UC-NRLF 1. I I lur EVOLUTION OF MAN ByWILHELA BOLSCHE LIBRA! ^^"^siBMtf^ THE LIBRARY OF THE UNIVERSITY OF CALIFORNIA GIFT OF Junius Adams THE EVOLUTION OF MAN BY WILHELM .BOLSCHE Translated by ERNEST UNTERMANN SIXTH THOUSAND CHICAGO CHARLES H. KERR & COMPANY 1906 Copyright, 1905 BY CHARLES H. KERK & COMPANY .Ijjgggj^ 241 THE HENNEBERRY CO., PRS., CHICAGO. 0 C PREFACE Whoever claims to be an educated man, a man who thinks, must acquire a knowledge of the outline of modern scientific research and of the theories concerning the descent of man. No thought is so essential and sublime as that about ourselves. One may be skeptical as to the value of these things, but before any discussion of them is possible, one must, above all, think. There must be no class distinction in view of these questions. Wherever great philosophies and movements in their interest have appeared in history, they have not addressed themselves merely to the kings of the spirit, but instinctively to the simple man of the people, to that place where the heart of the people is beating. Since natural science to-day claims to offer a new basis for a scientific world philosophy, it must again address itself to the common people. It may seem that scientific methods of expression and thought are an obstacle to popularization. If so, we must take so much the more pains to over- 457 PREFACE come this obstacle and find a popular interpreta- tion for our thoughts. The present little volume is addressed to the widest circle of readers, even to those who are as yet unacquainted with a goodly number of excellent but much more volu- minous works concerning the same subject. This little work is reduced to such a size that it may easily be perused in one leisure hour. Neverthe- less I think that the facts which it presents will furnish material for independent reflection in serious hours. As for its scientific basis, I have only to men- tion the name of Darwin. Whoever thinks him- self beyond this name in our days is specially in- vited to analyze his theories once more by the help of this short and comprehensive sketch. In its more intricate details my presentation of the matter is naturally based on certain ideas of Ernst Haeckel, but I must also give due credit to the great influence which the more recent re- searches of Herman Klaatsch of Heidelberg have exerted upon me. Whenever I have ventured beyond the line of facts, or combination of facts, I have done so from my own firm conviction that a thinking man is not dragged down by all these relations with the animal world, but is PREFACE rather strengthened and furthered in the con- sciousness of his own ethical powers. He then appears to me so much more triumphant above his animal nature, standing victoriously above the dark foundation of his own existence. Man and his history reach back into the primitive world of animal monsters, but this animal nature, this primitive world, lies prostrate at his feet over- come by himself. WlLHELM BOELSCHE. Friedrichshagen, New Year's Day, 1904. THE EVOLUTION OF MAN A lovely picture extends before my eyes. A virgin meadow stretches down a valley clad in emerald green. Innumerable blossoms of dande- lions and blue-bells rise from it like golden and violet flames. A gray granite wall, a witness of primordial days, forms the background to this fresh wave of full life. Above it, like a dark blue stage setting, rise the fir forest and the op- posite mountain wall. And far. far beyond it, al- most merging into the soft blue sky with a slightly deeper tint, appears the outline of the giant mountains. Now, a snow-white cloud, glistening in the sunlight, floats slowly and phan- tom-like towards me, coming down from the unknown distance beyond, and disappearing above me in the glittering light. The bright glow of the sun is diffused throughout it all, lending charm to the flowery meadow, the granite and the THE EVOLUTION OF MAN mountain forest — a great unity sunk in harmo- nious tranquility. Now, I hear far-off voices. Human beings are passing by, shielded from my sight by the great stone blocks. They are strang- ers, I do not know them. How much may be hidden by such distant voices — good and bad! What an infinite variety is comprised in this little word "man," how much that is noble and sublime — and how much that is brutal ! And yet, while those feeble undulations of the air which carry those voices toward me are still trembling in my ear, I am thinking of the simple message of the gospel, according to which all men without distinction are my brothers. Our civilization has at last risen to the point of impressing us with the fact that this many-headed mass of fif- teen hundred million people on the surface of this globe are bound by one common tie of sacred- ness which is expressed in the word, man ! They are all one unit, these human beings, one great family assembled on the surface of this globe, ready to share their sins, to forgive one another, to enjoy their pleasures together, to go hand in hand on their way through this great valley of riddles, the universe. But a clearer and sharper sound, not articu- lated into words, mingles with those indistinct 10 THE EVOLUTION OF MAN voices. It is the fine voice of a little baby, this monotonous and clear wail which sounds so help- less and yet stirs so much compassion! We all have grown up, we all have developed from such a small baby, such a bud of humanity. And my glance wanders once more over the green meadow. All those golden blossoms of dandelions and all the blue-bells have developed from a bud. Every one of those plants has grown up into the sunlight from some simple germ. And it seems to me that it is this same sun which neither of them can dispense with. The little rosebud of humanity in its cradle needs the sun quite as much as that brown, rough bud of yonder meadow flower. If the sun above us which is floating in the ice-cold solitude of space nmetfas ix millions miles a.wav were to be extinguished to-day, humanity would perish just as surely as the kingdom of meadow flowers. And from the depths of the human soul, whence also the lessons of the gospels have come, still another voice whispers into my inner ear. It is that same voice which was first heard in the wisdom of ancient India, and it said that the tie of common interest, of brotherhood, is not con- fined to man and man, that it comprises all living things of this globe, all things which grow up 11 THE EVOLUTION OF MAN under' tne rays of the sun in the silent grasp of secret, natural laws, and gradually develop to the summit of humanity. It is that other simple mes- sage which tells us : "Thou shalt not torture any animal uselessly; thou shalt hot wantonly break any flower, for they too are distant relations in the great flow of life, they too are still your brothers in the unfathomable recesses of nature. Helpless stands that flower, or that glittering lit- tle beetle before you, just like a trembling, little child. But the child grows up into a man, and who knows what this flower or that beetle may become some day, or what may have become of others like them, millions of years ago! It is such sentiments as these which every one of us feels in his or her best moments which seem to me fitting for the discussion of such a tremendous question as that of the evolution of man. Wherever the compassion of man can find its way, there the blessed and divine longing for understanding may also wend its steps without fear or shame. Whoever has so much love that he can feel it for an animal may also approach with perfectly clear conscience that other ques- tion, whether the blood relationship that freely binds him to other human beings does not per- 12 THE EVOLUTION OF MAN haps extend still further, whether he himself may not have developed from an animal. And he may recognize with calm conviction that this fact cannot have any more significance morally than that other fact which is affirmed a thousand times every day and sanctified by the deep love of every mother — the fact that even the greatest man must have first developed from a primitive, human bud, from a child which can neither speak nor walk, which germinates in the dark recesses of nature, just as that blue bell out there develops under the hot rays of the sun. And if the individual develops in this way, why should not all humanity have developed in this way, once upon a time? •*•* It was about a million years ago. If a man could have had the opportunity to wander through our present European continent, with a rifle in his hands, he would have seen in those days a very strange country. He might have imagined that he was in the interior of Africa as we know it to-day. He would have tramped for( weeks over immense prairies in Southern Europe, dotted sporadically with a few dense woods, and out of the wilderness of this green ' ocean of grass, he would have started before him innumerable herds of antelopes, giraffes and THE EVOLUTION OF MAN animals resembling wild horses. From his camp near a rippling spring, he could have watched in the clear moonlight, such colossal forms com- ing to drink and to bathe as were once seen by the first hunters who^ ventured into the interior of Africa by way of Cape Colony. There, he could have seen elephants of various species, with two and four tusks, or even with tusks bent downward like those of the walrus, massive rhinoceros, and ponderous hippopotami. Behind them he could have heard the roaring of lions, panthers, and giant wild-cats armed with saber- like teeth. Wandering further north into locali- ties which are now the scenes of a highly ad- vanced civilization, he would have entered the most impenetrable, primeval forest, similar to that in which Stanley in the heart of Africa experienced all the sensations of daring conquest of an absolutely wild tropical country. Out of the dense undergrowth, splendid palms rose to- ward the sunlight. Parrots of many colors shrieked, the features of a large anthropoid' ape, similar to our gorilla, might peep suddenly out of the thick covering of foliage, piercing the dar- ing intruder with sharp glances. And above it all, there trembled the atmosphere of a hot cli- mate. 14 #t-^£*-*-*. THE EVOLUTION OK MAN Our wanderer would have been still more surprised if he could have compared our present- day maps with the road traveled by him in those primeval days. Where the blue surface of the Mediterranean now extends so widely that a navigator cannot see the shores on either side, he would have advanced over dry ground from horizon to horizon through prairies inhabited by giraffes and forests peopled by monkeys. And where to-day the red rose of the Alps grows upon dizzy heights near the grim ice of the glaciers on mountain passes, there he would have found nothing but wooded hills in which his geologically trained eye might have discovered traces of a slow but irresistible rise. And where to-day the sun is sending its glowing rays down upon bare mountain ranges, as in the heart of France, he could have observed the horizon tinted blood-red, a reflection of the boiling lava of vol- canoes. A strange world in an immeasurably far off time! A million years is a tremendous period of time for human minds to grasp. If the history of human civilization is traced by written chronicles, it does not take us back beyond six thousand years. One might fill entire libraries with events 15 THE EVOLUTION OF MAN through which human beings have passed merely in a period of one thousand years. Here, we are supposed to place side by side thousands of thousands of years. What wonder then if the mirror of research transports us back to those primeval times into a different Europe, composed of different seas, countries, mountains and cli- mates. It is the so called "Tertiary Period" into which we have looked. Four great periods are distinguished by the historians of the earth, in speaking of the change and succession of animal and plant life as it is discovered in the course of the many million years during which it has developed. We may use the simple Latin numbers to designate these periods: Primus, the First, Secundus, the Sec- ond, Tertius, the Third, Quartus, the Fourth. There is the Primary period, the very first in which we discover traces of living beings on our earth. It was then that the forests were green, the fossil remains of which we now know as coal. Strange and uncouth newts/crawled about in their shade. The sea, the shores of which were covered by these trees, was alive with long forgotten crustaceans^and fishes. Then followed the Secondary period, in which the terrible giant THE EVOLUTION OF MAN saunans, typified by Ichthyosaurus, infested land and sea. After that we reach the third great period, the Tertiary period, when Europe had the climate and the fauna of present day Africa, such as giraffes, elephants and monkeys. And when this epoch came to an end, the Quaternary period began, with which our entire historical tradition is identified and in which we are stll living to-day. We do not meet any familiar objects until we reach this last period. The sur- face of the earth then assumes the form to which we are now accustomed. All things come closer to us. The things that lie beyond are strange to us, like an unknown creation, like a dream of some other planet. And yet man lived even in that Tertiary period. No song, no heroic story, gives any informa- tion about him. But where the voice of tradi- tion, the chronicles of conscious humanity are silent, there we find other witnesses that speak to us — the stones. The tradition of mankind ex- pires within the Quaternary period. There is an extreme moment when even the most ancient inscriptions of the Chinese, the Babylonians, and the Egyptians become mute. Written characters disappear and with them the earliest direct voice from the cradle of humanity about itself. But THE EVOLUTION OF MAN beyond that point we are made aware of a very important event in the development of this earth which took place in this Quaternary period, the traces of which are still visibly impressed in the rocks. It is the great ice age. For many thou- sand years, colossal masses of glacial ice were piled on top of the continents of Europe and North America. Large herds of mammoth, a species of elephant, covered with a thick coating of hair as a protection against the cold, grazed along the edge of these glaciers, just as in our day the musk-ox and the reindeer are doing in the countries near the North Pole. Undeniable and plain traces of human beings are still pre- served from that period. In the sand, which remained when the glaciers flowed into the caves which were formed by the mighty ice waters boring their way through the lime rocks, the crude and simple stone tools have been found with which the men of that period hunted the mammoth. The walls of such caves in France are still covered with colored pictures in which the men of that ice age have drawn unmistakable pictures of the mammoth. As it happens we are enabled to test the accuracy of those pictures, since well preserved bodies of mammoth with skin and hair are found in the 18 THE EVOLUTION OF MAN ice of Siberia. We have also found the skulls and bones of those men, so that we now have a fairly good idea of their characteristics, in spite of the fact that all written and oral traditions of the civilized nations now living have com- pletely forgotten their ancestors of the ice age. Even the most sublime symbolical picture of the evolution of civilization, the Bible, does not men- tion them anywhere. But those simple stone tools, especially knives and arrowheads, which give us such reliable information of man as the contemporary of the mammoth, are occasionally found also in the strata of rock which were already present when the ice age with its glaciers and mammoths be- gan. We find in them remains of that most prim- itive human civilization, together with bones of a giant elephant, who was not only larger and of different form than the mammoth, but also older — the so-called South-elephant (Elephas meridionalis.) But this South-elephant was still living in laurel groves and under magnolia blos- soms in France and Germany, instead of feeding on reindeer lichens on the edges of the glaciers. With this elephant we have come into the middle of the genuine Tertiary period. This Tertiary period, the more we follow it backwards, takes 19 THE EVOLUTION OF MAN us into a warmer climate instead of a colder one. In the middle of this period we meet with that very picture which I drew in the beginning. Eu- rope then had the giraffe plains and the primeval forests of the present day inhabited by anthro- poid apes, and there is no longer any doubt that the oldest tools of man, which we can distinguish as such, lead us even to the limit of this very hot, middle period of the Tertiary age. Man is even then a part of that picture ! He is himself almost a million years old on the surface of this globe, and had simple stone weapons and other tools which he used in his fight with the giant animals of that time. In other words, he possessed the indubitable beginnings of civ- ilization. It seems to me that we cannot trace matters up to this point without confronting this further question: Is it not possible that man may be still older? With this venerable age of one million years he is a part of the wonders of the primitive world, he drifts into the company of still stranger animals than the mammoth, into other climates than those of present-day Europe, the Alps of which were then in the first stage of formation and the seas of which had not yet found their 30 THE EVOLUTION OF MAN present level. So it really would not change matters very much even if we found that we must trace him further back into still more an- cient and strange landscapes of this globe. It is true that all traces of civilization disappear at this point. We do not know of a single piece of flint stone in the first half of the Tertiary period, or even of the saurian period following it, which would show the traces of the human hand. But long before we reach this point, we may observe a gradual divergence of these flint stone tools. They grow cruder and cruder. Is it too wild a speculation to suppose that men may have existed even beyond that time who may not have possessed sufficient civilization even to fashion the simplest stone tools ? In that case, we could not expect to find any stone tools as witnesses. But, one might say, there should at least be genuine human bones preserved in a fossil state in the solid rocks together with skeletons of the ichthyosaurians ? Still, this objection would not carry much weight. We know very well that not all of the living beings which once lived upon this earth left their fossil bones behind. The bones may have been destroyed, for human bones particularly are not very durable. Or they may ,./, THE EVOLUTION OF MAN be buried in certain places of the earth which we cannot investigate to-day, because they may be at the bottom of the sea, or covered by the perennial ice of polar regions. How often has not this earth been shaken through and through and turned inside out in these long, long periods ? Strata, which were once sediment at the bottom of the sea and which are still full of sea shells, are now found on the high summits of the Alps. On the other hand, entire mountain ranges, gr©und into sand, are now found in the flat sandstone of the plains, or at the bottom of the sea. Many of the remains of the primitive world have certainly been destroyed in this wild chaos, have been ground into powder, or broken to pieces. We get a vague conception of, this when we see that even the gigantic monsters of those primitive days have frequently left but one single bone, a thigh bone or skull of one single indi- vidual. That is to say, while thousands and thousands of individuals of this species lived once upon a time, only the scant remains of one single individual have come down to our time. Then too, there is still another possibility which is far more interesting. It is very probable that we may not recognize the man of those far dis- tant days, even if some of his bones were pre- 22 THE EVOLUTION OF MAN served. For man himself might have become transformed in his structure, and his bones might differ from ours. Might it not be possible that his bones might look so strange to us that scientists might have described them as belong- ing to some other being, little aware of the fact that these remains represented just the thing for which they were looking? Similar ideas have ever played a role in various tales and legends. There, we read that the men of the primitive world were gnomes, or again giants, Cyclopes with one eye, or fauns with goat's feet, tails and pointed ears. When mam- moth bones were first found, it was said that they were the actual remains of such old fabulous men, bones of the giants Gog and Magog, or of St. Christopher. Of course, this was nonsense, and the supposed human bones were nothing but honest mammoth bones with no relation to primitive man. But, we of to-day have really something better than mere remains to rely on, we have reliable scientific data for the theory that men with essentially different characteris- tics from ours existed not so very long ago. I mentioned, a while ago, that we have remains of skeletons of men who lived in the ice age, the age of mammoths. But these men of the 23 THE EVOLUTION OF MAN Two remains of skulls of palaeo-diluvian Men, with strongly protruding eye-bumps. Both cuts are side views. The cut at the top represents the skull-cap (the only part which was pre- served) found in 185C in a cave of the Neander valley near Dxisseldorf, Germany. The cut at the bottom represents a fairly complete skull, found in 1887 in a cave of Spy, near Namur, France, together with equally aged bones of Mammoths, Rhinoceros, and Cave-Bears. 24 THE EVOLUTION OF MAN ice age, who are still relatively close to us when compared to the more distant primitive periods, are not so very much behind in their civilization when compared to certain savage peoples of to- day. Even in our day, there are certain tribes, for instance in South America, who are not familiar with metals, who fashion all their tools and weapons out of stone, horn, or wood, and who therefore are actually living in the "Stone Age" similar to those primitive mammoth hunt- ers. Nevertheless, if one of us had met one of these primtive ice age men, we should have been somewhat startled by the features of that man. For his face, his size and his limbs would have appeared to us perceptibly different from ours, even from those of the savages of the present day. True, no one would have doubted that this was still a "man," but something strange, some- thing divergent, would certainly have startled us in this type of the "Ice-a^e man." We may still reconstruct this man tolerably well from the re- mains of his skeleton. It was in 1856 that such genuine human bones, with strangely divergent characteristics, were dis- covered for the first time and scientifically ana- lyzed. It was in the so-called Neander Valley near Dusseldorf (Rhineland). Some working 25 THE EVOLUTION OF MAN men were clearing out an old cave. They found an old and partly decayed skeleton. A phy- sician, Dr. Fuhlrott, happened along and saved as many of these bones as he could obtain. In this way they reached a museum, and they are now on exhibition in the Provincial Museum of Bonn. [The student is especially surprised by the con^ Istruction of the skull of this man, which is very flat in the part directly above the brain, and has thick and unsightly bumps right over the cavities of the eyes. Even the lowest Australian has no such bumps on his forehead to-day. For a long time the genuineness of this dis- covery was doubted, and no correct conclusions could be formed because the experts could not agree on the period to which this Neander Val- ley skull should be assigned. Some even doubted whether this man was really very old and whether he could have been a contemporary of the mam- moth. Rudolph Virchow then took part in the discussion and claimed that whatever might be the antiquity of these bones, and granting that they might be genuine bones of a contemporary of the mammoth, they certainly did not belong to a normal man, but rather to one who was dis- eased. The divergence from the present human type was attributed to the effects of disease. It REMAINS OF PITHECANTHROPUS ERECTUS, the mysterious being found by Eugene Dubois on the island of Java. The cut shows a skull-cap, seen from the side and from the top, with its bumps above the eyes; furthermore several views of the left thigh-bone, and two molar teeth. The thigh- bone has on its inner side some abnormal formations due, proba- bly, to some wound which this specimen received while alive. 27 THE EVOLUTION OF MAN was supposed that this Neander Valley man suf- fered from softening of the bones when a baby, from gout when an old man, and that at some time in his life his skull had been crushed by a blow and healed imperfectly. And in this way the bumps over the eyes and the other strange characteristics were supposed to have been pro- duced. But this very daring assumption, which looked far-fetched when examined in detail, was refuted when Professor Fraipont, in 1887, dis- covered two human skeletons in another cave near Namur (France), the so-called cave of Spy. These skeletons had skulls with the same strange bumps on them. One could not easily assume that all these individuals had endured the same improbable sufferings. Some time after that, a whole mass of remains of such bones, belonging to not less than ten individuals of different ages, were found near Krapina in Austria. They evi- dently represented the remains of a prehistoric cannibal feast, and the poor victims who had been roasted on that occasion had all of them the same structure of skull as that of the Neander Valley man. And, finally Schwalbe and Klaatsch have demonstrated scientifically that the Neander Valley bones were not at all diseased. It is quite certain, then, that a type of man 28 THE EVOLUTION OF MAN with such skulls has once existed, and the dis- coveries at Spy and Krapina have shown at the same time to what period that man belonged. They were found together with the bones of the mammoth and cave bear of exactly the same age. They were therefore remains of "Ice-age" men, and these ice-age men still showed this strange divergence from the present living type of "man." Now, let us imagine that these variations con- tinued far into the more primitive period. The traces of civilization, as we have seen, finally disappear altogether. Man himself, if present in those very primitive periods, would not have been advanced far enough to fashion the crudest weapons out of flint stone. And we may log- ically draw conclusions from this lack of ability as to his physical constitution. The man of the Ice-age was able to fashion weapons from flint- stones, and yet he was far behind us in the struc- ture of his skull. How far behind, then, in the structure of his skull, would be a man without knowledge of flint stone tools? The line of research here absolutely dissolves into nothing. Man diverges more and more from the present type of human beings. He finally varies to an extent which makes him absolutely 29 THE EVOLUTION OF MAN indistinguishable and hides "Man" in beings which are not at all like him. We must recall to mind the millions of years of the primitive world, the infinite succession of time, and think on and on along this line of natural development, just as we would in the case of a star which, once started on a defintie course from a certain point, continues to move and move incessantly in a certain fixed direc- tion. But now that we have gone so far, we feel a pardonable curiosity and a certain daring. Would it not be possible for our penetration, once we have conceived of these possibilities, to forge ahead still farther into the mystery of things, get at the facts of all these "possibilities," and ask what disguise man might have adopted ? What may be those strange primitive beings, the fossil remains of which we might perhaps find and in which he may be most likely hidden ? We have at least a starting point. We per- ceive, so to say, the mathematical point where the course begins to deviate, that is to say, we may start from these grotesque skulls of the ice- age with their crude bumps above the eyes. May we not speculate a little further as to the next physical transformation, and so forth? 30 THE EVOLUTION OF MAN It is precisely at this point that we meet with something which has the great advantage of not being merely a logical assumption, but rather a tangible scientific fact. The beautiful island of Java in the tropics has long been known on account of its violent vol- canic eruptions. As late as the Tertiary period there was an eruption of a certain volcano which buried an entire section of land with loose masses of ashes in the same way in which Mt. Vesuvius buried the city of Pompeii in historical times. On this occasion a multitude of living beings were buried. Their bones remained in that volcanic mass and were later on carried to a certain place by waters washing their way through this mass. The name of this place to-day is Trinil, and the old mass of volcanic ashes is now a part of the bed of the Bengavan river. In 1891 a Dutch physician, Eugen Dubois, made excavations in the banks of this river, and incidentally he dis- covered masses of old bones, mostly the bones of large mammals of the Tertiary period, such as elephants and hippopotami which do not live in Java in our day. But among those bones Dubois found also a thigh bone and skull cap and a pair of molar teeth of a peculiar creature which had evidently lived in those primitive days with 31 THE EVOLUTION OF MAN those animals at the time wherv the eruption of that volcano occurred. This creature must have had a strange likeness to human beings. It had almost the height of a man. Its upper thigh indicates that it had the habit of walking upright. Indeed, it was so manlike that a number of authorities in anatomy, for instance Rudolph Virchow, declared without hesitation that it was a genuine human bone. But matters were different with the skull. Flat, without a forehead, and with bumps above his eyes, this skull seemed in its fundamental plan to be an extreme exaggeration of the Neander Val- ley skull. But this exaggeration went so far that the human likeness receded against a new likeness. This Trinil skull resembled strikingly — a monkey skull. And it was -even possible to name the definite species of monkey which it resembled most nearly, a monkey living to this day in Southern Asia, the so-called gibbon. The gibbon is the nearest relative of the ourang- outang, the gorilla and the chimpanzee. The present living species are all of them much smal- ler than this strange creature of Trinil was. But that old skull was, in many respects, so like that of the gibbon that quite a number of grave ex- perts declared that it belonged to an extinct species of gibbon which had the size of a man. 32 THE EVOLUTION OF MAN Still, a few others did not agree with this idea. The cavity of the skull, so far as it was preserved, was filled with gypsum in order to find out how much space it contained for a brain. The figure ascertained by this means was ap- proximately half-way between a gorilla and the lowest Australian aborigine. That is to say, its brain capacity exceeded by far that of a gibbon without however coming anywhere near that of present-day man or even the ice-age man. What sort of a creature could this be? The scientists disagreed. "A very gibbon-like man," said some of them. "A very man-like gibbon," said the others. The discoverer Dubois took a middle course ; he baptized this creature with the double name of Pithej^njftror^^ This disagreement of the scientists is very in- structive in our research. We learn, as an actual fact, that in the Tertiary period there still existed on this globe certain creatures which stood about half-way between a man and a gibbon. Their skull exaggerated those characteristics, by which the ice-age man was distinguished from present- day man, to such an extent that this creature ap- proached a new station which we have long known by the name of. monkeys. In this way we are given a definite goal indicating the first 38 THE EVOLUTION OF MAN disguise in which we may look for man further back and discover him, so to say, by evidences which reveal his presence beyond that limit where he began to deviate entirely from the present type of. man. Is it perhaps possible that at a certain histor- ical stage man simply merges in the monkey? Here another very old and venerable line of reasoning, long used even in the most exact re- search of nature, comes to our aid. It was in 1735 that Linnaeus, a great^scientist, performed a monumental work. He then gave us the first comprehensive system of nature's forms. He arranged these forms in three great kingdoms, minerals, plants, animals. And within these kingdoms he arranged the various forms in systematic succession. In this way, he fur- nished us with a system of plants, and of ani- mals, which, in spite of its defects, gave us the first foundation for a comparative view and log- ical sequence by which we could hope to discover the natural connections of these forms in their main outlines. In performing this necessary work of genius, Linnaeus naturally had to solve the question: Where am I to place man? He did not hesitate for one moment. He placed man in the animal 34 THE EVOLUTION OF MAN kingdom on account of his physical structure, which showed that he belonged to the mammals/ and more definitely in the group of monkeys. Indeed, if we wish to build up any system even in our day, that is the only logical conclusion at which we can arrive. Man is not a simple mineral, he is a living being. Unless he is fed, he dies; that is to say, his form of existence is that of living beings who are compelled on pain of death to assimilate food. If we pinch his arm, he cries out ; in other words, he feels, and he has that peculiar faculty which we are accustomed to associate with the word "life," the faculty of suj)jjejrtiyjr feeling. Furthermore his food is of a definite kind. He cannot feed on pure mineral substances, he requires either vegetable or animal matter, he needs bread instead of stones, and of the elements of the air he can utilize only oxygen. This classes him with the other members of the animal kingdom in distinction from plants which feed on the soil. Again, in the animal kingdom there are two main groups. It is true that Linnaeus himself was not familiar with this distinction, but we have learned it since then. The individual body of the animal in one of these groups consists of only one so-called cell. It is one solitary little THE EVOLUTION OF MAN lump of animated substance. The individual body in the other gjroup of animals is composed of many such cells, which form a sort of co- operative association with division of labor. Well then, the body of man is built up with billions of such cells in the most wonderful manner. It consists of living building material, the cells, which make up its muscles, its blood, its skin and even its bones. In other words, man belongs to the group of animals that contain many cells, (lie does not belong to the uni-cellular low arch- Ityjges/ he is not a microscopically small in- ifusorium.ft This higher group of animals is again di- vided into a number of groups, among which we must make our choice. There are the sponges, the polypi, the jelly-fish, the worms, the star- fish, the echinoderms, the crustaceans, the in- sects, the snails, the shells, and, finally, ajjrpup which is distinguished by a spinal cord located above the digestive tract and protected by a more or less solid structure which serves at the same time for the support of the body, a backbone. We call this last group the vertebrates. No other group has this characteristic structure, and it is plain, at the first glance, that man can belong- only to this group, because he has a spinal cord THE EVOLUTION OF MAN AN OLD ORANG-OUTANG. His face is disfigured by peculiar excrescences on both cheeks. 37 THE EVOLUTION OF MAN and a backbone. Within this group of verte-X ibrates we distinguish the fish, which breathe ] in the water with gills instead of lungs; man i breathes through lungs, therefore he is not | a fish. Then follow the amphibians, that is to I say, the newts and frogs that breathe alternately • through gills and lungs. A frog, for instance, i breathes through gills, when a tadpole, and ac- quires his lungs later on. Human beings do t not have this double method of breathing. Fur- I thermore, the reptiles, that is to say, lizards, i crocodiles, turtles and related animals have blood which changes its temperature from warm to cold and vice versa. Their blood is cold when ' the air which they breathe is cold, but it is warm j when the sun shines upon them. These animals , do not yet possess their own heating apparatus within them. The human body heats itself, it is ! always warm, hence man is not a reptile. The I two last groups of vertebrates are always warm. These groups consist of birds and mammals. • Since we have to choose between these two, we . must investigate further. No bird suckles its ' young, but the human mother does that, and all i mammals do, therefore we belong on the side ' of the mammals. Now these mammals are again I divided into two great sections. Those of one 38 THE EVOLUTION OF MAN 1 section lay eggs, the Australian duckbills. The ! mammals of the other section have done away with that; the child when born is far more ma- ture. Every human mother testifies to the fact !< that human beings are not duckbills, but belong to a higher class. And now we come to a final choice. We look at the hands and teeth of man. Man is not a whale, the hands of which have turned into fins. He is not a carnivorous animal which has one-sidedly developed its eye- teeth and incisors. He is not an animal with hoofs which has laid special stress upon its molar I teeth. He is not a rodent, the best trumps of which are the incisors ; he is not a sloth, the teeth j of which have entirely degenerated, nor is he a bat, the hands of which are made into wings. There is only one single group of mammals, the teeth and hands of which resemble those of man, Kand that group is composed of monkeys. X Mark well: when Linnaeus placed man side by side with the monkeys in his system, he was not thinking of anything else but just an orderly arrangement, a systematic grouping of animals at a greater or smaller distance, just as a boy will stick his beetles into his collection, some closer, others farther apart. But since the days of Linnaeus a good many deep thinkers and clear 39 THE EVOLUTION OF MAN heads have asked the question whether this "sys- tem" might not have a deeper meaning and re- lation to nature? Now, when we remember that ye arrived at a certain station in our research which we named monkey-man, the probability of a deeper mean- ing of that system grows apace. We were look- ing for some primitive disguise by which man might have concealed his identity far back in the days of the primitive world, and we must cer- tainly say, when we think of this system, that of all the creatures of this globe, none is better fitted for such a disguise than is the monkey, that is to say, that animal which in spite of all the differences of its bony structure is still far more like us than all the other living beings of the earth together. Remember also that we were not speaking of monkeys in a general way; but indicated a certain species, the gibbon. Systematic zoology very early accomplished the separation of some species of monkeys from others, the so-called anthropoid apes. This word indicates that these apes are still closer to man in the system than any others. No other group in the system is so close to us. We now distinguish four species of these anth^grpoid apes. Two of them are living in Africa, tjjie gorilla and the i chimpanzee, and 40 THE EVOLUTION OF MAN two in Asia, the orang-outangand the gibbon, These four apes strangely resemble human be- ings, even externally. The layman is specially astonished to notice that they, like man, have not jin externally visible JaU. But scientists know that this occurs occasionally even among lower monkeys and so it is not considered a very convincing mark. But there is a very wonderful relation which should convince the most inveter- ate skeptic, and that is the following. Whoever has looked at a drop of blood through a very strong microscope knows that this; jpeculiar fluid is a mixture of two things, first, the so-called serum, and then the blood corpuscles floating round in it. Now when we compare the drops of blood of various animal species, we find that the red blood corpuscles have many different forms. Some of them are long, some are round, some are large and some small ; in brief they are different in fish, or newt, or bird, or mammal. This is no ground for sur- prise, for all these animals are very different in many other ways. The peculiar significance of this difference is that the attempt to inoculate an animal of one group with the living blood of another group always ends fatally. It is just as if these two kinds of blood carried on a war with one an- 41 THE EVOLUTION OF MAN other. The serum of one group destroys the blood corpuscles of another group. If an animal is inoculated with the blood of another group, it quickly feels the fatal effects of this struggle in its veins. It falls into convulsions and finally collapses entirely, just as a conflagration con- sumes a city in whose streets a violent civil war is raging. And this happens often in the case of animals which are relatively close to one another, for instance, many mammals. The blood of a cat kills a rabbit which is inoculated with it, and vice versa. But finally there is a cer- tain limit. The blood of a cat naturally does not kill another cat. Indeed, peace is guaranteed often among more distant relatives. Closely re- lated animals may mix their blood without dan- ger. A dog is so close to a wolf that the living blood of the one may mix with that of the other without harm. It is the same with a horse and a donkey. Now a short time ago a certain scien- tist, Friedenthal in Berlin, mixed human blood and monkey blood. At first one blood acted as a poison for the other; that is to say, as long as the objects of the experiment were man and a lower monkey. But when human blood came o the blood of the chimpanzee, peace was sud- denly established. The boundary of antagonisms had been crossed. The blood of man and that of 42 THE EVOLUTION OF MAN the anthropoid ape were so nearly akin that they agreed without difficultly. How could this be? Here it was not a question of comparing bone with bone. An answer came directly from the living. The secret of life, the most minute chemistry of the blood, testified to the most intir mate relationship, a consanguinity in the most Xdaring sense of the word. -* With this fact we have made another step ahead. The probability grows that man may have been concealed once upon a time in one of these creatures which we see represented by the anthropoid apes of to-day. Indeed ,the experi- ment with blood makes it almost evident that all four anthropoid apes now living are directly connected with this mysterious primeval fact. The question is only, what is this relation? We first of all feel tempted to ask whether these anthropoid apes themselves might not rep- resent that primitive stage for which we are looking. Are not these apes veritable primitive men that have not yet been transformed into genuine men to this day? One thinks involuntarily of the ludicrous tales of the negroes who say that the gorilla and the chimpanzee are really men, only they are too lazy to work, and for this reason pretend that 43 THE EVOLUTION OF MAN they are monkeys. And perhaps there is suffi- cient truth in this theory to justify the belief that these apes actually represent a type of primi- tive man who was arrested in his development against his will, and who went so far in his conservatism that he still illustrates the "mon- key stage" of man. Again one might ask at this point, how it is that a few of our crude and monkey-like great- grandfathers are still living in the form of iso- lated men of the woods, as a fixed primitive type, at a time when present-day genuine man has long arrived at his perfect form. But we meet with the same phenomenon within genuine humanity itself. Why does the native Australian with his Stone Age civilization still live in the bush, while over here civilized man has already risen to the full height of his evolution? And we have an illustration still closer at hand. In the plains where the modern metropolis steams and roars, progress walks with seven-leagued boots, while yonder in the remote mountain vil- lage ancient customs and institutions are still in full bloom. So, this would not be a very perti- nent objection. However, let us take a closer look at the anth- ronoui «H^es. We have four species." These four 44 THE EVOLUTION OF MAN species differ considerably from one another, some of them show even extreme differences. Do they possibly represent four different primi- tive stages of man? But every attempt to re- construct them from a continuously ascending line towards man is a complete failure. It is true that each species has a number of its own peculiar resemblances to man. But, it rather seems that these resemblances are distributed among them in a rather indiscriminate way, so that they all supplement one another in a funda- mental outline of man, but nevertheless do not form an ascending chain of evidence. We now remember that strange creature of Trinil, and our attention turns especially to the gibbon. Is it possible that he could be a genuine archetype, and that the orang-outang, the chimpanzee and the gorilla could be merely un- progressive branches? One thing cannot be de- nied: this gibbon possesses indeed very strange and portentous characteristics. It seems that this ape actually brings us closer to the secret of our descent. He is not a bestial gorilla, but a much more gentle and soulful creature. ^Te can. singthe^ music of the gc^le. — a very strange case in a mammal, which involuntarily reminds us that it is precisely in man that language and song 45 THE EVOLUTION OF MAN A GIBBON. The anthropoid apes of the gibbon group are living in southern Asia and in the Sunda Islands. They have extremely long arms, which are much longer than their legs. The gibbon has very significant relations with the human line of descent. THE EVOLUTION OF MAN have developed. Furthermore, if the gibbon descends from a tree to the ground which, by the way, he does not like to do, he walks habitu- ally on two legs and balances himself at the same time by stretching out his arms sideways, or folding them above his head, and these arms of the present-day gibbon are again a new clue in our research. Compared to the trunk and the legs these arms are excessively long. Any com- parison with man seems impossible in view of these arms. No other mammal has arms of such length. However, if we study the habits of gib- bon life, we easily recognize their purpose. The gibbon is the cleverest climber among the anthro- poid apes. He is an unexcelled acrobat, thanks to these arms. They represent an extreme but very adequate adaptation to his special needs. But when it comes to comparing him with mod- ern man, these arms of the gibbon certainly point away from us. The question arises whether the primitive man for whom we are looking could ever have had such spiderlike arms. The gorilla, chimpanzee and ourang-outang also have pretty long arms, but they are not nearly so long, and in that respect these apes seem to be much closer to man. Even a majority of the lower apes, such as Macacus, and even the ba- 47 THE EVOLUTION OF MAN boons, have a much closer resemblance to man in this one point. There seems to be only one way out of these strange contradictions. We must conclude that the living anthropoid apes are closely related to the archetype of man for which we are looking, but they do not represent its thorough-bred type. f Each one of them has developed along his own I line from this thorough-bred type simultaneously ^with man as we know him to-day. They did not change very much, but still they went far enough to acquire each his own peculiarities. All of them retain strong resemblances to the archetype, but one has preserved more of some characteris- tics and lost others, while the reverse is true of another species. Very likely the gibbon still re- sembles that archetype most closely, but even he has later acquired those enormous arms. It is highly interesting to know that we may mention a direct reason for our general assump- tion of probability, so that it becomes almost a certainty. Among living beings there is a very curious law, or at least a near approach to one. Young animals very frequently resemble the ancestors of their whole race more nearly than the adult animals. A frog in the tadpole stage still resembles a fish which breathes in the water 48 THE EVOLUTION OF MAN through gills. A great number of higher animals assume again in the egg, or in the mother's womb, certain forms which we meet on a much lower and more ancient plane. A bird in the egg shows for a while a great mass of vertebrae, in its tail which once characterized the extinct bird- lizard ( Archaopteryx) , a transition form be- tween lizard and bird, existing millions of years ago. * Haeckel has called this peculiar fact, which x recurs in innumerable cases and truly indicates a general and lawful connection, the "biogenetic principle," and this term has become fairly popu-^ lar to-day. x Well, then, the very first observers noticed that the gorilla, the chimpanzee, the orang- outang, resemble man more in proportion as they are younger. The giant gorilla, which is the most ferocious and bestial of all anthropoid apes in old age, resembles in its baby stage the human being so closely that even the layman who has never thought about these things is surprised. In view of the biogenetic law, this would indicate that these anthropoid apes are descended from an ancestor who was still more manlike than they are to-day. And the point is finally clinched by some facts which the scientist, Emil Selenka, has recently discovered in regard 49 THE EVOLUTION OF MAN to the gibbon. An unborn gibbon in its mother's womb at first has well proportioned arms just as if it were to become a human being. And it is only by gradual stages that the arms of the little ape develop into those enormous acrobatic limbs. If the biogenetic law is correct, then we would have in this case an exact proof that the ancestors of the present-day gibbon did not pos- sess those long arms and were, therefore, con- siderably more manlike. EMBRYOS OF A GIBBON, in an advanced state of development. Mark the likeness to a human embryo. (After Selenka.) Thousands of indications -thus point to the fact which occurred even to Darwin when he dis- cussed these things tentatively for the first time, so THE EVOLUTION OF MAN some thirty years ago. XA species of mammal X has once existed on this globe which contained the germs, not alone of man, but also of the gorilla, the chimpanzee, the orang-outang and the gibbon. All of them have later developed /from that type — unlike sons^of the same father. X No doubt this creature was, in some respects, a closer copy of the present anthropoid apes than of modern man, and it must have been closest to the gibbon of to-day. However, it was dis- tinguished from this gibbon, as we know him in his adult form by certain more manlike marks. And if we were to call that primitive being "man," because genuine man is descended from him and because he has such strong resemblances to human beings, then we might say of the pres- ent-day anthropoid apes that they are descended from man, instead of man being descended 'from the orang-outang, or the gorilla, as some lay- men frequently claim. That would be a much more correct statement, and would conform to the idea of Darwin, who gave rise to these dis- cussions. That primitive type is no longer living on this globe. Unless an unexpected discovery is made . in the partly unexplored forest regions of the in- terior of Africa, we may close the books in this 51 THE EVOLUTION OF MAN matter. At this point then, our steps must be directed exclusively towards the primitive world. But, what can be said in regard to those primi- tive bones and the possibility of fitting them into the picture which we have just drawn? Here we remember once more that famous Pithecanthropus of Trinil, who is half gibbon, half man. Is it possible that he could be the very type for which we are looking? There is one thing which gives rise to doubts, and that is the time to which he belongs. We have seen that it is almost a certainty that genuine man lived in the second third of the Tertiary period, that is to say, in those tropical forests of middle Europe. Recently, flintstone tools have been found in France in the strata of that period, which the scientist called the "Miocene Period." These tools are almost identical with certain stone tools of the crudest kind which every expert attributes to human hands. But the great forests of this Miocene period were inhabited by man-like apes. In Austria, Switzerland and France, there lived a genuine gibbon (Pliopithecus) and another species lived in France, closely resembling the chimpanzee, but yet standing by itself without being any closer to man (Dryopithecus) . A little later we also find genuine chimpanzees and 53 THE EVOLUTION OF MAN orang-outangs. So much we can tell by well preserved bones. It is evident that the unlike sons of that mysterious archetype had already branched of! at that period, and the types had become so plain that they could be separated into anthropoid apes and men. It seems, however, that the bones of Pithe- canthropus, which we know belonged to the ex- treme end of the Tertiary period, are apparently many thousand years younger than those bones of the Miocene period. If that creature of Trinil still contained in the germ a common thorough- bred type, then it follows that this type must have lived simultaneously with its unlike sons on the island of Java, even after the lapse of so many thousand years. Of course, such a thing would not be impossi- ble. Only we might ask whether that thorough- bred type could have been preserved in its orig- inal form during this entire period. We might be inclined to suspect at least some of the least typical characteristics and assume that this type might have developed a little further and adapted itself to the new conditions, while nevertheless it might still give us a far better idea of the ac- tual course of development than the present anthropoid apes. 53 THE EVOLUTION OF MAN It is also logical to ask whether Pithecan- thropus was not a long surviving "last Mohican" of a transition form from a genuine thorough- bred type to the genuine man. It all depends upon the weight which we lay upon the specific- ally genuine human marks. If any one is more attracted by the resemblance of that form to the present gibbon, he might argue that Pithecan- thropus was a transition form from the arche- type of past genuine man to the genuine gibbon. This last theory might be seriously considered from the moment that we could get a glimpse of the arms of that archetype, which we do not know as yet, provided they were to show a tendency toward the grotesque elongation of the genuine gibbon arm. Let us hope that the exca- vations in Java will be diligently pursued and that we may then be able to solve some of these more intricate problems. So much at least is certain, that the genuine common ancestor in question, who must have had at least a very close resemblance to Pithe- canthropus in the structure of his skull and legs, existed before the Miocene period, that is to say, in the first third of the Tertiary period. He rep*3 (resented the "Man" of that time — a creature! I which contained the possibilities of developmentj 54 THE EVOLUTION OF MAN -^ into genuine man and also those of development 1 into a gibbon, chimpanzee, gorilla and orang-j outang. Doubtless the greater part of his body was covered with strong hair, such as the present anthropoid apes have inherited from him. He is a real, genuine, living "Esau." The fact that the smooth "Jacob," man of to-day, has only a very slight indication of this hairy covering on most parts of his body, is not a proof to the con- trary. For we find the instructive law on the resemblances of the youthful forms to their an- cestors gives us a very satisfactory clue to our original ancestor: the body of the human being in the mother's womb is also, in its first stages, covered with thick woolly hair. Even the face is covered just as we see it to-day in the case of the adult gibbon, and only the inner surfaces of the hands and feet are left free. Evidently these free places were uncovered, even in the ancestor which this human embryo copies for a short time. This Esau-like covering of the human being does not disappear until immediately before birth, and in a few exceptional cases, this covering has even been retained during life. This is the origin of the renowned men with dog faces. Now we come to a new question. What is the ancestor of that archetype? In what other dis- 55 THE EVOLUTION OF MAN guise can we trace him further back? In the system, the four anthropoid apes are followed by the rest of the monkeys. This class again con- sists of at least three great groups which differ from one another. Some of them are the long- tailed monkeys of Asia and Africa, such as Macacus, baboons, etc., which make up the ma- jority of the popular monkeys in our zoological gardens. The second group lives exclusively in America, and the bright Capuchin monkey may be mentioned as a type. The third, also re- stricted to America, comprises a small number of little monkeys, having claws instead of nails on most of their fingers and toes and resembling much more a squirrel than a genuine monkey. The marmoset is one of them. These three groups can no more be used in the construction of a consecutive line of development than the four anthropoid apes. But a purely anatomical comparison leaves the impression that somewhere near them the next lower stage of man must be found. Even the very first experts who described the gibbon noticed that this same gibbon, aside from his strong resemblances to the other anthropoid apes and to man himself, also had certain other resemblances very plainly developed, and these 56 THE EVOLUTION OF MAN pointed towards the Macacus-like long-tailed monkeys. These characters could be inherited only from the archetype, and this_Jyge_aga_iri could only have inherited them from some still older type, which had a general and much great- er resemblance to the majority of the other mon- keys. That there was once upon a time a cer- tain ancestor who had an externally visible long tail is still evidenced by man himself. Not only is man in the tailed stage to this day, though the tail vertebrae are no longer externally visible, but these are certainly still better developed in man than in the anthropoid apes. Furthermore, the human embryo in the mother's womb once more reveals the persistency of that mysterious biogenetic law. It has a plainly, visible external tail. In exceptional cases this "embryo tail" is also preserved in adults, and in some cases we have those abnormal "tail men," whose existence has often been doubted, but who nevertheless exist. There is no reason why we should not as- sume that certain Macacus-like types, preceding the human type, carried a genuine tail for a con- stant characteristic. So far as we can judge from fossil remains of bones, genuine long-tailed monkeys, similar to those in present Asia, were already in existence in the middle of the Tertiary 57 THE EVOLUTION OF MAN period; in which both man and anthropoid apes were found. One species, Mesopithecus, lived in great numbers in Greece, where many bones of them have been found. This Grecian monkey had a very long tail. At the same time the form of its nose and the position of its eyes gave it a greater resemblance to the human being than any of the present long-tailed monkeys have. • On the other hand, the light hearted crowd of long- tailed monkeys has developed many characteris- tics which tend toward a direction leading away from man. There are, so to say, one-sidedly bestialized forms, an extreme exaggeration of which is the baboon family, for instance, the grotesque mandril. The conclusion is inevita- ble that once again, at this point, a line of descent originally close to man has gradually deviated into a bypath and produced many varieties of monkeys now living in Asia and Africa. There^ fore we should once more have to assume the existence of an archetype out of which de-j veloped, on the one hand, the original ancestor of man and of the anthropoid apes, and, on the other, that Grecian Mesopithecus and the many side lines of African and Asian long-tailed mon^ keys. Of course, this archetype would have to be still a great deal more ancient than the pre- 58 THE EVOLUTION OF MAN ceding one. It might have existed as early as the first third of the Tertiary period. By its ex- ternal characteristics, we should certainly have classed it among the genuine monkeys, and only a few slight anatomical marks would have be- trayed to the expert that he was not dealing with a monkey of later descent, but with one in which, so to say, the third generation of coming man was still concealed. Now, it is peculiar that we have actually found remains of monkey-like animals in the first third of the Tertiary period. They were discovered by the Spanish explorer Ameghino in Patagonia, the extreme end of South America, and were concealed in a layer of rock which must have been developed toward the end of that first third of the Tertiary period. We call this first third the "Eocene" period, or in English the dawn of the more recent period. When Ameghino first analyzed one of these Patagonian monkey skulls, it conjured up to his imagination the ghost of a very small man, so that he called it "Homun- culus," but it seems that after all this resem- blance to man is not much greater than that of the American monkeys of the Capuchin type, and that group of Eocene monkeys evidently be- longed to that class. It cannot be denied that the 59 THE EVOLUTION OF MAN present Capuchin monkey is in many respects, physically and mentally, man-like. It also has secret relations with the gibbon, and thus to the archetype of the Pithecanthropus kind. Thus, many things favor the more recent assumption that possibly these bright, gentle and highly in- telligent American Capuchin monkeys are the closest of any of the present monkey forms to that genuine monkey type of man which belongs to the Eocene period. On the other hand, the small and squirrel-like marmosets must be eliminated from our line of descent and regarded as a side line. Most likely they are a one-sided adaptation to special con- ditions in South America. But now that we have gotten so far, there can be no doubt as to the next question. If man can be traced so far back in monkeydom, he cannot but share all the vicissitudes of monkey life fur- ther back. Whatever may be the general descent; tof monkeys, that is at the same time the line of! jman's development. The prototype of monkeys! |is also that of man. The conventional system of mammals proceeds along a great downward scale. First we have the prosimiae, bats, insectivora! such as the hedge-hog, then ^rnivorjj ro^^itsf the large and 60 THREE SKULLS FOR COMPARISON. The skull at the top is thai of a young Gorilla, the one in the middle that of an old Gorilla, the one at the bottom that of a Man. Note how much more man-like the skull of the young gorilla is than that of the old one. 61 THREE SKULLS OF DIFFERENT MAMMALS. At the top is the skull of a carnivore, a Cat; in the middle that of a rodent, a Hare; and at the bottom that of an utura- fete/a How. 62 THE EVOLUTION OF MAN variegated group of ruminants/ etc. But this scale is only apparently a historical one. Who- ever were to imagine that man went through all these different stages in succession would not come to any definite result. For instance, if we compare the teeth of a rabbit with those of a monkey, we should have considerable difficulty in accepting the idea that the monkey could be descended from a rabbit. It is the same when we compare two styles of architecture. The one is simple and noble and the other a sort of bizarre caricature of the former. We do not take kindly to the idea that the simple style should have developed from the caricature. Just so, the rows of teeth of mon- keys, including those of man, give the impression of a simple temple of noble style, in which every- thing is developed in conformity with a definite and uniform system. But the teeth of a rabbit, of a horse, and even those of a cat, appear to us like a caricatured variation of that simple style, going to excess here, falling short there. Of course, the opposite idea that all these other groups of mammals should have developed from monkeys is equally improbable. The simplest historical premises oppose such an idea. Neither do the remains of bones of primitive animals 63 THE EVOLUTION OF MAN teach us that there were at a certain period, first, let us say, ruminants, later on, perhaps rodents, then carnivora, and finally monkeys. Nor do they show that there were at first no other higher mammals than monkeys, and then in successive periods ruminants, rodents, etc. We rather re- ceive the impression that all of these groups ap- peared simultaneously at a certain period. Now it is precisely the progress in our knowl- edge of extinct mammals which succeeded finally in leading us out of this labyrinth of contradic- tory assumptions. All those groups of mammals still appeared in the first third of the Tertiary period, the so- called Eocene period, to which we have re- peatedly referred. Monkeys, as we have seen, were among them. Hence, if we desire to learn more about the origin of these things, we must trace our steps further back, say to the beginning of this Eocene period. Now we have found in two places far distant from one another — in France near Cernays in the vicinity of Reims, and in North America in New Mexico — the bones of certain_^xtremely ojd, mammalsj)eloriging to just this period, and these bpnes^ explain the mystery very fully. On the one hand, all of these bones' have a very simple 64 THE EVOLUTION OF MAN and fundamental structure. They show a re- markable row of teeth without extremes, or cari- catured exaggerations, and the present monkey and human teeth are easily derived from them. Furthermore, these skeletons have four feet, or rather four hands, with five regular fingers, among them one very flexible thumb. This is another very good prototype of the monkey and human hand, which is so widely different from the claw of the lion, or from the shin and hoof of the horse. In place of nails, these five fingers had an indefinite sort of thing, half way between a claw and a hoof, which might easily have de- veloped into anything, say, a horse's hoof, a carnivore's claw, or the nail of a Simian, or a human hand. On the other side, these animals show the be- ginnings of certain divergences in the structure of their bones. Some of them have more of the rodent, others more of the carnivore, others of some dominating ruminant character. There is no doubt that these simultaneously represented a very ancient group of ancestors which was just then beginning to branch out into the various great side lines of mammals. And it is equally certain that one of these side lines was composed of monkeys. Of course this original side line 65 THE EVOLUTION OF MAN of monkeys must have resembled the original an- cestor in the structure of teeth and hands and must have been a straight continuation of its evolution in the best sense of the word. This ex- plains why man and monkey, who to this day possess the simple normal teeth and the primi- tive hand, give the impression, now that the an- cient group of ancestors has long become extinct, that carnivore, ruminants, etc., are nothing but very extreme caricatures of the archetype. Furthermore, the claim that the monkeys were really a side line of that very primitive ancestor, and the most direct side line at that, is substan- tiated by a study of those ancient bones of Cer- nays and New Mexico. Just as we still observe in those bones certain variations in the direction of carnivora, of rodents, of ruminants, so we also find a little group of animals which gradu- ally, but very decidedly, move in the direction of our monkeys. True, they are not yet genuine monkeys, but they certainly show an unmistakable resemblance to a certain group of mammals which have al- ways followed in the system directly after the monkeys, and which were often considered as some peculiar retinue of genuine monkeys, the so-called prosimiae. ' 66 THE EVOLUTION OF MAN TARSIUS SPECTRUM. This little lemur almost resembles a tree-toad and is related to the family-tree of man. THE EVOLUTION OF MAN To this day there is living in the Sunda Islands, that is to say, in the same locality where the gibbon and the orang are living, and where once upon a time Pithecanthropus struggled through his existence, a queer little creature, partly re- sembling a small monkey, partly a leap-mouse, with long stilted legs. This little creature is so funny in all its aspects that it has been called the "tree toad" among mammals. The official name of this little forest gnome is Tarsius spectrum. This Tarsius is counted among the prosimiae in the system. Quite a number of animals about the size of a cat belong to this group, some of them coming from Madagascar, and known as "Makis." Furthermore, there are the so-called Galagos and the very strange Finger-monkey. At a certain period there existed in Madagascar even some species of prosimiae, which were nearly as large as a man. Now this little Tarsius has a certain character which connects him very closely with the genuine monkeys, first of all with the American Capuchin monkey. Those who have been present at the birth of a human being will remember a certain bloody mass which is forthcoming after the birth of the child. This is the so-called placenta/ So long as the little human being rests in its mother's THE EVOLUTION OF MAN womb as an embryo, this placenta is its most important organ, because by its help, the nourish- ing juices from the blood of the mother pass into the body of the child and thus feed it. The various groups of mammals differ considerably in the method of forming this placenta in the mother's womb. Man and the anthropoid apes have their own peculiar method. This is another excellent proof of the close relationship between man and these apes, and it was a great acquire- ment for this science when Selenka demonstrated that these processes followed the same outline in the gibbon and the orang-outang as in man — a process which is otherwise found nowhere but in man. The Macacus-like, long-tailed monkeys follow a different method, and the American monkeys have another and more primitive one. Now, it is interesting to know that the prosimia Tarsius follows the model of the American mon- keys in forming this placenta, while the majority of the genuine prosimiae again go their own peculiar way. And since we have found in America very old bones of the species Tarsius, the probability grows that prosimiae of tfie Tarsius type may be the direct ancestors of the American monkeys. If so, it is at the same time the next station in the evolution of man. This 69 THE EVOLUTION OF MAN type of Tarsitis of the Tertiary period would certainly represent a further development of our old friends of Cernays and New Mexico, which show certain divergences from the original type in the direction of the prosimia. The scientific! name of these prosimiae is Lemuridae, and these very ancient ancestors indicating this direction ! have therefore been called "Pachylemuridae." Let us remark in passing that there is still a very little group of mammals, the so-called in- sectivora, such as the hedge-hog, moles, etc., that likewise have a placenta similar to that of Tarsius. It is among the hedge-hogs that this placenta is distinctly visible. The student can hardly fail to suppose that the hedge-hogs are likewise in some way closely related to the side line which branches off from the archetype in the direction of monkeys. However, this ques- tion is not yet settled. At any rate, the hedge- hogs give the impression of being members of a very ancient group, and they, more than any other living mammals of the present day, seem to have preserved most nearly, even in their ex- ternal structure, the actual form of that primeval group of Cernays and New Mexico. But, if we try to solve the question of the ancestors of that original group itself, we are 70 THE EVOLUTION OF MAN brought face to face with another historical fact. We have now arrived at the beginning of the Tertiary period. One step further back and we find ourselves in the age of the great saurians. The geological picture has now completely changed. We enter the Secondary period of the earth's history, that inconceivably long epoch in which the chalk cliffs of the Island of Rugen, the Jurassic' slate of Suabia, and the reddish sand- stone used in building the Strasburg Munster were formed. The greater part of the large fossil bones belonging to these days were the remains of giant reptiles, some of them resembling dragons. Those saurians swam around in the ocean like our present day whales, or they rolled around in the mud like our hippopotami. Some of them, resembling colossal kangaroos, grazed on the prairies like cows, lumbered about on their heavy hind legs, or jumped after their prey, and some of the most daring even rocked themselves on batlike wings high up in the air. It was not until gradually in the course of this geological period, which probably lasted many millions of years, that birds appeared for the first time — first of all, the lizard-bird Archaeopteryx. This transi- tion form shows very plainly in its structure that 71 THE EVOLUTION OF MAN birds are merely a side line of the great main branch of reptiles. Nevertheless, during this typical saurian age, there already existed some mammals, as is proved by the remains of their bones. True, they do not seem to have played a very prominent role. Their remains have been found, therefore, only in a few portions of the secondary strata. And all these scant remains belong to rather small animals, but such as they are they are well pre- served and teach us an important lesson. In our transition from the Tertiary period backward into more primitive times, we become aware of the fact that all higher mammals gradu- ally disappear, even that archetype of Cernays and New Mexico. Instead of them, the remains of mammal bones, wherever they may appear, be- long to representatives of a certain group of lower mammals, the so-called marsupials. /• The best known type of marsupial is the kan- garoo. But there are still a number of other rep- resentatives living, most of them in Australia, some of them also in America. These marsupial? have, among other peculiarities, a bony projec- tion in their lower jaw, and this always distin- guishes their jaw from that of any other mammal. The fossil lower jaws of these second- 73 THE EVOLUTION OF MAN ary mammals always have this very characteristic projection. They evidently belonged to a group of mammals whose last living representatives are the present-day marsupials. These bones are also found in Africa, Asia and Europe, proving that this race of marsupials formerly inhabited the entire earth. Under these conditions the assumption was justified that this primitive group of mammals represented the most ancient type, from which the tertiary archetype of the higher side lines might have developed. In that case, they once more show us another mile stone in the upward march of disguised man — a marsupial man as a contemporary of the Ichthyosaurian. This gen- eral conclusion is confirmed by a good many de- tails. Marsupials owe their name to a fact, which every child notices in the kangaroos of our zoo- logical gardens, that is to say, the female carries its young, which is born in an immature state, for a while in a protecting fold of its skin, the so-called pocket. In this pocket the young finds the milk-nipples which it uses in suckling as mammals do. To this day, the embryo of the highest mammals, including that of man, bears in the position and surroundings of its milk- 73 THE EVOLUTION OF MAN nipples certain indications which perceptibly point toward their original location in a pocket, a sure sign that the ancestors of all of them once went through the marsupial stage. A very good proof is furnished' by the present living mar- supials in their peculiar formation of that im- portant organ of propagation in the mother's womb, which we mentioned once before, the placenta. While in our previous remarks we mentioned only the different forms of this placenta, we now notice that the marsupials seem to have remained stationary at the point where the placenta was in its first stage of de- velopment. The majority of the marsupials have no placenta at all, and this is an indication of a former and still more ancient condition which is closely connected with the existence of a pocket and the premature birth of the young. The young was born so early and required the use of the milk-nipples so prematurely that it did not at all need a placenta connected with the nourish- ing juices of its mother's blood. On the other hand, a few species of Australian marsupials, the Perameles, show the beginning of a very simple and rudimentary placenta, and thus furnish an additional proof that this important organ went 74 THE EVOLUTION OF MAN through its first stages in the ranks of the mar- supials. In other words, the marsupials repre^\ pent the genuine ancient transition form from aj Uower to a higher mammal. We shall have to A SPECIES OF PERAMELES, a type of lower marsupials, which, however, forms a sort of placenta and thus approaches the higher mammals. assume that the progress from these marsupials with primitive placentas toward that archetype of Cernays and New Mexico took place during the chalk period, that is to say, the last great 75 THE EVOLUTION OF MAN division of the Secondary age. It is important/, *to note at the same time that a hand with five fingers and a flexible thumb, which have been so faithfully preserved by prosimiae, monkeys, an- thropoid apes and man, are found among the climbing species of marsupials, especially theX /American oppossum. Before the external nipples of the breast are formed in the human embryo, the milk gland is formed in the skin. If we remember the bio- genetic law, it seems to us that the milk gland existed in our ancestors at a certain stage be- fore the genuine nipple of the breasts came into existence. At the same time we see the human embryo at a certain early stage of its development with a very peculiar construction of its posterior opening. The opening for the products of the urinary and sexual organs is found in the rectum, so that there is only one single opening for all three things, the products of the digestive, the urinary, and the sex organs. It is not until the third month that a partition is formed in the rectum of the growing human being by means of which henceforth these excretions are divided and discharged through two openings, one for the products of the urinary and sex organs, the other for the products of digestion. This succession of 76 THE EVOLUTION OF MAN organs compels us to consider whether we have not to deal in this case with a very old inter-rela- tion of things. Could it be possible that mammals concealing man existed once upon a time which possessed milk glands, but no external nipples, and which had only one single opening for the products of the urinary, sex and digestive organs ? There are such mammals even in our day. They are known as Australian duckbills. One species of them, living on dry land, called Echidna, resembles a large hedge-hog and is protected by strong quills. It lives in Australia, Tasmania and New Guinea. Another kind, liv- ing in the water and called Ornithorhynchus, resembles in its pelt and habits the otter. It swims very well and lives in the little rivers and lakes of the Australian continent. Both duck- bills are without external nipples, but they have genuine milk glands. The milk percolates through a sieve-like place in the skin into the mouth of the young. At the same time the body of the duckbill has only one single opening for the products of the urinary, sex and digestive or- gans. Injhe_ system these duckbills follow after the marsupials. Neither of them has any placenta. Nor do they need it, and that fpr a very good 77 THE EVOLUTION OF MAN ECHIDNA HYSTRIX, the land duck-bill, an archetype of mammal living in Australia, which lays eggs like reptiles and birds, but suckles its young after they have hatched. Its egg, natural size, is shown to th« right. reason. They actually lay eggs in the regular way. The young is born in an egg with a parch- ment-like wall, just like a young turtle or lizard. But while it is hatched from this egg like a young bird, it licks up the milk of its mother mammal fashion. The terrestrial duckbill has furthermore the method of marsupials ; it carries first its egg and then its young in its pocket. The aquatic duckbill, on the other hand, does not use this method any more; it digs a hole in the bank of a river, makes a regular nest and there lays its eggs openly, just like a bird. The inevitable conclusion from these premises 78 THE EVOLUTION OF MAN ORNITHORHYNCHUS PARADOXUS, the -water duck-bill, a mammal of Australia which lays eggs and is closely related to the primeval ancestors of present-day mammals. is that these duckbills show us the more ancient group of ancestors below the marsupials. In other words, Australia has preserved for us a few "last Mohicans," witnesses to a certain stage in the development of mammals and of man in the far off days of the primitive world. And all that would now be required to complete the proof would be genuine historical testimony given by primitive fossil duckbill bones, such as furnished the required proof in the case of the marsupials. For a while it seemed as if this group would not be forthcoming. It is true that various little teeth and remains of small mammal bones, not belonging to any class represented by living mammals, not even the marsupials, were found in the strata of the age of the great saurians far 79 THE EVOLUTION OF MAN into this first third of the so-called Trias period. Most of the discoveries consisted of teeth, but neither of the two duckbills now living has any teeth. They are called duckbills because their toothless jaws are covered with a horny skin giving them the shape of bird's bills. The aquatic species especially has a genuine duck bill. However, one fine day the biogenetic law once more came to our rescue. A young duckbill de- velops in its first stages a sort of milk teeth, hav- ing the early characteristics of molar teeth. No teeth of any other living or extinct animal cor- respond to the form of these teeth of the young duckbill — with the sole exception of those fossil teeth of the saurian age. Hence we conclude that the toothless bills of the duckbills, in spite of the fact that they look so queer in a mammal, do not represent an ancient heritage. They are rather a newly acquired character, an adapta- tion, which these surviving Australians have ac- quired during the long period that has elapsed since then. Their ancestors in the saurian age, who were at the same time the genuine ancestors of the higher mammals, had teeth, and these are the very teeth which we now find in a fossil state. These ancient duckbills with teeth, as one might 80 THE EVOLUTION OF MAN call them, if this term were not self -contradictory, are known by the scientific name of AUotheria. When duckbills first became known, their bills were, of course, the first thing that gave rise to comment. Owing to their presence these mam- mals, which otherwise had all the marks of a mammal, gave a decided impression of a cross with birds. For this reason some people specu- lated from the beginning whether these queer creatures did not actually represent the transition of a mammal to a bird. In the light of the ex- planation given just now, we are not very much impressed with this speculation, for the bill ap- pears as something unessential and subsequently acquired, which has about the same significance as the whalebone in the jaws of the whale, or the exaggerated claws of the sloth. But the otherx /.characteristics of these duckbills concern us much more. There is above all t which had not been ascertained by the first ob- servers. This habit indeed indicates the descent of mammals from a lower class of vertebrates. But this lower class need not necessarily be birds, for reptiles, amphibians and fish also lay eggs. Indeed, the egg of a duckbill resembles much more that of a reptile, such as a lizard, or a turtle, than that of a bird. And if we consider 81 THE EVOLUTION OF MAN the structure of the skeleton, the resemblance to reptiles exceeds that to birds. The duckbill, the contemporary of saurians, seems to lead directly to the saurians, without touching the birds. The straight succession of our system misleads us in this instance. Birds represent a subsequent and one-sided branch line of reptiles, and have evidently nothing at all to do with the develop- ment of mammals. It is true that the birds have also permanently warm blood like mammals, and owing to this similarity they have been placed side by side in the system. A bird has often warmer blood than a mammal. But this again is one of those qualities which, though indicating a higher stage, were nevertheless acquired inde- pendently in widely different periods. In this connection we might point to the fact that the representatives of other dissimilar groups of animals have acquired the faculty of flying inde- pendently and at far distant stages. This is the case, for instance, with flies, bees, dragon-flies, butterflies, flying-fish, frogs, such as the flying frog of the Sunda Islands which flies by means of a skin between its separate toes, and lizards, such as the Australian flying lizard. There are, fur- thermore, the birds, and among mammals, the bats and the flying squirrels. There can be no 82 THE EVOLUTION OF MAN question whatever of any comparison between the one or the other of these groups in the matter of the flying apparatus. Each one of them, un- der pressure of conditions, has separately ac- quired this adaptation. A number of the old and extinct saurians, such as the Dinosaurians, the Pterodaktyls, or flying dragons, must have been in possession of permanently warm blood, so far as we are able to ascertain. A few snakes, such as the python, develop to this day warm blood, under certain conditions, for instance, when they have laid eggs and wish to give them a cer- tain amount of heat in hatching. So, it was na- tural that the bird should acquire for life a cer- tain faculty which appearel already among rep- tiles from which it is descended. As we have seen, the strange Archaeopteryx still represents an unmistakable transition form from the general reptile type to the bird. On the other hand, no visible line leads from birds to mammals. The bat is no more such a transition stage than a whale is a transition from mammals to fish. In both cases relatively highly developed mammals have acquired independent adaptations, the bats a flying apparatus and the whales a swimming apparatus. It is not difficult to imagine that the feathers 83 THE EVOLUTION OF MAN of the bird developed out of the scale of the lizard. But it seems quite improbable that either a scale or a feather should have been transformed to such an extent as to assume the characteristic form of hairy covering typical of mammals. Scales, as well as feathers, have evidently been from the very beginning essential means of pro- tecting the skin, either in defense against enemies, or in the case of birds, against the in- clemencies of the weather. We observe that scales serve that purpose occasionally even in mammals, for instance, among armadillos. Some whales likewise possessed something like that in former times. But the typical covering of the skin of mammals consists of hair. And it seems that originally hair had nothing to do with pro- tection such as is afforded by scales or feathers, but rather served a wider purpose, embracing not only protection but essentially feeling. The first hair consisted of very fine feelers and performed the functions of touch for the skin. It was not until later on, when mammals acquired warm blood, that hair assumed also the role of a non- conductor of heat. Now when we look about us to find the be- ginnings of sense organs of the skin which might have developed into hair, for instance, among 84 THE EVOLUTION QF MAN lower vertebrates than mammals, we are carried even beyond the scaly reptiles into the ranks of amphibians with naked skins. In distinction from reptiles, such as lizards, snakes, crocodiles and turtles, the amphibians embrace newts, toads and frogs. While these animals do not have any hair, they nevertheless have peculiar little sense organs precisely in those places of the skin which, among mammals, carry hair and which correspond pretty closely in their arrangement to the plan of the hairy cover- ing of embryos among mammals. According to the biogenetic law, this might very well indicate that the amphibians still show to-day the primi- tive form of a genuine hairy covering. We might well conclude from this fact that the most ancient mammals, for instance, those creatures belonging to the duckbill family which we dis- cover in the first third of the saurian period, the so-called Trias, are not descended from genuine reptiles, but rather from amphibians which oc- cupy a still lower position in the system. Now, it happens that the living representatives of amphibians still possess many a detail which might be regarded as an indication of the direct descent of mammals from them. It is remarka- ble that many frogs and toads have very signifi- es THE EVOLUTION OF MAN cant habits of primitive care for their offspring. Sometimes it is the males, sometimes the females, that carry the eggs round with them. The male of the European "Obstetric Toad" has the habit of taking the spawn from the female, wrapping it in strings round its hind legs and taking great care to protect it. The female of the Pipa of South America, on the other hand, carries it.c, eggs on its back, having little pockets in the skin of its back in which the eggs gradually mature and in which the young hatch. Among other toads, the skin has developed large hatching pockets in which first the eggs and later the young animals are carried about in just the same way that we observe among the land duckbills and the marsupials. Furthermore, various glands of the skin play an important role among amphibians. Everyone is acquainted with those glands of the toad which excrete a sharp juice serving as a protection against enemies. But such glands as those play a role in the formation of the pockets of the Pipa. It is not a very far- fetched idea that the young animal hiding in such a pocket might also begin to lick the excretions of its glands, which need not necessarily be caus- tic, but may serve as nutrition. If that is so, we should find ourselves at once at that stage which 86 THE EVOLUTION OF MAN is represented among mammals by the duckbill, the young of which licks, during its stay in the pocket, the percolating juice of a gland. On the other hand, it cannot be denied that the general construction of a duckbill has many A HUMAN EMBRYO, in the middle of the fifth week of its development. It is strongly magnified, its natural size being about one centimeter. Note the gill-opening on the neck, the fin-like limbs, and the plainly developed tail. points resembling those of saurians, in other words, of reptiles. The only marked difference in their skeleton is the way in which the lower jaw is attached to the skull. This separates rep- tiles and mammals very distinctly. Indeed, the 87 THE EVOLUTION OF MAN EMBRYO OF A LAND DUCK-BILL. Note the similarity to the embryos of man and monkey, both of them in the same stage of development. EMBRYO OF A MONKEY in about the same stage as the human embryo. Note the simi- larity between them. 88 THE EVOLUTION OF MAN attachment of the lower jaw to the skull in rep- tiles and mammals represents the two extremes of two independent methods. Now the study of the fossils of the primitive world gives us some clue toward a solution of these contradictory questions. The historical time which we should expect to represent the transformation of the most ancient duckbill- like mammals from the archetype next below them in the scale of evolution, would be about the transition from the Primary to the Second- ary period, that is to say, a time midway between the carboniferous and the first great saurian epoch. As it happens, it is precisely this time which again gives us some fossil testimony touching unmistakably on the question now un- der discussion. The present living representatives of am- phibians, such as newts, toads and frogs, were evidently not in existence at that early period. They are apparently a late bud on the branch of amphibian descent. But in their place there existed very strange and large amphibians, some of them resembling crocodiles with more or less solid bony armor. These amphibians possessed many reptilian marks, so that they give the im- pression that they were in transition from am- phibians to reptiles. 89 THE EVOLUTION OF MAN Simultaneously with them, there lived certain reptiles, small saurians which in many import- ant respects looked like amphibians and on their part represented a mixed group, the other end of the bridge, so to say. Thanks to a happy co- incidence a living grandchild of these amphibian reptiles of the primitive world is still found at this day in New Zealand. Its name is Hatteria punctata. Its entire construction is such that it represents a splendid illustration of the transition form combining the newt and the present-day lizard in an almost neutral shape. Finally, as a third count, we mention the fact that genuine large reptiles, some of them very grotesque in form, lived in those primitive days. The strange thing about them is that they have undeniable resemblances, especially in the struc- ture of their teeth to mammals. These are the so-called Theromorphoi. Their bones have been found mainly in South Africa, in Cape Colony. Their resemblance to mammals was so striking that their first discoverers naturally thought they had found typical transition forms from reptiles to mammals, and there are still many experts who share this view. Nevertheless, the genuine reptile marks, for instance, the adjustment of the lower jaw, typical of the saurians, are so unde- 90 THE EVOLUTION OF MAN niable that there are strong objections to an en- dorsement of that view. It is not credible that the reptilian type should have been so well de- HATTERIA PUNCTATA, a reptile living in New Zealand, which is a surviving type of the oldest primeval saurians. veloped by evolution in the first place and then continued on towards the mammal type. If we weigh all the facts, it appears most probable that a mixed group of ancestors existed 91 THE EVOLUTION OF MAN in those days of the latter part of the Primary period, but that this group combined in the germ amphibians, reptiles and mammals, just as we saw at a later stage that the oldest mammals of the Tertiary period took their departure from a mixed group which contained the possibility of evolution into Carnivora, ruminants, rodents and prosimiae. The members of this mixed group may have resembled the present-day amphibian newts, so far as the naked skin full of glands and sense organs was concerned, and they may have had p6ints of contact with them also as regards their mode of living and otherwise. Their lower jaw may have been so constructed that it might de- velop in the style of a genuine reptile as well as the other extreme of the genuine mammal, and the remainder of its bony structure may for many ages have resembled the living Hatteria, while other characteristics may have recalled the duckbill. Surely, their feet had five regular toes, one of them probably being a flexible thumb, in other words, the basis of the later "hand." The teeth of this group must have pointed in the di- rection of mammals. This mixed group branched off into the various side lines which we have already observed, each 92 THE EVOLUTION OF MAN one of them laying special emphasis on certain points of the old form, showing the naked newt in one place and the more reptile-like extinct armored amphibian in another, and a genuine reptile in still another place. The reptilians may at first have assumed such forms as we still ob- serve in Hatteria, and out of genuine reptiles developed the birds at a much later period. Still another side line would be represented by those Theromorphoi of Cape Colony which, on the whole, had a pronounced reptilian character, but still preserved in their teeth and in a few other points, such marks as have become typical later on only for mammals. Finally, running parallel with all the others the genuine mammals would have gone their own way. There is nothing of any consequence to pre- vent us from assuming that these mammals, which reached their highest stage in man, formed the central line or the crown of the entire line of descent. At any rate, they were the most intel- ligent line, and they may also have been the most favored physically and have deviated less from the characteristics of the great archetype. In view of all the facts known in this case, these conclusions seem certainly logical and sound. It is true that genuine fossil remains of this 93 THE EVOLUTION OF MAN hypothetical mixed group have not yet been lo- cated. But it must be remembered that our dis- cussion is now dealing with sections of the earth's history which are extending into eons of time in which all things are becoming indistinct and vague. To the extent that we venture into the dim past, our proofs must be founded more and more on circumstantial evidence. No one could expect that all the typical stages and their inter-relations should be distinctly seen, it must be sufficient to trace in its approximate outlines the logical course of the main growth. There are a great number of special witnesses to make a good case for our further investigation. We have now gotten far beyond the saurian period into the so-called Primary age. We are approaching those most ancient epochs which gives us any direct evidence of primitive life on earth by means of petrified specimens. We meet in that period numerous masses of mineral strata, which were once precipitated to the bot- tom of the sea in the form of mud. These strata bear no other fossil remains of animals than those of fishes. Evidently these were then the sole representatives of the animal world. We receive the impression that all animal life! I at that remote Primary age was concentrated in | 94 THE EVOLUTION OF MAN fishes, amphibians, reptiles and all other verte- 1 brates being contained in fish and no other verte- jbrate existing beside them. This historical testimony happens to coincide exactly with the conventional system in which the fish follow immediately after the reptiles and amphibians. A fish is distinguished from an adult newt, frog, lizard, turtle, bird or mammal, including man, by the way in which it breathes. All other vertebrates breathe through lungs in the open air. But fish represent a perfect adapt- ation to life in the water. Since a fish, however, also requires air for breathing, it has developed an organ which, being continuously surrounded by water, can assimilate the air contained in this water. This organ consists of the so-called gills located in the neck of the fish. Now, it is a fact well known to every school boy that the so-called tadpole hatches out of the eggs of newts, frogs and toads. This tadpole lives exclusively in the water exactly like a fish, and breathes only through regular gills. Not un- til the newt or frog abandons the early stage of the larva, does it acquire the faculty of breath- ing through genuine lungs and shed the gills, much in the same way that human children shed their milk teeth. The tadpole is nothing less 95 THE EVOLUTION OF MAN than an embryo set free. * And from the genetic law, which recognizes in the embryo the portrait of its ancestor, we conclude therefore that newts and frogs are descended from crea- tures which breathed through gills, that is to say, descended from fish, since they are the only vertebrates from which we may choosex But if these newts and frogs, according to the assumption that we made a while ago, are noth- ing but a side line of that main group from which mammals also developed once upon a time, nothing remains for us but to assume that this main group in its entirety leads back to a preceding station of water animals breathing through gills. Some one may object and ask how it is that no other animals besides frogs and newts, say, for instance, reptiles, birds and mammals up to man, have preserved breathing through gills in the embryonic stage. Why does not a young human being first become a tadpole before it be- comes a man? Well, in the first place, the bio- genetic law is not absolute. Very often it shows itself only in dim outlines. On account of subse- quent adaptation for purposes of protection, or for other reasons, some of those reversions to type have been subsequently eliminated. The 96 THE EVOLUTION OF MAN most useful character in the last analysis pre- vailed. And wherever a repetition of the charac- teristics of ancestors was too tedious, this or that stage was finally restricted or entirely eliminated. What good could an early tadpole stage in the water do a bird or a mammal ? On the contrary. We see often among certain frogs and newts a tendency to transfer the tadpole stage into the egg, or to go through it before the young is hatched at all. There is, for instance, a tree toad on the Island of Martinique which has become known through such a simplification of the evo- lutionary process. The tadpole of this little toad no longer hatches out of the egg. But granted that all this is so, should not the embryo of mammals, reptiles and birds show at least traces of a tadpole or fish stage in the mother's womb, or in the egg? It is the most remarkable proof of the reliability of the bio- genetic law that this is actually the case. No matter what embryo we may study, whether it is that of a lizard, a snake, a crocodile or that of the New Zealand Hatteria, or of a turtle, an ostrich, a stork, a chicken, a canary, a duckbill, a marsupial, a whale, a rabbit, a horse, or finally of a long-tailed American monkey or anthropoid gibbon — the embryo at a certain stage 97 THE EVOLUTION OF MAN of its development always shows a perceptible tadpole or fish stage. Its neck shows the marks of the gills and the characteristic intervals be- tween them by which the fish breathing in water permits it to circulate freely and flow around the breathing surfaces of the gills. Furthermore, the limbs which the embryos are just forming at this stage have likewise the plain outlines of fins. They push outward in the shape of round disks, and it is only the subsequent development which results in their further transformation, that is to say, into actual fins here, into the swift lower leg of 'a horse with a single toe there, or finally into the wings of a bird or the flying hand of a bat. If any strict scientific proof were still needed for our claim that all these higher vertebrates con- verge into a common archetype, it is obviously given everywhere by this common heritage of a gill and fin embryo, either in the egg or in the mother's womb. The gills and fins show that the oldest archetype, with which we are now dealing, was represented by a gill and fin animal — in other words, by a fish. There still remains this question to be an- swered: How is it with human beings in this respect? Every text-book on anatomy to-day gives a satisfactory answer. The embryo of 98 THE EVOLUTION OF MAN human beings at a certain stage is likewise pro- vided with traces of gills on its neck and with finlike disks in the places where arms and legs develop later on. This is as universally accepted as the fact first stated by Copernicus that the earth revolves around the sun. No man who has the least respect for truth can deny this fact. Nevertheless, there are people who find this very plain fact of embryology very little to their lik- ing, and who therefore frequently attempt to brand it as a "falsification." But every university text-book in the hands of every student of medi- cine, which is used as a basis for the state exami- nations, contains a statement of this simple fact, and if any student were to deny it during his examination he would be severely reprimanded by the state examiner. People who still refer to such undeniable and scientifically recognized facts as falsifications place themselves outside the pale of all moral premises and scientific re- search. It is a fact, then, that man is likewise de- scended from the fish. But, if we ask how it happened and what were the external causes which transformed in the far off primitive days a fish breathing through gills into a land animal breathing through lungs, 99 THE EVOLUTION OF MAN there is once more a living form which gives a direct clue. In a few small rivers of the eastern part of the Australian continent, a creature has been found which externally, so far as scales, fins and gills are concerned, resembles a large salmon or carp. But if we study its internal structure we find that it has also perfectly developed, serviceable lungs, and if we study its mode of liv- ing the logical purpose of this double supply of breathing organs becomes plain. During the dry season the little rivers of this region dry out almost completely. Nothing remains of them but a few pools of bad, brackish water in which the fishes are crowded together and encroach on one another's supply of air. Under these trying conditions, this strange animal swims to the sur- face of the water, draws air into its lungs and thus breathes after the manner of a genuine land animal, which dispenses altogether with water for breathing purposes. This paradoxical fellow who can change him- self at will into a fish and into a newt, has been called the "newt-fish," and its Latin name is Ceratodus. But this name was originally in- vented for the purpose of applying it to a band of fishlike creatures, which may be traced by fossil remains throughout a long evolution far back into 100 THE EVOLUTION OF MAN the earliest Primary period, and which are distin- guished by very peculiar teeth in the roof of the mouth. And the Australian Ceratodus of our day has exactly the same kind of teeth. Hence, CERATODUS FORSTERI, living in Queensland, Australia. It has gills, the regular breathing apparatus of fishes, but also lungs, the breathing organ of adult newts, frogs, reptiles, birds, and mammals. we logically conclude that it has preserved this peculiar double method of breathing from the days of primitive creation, and we refer to it as a last straggler of a real transition group from 101 THE EVOLUTION OF MAN primeval fish breathing through gills to the first primeval animal breathing through lungs — in other words, to that theoretical mixed group con- taining the principal characteristics of amphib- ians, reptiles and mammals. The fossil remains of those primitive relatives of Ceratodus are con- sidered as parts of creatures belonging to this transition group. /At all events, this Australian* )t Ceratodus shows very clearly what the conditions are in which a lung may develop. This is simply the outcome of lack of water, or lack of air in « /the water. / Some might ask how it happened that a new organ could develop just when it was needed most, very much like a fairy table which is set whenever the wish is expressed. The witchery of nature can never come out of the unknown; it has always some logical connection. Indeed, the lungs of Ceratodus on closer study reveal the fact that they are merely a transformation of an organ which all genuine fish carry with them — the so-called swimming bladder. This swimming bladder forms a sort of balloon filled with air in the body of the fish, and it serves in the first place as a means of overcoming the weight of the fish in the water. This organ fulfills a useful purpose in rising and sinking, and to this end it was pro- 102 THE EVOLUTION OF MAN vided with a valve for its regulation. Many fish therefore retained an open connection between the swimming bladder, the intestines and the mouth for the purpose of inhaling or exhaling air. This is the starting point of the lungs. The balloon or bladder in the vicinity of the aesopha- gus being filled or closed at will, it served at the same time to feed the arteries of its walls with A SHARK. The sharks are a primeval group of ancestors of fishes and are related to the family tree of man. oxygen. Once it had come into operation, it could under severe conditions be used as a sub- stitute for the gills in times when water was scarce. When these conditions continued for a long period of time, the swimming bladder as- sumed this role permanently and became a genu- ine lung, while the gills atrophied until nothing remained of them but traces in the embryo. Thus the land animal sprang into being, or to express 103 THE EVOLUTION OF MAN it with a view to the great line of our evolution, man emancipated himself from the stage of water animals. Now that we have found that Ceratodus is the living representative of another "bridge/' or at least one side of it, we are naturally anxious to find the other pillar of that bridge. That is to say, we should like to know what other species of fish helped to make this bridge, for there is a wide difference between fish and fish. When we mention fish to a layman, his first thought is of those kinds which he finds on his table and with which, under the present condi- tions of natural education, he is more familiar than with zoological literature. The overwhelm- ing majority of table fish consists of so-called "bony fish," that is to say, all of them have a more or less solid skeleton. All of the European river fish belong to this class, the trout, the pike, the carp, etc., and such sea fish as flounder, herring and codfish. If we find a jar of delicious caviar on our table, or if our meal is crowned with ex- pensive Russian sterlet, we meet another group of fish, the so-called Ganoids. The proudest rep- resentative of this class is the sturgeon, the eggs of which are used as caviar. These Ganoids are especially distinguished by the fact that some of 104 THE EVOLUTION OF MAN them have a very soft skeleton, consisting of cartilage instead of bones. This cartilage skeleton becomes permanent in a third group, which is not admitted to our table, but may be found on that of the Chinese, the sharks, which are known at least by name to everybody. Separated from these three groups of fish by a wide chasm, there is a fish-like creature which is very highly appreciated by gourmands— the lamprey. "' Finally, there remains one solitary and very strange little fish, the so-called lancet-fish or Am- phioxus, which is distinguished from all other fish by the extreme simplicity of its structure. A comparison of these five groups of fishes leads to the following conclusions in regard to the descent of man: If it is a fact that Ceratodus is actually a part of that bridge which connects with man, then the other end of that bridge could not be found among fish with a bony and solid skeleton, but among those which have a cartilaginous skeleton, the foremost ofk which are the sturgeons. Cera- todus itself still has a soft skeleton similar to that of the sturgeons. It is true that amphibians, reptiles and mammals have a very solid skeleton, 105 THE EVOLUTION OF MAN more solid even than that of the trout and her- ring. It is evidently a separate adaptation. The connecting link following the Ceratodus class is found below the entire class of bony fish which are once again a special side line. The Ceratodus class of fish have still other conspicuous relations to fish of the sturgeon class. A LAMPREY, the representative of a group of fish intermediate between the Amphioxus type and the Sharks. The historical evidence coincides with these marks, for sturgeons were present in extraordi- narily large numbers during the Primary age. In fact, there were so many species of them that they represented for a while the entire fish family on this earth. Wherever we see in museums 106 THE EVOLUTION OF MAN their beautiful resplendent scales, there we are face to face with another disguise of man, which takes us far back to the very beginning of the Primary age. Taking the soft skeleton as a basis for further research, it becomes evident that sharks are the next stage in our line of descent. The sharks played likewise a very prominent role in those primitive days, and to this day they are the most dangerous as well as the most intelligent of all fish. In a multitude of fine traits the shark is a genuine prototype of higher vertebrates trans- lated into fish life. The simple plan of four limbs is sharply outlined in its fins, that charac- teristic which has become so full of meaning in the subsequent evolution. Our teeth, which have become so typical in their present form as a mark distinguishing man from all other animals, may be derived by strict anatomical logic from a basic plan found among sharks, which is actually start- ling for the layman. The shark has a formidable set of teeth. But, in its mouth, it has also de- veloped a special trait in the way of thorny bristles which also appear in a less developed form in other parts of its body. The entire sur- face of the shark's skin is covered with peculiar and very fine, but rough, prickles, and the skin 107 THE EVOLUTION OF MAN of the mouth has developed these into specially strong and solid thorns, evidently for the good purpose of holding on to the food of these fish. This is a typical illustration of the genesis of "teeth" and without this clue it would be a very difficult problem to explain their origin. There are still further points of evidence. We have just seen that the shark has the basic plan of four limbs in the form of fins. The lamprey, on the other hand, has none of that as yet, but it has the beginning of a skull in a sort of skin and cartilage* pouch. Amphioxus, finally, has not even a trace of that. This would give us another chain of evidence. XThe line of evolution seems* to go upwards from Amphioxus by way of the lamprey to the shark, and other things which we observe at the same time fit very well into this x ^outline. Throughout this region in the process of evo- lution we find a number of details which do not become intelligible until we meet them again in very perfect forms in far higher stages. In the life processes of some sharks, for instance, a genuine placenta formation will suddenly appear, the embryo being nourished through a placenta. Like a flash of lightning the thought strikes us that nature at this stage suddenly tried some- 108 THE EVOLUTION OF MAN thing which was temporarily feasible and served as a means of adaptation, but was soon dropped, and did not reappear and become typical until the mammals arose. Again, in the development of the eggs of the lamprey, we see a sudden flaring up of almost the identical method which has later become typi- cal for the amphibians of the present day. All these things indicate that at this stage we meet once more one of those ancient mixed groups, typical for our line of descent, which contained the historical germs of all higher forms and were, so to say, reservoirs for all the possibilities of subsequent evolution. At the same time, we approach at this stage an entirely new and exceedingly significant point of departure, the source of all vertebrates in gen- eral. What is the characteristic mark of a verte- brate, including man ? The back bone, that great internal prop of the body. Well, then, we see the back bone growing softer and softer among the Ceratodus class, sturgeons and sharks, and it seems to dissolve more and more the further back we trace it. In the lamprey and finally in Am- phioxus .this backward formation is almost com- pleted. There the proud column has become 109 TPIE EVOLUTION OF MAN quite a thin thread of cartilage. It looks as if the backbone had gradually melted away like a piece of sugar in coffee. The spinal cord is no longer surrounded by solid bone, it extends through the body as a string of nerves, just as it does among worms or insects. And nothing indicates that typical characteristic which divides verte- brates absolutely from all other animals, but the position of this nerve string above the cartilagin- ous thread and above the digestive tract, while in all other animals the great nerve string is al- ways located below the digestive tract. The back- bone is here called merely the "chorda," and we are here evidently at the point where the verte- brates dissolve into invertebrates. And what does it matter? If man is disguised in a lamprey or an Amphioxus, then we may as well look for him entirely outside of the verte- brates. One species of lamprey, which bore their way into the bodies of other fish and live as para- sites upon them, were still mistaken for worms by Linnaeus himself. And the discoverer of Am- phioxus thought that he had found a snail, which it indeed resembles far more than a fish when we dig it up from its hiding place in wet sand and see its transparent and lancet-like little body. Any way, it makes no difference theoretically, no THE EVOLUTION OF MAN AMPHIOXUS LANCEOLATUS, and its development in the egg. This development shows cross- sections, first of a hollow ball, then of the stage when this ball begins to double up, until an almost closed body with skin and stomach membranes and one orifice is formed. The last stage represents the so-called Gastrula. Ill THE EVOLUTION OF MAN if we descend still further even into the world of the very low and entirely invertebrate animals. Of course, in practice we shall have to apply, still more than heretofore, what we have previously said about circumstantial evidence. In the first place, one source fails entirely at this point, that of geology. We are compelled to push backward far beyond even the Primary period into the very dimmest time. All direct proofs suddenly fail at this stage. There are no fossils beyond the Pri- mary ones. The minerals of more ancient epochs of the earth's development have been so trans- formed by a process of crystallization, the cause of which we do not yet understand, but which are in some way connected with pressure and heat, that impressions of fossil specimens of former living bones can no longer be discovered in them. Now these so-called crystalline slates are evidently the product of water, hardened sedi- ments of the sea, and there is no reason to assume that the sea in which they were formed contained no living beings at all. On the contrary, there are important reasons contradicting such an as- sumption. The animals of the Primary period are far too highly developed to represent the very first animals on this globe, unless we renounce the idea of development entirely and believe that the 112 THE EVOLUTION OF MAN first fauna and flora fell ready made from heaven. But the fact remains that from this time on we no longer find any remains of the ancient animals and plants, x If we wish to make further concluX vsions we can only rely on the now surviving lower and lowest creatures and look for points of con- tact with them in the embryonic stages of higher^ ^animal sx With this understanding, we now proceed to discuss the further evidences on which we may rely from now on. Among all the animals now known and living below Amphioxus, there is only one single small group which still shows a direct indication of a backbone, the so-called ascidians. These are small marine animals which -are surrounded by a cloak of wood-like substance, very much as snails sur- round themselves with a well-nigh closed house. To judge by their general construction, these ascidians would be most logically classed among the worms, save for a few points of contact with mollusks. Among these ascidians a fine thread of cartilage appears, which has about the same position as the "chorda" of Amphioxus. Most of them have this cartilage only in the embryonic or larval stage, but a few of them preserve it for life. There is a strong possibility that the ascid- 113 THE EVOLUTION OF MAN ians are very closely related to the vertebrates. It is true that on the one side they are buried deeply in the worm type far below Amphioxus. But on the other side they have the chorda, the first trace of a genuine backbone. But since some of them show this chorda only in the embryo stage, it seems evident that their ancestors had a still stronger hold on this rudiment, and were therefore still closer to the vertebrates than most of the present ascidians, which have evidently somewhat degenerated in this respect. So that Amphioxus and ascidians would be two branches of the common archetype which would, first of all, have developed the chorda. This archetype in order to produce the present-day ascidians must have been in all other respects unmistakably a wormlike animal. In short,jwej2just jpok for other traces of man — in worms. The term "worm" applies in the system to an enormous mass of different animals. There are hundreds of groups of fundamentally different worms. Some of them are of a higher order, with blood and sense organs and a genuine cen- tral nerve system. We would have to derive vertebrates most likely from them. If so, we should imagine a worm, which would not pos- sess a chorda like Amphioxus or the lamprey, but 114 THE EVOLUTION OF MAN would at least have a nerve string, which could later on develop into the spinal cord of a fish, and below which the digestive tract would extend in the form of a hose with one opening at each end of the body. The entire form would have no fin-legs, but would be a typical worm. This . is the outline to which most of the present higher worms actually correspond. At the same time we now find lower groups of worms which evidently belong further back in the scale. They have no complex nerve appara- tus, no blood system and no opening at the lower end of the body. We are justified in assuming that they represent an older type, a sub-stage of the worm type. In other words, within the worm family we should have to look for man in various disguises leading from the complex to the sim- ple. There is still something else to consider. In our system, apart from the vertebrates, there are still three other great groups of invertebrates which are of a higher organization than the worms. They are, first, the crustaceans' spiders and insects; then, the mollusks, such as snails, muscles and octopus and finally the echmoderms, ** such as star-fish, sea-urchins and related forms. Not even the most daring anatomical speculation can accomplish the miracle of deriving any one THE EVOLUTION OF MAN of these three classes from the other, and it is still less feasible to fit the vertebrates into any one of them. It would be impossible to develop an Amphioxus from a star-fish or an octopus. Some have attempted a theoretical line of descent from crustaceans to fish, but only by means of such a yawning chasm that no rational investi- gator went with them. The difference between these things is too great. On the other hand, it is remarkable that all those groups may easily be traced back, each by itself, to some higher worm. It is true that the worm type to which the line of crustaceans and insects attaches itself and to which, for instance, our leeches and earth worms belong, is very dif- ferent from an ascidian. Evidently there has been a great deal of individual evolution within the higher worm type. But nevertheless this picture presents a great deal of probability. The higher type of worm branched out into insects, mollusks, echinoderms and vertebrates, and it had four possibilities of evolution, among which only the vertebrate was destined to win the crown — the form of man. But this entire stage of worm life proceeded from some still lower worm which would therefore represent the next common sta- tion of all worms, and with them all men. 116 THE EVOLUTION OF MAN If we now try to get a conception of the worm in its lowest stage, we will find that its structure is wonderfully simple. Imagine for a moment that one limb after the other, one organ after another, is cut away from a man, arms and legs, the head, the spinal cord, the blood system, all the parts and organs between the stomach and skin, and that nothing remains finally but this skin and a stomach fitting it closely. Furthermore, let the"! (rectum be closed, which gave to the higher or- | Jganized worm the form of a hose, and only one i ( opening remains for both assimilation and excre-l tion. Now such animals actually exist on the lowest plane of worm life. In certain jelly-fish, there lives a little parasite named Pemmatodiscus, which is literally composed of nothing but skin and digestive tract. There is also another animal in our 'fresh waters which is a little above this stage, the so-called Hydra. In the case of the Hydra, its lower end has grown fast to the soil, while its mouth is surrounded by fine tentacles, and a few other details are slightly better de- veloped. Might it be possible that we could fol- low man down to this stage ? We sometimes speak of a man who consists only of "skin and bones." Well, that would still 117 THE EVOLUTION OF MAN be a vertebrate man. But now, we are asked to eliminate also the backbone. Man is to consist only of skin and stomach. These two organs are now supposed to contain the germs of everything which is later on developed to full bloom in the human body, such as the nerve system, the blood system, the nutritive system, the sexual system, etc. This idea has seemed rather daring to some people, in spite of the fact that a mere consid- eration of the zoological system suggests it. There we finally likewise arrive at the Hydra by going from complex to ever simpler forms. There is no escape from this logic, if we venture at all into this vague domain of circumstantial evidence. If we do this, the logical line of re- search is the one we have followed. But there is still another one, and it is peculiar that it leads to exactly the same result. We have not mentioned the embryo for some time, but now we shall ask it once more to act as a witness. It is an anatomical possibility which, like all such extreme speculations, is mainly proven by its plausibility that another group of genuine in- vertebrates may be derived from such skin-and- stomach creatures as the Hydra, without passing through that other line of development in the 118 THE EVOLUTION OF MAN evolution of worms. I speak of sponges, higher polypi and jelly-fish. These would form a definite circle, and all higher forms above the skin-and- stomach animals of the lowest class would be traceable to one common origin, viz., this skin- and-stomach animal itself. But now we recall that law which so frequently reproduces the por- trait of ancestors in the formation of the embryo. If the circumstantial evidence is to be conclusive, then the embryonic development of all animals from the jelly-fish to the vertebrate ought to re- produce such a portrait, representing the double cylinder of skin and stomach with one single orifice, a prototype of Pemmatodiscus, or Hydra. Here again all resistance is useless. It is unde- niable that such embryonic marks reappear in all nooks and corners of the higher classes of ani- mals. Itjs that gfcge whfct* %rwi ha* Hp