOR, BOTANIC GARDENS RIO DE JANEIRO FOURTH EDITION REVISED AND REWRITTEN CAMBRIDGE: AT THE UNIVERSITY PRESS 1919 First Edition, 1897. Second Edition, 1904. Third Edition, 1908. Reprinted, 1914. Fourth Edition, 1919. PREFACE IN this edition the work is completely revised, and as far as possible brought up to date. The most noteworthy new feature is the incorporation of all the parts into one general dictionary, and the omission of Part I of previous editions. When first written this had certain advantages, as being one of the few presentations in English of the ele- mentary facts and theories of ecology. But this advantage has long disappeared, and it seemed to me that the space would be better employed in increasing the number of genera dealt with. On consulting Sir David Prain and other bota- nists, I found that they agreed with this idea. By a slight addition to the total number of pages I have found it pos- sible to include all the genera, and hope that in this way the usefulness of the work to botanists in general may be greatly increased. I have of course attempted no criticism of those included, but have tried to indicate, as far as space would permit, the genera from which they have been segre- gated in many cases, or to which they are united by the other of the two chief recent editors of the vegetable kingdom (Bentham-Hooker, Engler-Prantl). It is obviously impossible to do very much in this direction. As it stands, the book is convenient for use, but a very slight addition to the facts given for each genus would add a line to the entry and, as there are roughly some 20,000 entries, this would add 400 pages to the book, and make it unwieldy. The same remark applies to the geographical distribution, which could not in general be given in great detail. Before criticising, again, the inclusion of many obsolete technical terms and synonyms, it must be remembered that o I vi PREFACE some who use this book wish to use it in connection with floras now long published, and also that it is impossible to reset such a book all through for each edition, so that it is necessary to insert a little "padding" on practically every page. Ecological, morphological, and other subjects have been dealt with by giving comparatively full accounts under se- lected genera. By referring, first to the subject, e.g. to Adventitious Buds, and then to the genera mentioned there, e.g. Begonia, Bryophyllum, &c., a fairly detailed account of each subject may be put together. In using the book, the subjects mentioned under a genus should always be looked up, to find other examples, e.g. under Aesculus look up Bud, Leaf, Leaf-mosaic, Cincinnus, Dichogamy, Change of Colour, >S:c. A book like this must obviously be a compilation, and I have to express my warmest thanks to Prof. A. Engler for permission to draw upon the vast mass of material con- tained in Die Natiirlichen Pflanzenfamilien. Owing to the war I have had no opportunity of asking him to renew this permission, and I trust that he will understand as much. When an article, as is frequently the case with the grouping of the sub-families within the family, is taken from the work mentioned, I have acknowledged the same by giving the name of the author. Otherwise I have in general: drawn upon the book for the genera accepted by its authors, for the number of species (which has been brought roughly up to date by aid of the Supplements to the Index Kewensis), and for their geographical distribution. The list of friends to whom I owe valuable suggestions, useful pieces of assistance, and the like, is very long, and I have no doubt that the following enumeration is incomplete, and must ask the pardon of those who do not figure in it, through some oversight on my part to note down their names at the moment the help was given. In the first place PREFACE vii I wish to thank Sir David Prain and the staff at Kew, more especially Drs Hill, Stapf, and Rolfe, and Mr S. A. Skan, whose detailed knowledge of the library has been of the very greatest assistance in easing my labour. The writing of this edition has occupied very much time during the last five years, and I am particularly grateful to Prof. Seward, who placed at my disposal a table large enough to enable me to spread out 20 works of reference at once, and to Dr Moss and others of the Cambridge staff. The first two of the five years were spent in Rio de Janeiro and I am much indebted for help to my colleagues there, particu- larly the late Dr Alberto Lofgren and Dr Achilles de Faria Lisboa. I also owe many useful suggestions to my col- leagues in other Botanic Gardens, e.g. Prof. O. Ames at Harvard, Prof. I. B. Balfour at Edinburgh, Prof. N. L. Brit- ton at New York, Mr I. H. Burkill at Singapore, Prof. A. Engler at Berlin, Dr J. H. Maiden at Sydney, the late Dr H. H. W. Pearson at Cape Town, Dr Sargant at Jamaica Plain, and others. The late Dr E. A. N. Arber, and Mrs Arber, have laid me under very many obligations, and so have Sir Francis Darwin, Professors Bower, Farmer, Goebel, Goodale, Henry, Lang, Oliver, Scott, and Yapp, and Messrs Davie, Lock, Lynch, Riddle, Small, Smith, and many more. To my wife my obligations are unmeasured. Finally for the illustrations I have to thank Herr Engelrnann for permis- sion to copy some of the late Prof. Eichler's figures, and Dr Rendle for the use of some of those in his book on Classification. J. C. WILLIS. CAMBRIDGE, April 4, 1919. INDEX TO THE IMPORTANT GENERAL ARTICLES, UNDER WHICH LISTS OF EXAMPLES (DESCRIBED AT GREATER LENGTH) WILL BE FOUND. GENERAL Abbreviations, Collecting, Concrescence, Description, Dimorphism, Literature, Nomenclature. VEGETATIVE ORGANS Adnate, Adventitious, Aerenchyma, Aerial Root, Branch, Bud, Bulb, Bulbil, Cauli(fiory), Concrescence, Corm, Iso- (bilateral, &c.), Leaf, Phyllo-(taxy, &c.), Poly-(morphism), Rhizome, Stem, Stipule, Sym- ( podium), Thorn, Tuber, Vegetative Reproduction. REPRODUCTIVE ORGANS Aestivation, Aggregate fruit, Andro-(phore, &c.), Anemo-(philous), Aniso-(phylly), Apetalous, Apo-(gamy), Aril, Asymmetrical, Bee-flowers, Berry, Bract, Butterfly-flowers, Carrion-flowers, Cincinnus, Cleistogamy, Cyme, Dichasial cyme, Dichogamy, Dioecism, Dispersal, Endo- (sperm, &c.), Epi-(gynous, &c.), Floral, Flower, Fly-flowers, Fruit, Geo- (carpic), Gyno-(dioecism, &c.), Heter-(ostylism), Inflorescence, Loose- pollen mechanisms, Mixed inflorescence, Nectary, Nut, Ovary, Perianth, Pollination, Receptacle, Seed, Sex distribution, Stamen, Staminode, Zygo-(morphism). CLASSIFICATION Nomenclature ; and cf. Key to Families at end of book. FORMS OF VEGETATION ; GEOGRAPHICAL DISTRIBUTION Beach-jungle, Chaparral, Climbing Plants, Dispersal, Epiphytes, Floral regions, Halo-(phytes), Insectivorous Plants, Mangroves, Myrme- cophilous Plants, Parasites, Pitcher Plants, Plant formations, Sapro- phytes, Water Plants, Xerophytes, Zones of Vegetation. ECONOMIC BOTANY Alcohol, Alkaloids, Arrowroot, Bamboo, Bark, Camphor, Cinnamon, Condiments, Cotton, Drugs, Dyes, Ebony, Economic Botany, Economic Products, Edible Products, Fibres, Fodder, Foliage Plants, Grass,. Gum, Guttapercha, Lac, Latex, Mucilage, Oil, Ornamental Plants,. Poison, Resins, Rubber, Sugar, Tan, Timber. EXPLANATORY INTRODUCTION The Index of English names, technical terms, &c., which formed Part III of former editions, is now incorporated with the list of genera, so that the work forms one dictionary from end to end, with the exception of the key to the families at the end of the book. All the genera of Bentham- Hooker, Engler-Prantl, and Linnaeus are now included, as well as all given in the Index Kewensis and Supplements (except many synonyms), together with a large number published since the last Supplement, and which, by the kindness of the Director of Ke\v, I have been able to obtain from the MS lists kept at Kew. The most recent of these are given in a little Supplement at the end of the main dictionary, and I hope to bring this supplement up to date at intervals during the currency of the edition, adding the entries later to the body of the work so far as the padding will allow. Besides the genera, all families and higher divisions are also included. The name of the genus is followed by the name of its author, often abbreviated, e.g. R.Br. (cf. Abbreviations), on the system explained under Nomenclature. The original description of the genus may be found by reference to the Index Kewensis. In the same way, the author is given after every species quoted, and the original description may be discovered from the same book. Owing to the continual changes that go on in many families and genera, names are often reduced to synonyms; a great number of such are given in this book, chiefly those used in well-known floras; e.g. Abildgaardia Vahl = Fimbristylis Vahl. Under some of the best known genera, e.g. Abies, a few specific synonyms have also been given, especially names frequently met with in gardens, and opposite to each of these is given the name now generally used; thus Abies alba Michx. must be looked for under Picea, Abies Douglasii Lindl. under Pseudo- tsuga, and so on. It is very difficult to decide when divergence of two forms is sufficient to entitle them to rank as genera, and this difficulty is the cause of much synonymy. A genus A is established by one author, and then it is discovered not to differ sufficiently from another genus/?, established by the same or another author, to remain as an independent genus. A is therefore merged in B and becomes a synonym. The x I NTH OD UCTION species of A retain as far as possible their old specific Qames when placed in B. When an entry such as " Abildgaardia Vahl = Fimbri- stylis Vahl p.p." is found, it means that the genus Abildgaardia as established by Vahl is merged in his Fimbristylis. Many of the species change their names, but some retain their specific names, when the name is not already occupied. This latter case is often indicated by putting the name of the old genus in brackets after that of the new, thus, F. (A.) fulvescetts. In many cases the names of some of the genera thus merged in other genera are indicated thus: Axinandra Thw. (BH. inch Naxiandra Krasser) ; no attempt however has been made to give all such cases or a fraction of them, but only a few of the more im- portant. In particular those have been given where the genus as here defined differs from the definition in Engler and Prantl's Natiirliche Pfianzenfamilien or Bentham and Hooker's Genera Plantarum by the inclusion and exclusion of other genera. The name of the genus is followed by that of the family to which it belongs, and after this is often a number (in brackets) indicating the section of the family, thus Acacia belongs to Subfamily I and Tribe 2 of Leguminosae. The general plan upon which the book has been con- structed, and the necessity for condensation, render it essential, if the full advantage is to be derived from its use, that the student should refer to the family as well as the genus. There he will find the important general characters possessed by its members, and should examine the genus to see in what it agrees, and in what it disagrees, with these. A further reference to the classification given at the end of the article upon the family will point out the special characters to be looked for in the genus as a member of some particular sub-family or tribe. In this way a large amount of information about the particular plant in question may be obtained, and at the same time the student will get into the way of regarding plants not as so many independent and disconnected units, but as related members of one great whole. In this way too he will soon acquire an appreciation of the relative importance of the different characters in classification and will learn to recognise the approximate relationships of most plants after a brief inspection, or even at sight. The families are those given by Engler in his Syllabus and in Die natiirlichen Pflanzenfamilien, but sufficient reference is made to Bentham and Hooker's system of classification to enable any one who may prefer to use that system to do so. The name of the family is followed by a statement of the number of INTR OD UCT1ON xi species in the genus, and its geographical distribution. The number, unless very small, is always only an approximation ; new research is always bringing new species to light, splitting up older ones, or com- bining two or more into one. This is all the information that is given about a very large number of the genera ; only when a genus presents some character of interest which is not common ta the order or group, is any particular mention made of it. The biological peculiarities of the most important genera are dealt with pretty fully, but much has been omitted. Thus in dealing with the pollination-methods of flowers a selection of important genera, illustrating the various methods, has been made for description ; so too with epiphytes, xerophytes, the morphology of parts, and so on. General discussions of all these subjects will be found under the title of the subject itself, and numerous examples are there quoted; these examples are mostly dealt with more fully. Numerous cross-references to other articles, e.g. Buds, Dichogamy, Fruit, Leaf, Parasite, Xerophyte, &c. (cf. Index), are made, and should be looked up. While in the morphology, &c. a selection has thus been made of genera for treatment, this is less the case with economic botany. This has been more fully treated, only a comparatively few genera being omitted. Space, however, has not permitted of a detailed description of economic products or the way in which they are obtained ; for this reference must be made to other works (see Literature). Turning now to the other articles upon the families, the same general principles apply to them. After the name of the family is given the order to which it belongs, marked EP. or BH. if necessary to distinguish between these systematists. This should be looked up in the key at the end ; this will show the families which are most nearly related to the one under consideration, and the characters that distinguish one from the other can be made out by comparison of their descriptions. The student should always endeavour to make out why a given family is classified in the position assigned to it. When the family as defined by Engler differs from that defined by Bentham and Hooker, as is so often the case, an attempt should be made to discover the reasons for the difference. After the position of the family in the system follows the number of its genera and species, the morphology and natural history of its vegeta- tive and reproductive organs, its economic products, and finally, in the case of the more important families, its classification into sub-families and tribes, with the more important genera belonging to each. The student should work through this part and study as many of the genera xii INTRODUCTION as possible before leaving the family. This is easily managed in dealing with the outdoor collection in our botanic gardens. No particular attempt is made in the book to avoid technical terms. When a term or abbreviation is used that the reader does not understand he should look it up in the Dictionary, or under Abbreviations. To save space, in many cases in which there are several words in use beginning with the same prefix, e.g. aniso-, apo-, endo-, epi-, geo-, gyno-, halo-, heter-, iso-, phyllo-, poly-, sym-, xero-, &c., all are given under the heading of the prefix. If a word is looked up as a whole, there- fore, and not found, reference should be made to the prefix before saying that it is omitted from the Dictionary. Many genera are described by different authors under different spellings, and the most common and important of such cases are in- cluded ; thus Prunella is also described under Brunella, Eleocharis also under Heleocharis, £c. \ A (fl. -class), fls. with freely exposed honey ; Acer, Euphorbia, Galitim, Hedera, Jlex, RhaniiiHs, Saxifrages, Umbelliferae. A-, An- (Gr. prefix), not. Aaron's Beard, Hypericum calycinuni L. AB (fl. -class), fls. with partially concealed honey; Caltha, Crataegus, Cruciferae, Fragaria, Poteniilla, Ranunculus, Scdtnit. Ab- (Lat. prefix), from; -axial (side), away from axis; -breviated, shortened; -errant, differing from type; -normal, varying from the rule; -original, strictly native; -ortion, imperfect or arrested development; -rupt, terminating suddenly; -sciss-layer, separation- layer for dropping the 1.; -sorption, taking up of fluids by r. or 1. Abaca, Manila hemp, Musa textilis Nee. Abasoloa La Llave. Compositae (5). i Mex. Abatia Ruiz et Pav. Flacourtiaceae (7) (Samydaceae, BH.}. 5 trop. S. Am. Fl. apet. L. opp. Abauria Becc. (Koompassia Maingay, EP.}. Legum. (n. 5). 2 Malaya. Abbevillea Berg. = Campomanesia Ruiz et Pav. p.p. (Myrt ). Abbottia F. Muell. Rubiaceae (n. 2). i N. Austr. Abbreviations. Descriptions of floral morphology are largely given in the terms of Floral Formulae, explained under that heading. When the name of a genus or family is repeated in the article dealing with it, it is represented by the initial letter only, e.g. A. for Abies. The name of a family is sometimes abbreviated by the omission of the terminal aceae, &c. ; e.g. Capparid., Compos. The term 'warm' is sometimes used instead of 'tropical and subtropical.' The expression BH. after a genus or family, &c., means "as defined by Bentham and Hooker in their Genera Plantaruni " ; EP. means "as defined by Engler and Prantl in the Pflanzenfamilien and Pflanzenreich" The following mathematical and other symbols are largely used : 2 , hermaphrodite © or O, annual tf , male © or Q, biennial ? , female 2+ , perennial ( ) enclosing P, K, C, A, or G, h,, tree or shrub united or concrescent > , more than G superior, 'G inferior, ovary < ,.less than oo, indefinite, numerous ±, more or less than x , hybrid _L , at right angles to §, section (of sp. or genus) ||, parallel to W. r A BBRE VIA TIONS = , equal to, merged in !, seen by author fi, micromillimetre, T(rVT mm. -ffi-, actinomorphic •|- , zygomorphic The following abbreviations are largely employed in this and other botanical books : N. hemisphere S. hemisphere , Old World New World A(ndroeceum) Abyss(inia) Achlam(ydeous) Actinom(orphic) Acum(inate) Adv(entitious) Afr(ica) Aggr(egate) Agr(icultural) Alb(umen) Alt(ernate) Am (erica) Amphitr(opous) Anatr(opous) Anemoph(ilous) Ann(ual) (als, &c. ) Ant(arcti)c Apet(alous) Apoc(ar)p(ous) Arch(ipelago) Avchichl (amydeae) Arct(ic) Arg(entina) Art(icle) As(ia) Assim(ilation) Asymm(etrical) Atl(antic) Austr(alia) Axill(ary) B(eatus), the late Beitr(age) Ber(ichte) Bot(any) Br(act) Braz(il) Brit(ain) Bull(etin) C(entral) (orolla) Cal(yx) (edonia) California) Campylotr(opous) Cap(itate) Caps(ule) c.c., cubic centimetre Cel(eberrimus) Centr(al) (alblatt) Char(acter) Chi(na) Cl(arissimus) (ements*) Cleist(ogamic) cm., centimetre Col(ony) Collected by) (ection) Concr(escence) Consp(icuous) Conv(olute) Cor(olla) Cosmop(olitan) Cot(yledon) C(om)p(oun)d C(ar)p(e)l Cult(ivated) Dehisc(ent) Dep(artment) Descr(iption) Dich(asial) Dichlam(ydeous) Dichot(omous) Dicot(yledon) Dim(inutive) Dioec(ious) Diplost(emonous) Distr(ibution) Dorsiv(entral) Ed(ible) (ition) Endosp(erm) England) Entomoph(ilous) * Research Methods in Ecology. Epig(ynous) Epipet(alous) Epiph(yte) Esp(ecially) Ess(ential) Eur(ope) Evap (oration) Evergr(een) Exalb(uminous) Exc(ept) Excl(uding) Exstip(ulate) Extr(orse) Extrafl(oral) Fam(ily) Fert(ilisation) Fl(ower)(in)g Fl(ore) pl(eno), double-flowered Fol(ium, a leaf) (iage) Fr(uit) Fri(gid) G(ynoeceum) Gen(us) Germin(ation) Ges(ellschaft) Gland(ular) Gr(eek) H(erbarium) Hab(itat) Hem(isphere) Herb(arium) Heterochlam(ydeous) Heterost(yled) Himal(aya) Hind(ustani) Homochlam(ydeous) Horiz(ontal) Hort(orum), of gardens ABB RE VIA TIONS Hybr(id) Hypog(ynous) I(sland) Ic(on), figure Imbr(icate) Inc(ertae) sed(is), of unknown position Incl(uding) Inconspic(uous) Ind(ia) Indeh(iscent) Indomal(aya) Ined(itus), unpublished Inferior) Infl(orescence) Interpet(iolar) Intr(orse) Invol(ucre) Irreg(ular) Isobil(ateral) Jap(an) Jard(in) K, calyx L(eaf) Lat(in) (eral) Laticif(erous) L(oco) c(itato), in the place quoted Linn(ean) Loc(ulus) Loculic(idal) m(etre) Madag(ascar) Mag(azine) Mal(aya) Masc(arenes) Mech(anism) Medit(erranean) Membr(anous) Met(amorphosed) Mex(ico) mm., millimetre Moluc(cas) Monoch(asial) Monochlam(ydeous) Monocot(yledon) Monoec(ious) Mus(eum) n(ovus), new N(atural) O(rder) Nat(ural) (uralised) Nat(iirlichen) Pfl(anz- enfamilien) Nearct(ic) Neotrop(ical) Nom(en), a name Nud(us), naked, without description N(ew) Z(ealand) Obdipl(ostemonous) Off(icinal) Opp(osite) Orn(amental) Orthotr(opous) Ov(ule) (ary) P(erianth) Pac(ific) Palaearct(ic) Palaeotrop(ical) Paras(ite) Pecl(icel) (uncle) Pen(insula) Pend(ulous) Perenn(ial) Perf(ume) Perig(ynous) Pet(al) Pfl(anzen) R(eich) Phil(ippines) Pl(ant) Plac(enta) Poll(icaris), inch Pollin(ation) Polyg(amous) Polyn(esia) Post(erior) p(ro) p(arte), in part Pref(ix) Prodr(omus) Protandr(ous) Protog(ynous) P(oin)t R(oot) Rad(ix) (ical) Recept(acle) Reg(ular) Repr(oduction) Repres(ented) Rev(iew) Rhiz(ome) Rudim(entary) S(eu), or Sandw(ich Is.) Sci(ence) Sem(en), a seed Sep(al) Septic(idal) Septifr(agal) Ser(ies) Sicc(us), dry Soc(iety) Sol(ilary) Sp(ecies) Sta(men) St(amino)d(e) Stip(ule) Subm(erged) Subtrop(ical) Succul(ent) Suff(ix) Sup(erior) Sympet(alous) Syn(onym) Sync(arpous) T(abula), a figure T(omus), a volume Tab(ula), a figure Tasm(ania) Temp(erate) Term(inal) Trans(actions) Transv(erse) Trop(ical) Undershr(ub) Usu(ally) Var(iety) Varieg(ated) Veg(etation) V(idi) S(iccam), dry specimen seen V(idi) V(ivam), living specimen seen W(est) I(ndies) Wiss(enschaft) Xero(phyte) Zygom(orphic) Of course many of these abbreviations also signify the adjectival and I 2 ABBREVIATIONS other forms of the word, e.g. albumen, albuminous, &c.; character, characterised, characteristic, &c. The following abbreviations of authors' names are in common use in giving the authority for genera or species : Achar(ius) Adans(on) Afz(elius) Ag(ardh) Ait(on) Alef(eld) Allem(ao) All(ioni) Anders(on) Andr(ews) Ant(oine) Arch(er) Ard(uino) Aresch(oug) Arn(ott) Aschers(on) Aubl(et) Auct(orum) Bab(ington) Bail(ey) Baill(on) Bak(er) Bal(ansa) Balb(is) Balf(our) Barb(osa) Rodr(igues) Barnad(es) Barn(eoud) Barr(elier) Bartl(ing) Batem(an) Bauh(in)* Baumg(arten) Beauv(ois) Becc(ari) Bedd(ome) Benj(amin) Benn(ett) Benth(am) B(entham and) H(ooker)t Berg(ius) Bernh(ardi) Bert(ero) Berth(elo) Berthol(ini) Bertol(oni) Bess(er) Bieb(erstein) Bigel(ow) Binn(endijk) Bisch(off) Bl(ume) Boeck(eler) Boerh(ave) Boiss(ier) Boj(er) Bomm(er) Bong(ard) Bonpl(and) Borck(hausen) Br(aun, own) Bref(eld) Brongn(iart) Brot(ero) Brunf(els) Buch(anan)- Ham(ilton) Burch(ell) Bur(eau) Burm(ann) Buxb(aum) Camb(essedes) Carr(iere) Carr(uthers) Casp(ary) Cass(ini) Cast(agne) Cav(anilles) C.DC., Casimir de Candolle Cerv(antes) Cham(isso) Champ(ion) Chapm(an) Chav(annes) Chois(y) Cl(ements) Clus(ius) Cogn(iaux) Colebr(ooke) Col(enso) Colm(eiro) Comm(elin) Comm(erson) Corn(uti) Coss(on) Cram(er) Cunn(ingham) Curt(is) Dalz(ell) Dav(enport) DC., A. P. de Candolle (17/8- 1841) Dec(ais)ne Ueless(ert) Del(ile) Dennst(aedt) De Not(aris) Desf(ontaines) Desr(ousseaux) Desv(aux) Dicks(on) Didrichs(en) Dietr(ich) Dill(enius) Dillw(yn) Dodon(aeus) Dougl(as) Drumm(ond) Dryand(er) Duch(artre) Dumort(ier) Dun(al) Eat(on) Eckl(on) Edgew(orth) Ehrenb(erg) Ehrh(art) Eichl(er) Ell(iott) Endl(icher)J Engelm(ann) Engl(er)§ Eschsch(oltz) Eschw(eiler) Ettingsh(ausen) Fabr(icius) Falc(oner) Fing(erhuth) Fisch(er) Flac(ourt) Forsk(al) Forst(er) Fourn(ier) Fourr(eau) Franch(et) Frem(ont) Fres(enius) Fr(ies) Gaertn(er) Gal(eotti) Gardn(er) Gasp(ari) Gaudich(aud) Gaud(in) Gies(enhagen) Gilb(ert) Gilib(ert) Gill(ies) Gis(eke) Gled(itsch) Gmel(in) Godr(on) Goldm(ann) G rah (am) Gren(ier) Grev(ille) Griffith) Griseb(ach) t Genera Plantarum, 1862—83. * Prodromus Theatri Botanici, 1620. J Ibid. 1836-40. § Natiirlicken PJlanzenfamilien, 1889-97; Das Pflanzenreich, i9oo-(in progress). ABB RE VIA TIONS Gronov(ius) Guett(ard) Guill(emin) Guss(one) Hack (el) Hall(er) Ham(ilton) Hanst(ein) Hartm(ann) Hartw(eg) Harv(ey) Hassk(arl) Havv(orth) Hedw(ig) Hegelm(aier) Heist(er) Hemsl(ey) Henfr(ey) Herb(ert) Herm(ann) Hern(andez) Hieron(ymus) Hildebr(and) Hill(ebrand) Hochst(etter) Hoffm(ann) Hoffm(an)s(eg)g Honck(eney) H(oo)k(er) Hook(er) Hook. f(ilius) * Horan(inow) Hort(orum), of gardens Houst(on) Houtt(uyn) Huds(on)- Humb(oldt) H(umboldt), B(onpland) & K(unth) Isn(ard) Jacks(on) Jacq(uin) Jenrn(an) Jord(an) Jungh(uhn) Juss(ieu) 1748-1836 Kaempf(er) Karst(en) Kaulf(uss) Kell(ogg) Kl(otzsch) Knaut(ius) Koel(er) Koen(ig) Koern(icke) Kon(ig) Korth(als) Kostel(etzky) K(un)tze, O. K(un)ze L(innaeus)f 1707-78 Labill(ardiere) Lag(asca) Lam(arck) Lamb(ert) Langsd(orff) Lapeyr(ouse) Laxm(ann) Leandr(o) Leavenw(orth) Ledeb(our) Lehm(ann) Lej(eune) Lem(aire) Lepr(ieur) Lesch(enault) Less(ing) Lestib(oudois) L. f(ilius) L'Herit(ier) Licht(enstein) Liebm(ann) Lindl(ey) 1799-1865 Lindm(ann) Linn(aeus)t Loefl(ing) Loes(ener) Loud (on) Lour(eiro) Ludw(ig) Luerss(en) Macfad(yen) Maing(ay) Mak(ino) Marcgr(af) Markh(am) Mart(ius) Mast(ers) Maxim(owicz) Medic(us) Meissn(er) Mert(ens) Mett(enius) Mey(er) Mich(au)x Mich(eli) Mig(uel) Mihi, of me Mik(an) Mill(er) Miq(uel) Mirb(el) Mitch(ell) Moc(ino) Moehr(ing) Mol(ina) Monn(ier) Moq(uin-Tandon) Morr(en) Muehlenb(erg) Muell-Arg(au) Muell(er) Muell, F(erd. v.) Murr(ay) Naud(in) Neck(er) Newm(an) Nied(en)z(u) Nor(onha) Nutt(all) Nyland(er) Nym(an) Oerst(ed) Oliv(er) Ort(ega) Parl(atore) Parm(entier) Pasq(uale) Pav(on) Perr(ottet) Pers(oon) Peterm(ann) Peyr(itsch) Pfeiff(er) Pfitz(er) Phil(ippi) Planch(on) Pluk(enet) Plum(ier) Poepp(ig) Poir(et) Poll(ich) Ponted (era) Pr(esl) Putterl(ick) Racib(orski) Radlk(ofer) Raf(inesque) R(obert) Br(own) Red(oute) Reich(ardt) R(ei)ch(en)b(ach) Reinw(ardt) Reiss(eck) Retz(ius) Reut(er) Rich(ard) Ridl(ey) Riv(inus)J Roem(er) Rohrb(ach) Roth(ert) Rottb(oell) Roxb(urgh) Rudb(eck) Rumph(ius) Rupp(ius) Rupr(echt) Sadeb(eck) Sald(anha) Salisb(ury) Sanguin(etti) Sauv(ageau) * SeeB. & H. t The starting point of modern nomenclature is the publication in 1753 of liis Species Plantarutn. \ Bachmann, 1652 — 1723. ABB RE VIA TIONS Soland(er) Torr(ey) Wallr(oth) Solms-Laub(ach) Tourn (efort) Walp(ers) Sond(er) 1656-1708 Walt(ers) Sonn(erat) Tratt(inick) Warb(urg) Spegazz(ini) Trautv(etter) Warm (ing) Splitg(erber) Trec(ul) Wats(on) Spr(engel) Trev(isano) Webb(er) Steinh(eil) Trin(ius) Wedd(ell) Steph(ens) Tul(asne) Welw(itsch) Sternb(erg) Turcz(aninew) Wendl(and) Steud(el) Turp(in) Wettst(ein) St Hil(aire) Tuss(ac) Wigg(ers) Sw(artz) Und(erwood) Wikstr(om) Tabern(aemon- Urb(an) Willcl(enovv) tanus) 1520-90 Vaill(ant) Willem(et) Targ(ioni) Vand(elli) Willk(omm) Toz(zetti) Vell(ozo) Wimm(er) Taub(ert) Vent(enat) Wulf(enius) Teysm(ann) Vieill(ard) Wydl(er) Thorns (on) Vill(ars) Zahlbr(uckner) Thonn(ing) Vis(iani) Zenk(er) Thou(ars) Viv(iani) Zeyh(er) Thunb(erg) Vog(el) Zipp(el) Thw(aites) Wahlenb(erg) Zoll(inger) Tod(aro) Wall(ich) Zucc(arini) Sauv(alle) Schau(er) Scheff(er) Scheidw(eiler) Schlecht(en)d(al) Schmid(el) Schnizl(ein) Schomb(urgk) Schrad(er) Schreb(er) Schult(es) Schum(acher) Schum(ann) Scop(oli) Scortech(ini) Seem(ann) Sendtn(er) Ser(inge) Seub(ert) Shuttl(eworth) Sibth(orpe) Sieb(old) Siegesb(eck) Smirn(ow) Sm(ith) Sod(iro) Abdominea J. J. Smith. Orchidaceae (n. 20). i Java. Abdra Greene (Draba L. p.p.). Cruciferae (4). i N. Am. Abele tree, Popnlus alba L. AbeliaR.Br. (Linnaea Gronov. p.p. EP.). Caprifoliaceae (3). 15 As., Mex. Sta. 4, didynamous. Abelmoscbus Medic. (Hibiscus L. p.p.). Malv. 12 trop. , Austr. Aberemoa Aubl. (Ditguetia St. Hil.). Anonaceae (i). 30 trop. Am. Aberia Hochst. (Dotyalis E. Mey. p.p. EP.}. Flacourtiaceae (4). (Bixineae, BH.). 12 Afr., Ceylon. A. caffra Harv. et Sond. (Ivei apple), and others, eel. fr. Abies (Tourn.) L. Synonymy: A. alba Michx. = Picea a.; do. Mill.= A. pectinata DC.; A. americana Mill. = Tsuga canadensis; A. californica Hort. = Pseudotsuga Douglasii; A. canadensis Michx. = Tsuga c.; do. Mill. = Picea alba; A. Cedrus Poir. =Cedrus Libani; A. Deodara Lindl. =C. D.; A. Douglasii Lindl. = Pseudotsuga D.; A. excelsh Link = A. pectinata; do. Poir. = Picea e. ; A. Kaempferi Lindl. = Pseudolarix K. ; A. Larix Poir. = Larix europaea; A. iiiontana Nym. = Picea excelsa; A. miicronata Rafin. = Pseudotsuga Douglasii; A.nigra Desf. or Duroi = Picea n. ; A. Omorika Nym. = Picea O.; A. orientalis Poir. = Picea o. ; A. pectinata Poir. = Picea rubra; A. Picea Lindl. = A. pectinata DC.; do. Mill. = Picea excelsa; A. rubra Poir. = Picea r. ; A. vulgaris Poir. = A. pectinata. Coniferae (Pinaceae, 2; see C. for generic characters). 24 N. temp. The firs are evergreen trees with needle 1. borne directly on the stems. ACACIA 7 No short shoots. On the main stem the symmetry is radial, whilst on the horizontal branches the 1. twist so as to get their surfaces all much in one plane. If the top bud or leader be destroyed, however, a branch bud below it takes up the vertical growth and radial symmetry. Cones large, arranged much like Finns; ? often brightly coloured, though wind-fertilised. The carpel-scales are large and appear on the outside of the cone between the ovuliferous scales. The cone ripens in one year. A. pectinata DC. (silver-fir, Mts. ofS. Eur.) yields a valuable wood, 'Strasburg' turpentine, &c. A. balsamea Mill. (E. N. Am., balsam fir) yields the turpentine known as Canada balsam. Many others yield useful timbers and resins. Handsome trees; commonly cult, are A. concolor Lindl. et Gord. (N. W. Am.), A.firma Sieb. et Zucc. (Japan), A. nobilis Lindl. (N. W. Am.), A. Nordmanniana Spach (Caucasus), A. Pinsapo Boiss. (Spain), A. Webbiana Lindl. (Himal.). Abietineae, a tribe ofConiferae Pinaceae. Abildgaardia Vahl. = Fimbristylis Vahl p.p. (Cyper.). Abiogenesis, spontaneous generation. Abobra Naud. Cucurbitaceae (3). t temp. S. Am. Abola Lindl Orchidaceae (n. 19). i Colombia. Abolboda Humb. et Bonpl. Xyridaceae. 10 S. Am. Abortion, imperfect or arrested development. Abroma Jacq. Sterculiaceae. 10 trop. As. to Austr. A. augusta L. (Indomal.) bark yields a good fibre. Abroma Juss. Nyctaginaceae. 30 N. Am. Anthocarp winged. Abrophyllum Hook. f. Saxifragaceae (v). i E. Austr. Abrotanella Cass. Compositae (7). 15 Rodriguez, Tierra del Fuego, Austr., N.Z., Auckland Is. Abrotanum (Tourn.) L.= Artemisia Tourn. p.p. (Compos.). Abrus L. Leguminosae (in. 9). 6 trop. A . precatorius L. has hard red seeds with black tips (crab's eyes), strung into necklaces, rosaries, &c., and used as weights (rati) in India (cf. Adenanthera). See Keiv Bull. 1890, p. i (Weather Plant). The roots are used in India as Indian liquorice. Absinthe, Artemisia. Absinthium Tourn. ex L. = Artemisia Tourn. p.p. (Compos.). Absolmsia O. Ktze. (Astrostemma p.p.Bff.). Asclepi.(ii. i). i Borneo. Abundance |CI.), the total number of individuals in an area. Abuta (Barr.) Aubl. Menispermaceae. totrop. S. Am. A.rufescens Aubl. (Guiana) yields white Pareira root. Abutilon Tourn. Malvaceae (2). 100 trop. and sub-trop. Noepicalyx. Fl. mech. like Malva silvestris, but some are self-sterile; the sta. do not move down, and the styles emerge through the anther-mass. Many visited by humming-birds. A. Avicennae Gaertn. cult, in China for fibre China jute. Abyssinian banana, Musa Ensete}. F. Gmel.; primrose, Primula. Acacallis Lindl. (Aganisia Lindl. EP.). Orchid, (n. 13). i N. Brazil. Acacia (Tourn.) L. Leguminosae (r. i). 500 trop. and sub-trop., mostly trees (wattles); typical leaf-form bipinnate with oo leaflets and small scaly slips. About 300 sp., forming the § Phyllodineae (chiefly in Austr., where they are char., and Polynes.), have simple leaf- like phyllodes, i.e. petioles flattened so as to place their surfaces 8 ACACIA vertically — this exposes less surface to radiation. A mere inspection, though it shows the phyllode to he a leaf-structure (it has an axillary bud), does not show that it is not a 1. turned edgewise, though it shows no twist at the base. Occasionally however there are reversions to type (i.e. to the ancestral form) on the plant, some phyllodes occurring with leaf-blades of the ordinary bipinnate type. This is still better seen in germinating seedlings. The first 1. are typical bipinnate 1., followed by others with slightly flattened stalks and less blade, and so on, until finally only phyllodes are produced. In A. alata R.Br. and others, the phyllodes are decurrent on the stem, like the 1. of thistles. In many the slips, are repres. by large thorns, swollen at the base. In A. spkaerocephala Cham, et Schlecht. (Cent. Am.), the thorns are inhabited by colonies of ants, which bore into them and clear out the internal tissue. The ants live on the A. and are fed by it. Extrafl. nectaries occur on the petioles, and yellow sausage-shaped food-bodies on the tips of the leaflets. These consist of parenchymatous cells containing food -st lifts, and are eaten by the ants. If attempt be made to interfere with the tree the ants rush out. (Cf. Cecropia, and see Nature, Aug. 1893, for an account of the leaf- cutting ants: and cf. Schimper, Plant Geogr. p. 140. This was the first case of myrmecophily (symbiosis with ants) discovered; see Belt's Naturalist in Nicaragua.) Other sp. are myrmecophilous also. A few are twiners, others hook-climbers. Most are xerophytes, often forming char, features in vegetation and scenery, e.g. the babul (A. arabica Willd.) with its low, spreading habit, is almost the only tree in many parts of the dry plains of India, and others are common in S. Afr. &c. In Austr. (esp. S. Austr.) the A. take a great part in the formation of the scrub, a concourse of shrubby plants of many genera, which covers the almost waterless country with a waste of veg. about 6 — 10 feet high, of a general bluish green effect, and with few herbs or grasses beneath. The fl. (diagram, see order) has oo long sta. , affording little pro- tection to the pollen. In A. hotnalophylla A. Cunn. (S. E. Austr.; Myall) the seed hangs out on a long red funicle. Many valuable products. A. Senegal Willd. (Soudan) yields the best gum-arabic; the gum exudes from the branches principally during the prevalence of the dry desert winds. Other sp. yield inferior qualities. A. catechu Willd. (E. Ind.) yields catechu orcutch (used in tanning), by digestion of the wood in hot water. With this the true khaki cloth is dyed and shrunk. A. decnrrens Willd. (Austr. ; black wattle) yields good tan bark; inferior barks from A. pycnantha Benth. (S. E. Austr.; golden wattle), A. dealbata Link (Austr.; silver wattle) &c. That of A. arabica Willd. is largely used in India. The wood of many is valuable, esp. Australian black-wood, A. mclanoxylon R.Br. Many have sweetly scented fls. ; those of A. Farmsiam Willd. (trop.) are the Cassie flowers of perfumery. A. annata R.Br. (temp. Austr.; kangaroo thorn), A. horrida Willd. (S. Afr.) &C. form good hedges or sandbinders. Acacia, false, Robinia Pseud-acacia L. Acaena L. Rosaceae (in. 9). 80 =fc, Mex., Calif., Polynes. Fr. hooked. Some sp. bud from junction of leaf and stalk. ACANTHACEAE 9 Acajou (W. I.), Guarea trichilioides L. Acaju, Cashew, Anacardium occidental L. Acalypha L. Euphorbiaceae (A. II. 2). 300 trop. , S. Afr. Anther lobes twisted ; stigmas branched. Several cult, for varieg. 1. Acampe Lindl. Orchidaceae (n. 16). 12 Indomal., China, (Afr. BH.). Acamptocladus Nash (Eragrostis Host, p^p.). Gramin. (n). i S.W. U.S. Acamptopappus A. Gray (AplopappusCa.ss.J3H.). Comp. (3). 2 N. Am. Acanthaceae (EP., BH.). Dicots. (Sympet. Tubiflorae). 140 gen., 2000 sp., esp. trop. but also Medit., U.S., Austr. 4 chief centres of distr. : Indomalaya, char, by Strobilanthes and Andrographidae, Afr. (Thunbergia and Barleria), Braz. (Mendoncia and Ruellia), and Centr. Am. (Aphelandreae and Odontonema). Many biological types — climbing plants, xerophytes, marsh plants, &c. — and much variety in habit. Very many in. damp places in trop. forests. Trees are rare ; most are shrubs or herbs with opp., usu. decussate and entire, exstip. 1., usu. thin. Cystoliths, visible as streaks or protuberances, are usu. common on 1. and stems. Infl. most commonly a dich. cyme, in its ultimate branchings tending to monoch., and frequently condensed in the leaf-axils as in Labiatae. Racemose infls. also occur, and sol. fl. are common. Bracts and bracteoles usu. present, o'ften coloured; the latter frequently large, ± enclosing the fl. Fl. $, hypog., zygom., usu. with nectariferous disc below ov. K (5 — 4), C (5—4), commonly two-lipped (upper lip sometimes not developed, e.g. in Acanthus). A rarely 5, usu. 4 or 2 epipet., usu. exserted ; i — 3 stds. frequently present ; anthers often with one lobe smaller than the other, or abortive; connective often long (rf. Salvia). The pollen exhibits great variety of patterns (see Nat. Pfl.); these are generally constant in the genus, and may be used in classification. G (2), 2-loc. with axile plac. each with 2 — oo usu. anatr. ov. in two rows. Style usu. long with two stigmas, the post, often smaller. The general arrangement of the fl. for visits of insects, protection of pollen, &c. is like Labiatae or Scrophulariaceae. Fr. a bi-loc. caps, (with few exceptions), usu..± stalked, loculic. to the very base. Seeds usu. exalb. Their modes of distribution are interesting (see Nat. Pji.). The capsules of § iv explode and the seeds are thrown out, largely by the aid of peculiar hook-like out- growths from their stalks (retinacula or jacnlators). Many have superficial scales and hairs which on wetting become mucilaginous (cf. Linum, Collomia), e.g. Crossandra, Ruellia, Blepharis. Classification and chief genera (after Lindau) : I. NELSONIOIDEAE (Ovules oo ; jaculators papilla- shaped) : Ebermaiera, Nelsonia. II. MENDONCIOIDEAE (Ov. 4, seeds not more than 2. Drupe; no jac.): Mendoncia. III. THUVBERGIOIDEAE (Ov. 4. Capsule; jac. papilla- like): Thunbergia. IV. ACANTHI01DEAE (Ov. 2— co . Capsule; jac. hook- shaped) : A. Contortae (cor. conv., or never ascendingly imbr.): Strobilanthes, Ruellia, Eranthemum, Barleria. io ACANTHACEAE B. Imbricatae (cor. ascendingly imbr., or with no upper lip) : Blepharis, Acanthus, Crossandra, Aphelandra, Andrographis, Dicliptera, Fittonia, Odontonema, Justicia, Beloperone. Acantbella Hook. f. Melastomaceae (i). i Orinoco valley. Acantbo- (Gr. prefix), thorny; -carpous. spiny-fruited. Acantbobotrya Eckl. et Zeyh. — Lebeckia Thunb. p.p. (Legum.). Acantbocardamum Thell. (Lepidium p.p. BH.}. Crucif. (2). i Persia. Acanthocarpus Lehm. Liliaceae (in). 3 S.W. Austr. (June. BH.} Acantbocepbalus Kar. et Kir. Compositae (13). 2 W. As. Acantbocereus Britton et Rose (Cereus p.p.)- Cact. (in. i). i S. Am. Acantbochiton Torr. Amarantaceae (2). i Texas, Arizona. Acantbocladus Kl. (Polygala L. p.p. EP.}. Polygalaceae. 3 S. Am. Acantbococos Barb. Rodr. (Cocos L. p.p.). Palmae (iv. 2). i Paraguay. Acantbodium Uelile = Blepharis Juss. (Acanth.). Acantbolepis Less. Compositae (11). r W. As. Acantbolimon Boiss. Plumbaginaceae. 80 E. Meclit., desert pi. Acanthomintba A. Gray. Labiatae (vi). 2 Calif. Acantbonema Hook. f. Gesneriaceae (i). i W. Afr. Acantbonycbia Rohrb. (Pentacaena Bartl.). Caryophyll. (i. 4). 5 Pac. Am. Acantbopale C. B. Clarke. Acanth. (iv. A). 7 trop. Afr. (Malaya?). Acantbopanax Miq. Araliaceae. 25 Ind. , Chi., Japan. Acantbopbippium Blume. Orchidaceae (n. 9). 7 Indomal. The axial outgrowth from the base of the column, common in O., is here very great and bends first downwards, then up, removing the insertion of the lateral sepals and labellum to a distance from the column. Acantbopboenix H. Wendl. Palmae (iv. t). 3 Mascarenes. Acantbopbyllum C. A. Mey. Caryophyllaceae (n. 2). 25 W. As., Siberia. Mostly desert xerophytes with prickly leaves. Acantbopsis Harv. Acanthaceae (iv. B). 78. Afr. Acantborbiza H. Wendl. Palmae (i. 2). 4 trop. Am. The adv. roots from the lowest nodes grow normally downwards, but those from the nodes above develope into thorny branches. Acantboscypbus Small (Oxytheca Nutt. p.p.). Polygon, (i. i). i N. Am. Acantbosicyos Welw. Cucurbitaceae (3). i S.W. Afr., A. horrida Welw., the Narras, a remarkable plant growing on sand dunes (cf. Welwitschia). The thick root is very long (up to 40 ft.). Above > ground is a thorny shrub, with long tendrils; the thorns are modified twigs. • (See Welwitsch, Trans. Linn. Soc. 27, 1869.) Acantbospermum Schrank. Compositae (5). 5 trop. Am., Galapagos. Acantbospbaera Warb. Moraceae (il). i Amazon valley. Acantbostacbys Link, Klotzsch, et Otto (Ananas Tourn. p.p.). Bromeliaceae (4). i (A. strobilacea L. K. et O.) Brazil. Acantbosyris Griseb. Santalaceae. 3 temp. S. Am. Furniture wood. Acanthotbamnus T. S. Brandegee. Celastraceae. i Mex. Acantbotreculia Engl. Moraceae (n). i Cameroons. Acantbura Lindau. Acanthaceae (iv. A), i Mattogrosso. Acantbus Tourn. ex L. Acanthaceae (iv. B). 25 trop. and sub-trop., As., Afr., Eur., mostly xero. with thorny 1. (those of A. spinosus L. furnished, it is supposed, the pattern for the decoration of the capitals of Corinthian columns). A. ilicifolhis L. is part of the palaeotrop. ACER ii mangrove (g.v.) veg. Fl. a large bee-fl.; there is no upper lip to the C, and the protection of the pollen, &c. is undertaken by the K. The anthers form a box by fitting closely together at the sides, and shed their pollen sideways into it, where it is held by hairs till an insect probing for honey forces the filaments of the sta. apart and receives a shower of pollen on its head (loose-pollen mechanism, cf. many Scrophulariaceae, Ericaceae, &c.). In the young flr. the style is behind the anthers, later on it bends down so as to touch a visiting insect. The fr. explodes ; large 'jaculators' on the seeds. Acarna All. = Atractylis L. p.p. (Compos.). Acarodomatia, cavities mite inhabited, Anamirta, Fraxinus, Parameria. Acaulescent, almost stemless; acaulis (Lat. ), without visible stem. Accessory branch, bud, supernumerary in same axil, see Buds ; organs of flower, the perianth. Acclimatisation, adaptation to new climate. Accrescent, enlarged and persistent. Accumbent, see Cruciferae. Aceituna. Symplocos. Acentra Phil. (Hybanthits Jacq. p.p. EP.) Violaceae. r Chili. Acer (Tourn.) L. Aceraceae (Sapind. BH. ). 1 1 5 N. temp., esp. in hill districts (A. psendoflafanits L., sycamore, and A. campestris L. , maple, in Brit., the latter native) and trop. nits.; many in China and Japan. Trees and shrubs, with opp. exstip. 1., deciduous or ever- green. L. often simple entire, more commonly 3- or 5-lobed, occa- sionally cpd. One may go through a collection of A. in an herbarium or elsewhere, comparing the 1. as to degree of development of the drip-tips (acum. apices to easily wetted 1., from which the water drips off rapidly after a shower, cf. Ficus) noting the kind of climate from which each has come. There is a correlation between length of tip and wetness of climate. Large winter buds, covered by scale 1. In many sp. transitional forms may be seen as the bud elongates in spring, between the scales and the green 1., showing that the scale = not the whole 1, but the leaf base. In the § Negundo there are no scales, but the bud is protected by the base of the petiole of the 1. in whose axil it arises. The 1. commonly exhibit varnish-like smears, of sticky consistence, known as honey-dew, the excretion of aphides which live on the 1.; the insect bores into the tissues, sucks their juices, and ejects a drop of honey-dew on an average once in half-an-hour. In passing under a tree infested with aphides one may sometimes feel the drops falling like a fine rain (see Pithecolobium). The fluid is rich in sugar. When the dew falls the hygroscopic honey-dew takes it up and spreads over the 1. ; then later in the day evap. reduces it to a varnish on the surface. Many other trees exhibit this phenomenon, e.g. lime, beech, oak (Btisgen, Der Honigthau, Jena). Fls. in racemes, sometimes contracted to corymbs or umbels, reg., polyg., not conspic. ; formula usu. KS, C5, A4 + 4, G (2). Apetaly in some. 3 cpls. are frequent, esp. in the end fl. of a raceme. 5 fls. protandrous ; honey freely exposed on the disc (fl. -class A), available to insects of all kinds. Fr. a samara. In germination, the long green cotyledons come above the soil almost at once. 12 ACER A. saccharum Marshall (A. saccharimim Wangenh.) and others of the E. U.S. yield maple sugar (i — 4 Ih. a tree) obtained by boring holes in February and March and collecting and evaporating the juice. Many yield good timber and charcoal. A number of Japanese sp., with prettily shaped or varieg. 1., cult, as orn. shrubs. Aceraceae (Sapind. p.p. BH.}. Dicots. (Archichl. Sapindales). i gen. (Acer, Dipteronia) with 120 sp. N. temp, and trop. mts. Trees and shrubs; 1. opp., petiolate, exstip., simple entire or more often palmately or pinnately lobed or cpd. Infl. racemose, corymbose, or fasciculate. Fls. reg., andromonoec., androdioec., dioec., &c., 5-4-merous, usu. dichlam. Disc annular or lobed or reduced to teeth, rarely absent. A 4 — ro, usu. 8, hypog., perig., or on disc; $ fir. with rudimentary G. G (2), 2-loc., lat. compressed ; styles 2, free or joined below; ov. 2 in each loc., orthotr. to anatr., with dorsal raphe. Fr. of 2 samaras, separating when ripe. Seeds usu. solitary, exalb., the cotyledons irreg. folded. Many yi^ld good timber, sugar, &c. (cf. Acer). Largely represented in the Tertiary. Aceranthus Morr. et Decne. (Epimedium Tourn. EP.}. Berberidaceae. 3 Chi., Jap. Aceras R.Br. Orchidaceae (n. i). i Eur. (incl. Brit.), Medit, A. anthropophora R.Br., the man-orchis. Like Orchis. Acerates Ell. = Gomphocarpus R. Br. p.p. (Asclepiad.). Aceriphyllum Engl. Saxifragaceae (i). i N. China. Acerose, needle-shaped. Acetabuliform, like a shallow saucer. Acetosa Tourn. ex Mill. = Rumex Linn. (Polygon.). Acetosella Moehr. =Oxalis Linn. (Oxalid.). -aceus (Lat. suffix), like. Achaenipodium T. S. Brandegee. Compositae (=j). i Mex. Achaetogeron A. Gray. Compositae (3). 10 Mex., Calif. Achantia A. Chevalier. Sterculiaceae. i trop. Afr. Acharia Thunb. Achariaceae. i S. Afr. Achariaceae (Passifloraceae, p.p. BH.}. Dicot. (Archichl. Parietales). 3 gen., 3 sp. S. Afr. Herbs or undershrubs with reg. monoec. fl. k;s— 5, C (3—5), A 3—5 epipet., G (3—5) with oo ov. on parietal plac. Caps. Endosp. Acharitea Benth. Verbenaceae (3). 2 Madag. Achatocarpus Triana. Phytolaccaceae (Amarant. BH.}. 10 trop. Am. Achene, a one-seeded small dry indehiscent fruit (q.v.) of i carpel. Achetaria Cham, et Schlecht. = Beyrichia Cham. (Scrophular.). AchiUea L. Compositae (7). 115 N. temp. A. Millefoliiun L. (yarrow or milfoil) and A. Ptarmica L. (sneezewort) in Brit. AcMlus Hemsl. Zingiberaceae (i). i Siam. No stds. Unisex, fl. Achimenes P.Br. Gesneraceae (n). 25 trop. Am., often cult. Achlaena Griseb. Gramineae (6). i Cuba. Achlamydeous, without a perianth (q.v.}. Achlamydosporeae (BH.}. The sixth series of Monochlamydeae. Achlys DC. Berberidaceae. 2 Japan and Pacif. N. Am. The perianth aborts early in development. Achneria Munro. Gramineae (9). 8 S. and trop. Afr. Ach.nopn.ora F. Muell. Compositae (3). i Austr. ACMOPYLE 13 Achradotypus Baill. Sapotaceae (i). 4 New Cal. Achras L. Sapotaceae (i). i W. Ind., trop. Am., A. Sapota L., cult. for ed. fr. (Sapodilla plum). The coagulated resinous latex (chicle gum) is used in U.S. for chewing-gum, statuettes, &c. AcliratiJiis O. Ktze. = Arachnites Phil. (Burmann.). Achroantries Raf. (Microstylis'N.vM. p.p. BH.). Orchid, (n. 4). i N. Am. Achudemia Blume. Urticaceae (2). 2 Java, Japan (?). Achyrachaena Schau. Compositae (5). i N.W. U.S. ^ Pappus of broad, silvery scales; fruit-heads used as 'everlastings.' Achyrantries L. Amarantaceae (2). 15 trop. and sub-trop. AcLyrocline Less. Compositae (4). 25 Madag., trop. Afr. and Am. Achyronia Royen ex L. = Aspalathus L. (Legumin.). Achyronychia Torr. et A. Gray. Caryophyll. (i. 4) (Illecebr. BH.}. 3 S.W. U.S., Mex. Achyropappus H. B. et K. — Schkuhria Roth. p.p. (Compos.). Achyrophorus Adans. = Hypochaeris Linn. p.p. (Compos.). Achyropsis Benth. et Hook. f. (AchyrantkesL. p.p. EP.). Amarant. (2). 3 Afr- Aciiyrospermum Blume. Labiatae (vi). 12 trop. Afr. to Malay Is. Achyrosteprms Kze. Compositae (inc. sed. ). Nomen. Achyrothalamus O. Hoffm. Compositae (12). 2 E. trop. Afr. Aciachne Benth. Gramineae (8). i trop. S. Am. Acianthus R.Br. Orchidaceae (n. 2). 12 Austr., N.Z., New Caled. Acicalyptus A. Gray. Myrtaceae (i). 3 Fiji (New Cal.?). Acicarpha Juss. Calyceraceae. 5 S. Am. Acicular, needle-shaped. Acidanthera Hochst. Iridaceae (ill). 16 trop. and S. Afr. Cult. Acidocroton Griseb. Euphorbiaceae (A. II. 3). i Cuba. Acidoton Sw. Euphorbiaceae (A. n. 2). 3 W. I. Acies, the edge. Acinaciform, scimitar-shaped. Acineta Lindl. Orchidaceae (n. 13). 10 Cent. Am., Mexico. An axial outgrowth carries out 2 sep. and the labellum (attached to the column). Acinodendron O. Ktze. (-drum L. ,#//.) = Miconia Ruiz et Pav. (Melastom.). Acinos Rupp., Moench. = Calamintha Lam. (Labiat.). Acioa Aubl. (Conepia BH.}. Ros. (vi. b). i N.E. S. Am. with ed. oily seed, 20 trop. Afr. Aciotis D. Don. Melastomaceae (i). 30 trop. Am., W. Ind. Aciphylla Forst. Umbelliferae (in. 5). 25 Austr., N.Z. Acis Salisb. = Leucojum L. (Amaryllid.). Acisanthera P.Br. Melastomaceae (i). 20 trop. Am., W.Ind. Ackama A. Cunn. Cunoniaceae (Saxifrag. BH.}. 2 N. Z., E. Austr., New Guinea. Acleisantlies A. Gray. Nyctaginaceae. 8 Mex., Texas. Aclisia E. Mey. =Pollia Thunb. p.p. (Commelin.). Acmadenia Bartl. et Wendl. f. Rutaceae (i). 15 S. Afr. Acmantriera Griseb. Malpighiaceae (n). 2 S. Am. Acmella Rich. = Spilanthes Jacq. (Compos.). Aemena DC. = Eugenia L. p.p. (£ff.) = Syzygium Gaertn. p.p. Acmopyle Pilger {Dacrydium Soland. p.p.). Taxaceae. i New Cal. 14 ACNIDA Acnida L. Amarantaceae (2). 3 U.S. Dioecious. Acnlstus Schott. Solanaceae (2)'. 20 trop. Am. Acoelorrhaphe H. Wendl. Palmaceae (i. 2). Nomen. Acokanthera G. Don. Apocynaceae (i. i). 3 Abyss., S. Afr., A. venenata G. Don, root and wood supplies Zulu arrow-poison ' Acom (W.I.), Dioscorea bulbifera L. Acomastylis Greene (Potentilla et Geum p.p.). Rosac. (in. 2). s N. Am. Acomis F. Muell. Compositae (4). 3 Austr. Acomosperma K. Schum. Asclepiadaceae(nomen). i Amazon valley. Aconceveibum Miq. Euphorbiaceae (B. n). i Java. Aconite, Aconitin, Aconitum ; winter-, Eranthis hyemalis Salisb. Aconitum Tourn. ex L. Ranunculaceae (2). 80 N. temp. A. Napcllns L. (aconite, monkshood, wolf's bane) in Brit. Fls. in racemes (see order). The post, sepal forms a large hood, enclosing the two 'petals' which are repres. by nectaries on long stalks. Fl. protandrous, adapted, by its structure and its blue colour, to bees. The distribution of A. largely coincides with that of the humble-bee (Bombus). Humble-bees often rob the flr. of its honey by biting through the hood. Fr. of follicles which open so far as to expose the seeds, which only escape when shaken by wind or otherwise (censer- mechanism}. All are poisonous; the tuberous roots contain alkaloids of the aconitin group (used in medicine). A. ferox Wall (root) furnishes the Bikh poison of Nepal. Acontias Schott = Xanthosoma Schott, p.p. (Arac.). Acophorum Gaudich. Gramineae. Nomen nudum. Acorellus Palla ex Kneuck (Cyperus L. p.p.). Cyper. (i). 3 S. Eur., Medit. Acoridium Nees et Meyen (Ceratoslylis Blume, Dendrochilum Blume). Orchidaceae (n. 3). 45 Indomal. Acorn, Qtiercus. Acorus L. Araceae (i). 2 N. temp., S.E. As. A. Calamus L. (sweet flag) Brit. Rhiz. sympodial; 1. isobil. Fl. $ , protog., with P. Used in flavouring. Acotyledones (Jussieu) = Cryptogamae. Acourtia D. Don=Perezia Lag. (Compositae). Acquired characters, non-hereditary, arising during life. Acradenia Kipp. Rutaceae (i). i Tasm. Acrandra Berg. (Campomanesia Ruiz et Pav. BH.}. Myrtaceae (i. i). 2 S. trop. Braz. Acranthera Arn. Rubiaceae (i. 7). 9 Indomal. Acranthous, cf. Orchidaceae. Acridocarpus Guill. et Perr. Malpighiaceae (i). 20 Afr., Madag., Arabia. Acriopsis Reinw. Orchidaceae (n. 1 6). 10 E. Indomal. Acrista O. F. Cook. Palmae (iv. i). i Porto Rico. Acritochaete Pilger. Gramineae (5). i Kilimanjaro. Acriulus Ridl. Cyperaceae (n). 2 Madag., Angola. Aero- (Gr. pref.), apical; -carpous, with terminal fruit; -gamae = Porogamae; see Chalazogamae ; -gens, ferns and mosses; -nychius (Lat.), curved like a claw; -petal, produced successively towards ACTINOLEMA 15 the apex; -phytium (Cl.), an alpine-plant-formation ; -scopic, facing the apex ; -spire, plumule; -tonic, cf. Orchidaceae. Acrobotrys K. Schum. et Krause. Rubiaceae(i. 3). i Colombia. Acrocarpidium Miq. = Peperomia Ruiz et Pav. (Piper.). Acrocarpus Wight ex Am. Leguminosae (n. 7). 3 Indomal. Acrocephalus Benth. Labiatae (vn). 40 Malay Arch, to trop. Afr. Acrochaene Lindl. Orchidaceae (n. 16). I Sikkim. Acroclinium A. Gray=Helipterum DC. p.p. (Comp.). Acrocoelium Baill. Icacinaceae (Olacineae, Bff.). i Congo. Acrocomia Mart. Palmae (iv. 2). 10 trop. Am., W. Ind. Acrodiclidium Nees. Lauraceae (n). 20 trop. Am. , W. I. A. Puchury Mez furnishes the ed. puchurim nuts. Acroglochin Schrad. Chenopodiaceae (A). i N. India, China. The fruit mass is prickly, many of the twigs not ending in fls. Acrolasia Presl. (Menfzelia L.). Loasaceae. 25 N. Am. Acrolopnia Pfitz. (Eulophia R. Br. p.p.). Orchidaceae (n. 5). 9 S. Afr. Acronychia Forst. Rutaceae (iv). 20 trop. As., Austr. Acropera Lindl. = Gongora Ruiz et Pav. p.p. (Orchid.). Acrophorus Presl. Polypodiaceae. i Indomal. Acrophyllum Benth. Cunoniaceae (Saxifrag. BH.). i New S. Wales. Acropogon Schlechter. Sterculiaceae. 3 New Cal. Acrosanthes Eckl. et Zeyh. Aizoaceae (n). 5 S. Afr. Acrosepalum Pierre (Ancistrocarpus Oliv. EP.). Tiliaceae. i trop. Afr. Acrospira Welw. (Dcbesia O. Ktze. EP.). Liliaceae (in). 3 Angola. Acrostemon Klotzsch (Eretitia D. Don, p.p.). Ericaceae (i v. 2). 9 S. Afr. Acrostichum L. Polypodiaceae. 4 trop. A. aureum L. is common on trop. coasts. Acrostylia Frappier. Orchidaceae (n. i). i Reunion. Acrotome Benth. Labiatae (vi. i). 58. and trop. Afr. Acrotrema jack. Dilleniaceae. 10 Indomal. Acrotriche R. Br. Epacridaceae (3). 8 temp. Austr. Acrymia Prain. Labiatae (i). i Malay Penin. Actad (Cl.), plant of a rocky sea-shore. Actaea (Tourn.) L. (Z>//.excl. Cirnicifuga L.). Ranunculaceae (2). 10 N. temp. A. spicata L. (bane-berry or herb-christopher), in Brit. Fls. in racemes (cf. Aconitum). Cpl. i. Berry. Actephila Blume. Euphorbiaceae (A. I. i). 10 Indomal., Austr. ActinantHus Ehrenb. (Oenanthe Tourn. p.p. EP.). Umbellif. (ill. 5). i W. As. Actinea Juss. = Cephalophora Cav. BH. — Actinella Pers. Actinella Nutt. Compositae (6). 20 Am. (dwarf sunflower). Actinella Pers. = Actinella Nutt. EP. Actinidia Lindl. Dilleniaceae (Ternstroem. BH.). 10 E. As. Actiniopteris Link. Polypodiaceae. i trop. Afr., As. It has the habit of a small palm with fan leaves. Actinocarya Benth. Boraginaceae (iv. i). i Tibet. Actinochloa Willd. = Bouteloua Lag. (Gramin. ). Actinodaphne Nees. Lauraceae (i). 50 E. Ind., Japan, N. Am. Actinodium Schau. Myitaceae (n. 3). i W. Austr. Actinokentia Dammer. Palmae (iv. i). i New Cal. . Bff.). Umbellif. (n. i). zE.Medit. 1 6 ACTINOLEPIS Actinolepis DC. (Eriophyllum Lag. p.p.). Compos. (6). 9 West U.S. Actinomeris Nutt. Compositae (5). 3 Atl. U.S. Actinomorphic (fl.), radially symmetrical, or symmetrical about any plane passing through the centre (includes cases like most Sympetalae, where there are only two carpels and more of other whorls). Actinophloeus Becc. (Drymophloeus Zipp. p.p.). Palmae (iv. i). i New Guinea. Actinopnora Wall. (Schoutenia Korth. BH.). Tiliaceae. 3 trop. As. Actinopnyllum Ruiz et Pav. =Sciadophyllum P. Br. BH. = Schefflera Forst. (Aral.). Actinorhytis Wendl. et Drude. Palmae (iv. i)- i Malaya. Actinoschoenus Benth. Cyperaceae (n). 3 China, Ceylon, Madag. Actinostemma Griff. Cucurbitaceae (i). 6 Ind. to Japan. Actinostemon Mart, ex Klotzsch. Euphorbiaceae (A. n. 7). 30 trop. Am. Actinostrobus Miq. Coniferae (Pinaceae 4; see C. for generic characters). 2 S.W. Austr. Actinotus Labill. Umbelliferae (i. i). 15 Austr. (flannel flower^. Actium (Cl.), rocky sea-shore formation. Actoplanes K. Schum. Marantaceae. 2 Malaya. Aculeate, Aculeatus (Latin), prickly. Acuminate, tapering to a point in hollow curves. Acura Hill. Compositae. Nomen. Acute, tapering to a sharp point in straight lines. Acutifolius (Lat.), with acute leaf. Acyclic, not in whorls. Ad- (Lat. pref.), to; -axial (side), the side towards the axis; -hesion, concrescence (q-v. ) of dissimilar organs, e.g. C and A; -justment (Cl.), functional response to stimuli ; -nascens (Lat.), growing upon something ; -pressed, appressed ; -verse, facing the main axis. -ad (Cl.), suffix denoting an ecad. Ada Lindl. Orchidaceae (n. 19). 2 Colombia. Cult. Adactylus Rolfe (Apostasia Blume p.p.). Orchidaceae (i. i ). 3 trop. As. Adamia Wall. = Dichroa Lour. (Saxifrag. ). Adam's needle, Yucca. Adansonia L. Bombacaceae (i). 10 palaeotrop. A. digitata L. is the baobab. Its height is not great, but the trunk may reach 30 feet in thickness. Fr. woody. (See Card. Chr. 1900, 57.) Adaphus Neck. Inc. sed. ( = Laurus Tourn. ?). Adaptable (CL), able to originate ecads. Adaptation, adjustment to conditions of life. Adder's tongue, Ophioglossmn. Addisonia Rusby. Compositae (2). i Bolivia. Adelia L. Euphorbiaceae (A. n. 2). 15 W.I., Colombia. Adelia P. Br. = Forestiera Poir (Olea.). Adeliopsis Benth. Menispermaceae. i N.E. Austr. Adelmeria Ridl. (Elmeria Ridl.). Zingiberaceae. 2 Phil. Is. Adelobotrys DC. Melastomaceae (i). 15 trop. Am., W.I. Adelodypsis Becc. (Dypsis Nor. p.p.). Palmae (iv. i). 2 Madag. Adelonema Schott \Homalonema p.p. BH.). Araceae (v). i Amazon, Adelonenga Becc. (Nenga p.p.). Palmae (iv. i). 2 New Guinea. ADENOSTEGIA 17 Adelopetaluui Fitzger. Orchidaceae (u. 3). i New S. Wales. Adelosa Blume. Verbenaceae (4). i Madag. Adelostemma Hook. f. Asclepiadaceae (n. i). i Burma. Adelostigma Steetz. Compositae (3). 2 trop. Afr. Aden- (Gr. pref.), a gland; -old, gland-like; -ophore, stalk supporting a gland. Adenandra Willd. Rutaceae (i). 25 S. Afr. Cult. orn. fl. Adenanthera Royen ex L. Leguminosae (i. 4). 5 trop. As., Austr. Seeds hard and bright red, or red and black (cf. Abrus). Adenanthos Labill. Proteaceae (i). 20 W. and S. Austr. Adenaria H. B. et K. Lythraceae. i Mex. to Arg. Adeneleutnera O. Ktze. = Ad_eneleutherophora Barb. Rodr. Adeneleutherophora Barb. Rodr. Orchidaceae (n. 6). i Brazil. Adenia Forsk. (MoJecca L. BH.}. Passiflor. 50 palaeotrop. Adenimesa Nieuwland = Conophora Nieuwland (Compositae). Adenium Roem. et Schult. Apocynaceae (n. i). 10 Arabia, Afr. Xerophytes with thick stems, and rather fleshy 1. Adenocalymna Mart. Bignoniaceae (i). 55 trop. Am. Adenocarpus DC. Leguminosae (in. 3). 10 Medit., trop. Afr. Adenocaulou Hook. Compositae (4). 3 Ind., Japan, U.S., Chili. Adenochilus Hook. f. Orchidaceae (n. 2). 2 Austr., N.Z. Adenochlaena Boiss. ex Baill. Euphorb. (A. n. 4). 2 Madag., Indomal. Adenocllne Turcz. Euphorbiaceae (A. n. 2). 5 S. Afr. Adenoderris J. Sm. Polypodiaceae. 2 W.I. Adenodolichos Harms. Leguminosae (in. 10). 12 trop. Afr. Adenogonum Welw. ex Hiern. (Engleria O. Hoffm. EP.}. Compositae (3). 2S. Afr. Adenogramma Reichb. Phytolaccaceae (Ficoideae BH.). 7 S. Afr. Adenogynum Reichb. f. et Zoll. (Chloradenia Baill. BH.). Euphorb. (A. II. 2). i Java, Timor. Adenolinum Reichb. =Linum L. (Lin.). Adenolisianthus Gilg. Gentianaceae (i). 2 Brazil. Adenoncos Blume = Sarcochilus R. Br. (Orchid.). Adenoon Ualz. Compositae (i). i Indomal. Adenopappus Benth. Compositae (6). i Mex. Adenopeltis Bert. Euphorbiaceae (A. n. 7). i Chili. Adenopetalum Klotzsch et Garcke= Euphorbia L. p.p. (Euphorb.). Adenophaedra Muell.-Arg. Euphorbiaceae (A. n. 2). i E. Brazil. Adenophora Fisch. Campanulaceae (i). 25 temp. Eur., As. Adenophyllum Pers. (Dysodia Cav. p.p. EP.). Compositae (6). 3 Mex. Adenoplea Radlk. (Buddleia Houst. p.p. EP.). Loganiaceae. 2 Madag. Adenoplusia Radlk. Loganiaceae. i Madag. Adenoporces Small (Tetrapteris Cav. p.p.). Malpighiaceae (i). i San Domingo. Adenopus Benth. Cucurbitaceae (3). 8 trop. Afr. Adenoropium Pohl.=Jatropha L. p.p. (Euphorb.). Adenosacme Wall. (Mycetia Reinw. EP.). Rubi. (i. 7). 7 Indomal. Adenosma Nees = Cardanthera Buch.-Ham. (BH.) — Synnema Benth. (Acanth.). Adenosma R. Br. Scrophular. (n. 6). 10 Indomal. , Austr. , China. Adenostegia Benth. = Cordylanthus Nutt. (Scrophular.). 12 N. Am. W. 2 1 8 ADENOSTEMMA Adenostemma Forst. Compositae (2). 6 trop. Am., i cosmotrop. Pappus glandular and sticky; fr. carried by animals. Adenostoma Blume. Scrophulariaceae. Nomen. Adenostoma Hook, et Am. Rosaceae (in. 3). i Calif. A. fascicu- latum H. et A. is one of the shrubs forming the chaparral or chamisal. Adenostyles Cass. Compositae (2). 6 alpine, Eur., As. Minor. Adenostylis Blume = Zeuxine Lindl. (Orchid.). Adesmia DC. (Patagoniitm Schrank). Leguminosae (in. 7). 90 S. Am. Leafstalks thorny, plants often with glandular hairs. Adhatoda Tourn. ex Medic. (Justicia L. p.p. EP.) Acanth. (iv. B). 100 trop. Adhunia Veil. Inc. sed. i Brazil. Adiantopsis Fee. Polypodiaceae. 15 trop. Am. As. Adiantum L. Polypodiaceae. 190 cosmop., esp. trop. Am. (maiden hair); A. Capilhis-veneris L. in Brit. (rare). Hothouse favourites. Some are climbing epiphytes. Adicea Raf. - Pilea Lindl. (Urtic.). Adina Salisb. Rubiaceae (i. 6). 15 trop. As., Afr. Adinandra Jack. Theaceae. 15 warm As., Afr. Adinobotrys Dunn. Leguminosae (in. 6). 5 Indomal., China. Adlumia Rafin. Papaveraceae (in), i E. N. Am. A leaf-climber. Adnaria Raf. = Gaylussacia H. B. et K. (Eric.). Adnate (adnation), concrescence of organs of different nature, e.g. axillary shoot to main shoot or leaf, stamens to petals, &c., Anthu- rinm, Apocynaceae, Araceae, Asclepiadaceae, Asclepias, Boraginaceae, Chailletia, Compositae, Citphea, Cyperaceae, Erythrochiton,Juglans, Passiflora, Pontederiaceae, Samolns, Solanaceae (figure), Sfathicarpa, Spat hiphy Hum, Tilia, Zostera ; anther, one joined to the filament by its whole length. Adolia Lam. (Scutia Comm. BH. £P.) Rhamnaceae. 3 trop., and S. Afr. Adolpnia Meissn. Rhamnaceae. 2 Mex., Calif. Adonis Dill, ex L. Ranunculaceae (3). 10 N. palaeotemp. A. aiitum- nalis L. , pheasant's eye, Brit. Adoxa L. Adoxaceae. i N. temp., incl. Brit., A. Moschatellina L., (moschatel). Rhiz. creeping, monopodial, bearing a flg. shoot with a few rad. 1., a pair of opp. cauline 1., and a small head of greenish fls., usu. 5 (a condensed dich. cyme). The term. fl. is usu. 4-merous, the lat. 5-merous (cf. Ruta, &c.). Fl. ? , reg., greenish, inconspic. P of 2 whorls; the outer usu. 3-merous, sometimes regarded as an invol. formed of bract and bracteoles, but quite probably a K. Sta. alt. with petals, divided almost to the base. G (3 — 5), rarely (2), semi- inf. with one pend. ov. in each loc. Drupe with several stones. Endosp. Chief visitors small flies, attracted by the musky smell. Adoxaceae (EP. ; Caprifol. p.p. BH.). Dicots. (Sympet. Rubiales). Only genus Adoxa (q.v.). Sometimes united to Saxifragaceae ; no very close relationships (see Schumann, Alorph. Stitdien). Adrastaea DC. (Hibbertia Andr. p.p. EP.). Dilleniaceae. i E. Austr. Adriana Gaudich. Euphorbiaceae (A. II. 4). 5 Austr. Adromlschus Lem. (Cotyledon Tourn. p.p. BH.). Crassul. 10 S. Afr. Adrorhizon Hook. f. Orchidaceae (11. 16). i Ceylon. AERIAL ROOTS 19 Adruc (W. Ind.), Cyperus articulatns L. Aduncate, bent like a hook. Adventina Raf. Compositae (inc. seel.). 2 N. Am. Adventitious (Cl.), invading from distant formations ; buds, arising elsewhere than normally in an axil, Begonia, Bryophylluin, Carda- mine, Cyslopteris, Linaria, Ophioglossum, Pteris, Pyrola; embryo, one formed without fertilisation, AUhornea, Citrus, Euonymns, Funkia, Nothoscorditin ; root, one developed from stem or leaf, Acan- thorhha, Araceae, Bromeliaceae, Gramineae, Orchidaceae, Palmae and most Monocotyledons, Peperomia, Podosteniaceae, Ranunculaceae, &c., and cf. Aerial roots ; shoot, one arising from root or leaf, Ailanthus, Anthuriitin, Podosteniaceae, Rafflesiaceae, Testudinaria. Adventive (Cl.), established temporarily. Adverse, facing main axis. Adynamandry, self-sterility. Adyseton Adans. =Alyssum L. (Crucifer.). Aechmandra Arn. = Melothria L. p.p. (Cucurbit.). Aechmantnera Nees. Acanthaceae (iv. A.). 2 Himalaya. Aechmea Ruiz et Pav. Bromeliaceae (4). 50 epiph., W.I., S. Am. Aechmolepis Decne. Asclepiadaceae (i). i Angola. Aedesia O. Hoffm. Compositae (i). 2 W. trop. Afr. Aegialitis R. Br. Plumbaginaceae. i trop. Austr. and As. Aegiceras Gaertn. Myrsinaceae (n. i). 2 palaeotrop. A. tnajus Gaertn. grows in mangrove swamps together with Rhizophora, &c., and exhibits a similar habit, vivipary, &c. Aegilops L. = Triticum L. p.p. (Gramin.). Aeginetia L. Orobanchaceae. 2 Ceylon to Japan and Phil. Is. Aegipnila Jacq. Verbenaceae (4). 40 trop. Am., W.I. Aegle Correa. Rutaceae (v). 3 Indo-mal. A. Marmelos Correa is the bael fruit, a valuable remedy for dysentery, &c. Aeglopsis Swingle. Rutaceae (v). i Ivory Coast. Aegopodium Knaut. ex L. Umbelliferae (ill. 5). 2 Eur., As. A.Poda- graria L. the goat, gout, or bishop's weed, in Brit. Aegopogon Beauv. Gramineae (3). 2 Braz. to Calif. Aeluropus Trin. Gramineae (10). 5 Medit. to Ind. Halophytes. Aeneus (Lat.), brass-coloured. Aeolanthus Mart. Labiatae (vn). 25 Afr. Aeonia (Oeonia) Lindl. Orchidaceae (n. 20). 5 Masc. Aeonium Webb et Berth. = Sempervivtim L. (Crassul.). Aequalis (Lat.), similar in size; Aequi- (Lat. pref.), equal. Aerangis Reichb. f. (Angraecutn Thou. p.p. BH.). Orchid, (n. 20). i Angola. Aeranthes Lindl. Orchidaceae (11. 20). 4 Madag., Masc. Aeranthus Rchb. f. = Mystacidium Lindl. (.#//.) = Macroplectrum Pfitz. Aerating roots, roots with aerenchyma. Aerenchyma, respiratory tissue formed by the phellogen; Avicennia, Bmgitiera, Herniiniera, Jitssieua, Neptitnia, Knmex, Sesbania, Sonneratia, Taxodiuin, Terminalia. Aeria O. F. Cook. Palmae (iv. i). i Porto Rico. Aerial roots, adventitious roots arising above ground, often forming bitttresses (Palmae, Pandanaceae), pillars (Araceae, Ficus), clasping 20 AERIAL ROOTS and climbing organs (Araceae, Hedera, Orchidaceae, Tecoma), water- absorbing organs (Orchidaceae, Velloziaceae), assimilating organs (Orchidaceae, Podostemaceae), thorns (Acanthorhiza), parasitic suckers (Cuscuta, Viscum), &c. Aerides Lour. Orchidaceae (11. 20). 20 E. As. Leaves fleshy. Aerophytes, epiphytes. Aerotropism, influence of gases on growth and curvature. Aeruginous, verdigris-coloured. Aerva Forsk. Amarantaceae (2). 12 trop. As., Afr. Aesandra Pierre (Payena A. DC. p.p. EP.). Sapotaceae (i). i trop. As. Aeschynanthus Jack. (Trichosporum D. Don.) Gesneraceae (i). 75 Indomal., China. Many epiphytes with fleshy leaves. Extreme protandry with movement of sta. Seeds with long hairs. Aeschynomene L. (incl. Herminiera Guill. et Perr. EP.). Legum. (in. 7). 70 warm. From the pith-like wood of A. aspera L. (shola, pith-plant) the solar helmets of trop. As. are made. Aesculus L. (incl. Pavia Boerh.). Hippocastanaceae (Sapind. BH.). 20 N. temp., S. Am. A. Hippocastanum L. (horse-chestnut) and several of § Pavia orn. trees. A. ohioensis Michx. (glabra Willd.) buckeye (U.S.). Trees with large winter buds, covered with resinous scale 1., containing next year's shoot and infl. very ad- vanced. The bud expands rapidly in spring. In A . pawiflora Walt, transitions from scale to perfect 1. may be seen, showing the former to = leaf bases. L. opp., exstip., palmate; the blades when young are hairy and hang downwards. Owing to different lengths of stalk, &c. , the 1., looked at from above, form a very good mosaic, and are all equally exposed to light. Infl. mixed, the primary structure racemose, the lat. branches cymose (cmcinni). Upper fls. i, anther 4-loc.) : Celosia. 2. Amaranteae (ov. t; anther 4-loc.): Amaranthus, Aerva, Ptilotus. 3. Gomphreneae (anther 2-loc.) : Gomphrena, Alternanthera, Iresine. Amarantellus Spegazz. Amarantaceae (2). i Arg. Amaranthus L. (incl. Blitum L.). Amarantaceae (2). 45 trop. and temp. Cult. orn. fl. (love-lies-bleeding, prince's feather). Infl. of c» fl. A. gangeticus L., &c., are pot herbs in India, &c.; A. cattdatus L., A. paniculatus L., &c., give ed. grain, used as a cereal in trop. As. Amarella Gilib., Raf.. Griseb. (Gentiana L. p.p.). Gent. (i). 15 N. Am. Amarenus C. Presl-Trifolium L. p.p. (Legum.). Amaroria A. Gray. Simarubaceae. i Fiji. Amarus (Lat.), bitter. Amaryllidaceae (EP., BH. incl. Velloziac.). Monocots. (Liliiflorae). 75 gen-> 7°° SP- mostly tr°P- or sub-trop. Living in dry climates, they AMBROSINIA 31 are usu. xero., many bulbous, leafing only in the rains, some covered with wax (Agave), ill have ordinary stems, many have rhiz. Infl. usu. on a scape, with spathe, always cymose, but often umbel- or head-like by condensation. Fl. $ , reg. or •[• (transv. so in Anigo- zanthos). P 3 + 3 petaloid, A 3 + 3, G (3), rarely £-inf., 3-loc. with axile plac. and oo anatr. ov. In some (Narcissus and allies) there is a conspic. corona, looking like an extra whorl of P, between the normal P and the sta., sometimes looked on as combined ligular out- growths of the 1. of P, sometimes as stipules of sta. (as seen in various stages in the series Caliphruria, Sprekelia, Eucharis, Narcissus). Caps, or berry. Veg. repr. by bulbils common. [BH. chars. (Atnaryllideae}, incl. fl. sol., A oo in bundles, placentae lamellate ± thickened at outer side.] Classification and chief genera (after Pax) : Anthers introrse. I. AA1ARYLLID01DEAE (bulbous, scapigerous) : Haeman- thus, Galanthus, Amaryllis, Crinum, Eucharis, Narcissus. II. AGAVOIDEAE (rhiz.; 1. fleshy, in rosettes): Polianthes, Agave, Furcrea. III. HYPOXIDOIDEAE (rhiz.; stem with small ordinary 1.): Alstroemeria, Bomarea, Anigozanthos. Anthers extrorse. IV. CAMPYNEMAT01DEAE: Campynema. Amaryllis L. (Belladonna Sweet). Amaryllidaceae (i). i Cape Col. A. bdladonna L., cult. orn. fl. (the A. of greenhouses = H ippeas- trum). Amasonia L. f. Verbenaceae (i). 6 Brazil to Trinidad. Amatnad (Cl. ), a sandhill pi.; amatMum, a sandhill formation. Amauria Benth. Compositae (6). 2 SW.U.S. Amauriella Rendle. Araceae (iv). i Nigeria. Amauriopsis Rydberg. Compositae (6). i U.S. Ambaitaa Barrere ex O. Ktze. = Cecropia L. (Moraceae). Ambelania Aubl. Apocynaceae (i. i). 6 Brazil to Venezuela. Amberboa Less., Pers.^Volutarella Cass., Centaurea L., &c. Ambiguous, of uncertain origin or doubtful position. Amtalogyna Raf. = Amaranthus L. p.p. (Amarant.). Amblostoma Scheidw. Orchidaceae (n. 6). 3 trop. S. Am. Amblyanthera Muell.-Arg. =Mandevilla Lindl. (Apocyn.). Amblyanthopsis Mez. (Ardisia Sw. p.p.). Myrsin. (n. i). 2 Indomal. Amblyanthus A. DC. Myrsinaceae (11. i). 3 Assam. Amblygonocarpus Maims. Leguminosae (i. 4). i Centr. Afr. Amblyocalyx Benth. Apocynaceae (i. 3). 2 Borneo. Amblyocarpum Fisch. et Mey. Compositae (4). i near Caspian Sea. Amblyopappus Hook, et Am. Compositae (6). 2 Chili. Amblystigma Benth. Asclepiadaceae (n. i). 2 Arg., Bolivia. Ambora Juss. = Tambourissa Sonner. (Monim.). Amborella Baill. Monimiaceae. i New Cal. Amboyna wood, Pterocarpus mdicits Willd. (?). Ambrosia L. Compositae (5). 15 Am., Afr. trop. Heads unisex. the ? one-flowered. Fr. enclosed in the invol. Ambrosinia L. Araceae (vn). i Medit. 32 AMBULIA Ambulia Lam. (Limnophila R. Br., Terebinthina Rumph. ex O. Ktze.). Scrophulariaceae (ll. 6). 30 palaeotrop. Amburana Schwacke et Taub. (Torresea Allem. EP.). Legum. (11. 3). i Minas Geraes, Brazil. Good timber. Ameghinoa Spegazz. Compositae (12). i Patagonia. Amelanchier Medic. Rosaceae (11). 10 N. temp. Ameletia DC. =Ammannia Houst. (Z?//.) = Rotala L. (Lythr.). Amellus L. Compositae (3). 10 S. Afr. Amentaceae, the catkinate families, Salicaceae, Juglandaceae, Betula- ceae, Fagaceae; amentaceous, catkin-bearing; amentum, a catkin. American aloe, Agave; -cowslip, Dodecatheon\ -ebony, Brya Ebemis DC.; -elemi, Bursera gummifera L. ; -fly-trap, Apocynum andro- saemifolium L. ; -laurel, Kalniia ; -mastic, Sckimis molle L. ; -water- weed, Elodea canadensis Michx. ; -witch-elder, Fothergilla. Amerimnon P. Br. = Dalbergia L. f. (Legum.). Amethystea L. Labiatae (I. i). i Siberia, E. Russia. Amherstia Wall. Leguminosae (n. 3). i Burma, A. nobilis, Wall., a tree often cult, for its splendid fl. Stalk and br. as well as pets, are bright pink. Sta. united in a tube. The young L, covered with brownish spots, hang down " as if poured out " ; later they stiffen, turn green and come to the horiz. position (Keeble, Ann. Bot. IX. 59). Ainianthlum A. Gray (Zygadenus Michx.). Liliaceae (i). i All. Am. Amicia H. B. et K. Leguminosae (in. 7), 5 Andes. In A. Zygomeris DC. the large stips. protect the bud. Ammannia (Houst.) L. Lythraceae. 20 cosmop. Ammi (Tourn. ) L. Umbelliferae (in. 5). 7 Medit., trop. Afr. Ammiopsis Boiss. Umbelliferae (in. 2). 2 NW. Afr. Ammobium R. Br. Compositae (4). i New S. Wales. A. alatum R. Br. cult, for the fl. heads, dried to form •' everlastings." Ammobroma Torr. Lennoaceae. i New Mex., Calif. Ammocallis Small = Vinca L. (BH.) = Lochnera Reichb. (Apocyn.). Ammocharis Herb. Amaryllidaceae (i). 2 S. and trop. Afr. Ammochloa Boiss. Gramineae (10). 2 Medit. Arnmochtliad (Cl.), a sandbank pi. ; -thium, a sandbank formation. Ammodaucus Coss. et Dur. (Daucus p.p.). Umbellif. (in. 8). i Algeria. Ammodendron Fisch. ex DC. Leguminosae (in. i). 5 W. As. Ammodenia Patrin = Arenaria L. p.p. (BH.) = Alsine Scop. p.p. Ammoniacum, gum-, Dorema ammoniacum D. Don. Ammophila Host. Gramineae (8). 4 N. temp. A. (Psamma) arundi- nacea Host, (marram) common on sandy coasts in Brit., and much used for sand-binding. After some years a light soil forms, in which other pi. take root. The 1. curl inwards in dry air. Ammoselinum Torr. et Gray. Umbelliferae (in. 5). 3 N. Am. Ainmospenna Hook. f. Cruciferae (2). 2 Medit. Ammothamnus Bunge. Leguminosae (in. i). 2 W. As. Amoenus (Lat. ), sweet, pleasant. Amomum L. Zingiberaceae (2). 150 paleotrop. Fl. usu. on scapes from the rhiz. ; wings and keel absent, standard folding round sta.- tube at base. Prolog, with persistent stigma. Amoora Roxb. Meliaceae (in). 20 Indomal. Amoreuxia Mo£. et Sesse. Cochlosperm. (Bixin. BH.). 3 Centr. Am. AM PHIL O CHI A 3 3 Amoria C. Presl = Trifolium Tourn. p.p. (Legum.). Amorpba L. Leguminosae (in. 6). 15 N. Am. Wings and keel o; stan Jard folds round base of sta.-tube. Prolog, with persistent stigma. Amorphocalyx Ivlotzsch = Sclerolobium Vog. p.p. (Legum.). i Guiana. Amorphopballus Blume (incl. Hydros/tie, Synantherias, BH.). Araceae (iv). 30 trop. As. Usu corm like rhiz., giving yearly a big 1. (up to 10 ft.) and infl. (in A. Titanum Becc. 3 ft. high), with $ fl. above and ? below. Its dirty red colour and foetid smell attract carrion flies, which sometimes lay eggs on the spadix. Amorphospermum F. Muell. {Lucumn p.p. BH.). Sapot. (i). i trop. E. Austr. Amorphous, shapeless. Ampacus Rumpn. ex O. Ktze. =Evodia Forst. (Rutac.). Ampalis Boj. Moraceae (i). 2 Madag. Ampelanus Raf. = Enslenia Nutt (Asclep.) 3 W. N. Am. Ampelidaceae, Ampelideae {BH.) = Vitaceae. Ampelocera Klotzsch. Ulmaceae. 2 trop. Am., W. I. Ampelocissus Planch. (Iritis L. p.p. BH.). Vitaceae. 65 trop. Ampelodaphne Meissn. Lauraceae (11). 5 Brazil, Guiana Ampelodesma Beauv. Grammeae (10). i Medit. When young used as fodder. The 1. used like esparto (Stipa). Ampelodonax Lojac. (Arundo Tourn. p.p.). Gramin. (10). i Sicily. Ampelopsis (L. C. Rich, in) Michx. p.p. ( Vitis L. p.p. BH.). Vitaceae. 24 temp, and subtrop. As. Am. For garden A. cf. Parthenocissits. Ampelosicyos Thou. =Telfairia Hook. (?) Cucurbitac. i Madag. Ampelothamnus Small (Andromeda p.p.). Ericaceae (n. i). i Florida. Amperea A. Juss. Euphorbiaceae (B. n). 6 Austr., Tasm. Arupherepbis H. B. et K. =Centratherum Cass. ^Compos.). Amphi- ((ir. pref.), both; -bious pi., pi. which can live in water or on land, Peflis, Pfffygonum ; -carpic, with two kinds of fr. , Aethionenia, Cardamine, Dickondra, Dimorphotheca ; -mixis, sexual repr. ; -tropous (ovule), turning both ways on stalk. Ampbiacbyris Nutt. (Gutierrezia Benth. p.p.). Comp. (3). 2 Calif. Ampbiantbus Torr. Scrophulariaceae (ll. 6). i Georgia. Ampbibecis Schrank-=Cemratherum Cass. (Comp.). Amphiblemma Naud. Melastomaceae (i). 5 trop. W. Afr. Cult. orn. Ampbiblestra Presl. Polypodiaceae. i Venezuela. Ampbibolis C. Agardh. (C_ymodoceaK.on.p.p. EP.). Potamoget. i Austr. Amphibromus Nees. Gramineae (9). i Austr. Amphicarpaea Ell. Leguminosae (in. 10). 18 trop. and N. Am., E. As. Some have cleist. fl. below, which give subterranean fr. like Arachis. Amphlcarpum Kunth (-on Raf.). Gramineae (5). 2 S.E. U.S. Amphicome Royle. Bignoniaceae (2). 2 Himal. AmpMcosmia Gardn. = Hemitelia R. Br. (Cyatheac.). Ampbidetes Fourn. Asclepiadaceae (n. i). 2 Rio de Janeiro. Ampbidonax Nees = Arundo L. and Zenkeria Trin. (Gram.) AmpWdoxa DC. Compositae (4). 6 S. and trop. Afr., Madag. Ampbiestes Sp. Moore. Acanthaceae (iv. B). i Madag. Amphigena Rolfe. Orchidaceae (n. i). 2 Cape Colony. Ampbiglossa DC. Compositae (4). 4 S. Afr. AmpMlocbia Mart. =Qualea Aubl. p.p. (Vochys.). W. -2 34 AMPHILOPHIS Ampbilopbis Nash (Andropogon L. p.p.). Gramineae (2). 4 U.S. Ainpbilopbium Kunth. Bignoniaceae (i). 10 warm Am. Ampbimas Pierre. Leguminosae (n. 8). 2 Gaboon. Arnpliiodon Huber. Leguminosae (in. 6). i Amazon valley. Ampbipogon R. Br. Gramineae (8). 6 Austr. Ampbirbapis DC. = Inula L., Microglossa DC., Solidago L. Ampbirrbox Spreng. Violaceae. 3 trop. Am. Ampbistelma Griseb. =Vincetoxicum (.£.//.);= Metastelma (Ascl.). Ampbitecna Miers. Bignoniaceae (4). 2 Mex. Ampbitbalea Eckl. et Zeyh. Leguminosae (in. 3). 10 S. Afr. Ampborantbus Sp. Moore =Phaeoptilum Radlk. p.p. (Nyctag.). Ampborcbis Thou. = Cynorchis Thou. p.p. (Orchid.). Ampborella T. S. Brandegee. Asclepiad. (n. i). i Mex. Ampboricarpus Vis. Compositae (i i). 2 S.E. Eur. Ampborocalyx Baker. Melastomaceae (i). i Madag. Amphymenium H. B. et K. = Pterocarpus L. p. p. ( Legum.). 1 5 trop.S.Am. Amplectecs (Lat.), embracing ; amplexicaul, clasping the stem. Amsinckia Lehm. Boraginaceae (iv. 2). 15 Pac. Am. Amsonia Walt. Apocynaceae (i. 3). 10 N. Am., Japan. Amydrium Schott (Epipremnum £P.). Arac. (ii). i Malay Arch. Amygdalopsis Carr. — Prunus Tourn. p.p. (Rosac ). Amygdalus (Tourn.) L. = Prunus Tourn. p.p. (Rosac.). Amylaceous, starchy; amyloid, starch-like. Amylocarpus Barb. Rodr. (Buctris'L. p.p.). Palmaceae(iv. 2). 2oBrazil. Amyris P. Br. Rutaceae (iv) (Burseraceae BH.). 12 trop. Am., W.I. Anabasis L. Chenopodinceae (B). 20 Medit., C. As. Anabata WiHd.=Faramea Aubl. (Logan.). Anacampseros L. Portulacaceae. 15 S. Afr. Xero. with fleshy 1., and buds protected by bundles of hair, representing slips. Anacamptis Rich. (Orchis p.p. BH.}. Orchid, (n. i). i Eur. N. Afr. Anacampt-orcbis x G. Camus, hybrid with Orchis. 2 S.W. Eur. Anacardiaceae (EP.; £>H.\nc\. C0rynocarpaceae,Julia.niaceae), Dicots. (Archichl. Sapindales). 60 gen., 500 sp., chiefly trop., but also Medit., E. As., Am. Trees and shrubs with alt. exstip. 1., and panicles of oo fl. Resin-passages occur, but the 1. are not gland- dotted (hence they cannot be confounded with Rutaceae). Recept. convex, flat, or concave ; gynophores, etc., occur. Fl. typically 5-merous, reg., hypog. to epig. ; A 10—5 or other number; G (3-1) rarely 5, each with i anatr. ov., often only one fertile. Usually drupe with resinous mesocarp ; embryo curved ; no endosperm. The fr. of Mangifera, Anacardium, Spondias, Pistacia, &c., are important. Rhus furnishes various useful products. [BH. cbars. include chars, of Corynocarpus and Julianiaceae, both rare; fam. in Sapindales.] Classification and chief genera (after Engler) : A. 5 free cpls., or i. L. simple, entire: 1. Mangifereae: Mangifera, Anacardium. B. Cpls. united. L. rarely simple : 2. Spotidieae (ovules in each cpl.) : Spondias. 3. Rhoideae (i ovule only, ovary free) : Pistacia, Rhus. 4. Semecarpeae (do., ovary sunk in axis) : Semecarpus. C. Cpl. i. ? fl. naked. L. simple, toothed: 5. Dobineae: Dobinea (only genus). ANASTROPHUS 35 Anacardium L. Anacardiaceae (i). 8 trop. Am.; A. occidental L. (cashew-nut) largely cult. Fl. polygamous. Each has i cpl. yielding a kidney-shaped nut with hard acrid coat. The nut (promotion nut, coffin-nail) is ed. Under it the axis swells up into a pear-like body, fleshy and ed. The stem yields a gum like arabic. Anacnaris Rich. = Elodea Michx. (Hydrochar.). Anacolosa Blume. Olacaceae (Olacin. BH.\. :o Indomal. Anacyclus L. Compositae (7). 15 Medit. Some offic. (radix pyrethri). Anadendrum Schott. Araceae (i). 6 Indomal. Anadenia R. Br. = Grevillea R. Br. p.p. (Proteac.). Anaectocalyx Triana. Melastomaceae (i). 2 Venezuela. Anagallis (Tourn.) L. Primulaceae. 18 Eur., As., Afr., S. Am., 2 Brit, (pimpernel). The fl. of A. ai-vensis L. (poor man's weather glass) closes in dull or cold weather. Anaglypha DC. Compositae (4). 2 S. Afr. Anagosperma Wettst. Scrophulariaceae (ill. 3). i N.Zealand. Anagyris L. Legum. (m. 2). 2 Medit. A. Lour. = Ormosia Jacks. Analogous (organs), agreeing in function, but not in descent, mode of origin, nor position; e.g. the r.-like 1. of Salvinia are analogous to r. Anamirta Colebr. Menispermaceae. 7 Indomal. The achenes of A. Coccnlus Wight et Am. (Cocculus indicus) are used to adulterate porter. In the angles between the big veins of the 1. are little cavities covered by hairs and inhabited by mites (acaro-domatia). Anamomis Griseb. = Eugenia L. p.p. (Myrt.). 5 W.I. Ananas Tourn. ex L. (Ananassa Lindl.). Bromeliaceae (4). 5 trop. Am., incl. A. sativus Schult., the pine-apple, largely cult, in Sandw. Is., Singapore, &c. Stem short and leafy, terrestrial, bearing a term. infl., which after fert. forms a common mass, fr. bracts, and axis, while the main axis grows beyond and forms a tuft of 1. — the crown of the pineapple. Some vars. cult. orn. 1. Anandrous, without sta. Ananthaeorus Underw. et Moxon ( Taenitis p.p.) Polypod. i trop. Am. Anantherix Nutt. = Asclepiodora A. Gray (BH. ) - Asclepias L. p.p. Anapeltis J. Sm. =Polypodium L. (Filic.). Anaphalis DC. Compositae (4). 40 As., Eur., Am. Hairy. Anaphrenium E. Mey. (Heeria Meissn. EP.}. Anacard. (3). 7 Afr. Anaphyllum Schott. Araceae (iv). 2 S. India. Anarrhinum Desf. (Simbulda EP.). Scroph. (n. 3). 12 Medit. Anarthria R. Br. Restionaceae. 6 S.W. Austr. Anarthrophyllum Benth. Leguminosae (in. 3). 12 Andes. Anartia Miers (Tabernaemontana L. p.p.). Apocyn. (i. 3). 7 trop. Am. Anastatica L. Cruciferae(4). i E. Medit., A. hierochuntina L. (rose of Jericho). While the seeds are ripening in the dry season the 1. fall off and the branches fold inwards, reducing the pi. to a ball of wickerwork, which rolls about with the pods closed until it reaches a wet spot, or the rainy season begins. Anastomosis, reunion of branches. Anastrabe E. Mey. Scrophulariaceae (n. 4). i S. Afr. Anastraphia D. Don. Compositae (12). 10 W. Ind. Anastropnus Schlecht.-Paspalum L. p.p. (Gram.), i U.S. 3 — 2 36 ANATHERUM Anatherum Beauv. = Andropogon L. p.p. (Gram.). Anatropanthus Schlecht. Asclepiadaceae (n. 3). i Borneo. Anatropous (ovule), reversed on stalk. Anaxagorea St. Hil. Anonaceae (i). 15 trop. As., Am. Anaxeton Gaertn. Compositae (4). 7 S. W. S.Afr. Ancad, a canon plant. Anceps (Lat.), ancipitous, two-edged. Anehietea A. St. Hil. Violaceae. 4 trop. S. Am. Anchomanes Schott. Araceae (iv). 5 trop. Afr. Anchonium DC. Cruciferae (4). 3 W. As. Anchovy pear, Grias caulijlora L. Ancnusa L. Boragin. (iv. 3). 45 Eur., N. Afr., W. As. A. officinalis L. was formerly offic., and is widely scattered (escape in Brit.). Ancistranthus Lindau. Acanthaceae (iv. B). i Cuba. Ancistrocarpus Oliv. Tiliaceae. 2 trop. W. Afr. Ancistrocarya Maxim. Boraginaceae (iv. 4). i Japan. Ancistrochilus Rolfe. Orchidaceae (n. 9). 2 trop. Afr. Ancistrocladaceae (EP. Dipterocarpaceae, p.p. BH.}. Dicots. (Archi- chl. Parietales.) Only gen. Ancistrocladus (q-v.}. Ancistrocladus Wall. Ancistrocladaceae. 12 palaeotrop. Sympodial lianes, each member ending in a watch-spring tendril. L. alt., lanceolate, entire, with minute stips. Racemose infl. of § reg. fl. K 5, teeth unequal; C (5) slightly united, convolute; A 5 or 10; G i-loc. with i basal erect semi-anatr. ov. Nut. Endosperm. Ancistrophyllum G. Mann et H. Wendl. Palmae (in). 4 W. Afr. Ancistrorhynchus Finet. Orchidaceae (n. 20). 2 trop. Afr. Ancistrum Forst. = Acaena L. (Rosac. ). Ancrumia Harv. Liliaceae (iv). i Chili. Ancylacanthus Lindau. Acanthaceae (iv. A), i New Guinea. Ancylanthos Desf. Rubiaceae (n. i). 4 trop. Afr. Ancylobothrys Pierre (Landolphia p.p.). Apocyn. (i. i). 6 trop. Afr. Ancylocladus Wall. = Willughbeia Roxb. (Apocyn.). Ancylogyne Nees = Sanchezia Ruiz et Pav. (Acanth.). Andersonia R. Br. (Sprengelia p.p. EP.}. Epacrid. (2). 20 W. Austr. Andes rose, Be/aria racemosa Vent, and other spp. Andira Lam. ( Vonacapoua Aubl.). Legum. (in. 8). 25 trop. Am., Afr. A. inermis H.B. et K. (angelin) is a rain-tree (cf. Pithecolobium) ; its wood (partridge-wood) is useful. Andiroba, Carapa spp. Andrachne L. Euphor'oiaceae (A. i. i). 15 trop. and subtrop. Andradaea Allem. Nyctaginaceae (? Phytolac.). i Rio de Janeiro. Andradia T. R. Sim. Leguminosae (n. 9). i trop. Afr. Andrea Mez. Bromeliaceae (4). i C. Brazil. Andreoskia (Andrzeiowskyd) Reichb. Crucif. (2). i E. Medit. Andriapetalum Pohl = Panopsis Salisb. (Proteac.). Andro- (Gr. pref.), male; -dioecious, 5 and (. and N. Z. The petioles of A. (Arch.) officinalis are offic., and are used in confectionery. Angelin tree (W. I.), Andira inermis H.B. et K. Angelocarpa Rupr. (Coelopfeurum Ledeb.). Umbel, (in. 5). r C. As. Angelonia Humb. et Bonpl. Scrophul. (11. i). 30 trop. Am., W. Ind. Angianthus Wendl. Compositae (4). 25 temp. Austr. Heads cpd. Angico gum, Piptadenia rigida Benth. Angiopetalum Reinw. Myrsinaceae (inc. sed.). i Java. Angiopteris Hoffm. Marattiaceae (i). 65 Madag., Indomal., A.evecta Hoffm. Large ferns with the sori not united into synangia as in most M. Annulus like that of Osmundaceae at apex of sporangium. The r. arise close to apex, and burrow down and outwards through the stem and leaf-bases, emerging some distance down. Angiospermae. One of the two great divisions of Spermaphyta, dis- tinguished from Gymnosperms by the fact that the cpls. are so infolded or arranged as to form an ovary in which the ovules are borne. Endosperm formed after, instead of before fert. All A. possess true fl., the essential parts of which are sta. and cpls. The former bear pollen-sacs ( = microsporangia of Pteridophyta), the latter ovules (megasporangia). The ovule is always enclosed in the cpl. ; it has two (or one) integuments, and usually one embryo- sac (more in some chalazogamic forms, Loranthaceae, &c.). The pollen-tube may enter by the micropyle or by the chalaza (cf. Chala- zogamae). Parthenogenesis, or development of the ovum into an embryo without fert., occurs in Antennaria, Akhemilla, Hieraciiim, Houttttynia, Thalictrum, IVikstroeiuia, &c., embryo formation by adv. budding in Alc/tornea, Citrus, Euonymus, Funkia,Nothoscordum, &c., apogamy (cf. Filicineae) in Balanophora. A. are divided into Mono- and Di-cotyledons (cf. classification at end of the book). Angkalanthus Balf. f. Acanthaceae (iv B). i Socotra. Angolaea Wedd. Podostemaceae. i Angola. ANISOMERIS 39 Angophora Cav. Myrtaceae (n. i). 5 E. Austr. AngorcMs Thou. = Angraecum Bory (Orchid.). Angostura bark, Cusparia febrifuga Humb. Angostyles Benth. Euphorbiaceae (A. II. 2). i N. Brazil. Angraecopsis Krzl. Orchidaceae (n. 20). i trop. E. Afr. Angraecum Bory. Orchidaceae (n. 20). 120 trop. Afr., Madag., Masc. Monopodial epiphytes, often cult. A. sesqinpfdale Thou, (wax-fl.) has an enormous spur a foot long, secreting honey at the bottom and is probably fert. by a moth with equal proboscis (cf. Yucca, Ficus). See Darwin's Orchids, p. 162. Some are leafless. Anguillaria R. Br. Lili. (i). 2 Austr., Tasm. A. Gaertn. = Ardisia. Anguillicarpus Burkil!. Cruciferae (2). i Beluchistan. Anguloa Ruiz et Pav. Orchid, (n. 12). 5 Peru, Colombia. Anguria (Tourn.) L. Cucurbit. (2). 25 tiop. Am. AnguriopsisJ. R. Johnston. Cucurbit. (2). i Venezuela. Angusti- (Lat. pref.), narrow-; -folius (Lat.), -leaved. Angylocalyx Taub. Leguminosae (in. i). 6 trop. Afr. Anhalonimn Lem. = Mammillaria Haw. (Bff.) = Ariocarpus Scheidw. Aniba Aubl. (Aydendron Nees, Ocotea BH). Lauraceae (n). 55 trop. Am. Anigozanthos Labill. Amaryll. (in) (Haemod. BH.}. 8 S.W. Austr. Fl. transversely -|- . Anil Ludw. ex O. Ktze. = Indigofera L. (Legum.). Animals, seed-dispersal by, cf. Seed-dispersal. Anime (resin), copal. Anisacantha R. Br. (Bassia All. p.p. EP.}. Chenopodiaceae (A). 6 Austr. Anisacanthus Nees. Acanthaceae (iv. B). 10 Am. Anisadenia Wall. Linaceae. 2 Himalaya. Anise, aniseed, Pimpinella Anisum L.; star-, -tree, Illiciiim verum Hook. f. Aniseia Choisy (Ipomoea L. p.p. BH.}. Convolv. (i). 15 trop. Aniselytroa Merrill. Gramineae (8). i Phil. Is. Aniserica N. E. Br. Ericaceae (iv. 2). i S. Afr. Aniso- (Gr. pref.), unequal- ; -merous, with — numbers in the whorls; -phylly, with — 1. at same node, Anisophyllea, Centradenia, Colum- nea, Gardenia, Gesneriaceae, Khigia, Melastoniaceae, Nyctaginaceae, Philadelphus, Randia, Sambitcus, Scrophularia, Strobilanthes, Taber- naemontana. Anisocarpus Nutt. (Madia Molina). Compositae (5). 5 W. U.S. Anisochaeta DC. Compositae (4). i S. Afr. Anisochilus Wall. Labiatae (vn). 20 trop. As., Afr. Anisocoma Torr. et Gray. Compositae (13). i W. U.S. Anisocycla Baill. Menispermaceae. 6 S. and trop. Afr., Madag. Anisodus Link et Otto = Scopola Jacq. p.p. (Solan.). Anisolobus A. DC. = Odontadenia Benth. (Apocyn.). Anisolotus Bernh. = Iiosaclda Dougl. (Legum.). 5 U.S. Anisomallon Baill. Icacin. (Olacin. BH.}. i New Caled. Anisomeles R. Br. Labiatae (vi). 6 Indomal. Anisomeria D. Don. Phytolaccaceae. 3 Chili. Anisomeris Presl (Chomelia Jacq.). Rubi. (n. 2). 17 trop. S. Am. 4o ANISONEMA Anlsonema A. Juss. = Phvllanthus L. p.p. (Euphorb.). Anisopappus Hook, et Am. Compos. (4). 8 S. China to S. Afr. Anisophyllea R. Br. Rhizophoraceae. 15 palaeotrop. It differs from other R. in its drupe fr., exalb. seed, and alt. exstip. sometimes anisophyllous 1. AnisophyUum Haw. = Euphorbia L. p.p. do Jacq. Inc. sed. i Baru. Anisopoda Baker Umbell. (in. 5). i Madag. Anisopogon R. Br. Gramineae (9). I Austr. Anisoptera Korth. Dipterocarpaceae. 20 E. Indomal. Anisopus N. E. Br. Asclepiadaceae (n. 3). 2 trop. W. Afr. Anisosciadium DC. (Kchinophora L. BH.). Umbell. (in. i). i W. As. Anisosperma Manso. Cucurb (i). i Brazil. Seeds medicinal. Anisostachya Nees (Jnsticia BH.}. Acanth. (iv. B). 5 trop. Afr., Madag. Anisostigma Schinz. Aizoaceae (n). i Namaland. Anisotes Nees. Acanthaceae (iv. B). 8 trop. Afr., Arabia. Anisothrix O. Hoffm. Compositae (4). i S. Afr. Anisotoma Eenzl. Asclepiadaceae (n. 3). i S. Afr. Anisotome Hook. f. = Aciphylla Forst. (Umbell.). Annatto, Bixa Orellatia L. Anneslea Wall. (Mountnorrisia Szysz.). Theac. 2 Indomal. Annesorrniza Cham, et Schlechtd. Umbelliferae (ill. 5). 10 S.Afr. A. capensis C. et S. has ed. roots. Annexed, adnate. Annona L. = Anona L. Annotinus (Lat.), applied to branches of last year's growth. Annual, annuus (Lat.), living one year only. Annulate, marked with rings. Annulus, Filicineae Lcptosp. Anochilus Rolfe. Orchidaceae (n. i). ? S. Afr. Anoda Cav. Malvaceae (2). 10 trop. Am. Cult. orn. Anodal, in the upward direction on the genetic spiral. Anodendron A. DC. Apocynaceae (n. i). 10 Ceylon to China. Anodiscus Benth. Gesneriaceae (n). i Peru. Anodopetalum A. Cunn. Cunoniaceae. i Tasmania. Anoectochilus Blume. Orchidaceae (11. 2). 25 Indomal. Cult. orn. 1. Anoectomaria x Rolfe. Hybrid of last with Haemaria. Anogeissus Wall. Combretaceae. 5 trop. Afr., As. Anogra Spach (Oenothera L. p.p.). Onagrac. (2). 20 N. Am. Anogramma Link. Polypodiaceae. 10 trop. Am., Japan. Anoiganthus Baker. Amaryllidaceae (I). 2 Natal and trop. Afr. Anomalanthus Klotzsch = Simochilus Klotzsch (Eric.). Anomanthodia Hook. f. (Randia p.p. EP.}. Rubiac. (i. 8). i trop. As. Anomatheca Ker-Gawl. = Lapeirousia Pourr. (Irid.). Anomianthus Zoll. Anonaceae (i). i trop. As. Anomochloa Brongn. Gramineae (6). i Brazil. Anomopanax Harms. Araliaceae (3). 5 Malay Arch. Anomospermum Miers. Menispermaceae. 8 Brazil, Guiana. Anomostephium DC. =Aspilia Thou (Compos.). Anomotassa K. Schum. Asclepiadaceae (n. 3). i Ecuador. Anona L. .Anonaceae (4). 70 trop. esp. Am. Fr. aggregate, often ANTENNARIA 41 very large, made up of the individual berries derived from the separate cpls., sunk in, and united with, the fleshy recept. That of some cult, sp. is ed., e.g. of A. Cherimolia Mill, (cherimoyer; trop. Am.), A. sqtianiosa L. (sweet sop, custard or sugar apple; E. Ind.), A. muricata L. (sour sop; trop. Am.) and A. reticnlata L. (custard-apple or bullock's heart; trop. Am.). A. (Geantheinuni) rhizantha Eichl. (Braz.) has rhizomes below the soil, bearing scale leaves only. The fls. are borne on branches of these above the ground. Anonaceae (EP. ; BH. incl. Eupomatiaceae). Dicots. (Archichl. Ra- nales). 46 gen. 820 sp. chiefly trop. (esp. Old World). Trees and shrubs (exc. one) with usu. two-ranked undivided exstip. 1. Stem sometimes sympodial, at least in infl. Oil passages present. Fls. reg. ? (rarely unisex.), solitary or in infl. of various types. Usu. formula P 3 + 3 + 3 (one or two outer whorls sepaloid); A oo (rarely few), spiral, hypog. ; G oo (Monodora is syncp.). Ovules usu. ao , ventral or basal, anatr. Fr. commonly an aggregate of berries, when many-seeded frequently constricted between the seeds. In Anona, &c. the berries coalesce with the receptacle. Ruminate en- dosperm (the chief character that separates A. from Magnoliaceae). Many yield ed. fr., e.g. Anona, Artabotrys. [BH. chars, incl. Po and fl. perig.] Classification and chief genera (after Prantl) : a. Apocarpous. 1. Uvarieae (P unjointed or clawed with claws against sta. ): Uvaria, Asirnina, Guatteria, Unona, Cananga. 2. Milhtseae (P valvate, usu. unequal, if clawed the claws away from sta.): Miliusa. 3. Hexalobeae (P equal, with crossfolds in bud) : Hexalobus. 4. Xylopieae (P hollow at base, ± constricted above it and again spread out or laterally compressed) : Xylopia, Arta- botrys, Anona. b. Syncarpous ; ov. uniloc. with parietal plac. 5. Monodoreae: Monodora (only genus). [Placed in Ranales by BH.} Anonidium Engl. et Diels. Anonaceae (i). 2 W. trop. Afr. Anonymus Walt. Inc. sed. (pro omnibus dubiis). Anoplophytum Beer = Tillandsia L. (Bromel.). Anopteris (Prantl) Diels. Polypodiaceae. i trop. Am. Anopterus Labill. Saxifragaceae (v). 2 E. Austr., Tasm. Anopyxis Pierre. Rhizophoraceae. i trop. Afr. Anotis DC. Rubiaceae (i. 2). 25 Indo-mal., i S. Am. Auotites Greene. Caryophyllaceae (n.' i). 20 N. Am. Anoumabia A. Chevalier. Sapindaceae. i Ivory Coast. Anplectrum A. Gray (Diplectria Rchb.). Melastom. (i). 20 Malaya. Anredera fuss. Basellaceae. i trop. Am. Ansellia Lindl. Orchidaceae (n. 5). 4 trop. Afr., Natal. Ante- (Lat. pref.), before; -petalous (sta.), opp. petals; -posed, opp. and not alt. with. Antelaea Gaertn. Inc. sed. 2 Java. Antennaria Gaertn. Compositae (4). 75 extra-trop., exc. Afr. A. 42 ANTENNARIA dioica Gaertn. (mountain everlasting, cat's foot) in Brit, is a small creeping dioec. perenn., hairy and semi-xero., occurring chiefly on hills and at the sea-shore, but not common in intermediate places. In A. alpina (L.) R. Br. only ? plants usu. occur, and show true parthenogenesis, the ovum developing into an embryo without fert. (not to be confused with the veg. budding of Alchornea). Anterior (side of fl.), facing the bract ; antero-posterior, median. Anthacanthus Nees. Acanthaceae (iv. B). 6 W. Ind. Anthaenantia Beauv. Gramineae (5). 8 warm Am. Anthagathis Harms. = Jollydora Pierre (Legum.). i trop. Afr. Anthelia Schott. Araceae (inc. sed.). i Celebes. Anthemis Mich, ex L. Compositae (7). 120 Eur., Medit. (4 Brit., chamomile). The fr. of A. arvensis L. has papillae on its upper surface which become sticky when wet (cf. Linum). Shows suborder chars, well. A. nobilis L. (chamomile) fl. offic. Anthephora Schreb. Gramineae (3). 10 trop. Am., trop. and S. Afr. Anther, cf. Stamen ; -idium, Filicineae Lepto, Pteridophyta ; -ozoid, spermatozoid, Pteridophyta. Anthericopsis Engl. Commelinac. (Liliac. BH. ). i E. trop. Afr. Anthericum L. Liliaceae (in). 130 Afr., Eur., Am. Antheropeas Rydberg. Compos. (6). 5 N. Am. Antherothamnus N. E. Br. Scrophular. (n. 4). i S. Afr. Antherotoma Hook. f. Melastom. (i). 2 trop. Afr., Madag. Antherura Lour. Rubiaceae (inc. sed. ). i S.E. As., Malay Arch. Antherylium Rohr. et Vahl (Ginoria Jacq. p.p. EP.). Lythr. 2 Mex., W. Ind. Anthesis, flower-opening. Anthistiria L. f. ( Themeda). Gramineae (2). 15 palaeotrop. A.vul- garis Hack, (kangaroo grass) covers large areas in Austr. and S. Afr. Antho- (Gr. pref.), flower- ; -carp, Nyctaginaceae ; -lysis, retrograde metamorphosis of fl. ; -philous, fl. -visiting ; -phore, stalk between K and C; -taxy, arrangement of fl. Anthobembix Perkins. Monimiaceae. 3 New Guinea. Anthobolus R. Br. Santalaceae. 5 Austr. Hypogynous. Anthobryum Phil. Frankeniac. (? Primul.). 2 Bolivia. Anthocarapa Pierre (Amoora p.p.). Meliac. (in). 2 Cochin China. Anthocephalus A. Rich. Rubiaceae (i. 6). 3 Malaya. Anthocercis Labill. Solanaceae (5). 20 Austr. Anthochlamys Fen/,1. Chenopodiaceae (A), i W. Centr. As. Anthochloa Nees et Meyen. Gramineae (10). 3 Am. Anthocleista Afzel. Loganiaceae. 25 trop. Afr. Anthodiscus G. F. W. Mey. . Caryocarac. (Ternstr. BH.}. 4 trop. S. Am. Anthodon Ruiz et Pav. = Salacia L. (Hippocrat.). Anthogonium Wall, ex Lindl. Orchid, (n. 9). i Himalaya, Burma. Antholoma Labill. Elaeocarp. 3 New Caled., New Guinea. Antnolyza L. Iridaceae (m). 25 Afr. Cult. orn. fl. Antnopogon Neck. (Gentiana L. p.p.) Gentian. 5 N. Am. Anthopterus Hook. Ericac. (in. 2) (Vaccin. BH.}. 10 Andes. Anthospermum L. Rubiaceae (n. 7). 35 Afr., Madag. Anthostema A. Juss. Euphorb. (A. n. 8). 3 trop. Afr., Madag. Fls. ANTIRRHOEA 43 in a cyathium like Euphorbia, but the axinns, Oleaceae, Parrot ia, Prin- glea, Sagina, Thaltctrutn, and cf. Cleistogamy. Apetlorhamnus Nieuwland (Kkamnns p.p.). Rhamn. i N. Am. Aphaenandra Miq. Rubiaceae (inc. sed.). i Sumatra. Aphaerema Miers. Elacourt. (8) (Samyd. BH.). i S. Paulo. Aplianactis Wedd. Compositae (5). 2 Andes. Aphanamixis Blume (Atnoora pp. BH.}. Meliac. (in). 8 Malaya. Aphanandrium Lindau (Neriacanthus). Acanth. (iv. B). i Columbia. Aphananthe Planch. • Ulmaceae. 5 Austr., E. As. Aphanelytrum Hackel. Gramineae (8). i Ecuador. Aphanes L. = Alchemilla L. (Ros.). Aphania Blume (Sapindus L. p.p. BH.}. Sapind. (i). 12 trop. As., Afr. Apbanisma Nutt. Chenopodiaceae (A), i California. Aphanocalyx Oliv. Leguminosae (n. 2). i Guinea. Aphanococeus Radlk. Sapindaceae (i). i Celebes. Aphanomyrtus Miq. Lauraceae (i). 3 Malaya. Aphanopetaluin Endi. Cunoniaceae. 2 S.E. and S.W. Austr. Aphanopleura Boiss. Umbell. (ill. 5). i Armenia. APO- 45 Aphanostephus DC. Compositae (3). 5 Mex., U.S. AphanostyHs Pierre (Clitandra p.p.). Apocyn. (i. i). 5 trop. Afr. Aphantochaeta A. Gray (Pentachaeta p. p. EP.). Compos. (3). 2 Calif. Aphelandra R. Br. Acanth. (iv. B). 60 warm Am. Cult. orn. fl. Aphelexis D. Don (Helickrysum p.p. EP.}. Compos. (4). 10 Madag. Aphelia R. Br. Centrolepidaceae. i S. Austr., Tasm. Apheliotropism, negative heliotropism. Aphloia Benn. (Neumannia A. Rich.). Flacourt. (4) (Bix. BH.). 4 S. Afr., Madag., Masc. Aphora Nutt. = Argithamnia P. Br. (BH. ) = Ditaxis Vahl p.p. Aphotic, dark. Aphragmus Andrz. (Braya p.p. BH.). Cruciferae (2). i Alaska. Aphyllanthes Tourn. ex L. Liliaceae (m). i. W. Meclit., A. mon- speliensis L. The sol. fl. is surrounded liy an invol. of or., and is regarded as the only remaining fl. of a head; the other fl. are only repres. by their bracts. Aphyllarum Sp. Moore. Araceae (vi). i Matto Grosso. Apfcyllon Mitch. (Orobanchc p.p. EP.). Orobanchaceae. 10 N. Am. Apiiyllorcliis Blume. Orchidaceae (n. 2). 12 Indomal. Aphylly, absence of 1. Apiastrum Nutt. Umbelliferae (in. 4). 2 N. Am. Apical (plac.), at upper end of ovary. Apicra Willd. Liliaceae (in). 10 S. Afr. Some, e.g. A.foliolosa\V]\\<\,, show extreme superposition of 1., forming almost solid masses of tissue. Many cult, orn fl. Apiculate, with small sharp point. Apinagia Tul. Podostemaceae. 16 Guiana, Brazil. Apio, Arracada xanthorhiza Bauer &c. Apiocarpus Montr. Staphyleaceae. i New Caled. Apiopetalum Baill. Araliaceae (3). 2 New Caled. Apios Moench. Leguminosae (in. 10). 8 N. Am.. E. As. A. tuberosa Moench is a climber with tuberous base. The keel of the fl. forms a tube which bends up and rests against a depression in the standard. When liberated by insects the tension of the keel makes it spring downwards, coiling up more closely, and causing the essential organs to emerge at the apex. Cult. orn. fl. Apium (Tourn.) L. Umbelliferae (in. 5). 4Ocosmop., 3 Brit. A.graveo- kns L. is the celery. Cult., and the etiolation of the leaf-stalks by heaping earth over them, render the garden form ed. A var. is the turnip-rooted celery or celeriac. Aplectrum Torr., Nutt. Orchidaceae (n. 9). 2 N. Am. Apleura Phil. Umbelliferae (i. 2). i Chili. AplolopMum Cham. (Haplolophium). Bignoniaceae (i). 3 Brazil. Aplopappus Cass. {Haplopappus Endl.). Compos. (3). 125 W. Am. Aplophyllum A. Juss. = Ruta L. (Rutaceae). Aplotaxis DC. = Saussurea DC. (Compos.). Apluda L. Gramineae (2). i Indomal., Madag. Apo- (Gr. pref.), from ; -carpous, with free cpls., Raminculaceae, Rosaceae, &c. ; -gamy, omission of sexual process, Angiospermae, Aspleninm, Balanophora, Elatostema, Filicineae Leptosp., Pteris, Todea ; -geotropic, -heliotropic, negatively g. or h., Bowenia ; 46 APO- -petalous, polypetalous ; -spory, Filicineae Leptosp., Dryopteris ; -tropous, anatr. with ventral raphe. Apocarpae (HH.}. The 6th series of Monocotyledons. Apochoris Duby. Primulaceae. i N. China. Apocopis Nees. Gramineae (2). 8 China to Madag. Apocynaceae (EP., BH.}. Dicots. (Sympet. Contortae). 130 gen., 1000 sp., mostly trop. ; a few temp.; Vinca minor gives the N. limit in Eur. Erect plants rare, mainly twining shrubs ; in the tropics many are large lianes. The stem has bi-collateral bundles ; latex always present. L. simple, usu. opp., entire, rarely with small interpet. stip. The primary type of infl. is a panicle, but in its later branchings it sometimes goes over into a dich. cyme or a cincinnus. Br. and bracteoles both present. Fir. 5 , reg., 5- or 4-merous. K (5), deeply lobed, quincuncial with odd sep. post.; C (5), usu. salver- or funnel-shaped, often hairy within, convolute (valvate in a few rare gen.) ; A 5, alt. with pets., epipet., with short incl. filaments; anther lobes full of pollen to the base, or empty at base and prolonged into rigid spines. Disc usu. present. G (2) or 2 (united by' style) or more, i- 01 2-loc. when syncp. Ov. GO , anatr., pend. ' Style usu. simple with thickened head. Berry, or more often 2 follicles. Seeds usu. flat and often with a crown of hairs serving for wind distr. Endosperm or none ; embryo straight. In the common A. with a large stylar head the stigma is at the edge or under surface of the head and self-fert. is almost impossible. An interesting mech. is found in Apocynum androsaemifohum. Many are showy ornamentals. Funtumia, Hancornia, Landolphia, Willugh- beia, &c. yield rubber. Several yield useful drugs, alkaloids, &c. Classification and chief genera (after K. Schumann) : I. PLUMIEROIDEAE (sta. free or only loosely joined to stylar head ; thecae full of pollen, rarely with spines ; seeds usu. without hairs) : i. Arduineae (syncarpous ; style not split at base) : Arduina, Allamanda, Landolphia. i. Pleiocarpeae (apocarpous ; style split at base ; more than 2 cpls.) : Pleiocarpa. 3. Plumiereae (ditto ; 2 cpls. ) : Plumiera, Amsonia, Vinca, Tabernaemontana, Cerbera. II. ECHITOIDEAE (sta. firmly joined to stylar head ; thecae empty at base, and with spines; seeds hairy) : 1. Echitideae (anthers included) : Apocynum, Nerium, Stro- phanthus, Dipladenia. 2. Parsonieae (anthers excluded): Parsonsia, Lyonsia, Wrightia. Apocynum (Tourn.) L. Apocyn. (n. i). 20 N. temp. Some cult., incl. A. androsaemifolium L. (American fly-trap), which has fl. mech. like Asclepiadaceae. Ov. crowned by disc, stigmatic below and surrounded by rigid lignified sta., at whose base is honey. An insect withdrawing its proboscis usu. brings it up the narrow slit between sta., at whose base is a drop of cement. Higher up the anthers open laterally, and thus the pollen is attached to the proboscis, but strength APPLE 47 is needed, and small insects are usu. caught, as may be seen in gardens. Seeds hairy (wind carriage). Apodanthera Arn. Cucurbitaceae (2). 20 warm Am. Apodanthes Poit. RafflesTac. (Cytinac. BH.). 5 trop. S. Am. Apodiscus Hutch. Euphorb. (A. I. i). i French Guiana. Apodocephala Baker. Compositae (i). 2 Madag. Apodolirion Baker. Amaryllidaceae (i). 6 S. Afr. Apodytes E. Mey. Icacinaceae. 14 S. Afr. to Malaya. A. dimidiata E. Mey. (S. Afr., white pear), good timber. Apogon Ell. (Serinia Raf.). Compositae (13). 3 N. Am. Apollonias Nees. Lauraceae (n). 2 Canaries, Madeira. Aponogeton L. f. Aponogetonaceae (Naiad. BH.). 15 palaeotrop., and S. Afr. Water pi. with sympodial tuberous rhiz. and basal L, usu. floating. Submerged 1. occur in some, e.g. A. (Otivirandra) fenestrale Hook. f. The whole tissue between the veins breaks up as the 1. grows, leaving a network of veins with holes between. The interior does not contain the usual intercellular spaces. The $ reg. fl. project above the water in spikes, sometimes divided longitudinally into 2 or 3 ; spathe early thrown off. P usu. 2, some- times 3 or even i, as in the much cult. A. distachyum Thunb. (Cape pondweed), where it is attached by a broad base, and looks like a br. In this sp. A oo , G 3 — 6, but usu. A 3 + 3, G 3, with 2 or oo ov. in each, anatr., erect. Fr. leathery. Embryo straight. Aponogetonaceae (£f.; Naiadcae p.p. BH.}. Monocots. (Helobieae). Only genus Aponogeton (q.v.}. Distinguished from Potamogetonaceae by coloured P and straight embryo, from Scheuchzeriaceae by P and sympodial structure. Apopetalum Pax. Rosaceae (i. i). i Bolivia. Apophyllum F. Muell. Capparidaceae (n). i N.E. Austr. Apoplanesia C. Presl. Leguminosae (in. 6). i Mex. Aporocactus Lem. (Cerats Mill. p.p.). Cactaceae (in. i). 2 N. Am. Aporosa Blume. Euphorb. (A. i. i). 35 Indomal. Aporosella Chodat. Euphorb. (A. I. i). i Paraguay. Aporrhiza Radlk. Sapindaceae (i). 3 Centr. Afr. Aporuellia C. B. Clarke. Acanth. (iv. A). 2 Malay Pen., Sumatra. Aporum Blume (Dendrobiuin Sw. p.p. BH.). Orchid, (n. 15). 12 trop. As. Aposeris Neck. (Hyoseris L. p.p. £H.). Compos. (13). i C. Eur. Mts. Apostasia Blume. Orchidaceae (i. i). 8 Indomal. Fl. almost reg. with 2 sta. ; ovary 3-loc. Appendicula Blume. Orchidaceae (n a. in.). 40 Malaya, Polynesia. Appendiculana O. Ktze. =next. Appendiculate, with appendages. Appendicularia DC. Melastomaceae (i). i Guiana. Apple, Pyrns Mains L. ; alligator-, Anona pahistris L. ; balsam-, Momordica bahamina L. ; custard-, Anona squamosa L., reticulata L. ; elephant-, Feronia Elephantum Correa ; kangaroo-, Solamim avicu- lare Forst. ; Kei-, Aberia caffra Harv. et Sond. ; love-, Lycopersictim escidentum Mill. ; Malay-, Eugenia malaccensis L. ; mammee-, Mammea americana L ; May-, Podophyllum pehatnin L. ; pine-, Ananas salivus Schult. f. ; rose-, Malay ; star-, Chrysophylluin 48 APPLE Cainito L. ; sugar-, Anona squamosa L.; thorn-. Daiur Stra- monium L. ; wood-, elephant-. Appressed, flattened down. Approximate, close together. Appunia Hook. f. Ruhiaceae (11. 9). 3 trop. S. Am. Aprevalia Bail!. Leguminosae (n. 7). i Madag. Apricot, Primus Armeniaea L. ; San Domingo-, RIammea americana L. Aptandra Miers. Olacaceae. 5 trop. S. Am., Afr. Apteria Nutt. Bunnanniaceae. 5 warm Am. Apteron Kurz. Khamnaceae. i Tenasserim. Apterous, wingless. Aptosimum Burchell Scrophular. (i. 2). 40 S. and trop. Afr. Aptotheca Mieis (Forsleronia) Apocyn. (inc. sed.). i Cuba. Apuleia Mart. Legum. (n. 5). 2 Brazil. A. praecox Mart excellent timber. Aquaticus (Lat.), living in water. Aquifoliaceae (EP. ; Ilicineae BH.}. Dicots. (Archichl. Sapindales ; Olacales BH.). 5 gen., 300 sp. temp, and trop. Shrubs and trees with leathery alt., 1. with minute or no stips and cymose infl. Fl. reg. unisexual, 3 — 6-merous ; no disc. Usu. K 4, €4, A 4, G usu. (4), 4-loc., each with i or 2 pend. anatr. ovules. Drupe. Endosp. Chief genus: Ilex. Aquifolium Tourn. ex Hall. = Ilex L. (Aquifol.). Aquilaria Lam. Thymelaeaceae. 8 Indomal., China. The wood of A. Agallocha Roxb. (Calambac, aloe-wood, eagle-wood), in about 8 °/0 of the trees, is saturated with resin (agar), used in India as a drug and perfume. Aquilegia (Tourn.) L. Ranunculaceae (2). 75 N. temp. A. vul garis L. (columbine), Brit., cult. orn. fl., with many others and many hybrids. Pets, with long spurs secreting honey (if. Delphinium). Fl. of class H, protandrous, visited by humble-bees. Sta. often 50 or more, in whorls of 5. Arabian coffee, Coffea arable a L. Arabidopsis Schur. (Sisymbr'nim L. Bff., Stenophragma EP.}. Cruc (2). 4 Eur , W. As. Arabis Linn. Cruciferae (4). 220 N. temp., S. Am., 5 Brit, (rock- cress). Cult. orn. fl. Araceae (EP.; Aroideae BH.). Monocots. (Spathiflorae). 105 gen., 1000 sp , trop. and temp. (92% trop ). Many types of veg. habit— herbs large and small, with aerial stems, tubers or rhiz., climbing shrubs, climbing epiph., marsh pi., one water pi. (Pistia) &c. In a few Pothoideae the stem is monopodial, but in most A. it is sym- podial. Each joint of the sympodium begins as a rule with one or more scale 1. before bearing fol. 1. Accessory (collateral) buds often found in the leaf axils. Sometimes, as in Anthurium, Philodendron &c., the axillary shoot is ' adnate ' to the main axis for some distance (cf. Solanaceae, Zostera &c. ). The buds usually appear in the 1. axils, but often get pushed to one side, and sometimes (e.g. Pothos) break through the leaf-bases as in Equisetum. L. of many types. Pinnately and palmately divided I. are frequent, but development not like that of such- leaves in Dicots. Holes are ARACEAE 49 present in the 1. of Monstera. See Monstera, Rhaphidophora, Philo- dendron, Helicodiceros, Dracontium, Zamioculcas, &c. Roots adv. and mostly formed above ground in the larger forms. Two types of aerial r. — climbing and absorbent. The former, like ivy, insensitive to gravity, show great negative heliotropism ; they cling closely to the support and force their way into the crevices. The latter, insensitive to light, respond markedly to gravity ; they grow down to the soil and enter it, branching out and taking up nourishment. The larger trop. A. show interesting stages in the development of epiphytism. The climbing forms grow to considerable size and form longer and longer aerial r. as they grow upwards. The original r. at the t.ase thus become of less and less importance and they often die away together with the lower end of the stem, so that the plant thus becomes an epiph. Of course, as it still obtains its water &c. from the soil, it is not an epiph. in the sense that e.g. many Orchids or Bromeliaceae are such, and it is evident that if this method of becoming epiph. were the only one found in the order, these plants could with no more justice be classed as true epiph. than the ivy which may often be seen in the ' bowls ' of pollard willows in Europe, and which has come there by climbing up the trunk and dying away below. It is found however that some sp. of Fhilodendron, Pathos, &c. are able to commence life as epiphytes. The fleshy fr. is eaten by birds and the seed dropped on a lofty branch. The seedling forms clasping r. and dangling aerial r. which grow steadily down to the soil, even if it be 100 feet or more away. It is hardly possible to suppose that these true epiph. sp. have been evolved in any other way than from former climbing sp. Lastly, some sp. of Anthurium &c. are true epiph. without connection with the soil (e.g. A. Hiigelii Schott = A. Hookcri Kunth.) ; they have clasping r. and also absorbent r. which ramify amongst the humus collected by the pi. itself. The aerial r. of some A. possess a velamen like Orchids. The 1. of Philodenaron cannifolium Schott have swollen petioles full of large intercellular spaces lined with mucilage. When rain falls these fill with water and act as reservoirs. [Cf. Schimper's Epiph. Veg. A merikas. ] Fls. without br., usu. massed together on a cylindrical spadix enclosed in a large spathe ; the spadix usu. terminates a joint of the sympodium (the 'continuation' bud is generally in the axil of the 1. next but one before the spathe), so that there is only one formed each year. Fl. $ or monoec. (dioec. in Arisaema), with or without P. Sta. typically 6 but usually fewer (down to i), often united into a synandrium (e.g. Colocasia, Spathicarpa) ; in Ariopsis the synandria again united to one another. Stds. often present, and these also may be fused into a synandrodium as in Colocasia. G with much variety of structure ; frequently reduced to i cpl. Berry. Outer integument of seed often fleshy. Endosperm or none. Fls. usu. protog. (even when monoec.). In many gen. (incl. most Eur.) the smell is disagreeable an 1 attracts carrion flies as pollen carriers (see. Arum, Dracunculus, Helicodiceros, &c. ). Many A. contain .latex, which is usually poisonous but is dispelled W. A 50 ARACEAE !>y heat. The rhizomes of many sp. contain much starch and are used as food (Caladium, Colocasia, Arum, &c.). Classification and chief genera (after Engler) : The grouping of the A. is very difficult and account has to be taken of histological as well as external characters. I. POTHOIDEAE (land pi. ; no latex or raphides ; 1. 2-ranked or spiral ; lat. veins of 2nd and 3rd order netted ; fls. usu. 5 ; ov. anatr. or amphitr.): Pothos, Anthurium, Acorus. II. MONSTEROIDEAE (land pi.; no latex; raphides; lat. veins of 3rd, 4th, and sometimes 2nd orders netted ; fl. 5, usu. naked ; ov. anatr. or amphitr.) : Rhaphidophora, Monstera, Spathiphyllum, Epipremnum. III. CALLO1DEAE (land or marsh pi.; latex; fl. usu. ?; ov. anatr. or orthotr. ; 1. never sagittate, usu. net-veined) : Symplocarpus, Calla. IV. LAS1OIDEAE (land or marsh pi.; latex; fl. $ or . BH.). Dicots. (Archichl. Aristolochiales). 5 gen., 200 sp., trop. and warm temp., except Austr. Herbs or shrubs, the latter usu. twining lianes. L. alt., stalked, often cordate, usu simple, exstip. Fl. $, epig., reg. or -|-. P usu. (3), petaloid; A 6 — 36, Iree, or united with the style into a gynostemium (cf. Asclepiads, Orchids, &c.). G 4— 6-loc. ; ov. oo in each loc., anatr., horiz. or pend. Caps. Embryo small in rich endosp. The A. are difficult to place in the system. They have been put near Dioscoreaceae, though not monocot. BH. place them in Multiovulatae Terrestres. Chief genera: Asarum, Aristolochia. ARTANEMA 55 Aristolochiales. The isth order of Dicot. Archichl. Aristotelia L'Herit. Elaeocarpaceae. 10 S. temp. Arjona Coinm. ex Cav. Santalaceae. 9 temp. S. Am. Armed, thorny. Armeniaca Tourri. ex Mill. =Prunus Tourn. (Rosac.). Armeria Linn. Plumbaginaceae. 60 N. temp, and andine. A.vulgaris Willd. (tlirift, sea pink) common on the coast of Brit, and in high mountain regions of Scotland, a fairly frequent phenomenon, due perhaps to similarity of conditions. Primary root perennial ; each year's shoot dies down all but a short piece, on which the following year's shoot arises as an axillary branch. Infl. a capitulum of cincinni, surrounded by a whorl of bracts, the outer forming a sheath round the top of the peduncle. After fert the K becomes a membranous funnel- like organ aiding seed-distribution by wind. Armoracia Gaertn. = Cochlearia L. and Nasturtium R. Br. (Cruc.). Arnatto, arnotto, Bixa Orellana L. Arnebia Forsk. Boraginaceae (iv 4). 12 Medit., trop. Afr., Himal. Some have black spots on the C, which fade as it grows older (see fam., and cf. Diervilla, Fumaria, &c.). Arnica Rupp. ex L. Compositae (8). 50 N. temp, and arctic. Tincture of arnica is prepared from all parts of the pi. Arnicastrum Greenman. Compositae (6). i Mex. Arnocrinura Endl. et Lehm. Liliaceae (in). 3 S.W. Austr. Arnoseris Gaertn. Compositae (13). i, A. pusilla Gaertn., Eur. (incl. Brit.). The bases of the invol. brs. enclose the ripe fr. (cf. Rhagadiolus). Arnottia A Rich. Orchidaceae (n. i). 2 Mauritius. Aroideae (/>//.) = Araceae. Arodendron Werth. (Typhonodorum p.p.). Araceae (v). i Zanzibar. Aronia Pers. = Amelanchier Lindl. (Rosac.). Aronicum Neck. = Doronicum L. p.p. (Compos.). Arpophyllum Llave et Lex. Orchidaceae (n. 6). 6 C. Am., W.I. Arrabidaea DC. Bignoniaceae (i). 100 S. Am. Arracacia Bancroft. "Umbelliferae (ill. 4). 45 Peru to Mex. A.xantho- rhiza Bauer and others cult. ed. tuberous r. Arrack, cf. Borassus, Cocos, &c. Arrhenatherum Beauv. Gramineae (9). 3 Eur., Medit. A. avenacetim Beauv. (false oat-grass, French rye-grass), Brit. Arrhostoxylmn Mart. = Ruellia L. p.p. (Acanth.)- Arrow-grass, Triglochin ; -head, Sagittaria sagittifolia L. Arrowroot, a pure starch obtained from various pi.; Bermuda-, W. Indian-, Maranta arundinacea L.; Brazilian-, Afaiiihot utilissima Pohl. ; E. Indian-, Curcuma angiislifolia Roxb., Tacca pinnatijida Forst. &c.; Portland-, Arum niaculatum L. Cf. Canna, Zea, &c. Arrowsmithia DC. Compositae (4). i S. Afr. Arsenococcus Small (Vacdnium p.p.)- Eric. (in. i). i Atl. U.S. Artabotrys R. Br. Anonaceae (4). 23 palaeotrop. Some cult, for sweetly scented fl. and ed. fr. They usually climb by aid of recurved hooks, which are modified infl. axes, and thicken and lignify when they clasp. Artanema D. Don. Scrophulariaceae (li. 6). 4 Indomal. 56 ARTANTHE Artanthe Miq. = Piper L. (Piper.). Artedia L. Umbelliferae (in. 8). i W. As. Artemisia Linn. Compositae (7). 260 .*, common on the arid soil of the western U.S., the Steppes, &c. 4 Brit, (wormwood). A. tri- dentata Nutt. and others form the ± halophytic ' sage-brush ' of the S.W. U.S. Fl. -heads small, inconspic., and w/«a?fert. (cf. Poterium, Rheum and Rumex, Plantago, Thalictrum, &c.). In A. vulgaris L. the marginal florets ?, the rest g. Head pend. ; the anther-tube projects beyond the C so that the dry powdery pollen is exposed to the wind. On the tips of the anthers are long bristles which together form a temporary pollen-holder. Afterwards the style emerges and the large hairy stigmas spread out. An interesting case of reacqui- sition of a character not found in most higher flowering pi. A. Abro- tanum L. (old man, southernwood), and others, cult. orn. 11. The flavouring matter of absinthe is derived from wormwood. Artemisiopsis Sp. Moore. Compositae (4). i Nyassaland. Arthraerua Schinz. Amarantaceae (2). i S.W. Afr. Arthratherum Beauv. = Aristida L. p.p. (Gramin.). Arthraxon Beauv. Gramineae (2). 15 palaeotrop., E. As. Arthrocarpum Balf. f. (Ormocarpum EP.). Legumin. (in. 7). i So- cotra. Arthroclianthus Baill. Leguminosae (ill. 7). 6 New Cal. Arthrocnemum Moq. Chenopodiaceae (A). 8 coasts |-x- . Arthrolobium Reichb. =Ornithopus L. and Scorpiurus L. Arthrophyllum Blume. Araliaceae (i). 10 Indomal. Arthrophytum Schrenk. Chenopodiaceae (B). i W. As. Arthropodium R. Br. Lili. (in). 8 Austr., N.Z., New Cal. Cult, orn. fl. Arthropogon Nees. Gramineae (4). 2 Brazil, Cuba. Arthropteris J. Sm. Polypodiaceae. 4 trop. and subtrop. Arthrosolen C. A. Mey. (Gnidia L. p.p. EP.). Thymel. 12 trop. and S. Afr. Arthrostema Ruiz et Pav. Melastomac. (i). 12 Cuba, W. trop. Am. Arthrostylidium Rupr. Gramin. (13). 16 trop. Am., W.I. Climbing. Arthrostylis R. Br. Cyperaceae (n). i Austr. Artfcrotaxis Endl. = Athrotaxis D. Don (Conif.). Arthrotriamnus Klotzsch et Garke= Euphorbia L. p.p. (Euph.). Artichoke, Cynara Scolynms L. ; Jerusalem-, Helianthas tuberosus. Articulate (1.), cut oft' by an absciss layer. Articulated, jointed. Artillery plant, Pilea. Artocarpus Forst. Moraceae (11). 60 Indomal., China. Many show good bud-protection by slips. A. laciniata Hort. has large drip-tips (Acer, Ficus). Fls. monoec., the 3 in pseudo-catkins, the ? in pseudo- heads. A multiple fr. is formed, the achenes being surrounded by the fleshy P and the common receptacle also becoming fleshy. The fr. contains much starch &c. and is a valuable food-stuff. Several sp. are cult, all over the trop., e.g. A. incisa L. (bread-fruit) and A. inte- grifolia L. (jak). The flesh has somewhat the texture of bread and is often roasted. The best cult, forms (cf. pear, banana, &c.) produce no seeds. The jak and others are cauliflural. Timber useful. ASCLEPIADACEAE 57 Artrolobium Desv. = CoronilIa L., Ornithopus L., &c. (Legum.). Arum (Tourn.) Linn. Araceae (vn). 15 Eur., Medit. A.maculatum'L. (cuckoo-pint, wake-robin, lords and ladies, Brit.) is a perenn. tuberous pi. with monoec. fls. ; V fls. at base of spadix (each of i cpl., naked) and S above (each of 2 — 4 sta.), and above these again rudimentary cf fls. repres. by hairs which project and close the mouth of the spathe. The foetid smell attracts flies, which enter the spathe, find the stigmas ripe, and are kept prisoners till the pollen is shed ; then the hairs wither and escape is possible (cf. Aristolochia). Fr. a berry. The starch of the tubers was formerly used as food under the name Port- land arrowroot, but it is difficult to get rid of the poisonous juices accompanying it. Other sp. are similarly used in Eur. Arum lily, Richardia africana Kunth. Aruncus Aclans. Rosaceae (I. i). 6 N. temp. Arundina Blume. Orchidaceae (n. 4). 8 S.E. As., Malay Arch. Arundinaceous, reed-like. Arundinaria Michx. Gramineae (13). 45 As., Am. Bamboos (g-v-)- Arundinella Raddi. Gramineae (4). 45 trop. Arundo Tourn. Gramineae (10). 12 trop. and temp. A. Phragmites L. (Brit.) = P. communis. The stems of A. Donax L. are used for sticks, fishing-rods, &c. Arvensis (Lat.), of arable land. Arytera Blume (Ratonia DC. p.p. BH.}. Sapind. (i). 20 E. As., Austr. Asaemia Harv. Compositae (7). i S. Afr. Asafoetida, Ferula Narthex Boiss., F. Assafoetida L., &c. Asagraea Baill. (Dalea L. p p. BH. ). Leguminosae (ill. 6). 2 N. Am. Asarabacca, Asarum europaeum L. Asarca Poepp. ex Lindl. = Chloraea Lindl. (Orchid.). Asarum (Tourn.) L. Aristolochiaceae. 30 N. temp. A. europaeum L. (asarabacca), an escape in Brit, (formerly medic.). Rhiz. below ground and creeping shoots above ; the latter are sympodial, each annual joint bearing several scale 1. below, then two green 1. and a terminal fl. Fl. reg. ; P (3), sometimes with 3 small teeth between the segments (perhaps remnants of a former inner whorl); A 12; G (6). The dark-brown, resinously scented fl. is visited by flies, and is very protog. ; when the stigmas are ripe the sta. are all bent away, but later on they move up to the centre and dehisce extr. The P lobes are bent in at first towards the centre of the fl. and form a sort of prison of it, but afterwards gradually straighten. Ascarina Forst. Chloranthaceae. 6 Polynesia. Ascending (aestivation), cf. Aest. ; (ovule), sloping upwards ; (stem), do. -ascens (Lat. suffix), tending towards. Aschamia Salisb. = Hippeastrum Herb. p.p. (Amaryllid.). Aschenbornia Schauer. Compositae (2). i Mex. Aschersoniodoxa Gilg-et Muschler (Draba p.p.). Cruc. (4). i Ecuador. Asciadium Griseb. Umbelliferae (inc. sed.). i Cuba. Ascidium, a pitcher. Asclepiadaceae (EP. BH.}. Dicots. (Sympet. Contortae). 220 gen., 1700 sp., mostly trop. (esp. Afr.), but a few temp. In veg. habit like Apocynaceae ; some perenn. herbs, but most are climbing shrubs 5 8 ASCLEPIADA CEA E or Hanes, with simple, entire, opp., exstip. 1. Latex is present. Many, esp. S. Afr. sp., xero. ; some, e.g. Periploca, with much reduced 1., others, e.g. Hoya. and still more, Stapelia, with fleshy stems. Epiphytes also occur ; cf. esp. Dischidia. Infl. usu. of many fls. and cymose or racemose (raceme or umbel). In the former case it is dich., but as in Caryophyll. the one branch tends to outgrow the other, and a monoch. (cincinnus) may arise in later branchings. When infl. axillary, there is usu. only one at each node; in the axil of the other 1. there is a veg. shoot, or nothing. In some gen. (cf. Asclepias) the infl. is extra-axillary. Fl. 5, reg , j-merous, usu. small. K 5, quincuncial, the odd sep. post.; C(5) usu. rotate or campanulate, with conv. or valvate aesti- vation. The essential organs (5 sta., 2 cpls. ) are complex. The sta. and style are usu. united to form a gynosteginm. The cpls. are free below as in Apocyn., but united at the tip with a common style ; ov. sup. The head of the- style is large and variously shaped, and the stigmatic surface is usually upon the edge or under side of it. To its margin are united the anthers of the 5 epipet. sta. ; the filaments of these are short or non-existent. The pollen in § i is united merely in tetrads, in the higher group, comprising the bulk of the order, into pollinia, as in Orchids. Usu. each anther contains two. In this group also there are curtain-like projections at the sides of the anthers, leaving a narrow slit between each pair of anthers. The pollen is removed from the anthers by a curious mechanism — the translator. This differs in the two suborders and so also does the fert. method. The translator always stands between two anthers and serves to carry away half the pollen from each of them. In the Periplocoideae it is a spoon- or funnel-like body with a sticky disc at the narrow end. Into it is shed the pollen from the two half anthers next to it, and as the sticky disc projects outwards in the male stage of the flower an insect will be likely to get it attached to its head, and carry it about like the pollinia of an orchid. In visiting a second flower the pollen may be placed on the stigmalic surface. In the Cynanchou/eae, on the other hand, there are pollinia, and the trans- lator has a different structure. It forms an inverted \-shaped organ, the foot of the Y being formed by the adhesive body (corpiisculuni] ; from this diverge the threads (retinaculn) which are attached to the pollinia, one in each anther. An insect in obtaining honey catches its leg in the slit between the anthers, and in drawing it up removes the pair of pollinia. The threads as they dry contract on the inner side till the pollinia meet, thus closely clasping the insect's leg. In drawing the leg through a similar slit in another fl. the pollinia catch on the stigmatic under-surface of the stylar head. (Cf. Apocynmn, which shows an approach to this mech.) The backs of the anthers as a rule bear appendages (ciiculli) forming a corona. In some cases it springs from the C. It may consist of small teeth, or be more complex, as in Asclepias and Ceropegia, and often takes up the functions of secreting and storing the honey. G (2), -z-loc., with oo anatr. ov. pend. from the ventral plac. Fr. a pair of follicles; seeds usu. crowned by a tuft of hairs for wind- ASPARAGOPSIS 59 carriage. Endosperm slight, cartilaginous. Some give useful fibres. Closely related to Apocyn., the only absolute distinction being the presence of translators in Ascl. ; otherwise the two sub-orders of each form a corresponding series, and the lower one in each is almost as nearly related to the corresponding one in the other order as to the higher group in its own order. Placed in Gentianales by BH. Classification and chief genera (after K. Schumann) : I. PERIPLOCOIDEAE (pollen in tetrads; translator spoon- like), i. Periploceae : Streptocaulon, Periploca. II. CYNANCHOlDEAE(vQ\\\m&\ corpusculum, &c.). 1. Asclepiadeae (pollinia pendulous on threads): Asclepias, Calotropis, Cynanchum. 2. St'canioneae (pollinia erect or horizontal, 4 in each anther) : Secamone (only genus). 3. Tylophoreae (do but 2 in each, erect): Ceropegia, Stapelia, Stephanotis, Hoya. 4. Gonolobeae (do. but 2 in each, horiz.) : Gonolobus. Asclepias L. Asclepiad. (n. i). ' 160 Am., Afr., esp. U.S. (silk-weeds). Herbs with umbellate infls. which spring from the stem between the petioles of the opp. 1. (cf. Cuphea), or above or below this. Two explanations exist, but which is right the evidence available does not show. Either the inn. is axillary to the 1. below it and is 'adnate ' to the stem (cf. Cuphea), or it is the termination of a shoot, and the stem is really a sympodiuin. The cuculli of the anthers form little pockets, into which honey is poured by the horn-like nectaries that project from them. Insects walking over the fl. and sipping honey frequently slip their legs down the sides of the gynostegium, and in drawing them up catch in the slit between two anthers and remove the pollinia (cf. family). The process may be watched on A. Cormtti Decne. in gardens. Asclepiodora A. Gray (Asclepias p.p. EP.). Asclep. (n. i). i N. Am. Ascocentrum Schlechter. Orchidaceae (n. 20). 4 New Guinea. Ascochilus Ridl. Orchidaceae (n. 20). 4 Malay Penins. Ascog-lossum Schlechter. Orchidaceae (n. 17). i New Guinea. Ascolepis Nees. Cyperaceae (i). 10 warm Afr., Am. Ascotainia Ridl. (Tainia p.p.). Orchid, (n. 7). 2 Malay Penins. Ascyrum L. Guttiferae (n). 5 N. Am., W. I., Himalaya. Asemnantha Hook. f. Rubiaceae (n. 3). i Yucatan. Aseptate, without partitions. Asexual, sexless ; -generation, Pteridophyta. Ash, Fraxinus excelsior L. ; mountain-, Pyrus Aitcuparia Ehrh. ; prickly-, Zantlwxyluin fraxineiun Willd. ; -pumpkin, fienincasa cerifera Savi. Asimina Adans. Anonaceae (2). 6 E. U.S. A. triloba Dun. (papaw) has ed. fr. Askidiosperma Steud. Restionaceae. i S. Afr. Aspalathus L. Leguminosae (in. 3). 160 S. Afr. Many are xero. with a heath-like habit. Asparagopsis Kunth — Asparagus Tourn. p.p. (Lili.). 60 ASPARAGUS Asparagus Tourn. (inch Myrsiphyllum Willd.). Liliaceae (vn). 1 20 |-*, mostly in dry places. Rhiz. with aerial shoots; 1. reduced to scales with linear green shoots in axils, usu. stem in tufts. These are small condensed cymes, of the type i shown in the diagram (figs, represent the branches of sue- 2 2 cessive orders). The number of shoots that develope 3 3 varies. In the infl. the same construction holds, the shoots 4 4 2, 2 bearing the fls. In the sub-genus M. there are flat leaf phylloclades (cf. Ruscus). Fr. a berry. A. officinalis L. cult., the young shoots being eaten. Aspasia Lindl. Ochidaceae (n. 19). 8 trop. Am. Aspen, Populus treniula L. Asper (Lat.), rough; -ifolius (Lat), rough-leaved. Asperella Humb. Gramineae (12). 2 As. (BH.), 4 N. Am , N. As., N.Z. (EP.). Asperifoliae = Boraginaceae. Aspermous, seedless. Asperugo (Tourn.) L. Boraginaceae (iv. 2). i Eur., As. Asperula L. Rubiaceae (n. n). 80 Eur., As., Austr., esp. Medit. A. odorata L. (woodruff) and A. cynanchica L. (squinancy-wort), Brit. Fls. homogamous ; fr. of woodruff hooked. Asphodel, Asphodelus; bog-, Nartheciitm ossifragiun Huds.; Scottish-, Tofiddia palnstris Huds. Asphodeline Rchb. Liliaceae (ill). 15 Medit. Asphodelus (Tourn.) L. Liliaceae (in). 12 Medit. (asphodel). L. isobil.; fls. protog. Aspicarpa Rich. Malpighiaceae (i). 5 Texas to Argentina. Aspidandra Hassk. Euphorbiaceae (inc. sed.). i Java. Aspidistra Ker-Gawl. Liliaceae (vn). 5 E. As. The large flat style forms a lid to the cavity made by the 8 P-leaves. Cult. orn. fl. Aspidium Sw. Synonymy: A. aemithtm Sw., cristatum Sw., dilatatum Willd., Filix-mas Sw., Oreopteris Sw., rigidum Sw., spinulosum Sw., Thelypteris Sw. = Dryopteris (same spec, names); A. Filix-foemina Sw. = Athyrium F.-f. Polypodiaceae, 200 cosmop. A. nculeatiitn Sw. (prickly shield-fern) and A. Lone hit is Sw. (holly-fern) in Brit. Aspidixia Van Tieghem ( Viscum p.p.). Loranthac. (n). 10 palaeotrop. Aspidocarya Hook. f. et Thorns. Menispermaceae. 5 Indomal. Aspidopterys A. Juss. Malpighiaceae (i). 20 trop. As. Aspidosperma Mart, et Zucc. Apocynac. (i. 3). 50 trop. and S. Am., W.I. Wood useful; bark (quebracha) used for tanning. Aspilia Thou. Compositae (5). 90 Brazil to Madagascar. Aspiliopsis Greenman. Compositae (5). i Mex. Aspleniopsis Mett. Polypodiaceae. i Melanesia. Asplenium L. Polypodiaceae. 540 cosmop.; n in Brit., including A. Filix-foemina Bernh. (lady-fern), A. Rnta-Diuraria L. (wall- spleenwort), A. Trichomanes L. (spleenwort) and A. Adiantum- nigruin (black spleenwort). The var. clarissima of the lady-fern shows apogamy (see Filicineae, Lepto.). A. bulbiferum Forst. and other sp. are 'viviparous,' producing young plants on their leaves by vegetative budding (not to be confounded with apospory). A. Nidus ASTHENO CHL OA 6 1 L. (the bird's nest fern) is an interesting epiphyte of the Old World tropics. It bears a rosette of leaves forming a nest in which humus collects; the roots ramify in this and obtain food and water. A. rhi- zophylhun Kunze is the walking fern, so called because the leaf-tips when they touch the soil bud into new individuals, and thus the plant spreads to some distance. Asprella Schreb. = Leersia Sw. (Gramin.). Assai palm, Euterpe edulis Mart. Assegai wood, Curdsia faginea Ait. Associations, plant, the grouping of pi. which occupies any definite uniform area and kind of land. Assurgent, ascending. Astartea DC. (Baeckea L. p.p.)- Myrtaceae (n. i). 5 Austr. Astelia Banks et Soland. Liliaceae (vi). 15 Polynes. Dioec. Cult, orn. fl. Astelma R. Br. — Hdichrysum, Helipterum (Compos.). Astelma Schlecht. Asclepiadaceae"(n. 3). i New Guinea. Astemma Less. Compositae (5). i Ecuador. Astemon Regel. Labiatae (inc. sed.). i S. Am. Astephania Oliv. Compositae (4). 2 E. Afr. Astephanocarpa Baker. Compositae (4). i Madag. Astephanus R. Br. Asclepiad. (n. i). 12 Madag., trop. Afr., warm Am. Aster Tourn. ex L. Compositae (3). 400 Am., As., Afr., Eur. ; 2 on Brit, coast (Michaelmas daisy), somewhat fleshy halophytes. Style sta. typical of § 3. The China aster of gardens (A. chinensis L.) is a Callistephus. Many cult. orn. fl. Aster, China, Callistephus hortensis Cass. Asteracantha Nees (Hygrophila BH.}. Acanth. (iv. A), i palaeotrop. Asteraceae = Compositae. Asterales (BH.}. The 2nd cohort of Dicotyledons (Gamopet.). Asteranthe Engl. et Diels (Asteranthopsis). Anon. (i). i Zanzibar. Asteranthera Hanst. Gesneriaceae (i). r Chili. Asteranthopsis O. Ktze. Anonaceae (i). i Zanzibar. Asteranthus Desf. Lecythidaceae. i Venezuela, N. Brazil. Asteriastigma Bedd. (Hydnocarpus p.p. EP.}. Flacourt. (3). r India. Asteriscium Cham, et Schlecht. Umbelliferae (I. 2). 27 Mex., S. Am. Asteriscus Moench = Odontospermum Neck. (Comp.). Asterocephalus (Vaill.) Arlans. =Scabiosa L. p.p. (Dipsac.). Asterochaete Nees. Cyperaceae (n). 4 Masc., S. Afr. Asterochlaena Garcke = Pavonia Cav. (Malv.). Asterogyne H. Wendl. Palmaceae (iv. i). 2 C. Am. Asteroid, star-shaped. Asterolasia F. Muell. Rutaceae (i). 7 Austr. Asterolinon Hoffmannsegg et Link. Primulaceae. 2 Medit., Abyss. Asteromaea Blume (Bottonia BH.}. Compositae (3). 3 E. As. Asteropeia Thou. Theaceae (Samyd. BH.}. 5 Madag. Asterophorum Sprague. Tiliaceae. i Ecuador. Asteropsis Less. (Podocoma Cass. BH.}. Compositae (3). i S. Brazil. Asterostemma Decne. Asclepiadaceae (n. 3). i Java. Asterostigma Fisch. et Mey. (Staurostigma £H.) . Arac. (vn). 5 Brazil. Asthenochloa Buese. Gramineae (inc. sed.). i Java. 62 ASTIANTHUS Astianthus D. Don (Tecoma Juss. p.p. BH.}. Bignon. (2). i C. Am., Mex. Astichous, not in rows. Astilbe Buch.-Ham. Saxifragaceae (i). 20 As., N. Am. Astiria Lindl. Sterculiaceae. i Masc. Astoma DC. Umbelliferae (in. 4). i Palestine. Astragalus Tourn. ex L. Leguminosae (in. 6). 1600 cosmop. exc. Austr. ; 3 Brit, (milk-vetch). Usu. on steppes, prairies, &c. and ± xero. often thorny ; the thorns commonly form by the stiffening of the petiole or midrib of the 1. when the blade falls off. A. gummifer Labill. and others yield gum-tragacanth, obtained by wounding the stem ; the gum exudes and hardens. Astrantia ( lourn.) L. Umbelliferae (n. i). 5 Eur., As. Astrebla F. Muell. Gramineae (n). 3 Austr. Astrephia Dufresne. Valerianaceae. 4 Chili. Astrocalyx Merrill. Melastomaceae (n). i Phil. Is. Astrocarpus Neck. Resedaceae. i S.W. Eur. Astrocaryum G. F. W. Mey. Palmae (iv. 2). 40 trop. Am. Several yield fibre and oil, or are cult, as decorative. Astrocasia Robinson et Millspaugh. Euphorb. (A. i. i). i C. Am. Astrochlaena Hallier f. Convolvulaceae (i). 20 Afr. Astrococcus Benth. Euphorbiaceae (A. n. 2). 2 Brazil. Astrodaucus Drude (Dauais p.p.). Umbelliferae (in. 2). 4 Medit. Astroloma R. Br. (Styphelia p.p. EP.). Epacridaceae. 25 Austr. Astronia Noronha. Melastomaceae (n). 30 Malay Arch., Polynes. Astronium Jacq. Anacardiaceae (3). 10 S. Am. Hard wood. Astrophiolate, with no strophiole. Astrophyllum Torr. et Gray. Rutaceae (i). i Mex., S.W. U.S. Astrostemma Benth. Asclepiadaceae (n. i). i Borneo. Astrothalamus C. B. Robinson. Urticaceae (3). i Phil. Is. Astrotheca Miers (Clusia L. p.p. BH.). Guttiferae (v). i Brazil. Astrotricha DC. Araliaceae (i). 6 Austr. Astydamia DC. Umbelliferae (in. 6). i Canaries. Asymmetrical (fl.), with no plane of symmetry, Canna, Marantaceae, Valerianaceae ; (1.), divided into unequal parts by midrib, Begonia, Ulmaceae. Asystasia Blume. Acanthaceae (iv. B). 30 palaeotrop. Asystasiella Lindau. Acanthaceae (iv. B). 3 trop. As , Afr. Ataenidia Gagnep. Marantaceae. i Gaboon. Atalantia Correa, Rutaceae (v). 18 trop. As., China, Austr. Atalaya Blume. Sapindaceae (i). 5 Austr., Malay Arch. Atamasco Raf. =Zephyranthes Herb. (BH.) = Amaryllis L. Atamisquea Miers. Capparidaceae (n). i Chili, Arg. Atamosco, Zephyranthes texana Herb. Atavism, cf. Reversion to type. Atelandra Bello (Meliosma Blume EP.}. Sabiaceae [Myrsinaceae (BH.)]. 2 Porto Rico. Atelanthera Hook. f. et Thorns. Cruciferae (4). i W. Tibet. Ateleia Moc. et Sesse. Leguminosae (in. i). 4 trop. Am., W.I. Atelophragma Rydb. (Astragalus p.p.). Legumin. (in. 6). 6 N. Am. Ater (Lat), black. -ATUS 63 Ateramnus P. Br. Inc. sed. Jamaica. Nomen delendum. Athamanta L. Umbelliferae (ill. 5). 10 Medit. Athanasia L. Compositae (7). 50 S. and trop. Afr., Madag. Athenaea Sendtn. vSolanaceae (2). 15 trop. Am. Atherandra Decne. Asclepiadaceae (i). 3 Malaya. Atheranthera Mast. Passifloraceae. i Angola. Atherolepsis Hook. f. Asclepiadaceae (i). 2 Burma. Atheropogon Muhl. ex Willd. = Bouteloua Lag. p.p. (Gramin.). Atlierosperma Labill. Monimiaceae. 2 Victoria, Tasmania. The strongly scented bark is sometimes used as a tea. Atherostemon Blume. Asclepiadaceae (i). i Burma, Malaya. Athrixia Ker-Gawl. Compositae (4). 20 Austr., Afr. Athroisma DC. Compositae (4). i E. Indomal. Athroostachys Benth. Gramineae (13). i Brazil. Climbing. Athrotaxis D. Don. Coniferae (Pinaceae ; see C. for gen. char.). 4 Tasmania, Austr. Athyana Radlk. Sapindaceae (i). i Arg., Paraguay. Athyrium Roth. Polypodiaceae. 120 cosmop. A. Filix-femina Roth, (lady fern) Brit. Athyrocarpus Schlecht. (Phaeospherion EP.}. Commel. 3 trop. Am. Athysanus Greene. Crucifcrae (4). 2 Calif. Atlantic cedar, Cedrtts atlantica Manetti. Atomostigma O. Ktze. Rosaceae (vi). i Matto Grosso. Atractocarpa Franchet. Gramineae (13). i Congo. Atractocarpus Schlechter et Krause. Rubiaceae (I. 8). i New Cal. Atractogyne Pierre. Rubiaceae (i. 8). i trop. Afr. Atractylis L. Composiiae (u). 20 Medit. to Japan. Atragene L. = Clematis L. (Ranunc.). Atrapnaxis L. Polygonaceae (n. i). 18 C. As., N. Afr., Greece. Atratus (Lat.), blackened. Atrichoseris A. Gray. Compositae (13). i Calif. Atriplex (Tourn.) L. Chenopodiaceae (A). 180 temp, and subtrop.; 6 Brit, (orache). Fls. unisexual or polyg., naked or with P. Atro- (Lat. pref.), black ; -purpureus, dark purple ; -virens, dark green. Atropa L. Solanaceae (2). 2 Eur., Medit., As. A. Belladonna L. (deadly nightshade) contains the alkaloid atropin, the basis of the drug belladonna used in medicine. Atropanthe Pascher (Sfopolia p.p.). Solanaceae (2). i China. Atropin, Atropa Belladonna L. Atropis Rupr. (Glyceria R Br. BH}. Gramineae (10). 30 temp. Atropous (ovule), in a line with funicle. Atroxima Stapf Polygalaceae. 4 trop. Afr. Atrutegia Bedd. (Goniothalamus Bl. hP.}. Anonac. (2). i S. India. Attalea H. B. et K. Palmae (iv. 2). 30 S. Arn., W. L, tiop. Afr. A. fimifera Mart (Brazil) yields Bahia Piassaba fibre (Kew Bull. 1889, p. 237). A. Cohune Mart. (Honduras) yields the ivory-like Cohune nuts. Attar of roses, Rosa, Pelargonium. Attenuate, tapering. -atus (Lat. suffix), provided with. 64 ATYLOSIA Atylosia Wight et Am. (Cantharospermiim W. et A. pp. EP.). Legu- minosae (in. to). 20 trop. As., Austr., Madag., Mascarenes. Aubergine, Solatium Melongena L. Aubletella Pierre (Chrysophytlnm L. p.p.). Sapotac. (i). i Guiana. Aubrietia Adans. Cruciferae (4). 15 Medit. Mis. Aubrya Baill. (Saccogtottis EP.). Humiriac. 10 S. Am., trop. Afr. Aujoumea Pierre. Burseraceae. i Gaboon. Yields resin AucubaThunb. Cornaceae. 3 Himal. to Japan. A.japonica1\mxk>. (Japan laurel) cult. orn. shrub ; it is dioecious. Aucubaephyllum Ahlburg (Grumilea p.p. EP.). Cornaceae. i Japan. Audibertia Benth. (Kamona Greene). Labiat. (vi). 10 N. Am. AudibertieUa Briq. Labiatae (vi). 10 Calif. Audouinla Brongn. Bruniaceae. i S. Afr. Augea Thunb. Zygophyllaceae. i S. Afr. Augia Lour. Inc. sed. i China. Augusta Leandr. = Stifftia Mikan. (Compos.). Augusta Pohl (Ucriana Spreng.). Rubiaceae (i. 3). i E. Brazil. Aulacocalyx Huok. f. Rubiaceae (11. i). 4 trop. Afr. Aulacocarpus Berg. Myrtaceae (i). 2 Brazil. Aulacodiscus Hook. f. (Pleiocarpidid). Rubiaceae (I. 7). i Malay Pen. Aulacolepis Hackel. Gramineae (8). i Japan. Aulacorhynchus Nees. Cyperaceae (inc. sed.). i S. Afr. Aulax Berg. Proteaceae (i). 2 Mts. S. Afr. Aulaya Harv. = Harveya Hook. (Scrophular.). Aulisconema Hua (Disporopsis $.?.). Liliaceae (vn). 2 China. Aulojusticia Lindau. Acanthaceae (iv B.). i S. Afr. Aulomyrcia Berg. = Myrcia DC. p.p. (Myrtac.). Aulospermum Coulter et Rose (Cymopterus p.p.). Umb. (ill. 6). 12 N. Am. Aulostephanus Schlechter. Asclepiad. (n. 3). i Natal. Aulostylis Schlechter. Orchidaceae (n. 9). i New Guinea. Aulotandra Gagnep. Zingiber. (i). 2 Madag., trop. Afr. Aurantium Tourn. ex Mill. = Citrus L. (Rutac.). Aureus (Lat.), golden. Auricled (1.), with two lobes overlapping stem. Auricula, Primula Auricula L. Auricula Tourn. ex Spach= Primula Tourn. Aurila Noronha. Inc. sed. Nomen. Austere, astringent. Australian black-wood, Acacia melanoxylon R. Br. ; chestnut, Casta- nospermum aitslrale A. Cunn.; -currant, Leucopogon\ -daisy, Vitta- dinia; -fuchsia, Correa; -heath, Epacris ; -honeysuckle, Banksia; red cedar, Cedrela australis F. Muell. Australina Gaudich. Urticaceae (5). 5 Austr., N.Z., S. Afr. Australis (Lat.), southern. Autochthonous, native. Autogamy, self-pollination, Capsella, Senecio, &c. Autrandra Pierre ex Prain (Erythrococca p.p.). Euph. (A. II. 2). i trop. Afr. Autrania C. Winckler et Barbey (Centaurea p.p. EP.). Comp. (tr). i Syria. AZALEA 65 Autumn crocus, Colchicum. Autunesia O. Hoffm. Compositae (i). i Angola. Auxemma Miers. Boraginaceae (i). 2 Brazil. Useful firewood. Auxopus Schlechter. Orchidaceae (II. 2). i Cameroons. Avellanita Phil. Euphorbiaceae (A. II. 3). i Chili. Avellinia Parl. Gramineae (10). 2 Medit. Avena L. Gramineae (9). 70 temp, and Mts. of trop.; 2 Brit. (oat-, grass). A. sativa L., the cult, oat, is perhaps derived from A.fatna L. It is cult, in Eur. to 69^° N. and forms the staple of the food of a large population. It occurs in two chief forms, the common oat with open spreading panicles, and the Tartarian oat with contracted one-sided panicles. [See De Candolle's Orig. of Cult. Pits. p. 373.] The 2-6-flowered spikelets form a loose panicle. The paleae are awned, the awn of the inf. palea being usually twisted and hygroscopic. In A. steriiis L. the awns cross, and when wetted try to uncurl and thus press on one another till a sort of explosion occurs jerking away the fruits. Avenastrum Jessm. (Avena L. p.p.). Gramineae (9). 10 S. Afr. Avens, Geitm. Averrhoa L. Oxalidaceae. 4 trop. As. (? ; long cult. ; origin uncertain). A. Bilitnbi L. (blimbing) and A. Carambola L. (carambola) cult, for fr., which is borne on the older stems (cauliflory). Averrlioidium Baill. Sapindaceae (11). 2 Brazil, Paraguay. Averse, turned back. Avicennia L. Verbenaceae (7). 3 trop., a constituent of the mangrove veg. (q.v.). The seeds germinate in the fr., and they have aerial r. projecting out of the mud like Sonneratia. Avocado, I'ersea gratissima Gaertn. f. Awl-wort, Subularia aquatica L. Awn, a thread-like organ on a fr., Anthoxanthum, Erodium, Gerania- ceae, Gramineae. Axanthes Blume=Urophyllum Wall. (Kubiac.). Axia Lour. (Boerhaavia L. p.p. EP.\. Nyctaginaceae (Valeriana- ceae? BH.}. i Cochinchina. Axial, axile, belonging to the axis ; -placentation, on the axis, cf. diagram of Guttiferae, Liliaceae, Axil, the upper angle between a 1. and the stem on which it is borne ; -lary, in the axil. Cf. Buds, Concrescence, &c. Axinaea Ruiz et Pav. Melastomaceae (i). 25 trop. Am. Axinandra Thw. (BH. incl. Naxiandra Krasser). Melastomaceae (ill)- (Lythraceae BH.} i Ceylon. Axiniphyllum Benth. Compositae (5). 2 Mex. Axis, the stem. Axonopus Beauv. = Paspalum L. (Gramin.). Axyris L. Chenopodiaceae (A). 6 Mid. and N. As. Ayapana, Eupatorium Ayapana Vent. Aydendron Nees. Lauraceae (n). 45 trop. Am. Ayenia Loefl. (Aniba Aubl.). Sterculiaceae. 25 trop. and subtrop. Am, Azadiracnta A. Juss. (Melia L. BH.) i Indomal. A. indica A. Juss. (nim) has astringent medicinal bark, and yields good timber. Azalea Gaertn. = Loiseleuria Desv. ; Azalea L. = Rhododendron L. W. C 66 AZALEASTRUM Azaleastnun Rydb. (Rhododendron p.p.)- Ericaceae (i. 2). i N. Am. Azara Ruiz et Pav. Flacourtiaceae. 20 Mcx., S. Am. Shrubs with alt. 1. ; one stip. is frequently almost as large as the 1. to which it belongs, giving the appearance of a pair of 1., not opp. Fl. apetalous ; outer sta. often without anthers. Azedarach Tourn. ex L. = Melia L. (Meliac.). Azeredia Arruda (Cochlospernmm p.p. EP.}. Cochlosperm. i Brazil. Azima Lam. Salvadoraceae. 3 S. Afr. to Phil. Is. In the axils are thorns (the 1. of an undeveloped shoot, cf. Cactaceae). Polypet. Azolla Lam. Salviniaceae. 4 trop. and subtrop. General structure like Salvinia. Two 1. are formed at each node, from (he dorsal half of a segment of the apical cell ; from the ventral half are formed roots and branches, but not at every node. The 1. are all alike ; each is bilobed and has a small cavity near the base, opening by a small pore, and inhabited by the Alga Anabaeita. The r. hang freely down in the water ; usually the root cap is thrown off after a time and the r. comes almost exactly to resemble the submerged 1. of Salvinia. The sporocarps are formed in pairs (4 in A. mlotica] on the ventral lobes of the first 1. of the branches. Each contains one sorus. The microspores are joined together into several masses in each sporangium by the hardened frothy mucilage (epispore, cf. Sal- vinia). Each of these massulae has its outer surface provided with curious barbed hairs (glochidia), and escapes on its own account. The megasporangium contains one spore. It sinks to the bottom ; decay of the indusium frees the spore and it germinates, giving rise to a ? prothallus which floats about on the water and may be anchored to a floating massula by the barbs. Azorella Lam. Umbelliferae (i. 2). 70 Andes to N.Z. Densely tufted xero. A. caespitosa Vahl. (balsam-bog, Falklands) forms tufts like Raoulia. Azureus (Lat), sky-blue. B (fl. -class), fls. with fully concealed honey, Calluna, many Caryophyl- laceae, Geranium, Gypsophila, Polemoniitm, Rnbiis, Thy tints. B' (fl. -class), like B, but aggregated into dense heads, Armeria, Coni- positae, Dipsaceae, Scabiosa. Babbagia F. Muell. (Osteocarpum EP.}. Chenopodiac. 4 Austr. Babiana Ker-Gawl. Iridaceae (in). 40 S. and trop. Afr., Socotra. Babingtonia Lindl. = Baeckea L. p.p. (Myrt.). Babul, Acacia arabica Willd. Bacca (Lat.), a berry; -cate, with berry; -iform, berry-like. Baccaurea Lour. Euphorbiaceae (A. I. i). 60 trop. Afr., As., Polynesia. Baccaureopsis Pax. Euphorbiac. (A. i. i). i trop. Afr. Baccharis L. Compositae (3). 380 Am., esp. campos. Many are leafless xero. with winged or cylindrical green stems (Bot. Jb. 27, 446). Bachelor's button, double fld. Centaurea, Lychnis, Ranunculus, &c.; also Jasione, &c. ; do. (W.I.), Goniphrena. Bachmannia Pax. Capparidaceae (11). 2 Pondoland. Bacillar, rod-shaped. Backhousia Hook, et Harv. Myrtaceae (n. i). 5 E. Austr. B. citrio- dora F. Muell. gives an essential oil almost entirely citral. BALANOPHORA CEA E 6 7 Baconia DC.^Pavetta L. (Rubi.). Bacopa Aubl. (Herpestis Gaertn. f.). Scroph. (il. 6). 50 warm. Bacterial colonies, Pavctta. Bacteroids, Legnminosae. Bactris Jacq. Palmae (iv. 2). 100 trop. Am., W.I. Fls. in groups of 3, one ? between two $. B. minor Jacq. (pupunha or peach palm, Brazil), ed. fr. Bacularia F. Muell (Linospadix p.p. EP.). Palm. (iv. i). 5 Austr., Malaya. Badiera DC. (Polygala p.p. EP.). Polygalaceae. 10 trop. Am., W.I. Badinjan (W.I.), Solatium Mclongena L. Badius (Lat.), chestnut brown. Badula Juss. (Ardisia Sw. p.p. BH.). Myrsin. (n). 5 E. Afr. Is. Badusa A. Gray. Rubiaceae (l. 5). i Fiji, Society Is. Baeckia L. Myrtaceae (n. i). 60 Austr. to China. Bael, Aegle Marmelos Correa. Baeobotrys Forst. = Maesa Forsk. (Myrsin.). Baeometra Salisb. Liliaceae (i). i S. Afr. Baeria Fisch. et Mey. Compositae (6). 20 Calif. Bagassa Aubl. Moraceae (i). 3 Guiana, N. Brazil. Bagnisia Becc. Burmanniaceae. 3 Malay Arch., N.Z. (?). Bahamas hemp, Agave rigida Mill. var. ; -grass, Cynodon Dactylon. Baliia Lag. Compositae (6). 15 N. Am. BaMa piassaba, Attaleafunifera Mart. Baikiea Benth. Leguminosae (n. 3). 6 trop. Afr. Baileya Harv. et A. Gray. Compositae (6). 3 Utah-Mexico. Baillonacanthus O. Ktze. = Solenoruellia Baill. (Acanth.). Baillonella Pierre (Jllh/iiisops L. p.p. EP.). Sapot. (2). i Gaboon. Baillonia Bocquillon. Verbenaceae (i). 2 S. Am. Baissea A. DC. Apocynaceae (n. i). 12 trop. Afr., As. Bajri, Pennisetum typhoideum Rich. Bakeria Andre. Bromeliaceae (3). i Colombia. Cult. orn. infl. Bakeridesia Hochreut. Malvaceae (2). i Mex. Bakeriella Dubard. Sapotaceae (i). n Afr. Bakerisideroxylon Engl. (Sideroxylon p.p.). Sapot. (i). 3 trop. Afr. Balaka Becc. (Ptychosperma p.p. EP.). Palmae (IV. l). 2 Fiji. Balangue Gaertn. Oleaceae. i Madag. Balania Nor. Inc. sed. Spp. o. Balania Van Tiegh. (Balanophora p.p. EP.). Balan. 2 E. As. Balaniella Van Tiegh. (Balanophora p.p.). Balan. 10 E. As. Balanites Delile. Zygophyllaceae. 2 Egypt, trop. Afr., Abyss. Oil. Balanocarpus Bedd. Dipterocarpaceae. 16 Indomal. Balanophora Forst. Balanophoraceae. 20 Indomal. Some apogamous (cf. Filicineae, Angiospermae; Treub in Ann. Buitenz. xv). Balanophoraceae (EP.; BH.). Dicots. (Archichl. Santalales). 15 gen., 40 sp., all but one trop. Parasites (no chlorophyll) on tree roots, to which the tuberous rhiz. is attached by suckers. From it springs the infl. (sometimes developed within the rhiz. and breaking through it), which comes above ground as a spike or head with scaly 1. and small unisexual fls. ^T.) = Sideroxylon (Dill.) L. Beaver poison (Am.), Cicuta maadata L. Bebbia Greene. Compositae (5). 3 S.W. U.S. Beccaxianthus Cogn. Melastomaceae (n). i Borneo. Beccariodendron Warb. (Mitrephora EP.}. Anonac. (2). i Oceania. Becium Landl.=Ocimum Tourn. (Labial.). Beckera Fresen. Gramineae (4). 3 Abyss. Beckeria Ridl. Rubiaceae (i. 7). 'i Malaya. Beckmaunia Host. Gramineae (n). i N. temp. Beckwithia Jepson (Ranunculus p.p.). Ranunc. (3). i Calif. Becquerelia Brongn. Cyperaceae (ll). 3 trop. S/Am. Beda nut, Terminalia belerica Roxb. Beddomea Hook. f. (Aglaia p.p. EP.). Meliaceae (in), i India. Bedfordia DC. Compositae (8). 2 temp. Austr., Tasm. Bed-straw, Galium. Bee-flowers, fl. esp. of class H, chiefly visited by bees ; Antirrhinum, Aquilegia, Boraginaceae, Borago, Calluna, Campanulaceae, Cobaea, Compositae, Digitalis, Gentiana, Labiatae, Lamium, Lavandula, Leguminosae, Lotus, Lychnis, Monarda, Onagraceae, Pedicidaris, Phacelia, Pyrus, Rhododendron, Sympkytum, Tiiia, Trifolium, Vacdnium. BELLI DA 75 Beech, Fagns sylvatica L. ; -fern, Diyopteris Phegopterh L.; -mast, the capsules of fr.; seaside- (W.I.), Exostemma. Beef-apple (W.I.), Sapota; -wood, Stenocarpus, Casuarina, &c. Beesha Kunth = Melocunna Trin. (Gramin.) ; do. Munro = Ochlandra. Beetroot, Beta vulgaris L. ; sugar-, a var. Befaria Mutis (Bejaria}. Ericaceae (i. i). 20 trop. and subtrop. Am. B. racemosa Vent, and others (Andes rose) form a consp. feature in the veg., taking the place of Rhododendrons. Begonia (Tourn.) L. Begoniaceae. 600 trop. and subtrop., esp. Am. Many cult, for handsome fls. and foliage. Most are perenn. herbs with thick rhiz. or tubers. Several climb by aid of roots like ivy. L. rad. or alt., in two ranks, with large stips. One side of the 1. is larger than the other, whence the name 'elephant's ear,' by which they are sometimes known. The surface of the 1. is easily wetted, and drip-tips are frequent (cf. Fiats}. In the axils groups of little tubers are frequently found ; these are not axillary branches, but are borne upon the true axillary branch, which does not lengthen. They also repr. easily by adv. buds which readily form on pieces of 1. cut off and placed on the soil under suitable conditions of moisture &c. (the common mode used in horticulture). A callus forms over the wound, and in it there develops a meristem which gives rise to one or more buds. Infl. axillary, clich. with a bostryx tendency. The first axes usually end in c?, the last and sometimes the last but one in ?, fls. In the i , P 2, valvate, or 4, decussate, corolline; A oo , free or not, the connective often elongated and the anthers variously shaped. In the ?, P 2 — 5 ; G usu. (i — 3), with 2 — 3 loc., and axile plac. often projecting far into them ; ovules oo , anatr.; styles ± free. Ovary usu. winged ; the wings persistent upon the capsular fr. No endosperm. Begoniaceae. Dicotyledons (Archichl. Parietales). 4 gen. with the characters of Begonia (q.v.). Placed in Passiflorales by BH. Begoniella Oliv. Begoniaceae. 3 Colombia. Behaimia Griseb. Leguminosae (in. 8). i Cuba. Behen-oil, Moringa pterygosperma Gaertn. Behnia Didrichsen. Liliaceae (x). i S. Afr. Behria Greene. Liliaceae (iv). i S. Calif. Behuria Cham. Melastomaceae (i). 7 S. Brazil. Beilschmiedia Nees. Lauraceae (11). 40 trop., and Austr., N.Z. Bejaria Zea (Befaria Mutis, q.v.). Ericac. (i. i). 15 warm Am. Belairia A. Rich. Leguminosae (ill. i). 2 Cuba. Belamcanda Adans. Iridaceae (n). i E. As. Cult. orn. fl. Belangera Cambess. Cunoniaceae. 10 S. Brazil, Paraguay. Belencita Karst. Capparidaceae (n). i Colombia. Belladonna, Atropa Belladonna L. Belladonna Sweet = Amaryllis L. (Amaryll.). Bell apple (W.I.), Passiflora lanrifolia L.; -flower, Campanula, Wahlen- bergia. Bellardia All. (Bartsia L. p.p. BH.}. Scroph. (in. 3). i Medit. Bellendena R. Br. Proteaceae (i). i Tasmania. Bellevalia Lapeyr. = Hyacinthus Tourn. p.p. (Liliac.). Bellida Ewart. Compositae (3). i Austr. 76 B ELLIS Bellis (Tourn.) L. Compositae (3). 15 Eur., Medit. B. percnnis L. (daisy) Brit. Multiplies and also hibernates by short rhiz. Kay florets ?. The head closes at night and in wet weather. Cult. orn. fl. Bellium L. Compositae (3). 6 Medit. Bellonia (Plum.) L. Gesneriaceae (n). 2 W.I. Bellucia Neck. Melastomaceae (i). 15 trop. Am. Fr. eel. Belly-ache bush (W.I.)tJatropha gossypifolia L. Belmontia E. Mey. Gentianaceae (i). 20 S. and trop. Afr., Maclag. Beloanthera Hassk. Inc. sed. i Java. Belonanthus Graebn. Valerianaceae. 2 Peru, Bolivia. Belonophora Hook. f. Rubiaceae (n. i). i W. Afr. Beloperone Nees. Acanth. (iv B). 45 warm Am., many cult. orn. fl. Belotia A. Rich. Tiliaceae. 'i Cuba, Mex. Bembicia Oliv. Flacourt. (7) (Samyd. BH.}. i Madag. Bembicina O. Ktze. = Bembicia Oliv. (Flacourt.). Bembix Lour. Malpighiaceae (inc. sed.). i Cochinchina. Bencomia Webb, et Berth. Rosaceae (in. 5). 2 Canaries, Madeira. Benevidesia Saldanha et Cogn. Melastomaceae (i). i S. Brazil. Bengal beans, Stizolobium (Mncuna) sp.; -kino, Butea frondosa Roxb. Benl seed (W. Afr.), Poly gala btityracea Heckel. Benincasa Savi. Cucurbitaceae (in). 2 trop. As. The fr. of B. cerifera Savi. is eaten in curries. It has a thick coating of wax. Benjamin, gum-, Styrax Benzoin Dryand.; -bush (Am.), Lindera Benzoin Meissn. Bennettia Miq. Flacourtiaceae (4). 2 S. As. ; do. R. Br. = Galearia. Ben nut, -oil, Moringa pterygosperma Gaertn. Bent-grass, Agrostis, Apera ; silky-, Cynositrus. Benthamantha Alef. = Cracca Benth. (Legum.). Benthamia Lindl. (1830) = Amsinckia Lehm. ; (i833) = Cornus Tourn. Benthamiella Spegazz. Solanaceae (inc. sed.). 10 Patagonia. Bentia Rolfe. Acanthaceae (iv. B). i S. Arabia. Bentinckia Berry. Pahnae (iv. i). 2 India. Benzoin Hayne= Styrax L. do. Nees— Lindera Thunb. Benzoin, gum, Styrax Benzoin Dryand. Benzonia Schum. Rubiaceae (inc. sed.). i W. Afr. Berardia Brongn. = Nebelia Neck. (j9ZT.) = Diberara Baill. (EP.). Berardia Vill. Compositae (12). i Alps. Berberidaceae (EP. ; BH. incl. Lardizabalaceae}. Dicotyledons (Archichl. Ranales). n gen., 135 sp., N. temp. (Berberis in trop. Mts. and S. Am.). Perennial herbs or shrubs, in the former case usu. with sympodial rhiz. Fls. in racemes, ?, reg. ; typical formula P 3 + 3 + 3 + 3» A 3 + 3, G i ; fl. sometimes 2-merous. Of the 4 outer whorls, the two outer are P proper, the two inner ' honey-leaves ' usu. with nectaries at the base (see Rammculaceae). The former are often termed the K, the latter the C. Anthers introrse, but in most cases open by two valves (/a; cress, Cardamine; -nut, Carya ; -orange, Citrus Anrantinm L. var. Biganuiia ; -root, Lnwisia ; -sweet, Solatium Dulcamara L. ; -wood, Picraena, Picramna, &c. Bitteria Borner (Carex p.p.). Cyperaceae (in). 5 N. temp. Bivinia Tul. (Calantica p.p. EP.). Flac. (9) (Samyd. BH.}. i Madag. Bivolva Van Tiegh. (Balanophora p.p.). Balan. 5 India, China. Bivonaea DC. Cruciferae (2). 4 W. Medit. Bixa L. Bixaceae. i trop. Am. B. Orellana L. cult for the seed ; the orange colouring matter of the outer layer of the testa (annatto, arnotto, roucou) is used in dyeing sweetmeats &c. Bixaceae (EP. ; Bixineae BH. incl. Flacourtiaceae and Cochlosper- maceae). Dicots. (Archichl. Parietales EP. BH.}. 3 gen. 6 sp. trop. Small trees or shrubs ; 1. alt. slip, entire. Fl. in panicles, 5 , reg. K 5, C 5, A oo , G (2), i-loc. with parietal plac. and oo anatr. ov. Style simple. Capsule splitting between plac. Seeds with red fleshy papillae. Endosp. starchy. Chief genus Bixa. Bixineae (BH.). Bixaceae, in wide sense. See above. Blachia Baill. Euphorbiaceae (A. II. 5). 8 Indomal. Black bead shrub (W.I.), Pithecolobium Unguis-cati Benth. ; -berry, Rubus fmticosus, &c.; -bindweed, Polygon ni/i Convolvulus L. ; -boy, Xanthorrhoea hastilis R. Br.; -bryony, l^annts communis L. ; -butt, Eucalyptus pilularis Sm. &.C.; -cap raspberry, Rubus occi- dentalis L. ; -dammar, Canarium ; -gram, Phaseolus Mungo L., var. radiatus ; -grass (Am.), Juncits Gerardi Loisel. ; -gum tree (Am.), Nyssa ; -jack (Am.), Quercus nigra L. ; -moss (Am.), Til- landsiansneoides'L.; -mustard, Brassica nigra Koch ; -oil, Celastrus; -snake-root, Ciinicifuga (Actaea) racemosa Nutt. ; -spleenwort, Asplenium Adiantum-nigruin L. ; -thorn, P minis spinosa L. ; -wattle, Acacia decurrens, Willd. ; -wood, Acacia inelanoxylon R.Br., Dalbergia latifolia Roxb., &c. Blackstonia Huds. = Chlora Ren. (Gentian.). Blackwellia Comm. ex Juss. = Homalium Jacq. p.p. (Flacourt.). Blackwellia Gaertn. Inc. sed. i sp. Habitat? Bladder-campion, Silene inflata Sm. ; -fern, Cystopterisfragilis Bernh. ; -nut, Staphylea pinnata L. ; -seed, Physospennum ; -senna, Colutea arborescens L. ; -wort, Utricitlaria. Blaeberry, Vacciniwn Myrtillus L. Blaeria L. Ericaceae (iv. i). 25 S. and trop. Afr. Blainvillea Cass. Compositae (5). 10 cosmotrop. Blairia Houst. ex L. = Priva Adans. (Verben. ). Blakea P. Br. Melastomaceae (i). 35 S. Am., W.I. Ed. fr. Blanchetia DC. Compositae (i). i Bahia. Raises perspiration. Blanchetiastrum Hassler. Malvaceae (3). i Brazil. 6—2 84 BLANCOA Blancoa Lindl. Amaryllid. (in), i S.W. Austr. (Haemodor. BH. ) Blandfordia Sm. Liliaceae (in). 4 E. Austr. Cult. orn. fl. Blanket flower, Gaillardia. Blastania Kotschy et Peyr. Cucurbit. (2). 3 trop. Afr. and As. Blastemanthus Planch. Ochnaceae. 3 N. Brazil, Guiana. K5 + 5. Blastocaulon Ruhland (Paepalanthus p-p.). Eriocaul. 3 Brazil. Blastochore (Cl.), a plant distr. by offshoots. Blastus Lour. Melastomaceae (i).. 4 E. As. Blatti Adans. (Sonneratia L. f. q.v.}. Sonnerat. 6 Indomal. Blattiaceae = Sonneratiaceae. Blazing star, Liatris sqitarrosa Willd. Blechnum L. Polypodiaceae. 220 cosmop. B. boreale Sw. (B. Spicant Roth.) in Brit, has fertile and barren 1., the latter larger. Cf. Bower, Studies, Ann. Bot. 28, 1914, p. 363. Blechum P. Br. Acanthaceae (IV. A). 4 trop. Am., W.I. Bleeding heart, Dicentra', (W.I.), Colocasia antiquornm. Bleekrodea Blume. Moraceae (i). 3 Madag., Borneo, Indochina. Blennodia R. Br. Cruciferae (4). 1 1 Austr. Blennosperma Less. Compositae (6). 2 Chili, California. Blepharacanthus Nees = Blepharis Juss. (Acanth.). Blepharandra Griseb. Malpighiaceae (n). i Guiana. Blepharanthera Schlechter. Asclep. (n. 3). 2 Damaraland. Blepharidachne Hackel (Eremochloe Wats.). Gramin. (10). 2 W.N. Am. Blephariglottis Rafin. = Habenaria Willd. (Orchid.). Blepharipappus Hook. Compositae (5). i W. U.S. Many = Layia. Blepharis hiss. Acanthaceae (iv. B). 80 palaeotrop., Medit., S. Afr. The seeds have hairs which swell up when wetted. Blepharispermum Wight ex DC. Compos. (4). 10 trop. As. and Afr. Blepharistemma Benth. in Wall. Rhizophoraceae. i India. Blepharizonia Greene. Compositae (5). 2 California. Blepharocalyx Berg. (Myrtus p.p. BH.}. Myrtac. (i). 3 warm S. Am. Blepharocarya F. Muell. Anacard. (3). (Sapind. BH.} i E. trop. Austr. Blepharodon Decne. Asclepiadaceae (n. i). 25 Mex. to Chili. Blepharoneuron Nash (Vilfa p.p-)- Gramin. (8). i S.W. U.S., Mex. Blephilia Rafin. Labiatae (vi). 2 N. Am. Bletia Ruiz et Pav. Orchid, (n. 9). 20 trop. Am., W.I. Cult. Bletilla Reichb. f. Orchidaceae (n a. n). i E. As. Blighia Kon. Sapindaceae (i). 3 trop. Afr. B. sapida Kon (akee, vegetable marrow) cult, for ed. fr. (fleshy arillate seed stalk). Blimbing, Averrhoa Biliinbi L. Blinding tree (Ceylon), Excoecaria Agallocha L. Blinks, Montia fontana L. Blinkworthia Choisy. Convolvulaceae (i). 2 Burma. Blitum (Tourn.) L. = Chenopodium L. p.p. (Chenopod.). Blood-flower, Hae»ianthus, (W.I.) Asclepias curassavica L. ; -leaf, Iresine ; -lily, Haemanthus ; -root, Sanguinaria caitadt'iisis L. ; -wood, Eucalyptus, (W.I.) Laplacea. Bloomeria Kellogg. Liliaceae (iv). 2 S. California. BOISDUVALIA 85 Blue-bell, Campanula rotundifolia L., Sdlla festalis Salisb. ; -berry (Am.), Vaccininm spp. ; -bottle, Centaurea Cyamis L. ; -flag (Am.), Iris ; -grass (Am.), Poa pratensis L. ; -gum, Eucalyptus Globulus Labill.; -lettuce (Am.), Mttlgedium ; -tangle (Am.), Gaylussacia frondosa Ton. et Gray; -weed (Am.), Echiuin. Bluet, Vaccinium pennsylvanicum Lam., Houstonia. Blumea DC. Compositae (4). 80 palaeotrop. Ai or ngai camphor is distilled from B. balsamifera DC. (S.W. China). Blumenbachia Schrad. Loasaceae. 6 temp. S. Am. Fr. very light, twisted, covered with grapnel hairs. Blumeodendron Muell.-Arg. =Mallotus Lour. p.p. (Euphorb.). Blunt, with rounded end. Blysmus Panz. =Scirpus Tourn. p.p. (Cyper.). Blyxa Noronha. Hydrocharidaceae. 10 •* warm. Blyxopsis O. Ktze. (Enhydrias Ridl.). Hydrocharid. i Malay Penin. Bo (Ceylon), Ficus religiosa L. Bobartia L. Iridaceae (n). 10 S. Afr. L. sword-like or centric. Bobea Gaudich. Rubiaceae (n. 2). 5 Sandwich Is. Bobua DC. = Symplocos L. p.p. (Symploc.). Bocagea St. Hil. Anonaceae (i). ro trop. Am. and As. Bocconia Plum, ex L. (BH. inch Macleya Reichb.). Papaveraceae (ti). 2 Mex. to Peru, W.I. Apetalous. Bocoa Aubl. (InocarpiiS Forst. Bff.}. Leguminosae (in. 8). i Guiana. The seeds are ed. Bocquillonia Baill. Euphorbiaceae (A. 11.2). 6 New Caled. Bodinieria Leveille et Vaniot. Ranunc. (2). i China. Bodinieriella Leveille. Ericaceae (n. i). i China. Boea Comm. ex Lam. Gesneriaceae (i). 25 trop. As. and Austr. Boebera Willd. = Dyssodia Cav. p.p. (Compos.). Boeberastrum Rydberg (Dyssodia p.p.). Compos. (10). 3 S.W. U.S. Boeckeleria T. Durand (Decalepis Boeck.). Cyper. (n). i S. Afr. Boehmeria Jacq. Urticaceae (3). 60 trop. and N. subtrop. B. nivea Gaudich. has good drip-tips (cf. Acer). It is cult, in China for the fibre (China grass, rhea) obtained from the inner bark (cf. Linum), perhaps the longest, toughest, and most silky of all veg. fibres, but most difficult to prepare. In the trop. the var. tenacissima (ramie) is cult. Boehmeriopsis Komarow. Urticaceae (3). i Corea. Boeica C. B. Clarke. Gesneriaceae (i). 6 S.E. As. Boenninghausenia Reichb. Rutaceae (n). i Khasias to Japan. Boerhaavia Vaill. ex L. Nyctaginaceae. 40 cosmop. Anthocarp often glandular, aiding in seed-dispersal. Boerlagea Cogn. Melastomaceae (i). i Borneo. Boerlagella Pierre. Sapotaceae (n). i Sumatra. Boerlagiodendron Harms. Araliaceae (i). 20 Malaya. Bog-asphodel, Narthecium 'ossifragum Huds. ; -bean, Menyanthes tri- foliata L. ; -myrtle, Myrica Gale L. ; -rush (\\\\.},Juncus. Bogoria J. J. Sm. Orchidaceae (11. 20). r Java. Bois fldele, Citharexylum ; -immortelle, Erythrina nmbrosa. Boisduvalia Spach (Oenothera p.p. BH.}. Onagr. (2). 8 W. coast Am- 86 BOISSIERA Boissiera Hochst. et Steud. Gramineae (10). i W. As. Bojeria DC. (Inula p.p. EP.}. Compositae (4). 3 Maclag., S. Afr. Bolandra A. Gray. Saxifragaceae (i). 2 Pac. N. Am. Bolanosa A. Gray. Compositae (i). i Mex. Bolax Comm. ex Juss. = Azorella Lam. (Umbell.). Bolbophyllaria Reichb. f. Orchidaceae (n. 16). 6 trop. Bolbophyllum Sprang. = Bulbophyllum Thou. (Orchid.). Bolboschoenus Palla (Scirfius p.p.). Cyper. (i). i cosmop. Bolboxalis Small (Oxalis p.p.). Oxalidaceae. i S. Afr. Boldo, Peumus Boldns Molina. Boldoa Cav. Nyctaginaceae. i Mex., W.I. Boldoa Endl. = Peumus Molina. (Monim.) Bole, straight main trunk. Boleum Desv. Cruciferae (2). i Spain. Bolivaria Cham, et Schlecht. = Menodora Humb. et Bonpl. p.p. Boll, a capsule, especially of cotton. Bollea Reichb. f. (Zygopetalum p.p. BH.}. Orchid, (n. 14). 3 W. trop. Am. Bollwilleria Zabel (Pyrus p.p.). Rosaceae (n). 2 Eur., Meclit. Boltonia L'Herit. Compositae (3). 4 U.S. Bolusanthus Harms (Lonchocarpus p.p.). Leguni. (ill. i). i S. Afr. Bolusia Benth. Leguminosae (ill. 6). i S. Afr. Bomarea Mirb. Amaryllidaceae (in). 90 Mex., C. and S. Am. L. like Alstroemeria. Often climbing. Umbels cymose. Cult. orn. fl. Bombacaceae (EP. ; Malvaceae^, p. BH. ). Dicots.(Archichl.Malvales). 20 gen., 140 sp., trop., esp. Am. Trees, often very large, with thick stems, sometimes egg-shaped owing to formation of water storage tissue ; 1. entire or palmate, with deciduous slips. Fl. § , often large, usu. reg. K (5), valvate, often with epicalyx ; C 5, conv., pets, asymmetric ; A 5 — oo , free or united into a tube, pollen smooth ; G (2 — 5), in the latter case the cpls. opp. the pets., multiloc. ; style simple, lobed or capitate ; ovulus 2 — oo in each loc.. erect, anatr. Capsule ; seeds smooth, often embedded in hairs springing from wall ; enclosp. little or o. The Adansonieae are ± myrmecophilous (cf. Acacia), with extrafloral nectaries on 1., K, or fl. stalk. Chief genera Adansonia, Bombax, Chorisia, Durio, Eriodendron. Bombax L. (incl. Pachira Aubl. EP.). Bombacaceae. 60 trop. B. inalabaricum DC. (cotton-tree, Ind., Ceylon) drops its 1. in Dec. and remains leafless till Apr., but fls. in Jan. The cotton is used for cushions, &c. Dug-out canoes are made of the soft wood. Bombay aloe, Agave \ -hemp, Crotalaria, Agave. Bombycidendron Zoll. ct Morr. (Hibiscus p.p.). Malv. (4). 4 Phil. Is. Bombycinus (Lat.), silky. Bombynia Nor. Inc. sed. Nomen. Bonace (W.I.), Daphnopsis. Bonamia Thou. Convolvulaceae (i). 3 Sandwich Is., Madag. Bonania A. Rich. Euphorbiaceae (A. n. 7). 6 Cuba. Bonannia Guss. Umbelliferae (in. 6). 2 S. Eur. Bonapartea Haw. = Agave L. (Amaryll.). Bonatea Willd. (Ilabenaria p.p.). Orchid, (n. i). ,s trop. and S. Afr. Bonatia Schlechter et Krause. Rubiaceae (i. 8). i New Calecl. BORA GIN A CEAE Medit. i E. Medit. Bonaveria Scop. (Securigera DC.). Legum. (in. 5). Bonavist (W.I.), Dolichos Lablab L. Boneset (Am.), Eupatorium. Bongardia C. A. Mey (Leontice p.p. EP. ). Berberid. Bonla Bal. Gramineae (13). i Tonquin. Boninia Planch. Rutaceae (i). 2 Bonin Is. Bonjeania Reichb. =Dorycnium L. p.p. (Legum.). Bonnaya Link et Otto (Ilysanthes p.p. EP.}. Scroph. (n. 6). 5 Warm. Bonnetia Mart, et Zucc. Theaceae. 5 trop. Am. Bonniera Cordemoy. Orchidaceae (n. 20). 2 Reunion. Bonnierella Viguier (Panax p.p.). Araliaceae (i). 2 Tahiti. Bonplandia Cav. Polemoniaceae. 2 Mex. Bontia L. Myoporaceae. i W.I. Bonyunia Schomb. Loganiaceae. 3 Guiana, Brazil. Boopis Juss. Calyceraceae. 25 Andes, Argentina. Boottia Wall. Hydrocharidaceae. 20 palaeotrop. Boquila Decne. Lardizabal (Berber. BH.}. i Chili. Borage, Borago officinalis L. Boraginaceae (EP., BH.). Dicots. (Sympet. Tubiflorae; Polemo- niales BH.}. 85 gen. 1600 sp., trop. and temp., esp. Medit. Most are herbs, perenn. by fleshy roots, rhiz., &c. ; a few shrubs and trees. L. usu. alt., exstip., generally, as well as the rest of the plant, covered with stout hairs (hence the name Asperifolieae, by which the B. are sometimes known). Infl. a coiled cincinnus, sometimes double, with a marked dorsiventrality. As the fls. open it uncoils, so that the newly opened fls. face always in the same direction. The morphology of this infl. is but imperfectly understood; adnation or concrescence occurs, and apparently dichotomy at the growing apex. The general agreement is, however, in favour of the view that the "boragoid," as it is sometimes called, is composed of dorsi-ventral monopodia. Cf. Nat. Pfl., Miiller in Flora, 94, Schumann, Morphol. Studien, n. Fl. ?, usu. reg., hypog., and 5-merous. rarely valvate, the odd sepal post.; C (5), funnel-shaped or tubular, the limb usu. flat; AS, epipet., alt. to petals, anthers introrse; G(2), on hypog. disc, usu. 4-loc. by "false" septum (see Labiatae), usu. with gynobasic style ; ovules i in each loc., erect, anatr. Fr. a drupe or 4 achenes. Seed with straight or curved embryo in usually slight endosp. ; the radicle directed upwards. [Cf. § V.] The floral arrangements are interesting. Most B. have a short tube, partly conceal- ing the honey; many (esp. IV. i and 3) have scales projecting inwards from the throat of the corolla, fully concealing the honey, protecting it from robbery and narrowing the entrance, so that visiting insects must take a definite track. " Many sp., in the course of their individual development, seem to K (5), imbr. or open, Floral diagram of Anchusa ; after Eichler. 88 BORAGINACEAE recapitulate to us the evolution of their colours — white, rosy, blue in several sp. of Myosotis; yellow, bluish, violet in M. versicolor; and red, violet, blue in Pulmonaria, Echium, &c. Here, white and yellow seem to have been the primitive colours." (Miiller.) Many B. are heterostyled, e.g. Pulmonaria. The fls. of many sp. are pendulous (and thus bee-flowers), e.g. Borago, Symphytum. Echium is gyno- dioecious. Classification and chief genera (after Giirke) : I. CORDIOIDEAE (drupe; style terminal: twice bi- lobed): Cordia. II. EHRETIOIDEAE (do.; style simple or bi-lobed or double ; no ring of hairs) : Ehretia. III. HELIOTROPIOIDEAE (do., do., but ring of hairs near tip of style): Tournefortia, Heliotropium. IV. BORAGINOIDEAE (style gynobasic : achenes). 1. Cynoglosseae (fl. reg. ; base of style more or less conical ; tips of achenes not projecting above pt. of attachment) : Omphalodes, Cynoglossum, Rindera. 2. Eritrichieae (do., but tips projecting above pt. of attach- ment): Echinospermum, Eritrichium, Cryptanthe. 3. Anchuseae (fl. reg.; base of style flat or slightly convex ; achenes with concave attachment surface) : Symphytum, Borago, Anchusa, Alkanna, Pulmonaria. 4. Lithospermeae (do., but surface of attachment flat) : Myo- sotis, Lithospermum, Arnebia, Cerinthe. 5. Echieae (fl. zygomorphic) : Echium. V. WELLSTEDIOIDEAE (4-merous ; ov. compressed, 2-loc. with one pend. ov. in each ; caps.) : Wellstedia. Boragineae (RH. ) = Boraginaceae. Boraginodes Post et O. Ktze. (Trichodesma p.p.). Borag. (iv. i). i Somaliland. Borago L. Boraginaceae (iv. 3). 3 Medit., Eur., As. B. officinalis L. Brit, (borage) cult, for bee feeding. It has a typical bee-fl. The blue pendulous fl. secretes honey below the ovary ; the elastic sta. form a cone and dehisce introrsely from apex to base, the pollen ripening gradually and trickling into the tip of the cone. Insects probing for honey dislocate the sta., receiving a shower of pollen (cf. Erica, Galanthus, Cyclamen). In older fls. the stigma, now ripe, projects beyond the sta. so as to be touched first. Boragoid, Boraginaceae. Borassodendron Becc. Palmae (n). i Perak. Borassus L. Palmae (n). 2 palaeotrop. B. fiabellifer L. (Palmyra palm) cult, in Ceylon, India, &c. Dioecious. Its uses are legion ; an old Tamil song enumerates 801. The wood of the trunk is very hard and durable, and resists salt water ; it is also used for rafters, well-sweeps, &c. The large fanshaped 1. are used as thatch, and made into olas or writing "paper" sheets, the writing being done upon them with a stylus. From the base of the 1. Palmyra fibre is collected, and used for making brushes, &c. The split 1. are woven into mats, baskets, &c. The fr. is eaten roasted, and the infl. is tapped for toddy (cf. Cocos, Agave) from which sugar or jaggery is BOTRYCHIUM 89 made, as well as vinegar, &c. The young seedlings are also eaten and yield a good flour when ground, and there are many other uses. Borbonia L. Leguminosae (ill. 3). 158. Afr. Borderea Miegeville (Dioscorea p.p. BH.}. Dioscor. i Pyrenees, i Chili. Borea Zipp. Inc. sed. Nomen. Boreava Jaub. et Spach. Cruciferae (2). 2 E. Medit. Borecole, Brassica oleracea L. var. Boree (Austr. ), Acacia pendula A. Cunn. Boretta Neck. = Daboecia D. Don (Eric.). Borneo camphor, Dryobalanops aromatica Gaertn. ; -rubber, Willugh- beia edulis Roxb., &c. Bommuellera Hausskn. Cruciferae (4). i E. Medit. Boronella Baill Rutaceae (in). 2 New Caled. Boronia Sm. Rutaceae (i). 65 Austr. Borreria G. F. W. Mey. (Spermacoce L.). Rubi. (n. 10). 95 trop. BorricMa Adans. Cumpositae (5). 6 warm Am., W.I. Borsczowia Bunge. Chenopodiaceae (B). i Aral plain. Borthwickia W. W. Smith. Capparidaceae (n). i Burma. Borya Labill. Liliaceae (in). 2 Queensland, W. Austr. Borzicactus Riccobono (Cereus p.p.). Cactac. (in. i). i Ecuador. Boschia Korth. Bombacaceae. 4 Malaya. Boschniakia C. A. Mey. Orobanchaceae. i N.E. As., N.W. N. Am. Boscia Lam. Capparidaceae (n). 20 trop. Afr. Bosea L. Amarantaceae (2). 3 Medit., India. Bosistoa F. Muell. Rutaceae (i). 2 E. Austr. Bosleria Aven Nelson. Solanaceae (2). i Nevada. Bosqueiopsis de Wild, et Durand. Morac. (n). 2 trop. Afr. Bosquiea Thou. Moraceae (n). 4 Madag., trop. Afr. Boss, a protuberance. Bossekia Necker (Riibus p.p.). Rosaceae (ill. 2). 2 N. Am. Bossiaea Vent. Leguminosae (in. 3). 35 Austr. Several xero. sp. have flattened green stems (phylloclades) with minute scaly 1. As in Acacia, &c., seedlings show transitions from 1. Bostrychanthera Benth. Labiatae (in), i China. Bostryx, a monoch. cyme where each lat. branch falls upon the same side of relatively main axis, Begonia, Bittomus, ffeuierocallis, Hypericum, Liliaceae. Boswellia Roxb. ex Colebr. Burseraceae. 10 trop. As. and Afr. B. Carteri Birdw. (Somaliland, &c.) and other sp. yield the resin frankincense or gum-olibanum, formerly offic., now used in incense. Other sp. also yield fragrant resins. B. serrata Roxb., an important tree on dry hills in India. Bothriochloa O. Ktze. (Andropogon p.p.). Gramin. (2). i Annam. Bothriocline Oliv. Compositae (i). 10 trop. Afr. Bothriospermum Bunge. Boragin. (iv. 2). 5 trop. and N.E. As. Bothriospora Hook. f. Rubiaceae (i. 8). i Guiana. Botryanthus Kunth - Muscari Mill. p.p. (Liliaceae). Botryceras Willd. (Laurophyllus EP.). Anacard. (3). i S. Afr. Botrychlum Sw. Ophioglossaceae. 40 cosmop. B. Lunaria Sw. (moonwort) in Brit. Habit like Ophioglossum, but the sterile as 90 BOTRYCHIUM well as the fertile part of the 1. is usu. branched. The r. appear one at the base of each 1., and branch monopodially. The spike is usu. much branched, the ultimate twigs being the sporangia. B. Lunaria has no veg. repr. like Ophioglossum, and each new pi. comes from a prothallus, which is small, not > i or 2 mm. long, oval, saprophytic, buried to a depth of i-io cm. In B. virginianum Sw. it is as mvfch as 20 mm. long, and seems to remain attached to the sporophyte for 5 or 6 years. The prothallus has a mycorhiza. (Jeffrey, Univ. of Toronto Studies, 1898; Bruchmann in Flora, 96, 1906, p. 203.) Botrymorus Miq. (Pipturns Wedd. EP.). Urtic. (3). i Malaya. Botryophora Hook. f. Euphorbiaceae (B. li). i Perak. Botryopleuron Hemsl. Scrophulariaceae (in. i). 5 China. Botryose, racemose. Bottionea Colla. Liliaceae (in), i Chili. Bottle-brush, Callistemon; -cod-root (W.I.), Capparis; -gourd, Lage- naria; -grass (Am.), Setaria viridis Beauv. Boucerosia Wight et Arn. (Caralluma p.p. EP.). Asclepiad. (n. 3). 30 trop. Afr. and As. Bouchardatia Baill. (Mdicope BH.}. Rutac. (i). i E. Austr. Bouchea Cham. Verbenaceae (2). 20 trop. Bouchetia DC. Solanaceae (4). 3 Texas to Brazil, Bouea Meissn. Anacardiaceae (i). 4 Malaya. Bouetia A. Chevalier. Labiatae (vn). i Dahomey. Bougainvillaea Comm. ex Juss. Nyctaginaceae. 12 S. Am. The group of 3 fls. is surrounded by 3 lilac or red persistent bracts. B. spectabilis Willd. is a splendid flowering creeper often cult. Bougueria Decne. Plantaginaceae. i Andes. Bourgeon, to bud or sprout. Bourgia Scop. Boraginaceae (inc. sed.). Nomen. Bournea Oliv. Gesneriaceae (i). i China. Bourreria P. Br. (Beureria Jacq.). Boragin. (n). 30 trop. Am., W. Ind. Bousigonia Pierre. Apocynaceae (i. i). 3 Cochinchina. Boussingaultia H. B. et K. Basellaceae. 10 trop. Am. Tubers ed. Bouteloua Lag. Gramineae (n). 40 Canada to S. Am., mainly in SW. U.S. (mesquit grasses, grama, side-oats). They form a large proportion of the herbage of the prairie, and are valuable as fodder. Boutonia DC. (Periblema DC. BH.}. Acanth. (iv. A), i Madag. Bouvardia Salisb. Rubiaceae (i. 5). 30 trop. Am. Some heterostyled like Primula. Cult. orn. perf. fls. Bouzetia Montr. Rutaceae (inc. sed.). i New Caled. Bowdichia H. B. et K. Legumin. (in. i). 2 trop. S. Am. Good wood. Bowenia Hook. Cycad. i Queensland, B. spectabilis Hook., easily recognized by the bipinnate 1. The upper part of the main r. gives rise to curiously branched apogeotropic r., which contain Anabaena (an alga) living in symbiosis, and branch exogenously (Ann. Bot., 1898). Bowlea Ilarv. Liliaceae (in), i S. Afr., B. vo/nbilis Harv., a xero. like Testudinaria, with a large partly underground stock (corm), BRACPIYRIS 91 giving off each year a much-branched climbing stem. This bears small 1., but they soon drop, and assim. is carried on by the green stem. Bowkeria Harv. Scrophulariaceae (n. 4). 7 S. Afr. Bowlesia Ruiz, et Pav. Umbelliferae (i. 2). 20 Am. Bowringia Champ. Leguminosae (ill. i). i Hongkong. Bowstring hemp, Sansevieria zeylanica Willd. Bow-wood, JMaclura aurantiaca Nutt. Box, Buxus', (Austr.) Eucalyptus; jasmine-, Phillyrea; Maracaibo or W. Indian-, Casearia praecox Griseb. ; -wood (W.I.), Vitex itinbrosa Sw. Boykinia Nutt. Saxifragaceae (i). 8 N. Am., Japan. Brabejaria Burm. f. Inc. sed. i S. Afr. Brabejum L. Proteaceae (i). i S. Afr., B. sfcllatifoliuni L. (wilde castanjes), whose seeds are eaten roasted. Bracea Britton. Apocynaceae (inc. sed.). i Bahamas. Bracea King = Sarcosperma Hook. f. (Sapot.). Brachialis (Lat.), a cubit long; brachiate, with spreading branches. Brachiaria Griseb. = Panicum L. p.p. (Gram.). Brachiolobos All. = Nasturtium R. Br. (Crucif.). Brachionidium Lindl. Orchidaceae (n. 8). 6 W.I., trop. S. Am. Brachistus Miers. Solanaceae (2). 20 Cent, and S. Am. Brachtia Reichb. f. Orchid, (n. 19). 3 Colombia. Brachy- (Gr. pref.), short. Brachyachaenium Baker (Dicome p.p.). Compos. (12). i Madag. Brachyactis Ledeb. (Aster p.p. EP.}. Compos. (3). 6 N. As., N. Am. Brachyandra Phil. Compositae (2). 2 Chili. Brachybotrys Maxim. Boraginaceae (iv. i). i China. Brachycarpaea DC. Cruciferae (i). i S. Afr. Brachychaeta Torr. et A. Gray. Compos. (3). r S. U.S. Brachychilum Petersen. Zingiberaceae (i). i Java. Brachychiton Schott et Endl. (Steradia L. p.p. BH.}. Sterculiaceae. n Austr. B. rnpcstris K. Sebum, (bottle tree) has swollen stems, B. acerifolius N. Muell. (flame tree) very fine fl. Brachyclados D. Don. Compositae (12). i S. Andes. Brachycome Cass. Compositae (3). 50 Austr., N.Z., N. Am., Afr. Brachycorythis Lindl. Orchidaceae (11. i). 12 W. and S. Afr. Brachyelytrum Beauv. Gramineae (8). 4 warm Am., Afr. Brachyglottis Forst. Compositae (8). i N.Z. Brachygyne Small (Seymeria p.p.). Scroph. (in. 2). i N. Am. Brachylaena R. Br. Compositae (4). 10 S. and trop. Afr. Shrubs. Brachylepis C. A. Mey. = Anabasis L. p.p. (Chenopod.). Brachylepis Wight et Am. Asclepiadaceae (i). i Nilgiris. Brachyloma Sond. Epacridaceae. 7 Austr. Brachylophon Oliv. Malpighiaceae (i). 3 Malay Penins. Brachynema Benth. Ebenaceae. i N. Brazil. Brachyotum Triana. Melastomaceae (i). 35 S. Am. Brachypodium Beauv. Gramineae (10). 10 temp., and Mts. of trop. 2 Brit, (false brome grass). Leaf reversed (cf. Alstroemeria). Brachypterys A. Juss. Malpighiaceae (i). 3 trop. S. Am., W.I. Brachyris Ntttt. = Gutierrezia Lag. p.p. (Compos.). 92 BRA CJfYSEMA Brachysema R. Br. Leguminosae (in. 2). 15 Austr. Brachysiphon A. Juss. Penaeaceae. 5 S. Afr. Brachystegia Benth. Leguminosae (n. 3). 20 trop. Afr. Brachystelma R. Br. Asclepiadaceae (n. 3). 50 palaeotrop. B.Bingeri A. Chev. has an ed. tuber. Brachystelmaria Schlechter. Asclepiad. (n. 3). 6 S. Afr. Brachystemma D. Don (Arenaria p.p. BH.). Caryoph. (i. i). i Himal. Brachysteplianus Nees. Acanthaceae (iv. B). 10 trop. Afr., Matlag. Bracnythalamus Gilg. Thymelaeaceae. 2 New Guinea. Brachytome Hook. f. (Kandia p.p. EP.). Rubiaceae (i. 8). 2 In- domal. Bracken, Pteridiuin aqnilinum (L.) Kuhn. Brackeniidgea A. Gray. Ochnaceae. 10 palaeotrop. Bract, the 1. in whose axil a fl. arises ; coloured-, Amherstia, Bougain- villaea, Castilleja, Euphorbia, Sahna, &c. ; persistent-, forming wings to fr., Bougainvillaea, Carpinus, Mirabilis, Spinada\ bract- eate, bearing bracts ; bracteole, a bractlet, borne on same axis as the fl. Bradburia Torr. et Gray. Compositae (3). 2 Texas, Mexico. Bradburya Rafin. (Centrosema Benth. Bff.}. Legu. (ill. 10). 30 Am. Bradleia Banks = Glochidion Forst. (Euph.). Bragantia Lour. (Apaina Lam. p.p.). Aristoloch. 5 Indomal. Brahea Mart. Palmae (i. 2). 4 Mexico, Texas. Decorative. Brainea J. Sm. Polypodiaceae. i E. As., B. insignis Sin., a dwarf tree-fern. The primary veins branch and rejoin repeatedly, forming small areas in the leaf; the veinlets run parallel and distinct. Brake, Pteridiwn aquilinum (L.) Kuhn ; curled rock-, Cryptogramme crispa R. Br. Bramble, Rnbus. Branch, an outgrowth of r. or shoot which repeats its structure ; usu. lat. (not dichotomous), exogenous in case of shoot, endogenous in r. In many ferns the branches are on the 1. bases, but in Equisetum and fl. plants are usu. axillary. Usu. only one in each axil ; if more than one, the others are accessory ; if side by side, collateral, Allium, Araceae, Crataegus, Liliaceae, Ahiscari, Quercus, Salix ; if one above the other, serial, Aristolochia, Calycanthus, Cercis, Colletia, Fuchsia, Gleditschia, Robinia, Syringa. Branches of two kinds — long and short shoots, or shoots of unlimited and limited growth — occur in Pinus and other Coniferae, Berberis, Cactaceae, Ginkgo, Spergula. Branching may be of two types — monopodial, Punts and other Coniferae, Orchidaceae, Paris, Pathos, or sympodial, Acorus, Aglao- nema, Ancistrocladus, Anonaceae, Anthurium, Araceae, Asariim, Eichhornia, Fagus, Iridaceae, Iris, Juncus, Liliaceae, Nartheciuin, Orchidaceae, Peperomia, Pistia, Polygonatttin, Pontederiaceae, Pota- mogeton, Ranunculus, Rhaphidophora, Ulmaceae, Vilis, Zostera. And cf. Buds, Concrescence, &c. Branda (W. L), Chione glabra DC. Brandegea Cogn. Cucurbitaceae (4). 2 California. Brandesia Mart. = Telanthera R. Br. (Bff.) = Alternanthera Forsk. Brandisia Hook. f. et Thorns. Scrophular. (n. 4). 3 Burma, China, BREAD-FRUIT 93 Brandy-bottle, Nuphar hiteum Sibth. et Sm. Brandzeia Baill. Leguminosae (ll. i). i Seychelles, Madagascar. Brasenia Schreb. Nymphaeaceae (n). t cosmop., exc. Eur. A 12 or more. Brassaia Endl. (Schefflera EP.}. Araliaceae (i). 3 Malaya. Brassaiopsis Dene, et Planch. Araliaceae (i). 10 Indomal. Brassavola R. Br. Orchidaceae (n. 6). 24 trop. Am. Brassia R. Br. Orchidaceae (n. 19). 30 trop. Am. Brassica (Tourn.) L. (BH. incl. Erucastruni Presl and Sinapis L.). Cruciferae (2). 85 Eur., Medit., As. 7 in Brit. Many forms are cult., some for the flr., others for the stem, root, leaf, or seed. B. nigra Koch is the black mustard, whose seeds yield the condiment. B. oleracea L. is the cabbage, with the various races derived from it, such as cauliflower and broccoli (fleshy infl.), kale or curly greens or borecole, brussels-sprouts (a form in which miniature cabbages are produced in all the leaf-axils on the main stem), kohl-rabi or knol- kohl (trop.) (a thickened stem, or conn, showing leaf scars on its surface), &c. B. campestris L. is the turnip, a biennial with thickened root, and a var. of it — B. Napus L. — is the rape, used in salads and in the preparation of rape- or colza-oil, expressed from the seeds. [See De Candolle's Orig. of Cultiv. Pits.] It is of interest to notice the great variety of morphology in the veg. organs, correlated with the different ways in which storage of reserve materials is effected, in the root, stem, leaf, flowerstalk, &c. Sauer-kraut, or salted cabbage, made by packing cabbage shreds in barrels with salt and pepper, and slightly fermenting, is a favourite food in Germany, esp. for winter use. The outer coat of the seed has mucilaginous cell-walls which swell when wetted (cf. Linum). Brassocattleya x Rolfe. Hybrid, Brassavola x Cattleya ; others are Brassolaelia, Brassoepidendrum, Brassocattlaelia (triple). Brathys Mutis ex L. f. = Hypericum Tourn. p.p. (Guttif.). Brauna, Melanoxylon Branna Schott. Bravaisia DC. Acanthaceae (iv. A). 2 trop. Am., W. I. Bravoa Lex. Amaryllidaceae (n). 5 Mexico. Rhizome with tuberous roots. Fl. zygomorphic by bending. Braya Sternb. et Hoppe. Cruciferae (4). 18 Eur., As., S. Am. Brayera Kunth. (Hagenia Willd.) Rosaceae (ill. 5). i Abyssinia. The dried 1 fls. (Koso) are used as a remedy for tapeworm. Brayodendron Small (Diospyros p.p.). Ebenaceae. i Texas, Mex. Brayopsis Gilg et Muschler (incl. Draba p.p.). Cruc. (2). 10 Andes. Brayulinea Small (Guilleminea p.p.). Amarant (3). i Ecuador. Brazil-cherry, Eugenia spp ; -nut, Bertholletia excelsa H. et B. ; -wood, Caesalpinia Sappan L. &c. ; -ian arrowroot, Manihot utilissima Pohl, M. Aipi Pohl ; -nutmeg, Cryptocarya moschata Nees & M. Braziletto (W.I.), Caesalpinia, Peltophorum, Sciadophyllum, Wein- mannia. Brazoria Engelm. et Gray. Labiatae (vi). 2 Texas. Brazzeia Baill. Tiliaceae. 2 W. trop. Afr. Bread-fruit, Artocarpus incisa L ; Nicobar-, Pandanus. ; -nut, Brosi- (Barbados) Artocarpus. 94 BREDEME YERA Bredemeyera Willd. Polygalaceae. 30 Austr., Tasmania. Bredia Blume. Melastomaceae (i). 3 Japan, China. Bremontiera DC. (Indigofera p.p. EP.}. Legum. (in. 6). i Masc. Breonia A. Rich. Rubiaceae (i. 6). 6 Madag., Mauritius. Bretschneidera Hemsi. Hippocast. (Sapind. BH.} i China. Brevi- (Lat. pref.), short; -lobous, -lobed, &c. Brevoortia Wood. Liliaceae (iv). i Calif. Breweria R. Br. Convolvulaceae (i). 25 trop. and subtrop. Brexia Noronha. Saxifragaceae (v). i Madag., Seychelles. Breynia Forst. Euphorbiaceae (A. i. i). 20 trop. As., Austr. Briar, Rosa ; -wood, Erica scoparia L. Bricchettia Pax. Euphorbiaceae (A. I. i). i Somaliland. Brickellia Ell. Compositae (2). 75 trop. and subtrop. Am. Bridelia Willd. Euphorbiaceae (A. 1.2). 40 palaeotrop. Bridgesia Bert, ex Cambess. Sapindaceae (i). i Chili. Bridgesia Hook, et Arn. = Ercilla A. Juss. (_5Zf.) = Phytolacca L. p.p. Brieya de Wild. Anonaceae (2). i Belgian Congo. Brighamia A. Gray. Campanulaceae (in), i Sandwich Is. Brillantaisia Beauv. Acanth. (iv. A). 20 trop. Afr., Madag. The posterior sta. are perfect (only case in family). Brinjal, egg-fruit, Solanutn Melongena L. Briquetia Hochreutiner. Malvaceae (2). i Paraguay. Brissonia Neck. = Indigofera L. and Tephrosia Pers. Bristle-fern, Trkhomanes. Britoa Berg. (Campomanesia BH.}. Myrtac. (i). 10 Brazil. Ed. fr. Brittenia Cogn. Melastomaceae (i). i Borneo. Brittonastrum Briq. Labiatae (vi). 15 Mex., SW. U.S. Brittonella Rusby (Mionandra EP}. Malpigh. (i). i Bolivia. Briza L. Gramineae (10). 12 temp. 2 Brit, (quake-grasses). Brizopyrum J. Presl (Distichlis BH}. Gram. (10). 7 S. Afr. Brizula Hieron. (Aphelia BH}. Centrolep. 5 S. Austr., Tasm. Broad bean, Vicia Faba L. ; -leaf tree (W. I.), Terminated latifolia Sw Brocchinia Schult. f. Bromeliaceae (3). 3 trop. Am., W. Ind. Broccoli, Brassica oleracea L., var. Brochoneura Warb. Myristicaceae. 4 Madag., E. Afr. Brodiaea Sm. Liliaceae (iv). 50 W. Am. Fls. in cymose umbels. The sta. have curious projecting appendages. Cult. orn. fl. Brombya F. Muell. (Mclicope p.p. EP}. Rutac. (i). i Austr. Brome grass, Bromns ; false-, Br achy podium. Bromelia Plum, ex L. Bromeliaceae (4). 10 W.I. , Brazil. Someed.fr. Bromeliaceae (EP.,BH.}. Monocotyledons (Farinosae; Epigynaej?//.). 40 gen., 1000 sp. trop. Am. Many terrestrial pi. (xero., living on rocks &c.), but the bulk of the sp., by virtue of their good seed- distribution and their xero. habit, have become epiph., forming a very char, feature in the veg. of trop. Am., more so than the orchids, which they surpass in number of individuals though not of sp. Most have a very reduced stem, bearing a rosette of fleshy 1. channelled on the upper surface and fitting closely together by their bases, so that the whole pi. forms a kind of funnel, usu. full of water. In this are dead 1., decaying animal matter and other debris (certain sp. of Utri- cularia live only in these pitchers). There are a number of adv. r. BR 0 UGHTONIA 9 5 which fasten the plant to its support, but which do not aid in its nutrition, or very little. The bases of the 1. are covered with scaly hairs by which the water in the pitcher is absorbed. Water is stored in the 1., which consist largely of water-tissue. They have a thick cuticle and often bear scaly hairs that reduce transpiration. Some show a totally different habit to this, e.g. Tillandsia usneoides (q.v.). [See Schimper, Epiph. Veg. Amerikas.} Infl. usu. out of the centre of the pitcher; bracts coloured. Fl. usu. ? , reg., 3-merous. P. 3 + 3 or (3) + (3), the outer whorl sepaloid, persistent, the inner petaloid ; A 6, introrse, often epipet. ; G (3), inf., semi-inf., or sup., 3-loc., with oo anatr. ov. on the axile plac. in each. Style i, stigmas 3. Berry or caps. ; seeds in the latter case very light, or winged. Embryo small, in mealy endosp. Classification and chief genera (after Wittmack) : 1. Tillandsieae (caps.; ov. sup.; 1. entire; seed hairy): Til- landsia. 2. Ptiyeae (caps. ; ov. sup. ; 1. thorny; usu. large stem) : Puya, Dyckia. 3. Pitcairnieae (caps.; ov. semi-inf. or almost sup.; 1. entire, or toothed at base, rarely at top) ; Pitcairnia. 4. Bromelieae (berry; ovary inf.; 1. with thorny teeth): Bro- melia, Ananas, Billbergia, Aechmea. Bromheadia Lindl. Orchidaceae (n. 5). 6 Malaya. Bromus Dill, ex L. Gramineae (10). 70 temp., and trop. Mts. 7 Brit. (brome-grass). Of little value as pasture. Brongniartia H. B. et K. Leguminosae (in. 6). 30 trop. Am. Brookea Benth. Scrophulariaceae (n. 4). i Borneo. Brook-lime, Veronica Beccabunga\; -weed, Satnolus Valerandi. Broom, Cytistis (Sarothamnus) scoparius Link.; -bush (W.I.), Par- theniuin ; -corn, Sorghum vulgare Pers. ; butcher's-, Ruscus acu- leattis L. ; -rape, Orobanche ; Spanish-, Spartium junceum L. ; -root, Epicampes ; -tree (W.I.), Baccharis; -weed (W.I.), Cor- chorus, Scoparia. Brosimopsis Sp. Moore. Moraceae (il). i Matto Grosso. Brosimum Sw. Moraceae (li). 10 trop. and S. temp. Am. Infl. remarkable, a spherical pseudo-head composed of one ? fl. and many tf fls. The former is sunk into the centre of the common recept. and its style projects at the top, whilst the latter occupy the whole of the outer surface. Each $ fl. has a rudim. P. and one sta., whose versatile anther in dehiscing passes from a shape somewhat like ^ to one like T. Achene embedded in the fleshy recept. The achene of B. Alicastruw Sw. is the bread-nut (not to be con- fused with Artocarpus, the bread-fruit), which is cooked and eaten in the W.I., &c. [The bread-nut of Barbados is, however, a seeded var. of the bread-fruit.] B. Galactodendron D. Don is the cow-tree or milk-tree of Venezuela. The milky latex flows in considerable quantities, tastes very like ordinary milk, and is used for the same purposes. The wood of several sp. is useful (snake-wood). Brossardia Boiss. Cruciferae (2). i Persia. Brotera Willd. (Cardopatium Juss. ). Compos, (u). 4 Medit. Broughtonia R. Br. (Epidendrum EP.}. Orchid, (n. 6). 2 W. Ind. 96 BROUSEMICHEA Brousemichea Bal. Gramineae (8). i Tonquin. Broussa tea, Vaccinimn Arctostaphylos L. Broussaisia Gaudich. Saxifragaceae (in). 2 Sandwich Is. Broussonetia L'Herit. Moraceae (i). 3 E. As., Polynes. Dioecious; . BH.). Dicots. (Archichl. Resales EP. BH.}. 12 gen., 50 sp. S. Afr. Heath-like shrubs, with alt. exstip. 1., and racemose infl. Fl. 5 usu. reg., 5-merous, generally perig. Sta. in one whorl. Cpls. (3—2) each with 3 or 4 ov., or i with i ov. Caps, with 2, or nut with i, seeds. Aril. Endosp. Chief gniera : Brunia, Berzelia. Brunneus (Lat.), brown. Brunnichia Banks. Polygonaceae (in. i). 3 N. Am., W. Afr. BUCK- BEAN 97 Brunonia Sm. Brunoniaceae. i Austr , Tasm. Herb with rad. entire exstip. 1. Blue fl. in heads, 5 . K (5), C (5), A 5, G i-loc. i-ovuled. Achi-ne. Exalb. Brunoniaceae (EP. ; Cioodeniaceae p.p. BH.}. Dicots. (Sympet. Cam- panulatae). Only genus Brunonia, q.v. Brunsvigia Heist. Amanllidaceae (i). 10 Afr. Cult. orn. fl. Brush (N.S. Wales, Queensland), forest ; -box, Tristunia. Brussels sprouts, Brassica oleracea L. var. Bruxanelia Demist. Rubiaceae (inc. sed.). i E. Ind. Bruyere, Erica sropana L. Brya P. Br. Leguminosae (in. 7). 5 Cent. Am., W. Ind B. Ebenus DC. yields the \vood Jamaica or American ebony, cocus or cocos wood, the heart wood turning black with age (ff. Diospyrosj. Bryanthus S. G. Gmel. Ericaceae (i. 3). i E. Siberia. BrylKinia F. Schmidt, Graniineae (loj. i Japan, Saghalien. Bryo- (Gr. pref. ), moss. Bryocarpum Hook. f. et Thorns Primulaceae. i Sikkim. Bryodes Benth. Scrophulariaceae (11. 6). i Mauritius. Bryomorphe Harv. Compositae (4). i Cape Colony. Bryonia L. Cucurbitaceae (3). 10 Eur., As., Afr. B. dioica Jacq. (Brit., white bryony) marks the N. limit of the family in Eur. 3 fl. larger. Honey secreted at the base of the P. Bryonopsis Am. Cucurbitaceae (3). 2 trop. Afr. and As. Bryony, bastard (W.I. ), Cissits; black-, Tamns comm-unis L. ; white-, Bryonia dtoica |acq. Bryophyllum Salisb. Crassul. 4 trop. In the notche. on the 1. of B. calycimim Salisb. adv. buds develops, giving rise to new pi. In B. proliferum Bowie there are simple and cpd. 1. on the same pi. K and C both gamophyllous. Cult. orn. Bryopsis Reiche (Lyallia^ Rcicheella p.p ). Caryo. (i. 3). i Chili. Buaze fibre, Securidaca longipedunculata Fres. (trop. Afr.). Bubon L. =Seseli L. (Umbel ). Bucco Wendl. = Agathosma Willd. (Rutac.). Bucephalandra Schott. Araceae (v). i Borneo. Bucephalon L. =Trophis P. Br. (Morac.). Buceragenia Greenman. Acanthaceae (iv. B). i Mexico. Buceras Hall, ex All. = Trigonella L. p.p. (Legum.). Buchanania Spre'ig. Anacard. (i). 20 trop. As. G 4 — 6, one fertile. Buchenavia Eichl. (Terminalia p.p. BH.}. Comhret. 8 Brazil., W.I. Bucbenroedera Eckl. et Zeyh. Leguminosae (in. 3). 15 S. Afr. Buchholzia FLngl. Capparidaceae (n). 3 trop. Afr. BucMngera Boiss. et Hohen. Cruciferae (4). i Persia. Buchloe Engelm. (Bulbilis Rafin.). Gramineae (n). i (B. dactyloides Engelm.) the buffalo-grass of the western prairies of the U.S., a good fodder. It is a small creeping grass. Buclinera L. Scrophulariaceae (in. 2) 60 trop. and subtrop. Buchnerodendron Giirke. Flacourtiaceae (2). 6 trop. Afr. Bucholzia Mart.--Telanthera R. Br. (BH.} — Alternanthera Forsk. Bucbu, Bai-o^ma betuiina Bartl. et Wendl. f. and others. Bucida L. (Terminalia p.p. BH.}. Combret. i Cent. Am., W I. Buck-bean, Menyanthes trifoliata L. ; -eye (Am.), Aesculus ohioensis w. 7 98. BUCK-BEAN Michx. ; -'a horn plantain, Plantago Coronopus L. ; -thorn, Rhamnus (W.I.), Rosa laevigata Michx., sea -thorn, Hippophae rhamnoides L. ; -wheat, Fagopymm esculentum Moench. Buckinghamia F. Muell. Proteaceae (11). i Queensland. Bucklandia R. Br. Hamamelidacc-ae. i Himal. to Java, B. populnea R. Br. The large slips, are folded against one another, enclosing and protecting the young axillary hud or infl. Fls. in heads in groups of 4, polyg. or monoec., sunk in the axis. The "calyx-tube" becomes visible as a ring alter flowering. Wood valued. Buckleya Torr. Santalaceae. 5 N. Am., China, Japan. Bucquetia DC. Melastomaceae (i). 2 Colombia, Ecuador. Bud, the much condensed undeveloped shoot end of the axis, composed of closely crowded young 1. with very short internodes, well seen in Brussels sprouts, Cabbage, Lettuce, Acer, Aesculus, Hippuris, Punts, Syringa, Ulmus; usu. axillary or terminal; abortion, Syringa\ accessory, extra buds in an axil, which may be collateral (side by side), Araceae, Crataegns, Muscari, Quercus, or serial (one above another), Aristolochia, ChrysophyJum, Colletia, Fraxinus. Fuchsia, Gledilsckia, Goethea, Gymnocladus, Juglandaceae, Lonicera, Meni- spcrmaceae, Oleaceae, Rhamnaccae. Sambucus, Theophrasta ; ad- ventitious, arising elsewhere than normally in an axil, Begonia, Bryophyllum, Cardamine, Cystopteris, Linaria, Ophioglossum, Ptt-ns, Pyrola ; dormant, branch buds which do not develope at once ; extra-axillary, Fagus, Monstera, Juglans ; -scales, the altered 1. which protect the (usu. winter) hud, Acer, Aesculus, Betula, &c. ; sub-petiolar-, Cladrastis, (ileditschia, Platanus, Rhus, Robinia, Wormia ; winter-, Acer, Aesculus, Betitla, Hottonia, Hydrocharis, Juglandaceae, Myriopkyllitm, Quercus, Rhododendron, Sophora, Utricularia, Viburnum. And cf. Aestivation, Flower bud, Vege- tative Repr., Vernation, and next art. -protection against cold, heat, radiation, &c. is obtained in many ways ; by stipules in Arto- carpus, Bucklandia, Cosmibuena, Cunonia, Dipterocarpus, Ficus, Magnoliaceae; the young 1. are pendulous in Aesculus, Amherstia, Bauhinia, Bioumea, Cinnamomum, Dryobalanops,Maniltoa, Saraca, Theobroma ; the young 1. red in Cinnamomum, Dryobalanops, Haematoxylon, Mesua, &c. ; the buds sub-petiolar in Cladrastis, Glcdilschia, Platanus, Rhus, Robinia, Wormia ; other ways occur in lockroma, Manihot, Philadelphia, Pathos, Spathodea, l^abernae- montana, &c. Buda Adans. = Spergularia Presl (Caryoph.). Buddleia Houst. Loganiaceae. 90 trop. and subtrop. Sometimes placed in Scrophul., but possesses slips (sometimes reduced to inter- peliolar lines). Buena Pohl = Cosmibuena Ruiz et Pav. and Cascarilla Wedd. Buergersiochloa Pileer. Gramineae (5). i New Guinea. Buettneria Loefl. Sterculiaceae. 60 trop. Buffalo-berry, Shepherdia argmtea Nutt. ; -wood, Burchellia. Bufonia Sauv. ex L. (Buffunia}. Caryophyllaceae (i. j). 20 Medit., Eur. Not unlike Juncus bufonius in habit. Buforrestia C. B. Clarke. Commelinaceae. 5 trop. W. Afr. Bugbane, Cimicifuga. BUPLEURUM 99 f Bugle, Ajitga reptans L. Bugloss, Lycopsis arvensis L. ; viper's-, Echium vulgare L. Buglossum (Tourn.) Adans. = Anchusa L. p p. (Borag.). Bugula Tourn. ex Mill. =Ajuga L. p.p. (Labiat.). Bulb, a modified stem, bearing a ± spherical mass of swollen 1. closely folded over one another, A Ilium, Galanthus, Li/iaceae, Lilium, Oxahs. Common in dry climates. Bulbil, a little bulb, usu. in place of a fl. in the infl. , Agave, Alliuiu, Cardamine, Gagea, Globba, Lilinin, Lycopodiuin, Oxalis, Remusatia, Saxifraga, Sfilla, and cf. Vegetative Repr. Bulbilis Rarin. = Buchloe Kngelm. (Gram.). Bulbine L. Liliaceae (in). 25 S. Afr., E. Austr. Bulbinella Kunth. Liliaceae (in). 15 S. Afr., N.Z., &c. Bulbinopsis Borzi. Liliaceae (in). 2 Austr. Bulbocodium L. Liliaceae (l). i Eur. Bulbophyllum Thou. Orchidaceae (n. 16). 350 trop., and S. temp. Epiph. with great reduction of 1. The 1. are often mere scales and assim. is perlormed by the tubers. In B. minulissimum F. Muell., &c., the tubers are hollow with stomata on inner surface (cf. 1. of Empetrum). For the fl. see Darwin's Orchids, p. 137. Cult. orn. fl. Bulbostylis DC. = Brickellia Ell. (Comp.). Bulbostylis Kunth (Kmbristylis p.p.)- Cyper. (i). 60 S. Am., Afr. Bull apple tree (W.I.), Sapota rugosa Griseb. ; -hoof (W.I.), Passi- ftora Murucuja L. ; -ock's heart, Anona reticulata L. ; -pine, Finns; -'s horn thorn, Aciicia sphaerocephala. Bullace, Prumis insititia L. Bullate, puckered. Bulleyia Schlechter. Orchidaceae (n. 3). i Yunnan. Bulliarda DC. = Tillaea Michx. (j6'^7.) = Crassula L. p.p. Bully-tree (W.I.), Dipholis, Alyrsine, Sapota, &c. Bulnesia C. Gay. Zygophyll. 6 Argentina, Chili. Timber. Bulrush, Typha, Sdrpus; -millet, Pennisetttm typhoideum. Bumelia Sw. Sapotaceae (i). 30 Am. Bunch-berry (Am.), Cornus canadensh L. Bunchosia Rich, ex Juss. Malpighiaceae (n). 40 trop. Am., W.I. Bungea C. A. Mey. Scrophulariaceae (in. 3). 3 As. Bunias (Tourn.) L. Cruciferae (4). 5 Medit, As. Bunioseris Jord. (Lactuca p.p.). Compositae (13). 2 France. Buniotrinia Stapf. et Wettst. Umbelliferae (in. 5). i Persia. Bunium L. (Carum L. p.p. BH.}. Umbell. (ill. 5). 30 N. palaeo- temp. Bunophila Willd. =Machaonia Humb. et Bonpl. (Rub.). Bunya-bunya pine, Araucaria Bidwillii Hook. Buphane Herb. Amaryllidaceae (i). 3 S. and trop. Afr. Cult. orn. fl. Buphthalmum L. Compositae (4). 4 Eur., As. minor. B. salici- foliiim L. is a char. pi. of the chalky Alps. Bupleurum (Tourn.) L. Umbelliferae (ill. 5). 100 Eur., As., Afr., N. Am. 4 Brit, (buplever or hare's ear). B. rotundifolinm L. has perfoliate L, whence the name throw-wax (thorow-wax) by which it is known. All sp. have entire 1., umisu. in this family. 7—2 too BUPLEVER Buplever, Bnphurum. Buprestis Spreng. = Bupleurum Totirn. (Umliel.). Bur or burr, a hooked fr. ; -bark (W . I.), Triumfctta; Bathurst or Noogoora, Xanthium : -dock, Arcthim Lappa L. ; -grass (Am.), Ceiuhrus; -marigold, Btdens; -reed (Am.), Spargamum; -weed, Mfdicago, Sparganium, Buraeavia Baill. Euphorbiaceae (A. i. i). 3 New Caled , Fiji. Burasaia Thou. Menispermaceae. 4 Madag Burbidgea Hook. f. Zingiheraceae (i). i Borneo. C-segments large, lat. stds. absent. The small labellum and petaloid sta. stand up in the cent i e of the fl. Burcharaia R. Br. (Reya O. Ktze.). Liliaceae (i). i Austr., Tasm. Burcbellia R. Br. Rubiaceae(i.S). i Cape Col. Buffalo wood, very hard. Burdachia Mart. Maipighiaceae (n). 2 N. Brazil. Bureavella Pierre. Sapotaceae (11). i Indomal. Burkea Benth. Leguminosae (n. i). i W. and S. Afr. Burlingtonia Lindl. = Rodriguezia Ruiz et Pav. (Orch.). Burmannia L. Burmanniaceae. 40 trop. and subtrop. Burinanniaceae (EP., BH.). Monocots. (Microspermae). 5 gen., 60 sp., trop. forest herbs, chiefly 'colourless' saprophytes. P (3 + 3), A 6 or 3, G (3), with parietal plac., or 3-loc. Caps. Seeds oo . Endosp. Chief genera: Burmannia, Thismia. (See Nat. P//., and Ann. of Bot. 1895.) Burmeistera Karst. et Triana. Campanulac. (in). 10 trop. S. Am. Burnatastrum Briq. Labiatae (vn). 2 S. Afr., Madag. Burnatia M. Mich. Alismaceae. i trop. Air. Burnet, Potcrium ; -saxifrage, Pimpinella Saxifraga L. Burnettia Lindl. Orchidaceae (n. 2). i Tasmania. Burning bush (Am.), Euonvnius atr-^piirpureus Jacq . Burn-nose (W.I.), Daphnopsis. Burragea Donn. Smith et Rose (Gaura p.p.). Onagr. (2). 2 Lower Calif. Burrielia DC. Compositae (6). i Calif. Bursa Weber in Wigg. = Capsella Medic. (Crucif.). Bursaria Cav. Pittosporaceae. i Austr. Bursera 'Jacq.' ex L. Burseraceae. 45 trop. Am. B. gummifera L. (birch tree, gommier, turpentine tree) furnishes the balsam resin known as American elemi, chibou, cachibou, or gomart. Burseraceae (EP., BH.). Dicots. (Archichl. Geraniales EP., BH.}. 13 gen-' 35° sp., trop. Shrubs and trees with alt., usu. cpd., dotted 1. Balsams and re.-ins occur, in lysigenous or schizogenous passages. Fls. small, generally unisex., with disc like Rutaceae, 5- or 4-merbus, obdiplo.st. when both whorls of sta. are present. Cpls. (5 — 3), ov. usu. i in each. Ovary multiloc. with one style. Drupe or caps. Seed exalb. Many of the order are useful on account of their resins, &c. Chief genera: Commiphora, Boswellia, Bursera, Canarium. Burtonia R. Br. Leguminosae (in. 2). 10 Austr. Buseria Th. Dur. (Leiochilus EP.). Ruhiaceae (il. 4). i Madag. Bush (Austr.), scrub, cf. Acacia; -clover, Lespedeza. Bush-wood, a forest in which the shrubs are so abundant as to keep the crowns of the trees from touching. BUXUS 10 r Bushiola Nieuvvland (Kochia p.p.). Chenopod. (A), i N. Am. Bussea Harms. Leguminos.ie (II. 7). i E. trop. Afr. Bustelma Fourn. Asclepiadaceae (u. i). i Brazil. Butayea Wiklem. (Pseudoblepkatis EP.). Acanth. (IV. B). i Congo. Butcher's broom, Ruscus aculeams L. Butea Keen, ex Roxb. Leguminosae (in. 10). 4 Ind , China. B. fiondosa Roxb (dhak or palas tree, or bastard teak), one of the handsomest of fig. trees. A red juice flows from incisions in the bark ; when dried it is known as Bengal kino and used as an astringent. The fls. yield a lugitive orange-red dye. The tree also yields lac (see fiats), and is very important for lac cult. Butomaceae (EP.; Alismaceae p.p. BH.}. Monocots. (Helobieae). 4 gen., 5 sp., trop. and temp. Water and marsh herbs with 1. of various types. Infl. usu. a cymose umbel. Fir. £ , reg., 2- or 3 merous, hypog. P 6, in two whorls, the outer sepaloid, the inner | etaloid ^exc. Butomus). Sta. 9-00, with introrse anthers. Cpls. 6-00, apocp., with oo anatr. ov. scattered over their inner walls (cf. Nymphaea), except on midrib and edges. Follicles; seed ex-alb.; embryo straight or horse-shoe shaped. Chief genera: Butomus, Hydrocleis. Butomopsis Kunth ( Tenagocharis EP. ). Butomaceae. i Austr. Butomus L. Butom. i temp. As., Eur. (incl. Brit.), B. unibellatus L. (flowering rush). Infl. a term. fl. surrounded by 3 bostryx- cymes. Butonica Lam. = Barringtonia Forst. p.p. (Lecyth.). Butter, cf. oils ; -bean, Phaseolus vtilgaris L. ; -bur, Petasites officinalis Moench. ; -and eggs (Am.), Linaria; -cup. Ranunculus; -nut, Caryocar, (\m.) Juglans ; shea-, Butyro sperniutn ; -tree, Pentadesma, Bassia ; -wort, Pinguicula. Butterfly flowers (class F), Cuphea, Daphne, Eupatoriiun , Gentiana. Lonicera, Lychnis, Onagraccae, Rubiaceae, &c. ; -orchis, Habcnaria, Oncidiiim. Button-bush (Am.), Cephalanthus\ -tree, (W.I.), Conocarpus ; -weed (W.I.)i Spermacoce, Borreria; -wood, Platanus occidental is. Buttonia MacKen. Scrophulariaceae (in. 2). i S. and trop. Afr. Butua bjchl. = Abuta Aubl. p.p. (Menisperm.). Butyrospermum Kotschy. Sapotaceae (i). 2 trop. Afr. The oily seeds of B, Parkii Kotschy when pressed yield shea butter. Buxaceae (EP. ', Euphorbiaceae p.p. BH.) Dicots (Archichl. Sapin- dales). 6 gen., 30 sp. temp, and trop. Evergreen shrubs with exstip. leathery 1. , and no latex. Fls. in heads or spikes, unisex., reg., apet. or naked. Sta. 4 — oo . G usu. (3), 3-loc., with 3 styles which are persistent on the fr. Ov. 2 — i in each loc., pend., anat., with dorsal raphe. Loculic. caps., or drupe. Seed with caruncle or none. Endosp. Chief genera: Buxus, Pachysandra. Buxanthus Van Tieghem (Buxus p.p.). Buxaceae. 2 trop. Afr. Buxella Van Tieghem (Buxus p.p.). Buxaceae. 2 S. Afr., Madag. Buxus L. Buxaceae. 20 palaeotemp. . W.I. B. Semperuirens L. (box) often cult. Fls. in heads, a term. ? flr. surrounded by a number of i.?00 SP-, chiefly localised in the dry regions of trop. Am., but spreading to a distance N. and S. (Opuntia missouriensis as far as 59° N.), and far up the mountains (to 12,000 ft. and even higher). Even in the damp forest regions some sp. appear as epiphytes. The only representative of the order in the Old World is Rhipsalis, found in Afr., Mauritius, &c., but several sp. of Opuntia, &c. are now nat. in S. Afr., Austr., &c. and becoming troublesome. Xero. of the most pronounced t>pe, exhibiting reduction of the transpiring surface, and also storage of water, often in great quantity. The veg. organs show great var. of type; the classification is perhaps better based upon them than upon the repr. organs. R. generally long and well-developed (in cultivation liable to decay). Stem CACTACEAE 103 fleshy, of various shapes, rarely bearing green 1. , and usu. provided with sharp barbed thorns, which give protection against animals. We may consider briefly some of the more important types of shoot found in C. (refer to genera for further details). The nearest approach to the ordinary plant-type is perhaps Pereskia, which has large green 1., somewhat fleshy, in whose axils are groups of thorns mixed with hairs ; the space occupied by these is termed the areole. About the morphology of the spines there has been much dispute; most authors regard them as repres. the 1. of the axillary shoot, whose stem is undeveloped, but there is also good evidence in favour of the view that they are "emergences." In some gen. they are provided with barbs. The next stage is found in Opuntia, where the stem has taken over the water-storing and assim. functions, but still bears 1. ; in some sp. these aid the stem functions throughout life, but in most they fall off very early, and the stem is usually flattened to expose more surface to air and light. Then we come to Leuchtenbergia, which has an aloe-like habit with the areoles on the tips of the apparent 1. ; the fl. arises either in the axil of the "leaf" or on the areole. Development shows that the apparent 1. is really a cpd. structure. The bud stands, not exactly in the axil, but on the base of the 1., and the two grow out together to form a leaf-cushion or mammilla^ at the outer end of which is the growing point and the rest of the 1. itself; the latter is represented by a small scale (often microscopic) and the former gives rise to the thorns, &c. on the areole. The same phenomenon is seen in Mammillaria, Cereus sp., &c. In some cases the growing point divides, during the growth of the mammilla, into two, one on the tip, the other in the axil, of the cushion. The latter gives rise to the fl. In Cereus, Echinocactus, &c. the stem is ± cylindrical, bearing ribs on which are the areoles at regular intervals; the rib is formed by the "fusion" of mammillae, i.e. by the growth of the tissue under them during their development (cf. formation of sympetalous corolla). In Phyllocactus, Epiphyllum, and sp. of Rhipsalis some or all of the shoots exhibit a flattened leafy form with areoles in notches on their edges. This form appears to be derived from the preceding by abortion of some of the ridges, and reversions are often seen (they appear if access ot light lie prevented). Lastly, other sp. of Rhipsalis show perfecily cylindrical stems. The bulk of the internal tissue consists of parenchyma in which water is stored ; the cell-sap is commonly mucilaginous, thus further obstructing evaporation. The cuticle is thick, and the ridges of the stem are usu. occupied by mechanical tissue, whilst the stomata are in the furrows. Everything thus goes to check transpiration to the utmost extent ; it is very difficult to dry a cactus for the herbarium, and its vitality is very great. Its growth is slow, but sp. of Cereus, &c. reach a great size. Veg. repr. is frequent in the mammillate forms, and occurs to some extent in others. In garden practice, cacti are often multiplied by cuttings, for a piece cut off and stuck into the soil will usually grow. Grafting is also largely resorted to. Fls. usu. solitary (exc. Pereskia), borne upon or near the areoles or in the axils of mammillae, large, brightly coloured, §? , reg. or •!•. P (oo ), showing gradual transition from sepaloid to petaloid 1., spirally 104 CACTACEAE arranged, often up the side of the ovary (ef. Nymphnea). Sta. cc , epipet. G (4 -co), uniloc. with parietal plac. and » anatr. ov. ; style simple. Berry, the flesh derived from the furicles. Endosp. or none. The fr. of many sp. is edible (e.g. Opuntia, &c.). Several are used in making hedges. Cochineal is cultivated on Nopalea, Opuntia, &.C. Classification and chief genera (after K. Schumann): I. PERESKIOIDEAE (habit of ordinary pi., with flat 1. and panicles ; no barbed thorns) : Pereskia (only genus). II. 0 PUN TIG WE 4E (succulents with round or flat leaf-like joints; 1. cylindrical, usu. falling very early; barbed thorns; fl. rotate): Opuntia, Nopalea (only genera). III. CEREOIDEAE (succulents; 1. reduced to scales, often very minute ; no barbed thorns) : 1. Echinocacteae ifl. funnel- or salver-shaped, in or near the areole) : Cereu.s, Phyllocactus, Epiphyllum, Echino- cactus, Melocactus, Leuchtenbergia. 2. Mammillarieae (do., but in axil of mammilla) : Mammil- laria, Pelecyphora. 3. Rhipsalideae (fl. rotate) : Rhipsalis. For lurther details refer to genera. Also Goebel, Pflanzeftb. Sch. and in Flora 1895, Ganong in f'/ora 1894, fiot. Gaz. 1895, Ann. Bot. 1898, Schumann in Nat. Pfl. and Gesammtbesehreibung tier Kakteeti, 1897-99, and Vochting in Pringsh. Jahrb. 1894. Cf. also Euphorbia and Stapelia. Cacteae (Bff.} ^-Cactaceae. Cactiflorae (Warming), the /th order of Choripetalae. Cactus L. =Cactaceae, esp. Mammillaria. Cactus, night-flowering:, Circus ; old man-, <"ereus senilis. Cadaba Forsk. Capparidaceae (ll). 20 palaeotrop. Disc prolonged post, into a tube ; both androphore and gynophore present. Cadalvena Fen/,1. Zin^iberaceae (II). i trop. Air. Cadellia F. Muell. Simarubaceae. 2 subtrop. Austr. Cadetia Gaudich. = Dendrobium Sw. (Orchid). Cadia Forsk. Leguminosae (III. i). 5 E. Afi., Madag., Arabia. Fl. almost reg. with free sta. Cadiscis E. Mey. Compositae (6). i S.W. Cape Colony. Cadjans. Cocos, Ni/>a, &c. Caducous, dropping early. Caecum, a prolongation of the embryo-sac, Casitarina, &r. Caelestina Cass. = Ageratum L. (Compos.). Caeruleus (Lat.), pale sky blue. Caesalpinia L. Leguminosae |ii. 7). 60 trop. and subtrop., often hook climbers. The pods of C. bonducella Fleming (nickar bean) are brought to Eur. by the Gulf Stream. Those of C. coriaria Willd. (divi-divi) are imported from Venezuela and W.I. for tanning. C. sappan L. (Indomal., cult.) and several Brazilian sp. yield a red dye Irom the wood (sappan, Brazil, or peach wood). C. pulcherrima Sw. (peacock fl., Barbados pride) is cult. orn. fl. Caesarea Cambess. =Viviania Cav. p.p. (Geran.). Caesia R. Br. Liliaceae (in). 10 Austr., S. Afr. C A LA THE A 105 Caesius (Lat.), lavender-coloured, or pale green and grey. Caespitose, in tufts. Caesulia Roxb Compositae (4) i N.E. India. Caffein, Coffea, Cola, Ilex. Cafta, Cat ha edulis Forsk. Caiophora Presl (Btumenbachia p.p. BH.). Loasaceae. 50 S. Am. Cajanus DC. Legum. (in. 10). i trop. Afr., As., C. indicus Spreng. (dhal, pigeon pea, or Congo pea) cult, in India, cScc. for its ed. seeds. Cajeput oil, A/elaleitca Leucaden a U shape. CampyliS Lour. Inc. sed i China. Campynema La 'ill. AmaryllMactrae (iv). Austr. Tasm. See fam. Campyneman tie Baill. Amaryllidaceae (iv). i New C.ded. Camwood, ttaplua nitida Afzel. Canada Dalsam, Abies balsaniea Mill. ; -pitch, Tsuga canadensis Carr. ; -rice, Ziz mia a^uatica L. Cinaigre, Rnnux hvmenosepalus Torr. Cauanga Rumph. ex Hook. f. et Thorns. Anonaceae (i). 3 trop. E. As to Austr. C. odorata Hook. f. is cult, for its rls., which yield the perfume known as yiang-ylang or Macassar oil. Canariastrum Engl. Burseraceae. i trop Afr. (?= Uapacd). Canariellum Engl. Burscrareae. i New Caled"nia. Canarina L. Campanulaceae (I. i). 3 Canary Is., trop. Afr. Like Campmula but usu. 6-merou>, and with ed. berry fr. Canarium (Rumph.) L. Barseraceae. 80 trop. As., Afr. C. commune L. (Java aim.ind; ed. see.!) furnishes the resin Manila Elemi (see Bur^era). C. strictum Roxb. (Malabar) and other sp. furnish some of the black dammar of commerce (ci. Agathis). Can iry creeper. T> opaeolum pere^rinum L. (cananen.tifH.ori ); -grass, -seed. Ptialaris canariensis L. ; -wMtewood, Liriodendron tulipi- Jera L. Canavalia DC. Leguminosae (in. 10). 12 trop. C. ensiformis DC. w. 8 CAN A FA LI A (sword or sabre bean, overlook) cult. ed. pods. C. obhisifolia DC. is a common trop. shore plant. Canbya Parry. Papaveraceae (11). 2 California, Mex. Cancellate, latticed. Cancrinia Kar. et Kir. Compositae (7). i Centr. As. Candidus (Lat.), pure white. Candle-nut, Aleurites ; -plant, Dictammts ; -tree, Parmentiera ; -wood (W. I.), Sciadophyllitm, Ainyris. Candollea Labill. in Ann. Mus. Par. 1805 (Stylidium Sw., q.v.}. Stylidiaceae. 85 Austr., N. Z., E. As. Candollea Labill. i8o6-Hibbertia Andr. p.p. (Dillen.). Candolleaceae = Stylidiaceae. Candy-tuft, Iberis amani, L. Cane, a commercial term for stems of grasses (esp. bamboos), climbing palms, &c. ; bamboo-, cf. bamboos ; -brake, Ariindinaria ; dumb-, Dieffenbachia; Malacca-, Calamus; rattan-, Calamus, and cf. Rattan; sugar-, Saccharum ojficinarum L. Tobago, Bactns minor Jacq. ; Whangee-, Phyllostachys. Canella P. Br. (Winterana L.). 2 W. Ind., trop. Am. C. alba Murr. yields Canella bark, used as a tonic and stimulant. Canella bark, see last. Canellaceae = Winteranaceae. Canephora Juss. Rubiaceae (i. 8). i Madagascar. Fls. in clusters at the top of a phyllodineous stalk with a 2-lobed calyculus. Canescent, grey or hoary. Canistrum Morren (Aechmea p.p. £ff.). Bromel. (4). 4 Brazil. Canker-berry (W. L), Solatium bahamense L. Canna L. Cannaceae. 40 trop. and subtrop. Am., C. indica L. cosmop. trop. Many sp. , vars. and hybrids, cult. C. indica (Indian shot) is the basis of most of these. Habit like Zingiberaceae or Marantaceae, but C. can be distinguished even when not in fl. by possessing neither the ligule of the former nor the pulvinus of the latter. Infl. term. usu. composed of 2-fl. cincinni. The two fls. are homodromous, but the bracteole is to the right in one and to the left in the other (behind one or other of the two lat. sepals in the diagram). Fl. $> , asymmetric, epig. K 3, C (3). The A is the most conspicuous part. There is a leafy sta. bearing half an anther on one edge, and a number of paaloid structures round it, usu. 3 but sometimes i or 4. One of these is the labellum (not = that of Zingibera- ceae), and is rolled back on itself outwards. The other two are often termed the wings (a /3 in diagram). When a fourth std. (7, cf. Marantaceae) is present it stands behind the fertile sta. Other sp. have only the labellum. style, 3-loc.; ov. in 2 rows in each loc., anatr. Seed with perisperm and straight embryo. As to the morphological explanation of the A, there are two views Floral diagram of Canna indica (after Eichler). The bracteole i* omitted. S = petaloid style; L = labellum; a/3 = staminodes. G (3) with petaloid Caps., usu. warty. CANTUFFA 115 Kichler (Btiitcndiag. \. p. 174) regards the labellum as a lat. sta. of the inner whorl, and the fertile sta. together with all the stds. as the post. sta. of the same whorl; the other sta. of the inner, and all the sta. of the outer, whorl are wanting. The older view looks upon /3, 7, as the 2 post. sta. of the outer whorl, and the labellum, a, and the fertile sta. as the 3 sta. of the inner whorl. (Cf. this fl. with those of Musaceae, Zingiberaceae and Marantaceae.) The pollen is shed upon the style in the bud ; insects alight on the labellum. touch first the term, stigma and then the pollen. The rhiz. of C. edulis Ker-Gawl. is ed., containing much starch. Cannabaceae (Warm ing) =§ iv. of Moraceae (q-v.). Cannabis (Tourn. ) L. Moraceae (iv). i Cent. As., C. saliva L., the hemp. Infl. like Humulus c? , dioec. Hemp is largely cult, both in temp, and trop. regions, in the former for the fibre, in the latter for the drug. A valuable fibre, used for ropes and other purposes, is obtained from the inner bark of the stem, much as flax is prepared from Linum, and for this purpose the plant is cult, in S. Eur., the eastern U.S., and other countries. In the trop., and esp. in India, the pi. is cult, for the sake of the narcotic resin which exudes from it, and which is used much like opium, both as a drug and as a stimulant. The drug occurs in three common forms, ganja, charas, and bhang. The first is the ? flg. tops with resin on them, packed together, the second, which comes from rather cooler climates, is the resin knocked off the twigs, bark, &c. , and the third, which is largely obtained from the wild plants, is the mature L, with their resinous deposit, packed together. Asiatics are much addicted to the use of hemp as a narcotic. It is smoked, with or without tobacco, and an intoxicating liquor, hashish, is made from it. The resin has an intoxicating stimulating effect. In small quantities it produces pleasant excitement, passing into delirium and catalepsy if the quantity be increased. The names given to the plant among them indicate this use of it, e.g. leaf of delusion, increaser of pleasure, cementer of friendship. The sale of ganja and charas is kept in check in India by a stringent licensing system, but that of bhang, which is collected from the wild plants, is not so easy to control. Cannaceae (EP.; Scitamineae p.p. BH.}. Monocotyledons (Scitami- neae). Only genus Canna (q.v.}. Cannomois Beauv. Restiaceae. 8 S. Afr. Cannon-ball tree, Couroitpita gnianensis Aubl. Canotia Torr. i Calif., New Mexico, doubtfully placed in Rutaceae, but perhaps belonging to Celastraceae. Canscora Lam. Gentianaceae (l). 18 palaeotrop. Cansjera Juss. Opiliaceae. 4 trop. As., Austr. Cantaloupe, melon, Cucumis Melo L. Canterbury bell, Campanula. Cantharospermum Wi^ht et Arn. (AtylosiaBH.). Leguminosae (in. ro). 20 Madag. , trop. As. and Austr. Canthium Lam. = Plectronia L. (Rubiaceae). Canthopsis Miq. (A'am/ia p.p. EP.). Rubiaceae (l. 8). i Timor. Cantua Juss. Polemoniaceae. 8 Peru, Bolivia. Cantuffa Gmel. (Pterolobium R. Br.). Legum. (n. 7). 5 palaeotrop. 8—2 n6 CANUS Canus (Lat.), grey-white. Caopia Adans. = Vismia Vand. (Guttif.). Caoutchouc, see Rubber. Capanemia Barb. Rodr. (Quekettia EP.). Orch. (il. 19). 2 Brazil. Caparrosa, Neea t keif era Oerst. Cape aster, Felicia; -chestnut, Calodendron; -cowslip, LachenaHa; -crocus, Gethyllis; -figwort, Phygelms ; -forget-me-not, Anchusa; gooseberry, Physalis; -honeysuckle, Tecoma; -jasmine, Gardenia; -lily, Crhium; -pondweed, Aponogeton; -primrose, Streptocarpus ; -tulip, Hiicinanthns. Caper, Capparis spinosa, L. Caperonia St Hil. Euphorbiaceae (A. n. 2). 33 trop. Am., Afr. Capet tree (W.I.), Capparis verrucosa Jacq. Capillary, hair-like. Capirona Spruce. Rubiaceae (i. 4). i S. Am. K like Mussaenda. Capitania Schweinf. Labiatae (vn). i E. Afr. Capitate, head -like. Capitularia J . V. Suringar. Cyper. (in), i New Guinea. Capitulum, a head of fl., Compositae, Comae., Dipsac., &c. Capnites Dum. = Corydalis Vent. (Papav.). Capnoides Tourn. ex Adans. = Corydalis Vent. (Papav.). Capnophyllum Gaertn. Umbelliferae (in. 6). 4 Medit., S. Afr. Capnorea Rafin.= Hesperochiron S. Wats. (Hydrophyll.). Capparidaceae (EP. , BH.). Dicots. (Archichl. Rhot-adales ; Parietales BH.). 40 gen., 450 sp., trop. and warm temp., many xero., with reduced, often inrolled, 1. (cf. Empetrum). Herbs or shrubs, with alt. simple or palmate 1., often with stips. (frequently repres. by thorns or glands). Fls. $ , reg., usu. in racemes, bracteate but with- out bracteoles- The P resembles that of Cruciferae (K.2 + 2, €4 diagonal), but great var. occurs in the A. In some sp. of Cleome there are 4 sta. in two whorls, but elsewhere there are more. Some sp. of Cleome, £c. show ictradynamous sta. In others, still further branching of the median sta. occurs and usu. the post. sta. is more branched than the ant. Staminody of some of the branches is frequent. Cpls. typically (i), transv. as in Cruciferae, wi'h parietal plac. In many sp. of sub-order II the number rises to 10 or 12 by the addition of a second whorl of cpls. and by decloublement. Ovules oo , campylotropous. A further complication is the presence of axial effigurations, &c. in the fls. A disc may occur between P and sta. (usually thicker at the post, sic'e), or a gynophore between sta. and ov., or both. Or the disc may grow up in the centre to form an androphore on which the sta. are borne and above them there may be a gynophore also. From the disc there often grow out structures of various shapes and sizes; these may be scales quite free from one another, or, as in Cadaba, &c., may be united into a tube. Or the scales may, as in Steriphoma, &c., alt. with and he joined to the sepals. Fr. a siliqua (with repluin), nut, berry or drupe. Seed exalb. with rmbryo folded in various ways as in Cruciferae. Few are useful : see Capparis, &c. CARAIPA 117 Classification and chief genera (after Pax): A. Mostly shrubs, with hairs or scales, rarely glandu'ar. No replum. I. DIPTERYGIOIDEAE (samara): Dipterygium (only genus). II. CAPPARIDOIDEAE (berry): Capparis, Cadaba, Maerua. III. ROYDS101DEAE (drupe): Roydsia. IV. EMBLINGIOIDEAE. Prostrate undershrubs. Calyx tube present. Petals (2). Nut. Emblmgia (only genus). B. Glandular annuals. Siliqua with replum. V. CLEOMOIDEAE : Cleome, Polanisia. Capparis (Tourn.) L. Capparidaceae (n). 150 trop. and sub-trop. (exc. N. Am.). Many climb by recurved stip. thorns. The rl.-buds of C. spinosa L. (Medit.) are known as capers (cf. Eugenia). Capraria (Tourn.) L. Scrophulanaceae (in. i). 4 warm Am. Capreolatus (Lat.), tendrilled. Caprification. cf. Ficns. Capriflcus Gasp. = Ficus Tourn. p.p. (Morac.). Caprifoliaceae (EP. ; BH. incl. Adoxaceae). Dicots. (Sympet. Rubi- ales EP., BH.). ir gen., 230 sp temp, (chiefly N.) and on trop. Mts. Mostly trees and shrubs with decussate usually exstip. leaves (see Sambucus). Fls. 9 , reg., or in cymes, usually -,-merous with the odd sepal post. Sta. in one whorl, epipet. G (2 — 5), multi- loc. with i — oo pend. ov. in each loc. Fr. usu. berry or drupe (caps, in Diervilla). Embryo small in fleshy endosp. [BH. chars. include herb with rad. 1. and small head of fl. ; P homochlam. ; G semi-inf.] Chief genera: Sambucus, Viburnum, Symphoricarpus, Linnaea, Lonicera, Diervilla. Caprifolium Tourn. ex L. = Lonicera L. p p. (Caprifol.). Capsella Medic. Cruciferae (4). 4 N. temp. C. Bursa-pastorts Medic, (shepherd's purse) in Brit, and a cosmop. weed, self-polli- nated. In early spring and lale autumn the sta. are often ± aborted. The 1. vary in shape and degree of division in various situations. Solms (Bat. Zeit. 1900, p. 167) describes C. Heegen. a new form which has an elongated ir., which has arisen from the shepherd's purse, and is almost generically distinct. Capsicum (Tourn. i L. Solanaceae (2). 30 C. and S. Am., i Japan. C. annunm L. cult.; its fr. are chillies or red peppers; dried and ground the v form Cayenne pepper. Other sp. are used, (^ee Irish in Rep. Miss. Rot Gdn., 1898, for revision of cult, forms.) Capsule, a dry dehiscent fruit of > i carpel. Capura Blanco (O/opko'-a Blume, EP}. Sapind. (i). r Phil. Is. Capura L. — Wikstrormia Endl. (Thymel.). Caracasia Szysz. Marcgraviaceae. 2 Venezuela. Caragana Lam. Leguminosae (in. 6). 40 Cent. As., China. Caraguata (Plum.) Lindl. (Gitzmania Ruiz et Pav.). Bromel. (i). 20 S. Am. Cult. orn. infl. Caraguata fibre, Bnwielw, Eryngium, Furcrea, &c. Caraipa Aubl. Guttifer ed of berries. The 1. of C. paltnata R. and P., gathered young, cut into thin strips and bleached, lorni the material of Panama l.ats. Carmenocania Wernham. Rubiaceae (i. 7). i trop. Am. Carmenta Nor. Inc. sed. Nomen. Carmichaelia R. Br. Legum. (in. 6). 20 N.Z., Lord Howe's I. X.. ro. with flat green stems (uhylloclades) and no green 1. (cf. Bo.ssiaea). Carminatia Mo9- Compositae (2). i Mex. Carnarvonia F. Muell. Proieaceae (n). i Queensland. Carnation, Dinnthus Caayophyllus L. Carna-uba, Cope mid a cerifera Mart Carnegiea Britton rt Rose (Lercus p.p.). Cactaceae (ill. i). i Texas ( Cereus gk antetis, q.v.). Carnegiea Perkins Moniiniaceae. i New Caled. Carneus (l.at. ), fi< sh-c<>louied. Carnivorous plants, see Insectivorous. Carnosus (I- at.), Heshy. Carob-tree, Ceratoma Siliqna L. Carolina allspice, Calycanthus ; -jasmine, Gelsenrium. Carolinea L. t. =Pachira Aubl. (BH.}= Bombax L. p.p. Carolinella Hemsl. Primulaceae. 3 China. CarolofritscMa Engl. Gesneriaceae (i). i trop. A^r. Caropodium Mapf. et \\ ettst. Umbel, (in. 5). i Persia. Caroxylon Thunb, Salsola L (Chenopod. ). Carp- (Gr. pref.), iruit ; -el, the megasporophyll of the fl., hearing the ovuU s. In ( .ymnospermae the ovule is exposed, but in Angiospermae the cpl. is infolded, and ihe uvules borne on thickened placentae. Cp s. may be free (apocarpous} or united (syncarpous}, in the latter case the ovary beinj^ uni- or multi-locular. 'I he tip nt the cpl. is the style, ending in the stigma ; -id diminutive of cpl. ; -ophore, Rosaceae, Umbel lijtrue ; -ophyll, carpel ; -OStrote (CL), pi. migrating by means of fr. ; -Otropic, cf. Movements. Carpacoce Sond Rubiaceae (n. 7*. 4 S. Afr. Carpentaria Becc. (Kmtia p.p. £/'). Palmae (iv. i). i New Guin. Carpenteria 1 <>rr. Saxifragaceae (in), i sp. Calif. Like Philadelphus, but ov. sup. ; sta. oo , cpls. 5 — 7. Carpesium L. Compositae (4). 16 S. Eur., As. CARVALHOA 121 Carpet plant, lonopsidium acaule ; -weed (Am.), Mollugo. Carpha Banks et Soland. Cyperaceae (i). 48. temp. Carphalea Ju>s. Kubiaceae (i. 2). i Madag. Carphephorus Cass. Composi'ae (2). 5 E. U.S. Carphobolus Sch-itt= Piptocarpha K. Br. (Comp.) Carpnochaete A. Gray. Compositae (2). 4 S.W. U. S., Mex. Carpinus L. Betulaceae (i). 21 N. temp., chiefly E. As. C. Betitlus L. Brit, (hornbeam). The young 1. hang downwards as the shoot expands. The ? catkins are term, on long shoots, the cT are them- selves short shoots. In the axil of each scale of the latter are 4 — 12 sta each split almost to the base. No bracteoles are present, so that it is doubtlul huw many rls. of the possible 3 (see fam.) are repies. In the ¥ there arc the 2 lat. fls. with all 6 biacieoles. On the top of the 2-loc. ovary is a small P. Fi. a i -seeded nut with a 3-lobed leafy wing on one side, whose centre lobe corresponds to the nract a or /3, the lat. lobes to the bra< teoles a', /3'; these unite and grow large after fert. The timber is little used. Carpoceras Link. = Thlaspi Tourn. (Crucif.). Carp^detus Foist. Saxiiragai eae (v). i New Zealand. Carpodinus R. Br. ex Sabine. Ap<>cyn. (i. i). 50 trop. Afr. Rubber is obtained by grating and boiling from the rhiz. of C. lanceolatus K. Sch. &c. (cf. Clitandra; Bot. Centr. 72, p. 116). Carpodiptera Griseb. Til aceae. 6 E Afr , Cuba. Carpolobia G. Don. Polygalaceae. 6 trop. W. Air. Carpolyza Salisb. (Hessea B^rg.). Amaryllidaceae (i). 4 S. Afr. Carponema Eckl. et Zeyh. Cruciferae (i). 4 S. Afr. Carpopogon Rox >. = Muruna Adans. (Legum.). Carpotroche Endl. Flacourtiaceae (2). 6 trop. Am. Carpoxylon H. Wendl. et Diude. 1'almae (iv. i). i New Hebrides. Carrichtera Adans. Cruciferae (2). i Medit. Carrierea Franch. Flacourtiaceae (4). i China. Carrion-flower (Am.), Smilax herbacea L, ; -flowers, fls. with a smell of carrion, visited esp. by carrion-loving flies, Amorphophallus, Araceae, Stapelia, &c. Carronia F. Muell. Menispermaceae. i New S. Wales. Carrot, Daucns Carola L. Carruthersia Seem. Apocynaceae (n. i). 4 Polynes a. Carsonia Greene (Cleomn p. p.). Cappaiid. (v). i N. Am. Cartagena bark, Cinchona cordifulia Mutis. Carteria Small. Orchidaceae (n. 2) i Florida, Bahamas. Cartnamus (Tourn.) L. Compositae (u). 25 Medit., Afr., As. C. tinctoruis L. (safflouer) cult, in Asia, &c. ; iis fls. are used in dyeing ; powdered and mixed with talc they form rouge. Cartiera Greene (St replant hits p.p.). Cru if (i). 6 N. Am. Cartilaginous, firm and tough ; endosp. of Liliaceac. Cartonema R. Br. Comm Imaceae. 6 irop. Austr. Carum Ru,jp. ex L (Btl. incl. Buniutn L.. f'eti-oseliniini Hoffm.). Umbelliferae (in. 5). 20 sp. ump. and sub-trop. 3 Brit., of which C. Larvi L. is ult. lor its fr. (caraway seeds). Caruncle, a small hard aril, Buxact'ae, Euphorbiaceae. Carvalhoa K. Schum. Apocynaceae (n. i). 2 E. trop. Afr. I 22 CARYA Carya Nutt. Juglandaceae. losp. N. Am., the hickory trees, cultivated for their wood, which is very tough and elastic, and for the edible fruit (pecans, like walnuts). Caryocar Linn. Caryocaraceae. 10 sp. trop. Am. The wood is very durable and is used in ship-building. The fruit is a large 4-seeded drupe ; the seeds are the Souari- or Butter-nuts of commerce. Caryocaraceae (Rhizoboleae)(EP.; Temstroemiaceae p.p. BH.}. Dicots. (Archichl. Parietales). i gen., 15 sp. trop. Am. Trees and shrubs with ternate opp. or alt. 1. with deciduous slips. Fls. $ in racemes. K (5 — 6), C (5 — 6), A oo , united into a ring or in 5 bundles. G 4- or 8 — 2O-loc. with as many styles, i ov. in each loc. Usu. drupe with oily mesocarp, and woody endocarp which splits into 4 meri- carps ; sometimes a leathery schizocarp. Little or no endosp. Genera : Anthodiscus, Caryocar. Caryodendron Karst. Euphorbiaceae (A. n. 2). 2 trop. S. Am. Caryophyllaceae (EP. ; BH. excl. Illecebraceae or Paronychiaceae, and Scleranthaceae]. Dicots. (Archichl. Centrospermae). 60 gen., isoosp. cosmop. (many Brit.), mostly herbs, a few undershrubs, with opp. simple usu. entire 1., often stip. ; the stem often swollen at the nodes, the branching dich. The infl. usu. term, the main axis and is typically a dich. cyme, but both in the veg. region and in the infl., of the two branches arising at any node, one (that in the axil of /3) tends to out- grow the other and after two or three branchings the weaker one often does not develope at all, so that a cincinnus arises. The whole infi. is very char., and such an one is often called a caryophyllaceous infl. Fls. 5 and reg., but often not isomerous. As a type, the formula of Lychnis may serve : K (5), C 5, A 5 + 5, G (5), with free central plac., uniloc. Ov. usu. ao , in double rows corresponding to the Floral diagrams of (i) Silene in_flata and (2) Paronychia sfi. (after Eichler), showing the ordinary type of fl. in Silenoideae and the most reduced type of Alsinoideae; a, |3 = bracteoles. cpls. , rarely few or i (Paronychieae), usu. campylotropous. In most cases the fl. is obdiplost. as may be recognised by the cpls. (when 5) being opp. the petals. Frequently, reduction of the number of parts occurs, e.g. G (3) or (2) or rarely (4); A 4 + 4, or 5, 3, 2, or i, and in other cases the C may abort (Sagina sp., Herniaria, &c.). The ovary, sta., and corolla are sometimes borne on an androphore CARYOPITYS 123 (e.g. Lychnis), an elongation of the axis between K and C. The petals sometimes have a ligule (e.g. Lychnis), and are often bifid. At the base of the ovary are often seen traces of the septa, which in the upper part do not develope ; in some cases the plac. is basal. Biologically, as well as morphologically, the fam. forms two distinct groups, a higher type, the Silenoideae, and a lower, the Alst- noideae. All secrete honey at the base of the sta., but while in the A. the fl. is wide open, so that short-tongued insects can reach the honey, in the S. a tube is formed by the gamosepalous K ; in this stand the claws of the petals and the sta., partly filling it up, and rendering the honey inaccessible to any but long-tongued insects, esp. bees and Lepidoptera. The latter class, esp. in the Alps (see Miiller's Alpen- btumen), are the chief visitors, and many of the S. are adapted to them —by length of tube, red and white colours, night-flowering in many sp., or emission of scent only at night, &c. The fls. are commonly protandr. Many A. are gynodioec. (cf. Labiatae). Fr. usu. a caps, containing several or oo seeds. It opens in nearly all cases by splitting from the apex into teeth which bend outwards, leaving an opening." The splitting may take place in as many, or in twice as many, lines as cpls. The seeds cannot escape from the capsule unless it be shaken, e.g. by wind or animals, and being small and light have a good chance of distr. Embryo usu. curved round the perisperm (in a few cases nearly straight). Classification and chief genera (after Pax) : I. ALSINOIDEAE (flr. polysepalous ; sta. often perig.). a. Fruit a capsule opening by teeth. 1. Alsineae (styles free to base ; 1. exstip.) : Stellaria, Ceras- tium, Sagina, Arenaria. 2. Spergiileae (do., but 1. stip.) : Spergula, Spergularia. 3. Polycarpeae (styles joined at base) : Drymaria, Polycarpon. b. Fruit an achene or nut. 4. Paronychieae (fls. all alike ; stipules) : Corrigiola, Paro- nychia, Illecebrum, Herniaria. 5. Dysphanieac (do., but 1. exstip. alt.) : Dysphania. 6. Scleranthcae (do., exstip. opp.) : Scleranthus. 7. Pterantheae (fls. in 3's, the 2 lat. ± abortive) : Pteranthus. II. SILENOIDEAE (fl. gamosepalous, hypog.) : 1. Lychnideae (calyx with commissural ribs) : Silene, Lychnis. 2. Diantheae (no commissural ribs) : Gypsophila, Dianthus. BH. separate off groups I, 4 — 7 as an independent fam. Illecebra- ceae (Monochlam. Curvembryae), retaining the rest (Caryophylleae) in Polypetalae Caryophyllinae. This is an unnatural separation of closely allied groups. See discussion of relationships of these tarns, in Nat. Pft. (Cary"phyllaceae, p. 68). Caryophyllata Tourn. = Geum Tourn. (Rosac.). Caryophyllatus (Lat.), with long claw. Caryophylleae (J3H.}. See above. Caryophyllinae (BH.}. The 41)1 cohort of Polypetalae. Caryophyllus L. = Eugenia L. (//ZT.)=Jambosa DC. p.p. (Myrt.). Caryopitys Small (Pinus p.p.). Pinaceae. 2 N. Am. i24 CARYOPSIS Caryopsis, achene with pericarp and testa united, Gramineae. Caryopteris Bungi . Verbenaceae (5). 5 Himal. to Japan. Caryospermum Blume (PerrotMia p.p. EP.). Celas't. 3 Indomal. Caryota L. Palmae (iv. i). 10 Indomal. Stem columnar; 1. bi- piunate. Infl. of a number of equal branches hanging down like a brush. They appear in descending order, the oldest in the ciown, the younger lower down in the axils of the old leaf-sheaths. Fls. in groups of 3, one ¥ between two rn, but Loth types oi fl. occur on one plant. It «ould appear to be simply a case ol variation in symmetry (cf. Exacum, Saintpaulia). In many sp. a division of labour takes place among the sta. (cf. Heeria); the insect visitors eat the pollen of the short sta. and carry away on their bodies that ot the long. I here is no honey. Fr. often chambered up by -false' sepia rum i g across it — outgrowths from the placenta. Many cult, for the 1.. which when dried form the drug senna. Alexandrian senna from C. aciitilotia Delile, Italian C. obovata Collad., Arabian C. anguslifolia Vahl. C. Fistula L. (purg ng Cassia, pudding pipe tree) has its setds embedded in laxative pulp. Cassia bark, Cinnaniotnum Cassia Blume ; -broom, Cassia. CASUARINA 125 Cassida Tourn. ex Adans. = Scutellaria Riv. (Labial.). Cassidispermum Hemsl. Sapotaceae. i Solomon Is. Cassie flowers, Acacia Farnesiana Willd. Cassine L. Ceiastraceae. 40 S. Afr., Madag. C. crocea Presl yields saffron-wood. Cassinia R. Br. Compositae (4). 20 S. Afr , Austr., N.Z. Cassinopsis Sond. Icacinaceae (Olacin. Bff.}. 4 S. Afr., Madag. Cassiope D. Don. Eric. (11. i). 7 boreal. L. much rolled back (see fam ; cf. Empetrum); in C'. Redo%vskii G. Don it is hollow. Cassipourea Aulil. Rh zophoraceae. 10 trop. Am. Cassupa Humb. et Bonpl. Rubiphily as in Acacia sphaerocephala (q.v.}. These ants protect the C. from the leaf-cutter ants. The internodes are hollow but do not communicate directly with the air. Near the top of each however is a thin place in the wall. A gravid ? ant burrows through this and brings up her brood inside the stem. The base of the leaf-stalk is swollen and hears lood bodies (cf. Acacia) on the lower side, upon which the ants feed. New ones form as the old are eaten. Several other sp. show similar features. An interesting point, that goes to show the adaptive nature of these phenomena, is that in one sp. the stem is covered with wax which prevents the leaf- cutters fn m climbing up, and there are neither food-bodies nor the thin places in the internodes. Cedar, < ed> ela, Ceitnts, 7'oona, &c. ; Atlantic, Cednts atlnntica Manetti; Australian red-, Toona; bastard-, Chicki-assia (W. I.), Giiazunia tomentosa H. B. K. ; '&erm\id.&-,JiinJfeinsbei»nid'ana\J.; Japanese-, Cr\ptonieria\ of Lebanon, Cfdrns; Oregon-, Cttpressiis Lawson/anah.. Murr. ; pencil-, Ju >n ferns; r^-.Junipei us: Siberian-, Pimti Ceinbra L. ; W. ndian-, Cedrela; white-, Chamaecyparis, Chtckrassia, l.ibocedms', yellow-, Chainaecyfaris; -wood, I oona. Cedrela P. Bi. (BH. incl Toona, q.v.}. Meliaceae. 100 trop. Am. Many yield valuable timber, e.g. C. odorata L. , the West Indian Cedar, used in cigar-boxes. Cedrelopsis Baill. Meliaceae (i). i Madng. Cedronella Riv. Labiatae (vi). i Canaries. Madeira. Cedrus ( I our '.) Mill, i on i ferae (Pinaceae; see C. for gen char.). 3, C. I ibani Barrel. (Cedar 01 Lebanon), C. a lantica Manetti (Atlantic Cedar; Algeria) and C. Deo^ara Loud. (Deodar; Him.d., gre- garious, and reaching to 40 ft in girth) ; all probably vars. of one sp. Handsome evergreen tret-s (often planted for orn.) with needle 1 and long and short shoots ; the latter m;iy grow for several years and even dtvelope into long shoots. Fls. sol , in the position of short shoots. The cone ripens in 2 — 3 years. Wood durable and valued for building, &c. Ceiba Gaertn. (Bomhax L. p.p.; Eriolendron DC. EP ). Bombac. 10 trop. Am. C. penlandra Gaertn. is the silk-cotton (cf. Erio- d end ion). Celandine. Ranunculus Ficaria L. ; greater-, Chelidoninm majiis L. ; W. Indian, Bouonia. Celastraceae (£/'., A'//.). Dicots. (Archichl. Sapindales; Celastrales BH.}. 38 gen. with 280 sp., trop. and temp. Trees or shrubs with simple, often lenthery, 1. and cymose (rarely racemose) infl. Kl. small, reg., usu. ? . K 4 — 5, free or united, C4 -5. There is usu. a well marked disc, on the upper side or edge of which are borne 4 — 5 strt. G (2 — 5), usu. with as many loculi, sometimes partly sunk in the disc. Ovules generally i in each loc., usu. erect, anatr. or apoir. Fr. a loculic. caps., samara, drupe, berry or indehi. caps. Seed usu. with brightly Coloured aril. Endosp. usu. present. C hi ej genera: Euonymus, Celastrus. Cassine. Celastrales (BH.}. The Qth cohort of Polypetalae. Celastrineae (BH.} = Celastraceae. CENTRA NTHERA 1 2 9 Celastrus L. Celastraceae. 30 trop. and suhtrop. Climbing shrubs with fruit like Eiionymus. Celebnia Nor. Inc. sed. Nomen. Celeriac, A pi it in g'-avtolens L. var. rapacfiini. Celery, Af>ium graveolens L. ; -pine, Phyltocladus. Cellulose, the carbohydrate of which cell walls are composed. Celmisia Cass. Compositae (3). 27 N.Z., Austr., &c. Celome Greene (Cleome L. p.p.). Capparid. (v). i N. Am. Celosia L. Amarantaceae (i). 35 trop. and temp., most interesting C. cristata L., the cock's-comb, a cult, (but now hereditary) mon- strosity, in which fasciation of the fls. of the infl. occurs. Celsa Veil. Zygophyllaceae. i Brazil. Celsia L. Scrophulariaceae (i. i). 40 Medit, Afr., As. Celtidaceae = Ulmaceae p.p. Celtis Tourn. Ulmaceae. 75 *. Like Ulmus, but with intr. anthers, drupe, and curved embryo. Fr. of nettle-tree (C. australis L.) ed.; wood useful for turning ; tree used as (odder in India. Cenarrlienes Labill. Proteaceae (i). i Tasmania. Cenchropsis Nash (Cenchnts p.p.). Gram. (5). i N. Am. Cenchrus L. Gramineae (5). 15 trop. and warm temp. Spikelet sur- rounded by invol. of sterile spikelets, which in some sp. become hard and prickly, surrounding the fr. and acting as a means of distribution by animals (cf. Tribulus, &c.). C. tribuloitles L. is a very troublesome pest in the wool-growing districts of N. Am. Cenia Comm. ex Juss. (CotiiLi p.p. EP.). Compositae (7). 9 S. Afr. Cenocentrum Gagnep. Malvaceae, i Indochina. Cenolophium Koch (Sdinuin p.p. BH.}. Umbel, (in. 5). i Eur., As. Cenostigma Tul. Leguminosae (n. 8). 3 Brazil, Paraguay. Censer-mechanism, Aconititm, and cf. Seed-dispersal. Centaurea L. Compositae (n). 600 cosmop., chiefly Medit. ; several Brit. e.g. C. nigra L. (knapweed), C. Scabiosa L., C. Cyanus L. (blue-bottle or cornflower). In the last two the outer fls. are neuter with enlarged C (cf. Hydrangea). C. Calritrapa L. (star-thistle) has long spiny invol. br. The fl. of C. shows the usual construction but the sta. are sensitive to contact and when touched (e.g. by insects probing) contract, thus forcing out the pollen at the top of tube. In C. montana L. and others there is a nectary on each br. of the invol. Numbers of ants are thus attracted. Centaurium Gilib. (Erythraea Borkh.). Gent. (i). 30 N. Am. Centaurodendron Johow. Compositae (n). i Juan Fernandez. Centauropsis Boj. Compositae (t). 3 Madagascar. Centaury, Erythraea Centaurium Pers. Centella L. (Hydrocotyle p.p. BH.}. Umbel, (i. i). 20 S. Afr. to As. Centema Hook. f. Amarantaceae (2). 5 trop. Afr. Centemopsis Schinz. Amarantaceae (2). 3 S. and trop. Afr. Centipeda Lour. Compositae (7). 5 Chili, Madag. , trop. As., Austr. Centotheca Desv. Gramineae (jo). 3 trop. As., Afr. Centradenia G. Don. Melastomaceae (i). 4 Mexico, Cent. Am. C. rosea Lindl. shows habitual anisophylly. Centradeniastrum Cogn. Melastomaceae (i). i Peru. Centranthera R. Br. Scrophular. (in. 2). 5 trop. As., China, Austr. w. y r3o CENTRANTHEROPSIS Centrantheropsis Boniti. Scrophular. (in. 2). i China. Centranthus DC. Valerianaceae. 12 Medit., Eur. C. rub>r DC. (red spur-valerian) cult. orn. C spurred at the base ; at the end of the spur honey is secre'ed. The tube of the C has a partition dividing it into two, one containing the style, the other, lined with downward- pointing hairs, leading to the spur. Fl. protandr. ; only long-tongued insects can obtain honey. Centratherum Cass. Compositae (i). 15 trop. Centric (1.), circular in section, with tissues distr. evenly all round, Alii tun, Bobartia, Eleocharis,Juncus, Littorella. Centrifugal, away from centre; -petal, towards centre. Gen'rilla Lindau. Acanthaceae (iv. B). i Cuba. Centrocarpha D. Don = Rudbeckia L. p.p. (Compos.). Centroglossa Barb. Rodr. (Zygostates Lindl. EP.). Orchidaceae (n. 19) 5 Brazil, Paraguay. Centrolepidaceae (EP., BH.). Monocotyledons (Farinosae; Glumaceae BH.). 6 gen. ,32 sp. Austr., N.Z., S. Am., Polynes., S.E. As. Small grass-like herbs with spikes of small fls. , $ or unisex., naked or with i — 3 hair-structures round them. A i — 2, G i — oo , each with one pend. orthotr. ov. Chief genus: Centrolepis. Centrolepis Lahill. Centrolepidaceae. 20 Austr., E. As. Centrolobium Mart. Leguminosae (ill. 7). 3 trop. Am. Pod winged. C. robiistum Mart, yields good timber (zebra wood). Centromadia Greene (Hemizonia p-p.). Compos. (5). 5 Calif. Centronia D. Don. Melastomaceae (i). 15 trop. Am. Centropetalum Lindl. Orchidaceae (n. 20). 4 Andes. Centroplacus Pierre. Euphorbiaceae (A. I. i). i trop. Afr. Centropogon Presl. Campanulaceae (in). 90 trop. Am., W.I. Centrosema Benth. (Bradbnrya EP.}. Legum. (ill. 10). 30 Am. Centrospennae. The ijth order of Dicots. (Archichl.). Cen'rostegia A. Gray. Polygonaceae (i. i). 2 California. Centrostigma Schlechter. Orchid, (n. i). 3 Nyassaland. Cen'unculus Dill, ex L. Primul. 3 temp, and subtrop., i Brit. Century, set of 100 dried plants. Century plant, Agave americana L. Cepa (Tourn.) L. = Allium Tourn. (Lili.). Cephaelis Sw. (Uragoga L. EP.). Rubiaceae (n. 5). 130 trop., esp. Brazil. For ipecacuanha cf. Uragoga. Cephal- (Gr. pref. ), head. Cephalacantnus I indau. Acanthaceae (iv. B). i Peru. Cephalandra Schrad. = Coccinia Wight et Arn. (Cucurb.). Cephalanthera Rich Orchidaceae (n. 2). 10 N. temp., 3 Brit. No rostellum ; the pollen germinates in situ, fertilising its own stigma (Darwin, Orchids, p. 80). The lat. stds (see fam.) are easily seen. Darwin regards C. as a degraded Epipactis (cf. Cephalopipactis). Cephalanthus L. Rubiaceae (I. 6). 8 warm countries. Cephalaralia Harms. Araliaceae (2). i Austr. Cephalaria Schrad. Dipsacaceae. 35 Medit., Afr. Cephalipterum A. Gray. Compositae (4). i S. and W. Austr. Cephalobembix Rydberg. Compositae (6). i Mexico. Cephalocarpus Nees. Cyper. (n). i Brazil. Habit of Dracaena. CERA TANDROPSIS 1 3 1 Cephalocereus Pfeiff. (em. K. Schum.; Cereus Mill. p.p. BH,}. Cactaceae (in. i). 24 Brazil, Mexico. Cephalocroton Hochst. Euphorh. (A. n. 4). 6 trop. Afr., Madag. Cephalocrotonopsis Pax (preceding, pp.). Euph. (A. n. 4). i Socotra. Cephalomappa Baill. Euphorbiaceae (A. n. 2). i Borneo. Cephalomedinilla Merrill. Melastomaceae (i). i Phil. Is. Cephalonema K. Schum. Tiliaceae. i trop. Afr. Cephalopappus Nees et Mart. Compositae (12). i Bahia. Cephalophilum Bonier (Polygoniini p.p.)- Polyg. (n. 2). i N. Am., N. As. Cephalophora Cav. (Helenium p.p. EP.}. Compos. (6). 12 temp. S. Am. Cephalopipactis x Aschers. et Graebn. Orchid. Hybrid between Cephalanthera and Epipactis. Cephalosphaera Warb. (Brochoneura p.p.). Myristic. i trop. Afr. Cephalostachyum Munro. Gramineae (13). 8 Indomal., Madag. Cephalostemon K. Schomb. Rapateaceae. i Brazil, Guiana. Cephalostigma A. DC. Campanulaceae (i). 7 trop. Cephalotaceae (EP.\ Saxifragaceae p.p. BH.}. Uicots. (Archichl. Kosales). Only genus Cephalotus (q-v.}~ Cephalotaxus Sieb. et Zucc. Coniferae (Taxaceae ; see C. for gen. char.). C. Fortunei Hook. cult. orn. shrub. All shoots of unlimited growth. Fls. dioec., the 35 "sp., trop. and subtrop. Herbs, shrubs, or trees, with opp. stip. 1. Fls. small, in spikes or cymes, 5 or unisex , sometimes with sepaloid P; A i — 3, united to one another and to ovary; 1 4 2 CHL ORA NTH A CEAE G i ; ov. few, pend., orthotr. Endosp. oily; no perispenn ; embryo minute. Chief genera : Chloranthus, Hedyo.smum. Chloranthus Sw. Chlorantli. ro E. As., E. Ind. P i, anterior; the centre sta. has a complete anther, the lat. each half (cf. Fumaria). Chloridion Stapf. Gramineae (5). i trop. Afr. Chloris Svv. Gramineae (iij. 40 trop. and warm temp. Several are useful pasture-grasses in Austr., &c. Chloro- (Gr. pref.), green, yellow; -phyll, the green colouring matter of 1. ; -in the fl. Deherainia ; -plastids, carriers of. Chlorocodon Hook. f. Asclepiadaceae (i). 2 E. and S.E. Afr. CMorocrambe Rydberg (Canlanthus p.p.). Cruc. (i). i N. Am. Chlorocyathus Oliv. Asclepiadaceae (i). i Delagoa Bay. Chlorocyperus Rikli = Cyperus L. (Cyperac.). Chlorogalum Kunth. Liliaceae(m). 3 Calif. C. poineridianum Kunth has a large bulb whose inner parts are used as a substitute for soap (cf. Saponaria). The outer layers yield a quantity of fibre. Chloromyrtus Pierre (Ettgema p.p. EP}. Myrt. (i). i trop. Afr. Chloropatane Engl. Mommiaceae. 2 trop. Afr. Chlorophora Gaudich. Moraceae (i). 3 W. Afr., trop. Am. The wood of the latter (C. tinctoria Gaudich.) forms the yellow dye fustic. Chlorophytum Ker-Gawl. Liliaceae (in). 50 trop. In C. comosum Baker infl. often replaced by veg. repr. ; long shoots develope in the axils of the br., weigh the stem down to the soil and take root. Cult. orn. Chloropyron Behr. = Cordylanthus Nutt. (Scroph.). Clilorosa Blume. Orchidaceae (n. 2). t Java. Chlorospatha Engl. Araceae (vi). i Colombia. Chloroxylon Rumph. Rutaceae (in) (Meliaceae, BH.}. i E. Ind., C. S-wietenia DC. (satinwood). Timber very lasting, largely used in veneering. The tree also yields a gum. Chloroxylum P. Br. Inc. sed. Quid? Chloryllis E. Mey. (Dolichos p.p. BH.}. Legum. (in. jo), i S. Afr. Choananthus Rendle. Amaryllidaceae (i). 2 Ruwenzori. Chocho, Sechium edule Sw. Chocolate, Theobroina. Chodanthus Hassler (Adenocalymma p.p.). Bign. (i). j Braz. Choisya H. B. et K. Rutaceae (i). i Mexico. Cult. orn. shrub. Choke-berry (Am.), Pyrus arbutifolia L. Chomelia Jacq., non L. (Anisorneris Presl EP.}. Rubiaceae (u. 2). 30 trop. S. Am., Afr. Chomelia L., non Jacq. ( Tare/tnaGaertn. BH.}. Rubi. (i. 8). 30 trop. As., Afr. Chondilophyllum Panch. ex Guillaumin (Meryta Forst. p.p.). Arali- aceae. i New Cale> Ionia. Chondodendron Ruiz et Pav. Menispermaceae. 10 Brazil, Peru, trop. Afr. C. tomentosum R. P. furnishes Radix Pareirae bravae. Chondrilla (Tourn.) L. Compositae (13). 20 N. temp. Chondrobollea x Hort. Orchidaceae. Hybrid of Chondrorhyncha and Bollea. Also Chondropetalum x Hort. Hybrid with Zygo- petalum. Chondrophylla A. Nelson (Gentiana p.p.). Gentian, (i). 2 N. Am. CHR OZOPHORA 1 43 Chondrorrhyncha Lindl. Orchid, (n. ft. n). 2 Colombia. Cult. Ciiondrosea Haw. — Saxifraga Tourn. p.p. (Saxitr.). Chondrostylis Boerlage. Euphorb. (A. II. 2). i Malaya. Chondrosum Desv. = Bouteloua Lag. p.p. (Gramin.). Chonemorpha G. Don. Apocynaceae (n. i). 2 Indomal. Chordospartium Cheesem. Leguminosae (in. 6). i N.Z. Choretrum R. Br. Santalaceae. 5 Austr. Chorilaena Endl. Rutaceae (I). 3 W. Austr. Chorilepis Van Tiegh. (Loranthus p.p.). Lorantli. (i). 3 Malaya, Phil. Is. Chorioluma Baill. (Sideroxylon p.p. EP.). Sapot. (i). i New Caled. Choriophyllum Benth. Euphorb. (A. I. r). 2 Malay Arch. Choripetalae (Warming) = Archichlamydeae. Choripetalous, polypetalous. Chorisia H. B. et K. Bombacaceae. 5 S. Am. C. speciosa St Hil. (paina de seda) gives a useful silky cotton from the pods. Chorisis, branching in floral organs. Chorispora R. Br. Cruciferae (4). 12 E. Medit., Centr. As. Choristega Van Tiegh. (Loranthus p.p.). Loranth. (i). 2 Celebes. Choristegeres Van Tiegh. (ditto). Loranth. (i). i Borneo. Choristigma F. Kurtz. Asclepiadaceae (n. i). i Argentina. Choristylis Harv. Saxifragaceae (v). i S. Afr. Cboritaenia Benth. (Pappea Sond. et Harv. EP.}. Umbelliferae (in. 6). i S. Afr. Chorizandra R. Br. Cyperaceae (n). 4 Austr. Chorizanthe R. Br. Polygon, (i. i). 35 Am. Some have an ochrea, usu. absent in this group. .Els. usu. single inside the invol. (cf. Eriogonum). Cborizema Labill. Leguminosae (ill. 2). 15 Austr. Chortolirion Berger (Haworthia p p.). Lili. (in). 4 Afr. Chowlee (India), Vigna Catjang \Va\p. Christ's thorn, Paliurus aculeatus Lam. Christensenia Maxon. Marattiaceae. i Phil. Is. Christiana DC. Tiliaceae. i Madag. to Guiana. Christisonia Gardn. Orobanchaceae. 10 trop. As. Roots parasitic on those of bamboos or Acanthaceae, united to a dense meshwork. The fig. shoots spring up, die, and decay, in a fortnight. Christmannia Dennst. Inc. sed. i E. Indies. Christmas gambol (W.I.), Ipomoea sidifolia Choisy; -pride (W.I.), Ruellia paniculata L ; -rose, Helleborus niger L. Christolea Cambess. Cruciferae (4). 2 W. and C. As. Christophine (W.I.), Sechium edule Sw. Christopteris Copeland. Polypodiaceae. 2 S.E. As. Chroilema Bernh. Compositae (3). i Chili. Chromanthus Phil. Portulacaceae. i Chili. Chromolepis Benth. Compositae (5). i Mexico. Chromoplastids, carriers of colouring matters. Chronopappus DC. Compositae (i). i Minas Geraes. Chrozophora Neck. Euphorbiaceae (A. n. 2). 10 Medit., W. As., N. Afr. C. tinctoria A. Juss. and C- verbascifolia Juss. are charac- teristic plants of the Medit. region. The former, once medicinal, is i44 CHROZOPHORA still sometimes used as the source of the dye turn-sole, tournesol, or bezetta rubra. Chrysactiuia A. Gray. Compositae (6). 3 Mex., S.W. U.S. Chrysalidocarpus H. Wendl. (Hvophorbe p.p. EP.). Palmae (iv. i). i Madag., C. lutescens H. Wendl. a favourite orn. palm, branching at the r. and forming tufts of stems. Chrysanthellum L C. Rich. Compositae (5). 4 trop. Chrysanthemum (Tourn ) L. (incl. Pyretkrum Hall.). Compositae (7). 150 Kur., As., Afr., Am. C. segetuin L. (corn-marigold) and C. Leu- canthemum L. (ox-eye or dog daisy) Brit. The autumn-flowering C. are cult, forms of C. indicnm L. and C. sinense Sabine (China, Japan). As in Dahlia, all florets have become ligulate (Hemsley in Card. Chron. 1889, p. 521, &c. ; Henry in Card. Citron., 1902, p. 301, and discussion by Hooker in Curtis, Bot. Mcale as to reduce the price of quinine from 12s. to is. an ounce. Decrease in price, the lack of any improvement in 'he barks, and attacks of disease, made the cult, die out in Ceylon, and lava, where improvement was taken in hand, now almost mono- polises it. India grows a good deal for supply to natives through the post offices. Several sp. are used, e.g. C. Calisaya Wedd. (yellow, and some crown, bark), C. Ledgeriana Moens (yellow bark, the richest in alkaloid), C. cordifolia Mutis (Cartagena bark), C. officinalis L. (condamined H. & B. ) (Loxa, crown or brown bark), C. siucinibra W. 10 146 CINCHONA Pav. (red bark). (Markham. Travels in Peru and India ; Reimers, Les quinquinas de culture, 1900.) Cinchonidin, Cinchona. Cincinnobotrys Gilg. Melastomaceae (i). 2 trop. Afr. Cincinnus, a monoch. cyme in which the successive lat. branches fall alt. on either side of the relatively main axis ; Bignomaceae, Boragi- naceae, Canna, Caryophyllaceae, Chenopodiaceae, Commclinaceae, Crassulaceat, Geranium, HManthemum, Heliconia, Hydrophyllaceae, Hyoscyamtts, Linaceae, Solanaceae, Strelitzia, Tradescantia, Urtica, Verbenaceae. Cineraria L. p.p. Compositae (8). 35 Afr., Madag. Many sp. of Senecio are often included in this genus. Cult. orn. fl. Cinereus (Lat.), ash grey. Cinga Nor. Inc. sed. Nomen. Cinna L. Gramineae (8). 2 N. temp. Cinnagrostis Griseb. Gramineae (S). i Argentina. Cinnamodendron Endl. Winteranaceae. 3 Brazil, W. Ind. Cinnamomeous, light yellowish brown. Cinnamomum (Tourn.) L. Lauraceae (i). 60 Indomal. Young leaves often red. C. zeylatucnm Nees (Ceylon) is the cinnamon. The pi. is coppiced in cult., and the bark of the twigs peeled off and rolled up is the spice. C. Cassia Blume (China, Japan) yields Cassia bark, often used to adulterate cinnamon. Its fl. buds are used as a spice (cf. Eugenia). C. Cnmphora T. Nees & Eberm. (China, Japan, For- mosa) is the camphor. The old trees are felled, and the wood cut into chips and distilled with steam, but in cult, the camphor is distilled from young shoots. Cinnamon, Cinnamomum zeylanicum Nees, (Am.) Canella; wild- (Ceylon), Lilsea zeylanica Nees, (W.I.) Canella alba Murr., Pimcnta acris Kostel. Cinnamosma Baill. Winteranaceae. i Madag. Cinquefoil, Poteniilla reptans L. Cionosicyos Griseb. Cucurbit. (3). i Jamaica. Cipadessa Blume. Meliaceae (in). 4 Indomal., Madag. Cipre (W.I.), Cordia Gerascanthus L. Cipura Aubl. Iridaceae (n). 5 trop. Am. Cipuropsis Ule. Bromeliaceae (i). i Peru. Circaea Tourn. ex L. Ona.nraceae (2). 9 N. temp, and arctic; 2 Brit. (enchanter's nightshade). Fl. dimerous with one whorl sta. Fr. hooked. Circaeaster Maxim. Chloranthaceae. i Himal., China. K2 — 3, C o, A i — 2, G i — 4; fr. hooked. (Hooker, Icones PL, t. 2366.) Circinate (aestivation), 1. rolled up like watch-springs, Filicineae Lep- tospot., Marsileaceae, petals Q{ Hamamelidaceae. Circinus Med. (Hymenocarpits Savi). Legum. (ill. 5). i Medit. Circumscissile, opening by splitting off a lid, Anagallis. Cirrhaea Lindl. Orchidaceae (n. 13). ? Brazil. Cirrhiferous, tendril-bearing : cirrhose, tendrilled. Cirrhopetalum Lindl. Orchidaceae (n. 16). 70 Indomal., Masc. Cirsium (Tourn.) Adans. (Cnictts L. p.p. EP.}. Compositae (u). 200 N. temp. Cissampelos L. Menispermaceae. 70 trop. and subtrop. f infl. CITRtfS 147 cymose. ? fi. with peculiar zygomorphic structure; cpl. one, witli one sepal and two petals at one side of it. Petals often united. Cissus L. (FiVzV Tourn. BH.}. Vitaceae. 275 trop. Cistaceae (EP., Cistineae BH.}. Dicots. (Archichl. Parietales EP., BH.}. 4 gen., i6osp., in dry sunny places, esp. on chalky or sandy soil, a few in S. Am., the rest N. temp. (esp. Medit. )• Shrubs and herbs with opp. rarely alt. leaves, often inrolled (cf. Ericaceae), with or without stipules. Glandular hairs usu. present. Fls. sol. or in cymose infl., 5 , reg. K 5, the two outer usu. smaller than the inner (sometimes regarded as bracteoles, but these are found lower down) ; C 5 or 3 or o, conv. (the petals overlap to right or left according as the 3 inner sepals overlap to left or right) ; A QO on a sub-ovarial disc (sta. developed in descending order); G (5 — 10 or 3) i-loc. with parietal (often projecting) plac. ; ov. oo or 2 on each, ascending, orthotr. ; styles free. Caps, loculic. Endosp. ; curved embryo. Genera: Cistus (ovules oo , capsule j-valved), Helianthemum (do., 3-valved), Hudsonia (ovules 2, pets. 5), Lechea (do., pets. 3 or o). Cistanche Hoffmgg. et Link. Orobanchaceae. 12 |^=. Cistanthera K. Schum. Tiliaceae. 3 trop. Afr. Cistiflorae (Warming). The loth cohort of Choripetalae. Cistineae (Bff.) = Cistaceae. Cistula Nor. Inc. sed. Nomen. Cistus (Tourn.) L. Cistaceae. 35 Medit. C. cretictts L. and C. ladanifems L. yield the resin ladanum (not laudanum), formerly offic. Many favourite cool-house shrubs (gum-cistus). Citharella Nor. Inc. sed. Nomen. Cithareloma Bunge. Cruciferae (4). 2 Turkestan. Citharexylum Mill. Verbenaceae (i). 35 warm Am. The common name fiddle-wood is a corruption of Bois-fidele. Citriobatus A. Cunn. Pittosporaceae. 2 S.W. Austr. Citriosma Ruiz et Pav. = Siparuna Aubl. (Monim.). Citron, Citrus medica L. Citronella oil, Cymbopogon Nardus Rendle, C. Winterianus Jowitt. Citropsis Swingle et Kellermann (Lt mania p.p.). Rutaceae (v). 4 trop. Afr. Citrullus Forsk. Cucurbitaceae (3). 4 ;=&. C. vnlgaris Schrad. (water- melon), C. Colocynthis Schrad. (colocynth), fr. a drug. Citrus L. Rutaceae (v). 10 sp. trop. Old World. Shrubs and trees with usu. simple L, which show a joint at the meeting place of blade and stalk, indicating their derivation from cpd. 1. like those of most of the fam. (cf. Berberis). Axillary thorns in some ( = metamorphosed 1. of the branch shoot). Fls. in corymbs, 5 . K and C 4 — 8 ; A oo in irreg. bundles, corresp. to an outer whorl only ; G (oo ) (6 or more). A second whorl sometimes appears. Fr. a berry with leathery epicarp, the flesh made up of large cells which grow out from the inner layer of the pericarp. Many cult, in warm countries, esp. California, the W. Indies, Brazil, the Medit. region, &c., for their fr. C. Medica L., the citron, is the parent sp. of several varieties, e.g. var. Liinonum the lemon, var acida the lime, var. Limefla the sweet lime. C. Aurantium L. is the orange, with its vars. Bergamia, the Bergamot orange (from which the IO 2 i48 CITRUS perfume is obtained), Bigaradia or a/nara the Seville or bitter orange, used in marmalade, C, decumana the shaddock, or pomelo, with its var. the grape-fruit, C. sinensis, the Malta or Portugal orange, C. suntara Engl., the suntara or kumquat, and others. C. nobilis Lour, is the true mandarin orange. Cf. De Candolle, Orig. oj Cult. Pits, p. 176; Engler in Nat. PJi.; Bonavia, Oranges... of India and Ceylon. Cladanthus Cass. Compositae (7). i S. Spain, Morocco. Claderia Hook. f. Orchidaceae (n. =,). i Perak. Cladium P. Br. Cyperaceae (n). 30 trop. and temp., esp. Austr. C. Mariscns R. Br. (gernianicuin Schrad.), Brit. Cladode, phylloclade of one. internode. Cladogynos Zipp. ex Span. Euphorb. (A. II. 2). i Malay Arch. Cladopus Moller. Podostemaceae. i Java. Cladostemon A. Br. et Vatke. Capparid. (ll). i Zanzibar. Cladostigma Radlk. Convolvul. (i). i Abyssinia. Ed. fr. Cladothamnus Brongn. Ericaceae (i. i). i N.W. N. Am. Cladothrix Nutt. Amarantaceae (3). 2 W. N.Am. Cladrastis Rafin. Leguminosae (in. i). i E.As., i E. N.Am. (cf. Epi- gaea) (C. tinctoria Kami., yellow-wood) ; its wood yields a yellow dye. Clambus Miers. Menispermaceae. i Mexico. Claoxylon A. Juss. Euphorbiaceae (A. n. 2). 60 palaeotrop. Clappia A. Gray. Compositae (6). 2 Texas, Mexico. Clarionea Lag. = Perez i a Lag. p.p. (Compos.). Clarisia Ruiz et Pav. Moraceae (n). 4 Peru, Brazil. Clarkeifedia Kuntze (Patrmia p.p.). Valer. i Himalaya. Clarkella Hook. f. Rubiaceae (i. 2). i Himalaya. Clarkia Pursh. Onagraceae (2). 8 W. N.Am. Cult. orn. fl. Mech. of fl. as in Epilobium. Clary, Salvia pratensis L. ; wild- (W. I.), Heliotr opium. Clastopus Bunge ex Boiss. (Vesicaria BH.). Cruc. (4). 3 Persia. Clathrate, latticed. Clathrospermum Planch. (Popo-wia EP.}. Anon. (i). 12 trop. Afr. Clathrotropis Harms (Diplotropis p.p.). Legum. (III. i). 2 Braz. Clausena Burm. f. Rutaceae (v). 20 palaeotrop. Some ed. fr. Clausia Trotzky (Hesperis BH.). Crucif. (4). 5 C. and N. As. Clavapetalum Pulle (Plalea p.p.). Icacinaceae. i Surinam. Clavate, club-shaped. Clavija Ruiz et Pav. Theophrastaceae. 40 trop. Am. Trees of palm- like habit, often with fls. on the old wood (cauliflory). Clavipodiurn Desv. ex Griming. Euphorb. (B. n). i Austr. Clavistylus J. J. Smith. Euphorb. (A. n. 2). i Java. Claw, a narrowed base of a petal, Cheiranihus, Caryopliyllac. Claytonia Gronov. ex L. Portulacaceae. 24 N. temp, and arctic ; 2 nat. in Brit. No slips. Fls. in sympodial cymes. Before pollin. the fl.-stalk is erect; fl. protandr., with outward movement of the sta. after dehisc. Honey, at base of each petal, accessible to short- tongued insects. After pollin., the stalk bends down through 180°, to return once more to the erect position when fr. ripe. The caps, contains 3 seeds and splits into 3 valves, the seeds lying across the lines of splitting. The inner surfaces of the valves contract as they dry and shoot out the seeds (cf. Buxus, Viola). CLETHRA 149 Cleanthe Salisb. Iridaceae (li). i Cape Colony. Cleanthes D. Don (Trixis BH.). Compos. (12). 3 S. Braz., Arg. Clearing-nut, Slrychnos potatorum L. f. Clearweed (Am.), Pi lea. Cleavers, Galium Aparine L. Cleft, cut halfway down. Cleghornia Wight (Baissea BH.}. Apocyn. (n. i). 4 Indomal. Cleidion Blume. Euphorbiaceae (A. n. 2). 18 trop. Cleisocratera Korth. (Psycholria p.p. EP.}. Rubi. (n. 5). i Borneo. Cleisostoma Blume. Orchidaceae (n. 20). 20 Indomal. Cleistachne Benth. Gramineae (2). 2 trop. As., Afr. Cleistanthus Hook. f. ex Planch. Euphorb. (A. I. 2). 40 palaeotrop. Cleistes L. C. Rich. (Pogonia BH.}. Orchid, (n. 2). 12 S. Am. Cleistochlamys Oliv. Anonaceae (i). i Mozambique. Cleistogamy, the production of closed self-pollinating fl., Amphicarfaea, Carda/iniie, Comiitelina, Halcnia, Lanihtm, Leersia, Lespedeza, Mal- pig/n'a, Montia, Otwiris, Oxalis, Paroc/ietus, Viola, esp. the last. Cleistoloranthus Merrill. Loranth. (i). j Phil. Is. Cleistopholis Pierre. Anonaceae (i). 5 trop. Afr. Clematicissus Planch. (Vitis p.p.). Vitaceae. i W. Austr. Clematis Dill, ex L. (incl. Atragene L.). Ranunculaceae (3). 220 cosmop. C. vitalba L. (traveller's joy) Brit. Mostly climbing shrubs with opp., usu. cpd., 1. Lower sides of petioles sensitive to contact. The petiole bends once round the support, thickens and lignifies. Fls. in cymes ; K coloured ; no pets, or honey secretion. The style often remains persistent upon the fr. and becomes hairy', thus forming a mech. for wind-distr. Clematoclethra Maxim. Dillen. (Ternstr. BH.}. 10 China. Clemensia Merrill. Meliaceae (n). i Phil. Is. Clementsia Rose (Sednm p.p.). Crassulaceae. i Rocky Mts. Cleobula Veil. Inc. sed. i Brazil. Cleobulia Mart. Leguminosae (in. 10). 3 Brazil. Cleome L. Cappariclaceae (v). 70 trop., subtrop. Disc usu. more developed on post, side ; may bear scales. Gynophore varies in length. Cleomella DC. Capparidaceae (v). 10 N. Am. Cleomodendron Pax. Capparidaceae (inc. sed.). i Somaliland. Cleonia L. Labiatae (vi). i W. Medit. Clermontia Gaudich. Campanulaceae (in). 12 Sandwich Is. The latex is used as bird lime. Some have ed. fr. Clerodendron L. Verbenaceae (4). 150 trop , subtrop. C. Thompsonae Balf., often cult. orn. fl., has red K and white C. The sta. project so as to form the landing place for insects, and when they are ripe the style is bent down. Afterwards the sta. roll up and the style takes their place. C. fistulosum Becc. has hollow internodes inhabited by ants (myrmecophily, ^.r'.). Clethra Gronov. Clethraceae. 12 trop., subtrop. Shrubs and trees with alt. 1.; fls. in racemes or panicles, without bracteoles, ?, reg. K 5, C 5, polypet. ; A 5 + 5, hypog. ; no disc ; anthers bent outwards in bud ; pollen in single grains ; ov. 3-loc. ; style with 3 stigmas. Caps. ; endosp. 1 5 o CLE THRA CEAE Clethraceae ( EP. ; Ericaceae p.p. BH.}. Dicots. (Sympet. Ericales). Only grnus Clethra, q.v. Clevelandia Greene ex Brandegee. Scroph. (in. 3). i Calif. Cleyera DC. (Eurya Thunb. p.p. EP.}. Theac. 9 warm Am., As. Clianthus Banks et Soland. (Donia G. Don). Leguminosae (in. 6). 2 Austr., N.Z. Cult. orn. fl. Clibadium L. Compositae (5). 1=, trop. Am., W.I. Clidemia D. Don (incl. Sagraea EP.}. Melastom. (i). 100 trop. Am. Ed. fr. Cliff-brake (Am.), Pellaea. Cliffortia L. Rosaceae (in. 5). 50 S. Afr. Cliftonia Banks et Gaertn. f. " Cyrillaceae. i S.E. U.S. Climacorachis Hemsl ,et Rose. Leguminosae (in 7). 2 Mex. Climbing fern (Am.), Lygodinm; -plants abound in trop. forests, where they usu. grow large and woody (lianes}, and are rarer else- where. Four chief groups: (i) twiners, whose tips nutate in search of support, Apios, Apocynaceae, Araliaceae, Aristolochiaceae, Basella, Bauhinia, Bignoniaceae, Bowiea, Calystegia, Camoensia, Ceropegia, Cassytha, Combretaceae, Connaraceae, Convolvulaceae, Cuscuta (sensitive stems, like tendrils), Cynanchum, Dipladenia, Freycinetia, Gnetum, Hoya, Ipomoea, Jasminum, Lardizabalaceae, Loasaceae, Lonicera, Lygodium, Malpighiaceae, Menispermaceae, Phaseolus, Phytocrene, Plumbago, Polygonum, Rhodochiton, Ruscus, Schi- zandra, Solanum, Tamus, Thunbergia, Wistaria; (2) climbers with sensitive organs, usu. tendrils > which may be modified stems, Antigonon, Landolphia, Passiflora, Vitis, leaves, Bignoniaceae, Cucurbitaceae, Cobaea, Corydalis, Leguminosae (Lathyrus, Vicia, &c.), Mutisia, sensitive hooks which clasp and become woody, Ancistrocladus, Artabotrys, Bauhinia, Gouania, Husjonia, Lan- dolphia, Paullinia, Strychno*, Uncaria, Unona, Uvaria ; sensitive I. occur in Gloriosa, Littonia, &c., petioles in Clematis, Dalbergia, Fumaria, Hablitzia, Maurandia, Rhodochiton, Tropaeolum, midrib in Nepenthes, /at, branches in Hippocratea, Macherium, Salacia, Securidaca, Uvaria ; (3) hook climbers, sprawling, and catching by hooks (cf. above), Caesalpinia, Calamus, Capparis, Combretaceae, Desmoncus, Dipladenia, Galium, Hugonia, Lycium, Pereskia, Plecto- comia, Smilax, Ventilago ; (4) root climbers with special negatively heliotropic adv. r. that adhere to the support, Araceae, Araliaceae, Begonia, Bignoniaceae, Clusia, Ficus, Hedera, Hoya, Kendrickia, Norantea, Piper, Rhus, Salacia, Sapindaceae, Tecoma Climbers are often of anatomical interest, presenting manyabnormal features, esp. the trop. lianes. For details, see Darwin, Climbing Plants; Schenk, Biologie und Anatomie der Lianen. Clinacanthus Nees. Acanthaceae (iv. B). i Malaya. Clinandrium (orchids), anther-bed. Clinogyne Salisb. Marantaceae. 20 trop. As. Afr. Clinopodium L. = Calamintha Tourn. (/?/f.) = Satureia L. p.p. Clinostigma Wendl. Palmaceae (iv. i). 5 Samoa, New Hebrides, &c. Clintonia Dougl. = Downingia Torr. (Campanul.). Clintonia Rafin. Liliaceae (vn). 6 E. As., N. Am. Clistax Mart. . \canthaceae (iv. B). 2 Brazil. CNEORUM 151 Clistoyucca Trel. (Yucca p.p.). Liliaceae (vi). i N. Am. Clitandra, Benth. Apocynaceae (i. i). 20 W. and C. trop. Afr. Rubber is obtained from the r. of C. Henriqueziana K. Sch. Clitoria L. Leguminosae (in. ro). 35 trop. and subtrop. Fls. inverted and the essential organs therefore touch an insect's back. Clivia Lindl. Amaryllidaceae (i). 5 Cape Col. Cult. orn. fl. Cloezia Brongn. et Gris. Myrtaceae (li. i). 6 New Caled. Cloiselia Sp. Moore. Compositae (12). i Madag. Tree. Clomenolepis Cass. Compositae (inc. sed.). Nomen. Clonodia Griseb. Malpighiaceae (n). i N. Brazil. Closia Remy. Compositae (6). 5 Chili. Closing of fl. in shade or cold, Aiiagallis, Bellis, Calandrinia, Eschscholtzia, Paeonia, Tragopogon. Clotbur (Am.), Xanthitim. Cloudberry, Rnbus Chainaeniorns L. Clove gilliflower, -pink, Diantlms Caryophyllns L., var. Clovenberry bush (W.I.), Sainvda serrulata L. Clover, Trifolium ; bush or Japanese-, LespeJeza. Cloves, Eugenia caryophyllata Thunb., (W.I.) Pimenta acns Kostel.; Madagascar-, Ravensara aroniatica Sonn. Clozelia A. Chevalier (Antrocaryon ; Pierre). Anacardiaceae (2). i Ivory Coast. Clubmoss, Lycopodiiini ; -rush, Scirpus. Clusia L. Guttiferae (v). 85 warm Am., mostly climbing epiph., clasping the host by anastomosing aerial r., and frequently strangling it (cf. Ficus). Fr. fleshy probably carried by birds. Clusiaceae (Warming) = Guttiferae. Clusianthemum Vieill. (Garcinia p.p. BH. ). Guttif. (v). 2 New Caled. Clusiella Planch, et Triana. Guttiferae (v). i Colombia. Cluster bean. Cya»iopsis\ -pine, Pinns Pinaster Ait. Cluytia Hoerh. ex L. Euphorbiaceae (A. n. 5). 50 Afi., Arabia. Cluytiandra Muell.-Arg. Euphorbiaceae (A. I. i). 5 trop. Afr. Clybatis Phil. (Leitceria p p. EP.}. Compos. (12). i Chili. Clypea Blume=Stephania Lour. (Menisp.). Clypeate, shield-shaped. Clypeola L. Cruciferae (4). 12 Medit. Clytos oma Miers (Pithtcocteniiim Mart. BH.). Bignoniaceae (i). 2 temp. S. Am. Cult. orn. fl., often under name Bignonia. Cnemidiscus Pierre. Sapindaceae (i). i Cochinchina. Cnemidophacos Rydb. (Astragalus p.p. ). Legum. (in. 6). i N. Am. Cnemidostachys Mart. = Sebastiana Spreng. p.p. (Euph ). Cneoraceae (EP.; Simarubaceae p.p. BH.). Dicots. (Archichl. Gera- niales). Only genus Cneorum, q.v. Near to Zygophyllaceae, but separated because only one whorl of sta. with no ligules, and no stipules, but oil-glands in the I. Cneoridium Hook. f. Rutaceae (i). i S. Calif. Cneorum L. Cneoraceae. 12 Medit., Canaries. Shrubs with alt. leathery exstip. 1. with oil-glands ; fl. sol. or in racemes, 3 — 4-merous, $ , reg. with column or bolster-like disc. A 3 — 4, G (3 — 4), lobed, with i ov. in each; style i. Schizocarp. 1 52 CNESMONE Cnesmone Blume. Euphorbiaceae (A. II. 2). i Indomal. Cnestidium Planch. Connaraceae. i Panama. Cnestis Juss. Connaraceae. 20 trop. Afr., As. Cnicothamnus Griseh. Compositae (12). i Argentina. Cnicus L. p.p. \_BH- and others incl. Cirsium, making 120 j&]. Com- positae (i i). i Medit. (C, Benedictus L., offic.). The genus is much confused with Carduus and Cirsium. Cnidium Cusson (Selimim BH.}. Umbellif. (in. 5). 20 N. palaeo- temp. Cnidoscolus Pohl.=Jatropha L. p.p. (Euphorb.). Coach. whip, Fouguieria splendens Engelm. Coadunate, adnate, connate. Coarctate, crowded together. Coaxana Coulter et Rose. Umbellif. (111.5). J Mexico. Cobaea Cav. Polemoniaceae. 9 trop. Am. C. scandens Cav. cult, orn. climber of very rapid growth. It climbs by aid of tendrils (leaf-structures) which are much branched, the branches ending in sharp hooks. The tendril nutates with great rapidity and is highly sensitive to contact (as maybe seen by rubbing one side and watching it for 5 min.); the hooks prevent the nutaton from dragging away a branch before it has had time to clasp its support (Darwin, Climbers, p. 106). The closed bud stands erect on an erect stalk, but when going to open, the tip of the stalk bends over. Fl. very protandr. with movement of sta. and styles. At first greenish with unpleasant smell (fly-fl.), it becomes purple with pleasant honey-like smell (bee-fl.). Afterwards the stalk goes through several contortions (cf. Linaria). Cobnut, Carylus, (W.I.) Omphalea triandra L. Cobresia Pers. (Kobresia Willd ). Cyper. (in). 5 N. palaeotemp. Coburgia Sweet = Stenomesson Herb. p.p. (Amaryll.). Coca, Ervthroxylum Coca Lam.; cocaine, ditto. Coccineus (Lat.), scarlet. Coccinia Wight et Arn. Cucurbitaceae (4). 20 trop. As. Afr. The fr. of C. itidica W. and A. is eaten as a veg. in India. Coccoceras Miq. Euphorbiaceae (A. II. 2). 3 Indomal. Coccocypselum P. Br. Rubiac. (i. 7). 10 trop. Am. Heterostyled. Coccoderma Miers. Menispermaceae. Nomen. Coccoglochidion K. Schum. Euphorb. (A. i. i). i New Guinea. Coccoloba L. (Coccolobis P. Br. ). Polygonaceae (in. i). 125 trop. and subtrop. Am. C. uvifera L., and others, ed. fr. (seaside grape). Cocconerion Baill. Euphorb. (inc. sed.). 2 New Caled. Coccothrinax Sargent. Palmaceae (i. 2). 10 warm Am. Coccule, portion of a divided coccus. Cocculus DC. Menispermaceae. 30 trop. and subtrop. Cocculus indicus, Anamirta Cocculus Wight et Arn. Coccus, n mericarp. Cochineal, Nopalea, Ofuntia. Cochlanthera Choisy (Clusia BH.}, Guttif. (v). i Venezuela. Cochlanthus Half. f. Asclepiadaceae (i). i Socotra. Cochlea, a closely coiled legume. Cochlear, spoon-shaped. cocas 153 Cochlearia Tourn. ex L. Cruciferae (2). 20 Eur., As. Minor. C. qffici- nalis L. (scurvy-grass) in Brit, with ± fleshy 1., chiefly at the seaside and on mts. (cf. Armeria). The thick root of C. Armoracia L. (horse- radish) is a condiment. Cochleate, coiled shell-shaped. Cochlianthus Benth. Leguminosae (in. 10). i Nepal. Cochlioda Lindl. Orchid, (n. 19). 5 trop. S. Am. Cult. Cochliostema Lem. Commelinaceae. i Ecuador, C. odoratissima Lem., cult. orn. perf. fl. The filaments of the fertile sta. develope both lat. and beyond the anthers into large wings. Anther-loculi spiral. Cochlospermaceae (EP.; Bixineae p.p. BH.}. Dicots. (Archichl. Parietales). 3 gen., 18 sp. trop. Trees and shrubs usu. with lobed 1. and racemose infl. of large $ , reg. or slightly •[• fl. K 4 — 5, C 4 — 5, A oo , G (3 — 5) with oo ov. in each on axile or parietal plac. Caps. Oily endosp. Coclilospermuin Kunth. Cochlospermaceae. 12 trop., mostly xero. ; some have stout tuberous underground stems ; many drop their 1. and flower in the dry season. Some cult. orn. Cochranea Miers. Boraginaceae (in). 10 Chili. Cockburnia Half. f. Globulariaceae. i Socotra. Cockle (Am.), Lychnis; -bur (Am.), Xanthimn. Cock's comb, Celosia cristata L. ; -head. (W.I.), Desmodium tortuosum DC.; -foot grass, Dactylis glomerata L. ; -spur (W.I.), Pisonia aculeata L. ; -spur thorn (Ceylon), Acacia eburnea Willd. Coco, Colocasia antiquorum Schott ; -de-mer, Lodoicea Seychellarum Labill.; -nut, Cocos nucifera L., Lodoicea; -plum, Chrysobalanus Icaco L. ; water nut, Nipa f net icons Thunb. ; -wood (W.I.), Inga vera Willd. Cocoa, Theobrotna Cacao L., and other spp. Cocops O. F. Cook. Palmaceae (iv. i). \ W. Indies. Cocos L. Palmae (IV. 2). 60 trop., esp. C. mtcifera L. (coconut), cult, throughout trop. It grows esp. well near to the sea, and its fibrous and woody fr. is capable of floating long distances uninjured, hence it forms a char, feature of marine island veg., and indeed probably became widely distr. in early times. It is a tall palm with large pinnate 1. and a dense monoec. infl. The .stem rarely stands vertically, but makes a gradual curve; this would appear to be due to helio- tropism. Fr. large, one-seeded. The outer layer of the pericarp is fibrous, the inner very hard (the shell of the coconuts sold in shops). At the base are 3 marks, corresponding to the 3 loc. of the ovary, two of which have become obliterated. Under one of these is the embryo. The thin testa is lined with white endosp., enclosing a large cavity, partly filled with a milky fluid. This palm furnishes many of the necessaries of life to the inhabitants of the tropics, and its products are largely exported from Ceylon, the Philippines, &c. The large 1. are woven into cadjans for thatching, mats, baskets, &c. ; their stalks and midribs make fences, brooms, yokes, and many other articles ol furniture. The bud or " cabbage " at the apex of the stem makes an excellent v< getable and is made into pickles and preserves. When flowering the infl. -axis is tapped for toddy, a drink like the Mexican pulque (cf. Agave), containing sugar. Evap. .of toddy i54 CO COS furnishes a sugar known as jaggery ; its fermentation gives an alcoholic drink, from which distillation produces the strong spirit known as arrack, while further fermentation gives vinegar. The fr. while young contain a pint or more of a sweetish watery fluid, a refreshing drink ; it decreases as the nut ripens. The kernels are eaten raw, or in curries, milk is expressed from them for flavouring, and oil is extracted by boiling or by pressure, in the latter case the kernels being first dried into what is known as copra. The refuse cake or poonac, left after the expression of the oil, is a valuable fattening food for cattle. The great use of the 'oil is for soap-making and margarine. In recent years a large industry has sprung up in desiccated coconut, largely used in confectionery, the kernel being sliced and dried in special desiccators. The outer wood of the stem (porcupine wood) is used for rafters, orn. articles. &c. The thick outer husk, rarely seen in Europe upon the nut, contains a large number of long stout fibres running lengthwise. The nut is placed in water till the soft tissues between these fibres decay, and the fibre (coir) is then beaten out ; or sometimes the fibre is obtained by special machinery. Cocos or cocus wood, Brya Ebcmts DC. Codia Forst. Cunoniaceae. 9 New Caledonia. Codiaeum Rumph. ex A. Juss. Euphorb. (A. n. 5). 6 Indomal., Polynes., Austr. C. variegatum Blume cult., esp. in trop., for its coloured 1. ; usu. known as Crotons, and also used as hedges. Some have curious 1 , often twisted, or with two blades separated by a length of petiole. Codlins and cream, Epilobium hirsntum L. Codon L. Hydrophyllaceae. 3 S. Afr. Codonacanthus Nees. Acanthaceae (iv. B). 2 Khasias, China. Codonanthe Hanst. Gesneriaceae (i) 10 trop. Am. Codonocarpus A Cunn. ex Hook. \Gyrostemon Desf. £f.). Phyto- laccaceae. 3 Austr. Codonocephalum Fenzl. Compositae (4). 2 W. As. Codonopsis Wall. Campanulaceae (i). 15 As. Cult. orn. fl. Codonorchis I.indl (Pogonia BH.}. On-hid, (n 2). 2 temp. Am. Codonosiplion Schlechter. Orchid, (n. 16). 2 New Guinea. Codonostigma Klotzsch (Scyphogyne JBH.). Eric (iv. 2). i S. Afr. Codonura K. Schum. Apocynaceae (n. i). i Cameroons. Coelachne R. Br Gramineae (9). 5 Indomal , China, Austr., Madag. Cortacnyrum Nees (Eragrostis p. p. BH.}. Gram. (n). 2 S.W. As. Coelanthum E. Mey. Aizoaceae (l). 2 S. Afr. Coelartnron Hook. f. (Amlropogon p.p. EP. ). Gramin. (2). i Indomal. Coelebogyne J. Sm . = Alchornea Sw. p.p. (Euphorb.). Coelia Lindl. Orchidaceae (11. 6). 5 trop. Am., W.I. Coelidium Vog. Leguminosae (ill. 3). 8 S. Afr. Coelina Nor. Inc. seel. Nomen. Coeliopsis Reichb. f. Orchid. (H. 13). i Panama. Coelocarpum Balf. f. Verbenaceae (i). 2 Socotra, Madag. Coelocaryon Warb. Myristicaceae. 5 trop. Afr. Coelococcua H. Wendl. (Metro.\ylou p.p.)- Palm. (in). 2 Polynes. Coelodepas Hassk. Euphorbiaceae (A. n. 2). ? Indomal. CO IX 155 Coelodiscus Bail). Euphorbiaceae (A. n. 2). 5 Indomal. Coeloglossum Harttn. (Habenaria p.p. BH.). Orch. (II. i). 2 N. temp. Coelogyne Lindl. (BH. incl. Pleione D. Don). Orchidaceae (it. 3). 1 20 Indomal. Coelonema Maxim. Cruciferae (4). i China. Coeloneurum Radlk. Solanaceae (4). 2 San Domingo. Coelopleurum Ledeb. (Archan^elica BH.}. Umbel, (in. 5). 6 N. Am., E. As. Coelopyrum Jack. Inc. sed. i Malaya. Coelorachis Brongn. (Rottbodlia L.). Gram. (2). 7 N. Am. Coelospermous, with boat-shaped seeds. Coelospermum Blume. Rubiac. (n. 9). 12 Malaya, Austr., Polyn. Coelostegia Benth. Bombacaceae. 3 Malaya. Coelostelma Fourn. Asclepiadaceae (II. 4). i Brazil. Coemansia March (Pentapanax p.p.). Araliac. (2). i Brazil. Coffea L. Rubiaceae (n. 4). 45 palaeotrop., esp. Afr. C. arabica L. (Arabian coffee) largely cult, in S. Brazil, Java, Jamaica, and else- where, often under the shade of large trees. C. liberica Hiern (Liberian coffee) cult. usu. at lower elevations; its produce is not so good. Other sp. nre also used. The fr. is a 2-seeded drupe, resembling a cherry. The pulp and the endocarp (which covers the two seeds like a layer of parchment) are mechanically removed. The seed, or coffee-bean, has a deep groove on the ventral side ; by soaking it in water the endosperm is softened and the embryo may be dissected out. The stimulating property depends on the presence of the alkaloid caffeine. Coffee cultivation was from 1850 to 1880 the mainstay of Ceylon agriculture, but was killed out largely by the attacks of a fungus (Hemileia vaslatrix) and the green bug. By far the largest cult, is that of Brazil, which in 1912 exported ^45 million worth of coffee. (Raoul, Culture du Cafeier, Paris.) Coffee, Coffea arabica L., &c. ; Kentucky-, Gymnodadus. Coffin nail, Anacardium occidentale L. Cogniauxia Baill. Cucurbitaceae (3). 4 trop. Afr. Cogswellia Spreng. (Peucedanum p.p.). Umbelliferae (in. 6). 70 N. Am. Cogwood, Zizyphus chloroicylon Oliv. ; (W.I.), Ceanothus. Cohesion, union of mi mbers of same whorl, eg. petals. Cohnia Kunth (Confyline p.p. BH.}. Lili. (vi). 3 Masc., New Caled. do. Reichb. f. =Cohniella Pfitz. Cohniella Pfitz. (Cohnia Reichb. f.). Orchid, (n. 19). r C. Am. Cohort, a group of allied fams., now termed an order. Cohosh (Am.), Cimicifnga; blue-, Caidophyllinn. Cohune nut, Atta'ea cohiuif Mart. Coilochilus Schlecht. Orchid, (u. 2). i New Caled. Coincya Rouy (Rapliainis p.p. BH.}. Crucif. (2). i Spain. Coinochlamys T. Anders. Loganiaceae. 5 W. Afr. Coir, Cocos nticifera L. Coix L. Gramineae (i). 6 India, China, esp. C. Lachryma L. (Job's tears) with inverted pear-shaped body at base of infl., the sheath of the br. of the infl., hollowed out and containing the i-fld. ? spikelet ; COIX the (f project beyond the mouth; cult, for food in Khasia Hills and Burma ; used in medicine in China. Cola Schott et Encll. Sterculiaceae. 50 Afr. C. vera K. Schumann and C. acuminata Schott et Endl. ( possibly identical) are the source of the kola nuts which form a principal article of trade in W. Africa. The nuts contain much caffein, and when chewed confer considerable power of sustaining fatigue ; they are consequently a staple in the diet of the negroes (cf. Erythroxylon). The tree is as yet rarely cult., but is very common in W. Afr. The nuts are skinned after keeping for a few days, and packed between 1. to keep them damp. Colax Lindl. (Lycasie Lindl. BH.}. Orchidaceae (n. 14). 3 Brazil. Cult. Colchicaceae (Warming) — Liliaceae (suborder i). ColcMcum L. Liliaceae (i). 45 Eur., W. As., N. Afr. C.autiimnale'L. (autumn crocus or meadow saffron), Brit. Below the soil is a large corm (fig. and description of corm &c. below). In autumn the fl. projects out of the soil. The P-tube is long, and the ovary remains below ground, protected from cold, &c. The protog. fl. is visited by A, pi. in fl. in autumn, xj. B, 1. and opening fr. in following summer, Xj. C, underground portion of fig. pi. cut lengthwise ; the thick outer line repres. the brown membrane enveloping the whole ; to the right is the corm formed from the base of last year's shoot, a withered portion remaining at the apex ; to the left is the fig. axis, a lat. shoot from the base of the corm ; from the base of the axis spring r. and above are the 1., sheathing and foliage; the fl. arises in the axil of one of the uppermost foliage 1., which will appear above ground with the fr. next spring, when the lower portion of the axis will swell to form a new corm. Reduced. D, section of upper part of fl. X J. E, ovary cut lengthwise. F, cross- section ovary. G, a single stigma. H, cross-section fr. I, ditto seed. BEGHI after Berg and Schmidt. E to G, and I enlarged, H x£. COLLECTING bees. In spring the 1. appear and the capsule is brought above ground by the lengthening of its stalk. The seeds and corms are used in medicine, in gout. Coldenia L. Boiaginaceae (n). 15 trop., subtrop. Colea Boj. Bignoniaceae (4). 18 Madag., Masc. Coleantnera Stschegl. Epacridaceae (3). 3 W. Austr. Coleanthus Seidl. Gramineae (8). i N. temp. Colebrookea Sm. Labiatae (vi). i India. Coleocoma-F. Muell. Compositae (4). i trop. Austr. Coleogyne Torr. Rosaceae (in. 3). i California. Coleonema Bartl. et Wendl. Rutaceae (i). 5 S. Afr. Coleosanthus Cass. = Brickellia Ell. (Compos.). Coleospadix Becc. (Ptychosperma EP.). Palm. (iv. i). 2 New Guin. Coleostachys A. Juss. Malpighiaceae (n). i N. S. Am. Coleotrype C. B. Clarke. Commelin. 3 S.E. Afr., Madag. Coleus Lour. Labiatae (vn). 150 palaeotrop. Many forms and hybrids with varieg. and coloured leaves, cult. C. elongatus Triinen is a peculiar sp. found only on the top of one mountain in Ceylon, and must have arisen by mutation {Ann. Perad. IV. r). Colic-root (Am.), Aletris. Colignonia Endl. Nyctaginaceae. 7 Andes. Collabium Blume. Orchidaceae (n. a. n). 3 Malaya. Colladonia DC. = Prangos Lindl. p.p. (Umbellif.). Collaea DC. = Galactia P. Br. p.p. (Legum.). Collar, junction of root and shoot. Collards (Am.), a form of cabbage. Collateral branches, buds, cf. Buds. Collecting (notes for field botanists, travellers and collectors). OUTFIT. Any or all of the following may be needed, according to the places to be visited, and the kind of work to be done. All that is needed should be taken from the start, as it is usually difficult to get suitable things quickly elsewhere; extra supplies of paper, &c. may be sent to the " Poste restante " at places to be visited en route. Portfolios for pressing plants as collected, lightly made of two strong cloth-covered pasteboards (17 in. xu in.) with encircling straps and handle, and to contain 30—50 sheets of paper. Specimens as collected are put at once into these, and time is thus saved in making large collections, e.g. on a journey in new country. Collecting tins or vasculums of various sizes for bringing plants home for further study. When slung on the back, the hinges should be on the lower side of the lid, and the bolt should slide downwards to fasten, otherwise it is liable to work loose. Small specimens are best carried in small round- cornered tobacco or tooth-powder tins, not among large ones in a general vasculum. Presses for drying pi.; each of two outer frames of \ inch iron rod 17^ x 1 1^ inches, filled in with stout wire netting soldered to the iron. The papers lie between these frames and the whole is strapped with two stout straps to obtain the pressure". 3 I i58 COLLECTING Lattices, 1 7 x 1 1 indie*, for admitting air between the masses of plants in the press, made of two sets of parallel thin laths fastened together. Drving paper in sheets 17* 1 1 inches in ample quantity ; slout Manila is best, blotting paper is too fragile. Mounting paper in sheets i6i v. io.j inches (standard si^e of Kew herbarium) or sheets of news- paper or other common paper for preservation of dry specimens removed from the press. Unfolded envelopes of thin paper cut into the shape shown, for seeds, flowers, &c.; place the specimen on i and fold over the wings 2, 3, 4, 5 in order. 2 must be the same size as i. Waxcloth for tying up bundles of dried and mounted specimens ; waterproof canvas for covers for presses, &c. in case of rain. Corrosive sublimate (mercuric chloride) and alcohol for poisoning specimens ; made up as required in the proportion of i part to 50. Large dish for poisoning. Naphthalin for keeping away insects. Kerosine tins or other square tins with large lids for preserving specimens in alcohol (lids that push in airtight, as in many tobacco tins, are the best) ; soldering apparatus for fastening up when full. Bottles with stoppers for preserving delicate specimens ; neckless glass tubes, with corks, of various sizes. Bottles are easily packed in joints of bamboo, tubes in small tins. Aluslin for wrapping alcohol specimens. Each should be wrapped with its label (in Indian ink, or better on metal) in a piece of muslin and packed in the tin ; specimens cannot then become mixed together, and can be closely packed . Alcohol for preserving ; ordinary methylated spirit is best for most things, but some require 70 % alcohol, and some absolute alcohol. Formalin, picric acid, chromic acid, glycerin, or other preservatives. Butterfly-net, killing-bottle, insect boxes, entomological pins, if eco- logical work is to be done. Hunting-knife, cutlass, or kukri for lopping creepers, &c. ; pruning shears for cutting branches ; strong pocket knives ; strong narrow-bladed trowels; geological hammer; strong rope for climbing, &c. Travelling microscope and lenses ; dissecting microscope; pocket lenses; microscope slides, cover-slips in alcohol or oil; scalpels; dissecting needles ; scissors large and small and with fine points; razors for section- cutting ; forceps; dishes; watch-glasses; camel-hairbrushes. Reagents and mount ants for simple microscopic work, e.g. iodine, glycerine, haematoxylin, gold-size, Canada balsam in xylol, alcohol, oil of cloves. Compass (prismatic by preference) ; spirit-level ; aneroid barometer ; thermometers (ordinary, maximum and minimum, wet and dry bulb) ; field glass (very useful for studying cliffs, ravines, trees, &c.) ; maps (geographical, geological, outlines for marking distribution, &c.). Photographic camera and lenses; tripod; films or plates in soldered tins ; chemicals and dishes for developing, fixing, &c. Drawing pencils (hard, medium, soft) ; drawing cards ; sketch-block; colours; brushes; india-rubber; ink ; compasses ; ruler; scales in inches and centimetres; gum and brush; pins ; pens ; pencils; stylograph. Spirit-lamp; tape-measure; string, twine and thread; thin wire; sheet'/tv?;/1 or zinc for labels (if latter, also solution of platinic chloride for writing on it) ; ghte-pot and glue for mounting. COLLECTING 159 Note-books with numbered detachable pages, so that the descr. ui" each specimen can be separated ; consecutively numbered and perforated labels for specimens, the numbers to correspond to those in note- book. The labels may be joined to the pages or in sheets like postage stamps. More than one of each number will usu. be needed. COLLECTING AND PRESERVING. The following hints will be found useful. Decide the general object of the work in advance, and collect principally for the furtherance of that object. In collecting for subsequent distr., collect enough specimens to go round, but do not seriously diminish a plant in its native locality. Do not collect immediately on arrival ; first become familiar with the plants aii'1 their local features and distribution. Better results are obtained by choosing certain localities as headquarters and working these thoroughly, than by rushing through a large district. Do not collect herbarium material in wet weather. Collect specimens which are as typical as possible, but also take some illustrating the range of variation, the difference of habit and size on different soils or situations, &c. Collect entire pi. if possible, incl. r. In shrubs or trees, twigs with 1. in all stages, portions of stem-bark and anything else necessary for a com- plete descr. Do not forget rad. 1., buds, fl., ripe and unripe fr., seeds. If a large collection is being made, it is quicker to use the portfolio than the vasculum, and to press each specimen as soon as obtained. Large flowers or heads (e.g. thistles), fruits, roots, tubers, &c. may be sliced in half before pressing, or the surface only sliced off. Notes and sketches should be made of the original appearance. L. of Conifers, Heaths, Succulents, &c. fall when dried, unless previously immersed a few seconds in boiling water. Do not immerse fl. Thurny and prickly plants should first be placed between boards and pressed down with the feet ; the prickles would otherwise tear the papers. Delicate water plants should be arranged upon sheets of white paper under water, and always remain on these sheets while drying. It saves time in drying delicate specimens to keep each always in a folded sheet of very thin paper. Place extra fl., small fragments, seeds, &c., in small envelopes, numbered to correspond with the specimens; do not have any small parts loose, or confusion may result. Place all specimens in the press the day they are collected. Withered plants may be soaked in water; if the stem be cut 2 — 3 in. above the former cut and tinder water it will often revive quickly. Label every specimen with its consecutive number in such a way that the number cannot be lost. Punched labels are best, tied on with thread. See that all envelopes, &c. have the same number. Spread out the specimens naturally. It many 1. &c. overlap, place bits of drying paper between. If stems have to be cut, mark the corre- sponding ends by stars on the paper. Spread out some fl., leave others unspread, and divide some in the antero-post. plane. Divide some fr. lengthwise and crosswise. 1 60 COLLECTING Arrange the specimens on the sheets so that they form a steady pile without lumps in the middle. Place a lattice upon every five inches of specimens. When all are ready place in the press and draw the straps as tight as possible, or better, place about 10 Ibs. weight upon it. Tighten the straps as the plants shrink. Change drying papers at least once daily: dry used paper in the sun or by the fire ; use warm driers where possiblr . See that petals, &c. do not stick to the paper: if necessary put slips of tissue paper under. In changing the papers, put the outer specimens inside, so that all shall dry evenly. Drying should be as rapid as possible to prevent loss of colour, blackening, &c. Fully dried plants no longer feel cold on the cheek, and are stiff and brittle. Dried specimens should be poisoned by a brief immersion in i % solution of mercuric chloride in alcohol. They should then be dried in the air, mounted (with glue) or laid between sheets of paper, and tied up in wax-cloth with a little naphthalin to keep out insects. Material for subsequent microscopic examination or for museums must usu. be preserved in alcohol. Cut into small portions, attach label (best of zinc written on with solution of platinic chloride, but paper and pencil or Indian ink will do temporarily) ; place in methylated spirit for a few days, wrapped in muslin, and finally preserve in large tin. A few inches of spirit at the bottom, enough to keep all specimen* moist when the tin is soldered, will suffice for most material. Specimens for embryo- logical, delicate histological, cytological, and other investigations, and delicate plants or organs, must be separately preserved from the fir t in absolute alcohol in bottles or tubes. Labels should be put inside these. Contents of bottles may be written on the ground surface of the stoppers, so as to be legible through the neck. Museum material may also be preserved in formalin (i part of ordinary solution to 10 or more of water). Some special preservatives, e.g. picric and chromic acids, are used in special cases. Specimens illustrative of economic uses of plants and their products should be collected in less known districts, e.g. samples of gums, resins, caoutchoucs, oils, fibres, timbers (portions of trunks, or slabs 8x4x4 inches), food-products, drugs, dyes, tans, &c. In all cases the exact origin should be verified, and herbarium specimens taken, bearing numbers to correspond with those placed on the products. From less known countries, endeavour to bring back living seeds (ripe, well dried, dry in canvas bags, or packed in charcoal in tins if to be long kept), bulbs and tubers (gathered when dry and with withered leaves), succulent plants (gathered dry and loosely packed), living plants (planted in earth in Wardian cases or sometimes in bamboo pots, if possible some weeks before moving). Cuttings may sometimes be brought in oiled silk wrappers; pseudobulbs in boxes with air-holes; tree lerns with the fronds removed, and a ball of earth round the root. RECORDING. The following hints are worth noting. Make all notes immediately upon observation of the facts ; never trust to memory, nor delay recording. Make all notes about individual specimens upon detachable sheets, CO LOG ASIA 161 numbered to correspond with the specimens ; never describe two or more on the same sheet ; never use the same number twice. Make no record till satisfied of its truth and accuracy. Accompany all notes with maps, drawings, sketches, or photographs as far as possible. Sketch and photograph all peculiarities of habit, characteristic forms of vegetation, and other features of interest. Mark all plants in such pictures with numbers corresponding to their numbers in your collection, and write a full description of each picture before leaving the spot. If a photograph is taken, make a rough sketch (from the picture on the focussing screen or finder) of the scene and put numbers to the plants. Label all specimens as collected with consecutive numbers, and subsequently with permanent labels (about 3x2 inches) giving name of herbarium, collection, tour or district in which they were collected, date, locality, and collector, as well as the number. As each specimen is gathered, record date, exact locality, elevation above sea, habit, colour of fl. and fr., scent, presence or absence of honey, floral mechanism, insect visits, and any other features and facts that cannot be ascertained from the specimens preserved. Note the comparative frequency of each species, the kind of situation and soil it affects, and the species with which it is found in association. Endeavour to note the chief general forms of vegetation and the local grouping of pi. in districts studied. Pay special attention to ecological and geographical questions. Record native names (question several different persons before deciding), economic uses, and points of general or ethnological interest. Further details of the subjects treated in this section may be found in Hints for Collectors (Kew Bulletin, 1914, p. 97), the Admiralty Manual of Scientific Enquiry, Dammer's Handbuch ftir Pflanzen- sam>nkr, Stuttgart, 1891, Asa Gray's Structural Botany, &c. Collective (fr.), resulting from several fl., Ficus, Morns, Platanns. Collet, collar. Colleters, glandular hairs. Colletia Comm. ex Juss. Rhamnaceae. 15 S. Am. Habit peculiar; in each axil are 2 serial buds ; the upper gives a triangular thorn, the lower fls. or a branch of unlimited growth. Colliguaja Molina. Euphorbiaceae (A. n. 7). 6 temp. S. Am. Collinsia Nutt. Scrophul. (n. 4). 25 N. Am., often cult. orn. fl. The fl. resembles, in shape and mech., that of Leguminosae. Collinsonia L. Labiatae (vi). 2 All. N. Am. Collinus (Lat.), on low hills. Collococcus P. Br. Inc. sed. Nomen. Collomia Nutt. Polemoniaceae. 20 W. Am. The seed coat has a covering of cells with mucilaginous walls which swell when wetted (cf. Brassica, Linum, &c.). Cult. orn. fl. Coliyris Vahl = Dischidia R. Br. (Asclep.). Colmeiroa F. Muell. Saxifragaceae (v). t Lord Howe I. Colobanthus Bartl. Caryophyllaceae (i. i). 15 S. Am., Austr., N.Z. Petals o. Sta. in one whorl. Colocasia Schott. Araceae (vi). 8 Indomal. Tuberous herbs or small shrubs. Monoec. Sta. in synandria. C. antiqnorum Schott (taro, W. II 1 62 COLO C ASIA coco, or scratch-coco), cult, in trop. for its rhiz., which when boiled loses its poisonous nature and forms valuable food. Colocynth, Citrullus Colocynthis Schrad. Colocynthis (Tourn.) L.= Citrullus Neck. (Cucurb.). Cologania Kunth (Amphicarpaea EP.). Legum. (m. 10). 30 Am., As. Colonist, weed of cult, land, rare elsewhere. Colophony, a form of resin. Coloptera Coulter et Rose. Umbell. (in. 6). 3 N.W. U.S. Colosantliera Pohl. Inc. sed. Nomen. Colour, change in fi, cf. Change; of young 1., Amktrstia, Brownea, Cinnaniomum, Dryobalanops, Haeinatoxylon. Colpias E. Mey. Scrophulariaceae (il. 3). i S. Afr. Colpodium Trin. Gramineae (10). 12 N. temp. Colpoon Berg. Santalaceae. 3 S. Afr. Colpothrinax Griseb. et H. Wendl. Palm. (i. 2). i Cuba. Colquhounia Wall. LaUatae (vi). 5 Indomal. Colt's foot, Tussilago Farfara L. ; (W.I.) Piper umbellatum L. Colubrina Rich, ex Brongn. Rhamnaceae. 20 trop., subtrop. Columbaria J. et C. Presl = Scabiosa Tourn. (Dipsac.). Columbia Pers. Tiliaceae. 15 trop. As. Columbine, Aqiiilegia vulgaris L. Columella, central axis of fr., Geranium, Thuja. Columellia Ruiz et Pav. Columell. 3 N. Andes. Shrubs with evergr. opp. exstip. 1. Fls. in cymes, 5, nearly reg. K 5, C (5), A 2, short and thick with irreg. broad connective and i twisted pollen sac. No disc. G (2), imperfectly 2-loc. ; ov. GO, anatr. ; style short and thick with broad 2 — 4-lobed stigma. Caps., enclosed in K. Endosp. Columelliaceae. Dicots. (Sympet. Tubiflorae ; Personates BH.}. Only genus Coluniellia (q.v.). [Van Tieghem, Ann. Sc. Nat. 8. xviii. 155.] Column, cf. Orchidaceae. Columnea Plum, ex L. Gesneraceae (i). 75 trop. Am., several climbers and epiphytes. Anisophylly is frequent. Columniferae (Warming). The J2th cohort of Choripetalae. Coluria R. Br. Rosaceae (ill. 2). 4 Siberia, China. Colutea (Tourn.) L. Leguminosae (in. 6). 12 S. Eur. to Himal. C. arborescens L. (bladder-senna) cult. Its 1. have similar properties to senna (Cassia) and are used to adulterate the latter. The pods are inflated and burst on being squeezed. Coluteocarpus Boiss. Cruciferae (2). i W. As. Mts. Colvillea Boj. ex Hook. Leguminosae (n. 7). i Madag. Colza, Brassica Napus L. Coma, a tuft of hairs. Comandra Nutt. Santalaceae. 4 Eur., N. Am. Comanthosphace Sp. Moore. Labiatae (vi). 4 Japan. Comarella Rydberg (Potentilla p.p.). Rosac. (m. 2). 2 N. Am. Coniarobatia Greene (Rubus p.p.). Rosac. (in. 2). i N.W. Am. Comarostaphylis Zucc. = Arctostaphylos Adans. p.p. (Eric.). Comarum L. = Potentilla L. (Rosac.). Combretaceae (EP., BH.). Dicots. (Archichl. Myrtiflorae; Myrtales BH-). i6gen., 450 sp. trop. and subtrop. Trees and shrubs with COMOCLADIA 163 alt. or opp. simple 1. and no stips. ; many climbers, some twining, some with hooks formed of the persistent bases of the petioles. Fls. usually sessile in racemose infls., $ , reg. Typical formula: K 5, C 5, A5 + 5» G i-loc. ; ov. 2 — 5, anatr., pend. ; style simple. There is a disc on the summit of the ovary, sometimes with various outgrowths. Fr. dry, i-seedecl, often winged at the angles. Seed exalb. ; coty- ledons usu. twisted spirally. Chief genera : Terminalia, Combretum, Quisqualis. Cornbretocarpus Hook. f. Rhizophoraceae. i Borneo. Combretodendron A. Chevalier. Combretaceae. i trop. Afr. Combretopsis K. Schum. (f.ophofyxis Hk. f.). Icac. i New Guin. Combretum L. Combretaceae. 330 trop. and subtrop., exc. Austr. and Polynes. The fruit of C. bittyrosum Tal. (trop. Afr.) yields a butter-like substance known as Chiquito, used as butter. Comesperma Labill. (Bredenieyera p.p. EP,}. Polygal. 25 Austr. Cometes L. Caryophyllaceae (i. 6). 2 W. As. Cometia Thou. Euphorbiaceae (A. i. i). 2 Madag. Comfrey, Syniphytiini offichtale L. Cominsia Hemsl. Marantaceae. 2 Austr., Solomon Is. Commelina L. Commelinaceae. 110 trop. C. coelestis Willd. cult, orn. fl. In the fl. there is division of labour between the sta. (cf. Heeria). The fl. stands horizontally and the sta. and style project beyond the C. The upper 3 sta. (in many sp.) are almost sterile, but the lobes are juicy. The two lat. lower sta. and the median one are fully fertile. Bees often climb up and pierce the upper anthers for honey. C. benghalensis L. has subterranean cleisto- gamic fls. The rhizome of some sp. is edible. Commelinaceae (EP., BH.). Monocots. (Farinosae ; Coronarieae BH.}. 25 gen., 300 sp., mostly trop. and subtrop. herbs with jointed stems and alt. sheathing 1. Infl. usu. a cincinnus of the Boraginaceae type. Fl. $, usu. reg., commonly blue. Typical formula K 3, C 3, A 3 + 3, G (3), but some sta. commonly absent or stds. K and C differ in colour and texture. Ov. 3-loc., with a few orthotr. ov. in each. Caps, loculic. or indeh. Endosp. fleshy; seed often arillate. Chief genera: Commelina, Tradescantia. Commensalism, living together for mutual benefit. Commersonia Forst. Sterculiaceae. 10 trop. As., Austr. Commersorchis Thou. Orchidaceae (inc. sed.). i Mascarenes. Commicarpus Standley (Boerhaavia p.p.). Nyctag. 2 N. Am., W.I. Commidendron Burch. Compositae (3). 5 St Helena. C. gumtni- fornni DC. yields a gum. Commiphora Jacq. (Bahamodendron Kunth). Burseraceae. 80 trop. As., Afr. Several spp. yield myrrh. The resin exudes from the tree and collects in lumps. It is used in medicine and in incense, &c. C. opobalsamuin Engl. is said to yield the resin balm of Gilead. Other sp. yield bdellium and other resins. Commissure (Umbelliferae), face by which carpels cohere. Common receptacle, the receptacle of all fl. on a head. Communis (Lat), social, general. Community (Cl.), a mixture of individuals of 2 or more spp. Comocladia P. Br. Anacardiaceae (3). 15 W.I., C. Am. II — 2 164 CO MO LI A Comolia DC. Melastomaceae (i). 20 S. Am. Comomyrsine Hook. f. (Weigeltia p.p. EP.). Myrsin. (n). 48. Am. Comopycna O. Ktze. = Pycnocoma Benth. (Euphorb.). Comoroa Oliv. (Teclea p.p. EP.). Rutaceae (iv). i Comoro Is. Comose, hairy in tufts. Comostemum Nees = Androtrichum Brongn. (Cyper. ). Comparettia Poepp. et End!. Orchidaceae (n. 19). 5 trop. Am. Cult. Compass-plants, Silphiiini laciniatuin L., Lactuca Scariola L. Comperia C. Koch (Orchis Bff.). Orchid. (11. i). i S.E. Eur. Compital, where veins intersect at an angle. Complanate, flattened. Complete (fl.), with two whorls of perianth. Complicate, folded on itself. Compositae (EP., BH.}. Dicots. (Sympet. Campanulatae ; Asterales BH.). The largest fam. of flg. pi., comprising about 900 genera, with over 13,000 sp. — more than io°/0 of the total. They are distr. over the greater part of the earth. Although so large a fam. they are well marked in their characters and cannot be confounded with any other, though they have a superficial likeness to Dipsaceae and Calyceraceae. Living in almost every conceivable situation, they present great variety in veg. habit, often within a single genus, e.g. Senecio (q.v.). Water and marsh plants and climbers are rare, and so also are epiphytes. This latter is interesting, for the distr. mech. of these pi. is admirably suited to an epiph. existence, and xero. is not uncommon. The enormous majority are herbaceous pi.; trees and shrubs are comparatively rare (about i|°/0). It is worthy of note that the latter often form an important feature in the Composite flora of oceanic islands (see Wallace's Island Life). L. usu. alt., frequently rad., opp. in Heliantbeae, whorled in a few cases, e.g. Zinnia vertidllata ; slips, rarely present. R. usu. a tap-root, sometimes tuberous as in Dahlia, &c., often thickened like that of a carrot, e.g. Taraxacum, Cichorium, &c. For details of veg. organs refer to individual gen.; e.g. Aster, Barnadesia, Bellis, Bidens, Cichorium, Dahlia, Espeletia, Gnaphalium, Helianthus, Helichrysum, Lactuca, Mutisia, Petasites, Senecio, Silphium, Taraxa- cum, &c. All tribes exc. 12 and 13 contain oil-passages in the root, stem, &c. In 13 (Cichorieae), laticiferous vessels are present, commonly containing a milky white latex (e.g. lettuce, dandelion). Infl. of racemose type, the fls. arranged in heads (capitiila), or rarely in spikes. These heads are again arranged in many cases into larger infls. — racemes, corymbs, &c., or even into cpd. heads (Echinops, &c.). In this last case, however, the smaller heads contain only one fl. each. Head surrounded by an invol. of bracts, usu. green, which performs for all the fls. of the head the functions that in most plants are performed by the calices of the individual fls., viz. protection of the bud and of the young fr. Fls. arranged upon a common receptacle — the enlarged end of the axis — of various shapes, most frequently flat, slightly convex or even spindle-shaped. The shape and surface-condition of the receptacle are chars, of COMPOSITAE 165 importance in classification of the fam. It may be smooth or hairy, &c. ; there may (Helianthus, &c.) or may not (Calendula, &c.) be, upon it, scaly br. belonging to the individual firs. In Cynareae these br. are divided so as to form numerous bristles. In the simplest case the fls. of a single head are all alike and §, but there are many deviations from this type. The fls. may be all actinomorphic (tubular') or all •]• (ligtilate) ', see below. Very commonly however, as in daisy or sunflower, there is a distinction into a disc of actinomorphic fls., and a marginal ray of -|- fls. Or, as in Centaurea sp., the outer florets may be actinomorphic but different in size from the central. The number of ray-florets varies in different sp., but according to definite rules. The distribution <>/" sexes among the fls. of a head varies much. The most common case is gynomonoecism, the ray-florets ?, the disc ?. The very large ray-florets of Centaurea sp. and others are completely sterile (cf. Hydrangea, Viburnum, &c.). Cf. also Tussi- lago, Petasites, &c. Tlas Jhnoer is fully epig., usu. j-merous. K absent in Ambrosia and its allies, Siegesbeckia, &c. ; in some cases it appears only as a slightly -globed rim upon the top of the inf. ovary (cf. Rubiaceae and Umbelliferae) ; usu. it takes the form of hairs or bristles— the pappus — and enlarges after fert. into a parachute (Dandelion) or into hooked bristles (Bidens) to aid in clistr. (see below). C (5), valvate in bud; actinom. (tubular) or •[• . Of the latter form there are two varieties, labiate (lipped) and ligulate (strap-shaped). The latter term, strictly speaking, should be applied to those corollas which are strap- shaped in form with 5 teeth at the end repres. the petals, but is usu. also given to those lipped forms where the lower lip is strap-shaped and ends in 3 teeth. Sta. 5, epipet. with short filaments, alt. with the petals. Anthers intr., cohering by their edges (syngenesious), forming a tube around the style (cf. Lobelia). G (2), with a simple style that forks at the end into two stigmas, an ant. 'and a post, (see diagram). The construction of the style and stigma is of importance in the classification. There is often a brush of hairs on the style below the stigmas. Only the inner (upper) surfaces of the stigmas are as a rule receptive to pollen. Ovary i-loc. with i erect, basal, anatr. ov., which gives an exalb. seed with straight embryo, enclosed in the dry indeh. pericarp. This fr. is usu. termed an achene, but of course is, if one adhere strictly to definitions, a pseudo-nut, as its pericarp is partly axial, and there is > one cpl. It is often crowned with a pappus (see below). Natural History of the Flower. Being massed together in heads, the individual fls. may be, and usu. are, comparatively very small, and the advantage is gained that a single insect visitor may fert. many fls. in a short time without having to fly from one to the other, while there is no loss of conspicuousness, and a considerable saving Floral Diagram of Composite fl. with pap- pus (after Eichler). The small outer lines repre- sent the pappus-bristles. 1 66 COMPOS1TAE of corolla-material, dvc. Throughout the fam., the same type of mech. of the individual fi. is found, the differences being slight and unimportant. It is simple, but effective. Honey is secreted by a ring-shaped nectary round the base of the style, and protected from rain and from short-lipped insects by the tube of the C. The depth of the tube varies within fairly wide limits, but is never so small as to permit the shortest-lipped insects to obtain the honey. As a fam., the C. all belong to Miiller's fl. class B', but there is considerable variety in the depth of tube, &c., and therefore also in the composition of the group of visiting insects to each. Thus the long-tubed purple- flowered Centaureas, &c. are mainly visited by bees and Lepidoptera, while the short-tubed yellow Leontodons or white Achilleas are visited mainly by flies. At the time when the fl. opens, the style, with its stigmas tightly closed against one another, is comparatively short, reaching up to, or projecting a small distance into, the anther tube. The pollen is shed into this, and as the style grows it presses the pollen little by little out at the upper end of the tube where it will come into contact with visiting insects. At last the style itself emerges and the stigmas separate. The fl. is now ? . Finally, in a great many cases, the stigmas curl so far back that they touch the pollen upon their own style, so that every fl. is certain to set seed, even though it be by self- fert. In a few cases, e.g. Senecio vitlgaris, insect visitors are very rare, and the fl. depends entirely on self-fert. The mech. is about the simplest and most perfect that exists for attaining the desired ends. A striking contrast is seen in the orchids ; they have bizarre fls. with most elaborate mechs., and an enormous number of seeds in every caps. An interesting modification of the mech. is found in Cynareae (see Centaurea) where the sta. are irritable. See also Artemisia (wind-fert.). The invol. bracts, or ray-florets, or both, often close up over the central fls. in cold or wet weather, thus protecting the fls. Natural History of the Fruit. The ripening fr.-head is generally protected from injury by the invol. bracts, whicli bend inwards over it, performing the function of a K. The calices of the individual fls. are thus rendered useless in this respect and are, in most C., used for purposes of distr. of the fr. In most cases, the K, after the fert. of the fl., grows into the familiar pappus, as seen in dandelions or thistles, usu. composed of fine hairs, often branched, but in some cases, e.g. Achyrachaena, leafy and membranous. The hairs are hygroscopic and spread out in dry air; this often helps to lever the fr. oft" the receptacle. In Adenostemma the pappus is sticky. In Bidens and others the pappus is formed of stout barbed bristles ; the fr. adheres to animals. In Arctium the invol. br. become hooked at the tips and cling to animals. In Xanthium the recept. is provided with hooks. In Siegesbeckia the bracts are sticky. A few genera, e.g. Helianthus, Bellis, &c., have no special arrangements at all, and the frs. remain upon the common receptacle till jerked off by wind or otherwise. General Considerations. The C. are generally regarded as occupying the highest position in the Veg. Kingdom. Their success COMPOSITAE 167 may he put down perhaps to the concurrence of several useful peculiarities, viz. 1 i) the massing of the fls. in heads, surrounded by invol. bracts: from this there results (a) greater conspicuousness, especially when ray-florets are developed; (A) a saving of material in the corollas, &c. ; (c) the fact that one insect visitor may fertilise many fls. in a short time without having to fly from one to another ; (2) the very simple and effective floral mechanism, which ensures ( 9 as a rule) on its lower surface (see Pinus, Taxus, &c.). In the ?, the cone (to avoid for the present the word flower) consists typically of an axis bearing leaf-like organs. The most familiar case is Pinus (i/.v.), where each 1. borne on the axis is a small scale, bearing on its upper surface a very large scale (these latter show on the outside of the cone) on the upper side of which, again, are the two ovules. We may diagrammatically represent it thus, using ' cover-scale ' to express the lower, ' ovuliferous scale ' (epi in at in m] the upper, of the two scales. In Cryptomeria, &c. we find a large scale borne directly on the axis, with a little flap on its upper side near the outer end, and the ovules at the base. The flap is, by both theorists, supposed to represent the ovuliferous scale, and so we have what is illustrated by the second diagram. Then in Cuprcsseae, &c. we find only one scale, and here the two theorists differ. In the other ovulif. sc.l fain., Taxaceae, still further difficulties meet us. In cover-scale/ Microcachrys the ovule is borne upon a 1. of the cover-scale ovules ovulif. scale cover-scale ovules cone, but in Phyllocladus it is axillary and in Taxus ^"vuKf sc 1 term, (see these gen.). In most C. there is only cover-scale} one integument, but in Taxaceae a second commonly appears, forming an aril, ± fleshy, round the seed as it ripens. Now as to the explanation of the facts. Eichler regards the whole cone as one 2 fl. with a number of cpls. (the 'cover-scales'). The cpl. may bear the ovule directly, as in the latter cases above mentioned, or may develope upon its upper surface a placenta (ovulif. scale) which bears the ovules. Cryptomeria thus represents a stage in this evolution, and the whole may be compared with the division of a 1. into a sterile and fertile part, as in Ophioglossum. The rival theory of Celakovsky regards each ovule or pair of ovules with its appurtenances as a 9 fl. (one cpl. to each ovule or pair) so that the cone is a spike of fls. A series may be thus drawn : Podo- carpus (one cpl., one ovule with two integuments, the whole in the axil of a cover-scale, which is therefore to be regarded as a bract], Taxus (fl. reduced to ovule, aril = outer integument); then in the I'mafeae we have spikes of fls. (cones), the cover scale being the bract, the ovuliferous scale the combined outer integuments of the ovules of two cpls. (or three, the keel on the middle of the scale in Pinus, &c. repres. the third) : a fusion of the bract with the fl. in its axil is supposed to have gone on, and we get next the Cryptomeria type, and finally that of Cupresseae. Ovules orthotr., exc. Podocarpus. For development of the ovule, fert., &c., see text-books. The cone often becomes hard and woody as the seeds ripen ; in other cases it becomes fleshy. The seeds contain an embryo with i — 15 cotyledons, and rich endosp. Natural History. The C. are entirely wind-fert. ; the pollen is light and powdery, sometimes provided with air-bladders (e.g. Pinus), 172 CO NI FERAE and is produced in enormous quantities. About the time it is shed the scales of the ? cones open to receive it and the grains adhere to the sticky fluid at the apex of the ovule. Fert. often does not take place for a long time afterwards (see Finns). The seeds in many genera with woody cones (e.g. Firms) are winged for wind-carriage ; in other genera they are animal-distributed, e.g. Juniperus (cone fleshy), Taxus (fleshy aril), cic. Further details of morphology, life history, >S:c. under Gyi/mo- sperniae, Pinus and other gen. See also Retinospora for the peculiar case of pi. retaining the ' seedling' form throughout life. Economically the C. are most important, furnishing the greater proportion of our timber, as well as resins, tars, turpentines, &c. See gen., esp. Abies, Pinus, Larix, Tsuga, Libocedrus, Juniperus, Taxus, &c. Classification and Key to Genera (after Eichler and Engler) : Fam. i. TAXACEAE. Mostly dioecious ; cone-formation imper- fect ; cpls. usu. few or even i terminal, with i — 2 ov. each ; seeds projecting beyond cpls., or even naked, with fleshy aril or drupaceous testa; chiefly"^. A. Anther with i pollen-sacs; cpls. i — co , with i ov., often very small ; ovuliierous scale in all but Pherosphaera, often united to integument. I. PODOCARPOIDEAE. a. Ovuliferous scale o; ov. at base of cpl., erect; 1. scaly. i . Pherosphaereae. i. Pherosphaera. b. Ovuliferous scale present ; 1. usu. linear. i. Podocarpeae. a. Ov. scale and integument separate. I. Cpls. many; ov. scale membranous, little enlarged. 1. Cpls. whorled, thick, obtuse; seeds free; 1. scaly. i. Microcachrys. 2. Cpls. spiral, imbr. ; seeds in groove at base, small; fr. cpls. united; 1. linear. 3. Saxegothaea. II. Cpls. i — few ; ov. scale developed; young ov. ± pend., ultimately mostly upright, rarely stationary, enclosed in ov. scale. 4. Dacrydium. p. Ov. scale completely curved into itself, united with integument of pend. ov., falling with seed; cpls. usu. small, much shorter than ov. 5. Podocarpus. B. Anther with 2 loc. ; cpls. with i ov. ; seeds surrounded by cupule; twigs phylloclades with tooth-like 1. -rudiments. II. PHYLLOCLADOIDEAE. Only genus 6. Phyllocladus. C. Anther with 3 — 8 loc.; cpls. with 2 ov., or fl. reduced to i ov. term, on axis covered with scale 1.; ov. scale o; seeds with cupule, exc. in 7. III. TAXOIDEAE. a. ? fl. of several decussate cpls.; cpl. with 2 ov. i . Cephalotaxeae. 7. Cephalotaxus. CONIFERAE 173 b. v fl. reduced to i ov. term, on axis covered with scale 1. 2. Taxcae. a. S fl. in pairs in axil of 1., each with 4 decussate scales ; anther 4-loc. 8. Torreya. /3. ? fl. usu. single, only one of pair developed, the common axis with scale 1. ; anther 6 — 8-loc. 9. Taxus. Younger ? fl. unknown ; position doubtful, ? Podocarpoideae ; sta. 2-loc. ; 2 fl. term, or 2—3 at tip of twig; cpls. 1—2. 10. Acmopyle. Fam. 2. PINACEAE. Mostly monoecious ; cones perfect ; seeds concealed between scales; testa woody or leathery; no aril. A. L. spirally arranged ; sta. with several long pollen-sacs ; cpls. simple, with i reversed ov. in centre. I. Arancarieae. Seeds free from cpl. i r. Agathis. ,, united lo cpl. 12. Araucaria. P>. L. spirally arranged ; cover and ov. scales, the latter usu. large, with 2 ov. II. Abieteae. a. Long shoots only present. a. Needles 4-angled ; cone pend.; scales persistent. 13. Picea. /3. Needles flat. I. L. with i central resin canal; cone pend.; scales persistent. 14- Tsuga. II. L. with 2 lat. resin canals; cone pend. or erect; sta. fl. sol. or umbellate. 15. Pseudotsuga. III. Cone upright with persistent scales. 1 6. Keteleeria. IV. Cone upright with deciduous scales. 17. Abies. b. Long and short shoots both present. a. Long shoots with scale 1. only; the green (needle) 1. on the short shoots. 18. Firms. ft. Needle 1. on both shoots. I. Needles evergr. ; fr. 2 — 3 yrs. in ripening. 19. Cedrus. II. Deciduous; fr. i yr. in ripening. Cone scales persistent. 20. Larix. Cone scales deciduous. 21. Psettdolarix. C. L. spirally arranged ; scales rudimentary or weakly developed into cover and ov. scales; ovules 2—8, axillary and erect, or reversed on edges of cpls. III. Taxodieae. a. Long shoots with scale L; short shoots = double needles. 22. Sciadopitys. b. Long shoots only, a. Seed reversed. I. Cpl. minute. China, Cochinchina. Ovules 3. 23. Cunninghamia. Ovules 2. 24. Taiwania. 174 CONIFERAE II. Cpl. with ridge-like inner scale. Tasmania, Austr. 25. Athrotaxis. III. Cpl. shield-shaped, no distinct ov. scale. N. Am. 26. Sequoia. /3. Seed erect. I. Cpl. with toothed scale. 27. Cryptomeria. II. Cpl. shield-shaped, no distinct scale. Cone scales persistent. N. Am. 28. Taxodium. Cone scales deciduous. China. 29. Glyptostrobus. D. L. opp. or whorled, rarely alt., often heterophyllous; ovule erect. IV. Citpresseae. a. Cone woody when ripe ; cpls. valvate. i. Actinostrobinae. a. Branchlets compressed ; cones sol., axis not produced beyond scales ; scales 4 ; N. Afr. 30. Tetraclinis. /3. Branchlets angular ; cones paniculate, axis produced beyond scale's. Scales 6 unequal ; Austr. 31. Callitris. ,, 8 equal ; Austr. 32. Actinostrobus. ,, 4 equal ; trop. and S. Afr. 33. Widdringtonia. y. Branchlets flattened or angular ; dioecious ; i cones sol. term.; axis produced; sharp point on back of each scale near top ; Chili, Tasm. 34. Fitzroya. b. Cone woody when ripe, cpls. imbr. ; whorls of all fl. 2-merous. 2. Thujopsidinae. a. Cpl. with 4—5 seeds. 35. Thujopsis. j3. Cpl. with 2 (i — 3) seeds. Cpls. 4, upper pair fertile. 36. Libocedrus. Cpls. 6 — 8, both upper pairs fertile. 37. Thuja. Cpls. 6—8 pair. 38. Fokienia. c. Cone woody when ripe; cpl. peltate; whorls of all fls. 2-merous. 3. Cupressinae. a. Cpls. strongly woody when ripe. 39. Cupressus. p. Cpls. slightly woody when ripe; twig system flat in one plane. 40. Chamaecyparis. d. Cone, berry or drupe-like when ripe. 4. Juniperinae. Only genus. 41. Juniperus. Commitella Rydberg (Hatcheia p.p.). Saxifr. (i). i W. U.S. Coniog-eton Blume = Buchanania Roxb. (Anacard.). Coniogramme Fee. Polypodiaceae. 3 warm |*. Conioselinum Fisch. (Ligusticum L. p.'p. BH.}. Umbelliferae (in. 6). 7 N. palaeotemp. ConiumL. Umbelliferae (in. 4). 2 N. temp., S, Afr. C. maailatitm L. (hemlock, very poisonous) Brit. CONVALLARIA 175 Conjugate, coupled. Connaraceae (EP., BH.}. Dicots. (Archich. Resales, EP., BH.). 16 gen., 160 sp., trop. ; closely allied to Leguminosae, chiefly dis- tinguished by the absence of stips. and the (usual) presence of > 2 free cpls. Mostly twining shrubs with alt. exstip. 1. and panicles of reg. fls. K 5 or (5), imbr. or valvate ; 05; A 10 or 5 sometimes joined below ; G 5 or i or 4, each with 2 erect orthotr. ov. Fr. usu. one follicle with one seed, album, or not, arillate. Chief genera: Connarus, Rourea, Cnestis. Connaropsis Planch. Oxalidaceae. 5 Malay Arch. Connarus L. Connaraceae. 70 trop. Am., Afr., As. Connate (1.), concrescent, Loniccra. Connective, the prolongation of the filament into the anther ; elongated in Begonia. Connellia N.E. Br. Bromeliaceae (2). 2 Guiana. Connivent, converging. Conobea Aubl. Scrophulariaceae (n. 6). 7 Am. Conocarpus L. Combretaceae. 2 trop. Am., Afr. Conocephalus Blume. Moraceae (in). 15 Indomal. The 1. of C. sitaveolens Blume possess water-secreting glands. Conoclinium DC. = Eupatorium Tourn. p.p. (Compos.). Conornitra Fenzl (Glossonema BH.}. Asclep. (n. i). i Kordofan. Conomorpha A. DC. Myrsinaceae (11). 40 trop. Am., W.I. Conophallus Schott = Amorphophallus Blume p.p. (Arac.). Conopnaryngia G. Don (Tabernaeinonta.no. L. p.p.; Plitmeria Tourn. p.p. BH.}. Apocynaceae (i. 3). 25 trop. and S.E. Afr. Conopholis Wallr. Orobanchaceae. i Carolina to Mex. Conopnora DC. Nieuwland (Cacalia p.p.). Comp. (8). 8 N. Am. Conopodium Koch. Umbelliferae (in. 5). 20 Eur., As., N. Afr. C. deniidatum Koch (Bimiiim flexuosuin With.) in Brit, (earth nut). The tuberous roots are ed. when roasted. Conospermum Sm. Proteaceae (i). 35 Austr. Conostegla D. Don. Melastomaceae (i). 15 trop. Am. Conostephium Benth. Epacridaceae (3). 5 W. Austr. Conostylis R. Br. Amaryllid. (in). (Haemodor., BH.) 35 W. Austr. Conothamnus Lindl. Myrtaceae (11. i). 3 W. Austr. Conradia Mart. = Pentarhaphia Lindl. (BH.} — Gesneria L. p.p. Conradina A. Gray. Labiatae (vi). i Florida. Conringia Heist, ex L. Cruciferae (4). 6 E. Medit. Constantia Rodrig. (Sophronitis p.p.)- Orchid, (n. 6). i Braz. Consuegria Muds. Inc. sed. Nomen. Contabescent (anther), shrivelling. Contarinia Vand. Verbenaceae (?). i Brazil. Contortae (Engler), the 5th order of Sympetalae. Contorted (aestivation), cf. Aestivation. Contortoduplicate, twisted and folded. Contrayerva (W.I.), Aristolochia. Conuleum A. Rich. Monimiaceae. i Guiana. Convallaria L. Liliaceae (vn). i N. temp. (incl. Brit.), C. rnajalis*L., lily of the valley, in woods. The stock developes a few scales and :76 CONVALLARIA two green 1. annually. The fls. are homogamous and fert. themselves in absence of insects. Cult. orn. fl. Convallariaceae (Warming) = Liliaceae (§ vi — xi). Convolute (aestivation), cf. Aestivation. Convolvulaceae (EP., BH. incl. Nolanaceae). Dicots. (Sympet. Tubi- florac ; Polemoniales BH.}. 40 gen., iooosp., trop. and temp , many annual herbs, others shrubs or (rarely) trees; several thorny xero., many climbing herbs or lianes, and one (Cuscuta) a climbing parasite. Some have tuberous roots (e.g. Ipomoea Batatas), other rhiz. or tuberous stems ; latex often present. L. alt., usu. petiolate, rarely with stips. Accessory buds often found in the leaf-axils. Infl. dich. with tendency to cincinnus or bostryx ; br. and bracteoles present. Fl. 5, reg., hypog., usu. 5-merous. K usu. polysepalous, imbr., the odd sep. post.; C sympet., of various shapes, usu. induplicate- valvate, sometimes conv. ; A 5, alt. with pets., epipet., on base of C, with usu. intr. anthers; G on a honey-secreting disc, (2), rarely (3 — si), with axile plac. ; ov. 2 in each loc. (rarely 4), erect, anatr. or semi-anatr., the micropyle facing out \vards and downwards; one integument. Berry, nut or caps. ; endosp. Fls. usu. large, brightly coloured. Extra-floral nectaries in many on the petiole. Few of economic value exc. for handsome fls. (see Ipomoea). The C. are closely related to Solanaceae, Boraginaceae and other Tubiflorae. See Nat. Pfi. and esp. note by Engler at p. i of art. C- [BH. cnars. incl. 1. sometimes paired in infl.; G (2 — 5), or 5, divided by irreg. longitudinal constrictions ; fr. of nutlets.] Classification and chief genera : I. CONVOLVULOIDEAE (independent green pi.): Di- chondra, Evolvulus, Porana, Ipomoea, Convolvulus, Argyreia. II. CUSCUTO1DEAE (leafless parasites) ; embryo without cotyledons) : Cuscuta (only genus). Convolvulus (Tourn.) L. Convolvulaceae (i). 180 chiefly temp. C. arvcnsis L. Brit, (bindweed) has sweetly scented fls. which are much more often visited by insects than the large but scentless fls. of Calystcgia sepium. Smaller fls. with short sta. appear on some stocks ; these appear to be due to the action of a fungus (cf. Lychnis and see Beth. z. Bot. Centr., 1893, p. 447). The root produces adv. stem buds, by which veg. repr. takes place. From incisions made in the rhiz. of C. scammonia L. a resinous juice flows, used as a purgative (scammony). Some yield rosewood oil. Cult. orn. fl. Convolvulus, Calystegia, Convolvulus. Conyza L. Compositae (3). 60 temp, and subtrop. Conzattia Rose. Leguminosae (II. 7). i Mexico. Cooba (Austr.), Acacia salicina Lindl. Cookia Sonner. = Clausena Burm. f. (Rutac.). Coondi. Carapa. Cooperia Herb. Amaryllid. (i). 2 Mex., Texas (evening star). Copaiba Adans. =Copaifera L. (Legum.). Copaifera L. Leguminosae (n. 2). 25 trop. Am., Afr. Several CORDIFOLIUS 177 S. Am. sp. yield the resin Balsam of Copaiba, and resins (copals) are also obtained from the Afr. sp. Timber good (purpleheart). Copaiva tree (W.I.), Copaifera. Copal, a hard resiri, Agathis, Copaifera, Hymenaea, Trachylobiitm ; Manila, Kauri-, Agathis. Copernicia Mart. Palmae (i. 2). 10 trop. Am., W.I. C. cerifera Mart, (wax- or Carna-uba-palm, Brazil) has its 1. coated with wax, removed by shaking ; it is used in making gramophone records, candles, &c. The wood, 1., &c., are also useful. Copianthus Hill. Amarantaceae?. i Indomal. Copisma E. Mey. = Rhynchosia Lour. p.p. (Legum.). Copper-beech, Fagus sylvatica L. var. Copra, dried endosp. of Cocas nudfera L. Coprosma Forst. Rubiaceae (n. 7). 45 N.Z., Austr., Malaya, Chili. • The stipules of some are glandular, and some have peculiar openings (Pdomatia) on the backs of the 1. Copse, coppice wood with occasional standard trees. Coptis Salisb. Ranunculaceae (2). 10 N. temp, and arctic. Coptoclieile Hoffmgg. Gesneriaceae (inc. sed.). i Brazil? Coptophyllum Korth. Rubiaceae (i. 7). r Sumatra. Coptosapelta Korth. Rubiaceae (i. 5). 4 Malaya. Coptosperma Hook. f. (Tarenna p.p.'). Ruhiac. (i. 8). i trop. Afr. Coquilla, coquillio nut, Attalea fimifera Mart. Coquito palm, Jubaea spectabilis H. B. et K. Coral-berry (Am.), Symphoricarpvs \ -creeper, Kennedya; -root, Corallorhiza ; -tree, Erythrina. Coralliokyphos Fleischm. et Rech. Orchid, (u. 2). i Samoa. Corallobotrys Hook. f. Ericaceae (in. i). i Khasias. Corallocarpus Welw. ex Benth. et Hook. f. Cucurb. (2). 20 trop. Corallodiscus Batalin. Gesneriaceae (i). r N. China. Corallonema Schlechter. Asclepiad. (n. i). i Bolivia. Corallorhiza Hall. Orchidaceae (n. 4). 15 N. temp. C. innata K. Br. (coral-root) Brit. Saprophytes with much branched fleshy rhiz., no r., and scaly 1. [C/. Epipogum.] Corallospartium Armstrong. Leguminosae (in. 6). i N.Z. Corbularia Salisb. = Narcissus Tourn. p.p. (Amaryll.). Corchoropsis Sieb. et Zucc. Tiliaceae. 2 Japan, China. Corchorus (Tourn.) L. Tiliaceae. 40 trop. C. capsularis L. and C. olitoiins L. (India, &c.) furnish the chief supply of the fibre jute or gunny; annuals about 10 feet high, little branched. The stems are cut and retted in water, and the fibre beaten out (cf. Linum). Cord-grass, Spartina. Cordate, heart-shaped. Cordeauxia Hemsl. Leguminosae (n. 3). i trop. Afr. Cordia L. Boragin. (i). 270 trop. Trees or shrubs; fr. ed.; that of C. Myxa L. (sebestens ; Egypt to Austr.) formerly medic. Some have good timber, e.g. C. Gerascanthits L. (trup. Am., W.I., prince- wood), and C. sebestana L. (trop. Am., W.I., aloewood). [Mez in Engl. Jahrb. XII.] Cordiaceae ( Warming) = Boraginaceae (§ i). Cordifolius (Lat.), cordate-leaved. W. 12 178 CORDOBIA Cordobia Niedenzu. Malpighiaceae. i S. Am. Cordyla Lour. Leguminosae (11. 9). i trop. Afr. Pods ed. Cordylanthus Nutt. (Adenostegia Benth.). Scroph. (in. 3). 12 N. Am. Cordyline Royen ex Adans. Liliaceae (vi). 15 trop., warm temp. Decorative ; habit of Dracaena. The 1. of some sp. yield fibre. Cordylocarpus Desf. (Rapistrum p.p. BH.}. Crucif. (2). i N. Afr. Cordylogyne E. Mey. Asclepiadaceae (n. i). i S. Afr. Corema D. Don. Empetraceae. 2 W. Eur., All. N. Am. Coreocarpus Benth. (Coreopsis f.^. EP.). Comp. (5). i California. Coreopsis L. Compositae (5). 80 Am., trop. Afr., Sandw. Is. Many cult. orn. ft Corethrodendron Fisch. et Basiner. Legum. (in. 7). i Soongaria. Corethrogyne DC. Compositae (3). 4 Calif. Coriaceous, leathery. ' Coriander, Coriandrum sativiitn L. Coriandrum (Tourn.) L. Umbelliferae (in. 3). 3 Medit. The fr. (coriander-seeds) of C. sativum L. are used in flavouring. Coriaria Niss. ex L. The only genus of Coriariaceae. 10 Medit. to Japan, N.Z., Chili to Mexico, mostly shrubs with opp. or whorled 1., sometimes becoming alt. at the ends of the shoots. The inconspic. protog. fls. are in racemose infls. K 5, C 5, A 5 + 5, 65. The petals are keeled on the inner side, and after fert. grow fleshy and enclose the cpls. forming a pseudo-drupe. Ov. i in each loc., pend., anatr. ; raphe dorsal. Enclosp. thin. C. myrtifolia L. (W. Medit.) yields tan, others a black dye. Coriareae (BH. ) = Coriariaceae. Coriariaceae (EP., BH.). Dicots. (Archichl. Sapindales). Only genus Coriaria (q-v.}. The only nearly related order is Empetraceae. Placed as anomalous order at end of Disciflorae BH. Coridothymus Reichb. f. (Thy inns p.p. BH.}. Labi. (vi). i Medit. Corion Mitch. = Spergularia J. et C. Presl (Caryoph.). Coriophyllus Rydberg (Cymoptems p.p.). Umbel, (in. 6). 4 N. Am. Coris Tourn. Primulaceae. 2 Medit. Fl. •!•• Corispermum B. Juss. ex L. Chenopodiaceae (A). 16 N. temp. Cork, the substance of which the water-tight bark is composed ; -oak, Quercus suberL,.; -tree, Indian, Millingtonia liortensis L. f . ; -wood, Ochroma, (Am.) Leitneria, (W.I.) A nona palitstris. Corm, the base of a stem swollen into a bulbous shape with reserves ; Colchicum (and fig.), Amorphofhallns, Arisaema, Boiviea, Crocus, Cyclamen, Eranthis, Hablitzia, Testudinaria. Cormonema Reissek. Rhamnaceae. 4 trop. Am. Cormophyte, a flowering plant or fern. Corn, in Engl. wheat, in Am. maize; broom-, Sorghum mtlgare Pers. ; Chinese- (W.I.), Setaria ilalica Beauv.; -cockle, Lychnis Gilhago Scop. ; -flower, Centaurea Cyanus L. ; Guinea-, Sorghum vulgare Pers.; Indian-, Zea Mays L. ; Kaffir-, Sorghum vulgare Pers.; -marigold, Chrysanthemum segetutn L. ; -salad, Valerianella. Corna Nor. Inc. sed. Nomen. Cornaceae (EP., BH. incl. Ganyaceae, Nyssaceae}. Dicots. (Arcliichl. CORREA 179 Umbelliflorae ; Umbellales BH.}. logen., 85 sp., N. and S. temp., and trop. Mts, Shrubs with opp. or rarely alt. leaves, usu. entire, exstip. Infl. dich., usu. condensed into corymbs or umbels, or even (Cornus) heads with invols. Fls. usu. §, reg., 4 — 5 — oo -merous. Iv 4 — 5, €4 — 5, A 4 — •-,, G(2); epig. disc ; simple style with lobed stigma; ovary i — 4-loc., with usu. i pend. ov. in each loc., the raphe dorsal. Fr. a berry or more often a drupe, with a i — 4-loc. stone or 2 separate stones. [BH. chars, include those of Garryaceae and Nys- saceae, both rare.] Chief genera: Cornus, Aucuba, Griselinia. Cornel, Cornus sanguined L. Cornelian cherry, Cornus mas L. Cornelia Rydberg (Cornus p.p.). Cornaceae. 3 N. temp, and arct. Corniola Adans. = Genista L. (Legum.). 1 Cornish moneywort, Sibthorpia eiiropaea L. Corniveum Nieuwland (Dicentra p.p.). Papav. (in). i N.W. Am. Cornucopiae L. Gramineae (8). i E. Medit., C. ciicullatiim L. Fls. in small heads ; when the fr. is ripe these bend over and break off with a sharp point ; they adhere to animals and are also said to burrow into the soil (cf. Stipa). Cornuella Pierre. Sapotaceae (inc. sed.). i Venezuela. Cornulaca Del. Chenopodiaceae (B). 5 Egypt to Afghanistan. Cornus (Tourn.) L. Cornaceae. 48 N. temp.; 2 Brit., C. sanguined L. (cornel or dogwood) and C. suecica. L., the latter a dwarf perenn. herb (Highlands). Annual stems are given off from the creeping perenn. stems. Fls. in umbels with invols. of 4 large white br. C. florida L. (N. Am.) and others yield useful wood. C. mas (mas- ciila) L. (Cornelian cherry, Eur., As. Minor) yields a fr. which makes good preserves. Its fls. appear in spring before the 1. Cornutia Plum, ex L. Verbenaceae (4). 8 trop. Am., W.I. Corokia A. Cunn. Cornaceae. 3 N. Zealand. Corolla, the_Lnner, usu. coloured, envelope of 1. of the fl. Corolline, belonging to the corolla. Coromandel wood, Diospyros quaesita Thw., &c. Corona, Amaryllidaceae, Asiiepiadaceae, Napoleona, Passifloraceae. Coronanthera Vieill. Gesneriaceae (i). 10 New Caled. Coronaria L. = Lychnis L. p.p. (Caryoph.). Coronarieae (BH.}. The 3rd series of Monocotyledons. Coronilla Tourn. ex L. Leguminosae (in. 7). 20 Eur., Medit., \V. As. The fl. resembles Lotus, but the honey is usu. secreted by the outer surface of the K, and insects poke between the claws of the petals. The buds are bent downwards, the open fls. project horiz., the ripening fr. down and the ripe fr. usu. up. Coronopus Rupp. ex L. (Senebiera DC.). Cruciferae (2). 20 subtrop., Eur. Coroya Pierre. Leguminosae (ill. 8). i Cochinchina. Corpse-plant (Am.), Monotropa. Corpusculum, Asclepiadaceae. Correa Andr. Rutac. (i). 5 temp. Austr. Cult. orn. shrubs (Australian fuchsia). Sympet. 12 2 i So CORRIGIOLA Corrigiola L. Caryophyllaceae (i. 4). 12 Medit., Ancles, S. Afr., Eur. i Brit. Corsia Becc. Burmanniaceae. i New Guinea. Corsiaceae (Beccari), Burmanniaceae p.p. Corsican pine, Pin us Laricio Poir. Cortaderia Stapf. Gramineae do). 10 S. Am. Pampas grass. Cortesia Cav. Boraginaceae (n). 2 temp. S. Am. Cortex, tissue between vascular bundles and epidermis. Cortia DC. Umbelliferae (ill. 5). 2 Himalaya. Cortusa L. Primulaceae. i Mts. of Eur. and As. Cult. orn. fl. Corunastylis Fitzgerald. Orchidaceae (n. 2). i Austr. Coryanthes Hoolc. Orchidaceae (n. 13). 6 trop. S. Am., epiphytic. Fl. pend.; seps. bent back and fairly large, pets, small. Labellum complex, forming a bucket-like organ with dome above ; the mouth faces upwards, and the edges are incurved ; there is also an overflow pipe projecting towards the seps. and closely covered in by the bent end of the column, with the stigma and anther. From the base of the column project two horns which secrete a thin watery fluid that drips into the bucket, keeping it full to the level of the overflow pipe. The dome (above) is composed of succulent tissue attractive to bees ; these light for places on it to drill the tissue ; every now and then one gets pushed off and falls into the bucket. It can neither fly nor climb out, and has to squeeze through the overflow pipe. In so doing it first passes the stigma, fertilising it if it bears any pollen, and then, passing the anther, is loaded with new pollinia. [Darwin's Orchids, p. 173, and cf. Stanhopea.] Corybas Salisb. = Corysanthes R. Br. (Orchid.). Corycium Swartz. Orchidaceae (n. i). 10 S. Afr. Corydalis Vent. Papaveraceae (in). 120 Medit., Eur., As. C.clavi- cidata DC. Brit., a (leaf) tendril-climbing annual. Most are perennial herbs with underground tubers. In C. cava Schweigg. et Kort., and others, the main axis forms a tuber, which dies away below, each annual shoot arising from the axil of a scale-1. of older date. In C. solida Sw., and others, the tuber is a swollen root-structure be- longing to the current annual shoot. Fls. transv. -|- (see fam. for diagram) ; twisting through 90° brings it vertical ; only one petal is spurred and contains the honey secreted by a staminal outgrowth. Its mech. resembles that of Leguminosae. The inner pets., united at the tip, enclose stigma and anthers ; the upper pet. covers the fl. Bees alighting push down the inner pet. and cause the essential organs to emerge. In some, e.g. C. oc/iroleuca Koch and C. In tea DC., the emergence is explosive (cf. Genista). The fls. of C. cava are self- sterile. Corylopsis Sieb. et Zucc. Hamamelidaceae. 10 China, Japan. Fls.?, in spikes with coloured bracts at base. Corylus (Tourn.) L. Betulaceae (i). 8 N. temp. C. avella>ia L., hazel-nut (Brit.). Shrubby (largely owing to extensive formation of suckers), with monoecious catkinale fls. (the ? catkin sessile and elliptical in outline, rather resembling a bud). Both are laid down in autumn ; the S catkins are visible all winter, but the ? are not obvious until the red stigmas come out early in the year. Anemoph. ; COR YTHOL OMA 1 8 1 the fact of fig. before the appearance of the 1. renders their chance of fert. greater. On the inner side of the br. in the $ catkin are found 2 scales and, adnate to these, 4 — 8 sta., each branched nearly to the base. Here only the central stem fl. of the possible 3 (cf. diagram of fam.) is pre- % 3 ft. % sent, with its bracteoles a, j3. In the ? catkin, a /3 on the other hand, we have the two lat. and not bract the central fl. as shown in the second diagram (* = missing fl.). At the time of fert. the ovary stem is minute, but the long red stigmas are easily a a' identified. After fert. the ovary (a-loc. at first) a ? (1. % ? fl. /3 gives a one-seeded nut. enclosed in a cup of /?' /3' green leafy nature, really the combined bract bract and bracteoles a, a', /3', very much developed. The fl. is chalazogamic (cf. Chalazoganiae}. The nuts of this and other sp. are valuable as dessert fr., &c. (hazel-nut, cob-nut, filbert), and have been cultivated from very early limes (cf. Goeschke, Die Hasclnnss, Berlin, 1887). Wood elastic, but cannot be obtained in large boards. Oil from the seeds. Corymb, a raceme in which the stalks of the lat. fl. elongate to bring all to one level, Cruciferaet Iberis. Corymbis Thou. Orchidaceae (ll. i}. 12 trop. Corymbium L. Compositae (i). 7 S. Afr. CorymborcMs Thou. (Corymbis BH.}. Orchid. (11. 4). 16 trop. Corymbostachys Lindau. Acanthaceae (iv. B). i Madag. Corynaea Hook. f. Balanophoraceae. 4 Andes. Corynanthe Welw. Rubiaceae (i. 5). ^ trop. Afr. Corynella DC. Leguminosae (in. 6). 3 W.I. Corynephorus Beauv. (IVciiigacrtneria Bernh.). Gramineae (9). 3 Eur. C. (IV. or Aim} catiescens Beauv., Brit. Corynephyllum Rose. Crassulaceae. i Mexico. Corynitis Spreng. Leguminosae (in. 6). 2 Cuba. Corynocarpaceae (EP., Anacardiaceae p.p. BH.}. Dicots. (Archichl. Sapindales). Only genus Corynocarpus, tj.r. Corynocarpus Forst. Corynocarpaceae. 3 New Hebrides, New Caled., N. Zealand. Trees or shrubs with alt. leathery 1. and panicles of $ fl. A in two whorls, inner sta. stds. ; G (2), one fertile, with one pend. ov. Compressed drupe. No endosp. Corynostylis Mart. (Calyptrion EP.}. Violaceae. 2 trop. S. Am. Corynotheca F. Muell. Liliaceae (in). 3 trop. and W. Austr. Corynula Hook. f. Rubiaceae (n. 7). i Colombia. Corypha L. Palmae (i. 2). 6 Ceylon, Indomal. The gigantic infl. terminates the life of the tree. C. umbraeulifera L. (talipot palm, Ceylon, S. Ind.) grows to a great size, up to 80 feet. The 1. are used as umbrellas, and for thatching, also as writing material (a metal stylus being used). Corypriantha Lem. = MamilIaria Haw. p.p. (Cact.). Coryphlum (CL), an alpine meadow formation. Corysanthes R. Br. Orchidaceae (n. 2). 20 N.Z. to Malaya. Corythea S. Wats. Euphorbiaceae (A. 11. 7). i Mexico. Corytholoma Decne. (Gesneria p.p.). Gesn. (u). 60 trop. Am. 1 82 COSCINIUM Coscinium Colebr. Menispermaceae. 4 Indomal. Cosmanthus Nolle = Phacelia Juss. p.p. (Hydrophyll.). Cosmea Willd. = Cosmos Cav. (Compos.). Cosmelia R. Br. Epacridaceae (2). i S.W. Austr. Cosmibuena Ruiz et Pav. (1802). Rubiaceae (i. 5). 6 trop. Am. Good bud-protection by the slips, of the last-opened 1. Cosmibuena Ruiz el Pav. (iy94) = Hirtella L. (Rosaceae). Cosmos Cav. (Cosmea Wi lid.). Compos. (5). 20 Am., W.I. Orn. fl. Cosmostigma Wight. Asclepiadaceae (11. 3). i Indomal. Cossinia (Cossignid) Comm. ex Lam. Sapind. (n). 3 Masc., New Cal. Cossonia Durieu (Kaffcnaldia BH.}. Cruciferae (2). 3 N. Afr. Costaea A. Rich. (Purdica BH.}. Cyrillaceae. 3 Cuba, Colombia. Costate (diminutive costulate), ribbed. Costera J. J. Smith. Ericaceae (in. i). i Borneo. Costularia C. B. Clarke. Cyperaceae (n). 9 Masc., S. Afr. Costus L. Zingiberaceae (n). 40 trop. Labellum very large, lat. stds. wanting, sepals and petals comparalively small. Projecting in the centre is the fertile petaloid sta. with anther on its ant. face ; the style reaches just above this. The fl. mech. thus resembles Iris. Cull. orn. pi. Cota J. Gay — Anlhemis Mich. p.p. (Compos.). Cotinus (Tourn.) L. (Rhus L. p.p. BH.}. Anacard. (3). 2 N. temp. Cotoneaster Rupp. Rosaceae (n). 30 N. temp. C. vulgaris Lindl. Brit. In the Alps visited solely by a wasp \Polistes gallica] whose nests are often attached to the rocks where the pi. grows. Fl. protog. with self-fert. in default of insects. Several cult. orn. shrubs. Cottea Ktmth. Gramineae (10). i trop. Am. to New Mexico. Cottendorffla Schult. f. Bromeliaceae (2). 2 trop. S. Am. Cotton, the fibre of Gossypiton, and loosely, of other IWalvaceae and Boinbacaceae. The fibre most used, and extensively cull, in ihe U.S., India, Brazil, trop. Afr., Japan, W. L, &c. It is the fine silky fibre enveloping the seed, and in G. barbadense and others comes clean away, but in G. herbaceum and most spp. leaves a ' fuzz ' adhering to ihe seed. There are many grades upon ihe market : the longest and finest 'staple' is that of Sea Island, G. barbadense L., and the next best Egyptian and some Brazilian and American. The bulk of the supply is American ' Upland,' and the poorest qualities come from India. For details see Watt, Wild and Cult. Cotton Plants of the World; Willis, Agriculture in the Tropics, -grass, Eriophoriuii ; -rose (Am.), Filago ; -sedge, Eriophornm ; silk-, Eriodendron, Chorisia ; -thistle, Onopordon ; -tree, Bombax ; -weed, Diotis ; -wood tree, Fopitlns. Cottonia Wight. Orchidaceae (n. 20). 2 Ceylon to China. Cottony, with sofl long hairs. Cotula (Tourn.) L. (incl. Ceiiiajnss. EP.}. Composilae (7). jocosmop., esp. 1^~. Cull. orn. fl. Cotylanthera Blume. Genlianaceae (i). 3 E. Indomal., Mariannes. Cotyledon Tourn. ex L. (incl. Echercria DC.). Crassulaceae. 100 Afr., Eur., As., Mexico, S. Am. C. Umbilicus Linn, (penny-wort) Brit. Pets, uniled inlo a lube. L. succulent. Cult. orn. fl. Cotyledon, a seed 1. CRAB GRASS 183 Cotylelobiopsis Heim. Dipterocarpaceae. i Borneo. Cotylelobium Pierre. Dipterocarpaceae. 5 Indomal. Cotyliform, dish- or wheel-shaped. Cotylodiscus Radlk. Sapindaceae (i). i Madagascar. Cotylonychia Stapf. Sterculiaceae. i trop. Afr. Coublandia Aubl. (Muellera L. f. ). Legum. (in. 8). 3 trop. Am. Couch grass, Agropyron repens Beauv. Coudenbergia March (Pentapanax EP.}. Araliac. (2). i Brazil. Couepia Aubl. Rosaceae (vi). 45 S. Am. Coula Baill. Olacaceae. 2 trop. Afr. Coulterella Vaizey et Rose. Compositae (5). i Lower Calif. Coulterophytum Robinson. Umbelliferae (in. 6). 5 Mex. Coiuna Aubl. Apocynaceae (i. i). 5 Brazil, Guiana. Coumarin, Anthoxantlnitn. Coumarouna Aubl. (Dipteryx, g.v.). Legum. (in. 8). 8 trop. Am. Country almond (Ceylon), Terniinalia Catappa L ; -walnut, Alatritei triloba Forst. Coupoui Aubl. Rubiaceae (inc. sed.) (Apocyn. Bff.}. i Guiana. Couralia Splitg. Bignoniaceae (2). 4 trop. S. Am. Courantia Lemaire (Cotyledon Tourn.). Crassul. i Mexico. Couratari Aubl. Lecythidaceae. 8 S. Am. The bark yields a soft fibre used for making clothing. Courbonia ISrongn. Capparidaceae (n). 5 trop. Afr. Courimari Aubl. Inc. sed. i Guiana. Couroupita Aubl. Lecythid. 9 trop. S. Am., W.I. The fls. of C. guianensis Aubl. are borne on the old stems and followed by large spherical woody caps, (whence the name cannon-ball tree). Good timber. Coursetia DC. Leguminosae (in. 6). 15 Brazil to Calif. Courtoisia Nees. Cyperaceae (i). 2 trop. As., Afr. Cousinia Cass. Compositae (n). 240 E. Medit., W. As. Coussapoa Aubl. Moraceae (in). 18 trop. S. Am. Coussarea Aubl. Rubiaceae (n. 8). 45 Brazil, Venezuela. Coutarea Aubl. Rubiaceae (i. 5). 6 trop. Am., W.I. Couthovia A. Gray. Loganiaceae. 5 Polynesia, Malay Arch. Coutoubea Aubl. Gentianaceae (i). 4 trop. S. Am., W.I. Covellia Gasp. = Ficus Tourn. p.p. (Morac.). Cover-scale, Coniferae. Covilhamia Kunth. Sterculiaceae. i Borneo. Covillea Vail. = Larrea Cav. (Zygophyll.). Cow-bane, Cicuta virosa L. ; -berry, Vacciniuin Vitis-Idaea L. ; -itch, Mucunaprurtens'DC.; -parsnip, Heracleum Sphondyliuin L. ; -pea, Vigna sinensis Endl.; -tree, Brosimum', -wheat, Melampyrum, Cowage, Mitcnna pruriens DC. Cowania D. Don. Rosaceae (in. i). 3 Mexico, S.W. U.S. Cowiea Wernham. Rubiaceae (i. 8). i Borneo. Cowrie pine, Agathis australis Steud. Cowslip, Primula vtris Lehm. Coxella Cheesem. et Hemsl. Umbellif. (in. 5). i Chatham Is. Crab grass (Am.), Panic urn, Eleusine; -'& eyes, Abrus precatorius L. ; -wood (W.I.), Carapa guianensis Aubl. 184 CRABBEA Crabbea Harv. Acanthaceae (iv. A). 12 trop. and S. Afr. Cracca Benth. ex Oerst. Leguminosae (in. 6). 6 trop. Am.; do. L. = Tephrosia Pers. ; do. (Riv.) Medic. = Vicia Tourn. p.p. Craibia Harms et Dunn. Leguminosae (in. 6). 15 trop. Afr. Craibiodendron W. W. Smith. Ericaceae (n. i). i Burma. Crambe Tourn. ex L. Cruciferae (2). 20 Eur., Meclit., As., Polynes., Patagonia. C. maritima L. (sea-kale) on coast of Brit, has 1. fleshy and waxy. The young 1. blanched form a veg. Cranberry, Vaccinium Oxycvccus L. Crane's bill, Geranium pratense L., &c. CranicMs Sw. Orchidaceae (11. 2). 25 trop. Am., W.I. Craniolaria L. Martyniaceae. 3 S. Am. Craniospennum Lehni. Boraginaceae (iv. 2). 3 temp. As. Craniotome Reichb. Labiatae (vi). i Himalaya. Cranocarpus Benth. Leguminosae (ill. 7). 2 Brazil. Crantzia Scop. (Alloplectus Mart. EP.). Gesneriaceae (i). 35 trop. Am.; do. Nutt. Umbelliferae (in. 5). i Am., Austr., N.Z.; do. Pohl. Inc. sed. Nomen. Craspedia Eorst. f. Compositae (4). 4 temp. Aust., JM.Z. Craspedodictyum Copeland (Gynmogramme p.p.). Polypodiaceae. 2 Malay Arch., Polynesia. Craspedorachis Benth. Gramineae (n). 2 trop. Afr. Craspidospermum Boj. ex DC. Apocynaceae (i. i). i Madag. Crassocephalum Moench. (Gytiura EP.}. Comp. (8). 20 warm Afr., As. Crassula Dill, ex L. (incl. Dinacria Harv., and Til/aea Michx. EP.). Crassulaceae. 150 S. Afr., a few Abyss, and Himal., chiefly succu- lent 1. xero. In C. lycopodioides Lam. the 1. are narrow and closely packed, giving to the pi. the habit of a Lycopodium. In C. {Rochea) falcala Wendl. the connate decussate 1. stand almost edgewise, and are very fleshy ; some of the epidermal cells are swollen above the rest into large bladders which meet one another over the whole surface. At first living, when the 1. is mature they are dead and full of air, their walls infiltrated with quantities of silica. A pro- tection against evap. is thus afforded. In C. ncniorosa Endl. there is veg. repr. by the formation of young plants in the infl. in place of fls. Crassulaceae (EP., BH.). Dicotyledons (Archichl. Resales -£/"., BH.). 15 gen., 450 sp. cosmop., chiefly S. Afr., a very natural group. Most are perenn. living in dry (esp. rocky) places and exhibit xero. chars., fleshy 1. and stem, often tufted growth, close packing of 1.. waxy surface, sunk stomata, &c. Veg. repr. frequent ; usu. by rhiz. or offsets ; some form bulbils, &c. (e.g. Crassula), others form adv. buds upon the 1. (e.g. Bryophyllum). Fls. usu. in cymes (cincinni), 5 or rarely unisex., actinom. with very reg. construction. Formula K«, C«, A M + «, Gn, where n represents any number from 3 to 30. K persistent; C sometimes (e.g. Cotyledon) gamopet.; A frequently obdipl. Insertion of parts usu. perig.. but recept. not deeply hollowed. Cpls. frequently slightly united at the base; at the base of each commonly a honey-secreting scale; ov. usu. . Fr. usu. a group of follicles with very small seeds. Endosp. none or very little. Fls. mostly protandr. and chiefly visited by flies, &c., their honey being C RE PIS 185 easily obtainable. Chief genera: Sedum, Sempervivum, Cotyledon, Bryophyllum, Crassula. Crassus (Lat.), thick. Crataegus Tourn. ex L. (Mespilns EP. The boundaries of these two genera and of Pyrus are ill-defined.) Rosaceae (n). 100 N. temp. Some hundreds of spp. have in recent years been described from the U.S., but there is some possibility that they may arise through hybridisation (cf. Journ. of ffered., June 1916). C. Oxyacantha L. (hawthorn or may) in Brit. The thorns are modified branches. Collateral buds appear in the axils. Fls. of class AB. The wood is a substitute for that of box in engraving, &c. Crataemespilus x G. Camus. Hybrid, Crataegus — Mespilus. i Eur. Crataeva L. Capparidaceae (11). 10 trop. Crateranthus E. G. Baker. Lecythidaceae. i Nigeria. Crateriform, cup-shaped. Crateriphytum Scheff. ex Koord. Loganiaceae. i Moluccas. Craterispermum Benth. Rubiaceae (n. i). 10 trop. Afr., Seychelles. Cratero siphon Engl. et Gilg. Thymelaeaceae. 2 Cameroons. Craterostemma K. Schum. Asclepiadaceae (n. 3). i S.E. Afr. Craterostigma Hochst. Scrophulariaceae (n. 6). 5 trop. and S. Afr. Cratoxylon Blume. Guttiferae (11). 12 Indomal. Cratylia Mart. Leguminosae (m. 10). 6 S. Am. Cratystylis Sp. Moore (Glearia p.p. in part.). Comp. (3). 3 Austr. Crawfurdia Wall. Gentianaceae (i). 10 warm As. Creaghia Scortech. (Mussaendopsis EP.). Rubiac. (i. 5). i Malaya. Creaghiella Stapf. Melastomaceae (i). i Borneo. Creeper, Canary, Tropaeolimi peregrinnm L. ; trumpet-, Tecoma radicans Juss. ; Virginian-, Parthenotissns. Creeping plant, one with stolons or runners, rooting at the nodes, Agropyron, Agrostis, Ajiiga, Fragaria, Ranunculus, &c. Cremanium D. Don = Miconia Ruiz et Pav. p.p. (Melast.). Cremanthodium Benth. Compositae (8). 15 Himal., China. Cremaspora Benth. Rubiaceae (n. i). 6 trop. Afr., Madag. Creraastra Lindl. Drchidaceae (n. 10). 2 Mepal, Japan. Cremastus Miers (Bignonia BH.}. Bignon. (i). 3 Brazil. Crenmium (Cl.), a cliff formation. Cremnophila Rose (Sedum p.p.). Crassulaceae. i Mex. Cremocarp, a mericarp. Cremocarpus Boiv. ex Baill. Rubiaceae (n. 7). i Comoros. Cremolobus DC. Cruciferae (i). 10 Andes. Crenamon Raf. Inc. sed. i N. Am. Crenate (L), teeth pointing forward, rounded ; notches sharp. Crenea Aubl. Lythraceae. 2 trop. Am. Crenium (CL), a spring formation. Creochiton Blume. Melastomaceae (i). 3 Java, Phil. Is. Creosote plant, Larrea mexicana Moric. Crepe-flower, Lagerstroemia, Crepidopsis Arv. Touv. Compositae (13). i Mex. Crepidospermum Hook. f. Burseraceae. 2 Northern S. Am. Crepinella Marshal. Araliaceae (i). i Brit. Guiana. Crepis (Vaill.) L. (incl. Barkhansia Moench.). Compositae (13). 1 86 CRE PIS 220 N. Hemisph., S. Afr., S. Am.; 6 in Brit, (hawk's beard). Like Hieracium. Crescentia L. Bignon. (iv). 5 trop. Am. Fls. on old stems, succeeded by gourd-like berries ; the epicarp is woody, and after removal of the pulp forms a calabash (C. Cnjete L., calabash tree, most used). Cress, Lepidium sativitin L. ; American-, Barbarca praecox R. Br. ; bitter-, Cardamine ; Indian-, Tropaeoluni; penny-, 7Yi/aipi; rock-, Arabis; water-, Nasturtium officinale R. Br. ; winter-, Barbarea; yellow-, Barbarea, Nasturtium. Cressa L. Convolvulaceae (i). 5 trop. and subtrop. Crest, a ridge or outgrowth. Cretaceous, chalky. Cribriform, sieve-like. Crinitus (Lat.), with soft hairs. Crinodendron Molina (Tricuspidaria R. et P.). Elaeocarp. 2 Chili. Crinum L. Amaryllidaceae (i). 70 trop. and subtrop., esp. on sea- coasts. Large bulbous pi. with showy fls. The seed of C. asiaticum L. (Goebel, Pftanzenbiol. Schild. \. p. 128) has a very thin corky covering and is suited to distr. by water and early germination. The ovule has no integuments, and the testa is replaced by a formation of cork at the outside of the endosp. Cult. orn. fl. Crioceras Pierre (Tabernaemontana p.p.). Apocyn. (i. 3). 2 trop. Afr. Crisp, crispate, crisped, curled. Cristaria (Heist.) Cav. Malvaceae (2). 30 Chili, Peru. Cristatella Nutt. Capparidaceae (v). 2 S.W. U.S. Cristatus (Lat.), crested. Crithmum L. Umbelliferae (in. 5). i C. niaritimiiin L., the samphire, on rocky coasts, Medit., Eur. (incl. Brit.). It has much divided and very fleshy 1. Used for making pickles. Crocidium Hook. Compositae (8). i W. N. Am. Crocion Nieuwland (Viola p.p.). Violaceae. 2 N. Am. Crockeria Greene ex A. Gray. Compositae (6). i Calif. Crocodiloides Adans. = Berkheya Ehrh. (Compos.). Crocopsis Pax. Amaryllidaceae (i). 2 S. Am. Crocosmia Planch. (Tritonia p.p. EP.}. Iridaceae (ill), i, C. aurea Planch., trop. and S. Afr. Cult. orn. fl. Crocus (Tourn.) L. Iridaceae (i). 60 Medit., Eur., 2 nat. in Brit. Below ground is a corm (cf. Colchicum), covered with a few scaly L, in whose axils may arise one or more buds, giving rise to new corms on the top of the old. The 1. are dorsiv., and curiously grooved on the back. The fl. is often single and term.; in some sp. there is a small cyme. The fl. closes at night and in dull weather. The tube of the P is so long that the ovary remains below the soil and is protected from the weather (cf. Colchicum). The fl. is protandr. and visited by bees and Lepidoptera. Honey is secreted by the ovary, and the anthers face outwards so as to touch any insect alighting on the petals and seeking honey. The stigmas are branched. Birds often bite off the fls. in gardens (? for honey) ; they seem to prefer the yellow fls., leaving the blue and white alone. The dried stigmas of C. sativits L. form saffron, once largely used as an orange yellow dye, but now chiefly employed in flavouring and CROWN-BARK 187 colouring dishes, liqueurs, &c. [See Kronfeld's Ceschichte des Sa/rans, &c., Wien 1892, or Beih. z. Dot. Centr., 1893, p. 71.] Crocus, autumn, Colchicum attfumnale L. Crocyllis E. Mey. Rubiaceae (n. 7). i S. Afr. Croftia King et Prain. Zingiberaceae (l). i India. Croftia Small (Schaueria p.p.)- Acanth. (iv. B). r N. Am. Croixia Pierre. Sapotaceae (n). i Malaya. Crookea Small (Ascyrum p.p.)- Guttif. (n). i N. Am. Croomia Torr. ex Torr. et A. Gray. Stemonaceae. 3 E. U.S., Japan. Crosnes, Stachys Sicboldi Miq. Crossandra Salisb. Acanthaceae (iv. B). 17 trop. As., Afr., Madag. The seeds of many sp. are covered with scales which spread out and become sticky when wetted (cf. Linum). Cult. orn. fl. Cross-fertilisation or -pollination, pollination from a distinct plant ; cf. Floral mechanisms, Flower classes, &c. Cross-wort, Galiitm. Crossandrella C. B. Clarke. Acanthaceae (iv. B). i Uganda. Crossonephelis Baill. Sapindaceae (l). r Nossi Be. Crossopetalum P. Br. = Myginda L. (Celastr.). Crossopteryx Fenzl. Rubiaceae (i. 5). i trop. Afr. Crossosoma Nutt. The only genus of Crossosomataceae. 3S.W. U.S., Mex. Shrubs with small stiff 1. and sol. 11. Like Rosaceae Spiraeoi- deae, but seeds kidney-shaped, with rich endosp. Aril. (Engler.) Crossosomataceae (EP.; Dillcniaceae p.p. BH.). Dicots. (Archichl., Rosales). Only genus (q.v.} Crossosoma. Crossostemma Planch, ex Benth. in Hook. Passifl. i W. trop. Afr. Crossostephium Less. Compositae (7). i China, Phil. Is. Crossostylis Foist. Rhizophoraceae. 6 Polynesia. Crossotropis Stapf. Gramineae (n). 3 trop. Afr., Arabia. Crotalaria Dill, ex L. Leguminosae (in. 3). 350 trop. and subtrop. C. jnncea L. (India, Austr.), an annual about 8 ft. high, is largely cult, for the fibre obtained from its stems by maceration in water (cf. Linum), known as Sunn-hemp, Bombay or Madras hemp, &c. C. retusa L. (trop.) is also employed. Croton L. Euphorbiaceae (A. II. i). 600 trop. and subtrop. Fls. mon- or dioecious, little reduced from the type of the fam. C. Tigliiiin L. (trop. As.) is the source of croton oil (a powerful purgative drug, expressed from the seeds). C. Cascarilla Benn. and C. Eluteria Benn. (Bahamas) yield Cascarilla bark, used as a tonic (cf. Cascarilla). C. laccifcrus L. (India, Ceylon) yields a lac, used in varnish-making, and several Brazilian spp. a dragon's blood resin. Croton (of trop. gardens), Codiaeum variegatum Bl. Croton-oil, Croton Tiglinni L. Crotonogyne Muell.-Arg. Euphorbiaceae (A. n. 2). to trop. Afr. Crotonogynopsis Pax. Euphorbiaceae (A. n. 2). i trop. Afr. Crotonopsis Michx. Euphorbiaceae (A. n. i). 2 N. Am. Crow-berry, Em pel nun nigrinn ' L. ; -foot, Ranunculus; -foot-grass, Dactyloctcn in in. Crowea Smith. Rutaceae (i). 4 Austr. Crown-bark, Cinchona ; -beard (Am.), Verbesina ; -Imperial, Frilil- laria imperialis L.; -palm (W.I.), Maximiliana. 1 88 CRUCIANELLA Crucianella L. Rubiaceae (n. n). 4 Eur., Medit. Cult. orn. fl. Cruciate, cross-shaped. Cruciferae (EP., BH.}. Dicots. (Archichl. Rhoeadales ; Panetales BH.}. 220 gen., 1900 sp., cosmop., but chiefly N. temp, and esp. Medit. ; a very natural fam., well marked off from others, though approaching Papaveraceae and Capparidaceae. Herbs, a few under: shrubs ; some ann., many perenn., forming each year a new shoot term, in the infl. L. usu. alt., exstip., with unicellular simple or branched hairs. For other peculiarities of veg. organs see gen., e.g. Brassica, Anastatica, Subularia, Vella, &c. Infl. usu. a raceme or corymb, and nearly always without bracts or bracteoles. Fl. usu. 5, reg., hypog., with typical formula K 2 + 2, €4, A 6, G (2). The K has two whorls, the C only one, alt. with the K as a whole. The petals usu. spread out in the form of a cross and are often clawed ; the sta. in two whorls, an outer of 2 short, an inner of 4 long, sta. (tetradynamous) ; anthers intr. The two cpls. are placed trans v., and have parietal plac., but the ov. is 2-loc. on account of the presence of an antero-post. partition, the repluin or so-called spurious septum, an outgrowth of the placentae. Stigmas 2, on short style, above the placentae (cf. Papaveraceae). Ovules anatr. Floral Diagram , J. (after Eichler). or campylotr. The explanation of the morphology of this fl. has given rise to much dispute. It is usu. regarded as a typically 2-merous fl. (cf. Papaveraceae), and the 4 petals and 4 inner sta. are supposed to be due to branching. Others regard it as 4-merous (e.g. Klein in Bot. Centr. 58, p. 197). Full discussion in Eichler's Bliithendia.gr. or in Asa Gray's Struct. Bot. p. 206. On the bases of the sta. are the nectaries, the honey being secreted into the often gibbous bases of the inner sepals. The sepals often stand almost straight up, and the petals are then provided with claws and spread out horiz. beyond the sepals. The honey is thus concealed to some extent and protected from rain. The majority of the order exhibit this construction more or less, thus coming into the biological fl. -class AB. In many gen. the fls. are arranged in corymbs, thus getting the advantage of many fls. massed together on one level (cf. Umbelliferae and Compositae). Insects visiting the fls. touch the anthers with one side of their bodies and the stigma with the other, and may in this way effect cross-fert., as they go sometimes to one, sometimes to the other, side of the fl. Dichogamy is frequent, but not well marked, and in almost all self-fert. ultimately occurs. fMiiller, Pert, of Fls.] Fr. a caps, of pod-like form ; if at least three times as long as broad it is called a siliqna, if shorter a silicula. It is divided into two by the replum and is usu. thin and membranous. The valves break away from below upwards, leaving the replum with the seeds pressed against it and adhering. The fr. may be flattened in two CRUPTNA 189 ways, either || or JL the replum ; this char, is of systematic importance. It may also be jointed between the seeds as in a lomentum (Legumi- nosae). Achene-like one-seeded fr. occur in a few gen. Others have subterranean fr. (Cardamine sp., &c.). The chars, of the seed are also of great importance in classification. The seed is exalb. : the testa is often mucilaginous, swelling up when wetted (e.g. the familiar case of mustard seed). The ovules being campylotr., the embryo sacs, and embryos, are curved, usu. with the radicle in one half of the seed, the cots, in the other. The shape of the embryo and the position of the radicle with regard to the cots, are important. The chief cases are: (i) radicle incumbent (or embryo notorhizal), i.e. lying on the back of one cot., the cots, not being folded on themselves; this may be shown thus o||, the o repres. the radicle; (2) decumbent (or embryo pleurorhizal}, o=, the radicle against the edges of the cots. ; (3) orthoplocous (cots, conduplicate), o> >; (4) spirolol'ous, as in (i) but cots, once folded, o||||; (5) diple- cotoboits, ditto twice or more folded, o||||||l|. For plants of economic value see esp. Brassica (which gives a number of valuable vegetables), Sinapis, Nasturtium, Lepidium, &c. All C. are harmless, and most are rich in sulphur compounds (to which the smell of boiling cabbages is due), and are thus useful in scurvy, &c. Classification and chief genera (after Frantl) : The grouping of the smaller divisions of the fam. and the defining of the genera is a most difficult task. Many classifications have been devised. Prantl (in Nat. Pfl.} bases his largely upon the hairs borne on the leaves. Others rely on characters of fruit and embryo, &c. In any case the identification of a cruciferous genus is a difficult matter. A. Hairs simple or none : no glandular hairs. 1. Thelypodieae (stigma equally developed all round; style un- divided or prolonged above middle of cpls., or turned back) : Pringlea, Thelypodium, Heliophila. 2. Sinapeae (stigma better developed over placentae) : Subu- laria, Lepidium, Iberis, Cochlearia, Alliaria, Sisymbrium, Cakile, Isatis, Vella, Sinapis, Brassica, Raphanus, Crambe, Nasturtium, Cardamine, Lunaria. B. Hairs branched (a few exceptions) : Sometimes also glandular hairs. 3. Schizopctaleae (stigma equal all round) : Schizopetalum, Physaria. 4. Hesperideae (stigma better developed over placentae) : Cap- sella, Draba, Arabis, Erysimum, Cheiranthus, Alyssum, Anastatica, Malcomia, Hesperis, Matthiola, Conringia. Cruckshankia Hook, et Am. Rubiaceae (i. 2). 5 Chili. Cruddasia Prain. Leguminosae (in. 10). i N. India. Crudia Schreb. (Apalaton Aubl.). Legumin. (n. 3). 20 trop. Cruentus (Lat.), blood coloured. Crula Nieuwland (Acer p.p.). Aceraceae. 7 E. As. Crumenaria Mart. Rhamnaceae. 5 trop. Brazil. Crunocallis Rydberg (Claytonia p.p.). Portul. i Arct. As. Am. Crupina Dill, ex L., Cass. Compositae (i i). 2 S. Eur. to Persia. 1 90 CRUSE A Crusea Cham, et Schlechtcl. Rubiaceae (n. 10). 10 Mex., C. Am. Crustaceous, hard and brittle. Cruzia Phil. Labiatae (vi). i Patagonia. Crymium (C1-), a polar barren formation. Cryosophila Blume (Copernicia BH.). Palmae (i. 2). i Mex. Cryphiacanthus Nees (Ruellia p.p.). Acanth. (iv. A). 10 S. Am. Crypsis Ait. Gramineae (8). i Medit. Cryptadenia Meissn. Thymelaeaceae. 5 Cape Col. Cult. orn. Cryptandra Sm. Rhamnaceae. 30 temp. Austr. Cryptanglum Schrad. Cyperaceae (n). 20 trop. Am. Cryptantha Lehm. (Eritrichium BH.). Borag. (iv. 2). 80 Pacif. Am. Cryptanthe Benth. et Hook. f. = preceding. Cryptanthopsis Ule. Bromeliaceae (4). 2 Brazil (Bahia). Cryptanthus Otto et Diet r. Bromeliaceae (4). 10 Brazil. Cult. orn. fol. Cryptarrhena R. Br. Orchidaceae (11. 19). 3 W.I., Guiana, Mex. Crypteronia Blume. Sonneratiaceae (Lythr. BH.). 4 Malaya. Cryptocarpus H. B. et K. Nyctaginaceae. 2 W. coast Am. Cryptocarya R. Br. Lauraceae (n). 60 trop. and subtrop. The fr. of C, moschata Nees et Mart. (Brazilian nutmegs) used as spice. Cryptocentrum Benth. (Mystacidium EP.). Orchid, (n. 20). i Ecuador. Cryptochilus Wall. Orchidaceae (n. 5). i Himalaya. Cryptochloris Benth. (Tetrapogon EP.). Gram in. (n). i Patagonia. Cryptocoryne Fisch. Araceae (vn). 20 Indomal. Marsh plants. Some sp. are apparently ' viviparous ' in their germination, like mangroves (see Goebel's Pflanzenbiol. Schild. I. p. 132). Cryptodiscus Schrenk. Umbelliferae (in. 4). 4 W. As. Cryptogamae. A term used to distinguish those pi. which do not produce seeds. All the higher C. exhibit alternation of generations (see Pteridophyta), and the distinction between them and Phanerogams depends on the fact that in C. the macrospore or spore falls out of its sporangium, germinates upon the ground or in water and gives rise to an independent ? (or 5 ) prothallus; in the P. on the other hand, it does not do so. The C. form 3 great groups, Thallophyta, Bryophyta, and Pteri- dophyta, each usu. regarded as equivalent to Spermaphyta. Cryptogamae Vascularea*= Pteridophyta. Cryptogramma R. Br. (Allosorus Bernh.). Polypodiaceae. 5 Eur., As., Am. C. crispa R. Br. (parsley-fern, curled rockbrake) Brit. Cryptogyne Hook. f. Sapotaceae (i). i Madag. Cryptolepis R. Br. Asclepiadaceae (i). 20 palaeotrop. Cryptomeria D. Don. Com'ferae (Pinaceae; see C. for gen. char.). i Japan, C. Japonica D. Don. (Japanese cedar), often cult. Timber good. Cryptophaseolus O. Ktze. (Canavalia p.p. EP.). Legum. (in. 10). i Annam. Cryptophoranthus Barb. Rodr. (Pleurothattis BH.). Orchid, (n. 8). 10 trop. Am., W.I. Cryptophragmium Nees (Gymnostachyum BH.). Acanth. (iv. B). 15 trop. As. Cryptopus Lindl. Orchidaceae (n. 20). i Madag. C UC URBITA CEAE 1 9 1 Cryptosepalum Benth. Leguminosae (n. 3). 8 trop. Afr. Cryptospora Kar. et Kir. Cruciferae (4) i W. As. Cryptostegia R. Br. Asclepiadaceae (i). 2 trop. Afr., Madag. Cryptostemma R. Br. (Arctotis p.p. EP.}. Comp. (10). 3 S. Afr. Cryptostephanus VVelw. Amaryllidaceae (i). 2 trop. Afr. Cryptostylis R. Br. Orchidaceae (n. 2). 7 Indomal. Cryptotaenia DC. Umbelliferae (in. 5). 3 Canada, Calabria, Came- roons (an almost impossible distribution; cf. Willis in Ann. Bot. 1916-7). Cryptotaeniopsis Dunn (Carum p.p. in part). Umb. (in. 5). 10 China. Ctenanthe Eichl. Marantaceae. 10 trop. Am. Ctenium Panz. Gramineae (n). 7 Am., Afr., Masc. Ctenodon Baill. (Aeschynomene p.p. EP.}. Legum. (in. 7). i Brazil. Ctenolophon Oliv. Linaceae (formerly Olac.). 2 Malay Peninsula. Ctenophryniurn K. Schum. Marantaceae. i Madag. Ctenophyllum Rydb. (Phaca p.p.). Leguminosae (in. 6). i N. Am. Ctenopsis De Notar. (Festiica p.p. EP.). Gramin. (10). i trop. Afr. Cuba bark (W.I.), bast, Hibiscus elatus Sw. ; -liemp (Aust.), Furcrea. Cubanthus Millspaugh (Pedilantkus p.p.). Euphorb. (A. n. 8). 2 Cuba. Cubeba Raf. = Piper L. (Piper.). Cubebs, Piper Cubeba L. Cubilia Blume. Sapindaceae (inc. sed.). 2 Phil. Is., Celebes. Cuckoo flower, Cardainine, &c. ; -pint, Arum maculatum L. Cucutaalus (Tourn.) L. Caryophyllaceae (n. i). i, C. bacdfer L., N. temp, (introd. in Brit.). Fr. a berry. Cucullaria Schreb. = Vochysia Juss. (Vochys.). Cucullate, hooded, cowled. Cucullus, Asclepiadaceae. Cucumber, Cuciimis sativus L.; calabash-, Lagenaria; squirting-, Ecballium; -tree (Am.), Magnolia. Cucumeropsis Naud. Cucurbitaceae (2). 2 trop. W. Afr. Cucumis (Tourn.) L. Cucurb. (3). 3 trop., subtrop. C. Melo L. (melon), C. sativus L. (cucumber) cult, from early times. Tendrils simple, regarded as of 1. nature (see fam.), the stem portion sup- pressed. Cucurbita (Tourn. ) L. Cucurbitaceae (3). 10 Am., but many so long cult, that their origin is doubtful. F~or tendrils see fam. Fls. monoecious. Germination interesting. On the lower side of the hypocotyl a peg is formed which holds one side of the testa firmly while the expansion of the plumule splits off the other side. The position of the peg is determined by gravity. C. Pepo L. is the pumpkin, with its vars. the vegetable marrow and squash; C. maxima Duchesne the giant pumpkin, cult, in N. Am Cucurbitaceae (EP., BH.}. Dicots. (Sympet. Cucurbitales ; Polypet. Passiflorales BH.}. 90 gen., 750 sp., wanting in the colder regions, most abundant in the trop., chiefly climbing ann. herbs with very rapid growth and abundance of sap in their steins and other tissues. L. alt. roundish, entire or lobed. They climb by tendrils, about whose morphological nature there has been much discussion ; they have been considered by various authors as "roots, stems, leaves, 192 CUC URBITA CEAE stipules, shoots, flower-stalks or organs sui generis." According to Muller (Nat. Pfl.} the tendrils of Cncurbita Pepo, with their frequent abnormalities, give a proof of their true nature. Every variety is found, from simple threads to long leafy tendrils, in which the 1. show all transitions to tendrils. Muller, therefore, considers the twining portion of the tendril to be a met. 1. , the lower stiff portion a stem. The tendrils of C. are very sensitive and show very well all the phenomena of tendril-climbing. Fls. diclinous, rarely 5 , in infls. of various types (see Nat. Pfl., or Eichler, Bluthendia.gr.). K and C typically (5) each, reg. ; A typically 5, but great var. is introduced by cohesions, &c.; it is almost always •[• . In Fevillea we find ~, sta. with biloc. anthers, the simplest type ; it is noteworthy that the usual 4-loc. anther never occurs in C., and no trace of the missing loc. is to be found in either lobe of the anther of Fevillea. In the rest of the fam. the A is more complex. In Thladiantha two pairs of sta. stand apart from the fifth sta. In Sicydium these pairs show union of their members at the base, and in others the union is more complete, until, as in Bryonia, &c., the A apparently has only 3 sta., of which 2 have 4-loc. anthers due to unions. The more the sta. depart from the simple type the more curved do the loculi of the anthers become, till in Cucurbita, &c. the pollen-sacs are twisted in a most extra- ordinary manner (cf. Columellia). In Cyclanthera the sta. are all united into a column with two ring-shaped pollen_chambers running round the top (cf. the flow ers of Cyclanthus). G i — io-loc. , with i — oo anatr. ov. in each loc. ; the most common type is, however, a 3-loc. ovary with axile plac. projecting deep into the cavity. Stigmas as many as cpls., usually forked. Fr. usu. fleshy, of the type of the melon or cucumber — berry-like, sometimes called a pepo. Seeds exalb. In Zanonia, Ecballium, Cyclanthera, &c. (q.v.), the mode of seed-dispersal is interesting. Several have ed. fr. , e.g. Cucurbita, Cucumis, Sechium, Lagenaria, &c. The relationships of this fam. have been much disputed. It has been placed near Passifloraceae, Loasaceae and Begoniaceae, but most probably it comes near to the Campanulaceae. Classification and chief genera (after Pax) : A. Pollen-sacs not fused into a ring. a. Sta. free or only united at base. i. Ftvilleeae (sta. 5, rarely 4) : Fevillea, Zanonia, Thladiantha. i. Melothrieae (sta. 3, rarely i or 4; pollen-sacs straight or slightly curved): Melothria, Telfairia. 3. Ciicurbiteae (do., but pollen-sacs S or U shaped): Acan- thosicyos, Momordica, Luffa, Bryonia, Ecballium, Cucumis, Lagenaria, Trichosanthes, Cucurbita. b. Sta. united into a column. 4. Sicyoideae: Echinocystis, Sechium, Sicyos. B. Pollen-sacs fused into a ring. 5. Cydanthereae: Cyclanthera (only genus). Cucurbitaceous, gourd-like. Cucurbitales (EP.). The pth order of Dicots. Sympetalae. Cucurbitella Walp. Cucurbitaceae (2). 4 temp. S. Am. CUPXESSUS 193 Cudrania Tree. Moraceae (n). 4 Japan to Austr., New Cal. Cudweed, Gnaphalium. Culcasia P. Beauv. Araceae (i). 15 trop. Afr. Culcitium Humb. et Bonpl. Compos. (8). 20 Andes. Like Espeletla. Cullenia Wight. Bombacaceae. i India, Ceylon. Cullumia R. Br. Compositae (10). 15 S. Afr. Cullumiopsis Drake del Castillo. Compositae (4). i Madag. Culm, the stem of a grass. Cultivation, see Economic and ornamental plants. Culver's physic, -root, Veronica virginica L. Cumbu, Pennisetiim tvphoidcum Rich. Cumin seed, Cuminiim Cyminuiii L. Cumingia Vidal. Bombacaceae. i Phil. Is. Cuminia Colla. Labiatae (vi). 3 Juan Fernandez. Cuminum (Tourn.) L. Umbelliferae (7). i C. Cyminum L., Medit. The fr. (cumin seeds) are sometimes used like caraway seeds. Cumminsia King ex Prain = Cathcartia Hk. f. (Papav.). Cuneal, cuneate, cuneiform, wedge-shaped. Cunila L. Labiatae (vi). 15 Am. Cunninghamia R. Br. Coniferae (Pinaceae; see C. for genus charac- ters). C. sinensis R. Br., the only sp., in S. China and Cochin- China. Cunonia L. Cunoniaceae. i, C. capensis L., S. Afr. It shows good bud-protection by stipules. Cunoniaceae (EP. ; Saxzfragactae p.p. BH.}. Dicot. (Archichl. Ro- sales). 21 gen., 120 sp., chiefly between 13° and 35° S- Shrubs and trees with opp. or whorled leathery 1., stip. (the slip, often united in pairs as in Rubiaceae). Fl. small, usu. $ . Receptacle usu. flat. K 4 — 5; C 4 — 5, usu. smaller than calyx, often absent ; A 8 — 10 or oo or 4 — 5; G usu. (2), rarely 2; ovary usu. 2-loc., generally with 00 — 2 ovules in 2 rows in each loc. Fruit usu. a capsule, rarely drupe or nut. Endosperm. Chief genera: Cunonia, Weinmannia. Cunuria Baill. Euphorbiaceae (A. II. 3). 2 N. Brazil. Cup, a hollow floral receptacle, Myrtaceae, Rosaceae, &c.; -flower, Nierembergia ; -plant (Am.), Silphium perfoliatum L. Cupania L. Sapindaceae (i). 32 warm Am. Wood of some is useful. C. sapida Voigt (C. edulis Schum. et Thonn.) = Blighia. Cupaniopsis Radlk. • Sapindaceae (i). 28 Austr., Polynesia. CupLea P. Br. Lythraceae. 200 Am. L. decussate; usu. there is one fl. at each node, standing between the two 1. ; this is really the axillary ft of the 1. below, and its peduncle is 'adnate' to the main stem. Many covered with sticky glandular hairs. Cult. orn. fl. Cupheanthus Seem. (Jambosa, Syzygium p.p. ? EP.). Myrtaceae (i). 1 New Caied. Cuphocarpus Decne. et Planch. Araliaceae (i). i Madag. Cupia DC. = Randia L. (Rubiac.). Cupirana Miers (Coupoui BH.). Apocynaceae (i. i). i Guiana. Cuprea bark, Reinijia. Cupreous, copper-coloured. Cupressaceae ( Warming) = Pinaceae § Cupresseae. Cupressus Tourn. ex L. [Synonymy: C. fastigiata DC. — C.semper- w. 13 i94 CUPRESSUS virens L. ; C. pendula Staunt. = C. fnnebris Endl.; C. nootkatensis Lamb. = Chamaecyparis nutkatensis Spach. ; C. thttjoides L = Ch. sphaeroidea Spach.; C. juniperoides L,. = Callitris arborea Schrad. ; C. japonica L. = Cryptomeria jap. Don. Coniferae (Pinaceae; see C. for gen. char.). 12 Medit., As., N.Am. The gen. habit is xero. , the 1. being much reduced and closely appressed to the stems. C. sempervirens L. is the cypress of the Medit. region; C.funebris Endl. the funereal cypress of China and Thibet, with ' weeping ' branches ; C. macrocarpa Hartn. (Monterey cypress, Calif.) is largely planted for timber and shade in warm countries. Several yield useful timber, e.g. C. Laivsoniana Murr. (Calif., Oregon), C. Lindleyi Klotzsch (Mexico), C. tornlosa Don (W.Himal.), C. sempervirens, &c. (see Camus, Les Cypres, Paris, 1914). Cupule, Fagaceae, Betulaceae. Cupuliferae (BH. ) = Betulaceae + Fagaceae ; (Warming) = Fagaceae. Curangajuss. Scrophulariaceae (n. 6). i Indomal. Curare, Strychnos toxifera Schomb. Curatella L. Dilleniaceae. 5 trop. Am., W. I. Curculigo Gaertn. Amaryllidaceae (in). 15 palaeotrop., S. Afr. Curcuma L. Zingiber. (i). 35 palaeotrop. C. angustifolia Roxb. tubers furnish east Indian arrowroot. C. longa L. yields the yellow dye turmeric (dried and ground rhiz. ). The tubers of C. Zedoaria Rose, yield zedoary, used in the East as a tonic and perfume. Curima O. F. Cook (Bactris EP.}. Palmae (iv. 2). 2 W. Indies. Curinila Roem. et Schult. Asclepiadaceae (inc. sed.). i Malaya. Curled rockbrake, Cryptogramma crispa R. Br. Curly greens, Brassica oleracea L. var. Curmeria Linden et Andre = Homalomena Schott p.p. (Arac.). Curraniodendron Merrill. Saxifragaceae (v). i Phil. Is. Currant, Kibes, Vitis ; Australian-, Leiicopogon ; black-, Ribes nigruin L. ; -bush (W.I.), Clidemia, Miconia, &c.; flowering-, Ribes sangni- neum Pursh; -tree (W.I.), Beureria, Jacquinia. Curroria Planch. Asclepiadaceae (i). i S. Afr. Curry-leaf (Ceyl., India), Murraya Koenigii Sprang. Curtia Cham, et Schlechtd. Gentian, (i). 10 Guiana to Uruguay. Curtisia Ait. Cornaceae. i S. Afr., C. faginea Ait., yielding a hard and useful timber (assegai-wood). Curvembryae. The ist series (BH.) of Monochlamydeae. The 7th cohort (Warming) of Choripetalae. Cuscuaria Schott (Scindapsiis p.p. EP.~). Araceae (n). i Malay Archipelago. Cuscus (khas-khas), Vetiveria zizanioides Stapf. Cuscuta (Tourn.) L. Convolvulaceae (n). 100 trop. and temp.; 3 Brit, (dodder, scald, &c.). Many have extended their boundaries through being carried about with their host plants. Leafless and rootless total parasites. The stem twines and is sensitive to contact like a tendril so that it clasps the support tightly ; it rarely makes more than three turns about the same branch of the host. At the points in close contact suckers are developed which penetrate the tissues of the host, growing into organic union with them and drawing off all the food CYATHEACEAE 195 materials required by the parasite, which has no green tissue of its own. The seeds of C. germinate later than those of the host plant ; a very short anchorage root is formed and the stem nutates in search of a host; as soon as it has clasped one the root dies away. Much damage is often done by these plants: most of the Brit. sp. confine themselves to particular host pi., but others attack a variety of pi. For details see Kerner's Nat. Hist, of Plants, vol. i., and papers by Peirce in Ann. of Bat. 1893 — 4. Cushion plants, with cushion-like growth reducing evap. Cusickia M. E. Jones. Umbelliferae (in. 6). i N.Am. Cusparia Humb. Rutaceae (i). 22 S. Am. C. febrifuga Humb. (C. trifoliata Eng.) yields Angostura or Cusparia bark, sometimes used in place of cinchona bark. Cuspidaria DC. Bignoniaceae (i). 5 Brazil, Bolivia. Cuspidate, with rigid point. Cussonia Thunb. Araliaceae (i). 25 S. and trop. Afr., Madag. Custard-apple, Anona sqtia»iosa L. , reticulata L. Cutandia Wilk. Gramineae (10). 6 Medit. Cutch, Acacia Catechu Willd., Rhizophora. Cuthbertia Small ( Tradescantia p-p-). Commelin. 2 N.Am. Cuticle, the thin detachable skin of a plant. Cutting grass (W.I.), Scleriaflagellum-nigrorum Berg. Cuttsia F. Muell. Saxifragaceae (v). i E.Austr. Cuviera DC. Rubiaceae (n. i). 10 trop. Afr. Several are ant- inhabited with hollow swellings of the stem above the nodes. Cyamopsis DC. Leguminosae (in. 6). 3 trop. Afr., As. C. psoraloides DC. is largely cultivated in India as fodder (guar). Cyanaeorchis Barb. Rodr. Orchidaceae (n. 7). i Brazil. Cyanandrium Stapf. Melastomaceae (i). 2 Borneo. Cyananthus Wall. Campanul. (i). 10 Mts. mid. and E. As. Ov. sup. Cyanastraceae (EP., Pontederiaceae p.p. BH. ). Monocots. (Farinosae). Only genus Cyanastrum, q.v. Cyanastrum Oliv. Cyanastraceae. 5 trop. Afr. Herbs with tuber or tuberous rhiz. , and racemes or panicles of $ reg. fls. P (3 + 3), A (6), G (3), 3-loc. with 2 ov. in each. Fr. i-seeded. Perisperm. Cyanea Gaudich. Campanulaceae (ill). 28 Sandw. Is. Cyanella L. Amaryllidaceae un.). (Haemodor. BH.} 7 S. Afr. Cyaneus (Lat.), full blue. Cyanocarpus Bailey. Proteaceae (11). 2 Queensland. Cyanodaphne Blume. Lauraceae (n). 2 Malay Archipelago. Cyanostegia Turcz. Verbenaceae (3). 4 W. Austr. Cyanothyrsus Harms. (Daniella p. p.), Legumin. (n. 3). 3 trop. Afr. Cyanotis D. Don. Commelinaceae. 35 palaeotrop. Cyanus (Toutn.) L. = Centaurea L. (Comp.). Cyathanthus Engl. Moraceae (i). i Cameroons. CyatheaSm. Cyatheaceae. 240 tiop. and subtrop. Tree ferns, forming a char, feature in the scenery of various regions. C. medullaris Sw. (N. Z.) and C. dcalbata Sw. are well known. Their pulpy pith is eaten by the natives. Cyatheaceae. Filicineae Leptosporangiatae (Homosporous). 9 gen. with 360 sp. chiefly trop. and subtrop., mostly tree ferns with stout 1 96 C YA THE A CEAE erect stems, covered with adv. roots and a palm-like crown of 1. at the top. These show circinate vernation, &c., very well. The sori are marginal or on the under side of the 1., naked or with a cup- shaped indusium ; the sporangia are shortly stalked and have a complete excentric annulus. Chief genera: Cyathea, Alsophila, Dicksonia, Hemitelia. CyatMum, an infl. reduced to look like a single fl., Euphorbia, Antho- sletna. Cyathocalyx Champ, ex Hook. f. et Thorns. Anon. (4). 9 Indomal. Cyathocephalum Nakai. Compositae (8). 2 Japan, Manchuria. Cyathochaeta Nees. Cyperaceae (n). 4 Austr. Cyathocline Cass. Compositae (3). 2 India. Cyathodes Labill. (Styphelia p.p. EP.}, Epacrid. 15 Austr. Polynes. Cyathogyne Muell.-Arg. Euphorbiaceae (A. i. i). 5 trop. Afr. Cyathopsis Brongn. et Gris. (Styphelia p.p. EP.). Epacridaceae (3). 2 Indomal. Cyathopus Stapf. Gramineae (8). i Indomal. Cyathoselinum Eenth. (Scseli p.p. EP.}. Umbellif. (in. 5). i Dal- matia. Cyatb-ostelma Fourn. Asclepiadaceae (n. i). 2 Brazil. Cyathostemma Griff. Anonaceae (i). 7 Malaya. Cyathula Lour. Amarantaceae (2). 10 Afr., As., S.Am. Cybele, a flora. Cybiantnus Mart. Myrsinaceae (ll). 35 trop. Am. Cybistax Mart. Bignoniaceae (2). 3 S. Am. The 1. of C. Spmcei K. Sch. are used as a blue dye, by boiling them with the cloth. Cycadaceae (chiefly after Eichler). Gymnospermae. 9 genera with about 75 sp., the survivors of a group of plants which in past ages figured more largely in the flora of the earth, reaching their maximum about the end of the Triassic and beginning of the Jurassic period. They represent the lowest type of living seed-plants and in appear- ance and habit are like tree-ferns. The stem is usu. short and stout, only growing to any noteworthy height in Cycas itself, and is often tuberously swollen; it shows a secondary growth in thickness. It has a long primary tap root. In some sp. a sort of felt-work of roots is formed at the base of the stem, and a number of short lat. branches of these stand erect and may emerge from the soil (see Nat. Pfl.}- The stem has usu. a crown of leaves, and its lower portion is covered with scales. There are, in all except a few sp. of Macrozamia, two sorts of 1., foliage- and scale-1., borne spirally upon the stem, and alt. with one another, as a rule several circles of scales before each circle of foliage 1., which they protect in the bud. The scales are really 1. bases whose blades abort. The foliage 1. are very char. They possess usu. a thickened, woody, ± sheathing base, which often persists after the fall of the rest of the 1. There is a stout rachis or petiole, frequently thorny at the base, the thorns being ' metamorphosed ' leaflets. Upon its upper side are two grooves, from which spring the leaflets, which may or may not be opp. to one another; there is usu. no term, leaflet. The leaflets may be entire or toothed and are usu. very rigid and leathery. Three types of nervature occur : CYCLADENIA 197 (1) midrib, no lateral nerves: Cycas. (2) midrib and lateral nerves : Stangeria. (3) numerous parallel or wavy, simple or forked nerves running longitudinally: the other genera. The fls. are dioec. and usu. take the form of cones ; these art: term., and so the stem becomes a sympodium, except in Cycas and some spp. of Encephalartos, &c., where the stem 'grows through' the fl or fls. The size of the cones varies considerably. Each consists essentially of a central axis bearing a number of fertile 1. or sporophylls; occasionally the lowest 1. are sterile as in Coniferae. In the ). Deflersia Schweinf. ex Penzig. Euphorb. (inc. fed.). i Erythrea. Deflexed, bent sharply outwards. Defoliation, leaf-casting. Degenerate fl., one which has gone back to an earlier type. Degenia Hayek. (Lesquerella p.p.). Crucif. (4). i Croatia. Deguelia Aubl. (Deiris Lour. BH.}. Legum. (in. 8). 40 trop. Deliaasia Blume. Lauraceae (n). 10 Malay Archipelago. Deherainia Decne. Theophrastaceae. 2 Mex., W. I., incl. D. smarag- dina Decne. with large green fls. (coloured by chlorophyll). Dehiscence, mode of opening; cf. Fruit, Stamen. Deianira Cham, et Schlecht. Gentianaceae (i). 5 Brazil. Deidamia Nor. ex Thou. Passifloraceae. 6 Madagascar. Deinacanthon Mez (Rhodostachys Phil.). Bromel. (4). i Argentina. Deinandra Greene = Hemizonia DC- (Comp.). Deinanthe Maxim. Saxifragaceae (in), i Japan. Deinbollia Schum. et Thonn. Sapindaceae (i). 15 warm Afr. , Madag. Dekindtia Gilg. Oleaceae. i trop. Afr. Dekinia Mart, et Gal. (Lepechinia p.p. E.P.). Labiat. (vi). i Mex. Delamerea Sp. Moore. Compositae (4). i Brit. E. Afr. Delaportea Thorel et Gagnep. Leguminosae (i. i). i Laos. Delarbrea Vieill. Araliaceae (2). 3 New Caled., New Guinea. Delavaya Franch. Sapindaceae (n). i Yunnan. Delima L. =Tetracera L. p.p. (Dillen.). Deliquescent (stem), breaking up into branches. DENDROPOGON 209 Delissea Gaudich. Campanulaceae (in). 7 Sandwich Islands. Delognaea Cogn. Cucurbitaceae (3). i Madagascar. Delopyrum Small (Polygonella p.p.)- Polygon. (11. 2). 2 U. S. Delostoma D. Don. Bignoniaceae (2). 5 trop. Andes. Delpechia Montr. Inc. sed. 2 New Caled. Delphinium Tourn. ex L. Ranunculaceae (2). 150 N. temp. D. Ajacis L. (larkspur) Brit. Several cult. orn. fl. Fls. -|- in racemes; the post, sepal is drawn out into a spur containing the spurs of the two post, petals, in which the honey is secreted. (Cf. with Aconitum, which is far more frequently robbed by humble-bees.) The fl. is protandr. with movement of sta., fert. by humble-bees. The open fl. projects horiz., but subsequently the stalk bends up and the follicles stand erect so that the seeds can only escape if shaken, e.g. by strong wind (censer-mechanism). Delphyodon K. Schum. Apocynaceae (n. i). i New Guinea. Delpinoa H. Ross (Agave p.p. EP. ). Amaryll. (n). i N. Am. Delpinoella Spegazz. Cruciferae (inc. sed.). i Patagonia. Delpinophytum Spegazz. Cruciferae (2). i Patagonia. Delpya Pierre ex Bonati (Vandellia p.p.). Scrophulariaceae (11. 6). i Cochin-China. Delpydora Pierre. Sapotaceae (i). i trop. Afr. Deltoid, the shape of an equilateral triangle. Dematophyllum Griseb. Zygophyllaceae(P). i Argentina. Demazeria Dum. Gramineae (10). 4 Medit., S. Afr. D.(Brizopyruni) sicitla Dum. is a cult, ornam. grass. Demersus (Lat.), sub-aqueous. Demeusia De Wild, et Durand. Amaryllidaceae (i). i Congo. Demidium DC. (Aniphidoxa EP.}. Compositae (4). i Madagascar. Democrita Veil. Inc. sed. i Brazil. Dendrobangia Rusby. Icacinaceae. i Bolivia. Dendrobium S\v. Orchidaceae (n. 15). 600 trop. As., Japan, Austr., Polynes. Epiphytes; cult. orn. fl. For floral mechanism see Dar- win's Orchids, p. 138. Dendrocalamus Nees. Gramineae (13). 12 Indomal., China. D. giganteus Munro (the giant bamboo), the largest known bamboo (cf. bamboos), grovvs with great rapidity (see Lock in Ann. Perad., II. 1904, p. 21 1), even as much as 46 cm. a day. D. strictus Nees (male bamboo) has solid stems, used for lances, &c. DendrocMlum Blume. Orchidaceae (n. 16). 3 Indomal. Dendrocolla Blume = Sarcochilus R. Br. p.p. (Orchid.). Dendroconche Copeland (Polypodium p.p.). Polypod. i Phil. Is. Dendrocousinia Millspaugh. Euphorbiaceae (A. n. 7). 2 N. Am. Dendroid, tree-like. Dendromecon Henth. Papaveraceae (n). ro California. Dendron (Gr.), a tree. Dendropanax Decne. et Planch. (Gilibertia Ruiz et Pav. EP.}. Aralia- ceae (i). 25 trop. and subtrop. Dendrophthoe Mart. = Loranthus L. p.p. (Loranth.). Dendrophthora Eichl. Loranthaceae (n). 20 W.I., trop. Am. Dendrophylax Reichb. f. Orchidaceae (n. 20). 3 W. Indies. Dendropogon Rafin.^Tillandsia L. (Bromel.). W. 14 2 IO DENDROSERIS Dendroseris D. Don. Compositae (13). 7 Juan Fernandez. Dendrosicyos Balf. f. Cucurbitaceae (•2). 2 Socotra, trop. Afr. Dendrosma Panch. et Sebert. Rutaceae (inc. sed.). i New Caled. Dendrostylis Karst. et Triana (iMayna EP.). Flac. (2). 8 S. Am. Denekia Thunb. Compositae (4). 2 trop. Afr. Denhamia Meissn. Celastraceae. 4 trop. Austr. Denisia Post et O- Ktze. (Phryma p.p.). Phrymaceae. i S. Afr. Denisonia F. Muell. Verbenaceae (3). i Australia. Denizen, a pi. probably foreign, but maintaining its place. Dennettia E. G. Baker. Anonaceae (i). i S. Nigeria. Dennstaedtia Bernh. Polypodiaceae. 60 trop., S. Am., Austr. Dens (Lat.), a tooth. Dentaria (Tourn.) L. (Cardamine p.p. BH.). Crucif. (2). 20 N. temp. Dentate (dim. denticulate), with small teeth pointing outwards. Dentella Forst. Rubiaceae (i. 2). i Indomal. Deodar, Cedrus Deodnra Loud. Deonia Pierre ex Pax. (Blachia EP.). Euphorb. (A. n. 5). i Cochin- China. Depauperate, diminutive. Dependent, hanging down. DeplancheaVieill. \Diplanthera BH.). Bignon. (2). 6 Malaya, Austr. Deppea Cham, et Sclilechtd. Rubiaceae (I. 3). 9 C. Am., Mex. Depresmenilia F. Muell. (Pityrodia p.p. EP.). Verben. (3). i Austr. Dermatobotrys Bolus. Scrophul. (inc. sed.). i Zululand. Dermatocalyx Oerst. Scrophulariaceae (n. 4). i Costa Rica. Deroemeria Reichb. f. (Habenaria p.p. BH.). Orchidaceae (n. i). 4 trop. Afr., Abyssinia. Derris Lour. (Deguelia Aubl.). Leguminosae (in. 8). 50 trop. Desbordesia Pierre ex Van Tiegh. {Irvingia p.p.). Simarubaceae. 5 trop. Afr. Descending (aestivation), see Aestivation. Deschampsia Beauv. Gramineae (9). 20 temp, and frigid. D. caespi- tosa Beauv. (Aira) and D. flexuosa Trin. in Brit, (hair grass); of tufted growth ; rough fodder grasses. Description of plants. Root, stem, leaf, flower, and fruit, &c. are described for floras, &c. in concise technical terms (original descrip- tions of new species must be in Latin), which are mostly given here under fl., 1., &c. The descriptions of families in this book may serve as examples for large groups, and as examples of a species described in full detail we may refer to any numbers of \\\ejourn. Linn. Sot. or to Lind ley's Descriptive Botany, from which we quote as instances: lilac (1.): 1. opp., exstip., roundish-cordate, very acute, thin, smooth, rather longer than the linear channelled petiole, buttercup (fl.) : fl. term., sol., on long angular and furrowed peduncles, reg., S, hypog. Sepals 5, polysep., oval, coloured at edge, reflexed, with shaggy hairs. Petals 5, polypet. , round- ish, concave, with wedge-shaped basal nectaries, bright yellow. Stamens co , polyandrous, spiral ; filament yellow, slender ; anther linear, adnate, extrorse. Carpels oo , apocarpous, superior, collected into a nearly spherical head, greenish ; stigmas sessile, recurved ; ovules solitary, ascending, anatropous. DEWEYA 211 But if all the species in a genus are known, it is obvious that for purposes of description to enable identification, such a description is much too long, and the skill of the describer will be shown in describing those characters only in full (or as fully as necessary) which are essential to the discrimination of the species among its congeners. Descurainia Webb et Berth. (Sisymbrium p.p. BH.) Cruciferae (4). 1 8 N. temp., S. Am. Desdemona Sp. Moore. Scrophulariaceae (inc. seel.), i Brazil. Desfontainia Ruiz et Pav. Loganiaceae. 2 Andes. Ovary 5-loc. Desmanthodium Benth. Compositae (5). 4 Mexico, C. Am. Desmanthus VVilld. Leguminosae (i. 3). 15 Am., Madagascar. Desmiograstis Borner. Cyperaceae (in), i N. temp. Desmochaeta DC. = Pupalia Juss. (Amarant.). Desraodium Desv. Leguminosae (in. 7). 170 trop. and subtrop. In D. gyrans DC. (telegraph plant), during the day, if the temperature be not below 72° F. the two small lat. leaflets of each 1. move steadily round in elliptical orbits. See Darwin's Movements of Plants. At night the leaves sleep, drooping downwards. Several are useful as fodder, and are cult. Desmogyne King et Prain. Ericaceae (in. r). i India. Desmoncus Mart. Palmae (iv. 2). 25 trop. Am., climbing palms with reedy stems, and hooks like Calamus. Desmonema Miers. Menispermaceae. 6 warm Afr. Desmoscelis Naud. Melastomaceae (i). 2 trop. S. Am. Desmostachya Stapf. Gramineae (10). r trop. As. and Afr. Desmostachys Planch. Icacinaceae. 3 Madagascar, trop. Afr. Desmothamnus Small (Andromeda p.p.). Eric. (n. i). i Florida. Desmotrichum Blume (Dendrobium p.p. ). Orch. (11. 15). 27 Malaya. Despeleza Nieuwland (Lespedcza p.p.). Legum. (in. 7). 4 U.S. Desplatzia Bocq. Tiliaceae. i W. trop. Afr. Desvauxia R. Br. — Centrolepis Labill. (Centrolep.). Detandra Miers (Sychnoscpalum Eichl. ). Menisp. 4 trop. S. Am. Detarium J uss. Legumin. (n. 2). 4 trop. Air. Pith of pod ed. Determinate, derinite, ending with a bud. Dethawia Endl. (Seseli p.p. BH.}. Umbellif. (in. 5). i Pyrenees. Detris Adans. = Felicia Cass. (Comp.). Deuterocohnia Mez. (Dyckia). Bromeliaceae (2). 3 S. Am. Deutzia Thunb. Saxifragaceae (ill). 20 N. temp, and trop. Ovary inf. 3 — 4-loc. The fruit splits septicidally into its cpls. which open each at its apex. The seed is provided with a winged testa, very light. Deverra DC. = Pituranthos Viv. (Umbell. ). Devil-in-a-bush, Nigdla; -'s bean (W.I.), Capparis jamaicensis Jacq. ; -'s bit scabious, Scabiosa Succisa L. ; -'s cotton, Abroma augusta L. f. Devillea Tul. et Wedd. Podostemaceae. i Brazil. Dewberry, Riilnts caesins L. Dewevrea M. Micheli. Leguminosae (in. 6). i trop. Afr. Dewevrella De Wild. Apocynaceae (n. i). i trop. Afr. Deweya Torr. et A. Gray (Arracacia BH.}. Umbell. (in. 4). 4 W. N. Am. 14—2 2 1 2 DE WILDEMANIA Dewildemania O. Hoffm. Compositae (i). i Congo. Dewindtia I)e Wild. Leguminosae (ll. 3). i trop. Afr. Dextrorse, to the right. Deyeuxia Clar. (Calania^rostii, 'p.p. EP.}. Gramineae (8). 120 temp. Dhak tree, Buteafrondosa Roxb. Dhal, pigeon pea, Cajaniis indicus Spreng. Di- (Gr. pref.), two; -adelpnous, in two groups; -androus, with two sta. ; -carpellary, with two cpls. ; -chlamydeous, with distinct K and C ; -chogamy, see article below ; -chotomy, actual forking of growing apex; -clinism (-clinous), with separate 3 and ? fl.; -coty- ledons, one of the great divisions of angiosperms ; -cyclic, in two whorls ; -dymous, twinned ; -dynamous, with two sta. longer than rest, Labiatae ; -merous, with two members in each whorl ; -midiate, halved ; -morphism, see article below ; -oecism, see article below ; -photic, with two surfaces unequally lighted ; -plecolobous, Cruci- ferae; -plocnlamydeous, see dichlamydeous ; -plostemonous, sta. in two whorls, outer alt. with C; -ptera, flies, &c. ; -Sticbous, in two ranks; -thecous, with two thecae; -varicate, very divergent ; -vergens (Lat.), separating. Dia- (Gr. pref.), transverse; -heliotropism, transv. h., e.g. in runners, &c. ; -phragm, a dividing membrane ; -tropism, a placing of organs transv. to a stimulus. Diacalpe Blume. Polypodiaceae. i trop. As. Diacarpa Sim. Sapindaceae. i E. Afr. Diacattleya x , Dialaelia x Hort. Orchidaceae. Hybrids of Diacrium with Cattleya and Laelia. Diacidia Griseb. Malpighiaceae (11). i trop. S. Am. Diacrium Benth. Orchidaceae (n. 6). 4 Mexico to Guiana. Diadenium Poepp. et Endl. Orchidaceae (n. 19). i Peru. Diagram, floral, see Floral Diagram. Dialiopsis Radlk. Sapindaceae (n). i trop. Afr. Dialium L. Leguminosae (11. 5). 15 trop. Petals 2, i, or o; sta. 2, or rarely 3. D. guineense Willd. (trop. Afr.; velvet tamarind) pod contains an ed. pulp ; wood useful, resists salt water. D. indum L. (Java ; tamarind plum) and others have also ed. fr. Dialyanthera Warb. Myristicaceae. 2 Peru, Colombia. Dialycarpa Mast. Bombacaceae. i Borneo. Dialyopsis Radlk. Sapindaceae (a), i trop. Afr. Dialypetalae, Polypetalae. Dialypetalum Benth. Campanulaceae (ill). 2 Madagascar. Diamorpha Nutt. Crassulaceae. 2 E. U.S. Diandriella Engl. Araceae (v). i New Guinea. Diandrolyra Stapf. Gramineae (5). j, habitat unknown. Dianella Lain. Liliaceae (in). u trop. As., Austr., Polynes., Masc. Dianthera Gronov. (Justicia p.p. EP.). Acanthaceae (iv. B). 80 trop. Dianthoseris Sch. Bip. Compositae (13). 2 Abyssinia. Diantbus L. Caryophyllaceae (n. 2). 250 Eur., As., Afr., esp. Medit., mostly in dry sunny situations (4 in Brit.; pinks). Genus readily known by the bracts under the K. Fls. very protandrous (class F), largely visited by butterflies. Many cult. orn. fl., e.g. D. barbatus L. DICER OSTYLIS 2 1 3 (Sweet William),/?. Caryophyllus\^,. (carnation, picotee, clove-pink), D. chinensis (China or Indian pink), &c. Diapedium Koen. = Dicliptera Juss. (Acanth.). Diapensia L. Diapensiaceae. 2, one Himal., D. lapponica L. circum- polar boreal. Tufted, like oo alpine and arctic pi.; fl. protog. Diapensiaceae (EP., BH.}. Dicots. (Sympet. Ericales EP., BH.). 6 gen., 9 sp., "^ , chiefly alpine and arctic evergr. under-shrubs, with rosettes of 1. ; fls. sol. or in racemes, with two bracteoles, £, actinom., without a disc. K (5) or 5, C (•;) nearly polypet., A 5, epipet., opp. sepals, with frequently 5 stds. opp. petals ; anthers transv., each lobe opening by longitudinal slit ; pollen simple ; G (3) with axile plac. bearing oo anatr. or amphitr. ov. ; style simple with 3-lobed capitate stigma. Fruit a loculic. caps. Embryo cylindrical, endosp. fleshy. Chief genera : Diapensia, Shortia, Galax. Diaperia Nutt. (Evax p.p.). Compositae (4). 4 N. Am. Diaphanantlie Schlechter. Orchid, (n. 20). 20 trop. Afr. Diapliycarpus Calest. (Canmi p.p.). Umbell. (in. 5). i Medit. Diarrhena Beauv. Gramineae uo). 3 N. Am., E. As. Diarthron Turcx. Thymelaeaceae. i C. As. Diascia Link et Otto. Scrophulariaceae (n. i). 30 S. Afr. Diaspasis R. Br. Goodeniaceae. i S.W. Austr. Diasperus L. = Phyllanthus L. (Euph.). Diaspis Niedenzu. Malpighiaceae (i). i Brit. E. Afr. Diastatea Scheidw. Campanulaceae (inc. sed.). i Mexico. Diastema Benth. Gesneriaceae (n). 18 trop. Am. Cult. orn. fl. Diateinacanthus Lindau. Acanthaceae (iv. B). i Centr. Am. Diatenopteryx Radlk. Sapindaceae (i). i S. Am. Diberara Baill. (Nebelia BH.}. Bruniaceae. 5 S. Afr. Dicaelospermum C. B. Clarke. Cucurbitaceae (2). i Indomal. Dicarpidium F. Muell. Bombacaceae. i Austr. Dicaryum Willd. (Geissanthus EP.}. Myrsinaceae (n). 2 S. Am. Dicella Griseb. Malpighiaceae (n). 3 Brazil, Paraguay. Dicellandra Hook. f. Melastomaceae (i). 2 trop. W. Afr. Dicellostyles Benth. Malvaceae (4). 2 Ceylon, Sikkim. Dicentra Bernh. Papaveraceae (in), 15 As., N. Am. D. Cucullaria Bernh. (Dutchman's breeches) and others cult. orn. fl. The rhiz. of many sp. (§ Cucullaria} resembles a succession of bulbs, on account of the fleshiness of the scale 1. and of the sheathing bases of the fol. 1. The materials formed in the 1. during the growing season are stored up in the fleshy base, which survives the winter, while the rest of the 1. dies. Fls. in racemes, pend. Each outer petal has a large pouch at its base. The inner petals are spoon-shaped and cohere at the tip, forming a hood which covers the anthers and stigma. The pend. position and complex structure of the fl. render it suited to bees, which hang on to it and probe for honey first one side, then the other, in the pouches of the petals. In so doing they push aside the hood and touch the stigma, on which there is usu. pollen from its own sta. Dicerandra Benth. = Ceranthera Ell. (Labiatae). Diceratella Boiss. Cruciferae (4). 3 W. As., trop. Afr. Dicerostylis Blume (Hylophila EP.}. Orchidaceae (n. 2). t Malaya. 214 DICHAEA Dichaea Lindl. Orchidaceae (n. 20). 10 trop. Am., W. I. Monopodial. creeping epiphytes with sheathing 1. Cult. orn. fl. Dichaelia^Harv. (Brachystelma BH.}. Asclep. (II. 3). 10 S. Afr. Dichaeopsis Pfitz. (Dichaea Lindl. p.p.). Orchidaceae (n. 20). 5 trop. Am. Dichaetanthera Endl. Melastomaceae (i). 16 Madagascar, Masc. Dichaetaria Nees (Gymnopogon Beauv.). Gramin. (n). i Indomal. Dichaetophora A. Gray (Boltonia p.p. BH.). Comp. (3). i Texas. Dichapetalaceae (EP.; Chailletiaceae BH.). Dicots. (Archichl. Gera- niales EP., BH.}. 3 gen., 100 sp. trop. Woody pi. with entire stip. 1. Fls. in cymose umbels, &c., sometimes epiphyllous, usu. reg., 5 or unisex., typically =,-merous. K and C free or united, the petals often bifid ; axis continued into a cup-like disc or scales ; A 5, sometimes epipet. ; 6(2—3), each with 2 ov" DruPe with '" or 2-loc. stone ; no endosp. ; sometimes a caruncle. Chief genus Dichapetalum. Dichapetalum Thou. (Chailletia DC.). Dichapet. 120 trop. Several have epiphyllous infl. (cf. Erythrochiton), probably arising by a de- velopment like infl. of Solanaceae, or thorns of Cactaceae. Dichasial cyme, one in which each successive branch bears two branches upon itself, Acanthaceae, Asclepiadaceae, Begonia, Bignoniaceae, Carvophyliaceae, Castanea, Convolvulaceae, Cornaceae, Labiatae, Linaceae, Sanicula, Saxifragaceae, Ulmus, Urlica, Verbenaceae. Dichazothece Lindau. Acanthaceae (iv. B). i Rio de Janeiro. Dichelacrine Endl. Gramineae (8). 3 Austr., N. Zealand. Dichelostemma Kunth. (Brodiea Sm.). Lili. (iv). 8 N. Am. Dicherantlius Webb. Caryophyllaceae (i. 6). i Canaries. DicMlanthe Thw. Rubiaceae (n. 2). 2 trop. As. Dichiloboea Stapf. Gesneriaceae (i). i E. trop. As. Dichilus DC. Leguminosae (ill. 3). 4 S. Afr. Dichoespermum Wight = Aneilema R. Br. p.p. (Commelm.). Dichogamy, ripening of sexes at different times ; sta. ripe before stigma, protandry, Aeschynanthus, Aconitum, Aquilegia, Bignonia, Borago, Campanula, Campanulaceae, Caryophyllaceae, Clerodendron, Com- positae, Crassulaceae, Crocus, Delphinium, . Dipsacaceae, Echium, Empetrum, Epilobium, Geraniaceae, Gesneriaceae, Gladiolus, La- biatae, Malva, Monarda, Oxalis, Phacelia, Phyteuma, Rosaceae, Saxifraga, Scabiosa, Stellaria, Teucrium, Thymus, Umbelliferae, Valeriana ; stigma ripe before sta., protogyiiy, Alopecurus, Amorpha, Anthurium, Aesculus, Araceae, Callitriche, Carex, Chimonanthus, Colchicum, Coriaria, Epimedium, Euphrasia, Ficus, Fragaria, Helle- borus, Humulus, Juncus, Magnolia, Mirabilis, Parietana, Pans, Plantago, Potamogeton. Pyrus, Scrophularia, Thalictrum. Dichoglottis Fisch. et Mey. =Gypsophila L. p.p. (Caryophyll.). Dichondra Forst. Convolvulaceae (i). 5 trop., some amphicarpic. Dichondropsis T. S. Brandegee. Convolvulaceae (i). i Mexico. Dichopog-on Kunth. Liliaceae (ill), i Austr. Cult. orn. Dichopsis Thw. (Palaquinm Blanco i/.v.). Sapotaceae (i). 50 Indomal. Dichorisandra Mikan. Commelinaceae. 30 trop. Am. Infl. racemose (cf. fam.); its branches often pierce the leaf-sheath. Dichosciadium Domin. (Azorella p.p.). Umbell. (I. 2). i Austr. DICTYO CHL OA 215 Dichostemma Pierre. Euphorbiaceae (A. n. 8). i trop. Afr. Dichotomantlies S. Kurz. Rosaceae (v). r Yunnan. Dichroa Lour. Saxifragaceae (in), i Himal. and China to Java. Dichrocephala L'Herit. ex DC. Compositae (3). 8 warm Afr., As. Dichrolepis Welw. (Eriocattlon p.p. EP.). Eriocaul. i trop. Afr. Dichromena Michx. Cyperaceae (i). 12 Am. Dickrostachys Wight et Am. Leguminosae (i. 4). 10 palaeotrop. Stipules often thorny. Upper fls. of infl. £ , yellow, lower neuter and white, rose, or purple. Dichrotrichum Reinw. Gesneriaceae (i). 10 Malay Archip. Dickinsia Franch. Umbelliferae (i. i). i China. Dicksonia L'Herit. (excl. Cibotiitm Kaulf.). Cyatheaceae. 25 trop. and subtrop. and S. Hemisph. Most are tree ferns, e.g. D. antarctica Labill. (Austr., N. Z.). Sori marginal. For the Tartarian lamb cf. Cibotium. Dicladanthera F. Muell. Acanthaceae (iv. B). i W. Austr. Diclidanthera Mart. Styracaceae. 2 Brazil. Diclidium Schrad. ex Nees=Mariscus Gaertn. (,#//. ) = Cyperus L. p.p. Diclidocarpus A. Gray ( Trichospermum BH.}. Tili. 2 Java, Fiji. Dicliptera Juss. Acanthaceae (iv. B). 70 trop. Diclis Benth. Scrophulariaceae (n. 3). 5 S- Afr., Madag. Diclytra Borckh. = Dicentra Bernh. (Papav. ). Dicoelia Benth. Euphorbiaceae (A. I. i). i Borneo. Dicoma Cass. Compositae (12). 30 Afr., Madag., trop. As. Dicoria Torr. et A. Gray. Compositae (5). 4 W. U.S. Dicorynia Benth. Leguminosae (n. 5). 2 Guiana, N. Brazil. Dicoryphe Thou. Hamamelidaceae. 13 Madag., Comoro Is. Dicotyledones. One of the two great divisions of Angiospermae. Dicraea (Du Pet. Th.) Tul. Podosternaceae. 7 Madag., Ceylon, India. Thallus (root) drifting from attached base, exogenously branched, with marginal secondary shoots. Fr. isolobous. Dicraeanthus Engl. Podostemaceae. i trop. Afr. Dicraeopetalum Harms. Leguminosae (in. i). i Somaliland. Dicranocarpus A. Gray. Compositae (5). i Texas, Mexico. Dicranolepis Planch. Thymelaeaceae. 15 trop. Afr. Dicranopteris Bernhardi = Gleichenia Sm. (Poly pod.). Dicranostyles Benth. Convolvulaceae (i). 2 trop. S. Am. Dicranotaenia Finet. Orchidaceae (n. 20). i Dahomey. Dicrastylis Drumm. Verbenaceae (3). 8 N. and W. Austr. Dicraurus Hook. f. Amarantaceae (3). i Texas, Mexico. Dicrypta Lindl. = Maxillaria Ruiz et Pav. (Orchid ). Dictamnus L. Rutaceae (i). i Eur. As., D. a/bus L. (D- Fraxinella Pers.), dittany or candle-plant. The ethereal oil secreted by the pi. is volatile and inflammable, so that on hot calm days the air round it may sometimes be ignited. Fl. •[• ; the unripe sta. are curved down, and bend up to dehisce. Fr. opens elastically. Dictyandra Welw. Rubiaceae (i. 8). 2 W. trop. Afr. Dictyanthus Decne. in DC. Asclepiadaceae (n. 4). 4 Mexico. Dictyocaryum H. Wendl. (Iriartea p.p. EP}. Palmae (iv. i). i trop. Am. Dictyochloa Camus (Ammochloa p.p.). Gramineae (10). i N. Afr. 216 DICTYOLOMA Dictyoloma A. Juss. Rutaceae ('li). 2 Brazil, Peru. Dictyoneura Blume (Cupania p.p. BH.). Sapind. (i). 3 Malay Arch. Dictyophleba Pierre (Landolphia p.p.). Apocyn. (i. i). T trop Afr. Dictyosperma Regel (Pirea EP.). Cruciferae (2). i Turkestan. Dictyosperma Wendl. et Drude. Palmae (iv. i). 3 Mascarenes. Dictyostega Miers. Burmanniaceae. 5 trop. Am. and Afr. Dictyoxiphium Hook. Polypodiaceae. i Cent. Am. Dicyclophora Boiss. (Pycnocycla BH.). Umbell. (in. i). i Persia. Dicymbe Spruce ex Benth. et Hook. f. Leguminosae (n. 8). i Braz. Dicypellium Nees. Lauraceae (i). i Brazil, D. caryophyllatum (Mart.) Nees. Wood valuable; bark (Cassia caryophyllata) smells like cloves. Dicyrta Regel (Achimenes p.p. EP.). Gesner. (n). -2 Guatemala. Didactyle Lindl. =Bulbophyllum Thou. (Orchid.)- Didelotia Baill. Leguminosae (li. 3). 4 trop. Afr. Didelta L'Herit. Compositae (10). 3 S.W. Afr. Didesmandra Stapf. Dilleniaceae. i Borneo. Didesmus Desv. = Rapistrum Tourn. p.p. (Crucif.). Didiciea King et Pantl. Orchidaceae (n. 6). i Sikkim. Didieraceae, a fam. sometimes created for Didierea. Didierea Baill. Sapindaceae (?). 4 Madag. Anomalous plants with the habit of cactus-like Euphorbias, and of doubtful affinity. See Nat. Pfi. m 5, p. 461, Kew Bull. 1898, p. 97. Didiplis Rafin. (Peplis p.p. EP.). Lythraceae. i N. Am. Didiscus DC. (Trachymene BH.). Umbell. (i. i). 10 Malaya, Austr. Didissandra C. B. Clarke. Gesneriaceae (i). 30 India, China. Didymaea Hook. f. Rubiaceae (n. n). i Mexico. Didymanthus Endl. Chenopodiaceae (A), i W. Aust. Didymeles Thou. Inc. sed. i Madagascar. Didymia Phil. (Mariscus p.p. BH.). Cyperaceae (n). i Chili. Didymocarpaceae = Gesneriaceae. Didymocarpus Wall. (Rottlera \z\\\). Gesneriaceae (i). 120 Indomal., China, Madag., Austr., trop. Afr. Didymochlaena Desv. Polypodiaceae. i trop. Didymochlamys Hook. f. Rubiaceae (1.7). i Colombia. Epiphytic. Didymopanax Decne. et Planch. Araliaceae (i). 24 trop. Am. Didymopelta Regel et Schmalh. (Astragalus p.p. BH.). Leguminosae (ill. 6). i Turkestan. Didymophysa Boiss. Cruciferae (2). i Persia. Didymoplexis Griff. (Leucorchis p.p. EP.). Orch. (n. 2). 10 Indomal. Didymosperma H. Wendl. et Drude. Palmae (iv. i). 8 Indomal. Didymotheca Hook. f. Phytolaccaceae. 4 Austr., Tasmania. Dieffenbachia Schott. Aracea (v). 12 trop. Am. Fls. monoec. , naked, the $ is a synandrium of 4 or 5 sta. D. Scgiiine Scott is the ' dumb cane' of the \V. Ind. , formerly used in torturing slaves; it renders speechless a person who chews a piece of the stem. Diellia Brackenridge. Polypodiaceae. 8 Hawaiian Is. Dielsia Gilg. Restionaceae. i Austr. Dielsina O. Ktze. (Polyceratocarpus Engl. et Diels). Anonaceae (4). i trop. Afr. Dielytra Cham, et Schlecht. = Dicentra Bernh. (Papav.). DIMORPHA NDRA 2 1 7 Dienia Lindl. = Microstylis Nutt. (Orchid.). Dierama C. Koch. Iridaceae (in). 4 trop. and S. Afr. DierviUa Tourn. ex L. ( Weigelia Thunh.). Caprifoliaceae. 8 E. As., N. Am. D. florida Sieb. et Succ. , and others, cult. orn. shrubs. Fl. adapted to bees; changes colour after fert. (? only the effect of age) ; cf. Ribes, Fumaria, &c. Dietes Salisb. = Moraea L. p.p. (Irid.). Dieudonnaea Cogn. Cucurbitaceae (3). i Peru. Diffuse, loosely spreading. Digera Forsk. Amarantaceae (2). i trop. As., Afr. Digitalin, Digitalis. Digitalis (Tourn.) L. Scrophulariaceae (in. i). 22 Eur., W. As., Canary Is. D. purpurea L. (foxglove), Brit. Fls. in racemes, one- sided by twisting of peduncles ; adapted to fert. by bees. The 1. are offic., containing the poisonous alkaloid digitalin. Digitaria Heist, ex Adans. = Panicum L. p.p. (Gram.). Digitate (1.), palmate with 5 (or 7) leaflets. Diglyphosa Blume (Ckrysoglossum BH.}. Orchid, (n. a. n). i Java. Dignathe Lindl. Orchidaceae (ll. 19). i Mexico. Dignathia Stapf. Gramineae (3). 2 E. trop. Afr. DigompMa Benth. Bignoniaceae (2). i Guiana. DigrapMs Trin. = Phalaris L. p.p. (Gramm.). Digyroloma Turcz. Acanthaceae (inc. sed.). i Madras. Diholcos K yd berg (Astragahis p.p.). Legumin. (ill. 6). 4 N. Am. Dilatris Berg. Haemodoraceae. 2 S. Afr. Dildo (W.I.), Cereus Swartzii Griseb. Dilkea Mast. Passifloraceae. 5 N. Brazil. Dill, Peucedanitm graveolens Benth. et Hook. f. Dillenia L. (incl. Wormia Rottb.). Dilleniaceae. 25 sp. Indomal. Dilleniaceae (EP. ; BH. incl. Crossosomataceae}. Dicots. (Archichl. Parietales ; Ranales BH.}. 12 gen., 320 sp., trop., esp. well repres. in the Austr. ' scrub' veg. Most are trees and shrubs (many lianes) with alt. usu. leathery 1. and cymose infl. Fl. usu. reg., ?. K 5, or 3, 4 or even oo , spiral, persistent after flowering ; C usu. 5 ; A oo , hypog., free or united at base; G. oo — i, free or ± united; styles usu. free; ov. oo — i, erect, anatr., with ventral raphe. Plac. usu. inconspic., unthickened. Fr. dehisc. or not. Seed always with funi- cular aril united to the testa. Endosp. copious; embryo small, straight. Chief genera: Dillenia, Hibbertia. Dillwynia Sm. in Kon. et Sims. Leguminosae (in. 2). 10 Austr. Dilobeia Thou. Pioteaceae (i). i Madagascar. Dilodendron Radlk. Sapindaceae (i). i S. Brazil. Seeds yield oil. Dilophia T. Thorns. Cruciferae (2). 5 Centr. As. Dimacria Lindl. in Sweet. = Pelargonium L'Herit. (Geran.). Dimeresia A. Gray. Compositae (4). i Oregon. Dimeria R. Br. Gramineae (2). -20 Indomal., S. China. Dimerocostus O. Ktze. Zingiberaceae (n). 4 W. S.Am. Dimerostemma Cass. Compositae (5). i Brazil. Dimetopis DC. = Trachymene Rudge (BH.} = Didiscus DC. (Umb.). Dimorphandra Schott. Leguminosae (n. i). 10 trop. Am. D. Mora Benth. et Hook, furnishes excellent timber. 2 1 8 DIMORPHANTHERA Dimorphanthera F. Muell. (Agapetes p.p.)- Eric. (in. 2). 4 Malaya, Australia. Dimorpliantlies Cass. =Conyza L. (Comp. ). Dimorphanthus Miq. = Aralia Tourn. Dimorphism, appearing in two forms, e.g. flower, Asperula, Cassia, Exacum, Saintpaulia, and cf. Dioecism, Hfterostylism; fruit, Car- damine, Chenopodium, Dimorphotheca ; inflorescence, Trifolium ; leaf, Anisophyllea, Bidens, Dischidia, Ficus, Hedera, Heteranthera, Platyceriuin, Polygonum, Polypodium, Ranunculus, Sagittaria, Salvinia, Trapa, and cf. Water-plants; plant, Littorella ; polltn, Faramea; root, Araceae, Ficus, Jussieua, Orchidaceae; shoot, Marc- gravia, Salacia; stamens, Heeria, Monochaetum. Dimorphocalyx Thw. Euphorbiaceae (A. n. 5). 6 Indomal. Dimorphochlamys Hook. f. Cucurbitaceae (3). 3 trop. W. Afr. Dimorphocoma F. Muell. et Tate. Compositae (7). i C. Austr. Dimorphotheca Vaill ex L. Compositae (10). 20 S. and trop. Afr. There are two kinds of fr. on the head (cf. Calendula). Dinacria Haw. (Crassula p.p. EP.). Crassulaceae. 2 S. Afr. Dinebra Jacq. Gramineae (n). i trop. Afr., As. Dinemagonum A. Juss. Malpighiaceae (i). 4 N. Chili. Dinemandra A. Juss. ex Encll. Malpighiaceae (i). 6 Peru, Chili. Dinklagea Gilg. Connaraceae. i Liberia. Dinochloa Buese. Gramineae (13). 6 Malay Archip. Dinophora Benth. Melastomaceae (i). 2 trop. W. Afr. Dinoseris Griseb. Compositae (i 2). i Argentina. Dintera Stapf. Scrophulariaceae (n. 6). i trop. Afr. Dinteracanthus C. B. Clarke ex Schinz. Acanth. (iv. A). 3 S. Afr. Dioclea H. B. et K. Leguminosae (in. 10). 20 trop. Diodia Gronov. Rubiaceae (11. 10). 35 trop. and subtrop. D. niari- t/i/ia Schum. et Thonn. is common to Afr. and Am. Dioecism (-ious), it\ Rtttaceae; -flower, Compositae; -old, like a disc. Discaria Hook. Rhamnaceae. 20 Andes, N.Z. , Austr. Dischidia R. Br. (incl. Conchophyllum Blume). Asclepiadaceae (n. 4). 70 Indomal., Polynes., Austr. Epiphytes, climbing by adv. r., and DISPERSAL 223 with fleshy 1. covered by wax. The curious pitcher-plant, D. Raffle- siana Wall., besides the ordinary 1., has pitcher-1. Each is a pitcher with incurved margin, about 10 cm. deep. Into it grows an adv. r. developed from the stem or petiole just beside it. The pitcher may hang with its mouth upwards or may stand horizontally or upside down. It usu. contains a lot of dtbris, largely carried into it by nesting ants. Most contain ± rain water, so that perhaps they act as humus collectors and water reservoirs. The inner surface is waxy, so that the water cannot be absorbed by the pitcher itself, but must be taken up by the roots. Developmental study shows the pitcher to be a 1. with its lower side invaginated. The existing sp. illustrate all stages. Many, e.g. D. bengalensis Colebr., have bi-convex 1.; others have the under surface concave, e.g. D. (C.) Colly ris Wall., and the roots are de- veloped under and sheltered by the concave 1. A further invagination would lead to D. Rafflesiana. (Treub in Ann. Buitenz. III. 1883, Haberlandt's Tropenreise, p. 168, and two papers in Ann. of Bot. 1893-) DiscMdiopsis Schlechter (Dischidia p.p.)- Asclep. (n. 3). 2 N. G., Phils. Dischisma Choisy. Scrophulariaceae (n. 7). 10 S. Afr. DiscMstocalyx (Distichocalyx) Lindau. Acanth. (iv. A). 7 trop. Afr. Disciflorae (RH.}. The 2nd series of Polypetalae. Disciphania Eichl. Menispermaceae. 8 trop. S. Am. Discocactus Pfeiff. (Echinocactits p.p. E.P.). Cact. (ill. i). 3 trop. Am., W.I. Discocalyx Mez. Myrsinaceae (n). 8 Malaya, Polynesia. Discocarpus Klotzsch Euphorbiaceae (A. I. i). 3 Brazil, Guiana. Discoglypremna Prain. Euphorbiaceae (A. n. 2). i trop. Afr. Discogyne Sciilechter. Saxifragaceae (v). i New Guinea. Discolobium Benth. Leguminosae (in. 7). 6 Brazil, Paraguay. Discoluma Baill. (Chrysophylhtm p.p.). Sapot. (i). i Brazil. Discontinuous distribution, cf. Geographical Distribution. Discophora Miers (Kutnmeria Mart.). Icacin. 2 trop. S. Am. , Discopleura DC. Umbelliferae (in. 5). 2 N. Am. Discopodium Hochst. Solanaceae (2). i trop. Afr. Discostigma Hassk. = Garcinia L. p.p. (Guttif.). Discrete, separate. Disella Greene (Si da p.p.)- Malvaceae (2). 4 N. Am. Diselma Hook. f. (Fitzroya p.p.). Coniferae. i Tasmania. Disemma Labill. = Passiflora L. p.p. (Passifl.). Disepalum Hook. f. Anonaceae (i). 4 Indomal. Disparage Gaertn. Compositae (4). 5 S- Afr. Dispersis Sw. Orchidaceae (n. i). 30 trop. Afr., Madag., As. Disperma C. B. Clarke. Acanthaceae (iv. A). 7 trop. Afr. Dispersal of seeds may be occasional, as by floating trees, ice, &c., tornados, mud on birds' feet, the voiding of the crops of carnivorous birds, &c. (see Origin of Species, chap, xil), or regular, by ways which may be classed in four heads. By wind are carried directly the spores of ferns, &c., seeds of Pyrola, Orchidaceae, some Caryo- phyllaceae, &c., by censer-mechanisms (see Aconitum), Campanula, 224 DISPERSAL Caryophyllaceae, Delphinium, Iridaceae, Liliaceae, Papaver; winged seeds in Bignoniaceae Bromeliaceae, Casuarina, Millingtonia, Pinus, Zanonia; winged fruits in Abronia, Aceraceae, Bignoniaceae, Car- pinus, Dipterocarpaceae, Fraxinus, Liriodendron, Malpighiaceae, Ptelea, Pterocarpus. Rumex, Serjania, Terminalia, Tripteris. Ulmus, Ventilago ; hairs, forming a parachute mechanism, in seeds of Apo- cynaceae, Asclepiadaceae, Epilobium, Gossypium, Salix, &c., in fruits of Anemone, Clematis, Compositae, Eriophorum, Typha, Valerianaceae, &c. By animals are carried, as inside passengers, the seeds in edible fruits ( an inch wide. Fls. unisexual, sometimes all of one sex on one receptacle, sometimes intermingled with several 90°. Down tree (W.I.), Ochroma Lagopus Sw. Downy, covered with fine soft hairs. Doxantha Miers (Bignonia p.p. BH.}. Bignon. (i). i U.S. Doyerea Grosourdy ex Bello (Corallocarpus p.p. EP.). Cucurbitaceae (2) (Myrtaceae BH.). 2 W.I., Venezuela. Draba Dill, ex L. Cruciferae (4). 150 N. temp, and arctic, and south-west. N. Am.; 5 in Brit, (whitlow-grass), incl. D. verna L. which occurs in Eur. in a vast number of vars. which breed true, and were distinguished as sp. by Jordan (cf. works on Mendelism). Most are tufted, ± xero. with hairy or fleshy 1. Dracaena Vand. Liliaceae (vi). 40 Old World trop. Mostly trees, whose stems branch and grow in thickness (extra-fascicular cambium). The famous dragon-tree of Teneriffe (D. Draco L.), blown down in 228 DRACAENA 1868, was 70 ft. high and 45 ft. in girth and was supposed to be 6000 years old. A resin exudes from the trunk of this sp. (dragon's blood); the original dragon's blood appears to be that of D. < inna- baH Half. f. (Socotra). \Cf. Pleomela, and see N. E. Brown in Kew Bull. 1914, p. 273.] Dracamina Nieuwland (Cardamine p.p.). Crucifer. (2). i Atl. U.S. Dracocephalum L. Labiatae (iv). 40 N. temp. Dracontioides Engl. Araceae (iv). i S. Brazil. Dracontium L. Araceae (iv). 10 trop. Am. The sympodial rhiz. gives rise yearly to one enormous 1. and an infl. The 1. has 3 chief divisions, and the lat. ones develope dichot. at first. Fl. ? with P. Dracontomelum Blume. Anacardiaceae (2). 6 Malaya to Fiji. Dracophyllum Labill. (excl. Sphe.notoma R. Br.). Epacridaceae (2). 25 N.Z., Austr., New Caled. The sheathing 1. leave ring-scars when they fall. Dracunculus (Tourn.) Adans. Araceae (vn). 2 Medit. Fert. like Arum. Dragon-root (Am.), Arisaema; -'s blood, -tree, Dracaena. Drakaea Lindl. Orchidaceae (n. 2). 4 Austr. Drake-Brockmania Stapf. Gramineae (10). i Brit. Somaliland. Draperia Torr. Hydrophyllaceae. i California. Drapetes Banks. Thymelaeaceae. 6 S. Am , N.Z. to New Guinea Drebbelia Zoll. Olacaceae. i Bali. Dregea E. Mey. Asclepiadaceae (11. 3). 5 S. Afr. to China. Prejera Nees (Jacobinia Bff.}. Acanthaceae (iv. B). 3 trop. Am. Drejerella Lindau. Acanthaceae (iv. B). 3 W. Indies. Drepananthus Maingay ex Hook. f. Anonaceae (4). 2 Malay Penin. Drepanium, a monoch. cyme with each successive branch on same side of relatively main axis, and in same plane, Juncus, Marantaceae. Drepanocarpus G. F. W. Mey. Leguminosae (in. S). 8 trop. Am., A IV. Drepanolobus Nutt. ex Torr. et Gray=Hosackia Dougl. p.p. Drepanostemma Jumelle et Perrier. Asclep. (n. i). i Madag. Driessenia Korth. Melastomaceae (i). 5 Malaya. Drimia Jacq. Liliaceae (v). 25 Afr. Drimiopsis L. et Paxt. Liliaceae (v). 5 S. and trop. Afr. Cult. orn. fl. Drimycarpus Hook. f. Anacardiaceae (4). i E. Hiinnl. Drimys Forst. Magnoliaceae. 10 S. Am., and N.Z. to Borneo. There is a distinction between calyx and corolla (cj. Illicium). The bark of D. IVinteri Forst. (Winter's bark) is medicinal. Drimyspermum Reinw. = Phaleria Jack (Thymel.). Drinks, cj. Alcohol ; infusion drinks from seeds of Coffee, T/iea, Theo- broma, Cola, Paitllinia, 1. of Thea, Ilex, Cat ha, Priva, Stachytarpheta, and root of Cichotium. Driodium (Cl.), a dry thicket formation. Drip-tip, <•/. Acer, and Artocarpus, Begonia, Boehnieria, Ficus. Droguetia Gaudich. Urticaceae (5). 4 trop. As., Afr., Madag. Droogmansia De Wild. (Desmodium p.p.). Legu. (in. 7). '4 trop. Afr. DRUDEOPHYTUM 229 Drooping1 of young' 1., Aesculus, Bauhinia, Cinnamomum, Dryoba- lanops; of young shoot, A/nherstia, Brownea, Saraca. Dropper, young immature bulb. Dropseed grass (Am.), Sporobohts, Muhlenbergia. Dropwort, Filipendttla vitlgaris Moench. ; water-, Ocnanthe. Drosera L. Droseraceae. 90 trop. and temp. 3 in Brit, (sundew). D. rotund if olia L. abundant in bogs. Herbs usu. with creeping rhiz. and rosettes of 1., insectivorous. The blade of the 1. is circular in some sp., elongated in others, and is set with curious tentacles; these are emergences containing vascular bundles and ending in swollen reddish heads which secrete a sticky glistening fluid. Flies and other insects mistaking it for honey are held by it. The tentacles are exceedingly sensitive to continued pressure even by the lightest bodies ; the result is to cause an inward and downward movement of the head of the tentacle, finally placing the fly upon the blade of the 1. At the same time the stimulus passes to the surrounding tentacles causing them also to bend downwards to the same point. The victim is thus smothered and now the glandular heads of the tentacles secrete a ferment which acts upon the proteitls and brings them into solution, when they are taken up by the 1. Afterwards the tentacles expand once more and recommence the secretion of the sticky fluid. The food thus obtained is of benefit to the pi., though it can live without it. D. is able to live in very poor soil. The extra materials obtained are devoted chiefly to seed-production. If the stimulus produced by the capture of an insect be very power- ful, the 1. itself may bend into a cup form, and this feature is very marked in some sp. , the 1. bending almost double over the prey. The fls. of the Brit. sp. rarely open, but self-pollinate in bud. Droseraceae (EP., BH.}. Dicots. (Archichl. Sarraceniales ; Resales BH.}. 5 gen., too sp. , Drosera cosmopolitan, the rest more local. Herbs, usu. with perenn. rhiz. and rosettes of 1. ; Aldrovanda a water-plant. All are insectivorous ; Dionaea and Aldrovanda have sensitive 1. which shut up when touched, the others catch their prey by sticky tentacles upon the 1. (see genera). Us. usu. in cin- cinni, rarely in racemes or sol., $, reg. , 5 — 4-merous, usu. hypog. K (5) ; C 5, imbr. or conv. ; A usu. 5, pollen in tetrads (cf. Erica- ceae); G (2, 3, or 5) ; plac. usu. parietal, rarely axile or free-central ; style long ; stigmas simple or branched ; ov. 3 — oo , anatr. Loculic. caps. ; seed with endosp. and small basal embryo. Genera: Dionaea, Aldrovanda, Drosophyllum, Drosera, Roridula. Drosophyllum Link. Droseraceae. D. liisitanicum Link, Morocco, Portugal, S. Spam. The 1. have glands of two kinds — stalked, secreting a sticky fluid (cf. Drosera), and sessile, which only secrete when stimulated by nitrogenous matter, and then secrete a digestive ferment. Insects alight on the glands and are entangled; they struggle for a while and finally sink down and die, and are digested by the ferment. The taller glands have no power of movement, but are able to secrete a ferment as well as the sessile ones. Drudea Griseb. Caryophyllaceae (i. 3). i Peru. Drudeophytum Coulter et Rose (Arracacia p.p.). Umb. (in. 4). ? N. Am. 23° DRUGS Drugs are obtained from oo pi. : cf. e.g. Abrus, Aconitum, Aegle, Aloe, Alpinia, Alstonia, Anckusa, Aralia (ginseng), Atropa (belladonna), Bamboos, Brayera, Brucea, Canella, Cannabis (hemp), Capsicum, Carica, Carum (caraway), Cascarilla, Cassia (senna), Cinchona (quinine, £c.), Cinnamomum (camphor), Cistus, Citrullus (colocynth), Citrus, Coix, Cola, Colchicum, Colutea, Commiphora, Convolvulus (scammony), Croton, Curcuma, Datura, Digitalis (digitalin), Dorema (gum-ammoniacum), Dracaena, Drimys, Elettaria (cardamom), Erythroxylon (cocaine), Eucalyptus, Eugenia, Ferula (asafoetida), Gentiana, Glycyrrhiza (liquorice), Guaiacumjpomoea ( jalap), Lewisia, Lindera, Mentha, Menyanthes, Myroxylon, Papaver (opium), Pence- dan urn (dill), Picraena, Pilocarpus, Piper, Podophyllum, Polygala, Pringlca, Rhammts (cascara), Rheum (rhubarb), Ricinus (castor-oil), Ruta (rue), Santalina, Sassafras, Schoenocaulon, Smilax (sarsa- parilla), Strophanthus, Strychnos (strychnine), Styrax, Tamarindus, Toluifera, Trigonella (fenugreek), Uragoga (ipecacuanha), Urginea (squill), Veratrum, Verbascum, Zingibcr (ginger), &c. See Dragen- dorff, Die Heilpflanzen, Stuttgart, 1898. Drummondita Harv. (Philotheca EP.). Rutaceae (i). Austr. Drupaceae ( Warming) = Rosaceae (sub-order v). Drupaceous, drupe-like; drupe, a fleshy fr. with hard endocarp, within which is the seed, Cornaceae,Juglans, Prunus. Drupatris Lour. Inc. sed. i Cochin-China. Drusa DC. (Bowlesia BH.}. Umbelliferae (i. 2). 3 Am., Canaries. Dry fr., esp. achenes and nuts (indehiscent), follicles, legumes and capsules (dehiscent); and schizocarps ; -ing pi., cf. Collecting. Dryadaea L. = Dryas L. (Ros.). Dryadorchls Schlechtei. Orchidaceae (n. 10). 2 New Guinea. Dryandra R. Br. Proteaceae (n). 50 Austr. Like Banksia. Dryas L. Rosaceae (in. 2). 2 arctic. D. octopetala L. (alpine in Brit.) is androdioec. in the Alps. Style feathery after fert. (cf. Clematis, Geum). Drymaria Willd. Caryophyllaceae (i. 3). 30 trop. and S. temp. Drymocallis Fourr. (Potentilla p.p. ). Rosac. (in. 2). 20 N. Am. Drymoda Lindl. Orchidaceae (n. 16). i Burma. Drymoglossum Presl. Polypodiaceae. 10 palaeotrop., Jap. , Chi. D. carnosum (Wall.) J. Sm. has succulent leaves. Drymonia Mart. Gesneriaceae (i). 15 trop. Am., W. Ind. Drymophila R. Br. Liliaceae (vn). 2 E. Austr., Tasm. Drymophloeus Zipp. Palmaceae (iv. i). 15 Malaya, N. Austr. Drymotaenium Makino. Polypodiaceae. 2 Japan, Formosa. Drynaria (Bory) J. Sm. Polypodiaceae. 20 palaeotrop. D. querci- folia (L.) J. Sm. has dimorphic 1., some projecting, assimilating and spore-bearing, the others small, close to rhiz., collecting humus. Dryobalanops Gaertn. f. Dipterocarpaceae. 4 Borneo, Sumatra. D. aromatica Gaertn. &c. yield Borneo or Sumatra camphor, used chiefly in China. The young 1. are red, and hang down. Dryopetalon A. Gray. Cruciferae (2). i Mexico. Dryopteris Adans. (incl. Cyclosorus, Lastrca, Leptogramma, Mcniscium, Phegopteris). 1000 cosmop. D. Filix-mas (L.) Schott DURO1A 231 (shield-fern) and others Brit. This sp. has a stout nearly erect rhiz. with large pinnate 1. The var. cristatum Moore shows apospory. Dryostachyum J. Sm. Polypodiaceae. 5 Malay Archipelago. Drypetes Vahl. Euphorbiaceae (A. I. i). 15 N. Brazil to W.I. Drypis L. Caryophyllaceae (n. i). r E. Medit. Duabanga Buch.-Ham. Sonneratiaceae. 2 Indomal. Dubautia Gaudich. Compositae (5). 6 Hawaiian Is. Duboisia R. Br. Solanaceae (5). 2 Austr., New Caled. Duboscia Bocq. Tiliaceae. i trop. W. Afr. Dubouzetia Panch. Elaeocarpaceae. 6 New Caled. Dubrueilia Gaudich. = Pilea Lindl. p.p. (Urtic.). Duchesnea Smith (Fragaria p.p. BH.}. Rosac. (in. 2). 2 S. As. Duck's meat, duckweed, Lemna. Ducosia Vieill. ex Guillaumin (Dubouzetia p.p.). Elaeo. i N. Cal. Ducrosia Boiss. Umbelliferae (in. 6). 3 VV. As. Dudleya Britton et Rose (Cotyledon p.p. EP.). Crassul. 60 Calif. Duffin bean, Phaseohis lunatus L. Dugesia A. Gray. Compositae (5). i Mexico. Dugezia Montr. Inc. sed. i New Caledonia. Duguetia A. St. Hil. (Aberemoa Aubl.). Anonaceae (i). 30 trop. S. Am., W.I. D. quitarensis Benth. &c. furnish Jamaica and Cuba lancewood. Fr. formed of the individual berries or achenes united to the fleshy recept. Dulacia Veil. (Liriosma BH.). Olacaceae. 15 trop. S. Am. Dulcis (Lat.), sweet. Dulichium Pers. (Websteria Wright). Cyper. (i). i All. N. Am. Dumasia DC. Leguminosae (ill. 10). 3 trop. As., Afr., Madag. Dumb-cane, Dieffenbachia Seguine Schott. Dumerilia Lag. ex DC.— Jungia L. (Comp.). Dumoria A. Chevalier. Sapotaceae (i). i trop. Afr. Good wood. Dumose, shrubby. Dunalia H. B. et K. Solanaceae (2). 8 W. S.Am. to Mexico. Dunbaria Wight et Arn. Leguminosae (ill. 10). 15 trop. As., Austr. Dune-plants, cf. Ammophila, Carex, Ety»uts, Hippophae, &c. Dunnia Tutcher. Rubiaceae (i. 4). i China. Duparquetia Baill. (Oligostemon BH.}. Legumin. (n. =,). i W. Afr. Dupontia R. Br. (Graphcphonun BH.). Gramin. (10). 2 Arctic. Duramen, the heart-wood. Durandea Planch. (Hiigonia EP.). Linaceae. 5 New Guinea, Polynes. Durandia Boeck. Cyperaceae (n). i Costa Rica. Duranta L. Verbenaceae (i). 10 trop. Am., W.I. Cult. orn. shrubs. Duravia Greene (Polygonum p.p.)- Polygon, (n. 2). 4 Calif. Durian, Durio zibelhinus Murr. Durieua Boiss. et Reut. = Daucus Tourn. p.p. (Umbell.). Durio Adans. Bombacaceae. 7 Indomal. D. zibet hinns Murr. pro- duces the durian fr., with delicate flavour and disagreeable smell. Seed with fleshy aril. Duroia L. f. Rubiaceae (i. 8). 10 S. Am. Myrmecophilous (cf. Acacia). D. petiolaris Hk. f. and D. hirsuta K. Sch. have stems swollen just below the infl. The swollen part is hollow and entrance 232 DUROIA is obtained by two longitudinal slits ; it is inhabited by ants, which bite through the thin tissue of the slits. D. saccifera Benth. et Hk. f. has 'ant-houses' on the 1. At the base, on the under side, are two pear-shaped organs formed by outgrowth of the 1. The entrance is upon the upper side, protected from rain by a little flap. Dusenia O. Hoffm. (Duseniella EP.}. Compos (12). i Patagonia. Duseniella K. Schum. Compositae (12). i Patagonia. Dussia Krug et Urb. Leguminosae (ill. i). i Martinique. Dutaillyea B.iill. Rutaceae (i). i New Caled. Dutch clover, Tn 'folium repens L. ; -grass (W.I.), Paniaim molle Sw.; rushes, Equist-tum. Dutchman's breeches, Dicentra; -pipe, Aristolochia. Duthiea Hackel. Giamineae (10). 2 Kashmir, Afghanistan. Duvalia Haw. Asclepiadaceae (il. 3). 15 S. Afr. Duvaliella Raill. Dipterocarpaceae. i Penang. Duvaljouvea Palla. (Cyperus p.p.) Cyperaceae (i). 2 As., Eur. Duvaua Kunth = Schinus L. p.p. (Anacard.). Duvaucellia Bowditch. Oleaceae. i trop. Afr. Duvernoya E. Mey (Adhatoda p.p. BH.}. Acanthaceae (iv. B). 25 Afr. Dwale, Atropa Belladonna L. Dyckia Schult. f. Bromeliaceae (2). 57 S. Am. Dyera Hook. f. Apocynaceae (I. 3). 3 Malaya. Dyerophyton O. Ktze. ( Vogella Lam.). Plumbaginaceae 3 palaeotrop. Dyer's greenweed, Genista tinctoria L. ; -weld, Reseda liitea, L. Dyes (vegetable), cf. esp. Alkainia, Baphia (cam wood), Bixa (an- natto), Caesalpinia (sappan), Carthamits (rouge), Chlorophora (fustic), Cladrastis, Crocus (saffron), Crozofihora, Curcuma (turmeric), Gar- rtnia, Haetnatoxylon (logwood), Indigo/era (indigo), Isatis (woad), LctK'SMn'a, Madura (tiistic), Morinda, Nopalea, Peganum (turkey red), Pterocarpiis, Reseda, Rhamitus, Rkiis, Rubia (madder). Dypsidium Bail!. (Dypsis p.p.) Palmae (iv. i). 3 Madagascar. Dypsis Noronha ex Thou. Palmae (iv i). 10 Madag. Dyschoriste Nees (Calophanes BH.}. Acanthaceae (iv. A). 50 trop. Dysodia Cav. Compositae (6). 35 Am. Dysodiopsis Rydberg (Dysodia p.p.). Compositae (6). i. S.W. U.S. Dysolobium Prain. Leguminosae (ill. 10). 4 Indomal. Dysophylla Blume. Labiatae (vi). 20 E. As., Austr. Dysopsis Baill. Euphorbiaceae (B. n). i Andes, Juan Fernandez. Dysoxylum Blume. Meliaceae (ill). 120 Indomal. D. Fraseranum Benth. (E. Austr.; Austr. mahogany) and others, good timber. Dysphania R. Br. Caryophyllaceae (I. 5). 3 Austr. L. alt. Dyssochroma Miers. Solanaceae (3). 4 trop. Am. E- (Lat. pref.), without : -bracteate, -calcarate, -costate, -dentate, -glandulose. -marginate (notched), -rostrate, -strophiolate, -val- vate, &c. And cf. Ex. Eagle-wood. Aijuilaria, Agallocha Roxb. (Burma). Earina Lindl. Orchidaceae (n a. mi. 6 N.Z., Polynesia. Earth-nut, Arachis hypogaca L , Conopodium demidatuni Koch. East Indian arrowroot, Curcuma, Tacca; -hemp, Crotalariajuncea'L.; -rosewood, Dalbergia latifolia Roxb.; walnut, Allrizzia Lebbtk Benth. Eastwoodia Brandegee. Compositae (3). i Lower California. ECHIDNOPSTS 233 Eatonella A. Gray. Compositae (6). 2 Nevada, California. Eatonia Kafin. Gramineae (10). 5. N. Am. Eau de Creole, Mammea. Ebenaceae (EP., BH.}. Dicots. (Sympet. Ebenales £P., BH.). 5 gen., 320 trop. (esp. Indomal.), trees and shrubs with alt., opp. or whorled, simple, leathery, usu. entire 1. Fls. axillary, sol. or in small cymes, reg. , usu. dioec., bracteolate, 3 — y-merous. (K) persistent; (C) convolute ; A epipet. at base of tube, usu. in 2 whorls but fre- quently so by branching; stds. usu. present in ? fls., (G) 2 — i6-lpc., with i — 2 anatr. ov. pend. in each loc. ; styles 2 — 8, free or united below. Fr. usu. a berry with fewer seeds than there were ovules, sometimes dehiscent. Embryo straight or slightly curved, in abun- dant cartilaginous endosp. Many yield valuable wood, e.g. Diospyros. Genera: Royena, Euclea, Maba, Diospyros, Tetraclis. Ebenales. The 4th order of Sympetalae (EP.), the 6th (Bff.). Ebeneous, black as ebony. Ebenus L. Leguminosae (in. 7). 14 Medit. to Beluchistan. Ebermaiera Nees (Staurogyne £P.). Acanthaceae (i). 40 trop. exc. Afr. Ebony, wood of many Ebenaceae, esp. Diospyros, blackened by a deposition of a gum-resin in the heartwood ; Gtirke gives a list of forms, Nat. Pfi. IV. i, p. 164; Ceylon and S. Indian ebony, the most important, is from D. Ebenum Koen. ; American-, green-, Jamaica-, W. Indian-, Brya Ebenus DC. Eburopetalum Becc. Anonaceae (4). i Borneo. Eburophyton A. A. Heller (Chloraea p.p.). Orch. (n. 2). i Calif. Ecad (Cl.), a habitat form due to origin by adaptation. Ecastaphyllum P. Br. (Dalbergia p.p. EP-). Legum. (in. 8). 6 trop. Am., W.I Ecballium A. Rich. Cucurbitaceae (3). i Medit., E.Elaterium A. Rich, (squirting cucumber). The ripe fr. is highly turgid; as it drops from the stalk, a hole is made in its lower end, and through this the contraction of the pericarp squirts the seeds, mixed with a watery fluid. A purgative (eiaterium) is prepared from the fr. Ecbolium Kurz. Acanthaceae (iv. B). 8 trop. Afr. Ecbolium Riv. ex L.=Justicia L. (Acanth.). Ecclinusa Mart. Sapotaceae (i). 6 N. Brazil. Eccremocarpus Ruiz et Pav. Bignoniaceae (3). 3 Peru. See fam. The valves of the fr. hang together at the top. Ecdeiocolea F. Muell. Restionaceae. i S.W. Austr. Ecdysanthera Hook, et Arn. Apocynaceae (n. i). 10 Indomal., China. Echeandia Ortega. Liliaceae (in). 6 Mexico to Guiana. Echenais Cass. =Cnicus Tourn. p.p. (j5//.) = Cirsium Adans. p.p. Echetrosis Phil. Compositae (7). t temp. S. Am. Echeveria DC. (Cotyledon p.p.). Crassulaceae. 70 Am. Echidiocarya A. Gray (Plagiobotrys p.p. EP.). Borag. (iv. 2). i. Arizona. Echidnium Schott. Araceae (iv). 2 trop. S. Am. Echidnopsis Hook. f. Asclepiadaceae (11. 3). 4 E. Afr. Xerophytes with succulent stems. 234 ECHfN- Echln- (Gr. pref.), spiny. Echinacanthus Nees. Acanthaceae (iv. A). 8 Himal., Java. Echinacea Moench. (Rudbeckia p.p.). Compositae (5). 2 N. Am. Echinantnus Cerv. (Tragus EP.}. Gramineae (3). None described. Echinaria Desf. Gramineae (10). i Medit. Echinocactus Link et Otto. Cactaceae (in. i). 250 Texas to Chili. Ribbed cacti (see fam.). Echinocarpus Blume (Sloanea p.p. EP.}. Tiliaceae. 6 E. As., Austr. Ecbinocephalum Gardn. (Melanthera p.p.). Comp. (5). 3 Brazil. Echlnocereus Engelm. (Cereus Haw. BH.}. Cactaceae (in. i). 50 Am. Echinochlaenia Borner. (Carex p.p .). Cyper. (in). 3 Tasm., N.Z. Echinochloa Beauv. = Panicum L. p.p. (Gramin.). Echinocystis Torr. et Gray. Cucurbitaceae (4). 25 Am. Tuberous climbing herbs. E. lobata Torr. et Gray often cult. Its tendrils are very sensitive and nutate rapidly ; they become straight and erect as they come round towards the main axis, thus avoiding contact. Echinodorus Rich. Alismaceae. 20 Am., Afr. Echinolaena Desv. (Panicum p.p. EP.). Gramineae (5). 2 S. Am., Madag. Echinopanax Decne. et Planch. (Fatsia BH.}. Araliaceae (i). i N. Am., Japan, E. horridus D. et P., an obstacle to travellers. Echinopepon. Naud. (Echinocystis Torr. et Gray). Cucurb. (4). 15 N. Am. Echinophora Tourn. ex L. Umbelliferae (in. i). 8 Medit. One cpl. is aborted. The umbel has one ? fl. in the centre, surrounded by s fls. The spiny stalks of the latter enclose the fr. Echlnopogon Beauv. Gramineae (8). i Austr., N.Z. Echinops L. Compositae (n). 75 E. Eur., Afr., As. The spherical head is really cpd., formed of oo small i-fld. heads, each with its own invol. The fls. are largely visited by bees. Echinopsilon Moq. =Chenolea Thunb. (j5//.). = Bassia All. p.p. Echinopsis Zucc. (Cereus p.p.). Cactaceae (in. i). 25 S. Am. Echinopterys A. Juss. Malpighiaceae (i, but forming a link between i and 2, as it has a flat torus), i Mexico. Mericarp spiny. Echlnospermum Sw. (Lappula p.p. EP.}. Bnraginaceae (iv. 2). 50 temp. E. Lappula Lehm. cult. The fls. change from white to red and blue (see fain.). Fr. hooked. Echinostachys Brongn. (Aechmea p.p.). Bromel. (4). 6S.Am. Cult. orn. infl. Echinothamnus Engl. Passifloraceae. i S.W. Afr. Echiochilon Desf. Boraginaceae (iv. 5). 2 N. Afr., Aden. Echioglossum Blume (Cleisostoma BH.}. Orch. (n. 20). 3 Indomal. Echioides (Tourn.). Desf. =Nonnea Medic. (Borag.). Echirospermum Saldanha da Gama. Leguminosae. Nomen. Echltes P. Br. Apocynaceae (n. i). 40 Am., W.I. Echium Tourn. ex L. Boraginaceae (iv. 5). 30 Eur., Medit. E. vulgare'L. (viper's bugloss) Brit, (offic.). FL •)-, protandr., gynodioec., bee -visited. Echyrospermum Schott. Leguminosae (inc. sed.). Nomen. Ecklonea Steud. (Trianoptiles EP.}. Cyperaceae (n). 2 Austr., S. Afr. EDIBLE PRODUCTS 235 Eclipta L. Compositae (5). 4 Austr. , S. Am. Eclopes Gaertn.= Relhania L'Herit. p.p. (Comp.). Ecology, relations of plants to their environment. Economic Botany, the study of pi. from the point of view of their uses to man ; -products, of value in the arts as food, &c., may be divided into groups: (i) gums, resins, rubbers or caoutchoucs, gutta-perchas, &c. , (2) oils, (3) dyes and tanning stuffs, (4) fibres, (5) drugs, (6) edible products, (7) timbers, and (8) miscellaneous, such as cork, vegetable ivory, weights, beads, &c. For more detail see under each of these heads, and consult Wiesner, Die Rohstoffe des Pflanzen- reichs, Vienna, 1903 ; Watt, The Commercial Products of India ; De Candolle, Origin of Cultivated Plants ; Kew Bulletin and other technical journals. Ecphymacalyx Pohl. Inc. sed. Nomen. Ecpoma K. Schum. Rubiaceae (i. 7). i trop. Afr. Ecpomanthera Pohl. Inc. sed. Nomen. Ectadiopsis Benth. Asclepiadaceae (i). 6 trop. and S. Afr. Ectadium E. Mey. Asclepiadaceae (i). 2 S. Afr. Ectasis D. Don= Erica L. p.p. (Eric.). Ectinocladus Benth. Apocynaceae (n. i). i W. Afr. Ectotropic, hyphae running between epidermal cells. Ectrosia R. Br. Gramineae (ro). 4 Austr. Edaphic, of the soil. Eddoes (W. I.), Colocasia antiqitorum Schott. Edelweiss, Leontopodiuvi alpinum Cass. Edgaria C. B. Clarke. Cucurbitaceae (2). i Sikkim. Edgeworthia Meissn. Thymelaeaceae. 2 Himalaya, China. Edible products, or foodstuffs in a wide sense, are obtained esp. from stores of reserves in pi., e.g. the seeds of cereals, Avena (oat), Coix, Eleusine, Euchlaena, Hordeum (barley), Oryza (rice), Panicum (millet), Paspalum, Pennisetum (bulrush millet), Secale (rye), Setaria, Sorghum (Guinea corn), Triticum (wheat), Zea (maize), Zizania, &c.; the seeds of many JLeguminosae, e.g. Arachis (peanut), Cajanus (cowpea), Cicer (chickpea), Uolichos, Glycine (soy), Lathyrus, Lens (lentil), Phaseolus (beans, &c. ), Pisum (pea), Psophocarpus, Vicia (bean), Voandzeia, &c. ; the seeds of Araucaria, Bertholletia (brazil- nut), Brosimum, Carya, Caryocar, Castanea (chestnut), Castano- spermum, Cocos (coconut), Corylus (hazelnut, filbert), Dioon, Fago- pyrum (buckwheat), Juglans (walnut), Lecythis, Nelumbium, Pistacia (pistachio), Telfairia, Trapa, &c. ; the underground reserves of Allium (onion, &c.), Alocasia, Araceae, Arracacia, Arum, Beta (beetroot), Brassica (turnip, &c.), Canna, Colocasia (taro), Commelina, Daucus (carrot), Dioscorea (yam), Helianthus (Jerusalem artichoke), Ipomoea (sweet potato), Manihot (cassava, tapioca), Maranta (arrowroot), Ophiopogon, Oxalis, Pachyrhizus, Peucedanum (parsnip), Plec- tranthus, Priva, Raphanus, Scilla, Scorzonera, Selinum, Stachys, Solanum (potato), Tragopogon, Tropaeolum, Ullucus, Xanthosoma, &c. ; from reserves in the stems of many trees, esp. palms, Acer (sugar), Alsophila, Arenga (sugar), Borassus (sugar, sago), Caryota (sugar, sago), Cycas, Cocos (sugar), Encephalartos, Metroxylon (sago), Oreodoxa, Saccharum (sugar), &c.; from reserves in I. and injl. in 236 EDIBLE PRODUCTS Brassica, Cynara, many palms, &c. Other 1. are also eaten, e.g. Apium (celery), Ceratopteris, Clienopodium, Crambe (sea-kale), Eremurus, Foeniculum (fennel), Lactuca (lettuce), Lepidium (ciess), Myrrhis, Rheum (rhubarb), Scorzonera, Spinacia (spinach), Tetra- gonia, &c.; similarly the young shoots of Asparagus, Bambusa, &c. This leads on to condiments or spices (see Condiments). ManyyiVj/y/r. are also eaten ; the greatest food value attaches to Artocarpus (bread-fr., jak), Musa (banana, plantain), and Phoenix (date), but others are Aberia, Achras, Anacardium (cashew), Ananas (pine-apple), Anona (custard-apple, &c.), Artaboirys, Averrhoa, Bactris, Benincasa, -Blighia, Borassus (palmyra- palm), Byrsonima, Carica (papaw), Celtis, Cereus, Chrysobalanus, Chrysophylhim, Citrullus, Citrus (orange, lemon, &c.), Coccinia, Coccoloba, Cucumis (melon, cucumber), Cucurbita (pumpkin, marrow), Debregeasia, Decaisnea, Dialium, Diospyros (kaki), Durio (durian), Eriobotrya (loquat), Eugenia (rose-apple, &c.), Feronia, Ficus (fig), Fragaria (strawberry), Garcinia (mangosteen), Gaylussacia (huckleberry), Glycosmis, Hymenaea, Juniperus, Lantana, Lapagevia, Litchi (litchi), Macadamia, Maclura, Mammea, Mangifera (mango), Mimu- sops, Monstera, Morus (mulberry), Myrtus, Nephelium (rambutan), Opuntia(pncklypear),Osmanthus, Pappea, Passiflora(granadilla,&c.), Pereskia, Persea (avocado), Peumus, Prunus (plum, cherry, apricot, peach, &c.), Psidium (guava), Punica (pomegranate), Pyrus (pear, apple, &.C.), Ribes (gooseberry, currant), Rubus (raspberry, &c.), Sechium, Spondias, Tamarindus (tamarind), Vaccinium (cranberry, bilberry, &c.), Vitellaria, Vitis (grape), Zanthoxylum, Zizyphus, &c. Drinks (q.v., and Alcohol), narcotics (y.v.), &c. may also be men- tioned here. For food for animals cf. Fodder, and Bee-flowers. Edithcolea N. E. Br. Asclepiadaceae (n. 3). i E. Afr., Socotra. Edmondia Cogn. Cucurbitaceae (3). i Venezuela. Edraianthus A. DC. = Wahlenbergia Schrad. (BH.) = Hedraianthus A. DC. Eflulis (Lat.), edible. Edwardsia, Salisb. = Sophora Linn. Edwinia A. A. Heller (JTamesia p.p.). Saxifnigaceae (in). 2 N. Am. Eel-grass, Zostera ; -trap pi. , Biovularia, Genlisea, Polypompholyx, Utriciilaria. Eenia Hiern et Sp. Moore. Compositae (4). i Damaraland. Effete, function less from age. Effigurations, outgrowths of receptacle, Capparidaceae, Orchidaceae, Passifloraceae. Effuse, expanded. Efulensia C. H. Wright (Deida/nia EP.). Passifloraceae. i trop. Afr. Eganthus Van Tiegh. Olacaceae. i Brazil. Egassea I'ierre ex Wildem. Scyiopetalaceae. 3 trop. Afr. Egeria Neraud. Rubiaceae. Nomen. Egg -apparatus, the ovum and synergidae ; -fruit, -pi., Solan um Melongena L. Eggersia Hook. f. (Neca p.p. El\}. Nyctaginaceae. i W. Indies. Eglantine, Rosa l'.^!antei ia L. Egletes Cass. Compositae (3). 6 trop. Am., Mexico. ELAEOCARPACEAE 237 Egyptian bean, lotus, Nclubium, Nyinphaea ; -lily, Richardia. Ehretia L. Boraginaceae (n). 40 trop., chiefly %.. Timber. Ehrharta Thunb. Gramineae (7). 25 S. Afr., .Ylasc., N.Z. Useful pasture grasses for sandy soil. Eichhornia Ivunth. Pontederiaceae. 5 S Am., W.I. The sympodium is very complex. Each shoot in turn is pushed to one side by the axillary shoot of its last 1. but one ; with this shoot it is combined, however, up to the last 1. of the axillary shoot. After leaving the axillary shoot, each shoot bears another 1. , and then ends in the infl., which is enclosed in a spathe, and at first glance appears to spring from the stalk of the last 1. In E. azarea Kunth. the fls. are di- morphic, in E. crassipes Solms trimorphic heterostyled. This last sp. has, when floating freely, large bladder-like swollen petioles, but in soil these are not nearly so large. They cause the plant to float high and it is easily blown about by wind, and has become a very troublesome weed (water hyacinth) in Florida, Java, Australia, &c. Eichleria Progel. Oxalidaceae. 2 S. Brazil. Eichlerodendron Briquet (Xylosma p. p.). Flacourt. (4). i Brazil. Eicosia Lilume. Orchidaceae. Nomen. Ekebergia Sparrm. Meliaceae (in). 12 S. and trop. Afr., Madag. Elachanthera F. Muell. Liliaceae (x). i Austr. Elachanthus F. Muell. Compositae (7). i temp. Austr. Elacholoma F. Muell. et Tate. Pedaliaceae. i Austr. Elaeagla Wed. Rubiaceae (i. 3). 2 N. Andes. Elaeagnaceae (EP., Bff.). Dicots. (Archichl. Myrtiflorae; Daph- nales /?//.). 3 gen., 25 sp. , chiefly on steppes and coasts, % ; much branched shrubs, often with leathery L, entire, opp. or alt., and covered, as are all parts, with scaly hairs. There are frequently thorns on the surface (reduced shoots). Infl. racemose; fls. § or unisexual, 2- or 4-merous. In the 3 the recept. is often flat, but in the ? or ? fl. it is tubular as in Thymelaeaceae, and may be fused with the ovary. No petals. Sta. as many, or twice as many, as sepals. G i with one erect anatr. ov. Pseudo-drupe. Seed with little or no endosp. Chief genera: Hippophae, Elaeagnus. Elaeagnus (Tourn.) L. Elaeagnaceae. 20 As., Eur., N. Am. (oleaster). The fr. of some is ed. Cult. orn. shrubs. Elaeis Jacq. Palmae (iv. 2). 2, one trop. Am., the other, E. guine- ensis [acq. trop. Afr. (oil-palm) from whose fr. the palm-oil, used for railway axles &c., is obtained by boiling. Elaeocarpaceae (EP. ; Tiliaceae p.p. BH.). Dicots. (Archichl. Mai- vales). 7 gen., 1 20 sp. trop. and subtrop., trees and shrubs with alt. or opp., stip. 1., and racemes, panicles or dichasia of fls. Disc usu. present. K 4 or 5, free or united, valvate ; C 4 or 5, rarely united, often o, the petals often much divided at the ends, valvate or imbr. but never conv.; A oo , free, on the disc, which is some- times developed to an androphore ; anthers 2-loc. usu. opening by two pores (sometimes confluent) at the apex ; G sessile, with 2 — oo (rarely i) loc.; ovules in each loc. oo or 2, anatr., pend. with ventral raphe; style simple, sometimes lobed at apex. Capsule or drupe; embryo straight, in abundant endosp. Chief genera: Elaeocarpus, 238 ELAEOCARPACEAE Sloanea, Aristotelia. The grounds upon which they are separated from Tiliaceae are chiefly anatomical. Elaeocarpus Burm. ex L. Elaeocarpaceae. 70 trop. Elaeodendron Jacq. f. (Cassine p.p. EP.). Celastraceae. 30 trop. and subtrop. E. croceum DC. (saffranhout, S. Afr.) good timber. Elaeogene Miq. Euphorbiaceae (inc. sed.). i Sumatra. Elaeoluma Baill. (Myrsine p.p. BH., Chrysophylhim p.p. EP.}. Sapotaceae (l). i Brazil Elaeophorbia Stapf. (Euphorbia p.p.). Euphorb. (A. n. 8). i trop. Afr. Elaeoselinum Koch ex DC. Umbelliferae (in. 7). 8 W. Medit. Elaphoglossum Schott. Polypodiaceae. 330 trop. and subtrop. Elaphrium Jacq. = Bursera Jacq. (Burs.). Elater, Equisetnni. Elaterioides O. Ktze. (Elateriospermum Bl.). Euph. (A. n. 3). 2 Malaya. Elateriopsis Ernst (Cyclanthera BH.}. Cucurb. (4). 6 S. Am. Elaterium [acq. Cucurbitaceae (4). 12 trop. Am. The fr. is explosive like that of Ecballium. Elaterium, Ecballium. Elatinaceae (EP., BH.). Dicots. (Archichl. Parietales; Guttiferales BH.). 2 gen., 30 sp. , trop. and temp. Undershrubs, herbs, or annual water-pi. ; the latter are able to live on land, altering their structure to suit the changed conditions (cf. Littorella). L. opp. or whorled, simple with interpet. stip. Fls. $ , reg., solitary or in dichasia, 2 — 6-merous. K hypog. , free or united ; C imbr. ; A in 2 whorls, or inner aborted; G syncarpous, multiloc., with simple style; plac. axile ; ov. oo , anatr. Capsule septifragal ; seed straight or curved; endosp. thin or none. Genera: Bergia, Elatine. Elatine Linn. Elatinaceae. 15 trop. and temp. E. hexandra DC. and E. Hydropiper L. (water pepper or pipe-wort) in Brit. Elatineae (BH.) = Elatinaceae. Elatinoides Wettst. (Linaria p.p .). Scroph. (n. 3). 24 Eur., As., Afr. Elatostema Forst. Urticaceae (2). 90 trop. As. E. acuminatitm Brongn. is apogamous. Some show water secretion from the 1. Elatostematoides C. B. Robinson. Urticaceae (2). 5 Phil. Is. Elattostachys Radlk. Sapindaceae (i). 10 Indomal., Polynesia. Elatus (Lat), tall. Elcismia Robinson (Celniisia p. p.). Compositae (4). 158. temp. Elcomarhiza Barb. Rodr. Asclepiadaceae (inc. sed.). i Brazil. Elder, Sambucus nigra L. ; American witch-, Fothergilla; dwarf- (W.I.), Pilta grandis Wedd. Elecampane, Inula Helenium L. Elegia L. Restionaceae. 20 S. Afr. Eleiotis DC. Leguminosae (in. 7). i India, Ceylon. Elemi, balsams, Burseraceae, Commiphora, Copaifera, Dipferocarpus, Litjiiidainkar, Pistacia, Styrax, Toluifera, etc. Eleocliaris R. Br. Cyperaceae (i). 90 cosmop. E. palustris R. Br. (spike-rush) on turfy moors in Brit. The green tissue is centric. The tubers of E. tuberosus Schult. (E. As.) are used as food. Eleogiton Link = Scirpus L. p.p. (Cyper.). E LODE A 239 Elephant-apple, Feronia Elephantum Correa; -'s ear, Begonia ; -'s foot (Am.), Ekphantopus. Elephantella Rydberg(P*a&V«/ar/jp.p.). Scrophul. (in. 3). i N. Am. Elepliantopus L. Compositae (i). 20 trop. E. scaber L. is an abundant and troublesome weed. Elephantorrhiza Benth. Leguminosae (i. 5). 2 trop. and S. Afr. Elettaria Maton. Zingiberaceae (i). i Indomal., E. Cardamomum Maton. Fls. on leafless shoots from the rhiz. Cult, in the mountains of Ceylon and S. India (cardamoms). The ripe fr. are picked and dried ; the seeds form a strongly flavoured spice, mainly used in India. Elettariopsis Baker. Zingiberaceae (2). 8 Malaya. Eleusine Gaertn. Gramineae (u). 10 trop. and subtrop. E. coracana Gaertn. (ragi, kurakkan) is cult, as a cereal in Ceylon, India, Africa, &c., and others are useful fodders. Eleutheranthera Poit. ex Bosc. Compositae (5). i trop. Am. Eleutheranthus (ELuthranthes) F. Muell. Rubi. (n. 7). i W. Austr. Eleutherine Herb. Iridaceae (n). 3 trop. Am., W.I. Eleutherococcus Maxim. (Acanthopanax p.p. EP.}. Aral. (i). 2 China. Eleutheropetalous, polypetalous. Eleutherospermum C. Koch (Plettrospermitm p.p. EP.). Umbel, (in. 4). 2 W. As. Eleutherostemon Herzog. Ericaceae (in. 2). i Bolivia. Eliaea Cambess. Guttilerae (n). i Madagascar. Elichrysum, cf. Helichrysum. Elionurus Humb. et Bonpl. ex Willd. Gramineae (2). 20 trop. and subtrop. Elisena Herb. Amaryllidaceae (i). 3 Peru. Elisma Buchen. Alismaceae. i Eur. (incl. Brit.), E. tiatans Buchen. Elissarrhena Miers (Anomospermum EP.). Menisperm. i Brazil. Elizabetha Schomb. Leguminosae (II. 3). 2 Brit. Guiana. Elk-horn fern, Platyceriiun. Elleanthus Presl. Orchidaceae (n. 7). 50 trop. Am., W. Ind. Ellertonia Wight. Apocynaceae (i. 3). 4 trop. As., Madagascar. Elliottia Muehlb. ex Nutt. Ericaceae (i. i). i S. All. U.S. Ellipanthus Hook. f. Connaraceae. 8 Indomal. Ellipeia Hook. f. et Thorns. Anonaceae (i). 10 Malaya. Elliptical, narrowish, tapering equally to both ends. Ellisia L. Hydrophyllaceae. 4 N. Am. Ellisiophyllum Maxim. Hydrophyllaceae. i Japan. Elm, Ulmus canipestris L. &c. ; Spanish- (W.I.) , Cordia, Hamelia. Elmera Ryclberg (Heuchera p.p.). Saxifragaceae (i). i N. Am. Elmeria Ridl. (Hornstedtia Retz.). Zingiberaceae (i). 2 Phil. Is. ElOdea Michx. Hydrocharitaceae. 6 Am. ; E. canadensis Michx. (American water- weed) arrived in Brit, about 1842 and rapidly spread over the inland waters of Eur. Only the ? plant is known in Eur., and the spreading is therefore due to veg. repr. chiefly by the breaking off of twigs. Submerged pi., slightly rooted, with whorls of 1., in whose axils are the usual squamulae. i fl. P 6, A 9; it breaks off as a bud and comes to the surface (cf. Vallisneria), where it opens. The ovary of the ? fl. grows to such a length_as to bring the fl. to the surface, where it is pollinated. P 6, stds. 3, G (3) ; enclosed below in 2 40 E LODE A a 2-leaved spathe. The pi. does not form a true winter bud; the 1. are merely a little more closely grouped together. Elongate, drawn out. Elsholtzia Willd. Labiatae (vi). 20 As., Eur., Abyss. Elutheria M. Roem. Meliaceae (n). 2 Peru to Venezuela. Elvasia DC. Ochnaceae. 5 N. Brazil, Guiana. Elvira Cass. Compositae (5). 3 trop. Am. Elymus L. Gramineae (12). 50 N. temp. S. Am. E. arenarius L. (lyme grass) on dunes in Brit. (cf. Ammophila); its 1. are coated with wax. Elyna Schrad. (Kobresia Willd. p.p.). Cyper. (in). 5 N. palaeotemp. Elynanthus Beauv. Cyperaceae (n). 30 %. Elynanthus Nees = Tetraria P. Beauv. (Cyper.). Elytranthe Blume (Loranthus p.p. BH.}. Loranth. (i). 50 Indomal. Elytraria Michx. (Tubiflora (imel.). Acanth. (i). 5 trop., subtrop. Elytropappus Cass. Compositae (4). 6 S. Afr. E. Rhinocerotis L. is a char. pi. of the karroo. Elytrophorus Beauv. Gramineae (10). 2 palaeotrop. Elytropus Muell -Arg. Apocynaceae (n. i). i Chili. Many bracts. Emarginate, notched. Embelia Burm. f. Myrsinaceae (il). 95 trop. and subtrop., exc. Am. Emblic myrobalan, Phyllanthus Emblica L. Emblingia F. Muell. Capparidaceae (iv). i W. Austr. Embolanthera Merrill. Hamamelidaceae. i Phil. Is. Embothrium Foist. Proteaceae (n). 5 Andes, Chili, E. Austr. Embryo, the young pi contained in the seed; adventitious-, Alchornea, Funkia, Nothoscordntu. Embryopteris Gaertn. = Diospyros L. (Eben.). Emeorhlza 1'ohl. Rubiaceae (n. 10). 2 S. Am. Emergences, surface outgrowths arising from other tissues as well as epidermis, Cactaceae, Diosera, A'osa, /fides. Emerus Tourn. ex Mill. = Corqnilla L. (Legum.). Emex Neck. Polygonaceae (i. 2). i Medit., S. Afr., Austr. The fr. is surrounded by the P, 3 of whose 1. are spiny. Emicocarpus K. Schum. et Schlechter. Asclep. (n. i). i S.E. Afr. Emilia Cass. (Settee io Tourn. p.p. EP.}. Compositae (8). 5 palaeo- trop. Emiliomarcelia Th. et H. Durand (Trichoscypha p.p.). Anacardiaceae (3). 4 Congo. Eminia Taub. (Rhynckosia p.p. ttH.}. Leguminosae (in. 10). 2 E. trop. Afr. Eminium Schott. Araceae (vn). 5 W. As. Emmenanthe Benth. Hydrophyllaceae. 6 N.W. N. Am. Emmenopteryx Oliv. Rubiaceae (i. 5). i China. Emmenosperma F. Muell. Rhamnaceae. 2 Austr. Emmeorrhiza Pohl. Rubiaceae (n. 10). 2 trop. S. Am. Einmer, Triticnm dicoccum Schrank. Emmotum Desv. Icacinaceae. 5 Brazil. Emorya Torr. Loganiaceae. i Texas. Empedoclea A. St. Hil. ( Tetracera p.p. EP.}. Dilleniaceae. i Brazil. Empetraceae (EP., fiff.). Dicots. (Archichl- Sapindales; anomalous ENDO- 241 Monochlamydeae Bff.). 3 gen., 4 sp., *. and Andes, occupying similar positions to Ericaceae; heath-like habit. The 1. are incurved backwards, forming a cavity on the under side partly filled up by hairs into which the stomata open. Infl. racemose, usu. dioec. In all but Corema the fls. are on 'short shoots' which arise lat. from the main axis and bear only scales below the infl. K 3, C 3, A 3, G (2 — 9). Loculi = cpls. ; ovules i in each, anatr. or nearly campylotr., erect on axile plac., with ventral raphe. Drupe with 2 — 9 stones; seed albuminous with no caruncle. Genera: Corema, Empetrum, Ceratiola. Empetrum (Tourn.) L. Empetraceae. i, E. nignun L. (crow-berry), on moors, N. temp. (incl. Brit.) and Andes. For habit, fl., &c. see fam. The flr. is dioec. and anemoph., but is sometimes $ and protandrous. Emplectanthus N.E. Br. Asclepiadaceae (u: 3). 2 S. Afr. Emplectocladus To IT. (Primus p.p. EP.}. Rosaceae (v). r N.W. Am. Empleuridium Sond. et Harv. Rutaceae (i). i S. Afr. Empleurum Ait. Rutaceae (i). i S. Afr. L. officinal (buchu). Empogona Hook. f. Rubiaceae (i. 8). 2 E. trop. and S. Afr. Enallagma Baill. Bignoniaceae (4). 4 C. Am., W.I. Enalus L. C. Rich. Hydrochar. i Indian and Pac. Oceans. See Enhalus. Enantia Oliv. Anonaceae (4). 3 W. Afr. Enantioblastae (Warming). The 4th cohort of Monocotyledons. Enantiophylla Coulter et Rose. Umbelliferae (in. 6). i Guatemala. Enantiostyly, Cassia (q-v.), Khugia, Leguminosae, Marantaceae, Saint f'aulia. Enargea Banks (Luzuriaga Ruiz et Pav.). Lili. (x). 3 S. Am., N.Z. Enarthrocarpus Labill. Cruciferae (2). 7 E. Medit., N. Afr. Encelia Adans. Compositae (5). 40 W. U.S. to Chili. Enceliopsis A. Nelson (Encelia p.p.). Compositae (5). 5 N. Am. Encephalartos Lehm. Cycadaceae. 5 Afr. The Kaffirs prepare a meal from the pith (cf. Cycas). Encephalosphaera Lindau. Acanthaceae (iv. B). i Colombia. Enchanter's Nightshade, Circaea. Encholirium Mart. (Dyckia BH.). Bromeliaceae (2). 6 Brazil. Enchosanthera King et Stapf (Anplectrum p.p.). Melastomaceae (i). i Indo-China. Enchylaena R. Br. Chenopodiaceae (A), i Austr. Enchysia Presl = Laurentia Neck. (Campan.). Enckea Kunth= Piper L. p.p. (Piper.). Encopa Griseb. Scrophulariaceae (n. 6). i Cuba. Encyclia Hook. = Epidendrum L. (Orchid.). Endacanthus Baill. Icacinaceae. i Madagascar. Endemic, confined to a small section of country, island, &c. Endiandra R. Br Lauraceae (n). 20 Indomal., Austr., Polynesia. Endive, Cichoriitm Endivia L. Endlicheria Nees (Ayendron BH. Aniba EP.}. Laur. (n). 3 trop. Am. Endo- (Gr. pref. ), within; -carp, the innermost layer of fr. in a drupe, &c.; -chylous, with internal water-storing tissue; -gamy, fusion of ? gametes; -genous, arising from internal tissues; -gens, Mono- cotyledons; -parasite, pi. living within the tissues of the host; w. 16 24 2 ENDO- -phytic, living within another; -pleura, inner seed coat, Cycadaceae; -rhizal, monocotyledonous; -sperm, the nutrient tissue outside 1-mbryo in sred, fleshy in Berbens, &c., bony in Phoenix, Phytelephas, horny in Coffea, &c., mealy in Frankeniaceae, ruminate in Anonaceae, and cf. Gymnospermae; -tropic, with hyphae entering the cells. Endoceilion Turcz. ex Herd. Compositae (inc. sed.). i Russia. Endodesmia Benth. Guttiferae (in), i W. trop. Afr. En.dolitn.odes Bartl. = Synisoocn Baill. (Rubiaceae). Endomallus Gagnep. Leguminosae (in. 10). i Cochin-China. Endonema A. Juss. Penaeaceae. 2 S. Afr. Endopogon Nees = Strobilanthes Blume p.p. (Acanth.). Endopogon Rafin. Rubiaceae (inc. sed.). i N. Am. Endorima Rafin. (Baldnina Nutt.). Cbmpositae (5). 3 N. Am. Endosiphon T. Anders, ex Benth. et Hook. f. Acanth. (iv A), i trop. Afr. Endospermum Benth. Euphorbiaceae (A. n. 6). 5 Malaya, China. Endosteira Turcz. Tiliaceae. i St Vincent. Endostemon N.E. Br. (Ocimuni p.p. ). Labiatae (vn). i S. and trop. Afr. Endressia J. Gay (Ligusticum p.p. BH.}. Umbellif. (ill. 5). i Pyrenees. Endusa Miers. Olacaceae. T. Peru. Enetophyton Nieuwland (Utricularia p.p.). Lentibular. i U.S. Engelhardtia Leschen. Juglandaceae. 10 Indomal., China. Engelmannia Torr. et Gray. Compositae (5). i S.W. U.S., Mex. Englerastrum Briquet. Labiatae (vn). 2 trop. Afr. Englerella Pierre. Sapotaceae (inc. sed.). i French Guiana. Engleria O. Hoffm. Compositae (3). 2 W. trop. Afr. Englerocharis Muschler. Cruciferae (4). 2 Peru. Englerodaphne Gilg. Thymelaeaceae. i E. Afr. Englerodendron Harms. Leguminosae (n. 3). i E. trop. Afr. Englerodoxa Horold. Ericaceae (in. 2). i Ecuador. Englerophytum Krause. .^apotaceae (i). r Cameroons. Englypha Chod. et Hassler. Aristolochiaceae. i Paraguay. Engrain, record produced on the organism. New Phyt. v. 200. Enhalus Rich. Hydr charitace^e. i In'omal., in salt water. The ? fls float horiz. at low water, and ca'ch the $ Us. which (cf. Vallis- neria) break off and float. As the tide rises, the ? fls. stand vertically, and the pollen, heavier than water, sinks down upon the stigmas. The testa bursts when the seed is ripe, and the embryo is freed. Enhydra {Enydraj Lour. Compositae (5). 9 trop. and subtrop. Enhydrias Ridl. Hydrocharitaceae. i Malaya. Enicosanthum Becc. Anonaceae (i). i Borneo. Enicostema Blume. Gentianaceae (i). i trop. Xero. and coast. Enkianthus (Enkyanthus) Lour. Ericaceae (n. i). 10 Himal. to Japan. Enkleia Griff. (Linostoma Wall.). Thymel. i Further India. Ennealophus N.E. Br. Iridaceae (n). i Amazon valley. Enneapogon Desv. ex Beauv.= Pappophorum Schreb. p.p. (Gram.). Enomegra Aven Nelson (Argemone p.p.). Papav. (n). 2 Rockies. Ensate, ensiform, sword -shaped. Enslenia Nutt. Asclepiadaceae (n. i). 3 N. U.S. to Colombia. EPA TIT IS 243 Entada Adans. (Pitsaetha L.). Leguminosae (l. 5). 12 trop. The seeds of E. scandens Benth. (nicker beans), a common trop. climber, are carried to Eur. by the Gulf Stream. Entandropliragma C. DC. Meliaceae (n). 6 Angola. Entelia R. Br. Tiliaceae. i N. Z. Wood very light. Enterolobium Mart. Leguminosae (i. i). 5 trop. Am., W.I. Enteropogon Nees. Gramineae (ti). 4 trop. Afr. and As." Enterosora Baker. Folypodiaceae. i Centr. Am., Jamaica. Enterospermum Hiern. Rubiaceae (i. 8). i E. Afr. coast. Enthomanthus Mo9- et Sesse ex Ramirez. Caprifol. 2 Mexico. Entire, without marginal notches. EntomopMly (fl.), insect-pollination; Mtiller has divided fl. into 8 fl.- classes (q.v.) according to their degrees of specialisation in regard to the visits of insects. Entoplocamia Stapf. Gramineae (n). 28. and trop. Afr. Entrecasteauxia Montr. Inc. sed. i New Caledonia. Enydra (Enhydrd) Lour. Compositae (5). 9 trop. and subtrop. Enzyme, an unorganised or soluble ferment. Eomecon Hance. Papaveraceae (n). 2 China. Epacridaceae (EP.\ Epacrideae BH.}. Dicotyledons (Sympet. Eri- ca\esEP.,BH.}. 21 gen., 350 sp., chiefly Austr. and Tasm., where they repres. the Ericaceae of other continents, but extending also to India, N.Z., Hawaii, and S. Am. They resemble Ericaceae (§ iv) closely in habit and appearance, being mostly small shrubs (a few small trees), the stems growing ± obliquely, sparingly branched, with narrow, entire, rigid 1., usu. alt. , rarely opp. or whorled. Sometimes the 1. are sheathing, in which cases either no scar is left on the stem when the 1. falls, or a ring mark is left, as in Dracophyllum. Fls. usu. in term, racemes or spikes. § , actinom. K 5, C (5), A 5, hypog. at edge of disc, or more often epipet. ; the anthers opening by one central longitudinal slit and without horns or other appendages; pollen simple or in tetrads; G (5) cpls. opp. petals, plac. axile; ov. in each loc. i — x , anatr., usu. pend.; style simple, sometimes in depression of top of ovary, with a capitate stigma. Capsule or stony drupe. Embryo straight, in copious endosp. Classification and chief genera (after Drude) : Each loc. with several seeds ; caps, loculic. 1. Prionoteae (sta. hypog., free; anther-loc. divided): Prio- notes, Lebetanthus (only gen.). 2. Epacrideae (sta. usu. epipet.; anther-loc. without separation after dehiscence): Sprengelia, Richea, Dracophyllum, Sphenotoma, Epacris. Each loc. with i seed: fr. indehiscent. 3. Styphelieae: Styphelia, Acrotriche, Pentachondra. Epacrideae (BH.} — Epacridaceae. Epacris Forst. Epacridaceae (2). 40 S.E. Austr., Tasm.. N.Z., New Caled. Cult. orn. fls. Epactium Willd. Inc. sed. i Peru. Epallage DC. Compositae (,s). 5 Madagascar. Epaltes Cass. Compositae (4). 12 trop. Epatitis Rafin. Compositae (inc. sed.). i Oregon. 16 — 2 244 EPERUA Eperua Aubl. Leguminosae (n. 3). 6 N. Braz., Guiana. E.purpurea Benth., a showy tree of the caatingas. E. fakata Aubl. (wallaba, Br. Guiana) yields a good timber. Ephebepogon Nees et Meyen (Pollinia p.p. EP.). Gram. (2). i China. Ephedra Tourn. ex L. Gnetaceae. 25 warm temp. Shrubs, much branched, with opp. connate 1. reduced to scales, so that the stem performs the work of assim. Fls. diclinous, with no trace of cpls. in chidia, Filices, e.g. Asplenium, Platycerium, Polypodium, &c., Ficus, Hydnophytitm, Marcgravia, Myrmecodia, Oncidin//i, Orchidaceae, Phalaenopsis, Phyllocaclus, Piper, Rhipsalis, Rhododendron, Scitticaria, Tillandsia, Vanilla, &c. See Schimper, Die epiph. Vegetation Amerikas, Jena, 1888, and Plant Geography; Goebel, Pfianzenbiol. Schilderungen. Epipogum S. G. Gmel. Orchidaceae (11." 2). i Eur. (incl. Brit.), As., E. aphvllnin Sw. a leafless saprophyte with branched rhiz. and no r. ; endotropic mycorhiza. Fl. as in Epipactis, but without twisting of the recept. Epipremnopsis Engl. Araceae (i). i Indomal. Epipremnum Schott. Araceae (n). 15 Indomal. Epiprinus Griff. Euphorbiaceae (A. n. 2). i Further India. EpirrMzantlies Blume = Salomonia Lour. p.p. (Polygal.). Episcia Mart. Gesneriaceae (i). 30 trop. Am., W.I. EpistepMum Kunth. Orchidaceae (n. »). 7 trop. S. Am. Epistylium Sw. = Phyllanthus L. p.p. (Euph.). Epitaberna K. Schum. Rubiaceae (i. 8). Cameroons. Epithema Blume. Gesneriaceae (i). to Indomal., trop. Afr. Epithymum Lunell (Citscuta p.p.). Convolv. (n). 5 W. U.S. Epitrachys C. Koch = Cnicus L. (5Ar.) = Cirsium Tourn. (Comp.). Equisetaceae. Pteridophyta (Equisetineae). An order with one sur- viving genus (Equisetum q.v.}, formerly well repres. Many large fossils. Equisetales (Eqnisdineae}. A main division of Pteridophyta (e/.v.), containing the single living fain. Equisetaceae. Equisetum L. Equisetaceae (only genus). 25 cosmop. ; 9 in Brit. (horsetails), chiefly in swampy places. Perenn. herbs with symppdial rhiz. which send up aerial shoots each year, of one or two kinds; in some sp. the ordinary green shoot bears the repr. spike at the end, while in the others there is a special rt-pr. shoot, usu. appearing early in the year, and often without chlorophyll, the ordinary shoots per- forming assim. work only. The stem is very distinctly jointed, and at the nodes are borne whorls of united 1. closely pressed against the stem, and of little or no use in assim. The branches emerge through the leaf-sheath and thus appear at first sight endogenous; in reality they are exogenous, but formed so much later than the 1. that their points of origin are already covered by the leaf-sheath, and so they are compelled to burrow through it. Stages in this process may easily be observed. The surface of the stem is grooved ; the ridges are occupied by mechanical tissue, whilst the green tissue and stomata are at the base of the furrows. This is a marked xero. structure and is repeated very closely in Casuarina. In several sp. the internodes EREMOLEPIS 247 of the rhiz. are swollen into tubers, which serve for hibernation and veg. repr. The spike is very like the 3 fl. of a Conifer, and has as much right to the title of flower. It is an axis with short internodes, bearing a dense mass of sporophylls. Each is shield-shaped and bears a number of sporangia upon the under side of the head (i.e. towards the stem), arranged like the horses of a 'merry -go-round.' The spores are of one kind only; each has, running round it, two spiral cuticularised bands of membrane, formed from the outer wall and termed elaters. These are hygroscopic, unfolding in damp air. In the rolling up again on drying, the elaters of one spore become entangled with those of others and cause them to adhere together, so that several prothalli may be formed near to one another when they germinate. This is advantageous, for the prothalli are dioec., though so far as we can tell the spores are all alike. The prothallus is fairly large, the 3 being smaller than the ? . The stems of E. hyemale L. (Dutch rushes) are used for polishing, the mechanical tissues contain much silica, as do those of most. Equitant (vernation), cf. Vernation. Eragrostis Host. Gramineae (10). 150 cosmop., mostly subtrop. Eranthemum L. (excl. Psenderanthemum Radlkt.). Acanthaceae (iv. A). 25 trop. As. Eranthis Salisb. Ranunculaceae (2). 7 N. palaeotemp. E. hyemalis Salisb. (winter aconite, nat. in Brit.) has a thick rhizome or row of tubers, one formed each year. The sol. term. fls. appear in February, before thel.; each has an invol. of three green 1., a 'calyx' of 6 seg- ments, and several honey-1. or petals. Brasilia R. Bruniaceae. Nomen. Ercilla A. Juss. Fhytolaccaceae. 2 Chili, Peru. E. voiubilis A. luss., climbs by adhesive discs, endogenous just above the axils. Erechtites Rafin. Compositae (8). 15 Am., Austr., N.Z. Erect (ovule), erect with stalk at base (fl. held erect). Eremaea Lindl. Myrtaceae (n. 2). 6 W. Austr. Eremaeopsis O. Ktze. = Eremaea Lindl. (Myrt). Eremalche Greene (Malvastrum p.p.). Malvaceae (2). 3 W. U.S. Eremantbus Less. Compositae (i). 20 Brazil. Eremia D. Don. Ericaceae (iv. 2). 30 S. Afr. Eremiastrum A. Gray. Compositae (3). 2 Calif., Arizona. Eremiopsis N.E. Br. Ericaceae (iv. 2). i S. Afr. Eremium (Cl.), a desert formation. Eremobium Boiss. (Alalcolmia BH.). Cruciferae (4). 3 N. Afr. Eremocarpus Benth. Euphorbiaceae (A. n. i). 2 Pac. N. Am. Eremocarya Greene. Boraginaceae (iv. 2). 3 Pac. N. Am. Eremocrilaena Baill. Chlaenaceae. i S. Madagascar. Eremochloa Buese. Gramineae (2). 6 SE. As. Eremochloe S. Wats. Gramineae (ro). 2 W. N.Am. Eremocitrus Swingle ( I'ripkasia p.p.) Rutaceae (v). i N. Austr. Eremocrinum M. S. Jones. Liliaceae (in), i California. Erernogxtfie Fenzl. = Arenaria L. p.p. (Carophyll.). Eremolaena Baill., cf. Eremochlaena. Eremolepis Griseb. Lonmthaceae (n). 5 trop. A., W.I. 248 EREMOLITHIA Eremolithia Jepson. Caryophyllaceae (i. 6). i California. Eremoluma Baill. (Lucuma p.p. EP.}. Sapotaceae (i. 6). i Guiana. Eremomastax Lindau (PaitloivillidmiaBH.}. Acanth. (iv. A), i W. Afr. Eremopanax Baill. Araliaceae (i). 3 New Caledonia. EremopMla K. Br. (Pholidia EP.}. Myoporaceae. 30 Austr. Eremophyton Beguinot. Cruciferae (2). i N. Afr. Eremosis Gleason ( Vernonia p.p.). Compositae (i). 15 Mex., C. Am. Eremosparton Fisch. et Mey. Leguminosae (ill. 6). i W. and C. As. Eremospatha G. Mann et H. Wendl. Palmaceae (ill. 2). 5 trop. Afr. Eremostachys Bunge. Lahiatae (vi). 40 mid. and W. As. Eremosyne Endl. Saxifragaceae (i). i S.W. Austr. Eremothamnus O. Hoffm. Compositae (8). i S. Afr. Eremurus Bieh. Liliaceae (ill). 20 alpine W. and Cent. As. Fl. protog. ; the petals crumple up before the essential organs are ripe. The 1. of E. aitrantiactis Baker are eaten in Afghanistan. Eria Lindl. Orchidaceae (n. 15). 240 trop. As. Epiphytes; cult. Eriachaenium Sch.-Bip. Compositae (9). i Tierra del Fuego. Eriachne R. Br. Gramineae (y). 25 trop. As., Austr. Eriadenia Miers. Apocynaceae (n. i). i trop. S. Am. Eriander H. Winkler. Rutaceae (iv). i trop. Afr. Eriandrostachys Baill. Sapindaceae (i). i Madagascar. Eriantlius Michx. Gramineae (2). 25 trop. E. Ravennae Beauv. orn. Eriastrum Wooton et Standley (Giiia p.p.)- Polemon. 2 New Mexico. Eriaxis Rchh. f. (Galeola BH.}. Orchidaceae (n. 2). i New Caledonia. Eribroma Pierre (Slerculia p.p. EP.}. Sterculiaceae. i trop. Afr. Erica (Tourn.) L. Ericaceae (iv). 500 Eur. (esp. Medit.) and S. Afr. (see fam.). 5 in Brit.; the two common heaths, E. cinerea L. and E. Tetralix L., cover great areas of moor. In habit like Calluna. Fl. bell-shaped and pendulous, visited and fert. mainly by bees. Honey is secreted by the disc, and insects hanging on to the fl. and probing for it must shake the sta. and receive a shower of the loose powdery pollen from the pores in the tips of the anthers. In the wider mouthed sp. the anthers have horn-like projections at the hack, which ensure contact with the insect's proboscis. The stigma pro- jects beyond the sta. so as to be touched first. Many S. Afr. sp. cult. E. scoparia L. is the heath of S. France, &c. (bruyere), several feet high; its rootstocks furnish 'briar' wood pipes. The roots of heaths possess endotropic mycorhiza. Ericaceae ( EP., BH. excl. Vacciniaceae, incl. Pyrolaceae p.p., CUthraceae. Dicots. (Sympet. Ericales EP., BH.}. 50 gen., 1 350 sp. Owing to their numbers and their social habit they form very char, parts of the veg. in many portions of the globe. Cosmop. except in deserts and in hot damp trop. regions. The Ericoideae are confined to Africa, Medit. and Europe, the two great masses of them being however separated by the Sahara, though sp. of Erica, &c. occur in each (see Drude in Nat. Pfl. or Pflanzengeog. for full discussion of the ERICACEAE 249 interesting distr. of E. See also gen., esp. Rhododendron, Vaccinium, Calluna, Erica). The distr. of E. on arctic moors and swamps and on dry moors in warmer regions (almost all grow in peaty soil) would lead us to expect, and we find, esp. in Ericoideae, xero. char. Woody plants from small undershrubs to large shrubs or even trees. Two types of habit — that of Ericoideae and that of the remaining tribes. In the latter there are usu. true winter-buds formed (e.g. Rhododendron), even though the 1. may last over the winter. The bud is covered with scale-]., and when its elongation occurs these drop off and a gap is left on the stem; the foliage 1. tend to form rosettes at the ends of the twigs. The 1. are generally elliptical, entire or nearly so, and leathery, frequently hairy. The upper epidermis is stoutly cuticu- larised, and there is often water storage tissue between it and the green tissue. In the Ericoideae there are no true winter-buds or scale-1. ; the plants are evergr., and the whorled 1. needle-like, often through being rolled back on themselves to form a groove or even a chamber on the under side (cf. Empetrum). The inrl. commonly terminates a strongly growing shoot, and a sympodial growth tends to be formed. The fls. may be sol., but are more often in racemose groupings, each with a bract and two bracteoles, g , actinom. or slightly zygom. K 4 — 5 ; C (4 — 5) or 4 — 5 (Ledeae}, usu. bell-shaped; A 8 — 10, obdipl., hypog. or rarely slightly epipet.; anthers intr., often with projecting appendages, the thecae often spreading at top, and opening by apical pores; pollen grains in tetrads. Below the gynoeceum is a fleshy disc secreting honey; G (4 — 5) sup. or inf., 4 — 5-loc., with axile plac. ; ov. in each loc. I — oo, anatr. ; style simple with capitate stigma. Capsule, drupe or berry. Embryo cylindrical, in copious endosp. The fls. of the Brit. sp. are mostly bee-rls. with 'loose-pollen' mechanism. The hanging position and the size are suited to bees. The stigma projects so as to be first touched, and in probing for the honey at the base of the fl the bee touches the sta. or their projecting horns, and by thus shaking them causes a shower of pollen to fall from the tips of the anthers. Calluna is partly anemoph. ; Kalmia has a curious explosive mechanism (see C., K., and Erica, &c.). [BH. chars. Shrubs and trees, 1. alt., opp., or whorled, exstip , usu. persistent. Infl. various. Fls. reg. $ . K (4 — 5) or 4 — 5; C usu. (4—5), A usu. 8 — 10, sometimes adnate to base of C., usu. dehisc. by pores; disc various or o ; G (i — 12), multi-loc., with style and peltate, capitate or lobed stigma; ov. usu. oo in each loc., anatr., on axile plac. Caps., rarely drupe or berry, with small seeds. Endosp.] Classification and chief genera (after Drude). I. RHODODENDROIDEAE (septicidal caps.; seed with ribbed loose coat, often winged; C falling after flowering ; sta. with upright or long adnate anthers, with no appendages): i. Ledeae (polypet. ) : Leclum. i. Rhododendreae (zygomorphic) : Rhododendron, Menzie- sia. 3. Phyllodoceae (actinomorphic) : Loiseleuria, Kalmia, Phyl- lodoce, Daboecia. 250 ERICACEAE II. ARBUTOIDEAE (berry or loculic. caps.; seed triangular or ovate, not winged; C falling; anthers much folded, with peg-like appendages, or prolonged into tubes, shedding the pollen upwards; ov. sup. ): 1 . Androinedeae (dry capsule with small calyx at base) : Cas- siope, Andromeda, Epigaea. 2. Gaultherieae (caps, or berry; K fleshy round caps., or leafy; anthers blunt at tip or with two short processes): Gaultheria, Pernettya. 3. Arbuteae (K as small disc at base of berry; anthers with two long processes) : Arbutus, Arctostaphylos. III. VACCINIOIDEAE (as II., but ov. inf.): i. Vatcinieae (ovary sharply denned from peduncle) : Gay- lussacia, Vaccinium. i. Thibaudieae (K decurrent on ovary and going over into peduncle): Pentapterygium, Agapetes, Paphia, Macleania, Thibaudia. IV. ERICOIDEAE(h. usu. loculic. caps , or nut ; seeds round, not winged ; C persistent after flowering ; anther with short connective, thecae spreading above, frequently appendaged): 1. Ericeae ( > i seed in each loc.) : Calluna, Erica. 2. Salaxideae (i seed in each loc., capsule or nut) : Eremia, Salaxis. Ericaceous, heath-like. Ericala Renealm. ex S. F. Gray = Gentiana Tourn. (Gent.). Ericales. The ist order of Sympetalae. The 4th cohort (BH.) of Gamopetalae. Ericameria Nutt. (Aplopappus BH.}. Compositae (3). 12 W. U.S. Ericetal, growing on moors. Erichsema Hemsl. Leguminosae (in. 2). i Austr. Ericinella Klotzsch. Ericaceae (iv. i). 4 S. and W. Afr. Ericoid (1.), narrow, nee 'le-like, ± rolled back. Erigenia Nutt. Umbelliferae (m. 4). i E. U.S. Erigeron L. Compositae (3). 150 cosmop., esp. N. Am.; 2 Brit, (flea-bane). Erinacea (Tourn.) Adans. Leguminosae (in. 3). i S.W. Eur. Erinna Phil. Liliaceae (iv). i Chili. Erinocarpus Nimmo ex J. Grah. Tiliaceae. i S. India. Erinus L. Scrophulariaceae (in. i). i Pyrenees, Alps. Erio- (Gr. pref.), woolly ; -phorous, very cottony, or woolly. Eriobotrya Lindl. Rosaceae (n). 12 subtrop. As. E. Japonica Lindl. has ed. fr. (loquat), largely cult, in As. Eriocaucanthus Chiov. Malpighiaceae (i). 3 trop. and S. Afr. Eriocaulaceae (EP., BH.). Monocots. (Farinosae, Glumaceae BH.). 6 gen., 540 sp. mostly trop. and subtrop. Perenn. herbs with grass- like 1. Fls. in heads (involucrate), inconspic., unisexual, 2- or 3- mcrous, reg. or -I- . P usu. sepaloid, in 2 whorls. .,5Ar. \ncl.Suxaceae}. Dicots.(Archichl.Geraniales; Unisexuales BH. ). 220 gen., 4000 sp., cosmop., except arctic. Few sp. have a very wide range ; the most widely-ranging genus is Euphorbia. Closely related to Geraniales by ihe structure of the gynoeceum, &c., although separated a good deal from the other fams. of the order by the amount of reduction in most of its fls. Diagram of central cya- thium of infl. of Euphorbia Peplus L. (after Eichler, modified). W. 258 E UPHORBIA CEAE Most are shrubs or trees, a few herbaceous (e.g. the Brit. sp.). Many are xero. ; a number of Australian sp. are of ericoid habit ; several, esp. S. Afr., Euphorbias are cactus-like; others resemble Lauraceae, or possess phylloclades (e.g. Phyllanthus sp.). A few are lianes. L. usu. alt.; some have opp. 1., some opp. 1. above and alt. below. Slips, usu. present, but may be repres. by branched hair- like bodies (Jatropha), glands, or thorns. Nearly all contain latex in special laticiferous cells. Infl. usu. complex ; almost every type occurs. Often the first branching is racemose and all subsequent ones cymose. In some cases, e.g. Dalechampia and Euphorbia (q-v-), the partial infls. are so condensed as to give the appearance of single fls. The fls. are always unisexual, monoec. or dioec. , reg., hypog. The P may be present as two whorls usu. j-merous; more often there is only one (calyx) and .frequently the fl. is naked. Sta. i — oo , free or united in various ways. Ricinus has branched sta. Phyllanthus cydanthera has the sta. united, with a ring-like common anther. G usu. (3), with axile placentae, and 3 loc. Styles usu. 2-lobed. The ovules are constant throughout the family and form its best distinctive feature; they are i or 2 in each loc., collateral, pendulous, ana- tropous, with ventral raphe. The micropyle is usu. covered by a caruncle, which is also found on the seed. The fruit is almost invariably a ' schizocarp-capsule. ' It splits into cpls. often elastically, and at the same time each cpl. opens ventrally, letting the seed escape. See albuminous. [BH. chars, incl. those of Buxaceae, esp. dorsal raphe, and loculic. caps, or drupe.] Most E. are poisonous. Several are important economic plants, e.g. Manihot (rubber, cassava), Hevea (rubber), Croton, Ricinus, &c. Classification and chief genera (after Pax) : A. PLATYLOBEAE (cotyledons much broader than radicle) : I. PHYLLANTHOIDEAE (ovules 2 per loc. ; no latex) : i. Phyllantheae (embryo large, little shorter than endosp. ; $ calyx imbricate) : Phyllanthus. i. Briddieae (do., but 3 calyx valvate): Bridelia. 3. Daphniphylleae (embryo short, 4 — 6 times shorter than en- dosp.) : Daphniphyllum. II. CKOTONOIDEAE (ovules i per loc. ; latex usu. present): 1. Crotoneae (sta. bent inwards in bud) : Croton. 2. Acalypheae (sta. erect in bud; fl. usu. apetalous; c? calyx valvate; infl. a raceme, spike, or panicle, axillary or term.): Mercurialis, Acalypha, Ricinus, Dalechampia, Tragia. 3. Jatropheae (do. ; infl. a dichasial panicle): Hevea, Jatropha. 4. Adnaneae (do.; infl. a simple term, spike or raceme): Manihot. 5. Cluytieae (s calyx imbr. ; i fls. with petals, in groups or cymes, these partial infls. axillary or in complex infls.): Codiaeum, Cluytia. 6. Gtlonieae (do. but apetalous) : Gelonium. 7. Hippomaneae (do.; apetalous; infl. axillary or term., spike- like, the partial infl. cymes) : Stillingia, Hura, Hippo- mane. E UR YSPERM UM 259 8. Eitphorbieae (cyathium) : Anthostema, Euphorbia. B. STENOLOBEAE (cotyledons as wide as radicle): I. PORANTHER01DEAE (ovules 2 per loc.) : Poranthera. II. KICINOCARPOIDEAE (ovules i per loc.): Ricinocarpus. Euphorbiodendron Millspaugh (Euphorbia p.p.). Euph. (A. n. 8). 1 2 trop. Am., W I. Euphorbiopsis Leveille (Euphorbia p.p.). Euph. (A. n. 8). i S.E. As. Euphoria Comm. ex Juss. Sapindaceae (i). 6 trop. and subtrop. As. Eupnoriantlms Radlk. Sapindaceae (i). i Malay Archipelago. Euphoriopsis Radlk. Sapindaceae (i). i Indomal. Euphrasia L. Scrophulariaceae (in. 3). 100 sp. extra-trop. E.offi- cinalis L. (eyebright) in Brit. Semi-parasites with loose-pollen fls. (see fam.). The 4 anthers lie close under the upper lip of the fl. ; the two upper cohere and also the upper to the lower on each side; the lower lobe of each has a projecting spine. Insects probing for honey shake the spines and receive a shower of pollen from among the anthers. The stigma protrudes beyond the sta. in most fls. so as to be touched first, but every stage can be found from highly protog. fls. with very protruding stigmas to almost homog. fls. whose stigma does not protrude and with self-fert. Euphrona Veil. Inc. sed. i Brazil. Euphronia Mart. Rosaceae (i. 2). i Brazil. Euphrosine Allem. Inc. sed. Nomen. Euphrosyne DC. Compositae (5). i Mexico. Euplassa Salisb. Proteaceae (il). 8 trop. Am. Eupomatia R. Br. Eupomatiaceae. 2 N.E. Austr., New Guinea. Fl. deeply perig., naked ; A oo , G oo . Oil cells. Eupomatiaceae (EP. Anonafeaep.p. BH.). Dicots. (Archichl. Ranales). Only genus Eupomatia, q.v. Euptelea Sieb. et Zucc. Trochodendraceae. 3 Japan to Bengal. Eureiandra Hook. f. Cucurbitaceae (3). 3 trop. Afr. EuroscMnus Hook. f. Anacardiaceae (3). 5 New Caled., E. Austr. Eurotia Adans. Chenopodiaceae (A). 2 Meclit., W. As., N. Am. Eurya Thunb. Theaceae. 4o Mexico, S. Am., W. and E. Ind. Euryale Salisb. Nymphaeaceae (in), i S.E. As. Fl. epigynous. The seeds and roots are eaten in China. Eurybia Cass. = Olearia Moench. (Comp.). Eurybropsis DC. =Vittadinia A. Rich. (Comp.). Eurycentrum Schlechter (Cystorchis p.p.). Orch. (n. 2). 2 N. G., Solomons. Eurycles Salisb. Amaryllidaceae (i). 2 N. Austr., Malaya. Eurycoma Jack. Simarubaceae. 3 S.E. As. Eurygania Klotzsch (Thibaudia p.p. EP.}. Eric. (m. 2). 8 Andes. Eurylepis D. Don = Erica Tourn. p.p. (Eric.). Eurylobium Hochst. Verbenaceae (2). i S. Afr. Euryloma D. Don = Erica Tourn. p.p. (Eric.). Euryops Cass. Compositae (8). 35 S. Afr. to Socotra. Eurypetalum Harms. Leguminosae (n. 2) i Guinea. Euryptera Nutt. (Peucedanum p p.). Umbellif. (in. 6). 6 N. Am. Eurysolen Prain Labiatae (in), i Indomal. Euryspennum Salisb. = Leucadendron R. Br. (Prot). 17 — 2 260 EURYSTEGIA Eurystegia D. Don = Erica Tourn. p.p. (Eric.). Eurystyles Wawra (? = Stenoptera EP.). Zingib. or Orchid. I Brazil. Eurytaenia Torr. et Gray. Umbelliferae (in. 5). i Texas. Euscapbis Sieb. et Zucc. Staphyleaceae. i Japan. Easideroxylon Teijsm. et Binn. Lauraceae (i). i Borneo. Eusporangiatae. .see Filicales. Eustegia Rafin. Melastomaceae (inc. sed.). 3, habitat? Eustegia R. Br. Asclepiadaceae (n. i). 5 S. Afr. Eustephia (."av. Amaryllidaceae (i). 2 Peru, Argentina. EustepMopsis R. Fries. Amaryllidaceae (i). 2 Argentina. EustigmaGardn. et Champ. Hamamelidaceae. 2 Hongkong, Tonquin. Eustoma Salisb. Gentianaceae (I. i). i Centr. Am., W.I. Eustrephus R. Br. Liliaceae (x). i E. Austr. Eutacta Link = Araucaria Juss. p.p. (Conif.). Eutaxia R. Br. Leguminosae (in. 2). 8 Austr. Euterpe Gaertn. Palmae (iv. i). 10 trop. Am. E. edit/is Mart. (Assai palm) ed. fr.; a beverage is prepared by soaking it in water. Eutetras A. Gray. Compositae (6). i Mexico. Euthamia Kll. (Solidago p.p.). Compositae (3). 10 N. Am. Euthemis Jack. Ochnaceae. 5 Indomal. Euthystachys A. DC. Verbenaceae (2). i S. Afr. Eutoca R. Br. = Phacelia Juss. p.p. (Hydrophyll. |. Eutrema R. Br. Cruciferae (2). 4 Siberia. Eutriana Trin. = Bouteloua Lag. (Gramin.). Euxena Calest. (Arabis p.p.). Cruciferae (4). t France. Euxolus Rafin. — Amarantus L. (Amarant). Euxylophora Huber. Rutaceae (i). i Amazon valley. Good wood. Euzomodendron Coss. Cruciferae (4). i S. Spain. Evacidium Pomel (Fitngop.p. EP.). Compositae (4). i N. W. Afr. Evandra R. Br. Cyperaceae (n). 2 S.W. Austr. Evansia Salisb. = Iris Tourn. (Irid.). Evax Gaertn. Compositae (4). 15 Medit., N. Am. Evelyna Poepp. et Endl. = Elleanthus Presl (Orchid.). Evening primrose, Oenotliera. Everardia Ridl. Cyperaceae (n). i Roraima (Brit. Guiana). Everettia Merrill. Melastomaceae (n). i Phil. Is. Everettiodendron Merrill. Euphorbiaceae (A. i. i). i Phil. Is. Evergreen, bearing green 1. throughout the year. Everlasting, Achyrackaena, Ammobium, Anfi'niiiiria, Gnaphalhun, Helichrysum, Helipterum ; -grass (Am.), Eriochloa ; mountain-, Antfniiana; -pea (Am.), Lathyrus. Eversmannia Bunge. Leguminosae (in. 7). i Persia. Evodia Forst. Rutaceae (i). 45 trop., exc. Am. Evodiauthus Oerst. (Carludovica p:p. BH.\ Cyclanthaceae. 2 Costa Rica, W.I. Evolution, the development of new forms from old. Evolvulus L. Convolvulaceae (i). 90 trop. and subtrop. Evonymus (Euonymus tj.v.) L. Celastraceae. 60 N. temp, and S.E. As. Evota Rolfe. Orchidaceae (n. i). 3 S.W. Cape Colony. Ewartia Beauverd. Compositae (4). 3 S.E. Austr., Tasmania. FAB A 261 Ex- (Lat. pref.), not, without, outside, e.g. -albuminous, without albumen ; -centric ; -eluded, exserted ; -current, with single straight main stem, Finns ; -Ogamy, tendency of allied gametes to avoid pairing ; -ogenous, arising from the external layers ; -Ogens, Coniferae and Dicotyledons ; -otropism, tendency of lat. r. to grow away from main r. ; -serted, protruding ; -stipulate, &c. Exacum L. Gentianaceae (i). 30 sp. palaeotrop. The style is bent to one side or other of the fl. ; both occur on the same plant (enantio- styly, ij.v.). Exarrhena R. Br. = Myosotis L. p.p. (Borag.). Excelsus (Lat.), lofty. Excoecaria L. Euphorbiaceae (A. n. 7). 30 sp. trop., exc. Am. For E. sebifera Muell.-Arg. see Sapium. Excoecariopsis Pax. Euphorbiaceae (A. n. 7). i S.W. Afr. Excremis Willd. Liliaceae (in), i Andes. Exechostilus K. Schum. Rubiaceae (n. i). i trop. Afr. Exfoliate, to peel off. Exoacantha Labill. Umbelliferae (in. 8). i Syria. Exocarpus Labill. Santalaceae. 15 Austr., Malaya, Madag. Timber. Exocarya Benth. Cyperaceae (n). i S.E. Austr. Exochaenium Griseb. in DC. (Belmontia E. Mey.). Gent. (i). 3 trop. Afr. Exochogyne C. B. Clarke. Cyperaceae (n). i Amazon valley. Exochorda Lindl. Rosaceae (i. i). 3 Centr. As. Cult. orn. shrubs. Exogonium Choisy (Ifomoea p.p. BH.). Convolvulaceae (i). 18 trop. Am. E. Purga Benth. (jalap) cult, for medic, resin. Exolobus Fourn. Asclepiadaceae (n. 4). 4 Brazil. Exomicrum Van Tiegh. = Ouratea Aubl. (Ochn.). Exomis Fenzl. Chenopodiaceae (A), i S. Afr., St Helena. ExorrMza Becc. (Kentia p.p. EP.). Palmaceae (iv. i). i Fiji. Exostemma Rich. Rubiaceae (i. 5). 20 W.I.. S. Am. Febrifugal alkaloids are contained in the bark. Exostyles Schott. Legnminosae (n. o)- 2 Brazil. Exotanthera Turcz. (Rinorea EP.}. Violaceae. i Madagascar. Exothea Macfadyen. Sapindaceae (n). 2 W.I. , C. Am. Exploding anthers, Broussonetia, Pilea, Urlica ; -pollen, &c., Cala- aenia, Compositae (Cynareae), Corytlalis, Cytisns. Genista, A'ahnia, Medicago, Posoqiteria, Pterostylis, Ulex, &c. ; -fruit, cf. Dispersal. Extra- (Lat. pref.), beyond; -axillary, outside the axil; -tropical, outside the tropics ; -floral nectaries, cf. Nectaries. Extrorse, opening away from centre of fl. Eye, a bud, Helianthus, Solatium ; -bright, Euphrasia officinalis L. Eylesia Sp. Moore. Scrophulariaceae (ill. 2). i trop. Afr. Eyrythalia Renealm. =Gentiana Tourn. (Gent.). Eysenhardtia H. B. et K. Leguminosae (in. 6). 5 Texas to Guate- mala. Eystathes Lour. Inc. sed. i Cochin-China. F (fl. -class), fls. suited to Lepidoptera, Angraeciini, Cuphea, Dianthus, Labiatae, Lonicera, Lychnis, Oenothera, Primula, and cf. Butterfly and Moth fls. Faba (Tourn.) L. = Vicia L. (Legum.). 262 FABERIA Faberia Hemsl. Compositae (13). i China. Fabiana Ruiz et Pav. Solanaceae (4). 15 S. S.Am. Cult. orn. shrubs. Fabricia Gaertn. = Leptospermum Forst. (Myrt.). Fabricia Scop. (Afysicarfus F.P.). Leguminosae (in. 7). 16 palaeo- trop. Facelis Cass. Compositae (4). Fades, external appearance. Facultative, incidental. Fadogla Schweinf. Rubiaceae (n. Fadyenia Hook. Polypodiaceae. 3 temp. S. Am. W. Ind. The i). 20 Afr. i, F. prolifera Hook. sterile 1. produce buds at the tips. Fagaceae (EP.; Citpuliferae p.p. />'//.). Dicots. (Archichl. Fagales). 5 gen., 350 sp. ; three chief centres of distr. — Fagus, Castanea § Eucastanea and Quercus N. extra-trop., Pasania and Castanopsis in trop. As. and Calif., Fagus § Nothofagus in S. Am., N.Z., and Floral diagrams of Castanea vulgaris, after Eichler. A, diagram of $ cyme in axil of catkin-leaf, the sta. and rudimentary gynoeceum only shown in the first fl. The sequence of the fls. is indicated by the figures i, 2, 3. B, diagram of ? partial infl. b = bract, aS = bractejles, a' (3' a, j3, = bracteoles of second order. S. Austr. Most are trees with simple 1. and scaly stipules that drop off as the 1. expand. The fls. come out in the axils of the 1. of the current year and are diclinous and anemoph., arranged in catkins or small spikes (exc. Fagus i ). In general there is a close resemblance to Betulaceae, and, as in that fain., the fls. are usu. in dich. cymes in the axils of the catkin-1.; there are often, however, > three fls. P bract-like, (4 — 7). i fl. with as many to twice as many or oo sta. undivided, with or without rudimentary style. _ ? fls. usu. in dich. of 3 in Castanea, i in Fagus, i in Quercus, &c. G usu. (3) with 3 styles (exc. sp. of Castanea) ; loculi 3, usu. visible before fert. Plac. axile, each bearing i pend. anatr. ov. with i integuments. Fr. a i -seeded nut. Seeds without endosp. The group of nuts is surrounded by a cup-like organ termed a cupule ; in the oak there is one nut in each cupule, in the beech two, in the chestnut three. About the morphology of this organ there has been much discussion. Eichler (see diagram above, fig. B, and Biii- thendiagr.) regards it as the combined bracteoles a' j8' a, /3, , Prantl (Engler's Bot. Jahrb. vm. 1887) as an axial outgrowth. See also Celukov>ky in Pringskeim's Jahrb. xxi. 1890, and cf. Betulaceae. The cupule only becomes clearly visible after fert. FALSE ACACIA 263 Some of the F. show signs of peculiar development of the embryo- sac, and other interesting features (see Chalazogamae). The order includes several important economic plants, chiefly valuable for their timber, e.g. oak (Quercus), beech (Fagus), chestnut (Castanea), &c. Classification and genera (after Prantl) : 1. Fageae (fls. in dich., rarely sol. in axils of foliage-1. ; lat. and single fr. 3-angled) : Fagus (^ fl. sol. or in dich. of 3 ; ? in 3 or sol.) : Nothofagus ( $ in dich. of many ; ? in 2s). 2. Castaneae ( ? fls. in dich. or single in the axils of catkin-1. ; fr. rounded at sides) : Style cylindrical, with pointed stigma, p. ), of starchy consistence. Farinosae (Engler). The 8th order of Monocotyledons. Farinosus (Lat.), covered with mealy powder. FarJjleberry (Am.), Vaccinium arboreum Marshall. Farmeria Willis. Podostemaceae. 2 Ceylon, S. India. Faroa Wehv. Gentianaceae (i). 12 Afr. Farquharia Stapf. Apocynaceae (u. i). i S. Nigeria. Farsetia Turra p.p. Cruciferae (4). 10 E. Medit., trop. Afr. Fasciation, lateral union of stems, branches, &c., Ce/osia. Fascicle, a tuft of branches [aA], fasciculate). Fascicularia Mez. Bromeliaceae (4). 4 Chili. Fastigiate, many branches || to stem, Populus (Lombardy poplar). Fatoua Gaudich. Moraceae (i). i Austr. to Japan, Polynesia. Fats, cf. oils. Fatsia Decne. et Planch, (excl. Echinopanax Decne. et Planch., and Tetrapanax C. Koch). Araliaceae. i Japan, cult. orn. fol. For F. papyrifera Benth. et Hook. f. see Tetrapanax. Faujasia Cass. Compositae (8). 3 Madagascar, Mascarenes. Faurea Harv. Proteaceae (i). 10 S. and trop. Afr., Madag. Fauria Franch. Saxifragaceae (i). i N. Japan. Fawcettia F. Muell. Menispermaceae. i S.E. Austr. Faxonanthus Greenman. Scrophulariaceae (i i). i Mexico. Faxonia T. S. Brandegee. Compositae (5). i Lower California. Faya Neck. = Crenea Aubl. (Lythrac.). Feather-foil (Am.), Hottonia: -grass, Stipa, (Am.) Leptochloa. Fedia Gaertn. p.p. Valerianaceae. i Medit. Other sp. cf. Valerianella. Fedtschenkoa Regel. Cruciferae (4). i Turkestan. Feeria Buser {Trachelium p.p.). Campanulaceae (i). i Morocco. Fegimanra Pierre. Anacardiaceae (i). 2 trop. Afr. Feijoa Berg (Ortkostemon EP.). Myrtaceae (i). 2 Brazil. Felicia Cass. Compositae (3). 60 S. Afr. to Abyssinia. Female fl. , Anctiba, Begonia, Brynnia, Rhamnus, Sagittaria. Fendlera Engelm. et Gray. Saxifragaceae (in), i Texas. Fendlerella A. A. Heller (Fendlera EP.). Saxifrag. (in), i N. Am. Fennel, Fofnicitlum ; -flower, Ni«ella sativa L. ; giant-, Ferula. Fenugreek, Trigonella Foenum-qraecum L. Fenzlia Benth. = Gilia Ruiz et Pav. p.p. (Polemon.). Fenzlia Encll. Myrtaceae (i). 2 Austr. -fer (Lat. suffix), bearing. Feral, wild. FIBRES 265 Ferdinanda Lag. (Zaluzania Pers. p.p.)- Compositae (5). i C. Am. Ferdinandusa Pohl. Ruhiaceae (r. 5). 10 W.I., trop. S. Am. Feretia Delile. Rubiaceae (i. 8). 2 Upper Nile, trop. Afr. Fergusonia Hook. f. Rubiaceae (n. 5). i Ceylon, S. India. Ferment, cf. Carica, Droseraccae. Fern, Filicineae; beech-, Drvopteris; bird's nest-, Aspleniitm\ bladder-, Cystopteris\ bristle-, THchomanes: elkhorn-, Platycerium; filmy, Hyinenophyllum (and fam.) ; hard-, Lomaria, Blechnum; hart's tongue, Pkyllitis; holly-, Polystichum ; lady-, Athyrium', maiden- hair-, Adiantiim ; northern-, Blechnum ; oak-, Dryopteris; parsley-, Cryptogramma; prickly shield-, Poly stic hum; royal-, Osmunda; shield-. Dryopteris; staghorn-, Platycerium; tree-, Cyathea, &c. ; walking-, Asplenium. Fernandezia Lindl. =Lockhartia Hook. (Orchid.). Fernandia Biill. Bignoniaceae (2). i Angola. Fernelia Comm. ex Lam. Rubiaceae (i. 8). 4 Mascarenes. Fernseea Baker. Bromeliaceae (4). t Itatiaia Mt (near Rio). Feronia Correa. Rutaceae (v). i India to Java, F. elephantum Correa (elephant-apple or wood-apple ; wood useful, and yields a gum ; fr. ed.). Feroniella Swingle. Rutaceae (v). i Indo-China. Ferraria L. Iridaceae (n). 10 Afr. Cult. orn. fl. Ferreirea Allem. Leguminosae (in. i). i Rio de Janeiro. Ferrugineus (Lat.), rust-coloured. Fertile, capable of giving fruit. Fertilisation, ihe union of $ and ? elements; cross-, between two fls. on different pi. ; self-, within the same fl. Ferula Tourn. ex L. Umbelliferae (in. 6). 60 Medit., Cent. As. F. communis L. cult, (giant-fennel). It only flowers after storing up materials for some years (cf. Fagus, Agave). F. Narthex Boiss. and F. Aisa-foetida L. are the sources of the drug asafoetida, obtained by notching the roots; used as a condiment in Pen-ia, &c. under the name 'food of the gods,' and as a stimulant in meHicine. F. galba- niflua Boiss. et Buhse and F. rubricaulis Boiss. are the sources of the medic, gum galbanum. Ferulago Koch. Umbelliferae (ill. 6). 40 Medit., S. Eur. Fescue-grass, Fcstuca. Festuca (Tourn. ). Gramineae (to). 100 cosmop. ; 5 in Brit, (fescue- grass). The 1. roll inwards in dry air (cf. Stipa). Many good pasture- grasses. When growing on mountains often viviparous (see fam.). Fever-bush (Am.), Lindera ; -few, Chrysanthemum, Matricaria ; -wort (Am.), Trios ten ni . Feuillea Gled. =Fevillea L. Fevillea L. Cucurbitaceae (i). 6 trop. Am. 5 sta. all alike. Fibigia Medic (Farsetia p.p. BH.}. Cruciferae (4). 12 E. Medit. Fibraurea Lour. Menispermaceae. 4 trop. and subtrop. As. Fibres, the strengthening tissues of plants, largely used in the arts for spinning, brush-making, plaiting, paper, rough weaving, tying, &c. ; those of the bast are most generally useful. The chief stem or leaf fibres are perhaps Abroma, Abutilon, Agave (sisal, &c.). Ampelo- desma, Arenga, Attalea (piassaba), Boehmeria (rhea, ramie), Borassus 266 FIBRES (palmyra), Broussonetia, Camelina, Cannabis (hemp), Carludovica, Caryota (kitul), Chlorogalum, Cocos (coconut, coir), Copernicia, Corchorus (jute), (Jordyline, Couratari, Crotalaria (Sunn-hemp), Cy- perus, Debregeasia. Eryngium (caraguata), Furcraea (Mauritius hemp), Hibiscus, Jubaea, l,aportea, Lardizabala, Leopoldinia (piassaba), Liiuim (flax), Lygeum, Maoutia, Marsdenia, Maurilia, Musa (Manila hemp), Pandanus, Phormium (NewZealandflax), Raphia(raffia), Sabal, Sansevieria (bowstring hemp), Spartium, Stipa (esparto), Tillandsia, Villebrunea, Yucca, &c. The chief surface fibres (on seeds, &c.) are Bombax, Calotropis, Chorisia, Cochlospermum, Eriodendron (kapok), Gossypium (cotton). Special or peculiar cases are Antiaris, Broussonetia, Lageita.Luffa, &c. See Dodge, Useful Fiber Plants, Washington, 1897. Fibrillose, with fibres. Fibrocentrum Pierre. Sapotaceae (inc. seel.), i Brazil. Fibrous root, one in tufts of uniform length, as in grasses. FJ.calb.oa Hiern. Ericaceae (n). i trop. Afr. Ficaria (Dill.) Hall = Ranunculus L. p.p. (Ranunc.). Ficinia Schrad. Cyperaceae (i). 50 S. and trop. Afr. Ficoidales (BH.}. The i4th cohort of Polypetalae. Ficoideae (BH.) = Aizoaceae. Ficus Tourn. ex L. Moraceae (n). 700 trop., chiefly E. Ind. and Polynes., &c. Trees and shrubs of the most various habit. In general alt. entire 1. with slips, which envelope the bud (acting as a protection to it against heat, &c.) and soon after their unfolding drop off. Adv. roots are very common. F. elastica Roxb. (indiarubber tree) grows as a stout independent tree, usu. commencing epiphytically, and often reaching a great size. At its base are developed buttress-roots, radiating out in all directions; their depth is often several feet, while their thickness is only a few inches. From the branches are given off adv. roots which grow downwards and enter the soil. These grow in thickness and form great pillars supporting the branches. The 1. are entire, and leathery, with a glossy surface. The slips, protecl the bud. Rubber is obtained by tapping (cf. Hevea). F. indica L. and F. benghalensis L. (banyan) show similar habit. The aerial roots form supporting pillars, and by their means the tree may reach immense size. (The banyan is sacred in India; the young roots are provided with tubes of bamboo to protect them, and the ground is prepared for them.) See plate in Nat. Pfi. of ihe famous Iree al Calculla. F. religiosa L. (Peepul or Bo-lree) is similar, bul ils 1. have a long acuminale apex, combined with an easily welted surface. From the apex (drip-tip] the rain drips off rapidly after a shower and the 1. is soon dry. In very wet trop. forests this property is of some importance. F. Sycomorusl^. (sycomore or mulberry fig) N. Afr. and F. Carica L. (fig) Eur. , Medit., are also erect trees. F. repens Rottl. is a small climbing sp. which takes hold of its support by aerial roots (as in ivy) ; these secrete a gummy substance conlaining caoutchouc, and then absorb the fluid constituents, leaving the caoutchouc as a cement, fastening the roots to their support Fl L1C ALES 267 (Darwin, Climbing Plants, p. 185). F. Thwaitesii Miq. and other climbing sp. are heterophyllous, the 1. on the climbing shoots small and different in shape. F. Benjainina L. and other sp climb up other trees giving off aerial clasping (negatively heliotropic) roots which surround the trunk. These roots thicken and unite into a network and finally often strangle the 'host' altogether. These sp. often become epiph. by the dying away of their lower portions, but like the Aroids they maintain communication with the ground by long aerial roots. Sometimes they commence as epiphytes and send down aerial roots to the soil. The infl. is hollowed out, and consists of a number of fls. inside a pear-shaped common recept., which opens by a narrow mouth at the top. Within the mouth, in most, are the • sporophylls -*• sporangia t I •*- •*- prothallus The other case, apospory, is found in Athyrium Filix-foemina var. clarissima, and in Polyslichmn augulare var. pulcherrimum, &c. Here spore-formation is replaced by a process of budding which gives rise to prothalli on the backs of the 1., so that the life-cycle runs Fern plant -*• -*• t I , ... , fspermatozoid •«- antheridium ) ., ,, fertilised ovum - { ^ _ archegonium ( ^ P™thall«s The latter must not be confused with the 'vivipary' of Asplenium bulhifenun, &c., where the leaf-tissue buds directly into new pi., which for a time remain attached, but ultimately grow independently {cf. Bryophyllum, &c.). A very large number of ferns are shade- and moisture-loving plants. Many however are xero. and alpine forms with reduced transpiration, exhibiting the familiar char, of such plants — reduced surface, thick cuticle, hairiness, incurving of leaves (cf. Ericaceae), and even, though rarely, succulence (Polypodium adnascens, Drymo- glossum carnosum, &c.). The tree ferns and many others have water storage tissue in the stem. Many are epiphytic, esp. in the trop., though they may be found growing in this way even in Britain. Cf- Platycerium, Polypodium, &c. The spores, consisting only of one cell, are much lighter than is possible for a seed, and may be carried by wind to enormous distances. 2. Hydropteridineae or Rhizocarpae. The two fams. of this group, though they have much in common, are probably derived from different stocks. For details cf. fams. As in the homosporous forms, so here the embryo gives rise directly to a new leafy pi., usu. aquatic, and exhibiting a creeping stem with a dorsiv. arrangement of the 1. Roots may or may not be formed. The stem grows by an apical cell. The sporangia are enclosed in capsular structures termed sporocarps. In the Salv. this body contains one sorus only, in the Mars, more than one. The sorus in the former has one kind of sporangium only, in the latter usu. both. The spores germinate in water; the megaspore gives rise to a small green $ prothallus which remains enclosed in the burst spore. Its free surface bears a few archegonia. The microspore gives rise FLACOURTIACEAE 271 (sometimes without escaping from the sporangium) to a rudimentary G (3), means calyx (corolla) of 3 concrescent sepals (petals), stamens in two whorls of 3 each, free, ovary of 3 concrescent carpels, inferior, and so on ; -kingdoms, see Floral Regions ; -leaves, the parts of the fl., esp. K and C ; -mechanisms, mechanism* to contrive as far as possible cross-fertilisation for a fl., and to ensure that the visiting insect shall receive pollen or touch the stigma, or to ensure self-fertilisation. They may be classed as follows (see individual headings for examples): Anemophily, see Pollination by Wind; Cleistogamy (production of self-fertilising fl.) ; Dichogamy (ripening of cf and ? at different times, with or without movements of sta. and style); Dioecism and other sex-distributions (q.v.)\ Explosive mechanisms ; Heterostylism ; Loose-pollen mechanisms ; Piston- mechanisms; Pollen-prepotency \ Pollination by Animals; Pollination by Water ; Pollination by Wind ; Self-sterility ; Sensitive stamens ; Sensitive stigmas ; Special mechanisms, such as Asclepiadaceae, Ficus, Orchidaceae, Salvia, Yucca, &c. ; Style-projection; Trap Flowers, &c. See Knuth, Handbook of Floral Pollination ; Oxford ; -regions, w. 18 274 FLORAL regions char, by the possession of a considerable number of local (endemic) forms ; the greater their number, and the higher their systematic rank, the more natural is the region. Engler divides the world as follows : I. Northern txtra-tropical Floral Kingdom, with 9 regions) (i) Arctic, (2) Sub-Arctic or Conifer, (3) Mid- European, (4) Macronesian (Azores, Madeira, Canaries, Cape Verdes), (5) Medi- terranean, including land all round that sea, (6) Central Asiatic, (7) Temp. E. Asiatic, (8) Pacific N. Am., (9) Atlantic N. Am.: II. Palaeotropic Floral Kingdom, with 9 regions, (i) N. Afr. — Indian desert, (2) Afr. forest and steppe, (3) S. W, S. Afr., (4) S. Atl. islands, (5) Madagascar and islands, (6) Nearer India, (7) Jlfousoonia (warmer Himal., Malaya, N. Austr., Polynesia) (this and 6 are usu. united in this book as Indomalaya, and sometimes called E. or W.), (8) E. China and S. Jap., (9) Hawaiian : III. Centr. and S. American Floral Kingdom, with 5 regions, (i) Mid- American Xero. (S.W. U.S., Mex.), (2) Trap. Am., (3) Andinf, (4) Galapagos, (5) Juan Fetnandez: IV. Austral Floral Kingdom, with 6 regions, (i) Antarctic S. Am., (2) Antarctic Continental, (3) Kerguelen, (4) New Zealand, (^Australia, (6) Tristan da Cunha, St Paul, Amsterdam : V. Oceanic Floral Kingdom, with 3 regions, Boreal, Tropical, and Austral; -symmetry shows two chief cases, if radial the fl. is regular or actinomorphic (even if the cpls. be fewer in number), Ran^^nc^^laceae, Kosaceae, &c., if not, some members being omitted in the outer whorls, or all members of a whorl not alike, it is irregular and may be zygomorphic (divisible into two halves, each the reflection of the other), Labiatae, Scropkulariaceae, &c., or asymmetrical (not so divisible), Valeria naceae. Cf. Floral Diagram, &c. Flores verbasci, Verbascum. Florestina Cass. Compositae (6). 3 Mexico. Floret, small fl. of a cluster. Floribundus (Lat. ), producing many fls. Florida Velvet Bean, Mucuna. Floridus (Lat.), showy. -florus (Lat. suffix), -flowered. Flos (Lat.), a flower. Floscopa Lour. Commelinaceae. 15 trop. and subtrop. Flotovia Spreng. r=Chuquiraga Juss. (Compos.). Flourensia DC. Composiiae (5). 10 Arizona to Argentina. Flower, a repr. short shoot, consisting of an axis (receptacle, q.v., thalanms, torus), bearing essential organs Qi.sporophylls, the androe- ceiiin or stamens (i/.v.), and gynoeceum or carpels (q.v.}, or only one of them in diclinous fl. ; also usu. some accessory organs or perianth (<].v.), most often divided into an outer green whorl or calyx, and inner coloured one or corolla', and insects, the relations between fls. and the insects which pollinate them, cf. Flower-classes, Floral Mechanisms, &c.; -axis, the receptacle; -bud, the young fl.. in which the 1. are packed in a definite way (aestivation, q.v.) ; -classes, the classes into which fls. may be divided according to their relations to insects, &c. ; W (wind-pollinated), Po. (offering pollen only), A (with freely exposed honey), AB (partly concealed honey), B (fully concealed honey), B' (do. in aggregated infls.), F (Lepidoptera fls. with long FODDER 275 tubes), H (bee fls. with long tubes, zygomorphism, &c.) ; see individual classes for examples ; colours of - (usu. in corolla) are due to chloro- plastids or bodies carrying chlorophyll (rare), Deherainea, chromo- plastids or bodies carrying colours, or to coloured cell-sap ; all colours change readily to white, and there are several cases of change (q.v.} of one colour to another ; bees are inclined to prefer blue, butterflies red and white ; -cup, a hollow receptacle ; -de-luce, Iris ; -descrip- tion, cf. Description ; -, doubling of, change of sta. to petals, or in Compositae of tubular to ligulate fl. ; -fence \Barbados), Caesalpinia pidcherriina Sw. ; -mechanisms, see Floral ; -movements, protecting against cold and wet, e.g. a bending downwards, Anemone, Bellis, Daucus, Fragaria, Linuin, Papaver, or a closing of the petals, Ana- gallis, Bellis, Calandrinia, Eschscholtzia, Tragopogon; -pride (W.I.), Caesalpinia pulcherri ma Sw. ; -tube, the concrescent portion. Flowering ash, Fraxinus ; -currant, Ribes sanguineum Pursh ; -fern, Osmnnda ; -rush, Butomus umbellatus L. Fluckigeria Rusby (Kohlerianthns Fritsch, EP.). Gesneriaceae (i). i Bolivia. Flueckigeria O. Ktze. (Ledenbergia Klotzsch). Phytolaccaceae. i trop. 8. Am., W.I. Flueggea Rich. = Ophiopogon Ker-Gawl. (Lili.). Flueggea Willd. Euphorbiaceae (A. I. i). 6 palaeotrop. Flueggeopsis K. Schum. (Phyllanthus p.p. EP.). Euphorb. (A. I. i). 3 Malaya. Fluitans (Lat.), floating. Fluviales= Helobiae. Fluviatilis (Lat.), growing in streams. Fly-flowers, Amorpkophallus, Araceae, Arum, Asariim, Cobaea, Com- positae, Crassulaceae, Cynanchum, Hedera, Helicodiceros, Paris, Stapelia, Umbelliferae, Veronica ; -orchis, Ophrys musdfera Huds. ; -trap, American, Apocynnm ; - -, Venus', Diouaea muscipula Ellis. Fockea Endl. Asclepiadaceae (11. 3). 6 Afr. Fodder. The food of grazing animals, &c. The grasses (fresh or dry) and Leguminosae (esp. the pods) are most generally useful. In- numerable pi. are used in different countries ; among the most important are Acacia, Agrostis, Alopecurus (fox-tail grass), Andro- pogon, Anthoxanthuin, Autliyllis, Arachis, Aristida, Astragalus, Atriplex, Avena (oat), Bonteloua (mesquit grass), Brassica, Briza, Bromus, Buchtoe( buffalo grass), Celt is, Cenchrus, Ceratonia (algaroba), Chionachne, Ckloris, Chrysopogon, Cicer (chick-pea), Cyamopsis, Cynodon (Bermuda grass), Cynosurns (dog's tail grass), Dactylis (cock's foot grass), Daucus, Deschampsia, Desmoaium, Dolichos (horse-gram), Ehrharta, Eleusine (ragi), Eragrostis, Eriochloa, Eruca, Ervum, Fagopyrum (buckwheat), Festztca (fescue), Galega, Glyceria, Gossypium (cotton-seed), Heteropogon, Hippocrepis, Holcus, Hordciun (barley), hnperala, hchaemum, Laihyrus, Leersia, Lespe- ateza, Loihun (rye gra>s), Lotus, Lupmus (lupin), Medicago (lucerne, cS:c.), Melica, Melilotus, Milium, Miicuna (Florida velvet bean), Muehlenbergia, Musa, Onobrychis (sainfoin), Ormtliopus, Oryza (rice), Panicum (millet, Guinea grass, &c.), Paspctlum, Penmsetnm (bajri), Phaseolus.(gi;a.m, beans), Phleuin (timothy), Pisum (pea), Poa (meadow l8—2 276 FODDER grass), Secale (rye), Sesamum (gingelly), Setaria (Italian millet), Sor- X/inm (Guinea corn), Sptrgula, Symp/iyt-inii, Trifolium (clover, &c.), Tripsaciim, Triticum (wheat), Vicia (vetch), Zea (maize), Zizyphus. Foeniculum Tourn. ex L. Umhelliferae (in. 5). 4 Medit, Eur., i Brit. (fennel). The young 1. of F. officinale All. are a good veg. when blanched like celery, and the fr. is a condiment. Foetid horehound, Ballot a nigra L. Foetidia Comm. ex Lam. Lecythidaceae. 3 Madag., Masc. Fokienia A. Henry et H. H. Thomas. Coniferae (Pinaceae, see C. for gen. char.), i E. China. Folia Jaborandi, Pilocarpus. Foliaceous, leaf-like, leaf-bearing. Foliage plants, cult, for orn. foliage, e.g. Beponia, Caladitim, Codiaeitm, Coleus, many Coniferae, Cordy/ine, Cortaderia, Dracaena, Eryngium, many Ferns, Fitlonia, Gitnnera, Gesneria, many Palmae, Panax, Pelargonium, Phormiiim, Rheum, Rhus, Selaginella, Smilax, Sone- rila, many trees. Folium (Lat), a leaf; -folius (Lat. suffix), leaved. Follicle, a dry dehisc. fr. of one cpl., dehisc. on ventral side only; Aconitum, Apocynaceae, Asclepiadaceae, Banksia, Crass ulaceae. Folotsia Costantin et Bois. Asclepiadaceae (n. i). i Madag. Fonna Lunell (Phlox p.p.). Polemoniaceae. 2 W. U.S. Fontainea Heckel. Euphorbiaceae (A. n. 5). i New Caled., E. Austr. Fontanesia Labill. Oleaceae. i Sicily, W. As. Fontanus (Lat.), growing in or near a spring. Food, see Edible Products, Fodder ; -bodies, Acacia, Cecropia ; - of the gods, Ferula. Fool's parsley, Acthusa Cynapium L. Foot, organ attaching fern plant to prothallus. Foramen, an aperture. Forbesia Eckl. (Cnrculigo p.p. BH.}. Amaryllid. (in). 6 Afr. Forbidden fruit (W.I.). Citrus aurantium L. var. paradisi. Forchhammeria Liebm. Capparidaceae (in). 3 Mex., W.I. Forcipella Bail!. Acanthaceae (iv. B). i Madag. Forcipella Small (Siphonychia EP.). Caryophyll. (i. 4). i N. Am. Fordia Hemsl. Leguminosae (m. 6). i S. China. Fordiophyton Stapf. Melastomaceae (i). 2 S. China. Forest, a close assemblage of trees, allowing no break in the overhead canopy; homogeneous (of one sp.), or diversified ; -oak, Casuarina. Forestiera Poir. Oleaceae. 15 Am., W.I. Forficaria Lindl. Orchidaceae (n. i). i S. Afr. Forgesia Comm. ex Juss. Saxifragaceae (v). i Bourbon. Forget-me-not, Myosotis. Fork-veined, veins forking into two, as in ferns. Forms of vegetation, groups of pi., of various fams , presenting a general resemblance in external habit, often correlated with resemblance in conditions of life, e.g. trees, epiphytes, &c. Formula, floral, see Floral formula. Forrestia A. Rich. Commelinaceae. 10 palaeotrop. Forsellesia Greene (Glossopetalon A. Gray). Celastr. 4 N. Am. Forskohlea L. Urticaceac (5). 5 Medit. to India. FRANKENIACEAE 277 Forstera L. f. (Phyllachne p.p. EP.}. Stylidiaceae. 4 Tasm., N.Z. Forsteronia G. F. W. Mey. Apocynaceae (n. 2). 30 trop. Am., W.I. ForsytMa Vahl. Oleaceae. 2 China. Cult. orn. fl. shrubs. Forsythiopsis Baker. Acanthaceae (iv. A). 2 Maclag. Fortunearia Rehder et Wilson. Hamamelidaceae. i China. Fortunella Swingle (Citnis p.p.). Rutaceae ^v). 4 E. As. (cumquats). Fortuynia Shuttl. ex Boiss. Cruciferae (2). 2 Persia, Afghanistan. Fothergilla Murr. Hamamelidaceae. 2 All. N. Am. (Am. witch elder), Cashmir. Fl. apet, A oo . Fouquieria H. B. et K. Fouquieriaceae. 5 warm N. Am., incl. F. splendens Engelm. (ocotilla, coach-whip), used for hedges. Shrubs with deciduous 1., the midribs persistent and thorny, and showy fl. in racemes, &c., ?, reg., 5-merous. C (5), A 10 — 15, G (3) with 4 — 6 ov. on plac. in middle of ventral side. Fr. spherical, 3-loc.; seeds with long hairs or wings. Fouquieriaceae (EP.\ Tamaricaceae p.p. BH.}. Dicots. (Archichl. Parietales). Only gen. Fouquieria (q.v.}. Fourcroya Sprang = Furcraea Vent (Amaryllid.). Four-o'clock, Mirabilis Jalapa L. Fourniera Scribner. Gramineae (3). i Mexico. Fournieria Van Tiegh. (Cespedesia EP.). Ochnaceae. i C. Am. Foveolaria Ruiz et Pav. Styraceae. i Peru. Fowl-meadow grass (Am.), Poa, Glyceria. Fox-glove, Digitalis purpurea L.; -grape, Vitis Labrnsca L. ; -tail grass, Alopecurus pratensis L., (Am.) Spartina patens Muhl. Fragaria (Tourn.) L. Rosaceae (in. 2). 10 *, Chili. F. vescaL,. (wild strawberry), Brit. Veg. repr. by runners is well shown. Fl. protog. (class AB), with epicalyx. Fr. of a number of achenes (the so-called seeds) upon a fleshy recept. The fl. bends down after fert, while the fr. ripens. In Am. the cult, forms tend to become dioec. or polyg. Several sp. cult, for the ed. fr. Franchetia Baill. Hamamelidaceae. i Madag. Franciscea Pohl = Brunfelsia L. p.p. (Solan.). Francoa Cav. Saxifragaceae (n). 2 Chili. Cult. orn. fl. (wedding flower) . Frangipani, Plumeria. Frangula Tourn. ex Hall. =Rhamnus L. p.p. (Rhamn.). Frangulinae (Warming), the i6th order of Choripetalae. Frankenia L. Frankeniaceae. 32 sea-coasts, temp, and subtrop. F. laevis L., sea-heath, in Brit. Halophytes with inrolled hairy 1. (cf. Empetrum). Frankeniaceae (EP., BH.}. Dicots. (Archichl. Parietales; Caryo- phyllinae BH. }. 4 gen., 60 sp. of salt-loving plants, trop. and temp, herbs with jointed stems; 1. opp., inrolled, exstip. (?). Fls. in dichasia, 5 , reg. K (4 — 7), C 4 — 2, A usu. 6 in two whorls, sta. slightly united at base; G usu. (3), i-loc. with parietal plac., only the lower parts of which bear ovules; ov. oo , anatr., ascending; style forked. Caps, loculic. Mealy endosp. ; embryo straight. Chief genera : Frankenia, Niederleinia. Closely related to Tamaricaceae and Gutti- ferae ; the agreement with Caryophyllaceae, near to which it is sometimes placed, e.g. by BH., is more in habit than in structure. 278 FRANKINCENSE Frankincense, BosweUia-Carteri Birdw. ; -pine, Finns Taeda L. Franklandia R. Br. Proteaceae (i). 2 W. Austr.' Franseria Cav. Compositae (5). 16 Am. Frantzia Pittier. Cucurhitaceae (4). 2 C. Am. Frasera Walt. (Swertia EP.}. Gentianaceae (i. 3). 8 N. Am. Fraunhofera Mart. Celastraceae. i Brazil. Fraxinus Tourn. ex L. Oleaceae. 50 sp. esp. N. Am., E. As., and Medit. F. excelsior L., ash, in Brit. Serial accessory buds in axils. Has large pinnate 1. with grooved petioles. Water is said to enter , this groove and be absorbed by the 1. ; the hollow is nsu. inhabited by acarids, forming a domatiutn. The fls. appear before the 1. in densely crowded short racemes. Each $ consists merely of 2 sta. ± i cpls., and is anemoph. ; but polygamy is the rule in this sp. and every possible combination of the three types of fl. ( 5 , one cpl. ; but in practice the large are nuts, the small achenes. True achenes in Ranunculus, Potentilla, &c.. but the name is also given to the fr. of Compositae, Gramineae (this variety, with pericarp and testa united, is sometimes called a caryopsis), Labiatae, &c. True nuts, from sup. ov., in Betulaceae, but the term is applied to the large one-carpelled fruit of Anacardium, &c. A var. of achene or nut is the winged one- seeded indeh. samara of ash, elm, Banisteria, Liriodendron, Ptelea, Seguieria, Ventilago, &c. Schizocarps various, e.g. the lomenlitm of many Leguminosae (a pod constricted between seeds, breaking into one-seeded portions), the schizocarps of Euphorbiaceae, Geraniaceae, Malpighiaceae, many Malvaceae, Sapindaceie, Umbelliferae. Dry dehisc. fr. of several kinds, esp. the follicle, legume, and capsule. Follicle of one cpl., dehisc. along ventral side only, Aco- nitum, Asclepiadaceae, Apocynaceae, Crassulaceae. Legume similar but dehisc. along both sides, as in most Leguminosae. Dry fr. of > i cpl. are capsules, but special forms have special names, e.g. the pod-liice siliqua of Cruciferae, the f>yxi* of Anagallis, &c. (capsule opening by a lid split off by circnmscissile dehisc'-nce), and others. The way in which it dehisces is of systematic importance. It usu. splits from apex down. If the splits, as in Epilobium, Iris, &c. , run down the midrib of each cpl., the dehisc. is loculicidal: if, as in Hypericum, the fr. breaks into its component cpls. , leaving the placental axis standing, it is septicidal; if the outer wail of the fr. breaks away, leaving the septa standing, it is septifiagal. The portions into which the fr. splits are termed valves. In some Campanulas, 280 FRUIT Papaver, &c., the dehiscence is porous, little openings forming in the pericarp. The commonest fleshy fr. are the berry and drupe. The former contains no hard part but the seeds ; these are surrounded by fleshy tissue and there is a firmer skin (epicarp] on the outside. Berries may be derived from sup. ovaries, as in Berbeiis, Solamim, Vitis, &c. or inf., as in Ribes, Vaccinium, &c. In rare cases the berry dehisces, Akebia, Myristica, or is constricted between the seeds, Maerua, Unona. The drupe (e.g. cherry) has a skin (epicarp] on the surface, then a fleshy mass of tissue (mesocarf) and a hard shell or stone (emfacarp}, all forming part of the pericarp; within the stone is the seed or kernel, usu. without a hard coat. Drupes from sup. ovaries occur in Prunus, &c., from inf. in Cornaceae, Juglans, &c. There may be one stone or pyrene (Prunus) or several (Cornus). Other fleshy fr. are the pome of Pyrus, &c. in which the fleshy receptacle encloses, and is united to, the core or product of the G proper ; the pepo or gourd of Cucurbitaceae, e.g. cucumber (a variety of the berry with hard epicarp). the peculiar pseudo-berry of Juni- perus (q.v.) &c. , the fr. of strawberry (fleshy recept. bearing achenes), rose (fleshy recept. enclosing achenes), Anacardium (fleshy recept. bearing nut), Gaultheria (caps, enclosed in fleshy calyx), Urera, &c. (achene in fleshy perianth), and so on. Aggregate fleshy fr. in Ano- naceae (berries), Rubus (drupes), &c. Multiple fleshy fr. frequent in Moraceae (e.g. mulberry, fig, bread-fruit), Ananas, Anona, Carlu- dovica, &c. The style and stigma often fall away as the fr. ripens, but fre- quently remain in a ± shrivelled or in an enlarged condition. Some- times the style forms a hook, as in Gcum, a plume, as in Clematis, or an awn, as in Geraniaceae (this name is applied to any long thread- like organ on a fruit; cf. Gramineae). Other interesting morphological features in fruits: cf. Hetero- carpy, Aesculus, Bertholletia, Chenopodium, Leontodon, Nymphaea- ceae, Nyctaginaceae, Palmae (e.g. Phytelephas, Lodoicea, &c. ), Pandanaceae, &c. Frutex, a shrub; frutescent, fruticose, shrubby. Fuchsia (Plum.) L. Onagraceae (2). 65 C. and S. Am., N.Z. Many cult. orn. fl. Many show two buds in each axil, one above the other. Fl. suited to bees, humming-birds, &c. Berry ed. Fuernrohria C. Koch. Umbelliferae (in. 3). i Armenia. Fuertesia LJrb. Loasaceae. i San Domingo. FuertesieUa Schlechter. Orchidaceae (n. 2). i San Domingo. Fugacious, falling early. Fugosia Juss. (Cienfnegosia EP.}. Malvaceae (4). 30 Am., Afr. , Au^tr. Fuirena Rottb. Cyperaceae (i). 25 trop. and subtrop. Fuliginous, sooty. Fuller's teasel, Dipsactts fullonum L. Fulvous, tawny. Fumana Spach (Hdianthennnn p.p. ). Cistaceae. 8 Medit., Eur., W. As. Fumaria Tourn. ex L. Papaveraceae (in). 40 Eur., As., Afr., chiefly GALAGANFA 281 Medit., i in Brit, (fumitory). Many climb by sensitive petioles (cf. Clematis). Fl. like Corydalis. F. capreolata L. var. pallidiftora Jord. (Brit.) shows colour-change in its fl. ; before pollination white, it gradually turns pink or carmine (cf. Ribes, Diervilla). Fumariaceae ( Warming) = § in of Papaveraceae. Fumariola Korshinsky. Papaveraceae (m). i Turkestan. Fumitory, Fnmaria. Funastrum Fourn. Asclepiadaceae (n. i). 2 trop. S. Am. Funereal cypress, Cuprcssus funebris Endl. Funicle, the stalk of the ovule; funiculate (ovule), stalked. Funifera, Leandr. ex C. A. Mey. Thymelaeaceae. 2 Brazil. Funkia Spreng. (Hosta Tratt. ). Liliaceae (in). 5 Japan, China. Embryos are formed in the seeds by outgrowth of the nucellus-tissue round the embryo-sac (cf. Alchornea). Seeds winged. Cult.orn. fl. Funtumia Stapf (Kickxia Blume p.p.). Apocynaceae (n. i). Strop. Afr. F, elastica Stapf is the chief source of Lagos or Ire rubber. Furcate, forked. Furcraea Vent. Amaryllidaceae (n). 20 trop. Am- Like Agave; infl. even larger. F. gigantea Vent, yields fibre (Mauritius hemp). Furcroya Rafin. = Furcraea Vent. (Amaryll.). Furfuraceous, with soft scales. Furze, Ulex eitropaeus L., &c. Fusaea W. E. Safford. Anonaceae (i). i Guiana, Venezuela. Fusanus R. Br. Santalaceae. 5 Austr., N.Z. Fuscous, dusky. Fusiform, spindle-shaped. Fustic, Chlorophora, Madura. Zanthoxyhtm ; young-, Rhus. Gabila Baill. (Pycnarrhena EP.}. Menispermaceae. i Timor. Gabunia K. Schum. (Tabernaemontana p.p.). Apocynaceae (i. 3). 7 W. Afr. Gad-bush (VV. I.), Arcettthobium gracile Engehn. Gaertnera Lam. Rubiaceae (11. 5). 30 trop. Afr. and As. Gagea Salisb. Liliaceae (iv). 30 N. temp. Old World. G. iutea Ker-Gawl. in Brit. Fl. protog In the 1. -axils of some are buds which, if fert. does not occur, develope into bulbils and drop off. Gagernia Klotzsch. Ochnaceae. i Guiana. Gagnebina Neck. Leguminosae (i. 4). i Mauritius, Madag. Gagnepainia K. Schum. (Hemiorchis p.p.). Zingiber. (i). 3 S.E. As. Gahnia Forst. Cyperaceae (n). 35 1%, esp. Austr. Gaiadendron G. Don (Loranthus p.p. BH.}. Loranth. (i). 4 Andes, Austr. Gaillardia Fouger. Compositae (6). 15 Am. Cult. orn. fl. Gaillonia A. Rich. Rubiaceae (11. 10). 12 Nubia to India. Gaimardia Gaudich. in Freyc. Centrolepid. 3 temp. S. Am., N.Z., Austr. Galactia P. Br. Leguminosae (in. 10). 70 trop. and subtrop. Latex, which is rare in the fatn., is found in this plant. Galactites Moench. (Lupsia EP.}. Comp. (n). 3 Medit., Canaries. Galactodendron Rchb. (-um Kunth) = Brosimum Sw. (Morac.). Galactoxylon Pierre (Bassia p.p. ). Sapotaceae (i). i Malay Arch. Galagania Lipsky. Umbelliferae (in. 5). i C. As. 282 GALANTHUS Galanthus L. Amaryllidaceae (l). 6 Eur., Medit. G. nivalis L. (snowdrop) in Brit. Bulb with i-fl. scape. P in two whorls. On the inner surface of the inner P-l. are green grooves secreting honey. The bud is erect, but the open fl. pendulous, visited by bees. The sta. dehisce by apical slits and lie close against the style. Each has a process outwards from the anther. The stigma projects and is first touched by an insect; in probing for honey it shakes the sta. and receives a shower of pollen (cf. Erica). Autogamy may occur in old fls. The fl. remains open a long time. Cult. orn. fl. Galapee tree (W.I.). Stiadophyliitm. Galarhoeus Haw. = Euphorbia L. p.p. (Euph.). Galatea, Galatella Cass. = Aster Tourn. p.p. (Comp.). Galax L. Diapensiaceae. i Virginia, Georgia. Galaxia Thunb. Iridaceae (l). 4 S. Afr. Galba (W. I.), Calophyllum Calaba Jacq. Galbanum, gum, Ferula. Galbulimima F. M. Bailey. Magnoliaceae. i Austr. Gale, sweet, Myrica Gale L. Galeana La Llave. Compositae (6). i Mexico. Galeandra Lindl. Orchidaceae (n. 5). 7 trop. Am. Epiphytes. Galearia Zoll. et Morr. Euphorbiaceae (A. n. 5). 12 Indomal. Galeata Wendl. Inc. sed. i Indomal. Galeate, helmet-shaped. Galedupa Lam. (Pongamia Vent.). Legum. (in. 8). i trop. As., Austr. Galega Tourn. ex L. Leguminosae (in. 6). 3 S. Eur., W. As. G officinalis L. sometimes cult, as a fodder-plant (goat's rue). Galenia L. Aizoaceae (n). 18 S. Afr. Galeobdolon Adans. = Lamium Tourn. p.p. (Labi.). Galeola Lour. Orchidaceae (n. 2). 10 Malay Archipelago. Galeopsis L. Labiatae (vi). 7 N. temp. |*. 3 in Brit., inch G. Tetrahit L. (hemp-nettle) with swollen upper ends to internodes, acting as pulvini. Galeorchis Rydberg (Orchis p.p.). Orchidaceae (n. i). i N.Am. Galeottia A. Rich. (Zygopetalum BH.}. Orchidaceae (n. 14). 2 Colombia. Galera Hlume (Epipogum BH.}. Orchidaceae (n. 2). 3 As. Galingale (Am.), Cyperus. Galiniera Delile. Rubiaceae (i. 8). i Abyssinia. Galinsoga Ruiz et Pav. Compositae (5). 5 Mexico to Argentina. G. parviflora Cav. now a common weed in Eur. and near Kevv. Galipea Aubl. Rutaceae (i). 6 S Am. Galium L. Rubiaceae (ll. n). 220 cosmop. ; to in Brit, (bed-straw, &c.). Herbs with whorls of 1. and slips, (see fam.) ; fls. in dichasial panicles, small with honey freely ex .osed (class A) on the epig. disc, usu. protandrous with ultimate self-pollination. G. Aparine L. (goose- grass or cleavers; Brit.) is a feeble hook-climber with small reflexed hooks on the stem. The schizocarp is also provided with hooks. Gallesia Casar. Phytolaccaceae. i Peru, Brazil. GaUitrichum Fourn. = Salvia Tourn. (Labiatae). Gallnuts, cf. Myrobalans. GARJAN OIL 283 Galopina Thunb. Rubiaceae (n. 7). 2 $. Afr. Galphimia Cav. Malpighiaceae (n). 12 warm Am. Cult. orn. shrubs. Galpinia N.E. Br. Lythraceae. i Transvaal. Galpinsia Britton (Oenothera BH.). Onagraceae. 6 N. Am. Galtonia Decne. Liliaceae (n). 2 S. Afr. Cult. orn. fl. (spire lily). Galvesia Domb. ex Juss. Scrophulariaceae (n. 3). 3 Peru to Calif. Galypola Nieuwland (Polygala p.p.). Polygalaceae. i U.S. Gama grass (Am.), Tripsacum, Gambeya Pierre (Chrysophyllum p.p. EP.). Sapot. (i). 5 trop. Afr., Am. Gambir, Uncarin Gambler Roxb. Gamblea C. B. Clarke. Araliaceae (i). i Sikkim. Gamboge, Garcinia Morella Desr., &c. Gamete, a sexual reproductive cell.. Gametophyte, the sexual generation. Gamo- (Gr. prefix), united ; -petalae (BH.), Sympetalae ; -petalous, -phyllous, -sepalous, with concrescent C, P, K; -tropic (movement), before fert. Gamocarpha DC. = Boopis Juss. (Calycer.). Gamogyne N.E. Br. Araceae (v). 2 Malaya. Gamolepis Less. Compositae (8). 12 S. Afr. Gamopoda Baker. Menispermaceae. i Maclag. Gamosepalum Hausskn. Cruciferae (4). 2 W. As. Gamosepalous. Ganja, Cannabis saliva L. GanophyUum Blume. Sapindaceae (n) (Burser. BH.). i Phils, to Austr. Ganua Pierre ex Dubard (Illipe p.p.). Sapot. (i). 6 Malaya. Ganymedes Salisb. = Narcissus Tourn. p.p. (Amaryll.). Garapatica Karst. (Alibertia BH.). Rubiaceae (i. 8). i Colombia. Fr. ed. Garberia A. Gray. Compositae (2). i Florida. Garcia Rohr. Euphorbiaceae (A. II. 3). i trop. Am. Garcilassa Poepp. et Endl. Compositae (5). i Peru. Garcinia L. Guttiferae (v). 200 palaeotrop. trees or shrubs with leathery 1. Sta. free or united into bundles or into a common mass. Berry ; seed arillate. The resin of G. Morella Desr. and other sp., obtained by cutting notches in the stem, forms gamboge. The fr. of many is ed., esp. that of G. Mangostana L. (mangosteen), the aril of the seed of which is a delicacy. Some yield useful timber. Gardenia Ellis. Rubiaceae (i. 8). 80 palaeotrop., largely cult. orn. perf. fl. Some sp. have apparently whorls of leaves, 3 in each, really a case of condensation of two whorls of 2 into one with extreme anisophylly of one whorl ; the fourth 1. is reduced to a minute scale. The stipules of many secrete a resinous fluid. Gardeniopsis Miq. Rubiaceae (n. 4). i Sumatra, Borneo. Gardneria Wall, ex Roxb. Loganiaceae. 3 India to Japan. Gardoquia Ruiz et Pav. (Saturda p.p. EP.}. Labiatae (vi). 30 W. Am. Garget (Am.), Phytolacca. Garhadiolus Jaub. et Spach (Rhagadioius BH.). Comp. (13). 4 W. As. Garidella Tourn. ex L. =Nigella L. (Ranunc.). Garjan oil, Dipterocarpus. 284 GARLIC Garlic, A/lfum ursinum L., &c. ; -pear tree (W.I.), Crataeva gynandra L. ; -shrub (W. I.), Bignonia alliacea Lam. Garnieria Brongn. et Gris I'roteaceae (i). i New Caledonia. Garnotia Brongn. in Duperr. Gramineae (8). 15 Indomal., China, Jap. Garnotiella Stapf. Gramineae (8). i Phil. Is. Garretia Welw. (Khaya EP.}. Meliaceae (n). r trop. Afr. Garrya Dougl. ex Lindl. Garryaceae. 15 N. Am., W.I. Shrubs with 4-angled twigs and opp. evergr. 1. Fl. in catkin-like panicles, 3 in axil of each bract, unisexual, i P 4, A 4 ; ? naked, G(2 — 3), uniloc. with 2 ovules. Fr. with thin pericarp and i — 2 seeds. Endo- sperm. Cult. orn. shrubs. Garryaceae (EP. ; Comaceae p.p. BH.). Dicots. (Archichl. Garryales). Only genus Garrya, q.v. Garryales. The 4th order of Dicots. Archichlamydeae. Garuga Roxb. Burseraceae. 6 Indomal. Garugandra Griseb. (Gleditschia p.p. EP.). Leguminosae (n. 7). (Anacardiaceae, BH.) i Argentina. Garuleum Cass. Compositae (9). s'S. Afr. Gaslondia Vieill. (Syzygium p.p. EP.). Myrtaceae (i). i New Caled. GaspariUo ( W. I . ), Esenbeckia. Gasteria Duval. Liliaceae (n). 35 S. Afr. Xero. with succulent 1. closely packed together, but often in the shade of long grass. Gastonia Comm. ex Lam. Araliaceae (i). 5 Madag., Masc. Gastranthus Moritz ex Benth. et Hook. f. Acanth. (iv B). i Venezuela. Gastridium Beauv. Gramineae (8). 2 Medit. Gastrochilus Wall. Zingiberaceae (i). 2 Himal., Burmah. Gastrocotyle Bunge. Boraginaceae (iv. 2). i Egypt to Panjab. Gastrodia R. Br. Orchidaceae (u. 2). 12 India to N.Z. Gastroglottis Bluine (Liparis p.p.). Orchidaceae (n. 4). 2 Peru. Gastrolepis Van Tiegh. Icacinaceae. i New Caledonia. Gastrolobium R. Br. Leguminosae (in. 2). 35 W. Austr. Gatesia A. Gray. Acanthaceae (iv. B). i U.S. Gaub tree, Diospyros Embryopteris Pers. Gaudichaudia H. B. et K. Malpighiaceae (i). 15 Mexico to Vene- zuela. Mericarp elevated on carpophore formed from wing of cpl. Gaudinia Beauv. Gramineae (9). 2 Medit. Gaultheria Kalm. ex L. Ericaceae (n. 2). 100 sp. Am., Japan to Himal. and Tasmania. The fruit resembles a berry, but is really a capsule, enclosed in the fleshy calyx but not adhering to it. G. proctttnbens L. (winter green, checker-berry or partridge-berry, U.S.). G. Shallon Pursh (sallal, shallon, N.W. Am.). Winter- green oil is distilled from the pi. of some sp. Gaura Linn. Onagraceae (2). 25 N. Am. The anthers are chambered up by horizontal septa in each foe. (cf. Circaea). Nut. Gaurella Small (Oenothera p.p.). Onagraceae (2). 2 N. Am. Gauropsis Presl (Clarkia p.p. EP.). Onagraceae (2). i Mexico. Gaussia H. Wendl. Palmaceae (iv. r). i Cuba. Gauze tree (W. L). Lagetta. Gavarretia Baill. Euphorbiaceae (A. n. 2). 2 N. Brazil. Gaya H. B. et K. Malvaceae (2). 8 trop. Am. GENIOSPOR UM 285 Gaylussacia H. B. et K. Ericaceae (in. i). 40 Am. (huckleberry). The 5 loc. of the ovary are made into 10 by partitions growing out from the midribs of the cpls., as in Linum. Gayoides Small (Abutilon p.p. £f., Sida p.p. BH.}. Malv. (2). 2 N. Am. Gayophytum A. Juss. Onagraceae (2). 6 Chili to Calif. Gazania Gaertn. Compositae (10). 24 Cape Colony. Cult. orn. fl. Gean. Prunits Avium L. Geanthemum R. E. Fries. Anonaceae (i). i Brazil. Cf. Anona. Geanthus Phil. Liliaceae (iv). 2 Chili. Gearum N.E. Br. Araceae (vn). i Goyaz. Geaya Costantin et Poisson. Ericaceae (iv. i). i Madag. Geigeria Giiesselich. Compositae (4). 30 S. and trop. Afr. Geijera Schott. Rutaceae (i). 5 E. Austr. Geissanthera Schlechter. Orchidaceae (n. 20). i New Guinea. Geissanthus Hook. f. Myrsinaceae (n). 25 equatorial S. Am. Geissaspis Wight et Arn. Leguminosae (in. 7). 8 trop. Afr., As. Geissois Labill. Cunoniaceae. 6 Austr. to Fiji. Geissolepis Robinson. Compositae (5). i Mexico. Geissoloma Lindl. ex Kunth. Geissolomataceae. G. marginal inn Kunth, Cape Col., a small xero. shrub, the onlysp. L. opp. evergr., with sol. axillary fls., 5 . K 4, C o, A 4 + 4, G (4) each with 2 pend. ov. Caps. 4-loc. Endosp. Geissolomataceae (EP. ; Penaeaceae p.p. BH.}. Dicots. (Archichl. Myrtiflorae). Only genus, Geissoloma, q.v. Geissomeria Lindl. Acanthaceae (iv. B). to trop. Am. Geissopappus Benth. Compositae (6). 3 trop. S. Am. Geissorhiza Ker. Iridaceae (m). 40 S. Afr., Madag. Cult. orn. fl. Geissospermum Allem. Apocynaceae (i. 3). 2 trop. Brazil. G. laeve Baill. has offic. bark, cortex Pereirae. Geissostegia Benth. = Erica Tourn. (Eric.). Geitonogamy, pollination from another fl. on same pi. Geitonoplesium A Cunn. Liliaceae (x). 2 E. Austr. Gelasine Herb. Iridaceae (n). 2 temp. S. Am. Geleznowla Turcz. Rutaceae (i). 3 W. Austr. Gelonium Roxb. Euphorbiaceae (A. n. 6). 15 warm As., Afr., Madag. Gelsemium Juss. Loganiaceae. 2 N. Am., As. G. sempervv-ens Ait. (Carolina jasmine) cult. The peduncle bears numerous bracteoles. Gemmate, in pairs. Gemmae, buds, Hymenophyllaceae. Gendarussa Nees=Justicia L. p.p. (Acanth.). General, used in sense opposed to partial. Generations, alternation of, Pteridophyta. Generic name, see Nomenclature. Genetyllis DC. = Darwinia p.p (Myrt.). Genianthus Hook. f. (Secamone p.p. EP.}. Asclepiad. (n. 2). 4 In- domal. Geniculate, bent sharply. Geniosporum Wall, ex Benth. Labiatae (vn). 15 Afr., Madag., Indomal. 286 GENIOSTEMON Geniostemon Engelm. et Gray. Gentianaceae (i). 2 Mexico. Geniostoma Forst. Loganiaceae. 25 Madag. to N.Z. Genip tree (W.I.), Melicocca, &c. Genipa (Tourn.) L. Rubiaceae (i. 8). 2 warm Am., W.I. Genista L. Leguminosae (in. 3). 90 Eur., N. Afr., W. As.; 3 in Brit. G. anglica L. (needle-gorse or petty whin) has large thorns (branches). The fl has an explosive mechanism, typical of many of the fam. (q.v. ). In G. tinctoria L., the dyer's greenweed (M tiller's Fert. of Fls. p. 189), there is no honey; the style and tube of sta. are enclosed in the keel, which is united along the top seam as well as the bottom. The sta. shed their pollen almost in the apex of the keel, but not so near it as to pollinate the stigma. When the fl. opens there is a tension of the sta.-tube on the lower side tending to bend it upwards ; this is resisted by an opposite one in the keel and wings, but if an insect alight on the wings and press them down, the upper seam of the keel gives way and 'an explosion follows. In it the style flies out, striking the under side of the insect, thus probably becoming cross-pollinated, and is followed by a shower of pollen which gives the insect a fresh coating to take to another fl. A yellow dye is obtained from the fls. of this sp. , which when mixed with woad gives a fine green (Kendal green). Genlisea A. St Hil. Lentibulariaceae. 12 trop. Am. and Afr. Gentian, Gentiana. Gentiana Tourn. ex L. Gentianaceae (i). 350 cosmop. exc. Afr., chiefly alpine ; 5 (gentian) in Brit. Most are alpine pi. of tufted growth. Fls. of interest (see Nat. Pfl., Muller'sfert. of Fls., Alpen- blumen, &c. ). The genus shows an ascending series of fls., adapted to higher and higher types of insects. G. lutea L. is a primitive type, with freely exposed honey, yellow homogamous fl. and short- tongued visitors. G. purpurea L. , G. Pneiinwnanthe L. (Brit.), &c. are blue long-tubed humble-bee fls. G. verna L. (Brit.), G. Amarella L. (Brit.) and G. cainpestris L. (Brit.) are long-tubed butterfly fls., sometimes protandr. The gentians form one of the most striking features of the flora of the Alps, occurring in large masses and with very conspicuous fls. ; G. acaulis L. is the most beautiful. In the Brit. Alts, they are rare. The root of G. lutea furnishes a tonic. Gentianaceae (EP., Bff.). Dicotyledons (Sympet. Contortae; Gen- tianales BH.). 80 gen., 800 sp. in every part of the globe and in great variety of situations — arctic and alpine pi., halophytes, sapro- phytes (Voyria, £c. ), marsh pi. (Menyanthes, &c.), water pi. (Lim- nanthemum), &c. They are mostly herbaceous (often perennial) ; a few shrubs. The perennial herbs have usu. a rhizome. L. opp., exstip., usu. entire. The infl. is usu. a dichasial cyme like Caryo- phyllaceae; as in that fam., the lat. branches often become mono- chasial. Other cyniose infls. also occur. Bracts and bracteoles present or not. Fls. reg., $, 4 — g-merous (rarely more). K usu. (5), imbr. ; C (•;), bell- or funnel-shaped, or sometimes salver-shaped, conv. (exc. Bartonia, Obolaria, &c., and § II); A as many as petals, alt. with them, epipet. ; anthers various, usu. introrse ; G with a glandular disc at base, (i), placed in the antero-posterior plane. GERANIA CEAE 2 8 7 Placentae usu. parietal, but they commonly project far into the cavity and spread out at their ends; occasionally the ovary is 2-Ioc. with axile plac. ; ovules usu. oo , anatr.; style simple; stigma simple or 2-lobed. Fr. usu. a septiciclal caps, with oo seeds rarely a berry (Chironia, &c.); seeds small; embryo small, in abundant endosp. The flowers of G. are insect-fertilised. The genus Gentiana has been very fully studied; see also Menyanthes (dimorphic). Classification and chief genera (after Engler): I. GENTIANOIDEAE (1. opp.: C conv. or imbr.) : Exacum, Erythraea, Chlora, Gentiana, Swertia. II. MENYAKTHOIDEAE (1. alt.; C induplicate-valvate) : Menyanthes, Limnanthemum. Gentianales (HH.} The 7th cohort of Gamopetalae. Gentianella Moench.= Gentiana Tourn. p.p. (Gent.). Gentilia A. Chevalier et Beille. Euphorbiaceae (A. I. 2). 2 trop. Afr. Genus, see Nomenclature. Genyorchis Schlechter. Orchidaceae (u. 16). 3 trop. Afr. Geo- (Gr. prefix), earth-; -carpic, producing subterranean fr., Ainphi- curpaeii, Arachis, Cardanritie, Trigonella, Voandzeia ; -graphical distribution, cf. textbooks; -philous, geocarpic ; -tropism, irrita- bility to gravity. Geobaianus Small. Rosaceae (vi). 2 Florida. Geocardia Standley^Geophila D. Don (Rubiac.). Geocarpon Mackenzie. Aizoaceae (n). i Missouri. Geocharis Ridl. Zingiberaceae (l). 2 Malaya. Geochorda Cham, et Schlecht. Scrophulariaceae (n. 6). i warm S. Am. Geococcus J. Drumm. ex Harv. Cruciferae (4). i N.W. Austr. Geodorum Jacks. Orchidaceae (n. 10). 10 Indomal. Geoffraea L. Leguminosae (in. 8). 4 trop. Am. Geoffraya Bonati. Scrophulariaceae (ll. 6). i Cambogia. Geomitra Becc. (Bagnisia p.p. EP.). Burmanniaceae. 2 Borneo. Geonoma Wild. Palmae (iv. i). 85 trop. Am. Geopanax Hemsl. Araliaceae (i). i Seychelles. Geophila D. Don. Rubiaceae (n. 5). 15 trop. Geoprumnon Rydberg (Astragahis p.p. EP.). Leguminosae (in. 6). 6 N. Am. Georchis Lindl. =Goodyera R. Br. (Orchid.). Georgina \Vilkl. = Dahlia Cav. (Comp.). Geosiris Baill. Burmanniaceae. i Madag. Geostachys Ridl. Zingiberaceae (i). 5 Malaya. Geracium Reichb. = Crepis Vaill. p.p. (Comp.). Geraniaceae (EP.\ BH. incl. Oxalidaeeae, Limnanthaceae, Tropaeo- laceae, Balsaminaceae). Dicots. (Archichl. Geraniales EP., BH.}. it gen., 650 sp., cosmop., mostly herbs, often hairy; Sarcocaulon fleshy. Fl. usu. reg. , $, 5-merous. K =,, imbr. with valvate tips, persistent ; C 5, imbi . or conv. ; A as many or 2 or 3 times as many as petals, united at base, obdipl. when > i whorl, anther usu. versatile ; G (5) or (2 — 3) or (3 — 5), with i — 2 or 2 — oo ovules in each on axile plac. ; ovules usu. pend. with ventral raphe and micropyle facing upwards ; style long with 5 stigmas. Fls. usu. protandr. Fr. usu. a 288 GERANIACEAE schizocarp, the cpls. splitting off from a central beak (the persistent style); each takes with it a strip of the tissue of the style, forming an awn, which is usu. hygroscopic (cf. Geranium, Erodium). Embryo straight or folded, in endosp Chief genera : Geranium, Erodium, Pelargonium, Sarcocaulon. [BH. chars. Herbs with alt. or opp. 1. Fl. 5, reg. or •!• . Recept. hardly disc-like, often with 5 glands alt. to C. K 5 or less, C 5 or less, or o, imbr., rarely conv. ; A twice as many as K, often ± connate, G (3 — 5, rarely i), multiloc.; ovules i — 2 per loc., pend. or not, raphe various. Caps., schizocarp, or rarely berry. Enclosp. little or o.] Geraniales. The 23rd order (Engler) of LHcots. (Archichl.). The 7th cohort (BH.) of Polypetalae. Geranium (Tourn.) L. Geraniaceae. 200 cosmop., esp. temp. (12 in Brit., incl. G. pratense L., meadow cranesbill, and G. Robertiatuim L., herb Robert). Infl. cymose, either dich. with cincinnus-tendency (by preference of the /3-bracteole), or a cincinnus alone, which is straightened out into a sympodium. The nectaries are at the base of the sta. These stand at first round the undeveloped style ; after dehiscence they move away, and finally the stigmas open. The fr. explodes, the awn twisting up so that the cpls. are carried up and outwards. In many sp. they open at the same time and the seeds-are shot out. The G. of greenhouses is really a Pelargonium. Geranium (of greenhouses), Pelargonium; -grass, -oil, Cymbopogon. Gerardia L. Scrophulariaceae (in. 2). 40 Am. Gerardiina Engl. Scrophulariaceae (in. 2). i trop. Afr. Gerardiopsis Engl. (Anticharis Endl.). Scrophular. (i. 2). i trop. Afr. Gerascanthus P. Br. = Cordia L. p.p. (Borag.). Gerbera Gronov. Compositae (12). 35 Afr., As. Cult. orn. fl. Germainia Bal. et Poitr. (Anthistiria p.p. BH.). Gram. (2). 2 S.E. As. German pellitory, Anacyclus. Germander, Teucrutm. Germen, the ovary. Germination, cf. Acacia, Cttcurbita, Streptocarpus, Ulex, Vivipary. Gerontogaeous, Old World. Geropogon L. = Tragopogon Tourn. (Comp.). Gerrardanthus Harv. ex Benth. et Hook. f. Cucurbitaceae ( i). 4 trop. Afr. Gerrardiana Oliver. Flacourtiaceae (9) (Samydaceae, BH.). i S. Afr. Gertrudia K. Schum. Flacourtiaceae (3). i New Guinea. Geruma Forsk. Inc. sed. i Arabia. Gesneria L. Gesneriaceae (n). 50 trop. Am., W.I. Gesneriaceae (EP., BH.). Dicots. (Sympet. Tubirlorae ; Personales BH.). 85 gen., 1100 sp. trop. and subtrop., mostly herbaceous or slightly woody ; shrubs and trees are rare. L. usu. opp., rarely whorled or alt., entire or toothed, never divided, exstip. Some are root-climbers, and amongst these are a few epiphytes, e.g. Aeschy- nanthus, usu. with fleshy water-storing 1. Many are tuberous, e.g. the Sinningia (Gloxinia) of greenhouses. A number repr. veg. by means of curious runners or suckers, covered with scale-leaves, usu. GETHYLLIS 289 Floral diagram of Gesneria, after Eichlcr; d= disc-gland. formed below ground, e.g. Nacgelia, Isoloma, &c. A peculiar mor- phology and life-history is found in Streptocarpus (g.v. ). Us- sol. or in cymose infls. of various types, ? , usu. markedly •[• . K (5), us>u. with very stout teeth, generally valvate ; C (5), often 2 -lipped, imhr. (in Kamondia, &c. it is nearly rotate and reg.); A usu. 4, didyna- mous, or 2, or 5 (Ramondia, &c.), alt. with corolla lobes; stds. often found. At the base of the flower-tube is a di*c, whose various shapes form important marks in distinguishing gen. ; it may be ring-shaped (thin or thick), 5-angled, 5-lobed, or re- duced to 5 or fewer glands. G sup., or ± inf. (see below); always (2), i-loc. with parietal plac. which sometimes project inwards so far that it becomes imperfectly 2-loc. ; ovules oo , anatr. ; style simple ; stigma often bilobed. Fr. usu. a caps , splitting loculic. into 2 valves, each of which may again split into 2 ; sometimes (Ramondia) the caps, is septicidal, or opens only at the tip, or the fr. may be ± fleshy or berry-like. Seeds small and numerous, with endosperm (§ II) or without (most of § i). Embryo straight. Fls. mostly protandrous; their large size and bright colours suit them to insects. Saintpaulia (enaceae (4). 5 Malay Archipelago. Gevuina (Gvevina) Molina. Pioteaceae (n). i Chili. Ed. nut. Gherkin, young fr. of Cucumis tativus L. Ghiesbreghtia A. Gray. Scroplmlariaceae (i. i). i Mexico. Ghikaea Volkens et Schweinf. Scrophulariaceae (ill. 2). i trop. Afr. Giant bamboo, Dendroca awns, Giganlochloa; -cactus, Ctreus; -fennel, Ferula ; -pumpkin, Cucnrhita ; -sugar pine, Finns. Gibbesia Small (Siphonychia EP.). Caryophyllaceae (i. 4). i N. Am. Gibbous, with projecting broad pouch. Gidgee (Austr. ), Acacia homatophylla A. Cunn. Giesekia (Gisekia) L. Phytolaccaceae. 5 Afr., India. Gifdoorn (S. Afr.), Sarcocanlon. Gifola Cass. = Filago L. (Comp.). Giganthemum VVelw. (Camoensia EP.}. Leguminosae (in. i). i trop. Afr. Gigantocnloa Kurz. Gramineae (13). 10 Indomal. Giant bamboos (q.v.), used in Java, &c. for building. Gigasiphon Drake del Castillo (Bauliinia p.p.). Leguminosae (n. 4). i Madag. Gigliolia Becc. Palmaceae (iv. i). 2 Borneo. Gigliolia Barb. Rodr. (Octomcria EP.). Orchidaceae (n. 6). 2 Brazil. Gilgia Pax. Euphorbiaceae (A. II. 2). i Somaliland. Gilgiochloa Pilger. Gramineae (9). i E. Afr. Gilia Ruiz et Pav. Polemoniaceae. 120 temp, and subtrop. Am. Gilibertia Ruiz et Pav. Araliaceae (i). 20 trop. Am., As., Japan, Chi. Gill (Am.), Kepeta Glechoma Benth. Gillbeea F. Muell. Cunoniaceae. i N.E. Austr. Gillenia Moench. Rosaceae (i. i). 2 N. Am. Cult. orn. fl. Gilletiella De Wild, et Durand. Acanthaceae (n). i Congo. Gillettia Rendle (Anlhericopsis Engl.). Commelinaceae. i E. Afr. Gilliesia Lindl. Liliaceae (iv). 3 Chili. Cult. orn. fl. Gilliflower. clove, Dianthus Caryophyllus L. ; wall-, wallflower. Gilruthia Ewart. Compositae (4). i W. Austr. Gin, Juiiiperits. Ginalloa Korth. Loranthaceae (il). 4 Indomal. Gingelly, gingili, Sesamum indicum L. Ginger, Zingiber ojfftcinale Rose. Gingidium F. Muell. = Aciphylla Forst. (Umbellif.). Ginginsia DC. = Pharnaceum L. (Aizo.). Ginkgo L. Ginkgoaceae. i, G. biloba L. (Salisburia adiantifolia Sm.), the maidenhair tree, perhaps found wild in W. China, but carefully preserved as sacred in temple gardens. It grows in the open in Eur., reaching 100 feet ; 1. deciduous in autumn, resembling those of maiden- GJELLER UP I A 2 9 1 hair fern, and very often with a deep median division, forked in venation (cf. ferns and cycads), scattered on long shoots, or crowded at the apex of short shoots, which sometimes elongate into long. Below the 1. on the short shoot are a few scale 1. Fls. dioecious, in the axils of the uppermost scales or lowest green 1. on a short shoot (position different from that usual in Coniferae with long and short shoots). 4 a stalked central axis, bearing scattered rather loosely disposed sta., each of which is a slender filament ending in an apical scale and two or more pollen-sacs with longitudinal opening. The pollen grain forms a rudimentary prothallus of a few cells, and the generative nuclei produce two large spirally coiled spermatozoids (cf. cycads). The ? has the form of a long stalk with two term, elliptical ovules enclosed at the base by a collar-like envelope repres. a reduced carpellary 1. Each ov. consists of a nucellus surrounded by one integument, which in the ripe seed forms a thick fleshy aril- like covering round a hard woody shell. In the mature ov. the greater part of the nucellus tissue is reduced to a thin papery layer enclosing a large embryo-sac with usually i archegonia. Fert. occurs before or after the ovule has fallen from the tree. The embryo has 2 cots. The seed is edible, and yields an oil, and the timber is useful. Ginkgo thus represents a very old type, with relationships to the Cycadales and the Filicales. Fossil species are found in the Carboniferous, 1'ermian, Triassic, and Jurassic, and in the Tertiary of England. For details see Coulter and Chamberlain, Morphology of Gymno- sperms; Fujii on floral morphology in Bot. Mag., Tokio, 1895; Seward and Gowan in Ann. Bot. 1900, p. 108 ; Ikeno on fertilisation in Ann. Set. Nat. Bot. xin. 1901, p. 303; Lyon on embryology in Minn. Bot. Stud. in. p. 275. Ginkgoaceae. The only fam. of Ginkgoales, with one gen. Ginkgo (q-v.), formerly placed in Coniferae. Ginkgoales. The 2nd class of Gymnosperms. Ginora L. (Ginoria Jacq.). Lythraceae. 7 Mex., W.I. Ginseng, Aralia Ginseng Baill. Giorgiella De Wild. Passifloraceae. i Congo. Gipsywort, Lycopus mropaeus L. Giraldia Baroni (Atractylis p.p. EP.). Compositae (n). i China. Giraldiella Damm. Liliaceae (v). i China. Girardinia Gaudich. Urticaceae (i). 6 trop. As., Afr. Stinging hairs. Gireoudia Klotzsch — Begonia L. p.p. (Begon.). Girgensohnia Bunge. Chenopodiaceae (B). 4 W. and C. As. Gironniera Gaudich. Ulmaceae. 8 Indomal., Polynesia. Gisekia (Giesekia) L. Phytolaccaceae (Aizoaceae BH.}. 5 trop. Afr. and As. Githago Adans. = Lychnis L. (#//.) = Agrostemma L. (Caryoph.). Githopsis Nutt. Campanulaceae (i). i California. Giulianettia Rolfe. Orchidaceae (n a. in), i New Guinea. Givotia Griff. Euphorbiaceae (A. n. 5). i S. India, Ceylon. Gjellerupia Lauterbach. Opiliaceae. i New Guinea. 19—2 292 GLABER Glaber (Lat.), glabrous, hairless. Glabraria L. = Litsia Lam. (Laur.). Glacial zone, see Zones of Vegetation. Gladdon, Iris foetidissima L. Gladiate, sword -shaped. Gladiolus (Tourn.) L. Iridaceae (in). 120 Afr., Eur., As. protandrous. L. isobil. Cult. orn. fl. Gland, an organ secreting fluid, e.g. nectary, oil-gland, water-pore; cf. Insectivorous PL, Plumbaginaceae, Saxifragaceae ; -ular hair, a sticky secreting hair, Citphea, &c. Glandonia Griseb. Malpighiaceae (n). i Amazon valley. Glans (Lat.), a nut. Glareal, growing on dry exposed ground. Glass-wort, Salsota; (Am., VV.I.) Salicomia. Glastonbury thorn, a var. of hawthorn. Glaucescent, becoming sea-green, or ± sea-green. Glaucidium Sieb. et Zucc. Ranunculaceae (i). 2 Japan, China. Glaucium Tourn. ex Hall. Papaveraceae (n). 12 Eur., As. G.flavum Crantz, yellow horned-poppy, on sea-shores in Brit. Glaucothea O. F. Cook. Palmaceae (i. 2). i California. Glaucous, sea-green, covered with a bloom like a plum. Glaux (Tourn.) L. Primulaceae. G. maritima L., sea milkwort, the only sp., N. temp, coasts (incl. Brit.), a halophyte with fleshy L The seedling dies after producing in the axil of one cot. a hibernating shoot, with a root of its own. From this fresh plants arise veg., the process being repeated for several years before flowering. Runners with scale 1. in whose axils renewal-shoots form appear before the flowering period. The fl. has no C, but a coloured K. Glaziocharis Taub. ex Warm. Burmanniaceae. i Rio de Janeiro. Glaziophyton Franch. (Arundinaria p.p. EP.). Gramin. (13). i Brazil. Glaziostelma Fourn. Asclepiadaceae (n. i). i Brazil. Glaziova Bur. Bignoniaceae (i). i Brazil. Tendrils with discs at tip (cf. Parthenocissus). Gleadovia Gamble et Prain. Orobanchaceae. i Indomal. Glechoma L. (Nepeta L. BH.}. Labiatae (vi). 6 |*. Glechon Spreng. Labiatae (vi). 12 Brazil, Paraguay. Gleditschia Clayton. Leguminosae (n. 7). n subtrop. Am., As. Stems usu. with stout branched thorns (stem structures, arising in 1. axils). The thorn comes from the uppermost of a series of sub- petiolar buds one above the other in the axil. No winter buds form, and the young apex of each twig dies off in winter, the next year's growth starting lat. Some used for hedges; some useful timber. Glehnia F. Schmidt. Umbelliferae (in. 6). i_E. As., W. N.Am. Gleichenia Sm. Gleicheniaceae. 100 trop., %.. Creeping rhiz.; 1. repeatedly branched in an apparently dichot. way. Gleicheniaceae. Filicales Leptosporangiatae. 2 gen., 100 sp., trop., subtrop. and S. temp., small ferns with creeping rhiz. and dichot. branched 1. Sorus of 2—8 sessile sporangia, without indusium, with complete transverse annulus, dehiscing longitudinally. Chief genus : Gleichenia. Gleniea Hook. f. Sapindaceae (i). i Ceylon. GLOSSORHYNCHA 293 Glinus L. (Mollugo L. p.p. BH.}. Aizoaceae (i). 6 trop. and subtrop. Gliricidia H. B. et K. Leguminosae (m. 6). 5 trop. Am., W.I. Glischrocolla A. DC. (Endonema BH.}. Penaeaceae. i S. Afr. Glischrothanmus Pilger. Aizoaceae (i). i Brazil. Globba L. Zingiberaceae (i). 60 Indomal. There is a short K; above this is the C tube, from the end of which spring 3 petals, a large labellum and i stds., also the slightly petaloid fertile sta., projecting beyond which is the style. The ovary is i-loc. with parietal plac. The lower cymes are usu. replaced by bulbils ; the mass of one of these consists of a root, springing lat. from the axis. Globe daisy, Globularia ; -flower, Trollins ; -thistle, Echitwps. Globose, almost spherical. Globularia Tourn. ex L. Globulariaceae. 18 Medit., Ear. Globulariaceae (EP.; Selagineae p.p. BH}. Dicots. (Sympet. Tubi- florae). 3 gen., 20 sp., Eur., Medit., Afr. Herbs or shrubs with alt., exstip., simple 1. and heads or spikes of fls. with or without invol. of bracts. Fl. 5 . K (5), persistent; C (5), median-zygo- morphic ; the upper lip of i petals is shorter than the 3-petalled lower lip; A 4, didynamous, epipetalous ; G i-loc., with i pend. anatr. ov. Fr. a one-seeded nut, free in base of calyx; embryo straight, in endosp. Chief genus : Globularia. Globulea Haw. = Crassula Dill. p.p. (Crassul.). Globulostylis Wernham. Rubiaceae (n. i). -2 Nigeria. Glochidia, Azolla. Glochidion Forst. Euphorbiaceae (A. i. i). 160 trop. As., Polynes. Glochidopleurum Koso-Poljansky. Umbelliferae (in. 5). i Cyprus. Glockeria Nees. Acanthaceae (iv. B). 5 Mexico, C. Am. Gloeocarpus Radlk. Sapindaceae (i). i Phil. Is. Gloeospermum Triana et Planch. Violaceae. 5 trop. Am. Glomera Blume. Orchidaceae (n. a. Hi). 12 Malay Archip., Poly- nesia. Glomerate, collected into heads. Glomeropitcairnia Mez. Broineliaceae (3). i Venezuela, W.I. Glomerule, a cluster of short-stalked fls. Gloriosa L. Liliaceae (i). 5 trop. As., Afr. They climb by aid of the L, whose tips twine like tendrils. Fl. pendulous, with sta. and style projecting horiz. Cult. orn. fl. Glory of the snow, Chiouodoxa. Glosocomia D. Don, Glossocomia Reichb. = Codonopsis Wall. (Cam- pan.). Glossocalyx Benth. Monimiaceae. 3 trop. W. Afr. Glossocardia Cass. Compositae (5). i Indomal. Glossocarya Wall. Verbenaceae (5). 3 Indomal. Glossochilus Nees. Acanthaceae (iv. A), i S. Afr. Glossodia R. Br. Orchidaceae (n. i). 5 Austr. Glossogyne Cass. Compositae (5). 5 Indomal., China. Glossolepis Gilg. Sapindaceae (i). i Cameroons. Glossonema Decne. Asclepiadaceae (n. i). 6 trop. Afr. and As. Glossopetalon A. Gray. Celastraceae. 3 S.W. U.S. Glossopholis Pierre. Menispermaceae. 3 trop. Afr. Glossorhyncha Ridl. Orchidaceae (n. 5). 3 Malay Archipelago. 294 GLOSSOSTELMA Glossostelma Schlechter. Asclepiadaceae (n. i). i Angola. Glossostemon Desf. Sterculiaceae. i Persia. Glossostephanus E. Mey. AsclepiaHaceae (n. i). i S. Afr. Glossostigrna Wight et Am. Scrophulariaceae (n. 6). 3 warm Glossostylis Cham, et Schlecht. =Alectra Thunb. (^//:) Berg. p.p. (Scrophular.). Glossula Lindl. (Habenaria p.p. EP.}. Orchidaceae (n. i). i China. Gloxinia L'Herit. Gesneriaceae (n). 6 trop. Am. G. speciosa Lodd., &.C., often cult. orn. fl., cf. Sinningia. Glucose, grape sugar. Glumaceae (Bff.). The 7th series of Monocotyledons. Glumaceous, glume-like. Glumicalyx Hiern. Scrophulariaceae (in. i). i S. Afr. Glumiflorae. The 4th order (Engler) or 2nd (Warming) of Monocoty- ledons. Gluta L. Anacardiaceae (i). 5 trop. As., Madag. The sap of G. Renghas L. yields a good varnish. Glyceria R. Br. Gramineae (10). 20 cosmop., esp. N. Am. 2 in Brit. Pasture grasses in wet meadows. Glycine L. (incl. Soja Moench.). Leguminosae (m. 10). 16 palaeo- trop. G. Sofa Sieb. et Zucc. and G. hispida Maxim, yield Soja beans, eaten in Japan, &c., and used as green fodder. An oil is obtained from the seeds. Glycosmis Correa. Rutaceae (v). 6 Indomal. Fr. ed. Glycyderas Cass. Compositae (3). i Madag. Glycyrrhiza Tourn. ex L. Leguminosae (in. 6). 12 temp, and sub- trop. An extract of the rhiz. of G. glabra L. is Spanish liquorice. Glyphaea Hook. f. Tiliaceae. 2 trop. Afr. Glyphosperma S. Wats- Liliaceae (in). I North Mexico. Glyptopetalum Thw. Celastraceae. 4 Indomal. Glyptopleura Eaton. Compositae (13). 2 Utah to California. Glyptostrobus Endl. (Taxodimn p.p. Bff.). Coniferae (Pinaceae; see C. for gen. char.). 2 China, G. pcndulus Endl. and G. hetero- phyllus Endl. Gmeiina L. Verbenaceae (4). 8 Indomal. Gnaphalium L. Compositae (4). 135 cosmop., 4 in Brit, (cudweed). G. snpiniiin L., alpine in Scotland, is a tufted hairy xero. [G. dioi- cum L. = Antennaria, G. Leontof>odium L. =Leontopodium.] Guaplialodes A. Gray. Compositae (4). 3 temp. Austr. Gnephosis Cass. Compositae (4). 15 temp. Austr. Gnetaceae. The onlyfam. of Gnetales, comprising 3 very distinct gen. with about 45 sp. trop. and subtrop. They are distinguished from the Coniferae by the absence of resin, by the presence of vessels in the secondary wood, and by the occurrence of a P. Classification and genera (after Engler) : I. EPHEDROIDEAE (sate, exstip., simple, evergr., leathery. Fls. dioec., in spikes which are frequently grouped into more com- plex infls. The spike bears decussate bracts, in wh< se axils are condensed partial inrls. of a large number of flrs. (cf. Labiatae), about 3 — 8 in the ?, but more (up to 40) in the sol. in the leaf-axils or in cymes, 298 GO ODENIA CEAE racemes, or spikes. K usu. 5, small; C (5) ; A 5, alt. with the petals, epipet. or not, with introrse sometimes syngenesious anthers; G (2), inf. or semi-inf. i- or 2-loc.; ovules i, 2, or oo in each usually ascending, anatr. ; style simple with 'pollen-cup' close under the stigma. Into this the pollen is shed in the bud; it then closes up, leaving only a narrow opening. The style bends down to stand in the mouth of the almost horizontal fl., so that insect- visitors come in contact with the cup and dust themselves with a little of the powdery pollen. As the sti^matic lobes grow up in the cup they keep forcing fresh pollen into the narrow slit, and finally emerge by it themselves and then receive the pollen of younger fls. from insect-visitors. The mechanism should be carefully compared with that of Campanulaceae and Compositae. Fr. usu. caps., sometimes a nut or drupe. Embryo straight, in fleshy endosp. The G. are very closely allied to Campanulaceae, differing chiefly in the absence of latex and the presence of the pollen-cup. They resemble Gentianaceae in a few points. C kief genera: Goodenia, Leschenaultia, Scaevola, Dampiera. [BH. chars, incl. those of Brunoniaceae. ] Goodenovieae (BH. ) = Goodeniaceae. Goodia Salisb. Leguminosae (ill. 3). 2 S. Austr. Good King Henry, Chenopodium Bonus- Henricns L. Goodyera R. Br. Orchidaceae (n. 2). 40 N. temp., trop. As., New Caled., Mascarenes ; G. repens R. Br. in Brit. Fl. as in Epipactis (Darwin, Orchids, p. 103). Gooringia Williams (Arenaria p.p.). Carophyll. (i. i). i Tibet. Gooseberry, Ribes Grossularia L. ; American- (W.I.), Heterotrichum ; Barbadoes- (W.I.), Peireskia aadeata Mill.; Cape-, Physalis\ Otaheite-, ditto; -tomato, ditto. Goose-foot, Chenopodium ; -grass, Ga/ium, (Am.) Eleusine indica. Gorceixia Baker. Compositae (i). i Rio de Janeiro. Gordonia Ellis. Theaceae. 20 Indomal., China, N.Am. Seeds winged. The bark of G. Lasianthus L. (loblolly-bay, S. U.S.) is employed for tanning. Sta. opp. to petals. Gorgoglossum F. C. Lehm. Orchidaceae (n. 13). i Colombia. Gorgonidium Schott. Araceae (vn). i Malay Archipelago. Gormania Britton ex Britton et Rose. Crassulaceae. 9 W. N.Am. Gorse, Ulex; needle-, Genista. Gorteria L. p.p. (p.p. -Berkheya Ehrh.). Compositae (to). 4 S. Afr. Gosela Choisy. Scrophulariaceae (n 7). i S. Afr. Gossweilera Sp. Moore. Compositae (i). i Portuguese W. Afr. Gossypianthus Hook. Amarantaceae (3). 2 Texas, Mexico. Gossypium L. Malvaceae (4). 12 trop. and subtrop. Epicalyx of 3 1. G (5). Loculic. caps. The seeds are covered with long hairs forming the material known as cotton (q.v.). The cult, forms are apparently reducible to 3 sp. , G. barbadense L. (trop. Am.), G. arbo- reum L. (Old World), and G. herbaceum L. (ditto). The cotton separates easily from the seed in the first sp., which is the Sea Island cotton of the U.S. ; in Egypt, India, &c. the other sp. are most used. From the seeds an oil is obtained by crushing (cotton-seed oil), and GRAMINEAE 299 the oil-cake left behind is largely used for feeding cattle, &c. The fls. are visited by bees and (in Am.) by humming-birds. Gothofreda Vent. = Oxypetalum R. Br. (Asclep. ). Gouania Jacq. Rhamnaceae. 45 trop. and subtrop. Some have watch-spring tendrils. The stalks of some sp. contain saponin. Goughia Wight = Daphniphyllum Blume (Euphorb. ). Goulardia Husnot (Agropyrum p.p.). Gramineae (12). 2 Europe. Gouldia A. Gray. Rubiaceae (i. 7). 5 Hawaiian Is. Goupia Aubl. Celastraceae. 2 Guiana. Gourd, Cucurbita; bitter-, Citrullus; bottle-, Lagenaria; snake-, Trichosanthes. Gourliea Gillies ex Hook. Leguminosae (in. i). i temp. S. Am. Pod ed. Goutweed, Aegopodium Podagraria L. Govenia Lindl. Orchidaceae (n. 10). 10 Brazil to W.I. Gowan, daisy. Goyazia Taub. Gesneriaceae. i Brazil. Grabowskia Schlechtd. Solanaceae (2). 6 S. Am. Gracilis (Lat), slender. Graderia Benth. Scrophulariaceae (in. 2). 3 Afr. Graeifea Seem. Tiliaceae. i Fiji. Graellsia Boiss. Cruciferae(2). i Persia. Graffenrieda DC. Melastomaceae (i). 18 trop. S. Am., W.I. Graft-hybrid, Cytisus. Grahamia Gill. Portulacaceae. i temp. S. Am. Graines d'Avignon, Rhamnus; grains of Paradise, Amomum. Gram, Cicer ; black and green, Phaseolns ; horse, Dolichos. Grama or gramma grass, Boutelotia. Gramerium Desv. (Panicum p.p. EP.). Gramineae (5). i Chili. Gramineae (EP., BH.}. Monocots. (Glumiflorae; Glumaceae BH.}. One of the largest orders of fig. pi., with about 350 gen. and 4000 sp. in all regions of the globe. In the temp, zones esp. they are a most important feature in the veg., forming prairies, steppes, &c. Most grasses are herbaceous with fibrous root, but a few, chiefly the bamboos (g.v.), reach a large size, even as much as 100 ft. Many are annual, but many perennial ; the latter commonly branch largely from their lower nodes and thus often give rise to a tufted habit (as seen in many common sp.) ; many possess rhiz. The stem has well-marked nodes, composed chiefly of softer tissues. If a stem be bent downwards (as occurs when wheat is Maid') these nodes recommence growth, growing more rapidly upon the lower side, so that the stem is once more brought to the vertical position. The stem is usu. hollow (exc. Zea, Saccharum, &c.) and circular in section. The 1. are alt., and with few exceptions, in 2-ranked phyllotaxy; they have a sheathing base, the edges of the sheath overlapping one another upon the side of the stem opp. to the blade (cf. Cyperaceae) ; there is no petiole (exc. in a few bamboos, &c.), and at the junction of blade and sheath there is a little membranous outgrowth, the ligule, upon the upper side of the leaf. The blade is usu. linear. Many xero. grasses have grooves along the upper side of the 1., with the stomata at the bases of the grooves; in most of these cases the 300 GRAMINEAE 1. rolls up upwards in dry air, enclosing the stomata completely and checking transpiration ; the lower surface, which thus becomes the outer, is covered with thick-walled cells and has no stomata. When the air again becomes moist the 1 unrolls. The infl. is complex; the unit of infl. is not the individual fl. but a spikelet, or small spike of fls. These are well seen in oats, where they are arranged in a panicle; in wheat the spikelets are sessile upon the main axis, forming a cpd. spike, usu. termed simply a spike ; this is more clearly seen in rye-grass. Each of these spikelets consists of one or more (usu. not more than 5) fls. one above the other on opp. sides of a very short axis, the whole enclosed in one or two or more larger leaves at the base of the axis, the inf. palea - inf. palea - - inf. palea - inf. palea - outer glume glumes. The first diagram gives a rough representation of the construction of a spikelet. The central line represents the axis (supposed elongated), which bears at the base usu. two leaves with nothing inner glume -j in their axils; these are the glumes Above them stand one or more leaves, the inferior paleae, in whose axils occur fls. (sometimes aborted). Upon the axis of the fl., opp. to the inf. palea, stands another leaf, the superior palea. Between the paleae the fl. itself is enclosed. The sup. palea, being upon the same axis as the fl., is evidently its bracteole. Owing to the short- ness of the floral axis, it appears to be in the axil of the inf. palea, or even upon the main axis of the spikelet. Above the sup. palea are two very small scales, the lodicules; they are opp. to the sup. palea (1 in fig.) and insignificant in size. They are sometimes supposed to repres. two of the three 1. of a reduced P, but it seems more probable that they really repres. a second bracteole and that the fl. is perfectly naked. The fl. itself has usu. 3 sta. with long filaments and versatile anthers, and i cpl. forming a i-loc. ovary, with i basal erect anatr. ovule, and i or more (usu. 2) stigmas, much branched. This ovary has some- times been regarded as formed of > i cpl., but the suture of the post, side of the ovary (cf. Prunus) seems to point clearly to its being of i cpl. only. The two stigmas may then be regarded as developments of the lat. parts of the cpl. whilst the central part (which usu. forms the stigma) remains undeveloped ; this also is the explanation of the two lodicules placed right and left of the proper position for an upper bracteole. All the fls. in the spikelet are enclosed as a rule within the glumes until ready to open. Then the glumes separate, and the hygroscopic lodicules force apart the paleae of the fls. Most sp. in Eur. are anemoph. and protog. ; the sta. grow very rapidly in b Floral diagram of a grass (after Eichler). b, inferior palea; v, superior palea ; 1, lodicule ; St., stigma. GRAMINEAE 301 warm weather and suspend the anthers clear of the paleae so that the loose powdery pollen is easily blown away and may be caught by the large stigma of another fl. The fruit is a caryopsis, i.e. an achene whose pericarp is com- pletely united to the seed-coat. Its construction can be well seen in maize (or wheat) ; at the broad end is the scar of the style, and on the under side at the pointed end is the embryo; on the upper side is the hilnm or point where the ovule was attached to the wall of the cpl. (the form of this scar is important in classification) ; the bulk of the seed consists of floury endosp. The embryo is straight, with its one cot. {scufellitm} completely enwrapping the radicle and plumule (this can be easily made out by dissecting soaked material). The radicle is towards the lower end of the fr. In germin. the cot. remains within the seed and extracts nourishment from the endo- sperm ; afterwards it withers away. Most grass frs. are sufficiently light to be dispersed by wind, esp. as the paleae often remain attached to them and become dry and chaffy. Others have hooks. Self-burying arrangement in Stipa (g.v.), effected by aid of the awn (this term is applied to any long thread-like outgrowth of glume or palea). Many grasses, e.g. sp. of Poa and Festuca, are viviparous, esp. on mountains. The spikelets are replaced by leafy shoots with adv. roots at their bases. These drop oft" and grow upon the soil (cf. Agave, Allium, &c.). From the economic point of view the G. are only rivalled in importance, if at all, by the Palmae and Leguminosae. The cereal grasses, e.g. Oryza, Triticum, Zea, Avena, Hordeum, &c. afford food to a large proportion of the earth's inhabitants (see Edible Products). Many grasses are valuable as fodder (q.v.} for domestic animals, or for hay. The bamboos (q.v ) supply many of the wants of the natives of trop. countries, Japan, &c. Many are used as sandbinders, &c. Classification and chief genera (after Hacke!) : the G. show near relationship only to Cyperaceae and perhaps Juncaceae, and are easily distinguished from these either by their veg. or floral characters. A. Spikelets i-flowered without elongation of the axis beyond the fl., or 2-flowered with the lower fl. imperfect; without measurable internode between the individual glumes or paleae, and when ripe fatting off from the stalk as a whole or together with certain parts of the axis of the spike, a. Hilum point-like; spikelets not compressed lat., but usually dorsally compressed or cylindrical. 1. Maydeae (inf. palea and, when present, sup. palea thin and membranous ; glumes firm, even leathery or cartilaginous, the lowest one the largest and overlapping the rest ; spike- lets usu. in racemes or spikes which become jointed when ripe; $ and ¥ spikelets in separate infls. or in separate parts of the same infl.); Euchlaena, Zea, Coix. 2. Andropogoneae (as I, but spikelets 5, or i and ? side by side in the same infl.): Saccharum, Andropogon, Sorghum. 3. Zoysieac (paleae membranous; glumes herbaceous, papery or leathery, the lowest usu. the largest ; spikelets falling singly or in groups from an unjointed spike-axis) : Zoysia. 3o2 GRAMINEAE 4. Tristegineae (paleae membranous ; glumes herbaceous or firm and papery, the lowest smaller or narrower than the rest; spikelets falling singly from the twigs of a panicle) : Arundinella. 5. Paniceae (paleae usu. cartilaginous, leathery or papery ; glumes more delicate, usu. herbaceous, the lowest usu. smaller ; spikelets falling singly from the twigs of a panicle or unjointed spike-axis) : Paspalum, Panicum, Setaria, Cenchrus, Pennisetum, Spinifex. b. Hilum linear ; spikelets lat. compressed. 6. Oryzeae: Zizania, Oryza, Lygeum, Leersia. B. Spikelets i — many-flowered; when i -flowered often with a pro- longation of the axis above the fl., their stalks usu. jointed above the glumes, so that they fall off leaving these on the axis ; when i- or more-flowered, always with distinct inter- nodes between the fls. a. Stem herbaceous annual. No petiole, or joint between blade and sheath. 7. Phalarideae (spikelets in panicles, spikelike panicles or racemes on distinct stalks — sometimes very short, not set in notches on the main axis; i -flowered, with 4 glumes and i -nerved sup. palea): Phalaris, Anthoxanthum. 8. Agrostideae (as 7, but spikelets with two or no glumes and 2-nerved sup. palea) : Aristida, Stipa. Phleum, Alopecurus, Sporobolus, Polypogon, Agrostis, Calamagrostis, Ammo- phila, Apera, Lagurus. 9. Aveneae (infl. as 7, but spikelets 2- — many-flowered ; inf. paleae usu. shorter than glumes, with twisted awn upon the dorsal side, more rarely with no awn or with an awn upon the tip as 10, but then always with 2 almost opp. fls. and no prolongation of the axis beyond them) : Holcus, Aira, Corynephorus, Deschampsia, Trisetum, Avena, Arrhenatherum, Danthonia. 10. Festuceae (as 9, but int. palea usu. longer than glume, without awn or with untwisted awn at tip) : Sesleiia, Gynerium, Arundo, Phiagmites, Eragrostis, Catabrosa, Melica, Briza, Dactylis, Cynosurus, Poa, Glyceria, Festuca, Bromus, Brachypodium. 11. Chlorideae (spikelets in 2 rows approximated to one another, forming a one-sided spike or raceme with un- jointed axis) : Cynodon, Spartina, Chloris, Bouteloua, Eleusine. 12. Hordeeae (spikelets in 2, or rarely more, opp. rows, forming a symmetrical, rarely one-sided, spike) : Nardus, Lolium, Agropyrum, Secale, Triticum, Hordeum, Elymus. b. Stem woody, at least below ; leaf often petiolate, finally separating from its sheath by a joint. 13. Bambuseae: Arundinaria, Bambusa, Dendrocalamus. Space will not permit of giving a key to the genera; the student should work through the commoner ones with a flora, or with Ward or Hutchinson's British Grasses. GRASS 3°3 Gramineous, relating to grasses, grass-like. Grammadenia Benth. Myrsinaceae (n). 10 trop. Am., W.I. Grammangis Reichb. f. Orchidaceae (n. 17). 2 Madag., Java. Granimanthes DC. Crassulaceae. i S. Afr. Grammatocarpus Presl (Scyphanthus EP.}. Loasaceae. 2 Chili. Grammatophyllum Blume. Orchidaceae (n. 17). 5 Malaya. G. spe- ciosum Bl. is about the largest known orchid. Grammatotheca C. Presl (Lobelia p.p. BH.}. Campanulaceae (ill), i S. Afr., i Austr. Grammica Lour. =Cuscuta L. (Convolv. ). Grammitis Sw. = Polypodium L. Grammosciadium DC. (Chaerophyllum p.p. BH.}. Umbelliferae (in. 8). 8 E. Medit. GranadiUa (Tour.) Rupp. =Passiflora L. p.p. (Passifl.). Granadilla, Passiflora quadrangularis L. ; -tree (W.I.), Brya. Grandidiera Jaub. Flacourtiaceae (2). i Zanzibar. Grandiflorus (Lat.), large fld.; grandis (Lat.), large. Grangea Adans. Compositae (3). 2 palaeolrop. Grangeria Comm. ex Juss. Rosaceae (vi). 3 Madag., Mauritius, Sun das. Grantia Boiss. Compositae (4). 5 E. Medit. Grape, Vitis vinifera L., &c.; -fruit, Citrus decumana Murr., var. ; -hyacinth, Muscari; seaside-, -tree, wild- (W.I.), Coccoloba. Graphephorum Desv. Gramineae (10). i N. Am. Graphistemma Champ, ex Benth. et Hook. f. Asclep. (n. i). i China. Grapple plant, Harpagophytum. Graptopetalum Rose. Crassulaceae. i Mexico. Graptophyllum Nees. Acanthaceae (iv. B). 5 Austr., Polynesia. G. horteme Nees (G. piclum Griff.) cult, for its prettily marked I. Grass, one of the Gramineae ; alkali-, Distichlis; arrow-, Triglochin\ barley-, Hordeum ; barnyard-, Panicum ; beard-, Polypogon ; bent-, Agrostis, Apera ; Bermuda-, Cynodon ; blue-, see Kentucky blue; brome-, Browns; buffalo-, Biichloe ; canary-, Phalaris; cat's tail-, Phleitm ; China-, Boehmeria; citronella-, Cymbopogon ; cock's foot, Dae ty Its ; cord-, Spartina ; cotton-, Eriophoritm ; couch-, Agropyron; crab- (Am.), Panicum ; darnel-, Lolium ; ditch- (Am.), Rtippia; dog's tail-, Cynosurus; dog's tooth-, Cynodon; eel-, Zostera ; esparto-, Lygeum, Slipa ; false brome-, Brachy podium ; false oat-, Arrhenathcrum • feather-, Stipa ; fescue-, Festttca ; florin-, Agrostis ; fowl-meadow- (Am.), Poa, Glyceria ; fox-tail, Alopecurus ; French rye, Arrhenatkerum ; gama-, Tripsacum ; goose-, Galium ; Geranium-, Andropogon ; grama-, Tripsacum ; Guinea-, Panicum; hair-, Deschampsia, Aira; hedgehog- (Am.), Cenchrns; herd's- (Am.), Phleum; holy- (Am.), Hierochloe; Indian- (Am.), Sorghum ; kangaroo-, Anthistiria ; Kentucky blue, Poa pra- tensis L. ; knot-, Polyqonum, Agropyron repens Beauv., (Am.) Paspa- lum distichum L. ; lemon-, Cymbopogon ; lyme-, Elymus ; marram-, Ammophila; mat-, Nardus ; Mauritius-, Panicum; meadow-, Poa ; melic-, Melica ; mesquite- (Am.), Bouteloua ; millet-, Milittm ; oat-, Avena; -of Parnassus, Pamassia; Pampas-, Cortaderia; 304 GRASS panic-. Panicum; quake-, Briza; reed-, Pkalaris; Eusa-, Cymbo- pogon; rush- (Am.), Vilfa\ rye-, Lolium; scorpion-, Myosotis ; scurvy-, Cochlearia; soft-, Holcus; sparrow-, Aspa>-agus\ sweet vernal-, AnthoAanthurn; timothy-, P/iletim : -tree, Xanthorrhoea ; twitch-, Agropyron; wheat-, A^ropyron; white bent-, Agrostis\ whitlow-, Draba ; -wrack, Zostera. Grass-cloth, Chinese, Boehmeria ; -hemp (Austr.), Agave rigida Mill. Gratiola (Rupp.). L. Scrophulariaceae (in. 6). 25 cosmop. Sta. 2. The dried plant of G. officinalis L. was formerly offic. Gratwickia F. Muell. Compositae (4). i Austr. Graveolens (Lat.), strongly scented. Gravesia Naud. Melastomaceae (i). 10 Madag. Cult. orn. fl. Crossed with Cassebeeria they give the fancy 'Bertolonias.' Gravisia Mez (Aechnua p.p. BH.}. Bromeliaceae (4). 4 trop. Am., W.I. Grayia Hook, et Arn. Chenopodiaceae (A). 2 \V. U.S. Greater celandine, Chelidoniiim majus L. Greek valerian (Am.), Polemonium. Green brier (Am.), Smilax; -gram, Phaseolus Mungo L. ; -heart, Nectatidra Rodiaei Hook. ; Kendal-, Genista ; -manure, Legumi- nosae; -weed, dyer's, Genista; -withe (W.I.), Vanilla claviculata Sw. Greenea Wight et Am. Rubiaceae (i. 3). 6 E. Indomal. Greenella A. Gray. Compositae (3). 3 S.W. U.S. Greeneocharis Guerke et Harms. Boraginaceae (iv. 2). 2 Pac. N. Am. Greeniopsis Merrill. Rubiaceae (i. 5). 2 Phil. Is. Greenmania Hieron. Compositae (5). i Colombia. Gregarious, growing in company, all of one sp. Greggia A. Gray. Cruciferae (4). 4 W. Am. Greigia Regel. Bromeliaceae (4). S. Am. Cult. orn. infl. Grenacheria Mez. Myrsinaceae (n). 6 Malaya. Grenadilla, Pas si flora quadrangularis L. Greslania Balansa. Gramineae (13)- 3 New Caledonia. Grevea Baill. Saxifragaceae (v). i Madag. Grevellina Baill. (Turraea p.p. EP.). Meliaceae (ill), i Madag. Grevia L. = Grewia L. (Tiliaceae). Grevillea R. Br. Proteaceae (n). 170 Austr. Trees and shrubs with racemose infls., 2 fls. in each axil. The style projects from the bud as a long loop, the stigma being held by the P until the pollen is shed upon it. Then the style straightens out, and the pollen may be removed ; presently the female stage supervenes. Some yield useful timber, and G. robnsta A. Cunn. (silky oak) and other sp. are now extensively employed as shade and timber trees in Ceylon and elsewhere. Grewia L. Tiliaceae. 120 As., Afr., Austr., esp. trop. Grewiella O. Ktze. (Grewiopsis EP.}. Tiliaceae. 2 trop. Afr. Grewiopsis De Wild, et Durand. Tiliaceae. a trop. Afr. Greyia Hook, et Harv. Melianthaceae. 3 S. Afr. Grias L. Lecythidaceae. 4 S. Am., W. Ind. G. cauliflora L., an- chovy pear, cult, in the W.I. GRUSONIA 305 Grlelum L. Rosaceae (iv). 28. Afr. Grifflanthus Merrill (Grijfithia Maingay). Anonaceae (i). 3 Phil. Is. Griffinia Ker-Gawl. Amaryllidaceae (i). 8 Brazil. Griffithella Warming. Podostemaceae. i VV. Ghats of India. Plants with the general veg. structure of Dicraea, but remarkable for the extraordinary polymorphism of their shoots, which may be cup or wineglass shaped, creeping or erect, and of many different forms, shapes, and sizes (cf. Willis, Ann. Perad., I., 1902, p. 364). They are of root nature. Griffithia J. M. Black. Compositae (4). i S. Austr. Grifflthia Maingay. Anonaceae (i). 3 Malaya. Griffitbia Wight et Arn. = Randia Houst. (Rubiac.). Grimthianthus Merrill. Anonaceae (i). 3 Phil. Is. Griffonia Bail I. (Bandeiraea Welw.). Legurn. (n. 4). 3 W. trop. Afr. Griffonia Hook. f. (Acioa Aubl.). Rosaceae (vi). 4 trop. Afr. Grigri palm (W.I.), Martinezia corallina Mart. Grimmeodendron Urb. Euphoibiaceae (A. n. 7). 2 W. Ind. Grindelia Wuld. Compositae (3). 35 Am. GrisebacMa Klotzsch (Eremia Don). Ericaceae (iv. 2). 25 S. Afr. Grisebachiella Lorentz. Apocynaceae (n. ij. i Argentina. Griselinia Forst. f. Cornaceae. 8 N.Z., S. Am. Griselinia Scop. = Pterocai pus L. (Legum.). Grisia Brongn. = Bikkia Reinw. p.p. (Rubi.). Grislea L. Lythraceae. i trop. S. Am. Grisollea Baill. Icacinaceae. 2 Madag., Seychelles. Grobya Lindl. Orchidaceae (n. 17). 2 Brazil. Gromwell, Lithospermum, Mertensia. Grona Lour. Leguminosae (in. 10). 4 trop. As. Gronophyllum Scheff. Palmaceae (iv. i). 2 Malaya. Gronovia Houst. ex L. Loasaceae. 2 trop. Am. A 5, no stds. G i. Grosourdya Reichb. f. Orchidaceae (n. 20). 3 Malay Archipelago. Grossera Pax. Euphorbiaceae (A. n. 2). 2 trop. Afr. Grossularia Tourn. ex Adans. (Kibes p.p.). Saxifrag. (vi). 40 N. Am. Ground-cherry (Am.), Physalis ; -hemlock (Am.), Taxus; -ivy, Nepeta Glechoma Benth. ; -laurel (Am.), Epigaea ; -nut, Arachis, Voandztia, (Am.) Afios\ -pine, Ajuga Chamnepitys Schreb., (Am.) Lycopodium dcndroitieum Michx. ; -pink (Am.), Phlox subitlala L. Groundsel, Senecio vulgaiis L.; -tree (Am.), Baccharis. Growth, permanent change of form ; for special cases see Albizzia, Bamboos, Dendrocalainus, Dracaena, Eucalyptus, Sequoia. Grubbia Berg. Grubbiaceae. 3 S. Afr. Woody pi. with opp. leathery 1. and small 5 reg. fls. P 4, A 4 + 4, G (2), 2-loc. below while young, later i-loc. with 2 pend. orthotr. ov. on central plac. Drupe. Oily enclosp. Grubbiaceae (EP.; Sanialaceae p.p. BH.}. Dicots. (Archichl. Santa- lales). Only gen. Grubbia, q.v. Gruinalis (Lat.), crane's bill shaped. Grumilea Gaertn. (Psychotria p.p. BH.}. Rubiaceae (n. 5). 45 |#. Grundelia L. 5 misprint for Gundelia. Grusonia Hort. Nicolai ex K. Schum. = Opuntia Tourn. (Cact.). W. 20 3o 6 GR YPO CARPHA Grypocarpha Greenman. Compositae (5). i Mexico. Guaco, Mikania amara Willd. Guadua Kunth (Bambusa p.p. EP.). Gramineac (13). 15 trop. Am. Guaduella Kranch. Gramineae (13). 2 trop. Afr. Guaiacum Plum, ex L. Zygophyllaceae. 4 N. and trop. Am. G. ojficinale L. yields lignum- vitae wood, from which is also obtained the medicinal resin guaiacum. Guamatela J. D. Smith. Rosaceae (in. 2). i C. Am. Guamia Merrill. Anonaceae (i). i Phil. Is. Guapira An hi. Verbenaceae (inc. sed.). i Guiana. Guapuronga, Marlierta. Guar, Cyaniopsis. Guarana, Paullinia Cttpana H. B. et K. Guardiola Cerv. ex Humb. et Bonpl. Compositae (5). 5 S.W. U.S., Mexico. Guarea Allem. ex L. Meliaceae (in). 100 trop. Am., Afr. The disc forms a gvnophore, and the sta. are completely united into a tube. Guatteria Ruiz et Pav. Anonaceae (i). 60 trop. Am. Berry stalked. Guava, Psidnim ; black- (W.I.), Gncttarda argentea Lam. Guayule, rarllicnium argentatum A. Gray. Guaza, ganja. Guazuma Plum, ex Aclans. Sterculiaceae. 5 trop. Am. Gueldenstaedtia Fisch. Leguminosae (in. 6). 10 Centr. As., China. Guelder rose, Viburnum Opiilus L. Guepinia Bast. = Teesdalia R. Br. (Cruc.). Guerkea K. Schum. Apocynaceae (n. i). 2 trop. Afr. Guernsey lily, Nerine. Guettarda L. Rubiaceae (n. 2). 50 trop. all but i (G. speciosa L. , which is common on trop. coasts) Am. Guevina (Geiwina) Molina. Proteaceae (ll). i Chili. Ed. nut. Guichenotia J . Gay. Sterculiaceae. 5 W Austr. Guiera Adans. Combretaceae. i trop. Afr. Guilandina L. = Caesalpinia L. (Legum.). Guild, a group of pi resembling one another ecologically. Guilielma Mart. = Bactris Jacq. p.p. (Palm.). GuillainiaVieill. (Alpinia p.p. EP.}. Zingiberaceae(l). 2 W. Polynes. Guilleminea H. B. et K. Amarantaceae (3). 3 warm Am. Guillenia Greene. Cruciferae (r). 6 W.N. Am. Guillonea Coss. Umbelliferae (in. 7). 2 Spain. Guimauve, marsh mallow. Guinea corn, Sorghum vulgare Pers. ; -grains, Amomnm Melegueta Rose.; -grass, Paniciim maximum Jacq.; -hen's weed (W.I.), Petiveria. Guioa Cav. (Cnpam'a p.p. BH.}. Sapindaceae (i). 35 Indomal. Guiraoa Coss. Cruciferae (2). i Spain. Guizotia Cass. Compositae (5). 8 Air. G. abyssinica Cass. (rantil or niger-seed) is cult, in India, &c, for its seeds, from which an oil is expressed. Gulubia Becc. (Kentia p.p. EP.). Palmaceae (iv. i). 2 New Guinea. Gum, products of disintegration of internal tissues, exuding from stems, &c., swelling or dissolving in water, insoluble in alcohol or ether; GUTTIFERAE 307 common in dry countries. Three chief classes: arabin (fully soluble), type gum-arabic (Acacia Senegal Willd.), bassorin (slightly soluble), type gum-tragacanth (Astragalus gummifer Labill.) and cerasin (swelling), type cherry-gum (Prunits Cerastes L.). Cf. above, and Ckloroxylon, Feronia, Piptadenia, &c. ; -ammoniacum, Dorema ; Angico-, Piptadenia ; -arabic, Acacia Senegal Willd. ; -benzoin, Styrax; blue-, Eucalyptus; chewing-, Achras Sapota L. ; Chicle-, ditto; -cistus, Cist us; -copal, cf. Copal; doctor's- (W.I.), Rhus Metopinm L. ; -elemi tree (W.I.), Dacryodes hexandra Griseb. ; -euphorbium, Euphorbia resmifera Berg. (Morocco) ; -galbanum, Ferula ; -guaiacum, Guaiacttm ; -kino, cf. Kino ; -lac, cf. Lac ; -ladanum, Cistus ; -olibanum, Boswellia ; -opopanax, Opopanax ; -plant (Am.) Grindelia ; -resins, resins containing a mixture of gum and resin, Boswellia (frankincense), Calopliyllum, Convolvulus, Dorema, Ferula, Garcinia (gamboge) ; -sandaracb., Tetraclinis • sweet-, Liquidambar ; -tragacanth, Astragalus ; -tree, Nyssa,(W.I.) Sapiitm . Gumbo, Hibiscus esculentus L. Gumillea Ruiz et Pav. Cunoniaceae. i Peru. L. alt. Gundelia (Tourn.) L. Compositae (10). i As. Min., Persia. Gundlachia A. Gray. Compositae (3). i Cuba, St. Domingo. Gundlea Steud. = Grumilea Gaertn. (Rubi.). Gunnera L. Haloragidaceae. 18 S. Am., Costa Rica, N.Z., Tasm., Sandw. Is., Java, S. Afr. Several are enormous herbs with 1. several feet across. PI. 2-merous. G i-loc. Stem polystelic. In the leaf- axils are ' squamulae intravaginales ' (cf. Potamogetonaceae). Cult, orn. fol. Gunnia F. Muell. Aizoaceae (n). 2 S. and W. Austr. Gunniopsis Pax (Aizoon p.p.)- Aizoaceae (n). i S.E. Austr. Gunny, Corchorus. Gunpowder plant, Pilea. Gurania Cogn. Cucurbitaceae (2). 55 trop. Am. Guraniopsis Cogn. Cucurbitaceae (2). i Peru. Gurjun balsam, Dipterocarpus. Gussonea A. Kich. = Angraecum Bory p.p. (BH.) = Mystacidium Lindl. p.p. Gustavia L. (Japarandiba Adans.). Lecythidaceae. 20 trop. Am. G. augusta L. is the stink-wood; the wood has a foetid smell. Gutenbergia Sch. Bip. Compositae (i). 10 trop. Afr. Guthriea Bolus. Achariaceae. i S. Afr. Gutierrezia Lag. Compositae (3). 20 Am. Gutta-jelutong, Dyera ; -percha, a substance like rubber, but softening with heat, in latex, esp. of Sapotaceae; cf. Bassia, Mimusops, Pala- quium, Payena ; -puteh, Palaquium ; -rambong, Ficns ; -sundek, Payena. Guttiferae (EP. ; BH.. excl. Hypericaceae, incl. Quiinaceae). Dicoty- ledons (Archichl. Parietales). 42 gen., 850 sp. chiefly trop. They are (exc. Hypericum) trees or shrubs with simple entire opp. exstip. 1. Oil glands or passages are always present, often showing as translucent dots upon the 1. The infl. is cymose, frequently umbellate. The fls. show considerable variety. The bracteoles are frequently close up 20- 3o8 GUTTIFERAE Floral diagram of Hypericum; after Eichler, ovary cut at base. S the " Schraubel " branch. to the calyx, and hardly to be distinguished from it. The axis is usu. convex, but exhibits many forms. The fl. may be cyclic or partly spiral, and is usu. reg., 5 , hypog. K imbricate ; C imbr. or conv.; A oo , free or united in various ways, frequently in bundles (usu. regarded as due to branching of originally simple papillae) ; very com- monly the outer ones or even all are staminodial ; G usu. (5) or (3), multi- or i-loc.; ovules oo , few, or i, anatr. ; styles free or united. Fr. often capsular, sometimes a berry or drupe. Seed exalb. Many yield useful timber ; the resins of Clusia, Garcinia, Calophyllum, &c., and the fr. of Garcinia and others are valuable. Classification and chief genera (after Engler) : the G. are closely allied to Theaceae, the only constant distinction being the presence of oil-glands, and to Dipterocarpaceae, which are chiefly distinguished by their alt. slip. 1. [BH. excl. Hypericoideae with Endodesmioideae as a separate fam.] I. KIELMEYEROIDEAE (1. alt. or opp.; fls. usu. 5, A oo, style i, G 3 — 5-loc.): Kielmeyera, Caraipa. II. HYPERICOIDEAE (1. opp.; fl. 5 ; sta. usu. oo , usu. in 5 or 3 — 8 bundles before the petals ; styles 3 — 5, usu. free; fr. a i- or 3-loc. septi- or loculicidal caps., or indeh.; embryo usu. straight with not very thick cots.) : Hypericum, Vismia. III. ENDODESMIOIDEAE (sta. united into a tube above, in 5 bundles below ; cpl. i ; drape ; cots, fleshy): Endo- desmia (only genus). IV. CALOPHYLLOIDEAE (1. opp. ; fls. $ or ylus ; - withe, Hamamelis. Head, a mass of sessile fls. on a common recept., Compositae ; com- pound -, Echinops ; cymose -, Dipsacaceae, Haenianthus. Headache-weed (W.I.), Hedyosmum nutans Sw. Hearnia F. Muell. (Aglaia p.p. EP.). Meliaceae (n). 6 E. Indomal. Heart pea (W.I.), Cardiosptrmum ; -'s ease, Viola ; -wood, the older wood of a tree trunk, in which no sap runs. Heath, Cal/una, Erica; prickly, Pernettya; St. Dabeoc's -, Daboecia; sea -, Frankenia, Heather, Callttna vulgaris Salisb., Erica. Heaven, tree of, Ailanthus. Hebanthe Mart. (Pjaffia EP.). Amarantaceae (3). 20 trop. Am. Hebecladus Miers. Solanaceae (2). 7 W. trop. S. Am. Hebeclinium DC. =Eupatorium Tourn. p.p. (Comp.). Hebecoccus Radlk. Sapindaceae (i). i Java. Hebenstretia L. Scrophulariaceae (n. 7). 30 S. and trop. Afr. The corolla is slit open along the anterior side, and the style and sta. project through the slit. Hebepetalum Benth. Linaceae. 2 trop. S. Am. Heberdenia Banks (Myrsine p.p. RH. ). Myrsinaceae (n). 2 Macro- nesia, Mexico. Hebestigma Urb. Leguminosae (ill. 6). i W.I. Hebongu Radlk. Simarubaceae. 2 Phil. Is. 318 HECASTOCLEIS Hecastoclels A. Gray. Compositae (12). i Nevada. Hechtia Klotzsch. Bromeliaceae (2). 15 Mexico. Cult. orn. infl. Hecistopteris J. Sm. Polypodiaceae. i trop. Am. Heckeldora Pierre. Meliaceae (in). 2 trop. Afr. Heckelia K. Schum. Menispermaceae. i New Guinea. Heckeria Kunth (Piper p.p. EP.}. Piperaceae. 8 trop. Hectorella Hook. f. Portulaceae. i N.Z. Hecubaea DC. (Heleninm p.p. EP.). Compositae (6). i Mexico. Hedeoma Pers. Labiatae (vi). 30 Am. Hedeomoides Briq. (Pogogyne p.p.). Labiatae (vi). 3 California. Hedera Tourn. ex L. Araliaceae. 3 temp. ;%. H. Helix L. (ivy) is a root climber. L. dimorphic, those on the climbing shoots lobed, those on the freely projecting shoots that bear the infl. not. The former form leaf- mosaics better. Fls. not very conspicuous, but coming out late in the year are largely visited for the freely exposed honey by flies and wasps. Hederella Stapf (Dissochaeta p.p. EH.}. Melastomac. (i). 4 Mai. Arch. Hederopsis C. B. Clarke. Araliaceae (i). i Malay Peninsula. Hedge-Log grass (Am.), Cenchrus; -hyssop (Am.), Gratiola; -mus- tard, Sisymbriitm, (W.I.) Ckenopodium ; -nettle (Am.), Stackys; -parsley, Caucalis. Hedona Lous. (Lychnis p.p.). Caryophyllaceae (n. i). i Tibet. Hedraeanthus A. DC. Campanulaceae (i). 12 S.E. Eur., Caucasus. Hedraianthera F. Muell. Celastraceae. i E. Austr. Hedwigia Sw. (Tetragastris EP.). Burseraceae. 3 trop. Am., W.I. H. balsamifera Sw. (Antilles) is known as pig's balsam, on account of a legend that wounded pigs rub against the trees to heal wounds with the resin. Hedycapnos Planch. =Dicentra Bernh. (Papav.). Hedycarya Forst. Monimiaceae. 10 Austr. to Fiji. Hedychium Koen. Zingiberaceae (i). 30 trop. As., Madag. Rhizome olten tuberous. The fl. has a long tube, at the end of which spring the narrow free parts of the petals and the larger staminodes and labellum. The stigma projects just beyond the anther. Cult, orn. fl. Hedyosmum Sw. Chloranthaceae. 20 trop. Am. Hedyotis L. (Oldenlandia p.p. EP.}. Rubiaceae (i. 2). 120 trop. Hedypnois Schreb. = Rhagadiolus Tourn. Hedypnois (Tourn.) Scop. (Leontodon BH.}. Compositae (13). 3 Medit. Hedysarum (Tourn.) L. Leguminosae (in. 7), 70 N. temp. Hedyscepe H. Wendl. et Drude (Kentia p.p. EP.). Palmaceae (iv. i). i Lord Howe's Island. Heeria Meissn. (Anaphrenium E. Mey. BH.}. Anacardiaceae (3). 12 Afr. Heeria Schlecht. (Heterocentron EP.}. Melastomaceae (i). 6 C. Am. Some sta. attract insects, the others pollinate them (cf. Com- melina). Helmia Link et Otto (Nesaea BH.). Lythraceae. 2 trop. Am. Heinsenia K. Schum. Rubiaceae (n. i). 2 trop. Afr. HELIGME 319 Helnsia DC. Rubiaceae (r. 8). 5 trop. Afr. Heisteria Jacq. Olacaceae. 25 warm Am., W. Afr. Hekistocarpa Hook. f. Rubiaceae (i. 2). i Nigeria. Heladena A. Juss. Malpighiaceae (n). 5 Brazil, Argentina. Helcia Lindl. (Trichopilia BH.}. Orchidaceae (n. 19). i Panama. Heldreichia Boiss. Cruciferae (2). 6 W. As. Helenia L. = Helenium L. Helenium L. Compositae (6). 30 W. Am. Heleocharis (Eleocharis} R. Br. Cyperaceae (i). 90 cosmop. Heleochloa Host. Gramineae (8). 8 N. palaeotemp. Heleog-iton Schult. = Scirpus L. (Cyper.). Heleophylax Beauv. =Scirpus L. p.p. (Cyper.). Helia Mart. (Lisianthits BH. ). Gentianaceae (i). 7 trop. S. Am. Heliamphora Benth. Sarraceniaceae. i Guiana, a pitcher plant (cf. Sarracenia). Helianthella Torr. et Gray. Compositae (5). 15 W. U.S., Mexico. Helianthemum Tourn. ex Hall. Cistaceae. 125 Eur., Medit., N. Am. H. vulgare Gaertn. and 3 others in Brit, (rock-rose). Infl. a cin- cinnus. The fl. contains no honey and is homogamous, with sensitive sta., which move outwards when touched. Helianthostylis Baill. Moraceae (n). i Amazon valley. Heliantnum Engelm. ex Britton (Echinodorus EP.\. Alism. ? N. Am. Helianthus L. Compositae (5). 60 Am. Good for chars, of § 5. In H. annuus L. (sunflower) the number of fls. upon the head is often enormous and they show very regular spiral arrangement, probably due (largely) to pressure in the bud. Ray florets neuter. The seeds give oil. H. tuberosus L. (Jerusalem artichoke) has sub- terranean tuberous stems, like potatoes, with well marked ' eyes ' (buds in axils of scale-1 ). Helichrysum Vaill. ex L. Compositae (4). 350 Eur., As., Afr., Austr. ; 150 in S. Afr. Many xero. with hairy surface, decurrent, &c. The dried fl. -heads of some sp. are ' everlastings.' Helicia Lour. Proteaceae (n). 30 Indomal. &c. Helicilla Moq. Chenopodiaceae (B). i China. Helicodiceros Schott. Araceae (vn). i Corsica, Sardinia, H. cri- nitus Schott (H. muscivorus Engl. ). The development of the pedate leaf is cymose ; the later formed branches grow more slowly than the earlier. The name nniscivorus is due to the number of flies captured ; attracted by the foul smell of the infl. (cf. Arum) they collect inside the spathe in enormous numbers ; it may often be seen tightly packed ; when it withers the top closes and they are caught. Heliconia L. Musaceae. 35 trop. Am. Fls. in cincinni ; odd sep. post. Helicophyllum Schott (Eminium EP.}. Araceae (vn). 3 W. As. Helicostylis Tree. Moraceae (n). 2 Guiana, N. Brazil. Helicteres Pluk. ex L. Sterculiaceae. 45 trop. (exc. Afr.). The fls. become zygomorphic if they happen to be in a horiz. position. Helictonema Pierre. Celastraceae. i trop. Afr. Helietta Tul. Rutaceae (iv). 4 trop. Am. Heligme Blume = Parsonsia R. Br. (Apocyn.). 320 HE LINUS Helinus E. Mey. ex Endl. Rhamnaceae. 4 palaeotrop. Helio- (Gr- pref.), sun- ; -trope, Heliotropium ; -tropism, irritability to light ; winter -trope, Petasiles. Heliocarpus L. Tiliaceae. 5 Mexico to Paraguay. Heliocaxya Bunge. Boraginaceae (iv. i). i Persia. Heliocereus Britton et Rose (Cereus p.p.). Cactaceae (in. i). 4 N. Am Heliophila Burm. f. ex L. Cruciferae (i). 70 S. Afr. Heliophytum DC. = Heliotropium L. p.p. (Boragin.). Heliopsis Pers. Compositae (5). 7 Am. Heliosperma ReichL. (Silene p.p. BH.). Caryophyllaceae (u. i). 5 Mountains of S. Eur. Heliotropium (Tourn.) L. Boraginaceae (in). 220 trop. and temp. H. peruviamim L. (cherry pie) and others (heliotrope) cult perf. fls. Helipterum UC. Compositae (4). 50 Austr., S. Afr. Xero. with persistent invol. of white scaly bracts. Cult. orn. fl. ; the dried flower-heads are sold as 'everlastings' (cf. Helichrysum, &c.). Hellebore, Helleborus ; white -, Veratrutn. Helleborine Tourn. ex Hall, Pers. = Serapias L. (Orchid.). Helleborine Hill (Epipactls p.p.). Orchidaceae (il. 2). 10 N. temp. Helleborine, Epipactis. Helleborus (Tourn.) L. Ranunculaceae (2). 15 Eur., Medit. ; 2 in Brit, (hellebore). PI. woody below, each shoot from the stock taking several years to reach maturity and flower. Fl. protog., opening early in the year. Cpls. slightly coherent at base. In H. tiiger L. (Christmas rose) the P turns green after the fl. has been fert. Hellenia Willd. — Alpinia L. (Zingib. ). Helleranthus Small {Verbena p.p.). Verbenaceae (i). i N. Am. Hellwigia Warb (Alpinia. p.p.). Zingiberaceae (i). i New Guinea. Helmholtzia F. Muell. Philydraceae. 2 E. Austr., Polynesia. Helmia Kunth=Dioscorea L. p.p. (Dioscor.). Helminthia Juss. = Picris L. p.p. (Comp.). Helminthocarpum A. Rich. Leguminosae (in. 5). i Abyssinia. Helminthostachys Kaulf. Ophioglossaceae. i, H. zeylamca Hook, f., Ceylon, Himal. to Queensland. Rhiz. dorsiv. with 2-ranked 1. on the upper side, and roots below, which do not bear any definite relation to the 1. Sporangia peltate, on -vsporangiophores from the sides of the fertile spike. [Cf. Fanner and Freeman, in Ann. of Bot. xin. 1899, p. 421 ; Lang on prothallus, do. xvi. 1902, p. 23.] Helmontia Cogn. Cucurbitaceae (2). 2 Brazil, Guiana. Helobieae. The 2nd order of Monocotyledons. Helodea Reichb. = Elodea Michx. (Hydrocharit.). Helodrium (Cl.), a thicket formation. Helogyne Nutt. Compositae (2). 2 Peru, Bolivia. Helonias L. Liliaceae (i). i east N. Am. Cult. orn. fl. Heloniopsis A. Gray. Liliaceae (i). 4 Japan, Formosa. Helopus Trin. = Eriochloa H. B. et K. (Gram.). Helosciadium Koch = Apium Tourn. p.p. (Umbellif.). Helosis Rich. Balanophoraceae. 3 trop. Am. Helwingia Willd. Cornaceae (Araliac. BH.). 3 Himal. to Japan. Helxine L. = Polygonum Tourn. (.5//.) = Fagopyrum Moench. HEMIZONELLA 3 2 1 Helxine Reg. Urticaceae (4). i Corsica, Sardinia. Hemandradenia Stapf. Connaraceae. -2 trop. Afr. Hemarthria R. Br. \Rottboellia p.p. EP.). Gramin. (2). 3 warm. Hemerocallis L. Liliaceae (in). 5 temp. Eur., As. Infl. a double bostryx. The fls. of H. fulva L. are self-sterile. Cult. orn. fl. Hemi- (Gr. pref.), half, partial; -cyclic, part in spirals, part in whorls; -parasite, a facultative saprophyte, a parasite which can exist as a saprophyte ; -ptera, bugs, &c. ; -tropous (insects), with tongues of medium length, suited to fl. -classes B and B'. Hemiandra R. Br. Labiatae (u). 3 S.VV. Austr. Hemiarrhena Benth. Scrophulariaceae (ill. 3). i trop. Austr. Hemiboea C. B. Clarke. Gesneriaceae (i). 3 China. Hemicarex Benth. (Kobresia Willd., Sclioeno.\iphium Nees, BH.}. Cyperaceae (in). 10 Himalaya, S. Afr. Hemicarpha Nees et Am. (Scipus p.p. BH.}. Cyper. (i). 3 trop. and subtrop. Hemicarpus F. Muell.=;Trachymene Rudge (j5//.) = Didiscus DC. Hemichaeiia Benth. Scrophulariaceae (n. 4). i C. Am. Hemichlaena Schrad. Cyperaceae (i). 3 S. Afr. Hemicliroa R. Br. Chenopodiaceae (A). 3 Austr. Hemicramtoe Webb. Cruciferae (2). i Morocco. Hemicyclia Wight et Am. Euphorbiaceae (A. I. i). 10 Indomal. Hemidesmus R. Br. Asclepiadaceae (i). i S. India. Hemidia Rafin. Inc. sed. Nomen. Hemidiodia K. Schum. (Spermacoce p.p.). Rubiaceae (n. 10). t Mexico to Brazil, Malay Archip. Hemigenia R. Br. Labiatae (n). 25 Austr. HemiglocMdion K. Schum. (Phyllanlhus p.p.). Euphorbiaceae (A. I. i). 4 New Guinea. Hemigraphis Nees. Acanthaceae (iv. A). 25 trop. As. Hemigyrosa Blume (Guioa p.p. EP.}. Sapind'aceae (i). 4 Indomal. Hemiliabsnaria Finet (Habenaria p.p.). Orchid, (n. i). 3 E. As. Hemiheisteria Van Tiegh. (Heisteria p.p.). Olacaceae. i S. Am. HemilopMa Franch. Cruciferae (2). i Yunnan. Hemimeris L. f., Thunb. Scrophulariaceae (n. i). 4 S. Afr. Hemhneris Pers. = Alonsoa Ruiz et Pav. (Scroph.). Hemionitis L. Polypodiaceae. 10 trop. Am. and As. Hemiorchis Kurz. Zingiberaceae (i). 3 E. Indomal. Hemiphora F. Muell. Verbenaceae (3). i W. Austr. Hemipriragma Wall. Scrophulariaceae (in. i). i Himalaya. Hemiphylacus S. Wats. Liliaceae (ill), i N. Mexico. Hemipilia Lindl. Orchidaceae (n. i). 5 Himal., China. Hemipogon Uecne. in DC. Asclepiadaceae (n. i). 7 S. Am. Hemisiphonia Urb. Scrophulariaceae (n. 6). i W. Indies. Hemistemma Juss. ex Thou. = Hibbertia Andr. p.p. (Dillen.). Hemistepta Bunge (Saussurea p.p.). Compositae (n). i E. As. Hemistylus Benth. Urticaceae (4). 4 S. Am. Hemitelia Br. Cyatheaceae. 75 trop. and S. temp. Tree-ferns. Hemithrinax Hook. f. ( l^hrinax p.p. EP.). Palmaceae (1.2). i Cuba. Hemitria Rafin. Loranthaceae (inc. sed.). i, habitat?. Hemizonella A. Gray. Compositae (5). 2 Pacif. U.S. W. 21 322 HEMIZONIA Hemizonia DC. Compositae (5). 25 west N. Am. Hemizygia Briq. (Ocimum p.p. BH.}. Labiatae (vn). 2 trop. and S. Afr. Hemlock, Conium maculatum L.; -spruce (Am.), Tsuga canadensis Carr. ; water-, Cicuta virosa L. Hemp, Cannabis, Moraceae; -agrimony, Enpatorium cannabinum L. ; bastard -, agrimony (W.I.), Ageratum conyzoides L.; Bombay -, Crotalaria juncea L.; bow-string -, Sansevieria zeylanica Willd., &c. ; China -, Abutilon ; Deccan -, Hibiscus cannabinus L. ; Madras -, cf. Bombay ; Manila -, Musa textilis Nee ; Mauritius -, Furcraea gigantea Vent.; -nettle, Galeopsis Tetrahit L. ; New Zealand -, Phormium tcnax Forst. ; sisal -, Agave sisalana Perrine ; sunn -, Crotalaria juncea L. Hemsleya Cogn. Cucurbitaceae (i). 2 S.E. As. Hen-and-chickens, Calendula officinalis L., var. Hen-bane, Hyoscyamus niger L. ; -bit, Lainium ainplexicatile L. Henckelia Spreng. = Didymocarpus Wall. (Gesner.). Henequen, Agavefourcroydes Lem. Henicosanthum Becc. Anonaceae (i). i Borneo. Henlea Griseb. Malpighiaceae. i Cuba. Henleophytum Karst. {Henlea Griseb.). Malpighiaceae. i Cuba. Henna, Lawsonia inermis L. Hennecartia Poisson. Monimiaceae. i Paraguay. Henonia Moq. Amarantaceae (i). i Madag. Henoonia Griseb. Solanaceae (4) (Sapotaceae BH.). i Cuba. Henophyton Coss. et Dur. Cruciferae (4). i Algeria. Henosis Hook. f. (Bulbophylhtm p.p.). Orchid, (u. 16). i Brazil. Henricia Cass. Compositae (3). i Madag. Henriettea DC. Melastomaceae (i). 12 trop. S. Am. Henriettella Naud. Melastomaceae (i). 20 trop. Am., W.I. Henriquezia Spruce ex Benth. Rubiaceae (i. 4). 5 N. Braz., Guiana. Henrya Hemsl. Asclepiadaceae (n. i). i Centr. China. Henslowia Blume. Santalaceae. 13 Indomal. , China. Hensmania Fitzgerald (Xerotes p.p.). Liliaceae (in), i Austr. Hepatica Dill, ex L. = Anemone L. p.p. (Ranunc. ). Heppiella Regel. Gesneriaceae (n). 10 trop. S. Am. Heptacodium Render. Caprifoliaceae. i China. Heptacyclum Engl. Menispermaceae. i trop. Afr. Heptandrous, with 7 stamens. Heptanthus Griseb. Compositae (5). 3 Cuba. Heptapleurum Gaertn. (Schefflera EP.). Araliaceae (i). 70 palaeo- trop. Heracleum L. Umbelliferae (in. 6). 70 N. temp, and trop. Mts. (//. Sphondylium L., cow-parsnip, in Brit.). Herb, a pi. with no woody part above ground ; - Christopher, Actaea spicata L. ; - Bennett, Geum ; - Paris, Paris qitadrifolia L. ; - Robert, Geranium Robertianum L. ; -aceous (1.). thin and green (stem), not woody above ground. Herbarium, a collection of dried plants. Herbertia Sweet (Alophia EP.). Iridaceae (n). 7 warm Am. Hercules' Club (Am.), Aralia spinosa L. HESPE RASTER 323 Herderia Cass. Compositae (i). 3 trop. Afr. Herd's grass (Am.), Phleumpratense L. Heritiera (Dryand.) Ait. Sterculiaceae. 4 palaeotrop. coasts. Herkogamous, 5 , but incapable of self-fertilisation. Hermannia L. (BH. excl. Mahernia L.). Sterculiaceae. 120 trop. and sub-trop., chiefly Afr. Hermaphrodite (fl.), with both stamens and carpels (functional). Hermas L. Umbelliferae (i. 2). 5 S. Afr. Hermbstaedtia Keichb. Amarantaceae (i). 10 trop. and S. Afr. Hermesia Humb. et Bonpl. = Alchornea Sw. p.p. (Euph.). Hermesias Loefl. (Brownea BH.). Legumin. (n. 3). 10 trop. Am , W.I. Hermibiccnia x G. Camus, Bergon, et A. Camus. Orchidaceae. Hybrid, Herminiurn — Gymnadenia (Bicchia). Hermidium S. Wats. Nyctaginaceae. i Nevada. Henniniera Guill. £t Perr. (Aesckynomene p.p. EP.). Leguminosae (ill. 7). i trop. Afr.,.//. elaphroxylon G. et P. Wood light, used for floats, canoes, &c. Cf. with the development of aerenchyma seen in other marsh plants (Lycopus, Jussieua, &c.). Herminium L. Orchidaceae (n. i). 8 temp. Eur., As. (H. Monor- chis R. Br., musk-orchis, in Brit.) Hermione Salisb. = Narcissus Tourn. i>.p. (Amaryll.). Hermodactylus Tourn. ex Mill. Iridaceae (n). i Medit. Hernandia Plum, ex L. Hernandiaceae. 10 trop. Hernandiaceae (EP. ; Lanraccae p.p. BH.). Dicotyledons (Archichl. Ranales). 4 gen. 25 sp. trop. Shrubs or trees with alt. exstip. 1., oil-cells, and cystoliths. Fl. 5 or unisexual reg. ?4 — 10, A in whorl before outer P, G i-loc., with i pend. anatr. ov. Chief genus : Heinandia. Herniaria (Tourn.) L. Caryophyllaceae (i. 4). 20 Medit., Eur., S. Afr. (H. glabra L., rupture-wort, in England.) Fl. apetalous. Herpestis Gaertn. f. (Bacopa EP.). Scrophulariaceae (11. 6) 50 trop. and subtrop., chiefly Am. Herpetacanthus Nees. Acanthaceae (iv. B). 5 Brazil. Herpetica Cook et Collins, Rafm. (Cassia p.p.). Legum. (n. 5). i Porto Rico. Herpetospermum Wall. Cucurbitaceae (3). 2 Himal. , China. Herpolirion Hook. f. Liliaceae (in), i N.Z., Tasm., S.E. Austr. Herpysma Lindl. Orchidaceae (n. 2). 2 Himal., Phil. Is. Herpyza Sauv. (Teramnus EP.). Leguminosae (ill. 10). i Cuba. Herrania Goudot (Theobroma p.p. EP.). Sterculiaceae. 5 trop. S. Am. Herreria Ruiz et Pav. Liliaceae (n). 3 S. Am. Herrickia Wooton et Standley. Compositae (3). i New Mexico. Herschelia Lindl. (Disa p.p. BH.). Orchid, (n. i). 3 S. and trop. Afr. Hertia Neck. (Othonnopsis p.p. B.H.). Compos. (8). 8 W. As., S. Afr. Herya Cordemoy. Celastraceae. i Bourbon. Hesperaloe Engelm. Liliaceae (vi). 2 Texas, Mexico. Hesperantha Ker-Gawl. Iridaceae (in). 25 S. and trop. Afr. Hesperaster Cockerell (Mcntzelia p.p.). Loasaceae. 10 N. Am. 21 2 324 HESPERASTRA GAL US Hesperastragalus A. A. Heller (Astragalus p.p.). Leguminosae (in. 6). 4 Calif. Hesperelaea A. Gray. Oleaceae. i Lower California. Hesperidanthus Rydherg (Streptanthus p.p.). Cruc. (i). i N. Am. Hesperidium, the berry of Citrus, &c. Hesperis L. Cruciferae (4). 25 Eur., Medit.; (i Brit.). Orn. fl. Hesperocallis A. Gray. Liliaceae (in), i Colorado desert. Hesperochiron S. Wats. Hydrophyllaceae. 2 W. U.S. Hesperochloa Rydberg (Festuca p.p.). Gramin. (10). i Rockies. Hesperocnide Torr. Urticaceae (i). 3 California, Hawaiian Is. Hesperodoria Greene (Bigelomia p.p.). Compos. (3). 2 W. U.S. Hesperogenia Coulter et Rose. Umbelliferae (in. 4). i N. Am. Hesperolinon Small (Linuni p.p.). Linaceae. 10 California. Hesperomannia A. Gray. Compositae (12). 3 Sandwich Is. Hesperomecon Greene (Mcconella Benth.). Papaver. (n). 7 Calif. Hesperomeles Lindl. = Osteomeles Lindl. (Ros.). Hesperonia Standley (Mirabilis p.p.). Nyctaginaceae. 9 N. Am. Hesperoschordum Lindl. =Brodiaea Sm. p.p. (Lili.). Hesperoxalis Small (Oxalis p.p.). Oxalidaceae. i N.W. U.S. Heaperoyucca Baker ( Yucca p.p. ). Liliaceae (vi). i Calif. Hessea Herb. Amaryllidaceae (i). 10 S. Afr. Cult. orn. fl. Hetaeria Blume. Orchidaceae (n. 2). 20 palaeotrop. Heter, hetero- (Gr. pref.), diverse; -carpous, producing more than one kind of fr., Aethionema, Calendula, Cardamine, Dimorphotheca', -chlamydeous, with P 1. of two kinds, K and C ; -cyclic, with different numbers in different whorls ; -dromous (aestivation), a right-hand-side fl. I he reflection of a left, Exacum, Marantaceae, Saint paulia ; -gamous, heterostyled, or in Compositae, fls. of dif- ferent sex in same head ; -merous, whorls with different numbers of members; -phylly, polymorphic 1., Bryophyllum, Capsella, Dis- chidia, Liriodendron, Hedera and other climbers, many epiphytes, insectivorous plants, and water-plants; -spory, presence of two kinds of spores, Pteridophyta ; -stylism, occurrence of two or more kinds of pi. of the same sp., one with e.g. long sta. and short style, the other with short sta. and long style, or long, mid and short sta. and style, Lythruin, Primula, Androsace, Boraginaceae, Bouvardia, Eichhornia, Erythroxylum, Fagopyrum, Hottonia, Lagerstroemia, Linum, Lythruin, Alenyanthes, Mitchella, Oldcnlandia, Oxalis, Pontederia, Primula, Psychotria, Pitlinonaria, Kudgea, Statice, Turneraceae. Heterachaena Fres. (Launaea p.p. EP.). Compositae (13). i Arabia, Abyssinia. Heterachne Benth. Gramineae (10). 2 N. Austr. Heteracia Fisch. et Mey. Compositae (13). i W. As. Heteradelphia Lindau. Acanthaceae (iv. A), i Sao Thome. Heteranthelium Hochst. (Agropyron p.p. BH.). Gramineae (12). i W. As. Heteranthera Ruiz et Pav. Pontederiaceae. 10 trop. and subtrop. Am., Afr. L. of two types— linear submerged and orbicular floating. Some have cleist. fls. Heterauthia Nees et Mart. Scrophulariaceae (inc. sed.). i Brazil. HEURNIA 325 Heteraiithoecia Stapf. Gramineae (5). t trop. Afr. Heterapithmos Turcz. Inc. sed. i Brazil. Heterixia Van Tiegh. (Viseum p.p.). Loranth. (n). i N.Z., Malaya. Heterocarpus Phil. (Cardamine p.p. BH.}. Crucif. (4). i Juan Fern. Heterocaryum A. DC. = Echinospermum Sw. (BH.} = Lappula Moench. Heterocentron Hook, et Arn. (Heeria BH.). Melastomaceae (i). 6 C. Am., Mex. Heterochaenia A. DC. Campanulaceae (i). i Mascarenes. Heterochaeta DC. = Aster Tourn. p.p. (Compos.). Heterocodon Nutt. Campanulaceae (i). i W. N.Am. Heterocoma DC. Compositae (i). r Brazil. Heterodendron (um) Desf. Sapindaceae (i). 4 Austr. Heteroderis Hoiss. (Crepis p.p. BH.}. Compositae (13). i W. As. Heterodraba Greene (Draba p.p. ). Cruciferae (4). 2 C*alif. Heterogaura Rothrock. Onagraceae (2). i California. Heterolaena Sch.-Bip. = Eupatorium Tourn. p.p. (Compos.). Heterolepis Cass. Compositae (4). 38. Afr. Heteromeles M. Roem. (Photinia Lincll.). Rosac. (n). 2 Calif. Heteromerae (JBH.). The 2nd series of Gamopetalae. Heteromma Benth. Compositae (3). i S. Afr. nits. Heteromorpha Cham, et Schlechtd. Umbelliferae (in. 5). 3 Afr. Heteronoma DC. = Arthrostema Ruiz et Pav. (Melastom.). Heteropanax Seem. Araliaceae (i). i Fiji. Heteropappus Less. Compositae (3). 5 China, Japan. Heteropetalum Benth. Anonaceae (2). i N. Brazil. Heterophragma DC. Bignoniaceae (2). 3 Indomal. Heterophyllaea Hook. f. Rubiaceae (i. 5). 2 Bolivia, Argentina. Heteropogon Pers. (Andropogon p.p. EP.}. Gramin. (2). 5 trop. Heteropsis Kunth. Araceae (i). 8 trop. S. Am. Heteropteris H. B. et K. Malpighiaceae (i). 100 trop. Am., i in trop. Afr. Fr. a samara (cf. Acer, Banisteria). Heteropyxis Ilarv. Inc. sed. (Myrt. ?, Lythr. ?). 2 S. Afr. Heterosciadium Lange. Umbelliferae (in. 2). i Spain. Heterosmilax Kunth. Liliaceae (xi). 5 E. As. Heterospathe Scheff. Palmaceae (iv. i). 4 Malay Archip. Heterospermum Cav. Compositae (5). 6 Arizona to Argentina. Heterostachys Ung. Sternb. Chenopodiaceae (A), i C. and S. Am. Heterostemma Wight et Arn. Asclepiadaceae (n. 3). r=; Indomal. Heterostemon Desf. Leguminosae (n. 3). 4 trop. Am. Heterothalamus Less. Compositae (3). 6 S. Am. Heterotheca Cass. Compositae (3). 5 W. U.S., Mexico. Heterothrix Rydberg (Streptanthus p.p.). Cruciferae (i). 2 N. Am. Heterotis Benth. = Dissotis Benth. (Melastom.). Heterotoma Zucc. Campanulaceae (m). 6 Mexico. CulK orn. fl. Heterotrichum Bieb. = Saussurea DC. (Compos.). Heterotrichum DC. Melastomaceae (i). 10 trop. Am., some ed. fr. Heterotropa Morr. et Dene. =Asarum L. p.p. (Aristoloch.). Heucbera L. Saxifragaceae (i). 30 N. Am., sometimes apet. Heuffelia Schur. = Avena L. p.p. (Gramin.). Heurnia Spreng. (ffiu-mia R. Br.) Asclep. (n. 3). 20 Afr. 326 HEURNIOPSIS Heurniopsis N.E. Kr. (Hnerniopsis). Asclep. (n. 3). i S. Afr. Hevea Auhl. Euphorbiaceae (A. II. 3). 20 trop. Am. //. brasiliensis Miill.-Arg. is the source of the best caoutchouc (Para rubber), largely exported from Brazil. The tree was introduced into Ceylon and the east in 1876, and in recent years a very large planting industry has grown up in it. Incisions, usu. herring-bone-like, are made in the bark, and the latex flows from them. The wound is renewed at intervals of one or two days by shaving off a thin slice from the lower side, when there is a larger flow of milk than at first. The milk is usu. coagulated with the aid of enough acid to neutralise its alkalinity, and the resulting 'biscuits' or sheets are dried, or made into crepe or other forms. Cf. Willis, Agriculture in the Tropics. Hewardia Hook. Liliaceae (inc. sed.). i Tasmania. Hewittia Wight et Arn. Convolvulaceae (i). 5 trop. Hex- (Gr. preT.), six ; -androus, with 6 sta. , &c. Hexadesmia Brongn. Orchid, (n. 6). 5 Brazil to Mexico and W.I. Hexaglochin Nieuwland (Triglochin p.p.). Juncag. i U.S. Hexaglottis Vent. Iridaceae (n). 3 Cape Colony. Hexalectris Rafin. Orchidaceae (n. 7). 2 Mex., S. U.S. Hexalobus A. DC. Anonaceae (3). 8 trop. Afr., Madag. Hexaptera Hook. Cruciferae (i). 6 temp. S. Am. Fr. winged. Hexapterella Urb. Burmanniaceae. i Lower Amazon. Hexasepalum Bartl. ex DC. (Diodia p.p. EP.). Rubi. (n. 10). i Mex. Hexatheca C. B. Clarke. Gesneriaceae (i). i Borneo. Hexisea Lindl. Orchidaceae (n. 6). 5 Brazil to Mex. and W.I. Heylandia DC. Leguminosae (in. 3). r S. India, Ceylon. Heynea Roxb. (IVdlmra p.p. EP.).' Meliaceae (in). 4 Indomal. Heywoodia Sim. Euphorbiaceae (A. I. i). i Cape Col. (Cape ebony). Hians (Lat.), gaping. Hibbertia Andr. (incl. Candollea Labill. 1806). Dilleniaceae. 100 Austr. , New Caled. &c. Mostly ericoid or climbing shrubs. Some have phylloclades. Infl. dich., but often, by reduction, coming to look like a raceme. The sta. &c. vary much in number in different sp. Hibernation, remaining ^quiescent during winter ; cf. Bulbs, Corms, Rhizomes, Tubers, Water-plants, Orchidaceae, &c. Hibiscadelphus Rock. Malvaceae (4). 3 Hawaiian Is. Hibiscus L. (excl. Abelmoschm Medic.). Malvaceae (4). 160 trop. and subtrop. The 5 ante-sepalous sta. are repres. by teeth at the top of the stamen-tube. Several are cult., esp. H. Rosa-sinensis L. (shoe-flower, fls. showy), H. Sabdariffa L. (Rozelle, fr. for jelly, &c.), H. (A.) esculentus L. (Okra or Bandakai, mucilaginous young fr. in soups, £c.). Hickory, Carya. Hicksbeachia F. Muell. Proteaceae (n). i Austr. Hicoria Rafin. = Carya Nutt. (Jugland.). Hidalgoa La Llave. Compositae (5). 2 W.I., C. Am. Hiemalis (Lat.), winter. Hieracium (Tourn.) L. Compositae (13). 450 *_, S. Afr., Andes; several in Brit, (hawk-weeds). Innumerable varieties have been raised by various botanists to specific rank (see London Cat. of Brit. Plants). Some are parthenogenetic. HIPPOMARATHRUM 327 Hiernia Sp. Moore. Scroph. (in. 2). Formerly Acanth. i Angola. Hierobotana Briq. ( Verbena p. p. ) . Verben. (i). i Colombia. Hierochloe S. G. Gmel. Gramineae (7). 13 temp, and cold, i Brit. Hieronima Allem. Euphorbiaceae (A. i. i). 10 trop. Am., W.I. Hieronymiella Pax. Amaryllidaceae (i). i Argentina. Higginsia Pers. = Hoffmannia S\v. (Rubi.). Higinbothamia Uline. Dioscoreaceae. i C. Am. Hilairella Van Tiegh. Ochnaceae. 2 Brazil. Hilaria H. B. et K. Gramineae (3). 5 C. Am. to S.W. U.S. Hilbertia Thouin. Inc. sed. Nomen. Hildebrandtia Vatke. Convolvulaceae (i). i Afr., As. K enlarged on fr. Hillebrandia Oliv. Begoniaceae. i Hawaiian Is. Hilleria Veil. (Mohlana BH.). Phytolaccaceae. 3 trop. S. Am. Hillia Jacq. Rubiaceae (I. 5). 5 Brazil to W.I. Hilum, the scar where stalk separates from the seed. Himalayan spruce, Picea Morinda Link. Himantochilus T. Anders. Acanthaceae (iv. B). 6 Afr. Himantoglossum Spreng. (Orchis p.p. BH.}. Orchid, (n. i). i Medit., mid-Eur. Himantophyllum Spreng. =Clivia Lindl. p.p. (Amaryll.). Himantostemma A. Gray. Asclepiadaceae (n. 4). i N. Am. Himeranthus Endl. (Jaborosa p.p. EP.}. Solanac. (2). 5 Argentina. Hindsia Benth. Rubiaceae (i. 5). 7 trop. S. Am. Hing (India), asafoetida, Ferula. Hinterhubera Sch. Bip. 1855. Compositae (3). 3 Andes. Hip, the fr. of Rosa. Hippeastrum Herb. Amaryllidaceae (i). 75 trop. and subtrop. Am. Cult. orn. fl. Hippeophyllum Schlechter (Oberonia p.p.). Orchid, (n. 4). 2 N.G. Hippia L. Compositae (7). 4 S. Afr. Hippia L. f. = Plagiocheilus Arn. (Comp.). Hippia F. W. Schmidt = Gentiana Tourn. p.p. (Gent.). Hippobromus Eckl. et Zeyh. Sapindaceae (n). i S. Afr. Hippocastanaceae (EP.; Sapindaceae p.p. BH.}. Dicots. (Archichl. Sapindales). Only genus Aesculus (q.v.). Hippocastanum Tourn. ex Rupp. = Aesculus L. p.p. (Hippocast.). Hippocratea L. Hippocrateaceae. 80 trop. Twining shrubs. Hippocrateaceae (EP.; Celastraceae p.p. BH.). Dicots. (Archichl. Sapindales). 3 gen., 150 sp. trop. and subtrop. Shrubs, mostly lianes, with opp. or alt. simple 1. Fls. in cymes, $ , reg., with disc. K 5, C 5, A 3 (rarely 5, 4, 2), G (3), with 2—10 anatr. ov. in each loc. Berry or schizocarp. No enclosp. Genera: Campylostemon, Hippocratea, Salacia. Hippocrepis L. Leguminosae (ill. 7). 12 Medit., Eur. H. coinosa L. to Scotland. Fl. mechanism like Lotus. Useful fodders. Hippodamia Decne. (Solenophora BH.}. Gesner. (n). 3 Mex., Costa Rica. Hippomane L. Euphorbiaceae (A. n. 7). i C. Am., W. Ind., Columbia (manchineel). Latex poisonous. Hippomarathrum Hoffmg. et Link. Umbelliferae (in. 4). 12 Medit. 328 HIPPOPHAE Hippophae L. Elaeagnaceae. i N. temp. |^r, H. rhainnoides L. (sea buckthorn) Brit. In the 3 fl. the hracteoles form a hood over the sta. in wet weather; when the air is drier, they separate at the sides, and the pollen may be blown away. Hipposelimim Britton et Rose (Ligusticuni p.p.). Umb. (in. 5). i S. Eur. Hippotis Ruiz et Pav. Rubiaceae (i. 7). 5 trop. S. Am. Hippoxylon Rafin. = Oroxylum Vent. (Bignon.). Hippuridaceae (EP. ; Haloragidaceae p.p. BH.}. Dicots. (Archichl. Myrtiflorae). Only genus Hippuris, q.-v. Hippuris L. Hippuridaceae. i, H. vulgaris L. (mare's-tail) almost cosmop. A water plant, with creeping rhiz. and erect shoots, whose upper parts usu. project above the water. L. linear, in whorls, the submerged ones longer and more flaccid than the aerial. Fl. sessile in axil of 1., § (or sometimes ? on some stocks; cf. Labiatae), con- sisting of i epig. sta. and i cpl., with i pend. ov. and no integuments, and a slight seam representing the K; wind fertilised. Hiptage Gaertn. Malpighiaceae (i). 10 Mauritius to China. Hiraea Jacq. (Mitscagma'Bert.). Malpighiaceae (i). 30 trop. Am. Hircinus (Lat.), with goaty smell. Hirpicium Cass. Compositae (10). 2 S. Afr. HirscMa Baker. Compositae (4). i S. Arabia. Hirsute, with long distinct hairs. Hirtella L. Rosaceae (vi). 40 S. and Cent. Am., i Madag. Fl. -|-, axis deeply hollowed on one side. The sta. and cpl. are not in the hollow, but on the other side of the surface of the axis. Hirtus (Lat.), hirsute. Hising-era Hellm. = Xylosma Forst. (Flac.). Hispid, with rough bristly hairs. Hispidella Barnad. ex Lam. Compositae (13). i Iberian Penins. Histiopteris (Agardh) J. Sm. Polypodiaceae. 2 warm, and ^. Hitchenia Wall. Zingiberaceae (i). 4 India. Hitoa Nadeaud. Rubiaceae (n. 4). i Society Is. Hladnikia Koch {Plenrospermum p.p. BH,}. Umbell. (in. 4). i Adriatic. Hoarea Sweet= Pelargonium L'Herit. p.p. (Geran.). Hoary, grey with fine pubescence. Hobble-bush (Am.), Viburnum. Hochstetteria DC. (Dicoma p.p. EP.}. Compositae (12). i trop. Afr., Arabia. Hockinia Gardn. Gentianaceae (i). i Rio de Janeiro. Hodgkinsonia F. Muell. Rubiaceae (n. 3). i S.E. Austr. Hodgsonia Hook. f. et Thorns. Cucurbitaceae (3). j Indomal. Hodgsoniola F. Muell. Liliaceae (in), i S.W. Austr. Hoeckia Engl. et Graebn. Valerianaceae. i China. Hoehnelia Schweinf. Compositae (i). i E. Afr. Hoelzelia Neck. = Swartzia Schreb. (Legum. ). Hoepfneria Vatke (Abrus p.p. EP.}. Leguminosae (in. 9). i trop. Afr. Hoffimannia Sw. Rubiaceae (i. 8). 24 trop. Am. Hoffmanniella Schlechter. Compositae. Nomen. i trop. Afr. Hoffmannseggia Cav. Leguminosae (n. 7). 20 S. Afr., S. Am. HOLOTHRIX 329 Hofmeisterella Reichb. f. in Walp. Orchid, (n. 19). i Ecuador. Hofrueisteria Walp. Compositae (2). 5 Calif, to Mexico. Hog gum (W.I.), Moronobea; false- (W.I.), Rhits Metopium L.; -plum, Spondias; -weed, (W.I.), Boerhaavia\ poisoned -meat or weed (W.I.), Aristolochia grandiflora Sw., (Am.) Ambrosia artcmi- siaefolia L. Hohenackeria Fisch. et Mey. Umbelliferae (in. 5). 2 Medit., W. As. Hohenbergia Schult. f. p.p. (Aechmea p.p. BH.}. Bromeliaceae (4). 1 8 trop. Am. Hoheria A. Cunn. Malvaceae (2). 3 New Zealand. Hoitzia Juss. = Loeselia L. (Polemon.). Holacantha A. Gray. Simarubaceae. i New Mexico. Holalafia Stapf. Apocynaceae (n. i). i Guinea. Holarrhena R. Br. Apocynaceae (i. 3). 10 palaeotrop. Holboellia Wall. Lardizabalaceae. 5 Himal., China. Holcophacos Rydberg (Astragalus p.p.). Leguminosae (ill. 6). 2 N. Am. Holcus L. Gramineae (9). 8 Eur., N. and S. Afr. 2 in Brit., //. mollis L., and H. lanatits L., Yorkshire fog or soft-grass. Holigarna Buch. -Ham. ex Roxb. Anacardiaceae (4). 5 Indomal. Hollandaea F. Muell. Proteaceae (n). 2 E. Austr. Hollisteria S. Wats. Polygonaceae (i. i). i Calif. Hollrungia K. Schum. Passifloraceae. i New Guinea. Holly, Ilex ; -fern, Aspidium Lonchitis Sw. ; -hock, Althaea; -oak Quercus Ilex L.; -rose (W.I. ), Turnera; sea-, Erynginm. Holmbergia Hicken (Chenopodium p.p.). Chenopodiaceae (A). i Argent. Holmia Borner — Cobresia Pers. p.p. (Cyper.). Holmskioldia Retz. Verbenaceae (4). 4 Madag. , trop. Afr., Himal. Holo- (Gr. pref.), complete. Holocalyx M. Micheli. Leguminosae (n. 9). 2 Brazil, Haraguay. Holocarpa Baker (Pentanisia EP.). Rubiaceae (n. i). i Madag. Holocarpha Greene (Heinizoitia p.p.)- Compositae (5). i Calif. Holochlamys Engl. Araceae (n). 2 New Guinea. Holochloa Nutt.= Heuchera L. p.p. (Saxifrag.). Holodictyum Maxon (Aspknimn p.p.). Polypodiaceae. 2 Mexico. Holodiscus Maxim (Spiraea p.p. BH.}. Rosaceae (i. 3). 5 W. N.Am. Orn. HolograpMs Nees. Acanthaceae (iv. B). i Mexico. Hologyne Pnlzer (Coelogyne p.p.). Orchid, (n. 3). 2 Malay Archip. Hololachna Ehrenb. Tamaricaceae. 2 C. As. Holophyllum Less. = Athanasia L. p.p. (Comp.). Holopleura Regel et Schmalh. Umbelliferae (inc. sed.). i Turkestan. Holoptelea Planch. Ulmaceae. i Indomal. Holoschoenus Link. = Scirpus L. p.p. (Cyper.). Holostemma R. Br. Asclepiadaceae (n. i). 3 Indomal., China. Holosteum Dill, ex L. Caryophyllaceae (i. i). 6 N. temp. |# (i Brit.). Holostigma Spach = Oenothera L. p.p. (#//.). = Chamissoa Link. Holostylis Duch. Aristolochiaceae. i S. Centr. Brazil. Holothrix L. C. Rich. Orchidaceae (n. i). 25 Afr., trop. and S. 330 HOLOZONIA Holozonia Greene (Lagophylla i>.p. EP.}. Compositae (5). i N. Am. Holstia Pax. Euphorbiaceae (A. 11.2). 2 trop. Afr. Holubia Oliv. Pedaliaceae. i S. Afr. Holy grass (Am.), Hierochloe. Homalanthus A. Juss. Euphorbiaceae (A. n. 7). 10 Indomal., Polynes. Homalium Jacq." Flacourtiaceae (9). 80 trop. After fert. the sepals or petals, or both, grow large and form wings (often hairy) to the fr. Homalobus Nutt. ex Torr. et Gray (Astragalus p.p.). Leguminosae (in. 6). 10 N. Am. Homalocalyx F. Muell. Myrtaceae (n. i). i N.E. Austr. Homalomena Schott. Araceae (v). 80 trop. As. and S. Am. Homalopetalum Rolfe. Orchidaceae (n. 6). i Jamaica. Homalosciadium Domin (Hydrocotyle p.p.). Umbellif. (i. i). i Austr. Homalostachys Boeck. (Scleria p.p. EP.). Cyperaceae (n). i China. Homeria Vent. Iridaceae (u). 8 S. Afr. Bulbils in axils of lower 1. Cult. orn. fl. Homilacanthus Sp. Moore. Acanthaceae (iv. B). i E. trop. Afr. Hominy, the meal of maize, Zea Mays L. Homochaete Benth. Compositae (4). i S. Afr. Homochroma DC. Compositae (3). i S. Afr. Homogyne Cass. Compositae (8). 3 Mts. of Eur. Homoianthus Bonpl. ex DC. = Perezia Lag. p.p. (Comp.). Homoio-, Homo- (Gr. pref.), alike, similar; -chlamydeous (P), with 1. of one kind only; -dromous (aestivation), all 1. turned the same way, not to r. in one, to 1. in another, fl.; -gamous (fl.) one in which sta. and stigma ripen together, and cf. Compositae; -geneous, uni- form ; -logous, equivalent by descent ; -morphous, uniform in shape ; -nym, the same specific name of the same pi., in another genus; -plastic, equivalent in structure and mode of origin, but of parallel, not common, descent ; -sporous, with spores of one kind only, Pteri- dophyta. Homolepis Chase (Panic it in p.p.). Gramineae (5). 3 trop. S. Am. Homonoia Lour. Euphorbiaceae (A. II. 2). 4 Indomal. Homonoma Bello (Nepsera EP.). Melastomaceae (i). i Porto Rico. Homopogon Stapf. Gramineae (2). i French Soudan. Homoranthus A. Cunn. ex Schau. Myrtaceae (n. 2). i E. Austr. Homozeugos Stapf. Gramineae (2). i W. Afr. Honckenya Bartl. - Arenaria Rupp. (#//.) =Alsine Scop. p.p. Honckenya Willd. Tiliaceae. 3 trop. W. Afr. Honesty, Lunaria biennis Moench. Honey, a sweet secretion formed (usu. in fl.) by nectaries; cf. especially Bee-flowers ; -dew, Acer, Tilia ; -guides, marks, &c. to show the way to the honey, Myosotis ; -leaves, Berberidaceae, Lardizabalaceae, Rannnculaceae ; -locust, Gleditschia ; -palm,/Hl>afa ; sham-, Lopezia, Parnassia; -suckle, Lonicera, (W.I.) Desmodiitm, Tecoma, (Austr.) Banksia ; - - French, Hedysanim. Hoodia Sweet. Asclepiadaceae (n. 3). 5 trop. and S. Afr. Cactus-like. Hook-Climbers, cf. Climbing Plants; -S on fruit, &c., aiding animal- dispersal, cf. Agrimonia (on receptacle), Bidens (pappus), Cenchrus (sterile spikelets), Emex (P), Geum (style), Tragoceros (C), Triglo- chin (cpl.), Uncinia (axis of origin), Xantkium (invol.). HORSFIELDIA 3 3 1 Hookera Salisb. = Brodiaea Sm. (Lili.). Hoop pine, Araitcaria; -tree (W.I. ), Melia; -withe (W.I.), Cohibrina, Rivma. Hoorebekia Cornelissen (Aplopappus p.p.)- Compos. (3). 7 W. Am. Hop, Hi i in it I us Lnpuliis L. Hopea L. = Symplocos L. (Symploc. ). Hopea Roxb. Dipterocarpaceae. 50 Indomal. Hopkinsia Fitzgerald. Restiaceae. i Austr. Hoplestigma Pierre. Flacourtiaceae (2). 2 trop. Afr. Hoplestigmataceae, a fam. sometimes made to contain the last. Hoplophyllum DC. Compositae (i). 2 S. Afr. Hoplophytum Beer (Aechmea p.p. BH.}. Bromeliaceae (4). 2 Brazil. Hoppea Willd. Gentianaceae (i). 2 India. Hoppia Nees (Bisboeckelera O. Ktze.). Cyperaceae (u). 5 Brazil, Guiana. Horaninovia Fisch. et Mey. Chenopodiaceae (B). 3 W. As. Hordeum (Tourn.) L. Gramineae (12). 20 temp. 4 in Brit, (barley- grass). Spikelets in groups of 3 on the main axis, forming a dense spike. Each is i -flowered when perfect, but commonly either the central or the two lat. fls. are aborted. The cult, barley is H. vul- gare L. (ff. sativuin Pers.). The most common form is the var. distichum or 2-rowed barley, where the central fl. of each group is fertile, but 6-rowed barley (var. hexastichum), and 4-rowed barley or bere, are also grown. The last is the most hardy and is cult, as far as 70° N. (in Norway). Horehound, Marrubinm vulgare L. ; foetid-, Ballota nigra L.; white-, MarrnbiiiDi vulgare L. Horkelia Cham, et Schlecht. (Potentilla p.p. BH.}. Rosaceae (ill. 2). 35 W. U.S. Horkeliella Rydberg (Horkelia p.p.). Rosaceae (in. 2). 3 N. Am. Hormidium Lindl. ex Heynh. (Epidendrum p.p. EP.}. Orchidaceae (n. 6). 6C. Am., Cuba. Horminum Mill. = Salvia Tourn. p.p. (Labiat.). Horminum (Tourn.) L. Labiatae (vi). i Mts. of S. Eur. Hormogyne A. DC. (Sideroxylon p.p. EP.}. Sapotaceae (f). i Austr. Hornea Baker. Snpindaceae (i). i Mauritius. Hornemannia Vahl. Ericaceae (in. 2). 2 Guiana, W.I. Hornera Jungh. Inc. sed. 2 Japan. Horn-beam, Carfinus; -nut, Trapa; - of plenty, Fedia; -wort, Cera- lophyllutn. Hornschuchia Nees. Anonaceae (i). 2 Brazil. Hornschuchia Spreng. Inc. sed. r Brazil. Hornstedtia Retz (Ainoiiimn p.p. BH.}. Zingiberaceae (i). 40 Indomal. Hornungia Bernh. (Gagea p.p.). Liliaceae (iv). 6 Eur. Horse Cassia (W.I.), Cassia polyphylla Jacq. ; -chestnut, Aesculus Hippocastanum L. ; -gram, Dolichos hiflorus L. ; -hair, vegetable, Tillandsia; -mint (Am.), Monarda ; -purslane (W.I. ), TriantAema ; -radish, Cochlearia Armoracia L. ; -radish tree, Moringa pterygo- sperma Gaertn. ; -tail, Equisttuiii; -wood (W.I. ), Calliandra. Horsfleldia Bl. ex DC. (Harmsiopanax EP.}. Araliaceae (2). i Java. Horsfieldia Chitilot (Monophyllaea Reichb.). Gesner. (i). i Java. 332 HORSFIELDIA Horsfieldia Willd. (Myrhtica p.p. /?//.). Myristicaceae. 50 palaeo- trop. Horsfordia A. Gray (Sida p.p. ). Malvaceae (2). 4 Mexico, Calif. Hortensis (Lat.), of gardens. Hortia Vand. Rutaceae (iv). 4 Brazil. Horticulture, cf. Ornamental Plants. Hortonia Wight. Monimiaceae. 2 Ceylon. Hortus siccus, a herbarium, or collection of dried pi. Hosackia Dougl. Leguminosae (in. 5). 30 W. N.Am. Hosea Dennst. Inc. sed. i Inclomal. Hosea Ridley. Verbenaceae (4). i Penang. Hosiea Hemsl. et E. H. Wilson (Natsiatnm p.p.). Icacinaceae. r China. Hoslundia Vahl. Labiatae (vn). 3 warm Afr. Host (of parasite), the pi. on which it feeds. Hosta Jacq. = Cornutia L. (Verben.). Hosta Tratt. (Fnnkia BH.). Liliaceae (in). 5 Japan, China. Cf. Funkia. Hoteia C. Morr. et Dene. = Astilbe Buch.-Hani. (Saxifr.). Hotnima A. Chevalier. Euphorbiaceae (A. II. 4). i trap. Afr. Hottentot bread, Testudinaria\ -fig, Mesembrycmthemiuu. Hottonia Boerh. ex L. Primulaceae. 2, one N. Am., the other, H. paluslris L. (water-violet), Siberia and Eur. (inch Brit.). Floating water pi. with finely-divided submerged 1. The fls. project above the water; they are dimorphic like Primula. Houlletia Brongn. Orchidaceae (n. 13). 5 trop. S. Am. Cult. Hound's tongue, Cynoglossitm officinale L. Hounea Baill. Flacourtiaceae (6) (Passifl. BH.). i trop. Afr., Madag. Houseleek, Sempervwum. Houstonia Gronov. ex L. Rubiaceae (i. 2). 20 west N. Am. Fls. heterostyled as in Primula; similar differences in stigma and pollen. Houttea Decne. (Vanhonttea p.p. EP.). Gesneriaceae (n). 3 Brazil. Houttuynia Thunb. Saururaceae. i Himalaya to Japan. Partheno- genetic. Hovea R. Br. Leguminosae (in. 3). 12 Austr. Hovenia Thunb. Rhamnaceae. i Japan to Himal. Fr. axis ed. Hoverdenia Nees in DC. Acanthaceae (iv. B). i Mexico. Howardia Klotzsch = Aristolochia L. p.p. (Arist.). Howea Becc. Palmae (iv). 2- Lord Howe's Island. Cult. orn. Howellia A. Gray. Campanulaceae (in). 2 N. Am. Howittia F. Muell. Malvaceae (2). i Austr. Hoya R. Br. Asclepiadaceae (n. 3). 100 Indomal., Austr. Twiners and root-climbers with fleshy 1. Cult. orn. fl. (wax-flower). Hoyopsis Leveille. Celastraceae. i China. Hua Pierre et de Wild. Sterculiaceae. i trop. Afr. Huanaca Cav. (Azorella p.p. EP.). Umbelliferae (i. 2). 6 S. Am., Austr., Tasm. Huberia DC. Melastomaceae (i). 10 Brazil, Peru. Huckleberry, Gaylnssacia; blue-, Vacdniumpennsylvanicum Lam. Hudsonia L. Cistaceae. 3 N. Am. Huegelia Benth. = Gilia Ruiz et Pav. (Polemon.). HUTERA 333 Huegelia R. Br. Inc. sed. Nomen. Huernia (Heurnia) R. Br. Asclepiadaceae (n. 3). 20 S. and trop. Afr. Huernlopsis N.E. Br. Asclepiadaceae (n. 3). I Cape Colony. Huertea Ruiz et Pav. Staphyleaceae. 2 Peru, Cuba. Hufelandia Nees (Bdlschiniedia p.p. BH.). Lauraceae (n). i C. Am. Hugeria Small (I'accinium p.p.)- Ericaceae (in. i). i N. Am. Hugonia L. Linaceae. ntrop. |^. The lower twigs of the infl. are modified into hooks for climbing. Hulletia King. Moraceae (in). 2 Malay Peninsula. Hulsea Torr. et A. Gray. Compositae (6). 6 W. U.S. Humata Cav. Polypodiaceae. 20 palaeotrop. Humbertia Lam. Convolvulaceae (i). i Madag. Humble-bees, cf. Bee-flowers; robbery by -, Aconitutn, Delphinium. Humblotia Baill. Euphorbiaceae (A. I. i). i Comoro Is. Humboldtia Vahl (Batschia Vahl). Leguminosae (n. 3). 4 Ceylon and S. India. H. laurifolia Vahl is myrmecophilous. The non- flowering twigs are normal, but those that bear fls. have hollow obconical internodes. In each of these, at the top, opposite the 1., is a slit leading to the cavity which is inhabited by ants. Humea Sm. Compositae (4). 4 S. Austr. Humifusus (Lat.), spreading on surface. Humilis (Lat.), dwarf. Eumiria Jaume St. Hil. Humiriaceae. 3 trop. Am. Humiriaceae (EP., BH.). Dicots. (Archichl. Geraniales). 3 gen., 20 sp., trop. Am., Afr. Shrubs with alt. 1. and $ , reg- fls., 5-merous with 10 — oo sta. and cup-like disc. Ovules i — 2 per cpl. Drupe. CAief genera: Humiria, Saccoglottis. • Humming-bird flowers, A bud Ion, Erythrina, Marcgravia. Humulus L. Moraceae (iv). 2 N. temp. Perennial climbing herbs. Infl. cymose, dioec., the .\>. EP.). Dicots. (Monochlam. Curvembryae). A fam. unnaturally divorced from its true relation- ships. Herbs, rarely shrubby, with 1. usu. opp. entire, slip. Infl. usu. cymose, fl. usu. £ , inconspic. P herbaceous or leathery, persistent (4 — 5)> A. 4 — 5, rarely more or less, opp. P; G i-loc. style i, rarely 2 — 3. Ovule i, rarely 2, amphitr. or anatr. Endosp. Includes the last four groups of § I of Caryophyllaceae. Illecebrum Rupp. ex L. Caryophyllaceae (i. 4). i W. Eur. (incl. Devon and Cornwall), Medit., W. Afr. ' Illegitimate fertilisation, Ly thru in, Primula. Illicium L. Magnoliaceae. 10 Atl. N. Am., As. I. verum Hook. fil. (star-anise ; China) is used for flavouring. There is a gradual transition in the spiral P from sepaloid to petaloid structure (cf. Nymphaea). The fr. is an aggregate of follicles. Illigera Blume. Hernandiaceae (Combret. BH.). 10 palaeotrop. Illipe Koenig, F. Muell. (Bassia p.p. BH.). Sapotaceae (i). 35 In- domal. Ilyphilos Small (Elatine p.p.). Elatinaceae. i W. U.S. Wild rice. Ilysanthes Rafin. Scrophulariaceae (n. 6). 25 trop. and subtrop. Imantina Hook. f. (Morindd p.p. EP.). Rubiaceae (n. 9). i New Caled. INDIAN ALMOND 341 Imantophyllum Benlh. et Hook. f. (Imatophyllum Hook. ) = Clivia Lindl. p.p. (Amaryll.). Imbricaria Comm. ex fuss. (Mimusops p.p. EP.). Sapotaceae (2). 7 palaeotrop. Imbricate (aestivation, y.v.), overlapping. Imhofia Herb. = Hessea Herb. p.p. (Amaryll.). Immersed (venation), below surface. Immobilis (Lat), immoveable. Immortelles, everlastings, q. z<. Imparipinnate (1.), pinnate with odd leaflet at end. Impatiens Riv. ex L. Balsaminaceae. 340 trop. and N. temp., esp. Mts. of India and Ceylon. /. Noli-tangere L., the touch-me-not, in Brit. The name is derived from the explosive fr. , a caps, with fleshy pericarp ; the outer layers of cells are highly turgid and thus a great strain is put upon the whole. Dehiscence is septifragal and is started by a touch when the fr. is ripe. The valves roll up inwards with violence (starting at the base) and the seeds are scattered in all directions. Many cult. orn. fl. Imperata Cyrilli. Gramineae (2). 6 trop. and subtrop. /. arun- dinacea Cyrilli (lalang) is a very troublesome weed in Malaya. Imperatorla (Tourn.) L. = Peucedanum Tourn. p.p. (Umbell.). Imphee, Sorghum zmlgare Pers., var. Inaequale (Lat.), unequal. Inarticulate, not jointed. Incanus (Lat.), hoary-white. Incarvillea Juss. Bignoniaceae (2). 5 E. and Cent. As. L. alt. Incense, Boswellia, Dactyodes, Styrax. Incertae sedis, of uncertain position. Numerous gen. here given are so described, usu. because their original descriptions leave much to theimagination, but sometimes because of real difficulty in placing them. Incised (L), notched at the margin. Included, not projecting. Incompletae (BH.} = Monochlamydeae. Incomplete (fl.), wanting one or more kinds of organs. Incumbent, Cruciferae. Incurved, bending inwards. Indefinite (growth), continuing till checked by the cold; (infl.), with the first axis not ending in a fl. Indehiscent, not opening. Indeterminate (infl.), indefinite. Indian almond, Terniinalia Catappa L.; -bean (Am.), Catalpa; -butter tree, Bassia biityracea Roxb.; -chickweed (Am.), Mollugo; -copal, Vateria indica L.; -cork tree, Millingtonia hortensis L. f.; -corn, Zea Mays L. ; -cress, Tropaeolunr, -date, Tamarindus; -fig, Opimtta; -grass (Am.), Sorghum ; -hemp, Cannabis sativa L. ; -lilac, Melia, Lagers froemia; -liquorice, Abnts precatorius L. ; -madder, Rubia cordifolia L.; -mallow (Am.), Abutilon ; -meal, ZcaMays L. ; -millet, Panicnm ; -mulberry, Morinda citrifolia L.; -physic (Am.), Gillenia; -pink, Spigelia; -pipe (Am.), Monotropa; -redwood, Chickrassia ta&u/arisA..]uss.',-Tice, Zizaniaaqnatica L. ; -rubber, cf. Rubber; -shot, Canna ; -tobacco,( Am.), Lobelia inflata L. ; -turnip (Am.), Arisaema. 342 INDIGENOUS Indigenous, genuinely native. Indigo, Indigofera\ China green-, Rhamnus chlorophora Decne. ; Chinese -, Polygonum tinctorium Ait. Indigofera L. Leguminosae (in. 6). 300 trop. /. kptostachya DC., tinctoria L. and Anil L. furnisli indigo. The plant is mown just before flowering, and soaked in water, whereby a yellowish solution is obtained. This on stirring and exposure to the air oxidises, and an insoluble precipitate of indigo is formed. The fls. are slightly explosive (cf. Genista). Indokingia Hemsl. Araliaceae (i). i Seychelles. Indomalaya, cf. Floral Regions, it (6) and (7). Indovethia Boerlage. Ochnaceae (Violaceae BH.}. i Borneo. Induplicate (aestivation), leaf margins turned inwards; (vernation), Palmae. Indusium, Filicales, Pteridophyta. Inermis (Lat.), unarmed, thornless. Inferae (BH.}. The ist series of Gamopetalae. Inferior (ovary), enclosed in the receptacle. luflexed, bent inwards. Inflorescence, the reproductive shoot, composed of, or bearing, a number of shoots of limited growth, termed flowers. Of two types, mono- podial or racemose, where the first axis does not as a rule terminate in a fl., but grows steadily onwards; and sympodial or cymose, where the main axis soon terminates in a fl., and the growth is taken up by the lat. axes in succession. The typical raceme has an axis growing indefinitely up the middle, and successively younger fls. on lat. branches, Cruciferae, Primus, Ribes. There are also the cpd. raceme or panicle, the raceme with sessile fls. or spike, with its var. the catkin or pendulous spike, the corymb, or raceme with all the fl. stalks elongating to the same level, Iberis, the umbel, which may be imagined as a corymb with all the fls. springing from one point, Umbelliferae, the cpd. umbel, the head, which is a common re- ceptacle with the fls. arranged in a dense mass with the youngest to the centre, Compositae, the cpd. head. The cyme may be mono-, di- or pleiochasial, according as each branch bears upon itself r, 2 or more branches; cf. dichasial cymes, &c. Many pi. possess mixed infls., with some branchings racemose, some cymose, Aesciilus, Be- tulaceae, Labiatae, Verbascum. Infundibuliform, funnel-shaped. Inga Scop. Leguminosae (i. i). 150 trop. and subtrop. Am., W.I. Ingenhouzia (Moc. et Sesse ex) DC. Malvaceae (4). i Mexico. Ink berry (W.I.), Randia aculeata L.; - nut, Semecarpus Anacardium L. f., Terminalia. Innate (anther), joined to filament by its base. Inobulbum Schlechter et Kranzlin (Dendrobium p.p.). Orchidaceae (n. 15). 2 New Caledonia. Inocarpus F\>rst. Leguminosae (in. 8). i Malaya, Polynesia. Seed ed. Inodes O. F. Cook (.SVz/Wp.p.). Palmaceae (i. 2). 9 N. Am., W.I. Inrolled leaf, Capparidcifcae, Einpctrum. Insect powder, Chrysanthemum. IPE CA CUANHA 343 Insectivorous plants, pi. which capture insects, &c., by special appa- ratus, and absorb the resulting products, whether after a special fermentation, or as humus. About 400 spp. belonging to Droseraceae (Aldrovanda, Dionaea, Drosera, Drosophylluin, &c.), Cephalotaceae (Cephalotus), Lentibulariaceae (Pingiticiila, Utricularia, &c.), Ne- penthaceae (Nepenthes), and Sarraceniaceae (Sarracenia, &c.). Cf. gen. mentioned for details, and Darwin, Insectivorous Plants. Insertion of leaves, mode of union with stem ; cf. Leaf. Insignis (Lat.), notable. Insolation, exposure to sun. Integrifolius (Lat.), simple -leafed. Integuments, the coats of the ovule-. Inter- (Lat. pref.), between ; -calary (growth), at a point between apex and base; -cellular spaces, air spaces in leaves, &c. ; -node, the space between a leaf and the next above it ; -petiolar stipules, Rubiaceae. Interruptedly pinnate, with alt. large and small leaflets, Rosaceae. Intrapetiolar stipules, Rubiaceae. Introrse (anther), opening towards centre of fl. Intruded, projecting forwards. Intsia Thou. (Afzelia p.p.). Leguminosae (n. 3). 8 palaeotrop. Inula L. Compositae (4). 100 Eur., As., Afr. (4 in Brit). The root of/. Heleniiim L., the elecampane, is officinal. Inulin, a carbohydrate. Inulopsis O. Hoffm. (Aplopappus p.p.). Compositae (3). i S. Brazil. Inversodicraea Engl. ex R. E. Fries. Podostemaceae. i Victoria Falls. Inverted flower, Clitoria, Campanulaceae, Orchidaceae. Involucel, secondary involucre, Uinbelliferae. Involucre, a whorl of bracts, usu. in condensed infls., Anemone, Com- positae, Eranthis, Unibelliferae. Involute (vernation), margins rolled inwards. Inyonia M. E. Jones. Compositae (inc. sed.). i N. Am. lochroma Benth. Solanaceae (n. 2). 15 trop. Am. /. macrocalyx Miers shows protection of the fl.-buds, as in Spathodea,- by watery secretion between K and C. Cult. orn. fl. lodanthus Torr. et Gray. Cruciferae (2). i Atl. N. Am. lodes Blume. Icacinaceae. 6 trop. As., Afr., Madag. * lodina Hook, et Am. Santalaceae. i temp. S. Am. lonactis Greene (Aster p. p.). Compositae (3). 3 N. Am. lone Lindl. (Bulbophyllum p.p. BII.). Orchid, (n. 16). 5 Indomal. lonidium Vent. Violaceae. 50 trop. and subtrop. The roots of /. Ipecacuanha Vent, are used in medicine (white Ipecacuanha) in the same way as the true drug (Uragoga). lonopsidium Rchb. (Cochlearia p.p. BH.). \ Portugal, /. acaule Rchb. with sol. fls. in the axils of radical leaves. lonopsis H. B. et K. Orchidaceae (n. 19). 10 trop. Am., epiphyte's. lonoxalis Small (Oxalis p.p.). Oxalidaceae. 70 N. Am. lostephane Benth. Compositae (5). 2 Mexico. Ipecacuanha Arruda = Psychotria L. (Bff.) — Uragoga L. Ipecacuanha, Uragoga Ipecacuanha Baill. ; bastard- (W. I.), Ascle- pias, curassavica L. ; wnite -, lonidium Ipecacuanha Vent. 344 JPHIGENIA Iphigenia Kunth. Liliaceae (i). 8 S. Aft. and Madag. to N.Z. IpMona Cass. Compositae (4). 10 S. Afr. to Turkestan. Ipnum Phil. (Diplachne p.p. EP.}. Gramineae (10). i temp. S. Am. Ipomoea L. (BH. incl. Aniseia, Batatas, Calonyction and Exogonium of Choisy, Mitta Cerv., Operculina Silva Manso, Pharbitis Choisy, and Quanwdit Moench.). Convolvulaceae (i. 4). 310 trop. and warm temp., chiefly climbing herbs or shrubs; many cult. orn. fls. (e.g. I. p2irpurea Roth, the morning glory). /. biloba Forst. is a char, creeping pi. of trop. beaches. /. Batatas Lam. (B. ednlis Choisy) is the sweet potato, largely cultivated in warm countries for its tubers, which are used like potatoes. /. (Exogonium) Purga Hayne is the jalap ; its rhizome gives off turnip-like roots about the size of apples. Worm- eaten tubers are most valuable, as the non-resinous parts are eaten. Ipomopsis Michx. = Gilia Ruiz et Pav. p.p. (Polemon.). Ipsea Lindl. (Pachystoma BH.). Orchidaceae (n. 9). 3 trop. Afr. and As. Ir6 rubber, Funtumia elastica Stapf. Iresine P. Br. Amarantaceae (3). 25 Am., Afr. Iriartea Ruiz et Pav. Palmae (iv. i). 10 trop. S. Am. The stem is supported on aerial roots (cf. Pandanus). Some of the branches of these roots are thorny (cf. Acanthorhiza). In /. ventricosa Mart. (Paxiuba palm), the stem has a peculiar egg-like thickening about half-way up (cf. Bombacaceae, Jatropha). Iriartella H. Wendl. (Iriartea p.p. EP.}. Palmaceae (iv. i). i Amazon. Iridaceae (EP., BH.). Monocotyledons (Liliiflorae ; Epigynae BH.). 57 gen., 800 sp. trop. and temp.; the chief centres of distr. S. Afr. and trop. Am. Chiefly herbs with a sympodial tuber or rhizome below ground. L. usu. equitant in two ranks. Infl. term., cymose (i fl. only in Crocoideae). Fl. ?, reg. or -|-. P 3 + 3, petaloid, united below into a long or short tube; A 3 (the outer whorl), with extr. anthers; G (3), 3-loc., with axile plac. (rarely i-loc. with parietal plac.); style usu. trifid and frequently ± petaloid. Ovules usu. oo , anatr. "Loculic. caps. Embryo small, in hard endosp. Classification and chief genera (after Pax) : I. CROCOIDEAE (fl. solitary, or several developed centri- fugally round a central one; plant small; 1. not exactly in \ phyllotaxy) : Crocus, Romulea. II. IRIDOIDEAE (fls. numerous, in spathes, several in each, usu. reg.; stem distinct; 1. equitant): Iris, Moraea, Ti- gridia, Sisyrinchium. III. IXIOIDEAE (similar, but spathes i -flowered; fl. often zygomorphic) : Ixia, Tritonia, Gladiolus, Freesia. Irideae (BH.) = Iridaceae. Iris Tourn. ex L. Iridaceae (n). 125 N. temp. 2 in Brit., /. Pseu- dacorus L., the yellow flag, and I.foetidissima L , the gladdon. Many cult. orn. fl. Most have a sympodial rhiz. with equitant isobilat. 1., and small cymes of fls. in spathes. P petaloid, the sepals usu. bending downwards at the outer ends; opp. to them and almost resting on them are the petaloid styles, under which are the sta. with their extr. anthers. Just above the anther, on the outer side of the style, is a ISOBERLINIA 345 little flap, whose upper surface is the stigma. Bees entering the fl. to get the honey secreted by the ovary rub off their pollen upon the stigma ; going farther in they get fresh pollen ; and when they come out close the stigma flap, which prevents self-fert. (cf. Viola). The flat seeds are suited to wind-dislr. The dried rhiz. of I . florentina L. (Orris root) smells like violets, and is used in perfumery; 'essence of violets' is made from it. Irlbachia Mart. (Lisianthus Bff.}. Gentianaceae (i). 3 trop. S. Am. Irmischia Schlechtd. (Metastelma BH.}. Asclepiad. (n. i). 3 Mex., W.I. Iron bark, Eucalyptus; -shrub (W.I.), Sauvagesia crecta L. ; -weed (Am.), }'ernonia; - wood, Mesna, &c. , applied to different woods in different countries. Irregular (fl.), one in which any whorl has members not all alike. Irritability, sensitiveness to stimuli. Irvlngella VanTiegh. (Irvingia p.p.)- Simarub. 10 trop. Afr. and As. Irvlngia Hook. f. Simarubaceae. 8 trop. Afr. and As. Butters from the seeds (cay-cay, dika, &c.). Iryanthera Warb. (Myristica p.p. BH.}. Myristicaceae. 4 N. trop. S. Am. Isabella Barb. Rodr. Orchidaceae (n. 6). i Brazil. Isachne R. Br. Gramineae (5). 30 trop. and subtrop. Isandra F. Muell. Solanaceae (inc. sed.). i Austr. Isanthera Nees. Gesneriaceae (i). 3 Indomal. Isanthus L. C. Rich, in Michx. Labiatae (i). i N. Am. Isatis Tourn. ex L. Cruciferae (2). 50 Medit., Eur., As. /. tinctoria L. is the woad, largely used as a dye before the introduction of indigo. It is prepared by grinding the leaves to a paste and fermenting them. [Nature, 55, pp. 36.795 61, pp. 331, 563.] Ischaemum L. Gramineae (2). 40 trop. and subtrop. Ischarum Blume=Biarum Schott (Araceae). Ischnoa F. Muell. Compositae (7). i New Guinea. Ischnocentrum Schlechter. Orchidaceae (n. a. in). New Guinea. Ischnochloa Hook. f. Gramineae (2). i N.W. Himalaya. Ischnogyne Schlechter. Orchidaceae (n. 16). i Yunnan. Ischnolepis Jumelle et Perrier. Asclepiadaceae (i). i Madag. Ischnosiphon Koern. Marantaceae. 20 trop. Am. Ischnostemma King et Gamble. Asclepiadaceae (n. i). i Malaya. Ischnurus Balf. f. Gramineae (12). i Socotra. Iseilema Anderss. (Anthistiria p.p. BH. ). Gramineae (2). 5 Indomal. Isertia Schreb. Rubiaceae (i. 7). 10 S. Am. Isidorea A. Rich. Rubiaceae (i. i). i W.I. Island floras, cf. Wallace, Island Life, Willis in Ann. Bot. 1916-17. Ismeue Salisb. = Hymenocallis Salisb. (Amaryllid.). Isnardia L. = Ludwigia L. (Onagrac.). Iso- (Gr. pref.), equal, especially in number; -bilateral (L), with symmetrical internal structure, and edge to the light, Acorns, Aspho- dehis, Eucalyptus, Gladiolus, Nartheciuin ; -diametric, of equal diameters ; -merous, with equal numbers in every whorl ; -sporous, with all spores alike ; -stemonous, with sta. in one whorl, as many as P. Isoberlinia Craib et Stapf. Leguminosae (n. 3). 2 Nigeria. 346 1SOCARPHA Isocarpha R. Br. Compositae (5). 5 Texas to Peru, W.I. IsocMlus R. Br. Orchidaceae (n. 6). 5 trop. Am., W.I. Isochoriste Miq. Acanthaceae (iv. B). 2 Java, Angola. Isocoma Nutt. (Bigelowia p.p.). Compositae (3). 10 N. Am. Isodendrion A. Gray. Violaceae. 3 Hawaiian Is. Isodesmia Gardn. Leguminosae (in. 7). 2 Brazil. Isoetaceae. Isoetales. Only genus Isoetes (., ./5^.) Dicots. (Archichl. Juglandales; Unisex. BH.}. 6 gen., 40 sp., N. temp., trop. As. Trees; alt. stip. 1., with brown hairy winter buds ; the buds arise rather high up in the leaf axils, and sometimes several, appear in descending order. Infl. monoecious, the 3°° SP- m damp and cold places, temp, and frigid zones and trop. Mts. Usu. creeping sympodial rhiz., one joint of the sympodium appearing above ground each year as a leafy shoot. The stem does not often lengthen above ground, except to bear the infl.; 1. usu. narrow, occasionally centric (Juncus). Infl. usu. a crowded mass of fls. borne in cymes of various types, usu. monochasial. Fl. £ , regular wind-fertilised. P 3 + 3, sepaloid, with the odd leaf of the inner whorl post.; A 3 + 3 (or the inner wanting), anthers dehiscing lat., pollen in tetrads; G (3); plac. axile or parietal, with oo or few anatr. ov. ; style simple, with 3 brush-like stigmas. Loculic. caps. Embryo straight, in starchy endosp. Chief genera: Prionium, Juncus, Luzula. Juncaginaceae = Scheuchzeriaceae. Juncella F. Muell. (Trithuria BH.). Centrolepidaceae. 2 S. Austr., Tasm. Juncellus C. B. Clarke (Cyperus p.p. EP.). Cyperaceae (i). 10 |*. Junceus (Lat.), rush-like. Juncoides Dill. — Luzulu DC (June.). Juncus (Tourn.) L. Juncaceae. 225 cosmop. but chiefly in cold, wet places. 1 8 sp. of rush are found in Brit. Most have a sympodial KAGENECKIA 351 rhiz. giving off one leafy shoot each year. The I. are of various types, with large sheathing bases. Some are flat and grass-like, others needle-like, and still others centric in structure and standing erect. The infl. is a dense head or panicle, of cymose construction (usu. rhipidia or drepania). In some sp. it appears to be lat. on a leaf-like cylindrical stem, but is really only pushed to one side by the bract of the infl. Fl. protog. and wind-fert. Rushes are largely used for making baskets, chair bottoms, &c. J. squarrosus L. is common on hill pastures in Brit. ; it is eaten by sheep and forms a valuable part of their fodder when grass is scarce. June-berry (Am.), Amelanchier. Jungia L. f. Compositae (12). 15 S. Am. Jungle, low or thin forest. Juniper, Juniper us. Juniperus Tourn. ex L. Coniferae (Pinac. 42 ; see C. for genus cha- racters). 30 *. The juniper,/, communis L. (Asia and Eur. incl. Brit.), and J. Oxycedrus L., &c. have needle 1. throughout life; others, such as J. Sabina L. , the savin (Eur. As.), have small 1. closely appressed, as in Cupressus. Seedling forms of these are known (see Retinospora). The cone consists of i — 4 whorls of scales, one only being fertile, as a rule.. In ripening the whole becomes a fleshy mass enclosing the hard seeds, and forming a good imitation of a true berry. The fruit is eaten by birds. That of/", communis is used in making gin. The wood of/, virginiana L. is the red cedar used for pencils ; others also give useful timber. Juno Tratt. = Iris L. p.p. (Irid.). Junodia Pax. Euphorbiaceae (A. i. i). i trop. Afr. Jurinea Cass. Compositae (n). 50 Medit., Eur., As. Juruasia Lindau. Acanthaceae (iv. B). 2 Amazonas. Jussieua L. (Jussiaea). Onagraceae (l). 40 trop. ; water and marsh plants. Aerating tissue is well developed (cf. Sonneratia, Sesbania). In/, repens L. (/. diffusa Forsk.), when growing in water, two forms of root develope, ordinary anchorage roots and erect spongy roots which grow upwards, often till they reach the surface of the water. The bulk of the tissue consists of aerenchyma. In /. suffruticosa L. (/. salicifolia H. B. et K.) there is an erect stem, whose lower part is covered with aerenchyma if growing in water (cf. Lycopus). If the plants be grown on land none of these phenomena appear. [Figs., &c., in Goebel's Pflanzenbiol. Schild. II. 256.] Justenia Hiern. Rubiaceae (i. 2). i trop. Afr. Justicia Houst. ex L. Acanthaceae (iv. B). 300 trop. Jute, Corchorus; China -, Abittilon Avicennae Gaertn. Kadsura Kaempf. ex Juss. Magnoliaceae. 8 trop. As., China, Japan. Fls. unisexual, spiral throughout. Climbing shrubs with no sti- pules. Kadua Cham, et Schlecht. Rubiaceae (i. 2). 16 Hawaiian Is. Kaempferia L. Zingiberaceae (l). 55 trop. As. and Afr. Cult. orn. fl. Kaernbachia Schlechter. Cunoniaceae. i New Guinea. Kaffir boom, Erythrina ; - bread, Encephalarlos ; - corn, Sorghum vul- garePers.; -lily, Clivia; -thorn, Lycitt/ti. Kageneckia Ruiz et Pav. Rosaceae (i. 2). 3 Chili. 352 KAKOSMANTHUS Kakosmanthus Hassk. (Payena p.p. BH.). Sapotaceae (i). 4 Malay Arch. Kalaliaria Baill. Verbenaceae (4). 2 trop. and S. Afr. Kalanchoe Adans. Crassulaceae. 70 trop. Like Bryophyllum. Kale, Brassica oleracea L. var. ; sea-, Crambe maritima L. Kalidium Moq. in DC. Chenopodiaceae (A). 4 S. Russia, W. As. Kaliphora Hook. f. Cornaceae. 4 Madag. Kallstroemia Scop. (Trilndiis p.p.). Zygophyllaceae. 10 Am., Austr. Kalmia L. Ericaceae (i. 3). 6 N. Am., Cuba. The anthers are held in pockets of the C, and the filaments are bent like bows when the fl. is open. An insect probing for honey releases them, and the anthers strike against him, loading him with pollen. Kalmiella Small (Kalmia p.p.). Ericaceae (i. 3). i N. Am. Kalopanax Miq. (Acanthopanax BH.). Araliaceae (i). 2 E. As. Kalosanthes Haw. = Rochea DC. (Crass.). Kalymopetalon Pohl. Inc. sed. Nomen. Kampmannia Steud. Gramineae (inc. sed.). i N. Zealand. Kanahia R. Br. Asclepiadaceae (n. i). 3 E. Afr., Arabia. Kandelia Wight et Am. Rhizophoraceae. i trop. As. Kangaroo apple, Solatium aviculare Forst.; -grass, Anthistiria\ -thorn, Acacia armata R.Br. Kania Schlechter. Saxifragaceae (ill), i New Guinea. Kaniata, Mallotus. Kanimia Gardn. Compositae (2). 7 trop. S. Am. Kapa cloth, Broitssonetia papyri/era Vent. Kapok Eriodendron aiifractuosum DC., Bombax. Karatas (Plum.) Mill. Bromeliaceae (4). 3 W. Ind. Karite, Buty rasper mum Parkii Kotschy. Karlea Pierre (Maesopsis EP.). Rhamnaceae (formerly in Styraceae and Sapotaceae). i trop. Afr. Karpaton Rafin. Inc. sed. i N. Am. » Karri, Eucalyptus diversicolor F. Muell. Karwinskia Zucc. Rhamnaceae. 3 warmer N. Am. Katabolism. the breaking down of materials in the pi. Katafa Costantin et Poiss. (Cedrelopsis Baill.). Meliaceae (i). (Authors place as new tribe of Celastraceae.) i Madag. Kaufmannia Regel. Primulaceae. i Turkestan. Kaulfussia Blume (Christensenia Maxon). Marattiaceae. 2 S.E. As. The palmate 1. has large pores below, due to tearing apart of guard cells of stomata. [Campbell in Ann. Bzg. 2. vn. 69.] Kauri, - copal, Agathis australis Steud. Kayea Wall. Guttiferae (iv). 12 Indomal. Kedrostis Medic. Cucurbitaceae (2). 15 trop. Afr. and As. Keel, cf. Leguminosae. Keenania Hook. f. Rubiaceae (i. 7). 2 Indomal. Keerlia A. Gray et Engelm. Compositae (3). 3 Texas, Mexico. Kefersteinia Reichb. f. (Zygopetalum BH.}. Orchid, (n. 14). 6 trop. S. Am. Kegelia Reichb. f. Orchidaceae (li. 13). i Guiana. Kei-apple, Aberia (Doryalis) caffra Harv. et Sond. Keiria Bowdich. Oleaceae. i Madeira. KIGELIA 353 Keiskea Miq. Labiatae (vi). i Japan. Keithia Benth. (Hedeoma p.p. EP.}. Labiatae (vi). 9 trop. Am. Keithia Spreng. Capparidaceae (inc. seel.), i Brazil. Keitia Regel. Iridaceae (n). t Natal. Kelleronia Schinz. Zygophyllaceae. 2 E. trop. Afr. Kelloggia Torr. ex Benth. et Hook. f. Rubiaceae (n. 7). W. N. Am. Kelseya Rydberg (Eriogyiiia p.p.). Rosaceae (i. i). i Montana. Kendal green, Genista tinctoria L. Kendrickia Hook. f. Melastomaceae (i). i Ceylon, S. India. Kenguel seed, Silylnim Mariamtm Gaertn. Kennedya Vent. (incl. Hardenbergia Benth.). Leguminosae (ill. 10). 15 Austr. The fls. of some are almost black. Cult. orn. fl. Kenopleurum Candargy. Umbelliferae (in. 6). i Lesbos. Kentia Blume. Palmae (iv. i). 10 Moluccas to N.Z. (not in Austr.). Fls. in groups of 3 (2 male) on the spadix. Kentiopsis Brongn. Palmaceae (iv. i). 2 New Caledonia. Kentranthus Neck. = Centranthus DC. (Valerian.). Kentrochrosia Lauterb. et K. Schum. Apocynaceae (i. 3). i New Guinea. Kentrophyllum Neck. =Carthamus L. (Comp.). Kentrosphaera Volkens. Amarantaceae (2). i Kilimandjaro. Keracia Calest. (Hohenackeria p.p.). Umbelliferae (111.5). ' W.Medit. Keramanthus Hook. f. (Adenia p.p. EP.}. Passifloraceae. i trop. Afr. Keraselma Neck. = Euphorbia L. (Euph.). Keratto, Agave. Keraudrenia J. Gay. Sterculiaceae. 8 Austr., Madag. KerberaE. Fourn. (Alclinia EP.}. Asclepiadaceae(n, i). 2Braz., Urug. Kerguelen cabbage, Pringlea antiscorbulica R. Br. Kerinozoma Steud. Gramineae (12). i Java. Kermadecia Brongn. et Gris. Proteaceae (n). 5 New Cal., E. Austr. Kernera Medic. (Cochlearia L. p.p.). Cruciferae (2). 5 Alps. Kerneria Moench. =Bidens Tourn. p.p. (Comp.). ' Kerria DC. Rosaceae (in), i E. As., K. japonica DC., often cult. orn. shrub. Kerstingia K. Schum. Rubiaceae (n. j). i Togoland. Kerstingiella Harms. Leguminosae (in. 10). i Togoland. Geocarpic. Keteleeria Carr. Coniferae (Pinac. ; see C. for gen. char.). 4 China. Keyserlingia Bunge ex Boiss. (Sophora p.p. EP.}. Legum. (in. i). 2 W.As. Keysseria Lauterb. Compositae (3). i New Guinea. Khas-khas, Vetiveria zizanioides Stapf. Khat, Cat ha cdnlis Forsk. Knaya A. Juss. Meliaceae (n). 4 trop. Afr. Kibara Endl. Monimiaceae. 15 Indomal. Kibessia DC. Melastomaceae (n). 16 Malaya. Kickxia Blume. Apocynaceae (11. i). 2 Java. Kidney-bean, Phaseolus vulgaris L. ; -fern, Trichomanes ; -vetch, An thy I Us vulneraria L. Kielmeyera Mart. Guttiferae (i). r 8 S. Brazil, char, of campos. Kigelia DC. Bignoniaceae (4). 3 trop. Afr., Madag. The infls. are borne on old wood, hanging down on very long stalks. w. 23 354 KIGELIANTHE Kigelianthe Baill. Bignoniaceae (2). 2 Madag. Kiggelaria L. Flacourtiaceae (3). 5 S. and trop. Afr. K. Dregeana Turcz. yields a good timber (Natal mahogany). Kinepetalum Schlechter. Asclepiadaceae (n. 3). i Afr. Kinetostigma Dammer. Palmaceae (iv. i). i C. Am. King-cup, Caltha palustris L. Kingdonia Balf. f. et W. W. Smith. Ranunculaceae (3). i China. Kingia R. Br. Liliaceae (in) (June. BH.}. i W. Austr., a char. pi. Kingiodendron Harms. (Hardivickia p.p.). Leguminosae (n. 2). i Indomal. Kingstonia Hook. f. et Thorns. Anonaceae (i). Malay Peninsula. Kinia Rafin. Inc. sed. i Borneo. Kinnikinnik (Am.), Cornus sericea L. Kino, a resin-like substance, soluble in water, astringent, used medi- cinally and in tanning, Butea, Eucalyptus, Pterocarpus, &c. Kirengeshoma Yatabe. Saxifragaceae (in), i Japan. Kirganelia Juss. = Phyllanthus L. p.p. (Euph.). Kirilowia Bunge. Chenopodiaceae (A), i Turkestan, Afghanistan. Kirkia Oliv. Simarubaceae. 4 trop. and S. Afr. Kissenia R. Br. ex T. Anders. Loasaceae. i Arabia, S. Afr. KissodendronSeem. (f/edt'rap.p. BH.}. Araliaceae(i). i Austr., N.G. Kitaibelia Willd. Malvaceae (i). i lower Danube. Cult, orn. fl. Kitchingia Baker. Crassulaceae. 10 Madag. Kitul (Ceylon), Caryota itrens L. Klaineanthus Pierre ex Prain. Euphorbiaceae (A. n. i). i Gaboon. Klainedoxa Pierre. Simarubaceae. 6 trop. Afr. Klaprothia H. B. et K. Loasaceae. i N. trop. S. Am. Klattia Baker. Iridaceae (n). 2 Cape Colony. Kleinhovia L. Sterculiaceae. i trop. As. Kleinia Jacq. = Porophyllum Vaill. (Comp.). Kleinia L. = Senecio Tourn. (Comp.). Klopstockia Karst. = Ceroxylon Humb. p.p. (Palm.). Klossia Ridley. Rubiaceae (i. 2). i Malay Peninsula. Klotzschia Cham, et Schlechtd. Umbelliferae (i. 2). 2 S. Brazil. Klugia Schlechtd. Gesneriaceae (i). 4 trop. As. Markedly aniso- phyllous, with heterodromous fls. (cf. Cassia, Saintpaulia). Knappia Sm. = Mibora Adans. (Gram.). Knapweed, Centaurea Cyanus L. Knautia L. (Scabiosa p.p. BH.}. Dipsaceae. 25 Medit., Eur. Knawel, Scleranthus annuus L. Knees, cf. Taxodium. Kneiffia Spach (Oenothera p.p. BH. }. Onagraceae(2). 10 temp. N. Am. Cult. orn. fl. Knema Lour. (Myristica p.p. BH.}. Myristicaceae. 40 Indomal. Knesebeckia Klotzsch = Begonia L. p.p. (Begon.). Knife-grass (W.I.), Scleria latifolia Sw. Knightia R. Br. Proteaceae (n). 3 New Zealand, New Caled. K. excelsa R. Br. (rewa; N.Z.) furnishes a beautiful timber. Kniphofia Moench. Liliaceae (ill). 25 S. Afr., cult. orn. fl. Bees sometimes force their way into fls. and are unable to return. Knol-kohl, Brassica oleracea L. , var. KOSLOVIA 355 Knotgrass, Polygnnum, (Am.) Paspahnn distichum L. ; -weed (Am.) Polygonitin. Knowltonia Salisb. (Anemone p.p. EP.}. Ranunculaceae (3). 6 S. Afr. Knoxia L. Rubiaceae (li. i). 9 Indomal. Koanophyllon Arruda. Compositae (inc. sed.). i Peru. Kobresia Willd. Cyperaceae (in). 15 N. temp. |# (i Brit.). Kochia Roth. Chenopodiaceae (A). 35 N. temp., Austr., S. Afr. Kochiophyton Schlechter ex Cogn. (AcacaUis Lindl.). Orchidaceae (n. 13). i Amazonas. Koeberlinia Zucc. Capparidaceae (n) (Simarub. BH.}. i Texas, Mexico. A leafless xerophyte with thorny twigs. Cf. next article. Koeberliniaceae. Dicot. (Archichl. Parietales). A fam. made for the genus Koeberlinia (y.v.), now placed in Capparidaceae. See Nat. Pfl. Koechlea Endl. (Cirshun p.p. EP.}. Compositae (11). i Mt. Taurus. Koehneola Urb. (Tetranthus p.p-)- Compositae (5). i Cuba. Koeleria Pers. Gramineae (10). 25 temp, (i Brit.). Koellensteinia Reichb. f. (Aganisia p.p. BH.}. Orchid, (n. 14). i Colombia. Koellia Moench. (Pycnanthemum Michx.). Labiatae (vi). ic, S. U.S. Koellikeria Regel. Gesneriaceae (n). i Colombia. Koelpinia Pall. Compositae (13). 3 N. Afr. to E. As. Koelreuteria Laxm. Sapindaceae (n). 3 China. The capsule is large and bladdery and may be blown about by wind (cf. Colutea). Koenigla L. (Polygonwn p.p. BH.}. Polygon, (i. i). i Arctic, Himal. Koernickea Klotzsch (Paulh'niap.p. EP.}. Sapindaceae (i). i Guiana. Kohautia Cham, et Schlechtd. =Oldenlandia L. p.p. (Rubi.). Kohleria Regel (Isoloma BH.}. Gesner. (n). 40 trop. Am. Cult. orn. fl. KoMerianthus Fritsch (Fhtckigeria p.p.). Gesneriaceae (i). i Bolivia. Kohl-rabi, Brassica oleracea L., var. Kokia Lewton (Gossypiuin p.p.). Malvaceae (4). i Hawaiian Is. Kokoona Thw. Celastraceae. 3 Indomal. Kola nut, Cola aciiminata Schott. et Endl., C. vera K. Schum. Kolkwitzia Graebn. Caprifoliaceae. i China. Kolobochilus Lindau. Acanthaceae (iv. B). 2 Costa Rica. Kolobopetaliun Engl. Menispermaceae. 4 trop. Afr. Kolowratia Presl (Alpinia p.p. BH.}. Zingiberaceae (l). i Phil. Is. Komaroffia O. Ktze. (Nigella p.p.)- Ranunculaceae (i}. i Turkestan. Kompitsia Costantin et Galland. Asclepiadaceae (i). i Madag. Koniga R. Br. = Alyssum Tourn. (BH.} = Lobularia Desv. Koompassia Maingay (Abauria Becc.). Leguminosae (n. ,s). 2 Malaya. Koordersina O. Ktze. = Koordersiodendron Engl. (Anacard.). Koordersiodendron Engl. Anacardiaceae (2). i Celebes. Kopsia Blume. Apocynaceae (i. 3). 10 Indomal. Korshinskia Lipsky. Umbelliferae (in. 5). i C. As. Korthalsella Van Tiegh. (Bifaria p.p.). Loranthaceae (n). 20 Poly- nesia to Madag. Korthalsia Blume. Palmae (m). 20 Indomal. Some, e.g. K. horrida Becc., are said to be myrmecophilous (cf. Cecropia), the ants living in the sheaths of the leaves. Koslovia Lipsky (Albertia p.p.). Umbelliferae (in. 4). i Turkestan. 23—2 356 KOSMOSIPHON Kosmosiphon Lindau. Acanthaceac (iv. A), i trop. Afr. Koso, Brayera, Hagenia. Kosteletzkya C. Presl. Malvaceae (4). %•%_. Kostyczewa Korshinsky. Leguminosae (in. 6). i Turkestan. Kotchubaea Fisch. Rubiaceae (i. 8). i N. Brazil, Guiana. Kraenzlinella O. Ktze. (Otopetalum Lehm. et Kranzl.). Orchidaceae (n. 1 6). i Ecuador. Kralikia Coss. et Dur. Gramineae (12). i Algeria. Krameria Loefl. Leguminosae (n. 6). 13 Mexico to Chili. Placed in Polygalaceae by BH., but has a pet., not a sep., post. K and C 4 — 5; A 4, anthers opening by pores. Eichler, Bliithendiag. p. 522. Krascheninikowia Coss. et Dur. = Stellaria L. p.p. (Caryophyll.). Kraunhia Kafin. (Wistaria Nutt.). Leguminosae (in. 6). 4 E. As., E. N. Am. Kraussia Harv. (Tricalysia p.p. EP.). Rubiaceae (i. 8). 3 trop. and S. Afr. Kreysigia Reichb. Liliaceae (i). i S.E. Austr. Krigia Schreb. Compositae (13). 6 N. Am. Krokeria Moench. = Lotus L. p.p. (Legum.). Krugella Pierre (Poitteria p.p. EP.). Sapotaceae (i). i Trinidad. Krugia Urb. (Marlierea p.p.). Myrtaceae (i). i W.I. Krugiodendron Urb. (CeanotJnis p.p.). Rhamnaceae. i W.I. Krynitzkia Fisch. et Mey. (Cryptanthe EP.). Boragin. (iv. 2). 45 W. Am. Kugia Berl. Inc. sed. Nomen. KubUaasseltia J. J. Smith. Orchidaceae (n. 2). 2 Malay Archip. Kuhlia H. B. et K. (Banara EP.). Flacourtiaceae (5). 6 Am. Kuhnia L. Compositae (2). 3 Arizona to Mexico. Kuhnistera Lam. (Petalostemon Michx.). Leguminosae (in. 6). 20 N. Am. KumlieniaGreene(6Uj£ra//«.r EP.). Ranunculaceae(3). iN.W.Am. Kummeria Mart. (Discofhora Miers). Icacinaceae. 2 Brazil, Guiana. Kummerowia Schindler (Lespedeza p.p., q.v.). Legumin. (ill. 7). i Japan. Kumquat, Citrus japonica Thunb. Kundmannia Scop. Umbelliferae (in. 5). i Medit., S. Eur. Kunstleria Prain. Leguminosae (ill. 8). 5 Malay Peninsula. Kunthla Humb. et Bonpl. (Chamaedorea p.p. BH.). Palm. (iv. i). i N.W. S. Am. Kunzea Reichb. Myrtaceae (n. i). 18 Austr. Kunzmannia Klotzsch et Schomb. Rutaceae (inc. sed.). i Guiana. Kurakkan (Ceylon), Eleusine Coracana Gaertn. Kurdee seed, Carthamus tinctorius L. Kurrimia Wall. Celastraceae. 8 Indomal. Kurzamra O. Ktze. (Soliera Clos). Labiatae (vi). i Chili. Kuschakewiczia Regel et Smirn. Boraginaceae (iv. i). i Turkestan. Kutchubaea Fisch. ex DC. Rubiaceae (i. S). i N. Brazil, Guiana. Kutira gum, Cochlospermum, Sterciilia. Kydia Roxb. Malvaceae (2). 2 India. Kyllinga Rottb. Cyperaceae (i). 50 trop. and subtrop. Some have aromatic roots. LABIATAE 357 Kyrstenia Neck. (Eupaloiiutn Tourn. p.p.)- Compositae (2). 50 N. Am. Labatia Sw. Sapotaceae (i). 6 trop. S. Am., W.L Labdanum, Cislits creticus L. , C. ladaniferus L. Labellum, Caniia, Muranfaceae, Orchidaceae, Stylidiaceae, Zingiberaceae. Labiatae (EP., J3H.}. Dicotyledons (Sympet. Tubiflorae; Lamiales BH.}. 200 gen. 3000 sp., cosmop. ; chief centre the Medit. region. Some small groups are localised in their distribution, e.g. § II. in Austr. and Tasmania, III. in India, Malaya, China, &c., VIII. in Centr. Am., whereas the large ones, such as I. and IV., are cosmop. Most L. are land-plants, and herbs or undershrubs, similar in habit and structure. Stem usu. square, with decussate simple exstip. L, often hairy and with epidermal glands secreting volatile oils, which give char, scents to many. A few marsh-plants (Mentha, Lycopus, &c. ), a few climbers (Stenogyne sp., Scutellaria. &c.), and a few small trees (Hyptissp.). Manyxero. with reduced, sometimes infolded, 1., hairi- ness, thick cuticles, &c. , e.g. Rosmarinus. The axis of the first order is not closed by a fl. but only those of later orders; thus the primary form of the infl. is racemose, and a simple raceme actually occurs in Scutellaria, &c. Usu. however a di- chasial cyme, becoming cincinnal in its later branchings, occurs in the axil of each 1. upon the upper part of the main axis. In Teucrium, Nepeta sp. , &c., the construction of this cyme is easily seen ; but in most L. it is closely 'condensed' into the axil, so that all the fls. are sessile; but it is easily seen that the central fl. opens first and then those on either side of it (see diagram). The two condensed cymes at each node overlap the leaf- axils and often form what looks like v\ c T , , c a ... . a . r * loral diagram of Lamuim album a whorl of Ms. ; this infl. IS often with indication of dich. double called a verticillaster or false whorl. cincinnus at the sides. (After Fl. 5 or gynodioec., -|-,hypO!J., Eichler.) The asterisk repre- • J r ° sents the missing posterior sta 5-merous with suppression in some whorls. Usu. formula K (5), C (5), A 4, G (2). K tubular, bell- or funnel-shaped, sometimes 2-lipped, persistent in fr. ; C usu. 2-lipped with no clear indication of the individual petals; A 4, didynamous, or of nearly equal length, sometimes 2, epipet. with intr. anthers. G on a nectariferous disc (often developed on anterior side only), of (2) cpls. placed antero-post. Early in development a constriction appears in the ovary in the antero-post. line, dividing each cpl. into 2 loculi, so that the ovary becomes 4-loc. as it matures. Each of the 4 portions is nearly independent of the rest, and the style springs between them from the base of the ovary (i.e. is gynobastc) ; stigma 2-lobed. Placentae axile, each with i basal erect anatr. ovule with ventral raphe. Fr. usu. a group of 4 achenes or nutlets, each containing one seed ; sometimes a drupe. Seed with no enclosp. or very little; the radicle of the embryo points downwards (cf. Boraginaceae). 358 LAB I AT A E The fls. belong in general to classes H. and F. The 2-lipped C ensures that the visiting insect shall take a definite position in regard to the anthers and stigma whilst probing for the honey at the base of the fl. The lower lip acts as a flag to attract, and also as a landing- place, whilst the upper lip shelters the essential organs, which are usu. placed so as to touch the insect's back. The length of the C-tube varies very much, and with it the kind of visitors. Most Brit. sp. are bee fls., the long-tubed red fls. of Monarda &c. are butterfly fls., and a few sp. of Sal via &c. are humming-bird fls. The pollinatiun-mech. is usu. simple; in Lamium, &c. the fl. is homo- gamous, the stigma merely projecting beyond the anthers so as to be touched first, but usu. the fl. is dichogamous (protandr.), often with movements of the essential organs, e.g. in Teucrium, &c. The lever- mechanism of Salvia is almost unique. Thymus, Origanum, and their allies, have nearly regular fls. visited by a more miscellaneous selection of insects. In many L., esp. g VI., interesting distrs. of sex appear, esp. gynodioecism. A few disperse their fr. by aid of the persistent bladdery K, or by hooks formed from the K teeth. The stalks are often hygroscopic and move in such a way as to favour dispersal in wet weather. Useful on account of their volatile oils; many, e.g. Thymus, Ocimum, Origanum, Salvia, &c., used as condiments. Oils and perfumes are obtained by distillation from Rosmarinus, Pogostemon, Lavandula, &c. Food products from Stachys sp. Classification and chief genera (after Briquet, from whose account much of the above is condensed) ; closely allied to Verbenaceae ; from Boraginaceae the position of the radicle sharply separates them, whilst the similarity to Scrophulariaceae, &c. is largely in minor chars. A. Style not gynobasic. Nutlets with lateral-ventral attachment and usu. large surface of contact (often >\ as high as ovary). I. AJUGOIDEAE (seed exalb.) ; i . Ajugeac (corolla various ; upper lip if present rarely concave ; sta. 4 or 2 ; anther 2-loc. ; nutlets ± wrinkled): Ajuga, Teucrium. 2. Rosmarineae (corolla strongly 2-lipped; upper lip very con- cave and arched ; sta. 2 ; anthers i-loc. ; nutlets smooth) : Rosmarinus (only genus). II. PROSTANTHERO1DEAE (seed albuminous): Prostan- thera. B. Style perfectly gynobasic. Nutlets with basal attachment and usu. small surface of contact, rarely with ± basal-dorsal attachment. III. PRASIOIDEAE (nutlet drupaceous with fleshy or very thick exocarp and hard endocarp) : Stenogyne, Gomphostemma. IV. SCUTELLARIOIDEAE (nutlet dry; seed ± transversal; embryo with curved radicle lying on one cot. ) : Scutellaria. V. LA VANDULOIDEAE (nutlet dry ; seed erect ; embryo with short straight superior radicle ; disc-lobes opp. toovary-lobes; nutlets with ± distinct dorsal-basal attachment ; sta. 4 in- cluded; anthers i-loc. at tip through union of thecae): Lavan- dula (only genus). LACHNOLOMA 359 VI. STACHYDOIDEAE (ditto, but disc-lobes, when distinct, alt. with ovary-lobes ; nut with small basal attachment ; sta. ascending or spreading and projecting straight forwards) : Marrubium, Nepeta, Dracocephalum, Prunella, Phlomis, Galeopsis, Lamium, Ballota, Stachys, Salvia, Monarda, Ziziphora, Horminum, Calamintha, Satureia, Origanum, Thymus, Mentha, Pogostemon. VII. OCIMOIDEAE (as VI., but sta. descending, lying upon under lip or enclosed by it) : Hyptis, Ocimum. VIII. CATOPHERIOIDEAE (nutlet dry; seed erect; embryo with curved radicle lying against the cotyledons): Catopheria. Labiate (C), with projecting lip. Labichea Gaudich. ex DC. Leguminosae (n. 5). 5 Austr. Labidostelma Schlechter. Asclepiadaceae (n. 4). i Guatemala. Labile, plastic. Labisia Lindl. Myrsinaceae (n). 5 Malay Archip. Lablab Adans. (Dolichos Bff.}. Leguminosae (ill. 10). i trop. Afr., L. vulgaris Sair, largely cult, in trop. for ed. pods. Labordia Gaudich. Loganiaceae. 9 Hawaiian Is. Labourdonnaisia Boj. Sapotaceae (i). 3 Mauritius, Natal. Labrador tea (Am. ), Ledum palustre L. Labramia A. DC. (Mimusops p.p. EP.). Sapotaceae (2). i Madag. Laburnum L. Leguminosae (in. 3). 3 Eur. , W. As. L. zntlgare J. Presl, the common laburnum. The fl. has a simple Trifolium- mech. There is no free honey ; bees pierce the swelling at the base of the vexillum (cf. Orchis). All parts are poisonous. Lac, a resin formed as an excretion from the skin of the lac insect, living on Butea, Croton, Ficus, Schleichera, &c. Lacaena Lindl. Orchidaceae (n. 13). 2 Mexico, C. Am. Cult. orn. fl. Lacaitaea Brand ( Trichodesma p. p.). Boragin. (iv. i). i Sikkim. Laccodiscus Radlk. (Cupania p.p. Bff.). Sapindaceae (i). i trop. W. Afr. Laccopetalum Ulbrich (Anemone p.p.). Ranunculaceae (3). i Peru. Laccosperma G. Mann et H. Wendl. (Ancistrophyllum p.p.). Palma- ceae (in), r W. Afr. Lace-bark (W.I.), Lagetta Lintearia Lam. Lacerate, deeply and irregularly divided. Lachemilla Rydberg (Alcheinilla p.p.). Rosaceae (in. 5). 15 N. Am. Lachenalia Jacq. Liliaceae (v). 35 S. Afr. L. tricolor Jacq. a little bulbous plant with two leaves, cult. orn. fl. Lachnaea L. Thymelaeaceae. 18 S. Afr. Lachnagrostis Trin. = Deyeuxia Clar. (Bff.) = Calamagrostis p.p. Lacb.nantn.es Ell. Haemodoraceae. i N. Am., L. tinctoria Ell., the paint-root. The roots yield red dye. Orig. of Species, 6th ed., p. 9. Lachnastoma Korth. (Coffea p.p.). Rubiaceae (n. 4). i Java. Lachnocapsa Balf. f. Cruciferae (4). i Socotra. Lachnocaulon Kunth. Eriocaulaceae. 4 S.E. N.Am. Lacnnocephalus Turcz. (Mallophora p.p.). Verben. (3). 5 temp. W. Austr. Lachnochloa Steud. Gramineae (inc. sed.). i Senegambia. Lachnoloma Bunge. Cruciferae (r). i Turkestan. 360 LACHNOPHYLLUM Lachnophyllum Bunge. Compositae (3). 2 W. As. Lachnorhiza (Vernonia p.p. EP. ). Compositae (i). i Cuba. Lachnospermum Willd. Compositae (4). i S. Afr. Lachnostachys Hook. Verbenaceae (3). 8 Austr. Lachnostoma H. B. et K. Asclepiada'ceae (n. 4). 6 warm Am. Lachnostylis Turcz. Euphorbiaceae (A. I. i). i Cape Colony. Laciala O. Ktze. (Schizoptera EP.}. Compositae (5). i Ecuador. Laciniaria Hill = Liatris Schreb. (Comp.). Laciniate. divided into several long segments. Lacis Schreb. Podostemaceae. r Amazon. Lacistema Sw. Lacistemaceae. 16 trop. Am., W.I. Shrubs with distichous exstip. 1., and spikes of very small fls. $, naked or with one whorl of P, with concave axis. A i, 0(2 — 3) with parietal plac. and i — -2 pend. ov. on each. One-seeded caps. Endosp. Lacistemaceae (EP., BH.). Dicots. (Archichl. Piperales; Monochlam. Anom. BH.}. Only genus Lacistema, q.v. Closelyallied to Piperaceae. Lacmellia Karst. Apocynaceae (i. i). i trop. S. Am. Lacquer, Japan, Rhus vernicifera DC. Lactoridaceae (EP. ; Piperaceae p.p. BH.}. Dicotyledons (Archichl. Ranales). Only gen. Lactoris, q.v. Lactoris Phil. Lactoridaceae. i sp., L. fernandeziana Phil., Juan Fernandez. Shrub with 3-merous fls. P 3, A 3 + 3, 63. Lactuca (Tourn.) L. (excl. Mulgedium Cass.). Compositae (13)- 10%, chiefly N. temp. Old World; 4 in Brit. L. Scariola L. (prickly lettuce) is a compass-plant in dry exposed places (see Silphium); rapidly spreading in the U.S. as a weed. L. sativa L. is the lettuce of salads. Fl. like Hieracium. Lacuna, an open space in tissue. Lacustris (Lat.), living in lakes. Ladanum, Cistus creticus L., C. ladaniferits L. Ladders, cf. Bamboos. Ladenfoergia Klotzsch (Cascarilla Wedd. p.p.). Rubiaceae (i. 5). 10 S. Am. Bark astringent, containing alkaloids. Lady-fern, Athyrhun Filix-femina Roth. ; -'s fingers, Anthyllis vul- neraria~L.\ -slipper orchid, Cypripedinm\ -thumb (Am.), Polygonum Persicaria L. ; -tresses, Spiranthes aiitu)>malis Rich. Ladyginia Lipsky. Umbelliferae (ill. 6). i C. As. Laelia Lindl. Orchidaceae (n. 6). 25 trop. Am. Often epiphytic. Laeliocattleya x Rolfe. Orchid hybrid, Laelia and Cattleya. Laestadia Kunth. Compositae (3). 4 trop. Andes. Laetia Loefl. Flacourtiaceae (7). 10 trop. Am., W.I. Laevigatus, laevis (Lat.), smooth. Lafoenaia Vand. Lythraceae. 10 trop. Am. Lafuentia Lag. Scrophulariaceae (in. i). i S. Spain. Lagarinthus £. Mey. = Schizoglossum E. Mey. (Asclep.). Lagarosiphon Harv. Hydrocharidaceae. 9 Afr., Madag. Lagascea Cav. Compositae (5). 10 C. Am. Lagenandra Dalz. Araceae (vn). 5 Ceylon, S. India. Lagenanthus Gilg. Gentianaceae (i). i Colombia. Lagenaria Ser. Cucurbitaceae (3). i palaeotrop., L. vulgaris Ser. (calabash-cucumber). The outer woody pericarp makes a flask. LAMPRODITHYROS 361 Lagenia E. Fourn. (Arattjia p.p.). Asclepiadaceae (n. i). 2 Brazil. Lagenias E. Mey. (Sebaea p.p. BH.~). Gentianaceae (i). i Cape Colony. Lagenocarpus Nees. Cyperaceae (n). 15 trop. S. Am. Lagenophora Cass. Compositae (3). 15 japan to N.Z., Chili. Lagerstroemla L. Lythraceae. 25 palaeotrop. Some heterostyled like Lythrum. Cult. orn. fl. trees. Some provide good timber. Lagetta Juss. Thymelaeaceae. 3 W. Ind. L. Lintearia Lam. is the lace tree. Its bast-fibres on removal from the stem (by macera- tion, &c.) form a network used for making dresses, &c. Laggera Sch.-Bip. ex Hochst. Compositae (4). 12 palaeotrop. Lagoa Durand. Asclepiadaceae (II. i). i Brazil. Lagochilium Nees= Aphelandra R. Br. (Acanth.). Lagochilus Bunge. Labiatae (vi). 15 W. As. Lagoecia L. Umbelliferae(n. 2). i Medit. One of the usu. two loc. of the ovary is aborted. Lagophylla Nutt. Compositae (5). 6 W. N.Am. Lagos rubber, Funtumia elastica Stapf. Lagoseris Hoffmgg. et Link = Crepis L. p.p. (Comp.). Lagotis J. Gaertn. Scrophulariaceae (in. i). 10 N. and C. As. Laguna Cav. = Hibiscus L. (Malv.). Lagunaria G. Don. Malvaceae (4). i E. Austr., Norfolk L, Howe I. Laguncularia Gaertn. f. Combretaceae. i W. trop. Afr., Am. (mangrove). Lagurus L. Gramineae (8). j Medit., L. ovatus L., cult. orn. Laliia Hassk. (Dttrio p.p. EP.}. Bombacaceae. i Borneo. Lallemantia Fisch. et Mey. Labiatae (vi). 4 W. As. Lamarchea Gaudich. Myrtaceae (n. i). i W. Austr. Lamarckia Moench. Gramineae (10). i Medit. Cult. orn. Lamb-kill (Am.), Kalmia angustifolia L. ; -'a lettuce, Vakrianella, (\m.) Fedia; -quarters, Chenopodiuni. Lambertia Sm. Proteaceae. 8 Austr. Lamellisepalum Engl. Rhamnaceae. i trop. Afr. Lamiacanthus O. Ktze. Acanthaceae (iv. A), i Java. Lamiales (/?//.). The roth order of Gamopetalae. Lamina, the blade of a I. Lamium (Tourn.) L. Labiatae (vi). 40 Eur., As., extratrop. Afr. 5 in Brit., incl. L. album L. (white dead-nettle), L. amplexicmtle L. (hen- bit), L. purpiireum L. (purple dead-nettle) and Z. Galeobdolon Crantz (yellow archangel). L. album has sympodial rhizomes and large white homogamous humble-bee fls. L. amplexicaitle\\v& cleist. fls. in spring and autumn ; they look like ordinary buds with a small C, and are pollinated without opening. Lamourouxia H. B. et K. Scrophulariaceae (in. 3). 20 trop. Am. Lampaya Phil. Verbenaceae (i). i Chili. Lampocarya R. Br. = Gahnia Forst. p.p. (Cyper.). Lamprachaenium Benth. Compositae (i). i Indomal. Lamprocaulos Mast. (Elegia p.p. EP.}. Restionaceae. 2 S. Afr. Lamprochlaenia Borner = Carex L. p.p. (Cyper.). Lamprococcus Beer (Aechmea p.p. BH.). Bromeliaceae (4). 8 trop. Am. Cult. orn. infl. Lamprodithyros Hassk. =Aneilema R. Br. p.p. (Commelin.). 362 LAMPROLOBIUM Lamprolobium Benlh. Leguminosae (in. 6). i Queensland. Lamprospermum Klotzsch (Matayba EP. ). Sapindaceae (i). 2 Guiana. Lamprothamnus Iliern. Rubiaceae (11. i). i Zanzibar. Lamprothyrsus Pilger. Graniineae (9). i S. Am. Lamprotis D. Don = Erica Tourn. p.p. (Eric.). Lampsana (Tourn.) Rupp. = Lapsana L. (Comp.). Lanaria Ait. Amaryllidaceae (in) (Haemodor. BH.}. i S. Afr. Lanatus (Lat.), woolly. Lance, Dendrocalaimis ; -wood, Duguetia, Oxandra, Tonmefortia, &c. Lancea Hook. f. et Thorns. Scrophulariaceae (n. 6). 2 Tibet, China. Lanceolate (1.), 3 times as long as broad, tapering gradually. Landolphia Beauv. Apocynaceae (i. i). 30 trop. and S. Afr. Several are lianes with curious hook tendrils like Strychnos. Fr. a large berry full of an acid pulp composed of the hair-structures on the seeds. Several, e.g. L. Kirkii Dyer, L. comorensis Benth. et Hook, f., &c., yield rubber, the coagulated latex. It is known in trade as African and Madagascar rubber. Landtia Less. (Arctotis p.p. EP.}. Compositae(io). 4 S. Afr., Abyss. Landukia Planch. ( Vitis p.p.). Vitaceae. i Java. Lanessania Baill. Moraceae (n). i N. Brazil. Langlassea H. Wolff. Umbelliferae (in. 6). i Mexico. Langloisia Greene (Gilia p.p.). Polemoniaceae. 4 S.W. U.S. Langsdorffia Mart. Balanophoraceae. i trop. Am. L. kypogaea Mart. LamumLindl.(.£//Vfc«^rtt/wp.p. EP.}. Orchidaceae (n.6). 4 trop. Am. Lankesteria Lindl. Acanthaceae (iv. A). 5 trop. W. Afr. Lannea A. Rich (Odina BH., Calesium EP.}. Anacardiaceae (2). 15 palaeotrop. Lanneoma Delile (Lannea p.p.). Anacardiaceae (2). i Abyss., E. Afr. Lanose, lanuginose, woolly. Lansbergia De Vriese = Trimezia Salisb. (Irid.). Lansium Rumph. Meliaceae(ni). 4lndomal. L.domesticum Jacked.fr. Lantana L. Verbenaceae (i). 60 trop. and subtrop. Shrubs, often used for hedges. Some have ed. fr. Lantanopsis Wright. Compositae (5). 3 Cuba, S. Domingo. Lapageria Ruiz et Pav. Liliaceae (x). i Chili, L. rosea Ruiz et Pav., a climbing shrub with ed. fr., cult. orn. fl. Lapatero (W.I.), Copaifera officinalis L. Lapathum (Tourn.) Adans. = Rumex L. p.p. (Polygon.). Lapeyrousia Pourr. Iridaceae (in). 30 S. Afr. to Abyss. Cult. orn. fl. Laphamia A. Gray. Compositae (6). 15 S. U.S., Mexico. Lapiedra Lag. Amaryllidaceae (i). 2 Spain, Morocco. Lapithea Griseb. (Sabbatia BH.}. Gentian, (i). i Carolina to Texas. Laplacea H. B. et K. (Haemocharis EP.}. Theaceae. 25 trop. As., Am., W.I. Laportea Gaudich. Urlicaceae (i). 30 trop. Many sting violently. Lappa (Tourn.) Rupp. = Arctium L. (Comp.). Lappago Schreb. =Tragus Hall. (Gram.). Lappula Moench. (Echinosperimtm BH.). Boragin. (iv. 2). 60 temp. Lapsana L. Compositae (13). 9 N. temp. %. /.. communis L. (nipplewort) in Brit. The fls. are inconspicuous and pollinate them- selves. There is no pappus. LA STOPE TA LUM 363 Larch, Larix. Laretia Gill, et Hook. Umbelliferae (i. 2). i Andes of Chili. Laricopsis Kent. = Pseudolarix Gord. Coniferae (Pinac.). Lardizabala Ruiz et Pav. Lardizabalaceae. 2 Chili. Tough fibre from the stems of L. biternata Ruiz et Pav. Lardizabalaceae (EP. ; Berberidaceae p.p. Bff.). Dicots. (Archichl. Ranales). 7 gen. 15 sp. Himal. to Japan, Chili. Mostly climbing shrubs with palmate 1. Fls. in racemes, usu. in the axils of the scale-1. at the bases of the branches, polygamous or diclinous. Usual for- mula P 3 -t- 3, A 3 + 3, G 3 or more. 2 whorls of small honey-leaves (see Ranunculaceae) often occur between P and A ; sta. sometimes united ; anthers extrorse ; ovules oo in longitudinal rows on the lat. walls (cf. Nymphaeaceae), anatr. The fl. of either sex shows rudi- ments of the organs of the other sex. Berry. Embryo small and straight, in copious endosp. Chief genera: Decaisnea, Akebia, Lardizabala. Larix Tourn. ex Adans. Coniferae (Pinac. 20; see C. for genus cha- racters). 8 Eur., N. As., N. Am. The general chars, are those of Cedrus, but the 1. are deciduous, and the cones ripen in a single year. L. europaea DC. (L. decidua Mill.) is the common larch, cult, on a large scale for its wood, bark (used in tanning) and turpentine (Venice t.). Others are also important, e.g. L. americana Michx. (tamarack). Larkspur, Delphinium. Larochea Pers. = Crassula Dill. p.p. (Crass.). Larrea Cav. Zygophyllaceae. 4 subtrop. Am. Xerophytes. L. mexi- cana Moric. (Mexico, &c.) is the creosote plant, which forms a dense scrub-veg. and binds the drifting sand together. Its strong smell prevents it from being eaten by animals. Larrea scrub, a scrub association char, by Larrea mexicana Moric. Lasallea Greene (Aster p.p.). Compositae (3). 3 N. Am. Lascadium Rafin. Inc. sed. i S. U.S. Laseguea A. DC. Apocynaceae (n. i). 10 trop. S. Am. Laserpitium L. Umbelliferae (ill. 7). 35 Eur., N. Afr., As. Lasia Lour. Araceae (iv). 2 Indomal. Lasiacis Hitchcock (Panicum p.p.). Gramineae (5). 6 N. Am. Lasiadenia Benth. Thymelaeaceae. i Guiana. Lasiagrostis Link — Stipa L. (Gram.). Lasiandra DC. =Tibouchina Aubl. (Melast). Lasianthemum Klotzsch ( Talisia p.p. EP.}. Sapindaceae (i). 2 Guiana. Lasianthera Beauv. Icacinaceae. i trop. W. Afr. Lasianthus Jack. Rubiaceae (it. 5). 90 Indomal. Lasiocarpus Liebm. Malpighiaceae (i). r Mexico. Lasiochloa Kunth. Gramineae (ro). 4 S. Afr. Lasiocladus Boj. ex Nees. Acanthaceae (iv. B). 2 Madag. Lasiococca Hook. f. Euphorbiaceae (A. II. 2). i Sikkim. Lasiocoma Bolus. Compositae (8). i S. Afr. Lasiocorys Benth. (Lencas p.p. EP.). Labiatae (vi). 7 Afr. Lasiocroton Griseb. Euphorbiaceae (A. n. 2). i Jamaica. Lasiodiseus Hook. f. Rhamnaceae. 5 trop. Afr., Madag. Lasiopetalum Sm. Sterculiaceae. 25 Austr. 364 LASIOPOGON Lasiopogon Cass. Compositae (4). 3 S. Afr., Medit. Lasiorrliiza Lag. = Leuceria Lag. p.p. (Comp.). Lasiosiphon Fresen. (Giiidia EP.). Thymelaeaceae. 20 palaeotrop. Lasiospermum Lag. Compositae (7). 3 S. Afr. Lasiospora Cass. = Scorzonera Tourn. p.p. (Comp.). Lasiostelma Benth. Asclepiadaceae (n. 3). 4 S. Afr. Lasiostoma Schreb. — Strychnos L. (Logan.). Lassa O. Ktze. = Pavonia Cav. (Malv.). Lassonia Buchoz. Magnoliaceae. i China. Lastarriaca Remy. Polygonaceae (i. i). 2 Calif., Chili. Lasthenia Cass. Compositae (6). 10 W. Am. Lastrea Presl — Dryopteris Adans. (Folypod.). Latace Phil. Liliaceae (iv). i Chili. Latanla Comm. ex Juss. Palmae (n). 3 E. Afr., Mascarenes. Lateral branching, cf. Branch. Latex, a milky fluid contained in special vessels or cells of the tissues, Alismaceae, Apocynaceae, Araceae, Asclepiadaceae, Eitphotbiaceae, Compositae (Cichorieae), Galactia, JMoraceae, Sapotaceae, &c. Cf. Guttapercha, Rubber, Balata ; laticiferous, latex-bearing. Lathraea L. Orobanchaceae. 5 temp. Eur., As. L. Si/iiainaria L. in Brit, (tooth-wort) is a total parasite living upon the roots of hazel, beech, &c. It has a thick rhiz. bearing 4 rows of tooth-like scaly 1. The fl. shoot comes above ground and bears a raceme of purplish fls., all bent round to the same side of the infl., protogynous. The scales upon the rhiz. are hollowed, each containing a branched cavity open- ing to the outside by a narrow slit at the base of the back of the 1. This arises by adevelopment similar to that which forms the chambers in the 1. of Empetrum, Cassiope, &c. In the small lat. cavities opening out of the main one there are found peculiar glandular organs, resembling those of insectivorous plants. Small insects, &c. are often found in these leaves (cf. bladders of Utricularia, &c.) and it has been supposed that these organs absorb their proteids like the glands of Drosera, &c. This however is doubtful. L. Clandestina L. is parasitic upon willows. The capsule of L. splits explosively. [For details see Heinricher, Die Schitppenwurz, Stuttgart, 1908.] Lathriogyna Eckl. et Zeyh. Leguminosae (in. 3). i S. Afr. Lathrophytum Eichl. Balanophoraceae. i Rio de Janeiro. Lathyrus (Tourn.). L. (incl. Orobus L.). Leguminosae (in. 9). no N. temp., and Mts. oftrop. Afr. and S. Am. 10 in Brit, (pea), including L. Aphaca L. and L. Nissolia L. The former has large green stipules performing assim. functions, whilst the 1. is transformed into a tendril; the latter has its petioles flattened into phyllodes and has no 1. blade at all (see Acacia). L. macrorrhizns Wimm. has tuberous roots which may be eaten like potatoes. L. sativus L. ( Jarosse) and L. Cicera L. are cult, in S. Eur. as fodder and are also eaten like chick-pea (Cicer). L. odoratus L. is the sweet-pea. The fl. is like that of Vicia; on the style is a tuft of hairs that brushes the pollen out of the apex of the keel, where it is shed by the anthers. L. latifolius (everlasting pea) also cult. orn. fl. Lati- (Lat. pref.), broad; -folius, broad-leaved, &c. &c. Latipes Kunth. Gramineae (3). i Senegal to Sincl. LA UKIUM 365 Latouchea Franch. Gentianaceae (i). i China. Latouria Blume (Dendrobium p.p. BH.). Orchid, (n. 15). i New Guinea. Latreillea DC. = Ichthyothere Marl. (Comp.). Latrobea Meissn. Leguminosae (in. 2). 6 W. Austr. Latua Phil. Solanaceae (2). i Chili. Laubertia A. DC Apocynaceae (n. i). 2 Peru, Bolivia. Laugeria Vahl (Gnettarda p.p. EP.}. Rubiaceae (n. 2). 2 W.I. Launaea Cass. Compositae (13). 30 trop. and subtrop. L. pinnatifida Cass. a char, plant of sandy trop. beaches. Lauraceae (EP., BH'mcl. Hcrnandiaceae). Dicots. (Archichl. Ranales; Daphnales BH.). 40 gen. 1000 sp. tiop. and subtrop.; chief centres of distr. S.E. As. and Brazil. Trees and shrubs with leathery evergr. alt. exstip. 1. The tissues contain numerous oil-'cavities. Cassytha is an interesting parasite. Infl. racemose, cymose, or mixed. Fl.actinom., apet., usu. 3-merous, 5 or monoec. Formula usu. P2n, A4n, Gn. P in two whorls, peiig. ; A perig. or epig., in 3 or 4 whorls, some of which are commonly reduced to stds. ; anther usu. 4-loc. opening by valves (cf. Berberidaceae), usu. intr., but in many cases those of the third whorl extr. The axis is ± concave, and the ovary is free from it at the sides. G i (Payer, Baillon) or more probably 3 (Eichler), forming a i-loc. ovary, with i pend. anatr. ov. Fr. a berry, often ± enclosed by the cup-like recept., which also becomes fleshy in these cases. Embryo straight ; seed exalb. The position of the L. in the system is doubtful ; they apparently form the connecting link between the Ranales (to the more typical fams. of which they are linked by Monimiaceae and Calycanthaceae) and theThymelaeales. Important economic plants are found in nearly all the genera mentioned below. [See Nat. Pfl. and Mez. in Bot.Jahresb. 1889, p. 459, and Bot. Centr. 54, p. 275 (abstracts).] Classification and chief genera (after Pax) : I. PERSEOIDEAE (anther 4-loc.) : Cinnamomum, Persea, Sassafras, Litsea. II. LAUROIDEAE (anther 2-loc.): Cryptocarya, Lindera, Lauras, Cassytha. Laurel, Laurus nob His L. ; Alexandrian-, Calophyllum Inophyllum^.; bay-, Laurus nobilis L.; cherry-, Prumis Laurocerasusi,. ; Japan -, Aucuba japonica Thunb. ; Portugal-. Prumts lusitanica L. ; sea- side- (W. L), Phyllanlhus; spurge-, Daphne Lanreola L. ; West Indian- (W.I.), Prumts occidentalis Sw. Laurelia Juss. Monimiaceae. i N.Z. , i Chili. L. Novae- Zealandiae A. Cunn. supplies a useful timber. Tht frs. of L. arotnalica Juss. are used as a spice under the name Peruvian nutmegs. Laurembergia Berg. (Serpicula L.). Haloragidaceae. 20 palaeotrop. and S. Afr. Laurentia Michx. ex Adans. Campanulaceae(m). 10 Meclit., S. Afr. , N. Am. Laurestinus (Am.), Viburnum. Lauridia Eckl. et Zeyh. (Elaeodendron BH.}. Celastraceae. i S.Afr. Laurineae (BH.), Lauraceae. Laurium (CL), a drain formation. 366 LAUROCERASUS Laurocerasus (Tourn.), M. Roem. = Prunus L. p.p. (Ros.). Laurophyllus Thunb. (Botryceras BH.). Anacardiaceae (3). iS.Afr. Lauras (Tourn.) L. Lauraceae (n). 2, L. ttobilis L. the true laurel or sweet bay, Medit. (1. aromatic, used in condiments, &c., berries in veterinary medicine), and L. canariensis Webb et Berth., Canaries and Madeira. Fls. unisexual by abortion. Lautembergia Baill. (Alchorneay.p. BH.). Euph. (A. 11.2). 2 Madag. Lauterrjachia Perkins. Monimiaceae. i New Guinea. Lavalleopsis Van Tiegh. Olacaceae. 3 W. trop. Afr. Lavandula Tourn. ex L. Labiatae (v, q.v. for gen. char.). 20 Medit. to India. From L. veraDC. (lavender) is obtained oil of lavender, by distillation of the fls. ; it is used in painting, and in the manufacture of lavender water. L. Spica Cav. and L. Stoechas L. are also used. The protandr. fls. are visited by bees and form a good source of honey. Lavatera L. Malvaceae (2). 20 Medit., Austr., mid-As. L. arborea L., tree-mallow, on rocks on the Brit, coast. Lavauxia Spach (Oenothera p.p. BH.}. Onagraceae (2). 8 W. Am. Lavender, Lavandula vera DC. &c.; cotton-, Santalina; sea-, Statice. Lavenia Sw. = Adenostemma Forst. (Comp. ). Laviclia Phil. Compositae (12). i Patagonia. Lavigeria Pierre (Jcacina p.p.). Icacinaceae. i Cochinchina. Lavoisiera DC. Melastomaceae (i). 50 Brazil. Lavradia Veil, ex Vand. Ochnaceae. 6 Brazil. Lawia Griff, ex Tul. Tristichaceae. i Ceylon to Bombay Ghats. Thallus of shoot nature, creeping, with endogenous shoots on upper side. Lav/sonia L. Lythraceae. i palaeotrop. L. inerniis L. The powdered 1. form the cosmetic, henna, used in the East to stain the finger-nails, &c., red. Laxmannia R. Br. (Bartlingia F. Muell.). Liliaceae (m). 8 Austr. Laxus (Lat.), loose. Layia Hook, et Arn. Compositae (5). 14 W. N. Am. Cult. orn. fl. Leader, top bud of a monopodial shoot, Abies. Lead-tree (W.I.), Leucaena glanca Benth. ; -wort (W.I.), Plumbago. Leaf, a thin green expanded organ, borne on the stem at the nodes. The interior of an ordinary 1. is made up of a mass of spongy green tissue (iiiesophylt), covered by an epidermis which contains stoniata leading to the intercellular spaces in the interior, and whose outer wall is covered by a cuticle of waxy or corky nature. The vascular bundles that run through the 1. and hold it outstretched are called the veins or nerves. The 1. shows distinction into a base abutting on the stem and often bearing a pair of green or membranous expansions (stipules), and z.blade or lamina, often with a stalk or petiole between. Such a 1. has an upper or ventral surface, and a lower or dorsal, and is markedly dorsiventral in structure; it places itself ± horiz. in what is called \tefi.\ed light position. Others (isobi 'lateral, q.v.) have symmetrical structure on both sides, and place themselves with their edges to the light (phy/lodcs (q.v.) also come into this category), and yet others (centric, q.v.) have symmetrical structure all round, and stand with their apices to the light. Many 1. exhibit sleep movements LEAF 367 at night, or movements during heat or dryness (cf. Movements). The fall of a 1. is often effected by an absciss-layer, or stratum of cork which separates it from the stem, and then splits, leaving a leaf-scar upon the stem. L. with such a layer are called articulate, without, non-articulate. Under other than mesophytic conditions such structure as just described shows many modifications; cf. Climbing Plants, Insecti- vorous Plants, Parasites, Saprophytes, Water Plants, Xerophytes. It is also modified for Storage (g.v-) of reserves, or for Vegetative Reproduction (q-v.). In external form 1. show very great variety, and the grouping of gen. into sp. is largely determined by this, so that it is necessary to understand the use of the technical terms employed in such work, before one can use a flora with any advantage. An outline of such terms and their use follows. Descriptive Terms. The student should practise describing leafy shoots until expert in handling terminology, but there is no need to commit the terms to memory. At first he should describe in detail in the order given below, but afterwards try to render his descriptions short and pithy without sacrifice of essentials; this can only be well done by comparison with related forms to see what points are common to all. L. as to phyllotaxy (q.v-} or arrangement may be radical, or on the subaerial stem (cauline) ; whorled (i'erticillate), opposite (and then decussate if each pair is _L the next, connate if the two are con- crescent as in Lonicera, anisophyllous if unequal in size or shape), or alternate (the phyllotaxy fraction may be given, or the number of ranks described by the terms di-, tri-stichous, &c.). With regard to insertion or mode of union with the stem the 1. may be petiolate or sessile (i.e. with or without stalk respectively ; the petiole is descr. like a stem), auricled (with two lobes of the blade overlapping the stem), amplexicaul (the lobes clasping the stem), sheathing (as in Grasses, the leaf-base forming a tube round the stem), perfoliate (the leaf united round the stem, as in Bupleurum), Recurrent (continued by a wing on the stem, as in thistles), &c. It may bear a ligitle or scale at the upper end of the leaf-base or sheath, as in Grasses. It may be stipulate or exstiptilate (with or without slip, respectively); the shape, &c. of the slip, is described as if they were 1. and they may be free or adnate (F in fig., concrescent with the leaf- base or petiole, as in rose), united to other slip., inter- or intra-petiolar, branched, &c. (see Rubiaceae), ochreate (sheathing, as in Polygo- naceae), or modified in various ways. The venation (arrangement of the veins) may be pinnate or palmate ; in the former case there is a midrib with lat. veins branching from it, in the latter several equal veins spread out in the 1. like the ribs of a fan, from one point. The further ramification of the veins is descr. by net-veined (irreg. mesh- work, as in most Dicots.), parallel-veined (meshes more or less rect- angular, as in most Monocots.), fork-veined (veins forking into two, as in Ferns). L. are divided into simple and compound, as the stalk bears one or several separate leaflets. In the latter case the leaflet is desc. as if 368 LEAF it were a 1., and the common stalk is called the rachis. If the leaflets spring from the sides of the rachis, as in the pea, the 1. is pinnate (F), if all from one point palmate (E). If the leaflets of a pinnate 1., as in many Acacias, are again pinnately cpd., the 1. is bif innate. A 1. with 3 leaflets (as in clover) is ternate or tri -foliolate, with 3 ternate leaflets bitemate. Pinnate 1. may be equally (part-} pinnate (with an even number of leaflets), unequally (impart-} pinnate (with an odd leaflet at the end), or interruptedly pinnate (large and small leaflets alt. as in many Rosaceae). A palmate 1. with 5 or 7 leaflets is often called digitate. The leaflet may have stipule-like organs, or stipels (adjective stipellate). The leaf may be dorsiventral, isobilateral, or centric, or replaced by a phyllode, scale, pitcher, or other organ. The shape of the leaf-blade or leaflet itself, if simple, or the out- line of a cpd. 1. , may be needle-shaped or acicular as in Pinus, subulate or awl-shaped, tubular as in onion, linear (long and narrow as in Grasses), lanceolate (about 3 times as long as broad, tapering gradually towards the tip ; A in fig.), ovate (about twice as long as broad, and tapering towards the tip; B), cordate (similar, but heart-shaped at the base; C), elliptical (tapering equally to base and tip, and some- what narrow), oval (do. but wider), oblong (sides || for some distance, the ends tapering rapidly; F), reniform (kidney-shaped), orbicular (cir- cular in outline ; if the petiole is inserted at the middle of the blade, as in Tropaeolum, this leaf is termed peltate}, hastate (with two pointed jWl 1 ' u -Al • v a / AUM-C ;.ag e a. FORMS OF LEAVES. A, subsessile, exstip., lanceolate with cuneate base, entire, acute. B, sessile, exstip., ovate, serrate below, entire above, acum. C, petiolate, exstip., cordate, crenate, obtuse. D, ses- sile, exstip., somewhat obovate, pinnatifid sinuate, obtuse. E, petio- late, exstip., palmate (digitate), with obtuse oblanceolate leaflets. F, impari-pinnate with triangular adnate slips.; leaflets shortly stalked, oblong, apiculate, the laterals oblique. G, a, plicate 1. in section, b, mucronate apex, c, dentate spiny margin, d, retuse apex, e, glandu- lar hair. LEAF 369 lobes sticking out horiz. at the base), sagittate (two lobes projecting towards the stem), spatulate (spoon -shaped, as in daisy), &c. If a 1. be of lanceolate shape but the general tapering be towards the base, it is called oblanceolate (E) ; so also obovate or obcordate. If wedge- shaped, tapering to the base, it is cuneate. The 1. may be oblique or asymmetrical (F), when the midrib divides it into unequal halves, as in Begonia. The 1. (or leaflet) may be entire, i.e. without notches in the margin (A), or incised; the margin may also be fringed or fimbriate, cartilaginous, membranous, wavy or undulate (as in holly), curled or crisped (as in sea-kale), spiny (Gc), glandular (with sticky hairs or glands ; G), apiculate (do. with small point; F), truncate or praemorse( broad, straight end, as if bitten oK),cirrhose (tendrilled). The surface of the 1., as of other parts, may be glabrous (without hairs), pilose (soft, scattered hairs), downy or pubescent (fine, soft hairs), hairy (coarser), hispid (rough, bristly), tomentose (with a cottony felt), woolly, glandular-hairy; scabrous (rough), smooth, prickly, glaucous (with bluish waxy gloss), reticulate (netted), rugose (ridged or wrinkled), squarrose (roughly scurfy), &c. Hairs may be spreading m a//n?.sW(nattened down), jz;////V(unbranched), glandular (Ge), bifid, stellate (like a starfish), squamate (scaly), &c.; prickles may be straight or curved, bent backwards (retrorse), &c. The 1. may be doited with oil-glands, or variegated in colour, green, red, &c. ; frequent shades are fulvous (tawny), rubiginose (rust-coloured), &c. The texture maybe thin or herbaceous, coriaceous (leathery), siuculent or fleshy, membranous or scarious (thin, dry, not green, and flexible or stiff respectively), &c. L. may be evergreen or deciduous (falling in winter) ; in this case articulate (cut off by special absciss layer and falling early), or non- articulate (hanging on to the stem, though dead, for an indefinite period, as in beech hedges, oak, &c.). The vernation, or folding in bud, is descr. like the aestivation of flower-buds (y.v.) ; the following terms are also used to describe the folding of the individual 1., con- duplicate (folded lengthwise as in oak), plicate (folded several times ; W. 24 37° LEAF Ga), involute (margins rolled inwards), revolitte (outwards), redinate (apex bent down to base), com'ohde (rolled spirally, as in Musa) ; conduplicate 1. overlapping younger ones thus < < • > , as in Iris, are called equitant. When a character is not accurately described by one of the terms given, but is rather half-way between two, both are used; thus a 1. may be linear-lanceolate or ovate-cordate. Sub-, meaning 'nearly,' is often used as a prefix to adjectives, e.g. subsessile, subacute. As an example of the use of these technical terms, we quote from Lindley the descriptions of two leaves : Lilac: leaves opposite, exstipulate, roundish-cordate, very acute, thin, smooth, rather longer than the linear channelled petiole. Garden Strawberry : leaves all radical, ternate, dark-green, some- what shining, very coarsely serrated; with strong parallel oblique veins, silky beneath ; leaflets nearly sessile, roundish oblong, entire towards the base, shorter than the semi-cylindrical hairy petioles; stipules membranous, lanceolate, acuminate, half adnate. For other details of leaf structure and modification, see Anisophylly, Bud, Climbing Plants, Concrescence, Dimorphism, Driptip, Insecti- vorous Plants, Parasites, Phyllodes, Phyllotaxy, Saprophytes, Stipule, Storage, Vegetative Reproduction, Water-plants, Xerophytes, and articles below. Leaf, climbing, cf. Climbing Plants ; - colour, Amherstia, Brownea, Castilleja, Cinnamoimim, Haematoxylon, Saraca; -cushion, Cacta- ceae; -cutting ants, Acacia; -fall in dry season, Bombax, Caatingas, Cochlospernmm ; - mosaic, the fitting in of 1. with one another to make the best use of light and air available, Aesat/us, Hedera, Tilia- ceae ; - movements, cf. Movement. Leaudra Raddi. Melastomaceae (i). 200 trop. Am., W.I. Leather-leaf (Am.), Cassandra; -wood, Cyril la. Leavenworthia Torr. Cruciferae (2). 4 Atl. N. Am. Lebeckia Thumb. Leguminosae (in. 3). 258. Afr. Letaetanthus Endl. Epacridaceae (i). i Fuegia, Patagonia. Lebidiera Baill. =Cleistanthus Hook. f. (Euph.). Lebidieropsis Muell.-Arg. (Cleistanthus p.p. BH.). Euphorbiaceae (A. i. 2). i Indomal. Lecananthus Jack. Rubiaceae (i. 7). 3 Malaya. Lecaniodiscus Planch, ex Benth. Sapindaceae (i). 2 trop. Afr. Lecanopteris Reinw. Polypodiaceae. 7 Malaya. Lecanorchis Blume. Orchidaceae (n. 2). 4 Malaya, Japan. Lecanosperma Rusby. Rubiaceae (i. 5). i Bolivia. Lecanthus Wedd. Urticaceae (2). i Indomal., Abyssinia. Lechea Kalm. ex L. Cistaceae. 4 N. Am., W. I. Lechlera Miq. Inc. sed. Nomen. Lechuguilla, Agave. Leciscium Gaertn. Myrtaceae (inc. sed.). Nomen. Lecocarpus Decne. Compositae (5). i Galapagos. Lecokia DC. Umbelliferae (in. 4). i Crete to Persia. Lecomtea Pierre ex Van Tiegh. (Harmandia p.p.)- Olac. i trop. Afr. Lecomte'doxa Dubard (Mimusops p.p.). Sapotaceae (n). i Gaboon. Lecontea A. Rich. (Paederia p.p. EP.}. Rubiaceae (n. 6). 2 Madag. LEG UMINOSAE 3 7 1 Lecostemon (Mo£. et Sesse ex) DC. Rosaceae (vi). 6 Brazil to Mexico. Lectandra J. J. Smith. Orchidaceae (n. a. in), i Java. Lecticula Barnhart ( Utricularia p. p. ). Lontibulariaceae. i E. U.S. LecytMdaceae (EP.; Myrtaceae p.p. BH.} Dicots. (Archichl. Myrti- florae). 1 8 gen. 140 sp. trop. trees ; 1. generally in bunches at the ends of the twigs, simple, exstip. Fls. single or in racemose infls. , $, perig. or epig., with complete fusion of recept. and ovary. K4 — 6, C 4 — 6, A oo in several whorls, the sta. more or less united at base. G with 2 — 6 or more loc., in each i — oo anatr. ov. There is usually an intra-staminal disc as well as one under petals and sta. Berry or capsule. No endosp. Bertholletia and others are economic plants. Chief genera: Barringtonia, Napoleona, Couroupita, Lecythis, Ber- tholletia. Lecythis Loefl. Lecythidaceae. 30 trop. S. Am. Fr. a huge wooden caps., opening by a lid. The oily seeds are eaten as Sapucaia nuts. The empty fr. is termed a 'monkey-pot'; it is filled with sugar, the monkey inserts its hand, clasps it, and then cannot withdraw it. Lecythopsis Schrank (Conratari EP.). Lecythidaceae. 2 Brazil. Leda C. B. Clarke (Leptostachya p.p.). Acanthaceae (iv. B). 5 Malay Penins. Ledebouriella H. Wolff (Trinia p.p.). Umbelliferae (ui. 5). 2 As. Ledenbergia Klotzsch (Flueckigeria O. Ktze.). Phytolaccaceae. i trop. S. Am., W. I. Ledermannia Mildbr. et Burret. Tiliaceae. i trop. Afr. Ledermannlella Engl. Podostemaceae. r Cameroons. Ledocarpon Desf. — Balbisia Cav. (Geran. ). Ledonia Spach = Cistus Tourn. p.p. (Cist.). Ledothamnus Meissn. Ericaceae (i. 3). i Guiana. Lecluni Rupp. ex L. Ericaceae (i. i). 3 sp., two in N. Am., Z. palustre L. circumpolar. The leaves are rolled back (cf. Empetrum). Seeds winged at ends. This sp. is used in Labrador as tea. Leea Royen ex L. Vitaceae. 45 palaeotrop. Leechee, Litchi chinensis Sonner. Leek, Allium Porniin L. Leersia Soland. ex Sw. Gramineae (6). 5 N. temp, and trop. \%. Marsh grasses similar to Oryza, and used as fodder in As. L. oryzoidesSvi. (Eur.) has cleistogamic fls. (Darwin, Forms of Fls., p. 335). Lefebvrea A. Rich. (Peucedanum p.p. EP.). Umbellif. (in. 6). 4 trop. Afr. Lefrovia Franch. Compositae (12). i Bolivia. Legendrea Webb, et Berth. Convolvulaceae (i). i Canaries. Legitimate pollination, Lythrum, Primula. Legnephora Miers=Pericampylus Miers (Menisperm.). Legume, a fr. of one cpl. , dehiscing on both sides, Legnminosae. Leguminosae. Dicotyledons (Archichl. Resales). The second largest fam. of flowering plants, with 500 gen. and 12,000 sp., cosmop. Mimosoideae and Caesalpinioideae are mostly trop., Papilionatae mostly temp, (abundant on steppes, &c.). The account here given is largely condensed from that of Taubert in Nat. Pfl. 24 — 2 372 LEGUMINOSAE Living in every soil and climate they show great variety in habit — trees, shrubs, herbs, water-plants, xerophytes, climbers, &c. The roots of most exhibit peculiar tubercles — metamorphosed lat. roots containing peculiar bacterial organisms (Rhizobium sp.). Plants pro- vided with these are able to take up much more atmospheric nitrogen. The plant appears actually to consume the 'bacteroids' which live in its cells, after they have stored up in themselves a considerable amount of nitrogenous material. Hence the value of the L. as a crop on poor soil, or as preceding wheat in the rotation of crops ; for instead of impoverishing the soil they enrich it, either by the nitrogen con- tained in their roots and liberated as these decay, or by that of the whole pi. if ploughed in as 'green manure.' Stem commonly erect; many climbers. Some, e.g. Vicia, climb by leaf-tendrils, some, e.g. Bauhinia, by stem-tendrils, some by hooks (modified in Caesalpinia, &c., emergences in Acacia, &c.), some by twining. Creeping stems, rooting at the nodes, also occur. Thorns, usu. modified branches (e.g. Gleditschia) or stipules (Acacia), are common. The stems of the erect trop. sp. often branch so that the branches run parallel and erect, and bear crowns of 1. at the top. The stems of many lianes are peculiarly shaped, often flat, or corrugated in various ways, owing to peculiar growth in thickness. L. usually alt., slip., and nearly always cpd. Many have very small 1., e.g. Ulex, or scaly 1. and flat stems, e.g. Carmichaelia. The stipules vary much in size, &c. (see Acacia, Lathyrus, Vicia). The 1. usu. perform sleep-movements at night, some moving upwards, some downwards, or in other ways, but finally usu. placing the leaflet edge- wise to the sky. In Mimosa and Neptunia the 1. are sensitive to a touch and at once assume the sleep-position, recovering after a time. In Desmodium gyrans the lat. leaflets execute continuous spontaneous movements as long as the temperature is high enough. Infl. apparently always racemose, but with variety; simple raceme very common, also panicle and spike. Dorsi ventral racemes, re- sembling the cymes of Boraginaceae, also occur (e.g. Dalbergia). The fls. are regular (and then frequently polygamous) or irreg. (and then usu. $ ) ; recept. usu. convex or flat, so that at most the fl. is slightly perig. K developed in ascending order, usu. 5-merous, the odd (oldest) sepal anterior ; the sepals ± united. C polypetalous, alt. with the K; aestivation valvate(Mimosoideae), ascending imbr. (Caesalpinioideae), Floral diagrams of Vicia Faba (Papilionatae) and Acacia latifolia (Mimosoideae), after Eichler (modified). LEGUMINOSAE 373 or descending (Papilionatae). In many cases it is -|- to a high degree, having a large petal posterior (vexillum or standard), two lateral (alae or wings), and two anterior ± joined to form a keel or carina. A typically of 10 sta., free or united into a tube; in the latter case the tenth sta. (the posterior one) often remains free, so as to leave a slit in the tube, only covered loosely by this sta. Many variations are found. In cases where a keel is present, the sta. are enclosed in it. G typically of one cpl. with its ventral side directly posterior; long style and terminal stigma. There are two rows of ovules (alt. with one another so as to stand in one vertical rank), anatr. or amphitr., ascending or pend. Fertilisation (Papilionatae). The keel encloses the essential organs, protecting them from rain, &c. and rendering the fl. complex. Honey is secreted by the inner sides of the sta. near their base, and accu- mulates in the stamen-tube round the base of the ovary. The tenth sta. is free of the tube, and at the base, on either side of it, are two openings leading to the honey. The honey is thus concealed and at some depth, so that a clever insect with a tongue of moderate length is required. All this points to the P. being bee-flowers (class H), as in fact is the case. Insects alight upon the wings and depress them by their weight, whilst they probe for honey under the standard. The wings are always joined to the keel, usually by a protuberance in the former fitting into a suitable hollow in the latter, so that the keel is thus depressed likewise. This causes the emergence of the essential organs, the stigma usu. coming first, so that a fair chance of cross- fert. exists. Self-pollination usu. occurs when the insect flies off, leaving the keel to return to its former position. "Four different types of structure may be distinguished (in Papi- lionatae) according to the manner in which the pollen is applied to the bee : (i) P. in which the sta. and stigma emerge from the carina and again return within it. They admit repeated visits; e.g. Tri- folium, Onobrychis. (i) P. whose essential organs are confined under tension and explode. In these only one insect's visit is effective ; e.g. Medicago, Genista, Ulex. (3) P. with a piston mechanism which squeezes the pollen in small quantities out of the apex of the carina, and not only permits but requires numerous insect visits; e.g. Lotus, Ononis, Lupinus. (4) P. with a brush of hairs upon the style which sweeps the pollen in small portions out of the apex of the carina. They for the most part require repeated insect visits ; e.g. Lathyrus, Vicia." (Miiller.) Cleistogamy is fairly common. In several cases the stigma in the unvisited fl. lies in the keel among the pollen, but it has been shown that it only becomes receptive (if young) when rubbed, so that autogamy does not necessarily occur. For the phenomenon of enantiostyly (right- and left-styled fls.) see Cassia. Some have fls. which after fertilisation bury themselves in the earth and there ripen their fruit; e.g. Arachis, Lathyrus, Trifolium, Vicia, Voandzeia, &c. Fr. typically a legume, or pod opening by both sutures. In some the pod is constricted between the seeds, forming a lomentum which breaks up into indeh. one-seeded portions. The pods frequently open explosively, the valves twisting up spirally, e.g. in Ulex, Cytisus sp., 374 LEG U MINOS A E &c. In Colutea, &c. they are inflated. Some are eaten by animals, but the seed-coats are hard enough to preserve the seeds from injury. Some have a coloured fleshy aril (Acacia sp., &c.). Still others have hooked pods, e.g. Medicago, Mimosa. The seed is exalb. ; usu. large store of reserves in the cot. Economically the L. are most important. The seeds of many sp. form important food-stuffs, e.g. of Arachis (see Edible Products for common names), Cajanus, Cicer, Dolichos, Glycine, Lathyrus, Lens, Lotus, Lupinus, Phaseolus, Pisum, Vicia, Voandzeia, &c. The pods of Cevatonia, Tamarindus, Phaseolus, Prosopis, &c. are also eaten. A great number are valuable as fodder, and known as artificial grasses, e.g. Trifolium, Medicago, Onobrychis, Lotus, Vicia, &c. Many trop. and subtrop. sp. yield valuable timber, e.g. Acacia, Albizzia, Dal- bergia, Gleditschia, Hymenaea, Melanoxylon, Pericopsis, Pterocarpus, Robinia, Sophora, &c. ; Crotalaria and others are sources of fibre; Acacia, Genista, Haematoxylon, Indigofera, &c. yield dyes; gums and resins are obtained from Acacia, Astragalus, Copaifera, Hyme- naea, &c. ; oil is expressed from the seeds of Arachis and Voandzeia ; kino is obtained from Pterocarpus, and so on. See Economic Products and genera. Classification and chief genera (after Taubert) : the L. are nearly related to Rosaceae (especially Chrysobalaneae) and Connaraceae. Warming erects L. into a separate order, and considers each of the three chief divisions as an independent fam. The primary division is; I. MIMOSOIDEAE. Fls. regular, corolla valvate. II. CAESALPINIOIDEAE. Zygomorphic ; corolla-aestiva- tion imbricate ascending. III. PAPILIONATAE. Zygomorphic papilionaceous ; corolla- aestivation imbricate descending. These are again subdivided : the following key enables one to ascertain to which of the subdivisions any Leguminous plant belongs. I. MIMOSOIDEAE. A. Calyx valvate. a. Sta. more than 10. j. Ingeae (sta. united): Inga, Calliandra, Pithecolobium, Al- bizzia. •2. Acacieae (sta. free) : Acacia (only gen.). l>. Sta. as many or twice as many as petals. 3. Enniiinoseae (anther glandless) : Mimosa. 4. Adenanthereae (anther in bud crowned by a gland ; endo- sperm) : Neptunia, Prosopis. 5. Piptadenieae (ditto, no endosperm) : Piptadenia, Entada B. Calyx imbricate. 6. Parkicae : Parkia. II. CAESALPINIOIDEAE. A. Calyx in bud quite undivided or tubular below. a. Leaves simple or one pair of leaflets. Sta. 10 or fewer. 4. Bauhinicae: Bauhinia, Cercis. b. Leaves once pinnate (exceptions). Sta. oo or rarely 9 — 13. LEGUM1NOSAE 375 9. Swartzieae (Tounateae)'. Swartzia. c. Leaves bipinnate, or once pinnate; sta. 5. 1. Dimorphandreae: Dimorphandra. B. Calyx in bud quite polysepalous or very nearly so. a. Two anterior petals modified to large glands, anthers opening by pores. 6. Kramerieae: Krameria (only gen.). b. Two anterior petals developed or not, but not glandular, a. Leaves some or all bipinnate. 7. Eucaesalpinieae: Caesalpinia, Haematoxylon, Gleditschia, Poinciana. ^. Leaves once pinnate. 5. Cassieae (anthers basifixed, opening by terminal pores) : Cassia, Ceratonia. 3. Amherstieae (dorsifixed, no pores; ovary adnate behind to torus) : Hymenaea, Tamarindus. 8. Sclerolobiea*: (ovary free, ovules > 3): Sclerolobium. 2. Cynometreae (ovules i or 2) : Copaiba. III. P API LION A TA E. A. Sta. free. 1. Sophoreae (pinnate leai) : Sophora, Myroxylon. 2. Podalyrieae (simple or palmate) : Anagyris. B. Sta. united. a. Lomentum. 7. Hedysareae: Coronilla, Onobrychis, Arachis, Desmodium. b. Legume or indehiscent pod. a. Leaves absent. 6. Galegeae: Robinia, Colutea, Indigofera, Tephrosia, Astra- galus, Oxytropis. (Cf. below.) p. Leaves present. I . Leaf or leaflets "without stipels. (1) Lf. simple or palmate. Lf. with 3 entire leaflets. 3. Genisteae: (shrubs) Genista, Lupinus, Ulex, Cytisus, Cro- talaria. 6. Galegeae: (herbs) as above. * Lf. with 5 entire leaflets. 5. Loleae: Anthyllis, Lotus. *** Lf. with 3 toothed leaflets. 4. Trifolieae : Ononis, Medicago, Trifolium. (2) Leaf pinnate. * Leaf-stalk ending in bristle or tendril. 9. Vicieae: Abrus, Vicia, Lathyrus, Pisum. ** Leaf-stalk not ending so. t Pod dehiscing in 2 valves. § Sta. filamentous. 10. Phaseoleae : (ovary surrounded by disc) Apios, Phaseolus. 6. Galegeae: (no disc) as before. §§ Some or all sta. broadened at apex. 5. Loteae : as before, ft Pod indehiscent. 376 LEGUMINOSAE 8. Dalbergieae: Dalbergia, Pterocarpus. II. Leaves or leaflets with stipels. 10. Phaseoleae: (pod dehiscent) as before. 8. Dalbergieae: (pod indehiscent) as before. Lehmanniella Gilg. Gentianaceae (i). 2 Colombia. Leianthus Griseb. (Lisianthus EP.}. Gentianaceae (i). 10 W.I. , C. Am. Leibergia Coulter et Rose. Umbelliferae (ill. 5). i Idaho. Leiboldla Schlecht. ( Vernonia p.p.). Compositae (i). 4 N. Am. Leichhardtia F. Muell. Menispermaceae. i N.E. Austr. Leichtlinia H. Ross (Agave p. p. EP.}. Amaryllidaceae (n). i Mexico. Leidesia Muell. -Arg. in DC. Euphorbiaceae ( A. n. 2). 2 Cape Colony. Leighia Cass. = Viguiera H.B. et K. (Comp.). LeimantMum Willd. =Melanthium L. (Lili.j. Leiocarpodicraea Engl. (Dicraea p.p.). Podostemaceae. 3 trop. Afr. LeiocMlus Hook. (Buseria EP. ) . Rubiaceae (n. 4). i Maclag. Leioclusia Baill. Guttiferae (inc. sed.). i Madag. Leiogyne K. Schum. {Neves- Arniondia EP.). Bignoniaceae (i). i E. Brazil. Leiopliaca Lindau. Acanthaceae (iv. A), i trop. Afr. Leiophyllum Hedw. f. Ericaceae (1.3). i Atl. U.S. Leioptyx Pierre ex De Wild. Meliaceae. i trop. Afr. Leiostemon Rafin. (Pentstenion p.p.). Scrophular. (n. 4). 2 N.W. N. Am. Leiothrix Ruhland (Paepalanthus p.p.). Eriocaulonaceae. 30 S. Am. Leiothylax Wmg. (Dicraea p.p.). Podostemaceae. 3 trop. Afr. Leiphaimos Cham, et Schlechtd. (Voyria Bff.). Gentianaceae (i). 20 trop. Am. and Afr., W.I. Leitgebia Eichl. Ochnaceae. i Brit. Guiana. Leitneria Chapm. Leitneriaceae. 2 N. Am. Shrubs with spikes of dioecious fls. 3 naked with 3 — 12 sta. ? with scaly P, i cpl. and long style. G i-loc., with i amphitr. ov. Fr. drupaceous. Seed albumi- nous. Embryo straight. Leitneriaceae (EP., BH.). Dicots. (Archichl. Leitneriales; Unisexuales BH.). Only genus Leitneria, q.v. Leitneriales. The yth order of Dicots. Archichlamydeae. Leitnerieae (BH. ) = Leitneriaceae. Lemairea de Vriese. Goodeniaceae (inc. sed.). i Amboina. Lemaireocereus Britton et Rose (Cereus p.p.). Cactaceae (in. i). 13 trop. and subtrop. Am. Lembertia Green (Eatonella p. p.). Compositae (6). i California. Lemmonia A. Gray. Hydrophyllaceae. i California. Lemna L. Lemnaceae. 6 cosmop. ; 4 in Brit, (duckweed). The plant consists in most, e.g. L. minor L., of a flat green floating blade, the stem, which performs leaf-functions. From the under side hangs down a long adv. root, with well-marked root-cap, visible to the eye. No 1. The stems are oval and slightly turned up at the ends, so that if two are placed near together in water, they will run against one another and adhere by the tips. In the post, portion on either side is a groove under the edge. In this arise branches which may either (as in L. trisulca L., &c.) remain in union with the parent shoot, or become detached and give rise to new plants. In autumn a number LEOCUS 377 of these are formed ready to start growth in the next spring, whilst the mother plants sink to the bottom. The fls. are also borne in these grooves. There is a very reduced spathe, with 2 i p.p.). Ranunculaceae (3). i N. Am. Leucocoryne Lindl. Liliaceae (iv). 8 Chili. Cult. orn. fl. Leucocrinum Nutt. ex A. Gray. Liliaceae (in), i W. U.S. Leucocroton Griseb. Euphorbiaceae (A. II. 2). 4 Cuba. Leucogenes Beauverd. Compositae (4). 2 New Zealand. Leucojum L. Amaryllidaceae (i). 10 S. Eur. (snow-flake). Leucolaena R. Br. = Xanthosia Rudge p.p. (Umbell.). 382 LEUCOLENA Leucolena Ridley. Orchidaceae (n. 5). i Malay Peninsula. Leucomeris D. Don. Compositae (12). 2 Nepal, Burma. Leucomphalos Benth. Leguminosae (ill. i). i W. trop. Afr. Leuconotis Jack. Apocynaceae (i. i). 10 Malaya. Leucopholis Gardn. Compositae (4). 3 S. Brazil. Leucophrys Rendle (Panicnin p.p.). Gramineae (5). i trop. Afr. Leucophyllum Humb. et Bonpl. Scrophulariaceae (i. i). 3 Texas, Mexico. Leucopitys Nieuwland (Finns p.p.). Coniferae (Pinaceae). 2 N. temp. Leucopogon R. Br. (Slyphdia Sol.). Epacridaceae (3). 130 Austr., Malaya. Leucopsis Baker (Aster p.p. EP.}. Compositae (3). 10 trop. Am. Leucorchis Blume (Duiymoplexis BH.). Orchidaceae (n. 2). 3 In- domal. Leucosalpa Scott Elliot. Scrophulariaceae (in. 2). i Madag. Leucosceptrum Sin. Labiatae (i). 2 Himalaya, China. Leucosidea Eckl. et Zeyh. Rosaceae (in. 5). i S. Afr. Leucosmia Benth. Thymelaeaceae. 4 Fiji. Leucospermum R. Br. Proteaceae (i). 30 S. Afr. Leucosphaera Gilg. Amarantaceae (2). 2 S.W. Afr. Leucostegane Prain. Leguminosae (n. 3). i Indomal. Leucostegia Presl — Davallia Sm. (Polypod.). Leucosyke Zoll. et Morr. Urticaceae (3). 10 Malaya, Polynesia. Leucosyris Greene (Aster p.p.). Compositae (3). 2 N. Am. Leucothoe D. Don. Ericaceae (n. i). 35 Am. Like Andromeda. Leunisia Phil. Compositae (12). i Chili. Leurocline Sp. Moore. Boraginaceae (iv. 5). 2 trop. Afr. Leuzea DC. (Centaurea p.p. EP.). Compositae (n). 3 Medit. Levenhookia R. Br. Stylidiaceae. 7 Austr. The labellum is shoe- shaped and at first embraces the column, but if touched it springs downwards. Lever- wood, Ostrya. Levieria Becc. Monimiaceae. 4 New Guinea, Queensland. Levigatus (Lat. ), smooth, slippery. Levisticum Riv. ex L. Umbelliferae (ill. 6). 2 Eur., W. As. Levya Bur. ex Baill. Bignoniaceae (i). i Nicaragua. Lewisia Pursh. Portulacaceae. 2 California. L. rediviva Pursh (bitter- root) with thick rhiz., fleshy roots and 1., is very xero. Two years' drying will hardly kill it. K.4 — 8, C 8— 16; A oo . Leycesteria Wall. Caprifoliaceae. 3 Himalaya, China. Leyssera L. Compositae (4). 5 S. Afr,, Medit. Lhotzkya Schau. Myrtaceae (n. 2). 10 Austr. Liabum Adans. Compositae (8). 60 Am. and W. Ind. Liane, a woody climber, Ancislrocladus, Apocynaceae, Aristolockiaceae, Asclepiadaceae, Bignoniaceae, Hippocrateaceae, Landolphia, Legumi- nosae, Malpighiaceae, &c. Cf. Climbing Plants. Liatris Schreb. Compositae (2). 15 N. Am. Libanotis Riv. ex Hall. =Seseli L. (ytf^.J^Athamanta L. Liber, phloem. Liberian coffee, Coffea liberica Hiern. LI LI ACE AE 383 Libertia Spreng. Iridaceae (n). 8 Chili, Austr., N.Z. Cult. orn. fl. Libocedrus Endl. Coniferae (Pinac. 37; see C. for gen. char.). 8, 2 in Chili, i in N.Z., i each in New Caled., Japan, China, Calif. L. Doniana Endl. (N. Z.), L. tclragona Endl. (Chili) and L. decur- rens Torr. (Calif., white cedar) yield valuable timber. Licania Aubl. Rosaceae (vi). 45 S. Am. Lichtensteinia Cham, et Schlechtd. Umbelliferae (in. 5). 5 S. Afr. Licuala Thunb. Palmae (i. 2). 40 Indomal. Lidbeckia Berg. Compositae (7). 3 S.W. Cape Colony. Liebrechtsia Wildem. (I'igna p.p. EP.}. Legumin. (in. 10). 6 trop. Afr. Lietzia Regel. Gesneriaceae (n). i Brazil. Lifago Schweinf. et Muschler.. Compositae (4). i Algeria. Life-history, the course of development and metamorphosis. Ligea Poit. ex Tul. Podostemaceae. 5 Brazil, Guiana. Ligeria Decne. = Sinningia Nees p.p. (Gesn.). Light, effects of, see Physiological textbooks. Lightfootia L'Herit. Campanulaceae (i). 45 Afr. Lightia Schomb. Trigoniaceae. 2 Guiana, Amazon valley. Lign-aloes, eagle-wood, Aquilaiia Agallocha Roxb. (Burma). Ligneus (Lat.), woody; -ification, becoming woody. Lignum vitae, Gnaiacuin officinale L. Ligularia Cass. (Scnecio p.p. BH.). Compositae (8). 35 Eur., As. Ligulate (C), strap-shaped, Compositae. Ligule, a scale at upper end of leaf-sheath, Gramineae. Liguliflorate, with ligulate flowers. Ligusticella Coulter et Rose (Ligitstifiim p.p.). Umbelliferae (in. 5). i Colorado. Ligusticum L. Umbelliferae (in. 5). 55 N. Hemisph., Chili, N.Z. L. scoticuin L. (lovage) in Brit.; it is sometimes used as a pot-herb. Ligustrum (Tourn.) L. Oleaceae. 35, esp. E. As. L. vulgare L. (piivet) in Brit. Lijndenia Zoll. et Morr. (Memecylon p.p. BH). Melastom. (in), i Java. Lilac, Syringa vulgaris L. ; Californian -, Ceanothus ; Indian -, Melia Lagers t roe mia; West Indian -, Melia. Lilaea Humb. et Bonpl. Scheuchzeriaceae (Naiad. BH.}. i Rockies, Andes. Liliaceae (EP., BH.). Monocots. (Liliiflorae; Coronarieae BH.}. One of the largest fams. of flowering plants; 200 gen., 2700 sp., cosmop. ; the smaller groups often confined to definite floral regions. Most are herbs with sympodial rhiz. or bulbs; a few trop. and warm temp, forms, e.g. Yucca, Dracaena, &c., are shrubs or trees, often with an unusual mode of growth in thickness. Many are xero.; some, e.g. Aloe and Gasteria, are succulent ; others, e.g. Phormium, have hard isobil. 1. ; others, e.g. Dasylirion, have tuberous stems and narrow 1. ; Bowiea only produces leafy shoots in the wet season. Smilax, Gloriosa, &c., are climbing pi., the former with peculiar stipular tendrils. Ruscus exhibits phylloclades. Infl. most commonly racemose ; fls. ,vith no bracteoles ; when the latter occur, the further branching from their axils usually takes a 384 LILIACEAE Diagram of Convallaria (after Eichler). cymose form, especially that of a bostryx, as in Hemerocallis. The apparent umbels or heads of Allium, Agapanthus, &c. are really cymose. Sol. term. fls. occur in tulip, &c. Fls. usu. § , reg., penta- cyclic, 3-merous (rarely 2, 4, or 5), hypog. P 3 + 3, free or united, petaloid or sometimes sepaloid; A 3 + 3 or fewer, rarely more, usu- ally with introrse anthers; G (3) usu. sup., rarely inf. or semi-inf., 3-loc. with axile, or rarely i-loc. with parietal plac.; ovules usu. oo , in two rows in each loc., anatr. Fr. usu. capsular, loculic. or septic., sometimes a berry. Seed with straight or curved embryo, in abundant fleshy or cartilaginous, never floury, endosp. Fls. usu. insect-pollinated. Honey in Scilla, Allium, &c., is secreted by glands in the ovary- wall between the cpls.; in other cases by glands on the bases of the perianth-1. (see Muller's Pert, of Fls. ). Yucca (q. v. ) has a unique pollination-method. Economically the L. are of no great value. The chief food plants are Alli- um and Asparagus ; Phormium, Yucca, and Sansevieria yield useful fibre ; Smilax, Urginea, Aloe, Colchicum, Veratrum, &c., are medicinal. Xanthorrhoea and Dracaena yield resins ; Chlorogalum is used as soap. Many are favourite garden and greenhouse plants, e.g. Convallaria, Tulipa, Fritillaria, Lilium, Agapanthus, Kniphofia, Funkia, Hyacinthus, Gloriosa, and many more. Classification and chief genera (after Engler): the L. are closely allied to Juncaceae; usu. they can be distinguished by their petaloid P, but many L. have a sepaloid P, e.g. Xanthorrhoea, Kingia, &c., and in these cases almost the only distinction is the absence in L. of the long thread-like twisted stigmas of J. BH. unite these genera, and some others, to Juncaceae, and place sub-fams. vin and ix in Haemodoraceae. Warming splits up the family into Colchicaceae, Liliaceae, and Convallariaceae. I. MELANTHI01DEAE (rhiz., or bulb covered with scale-1. and with term, infl.; anthers extr. or intr.; caps, loculic. or septic.; fr. never a berry): Tofieldia, Narthecium, Veratrum, Gloriosa, Col- chicum. II. HRRRERIOIDEAE (tuber, with climbing stem; 1. in tufts ; small-flowered racemes at base of these or in panicles at ends of twigs; septic, caps.) : Herreria (only genus). III. ASPHODELO1DEAE (rhiz. with radical 1., rarely stem with crown of 1. or leafy branched stem or bulb; infl. usu. term., a simple or cpd. raceme or spike; P or (P); anthers intr.; caps., rarely berry): Asphodelus, Chlorogalum, Bowiea, Funkia, Hemerocallis, Phormium, Kniphofia, Aloe, Gasteria, Haworthia, Aphyllanthes, Lomandra, Xanthorrhoea, Kingia. LIME 385 IV. ALLIOIDEAE (bulb or short rhiz.; cymose umbel ± enclosed by two broad or rarely narrow 1., sometimes joined; infl. rarely of i fl.): Agapanthus, Gagea, Allium, Brodiaea. V. LILIOIDEAE (bulb; infl. term., racemose; P or (P) ; anthers intr. ; caps, loculic., except in Calochortus): Lilium, Fritillaria, Tulipa, Scilla, Ornithogalum, Hyacinthus, Muscari. VI. DRACAEN01DEAE (stem erect with leafy crown, except in Astelia; 1. sometimes leathery, never fleshy; P free or united at base; anthers intr.; berry or caps.) : Yucca, Dasylirion, Dracaena. VII. ASPARAGOIDEAE (rhiz. subterranean, sympodial ; berry) : Asparagus, Ruscus, Polygonatum, Convallaria, Trillium. VIII. OPHIOPOGONOIDEAE (short rhiz., sometimes with suckers, with narrow or lanceolate radical 1.; P or (P) ; anthers intr. or semi-intr.; ovary sup. or |-inf.; fr. with thin pericarp and i — 3 seeds with fleshy coats): Sansevieria, Ophiopogon. IX. ALETR01DEAE (short rhiz. with narrow or lanceolate radical 1.; (P); anthers semi-intr.; caps, loculic.; seeds , with thin testa): Aletris (only genus). X. LUZURIAGOIDEAE (shrubs or undershrubs with erect or climbing twigs; infl. -twigs usu. many-flowered, cymose, rarely i- flowered, with scaly bract at base; both whorls of P alike or not; berry with spherical seeds): Luzuriaga, Lapageria. XI. SMI LA COIDEAE (climbing shrubs with net-veined 1. ; fls. small in axillary umbels or racemes or term, panicles; loc. with i or 2 orthotr. or semi-anatr. ovules): Smilax. Liliago (Caesalp.) L. = Anthericum L. (Lili.). Liliiflorae. The c;th order of Monocotyledons. Lilium Tourn. ex Linn. Liliaceae (v). 50 N. temp. Herbs with scaly bulbs, leafy stems and fls. in racemes. Honey secreted in long grooves at the bases of the P-leaves. The fls. of many sp. are visited by Lepidoptera. L. Mart agon L. gives off its scent at night (cf. Oenothera). L. bitlbiferuin L. is reproduced veg. by bulbils in the leaf-axils. In most sp. with hanging fls. the caps, when ripe stands upwards, so that the seeds can only escape when it is shaken. Many sp. of lily cult. orn. fl. Lilloa Speg. (Synandrospactix EP.}. Araceae (vn). i Argentina. Lily, Lilium, used of most Liliaceae, Amaryllidaceae, &c. ; arum -, Richardia africana Kunth ; Bourbon -, Lilium candidiim L. ; Chinese sacred-, Narcissus; Jacobean-, Sprekeliaformosissima Herb. ; Kaffir -, Clivia ; - of the valley, Convallaria majalis L. ; queen -, Phaedranafsa; spire-, Galtonia; swamp-, Zephyranthes; -thorn (W.I.), Catesbaea; water-, Nuphar, Nymphaea. Lima bean, Phaseolus lunatits L. Limacia Lour. Menispermaceae. 20 trop. As. and Afr. Limaciopsis Engl. Menispermaceae. i trop. Afr. Limatodes Lindl. (Phajtis p.p. BH.}. Orchidaceae (n. Q). i Burma. Crossed by Preptanthe veslita is the Calanthe Veitchii Hort. Limb, the free portion of a concrescent corolla; one branch of a sym- poclium. Lime, Citrus Medica L. , var. acida, var. Limetta, Tilia ; - myrtle (W.I.), Triphasia. W. 25 3 86 LIMEUM Limeum L. Phytolaccaceae (Aizoa eae BH.}. 15 S- Afr. to India. Limit of trees, either N. or S., or upwards in the Mts., beyond which they do not naturally occur. Limivasculum B6rner=Carex p.p. (Cyper.). Limnanthaceae (EP.; Geraniaceae p.p. BH.). Dicot. (Archichl. Sapindales). A very small fam. (2 gen., 5 sp., N. Am.) sometimes united to Geraniaceae, but with the ovules as in ord. Sapindales. Herbs with exstip. alt. 1. and reg. 5 fls., 3— 5-merous, with two whorls of sta. Ovary 3—5-100., ovules i in each loc., ascending, the micropyle facing outwards and downwards. Fruit a schizocarp. Seeds exalbuminous. Genera: Limnanthes, Floerkea. Limnanthemum S. P. Gmel. Gentianaceae (n). 20 trop. and temp. L. (Villarsia) nvmphaeoidcs Hoffmgg. et Link, S. England, is a water- plant with habit of Nymphaea. The infl. appears to spring from the top of the leaf-stalk, but really the floating 1. springs from the infl. axis. This is an advance upon the Nymphaea construction, as the materials going from 1. to seeds have not to travel to the bottom of the pond and up again. Limnanthes R. Br. Limnanthaceae. 4 Pacific N. Am. Limnas Trin. Gramineae (8) i E. Siberia. Limnia L. (Claytonia Gronov.). Portulacaceae. 10 N. Am. Limnium (Cl.)i a lake formation. Limnobium Rich. (Hydromystria G. F. W. Mey., Trianea Karst.). Hydrocharitaceae. 3 Am. L. (H.) stoloniferum Griseb. (T. bogo- tmsis Karst.), a small floating plant often cult. It reproduces veg. by ' runners' (cf. Hydrocharis). Its root-hairs are used to show circula- tion of protoplasm. Only the ? pi. is known in Eur. Limnocharis Humb. et Bonpl. Butomaceae. 2 trop. S. Am., W.I. Limnochloa Beauv. ex Lestib. = Eleocharis R. Br. (Cyper.). Limnodium (Cl. ), a salt marsh formation. Limnophila R. Br. (Ambulia Lam. EP.}. Scrophular. (H. 6). 30 palaeotrop. Limnophyton Miq. Alismaceae. 2 palaeotrop. Limnbrchis Rydberg (Habenaria p.p.). Orchid, (n. i). 24 N. Am., Eur. Limnosipanea Hook. f. Rubiaceae (i. 3). 48. and C. Am. Limodorum (Tourn.) L. Orchidaceae (n. 3), i S. Eur. A leafless saprophyte with no chlorophyll (cf. Epipogum). The 4 lat. sta. are sometimes fertile. Limonia L. Rutaceae (v). 10 trop. Afr. and As. Some have thorns in the leaf-axils (1. of branch, as in Cactaceae). The fr. of L. acidis- sinia L. is used in Japan as a substitute for soap. Limoniastrum Moench. Plumbaginaceae. 3 Medit. Limonium Tourn. ex Mill. =Statice L. (Plumbag.). Limosella L. Scrophulariaceae (n. 6). 7 cosmop. L. aquatica L. , mud- wort, in Brit, multiplies by runners. Linaceae (hJ\ ; BH. add Erythroxylaceae}. Dicot. (Archichl. Gera- niales). 9 gen., 150 sp., cosmop. Mo.-,t are herbs and shrubs with all. entire often stip. 1. Infl. cymose, a dichasium or cincinnus, the latter usu. straightening out very much and looking like a raceme. Fl. ?, reg., usu. p-merous. K 5 , quincuncial ; €5, imbr. or conv. ; LINNAEA 387 AS, 10 or more, often with stds., united at base into a ring; G (2— 3 — 5), multi-loc. , often with extra partitions projecting from the mid- ribs of the cpls., but not united to the axile plac. ; ovules i or 2 per loc., pend. , anatr., with the micropyle facing outwards and upwards. Septic, caps., or drupe. Embryo usu. straight, in fleshy endosp. Linum (flax, linseed) is economically important. Chief genera: Radiola, Linum, Hugonia. [BH. chars, incl. ov. i-loc.] Linaloa, Bursera delpechiana Poiss. (Mexico). Linanthus Benth. (Gilia p.p.). Polemoniaceae. 15 N. Am. Linaria Tourn. ex Mill. (BH. incl. Cymbalaria Medic., Elatine Rupp., Elatiuoides Wettst.). Scrophulariaceae (11. 3). 100 *_ and S. Am., chiefly extra-trop. 7 in Brit, (toad-flax), esp. L. Bulgaria Mill, (yellow toad-flax). The pi. is a perennial, each year's growth arising from an adv. bud upon the summit of the root. The fl. is closed at the mouth ; honey is secreted by the nectary at the base of the ovary and collects in the spur. The only visitors are the larger bees, which are able to open the fl. , and whose tongues are long enough to reach the honey. Peloria of the fl. is frequent ; a term. fl. appears upon the raceme and is symmetrical, with 5 spurs upon the C and a tubular mouth. Sometimes fls. of this type occur all down the raceme. Another interesting sp. is L. Cymbalaria Mill., the ivy-leaved toad- flax, found on walls in Brit. Before fert. the fls. are positively helio- tropic and stand erect ; after it they become negatively heliotropic and bend downwards, seeking out the dark crannies in the substratum, where the seeds ripen. Linariopsis Welw. Pedaliaceae. i trop. S.W. Afr. Linconia L. Bruniaceae. 3 S. Afr. Lindackeria Presl = Oncoba Forsk. (Flacourt.). Lindauea Rendle. Acanthaceae (iv. B). i Somaliland. Lindelofla Lehm. Boraginaceae (iv. i). 2 W. Himal. Linden, Tilia. Lindenbergia Lehm. Scrophulariaceae (n. 6). 12 trop. As. and Afr. Lindenia Benth. Rubiaceae (i. 3). 3 C. Am., Fiji, New Caledonia. Lindera Thunb. Lauraceae (n). 70 sp. Japan to Java. L. Benzoin Meissn. has aromatic bark (antifebrile). Lindernia All. (Yandellia L. BH.}. Scrophular. (n. 6). 30 warm. Lindheimera A. Gray et Engelm. Compositae (5). i Texas. Lindleya H. B. et K. Rosaceae (i. 2). i Mexico. Lindleyella Rydberg (Lindleya H. B. et K. ). Rosaceae (i. 2). 2 N. Am. Lindleyella Schlechter. Orchidaceae (a. 12). i W.I., Guiana. Lindmania Mez. (Cottendorfia p.p.). Bromeliaceae (2). 5 S. Am. Lindsaya Dryand. Polypodiaceae. 90 trop. and subtrop., exc. Afr. Lineae (BH. ) = Linaceae. Linear (1.), long and narrow, as in grasses. Lineate (dimin. lineolate), marked with (fine) lines. Ling, Callitna vulgaris Salisb. Lingelsheimia Pax. Euphorbiaceae (A. I. i). 2 C. Afr. Linnaea Gronov. in L. (incl. Abelia R. Br.). Caprifoliaceae (in). 20 *, L. borealis L. in Scotland. Ovary covered with glandular hairs. Sta. 4, didynamous. Two loculi are oo -ovulate and sterih, the other i -ovulate and fertile. Some (Abelia) cult. orn. fl. ; ed. fr. 25—2 383 L1NNAEOPSIS Linnaeopsis Engl. Gesneriaceae (i). i trop. Afr. Linnaeus' system, the artificial system, by which plants were divided into classes, &c. on purely artificial lines, by the number of stamens and cpls., &c. ; -species, cf. Nomenclature. Linocalyx Lindau. Acanthaceae (iv. B). i trop. Afr. Linochilus Benth. = Diplostephium H. B. et K. (Comp.). Linociera Sw. (Mayepa Aubl. EP.). Oleaceae. 50 trop., subtrop. Linodendron Griseb. (Lasiadenia BH}. Thymelaeaceae. 4 Cuba. Linospadix Becc. Palmaceae (iv. i). 8 New Guinea, Austr. Linostoma Wall. Thymelaeaceae. 2 Further India. Linosyris Cass.= Aster Tourn. p.p. ; do. Torr. et Gray=Bigelovia DC. Linseed, Lint, Linum ttsitatissimum L. Lintonia Stapf. Gramineae (9). i Brit. E. Afr. Linum Tourn. ex L. Linaceae. 95 temp, and subtrop., esp. Medit. 4 in Brit., L. catharticum L. (purging flax), common, L. usitatissimuin L. (common flax or linseed), an introduction, cult, in Ireland and elsewhere. Fls. in sympodial cincinni. Several are heterostyled (dimorphic), e.g. the common red one of gardens, L. grandiflonun Desf. Illegitimate pollination in this sp. produces absolutely no seed at all. The seed has a mucilaginous testa which swells on wetting. Flax is the fibre of Linum usitatissimnm L., obtained by rotting off the softer tissues in water; linen is made from it. The shorter fibres form tow, and scraped linen lint. The seeds (linseed) yield an oil by pressure, and the remaining ' cake ' (cf. Gossypium) is used for cattle-feeding, &c. Lip, a projecting anterior part of an irregular corolla or calyx. Liparia L. Leguminosae (in. 3). 48. Afr. Liparis Rich. (BH. incl. Cestichis Thou.). Orchidaceae (li. 4). 100 trop. and temp, (i in Brit., rare). Liparophyllum Hook. f. Gentianaceae (n). i Tasm., New Zealand. Lip-fern (Am.), Cheilanthes. Lipocarpna R. Br. in Tuckey. Cyperaceae (i). 7 trop. Lipochaeta DC. Compositae (5). 12 Hawaiian and Galapagos Is. Lipostoma D. Don (Coccocypselum BH.}. Rubiaceae (i. 2). 2 Brazil. Lipozygis E. Mey. -— Lotononis Eckl. et Zeyh. p.p. (Legum.). Lippia Houst. ex L. Verbenaceae (i). 120 trop. Am., Afr. The 1. of L. citriodora H. B. et K. yield an aromatic oil used in perfumery under the name Verbena-oil. Some have axillary thorns. Liquidambar L. Hamamelidaceae. 4 Medit., As., N. Am. Fls. monoec., apet., the cf in upright spikes, the ? in heads on pend. stalks. The seeds are easily shaken out in strong winds. Storax (a fragrant balsam) is obtained from all, but chiefly from L. orientalis Mill. (As. min.). L. styraciflua L., sweet gum, N. Am.; wood useful (satin walnut). Liquorice, Glycyrrhiza glabra L. ; Indian-, -vine (W.I.), Abrus. Lirayea Pierre (Afromendoncia EP.}. Acanthaceae (n). i trop. Afr. Liriodendron L. Magnoliaceae. i N. Am., L. tulipifera L., the tulip-tree, often cult, in parks. The 1. is polymorphic. Fr. a samara ; the aggregate of samaras upon the recept. looks like a pine-cone. The wood is useful (canary whitewood). Liriope Lour. Liliaceae (vin) (Haemodor. BH.}. 2 E. As. LITERATURE 389 Liriosma Poepp. et Encll. Olacaceae. 15 trop. S. Am. Lisaea Hoiss. (Caucalis p.p. BH.). Umbelliferae (m. 2). 3 W. As. Lisianthus L. Gentianaceae (i). 15 W.I. , Cent. Am. Lissanthe R. Br. (Styphelia p.p. EP.). Epacridaceae (3). 4 Austr. Lissocarpa Benth. Styracaceae. i Brazil. Lissochilus R. Br. Orchidaceae (n. 10). 50 trop. Afr. Listera R. Br. Orchidaceae (n. 2). 20 N. temp. L. ovata R. Br. (tway-blade) and L. cordata R. Br. in Brit. The labellum is bent downwards and forked into two. The rostellum on being touched ruptures violently and ejects a viscid fluid which cements the pollinia to the insect as in Epipactis (see Darwin, Orchids, p. 115). Listia E. Mey. Leguminosae (in. 3). i S. Afr. Listrostacbys Reichb. f. (Angraeatm p.p. BH.). Orchidaceae (ll. 20). 50 trop. Afr. Litanthus Harv. Liliaceae (v). i S. Afr. Litanuni Nieuwland (Talinum p.p.). Portulacaceae. i N. Am. Litchi Sonner. (Nephelium p.p. BH.). Sapindaceae (i). i China, L. chinensis Sonner. (litchi or leechee), cult, for ed. fr., a one-seeded nut with fleshy aril. Literature : the following general works of reference may be mentioned, as providing further information upon the various subjects of this dictionary : Admiralty Manual of Scientific Enquiry. Bailey, L. H., Universal Cyclopaedia of Horticulture. Baker, J. G., Handbook of the Fern Allies. Bentham and Hooker, Genera Plantarum. London. British Pharmacopoeia. Christensen, Index Filicum. Copenhagen, 1906. Clements, Research Methods in Ecology. Lincoln (Nebr.), 1905. Darwin, Origin of Species, Naturalist's Voyage, &c. De Candolle, Origin of Cultivated Plants, Engl. ed. De Dalla Torre and Harms, Genera Siphonogamarum, De Vries, The Mutation Theory, Engl. ed. Durand, Index Generum Phanerogamarum. Eichler, Bluthendiagramme. Leipzig. Engler, Das Pflanzenreich. Leipzig (in course of publication) ; Syllabus der Vorlesungen. Engler and Prantl, Die Natiirlichen Pfianzcnfamilien. Leipzig. Goebel, Organography of Plants, Engl. ed. Oxford. Hooker, W. J., Synopsis Filicum. Hooker and Jackson, Index Kewensis, and Supplements. Jackson, Dictionary of Botanical Terms. Kew : Official Guides to the Museums. Knuth, Handbook of Floral Pollination, Engl. ed. Oxford. Linnaeus, Genera Plantarum. Sanders, Encyclopaedia of Gardening. London, 1912. Schimper, Geography of Plants, Engl. ed. Oxford, 1903. Semler, Die tropische Agrikultur. Wismar, 1897 — The Treasury of Botany. Wallace, Travels on the Amazon, Island Life, Malay Archipelago, &c. 390 LITERATURE Ward, Trees. Cambridge. \\3.\mmg,Oecology of Plants, \Lng\. ed. Oxford; Systematic Botany. Watt, Dictionary of the Economic Products of India; Commercial Products of India, London, 1908. Wiesner, Die Rohstoffe des Pflaiizenreichs. Leipzig. Willis, Agriculture in the Tropics. Cambridge, 1914- Numerous more special works are mentioned under different heads throughout the book. Litliobium Bong. Melastomaceae (i). i Minas Geraes. Lithocardium L. = Corclia L. (Borag.). Lithophragma Torr. et Gray (Tellima R. Br. p.p.). Saxifragaceae (i). 15 N. Am. Lithophytum T. S. Brandegee. Solanaceae? i California. Litnospermum (Tourn.) L. Boraginaceae (iv. 4). 50 temp.; 3 in Brit, (gromwell). Lithraea Miers. Anacardiaceae (3). 3 S. Am. Litobrochia Presl = Pteris L. Litosanthes Blume. Rubiaceae (n. 5). 3 Malay Archipelago. Litsea Lam. Lauraceae (n). 150 trop. As., Austr. L. and bark medicinal. Littledalea Hemsl. Gramineae (10). i Tibet. Littonia Hook. Liliaceae (i). 4 S. and trop. Afr. Like Gloriosa. Littoralis (Lat.), growing on the beach. Littorella Berg. Plantaginaceae. i sp., i in S. Am., and /,. lacustris L. (shore-weed) in Eur. (incl. Brit.). This pi. exhibits two forms, one in water, another on land. The land form has a rosette of narrow 1. about 3 cm. long, which spread out upon the ground and show distinct dorsiventral structure. Fls. in groups of 3, one cf on a long stalk between two sessile ¥ , which are ripe before the sta. emerge from the former. Both sta. and style are very long and the fls. are wind-pollinated. Fr. a nut. The water form has much larger 1. which grow erect and are cylindrical (centric) in form and internal structure; no fls. are produced, but the plant multiplies largely by runners. It is often mistaken for Isoetes. Live-long, Sedum . Liveus (Lat.), pale lead colour. Livistona R. Br. Palmae (i. 2). 20 Indomal., Austr. Tall trees with fan leaves and panicles of $ fls. Fr. a berry. Lizard's tail (Am.), Saururus. Llagunoa Ruiz et Pav. Sapindaceae (n). 2 Ancles. Llanosia Blanco (Ternstroemia p.p.). Theaceae. i Phil. Is. Llavea Lagasca. Polypodiaceae. i trop. Am. Llavea Liebm. Inc. sed. 2 Mexico. Lloydia Salisb. Liliaceae (v). 5 N. temp. ; i on Snowdon. Loasa Adans. Loasaceae. 100 Mexico and S. Am., chiefly Mts. of Chili and Peru. Several cult. orn. fl. ; they possess stinging hairs. The fls. are generally yellow and face downwards. The nectaries, formed of combined stds. (see fam.), are large and conspicuous. The petals are boat-shaped and conceal the groups of sta. Loasaceae (EP., BH.}. Dicotyledons (Archichl. Parietales; Passi- florales BH.). 13 gen., 120 sp. Andine plants, many cult. orn. fl. LO DO ICE A 391 Mostly herbs, frequently twining, with opp. or alt., rarely slip., 1. The epidermis bears hairs of various kinds; esp. common are grapple- hairs and stinging-hairs. Fls. usu. in cymes, often sympodial, yellow (rarely white or red), 5 , usu. j-merous. Receptacle deeply hollowed out, so that the fl. is epig. K 5, imbr. ; C 5, free or united ; A 5 — oo . In the genera with oo sta. there is much difference as to the arrange- ment. In Mentzelia they are evenly distributed round the style, the outermost in some sp. being sterile. In other gen. it is the ante- sepalous sta. that are sterile, and in some, e.g. Loasa, Blumenbachia, 3 or more of the stds. are united to form a large coloured nectary, whose mouth is towards the centre of the fl. and partly obstructed by the other stds. G i or more commonly (3 — 5), with parietal plac. ; ovules i, several, or oo, anatr., with one integument ; style simple. Fr. various, often a caps., sometimes spirally twisted. Endosp. or not. Chief genera: Gronovia, Mentzelia, Loasa, Blumenbachia. Loaseae (/?.#". ) = Loasaceae. Lobe (1.), portion of a divided (not cpd.) 1. or stigma. Lobelia Plum, ex L. Campanulaceae (in). 220 trop. and temp. ; 2 in Brit., one in lakes (L. Dort manna L.); several cult. orn. fl. The fl. (see fam.) is twisted upon its axis through 180°, and is -j- . The anthers are syngenesious as in Compositae, and the style pushes through the tube thus formed, driving the pollen out at the top. Finally it emerges, the stigmas separate, and the ? stage begins. [See fam. and cf. Campanula, Phyteuma, Jasione and Compositae.] Loblolly bay, Gordonia Lasianthus L. ; - pine, Finns Taeda L. ; - sweetwood (W.I.), Sciadophyllum; -tree (W. I.), Cupania, Pisonia. Lobogyne Schlechter. Orchidaceae (n. a. in). 2 New Guinea, Poly- nesia. Lobostemon Lehm. Boraginaceae (iv. 5). 50 S. Afr. Lobostephanus N.E. Br. (Emicocarpus p.p. EP.). Asclepiadaceae (n. i). i S. Afr. Lobularia Desv. (Alyssum p.p. BH.}. Cruciferae (4). 4 Medit. Locellate, divided into small compartments. Lochia Balf. f. Caryophyllaceae (i. 4). i Socotra. Loclimium (01.), a thicket formation. Lochnera Reichb. (Vinca p.p. Bff.). Apocynaceae (i. 3). 3 trop. L. rosea Reichb., one of the commonest trop. weeds. Lockhartia Hook. Orchidaceae (n. 19). 20 trop. Am., W.I. No tubers; 1. crowded together. Loculament, loculus, a cavity in the ovary, usu. containing ovules ; loculicidal (fr.), splitting down midrib of each cpl. Locust, honey-, Gleditsckia; -tree, Robinia Pseudacacia L. (W. I.), Byrsonima, Hytnenaea. Loddigesia Sims. Leguminosae (ill. 3). i S. Afr. Lodh bark, Symplocos racemosa Roxb. (India). Lodhra Guill. =Symplocos Jacq. p.p. (Symploc.). Lodicularia Beauv. = Hemarthria R. Br. (£//.). = Rottboellia L. f. Lodicule, Gramineae. Lodoicea Comm. Palmae (n). i Seychelles, L. SecheUarum Labill., the double coco-nut or Coco de mer. Dioec. The fr. is one of the largest known and takes 10 years to ripen. The nut is bilobecl. The 392 LO DO ICE A fr. used to be found floating in the Indian Ocean long before the tree was discovered (see Treas. of Bot). Loeflingia L. Caryophyllaceae (i. 3). 5 Medit., C. As., N. Am. Loeselia L. Polemoniaceae. 18 Am. Loesenera Harms. Leguminosae (n. 3). i trop. Afr. Loewia Urb. Turneraceae. 2 trop. Afr. Loganberry, a supposed blackberry-raspberry hybrid, but cf. Journ. of Heredity, Nov. 1906, p. 504. Logania R. Br. Loganiaceae. 21 Austr., N.Z. Loganiaceae (EP., BH.). Dicots. (Sympet. Contortae ; Gentianales BH.}. 32 gen., 350 sp., trop. ; a few warm temp. (Am., As., N.Z., but none in Eur.). Trees, shrubs, and herbs with opp. slip. L; slips. often very much reduced. Many climbing plants, esp. Strychnos. Infl. as a rule cymose, various ; the fls. with bracts and bracteoles, usu. reg., £ , and 4 — 5-merous, with occasional increase in number in C and A. Disc small or absent. K (4 — 5), imbr. ; C (4 — ,s)> valv., imbr., or conv. ; A 4— 5, rarely i, epipet.; G (2), anteropost., 2-loc., or rarely imperfectly so, or i- or more-loc. ; style simple ; ovules usu. oo , amphi- or ana-tr. Caps., berry, or drupe. Endosp. Nearly allied to Apocynaceae, Gentianaceae, Solanaceae, Scrophulariaceae and Rubiaceae. See discussion in Nat. Pfl. Chief genera: Logania, Spigelia, Strychnos, Fagraea, Buddleia, Desfontainea. Logwood, Campeachy -, Haematoxylon campechianum L., Ceanothns Chloroxylon Nees ; bastard- (W.I.), Acacia Berteriana Sprang. Loheria Merrill. Myrsinaceae (n). i Phil. Is. Loiseleuria Desv. Ericaceae (i. 3). i north circumpolar, L. procumbent Desv. (Azalea procumbeus L. ), the trailing Azalea. Highlands of Scotland. The 1. are very wiry, and rolled back at the margins, thus reducing transpiration. Fls. reg., protogynous ; they come out shortly after the melting of the snow. Lolium L. Gramineae (12). 6 Eur., N. Air., temp. As. L. perenne L. in Brit, (rye-grass). Spikelets in a 2-ranked spike, and placed edgewise (this distinguishes the subtribe Lolieae from the Leplureae, to which Triticum and Hordeum belong). The rye-grasses are valu- able pasture and fodder. Lomandra Labill. (Xerotes R. Br. BH.). Liliaceae (in) (Juncaceae BJf.}. 30 Austr. Dioecious. Lomaria Willd. = Blechnum L. (Polypod.). Lomatia R. Br. Proteaceae (n). 9 E. Austr., Tasm., Chili. Lomatium Rafin. (Peucedanwu p.p. BH.\. Umbellif. (in. 6). 60 W. N.Am. Lomatophyllum Willd. Liliaceae (in). 3 Masc. Cult. orn. fl. Lomatozona Baker. Compositae (2). i Goyaz. Lombardy poplar, Populus nigra L., var. pyramiiialis Spach. Lomentose, like a lomentum, Raphanus; lomentum, a pod constricted between the seeds, breaking into i -seeded portions, Leguminosae. Lommelia Jumelle et Perrier. Palmaceae (iv. i). i Madag. Lonas Adans. Compositae (7). i S.W. Medit. Lonchitis L. Polypodiaceae. 8 trop. Am. and Afr., Madag. Lonchocarpus H. B. et K. Leguminosae (in. 8). 75 trop. Am., Afr., Austr. LOPEZ f A 393 Lonchomera Hook. f. et Thorns. (Mezzellia p.p.). Anonaceac (i). i Malay Peninsula. Lonchophora Dur. Cruciferae (4). i N. Afr. Lonchostephus Tul. Podostemaceae. i Amazon. Lonchostoma Wikslr. Bruniaceae. 3 S. Afr. London pride, Saxifraga 2i»ibrosa L. Long and short shoots, Berberis, Coniferae, Ginkgo, Phyllocladits, Pi nits. Longan, Nephelium Longana Cambess. Longetia Baill. Euphorbiaceae (A. I. i). i New Caled. Longi- (Lat. pref.), long. Long-moss, Tillandsia. Long-tongued insects, bees and Hymenoptera. Lonicera L. Caprifoliaceae. 100 %. L. Periclymenum L. (honey- suckle or woodbine), and others, in Brit. Mostly erect shrubs, a few twining, with opp. frequently connate 1. In the axils of many (e,g. L. tatarica L.) are serial buds, of which the lowest gives rise to the fls. usu. in pairs, the central fl. of the small dichasium not being developed. The fl. is frequently •[• , and gives rise to a berry. In some the pair of fls. produces two independent berries, in others the berries fuse into one as they form. Some sp. exhibit the 'fusion' even earlier; and one finds two corollas seated upon what at first glance appears a single inf. ovary. Dissection shows that in most cases the two ovaries are side by side, free from one another, in a com- mon hollow axis ; in a few cases, however, the union is more complete. The fl. of the honeysuckle is visited chiefly by hawk-moths (at night). The fl. opens in the evening, the anthers having dehisced shortly before this. The style projects beyond the anthers. The fl. moves into a horiz. position at the same time. At first the style is bent downwards and the sta. form the alighting place for insects. Later on the style moves up to a horiz. position, the sta. shrivel and bend down, and this is complete by the second evening when the next crop of buds is opening. At the same time the fl. has changed from white to yellow. The length of the tube keeps out all but very long- tongued insects (class F). Loofah, Luffa cylindrica M. Roem. Looking glass tree, Heritiera littoralis Ait. Loose pollen mechanism, cf. Acanthus, and Barlsia, Borago, Calluna, Cyclamen, Erica, Ericaceae, Euphrasia, Galanthus, Melampyrum, Pedicularis, Scrophnlariaceae, Solanwti. Loosestrife, purple, Lythrum, yellow, Lysimachia. Lopezia Cav. Onagraceae (2). 15 C. Am. Fl. -|- . The two upper petals are bent upwards a little way from the base, and at the bend there seems to be a drop of honey. In reality this is a dry glossy piece of hard tissue ; like the similar bodies in Parnassia it deceives flies. There are real nectaries at the base of the fl. There are two sta., of which the post, only is fertile; it is enclosed at first in the ant. one, which is a spoon-shaped petaloid std. In the early stage of the fl., the style is undeveloped and insects alight on the sta.; later the style grows out into the place first occupied by the sta., which now bends upwards out of the way. In L. coronata Andr., &c. there 394 LOPE/ 1.1 is an upward tension in the sta., a downward in the std., and an explo- sion occurs when an insect alights. Cult. orn. fl. Lophaeme Stapf. Gramineae (i i). i S. Afr. Lophactis Rafin. Compositae (inc. sed. ). i N.Ain. Lophanthera A. Juss. Malpighiaceae (n). i N. trop. S. Am. Lophanthus Benth. Labiatae (vi). 2 C. As., China. Fl. stalk some- times resupinate like that of Lobelia. Lophatherum Brongn. Gramineae (10). 4 E. As. Lophiocarpus Aliq. (Sagittaria p.p. BH.}. Alismaceae. 4 trop. Lophiocarpus Turcz. Chenopodiaceae (inc. sed. ). 28. Afr. LopMola Ker-Gawl. Amaryllidaceae (in), i All. N. Am. Lophira Banks ex Gaertn. f. Ochnaceae. 2 trop. Afr. The fr. of Z. alata Banks (African oak) has one sep. much, a second less, elongated. The seeds yield an oil on pressure; timber good. Lophium (Cl.), a hill formation. Lophocarpus Boeck. Cyperaceae (n). i Tonquin. Lophocereus Britton et Rose (Cereus p.p.). Cactaceae (in. iU 2 S.W. U.S. Lophogyne Tul. Podostemaceae. 2 Rio de Janeiro. Lopholaena DC. Compositae (S). 4 S. and trop. Afr. Lopholepis Decne. Gramineae (3). i India. Lophopappus Rusby. Compositae (12). i Bolivia. Lophopetalum Wight ex Arn. Celastraceae. 12 Indomal. Lophophora Coult. (Anhaloniitm p.p.). Cactaceae (in. 2). i N.Am. Lophophyllum Griff. (Peramphora p.p.£P.). Menisperm. i Indomal. Lophophytum Schott et Endl. Balanopho:aceae. 4 trop. S. Am. Lophopogon Hack. Gramineae (2). 2 Indomal. Lophopterys A. Juss. Malpighiaceae (i). i Guiana. Lophopyxis Hook. f. Icacinaceae (formerly Euph.). 3 Malaya. Lophoschoenus Stapf (Cyclocampe p.p.). Cyper. (n). 3 Seychelles, Borneo, New Caledonia. Lophosciadium DC. Umbelliferae (in. 6). 5 S. Eur., W. As. Lophospermum D. Don=Maurandia Orteg. p.p. (Scroph.). Lophostachys Pohl. Acanthaceae (iv. A). 12 Brazil, Peru. Lophostigma Radlk. Sapindaceae (i). i Bolivia. Lophostoma Meissn. (Linostoma BH.}. Thymelaeaceae. 3 Amazonas. Lophotocarpus Durand (Lophiocarpus p.p.). Alismaceae. i trop. Lopriorea Schinz. (Psilotrichum p.p.). Amarantaceae (2). i E. Afr. Lopseed (Am.), Phryma. Loquat, Eriobotrya japonica Lindl. Loranthaceae (EP., BH.}. Dicots. (Archichl. Santalales; Achlamydo- sporae BH.}. 21 gen., 520 sp., trop. and temp. The only genus in Brit, is Viscum, the mistletoe. An interesting fam. of parasites with green 1. Mostly small semi-parasitic shrubs attached to their hosts by suckers or haustoria — usu. regarded as modified adv. roots. A few root in the earth, e.g. the W. Austr. Nuytsia, which grows into a small tree 30 feet high. Most are fairly omnivorous in their choice of hosts, but a few are restricted to one or two. Where the parasitic root joins the host, there is not uncommonly an outgrowth, often of considerable size and complicated in shape. The parasitic root often branches within the tissue of the host, as in mistletoe. The stem is LORENTZIA 395 sympodial, often dichasial, e.g. in Viscum, and the 1. u.su. evergr. and leathery. Infl. cymose, the fls. usu. in little groups of 3 (or 2, by abortion of the central fl.). When the fls. are stalked, the bracts of the lateral fls. are always united to their stalks, up to the point of origin of the fl. (see Viscum and Loranthus). Infl. sometimes in spikes, with the fls. on the internodes as well as on the nodes. The recept. is hollowed out, and the P springs from its margin. In the Loranthoideae there is below the P an outgrowth of the axis in the form of a small fringe — the calycttlus. Some look upon it as a K, many as an outgrowth of the axis; and this is perhaps the safest view. P either sepaloid or petaloid. Fls. $ or unisexual. Sta. as many as, and (as in Proteaceae) united with, the P-leaves. The pollen is often developed in a great number of loculi, separate from one another, though often becoming continuous when mature. Ovary i-loc., sunk in, and united with, the receptacle, the ovules not differentiated from the placenta. Embryo-sacs > one, curiously lengthened (cf. Casuarina). Fr. a pseudo-berry or -drupe, the fleshy part really the receptacle. Round the seed is a layer of viscin, a very sticky substance. [For full details of the many interesting features of this fam., the infl., fl., pollen, development and structure of ovule and embryo-sac, fruit, seed, germination, haustoria, &c., see Engler in Nat. Pfl. and papers by Wiesner in Sitz. k. Akad. Wren, cm. 1894, and Keeble in Trans. Linn. Soc. v. 1896.] Classification and chief genera (after Engler) : I. LORANTHOIDEAE (\v\t\\cz\yc\i\\\s): Stiulhanthus, Lor- anthus, Psittacanthus. II. VISCOIDEAE (without calyc.) : Arceuthobium, Viscum. Diagram of Loranthus (after Eichler). c, calyculus. Loranthus L. Loranthaceae (i). 300 |* trop. and subtrop. Semi- parasites. $ or unisexual fls. in small cymes, the bracts adnate to the peduncles. Fr. like that of Viscum. See Nat. Pfl., Eichler's Bliithendiag. and papers by Wiesner and Keeble (cf. fam.). Lorate, loriform, strap-shaped. Lords and ladies, Arum niacitlatitm L. Lorentea Lag. = Pectis L. p.p. (Comp.). Lorentzia Griseb. (Pascalia BH., Wedelia p.p. EP.). Compositae (5). i S. temp. Am. Lorentzia Hieron. (Ayenia Loefl.). Sterculiaceae. 2 Argentina. 396 LOREYA Loreya DC. Melastomaceae (i). 8 S. Am. Loropetalum R. Br. Hamamelidaceae. i Khasias, S. China. Lorostelma Fourn. Asclepiadaceae (n. 3). i Brazil. Lortia Rendle. Euphorbiaceae (A. I. 8). 2 trop. Afr. Lote fruit, Zizy pirns Lotus Lam. (Medit.). Lotononis Eckl. et Zeyh. Leguminosae (in. 3). 65 Afr., Medit. Lotoxalis Small (Oxalis p.p.). Oxalidaceae. isN.Am. Lotus (Tourn.) L. Leguminosae (in. 5). 15 temp. Eur., As., S. Afr., Austr. L. comiculatus L., bird's foot trefoil, and others, in Brit. The floral mechanism (class H) is typical of many of the fam. (q.v.). The keel is united above and below, leaving only a small opening at the apex. The pollen is shed in bud into the tip of the keel, and the filaments of five sta. thicken out below the anthers, together forming a piston, which, when the keel is depressed, forces the pollen out in a stream at the apex. The style is immersed in the pollen, but only becomes receptive on being rubbed, so that the fl. has a chance of cross-fert. The plant is useful for pasturage. Lotus, Zizyphus; -berry (W.I.), Byrsonima; sacred-, Nymphaea, Nehinibium. Loudonia Lindl. Haloragidaceae. 3 W. Austr. Louisiana grass (Am.), Paspaluw platycaule Poir. Lourea Neck. Leguminosae (m. 7). 4 trop. As. and Austr. Loureira Meissn. Inc. sed. i Cochinchina. Lourya Baill. Liliaceae (vin). i Cochinchina. Lousewort, Pedicularis. Louteridium S. Watson. Acanthaceae (iv. A). C.Am. Louvelia Jumelle et Perrier. Palmaceae (iv. i). i Madag. Lovage, Ligusticum scoticum L. Love apple, Lycopersicum esciilentniii Mill.; -grass (Ceylon), Andro- pogon aciculaftis Retz. ; -in a mist, Nigel/a, (W. L), Passi flora foetida L. ; - lies bleeding, Amaranthus. Lovoa Harms. Meliaceae (n). 2 trop. Afr. LovomaO. F. Cook (Ptychospernia p.p.). Palmaceae ( I v. i). i N. Austr. Lowia Scortech. Musaceae. 2 Malaya. Lowiara x . Orchidaceae. Hybrid between Brassavola, Laeiia, and Sophronitis. Loxanthera Blume (Loranthus p.p. BH.}, Loranth. (i). 2 Borneo, Java. Loxocalyx Hemsl. Labiatae (vi). i China. Loxocarpus R. Br. (Didymocarpus p.p.). Gesneriaceae (i). 3 Malaya. Loxocarya R. Br. Restionaceae. 18 S.W. Austr. Loxococcus H. Wendl. et Drude. Palmae (iv. i). i Ceylon. Loxodiscus Hook. f. Sapindaceae (11). i New Caledonia. Loxogramuie (Blume) Presl = Polypodium L. (Polypod.). Loxonia Jack. Gesneriaceae (i). i Sumatra, Java. Loxopterygium Hook. f. Anacardiaceae (3). 5 trop. S. Am. Loxostemon Hook. f. et Thorns. Crucifera'e (4). 2 Sikkim to Yunnan. Loxostigma C. B. Clarke. Gesneriaceae (i). 2 Himalaya, China. Loxostylis Spreng. ex Reichb. Anacardiaceae (3). i Cape Colony. Loxothysanus Robinson (Baliia p.p.). Compositae (6). 2 Mexico. LUPIN US 397 Loxsoma Br. Hymenophyllaceae? i N.Z. It agrees with Tricho- manes in the sorus, but differs widely in the veg. habit, which is that of Cyatheaceae, and is probably better placed in a separate fam.; cf. Gwynne-Vaughan, Ann. Bot. Xiv. 1901, p. 71. Loxsomaceae. Cf. Filicineae, and Loxsoma. Lozanella Greenman. Ulmaceae. i Mexico. Lubinia Comm. ex Vent (Lysimackia p.p.). Primulaceae. 4 S. Afr., Masc. Is., Japan. Lucaea Kunth = Arthraxon Beauv. (Gram.). Lucens, lucidus (Lat.), with shining surface. Lucerne, Medicago saliva L. Lucilia Cass. Compositae (4). 25 S. Am. Luciliopsis Wecld. Compositae (4). 2 Bolivia. Lucinaea DC. Rubiaceae (i. 7). 10 Malaya. Luculia Sweet. Rubiaceae (i. 5). i Himal., Khasias. Lucuma Molina (inch Vitdlaria Gaertn. f.). Sapotaceae (i). 50 trop. Am. The fr. of L. hi/era Molina is ed. Lucya DC. Rubiaceae (i. 2). i W.I. Ludia Comm. ex Juss. Flacourtiaceae (4). 2 E. Afr., Madag., Masc. LudoviaBrongn. Cyclanthaceae. 2 trop. S. Am. S fls. as in Carludovica, ? sunk to stigmas with rudimentary P. Climbers. Ludovica Vieill. ex Guillaumin = Bikkia Reinw. (Rubi.). Ludwigia L. Onagraceae (2). 20 cosmop. ; i Brit. Ludwigiantha Small (Ludwigia p.p. ). Onagraceae (2). i N. Am. Lueddemannia Reichb. f. (Acineta p.p. EP.}. Orchid, (n. 13). 4 N. S.Am. Luederitzia K. Schum. (Pavonia p.p.). Malvaceae (3). 2 S. Afr., Somaliland. Luehea Willd. Tiliaceae. 18 trop. Am., W.I. Luerssenia Kuhn. Polypodiaceae. i Sumatra. Luffa (Tourn.) L. Cucurbitaceae (3). 7 trop. L. cylindrica M. Roem. (L.aegyptiaca Mill.) furnishes the loofah or bath sponge (the vascular bundle net of the pericarp). Fr. of most ed. Lugonia Wedd. Asclepiadaceae (n. i). 2 Peru, Bolivia. Luina Benth. Compositae (8). i N.W. U.S. Luisia Gaudich. Orchidaceae (n. 20). 15 trop. As. to Japan. Cult. Lumnitzera Willd. Combretaceae. 2 palaeotrop. , in mangrove (q-v.) swamps. Fr. floated by ocean currents. Lunania Hook. Flacourtiaceae (7). 5 W.I., Brazil. Lunaria Tourn. ex L. Cruciferae (2). 2 Eur. L. biennis Moench. (L. annua L.) is the honesty of gardens. Lunasia Blanco. Kutaceae (i). 5 Malay Archipelago. Lunate, half-moon-shaped. Lundia DC. Bignoniaceae (i). 10 E. S.Am. Lunellia Nieuwland. Scrophulariacetfe (in. i). 3 U.S. Lung-wort, Pulmonaria, (Am ) Mertensia. Lupin, Lupinus. Lupinaster Buxb. =Trifolium Tourn. p.p. (Legum.). Lupinus (Tourn.) L. Leguminosae (in. 3). 150 Am., Medit. Floral mechanism like Lotus. The fr. explodes, its valves twisting spirally. Several are cult. orn. rl., or used as fodder. 398 LUPSIA Lupsia Necker (Galactites BH.). Compositae (n). 3 Medit. Luridus (Lat.), dingy brown or yellow. Luteus(Lat.), yellow; luteolus, pale yellow; lutescens (Lat.), yellowish. Luvunga Buch.-Ham. Rutaceae (v). 5 Indomal. Luxembergia A. St Hil. Ochnaceae. 8 Brazil. Luziolajuss. Gramineae (6). 6 Brazil to Alabama. Luzonia Elmer. Leguminosae (in. 10). i Luzon. Luzula DC. Juncaceae. 65 temp., chiefly Old World; 6 in Brit, (wood-rush). Rhiz. as in Juncus; 1. usu. flat. Luzuriaga Ruiz et Pav. (Enargea Banks). Liliaceae (x). 3 S. Am., N.Z. Lyallia Hook. f. Caryophyllaceae (1.3). i Kerguelen. Lycaste Lindl. Orchidaceae (n. 12). 30 trop. Am. Epiphytes. A chin is formed by an axial outgrowth from the column. Lychniothyrsus Lindau. Acanthaceae (iv. A), i Brazil. Lychnis (Tourn.) L. (BH. incl. Agrosteinma L., Gitkago Adans., Melandriuni Roehl, Viscaria Riv.). Caryophyllaceae (ii. i). loN. temp. |^: ; 3 in Brit., incl. L. Flos-cuciili L. (ragged robin). Fls. protandrous, suited to bees and Lepidoptera. Thefls. often show the sta. filled with a black or brown powder, instead of pollen ; this is the spores of the fungus Ustilago antherantm, which are thus dis- tributed from plant to plant, like pollen, by the visiting insects. [For L. dioica L. see Melandrium.] Lychnodiscus Radlk. Sapindaceae (i). 2 W. Afr. Lychnophora Mart. Compositae (i). 17 S. trop. Brazil. Lychnophoriopsis Sch. Bip. Compositae (i). 2 Minas Geraes. Lycium L. Solanaceae (2). 75 temp. Many have thorny twigs; L. of nun L. (Kaffir thorn) is used for hedges in S. Afr. L. barbarian L. often cult, under the name tea-plant. Lycomormium Reichb. f. Orchidaceae (ii. 13). i Peru. Lycopersicum Hill (Solanum p.p. EP.). Solanaceae (2). 10 S. Am. L. esculent inn Mill. (Solatium Lycopersicum L.) is the tomato or love-apple. Lycopodiaceae. Lycopodiales, Eligulatae. 2 gen., 190 sp., trop. and temp. The fertilised ovum gives rise directly to the leafy plant ; the embryo has a suspensor and a foot; and its upper part at first forms a tuber-like organ, the protocor/n, from which the 1. and stem develope. In P. the stem is short and unbranched, in L. long and much branched, bearing small simple 1., and roots developed in aero- petal succession. The sporangia are axillary, and form as a rule a dense terminal cone or strobilus. The spores are all of one kind and give rise on germ, to fairly large monoec. prothalli. Classification and genera : Small pi. with a few rad. 1. and head of sphorophyllson leafless stalk: Phylloglossum. Larger pi. with no rad. 1. and sporophylls on leafy stalk : Lycopodium. Lycopodiales. One of the main divisions of Pteridophyta. Mostly leafy plants, with well developed stems and small unbranched 1. The sporophylls are usu. massed together into cones, recalling those of the Gymnosperms. Sporangia single at base of 1. on upper side, or in axil, i-loc. They are classified as follows. L YROCARPA 399 A. ELIGULA TAB (1. with ligule : spores all alike). B. L[GULA7"AE (1. with ligules: spores of two kinds). Lycopodium L. Lycopodiaceae. 185 trop. and temp.; 5 in Brit., chiefly in mountain districts (club-mosses). The commonest is L. flavatum L. (often called stag-horn moss) ; the others are L. Selago L., L. alpimiin L., L. aniiotinuin L., and L. inundatnin L. All but the last are xero. evergr. pi. with hard wiry 1. The stem branches frequently, apparently dichotomously, but in reality usu. in a mono- podial manner. Upon it are borne the roots, which branch dichoto- mously, and are developed in acropetal succession. The 1., narrow and unbranched, are usu. placed spirally upon the stem, but in some form four ranks, as in most Selaginellas. Many have veg. repr. by small bulbils in the 1. axils. [Prothalii, see Bruchmann in Bot. Zeit. 1899, p. 6.] Sporangia all alike in structure, containing spores of one kind only ; placed upon the bases of 1. which are usu. crowded together to form a terminal spike or strobilus. In Z . Selago some or all of the sporangia are often replaced by small bulbils (cf. Polygonutn vivi- paruni, Allium sp., Globba. &c.). Lycopsis L. Boraginaceae (iv. 3). 3 Eur., As. L. arvemis L., small bugloss, in Brit. Lycopus Tourn. ex L. Labiatae (vi). 10 N. temp. L. curopaeus L., gipsy wort, in Brit. Lycoris Herb. Amaryllidaceae (i). 4 Japan. Lycoseris Cass. Compositae (12). 10 C. Am. to Peru. Lycurus H. B. et K. Gramineae (8). 2 Mexico, New Mexico. Lygeum L. Gramineae (6). i Medit., L. Spartuin Loefl., one of the esparto-furnishing grasses (cf. Stipa and Ampelodesm'a). Lyginia K. Br. Restionaceae. i S.W. Austr. Lygisma Hook. f. Asclepiadaceae (n. 3). i Burma. Lygistum P. Br. = Manettia L. p.p. (Rubi.). Lygodesmia D. Don. Compositae (13). 6 N. Am. Lygodisodea Ruiz et Pav. (Paederia p.p. EP.). Rubi. (n. 6). 4 trop. Am. Lygodium Sw. Schizaeaceae. 25 trop. and subtrop. Twining ferns. The stem remains comparatively undeveloped, but the 1. has un- limited apical growth, and the long midrib twines around supports like the stem of the hop, bearing pinnae at intervals. The 1. are borne on the stem in one dorsal row. The sporangia are in a double row on the back of the fertile pinnae, and each is surrounded by a cup-like indusium Lyme grass, Elynnts arenarins L. Lyonia Nutt. Ericaceae (n. i). 16 E. As., N. Am., i circumpolar. Lyonnetia Cass. = Anthemis Michx. p.p. (Comp.). Lyonothamnus A. Gray. Rosaceae (inc. sed.). i California. Lyonsia R. Br. Apocynaceae (u. 2). 15 Austr., New Guinea. Lyperanthus R. Br. Oichidaceae (n. 2). 8 Austr., N. Z., New Caled. Lyperia Benth. (Chaenostoma p.p. E/3.}. Scrophulariaceae (u. •;). 33 S. Afr. Lyrate (1.), dhided with large term, lobe, as in mustard. Lyrocarpa Hook, et Harv. Cruciferae (3). i California. 400 L YSIAS Lysias Salisb. (Habenaria p.p. BH., Platanthera p.p. EP.). Orchid- aceae (n. i). 2 N. Am. Lysicarpus F. Muell Myrtaceae (n. i). r Queensland. Lysichiton Schott. Araceae (in), i temp. E. As. Lysidice Hance. Leguminosae (u. 3). r S. China. Lysiella Rydberg (Habenaria p.p.). Orchid. (11. i). £ N. Am. Lysiloma Benth. Leguminosae (i. i). 10 trop. Am., W.I. LysimacMa (Tourn.) L. Primulaceae. 60 temp, and subtrop. ; 4 in Brit. L. vulgaris L., yellow loosestrife, is said by Mtiller to occur in two forms, one in sunny places with large fls. suited to crossing, and one in shady spots with small self-fert. fls. L. neinoru»i L. (yellow pimpernel) and L. Nuiiinntlnria L. (moneywort) also in Brit. Lysimachiopsis A. A. Heller (Lysimachia p.p.). Primul. 4 Hawaiian Is. Lysinema R. Br. Epacridaceae (2). 5 W. Austr. Lysionotus D. Don. Gesneriaceae (i). 5 Himalaya, China. Lysiosepalum F. Muell. Sterculiaceae. "2 W. Austr. Lysiostyles Benth. Convolvulaceae (i). i Guiana. Lysipomia H. B. et K. Campanulaceae (in). 7 Andes. Lytanthus Wettst. (Globularia p.p.). Globular. 2 Canaries, Azores. Lythraceae (EP. ; BH. incl. Oliniaceae, Pitnicaceae). Dicots. (Archichl. Myrtiflorae). 21 gen., 500 sp., all zones but frigid. Herbs, shrubs, or trees ; 1. usu. opp., entire, simple, with very small stipules or none. Fls. in racemes, panicles, or dichasial cymes, ? , reg. or -|- , usu 4- or 6-merous. The axis (' calyx-tube') is hollow, generally tubular. The sepals are valvate, and frequently possess an epicalyx, formed, as in Potentilla, of combined slips. Petals crumpled in bud, sometimes absent. Sta. inserted (often very low down) on calyx-tube, typically twice as many as sepals, but sometimes fewer or oo . G with simple style and usu. capitate stigma ; 2 — 6-loc. , at the base at least, rarely i-loc. with parietal placenta. Ovules usu. oo , anatr., ascending. The fls. of Lythrum (ij.v.) and others are heterostyled. Dry fr., usu. cap- sular. No endosp. A few yield dyes (Lawsonia, &c. ), or are medicinal. Chief genera: Peplis, Lythrum, Cuphea, Lagerstroemia. Lythrum L. Lythraceae. 23 cosmop. ; 2 in Brit., incl. L. Salicaria L. (purple loosestrife). The 6-merous fls. are sol. or in small axillary dichasia like Labiatae. Each has 12 sta. in two whorls of different length, and the style again is of different length to any of the sta. Three forms of fl. occur (frimorpkism), each on a separate pi.; they are distinguished as long- mid- and short-styled fls. The diagram illus- trates the arrangement of parts (S= stigma, C A A A = anthers, B = base of fl.), as seen in side ^a g 3 ^3 view. It is evident that an insect visiting \ '2 .2 „ 2 the fls. will tend on the whole to transfer j,1 V,1 p pollen from A3 to S3, A2 to S2) Ax to S1( , ., , rather than from sta. of one length to style , ""' a , of another, for it will enter these fls. in the Styled same way and to the same depth. The sta. and style project so far that an insect can alight directly upon them. Darwin (Forms of Flowers) showed by a long series of experiments that the best results are obtained by pollinating 83 from AS, or Sj from Aj, &c. , i.e. by MACLUR'A 4° i crossing two plants. The number of seeds thus obtained is much greater and their fertility higher than if S2 or Sj be fertilised from A3, or any other such union be made. Fertilisation of a stigma by sta. of corresponding length Darwin terms legitimate, by sta. of a different length illegitimate. The offspring of illegitimate fert. are few, and have the sterility and other sexual characters of hybrids. As in nearly all other heterostyled pi., the longer the sta. the larger the pollen grains, and the longer the style the larger the papillae of the stigma. Maba Forst. Ebenaceae. 70 trop. and subtrop. The wood of some is used as a substitute for ebony. Fr. of some ed. Mabea Aubl. Euphorbiaceae (A. n. 7). 16 trop. Am. Mabee bark (W.I.), Ceanothus reclinatus L'Herit. Macadamia F. Muell. Proteaceae (n). s Austr. (nut-tree). Seeds ed. Macairea DC. Melastomaceae (i). 20 trop. S. Am. Macaranga Thou. Euphorbiaceae (A. n. 2). roo palaeotrop. M. caladifolia Becc. has hollow peduncles inhabited by ants. Macarisia Thou. Rhizophoraceae. 2 Madag. Macarthuria Huegel ex Endl. Aizoaceae (i). 3 Austr , S.E. and S.W. Macary butter (W.I.), Picramnia Antidesma Sw. Macassar oil, Cananga odorata Hook. f. Macaw bush (W. I.), Solatium mammosiim L. ; -tree, Acrocomia. Macbridea Ell. ex Nutt. Labiatae (vi). 2 S.W. U.S. MaccMe, maqui, the copse association of Mediterranean coasts. Macdougalia A. A. Heller. Compositae (6). i N. Am. Mace, Myristica fragrans Houtt.; -reed, Typha. Macfadyena A. DC. Bignoniaceae (i). 3 S. Am. Macgregoria F. Muell. Stackhousiaceae. i E. Austr. Macgregorianthus Merrill. Thymelaeaceae. r Luzon. Machadoa Welw. ex Benth. et Hook. f. Passifloraceae. i trop. Afr. Machaeranthera Nees (Aster p.p. BH.}. Compositae (3). 45 N. Am. Machaerium Pers. Leguminosae (in. 8). 65 trop. Am. Resembles Dalbergia. Many are lianes, climbing by sensitive lateral shoots, and provided with recurved stipular thorns. Some of the jacaranda tim- bers (rosewoods) are furnished by this gen. (cf. Dalbergia). Machaerocarpus Small (Damasonium p.p.). Alism. i California. Machaonia Humb. et Bonpl. Rubiaceae (n. 2). 12 trop. Am., W. I. Machilus Nees. Lauraceae (i). 20 S.E. As. Mackay bean, Entada scandens Benth. Mackaya Harv. (Asystasia BfL). Acanthaceae (iv. B). \ S. Afr. Cult. orn. fl. Mackinlaya F. Muell. Araliaceae (3). 2 Queensland. Macleania Hook. Ericaceae (in. 2). 15 W. trop. Am. Cult. orn. fl. Macleya Rchb. (Bocconia p.p.). Papaveraceae (n). i China, Japan. Cult. orn. fl. Macludrania x Andr£. Urticaceae. Hybrid Maclura-Cudrania. Madura Nutt. Moraceae (i). i S.W. U.S., M. aurantiaca Nutt. (bow- wood or Osage orange). The tree bears thorns (branches). Fls. dioec., the cf in pseudo-racemes, the ? in pseudo-heads ; individual fls. like Morus. After fert. each ? fl. produces an achene enclosed in the fleshy P, and at the same time the common recept. swells up into a W. 26 402 MACLURA fleshy mass, so that a large yellow multiple fr. is formed. The wood is used for bows, carriage-poles, &c. The 1. are used for feeding silkworms. [J7. tinctoria D. Don, &c. = Chlorophora.] Macnatoia Benth. Ericaceae (iv. i). i S.W. Cape Colony. Macnemaraea \Villem. Inc. sed. i China. Macodes Lindl. Orchidaceae (n. 2). 2 Malaya. Cult. orn. 1. Macomariax Rolfe. Orchid hybrid, Macodes and Haemaria. Macoubea Aubl. Inc. sed. i Guiana. Macowania Oliv. Compositae (4). 2 S. Afr. Macphersonia Blume. Sapindaceae (i). 5 trop. E. Afr., Madag. Macqui berry, Aristolochia Maqui L'Herit. Macrachaenium Hook. f. Compositae (12). i Patagonia. Macradenia R. Br. Orchidaceae (n. 19). 5 W.I., Guiana, C. Am. Macranthera Torr. ex Benth. Scrophulariaceae (in. 2). 2 S. U.S. Macranthisiphon Bur. Bignoniaceae (i). i Ecuador, Peru. Macreightia A. DC. in DC. = Maba Forst. p.p. (Eben.). Macro- (Gr. prel.), long, large ; -podous (embryo), without cotyledons ; -sporangium, that which holds -spores, the larger when there are two kinds, Ptertdophyta. Macrocalyx Costantin et Poisson. Malvaceae (4) i Madag. Macrocalyx Miers. Rubiaceae (inc. sed.). Nomen. Macrocarpaea Gilg (Lisianthus BH.}. Gentian, (i). 10 Cuba, trop. S. Am. Macrocarpium Nakai (Cormis p.p.). Cornaceae. i Japan. Macrocentrum Hook. f. Melastomaceae (i). 3 Guiana, E. Brazil. Macrochaetium Steud. (Telraria p.p. EP.). Cyperaceae (n). i S. Afr. Macrochordion de Vriese (Aechmea p.p. BH.). Bromeli. (4). 6 S. Am. Cult. orn. fl. Macroclinidium Maxim. Compositae (12). 2 Japan. Macroclinium Barb. Rodr. = Ornithocephalus Hook. p.p. (Orch.). Macrocnemum P. Br. Rubiaceae (i. 5). 10 W.I., S. Am. Macrococculus Becc. Menispermaceae. i New Guinea. Macrodendron Taub. Cunoniaceae. i Rio fie Janeiro. Macrodiscus Bur. (Distictis Mart.). Bignoniaceae (i). r W.I. Macrolobium Schreb. ( Vouapa Aubl.). Legumin. (11.3). 20 trop. Am., Afr. Macrolotus Harms (Argyrolobmm p.p. EP.}. Legum. (in. 3). i trop. Afr. Macromeria D. Don. Boraginaceae (iv. 4). 8 C. and S. Am. Macropanax Miq. Araliaceae (i). 3 Sikkim to Java. Macropelma K. Schum. Asclepiadaceae (i). i E. Afr. Macropeplus Perkins. Monimiaceae. i E. Brazil. Macropetalum Burch. ex Decne. Asclepiadaceae (n. 3). 2 Cape Colony. Macrophloga Becc. (Chrysalidocarpus p.p.). Palm. (iv. i). r Madag. Macropidia J. Drumm. ex Harv. Amaryllidaceae (in), i W. Austr. Macropiper Miq. (Piper p.p. BH.]. Piperaceae. 6 Polynesia. Macroplectrum Pfitz. (Angraecum p.p. ). Orchidaceae (n. 20). i Madag., Masc. (A. sesquipedale, q.v.). MacropodandraGilg. Buxaceae. i trop. C. Afr. Macropodium (R. lir. in) Ait. Cruciferae (4). i C. As. MA GNOLIA CEAE 403 Macropsychanthus Harms. Leguminosae (in. 10). 2 New Guinea, Phil. Is. Macropterantb.es F. Mue.ll. Combretaceae. 4 N. Austr. Macrorhamnus Baill. Rhamnaceae. i Madag. Macrorh.yncb.us Less. =Troximon Null. (Comp.). Macrorungia C. B. Clarke. Acanthaceae (iv. B). 4 trop. and S. Afr. Macroscepis H. B. et K. Asclepiadaceae (n. i). 7 trop. Am. Macrosepalum Regel et Schmalh. Crassulaceae. i Turkestan. Macrosiphonia Muell.-Ajg. Apocynaceae (11. i). 10 trop. Am. Xero. Macrosolen Blume = Loranthus L. p.p. (BH.) — Elytranthe Blume p.p. Macrosphyra Hook. f. Rubiaceae (i. 8). 2 W. trop. Afr. Macrostegia Nees in DC. Acanthaceae (iv. A), r Peru. Macrostylis Bartl. et Wendl. Rutaceae (i). 10 S.W. Cape Colony. Macrotoiuia DC. Boraginaceae (iv. 4). 8 Medit. to Himalaya. Macrotorus Perkins (Mollinedia p.p.)- Monimiaceae. i Rio de Janeiro. Macrotropis DC. =Onnosia Jacks. (Legum.). Macrozamia Miq. Cycadaceae, q.v. 15 Austr. Macrozanonia Cogn. (Zanonia p.p.). Cucurbitaceae (i). i Java. Maculatus (Lat.), spotted. Mad-apple (W.I.), Solatium Melongena L. Madagascar clove, Ravensara aromatica J. F. Gmel.; -plum, Fla- conrtia\ -rubber, Landolphia, &c. Madar fibre, Calotropis gigantea Ait. Madarosperma Benth. Asclepiadaceae (u. i). i Upper Amazon. Maddenia Hook. f. et Thorns. Rosaceae (v). 2 Himalaya. Madder, Rubia tinctorum L. ; field -, Sherardia arvensis L. ; Indian - (W.I.), Oldenlandia umbellata L. Madia Molina. Compositae (5). 15 W. Am. M. sativa Mol. (madi, Chili; tanveed, U.S.), cult, for the oil from the seed. Madras hemp, Crolalariajuncea L. Madre de cacao (W.I. ), Erythrina nmbrosa, H. B. et K. Madronella Greene (AfonarJel/a p.p.). Labiatae (vi). 35 N. Am. Maerua Forsk. Capparidaceae (n). 30 trop. Afr., As. The fr. is a berry, constricted between the seeds like a lomentum. Maesa Forsk. Myrsinaceae (i). 35 trop., except Am. Maesobotrya Benth. in Hook. Euphorbiaceae (A. i. i). 3 trop. Afr. Maesopsis Engl. Rhamnaceae. 2 Irop. E. Afr. Mafekingia Baill. (Raphiacme EP.). Asclepiadaceae (i). i S. Afr. Mafootoo-withe (W.I.), Entada scandens Benth. Maga Urb. (Thespesia p.p.). Malvaceae (4). r Porto Rico. Magnistipula Engl. Rosaceae (vi). 3 trop. Afr. Magnolia L. Magnoliaceae. 21 As., N. Am. (esp. trop.). Trees with sheathing slips, covering the bud, and term. fls. P petaloid, except sometimes the outermost L, and in whorls. Sta. and cpls. oo , on a lengthened torus. Protogynous. Fr. an aggregate of follicles ; each dehisces by its dorsal suture, and the seed dangles out of it on a long thread formed by the unravelling of the spiral vessels of the funicle. The outer integument of the ovule becomes fleshy as it ripens, and the seeds may thus be distr. by birds. Several cult. orn. fl. Magnoliaceae (£'/".; BH. incl. Trochodendraceae}. Dicots. (Archichl. Ranales). 9 gen., 70 sp., trop. and subtrop. trees and shrubs (some 26 — 2 404 MAGNOLIACEAE climbing). They have alt. 1., which in the Magnolieae have big slips., united to form a thimble-like hood, covering in the bud all the younger 1. As each 1. expands it throws off the hood of the next older 1. The fls. are term, (in most) or axillary, usu. sol., $ or uni- sexual. In the Magnolieae the P is cyclic, in the rest of the fam. it is spiral, as are the sla. and cpls. throughout. P usu. petaloid ; A oo , hypog. ; G usu. oo , on a long torus. Follicle, berry, or samara ; seed albuminous, the endosp. not ruminate. The timber of most M. is good; Illicium, &c., furnish economic products. Magnolia and Liriodendron are cult, for their fls., &c. Chief genera : Magnolia, Liriodendron, Kadsura, Schizandra, Illicium, Drimys. Magonia A. St. Hil. Sapindaceae (n). 2 Brazil. Maguey, Agave americana L. Magydaris Koch ex DC. Umbelliferae (in. 4). 2 All. Eur. Mahafalia Jumelle et Perrier. Asclepiadaceae (IF. 4). i Madag. Mahernia L. (Hermannia EP.). Sterculiaceae. 80 Afr. Mahoe (W. I.), Paritium, Thespesia, Hibiscus. Mahogany, strictly speaking the timber of Simetenia Mahogoni Jacq. (Honduras, &c.) or Khaya senegalensis A. Juss., but the commercial timbers include many that resemble these, e.g. those of Cedrela, Melia, Dysoxylum and other Meliaceae, and of Kiggelaria, &c. See Stone, Timbers of Commerce. Mahonia Nutt. (Berberis p.p.). Berberidaceae. 40 *. Cult. orn. shrubs. Mahua, mahwa, Bassia latifolia Roxb. Mahurea Aubl. Guttiferae (i) (Ternstr. BH.). 4 Guiana, Venezuela. Mahya Cordem. (Sphacele p.p.). Labiatae (vi). i Bourbon. Maianthemum (Weber in) Wigg. Liliaceae (vil). i N. temp, (inch Brit.). Fl. 2-merous, protogynous. Maidenhair fern, Adiantum; -tree, Ginkgo biloba L. Maidenia Rendle. Hydrocharidaceae. i N.W. Austr. Maieta Aubl. Melastomaceae (i). 8 trop. Am. Some have bladdery outgrowths of the 1. inhabited by ants (cf. Duroia). Fr. ed. Maihuenia Phil. (Opuntia p.p.). Cactaceae (il). 3 Chili, Argentina. Maillardia Frapp, et Duch. Moraceae (i). i Bourbon. Maillea Parl. (Phleum p.p. EP.}. Gramineae (8). i Medit. Is. Maingaya Oliv. Hamamelidaceae. r Penang. Mairea Nees. Compositae (3). 10 Cape Colony. Maize, Zea Mays L. ; water -, Victoria regia Lindl. Majidea J. Kirk ex Oliv. (Harpullia p.p. EP.). Sapind. (n). i trop. Afr. Majoe-bitter (W.I.), Picramnia Antidesma Sw. Majorana (Tourn.) Rupp. (Origanum p.p. BH.). Labiatae (vi). 6 E. Medit. Majus (Lat.), greater. Malabaila Hoffm. Umbelliferae (in. 6). 25 Medit., W. As. Malacantha Pierre (Chrysophyllum p.p.). Sapot. (i). 5 W. trop. Afr. Malacca cane, Calamus. Malachium Fries = Stellaria L. p.p. (Caryophyll.). Malachra L. Malvaceae (3). 6 trop. and subtrop. Am. MALPIGHIA 405 Malacocarpus Salm-Dyck (Echinocactus p.p. BH.}. Cactaceae (ill. i). 8 S. Brazil, Uruguay. Malacochaete Nees = Scirpus Tourn. p.p. (Cyper.). Malacolepis A. A. Heller (Malacothri* p. p.). Compos. (13). i Calif. Malacomeles Decne. (Naegelia Lindl.). Rosaceae (n). i Mexico. Malacophilous, fert. by snails. Malacothamnus Greene (Malvastriini p.p.). Malvaceae (2). 9 N. Am. Malacothrix DC. Compositae (13). 15 Calif. Malaisia Blanco. Moraceae (i). i Malaya, Austr., Polynesia. Malanea Aubl. Rubiaceae (n. 2). 10 trop. S. Am., W.I. Malaxis Soland. ex Sw. Orchidaceae (n. 4). i N. temp. (incl. Brit.), M. paludosa Sw. The fl. is twisted through 360°, so that the la- bellum is again uppermost. See Darwin's Orchids, p. 130. Malay apple, Eugenia malaccensis L. Malcomia R. Br. (Malcolmia Spreng.). Cruciferae (4). 30 Medit. Male bamboo, Dendrocalamus stnctus Nees ; - fl. , with sta. only. Malesherbia Ruiz et Pav. Malesherbiaceae. 20 W. S. Am. Herbs or undershrubs with alt. often deeply lobed exstip. 1. , often very hairy. Fls. £ , reg. , 5-merous ; axis tubular, running out into a disc and gyno- phore, with which the sta. are concrescent. G (3) with parietal plac. and oo ov. ; styles 3 — 4 below apex of ovary. Caps. No aril. Malesherbiaceae (EP. ; Passifloraceae p.p. BH.}. Dicots. (Archichl. Parietales). i gen., Malesherbia, q.v. It differs from Passifl. in having no aril and the styles more deeply inserted and widely separated ; from Turneraceae in having no aril, in the aestivation of the C, and in the persistent recept. Malinvaudia Fourn. Asclepiadaceae (n. 4). i S. Brazil. Mallee (scrub), Eucalyptus, dwarf sp. like E. dumosa Cunn. , &c. Malleola J. J. Sm. et Schlechter (Saccolabium p.p.). Orchid, (n. 20). 6 Malaya. Mallinoa Coult. Compositae (2). i Guatemala. Mallophora Endl. Verbenaceae (3). i W. Austr. Mallostoma Karst. (Arcythophyllnm EP.). Rubiaceae (i. 2). 51 trop. Am. Mallotonia Britton (Tournefortia p.p.). Boraginaceae (in), i W.I. Mallotopus Franch. et Sav. Compositae (8). i Japan. Mallotus Lour. Euphorbiaceae (A. n. 2). 80 palaeotrop. Kamala dye is obtained from the caps, of M. philippinetKis Miill.-Arg. (Ceylon — Austr.) . Mallow, Malva and other Malvaceae ; marsh -, Althaea offidnalis L. , musk -, rose -, Hibisats ; tree -, Lavatera. Malmea Fries. Anonaceae (i). i Brazil. Malope L. Malvaceae (i). 3 Medit. The 3 1. of the epicalyx are very large. Cpls. oo , in vertical rows (see fain.). Cult. orn. fl. Malortiea H. Wendl. (Reinhardtia p.p. EP.}. Palmaceae (iv. I.) C\ . Am. Malouetia A. DC. Apocynaceae (n. 2). 20 S. Am., Afr. Malperia Wats. Compositae (2). i Mexico. Malpighla Plum, ex L. Malpighiaceae (n). 25 trop. Am., W.I. Not climbers. Some have stinging hairs ; some have cleist. fls. 406 MALPIGHIACEAE Malpighiaceae (EP., BH.}. Dicots. (Archichl. Geraniales,). 55 gen., 650 sp., trop., esp. S. Am. Shrubs or small trees, usu. climbing, forming a marked feature among the trop. lianes. Stem-anatomy peculiar. L. usu. opp., entire, slip., frequently gland-dotted; pi. usu. covered with peculiar branched unicellular hairs. Infl. race- mose. Fl. § , obliquely •[• . K (5), imbr. , often with large glands at the base of (outside) the sepals; C 5, petals usually clawed, imbr.; A 5 + 5, obdiplost., often fewer, joined in a ring at the base; anthers opening intr. by longitudinal splits; G (3), obliquely placed in the fl., 3-loc. with axile plac. ; one ovule in each loc., pend., semi-anatr. , with ventral raphe. Fr. typically a schizocarp breaking into 3 meri- carps, but frequently one or more of the loc. abort. The mericarps are often winged, in some cases, e.g. Banisteria, like those of Acer. Seed exalbum. Classification and chief genera (after Niedenzu) : I. PYRAMIDO TORAE (torus pyramidal ; mericarps usually winged): Tetrapteris, Banisteria, Acridocarpus. II. PLANITORAE (torus flat or concave; mericarps not winged) : Malpighia, Bunchosia, Byrsonima. Malt, barley steeped to start germ., and then kiln-dried. Maltebrunia Kunth. Gramineae (6). 2 Madag., S. Afr. Malus Tourn. ex L. = Pyrus Tourn. p.p. (Ros.). Malva (Tourn.) L. (BH. excl. Callirhoe Nutt.). Malvaceae (2). 30 N. temp.; 3 in Brit. Fl. of the ordinary type of the fam., with oocpls. Two Brit. sp. M. sylvestris L. and M. rotiindifolia L. (large and small mallow) afford a contrast in floral mech., &c. Honey is secreted in little pockets in the recept., covered with hairs which exclude rain and very short-tongued insects. The large mallow is very protandr.; the sta. stand up at first in the middle of the fl., and afterwards bend outwards and downwards whilst the styles lengthen and occupy the original positions of the sta. The small mallow has much smaller fls., much less visited by insects; they go through stages similar to those described above, but at the end of the 9 stage the styles bend downwards, twist in among the anthers and pollinate themselves. The 1. in autumn may usu. be seen covered with brown spots caused by the fungus Pnccinia malvacearum (cf. Berberis). Malvaceae (EP.; B.H. incl. Bombacaceae). Dicots. (Archichl. Mai- vales). 35 gen., 700 sp., trop. and temp. Herbs, shrubs, or trees, with alt. stip. 1. Fls. sol. or in cpd. cymose infls. made up of cin- cinni, $ , reg., usu. 5-merous. Epicalyx often present ; probably an aggregation of bracteoles, but perhaps stipular like that of some Rosaceae (q.v.). K 5 or (5), valvate; C 5 conv., the petals usu. asymmetrical; A usu. oo , owing to branching of the inner whorl of sta. (the outer is usu. absent), all united below into a tube which is joined to the petals and at first sight makes the C appear gatno- petalous; the anthers are monothecous (i.e. each = half an anther), the pollen grains spiny. G (i — oo ) frequently (5), multi-loc., wilh axile placentae. In § I a division of the cpls. by horiz. transv. walls occurs, producing vertical rows of one-ovuled portions. Ovules j — oo in each cpl. , anatr. , usually ascending, sometimes pend. Malva- MANGEL WURZEL 407 viscus has a berry, the rest of the order dry fr., either caps, or schizocarps. Embryo usually curved, surrounded by endosp. The fls. are generally protandr. (see Malva and Goethea). Gossypium (cotton), Hibiscus, and others arc of economic value. Many are garden favourites. [BH. chars, incl. 1. often palmately lobed or cpd.; A 5 — oo , free or united, pollen smooth ; endosp. little or none.] Classification and chief genera (after Schumann) : A. Cpls. in vert. rows. 1. Malopeae: Malope, Kitaibelia. B. Cpls. in one plane. 2. Malveae (schizocarp; styles as many as cpls. ) : Abu- tilon, Lavatera, Althaea, Malva, Anoda. 3. Ureneae (schizocarp; styles twice as many as cpls.) : Urena, Goethea, Pavonia. 4. Hibisceae (capsule) : Hibiscus, Gossypium. Malvales. The 26th order (EP.) of Archichlamydeae. The 6th cohort (BH.) of Polypetalae. Malvastrum A. Gray. Malvaceae (2). 85 Am. and S. Afr. Malvaviscus Dill, ex Adans. Malvaceae (3). 12 warm Am. Mammea L. Guttiferae (iv). i W. Ind., M. americana L., cult, for ed. fr., the Mammee or St Domingo apricot. The fls. are used in preparing a liqueur (eau de Creole). Mammee, Mannnea americana L. ; - sapote, Liicuina inatnmosa Gaertn. f. Mammilla, Cactaceae. Mammillaria Haw. (incl. Anhalonium Lem.). Cactaceae (in. 2). 400 trop. Am. Mostly small plants of very condensed form, often almost spherical in outline, with well-marked mammillae (see fam. ). There is a division of the growing point into two in the course of formation of the mammilla, and the part at the base of the mammilla (i.e. in the axil) gives rise to the fl. The G elongates after fert. so that the tip of the long red berry is raised clear of the thorns. In some sp. veg. repr. and dispersal occurs by the mammillae breaking off and blowing about or adhering. to animals, Fr. ed. Mammoth tree, Sequoia gigantea Lindl. et Gord. Man-of-the-earth (Am.), Ipomoea pandurata Meyer ; - orchis, Aceras anthropophora R. Br. Mana grass (Ceylon), Cymbopogon confertiflorns Stapf. Managa Aubl. Inc. sed. i Guiana. Manatu-grass (W.I.), Thalassia. Manchineel, Hippomane Mancinella L. Mancoa Wedd. Cruciferae (3). i Andes. Mandevilla Lindl. Apocynaceae (11. i). 45 trop. and subtrop. Am. Mandioca, Manihot Aipi Pohl. Mandragora (Tourn.) L. Solanaceae (2). 3 Medit. to Himal. (man- drake). For superstitions connected with this, cf. Treas. of Bat. Mandrake, Mandragora, (Am.) Podophylhun. Manettia Mutis. Rubiaceae (i. 5). 30 trop. Am. Manfreda Salisb. (Agave p.p.). Amaryllidaceae (n). 20 N. Am. Mangatoeira rubber, Hancornia speciosa Gomez. Mangel (mangold) wnrzel, Beta vulgaris L. var. 408 MANGIFERA Mangifera L. Anacardiaceae (i). 30 Inclomal. M. indica L. is the mango, everywhere cult, in the trop. for its fr. , a large drupe derived from the i cpl. of the fl. Numerous vars. occur. Manglietia Bhime (Magnolia p.p. EP.}. Magnoliaceae. 3 Indomal. Manglilla Juss. = Myrsine L. (BH.) = Rapanea Aubl. Mango, Mangifera indica L. Mangonia Schott. Araceae (vn). i Brazil. Mangosteen, Garcinia Mangostana L. Mangroves, the association of pi. of the muddy swamps at the mouths of rivers and elsewhere in the trop., over which the tide flows daily, leaving the mud hare at low water; chiefly %. Chief gen. Rhizo- phora ; others are Acanthus, Aegiceras, Avicennia, Bruguiera, Carapa, Ceriops, Conocarpus, Laguncularia, Kandelia, Lumnitzera, Scyphi- phora, Sonneratia, &c. Usu. much branched, with aerial roots, both flying buttress and pillar roots ; aerating roots rise from the mud in Avicennia, Bniguiera, Sonneratia, &c. Many show viviparous germination. Manicaria Gaertn. Palmaceae (iv. i). i trop. S. Am. Manicoba rubber Gaertn., Manihot Glaziovii Muell.-Arg. Manihot Tourn. ex Adans. Euphorbiaceae (A. n. 4). 150 S. Am. to Mexico. Shrubs and herbs with monoec. fls. J7. Glaziovii Miill.- Arg. and other sp. show hud-protection well. The petiole of the young leaf curls upwards and inwards, so that the leaf is brought above the bud. M. utilissima Pohl is the bitter, M. Aipi Pohl (M. palmata Miill.-Arg. ) the sweet cassava or mandioc ; both are exten- sively cult, in the trop. for their large tuberous roots, which contain much starch, &c., and form a valuable food-stuff. The bitter cassava is the one usually cult.: its poisonous juice is squeezed out, and finally dissipated in the drying. The ground roots form mandioc or cassava meal, sometimes called Brazilian arrowroot. By a special mode of preparation, tapioca is prepared from the root. The poisonous juice, evaporated to a syrup and thus rendered harmless, forms an antiseptic, known as cassareep, used in preserving meat, &c. M. Glaziovii is the Ceara rubber ; rubber- is obtained by tapping the stem of the tree in the usual way. Several other sp. also yield rubber. Manila copal, A gat his Dammara Rich. ; - elemi, Canarium commune L. ; - hemp, Mnsa textilis Nee. Maniltoa SchefiF. Leguminosae (n. 2). i Colombia. Manioca, Manihot Aipi Pohl. Manisuris L. = Rottboellia L. f. (Gram.). Manisuris S\v. Gramineae (2). i trop. Manjack (W.I.), Cordia macrophylla L. Manna D. Don = Alhagi Tourn. (Legum.). Manna, Alhagi, Tamarix; - grass (Am.), Glycerin. Mannia Hook. f. Simarubaceae. i trop. W. Afr. Manniella Reichb. f. Orchidaceae (n. 2). i trop. Afr. Manniophyton Muell.-Arg. Euphorbiaceae (A. n. 2). i trop. VV. Afr. Manochlaenia B6rner=:Carex Dill. p.p. 6 in some. Megacaryon Boiss. Boraginaceae (iv. 5). i W. As. Megaclinium Lindl. Orchidaceae (ll. 16). 20 trop. and subtrop. Afr. Cult. 4i 6 MEGADENIA Megadenia Maxim. Cruciferae (2). i China. Megalachne Steud. Gramineae (10). i Juan Fernandez. Megalochlamys Lindau. Acanthaceae (iv. B). 2 Afr. Megalodonta Greene (Bidens p.p.). Compositae (5). 3 N. Am. Megalopus K. Schum. Rubiaceae (n. 5). i Cameroons. Megalostylis Sp. Moore. Euphorbiaceae (A. II. 2). i Upper Amazon. Megaphyllaea Hemsl. Meliaceae (in), i Perak. Megapterium Spach (Oenothera p.p. BH.}. Onagraceae (2). 3 Missouri valley. Cult. orn. fl. MegarrMza Torr. et Gray— Echinocystis Torr. p.p. (Cucurb.). Megaskepasma Lindau. Acanthaceae (iv. B). i Venezuela. Megastachya Beauv. = Eragrostis Beauv. p.p. (Gram.). Megastigma Hook. f. Rutaceae (i). 2 Mexico, Guatemala. Megastyles Schlechter. Orchidaceae (n. 2). 7 Indomal. Megistostegium Hochr. (Macrocalyx Cost, et Poiss.). Malvaceae (4). i Madag. Megistostigma Hook. f. Euphorbiaceae (A. n. 2). i Malacca. Meibomia Heist, ex Adans. (Des»iodnt»i^.p.}. Legumin. (111.7). 6 Am. Meiocarpidium Engl. et DiehiUvariap.p.). Anonaceae (i). 2 W. Afr. Meiogyne Miq. (Unona p.p. BH.}. Anonaceae (4). 2 trop. As. Meionectes R. Br. Haloragidaceae. i S. Austr., Tasm. Meiracyllium Reichb. f. Orchidaceae (n. 6). 2 Mexico. Mela-, melano- (Gr. pref.), black. Meladendron Molina. Solanaceae (inc. sed.). i Chili. Melalema Hook. f. Compositae (8). i Patagonia. Melaleuca L. Myrtaceae (n. i). 100 Austr. i to India. The 1. of M. Leucadendron L. (Austr., Indomal.) yield Cajeputoil. Sta. in antepet. bundles. Several yield oil ; timber useful. Melampodium L. Compositae (5). 25 Am. Melampyrum (Tourn.) L. Scrophulariaceae (in. 3). 35 N. temp.; 4 in Brit, (cow-wheat). Semi-parasites (see fain.). The fl. has a loose-pollen mechanism ; the 4 anthers lie close together and form a pollen-box ; the filaments of the sta. are covered with sharp teeth. Melananthos Pohl. Inc. sed. Nomen. Melananthus Walp. Solanaceae (5). 2 Brazil, C. Am. Melancium Naud. Cucurbitaceae (2). i E. and S. Brazil. Melandrium Roehl (Lychnis p.p. BH.}. Caryophyllaceae (n. i). 60 *., S. Afr., S. Am. M. ntbrum Garck. (Lychnis dioica L.) isdioec. and the ? pi. is stouter and coarser in growth than the 3 . Melanobatus Greene (Riibus p.p.). Rosaceae (in. 2). 7 N. Am. Melanocenchris Nees. Gramineae (i i). 3 trop. As. and Afr. Melanochyla Hook. f. Anacardiaceae (4). 4 Malaya. Melanococca Blume. Rutaceae (inc. sed.). i New Guinea. Melanodendron DC. Compositae (3). i St Helena. Tree. Melanodiscus Radlk. Sapindaceae (i). 2 trop. Afr. Melanophylla Baker. Cornaceae. 3 Madag. Melanopsidium Cels. (BiJliottia p.p. BH.}. Rubi. (i. 8). i Rio de Janeiro. Melanorrhoea Wall. Anacardiaceae (i). 6 Malaya. M. nsitata Wall. (Theetsee) yields a valuable black varnish, obtained by tapping the stem ; the sap turns black on exposure to air. MELEGUETA PEPPER 417 Melanosciadium Boiss. Umbelliferae (in. 5). i China. Melanoselinum Hoffm. (Thapsia p.p. BH.). Umbelliferae (in. 7). 2 Madeira. Melanoseris Decne. = Lactuca Tourn. (Comp.). Melanotis Neck. Inc. sed. Nomen. Melanoxylon Schott. Leguminosae (n. 8). i S-E. Brazil, M. Brauna Schott (Brauna). Timber useful. Melanthera Rohr. Compositae (5). 20 Afr., Madag., Am. Melanthesa Blume- Breynia Forst. (Euph.). MelantMum Clayton ex L. Liliaceae (i). 3 N. Am. Melasma Berg. Scrophulariaceae (in. 2). 25 Am., Afr., Madag. Melasphaerula Ker-Gawl. Iridaceae (in), i Cape Colony. Melastoma Burm. ex L. Melastomaceae (i). 40 As., trop. and E. Melastomaceae (EP., BH.}. Dicotyledons (Archichl. Myrtiflorae ; Myrtales, />//.)• I5°gen-i 2500 sp., trop. and subtrop. A very natural family, usu. easy to recognize, even when not in fl., by the peculiar leaf-veining, &c. The habit differs much, as they exist under varied conditions. Some are herbs, others trees or shrubs ; some climb, usu. by roots ; some are epiph., water or marsh pi. L. nearly always decussate (stem often 4-angled), but one 1. generally much larger than the other; the lesser in some cases withers away as it grows older, and drops off. The veins of the 1., which is usu. simple and entire or nearly so, diverge fiom the base and converge again at the apex, as in many monocot. L, so that there is no true midrib. Many are myrmecophilous, e.g. Tococa, Maieta, &.c. (q.v-). Infl. cymose, but in great variety. Fl. usu. very char., easily recognized by the curious appendages of the anthers. Recept. (' calyx- tube ') tubular or bell-shaped, commonly ± united with the ovary, often brightly coloured. K 4 or 5 ; C 4 or 5, perig. ; P. usu. reg. , but irregu- larity often occurs in the A. Sta. usu. twice as many as petals, standing (when mature) in one whorl, bent down in bud so that the anthers come between the ovary and the recept. ; the anther-loculi open by a common apical pore. The connective is developed in various ways and usu. provided with curious appendages, frequently of sickle-like form, giving a char, aspect to the fl. G. sup. or inf., usu. 4 — 5-loc., with a simple style and stigma: ovules co , anatr., on axile plac. Fr. a berry or loculic. caps. Seed exalbum. ; one cot. larger than the other. The M. are of little economic importance; a few yield colouring matters. Classification and chief genera (after Krasser) : A. Fruit many-seeded. Embryo very small. I. MELASTOMATOIDEAE (ovules on slightly projecting placentae in inner angle of loc.) : Tibouchina, Centra- denia, Melastoma, Monochaetum, Medinilla, Leandra, Miconia, Tococa, Maieta. II. ASTRONIOIDEAE (ovules on a placenta at base or on wall of loc.) : Kibessia. B. Fruit i — 5-seeded. Embryo large. III. MEMECYLOIDEAE: Memecylon. Meleagrinex Arruda. Inc. sed. i Brazil. Melegueta pepper, A mo mum melegueta Roscoe. w. 27 4i8 MEL-GRASS Mel-grass, Ammophila arundinacea Host. Melhania Forsk. Sterculiaceae. 25 Afr., As., Austr. Meliadelpha Radlk. Meliaceae (inc. sed.). 2 Polynesia. Melia L. (BH. incl. Azadirachta A. Juss. q.v. for M. Azadirachta L., the nim). Meliaceae (ill). 12 palaeotrop. Some useful for timber. M. Azedarach L., the beadtree, cult, for its fls. Meliaceae (EP., BH.}. Dicots. (Archichl. Geraniales). 40 gen., 600 sp., trop and subtrop. Mostly trees and shrubs, with alt. exstip. pinnate L, and cymose panicles of 5 reg. fls. K (4 — 5) or 4 — 5; €4 — 5; A 8 — 10 usu. united below into a tube, or sometimes united all their length, in which case the anthers are sessile on the tube; disc present or not ; G 2 — 5-loc. or rarely i-loc. or more than 5-loc. ; style present or not; ov. in each loc. i, 2, or more, usu. pend. and anatr., with ventral raphe. Capsule, berry, or drupe; seeds often winged, with endosp. Many, e.g. Swietenia (mahogany), Cedrela, &c., yield valu- able timber; the seeds of several are used as sources of oils; others have ed. fr. Classification and chief genera (after Engler) : I. CEDRELOIDEAE (sta. free) : Cedrela, Pteroxylon. II. SWIETEN1OIDEAE (sta. in a tube ; seeds winged) : Swie- tenia, Khaya. III. MELIOIDEAE (sta. in a tube; seeds not winged) : Carapa, Melia, Azadirachta, Trichilia, Guarea. Melianthaceae (EP.\ Sapindaceae p.p. BH.). Dicots. (Archichl. Sa- pindales). 3 gen., 17 sp., Afr. (trop. and S.). Trees and shrubs with alt. usu. stip. 1., and racemes of $ , median-zygom. fls., whose stalks twist through 180° at the time of flowering. K 5 or (5), sometimes 4 by union of two sepals; C 4 or 5; disc extra-staminal ; A 5 or 4 or 10, free or united at base; G (4 — 5), 4 — 5-loc. with one basal or many axile ov. in each loc. ; ovules erect or pend., anatr. with ventral or dorsal raphe according as they are erect or pendulous respectively. Capsule; seed sometimes with aril; endosp. fleshy or horny. Chief genus: Melianthus. Melianthus L. Melianthaceae. 5 S. Afr. Fls. very rich in honey. Melica L. Gramineae (10). 40 temp., exc. Austr.; 2 in Brit, (melic- grass). Melic-grass, Melica. Melicnrus R. Br. (Styphelia p.p. EP.). Epacridaceae (3). 2 Austr. Melicocca L. Sapindaceae (i). 2 trop. Am., W.I. M. bijuga L. (W. I. ) for timber and fr. Melicope Forst. Rutaceae (i). 12 trop. As., warm Austr. Melicytus Forst. Violaceae. 4 New Zealand, Norfolk I. Melientha Pierre. Opiliaceae. i Cambodia. Melilot, Melilotus. Melilotus Tourn. ex Hall. Leguminosae (ill. 4). 20 temp, and sub- trop. |^. 3 in Brit, (melilot). The fls. contain much honey, and are visited by bees. Melinia Decne. in DC. Asclepiadaceae (n. i). 6 E. temp. S. Am. Melinis Beauv. Gramineae (4). 5 trop. S. Am., Afr., Madag. Fodder. MelioscMnzia K. Schum. (Chisocheton p.p. EP.). Melia. (in), i N.G. Meliosma Blume. Sabiaceae. 55 trop. and subtrop. As. and Am. MENISPERMACEAE 419 Melissa Tourn. ex L. Labiatae (vi). 4 Eur. , W. As. M. officinalis L. (balm) cult. Melittacanthus Sp. Moore. Acanthaceae (iv. B). r Madagascar. Melitella Sommier. Compositae (13). i Malta. Melittis L. Labiatae (vi). i Eur. (incl. Brit.), M. Melissophylluin L., the bastard-balm. Mellera Sp. Moore. Acanthaceae (iv. A). 3 trop. E. Afr. Melleus (Lat.), of honey colour or taste. Mellichampia A. Gray. Asclepiadaceae (n. i). r Mexico. Melliniella Harms. Leguminosae (in. 7). i E. trop. Afr. Mellissia Hook. f. Solanaceae (2). i St Helena. Melloa Bur. Bignoniaceae (i)- 2 Brazil, Venezuela. Melo (Tourn.) L. — Cucumis Tourn. (Cucurb.). Melocactus (Tourn.) Link et Otto. Cactaceae (in. i). 30 W.I., S. Am. Ribbed plants, like Cereus. Fls. produced at the top. Melocalamus Benth. Gramineae (13). i Burma. Melocaima Trin. Gramineae (13). i Indomal. Exalbum. seed ed. MelocMa Dill, ex L. Sterculiaceae. 60 trop. Melochia Rottb. Inc. sed. i Guiana. Melodinus Forst. Apocynaceae (i. i). 25 Indomal., Polynesia. Melodorum Hook. f. et Thorns. Anonaceae (4). 30 palaeotrop. Melolobium Eckl. et Zeyh. Leguminosae (in. 3). 12 S. Afr. Melon, Cucumis Melo L. ; -cactus, Melocactus ; -thick (W.I.), Melo- cactus; -, water, Citnillus vulgaris Schrad.; -, white gourd, Benin- casa cerifera Savi. Melosperma Benth. Scrophulariaceae (n. 6). i Chili. Melothria L. Cucurbitaceae (2). 60 trop. and subtrop. Membranous (L), thin, dry, not green, flexible. Memecylanthus Gilg et Schlechter. Caprifoliaceae. i New Caled. Memecylon L. Melastomaceae (in). 120 palaeotrop. Memora Miers (Adenocalymma p.p. BH.). Bignoniaceae (i). 20 S. Am. Memorialis Buch.-Ham. (Poitzolzia p.p. BH.}. Urtic. (3). 13 Indo- mal. Menabea Baill. Asclepiadaceae (11. 2). i Madagascar. Menais Loefl. Inc. sed. i S. Am. Menaphronocalyx Pohl. Inc. sed. Nomen. Mendoncia Veil. Acanthaceae (11). 24 trop. Am. Mendozia Ruiz et Pav. = Mendoncia Veil. (Acanth.). Menepetalum Loes. Celestraceae. 5 New Caledonia. Menestoria DC. Rubiaceae. Genus delendum ; farrago. Meniscium Schreb. = Dryopteris Adans. (Polypod.). Meniscoid, watch-glass-shaped. Menispermaceae (EP., BH.). Dicots. (Archichl. Ranales). 63 gen., with 360 sp. , trop. and warm temp. Mostly climbing shrubs with alt. simple 1., in whose axils are usu. serial buds. The stem-anatomy is of interest. Fls. in axillary racemes (exc. Cissampelos, &c.), uni- sexual, usu. dieec. Formula usu. K3 + 3, C 3 + 3, A 3 + 3, 63, but many exceptions. K and A often > 6 ; sometimes only i cpl. Ovules i in each cpl., ventral, pend., semi-anatr. Achene; seed album. The classification of the genera is largely based on the structure of 27—2 420 MENISPERMA CEAE the seed. A few are or have been medicinal, on account of the bitter principle in the roots. Chief genera: Menispermum, Cocculus, Cissampelos, Jateorhiza, Tinospora, Anamirta. Menispermum (Tourn. ) L. Menispermaceae. 3 temp. E. As., Atl. N. Am. (moon-seed). Menkea Lehm. Cruciferae (3). 2 Austr. Menodora Humb. et Bonpl. Oleaceae. 15 warm Am., Afr. Menonvillea R. Br. Cruciferae (i). 4 Chili, Peru. Mentha (Tourn.) L. Labiatae (vi). 15 I*. 6 in Brit, (mint), incl. M. piperita L. (peppermint), M. Pidegium L. (penny-royal). From the former an oil is obtained by distillation and used in medicine, &c. M. viridis L. is cultivated as a pot-herb for flavouring. Menthol, a substance extracted from oil of peppermint, &c. Mentum, a chin, cf. Orchidaceae. Mentzelia Plum, ex L. (BH. incl. Eucnide Zucc. ). Loasaceae. 55 warm Am. No stinging hairs. In some the outer sta. are sterile. Menyanthes (Tourn.) L. Gentianaceae (n). i N. temp. (incl. Brit.), M. trifoliata L., the buck- or bog-bean. A bog plant with creeping rhiz. and alt. 1. Fls. dimorphic heterostyled (ff. Primula). The rhizome has bitter tonic properties. Menziesia Sm. Ericaceae (i. 2). 7 N. temp. [M. polifolia Juss.= Daboecia polifolia.] Meoschium Beauv. = Ischaemum L. p.p. (Gram.). Mephitidia Reinw. ex Blume = Lasianthus Jack. (Rubi.). Merathrepta Rafin. (Danlhonia p.p.). Gramineae (9). 9 N. Am. Merciera A. DC. Campanulaceae (i). 4 S. Afr. Merckia Fisch. (Arenaria p.p. BH.}. Caryophyll. (i. i). i N.E. As., N.W. Am. Mercurialis (Tourn.) L. Euphorbiaceae (A. n. 2). 7 Medit., Eur., E. As. 2 in Brit., M. perennis L. and M. annua L. (mercury). Uioec. anemoph. fls. Veg. repr. by rhizomes. C absent. Cpls. (2). Mercury, Mercurialis. Merendera Ram. Liliaceae (i). 10 Medit., Abyssinia. Meretricia Neraud. Rubiaceae. Nomen. Meriandra Benth. Labiatae (vi). 2 Himalaya, Abyssinia. Meriania Sw. Melastomaceae (i). 30 W.I., trop. Am. Mericarp, the one-seeded portion of a schizocarp, Umbelliferae. Mericarpaea Boiss. Rubiaceae (n. n). i W. As. Meringurus Murbeck. Gramineae (12). i Tunis. Merinthopodium Donnell Smith. Solanaceae (4). C. Am. Merinthosorus Copeland (AcrostzcAump.p.). Polypodiaceae. i Phil. Is. MeriolixRann.(Cte« i fl. The fl. MIXED 427 opens in the evening and is protog. (in M.Jalapa L. and other sp.), with ultimate autogamy on withering. The invol. often forms a parachute on the fr. The tuberous roots of M. Jalapa L. (false jalap, four-o'clock, marvel of Peru) were formerly used as jalap. Mirasolia Sch.-Bip. (Titlwnia p.p. EP., Gymnolomia p.p. BH.}. Compositae (5). i Mexico, C. Am. Mirbelia Sm. Leguminosae (in. 2). 16 Austr. Mirtana Pierre (Anamirta p-p.)- Menispermaceae. i Cochinchina. Misanteca (Miscanteca} Cham, et Schlechtd. Lauraceae (n). 4 trop. Am., W.I. Miscanthus Anderss. Gramineae (2). 10 S. and E. As. Miscellaneous useful products may be roughly grouped into such things as beads (Abrus), cork (Quercus), teasels (Dipsactis), vegetable ivory Phytelephas], weights (Abnis, &c.). Cf. these heads. Miscnobulbum Schlechter (Tainia p.p.). Orchidaceae (n. 9). 5 N.G. Mischocarpus Blume (Ratonia p.p. BH.). Sapindaceae (l). 12 Indomal. Mischocodon Radlk. Sapindaceae (i). i New Guinea. Mischodon Thw. Euphorbiaceae (A. i. i). i Ceylon, S. India. Mischophloeus Scheff. Palmaceae (iv. i). i Ternate. Miscolobium Vog. = Dalbergia L. (Legum.). Missiessya Gaudich. = Leucosyke Zoll. (Uitic.). Mission grass, Stetwtaphrnm. Mistletoe, Viscnm, esp. V. album L., Loranthaceae, (Am.) Phoradendron. Mistus, mixtus (Lat.), cross-bred between forms of a sp. Mitchella L. Rubiaceae (n. 7). 2 N. Am. (M. re fens L.) and Japan. Dimorphic heterostyled. The fls. are in pairs with united ovaries. Occasionally K and C also fuse and give a double ovary surmounted by a lo-lobed K and C (cf. Lonicera). Mitella Tourn. ex L. Saxifragaceae (i). 10 N. Am., Japan. The inconspic. greenish fls. stand in unilateral racemes. Mitellastra Howell (Mitella p.p.). Saxifragaceae (i). N.W. N. Am. Mitolepis Balf. f. Asclepiadaceae (i). i Socotra. Mitophyllum Greene (Streptanthus p.p.). Cruciferae (i). i Calif. Mitostemma Mast. Passifloraceae. 2 Brazil, Guiana. Mitostigma Decne. in DC. Asclepiadaceae (n. i). 6 S. Am. Mitozus Miers (Echttes R. Br. p.p.). Apocynaceae (n. i). 20 S. Am. Mitracarpum Zucc. Rubiaceae (n. 10). 15 S. Am., Afr. Mitragyna Korth. Rubiaceae (i. 6). 12 trop. As., Afr. Mitranthes Berg (Calyptranthes BH.}. Myrtaceae (i). 4 trop. Am., W.I. Mitraria Cav. Gesneriaceae (i). i Chili. Mitrasacme Labill. Loganiaceae. 28 Austr., N.Z., trop. As. Mitrastemma (Mitrastcinon} Makino. Rafflesiaceae. i Japan. Mitratheca K. Schum. Rubiaceae (i. 2). i trop. Afr. Mitre-flower, Mitraria; -wort (Am.), Mid cola. Mitreola L. Loganiaceae. 4 Am., Indomal., Austr. Mitrephora Hook. f. et Thorns. Anonaceae (2). 25 trop. As. Mitriostigma Hochst. (Randia p.p. EP.}. Rubiaceae (i. 8). 3 trop. and S. Afr. Mixed (infl.), partly racemose, partly cymose, Aesculus, Betulaceae, Ceratostigma, Labiatae, Marina, Statice, Verbascutn. 428 MIYOSHIA Miyoshia Makino (Protolirion Ridl.)- Liliaceae (i). i Japan. Mnassea Veil. Inc. sed. i Brazil. Mnemion Spach = Viola Tourn. (Viol.). Mniochloa Chase (Digifartap.p.). Gramineae (5). i Cuba. Mniodes A. Gray. Compositae (4). 2 Peru. Mniopsis Mart. Podostemaceae. 4 Brazil. Mniothamnus Niedenzu (Berzelia p.p.). Bruniaceae. i S. Afr. Moa, Bassia latifolia Roxb. Moccasin flower (Am.), Cypripediiim. Mocinna Cerv. ex La Llave = Carica L. (Caric.). Mocker nut (Am.), Carya tomentosa Nutt. Mock-orange (Am.), Philadelphus. Mocquerysia Hua. Flacourtiaceae (10). i trop. W. Afr. Modecca Lam. (Adenia Forsk. EP.). Passifloraceae. 50 palaeotrop. Modiola Moench. Malvaceae (2). i Am., S. Afr. (?). Modiolastrum K. Schum. (Modiola Moench). Malvac. (2). i S. Am. Moehringia L. (Arenaria p.p. BH.). Caryophyliaceae (i. i). 20 N. temp. Moenchia Ehrh. (Cerastiin/i p.p. BH.}. Caryophyliaceae (i. i). 5 Eur., Medit. Moerenhoutia Blume. Orchidaceae (n. 2). 2 Polynesia. Moghania Jaume St Hil. (Flemingia Roxb.). Legum. (in. 10). 20 palaeotrop. Mogiphanes Mart. (Alternanthera p.p. EP.). Amarantaceae (3). 12 trop. Am. Mogorium Tuss.=Jasminum Tourn. (Oleac.). Mohadenium Pax. Euphorbiaceae (A. n. 8). i E. Afr. Mohavea A. Gray. Scrophulariaceae (n. 3). 2 S.W. U.S. Mohlana Mart. Phytolaccaceae. 2 S. Am., trop. Afr., Madag. Mohria Sw. Schizaeaceae. 3 trop. and S. Afr. Sporangia on under side of ordinary L, margins turned back over them (cf. Pteris). Moldenhauera Schrad. Leguminosae (u. 7). 3 Brazil, Venezuela. Molina Ruiz et Pa\r. = Baccharis L. (Comp.). Molinaea Comin. ex Juss. (Cupania p.p. BH.). Sapindaceae (i). 8 Madagascar, Mascarene Is. Molineria Parl. (Aira p.p. BH.). Gramineae (9). 3 W. and S. Eur., As. Min. Molineriella Rouy (Aira p.p.). Gramineae (9). i Medit. Molinia Schrank. Gramineae (10). i Eur. (incl. Brit.), As., M. caerulea Moench, char, of wet grass moors. Mollera O. Hoffm. Compositae (4). 2 trop. Afr. Mollia Mart. Tiliaceae. 7 trop. S. Am. Mollinedia Ruiz et Pav. Monimiaceae. 70 trop. Am. Mollis (Lat.), soft, pubescent. Mollugo L. Aizoaceae (i). 15 trop., and N. Am. Molopanthera Turcz. Rubiaceae (i. 5). i E. Brazil. Molospermum Koch. Umbelliferae (in. 2). i W. Medit. Moltkia Lehm. Boraginaceae (iv. 4). 6 Himalaya to Medit. Moluccella L. Labiatae (vi). 2 Medit. Moly, Allintn Moly L. Mombin, Spondias. MONIZIA 429 Momisia F. G. Dietr. = Celtis Tourn. p.p. (Ulm.). Momordica (Tourn.) L. Cucurbitaceae (3). 25 palaeotrop. Monachanthus Lindl.=Catasetum Rich. (Orchid.). Monachochlamys Baker. Acanthaceae (n). i Madag. Monachosorum Kunze. Polypodiaceae. 2 E. warm As. Monachyron Parl. ( Tricholaena p.p.). Gramineae (5). i Cape Verde Is. Monactinocephalus Klatt (Inula p.p. EP.). Compositae (4). i S. Afr. Monactis H. B. et K. Compositae (5). 2 trop. S. Am. Monadelphanthus Karet (Capirona p.p. EP.}. Rubi. (i. 4). i Co- lombia. Monadelphous (A), concrescent in one bundle. Monadenia Lindl. (Disa p.p. BH.}. Orchidaceae (n. i). 12 Cape Col. Monadeniuin Pax. Euphorbiaceae (A. n. 8). 5 trop. Afr. Monandrous, with one sta. Monanthes Haw. Crassulaceae. 3 Morocco, Canaries. Cult. orn. pi. Monanthochloe Engelm. Gramineae (10). i S. U.S. and Argentina. Monanthotaxis Baill. Anonaceae (i). i Congo. Monarda L. Labiatae (vi). 20 N. Am. Sta. 2. Fl. protandrous, visited by bees (and humming-birds in the red sp.). The 1. of some are used medicinally in the form of tea (Oswego-tea). Monardella Benth. Labiatae (vi). 25 W. N.Am. Monarrhenus Cass. Compositae (4). 3 Madagascar, Mascarene Is. Monarthrocarpus Merrill (Desmodium p.p.). Leguminosae (in. 7). i Phil. Is. Monechma Hochst. (Justida p.p.). Acanthaceae (iv. B). 50 trop. Afr. Monelasum Van Tiegh. = Ouratea Aubl. (Ochn.). Monelytrum Hack. Gramineae (3). i S.W. Afr. Monenteles Labill. = Pterocaulon Ell. (Comp.). Monerma Beauv. (Psilurus p.p. BH.). Gramineae (12). 3 warm |%. Moneses Salisb. (Pyrola p.p.). Pyrolaceae. i boreal and arctic. Money wort, Lysimachia Nitinmularia L. ; Cornish. - -, Sibthorpia enropaca L. Moniliform, like a row of beads. Monimia Thou. Monimiaceae. 3 Madag. , Mascarenes. Monimiaceae (EP., BH.). Dicots. (Archichl. Ranales ; Micrembryae BH.). 30 gen., 200 sp., chiefly S. trop., and esp. in the ' oceanic ' floral regions (Madag., Austr., Polynes.). Shrubs and trees, with leathery evergr. 1., often resiniferous with aromatic scent, usu. opp., exstip. Fls. sol. or in cymes., perigynous, commonly unisexual, reg.; often the two sexes differ in the hollowing of the axis. Frequently the bud opens by throwing off the outer ends of the P-leaves as a sort of lid. P 4 — oo , simple, or o ; A oo or few, the anthers intr. or extr. , opening by slits or valves ; G. usu. oo , sometimes few or i, each with I usu. basal erect anatr. ovule. Fr. of achenes, often ± enclosed in or borne on a fleshy recept. Embryo straight, in copious endosp. The fam. forms a connecting link between Lauraceae and the other Ranales, being closely allied on one side to L. , on the other to Caly- canthaceae. Chief genera: Hedycaria, Peumus, Ta;i,bourissa, Laurelia. Monixus Finet (Angraecum p.p.). Orchid, (n. 20). 8 trop. Afr., Madag. Monizia Lowe = Thapsia L. (.S^y.^Melanoselinum Hoffm. 430 MONKEY APPLE Monkey apple (W.I.), Anona palustris L.; -bread, Adansonia digi- tata L.; -flower, Mimulus luteus L.; -pot, Lecythis ; -puzzle, Arau- caria imbricata Cav. Monkshood, Aconitntn. Monnieria L. Butaceae (i). 2 trop. S. Am. Monnina Ruiz et Pav. Polygalaceae. 75 Mexico to Chili. One of the two cpls. is usu. rudimentary. Fr. indehiscent. Mono- (Gr. pref.), one; -carpellary, of i cpl.; -carpic, once-fruiting, Agave, Corypha, &c.; -chasial cyme, one in which each successive branch bears one branch upon itself, and of four types, bostryx, cin- cinnus, drepaniiiin, rhipidtuui, q.v.; -Chlamydeous, with one whorl of P; -clinous, hermaphrodite; -ecious, with i and ? fls. on the same pi. ; -graph, a systematic account of a group ; -petalous, sympetalous ; -podial branching, -podium, where the same growing point continues in a straight line from year to year, and forms branches in regular succession, Coniferae, Paris, Pot/ws; -spermous, one-seeded ; -sym- metrical, zygomorphic ; -thecal, Malvaceae ; -tocous, fruiting once only; -typic (genus), with one species. Monocarpia Miq. Anonaceae (i). 2 trop. As. Monocera Jack^Elaeocarpus Burm. p.p. (Elaeocarp.). Monocnaete Doell. Gramineae (n). i Brazil. Monocnaetum Naud. Melastomaceae (i). 30 W. trop. Am. A di- morphous. The style, at first bent clown., moves slowly up till horiz. Monochasma Maxim ex Franch. et Sav. Scroph. (in. 3). i Japan, China. Monochilus Fisch. et Mey. Verbenaceae (i). i Brazil. Monochilus Wall, ex Lindl. =Zeuxine Lindl. (Orchid.). Monocnlamydeae (BH.). One of the chief divisions of Dicots. Monochoria C. Presl. Pontecleriaceae. 4 E. Afr. to Austr. Monococcus F. Muell. Phytolaccaceae. i Austr., New Caled. Monocosmia Fenzl. Portulacaceae. i Chili. Monocostus K. Schum. Zingiberaceae (n). i Peru. Monocotyledones. One of the two great divisions of Angiospermae. Their classification is less difficult than that of the Dicotyledons, and a comparison should be made of the ways in which it is done in the various systems. On the origin of M., one of the great unsolved problems in phylogeny, cf. Seward, Geological History, in Ann. of Bot. X, 1896, p. 205 ; Miss Sargant, Theory of origin of M., in do. XVII, 1903, p. i, and esp. review by Bancroft, in New Pkytol. 13, 1914, p. 285. Monodora Dun. Anonaceae (5). 10 trop. Afr., Madag. Berry with woody epicarp. Seeds of M. Myristica Dun. sometimes used as nutmegs. Monogramma Schk. Polypodiaceae. 15 trop. and subtrop. Monolena Triana. Melastomaceae (i). 4 trop. S. and C. Am. Cult, orn. fl. Monolepis Schrad. Chenopodiaceae (A). 3 N. Am. Monolopia DC. Compositae (6). 5 Calif. Monomeria Lindl. Orchidaceae (n. t6). 2 Nepal, Burma. Monoon Miq.= Polyalthia Blume p.p. (Anon.). MONTEZUMA 43 1 Monopetalanthus Harms. Leguminosae (n. 2). i trop. Afr. Monophrynium K. Schum. (Phrynium p.p.). Marantaceae. 2 Phil. Is. Monophyllaea R. Br. Gesneriaceae (i). 7 Malay Archipelago. Monophyllanthe K. Schum. Marantaceae. i French Guiana. Monoporandra Thw. Dipterocarpaceae. 2 Ceylon. Monoporus A. DC. (Ardisia p.p.). Myrsinaceae (n). 6 Madagascar. Monopsis Salisb. (Lobelia p.p. BH.}. Campanul. (ill). 9 S. Afr., Abyss. Monopteryx Spruce. Leguminosae (in. i). 2 Amazon valley. Monoptilon Torr. et Gray. Compositae (3). i California, Utah. Monopyle Moritz ex B. et H. f. Gesneriaceae (n). 6 C. Am. to Peru. Monopyrena Spegazzini. Verbenaceae (i). i Patagonia. Monosepalum Schlechter (B-ulbophyllum p.p.). Orchidaceae (n. 16). 3 New Guinea. Monosis DC.=Vernonia Schreb. p.p. (Comp.). Monostachya Merrill. Gramineae (10). i Luzon. Monosternma Turcz. (Sarcostemma p.p.). Asclepiad. (n. i). i S. Afr. Monotagma K. Schum. (Ischnosiphon p.p.). Marant. 8 trop. and S. Am. Monotaxis Brongn. Euphorbiaceae (B. n). 7 Austr. Monotes A. DC. Dipterocarpaceae. 6 trop. Afr. Monothecium Hochst. Acanthaceae (iv. B). 3 palaeotmp. Monotoca R. Br. Epacridaceae (3). 6 Austr. Monotropa L. (incl. Hypopitys Dill.). Pyrolaceae. 3 N. temp. M. Hypopitys Walt, (yellow bird's-nest), in fir, birch and beech woods in Brit., a yellowish saprophyte with scaly 1. and a short term, raceme of fls. Below the soil is found a very much branched root system, the roots being covered with a superficial mycorhiza by whose aid absorption takes place. Buds are formed adv. upon the roots and lengthen into the flowering shoots. Monotropeae (Bff.; Pyrolaceae p.p. EP.}. Dicots. (Gamopet. Ericales). A fam. containing the saprophytic Pyrolaceae only. Monotropsis Schwein. ex Ell. = Sch weinitzia Ell. (Pyrol.). Monoxalis Small (Oxalis p.p.). Oxalidaceae. i N. Am. Monsonia L. Geraniaceae. 12 Afr., As. Monstera Adans. Araceae (n). 27 trop. Am. Climbing shrubs with pinnatifid 1., full of round holes. When very young the 1. is entire ; then the tissue between the veins ceases to grow rapidly, becomes dry and tears away, thus leaving holes between the ribs ; at the edge the marginal part usually breaks, and thus the outermost hole gives rise to a notch in the 1., which becomes pinnated. Beginning as a climber the pi. usu. ends as an epiph. with aerial roots to the soil. Fls. . § The fr. of M. deliciosa Liebm. is ed. Monstrosities, marked aberrant variations suddenly appearing. Montagnaea DC. =Montanoa Cerv. (££f.) = Eriocoma. H. B. et K. (Comp.). Montanoa Cerv. Compositae (5). 25 Mexico to Colombia. Cult. orn. fl. Montanus (Lat), mountain. Montbretia DC. =Tritonia Ker-Gawl p.p. (Irid.). Monterey cypress, Cupressus macrocarpa Hartn. Montezuma (Moc. et Sesse ex) DC. Bombacaceae. i Mexico. 432 MONTI A Montia Mich, ex L. Portulacaceae. i cosmop. M.fontana L. (blinks), an annual herb, usu. in wet places, with small cymes of fls. In bad weather or when submerged they become cleistogamic. The stalk moves like that of Claytonia, and the fr. explodes in the same way. Eaten as salad. [Am. authors incl. Claytonia p.p.] Montinia Thunb. Saxifragaceae (v). i S. Afr. Montiopsis O. Ktze. Portulacaceae. i Bolivia. Montolivaea Reichb. f. (Habenaria p.p.). Orchid, (n. i). i Abyssinia. Montrichardia Crueg. Araceae (iv). i trop. S. Am., W.I. Montrouziera Planch, ex Planch, et Triana. Guttif. (v). 3 New Caled. Monttea C. Gay. Scrophulariaceae (n. 6). 3 Chili. Moonia Arn. (Chrysogonum p.p. />'//.). Compositae (5). 5 Indomal. Moonseed, Mentspermum ; -wort, Botrychntm. Moorea Lemaire (Cortaderia p.p.). Gramineae (10). 58. Am. Moorea (Neomoorea] Rolfe. Orchidaceae (II. 13). i S. Am. Moquilea Aubl. Rosaceae (vi). 20 S. and C. Am. Some apet. Moquinia DC. Compositae (12). 10 S. Am. Dioecious shrubs. Mora Schomb. ex Benth. = Dimorphandra Schott (Legum.). Moraceae (EP.; Urticaceae p.p. BH.}. Dicots. (Archichl. Urticales). 55 gen., 800 sp., trop. and subtrop., a few temp. Most are trees or shrubs with stip. 1., and with latex. [See Ficus, Cecropia, Madura, Humulus.] Infl. cymose, usu. in the form of (pseudo-) racemes, spikes, umbels or heads (cf. Urticaceae, and paper there cited). Fls. unisexual. P usu. 4 or (4), persistent; A in ov. in each loc. Berry, caps., or schizocarp. Seed with straight embyro and mealy perisperm. Fls. rich in honey, and visited by bees and birds. Musa is an important economic genus. • Classification and genera: closely related to the other Scitamineae (joined by BH.), less closely to Liliiflorae and Orchids. I. MUSOIDEAE (1. alt., fl. collateral, post. P leaf free): Musa. II. STREL1T7.I01DEAE (1. in i ranks, fl. in cymes, K free) : Ravenala, Strelitzia, Heliconia. III. LOWIOIDEAE (1. in 2 ranks, fl. in panicles from rhiz., K tubular) : Lowia, Orchidantha. Musanga C. Sm. ex R. Br. Moraceae (in), i Congo. Muscadinia Small (Vitis p.p.). Vitaceae. 2 N. Am. Muscari Tourn. ex Mill. Liliaceae (v). 40 Medit., Eur., As. M, race- mosum Mill, (grape-hyacinth) in Brit. Collateral buds in axils. Upper fls. of the raceme neuter, giving extra conspicuousness to the infl. (cf. Centaurea Cyanus}. Muscaria Haw. =Saxifraga Tourn. p.p. (Saxifr.). Muschleria Sp. Moore. Compositae (i). i Angola. Museniopsis Coulter et Rose. Umbelliferae (in. 5). 10 Mex., W. U.S. Musenium Nutt. (Mitsineitm Rafin.). Umbell. (in. 4). 3 N. Am. Musgravea F. Muell. Proteaceae (n). i Queensland. Muskit, cf. Mesquite. Musk mallow, Alalva moschata L., Hibiscus ; - melon, L'ltfitmis Melo L. ; - orchis, Herininium Monorchis R. Br. ; - plant, Minnilns inoschatus Dougl. ; - thistle, Car duns nntans L. Musquash root (Am.), Cicitta inaculata L. Mussaenda Burm. ex L. Rubiaceae (i. 7). 35 palaeotrop. One sepal 438 MUSSAENDA is large, leafy, and brightly coloured, and helps to make the fl. con- spicuous (cf. Euphorbia, Salvia). Mussaendopsis Baill. Rubiaceae (i. 5). i Malaya. Mussatia Bur. (Bignonia p.p.). Bignoniaceae (i). i Guiana. MusscMa Dum. Campanulaceae (i. i). 2 Madeira. The capsule opens by many transv. slits between the ribs. Cult. orn. fl. Mustard, Brassica nigra Koch, Sinapis; hedge-, Sisymbrium offici- wa/^Scop.; treacle-, Ery simian cheiranthoid<:s L. ; -\xtt,Salvadora persica L. Mutabilis (Lat), changeable (in colour, &c.). Mutant, form arising by mutation; cf. de Vries, Mutation Theory. Muticous, blunt. Mutisia L. f. Compositae (12). 55 S. Am. Many climbers (a rare habit in C.) with ends of leaf-midribs prolonged into tendrils. All are shrubby with large heads of fls. Shows well chars, of § 12. Mutumocarpon Pohl. Inc. sed. Nomen. Myagrum (Tourn.) L. Cruciferae (2). i Medit., mid-Eur. Myall (Austr.), Acacia homalophylla A. Cunn. Myanthus Lindl. = Catasetum Rich. (Orchid.). Mycelium, Rafflesiaceae. Mycetia Reinw. (Adenosacme Bff.). Rubiaceae (1.7). 5 Indomal. MycorMza, a fungus whose hyphae replace root-hairs in absorption (cf. Saprophytes), Botrychium, Epipogiim, Neotfia, Orchidaceae. Myginda Jacq. Celastraceae. 15 trop. Am., W.I. Myodocarpus Brongn. et Gris. Araliaceae (2). 12 New Caledonia. Myonima Comm. ex Juss. Rubiaceae (n. 4). 5 Mauritius, Bourbon. Myopordon Boiss. Compositae (n). 2 Persia. Myoporaceae (EP., Bff.). Dicots. (Sympet. Tubiflorae; Lamiales Bff.}. 5 gen., 100 sp. chiefly Austr. and neighbouring Is. (i in each of following: — Sandvv. Is., E. As., Mauritius, Afr., W.I.). Most are trees or shrubs, with alt. or opp. entire exstip. 1. , often covered with woolly or glandular hairs, frequently very reduced in size. Fls. sol., or in cymose groups, axillary, 5, reg., or -|- . K(s), C (5), A4, didynamous; anther loculi confluent; G (2), 2-loc. or by segmentation 3 — io-loc., in the former case with i — 8, in the latter with i, pend. anatr. ovule in each loc. Drupe. Endosperm. Chief genera: Pholidia, Myoporum. Myoporineae (BH.) = Myoporaceae. Myoporum Banks et Soland. Myoporaceae. 25 Austr., E. As., Sandw. Is., Mauritius. M. laetnin Forst. f. (N.Z.) yields useful timber. Myoschilos Ruiz et Pav. Santalaceae. i Chili. Myoseris Link = Pterotheca Cass. (Comp.). Myosotidium Hook. Boraginaceae (iv. i). i sp. Chatham Is. Myosotis L. Boraginaceae (iv. 4). 35 sp. |^ temp. 8 in Brit, (scorpion-grass, forget-me-not). The corolla-mouth is nearly closed by scales, and in some there is a coloured ring at the entrance form- ing a honey guide (see Life of Sprengel, in Nat. Science, Apr. 1893). The colour of the C changes as it grows older (see fam.). Myosurandra Baill. Hamamelidaceae. i Madagascar. Myosurus L. Ranunculaceae (3). 7 temp. M. minimus L. (mouse- tail) in Brit. Recept. much elongated. MYRMECODTA 439 Myracrodruon Allem. = Astronium Jacq. p.p. (Anacard.). Myrcsugenia Berg. (Myrtus p.p. BH.). Myrtaceae (i). 15 temp. S. Am. Myrcia DC. Myrtaceae (i). 500 trop. S. Am., W.I. Myrcialeucas Roj. Myrtaceae (i). i Argentina. Myrciaria Berg. (Eugenia p.p. BH.}. Myrtaceae (i). 65 trop. S. Am., W.I. Myriactis Less. Compositae (3). 3 Java to Persia. Myrialepis Becc. (Plectocomiopsis p.p"). Palm. (in. 2). i Perak. Myrianthemum Gilg. Melastomaceae (i). i trop. Afr. Myrianthus Beauv. Moraceae (in). 5 trop. Afr. Myriaspora DC. Melastomaceae (i). 2 trop. S.Am. Myrica L. Myricaceae. 40 ^ and Andes, esp. subtrop. M. Gale L. (sweet gale or bog-myrtle) in Brit, in mountain bogs. Its 1. have a pleasant resinous smell when rubbed or on hot days. Fls. in short catkins, achlam. The J has usu. 2 bracteoles and 4 sta. (2 — 16); the 1 2 — -4 bracteoles and (2) cpls. with one erect orthotr. ov. Nut, the exocarp secreting wax. No endosp. M. cerifera L. (N. Am., wax- myrtle, bay-berry) and others used as sources of wax ; the frs. are boiled. Myricaceae (EP., BH.). Dicots. (Archichl. Myricales; Unisexuales BH.). Only genus Myrica (q.v-)- Myricaria Desv. Tamaricaceae. to Scandinavia to China. Myriocarpa Benth. Urticaceae. 8 trop. Am. Fls. oo , in catkins. Myriocephalus Benth. Compositae (4). 8 temp. Austr. Myrioneuron R. Br. Rubiaceae (i. 7). 8 Assam to Borneo. Myriophyllum Ponted. ex L. Haloragidaceae. 2ocosmop.; 2 in Brit, (water milfoil). Submerged water plants, with usu. whorled much- divided L, borne on shoots that spring from the rhizome-like creeping stems. Land forms occasionally produced in some. The infl. projects above water; fls. wind-fertilised. Hibernation by winter-buds as in Utricularia. Myriopteron Griff. Asclepiadaceae (i). i E. Indomal. Myriostachya Hook. f. Gramineae (10). i Indomal. Myripnois Bunge. Compositae (12). 2 N. China. Myristica L. Myristicaceae. 80 palaeotrop. Trees with 2-ranked exstip. evergr. 1. and dioec. reg. fls. Fr. a berry ; splits by both sutures, disclosing a large seed — the nutmeg — with a curious branched red aril — the mace — around it. The nutmeg of commerce is the seed of M. fragrans Houtt. (M. moschata Thunb.), Moluccas. Myristicaceae (EP., BH.}. Dicots. (Archichl. Ranales; Micrembryae BH.}. 18 gen., 275 sp., trop., esp. As. Trees and shrubs with simple evergr. 1. with oil cells, and racemes of fls., unisexual, reg., usu. 3-merous. P (3), simple (cf. Monodora); A (3 — 18), extr.; G i, with i basal anatr. ov. Fleshy fr. Aril. Endosp. ruminate. Myrmechis Blume. Orchidaceae (n. 2). 3 Java to Japan. Myrmecodia Jack. Rubiaceae (n. 5). 20 E. Indomal. Epiph. with leafy stems. The base forms a large tuber, fastened to the support by adv. roots, composed of a large mass of tissue, chiefly cork, pene- trated by numerous communicating galleries and chambers, inhabited by ants. These galleries are formed in a peculiar way ; after germ, the 440 MYRMECODIA hypocotyl swells into a small parenchymatous tuber, and in this, in an axial direction, there appears a hollow cylinder of phellogen which proceeds to form cork on its inner side and parenchyma on the outer, thus adding to the bulk of the tuber and at the same time forming in it a hollow space (for the tissue within the cylinder of cork dies and dries up) open to the surface. Near the outer surface of the tuber is a phellogen layer acting in the ordinary way, forming a bark. As the tuber grows more pliellogens appear like the first, adding more paren- chyma and forming new cavities which are always in communication with the old. It has not been proved that the ants are of any service to the plant (see Acacia, Cecropia). The tuber is perhaps a water- storage apparatus. MyrmecopMly, symbiosis with ants, cf. Acacia, and Bambacaceae, Cecropia, Clerodendron, Cuviera, Duroia, Humboldtia, Hydnophytmn, Korthalsia, Macaranga, Maieta, Myrmecodia, Nauclea, Rubiaceae, Triplaris. Myrmedoma Becc. Rubiaceae (n. 5). i New Guinea. Epiphytic. Myrmephytum Becc. Rubiaceae (n. 5). i Celebes. Epiphytic. Mynnidone Mart. Melastomaceae (i). 2 trop. S.Am. Myrobalans (gall-nuts), astringent fr. used for tanning and in medicine : chebulic -, Terminalia Chebula Retz. ; emblic -, Pkyllanthus Em- blica L. Myrobalanus Gaertn. = Terminalia L. p.p. (Combret.). Myrocarpus Allem. Leguminosae (in. i). 2 Rio de Janeiro. Yield a balsam like balsam of Peru. Myrodia Sw. (Quararibea p.p.EP.). Bombacaceae (Sterculiaceae j5//.). 7 trop. Am. Myrosma L. f. Marantaceae. 8 S. Am. Myrospermum Jacq. Leguminosae (in. i). i trop. Am., W.I. Myrothamnaceae (EP., Hamamelidaceae p.p. BH.}. Dicots. (Archichl. Resales). Only genus Myrothamnus (q-v.). Myrothamnus Welw. Myrothamnaceae. 2 Afr., Madag. Xero. shrubs with opp. 1. and spikes of achlam., reg. , dioec. fl. i 4 — 8 sta. ; ¥ G (4 — 3). Septic, caps. Endosp. Myroxylon J. et G. Forst (Xylosma Forst. f.). Flacourtiaceae (4). 45 trop. (exc. Afr.). Myroxylon L. f. (Toluifcra L.). Leguminosae (in. i). Strop. S.Am. M. Pereirae Klotzsch yields the medicinal balsam of Peru, M. tolui- f , anatr., in many rows. Capsule leathery, loculic. Seeds light with long hair-like processes at the ends ; embryo straight, in fleshy endosp. Many sp. and hybrids cult. [See Goebel's Pflanzenbiol. Sch., Macfarlane in Ann. of Bot. ill. and Vll., and cf. Sarracenia, Cephalotus.] Nepeta Riv. ex L. (incl. Glechoma L.). Labiatae (vi). 150 _*. N. Glechoma Benin, (ground-ivy) and N. Cataria L. (cat-mint) in Brit. Fls. gynodioecious. Nephelaphyllum Blume. Orchidaceae (n. a. n). 6 E. As. Nephelium L. (excl. Litchi Sonner.). Sapindaceae (i). 25 Indomal. N. lappaceum L. (rambutan) cult. ed. fr. N. Longana Cambess. (longan) and others also used. [N. Litchi Cambess, see Litchi.] Nephelochloa Boiss. Gramineae (10). i W. As. Nephradenia Decne. Asclepiadaceae (n. 3). 5 Braz. to Mex. Nephrocarpus Dammer. Palmaceae (iv. i). i New Caled. Nephrocarya Candargy. Boraginaceae (iv. 3). i Greece. Nephrodium Rich. =Dryopteris Adans. p.p. (Polypod. ). Nephrolepis Schott. Polypodiaceae. 18 trop., and Japan, N.Z. They produce runners like strawberry, but not axillary, which root and give new pi. Nephropetalum Robinson et Greenman. Sterculiaceae. i N. Am. Nephrophyllidium Gilg (Menyanthes p.p. Bff., Fauria p.p. EP.). Gentianaceae (n). i N.W. Am., Japan. Nephrophyllum A. Rich. Convolvulaceae (i). i Abyssinia. Nephrosperma Balf. f. Palmaceae (iv. i). i Seychelles. Nephrostigma Griff. Anonaceae (inc. sed.). Nomen. Nephthytis Schott. Araceae (iv). 4 trop. W. Afr. Nepsera Naud. Melastomaceae (i). i trop. S. Am., W.I. Neptunia Lour. Leguminosae (i. 4). 10 trop. and subtrop. N.oleracea Lour, has a floating stem, rooting at the nodes, and covered by aeren- chyma. The 1. are sensitive like those of Mimosa. Fls. in heads, the lower c? , or neuter with petaloid stds. Neraudia Gaudich. Urticaceae (3). 3 Hawaiian Is. Neriacanthus Benth. Acanthaceae (iv. B). i Jamaica. Neriandra A. DC. = Skytanthus Meyen (Apocyn.). Nerine Herb. Amaryllidaceae(i). 15 Cape Colony. Cult. orn. fl. NE W JERSE Y TEA 449 Nerisyrenia Greene (Greggia EP.). Cruciferae (4). 2 W. N. Am. Nerium L. Apocynaceae (n. i). 3 Medit. to Japan. N. Oleander L. (oleander) has pits on the lower surface of the evergr. 1., in which the stomata are sunk (several in each) and covered with hairs, reducing transpiration. Fls. suited to long-tongued moths. Nerophila Naud. Melastomaceae (i). i Senegambia. Nertera Banks et Soland. Rubiaceae (n. 7). 10 Andes, N.Z., Austr., Sandw. Is., Malaya. Nerve (L), the vascular bundles which run through it. Nervilia Comm. ex Gaudich. (Pogonia p.p. BH.). Orchidaceae (n. 2). 25 trop. and subtrop. %. Nesaea Comm. ex Juss. Lythraceae. 30 Afr., Austr., As., N. Am. Nesiota Hook. f. Rhamnaceae. i St Helena. Neslia Desv. Cruciferae (4). i Eur., N. As., Medit. Nesodoxa Calest. (Ercmopanax p.p.). Araliaceae (i). i New Caled. Nesodraba Greene (Draba p.p.)- Cruciferae (4). 4 W. N. Am. Nesogenes A. DC. Verbenaceae (3). 2 Roclrigues, Polynesia. Nesogordonia Baill. Inc. sed. i Madag. Nesothamnus Rydberg \Ptrityle p.p.). Compos. (6). i Lower Calif. Nestlera Spreng. Compositae (4). 10 Cape Colony. Nettle, Urtica; dead -, Lamium ; devil or fever -, Laporiea ; hemp-, Galeopsis ; Nilgiri -, Girardinia; Spanish- (W.I.), Bidens; -tree, Celtis, (W.I.) Pilea. Nettoa Baill. Tiliaceae. i Austr. Net-veined, with irreg. network of veins, most Dicots. Neuburgla Blume. Apocynaceae (i. i). i Malaya. Neumannia A. Rich. (Aphloia BH.). Flacourtiaceae (4). 4 Madag., S. Afr. Neuracanthus Nees. Acanthaceae (iv. A). 10 palaeotrop. Neurachne R. Br. Gramineae (3). 3 Austr. Neurada L. Rosaceae (iv). i Medit. to E. Ind. Neurocalyx Hook. Rubiaceae (i. 2). 6 Ceylon, Borneo. Neurocarpaea P. Br. (Pentas p.p.). Rubiaceae (i. 2). 3 Afr., Madag. Neurocarpum Desv. = Clitoria L. p.p. (Legum.). Neurolaena R. Br. Compositae (8). 2 W.I., Colombia. Neurolobium Baill. Apocynaceae (i. 3). i Brazil. Neuroloma Anclrz. ex DC. =Parrya R. Br. (Crucif.). Neuropeltis Wall. Convolvulaceae (i). 4 trop. As. and Afr. Neurosoria Mett. Polypodiaceae. i trop. Austr. Neurotheca Salisb. ex B. et H. f. Gentian, (i). 5 trop. Afr., S. Am. Neustanthus Benth. = Pueraria DC. p p. (Legum.). Neuter, without fertile sporophylls, Centaurea, Hydrangea, Muscat-i, Viburnum. Neuwiedia Blume. Orchidaceae (i. i). 6 E. Indomal. Nevesarmondia K. Schum. (Pithecocteniiim p.p.). Bignoniaceae (i). i Brazil. Neviusia A. Gray. Rosaceae (in. i). i Alabama. Newberrya Torr. Pyrolaceae. i Oregon, Washington. Newbouldia Seem. Bignoniaceae (2). 3 trop. W. Afr. Newcastlia F. Muell. Verbenaceae (3). 7 trop. Austr. New Jersey tea, Ceanothus. W. 29 450 NE WTONIA Newtonia Baill. Leguminosae (i. 4). 2 trop. W. Afr. Newtonia O. Hoffm. (Antutiesia BH., Gongrothamnus EP.). Com- positae (8). i Angola. New Zealand flax, hemp, Phormium tenax Forsk.; -- daisy bush, Oleandra ; - - holly, Osmanthiis ; - - pincushion, Raoulia ; - - spinach, Tetragonia expansa Miirr. Neyraudia Hook. f. Gramineae (10). i trop. As., Afr., Madag. Ngai camphor, Blmnea balsam if era DC. Nicandra Adans. Solanaceae (i). i Peru, JV. physaloides Gaertn. Ov. divided in an irreg. way by plac. Berry nearly juiceless and with co seeds, enclosed in the enlarged K. Cult. orn. fl. Nicker bean, Caesalpinia bouducella Flem., Entada. Niclouxia Battandier. Compositae (4). i S.W. Sahara. Nicobar breadfruit, Pandanns Leram Jones. Nicodemia Tenore Loganiaceae. 3 Madagascar, Mascarene Is. Nicolasia Sp. Moore. Compositae (4). 3 S.W. trop. Afr. Nicolletia A. Gray. Compositae (6). 2 S.W. U.S. Nicolsonia DC. = Desmodium Desv. p.p. (Legum.). Nicoteba Lindau (Justicia p.p.). Acanthaceae (IV. B). 5 palaeotrop. Nicotiana L. Solanaceae (4). 45 Am., Polynes., i Austr. N. Taba- citm L., cult, in warm countries, esp. U.S., Cuba, Sumatra, Egypt, Brazil, etc., is the tobacco, grown as an annual crop; the 1. are gathered, hung up and slowly dried, then packed in heaps and fermented slightly. Different varieties are grown, and usu. in different places, for cigar, cigarette, and pipe tobacco. N. rustica L. and others are also used. Nidorella Cass. Compositae (3). 20 Abyssinia to S. Afr. Nidularium Lem. (A'arafas BH.}. Bromeliaceae (4). 15 Brazil. Niebuhria DC. (Maerua p p. EP.). Capparidaceae (a). 12 trop. As., Afr. Niederleinia Hieron. Frankeniaceae. i temp. S. Am. Niedzwedzkia B. Fedtschenko = Sesamum L. p.p (Pedal.). Niemeyera F. Muell. Sapotaceae (i). i trop. E. Austr. Nierembergia Ruiz et Pav. Solanaceae (4). 20 trop. and subtrop. Am. Nietneria Klotzsch et R. Schomb. Liliaceae (j). i Brit. Guiana. Nigella (Tourn.) L. Ranunculaceae (2). 16 Medit., Eur., often cult, orn. fl. (love-in-a-mist, devil-in-a-bush). Annuals. Alt. with the K is an invol. of 5 1. Within the coloured K are 5 — 8 nectaries, pocket - like structures with lids which prevent small insects from reaching the honey. The cpls. are more or less completely united but have separate styles; they give a caps. fr. Fl. protandrous. Niger (Lat.), black. Niger seed, Guizotia abyssinica Cass. Night-flowering cactus, Cereus gratutiflonis Mill., &c. Nightshade, Solatium ; deadly -, Atropa Belladonna L. ; enchanter's -, Circaea lutetiana L. Nigrescens, nigricans (Lat.), blackish. Nigribicchia, x E. G. Camus, Beyer, et R. Camus. Orchidaceae. Hybrid Nigritella-Habenaria (Bicchia). Nigritella Rich. (Habenaria p.p. BH.}. Orchidaceae (n. i). i Mts. of Eur. NOMENCLATURE 451 Nilgiri nettle, Girardinia heterophylla Decne. Nim, neem, Azaciirachta indica A. Juss. (Afelia Azadirachta L.). Nimble Will (Am.), Rhiehlenbergia diffusa Schreb. Niopo tree (W. I.), Piptadenia. Nipa Thunb. Palmae ( vi). i palaeotrop., N.fritticans Thunb., a low- growing palm with monoec. infl. Fr. woody, combined into a dense head ; each contains one seed. It grows in brackish water and is very char, upon trop. coasts. [See Phytelephas.] Niphaea Lindl. Gesneriaceae (n). 2 Guatemala, Cuba. Niphobolus Kaulf. =Cyclophorus Desv. (Polypod.). Nipplewort, Lapsana communis L. Nirarathamnos Balf. f. Umhelliferae (in. 5). i Socotra. Nirwamia Rafin. Euphorbiaceae (inc. sed.). i Japan. Nisa Noronha= Homalium Jacq. p.p. (Flac.). Nissolia Jacq. Leguminosae (in. 7). 7 trop. and subtrop. Am. Nitidus (Lat.), lustrous, smooth and shining. Nitraria L. Zygophyllaceae. 3 palaeotrop. Nitrophila S. Wals. Chenopodiaceae (A), i W. N. Am. Nivalis (Lat.), growing near snow; niveus (Lat.), snow-white. Nivenia R. Br. (Paranomns p.p. EP.). Proteaceae (i). 12 S. Afr. Nivenia Vent. (Aris.'ea BH.}. Iridaceae (n). 2 S. Afr. Cult. orn. fl. Noaea Moq. in DC. Chenopodiaceae (B). 7 W. As. Noccaea Moench (Htttchinsia p.p. EP.). Cruciferae (4). 2 Alps. Node, the joint where a leaf springs from the stem. Nodocarpaea A. Gray. Rubiaceae (n. 10). i Cuba. No-eye pea (W.I.), Cajanus indicus Spreng. Noisettia H. B. et K. Violaceae. i Brazil, Peru, Guiana. Nolana L. Nolanaceae. 20 Chili, Peru. Many are shore plants with fleshy 1. Nolanaceae (EP. • Convolvulaceae p.p. BH.). Dicots. (Sympet. Tubi- florae). 3 gen., 30 sp. , W. coast of S. Am. Herbs or low shrubs with simple L, often covered with glandular hairs. The 1. in the veg. region, are alt., but in the infl. portion they become paired in the same way as in Solanaceae (q.v. )• Many are sea-shore plants with fleshy 1. Fls. sol. in the leaf-axils, 5 , reg. K (5); C (5); A 5, alt. with petals ; G typically 5, only united in Alona, usu. free and divided by irreg. longitudinal constrictions into 5 or 10 portions standing in a row, or by longitudinal and transv. constrictions into 10 — 30 portions in 2 or 3 rows. The fr. consists of a corresponding number of r — 7- seeded nutlets. Stipe i. Seed album. Genera: Nolana, Alona, Dolia. Nolina Michx. Liliaceae (vi). 10 S.W. N. Am. Xero. Cult. Nolletia Cass. Compositae (3). 4 Morocco, S. Afr. Noltea Reichb. Rhamnaceae. i S. Afr. Cult. orn. shrub. Nomaphila Blume (Hygrophila p.p. EP.). Acanth. (iv. A). 8 palaeo- trop. Nomenclature (with esp. reference to classification). The unit in classifi- cation is the species, which was described by A. L. de Jussieu as "the perennial succession of similar individuals perpetuated by generation." All marsh marigolds (Caltha palustris) form one sp., or all pi. of Indian corn (Zea Mays) or coconut (Cocos mid/era). But exactly to 29 2 452 NOMENCLATURE define a sp. is impossible. Each man in practice arrives at his own conception somewhere between (or at) the extremes usu. called Lin- nean and Jordanian sp. Draba (Erop/iila) verna for example (Linnean) is distinguished from other D. by absence of petiole and oblong- elliptical pod, but Jordan, studying the sp. in great detail, split it into a great number of forms, to which he gave specific rank, when he found that each one continued to breed true. These were distinguished by "small" characters, such as differences in hairiness, fruit-shape, leaf-form, &c. Cf. Jordan, Diagnoses i cpl. ; Areca -, Areca; Australian chest-, Castanospermum australe A. Cunn.: Bambarra ground-, Voandzeia subterranea Thou.; betel-, Areca Catechu L. ; bladder -, Staphylca ; Brazil-, Be'-tholletia excelsa Humb. et Bonpl. ; bread -, Brosinntm ; butter -, Caryjcar nuciferum L. ; candle -, Aleurites triloba Forst. ; cashew -, Anaeardium occidentale L. ; chest-, Castanea vulgaris Lam. ; cob-. Corylus ; COCO-, Cocos nu- cifera L. ; cola-, Cola vera K. Schum. ; coquilla -, Attalea funifera Mart. ; earth-, Arachis hypogaea L., Couopodiitm dennJatiim Koch; -grass, Cypents; ground-, Arachis hypogaea L. ; hazel-, Corylus Avellana L. ; hog-, Spondias ; horn-, Trapa ; horse-chest-, Aesculus Hippocastamnn^.; ivory-, Phytelephas ; kola-, see cola ; marking-, Semecarpus ; pea-, Arachis hypogaea L. ; pistachio-, Pistachia vera L. ; pecan-, Carya; physic -, Jatropha Curcas L. ; -rush (Am.), Scleria; Sapucaia-, Lecythis; Souari-, Caryocar nuciferum L. ; -tree (Austr. ), Macadamia. Nutans (Lat.), nodding. Nutation, lateral swaying of tip of a growing organ. 456 NUTLETS Nutlets, Labiatae (fr.). Nutmeg, Myristica fragrans Houtt., Alonodora Myristica Dun. ; Bra- zilian-, Cryptocarya; calabash-, Monodora Myristica Dun. ; -grass (Am.), Cyferus rotnndus L. ; Peruvian-, Laurelia aromatica Juss. Nuttallia DC. = Nemopanthes Rafin. (Aquifol.). Nuttallia Rafin. = Mentzelia Plum. p.p. (Loas.). Nuttallia Torr. et Gray. Rosaceae (v). i N.W. Am. Like Prunus, but with 5 free cpls. Nuxia Comm. ex Lam. Loganiaceae. 20 Afr., Madag. Nuytsia R. Br. Loranthaceae (i). i VV. Austr., a small tree, doubt- hilly parasitic on roots. Cotyledons 3. Nyctaginaceae (EP., BH.}. Dicots. (Archichl. Centrospermae ; Cur- vembryae BH.}. 20 gen., 160 sp., mostly trop. and esp. Am. Trees, shrubs or herbs with opp. (often unequal) 1. and no slips. Fls. in cymes, $ or unisexual, and with much variety. At the base of the fls. are usu. several bracts, often large and coloured. In Bougainvillaea 3 large conspicuous bracts enclose a group of 3 fls. In Abronia the number of bracts and fls. is larger, while in Mirabilis there is only one fl. and the involucre resembles a calyx. P usu. (5), petaloid, peisistent upon the ripe fr. ; usu. the upper part drops away and the fr. remains in the lower part, which is termed the anthocarp, and may become glandular, or form an umbrella-like wing, or otherwise serve for seed- dispersal. A typically 5, alt. with the P, but often 3, 8, 10 or other numbers, or raised to -20 or 30 by branching; filaments often of un- equal length ; G i, with long style and i basal erect ana-campylotr. ov. Achene enclosed in the P. The N. are of slight economic value ; see Mirabilis, Neea, &c. Chief genera: Mirabilis, Bougainvillaea, Pisonia, Neea, Reichenbachia. Nyctagineae (BH.) — Nyctaginaceae. Nyctaginia Choisy. Nyctaginaceae. i Texas, Mexico. Nyctago Juss. = Mirabilis L. p.p. (Nyct). Nyctanthes L. Oleaceae. i Indomal. Nycterinia D. Don = Zaluzianskya F. W. Schmidt (Scroph.). Nycterium Vent. = Solanum Tourn. p.p. (Sol.). Nycticalos Teijsm. et Binnend. Bignoniaceae (i). i Malaya. Nyctocereus Britton et Rose (Ceretis p.p.). Cactaceae (in. i). 3 Mexico, Nicaragua. Nyctophyla Zipp. Inc. sed. i Timor. Nymania K. Schum. Euphorbiaceae (A. i. i). i New Guinea. Nymania S. O. Lindb. Meliaceae (in), i S. Afr. Nymphaea (Tourn.) L. (Castalia Salisb.). Nymphaeaceae (in). 40 trop. and temp. Ar. alba L. (white water-lily) in Brit. Many cult., e.g. N. Lotus L. sometimes supposed to be the sacred lotus of Egypt (see Nelumbium), They grow in shallow water. There is a stout creeping rhiz. ; at the tip it is bent up, and bears slip. 1. and fls. on long stalks. The peduncle occupies the position of one of the 1. of the spiral, and there is no bract at its base. The 1. is large and floats on the surface ; it is nearly circular, entire, and leathery, with stomata, cuticle and palisade tissue on the upper side. Fl. 5 , reg., acyclic ; floats on the surface. The 4 outermost floral 1. exhibit a peculiar aestivation, the ant. being entirely outside, the post. NYSSANTHES 457 inside the lat. 1. According to Caspary (Eichler, Blnthetidiag. n. 184) the ant. 1. repres. the bract (' adnate ' to the peduncle ; ff. Solanaceae), the two lat. 1. the bracteoles, the post, a true sepal. Most authors regard the 4 1. as a K. C well developed ; 4 outer petals alt. with the sepals, and 4 inner alt. with the outer. These 8 form the starting points of as many spirals of petals, usu. 4 in each, alt. approximately with the outer 8 and with one another. As we pass inwards the petals become narrower and show transition forms to the sta., which to the number of 50 or 100 continue the fl. inwards. Whilst the K is hypog. the petals and sta. are inserted up the sides of the G, which has 10—20 loculi, each containing oo ov. scattered over the whole carpellary surface (cf. Butomus). The sessile stigmas form a number of rays upon the upper surface of the G, as in a poppy. The fr. is a large berry containing oo seeds, each covered by a spongy aril. Between aril and seed are air-bubbles. The fr. ripens under water and when it dehisces the mass of seeds floats up; then the individual seeds part company, each drifting about until the air escapes (by decay of the aril or otherwise), when it sinks to the bottom of the pond. There is a large perisperm round the endosp. proper. [Conard, The Waterlilies, Washington, 1905.] Nymphaeaceae (EP., BH.}. Dicots. (Archichl. Ranales). 8 gen., 60 sp., cosmop. Water or marsh pi. usu. with rhiz., and with submerged, floating, and aerial 1. and sol. usu. large fls. of great variety of pattern. Cabomba is most simple and agrees in type with the other Ranales ; whilst in Nuphar, Nymphaea, Victoria, Nelumbium, &c., great modifi- cation appears, esp. in the G. In Nelumbium the ovary is still apocp., though the cpls. are connected by the curious torus, whilst in the other gen. it is syncp. ; in Nuphar it is sup., in Nyrnphaea semi-inf. , in Victoria inf. The P too shows much variety, from the simple Cabomba-type to Nuphar, &c. Ovules usu. anatr. The seed has both endosperm and perisperm (exc. Nelumbium), and is often aril- late. For full details of the floral structure see chief gen., and Eichler's Bliithendiagramme. Classification and genera (after Caspary); I. NELOMBONOIDEAE (seed exalbuminous ; cpls. free in obconical receptacle) : Nelumbium (only genus). II. CABOMBOIDEAE (endosperm and perisperm; cpls. free): Cabomba, Brasenia. III. NYMPH AEOIDEAE (do., but cpls. united); Victoria, Euryale, Nymphaea, Nuphar, Barclaya. Nyropnylla Neck. Lauraceae. Nomen. Nyssa Gronov. ex L. Nyssaceae. 6 N. Am., Himal., Malaya. N.syl- vatica Marsh and others in N. Am. (tupelo, pepperidge, gum-tree) yield timber and ed. fr. Nyssaceae (EP.; Cornaceaep.p. BH.}. Dicots. (Archichl. Myrtiflorae). 3 gen., 10 sp., E. A?., E. N. Am. Shrubs with alt. exstip. 1. and small 5 or unisexual fls. in heads, racemes, umbels, or sol., with flat or hollow recept. K 5 or more, C 5 or o, A twice as many or less, G usu. i-loc., sometimes 6 — lo-loc. with one ov. in each. Usu. drupe. Endosp. Nyssanthes R. Br. Amarantaceae (2). 2 Austr. 458 NYSSOPSfS Nyssopsis O. Ktze. = Camptotheca Decne. (Corn.)- Oak, QIU-/-CHS ; American turkey -, Querc us obiusiloba ; Ceylon -, Schlei- chera trijuga Willcl.; cork-, Quercus Suber L. ; Dominica-, Ilex sideroxyloidesGrKeb.; -fern, Dryopteris Linneana C. Chr. ; forest-, Casuarina ; dyer's -, Quercus linctoria Bartr. ; live -, Quercus virginiana Mill, and other evergr. sp. ; patana - (Ceylon), Careya arboiea Roxb. ; holly-, Quercus Ilex L. ; Quebec-, Quercus alba L. ; she-, Casuarina; silky-, Grevillea; Turkey-, Quercus Cerris L. ; white -, Quercus alba L. Oakesia S. Wats. (Uvularia EP.}. Liliaceae (i). 2 N. Am. Oakesiella Small (Uvularia p.p.). Liliaceae (i). 3 N. Am. Oat, Avena saliva L.; -grass, Arena fatua L., &c.; side-, Boute- loiia. Oaxacania Robinson et Greenman. Compositae (2). i Mexico. Ob- (Lat. pref.), inverted ; -diplostemonous (sta.), in two whorls, the outer opp. to the pets., Burseraceae, Caryophyllaceae, Crassulaceae, Oxalidaceae, Saxifragaceae, Zygophyllaceae ; -lanceolate, -lique, -long, -ovate, &c. , see Leaf; -solete, aborted. Obbea Hook. f. Rubiaceae (11. 2). i Hawaiian Is. Obeliscaria Cass. = Lepachys Rafin. (BH. ) . = Rudbeckia L. p.p. Oberonia Lindl. Orchidaceae (n. 4). 80 palaeotrop. Obetia Gaudich. Urticaceae (i). i Madagascar, Bourbon. Obione Gaertn. = Atriplex L. p.p. (Chenop.). Obolaria L. Gentianaceae (i). i N. Am. Saprophyte (<./. Bartonia) of a purplish green colour with scaly 1. Occidentalis (Lat.), western. Oceanium (Cl.), an ocean formation. Oceanorus Small (Amiauthiitm A. Gray). Liliaceae (i). i N. Am. Ochagavia Phil. Bromeliaceae (4). i Juan Fernandez. Ochanostachys Mast. Olacaceae. i Malay Peninsula, Borneo. Ochlandra Thw. Gramineae (13). 6 India, Ceylon. Ochna L. Ochnaceae. 45 trop. As., Afr., Cape Col. K coloured. Cpls. 3 — 15, free below, but with a common style. After fert. the style falls and each cpl. gives a drupe, while the recept. becomes fleshy under them. The 1. shows veining well. Ochnaceae (EP.\ BH. place Sanvagesia, &c. in Violaceae). Dicots. (Archichl. Parietales ; Geranialer BH.}. 17 gen., 210 sp., trop. Most are trees or shrubs with alt. usu. simple slip. 1. and panicles, racemes or cymes (Sauvagesia, &c.) of? , usu. reg. fls. K 5, free or united at base, imbr.; C 5, rarely 10, contorted ; A 5, 10, or oo , hypog. or on an elongated axis ; G (2 — 5), rarely (10 — 15), often free below with common style (cf. Apocynaceae). Ovules i — 2 — oo in each cpl., erect or rarely pend., always with ventral raphe. The axis swells and becomes fleshy under the fr. , which is usually a cluster of drupes, but sometimes a berry or capsule. Endosp. or not. Chief genera: Ochna, Gomphia, Sauvagesia. Ochocoa Pierre (Scyphocephalium EP. ). Myristicaceae. i trop. Afr. Ochra, Hibiscus esculentns L. Ochradenus Delile. Resedaceae. 5 S. Medit. Ochrea, ocrea, sheathing stipule, Polygonaceae. Ochrocarpus Thou. Guttiferae (iv). 10 palaeotrop. OD ONTONEMA 459 Ochroma Sw. Bombacaceae. i trop. S. Am., W.I., O. Lagopus Sw. (balsa, corkwood). Wood very light. Seeds embedded in hairs. Ochronerium Baill. Apocynaceae (n. i). i Madagascar. Ochropteris J. Sm. Polypodiaceae. i Madagascar, Mascarene Is. Ochrosia Juss. Apocynaceae (i. 3). 15 palaeotrop. Ochthocliaris Blume. Melastomaceae (i). 5 Malaya. Ochthocosmus Benth. (PAyltocosmusKlotzscti). Lin. 3 trop. Am. , Afr. Ochthodium DC. Cruciferae (2). i W. As. Ocimum L. Labiatae (vil). 60 sp. trop. and warm temp. 0. Basi- licum L. is the basil, sacred in the Hindu religion (tulsi). Oclemena Greene (Aster L. p.p.). Compositae (3). 2 N. Am. Ocotea Aubl. Lauraceae (i). 230 trop. and subtrop. O. bullata E. Mey. (S. Afr.) yields a useful timber (stinkwood). Ocotilla, Fouquieria splendens Engelm. Ocrearia Small (Saxifraga p.p.). Saxifragaceae (i). i W. N.Am. Octadesmia Benth. Orchidaceae (n. 6). 3 Jamaica, S. Domingo. Octarrhena Thw. = Phreatia Lindl. (Orchid'.). Octas Jack. Inc. sed. i Malaya. Octella Rafin. = Melastoma L., &c., p.p. (Melast.). Octoceras Bunge. Cruciferae (4). i W. As. Octodon Thonn. (Borreria p.p. £P.). Rubiaceae (n. 10). 2 trop. Afr. Octoknema Pierre. Olacaceae. 4 trop. Afr. Octolepis Oliv. Thymelaeaceae, once Flacourtiaceae. 4 W. Afr. Octolobus Welw. Sterculiaceae. i Angola. Octomelis Miq. Datiscaceae. 2 Malay Archipelago. Octomeria R. Br. Orchidaceae (n. 8). 15 Brazil to W.I. Octopleura Griseb. (Ossaea p.p. EP.). Melastom. (i). 5 trop. Am., W.I. Octotheca R. Viguier. Araliaceae (i). i New Caledonia. Octotropis Bedd. Rubiaceae (n. i). i Travancore. Odacmis Rafin. Inc. sed. i N. Am. Odiua Roxb. (Calesiuin Adans. ). Anacardiaceae (2). 15 trop. Afr., As. Odonia Bertol. (Galactia P. Br.). Leguminosae (in. 10). 8 trop. Am. Odont-, odonto- (Gr. pref.), tooth. Odontadenia Benth. Apocynaceae (n. j). 20 trop. S. Am. Odontandra Willd. ex Roem. et Schult. (Trichilia p.p. BH.). Melia- ceae (ill). 4 trop. S. Am. Odontanthera Wight. Asclepiadaceae (inc. sed.). Nomen. Odontarrhena C. A. Mey. = Alyssum Tourn. p.p. (Crucif.). Odontelytrum Hack. Gramineae (5). i trop. Afr. Odontioda x Rolfe. Orchidaceae. Hybrid, Odontoglossum-Cochlioda. Odontites (Riv.) Hall (Barlsia BH.). Scrophulariaceae (in. 3). 20 Medit., S. Eur., W. As. Semiparasites (see fam. ). Odontocarya Miers. Menispermaceae. 4 trop. S. Am., W.I. Oclontochilus Blume. Orchidaceae (n. 2). 10 Indomal, Polynesia. Odontocidium x . Orchidaceae. Hybrid, Odontoglossum-Oncidium. Odontocyclus Turcz. Cruciferae (inc. sed.). i Kurile Is. Odontoglossum H. B. et K. Orchidaceae (n. 19). too Mts. of trop. Am. Epiphytes. Many cult. orn. fl. Many hybrids. Odontonema Nees (Thyrsacanthus BH.). Acanthaceae (iv. B). 25 trop. Am. 460 ODONTONEMELLA Odontonemella Lindau (Eranthemum p.p.). Acanth. (iv. B). 2 Indo- mal. Odontonia x Rolfe. Orchidaceae. Hybrid, Odontoglossum-Miltonia. Odontonychia Small (Siphonychia p.p.). Caryoph. (i. 4). 2 N. Am. Odontosoria (Presl) Fee. Polypodiaceae. 20 trop. and subtrop., exc. Afr. Odontospermum Neck. (incl. Asteriscus Moench). Compositae (4). 12 Medit. O. (A.) pygmaeitm O. Hoffm. is a xero. whose fr.-heads close in dry weather (cf. Anastatica, Mesembryanthemum) ; the seeds only escape in damp weather suitable for germination. Odontostelma Rendle (Schizoglossum EP.}. Asclep. (n. i). i Angola. Odontostomum Torr. Liliaceae (in) (Haemodor. BH.}. i California. Odontotecoma Bur. et K. Schum. (Tecama p.p.). Bignoniaceae (2). i Brazil. OdontycMum K. Schum. (Hedychium p.p.). Zingiber (i). i Mai. Pen. Odostemon Rafin. (Berberis p.p.). Berberidaceae. 7 N. Am. Odyendea Engl. (Quassia p.p.). Simarubaceae. 2 trop. W. Afr. Oeceoclades Lindl.= Saccolabium Blume (Orchid.). Oecology, ecology. Oecopetalum Greenman et C. H. Thompson. Icacinaceae. i Mexico. Oedematopus Planch, et Triana (Havetiopsis BH.}. Guttiferae (v). 3 Amazon valley. Oedera L. Compositae (7). Cape Colony. Oegroe Phil. Compositae (inc. seel.), i Chili. Oenanthe (Tourn.) L. Umbelliferae (in. 5). 35 N. temp. \%. 7 in Brit, (water drop-wort). Oenocarpus Mart. Palmaceae (iv. i). 8 N. S- Am. Oenone Tul. (Ligta Tul.). Podostemaceae. 8 Guiana, Brazil. Oenosciadium Pomel (Oenanthe p.p. EP.}, Umbellif. (in. 5). i N. Afr. Oenothera L. (BH. incl. Godetia Spach, Onagra Tourn., Xylopleurum Spach). Onagraceae (2). 30 Am. O. (Onagra) biennis L. (evening primrose), &c., cult. orn. fl. The fls. of O. biennis emit scent at evening and are visited by nocturnal moths, to which they are suited by the long tubes. Oenotheraceae (Warming) = Onagraceae. Oenotheridium Keiche (Godetia p.p.). Onagraceae (2). i Chili. Oeonia (Aeonia} Lindl. Orchidaceae (n. 20). 5 Mascarene Is. Oeoniella Schlechter (Epidendritm p.p.). Orchid, (n. 6). 2 Madag., Masc. Ofaiston Rafin. Chenopodiaceae (B). i W. As. Officinalis (Lat.), medicinal. Offset, a short runner, bending up at the end, Agave, Sempervivum. Oftia Adans. Myoporaceae. 2 S. Afr. Ogeechee lime (Am.), Nyssa. Oianthus Benth. Asclepiadaceae (n. 3). 3 India. -oides (Gr. suff.), -like. Oil occurs in plants in two forms, the fixed oils, or non-nitrogenous reserves in seeds, and the volatile oils, which give the perfume to many fls. and 1. The former are obtained by pressure, the latter by distillation. Fixed oils from Alcnrites, Arachis (groundnut), Argania, OLACACEAE 461 Barringtonia, Brassica (rape, colza), Calophyllum, Carya, Cocos (coconut), Corylus, Croton, Elaeis (palm oil), Eruca, Fagus, Ginkgo, Glycine (soja, soy), Gossypium (cottonseed), Guizotia (nigerseed), Helianthus, Juglans (walnut). Linum (linseed), Melia, Moringa, Olea (olive), Papaver, Polygala, Ricinus (castor), Sapium, Schleichera, Sesamum (gingelly), Theobroma (cacao-butter), Tilia, Vateria, &c., &c. Some of these are drying oils, like linseed, and used in painting, some remain fluid, some are solid or fatty, esp. in Europe. More solid fatty bodies are obtained from Bassia, Butyrospermum, Caryocar, Pentadesma, &c. Volatile oils from Acacia, Backliousia, Calamintha, Cananga, Cinnamomum (cinnamon), Citrus (lemon, &c.), Cymbo- pogon (citronella, geranium, lemongrass), Dictamnus, Eucalyptus, Ettgenia (clove), Gaultheria (wintergreen), Jasminum, Labiatae, Lavandiila (lavender), Lippia, Melaletica, Mentha (peppermint), Nardostachys (spikenard), Origanum, Pelargonium, Pogostemon (patchouli), Reseda, Rosa, Roseniarinus, Santaluni (sandalwood), Sassafras, Thymus, Viola, &LC., &c. Andiroba -, Carapa ; argan -, Argania ; bay -, Laurus ; ben -, Moringa; bergamot-, Citrus; birch-, Betula; cajeput-, Melaleuca; camphor -, Cinnamomum ; caraway -, Cantin ; castor -, Ricinus ; chaulmoogra -, Gynocardia ; citron -, Citrus ; citronella -, Cymbo- pogon ; clove -, Eugenia ; coconut -, Cocos ; cohune -, Attalea ; colza -, Brassica ; cottonseed -, Gossypium ; croton -, Croton ; cumin -, Cuminum ; Florence (fine olive) -, Olea ; geranium -, Pelargonium, Cymbopogon ; gingelly -, Sesamum ; groundnut -, Arachis; hempseed -, Linum; illupi -, Bassia; jasmine -, Jas- minum; juniper -, Juniperus ; kekuna -, Aleurites; khus-khus -, Vetiveria; lavender-, Lavandula; lemon-, Citrus; lemongrass-, Cymbopogon; linseed -, Linum ; Macassar-, Cananga; margosa-, Azadirachta ; marjoram-, Origanum ; mustard -, Brassica ; neroli -, Citrus ; nim -, Azadirachta ; olive -, Olea ; palm -, Elaeis ; pat- chouli -, Pogoslemon ; peanut -, Arachis ; pimento -, Pimenta ; poppy-, Papaver; rantil-, Guizotia; rape -, Brassica ; rosemary-, Rosmarinus ; rue -, Ruta ; rusa -, Cymbopogon ; sandalwood -, Santahim; sanderswood-, Santaluni ; sassafras-, Sassafras; savin-, Juniperus; sunflower-, Helianthus; thyme-, Thymus; tonquin -, Dipteryx; turpentine-, Pinus, &c.; verbena-, Lippia; walnut-, Juglans ; wintergreen -, Gaultheria ; wood -, Dipterocarpus, Aleu- rites ; - glands or passages, Guttiferae, Lauraceae, Monimiaceae, Myrtaceae. Oilapetalum Pohl. Inc. sed. Nomen. Oionychion Nieuwland (Viola -p.-p. ). Violaceae. i N. Am. Oiospermum Less. Compositae (i). i Bahia. Oistonema Schlechter. Asclepiadaceae (n. 3). i Borneo. Okenia Schlecht. et Cham. Nyctaginaceae. i S. Mexico. Okra, Hibiscus esculentus L. Ola (Ceylon), Borassus, Corypha, &c. Olacaceae (EP.; Olacineae p.p. BH.). Dicots. (Archichl. Santalales). 25 gen., 1 20 sp., trop. Most are shrubs or trees with alt. entire 1. and small £ reg. fls. There is a distinct K, resembling the calyculus of Loranthaceae, but probably not equivalent to it. C 4 — 6 ; A as 462 OLACACEAE many or 2 or 3 times as many ; G partly sunk in the disc, or free, 2 — 5-loc. at base, i-loc. above, with free plac. and i ovule hanging down into each loc. (occasionally i-loc. i-ovuled). Drupe or nut, one-seeded. Seed with testa and endosp. C hie/ genera : Ximenia, Olax. Olacales (BH.). The 8th order of Polypetalae. Olacineae (BH., Olacaceae + Icac inaceae £P.). Trees and shrubs with usu. alt. 1. and axillary infl. of $or unisexual reg. fls. K, C (4 — 5 — 6), A 4 — 10, rarely 12, G free or partly in disc, (3 — 5), i- or multi-loc. with few ovules. Drupe i -seeded. Endosp. Olax L. Olacaceae. 35 palaeotrop. Oldenburgia Less. Compositae (12). 3 Cape Colony. Oldenlandia L. (incl. Hedyotis L.). Rubiaceae (i. 2). 180 trop. Some are heterostyled (dimorphic). Oldfieldia Hook. Euphorbiaceae (A. I. i). i trop. W. Afr. , O. afri- cana Benth. et Hook, f., the African oak. Good timber. Old maid (W.I.), Vinca rosea L. ; -man, Artemisia Abrotanum L.; - - 's beard, Tillandsia; --cactus, Cereiis senilis Salm-Dyck; - woman's bitter (W.I.), Picramnia ; - witch grass (Am.), Panicum capillare L. Olea (Tourn.) L. Oleaceae. 35 Medit., S. Afr., E. Ind., Austr., Polynes. O. europaea L. (olive), cult, in Medit. region from early ages. The wild form has thorny twigs and a small fr., the cult, form (var. saliva DC.) is smooth and has a large drupe with oily flesh. The oil is obtained by bruising and pressing the fruit. Several yield good timber, e.g. the olive, O. laurifolia Lem. (S. Afr. ; black ironvvood), &c. Oleaceae (EP., BH.}. Dicots. (Sympet. Contortae; Gentianales BH.). 21 gen., 400 sp., trop. and warm temp., esp. E. Ind. Shrubs and trees usu. with opp. L, which are exstip., simple or pinnate, often entire. Serial accessory buds occur in the leaf-axils of many sp. (e.g. Syringa) in both fig. "and veg. parts. The infl. is racemose or cymose, often bracteolate. Fls. ? , rarely unisexual, reg., 2 — 6-merous, sometimes poly- or a-petalous (Fraxinus, &c.). K typically (4), valvate ; C (4) valvate or imbr., rarely conv. ; A 2, epipet. usu. transv. placed, and alt. with cpls. ; no disc; G (2); stigma 2-lobed on simple style ; ov. 2-loc. with 2 anatr. ov. in each loc. Berry, drupe, or caps., or schizocarp, with i — 4 seeds. Endosp. or none, embryo straight. Olea, Fraxinus, &c. , are of economic value. Oleander, Neriitm Oleander L. Oleandra Cav. Polypodiaceae. 10 trop. Olearia Moench. Compositae (3). 90 Austr., N.Z. , New Guinea. Replaces Aster, and closely resembles it, but all trees or shrubs. Oleaster, Elaeagnus, Olea. Oleiferus (Lat.), oil-bearing. Oleine, Cocos. Oleoxylon Roxb. Dipterocarpaceae. i Burma. Oleraceus (Lat.), esculent. Olibanum, Bosiuellia Carteri Birdw., £c. Oligandra Less. Compositae (4). 3 trop. S. Am. Oliganthes Cass. Compositae (i). ' 8 trop. Am. Oligarrnena R. Br. Epacridaceae (3). i W. Austr. ON AGRA 463 Oligo- (Gr. pref.), few ; -merous, with fewer members in whorl. Oligobotrya Baker. Liliaceae (vn). i China. Oligocarpus Less. Compositae (9). 3 S. Afr. Oligocladus Chodat et Wilczek. Umbelliferae (ill. 6). i Argentina. Oligodora DC. (Athanasia p.p. EP. ). Compositae (4). i S. Afr. Oligogynium Engl. (Nephthytis p.p. BH.). Araceae (iv). i W. Afr. Oligolobos Gagnep. Hydrocharidaceae. i China, Indochina. Oligomeris Cambess. Resedaceae. 5 Africa, India, S.W. U.S. Oligonema S. Wats. Compositae (3). i Mexico. Oligoneuron Small (Solidago p.p.). Compositae (3). 4 N. Am. Oligosporus Cass. = Artemisia L. p.p. (Comp.). Oligostemon Benth. (Duparqiidia EP.). Leguminosae (n. s). i W. Afr. Oligothrix DC. Compositae (8). 2 trop. and S. Afr. Olinia Thiinb. Oliniaceae. 6 Afr-_ Shrubs with opp. entire 1. and panicles of? fls. K, C, A 4—5, G (3 — 5), 3 — 5-loc. each with 2 — 3 ovules ; short style. Drupe ; no endosp. Oliniaceae (EP. ; ' Lythra^eae p.p. BH.). Dicots. (Archichl. Thyme- laeales). Only genus Olinia ( r/u's mascula L.; fly-, Ophrys mitscifera Huds.; frog-, Habenaria (Coeloglossum) viridis R. Br.; lady's slipper-, Cypripedium ; man-, Aceras anthropophora R. Br.; musk -, Herminium Monorclns R. Br.; scented-, Gymna- denia conopsea R. Br.; spider-, Ophrys aranifera Huds.; spotted-, Orchis maculata L. A — D. Orchis mascula (Purple Orchis). A. Front view of anther and top of column ; a, anther ; s, s, pair of stigmatic surfaces, one on each side of the rostellum, r. The pollen-sacs have split lengthwise, exposing the pollinia in the upper portion. On either side of the anther is a blunt outgrowth representing a sterile stamen. B. Dissection of flower, side view, showing part of lip and the spur (s/>) and the relation of rostellum (>-) and stigma (s) to entrance of spur. C. Base of pair of pollinia, front view ; c, caudicles ; r, rostellum ; g, gland ; »i, membranous disc. D. Single pollinium, the pollen-containing portion separated, showing arrangement of packets of pollen on the two main axes. E. Pollinia of O. pyramidalis attached to a common gland (jr). All enlarged. A, D, from original drawing by Fr. Bauer. B, C, E, after Darwin. Orchiserapias x E. G. Camus. Orchidaceae. Hybrid, Orchis-Serapias. Orcuttia Vasey. Gramineae (10). i California. 472 ORDEAL BARK Ordeal bark, Erythrophlaeum ; - bean, Physostigma. Oreacanthus Benth. Acanthaceae (iv. B). i Cameroons. Oreantbes Benth. Ericaceae (in. -2). i Ecuador. Oregon cedar, Chamaeeyparis Lawsoniana Parl. Oreinotinus Oerst. = Viburnum L. p.p. (Caprif.). Oreiostachys Gamble. Gramineae (13). i Java. Oreo- (Gr. pref.), mountain. Oreobambos K. Schum. Gramineae (13). i E. Afr. Oreobatus Rydberg (Rubus p.p.). Rosaceae (in. 2). 2 N. Am. Oreobliton Dur. et Moq. Chenopodiaceae (A), i Algeria. Oreobolus R. Br. Cyperaceae (n). 3 S. Am., Austr., N.Z., Polynesia. Oreocallis Small (Leucothoe 'p.p.). Ericaceae (n. i). i N.W. U.S. Oreocarya Greene (Krynitzkia p.p.). Boraginaceae (iv. 2). 20 Pac. N. Am. Oreocereus Riccob. (Pilocerens p.p.). Cactaceae (ill. r). i Boliv. Oreocbaris Benth. Gesneriaceae (i). 8 China, Japan. Oreochloa Link. Gramineae (to). 2 S. Eur. Oreocbrysum Rydberg (Aplopappns p.p.). Compositae (3). i N. Am. Oreocnida Miq. =Villebrunea Gaudich. (Urtic.). Oreodaphne Nees et Mart. = Ocotea Aubl. p.p. (Laur.). Oreodoxa Willd. Palmae (iv. i). 6 trop. Am., W.I. Monoec.; fls. in groups of 3, a ? between two S • O. oleracea Mart, is the cabbage palm ; the young head of 1. is cut out and eaten. The fr. yields an oil, and a form of sago is obtained from the stem (see Metroxylon). The 1. are used for thatch, &c. 0. regia H. B. et K. is the royal palm. Both are extensively used for avenues. Oreograstis K. Schum. Cyperaceae (i). i E. trop. Afr. Oreolirion E. P. Bickn. (Sisyrinchium p.p.). Iridaceae (n). 2 N. Am. Oreomitra Diels. Anonaceae (2). i New Guinea. Oreomunnea Oerst. (Engelhardtia Bff.}. Juglandaceae. i C. Am. Oreomyrrbis Endl. Umbelliferae (in. 4). 6 S. temp. Oreopanax Decne. et Planch. Araliaceae (i). 80 trop. Am. Oreophlla D. Don = Hypochoeris L. p.p. (Comp.). Oreopbylax Endl. = Gentiana L. p.p. (Gent.). Oreopolus Schlecht. (Cruckshankia p.p. EP.). Rubi. (i. 2). i Andes. Oreorcbis Lindl. Orchiclaceae (n. 5). 5 Himalaya to Japan. Oreosciadium Wecld. (Afitim p.p. EP.). Umbellif. (in. 5). 5 Andes. Oreoselinum (Tourn. ) Adans. = Peucedanum Tourn. (Umbel.). Oreoseris DC. =Gerbera Gronov. p.p. (Comp.). Oreosolen Hook. f. Scrophulariaceae (in. i). 2 Himalaya. Oreosphacus T'hil. Labiatae (vi). i Chili. Oreostemma Greene (Aster p.p.). Compositae (3). 4 N. Am. Oreostylidium Berggr. Stylidiaceae. i New Zealand. Oreosyce Hook. f. Cucurbitaceae (2). 2 W. trop. Afr. Oreotbyrsus Lindau. Acanthaceae (iv. B). 2 New Guinea. Oreoxis Rafin. (Cymopterus p.p.). Umbellif. (in. 5). i Colorado. Oresitropbe Bunge. Saxifragaceae (i). i China. Orestia Ridl. Orchidaceae (n. 4). i St Thomas I., W. Afr. Orgadium (CL), an open woodland formation. Organs, the parts of a plant regarded as performing functions. Orias Dode. Lythraceae. i Cochinchina. OR O BANG HA CEAE 4 7 3 Oricia Pierre. Rutaceae (iv). 3 trop. Afr. Orientate (Lat.), eastern. Orientation, definite position with regard to stimuli. Origanum Tourn. ex L. Labiatae (vr). 7 Eur., Medit. O. vitlgare L. (marjoram) in Brit., used as a flavouring herb. O. Majorana L. yields oil of marjoram by distillation. Orites R. Br. Proteaceae (n). 6 temp. E. Austr. Oritrephes Ridl. Melastomaceae (i). i Malay Peninsula. Orixa Thunb. \Celastrus p.p. BH.}. Rutaceae (i). i Japan. Orlaya Hoffm. (Daunts p.p. BH.}. Umbelliferae (171/2). i Medit. Orleanisia Barb. Rodr. Orchidaceae (n. 6). i Brazil. Ormocarpum Beauv. (Diphaca Lour.). Leguminosae (in. 7). 10 trop. and subtrop. |%. Ormosciadium Boiss. Umbelliferae (in. 6). i W. As. Ormosia Jacks. Leguminosae (ill. i). 25 trop. The seeds of 0. dasycarpa Jacks, (bead or necklace tree) show the same red and black surface as Abrus precatorius. Ornamental plants, whether cult, for fl., fr. or 1., or as ornamental shrubs or trees,, are very numerous, and cannot be listed. Against most important genera thus employed are placed such notes as "cult. orn. fl." So far as the cold zones are concerned they may be roughly classified into hardy pi., which will stand the winter out of doors, half-hardy, which require protection during winter, or if annual to be germinated under glass and planted out, cool house plants from warmer climates than England, succitlent house plants from dry climates, and stove plants from hot moist trop. climates. Or in another way, they may be grouped into ornamental trees, shrubs, and herbs, with the minor groups of water-plants, climbers, &c. Ornanthes Rafin. =Fraxinus Tourn. (Olea.). Ornithidium Salisb. Orchidaceae (u. 1 8). 25 trop. Am., W.I. Cult. Ornitho- (Gr. pref.), bird ; -phily, fert. by birds. Ornithobaea Parish. Gesneriaceae (i). i S. Burma. Ornithocarpa Rose. Cruciferae (3). i Mexico. Ornithocephalus Hook. Orchidaceae (n. 19). 20 trop. Am., W.I. Ornithochilus Wall, ex Lindl. Orchidaceae (n. 20). 2 E. As. Ornithogalum (Tourn.) L. Liliaceae (v). 90 temp. |*. O. iirnbel- latum L. (star-of-Bethlehem) in Brit. Ornithoglossum Salisb. Liliaceae (i). 2 S. Afr. Ornithophora Barb. Rodr. (Sigmatostalix p.p. EP.}. Orchidaceae (n. 19). i Brazil. Ornitnopus L. Leguminosae (in. 7). 8 Medit., W. As., trop. Afr., S. Brazil. O. perpusillits L. in Brit, (bird's foot). O. sativns Brot. (seradella, serratella) affords good fodder. Ornithostaphylos Small (Arctostaphylos p.p.). Ericaceae (11. 3). i Lower California. Ornithoxanthum Link = Gagea Salisb. (Lili.). Ornitrophe Comm. ex Juss. = Schmidelia L. (BH.}~ Allophylus L. Ornus Neck. = Fraxinus Tourn. (Olea.). Orobanchaceae (EP., BH.}. Dicots. (Sympet. Tubiflorae ; Personales BH.}. 12 gen., 140 sp., chiefly N. temp. %; a few Am. and trop. All are parasitic herbs with little or no chlorophyll, attached by 474 OROBANCHACEAE suckers formed upon their roots to the roots of other plants (the seeds of Orobanche only germinate when in contact with a root of a host). For details see genera. Infl. term., a raceme or spike (exc. Phelipaea, which has a sol. term. fl. ). Fl. g , -|- . K (2— 5) hypog., C (5), imbr., 2-lipped ; A 4, didynamous, epipet. ; anthers opening longitudinally ; G usu. (2), rarely (3), i-loc. Placentae parietal, often T-shaped in section or branched; ovules oo , anatr. ; style i. Loculic. caps.; seeds small, with minute undifferentiated embryo in oily endosp. Chief genera : Orobanche, Christisonia, Lathraea, Phelipaea. Orobanche (Tourn.) L. Orobanchaceae. 90 temp, and subtrop.; 7 in Brit, (broom-rape). Parasitic by their roots upon the roots of other pi.; no green tissue. O. ramosa L. is common on hemp. O. major L. (0. elatior Sutton) on Centaurea, &c. (in Brit.), O. minor Sutton on clover. Some are confined to one host, e.g. 0. Hederae Duby to ivy, others are more general in their attacks. Orobus (Tourn.) L. = Lathyrus Tourn. p.p. (Legum.). Orochaenactis Coville (Chaenactis p.p.). Compositae (6). i Calif. Orogenia S. Wats. Umbelliferae (in. 4). 2 W. N. Am. Orontium L. Araceae (in), i All. N. Am. Aquatic. Oropetium Trin. Gramineae (12). i Indomal. Orophaca Britton (Astragalus p.p.). Leguminosae (in. 6). 3 N. Am. Orophea Blume. Anonaceae (2). 32 Indomal. Orophochilus Lindau. Acanthaceae (iv. A), i Peru. Orophytium (Cl.), a subalpine plant formation. Oroxylon Vent. Bigrioniaceae (i). i Indomal. Orphanidesia Boiss. et Bal. Ericaceae (n. i). i W. As. Orphium E. Mey. Gentianaceae (i). i Cape Colony. Orpine, Sedum Telephium L. Orris root, Iris ftorentina L. Ortega L. Caryophyllaceae (i. 3). 2 Spain, Italy. Ortgiesia Regel. Bromeliaceae (4). 2 trop. Am. Orthaea Klotzsch (Thibaudia p.p. EP. ). Ericaceae (in. 2). i Peru. Orthantha Kerner. Scrophulariaceae (ill. 3). 3 Eur. , W. As. Orthanthera Wight. Asclepiadaceae (n. 3). 4 Afr., India. Orthechites Urb. (Echites p.p.). Apocynaceae (ll. i). i Jamaica. Ortho- (Gr. pref.), upright, straight; -plocous, Cruciferae; -stichies, . straight ranks ; -tropous (ov.), in a straight line with the funicle ; -tropic, placing itself in line with the stimulus. Orthocarpus Nutt. Scrophulariaceae (in. 3). 30 W. Am. Orthoceras R. Br. Orchidaceae (n. 2). i .S. E. Austr., New Zealand. Orthoclada Beauv. Gramineae (10). i trop. Am. L. petiolate. Orthogoneuron Gilg. Melastomaceae (i). i trop. Afr. Orthogynium Baill. Menispermaceae. i Madagascar. Ortliopappus Gleason (Elephantopus p-p.)- Compos, (i). i trop. Am. Ortnopenthea Rolfe. Orchidaceae (n. i). 10 S. Afr. Orthophytum Beer (Prantleia EP., Pitcairnia BH.}. Bromeliaceae (4). 2 C. Brazil. Orthopogon R. Br. = Oplismenus Beauv. (Gram.). Orthopterygium Hemsl. (Juliania p.p.). Julianiaceae. i Peru. Orthorrhiza Stapf (Chorispora p.p.). Cruciferae (4). i Persia. OSMUND A CEAE 475 Orthosia Decne. ( Vincetoxicum Moench Bff., Cynanclmm L. p.p. EP.). Asclepiadaceae (II. i). 12 S. Am. Orthosiphon Benth. Labiatae (vn). 50 Indomal., trop. Afr. Orthostemon Berg. (Feijoa p.p. BH.]. Myrtaceae (i). i subtrop. S. Am. Orthostemon R. Br. (Canscora p.p.). Gentianaceae (i). i trop. Afr. Orthotactus Nees=Dianthera Gronov. (BH.) = Justicia L. Orthrosanthus Sweet. Iridaceae (n). 7 Austr., S. Am. to Mexico. Orumbella Coulter et Rose (Ligusticum p.p.). Umbelliferae (ill. 5). i Alaska. Orychopliragmus Bunge (Moricandia p.p. BH.}. Crucif. (4). [ N. China. Oryctanthus Eichl. (LoraiitAusp.p.gff.). Loranth.(i). 10 trop. Am. Oryctes S. Wats. Solanaceae (2). i Nevada. Orygia Forsk. Aizoacae (i). i Afr. to Mysore. Oryza L. Gramineae (6). 6 trop., incl. 0. sativa L. (rice), one of the chief food plants of the world, an annual, wild in Indomal., S. Am. The cult, rices are probably derived polyphyletically from some of these. The main kinds are hill and swamp rice, the former chiefly grown by wild tribes. Swamp rice is chiefly cult, in Bengal, S. India, S.E. As., Japan, China and S. Am., and occurs in oo vars. It is cult. in shallow water till nearly ripe, when the water is drained off. The grain in the husk is known as paddy. Oryzopsis Michx. Gramineae (8). 15 N. temp. Osage orange, !\Iaclura atirantiaca JSfutt. Osbeckia L. Melastomaceae (i). 50 palaeotrop. Osbornia F. Muell. Myrtaceae (n. i). i N.E. Austr. Oschatzia Walp. (Azorella p.p. BH.). Umbelliferae (i. i). 2 Austr. Oserya Tul. et Wedd. Podostemaceae. 5 Brazil to Mexico. Osier, Salix vitninalis L. Osmanthus Lour. Oleaceae. 10 E. and S. As., Polynes., N. Am. O. fragrans Lour. (Oleafragrans Thunb.), often cult. , has ed. fr., and its 1. are used to perfume tea. Osmelia Thw. Flacourtiaceae (7). 3 Indomal. Osmia Sch.-Bip. (Eupatorinin p.p.). Compositae (2). 3 N. Am. Osmites L. Compositae (4). 6 Cape Colony. Osmitopsis Cass. Compositae (4). i Cape Colony. Osmohydropliora Barb. Rodr. Bignoniaceae (i). i Amazonas. OsmorMza Rafin. Umbelliferae (in. 2). 15 As., Am. Osmoxylon Miq. Araliaceae (i). 2 Malay Archipelago. Osmunda L. Osmundaceae. 10 temp, and trop. O. regalis L. (royal fern) in Brit, has a root-stock sometimes a foot high, like the stem of a tree fern, bearing scale 1. below the soil and ordinary 1. above. The fronds are large ( i — 10 feet) ; the lower pinnae are veg., the upper are repr. only and form a sort of panicle. They are densely covered with sori, which have no indusium and have a peculiar annulus con- sisting of a round group of cells at one side of the apex. The sporangium dehisces longitudinally. Other sp. have the fertile pinnae on the lower part of the 1., others again have separate veg. and repr. 1. Osmuudaceae. Filicales Leptosporangiatae. 2 gen., 12 sp. , trop. and temp. Short-stemmed ferns, with naked sori. The sporangia are 476 OSM UNDA CEAE shortly stalked and have an annulus, consisting of a roundish group of cells at one side of the apex ; they open by a longitudinal fissure. Genera; Osmunda (sori on special pinnae), Todea (sori on backs of ordinary pinnae). Ossaea DC. Melastomaceae (i). 55 trop. Am., W.I. Ostenia Buchenau. Butomaceae. i Uruguay. Osteocarpum F. lko&\\.(Threlkeldia BH.). Chenopod. (A). 5 Austr. Osteocarpus Phil. (Alona p.p.). Nolanaceae. 4 Chili. Osteomeles Lindl. Rosaceae (n). to Andes, Polynesia. Osteophloeum Warb. (Myristica p.p.). Myristic. i Amazonas. Osteospermum L. Compositae (9). 40 S. Afr. Osterdamia Neck = Zoysia Willd. (Gram.). Ostiolate, with a mouth. Ostodes Blume. Euphorbiaceae (A. n. 5). 6 Indomal. Ostrearia Baill. ex Niedenzu. Hamamelidaceae. i Queensland. Ostrich fern, Maltettcda Striithiopteris. Ostrowskia Regel. Campanulaceae (i). I Turkestan. Ostrya Mich, ex L. Betulaceae. 2 N. temp. Like Carpinus. 0. virginica Willd. (lever-wood) furnishes a hard wood. Ostryocarpus Hook. f. Leguminosae (ill. 8). i trop. W. Afr. Ostryoderris Dunn. Leguminosae (in. 8). 3 trop. W. Afr. Ostryopsis Decne. Betulaceae. i E. Mongolia. Oswego tea, Monarda. Osyricera Blume. Orchidaceae (n. 16). 2 Java, New Guinea. Osyridicarpos A. DC. Santalaceae. 3 Abyssinia to S. Afr. Osyris L. Santalaceae. 7 \3fc. Otacanthus Lindl. Scrophular. (n. 6), (Acanth. BH.). i Brazil. Otaheite apple (W.I.), Etigcnia malaccensis L. ; -gooseberry (W.I.), Phyllanthus distichns Muell.-Arg. Otantnera Blume. Melastomaceae (i). 10 Malaya, trop. Austr. Othake Rafin. (Gaillardia BH.}. Compositae (6). 6 S.W. U.S., Mexico. Otherodendron Makino (Elaeodendron p.p.). Celastraceae. i Japan. Othonna L. Compositae (8). 80 S. Afr. Xero. with swollen roots and often fleshy 1. Otnonnopsis Jaub. et Spach (Hertia EP.). Compos. (8). 8 Afr., W. As. Otiophora Zucc. Rubiaceae (II. 7). Madag., trop. Afr. Otocalyx T. S. Brandegee (inc. sed.). i Mexico. Otochilus Lindl. Orchidaceae (n. 3). 4 Himalaya, Burma. Otochlamys DC. Compositae (7). i Cape Colony. Otomeria Benth. Rubiaceae (i. 2). 7 trop. Afr., Madag. Otonephelium Radlk. (Nephelium p.p. BH.). Sapindac. (i). i Malabar. Otopappus Benth. (Zexmenia p.p. EP.). Compositae (.s). 6 C. Am. Otopetalum Miq. Apocynaceae (i. i). i Java. Otopetalum F. C. Lehm. et Kranzl. Orchidaceae (n. 16). i Ecuador. Otophora Blume. Sapindaceae (i). 12 trop. As. Otoptera DC (Vigna p.p. BH.). Leguminosae (in. 10). i S. Afr. Otostegia Benth. Labiatae (vi). jo W. As., Abyssinia. Otoxalis Small (Oxalis p.p.). Oxalidaceae. i Guatemala. Ottelia Pers. Hydrochariclaceae. 15 trop. and subtrop. Otto of rose, Rosa. OVARY 477 Ottoa H. B. et K. Umbelliferae (in. 4). i Mexico. Ottonia Spreng. = Piper L. p.p. (Pip.). Ottoschulzia Urb. (Poraqueiba p.p.). Icacinaceae. 3 W.I. Oubanguia Baill. Tiliaceae. 5 trop. Afr. Oudemansia Miq. = Helicteres Pluk. (Stercul.). Ougeinia Benth. Leguminosae (in. 7). i India. Ouratea Aubl. (Gomphia Bff.). Ochnaceae. 150 trop. Ourisia Comm. ex Juss. Scrophulariaceae (in. i). 20 Am., N.Z. Ourouparia Aubl. ( Uncaria Schreb.). Rubiaceae (i. 6) . 30 trop., esp. As. Outea Aubl.=Macrolobium Schreb. (Legum. ). Outfit for collecting, cf. Collecting. Ouvirandra Thou. = Aponogeton Thunb. (Apon.). Oval (1. ), widish, tapering equally to base and tip. Ovary, the hollow chamber formed by the infolded sporophylls, in which the ovules are borne on thickened placentae or cushions. It may be superior (on apex of torus above sta.) or inferior (sunk in torus below sta.) or intermediate, apocarpous (free cpls.) or syncarpous (united), in spirals or whorled, radial or-|-in symmetry. It may be unilocular ( i -chambered), or hi- tri- locular, &c. (often simply called multilcc.). The imaginary joins of the cpls. are called the sutures (ventral if at the centre), and the midrib of the cpl. is called the dorsal suture. The placenta may bear one, two or more rows of ovules, or one or two ovules only. The partitions of the ovary are called the septa, and false septa, chambering it into more loc. than normal, are found in Astragalus, Cmciferae, Gaylussacia, Linum, &c. DIAGRAMMATIC LONGITUDINAL SECTIONS OF FLOWERS, to show types of re- ceptacle, ovary, stigma, &c. A, hypogynous flower with unilocular ovary, parietal placentation, and numerous ascending anatropous ovules with raphe downwards; stigma sessile, bilobed. B, perigynous flower with multi-locular ovary, a\-\\f nlar-en- tation, and numerous horizontal anatropous ovules with raphe upwar.ls ; style with bifid stigma. C, epigynous flower with multi-locular ovary, apical placentation, and solitary pendulous anatropous ovules with ventral raphe; style long with capitate stigma, r, receptacle; k, calyx; c, corolla; a, stamens; a, ovary; st, stigma. The style, if present, may be long or short, cylindrical, filiform, ribbed, &c. ; it may be terminal, lateral, or basal. The stigma or stigmas may be sessile or on a style, simple (and then often capitate or head-like), lobed, -fid, radiate, &c. The ovule may be sessile or stalked (on a.fttntcZe); erect, ascending, 478 OVARY horizontal, or pendulous. It maybe ortho- or atropous (in a line with stalk, micropyle away from the latter), anatropous (reversed, and attached to stalk along side), amphitropous (stalk attached to middle of ovule), or campylotropous (ovule itself curved into U shape). Ovate (1.), about twice as long as broad, tapering to the tip. Overlook (W.I.), Canavalia ensiformis DC. Ovidia Meissn. Thymelaeaceae. 4 Chili. Ovieda L. = Clerodendron L. (Verben.). Ovoid, solid oval. Ovule, the possible future seed, cf. Ovary. Ovuliferous scale, cf. Coniferae. Ovum, the ? sexual cell, Angiosperm, ^ — 5, over f of the depth. Palmerella A. Gray. Campanulaceae (in). 2 Mexico, California. Palmeria F. Muell. Monimiaceae. 7 Austr., New Guinea, Celebes. Palmetto, Sabal, Thrinax; small-, Carludovica. Palmiet, Pr ionium Palmita E. Mey. Palmirah, Palmyra palm, Borasstis Jlabettifer L. Palmiste, Oreodoxa. Palmorchis Barb. Rodr. (Sobralia p.p. EP.). Orchid, (n. 7). 2 Brazil. Palmstruckia Sond. Cruciferae (i)- i S. Afr. Palovea Aubl. Leguminosae (II. 3). i French Guiana. Palta, avocado, Persea gratissima Gaertn. f. Paltonium Presl. Polypodiaceae. 2 trop. Am., China. Paludose, palustris (Lat.), of marshes. PANDANUS 485 Palumbina Reichb. f. (Oncidinni p.p. BH.). Orchid, (n. 19). i Guatem. Pamburus Swingle. Rutaceae (v. ). i India. Pampas, the grassy plains of S. temp. Am. ; - grass, Cortaderia argentea Stapf. Pampelmousse, shaddock, Citrus decumana Murr. Pamphalea DC. Compositae (12). 5 temp. S. Am. Pampliiiia Mart. Styracaceae. 3 Brazil. Panama hats, Carludovica; -rubber, Castilloa elastica Cerv., &c. Panargyrus Lag. — Nassauvia Juss. p.p. (Comp.). Panax Linn. (BH. inch Cheirodendron Nutt., Nothopanax Seem.). Araliaceae (2). 6 trop. and E. As., N. Am. For ginseng see Aralia. Pancheria Brongn. et Gris. Cunoniaceae. 10 New Caled. Pancovia Willd. (Erioglossum BH.). Sapindaceae (i). 3 trop. W. Afr. Pancratium Dill, ex L. Amaryllidaceae (i). 12 Medit., trop. As. Panda Pierre. Pandaceae. i trop. W. Afr. Fl. cyclic, dioec., hetero- chlam . G (3) , each with one pend. orthotr. ov. Drupe with 3 one-seeded chambers. Pandaceae (EP.}. Dicots. (Archichl. Pandales). Only gen. Panda, q.v. Pandales. The 22iid. order of Dicots. Archichl. Only fam. Pandaceae. Pandanaceae (EP., BH.). Monocots. (Pandanales; Nudiflorae BH.). 3 gen., 225 sp. , char. pi. of the Old World tropics, but a few warm temp. Mostly sea-coast or marsh pi. with tall stems supported upon aerial roots, frequently branched; buds are found in all axils, and the branching appears dichotomous ; some are climbers. The aerial roots have marked root-caps of membranous texture. L. in 3-ranked phyllo- taxy, but stem usu. twisted so that they appear to run in well-marked spirals, whence the name of screw-pines. L. parallel-veined, long, and narrow, with open sheath and usu. thorny margin ; generally sharply bent downwards at the middle, and corrugated like a palm 1. Infl. term., with a few bract-like 1. at the base going^ gradually over into the foliage L, usu. a racemose spadix with neither bracts nor bracteoles to the individual fls., which are somewhat difficult to make out. The ,Aizoaceae, Canna, Caryophyllaceae, Nymphaeaceae, Phytolaccaceae, Piperaceae. Periandra Mart, ex Benth. Leguminosae (in. 10). 6 Brazil. Perianth, the outer covering of the fl., composed of non-reproductive 1., usu. divided into an outer greenish whorl, the K, and an inner, coloured, the C. Concrescence is frequent, and the petals may be aborted (only recognisable for certain when many closely related forms possess them), thus giving an apctalous fl. Fl. with perianth, chlamy- deous, without, naked or achlamydcous; with one whorl, haplo- or mono-chlamydeous or ' incomplete'1 (apetaloiis if the phenomenon is clue to suppression of C), with two whorls, di- or diplo-chlamydeous or 'complete? P with 1. of one kind only (tepals] homo-chlamydeotis, of two kinds (sepals forming a calyx, and petals a corolla), heterochlamydeous. The P (K, C) may be hypo-, peri-, or epi-gynoits; of free organs {poly-phyllous, -sepalons, -petalous) or concrescent (gamo-phylloits, -sepalous, gamo- or sym-petalous] ; in the latter case the concrescent part or tithe bears the free lobes, teeth, or segments together forming the limb. It may also be regular (actinomorphic) or irregular (zygo- morphic, or asymmetrical}. If it fall as the bud opens, it is caducous, just after fert, deciduous; if it remain unwithered round the fr., per- sistent; withered, marcescent; enlarged, as in Physalis, accrescent. A homochlam. P may be sepaloid (looking like a K in colour and texture) or petaloid (like a C). The aestivation is described in the terms given under Aestivation. The sepals are commonly leafy and green, but sometimes ± woody (some Myrtaceae, &c.), or brightly coloured (Clerodendron, some Ranunculaceae, &c.). In many epig. fls. they are much reduced, e.g. in Umbelliferae and Rubiaceae. In Compositae they are frequently repres. by a pappus of hair or bristles. If concrescent, the K is de- scribed by the terms given for the C. In Malvaceae, some Rosaceae (e.g. Potentilla) and Lythraceae there is an epicalyx of apparent stipular nature (see also Dipsaceae). PERITOMA 499 The petals are usu. of some other colour than green, and of delicate texture. They may be narrowed at the base, as in wallflower, into a claw (unguiculate), fringed with hair-like teeth, as in pinks (fimbriate), bi-Jid, tri-fid, &c. (cf. 1.), or divided into several long segments (laciniate) ; they may be spurred (with long hollow projection, as in Viola), saccate or gibbous (with projecting broad pouch), scaphoid (boat-shaped, as in Loasa), &c. The general form of the sympetalous corolla may be tubular, funnel-shaped, urceolate (urn-shaped), cam- pamdate (bell-shaped as in Canterbury bells), rotate (wheel-shaped with little or no tube, as in Veronica), salver-shaped (ditto, but with long tubular portion, as in primrose), spurred, saccate, gibbous, ven- tricose (swollen out all round in the basal part); if irreg., it may be labiate or bilabiate (with two projecting lips, as in Labiatae), personate (labiate, mouth closed by projecting lobe, as in Antirrhinum, &c.), helmet-shaped, ligulate (strap-shaped, as in dandelion, &c.), &c. The shape, texture, &c. of the individual sepals, petals, calyx-lobes, corolla-lobes, are described as if they were leaves. Perianthomega Bur. (Bignonia p.p.)- Bignon. (i). i C. Brazil. Perianthostelma Baill. Asclepiadaceae (n. i). 3 trop. Afr. Peritalema DC. (Boutonia DC. EP.}. Acanthaceae (iv. A), i Madag. Periblepharis Van Tiegh. (Luxenibtirgia p.p.). Ochn. i Brazil. Pericampylus Miers. Menispermaceae. 6 E. Indomal. Perichasma Miers (Stephania p.p. BH.}. Menispermaceae. i trop. W. Afr. Perichlaena Baill. Bignoniaceae (2). i Madag. Periclymenum Tourn. ex Rupp. = Lonicera L. p.p. (Caprifol.). Pericome A. Gray. Compositae (6). 2 S. U.S., Mexico. Pericopsis Thw. Leguminosae (in. i). i Ceylon, yielding a pretty cabinet wood (nedun). Perictenia Miers (Odontadenia p.p. EP.). Apocyn. (n. i). i Peru. Peridiscus Benth. Flacourtiaceae (inc. sed.). i Braz., Venezuela. Periestes Baill. Acanthaceae (iv. B). 2 Madagascar, Comoros. Perilla Linn. Labiatae (vi). 3 India to Japan. Perillula Maxim. Labiatae (vi). i Japan. Perilomia II. B. et K. Labiatae (vi). 8 Chili to Mexico. Perinerion Baill. (Baissea A. DC. p.p.). Apocyn. (n. i). i Angola. Periodicity, usu. correlated with periodicity in climate, e.g. winter and summer, wet and dry, or hot and cool seasons. And cf. Agave, Fag us, &.c. Periornphale Baill. Gesneriaceae (inc. sed.). 2 New Caledonia. Peripeplus Pierre. Rubiaceae (n. 5). i Gaboon. Periphragmos Ruiz et Pav. (Cantua Juss.). Polemon. 2 Andes. Periploca Tourn. ex L. Asclepiadaceae (i). 12 temp. \3fc, trop. Afr. Peripterygia Loes. (Pterocelastrus^.^.). Celastr. i New Caled. Peristeria Hook. Orchidaceae (n. 13). 5 C. Am., incl. P. elata Hook, (dove orchid). Cult. orn. fl. Peristethium Van Tiegh. (Loranthus p.p.). Loranth. (i). i Andes. Peristrophe Nees. Acanthaceae (iv. B). 15 palaeotrop. VetiatylasElume(ffa6enariap.p.). Orchidaceae (ii. t). 80 warm |^. Perithrix Pierre (/{alcsanthiis EP.). Asclepiad. (i). t Cameroons. Peritoma DC. = Cleome L. (Cappar.). 32—2 500 PERITYLE Perityle Benth. Compositae (6). 15 S.W. U.S., Mexico. Periwinkle, Vinca. Perizoma Miers (Salpichroa p.p.). Solanaceae (2). i S. E. U.S. Pernambuco rubber, Hancornia speciosa Gomes. Pernettya Gaudich. Ericaceae (n. 2). 30 S. Am., Mexico, Tasm., N.Z. Pernettyopsis King et Gamble. Ericaceae (n. 2). 2 Malay Penins. Peronema Jack. Verbenaceae (5). i Malay Archipelago. Perotis Ait. Gramineae (3). 5 palaeotrop. Perotriche Cass. Compositae (4). i S. Cape Colony. Perovskia Karel. Labiatae (vi). 4 W. As. Perpusillus (Lat.), very small. Perralderia Coss. (Grantia BH.}. Compositae (4). 2 N. W. Afr. Perriera Courchet. Simarubaceae. i Madagascar. Perrieranthus Hochr. Malvaceae (4). i Madag. Perrierophytum Hochr. Malvaceae (4). i Madag. Perrottetia H. B. et K. Celastraceae. 10 Mexico, C. Am., Polynesia. Persea Plum, ex L. Lauraceae (i). 20 trop. Am. The fr. of P. gratissima Gaertn. f. (aguacate, avocado, alligator pear, palta) is ed. Persian berries, yellow berries, Rhamnus infectoria L.; -insect powder, Chrysanthemum roseum Adam.; -lilac, Melia Azedarach L. Persica (Tourn.) Mill. = Primus L. (Ros.). Persicaria (Tourn.) L. (Polygonum p.p.). Polygonaceae (n. 2). 75 N. Am. Persimmon, Diospyros virgin iana L., D. Kaki L. f., &c. Persistent (P, K, C), remaining unwithered round the fl., Corylaceae, Fagaceae, Physalis. Personales (BH.). The Qth order of Gamopetalae. Personatae (Warming). The 5th order of Sympetalae. Personate (C), labiate, mouth closed by projecting lobe, Antirrhinum. Persoonia Sm. Proteaceae (i). 60 Austr., N.Z. Pertusate, with slits. Pertya Sch.-Bip. Compositae (12). 5 Japan to Afghanistan. Peru, balsam of, Myroxylon Pereirae Klotzsch ; - bark, Cinchona • marvel of-, Mirabilis; -vian nutmeg, Lauretta. Perularia Lindl. (ffabenariap.p. BH.}. Orchid, (n. i). i N. Am., As. Pervillaea Decne. Asclepiadaceae (n. 3). i Madag. Perymenium Schrad. Compositae (5). 25 C. Am. Pescatoria Rchb. f. (Zygopetalum p.p. BH.}. Orchidaceae (n. 14). 10 Colombia. Cult. orn. fl. Peschiera A. DC. = Tabernaemontana Plum. (Apocyn.). Pessopteris Underwood (Polypodium p.p.). Polypod. i trop. Am. Pestallozzia Zoll. et Morr. =Gynostemma Blume (Cucurb.j. Petagnia Guss. Umbelliferae (in. 2). i Sicily. Petal, a 1. of the C, usu. coloured; -ody, change of sta. to pet.; -oid, petal-like ; -omania, abnormal increase of petals. Petalacte D. Don. Compositae (4). i W. Cape Colony. Petalactella N. E. Br. Compositae (4). i S. Afr. Petalidium Nees (EP. excl. Pseiidobarleria). Acanth. (iv. A), r Indomal. Petalinia Becc. (Ochanostachys p.p. EP.). Olacaceae. i Banka. PETROSELINUM 501 Petalodiscus Baill. (Savia BH.}. Euphorbiaceae (A. I. i). 5 Madag. Petalolophus K. Schum. Anonaceae (4). i New Guinea. Petalonema Gilg. Melastomaceae (i). i trop. E. Afr. Petalonyx A. Gray. Loasaceae. 4 Mexico, S.W. U.S. Petalostelma Fourn. Asclepiadaceae (n. 3). i Brazil. Petalostemma R. Br. Inc. sed. i Abyssinia. Petalostemon Michx. (Kuhnistera EP.}. Legumin. (in. 6). 25 N. Am. Petalostigma F. Muell. Euphorbiaceae (A. I. i). i E. Austr. Petalostylis R. Br. Leguminosae (n. 5). i Austr. Petasites (Tourn.) L. Compositae (8). 15 N. temp. P. .offidnalis Moench (butter-bur) in Brit, spreads largely by rhiz. It is dioecious (cf. Tussilago, its close ally). The 3 head has about 30 fls. with the usual mech. of Compositae, the style acting as pollen-presenter, though the ovary is not fertile. Occasionally a few $ fls. are found. The 9 head consists of about 150 $ fls. surrounding i — 3 c? fls. Only the male fls. secrete honey. P. fragrans Presl (winter heliotrope) cult. perf. fls., which appear in Feb. Petastoma Miers (Bignonia p.p. BH.}. Bignoniaceae (i). 8 trop. Am. Peteria A. Gray. Leguminosae (ill. 6). i New Mexico. Petermannia Y. Muell. Dioscoreaceae. i New S. Wales. Petersia Welw. Lecythidaceae. 2 W. trop. Afr. Petersianthus Merrill (Petersia Welw.). Lecythid. 3 Afr., Phil. I. Petesia P. Br. = Rondeletia L. (Rubi.). Petesioides Jacq. (Wallenia Sw. ). Myrsinaceae (11). 4 W.I. Petiole, leafstalk ; sensitive -, cf. Climbing Plants. Petitia Jacq. Verbenaceae (4). 4 Mexico, W.I. Petitmenginia Bonati. Scrophulariaceae (in. 2). i Cambogia. Petiveria Plum, ex L. Phytolaccaceae. i warm Am., W.I. Petlomelia Nieuwland (Fraxinus p.p.). Oleaceae. i N. Am. Petraeus (Lat.), growing on rocks; petrium (Cl.), a rock formation; -odium (Cl. ), a boulder field formation. Petraeovitex Oliv. Verbenaceae (5). 3 Malay Archipelago. Petrea Houst. ex L. Verbenaceae (i). 12 trop. Am., W. Ind. Cult, orn. fl., climber. Petrobium R. Br. Compositae (5). i St Helena. Petrocallis R. Br. (Draba p.p. BH.). Cruciferae (2). i Mts. S. Eur. Petrocarya Schreb. = Parinarium Aubl. (Ros.). Petrocodon Hance. Gesneriaceae (i). i Chili. Petrocoptis A. Br. (Lychnis p.p. BH.}. Caryophyll. (n. i). 2 Pyrenees. Petrocosmea Oliv. Gesneriaceae (i). 3 China. Petrollinia Chiov. (Iniila p.p.). Compositae (4). i E. trop. Afr. Petromecon Green (Eschscholtzia p.p.). Papaver. (n). 2 Calif. Petronia Barb. Rodr. (Promenaea p.p. EP.}. Orchid, (n. /3. n). i Braz. Petrophila R. Br. Proteaceae (i). 35 Austr. Petrophyes Webb et Berth. = Monanthes Haw. (Crassul.). Petrophyton Rydberg (Spiraea p.p.). Rosaceae (i. i). 5 N. Am. Petrosavia Becc. Liliaceae (i). i Borneo. Petroselinum Hoffm. (Canim p.p. BH.}. Umbelliferae (in. 5). 5 Eur., Medit. P. sativum Hoffm. is the parsley, cult, as condiment. 502 PETROSIMONIA Petrosimonia Bunge. Chenopodiaceae (B). 7 Greece to C. As. Petrusia Baill. Olacaceae. i Madagascar. Petteria C. Presl. Leguminosae (in. 3). i S.E. Eur. Petty whin, Genista anglica L. Petunga DC. Rubiaceae (i. 8). 6 E. Indomal. Petunia Juss. Solanaceae (4). 15 S. and warm N. Am. P. vio- lacea Lindl. and others often cult. orn. fl. Peucedanum (Tourn.) L. (BH. incl. Anethiim Tourn., Pas/i/iaca L.). Umhelliferae (ill. 6). 180 Eur., As., Afr., Am.; 4 in Brit. P. sativum Benth.-et Hook. f. (parsnip), cf. Pastinaca. P. (A.) 'graveolens Benth. et Hook. f. (Medit.) is the dill; fr. a condiment; P. offidnale L. (Brit.) is the sulphur-root used in veterinary practice; P. OstriitJiiiim Koch (Brit.) is also used. Peucephyllum A. Gray. Compositae (S). i S.W. U.S. Peumus Molina. Monimiaceae. i Chili, P. Boldus Molina, the Boldo. Wood hard ; bark yields dye; fr. ed. Peutalis Rafin. = Polygonum Tourn. (Polyg.). Peyrousea DC. Compositae (7). i Cape. Pezisicarpus Vernet. Apocynaceae (n. i). i Siam. Pfaffia Mart. Amarantaceae (3). 20 warm S. Am. Pfeiffera Salm-Dyck (Rhipsali's p.p.). Cact. (in. 3). i S. Braz. Phaca L. (Astragalus p.p.). Leguminosae (in. 6). 60 N. Am. Phacelia Juss. (incl. Cosniantlnts Nolte, Entoca R. Br., Whitlavia Harv.). Hydrophyllaceae. 100 N. Am., Andes, often cult. orn. fl. The fl. is a bee-flower with honey secreted below the ovary and guarded by stipule-like flaps at the base of the sta. The large- flowered sp. are highly protandrous. The anther as it dehisces turns inside out. Phacellanthus Sieb. et Zucc. Orobanchaceae. i Japan. Phacellaria Benth. Santalaceae. 2 Further India. Phacellothrix F. Muell. Compositae (4). i E. trop. Austr. Phacelophrynium K. Schum. (Phrynium p.p.). Marantaceae. 6 Ma- laya. Phacopsis Rydberg (Astragalus p.p.). Legum. (in. 6). 2 N. Am. Phaeanthus Hook. f. et Thorns. Anonaceae (2). 7 Malaya. Phaecasium Cass. (Crepis p.p. EP.~). Compositae (13). 3 Eur., W. As. Phaedranassa Herb. Amaryllidaceae (i). 4 Andes. Phaedranthus Miers (Bigiwnia p.p. BH.}. Bignoniaceae (i). i Mex. Phaenixopus Cass. =Lactuca Tourn. p.p. (Comp. ). Phaenocoma D. Don. Compositae (4). i Cape Colony. Cult. orn. fl. Phaenogams, Phanerogams, Spermaphyta. Phaenohoffmannia O. Ktze. (Fleiospora Harv.). Leguminosae (in. 3). i S. Afr. Phaenology, study of the periodic phenomena of vegetation. Phaenopyrum M. Roem. = Crataegus ' Tourn. (BH.) = Cotoneaster Rupp. Phaenosperma Munro ex Benth. Gramineae (4). i China. Phaeocephalus Sp. Moore. Compositae (7). i Cape Colony. Phaeomeria Lindl. (Amomump.p.ffff.). Zingiber. (i). 16 Indomal. Phaeoneuron Gilg. Melastomaceae (i). 4 trop. Afr. Phaeopappus Boiss. =Centaurea L. p.p. (Comp.). FHELLANDRIUM 503 Phaeoptilum Radlk. Nyctaginaceae. i S. Afr. Phaeospheriona Hassk. (Athyrocarpus BH.). Commelin. 4 trop. Am. Phaeostemma Fourn. Asclepiadaceae (11. 4). 2 Brazil. Phagnalon Cass. Compositae (4). 20 Canaries to Himalaya. Phaiocalanthe x Rolfe. Orchidaceae. Hybrid, Phaius-Calanthe. Phaiocymrjidium x . Orchidaceae. Hybrid, Phaius-Cymbidium. Phaius Lour. (P/iajns Hassk.). Orchidaceae (n. 9). 16 trop. As., Malaya, Austr., &c. Terrestrial. Phalacraea DC. = Piqueria Cav. (Comp. ). Phalacrocarpum Willk. (Chrysanthemum p.p. BH.). Comp. (7). i Spain. Phalacroderis DC. (Rodigia EP.). Compositae (13). i Aegean. Phalacrodiscus Bess. = Chrysanthemum Tourn. p.p. (Comp.). Pnalacroceris A. Gray. Compositae (13). i California. Phalaenopsis Blume. Orchidaceae (n. 20). 40 Indomal. Epiph. with flattened aerial roots. Phalangium (Tourn.) Adans. =Anthericum L. p.p. (Lili.). Phalaris L. Gramineae (7). 10 Eur. P. canariensis L. (canary grass) seeds are used for cage-birds. P. ariutdinacea L. (reed-grass) is com- mon in Brit. Phaleria Jack. Thymelaeaceae. 12 Indomal. Phanera Lour. =Bauhinia L. p.p. (Leguni.). Phanerogamae, see Spermaphyta. Phanerophleoia Presl. Polypodiaceae. 10 trop. Am. Phania DC. Compositae (2). 3 W.I. Pnanopyrum Nash (Paiiicinii p.p.). Gramineae (5). i N. Am. Pliarbitis Choisy (Ipomoea p.p. BH.}. Convolvulaceae (i). 6 trop. and subtrop. Cult. orn. fl. Pharetranthus F. W. Rlatt (Coreopsis p.p. EP.). Comp. (5). i Phil. Pharmacosycea Miq. = Ficus Tourn. p.p. (Mor. ). Pharnaceum L. Aizoaceae (i). 16 S. Afr. Pharus P. Br. Gramineae (6). 5 trop. Am. Phaseolus (Tourn.) L. Leguminosae (m. 10). 160 trop. and warm temp. Fl. mech. like Vicia, but complicated by the spiral coiling of the keel with the inclosed style. P. inultiflorus Willd. (Mexico) is the scarlet-runner, P. vulgaris L. the French or kidney bean, P. lunatus L. the Lima or duffin bean similarly used in the trop., P. aciitifolius A. Gray the tepary of the S.W. U.S., P. Mungo L. (P. Max L.) the 'green gram' of India, used like kidney beans, or roasted, and as horse food. The var. radiatus of the last, with darker beans (black gram), is a highly valued pulse and horse food in India. Phaulanthus K.idl.(Anertncleistttsp.'p.). Melastomaceae (i). 6 Malay Peninsula, Assam. Phaulothamnus A. Gray. Phytolaccaceae. i North Mexico. Pliaylopsis Willd. (Micranthus EP.). Acanthaceae (iv. A). 12 pa- laeotrop. Pheasant's eye, Adonis autumnalis L. Phebalium Vent. Rutaceae'(i). 35 Austr., N.Z. Phegopteris Fee = Dryopteris Adans. (Polypod.). Phellandrium (Tourn.). L. =Oenanthe Tourn. p.p. (Umbellif.). 504 PHELYPAEA Phelypaea Tourn. ex L. Orobanchaceae. 2 Cent. As. Phelline Labill. Aquifoliaceae. 12 New Caledonia. Phellium (Cl.), a rock field formation. Phellodendron Rupr. Rutaceae (iv). 4 E. temp. As. Phellogen, formative tissue of cork (bark). Phellolopliium Baker. Umbelliferae (in. 5). i Madag. Phellopterus Benth. in B. et H. f. =Glehnia F. Schmidt (Umb. ). Phellopterus Nutt. (Cymopterus Rafin. p.p.)- Umbelliferae (in. 6). 4 E. As., W. N.Am. Phenax Wedd. Urticaceae (3). 12 trop. Am. Pberosphaera Archer (Dacrydium p.p.). Coniferae (Taxaceae i ; see C. for gen. chars.), i Austr., i Tasmania. Shrubs. PherotricMs Decne. Asclepiadaceae (n. 4). 2 Mexico. PMalacanthus Benth. Acanthaceae (iv. B). i E. Bengal. PMalanthus Griseb. Rubiaceae (n. 3). 4 W.I. Phialocarpus Deflers. Cucurbitaceae (2). i Arabia. Phialodiscus Radlk. (Blighia p.p.). Sapindaceae (i). 5 trop. Afr, PMlactis Schrad. Compositae (5). 2 Mexico. Philadelphia (Riv.) L. Saxifragaceae (in). 20 N. temp. Shrubs with opp. 1. ; the buds arise closely protected by the 1. -bases through which in many they have to break. Fls. conspicuous, strongly scented, protogynous. Sta. 20 — 40; ovary inf., usu. 4-loc. Several sp. cult, orn. fl. shrub (Syringa). Philastrea Pierre. Sterculiaceae. i Cambodia. PMIbornea Hallier (Durandea p-p.). Linaceae. 2 Borneo. PMlesia Comm. ex Juss. Liliaceae (x). i S. Chili, a much-branched shrub with petioled, i-nerved, rolled-back 1., not easily recognised as a Monocot. Philgamia Baill. Malpighiaceae (inc. sed.). i Madag. Philibertella Vail. (Philibertia p.p.). Asclep. (n. i). 10 N. Am. Philibertia H. B. et K. Asclepiadaceae (n. i). 35 Am. Philippia Klotzsch. Ericaceae (iv. 2). 20 S. Afr., Madag., Masc. Philippiella Spegazz. Caryophyllaceae (i. 4). i Patagonia. Phillipsia Rolfe (Satanoi rater p.p. JSP.). Acanthaceae (iv. A), j Somaliland. Phillyrea L. Oleaceae. 6 Medit. Phillyrophyllum O. Hoffm. Compositae (4). i Kalahari desert. Philodendron Schott. Araceae (v). 225 trop. Am. Most are shrubs, usu. climbing, often epiph., with both clasping roots and aerial roots reaching the soil (see fam.). The latter sometimes twine as they descend. The pinnation of the 1. is due to a delayed development of the portions between the ribs, and not to a process such as occurs in Monstera (q.v.). Monoecious. PMlodice Mart. Eriocaulonaceae. 2 Brazil. Philoglossa DC. Compositae (5). 2 Peru, Ecuador. Philogyne Salisb. = Narcissus Tourn. p.p. (Amaryll.). Philonotion Schott. Araceae (v). i Amazonas. Philotheca Rudge. Rutaceae (i). 5 Austr. PMlotria Rafin. (Elodea p.p.). Hydrocharidaceae. 5 N. Am. -philous (Gr. suff.), loving, dwelling in. Philoxerus R. Br. (Iresine p.p. EP.). Amarant. (3). 10 trop. exc. As. PHORMIUM 505 Philydraceae (EP., F>H.}. Monocots. (Farinosae; Coronarieae./5/f.). 3 gen., 4 sp. , Indomal., Austr. Herbs witli 2-ranked sheathing narrow 1. and fls. in spikes, 5 , -|- , homochlam. A i, ant. 0(3), style i, plac. axile or parietal, with oo anatr. ov. Caps. Endosp. Philydrum Banks. Philydraceae. i E. As., Austr. Philyrophyllurn O. Hoffm. Conipositae (4). i Kalahari desert. Phinaea Benth. Gesneriaceae (11). 4 Colombia. Fhippsia R. Br. Gramineae (8). r arctic, r Indomal., i Argentina (if. Cryptotaenia). Phitopis Hook. f. Rubiaceae (i. 3). i Peru. Phlebanthia Reichb. Caryophyllaceae (inc. sed.). Nomen. Phlebocalymna Griff, ex Miers (Gonocaryiun EP.). Olacaceae. 4 E. As., Austr. Phlebocarya R. Br. Amaryllidaceae (in) (Haemod. BH.). 3 W. Austr. Phlebochiton Wall. (Tapiria BH., Pegia EP.). Anacardiaceae (2). i E. Himalaya. Phlebolithis Gaertn. Sapotaceae (inc. sed.). i, habitat? Phlebotaenia Griseb. (Polygala p.p. EP.). Polygalaceae. i Cuba. Phleum L. Gramineae (8). 10 temp, except Austr. P. prate use L., timothy-grass (Brit.), is a valuable fodder. Phloem, the outer part of the vascular bundle, carrying prepared food. Phloga Nor. ex Thou. Palmae (iv. i). 2 Madag. Phlogacanthus Nees. Acanthaceae(iv. B). 15 Indomal. Cult. orn. fl. Phlogella Baill. Palmae (iv. i). i Comoros. Phlomis L. Labiatae (vi). 65 Medit. to China. The upper lip of the fl. is raised by an entering insect. Phlox L. Polemoniaceae. 50 N. Am., Siberia. Cult. orn. fl. Phoberos Lour. = Scolopia Schreb. (Flac.). Phocea Seem. Celastraceae. i New Caledonia. Phoebanthus Blake (Helianthdla p.p.). Compositae (5). i N. Am. Phoebe Nees. Lauraceae (i). 55 Indomal., trop. Am., W.I. Phoenicanthemum Blume = Loranthus L. p.p. (Loranth.). Phoenicaulis Nutt. (Clieiranthus^.^.BH.). Cruciferae(3). i Pacific Am. Phoeniceus (Lat), scarlet. Phoenicophorium H. Wendl. (Stevensonia BH.}. Palmae (iv. i). t Seychelles. Phoenicospermum Miq. (Sloanca p.p. EP.). Elaeocarp. i Java. Phoenix L. Palmae (i. i). 12 trop. Afr., As., incl. P. dactylifera L. (date palm, Afr., S.W. As.). It has a columnar stem covered with old 1. -bases; the 1. are pinnate. Fls. dioec. ; the Arabs fert. the ? spadix by hanging a ,). i S. Am. Phrodus Miers. Solanaceae (2). 4 Chili. Phryganocydia Mart. {MacfadyenaBH.}. Bignoniaceae(i). i E. S. Am. Phrygilanthus Eichl. (Loranthns p.p. BH.). Loranthaceae (i). 20 S. Am., E. Austr. Phrygiobureaua O. Ktze. = rhryganocydia Mart. (Bignon. ). Pliryma L. Phrymaceae. i E. As., N. Am. Herb with opp. 1. and small axillary fls., •]-. Cpl. i with one erect orthotr. ov. Phrymaceae (EP.; Verbenaceae p.p. BH.}. Dicots. (Sympet. Tubiflorae). Only gen. Phryma, q.v. ; the chief distinction from Verbenaceae is the erect orthotr. ovule ; no transitions between this and other V. Phrynium Loefl. Marantaceae. 12 Indomal. Phtheirospermum Bunge. Scrophulariaceae (in. 3). 4 E. As. Phthirusa Mart. (Loranthns p.p. BH.}. Loranth. (i). 35 trop. Am. Phuodendron Graebn. Valerianaceae. i Brazil. Phuopsis Benth. et Hook. f. Rubiaceae (n. n). i Caucasus. Phycella Lindl. = Hippeastrum Herb. p.p. (Amaryll.)- Phygelius E. Mey. Scrophulariaceae (n. 4). 2 S. Afr. Cult. orn. fl. Phyla Lour. (Lippia p.p.). Verbenaceae (i). 6 N. Am. Phylacium Bennett. Leguminosae (in. 7). i Malay Archipelago. Phylica L. Rhamnaceae. 70 S. Afr., JMadag., &c. Mostly xero. shrubs, often of heath-like habit with 1. rolled back (cf. Empetrum). Phyllacantlia Hook. f. Rubiaceae (i. 8). i Cuba. Phyllachne Forst. Stylidiaceae. 9 Tasm., N.Z., S. Am. Phyllactinia Benth. (Pasaccardoa EP.}. Compositae (12). i trop. Afr. Phyllactis Pers. (Valeriana p.p. EP.}. Valerian. 25 Mexico, S. Am. Phyllagathis Bliune. Melastomaceae (i). 5 S.E. As. Phyllanthera Blume. Asclepiadaceae (i). 2 Malay Penins. , Java. Phyllanthodeudron Hemsl. (Phyllanthus p.p. EP.}. Euphorbiaceae (A. i. i). 2 Siam. Phyllanthus L. Euphorbiaceae (A. i. i). 460 temp, and trop., exc. Eur. and N. As. The trop. Am. § Xylophylla has flat green phyllo- clades bearing fls. on the margins. The ultimate shoots in § Eu- phyllaiithus look like pinnate 1. In P. cyclanthera Baill. the S fl. has its 3 sta. united into a synandrium with ring-like anther at top. PHYLLO- 5°7 Phyllarthron DC. Bignoniaceae (4). 6 Madag. , Mascarenes. The 1. is reduced to a jointed winged petiole. Phyllepidum Rafin. Amarantaceae (inc. sed.). Gen. dubium. i N. Am. Phyllis L. Rubiaceae (n. 7). i Canaries, Madeira. Phyllitis Ludwig. Polypodiaceae. 10 trop. and subtrop. Phyllo- (Gr. pref.), -phyllous (suff. ), leaf; -clade, a stem structure usu. ± flattened and serving 1. purposes, Asparagus, Baccharis, Bos- siaea, Carmichaelia, ffibbertia, Lathyrus, Leinna, Miiehlenbeckia, Oxalis, Phyllanthus, Phyllocladns, RHSCHS, Semele; -de, a petiole flattened and green, taking over 1. -functions, Acacia, Oxalis; -dy, change of fl. organs to 1. ; -me, a leaf structure; -taxy, the arrange- ment of the 1. upon the stem. It follows definite rules, esp. in flowering-plants, though it varies within certain narrow limits. The 1. may be several at each node (in whorls), or two (usu. opposite), or one (alternate). When the stem is so short that the 1., as in the primrose or dandelion, are all crowded together and spring from the level of the ground, they are said to be ' radical. ' In the first two cases the I. at one node usu. stand above the gaps at the node below. In the case of alt. 1. there is found to be a fairly constant angle between each 1. and the next one above it, e.g. in Plantago (fig.) this angle is f of the whole circumference measured the nearest way. This fraction f represents the phyllotaxy. Twist- ing excepted the 1. will stand in 8 vertical rows, each divided from the next by \ of the circum- ference. Leaf 2 will be f , 3 will be f , 4 will be f, 5 will be V-> 6 will be Y-, 7 will be V, 8 will be -V~> and 9 will be -\4-, of the circumference from leaf i, i.e. immediately over it, and three turns of the spiral aboveit. Hence the rule for determining phyllotaxy: start from any 1. A and draw a spiral round the stem, passing by the nearest way through all consecutive 1. to the 1. B exactly above A; then the number of 1. from A to B is the denominator, the number of turns of the spiral the numerator, of the fraction representing the phyllotaxy. In Gramineae the phyllotaxy is 5, i.e. alt. on opp. sides of the stem, in Cyperaceae \. Nearly all other actual arrangements are terms of the continued fraction starting from |, g. If we add the numerators together to make a new numerator, and treat the denomi- nators in the same way, we get the next arrangement f . This with \ gives f , and then fa and so on. It is rare to find a stem that shows PHYLLOTAXY. 508 PHYLLO- the phyllotaxy very clearly ; usually in the course of growth more or less twisting occurs (cf. Pandanaceae). The benefit of the phyllotaxy is that the leaves are spread out to occupy the available space to advantage. Phyllotaxies of these types give shoots of radial symmetry ; there are also bilateral arrangements, esp. upon horizontal shoots. Some- times the dorsiventrality is attained by the twisting of the leaf-stalks from the positions in which they arose, but more commonly there is a more or less tsvo-ranked (distichous) phyllotaxy, the 1. arising upon the sides of the axis, and merely having to twist at their bases to place themselves horizontally ; e.g. in the yew (Taxus), lime (Tilia), Abies, Anona, Betulaceae, Pinus, Ulmus, &c. In other plants, again, alterations of phyllotaxy occur for which no explanation can be given ; e.g. in Baptisia, Eucalyptus, Nolana- ceae, Quisqualis, Solanaceae, Thelygonum, &c. Phylloboea Benth. Gesneriaceae (i). 2 Further India, China. Phyllobotrium Muell.-Arg. Flacourtiaceae (10). i Gaboon. Phyllocactus Link. (EP. incl. Epiphylhim p.p.). Cactaceae (in. i). i6trop. Am., W.I. , often epiphytic. Plat-stemmed plants (seefam. ). Phyllocalyx Berg. = Eugenia Mich. p.p. (Myrt.). Phyllocarpus Riedel ex Endl. Leguminosae (n. 8). Rio de Janeiro. Phyllochlamys Bur. in DC. Moraceae (i). i Indomal. Phyllocladus Rich. Coniferae (Taxaceae, 6; see C. for gen. char.). 6 Tasm., N.Z., Borneo (celery pine). The 'short shoots' are repre- sented by flat green leaf-like structures — phylloclades — whose stem- nature is easily recognized by their position in the axils of the scale 1. on the 'long shoots.' The edges of the phylloclades also bear scales. The fls. (mon- or di-oec.) occupy the position of phylloclades. Each cpl. has one axillary erect ovule. The seed has a small basal aril. The timber is useful : the bark of P. trichomanoides D. Don is used for tanning. Phylloclinium Baill. Flacourtiaceae (10). i Congo. Phyllocomos Mast. Restionaceae. i S. Afr. Phyllocosmus K\otzsch(Oi://t/iocosmus£P.). Linaceae. 5trop. Am., Afr. Phyllocrater Wernham. Rubiaceae (i. 2). i Borneo. Phylloctenium Baill. Bignoniaceae (4). i Madag. Phyllodes Lour. = Phrynium Loefl. (Marant). Phyllodium. Desv. = Desmodium Desv. (Legum.). Phyllodoce Salisb. (Bryanthus BH.}. Ericaceae (i. 3). 10 N. cir- cumpolar and temp. Phylloglossum Kunze. Lycopodiaceae. i Austr. and N.Z., P. Dntni- mondii Kunze. The embryo forms a protocorm (see fam.), which produces a crown of sterile 1. and a short unbranchecl stem, bearing at its apex a single cone of sporangia, like the cone of Lycopodium. "At the end of the growing season a new protocorm is formed. This arises directly from the apex of the old one where no strobilus is developed, but in the latter case grows out upon a sort of peduncle from near the base of one of the 1." (Campbell). Prothallus sub- terranean. [Cf. Wernham in Ann. Bot. 1910, p. 335.] Phyllogonum Coville. Hydnoraceae. i California. Phyllomelia Griseb. Rubiaceae (ll. 4). i Cuba. PHYSOSTEGIA 509 Phyllonoma Willd. ex Schult. Saxifrag. (v). i Mex. to Colombia. Phyllophiorrhiza O. Ktze. = Ophiorhizophyllum S. Kurz (Acan.). Phyllopodium Benth. Scrophulariaceae (n. 5). 158. Afr. Pnyllorhachis Trimen. Gramineae (5). i Angola. Phylloscirpus C. B. Clarke. Cyperaceae (i). i Argentina. Phyllosma Bolus. Rutaceae (i). i S. Afr. Phyllospadix Hook. Potamogetonaceae. 2 W. coast N. Am., Japan. Dioecious. Phyllostachys Sieb. et Zucc. Gramineae (13). 6 Japan to Himalaya. The stripped stems are Whangee canes. Phyllostegia Benth. Labiatae (ill). 17 Hawaii, Tahiti. Phyllostylon Capanema ex B. et H. f. Ulmaceae. i Rio, i Paraguay, i Cuba. Cf. Cryptotaenia. Good timber. Phyllota Benth. Leguminosae (ill. 2). 10 Austr. Phyllotricrium Thorel ex Lecomte. Sapind. (inc. sed.). i S.E. As. Phylloxera, the root and leaf louse of the vine (Vitis). Phylloxylon Baill. Leguminosae (in. 8). 2 Madag., Mauritius. Phylogeny, line of descent from other forms. Phylon (Gr.), line of descent. Phymaspermum Less. Compositae (7). 5 Cape Colony. Phymatidium Lindl. Orchidaceae (n. 19). 2 Brazil. Phymatocarpus F. Muell. Myrtaceae (n. i). 2 W. Austr. Phymatodes Presl = Poly podium L. (Polypod.). Physacanthus Benth. Acanthaceae (iv. A). 3 W. trop. Afr. Physaliastrum Makino (Chamaesaracha p.p.). Solan. (2). 2 Japan. Physalidium Fenzl. Cruciferae (2). t Persia. Physalis L. Solanaceae (2). 50 cosmop. The berry of P. Alkekengi L. (winter cherry) is edible, also that of P. peruviana L. (strawberry or gooseberry tomato, or cape gooseberry). It is enclosed in the bladdery persistent calyx, which becomes red. Physaria A. Gray. Cruciferae (3). 6 Pac. N. Am. Physedra Hook. f. Cucurbitaceae (3). 5 trop. W. Afr. Physena Nor. ex Thou. Flacourtiaceae (inc. sed.). 2 Madag. Physic-nut, Jatroph a Ciircas L. Physocalymma Pohl. Lythraceae. i trop. S. Am. Timber valuable. Physocalyx Pohl. Scrophulariaceae (in. 2). 2 Brazil. Physocarpus Maxim, (-pa Rafin.) (Neillia BH.}. Rosaceae (i. i). 3 N. Am., N.E. As. Physocaulis Tausch (Chaerophyllnni p.p. BH}. Umbelliferae (in. 2). i Medit. Physochlaina G. Don. Solanaceae (2). 5 C. As. Physodium Presl. Sterculiaceae. 2 Mexico. Physoleucas Jaub. et Spach (Lencas p.p. BH.}. Labi.(vi). i Arabia. PhysoloMum Benth. = Kennedya Vent. p.p. (Legum.). Physopodium Desv. Lythraceae. i Bourbon. Physopsis Turcz. Verbenaceae (3). i W. Austr. Physoptychis Boiss. ( Vesicaria BH.}. Cruciferae (4). i Persia. Physorrhynchus Hook. Cruciferae (2). 2 Persia, Afghanistan. Physosiprion Lindl. Orchidaceae (n. 8). 5 trop. Am. Cult. orn. fl. Physospermum Cusson. Umbelliferae (ill. 4). 5 Eur. Physostegia Benth. Labiatae (vi). 5 N. Am. 5 1 o PHYSOSTELMA Physostelma Wight. Asclepiadaceae (n. 3). 2 Malaya. Physostemon Mart. (Cleome p.p. BH.}. Capparid. (v). 4 trop. S. Am. Physostigma Balf. Leguminosae (in. 10). 2 trop. Afr. The keel is spurred. P. -venenosuni Balf. is the ordeal bean of Calabar. Physotrichia Hiern. Umbelliferae (in. 5). 2 trop. Afr. Physurus L. C. Rich. Orchidaceae (n. 2). 25 warm As., Am. Phyt-, Phyto- (Gr. pref.), plant-; -logy, botany. Phytelephas Ruiz et Pav. Palmae(v). 4 trop. Am. Like Nipa, widely different from other palms; with affinities to Pandanaceae and Cyclan- thaceae. Short-stemmed with large pinnate rad. 1., and dioec. infls. c? infl. a sausage-shaped spadix ; the fl. has an irreg. P and oo sta. with long filaments. ? spadix simple with spathe of several 1., and about 6 fls.; the fl. has an irreg. P (an outer whorl of 3 and inner of 5 — 10 longer 1.), numerous stds. and usu. a 5-loc. ov. with long style and stigmas. Each fl. gives a berry, and the actual fr. consists of 6 or more of these united together. The outer coat is hard, with woody protuberances. Each partial fr. contains several seeds; the endosp. (cellulose) is very hard (vegetable-ivory) and is used for turning into billiard balls, £c. [Compare this fr. with Pan- danus and Carludovica.] Phyteuma L. Campanulaceae (i. i). 45 Medit., Eur., As. P. orbi- culare L. and P. spicatitin L. (rampion) in Brit. Fl. mech. interest- ing (see fain. ). The fls. are small, and massed together in heads. A tube is formed by the coherence of the tips of the long thin petals, within which the anthers are held. The style pushes up through this and drives the pollen gradually out at the end, where it is exposed to insects. Finally the style emerges, the stigmas open and the petals separate and fall back. [Compare with Campanula, Jasione and Compositae.] Phytocrene Wall. Icacinaceae. 7 Indomal. Twining shrubs with very large vessels in the stem. If the stem be cut a quantity of water escapes, which is drunk by the Malabar natives. Fls. dioec. Phytolacca Tourn. ex L. (incl. Pircunia Bert.). Phytolaccaceae. 26 trop. and subtrop. Herbs with fleshy roots, or shrubs or trees. Fls. reg. ; P 5, A 10 — 20, G 7 — 10 or (7 — 10); in the latter case fr. a berry, in the former an aggregate of achenes or drupes. Phytolaccaceae (EP. , BH.). Dicots. (Archichl. Centrospermae; Cur- vembryae BH.}. 20 gen., 60 sp., chiefly trop. Am. and S. Afr. Herbs, shrubs, or trees, with racemose or cymose infls. of_ regular inconspic. $ fls. P 4 — 5, A 4 — 5 or more (to oo ), G, rarely G, i — oo or (i — oo ), ovules i in each cpl., amphi- or campylo-tropous. Drupe or nut, rarely capsule. Seed with perisperm, often arillate. The fls. exhibit great variety in structure, owing to branching of sta. and different numbers and arrangements of cpls. Chief genera: Seguiera, Rivina, Phytolacca. Phytosalpinx Lunell = Lycopus Tourn. (Labi.). Piaranthus R. Br. Asclepiadaceae (n. 3). 128. Afr. Piassaba fibre, Attalea fun if era Mart. (Bahia), Leopoldinia Piassaba Wallace. Picardaea Urb. Rubiaceae (i. i). i Haiti. Picconia DC. (Notelaea p.p.). Oleaceae. i Canaries, Madeira. PILIFEROUS 511 Picea Link. Synonymy : P. vulgaris Link (Finns Abies L.) = P. excelsa Link; P. canadensis Link = Tsuga canadensis; P. Pinsapo Loud. = Abies Pinsapo; P. rn'ora A. Dietr. =P. nigra Link. Coniferae (Pinaceae 13; see C. for gen. char.). 35 j&. Long shoots only with needle 1. Fls. single. Cones ripening in one year. P. excelsa Link, the Norway spruce or spruce-fir, found in Eur. from the Pyrenees to 68° N., furnishes valuable wood, resin, and turpentine. P. alba Link (silver fir, N. Am.), P. Morinda Link (Himalayan spruce), and others are also valuable. Pichleria Stapf et Wettst. Umbelliferae (in. 5). 2 Persia. Pichonia Pierre (Epiluma p.p. EP.). Sapotaceae (i). 2 New Caled. Pickerel weed (Am.), Pontederia. Pickeringia Nutt. ex Torr. et Gray. Legumin. (in. 2). i Calif. Picotee, var. of carnation, Dianthns Caryophyllns L. Picradenia Hook. (Actinella p.p.). Compositae (6). 20 N. Am. Picradeniopsis Rydberg (Bahia p.p.). Compositae (6). i N. Am. Picraena Lindl. (Picrasma p.p. EP.). Simarubaceae. 2 W.I. , Brazil. Picralima Pierre. Apocynaceae (i. 3). i Gaboon. Picramnia Sw. Simarubaceae. 30 trop. Am., W.I. Picrasma Blume. Simarubaceae. 8 trop. and subtrop. The bitter wood and bark are used as a substitute for quassia. Picrella Baill. Simarubaceae. i Mexico. Picridium Desf. (Rcichardia EP.}. Compositae (13). 10 Medit. Picris Linn. Compositae (13). 36 Medit., W. As., Abyss. 2 in Brit. Picrocardia Radlk. Simarubaceae. i New Caled. Picrodendron Planch. Simarubaceae. i W.I. Picrolemma Hook. f. Simarubaceae. i Amazonas. Picrorrhiza Royle ex Benth. Scrophulariaceae (in. i). i Himal. Picrosia D. Don. Compositae (13). i warm S. Am. Pictetia DC. Leguminosae (in. 7). 6 W.I., Mexico. Pictus (Lat.), coloured. Piddingtonia A. DC. = Pratia Gaudich. (Campan.). Pierardia Roxb. = Baccaurea Lour. p.p. (Euph.). Pieris D. Don (Lyonia p.p. EP.). Ericaceae (11. i). 10 N. Am., E. As. Pierrea Hance. Dipterocarpaceae. r Malaya. Pierreanthus Bonati = Delpya Pierre (Scroph.). Pierreodendron Engl. Simarubaceae. i trop. Afr. Pierrina Engl. Scytopetalaceae. 2 trop. Afr. Pig lily (Afr.), Zantedesc/iia; - nut (Am.), Carya pordna Nutt., (W.I.) Omphalea; - weed (Am.), Chenopodnim ; -'s balsam, Hedivigia. Pigafettia Becc. Palmae (in). 3 Malay Archipelago. Pigea DC. = IonicliunV Vent. (BH. ).= Hybanthus Jacq. (Viol.). Pigeon berry (Am.), Phytolacca ; -pea, Cajanns indie us Spreng. Pignons, ed. seeds of Phms Pinea L. Pilea Lindl. Urticaceae (2). 140 trop., cult, (artillery plant), so called from the puffs of pollen ejected by the exploding sta. (cf. Urtica). Pileanthus Labill. Myrtaceae (11. 2). 3 W. Austr. Pileostegia Hook. f. et Thorns. Saxifragaceae (in), i S.E. As. Pileus Ramirez. Passifloraceae. i Mexico. Pilewort, Ranunculus Ficaria L. Piliferous, hair-bearing. 5i2 PILIOCALYX Piliocalyx Brongn. et Gris. Myrtaceae (i). 4 New Caledonia. Pillansia L. Bolus. Iridaceae (ill), i S. Afr. Pillar roots, cf. Aerial roots. Pillwort, Pilularia. Pilocarpus Vahl. Rutaceae (i). 12 trop. Am., W.I. The leaves of P. pennatifolius Lem. are the officinal 'folia Jaborandi.' Pilocereus Lem. (Cemis p.p. BH.}. Cactaceae (in. i). 50 Mex., S. Am. Pilogyne Schrad.^Melothria L. p.p. (Cucurb.). Pilophyllum Schlechter (Chrysoglossum p.p.). Orchidaceae (n. a. 11). 2 Java, New Guinea. Pilose, with soft scattered hairs. Pilosella (Rupp.) Sch.-Bip. = Hieracium Tourn. p.p. (Comp.). Pilosperma Planch, et Triana. Guttiferae (v). i Colombia. Pilostigma Costantin. Asclepiadaceae (n. i). i Indochina. Pilostyles Guill. (Apodanthes p.p. Jiff.). Rafflesiaceae. 8 S. Am. to W. As. Parasites on Leguminosae. Pilouratea Van Tiegh. (Gomphia p.p.). Ochnaceae. i Brazil. Pilularia L. Marsiliaceae. 6 N. and S. temp. P. globiilifera L. , the pill-wort, on the margins of lakes in Brit. Creeping rhiz. bearing roots on the lower surface and linear erect 1. on the upper. The pea- shaped sporocarp, borne on the ventral side of a 1. -stalk, has a hard outer coat and consists of four sori, each containing micro- and mega- sporangia. Life history like Marsilia. Pilumna Lindl. =Trichopilia Lindl. (Orchid.). Pimelandra A. DC. (Ardisia p.p.). Myrsinaceae (n). 7 Indomal. Pimelea Banks et Soland. Thymelaeaceae. 80 Austr., Tasm., N.Z., Timor. Fls. in heads, protandrous. Cult. orn. fl. Pimeleodendron Hassk. Euphorbiaceae (A. n. 7). 3 Malay Archip. Pimenta Lindl. Myrtaceae (i). 5 'trop. Am., W.I. The unripe fr. of P. officinalis Lindl., rapidly dried, form allspice. Pimentelea Wedd. Rubiaceae (i. 5). i Peru. Pimento, Pimenta officinalis Lindl. Piraia Seem. Sterculiaceae. i Fiji. Pimpernel, Anagallis a>~uensis L. ; yellow-, Lysimachia nemontrn. Pimpinella (Riv.) L. Umbelliferae (in. 5). 100 %? and S. Afr. i in Brit., incl. P. Saxifraga L. (burnet-saxifrage). P. Anisniii L. (Medit., anise), fr. (aniseed) are used in flavouring. Pin-eyed (in Primula, &c.), long-styled. Pinaceae, see Coniferae. Pinanga Blume. Palmae (iv. i). 50 Indomal. Pinaropappus Less. Compositae (13). i Texas, Mexico. Pinarophyllon T. S. Brandegee. Rubiaceae (i. 8). i Mexico. Pinaster, Pin us Pinaster Ait. Pinckneya Michx. Rubiaceae (i. i). i S. U.S. Cinchonin in bark. Pindarea Barb. Rodr. Palmae (iv. 2). i Brazil. Finder (W.I.), ground nut, Arachis hypogaea L. Pine, Pinns\ -apple, Ananas sativus Schult. ; Bhotan-, blue-, Pimus excelsa Wall.; Bunya-Bunya -, Araucaria Bidwilli Hook.; celery-, Phyllocladus ; cluster -, Pinus Pinaster Ait. ; Corsican -, Pin/is Laricio Poir. ; cypress-, Callitris; digger -, Pinus Sabiniaiia Dougl. ; frank- PIN US 513 incense-, Pinus Taeda L. ; golden-, Psetidolarix ; hoop -, -Arau- caria Cunninghamii Ait.; huon -, Dacrydium Franklinii Hook, f . ; loblolly -, Finns Taeda L. ; Norfolk Island -, Araucana excelsa R. Br. ; Oregon-, Pseudotsuga Douglasii Carr. ; parasol-, Sciadopitys; pitch -, Pinus palustris Mill., &c. ; -sap (Am.), Afonotropa; screw -, Pandanus; stone -, Pinus Pinea L. ; sugar -, Pinus Lambertiana Dough; Weymouth -, Finns Strobus L. ; wild- (W.I.), Tillandsia; yellow -, Pinus echinata Mill. Pineda Ruiz et Pav. (Hanaro, p.p. BH.}. Flacourt. (5). 3 N.W. S. Am. Pinelea Lindl. (Ristrepia p.p. EP.). Orchid, (n. 8). i Brazil. Pinellia Tenore. Araceae (vn). 3 Japan, China. Cult. Piney varnish, Valeria indica L. Pingerin, pinguin (W.I.), Bromelia Pinguin L. Pinguicula Tourn. ex L. Lentibulariaceae. 30 N. extra-trop., Andes, Antarctic zone; 3 in Brit, (butterwort), incl. P. vulgaris L. It has a rhiz. with a rosette of rad. 1. arranged in f phyllotaxy. The 1. are covered with glands, some sessile, some on stalks, secreting a sticky fluid to which small insects adhere. Rain washes them against the edge of the 1., which is slightly upturned: when stimulated by the presence of proteid bodies it rolls over upon itself and encloses them, and then the sessile glands secrete a ferment, digest the prey, and absorb the products, after which the 1. unrolls again. P. lusitanica L. is found on the western shores of Brit, and is one of a few sp. which have migrated thus far up the Atl. coasts (common in Portugal). Pinillosia Ossa in DC. Compositae (5). 2 Cuba. Pink, Dianthus, esp. D. monspessulanus L. ; Cheddar -, D caesiits Sin.; Chinese -, D. chinensis L. ; clove -, D. caryophyllus L. Pinna, leaflet of a pinnate 1. ; pinnate, cpd., leaflets on either side of stalk, like a feather ; pinnati-fid, -partite, -sect, pinnately notched to i— i, \— |, >| depth. Pintoa C. Gay. Zygophyllaceae. i Chili. Pinus (Tourn) L. Synonymy: P. Abies L. (P. excdsa Lam.) = Picea excelsa; P. alba Ait. = Picea alba; P. bahamea L. = Abies balsamea; P. Cedrus L. = Cedrus Libani ; P. Douglasii Lamb. — Pseudotsuga Douglasii; P. Larix L. = Larix europaea ; P. maritima Lam. = P. Pinaster; P. nigra Ait. = Picea nigra; P. Picea L. = Abies pectinata. Coniferae (Pinaceae 18: see C. for gen. chars.). 70 N. temp, and on Mts. in the N. trop. Evergr. , resinous trees with both long and short shoots (see Coniferae). If a tree be examined in winter the main axes will be found each with a group of buds at the end, one term., the rest lat., covered with resinous scale 1. Each gives rise in spring to a ' long shoot ' or shoot of unlimited growth ; the term, bud continues the main axis of all, forming a year's growth before branching in a similar way again. The large branches thus form rough whorls marking each year's growth. On the stem of a long shoot no green 1. are directly borne, but only scales, first the bud scales above mentioned and then others in whose axils arise the 'short shoots,' or shoots of limited growth. Each. .of these has a few scale 1. at the base of a very short stem and ends with 2 or more green 1. of needle shape. When there are two, their upper flat sides face one another. These needle 1. exhibit xero. characters; they are thick in proportion to surface exposed, they w- 33 514 PINUS have a very stout epidermis with a hypoderm of thick walled tissue under it, and the stomata are placed at the bottom of deep pits ; the intercellular spaces too are veiy small. The fls. take the form ol the familiar cones, the 3 grouped together in spikes. Each fl., whether 3 or ? , occupies the position of a short shoot and is of limited growth — an axis with a few scale-1. below bear- ing a number of sporophylls. In the iov.] Piperales. The 2nd order of Archichlamydeae. 33—2 516 PI PERI A Piperia Rydberg (Habenaria p.p.)- Orchidaceae (ll. i)- 10 N. Am. Pipe vine (Am.), AristolochiaSipho L'Herit. ;-wort, Elatine,Erioeaidon. Piptadenia Benth. Leguminosae (i. 5). 45 trop., esp. Am. P. rigida Benth. (Brazil) yields Angico gum, used like gum-arabic. Piptanthocereus Riccob. (Cerent p.p.). Cactaceae (in. i). 10 Argen- tina to Mexico. Piptanthus Sweet. Leguminosae (ill. 2). 2 Himalaya, China. Piptatherum Beauv. = Oryzopsis Michx. p.p. (Gram.) Piptocalyx Oliv. ex Benth. Monimiaceae. i New S. Wales. Piptocalyx Torr. (Krynitzkla p.p. BH., Greeneoc&arisp.p. EP.}. Bora- ginaceae (iv. 2). 2 Pac. N. Am. Piptocarpha R. Br. Compositae (i). 30 trop. Am. Piptocelus C. Presl. Inc. sed. i Ecuador. PiptocriaetiumJ.Presl(<9r)/3(?/j-/j-p.p.j5Ar.). Gramin.(8). lotemp.S.Am. Piptocoma Cass. Compositae (ij. i S. Domingo. Piptolepis Sch.-Bip. Compositae (i). 8 Minas Geraes. Piptoptera Bunge. Chenopodiaceae (B). i Turkestan. Piptosaccos Turcz. Inc. sed. i Malaya. Piptospatha N E. Br. Araceae (v). 8 Malaya. Piptostigma Oliv. Anonaceae (2). 4 W. Afr. Piptothrix A. Gray. Compositae (2). 3 Mexico. Pipturus Wedd. Urticaceae (3). 8 Mascarenes to Australia. Pipul (peepul) Fiats religiosa L. Piqueria Cav. Compositae (2). 15 Bolivia to Mexico. Piranhea Baill. Euphorbiaceae (A. i. i). i Brazil. Piratinera Aubl. =Brosimum Sw. (Morac.) Pircunia Beit. = Phytolacca L. (Phytol.) Pirea Uurand (Dictyosperma p.p.). Cruciferae (2). i W. As. Pirigara Aubl. =Gustavia L. (Lecythid.) Piriqueta Aubl. Turneraceae. 25 warm Am., Afr., Madag. Pirola Neck . = Pyrola Tourn. (Pyrol. ) Pirolaceae = Pyrolaceae. Pironneauella 2 fls.; -mery, with more whorls than normal. Pleiocardia Greene (Streptanthus p.p.). Cruciferae (i). 10 Calif. Pleiocarpa Benth. Apocynaceae (i. 2). 10 W. Afr. Pleiocarpidia K. Schum. (Aulacodisciis BH.}. Rubi. (i. 7). i Malaya. Pleioceras Baill. Apocynaceae (n. 2). 3 W. trop. Afr. Pleiocniton Naud. ex A. Gray. Melastomaceae (7). 7 S. Brazil. Pleiococca F. Muell. Rutaceae (i). i E. Austr. Pleiogynium Engl. Anacardiaceae (2). i Queensland. Pleiomeris A. DC. (Jlfyi-smep.p. BH.}. Myrsinaceae(n). i Canaries, Madeira. Pleione D. Don (Coelogyne p.p. BH.}. Orchidaceae (n. 3). 15 Indo- mal. Cult. Pleiospora Harv. Leguminosae (in. 3). 2 S. Afr. Pleiostachya K. Schum. (Ischnosiphon p.p.). Marant. 2 Ecuador, C. Am. 522 PLEIOTAENIA Pleiotaenia Coulter et Rose (Polytaenia DC.). Umbelliferae (in. 6). i N. Am. Pleiotaxis Steetz. Compositae (12). 10 trop. Afr. Plenckia Reissek. Celastraceae. i Brazil. Plenus (Lat.), full. Pleodendron Van Tiegh. (Cinnamodendron p.p.). Winteranaceae. i W.I. Pleodiporoclina Van Tiegh. (pchna p.p.). i trop. Afr. Pleogyne Miers. Menispermaceae. i trop. E. Austr. Pleomele Salisb. (Dracaena p.p.). Liliaceae (vi). 100 palaeotrop. Pleonotoma Miers (Bignonia p.p.). Bignoniaceae (i). 6 Brazil. Pleopeltis Humb. et Bonpl. = Polypodium L. (Polypod.). Pleopetalum Van Tiegh. (Och>ia p.p.). Ochnaceae. 4 Indomal. Pleouratea Van Tiegh. (Goinphia p.p.). Ochnaceae. i Brazil. Plerandra A. Gray. Araliaceae (i). 6 New Guinea to Fiji. Plerandropsis R. Viguier. Araliaceae (i). i Tonquin. Pleroma D. Don = Tibouchina Aubl. p.p. (Melast.). Plesiatropha Pierre ex Hutchinson=Mildbraedia Pax (Euph.). Plesmonium Schott. Araceae (iv). i N. India. Plethadenia Urb. Rutaceae (i). i S. Domingo. Plethiandra Hook. f. Melastomaceae (n). 2 Borneo. Pleur-, pleuro- (Gr. pref.), side-; -anthous, Orchidaceae ; -rhizal, Cruciferae. Pleurandra Labill. = Hibbertia Andr. p.p. (Dillen.). Pleurandropsis Baill. (Asterolasia p.p. BH.). Rutac. (i). i W. Austr. Pleuranthium Benth. (Epidendnim p.p. EP.). Orchid, (n. 6). i Cuba. Pleuranthodes Weberb. (Gonania p.p.). Rhamnaceae. i Hawaiian Is. Pleuricospora A. Gray. Pyrolaceae. i California. Pleurisanthes Baill. Icacinaceae. i French Guiana. Pleuroblepharis Baill. Acanthaceae (iv. B). i Madag. Pleuroblepharon Kunze. Orchidaceae. Nomen. Pleurobotryum Barb. Rodr. (Plewoihallis p.p. ). Orch. (n. 8). i Braz. Pleurocalyptus Brongn. et Gris. Myrtaceae (n. i). i New Caled. Pleurocarpaea Benth. Compositae (i). i trop. Austr. Pleurocoffea Baill. Rubiaceae (n. 4). i Madag. Pleurogyna Eschsch. ex Cham, et Schlechtd. Gentianaceae (i). 7 N. temp, and cold. Pleuropetalum Hook. f. Amarantaceae (i). 2 C. Am., Galapagos. Pleurophora D. Don. Lythraceae. 5 S. Am. Pleuro phragma Rydberg (Thelypodium p.p.). Crucif. (i). 3 N. Am. Pleurophyllum Hook. f. Compositae (3). 2 S. islands off N.Z. Pleuropogon R Br. Gramineae (10). 3 N. temp, and polar. Pleuropterantlia Franch. Amarantaceae (2). (Chenop. BH.) i Somaliland. Pleuropterygium Gross. (Polygonum p.p.). Polygonaceae (n. 2). 3 Japan, Saghalien. Pleuroridgea Van Tiegh. (Ochna p.p.). Ochnaceae. 5 trop. Afr. Pleurosorus Fee. Polypodiaceae. 3 Spain, Chili, N.Z. Pleurospermum Hoffm. Umbelliferae (ill. 4). 25 N. As., E. Eur. Pleurostachys Brongn. (Rhynchospora p.p. EP.). Cyper. (n). 30 S. Am. PO 523 Pleurostelma Baill. Asclepiadaceae (n. i). i Madag. Pleurostelma Schlechter (Schlechterella EP.). Asclep. (i). i E. Afr. Pleurostemon Raf. Inc. sed. i N. Am. Pleurostylia Wight et Arn. Celastraceae. 2 S. Afr. to India. Pleurothallis R. Br. Orchidaceae (n. 8). 500 trop. Am., W.I. Pleurothyrium Nees. Lauraceae (i). i Peru. Plicate (vernation), folded several times. Plicouratea Van Tiegh. (Gomphia p.p.). Ochnaceae. 8 Brazil. Plinthus Fenzl. Aizoaceae (u). i Cape Colony. Pliogynopsis O. Ktze. (Pleiogynium Engl.). Anacard. (2). i N.E. Austr. Plocama Ait. Rubiaceae (n. /). i Canaries. Plocaniophyllon T. S. Brandegee. Rubiaceae (i. 8). i Mexico. Plocoglottis Hlume. Orchidaceae (119). 10 Malay Archipelago. Plocosperma Benth. Loganiaceae. 2 Guatemala, Mexico. Pluchea Cass. Compositae (4). 30 trop. and subtrop. Plukenetia L. Euphorbiaceae (A. n. 2). 12 warm Am., Afr., As. Plum, Primus domestica L.; coco-, Chrysobalanus; date-, Diospyi-os Kaki L. f. ; hog-, Spondias; Madagascar-, Flaconrtia ; marma- lade- (W.I.), Lucitma iiiammosa Gaeitn. f.; sapodilla -, Achras Sapota L.; tamarind-, Dialiuin inditin L. ; -tree, seaside (W.I.), Ximenia. Plumbagella Spach (Plumbago p.p. J3H.). Plumbaginaceae. i E. As. Plumbaginaceae (EP., BH.). Dicotys. (Sympet. Plumbaginales ; Pri- mulales BH.}. 10 gen., 280 sp. cosmop., but esp. on salt steppes and sea-coast. Perennial herbs or shiubs with narrow I., on whose surface water glands occur, or sometimes chalk glands (cf. Saxifraga). Infl. of various types, racemose and cymose (see Plumbago, Ceratostigma, Statice, Armeria), bracteolate. Fls. reg., $ , 5-merous, the odd sepal post. K persistent, C often nearly polypetalous, conv., A 5, epipet. and opp. the petals, G (5), i-loc., with basal placenta, and one anatr. ov. , whose stalk curves up to the top of the loc. and causes the micropyle to be directed upwards. Styles or stigmas 5. Nut; embryo straight, in floury endosp. The fam. is distinguished from Primulaceae by the ovary and styles. Chief genera: Plumbago, Ceratostigma, Acantholimon, Armeria, Statice, Limoniastrum. Plumbaginales. The 3rd order of Dicots. Sympetalae. Plumbago Tourn. ex L. Plumbaginaceae. 10 cosmop. Racemose infl. K with glandular hairs, aiding seed-dispersal. Plumbeus (Lat.), lead-coloured. Plumeria Tourn. ex L. Apocynaceae (i. 3). 45 warm Am. Several cult. orn. perf. fls. (offered in Buddhist temples), esp. P. acutifolia Poir (temple tree, frangipani). Plummera A. Gray. Compositae (6). i Arizona. Plumose, feathered. Plumule, the stem bud of the embryo pi. in the seed. Pluri- (Lat. pref.), many-. Pneumatophore, erect root with aerenchyma; cf. Mangroves. Pneumonanthe Gleditsch (Gentiana p.p.). Gentian, (i). 25 N. Am. Po (fl. -class), fls. offering pollen only to visitors, Anemone, Hypericum, Papaver, Rosa, Thalictrmn, Verbascuin. 524 POA Poa L. Gramineae (to). 150 cosmop. ; 8 in Brit, (meadow grass). Many are useful pasture-grasses. Poacynum Baill. (Apocynum p.p.). Apocynaceae (11. i). i trop. Afr. L. alt. Poaephyllum Ridley (Agrostophyllum p.p.). Orchid. (11.4). i Malaya. Poagrostis Stapf. Gramineae (9). i S. Afr. Pochota Goyena. Bombacaceae. i Nicaragua. Pockwood tree (W.I.), Guaiacum. Pocockia Ser.=Trigonella L. p.p. (Legum.). Podachaenium Benth. (Ferdinanda p.p. JSP.), Compositae (5). i C. Am. Cult. orn. 1. Podadenia Thw. Euphorbiaceae (A. n. 2). i Ceylon. Podagrostis Scribner et Merrill (Agrostis p.p.). Gramineae (8). r Alaska to Oregon. Podalyria Lam. Leguminosae (in. 2). -208. Afr. Podandra Baill. Asclepiadaceae (n. i). i Bolivia. Podandria Rolfe (Habenaria p.p. EP.}. Orchidaceae (n. i). i trop. Afr. Pedantries Haw. (Stapelia p.p.). Asclepiadaceae (n. 3). 8 S. Afr. Podanthum Boiss. (Phytenma p.p.). Campanulaceae (i. i). 50 Medit. Podanthus Lag. Compositae (5). 2 Chili, Argentina. Podistera S. Wats. {Cyviopteriis p.p.). Umbelliferae (in. 5). i N. Am. Podo- (Gr. pref.), stalk. Podocarpus (L'Herit.) Pers. Coniferae (Taxac. 5 ; see C. for gen. chars.). 60 E. As. and S. temp, and trop. Dioec.; 9 peculiar. There are usu. three pairs of scales, decussate. One of the middle pair projects above all the rest, bearing an anatr. ov. The other 5 are sometimes united to form the so-called recept. The fr. usu. consists of a fleshy mass (the 'recept.') bearing an arillate seed. In some the sterile scales do not become fleshy. Timber valuable. Podochilus Blume. Orchidaceae (n. a. ill). 12 Inclomal. Epiphytes. Podochrea Fourr.= Astragalus L. p.p. (Legum.). Podochrosia Baill. Apocynaceae (i. 3). i New Caledonia. Podococcus G. Mann et H. Wendl. Palmae (iv. i). i W. trop. Afr. Podocoma Cass. Compositae (3). 7 Austr., S. Am. Podogynium Taub. Leguminosae (11. 2). i E. trop. Afr. Podolasia N.E. Br. Araceae (iv). i Borneo. Podolepis Labill. Compositae (4). 15 Austr. Podolobium R. Br. = Oxylobium Andr. (Legum.). Podonephelium Baill. Sapindaceae (i). i New Caledonia. Podoon Baill. Phytolaccaceae. i Yunnan. Podopetalum F. Muell. Leguminosae (in. i). i Queensland. Podophania Baill. Compositae (2). i Mexico. Podophorus Phil. Gramineae (8). i Juan Fernandez. Podophyllum L. Berberidaceae. 5 N. temp. (May apple). In P. peltatum L. the rhiz. sends up yearly a shoot bearing t\vo large peltate 1., which hang down when young like a closed umbrella. A drug is prepared from the rhiz. Fr. a berry, whose flesh consists chiefly of the plac. which grows up round the seeds (wrongly termed aril). Podopogon Ehrenb. Inc. sed. Nomen. Podopterus Humb. et Bonpl. Polygonaceae (in. i). i Mexico. POGOGYNE 525 Podorungia Baill. Acanthaceae (iv. B). i Madag. Podosciadium A. Gray (Eulophus p.p. EP.}. Umbelliferae (in. 5). 2 Calif. Podosemum Desv. = Muehlenbergia Schreb. p.p. (Gram.). Podosperma Labill. = Podotheca Cass. (Comp.). Podospermum DC. = Scorzonera L. p.p. (Comp.). Podostelma K. Schum. A>-clepiadaceae (n. i). i Abyssinia. Podostemma Greene (Asclepias p.p.). Asclepiad. (n. i). 6 N. Am. Podostemaceae (EP. and BH. inch Tristichaceae). Dicotyledons (Archichl. Rosales ; Multiovulatae aquaticae BH. ). 22 gen., 100 sp. trop. A remarkable fam. of plants living only in rushing water, and growing only on rocks in rivers. The morphology of the veg. organs is extremely varied and complex. The seeds are shed on the rocks during the drier season of the year, and germinate when the rains cause them to be submerged. The primary axis is usu. small, and from the base there buds out a green thallus, usu. of adv. root nature. In Podostemon, &c. it is ± filamentous, creeping on the rock, and attached to it by hairs or exogenous projections termed haptera. In Dicraea it is ± freely swimming and often ribbon-like or sea-weed-like. In Hydrobryum it is ± flattened, creeping, lichen-like. Other compli- cations occur in Castelnavia and others. From the thallus in most cases endogenous secondary .shoots arise, and remain veg. (1. alt., simple or much divided) till the latter part of the rainy season, when they form fls. which open when exposed by the fall of the water. The plants die after shedding their seed, unless an early rise of water occurs. Their outer tissues are usually very siliceous. Fls. simple, 5 , reg. or not, naked, enclosed in a spathe, hypog. A i-2-oc , G usu. (2) 2-loc. with thick axile plac. Ovules oo (exc. Farmeria), anatr. Caps.; exalbum. seeds. The more dorsiventral the veg. organs, the more dorsiventral in general is the fl., the phenomenon showing pro- gressively in A, G, fr. , seed, and embryo. Chief genera : Rhyncholacis, Mourera, Podostemon, Dicraea, Hydrobryum, Castelnavia. [For full details see Willis in Ann. Pcrad. i, 1902 and literature there quoted.] Podostemon Tul. Podostemaceae. 12 trop. and subtrop. Podostigma Ell. Asclepiadaceae (n. i). i S.E. U.S. Podotheca Cass. Compositae (4). 5 temp. Austr. Podranea Sprague (Tecoma p.p.). Bignoniaceae (2). 2 warm Afr. Poecilandra Tul. Ochnaceae. i Guiana, N. Brazil. Poecilanthe Benth. Leguminosae (in. 6). 3 Brazil. Poecilochroma Miers. Solanaceae (2). 8 Peru, Ecuador. Poeciloneuron Bedd. Guttiferae (iv). 2 S. India. Poecilopteris Presl— Leptochilus Kaulf. Poecilostachys Hack. Gramineae (10). i Madag. Poederiopsis Rusby (Pacderia p.p.). Rubiaceae (il. 6). I Boliv. Poeppigia C. Presl. Leguminosae (n. 8). i trop. Am., VV.I. Poga Pierre. Rhizophoraceae. i Gaboon. Poggea Guerke. Flacourtiaceae (2). 2 W. trop. Afr. Poggeophyton Pax. Euphorbiaceae (A. n. 2). i W. trop. Afr. Pogochloa Sp. Moore. Gramineae (10). i Matto Grosso. Pogocybe Pierre. Leguminosae (li. 7). i Cochinchina. Pogogyne Benth. Labiatae (vi). 3 California. 526 -POGON -pogon (Gr. suff.), a beard of hairs. Pogonanthera Blume. Melastomaceae (i). 3 Malaya. Pogonanthus Montr. Inc. sed. i New Caled. Pogonarthria Stapf. Gramineae (10). 2 trop. and S. Afr. Pogonatherum Beauv. Gramineae (2). 2 India to Japan. Sta. 2. Pogonia Andr. = Myoporum Banks et Soland. (Myopor.). Pogonia Juss. Orchidaceae (n. 2). 5 E. As. Pogoniopsis Reichb. f. Orchidaceae (II. 2). i Brazil. Pogonophora Miers. Euphorbiaceae (A. n. 5). i Guiana, Brazil. Pogonopus Klotzsch. Rubiaceae (i. i). 3 S. and C. Am. Pogonotrophe Miq. = Ficus Tourn. p.p. (Mor. ). Pogopetalum Benth. =Emmotum Desv. (Icacin.). Pogostemon Desf. Labiatae (vi). 35 Indomal. P. Patchoiily Pellet yields the well-known perfume by distillation. Poicilla Griseb. Asclepiadaceae (n. 4). i Cuba. Poicillopsis Schlechter (Poicilla p.p.). Asclepiadaceae (n. 4). i S. Domingo. Poikilacanthus Lindau (Adhatoda p.p.). Acanth. (iv. B). 4 S. Am. PoiMlospermum Zipp. ex Miq. Urticaceae (3). i Amboina. Poinciana Tourn. ex Linn. Leguminosae (n. 7). 3 trop. Afr., Madag., As. P. regia Boj. cult. orn. flg. tree (flamboyante). Poinsettia R. Grab. = Euphorbia L. p.p. (Euph.). Poiretia Vent. Leguminosae (ill. 7). 5 trop. Am. Poison, Acokanthera,Acouitum, Antiaris, Erythrophloeum, Physostignia, Strychnos, Toxicodendrttm , &c.; -berry (W.I.), Cestnim ; bikh -, Aconititm ; curare-, Strychnos toxifera Schomb. ; -dogwood (Am.), Rhus venenata DC.; -hemlock (Am.), Contain ; -ivy, Rlnts Toxi- codendron L.; -oak, jRhus ; - sumacb. (Am.), Rhus; wourali -, Strychnos. Poissonia Baill. (Coursetia p.p.). Leguminosae (ill. 6). i Peru. Poitaea Vent. Leguminosae (in. 6). 5 W.I. Poium (Cl.), a meadow formation. Poivrea Comm. ex Thou. = Combretuna L. (Combr.). Pokeweed, Phytolacca. Polakia Stapf. Labiatae (vi). i Persia. Polakowskia Pittier. Cucurbitaceae (4). i Costa Rica. Polanisia Rafin. (CUome p.p. BH.}. Capparid. (v). 30 trop., subtrop. Polemannia Eckl. et Zeyh. Umbelliferae (in. 5). 2 S. Afr. Polemoniaceae (£f-, BH.\ Dicots. (Sympet. Tubiflorae ; Pole- moniales BH}. 8 gen., 200 sp., chiefly N. Am.; a few in Chili, Peru, Eur, N. As. Herbs (rarely shrubby below), glabrous or shortly hairy, with usu. opp. exstip. 1. Els. in cymes (sometimes condensed into involucrate heads), $ , reg. or slightly -|- , with or without bracteoles. K (5), valvate or imbr., persistent ; C (5), bell- funnel- or plate-shaped, usu. conv. ; A 5, epipet., alt. with petals ; G (3) or rarely (2 — 5), on a disc, multiloc., with simple style ± lobed at tip. Ovules i — oo in each loc., anatr., sessile. Fr. usu. a loculic. caps. Embryo straight, in endosp. Chief genera : Cobaea, Cantua, Phlox, Collomia, Gilia, Polemonium (mostly favourite border plants). Polemoniales (Bff.}. The 8th order of Gamopetalae. Polemoniella A. A. Heller (Polemonium p-p-)- Polemon. i N. Am. POLY A DO A 527 Polemonium (Tourn.) L. Polemoniaceae. 20 N. temp., Chili. P. caentlenm L. (Jacob's ladder) in Brit. Polianthes L. Amaryllidaceae (n). 3 C. Am. P. tuberosa L. (tuberose) cult, for scented fls. Poliomintha A. Gray (Hedeoma p.p. EP.). Labiatae (vi). 5 S.W. N. Am. Poliothyrsis Oliv. Flacourtiaceae (4). i China. Polish wheat, Triticitm polonicum L. Politus (Lat.), polished. Polium Tourn. ex L. = Teucrium Tourn. p.p. (Labial.). Pollards, Salix, &c. Pollen, the microspores of fl. pi.; - as food for insects, cf. Po ; - cup, Goodemaceae - dimorphism, Faramea ; - sac, cf. Cucurbitaceae. Pollia Thunb. Commelinaceae. 16 palaeotrop. Pollichia (Solancl. in) Ait. Caryophyll. (i. 4). 2 trop. and S. Afr. Pollination, the placing of pollen upon the stigma or ovule, may be effected by wind, animals, or water, and may be cross or self-fert. Wind-pollinated fls. (class W) show abundance of pollen, ripening simultaneously, dry and incoherent, freely accessible to the wind (catkins, pend. sta., explosive anthers, &c.); large much-branched stigmas to catch it; often flowering before the 1. appear, &c.; Artemisia, Betulaceae, Calluna, Carex, Casiiarina, Coni/erae, Corylus, Cycada- ceae, Cyperaceae, Elaeagnaceae, Empetrum, Fiaxinns, Gra/nineae, Humuhts, Juglandaceae, Jttncaceat', Littorella, Mercurialis, Rlyro- tharnnaceae, Platanns, Potamogeton, Quercus, Rumtx, Spargdnium, Spinacia. Thalictnun, Triglochiii, Typkaceae, Ulmaceae, Urticaceae, Zea. Pollination by animals is usu. by insects; the most important are Coleoptera (beetles), very short tongued, Diptera (flies), mostly short tongued, but the Syrphidae or hover-flies long tongued and clever at finding concealed honey, Hymenoptera (ants, wasps, bees, &c.) all short tongued but the bees, and Lepidoptera (butterflies and moths) all long tongued. Other animals are very rarely employed; bats in Freycinetia Cf. Flower classes, Floral Mechanisms. Pollination by water in Ceratophylliitn, Enhalus, Najas, Riippia, Vallisneria, Zanmchellia, Zostera, &c. ; legitimate and illegitimate, Ly thrum, Primula. Pollinia Trin. Gramineae (2). 40 warm |^. Pollinium, a number of pollen-grains united into a mass (a definite number of masses per anther), Asdepiad., Orchid. Polpoda C. Presl. 1'hytolaccaceae (Aizoaceae J3H.). i S. Afr. Poly- (Gr. pref.), many-; -adelphous, -androus, &c.; -carpic, fruiting many times ; -embryony, formation of > i embryo in an ovule ; -gamy, $ , a; -wood, Cocos nucifera L. Pores, dehiscence by, cf. Stamen. Porlieria Ruiz et Pav. Zygophyllaceae. 3 Mexico, Andes. The leaflets of P. hygrometrica R. et P. spread out horiz. at night, but in the day fold up in pairs, reducing transpiration. Good timber. Porochna Van Tiegh. (Ochna p.p.)- Ochnaceae. 12 trop. Afr. Porocystis Radlk. Sapindaceae (i). i Amazon valley, Guiana. Porogamae, see Chalazogamae ; porogamic (feit.), by micropyle. Porophyllum Vaill. ex L. Compositae (6). 28 warm Am. Porospermum F. Muell. Araliaceae (2). i N.E. Austr. Porotheca K. Schum. Menispermaceae. i New Guinea. Porpax Lindl. (Eria p.p. BH.}. Orchidaceae (n. 15). 6 Indomal. Porphyranthus Engl. Burseraceae. i Cameroons. Porphyrocodon Hook. f. Cruciferae (4). i Colombia. Porphyrocoma Scheidw. (Dianthera p.p. BH.}. Acanthaceae (iv. B). 2 trop. S. Am. Cult. orn. fl. Porphyrodesme Schlechter (Saccolabituit p.p.). Orchidaceae (u. 20). i New Guinea. Porphyroglottis Ridl. Orchidaceae (n. a. n). i Borneo. Porphyrospatha Engl. Araceae (vi). 3 C. Am. Porphyrostemma Benth. ex Oliv. Compositae (4). i E. Centr. Afr. Porrum (Tourn.) L. = Allium Tourn. p.p. (Lili.). Portea C. Koch. Bromeliaceae (4). 5 Brazil. Cult. orn. infl. Portenschlagia Vis. Umbelliferae (in. 5). i Dalmatia. Porteria Hook.= Phyllactis Pers. (#Zf.)=Valeriana Tourn. p.p. Portesia Cav. = Trichilia P. Br. (Meli.). Portland arrowroot. Arum ma.cn/atum L. Portlandia P. Br. Rubiaceae (i. i). 8 W.I. , Mexico. Portugal laurel, Primus lusitanica L. Portulaca L. Portulacaceae. 20 trop. and subtrop. (purslane). Thefl. has a semi-inf. ovary and 4 — costa. It remains closed in bad weather. The sta. of P. oleracea L. are sensitive to contact and move toward the side touched. Portulacaceae (EP., BH.}. Dicots. (Archichl. Centrospermae ; Caryo- phyllinae BH.}. 17 gen., 225 sp., cosmop., but esp. Am. Most are 534 FOR TULA CA CEAE annual herbs, often with fleshy 1., and with stipules (sometimes repres. by axillary bundles of hairs). Fls. usu. in cymes (often dich. with tendency to cincinni), reg., 5 . K 2, the lower sepal (usu. ant.) over- lapping the upper (the two are often regarded as bracteoles); €5; A 5 + 5. or 5 °PP- the petals, or some other number; G (2 — 8) usu. (3), sup. exc. in Portulaca, i-loc. with several stigmas and 2 — oo campylotr. ov. on a central basal plac. The fls. secrete honey and are mostly insect-pollinated. Caps, with album, seeds; that of Claytonia and Montia is explosive ; embryo more or less curved round the peri- sperm. C/z/^£ttmz.-Calandrinia,Claytonia,Montia,Portulaca,Lewisia. Portulacaria Jacq. Portulacaceae. 2 S. Afr. Portulaceae (BH. ) = Portulacaceae. Posadaea Cogn. Cucurbitaceae (2). Posidouia Kon. Potamogetonaceae. ing glass). Poskea Vatke. Boraginaceae (11). 1 Colombia. 2 Austr., Medit. (used for pack- Somaliland. Posoqueria Aubl. Rubiaceae (i. 8). 5 S. Am., W.I. Posterior, the side facing to the stem on which the flower is a branch (upper side in most floral diagrams). Postea Boiss. et Blanche. Compositae (4). 4 Syria, Persia. Posticous, on the posterior side. Potalia Aubl. Loganiaceae. i trop. S. Am. Potameia Thou. Lauraceae (il). i Maclag. Potamium (Cl.), a river formation. Potamobryon Liebm. Podostemaceae. 3 Mexico. Potamogeton (Tourn. ) L. Potamogetonaceae. 90 cosmop. n or more (cf. Rubus) in Brit, (pond-weed). Water pi. with creeping sympodial rhiz. and erect leafy branches ; all submerged or some floating. A series of types occurs, beginning with the floating sp. and ending with the narrow-leafed submerged ones. There can be no doubt of the origin of the P. from land pi., and Schenck looks upon P. natans L. as the sp. least modified to suit a water existence, i.e. the nearest to the ancestral type. The upper 1. are ovate, leathery, and float ; the lowersubmerged, sometimes linear. Then come such as P. hetero- phyllus Schreb. where the sub- merged 1. are all narrow. Next P. lucens L., P. crispus L. , &c. with all the 1. lanceolate and sub- merged. Then in P. obtnsifolins Mert. et Koch, P. pusilhis L.,.'p. BH.}. Monocots. (Helobieae). 9 gen., i 20 sp., cosmop., all water plants, several marine. There is in most a creeping stem or rhiz., mono- or sym-podial, attached to the soil by adv. roots, and sending off erect branches upwards into the water. These usu. have ribbon leaves, submerged (exceptions in Potamogeton), arranged in \ phyllotaxy. The base is sheathing, and within the sheath are the small scales (sijnamulae intravaginales] which occur in most of the Helobieae. The infl. is a spike or cyme, or the fls. solitary. The fls. possess no true P (exc. Zannichellia ? ), but Potamogeton shows leaf-like outgrowths from the sta. which perform the P functions ; similar structures occur in Zostera and Ruppia. In several gen. it is not easy to decide what is to be regarded as the fl. (see Zostera). Fl. § or unisex., reg., i — 4-merous. Cpls. free or only i. Ovules i in each cpl., pend. orthotr. Fruit one- seeded. No endosp. Embryo with well-developed hypocotyl. For details see genera. Chief genera: A. Fls. in spikes: 1. saltwater: Zostera, Phyllospadix, Posidonia, Ruppia. 2. fresh or brackish : Potamogeton. B. Fls. in cymes or sol. : Cymodocea, Zannichellia. Potamopbila R. Br. Gramineae (6). i Austr. Potaninia Maxim. Rosaceae (in. 2). i Mongolia. Potato, Solatium tuberosum L. ; sweet-, Ipomoea Batatas Lam. Potentilla L. (incl. Comarum L., Tormentilla L.). Rosaceae (ill. 2). 225, chiefly N. temp, and arctic, a few S. temp, and Andes; 9 in Brit., incl. P. Anserina L. (silverweed), P. reptans L. (cinquefoil), P. Coniarum Nestl., P. Tormentilla Neck, (tormentil), &c. Herbs, usu. with creeping stems which root at the nodes and thus multiply the plant veg. There is a well-marked epicalyx of small green 1. outside and alt. with the seps. These are the slips, of the seps. united in pairs; often one or more may be seen with two lobes or even completely divided. Fls. of class AB, homogamous, fly-visited; honey secreted by a ring-shaped nectary within the sta. Poteranthera Bong. Melastomaceae (i). 4 Brazil, Guiana. 536 PO TERIDIUM Potericlium Spach, Rydberg (Sangnisorba p.p.)- Ros. (in. 5). i N. Am. Poterium L. (BH. inch Sangnisorba, q.v.im Brit. sp.). Rosaceae (in. 5). i Italy, E. Medit. Potherbs, cf. Edible Products. Pothoidium Schott. Araceae (i). 2 Malaya. Pothos L Araceae (i). 80 trop. As., Madag. Monopodial (seefam.). Stem climbing, with adv. roots. The buds break through the axils, so that the branching seems infra-axillary. Fl £ . P 3 + 3. Pothuava Gaudich. (AecJiniea p p. BH. ). Bromeliaceae (4). i S. Am. Pottingeria Prain. Saxifragaceae (v). i India. Pottsia Hook, et Arn. Apocynaceae (n. 2). 3 India, China. Pouchetia A. Rich. Rubiaceae (i. 8). 3 W. trop. Afr. Poulsenia Kggers. Moraceae (11). i Ecuador. Pounce, Tetraclinis qnadrivalvis Mast. Poupartia Comm. (Spondiasp.p.HJ?.). Anacardiaceae (2). 4 Madag., Masc. Pourouma Aubl. Moraceae (in). 20 trop. S. Am. Ed. fr. Pourretia Ruiz et Pav. = Puya Molina (Bromel.). Pourthiaea Decne. Rosaceae (n). sE.As. Pouslowia Wight. Inc. sed. Nomen. Pouteria Aubl. Sapotaceae (i). 30 trop. Am. Pouzolzia Gaudich. Urticaceae (3). 40 palaeotrop. The root of P. tuberosa Wight is eaten in India. Poverty grass (Am.), Aristida dichotoma Michx. Pozoa Lag. (Azorella p.p. BH.). Umbelliferae (i. 2). 4 Austr., S. Am. Pradosia Liais (Lucuma p.p. BH.}. Sapotaceae (i). i Rio de Janeiro. Hard wood. Praecox (Lat.), appearing early. Praemorse, as if bitten off. Praesepium Spreng. Rosaceae. Nomen. Prageluria N.E. Br. = Pergularia (L.) N.E. Br. Prairie, the grass country E. of the Rocky Mts. ; - clover (Am.), Petalo- stemon ; - grass (Am.), Sporobolus asper Kunth ; - turnip, Psoralea. Prainea King ex Hook. f. Moraceae (in). 3 Malaya. Prangos Lindl. Umbelliferae (in. 4). 36 Medit., C. As. Prantleia Mez (Orthophytum Beer). Bromeliaceae (4). 2 C. Brazil. Praravinia Korth. Rubiaceae (i. 7). i Borneo. Prasanthea Decne = Paliavana Vand. (Gesner.). Prasium L. Labiatae (in), i Medit. Prasophyllum R. Br. Orchidaceae (n. 2). 32 Austr., N.Z. Pratal, growing in meadows; pratensis (Lat.), of meadows. Pratia Gaudich. Campanul. (ni;. 16 S. Am., Austr., N.Z., trop. As. Prefoliation, vernation. Premna L. Verbenaceae (4). 45 trop. and subtrop. |%. Prenanthella Rydberg (Prenanthes p.p.). Compositae (13). i N.W. Am. Prenanthes Vaill. ex L. Compositae (13). 27 N. temp, and trop. Afr. Preptanthe Reichb. f. (Calanthe p.p. BH.}. Orchid, (n. 9). 2 trop. As. Prepusa Mart. Gentianaceae (i). 3 Brazil. Prescottia Lindl. Orchidaceae (n. 2). 22 trop. Am., W.I. PRIMULA CEAE 5 3 7 Preserving-, cf. Collecting. Preslia Opiz. Labiatae (vi). i W. Medit. Prestoea Hook. f. Palmae (iv. i). 2 Trinidad, Colombia. Prestonia R. Br. Apocynaceae (n. 2). 30 trop. Am. Pretrea f. Gay. Pedaliaceae. i S. and trop. Afr. Pretreothamnus Engl. Pedaliaceae. i Somaliland. Preussiella Gilg. Melastomaceae (i). i W. trop. Afr. Prevostea Choisy (Breweria p.p. BH.}. Convolv. (i). 10 trop. Afr., Am. Prickly • ash, Zanthoxylum fraxineitin Willd. ; -heath, Pernettya', -lettuce, Lactuca; - pear, Opuntia; -pole (W. I.), Bactris', -poppy (Am.), Argemone; -shield-fern, Aspidinm acnhatum Sw. ; -withe (W.I.), Cereus triangularis Mill. Priestleya DC. Leguminosae (in. 3). 158. Afr. Prim (Am.), privet, Li gust rum vulgare L. Primine, outer coat of an ovule. Primrose, Primula vulgaris Huds. ; Cape-, Streptocarpus ; Chinese-, Primula sineiisis Sabine; evening-, Oenothera; -willow (W.I.), Jussieua. Primula L. Primulaceae. 210 N. hemisph. chiefly in hilly districts. A few elsewhere, e.g. P. farinosa L. , var. magellanica Hook., at the Str. of Magelhaen. The rhizome is a sym podium, each joint termi- nating in an infl. In some sp. this consists of successive whorls of fls. arranged up a long stalk, e.g. japonica A. Gray. A few of the more important sp. are: P. sinensis Sabine, the Chinese prim- rose, P. elatior Hill, the oxlip (Brit.); P. vulgaris Huds. (P. acaulis Hill), the primrose (Brit.), P. veris Lehm. (P. offidnalis Jacq.), the cowslip (Brit.), P. farinosa, L. (Brit.), P. japonica A. Gray, P. Au- ricula L., the auricula with its many forms. A great many hybrids occur, and garden vars. In the double crowned cowslip the K has become petaloid, so that the fl. looks as if it had two Cs, one within the other. The fls. are dimorphic, heterostyled. On one pi. are long-styled fls. with sta. halfway up the tube and the stigma at its mouth; on another plant are short-styled fls., with stigma halfway up and anthers at the mouth. The depth and narrowness of the tube suit the fl. to bees or butterflies, and these tend to carry pollen from long sta. to long style or from short to short. These 'legitimate' pollinations (see Lythrum) which are at the same time crossings, are the only ones which produce a full complement of fertile seed. The fl. stalks in umbellate forms, e.g. cowslip, stand close and erect till the fls. open, then spread out, and close up again as the fr. ripens ; thus the caps, is held erect and the seeds must be shaken out. Primulaceae (EP.,BH.). Dicots. (Sympet. Primulales). 28gen., 35osp. co.smop., but esp. N. temp. 9 Brit, genera. Herbaceous pi., commonly perenn., with rhiz. or tubers; 1. opp. or alt., exstip. Fls. often borne on scapes, which when > i -flowered are term. ; they are usually actinom., 5 , often heterostyled, and j-merous, without bracteoles, the odd (4th) sepal post. K (5) persistent; C (5), reg. (exc. Coris), or 5, or o (Glaux) ; A 5, epipet. and opp. the pets.; occasionally 5 stds. alt. 'with the pets.; anthers intr. The presence of the stds. here 538 PRIMULACEAE Floral diagram of Primula acaulis (after Eichler). as in Myrsinaceae explains the antepetalous position of the sta. as due to the abortion of the originally outer whorl. Much discussion took place on this subject in former times, espe- cially after Pfeffer's discovery of the peculiar development of the corolla from the backs of the sta. (see Eichler, Bliithendiag. or Asa Gray, Struct. Bot. p. 197). Ovary superior or half-inferior (Samolus), syncarpous with free central placenta, typically of 5 cpls., but this is not easily proved, as no partitions (cf. Caryophyllaceae) are found in the ovary and the style and stigma are simple. The capsule splits into 5 valves, and monstrous fls. with 5 leaves in place of the ovary occur ; hence \ve may perhaps assume 5 cpls. Ovules oo , spirally or in whorls on the placenta, semi-anatro- pous. The morphology of the free-central placenta has also been a subject of dispute (see Pax in Nat. Pfl., or Eichler, Bliithendiag.}. A great number of the P. have heterostyled flowers (Primula, Hottonia, Glaux, Androsace, &c., q. v. for details). The fruit is a capsule, dehiscing in various ways, but usually by teeth at the tip, one opposite to each sepal. Seeds few or many ; embryo small, in fleshy endosperm. Chief genera: Primula, Andro- sace, Soldanella, Hottonia, Samolus, Lysimachia, Trientalis, Glaux, Anagallis, Cyclamen. Primulales. The 2nd order (EP.) of Sympetalae. The 5th order (BH.) of Gamopetalae. Primulina Hance. Gesneriaceae (i). i China. Primulinae (Warming). The 3rd order of Sympetalae. Prince's feather, Amaranthus. Princewood bark (W. I.), Exostemma. Principes. The 5th order of Monocotyledons. Pringlea Anders. "ex Hook. f. Cruciferae (i). P. antiscorbutica R. Br. (Kerguelen cabbage), the only sp., has the habit of a cabbage, with the fls. borne on lat. axes. It grows only "on the tempestuous shores of Kerguelen's I., where winged insects cannot exist, because at every flight they run the risk of being drowned. Under these circumstances the plant has become modified for fert. by wind, acquiring exserted anthers and long filiform stigmatic papillae. It still retains traces of its descent from entomophilous ancestors ; for while on the great part of the island it is devoid of petals, it occurs abundantly in shaded places with petals" (Miiller). Pringleochloa Scribner (Opizia p.p. EP.). Gramineae (i i). i Mexico. Pringleopnytum A. Gray (Ber«inia p.p. EP.). Acanth. (iv. B). i Calif. Prinos Gronov. ex L. = Ilex Tourn. p.p. (Aquifol.). Prinsepia Royle. Rosaceae (v). i Himalaya. PROSOPANCHE 539 Printzia Cass. Compositae (4). 5 S. Afr. Prionacline Nees. Gramineae (9). r S. Afr. Prionanthium Desv. (Prionachne p.p. EP.). Gram. (9). 3 Indomal., S. Afr. Prionitis L. = BarIeria L. p.p. (Acanth.). Prionium E. Mey. Juncaceae. i Cape Col., P. Palmita E. Mey. (P. serrattim Buchen.), the palmiet, a shrubby aloe-like plant with a stem i — 2 metres high, covered with the fibrous remains of old 1. It grows on the edges of streams, sometimes almost blocking them up. Veg. propagation takes place by formation of runners. Adv. roots form between the 1. [Buchenau in Bibl. Bot., No. 27.] Prionophyllum C. Koch (Dyckia p.p. BH.}. Bromel. (2). 2 S. Braz., Urug. Prionosciadium S. Wats. Umbelliferae (in. 6). [o Mexico, C. Am. Prionotes R. Br. Epacridaceae (i). i Tasmania. Prioria Griseb. Leguminosae (u. 2). i Panama, Jamaica. Priotropis Wight et Arn. Leguminosae (in. 3). 2 E. Himal., Socotra. Prismatocarpus L'Herit. Campanulaceae (i). 158. Afr. Prismatomeris Thw. Rubiaceae (n. 9). i Ceylon, India. Pritchardia Seem, et H. Wendl. (excl. Washin^lonia H. Wendl.). Palmae (i. 2). 5 Fiji, Hawaiian Is. Pritchardiopsis Becc. Palmae (i. 2). r New Caledonia. Pritzelia Klotzsch = Begonia L. p.p. (Begon.). Pritzelia F. Muell. Philydraceae. i New Guinea. Priva Adans. Verbenaceae (i). 10 trop. and subtrop. The 1. of P. echinata Juss. are used as tea; tubers of P. laevis Juss. ed. Privet, Ligustrum vulgare L. Probletostemon K. Schum. Rubiaceae (i. 8). i Sierra Leone. Proboscella Van Tiegh. Ochnaceae. 2 trop. Afr. Proboscidea Schmid. (Alartynia p.p.). Martyniaceae. 6 warm Am. Procerus (Lat.), lofty. Process, a projecting appendage. Prochnyantb.es S. Wats. Amaryllidaceae (n). i Mexico. Prockia P. Br. ex L. Flacourtiaceae (5). 4 trop. Am. Prockiopsis Baill. Flacourtiaceae (2). i Madag. Procris Comm. ex Juss. Urticaceae (2). 5 palaeotrop. Procumbent, prostrate. Proliferous, bearing offshoots. Proliflcation, formation of buds in the axils of floral 1. Promenaea Lindl. (Zygopetalum p.p. BH.}. Orchid, (n. /3. 11). 5 Braz. Promotion nut, Anacardium accident ale L. Pronaya Huegel. Pittosporaceae. i W. Austr. Pronuba (moth), cf. Yucca. Propulsive mechanisms, cf. Dispersal of Seeds. Proscephalium Korth. Rubiaceae (n. 5). i Java. Proserpinaca L. Haloiagidaceae. 2 N. Am. g-merous. Ann. Bot. 18, 579- Prosopanche de Bary. Hydnoraceae. i Argentina. 540 PRO SO PIS Prosopis L. Leguminosae (l. 4). 30 trop. and subtrop., some xero., without 1., many thorny, the thorns being epidermal, or metamorphosed branches or stipules. P. juliflora DC. (trop. Am.) is the mezquit tree (fodder, &c.). P. alba Griseb. has sweet succulent pods (algaroba blanca), used as food. Prosopostelma Baill. Asclepiadaceae (n. t). i W. trop. Afr. Prostanthera Labill. Labiatae (n). 40 Austr. Prostnecidiscus J. Donnell Smith. Asclepiadaceae (n. i). i Guatem. Protamomum Ridl. Musaceae. i Malaya. Protandry, cf. Dichogamy. Protanthera Rafin. Liliaceae. Nomen. 2 N. Am. Protarum Engl. Araceae (vn). i Seychelles. Protea L. Proteaceae (i). 75 S. and trop. Afr. Fls. in showy heads, often with coloured bracts. Proteaceae (EP. , BH.). Dicotyledons ( Archichl. Proteales; Daphnales BH.}. 50 gen., 960 sp. "which have a very char, distr. ; there are in AustiC 591, trop. E. As. 25, New Caled. 27, N.Z. 2, Chili 7, trop. S. Am. 36, south-west Cape Col. 262, Madag. 2, Mts. of trop. Afr. 5" (Engler). The great majority live in regions where there is annually a long dry season. Correlated with this is the fact that they are mostly xero. Nearly all shrubs and trees with entire or much-divided exstip. 1., which have commonly a thick cuticle and often a covering of hairs further checking transpiration. The fls. are borne in racemes, spikes, heads, &c., and are often very showy ; many have their pollen freely exposed, though they are not wind-fert.— a peculiarity perhaps con- nected with their life in a dry climate (compare the Acacias of Australia). The fls. are usu. 5 , often -|- . P (4), corolline, valvate ; the 1. commonly bent or rolled back when open; sta. 4, inserted on the tepals, and usu. with only the anthers free; G i, ovules many or few or one, pend. or not, the micropyle facing the base of the ovary. Style term., long, often bent inwards. Follicle, capsule, drupe or nut; seed exalbum. The ovary is sometimes borne on a gynophore and at its base are commonly nectarial outgrowths. The fls. are protandrous and adapted to insect-fert. Classification and chief genera (see discussion in Nat. Pfl-Y I. P£tfSOON/OJD£A£ (fls. single \na\i\s of bracts; ovules seldom, few or 2; drupe or nut, one seeded): Persoonia, Protea, Leucadendron. II. GREVILLOIDEAE (fls. in pairs; ovules several or 2 ; fr. usu. dehisc., many seeded): Grevillea, Hakea, Bftnksia. Proteales. The i3th order of Archichlamydeae. Protection against grazing animals is obtained by stinging hairs (Loasaceae, Urticaceae, &c.), by prickles, &c., or by disagreeable taste; - of buds, see Bud ; - of honey, cf. Flower-classes ; - of flowers, see Flower-movements; - of leaves, see Movements; - of young leaves, Ainherslia, Brownea, &c. Proteids, complex organic bodies, containing C, H, O, and N. Proteocarpus'B6rner = Carex L. p.p. (Cyper.). Proteopsis Mart, et Zucc. Compositae (i). 2 campos of S. Braz. Proterandry, -ogyny, protandry, &c. Cf. Dichogamy. PSEUD-, PSEUDO- 541 Prothallus, the sexual pi. ; Pteridophyta, Filicineae, &c. Protium Barm. f. Burseraceae. 50 trop. Am. Some yield good balsam resins. Proto- (Gr. pref.), first, earliest; -corm, Lycopodiaceae; -gyny, see Dichogamy; -plasm, the living substance of pi. Protolirion Ridl. (Petrosavia p.p.). Liliaceae (i). i Mai. Penin. Protomegabaria Hutchinson. Euphorbiaceae (A. I. i). 2 trop. Afr. Protorlius Engl. Anacarcliaceae (3). 9 Madag., S. Afr. Protoschwenkia Solereder. Solanaceae (5). i Bolivia. Proustia Lag. Compositae (12). 10 Andes, temp. S. Am. Proximal, nearest to axis. Pruinose, with waxy powdery secretion on the surface. Prumnopitys Phil. = Podocarpus L'Herit. p.p. (Conif.). Prune, Primus domestica L. Prunella L. (Brnnella Tourn.). Labiatae (vi). P. vulgaris L. (self- heal) cosmop (incl. Brit.), 5 Medit. Eur. The fr. K is closed and points up in dry air, but opens and stands horiz. in damp. Prunus (Tourn.) L. (incl. Ai/iygdalus'Yovun., Cerasus Tourn.). Rosaceae (v). 85 N. temp.; a few trop. /'. insititia L (bullace), P. spinosa L. (sloe'or blackthorn), P. avium L. (gean), P. Padits L. (bird-cherry), in Brit. The fl.-buds are laid down in August or September of the preceding year. There is i cpl., which gives rise to a drupe, while the hollow recept. usu. falls away. Many sp. are cult, for their fr., e.g. P. Armeniaca L. (apricot), P. domestica L. (plum, prune), P. Amyg- dalus Stokes (almond), P. Persica Stokes (peach, with its smooth- fruited variety the nectarine), P. Cerasus L. (cherry), &c. P. Lauro- cerasus L. is the cherry laurel ; it has extra-floral nectaries on the backs of the L, showing as brownish patches against the midribs. The spines of some sp. are axillary, as in Crataegus. Pruriens (Lat.), causing itching. Przewalskia Maxim. Solanaceae (2). 2 C. As. Psacalium Cass. = Senecio Tourn. p.p. (BfJ) — Cacalia L. p.p. Psamma Beauv. =Ammophila Host. (Gram.). Psammanthe Reichb. Caryophyllaceae (inc. sed.). Nomen. Psammisia Klotzsch. Ericaceae (m. 2). 35 Andes, Guiana. Psammogeton Edgew. Umbelliferae (in. 2). 4 W. As. Psammogonum Nieuwland (Polygonella p.p.). Polygon, (n. 2). 2 N. Am. Psammomoya Diels et Loesener. Celastraceae. 2 Austr. Psammophilous, sand-loving. Psammotropha Eckl. et Zeyh. Phytolaccaceae (Aizo. Bff.). 5 S. Afr. Psathura Comm. ex Juss. Rubiaceae (n. 5). 6 Madag., Mascarenes. Psathyranthus Ule. Loranthaceae (i). i Upper Amazon. Psathyrotes A. Gray. Compositae (8). 4 W. U.S., North Mexico. Psedera Necker ex Greene (Ampelopsis, &c. p.p.). Vitaceae. 10 N. temp, and subtrop. Psednotrichia Hiern. Compositae (3). i trop. Afr. Pselium Lour. (Pericampylns BH.}. Menispermaceae. i Cochinchina. Psephellus Cass. = Centaurea L. p.p. (Comp.). Pseud-, pseudo- (Gr. pref.), false; -axis, a sympodium; -berry, Gaul- theria, Urera, Viscum ; -bulb, Orchidaceae ; -carp, the product of the 542 PSEUD-, PSEUDO- ovary, together with any other organ that developes into the fr., Pyrus, fiosa. &.c. ; -catkin, Humulus; -head, Madura, Moraceae, Urticaceae; -panicle, Humulus; -raceme, Hyperiaun, Madura, Moraceae; -spike, Moraceae, Morus ; -umbel, Hypericum, Motaceae. Pseudabutilon R. E. Fries (Abutilon p.p.)- Malv. (2). 10 warm Am. Pseudagrostistachys Pax et K. Hoffm. (Agrostistachys p.p.). Euphor- hiaceae (A n. 2). i W. Afr. Pseudais Decne. Thymelaeaceae. i Moluccas. Pseudalomia Zoll. et Morr. Compositae. Nomen. Pseudanthistiria Hook. f. (Andropogon p.p. EP.). Gramin. (2). 4 Indomal. Pseudanthus Sieb. ex Spreng. Euphorbiaceae (B. i). 7 Austr. Pseudarthria Wight et Arn. Leguminosae (in. 7). 5 Afr., trop. As. Pseuderant&emum Radlkf. Acanth. (iv. B). 60 trop. Cult. orn. fl. Pseuderia Schlechter. Orchidaceae (n. 15). 7 Malay Archipelago. Pseudibatia Malme. Asclepiadaceae (n. 4). 4 S. Am. Pseudima Radlk. (Sapindits p.p. BH.}. Sapindaceae (i). i N. trop. S. Am. Pseudixus Hayata. Loranthaceae (n). i E. As., Austr. Pseudobaeckea Niedenzu. Bruniaceae. 7 S. Afr. Pseudobahia A. Gray, Rydberg (Monolopia p.p.). Comp. (6). 2 Calif. Pseudobarleria T. Anders. (Petalidiuin BH.). Acanth. (iv. A). 15 Afr. Pseudobastardia Hassler (Sida p.p.). Malvaceae (2). 2 Brazil. Pseudoblepharis Baill. Acanthaceae (iv. B). 5 trop. Afr. Pseudobotrys Moes. Icacinaceae. i New Guinea. Pseudobravoa Rose (Bravoa p.p.). Amaryllidaceae (n). i Mexico. Pseudobraya Korshmsky. Cruciferae (4). i C. As. Pseudobromus K. Schum. (Brachydylnun p.p.). Gramin. (8). 2 Afr. Pseudocadia Harms (Cadia p.p.). Leguminosae (in. i). i trop. Afr. Pseudocalyx Radlk. Acanthaceae (in), i Madagascar. Pseudocarapa Hemsl. Meliaceae (in), i Ceylon. Pseudocarpidium Millsp. ( Vitex p.p.)- Verben. (4). 3 Cuba, Florida. Pseudocaryophyllus Berg. = Myrtus Tourn. p.p. (Myrt.). Pseudocedrela Harms (Cedrela p.p.). Meliaceae (11). 3 trop. Afr. Pseudocentrum Lindl. Orchidaceae (n. 2). 8 trop. Am., W.I. Pseudoclirosia Blume. Apocynaceae (i. 3). i New Guinea. PseudocincLona A. Chevalier ex E. Perrot. Rubi. (i. 5). i trop. Afr. Pseudoclinium O. Ktze. (Leptoclinium Gardn.). Comp. (2). i Goyaz. Pseudoconnarus Radlk. (Connarus p.p. ). Connaraceae. i Amazonas. Pseudocroton Muell.-Arg. Euphorbiaceae (A. II. 2). i C. Am. Pseudocydonia C. K. Schneider (Chaenomeles-p.p.). Ros. (n). I China. Pseudocymopterus Coulter et Rose (Cymopterus p.p.). Umbelliferae (in. 6). 5 S.W. U.S. Pseudocynometra O. Ktze. = Maniltoa Scheff. (Legum.). Pseudocytisus O. Ktze. = Vella DC. (Crucif.). Pseudodracontium N.E. Br. Araceae (iv). 3 Cochinchina. Pseudoernestia Krasser (Ernestia p.p.). Melastom. (i). i Venezuela. Pseudoeugenia Scortech. (Eugenia p.p. EP.). Myrt. (i). 2 Mai. Penin. Pseudogaltonia O. Ktze. (ffyacinttius p.p.). Liliaceae v). 2 trop. Afr. PSIDIASTRUM 543 Pseudogardneria Raciborski (Gardneria p.p.). Logan. 2 E. As. Pseudohamelia Wernham. Rubiaceae (i. 7). i trop. Am. Pseudohydrosme Engl. Araceae (iv). 2 W. trop. Afr. Pseudolachnostylis Pax. Euphorbiaceae (A. I. i). 2 trop. Afr. Pseudolarix Gord. Coniferae (Pinaceae, 21; see C. for gen. char.). i China, P. Kaeinpferi Gord., the golden pine. Like Larix, but dis- tinguished chiefly by the deciduous fruit-scales. Pseudoliparis Finet (Rlicrostylis p.p.). Orchid, (n. 4). i N. G. Pseudolmedia Tree. Moraceae (n). 5 trop. Am., W.I. Pseudolopezia Rose (l.opezia p.p.). Onagraceae (2). 2 N. Am. Pseudomacriaerium Hassler. Leguminosae (in. 8). i Paraguay. Pseudomacodes Rolfe. Orchidaceae (n. 2). i Solomon Is. Pseudomarsdenia Baill. Asclepiadaceae (it. 3). i Mexico. Pseudomorus Bur. Moraceae (i). i Norfolk I. Pseudomussaenda Wernham. Rubiaceae (i. 7). 4 trop. Afr. Pseudonephelium Radlk. (Nephelinm p.p. BH.). Sapind. (r). i Borneo. Pseudopanax C. Koch. Araliaceae (i). 6 temp. S. Am., N.Z. Pseudopavonia Hassler. Malvaceae (3). i Paraguay. Pssudopentatropis Costantin. Asclepiadaceae (n. i). i Indochina. Pseudophoenix H. Wendl. et Drude. Palmae (iv. i). i Florida. Pseudoprosopis Harms (Prosopis p.p.). Legumin. (i. 4). i trop. Afr. Pseudopteris Baill. Sapindaceae (i). i Madag. Pseudopteryxia Rydberg (Cymopterns p.p.). Umbell. (in. 5). 3 Rockies. Pseudopyxis Miq. Rubiaceae (u. 6). i Japan. Pseudoreoxis Rydberg (Cymopterus^.p.). Umbell. (111.5). 2 Rockies. Pseudorlaya Murb. (Daucm p.p.). Umbelliferae (in. 8). \ W. Eur. Pseudorobanche Rouy (Alectra p.p.). Scrophular. (in. 2). i S. Afr. Pseudosarcolobus Costantin. Asclepiadaceae (n. 3). t Indochina. Pseudosassafras Lecomte. Lauraceae (i). i China. Pseudosciadium Baill. Araliaceae (3). i New Caled. Pseudoseris Baill. (Gerbera p.p. EP.). Compositae (12). 2 Madag. Pseudosmodingium Engl. Anacardiaceae (3). 3 Mexico. Pseudosopubia Engl. (Sopubia p.p.). Scrophul. (in. 2). 5 trop. Afr. Pseudospondias Engl. Anacardiaceae (2). 2 W. and C. trop. Afr. Pseudostachyum Munro. Gramineae (13). i Himalaya. Pseudostenosiplionium Lindau (Strobilanthes p.p.). Acanthaceae (iv. A). 5 Ceylon. Pseudostonium O. Ktze. = Pseudostenosiphonium Lindau (Acan.). Pseudostreblus Bur. in DC. Moraceae (i). 2 Indomal. Pseudostriga Bonati. Scrophulariaceae (in. 2). i Cambodia. Pseudotaenidia Mackenzie. Umbelliferae (in. 6). i N. Am. Pseudotragia Pax. Euphorbiaceae (A. 11. 2). 2 S.W. Afr. Pseudotrophis Warburg. Moraceae (i). 2 New Guinea, Phil. Is. Pseudotsuga Carr. (Tsiiga Carr. p.p. BH.). Coniferae (Pinaceae, 15; see C. for gen. char.). 3 W. N. Am., E. As., incl. P. Do2tglasiiCz.rr. (P. mucronata Sudw.), the Douglas fir of W. N. Am., useful for masts, &c. Pseudoxalis Rose (Oxalis p.p.). Oxalidaceae. i Mexico. Psiadia Tacq. Compositae (3). 35 Arabia, Afr. , Madag. Psidiastrum Bello. (Eugenia p.p. EP.). Myrtaceae (l). i Porto Rico. 544 PSWIOPSIS Paidlopais Berg. Myrtaceae (i). i Venezuela. Ed. fr. Psidium L. Myrtaceae (i). no trop. Am., W.I. Many yielded, fr., e.g. P. Guajava L. , the guava. Psila Phil. Compositae (4). i Chili. Psilactis A. Gray. Compositae (3). 3 Mexico. Psilanthele Lindau. Acanthaceae (iv. B). 3 trop. S. Am., Jamaica. Psilanthus Hook. f. Rubiaceae (n. 4). 2 trop. W. Afr. Psilium (Cl.), a prairie formation. Psilobium Jack. Rubiaceae (i. 7). 2 Sumatra. Psilocarphus Nutt. Compositae (4). 4 \V. U.S., Chili. Psilocarya Torr. (Ryncospora Bff.). Cyperaceae (i). 6 Am., Austr. PsilocMlus Rodr. (Cleistes p.p. EP.). Orchid, (n. 2). i Brazil. PsUonema C. A. Mey. = Alyssum L. (Crucif.). Psilopeganum Hemsley. Rutaceae (i). i C. China. Psilostachys Hochst. (Psilotrichium £P.). Amarant. (2). 4 trop. |#. Psilostroplie DC. (Riddellia Nutt.). Compositae (6). 6 S.W. U.S. Psilotaceae. Psilotales Two gen. (Psilotum, Tmesipteris) with 4 sp. trop. and subtrop. The mature sporophyte has no roots, their functions being performed by the branched rhiz. The aerial branches bear only scale-1. in Psilotum. The sporangia are 2- or 3-loc., borne on small two-lobed sporophylls. "There has been much disagreement as to the morphological nature of the sporangiophores of the Psilotaceae. The two chief views are the following: (i) that the whole sporangio- phore is a single foliar member; (2) that it is a reduced axis bearing a terminal synangium and two 1. The recent very careful researches of Bovver upon the origin of the sporangiophore and synangium confirm the former view" (Campbell ; see also Bower in Phil. Trans. CLXXXV, 1894, p. 473). "The fully-developed synangium (2-loc. in Tmesipteris, 3-loc. in Psilotum) has the outer walls of the loculi composed of a superficial layer of large cells, beneath which are several layers of smaller ones. The cells composing the septa are narrow tabular ones; occasionally the septum is partially absent.... Bovver regards the whole synangium as homologous with the single sporangium of Lycopodium" (Camp- bell). The prothallus has not been seen. Psilotales. The 5th class of Pteridophyta. Psilotrichum Blume Amarantaceae (2). 15 palaeotrop. Psilotum Svv. Psilotaceae. 3 trop. They are probably =t saprophy tic in their habit, and have neither roots nor green L, but only green stems. See fam. for details of sporangia, &c. Veg. repr. is common, small gemmae being formed upon the rhiz. At first no structural differentia- tion is visible in these, but apical cells are formed later. Psiloxylon Thou, ex Tul. (Fropiera BH.}. Flacourt. (inc. sed.). i Masc. Psilurus Trin. Gramineae (12). i S. Eur. to Afghanistan. Psittacanthus Mart. (Loranthus p.p. BH.}. Loranthaceae (i). 50 trop. Am. Psophocarpus Neck. Leguminosae (ill. 10). 5 palaeotrop. P. tetra- gonolobus DC. and others cult. ed. pods. Psoralea L. Leguminosae (in. 6). 100 trop. and subtrop. P. esculenta Pursh (N. Am. prairie turnip) has ed. tuberous root. PTERIDOPHYTA 545 Psorospermum Spach. Guttiferae (11). 20 trop. Afr. , Madag. Psychine Desf. Cruciferae (2). i N. Afr. Psychotria L. (excl. Mapourea Aubl.). Ruhiaceae (n. 5). 400 trop. Some heterostyled. Many have infl.-axis brightly coloured. [For P. Ipecacuanha Stokes see Uragoga.] Psychrobatia Greene (Rubus p.p.). Rosaceae (in. 2). i N.W. Am. Psychrogeton Boiss. (Aster -p.p. BH.}. Compositae (3). i Afghanistan. Psychrophyton Beauverd (Raoulia p.p.). Compositae (4). 9 N.Z. Psyllium Tourn. ex Juss.= Plantago L. p.p. (Plantag. ). Psyllocarpus Mart. Rubiaceae (n. 10). 5 Brazil. Psyllothamnus Oliv. Caryophyllaceae (i. 4). i Aden. Ptaeroxylon Eckl. et Zeyh. Meliaceae (i) (Sapind. BH.}. \ S. Afr., P. ittilc: E. et Z. (sneeze wood, cape mahogany), timber. Ptarmica (Tourn.) Neck. = Achillea L. p.p (Comp.). Ptelea L. Rutaceae (iv). 60 N. Am. P. trifoliata L. cult. orn. shrub (shrubby trefoil). Fls. monoec. Fr. winged (if. Ulmus). Pteleocarpa Oliv. Boragin. (n) (Olacin. BH.}. 2 Malay Pen., Borneo. Ptelidium Thou. Celastraceae. i Madag. Pteleopsis Engl. Comhretaceae. 4 E. trop. Afr. Pteralyxia K. Sclium. ( Vallesia p.p.)- Apocyn. (i. 3). i Hawaiian Is. Pterandra A. Juss. Malpighiaceae (n). i Brazil. Pteranthus Forsk. Caryophyllaceae (i. 6). r Medit. Pterichis Lindl. Orchidaceae (n. 2). 6 trop. S. Am. Pteridium Gled. i cosmop., P aquiliniini Kuhn, the bracken. It has a creeping rhiz. , bearing 2 ranks of 1. At the base of the 1. is a nectary, which ants visit. The sori are confluent along the 1. margin, which is curved over them, while there is also a true indusium on the inner side of the sori (the char, of distinction from Pteris, in which the bracken used to be placed). Adv. buds appear on the back of the 1. stalk, near the base. Pteridocalyx Wernham. Rubiaceae (i. 3). i Brit. Guiana. Pteridophyllum Sieb. et Zucc. Papaveraceae (i). i Japan. Pteridophyta Vasailnr Cryptogams, one of the four chief divisions of the Vegetable Kingdom. They have a well marked alternation of generations, the gametophyte (oophyte or sexual generation) being insignificant in size compared to the sporophyte (asexual generation), but still capable of independent growth. The life history of a typical Pteridophyte may be shown diagrammatically as follows, taking the fern as an example : Fern-plant -*• sporophylls -*• sporangia ->• spores t f I spermatozoid •*- antheridia ) fertilised ovum •*- •{ .}••*- prothallus. | ovum -*- archegonia | The 'plant' or asexual generation alternates with the prothallus or sexual generation. In many P. there are 3 and 9 prothalli. The pro- thallus corresponds to the 'plant' in a moss or liverwort, whilst the sporogonium of these latter = 'plant' in a fern or lycopod. The plant itself takes various forms in the different groups. Except in tree ferns and in fossil forms it does not attain great size. There is an erect stem in many ferns, &c. ; others have creeping stems (e.g. Lycopodium and Selaginella), rhiz. (many Ferns), or floating w. 35 546 PTERIDOPHYTA stems (Hydropterideae). The 1. are simple, except in many ferns. There is no primary tap-root, but roots are formed as required from the stem or 1. Internally there are well marked vascular bundles in both stem and leaf, and many anatomical features of fig. pi. may be found here also. The sporangia arise upon the 1., either on the ordinary foliage as in most ferns, or on specially differentiated 1. as in Osmunda, Equisetum, Lycopodiales Hydropteridineae, &c. They may be sol. or in groups (son) ; in the latter case often protected by a special outgrowth of the 1., the indusiinn. The spores are formed by a com- plicated process from a single cell or row or layer of cells — the archesporium — inside the sporangium : each has a thick waterproof outside wall. The spores maybe of one kind only, in which case the plant is termed homo- or iso-spomus, or of two kinds (heterosporous}, when the smaller spore is termed the microspore, the larger the mega- ox macro-spore ; the former gives rise to a cf prothallus, the latter to a? . Falling upon the soil (or into the water, in the case of the Hydropteridineae) the spores germinate under suitable conditions, giving rise to the sevual plants or prothalli. The prothallus is a small body without distinction into stem and 1. ; it absorbs materials from the soil (usually by rhizoids) and, being green, assimilates in the ordinary way. It bears the sexual organs — antheridia ( 3 ) and archegonia ( ? )• In the homosporous forms these are both found on the same prothallus, except in Equisetum, where, though the spores are absolutely similar so far as we can tell, there are separate 3 and 9 prothalli, as in the heterosporous forms. In the antheridia are developed the motile 3 cells or spermatozoids. Pert, takes place by aid of water. The mucilage contained in the neck of the archegonium is attractive to the spermatozoids, which swim up the neck of the archegonium. One of them finally fuses with the ovum or ? cell at the base of the archegonium, and the fertilised ovum (zygote) then develops into a new 'plant' or asexual generation, being nourished by the pro- thallus until it can assimilate for itself. Classification (after Engler): I. Filicates. L. usu. more strongly developed than stem, often big and much branched, usu. circinate in vernation. Spo- rangia on normal or special 1., on edge or lower side, usu. in sori. Spermatozoid polyciliate. 1. Filicales Leptosporangiatae. 2. Marattiales. 3. Ophioglossales. [II. Sphenophyllales. Fossil plants only.] III. Equisetales. Homosporous (some fossils heterosporous), with dioec. prothalli. Stem much branched with jointed internodes and small sheathing whorls of 1. (not green). Sporangia on peltate sporangiophores, forming a term, spike. Spermatozoids polyciliate. IV. Lycopodiales. Homosporous or heterosporous ; in the latter case the ? prothallus remains enclosed in the spore till fert. Stem simple or branched ; 1. many, small, usu. alt., entire. PTEROPYRUM 547 Sporangia singly on upper side of leaf-bases, or in their axils. Spermatozoids biciliate. i. Lycopodiales eligulatae. •2. Lycopodiales ligulatae. V. Psiloiales. Sporophylls bipartite. Sporangia 2 — 3'loc. Spermatozoids biciliate. VI. IsoetaUs. Stem short, growing in thickness, with numerous long 1. with ligula over a basal groove, in which is a sporangium ; megaspures on outer sporophylls, micro- spores on inner. Spermatozoids polyciliate. [VII. Cycadojilices . Fossil plants only.] Pterigeron A. Gray. Compositae (-(). 7 Austr. Pterigostachyum Nees ex Steud. (Dimeria R. Br. ). Gram. (2). 12 S.E. As. Pteris L. Polypodiaceae. 160 cosmop. [For P. aquilina L., the bracken, see Pteridium.] Apogamy occurs in P. crelica L. (see Filicales Leptosporangiatae). Pterisanthes Blume. Vitaceae. 12 trop. As. Pternandra Jack. Melastomaceae (ii). 6 Malaya. Pternopetaluin Franch. Umbelliferae (ill. 5). i Tibet. Ptero- (Gr. pref.), wing. Pterocactus K. Sebum. Cactaceae (n). 3 Argentina. Pterocarpus L. Leguminosae (in. 8). 24 trop. Fruit winged. Several, esp. P. Marsnfium Roxb., furnish kino, an astringent resin. P. santalimis L. f. yields red sandal- wood. Pterocarya Kunth. Juglandaceae. 4 N. temp. j^. Pterocaulon Ell. Compositae (4). 15 Argentina to Texas. Pterocelastrus Meissn. Celastraceae. 7 Cape Colony. Pteroceltis Maxim. Ulmaceae. i N. China. Pterocephalus Vaill. ex Adans. (Scabiosa p.p. BH.). Dipsacaceae. 20 Medit., Indomal., trop. Afr. Pterochrosia Baill. Apocynaceae (i. 3). i New Caled. Pterocladon Hook. f. Melastomaceae (i). i Peru. Pterococcus Pall. = Calligonum L. p.p. (Polygon.). Pterocoelion Turcz. Tiliaceae (inc. sed. ). i Java. Pterocymbium R. Br. (Stfi-fidia p.p. BH.}. Stercul. 5 Burma, Malaya. Pterodiscus Hook. Pedaliaceae. 10 S. and trop. Afr. Pterodon Vog. Leguminosae (in. 8). 4 Brazil, Bolivia. Pterogastra JS'aud. Melastomaceae (i). 3 N. trop. S. Am. Pteroglossaspis Reichb. f. Orchidaceae (11. 10). 3 E. trop. Afr. Pterogonium Fee. Polypodiaceae. 2 Guiana. Pterogyne Tul. Leguminosae (n. 2). i Brazil. Pterolepis Miq. Melastomaceae |i). 30 trop. Am. Pterolobium R. Br. (Cantuffa EP.}. Leguminosae (n. 7). 5 palaeo- trop. Pteronema Pierre. Simarubaceae. i Burma. Pteroneurum DC. =Cardamine Tourn. p.p. (Crucif.). Pteronia L. Compositae (3). 60 S. Afr. Pteropetalum Pax (huadenia p.p. EP.}. Capparid. (n). i Togoland. Pteropogon DC. =Helipterum DC. p.p. (Comp.).. Pteropyrum Jaub. et Spach. Polygonaceae (n. i). 5 S.W. As. 35-2 548 PTER OR HA CHIS Pterorhachis Harms. Meliaceae (in), i Cameroons. Pteroscleria Nees. Cyperaceae (n). 3 trap. Am., W. I. Pterosicyos T. S. Brandegee. Cucurhitaceae (4). i Mexico. Pterospermum Schreb. Sterculiaceae. 20 trop. As. Pterospora Nutt. Pyrolaceae. i N. Am. Pterostegia Fisch. et Mey. Polygonaceae (i. i}. i California. Pterostemma Kranzl. Orchidaceae (n. 19). i Colombia. Pterostemon Schau. Saxifragaceae (iv). i Mexico. Pterostigma Benth. = Adenosma R. Br. p.p. (Scroph.). Pterostylis R. Br. Orchidaceae (n. 2). 45 Austr., N.Z. , New Caled. The median sepal, with the petals, forms a hood over the rest of the fl. The flap of the labellum hangs out below and is irritable. If an insect land on it, it instantly moves up and imprisons the visitor against the column ; the only mode of escape is by squeezing past the stigma and anther. After half-an-hour the lip goes down again and is ready for another capture (Darwin, Orchids, p. 86). Pterostyrax Sieb- et Zucc. (Halesia p.p.). Styrac. 3 Japan, China. Pterotaberna Stapf ( Tabernaemontana p.p.). Apocyn. (i. 3). i W. Afr. Pterotlaeca Cass. Compositae ( 13). 10 Medit., W. As. Pterothrix DC. Compositae (4). 3 Cape Colony. Pterotropia Hillebrand. Araliaceae (i). 3 Hawaiian Is. Pterotum Lour. Inc. sed. i Cochinchina. Pteroxygonum Dammer et Diels. Polygonaceae (n. 2). i China. Pterygiella Oliv. Scrophulariaceae (in. 3). 2 China. Pterygodiurn Sw. Orchidaceae (n. i). 158. Afr. Pterygopappus Hook. f. Compositae (4). i Tasmania. Pterygopodium Harms. Legurninosae (ill. 8). i Cameroons. Pterygota Schott. et Endl. (Sterciilia p.p. BH.}. Stercul. 4 trop. %. Pteryxia Nutt. (Cymopterns p.p-)- Umbelliferae (in. 5). 7 N. Am. Ptilimnium Rafin. (Discopleura p.p.). Umbell. (ill. 5). 3 N. Am. Ptilocalais A. Gray, Greene (Microseris p.p.). Comp. (13). i N. Am. Ptilochaeta Turcz. Malpighiaceae (i). 3 Brazil, Argentina. Ptilotrichum C. A. Mey. (Alyssum p.p. BH.}. Cruciferae (4). 10 Medit. Ptilotus R. Br. Amarantaceae (2). 60 Austr. Ptychandra Scheff. Palmae (iv. i). 2 Moluccas. Ptychanthera Decne. Asclepiadaceae (n. 4). i S. Domingo. Ptychococcus Becc. Palmae (iv. i). 3 New Guinea, .Moluccas. Ptychogyne Pfitz. (Panisca p. p.). Orchid, (n. 3). 2 Mai. Pen., Java. Ptychomeria Benth. =Gymnosiphon Blume (Bunnann). Ptychopetalum Benth. Olacaceae. 6 trop. Am., W. Afr. Ptychopyxis Miq. Euphorbiaceae (inc. sed.). i Malaya. Ptychoraphis Becc. Palmae (iv. i). 3 Malaya. Ptycbosema Benth. Leguminosae (ill. 6). i W. Austr. PtychospermaLabill. Palmae (iv. i). 15 E. Indomal. Fls. in threes, 2 (f and i ? . P. (Seaforthia) elegans Blume cult. orn. Ptychotis Koch (Carum p.p. BH.}. Umbelliferae (ill. 5). 6 Eur., Afr. Ptyssiglottis T. Anders. Acanthaceae (iv. B). 5 Indomal. Puberulous, minutely pubescent. Pubescent, covered with fine soft hairs, downy. Puccinia, if. Malva. PURGING BUCKTHORN 549 Puccoon (Am.), Lithospermum, Sanguinaria ; yellow-, Hydrastis. Pudding-pipe tree, Cassia. Puelia F ranch. Gramineae (13). 4 W. trop. Afr. Pueraria DC. Legmninosae (in. 10). 15 trop. As. to Japan. Fibre. Pugionium Gaertn. Cruciferae (4). 2 Mongolia. Pugiopappus A. Gray (Leptosyne BH., Coreopsis p.p. EP.}. Compositae (5). 3 California. Pulchellus (Lat.), beautiful. Pulicaria Gaertn. Compositae (4). 30 Eur., As., Afr. P.dysenterica Gaertn. (Inn la dysenterica L.) in Brit, (flea-bane). Pullea Schlechter. Cunoniaceae. 2 New Guinea. Pulmonaria (Tourn.) L. Boraginaceae (IV. 3). 10 Eur. P.officinalis L. (lung-wort, formerly officinal) and P. angustifolia L. in Brit. Both have dimorphic heterostyled fls. which change from red to blue as they grow older (see fam.). Pulque, Agave. Pulsatilla (Tourn.) L. = Anemone Tourn. p.p. (Ranunc.). Pultenaea Sm. Leguminosae (in. 2). 80 Austr. Pulvinaria Fourn. Asclepiadaceae (n. i). i Brazil. Pulvinate, cushion-shaped. Pulvinus, a swollen joint, often concerned in movement, Galeopsis, Marantaceae, Mimosa, Neptitnia. Pumelo, Citrus decuinana Murr. Pumilus (Lat.), low, small. Pumpkin, Cncnrbita Pepo L. Punctate, clotted. Punctum vegetationis (Lat.), growing point. Pungens (Lat.), sharp-pointed. Punica (Tourn.) L. Punicaceae (only genus). 2, one in Socotra, the other, P. Granatum L., the pomegranate, from the Balkans to the Himalayas, and cult. The young twigs have four wings, composed simply of epidermis and cortical parenchyma ; these are early thrown off. The fl. is 5 i reg., perig. K 5 — 8, valvate ; C 5 — 8, imbr. ; A oo ; G adnate to receptacle. The mature ovary has a peculiar struc- ture, due to a development like that in Mesembryanthemum. Two whorls of cpls. with basal plac. are laid down, and then a peripheral growth tilts them up from || • || to =•= so that two layers of loculi are formed and the placentation appears to be parietal. Ovules oo , anatr. The arrangement is also seen in the fr. , commonly termed a berry, but not strictly so. The pericarp (axial in part) is leathery, and the fleshy inner part round the seeds is really the outer layers of the seed coats. Punicaceae (EP.\ Lythraceae p.p. BH.}. Dicots. (Archichl. Myrtiflorae). Only genus Punica (q.v.). Puniceus (Lat.), bright carmine. Pupalia Juss. Amarantaceae (2). 7 Afr., Madag., As. Pupunha palm, Bactris. Purdiaea Planch. (Costaea EP.}. Cyrillaceae. 3 Cuba, Colombia. Purdieanthus Gilg. Gentianaceae (i). i Colombia. Purging buckthorn, Rkamnuscatkartical^.; -cassia, Cassia Fistula L.; - flax, Linum catharticum L. 55° PURGOSEA Purgosea Haw. =Crassula L. (Crassul.). Purple heart (W.I.), Copaifera; -lip (W.I.), Vanilla; -loosestrife, Lythrum Salicaria L.; - top (Austr.), Verbena. Purpurella Naud. (Tibouchina BH.}. Melnstomaceae (8). 12 trop. S. Am. Purpureus (Lat.), purple. Purpusia T. S. Brandegee. Rosaceae (in. 2). i N. Am. PursMa DC. Rosaceae (in. 3). i Pac. U.S. Purslane, Porlnlaca; sea-, Arcnana p:ploides L. Pusaetha L. (Entada Aclans.). Leguminosae (l. 5). 12 trop. Puschkinia Adams. Liliaceae (v). 2 W. As. Cult. orn. fl. Pusillus (Lat.), small, weak, slender. Pustule, a pimple or blister. Putamen, the stone of a drupe. Putoria Pers. Rubiaceae (n. 7). 3 Medit. Putranjiva Wall. Euphorbiaceae (A. i. i). 4 Indomal. Putterlickia Endl. Celastraceae. 2 Cape Colony. Puya Molina. Bromeliaceae (2). 25 Andes. Some 3 in. high, thick stem. Pycnandra Benth. Sapotaceae (i). i New Caled. Pycnanthemum Michx. Labiatae (vi). 15 N. Am. Pycnanthus Warb. (Myristica p.p.). Myristicaceae. 5 trop. Afr. Pycnarrliena Miers. Menispermaceae. 20 E. Indomal. Pycnobotrya Benth. Apocynaceae (n. i). i Gaboon. Pycnobregma Baill. Asclepiadaceae (n. 4). i Colombia. Pycnocoma Benth. Euphorbiaceae (A. n. 2). 12 trop. Afr., Masc. Pycnocomon Hoffmgg. et Link (Scabiosa p.p. BH.}. Dipsac. 2 Medit. Pycnocycla Lindl. Umbelliferae (in. i). 7 Abyssinia to N.W. India. Pycnoneurum Decne. Asclepiadaceae (11. i). 2 Madag. Pycnophyllum Remy. Caryophyllaceae (i. 3). 3 Andes. PycnorhacMs Benth. Asclepiadaceae (n. 3). i Malay Peninsula. Pycnosphaera Gilg (Faroa p.p.). Gentianaceae (i). 2 trop. Afr. Pycnospora R. Br. ex Wight et Am. Legumin. (ill. 7). i trop. As., Austr. Pycnostachys Hook. Labiatae (vn). 15 trop. and S. Afr., Madag. Pycnostelma Bunge ex Decne. Asclepiadaceae (n. i). 2 China. Pycnostylis Pierre. Menispermaceae. i trop. Afr. Pycnothymus Small (Satureia p.p.). Labiatae (vi). i N. Am. Pycreus Beauv. (Cy ferns p.p. EP.). Cyperaceae (i). 100 cosmop. Pygeum Gaertn. Rosaceae (v). 20 palaeotrop. Pygmaeopremna Merrill. Verbenaceae (4). i Luzon. Pygmaeus (Lat.), dwarf. Pynaertia De Wild. Meliaceae (in), i Congo. Pyramia Cham. Melastomaceae (i). 3 S. Brazil. Pyramidium Boiss. Cruciferae (4). i Afghanistan. Pyramidocarpus Oliv. Flacourtiaceae (i). i W. trop. Afr. Pyramidoptera Boiss. Umbelliferae (in. 5). i Afghanistan. Pyramidostylium Mart. = Salacia L. (Hippocrat.). Pyrecnia Nor. Inc. sed. Nomen. Pyrenacantha Wight. Icacinaceae. 10 S. and trop. Afr., W. As. Pyrenaria Bin me. Theaceae. i c Indomal. QUAKE-GRASS 551 Pyrene, a single stone of a drupe. Pyrethrurn Hall. = Chrysanthemum Tourn. (Comp.). Pyriform, pear-shaped. Pyrola (Tourn.) L. (incl. Moneses Salisb.). Pyrolaceae. i=. N. temp. (5 Brit., wintergreen). Evergreens with creeping stocks. P. (Moneses) itniftora L. has adv. buds on the roots, and a solitary term. fl. The fls. of 'P. minor L. are in racemes, pend., without discs. There is no honey ; the stigma projects beyond the anthers, but pollen may at last fall upon it from them. P.-rotundifolia L. is similar. The seeds of P. are very light and are distr. by wind. Pyrolaceae (EP.; Ericaceae p. p. , Monotropeae BH.}. Dicots. (Sympet. Ericales). 10 gen., 30 sp., cold N. temp, and arctic. The two Brit, genera represent the two types of habit found in the order — evergreen plants with sympodial growth from rhiz. (Pyrola), and saprophytes (Monotropa). The infl. is term.; it may be a true raceme (Pyrola), or a cyme, leafless or with scaly bracts. Fl. 5 actinom. K 4 — 5 ; C (4 — 5) or 4 — 5; A 8 — 10, obdiplost. ; G (4 — =,). The petals and sta. are often at the edge of a nectariferous disc. Anthers intr., opening by apical pores or transv. valves; pollen simple or in tetrads. Cpls. opp. petals; ovary imperfectly 4 — 5-loc. Style simple; ovules minute, oo , anatr. , on thick fleshy plac. Capsule. Seeds oo , small, in loose testa. Embryo of few cells, without differentiation of cotyle- dons. Chief genera : Pyrola, Chimaphila, Monotropa, Sarcodes. BH. unite Pyrola and the green-leaved forms to Ericaceae, making a fam. Monotropeae for the saprophytes. Pyronia x Veitch. Rosaceae. Hybrid, Pyrus-Cydonia (pear and quince). See_/0«;'M. of Hered. 1916, p. 416. Pyrostegia C. Presl (Bignonia p.p. BH.}. Bignon. (i). 10 S. Am. Pyrostria Comm. ex Juss. Rubiaceae (ll. i). 8 Mauritius, Rodrigues. Pyrrhopappus DC. (Sitilias Rafin.). Compositae (13). 6 N. Am. Pyrrocoma Hook. (Aplopappus Cass.). Compositae (3). 20 N. Am. Pyrularia Michx. Santalaceae. 2 N. Am., Himalaya. Pyrus (Tourn.) L. (BH. incl. Cydonia, Mespilus}. Rosaceae (n). 65 N. temp.; 6 in Brit., incl. P. Aitcnparia Ehrh., the rowan or mountain ash, P. Alalus L. the apple, &c. The recept is hollowed out and united to the syncarpous ovary. The fls. are protogynous. and are visited by bees and many other insects. Several var. of pear (P. cominunis L.) are self-sterile. After fert. the fr. becomes a large fleshy pseudocarp (pome), the flesh consisting of the enlarged recept., while the gynaeceum forms the core. Several are cultivated for their fruit, e.g: P. Mat 'us L. (apple), P. cominunis L. (pear), P. germanica Hook, f. (medlar). P. japonica Thunb. is often grown upon walls. Pyxidantliera Michx. Diapensiaceae. i E. U.S. Pyxidanthus Naud. (Blakea BH.}. Melastomaceae (i). 3 N. trop. S. Am. Pyxis, capsule opening by a lid that splits off, Anagallis. Quadrania Nor. Inc. sed. Nomen. Quadrasia Elmer. Flacourtiaceae (4). i Phil. Is. Quadri- (Lat. pref.), four. Quaternate, arranged in fours. Quake-grass, Briza. 552 QUA LEA Qualea Aubl. Vochysiaceae. 32 trop. Am. Quamash, Camassia. Quamoclidion Choisy (Mirabilis p.p.)- Nyctaginaceae. 4 N. Am. Quamoclit Tourn. ex Moench (Ipomoea p.p. BH.}. Convolvulaceae (i). 12 trop. Am. Cult. orn. fl. Quapoya Auhl. (Clusin p.p. BH.}. Guttiferae (iv). 3 Guiana, Peru. Quaqua N.E. Br. (Caralhinia EP.}. Asclepiadaceae (n. 3). i S. Afr. Quaquaversal, bending every way. Quararibea Aubl. Bombacaceae. 7 warm Am. Quartinia Endl. (Rotala p.p. EP.}. Lythraceae. i Abyssinia. Quassia L. Simarubaceae. 2 N.E. Brazil, W. Afr., Q. amara L. (Am.), is the source of quassia wood. Quassia bark, Picraena, Quassia ; - wood, Quassia. Quebec oak, Qitercus alba L. Quebrachia Griseb. (Schinopsis EP.}. Anacardiaceae (3). 5 S. Am. Quebracho, Schinopsis, Aspidosperma. Queen-of-the-meadow, Uimaria Spiraea-Ulmaria Hill. Queensland nut, Macadamia termfolia F. Muell. Quekettia Lindl. Orchidaceae (n. 19). 3 Brazil. Quelcliia N.E. Br. Compositae (12). i Roraima (Brit. Guiana). Queltia Salisb. = Narcissus Tourn. p.p. (Amaryll. )• Quercitron bark, Qiic-rcus tinctoria Bartr. Quercus (Tourn.) L. (BII. incl. Pasania Oerst.). F^agaceae. 300 N. temp., Indomal., Pacific coasts, &c. The oaks are evergreen or deciduous trees, in the latter case esp. with well-developed winter buds. The cupule contains i ? fl. only (see fain.), and forms the acorn- cup at the base of the nut in fr. Thea. Retiniphyllum Humb. et Bonpl. Rubiaceae (i. 8). 10 trop. S. Am. Retinispora Sieb. et Zucc. = Thuya L., &c. Seedlings of many sp. of the genera Chamaecyparis, Cupressus, Thuya, &c., exhibit, instead of the decussate appressed 1. of the mature plant, spreading needle-1. (often in whorls of 4) like those of Abies, &c. (cf. Pinus, Acacia, &c.). If now these young seedlings be used as offsets, the new pi. thus formed retain throughout life this form of foliage; and pi. are thus obtained of totally different habit from that usual in these genera. To these 'seedling forms' the name R. was given. Many in gardens. The synonymy of some is: If. decussata hort. = Thuya orientalis ; R. filifera Fowles = Cupressus obtusa; K. jimiperoides Carr. = Thuya orientalis; R. obtusa Sieb. et Zucc. = Cupressus obtusa; R. pisifera W. 36 562 RETINISPORA Sieb. et Zucc. — Cupressus pisifera; R. recurvata hort. and R. rigida Carr. = Thuya orientalis ; R. squarrosa Sieb. et Zucc. and R. stricta hort. = Cupressus pisifera. For further synonymy see Index Kewensis. Retinodendropsis Heim. Dipterocarpaceae. i Borneo. Retting, rutting away the softer tissues, cf. Linum, Cocos. Retzia Thunb. Loganiaceae (Solan. BH.). i S. Afr. Reussia Endl. Pontederiaceae. 2 S. Am. Reutera Boiss. = Pimpinella Riv. p.p. (Umbell.). Reverchonia A. Gray. Euphorbiaceae (A. i. i). i Texas. Reya O. Ktze. (Burchardia R. Br.). Liliaceae (i). i Austr., Tasm. Reyesia Clos (Salpiglossis p.p. EP.). Solanaceae (5). i Chili. Reynaudia Kunth. Gramineae (6). i Cuba, S. Domingo. Reynoldsia A. Gray (Trevesia BH.). Araliaceae (i). 4 Polynesia. Reynosia Griseb. Rhamnaceae. 5 W.I., Florida. Reynoutria Houtt. Inc. sed. i Japan. Rhabdadenia Muell.-Arg. Apocynaceae (11. i). 7 trop. Am., W.I. Rhabdia Mart. Boraginaceae (n). i trop. Rhabdodendron Gilg et Pilger. Rubiaceae (inc. sed.). 7 Amazon. Rhabdophyllum Van Tiegh. (Oiiratea Aubl.). Ochnac. 25 trop. Afr. Rhabdosciadium Boiss. Umbelliferae (ill. 2). 3 Persia. Rhabdostigma Hook. f. Rubiaceae (n. i). i E. Afr. Rhabdothamnopsis Hemsl. Gesneriaceae (i). i China. Rhabdothamnus A. Cunn. Gesneriaceae (i). i New Zealand. Rhabdotheca Cass. = Launaea Cass. p.p. (Comp.). Rhachicallis DC. Rubiaceae (i. 3). i W.I. Rhachidospermum Vasey (Jouvea p.p. EP.). Gramineae (12). i Calif. Rhachis, rachis ; rhaphe, raphe. Rhacodiscus Lindau. Acanthaceae (iv. B). 3 S. Am. Rhacoma Adans. (Leuzea BH. , Centanrea p.p.). Comp. ( 1 1). 8 W.I., S. Am. Rhacoma L. (Myginda BH.). Celastraceae. 15 W.I., trop. Am. Rhadamanthus Salisb. Liliaceae (v). 2 S. Afr. Rhagadiolus Tourn. ex Scop. (BH. incl. Hedypnois and Garhadiolus). Compositae (13). i Medit. Fr. linear, without pappus, completely enwrapped in an involucral bract. Rhagodia R. Br. Chenopodiaceae (A). 1 2 Austr. Rhamnaceae (EP., BH.). Dicots. (Archichl. Rhamnales; Celastrales BH.). 40 gen., 500 sp., cosmop. Mostly trees or shrubs, often climbing (by aid of hooks in Ventilago, tendrils in Gouania, &c., twining stems in Berchemia): thorns occur in some, and especially in Colletia, &c. (q.v.). In these pi. too, serial buds occur in the 1. -axils. L. simple, usu. with stip., never lobed or divided. Infl. cymose, usu. a corymb. Fl. inconspic., 5 or rarely unisexual, reg., sometimes apet. Recept. hollow, free from or united to the ovary. K 5 — 4, valvate; C 5 — 4, usu. small, often strongly concave, frequently clawed at base ; A 5 — 4, •alt. with sepals, usu enclosed by the petals, at any rate at first. Disc usu. well developed, intra-staminal ; G free or ± united to recept. , 3 — 2- (rarely by abortion i-) loc. (sometimes 4- or typically i-loc); in each loc. i (rarely 2) basal ovule with downwardly-directed micropyle; style simple or divided. Fr. dry, splitting into dehisc. or indeh. RHAPHISPERMUM 563 mericarps, or a drupe with i or several stones, or a nut. Endosp. little or none. Many of the dry fr. show special adaptations for wind- carriage, e.g. Paliurus, Ventilago. Closely related to Vitaceae, from which it is chiefly distinguished by the small petals, the recept., the endocarp and simple 1. ; it also approaches Celastraceae, the chief distinction being the antepetalous sta. Few are of economic value; see Zizyphus,' Rhamnus, Hovenia. Chief genera: Ventilago, Paliurus, Zizyphus, Rhamnus, Hovenia, Ceanothus, Phylica, Colletia, Gouania. Rhamnales. The 25th order of Archichlamydeae. Rhamneae (£//.) = Rhamnaceae. Rhamnella Miq. (Microrhaninus BH.}. Rhamnaceae. i Japan, China. Rhamnidium Reissek. Rhamnaceae. 5 trop. S. Am., W.I. Rliamnoneuron Gilg. Thymelaeaceae. i Tonquin. Rhamnus Tourn. ex L. Rhamnaceae. 75 cosmop. , 2 in Brit., R. cathartica L. (common or purging buckthorn) and R. Fraugula L. (alder buckthorn). Shrubs with alt. or opp. 1. and small cymose clusters of fls. Two sections. To § i, Eiirhamnus (fls. usu. 4-merous, polyg. or dioec.), belong R. A latent us L. (Medit.) and A', cathartica (Eur., As., Medit.), whose berries are purgative; the juice of the fr. is mixed with alum and evaporated, thus forming the paint known as sap-green; also R. infectoria L. (Mts. of S. Eur.) whose berries (graines d'Avignon or 'yellow berries') yield useful green and yellow dye-stuffs, and R. chlorophora Decne. from whose bark the Chinese prepare the dye known as 'Chinese green indigo' used in dyeing silk {R. utilis Decne. is also employed). To § 2, Fraugula (fls. .usu. 5-merous, $), belong R. Fraugula (Eur., As., N. Afr.) whose bark is officinal (cathartic) and whose wood forms one of the best charcoals, R. Purshiana DC. in N. Am., whose bark (Cascara sagrada) is largely used as a cathartic, &c. Rhamphicarpa Benth. Scrophulariaceae (in. 2). 8 palaeotrop. Rhamphldia Lindl. = Hetaeria Blume (Orchid.). Rhamphogyne Sp. Moore (Abrotanellap.^.). Compos. (3). i Rodrigues. Rhanterium Desf. Compositae (4). 3 N.W. Afr. to Beluchistan. Rhapbanistrocarpus Pax. Cucurbitaceae (3). i Mombasa. Rhaphanocarpus Hook. f. Cucurbitaceae (3). 2 trop. E. Afr. Rhaphanus L. (Raplianus). Cruciferae (2). 10 Medit., Eur., Java. Rhaphidanthe Hiern. Ebenaceae. i trop. W. Afr. RhapMdiocystis Hook. f. Cucurbitaceae (3). 3 trop. Afr., Madag. Rhaphldophora Hassk. Araceae (n). 60 Indomal. Sympodial climbing stems with clasping roots and pend. aerial roots. The pin- nation of the 1. arises like that in Monstera, i.e. by long holes arising between the ribs, and the margin finally breaking. Fls. 5 • Rhaphidorhynchus Finet (Angraecum p.p.). Orchidaceae (n. 20). 22 trop. Afr., Madag., Masc. Rhaphidospora Nees (Jitsticia p.p. Bff.). Acanth. (iv. B). 6 palaeo- trop. Rhaphiolepis Lincll. Rosaceae (n). 4 subtrop. E. As. Rhaphlophallus Schott (Amorphophallns^.p.). Araceae (iv). i S. Ind. Rhaphiostylis Planch. (Apodytes BH.). Icacinaceae. 7 trop. W. Afr. RhapMs Lour. = Chrysopogon Trin. = Andropogon L. p.p. RhapMspermum Benth. Scrophulariaceae (in. 2). i Madag. 36—2 564 RHAPHITAMNUS Rhaphitanimis Miers. Verbenaceae (i). i Chili. Rhapidophyllum H. Wendl. et Drude. Palmae (i. 2). i S.E. U.S. Rhapis L. f. Palmae (i. 2). 5 E. As. Cult. orn. Rhaponticum (Vaill.J Hall. =Centaurea L. p.p. (Comp.). Rhaptonema Miers. Menispermaceae. i Madag. Rhaptopetalum Oliv. Scytopetalaceae (Olacin. BH.}. 5 trap. Afr. Rhatany root, Krameria. Rhazya Decne. Apocynaceae (i. 3). 2 W. As. Rliea. Boehnieria nivea Gaudich. Rheedia L. Guttiferae (v). 17 trop. Am., Madag. Rhektophyllum N'.E. Br. Araceae (iv). i trop. W. Afr. Rheotropism, sensitiveness to water stimulus. Rhetinodendron Meissn. Compositae (8). i Juan Fernandez. Rhetinosperma Radlk. Sapindaceae (i). i Queensland. Rheum L. Polygonaceae (I. 2). 20 temp, and subtrop. As. Fls. like Rumex, but coloured and entomophilous, though they exhibit traces of anemophily in very large stigmas (cf. Poterium, &c.). R. ojficinale Baill. furnishes medicinal rhubarb; R. Rhaponticum L. is the rhubarb used as a vegetable. Rhexia L. Melastomaceae (i). 7 E. U.S. Rhigiocarya Miers. Menispermaceae. i trop. W. Afr. Rhigiophyllum Hochst. Campanulaceae (i. i). i S. Afr. Rhigospira Miers (Tabernaemontana L.). Apocynaceae (i. 3). 7 Brazil. Rnigozum Burch. Bignoniaceae (2). 5 S. Afr. Rhinacanthus Nees. Acanthaceae (iv. B). 6 palaeotrop. Rhinanthus L. (incl. Fistularia L.). Scrophulariaceae (in. 3). 9 Eur., Medit., M. Am.; 2 in Brit, (yellow-rattle), common in damp pastures. Semi-parasites with loose pollen fls. (see fam.). Rhinopteryx Niedenzu. Malpighiaceae (i). i Gambia. Rbipidium, a monoch. cyme where each successive branch falls alt. on one side and the other of the relatively main axis, and all fls. are in one plane, Iris, Juncus. Rhipogonum Forst. Liliaceae (xi). 5 E. Austr., N.Z. Rhipsalis Gaertn. (incl. Hariota, Pfeijfera). Cactaceae (ill. 3). 50 Brazil, Argentina. R. Cassytha L. is found in Ceylon and Madag., but quite possibly introduced (the only cactus out of Am. ; see fam.). Epiph., rarely thorny, sometimes of Cereus-like structure, sometimes Phyllocactus-like, or with cylindrical stems (compare sp. of Euphorbia). Fr. fleshy. Rhiz- rMzo- (Gr. pref.), root; -carp, pi. with sporangia on root-like processes, Marsileaceae\ -me, see separate article; -opliore, Selaginella. Rhizanthemum VanTiegh. (Loranf/ntsp.p.). Loranth. (i). 2 Malaya. Rhizanthous. flowering from the root. Rrjizofooleae = Caryocaraceae. Rhizocephalum Wedd. Campanulaceae (ill). 4 Andes. Rliizoma galangae, Alpinia officinarum Hance. Rhizome, an underground creeping stem, usu. thickened, and mono- or sympodial, Acorus, Alismaceae, Anemone, Asparagus, Balanophoraceae, Bambusit, Begonia, Bellis, Cyperaceae,Dioscorea,Equisetum, Eranthis, Gmmhieae, Iridaceae, Juncaceae, Lamiitm, Liliaceae, Mnsa, Ophio- glossaceae, Petasites, Fteris, Ranunculaceae, Typha, Zingiber. R HO DOR A 565 Rhlzophora L. Rhizophoraceae. 3, R. Mangle L. in Am., K. inuc- ronata Lam. and another, Japan to E. Afr. These mangroves (?•£'.) are moderate-sized trees with a great development of roots from the stem and branches. On the sub-aerial parts of the roots are large lenticels, probably serving in the same way as the aerenchyma of Bruguiera, &c. The seed germinates upon the tree, the hypocotyl projects at the micropyle and grows rapidly. The bark is used for tanning, yielding a substance known as cutch (cf. Acacia). Rnizophoraceae (EP., BH.}. Dicots. (Archichl. Myrtiflorae ; Myrtales BH.}. 12 gen., 60 sp., trop., mostly Old World. See Mangroves. Trees usu. with opp. slip. 1. ; fls. sol. or in cymes, &c. , 5 , hypo- to epi-gynous, reg. K usu. 4 — 8 ; C 4 — 8 ; A 8 — oo , inserted on outer edge of perig. or epig. disc; G (2 — 5), i — 5-loc. with usu. 2 anatr. pend. ov. in each loc. Fr. a slightly soft berry. Chief genera: Rhizophora, Bruguiera, Weiht-a. Rhodamnia Jack. Myrtaceae (i). 12 Malay Penins. to N.S. Wales. Rhodanthe Lindl. = Helipterum DC. p.p. (Comp.). Rhodax Spach = Heliantherrium Tourn. (Cist.). Rhodea Endl. = Rohdea Roth. (Lili.). Rhodes grass, Chloris. Rhodiola L. = Sedum Tourn. p.p. (Crass.). Rhodo- (Gr. pref.), rose red. Rhodocalyx Muell.-Arg. Apocynaceae (n. i). i campos of Minas Geraes. K coloured. Rhodochiton Zucc. Scrophulariaceae (n. 3). i Mexico, R. vohibile Zucc. , a twiner with sensitive petioles (cf. Clematis). Cult. orn. Rhodochlaena Thou. Chlaenaceae. 3 Madag. Rhodoclada Baker = Asteiopeia Thou. (Theaceae). E Madag. Rhodocodon Baker. Liliaceae (v). i Madag. Rhodocolea Baill. Bignoniaceae (4). i Madag. Rhododendron L. (incl. Azalea L.). Ericaceae (i. 2). 250 N. temp., &c. ''One sp. (R. Lochae F. Muell.) is found in trop. Austr. , the greatest richness of sp. is in E. As., from S. China to the Himal. and Japan; a second and lesser abundance is found in temp. N. Am., and a few sp. in the arctic regions. 4 sp. in Mid. and S. Eur. , 5 in Caucasus." (Drude.) Many sp. and hybrids cult. Shrubs and small trees with leathery 1. ; the 1. of § Azalea last one year, those of the other subgenera usu. more. Large winter buds are formed covered with scale 1. ; the larger and stouter ones contain infls., the slender ones merely 1. The branch bearing an infl is continued by the formation of a bud in one of the upper axils. Some of the Indian sp. are epiph. The C is slightly -|-, and the sta. and styles bend up- wards to touch the under surface of a visiting insect. R. ferrugineum L. (alpine rose) is protandr. and visited by humble-bees. Rhodogeron Griseb. Compositae (4). i Cuba. Rhodohypoxis Nel. Amaryllidaceae (in). 2 S.E. Afr. Rhodolaena Thou. Chlaenaceae. 3 Madag. Rhodoleia Champ, ex Hook. Hamamelidaceae. 2 Hongkong to Java. Rhodomyrtus Reichb. Myrt. (i). 6 trop. and E. As., Austr. Ed. fr. Rhodopis Urb. (Rudolphia p.p. ). Leguminosae (in. 10). r W.I. Rhodora L. = Rhododendron L. p.p. (Eric.). 566 RHODOSCIADIUM Rhodosciadium S. Wats. Umbelliferae (in. 6). 5 Mexico. Rhodosepala Baker. Melastomaceae (i). 3 Madag. Rhodospatha Poepp. et Endl. Araceae (n). n S. Am. Rhodospbaera Engl. Anacardiaceae (3). i E. Austr. Rhodostachys Phil. Hromeliaceae (4). 7 S. Am. Cult. orn. infl. Rhodothamnus Reiclib. Ericaceae (i. 3). i E. Alps, E. Siberia. Rhodotypos Sieb. et Zucc. Rosaceae (in. i). i Japan, R. kerrioides S. et Z., a cult. orn. fig. shrub. It has opp. 1., found in no other plant of the fam., except in seedlings of Primus. There is an epicalyx (see Potentilla). Rhoeadales. The igth order of Archichlamydeae. Rhoeadinae (Warming). The 9th order of Choripetalae. Rhoeidium Greene (Khus p.p. ). Anacardiaceae (3). 7 Mexico, Texas. Rhoeo Hance. Coinmelinaceae. i C. Am., Mexico. Cult. orn. fl. and 1. Rhoicissus Planch. ( Vitis p.p.). Vitaceae. 12 trop. and S. Afr. Rhoium (Cl.), a creek formation. Rhombochlamys Lindau. Acanthaceae (iv. B). 2 Colombia. Rhomboidal, of rhomboid shape. Rhombonema Schlechter. Asclepiadaceae (n. i). i S. Afr. Rhopalandria Stapf. Menispermaceae. i W. trop. Afr. Rhopaloblaste Scheff. Palmae (iv. i). 2 Moluccas, New Guinea. Rhopalobrachium Schlechter et Krause. Rubiaceae (n. i). 2NewCal. Rhopalocarpus Boj. Flacourtiaceae (inc. sed.). 3 Madag. Rhopalocnemis Jungh. Balanophoraceae. i E. Indomal. Rhopalopilia Pierre. Opiliaceae. 3 W. trop. Afr. Rhopalostylis H. Wendl. et Drude (Kentia p.p. EP.). Palmae (iv i) 2 Norfolk I., New Zealand. Rhubarb, Rheum officinaleEzM.; -(vegetable) R. Rhaponticum L. Rhus (Tourn.) L. (incl. Cotinus Tourn. and Toxicodendron Tourn.). Anacardiaceae (3). 130 subtrop. and warm temp. R. Coriaria L. is the sumac (S. Eur.); its 1., ground fine, are used for tanning and dyeing. R. Toxicodendron L. (N. Am., poison-ivy) climbs like ivy. Its juice produces ulcerations or erysipelas. R. Cotinus L. (Medit. to China) is the wig-tree, often cult, in shrubberies. Fls. polyg. The stalk of each drupe remains smooth, but the sterile parts of the panicle lengthen and become hairy. Then when ripe the stalks become detached at their joints, and the whole infl., with the fr. on it, falls to the ground and may be blown about. The wood yields the yellow dye 'young fustic.' R. verniciftra DC. is the lacquer-tree. Japan lacquer is obtained from notches in the stem. A', sticcedanea L. is the wax-tree of Japan ; its crushed berries yield wax. Rhynchanthera DC. Melastomaceae (i). 35 trop. Am. Rhynchanthus Hook. f. Zingiberaceae (i).' "i Burma. Rhynchelytrum Hochst. (Tricholaena EP.). Gramineae (5). i trop. Afr. Rhynchocalyx Oliv. Lythraceae. i Natal. Rhynchocarpa Schrad. = Kedrostis Medic. (Cucurb.). Rbynchocorys Griseb. Scrophulariaceae (in. 3). 28. Eur. to Persia. Rbynchodia Benlh. Apocynaceae (n. i). 2 E. Indomal. Rbynchodium C. Presl= Psoralea L. (Legum.). RICH ELL A 567 Rhynchoglossum Blume. Gesneriaceae fi). 2 Malay Archipelago. Rhyncholacis Tul. Podostemaceae. 7 Guiana, Brazil. Rhynchopera B6rner = Caiex L p.p. (Cyper.). Rhynchopyle Engl. Araceae (v). 4 Borneo. Rhynchoryza Baill. (Oryza p.p. £/>.). Gramineae (6). i Brazil. Rhynchosia Lour. Leguminosae (in. 10). 120 trop. and subtrop. Rhynchospermum Reinw. Compositae (3). i Himalaya to Japan. Rhynchospora Vahl. Cyperaceae (n). 150 cosmop., esp. trop. Rhynchosporous (fr.), ending in a beak. Rhynchostigma Benth. (Toxocarpus BH.}. Asclepiadaceae (n. 3). 3 trop. VV. Afr. Rhynchostylis Blume. Orchidaceae (n. 20). 2 Inclomal. Cult. orn. fl. Rhynchotechum Blume. Gesneriaceae (i). 9 Indomal. Rhynchotheca Ruiz et Pav. Geraniaceae. 2 Peru. Rhyncho ;ropis Harms (Indigofera p.p.)- Leguminosae (ill. 6). 2 W. Afr. Rhysopterus Coulter et Rose (Cymopterus p.p.). Umbelliferae (in. 5). 3 N. Am. Rnysotoechia Radlk. Sapindaceae (i). 7 Austr., Malay Archip. Rhyssocarpus Endl. (Melanopsidinm EP.}. Rubiaceae (i. 8). i trop. Am. Rhyssolobium. E. Mey. Asclepiadaceae (11. 3). i S. Afr. Rhyssopteryx Blume. Malpighiaceae (i). 7 Malaya, N. Austr. Rhyssostelma Decne. Asclepiadaceae (11. i). i temp. S. Am. Rhytachne Desv. Gramineae (2). 5 trop. Afr. Rhyticarpus Sond. Umbelliferae (in. 5). 3 S. Afr. Rhyticaryum Becc. Icacinaceae. 3 New Guinea. Rhyticocos Becc. (Cocos p.p.). Palmae (iv. 2). i \V.I. Rhytidantriera Van Tiegh. Ochnaceae. 3 Colombia. Rhytidophylluin Mart. Gesneriaceae (n). 12 W.I. Rhytiglossa Nees = Dianthera Gronov. (^5/7^.)=Justicia p.p. Rhytispennum Link= Lithospermum L. (Borag.). Rib grass, -wort plantain, Piantago. Ribbon grass, Pkalaris arundinacea L. var. variegata. Ribeiria Arruda. Rosaceae (inc. sed.). Nomen. Ribes L. Saxifragaceae (vi). 60 N. temp, and Andine; 4 in Brit. Shrubs, often with spines (emergences), and with racemes of fls. on 'short shoots.' G with two parietal plac. Fls. usu. homogamous, with self-pollination in default of insect-visits. R. alpinuin L. is dioec. In R. sanguineum Pursh (cult. orn. shrub, flowering currant) the petals change from white to pink as the fls. grow older, and in J?. aiireum Pursh from yellow to carmine (see Fumaria, Boraginaceae). R. rubrum L. is the red, R. nignun L. the black currant, R. Grossu- laria L. the gooseberry, all cult, for fr. Rice, Oryza saliva L. ; Canada, Indian, Tuscarora-, Zizania aquatica L. ; -paper, Tetrapanax papyri/era C. Koch. Ricnardia Houst. ex L. = Richardsonia Kunth. Richardia Kunth = Zantedeschia Spreng. (Arac.). Richardsonia Kunth. Rubiaceae (II. 10). 9 warm Am. Richea R. Br. Epacridaceae (2). 8 Tasmania, Victoria. Richella A. Gray. Anonaceae (2). i Fiji. 568 RICHER! A Richeria Vahl. Euphorbiaceae (A. I. i). 3 Brazil, Peru. Richthofenia Hosseus. Rafflesiaceae. i Siam. Richweed (Am ), Pilea. Ricinocarpos Uesf. E "phorbiaceae (B. n). 13 Austr. Ricinodendron Muell.-Arg. Euphorbiaceae (A. n. 5). 2 trop. W. Afr. Ricinus (Tourn.) L. Euphorbiaceae (A. II. 2). r trop. Afr., R. com- nnmis L. (castor-oil), a shrub in trop., a herb in Eur. Monoec. The 5, often only united by the style, and ± divided when ripe; loculic. de- hiscence usu. with separation of the endocarp ; rarely 4 — i fleshy drupes. i. Zanthoxyleae (woody plants, usu. with small greenish, reg., often unisexual fls. ; cpl. rarely with > 2 ovules ; embryo with flat cots, in endosp.) : Zanthoxylum, Fagara, Choisya. i. Riiteae (herbs or undershrubs, rarely shrubs, with moderate sized § fls., sometimes slightly -I- ; cpls. usu. with > 2 ov. ; endosp.): Ruta, Dictamnus. 3. Boronieae (undershrubs and shrubs, with reg. usu. § fls.; endosp. fleshy, otherwise as 2) : Boronia, Eriostemon, Correa. 4. Diosmeae (undershrubs and shrubs, rarely trees with simple 1. ; exalbum., embryo usu. straight with fleshy cots.) : Calodendron, Adenandra, Uiosma. 5. Cusparieae (shrubs and trees with reg. or •]• fls. ; endosp. little or o; embryo curved, with radicle between the cots.) : Almeidea, Galipea, Cusparia. 6. Dictyolomeae (fls. reg., haplostemonous ; sta. with scales at base; cpls. with oo ovules, only united at the base; trees with double pinnate 1.) : Dictyoloma. II. FLINDERSIOIDEAE. Cpls. (5—3), each with 2—8 2-ranked erect ovules ; capsule loculic. or septic, with persistent endo- carp; seed winged, exalbum. ; woody plants with lysigenous glands. Flindersieae: Flindersia, Chloroxylon. III. SPATHELIO1DEAE. Cpls. (3) each with 2 pend. ov.; drupe winged ; secretory cells and lysigenous oil-glands at margins of 1. Spathdieae: Spathelia. IV. TODDALIOIDEAE. Cpls. (5—2) or i, each with 2—1 ov. ; drupe or dry winged fr. ; endosp. or o; 1. and bark with lysi- genous oil-glands. Toddalieae: Ptelea, Toddalia, Skimmia. V. AURANT10IDEAE. Berry, often with periderm, and SACCARDOPHYTUM 579 with pulp derived from sappy emergences of cpl. wall. Seeds exal- bum., often with 2 or more embryos. Lysigenous oil-glands. Aurantieae: Glycosmis, Limonia, Atalantia, Feronia, Aegle, Citrus. Ruthea Bolle (Lichtensleinia BH.). Umbelliferae (in. 5). 4 Afr. Rutidea DC. Rubiaceae (n. 4). 15 trop. Afr., Madag. Rutidosis DC. Compositae (4). 6 Austr. Rutilans (Lat.), with glowing fls. Rutilia Veil. Inc. sed. i Brazil. Ruttya Harv. Acanthaceae (iv. B). 3 S. Afr. Ruyschia Jacq. Marcgraviaceae. 3 trop. Am. Ryania Vahl (Patrisia Rich.). Flacourtiaceae (7). 10 N. trop. S. Am. Rydbergia Greene (Actinella p.p.)- Compositae (6). 2 N. Am. Rye, Secale cereale L. ; -grass, Lolium perenne L.; French--, Arrhe- natherum. Rylstonea R. T. Baker. Myrtaceae (inc. sed.). i Austr. Ryncospora Vahl. Cyperaceae (n). 175 temp, and trop. ; 2 in Brit, in wet bogs (beak rush). Ryparosa Bl. Flacourtiaceae (3). loS.E.As. Ryssopterys Blume ex A. Juss. Malpighiaceae (i). 7 E. Indomal. Ryssosciadium O. Ktze. = Rhysopterus Coulter et Rose (Umbell.). Ryticaryum Becc. Icacinaceae. 4 New Guinea. Rytidocarpus Coss. Cruciferae. i Morocco. Rytidptus Hook. f. (Bobca p.p. EP.}. Rubiaceae (n. 2). i Ha- waiian Is. Sabadilla Brandb. et Ratzebg. (Schoenocaulon A. Gray BH.}. Lilia- ceae (i). 5 Georgia to Venezuela. SabalAdans. Palmae (i. 2). 10 warmer Am., W.I. S. Palmetto Lodd., &c. (palmetto or thatch palm), 1. are used for thatching; the wood is also useful. Sabaudia Buscalioni et Muschler. Labiatae (vn). i C. Afr. Sabazia Cass. Compositae (5). 2 Mexico. Sabbata Veil. Compositae (inc. sed.). 2 Brazil. Sabbatia Adans. Gentianaceae (i). 12 N. Am. (rose pinks). Sabia Colebr. Sabiaceae. 20 S. and E. As. Sabiaceae (EP., BH.}. Dicots. (Archichl. Sapindales). 4 gen., 65 sp., trop. and E. As. Trees, shrubs or lianes with alt. exstip. impari- pinnate or simple 1. Infl. a panicle or cymose panicle, with bracts and bracteoles. Us. usu. $. K (3—5), imbr. or free; C 4—5, sometimes united at base, imbr., the inner 2 much reduced ; A 5, opp. petals, all or only i fertile, the rest being staminodial; G (2) 2-loc. ; in each loc. usu. i axile pend. or horiz. semi-anatr. ov. with micro- pyle upwards. Indeh. fr. with exalbum. seeds. Chief genera: Sabia, Meliosma. Sabicea Aubl. Rubiaceae (i. 7). 30 trop. Am., Afr., Madag. Sabiiia Hall = Juniperus Tourn. p.p. (Conif.). Sabinea DC. Leguminosae (in. 6). 3 Panama, W.I. Sabre bean, Canavalia ensifori/iis DC. Sabulina Reichb. = Arenaria Rupp. (BH.}. = Alsine Scop. p.p. Sac, a pouch ; saccate, with a pouch. Saccardophytum Spegazzini. Solanaceae (inc. sed.). i Patagonia. 37—2 580 SACCELLIUM Saccellium Humb. et Bonpl. Boraginaceae (u). i Peru, Argentina. Saccharodendron Nieuwland (Acer p.p.). Aceraceae. 5 N. Am. Sacchamm L. Gramineae (2). 12 trop., subtrop., incl. S. officinantm L. (sugar cane), a native (?) of trop. E. As., now cult, in most warm regions, esp. Java, Hawaii, U.S., &c. From the rhiz. there spring each year shoots which may reach 12 — 15 feet and a thickness of 2 inches; the outer tissues have much silica in their cell-walls. The infl. is a dense woolly spike, the first and second glumes of each spikelet being covered with long hairs. The cult, form has always been veg. pro- pagated (pieces of the haulm, each bearing a bud, are planted), but recently a more vigorous race has been raised from seed. The sugar is contained in the soft central tissues of the stem ; the canes are cut before flowering and crushed between rollers to extract the juice; afterwards it is boiled down under reduced pressure and laid out to crystallise. Saccocalyx Coss. et Dur. (Satureia p.p. BH. ). Labiat. (vi). i Alger. Saccoglossum Schlechter. Orchidaceae (n. 16). 2 New Guinea. Saccoglottis Endl. Humiriaceae. 10 trop. S. Am., Afr. Saccolabium Blume. Orchidaceae (n. 20). 50 Indomal. Cult. orn. fl. Saccolaria Kuhlmann. Lentibulariaceae. i Upper Amazon. Saccolepis Nash (Panicitm p.p.). Gramineae. 5 N. Am. Saccoloma Kaulf. Polypodiaceae. 8 trop. Am., E. As. Saccopetalum Bennett. Anonaceae (2). 5 trop. As., Austr. Saccostoma Wall. Labiatae (inc. sed.). i Indomal. Sachsia Griseb. Compositae (4). 3 Cuba. Sack tree (Ceylon), Antiaris toxicaria Leschen. Sacleuxia Baill. Asclepiadaceae (n. i). i Zanzibar. Sacoglottis Mart. - Saccoglottis Endl. (Humir.). Sacred bean or lotus, Ndnmbium speciositm Willd. Sacrosphendatnus Nieuwland (Acer p.p.). Aceraceae. i N. Am. Sadiria Mez (Pimelandra p.p.). Myrsinaceae (n). 4 Indomal. Sadleria Kaulf. Polypodiaceae. 4 Hawaiian Is. Safflower, Carthanius tinctorius L. Saffordia Maxon. Polypodiaceae. i Peru. Saffordiella Merrill (Leptospermum p.p.). Myrt. (n. i). i Phil. Is. Saffranhout, Elaeodendron crocenm DC. Saffron, Crocus sativus L. ; meadow-, Colchicum autumnale L. Sage, Salvia, esp. S. officinalis L. ; -brush, Artemisia tridentataNutt., &c. ; -rose (W.I.), Turnera; wood-, Tencriiim. Sageraea Dalzell (Bocagea BH.}. Anonaceae (i). 6 Indomal. Sageretia Brongn. Rhamnaceae. 12 warm E. As , N. Am. Sagina L. Caryophyllaceae (I. [). 20 N. temp.; 4 in Brit, (pearl- wort). Small herbs with inconspic., sometimes apet. fls. Sagittaria Rupp. ex L. Alismaceae. 33 Am., temp. |-x-, incl. S. sagit- tifolia L. (arrow-head) in Eur. (incl. Brit.), a water-plant with a short rhiz. bearing 1. of various types, the number of each kind depending on the depth of the water, &c. The fully submerged 1. are ribbon- shaped, the floating ones have an ovate blade, whilst those (usually the majority) that project above water are arrow-shaped (sagittate). In the axils are formed the ' renewal' shoots which last over the winter, short branches which burrow into the mud and swell up at the ends SALICORNIA 581 each into a large bud whose central axis is swollen with reserve- materials; in spring this develops into a new plant. The diclinous racemose infl. projects above water; the S fls. are lower down than the itans Engl. (Braz.), good timber. Sails, cf. Bamboos. Sainfoin, Oiiobrychis sativa Lam. St Dabeoc's heath, Daboeda ; - Ignatius' beans, Strychnos Ignatii Berg.; - John's oread, Ceratonia; -- wort, Hypericuin. Saintpaulia H. Wendl. Gesneriaceae (i). 3 E. Afr., incl. S. ionantha H. Wendl. (Afr. violet). The fl. is like that of Exacum, with similar dimorphic symmetry. In some the style projects to the left over the C, in others to the right (cf. Exacum, Cassia). Sakersia Hook. f. Melastomaceae (i). i Cameroons. Sal, Shorea robusta Gaertn. f. Salacia L. Hippocrateaceae. 100 trop., often lianes with dimorphic branches, one form suited for climbing. Salacicrater Loes. Hippocrateaceae. t New Guinea. Salacistis Reichb. f. (Hetaeria BH.). Orchidaceae (n. 2). i Java. Salad oil, nominally olive oil, Olea europea L. Salaxis Salisb. Ericaceae (iv. 2). 24 Cape Colony. Salazaria Torr. Labiatae (iv). i S.W. U.S., Mexico. Saldanhaea Bur. Bignoniaceae (i). 2 Brazil. Saldania Sim. Leguminosae (in. 8). i S.E. Afr. Saldinia A. Rich. Rubiaceae (n. 5). 2 Madag. Salep, the dried tubers of some sp. of Orchis and (in India) Eulo- phia. Salicaceae (EP., BH.). Dicots. (Archichl. Salicales; anom. Incom- pletae BH.). 2 gen., 180 sp., N. temp., trop. and subtrop. Shrubs or trees with stip. 1. and much veg. repr. by suckers. Fls. naked, in catkins or spikes, dioec. (many hybrids exist). The catkins arise in autumn and remain as buds through the winter, developing in early spring. The i fl. consists of 2 — 30 sta. in the axil of a bract, the ? usu. of (2) cpls. transv. placed, syncarpous with parietal plac. ; ovules oo , anatr. Seeds exalbum. with basal tufts of hairs. Genera: Salix, Populus. Salicales. The 3rd order of Archichlamydeae. Salicaria Tourn. ex Mill. = Lythrum L. (Lythr.). Saliciflorae (Warming). The ist order of Choripetalae. Salicornia (Tourn.) L. Chenopodiaceae (A). 10 on sea-coasts. S. herbacea L. (saltwort) cosmop., incl. Brit. Succulent herbs, with the habit of a cactus, leafless and with jointed nodes. Fls. in groups of 3 582 SALICORNIA or more, one group sunk in the tissue on either side of each internode. P fleshy; sta. i or 2. Salisburia Sm. = Ginkgo L. (Ginkg.). Salix (Tourn.) L. Salicaceae. 160 cosmop. The sp. are difficult to separate and there are many hybrids (cf. Ruhus, Rosa), 12 in Brit, (willow, sallow, &c.). Branching monopodial, but the term, bud usu. dies, and the next lat. bud continues the axis. Some have collateral buds in the axils. There is extensive veg. repr. by suckers. Some, e.g. S. alba L., are often pollarded, or cut off at a height of 8 feet or so; from the callus formed upon the wounds new shoots spring, and thus the 'crown' of shoots is produced. Among the Brit. sp. are S. herbacea L., the dwarf or arctic willow, a creeping alpine and arctic form, and S. Janata L., &c. alpine forms with very woolly 1. The fls. contain honey, and as they appear in early spring, before the L, and when they have but few competitors, they receive a great many visits from insects, especially from bees. S. viminalis L. is the osier, whose twigs are used in making baskets, &c. S. babylonica L. is the weeping willow. S. caprea L. useful wood. Sallal, Gaultheria Shallon Pursh. Sallow, Salix. Salmea DC. Compositae (5). 12 Mexico, W.I. Salmeopsis Benth. Compositae (5). i S. Brazil, Paraguay. Salmia Willd. = Carludovica Ruiz et Pav. (Cyclanth.). Salomonia Heist. (Polygonatum p.p.). Liliaceae (vn). 2 N. Am. Salomonia Lour. Polygalaceae. 5 E. As. Some are parasitic. Salpichroa Miers. Solanaceae (2). 12 warm Am. Salpiglossis Ruiz et Pav. Solanaceae (5). 8 S. Am. Cult. orn. fl. Salpinga Mart. Melastomaceae (i). 3 S. Brazil. Cult. orn. fl. Salpingacantlius Sp. Moore (Ruellia p.p. EP.). Acanth. (iv. A), i Brazil. Salpingia Torr. et Gray (Oenothera p.p.). Onagr. (2). 4 Mexico, Texas. Salpinxantha Hook. f. (Geissomeria BH.}. Acanthaceae (iv. B). i Jamaica. Salsify, Tragopogon porrifolius L. Salsola L. Chenopodiaceae (B). 40 cosmop., maritime or on salt steppes. S. Kali L. (glass-wort) in Brit., a very fleshy plant with 1. ending in spines. A var. tragus Moq. of this sp. (Russian thistle) has in recent years become a pest of agriculture in N. Am. Salsuginosus (Lat.), growing where inundated by salt water. Salt-bush, A triplex, &c. ; -wort, Salsola; -steppe plants (halophytes), Chenopodiaceae, Frankeniaceae, &c. Saltia R. Br. Amarantaceae (2). i S. Arabia. Salvador tea, Gaultheria. Salvadora Garcin. ex L. Salvadoraceae. 2 W. As., Afr. S. persica L. is said to be the mustard of the Bible. Its 1. taste like mustard. Salvadoraceae (EP., BH.). Dicots. (Sympet. Contortae ; Gentianales BH.}. 3 gen., 6 sp., As., Afr. Shrubs and trees with opp. entire slip. 1. and racemose infls. Fls. $ or unisexual, reg. K (2 — 4); C (4 — 5) or 4 — 5, with teeth or glands on the inner side; A 4 — 5, epipet. or not; G (2), i — 2-loc. with 1—2 erect anatr. ov. in each. SAL VINIA CEAE 583 Fr. a i-seeded berry or drupe. Seed exalbum. Genera: Azima, Dobera, Salvadora. The relationships are doubtful, for we do not know if the polypetaly of A. and D. is original or secondary. If the former, the order must perhaps be placed near Celastraceae. Salver-shaped (C),±flat, with long tubular portion, Primula. Salvertia A. St Hil. Vochysiaceae. i campos of S. Brazil. Salvia (Tourn.) L. Labiatae (vi). 550 trop. and temp. 6". Verbe- naca L. (sage) and S.pratensis L. (clary) in Brit. The sta. are reduced to 2 (the ant.), each of which has a sort of T-shape, the connective of the versatile anther being greatly elongated. The stalks of the sta. stand up together across the mouth of the fl., and a bee, in pushing down towards the honey, comes into contact with the inner end of the anther, and raising it causes the outer to descend upon its back and to rub it with pollen. In some forms of S. both ends of the lever bear fertile anthers; but in most the useless half-anther at the inner end is aborted, and the outer half of the connective is much longer than the inner (compare S. officinalis with S. pratensis}. The fl. is protandrous, and in the later stage the style bends down and places the stigma in position to be touched first by an entering insect. Some have coloured bracts at the top of the infl., adding to its conspicuous- ness. .9. officinalis L. (Medit.) is the garden sage. Salviacantlms Lindau. Acanthaceae (iv. B). i Cameroons. Salviastram Scheele. Labiatae (vi). 3 Texas, New Mexico. Salvinia (Mich.) Schreb. Salviniaceae. 10 trop. and warm temp., incl. S. natans (L.) All. The pi. floats freely on the water; at each node is a whorl of three L, and the whorls alt. with one another. There are two floating 1. derived from the upper half of a segment of the apical cell (see fam.), and a submerged 1. derived from the lower. There are no roots, their function being performed by the finely divided submerged 1. (cf. Trapa, Ranunculus, Cabomba). The sporocarps are borne several together as outgrowths from the base of a submerged 1. The microspores germinate inside the sporangium, the prothalli emerging through its wall as fine tubes, at the end of which the antheridia form. Salviniaceae. Filicales Leptosporangiatae. Two genera, Salvinia (1. in whorls of 3) and Azolla (1. in two ranks), with 15 sp., trop. and temp. Water plants, with a stem floating upon the water, and growing by a two-sided apical cell (3-sided in the young embryo, as in other Filicales). A dorsiventral construction thus arises; segments are cut off right and left from the apical cell, and the first division of each of these segments divides it into a dorsal and a ventral half. In S. the dorsal halves give rise to the floating, the ventral to the submerged 1. ; in A. the former give rise to the 1., the latter to the branches and roots. The sporangia are grouped into sori; the sorus is enclosed in a highly developed indusium, forming a sporocarp. Each contains only one kind of sporangium (micro- or mega-sporangia). The sporocarp is an outgrowth of a 1., — in S. of a submerged L, in A. of the ventral lobe of an ordinary 1. The spore is covered with an epispore, consisting of hardened frothy mucilage. It sinks, when set free from the sporangium. On germ, the microspore forms a rudimentary <$ prothallus consisting of one (Pmore) veg. cell and an antheridium. The megaspore forms 584 SALVINIACEAE a ? prothallus, which remains enclosed in the burst spore, and has two parts, an upper small-celled green part on which are borne the arche- gonia, and a lower colourless part (of one or more large cells), in which reserves are stored up for the use of the young plant which will be formed from a fert. ovum (cf. Selaginella and Phanerogams). Salzmannia DC. Rubiaceae (n. 3). i E. Brazil. Samadera Gaertn. Simarubaceae. 7 Madag. , Indomal. Samaii. Pithecolobinni Sam an Benth. Samara L. = Embelia Burm. p.p. (Myrsin.). Samara, a winged i -seeded indeh. achene or nut, Acer, Banisteria, fraxinns, Liriodcndron, Ptelea, Pterocarpus, L/hntts, Ventilago. Sambucus (Tourn.) L. Caprifoliaceae. 20 N. temp., S. Am., As. to Austr. S. nigra L. (elder) and S. Ebuhts L. in Brit. Differs from the rest of the fam. in having cpd. 1. and extr. anthers; it also possesses well-marked stipules. Lenticels show clearly in the bark. S. nigra L. shows serial adv. buds. There has been some discussion about the proper position to be assigned to it; it has been put in Valerianaceae or in a new family, Sanibucaceae, forming a link between Rubiales and Aggregatae. A wine is prepared from elder berries. Sameraria Desv. (hatis BH.}. Cruciferae (2). 7 E. Medit. Samoa millet, Panicum. Samolus (Tourn.) L. Primulaceae. 10 cosmop., esp. ^; S. Valerandi L. (brook-weed) in Brit. The whole pi. dies down in autumn, but young shoots form in summer and take root. The bracts of the fls. are 'adnate' to the axes, so as to look like sol. bracteoles (cf. Sola- naceae). Samphire, Crithmum maritimum L., (Am.) Salicornia. Samuela Trelease. Liliaceae (vi). 2 Mexico, California. Samyda L. F'lacourtiaceae (7). 4 W.I., Mexico. Samydaceae (BH.}. A fam. in Passiflorales, comprising the gen. Samyda, Casearia, Banara, Abatia, Homalium, &c. ; placed in Fla- courtiaceae by Engler. Sancliezia Ruiz et Pav. Acanthaceae (iv. A). 10 trop. S. Am. San Domingo apricot, Mammea ainericana L. Sand-binding1 plants, Acacia, Ammophila, Carex, Distichlis, Elymus, Larrea, Lepidosperma, Stenotaphriun, &c. ; - bur, Cenclirus tribuloides L. ; -box tree, Hitra; -dunes, flora of, cf. Tansley, Types of Brit. Veg.; -spurrey (Am.), Spergidaria; -wort, Arenaria. Sandalwood, Santaium album L. ; red -, Pterocarpus sanlalinus L. Sandarach, Tetraclinis quadrivalvis Vent. Sandbergia Greene (Arabis p.p.). Cruciferae (4). i N.W. N. Am. Sanderella O. Ktze. = Parlatorea Barb. Rodr. (Orch.). Sanders wood, Sandalwood. Sandersonia Hook. Liliaceae (i). i Natal. Sandoricum Cav. Meliaceae (in). 6 Mauritius, Indomal. Some have ed. fr. Sanguinaria Dill, ex L. Papaveraceae (n). i All. N. Am., S. catia- densis L. , the blood-root. It has a thick rhiz. giving off annually one 1. and a i-flowered scape. The rhiz. is used in medicine. Sanguineus (Lat.), blood-coloured. Sanguisorba Rupp. ex L. (Poteriiim BH.}. 30 N. temp. 2 Brit., SAPINDA CEAE 585 S. minor Scop, and 6". officinalis L., anemophilous, the former with long pend. sta. Sanhilaria Baill. Bignoniaceae (i). r S. Brazil. Sanicle, Sanicula europea L. Sanicula (Tourn.) L. Umbelliferae (n. i). 20 cosmop. exc. Austr. 6". enropaea L. (sanicle) in Brit. Fls. in cymose umbels, themselves arranged in dichasia. Fr. hooked, animal- distributed. Sansevleria Thunb. Liliaceae (vi). 12 trop. Afr., As. Xero. with fleshy 1. S. zeylanica Willd. yields fibre (bow-string hemp). Santalaceae (EP.; BH. incl. Grubbiaceae, Myzodendraceae). Dicots. (Archichl. Santalales; Achlamydosporeae BH.}. 26 gen., 250 sp., of semi-parasitic shrubs, trees and herbs, resembling Loranthaceae in many ways, trop. and temp. Some are stem-parasites like mistletoe, others root-parasites like Rhinanthus (e.g. Thesium). L. opp. The total infl. may be a raceme, spike, head, &c., but often, instead of the single fl. in each axil, there is a little cyme of 3, as in Loranthaceae. Fls. $ or unisexual, with perig. or epig. disc and a simple P (sepaloid or petaloid); sta. =, and inserted on, the P-leaves. G i-loc., with a central placenta bearing i — 3 ovules. Nut or drupe; seed i, with no testa, and much endosp. Chief genera: Santalum, Thesium. Santalales. The 1 4th order of Archichlamydeae. Santalina Baill. Rubiaceae (i. 8). i Madag. Santalodes O. Ktze., Santaloides Schellenb. = Rourea Aubl. p.p. Santalum L. Santalaceae. 10 Indomal. Parasitic trees. S. album L. (S. Ind., esp. Mysore) furnishes the true sandal-wood (yellow or white). Oil is distilled from it. Santiria Blume. Burseraceae. ^,30 Malaya. Santiridium Pierre = Pachylobus G. Don p.p. (Burser.). Santiriopsis Engl. (Santiria p.p.). Burseraceae. i S. Thome. Santolina Tourn. ex L. Compositae (7). 8 S.W. Eur. 6". Chamae- cyparissus L. is officinal. Sanvitalia Gualt. in Lam. Compositae (5). 8 warm Am. Cult. orn. fl. Sap-green, Rhamnus cathartica L. ; -wood, the young outer wood. Sapidus (Lat.), with pleasant taste. Sapindaceae (EP.~, BH. incl. Aceraceae, Hippocastanaceae, Meliantha- ceae, Staphvleaceae}. Dicots. (Archichl. Sapindales). 120 gen., 1000 sp., trop. and subtrop. 5 gen. (Serjania, Paullinia, &c.) with 300 sp. are lianes, the rest erect trees or shrubs. The liane.s climb by tendrils, which are met. infl. -axes and are usu. branched or sometimes watch- spring-like; their stems often show peculiar internal anatomy. L. alt., slip, in the climbing sp., usu. cpcl., pinnate; in the climbing sp. there is usu. a true term, leaflet, but not in the erect; in these one of the last pair of leaflets often becomes term., so that the 1. is asymmetric. The tissues of the plants usually contain resinous or latex-like secretions in special cells. The infl. is cymose, usu. a cincinnus, with bracts and bracteoles. Fl. unisexual (the sta. are apparently well developed in the ? so that it is easily mistaken for 5, but the pollen is useless, and the anthers do not open), generally monoec., reg. or often obliquely -|- , 5- or 4- merous. K usu. •;, rarely (5), imbr. or rarely valvate or open, some- times apparently 4-merous by union of 2 sepals; C usu. 5, imbr., with 5 86 SAPINDA CEAE well-marked disc between it and the sta. ; A usu. 5 + 5 in one whorl, often with i absent, more rarely 5, 4, or oo , inserted within or rarely upon the disc round the rudimentary ovary; G in ? fl. usu. (3), 3-loc. with term, style; ovules usu. i in each loc., ascending, with ventral raphe. Fr. a caps., nut, berry, drupe, schizocarp, or samara, usu. large, often red ; seed often arillate, with no endosp. ; embryo usu. curved. Many S. are of economic value ; several yield valuable timber ; Nephelium, Litchi, and others furnish ed. fr. Classification and chief genera (after Radlkofer) : I. EUSAPINDACEAE (ov. sol. in loc., erect or ascending, micropyle down) : Serjania, Paullinia, Sapindus, Talisia, Schleichera, Litchi, Nephelium, Pappea, Cupania, Blighia. II. D VSSAPINDACEAE (ov. usu. 2 or several in each loc., in the first case erect or pend., in the second horiz. , rarely i pend. with micropyle up) : Koelreuteria, Dodonaea. Sapindales. The 24th order (EP.) of Archichlamydeae. The loth (BH.) of Polypetalae. Sapindus Tourn. ex L. Sapindaceae (i). n trop. and subtrop. exc. Afr. and Austr. The berries of S. Saponaria L. (Am.) contain saponin, form a lather with water, and may be used as soap. Sapium P. Br. Euphorbiaceae (A. n. 7). 35 trop. Seeds of S. sebiferum Roxb. (tallow-tree, China) are coated with fat; they also yield an oil by pressure. Some yield rubber (Bolivian, Colombian). Sapodilla plum, Achras Sapota L. Saponaria L. Caryophyllaceae (n. 2). 20 N. temp., chiefly Medit. S. officinalis L. (soapvvort) in Brit. Its leaves lather if rubbed with water. Fls. protandrous, butterfly-visjted. Saponin, Cklorogalum, Gonania, Limoiiia, Sapindus, Saponaria. Sapota Plum, ex Mill. = Achras L. (Sapot.). Sapotaceae (EP., BH.). Dicots. (Sympet. Ebenales). 35 gen., 600 sp. , trop., mostly trees with entire leathery 1., sometimes slip. They are commonly hairy with 2-shanked hairs, and contain secretory passages in pith, cortex and 1. Fls. sol. or in cymose bunches in the 1. axils or on old stems, bracteolate, $ , reg. or not. K 2 + 2, 3 + 3, 4 + 4, or 5 ; C usu. equal in number to sepals, and alt. with the K as a whole, as in Cruciferae, rarely in 2 whorls. In Miniusopeae the petals have dorsal appendages like themselves, giving the appearance of more than one whorl. Sta. in 2 or 3 whorls, but frequently the outer staminodial or absent ; anthers commonly extr. G, syncarpous, multiloc. ; cpls. = or twice the number of sta. in a whorl, or more ; ovules at base of axile placenta, one in each loc., anatr. with micropyle facing down ; style simple. Beriy, the flesh sometimes sclerenchy- matous near the surface. Seeds few or one, usually album. ; endosp. oily ; testa hard and rich in tannin. Many S. furnish useful products, esp. gutta-percha and balata ; see all genera below. A fam. of conspic. economic value. Classification ana chief genera (after Engler): I. Palaqiiieae (petals without appendages): Bassia, Payena, Palaquium, Achras, Butyrospermum, Sideroxylon, Chryso- phyllum. II. Mimusofeae (petals with appendages — see above) : Mimusops. SARCOPHYTE 587 Sapote, Liicitma mammosa Gaertn. f. Sappan wood, Caesalpinia sappan L. Sapranthus Seem. (Porcdia BH., Uvaria p.p. EP.). Anon. (i). i C. Am. Sapria Griff. Rafflesiaceae. i Assam. Saprophytes, plants which grow upon decaying organic matter -and absorb the products of decay, Burmanniaceae, some Gentianaceae (Bartonia, Obolaria, &c.), Monotropa, some Orckidaceae (Corallorhiza, Epipogoti, Limodonim, Neottia, &c.), Trinridaceae, the prothallus of Ophioglossaceae. Saprosma Blume. Rubiaceae (n. 5). 12 Indomal. Sapu (Ceylon), Michdia Champaca L. Sapucaia nut, Lecythis. Saraca L. Leguminosae (n. 3). TO trop. As. Young shoots pend. (cf. Amherstia, Brownea). Fls. (scented at night) as temple of- ferings. Saracha Ruiz et Pav. Solanaceae (2). 12 Bolivia to Mexico. Saranthe Eichl. (Myrosma BH.). Marantaceae. 8 Brazil. Sararanga Hemsl. Pandanaceae. i Solomon Is. Sarcaiituidion Baill. Icacinaceae. i New Caledonia. Sarcanthus Lindl. Orchidaceae (n. 20). 30 Indomal., China. Cult. Sarcaulis Radlk. Sapotaceae (i). i N. Brazil, Guiana. Sarcinantiius Oerst. (Cariudovica p.p. BH.). Cyclanth. i Costa Rica. Sarcobatus Nees. Chenopodiaceae (B). 2 N. Am. Sarcocalyx Walp. = Aspalathus L. (Legum.). Sarcocapnos DC. Papaveraceae (in). 3 Medit. Sarcocarp, the fleshy part of a 'drupe ; -testa, of a seedcoat. Sarcocaulon Sweet. Geraniaceae. 5 S. Afr. Xero. ; fleshy stems. When the 1. falls the base of the petiole hardens to a thorn. Sarcocepfcalus Afzel. ex R. Br. Rubiaceae (i. 6). 10 palaeotrop. Sarcocnilus R. Br. Orchidaceae (n. 20). 40 Indomal. Polynesia. SarcocMaena Thou. Chlaenaceae. 4 Madag. Sarcochlamys Gaudich. Urticaceae (3). i Indomal. Sarcococca Lindl. Buxaceae. 5 Indomal. Sarcocodon N. E. Br. (Carallitma p.p. EP.). Asclep. (n. 3). 4 S.W. As. Sarcocolla L. Penaeaceae. 4 Cape Colony. Sarcodes Torr. Pyrolaceae. i California. Sarcodraba Gilg et Muschler (Draba p.p.). Crucif. (4). i Patag. Sarcodum Lour. Leguminosae (in. 6). i Cochinchina. Sarcoglottis Presl (Spiranthes p.p. BH.). Orchid, (n. 2). 10 trop., subtrop. Sarcolaena Thou. Chlaenaceae. 4 Madag. Sarcolobus R. Br. Asclepiadaceae (n. 3). 4 Malaya. Sarcomelicope Engl. (Evodia p.p.). Rutaceae (i). i New Caled. Sarcomphalus P. Br. Rhamnaceae. 5 W.I. S. laurinus Griseb. timber. Sarcopetalum F. Muell. Menispermaceae. i E. Austr. Sarcoplirynium K. Schum. Marantaceae. 12 W. trop. Afr. Sarcophyte Sparrm. Balanophoraceae. i Cape Colony. 588 SARCOPILEA Sarcopilea Urb. Urticaceae (2). i S. Domingo. Sarcopodium Limll. (Bulbophyllum p.p.)- Orchid, (n. 16). 20 E. Indomal. Sarcopteryx Radlk. Sapindaceae (i). 8 E. Indomal. Sarcopyramis Wall. Melastomaceae (i). i India. Sarcosperma Hook. f. Sapotaceae (i). 4 E. Indomal. Sarcostemma R. Br. Asclepiadaceae (n. i). 12 trop. and subtrop. %. Leafless xero. with slightly fleshy stems. Sarcostigma Wight et Arn. Icacinaceae. 3 Indomal. Sarcotheca Blume. Linaceae. i Borneo, Malay Peninsula. SarcotoecMa Radlk. (Cnpania p.p.; Ratonia BH.}. Sapindaceae (i). 2 Austr. Sarcozygium Bunge (Zygophyllum p.p..£P.). Zygophyll. i Mongolia. Sarga Ewart. Gramineae (8). i N.W. Austr. Sargentia S. Wats. Rutaceae (iv). i Mexico, Calif. Fr. ed. Sargentodoxa Rehder et Wilson. Lardizahalaceae. i China. Sarmentose, forming long runners. Sarmienta Ruiz et Pav. Gesneriaceae (i). i Chili. Sarothamnus Wimm. = Cytisus L. p.p. (Legum.). Sarotheca Nees = Justicia L. (Acanth.). Sarothra L. = Hypericum Tourn. (Guttif.). SarothrocMlus Schlechter (Trichoglottis p.p.). Orchiclaceae (n. 20). i Siam, Burma. Sarracenia L. Sarraceniaceae. 7 All. N. Am. (side-saddle fl.), in sunny marshy places. Low herbs with rosettes of rad. 1. ; each 1. is repres. by a long narrow pitcher with a flat green wing of tissue on the ventral side, serving chiefly for assim. The general structure of the pitcher is similar to that found in Nepenthes ; it has a fixed lid projecting over the mouth, and the lip is usu. turned down inwards. The mouth bears numerous honey-glands ; below these comes the 'slide-zone,' then the zone of hairs (cf. Nepenthes), and at the bottom is water in which the insects are drowned. The pitchers are often brightly coloured. In S. the entire 1. is a pitcher, while in Nepenthes it is only part of the L, and in Cephalotus only certain 1. Many cult. orn. ; many hybrids. Sarraceniaceae (EP., BH.}. Dicots. (Archichl. Sarraceniales ; Parie- tales BH.). 3 gen., 9sp., Am. Insectivorous pitcher-plants (see gen.) with rosettes of rad. 1. and 5 reg. fls. K 9 — 8 — 5, spiral, if > 5 the outer 3 small; C as many as inner sepals and alt. with them, or o; A oo ; G (6 — 5 — 3) with oo anatr. ov. on inrolled cpl. -walls. Loculic. caps, with oo seeds; endosp. fleshy. Genera: Heliamphora (raceme ; ovary 3-loc.), Sarracenia (fl. sol.; ovary 5-loc., the top of the pitcher simple), Darlingtonia (ditto, but the top of the pitcher fish-tail- shaped). Sarraceniales. The 2oth order of Dicots. Archichl. Sarsaparilla O. Ktze. Liliaceae (xi). i trop. Am. Sarsaparilla, Smilax. Sartwellia A. Gray. Compositae (6). 2 Texas, Mexico. Saruma Oliv. Aristolochiaceae. i S.W. China. Sasa Makino et Shibata (Bawbusa p.p.). Gramin. (13). 8 Japan. Sassafras L. Lauraceae. i Canada to Florida, S. ojficinale Nees et SAXIFRAGA 589 Eberm. (La urns Sassafras L.). The wood and bark yield oil of sassafras, used in medicine. Sassafridium Meissn. Lauraceae (i). 3 trop. Am., W.I. Satanocrater Schweinf. Acanthaceae (iv. A). 2 trop. Afr. Satinwood, Chloroxylon Swietenia DC. Sativus (Lat.), cultivated or planted. Sattadia Fourn. Asclepiadaceae (n. i). i C. Brazil. Satureia L. (incl. Calainintha Lam.). Labiatae (vi). 130 warm regions. Fls. gynodioec. S. hortensis L. and S. montana L. (summer and winter savories) cult, condiments. Saturna Nor. Euphorbiaceae (inc. sed.). Nomen. Satyria Klotzsch. Ericaceae (in. 2). 4 trop. Am. Satyrium L. Orchidaceae (n. i). 70 S. Afr. to trop. As. Fl. not twisted, so that labellum uppermost ; it is prolonged backwards into two spurs. The actual summit of the column is occupied by the stigma, the anther being bent round 1 to it. Sail, Albizzia stipitlata Boiv. Sauerkraut, Brassica oleraceae L. Saundersia Reichb. f. Orchidaceae (n. 19). i Brazil. Saurauia Willd. Dilleniaceae. 75 trop. As., Am. Saurog-lossum Lindl. (Spiranthes p.p. Bff.}. Orchid, (n. 2). 3 trop. Am. Sauromatum Schott. Araceae (vn). 6 palaeotrop. L. pedate (cy- mosely branched). Sauropus Blume. Euphorbiaceae (A. i. i). 20 Indomal. Saururaceae (EP. ; Piperaceae p.p. BH.). Dicots. (Archichl. Piperales). 3 gen., 4 sp., E. As. and N. Am. Herbs with $ fls., apparently primitively naked. A 6 or fewer; G 3 — 4 or (3 — 4), in the latter case with parietal plac. ; ov. orthotr. Endo- and peri-sperm. Chief genera: Saururus, Houttuynia. Saururus Plum, ex L. Saururaceae. i Japan to Phil. Is., i in E. U.S. Bog pi. with spikes of fls., br. usu. adnate to axis of its fl. Saussurea DC. Compositae (n). 125 N. temp. S. alpina DC. alpine in Brit., with hairy 1. ; its fls. a,re blue, with sweet scent (the latter unusual in the order). Many have 3 cpls. Sautiera Decne. Acanthaceae (iv. A), i Timor. Sauvagesia L. Ochnaceae (Viol. BH.}. 12 trop., esp. Brazil. 5 fertile sta., surrounded by oo stds. Cpls. 3. This gen. and a few others sometimes formed into a separate fam. or placed in Violaceae. Sauvallea Wright. Commelinaceae. i Cuba. Savannah, grass country broken by patches of forest or copse; -flower (W.I.), Echites; -wattle (W. L), Citliarexvlum. Savia Willd. Euphorbiaceae (A. i. i). 4 W. Ind. Savignya DC. Cruciferae (2). 2 E. Meclit. Savin, fit ni penis Sabina L. Savoury, Satureia. Saw wort, Serratula tinctoria L. Saxatilis (Lat.), growing on rocks. Saxegothaea Lindl. Coniferae (Taxac. 3; see C. for gen. char.). i Andes of Patagonia. Fr. a many-seeded 'berry' like Juniperus. Saxifraga Tourn. ex L. Saxifragaceae (i). 225 N. temp., Arctic, 59° SAXIFRAGA Andes, chiefly alpine. 13 in Brit, (saxifrage). Most show xero. char., such as tufted growth, close packing of 1. (esp. well shown in S. oppositifolia L.), succulence, hairiness, &c. Many are veg. repr. by offsets, or (e.g. S. granulata L.) by bulbils in the lower leaf-axils. Many exhibit chalk-glands at the tips or edges of the 1. (e.g. S. oppositi- folia at the tip) ; these are water-pores with nectary-like tissue beneath, secreting water containing chalk in solution. As the water evaporates, the chalk forms an incrustation. Fls. usu. in clich. cymes with cincinnus tendency. Every stage occurs from hypogyny to epigyny. Honey only partially concealed ; fls. visited by miscellaneous insects. Most are protandrous. A few, e.g. S. sarmentosa Linn, f., have -j- fls. Saxifragaceae (EP. ; BH. incl. Cephalotaceae, Ctmoniaceae). Dicots. (Archichl. Resales). 70 gen., 650 sp. cosmop., chiefly temp. Most are perenn. herbs, a few shrubs or trees, with usu. alt., rarely stip. 1. Many alpine and arctic forms of xero. habit. Infl. of various kinds, both racemose and cymose. FI. usu. g, reg., cyclic, 5-merous (exc. cpls.). Recept. flat or hollowed to various depths, so that sta. and P may be peri- or epi- gynous. K usu. 5; C 5, imbr. or valvate, sometimes (5) or o; A usu- 5 + 5. obdiplost.; cpls. rarely free and as many as petals, usually fewer and joined below, often 2 ; plac. parietal or axile, with several rows of anatr. ov.; styles as many as cpls. Fls. mostly protandrous. Capsule or berry. Seed with rich endosp. round a small embryo. Of little economic importance; Ribes yields valuable fr. Many favourites in horticulture, e.g. Saxifraga, Francoa, Philadelphia, Deutzia, Hydrangea, Escallonia. Classification and chief genera : I. SAXIFRAGOIDEAE (herbs of various habit ; 1. alt. ; P 5- or rarely 4-merous; G usu. (2), hypo- or epi-gynous, i- or 2-loc.) : Saxifraga, Tellima, Chrysosplenium, Parnassia. II. FRANCOIDEAE (perenn. herbs with rad. 1. and spikes or racemes on naked scapes; fl. 4-merous; G4-loc.): Francoa. III. HYDKANGEOIDEAE (shrubs or trees ; 1. usu. opp., simple ; Pusu. s-merous; sta. epig. ; 63— 5-loc.): Philadelphus, Deutzia, Hydrangea. IV. PTEROSTEMONOIDEAE (shrubs with alt. stip. simple 1. ; sta. 10 ; G 5-loc.; ov. 4 — 6, on axile plac.): Pterostemon. V. ESCALLONIOIDEAE (shrubs or trees, rarely herbs; with simple alt. exstip. often leathery and gland-dotted 1. ; sta. = pets. ; G superior to inferior; ov. oo ): Brexia. Escallonia. VI. RIBESIOfDEAE (shrubs with alt. simple exstip. 1. and racemes of fls. ; G i-loc. with 2 parietal plac.; berry): Ribes. VII. BAUEROIDEAE (shrubs with opp. 3-foliate exstip. 1. and simple axillary fls.; ovary semi-inf. with 2 parietal plac.; loculic. caps.): Bauera. Saxifrage, Saxifmga; burnet -, Pnnpinella ; golden-, Chrysosplenium. Saxifrageae (BH.} - Saxifragaceae. Saxifragella Engl. Saxifragaceae (i). i Antarctic S. Am. Saxifraginae (Warming). The i8th order of Choripetalae. Saxifragopsis Small (Saxifraga p.p.). Saxifr. (i). i Calif. Saxofridericia R. Schomb. Rapateaceae. 5 Guiana, N. Brazil. SCHAFFNERA 591 Sayeria Kranzl. Orchidaceae (n. 15). i New Guinea. Scabiosa (Tourn.) L. (/>//. incl. Knautia L., Pterocephalus Vaill., Sitccisa Neck.). Dipsacaceae. 60 Eur., Medit. ; 3 in Brit., incl. S. (A'.) arvemis L. (scabious) and S. Succisa L. (devil's-bit scabious). The former has a large head of fls. (class B'); the C is drawn out upon the outer side (cf. Compositae), and this the more the further from the centre of the head. Honey is secreted by the upper surface of the ovary, and protected from rain by hairs. The sta. are ripe first, while the style with immature stigmas is quite enclosed in the C; later the sta. wither and the style occupies their place. The stigmas of the various fls. on the head ripen nearly together. Scabious, Scabiosa ; sheep's bit -, Jasione montana L. Scabrous (dim. scabrid, scaberulous), scurfy. Scaevola L. Goodeniaceae. 65 Austr., Polynes. , trop. coasts. 6". Koenigii Vahl furnishes a kind of rice paper; its pith is squeezed flat. It is a char. pi. of trop. beach jungle. Scald, Cuscnta. Scale leaves, the scaly 1. on the outside of buds, rhizomes, &c., Araceae, Casuarina. Scalesia Arn. Compositae (5). 15 Galapagos. Scaligeria DC. (Conopodium p.p. BH.}. Umbellif. (in. 4). 10 E. Medit. Scalloped, crenate. Scaly bulb, one with overlapping L, Liliuin. Scammony, Convolvulus Scamnionia L. Scandens (Lat.), climbing. Scaadivepres Loes. Celastraceae. i Mexico. Scandix Tourn. ex L. Umbelliferae (in. 2). 12 Eur., Medit. S. Pecten- Veneris L. (Venus' comb) in Brit. The ripe mericarps separate with a jerk, Scape, a stalk from the base of the pi. bearing only fls., Amaryllidaceae, Taraxacum. Scapha Nor. = Saurauja Willd. (Dillen.). Scapruspatha Brongn. Araceae (vn). i Bolivia. Scaphocnlamys Baker (Kaemffera p.p. EP.}. Zingib. (i). 'i Malay Pen. Scaphoid, boat-shaped, petals of Loasa. Scaphopetalum Mast. Sterculiaceae. 5 trop. W. Afr. Scaphosepalum Pfitz. Orchidaceae (11. 8). 10 N. trop. S. Am. Scaphyglottis Poepp. et Endl. Orchidaceae (n. 6). 10 trop. Am. Scapigerous, scape-bearing. Scarious, thin, dry, not green, stiff. Scarlet runner, PAaseolus multiflorns Willd. Scelochilus Klotzsch. Orchidaceae (II. 19). 4 Andes. Scented fls., cf. Perfumed; -orchis, Gymnadenia conopsea R. Br. Scepa Lindl. = Aporosa Blume (Euph.). Sceptrocnide Maxim. Urticaceae (i). i Japan. Schachtia Karst. (Duroia p.p. EP.}. Rubiaceae (i. 8). i Colombia. Schaefferia facq. Celastraceae. 3 Texas to Colombia, W.I. Schaetzellia Sch.-Bip. Compositae (2). i Mexico. Schaffnera Benth. Gramineae (3). i Mexico. 592 SCHAFFNERELLA Schaffnerella Nash (Schaffnera p-p.). Gramineae (3). i Mexico. Schaueria Nees. Acanthaceae (iv. B). 8 trop. Am. Schedonnardus Steud. Gramineae (n). i N. Am. Schedonorus Beauv. = Bromus p.p. (Gram. ). Scheelea Karst. (Attalea p.p. EP.). Palmae (iv. 2). 5 trop. Am.^ Schefferomitra Diels (Mitrephora p.p.). Anon. (2). i New Guinea. Schefflera Forst. (incl. Heptaplenrnm Gaertn.). Araliaceae (i). 150 trop. Schefflerodendron Harms. Leguminosae (in. 6). 2 trop. Afr. Schelhammera K. Br. Liliaceae (i). 2 E. Austr. Schenckia K. Schum. Rubiaceae (i. 3). i S. Brazil. Scheucrileria Heynh. Compositae (inc. sed.). Nomen. Scheuchzeria L. Scheuchzeriaceae. i N. temp, and arctic, incl. Brit. ; S. palustris L., a marsh plant. Scheuchzeriaceae (EP.; Naiadeae p.p. BH.}(Juncaginaceae). Mono- cots. (Helobieae). 4 gen., 12 sp. temp. Perenn. marsh herbs of grass-like habit ; squamulae intravaginales (cf. Potamogetonaceae) in axils of sheathing 1. Fls. 5 or 3 ? in racemes or spikes, reg., greenish, wind-fert., protog. P 3 + 3, homochlam., A 3+ 3, extr., 63 + 3 some- times united, but the outer whorl often abortive; stigmas sessile; i anatr. ov. in each cpl Achene or schizocarp ; exalbum. ; embryo straight. Chief genera: Triglochin, Scheuchzeria, Lilaea. ScMckendautzia Pax. Amaryllidaceae (in). 2 Argentina. Schickendantziella Spegazzini. Liliaceae (inc. sed.). i Argentina. Schieckia Karst. (Celaslnis p.p. EP.). Celastraceae. i trop. S. Am. Schiedia Cham, et Schlecht. Caryophyllaceae (i. i). n Hawaiian Is. ScMedeophytum H. Wolff. Umbelliferae (in. 4). i Mexico. ScMekia Meissn. Haemodoraceae. i warm S. Am. ScMUeria Kunth = Piper L. p.p. (Pip-). Schima Reinw. ex Blume. Theaceae. 10 E. Indomal. 6". Wallichii Choisy good timber. ScMmmelia Holmes (Amyris p-p.). Rutaceae (iv). £ W.I. Schimpera Hochst. Cruciferae (2). 2 E. Medit. Schindleria H. Walter ( Villamilla p-p-)- Phytolacc. 5 Peru, Bolivia. ScMnnongla Schrank. Iridaceae (inc. sed.). i S. Afr. ScMnopsis Engl. (Qitebrachia BH.}. Anacardiaceae (3). 5 S. Am. The wood (quebracho) is hard and rich in tannin ; used for tanning. Schinus L. Anacardiaceae (3). 12 Mex. to Argent. 6". Molle L. yields American mastic (resin); cult, for shade, &c. (pepper-tree). Schinziella Gilg (Canscora p.p.). Gentianaceae (i). 2 trop. W. Afr. Schismatoclaea Baker. Rubiaceae (i. 5). 4 Madag. Schismatoglottis Zoll. et Mor. Araceae (v). 75 Malaya. At top of spadix, above thecffls., are sterile fls. consisting of stds. Schismus Beauv. Gramineae (10). 5 S. Afr., Medit. Schistocarpaea F. Muell. Rhamnaceae. i Austr. Schistocarpha Less. Compositae (8). 6 Mexico to Peru. Schistocaryum Franch. Boraginaceae (iv. 2). i Yunnan. Schistogyne Hook, et Arn. Asclepiadaceae (n. r). 4 S. Am. Schistonema Schlechter. Asclepiadaceae (n. i). i Peru. ScMstostephium Less. Compositae (7). 6 S.E. Afr. ScMstostigma Lauterb. Euphorbiaceae (A. n. 5). i New Guinea, SCHIZOSTA CH YUM 5 93 Schivereckia Andrz. ex DC. (Alyssum p.p. BH.}. Crucif. (4). 2 E. Medit. Schizachue Hack. Gramineae (10). i Saghalien. Schizachyrium Nees (Andropogon p.p. ). Gramineae (2). 8 N. Am. Schizaea Sm. Schizaeaceae. 25 trop. and subtrop. Sporangia in a double row on lower surface of each of the reduced fertile pinnae. Schizaeaceae. Filicales Leptosporangiatae. 5 gen., 70 sp., chiefly trop. Am.; a few subtrop. or temp., mostly small ferns with but little stem. Lygodium is a curious leaf-climber. As in Osmunda, the sporangia are borne (exc. in Mohria) on special pinnae of the leaf, distinct from the ordinary veg. pinnae. The sporangia are sessile, usu. without indusium; at the apex is a cap-like annulus, and the sporangium dehisces longitudinally. Chief genera: Schizaea, Aneimia, Lygodium, Mohria. Schizandra Michx. Magnoliaceae. 7 trop. and warm temp. As., N. Am. Climbing shrubs with exstip. 1. and spiral fls. Schizanthus Ruiz et Pav. Solanaceae (5). 15 Chili. Fl. •[• ; stalk curved, and the two really upper petals form the lower lip which is 3— 4-lobed, while the lat. petals are 4-lobed and the lowest petal forms the simple or slightly 2-lobed upper lip. Sta. 4, 2 fertile and 2 staminodial. Fl. like the papilionate Leguminosae (cf. Collinsia), and fert. in a similar way, usu. by an explosive movement (cf. Genista). Scnizeilema Domin (Pozoa Hook. f.). Umbel, (i. 2). 12 N.Z., Austr. Schizo- (Gr. pref.), split-; -carp, a fr. that splits up, without opening of the cpls., into i -seeded mericarps, Eitphorb., Gerart., Malpigh., Malv., Sapind., Tropaeol., Umbellif.; -genous, formed by splitting. Schizobasis Baker. Liliaceae (in). 5 S. Afr. Schizocalyx VVedd. Rubiaceae (i. 5). i Colombia. ScMzocapsa Hance. Taccaceae. i S.E. China. ScMzocarpum Schrad. Cucurbitaceae (3). 4 Mexico. ScMzocasia Schott. Araceae (vi). 2 Phil. Is., New Guinea. ScMzochilus Sond. Orchidaceae (it. i). 4 Cape Colony. Schizochlaena Thou. Chlaenaceae. 5 Madag. Sdrizocodon Sieb. et Zucc. Diapensiaceae. 2 Japan. Schlzodium Lindl. (Disa p.p. BH.}. Orchidaceae (n. i). ro S. Afr. ScMzoglossum E. i\ley. Asclepiadaceae (n. i). 50 trop. and S. Afr. Schizolaena Thou. Chlaenaceae. 5 Madag. ScMzolepis Schrad. ex Nees = Scleria Berg. (Cyper.). Schizolobium Vog. Leguminosae (n. 7). i S. Brazil. Schizoloma Gaudich. Polypodiaceae. 15 trop. and subtrop. Schizomeria D. .Don. Cunoniaceae. i E. Austr. Schizomeryta R. Viguier (Meryta p.p.). Aral. (i). i New Caled. Schizonepeta Briq. (Nepeta p.p.). Labiatae (vi). 4 temp. As. Schizonotus A. Gray. Asclepiadaceae (n. i). i California. Schizopepon Maxim. Cucurbitaceae (2). i China, Japan. Schizopetalon Sims. Cruciferae (3). 5 Chili. Schizophragma Sieb. et Zucc. Saxifragaceae (m). i Japan. Schizopremna Baill. Verbenaceae (4). i Timor. Schizoptera Turcz. Compositae (5). 2 Ecuador, Mexico. Schizoscyphus (Schizosiphon) K. Schum. Legum. (n. 2). i N. Guin. Schizostacriyum Nees. Gramineae (13). 10 E. As. w. 38 594 SCHIZOSTEPHANUS Schizostephanus Huchst. (rincetoxicum p.p. BH.}. Asclepiadaceae (II. i). i E. Afr. ScMzostigma Am. Rubiaceae (i. 7). i Ceylon. Ovary 5 — y-loc. Schizostylis Backh. et Harv. Iridaceae (in). 2 S. Afr. S. coccinea Eackh. et Harv. often cult, for its handsome fls. Schizotrichia Benth. Compositae (6). i Peru. Schizozygia Baill. Apocynaceae (i. 3). i Zanzibar. Schkuriria Roth. Compositae (6). u W. Am. Schlecritendalia Less. Compositae (12). i Brazil. A plant of very unusual habit (for this fam.), and with peculiar anatomy. Schlechterella K. Schum. Asclepiadaceae (i). i E. Afr. Schlechteria Bolus. Cruciferae (4). i W. Cape Colony. Schlechterina Harms. Passifloraceae. i S. Afr. Schlegelia Miq. Bignoniaceae (4). 3 W.I., Guiana. Schleichera Willd. Sapindaceae (i). i trop. As., S. trijuga Willd. (Ceylon oak). Useful limber ; aril of seed ed. ; oil expressed from seed itself. Furnishes the best lac (Mirzapore lac). Schleidenia Endl. = Heliotropium Tourn. (Borag.). Schleinitzia Warb. (Piptadenia p.p. EP.}. Legum. (i. 5). i N.G. Schleropelta Buckley = Hilaria H. B. et K. p.p. (i N. Am.). Schlimmia Planch, et Linden. Orchidaceae (n. 13). 2 Colombia. Scrilumbergeria E. Morr. Bromeliaceae (i). 3 S. Am. Schmalliausenia C. Winkler. Compositae (n). i C. As. Schmalzia Desv. (Rhus p.p.). Anacardiaceae (3). 50 N. Am. Schmidelia L. Sapindaceae (i). 55 S. Afr. and trop. Schmidtia Steud. C-ramineae (10). 2 Afr. Scrmella Raddi = Bauhinia L. p.p. (Legum.). Schoberia C. A. Mey. = Suaeda Forst. p.p. (Chenop.). Schoenefeldia Kunth. Gramineae (n). 4 trop. Afr. and As. Schoenia Steetz. Compositae (4). i temp. Austr. Schoenlandia Cornu. Pontederiaceae. i trop. Afr. Schoenobiblus Mart. Thymelaeaceae. 3 trop. S. Am., W.I. Schoenocaulon A. Gray. Liliaceae (i). 6 Am. Veratrin from seeds. Schoenocephalium Seub. Rapateaceae. 2 N. Brazil. Schoenocrambe Greene (Sisymbriiim p.p.). Cruciferae (2). 3 N. Am. Schoenodendron Engl. Cyperaceae (inc. sed.). i Cameroons. Tree. Schoenolaena Bunge (Xanthosia BH.}. Umbelliferae (i. i). 2 W. Austr. Schoenolirion Durand. Liliaceae (in). 4 N. Am. SchoenorcMs Reinw. Orchidaceae (n. 20). i Java. SchoenoxipMum Nees. Cyperaceae (in). 2 S. Afr. Schoenus L. Cyperaceae (n). 70 Austr., N.Z., a few in Malaya, Am., Eur.; JK nigricans L. in Brit. Scnoepfla Schreb. Olacaceae. 15 trop. Schoepfianthus Engl. ex De Wild. Olacaceae. i trop. Afr. Nornen. Schollera Roth = Vaccinium L. p.p. (Eric.). Scholtzia Schau. (Baeckea p.p. EP.}. Myrtaceae (n. i). 12 Austr. Schombocattleya, Schombolaelia, Schombolaeliocattleya, Schom- burgkiocattleya Hurt. Orchidaceae. Hybrids of Schomburgkia. Schomburgkia Lindl. Orchidaceae (11. 6). 12 trop. Am. Cult. Schotia Jacq. {Theodora Medic.). Legum. (n. 3). 6 trop. and S. Afr. SCIOPHYTIUM 595 Schoutenia Korth. Tiliaceae. 5 trop. E. As. Scbouwia DC. Cruciferae (2). i Arabia, N. Afr. Scbradera Vahl. Rubiaceae (r. 7). 5 W.I. to Amazon. Scbrankia Willd. Leguminosae (i. 3). 7 warm Am. Scbrebera Roxb. (Nathusia Hochst. ). Oleaceae. 6 Afr., India. Scbrebera Thunb. (Hartogia L. f. ). Celastraceae. 2 S. Afr., Madag. Scbrenkia Regel et Schmalh. Umbelliferae (in. 3). •; N. As. Scbstscburowskia Schrenk. Umbelliferae (inc. sed.). r Kokania. Scbubea Pax. Euphorbiaceae (A. 11. 2). i Cameroons. Scbubertia Mart. (Araujia BH.). Asclepiadaceae (n. i). 6 S. Am. Scbultesia Mart. Gentianaceae (i). 18 trop. Am., Afr. Schultzia Sprang. Umbelliferae (in. 5). 4 C. As., W. India. Scbumacberia Vahl. Dilleniaceae. 3 Ceylon. Schumannia Kuntze (Ferula p.p.). Umbellif. (in. 6). i Turkestan. Scbumanniantbus Gagnep. (Phrynium p.p.) Marantaceae. i S.E. As. Scbumamriopbyton Harms (Tetrasfigma p.p.)- Rubi. (i. 8). i W. Afr. Scbuurmansia Bkime. Ochnaceae (Viol. BH.}. 5 Malay Archipelago. Scbuurmansiella H. Hallier (Schuurmansia p.p.). Ochn. i Borneo. Scbwabea Endl. Acanthaceae (iv. B). 5 Afr. Scbwackaea Cogn. Melastomaceae (i). i C. Am. Scbwalbea L. Scrophulariaceae (in. 3). r E. N.Am. Scbwannia Endl. Malpighiaceae (i). ' 6 Brazil. Scbwartzkopffia Kranzl. Orchidaceae (11. i). i W. trop. Afr. Scbweiggeria Spreng. Violaceae. i Brazil. Scb.weinfurtb.ia A. Br. Scrophular. (n. 3). 3 E. Afr. to W. India. Scbweinitzia Ell. Pyrolaceae. i N. Am. Schwendenera K. Schum. Rubiaceae (11. 10). i Sao Paulo. Scbwenkia L. Solanaceae (5). 20 trop. Am., Afr. Scbychowskya Endl. (Fleurya p.p.). Urtic. (i). i palaeotrop. Sciadodendron Griseb. Araliaceae (2). i C. Am. Sciadonardus Steud. Gramineae (nomen). i N. Am. Sciadopanax Seem. (Panax p.p. BH.}. Araliaceae (i). i Madag. SciadopbyUum P. Br. (Schefflera p.p. EP.\. Arali. (i). 25 trop. Am. Sciadopitys Sieb. et Zucc. Coniferae (Pinaceae 22 ; see'C. for gen. char.), i Japan, S. verticillata Sieb. et Zucc., the parasol-pine or umbrella-fir, planted round temples. Short shoots crowded together at ends of annual long shoots. Each like Pinus except that the two green needle-1. are 'fused' into a single needle grooved down the centre, so that at first glance they seem to be whorls of ordinary 1. at the tip of each year's growth. The cones take two years to ripen. The wood is useful. Sciadotenia Miers. Menispermaceae. 10 trop. S. Am. Sciapbila Blume. Triuridaceae. 10 trop. Cf. Flora, 101, p. 395. Scilla L. Liliaceae (v). 100 temp. |*. S. festalis Salisb. (S. nutans Sm.), wild hyacinth or English bluebell, in Brit. Bulbs with racemes. Scillopsis Lem. = Lachenalia Jacq. p.p. (Lili.). Scindapsus Schott. Araceae (n). 20 Indomal. Scion, young shoot, twig used for grafting. Sciopbytium (Cl.), a shade formation. 38-2 596 SCIRPODENDRON Scirpodendron Zippel. Cyperaceae (u). 2 Indomal. Scirpus (Tourn.) L. Cyperaceae (i). 200 cosmop. char., of wet moors, bogs and marshes ; 15 in Brit. Stem usu. erect and angular, bearing 3 ranks of 1. reduced to sheaths, and performing assim. Its base often gives rise to creeping rhiz. or to shoots ending in tubers like potatoes. The racemose many-flowered spikelets are aggregated into a terminal tuft. Fl. $ , with 6 P-scales in two whorls; in many sp. protog. ; in all wind-pollinated. S. lacustris L., sometimes termed bulrush, is used for matting, chair-seats, &c. Scitamineae. The roth order (EP.) of Monocots. A fam.^ (BH.) in Epigynae, incl. Musaceae, Marantaceae, Cannaceae, and Zingibera- ceae of Engler. The 6th order (Warming) of Monocots. Sclarea Tourn. ex Mill. = Salvia L. p.p. (Labi.). Scler-, Sclero-(Gr. pref.), hard ; -caulous, with dry hard stem ; -enchyma, tissue with thick hard walls; -phyllous, with thick hard 1.; -sis, hardening by lignification ; -testa, woody layer of seed coat. Sclerachne R. Br. Gramineae (i). i Java. Scleranthaceae or Illecebraceae. See Caryophyllaceae. Scleranthus L. Caryophyllaceae (i. 6). 10 Eur., As., Afr., Austr.; S. animus L. (knawel), &c. in Brit. Fls. apet., self-fert. Scleria Berg. Cyperaceae (n)- 100 trop. Sclerocalyx Nees (Gymnacanlhus EP. ). Acanthaceae (iv. A), i Mexico. Sclerocarpus Jacq. Compositae (5). 7 Texas, C. Am., trop. Afr. Sclerocarya Hochst. Anacardiaceae (2). 3 trop. Afr. Ed. fr. SclerocepHalus Boiss. Caryophyll. (i. 4). i Medit., a char, plant. SclerocMton Harv. Acanthaceae (iv. B). 5 Afr. Sclerochlaena Baill. Chlaenaceae. i Madag. Sclerochloa Beauv. Gramineae (10). i Eur., As. Sclerochorton Boiss. Umbelliferae (in. 5). i W. As. Sclerodactylon Stapf. Gramineae (10). i Madag. Sclerodictyon Pierre. Apocynaceae (i. i).. i Gaboon. Sclerolaena R. Br. (Bassia p.p. EP. ). Chenopodiaceae (A). 6 Austr. Sclerolepis Cass. Compositae (2). i W. U.S. Sclerolobium Vog. Leguminosae (il. 8). 15 Brazil, Guiana. Scleromelum K. Schum. et Lauterb. Santalaceae. i New Guinea. Scleromitrion Wight et Arn. = Hedyotis L. (BH.} = Oldenlandia L. Scleronema Benth. (Catostemma BH.}. Bombacaceae. i Brazil, Ecuador. Scleroolaena Baill. (Xylochlaena EP.). Chlaenaceae. i Madag. Scleropliylax Miers. Solanaceae (4). 4 Argentina. Scleropoa Griseb. (Festuca p.p. BH.). Gramineae (10). 2 Medit. Scleropogon Phil. Gramineae (10). 3 Chili, Mexico, Texas. Scleropyrum Arn. Santalaceae. 2 Indomal. Sclerosia Klotzsch. Ochnaceae (nomen). i Guiana. Sclerosperma G. Mann et H. Wendl. Palmae (iv. i). i W. trop. Afr. Sclerostylis Blume — Atalantia Correa (Rut.). Sclerotheca A. DC. Campanulaceae (in). 2 Society Is. Sclerothrix C. Presl. Loasaceae. i Mexico to Brazil and Peru. Scobedla Labill. ex Steud. Labiatae (nomen). i, habitat? Scoke (Am.), Phytolcuca. SCR OP HULA RIA CEAE 597 Scoliopus Torr. Liliaceae (vil). i W. N. Am. Scoliotheca Baill. Gesneriaceae (2). i Colombia. Scolochloa Link (Graphephorum BH.). Gramineae (10). 2 N. temp. Scolopendrium Adans. = Phyllitis Ludwig (Polypod.). 6". vnlgare Sm. (hart's tongue fern, Brit.) = P. Scolopendrium. Scolopia Schreb. Flacourtiaceae (5). i8warm|=£; S. Afr. sp. timber. Scolosanthes Vahl. Rubiaceae (n. 3). 6 W.I. Scolymus Tourn. ex L. Compositae (13). 3 Medit. Scoparia L. Scrophulariaceae (in. i). 10 trop. Am. Scopolia Jacq. Solanaceae (2). 4 Eur., As. Scopularia Lindl. (Holothrix BH.}. Grchidaceae (n. i). i Cape Col. Scopulophila M. E. Jones. Caryophyllaceae (i. b). i W. U. S. Scorodendron Blume (Lepisanthes p.p. EP.). Sapindaceae (i). i Timor. Scorodocarpus Becc. Olacaceae. i Borneo. Scorodonia Adans. =Teucrium L. p.p. (Labi.). Scorodophloeus Harms. Leguminosae (n. 2). i Cameroons. Scorpaena Nor. Inc. sed. Nomen. Scorpioid, inn. with fls. in 2 ranks, coiled like a scorpion's tail, Bora- gtnaceae. Scorpion grass, Rlyosotis. Scorpiurus L. Legumin. (ill. 7). 6 Mcdit. Pod twisted, indeh. Scortechinia Hook. f. Euphorbiaceae (inc. sed.). 4 Malaya. Scorzonella Nutt. (Klicroseris p.p.). Compositae (13). 3 Calif. Scorzonera (Tourn.) L. Compositae (13). 100 Medit., C. Eur. and As. Roots of 6'. hispanica L., &c. are eaten as vegetables. Scotch attorney (W. I. ), Clusia; -fir, Pituissylvestris'L.; -grass (W. I.), Panicnm molle Sw. ; -tish asphodel, Tofieldia. Scottellia Oliv. Bixaceae. 3 trop. W. Afr. Scouring rush (Am.), Equisctum. Scratch coco (W.J.), Colocasia antiqiiontm Schott. Screw pine, Pandanaceae, Pandanus; -tree (W.I.), Helicteres, Scribneria Hackel (Lfpturns p.p.). Gramineae (12). i Calif., Oregon. Scrobiculatus (Lat.), pitted. Scrofella Maxim. Scrophulariaceae (n. 4). i China. Scrophularia Tourn. ex L. Scrophulariaceae (11. 4). 120 N. temp.; 3 in Brit., incl. S. aquatica L. and S. nodosa L., the fig-worts. Perenn. herbs with opp. 1., which on the lat. twigs are commonly anisophyllous. Fls. in tall infls. whose primary branching is race- mose; the lat. shoots are dichasial. Sta. and style arranged along the lower lip of the C (upper usual in such fls.); The posterior sta. , usu. absent in the fam., is repres. by a std. Fl. markedly protog., largely visited by wasps. Scrophulariaceae (EP., BH. unite n. 7 to Globitlariaceae to form fam. Selagineae). Dicots. (Sympet. Tubiflorae; Personales BH.). 200 gen., 2600 sp., cosmop. Most are herbs and undershrubs, a few shrubs or trees (e.g. Paulownia), with alt., opp., or whorled exstip. 1. Many exhibit interesting features in the veg. organs. Several are climbers (e.g. Maurandia, Rhodochiton, &c. ). The Veronicas of N.Z. are xero. with resemblance in habit to certain Coniferae. A 598 SCROPHULARIA CEAE number of sp. in in. 2 and 3 (below), e.g. Euphrasia, Bartsia, Pedi- cularis, grow in swampy grass-land and are parasitic by their roots upon the roots of the grasses. Suckers are formed at the points of contact, in spring; they absorb food till the summer, and later absorb organic compounds from the dead parts of the host, and function for storage of reserve-materials. The plants possess green 1. of their own, and so are able to assimilate. Infl. racemose or cymose, in the former case usu. a spike or raceme, axillary or term, (every variety in sp. of Veronica). Sol. Floral diagrams of A, Vcrbascuin nigrum, B, Linaria vulgaris, C, Veronica Chaniaedrys ; after Eichler. axillary fls. in many, e.g. Linaria. Cymose infls. usu. dichasia, often united into complex corymbs, &c. Bracts and bracteoles usu. present. In Castilleja the upper 1. and bracts brightly coloured. Fl. 5 > •!• > sometimes nearly reg. (Verbascum, &c.) ; considerable variety in structure, as illustrated by the floral diagrams given. The bulk of the fam. show the Linaria type. K (5), of various aestiva- tions ; C(5), median -|- , often 2-lipped ; A4 (sometimes 2), didynamous, epipet., the post. sta. sometimes repres. by a std. (e.g. in Scrophularia and Pentstemon). Verbascum and its allies have an actinom. C and 5 sta. ; Veronica (q-v.) shows 4 sepals (the post, one of the typical 5 absent), 4 petals (the post, pair of the .s united), and 2 sta., the C. rotate. Other variations occur in the Selagineae, &c. Below the ovary is a honey-secreting disc. G (2), medianly placed (not obliquely as in Solanaceae), 2-loc., with axileplac.; ov. usu. oo , less commonly few (e.g. Veronica, &c.), anatr. ; style simple or bilobed. Fr. sur- rounded below by the persistent K, usu. a capsule (dehisc. in various ways) or a berry. Seeds usu. numerous, small, with endosp. Embryo straight or slightly curved. Most have fls. ± adapted to insect-visits. Miiller divides them into' 4 types: (i) the Verbascum or Veronica type (see gen.) with open fl. and short tube (bees and flies), (2) the Scrophularia type (wasps), (3) the Digitalis and Linaria type with long wide tubes and the essential organs so placed as to touch the back of the insect (bees), and (4) the Euphrasia type or 'loose- pollen' fl., where the pollen is loose and powdery, and the anthers (protected by upper lip) have spines, &c., so that they may be shaken upon the entrance of the insect, which thus receives a shower of pollen. The fls. are seldom markedly dichogamous, but the stigma usu. projects beyond the sta. SCUTCH GRASS 599 so as to be first touched. Most are capable of self fert. in default of visits. For further details see gen. In Linaria, &c. (fl-v. ) there sometimes appears a terminal fl. to the raceme, and this exhibits peloria, having a symmetrical C with spurs to all the petals (cf. Ruta, or compare Aquilegia with Delphinium). A number are or have been officinal, e.g. Digitalis ; most are poisonous. Many favourite cult. orn. plants, e.g. Calceolaria, Mimulus, Pentstemon, Antirrhinum, Linaria, Veronica, Collinsia, &c. Classification and chief genera (after von Wettstein) : A. Two post. C-teeth (or upper lip) cover lat. teeth in bud. I. PSEUDOSOLANEAE (all 1. usu. alt. ; 5 sta. often present): i. lrerbasceae (C with very short tube or none, rotate or shortly campanulate) : Verbascum, Celsia. i. Aptosinuae (C with long tube): Aptosimum. II. ANTIRKHINOIDEAE (lower 1. at least opp. ; the 5th sta. wanting or staminoclial) : a. C 2-lipped ; lower lip concave, bladder-like. 1. Calccolarieae: Calceolaria. /3. C almost actinom., or 2-lipped with flat or convex lips. 2. Henii liter ideae (dehisc. caps. ; C spurred or saccate at base, with no tube) : Alonsoa. 3. Antirrhineae (as i, but with tube): Linaria, Antirrhinum, Maurandia, Rhodochiton. 4. C/ieloncaL' (dehisc. caps, or many-seeded berry ; C not spurred or saccate; infl. cymose, cpd.) : Russelia, Wightia, Col- linsia, Scrophularia, Chelone, Pentstemon, Paulownia. 5. Mamtleae (dehisc. caps. ; C as in 4 ; infl. not cymose, usu. simple; anthers finally i-loc.): Zaluzianskia, Lyperia. 6. Gratiolcae (as 5, but anthers finally 2-loc.): Mimulus, Gra- tiola, Torenia. 7. Selagineae (drupe or indeh. few-seeded caps.) : Hebenstretia, Selago. B. Two post, teeth (or upper lip) of C covered in bud by one or both of the lat. teeth. III. RHINANTHOIDEAE. a. C-teeth all flat and divergent, or the 2 upper erect. 1. Digitaleac fanther-loc. finally united at tip ; 2 upper C-lobes often erect ; not paras.) : Veronica, Digitalis. 2. Geraniicae (anther-loc. always separate, one often reduced; C-lobes all flat, divergent; often paras.): Gerardia. b. 2 upper C-teeth form a helmet-like upper lip. Often paras. 3. Rh inantheaf : Castilleja, Melampyrum, Tozzia, Euphrasia, Bartsia, Pedicularis, Rhinanthus. Scrophularineae (BH.}~ Scrophulariaceae. Scrub, cf. Acacia ; -myrtle (Austr.), Backhoiisia. Scubalia Nor. Inc. sed. Nomen. Scurrula L. = Loranthus L. (Lor.). Scurvy grass, Cochlcaria officinal is L. Scutachne Hitchc. et Chase. Gramineae (5). 2 Cuba. Scutate, scutiform, buckler-shaped. Scutch grass (Am.), Cynodon Dae ty Ion Pers. 6oc SCUTELLARIA Scutellaria Riv. ex L. Labiatae (iv). 200 cusmop. exc. S. Afr. 2 in Brit., S. galericulata L. and 6". minor Huds. (skull-cap). Scutellum, Gramineae. Scutia Comm. ex Brongn. Rhamnaceae. 3 S. Afr., trop. Scuticaria Lindl. Orchidaceae (n. 18). 2 trop. S. Am. Epiphytes, with no tubers, but long pendulous fleshy cylindrical stems. Cult, orn. fl. Scutinanthe Thw. (Canarium p.p. BH.}. Burseraceae. i Ceylon. Scybalium Schott et Endl. Balanophoraceae. 4 trop. Am., W.I. Scyphanthus D. Don (Grammatocarpus BH.}. Loasaceae. 2 Chili. Scypharia Miers (Colletia BH.}. Rhamnaceae. 4 W. trop. Am. Scyphellandra Thw. (Ahodeia p.p. BH.}. Violaceae. i Ceylon. Scyphiphora Gaertn. f. Rubiaceae (i. 8). i Indomal. Scyphlphorous, cup-bearing. Scyphocephalium Warb. Myristicaceae. 6 trop. W. Afr. Scyphochlamys Balf. f. Rubiaceae (n. i). i Rodrigues. Scyphocoronis A. Gray. Compositae (4). i W. Austr. Scyphogyne Brongn. Ericaceae (iv. 2). 10 S. Afr. Scy phony chium Radlk. (Cupania p. p. ). Sapind. (i). i N.E. Brazil. Scyphopetalum Hiern (Paranephdinm p.p. EP.}. Sapind. (i). i Burma. Scyphostachys Thw. Rubiaceae (i. 8). 2 Ceylon. . Scyphostegia Stapf. Monimiaceae (??). i Borneo. Scyphostelma Baill. Asclepiadaceae (n. i). i Colombia. Scyphostrychnos Sp. Moore. Loganiaceae. i Nigeria. Scyphosyce Baill. Moraceae (n). 2 trop. W. Afr. Scytalia Gaertn. = Nephelium L. (Sapind.). Scytanthus T. Anders. (Thomandcrsia EP.}. Acanth. (iv. B). i W. Afr. Scytanthus Liebm. (Cytinns p.p. BH}. Rafflesiaceae. 4 Mexico. Scytopetalaceae (EP.}. Dicots. (Archichl. Malvales). Only gen. Scytopetalum, tj.v. Scytopetalum Pierre. Scytopetalaceae. 10 trop. W. Afr. Trees or shrubs with alt. leathery 1. and long-stalked fls. K dish-like, C 3 — 7, valvate, A oo , G (4 — 6), each with 2 — 6 pend. ov. Fr. woody or drupaceous, i -seeded. Sea bean, Entada scandens Benth. ; - beet, Bda vulgaris L. ; - blite, Sueu'iia marit'nna Dum. ; -coast and salt-steppe pi., cf. Halophytes; -buckthorn, Hifpophae rhamnoi&s L. ; -grape, Coccoloba uvifera L. &c. ; -heath, Fraiikcuia; -holly, Erynginui ; -Island cotton, Gossypinm barbadensc L. ; - kale, Crainbe niaritiina L. ; - lavender, Statice', -milkwort, Glaitx inariti»ia L. : -pink, Armeria vitlgaris Willd. ; -purslane, Arenaria peploides L. ; -side grape, Coccoloba \ -rocket, Cakile t/iarithna Scop. Seaforthia R. Br. = Ptychosperma Labill. (Palm.). Sebaceus (Lat.), like lumps of tallow. Sebaea Soland. ex R. Br. Gentianaceae (i). 100 warm |^. Sebastiania Spreng. Euphorbiaceae (A. n. 7). 45 trop., U.S. Sebastiano-Schaueria Nees. Acanthaceae (iv. B). i Brazil. Sebertia Pierre. Sapotaceae (ll). i New Caledonia. Sebestens, fr. of Coniia J/iuv? I .. SEED 60 1 Sebicea Pierre ex Diels^Tiliacora Colebr. (Menisp.). Sebizia Mart. Vitaceae. Nomen. Secale (Tourn.) L. Gramineae (12). 2 Medit., Eur., As. S. cereale\^., the rye, is largely cult, in N. Eur. as a cereal, forming a staple food. There are no well-marked races. The hardy winter ryes are the best. Also used as fodder. Secamone R. Br. Asclepiadaceae (n. 2). 60 palaeotrop. Secamonopsis Jumelle. Asclepiadaceae (n. i). i Madag. Sechiopsis Naud. Cucurbitaceae (4). i Mexico. SecMum P. Br. Cucurbitaceae (4). i trop. Am., S. edule Sw., cult, for its ed. fr. (chocho), containing one enormous seed. Secondary shoots, adv. shoots when primaries occur, Ailantkus, An- thurium, Podostemaceae, Testudinaria, Tristichaceae. Secondatia A. DC. Apocynaceae (n. i). 5 trop. S. Am. Secretaria Muell.-Arg. (Hyeronyma BH.). Euph. (A. I. i). i Gui. -sect, divided to the base. Secula Small. Legiiminosae (in. 7). i W.I., Florida. Secund, all directed to the one side. Secundine, inner coat of ovule. Securidaca L. Polygakceae. 32 trop., exc. Austr. Climbers. Securigera DC. (Bonavcria Scop.). Leguminosae (in. 5). i Medit. Securinega C'omm. ex Juss. Euphorb. (A. I. i). 10 temp, and sub- trop. Sedastruru Rose. Crassulaceae. 7 Mexico. Seddera Hochst (Breweria p.p.)- Convolvulaceae (i). 6 palaeotrop. Sedella Britton et Rose (Seditm p.p.). Crassulaceae. 2 Calif. Sedge, Carex,Cyperaceae; cotton-, Eriophorum; sword-; Lepidosperma. Sedum Tourn. ex L. Crassulaceae. 150 N. temp., i in Peru; 9 in Brit., incl. S. Telephiuin L. (orpine or livelong), 5". Rhodiola DC. (rose-root), S. anglicum Huds. and 6". acre L. (stonecrop or wall- pepper). Fleshy-leaved xero. Cult. orn. fl. and fol. Seed, the product of the ovule after fert. ; it contains the embryo, and if nothing else, is exalbuminons, but may contain reserve material out- side the embryo (albuminous), which may be derived from the interior of the embryo-sac (endosperm), or from tissue outside this (perispertu). Usually mentioned after the fr. in descriptions. In form, &c. it may be large or small ; spherical, ellipsoidal, &c. ; anatr., amphitr., &c. like the original ovule. The testa may be smooth, or covered with small or large tubercles, papillae, granules, ribs, &c. ; green, brown, or of other colours ; thin or thick, woody (as in Bertkolletia), with a fleshy outer layer (as in Bijca, Cycas, Magnolia, Moraea, &c.), hooked or winged (see below), or provided with hairs. It is usu. firm and tough, allowing very little evap. of water from the seed. Its outer cells sometimes have mucilaginous walls and swell when wetted, as in Brassica, Collomia, Liniun, Plantago, &c.; sometimes there are scales or hairs upon it and these swell, as in many Acanthaceae. The testa is usu. marked with a scar, the hilum, where the stalk separated from the seed. Many are covered by an extra coat (aril] that is developed like the original integuments, i.e. by a cup-like growth of tissue around the seed from the top of the stalk or rarely from the micropyle. Its 602 SEED development may be studied in fr. of Enonymus. Arillate seeds in Celastraceae, Commelinaceae, &c., Dilleniaceae, Myristica, Nymphaea- ceae, Passiflora, Sapindaceae, Taxus, &c. In Euphorbiaceae the aril remains small and hard and is called a caruncle, but usu. it is fleshy. The endosp. may be starchy, farinaceous, floury, or mealy (the cells containing starch and when powdered forming a floury dust) as in Triticum, oily as in Papaver, Ricinus, &c., fleshy as in Berberis, horny as in Coffea, bony as in Phoenix and Phytelephas, ruminate (marked by wavy transverse lines which give it a marbled look) as in Anonaceae, mucilaginous, &c. The embryo (rarely embryos) may be straight, curved, twisted, &c. (and cf. Cruciferae). It may have one, two, or several cotyledons (im- portant in classification), or rarely none, as in Cuscuta, &c. Mention must be made of adventitious embryos of Alchornea, Funkia, Nothoscordum, &c., and of the peculiar cases of Ginkgo, and Gnetum, where the seed 'ripens' before fertilisation. Seemannaralia R. Viguier. Araliaceae (i). i S. Afr. Seemannia Regel. Gesneriaceae (n). 5 Peru, Bolivia. Seetzenia R. Br. Zygophyllaceae. i Afr., As., in deserts. Segetalis (Lat.), growing in grain fields. Segments, the free portions of a concrescent K or C. Segregate, kept separate. Seguieria Loefl. Phytolaccaceae. 25 S. Am. L. leathery; stipules thorny. Powerful odour of garlic. Cpl. i. Fr. a samara. Seidelia Baill. Euphorbiaceae (A. n. 2). i S. Afr. Seidlitzia Bunge et Boiss. Chenopodiaceae (B). i Armenia, Persia. Selagineae (Bff.). See Scrophulariaceae and Globulariaceae. Selaginella Spring. The only genus of Selaginellaceae. 700 chiefly trop. ; a few temp. e.g. selaginoidcs Link on boggy hill sides in Brit. Most live in damp places, esp. in forests, but a few xero. The embryo has a suspensor, and grows directly into the leafy plant which shows a habit very similar to Lycopodium — much-branched stem, often creeping, bearing roots on lower side and 1. on upper, with term, cones of sporangia. L. spirally arranged as in most Lycopodiums (e.g. in S. selaginoides], or more commonly in 4 ranks, two outer ones of large, two inner ones of small L, thus giving the stem a dorsiventral structure. The roots in some, e.g. S. Kraussiana A. Br. and S. Martensii Spring, are borne on rhizophcres, anomalous stem branches developed at the nodes and exhibiting a sort of intermediate structure between stem and root. The sporangia are placed at the bases of the 1. in term, cones of radial symmetry. The mega-sporangia contain 4 large spores and can easily be distinguished by eye. On germ, a microspore produces a rudimentary S prothallus bearing an anthe- ridium. The megaspore forms a ? prothallus, which remains en- closed in the burst spore, and has an upper small-celled green portion and a lower large-celled storage portion as in Salviniaceae, &c. S. lepidophytta Spring is a xero. which curls into a ball in the dry season, and may be rolled about by wind (cf. Anastatica). Selaginellaceae. Lycopodiales. Only genus Selaginella (q.v.). Selago L. Scrophulariaceae (n. 7). 140 S. and trop. Afr., Madag. Selenia Nutt. Cruciferae (2). 2 S.W. U.S. SENECIO 603 Selenicereus Britton et Rose (Cereus p.p.)- Cactaceae (ill. i). 12 W.I., C.Am., Mexico. Selenipedium Keichb. f. Orchidaceae (i. 2). 2 N. trop. S. Am. Ov. 3-loc. Selenocera Zipp. ex Span. Rubiaceae (inc. sed.). i Timor. Self coloured, of uniform tint ; - fertilisation, from the pollen of the same fl., Cephalanthera, Ophrys, Senecio, many annuals, Cleisto- gamic fls., &c. ; -burying fl., Trifolium, Trigondla^ft., Arachis., Voandzeia, seed, Erodium ; - heal, Prunella ; - sterility, sterility to its own pollen, Abittiloii, Corydalis. Selinocarpus A. Gray. Nyctaginaceae. 4 S.W. U.S., Mexico. Selinum L. Umbelliferae (in. 5). i6j*. Selkirkia Ilemsl. Boraginaceae (iv. i). i Juan Fernandez. Selleophytum Urb. Compositae (5). i Haiti. SellieraCav. Goodeniaceae. 2 Austr., N.Z., temp. S. Am. Fr. indeh. Selliguea Bory = Polypodium L. (Polypod. ). Selloa H. B. et K. Compositae (5). i Mexico. Sellocharis Taub. Leguminosae (ill. 3). i S.E. Brazil. Selwynia F. Muell. (Coccnhis p.p. EP.}. Menispermaceae. i Austr. Selysia Cogn. Cucurbitaceae (2). 2 Brazil, Colombia. Semaphore plant, Dcsmodium gyrans DC. Semecarpus L. f. Anacardiaceae (4). 40 Indomal. The young fr. yields a black resin used as marking ink, &c. Semeiandra Hook, et Am. Onagraceae (2). i Mexico. Semele Kunth. Liliaceae (vn). i Canaries, S. androgyna Kunth (Ruscus androgytnis L.). A climbing shrub with leaf-like phylloclades in the axils of scale-1. Fls. in little cymes (cf. Asparagus) on edges of phylloclades. The new shoots rise from the soil, and grow long before the lat. branches, bearing the phylloclades, begin to unfold. Semenovia Regel et Herder. Umbelliferae (inc. sed.). i C. As. Semiaquilegia Makino (fsopyrnm p.p.). Ranunc. (2). i Japan. Semibegoniella C. DC. Begoniaceae. 2 Ecuador. Seminiferous, seed-bearing. Semiramisia Klotzsch (Thibaudia p.p. EP.). Eric. (in. 2). 2 S.Am. Semonvillea J. Gay. Phytolaccaceae (Aizo. BH.}. i S. and trop. Afr. Sempervirens (Lat.), evergreen. Sempervivum Rupp. ex L. Crassulaceae. 50 S. Eur., Himal., Abyss., &c. S. tectorum L. (houseleek, planted on cottages to keep slates in position) is a xero. with fleshy I. and veg. repr. by offsets. Senaea Taub. Gentianaceae (i). i Minas Geraes. Senapea Aubl. Inc. sed. i Guiana. Senebiera DC. (Coronopus Rupp.). Cruciferae (2). 12 subtrop., Eur.; one nat. in Brit., S. Jidytna Pers. Senecio (Tourn.) L. (incl. Cineraria L. p.p., Kleinia Haw., Ligularia Cass., &c.). Compositae (8). 1450 cosmop., 9 in Brit, (ragwort, groundsel, &c.). Shows § chars, well. The gen. includes pi. of most various habit. Some are climbers, e.g. S. macroglossns DC. (S. Afr.), which is remarkably like ivy. Many are xero. some with fleshy L, others with fleshy stems, others with hairy or inrolled 1. (cf. Empetrum). The fls. of 51. vulgaris L. (groundsel) are regularly self-fert. and are very inconspic.; there are no ray-florets. In S. 604 SENECIO Jacobaea L. (rag-wort) there are ray-florets, and the conspic. fls. are largely visited by insects. The. fleshy stems of S. (K.) articulatus Sch. Bip. (S. Afr.) separate at the joints and grow into new pi. S. Johnstoni Oliv. is a remarkable tree on Kilimandjaro. Senefeldera Mart. Euphorbiaceae (A. n. 7). 4 Brazil. Senega snake root, Poly gala Senega L. Senkenbergia Schau. (Boerhaavia p.p. EP.}. Nyctagin. 3 N. Am. Senna Touin. ex Mill. = Cassia Tourn. p.p. (Legum.). Senna, Alexandrian, Arabian, Italian, Cassia ; bladder-, Colutea. Sennenia Pau ex Sennen (Trisetum p.p.). Gramin. (9). i Eur. Senra Cav. Malvaceae (4). i E. Afr., Arabia. Sensitive fern (Am.), Onoclea; -1. and petioles, cf. Climbing PI., Insectivorous PI., Movements; - plant, Alimosa piidica L., Neptunia, &c. ; - sta., Berberis, Centattrea and many Conipositae, Portitlaca, Sparmannia ; - stigma, Martynia, Mininlus, Strobilanthes. Sepal, cf. Perianth; -oid (P), all of sepal-like 1. Sepalosiphon Schlechter. Orchidaceae (n. a. in), i New Guinea. Septal, of hedgerows. Septas L. =Crassula Dill. p.p. (Crass.). Septate, divided by a partition wall or walls. Septemfid, in seven divisions. Septicidal, septifragal, cf. Fruit. Septum, a partition, e.g. in a multiloc. ovary. Sequoia Endl. (Washingtonia Winsl., Wellingtonia Lindl.). Coniferae (Pinaceae 26; see C. for gen. char.). 2 N.W. Am. S. gigaiitea Lindl. et Gord. is the mammoth tree of Calif., discovered in the Sierra Nevada in 1850. The tallest is 320 feet, the thickest 35 feet (Sargent; cf. Eucalyptus); the age of the largest is about 1500 years. In some museums are sections of a tree cut down in 1882 a'nd show- ing 'SSS annual rings. 6". sempervirens Endl., the redwood, is even taller, though not so thick (340 and 28 feet), and is valued for its timber. &c. Seradella, serratella, Ornithopus sativus Brot. Seraphyta Fisch. et Mey. Orchidaceae (n. 6). i W.I. Serapias L. Orchidaceae (n. i). 5 Medit. Serenoa Hook. f. Palmae (i. 2). i S.E. U.S. Serial (branches, buds) one above another at same node, cf. Bud. Serianthes Benth. Leguminosae (i. i). 5 trop. As., Polynesia. Sericeous, silky. Sericocarpus Nees. Compositae (3). 5 U.S. Sericocoma Fenzl. Amarantaceae (2). 15 S. and trop. Afr. Sericocomopsis Schinz. Amarantaceae (2). 4 E. trop. Afr. Sericodes A. Gray. Zygophyllaceae. i Mexico. Sericographis Nees=Jacobinia Moric. (Acanth.) Sericorema Lopriore (Sericocoma p.p.). Amarant. (2). 2 Afr. Sericospora Nees. Acanthaceae (inc. sed.). i Antilles. Sericostachys Gilget Lopriore. Amarant. (2). 2 C. and W. Afr. Sericostoma Stocks. Boraginaceae (iv. 4). 3 E. Afr. to N.W. India. Sericotheca Rafin. (Spiraea p.p.). Rosaceae fi. i). 74 N. and C. Am. Serldia Juss. = Centaurea L. p.p. (Comp.). Series, cf. Nomenclature. SEX DISTRIBUTION 605 Seringia J. Gay. Sterculiaceae. i E. Austr., New Guinea. Serinia Rafin. (Krigia BH.}. Compositae (13). 3 S. U.S. Seriola L. = Hypochoeris L. p.p. (Comp.). Seriphium L.= Stoebe L. (Comp.). Seris Less. Compositae (12). 3 campos of S. and E. Brazil. Serissa Comm. ex Juss. Rubiaceae (n. 7). i China, Japan, cult. medicinal. Serjania Plum, ex Schum. Sapindaceae (i). 175 warm Am. Lianes with watch-spring tendrils and stip. 1. Fr. a 3-winged schizocarp. Serotinus (Lat.), late in the season. Serpicula L. Haloragidaceae. 3 trop. and subtrop. Land or marsh pi. Serrafalcus Parl. = Bromus L. p.p. (Gramin. ). Serrastylis Rolfe (Macradenia p.p.EP.). Orchid. (n. 18). i Colombia. Serrate, margin with small teeth pointing forward ; cf. Leaf, Fig. B. Serratula Dill, ex L. Compositae (i i). 40 Eur. to Japan. S. tinctorial. (saw-wort) in Brit, is dioec. Serresia Montr. Inc. sed. i New Caled. Serruria Salisb. Proteaceae (i). 50 S. Afr. Sersalisia R. Br. (Lucuma p.p.). Sapotaceae (i). 6 frop. Afr. Sertifera Lindl. Orchidaceae (n. 7). 2 Ecuador, Peru. Service berry, Amelanchier. Sesame, Sesamum indicum L. ; -grass (Am.), Tripsarum. Sesamothamnus Welw. Pedaliaceae. 5 trop. Afr. Sesamum L. Pedaliaceae. 15 trop. Afr., As. S. indicum L. largely cult, in India, &c. for the oil from seeds (gingili, sesame, &c.). Sesban (W.I.), Sesbania. Sesbania Scop. Leguminosae (in. 6). 24 trop. and subtrop. S. acu- leata Poir. is a marsh plant, giving off floating roots from the base of the stem, covered with spongy aerenchyma (cf. Neptunia). Seseli L. Umbelliferae (in. 5). 60 Eur., Afr., As., Austr.; i Brit. Sesleria Scop. Gramineae (10). 10 Eur., W. As. ; i Brit. Sesqui- (Lat. pref.), one and a half; -pedalls, i| feet. Sessea Ruiz et Pav. Solanaceae (4). 5 Andes. Sessile, without stalk. Sesuvium L. Aizoaceae (n). 5 trop. and subtrop. Halophytes. Seta, a bristle. Setaria Beauv. Gramineae (5). 30 trop. and warm temp. S. italica Beauv. (Italian millet) is cult, as a cereal in As. Setariopsis Scribnerex Millsp. (Setaria p.p.'). Gramineae (5). 2 Mexico. Setcnellanthus T. S. Brandegee. Capparidaceae (inc. sed.). i Mex. Setcreasea is.. Schum. et Sydow (Treleasea Rose). Commel. 4 N. Am. Sethia H. B. et K. = Erythroxylum P. Br. p.p. (Erythr.). Setilobus Baill. Bignoniaceae (i). i Brazil. Setiscapella Barnhart (Utricularia p.p .). Lentib. 12 trop. Setouratea Van Tiegh. (Gomphia p.p.). Ochnaceae. 8 Brazil. Seville orange, Citrus Aurantiuni L., var. Bigaradia or amara. Seven-year vine (W.I.), Ipomoea tuberosa L. Sewerzowia Regel et Schmalh. (Astragalus p.p. BH.}. Leguminosae (in. 6). i Turkestan. Sex distribution ; sta. and cpls. in one 5 fl., t/ionoclinoiis, in separate 3 and ? fls., diclinous; i and ? fls. on same pi., monoecious, on separate pi., 606 SEX DISTRIBUTION dioecious. Gynomonoecisin ( $ and ? on same pi., as in some Compo- sitae, Labiatae, &c.), gynodioecism ( $ and ? on separate pi., as in Nepeta, Thymus, and other Labiatae, Plantago,manyCaryophyllaceae, &c.), andromonoecism ( $ and 30 ft. high, much branched to ground. Shrubby trefoil, Pldea trifoliata L. Shuteria Wight et Arn. Leguminosae (in. 10). 5 trop. Afr., As. Shuttleworthia Meissn. = Verbena L. (Verben.j. Sibangea Oliv. Euphorbiaceae (A. I. i). i Gaboon. Sibara Greene (Cardamine p.p.). Cruciferae (2). 6 Calif. Sibbaldia L. (Potent ilia p.p. BH.). Rosaceae (in. 2). 8 N. temp. j*. Sibbaldiopsis Rydberg (Potentilla p.p.)- Rosaceae (in. 2). i N. Am. Siberian cedar, Finns Ceinbra L. Sibiraea Maxim. (Spiraea p.p. BH.). Rosaceae (i. i). i Siberia. Sibthorpia L. Scrophulariaceae (in. i). 6 Ear., Medit., Nepal, Andes. S. enropaea L. (Cornish money-wort) in S. England. Sicana Naud. Cucurbitaceae (3). i trop. Am., W.I. Fr. ed. Siccus (Lat.), dry, juiceless. Sickingia Willd. Rubiaceae (i. 3). 14 trop. S. Am. Some medicinal. Sicklepod (Am.), Arabis canadensis L. Sicydium Schlechtd. Cucurbitaceae (2). 6 trop. Am. Sicyomorpha Miers. Celastraceae. 2 Peru. Sicyos L. Cucurbitaceae (4). 30 trop. Am., Polynes., Austr. Sicyosperma A. Gray. Cucurbitaceae (4). i New Mexico, Texas. Sida L. Malvaceae (2). 70 cosmop. Sidalcea A. Gray. Malvaceae (2). 12 N.W. Am. Sidanoda Wooton et Standley (Anoda p.p.). Malv. (2). i New Mexico. Sidastrum E. G. Baker (Sida p.p.). Malvaceae (2). i trop. Am. Side oats, Routdoua ; - saddle flower, Sarracenia. Sideranthus Nutt. (Aplopappits Cass.). Compositae (3). 12 N. Am. Sideritis Tourn. ex L. Labiatae (vi). 45 N. temp. |*. Siderocarpus Small (Acacia p.p.). Leguminosae (i. 2). i N. Am. Sideroxylon (Dill.) L. Sapotaceae (i). 90 palaeotrop. Siebera J- Gay. Compositae (n). i W. As. Siegesbeckia L. Compositae (5). 4 trop. and warm. temp. Heads small, with invol. of =, bracts, covered with very sticky glandular hairs, aiding in distr., the whole head breaking off. Sieglingia Bernh. = Triodia R. Br. p.p. (Gramin.). Sievekingia Reichb. f. Orchidaceae (11. 13). i Costa Rica. Sieversia Willd. (Gcum p.p.). Rosaceae (in. 2). 10 N. Am. Sigmatochilus Rolfe. Orchidaceae (11. 3). r Borneo. Sigmatogyne Pntzer (Panisea p.p.). Orchidaceae (n. 3). 2 Himalaya. Sigmatosiphon Engl. Pedaliaceae. i S.W. Afr. Sigmatostalix Reichb. f. Orchidaceae (n. 19). 6 trop. S. Am. Sigmoid, S-shaped. Silaus Bernh. Umbelliferae (in. 5). 8 N. temp. ^ (i Brit.). Silene L. Caryophyllaceae (n. i). 320 N. temp., esp. Medit. (7 in Brit.). Els. of many (class F) adapted to butterflies, e.g. S. acazilis L. (moss-campion, a tufted alpine) ; others to moths, e.g. S. inflata Sm. (bladder campion), which emits scent at night. Siler Crantz. Umbelliferae (in. 7). i Eur., Siberia. Silicula, Cruc'iferae; siliqua, Cruciferae, Capparidaceae. 6o8 SILIQUAMOMUM Siliquamomum Baill. Zingiberaceae (i). i Tonquin. Silk cotton tree, Eriodendron ; -grass (W.I.), Mditlarium; -weed, Asclepias. Silkworms, plants for, Ailanthus, Lactuca, Madura, Morns. Silky bent grass, Apera; -oak (Austr.), Grevillea. Silphiosperma Steetz (Brachycome Cass.). Compositae (3). i Austr. Silphium L. Compositae (5). 13 E. U.S. S. laciniatum L. is the ' compass-plant ' of the prairies. In an exposed position its 1. turn their edges to N. and S. and avoid the mid-day radiation. (Cf. Lactuca.) Cult. orn. rl. Silvaea Phil. Portulacaceae. 4 Chili. Silver bell tree (Am.), Halesia; -berry (Am.), Elaeagnus argentais Pursh ; -fir, Abies, Ficea; -tree, Leitcadendron; -weed, Potentilla Amentia L. Silvia Allem. (Neosilvia Pax; Endiandra BH., Mezilaurus EP.). Lauraceae (n). 2 Brazil. Wood useful. Silvia Benth. Scrophulariaceae (ill. 2). 2 Mexico. Silviantnus Hook. f. Rubiaceae (i. 2). i E. Bengal. Silvorchis J. J. Smith. Orchidaceae (li. i). i Java. Silybum Vaill. ex Adans. Compositae (u). 2 Medit. S. Marianum Gaertn. (milk-thistle, Brit.) is now widely distributed over the pampas, where it was introduced. Simaba Aubl. Simarubaceae. 20 trop. S. Am. Simaruba Aubl. Simarubaceae. 6 trop. Am., W.I. Simarubaceae (EP.; BH. incl. Brunelliaceae, Cneoraceae, Koeberlini- aceae). Dicots. (Archichl. Geraniales). 28 gen., 125 sp., trop. and subtrop. Shrubs and trees with alt. pinnate or simple 1., never gland- dotted. Fls. small, reg. , ?. often oo , in axillary compound panicles or cymose spikes. K and C 3 — -y-merous. K free or more often united; C imbr. or rarely valv.; disc between sta. and ovary ring- or cup-like, sometimes enlarged into a gynophore ; A twice as many as petals, obdiplost. , often with scales at the base; G (4 — 5) or less, often free below and united by the style or stigma; ovules usu. I in each loc. as in Rutaceae. Schizocarp or caps. ; endosp. thin or none : embryo with thick cots. A few yield useful timber. Chief genera : Simaruha, Ailanthus. Simarubeae ( BH.) = Simarubaceae. Simarubopsis Engl. Simarubaceae. r Togoland. Simbuleta Forsk. (Anarrhinum Desf.). Scrophulariaceae (n. 3). 12 Medit., W.As., Abyssinia. Simethis Kunth. Liliaceae (in), i Brit., W. and S. Eur. Simuiondsia Nutt. Buxaceae. i California. Simocheilus Klotzsch. Ericaceae (iv. 2). 40 S. Afr. Simple fruit, one indivisible fr. from one rl. ; -L, with r blade. Simplicia T. Kirk. Gramineae (8). • i New Zealand. Simsia R. Br. Proteaceae (i). 5 W. Austr. Sinapis L. (Brassica p.p. BH.). Cruciferae (2). 5 Medit., Eur. S. ari'ensis L. (charlock) is an abundant weed of cult. ; our cornfields are yellow with it in summer. 6". alba L. is the white mustard. Sinapodendron Lowe (Brassica p.p. BH.). Crucif. (2). 5 Madeira, &c. Sincoraea Ule. Bromeliaceae (4). i Bahia. SIUM 609 Sindechites Oliv. Apocynaceae (n. i). j S.W. China. Sindora Miq. Leguminosae (n. 2). 6 Malaya. Sinensis (Lat.), Chinese. Singana Aubl. Leguminosae (inc. sed.). i Guiana. Sinistrorse, turned to the left. Sinningia Nees. Gesneriaceae (n). 20 Brazil. S. speciosa Hiern, &c. cult, (generally known as Gloxinias). Tuberous plants. Usual pro- pagation by planting 1. on the soil ; from the base of the petiole a new pi. arises by budding (cf. Begonia, Streptocarpus). Sinofranchetia Hemsl. (Parvatia p p.). Lardizabalaceae. i China. Sinomenium Diels (Cocculus p.p.). Menispermaceae. i E. As. Sinowilsonia Hemsl. Hamamelidaceae. i China. Siuuate (1.), with teeth and notches rounded (Leaf, fig. D). Sinus, bay, re-entrant angle. Siolmatra Baill. Cucurbitaceae (i). 2 Brazil, Paraguay. Sipanea Aubl. Rubiaceae (i. 3). 3 S. Am. Siparuna Aubl Monimiaceae. 90 trop. Am. Siphanthera Pohl. Melastomaceae (i). 12 Brazil, Guiana. Siphoboea Baill. Gesneriaceae (i). i Phil. Is. Siphocampylus Pohl. Campanulaceae (in). 1 10 trop. Am., W.I. Siphocodon Turcz. Campanulaceae (i). i S. Afr. Siphocolea Baill. Bignoniaceae (4). 3 Madag. Siphonandrium K. Schum. Rubiaceae (i. 7). i New Guinea. Siphonanthus L. = Clerodendron L. p.p. (Verben.). Siph.on.ella A. A. Heller (Gilia p.p.)- Polemoniaceae. i W. N. Am. Siphonella Small (Fedia p.p.). Valerianaceae. 2 N. Am. Siphoaia Kich. = Hevea Aubl. 6\ elastica Pers. = H. guianensis. Siphonidium Armstr. Scrophulariaceae (in. 3). i New Zealand. Siphonochilus Wood et Franks. Zingiberaceae (i). i Natal. Siphonodon Griff. Celastraceae. 2 Malaya, Austr. Siphonogamy, fertilisation by pollen tube. Siphonoglossa Oerst. Acanthaceae (iv. B) 3 Am. Siphonostegia Benth. Scrophuln'riaceae (in. 3). 3 As. Siphonostelma Schlechter. Asclepiadaceae (n. 3). i S.W. Afr. Siphonychia Torr. et A. Gray. Caryophyllaceae (I. 4). i Atl. N. Am. Sipolisia Glaziou. Compositae (i). i Minas Geraes. Siris, Albizzia Lebbek Benth. Sirium L. = Santalum L. (Santal.). Sisal hemp, Agave sisalana Perrine. Sison L. Umbelliferae (in. 5). 2 Eur., incl. Brit., Medit. Sissoo, Dalbergia Sissoo Roxb. Sisymbrium (Tourn.) L. (BH. incl. Alliaria). Cruciferae (2). 50 N. temp. \% ; 3 Brit., incl. S. officinale Scop, (hedge-mustard). Sisyndite E. Mey. Zygophyllaceae. i S. Afr. Sisyranthus E. Mey. Asclepiadaceae (11. 3). 8 S. Afr. SisyrincMum L. Iridaceae (n). 75 Am. Cult. orn. fl. Sisyrolepis Radlk. Sapindaceae (inc. sed.). i Siam. Sitanion Rafin. (Elymus L. p.p.). Gramineae (12). 12 N. Am. Sitilias Rafin. (Pyrrhopappns DC.). Compositae (13). 6 U.S., Mexico. Sitka cypress, Chamaecyparis mttkaensis Lindl. et Gord. Slum (Tourn.) L. Umbelliferae (ill. 5). to N- temp., trop. Afr. w. 39 610 SIUM i in Brit, (water-parsnip). S. Sisarum L. (skirret) cult, for tuberous roots. Skimmia Thunb. Rutaceae (iv). i Himal. to Japan, 5". japonica Thunb., often cult, for its handsome foliage and red berries. Skirret, Sinm Sisarum L. Skolemora Arruda. Inc. sed. i Brazil. Skull cap, Scittellaria. Skunk cabbage, Symplocarpus foetidus Nutt. Skytanthus Meyen. Apocynaceae (i. 3). 3 Brazil, Chili. Slackia Griff. Gesneriaceae (i). i Burma. Sladenia Kurz. Dilleniaceae (Ternstr. Bff.}. i Yunnan. Sleep movements, cf. Movement. Sloanea L. Elaeocarpaceae. 45 trop. Sloe, Prunus spinosa L. Sloetia Teijsm. et Binn. Moraceae (i). 2 Malaya. Sloetiopsis Engl. Moraceae (i). i Usambara. Small reed, Calatnagrostis. Smallia Nieuwland = Triorchos Small et Nash (Orchid.). Smartweed (Am.), Polygonum. Smeathmannia (Soland.) R. Br. (Paropsia p.p.). Flacourtiaceae (6) (Passiflor. BH.}. 6 trop. Afr. Smelophyllum Radlk. Sapindaceae (i). i S. Afr. Smelowskia C. A. Mey. Cruciferaej4). 8 temp. As., Pac. N. Am. Smilacina Desf. Liliaceae (vii). 20 Himalaya to C. Am. Smilax (Tourn.) L. Liliaceae (xi). 210 trop. and subtrop. Most are climbing shrubs with net-veined 1. At base of 1. spring two tendrils, one on either side, usu. regarded as modified stip., though these organs scarcely occur in Monocots. Stems often furnished with recurved hooks which aid in climbing. Fls. dioec., in umbels. The dried roots of several S. Am. sp. form sarsaparilla. Smirnowia Bunge. Leguminosae (ill. 6). i Turkestan. Smithia Ait. Leguminosae (in. 7). 35 trop. As., Afr. Smithiantha O. Ktze. (Naegelia Regel). Gesneriaceae (n). 6 Mexico. Cult. Smodingium E. Mey. Anacardiaceae (3). i S. Afr. Smyrniopsis Boiss. (Smyrninm p.p. BH.}. Umbellif. (in. 4). 3 E. Medit. Smyrnium (Tourn.) L. Umbelliferae (in. 4). 7 Medit., Eur., Brit. S. Olusatrum L. (alexanders), formerly used like celery. Smythea Seem. Rhamnaceae. 4 Malay Archipelago, Polynesia. Snail flower, Arisaema. Snake gourd, Trichosanthes angnina L. ; -head (Am.), Ckelone; -'& head, Fritillaria\ -root, Cimiciftiga; black-- (Am.), Sanicula; button --(Am.), Liatris; Senega--, Polygala; -wood, Ophioxylon, Ophiocaryon. Snapdragon, Antirrhinum majus L., (W.I.) Rnellia. Sneeze wood, Pteroxylon, (Am.) Heleninm ; -wort, Achillea. Snow ball tree, I'iburmim Opitlus L. ; -berry, Symphoricarpus race- mosus Michx. ; -drop, Galanthns nivalis L. ; -drop tree, Ckionantktts, Halesia; -flake, Leucojnm. Soap, Cklorogvlum, Liliaceae, Limonia, Sapindits, Saponaria; -berry SOLANACEAE 6n tree (W.I.), Sapindits; -tree, Quilla/a; -wood (W.I.), Ctethra', -wort, Saponaria. Soaresia Sch.-Bip. Compositae (i). i campos of S. Brazil. Sobole, a shoot from the ground. Sobolewskia Marsch.-Bieb. Cruciferae (2). 3 W. As. Sobralia Ruiz et Pav. Orchidaceae (n. 7). 33 Peru to Mexico. Social habit, forming homogeneous forests, pine, birch, beech. Societies, Plant, see Plant societies. Socotora Balf. f. (Periploca p.p. EP.}. Asclepiadaceae (i). i Socotra. Socotranthus O. Ktze. =Cochlanthus Balf. f. (Asclep.). Socratea Karst. (Iriartea p.p. EP.}. Palmae (IV. i). i Brazil. Sodiroa Andre. Bromeliaceae (i). 3 Colombia, Ecuador. Soemmeringla Mart. Leguminosae (in. 7). i N.E. Brazil. Soft grass, Holcus. Sohnreyia Krause. Rutaceae (i). i Manaos. Soja bean, Glycine. Soja Moench = Glycine L. p.p. (Legum.). Sola, Acschynoinene aspera L. Solanaceae (EP., BH. incl. Nolanaceae). Dicots. (Sympet. Tubiflorae; Polemoniales BH.}. 72 gen., 1500 sp. trop. and temp.; chief centre C. and S. Am., where there are 36 local gen.; in Eur. and As. only § 2 is repres. Herbs, shrubs or small trees; 1. in the non-flowering part usu. alt., but in the infl. -portion alt. or in pairs ; the arrangement in pairs is due to the mode of branching and adnation as illustrated in the figure. In Datura the branching is dichasial, and the bracts are adnate to their axillary shoots up to the point at which the next Branching in Solanaceae (after Eichler); Datura Stramonium (left) and Atropa Belladonna (right), i, 2, 3, firs, or infls. of successive orders; 6, bract of i, a |3 bracts of 2, and so on. branches arise, so that a looks like the bracteole of 2, rather than its bract. In Atropa the branching is cincinnal, one of the two branches at a node remaining undeveloped, and the bract is again adnate to its axillary branch. Of the pair of 1. thus found at any node, one is usu. smaller than the other. In Solanum, &c. further complications occur (see Eichler's Bliithendiag}. Fls. sol. or in cymes, ?, sometimes •]• . K (5), persistent; C (5), of various forms, rarely 2-lipped, usually folded and conv.; A 5, alt. 39—2 6i2 SOLANACEAE with petals, epipet., or fewer in -|- fls. , often opening by pores; G (2), obliquely placed in the f). (the post. cpl. to the right, the ant. to the left, when shown in a floral diagram), 2-loc., sometimes with secondary divisions (e.g. Datura), upon a hypog. disc; ov. i — oo in each loc. , anatr. or slightly amphitr., on axile plac. (most often the plac. are swollen and the ov. numerous) ; style simple, with 2-lobed stigma. Berry or caps. Embryo curved or straight, in endosp. Fls. conspic., insect-visited ; some, e.g, Nicotiana, suited to Lepidoptera. A few are economically important, e.g. Solanum (potato), Nicotiana (tobacco), Lycopersicum, Capsicum, &c. ; Datura, Atropa, &c. are medicinal; several are favourites in horticulture. Classification and chief genera (after von Wettstein) : the S. are nearly related to Scrophulariaceae, the most general distinction being the oblique ovary : this however is by no means easily made out, and the zygomorphism of the fl. is most often used as a distinction. Cer- tain genera of S. are nearly related to various Boraginaceae, Gesne- riaceae, Nolanaceae, &c., and it is possible that the S. are not really a simple monophyletic family ; they occupy a middle place between the Tubuliflorae with actinom. and those with zygom. fls. A. Embryo clearly curved, through more than a semicircle. All 5 sta. fertile, equal or only slightly different in length. i. Nicandreae (ovary 3 — 5-loc., the walls of the loc. dividing the placentae irregularly) : Nicandra (only genus). i. Solaneae (ovary 2-loc.) : Lycium, Atropa, Hyoscyamus, Phy- salis, Capsicum, Solanum, Lycopersicum, Mandragora. 3. Datureae (ovary 4-loc., the walls dividing the placentae equally) : Datura, Solandra (only genera). B. Embryo straight or slightly curved (less than a semicircle). 4. Cestreae (all 5 sta. fertile) ; Cestrum, Nicotiana, Petunia. 5. Salpiglossideae (2 or 4 sta. fertile, of different lengths): Salpi- glossis, Schizanthus. Solandra L. = Hydrocotyle Tourn. (BH.} - Centella L. p.p. (Umb.). Solandra Sw. Solanaceae (3). 6 trop. Am. Solanopsis Borner (Solanum p.p.). Solanaceae (2). 2 Am. Solanum (Tourn.) L. (incl. Lycopersicum Hill). Solanaceae (2). 1225 trop. and temp. .5". Dulcamara L. (bittersweet, nightshade) and S. nigrum L. in Brit. The fls. are small, with a cone of anthers opening at the tip as in Borago. S. tuberosum L. (S. Am.) is the potato. From the axils of the lowest 1. there spring branches which grow horiz. underground and swell up at the ends into tubers (potatoes). That these are stem structures is shown by their origin and by their possession of buds — the 'eyes.' Each eye is a small bud in the axil of an aborted 1. (repres. by a semicircular rim). When the parent plant dies down in autumn the tubers become detached, and in the next season they form new plants by the development of the eyes, at the expense of the starch and other reserves stored in the tuber. By heaping earth against the stem, so as to cover more of the leaf-axils, more of the axillary shoots are made to become tuber-bearing ; hence the value of ridging potatoes. S. Lycopersicum L. (Am.) is the tomato, cult, for ed. fr. S. Melongena L., the egg-fruit, is cult, in warm countries for ed. fr. SONNERA TIA CEAE 6 1 3 Solaria Phil. Liliaceae (iv). i Chili. Soldanella L. Primulaceae. 5 Alps of Eur. The fls. expand at very low temperatures, often coming up through the snow ; they have a mechanism like that of Erica. Soldier plant (W.I.). Calliandra. Solea Spreng. = lonidium Vent. (BH.} = Hybanthus Jacq. Solena Willd. = Posoqueria Aubl. (Rubi.). Solenandra Hook. f. Rubiaceae (I. 5). i Cuba. Solenanthus Ledeb. Boraginaceae (iv. i). 15 Medit., C. As. Solenidium Lindl. Orchidaceae (n. 19). i Colombia. Solenixora Bail!. Rubiaceae (n. 4). i Madag. Solenocarpus Wight et Arn. Anacardiaceae (2). i India. Solenocentrum Schlechter. Orchidaceae (n. 3). i Costa Rica. Solenomelus Miers. Iridaceae (ll). 2 Chili. Solenophora Benth. Gesneriaceae (n). 2 Mexico, C. Am. Solenoruellia Baill. Acanthaceae (iv. B). i Mexico. Solenospermum Zoll. Celastraceae. i Java. Solenostemma Hayne. Asclepiadaceae (n. i). i Egypt, Arabia. Solenostemon Thonn. (Plectranthtis p.p. BH.}. Labi. (vn). 8 W. Afr. Solenosterigma Klotzsch ex K. Krause— Philodendron Schott. Solenostigma Endl. = Celtis Toiun. p.p. (Ulm.). Solenostyles Host. Acanthaceae (inc. sed.). i, habitat?. Solenotus Stev. = Astragalus Tourn. (Legum.). Solfia Rechinger. Palmae (iv. i). i Samoa. Solidago (Vaill.) L. Compositae (3). 90 Am.; i in Eur. (incl. Brit.), S. Virgaurea L., the golden rod. Soliera Clos (Kurzamra EP.). Labiatae (vi). i Chili. Solitary (fl.), one per axil. Soliva Ruiz et Pav. Compositae (7). 6 Am., Austr. Sollya Lindl. Pittosporaceae. 2 W. Austr. Twiners. Solmsia Baill. Gonystilaceae. 2 New Caledonia. Solms-Laubachia Muschler. Cruciferae (4). i China. Solomon's seal, Polygonatum. Somalia Oliv. (Barleria p.p. EP}. Acanthaceae (iv. B). i trop. Afr. Sommera Schlechtend. Rubiaceae (i. 7). 2 Amazon valley, Mexico. Sommerfeltia Less. Compositae (3). i S. Andes. Char. pi. Sommieria Becc. Palmae (iv. i). 2 New Guinea. Somphoxylon Eichl. Menispermaceae. i Guiana. Sonchus (Tourn.) L. Compositae (13). 45 %, 3 Brit, (sow-thistle). Sondaria Dennst. Rhamnaceae (inc. sed.). i Indomal. Sonerila Roxb. (Cassebeeria Dennst.). Melastom. (i). 70 warm As. Sonnea Greene. Boraginaceae (iv. 2). 6 Pac. N. Am. Sonneratia L. f. (Blatti Adans.). Sonneratiaceae. 6 Indomal. Man- groves (q-v.), with the general habit of Rhizophoraceae. Aerial roots spring vertically out of the mud, arising as lat., negatively geotropic branches upon the ordinary roots ; they are provided with aerenchyma, and appear to be respiratory organs. Sonneratiaceae (Blattiaceae ; EP.\ Lythraceae p.p. BH.). Dicots. (Archichl. Myrtiflorae). 3 gen., 12 sp. trop. As. and Afr. Shrubs and trees with opp. entire exstip. 1. and conspic. fls., heterochlam. orapet., 5 or c? ? , reg. K 4-8, C 4-8 or o, A oo , G (4-15), with one style and 6 1 4 SONNERA TIA CEAE multiloc. ov. with oo ovules. Caps, or berry ; oo seeds, exalbum. Chief gen. Sonneratia, Duabanga. Sonzaya Marchand (Canarium p.p. EP.). Burseraceae. i Austr. Sophia L. 1735 (Sisymbrium p.p.)- Cruciferae (2). 20 N. Am. Sophia L. i775 = Bombax L. p.p. (Bomb.). Sophoclesia Klotzsch. Ericaceae (in. 2). 10 Andes, Guiana, W.I. Sophora L. Leguminosae (in. i). 25 trop. and warm temp. Winter- buds naked. The wood is very hard. Sophrocattleya, Sophrolaelia, Sophrolaeliocattleya, Sophrocattlaelia x Hort. Orchidaceae. Hybrids of Sophronitis. Sophronanthe Benth. = Gratiola Rupp. (Scroph.). Sophronitis Lindl. Orchidaceae (n. 6). 4 S.E.Brazil. Cult. orn. fl. Sopubia Buch. -Ham. Scrophulariaceae (in. 2). 20 palaeotrop. Sorbaria A. Br. (Spiraea p.p. BH.}. Rosaceae (i. i). 5 N. As., N. Am. Sorbaronia x C. K. Schneider. Rosaceae. Hybrid Sorbus-Aronia. Sorbopyrus x C. K. Schneider. Rosaceae. Hybrid Sorbus-Pyrus. Sorbus (Tourn.) L. = Pyrus Tourn. p.p. (Ros.). Sordidus (Lat.), dirty white. Sorghastrum Nash (Andropogon p.p.). Gramineae (2). 4 N. Am. Sorgho, Sorghum vulgare Pers. Sorghum L. (Andropogon, p.p.). Gramineae (2). 13 trop. and sub- trop. .S. vulgare Pers. (millet or guinea corn), largely cult, in Medit. &c. as a cereal. From the haulm of the var. saccharatum Koern.-sugar is sometimes prepared. Soriferous, bearing sori. Sorindela Thou. Anacardiaceae (3). 10 trop. Aft., Madag. Sorocea A. St Hil. Moraceae (n). 12 trop. Am. Sorocephalus R. Br. Proteaceae (i). 10 S. Afr. Sorosis, a fleshy multiple fruit, Ananas, &c. Sorrel, Rumex ; wood -, Oxalis acetosella L. Sorus, a group of sporangia, Cycadaceae, Filicales, Pteridophyta. Souari nut, Caryocar. Souchong, Thea. Soulamea Lam. Simarubaceae. i Moluccas to Fiji. Soulangia Brongn. = Phylica L. (Rhamn.). Souleyetia Gaudich. (Pandanus p. p. EP.}. Pandanaceae. i, habitat?. Souliea F ranch. Ranunculaceae (2). i China. Sour grass, Paniann conjugation Berg.; - sop, Anona muricata L. Souroubea Aubl. (Ruyschia BH. ). Marcgraviaceae. 6 trop. Am. Southern cold zone, see Zones of Veg. Southernwood, Artemisia Abrotanum L. Sow-bread, Cyclamen ; -thistle, Sonchus, Sowerbaea Smith. Liliaceae (ill). 4 Austr. Soy bean, Glycine Soja Sieb. et Zucc., G. hispida Maxim. Soyauxia Oliv. Flacourtiaceae (6) (Passiflor. £ff.). 5 W. trop. Afr. Soyeria Monn. = Crepis Vaill. p.p. (Coinp.). Soymida A. Juss. Meliaceae (n). i Indomal. Astringent bark. Wood. Spachea A. Juss. Malpighiaceae (11). 10 W.I., trop. S. Am. Spadiceus (Lat.), chestnut-coloured, or bearing a spadix. Spadiciflorae (Warming). The 3rd order of Monocots. SPATHODEA 615 Spadix, a spike with fls. ± sunk in tissue, enclosed in a large 1. or spathe, Araceae, Cyclanthaceae, Palmae, Zostera. Spananthe Juss. Umbelliferae (i. 2). i trap. Am. Spanish bayonet, Yucca ; - berries, Rhanmiis infectoria L. ; - broom, Spartiumjunceum\j.\ -dagger, Yucca ; -liquorice, Glycyrrhiza gla- bra L. ; - needle, Bideiis ; - plum, Spondias. Spanoghea Blume. Sapindaceae (i). 2 Malay Archipelago. Sparattanthelium Mart. Hernandiaceae. 5 trop. Am. Sparattosperma Mart. Bignoniaceae (2). 3 Brazil. Sparattosyce Bur. Moraceae (n). 2 New Caledonia. Sparaxis Ker. Iridaceae (ill). 6 Cape Colony. Cult. orn. fl. Sparganiaceae (EP. ; Typhaceae p.p. BH.}. Monocots. (Pandanales). Only gen. Sparganium (q.v.}. Sparganium (Tourn.) L. Sparganiaceae. 15 N. temp., Austr. , N.Z. ; 3 in Brit, (bur-weed), in ponds. Creeping rhiz. and stem projecting above water with the 1. and fls. These in spherical heads, the i usu. higher up than the ?. P 3 — 6, scaly, sepaloid ; Zf.) = Lindernia All. Tium Medic. (Astragalus p.p.). Leguminosae (ill. 6). 10 N.Am. Tmesipteris Bernh. Psilotaceae. i Austr., N.Z., Polynes., T. tan- nensis Bernh. It grows as an epiphyte (? parasite) on the trunks of tree ferns. The rhizome bears large lanceolate green 1. Toadflax, Linaria; bastard--, Thesimn. Tobacco, Nicotiana. Tococa Aubl. Melastomaceae (i). 40 trop. S. Am. Tocoyena Aubl. Rubiaceae (i. 8). 8 S. Am., Cuba. Todaroa Parl. Umbelliferae (ill. 5). 2 Teneriffe. Toddalia Ju.ss. Rutaceae (iv). i trop. As., Afr., Madag. Toddaliopsis Engl. Rutaceae (iv). i Zanzibar. Toddy, Borassus, Caryota, Cocos, &c.: - palm, Caryota urens L. Todea Willd. Osmundaceae. i S. Afr., Austr., T. africana Willd. (T. barbara Moore); apogamous (see Filicales). ToecMma Radlk. Sapindaceae (i). 6 Austr., New Guinea. Tofieldia Huds. Liliaceae (i). 15 N. temp., Andes. T. palustris Huds. (Scottish asphodel) in Brit. 3-lobed invol. (calyculus) beneath the K. Toltaonia O. Ktze. Compositae (3). i Annam. Tolmiea Torr. et Gray. Saxifragaceae (i). i Pac. N. Am., T. Men- ziesii Torr. et Gray. Adv. buds on upper part of petiole. Axial cup split down ant. side. Pets, thread-like; only 3 post. sta. occur. Tolpis Adans. Compositae (13). 15 Azores, Canaries, Medit. Tolu, balsam of, Myroxylon toluiferum H. B. et K. Toluifera L. (Myroxyloii Forst., q.v.}. Legum. (ill. i). 6 trop. S. Am. Tomato, Solanwn Lycopersicum L.; gooseberry -, strawberry -, Phy- salis ; tree -, Cyphomandra. Tomentose, with cottony felt of hairs. Tomex Thunb. = Litsea Lam. (Laur.). Tonalantrius T. S. Brandegee. Compositae (5). i Mexico. 654 TOND1N Tondin G. W. Schilling. Inc. sed. i Guiana. Tonduzia Pittier (Ramvoljia p.p.)- Apocynaceae (i. 3). 2 C. Am. Tonella Nutt. ex A. Gray. Scrophulariaceae (II. 4). 2 W. U.S. Tonestus A. Nelson (Aplopappus p.p.)- Compositae (3). 3 N. Am. Tonina Aubl. Eriocaulonaceae. i trop. S. Am., W.I. Tonka or Tonctuin bean, Dipteryx odorata Willd. Tontelea Aubl. = Salacia L. (Hippocrat.). Tooart, towart (Austr.), Eucalyptus gomphocephala DC. Toon, Cedrela Toona Roxb. Toona M. Roem. (Cedrela p.p. £//.). Meliaceae (i). 8 As., Austr. Toothache grass (Am.), Gtenium; -tree, Zanthoxylum; -weed, Spi- lanthes; -wort (Am.), Dentaria. Toothwort, Lathraea squamaria L. Topee tampo, Calathea Alloida Lindl. Topiary, formal ornamental gardening. Topobea Aubl. Melastomaceae (i). 20 warm Am. Ed. fr. Topographical agents in distr., mountains, rivers, seas, soil, &c. Tordylium Tourn. ex L. Umbelliferae (in. 6). 16 Eur. (i Brit.), N. Afr., As. Torenia L. Scrophulariaceae (n. 6). 22 trop. Torgesia Bornmtiller. Gramineae (8). i Palestine. Toricellia DC. Cornaceae. 2 Himalaya, China. Torilis Adans. (Caucalis BH.}. Umbellif. (in. 2). 2} Medit , N. As., Afr. Tormentil, Potentilla Torment ilia Neck. Tormentilla (Tourn.) L. =Potentilla L. (Ros.). Tornabenea Parl. ex Webb. Umbelliferae (in. 7). 3 Cape Verde Is. Tornelia Gutierrez = Monstera Adans. (Arac.). Torralbasia Krug. et Urb. (Enonymus p.p.). Celastraceae. r W.I. Torresia Allem. (Awburana Schwacke et Taub.). Legum. (in. i). i Braz. TorreyaArn. Coniferae(Taxaceae 8; see C. for gen. char.). 4 N. Am., China, Japan. Like Taxus. The timber is useful. Torricellia DC. Cornaceae. 2 Himalaya, China. Torrubia Veil. (Piscina Plum.). Nyctaginaceae. 9 Am., W.I. Torulinium Desv. (Mariscus BH. , Cyperus\>.\>. EP.). Cyperaceae (i). 6 W.I., Colombia to Chili. Torulose, cylindrical with slight contractions. Torus, floral receptacle, q.v. Total parasites, those that take all food from the host. Touch me not, Impatiens. Touchardia Gaudich. Urticaceae (3). i Hawaiian Is. Touchiroa Aubl. (Crudia Schreb.). Legum. (11. 3). 2 Borneo, Burma. Toulicia Aubl. Sapindaceae (i). 10 trop. S. Am. Touloucouna, Caraipa. Tounatea Aubl. (Swartzia Schreb.). Leguminosae (11. 9). 60 trop. Am., Afr. Tournefortia L. Boraginaceae (m). 120 trop. and subtrop. Trees and shrubs. Tournefortiopsis Rusby. Rubiaceae (11. 2). i Bolivia. Tournesol, Chrozophora tinctoria A. Juss. TRA CH YSPERMUM 6 5 5 Tourneuxia Coss. Compositae (13). i Algeria. Tournonia Moq. Basellaceae. i Colombia. Touroulia Aubl. Quiinaceae. 3 Guiana, Brazil. Tourrettia Fougeroux. Bignoniaceae (5). i Peru to Mexico. Tous les mois, ^Caniia edulis Ker-Gawl. Touteria Eaton et Wright (Mentzelia p.p.). Loasaceae. 17 N. Am. Tovaria Neck. = SmiIacina Desf. (Lili.). Tovaria Ruiz et Pav. Tovariaceae. 2 W.I., S. Am. Herbs with term, racemes. Fl. ? , reg. K, C, A, 8, G (6 — 8) with plac. reaching to centre, and oo ovules. Berry. Endosp. Tovariaceae (EP.,Capparidaceae p. p. BH.]. Dicots. (Arcbichl. Rhoea- dales). Only genus Tovaria, q.v. Tovomita Aubl. Guttiferae (v). 30 trop. Am. Tovomitopsis Planch, et Triana ( Chrysuchlamys Bff.}. Guttiferae (v). 10 trop. Am. Tow, Linum usitatissiiniiui L. Towel gourd, Luffa. Townsendia Hook. Compositae (3). 18 Rocky Mts. Townsoaia Cheeseman. Orchidaceae (n. 2). i New Zealand. Toxanthera Hook. f. Cucurbitaceae (2). 2 S. and trop. Afr. Toxanthes Turcz. Compositae (4). 2 S. and W. Austr. Toxicodendron (Tourn.) L. (Kfnts p.p.). Anacard. (3). 35 N. Am. Toxicodendrum Thunb. Euphorbiaceae (A I. i). i Cape Colony. Toxicophlaea Harv. = Acokanthera G. Don (Apocyn.). Toxicoscordion Rydberg (Zygadenns p.p.). Liliaceae (i). 10 N. Am. Toxocarpus Wight et Am. (Secamone EP.}. Asclepiadaceae (u. i). 22 palaeotrop. Tozzettia Savi = Alopecurus L. (Gram.). Tozzia L. Scrophulariaceae (in. 3). i Alps, i Carpathians. Semi- parasites, with loose-pollen fls. (see fam.). Trabeculae, Isoetes. Tracaulon Rafin. (Polygonum p.p.). Polygon, (n. 2). 20 Am., As., Austr. Tracheliopsis Buser (Campanula p.p.). Campan. (i. i). 3 Medit. Trachelium Tourn. ex L. Campanul. (i). 7 Medit. Cult. orn. fl. Trachelospermum Lem. Apocynaceae (n. i). 8 India to Japan. Trachyandra Kunth = Anthericum L. p.p. (Lili.). Trachycarpus H. Wendl. Palmae (i. 2). 5 E. As. Trachydium Lindl. Umbelliferae (ill. 4). 12 E. Afr. to China. Trachylorjium Hayne. Leguminosae (n. 3). 3 trop As., E. Afr. These yield copal, which is dug up from the soil near the roots or in a half- fossilised condition from places where trees once existed. Trachymene DC. = Platysace Bunge (Umbell.). Trachyinene Rudge. Umbelliferae (i. i). 14 Austr. to Borneo. Trachynotia Michx. -Spartina Schreb. (Gram.). Trachyphrynium Benth. Marantaceae. 6 trop. Afr. Trachypogon Nees. Gramineae (2). 2 Am., S. Afr., Madag. Trachypteris Andree. Polypodinceae. i trop. Am. Trachys Pers. Gramineae (3). i coast of India. Trachyspermum Link (Canon p.p. £H.). Umbellif. (in. 6). 12 Afr., S. As. 656 TRA CHYSTEMON Trachystemon D. Don. Boraginaceae (iv. 3). 2 Medit. Trachystigma C. B. Clarke (Didymocarpus p.p. EP.). Gesneriaceae (i). i trop. Afr. Tracyanthus Small (Zygademts p.p.)- Liliaceae (8). i N. Am. Tradescantella Small (Tradescantia p.p.). Commelin. 2 N. Am. Tradescantia Rupp. ex L. Commelinaceae. 35 trop. and N. Am. T. virginiana L. (spider-wort), &c. Cult. orn. fl. 6 perfect sta. covered with hairs. Protandr. Infl. a cincinnus. Tragacanth, Astragalus, esp. gummifer Labill. Tragacantha Tourn. ex L. = Astragalus Tourn. (Legum.). Traganthes Wallr. (Eupatornnn p.p.). Compositae (2). 5 N. Am. Traganum Delile. Chenopodiaceae (B). i Meclit. Tragia Plum, ex L. Euphorbiaceae (A. n. 2). 55 trop., subtrop. Tragium Sprang. = Pimpinella Riv. p.p. (Umbell.). Tragoceros H. B. et K. Compositae (5). 4 Mexico. The C of ? fl. becomes rigid aftei fert., and forms a double hook upon the fr. Tragopogon (Tourn.) L. Compositae (13). 35 N. temp. |* (T. pra- tensis L., goat's beard, in Brit.). The fl. -heads of the Brit. sp. close at midday, whence its name of ' John-go-to-bed-at-noon.' T. porri- foluis L. (salsify) sometimes grown as a vegetable. Tragopyrum Bieb. = Atraphaxis L. (Polypod.). Tragoselinum Tourn. ex Hall. = Pimpinella Riv. p.p. (Umb.). Tragus Hall. Gramineae (3). 2 trop. and subtrop. Trailing arbutus, Epigaea repens L. ; - azalea, Loisdetiria. Trailliaedoxa W. W. Smith et Forrest. Rubi. (inc. sed.). i W. China. Tralliana Lour. Inc. sed. i Cochinchina. Translator, Asclepiadaceac. Transpiration, evaporation of water from plant. Trap flowers, fls. which entrap visitors, allowing them to escape by another road, or at another time, past the essential organs, Aristo- lochia. Arum, Asannn, Ceropegia, Coryanthes, Cypnpedium, Mag- nolia. Trapa L. Onagraceae (i), sometimes in a special fam. Hydrocaryaceae. 3 \$fc (horn-nut), incl. 7'. natans L. Water pi. with floating 1. and submerged adv. roots which contain chlorophyll and assimilate. Fl. 5 , 4-merous, perig., with a disc, above the sta. G 2-loc. , with one anatr. pend. ov. in each; raphe ventral. Seed large, exalbum. , in a horned nut; used as food in China, &c. Trapella Oliv. Pedaliaceae. 2 China, Japan. Trapeziform, of unsymmetrical 4-sided shape. Trasus S. F. Gray = Carex Dill. p.p. (Cyper.). Trattinickia Willd. Burseraceae. i trop. S. Am. Traunia K. Schum. Asclepiadaceae (n. 3). i Kilimandjaro. Trautvetteria Fisch. et Mey. Ranunculaceae (3). i Japan, N. Am. Traveller's joy, Clematis; -tree, Ravenala. Treacle mustard, Erysimum. Tread softly (A.m.),Jatropha. Trechonaetes Miers. Solanaceae (2). i Chili. Treculia Decne. ex Tree. Moraceae (n). 6 W. trop. Afr. The seeds of T. africana Decne. (okvva) are ground into meal. Tree, a woody plant usu. with only one =.tem at base, and > 30 feet high ; TRIAS 657 - fern, Alsophila, Brained, Cyathea, Dicksonia, Heinitelia, &c.; - of heaven, Atlant/ins; -tomato, Cyphomandra. Trefoil, Tri/alinm ; bird's foot-, Lotus; shrubby-, Ptelea, Treichelia Vatke. Campanulaceae (i). i S. Afr. Treleasea Rose (Setcreasea EP.}. Commelinaceae. 5 N. Am. Trema Lour. Ulmaceae. 30 trop. Tremacanthus Sp. Moore. Acanthaceae (iv. A), i Matto Grosso. Tremandra R. Br. Tremandraceae. 2 W. Austr. Tremandraceae (EP., BH.}. Uicots. (Archichl. Gerauiales; Polygalmae BH.). 2 gen., 26 .sp. Austr. Herbs with whorled, alt. oropp., exstip. 1. Fls. dichlam., reg. K 4 — -5, rarely (4 — 5), valvate; C 4 — 5, val- vate; A 8, 10, or rarely 6; G (2), medianly placed; style and stigma simple; ovules i or 2 in each loc., anatr. Caps., loculic. or also septic. Album, seed, with or without aril. Genera: Tetratheca, Tremandra. Tremandreae (BH.} = Tremandraceae. Tremanthera F. Muell. Theaceae. i New Guinea. Trematolobelia Zahlbr ex Rock (Lobelia p.p.). Campanulaceae (in), t Hawaiian Is. Trematosperma Urb. Icacinaceae. i Somaliland. Trembleya DC. Melastomaceae (i). 14 S. Brazil. Trentepohlia Roth. = Heliophila Burm. f. (Crucif.). Trepocarpus Nutt. ex DC. Umbelliferae (in. 5). i S. U.S. TresantheraKarst. (Rustia BH.}. Rubiaceae (i. i). 2 Venezuela, W.I. Tretocarya Maxim. Boraginaceae (iv. 2). 2 China, Himalaya. Treubella i'ierre (Palaquiiun p.p.). Sapotaceae (i). i Sumatra. Treubia Pierre (Lophopyxis EP.}. Icacinaceae. i Ceram. Treutlera Hook. f. Asclepiadaceae (n. 3). i Sikkim. Trevesia Vis. Araliaceae (i). 4 Indomal. Trevirana Willd. = Achimenes P. Br. p.p. (Gesn.). Trevoa Miers. Rhamnaceae. 3 Andes. Trevoria F. C. Lehmann. Orchidaceae (n. 13). 2 Ecuador, Colombia. Trewia L. Euphorbiaceae (A. 11.2). 2 Indomal. Tri- (Gr. and Lat. pref), three; -adelphous, -androus, -carpellary, -chotomous (branching into 3), -cyclic (with 3 whorls), -farious (facing 3 ways), -fid, -foliate, (with 3 leaflets), -gonous (3-angled), -lobous, -locular, -merous (parts in 35), -morphism (with 3 forms, LytJiruni, &c.), -oecism ( 5 , 6 sp. trop. Herbs of rapid water in hill streams (cf. Podostemaceae) with creeping thread-like roots giving off (exc. Lawia, where the primary axis is flattened into a thallus and gives off) large numbers of secondary shoots with minute delicate simple exstip. 1. P 3—5 or (3—5)» reg-> sepaloid, A 3, 5, co , or i, G (2—3) 2— 3-loc. with oo anatr. ov. Caps. Genera: Tristicha, Lawia, Weddellina. Tristichocalyx F. Muell. (Legnephora p.p.). Menispermaceae. 2 Austr. Tristira Radlk. Sapindaceae (i). 2 Malaya. Tristiropsis Radlk. Sapindaceae (i). 4 Mariannes, Solomons, N.G. Tristis (Lat.), dull coloured. Trisyngyne Baill. Euphorbiaceae (A. n. 7). 2 New Caled. Tritaxis Brill. Euphorbiaceae (A. II. 3). 4 Indomal. Triteleia Uougl (Brodiaea p.p.). Liliaceae (iv). 20 W. Am. Trithrinax Mart. Palmae (i. 2). S. Am. Tritliuria Hook. f. (Juncella EP.). Centrolepidaceae. 2 Austr., N. Z. Triticum L. Gramineae (12). 15 Medit., Eur., W. As. Spikelets 2 — 5 -flowered, in a dense spike. 3 sp. of wheat, with numerous sub- species, are cult, but the parent forms cannot be traced, and there is dispute as to the specific rank of these forms. T. monococcum L. is the one-grained wheat or small spelt (the grains of spelt do not fall out of the glumes when threshed), T. polonicitm L. the Polish wheat, TROPAEOLUM 663 and T. sativum Lam. the wheat proper, with 3 races, the ordinary spelt, the earliest cultivated (T. spelta L.), the 2 -grained wheat (T. dicoccum Schr.), and T. sativum tenax, of which there are 4 sub-races, (1) the hard or flint wheat (T. durum Desf.) used for macaroni, &c., (2) the turgid or rivet wheat (T. turgidwn L.), (3) the dwarf wheat (T. compactum Host.), and (4) the common wheat (T. vulgare Vill.) in oo vars. All are sometimes considered as forms of T. vulgare. Cf. Percival, Agric. Botany; Kornicke and Werner, Handb. d. Getrei- debaucs, Bonn, 1885, &c. Tritoma Ker-Gawl. = Kniphofia Moench (Lili.). Tritomopterys Niedenzu (Gandichaudia p.p.)- Malpighiaceae (i). 8 Mexico, C. Am. Tritonia Ker-Gawl. Iridaceae (in). 30 S. and trop. Afr. Cult. orn. Triumfetta Plum, ex L. Tiliaceae. 75 trop. Herbs or shrubs, often with extrafloral nectaries at base of 1. On each internode of infl. are usu. at least three 3-flowered dichasial cymes. The first and oldest is opp. to the 1. ; the rest stand alt. right and left between the first and the 1. Fruit with hooked spines (animal distr.). Triuridaceae (EP., BH.}. Monocots. (Triuridales ; Apocarpae BH.}. 2 gen., 40 sp. , trop. As., Afr., Am. Little saprophytes with scale 1. and small fls. on long stalks, $ or .;. Urticaceae p.p. BH.}. Dicots. (Archichl. Urticales). r3 gen-> J3.° sP-> tr°P- and temp. Trees with sympodial stems, bearing 2-ranked simple often asymmetrical 1 with stips. Fls. usu. in cymose clusters, generally unisexual. P 4 — 5, free or united, sepaloid,' theo- retically belonging to two whorls; A 4— 5, opp. the perianth-1., in two whorls; G rudimentary in i fl., in the ? of (2) cpls., sometimes 2-loc. but usu. i-loc., the second loc. aborting; ov. i per loc. , anatr. or amphitr., pend., style linear or bifid. Nut, samara or drupe. Seed usu. with no endosp. The wood of many is useful. Chief genera: Ulmus, Celtis. Ulmaria (Tourn.) Hill (Spiraea p.p. BH.). Rosaceae (in. 4). 10 N. temp. U. palustris Moench (Spiraea Ulmaria, meadow-sweet) and U. Filipendiila Hill (S. Filipendula, dropwort) in Brit. Ulmus (Tourn.) L. Ulmaceae. 18 N. temp, and Mts. of trop. Asia. 668 ULMUS U. montana With, (vvych elm) and U. campestris L. (elm) in Brit. Growth sympodial, the term, bud being suppressed. L asymmetri- cal, one side larger than the other (cf. Begonia). The fls. are § and come out before the 1. as little reddish tufts, each a short axis with a number of 1., beginning 2-ranked at the base and going over to 5-ranked above. There are no fls. in the axils of the lowest to or 12 ; in the axils of the upper 1. are fls. arranged in small dich. cymes (cf. Betulaceae), which are reduced, in U. campestris and others, to the one central fl. Each fl. has ?4 — 8 and as many sta. with i-loc. ovary. [See Chalazogamae.] Fr. a samara. The elm supplies a valuable timber. Umbel, an infl. in which the stalks of the fls. all spring from the top of the main stalk. Umbellales (BH.}. The i5th order of Polypetalae. Umbelliferae (EP., BH.}. Dicots. (Archichl. Umbelliflorae). 200 gen., 2700 sp., cosmop., chiefly N. temp. Many in Brit. Most can be recognized by habit ; herbs with stout stems, hollow internodes, and alt. exstip. sheathing 1. with their blades much divided pinnately. A few, e.g. Hydrocotyle and Bupleurum, have entire 1. Infl. usu. a cpd. umbel. At the top of the stalk of each partial umbel, an invol. of bracts is often found (the bracts of the outer fls.), and a similar larger invol. often occurs at the top of the main stalk bearing the cpd. umbel ; the latter is sometimes termed the involucre in contradis- tinction to the involucels of the partial umbels. A term. fl. often occurs, e.g. in Daucus. In a number of genera (e.g. Astrantia, Hydro- cotyle) simple umbels occur, cymose in type (as the non-centripetal order of opening of fls. shows) and often arranged in cymose groupings, e.g. in Sanicula. Eryngium has a cymose head. Some sp. of Xan- thosia and Azorella have such cymose infls. reduced to single fls., and these infls. have commonly invols. of bracts. Fl. usu. 5 and reg. (see below), epig. K 5, usu. very small, the odd sepal post. ; €5 (rarely o), usu. white or yellow; A -,, intr. ; G (2), antero-post., 2-loc. ; in each loc. one pend. ovule, anatr., with ventral raphe. On top of the ovary is an epig. disc, prolonged into two short styles. The massing of the fls. into dense infls. makes them conspic. (cf. Compositae), and this is aided by the zygomorphism of the C often seen; the outer petals of the outer fls. are drawn out (cf. Cruciferae) so as to form a sort of ray. Honey is secreted by the disc; it is accessible to all insects (fam. in class A). The chief visitors are flies ; fls. very protandrous, the cf stage being most commonly over before the ? begins. The ovary ripens into a very char, fruit, a dry schizocarp, which splits down the septum between cpls. into 2 mericarps, each containing one seed. The two are generally held together at first by a thin stalk (carpophore} running up between them. Floral diagram. The structure of the pericarp is of great import- ance in determining the gen. It is nearly always necessary to have ripe fr. in order to identify one of the U. The shape is often im- UMBELLIFERAE 669 portant ; the outer surface of each mericarp has generally 5 projecting primary ridges, two of which (the lateral ridges) are at the edges where the splitting takes place. Between these are sometimes secondary ridges, 4 to each mericarp. In the furrows are often found oil-cavities (seen as small openings in cross-section) known as vittae. The seed is often united to the pericarp ; it is album, with small embryo in oily endosp., which is usu. cartilaginous in texture. The shape of the endosp. as seen in cross-section is of importance ; it may be crescentic, or ventrally grooved, or concave on ventral side. The fr. often shows adaptations for distr. ; in many (e.g. Heracleum and allies) the meri- carp is thin and flat, suited to wind-carriage ; in others (e.g. Daucus) it has hooks. See also Scandix. Many U. are economically useful, but as a rule they are poisonous. See Daucus (carrot), Pastinaca (parsnip), Apium (celery), Crithmum (samphire), Foehiculum (fennel), Archangelica, Carum, Ferula, Pim- pinella, Coriandrum, Petroselinum, &c. Classification and chief genera (after Engler) : I. HYDROCOTYLOIDEAE (fr. with no free carpophore, and woody endocarp : vittae none or in main ribs). 1. Hydrocotyleae (fr. with narrow surface of union, lat. flat: tened) : Hydrocotyle, Azorella. 2. Mulineae (fr. with flattened or rounded back; ^T) : Bovv- lesia. II. .SANICULOIDEAE (endocarp soft, exocarp rarely smooth; style long with capitate stigmas, surrounded by ring-like disc ; vittae various). 1. Sanicnleae (ov. 2-loc. ; fr. 2-seeded, with broad surface of union ; vittae) : Eryngium, Astrantia, Sanicula. 2. Lagoecicae (ov. i-loc. ; fr. i -seeded ; vittae indistinct); Lagoecia. III. APIOIDEAE (endocarp soft, sometimes hardened by sub- epidermal fibre layers; style on apex of disc; vittae). A. Primary ridges projecting, the lat. sometimes wing-like; no secondary ridges. a. Secondary umbels each with i or few ? fls. surrounded by //., Cephaelis Sw.). Rubiaceae (n. 5). 130 trop. U. (C.) Ipecacuanha Baill. (/*. Ipecacuanha Stokes) is the ipecacuanha (Brazil), a herb with decumbent stem, and roots thickened somewhat like rows of beads. Root used in medicine. Uralepsis Nutt. = Triodia R. Br. p.p. (Gram.). Urandra Thw. (Lasiant/ius p.p.). Icacinaceae. 10 Indoinal. Uraria Desv. Leguminosae (in. 7). 12 palaeotrop. Urbania Phil. Verbenaceae (i). 2 Chili. Urbanodendron Mez (Aydendron p.p.). Lauraceae (n). i E. Brazil. Urbanodoxa Muschler (Cremolobus p-p.). Cruciferae (i). i Peru. Urbanosciadium H. Wolff. Umbelliferae (in. 4). i Peru. Urbinella Greenman. Compositae (6). i Mexico. Urbinia Rose (Echmcria p.p.). Crassulaceae. 3 Mexico. Urceocharis x Mast. Hybrid Urceolina - Eucharis (Gard. Chr, 1892). Urceola Roxb. Apocynaceae (n. i). 8 E. Indoinal. Urceolate, urn-shaped. Urceolina Reiclib. Amaryllidaceae (i). 3 Andes. Urechites Muell.-Arg. Apocynaceae (n. i). 5 W.I., Florida. Ureljrtrum Hack. Giamineae (2). 3 trop. and S. Afr. Urena Dill, ex L. Malvaceae (3). 3 trop. and subtrop. Schizocarp, the individual cpls. provided with hooks. Useful fibre. Urens (Lat.), stinging. Urera Gaudich. Urticaceae (i). 22 trop. Stinging hairs powerful. Achene enclosed in persistent fleshy P. (pseudo-berry). Urginea Steinh. Liliaceae (v). 40 Medit., Afr., India. U. Scilla Steinh. ( U. maritima Baker, squill), large bulbs used in medicine. Urmenetia Phil. (Onoseris p.p. EP.). Compositae (12). i Chili. Urnularia Stapf ( Willughbeia, &c. p.p.). Apocyn (i. i). 5 Malaya. Urobotrya Stapf. Opiliaceae. 4 trop. Afr. Urochlaena Nees Gramineae (10). 2 S. Afr. Urodesmium Naud. (Pachyloma DC.). Melastomaceae (i). 2 Brazil. Uropappus Nutt. = Microseris D. Don, p.p. (Comp.). Uropetalon Ker-Gawl. = Dipcadi Medic, p.p. (Lili.). Urophyllum Wall. Rubiaceae (i. 7). 40 trop. Afr. to Japan. Uroskinnera Lindl. Scrophularioceae (n. 4). 2 C. Am,, Mexico. Urospatha Schott. Araceae (iv). 15 trop. S. Am. Urospermum Scop. Compositae (13)". 2 Medit. Urostephanus Robinson^et Greenman. Asclepiadaceae (n. 4). i Mexico. Urostigma Gasp. = Ficus Tourn. p.p. (Mor.). Urotheca Gilg. Melastomaceae (i). i E trop. Afr. Uralnea Gaertn. Compositae (10). 60 S. Afr., Abyssinia. Urtica (Tourn.) L. Urticaceae (i). 30 temp, (nettles). Herbs with opp. 1. and slips, (sometimes united in pairs between the petioles, as in Rubiaceae), usu. covered with stinging hairs. The various types of infl. are well shown in the 3 Brit. sp. In general the infl. is a dich. cyme with tendency to cincinnus by preference of the /3-bracteole. In U. pihtlifera L. (Roman nettle) the rf and ? infls. spring side by side from each node, the i catkin-like, the 9 a pseudo-head. In U. iirens 672 URTICA L. (small nettle) a panicle is formed containing both i and ? fls. In U. dioica L. (large or common nettle) there is a panicle, but each sex is confined to its own plant. P 4; A 4, opp. to P leaves. The sta. are bent down inwards in the bud, and when ripe spring violently up- wards and bend out of the fl., the anther turning inside out, so that the loose powdery pollen is ejected as a little cloud, and may be borne by wind to the stigma. The ? fl. has a i-loc., i-ovuled ovary with a large brush-like stigma. Achene enclosed in the persistent P. Young tops eaten like spinach. Useful fibre from stems. Urticaceae (EP. ; BH. incl. Moraceae, Ulmaceae, Cynocrambaceae}. Dicots. (Archichl. Urticales). 41 gen., 480 sp., trop. and temp. Most are herbs or undershruhs, with no latex, and with alt. or opp. slip. 1. Intl. cymose often ' condensed ' into pseudo-heads, &c. Fls. usu. uni- sexual and reg. P 4 — 5, free or united, sepaloid; sta. as many, bent down inwards in bud and exploding when ripe; G i-loc. with i erect basal orthotr.ov.and i style. Achene. Seed usu. with rich oilyendosp.; embryo straight. Boehmeria, Urtica, Maoutia and others are used as sources of fibre. Classification and chief genera (after Engler) : A. With stinging hairs. P (4—5) in ? . L. alt. or opp. 1. Ureieae: Urtica, Urera, Laportea. B. No stinging hairs. 2. ProcriJeae (P of ? 3-merous, stigma paint-brush-like): Pilea, Pellionia, Elatostema. 3. Boehmerieae ( i usu. with 4—5 sta. No invol.): Boehmeria, Maoutia. 4. Parielarieae (P present ; bracts often united in invol. ) : Parie- taria. 5. Forskohleeae ( 3 fl. reduced to ( sta.). Ulmaceae are distinguished by infl., aestivation of sta., and ovule, and Moraceae by presence of latex, and also usu. by ovule, embryo, &c. Urticales. The 1 2th order of Archichlamycleae. Urticastrum Fabricius = Laportea Gaudich. (Urtic. ). Urticiflorae (Warming). The 4th order of Choripetalae. Urucury nut, Maximiliana. Urvillea H. B. et K. Sapind. (i). 10 warm Am. Lianeslike Serjania. Usteria Willd. Loganiaceae. i trop. W. Afr. Ustilago antherarum, Lychnis. Utleria Bedd. Asclepiadaceae (i). i S. India. Utricle, Cyperaceae. Utricularia L. Lentibulariaceae. 210 trop. and temp., the latter all aquatic. Some Brazilian sp. are confined to the pitchers of the Brome- liaceae. 3 in Brit. ; U. vulgaris L., the bladder-wort, common. The morphology is intere.-ting, for the usual distinctions drawn be- tween root, stem and 1. cannot be applied here. The common bladder- wort is a submerged water pi. with finely-divided 1. ; it never has roots, even in the embryo. The fls. project above water on short shoots, and there are also short shoots with small 1., which arise from the main axis and grow upwards to the surface. Upon the ordinary submerged 1. are borne the bladders, curious hollow structures with VAHADENIA 673 trap-door entrances. Small Crustacea and other animals push their way into the bladders and are not able to escape, for the doors only open from outside. The plant takes up the products of the decay of the organisms thus captured; it is very doubtful whether any special ferment is secreted. Other sp. are land pi. with peculiar runners, which develop in the moss or other substratum, on which they grow, and there bear the bladders. Others again, e.g. U. montana Poir., are epiph. with water storage in tuberous branches. The 1. of all these forms are simple. Goebel (PJianzenbiol. Sch.} has investigated the development of U. and finds that all these parts — 1., bladders, runners, water-shoots, erect shoots, &c. — are practically equivalent to one another, and that the same rudiment at the growing point may give rise to any one of them, or that they may themselves change from one to another type. Similarly on germ, a lot of spirally-arranged primary 1. are produced, and then one or two water-shoots appear lat. on the growing point, bearing no direct relation to the 1. in position, but apparently homologous with them. "LikeGenlisea, U. possessed originally a leaf-rosette, ending with an infl., and consisting partly of bladders. Then were added the swimming water-shoots or (in land forms) runners, which though externally unlike leaves (since they develop indefinitely and produce leaves and infls.) yet are originally homologous with them." For further details see Goebel, loc. dt. Hibernation in Brit, sp., &c. by winter buds full of reserves, which drop off and sink. Uvaria L. Anonaceae (i). 100 trop. Mostly lianes with recurved hooks (infl. -axes). The connective of the anther is usu. leafy. Uvariastrum Engl. Anonaceae (i). 2 W. trop. Afr. Uvariopsis Engl. ex Engl. et Diels. Anonaceae (i). i Cameroons. Uvarius (Lat.), like a bunch of grapes. Uva-Ursi Tourn. ex Moench = Arctostaphylos Adans. p.p. (Eric.). Uvifera L., O. Ktze. = Coccoloba L. (Polygon.). Uvularia L. Liliaceae (i). 4 E. N. Am. Cult. orn. fl. Vaccaria Medic. (Saponaria p.p. BH.}. Caryophyll. (n. 2). 3 Eur., W. As. Vacciniaceae (BH.}~ Ericaceae (§ in. Vacdnioideae). Vacciniopsis Rusby. Ericaceae (in. 2). i Bolivia. Vaccinium L. (incl. Oxycoccus Tourn.). Ericaceae (in. i). 120^, Andes, Madag. 4 in Brit. V. Myrlillus L. the whortle-, bil- or blae-berry, common in hilly districts. V. uliginosum L. at high levels. Both have deciduous 1. and blue berries. V. Vitis-Idaea L., the cow- or whimberry (often called cranberry by error), also a mountain sp., evergr. V. Oxycoccus L., the cranberry, in mountain bogs, a trailing evergr. with 1. edges rolled back. The fls. resemble Erica, both in structure and mech., but ov. inf. ; largely visited by bumble-bees. The fleshy fr. is ed. (used for jams, &c.) and is much distr. by birds. That of the N. Am. sp. V. pennsylvanicum Lam. is called blue huckle- berry. Vagaria Herb. Amaryllidaceae (i). i Syria. Vaginate, sheathed. Vagnera Adans. (Smiladna p.p.). Liliaceae (vn). 10 N. Am. Vahadenia Stapf (Landolphia p.p.). Apocynaceae (i. i). i Congo. w. 43 674 VAHEA Vahea Lam. (Landolphia p.p.)- Apocynaceae (i. i). 2 trop. Afr., Madag. Vahlia Thunb. Saxifragaceae (i). 4 warm Afr., As. Fls. in pairs (cymes) ; G. Vahy, Landolphia madagascariensis Benth. et Hook. f. Vailia Rushy. Asclepiadaceae (n. r). i Bolivia. Vaillantia Tourn. ex L. Rubiaceae (n. n). 2 Medit. Valcarcelia Lag. Leguminosae. Nomen. Valdivia Remy. Saxifragaceae (v). i Chili. Valentiana Rafin. Inc. sed. i Abyssinia. Valentina Speg. Boraginaceae (in), i Patagonia. Valentiniella Speg. (Valentino p-p.). Boraginaceae. i Patagonia. Valenzuelia Bert. Sapindaceae (i). 2 Chili, Argentina. Valerian, Valeriana ; red spur -, Centranthus. Valeriana Tourn. ex L. Valerianaceae. 180 Eur. , As., Afr., Am. V. officinalis L. and V. dioica L. in Brit, (valerian). Fls. protandr. The K forms a pappus upon the fr. Valerianaceae (EP., BH.). Dicots. (Sympet. Rubiales; Asterales^ZT.). 8 gen., 350 sp., Eur., As., Afr., Am. Herbs with exstip. 1. and dich. branching. Fls. in cymose panicles, &c. , £ or unisexual, asymmetric, usu. 5-merous. K sup., little developed at time of flowering, after- wards often forming a pappus as in Compositae ; C usu. (5), often spurred at base; A i — 4, epipet., alt. with petals; anthers intr. ; G (3); only i loc. is fertile, and contains i pend. anatr. ov. Achene. Seed exalbum. Chief genera: Valerianella, Valeriana, Centranthus. Valerianella Tourn. ex Hall. (BH. incl. Pledritis DC.). Valerianaceae. 50 N. temp.; 4 Brit, (corn-salad or lamb's lettuce). Seed-dispersal mech. various. In V. Auricula DC. the sterile loc. of the fr., in V. vesicaria Moench the K, is inflated, in V. discoidea Loisel. it forms a parachute, whilst in others it is provided with hooks. Valerianopsis C. A. Muell. (Valeriana p.p. EP.). Valer. 7 Brazil. Valetonia Durand ex Engl. Icacinaceae. i Brazil. Validallium Small (Allhun p.p.). Liliaceae (iv). i N. Am. Vallaris Burm. f. Apocynaceae (11. 2). 6 Indomal. Vallea Mutis ex L. f. Elaeocarpaceae. 3 Colombia, Peru. Vallecula (Lat.), grooves in fruit. Vallesia Ruiz et Pav. Apocynaceae (i. 3). 2 Florida to Argentina. Vallisneria Mich, ex L. Hydrocharitaceae. 3 trop. and subtrop. V. spiralis L. (Eur.) a dioec. submerged water-pi, with ribbon 1. i fls. in dense spikes enclosed in spathes; when ready to open the fls. break off and float up to the surface, where they open. ? fl. sol. on very long stalk, which brings it to the surface; it has green P, inf. ov. and 3 large stigmas. Pollination occurs on the surface (cf. Elodea) ; and after it the stalk curls up into a close spiral, dragging the young fr. to the bottom to ripen. Veg. repr. by runners, rooting at the ends. Vallota Herb. Amaryllidaceae (i). i Cape Colony. Cult. orn. fl. Valonia, Qiierciis Aegilops L. Valvate (aestivation), 1. touching, not overlapping. Valves, the portions into which a fruit splits. Valvular dehiscence (anther), Berberis, Lauraceae. Vanclevea Greene (Grindelia p.p.). Compositae (3). i Utah. VEGETATION 675 Vancouveria C. Morr. et Dene. (Epimedium p.p. EP.). Berberidaceae. 2 N.W. Am. Vanda Jones. Orchidaceae (n. 20). 25 Indomal. Epiph. with fleshy 1., sometimes cylindrical. Cult. orn. fl. Vandellia L. (Lindemia All.). Scrophul. (11. 6). 30 trop., subtrop. Vandopsis Pfitz. (Vanda p.p.). Orchidaceae (n. 20). 3 E. Indomal. Vangueria Juss. Rubiaceae (n. i). 40 trop. Afr., As., Madag. Vanhouttea Lem. (Hoiittea Decne.). Gesneriaceae (n). 4 Brazil. Vanilla Plum, ex L. Orchidaceae (11. 2). 30 trop. Climbers with fleshy 1. and thin velamen (see fam.). V. plan ifolia Andr. (Mex.) is cult. ; its pods form the spice vanilla. Vanilla, Vanilla planifolia Andr. ; -grass (Am.), Hierochloe. Vanillosma Spach=Piptocarpha R. Br. p.p. (Comp.). Vanillosmopsis Sch.-Bip. Compositae (i). 7 Brazil. Vaniotia Leveille. Scrophulariaceae (in. i). i China. Vanoverberghia Merrill. Zingiberaceae (i). i Luzon. Vantanea Aubl. Humiriaceae. 5 Brazil, Guiana. Varangevillea Baill. Verbenaceae (*). i Madag. Vargasia Ernst (Caracasia Szysz.). Marcgraviaceae. 2 Venezuela. Variation expresses the fact that no two beings are exactly alike, and that there may exist constant slight differences between two forms, expressed in the idea of varieties, e.g. a pink and a white variety of the same rose. Cf. Nomenclature. Varilla A. Gray. Compositae (5). 2 Texas, Mexico. Varnish, a solution of resin in oil of turpentine, alcohol, or other solvent ; cf. resins; Chinese - tree, Aleurites Fordii Hemsl. Varronia P. Br. (Cordia p.p.). Boraginaceae (i). 4 W.I. Varthemia DC. (Iphiona BH'.). Compositae (4). 4 Egypt to N.W. India. Vasconcellea A. St Hil. = Carica L. p.p. (Caric.). Vasconcellosia Caruel (Carica p.p. EP.). Caricaceae. i Brazil. Vaselia Van Tiegh. (Efaasia p.p.). Ochnaceae. i Brazil. Vaseyanthus Cogn. Cucurbitaceae (4). 2 Calif. Vasivaea Baill. Tiliaceae. i Amazon valley. Vassobia Rusby. Solanaceae (2). i Bolivia. Vateria L. Dipterocarpaceae. 3 Seychelles, S. India. V. itidica L. yields a gum-resin (Indian copal, white dammar). Vatica L. Dipterocarpaceae. 45 Indomal. Several yield resins and useful timbers. Vaupelia Brand (Trichodesma p.p.). Boragin. (iv. i). 6 trop. Afr. Vauquelinia Correa ex Humb. et Bonpl. Rosaceae (i. 2). 4 Mex., Ariz. Vausagesia Baill. Ochnaceae. 2 W. trop. Afr. Vavaea Benth. Meliaceae (in). 5 Polynesia. Vazea Fr. Allem. ex Mart. Olacaceae (nomen). i Brazil. Veatchia A. Gray. Anacardiaceae (3). i California. Vegaea Urb. Myrsinaceae (n). i S. Domingo. Vegetable (for eating), cf. Edible Products; -horsehair, Tillandsia; -ivory, Pahnae, Phytelephas; -marrow, Cucitrbita Pepo L., var. ; - oyster, Tragopogon porrifoliiis L. ; - sheep, Raoulia. Vegetation, see Forms of-, Zones of -. 43—2 676 VEGETATIVE REPRODUCTION Vegetative reproduction, by the detachment of portions of the veg. system, which may grow into new plants; a branch may be detached without modification (water pi., rhizomes, &c.), or may be specialised, with supplies of reserves; special cases are bulbs, bulbils, conns, tubers, buds, &c. ; Agave (bulbils), Agropyron (rhizomes), Ajuga (runners), A/Hum, Asplenium, Begonia (adv. buds, tubers), Bettis (rhizomes), Bertolonia, Bryophyllitm (adv. buds), Cactaceae (mammillae), Carda- mine, Carex (off-shoots), Chlorophytnm(^QC\^\ shoots), Crassula (pi. in placeof fl.), Crassulaceae, Cystopteris,Epilobium,Fadyenia, Fragaria (runners), Gagea, Gesneriaceae (suckers), Globba, Hymenophyllaceae (gemmae), Liliiim, Limnobinm, Limosella (runners), Lycopodium (bulbils), Mannnillaria (mammillae), Mercurialis (rhizomes), Nastitr- tinm, Nephrolepis, Ophioglossitm (adv. buds), Opuntia (branches), Oxalis,Polygomt»i, Potentilla, Prionium, Psilotiiw (gemmae), Ranun- culus, Remnsatia (hooked bulbils), Rosaceae, Ritbus, Saccharum, Salicaceae (suckers), Sempervivum (offsets), Senecio (stem joints), Sinnitigia, Tnssilago, Vallisneria, Zingiber. Veins (of 1.), the stiff vascular bundles running throughout. Veitchia H. Wendl. Palmae (iv. i). 4 Fiji, New Hebrides. Velaea DC. (Arracacia BH.). Umbelliferae (in. 4). 20 N. Am. Velamen, Araceae, Orchidaceae. Velezia L. Caryophyllaceae (n. 2). 4 Medit. to Afghanistan. Vella L. Cruciferae (2). 3 W. Medit. Thorns = stems. Velleia Sm. Goodeniaceae. 12 Austr. Ovary ± sup. Vellozia Vand. Velloziaceae. 40 Brazil, esp. campos. Velloziaceae (EP. ; AmaryUidateae p.p. BH.). Monocotyledons (Lilii- florae). 2 gen., 70 sp., Brazil, Afr., Madag. Xero., chiefly of rocky places or dry campos. Perenn. with dichot. branched stems and 1. in rosettes (cf. Aloe). Upper parts of stems clothed with fibrous sheaths of old 1., lower parts with adv. roots. The stem is thin, but its coating of roots may be inches deep. Water poured over the roots disappears as if into a sponge, and the pi. is thus able to supply itself from dew, &c. during the dry season. The 1. also are xero. Fls. sol., term., reg. P 3 + 3, coralline; A 3 + 3, or oo, in bundles. G 3-loc., with placenta in the form of lamellae, ± peltately widened or thickened at the outer side. Ovules oo . Caps. Endosp. Genera: Vellozia, Barbacenia. Chief differences from Amaryllid. in A and plac. Velloziella Baill. (Digitalis p.p.)- Scrophul. (in. i). i Brazil. Velophylla Benj. Clarke. Podostemaceae. i Brazil. Veltheimia Gleclitsch. Liliaceae (v). 3 S. Afr. Cult. orn. fl. Velutinus (Lat.), velvety. Velvet bean, Muatna; -bur (W.I.), Priva; -grass (Am.), Holcus lanatns L. ; -leaf (W.I.), Cissampehs Pareira L. ; -seed (W.I.), Guettarda ; - tamarind, Dialium. Velvitsia Miern. Scrophulariaceae (ill. 2). i Angola. Venation, arrangemeut of the veins in a 1. Venegasia DC. Compositae (6), i California. Venenatus (Lat.), poisonous. Venice turpentine, Larix europea DC. Venidium Less. (Arclotis p.p. EP.}. Compositae (10). 18 S. Afr. Venose, with veins. VERBENA CEAE 677 Ventenata Koel. Gramineae (9). 3 Eur., As. Minor. VentilagoGaertn. Rhamnaceae. ippalaeotrop. Some climb by hooks. Fr. with wing on upper end, formed from style after fert. Ventral (surface), upper, towards axis, but not consistently used. Ventricose (C), with basal part swollen out all round. Venulose, thickly veined. Venus' comb, Scandix ; - flytrap, Dionaea ; - looking glass, Specularia. Veprecella Naud. Melastomaceae (i). 20 Madag. Vepris Comm. ex A. Juss. (Toddalia BH.}. Rutaceae (iv). 6 palaeo- trop. Veratrilla Franch. Gentianaceae (i). i China. Veratrin, Sckoenocaulon, Veratrum. Veratrum (Tourn.) L. Liliaceae (i). 10 N. temp. Rhiz. with leafy stem and racemes, lower fls. 5 » but upper usu. , spathes (not indicated in the upper shoots). B. Spathe of Z. nana with flattened spadix taken out; v, the retinaculum; X2. C and D. Half-anther and pistil of same, more enlarged. E. Fruit of Z. marina, the thin pericarp turned back to show the seed, X2j. F. Diagram of a main axis, i, with its fore-leaf (/>') and the axil- lary shoot 2, with its fore-leaf (b-) ; sp, spathe borne on 2, surrounding the spadix; s, intravaginal scales; v, bracteole. G. Diagram of part of spadix with two flowers ; v, bracteole. A, F, G, after Eichler; B, C, D, from English Botany; E, after Le Maout and Decaisne. above its point of origin. This is most easily seen in the infl. region ; the branching is that of a rhipidium, but shoot i, which springs from the axil of a 1. on shoot i, is adnate to i up to the point at which the first 1. is borne on i; this 1. occupies the angle between the two shoots where they separate. Shoot i (and 2, 3, &c. successively) is pushed aside and bears an infl. (Eichler's Bliithendiag. or Nat. Pft. for details and figs.) L. long, linear, sheathing at base. yoo ZOSTER A Infl. a flattened spadix, enclosed at flowering time in a spathe (the sheath of the uppermost 1.). This is open down one side, and on the corresponding side of the spadix the fls. are borne, the essential organs forming two vertical rows, each composed of a cpl. and a sta. alt. On the outer side of the spadix next the sta. is often a small 1. (retinacnltim of systematic ret ^J; ^ ' works). The midrib of the cpl. faces outwards. ' cpi' sta] ret. Each cpl. contains one ovule and has two flat stigmas. The sta. consists of two half anthers, joined by a small connective. It is difficult to decide what is the actual 'flower' in this plant ; the usual view is that each sta. witli the cpl. on the same level forms a fl., the retinaculum representing the bract. Fert. peculiar, Z. being one of the water pi. most completely modified from the ancestral land-pi, type. Fl. submerged like the rest of the pi. The pollen grains are long threads of the same specific gravity as salt water, so that when discharged they float freely at any depth. The stigmas are very large, and thus have a good chance of catching some of the grains. The whole mech. is similar in principle to that of a wind-fert. pi. Fr. an achene. In winter it hibernates without any special modification. The pi. is largely used for packing glass, stuffing cushions, &c.,.esp. in Venice. [For further details see fam., and Schenk's Wassergewachse. ] Zosterella Small. Pontederiaceae. i Pennsylvania. Zoysia Willd. Gramineae (3). 3 Masc. Is. to New Zealand. Zozimia Hoffm. Umbelliferae (ill. 6). 6 W- As. Zschokkea Muell.-Arg. (Lactnellia BH.}. Apocynaceae (i. i). 8 trop. S. Am. Zuccagnia Cav. Leguminosae (n. 7). i Chili. Zuccarlnia Blume. Kubiaceae (i. 8). i Java. Zuccarinia Maerklin. Inc. sed. Nomen. Zuckia Standley. Chenopodiaceae (A), i S.W. U.S. Zuelania A. Rich. (Casearia p.p. BH.}. Flacourtiaceae (7). i W.I. Zwackhia Sendtner. Boraginaceae (iv. 5). i S.E. Eur. Zwartbast. Royena lucida L. Zwingera Schreb. = Simaba Aubl. (Simarub.). Zycona O. Ktze. = Allendea La Llave (Comp.). Zygadenus Michx. Liliaceae (i). 12 N. Am., Siberia. Zygalchemilla Rydberg (Alchemilla p.p.). Rosaceae (in. 5). i trop. Am. Zyganthera N. E. Br. (Pseitdohydrosme p.p.). Araceae(iv). i Gaboon. Zygella Sp. Moore. Iridaceae (n). i Matto Grosso. Zygia Walp. = Albizzia Durazz. p.p. (Legum.). Zygo- (Gr. pref. ), yoke-; -morpMsm, symmetry about one plane through axis only, cf. floral diagrams of Labiatae, Papaveraceae, Scrophulariaceae ; usu. antero-post., it is diagonal in Solaiiaceae, transverse in Atiigozanthos, Haemodoraceae, some Papaveraceae, right and left-handed in Cassia, Exacttm, Saintpanlia ; -te, the cell pro- duced by the union of the sexual gametes. Zygobatemannia x Rolfe, Zygocolax x Rolfe, Zygonisia x Rolfe, Zygo- mena x Hort. Orchidaceae. Hybrids of Zygopetalum with Bate- mannia, Colax, Aganisia, and Zygosepalum (Menadenium). ZY GO STIGMA 701 Zygodia Benth. Apocynaceae (n. j). 4 trop. Afr. Zygogynum Baill. Magnoliaceae. 3 New Caled. Zygomenes Salisb. = Cyanotis D. Don (Coinmel.). Zygonerion Baill. (Strophcuithns p.p.). Apocyn. (n. i). i Angola. Zygoon Hiern. Rubiaceae (i. 8). i Zambesi. Zygopetalum Hook. (incl. Pescatorea Rchb. f.)'. Orchidaceae (n. 14). 20 trop. Am. The base of the column forms a chin. Cult. orn. fl. Zygophyllaceae (EP., BH.}. Dicots. (Archichl. Geraniales). 22 gen., 160 sp., xero- or halo-phytes, trop. and subtrop. Most are woody perennials ; 1. opp. . slip., usu. hairy, fleshy or leathery. Fls. in cymes, reg., $ . K 5 ; C 5 ; A 5 + 5, obdiplost. and with ligular appendages ; £i(5)> 5-loc. with i or more pend. ov. in each loc. Fr. usu. a caps. Seeds with or without endosp. Guaiacum, Peganum, &c. furnish useful products. Chief genera: Zygophyllum, Guaiacum, Forlieria, Larrea, Peganum, Tribulus. Closely related to Rutaceae. Zygophyllidium Small (Euphorbia p.p.). Euph. (A. n. 8). i N. Am. Zygophyllum L. Zygophyllaceae. 60 |#, deserts and steppes. L. and twigs fleshy. Zygoruellia Baill. Acanthaceae (iv. A), i Madag. Zygosepalum Reichb. f. (Zygopetalum Bfl.). Orchid, (u. r4). 2 trop. S. Am. Zygostates Lindl. Orchidaceae (n. 19). 4 Brazil. Zygostelma Benth. Asclepiadaceae (i). i Siam. Zygostelma Fourn. Asclepiadaceae (n. i). i Brazil. Zygostigma Griseb. Gentianaceae (i). 2 Brazil, Argentina. Suggestions for additions to and corrections of the preceding pages can be noted on this slip, which can be detached when desired, signed, and sent to DrJ. C. Willis, cjo The University Press, Cambridge SUPPLEMENT. TEXT CORRECTIONS, AND NEW GENERA. w. 45 SUPPLEMENT. TEXT CORRECTIONS, AND NEW GENERA. This supplement will be brought up to date at intervals. Abroma, A. augnsta for L. read L. f. Acaena. For (in. 9) read (in. 5) and place A. before Acajou. Acanthura. For (iv. B) read (iv. A). Accidental, adventitious. Achradelphia O. F. Cook (Achras p.p.). i S. Am. (A. mammosa L.). Acmispon Rafin. (Hosackia p.p.). Leguminosae (ill. 5). 6 N.Am. Acrostigma O. F. Cook et Doyle. Palmae (iv. i). i Colombia. Acrosynanthus Urb. Rubiaceae (i. 3). i Cuba. Adelocaryum Brand. Boraginaceae (iv. i). i Tibet. AdenoracMs Nieuwland (Aroma p.p.). Rosaceae (n). 3 N. Am. Adhesion, union of members of different whorls, e.g. C and A. Adjustment (Cl.), functional response to stimuli. Adnascens (Lat.), growing upon something. Adoxa, add P homochlam. Adverse, facing main axis. Aeginalitis, read Aegialitis. Afrotrewia Pax et K. Hoffm. Euphorbiaceae (A. n. 2). i Cameroons. Agaloma Nieuwland (Euphorbia p.p.). Euphorbiaceae (A. n. 8). 14 N. Am. Agrostemma, add Caryophyllaceae (n. i). Aiolon Lunell (Anemone p.p.). Ranunculaceae (3). i N. Am. Aistopetalum Schlechter. Cunoniaceae. 2 New Guinea. Aizoaceae. Line 7, add fl. £ reg. Ajowan (for thymol), Caruin copticuin Benth. et Hook. f. Ajugoides Makino (Stachys p.p. ). Labiatae (vi). i Japan. Alatus (Lat.), winged. Allioniaceae = Nyctaginaceae. Alpine rose, Rhododendron ferrugineum L. Alternifolius (Lat.), alternate-leaved. Amaryllidaceae. For G read G. Ambianella A. Chevalier (Mimusops p.p.). Sapotaceae (2). i Congo. Amblynotopsis Macbride (Antiphytum p.p-)- Boraginaceae (iv. 4). 4 trop. N. Am. Amelanchus Franz Miiller ex V. Ros. (Amelanchier p.p.). Rosaceae (n). 3 N. temp. Amides, crystallisable bodies, containing C, H, Oand N; asparagin, &c 708 SUPPLEMENT Andropterum Stapf. Gramineae (2). i Nyassaland. Anelsonia Macbride et Payson (Draba p.p.). Cruciferae (4). i N.W. U.S. Animals. For Seed-dispersal read Dispersal of Seeds. Anota Schlechter (Saccolabium p.p.). Orchidaceae (n. 20). i Malaya. Antheroporum Gagnepain. Leguminosae (in. 8). 2 Sikkim, Cochin- china. Apodiscus. For Guiana read Guinea. Aranella Barnhart ex Small (Utricularia p.p.). Lentil mlariaceae. i Florida, Cuba. Arisanorchis Hayata. Orchidaceae (n. 2). i Formosa. Aristopetalum Schlechter. Cunoniaceae. 2 New Guinea. Artemisiastrum Kydberg (Artemisia p.p.). Compositae (7). i Calif. Azolla. After A. niloticci read Decne. Balanophoraceae. For G read G. Baliospermum : for 4 sp. read 10. Barombia Schlechter. Orchidaceae (11. 20). i Cameroons. Bathiea Schlechter. Orchidaceae (u. 20). i Madag. Beckeria Ridley, read Becheria. Beesia Balf. f. et W. W. Smith. Ranunculaceae (i). i Burma, Yunnan. Belandra Blake. Apocynaceae (n. i). i Brit. Honduras. Betchea Schlechter. Cunoniaceae. 5 New Guinea, N. E. Austr. Blandibractea Wernham. Rubiaceae (i. 3). i Brazil. Bombacopsis Pittier (Pachira p.p.). Bombacaceae. 2 Panama, Costa Rica. Bovleria err. = Borreria. Brassica. Line 2, after Cruciferae for (n. 10) read (2). Broussonetta, read Broussonnetia. Callus (in a grass), the swollen base of the inferior palea, adnate to the axis. Campos, open grassy plains of Brazil, with groups or patches of trees. Camptophytum Pierre ex A. Chevalier. Rubiaceae (inc. sed.). i Congo. Capillipedium Stapf. Gramineae (2). 6 warm |*. Capitanopsis Sp. Moore. Labiatae (vn). i Madag. Capporea, read Capnorea. Carenophila Ridley. Zingiberaceae (i). i Malay Peninsula. Carpentia Ewart in Ewart et Davies. Convolv. (i.) i N. Austr. Caryophyllaceae. In last para, after "4 groups" read of I. Catopsis. Before W.I. read 10. Celeonia, read Celebnia. Censer-mechanism. For Seed-dispersal read Dispersal of Seeds. Chaboisseae, read Chaboissea. Chaetothyla, read Chaetothylax. Chamaecyparis. 6". Lawsoiriana is from N.W. N. Am. Chamartemisia Rydberg(77r;/rt^«wp.p.). Compositae (7). i Nevada. Charlock, Sinapis alba L. Chasmopodium Stapf. Gramineae (2). 2 trop. Afr. Chloranthaceae. Before perisperm omit no. Chorilepidella Van Tiegh. (Loranthns p.p.). Loranthaceae. i Phil. Is. Chrysophae Koso-Poliansky. Umbelliferae (in. 5). 2 E. Medit. SUPPLEMENT 709 Clarorivinia Pax el K. Hoffm. (Mailotusp.p.). Euphurbiaceae (A. n. 2). i New Guinea. Cocos. For Palmae (iv. 7) read (iv. 2). Coelopyrena Yalelon. Rubiaceae (n. 5). i Ainboina. Columelliaceae, for G read G. Compositae. In classification, lini/i-ac, read (as 3. Conceveibastrum Pax et K. Hoffm. (Alchornca p.p.). Euphorbiaceae (A. II. 2). i Brazil. Conocarpus, add W.I. _ Cornaceae, for G read G. Corollonema Schlechter. Asclepiadaceae (n. i). i Bolivia. Costaricia II. Christ. Polypodiaceae. i Costa Rica. Cremnobates Ridley. Rhizophoraceae. i New Guinea. Cymbidiella Rolfe (Cymbidium p p.). Orchiclaceae (n. 17). 3 Madag. Cymopterus. For (in. 6) read(\\\. 5). Cynopaema Lunell (Apocynum p.p.). Apocynaceae (n. i). 3 N. Am. Dasypetalum Pierre ex A. Chevalier. Flacourtiaceae (i). i Congo. Davidia. For Cornaceae rra^Nyssaceae. Delphiniastrum Nieuwland = Delphinium Tourn. p.p. Desmopsis Safford (Unona p.p.). Anonaceae (i). =, C. Am. Desmos Safford (Unona p.p.). Anonaceae (i). 30 |^r. Deuteromallotus Pax et K. Hoffm. (Mallotm p.p'). Euph. (A. II. 2). i Madag. Digitalis. For (in. n) nad (\\\. i). Diodella Small. Rubiaceae (n. 10). i Florida, W.I. Dischidia. For Asclepiadaceae (n. 4) read (u. 3). Doliehostegia Schlechter. Asclepiadaceae. i Phil. Is. Echinaceae, read Echinacea. Ecliptostelma T. S. Brandegee. Asclepiadaceae (n. i). i Mexico. Eleutherostemon Herzog (Themistoclesia p.p.). Ericaceae (in. 2). t Bolivia. Enanthe Schlechter (Vanda p.p.). Orchidaceae (n. 20). i Phil. Is. Endospermum. For 5 sp. read to. Eremopogon Stapf. Gramineae (2). 4 warm ^. Euclasta Franch. Gramineae (2). i trop. Afr., Am. False asphodel, read Tofieldia. Fenixia Merrill. Compositae (=;). i Phil. Is. Festuca, After (Tourn.) read L. Floccose, bearing tufts of hair. Gibbsia Rendle in L. S. Gibbs. Urticaceae (3). 2 N.W. Dutch New Guinea. Giorgiella De Wild. Passifloraceae. i Congo. Glume, cf. Cyperaceae, Gramineae. Gonystilus. For entirely read entire. Gramineae, at end, to § 13, add CL Bamboos. Greengage, a var. of plum. Guerreroia Merrill. Compositae (5). i Phil. Is. Haemodoraceae, line ,s, after transv. read -\- . Hecatostemon Blake. Flacourtiaceae (7). i Venezuela. Helleborine. For Secapias read Serapias. 7io SUPPLEMENT Hernandiaceae, add fr. winged, no enclosp. Hippia F. W. Schmidt, read Hippion. Hoyella Ridley. Asclepiadaceae (n. 3). i Sumatra. Hydropectis Rydberg (Pectis p.p.)- Compositae (6). i Mexicu. Hyperapsis, read Ilyperaspis. Hypogyniura Nees in Stapf. Gramineae (2). 2 trop. Am., W. and S. Afr. Idenburgia L. S. Gibbs. Monimiaceae. 2 N.W. Dutch New Guinea. Ilocania Merrill. Cucurbitaceae (3). i Phil. Is. Jardinea Steud. Gramineae (2). 3 trop. Afr. Jolly brown, read Jolly, brown. Juglandaceae. For stip. read exstip. Jumellea Schlechter (Angraecum p.p.). Orchidaceae (n. 20). 12 Ma- dag., Masc. Kingiella Rolfe. Orchidaceae (n. 20). 5 Indomal. Klaineastrum Pierre ex A. Chevalier. Melastomaceae (in), J Gaboon. Koordersiochloa Merrill. Gramineae (9). i Java. Lacaenae, read Lacaena. Laretia, Laricopsis (after Larch) should follow Lardizabalaceae. Leitneriaceae, for 22 sta. read 12. Leptocentrum Schlechter. Orchidaceae (n. 20). i W. Afr. Leucocorema Ridley. Icacinaceae. i New Guinea. Linoma O. F. Cook (Dictyosperma p.p.)- Falmae (iv. i). i Masca- renes. Litosiphon Pierre ex A. Chevalier (Lovoa p.p.). Meliaceae (n). i Congo. Loranthaceae, p. 395, line 12 of 3rd para, for vicsin read viscin. Loxsomopsis Christ. Polypodiaceae. 3 trop. Am. Macrocatalpa Britton. Bignoniaceae (2). 3 W.I. Mahoberberis C. K. Schneider. Hybrid, Mahonia-Berberis. Mairella Leveille = Phelypaea Tourn. p.p. (Orob.). Manuleopsis Thellung. Scrophulariaceae (ll. 5). i Hereroland. Meiomeria Standley (Chenopodinm p.p.). Chenopodiaceae (A), i Mexico. Meliaceae. BH. incl. § n of Rutaceae. Meliadelpha should go below Meliaceae. Melioschizzia, read Melioschinzia. MiscantMdium Stapf. Gramineae (2). 4 trop. and S. Afr. Mischopleura Wernham in Ridley. Ericaceae (i. 2). 2 New Guinea. Monelasum, read Monelasmum. Monotroppa, read Mpnotropa. Mormodes. For Orchidaceae (17) read (n. ti). Musa, add The sp. cult, in the Canaries is M. Cavcndishii Lambert ex Paxt. (China). Musaceae, line 6, for =1. read =. Myricaceae, for nut, read drupe or nut. MyrmecopMla Rolfe (Schomburgkia p.p.). Orchidaceae (ll. 6). 7 C. and trop. S. Am. Myzodendraceae, line 2, after very, read small. Neomanniophyton Pax et K. Hoffm. Euphorbiaceae (A. n. 2). 12 W. Afr. SUPPLEMENT 711 Neotrewia Pax et K. Hoffm. (Mai lot us p.p.). Euphorbiaceae (A. n. 2). i Phil. Is. Neowollastonia Wernham in Ridley. Apocynaceae (i. 3). i New Guinea. Nephelochloa Boiss. Gramineae (10). i W. As. Nephradenia Decne. in DC. Asclepiadaceae (n. 3). 5 Brazil to Mex. Notomia, read Notonia. Nyctaginaceae, add seed with perisperm. Nyssaceae. After alt. exstip. read \. Octoknemataceae (Olacaceae p.p.). Dicots. (Santalales). Only genus Octoknema. Woody pi. with alt. 1. and stellate hairs. Fls. ? , P 2 — 3, A 2 — 3, anteposecl, G i-loc. with thread-like plac. pressed against the wall, and 3 pend. ov. Fr. drupaceous, i-seeded. Oreophila D. Don = Hypochoeris L. (Comp.). Oxycarpha Blake. Compositae (=,). r Venezuela. Pachystoma (Klotzsch) Muell.-Arg., read Pachystrotna. Palmervandenbroekia L. S. Gibbs. Araliaceae (r). i N.W. Dutch New Guinea. Papuzilla Ridley. Cruciferae (2). i New Guinea. Parasyringa W. W. Smith (Syringa p.p.). Oleaceae. i Yunnan. Pentaphylax. For caps, read Caps. Pentastira Ridley. Icacinaceae. 2 New Guinea. Petalocaryum Pierre ex A. Chevalier. Olacaceae. i Gaboon. Phaenoeuron, read Phaeoneuron. Phalacroceris, read Phalacroseris. Phyllomolia, read Phyllomelia. Pfcyllitis. Add Cf. Scolopendrium. Pierreodendron A. Chevalier. Sapotaceae. i Gaboon. Plastolaena Pierre ex A. Chevalier (Schumanniophyton Harms). Rubiaceae (i. 8). i Congo. Platanus, for G 4 or 3 read i, 4, or 3. Pleurodiscus Pierre ex A. Chevalier. Sapindaceae (i). 2 W. trop. Afr. Pocillaria Ridley. Icacinaceae. i New Guinea. Poikilogyne E. G. Baker in L. S. Gibbs. Melastomaceae (n). i N.W. Dutch New Guinea. Pontederiaceae. For G read G_. Postea, read Postia. Proteaceae. For G read G. Psychanthus Ridley (Alpinia p. p. ). Zingilieraceae ( i ). 6 New Guinea. Pyrogennema Lunell (Epilobium p.p.). Onagraceae (2). i (E. angnsti- foliuni L.) N. temp. Pyrsonota Ridley. Cunoniaceae. i New Guinea. Raritetoe Wernham. Rubiaceae (i. 8). r Colombia. Resinocaulon Lunell (Silphium p.p.). Compositae (5). i E. U.S. Rhysolepis Blake (Vigidera p.p.). Compositae (5). 2 Mexico. Riseleya Hemsl. Euphorbiaceae (A. i. i). i Seychelles. Ropoutea, read Ropourea. Rosa. For (in. 10) read (in. 6). Rossittia Ewart in Ewart et Davies. Rutaceae (i). i N. Austi. Sagentia (after Sarga), read Sargentia. 7i2 SUPPLEMENT Salacicrater, read Salacicratea. Samanea Merrill (Pithecolobium p.p.). Leguminosae (i. i). i S. Am. (P. Sam an Benth.). Santalaceae. For L. opp. read opp. or alt. Saugetia Hitchcock et Chase. Gramineae (t i). i Culm. Scheuchzeriaceae. For 12 sp. read 15. Sehismocarpus Blake. Loasaceae. i Mexico. Schizandria, read Schizandra. Seed-dispersal, see Dispersal of Seeds. Setosa Ewart in Ewart et Davies. Gramineae (3). i N. Austr. Simarubaceae before 5 read rarely. Simonenium Diels (Cocculus p.p.). Menisp. i E. As. Spathia Ewart in Ewart et Davies. Gramineae (2). i N. Austr. Sperauksia, read Speranskia, and 3 China. Stemonaceae. For G read G. Ternncius Lunell (Thlaspi p.p.). Cruciferae (2). i N. temp. |*. Tesmannia, read Tessmannia. Thuranthos C. H. Wright. Liliaceae (v). i S. Afr. Thylacophora Ridley. Zingiberaceae (i). i New Guinea. Thyrsia Stapf. Gramineae (2). 4 trop. Afr., India. Tiliaceae, add fr. usu. a caps, or schizocarp ; endosp. Trailliaedoxa W. W. Smith et Forrest. Rubiaceae (inc. sed.). i Yunnan. Trifidacanthus Merrill. Leguminosae (in. 7). i Ph. Is. Trillesianthus Pierre ex A. Chevalier. Tiliaceae. r Congo. Trithrinax, add 3 (species). Triuris Miers. Triuridaceae. 2 Brazil. Trophisomia Roj. Moraceae (n). i Chaco (Argentina). Turricula Macbride (Naina p.p.). Hydrophyllaceae. i California. Usoricum Lunell (Onagra p.p.). Onagraceae'(2). i N. Am. Veconcibea Pax et H. Hoffin. (Conceveiba p.p.). Euphorbiaceae (A. n. 2). 2 Brazil. Vesicarpa Rydberg (Artemisia\>.y.}. Compositae (7). i W. U.S. Volkensinia Schin/. \Kentrosphaera p.p.). Amarantaceae (2). i trop. Afr. Wine-berry, Rubus phoenicolasius Maxim. (|apan). Zingiber, add 55 Masc., trop. As. 1 6 Dec. 1918. KEY TO THE FAMILIES OF FLOWERING PLANTS BASED ON ENGLER'S CLASSIFICATION AS GIVEN IN DIE NATURLICHEN PFLANZENFAMILIEN, AND REVISED IN HIS SYLLABUS, ED. 7. W. 46 ii MONOCOTS. PANDAN ALES— TRIURID ALES A; a; a; (3 MONOCOTYLEDONEAE [Embryo with one cot.; stem with closed bundles; 1. usu. |]-veined; fl. usu. 3-merous.] A. Orders with predominant variability in number of floral parts (Orders i — 7) : a. Typically achlamydeous fls. appear (Orders i — 4). a. Fls. usu. naked. Great variability in number of sta. and cpls. 1. PANDAN ALES. Marsh herbs, or trees, with linear 1., and cpd. heads or spikes of naked, haplo- or homo-chlamydeous — i, G oo — i. Endosp. P. Naked fls. occur, but also all stages from achlam. to hetero- cnlam. fls. , and from hypog. to epig. Number of essential organs definite or not (Orders 2, 3). 2. HELOBIAE. Water or marsh pi. with scales in axils, and cyclic or hemicyclic fls. P in o, i, or 2 whorls, h >mo- or httero-chlam., hypog. or epig. A oo — i, G oo — i , free or united. Endosp. little or none. i. Potamogetonineae: fl. hypog., achlam., haplo-, or homo- chlam. (fams. i — 4). 2. Alismatitieae : fl. hypog. , usu. heterochlam.; ov. on ventral suture (fam. 5). 3. Butomineae: fl. hypog. or epig., usu. heterochlam.; ov. on inner surface of cpls. (fams. 6, 7). JRIDALES. Saprophytes with scale 1. and small long- Iked homochlam. 5 or j ? fls. P 3 — 8, valv., petaloid; A. 3, 4, or 6; ? , i stds. G oo each with i basal ov. ; oo styles. •ir-orr\ tVii^L- T^nr1r\cn TRIURIDALES. stalked sta. racemed or umbelled on short or long axis, ? of (oo — i) cpls. with sessile stigs. and oo — i ov. ; heads of berries or drupes; endosp. oily. Sparganiaceae : rhiz. herbs with 2-ranked 1. and fls. in 3 ? heads, ? heads lower. P 3 — 6, sepaloid, A 3 — 6, G (i — 2), each with i pend. ov. ; fr. drupaceous; endosp. floury. Potamogetonaceae : submerged or floating herbs of fresh or salt water, with usu. 2-ranked 1. and sol. or spiked $ or ¥ <£ reg. fls. P usually o, A 4 — i, G^ — i each with i pend. ov. ; fr. i-seeded. Najadaceae: submerged herbs with opp. linear toothed 1. and 3 ? fls., 3 P 2, A i term.; ? P i or o, G i, with i basal anatr. ov. Aponogetonaceae ; tuber-rhiz. water herbs with submerged or float- ing 1. and spikes (in caducous spathes) of $ reg. fls. P 3 — i peta- loid, A 3 + 3 or more, G 3 — 6 ; fr. leathery, seeds 2 or oo . Scheuchzeriaceae : marsh herbs with narrow 1. and racemes or spikes of 5 or rf ? reg. fls.; P usu. 3 + 3, homochlam., bractlike, A 3 + 3, G 3 + 3 sometimes united, outer often absent, i or 2 anatr. ov. in each. [Lilaea 5 3 ? , A i, G i.] Alismaceae : water or marsh herbs with rad. 1., latex, and much branched infl. of reg. heterochlam. § or 3 cpls. 5. PRINCIPES. Tree-like or woody pi., sometimes climbing, with fan or feather 1., and reg. usu. ov. ; caps.; endosp. 2. Stemonaceae: perenn. herbs with rhiz. and often climbing stem and axillary infls. of homochlam. $ reg. 2-merous fls. ; P sepaloid, G (2), i-loc. ; caps. 3. Liliaceae: herbs with rhiz. or bulbs, shrubs, or trees with infl. of usu. racemose type, of usu. homochlam. $ reg. usu. 3-merous fls. ; P or (P) 3 + 3, petaloid, A 3 + 3, G to G 3— (2— 5)-loc.; fr. various; endosp. fleshy or cartilaginous. 4. Haemodoraceae : perenn. herbs with 2-ranked 1. and simple or cpd. infl. of $ reg. or -|- fls.; P 3 + 3, A 3, G to G, 3-loc. with few ov. ; caps. viii MONOCOTS. LILIIFLORAE-MICROSPERMAE B; a; b 3. Iridineae: as last, but inner sta. aborted (fam. 9). b. Fls. homo\petaloid} or heterochlam., epig., usu. -\- (Orders 10, u). 10. SCITAM1NEAE. Trop. herbs, sometimes very large or woody, with cyclic, homo- or hetero-chlam. usu. -|- 3-merous fis.; A typi- cally 3 + 3, but often with great reduction, even to i sta., G usu. 3-loc. with large ov. ; usu. aril, peri- and endosp. ii. MICROS PERM AE. Fls. cyclic, homo- or hetero-chlam., 3-merous, typically diplostemonous, but commonly with great reduction in A, G 3- or i-loc. with oo small ov.; endosp. or o. 1. Burmanniineae: fls. usu. reg.; endosp. 2. Gynandrae: fls. always •]• ; no endosp. AMAR YLLIDA CEAE-ORCHIDACEAE ix 5. Amaryllidaceae : herbs or shrubs of various habit and cymose infl. on scape, of $ reg. or •[• fls.;_P 3 + 3 petaloid, A 3 + 3 usu. intr., often with stipular corona, G (3), rarely ^-in(., 3-loc. with oo ov.; caps, or berry. 6. Velloziaceae : herbs or shrubs with linear crowded 1. and term. sol. j? YeS- 3-merous fls. on long stalks; P petaloid, A 6 or 6 bundles, G (3), 3-loc. with oo ov. on lamellar plac.; caps.; endosp. 7. Taccaceae : perenn. herbs with tubers and large entire or cymosely branched 1., and cymose umbels of £ reg. fls. with long thread-like br. ; P 3 + 3, petaloid, A 3 + 3, G (3), i-loc. with parietal plac., 6 petaloid stigs. and oo ov.; caps, or berry. 8. Dioscoreaceae : climbing herbs with usu. tuberous rhiz. and alt. or opp. often sagittate L; fls. in racemes, homochlam., $ or 3 ¥ , reg. ; P sepaloid, usu. united, with tube, A 3 + 3, inner sometimes stds., G (3), 3- or i-loc., usu. with 2 ov. to each, styles 3 or 6; caps, or berry ; endosp. 9. Iridaceae: perenn. herbs or undershrubs with equitant 1. and term. cymose infl. of $ reg. or -I- fls.; P 3 + 3 homo- or hetero-chlam., A 3 extr., G (3), 3-loc. with 3 styles sometimes divided and leafy, ov. oo, rarely i-loc. ; caps.; endosp. IQ i. Musaceae : very large herbs with 'false' stem, or trees, with cpd. infl. with large often petaloid br. and $ or 3'1°C< with i — oo ov. in each; berry or caps.; endosp. and perisp. i. Zingiberaceae : perenn. herbs with tuberous rhiz. and lane, petiolate 1., with ligule and simple or cpd. infls. of usu. $ -|- fls.; K (3), C (3) forming tube below, A i (of inner whorl, with labellum opp. to it of 2 inner stds., and sometimes 2 outer stds.) G (3), usu. 3-loc. with oo ov. ; caps.; usu. aril; endosp. and perisp. 3. Cannaceae : perenn. herbs with large 1. and cpd. infl. of showy heterochlam. $ asymmetric fls.; K 3, C (3), A i — 5, only half of i inner sta. fertile, the other half, and rest, petaloid stds., G (3), 3-loc. with oo ov. ; caps. ; endosp. and perisp. 4. Marantaceae: perenn. herbs with 2-ranked 1. with pulvinus at end of stalk, and heterochlam. $ asymmetric fls.; P 3 + 3, A 4 — 5, only i inner half fertile, as in last, the_2 other inner and i — 2 outer petaloid (i inner usu. hoodlike), G (3), 3-loc. or i-loc. by suppression, with i ov. in each; aril; caps.; endosp. and perisp. II 1. Burmanniaceae : green or saproph. herbs with sol. or cymose fls.; p(3 + 3) or 3 + 3. A 3 + 3 or 3, G (3), 3- or i-loc. ; caps., oo seeds; endosp. 2. Orchidaceae: perenn. herbs of various form, often epiph. with pseudobulbs, and $ , i , usu. resupinated, homo- or hetero-chlam. fls.; P 3 + 3, A i or 2, united with style of G (3) i-loc. to form a column ; pollen in tetrads usu. united to pollinia, stigmas 3, the third usu. rudimentary or forming a rostellum, ovules oo ; caps.; no endosp. x DICOTS. VERTICILLATAE—JUGLANDALES A; B; a DICOTYLEDONEAE [Embryo with two cots.; stem with open bundles; 1. usu. net-veined; fl. usu. not 3-merous.] Archichlamydeae (Orders i — 30) (Fl. achlam., haplochlam., or diplochlam., usu. polypet., rarely sympet. or apet.) A. Ov. with 20 or more embryo sacs, and chalazogamic fert. (Order i). 1. VERTICILLATAE. Woody pi. of Equisetum habit; 3 fls. in catkin-like spikes, ? in heads, at end of twigs; c? with 2 median bractlike P and a central sta., ? naked, G (2) with 2 threadlike stigmas, 2-loc., the post, sterile, the other with 2 — 4 erect orthotr. ov.; fr. indeh. ; no endosp. B. Ov. usu. with only i embryo sac (Orders 2 — 30). a. Fls. naked or with haplochlam. bract-like P (Orders 2 — 12). 2. PIPERALES. L. simple, stip. or not, and spikes of small achlam. or haplochlam. ? or $ ? fls.; A i — to, G i — 4, free or united. 3. SALICALES. Woody with simple alt. stip. 1. and spikes of dioec. achlam. fls., disc cup-like or reduced to scales; A 2 — , G (2), i-loc. with parietal plac. and oo anatr. ov. ; caps, with oo seeds, seeds small with basal tuft of hairs and no endosp. 4. GARRYALES. Woody pi. with opp. evergr. 1. and fls. in catkin- like panicles, $ ? ; G (2 — i) with i ov. ; drupe or nut. b. Usti. with, sepaloid or petaloid P, rarely heterochlam. (Orders 13-16). 13. PROTEALES. Woody with alt. exstip. 1. and spikes or racemes of cyclic homo- (apparently haplo-)chlam. 2 + 2-merous5 or 3 ? reg. or -|- fls. ; P petaloid ; sta. anteposed and usu. adherent to P, G i ; fr. various, no endosp. 14. SANTALALES. Herbs, shrubs, trees, often paras., with cyclic, usu. homochlam. fls.; A anteposed, in i or 2 whorls, G, rarely G (2 — 3), rarely i, each with i pend. ov. (or ov. not differentiated). i. 'Santalineae: ov. differentiated from plac., often without integ. (fams. I — 6). BA TIDA CEAE—SANTALA CEA E xiii 9 i. Batidaceae: 10 i. Julianiaceae : II 1. Betulaceae: shrubs and trees with alt. simple 1. with caducous slips. and monoec. anemoph. fls._in catkins, typically 3 fls. per axil ; P sepaloid or o, A 2 — to, G (2), 2-loc. each with i pend. ov. ; nut; no endosp. 2. Fagaceae: trees, rarely shrubs, with simple 1. and caducous scaly slips., and usu. catkins or_ small spikes of i ? fls. ; P sepaloid (4 — 7), A 4 — 7 or 8 — 14, G usu. (3), 3-loc. , 3-styled, each with 2 pend. ov. ; nut; no endosp. 12 1. Ulmaceae: trees and shrubs with 2-ranked simple slip. 1. and axill. cymes of homochlam. $ or $ ? fls.; P 4 — 5, sepaloid, A (4 — 5) or (8 — 10), G(2), slyles 2, usu. i-loc. wilh i pend. ov.; nut or drupe ; usu. no endosp. 2. Moraceae : usu. trees and shrubs with slip. 1., lalex, and cymes of small i ? fls., oflen head-like ; P usu. 4 or (4), persistent, rarely o, A as many, opp. P, G (2), i-loc. with usu. i pend. ov. ; nut or drupe ; endosp. or not. 3. Urticaceae: usu. herbs with opp. or alt. slip. 1., no latex, and cy- mose infls. of small homochlam. usu. S ? fls., P usu. 4 — -5, A 4 — 5 opp. P, bent inwards in bud and exploding, G i-loc. with i basal ov. and i style; nut or drupe; endosp. i . Proteaceae : 14 Myzodendraceae : semiparas. undershrubs with alt. 1. and minute naked c? ? fls. ; A 2 — 3 — i with monothecous anlhers, G (3) wilh axile plac. and 3 ov.; fr. with 3 feathery bristles in angles. Santalaceae: semiparas. herbs, shrubs, trees with opp. or alt. Land small 5 or S ¥ homochlam. fls. with perig._or epig. disc ; P usu. 2 + 2 or 2 + 3, A as many, inserted on P, G i-loc. with axile plac. and i — 3 ov.; nut or drupe, i-seeded; endosp. xiv D I COTS. SANTALALES—CENTROSPERMAE B; b; c 2. Loranthineae : ov. usu. not differentiated (fam. 7). Balanophorlneae : plac. central with pend. ov. with no in- teg. ; chlorophyll-less paras, (fam. 8). 15. ARISTOLOCHIALES. Fls. cyclic, homo- or haplo-chlam, reg. or •)•; P petaloid, G usu. inf. 3 — 6-loc. with axile plac., or i-loc. \vith parietal, and oo ov. 16. POLYGON ALES. L. usu. ochreate, fls. haplo- to hetero-chlam., § reg.; G i-loc. with usu. i basal erect ov.; nut; endosp. c. P haplochlatn., sepaloid 'or petaloid, sometimes heterochlam. (Order I?)- 17. CENTROSPERMAE. Usu. herbs with spiral or cyclic homo- or hetero-chlam. _fls.; A usu. =and opp. P, but also oo — i, G (oo — i) or free, rarely G, usu. i-loc. with oo — i campylotr. ov.; perisperm. i. Chenopodiineae: P bracteoid, not > 5, Aanteposed; ovule usu. i (fams. i, 2). OPILIACEAE—AMARANTACEAE xv 3. Opiliaceae: fls. § heterochlam. with slight seam-like K; G with i ov. with no integument. 4. Grubbiaceae: trees or shrubs with opp. leathery 1. and small g reg. fls.; P 4 sepaloid, A 4 + 4, G (i), i — 5-loc. below when young, later i-loc. with i pend ov. on central plac.; drupe; oily endosp. 5. Olacaceae: trees and shrubs with usu. alt. entire 1. and small ? reg. fls.; K 4 — 6, very small, C 4 — 6, A as many or i — 3 times as many, G (2 — 5), i — 5-loc. at base, i-loc. above, with i ov. pend. into each loc.; i -seeded drupe or nut; endosp. 6. Octoknemataceae : woody with alt. 1. and $ fls.; P 2 + 3, A 2 + 3, anteposed, G i-loc. with 3 pend. ov. ; drupe i -seeded. 7. Loranthaceae: woody semiparas., usu. on trees, with usu. reg. 2 — 3- merous,_usu. homochlam. £ or S ? fls.; P in two whorls, A as many, G i-loc. usu. without differentiation of ov. and plac.; layer of viscin round seed; endosp. 8. Balanophoraceae : fleshy root paras, with tuberous rhiz. from which stems rise endog., and small fls. in spikes or heads, homochlam. or naked, usu. S ? ; P in S 3 — 4 (2 — 8), united below, A as many or i — 2; P in ¥ usu. o; G (i — 2), rarely (3 — 5); nut or drupe; endosp. 15 1. Aristolochiaceae: herbs or climbing shrubs with alt. exstip. 1. and homochlam. 5 reg. or ± fls.; P usu. (3), petaloid, A 6—36, free or united with style, G, rarely G, 4 — 6-loc. with oo ov. ; caps. ; endosp. 2. Rafflesiaceae : thalloid parasites, shoots very short with term. fl. or raceme, usu. G 3 or rnore with oo ov.; berry; endosp. 7. Berberidaceae : herbs or shrubs with simple or cpd. 1. and $ reg. homo- or hetero-chlam. 3 — 2-merous fls.; P in 2 — 4 whorls, often with 2 whorls of honey-1., A in two, G i, rarely more, with oo -^ i ov. ; berry ; endosp. w. 47 xviii D I COTS. RANALES—RHOEADALES B; d; a; ft 4. Magnoliineae: fls. with P, spiral to cyclic; ov. on ventral suture; oil cells (fams. 9 — 18). P. Syncarpy and hypogyny the rule (Orders 19, 20). 19. RHOEADALES. Usu. herbs with racemes of fls., cyclic (exc. sometimes the A), heterochlam., rarely homochlam. or apet., hypog., reg. or -|- ; G (00—2), ov. with 2 integ. 1. Rhoeadineae: fl. heterochlam, K usu. 2 (fam. i). 2. Capparidineae: fls. heterochlam., Kusu. 4 or more (fams. 2-4). 3. R&edineae: fls. heterochlam, spirocyclic (fam. 5). MENISPER MA CEAE— RESEDA CEA E xix 8. Menispermaceae: climbing shrubs with usu. alt. simple 1. and small usu. reg. i present, some with dorsal raphe and micropyle down. i. Geraniineae: fls. heterochlam. rarely apet., usu. reg. and obdipl., rarely haplostemonous and in -I- fls. usu. abor- tion of some sta.; anthers opening longitud., G iso- or oligo-merous ; ov. with 2 integ. (fams. i — 12). A. No secretory cells or passages (fams. i — 7). BR UNI A CEAE— OX A LI DA CEAE xxiii 11. Bruniaceae: heath-like undershrubs with alt. exstip. 1. and cpd. spikes, racemes and heads of usu. reg. and perig. 5 fls.; K, C, A 5, G (3 — 2) each with 3 — 4 ov. or i with i; caps.; aril; endosp. 12. Hamamelidaceae : woody with usu. alt. stip. 1. and spikes or heads of 5 ore? 9 reg., heterochlam. apet. or naked fls. surrounded by br. ; K, C, A 4 — 5, G (2) with i — oo pend. ov.; caps.; endosp. 13. Eucommiaceae : trees with alt. extip. 1. and latex, and naked i ? reg. fls.; A 6— 10, G (2), one aborting, with 2 pend. ov.; samara ; endosp. 14. Platanaceae : woody with alt. 3 — .s-lobed stip. 1. and pend. spherical heads of free or united below, G (4 — 2) with oo erect ov. on parietal plac.; caps.; endosp. J2. Tamaricaceae : shrubs or herbs with small alt. exstip. 1. and ? reg. 4 — 6-merous fls.; A" as many or twice as many as C, or oo in groups, G (5 — 2) with oo ascending ov. on basal plac.; style divided; caps.; seed hairy; endosperm or none. 13. Fouquieriaceae : shrubs with decid. 1. and thorny midrib, and ra- cemes or panicles of showy 5 reg. fls.; K 5, C (5), A 10 — 15, G (3), each with 4 — 6 ov.; seeds hairy or winged. 14. Cistaceae: herbs and shrubs with usu. opp. 1. with glandular hairs and ethereal oil, and ? reg. fls.; K.S— 3, C 5— 3— o, A oo , G (5 — 10) with oo or 2 ov. on parietal plac.; caps.; endosp. 15. Bixaceae: woody pi. with alt. simple 1. and showy % reg. fls. in panicles; K, C 5, A oo , G (2) each with oo ov. on parietal plac.; style i; caps.; endosp. W. 48 xxxiv DICOTS. PARIETALES B; d; * 5. Cochlospermineae: as last, but endosp. of kidney-shaped seed oily (fam. 16). 6. Flacourtiineae: G free on convex axis, or in tubular axis rarely united at sides to G ; endosp. oily (fams. 17 — 24). 7. Papayincae: G free in tubular or bell-shaped axis; endosp. oily; latex (fam. 25). 8. Loashieae: G sunk in and united to axis ; endosp. oily, rarely none (fam. 26). 9. Datiscineae : G sunk in and united to axis; endosp. thin, embryo oily; fls. in racemes (fam. 27). 10. Begoniineae: as last, but no endosp.; fls. in dichasia or scorpioid cymes (fam. 28). COCHLOSPERMA CEAE— BEGONIA CEAE xxxv 1 6. Cochlospermaceae: woody, usu. with lobed or cpd. 1. and showy ? reg. or -|- fls. in racemes or panicles; K, C 4—5, A oo , G (3 — 5) each with oo ov. on parietal or almost central plac.; caps.; endosp. 17. Winteranaceae : woody pi. with alt. exstip. 1. and cymose umbels of $ reg. fls.; K 4 — 5, C 4 — 5 or o, A (20 or less), G (2 — 5) with 2 — oo ov. on parietal plac.; berry; endosp. 18. Violaceae: herbs, or woody, with alt. stip. 1. and ? reg. or •[• fls.; K, C, A 5, G (3), each with i — oo ov. on parietal plac.; caps, or berry ; endosp. 19. Flacourtiaceae : usu. woody pi. with alt. stip. simple 1., and £ or i ? reg. fls.; R 2 — 15, C 10 — o, A usu. oo , G or semi-inf. (2— 10) usu. with GO ov. on parietal plac.; berry or caps.; often aril; endosp. 20. Stachyuraceae: small shrubs with alt. 1. and racemes of small £ or polyg. reg. fls.; K, C 4, A 8, G (4) with oo ov. ; berry; aril; endosp. 21. Turneraceae: herbs, trees or shrubs with alt. 1., stip. or not. and axillary or racemed or cymed 5 reg. fls. with tubular axis; K, C, A 5, G (3), each with 3 — oo ov. on parietal plac.; style divided ; caps.; aril; endosp. 22. Malesherbiaceae : herbs or undershrubs with alt. exstip. usu. very hairy 1. and racemes or cymes of $ reg. j-merous fls. with tubular axis and gynophore; A 5, concrescent with gynophore, G (3) with oo ov. on parietal plac.; caps.; no aril. 23. Passifloraceae : herbs or shrubs often climbing by tendrils with simple usu. palmately lobed 1., stip. or not, and fls. sol. or in racemes or cymes, reg., 5 or i ? , with axis often ± tubular ending in effigurations ; K, C 5, rarely 3 — 8, A usu. 5 or 4 — 8, rarely oo , united to prolongation of axis, G (3 — 5) usu. with oo ov. on parietal plac.; caps, or berry ; usu. aril and endosp. 24. Achariaceae: herbs or undershrubs with simple or lobed 1. and single fls. or few in an axil, c? ? reg., 3 — 5-merous; K, C, A 3 — 5, G as last ; caps. ; endosp. 25. Caricaceae : woody pi. with simple or cpd. exstip. 1. and axillary infls. of , G (4 — 15) united to hollow axis, with i style, 4 — i5-loc. with oo ov. ; caps, or berry-like fr. ; no endosp. 8. Punicaceae: woody pi. with entire 1. and showy axillary $ reg. fls. with top-shaped axis ; K, C 5 — 7, A oo , G (9) in superposed whorls with oo ov. united to axis, i style; berry-like fr. ; no endosp. 9. Lecythidaceae : woody pi. with alt. entire exstip. 1. and $ fls. with hollow axis ; K usu. 4 — 6, C 4 — 6, rarely more or o, A oo , ± united at base, bent inwards in bud, G (2—6) each with oo— r ov. ; style i ; fleshy or woody fr.; no endosp. 10. Rhizophoraceae : woody pi. usu. with opp: stip. 1. and usu. ? reg. fls., sol. or in cymose infls. , hypog. or epig.; K 3 — 16, usu. 4 — 8, C as many or o, A 8 — oo , G usu. (2 — 5), rarely 6, each with 2 — 4 — oo pend. axile ov. ; fr. usu. with i seed per loc.; sometimes viviparous. xxxviii DICOTS. MYRTIFLORAE-UMBELLIFLORAE B; d; £ 3. Hippuridineae : fls. epig. with i sta.; i cpl. with i ov. and no integ. (fam. 18). 4. Cynomoriineae: root paras, with epig. fls. with i sta., cpl. with i ov. with i integ. (fam. 19). 30. UMBELLIFLORAE. Fls. usu. in umbels, cyclic, heterochlam.; usu. with i whorl of sta., epig., 4 — 5-, rarely co.-merous, ? reg. ; G (5 — i) or (oo ) each with i (rarely 2) pend. ov. with i integ., rich endosp. NYSSACEAE-CORNACEAE xxxix 11. Nyssaceae : shrubs with alt. exstip. 1. and small £ or i-loc. but with only i ovule): 87. Valerianeae. 88. Dipsaceae. 89. Calycereae. 90. Com- positae. Order 3. Campanales (fl. usu. irregular; sta. usu. epig. ; ovary 2 — 6-loc., with usu. oo ovules in each loc.) : 91. Stylidieae. 92. Goodenovieae. 93. Campanulaceae. SERIES II. HETEROMERAE. Ovary usu. superior; sta. epipet. or free from corolla, opp. or alt. to its segments, or twice as many, or oo ; cpls. >2. Order 4. Ericales (fl. usu. regular and hypog. ; sta. as many or twice as many as petals ; ovary i — oo -loc. with i — oo ovules in each loc.; seeds minute) : 94. Ericaceae. 95. Vaccinieae. 96. Monotropeae. 97. Epa- crideae. 98. Diapensiaceae. 99. Lennoaceae. Order 5. Primulales (corolla usu. regular and hypog., sta. usu. = and opp. to corolla-lobes; ovary i-loc. with free-central or basal placenta and i — oo ovules) : 100. Plumbagineae. 101. Primulaceae. 102. Myrsineae. Order 6. Ebenales (corolla usu. hypog. ; sta. usu. more than corolla- lobes, or if as many, then opposite to them, except in 103, often oo ; ovary i — oo-loc.; ovules usu. few ; trees or shrubs) : 103. Sapotaceae. 104. Ebenaceae. 105. Styraceae. SERIES Iff. BICARPELLATAE. Ovary usu. superior ; sta. as many as or fewer than corolla-lobes, alt. to them ; cpls. 2, rarely i or 3. BENTHAM-HOOKER SYSTEM liii Order 7. Gentianales (corolla regular, hypog. ; sta. epipet. ; 1. gene- rally opp.): 106. Oleaceae. 107. Salvadoraceae. 108. Apocynaceae. 109. Asclepiadaceae. no. Loganiaceae. in. Gentianaceae. Order 8. Polemoniales (corolla regular, hypog.; sta. = corolla-lobes, epipet.; ovary i — 5-loc. ; 1. generally alt.) : 112. Polemoniaceae. 113. Hydrophyllaceae. 114. Boragineae. 115. Convolvulaceae. 116. Solanaceae. Order 9. Personates (fl. usu. very irregular ; corolla hypog., often 2-lipped ; sta. generally fewer than corolla-lobes, usu. 4, didynamous, or 2 ; ovary 1—2- or rarely 4-loc. ; ovules usu. oo ) : 117. Scrophularineae. nS. Orobanchaceae. 119. Lentibularieae. 120. Columelliaceae. 121. Gesneraceae. 122. Bignoniaceae. 123. Pedalineae. 124. Acanthaceae. Order 10. Lamiales (corolla usu. 2-lipped, hypog., rarely regular ; sta. as in preceding; ovary 2 — 4-loc.; ovules solitary in loc., or rarely > i in fams. 125 and 127 ; fruit a drupe or nutlets) : 125. Myoporineae. 126. Selagineae. 127. Verbenaceae. 128. Labiatae. Anomalous Fam. 129. Plantagineae. III. Monochlamydeae or Incompletae (fl. usu. with one whorl of perianth, commonly sepaloid, or none) : SERIES I. CUR VEMBR YA E. Terrestrial plants with usu. § fls. ; stat generally = perianth-segments ; ovule usu. solitary; embryo curved in floury endosp. 130. Nyctagineae. 131. Illecebraceae. 132. Amarantaceae. 133. Chenopodiaceae. 134. Phytolaccaceae. 135. Batideae. 136. Poly- gonaceae. SERIES II. MULTIOVULATAE AQUATfCAE. Aquatic plants with syncarpous ovary and oo ovules. 137. Podostemaceae. SERIES HI. MUL TIO VULA TAE TERRESTRES. Terres- trial plants with syncarpous ovary and oo ovules. 138. Nepenthaceae. 139. Cytinaceae. 140. Aristolochieae. SERIES IV. MICREMBRYAE. Ovary syn- or apo-carpous ; ovules usu. solitary; embryo very small, surrounded by endosp. 141. Piperaceae. 142. Chloranthaceae. 143. Myristiceae. 144. Monimiaceae. SERIES V. DAPHNALES. Ovary usu. of i cpl. ; ovules soli- tary or few ; perianth perfect, sepaloid, in i or 2 whorls ; sta. perig. 145. Laurineae. 146. Proteaceae. 147. Thymelaeaceae. 148. Penaeaceae. 149. Elaeagnaceae. SERIES VI. ACHLAMYDOSPOREAE. Ovary i-loc., 1—3- ovuled ; ovules not apparent till after fert. ; seed with endosp., but no testa, adnate to receptacle or pericarp. 150. Loranthaceae. 151. Santalaceae. 152. Balanophoreae. SERIES VII. UNISEXUALES. Fls. unisex. ; ovary syncarpous or of i cpl. ; ovules solitary or 2 per cpl. ; endosp. or none ; perianth sepaloid or much reduced or absent. liv BENTHAM-HOOKER SYSTEM 153. Euphorbiaceae. 154. Balanopseae. 155. Urticaceae. 156. Platanaceae. 157. Leitnerieae. 158. Juglandeae. 159. Myricaceae. 1 60. Casuarineae. 161. Cupuliferae. SERIES VIII. ANOMALOUS FAMILIES. Unisex, fams. of doubtful or unknown affinities. •• 162. Salicaceae. 163. Lacistemaceae. 164. Empetraceae. 165. Ceratophylleae. II. MONOCOTYLEDON ES (as in Engler). SERIES I. MICROSPERMAE. Inner perianth petaloid ; ovary inferior with 3 parietal or rarely axile placentae ; seeds minute, exalb. 169. Hydrocharideae. 170. Burmanniaceae. 171. Orchideae. SERIES II. EPIGYNAE. Perianth partly petaloid ; ovary usu. inferior ; endosp. abundant. 172. Scitamineae. 173. Bromeliaceae. 174. Haemodoraceae. 175. Irideae. 176. Amaryllideae. 177. Taccaceae. 178. Diosco- reaceae. SERIES III. CORONARIEA E. Inner perianth petaloid ; ovary usu. free, superior ; endosp. abundant. 179. Roxburghiaceae. 180. Liliaceae. 181. Pontederiaceae. 182. Philydraceae. 183. Xyrideae. 184. Mayacaceae. 185. Comme- linaceae. 186. Rapateaceae. SERIES IV. CALYCIlVAE. Perianth sepaloid, herbaceous or membranous ; ovary &c. as in III. 187. Flagellarieae. 188. Juncaceae. 189. Palmae. SERIES V. NUDIFLORAE. Perianth none, or represented by hairs or scales ; cpl. i or several syncarpous ; ovary superior ; ovules i — co ; endosp. usu. present. 190. Pandaneae. 191. Cyclanthaceae. 192. Typhaceae. 193. Aroideae. 194. Lemnaceae. SERIES VI. APOCARPAE. Perianth in i or 2 whorls, or none; ovary superior, apocarp. ; no endosp. 195. Triurideae. 196. Alismaceae. 197. Naiadaceae. SERIES VII. GLUM ACE A E. Fls. solitary, sessile in the axils of bracts and arranged in heads or spikelets with bracts ; perianth of scales, or none; ovary usu. i-loc., i-ovuled; endosp. 198. Eriocauleae. 199. Centrolepideae. 200. Restiaceae. 201. Cyperaceae. 202. Gramineae. INDEX TO ORDERS AND GROUPS OF HIGHER RANK. Achlamydosporeae, liii Apocarpae, liv Archichlamydeae, x Aristolochiales, xiv Asterales, Hi Balanopsidales, x Batidales, xii Bicarpellatae, Hi Calycinae, liv Campanales, Hi Campanulatae, xlviii Caryophyllinae, 1 Celastrales, li Centrospermae, xiv Contortae, xlii Coronarieae, liv Cucurbitales, xlviii Curvembryae, liii Daphnales, liii Dicotyledones, x, 1 Disciflorae, li Ebenales, xlii, Hi Epigynae, liv Ericales, xl, Hi Fagales, xii Farinosae, iv Ficoidales, Hi Gamopetalae, Hi Garryales, x Gentianales, liii Geraniales, xxii, li Glumaceae, liv Glumiflorae, iv Guttiferales, 1 Helobiae, ii Heteromerae, Hi Incompletae, liii Inferae, Hi Juglandales, x Julianiales, xii Lamiales, liii Leitneriales, x Liliiflorae, vi Malvales, xxx, 1 Micrembryae, liii Microspermae, viii, liv Monochlamydeae, liii Monocotyledones, ii, liv Multiovulatae, liii Myricales, x Myrtales, li Myrtiflorae, xxxvi Nudiflorae, liv Olacales, li Opuntiales, xxxvi Pandales, xxii Pandanales, ii Parietales, xxx, 1 Passiflorales, Hi Personatae, liii Piperales, x Plantaginales, xlvi Plumbaginales, xl Polemoniales, liii Polygalinae, 1 Polygonales, xiv Polypetalae, 1 Primulales, xl, Hi Principes, iv Proteales, xii Ranales, xvi, 1 Rhamnales, xxx Rhoeadales, xviii Resales, xx, li Rubiales, xlvi, Hi Salicales, x Santalales, xii Sapindales, xxvi, li Sarraceniales, xx Scitamineae, viii Spathiflorae, iv Sympetalae, xl Synanthae, iv Thalamiflorae, 1 Triuridales, ii Tubiflorae, xliv Umbellales, Hi Umbelliflorae, xxxviii Unisexuales, liii Urticales, xii Verticillatae, x CAMBRIDGE : PRINTED BY J. E. PEACE, M.A., AT THE UNIVERSITY PRESS