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Gre
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The Evolution of the Earth
AND ITS INHABITANTS
cA series of Lectures Delivered before the
Yale (hapter of the Sigma Xi during the
eAcademic Year 1916-1917
“Foseph Barrell
Charles Schuchert
Lorande Loss Woodruff
‘Richard Swann Lull
Ellsworth Huntington
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NEW HAVEN
YALE UNIVERSITY PRESS
London : Humphrey NGlford : Oxford University Press
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COPYRIGHT, 1918, BY
YALE UNIVERSITY PRESS
First published, 1918.
Second printing, 1919.
Third printing, 1920.
Fourth printing, 1922.
Fifth printing, 1923.
PREFACE
DuRincG the collegiate year 1916-1917, the conduct of the
Yale Chapter of the honorary scientific society of the Sigma Xi
was entrusted to my guidance, and in order to render the meet- _
ings of the chapter as profitable as possible, a symposium was
proposed on the geological and biological evidences for the
_ evolution of our planet and the earth-borne life. I therefore
asked such of my colleagues as were authorities on the several
subjects involved to prepare addresses to be delivered before
_ the chapter and later to be published in the form of the pres-
ent book. The course of lectures was as follows:
THE EVOLUTION OF THE EARTH AND ITS INHABITANTS
Lecture I. The Origin of the Earth, November 23, 1916,
Professor Joseph Barrell.
Lecture I]. The Earth’s Changing Surface and Climate,
December 13, 1916, Professor Charles Schuchert.
‘Lecture III. The Origin of Life, January 24, 1917, Pro-
fessor Lorande Loss Woodruff.
Lecture IV. The Pulse of Life, February 15, 1917, Professor
Richard Swann Lull. |
Lecture V. Climate and Civilization, April 20, 1917, Dector
Ellsworth Huntington.
The scope of the combined essays is of necessity very broad,
ranging as it does from a conception of the universe to the
trend of modern civilization. ‘Thus the first chapter deals not
alone with the genesis of the earth but of the parent solar
system, and, the earth having been established, its history ts
traced until the time of its becoming a fit environment for the
abode of life. The second lecture deals with the changing lines
vi | PREFACE
of demarcation between land and sea, the rise and growth of
continents, the formation and severance of land-bridges, and
the climatic changes which are recorded for geologic time.
The physical environment once established, Professor Wood-
ruff tells what we know and do not know of the origin of life.
This is largely an academic discussion of the several theories
which have been advanced to account for the evolution of life-
less into living matter, for from the nature of the problem
evidences of direct observation are not available. ‘The lecture
on the pulse of life attempts to link up cause and effect; to find
those forces which are responsible for the more or less
rhythmic accelerations of evolution shown by the fossil record. |
The main cause is found to be climatic change, which in turn —
has as a chief controlling factor earth shrinkage and the con- ~
sequent warping of the crust discussed in the second lecture.
The pulse of life applies not alone to the evolution of animals
and plants, but also to mankind. How climatic changes have
influenced the growth of civilization and the formation of
racial characteristics of mentality is set forth in the last lecture, —
that by Doctor Huntington. In so far as possible, these essays
are the fruits of the original research of their several authors,
which in certain instances are set forth here for the first time.
The treatment of the entire subject and the marshaling of the
facts thus assembled are entirely new.
I am deeply grateful to my, colleagues, not only for their
having accepted the tasks thus laid upon them, which in several
instances implied new and extensive research, but also for the
success with which the lectures were presented, as attested by
_ the society.
RICHARD SWANN LULL,
President, Yale Chapter, Sigma XI.
1916-1917.
Yale University,
December 1, 1917.
Preface . .
Chapter
Chapter
Chapter
Chapter
Chapter
Index
CONTENTS
ee ne enigee ak RICHARD SWANN LULL
The Origin of the Earth . . JOSEPH BARRELL
The Earth’s Changing Surface and
Climate. . . . #. + CHARLES SCHUCHERT
The Origin of Life . LORANDE LOSS WOODRUFF
The Pulse of Life . RICHARD SWANN LULL
Climate and the Evolution of Civilization
ELLSWORTH HUNTINGTON
ILLUSTRATIONS
Restorations of: A. the hypothetical proavian, B. the
toothed bird Archaeopteryx . . . . Frontispiece
A. Nebulosity in the Pleiades. B. Spiral nebula in
Ursa Major . . . . facing page
Diagram to illustrate tidal ices ‘
Origin of a spiral nebula according to Ciaihentin
and Moulton . :
The western and eastern jeateaeres Wocize dave
Diagram showing the times and probable extent of the
more or less marked climatic changes in the geologic
history of North America, and of its elevation into
chains of mountains j ane ty
Glacially striated diabase, ‘cide in pees. Penal
time. South Africa . . . . . facing page
Lower Cambrian quartzite, striated in early Permian
time. South Australia . . . . facing page
Glacial bowlder beds beneath thick formations of
dolomite on the upper Yangtse River, China. Age
of till, late Proterozoic . . . . facing page
Eroded exposure of the oldest known glacial deposits
(Lower Huronian), near Cobalt, Ontario
facing page
Striated bowlder taken from the oldest known glacial
deposits Sits Bilger g th te itor RI ae
Generalized map oe North America in Paleozoic time
The marine inundation in Pennsylvanian time .
Earlier and maximal phases of the Silurian flood
. Great Middle Cretaceous sees of the oceans
over the lands .
Diagram showing time aba extent ‘at ‘te men
floods that have inundated North America since the
close of the Proterozoic era
62
77
79
x
Figure 14.
Figure 15.
Figure 16.
Figure 17.
Figure 18.
Figure 19.
Plate III.
Figure 20.
Figure 21.
Figure 22.
Figure 23.
Figure 24.
Figure 25.
Plate IV.
Figure 26.
Figure 27.
Figure 28.
Figure 29.
Figure 30.
Figure 31.
Figure 32.
Figure 33.
ILLUSTRATIONS
Diagram of the pulse of life .
Squid, Loligo sp., 4a yagi a fish }
Fish forms . facing pt
Laurentian peneplain as seen from the south end of
Lake Michikamau, Labrador . facing page
Lung-fish, Neoceratodus, breathing .
African fringe-finned ganoid, Polypterus delhexi
Cast of the oldest known fossil footprint, Thinopus
antiquus, from the Upper Devonian of Pennsylvania
facing page
Development of the hind foot of a salamander,
Triton teniatus : AA aie
Foot of a reptile, Ranodon penne ‘ :
Restoration of the Permian stegocephalian, Cases
aspidephorus ~iadile
Diagram showing the elation ei the extta-equliry ae
membranes: A. bird or reptile, B. mammal
facing page
Skull of cynodont reptile, Nythosaurus larvatus .
Tooth of a carnivorous dinosaur, Allosaurus, and jaw
of a contemporary mammal, Diplocynodon
facing page
Restoration of archaic mammals: A. Phenacodus pri-
maevus, B. Coryphodon, C. Dinoceras facing page
Restoration of the creodont, Dromocyon .
Map showing the geographical distribution af the
primates, living and extinct, and their indicated dis-
persal from Holarctica Ut
Gorilla, Gorilla gorilla facing page
Gibbon, Hylobates lar facing page
Photograph of a submarine, twenty feet below the
surface, taken from the aeroplane whose shadow is
shown in the picture facing page
Climograph of the eth cice tn United States .
Climograph of France and Italy .
Climograph of Southern California
124
124
Figure 34.
Figure 35.
Figure 36.
Figure 37.
Figure 38.
ILLUSTRATIONS
Climograph of deaths among colored people in the
eastern United States, 1912-1915. sae hesiattay
_ diseases
Climograph of dias sa white sbonie in the eastern
United States, 1912-1915. Non-contagious diseases
The distribution of human energy on the basis of
climate Be ig
The distribution of Ecilization
Changes of climate in the abniienl « zone darling
historic times, on the basis of the growth of trees in
NE a
rigs
Ewen
ee
CHAPTER I
THE ORIGIN OF THE EARTH?
JOSEPH BARRELL
PROFESSOR OF STRUCTURAL GEOLOGY IN YALE UNIVERSITY
INTRODUCTION
THE logic of all branches of science points to the existence
of some system of evolution of the universe, its complete
nature hidden in the vastnesses of time and space, but never-
theless developed in accordance with Nature’s laws. The
earth is one of the celestial host, its beginnings are bound up
with that of other bodies. In the history written in the struc-
ture of the earth and in the relations of the earth to the planets,
stars, and nebulz lies concealed the story of its genesis. “Two
chief methods of approach, the geologic and astronomic, lead
toward the solution of this fundamental problem.
The history of the earth is read in the rocks which have
been thrust up by internal forces and beveled across by erosion.
The nearer events are clearly recorded in the sequence and
nature of the sedimentary rocks and their fossils. But the
oldest formations have been folded, mashed, and crystallized
out of all resemblance to their original nature, and intruded
by molten masses now solidified into granite and other
_ 1 Also presented before the Geological Society of Boston, January 19, 1917.
Some pages of the following article have been drawn from one by the writer
entitled “Origin of the Solar System Under the Planetesimal Hypothesis,” pub-
lished as Chapter XXV in Pirsson and Schuchert’s “Text-book of Geology,”
1915. For permission to use this material grateful acknowledgment is made to
the authors and publishers of that work.
2 EVOLUTION OF THE EARTH
igneous rocks. Fossils, the time markers of geology, if once
existent, have been destroyed, and, as in the dawn of human
history, vast periods of time are dimly sensed through the
disordered and illegible record. This crystallized and intri-
cately distorted series of the oldest terrestrial rocks tells of an
earth surface on which air and water played their parts, much
as now. But it was a surface repeatedly overwhelmed by out-
pourings of basaltic lava on a vaster scale than those of later
ages, and the crust was recurrently broken up and engulfed in
the floods of rising granitic magmas. Here the geologic
record begins, but the nature of its beginning points clearly to
the existence of a prehistoric eon. At the farther bounds of
this unrecorded time, forever hidden from direct observation,
lies the origin of the earth.
But the mind of man will not be baffled. Since he may not
see directly he will see by inference. Convergent lines of
evidence derived from various fields of knowledge may be
followed part way toward this goal, like those rays perceived
through the telescope on the. full moon near the margin of its
visible hemisphere, which converge toward craters on the side
of the moon that no eye shall ever see.
Leaving the geologic field of evidence, the problem of the
origin of the earth may be approached from the astronomic
side. The relationships of the earth to the stars and the
planets are displayed in the depths of the heavens, and vestiges
must there exist of the cosmic conditions which gave birth to
our world and the other planets of our system. ‘The forces
of nature are found to obey the same laws as far as the tele-
scope can penetrate. ‘The spectroscope detects the familiar
chemical elements in distant stars. These instruments give
assurance of the unity of the cosmos, but the diversity of
objects indicates various stages and various types of evolution.
Which approaches nearest to that of our solar system? We
must be content to study very much larger and therefore unlike
AIND ITS INHABITANTS 3
systems, since from the distances of even the nearer stars the
earth and her sister planets would be hopelessly invisible in the
most powerful telescope. We cannot, then, follow into the
planetary stage the evolution of other systems comparable to
our own. Yet in nebula, in stars, and in the inherited motions
and configurations of our planetary system are clues which
pieced together lead up toward the origin of the earth.
The problem of the origin of the earth is within the domain
of scientific investigation, but as yet the pictures which may
be drawn are varied. The vague outlines shift and change
but become clearer with the growth of knowledge. Where
the solution of a problem is not yet definitive and certain, the
method of multiple working hypotheses should be used. All
facts and theories should be matched to these several hypoth-
eses to determine which one of them shall be selected and
modified, and which shall meet the fate of the unfit. At the
present stage of investigation any one view should not be
regarded as established beyond question, even though the
assembled evidence seems strongly to support it. In a single
presentation, however, all hypotheses cannot be equally treated
and each investigator, while recognizing the existence of other
hypotheses, may properly emphasize that one which seems
to him most in accordance with the various categories of facts
and more firmly established inferences.
The hypotheses of earth origin begin more especially in
the astronomic field in the search for initial causes; they end
in the geologic field where they dovetail into the known rela-
‘tions. The surviving hypothesis must give a sound explana-
‘tion of those broader terrestrial conditions of atmosphere,
_hydrosphere, and lithosphere, of ocean basins and continental
platforms, which had become established by the beginning of
the geologic record. But, although much has been learned, it
is still unsettled among geologists as to how far those funda-
“mental conditions in early geologic times were different from
4 EVOLUTION OF THE EARTH
those of the present. On the whole, the problem of the genesis
of the earth appears to lie somewhat more in the field of the
geologist than in that of the astronomer.
THE PLACE OF THE EARTH IN THE UNIVERSE
The earth, a member of the solar system-, The earth is
but one among the planets which together with the sun con-
stitute the solar system. It is neither the largest nor the
smallest, neither nearest to nor most remote from the sun.
The sun is a star and is but one among the millions of stars,
and, though apparently so great as seen from the earth, is
mediocre among them in size and brightness. The origin of
the earth is obviously bound up with the origin of the other
planets and all in the history of the sun. A presentation of
the significant facts of magnitudes, motions, and distribution
of these bodies, familiar though they are to most readers,
should therefore precede the consideration of the genesis of
the earth. |
The planets visible to the unassisted eye are, besides the
earth, five in number, distinguished by the ancients from the
stars by their steady light and by their wanderings through
the zodiacal path in the sky, wanderings produced as a result
of the combined effect of their motions and that of the earth in
nearly circular paths about the sun, their common center. The
telescope has added to the number of planets two large ones,
Uranus and Neptune, invisible to the naked eye because of
their distance from sun and earth, and, in one zone of inter-
mediate distance, a swarm of smaller bodies, the asteroids,
better called planetoids. In size the planets sink to vanishing
insignificance in comparison with the sun or any other star.
Their distances from the sun and from each other are also
almost infinitesimal in comparison with the distances which
separate the stars. ‘They shine by light reflected from the
AND ITS INHABITANTS — 5
sun, and as all the planets and planetoids are attendant upon
the sun, they form a common system, the solar system. The
facts regarding their size, their distance from the sun, and the
inclination and position of the planes of their orbits with
respect to the orbit of the earth are tabulated as follows:
PRINCIPAL ELEMENTS OF THE SOLAR SYSTEM
Mean distance | Inclination
ee Diameter, Density, in millions of of plane of
miles water=1 miles from orbit to
the sun earth’s orbit
UME gidccc ioe a" alw'sre 866,400 1.39 Be a 7° 359
[ Mercury......... 3,030 ? 36 7 00
Mi OUR as die's cbse a's 7,700 4.85 67 3 24
fs Oe ee 7,918 5.58 93 0 00
oe | Moon...... ER 2,162 3.40 - 0.2392 5 09
DOES oii Oiein cic oes 4,230 4.01 141 $2 S52
0 to
Planetoids........ 0 to 485 ? 200 to 400 34 42
ie EE 86,500 1.33 483 £89
Major } Saturn........... 73,000 0.72 886 2 30
planets | Uranus........... 31,900 1.22 1,782 0 46
Neptune......... 34,800 2.11 2,792 1 47
1 Inclination of sun’s equator to earth’s orbit.
2 Mean distance of moon from earth.
Notable Planetary Relations. That the orderly nature of
this system implies some mode of evolution was seen by the
framers of the nebular hypothesis. The more notable of
these relations, following the summary by Young, are:
1. The orbits are all nearly circular.
2. hey are nearly in one plane (excepting the cases of
.
some of the little planetoids). :
3. The revolution of all is in the same direction.
_ 4. There is a curiously regular progression of distances
from the sun (expressed by Bode’s law, which, however, breaks
down at Neptune). See foregoing table.
6 EVOLUTION OF THE EARTH
5. There is a rough progression of density, increasing both:
ways from Saturn, the least dense of all the planets in the
system. |
As regards the planets themselves, we have: |
6. The plane of the planets’ rotation, the plane of their
equators, roughly coinciding with those of the orbits (probably
excepting Uranus).
7.. The direction of the rotation of the planets about thei
polar axes the same as that of their revolution in their orbits
(excepting probably Uranus and Neptune).
8. The plane of orbital revolution of the satellites of eac
planet coinciding nearly with that of the planet’s rotation, its
equatorial plane. -
9. The direction of the satellites’ revolution in their orbits
also coinciding with that of the planet’s rotation about its axis,
with exceptions in the case of the ninth satellite of Saturn and
probably the seventh of Jupiter. : .
10. The largest planets rotating most swiftly. :
The sun, a member of the stellar system. The sun, as has
been stated, is but a star and a member of the stellar syste
What are the orders of magnitude in number, in size, in dis-
tance, in speed, in duration among these countless orbs, and
how do these relations enter into the problem of the origin o
the earth as one of that retinue of planets which attend upon
the sun? :
The luminous stars of our system are estimated to be more
than a hundred million in number. The number of the dark
stars is unknown. Giving no ray of light to reveal their exist-
ence, they may for all we know be as numerous or more
numerous than those in the radiant stages of their existence.
The few stars whose sizes are known range in diameter from
somewhat below a million to upward of ten million miles and
more. {
Many of the stars are in reality double or multiple stars,
consisting of companions so close that the two or more appear
as one star to the naked eye, or even under the highest power
AND ITS INHABITANTS 7
of the telescope, the evidence of their composite nature being
revealed only through the analysis of their light by the spectro-
scope. Jhese double stars revolve swiftly about each other,
but such internal motions must be sharply distinguished in
thought from the streaming or drifting of the stars as parts of ©
the great stellar system. Relatively to the sun they are found
to move through space with speeds averaging between 10 and
30 miles per second, but ranging from less than 10 to more
than 200 miles per second. They do not move, however,
singly and in closed orbits, but rather in broadly scattered
groups whose paths are almost straight lines. ‘These courses
of the stars must slowly curve under the aggregate attraction
of the millions of stars, but can never return into themselves.
The paths of groups of stars intersect other groups and are
to some extent interwoven among themselves. These groups
have been found to be integrated into two greater groups inter-
meshed among each other and forming two great star streams
whose average motions are in opposite directions. With the
passage of millions of years, the stars thus continually enter
into new relations and build new configurations in the skies:
a myriad host of stellar fireflies, the living and the dead,
streaming through space hundreds of millions of miles per
year. |
Although the stars are so great in number, their distances
from each other average tens of millions of millions of miles,
those in our part of the stellar system averaging between sixty
and eighty trillions. ‘The star nearest to the sun, a Centauri,
happens, however, to be at a lesser distance of about twenty-
six trillions of miles.
To bring down the dimensions of the universe to finite
comprehension, we must divide the scale of nature by a
thousand million. Then the earth would be represented by a
pebble half an inch in diameter, circling once a year about a
sun 4.5 feet in diameter, at a distance of 500 feet. The nearest
8 EVOLUTION OF THE EARTH
star, a Centauri, would on this same scale be seen as two
spheres revolving about each other at a distance apart equal
to 2 miles, and each comparable in size to the sun. This
double star would be situated at a distance of about 25,000
miles from our planetary system with its sun, but the other
stars in this part of the stellar system would be separated
from each other on the average by more than twice this dis-
tance. The Galaxy, or Milky Way, is the cloud-like zone of
faint stars which extends as a belt around the sky. The stars
in it appear faint and close together because of their remote-
ness. They seem to constitute the outer zone of our stellar
system, and its dimensions are only vaguely known. On this
diminutive scale the Milky Way might be found to be encom-
passed by a circle of a hundred million miles diameter, or it
might be more or less. |
The nebule. All hypotheses of earth origin derive the
planets and the sun from an antecedent nebulous or meteoritic
state. The cloudy patches of light known as nebula, which
are revealed especially by stellar photography, are, however,
of several very different natures and it is a vital question as
to which, if. any, of these types, could have given birth to our
planetary system. |
First are the irregular nebule, diffuse clouds of luminol
matter, pervading whole groups of stars as in Orion and the
Pleiades, shown in Plate I, A, denser about certain stars, but
nevertheless enormously attenuated. This kind of nebulosity
is associated with certain regions of the Milky Way. From
the characteristics of their spectra, the stars in such nebule
are regarded as young stars and the nebulous matter may
represent the remains of an antecedent stage.
The planetary nebule are a distinct type, comparatively
few in number, and also found associated with the Milky Way.
They show in the telescope faint, greenish, circular discs from
which they derive their name rather than from any known
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AND ITS INHABITANTS 9
relationships to planetary systems. [hey seem to be related
in their origin to New Stars and these in turn are thought to
be produced by stars sweeping through clouds of meteoric or
gaseous matter and attaining temporarily, from the swift
impacts, an enormous brilliancy. The impact is so super-
ficial, however, that the extreme brilliance is usually lost
in a few days or weeks, and the star subsides through a
stage like a planetary nebula into a peculiar type of star
known as the Wolf-Rayet stars. The origin of the true plane-
tary nebule has not, however, been observed, as they appear
_ to possess a longer life than those which have originated in
the past few centuries from new stars.
The ring nebule are few and special, having the form of a
vortex ring.
The stellar nebule form another small group which look
in the telescope like hazy stars.
By far the greatest number of the nebulz are classified as
spiral nebula, more than 120,000 of which have been made
Sowa by photography in connection with the greater tele-
scopes. Their actual number must of course be far greater.
These objects, unlike the other forms of nebule, avoid the
Milky Way, and are scattered over regions where the stars
are more widely spaced. They are very remote and may be
entirely beyond the stellar system. This implies enormous
magnitude. It seems probable that in general they possess
high internal velocities, which implies in turn enormous
masses to generate such velocities. “These nebule possess
spectra similar to those of stars rather than, like the other
_ types of nebule, spectra of diffuse clouds of gas. Some
_ astronomers look upon them, therefore, as possibly systems
_ of stars rather than true nebule; systems so remote as to give
the appearance of faint cloud-like spirals, even when viewed
under the highest powers of the telescope. A typical spiral
nebula is shown in Plate I, B.
Bh :
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10 EVOLUTION OF THE EARTH
HyporTuHEsis OF PLANETS DERIVED FROM A PRIMAL NEBULA
The original hypothesis of Kant and Laplace. In 1754
and 1755 Immanuel Kant, the philosopher, published the most
remarkable papers which had appeared up to that time upon
the evolution of the solar system. He concéived matter to
have been originally diffused and cold. From a position of
rest it began to converge under the influence of gravitation
and gave rise to the sun. In some manner he held that the
matter in converging acquired a movement of rotation.
Certain nuclei grew up independently from the center and
gave rise to the planets and satellites. In 1785 he developed
the idea that the contraction of the sun’s mass would develop ~
its heat, a view elaborated by Helmholtz in 1854 and generally ©
held by astronomers at the present time. Thus Kant sought,
and with a large measure of success, to evolve the present state
of the universe from the simplest condition by means of
mechanical laws alone.
In 1796 Laplace, one of the most eminent of French
astronomers, published a general work on astronomy, and im
a short note at the end of the appendix proposed a theory of
the origin of the solar system which shortly became widely
known as the nebular hypothesis. He was evidently unaware
of Kant’s work published forty-one years previously. Laplace
is most noted for his mathematical work on celestial mechanics, —
yet he did not develop his hypothesis along such lines and
apparently did not attach much importance to it. Neverithe-—
less, it became the dominant idea in cosmic evolution through-
out the next century.
Laplace postulated an original nebula as a very hot, gaseous
mass extending beyond the orbit of the farthest planet and
possessing a uniform rotation throughout, as if it were a solid
body. Its size was the result of a balance between expansion
from its heat and contraction from its gravitation. As it lost
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AND ITS INHABITANTS II
heat it contracted and, with the same energy of rotation that
it possessed before, necessarily revolved on its axis in a shorter
time. At last a stage was reached where, in the equatorial
belt, centrifugal force balanced gravitation and the matter
subjected to this balance of forces could sink in no further.
It is thought to have existed as a ring, left behind by the
condensing mass. ‘he ring, however, was unstable; it broke
up and gathered into one body. During the further shrinking
of the main mass other rings were in turn abandoned. Each
gathered into a subordinate nebula, passed through an inde-
pendent evolution, and the whole gave rise to the system of
planets and their satellites.
_ Modifications of the nebular hypothesis. During the first
half of the nineteenth century the nebular hypothesis was
accepted by astronomers almost without question, but during
the second half many serious dynamical objections were de-
veloped and a process of modification began, until now not
much remains of the original conception of Laplace. A
_ rather full statement of the hypothesis and the objections to
it has been given recently by Campbell.” A briefer summary
and a citation of but a few of the modifications in the general
concept must here suffice.
George Darwin, Lockyer, Faye, Fouché, Poincaré, and
others have taken part in this work, and in the opinion of these
mathematicians and astronomers the framework of the result-
_ ing structure is still sound, though subject of course to further
modification as knowledge increases. It was shown that the
original nebula need not have been hot, but, as perceived by
Kant, would develop heat from its self-condensation. A loose
swarm of cold meteorites would suffice as well as an original
gas for the initial state. The mass could never have revolved
asa unit body, as if it were a solid. On the contrary, the inner
2 Campbell, W. W., “The Evolution of the Stars and the Formation of the
Earth.” Scientific Monthly, vol. 1, 1915, pp. 189-194.
12 EVOLUTION OF THE EARTH
parts would be condensed and revolve fast while the outer
parts were still diffuse and revolved slowly. The mode sug-
gested by Laplace for the separation of the rings is also’
dynamically very unsatisfactory. Moulton has shown that the
growth of the planets and the development of rotation in the
same direction as their orbital motion could be much better
attained from an initial state in which the component particles
revolved in the same plane but independently and in highly
elliptic orbits about the central nucleus. This is a wide de-
parture from the idea of a circular ring revolving as a unit
body. | .
Still more fundamental objections, emphasized by Cham-
berlin and Moulton, are found in certain of the existing
dynamical relations of the solar system. It would be expected
that in condensation the central mass would continually aban-
don matter from its equatorial zone, the inner planets would
presumably have possessed the greater masses, and the final
sun would now show a high speed of rotation, giving an equa-
torial diameter far greater than the polar. Such expectations
are contrary to the facts. The sun revolves so slowly on its
axis, once in twenty-five days, that it has no measurable equa-
torial bulge. In other words, centrifugal force is negligible
in the sun. Furthermore, the equatorial plane of the sun,
instead of lying precisely in the mean plane of the planets’
orbits, is inclined seven degrees to such a mean plane. :
A hypothesis to gain scientific credence must emerge success-
ful from the test of observed facts and mathematical theory.
The nebular hypothesis has not done so. It is on the defensive
and has lost standing during the past generation. Neverthe-
less, it would be premature to abandon it entirely. It has the
advantage of simplicity in that satellites, planets, and sun are
explained as the products of a single process, convergence in a
rotating nebula. But nature is often found to be complex in
her operations, so that this advantage is of doubtful weight.
AND ITS INHABITANTS 13
HYypoTuHeEsis oF PLANETS DERIVED FROM A SMALL
SECONDARY NEBULA
Distinctive features of the planetesimal hypothesis. The
planetesimal or spiral nebula hypothesis of Chamberlin and
Moulton postulates the sun already in existence from the in-
gathering of a primal nebula. It was at some later stage dis-
rupted through the tidal forces produced by the close approach
and passage of another star. The result was a secondary
nebula, but one essentially unlike the primary. The secondary
nebula was developed in a plane and initially possessed a
spiral form with the sun at its center. All of its parts moved
with freedom and independence in elliptic orbits, a point of
difference from the Laplacian hypothesis. The nebula con-
tained only a minute fraction of the solar matter, but was
endowed by the passing star with a great rotational energy,
so that, although so insignificant in mass, the planetary matter
dominates enormously over the sun in the moment of momen-
tum of the whole system. Thus the planetesimal hypothesis
is a bold and frank abandonment of the terms of the original
or nebular theory. It is too early as yet to predict what will
be the ultimate fate of this hypothesis of a secondary, and, in a
measure, an accidental origin of the planets, but, as expounded
by its originators, it must be regarded as dynamically more
satisfactory than the present form of the hypothesis of primary
origin. ‘The essential features will, therefore, be presented as
the more probable preliminary steps in the genesis of the earth.
Forces of tidal disruption. The power of stars to disrupt
each other without coming into actual contact, merely through
relatively close approach, must be understood, as it is the basis
of the planetesimal hypothesis.
The sun and moon raise terrestrial tides by virtue of the
pull of gravity and thus medify that spheroidal form of the
earth which is given by its own gravity and the centrifugal
14 EVOLUTION OF THE EARTH
force of its revolution. Suppose, in other examples of ©
heavenly bodies, the tidal pull to be many times stronger, the —
self-gravitative cohesion to be many times weaker. A limit
will be reached at which the body may be pulled to pieces. This —
phenomenon, which has been actually observed in the case of —
comets passing close to the sun, has been called tidal dis-
ruption.
=
Fic. 1.—Diagram to illustrate tidal forces.
Let M and N in Figure 1 bé two-bodies passing each other
in space, and consider the action of the larger on the smaller. —
According to Newton’s law, the bodies attract each other
directly as their masses and inversely as the square of their
distances, causing them to swing toward each other while
passing by, but soon losing influence as they separate in their —
journeys through space. Consider three particles, a, b, c, on —
the line of attraction, taking them as separate parts of the
smaller body. But a is nearer to M than is b, and b is nearer
than c. Therefore if we represent the relative attractions by ©
lines, these lines will correspond to the distances which the —
particles would move in a given time if free to obey the attrac-
AND ITS INHABITANTS 15
tion of the other body. The line at a is longer than that at b
and the latter is longer than that atc. If N was not bound
together by its own gravity or rigidity, a, b, and c would
therefore drift apart and fan out while passing M. Consider
that rigidity is negligible, as in a fluid globe; then, if a minus
b, or b minus c are quantities which become greater than the
self-gravitative force of N holding together a and c, the unity
of the body becomes destroyed. ‘The problem, however, is
not quite so simple, since the influence of all other points in
N must also be considered.
As the nearer part of the body is pulled from the center,
and as the center is pulled from the farther side, there will,
further, be two simultaneous tides of approximately equal
height, but on opposite sides of the distorted body. They
tend to be always on the line joining the two bodies. Thus, on
the earth there are two tides on opposite sides, but the revolu-
tion of the earth on its axis, like a car wheel under two oppo-
site brake-shoes, gives an apparent effect, to one on the
surface of the earth, of a revolution of the tidal wave. As
a result of the equal tides at opposite ends of a diameter there
are, on any part of the ocean, two high tides in twenty-four
hours.
Mode of tidal disruption in stars. A star is characterized
by its enormous size and mass and by the possession of a
gaseous constitution. The diameter and density are dependent
upon the balance at every point between the tremendous ex-
pansive forces of internal heat and the equally great compres-
sive forces due to its own gravity. If it contracts, then its
surface and each component shell below comes nearer to the
center, the effect of,gravity upon any shell accordingly increases
inversely as the square of the new radius, and a higher internal
temperature becomes necessary to balance the higher gravita-
tive force. From this there results the paradox known as
Lane’s law,—that so long as a body maintains a gaseous
16 EVOLUTION OF THE EARTH
constitution its temperature must rise as it contracts, even
though at the same time it is radiating heat. The temperature
of the interior must furthermore be higher than the tempera-
ture of the surface, because of the greater compression with
depth, as is illustrated in the different strata of the terrestrial
atmosphere. In those convective or slow boiling movements
which are necessary in the sun and other stars in order that
they should be able to maintain their surface radiation, there
is then a constant liberation of energy from the depths and»
a system of balanced motion which if disturbed could lead in
any star to an explosive blowing out of material from it on an
enormous scale. |
VX eee eee
The tide-generating force varies directly with the mass of —
the disturbing body and also with the radius of the body dis-
turbed. It varies approximately inversely with the cube of
the distance between the centers. The deforming force is,
furthermore, greatest in the interior because the tidal forces
acting on the zone at right angles to the line of attraction —
have a component which tends to squeeze in the points d, d’
’
:
f
i
‘
f
of Figure 1 toward the center. The gravitative control is
accordingly weakened along the line a, b, c, and is strengthened
in the directions at right angles. Now apply this’ principle
to the gaseous balanced nature of a star, and it is seen that the —
expansion in the line a, b, c is no longer exactly balanced by ©
the gravitative compression, and the unbalancing is greatest
in the center, where also is the region of highest compression
and highest temperature. The effect is as if one squeezed a
syringe bulb with orifices for exit at both ends, a bulb, however,
like an air rifle, filled with gas compressed to an explosive
degree.
Tidal disruption of the ancestral sun. The sun is occa-—
sionally observed to shoot out streams of gas, known as solar —
prominences, to heights of nearly 300,000 miles, and at —
velocities ranging up to 300 miles per second. Such phe-
AND ITS INHABITANTS 17
nomena indicate the enormous elastic and explosive energy
resident in the sun’s interior, an expansive potency held in
_ restraint by the equally prodigious power of the sun’s gravity.
Supposing then that the ancestral sun was subjected to tidal
disruption by the approach of another and possibly much more
massive star, it remains to be seen how the nebula resulting
from tidal disruption can become the embryo of an orderly
planetary system. If the matter were shot out from great
depths in the sun by its normal expansive forces plus the tidal
forces, the velocity of departure might rise high above the
observed velocities of 300 miles per second. If 400 miles or
more, it would be above the “‘critical velocity” of the sun. The
gravitative attraction of the latter could then never reclaim
that matter, because the decrease in the outward velocity due
to the solar attraction would never bring the velocity down to
zero, and could therefore never reverse the motion of the
_ escaping matter and bring it back to the sun.
It is doubtful if the sun could have drawn back to itself
material expelled with a velocity of even 300 miles per second,
for the passing star, by lowering the gravitative power of the
sun on the line passing through the two, would temporarily
decrease on that line the critical velocity. In other words, it
would help to drag matter away from the sun, even though
that matter could not catch up to the passing star, but would
be left wandering in interstellar space, forming possibly come-
tary_and meteoric material for other systems. But some,
or possibly all, of the matter of the exploded sun may have
had lesser velocities of escape and would consequently remain
within its gravitative control. In so far as it was not deflected
sideways by some extraneous force, it would fall back on the
surface of the sun as the water of a geyser falls back into its
pool. But the gravitative pull of the passing star would serve
‘as such an extraneous force, analogous to the wind which
blows part of the geyser water, as it rises and falls, to one
18 EVOLUTION OF THE EARTH
side of the basin. The matter shot out toward the passing star
would be attracted sideways after it as the star receded into
space. On falling back toward the sun it would consequently
_ pass to one side and elliptical orbits of the separate particles
would become established. The material shot out in the
reverse direction, from the opposite side of the sun, would
meet much the same conditions except that the sidewise pull ©
of the passing star would be less on it than on the sun. It is”
seen that the lateral or deflecting force acting on both arms of !
the nebula would be due to the difference between the pull of
the passing star on each arm and on the central body. The
initial spiral arms do not then represent the path along which
the material was shot out, but mark the rotation around the
central body or sun, both of the axis of expulsion during the
passing of the star and of the matter after it is expelled, as |
shown in Figure 2. .
Fic. 2.—Origin of a spiral nebula according to Chamberlin and Moulton.
The spiral nebula would be developed in a plane. That
plane is established by the hyperbolic orbit of the passing star
with the sun at the focus of the orbit. The new system would”
thus show in its nature features imposed by both its parents.
From such a nebulous fiery birth Moulton especially has _
shown how, in accordance with the laws of celestial mechanics,
a planetary system could result. .
The matter which has converged into the planets would be
that residue of the solar tidal disruption which did not pass
beyond gravitative control and did not fall back into the body
AND ITS INHABITANTS 19
ée the sun. ‘This residue is only a very small fraction of the
sun’ smass. It would appear probable, however, that the solar
disruption was very great in order to give an axial revolution
to the reaggregated matter forming the present sun, so that
lits equator should be, as observed, only seven degrees from
\the mean plane of all of the planetary orbits. The present
revolution of the sun is probably due then to the whirl pro-
duced during tidal disruption and not to an axial rotation
belonging to the sun before the event took place.
This brings us to the final stage in the evolution of the
planets according to the planetesimal hypothesis. In the arms
of the spiral nebula were knots or nuclei of matter constituting
the cores of the planets. Four small knots, the earth-moon
knot being a double one, represented the beginnings of the
four small inner planets (see table, page 5). In the zone of
the planetoids there was, however, no dominating nucleus, and
they have therefore remained to this day largely in the plan-
etesimal state. Four greater nuclei beyond were the begin-
nings of the major planets. Smaller nuclei associated with
the larger marked the presence of satellites.
The orbits of the planetary nuclei and of the scattered
\planetesimal swarm were highly eccentric, having the form
of a tangle of ellipses of all forms and sizes but lying in nearly
a common plane and with a common direction of revolution
about the central body. Collisions would inevitably occur at
the crossing of the paths in the course of numberless revolu-
jtions and the nuclei would have sufficient mass and conse-
quent gravitative power to retain the matter colliding with
\them. In this way, each planet would in the course of time
‘clear up an orbital zone, and these zones, because of the eccen-
|tricities of the orbits of the component particles, overlapped
each other with the exception of a region between Mars
and Jupiter. But Moulton has shown that in such planetary
growth by accretion, an axial revolution would arise in the
Pe
20 EVOLUTION OF THE EARTH
same direction as the orbital revolution, and that the incor-
poration of all the planetesimals would cause the eccentrici-
ties to cancel out, giving to the whole mass a nearly circular
instead of a highly elliptical orbit. This would lead us to
believe that the original nucleus was but a small part of the
completed planet, so that its original ellipticity of orbit was
submerged beneath the average influence of the added masses.
Outstanding difficulties of the planetesimal hypothesis. The
disruption or spiral nebula hypothesis explains the features
of the solar system more successfully than the older nebular
hypothesis has thus far been able to do, but there are difficul-
ties still remaining, though these may perhaps be the result of
special conditions.
The most striking departure of the real system from that
expectation deduced from the hypothesis is found in the rota-
tion of the sun. The passing of a star able to drag matter
from the sun to the distance of the planet Neptune would be
expected to lead to an enormous tidal distortion of the sun
mass. This great tidal wave would involve a lifting on
revolution about the sun, tending to give it a certain energy’
of rotation. A very little stronger action and the sun would)
in fact have been literally pulled to pieces and its matter
scattered beyond its gravitative control. It is possible that!
it may in this way have lost a part of its mass. Considerable
quantities of the expelled matter should have fallen back.
obliquely in the sun and tended further to increase its velocity:
of rotation. ‘The path of the approaching star could hav "
had no relation to the previous equatorial plane of the sun.
The probabilities would consequently be that the final rotation
would be a resultant between the older and newer forces and
lie in an intermediate plane at a considerable angle to the
plane of the planets’ orbits. Now, as a matter of fact, the
sun, as has been previously noted, revolves but once in twenty-
five days and its equator is inclined but seven degrees to “te
———i
ae ee
AND ITS INHABITANTS 21
mean plane of the planets’ orbits.. I'o explain this, Chamber-
3 supposes that the sun had originally a rotation in a plane
not greatly different from that in which the passing star ap-
roached, but rotated in the opposite direction. The whirl
ave been a little greater than its initial rotation, but, being —
ire to the solar mass by the tidal disruption is assumed to |
n the opposite direction, the resultant was at a slow speed
and yet 1 et nearly in the plane of the planetary orbits.? Campbell
points out that the chances are highly against such a special
‘arrangement. If ina number of solar systems such an arrange-
‘ment prevailed, it would constitute a conclusive proof against
‘the hypothesis, but in the one example the exceptional com-
bination may have occurred and it cannot be urged as a
disproof.*
_ Turning to another aspect of the hypothesis, the innumer-
able spiral nebule of the heavens, although good illustrations
of the initial hypothetical form of the solar planetary system,
do not appear to be stages in a similar evolution in the way
} that Chamberlin and Moulton have conceived them to be.
They are, as previously stated, of a much vaster order of
magnitude, they avoid the region where the stars are clustered,
are at remote stellar distances, and by their very number show
a notable duration of their form. On the other hand, the
postulated originally spiral form of the solar nebula would
have been evanescent. Within a century from the time of
origin all except the outer nuclei would have completed many
‘revolutions about the sun. But the different periodic times of
‘the nuclei would in a few revolutions have caused the initial
‘spiral form to disappear. It would become wound up and
Marther blended together owing to the high ellipticities of the
“constituent orbits.
8 8 Chamberlin, T. C., “The Origin of the Earth,” 1916, pp. 130-132.
| #Campbell, W. W., “The Evolution of the Stars and the Formation of the
| Earth.” Scientific Monthly, vol. 1, 1915, p. 241.
i)
—
22 EVOLUTION OF THE EARTH
The temporary dispersal of the solar mass would lead te
an enormous increase in its radiant energy. The planetesimal
matter would as a consequence of its dispersion and increased
radiating surface cool with great rapidity, except in the nuclei
of masses, and cease to be self-luminous, the smaller particles
almost immediately becoming cold except as they were heatec
by the larger and profoundly disturbed solar mass.
The new stars which have been observed are not regarded”
as of this nature since they are not expanded into a spiral. In
fact, as previously mentioned, no examples are known which
serve as illustrations, in the terms of this hypothesis, of the
birth stage of planetary systems. This lack of examples may
be connected with the small scale as well as the temporary
character of such a nebula. The whole solar system, extend-
ing to the orbit of Neptune, would subtend slightly less than
half a minute of arc as seen from the nearest star. Tne”
average star is hundreds of times farther away and at the
greater distances a nebula of this order would not betray its”
existence by its form but only by the temporary great increase
in radiance at the time of its birth. |
Chances of close approach. Whether the chance is great
or small of a planetary system being generated from any
particular star by tidal disruption cannot be used as an argu-"
ment for or against this hypothesis unless it were known what
proportion of the stars possessed planetary systems. But tha
knowledge is hopelessly concealed from us in the depths of
space, since such a system as ours after its temporary initial)
brilliance would, as shown, be invisible in the most powerful
telescopes even if it existed about the nearest star.
It is not known how near an approach would be necessary
to generate such a solar system; but the chances of close”
approach must be very small for any particular star. The
motion at any instant of any individual star is the result of it:
motion inherited from the past plus the attractions of all the
is,
Ne OE aii oe
AND ITS INHABITANTS 23
fmatter in the universe pulling upon it in the present. If two
dies without previous motion were to be attracted toward
‘each other, and were able to ignore the gravitative pull of
all other bodies, they would move in a straight line toward
‘each other’s centers with ever increasing velocity until colli-
: would result. But the least inherited motion in any
other direction, or the least deflecting pull upon one of them
more than upon the other by other bodies would prevent to
that degree a central collision, or in almost all cases any
collision whatever. Now the velocities of the stars through
‘space at great distances from each other are so great that
‘individual stars can have almost no attractive influence upon
‘each other. They must move in nearly straight lines past
each other unless they happen to pass within a thousandth
‘part of their average distance. It is seen then that the
‘chances of close approach depend primarily upon the acci-
‘dental crossing of their paths and only secondarily upon their
‘mutually attracting each other. For this reason the chance
of actual collisions may be regarded as negligible, even con-
‘sidering the vast number of stars. Approach sufficiently
near to generate strong tidal forces would, however, have
occurred during their long lifetimes as radiant bodies to a
considerable number out of the hundred million or more of
‘stars which are known to exist in the stellar system, but for
‘any one individual star, where the spacing is of the magni-
| tude existing in our part of the stellar system, the chance of
| such approach even in a billion’: years would be very small.
In fact, it has been estimated as only one chance in 1,800 in
‘that time. It is possible, however, that this happened once
to our star, that is, to our sun of that time, in the distant
“past and from that disruptive tidal force was born our
“$system of planets. If such an event was in fact a necessary
antecedent condition, fortunate indeed has been our planetary
fate, for not only did this happen so early in the sun’s career
24 EVOLUTION OF THE EARTH
that its radiant energy has been able to endure through all
the ages needed for organic evolution, but the spacing of
the stars is so wide and the chance of approach so rare that
no other of them has since advanced sufficiently near to
throw this system into disorder, or to disrupt and sweep
away the earth and its sister planets as a wasted effort, and 9
start the re-creation of a new heaven and a new earth.
HypoTHEsis OF EARTH-GROWTH BY SLOW ACCRETION
OF PLANETESIMALS |
Under the terms of either nebular or planetesimal hypoth- 7
esis a scattered state of the planetary material is implied
as a stage antecedent to the origin of the planets. Was this
growth of the planets geologically slow or rapid? Did it
take tens or hundreds of millions of years, or was it on the
contrary largely accomplished in tens or hundreds of thou-
sands of years? Was the material largely in dust-like or 7
molecular form, or was it to a large extent in nuclei of con-
siderable size? From these different postulates very diver-
gent consequences may be traced in the formative stages of ©
the earth; and finally the present nature of the earth itself 7
may speak in favor of one or the other of these views.
Chamberlin, who has been the chief writer on this sub-
ject, adopts the hypothesis that the stages of earth-growth
were very prolonged, even geologically speaking, and that 7
the accretion was dominantly of dust-like or molecular par- 7
ticles. According to him the building up of the planets fol- 7
lowed three stages: first, the direct condensation of the 7)
nuclear knots of the spirals into liquid or solid cores; second, ©
the less direct collection of the outer, or orbital and satelli-
tesimal matter; third, the still slower gathering up of the —
planetesimal material scattered over the zone between ©
adjacent planets. This third factor, in Chamberlin’s view, is ©
AND ITS INHABITANTS 25
regarded as very important and he believes this diffused
matter contributed much of the earth substance, very slowly
and in a dust-like form. This is one of the critical points
in the details of the theory upon which turns much of the
development of his following argument.
Chamberlin conceives the earth to have been built up as
a solid body, not to have been fluid or viscous at any time
later than the early nuclear stage and to have begun to
_hold an ocean by the time it contained 30 or 40 per cent
of the present mass. Such liquid rock as was generated
by compression or radioactivity during earth-growth is re-
garded as having been kneaded and squeezed to the surface,
where it solidified approximately as fast as it was formed.
In earth-growth, the denser planetesimal dust, he argues,
tended to be somewhat segregated into the primitive ocean
basins and served to maintain in them, as the earth was
built outward, a greater density than in the elevated zones
between, establishing thus a relation between density and
elevation.
It seems a debatable question if such a large proportion
of the added material was necessarily dust-like and capable
of being weathered, sorted, and distributed by the primitive
atmosphere and ocean. In fact, from this beginning of
earth-growth the preponderance of the evidence appears to
the writer to be against those sub-hypotheses which Cham-
berlin has followed. This evidence, its bearings and con-
clusions, will form the following parts of this article. It will
be of ultimate value to both lines of argument that each may
be weighed against the other.
HYPOTHESIS OF EARTH-GROWTH BY RAPID INFALL OF
PLANETOIDS
Preliminary statement. Alternative views quite different
from those which have been presented under the previous
26 | EVOLUTION OF THE EARTH
heading will now be discussed. It appears to the writer
that the chemical character of the igneous rocks, the limited
depth of density variations in the crust, the limited amount
of salt in the sea, the rotation periods of the moon and
planets,—all point to a molten condition of the earth at the
completion of its growth. In the limited space available the
more technical aspects of the arguments must, however, be
omitted. The questions raised by this conclusion are: What
mode of growth would have favored a molten state and
how far did this precede the beginning of the geologic
record, as given by the oldest rocks exposed at the surface
of the globe?
Up to this point the method of alternative hypothesis |
has been pursued, and from the standpoint of scientific pro-—
cedure it should be continued to the end. ‘The limitations —
of space in a single essay, however, forbid. This subject, -
which for complete analysis should be developed in a volume, —
must be compressed into a few pages. ‘The judicial style
must often be abandoned for the declarative. Descriptions —
of some things which no eye has ever seen will be given
graphically as though viewed by a witness. This change in
method necessitated by limitations of space is, however,
least objectionable in the closing parts of the subject, since
the foundation hypotheses have been already presented ‘and
the argument leads from them toward the established facts
of the geologic record.
Significance of the planetoids. The belt of asteroids, better
called planetoids, appears to have remained more nearly in
its original state than have other parts of the solar system.
The lack of aggregation into a planet may be due in part to
the absence of any dominating center. More than eight hun-
dred of these bodies have now been discovered and listed and
countless others must be so small that they will largely remain
unknown. The diameters range from a maximum of 485
:
AND ITS INHABITANTS 27
miles in increasing numbers down to 15 to 20 miles, the limit
of telescopic visibility.
At some diameter below the limit of visibility in the tele-
scope, although the number may be increasingly great, the
summation of their masses must begin to fall off, since other-
wise the combined bulk would produce a perceptible glow in
the sky. Furthermore, Leverrier demonstrated from the
limited perturbations of Mars in its orbit that the whole
amount of matter distributed between the orbits of Mars and
Jupiter cannot exceed about one-fourth of the mass of the
earth. It may be less. In fact, later calculations limit it to
less than one-hundredth of the mass of the earth. The rate
of increase in numbers in the smaller visible sizes suggests in
connection with the limitation in aggregate mass that a con-
siderable part, perhaps a larger part of the matter, is not in
dust-like or molecular form but is in fragments of appreciable
size ranging up to some miles in diameter. These masses,
owing to their small diameters and weak gravitative force,
would possess almost no power to grow by accretion. ‘They
must retain almost the original state of the nebula, or better,
the meteoritic swarm, and are perhaps as likely to have suf-
fered occasional shattering and scattering by impact as to have
grown from a lower order of size. Their evidence favors the
hypothesis that the scattered matter which was added to the
nucleus to form the earth was largely of such size that the
individual planetoids would have plowed through a primordial
atmosphere and ocean, if such existed, and have penetrated
beneath the surface of the liquid or solid body below. The
energy of impact from dust-like material would be absorbed
at the surface and, as heat, quickly radiated into space. The
‘accretion of dust would favor the growth of an earth solid
throughout. Larger masses would, on the other hand, carry
the energy of impact into the earth. They would not strike
with the high velocities of the meteors which collide with the
28 EVOLUTION OF THE EARTH
earth, since the different planetoids were traveling in the same
general direction, but nevertheless a state of incandescence and
liquidity would be likely to result from the sizes of the masses
involved. If in addition the infall of masses was sufficiently
rapid to bury the heat of previous infalls before it could be
dissipated by conduction to the surface, a general heating and
liquefaction of the earth would tend to take place, both from —
the increased compression of the deeper nucleus and the effects
of impact at higher levels.
The fact that the planets have cleared up the zones about
them, whereas the planetoids have remained permanently in
a scattered state, is an argument for holding that the existence
of dominating nuclei determined the growth of the planets.
It is likely that the nuclei were of various sizes, were clustered
to various degrees, and many of them united by their impact.
A somewhat limited number and considerable size of the units”
as well as their grouping would be in accord with the lack of
relation of the amount of eccentricity and inclination of orbit
to the masses of the several planets. |
Indications of primordial tidal retardation. The moon
keeps the same face turned always toward the earth. Con-
sequently, from a point in outer space, it would be seen to”
rotate on its axis in exactly the same time that it completed its”
orbital revolution. Mercury is known also to keep the same
face turned always toward the sun and the same relation is”
probably true of Venus. The other planets revolve many
time on their axes during the period of revolution, the earth,
for example, 366 times. The exact correspondence in the
moon, Venus, and Mercury between their times of axial rota-
tion and orbital revolution points.to some causal relation be-
tween the two periods. That relation is one of tidal forces.
The moon distorts slightly the earth’s figure, but as the tidal
forces due to the moon are weak and the earth is very rigid,
this distortion in figure is expressed mostly by the rise and
AND ITS INHABITANTS 29
fall of the earth’s fluid envelope. In so far as the body of
the earth yields, it is an elastic yielding which involves no
measurable expenditure of energy. The oceanic tidal waves
tend to continually face the moon and the earth revolves be-
“neath them, like a wheel revolving between two opposite
‘brake-shoes. This generates tidal friction which tends to slow
down the axial rotation of the earth. There is no question as
to the correctness of this theory, but there is a very serious
question whether the forces are not so weak as to be without
any geologic consequences, at least under the present rigid
condition of the earth’s interior.
Some of the latest work has been given to measuring directly
the retardative influence of the tides, if such exists. Mac-
Millan has made an estimate of the loss of energy by friction
of the oceanic tides. He used the formule employed by en-
gineers for the loss of head due to friction and viscosity, and
applied them to the ocean. His conclusion is that the day
would be lengthened by one second in about 500,000 years.
Even if this figure be in error tenfold or a hundred-fold it is
still in great contrast with the conclusion of Adams in the
_ middle of the last century, that the earth was losing time at the
rate of 22 seconds per century, a figure raised to 23.4 seconds
by Darwin and lowered to 8.3 seconds by Newcomb. Mac-
Millan’s method brought to bear as a retardative agency prac-
tically all the friction of the tides, irrespective of their positions
or directions of motions, and seems to show that thé water
tides do not have and have never had an appreciable effect on
_the earth’s rotation.®
_ Tidal retardation if it has ever been an important factor in
_ planetary history must then be chiefly due to a body tide. In
so far as there is a mere elastic yielding of the body no energy
® MacMillan, W. D., “On the Loss of Energy by Friction of the Tides.” In
“The Tidal and Other Problems.” Carnegie Institution of Washington, Pub.
No. 107, 1909, pp. 71-75.
30 EVOLUTION OF THE EARTH
is consumed, but a viscous drag will produce retardation.
Recent measurements of the rigidity of the earth under tidal
stresses, both by the horizontal pendulum and by the water
level in a horizontal pipe, show that the earth as a whole is”
more rigid than steel and that under the exceedingly small tidal
stresses the yielding is essentially elastic. he estimates of
viscosity are so small that they are within the limits of error
of the measurements. The smallness of the tidal strains in
the earth may be appreciated by citing Darwin’s calculations. :
According to this investigator, the tides raised by the moon -
upon the earth generate a stress-difference of 16 grams per
square centimeter at the poles, 48 grams at the equator, and
128 grams at the center of the earth. Thus the earth is
stressed by the lunar tidal forces even at the center to only
about one part in fifteen thousand of the strength which
granite has at the surface of the earth.
The tidal force exerted by the earth on the moon is about ©
twenty-two times as great as the lunar tidal force on the earth, ©
and reaches about one part in six hundred or seven hundred of
the strength of granite. If the moon were once nearer the ~
earth, the tidal stress-difference was much greater, varying
inversely with the cube of the distance. ‘Tidal retardation must
have acted efficiently upon the moon, nevertheless, until the
moon was at its present distance and the stresses reduced to
their present magnitude, in order to have reduced its period
of rotation to the same value as its final orbital period about
the earth. The action must have been that of a viscous body
tide since the moon has never been able to hold to itself an
ocean envelope. ‘The tidal force exerted by the earth upon
the body of the moon consequently must have produced a :
notable viscous yielding and continued to do this in spite of
increasing distance of the moon and increasing rigidity.
The far greater mass of the earth prevented such large
effects of tidal retardation from being felt, but its period of ©
| AND ITS INHABITANTS 31
‘ revolution compared. to those of Mars and the outer planets
suggests that the tidal forces of the moon and sun have pro-
_ duced a notable slowing down of the earth also. The largest
planet, Jupiter, 86,500 miles in diameter, revolves the most
rapidly, completing one revolution in 9 hours 55 minutes;
Saturn, the next largest, revolves in 10 hours 14 minutes.
Uranus also shows by the pronounced polar flattening of its
disc that it revolves in some similar short period. Mars, with
a diameter approximately half of that of the earth and a
_ twentieth of that of Jupiter, revolves in 24 hours 37 minutes.
These planets can never have suffered largely from tidal
_ retardation and a rough rule appears to prevail that the larger
the planet the more rapidly it rotates. Judging from its mass,
the earth may consequently have originally rotated in a period
- of between 15 and 20 hours. This argument is only of sug-
_ gestive value, but it is in accord with other lines of argument.
If the moon passed through a viscous state sufficiently pro-
longed for it to respond completely to tidal control in its
rotation period, the presumption is clearly that the earth, a
larger body and better able to retain its heat, also passed
through a similar stage of viscosity. The present rigid and
elastic condition of the earth appears then to be a secondary
feature and the present ineffectiveness of the tides cannot be
safely used as an argument against the strong indications of a
primordial tidal retardation.
Significance of the oceanic salt. Sodium derived from the
weathering of igneous rocks has been stored through all geo-
logical time in the ocean as sodium chloride. The ocean has
grown more salty since it first gathered on the earth, yet it is
so far undersaturated that sea water must be nine-tenths
evaporated before sodium chloride begins to be precipitated.
Furthermore, the indications are that it never was saturated,
even though in primordial times the sea water may have been
less in volume. Concentration to a degree which eliminates
ac > tee EVOLUTION OF THE EARTH
part of the sodium chloride raises the percentage content of
bromine, magnesium chloride, and magnesium sulphate, so
that sodium becomes subordinate to magnesium and the ratio
of bromine to chlorine is increased Subsequent dilution
would not change this ratio and the introduction of new salts
could never bring it back to the original composition. The
evidence from the sea itself is substantiated by the testimony
of the sedimentary rocks. The amount stored as impregna-
tions or as salt deposits in the sediments is quantitatively
negligible, either as compared to the volume of the sediments
or the mass of the oceanic salts. Salt deposits, furthermore,
so far as known, began to be present only in the Paleozoic, in
the later half of geologic time, the great masses of earlier
sedimentary strata being barren of them.
It has been calculated that the total sodium in the ocean
would be derived from the weathering and erosion over all
the earth of a mantle of igneous rock of average composition
only 2,300 feet thick, corresponding to 6,500 feet as the
average thickness of erosion if restricted to the area of the con-
tinental platforms, including the lands and extending out to
a depth of 600 feet below sea-level.
Daly has noted the significance of these facts upon the
hypotheses of earth-growth.* Chamberlin supposes an ocean
to have existed for long geologic ages upon the surface of
the earth during its growth from a body about half of its
present diameter and one-eighth of its present volume. The
planetesimal material, he holds, was weathered and sorted
into lighter and heavier portions, leading to the development
of lighter protuberant and heavy depressed areas. The
limited quantity of salt in the sea, however, is distinctly against
such a hypothesis of oceanic antiquity and continental build-
ing. The amount of erosion in evidence where the older
rocks are exposed as crystalline masses and from the great
® Daly, R. A., “Igneous Rocks and Their Origin,” 1914, pp. 159, 163, 164.
AND ITS INHABITANTS 33
uplifted tracts of later times is apparently about sufficient to .
account for all the salt in the sea. In fact, the estimates of
_ erosion through known geologic time based on the nature of
rock exposures and the thickness of sediments have fully
equaled or exceeded the amount given by the quantity of salt
in solution. Weathering, erosion, and the accumulation of
_ salt had therefore played no considerable part previous to the
time recorded by the oldest rocks. ‘The earlier physical con-
- ditions must have been very different from those which later
_ prevailed.
Ao
FAVORED HyYPoTHESIs OF AN EARTH INITIALLY MOLTEN
Indications of a primordial molten state. The indications
_ of primordial tidal retardation and the limited amount of salts
in the sea both point to the cgnclusi was molten
at the completion of its growth. A molten state suggests a
rapid earth-growth due to an original clustering of the matter
whose convergence built up the planet. Larger nuclei hun-
dreds of miles in diameter and smaller ones able to the
planetoids moved in elliptic and nearly intersecting orbits.
~_Mutual perturbations kept modifying these orbits and provid-_
ing new chances for collisions, union, and growth. Such colli-
sions led to_a development _of energy of impact suthcienf fo
_produce in the growing earth a molten state, at least in the
_ outer portions. The earth kept growing at the same time
—by sweeping up large quantities of finer material, but a molten
_state suggests that the greater growth was due to the intall
“ot Targer nuclei. Finally, but one outstanding nucleus, the
“moon, was left beside the earth, and the earth-moon system
attained a condition of stability and completed growth.
If the composition of the earth as a whole is similar to
that of the meteorites, those samples of matter which come to
us from the heavens, the most abundent-material in the deep
body of the earth is metallic iron. Now the blast furnace
34 EVOLUTION OF THE EARTH
/ i have el
_ makes familiar the fact that slag is insoluble in iron and,
being lighter, gathers in the upper part of the crucible, like
cream upon milk. The slag is similar in composition to
basaltic 1 igneous rocks. The density of the deep interior sug-
ests that it is layered like the crucible of the blast furnace —
and that the silicate rocks form an envelope some hundreds
of miles thick, grading down into a great metallic core. The
silicate envelope ultimately differentiated further, resulting
in a rise of the more siliceous and lighter fraction into an outer _
layer, perhaps 50 to 75 miles in thickness. This in turn crys-
tallized into a primordial, universal, granitic crust above a
thicker basaltic shell below. |
The primordial atmosphere. Granting the conclusions in |
regard to the initial fluid state of the earth, let the primordial
atmospheric conditions be pictured. A gaseous envelope exist-
ing in equilibrium with rock magma would be dominantly
water-vapor, followed in order of importance by carbon
dioxide and carbon monoxide, chlorine and hydrochloric:acid, —
with some nitrogen, but no free oxygen.
The present atmosphere of nitrogen and oxygen penetrates
by solution into the liquid envelope of water. The primitive -
atmosphere of water-gas penetrated in the same manner by —
solution into the ocean of molten rock. But such an admixture —
of molten rock and water-gas is known to have a remarkable —
effect upon the melting point of silicate magmas. Under dry
fusion the exalted temperature of 1300° to 1500° C.,
dazzling white heat, is necessary to make granite thinly molten.
But if the fluid rock has dissolved an abundance of water- —
vapor the mixture stays fluid until it has cooled below a tem-
perature of 800° C. ‘The surface of the earth when molten
was perhaps no hotter than this. At such a comparatively
low temperature and even at somewhat higher temperatures
there would be but little dissociation of water into its com-
ponent gases, and the earth would be capable of holding to
' AND ITS INHABITANTS 35
Tisclf, even in its molten stage, an envelope of water in the
form of a deep and heavy atmosphere of water-gas. Such an
envelope, including also an abundance of carbon dioxide and
carbon monoxide, formed an effective thermal blanket, pre-
nting a precipitous chilling one freezing at the surface of
the ocean of molten rock.
_ The effectiveness of the bis depended upon the pecu-
arity of both water-gas and carbon dioxide in being opaque
E the slow vibrations of dark heat, absorbing these near
the bottom of the primitive atmosphere and reradiating them
from higher levels as long, slow heat waves. Strong convec-
tion currents carried up these heated gases from the super-
heated base to the higher levels of the atmosphere. There the
uilling condensed the water-vapor into a thick and universal —
canopy of cloud, boiling up like thunder-heads from below,
shedding continuously a downpour of acid rain, rain dissipated
again into vapor as fast as the drops fell into the deeper and
hotter strata of the atmosphere. The intensity of the vertical
convection maintained a high electric tension. Incessant
flashes of lightning linked as with living, fiery tentacles the
cloudy heavens to the lurid molten earth. Tremendous re-
verberations of thunder, unsensed by mortal ears, shook the
atmosphere in the worldwide primeval storm.
_ The sunlight of the Primordial Era illumined and was
teflected from the outer side of the mantle of cloud. The
planet shone brilliantly by this reflected light, having an ex-
ternal appearance similar to that which Jupiter and Saturn
still possess. Above the zone of cloud the carbon dioxide and
other gases, with very minor amounts of water-vapor, ex-
tended with diminishing density as an upper transparent
envelope.
_ During the more rapid growth-stages the molecular and
dust-like matter swept up by the earth settled like a never
ceasing cloud of volcanic ash. The planetesimals of sand and
36 EVOLUTION OF THE EARTH
gravel size were swept up by the earth many millions of times
more abundantly than are meteors at the present time. Those.
meeting the earth with the higher velocities were consumed by
impact. Over the hemisphere of night the otherwise invisibl
atmosphere above the cloud canopy scintillated with incessant
flashes of light and trails of luminous dust. Bodies of larger
size gave in their dissolution a still more brilliant display and
penetrated to greater depths. At longer intervals, with titanic
rush and roar, a greater projectile, tens or even hundreds of
miles in diameter, cleaved through the canopy of cloud, leaving
a tumultuous maelstrom behind, drove almost unchecked
through the deep and dense atmosphere below, and, with
worldwide commotion, was engulfed, with development of
fervid heat, within the molten sea. |
THE PASSAGE OF THE MOLTEN INTO THE RIGID EARTH
Density stratification by fractional crystallization, The
central parts of the earth were compressed during the growth
stages by the increasing load above. This compression
developed heat, but also raised the fusion point and made for
a greater rigidity. It is not known, consequently, whether
during earth-growth the center tended toward a liquid or
solid state. The outer part, however, with a thickness of
perhaps the outer quarter of the radius, comprising about one-
half of the volume of the sphere, seems to have passed into
a truly molten condition. | |
The heavy atmosphere and canopy of cloud prevented a
rapid radiation from the molten surface, probably sufficien
during the highly liquid stage to prevent a crusting over of
frozen rock. The method of solidification approached neare
to that which occurs in a large reservoir of magma intruded
into the crust than to the freezing of a modern lava stream in
contact with the air.
AND ITS INHABITANTS 37
_ At last the rapid generation of heat by impact lessened, and
the fluid sphere, seething with slow convection currents, began
‘to cool. Certain compounds in the mutual solution of rock
elements became insoluble and fractional crystallization was
initiated. ‘The heavy basic crystals were the first to form:
crystals of metallic sulphides, magnetite, hornblende or
pyroxene, and olivine. These crystals, because of their high
‘specific gravity, tended to work downward in the convective
movement. At first they were dissolved again in the abyss,
but as time went on they remained undissolved and accumu-
lated in the deeper parts of the fluid zone. The remaining |
magma was more siliceous, of lighter gravity, and in crystalli-
zation gave to the upper shell a higher proportion of feldspar
and quartz. The original crust of the earth was consequently
a granite. [he process of fractional crystallization may,
‘however, not be a sufficient explanation. An immiscibility of
the complex mineral solutions may have developed upon the
lowering of the temperature. In one way or the other or by a
combination of several causes a density stratification is sug-
gested by a number of lines of evidence as existing in the earth.
The conclusion, then, is more than a mere inference from
theories of crystallization.
The gathering of the ocean waters. At last the ocean of
molten rock had shallowed, crystallization went forward in
separate basins, convection became hindered, the surface
froze as in a lava'caldron. Then rain, ever descending from
the shield of perpetual cloud, but never heretofore reaching
the bottom of the atmosphere, at last began to splash on the
hot surface of the earth. The raindrops at first were dis-
‘sipated by contact and sent flying back as scattered molecules
of gas. But, owing to the low conductivity of rocks, the tran-
sition stage was very brief, and perhaps even in a few thousand
years from the time when the crustal congelation of the earth
had taken place a permanent ocean of acid water began to
oo
EVOLUTION OF THE EARTH |
fest upon the surface. For a while the balance swung, as one
section or another of the crust was broken through and lavas
would pour out abundantly. Rapidly, however, from th
geologic standpoint, as the surface cooled, the atmosphere of
water-vapor condensed in a never ceasing deluge until an
ocean, probably universal in its extent, had gathered to a meat
depth of several thousand feet. The remaining atmospher
was comparatively rare and cold. Carbon dioxide became thi
dominant gas, and water-vapor subordinate. Solar heat begai
to play the principal part in warming the equatorial zone. /
system of planetary winds developed in accordance with the
new order of nature. The cloud canopy became thin and
broken, resolving itself into climatic belts. Sunlight for the
first time began to pierce the lower atmosphere and illumin
from without the surface of the earth.
During the earlier time, when the water could exist only a
gas in the atmosphere, the great pressure of this envelope ha
kept much, perhaps most, of the gases in the molten rocks
With the great fall in atmospheric pressure which accompaniet
the gathering of the ocean, magmas which broke through the
higher levels of the crust into the regions of this decreased
pressure were able to give off great volumes of gases which it
the earlier stage had been repressed. These gases, freed foi
the first time, are termed juvenile and from this time forwar
juvenile waters were added to the ocean. In the first age
following the solidification of the earth the additions wer
large in volume, but igneous action continues to bring ney
magmas to the surface recurrently from age to age. Thes
give off the gases which have been repressed in them sine
the origin of the earth. Thus, in intermittent and lessenec
rate, the surface waters have increased through geologic time
As Suess has said, the body of the earth has given forth its
oceans. :
a i
AND ITS INHABITANTS 39
THE ORIGIN OF OCEAN BASINS
_ The relations of crustal density to ocean basins. The fluid
‘earth had a surface as level as the ocean, and the process of
‘solidification which has been outlined does not account for
those marked variations in density and in surface form which
‘are expressed by the outer crust of the solid earth being divided
into continents standing high above the ocean floors. A sketch
of the formative period is therefore not complete unless the
processes are briefly discussed which are thought to have
‘shaped the earth’s surface, giving rise to the existence of lands
even before the period of the oldest known rocks.
Reasons will be given below for holding that the ocean
basins have been formed by subsidence of broad areas of the
crust, owing to the weight of magmas of high specific gravity
rising widely and in enormous volume from a deep core of
greater density into these portions of an originally lighter
crust. This regional subsidence was especially characteristic
of primordial times, but the process did not wholly cease then;
since certain lines of evidence suggest that some ocean basins
have been extended in later geologic ages, breaking into once
wider continental platforms. ‘The resultant increase in the
volume of the ocean basins has led to a drawing off of the.
ocean waters from the continental areas, and a marked diminu-
tion of the shallow seas of earlier ages.
The cause of the continued generation of new bodies of
molten rock in the sub-crustal shell, adequate to account for
the observed results of later geologic time, is thought to Ife
in the slow accumulation of heat from radioactivity in these
depths below the crust. ‘This is discussed in the subsequent
topic on the rise of basic magmas.
Some very thick bodies of intrusive rock are observed to
be more dense and basic below, lighter and more siliceous
above. The lower part is a gabbro, whereas the upper may
40 EVOLUTION OF THE EARTH
be a granite. The separation has taken place after the in-
trusion of the magma, the denser material sinking, the lighter
rising. ‘There are indications that the process goes forward
ona larger scale also, a scale so large that the dark and heavy
base is never seen, erosion of later ages being restricted en-
tirely to the granite zone. Such a splitting in composition is
indicated in that the earlier intrusions in a period of igneous
activity are intermediate or basic and the later products are
vast bodies of granite. The greater density in the earth’s
interior suggests a primal density stratification on even a larger
scale, which has been discussed under a previous topic.
But in the outer shell, 50 to 75 miles thick, the density is
far from being uniform. In recent years it has been proved
by means of precise geodetic measurements on the local in-
tensity of gravity and deflections of the vertical that the crust.
beneath the continents is notably less dense than that beneath
the oceans. The most of this difference in density exists prob-
ably within the outer 50 miles. The continents stand high,
consequently, for the same reason that an iceberg rises above
the surface of the sea: it is the position of equilibrium. At a
certain depth the downward pressures given by the thicker
continental and the thinner oceanic crust are the same and a
condition of hydrostatic equilibrium prevails in the sub-crustal
shell.
This condition of equal pressures at a certain depth is called
isostasy. It is not inconsistent with a solid and rigid condi-
tion of the earth, but does mean that at a greater depth,
apparently from 50 to 300 miles or more, hot but solid rock
can slowly yield and flow by recrystallization. ‘The process
is physically the same as that by which a glacier flows under
the slight stimulus of an almost level surface slope. The condi-
tion for such ready recrystallization is found in temperatures
which are close to those of fusion. At such temperatures
molecules under strain pass readily from the solid to the liquid
‘sa1aydstwiay U19}sva pue UIIIsAM BY —"T] ILVIg
AND ITS INHABITANTS 41
state, shift into positions which ease the strain, and again enter
into a crystalline solid condition. The proof that such a pro-
cess exists in the earth is based on several lines of evidence.
First, evidence of a broad isostatic equilibrium notwith-
standing the agencies of mountain folding, of erosion, and of
sedimentation, all of which work through geologic time tend-
ing to destroy those relations of elevation which are needed
to maintain isostasy, giving equal pressures by broad crustal
areas of unlike density upon the yielding zone below.
_ Second, the evidence of increasing temperature with depth,
giving temperatures close to those of fusion at depths below
40 to 50 miles. |
_ Third, the evidence from tides and earthquakes that the
earth as a whole is more rigid than steel and cannot possess a
fluid shell beneath the crust.
Fourth, the physical principle that at temperatures close
to fusion a crystalline substance is incapable of supporting
permanent shearing stresses, but yields slowly by recrystalliza-
tion, notwithstanding the fact that under short stresses the
same substance may be as rigid as steel.
_ The conclusion to which this argument leads is that an outer
crust or lithosphere, the rock sphere, 50 to 75 miles thick and
very strong, is marked by broad variations in density amount-
ing to as much as 5 per cent, and more local variations up to
IO per cent, which correspond to the broader relief of the
earth’s surface. Below this lies a thick, hot, basic, rigid yet
weak shell which the writer has named the asthenosphere,"
the sphere of weakness. The problem of the origin of the
ocean basins and of continental platforms resolves itself con-
sequently into the origin of the density differences in the
lithosphere and the maintenance of the heated and weak con-
dition in the asthenosphere.
TBarrell, Joseph, “The Strength of the Earth’s Crust.” Jour. Geology, vols.
22, 23, 1914, 1915.
42 EVOLUTION OF THE EARTH |
Rise of basic magmas from the asthenosphere. ‘The series
of radioactive elements slowly break down into elements of
lower atomic weight and give off in the process enormous quan-
tities of energy. Uranium, in degenerating through radium to
the stable element lead, develops more than a million times |
the heat given by the combustion of an equal weight of coal,
but the disintegration of the element and the liberation of its
heat are so slow that the whole duration of geologic time has”
not sufficed to eliminate uranium from the crust of the earth.
Therefore it has acted as a permanent generator of heat in
the rocks which contain it. |
Uranium and thorium, the parents of the radioactive series,
are widely though sparsely diffused through the lithosphere.
It has been calculated that, if they extend in their surface
amount to a depth of 40 miles, they must supply heat to the
surface as fast as it is lost by radiation into space. The earth
therefore appears not to be growing colder, though losing heat.
The small content of’ radioactive elements in the basaltic
shell below the granitic crust of the continents would then
supply that slow increment of heat which is necessary to
generate new molten rocks. The granitic shell above, though
somewhat richer in radioactive elements, is sufficiently near
the surface to lose its excess heat by conduction. ‘The excess
heat generated in the asthenosphere is, on the contrary, so”
deeply buried that it cannot escape in that manner but must
slowly transform some of the solid rock into liquid form.
Reservoirs gather, until their mass, combined with their de-
creased density in the fluid form, enables them to work their
way through the crust above and demonstrate their existence
in igneous activity at the surface of the earth. The magma
which thus comes from the greatest depth and in greatest
volume would, because of the initial density stratification,
produce a notable increase in the density of the outer crust.
AND ITS INHABITANTS 43
In order to reéstablish isostatic equilibrium such a region must
subside.
Most of the igneous rock of later geologic ages which has
been intruded into the outer continental crust clearly has aot
_ increased the density sufficiently to produce a foundering and
would appear therefore either to have come from somewhat
higher levels or to have risen in lesser quantity. In some
~
regions, however, as in that of the Lake Superior basin, large
masses of basic magma do seem to have overweighted the crust
in an early geologic period and produced a tendency to settle
as a basin. The same effect may have taken place to even a
larger degree in some regions of notable subsidence, as in the
Mediterranean basins. In the earliest times, following the
_ solidification of the earth, the forms and relations of the ocean
basins suggest that dense’ molten matter from the depths of
the earth broke into or through the outer crust, on a gigantic
scale, eruption following eruption until the widespread floods
had weighted down broad areas and caused their subsidence
into ocean basins. |
As seen in the lava plains of the moon, such an action, once
started at a certain point, is conceived to have gone forward
with widening radius, leading to the origin of the many rudely
circular outlines characteristic of the ocean basins. The
process left great angular segments of the original lighter crust
as continental platforms standing in relief between the coales-
cent basins. The waters gathered into the basins and the
continents emerged.
THE REIGN OF SURFACE PROCESSES AND BEGINNING OF THE
ARCHEAN
It is possible that shallow ocean basins began to form nearly
as fast as the waters gathered, tending to maintain some land
areas above the level of the primordial sea. Or the lands
may have emerged later, as the ocean basins spread and deep-
44 EVOLUTION OF THE EARTH |
ened. With the separation of the lands from the seas, ero-
sion began, carbon dioxide was abstracted from the atmosphere )
to make carbonates, and a further cause of atmospheric
depletion was initiated. Thinner, rarer, and colder grew the ©
gaseous envelope, until an oscillating balance was established —
between the supplies of new gases from the uprising molten —
rocks and the losses involved in the weathering of their solid
forms. Nitrogen was at first relatively small in quantity and
oxygen not present in more than a trace. An evolution in
atmospheric composition had still to go forward through ©
following ages to transform it into a gaseous medium for the
support of the higher life. But even in the early periods
following the gathering of the oceans and the emergence of the -
lands, the sun warmed the atmosphere and earth. An environ- |
ment suitable for the lowest organisms had arisen and the
earliest forms of life may not have been long in coming into
existence. The reign of the surface processes had begun, but,
at age-long intervals, the still youthful energies of the interior ©
broke forth. Magmas in great volume ascended, now seen as ~
the most ancient granite-gneisses. In the great crustal over-—
turning of these earliest revolutions the foundation rocks ap-
pear to have been everywhere destroyed. The oldest rocks pre-
served are mashed and crystallized sediments and lava sheets
resting as fragments of a cover on the reservoirs of younger
magma. Such sediments, altered and intruded, are the
oldest Archean rocks. It is not known how close they stand
in point of time to the formative processes whose description
has been attempted. With these oldest rocks, the dimly
known, heroic and mythical eon of the earth is closed and the
first historic eon opens as the remote and long enduring
Archean division of geologic time.
CHAPTER II
THE EARTH’S CHANGING SURFACE AND CLI-
MATE DURING GEOLOGIC TIME
CHARLES SCHUCHERT
PROFESSOR OF PALEONTOLOGY IN YALE UNIVERSITY
UNIFORMITY OF NATURE. The previous lecturer in this course
had to seek for the probable origin of the earth in far-off space
among the stars, examining them through the great telescopes
of our time, through the light-sensing chemical plate of the
photographic camera, and through the even more wonderful
spectroscope. With this knowledge in hand, postulate upon
_ postulate has been tried out through that talismanic study,
mathematics, and so through astronomy and the science of
numbers there is revealed an earth evolution still hazy, to be
further established through geodesy, mechanics, and chemistry
before the geologist comes to be its interpreter. “Then, hand
in hand with the geologist, the paleontologist, or student of
ancient life, reveals the organic hordes that have gone on and
through whose fossil remains is unraveled the history of the
earth. In all of this we see the brotherhood of the sciences
_ and the fundamental postulate of the uniformity of nature.
The geological time-table. As some geologic terms must
be used in this lecture, it is desirable to define them here. The
geologic history of the earth is now divided as follows:
Present time.
_ Psychozoic era. Age of man or Age of reason.
ncludes the present or ‘‘Recent time,’”’ and the time
during which. man attained his highest civilization,
estimated to be probably less than 30,000 years.
46 EVOLUTION OF THE EARTH
Geologic time.
Cenozoic era. Age of mammal dominance.
Glacial or Pleistocene time. Last great ice age.
Late Cenozoic or Pliocene and Miocene time. Rise of
higher primates: apes and man.
Early Cenozoic or Oligocene and Eocene time. Rise
of higher mammals, including primates.
Mesozoic era. Age of reptile dominance.
Cretaceous period. Rise of archaic mammals.
Comanchian period. Rise of flowering plants and —
higher insects.
Jurassic period. Rise of birds and flying reptiles.
Triassic period. Rise of dinosaurs and primitive mam-
mals.
Paleozoic era. Age of fish dominance. .
Permian period. Rise of reptiles. Another great ice —
age.
Pennsylvanian period. Rise of insects and first time —
of marked coal accumulation.
Mississippian period. Rise of marine sharks. 4
Devonian period. First known amphibians and marine
fishes. |
Silurian period. First known land floras.
Ordovician period. First known fresh-water fishes. .
Cambrian period. First abundance of marine fossils, —
and dominance of trilobites.
Proterozoic era. Age of invertebrate dominance. An
early and a late ice age.
Archeozoicera. Originof protoplasm and of simplest life. —
Cosmic time.
Formative era. Birth and growth of the earth out of
the spiral nebula of the sun. Beginnings of the atmos-—
phere and hydrosphere, and of continental platforms —
and oceanic basins. No known geological record.
Origin of the sun and earth. Professor Barrell, in the pre-
vious lecture, pointed out that the sun is a star, and one of the”
smaller among the countless millions seen through the tele-
scope. It may have had its origin in a diffuse nebula like
Orion, and, according to the disruption hypothesis of Cham-—
AND ITS INHABITANTS 47
berlin and Moulton, had a long antecedent history before it
gave birth to its planetary system. ‘Therefore before there
_ was an earth, the sun had been condensing and moving undis-
turbed in space for countless ages, until it came within the
influence of another and a greater star. The gravitative
influence of this mighty passing intruder acted catastrophically
upon the sun and caused its partial disruption. As a conse-
quence, from opposite sides of the sun there streamed rapidly
outward materials of its own mass that soon arranged them-
selves into a two-armed luminous spiral nebula. The mass
which was so extruded, and which did not fall back again into
_ the sun, was less than 1 per cent of the sun’s mass, and yet it
_ gave rise to the planetary system which encircles the parent
body. From this we also the more readily appreciate the
great size of the parent sun, whose volume is 1,300,000 times
greater than that of the earth.
Growing earth during the Formative era. In the solar
nebula there existed eight knots of more or less loosely aggre-
gated denser and hotter matter, the nuclei of the future minor
and major planets and their satellites, and these in the course
of time attracted to themselves most of the surrounding nebu-
lous material, usually spoken of as the planetesimals. The
earth-knot, it is estimated, may have had an original diameter
of about 5,500 miles, and the moon-knot perhaps half its
_ present diameter of 2,162 miles. In the course of cosmic time,
the earth and moon, revolving as companions about the sun,
gathered into themselves the planetesimals that lay within
_ their spheres of influence.
The speaker holds to the postulate of Barrell that the plan-
_ etesimals conceived by Chamberlin to have been of the size of
sand and dust were probably for the most part much larger
and more like the planetoids in dimensions. From time to
time, as one or more of the planetoid-like bodies, singly or in
combination, plunged in upon the earth with velocities up to
48 EVOLUTION OF THE EARTH
tens of miles per second, their impact with and engulfment by
the earth engendered much additional heat. In fact, they
liquefied themselves and their earth surroundings as a result
of their great impacts. Furthermore, the high internal heat
of the earth was accentuated by the condensation accompany-
ing the increase of mass from an original earth diameter of -
5,500 miles to one initially larger than 8,100 miles. At no time, |
however, does it appear that the earth was necessarily in an
astral condition, shining like a star. :
The present diameter of the earth is 7,918 miles, but at the
close of the growing period it must have been 200 and possibly
even 400 miles greater, for it is well known to geologists that
throughout geologic time it has been losing volume, due in.
part to the loss of heat into space, but probably in greater
degree to internal molecular rearrangements. It is now known ©
that the earth does not shrink all the time because of these -
changes, but does so only periodically, and at these times the »
surface of the planet becomes wrinkled, giving rise to more or
less high ranges of mountains. Even though the earth has _
been dissipating its inherited and derived energy for at least
some hundreds of millions of years, and in so doing repeatedly
giving birth to new series of mountains, our mundane sphere ©
is still far from having attained the internal stability that will, -
when achieved, result in a featureless earth, an ie
devoid of the carbon dioxide that is the basis of life, and a
universal lifeless ocean—the facial expression which the earth |
will have in its old age.
Volcanism during the Formative era. While the earth was :
internally very hot and viscous, the body-mass was in slow
movement, with the heat ever rising toward the surface. |
These movements simulated a slow boiling process, and the
upwellings of the molten matter repeatedly broke through the
outer cold shell and engulfed it. As the cold crustal masses
descended, they liquefied, and in expanding brought about
|
:
—
|
:
| AND ITS INHABITANTS 49
increased pressures that again found relief in the rising tongues
‘of heated matter. This condition is believed to have continued
for a very long time, and to have been particularly active
during the last half of the Formative era and the first era of
_ geologic time, an age that may have. endured for one-eighth
to possibly one-fourth of the earth’s entire history. The
boiling process not only disseminated the internal heat, but also
‘brought about an irregular sorting out of the heavier and
lighter substances, so that the more basic and metalliferous
materials as a rule sank to ever lower levels, while the less
heavy acidic ones rose as granitic rocks toward the surface of
‘the earth. Finally, at the close of this first era, the Archeo-
zoic, the lithosphere or outer rocky and more or less rigid
shell may have had a thickness of 50 miles. Through such a
_ rocky mass little of the internal heat rises to the surface, and
life would be impossible on the earth if it were not for the
constant flow of the sun’s radiant warmth, made equable and
usable by the atmosphere. These conditions were already
present in the Archeozoic era.
Origin of continents and oceanic basins. It is well known
to geodesists and geologists that the continents are built of
lighter materials, essentially of granites, while the greater
Oceanic areas have the heavier basaltic rocks beneath them,
and that the difference in specific gravity amounts to about
3 per cent. We have seen that these differences came about
seemingly as a result of the internal boiling process that pre-
vailed during the Formative era and the Archeozoic, and so
it appears that the higher continental masses and the depressed
Oceanic basins came into being very early in the history of
the earth. The same interrelation of masses is also to be
seen in the moon.
This is the igneous theory of the origin of oceanic basins
and continental platforms of Barrell, and it holds that the
differentiation took place after the earth had attained its com-
50 EVOLUTION OF THE EARTH
plete growth, in other words during the closing time of the
Formative era and during the Archeozoic era. ‘The differ-
ences between the densities of the lands and oceans are held ©
by Barrell to be only skin deep—in the outer one-fiftieth of —
the earth’s present diameter, or in the outer shell of about —
150 miles in thickness.
On the other hand, it is true that the lands throughout
geologic time ‘have repeatedly gone down as well as up, but
the sum of their movements has been upward,- and for this —
reason geologists speak of them as the positive areas of the
earth’s surface. In the same way, the lower-lying fields have
also risen locally at times, but their movement has in the
aggregate been downward, bringing about the increasingly
greater oceanic basins, the negative areas, by far the largest
of the earth’s surface. We shall see later on that the oceanic”
fields have become larger at the expense of the lands to
accommodate the ever increasing volume of water, and it~
therefore naturally follows that the continents in former geo- |
logic times were considerably larger than they are now, pos-
sibly even 25 per cent larger. But the newer geology no™
longer holds to the theory that the oceans and lands have
repeatedly changed places; quite the contrary, we agree with
Dana that the present positions of the land and water areas
have been more or less permanent throughout geologic time.
Origin of the atmosphere. It was not so long ago that
most geologists held with the French astronomer Laplace
and with Dana that the earth was originally very hot, and that
all of the water of the hydrosphere, the salts of the oceans,
and the carbon dioxide of the air and oceans had been parts
of the primal gaseous envelope. Not only this, but that the
latter also included all the carbon locked up in the rocks,
estimated to be 30,000 times greater than the amount now in
the atmosphere. Accordingly, as the earth was held to have
been cooling throughout its history, it was thought that only
AND ITS INHABITANTS 51
recently, geologically speaking, was the present thin condition
of the atmosphere brought about.
_ Even though more than sixty years ago a few English
geologists who were familiar with the geology of India and
Africa began to point out that the earth had passed through
-acold period toward the close of the Paleozoic era (see Figs.
4and 5), this discovery did not at once break down the teach-
ings of Laplace and Dana. When, however, still older glacial
periods began to be discerned in the stratified rocks and espe-
cially in the oldest ones near the beginning of the Proterozoic
era (see Figs. 7 and 8)—a record that is remarkably well
preserved in Canada—the final straw was at hand to break
down the theory that the climates of the earth had passed
progressively from an astral through a torrid to a temperate
condition, along with a slow clarifying of the atmosphere (see
Figs. 3-8).
_ The newer geology holds, with Chamberlin, that the grow-
ing earth was originally too small in mass to hold an atmos-
phere and accordingly there was a time when our planet had
|mone. As it grew in diameter, the earth was more and more
| able to hold an atmosphere and a hydrosphere, and during
| the long growing period there was finally originated an atmos-
| phere probably not much unlike the present one, which in a
| roughly approximate way may be said to be composed of
| four-fifths nitrogen and one-fifth oxygen. It should, however,
| be noted here that at first the atmosphere had but little oxygen.
| Since life came to be, using the carbon and freeing most of
the oxygen of the carbon dioxide of the atmosphere, the
quantity of oxygen has more or less steadily increased, though
| at all times much of it has been consumed in the oxidation
| processes of the rocks and minerals.
| The most constant accessory constituent of air is carbon
dioxide, one of the three fundamental materials at the basis
| of life. In the present atmosphere there are about three vol-
52 EVOLUTION OF THE EARTH
umes of this gas to 10,000 of air, and there is as much more
in living things as there is in the atmosphere. On the other
hand, there is in the oceans of today, according to F. W
Clarke, the geochemist, from 18 to 27 times more carbor
dioxide than in the air (Johnston and Williamson say that
at 15° C. there is about 70 times more), while the still vaster
volumes locked up in the sedimentary rocks and in the fuel
and carbonaceous deposits of the earth are computed to be
30,000 times greater than the volume in the present atmos
phere. These facts are brought forward at this time to shoy
that the constituents of the atmosphere have always variec
because of the constant loss of carbon dioxide and oxygen to
the sedimentary rocks, but that at the same time there ha:
always been a resupply of carbon dioxide through the e
active volcanoes and the mineral springs, and of oxyger
through the life activities of plants. Holmes says:* “Ever
now, the outer 70 miles of the earth’s crust would be competent
to supply all the nitrogen of the atmosphere, the water of tl
oceans and the vast quantity of carbon dioxide represented br
limestones and carbonaceous deposits.”’
It has been well said that if volcanism should cease it would
not be long before the existence of life would be impossible
because of the absence of carbon. We should add here that
if there were again as much life as there is at present, all the
carbon of the atmosphere would be in the living plants ane
animals, and, if such a condition were possible, death woulc¢
come to them all. Therefore life and its abundance at an}
time are conditioned by the amount of this gas present in th
atmosphere. a
Climates of the past. The quantity of carbon dioxide i
the atmosphere is to a certain extent also a climatic regulator,
though the greater factor in this matter is water-vapor, whicl
is also the most variable constituent of the atmosphere. Witl
1 Holmes, A., “The Age of the Earth,” 1913, p. 30.
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54 EVOLUTION OF THE EARTH a
a greater abundance of both, and especially of aqueous vapor,
there results a thickened atmospheric blanket that not only a
holds in more of the earth’s warmth but takes up greater
amounts of the sun’s radiations, and when there is less of this j
gas and of water-vapor the climates are thinner and cooler.
The amount of carbon dioxide and water-vapor in the atmos-
phere has varied much throughout the geologic ages. “a
Huxley in his interesting book Physiography has well saic
that without the sun ‘“There could neither be rain nor rivers.
. Rain is dependent for its distribution upon ae
the atmosphere, but these currents are due to disturbances of
equilibrium which are brought about by means of solar heat.
Without the sun, then, there could be no winds.” The currents
of the sea in their final analysis are also due to the heat of
the sun. Therefore through the radiant action of the sun,
water-vapor is added to the atmosphere, and the amount of
it is at present and always has been the latter’s most variable
constituent, and, as rain, is of the greatest importance geologi-
cally. Clarke states? that water “is not merely a solvent anc
disintegrator of rocks, but it is also a carrier, distributing othet
substances and making them more active. To the circulatio n
of atmospheric moisture we owe our rivers, and through them
erosion is effected. The process of erosion is partly chemical
and partly mechanical, and the two modes of action reénforet
each other. By flowing streams the rocks are ground to sand
and so new surfaces are exposed to chemical attack. On thi
other hand, chemical solution weakens the rocks and render
them easter to remove mechanically. . . . It is through the
agency of rain or snow that the atmosphere produces its grea
est geological effects. . . . Aqueous vapor dissolves and es
centrates the other ter eeuicnts of air, and brings them to
ground in rain.’ a
: k
2 Clarke, F. W., “The Data of Geochemistry.” U.S. Geol. Survey, Bull. 49
1911, p. 48.
Fic. 4.—Glacially striated diabase, made in early Permian
time. Vaal River north of Kimberley, South Africa.
Photograph by W. M. Davis.
Fic. 5.—Lower Cambrian quartzite, striated in early
Permian time. Inman Valley, South Australia.
Photograph by T. W. E. David.
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a AND ITS INHABITANTS 55
_ Weare living at a time when the earth has marked climatic
differences, varying between icy polar climates and hot moist
__ or dry tropical conditions. This has, however, not always
been the case, for not long ago geologically the temperature
of the entire earth was even colder than it is now, although
| most of the climates of the past have been warm and fairly
| equable the world over (see Fig. 3). ‘The ancient plants and
animals are “self-registering thermometers” with regard to
the climates of the past, and they indicate that warm climates
persisted during long geological ages, and that even though
there were at all times zonal belts and fluctuations in the tem-
perature, the polar areas were usually inhabited, as is now
well known, by plants and animals of kinds that were adjusted
‘to winterless environments. [hese temperature fluctuations
were greatest during the closing and opening stages of the
periods and eras.
The very long warm times were separated by short periods
of cool to cold climates. Geologists now know of seven
periods of decided temperature changes (earliest and latest
Proterozoic, Silurian, Permian, ‘Triassic-Lias, Cretaceous-
Eocene, and Pleistocene), and of these at least four (those in
italics) were glacial climates). Cooled climates occur when
the lands are largest and most emergent, during the closing
stages of periods and eras, and cold climates nearly always
exist during or immediately following the times when the
‘earth is undergoing most marked mountain making (see
Fig. 3).
Origin of the earth’s waters. ‘There was a time when the
earth was too small in mass to hold a hydrosphere, the
envelope of water that lies beneath the atmosphere and above
the rocky surface of the planet. As the earth grew in mass,
it became more and more possible for it to have standing
bodies of water and clouds of water-vapor floating with the
winds of the atmosphere. All of this water, the newer geology
56 EVOLUTION OF THE EARTH
thinks, came forth out of the earth itself, through volcanic
activity and thermal springs. Even though some of the sur+
ficial waters vanish through soaking into the cold earth-shell,
their volume has increased throughout geologic time; the
greatest amount was added during the later part of the long
growing period of the earth and during the Archeozoic era,
when from 25 to 50 per cent of the present volume is believed
to have come into existence. The rest has been added during
subsequent geologic time.
Source of the salts of the oceans. It is well known that the
seas and oceans are salty, and that on the average there are
3.5 pounds of saline matter to every 100 pounds of marine
water, and in each 100 pounds of sea salts there are nearly 78
pounds of sodium chloride or our table salt. Clarke has esti-
mated that if all of the saline matter of the oceans were con-
centrated, and the volume placed on the United States, its
surface would be completely covered to a depth of 1.6 miles.
As all of this saline matter has been leached out of the rocks
of the dry lands since the earth has had rains, and as very little
of it, comparatively, has been taken out of the ocean by the
accumulating rocks, it has been further estimated that it repre-
sents the breaking down of a mass of average igneous rock
equal to at least 6,900 feet in thickness over all the continental
platforms. Probably it is more correct to state that the con-
tinents have suffered erosion of igneous rocks amounting to
between 1 and 2 miles of average depth. Of course all
erosion throughout geologic time was far greater, perhaps,
as Barrell states, from 50 to even 100 per cent higher. It
included the reworking of older materials, igneous and-sedi-
mentary. Furthermore, ° ‘more than a half, perhaps four-
fifths, of the erosion of igneous rocks was accomplished before
the beginning of the Paleozoic” (Barrell).
Various computations have been made as to how long it
has taken the salts in the oceans to accumulate and the best
ge ne Ca oe
Fic. 6.—Glacial bowlder beds beneath thick formations of
dolomite on the upper Yangtse River, China, in latitude
31°. N. Age of till, late Proterozoic. Photograph by
Bailey Willis. Published in ‘‘Researches in China,”’
Publication No. 54, Vol. I, 1907, of the Carnegie
Institution.
Fic. 7.—Eroded exposure of the oldest known glacial deposits (Lower Huronian),
near Cobalt, Ontario. Photograph by A. P. Coleman.
Fic. 8.—Striated bowlder.taken from the oldest known glacial deposits.
Photograph by A. P. Coleman.
ee a ,
:
AND ITS INHABITANTS 57
: of the calculations arrive at about 100 million years. This
~ figure is based on the postulate that the present rate has been
‘continuous throughout geologic time; this rate, however,
‘appears to be excessive, and renders the postulate erroneous.
The rate was highly variable during geologic time, as will be
-seen from the evidence throughout this chapter, so that the
Tatest calculation of Holmes*® gives the time as between 210
and 340 million years, with the warning that these high figures
“must not be supposed to possess any serious value.
a For the present we can only conclude that our knowledge of
the part played by sodium and chlorine in the constant redis-
3 Bee ution of the materials of the earth’s crust is still lamentably
imperfect, and that quantitative deductions drawn from it
~ must be regarded as being purely provisional.”
Origin of the sedimentary strata. ‘The great variability of
geologic climates also leads to variations in the power of the
atmosphere to weather and break down the high places of
the lands. The earliest atmosphere, far richer in carbon
monoxide and carbon dioxide than subsequent climates, was
a most active weathering agent in dissolving and breaking up
the oldest rocks of the lithosphere, the igneous basaltic and
granitic rocks. Then, too, there had not yet developed a
vegetation on the most ancient lands to hold the fast-forming
soils, and these conditions made it inevitable that all of this
loose and solvent material should be rapidly washed by the
rains and rivers into the oceanic basins. Therefore the strati-
fied or water-laid rocks accumulated during the earlier geologic
eras more quickly than in the later ones. On the other hand,
the erosive power of both air and water has varied much
_ throughout geologic time because of the periodically varying
height of the land and the ever changing humidity and tem-
"perature of the air. Accordingly, the rivers carry their loads
of sediments for long times very slowly and at others much
% Holmes, A., op. cit., pp. 74-75,
58 EVOLUTION OF THE EARTH
more quickly. If the erosion on the land is rapid and the
rivers are not long, the land débris is apt to be coarser in grain
and less well assorted, but of whatever nature the waste of —
the lands is, the rivers finally deliver it to the seas and oceans;
and here the materials are subjected to a long process of re-—
working and assorting into muds and sandstones. As all of
this work is done in waters, as the character of the sediments ©
varies, and as the wave-power is variable with the winds from .
day to day, it follows that the materials must be laid down in
horizontal sheets of different kinds of rock that make up the —
strata of the stratified rocks. From this it is apparent why
the sedimentary rocks of the earth’s outer shell not only are .
of different kinds, but why they so often occur in more or less -
long cycles of deposits that begin with coarse sandstones or
even conglomerates and pass upward into the much finer-—
grained mudstones and finally into limestones. The limestones
are in the main the deposits due to the chemical activities of :
organisms or the accumulating debris of plants and animals.
Geologists therefore speak of limestones and chalks as organic
deposits. So throughout the geologic ages, cycle upon cycle”
of sedimentation succeeded one another, the débris of suc-
cessive ranges of once majestic mountains, and in this orderly
sequence of deposits lies buried much of the life of the past.
The United States Geological Survey stated in 1912 that
the surface of the United States is being removed by solvent
denudation at the average rate of .o013 of an inch a year,
or I inch in 760 years. This means a removal by the streams
of over 270 million tons of dissolved matter and 513 million -
‘tons of suspended matter annually. In other words, there is”
delivered into the seas and oceans each year from the area |
of the United States alone 783 million tons of rock materials.
The amounts removed from different drainage basins show
interesting comparisons. In respect to dissolved matter, the
southern Pacific basin heads the list with 177 tons per square
AND ITS INHABITANTS 59
mile per year, the northern Atlantic basin being next with 130
tons. The rate for the Hudson Bay basin, where the topog-
raphy is nearly flat and the rocks are mainly granites, is lowest,
with 28 tons.
‘Holmes states* that the known denudation rates vaiy be-
tween I foot in 400 years (Irawadi basin) to 1 foot in 47,000
years (Hudson Bay region). For North America the rate
for solvent denudation is 1 foot in 30,000 years, and for
~ mechanical removal 1 foot in 12,000 years. ‘Taking both
——————— tlie te
together, the average rate of denudation is found to be 1 foot
in 8,600 years.”
The ancient life is known to us in the countless fossils of
the rocks, for they are the remains of plants and animals
living on the lands and in the seas at the time when the sedi-
ments were accumulating. Most of them, like the shellfish,
lived on or in the bottom of the seas, other active forms swam
about in the water, while the remains of the plants and animals
of the lands were drifted by the streams into the marine areas,
where they eventually sank to the bottom and came to be
covered over—buried—by the accumulating sands, muds, or
limestones. It should, however, be added that but little of
the life of any time is preserved as fossils, for much of it is
too soft to be preservable, or it is eaten or destroyed by other
organisms or reduced by the solvent powers of bacteria and
the acids of the waters.
The record of the physical forces which automatically en-
tomb the successive upwellings of life is found upon all the
continents, but is woefully incomplete in any one. For many
cycles the marine record is best in North America, at other
times in Europe or the Mediterranean countries or .in Asia.
The land or fresh-water record is even more scattered and
imperfect, with the longest deposition in Africa, a .continent
that, south of the Sahara desert, has been invaded but little
_*Holmes, A., of. cit., p. 57.
60 EVOLUTION OF THE EARTH
by the ancient oceans. Out of all these widely scattered
records the geologist pieces together the geological column,
embodying the history of the earth, and yet at best we seem
to have recovered far less than half of it. The record may
be complete in the oceanic basins, but here it is forever hidden
from hammer and mind. However, if we piece together all of
the thicker known geological formations of sandstones, mud-
stones, and limestones into a superposed sequence, we get a
pile of about 53 miles in thickness as a mean estimate, with
the maximum thicknesses attaining to over 67 miles (Sollas
gives a maximum of 63.5 miles). This means the more or
less rapid wearing away almost to sea-level, one after another,
of more than twenty ranges of mountains like the present
European Alps or the American Rockies. During the in-
credibly long intermediate times, when the lands were planed
to a low relief, there was very little erosion.
Ratios of muds, sandstones, and limestones. We have seen
that the continents have lost through atmospheric erosion a
layer of igneous rock between 1 and 2 miles thick in the
course of geologic time. ‘This amount of eroded average
igneous rock should theoretically resolve itself into 30 per cent
of solution materials and 70 per cent of detritals, or 80 per
cent of mudstones, I1 per cent of sandstones, and 9 per cent
of limestones. Holmes gives 70 per cent of shales (20 per
cent quartz), 16 per cent of sandstones (75 per cent quartz),
and 14 per cent limestones (75 per cent calcium carbonate).
However, actual observations of the stratified rocks of Amer
ica and Europe show quite different percentages for these three
categories of water-laid sediments, and the difference is dut
to a great increase of pore-space in the fragmented material
and to the addition of material extracted out of the atmos
phere and hydrosphere during the weathering processes
Accordingly, Leith and Mead*® have shown that the cubica
5 Leith, C. K., and Mead, W. J., “Metamorphic Geology,” 1915, pp. 59-97.
AND ITS INHABITANTS 61
content of average igneous rocks thus increases in volume by
at least 28 per cent, and that on the continents we appear to
have 48 per cent of mudstones, 32 per cent of sandstones, and
20 per cent of limestones. This marked difference between
‘chemical theory and actual presence is further explained by
the unknown quantitative loss from the continents of the finest
muds, sands, and solvent materials that the lands have per-
manently contributed to the oceanic basins. On the basis of
the circulation of radium, Holmes estimates that the loss from
the lands to the deep-sea deposits is about one-thirtieth of all
the material eroded, i.e., about 300 million tons of the 9,000
| million tons annually.
_ Distribution of the sedimentary rocks. We have seen that,
according to theory, all of the continents have lost a layer of
original i igneous rock that is on the average between 1 and
2 miles in thickness, and it is this material, removed and
reworked time and again by the weathering processes and the
| further chemical and assorting agencies of the rivers and
| Oceans, that has gone into building the known geological
column, with its maximum summation thickness of 67 miles.
| In no one place, however, can be seen more than a small
| part of this record, for usually the local thickness is under 1
| mile, though there are limited regions where as much as 20
miles of it is present. This is because the deposition of the
geologic formations in the periodically rising and flooding
| oceans, as described later in this lecture, takes place at any
| given time in very limited areas, most commonly, as at
| present, along the margins of the continents, or in long and
| Marrow troughs—the geosynclines (see Fig. 9)—within the
continental borders, and only periodically over the wider inner ~
portions of the lands. We may add that over the surface of
one-third of North America the rocks are all of igneous or
crystalline character, that. over more than one-half of it the
thicknesses of the sedimentary rocks are under 1 mile, and
LW
4
»
Dp
A
eset ts,
LANDS, GEOSYNCLINES,
AND EPEIRIC
SEAS OF N. AMERICA.
CHARLES SCHUCHERT
@CALE OF STATUTE MILES
100 0 | 200° 400 600
positive elements, and in lightest shading the extensive neutral medial area of
continent. ‘The two heavy black lines indicate the Cincinnati and Ozark (
kakee) uplifts. From the Pirsson-Schuchert ‘“Text-book of Geology,”’ publishe¢
by John Wiley & Sons, Inc.
—
AND ITS INHABITANTS 63
that over the remaining small portion they vary from 1 to
20 miles; yet the greater depths do not cover more than one-
eighth of the continent, and they occur in the long and narrow
troughs of sedimentation, of which the two best known lie in
the Appalachian and Rocky Mountain regions (see Fig. 9).
_ Eras of geological time. Now let us see how geologists
divide this great pile of at least 53 miles of sediments and
_ their included fossils. (A synopsis of geologic time is pre-
| sented on page 46, and in Fig. 3.) The first era, or Archeo-
‘zoic time, with the oldest known strata, has no proved fossils,
though a very low form of water-living alga appears to have
been present. Even if we exclude the probable fossil evidence,
the presence in Ontario of 18 miles of sediments, of which
about one-half is impure limestone along with much graphite,
indicates unmistakably that life was already in existence. As
"we are treating of the oldest known time in the history of the
earth, this is the place to emphasize what results were then
attained and what natural processes were at work.
At the very beginning of the Archeozoic era, the earth had
‘a rocky and fairly stable exterior known as the lithosphere, an
atmosphere and a hydrosphere; protruding continents, and
oceanic basins that were filled with slightly saline water; twice-
daily tides, wear and tear of the ocean waves against the
| lands, reduction of the high lands by the atmosphere, and,
through the rains, washing of the soil and solution materials
into the seas and oceans. ‘The sun shone then as now and,
made life possible, and the man in the moon was as clearly
developed as he is today, remaining so ever since because this
satellite has had neither atmosphere nor water to wash away,
his face. We therefore see that the earth was then very much
‘as it is now, with these great differences, that at first the
atmosphere was almost devoid of free oxygen and probably
richer in carbon dioxide, that the lands were not covered with
verdure, and that what there was of life was of a very low
64 EVOLUTION OF THE EARTH
order and possibly more prevalent on the lands than in the
oceans.
Then follows the second era of geologic time, the Protero-
zoic, with a thickness in south-central Canada of nearly 14
miles of coarse sediments almost devoid of fossils, and 4 miles
of lavas. Probably more than three-fourths of this mass is
of fresh-water and volcanic origin, and less than one-fourth
may be of marine origin. In the Rocky Mountains there are
upwards of 7 miles of less coarse sediments that include about
1.5 miles of possibly marine limestones. In this connection it
should also be noted that one of the remarkable recent dis-
coveries in geology is that we know very little of marine Pro-
terozoic sediments (in the sense of marine and fossiliferous
Paleozoic formations) and that there is as yet even no hint
as to where geologists are to look for them.
The Archeozoic and Proterozoic eras have the most ancient
strata of the earth and their combined thickness is 32 miles as
against about 21 miles for all subsequent, fossiliferous strata.
In this we discern the astounding fact that at least one-half
of geological time and the greater part of the earth’s history
had passed before organisms became sufficiently endowed with
hard parts to be abundantly preserved as fossils in the sedi
mentary rocks. It also means that the evolution of life for a
incredibly long time was exceedingly slow in rising from the
simple unicellular forms into the multicellular forms of plants
and animals. It may be that the Archeozoic was entirely
occupied in originating only the unicellular plants and animals,
In the Proterozoic, organic differentiation appears to have
gone on faster, because at the very beginning of the Paleozoic
era are found all of the main kinds of marine animals other
than fishes. The subject of the origin and progress of life
from the unicellular forms to the most complex modern verte’
brates will be developed by Professors Woodruff and Lull 1 in |
subsequent lectures. 4
AND ITS INHABITANTS 65
The third era of geologic time, the Paleozoic, is based upon
a thickness of about 4.75 miles of muds and sands and 3.4
miles of limestones. These thicknesses are in the Appalachian
“region, and form one of the most complete of all known
Paleozoic records. In the Cordilleran area, where the record
is far less complete, there are about 2 miles of coarse deposits
and 3 miles of limestones. Toward the medial regions of the
continent the thick lateral sections thin down to about 1 mile,
essentially of limestones.
Shortly after the opening of the Paleozoic appear the fishes,
_ the first of the vertebrates, and now, surprising as it may seem,
in the fresh waters of the land and not inthe seas. Before half
_ of the Paleozoic is gone, double-breathing is perfected, and
‘the amphibia begin to people the dry lands, and long before
the close of the era give rise to the reptiles. It is also in the
_middle Paleozoic that the most primitive land floras arise,
and not long afterward, in the Coal Measures, their remains,
falling into great swamps, create the world’s greatest supply
of fuel in the soft and hard coals (see Fig. 10).
The fourth era, known as the Mesozoic, has, in western
North America, about 7.5 miles of sediments, of which not
more than 1.25 miles are limestones. On the other hand, in
the lands bordering the Gulf of Mexico and chiefly in Mexico
there are less than 3 miles of Mesozoic sediments, but here
the limestones make up more than one-half of the total
thickness.
The Mesozoic is the Age of Reptiles, and yet the little
mammals and the toothed birds are storing up intelligence and
strength to replace the reptiles when the cycads and conifers
shall give way to the higher flowering plants. Then follows .
the Cenozoic era, the Age of Mammals, with a thickness in
California of about 5 miles of coarse sediments, contrasting
with about 4 miles of fresh-water detritals in the Rocky
Mountain area, and in these rocks occurs a most wonderful
66 EVOLUTION OF THE EARTH
array of modernized animals. The era closes with a very
decided glacial climate and another marked change in the
animal world, out of which man rises to his present primacy
among organisms. Geologists therefore call the present time
the Psychozoic era or the Age of Reason.
Fic. 10.—The marine inundation in Pennsylvanian time. Coal
swamps in partially marine areas indicated by broken lines.
From the Pirsson-Schuchert ‘“Text-book of Geology,’’ pub-
lished by John Wiley & Sons, Inc.
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68 EVOLUTION OF THE EARTH
Relative duration of the eras. The preceding table gives
the generalized thicknesses of the sedimentary rocks of North |
America in terms of the detritals (muds and sands) and the
solution materials (limestones and dolomites). To attain
more clearly to the relative duration of each of the eras it is”
desirable to restate the probable time taken by the far more |
slowly accumulating solution materials in terms of those of
the detritals. Geologists differ greatly as to the time required
to deposit any kind of sediment, but most of them will agree
that limestones take from four to ten times longer than the
coarser materials. On the other hand, the rate of deposition
is exceedingly variable from place to place and from time to
time. Further, the limestones of pre-Cambrian time are on
the average far less pure than those of the later eras. Ac-
_ cordingly, on the basis of per annum rate of sedimentary accu-
mulation there are as yet no reliable estimates, and as we are
just beginning to appreciate that the geologic record is very
incomplete, any percentages given at present must be taken as
only suggestive of geologic time. A far more reliable stan-
dard is that of the rate of disintegration of radioactive
minerals, though here the estimates of geologic time, of not
less than 1,600 million years, appear to the writer to be
excessive by 50 per cent.
Now let us compare the above results with the ratios ob-
tained by other geologists, but before doing so it should be
said that their statements relate almost always to post
Proterozoic time. If, however, we follow Dana’s ratios, it
is clear that he held in his famous text-book that about one:
half of geologic time lies back of the Cambrian. It is on this
basis that the following table has been prepared, and th
figures in parentheses restate the ratios in percentages of al
geologic time.
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70 EVOLUTION OF THE EARTH
Periodic reélevation of the lands. We have thus far pre-
sented the sedimentary evidence in the main as if it had con-
tinuously and regularly formed, without stoppages or “‘breaks.”
The geologic record, however, was not laid down in this way,
because the surface of the earth does not remain stationary.
Even though to humanity the “everlasting hills” appear to
be permanent features of the earth’s surface, they are all
doomed to be carried away by the rain and transported by the
rivers into the seas and oceans. Erosion of the land goes on
until all is worn to near sea-level, and then there is but a very
sluggish run-off of the rain that falls upon the planed con-
tinents. The earth has repeatedly had such a low relief; in
fact, this has been the condition during the greater part of
its history. .
The continents are repeatedly reélevated in relation to th 2
strand, and this goes on many times in a small way and less
often on a large scale (see Fig. 3). We have said that in the
course of the geologic ages the earth has shrunk at least 200
and possibly 400 miles in diameter. The earth is still shrink-
ing all the time, and this gives rise on the surface to small
warpings whose difference of elevation is usually not more tha
a few hundred feet. These are but surficial symptoms, due to
internal readjustment, remarkable as it may seem, in an earth
as rigid as steel—local accommodations of the earth’s mas
to loss of heat and molecular rearrangements. ‘These altera
tions finally set up strains in the. lithosphere which are tot
great for its strength to bear, and then there is a time 6
breaking and greater readjustment between the relativel}
settling and rising masses. At these times ranges of moun
tains are slowly raised up near the margins of the continents
due to the shortening, folding, and breaking of the earth's
crust, and the ranges have lengths of between 1,000 and 1,50%
miles. ‘These are the minor shrinkage movements, the “di
turbances,” which are coming more and more to be regardet
AND ITS INHABITANTS 71
as the basis for dividing the eras into periods of time, and how
often they occurred is one of the things that geologists are
trying to ascertain. In North America we know of at least
eight of these minor crustal readjustments, but in all the world
there are many more than this.
Finally, there comes a time of major shrinking that adjusts
all of the strains and stresses set up in the earth’s mass by the
minor, incompletely adjusted shrinkages. The earth has just
passed through one of these major readjustments, and accord-
ingly we see all of the continents standing far higher above
sea-level than has been the rule throughout geologic time, and
in many of them rise majestic ranges of mountains (see
Fig. 3). A grander, more diversified, and more beautiful
geography than the present one the earth has never had; this
statement is made advisedly and with the knowledge that our
planet has undergone at least six of these major readjustments
of its mass. These greater movements are the “revolutions”
that close the eras. Because the lands are then high they are
subject to more active erosion and in the last analysis all of
the broken-up detrital and dissolved material is carried away
by the streams to the oceans. At these times the continents are
also largest and the materials received by the oceans are laid
down on the outermost edges of the lands, where subsequent
transgressions by the sea cover and hide them from our
observation. After a long time the sea again comes to press
further and further upon the land and spreads more forma-
tions of stratified rocks over those left by the previous flood-
ings, the older geologic formations (see Fig. 11). Therefore
there is upon the present continents between each two such
successive formations a ‘“‘break’’ in deposition, a hiatus in the
geologic record, a “time interval” when no record other than
erosion is at hand. These ‘breaks’? in sedimentation are
representative of loss of record and are called ‘“‘intervals”’;
they are regarded as the closing times of the eras.
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AND ITS INHABITANTS 73
During these intervals when the continents are highest and
largest, the oceans are smallest in areal extent, the separate
continents are often united by land-bridges as Panama unites
the Americas, such bridges alter the direction of the great
ocean streams like the Gulf Stream, the high mountain ranges
alter the directions of the air currents and take out of them
their moisture so that great desert areas arise like our inter-
Rocky Mountain country, and because of these vast changes
nearly all of the major movements are accompanied by glacial
climates. Such mighty changes in the geography, topography,
and climate of the earth react strikingly upon the life of the
polar and temperate belts and so disarrange it that the major
“«ntervals” following the ‘revolutions’ are spoken of as the
critical times in the earth’s history—critical not only for the
geography, separating or uniting continental masses, but also
for life, since as a result of the lands then becoming high and
cool to cold in places and elsewhere moist or dry, vast domains
of organisms are forced to adapt themselves to the altered
conditions, or to migrate into more favorable areas, or to die
out and make room for the rising hordes of fitter types.
In this way cycle after cycle of organisms appear and vanish,
and their coming and going is brought about by the changing
environment. ‘Thus in the Mesozoic era, we see the lands
mastered by the cycads and conifers among the plants and the
dinosaurs among the animals, the air by the flying dragons,
and the seas and oceans by other reptiles and hordes of ammo-
nites and squid-like molluscs. The Great Reaper then steps in
and through struggle and the elimination of countless organ-
isms, resulting in regressive and progressive evolution, the
lands in the Cenozoic begin to bloom with more and more
flowering plants and grand hardwood forests, the atmosphere
is scented with sweet odors, a vast crowd of new kinds of
insects appear, and the places of the once dominant reptiles
of the lands and seas are taken by the mammals. Out of these
74 EVOLUTION OF THE EARTH
struggles there rises a greater intelligence, seen in nearly all ©
of the mammal stocks, but particularly in one, the monkey-
ape-man line. Brute-man appears on the scene with the intro-
duction of the last glacial climate, a most trying time for all ©
things endowed with life, and finally there results the domi-
‘nance of reasoning man over all of his brute associates. :
The Cenozoic era was a time of especially marked geo-
graphic alteration, as is especially well seen in the evolution
and spread of the elephant stock or Proboscidea. ‘These
animals arose in Africa early in the era, but there was no
means by which they could spread into other continents, because
Africa remained isolated. At about the middle of the Ceno-
zoic, the Alps were rising and these crustal alterations also
gave birth to a land-bridge across the Mediterranean connect-
ing Europe and Africa. Across this bridge the long-faced -
elephants, now all gone, spread first into Europe and thence
into Asia. A little later much of Asia began to rise, and the
culmination is seen in the grandest of all mountains, the present
Himalayas. ‘These alterations permitted the elephant stock
to spread across Asia and the Nome bridge into Alaska and
wider North America, and they had no sooner arrived there
than they were on their way across the newly arisen Panama _
bridge, thence to spread all the way south into distant Argen-
tina. North America was then peopled with many kinds of —
camels and horses, and they, along with many other animals,
migrated into South America. Wanderlust was upon the
world, and from South America there migrated into our coun-
try great sloths, a claw of one of which was found by Thomas
Jefferson in Virginia, and described as that of a huge lion.
The great Democrat may be excused for his error, since in
those days fossil sloths were unknown in North America;
rather should he be praised, for he is the only president who
has described a fossil, other than live ones! |
Times of volcanic activity. In all that has been said we
AND ITS INHABITANTS 75
see the periodic rejuvenation of the lands and the renewal of
active erosion. During these times of crustal unrest and
subsequent internal readjustment the molten portions deep
lown in the lithosphere are forced to rise as great hot tongues
into the cores of the mountain ranges, and in many places they
break through and give rise to active volcanoes. Beneath lie
great and highly heated masses that subsequently cool into
granitic rocks, and when long afterward the mountains are
worn away we discern among their roots these cores of granite.
However, during all the time of crustal rising and readjust-
ments the volcanoes are more or less active, spewing violently
01 quietly into the atmosphere tremendous volumes of rocks
lown into volcanic ashes and of lavas, and great quantities of
ises that increase the volume of the atmosphere, and much
avenile water is added by them and the thermal springs to the
adose water of the hydrosphere. Even though the litho-
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78 EVOLUTION OF THE EARTH
square miles. The greatest floods have spread from the Arctic
Ocean, others from the Pacific, and those of least amount from
the Atlantic. On the other hand, the number of floods was
greater from the south than from the north, and the waters
were most persistent in the Appalachian, lower Mississippi,
and Rocky Mountain areas. Where the greatest thicknesses
of sediments accumulate, out of this continental débris there
will arise, phcenix-like, a future mountain range. After about
40,000 feet of strata had been deposited in the Appalachian
trough, there arose near the close of the Paleozoic era the
majestic Appalachian Mountains, some of which may have
towered to at least 20,000 feet above the sea. In fact, the
original mountains were destroyed by erosion during the Meso
zoic era and the reélevated Appalachians of today are due te
later uplifts of more than 2,000 feet; subsequent erosion ha
developed them into their present interesting forms. Toda
one may travel comfortably over the roots of these once gran
mountains by way of the Pennsylvania, the Baltimore anc
Ohio, and the Chesapeake and Ohio railways.
Areal variability of organic habitats. We have seen tha
the dry lands become alternately larger and smaller, and in th
same way the shallow-water areas of the oceans are vasth
increased when they spread over the lands, and are agai
greatly decreased when the land areas increase in size. Thv
when the lands are widely flooded there is a great increase 1
the quantity of marine life, but little is originated that is ney
while on the lands the climate becomes insular, warm af
moist, and productive of the greatest amount of life in th
then restricted areas. On the other hand, when crustal reat
justments occur, the lands are largest, driest, and coolest, ar
these changing environments react on the life and bring abot
great alterations in the composition of the floras and fauna
At these times the shallow-water marine areas are smalle
and most variable in turbidity and salinity—conditians that
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80 EVOLUTION OF THE EARTH
set up renewed evolution, resulting in the rise of new stocks
In these various periodic physical changes we see the stimulus
that quickens organic evolution. Each cycle begins and closes
with a comparatively short time of accelerated evolution, while
the far longer intermediate tranquil periods lead to but little
that is striking in the way of new organisms.
CONCLUSION
From this summary of the earth’s changing surface and
climate during geologic time the following facts may bk
selected for emphasis:
First, geologic time is very, very long, indeed so vast as te
be beyond human comprehension. No geologist today thinks
that the evolution of the earth and its life could have taker
place in less than 100 million years. My own view as a st
dent of Historical Geology is that geologic time endured abou i
800 million years. In all of this we perceive how slowly th
physical and organic processes bring about the results of nature.
Second, the constant shrinking of the earth leads to an in
stability of surface that brings about periodic changes, no
only in the areal space relations of the water and land, bu
in the shapes and heights of the lands as well. Third,
sooner are the lands elevated above the sea-level than thi
weathering processes become more active and through tht
agency of the rain and wind all high places are, according t
human standards, slowly but surely moved into the seas an¢
oceans. Fourth, as a result of the transference of the high
lands into the water areas the latter are to a certain exte
displaced and periodically flood more or less of the land
Fifth, due to these surficial changes the atmosphere and th
climate are constantly changing in a small way, but every now
and then when the lands are largest, highest, and driest, a col
period appears and disarranges the entire organic world, bot
on the lands and in the waters. Sixth, when these ‘‘critica
AND ITS INHABITANTS Sr
periods” are upon the world, the face of the earth is scenically
grand and beautiful and at the same time the struggle for
existence among the living is most intense. The rulers of
the various domains find themselves overtrained and over-
specialized, and succumb one after another to the changing
environment. Their places are taken by the small, less
specialized, and heretofore little known stocks, which quickly
adapt themselves to their environments and become the
dominators of the organisms about them. In all of this un-
ceasing organic struggle most of the unadaptive families fail
to continue; others are pushed by the pulse of life into the
less desirable places, where they continue to exist as the static
| forms—the living fossils that tell us so much that is most
interesting of once prominent stocks of plants and animals;
but at all times much of life quickly responds to the changing
environment and is remolded into the more fit, active, and
| alert types. The experiences thus gained are passed on to
successive generations, and so the organic world as a whole
| advances to ever better living mechanisms endowed with higher
| and higher mentality.
The greatest mentality in the sea has been repeatedly de-
| rived from the continents, first in the fishes, then in the reptiles,
| and lastly in the mammals, and they have adapted themselves
| to the sea because of the ease with which they can there prey
upon the less alert and intelligent. Such adapted stocks in the
course of geologic time grow larger and larger, as, for
instance, the whales of today. Out of them, however, comes
no higher mentality. They represent an adaptation in the
wrong direction, that is, to an easier life, for the highest
Organisms with the greatest mentality have been developed
only on the land where the struggle for existence is fiercest
because of the constant necessity of adaptation to an environ-
ment subject to intense changes. Organic supremacy is at-
tained only through constant vigilance.
CHAPTER III
THE ORIGIN OF LIFE?
LORANDE LOSS WOODRUFF
PROFESSOR OF PROTOZOOLOGY IN YALE UNIVERSITY
PROTOPLASM. ‘The phenomena: to: which we apply the term
life have as their physical basis an inconceivably compl
physicochemical organization called protoplasm. The con-
ception of a common basis of all vital activities—that all living
nature is one—was the culmination of a long series of re-
searches during the first half of the nineteenth century, and
forms the corner-stone of modern biology. At the presen
time biology is the study of the properties of protoplasm
because to it, in the last analysis, the multifarious activities 0:
animals and plants must be referred. |
Since we are only familiar with life as a manifestation of
protoplasmic activity, the problem of the origin of life natu
rally resolves itself into the problem of the origin of proto-
plasm, and it is obviously impossible to discuss the origin 0!
living matter without an insight into its composition anc
organization. ‘The crude attempts of the alchemists of the
Middle Ages to produce artificially in crucible or retort ai
homunculus complete, though they had not the slightest com
ception of its composition, affords, in contrast with the
present trend of intensive analysis of protoplasm, a vivid illus
tration of the progress of scientific method in the interi
Obviously, analysis must precede synthesis even though tht
latter may never be achieved.
1 This lecture is published cenentinily as it was presented to a general uni
versity audience except that some of the more important references to th
literature of the subject have been inserted as footnotes.
_— ee a ee ee ee eee
a
Fae
AND ITS INHABITANTS 83
From one point of view it is impossible to analyze proto-
plasm because the least disturbance of its fundamental organi-
zation results in a cessation of those phenomena characteristic
f life, leaving matter in the non-living state before us. How-
er, since in the transformation of matter from the living to
he lifeless condition there is certainly no loss of weight, it
‘follows that the complete material basis of protoplasmic
‘organization remains for examination, and we may assume,
as a working hypothesis, that the properties of protoplasm are
a resultant of the properties of its constituent elements. But
it may be noted in passing that an analysis of the débris from
a destroyed factory probably would give one very little insight
in 0 the modus operandi of the intact organization.
_ Physical characteristics of protoplasm. Since the funda-
ie activities of all forms of protoplasm are of the same
nature, biologists have naturally tried to discover a correspond-
ing fundamental physicochemical basis in all living matter,
and thus far research, broadly speaking, indicates that such
exists. Living protoplasm ordinarily appears under a moder-
ately high power of the microscope as a viscid, granular fluid.
en studied with the highest magnifications, usually after
ee
being killed and stained, it often exhibits an extremely complex
structure which, however, cannot be briefly described, as it is
subject to considerable variation not only i in the protoplasm of
different organisms and tissues, but in the same microscopic
t mass under varying physiological conditions. Most fre-
e though, it seems to present a foam-like appearance,
due perhaps to closely crowded minute drops of a liquid alveo-
substance suspended in a continuous interalveolar substance,
also liquid but of a different physical nature. But it seems
clear, since studies of protoplasmic structure have revealed by
no means a complete correlation between its morphological
organization and its activities, that the key to the latter must
lie i in an ultra-microscopic architecture whose varying phases
84 EVOLUTION OF THE EARTH
are as it were reflected in the changing picture which the micro-
scope is capable of resolving. :
Chemical characteristics of protoplasm. Chemical analysis
shows that protoplasm is a colloidal complex comprising chiefly
the elements carbon, hydrogen, oxygen, nitrogen, sulphur,
phosphorus, potassium, calcium, sodium, chlorine, iron, and
magnesium—all of which are commonly found in the inor-
ganic world. Indeed, there is no chemical element present
which is peculiar to living matter. 4
But there are combinations of elements which are distinctly
characteristic of protoplasm, not being found in nature except
as the result of protoplasmic activity. “These chemical com-
binations are the proteins, carbohydrates and fats, and of
them the proteins are the most significant because they are
universally present as a part of all living matter and form
quantitatively its chief organic constituent. Proteins invari-
ably consist of the elements carbon, oxygen, hydrogen, nitro-
gen, sulphur, with sometimes phosphorus or iron; the nitrogen,
however, particularly distinguishing them from the othet
organic compounds of the living complex. Although the
stereochemical composition of a protein molecule has not been
determined, analyses show that it may be split up into a large
number of simpler, though still very complex molecules, anc
it is evident that its few elements may be represented by hun
dreds and even thousands of atoms. As Underhill has said
“Viewed from the chemical standpoint protein is seen as a hu g
molecule, complex in structure, labile in character and there
fore prone to chemical change. So large and intricate is tht
make-up of the molecule that chemists for generations havi
been baffled in their attempts to gain any adequate conceptio
of its nature. At the present stage of our knowledge it i
impossible to form any satisfactory definition of a protet
based either on its chemical or physiological properties.’
2 Underhill, F. P., “The Physiology of the Amino Acids,” 1915, published
Yale University Press.
ee ee
AND ITS INHABITANTS 85
_ It must suffice then to emphasize that it is the presence of
Broteins and the power of forming them which is the diag-
sti chemical characteristic of matter which is alive. It will
be recognized, of course, that proteins are not “‘alive,” and even
if the biochemist should succeed in artificially synthesizing a
; rotein, which has not thus far been accomplished, by group-
g together amino acids, which already have been made syn-
etically, this would not be in any sense the artificial produc-
on of life. But since proteins are the one molecular system of
which we are aware which is diagnostic of protoplasm, we have
e right to assume, for the present at least, that they represent
in a way the quintessence of that arrangement of matter which
exhibits life, while clearly recognizing that they are but an
integral part of the more complex, physicochemical structure
of protoplasm. Protein is, as it were, “the chemical nucleus
or pivot around which revolve a multitude of reactions char-
acteristic of biological phenomena.”
_ The vehicle of life manifestations is thus clearly a physico-
chemical complex and one of its most conspicuous and funda-
mental phenomena is what has been described as “energy
traffic” or the function of trading in energy. This consists in
appropriating energy from the environment, storing it in a
state of higher potential and later expending it in the kinetic
form. The lability of many of the molecular aggregates of
protoplasm, that is, their tendency to change in composition
toward greater or less complexity, coinciding with deoxidative
and oxidative processes, is an important characteristic and one
which endows the protoplasmic system as a whole with its
perennial plasticity—‘“‘its peculiar proneness to change its com-
position under the stimulus of slight changes in the energy-
equilibrium between itself and its surroundings.’
Definitions of life. This continual flux, which is now em-
8 Allen, F. J., “What is Life?” Proc. Birmingham Nat. Hist. and Philos. Soc.,
vol. 11, pp. 44-67, 1899.
86 EVOLUTION OF THE EARTH |
phasized as characteristic of protoplasm, was recognized as
a characteristic of organic individuals long before it wat
recognized in their physical basis, and is expressed 1 in neark
all definitions of life. Aristotle described life as “the asse
blage of the operations of nutrition, growth and destruction” ;
Bichat as “‘the sum total of the functions which resist . death”
De Blainville as “the twofold internal movement of composi
tion and decomposition, at once general” and continuous”
Lewes as ‘“‘a series of definite and successive ance both o:
structure and composition, which take place within an indi
vida without destroying its identity”; while’Spencer defines i
“the definite combination of heterogeneous changes, dot!
simultaneous and successive, in correspondence. with exter
coexistences and sequences’ “and as “‘the Sontmuaas “ad aaa
of internal relations to external relations.”
Obviously these statements are far from satisfactory, bu
no adequate, concise and yet comprehensive definition of lif
ever has been formulated, though it has been. essayed b
the best minds for generations. The fact is that life is to
complex to be described concisely, and so unique that it i
impossible to resort to the lexicographer’s trick of comparin
it with something else. Although many physicochemic:
phenomena contribute to life, not one of them will serve ¢
a criterion and all of them collectively do not offer, as yet,
satisfactory explanation. Whether behind the physical ph
nomena there are intangible metaphysical factors is a questic
outside the realm of biology. Since the limitations of ot
senses confine our perception to that part of our environmen
perhaps a very small ‘part, denoted by the fundamental co
cepts of science, matter, and energy acting in space and tim
we are forced to interpret, in so far as we are able, the li
processes in these terms. Thus far, however, the study
protoplasm has given no reason for abandoning the productiy
working hypothesis that life phenomena are an expression ¢
AND ITS INHABITANTS 87
a complex interaction of physicochemical laws which do not
differ fundamentally from the so-called laws operating in the
jnorganic world.
Individuality of organisms. We have seen that, although
all protoplasm has a similar fundamental physicochemical
basis, nevertheless there is a considerable, indeed a great, di-
versity in the minor details of its composition, not only in
different species of animals and plants, but also in the various
parts of the same animal or plant. In fact, organisms are
organisms because of specific local differentiations in the imme-
diate substratum of their vital activities.
_ This specification of protoplasm is rendered possible by the
fact that the structural units of all organisms are microscopic
masses of protoplasm, termed cells. The bodies of all higher
animals and plants are congeries of millions of these proto-
plasmic units, while many of the simpler forms of life consist
| of but a single one. Therefore it is apparent that a funda-
mental characteristic of organisms as we know them today is
not only that the material basis of their activities which we
call life is the somewhat protean, though no less real, proto-
plasm, but also that this physical basis is individualized into
structural moieties, the cells. It is not an overestimation to
say that the recognition of this dual similarity of physico-
chemical and morphological organization of all living things
must be considered the greatest single generalizatioa which
biological science has attained, and must be ranked with the
theory of organic evolution, of which it is the corner-stone, in
its far-reaching bearings.‘
*See Allen, F. J., op. cit.
Minchin, E. A., “The Evolution of the Cell.” Amer. Nat., vol. 50, 1916,
pp. 5-38, 106-118, 271-283.
_ Conklin, E. G., “The Basis of Individuality in Organisms from the
_ Standpoint of Cytology and Embryology.” Science, new ser., vol. 43,
1916, pp. 523-527.
Driesch, H., “The Problem of Individuality,” 1914.
88 EVOLUTION OF THE EARTH
Obviously, then, the biologist in dealing with living sub-
stance has it only in the form of cells, and these, singly or col-
lectively, constitute individual organisms. Thus in the final
analysis protoplasm is known to us only in the form of living
individuals, and the expressions ‘“‘protoplasm”’ and “‘life” are
merely abstractions, one indicating that all individuals have
to a certain extent a common organizational foundation and
the other that they exhibit certain characteristic actions and
reactions. The living organism is a microcosm which exhibits
a permanence and continuity of individuality correlated with
specific behavior, and this it transmits to other matter which
it makes a part of itself and to its offspring in reproduction.
It is apparent that the phenomena which we call life are
dependent upon the interplay and interchange between the
highly organized protoplasmic complex and its environment.
Although the organism, whether animal or plant, is an indi
vidual, still it retains its individuality—lives—solely by its
powers of developing and maintaining exquisite adjustments to
its surroundings, and therefore the concept protoplasm has
little or no content unless environmental relations are included.
This fitness of the organism, as Henderson has recently empha-
sized, is but one part of a reciprocal relationship of which
the fitness of the environment is the other. ‘The fitness of the
environment results from characteristics which constitute a
series of maxima—unique or nearly unique properties of
water, carbonic acid, the compounds of carbon, hydrogen,
and oxygen and [for primordial life] the ocean.” ‘No other
environment consisting of primary constituents made up of
other known elements, or lacking water and carbonic acid,
could possess a like number of fit characteristics’ for proto-
plasmic phenomena. The properties of matter and the course
of cosmic evolution are intimately related to the structure of
the living being and to its activities. Indeed, ‘‘the whole evolu-
tionary process, both cosmic and organic, is one, and the biolo-
AND ITS INHABITANTS 89
_ gist may now rightly regard the universe in its very essence as
_biocentric.’”®
\ History of the establishment of biogenesis. The previous
lectures in.this course have described the theories in regard to
the origin of this environment on the earth, so we turn now
“to the question of the origin of life—that primeval question
_ which has been asked by all ages and the answers to which, if
_ they do no more, at least epitomize the philosophic and scien-
; " tific perspective of the times.
_ One of the greatest intellectual characteristics of the Greeks
_ was their scientific curiosity, and therefore it is not strange
_ that the first biological question which they propounded was
_ in regard to the origin of life. Even to the best minds of the
ancients there was little incongruity’ in the idea of animals
and plants arising de novo from earth or water. Although
Aristotle, whose scientific studies formed the foundations of
natural history, devoted a great deal of thought and ingenuity
to the subject of the origin of animals, he apparently accepted
| with little reservation the statements that even such highly
developed organisms as worms, insects, and some fishes could
come into being from mud. Such ideas are voiced over and
over again through more than twenty centuries by philosopher
and poet, theologian and layman.
Lucretius, in his De rerum natura, says that “with good
reason the earth has gotten the name of mother, since all
things are produced out of the earth, forming in rain water
and in the warm vapors raised by the sun.”’ Virgil in the
fourth Georgic, which is devoted to a discussion of bees,
graphically describes a simple method used in Egypt for obtain-
ing bees from the dead bodies of bullocks.
Coming to the medieval period and later we find Cardan
stating that water engenders fishes and that various types of
animals arise from fermentation. Swan in his Speculum mundi
_ 5 Henderson, L. J., “The Fitness of the Environment,” 1913.
go EVOLUTION OF THE EARTH
says that a dead horse breeds wasps, a mule produces hornets,
while from an ass arise bumble bees. Van Helmont, one of
the founders of chemical physiology, gives particularly specific
directions for the experimental production of scorpions and
mice, while Kircher actually figures animals-which he avers
arose under his own eyes through the influence of water on
the stems of plants; an instance, perhaps, not of spontaneous”
generation but of the transformation of plant into animal,
which was also a ‘notion prevalent at the time. The ironical”
reflections of one Ross, aroused by the scepticism of Sir
Thomas Browne in regard to mice arising by putrefaction, are
quite typical of seventeenth-century opinion. He says: “So
we may doubt whether in cheese and timber worms are gen-
erated, or if beetles and wasps in cow-dung, or if butterflies,
locusts, shellfish, snails, eels, and such life be procreated of
putrefied matter, which is to receive the form of that creature
to which it is by formative power disposed. ‘To question this
is to question reason, sense, and experience. If he doubts
this, let him go to Egypt, and there he will find the fields —
swarming with mice begot of the mud of Nylus, to the great
calamity of the inhabitants.”
That such ideas of the origin of life were prevalent and their
truth untested by experiments is an eloquent commentary on
the general state of the scientific method before the Renais-
sance. It was Francesco Redi, an eminent Italian scholar, —
physician, and naturalist, who, not content with tradition, but
with great faith in observation, made a study of the origin ©
of maggots in decaying animal matter. By the simplest of
experiments he found that maggots never developed in meat ~
to which flies were not allowed access and that the seeming
transformation of meat into maggots was the result of flies
laying their eggs on the meat. When this was prevented, no —
matter how he varied the materials of his experiments, he
obtained the same result, and naturally concluded that the sup- —
B
2. i rr ———————e
AND ITS INHABITANTS gI
posed appearance of life where life had not been previously
was due to the introduction of foreign living material.®
One may imagine that the practical man of affairs, who
scofied at Redi toiling under the Italian sun with meat and
“maggots to satisfy a scientific curiosity, little dreamed that
the practical results which germinated from this folly would
be among the most important factors in twentieth- -century
‘civilization. Indeed, it is difficult to overestimate the i impor-
tance of Redi’s conclusion from either the theoretical or prac-
tical viewpoint, for with it was definitely formulated the theory,
which has gained content and impetus as the years have rolled
‘on, that matter does not assume the living state, at the present
time at least, except under the direct influence of preéxisting
living matter.
The influence of this work gradually became apparent in
scholarly literature; Derham, for instance, stating that ‘‘Spon-
taneous generation is a doctrine so generally exploded that I
shall not undertake to disprove it. It is so evident that all
animals, yea and vegetables, too, owe their production to
parent animals and vegetables, that I have often admired at
the sloth and prejudices of the ancient philosophers in so easily
taking upon trust the Aristotelian, or rather A‘gyptian doc-
trine of equivocal generation.’” Another writes: “I would
as soon say that rocks and woods engender stags and ele-
phants as affirm that a piece of cheese generates mites. Stags
are born and live in woods, and mites in cheese, but they both
owe their being to that of other animals.’ And again, Henry
Baker, the versatile microscopist of the Royal Society, says:
“Nothing seems now more contrary to reason, than that chance
and nastiness should give a being to uniformity, regularity
and beauty . . . and create living animals. . . . This, however,
© Redi, F., “Esperienze Intorno Alla Generazione Degl’ Insetti,” 1668. Eng-
lish translation by M. Bigelow, 1909.
*Derham, W., “Physico-Theology,” 1713,
92 EVOLUTION OF THE EARTH
was the opinion not only of the ignorant and illiterate, but off
the most learned grave philosophers of preceding ages; and
would probably still have been taught and believed had not
microscopes discovered the manner how all these things are
generated, and restored to God the glory of his own =
work.’’® |
Although it is relatively easy to follow when one’s eyes are
opened, it is not to be imagined, of course, that one experiment,
or even the long series of studies made by Redi, could annihi-
late a time-honored belief. Indeed, the history of the establish-
ment of the theory extends down even to the present time,
for no sooner had experiment apparently disposed of it com-
pletely than it arose again, pheenix-like, with fresh vigor in
a slightly different phase demanding further study. |
Among others, a Scotch priest, ‘Turbervill Needham, studied
the problem and believed that various minute organisms,
which improvements in the microscope were bringing to the
fore, appeared spontaneously in infusions which he boiled ane
corked up in flasks.®° His results attained considerable noto-
riety, since Buffon, the famous French naturalist, found in
them a substantiation of his theory that all organisms are
aggregates of indestructible units, which upon the death of
the individual are disseminated in nature and later are e
ployed as moieties in the organization of arising generations
Needham’s results, however, were soon shown by Lazzar
Spallanzani* to have been obtained by inadequate sterilizatio:
and sealing of the infusions which he studied; but at this poin
objections came from another source—the chemists who had
recently discovered oxygen and the importance of this elemen
8 Baker, H., “The Microscope Made Easy,” 1742.
9 Needham, T., “A Summary of Some Late Observations upon the Generation,
Composition, and Decomposition of Animal and Vegetable Substances.” Philo:
Trans. Roy. Soc. London, 1748. ¢
10 Spallanzani, L., “Expériences pour Servir a l’Histoire de la Génération de
Animaux et des Plantes ” 1786.
.
4
;
AND ITS INHABITANTS 93
in the free state for the processes of life and for putrefaction
and fermentation of organic matter. The treatment to which
Spallanzani subjected his infusions might well have so changed
‘the organic matter, they argued, that it was incapable of pro-
‘ducing life. This objection was met by a series of experiments
by various investigators during the first half of the last
Bentury, which showed conclusively that thoroughly sterilized
fusions never developed living organisms when air was
admitted which had. been rendered sterile by heat or by having
all suspended matter removed, while the studies of Schwann
and Cagniard de la Tour intimated that fermentation and also
putrefactive changes in organic infusions are themselves the
result of micro-organisms.
So far it was clear that organic infusions in which no life
is present do not develop living organisms except when con-
taminated with material from an extraneous source, which is
ordinarily the atmosphere, but it remained then to be shown
that living organisms are practically ubiquitous in the atmos-
phere and thus are available for the initiation of the micro-
flora and fauna of infusions. This was put upon a broad
empirical basis chiefly by the labors of Pasteur. Tyndall,
who himself made a notable contribution to the solution of
the problem by researches in experimental physics, confidently
| asserted that there seems to be no flaw in the reasoning, and
it is so simple as to render it unlikely that the notion of life
| developing from dust can ever again gain currency among the
| members of a great scientific profession.
’ We thus reach the general conclusion that, so far as human
| observation and experimentation go, no form of life arises
today except from preéxisting life. But since life is present on
the earth now and the paleontological record indicates that it
has been present without interruption for some hundreds of
millions of years, we have to consider the following alterna-
| tive: Either life was transported to this planet from some
| The magnitude of the problem may be inferred from a recent
statement by the dean of American biologists that ‘‘the study
94 EVOLUTION OF THE EARTH
other part of the universe; or life arose spontaneously from
non-living matter at one period at least in the past as a natural
result of the evolution of the earth and its elements.
THEORIES OF THE ORIGIN OF LIFE Ss
It will be useless to offer any extended discussion or specific
criticisms of the individual theories of the origin of life, for
each one rests on many uncertain postulates, which have <
greater or less degree of probability dependent, to a con-
siderable extent, on the personal equation of the critic. It
should be said, however, in justice to the authors of the theories
which we shall outline, that the theories are not advanced as
final solutions of the problem of the origin of life, but rather
as gropings toward a formulation of the conditions which con-
ceivably might have attended and contributed to its genesis.
of the cell has on the whole seemed to widen rather than to
narrow the enormous gap that separates even the lowest forms
of life from the inorganic world.’’™
Vitalism. ‘The vitalistic conception that life phenomena are
in part at least the resultant of manifestations of matter and
energy which transcend and differ intrinsically in kind from
those displayed in the inorganic world—a denial, as it were,
in the organism of the full sufficiency of known fundamenta
laws of matter and energy—has arisen many times in th
development of biological thought, either as a reaction against
premature conclusions of the nascent sciences or from af
overwhelming appreciation of the complexity of life phe
nomena. Vitalism goes back as far as ‘the history of science
is recorded, but it attained its most concrete formulation as a)
doctrine during the early part of the eighteenth centu
in opposition to the obviously inadequate explanations which
11 Wilson, E. B., “The Cell in Development and Inheritance,” 2d ed., 1900;
“The Problem of Development,” Science, new ser., vol. 21, 1905, pp. 281-294.
4
] AND ITS INHABITANTS 95
chemistry and physics could offer for the phenomena of irrita-
bility of living matter then prominently before the professional
biologist. The vitalists at that period abandoned almost
- os :
completely all attempts to explain life processes on a physico-
chemical basis and assumed that an all-controlling, unknown,
-and unknowable, mystical, hyper-mechanical force was re-
sponsible for all living processes. It is apparent, of course,
that such an assumption in such a form is a negation of the
scientific method, and at once removes the problem from the
realm of scientific investigation. No biologist at the present
day subscribes to vitalism in this form; some uphold vitalism
(if it must still be called vitalism) in its modern aspect, while
all will undoubtedly admit that we are at the present time
“utterly unable to give an adequate explanation of the funda-
“mental life processes in terms of physics and chemistry.
Whether we shall ever be able so to do is unprofitable to
7
‘speculate about, though certainly the twentieth century finds
relatively few representative scientists who really expect a
“scientific explanation of life ever to be attained or who expect
that protoplasm will ever be artificially synthesized. But,
“In ultimate analysis everything is incomprehensible, and
the whole object of science is simply to reduce the fundamental
incomprehensibilities to the smallest possible number.’
Cosmozoa theory. ‘The establishment of the fact that, so
| far as we can determine, life does not arise except from
| preéxisting life at the present time, and the dawning realiza-
tion of the intrinsic and unique complexity of the architecture
of matter in the living state, which has thwarted the attempts
of alchemist and biochemist to synthesize even one of its chief
| molecular aggregations, have led several scientists to suggest
and elaborate the hypothesis that life has never arisen de novo
on the earth but has been transported hither from elsewhere
in the universe.
)
)
12 Huxley, T. H., “Darwiniana.” “Collected Essays,” vol. 2, 1893, p. 165.
~
96 EVOLUTION OF THE EARTH :
Such an idea was advanced by Richter, and later inde-
pendently suggested and discussed by Helmholtz and Kelvin.*
On the assumption that some of the heavenly bodies have
always been the abode of life, and from the fact that smal 1
solid particles, which presumably have formerly been an
integral part of such bodies, are moving everywhere in space, —
Richter pictures these particles as the vehicles which dis-
seminate the simplest forms of life through interstellar space
to find lodgment and development upon such planets as afford
a suitable environment. Clearly, from this point of view, life
is as old as the universe itself—living matter has never
originated but has been transported from world to world.
Hence the acceptance of this permanent dualism of living and
lifeless matter does not answer the broad question as to the
origin of life, but transfers its origin to a ‘“‘conveniently inac-
cessible corner of the universe where its solution is impossible.”
However, the question before us is the origin of life upon the
earth, and the plausibility of this cosmozoa theory depends on
two assumptions—that life exists elsewhere in the universe
and that life can be maintained during the interstellar voyage.
Neither assumption has, of course, any empirical foundation
whatsoever, though the second offers at least something
tangible for discussion. J
Many of the lower forms of life, such as the bacteria anc
protozoa, have the power of developing, particularly unde:
the influence of unfavorable environmental conditions, pro
tective coverings of various sorts about themselves and of
assuming a resting condition in which all the metabolic activi
ties characteristic of active life are reduced to the lowest ebb
In this spore or encysted state they are immune to extremes 0
temperature and desiccation to which they readily succuml
during vegetable life. It has recently been found, for example
13 Verworn, M., “Allgemeine Physiologie,” 5th ed., 1909. English translatio
of 2d edition by F, S. Lee, 1899.
AND ITS INHABITANTS 97
that certain types of bacteria can successfully endure a
temperature of nearly —200° C. for six months, and about
—250° C. for shorter periods, that is, a temperature con-
| siderably lower than that at which any chemical reactions
are known to occur; and again the spores of other bacteria
can withstand a temperature as high as 120° C. for a short
| time. The cysts of some relatively highly specialized protozoa
‘can retain their vitality for at least fifty years, and the seeds
‘of the higher plants, particularly those with a thick and im-
permeable integument, have been found to retain the power of
germination for nearly a century, though the statement that
grain from Egyptian tombs still maintains its power of growth
has been disproved. The ability to continue dormant life
clearly seems to depend to a large extent on the presence
‘of a dry enclosing membrane, since under these conditions the
entrance of gases is impossible and the chemical processes
within are at a minimum. Thus it is apparent that certain
Organisms may survive unfavorable conditions for a long
period, and we have no reason for believing that the limits of
endurance of other forms do not greatly exceed that of the
few which have thus far been studied.
On the other hand, the exigencies to which living matter
would be exposed when started on its interstellar journey are
not inconsequential. Meteors in their fall through the earth’s
atmosphere become incandescent and, if they are the vehicle
of transfer, it would only be conceivable for life to survive
far below the surface where the temperature is lower. To
avoid this and other difficulties it has been suggested by
Arrhenius that the radiation pressure of light is sufficient to
Overcome the attraction of gravitation for particles of the
extraordinary minuteness of some of the lowest forms of life,
and that isolated germs might make the journey to the earth.
But on the assumption that such an individual organism were
forced out into space by the mechanical pressure of the light
98 EVOLUTION OF THE EARTH
waves from the sun of the nearest solar system, it would re-
quire many thousand years to reach the earth. Only on the
view that cells are potentially immortal and can remain in a
dormant condition nearly indefinitely can we believe that lif
has reached the earth from other planets. Arrhenius maintains
that this is possible owing to the exceedingly low temperature
and absence of water-vapor which must prevail in cosmic space;
and Loeb states that there is no reason why spores should lose
appreciably more of their germinating power in ten thousand
years than in six months.** ;
Without further discussion it is apparent that the theory is
one which cannot be proved or disproved. At first thought,
and as first outlined by Richter, it commanded little serious
attention; but with its strictly scientific formulation by later
physicists and biologists, and especially in view of our increas-
ing appreciation of the potentialities of life in the latent state,
we are justified perhaps in seriously questioning, with Helm-
holtz, ‘‘whether after all life has ever arisen, whether it maj
not be even as old as matter, and whether its germs, passing
from one world to another, may not have developed where
they found favorable soil.”” But the majority of biologists
doubtedly would agree with Schafer that, ‘knowing what we
know, and believing what we believe, as to the part played by
evolution in the development of terrestrial matter, we are,
without denying the possibility of the existence of life in othe
parts of the universe, justified in regarding these cosmic theories
as inherently improbable—at least in comparison with the solu
tion of the problem which the evolutionary hypothesis offers.’”
14 Arrhenius, S., “Worlds in the Making.” English translation by H. Borns
1908.
Loeb, J., “The Organism as a Whole from a Physicochemical Viewpoint,’
1916.
Woodruff, L. L., “A Pedigreed Race of Paramecium.” Proc. Soc. Exp. Biol
and Med., vol. 9, 1912.
15 Schafer, E. A., “Life: Its Nature, Origin and Maintenance.” Presidentia
address, Brit. Assoc. Adv. Sci. Science, new ser., vol. 36, 1912, pp. 289-312,
a AND ITS INHABITANTS 9
_ With this in mind we may briefly survey some of the modern
scientific conjectures which attempt to formulate the phe-
nomena attendant upon the evolutionary origin of matter in
the living state from the inorganic upon the earth.
| Pfliiger’s theory. Assuming that the surface of the earth
was once incandescent, Pfliiger has offered a suggestive theory
‘the crucial points of which turn on the chemical characteristics
of the proteins. He emphasizes that there is a fundamental
difference in the nitrogenous radicals of what he terms “‘dead”
proteins, as egg albumin, and “living” proteins taking an active
part in the economy of protoplasm, since the nitrogenous
omposition products of the latter either contain the cyano-
a ; a radical or can be artificially produced from compounds of
cyanogen by atomic rearrangement. ‘This suggests to Pfliiger
the probability that the cyanogen radical is an integral part
of the molecular complex of living proteins and since in the
formation of cyanogen a large amount of heat is absorbed, it
follows that this radical possesses a large amount of internal
energy and thus with it there is “introduced into the living
‘matter energetic internal motion.’
Pfliiger’s idea that the protein molecules of living proto-
plasm owe their diagnostic characteristics, in particular their
bility, to cyanogen is supported, be believes, by certain
analogies, which we need not consider, between cyanogen com-
pounds and “‘living’’ proteins. Indeed, he says, the “‘similar-
‘ity i is so great that I might term cyanic acid a half living mole-
cule.” “When we think of the beginning of organic life, we
must not think primarily of carbonic acid and ammonia; for
they are the end of life, not the beginning.” ‘The beginning
lies rather in cyanogen.” Pointing out that cyanogen and its
compounds arise only in an incandescent heat, he holds that
“life is derived from fire, and its fundamental conditions were
laid down at a time when the earth was still an incandescent
ball.” Thus living material owes its genesis to cyanogen com-
100 EVOLUTION OF THE EARTH
pounds which, on account of their tendency to decomposition,
entered into relations with carbon compounds arising at similar
temperatures. When the temperature conditions of the earth’s |
surface permitted the precipitation of water, this with the
salts and gases in solution joined the growing cyanogen-
carbon complex and gave rise to the highly labile protein.
molecules so characteristic of protoplasm. Thus, according to
Pfliiger’s hypothesis, arose a relatively simple, homogeneous
material from which has been evolved the highly differentiated
PROLOE aa masses or cells of organic life today.*®
Moore’s theory. Moore essays to picture with rather bold
strokes the origin of life from the inorganic elements of the
cooling earth, by a continuation of the slow process of com-
plexification which he sees inherent in matter. ‘This note,”
he says, “‘cannot be too strongly sounded that as matter is
allowed capacity for assuming complex forms those complex
forms appear. As soon as oxides can be there, oxides appear;
when temperature admits of carbonates, then carbonates are
forthwith formed. . . . Next in order of development prio
to life inorganic colloids begin to appear in solution, or sus-
pension, in the waters of the cooling globe, alumina and silica
deposited in colloidal form are seen in many sedimentary
rocks. Single molecules existing in solution, and capable of
forming colloids, with alterations in temperature, and in
chemical reaction of the environment, begin to form com-)
plexes, or solution aggregates, in which the unit of chemical
structure passes from the atom to the molecule.” |
Accompanying these structural changes, the energy types
and phases inhabiting the unit of structure also vary. The
rates of vibration or of phasic activity in the colloidal aggre=
gates become slower than in the simpler molecules of the
16 Cf. Verworn, of. cit.
Pfliger, E., “Ueber die physiologische Verbrennung in den lebendigen
Organismen.” Arch, f. d. ges. Physiol., vol. 10, 1875. q
? AND ITS INHABITANTS IOI
cr stalloids. The characteristics of colloids, slowness of
reaction, metastable equilibrium, delicacy of union, and in-
= reactivity of specific type, become present in the forms
‘yironment. As the complexity of structure increases, the
ature of the equilibrium in the colloidal aggregates ap-
proaches more and more towards that labile, easily destroyed,
but also more readily constructive condition which is char-
acteristic of life.
_ Moore then states in a general way as a “law universal in
Ml ts application to all matter, although varying in intensity in
‘different types of matter, and holding throughout all space as
“generally as the law. of gravitation—a law which might be
called the Law of Complexity—that matter so far as its energy
‘environment will permit tends to assume more and more
complex forms in labile equilibrium. Atoms, molecules, col-
loids and living organisms arise as a result of the operations
‘of this law, and in the higher regions of complexity it induces
organic evolution and all the many thousands of living forms.”
Tn this manner he conceives that the hiatus between non-living
and living things can be bridged over, and that life arose as
an orderly development, which comes to every earth in the
universe in the maturity of creation when the conditions arrive
within the suitable limits.*’
_ Allen’s theory. We may now consider certain theories
which conform with the primal earth conditions as postulated
by the planetesimal theory of its origin. Allen, for example,
maintains that it is simplest to believe that the circumstances
which support life would also favor its origin and that if life
| formerly existed actively outside the range of the freezing
and boiling points of water, it must have been quite different
from that which now exists. Life then arose at the period
when the physical conditions of the earth came to be nearly
1, Moore, B., “The Origin and Nature of Life,” 1912.
102 . EVOLUTION OF THE EARTH
what they are at present. At such a time, as now, dis-
charges of lightning passing through the damp air produced
ammonia and oxides of nitrogen, which were carried down by
rain into the streams and pools. If this process went on un:
modified, the result would be a large accumulation of nitrogen
compounds in the terrestrial waters, which would also contain
carbon dioxide probably in larger amounts than the waters o:
the present day, because since that time much carbon dioxide
has been withdrawn from the air and water and locked up in
organic matter. In the same waters the usual metallic ele-
ments, in the form of chlorides, sulphates, phosphates, etc.,
would also be present.
Thus there must have been brought together an abundance
of such raw materials as are required for the production of
living matter, and, provisionally, Allen imagines some suck
reactions as the following to have occurred: solar energy
acting on the water or damp earth containing the raw material;
just mentioned, caused dissociation and rearrangement of thé
atoms, the nitrogen abstracting oxygen from its compound
with carbon, hydrogen, sulphur, and other elements and de:
livering it to the atmosphere. Not much energy would bi
absorbed by a transparent liquid; but such actions as th
above would occur particularly in water containing compound
of iron in solution or suspension since these compounds woul
absorb the solar energy. In this way compounds of nitroger
carbon, etc., accumulated in the water or damp earth; an
further reactions, anabolic and catabolic, occurred among ther
by virtue of the lability of the nitrogen compounds. Life a
this stage was of the humblest kind since there were no defini
organisms, only diffuse substances trading in energy, an
between this stage and the evolution of cellular organisms 4
immense period probably elapsed. |
Allen questions whether the earliest organisms were able ;
utilize the energy of sunshine or whether their whole energ
q
AND ITS INHABITANTS 103
was derived from the nitrogen compounds produced in the
atmosphere by lightning, but in any case he thinks the faculty
of appropriating the energy of sunshine must have developed
at an early period, if not at the very beginning. In the func-
tions of life, constructive process precedes destructive, and it
therefore seems. reasonable to suppose that the earliest forms
| of life were concerned more in accumulating than in dispersing
energy, and that the energy-dispersing organisms, like bacteria
and fungi, have had a comparatively late origin.
_ Allen’s view of the origin of life “implies that the first
attempts at life are still continuing, and that if by chance all
life were wiped out, another cycle would begin.” Such he be-
lieves to be the case, and to the question, ‘‘Why do we not find
evidence of these processes in the form of primitive vital sub-
stances in water or elsewhere ?”’ he answers that as soon as any
such substances begin to be formed, they are seized and assimi-
lated by the already developed organisms.*
Troland’s enzyme theory. With the increasing realization
of the importance of enzymes in the economy of organisms it
is not strange that in these chemical bodies has been sought
the key to life’s origin, and accordingly we find Troland stating
that life is something which has been built up about the
enzyme. This author assumes that, at some moment in earth
history, a small amount of a certain autocatalytic enzyme
suddenly appeared at a definite point within the yet warm
ocean waters which contained in solution various substances
reacting very slowly to produce an oily liquid immiscible with
water. If, when this occurred, the enzyme became related to
the reaction in such a way as to greatly increase its rate by
reducing the chemical friction involved, Troland believes it is
obvious that the enzyme would become enveloped in the oily
material resulting from the reaction, and the little oil drop
| would increase until it was split into smaller globules by water
48 Allen, F. J., of. cit.
104 EVOLUTION OF THE EARTH
currents, provided the original substances which combined
were soluble in oil as well as in water. Thus arose, according
to his theory, the first and simplest life-substance, possessing
the power of indefinitely continued growth, by virtue of the
postulated autocatalytic character of the initial enzyme. Tro-
land is convinced that this theory of the origin of life satisfies
most of the objections which have been advanced against
ordinary chemical hypotheses, since the characteristic catalytic
power of the enzyme accounts for the elevation of the rate of
synthetic action from, practically zero to one making possible
rapid growth of the protoplasmic mass. The same catalytic
property explains the localization of the reaction in a definite
region, for catalysis can occur only where the catalyzer itself
is present. ‘The theory also provides a basis for the per-
manent growth of the primitive organism—if such we dare to”
call it—as well as for its reproduction, without, in general, the
loss of its specific individuality.” Troland suggests that the
most fundamental objection which can be raised against’ his
ideas is the fortuitous formation of the original enzyme; but
he insists that since only a single molecule of the enzyme was
required, and since multitudinous chemical reactions undoubt-
edly took place in the primordial ocean, this objection of
improbability is almost absurd.*® |
Osborn’s theories. The most recent consideration of the
origin of life is that by Osborn, published during the past year.
Starting with the postulate that “the primal earth, air, and
water contained all the chemical elements and three of the
most simple but important chemical compounds, namely,
water, nitrates, and carbon dioxide, which are known to be
essential to the prechlorophyllic and chlorophyllic stages of
the life process,” Osborn suggests that an initial step in the”
origin of life was the codrdinating or bringing together of
19 Troland, L. T., “The Chemical Origin and Regulation of Life.” Th
Monist, vol. 24, 1914, pp. 92-133. {
ae gn
AND ITS INHABITANTS 105
_ these elements, which, so far as we know, had never been in
- combined action before. He states that the thermal condi-
tions of present-day living matter point to the probability that
this codrdinating of the ‘“‘life elements’ was initiated when
‘portions at least of the earth’s surface and waters had tem-
_ peratures of between 6° and 89° C. and before the atmospheric
vapors admitted a regular supply of sunlight. The earliest
' function of living matter appears to have been to capture and
_ transform the electric energy of the chemical elements char-
acteristic of protoplasm, and this power probably developed
2 only in the presence of heat energy derived from the earth or
_ from the sun. Frankly admitting, however, that both the time
_and place of the origin of life is a matter of pure speculation,
-in which we have as yet no observations or uniformitarian
_ reasoning to guide us, Osborn advances five hypotheses in
regard to it as follows:
An early step in the organization of living matter was the
assemblage of several of the chemical elements essential to
life. Of these the four most important elements were ob-
tained from their previous combination in water, from the
nitrogen compounds of volcanic emanations or from the
atmosphere, consisting largely of nitrogen, and from atmos-
pheric carbon dioxide. The remaining elements came from
the earth.
Whether or no there was a sudden or a more or less serial
grouping of these elements, one by one, Osborn is led to a
second hypothesis that ‘‘they were gradually bound by a new
form of mutual attraction, whereby the actions and reactions
of a group of life elements established a new form of unity in
the cosmos, an organic unity or organism quite distinct from
the larger and smaller aggregations of inorganic matter pre-
viously held or brought together by the forces of gravity.”
This leads to the hypothesis that since all living cells are
colloidal the grouping of the ‘“‘life elements” took place in a
106 EVOLUTION OF THE EARTH
state of colloidal suspension, for it is in this state that the —
life elements best display their incessant action, reaction and
interaction.
With this assemblage, mutual attraction, and colloidal con- |
dition, a fourth hypothesis is that there arose the rudiments of ~
competition and selection. ‘‘Was there any stage in this group- —
ing, assemblage, and organization of life forms, however ~
remote or rudimentary, when the law. of natural selection did —
not operate between different unit aggregations of matter?
Probably not, because each of the chemical life elements pos-_
sesses its peculiar properties which in living compounds best
serve certain functions. ‘This codperation was also an appli-.
cation of energy new to the cosmos.’ In other words, every
“life element”’ has its single and multiple services to render to
the organism. .
And as a fifth hypothesis relating to the origin of organisms,
Osborn advances the idea that the evolution and specialization —
of various enzymes has proceeded step by step with the evolu- |
tion of plant and animal functions, ‘‘since in the evolution from ~
the single-celled to the many-celled forms of life and the multi-
plication of these cells into hundreds of millions, into billions, —
and into trillions, as in the larger plants and animals, bio-
chemical coédrdination and correlation become increasingly
essential.’’?° : '
Conclusion. Such are the principal attempts of modern
biologists to formulate in some concrete way the evolution of
matter in the living state from the elements of the earth.
Their consideration is valuable in that it indicates the uni-
formitarian trend which, at the present day, biological thought -
follows in this field, and also the diversity of results arrived 7
at by the scientific imagination when it is largely untrammeled
20 Osborn, H. F., “The Origin and Evolution of Life upon the Earth.” Scien-_
tific Monthly, vol. 3, 1916, pp. 5-22, 170-190, 289-307, 313-334, 502-513, 601-614.
AND ITS INHABITANTS 107
by facts. One, however, cannot but admire the scientific
caution of Huxley when he said:
_ “Looking back through the prodigious vista of the past, I
‘find no record of the commencement of life, and therefore I
“am devoid of any means of forming a definite conclusion as
‘to the conditions of its appearance. Belief, in the scientific
sense of the word, is a serious matter, and needs strong founda-
tions. To say, therefore, in the admitted absence of evidence,
that I have any belief as to the mode in which existing forms of
life have originated, would be using words in a wrong sense.
But expectation is permissible where belief is not; and if it
were given to me to look beyond the abyss of geologically
recorded time to the still more remote period when the earth
was passing through physical and chemical conditions, which it
can no more see again than a man can recall his infancy, I
should expect to be a witness of the evolution of living proto-
plasm from not living matter. I should expect to see it appear
under forms of great simplicity, endowed, like existing fungi,
with the power of determining the formation of new proto-
plasm from such matters as ammonium carbonates, oxalates
and tartrates, alkaline and earthy phosphates, and water,
without the aid of light. That is the expectation to which
analogical reasoning leads me; but I beg you once more to
recollect that I have no right to call my opinion anything but
an act of philosophical faith.””
Thus since biologists are at the present time absolutely
unable, and probably will be for all time unable, to obtain
empirical evidence on any of the crucial questions relating to
the origin of life on the earth, their endeavors are and must
be directed chiefly toward an intensive analysis of life mani-
festations as exhibited in the physical basis of individual
21 Huxley, T. H., “Biogenesis and Abiogenesis.” Presidential address, Brit.
Assoc. Adv. Sci. 1870. “Collected Essays,” vol. 8.
108 EVOLUTION OF THE EARTH .
organisms, in their bionomic relations, and in the kaleidoscopic
panorama which geological history presents of their evolution
In the closing words of Darwin’s Origin of Species: .
“Tt is interesting to contemplate a tangled bank, clothed with
many plants of many kinds, with birds singing on the bushes,
with various insects flitting about, and with worms crawling
through the damp earth, and to reflect that these elaborately
constructed forms, so different from each other, and dependent
upon each other in so complex a manner, have all been pro-
duced by laws acting around us. . . . There is grandeur ir
this view of life with its several powers, having been originally
breathed by the Creator into a few forms or into one; anc
that, whilst this planet has gone cycling on according to the
fixed law of gravity, from so simple a beginning endless forms
most beautiful and most wonderful have been, and are being
evolved.” |
t CHAPTER IV
THE PULSE OF LIFE
RICHARD SWANN LULL
PROFESSOR OF VERTEBRATE PALEONTOLOGY IN YALE UNIVERSITY
THE stream of life flows so slowly that the imagination fails
‘to grasp the immensity of time required for its passage, but
like many another stream, it pulses as it flows. There are
‘times of quickening, the expression points of evolution, and
these are found to be coincident with geologic change. These
coincidences are so frequent and so exact that the laws of
‘chance may not be invoked to account for them. They stand
to each other in the relation of cause and effect.
This does not imply the acceptance of any one philosophical
factor of evolution, for whether the creature is directly modi-
fied by environmental change, or indirectly through induced
habit, or whether nature merely sets a standard to which the
Organism must attain if it would survive, matters not; the
fact remains that changing environmental conditions stimulate
the sluggish evolutionary stream to quickened movement.
Whenever it has been possible to connect cause and effect, the
immediate influence is found to be generally one of climate, !
back of which lies, as the main cause, earth shrinkage and a |
‘consequent warping of the crust, with the elevation and spread
‘of the lands and the formation of mountain ranges. In addi-
tion to this mundane cause, there are the complex rhythms in
solar energy and the consequent variation in the amount of
solar-derived heat. For example, the most generally accepted
I10 EVOLUTION OF THE EARTH |
single cause of the last or Pleistocene glacial period is the
great continental elevation which formed the Cascadian reval
lution, but, so far as our knowledge goes, that would not
account for the successive advances and retreats of the ice
mantle, with the attendant climatic variation; and some other
factor such as the rhythms of solar energy must be invoked as
of supplemental influence. - Nevertheless, the ultimate source
of profound and far-reaching crises in the evolution of the
organic world may have been, geologically speaking, of a very
simple character.
Through the collaboration of my colleagues, Professors
Schuchert and Barrell, the appended chart, Figure 14, has
been prepared to show the relation between the changing con-
tinental elevation and climate. To this I have added a curve
representing the consequent acceleration and retardation of
the evolutionary stream. ‘The climatic records of pre-Paleo-
zoic time are so vague and unreliable that it was not thought
wise to include the vast Archeozoic and Proterozoic eras
within the scheme. The time values of the several included
eras are not proportionally indicated, too much, space by far
being given to the Cenozoic because of its biologic interest.
As a matter of fact, its duration is more nearly comparable
to that of Permian time. a
The curves are necessarily Cater for there was
neither space nor necessity for greater detail. In the altitu-
dinal curve, the upslope of the diastrophic peaks signifies
rising diastrophism; the downslope, the period of erosion
before the continents are low enough to have mantles of sedi-
ment spread upon them. A greater or lesser part of the peak
and downslope represents, therefore, the unrecorded interval
between eras or periods, as the case may be. The tangential.
lines drawn to the diastrophic peaks show the relation of a
revolutions. Were the Cenozoic compressed into its normal
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ries ¢ EVOLUTION OF THE EARTH
relative length, the slope of the tangent drawn to its peaks
would correspond more nearly to those of the Paleozoic.
In the climatic line, the minor undulations indicate in a very
general way the climatic oscillations which must have existed
but of which we have as yet no very exact data; the larger
movements, on the other hand, are based upon observed fact.
The optimum embraces a normal range of temperature and
degree of moisture; a rising curve, increased aridity; its de-
pression, a lowering of temperature. Where the line is single
there is a supposed uniformity of conditions the world over;
where it splits, aridity and cold are differentiated. It is highly
probable that with more detailed knowledge the divergence of
the moisture and temperature curves will be more extensive.
The life line is dotted where theory precedes the recorded
fact, solid when actual fossils attest the truth of the assumed
evolution. The numbers indicate the times of these first
records as follows: 1, first vertebrates; 2, lung-fishes; 3, verte-
brate footprints; 4, reptiles; 5, dinosaurs; 6, mammals; 7,
birds; 8, amphibious dinosaurs; 9, archaic mammals; 10, mod-
ernized mammals; 11, man. (Vide infra, page 144.)
Crises OF EVOLUTION ~ q
Origin of life. The first great crisis in the evolution: of
organic beings was the origin of life: the marvelously subtle
combinations attained by certain very familiar inorganic ele-
ments, increasing their molecular complexity until a substance
was produced endowed with the attributes of life. Of this
momentous event we have no record, nor.does the geologic
cause come either within the scope of our knowledge or con-
jecture, for the time was too remote and the first living sub-
stance too delicate to leave any decipherable record upon the
rocks. All we can say of it is that in the fullness of time, when ~
the earth had, in the course of its physical evolution, become
adapted as the abode of life, living substance came into being.
f
,
t AND ITS INHABITANTS 113
rs Establishment of the lime-secreting habit. The first re-
corded crisis, that of the assumption of the lime-secreting
habit—perfected by animals in the Upper Cambrian, by plants,
“marine alge, much earlier—has again no accepted geologic
‘cause. The importance of this crisis is doubtless not as great
as that of many unrecorded ones which had gone before, its
Seeenncance lying mainly in the fact that the development of
yard parts enabled the organisms which bore them to write
re mperishable records of their existence upon the rocks. Pre-
vious to this, the indications of life are indirect, such as the
accumulation of graphite, which is never produced in nature
except as the result of organic activity; or the records are in
the nature of the rare fossils of algz, of Protozoa (Radio-
aria), and of the burrows of annelid worms. It may be
"assumed that the presence of Protozoa, on the one hand, and
of Annelida, on the other, in late Proterozoic sediments implies
‘the existence of all the intermediate phyla of inverte-
brates—sponges, celenterates, flat and round worms—and the
probabilities are that all the other invertebrate groups except
the Arthropoda were also present.
_ The culmination of this crisis, like several which are yet to
be discussed, did not come until long after its inception, for
while certain creatures like the Archeocyathine (probable
corals) and brachiopods show limy skeletons in the early
Cambrian, others were as yet chitinous, and it is not until the
“Middle Ordovician that the process is entirely complete.
The definite fossil record thus established shows us that
the evolution of great invertebrate groups occurred largely
‘before the close of the Proterozoic, hence we may not speak
| confidently of cause and effect. Vertebrate evolution, on the
| other hand, lies within the period of recorded evolutionary
history, and while there is little direct evidence regarding the
| origin of this important phylum, the evolutionary crises
through which it passed are entirely within the scope of our
114 EVOLUTION OF THE EARTH
observation. That the invertebrates also have advanced si
the Proterozoic is certainly true, but their evolution is lar
one of detail and does not represent the establishment of ;
new principle or type, that is, if we except the insects. Wy
the vertebrates, however, this is not true, for some of 1
most momentous advances in the evolution of higher for
lie within the range of their fossil record.
Origin of vertebrates. ‘The distinction between the ve
brates and invertebrates is largely dynamic, for the former
principally motor types, the latter largely quiescent, slugs
forms, often actually sedentary, that is, fixed in their mode
life. A review of invertebrates, especially the aquatic for ort
will serve to emphasize this point. The sponges are enti
sedentary, while the celenterates are either fixed focal
the corals or capable of very slight creeping locomotion
the sea-anemones; or are feeble swimmers like the jelly-fisk
subject to the whim of tidal and other marine currents, or
the turmoil of a wind-swept sea. Of the worms, using the te
in its old collective sense, some again are fixed, some crawlt
some feeble swimmers, and the same is true of echinode
and many molluscs. The arthropods are perhaps the m
venturesome of marine invertebrates except the cephalo
molluscs, but even to them do the same three condi
fixedness, crawling, or none too effective swimming <
Such as are pelagic are, like the jelly-fish, so non-resistan
to be largely the victims of circumstance. [
Of all aquatic invertebrates, the cephalopods alone |
developed locomotive powers of marked significance, but t
locomotion is strikingly different from that of a vertebrat
that it is the perfection of a method of jet propulsion w
other invertebrates have also developed, although none
carried it so far. Briefly, the perfected cephalopod, suc
the squid (see Fig. 15), has an elongated, spindle-sh;
body, at the hinder pointed end of which are two horizo
rt
AND ITS INHABITANTS IIs
stabilizing fins that may be used for slow forward swimming
by means of wave-like undulations. The body is enclosed in
a muscular fold of the body-wall known as the mantle, but
dees not fill it, leaving a space beneath known as the mantle
cavity. There is on the under side of the well-developed
head, with its huge eyes and long tentacles, a tubular funnel
(Fig. 15, f) whose larger end connects with the mantle
Fic. 15.—Squid, Lo/igo sp., capturing a fish: f, funnel. After Doflein, from
Lull’s ‘‘Organic Evolution,’’ published by the Macmillan Company.
cavity, the smaller one pointing forward beneath the head.
Relaxation of the mantle permits the ingress of water into'the
cavity; closure of the muscular margin around the animal’s
neck, on the other hand, prohibits its egress except through
the funnel. A forcible contraction of the entire mantle forces
the water out in a sharp jet which impinges against the sur-
rounding water and thus drives the creature backward at a
very rapid rate. As compared with the easy undulatory move-
ments of an aquatic vertebrate, however, this jet propulsion
mode is.highly costly of effort, and its adoption by mankind
for steam-propelled craft was to meet peculiar conditions of
short choppy seas and a disastrously ‘“‘racing’’ screw propeller
rather than because of any inherent virtue in the basic idea.
Its rejection from ordinary usage is due largely to its extreme
lack of economy.
The cephalopods, being predaceous forms, must overtake
their prey, and in turn must flee from devouring Cetacea for
which they form a principal dietary staple, hence their urgent
need of locomotion; but for the great host of invertebrate
forms their static habitat places no premium upon locomotive
—
116 EVOLUTION OF THE EARTH !
powers, and the evolution of so marked an’ attribute as high
speed must have some powerful incentive. The thesis has been
advanced therefore that the main distinction between verte-
brate—or more properly chordate—and invertebrate is largely
a response to habitat, the invertebrate being the product of
static waters, marine or sluggish terrestrial, where an effort
less existence, carried hither and yon by wave and tide, wouk C
not remove it from the environment. The chordate, on the
other hand, is the outcome of dynamic or flowing terrestria al
waters which enforced swimming powers as the only means of!
resisting eviction from the realm. ‘The distinction is not on ;
of contour, for the fusiform shape of the speedy animal is th
result of swift movement through a more or less resista
medium, whatever the motive power; but the segmental body
muscles, the internal resistant axis, and the fin-like expansions
to resist the thrust of the muscles, all are the direct outcome 0
the mode of locomotion, that by lateral undulation of a primi.
tively elongated body. (See Fig. 16.) ‘That this locomotive
device might have been developed in static waters is not denied, |
for some marine worm-like organisms swim by a wriggling
movement, though invariably in the dorso-ventral plane, a
there is in the sea little incentive to enforce its rapid perfec.
tion, such as dynamic waters would produce. Hence thi
assumption that the vertebrates are the outcome of terrestrial
waters. To illustrate the means whereby the evolution wa:
initiated, Chamberlin’ has discussed the peculiar habit of a
stream-borne lamprey, Petromyzon (Fig. 16, E, left-hand eel) '
which adheres to the bottom by its suctorial mouth and allow:
its body to undulate in the pulsating current as a flag is is
whipped in the breeze or a rope of grass in the stream. Unles:
the stream be very shallow so as to cause distinct vertical dis
placements of the water, this undulation is always lateral o
1 Chamberlin, T. C., “On the Habitat of the Early Vertebrates.” Jo ur.
Geology, vol. 8, 1900, pp. 400-412. .
1G ee —Fish forms. From ‘The Origin and Evolution of Life,’? by Henry
_ Fairfield Osborn; copyright, 1917, by Charles Scribner’s Sons.
-
Sa gis a Ses) BID 8
abd &
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4
AOR age tn ene a:
[ eee ae
AND ITS INHABITANTS 117
1 ‘the horizontal plane, as that is the direction of least re-
ce because of the inhibitive force of gravity. When the
mprey wishes to swim to maintain its position in the current,
_ simply reproduces i in the active voice the motions imparted
© it in the passive voice by the stream itself. More forcible
ction drives it ahead. Such motion, consisting as it does, of
“series of reversed curves, requires segmented muscles on
ither side, along which alternating waves of contraction may
ass. An axial stiffening to resist the longitudinal compres-
on of such muscles is also necessary. This is first a pliant
ellular membrane rendered resistant by being tensely filled
ith liquid, next a supple rod of cartilage, and finally a bony
xis, segmented for the sake of flexibility. The economic eff-
a y of this undulatory mode of progression as compared
h the jet propulsive method of the cephalopod is evident to
yone who has observed the quick dart of a trout or the
; n: marvelously effortless progress of a school of porpoises at the
_ bow of a ship.
DAs time went on, the assumption of the compact spindle
pape of a swiftly swimming fish became, as we have argued,
rely a speed response and not necessarily an exclusively
am-borne attribute. Depressed grovelling forms (see Fig.
6, C), or compressed highly specialized forms, are, however,
sither marine or static fresh-water in habitat, never stream
wellers.
Tt has been argued that as the most primitive existing
lordates, Amphioxus, the tunicates, and the adelochordates
Balanoglossus), are marine, the ancestral chordates must
ave been also, but their distribution corroborates the hy-
ithesis of fresh-water origin instead of denying it, for as
Matthew” has shown, the most ancient members of a group
re not to be found at the old center of, evolution, but rather
2 Matthew, W. D., “Climate and Evolution.” Ann. New York Acad. Sci., vol.
, 1915, p. 180.
118 EVOLUTION OF THE EARTH F.
at the periphery of their migratory area. This is certainh
true of these protochordates which, save for a few pelag
forms, are all found near the continental margins in t
shallow seas. They are all degenerate and more or les
sedentary, and hence could not maintain themselves in thei
fluviatile habitat and were swept into the sea—descendants o
the unsuccessful early tyros in the art of undulatory swimmin;
But all of this requires as an impelling cause a change ¢
habitat on the part of the invertebrate progenitors, eithend
a migration of the stock from static to dynamic waters, or |
a change in the character of the waters themselves. T
former view entails the necessity of some force to impel tl
migration, either the need for safety or food or the lack
salinity on the part of flowing waters, none of which seer
adequate to compel so radical a change, and one along a li
of so great resistance as the evolution of the invertelam
ancestor into the chordate implied. On the other hand, mig
tion from the sea into sluggish terrestrial waters is mi
readily imagined, .but affords no stimulus for chordate ev al
tion. The picture of the Laurentian peneplain (see Fig. 17
with its low-lying lands and slow drainage, shown by glac
lakes, the result of recent choking of sluggish streams, s st
gests the solution, for while intermigration under such
ditions would doubtless be relatively easy, it would n i :
have stimulated chordate evolution. Geology records a gr
diastrophic movement toward the close of the Protenaaa 0
the so-called Grand Canyon revolution, partial evidence f
which may be seen in the 8,000 to 12,000 feet of sedi nent
more or less conglomeratic, which were swept from mount uit
to the eastward and accumulated in the southern Appalach uu
region during the early part of Lower Cambrian time. ;
great upheaval changed the face of nature in many reg
and quickened the static terrestrial waters to rapid and w
spread movement over all the uplifted lands. Such inver
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SMI[GOId,, WO “Topeiqey ‘newex!yoty axe] Jo pus yynos ay} wos uaas sv urejdaued uenusiney—-y] ‘org
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AND ITS INHABITANTS 11g
_brate stocks as could neither cling to the bottom nor stem the
Muickening current were swept to the encircling sea and lost
to the limnobiotic fauna; such as could, remained to people the
fluviatile realm with creatures of a markedly higher sort.
5 The first recorded chordate comes from the Ordovician, but
is an armored fish-like form, an ostracoderm, of groyvelling
abit and implied quiescent habitat (see Fig. 16, C). Such
“types, Patten* to the contrary, can hardly be considered as the
first expression of chordate evolution, much less annectant
forms between the vertebrates and the higher arthropods
which they so closely resemble; but they were rather a highly
specialized offshoot of the primitive chordate stem, derived
after the physical stimulus of quickened drainage had spent its
force and times of quiescent waters had again appeared.
The evidence points, therefore, to Chamberlin’s conclusion,
that the place of chordate origin was the flowing land waters,
to which may be added as the impelling cause a diastrophic
movement which quickened the drainage, and, as the time of
its inception, the Epi-Proterozoic interval. This crisis is there-
fore the direct outcome of earth movement without the inter-
vention of the climatic factor.
_ Emergence of terrestrial vertebrates. The waters, while
‘a very necessary stimulus to early chordate evolution, afford
too restricted an environment for the evolution of higher
forms, and as a consequence all vertebrates whose ancestry
‘can be traced back through an unending line of water inhab-
itants since the beginnings of life on earth are but fishes, and
no matter to what degree they may have been specialized,
they could not have risen nor can they ever rise to a higher
plane. The emergence from the limiting waters to the limitless
air was absolutely essential to further development and consti-
tuted one of the greatest crises in organic evolution.
To know the route of. such migration is of the utmost im-
*Patten, W., “The Evolution of the Vertebrates and Their Kin,” 1912.
120 EVOLUTION OF THE EARTH
portance if we would find a cause. In search of this, the mind
turns at once to the land-encircling strand or tidal zone, stretch-
ing as it does over the many thousands of miles of the coast,
the omnipresent frontier between land and sea. Here th
conditions are varied, here there is food and light, and heral
the stimulus to double-breathing is enforced by the twice daily
baring of the zone and its consequent transition from aquatic
to terrestrial condition and back again. with monotonous regu
larity. But in spite of its convenience, few indeed have beer
the stragglers of the vast army of aquatic life which have ever
essayed its passage—land crabs and other crustaceans, certain
molluscs, a few of the higher fishes such as the mud-skippers
Periophthalmus and Boleophthalmus, all of which are impelled
to a temporary or permanent migration shoreward by the nee d
of food. And while they have developed accessory respiratory
devices, the latter are merely spongy or other extensions of
modifications of the water-breathing gills, and in no instance i
there developed anything comparable in structure or ultimate
efficiency to the terrestrial vertebrate’s lung. Such structures
as the latter have been developed only in fresh waters, al
fishes which show it or its homologue, the swim-bladder, being
either fresh-water or the descendants of fresh-water forms.
The sharks forsook their ancestral waters before or when the
need of such a structure was felt and have never developed it:
on the other hand, all other fishes except the cyclostomes eithet
have it or have lost it through specialization. In the moderr
fish, the swim-bladder has a hydrostatic function in that i
serves to alter the specific gravity of the creature and thu
enables it to maintain any given level in the water without
further effort. As such, it is analogous to the water ballas
tanks in a submersible ship, although the mechanism whereby
the effect is produced naturally varies. ‘
The lungs of higher vertebrates are therefore identical with
the swim-bladder of the fish and have retained and vastly im.
AND ITS INHABITANTS 121
Bproved the primal function of air breathing. This function
is inconceivable as a prime requisite to marine life, and could
- only have arisen in terrestrial waters beyond the limits of the
tidal zone,* where, due to increasing aridity of climate, the
waters became reduced to stagnant pools which had neither
| sufficient flowage nor wave action to renew the supply of
respirable air exhausted by the contained life. Such conditions
prevail today in Australia, where the recurrent droughts cause
the rivers to dwindle until only isolated pools are left. Within
certain of these rivers, the Mary, Burnett, and Dawson, dwells
an ancient relic form, the lung-fish or dipnoan Neoceratodus,
which in times of habitat stagnation rises to the surface, gulps
SRE ane
ee ee
(eb Re Sets Me 7
+ Fic. 18.—Lung-fish, Neoceratodus, breathing. After Dean.
air (Fig. 18) into its lung, a highly vascular outgrowth or
diverticulum of the alimentary canal, and thus aérates the
. contained blood. Other dipnoans also exist, as the African
lung-fish, Protopterus, of the Nile, which, during times of
_ drought, forms for itself a cocoon-like case of slime-hardened
*See Barrell, J., “Influence of Silurian-Devonian Climates on the Rise of Air-
_ breathing Vertebrates.” Bull. Geol. Soc. America, vol. 27, 1916, pp. 387-436.
122 EVOLUTION OF THE EARTH
mud in the river bottom and estivates therein; and the South -
American lung-fish, Lepidosiren, of the Amazon River and its
affuents, which lives near the margin of the water, using its
lung almost with the regularity of a mammal, and also forming —
a burrow for its habitation during the dry season. Of these
dipnoans the latter two belong to a group of which we have
no fossil record; Neoceratodus, on the other hand, is a relic
of what was formerly a large and widespread group. Yet
another order of fishes in which the air-bladder has a respira-
tory value is the Crossopterygii, or fringe-finned ganoids, again
an important and numerous group in the geologic past but now
are WA CMe Le) Ke LLLE Z
S ROR ane Cae
‘ Sy Lad int R») SRY) Ke
yy rr
BAS Ses DRS RO
Fic. 19.—African fringe-finned ganoid, Polpterus delhexi. After Jordan, from
Lull’s ‘‘Organic Evolution,’’ published by the Macmillan Company.
represented by but two genera, Polypterus (Fig. 19) and-
Erpetoichthys, both tropical African in distribution. While
these living genera have not so effective a respiratory device
as the dipnoans, nevertheless they present fewer anatomical
difficulties to stand in the way of relationship with the am-
phibia. Without rehearsing the technical arguments, it may
suffice to say that the generally accepted view is that the terres-
trial vertebrates were derived either from ancient crossop-
terygians or from a group ancestral to both them and dipnoans.
The geologic cause which lies back of the emergence is
apparent. Diastrophic movement during the Silurian period
( see Fig. 14) initiated a widespread aridity which culminated
in the latter part of the period, continued with varying intensity
into and through Devonian time, and rose again to greater
AND ITS INHABITANTS 123
‘severity in the latter part of that period. This meant, as in
Australia today, the reduction of rivers and other bodies of
fresh water and the entailed concentration of their fauna,
which is borne out by the mode of occurrence of the Lower
Devonian (Old Red Sandstone) fishes—innumerable speci-
‘mens in very restricted areas. Add to this the diminution of
aération of these waters and it will be seen that a high premium
would be placed upon powers of air breathing or of zstivation.
Still further desiccation would necessitate some sort of activity
‘during the increasingly long droughts, for the periods of torpor
_would otherwise bear too great a ratio to the creature’s life
span. Thus a premium would be placed upon ability to crawl
poe ee
=”
ashore and maintain an active life, while the less fit would sleep
the sleep that knows no waking, to their racial extinction.
Aridity, therefore, would place a premium, first upon lung
breathing, ‘the first recorded lung-breathing fish appearing in
the Lower Devonian, although they must have existed in the
Silurian; and later upon emergence, the entire process up to the
perfection of the terrestrial limb covering more than the whole
Devonian period, as the earliest known footprint of a terres-
trial vertebrate is found in Upper Devonian rocks.
Evolution of the terrestrial foot. The question of the evo-
| lution of the pentadactyl hand and foot, which is the vertebrate
standard, from the ancestral fish fin is not fully solved, but
| much light is thrown upon it by the above-mentioned footprint,
known to science as Thinopus antiquus and preserved in the
Peabody Museum at Yale. The slab of sandstone bears a
single track (Pl. III), that of a right foot having two well-
developed digits with distinct phalangeal impressions. On the
outer side of the second digit is a budding third, while lower
down on the side of the foot may be seen the rudiment of a
fourth. Rabl> has shown that the developing foot of the
5Rabl, C., “Gedanken und Studien tiber. den Ursprung der Extremitaten.”
Zeits. fiir wiss. Zoologie, vol. 70, 1901, pp. 474-558.
ONG
Fic. 20.—Development of the hind foot of a
salamander, Triton teniatus. After Strasser,
from. Lull’s “Organic Evolution,’’ published
by the Macmillan Company.
Fic. 21.—Foot of a reptile, Ranodon
sibericus: f, fibula; 4, femur; 4,
tibia; I, Ii, II, IV., digits 1-5.
After Wiedersheim, from Lull’s
‘*Organic Evolution,”’ published by
the Macmillan Company.
PLaTE III.—Cast of the oldest known fossil footprint, Thinopus
antiquus, from the Upper Devonian of Pennsylvania. Original
in the Yale University Museum.
AND ITS INHABITANTS 125
embryonic amphibian, Triton (Fig. 20), passes through a
series of stages of which one corresponds precisely to the
degree of development of the Devonian track, which, from
E size, is probably that of an adult although yet in the
dolescence of its race. Comparative anatomy corroborates
this belief and completes the tale of evidence (Fig. 21), in
that the principal axes of the foot, as shown by the distribution
of nerves and blood-vessels, lie in the first and second digits,
the lesser axes of digits III, IV, and V arising as lateral
branches from that of digit II.
With the opening of Mississippian time came increased
moisture and in the succeeding widely extended swampy forests
of the coal period, amphibia throve mightily and developed
into the many sorts of so-called Stegocephalia or armored
forms. That they still returned to their ancestral waters to
bring forth their young, and that the latter bore gills upon the
eck for aquatic respiration is evidenced by the actual traces
f such structures in many fossil forms.
Fic. 22.—Restoration of the Permian stegocephalian, Cacops
aspidephorus. After Williston, from the Pirsson-Schuchert
**Text-book of Geology,’’ published by John Wiley &
Sons, Inc.
Origin of reptiles. During the latter half of ‘the Mississip-
pian, however, came a diastrophic movement with a wave of
aridity, making the return to the natal waters increasingly
dificult until many forms were forced to abandon it forever,
and the reptiles came into being. There are today well-
recognized criteria whereby a reptile may be distinguished
126 EVOLUTION OF THE EARTH
anatomically from an amphibian, but with certain forms, such
as Cacops (Fig. 22) from the Permo-Carboniferous red beds
of Texas, the distinction is by no means so clear, for such as
this were transitional, Cacops being:‘yet on oe ape side
of the frontier. |
The change of habit induced by this ire meant a radical
change in the character of the vertebrate egg, with the develop-
ment of certain membranes (Fig. 23) whereby the contained
young could respire air, unnecessary in the water-laid egg.
Thus the terrestrial egg, typified by that of bird or reptile, is
large, with an abundance of nutrient “‘yolk”’ for the developing
embryo, and enclosed in a protective though porous shell.
Two membranes are developed, the first of which, the amnion,
is merely protective, in that the cavity between its two layers
contains a fluid which serves to absorb mechanical shocks and
to guard the embryo against too rapid temperature changes.
The other membrane is known as the allantois (see Fig. 23).
This, when fully developed, lines the inside of the shell and
is abundantly supplied with blood-vessels, veins, arteries, and
capillaries, which connect with the embryonic circulation.
This is the respiratory device, for the blood coursing through
the allantoic capillaries is thus in close osmotic relationship.
with the air which penetrates the porous shell, and thus the
blood is purged of its carbonic acid gas and other volatile
impurities and receives its life-sustaining supply of oxygen.
Such an egg must be laid on land or retained within the mother:
until it hatches, for the contained young would drown if the
egg were submerged, as surely as would the adult. !
But the development of these membranes has a still deeper
significance, because the allantois through an added function
becomes the intermediary between the unborn young and its”
mammalian mother, and hence its development, due to that
far-off climatic change, has rendered possible the highe
(placental) mammal with all which that implies to humanity. ~
i AND ITS INHABITANTS 127
_ Origin of warm blood. ‘Toward the close of the Pennsyl-
_ yanian commenced that series of earth movements which was
; to culminate in the Appalachian revolution and the passing of
’ the Paleozoic, and as attendant phenomena came increasing
i aridity and, early in Permian time, glaciation, which, in the
southern hemisphere, exceeded in intensity and extent the
_ so-called glacial period of the Pleistocene.
; Aridity has its influence upon terrestrial types, at any rate
— upon the more progressive of them, in that it places a premium
upon traveling powers, especially upon speed, for not only
are food and water scarce and far between, but the strife
_ between the pursuer and pursued becomes intensified—neither
can afford to be outdistanced by the other. This means in-
creased metabolism, which in turn generally implies not only
_ greater motive powers but higher temperature. With in-
_ creasing cold a premium would be placed upon such creatures
_ as could maintain their activity beyond the limits of the shorten-
ing summers, and this could only be accomplished by the de-
_velopment of some mechanism whereby a relatively constant
temperature could be maintained within the animal regardless
_ of external conditions; in other words, warm, as opposed to
‘cold (really variably temperatured or poikilothermous) blood.
_ This crisis means much, for the cold-blooded reptile has its
most decided limitations. On the other hand, the evolution of
the bird and mammal, the latter particularly, was rendered
- possible by the concurrence of these two factors, aridity and
cold. Thus while actual recorded mammals (Upper Triassic)
and birds (Upper Jurassic) are younger in time, their incep-
tion could hardly have been later than the Permian. Not that
warm blood was at once attained, for that we believe to have
_been a relatively slow process, just as was the emergence.
Indeed, in the existing egg-laying monotreme mammals the
blood still ranges in temperature through at least 30° F., so
‘ with them the mechanism is not yet perfected.
128 EVOLUTION OF THE EARTH
Origin of mammals. ‘These forms are clearly derived from
a reptilian stock known as the cynodonts or theriodonts (Fig
24), in which the dentition has become differentiated as in the
mammals into incisor, canine, and molar teeth. The reptiles
differ from the mammals in that the lower jaw is still a
complex of several bones, while in the mammal there is but
one on either side. Many cynodonts were long of limb and
must have traveled with the body well off the ground, whict
seems to have been a prerequisite to the development of warm
blood. ‘These cynodonts are widespread, but the evidence
points to certain of those of Africa as nearer the direct an -
tors of the mammals; the record of the actual transition
however, is as yet unrevealed.
Fic. 24.—Skull of cynodont reptile, Nythosaurus
larvatus. Note the mammal-like tooth differ-
entiation, but complex reptilian jaw. After
Broom, from the Pirsson-Schuchert ‘*Text-
book of Geology,’ published by John Wiley
& Sons, Inc. .
Origin of birds. Birds, on the other hand, came from
another, unrelated reptilian stock, that which also produced
the dinosaurs. ‘True flight, such as that which the birds de-
veloped, has been thrice evolved among the vertebrates and
once among the invertebrates. In the two other vertebrate
groups, however, the flying mechanism involves not only ni
fore limbs, as in the bird, but also the hinder pair. With th
birds, moreover, the hind limbs, as in dinosaurs, show a dis-
or
Ppietea
ty
ton
| =
Es wr 3
\ htt
z St
ie
> %
z it a
% :
we
s
Fic. 23—Diagram showing the relation of the extra-
embryonal membranes. A, bird or reptile, with
functional yolk-sac and respiratory allantois; B,
mammal with functionless yolk-sac and with the
allantois converted into an umbilical cord and
placenta. From Wilder’s ‘‘History of the Human
Body,’’ published by Henry Holt & Company.
aa AND ITS INHABITANTS 129
‘tinct adaptation of another sort, that for swift movement,
sence the bird-like hind limbs of the dinosaur—or perhaps the
“converse of this statement would be more nearly true. Swift
“movement, which in this instance implies bipedality, evidently
receded flight, and its impression on the bird was so great
hat it has never been entirely relinquished despite the attain-
ment of the higher faculty. Whether birds passed through an
rboreal condition or whether they arose direct from cursorial
ypes is unknown. It hardly seems probable, however, that
“more than one evolution occurred. We may conceive of a
‘proavian as being at least partially arboreal and launching
itself into the air from a convenient tree, sustained for a brief
‘soaring glide on motionless fore limbs and tail, the scales of
which had metamorphosed into buoyant feathers (see Fron-
‘tispiece, A). But the first recorded instance, Archeopteryx
long lizard-like tail of its reptilian forebears, had already
ained the power of sustained flight, though by no means so
Origin of dinosaurs. The same primal influence—aridity—
| which produced the bird also gave rise to the dinosaur.
Whether the latter came within the further influence of glacial
cold and evolved a constant temperature of blood is not known;
the total absence of any heat-retaining clothing in the dino-
That their temperature rose decidedly during their periods of
| activity is, however, a reasonable assumption. Possibly the
| early dinosaur-like forms which dwelt within the influence of
‘the Permian cold became the birds, while those beyond its
influence remained dinosaurs and as such were destined to
‘dominate the lands as no creatures before them had ever done.
130 EVOLUTION OF THE EARTH
of the far-famed Connecticut Valley,* for here there is a pro-
fusion of trails of creatures great and small, quadrupeds and
bipeds, may of which were long of limb, compact of foot, and
impress one as being concerned with weighty matters—ques-
tions of food or safety which would brook no delay. In
several instances the quadrupedal resting posture is clearly
indicated by a creature whose normal gait is that of a biped,
but in no instance is the impressed hand that of a carnivorous
or theropod dinosaur, whose sharply curved grasping talons
borne on the fore limbs would have left a highly characteristic
impression such as is never seen. That carnivores were pres-
ent is an irresistible conclusion, and the inference is that certain
numerous footprints made by the hind feet only, in which the
claws, including that of a grasping hallux, were comparatively
sharp, are attributable to such forms. That aridity does give
rise to bipedality is evidenced by the present existence of
several bipedal lizards, notably in our own Southwest, and in
Australia where the frilled lizard, Chlamydosaurus, reaches
a length of five feet and is exceedingly swift of foot. |
Rise of sauropod dinosaurs. Climatic oscillation during the c
Jurassic gave rise to humid conditions and this, coupled witt
extensive low-lying delta lands along the shores of shallow
seas, tempted certain of the increasingly large Theropoda te
forsake the strenuous life of a carnivore for the slothful ease
of an amphibious herbivore. These huge creatures, because
of the increased burden of the flesh, had reacquired a quad.
rupedal gait as had the armored dinosaurs (stegosaurs).
They suffered no very great alteration in their dental battery,
the teeth of which became spoon-like (Brontosaurus) or, col
lectively, rake-like (Diplodocus), fit only for securing some
sort of abundant vegetation, not for the rending of flesh. The
fate of these forms and that of the armored Stegosaurus wer
8 See Lull, R. S., “Triassic Life of the Connecticut Valley.” Connecticut Stag
Geol. Nat. Hist. Burect: Bull. 24, 1915.
3 AND ITS INHABITANTS 131
curiously linked together, for although unrelated, their habitat
and habits seem to have been very similar and their extinction
as apparently simultaneous. What was the physical cause
of this extinction, which is believed to have occurred in the
‘Comanchian, we do not know, for there is little evidence of
‘climatic change. It may have been a temporary restriction
of habitat, but curiously enough, these dinosaurs managed to
‘survive the uplift at the close of the Jurassic, but are ap-
P oped unknown, with a possible exception in Patagonia,
after the much less extensive Comanchian movement.
All of the most impressive characteristics of both Sauro-
‘ F oda and Stegosauria, huge size, small brain, deficient teeth,
and, in the latter, huge upstanding plates and spines, are to
_ the paleontologist indications of the overspecialization which
1 he interprets as racial senility. With such forms these char-
f cteristics, when coupled with a long period of adolescence
and the consequent slow breeding which may safely be in-
} Wecred, render their extinction imminent.
__ Gadow,’ in speaking of the habits of recent crocodiles, says:
y q “In cooler countries they hibernate in the ground; and in hot
_ countries, which are subject to drought, some kinds estivate
ze in the hardened mud; or they migrate. When, during a pro-
longed drought on the island of Marajé, at the mouth of the
Amazon, the swamps and lakes were dried up, the alligators
‘migrated towards the nearest rivers, and many perished in the
‘ a empt. On one farm were found 8,500 dead, and at the end
_ of Lake Arary more than 4,000. Such occurrences in bygone
times may perhaps explain the masses of bones found here and
are in a fossil state.”
_ From this it will readily be seen how small a climatic change
“might account for the serious depletion of an army of huge
_ forms, perhaps never very numerous as to individuals, and
17 Gadow, H., “Amphibia and Reptiles.” Cambridge Natural History, vol. 8,
1909, p. 447.
>
eat
rae
ee a aoee
oe »
I 42 EVOLUTION OF THE EARTH .
this may have been just enough to give the final death blow to
an expiring race.
Final extinction of dinosaurs. In the later Cretaceous t ie
amphibious habitat was again widespread, if indeed it ke
ever seriously diminished after its first expansion in the Juras-
sic, for here we find the final eflorescence of dinosaurs. The
Sauropoda and Stegosauria had gone, but in their place were
unarmored duck-billed dinosaurs or trachodons, fairly rapid
runners ashore but, judging from webbed feet and compressed
tail, as quick as crocodiles when circumstances forced their
retreat to the waters. Heavily armored dinosaurs, the nodo-
saurs (ankylosaurs) were present, as were the horned Ceratop-
sia, some of which were highly grotesque beings. And to main-.
tain the balance of power there were carnivores, both small and
great, the latter the mightiest beasts of prey which ever walked
the earth. Then comes their dramatic extinction, the world
over, although they may have lost their world-wide dominance
some time before, as we only know these late Cretaceous forms
doubtful Patagonia excepted, from a few European localities
and from western North America; but there they were in ng
climax of their grandeur and there is little save the tendency y
to overspecialization once more to warn the observer of their
coming dissolution. But so far as our records go, not one
dinosaur of all the hosts that were survived the Mesozoic, for
undoubted post-Cretaceous rocks have not yielded a fragmiegy
of their remains.
Why they became extinct no one knows. Our chart show:
a lowering of temperature toward the close of the Cretaceous
and to such climatic changes reptiles are highly sensitive.
However that may be, the great Laramide revolution which
marks the close of the Mesozoic must have brought in a long
chain of attendant events in consequence of which the dinosaurs
perished. Of all factors of which we have knowledge, th
draining of the low-lying coastal lands, with a consequent
Fic. 25.—Tooth of a carnivorous dinosaur,
Allosaurus, and (below) jaw of a contem-
porary mammal, Diplocynodon. Originals
In Yale University Museum. From Lull’s
“Organic Evolution,’’ published by the
| Macmillan Company.
— |) AND ITS INHABITANTS 133
obliteration of the amphibious habitat, seems to have been the
most significant. Dinosaurs have been spoken of as associated
vith peneplanation and the times of their expansive evolution
seem to be coincident with periods of degradation rather than
vith diastrophism, the latter having a restrictive influence.
_ Rise of mammals. Perhaps the most remarkable thing
thich the history of the Mesozoic brings forth is the immense
fetiod of evolutionary stagnation on the part of the mammals.
T ey 2re first actually recorded in the Upper Triassic rocks
f three rather remote localities, North Carolina, Germany,
ind South Africa, and are already differentiated in dietary
Bravics. _ During the Mesozoic, they develop in numbers and
ee certain extent in tooth specialization. They do not, how-
ever, increase markedly in size, but are humble folk, so far
ri as our records have revealed them, until the extinction of the
enosaurs has been accomplished. One cannot but associate
the idea of mammalian suppression with that of dinosaurian
| ¥ ‘dominance in the relation of cause and effect, unless it shall
| “tome day be revealed that the mammals were undergoing a
marked evolution beyond the temperature-limited habitat of
the reptiles. That the former showed no marked evolutionary
advance in the place where the dinosaurs actually occurred is
an attested fact, and the significance of the dinosaurian check
4 is no more graphically shown than by two specimens in the
Yale Museum. These are both of Morrison age, from what
is known i in our records as Quarry 9 at Como Bluff, Wyoming,
| Beccary which has produced a number of the rare mammalian
“specimens. The striking thing, however, is the association of
i. jaws, especially the type of Diplocynodon Marsh, with
‘the tooth of a carnivorous dinosaur, possibly A/losaurus. The
‘figure here reproduced (Fig. 25) is from a simultaneous
photograph of these two specimens, which are therefore on
exactly the same scale. The single dinosaurian tooth greatly
¥
L
‘exceeds not only the tooth of the mammal, but the containing
ate ea
134 EVOLUTION OF THE EARTH
jaw or even the entire creature as the imagination conjures
it up.
The archaic mammals. Dinosaurian extinction heralded
the expansion of the mammals, and-thus the basic cause of the
overthrow of the reptilian dynasty, the Laramide revolution,
was the enabling act in the evolution of the higher race.
Nature now began once more to people land and sea with
beasts of diverse sorts, both small and great; but it was onl
the warm-blooded furry forms whose privilege it was thu
to expand.
The first attempt, following hard upon the dinosaurian ex
tinction, proved to be brief of duration, as though Nature tool
the stock she had at hand without waiting for the coming o
more plastic types. These creatures of the first mammaliar
expansion have been called archaic, in that while capable of
certain degree of specialization, they were more or less stati
in three very essential structures, feet, teeth, and brain. Ce
tain of them were herbivores, some light-limbed, fairly speedy
suggestive of the later cursorial ungulates, although neve
attaining perfection as speed-adapted types. These were th
condylarths (Pl. IV, A). Others were slow-moving, ponde:
ous forms relying upon weapons rather than upon fleetness for
defense. Among these latter, the amblypods, were the swarm
dwelling Coryphodon (Pl. IV, B) and the later Dinocera
(Pl. IV, C), with conservative molar teeth and feet andl
surdly small brain, coupled with elephantine bulk and propo
tions of body and limbs, but with what was superficially
highly specialized skull having many horn-like prominenc
and sabre-like canine teeth—a veneer of specialization over
primitive type. q
The flesh-eaters were in some respects better equipped h
their plant-feeding contemporaries, but they, like the other:
were, if one may judge from skull capacity, notoriously dull al
|
}
AND ITS INHABITANTS 135
stupid compared with the shrewdness of their modern sup-
planters (Fig. 26).
iv Incursion of the modernized mammals. The archaic mam-
mals barely survived the Eocene, only one group, the hyzno-
.. being found in Oligocene rocks: Early in the Eocene,
“however, are seen the vanguard of an army of invaders, none
_ of which seem directly related to the native mammals. Their
Fic, 26.—Restoration of the creodont, Dromocyon. After Osborn, from
Lull’s ‘Organic Evolution,’’ published by the Macmillan Company.
| simultaneous appearance in North America and Europe
points to a contiguous center of evolution somewhere to the
north, either a circumpolar land or the northern part of what
| isnow Asia. Here they underwent their primal evolution and
| here they were endowed with the highest potentialities along
| the three directions wherein the archaic mammals failed.
| Climatic oscillation in the north in the early Tertiary drove
| these modernized mammals south along the three continental
radii, not all at once, but in a series of drives, until the competi-
136 ~ EVOLUTION OF THE EARTH
tion became too severe for the native inhabitants to endure. —
Assimilation of the native stocks by such an invading army is
impossible among animals, however it may be with mankind. |
The archaic mammals, therefore, had but little choice. Some
lingered on, enduring the competition until it became greater
than they could bear, others may have migrated still farther
south to find asylum, which served to postpone their inevitable
fate. Yet others, a very few, may have evolved into higher
types, such as the family Miacide of the archaic carnivores,
although whether they deserve the stigma of genetic relation-
ship with the other archaics in view of this potentiality is
somewhat doubtful. | !
Rise of grazing mammals. The modernized invaders are
now established in their kingdom; they are the early odd-
toed ungulates—horses, rhinoceroses, tapirs; the even-toed un-
gulates, such as camels, deer, and swine; the rodents; the carni-
vores, insectivores, and primates; and in the Old World the
proboscideans or elephants and mastodons. Continental ele-
vation in Europe and more especially in Asia during the
Miocene brought in its train a marked increase in aridity which
in turn had a more or less profound effect upon the flora of
the temperate zones, for it meant a diminution of shrubby
and herbaceous plants and a wide expansion of the harsher
grasses, which now become the dominant note in the world’s
flora. ]
This could not but affect the mammals most profoundly,
especially the hoofed forms. Floral differentiation during the
Oligocene had already made its impress upon certain groups,
such as the horses, so that they in turn were differentiating
along several lines, some with short-crowned teeth suited te
tender herbage, others with grinders whose length and com
plexity forecast the grazing teeth of their successors and whos
dietary choice led in the direction of the coming grasses. Ther
came the floral change of the Miocene and with it a rapid
ys ,
ai hk WW & ae . ~~ z
eet ea = > =e
f/ j XS: s > ‘
> = ws > al
ei “Ss FF ‘
NU r f * 2 tZB ’
. ; TSF Z "
Wog ‘: : oA 1 ois
2 2
Xf i } ES ~ eee “8. 4 4
47, ANS 1 b= " ‘
Win RGA
I My AG ) .
/
Pra
H é
A, : eS
Fic. A.
DPM -F
a) BB Wes a Bs.
: N
FRR yh) WY. 3
“WN =
oe .* ws ~s .
Kony eye aie,
SIA
Fic. C.
PLate IV.—Restorations of archaic mammals. A, cursorial type, Phenacodus
primevus; B, swamp-dwelling amblypod, Coryphodon; C, four-horned
amblypod, Dinoceras, the culmination of its race. After Lull.
we
AND ITS INHABITANTS 137
‘expansion of grazing forms—horses, camels, deer—and the
Meescriction and often the extinction of browsing types. It is
_ true that browsing forms are still extant, but not in their old
_ profusion nor in their old homes, while the grazing forms are
‘numerous and characteristic of the widespread steppes the
world over. |
5; Origin of man. We have observed the influence of geologic
‘change in the evolution of the brute, and we have now to
i aquire whether mankind in his long upward course has been
| "amenable to those same laws or whether he has been a thing
“4 part from other forms of life, whose development has been
controlled by other influences. As the primates, the group to
ipbich mankind belongs, are to be classed with the modernized
‘mammals, their course of evolution up to the point of their
d differentiation as primates must have been one with all the
‘test and hence the result of the same chain of causes. And
4 their differentiation from the other mammals when they came
tC 0 the parting of the ways seems to have been due to the de-
parture of the latter from their primal mode of life and
structure rather than to any special evolution of the primates
‘themselves, for in many ways they are among the most primi-
tive of the modernized hosts, and their tree habitations may
well have been a very ancient habitat of the whole mammalian
_race.®
They throve in their northern home just as did their other
| compatriots, and like them drove southward along the several
~ continental radii, the rear guard drawing in toward the equator
| with the northern limits of the tropical forests within which
| they dwelt and upon which, with rare exceptions, they are
| _ dependent to this day for food and safety. They reached
' North America, as the map (Fig. 27) indicates, early in
* Eocene time (Wasatch) and became so abundant as to form a
|< 8 Matthew, W. D., “The Arboreal Ancestry of the Mammalia.” Amer. Nat.,
vol. 38, 1904, pp. 811-818.
138 EVOLUTION OF THE EARTH
high percentage of certain fossil faunas. Here they throve
until the close of the Eocene, when they died out, and North
America knew no more primates until the coming of man.
They had, however, crossed the isthmian land-bridge or its
equivalent into South America, where they still persist.
DISTRIBUTION OF PRIMAT
ES Movern Anturopoinea {Mont ys APES BAB
Tn ” Lemuropta (LEMURS ,LORIS, TARSIER
E, Eocene (ano Oligocene Lemuroins
0, Oricocene AntHropoins
| M, Miocene
P, Puocene ”
Fic. 27.—Map showing the geographical distribution of the primates, living
and extinct, and their indicated dispersal from Holarctica. After Matthew,
from ‘‘Problems of American Geology,’’ 1915, published by Yale Uni-
versity Press.
In Europe, the history of primates below man was similar —
to that in North America, marked by Eocene abundance and —
synchronous extinction, but during the Miocene, at the time —
of the proboscidean migrations, they reappeared, probably ~
from Africa, to suffer a second extinction during the Pleisto-
AND ITS INHABITANTS 139
zene. While the sub-human primates are now extinct on the
“Asiatic mainland north of the great Himalayan uplift, they
till persist in tropical Asia, in India, Indo-China, the East
indies, Japan, and the Philippines. ‘ted are also extant in
ropical Africa.
_ Anthropoid apes. Man’s nearest blood relatives, whatever
lay be his prejudice in the matter, are the so-called anthropoid
i‘. man-like apes: the orang, chimpanzee, gorilla, and gibbon,
i descendants from the same stock which gave rise to hu-
lanity and perhaps fallen to their present condition through
being the victims of circumstances. That man owes his higher
estate, at least in part, to his past environments we shall
‘endeavor to show.
% eof the great apes, the orang-utan is one of the most familiar,
or it and the chimpanzee are the kinds most frequently seen
| n captivity. The orang is readily distinguished, however, by
its reddish hair. It is rarely more than four feet tall but is
C elatively of great girth, which, together with an arm spread
: f upward of seven and a half feet, gives it a remarkable
ppearance. ‘The jaws are powerful, with large canine teeth,
but the hands are the chief weapons of defense. The great
of the orang renders it less agile than the gibbon, for
4 : ance (see page 140), and it climbs somewhat laboriously as
does a man. It is highly intelligent but sluggish, stirred to
r ction only by some powerful incentive such as hunger.
‘Orangs are today confined to the dense somber forests of
Bomeo and Sumatra, but the fossil remains of their ancestors
some from the Siwalik Hills on the Asiatic mainland and thus
| etray the course of their migration.
, B The chimpanzee and gorilla are both African in distribution,
| alt hough also of Asiatic stock. ‘They are perhaps the nearest
to mankind in blood relationship. The chimpanzee may
readily be distinguished from the orang by its black hair. It is
so taller, although never exceeding five feet, and is less bulky
as
im
= ina = J
.
140 EVOLUTION OF THE EARTH
and hence more active, swinging from tree to tree by the han¢
with great agility. They rest in the sitting posture, sometim
stand or walk, but run on all fours. Jaws and teeth are *
chief defense.
The gorilla (Fig. 28) is by far the most. formidable of 4
the great apes, for its huge size and strength and its unparz
leled ferocity make it a veritable terror. An authentic rf
corded size makes the gorilla five feet one and one-half inche
tall and 418 pounds in weight. Its lower limbs, whi
enormously powerful, are disproportionately short. If
limbs bore a human proportion to the torso, the creatur
would stand at least seven feet in height, with a weight 1
half a thousand pounds! The huge size of this ape has fore
it to become partially terrestrial, but instead of becomi
more man-like as an adaptation to ground-living, it has tak
a yet more brutal aspect, more like a bear than a human bein
All of these apes, the orang, chimpanzee, and gorilla, ;
degenerating from the higher condition of their comm
ancestor with mankind, the chimpanzee least, the gorilla me
of all. The gibbons (Fig. 29), however, of which there ar
several species, while the most remote from mankind in actt
relationship, have probably retained in greater degree th
any others the habits and development of the anthropoid stem
form. They are wonderful acrobats, their relatively small siz
and immensely long and powerful arms lending themselves t
the full measure of arboreal progression. The gibbons ar
oriental in distribution, living in the wooded regions of south
eastern Asia and the islands of Sumatra, Borneo, and Java. _
It were well to dwell for a moment upon the locomotiy
methods of these apes which, instead of running upon th
upper side of the branches, as do most arboreal forms, swim
beneath them by means of their hands. This method of loce
motion has been called brachiation (Lat. brachium, arm) am
in all probability took its rise with the earliest anthropoids
Fic. 28.—Gorilla, Gorilla gorilla. From ‘“The
American Natural History,’? by William T.
Hornaday; copyright, 1904, 1914, by William
T. Hornaday; published by Charles Scribner’s
Sons.
AND ITS INHABITANTS EAI
aching its highest development in the modern gibbon. On
1e ground, the gibbon walks erect, either touching the knuckles
f the hands to the ground or with the arms held above the
ead. The gait is quick, waddling, with no elasticity of step,
nd they are soon overtaken. But in the trees they are virtu-
lly transformed, for their hand leaps are prodigious, twelve,
ighteen, one authority says no less than forty feet being
eared, and that for hours at a time. Fully to appreciate
hat this means one should compare it with the precise
iechanical stride of a racehorse, for whereas in the horse
tere is a practical uniformity of conditions stride after stride,
vith the gibbon no two hand leaps can be the same, and each
me such a thing is essayed a problem must be solved, for in
rder neither to over- nor under-shoot the mark, the right
+ oo of force must be used, and this varies with the distance
ind the ability of the objective branch or branches to bear
&. creature's weight. Thus, as with a gun pointer, the aim,
stance, trajectory, windage, and in addition the varying
force, all must be taken into consideration; in the present
instance, moreover, the problem must be solved and its prac-
tical application brought about instantly, and the penalty for
error either in solution or application may be death. Even
though the process becomes automatic, the result of much
i experience, there is, nevertheless, a high premium placed upon
ee tal acuteness, and the weeding out of the unfit is ruthless.
A \dd to this the fact that the hands may be used to bring objects
before the face for examination, thus inciting the powers of
Observation, and two great stimuli to higher mentality are
- tained There is reason to believe that the human precursor, |
| before leaving the sheltered life of an arboreal primate, pro-
4 ressed and acted much as do the gibbons, with a consequent
kening of intellect as time went on.
| Descent from the trees.. But while tree-life had much to do
witl the prehuman evolution of our ancestors, the arboreal
142 EVOLUTION OF THE EARTH
is, after all, a limited environment and the descent to the tet
restrial habitat was as necessary to further evolution manwards
as was that older emergence .upon land on the part of ou
piscine ancestry. In the former case as in the latter, the actual
attainment of the terrestrial habitat is supposed to have been
forced by geologic change of a very similar character. Th
presumption is that central Asia was the evolutionary cente
of the anthropoids and so far as our records go this is born
out by the fossils, the oldest of which are found in the Siwali
Hills of northern India in rocks of Lower Pliocene age. Fr on
this center of radiation these primates took their departure
the gorilla and chimpanzee southwestwardly toward the Darl
Continent, while the gibbons and orang went toward the south
east. The prehuman, on the other hand, remained in centra
Asia nearest the dispersal center, which, we have seen, 1
generally true of the latest and most highly specialized of
race. The forms which migrated southward felt the enerva
ing languor of the tropics and remained static, if they did no
actually retrogress, whereas the prehuman in the more invig
orating and variable climate progressed in his evolution towar
a higher type. Central Asia has proved to be the theate
wherein the highest mammalian evolution could be attainec
for here the forms which man has chosen for his companion
the domestic animals, almost without exception attained thei
evolutionary completion.
The actual descent from the trees seems to have been dt
to a chain of events which in many ways parallel those which
impelled the emergence.? The Miocene uplift, with the co
sequent aridity, has been mentioned. As a direct result, th
stupendous barrier of the Himalayas began to arise, cuttif
off the forests of central Asia from their old-time contin
with those in India, and thereby severing the lines of com
® Barrell, J., “Probable Relations of Climatic Change to the Origin of t P|
Tertiary Ape-man.” Scientific Monthly, January, 1917, pp. 16-26. =
Fic. 29.—Gibbon, Hylobates lar.
Fret ect mag
| AND ITS INHABITANTS 143
munication by which the anthropoids had left the cradle of
their evolution. Miocene and Pliocene aridity diminished the
northern forests, as Devonian aridity had diminished the
terrestrial waters, until finally there were detached wooded
areas within which the contained primates were as isolated as
is the orang on sea-girt Borneo and Sumatra today. Further
diminution compelled the descent to the ground on the part of
the larger and more intelligent forms among them and the
destruction of such as could not meet terrestrial competition.
Thus a man-like tree-born primate became an earth-borne
creature which from sheer necessity ultimately arose to man’s
estate. This meant the assumption of the erect posture and
the lengthening and strengthening of the lower limbs for speed,
while the hands, released from the fetters of their earlier loco-
otive function, became the organs of the mind. Now man
had to compete with the mighty carnivores, and as nature had
ill endowed him with defensive weapons he had to devise crude
armaments of stick or stone with which to insure his survival.
[he dwindling forests, especially as their tropical character
nad gone, no longer supplied an easily gained livelihood, and
sustenance had to be sought from other sources, mainly from
among the feebler of man’s fellow creatures. It is probable
at he also utilized the seeds of grasses which grew along
the margins of the forests, the ancestors of our domesticated
2ins.
Increasing severity of winters, prophetic of the Glacial
Period, necessitated some further means of keeping warm,
and man utilized for clothing the skins of beasts which he had
slain and later harnessed fire to his use. In common with the
other primates, the precursors of mankind were doubtless
gregarious, probably submitting to the leadership of such as
iby strength or cunning could enforce their authority, but not
jantil the terrestrial life was attained was the higher communal
ife with codperative division of labor possible.
144 EVOLUTION OF THE EARTH
By means of this codperation and his individual prowes
man has attained dominion over the organic world, whil
through his subduing of nature’s forces he conquers all o:
her varied realms from the equator to the poles, adapting him
self to environments which mold their creature denizens most
profoundly and through which he passes unimpressed by the
laws which compel their adaptation, for he is swifter than th
eagle in his flight and he passes through the waste of the
waters unscathed by nature, a victim, if such he be, only t
others of his own kind (Fig. 30).
SUMMARY 4
As the physician, by a clever device, can record graphically)
the pulsations in the blood stream which are synchronous wit
the throbbing of the human heart, so I have drawn a curve te
show the correspondence between the pulse of life and thi
heavings of the earth’s broad breast. In this curve (Fig. 14)
the evolutionary crises and periods of expansive evolution ar
represented by upward movements, while extinctions are indi
cated by the downward plunge of the racial line. ‘This curve
is broken where as yet we have no tangible evidence to validat
our theory, solid when the fossil record changes theory int
fact. if
Thus the first recorded crisis, the origin of vertebrates, i
drawn coincident with the revolution at the close of the Pro
-terozoic era, but the actual vertebrate record begins in mid
Ordovician time at the numeral 1. The development of lung
is connected with the disturbance at the close of the Siluriai
uplift, the first recorded lung-fishes appearing at the numeral 2,
in the Lower Devonian, while the first footprint at 3, in th
Upper Devonian, completes the evolutionary movement. Th
Mississippian is a time of expansive evolution of the stego
cephalians, while the establishment of the reptiles was th
result of aridity toward its close, hence synchronous with th
°
Fic. 30.—Photograph of a submarine,
twenty feet below the surface, taken
from the aeroplane whose shadow is
shown in the picture. From London
Illustrated News.
iL AND ITS INHABITANTS 145
uachita uplift. The culmination of the wide disturbances at
the close of the Pennsylvanian brought on aridity and glacia-
on in the early Permian, with the beginnings of the warm-
ooded stocks, birds and mammals, although the first recorded
rd at 7 is Upper Jurassic and the first known mammals at 6
rp per Triassic. Aridity has been regarded as the chief incen-
tive to dinosaurian evolution, the actual record at 5 beginning
in the Lower Trias. The differentiation of the Sauropoda, a
res ponse to the expansion of the amphibious aquatic realm, may
uve been Triassic, but in greater probability the subsidence
hich followed the Palisade diastrophic movement late in
le Triassic may be nearer to the actual time. They first
& | spear in the Lower Jurassic, but do not culminate until early
omanchian, shortly after which they suffer extinction from
own causes, geologically speaking. The other dinosaurs
radually i increase in numbers, size, and specialization, until
th oe of the Cretaceous, when, with the movements which
ninate in the Laramide revolution, en plunge into extinc-
#1
Nn.
| The curve of mammalian evolution is perhaps the most
striki ing of all, consisting as it does of minor fluctuations which
nay represent accident of preservation and recovery rather
periods of evolutionary expansion, as is equally true of the
z osaurian curve. Then with the Laramide revolution and
he coincident dinosaurian extinction comes the rapid mamma-
Tian deployment, the archaic mammals, first known in the basal
cene at 9, becoming fewer and less important, although
er casing in specialization until their final extinction in the
Middle Oligocene. With the fluctuating climate of the north-
ern area come the modernized forms, first known in the Lower
Eocene at 10, then the differentiation of browsing and grazing
for s, followed by a restrictive evolution of the former and
wide expansion of the latter consequent upon the Miocene
lift and its resultant aridity. While the primate line is solid,
146 EVOLUTION OF THE EARTH |.
the derived human line is broken to the point 11, the
actual record of man; the descent from the trees is, howeve
made coincident with the Himalayan uplift and with the i
creasing aridity of the Miocene.
Man’s intellectual and spiritual rise and his dominanen ove
the forces of nature and over the brute creation are shor
graphically by the height to which his line ascends above the
highest crest of the evolutionary crises which have gone befor
For this dominance the Cascadian revolution with its recur e
periods of devastating cold, out of which we are emergin
must be looked upon as a contributory cause. 7
Thus time has wrought great changes in earth and sea, a
these changes, acting directly or through climate, have alw
found somewhere in the unending chain of living beings cert:
groups whose plasticity permitted their adaptation to the ne)
arising conditions. 3 i
The great heart of nature beats, its throbbing stiniul eee :
pulse of life, and not until that heart is stilled forever will t
rhythmic tide of evolution cease to flow.
CHAPTER V
CLIMATE AND THE EVOLUTION OF
CIVILIZATION
ELLSWORTH HUNTINGTON
RESEARCH ASSOCIATE IN GEOGRAPHY IN YALE UNIVERSITY
FACTORS IN HUMAN PROGRESS
N progress depends upon three great factors. The
I taherent mental capacity, the second material resources,
oh third energy. If any of these is lacking, civilization
lates or even retrogrades. Where all are present civiliza-
| | moves onward. How far do these three depend upon
sical environment? How far does man’s higher evolution
’ with the evolution of plants and animals discussed in
vious lectures? At first sight it appears as if there were
teat gap between the evolution of man’s body which this
rse of lectures has thus far considered, and of the mind
ich is the subject now before us. In one sense there is
oubtedly such a gap. Yet the more we study the matter,
more we see that from the lowest protozoan to the highest
osopher a marvelous unity pervades all nature.
ll progress in civilization, whether material or moral,
arises from ideas in the minds of individuals. Therefore the
st requisite of any advance is men with unusually gifted
minds. Some races seem to be capable of producing such men
jn far greater numbers than do others. We rightly think of
me¢ient Greece as preéminent in this respect. Galton, the
founder of modern eugenics, has said that the average Athe-
eS a = «an = = ee
148 EVOLUTION OF THE EARTH
nian in the age of Pericles was as much superior to the averag
Englishman of today as the Englishman is superior to th
African. Although a few students still claim that all race
possess equal possibilities if given equal opportunities, thei
conclusions lack statistical foundation. Judged by thei
achievements and by all the exact tests yet available, race
differ as do individuals, although not to so great an extent. —
What causes these racial differences? We cannot answe
until the biologists give us more light on the origin of the ney
forms called mutants. If it be asked, however, what pr
serves the mutants and thus gives rise to new racial qualitie:
we can answer with considerable certainty. Environment b
means of natural selection allows some types to perpetuat
themselves indefinitely, while it rigidly exterminates other
Among the various environmental factors, climate is appar
ently the most important. As Professor Lull has said in
previous lecture: “Changing environmental conditions stim
late the sluggish evolutionary stream to quickened movemen
Whenever it has been possible to connect cause and effect,
the immediate influence is found to be generally one of
climate.” e
Inherent mental capacity. The American Indians seem t
furnish one of the best examples of the influence of clima
upon mental capacities. Practically all authorities agree that
the Indian is endowed with a relatively conservative type «
mind. He has great powers of observation, of patience,
endurance, and also of action when some crisis suddenly sti
him. He is lacking, however, in originality, in the power |
adaptation, and in the quick insight and inventiveness whit
make the Japanese so competent in seizing what they want
European civilization. It has been suggested that the mental
capacities and traits of the Indians have been modified by t
1 For a fuller discussion of this matter see “The Red Man’s Continent,” vale :
University Press, 1918.
z AND ITS INHABITANTS 149
yay of Bering Strait. If that is the case, they must have spent
many generations in the extremely trying environment of the
ar North where the January temperature averages 10° F.
below zero, and where the winter night lasts months. Such an
upon itself and in time gives way. ‘The type that survives is
the phlegmatic man who can sit idly for weeks inside his stuffy
hut, and not care whether anything happens or not. When
they left the primitive home of man in Asia, the ancestors of
the Indians presumably had minds like those of their neighbors
who became the fathers of other races. When they emerged
from their long sojourn in the Far North, however, they had
lost some of their most valuable qualities.
_ In the same way the European Nordics possess the type of
mind to be expected of a race that has always lived in a stimu-
lating climate. The Japanese show similar characteristics to
an almost equal degree. It is significant that although these
two races pushed out from central Asia in opposite directions,
neither was ever forced far to the north or south, and each
finds its present home in one of the world’s best climates.
Neither race, however, has evolved in a uniform climate, for
changes due to the Glacial Period, especially in Europe, have
forced them to endure repeated epochs of stress. The stress,
however, was of a stimulating kind because it was apparently
characterized by variability and not by the monotonous uni-
formity of the Far North or the equator.. The African
negroes, on the other hand, have by no means been so fortu-
nate. Because their migrations led them into southerly regions
|they suffered a repressive evolution much like that of the
Indians. In tropical regions the energetic types unfortunately
150 EVOLUTION OF THE EARTH
kill themselves by overexertion. Activity accelerates the pro
esses of metabolism and generates toxic poisons. In the right”
kind of climate these are eliminated during periods of rest.
a warm climate, however, the high temperature appears to’
cause excessive chemical activity of the protoplasm just as does
exercise. Hence people feel tired even without exertion. When
bacteria attack such people, they find a ready prey, and when
the effects of activity and of heat are combined the result is often -
fatal. The exact mechanism of the process has not yet been de-
termined, but some such poisoning of the system and consequen
elimination of unduly active types appears to be one of thx
reasons why the negro has acquired a comparatively indolen
character.
Among the backward natives of Australia the elimination o
energetic, nervously alert people has gone farther than among
the negroes. The Australians in crossing the torrid zone were
subjected to all the evils which have weakened the mental
powers of the negroes. They also suffered a terrible handicap _
because their tropical experience was the precursor of af
equally strenuous repression by the desert. There sudden and
intense activity is at a premium when the water dries up and
a long march must be made to a new supply. The most
essential of all qualities, however, is the ability to endure |
hunger, thirst, and heat indefinitely, a kind of endurance which |
is much harder on people with alert nerves than upon those
of stolid disposition. Moreover, mental alertness loses muc
of its importance as a factor in natural selection when th
environment becomes so poor that there are almost no materia
resources. It is by no means strange, then, that the Aus
tralian bushmen, even more than their fellow sufferers, t
Hottentots of South Africa, show, as it were, the combi
weaknesses of the tropical negroes and of the desert peop
of Arabia. One might go on to discuss this theme in relatio
to all the races of the earth. Such a discussion would appat
ently strengthen the conclusion that while the mental inhe it.
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OS SS a TN aD aN eal
AND ITS INHABITANTS 151
‘tg nce of an individual may show no apparent relation to his
4 ipress of the ieatit environments through which his race
has passed.
Material resources. Material resources are no less im-
portant than innate capacity in their effect upon civilization.
1 hey are, if anything, still more closely connected with climate.
A Socrates with no resources except the sands of Arabia to
support himself and his ideas would never have been heard of.
ohammed had to live in an oasis and not in the sand. His
religion would not today dominate nearly a seventh of man-
kind if it had not speedily spread to places with abundant
material resources. The Eskimos show the importance of
material resources still more clearly. Though possessing the
same power of passive, nerveless endurance which is character-
ow of the Indians, these people of the snow and ice seem to
“have a strain of inventive ability. Their stone lamps, their
drills for lighting a fire by twirling one stick upon another,
_their clever boats which make a man and his canoe parts of
a single watertight structure, all show the earmarks of in-
‘genious minds. Nevertheless, civilization has not made prog-
“ress among the Eskimos. Even if they were not afflicted with
the inertia of the North, the absence of material resources
almost forbids a high civilization. Here, as in so many other
“cases, climate is the factor which mainly determines the re-
“sources. No crops will grow and even the reindeer does not
thrive i in many parts of the Eskimo coasts, hence hunting is
- the only possible mode of life. Hunters must be nomadic.
‘They cannot accumulate any large amount of the material
Bessources which are needed as aids to progress. If the white
“man with his claims to superiority were placed in the home
“of the Eskimos with no outside resources, would it be more
than a few generations before his mode of life and manner of
thought would be much like theirs? The retrogression of the
oo
|
2
Teo EVOLUTION OF THE EARTH |
white fishermen in northern Newfoundland and Labradoi
where the conditions are much better than in the home of the -
Eskimos, is a pointed answer to this question. |
The effect of climate upon material resources and of ma- |
terial resources upon civilization is well illustrated by a com- —
parison of the Indians of California and Utah with a
Pueblo neighbors in New Mexico. The pre-Columbian in-
habitants of California and Utah were abject savages. a
|
dwelt in flimsy brush huts, and ate rabbits, lizards, grass-
hoppers, acorns, and other equally poor kinds of food.
Because food was so scanty they lived in very small comma
ties, and were forced frequently to move from place to place. |
Because of their wandering, isolated life they had only the
rudiments of social and political organization. All in all,
they were well-nigh the lowest of the American aborigines.
The Pueblo Indians, on the other hand, had risen far beyond
mere savagery and were well along in the stage of culture -
known as barbarism. They had an excellent diet of corn,
beans, and squashes, with enough meat to keep them in health. |
Not being obliged to wander, they lived in compact, well-made
adobe houses. Their villages were large enough and neal
enough to one another so that intercourse was frequent, hence
they had a highly organized social and governmental system.
They had also developed a series of complex religious cere-
monials that did much credit to their mental powers. Among
the aboriginal people of America they stood not far from |
the top. é |
The contrast between these two neighboring types was ap- :
parently not due to racial differences. The Hopis, who were —
among the most advanced Pueblo people, were apparently _
of the same race as the Utes, who were among the lowest of —
the other type. Material resources seem to have been the _
main cause of the contrast. ‘The Pueblos had behind them |
the resources of agriculture, while their savage neighbors —
bey
f
: AND ITS INHABITANTS 153
could not raise crops. This difference was due to climate. At
‘first sight the climates of the two regions appear closely
similar. Both are dry. Salt Lake City, in the home of the
| Utes, for example, has an average rainfall of sixteen inches
per year. Los Angeles, where lived some of the lowest Cali-
fc ornia Indians, has the same. Santa Fé, in the center of the
_ Pueblo district, receives a yearly average of nearly fifteen
inches. Why, then, was agriculture and hence civilization so
different in the two types? The answer is found in the
_ season of rainfall. At Santa Fé the months of June, July, and
August have an average of 8.4 inches of rainfall, whereas at
- Salt Lake City the same months average only 2.2 inches, and
_ at Los Angeles are practically rainless.
| s _ Today the greatness of California depends upon its farms
more than upon any other material feature; and the farms of
Utah are also by no means to be despised. It must be remem-
yered, however, that wheat, barley, beets, grapes, oranges, and
Bother orchard fruits are the staples of agriculture in these
} # ‘regions. They are raised largely by means of elaborate sys-
_ tems of irrigation which utilize the winter snows upon the
mountains. Where irrigation is not practised, cattle are the
‘great agricultural resource. The other foundation of pros-
perity is mining. All these things are essentially European.
The crops which place parts of California and Utah among
‘¢ he world’s garden plots are not indigenous to America. The
_ cattle and horses which browse on a thousand brown hillsides
5 had no counterpart in the New World before Columbus. Since
iron tools were unknown, the art of mining was impossible. So
¢ ar as anyone has yet pointed out, the poor Indians had no
& plants that would serve in place of those with which the early
¥ people of the Old World were blessed. The absence of sum-
ia mer rain which is typical of all subtropical climates caused the
" vegetation to be scanty, and hence wild animals, wild seeds, and
fruits were also scarce. Thus the people were condemned to be
154 EVOLUTION OF THE EARTH
nomadic hunters and to remain in the lowest stage of civiliza-
tion. :
Turn now to the Pueblo region. A rainfall of six or eight —
inches in the three summer months seems insignificant. Yet
it made all the difference between savagery and barbarism. _
In its absence the Utes and the California Indians were
as much below the Hopis and the other Pueblo tribes as the _
Pueblos are below modern Europeans. The one great cereal
of the New World is Indian corn. The two other staple —
crops capable of being raised in the dry regions of pre- |
Columbian America were beans and pumpkins. All of these —
can grow where there is moisture in summer, but not otherwise. _
To raise them by irrigation in places like Utah and California —
is practically impossible for primitive people. On all except
the larger streams, whose control is beyond the power of —
beginners, the floods come to an end long before the time when —
the crops most need them. Where the Pueblos live, however, —
the summer rains produce floods just when the corn, beans, —
and pumpkins are most eager for water. Hence these crops
provided the early Americans with the basis for a well-
developed system of agriculture, and progress was assured. |
Even if the Utes and the Indians of California had been more _
inventive than the Pueblo Indians and had produced men with _
minds of unusual brilliancy, the lack of material resources —
adapted to the early stages of development would probably ©
have kept them permanently in savagery. 5
The material resources thus far mentioned depend upon
climate. In this they are like nine-tenths of the world’s re-
sources. All kinds of food, all kinds of clothing, and a large
part of the materials used for shelter depend absolutely upon
climate. Perhaps there is some exception to this statement,
but I have searched for it in vain. . The fish of the sea, the
fowl of the air, and the beasts of the field, even though they |
may prey upon other animals, are in the last analysis wholly }
e AND ITS INHABITANTS 155
_ dependent upon the vegetation which feeds either them or
_ their prey. Vegetation is absolutely dependent upon tempera-
' ture, rainfall, winds, ocean currents, and other climatic con-
_ ditions. Soil, to be sure, is of much importance, but all soils
_ would be moderately fertile if they had never been subjected
to any except a favorable climate. There would be no great —
__wastes of sand, salt, gravel, or naked rock, for these arise
_ only under aridity, glaciation, high wind, or other extremes.
; In spite of the overwhelming preponderance of climate in
_ determining the nature of material resources, certain articles,
__ such as iron and coal, occur with little reference to heat or cold,
_ moisture or drought. Their use, however, depends largely
upon climatic conditions. Iron was well known in Mediter-
_ ranean lands at the time of their highest civilization. Never-
theless, its use was strictly limited by the scarcity of fuel in a
_ subtropical climate. Only when iron was smelted in countries
like England where the climate fostered dense forests did its
__use increase rapidly. When the value of coal was at last
realized it added wonderfully to man’s material resources.
_ Yet even here the limitations of climate are strict. In Alaska
_ today the presence of coal does little to advance civilization.
"It brings a group of coal miners whose chief desire is to make
P money enough to enable them to get home again. Because
_ Alaska is so cold and inhospitable the regions with better
| i climates farther south are the ones that profit by its coal. So,
__ too, with the iron mines of Gellivara in northern Sweden. A
. _ few small towns have sprung up, but the places chiefly benefited
4 are the cities of southern Sweden where the climate is less
_ Severe. Thus it appears that all kinds of resources are strictly
| limited by climate either in their occurrence or in their use.
| Energy. The importance of energy in advancing civiliza-
tion is not so well understood as that of inherent mental
v "capacity and material resources. Darwin says that the latent
_ capacities with which most men are endowed are so great
en -z 0 .
156 EVOLUTION OF THE EARTH
that inherent ability is less important than the zeal and deter-
mination, or in other words, the energy with which that ability —
is used. In our own experience have we not scores of times —
had good ideas—perhaps revolutionary ideas—with which we —
toyed for an hour or a day, only to forget them because we had ~
not the energy to undertake the hard work necessary to bring —
them to fruition? Such energy is needed not only by the indi- —
viduals of unusual capacity who make the great inventions, —
but also by the people around them who must put the new —
ideas into effect. ‘We have but partly solved the mysteries ©
of the progress of civilization —when we have pointed out —
that each tangible stage of progress owed its initiative to a —
new invention or discovery of science. To go to the root of —
the matter we must needs explain how it came about that a
given generation of men was in a mental mood to receive the —
new invention or discovery.” The necessary mental mood
is ‘alertness,’ which is merely a manifestation of energy.
Therefore it would seem that in the march of civilization —
energy is quite as important as either inherent mental capacity _
or material resources. Energy is partly the result of proper —
food, clothing, and shelter, partly the result of inherited
ability, and partly the result of freedom from specific disease. _
All these things depend in large measure upon climate. Back —
of all, however, lies something else. Everyone knows that |
he may be well fed, well dressed, well housed, of good ancestry, —
and free from specific disease, and yet may not feel like wor
even when he has no special occasion to be tired. Apparently
the thing that is needed is the stimulus of the right kind of —
climate. Health is one thing; full efficiency is quite another.
Effect of climate upon man’s efficiency. This matter is so
AND ITS INHABITANTS 157
merable indirect effects through inheritance and material re-
ssources. In this he appears to be like all other organisms.
For every species of living being there is apparently a certain
optimum or most favorable condition. As the temperature,
humidity, or other climatic elements depart more and more
widely from the optimum, the animal’s reproductive rate
diminishes, its general strength declines, it becomes more
susceptible to disease, and its life is shortened. Between the
optimum conditions and those which cause death there may be
| a wide range in which life and even health are possible, but
in which the organism is not at its best.
One of the great lessons of biology is that man’s physio-
logical nature is essentially the same as that of the lower
animals. A law which applies universally to them applies also
to him. This is eminently true in respect to climate. By
a easuring the rate of reproduction of infusoria, the rate of
growth of plants, the amount of oxygen consumed by crayfish,
or the length of life of the boll weevil we can arrive at an exact
estimate of the effect of climate upon these various organisms.
Inthe same way we can measure man’s response to climate and
find out just what conditions are the best and how much harm
is done by departures from the ideal. Thus far man’s physical
relation to climate has been tested chiefly in the following
ways: (1) by frequent measurements of the weight of healthy
ersons or of those suffering from tubercular or other diseases,
(2) by daily or weekly tests of people’s strength, (3) by
_ €xamination of the amount of work done by specially chosen
subjects or by piece workers in factories day after day
throughout long periods, (4) by measuring the amount of
tarbon dioxide given off in the breath of people who are at
fest under various climatic conditions, (5) by tabulating
| people’s estimates of their own feelings of comfort or dis-
Bemfort at different temperatures and humidities, and (6) by
ertaining the amount of sickness and death at different
158 EVOLUTION OF THE EARTH
seasons and comparing these with the weather. All of these
methods yield substantially the same results. So far as
temperature is concerned they indicate that people’s health is
best and their physical energy greatest when the average
temperature for day and night together is from 60° to 68° F.,
which means when the noon temperature ranges from about
65° to.7E". 5
Among the methods mentioned above the study of deaths
seems to be the best thus far employed. As this method has
never before been used on a large scale, it may be well to con-
sider it somewhat fully. Death is one of the few occurrences -
which takes place in all parts of the world and can easily be
reduced to accurate statistics. Such statistics are kept by all
of the more advanced governments, so an enormous body of
valuable facts is easily available, and needs only to be tabulated -
in order to give most significant results. Accordingly, I shall 4
here give a résumé of the detailed study of about 9,000,000
deaths in Italy, France, and the United States, and of the more
cursory but no less exact study of a much larger number in-
Belgium, Germany, Sweden, Finland, and Japan. The most
important result of these studies is a series of “climographs”
like Figures 31 to 33. Figure 31 is based on 2,500,000 deaths |
in the part of the United States north of the fortieth parallel |
and east of the Missouri River during the years 1900-1912.
Figure 32 is based on 2,200,000 deaths in France from 1901 |
to 1910, and 1,500,000 in Italy from 1899 to 1913. Figure’
33 is based on 142,000 deaths in four California cities, namely, |
San Francisco, Sacramento, Los Angeles, and San Diego, from)
1900 to 1912. The diagrams show to what extent the deaths
during months with any specified temperature and humidity
fell short of or exceeded the normal. Temperature is indi-
cated at the left and humidity at the top. The heaviest shad-)
ing means that on an average the deaths during months having
the conditions included within its area were at least 10 per cent
. P 7 a ce" a er?) ‘ es I sD i 5 ae ~~. . id ee J = ‘
a ee , a of. : e eo ae = ara i | ee ee a --—s
“ . es °r ' - as . pe eee , - 4 — eos ow oo
Ll T
re ee Bee ae ee ee
See . ant
Bolina, on the contrary, the white and the colored deaths in
the towns for which statistics are available each amount to
about 4,000 per year. In spite of this, the two diagrams show
the same general features. This.is highly significant. It
‘shows that in spite of the great outward difference between
y whites and negroes, the two races are fundamentally alike in
their climatic response. Even though his ancestors lived in
Africa for thousands of years, the negro appears to have
acquired little more than an external adaptation toa hot climate.
His case is even more striking than that of the Finn, who has
lived in a cold climate for many generations. The average
temperature in the places wherethese two races have spent most
of their lives differs by about 40°. Nevertheless, the tempera-
ures at which each is most healthy and energetic differ by no
more than 4°. It appears that not only the Nordic, Mediterra-
nean, and Mongoloid races are (almost) alike in their response
to climate, but even the negro joins them. It seems scarcely
going too far to query whether all mankind may not shownearly
the same adjustment to one special kind of climate. Men may
have black skins to protect them from the heat of the tropics,
or fair skins adapted to a cloudy, northern home, but so far as
actual temperature and humidity are concerned they may not be
essentially different. Possibly man’s adjustment to climate is
somewhat like the temperature of the blood, which is almost
unchangeable, no matter in what climate man may live. Pos-
sibly this seeming uniformity in man’s adaptation to climate
may indicate that the conditions under which he is now at
his best are those under which he took the most important
172 EVOLUTION OF THE EARTH
steps in his final physical evolution, and acquired his present
invariable inner temperature.
In this connection it is important to note that among white
men the best conditions for mind and for body do not appear
to be quite the same. Variability seems, if anything, to be
> J aaa
even more stimulating to mental than to physical activity. |
Tests of school children made by Lehman and Pedersen in
Copenhagen and my own study of.the marks of about 1,600
students at West Point and Annapolis indicate that mental
activity is greatest at a temperature decidedly lower than the
optimum for physical energy. For negroes the difference, as _
indicated by daily tests of twenty-two students at Hampton
Institute for sixteen months, is much less. Perhaps this indi-
cates that after the separation of the white and negro races
there came a period of low temperature and of climatic stress
which modified the white man’s mental response, but did not
affect the negro because he had gone too far south.
If the conclusions here presented are sound, a knowledge of
the temperature, humidity, and variability of any region enables
us to determine what effect its climate would have upon the
_physical and mental energy of all mankind. It is thus possible
to prepare a map showing the distribution of either kind of
energy or of the two combined. Such a map, in which physical
and mental energy are regarded as of equal importance, is
shown in Figure 36.5 The map is subject to correction when
fuller data are available, but that will not change its main
outlines. The heavily shaded areas are those of great energy.
The agreement of these areas with the places where civiliza-
tion is highest is too marked to escape notice. In order to ©
bring the matter more clearly before us, Figure 37 has been
prepared. This is a map showing the distribution of civiliza-
tion according to the opinion of fifty competent judges in
fifteen different countries. The judges have been grouped in
5'The map is fully discussed in “Civilization and Climate.”
700 120 140
60
20 40 60
o.
.
.
SS ee
te.
ima
Fic. 36.—The distribution of human energy on the basis of cl
From ‘‘Civilization and Climate.’’
140 160
120
Es
Re
Ne
:
1S > .
e
Y)
ue pany
— ee ee le ee ST
De es aes cw dee ‘acs PEaiveibicie Se. sepns ease akon
te.””
From ‘‘Civilization and Cli
Fic. 37.—The distribution of civilization.
174 EVOLUTION OF THE EARTH
such a way that the opinion of each of five groups, namely ~
Americans, British, Teutonic Europeans, Latin Europeans, and
Asiatics, has equal weight. The agreement of the two maps
is surprising. It indicates that at the present time the distribu- —
tion of climatic energy and of civilization is almost identical.
Such differences as are discernible occur almost wholly where -
exact information is lacking or where the presence of Euro- —
peans as colonial rulers raises the apparent standard of
civilization.
This brings us to the climax of our discussion. At the
beginning of this lecture we saw that human progress, that is, —
the growth of civilization, depends in apparently equal measure ©
upon inherent mental capacity, material resources, and energy.
We then saw that although inherent capacity has no relation
to present climatic conditions, it is closely intertwined with —
those of the past. We also saw that although some resources
like the metals and coal occur without respect to climate, their —
utilization is largely controlled by climatic conditions, while
the vast majority of resources depend directly upon climate
through its effect on vegetation. Finally we have seen that —
human energy, even more than either of the other conditions, —
appears to be dependent directly upon the physiological effect —
of climate upon man’s body. Thus in the evolution of civiliza- —
tion, as in the broader field of the evolution of life, the most ~
obvious controlling factor, although by no means the only one, ©
appears to be climate.
STEPS IN THE EVOLUTION OF CIVILIZATION
Invention of tools and speech, and discovery of use of fire. ©
In the light of this conclusion let us briefly examine some of |
the great steps in the evolution of civilization. Three of the
earliest and greatest steps were the invention of tools, the in- —
vention of speech, and the discovery of the use of fire. As to ©
the relation of tools and speech to climate, nothing definite —
_
ag a ee ee ee ee ee ee
. .
1 a aie sr a ie ae Sa Tid Pie a ete aioe te seg Se eye
i AND ITS INHABITANTS 175
a can be said. They probably became a part of man’s cultural
| inheritance so early that he still lived in his primeval home,
and had perhaps not begun to differentiate into races. If that
is the case, the climate was presumably of the kind which we
_ have found to be most stimulating. That is, it had a tempera-
_ ture about like that of modern Greece, but with much more |
_ variability from day to day. As to fire, the conditions of its
_ discovery were probably like those of the first use of tools
_and of speech. In this case, however, we can go farther.
_ Presumably after man discovered that he could produce fire
artificially he first employed it primarily as a source of heat.
_ Only in a climate which had a distinct cold season would the
incentive to its use be great. Only in a climate where there
_ was plenty of wood and also a dry season to prepare the wood
for fuel would he find it easy to use the new discovery. More-
over, only in a relatively stimulating climate would early men
| probably have had the energy to develop the highly laborious
art of firemaking, for few things require greater determination
_ and persistence. When some happy accident taught man the
' value of heat in cooking food, the discovery doubtless spread
_ to warm regions, but it seems no great stretch of the truth to
_ infer that tropical man, if left to himself, would never have
_ become a user of fire.
| Discovery of use of iron. One of the next great steps in
_ human progress was the discovery of the use of iron. This
discovery required the concurrence of four important con-
ditions, all of which are much less likely to occur within the
_ tropics than without. First, there must be bits of ore lying
_ about where fires are likely to be made. Of course this may
occur in any part of the world, but it is far less likely to happen
_ within the tropics than farther north because the tropical soil
is generally so deep that rocks are rare. In the next place the
fires must be hot enough and of long enough duration to melt
the ore. This, too, may occur within the tropics, but is much
176 EVOLUTION OF THE EARTH .
less common there than in colder climates. In warm regions
cooking is apt to be done over small fires that do not make
people uncomfortable, or else in pits dug in the hot ashes;
and the temperature is rarely high enough to melt iron ore.
Farther north, however, where the winters are chilly, huge —
fires must have been common from the earliest times. There, —
too, in the fairly dry areas many stones lie on the surface
almost everywhere. The next necessity is that when some bits —
of ore happened to be in a particularly hot fire a man of
unusual genius should be present and observe the molten ~
metal. This might occur anywhere, but we have already seen —
reasons for believing that quickness of intellect is fostered in
some climates more than in others. Finally, the fourth requi-
site, and much the most important, is that the man of genius
have the zeal and determination, as Darwin puts it, to bring —
to fruition the ideas engendered by his observations. A singl |
mind, however, was not enough to consummate the great dis-
covery. It was necessary that the generation of men who
lived with the genius should be ‘in a mental mood to receive
the new invention or discovery.” ‘That mood, together with ;
the necessary energy on the part of the genius, is rarely found 4
within the tropics. It is common in regions blessed with a |
stimulating climate. ‘Thus each of the conditions controlling
the discovery of the art of smelting is so much stronger outside
the tropics than within them that it seems highly probable |
that the art arose in a stimulating extra-tropical climate. It —
is generally supposed that the use of iron originated in North ©
Africa perhaps 6,000 years ago. This is eminently consiste - ii
with our conclusions. At that time, as is almost universally —
admitted by geologists, the climate of the world was inter-
mediate between that of the Glacial Period and the present. “4 }
Therefore North Africa was then a decidedly more stimulat-—
ing place than it is today. This same line of reasoning applies
to other great steps in the early development of civilization, |
AND ITS INHABITANTS 177
for many of them were influenced by the glacial stages or minor
( epochs which succeeded the last glacial epoch.
; Taming of wild animals. The taming of wild animals was
_ one of these steps. The dog was apparently the first domestic
animal. His contribution to civilization, however, is slight,
i for instead of lifting man out of the hunting stage, he pre- —
; serves it. If the hunter becomes a tiller of the soil he does not
f want dogs, for he no longer has a frequent surplus of meat
_ which will spoil if the dogs do not eat it. While he remains a
§ hunter, however, the dogs not only help their master to find
_ wild animals, but serve as a reserve supply of food in times of
( scarcity. Among the American Indians no delicacy was for-
merly more esteemed than a fat young puppy. The taming of
| the dog was in itself no great feat. It was easy to bring home a
: wild puppy, which grew up as tame as though its ancestors
_ had long been domesticated. This, however, was the step
_ which apparently led to the domestication of other and more
_ useful animals. It probably did not occur within the tropics,
_ for today the wolf-like creatures from which the dog is sup-
4 posed to be descended do not appear to be found within twenty-
_ five degrees of the equator.
_ Other domestic animals far surpass the dog in utility.
_ Lowest among these stands the pig. This creature is highly
useful as a source of food, but cannot easily be domesticated
unless the art of agriculture is well developed, for it is not
readily herded, and its food is primarily the products of agri-
culture rather than grass. Therefore, though the pig might
have helped civilization, it never had the chance, because be-
fore it was kept in large numbers the tillage of the ground had
_ already done all, and more than all, that the pig could do in
' this line. So far as climate goes it might have been domesti-
_ cated almost anywhere from the equator to the temperate
_ zone.
The horse, the ox, and the sheep stand in a different cate-
7
1
%
178 EVOLUTION OF THE EARTH
gory from the dog and the pig. Aside from agriculture —
nothing did more than their domestication to give man that
sense of ownership and feeling of responsibility which are —
among the most essential prerequisites of high civilization. As —
soon as herds were in his care, man was compelled to watch
them day by day. Carelessness was fatal. He must defend
them from wild animals, drive them to new supplies of grass
and water, and plan to sustain them through the winter. Not ~
only did man thus become a property owner by reason of his —
beasts, but he himself was able to increase in numbers to a
degree utterly impossible while he still depended on the chase.
Therefore the contact of family with family greatly increased.
That necessitated either war and destruction, or some kind of
mutual agreement whereby was laid the basis of a rudimentary
social and political organization. As to the climate under
which domestication of the grass-eating animals took place, itis —
almost certain that the horse, ox, and sheep all were first tamed _
in the subtropical grasslands of central Asia thirty or forty
degrees from the equator. Since this occurred during the ~
change from the last glacial epoch to the present, it presumably |
took place in a climate closely resembling that which we have
found to be ideal. Two other animals, the llama and the |
camel, have been domesticated under less favorable circum- ~
stances, but neither is truly tropical. The llama of Peru lives
where it is decidedly cool. The camel may frequent the hot ~
desert, but the only known wild camels are in the deserts of
central Asia where the winter temperature may be 20° below
zero. Neither of these animals can vie with the horse, ox,
and sheep as contributors to civilization. In the case of the €
llama this may be due to small size, but probably in both cases
a more important factor is the fact that neither animal flour- ~
ishes in the kind of climate in which man is at his best. ft
Rise of agriculture. During the whole course of human ~
history probably no one thing has had a greater influence upon ©
AND ITS INHABITANTS 179
civilization than has the art of agriculture. It is impossible
‘to say exactly where agriculture arose. Perhaps it developed
‘in different ways in many different regions. The great staples
of agriculture are the cereals—wheat, barley, rice, millet,
maize, rye, and oats. The majority of these are of subtropical
origin. Rice, however, which supports more people than any
~ other one kind of food, is probably of tropical origin. The
same may be true of maize. This conclusion, however, is mere
guesswork, for that grain has never been found in its indige-
nous form. Millet, which well-nigh rivals maize in the num-
ber of people it supports, is of great importance within the
‘tropics, but apparently it originated farther north in a sub-
tropical climate, such as that of Egypt. The other grains all
appear to have originated in regions well beyond the limits
of the tropics.
_ Whatever may be said of the origin of agriculture, it needs
‘little demonstration to show that its effect in advancing civiliza-
| tion has been far greater in relatively cool climates than in
' those that are warm. It is a commonplace of history that
_ the great civilizations of early times arose in fertile plains
| where agriculture was carried on by means of irrigation.
_ These were located more than twenty-five degrees from the
" equator, as we see in Egypt, Mesopotamia, northern India,
and China. It is hard to overestimate the effect of irrigation
upon the early progress of civilization. Because of the neces-
» sity of making ditches and reservoirs, and of having everything
"ready when the water is turned on a field, the power to pre-
pare consciously for the distant future is enormously stimu-
lated. The man who depends on irrigation cannot neglect his
fields, for a week of carelessness may cost his whole crop.
He must learn, too, to live at peace with his neighbors. If
men who use the same stream for their water-supply quarrel
with one another and break down one another’s dams and
ditches the chances are that all will suffer famine. They must
;
o
h
180 EVOLUTION OF THE EARTH
agree, and therefore must organize a well-regulated system 0
government. The fact that in irrigated regions many p 20
ple live close together on a small area also tends to caus
society to be thoroughly and wisely organized. Moreove ver
the mere fact that water is precious leads to economy in its ust
and to intensive agriculture. Hence man’s faculty for planning
and for devising new schemes to increase his crops is constantl
stimulated. The Hopis, whom we have already cited, are a
admirable example of this. Could there be any ai
velopment of agricultural science than their way of putting 1g
seed into the midst of a ball of wet clay and burying it in tl
dry sand? The stalk of corn that grows from such a seed m r
be only a foot or two high, but it bears an ear with a hund: rc
grains to replace the one grain that was planted. i
it need hardly be said, is rarely practised in tropical countri
So far as it exists there it is either a relatively late develll
‘ment as in Java, or is confined to the highlands as in Peru, ¢
to certain dry regions as in India. Everywhere it is a ana
response to special types of climate.
So it is with all agriculture. Its type depends upon the kin
of climate. After man has reached a fairly high stage o
culture the most stimulating of all kinds of agriculture appe ar j
to be that which prevails in temperate regions where the a is
rain at all seasons. There each farmer lives on his own ind
pendent farm. He may not have the stimulus of close conté
with his neighbors which is found in irrigated vubceail
regions, but he has the great stimulus of being wholly ind
pendent and of being constantly urged to meet his own needs. —
Moreover, he must be industrious and alert to a degree d
manded of no other kind of farmer. The constant variety
weather to which he is subjected demands that he shall
ready to cut or harvest his hay instantly when the right tin
comes. He cannot put off cutting his grain as can the st
tropical farmer, for when once it is ripe the rain may cor
i" AND ITS INHABITANTS 181
-andruinit. In the fall, no matter how tired he is, he is forced
to gather his crops before a sudden frost comes and spoils
some of them. Thus while tropical agriculture is a help
_ toward civilization, that which prevails in subtropical regions
is far more helpful, while that of the stormy temperate zone is
still more so.
Invention of writing. We come now to the invention of
writing, the great step which marks the boundary line between
_ barbarism and civilization. Twice at least this step has been
taken, once among the Mayas in Guatemala, and once in Asia.
' Perhaps the writing of the Chaldeans may be of different
origin from that of the Chinese, although we cannot speak
_ with certainty. At least the line of development of the western
- forms of writing in Mesopotamia, Egypt, and the adjacent
" lands, has been very different from that on the eastern side of
_ the continent. However this may be, it seems certain that
_ writing was invented in regions with a variable and stimulating
climate, as will soon appear more fully. The invention marks
an era in man’s development as truly as does the discovery of
pene use of fire or the invention of the art of smelting iron.
When man learned to speak, the individual thereby became
able to avail himself of the experience of all the people around
him and of many people whose experiences had been handed
‘down in memory. When he learned the art of writing he no
' longer needed to trust to memory. The minds of all the ages
_ became available to the mind of today. During.the hundreds
_ of thousands of years of the previous existence of the human
ace many a great man must have lived in vain, because among
his immediate associates there was none to comprehend and
carry out his ideas. When writing was invented, it enabled
such men to spread their thoughts a hundred times more
"widely both in place and time. Who knows how greatly our
modern discoveries are influenced by thoughts recorded by the
ancient Greeks and Egyptians?
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182 EVOLUTION OF THE EARTH
Contribution of tropical countries. In these later days sine
historical records have been kept, the contribution of tropica
countries to civilization has been as meager as appears to ha
been the case in the remote past. So far as we are aware, 7
truly great man except Mohammed has arisen within twenty
five degrees of the equator. Gautama, the founder of Buc
dhism, lived north of this limit. Moreover, his home y
among the mountains where the climate is more stimulatir
than in the lowlands. Mesopotamia had its great men whos
names are still known, but that country, though hot in su a
lies from thirty to thirty-five degrees north of the equator. Ss
too with Egypt, for its great men almost without exceptiot
lived from Thebes northward, and the latitude of Thebes i
more than twenty-five degrees north. In modern times it t
even harder than in the past to find great men who lived i
tropical or even in warm countries. Diaz in the high, coo
but nevertheless tropical plateau of Mexico may be cited as a
example, but a hundred years from now only the special stu
dent will have heard of him, while men like Lincoln, Pasteur
and Humboldt will still be admired by thousands, yea, millions r
of people. Even if we go back to Mohammed as an exampl
of a truly great man arising within the tropics, we find :
his ideas came to fruition only when they were carried no ‘
ward to a place where the minds of that generation were mor
alert than in southern Arabia. [
We have now seen that civilization is today highest in place
where climatic energy is also high. We have likewise seen na
in the past no great steps in human progress seem to have beet
taken in tropical countries, nor do great men appear to hay
sprung up there in appreciable numbers. These consideration
apparently lead to the conclusion that when the progress «
human civilization is viewed in the broadest light its grea
movements depend upon climate almost as closely as do the
pulsations of the evolutionary stream in geological times. On
fy
AND ITS INHABITANTS 183
great objection, however, has hitherto seemed to controvert
any such conclusion. ‘Tropical countries may indeed have
_ made no appreciable contribution to civilization at any time,
but no later than two or three thousand years ago many
_ countries where climatic energy is now comparatively low were
the seats of the highest civilization. How was this possible?
; The answer is that a great mass of evidence seems explicable
_ only on the hypothesis of pulsatory changes of climate during
historic times. This evidence has been discussed so fully in
other publications that it seems unnecessary to repeat it here.°
| It will be enough to state the main conclusions with almost no
' details of proof. With these in mind we can apply them to
_ concrete instances and see how climate appears to have been
related to historic crises.
HIsTORICAL CHANGES OF CLIMATE
_ During the past few decades the opinion of geographers as
_ to historical changes of climate has followed a course almost
identical with that of geologists as to earlier and greater
_ changes. Formerly climatic uniformity was supposed to be
_ the “normal” condition and variations were supposed to be
rare and exceptional. Today geologists universally believe
_ that glacial periods have occurred irregularly from the earliest
times to the most recent, and that these have been broken into
_ alternate glacial and interglacial epochs, while other less ex-
_ treme changes have been frequent. Coming to historical times
_ a belief in similar but smaller climatic pulsations is now almost
- universal among American geographers, as is indicated by
® See “The Pulse of Asia,” 1907; “Palestine and Its Transformation,” 1911;
and “The Climatic Factor,” 1914. Briefer treatments embodying further modi-
fications of the original hypothesis, together with replies to certain criticisms, are
found in “The Solar Hypothesis of Climatic Changes,” Bull. Geol. Soc. America,
vol. 25, 1914, pp. 477-590; in “Civilization and Climate,” 1915, ch. XI; and in
“Climatic Change and Agricultural Decline as Elements in the Fall of Rome,”
Quart. Jour. Econ., vol. 31, 1917, pp. 173-208.
184 EVOLUTION OF THE EARTH
the answer to a series of questions recently sent out by the
writer. As Dr. W. D. Matthew, one of the most careful stu-
dents of the matter, says, “‘I should demand proof before I~
would admit that there has been uo change within 2,000
_years.”’
The weakness of the hypothesis of climatic uniformity is
indicated by a consideration of the strongest argument in its
favor. It runs thus: The palm and the vine grow together _
only within a most limited range of temperature. Today they ~
grow together in essentially the same places as 2,000 or 3,000 ©
years ago. Hence no change of temperature and no change
of climate. This argument is weak in two respects. In the
first place, students of the Glacial Period agree that at the
height of the last epoch of advancing ice, perhaps 30,000 years
ago, the mean temperature of the earth was only about 10°, or ~
at most 20° F., lower than now. On that basis an historic —
change of climate one-tenth as great as the enormous change
since the height of glaciation would mean a change of only one
or two degrees in temperature, an amount too small to detect
by means of vegetation. In the second place, the fact that
there has been no appreciable change of temperature in his-
toric times does not mean that there has been no change in the |
distribution of rainfall. The records of the United States
Weather Bureau show that during the years 1875 to 1884 the
region from Galveston to New Orleans had 40 per cent more ~
rainfall than during the ten years from 1890 to 1899. The ~
difference in temperature between these two periods was only
0.42° F. Curiously enough, the wet period was the warmer,
although in other cases the reverse has been true. If such ,
important changes in the distribution of rainfall can occur in
our own day with almost no change in temperature, there is —
no reason why much larger changes may not have occurred ©
similarly in the past. That such changes have occurred is indi-
cated by almost innumerable waterless ruins like those which
ea AND ITS INHABITANTS 185
everywhere dot the country from Mongolia through Turkestan
and Persia to Turkey and North Africa, and also in our own
Southwest. In the deserts, roads too dry for caravans, but
known once to have been much used, also point to a change in
rainfall. So, too, do abandoned springs and the scanty supply
_of water in many aqueducts. One of the most significant pieces
of evidence is the amount of salt dissolved in the waters of |
Owens Lake in southern California, and of Pyramid and Win-
-nemucka lakes in Nevada. This, as Gale has shown, indicates
that from 2,000 to 4,000 years ago these lakes must have been
so high that they overflowed and were fresh. About 2,000 or
2,500 years ago Owens Lake appears to have been two and
one-half times as large as now and to have stood 180 feet
above its present level.
" It is not enough to conclude merely that the climate of the
_ present is different from that which prevailed 2,000 or 3,000
‘Years ago. We must know the nature of the change. Geolo-
_ gists formerly thought that climatic changes proceed very
slowly and uniformly in one direction. They now believe
_ that they are highly irregular. Even when the general change
during thousands of years is decidedly in one direction, it is
_ marked by great irregularities. There is now a general con-
viction that the same is true of historic times. Pulsatory
changes have apparently occurred whereby certain centuries
_have been moister than the present and others drier. No
_ other hypothesis seems adequate to explain the fact that lakes
_whose waters are known to have stood many feet above their
_ present level also contain ruins buried beneath their waters.
Equally strong evidence is afforded by the fact that the growth
; of the big trees of California indicates constant pulsations
whose main features agree with those inferred from other
kinds of evidence in Mediterranean regions and central Asia.
In our own day exact records show an unmistakable harmony
_ between spring rainfall and tree growth in California on the
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Fic. 38.—Changes of climate in the subtropical zone during historic times, on the basis of the
:
13001 1000 800 600 400 SHO RGAD. 20 400 600-800
1606
* AND ITS INHABITANTS 187
one hand, and Mediterranean rainfall on the other. Hence
‘it seems legitimate to infer that the main fluctuations in tree
growth, which are portrayed in Figure 38, represent climatic
_ pulsations of world-wide extent.
Even yet we have not fully explained the nature of climatic
changes. They appear to have been not only pulsatory, but to
have produced different effects in different parts of the world.
It is an interesting coincidence that independently and on
wholly different lines of evidence the German glaciologist,
Penck, and the writer at almost the same time announced the
‘conclusion that climatic changes are the result of an alternate
_ equatorward and poleward shifting of the climatic zones. The
” matter seems to be even more complicated than this, however,
’ for changes in the interior of continents seem to be different
’ from those on their borders or over the oceans. The net
" result is that when such regions as the subtropical zone which
embraces California and the Mediterranean lands enjoy un-
usually abundant rainfall, the northern border of the zone of
equatorial rains becomes drier. The chief basis for this con-
' clusion, so far as historic times are concerned, is the fact that
the Maya ruins in the dense forests of Central America seem
to indicate periods of relative dryness at the times when tree
growth in California indicates unusual moisture, and vice
versa.
_ One more point needs emphasis if we would understand the
“nature of these pulsatory shiftings of the climatic zones.
Apparently the most essential feature is a change in the tracks
of storms. At certain periods the total number of storms
appears to increase. Of even greater importance, however, is
the fact that at such times the storms show a tendency to follow
two belts instead of the one to which they are now largely
confined. The present belt of storms is well known. From
British Columbia it runs as a band hundreds of miles wide
centering along the international boundary line, but with a
188 EVOLUTION OF THE EARTH '
southward tendency. Its center sweeps across the Great Lakes
and then swings northeastward down the St. Lawrence Rive:
to the Atlantic south of Newfoundland. It crosses the Atlant nti
to Europe, where it splits up, and almost dies out in Siberia a
central Asia. On the eastern border of Asia it is joined
an important and vigorous branch from Japan, and then s
the Pacific coast of North America to British Columbus It
course may be judged from Figure 36, where in general th
dark areas of high energy are also stormy areas. The other
or minor storm belt, appears to start in California, cros
Arizona and New Mexico to the Gulf of Mexico, and thet
swing. northeastward into the Atlantic Ocean. In the Ol
World its course lies along the Mediterranean, beginning a ap-
parently east of Spain. Beyond the Mediterranean it passe
across Syria to Mesopotamia and Persia, and then in an attenu
ated form to India. It probably revives again in China an
joins the strongly developed Japanese area of storms. In ch
past, at times when rainfall has been abundant in Mediterra
nean lands, many storms appear to have followed this soutlay “I
belt. Thus increased rainfall and increased variability 0
curred during certain centuries, while the opposite occurr ec
in others. The centuries which saw increased rainfall in hi
subtropical belt saw decreased rainfall farther south in place
like Yucatan, on the northern border of the torrid zone, bu
with this decreased rainfall also went greater variability be
cause of the storms whose centers passed a little farther nortl
Hence it appears that not only in our own day, but throughow
historic times the highest civilization has been found in th
regions of greatest climatic energy. | :
The possible exceptions to this conclusion are the areas
high civilization within twenty-five degrees of the equator.
These include Central America, the highlands of Peru, Yemi
and Rhodesia, Indo-China, and some of the East Indies lik
Sumatra and Java. None of these places, however, made any
f AND ITS INHABITANTS 189
great impression on the world. In the plateaus of Peru,
Yemen, and Rhodesia, civilization took a few steps and then
halted. This corresponds closely with their climate. In mean
‘temperature and humidity they are admirable. Therefore to
that extent they are stimulating. A competent race migrating
to any one of them would be invigorated and would be led to
‘make important advances. There is no evidence, however,
‘that any of them ever enjoyed any great degree of climatic
variability due to strong seasonal changes and frequent storms.
Therefore they lacked the stimulus needed to keep them
ie steadily progressing.
_ The only tropical center of civilization which developed
any really remarkable ideas was Central America, where the
“Mayas evolved a decidedly high type of architecture, a re-
markably accurate calendar based on extensive astronomical
knowledge, and a system of writing more advanced than that
which the Chinese have even yet attained. It is perhaps sig-
' nificant that the center of Maya culture in Yucatan and Guate-
mala lies in the longitude where storm tracks today swing
farthest south and where they apparently swung much farther
: Roach in the past. To this day the Yucatecan descendants of
the Mayas say that they work fastest when the temperature is
lowered by the “‘northers” which follow in the wake of storms
whose centers pass farther north. In the past such northers
Were apparently much more frequent than now. Hence it
looks as if the relatively active Maya civilization were asso-
ciated with variability anal as are the high civilizations of
today."
As to the cultural center in southeastern Asia, we cannot
; Deak with much certainty. It never showed anything like the
Sriginality of the Central American center, for most of its
deas were borrowed from places farther north. It did not
7 This subject is fully discussed in the author’s volume, “The Climatic
| Factor.” Pub. No. 192, Carnegie Institution of Washington.
190 EVOLUTION OF THE EARTH q
last so long as the other. It is worth noting, however, —
relation to the continent of Asia is almost the same as tl
relation of the Maya Civilization to the continent of Americ
There is some reason to think that when the southern sto rrr
belt was intensified even this region, like its American coun’
part, may have had a more variable climate than at preseiell
Effect of climate upon Roman history. The ca
Rome furnishes a good example of the way in which clim
changes appear to influence the march of history.* The gold de
age of Rome occurred 400 or 500 years B. Cc. In the fourt t
century B. C., as appears in Figure 38, storms and rainfal
seem to have been abundant in subtropical lands like Cali
fornia and Italy. In Rome five acres of land was then cor n
sidered enough to support an average family, although tl
presumably does not include the land used for pasturage. Gu
tivation, as we know from numerous classical accounts, .
highly intensive, so that the most advanced methods of
culture were developed. ‘The crops rarely failed and the
was widespread prosperity. “The farmers were iidepedill
and sturdy, and the difference between rich and poor was sligh
The towns were small and the conditions were all highh
favorable to a strong democratic form of government. ’
Turning back to Figure 38 we see that during the thir
century B. Cc. there was a marked diminution of rainfall. I
Rome this was accompanied by two occurrences of siniste
omen. One was a serious decline in agriculture. The smal
tracts of land which had hitherto been the rule were no longe
large enough to furnish a living for the farmer and his famil
Crops that had previously been profitable ceased to be wort
while. Hence there arose much discontent and the agraria
troubles with which the names of the Gracci are closely asso=—
8 See Simkhovitch, V. G., “Rome’s Fall Reconsidered,” Pol. Sci. Quart., J ar
1916; and Huntington, Ellsworth, “Climatic Change and Agricultural Exhav
tion as Elements in the Fall of Rome,” Quart. Jour. Econ., vol. 31, 1917, pp. 17: 3
208. c
| AND ITS INHABITANTS IgI
ciated. The other sinister event was an increase in malaria,
which Jones® has shown to have played an important part in
_ diminishing the vigor of both the Greeks and Romans. Its
increase was apparently due to the fact that the decline in rain-
fall caused a diminution in the amount of vegetation on the
mountains. Hence the streams washed down an undue amount
of silt which filled up their beds in the lowlands and caused
+hem to wander widely and form great swamps. These were
ideal breeding places for the anopheles mosquito which is the
carrier of malaria.
_ After the period of depression in the second century B. C.,
_ Rome recovered somewhat during a period of favorable
_ climate culminating about the time of Christ. Yet she never
was quite restored to her former energy and glory. A century
_or two later, in the early part of the Christian era, there began
a climatic decline which culminated in the seventh century.
' Since crops were no longer profitable, the land was used for
_ grazing purposes, the methods of agriculture became slipshod
and unprofitable, and the farms fell into the hands of a few
large owners. The great number of sheep and goats not only
_added to the difficulties of agriculture by their ravages upon
the fields, but ate up the seedling trees and thus prevented the
_ growth of new forests. Hence the soil was washed away from
the hillsides, with distressing consequences.
_ With this agricultural decline there arose political diff-
culties. Taxes which had previously been easily paid became
‘onerous. Agrariart reforms were even more necessary than in
_ the days of the Gracci, but they were much harder to make.
~ The people flocked to the city in order to get work and thereby
_ share in some of the wealth which came to Rome because of
her conquests. This, however, only increased the evils.
Democracy gave place to plutocracy, and thus to despotism.
At this same time the nomads in the drier parts of the lands
9 Jones, J. H. S., “Malaria, a Neglected Factor in History.”
7
ey
ay
192 EVOLUTION OF THE EARTH 7
surrounding the Roman Empire were also suffering fro 01
drought and famine. Therefore they began to make rai
which set more prosperous people also into motion, and soc
barbarian invasions threatened the very existence of | th
Roman Empire. If the climate had remained favorable then
might have been an occasional raid, but there is no reasaay
think that there would have been the great disasters whic
finally became so terrible.
Worst of all, the climatic changes seem to have had a
effect upon the energy of the Romans themselves. We hay
already seen how close is the connection between climate an
energy. As the variability of the Italian climate decreased
because the storms diminished in. number, the energy of th
people seems also to have declined. This was particularh
unfortunate because special energy was needed to resist the
ravages of malaria, to overcome the agricultural difficulties, tc
engender the self-control and patience which are essential when
political difficulties arise because of undue. taxation or other
causes. Most of all, energy was needed to resist the ovasiel
of barbarians, but instead of this the people’s energy declined.
They became prone to sit still and wait for someone to feed
them, prone to think only of pleasure, and hence ready for
the disaster which finally overtook the Roman Empire.
It would be interesting to carry this discussion farther ang
see how these same principles apply to almost every countr
It is easy to attribute too much to climate, and I am well aware
that many of my hearers will think that I have done so in thi
lecture. It must be remembered, however, that this is meni
a brief attempt to show the importance of one of the great :
factors in the evolution of civilization. The fact that the
importance of climate is here emphasized does not mean that _
I deem it a whit more important than the other factors. 50
far as inherent mental capacity is concerned climate is in one __
sense a minor factor. It is more important as regards material
\] AND ITS INHABITANTS 193
resources, but is far from being the sole factor. Even where
energy is considered, the effect of climate may readily be neu-
tralized by several other factors such as lack of resources or
lack of ability. The point to be emphasized is that climatic
energy is one of the great factors which must be reckoned with
in any attempt to understand the evolution of civilization.
Hitherto, its importance has not been realized. Therefore
today we must emphasize it until it takes its true place with
inherent mental capacity and material resources. It is not a
‘determiner of civilization, but a condition which prevents
| civilization from advancing i in some places and stimulates it to
_ the greatest activity in others. Without an understanding of
‘its part in evolution we cannot rightly comprehend our own
‘present development in any of the great branches of human
_»progress. |
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ctivity, mental, optimum temperature
for, 172
_ physical, optimum temperature for,
172
_Adelochordates, 117
Africans, 148, 149
Agriculture, 153, 154, 177, 178, 180
\ir currents, changes in, 73
Alaska, coal in, 155
Alchemists, 82, 95
Alge, 63, 113
Allantois, 126, 128
Allen, F. J., 101
Allosaurus, 133
a Centauri, 7, 8
Alps, rise of, 74
Alumina, 100
Alveolar substance, of protoplasm, 83
Amblypods, 134, 136
American Indians, 148, 151, 153, 177
Amino acids, 85
Ammonia, 99, 102
Ammonites, 73
Ammonium carbonates, 107
Amnion, 126, 128
origin of, 65, 111
Amphioxus, 117
Animals, domestic, 177, 178
-lime-secreting, 113
wild, taming of, 177
Ankylosaurs, 132
Annelids, 113
pes, anthropoid, 139, 141
origin of, 111
Appalachian Mts., 78
_ revolution, 111, 127
trough, 78
rabians, 150
. _ INDEX
Arboreal habitat, forsaking of, for
terrestrial, 142
Archzocyathine, 113
Archaeopteryx, 129, frontispiece
Archean, beginning of, 43
Archeozoic era, 46, 49, 50, 56, 63
era, time ratio of, 69
sediments, thickness of, 63, 67
Arctic climate, effect of, on man, 149
Aridity, 123, 126, 127, 129, 136
Devonian, 143
Miocene, 143, 146
Mississippian, 145
Permian, 145
Pliocene, 143
Aristotle, 86, 89, 91
Arrhenius, S., 97, 98
Arthropods, 114
Asia, as home of domestic animals, 178
as primitive home of man, 149
as theater of mammalian evolution,
142
Asteroids, see planetoids
Asthenosphere, 41
Athenians, 147
Atmosphere, 54, 57, 75, 80
in Archeozoic, 63
of growing earth, 51
of water-gas, 35
origin of, 50
primordial, 34 ’
Atmospheric blanket, of earth, 54
Atoms, 100, 101, 102
Australians, native, 150
Azoic era, time ratio of, 69
Bacteria, 96, 97
Baker, H., 91
Balanoglossus, 117
196
Barbarism, 152, 154
Barrell, J., 47, 49, 50, 56, 67, 69, 110
Beans, 154
Bees, 90
Beetles, 90
Belgium, optimum temperature in, as
shown by death-rate, 167-169
Bichat, M. F. X., 86
Biochemists, 85, 95
Biogenesis, history of, 89
Biology, 82
Bipedality, cause of, 129, 130
Birds, evolution of, 127
first recorded, 111, 129
origin of, 111, 128, 145 |
toothed, 65, 129, frontispiece
Blainville, D. de, 86
Boleophthalmus, 120
Brachiation, 140
Brachiopods, 113
Breaks, geologic, 70, 71
Brontosaurus, 130
Browne, Sir Thomas, 90
Browsing mammals, origin of, 111
Buffon, Comte de, 92 “
Butterflies, 90
Cacops, 125, 126
California, agriculture of, 153
deaths in, 158, 161, 164, 165
Indians of, 152, 153, 154
rainfall in, 187
rate of growth of big trees in, 185,
186
southern, climate of, 166
storms in, 166
Cambrian period, 46, 79, 111, 113, 118
Camels, 74, 136, 137, 178
Campbell, W. W., 11, 21
Canadian period, 76
Carbohydrates, 84
Carbon, 75, 102
compounds, 100
INDEX
Carbon dioxide, 34, 35, 38, 44, 50,
52, 54, 57, 102, 104, 105 .
dioxide, as climate regulator, 52_
dioxide, in Archeozoic, 63 a.
dioxide, resupply of, throuaig
canoes, 52 i
monoxide, 34, 35 “
Carbonic acid, 99
Cardan, G., 89
Carnivores, 136 B=
Cascadian revolution, 110, 111, 146
Catalysis, 104 ;
Cells, 87, 94, 102, 105, 106
potential immortality of, 98 a
Cenozoic era, 46, 65, 73, 74, 76 Ry
110, 111 Bt
~| ao
oe
era, time ratio of, 69 ix
sediments, thickness of, 65, 67
Central America, 187, 188, 189
Cephalopods, 114, 115
Ceratopsia, 132
Cereals, 179
Cetacea, 115
Chaldeans, 181
Chalks, 58
Chamberlin, T. C., 21, 24, 25, 47, 5 5
116, 119 | 4
Chamberlin and Moulton, on the nebi
lar hypothesis, 12 2
planetesimal hypothesis of, 13,
Chazyian period, 76 ,
Chemical elements, essential to d
105 I
Chimpanzee, 139, 140, 142
China, 179
Chinese, 181
Chlamydosaurus, 130
Chordates, evolution of, cause bs
first recorded, 119
origin of, according to Chambe erli
119 a,
Cincinnati uplift, 62
Cincinnatian period, 76
y!
Dy "
Civilization, distribution of, 173
- evolution of, 147, 174
high, areas of, 188
Civilizations, early, 179
Clarke, F. W., 52, 54, 56
Climate, arctic, effect of, on man, 149
as factor in racial differences, 148
_ changes in, 53, 78, 80, 111
_ changes in, as cause of extinction,
131
_ changes in, causes of, 187
changes in, effects of, 187
changes in, evidences for, 184, 185
changes in, historical, 183, 186
_ Cretaceous, 55, 132
effect of, on clothing, 154
: effect of, on death-rate, 158
effect of, on discovery of use of fire,
= 175
effect of, on energy, 156
effect of, on food, 154
__ effect of, on man, 153
3 effect of, on materials for shelter,
154
. effect of, on Roman history, 190
_ Eocene, 55
_ Liassic, 55
- Permian, 51, 55, 127, 129, 145
Pleistocene, 55, 66, 110
_ Triassic, 55
_ tropical, effect of, on man, 149
Climates, ancient, 51, 52, 55
- glacial, 55, 66, 73, 110, 127, 143
limographs, 159-161, 168, 169
Clothing, dependence of, on climate,
154
effect of, on energy, 156
“necessitated by Glacial period, 143
Yoal, 65, 66, 155, 174
Jelenterates, 113, 114
Sold, increase in, influence of, 127
INDEX
197
Collisions, among planetary nuclei, 33
among stars, 23
Colloidal suspension, life elements in,
106
Colloids, 100, 101
characteristics of, 101
Comanchian period, 46, 76, 79, 111,
131, 145
Communal life, 144
Complexification, in matter, 100
Complexity, law of, 101
Condylarths, 134, 136
Conglomerates, 58
Conifers, 65, 73
Connecticut Valley footprints, 130
Continents, areal variability of, 78
elevation of, 71, 73, 75, 136
in Archeozoic, 63
origin of, 49
permanency of, 50
when largest, 73, 76
Convection currents, 35, 37
Coérdination, biochemical, 106
Corals, 114
Coryphodon, 136
Cosmic time, 46, 47
Cosmozoa theory, 95
Crabs, land, 120
Creator, 108
Creodonts, 135
Cretaceous period, 46, 76, 79, 111, 132,
145
period, climate of, 55, 132
period, in North America, 77
Critical periods, 73, 81
Crocodiles, 131
Croixian period, 76
Crops, of early Americans, 154
Crossopterygii, 122
Crust of earth, granitic, 34
Crustaceans, 120
Crustal density, relation of, to ocean
basins, 39
Crystallization, fractional, 37
198
Crystalloids, 101
Crystals, basic, 37
Culture, centers of, 188, 189.
Cyanogen, 99
Cycads, 65, 73
Cyclostomes, 120
Cynodonts, 128
Cysts, 96, 97
Daly, R. A., 32
Dana, J. D., 50, 69
Darwin, C. R., 108, 155
Death-rate, cause of differences in, 163
effect of storms on, 166
effect of temperature on, 164
Deaths, statistics of, 158, 159, 160, 161
Deer, 136, 137
Denudation, 58
rates of, 59
Deposition, rate of, 68
Deposits, cycles of, 58
glacial, 54, 56
organic, 58
salt, 32
Derham, W., 91
Deserts, 73, 185
effect of, on man, 150
Devonian, Lower, 123, 144
period, 46, 76, 79, 111, 122
period, aridity in, 143
period, diastrophism in, 122
Upper, 145
Diastrophism, 50, 110, 118, 133
Devonian, 122
Miocene, 146
Mississippian, 125
Pennsylvanian, 127, 145
Silurian, 122, 144
Diaz, 182
Dinocerata, 136
Dinosaurs, 73
ankylosaurs, 132
armored, 130, 132
associated with peneplanation, 133
INDEX
Dinosaurs, bipedal, 129
carnivorous, 130, 132
Ceratopsia, 132
duck-billed, 132
effect of, on mammals, 133
evolution of, 145
expansion of, 145
extinction of, 131, 132, 134
nodosaurs, 132
origin of, 111, 129
Sauropoda, 111, 130, 132, 145
Stegosauria, 130
Theropoda, 130
trachodons, 132
unarmored, 132
Diplocynodon, 133
Diplodocus, 130 ‘9
Dipnoans, 121, 122 a
Disruption hypothesis, of Chamberlin
and Moulton, 46 'y
Disturbances, 70 a
Dog, 177 |
Double-breathing, 120
Dromocyon, 135
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nab
Earth, 105
age of, 57
as a planet, 5
atmospheric blanket of, 54
crustal readjustments in, 78
diameter of, 48
effect of moon on, 28
effect of, on moon, 30
evolution of, 45, 94
granitic crust of, 34 . 4
growing, atmosphere of, 51 3
-growth, by rapid infall of plan-
etoids, 25 -_
-growth, by slow accretion of plat -
etesimals, 24 e
-growth, in Formative era, 47
internal heat of, 48, 49
-moon knot, 19
negative areas in surface of, 50
‘Earth, origin of, 46
outer shell of, 40
state, 36
place of, in universe, 4
positive areas in surface of, 50
primordial molten state of, 26, 33
readjustments in, 70, 71
shrinkage of, 48, 70, 71, 80, 109
sub-crustal shell of, 40
_ tidal strains in, 30
_ warping of, 70, 109
Echinoderms, 114
Eels, 90 . .
Efficiency, effect of climate on, 156
Eggs, 126
Egypt,-90, 97, 179, 181, 182
Elephants, see Proboscidea
Energy, as factor in human progress,
147, 155, 173, 174, 192
distribution of, on basis of climate,
173
electric, of chemical elements, 105
heat, 105
to what due, 156
“Energy traffic,” 85, 102
~ England, iron in, 155
- Englishmen, 148
' Environment, changes in, 73
_ fitness of, 88
Enzyme theory, of Troland, 103
Enzymes, autocatalytic, 103
catalytic power of, 104
evolution of, 106
Eocene time, 46, 111, 137, 138, 145, 146
time, climate of, 55
Eopaleozoic time, 79
_Epi-Proterozoic interval, 119
Eras, duration of, 67, 68
Erosion, 32, 40, 54, 56, 57, 58, 60, 70,
76, 78, 110
Erpetoichthys, 122
Eskimos, 151
Europeans, modern, 154
- INDEX
passage of, from molten into rigid ©
199
Evolution, 73, 98
crises of, 112
cycles of, 80, $1
organic, 101
Fats, 84
Finland, optimum temperature in, as
shown by death-rate, 167, 169
Finns, 169, 171
Fire, discovery of use of, 174, 175
life derived from, 99
Fish forms, 116
Fishes, $1, 89, 119, 123
Age of, 46
origin of, 65
Flight, true, evolution of, 128
Floras, origin of, 65
Flying dragons, 73
Food, dependence of, on climate, 154,
155
effect of, on energy, 156
use of fire in cooking, 175
Foot, reptilian, 124
terrestrial, evolution of, 123
terrestrial, origin of, 111
Footprint, earliest known, 123, 124,
144
Footprints, Connecticut Valley, 130
Upper Triassic, 129
Formative era, 46, 47, 49, 50
era, earth-growth in, 47
era, volcanism in, 48
Fossils, 59
living, $1
France, deaths in, 160, 164, 165
Gabbro, 39
Gadow, H., 131
Galaxy, see Milky Way
Gale, H. S., 185
Galton, F., 147
Ganoids, 122
Gautama, 182
200
Geikie, A., 76
Generation, equivocal, 91
spontaneous, 90, 91
Geologic column, 60, 67
history, 108
record, 70
time, eras of, 63
time, length of, 80
time, ratios of, 69
time-table, 45
Geosynclines, 61, 62
Germany, optimum temperature in, as
shown by death-rate, 167
Gibbon, 139, 140, 141, 142, 144
Gills, 120
Glacial deposits, 54, 56
period, 46, 184
period, effect of, on man, 149
periods, 55, 183
Gorilla, 139, 140, 142
Grand Canyon revolution, 111, 118
Granite, 37, 40, 49, 75
Graphite, 63, 113
Grasses, expansion of, 136
used for food by man, 143
Gravitation, 97
Grazing mammals, expansion of, 137
mammals, origin of, 111
mammals, rise of, 136
Greece, 147, 175
Greeks, 89, 191
Habitats, organic, variability in, 78
Hands, as organs of the mind, 143
Haughton, S., 69
Health, best climatic conditions for,
158
effect of storms on, 166
Helmholtz, H. L. F. von, 10, 96, 98
Helmont, J. B. van, 90
Henderson, L. J., 88
Herds, 178
Himalayan uplift, 146
Himalayas, 74
INDEX .
Imagination, scientific, 106
Holmes, A., 52, 57, 60, 61
Homo (Eoanthropus) dawsoni, 142
Hopi Indians, 152, 154, 180
Hornets, 90
Horses, 74, 136, 137, 177, 178
Hottentots, 150
Human progress, factors in, 147
Humboldt, A. von, 182
Hunters, 151, 154, 177
Huxley, T. H., 54, 107
Hydrogen, 102
Hydrosphere, 50, 51, 55, 63, 75
Hylobates lar, 144
Ice age, Permian, 51
age, Proterozoic, 51
Igneous rocks, 43, 75 |
rocks, as source of oceanic salts, 56 —
rocks, erosion of, 32
Impact, effects of, 28
India, 179
irrigation in, 180
Indian corn, 154
Indians, American, 148, 151, 153, 177 :
California, 152, 153, 154
Hopi, 152, 154, 180
New Mexico, 152
Pueblo, 152, 154
Utah, 152
Ute, 152, 153, 154
Indo-China, 188
Infusions, organic, 93
sterilization of, 92
Insectivores, 136
Insects, 73, 89
Intervals, time, 71, 73
Invertebrates, distinguished from ver- —
tebrates, 114 |
evolution of, 114
Iron, 155
compounds of, 102
discovery of use of, 175 ;
limits to use of, 155
INDEX
Irrigation, 153, 154, 179, 180
Irritability of living matter, 95
Isostasy, 40, 41
Italy, deaths in, 158, 160, 164, 165
Japan, optimum temperature in, as
shown by death-rate, 167
Japanese, 149, 169
Java, 188
irrigation in, 180
Jefferson, Thomas, 74
Jelly-fishes, 114
Johnston and Williamson, 52
Jupiter, 5, 6, 31,
Jurassic, Lower, 145
period, 46, 76, 79, 111, 130, 131
Upper, 127, 129, 145
Juvenile waters, 38, 75
Kankakee uplift, 62
Kant, hypothesis of, concerning evolu-
tion of solar system, 10
Kelvin, 96
Kircher, 90
Labrador, fishermen of, 152
Lamprey, 116
Land-bridges, 73, 74, 138
Land waters, as place of chordate
origin, 119
Lands, contributions of, to oceans, 61
elevation of, 70, 109, 110, 111
Lane’s law, 15
Laplace, P. S. de, 10, 50
Laramide revolution, 111, 132, 134, 145
La Tour, C. de, 93
Laurentian peneplain, 118
Lava, 2, 38, 44, 75
Lavas, of moon, 43
Proterozoic, 64 |
Law of Complexity, 101
Lehman and Pedersen, 172
Leith and Mead, 60
Lepidosiren, 122
20I
Leverrier, U. J. J., 27
Lewes, 86
Lias, climate of, 55
Life, 88
ancient, 59
Archeozoic, 63
as manifestation of protoplasmic.
activity, 82
changes in, 73
chemical elements essential to, 105
chemical elements of, grouping of,
105
communal, 144
cycles in, 73
definitions of, 85
dormant, 97
elements, coérdination of, 105
elements, grouping of, 105
elements, in colloidal suspension, 106
evolution of, 64
fresh-water record of, 59
land record of, 59
marine, increase in, 78
marine, record of, 59
materials at basis of, 51
origin of, 63, 82, 94, 112
paleontological record of, 93
pulse of, 81, 109, 111, 144, 146
scientific explanation of, 95
thermometers, 55
transportation of, through space, 96
vehicle of manifestations of, 85
Light, radiation pressure of, 97
Lightning, 102, 103
Limbs, change in, in primates, 143
Lime-secretion, establishment of, 113
Limestones, 58, 60, 61, 67, 68
Lincoln, A., 182
Lithosphere, 41, 49, 50, 75, 76
density differences in, 41
Lizards, bipedal, 130
frilled, 130
Llama, 178
Locusts, 90
202
Loeb, J., 98
Loligo, 115
Los Angeles, deaths in, 158, 161
rainfall at, 153
Lucretius, 89
Lull, R. S., 148
Lung-breathing, first recorded, 123
place of origin of, 121
Lung-fishes, 121, 122
first recorded, 144
Lungs, development of, 144
of higher vertebrates, 120
origin of, 111
Maggots, 90
Magmas, 2, 34, 37, 38, 39, 40, 42, 43, 44
Maize, 179
Malaria, 191
Mammals, 65, 73, 81
Age of, 46
archaic, 111, 134, 135, 136, 145
browsing, origin of, 111
evolution of, 127, 145
expansion of, 134
fate of, 136
first recorded, 111, 127
grazing, rise of, 111, 136, 137
modernized, 135, 145
modernized, origin of, 111
modernized, source of, 135
origin of, 111, 128, 145
placental, 126
rise of, 133
Man, Age of, 45
best climatic conditions for, 158
brute-, 74
dominance of, over organic world,
144, 146
increase of, dependent on domestic
animals, 178
origin of, 111, 137
physiological adjustment of, to cli-
mate, 156
reasoning, 74
INDEX
Miacida, 136
Man, response of, to climate, 157, 74
rise of, 146
Mars, 5, 27, 31
Material resources, as factor in hun
progress, 147, 151, 152, 154, 18 5,
174, 193
resources, limited by climate, 155
Matter, complexification in, 100
evolution of, 106
living, irritability of, 95
Matthew, W. D., 117, 184
Mayas, 181, 187, 189
Mediterranean lands, iron in, 155
lands, rainfall in, 187, 188
peoples, 169,171. a
Membranes, extra-embryonal, 126, 12! .
Mental activity, optimum temperaturs
for, 172 a
alertness, a manifestation of ener zy
156 .
alertness, effect of lack of materia 1
resources on, 150 om
capacity, as factor in human prog-
ress, 147, 148, 174, 192
Mentality, 81
Mercury, 28
Mesopotamia, 179, 181, 182 .
Mesozoic era, 46, 65, 73, 76, 78, 79, 111, a
132, 133 1
era, time ratio of, 69
sediments, thickness of, 65, 67
Meteorites, 33
Meteors, 27, 97
aoe:
A
ag
”
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Fi ota i
SN cota
Se es
Mice, 90
Microcosm, organism as, 88
Milky Way, 8
Millet, 179
Mining, 153
Miocene time, 46, 111, 136, 138
time, aridity in, 143, 146
time, diastrophism in, 142, 146 Tae
Mississippian period, 46, 76, 79, a“
125, 145
‘
INDEX
Mississippian period, aridity in, 145
period, diastrophism in, 125
Mohammed, 151, 182
Mohawkian period, 76
Molecules, 84, 100
_ Molluscs, 114, 120
_ Molten stage, in earth’s history, 26, 33
- Mongolia, 185
Mongoloid races, 171
Monotremes, 127
- Moon, 2, 5, 13, 28, 30, 33, 43
effect of earth on, 30
effect of, on earth, 28
in Archeozoic, 63
- Moon-knot, 47
Moore, B., 100
- Moulton, F. R., 12, 18, 19
_ Mountain-making, 53, 70, 78, 109
Mountains, 71
cycles of, 60
origin of, 48
Muds, 58, 60
a Mud-skippers, 120
Mudstones, 58, 61, 67, 68
Mutants, 148
_ Nature, uniformity of, 45
Nebulz, 8
irregular, 8
planetary, 8
ring, 9
solar, 21, 47
spiral, 8, 9, 13, 18, 20, 21, 47
stellar, 9
~ Nebular hypothesis of Laplace, 10
hypothesis of Laplace, modifications
of, 11
_ Needham, T., 92
Negative areas, in earth’s surface, 50
Negroes, American, adaptation of, to
_ climate, 171
death-rate of, 164, 168, 169, 171
effect of tropical climate on, 149
Neoceratodus, 121, 122
203
Neogene time, 79
Neopaleozoic time, 79
Neptune, 4, 5, 6
Neutral medial area of North America,
62
New Mexico, Indians of, 152
New York City, death-rate in, 171
Newfoundland, fishermen of, 152
Nitrogen, 44, 102, 103
compounds, 104, 105
in primordial atmosphere, 34
Nodosaurs, 132
Nome land-bridge, 74
Nordics, 149, 169, 171
North America, in Paleozoic time, map
of, 62
North Carolina, death-rate in, 171
Nuclei, planetary, 19, 33
Nythosaurus larvatus, 128
Oceanic basins, 43
basins, in Archeozoic, 63
basins, increase in volume of, 39
basins, origin of, 39, 49
basins, relations of crustal density
to, 39
currents, 54
currents, changes in, 73
floodings, 61, 71, 75, 76, 79
floodings, Cretaceous, 77
floodings, Pennsylvanian, 66
floodings, Silurian, 72
level, inconstancy of, 76
salts, 31, 50, 56
waters, gathering of, 37
waters, increase in volume of, 50
waters, origin of, 55
Oceans, carbon dioxide in, 52
increase in area of, 78
origin of, 37
when smallest, 73
Oligocene time, 46, 111, 135, 136, 145
Orang, 139, 140, 142
Orbits, planetary, 5, 19
204
Ordovician period, 46, 79, 111, 113,
119, 144
Organic deposits, 58
habitats, areal variability in, 78
Organism, 105
Organisms, bionomic relations of, 108
individuality of, 87
origin of, 106
Orion, 8
Osborn, H. F., 104
Ostracoderms, 119
origin of, 111
Over-specialization, indications of, 131
Owens Lake, 185
Ox, 177, 178
Oxalates, 107
Oxides, 100
Oxygen, 44, 51, 102
importance of, for life, 92
in primordial atmosphere, 34
Ozark uplift, 62
Paleogene time, 79
Paleogeography, 66, 72, 77
Paleozoic era, 46, 64, 65, 76, 78, 79,
111, 127
era, North America in, 62
era, time ratio of, 69
sediments, thickness of, 65, 67
Palms, 184
Panama land-bridge, 74
Pasteur, L., 93, 182
Patten, W., 119
Peabody Museum of Yale University,
123, 133
Penck, A., 187
Peneplanation, 76
Pennsylvanian period, 46, 66, 76, 79,
111,..127
period, diastrophism in, 127, 145
period, North America in, 66
Periodicity, 70, 75
Periophthalmus, 120
INDEX | 3
=!
“~—
Permian period, 46, 76, 79, 111,127 _
period, aridity in, 145 Z
period, climate of, 51, 55, 127,
145 a ,
Persia, 185
Peru, 180, 188, 189
Petromyzon, 116 ~
Pfliiger, E., 99 ae
Phasic activity, 100 a
Phenacodus primavus, 136
Phosphates, 107
Physical activity, optimum ‘temy
ture for, 172 —
Pig, 177 3
Planetary nuclei, 19, 33 a
system, 47 Sa
Planetesimal dust, 25 . ,
hypothesis, 13, 101
hypothesis, outstanding difficulties @
20
material, 32
Planetesimals, 35, 47
earth-growth by slow accretion o:
24 z
Planetoids, 4, 5, 26, 28
earth-growth by rapid infall of, 26
significance of, 26
Planets, 4, 5, 13, 18, 19, 24, 28, 31
relations of, 5, 6
Plants, flowering, 65, 73
herbaceous, 136
lime-secreting, 113
Pleiades, 8
Pleistocene time, 46, 111, 138
time, climate of, 55, 66, 110 x
Pliocene time, 46, 111 4
time, aridity in, 143
Polypterus delhezi, 122
Positive areas, in earth’s surface, 50 —
areas, in North America, 62 =
Posture, erect, assumption of, 143
Primates, 136, 137, 143
American, 137
descent of, from trees, 143, 146
INDEX
Primates, distribution of, 138, 139
Eocene, 137
European, 138
evolution of, 146
origin of, 111
radiation of, 137, 138, 142
Proavian, 129, frontispiece
Proboscidea, 74, 136
Proteins, 84, 85, 99, 100
Proterozoic era, 46, 113, 118, 144
era, climates of, 51, 55
era, time ratio of, 69
lavas, 64
sediments, thickness of, 64, 67
Protochordates, 118
Protoplasm, 82, 88, 99, 100, 105
alveolar substance of, 83
chemical characteristics of, 84
combinations of elements in, 84
lability of, 85
living, evolution of, 107
physical characteristics of, $3
variation in, 87
Protopterus, 121
Protozoa, 96, 97, 113
Psychozoic era, 45, 66
Pueblo Indians, 152, 154
Pulse of life, 81, 109, 111, 144, 146
Pumpkins, 154
Pyramid Lake, 185
Rabl, C., 123
Racial differences, causes of, 148
Radioactive elements, 42
minerals, rate of disintegration’ of,
as basis for computing geologic
time, 67, 68.
Radiolaria, 113
Rain, 54
first, on earth, 37
Rainfall, 153, 154, 184
Ranodon sibericus, 124
Recrystallization, 40
Redi, F., 90, 91, 92
205
Reptiles, 65, 73, 81, 132, 145
Age of, 46
foot of, 124
origin of, 65, 111, 125
Revolutions, 71
Rhinoceroses, 136
Rhodesia, 188, 189
Rhythms, in solar energy, 109
Rice, 179
Richter, 96, 98
Rivers, 54, 57, 76
Rocks, acidic, 49
basaltic, 49
basic, 49
granitic, 49
sedimentary, 57, 61, 67, 68
stratified, composition of, 60
Rodents, 136
Roman history, effect of climate on,
190
Ross, 90
Sacramento, deaths in, 158, 161
Salamander, foot of, 124
Salt deposits, 32
evidence of, for climatic change, 185
oceanic, derivation of, 32
oceanic, significance of, 31
Salt Lake City, rainfall at, 153
San Diego, deaths in, 158, 161 *
San Francisco, deaths in, 158, 161
Sandstones, 58, 60, 61, 67, 68
Santa Fé, rainfall at, 153
Satellites, 6, 19
Saturn, 5, 6, 31
Sauropoda, 111, 130, 132, 145
characteristics of, 131
Savagery, 152, 154
Schafer, E. A., 98
Schuchert, C., 110
Schwann, 93
Scorpions, 90
Sea-anemones, 114
206
Sedimentary rocks, distribution of, 61
rocks, origin of, 57
rocks, thicknesses of, 61, 67, 68
Seeds, 97
Selection, natural, 106
Sharks, 120°
Sheep, 177, 178
Shellfish, 90
Shelter, dependence of, on climate, 154
effect of, on energy, 156
Silica, 100
Silurian period, 46, 76, 79, 111, 122, 123
period, climate of, 55
period, diastrophism in, 122, 144
period, North America in, 72
Sloths, fossil, 74
Snails, 90
Snow, 54
Socrates, 151
Soil, 155
Solar energy, 102
energy, rhythms in, 109
prominences, 16
system, elements of, 5
system, Kant’s hypothesis of evolu-
tion of, 10
system, Laplace’s theory of origin of,
10
system, plantesimal hypothesis of
origin of, 13
Sollas, W. J., 60
Space, interstellar, transportation of
life through, 96
Spallanzani, L., 92
Speech, invention of, 174
Spencer, H., 86
Sponges, 113, 114
Spontaneous generation, 90, 91
Spores, 96, 97, 98
Squids, 73, 114, 115
Star streams, 7
Stars, 6, 13, 15, 23
chances of close approach among, 22
INDEX y
‘"
Stars, mode of tidal disruption in, 15 x
new, 9
Wolf-Rayet, 9 4
Static forms, $1
Stegocephalia, 125, 145
Stegosauria, 130, 132
characteristics of, 131
Stegosaurus, 130
Stellar system, sun a member of, 6 —
Stockholm, optimum temperature a
as shown by death-rate, 167, a |
Storms, changes in track of, 187.
effect of, on human energy, 166
present belt of, 187 B -
Strand-line, 76, 120 )
Stratification, density, 36 , a
Struggle for existence, 73, 81 ag
Suess, E., 38 or ,,
Sulphur, 102
Sumatra, 188 4
Sun, 4, 10, 12, 13, 19, 20, 44, 98, 1¢ 5
ancestral, tidal disruption of, 16 5
as life factor, 54 es
as member of stellar system, 6 i.
critical velocity of, 17 om
origin of, 46 a
prominences shot out from, 16
rotation of, 20
Sunshine, 102
Surface processes, in earth, 43
Swan, 89 .
Sweden, iron in, 155 ne
Swim-bladder, 120 a
Swine, 136 .
Tapirs, 136
Tartrates, 107 a
Temperature, changes in, effect of,
165, 172 4
effect of, on death-rate, 163, 164, 165.
optimum, 167
Terrestrial foot, evolution of, 123
foot, origin of, 111
types, influence of aridity on, 127
'
ea em
i aan ae 1a
a 4 :
——-
De x
Re
INDEX
Terrestrial types, influence of increas-
ing cold on, 127
vertebrates, eggs of, 126
vertebrates, emergence of, 119, 122
waters, as place of chordate origin,
119
waters, as place of origin of lung-
breathers, 121
Tertiary, climatic oscillation in, 135
Theriodonts, 128
Thermal springs, 56, 75
Theropoda, 130
Thinopus antiquus, 123, 124
Thorium, 42
Tidal disruption, forces of, 13, 14
disruption, in ancestral sun, 16, 17
disruption, mode of, in stars, 15
force, of earth on moon, 30
retardations, 28
strains, in earth, 30
Tide-generating force, 16
Tides, body, 29
in Archeozoic, 63
loss of energy due to, 29
Tools, invention of, 174
Trachodons, 132
Trees, growth of, as evidence of cli-
matic change, 185, 186
Triassic period, 46, 76, 79, 111
period, climate of, 55
Upper, 127, 129, 133, 145
Triton, 124,125
Troland, L. T., enzyme theory of, 103
Tropical climates, effect of, on man,
149
Tropics, contribution of, to civiliza-
tion, 182
Tunicates, 117
Turkestan, 185
Turkey, 185
Tyndall, J., 93
Underhill, F. P., 84
Ungulates, 136
207
Uniformitarian trend, of biologic
thought, 106
United States, eastern, deaths in, 158,
159, 161, 164, 165, 168, 169, 170,
171
northeastern, storms in, 166
southwestern, ruins in, 185
Weather Bureau, 184
Uranium, 42, 67, 69
Uranus, 4, 5, 6, 31
Utah, 152, 153, 154
Indians of, 152
Ute Indians, 152, 153, 154
Vadose water, 75
Vegetation, dependence of, on climate,
155
Venus, 5, 28
Vertebrates, distinguished from in-
vertebrates, 114
egg of, change in, 126
evolution of, 113
first recorded, 144
origin of, 111, 114, 144
terrestrial, emergence of, 119, 122
Virgil, 89
Vitalism, 94
Volcanic activity, as source of earth’s
water, 56
activity, times of, 74
ashes, 75
Volcanism, in Formative era, 48
Volcanoes, as source of carbon dioxide,
52
Walcott, C. D., 69
Warm blood, origin of, 111, 127, 145
Wasps, 90
Water, precipitation of, 100
Water-vapor, 54, 98
-vapor, as climate regulator, 52
-yapor, in primordial atmosphere,
34, 35, 38
Waters, juvenile, 38, 75
; Waters, anit? as place of chordate
origin, 119
oceanic, gathering of, 37
- vadose, 75
Waucobian period, 76
Weathering, 32, 33, 80
Whales, 81
Williams, H. S., 69
Wilson, E. B., 94
Ar
Wd oa el Pe ee
Winnemucka Lake, 185 :
| Worms, 89, 90, 113, rr
ng ees
4 ‘a
Writing, invention of, 11
Yemen, 138, 189 me
Young, ClAns, |
Yucatan, rainfall i in, 188
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