FLORA MALESIANA
SERIES I - SPERMATOPHYTA
Flowering Plants Vol. 6, part 6
INDEX TO REVISED FAMILIES
Aceracede,: +. 4: 3,592 Elatinaceae 4: 203 Pentaphragmataceae. 4: 517 Actinidiaceae s.str. . 4: 37 lEpacridaceae . 6: 422 Pentaphylacaceae. 53-121 Aizoaceae ees ia 207n subricaceae. : 6: 469 Philydraceae 4: 5 Alismataceae . . . 5:317 + #Erythroxylaceae 5: 543 Phytolaccaceae 4: 229 Amaranthaceae . 4: 69,593 Ficoidaceae 4: 267 __Pittosporaceae 5: 345 Ancistrocladaceae . 4: 8 _ Flacourtiaceae 5: 1 Plumbaginaceae 4: 107 Aponogetonaceae 4: 11, 7: 213 Flagellariaceae 4: 245 Podostemaceae te G5 Basellaceae - 5:300 Geraniaceae 6: 445, Polemoniaceae . . 4: 195 Batidaceae 5: 414 Gnetaceae. 4: 336 Pontederiaceae aS oS Betulaceae 5: 207 Gonystylaceae 4: 349 Portulacaceae TWA Bixaceae s. str. 4: 239 Goodeniaceae 5: 335 Primulacea 6: 173 Burmanniaceae . 4: 13,592 Haemodoraceae 5: 111 Proteaceae 5: 147 Burseraceae 5: 209 Haloragaceae 7: 239 Punicaceae 4: 226 Butomaceae 5: 118 Hamamelidaceae . 5: 363 Restionaceae . 5: 416 Byblidaceae 7: 135 Hippocrateaceae . 6: 389 Rhizophoraceae i TS ae Callitrichaceae 4: 251 Hydrocaryaceae 4: 43 - Salicaceac,. (..°) Sieceesemn Campanulaceae 6: 107 Hydrocharitaceae. 5: 381 Salvadoraceae. «.) eaaaees Cannabinaceae 4: 223 Hydrophyllaceae . 4: 207 Sarcospermaceae. . 4: 32 Capparidaceae 6: 61 Icacinaceae 7: 21 Saururaceae :..’ sean Caprifoliaceae . 4: 175,598 Juglandaceae . 6: 143 Scyphostegiaceae. . 5: 297 Cardiopteridaceae 7: 93 Juncaceae . 4:210 Simaroubaceae . . 6: 193 Celastraceae . . 6: 227, 389 #Juncaginaceae. 4: 57 Sonneratiaceae . 4: 280, 513 Centrolepidaceae §: 421 Lemnaceae 7: 219 Sparganiaceae <) (eases Ceratophyllaceae 4: 41 Loganiaceae 6: 293°. Sphenocleaceae’ | eae, Chenopodiaceae . 4: 99,594 Lophopyxidaceae 7: 89 Stackhousiaceae < )-yAewa Clethraceae 7: 139 Malpighiaceae 5: 125 Staphyleaceae « “EL Geeeas Cochlospermaceae 4: 61 Martyniaceae . 4: 216, Stylidiaceae’.. 2 ieee Combretaceae 4: 533 Molluginaceae 4: 267 | Styracacede . -. -eaaaeeas Connaraceae . . . 5:495 Moringaceae . 4: 45 Thymelaeaceae 4: 349, 6: 1 Convolvulaceae . 4: 338,599 Myoporaceae . 4: 265. -Trigonidceae.. 4. eee Corynocarpaceae 4: 262 Myricaceae 4: 277 -’Turneraceae .° 3 Seer Crassulaceae . 4: 197 Najadaceae 6: 157 Typhaceae . eA ae SSSA Datiscaceae 4: 382 Nyctaginaceae : 62450 # Umbelliferae . ~ 41133595 Dichapetalaceae . 5: 305... Nyssaceae.... . |... . y42029- ‘Walerianaceae’:) 2 aeeenaeecee Dilleniaceae 4: 141 Ochnaceae 1) Meee Pa eba Oat AVG laceae ids Sen coe Dioscoreaceae 4° 293 « Oxalidaceae wo” Sm feasts 151 Xyridaceae . . 4: 366, 598 Dipsacaceae 4: 290 Papaveraceae. . . 5:114 # Zygophyllaceae, . . 4: 64 Droseraceae A: 37) -Pedaliaceae 2 0-0, ee 42 216
TAXONOMICAL REVISIONS
REPUBLIK INDONESIA REPUBLIC OF INDONESIA LEMBAGA ILMU PENGETAHUAN INDONESIA (L.LP.L.) / INDONESIAN INSTITUTE OF SCIENCES
FLORA MALESIANA
BEING AN ILLUSTRATED SYSTEMATIC ACCOUNT OF THE MALESIAN FLORA | INCLUDING KEYS FORDETERMINATION | DIAGNOSTIC DESCRIPTIONS | REFERENCES TO THE LITERATURE|SYNONYMY]/AND DISTRIBUTION | AND NOTES ON THE 'ECOLOGY OF ITS WILD AND COMMONLY CULTIVATED PLANTS
PUBLISHED UNDER THE AUSPICES OF LEMBAGA BIOLOGI NASIONAL BOTANIC GARDENS OF INDONESIA / BOGOR / JAVA AND OF THE RIJSKSHERBARIUM / LEYDEN / NETHERLANDS
PREPARED ON AN INTERNATIONAL CO-OPERATIVE BASIS UNDER THE SUPERVISION OF SEVERAL DIRECTORS OF BOTANIC GARDENS / KEEPERS OF HERBARIA AND VARIOUS PROMINENT BOTANISTS
FOR: THE PROMOTION. OF BOTANICAL SCIENCE AND THE CULTURAL ADVANCEMENT OF THE PEOPLES OF SOUTH-EASTERN ASIA TO THE SOUTHWEST PACIFIC REGION
SERIES I
VOLUME 6 SPERMATOPHYTA
GENERAL EDITOR:
Dr CrG. G. IVAN STEENIS DIRECTOR OF THE FOUNDATION ‘FLORA MALESIANA’
PUBLISHED BY WOLTERS-NOORDHOFF PUBLISHING, GRONINGEN, THE NETHERLANDS PRINTED IN THE NETHERLANDS 1960-1972
COPYRIGHT 1972
All rights reserved, including the right to reproduce this book or parts thereof in any form
ISBN 9001 31822 3
Printed in the Netherlands
Title-page Contents . Dedication
Abbreviations and signs.
CONTENTS
TAXONOMICAL REVISIONS
in alphabetical sequence
Campoanulaceae by B. Moeliono & P. Tuyn .
Capparidaceae by M. Jacobs . Celastraceae-I by Ding Hou . Celastraceae-II by Ding Hou . Epacridaceae by H. Sleumer . Ericaceae by H. Sleumer
Geraniaceae by R. C. Carolin. Juglandaceae by M. Jacobs Loganiaceae by P. W. Leenhouts . Najadaceae by W. J. J. O. de Wilde . Nyctaginaceae by J. F. Stemmerik Primulaceae by P. A. J. Bentvelzen Simaroubaceae by H. P. Nooteboom.
Staphyleaceae by B. L. van der Linden .
Thymelaeaceae by Ding Hou .
Addenda, corrigenda, et emendanda by C. G. G. J. van Steenis and collaborators .
ADDENDA to volumes 4, 5 and 6
INDEX
Index to scientific plant names by M. J. van Steenis-Kruseman
Page (3) (5) (7)
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915
985
Dedicated to the memory of ELMER DREW MERRILL
DEDICATION
The completion of the sixth volume of this Flora gives me the privilege to dedicate this to the memory of ELMER DREW MERRILL, a man who has achieved more for the knowledge of the Malesian flora than any other individual botanist.
It is neither my intention to give nor is it the proper place for a full biography of this most distinguished American scientist, as it would for the greater part be duplication of his own ‘Autobiographical’ (1953), the scholarly essay by RossBins (1958), and the vivid life sketch by ScCHULTES (1957), which together give the story of his life, his ambitions, his personality, his immense drive, his multiple interests, his capacity for establishing botanical periodicals as well as successfully filling the posts of Dean of a Faculty of Agriculture, director of the Bureau of Science at Manila, director of the New York Botanical Gardens, and administrator of Botanical Collections of Harvard University.
It is my purpose to review MERRILL’s aims and vision, ambitions and achievements in the light of his time, to explain the value of his pioneer works for Indo-Malesian botany, how he used opportunities and had to bow to unforeseen events and circumstances which in no mean way influenced his career. Naturally MERRILL’s personality pervades the story, that of a straight- forward, righteous person, unbiassed in scientific matters, appreciating any progress in bio- logical science. It is of course especially his great achievements with regard to the knowledge of the Malesian flora which are the main theme and I will try to elucidate several aspects which he pursued.
A glance through his immense bibliography, containing some 550 entries, among which ten very large books, reveals his fantastic productivity, largely centered on the flora of Malesia, East Asia and the Pacific.
Scanning the herbarium one becomes aware of the fact that during his lifetime he must have pre-identified, named or definitely identified over half a million specimens from the East, including the Philippines, Sumatra, Borneo, Amboyna, New Guinea, Melanesia, Micronesia, southern China (including Hainan), Indo-China and Burma, made possible by his unequalled knowledge of forms, his cast-iron memory and his zealous devotion.
It is with awe that one observes such a great achievement and then one wonders how one man could find time to do all this. The answer is given by SCHULTES who wrote that ‘his reaction to added work was to lengthen the day’.
As Rossins wrote, ‘he was a man in a hurry who saw clearly a program of research which absorbed him, but which was greater than any individual could complete within the limits of a single life-time. The demands of this program possessed him. It led him to make various in- novations and modifications of convential procedures in herbarium methods with increased efficiency in the use of this, for him, essential tool and to advocate briefer citations, one-name periodicals, and other means of economizing time and effort. It induced him, at least in part, to make quick decisions without long considerations of pros and cons and to act at times without regard to the feelings of others.’
‘In spite of his absorption with his speciality MERRILL was not a recluse. Nothing pleased him more than to light his pipe and sit down to talk with a group of gardeners or with a student or his colleagues, mainly, of course, about plants. He played an active part in many organizations and valued the associations he made in them. He joined the Masonic order and eventually became a thirty-third degree Mason. He enjoyed an evening cocktail in his later years, and loved to have visitors and dinner guests. Though he had no hobbies outside his profession, he was interested in sports, especially baseball, football, and tennis. The ‘Autobiographical’ accounts of his adventures in collecting in the Philippines and in China reveal some of the human aspects of his character.’
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Such human aspects became evident to us very shortly after the war as he sent food parcels to his German colleagues and distributed amongst other botanical material on loan or for gift to colleagues in other freed countries envelopes with pounds of tobacco labelled with dry humour Nicotiana tabacum L. in parcels marked as ‘botanical specimens for scientific study only’.
‘His energy was boundless. It was his habit, at the University of California, to arrive at the herbarium at 6:00 a.m. and work on shipments of plants until he left for the Dean’s office at 9:00 a.m.; he continued at noon after office hours, or on sundays and holidays’ (RosBINs, 1958).
As will appear later, his drive was probably born from the challenge he had to face in the Philippines in his early days and which activated his innate energy: it became a life-long habit.
MERRILL was born in East Auburn, Maine, October 15, 1876. His parents were of limited means and belonged to the industrious people of New England. One of his ancestors was NATHANIEL MERRILL who immigrated in 1635 and was of Huguenot descent (originally DE MERLE). In his family there was also English and Scottish blood, a ‘melting pot’ feature not unusual in the United States. As a young boy he was interested in natural history, in birds, rocks, minerals, fungi, and local woods; before reaching highschool age he became interested in col- lecting and naming plants, but he went in for engineering in 1894 in Maine State College at Orono which in his senior year became the University of Maine. He took the general science course, but remained interested in biological work, particularly in the classification of plants and worked under Prof. F. L. HARVeEy especially on cryptogams; he attended, however, only two semesters on botany. His private herbarium contained some 2000 specimens. In 1898 he got his B.Sc. and became assistant in natural history. While working for his M.Sc. he accepted a post as assistant-agrostologist in the U.S. Department of Agriculture, Washington, as assistant to F. LAMSON-SCRIBNER, then the leading authority on North American grasses, composing amongst others a Manual of the Grasses of Alaska.
In 1902, following the establishment of the sovereignty of the United States over the Phil- ippines, his chief accepted a post as director of the newly established Department of Agriculture in Manila. LAMSON-SCRIBNER was Obviously so very satisfied with MERRILL’s person and capacity that he asked him to become botanist in his department. MERRILL, however, was completely satisfied with being an agrostologist in his own country and refused twice, arguing that he knew nothing about tropical plants from the East. Finally ScRIBNER convinced him that ‘nobody in the United States knew anything about the Philippine flora and that he had as good a chance as any one’. He accepted on Monday afternoon, February 20, 1902, and had to agree to be ready to sail from New York at 1:00 p.m. February 22! He nearly missed the boat because of a heavy sleet storm delaying the train from Washington D.C. to New York, but he made it. He, least of all, could not anticipate that he would remain for twenty-two years in the Philippines. Slightly over two months later he reached Manila where the new personnel had to start work ina vacant building, ‘without a chair or table, much less a botanical publication or a botanical specimen’.
With a huge program before him MERRILL started energetically: one month after arrival he made a six weeks trip, partly under military guard, and during his first years he spent approxi- mately one half of his time in the field; in the next twenty-two years he explored in almost all parts of the extensive archipelago.
In September 1902 he paid a two-months visit to the Herbarium Bogoriense, taking with him his botanical specimens, for no trustworthy work could be done in Manila in the complete absence of authentically named material. Here he became acquainted with the literature of Malesia of which he acquired an excellent working knowledge. He himself said that it was of infinite value for building up the botanical library at Manila. He wrote also an extensive report
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Dedication
on this visit explaining the methodologies of collecting, forest plotting, numbering of trees in forest reserves, herbarium methods and techniques, efc. used in Java, which served as a model for the work in the Philippines.
At that time the responsibility for taxonomical research work in the whole of Malesia vir- tually hinged on two systematists, RIDLEY in Malaya and VALETON at Bogor who collaborated with Koorpers, the organiser of the collecting; to these MERRILL was now added as a third.
In comparison with Malaya and Java, work in the Philippines had the great disadvantage of having a much less solid basis; this consisted only of BLANCO’s, FERNANDEZ-VILLAR’s and VIDAL’s early works, and at Manila there were no voucher specimens of these authors’ collec- tions, nor of the vital collections of CUMING and HAENKE for comparison.
This vacuum led him to realize that he had to start from scratch and had to build up a col- lection and library, to interpret plants from older works, enabling the description of new plants and accounts of collections, that it would be compulsory to ask assistance from foreign botanical specialists in various groups, that he had to stimulate interest in the botany of the Philippines by distributing duplicates on a large scale and by bringing Philippine plants into ‘circulation’, and that he had to bring into being a publication medium for scientific results and for the correlation of Philippine botany with that of adjacent countries.
This clear vision of the items of the enormous amount of pioneer work ahead would have discouraged any average scientist, but to MERRILL it meant a great challenge, to create a centre from scratch.
It became almost a one man show. In the course of twenty years he had never a permanent collaborator, except two honorary collaborators, viz E. B. COPELAND for the Pteridophytes and Oakes Ames for the Orchidaceae. Another honorary American collaborator, Miss JANET PER- KINS, started a series of publications under the name ‘Fragmenta Florae Philippinae’; she settled at Berlin where she worked in conjunction with OTro WARBURG who had made large Philippine collections himself. These collections, together with those of MERRILL and Capt. AHERN formed her basic material, but after 3 fascicles (1904-1905) this promising series was abandoned. Another American botanist who later joined the Bureau of Science at Manila and was employed for three years (1908-1911) was CHARLES BUDD ROBINSON, a critical, promising botanist. He returned in 1912 for another period, but unfortunately met a premature death in 1913 on the island of Amboyna while re-collecting Rumphian plants.
MERRILL of course worked in close collaboration with the members of the staff of the Forestry Institute. They were mostly collectors describing forest composition and timbers, such as H. N. WHITFORD, H. M. CurrAn, and others. An exception was F. W. Foxwortuy who actually served for some years as botanist at the Bureau of Science and who took care of a revision of Philippine Dipterocarpaceae, the most important timber family.
An American botanist of great impact on the development of Philippine botany was ADOLPH DANIEL EDWARD ELMER (1870-1942). He was originally in the employ of the Bureau of Science in Manila (1904-1905), but settled later as a professional free-lance plant collector and collected over 20.000 numbers in the Philippines in a large number of sets. ELMER published lavishly in a series of his own, the ‘Leaflets of Philippine Botany’, ten volumes, together covering 3936 pages in print, among which descriptions of over 1500 new species. What MERRILL’s relation to ELMER was and whether he appreciated him or not are not clear, as he does not mention him in his ‘Autobiographical’. At the time of his stay in Leyden, where I had the privilege to have him as our guest, Oct-Nov. 1951, I omitted to unearth this. I can hardly believe that ELMER’s super- ficial uncorrelated descriptions can have been very welcome. In Ficus MERRILL reduced 31 of the 70 described species by ELMER before 1923 and of the total of 85 novelties of ELMER’s Ficus only
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FLORA MALESIANA
13 are still accepted in CORNER’s monograph. MERRILL had to tolerate ELMER’s work because the latter was an entirely independent man whose virtue lies mainly in his profuse collecting.
Why MerriLL never had a larger permanent staff of at least 2-3 qualified botanists is another question left unanswered. He could easily have claimed staff for the Phanerogams as besides his collecting work, he had curatorial and organisational tasks. Since 1912 he also had an associate professorship in the University of the Philippines and served as head of the Department on a half-time basis and without additional compensation (sic), “which until 1918 seriously interfered with productive work in systematical botany’. To make botany popular and to frame a textbook of systematic botany for both residents and students, he had already prepared a ‘Flora of Manila’ (1912), still a very useful book, covering some thousand species. During the academic year his teaching duties never occupied less than eighteen hours per week, and during certain semesters even thirty-six. It is amusing to read in his ‘Autobiographical’ how he got rid of it, early in 1919, ‘his full time being for the first time in many years available for what he most desired to do’. But after two weeks he was appointed as director of the Bureau of Science, a post neither sollicited nor desired, but which he could not refuse under the circumstances, it being an order.
In passing I remark that of the Filipino students he taught not one specialized in plant system- atics as far as I know. Those with a biological tendency probably all went in for more applied branches, such as pharmacy, entomology, agriculture, forestry, fisheries, etc., so that when MER- RILL left the Bureau of Science in 1923 he had no immediate successor as botanist. It was five years later when Dr. E. QUISUMBING was engaged as such. He had received his primary education at the College of Agriculture, University of the Philippines and at Chicago University. He worked together with MERRILL, then Dean of the Faculty of Agriculture, University of California at Berkeley, for two years, 1926-1928, from which a joint paper “New Philippine Plants’ emanated; from this it appears also that MERRILL was quite capable of making botanical drawings. The main thing was that MERRILL induced him in these two years to start revisional work for the initiation of a ‘Flora of the Philippines’ and along this line QuUISUMBING elaborated the first sample, Philippine Piperaceae, while at Berkeley, under MERRILL’s supervision, as a final coach- ing and a start towards this new goal.
Notwithstanding the time-consuming handicaps connected with education and administration MERRILL poured all his available energy into the botanical aims set forth above. He had by necessity been induced to take on administrative duties in the Philippines and had shown his capacity to meet this challenge. This experience served him later enabling him to fulfill other administrative duties in California, New York and Harvard in a successful way. That he could pursue his own botanical work simultaneously in free time and leisure hours is due to the fact that his heart was in botany and that he regarded work in plant systematics and floristics as his ‘safety valve’ ‘when he could immediately forget his administrative problems’. The taking of vacations was rather foreign to him and in the entire period in the Philippines, from 1902 to 1923, he only took vacation in 1905, while in other years he used his annual month’s leave for making collecting trips, mainly in China.
He built up a collection, personally, with various famous skilled Filipino collectors (RAMos, EDANO, SULIT and many others) from all parts of the Philippines (the Bureau of Science = B.S. series), acquiring huge collections from private collectors (ELMER, Mrs. CLEMENS, WENZEL, LOHER, VANOVERBERGH, efc.) and from the Forestry Service officers (the F.B. series). Besides this he acquired large collections from surrounding regions, Guam, China, Indo-China, Amboy- na, and in addition very large sets of duplicates in exchange for the material which he widely distributed ‘on a free exchange basis’, that is, the liberal but in the long run profitable principle of sending out duplicates as many as each institution has available, thus not on a precise | : 1
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Dedication
specimen basis. He estimated that in 1922 he had sent out more than 500.000 duplicates, the reference collection at Manila amounting to c. 275.000 mounted specimens, containing repre- sentatives of practically all Philippine species, in the form of types, isotypes, fragments of types, material critically compared with originals, photographs of types and sketches.
The library, similarly built up from scratch, was after a decade admittedly ranked on the same level with those at Bogor and Calcutta and far larger than the one at Singapore.
He had also to provide for a publication medium and was instrumental in founding the Philippine Journal of Science, Botany Supplement, which was filled mostly by his own contri- butions but also served for revisions prepared by his associates, Philippine and foreign.
Interpreting the older works he found a great necessity, both for botanical and nomenclatural reasons. He started with the work of BLANco, for which purpose he had special collections made in Blanco’s classical areas, from which emanated his ‘Species Blancoanae’ (1918), followed by a similar key work on ‘Rumphius’ Herbarium Amboinense’ (1917), both unrivalled for their critical standard. He tried to correlate both interpretations with material from the locus classicus. Only first-class botanists with a very wide knowledge of plants are capable of composing such works. In 1921 he published an evaluation of BURMAN’s ‘Flora Indica’, but this was not sustained by a study of the types at Geneva; unfortunately BURMAN’s herbarium is dispersed through the general Herbarium by HocHREUTINER. For his “Commentary on Loureiro’s Flora Cochinchi- nensis’ (1935) he had at his disposal earlier collections made by Mrs. CLEMENs in the locus classicus near Hué; he also tried to locate types of LouUREIRO in the Herbarium of the British Museum, London. At the end of his life he was working on a very large work on the location and evaluation of the RoxBURGH plants, the MSS for which are left unfinished (a complete copy has been deposited in the Arnold Arboretum, a less complete one at Leyden). We would have preferred that he had started this work earlier instead of his immense evaluation of the works of RAFINESQUE which occupied him for several years onwards of 1940. Admittedly the location and evaluation of the ROXBURGH types could only be performed by prolonged visits to European herbaria, which were impossible during the war and which, moreover, MERRILL could not make while he was Administrator of the Harvard Collections. All these works are extremely important for typification and nomenclature, often of very common tropical plants as many errors or omissions were made in their former interpretation, if indeed any was ever made.
MERRILL contributed lavishly to describing new species — some 4000, of which 3000 from the Philippines — and several new genera of plants. They were partly published in a series of New and Noteworthy Philippine Plants (18 numbers, 1904-1922), partly in accounts of particular collections from certain islands, districts or mountains. It is quite certain that only a fraction of these will stand the test of time. To understand this we must consider his vacuum position and the dilemma with which he was faced, either to do critical-botanical work or to do the best he could in determining plants by reading descriptions, comparisons with available material and if nothing fitted to describe the plant as new. Critical-botanical work of course goes slowly, at a rate of some 20-100 accepted species a year depending on the group and the botanist’s capacities and zeal; it also requires that one has access to a large library and can borrow types and authentic material. The absence of the latter facility is the greatest handicap for monographic work in the tropics, as the largely European-centred ‘type herbaria’ are not prepared, and rightly so, to make large loans to the tropics, an affair which is too risky in several respects. This necessitates that workers in the tropics, after having prepared preliminary MSS, must pay prolonged visits to these institutions for checking type and authentic material and establish identities and names. And although MERRILL gradually assembled at Manila a huge herbarium and photographs or fragments of types from European herbaria, and attached figures, descriptions, notes, pencil
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FLORA MALESIANA
sketches, carbon leaf impressions, etc. to the sheets, framing in this way a new tool of informa- tion, a sort of combined plant file and library, he had little of this during the first decade of his work at Manila. At that time he had to be content to study descriptions, an art which he fully mastered, an art which is gradually becoming extinct among contemporary botanists who adhere to the examination of type specimens. Moreover, the incoming Philippine material was over- whelming, all completely unnamed. To cope with this he asked the co-operation of all available specialists in the world to elaborate certain groups. The bulk he had to name himself and this could be done only in a superficial way, thus it amounted to mere floristic name-giving and was not truly systematic, comparative, either regional or semi-monographic. Besides all the already mentioned handicaps to work of lasting value, the Philippine flora offers an additional com- plication in its specific variability, which is in part certainly a consequence of the archipelagic conditions due to fragmentation of once continuous populations before the block-sinking disrupted these in past geologic time. With scanty material a botanist will under such circum- stances be inclined to describe more species than there really are. It must be admitted, however, that the specialists to whom he entrusted the work on as many groups as possible were faced by exactly the same difficulty and their work suffers from the same evil. By working on this level MERRILL certainly created problems for others to solve. I have not scanned families on that point, but the six sedges he described as new are now all in the synonymy.
He himself was of course fully aware that there were far less species in the Philippine flora than listed at the time of his enumeration (1923), and being honest and wise to a high degree, he frankly admitted during a discussion on the virtue of different policies for unravelling the botan- ical treasures of Malesia, that ‘many of my new species, and even new genera, were optimistically proposed’, adding that he had seen no other way to create a botanical basis.
Of course nowadays, half a century later, ‘collection description’ is an obsolete procedure, detested as an inferior sort of botany creating unnecessary duplication and spending valuable time in a useless way. Nowadays it is clear that thorough work is needed and this can be done because in general collections have accumulated sufficiently to provide a solid basis, which we hope is about as good as, or slightly better, than the basis collections which were used for the compilation of the ‘Flora of British India’. Fortunately collections are becoming more ample every year due to enthousiasts in Malaya, Sarawak, Sabah, the Philippines, and Lae, to whom we cannot be thankful enough. In MERRILL’s ‘period’ the time was simply not ripe for launching comprehensive botanical works. To have a fair judgement of his pioneer work the commentary just given should be considered.
Several facets of his work were intended to cope with the urgencies of the ‘vacuum position’, e.g. the building up of collections, the library, a publication medium, the nomenclatural evalua- tion of old works, the description of supposed novelties, the framing of enumerations, as pre- cursors to Floras, and bibliographies. Having a vast knowledge of the literature he was in the latter extremely successful, in accuracy and completeness. His enumerations of the floras of Borneo, Hainan, Banguey I., Guam, and some others are basic for later work, especially that of Borneo.
After the passing of Taft’s Law (1917) it was American policy to prepare the Philippines for independence and MERRILL foresaw that his future career would not allow him to compile his ultimate goal, a ‘Flora of the Philippines’ for which his ‘Flora of Manila’ had served as a model — actually in 1922 although he was director of the Bureau of Science he was on a year-contract, without pension and could be dismissed with a year’s salary as bonus (sic) — so he prepared his ‘Enumeration of Philippine Flowering Plants’, to synthesize what was achieved and leave a basis. This is still a most useful work, without which we would be much the poorer. The introductory
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Dedication
essays are a masterpiece of work, the enumeration is complete, and the bibliography is still a reliable source of information. Of equal value are his bibliographies of Borneo, the Pacific, and East Asia.
MERRILL’s ability of mastering and analyzing literature entailed of course changes in name- giving according to the principle of priority and typification. As a good botanist of course he favoured stability of nomenclature, which cannot always be realized, however, particularly not at species level. He listed especially, in his works on LouREIRO and RAFINESQUE, the cases where conservation of generic names was necessary.
A corollary of his bibliographic interest was establishing new journals; he was instrumental in founding the ‘Philippine Journal of Science’ and many others, later pointing out the virtue of one-name periodicals of which he founded several, a procedure later widely accepted (Blumea, Willdenowia, Persoonia, Adansonia, e/c.).
Shortly after the war he also started a promising offset reprinting of rare early works, the desirability of which probably emanated from his study of the rare works of RAFINESQUE. He told me that with a subscription of 150 copies this could be done without loss at a very low cost on a non-profit basis. It is a pity that this was not continued by somebody for the benefit of botany, as there was and is a need for such inexpensive reprints. Before the war I had already tried to raise the interest of Chronica Botanica for such a purpose and later made similar pro- posals to the I.A.P.T., but found no sympathy for the idea. The result is that large firms nowadays produce such offset prints at prohibitive prices on a profit basis, a most undesirable situation for botany in general and the Philippines especially as the reprints of the ‘Flora Manila’ and the ‘Enumeration’ fall entirely beyond the financial capacity of most Philippine residents for whom they are intended. That a low-price large sale is still possible is apparent from the excellent offset reprinting in Formosa for educational purposes.
Another corollary which proved to be of great scientific interest was his careful recording of vernacular plant names, an intricate affair because of the many dozens of native languages in the Philippines. He examined these names together with linguists and found that several of them were corrupted Spanish names; thus he contributed to both linguistic and botanic science. He found also that many early plant introductions had never spread and still retained station in the coastal hinterland of Cagayan Bay where the Spanish galleons had anchored. This, combined with the vernacular names, brought him to a consideration of early post-Columbian transtropical transport of cultivated plants and aliens. In this, he found that a clear distinction must be made between the galleon routes of the Portuguese from Brazil to Goa and Macao and that of the Spanish from Acapulco in West Mexico to Manila via Guam. Both galleon routes proved very important indeed, as they persisted for centuries. In this way he was able to unearth the origin of a number of plants on which there was no unanimity of opinion, including such important crops as maize, and of sweet potato and tobacco in the New Guinea highlands. Later he was able to check this in the herbarium by studying the lists of the early BANKs & SOLANDER col- lections made during the first of Cook’s voyages in the Pacific. On this subject he published a book which, notwithstanding acid criticisms on ‘certain diffusionists’, is a magnificent source on the subject of ethnobotany affecting all tropical botany. In passing I remark that a similar study should still be made on the ComMMeErRSON collections, as a second proof for his theory.
I do not know of course what MERRILL’s reaction would have been to a dedication of a volume of our Flora to him. In comparison with the great many honours fallen to him, honorary docto- rates and honorary memberships, memberships of academies of science, the gold medal of the French Ministry of Agriculture, the gold medal of the Linnean Society of London, the Geoffrey St. Hilaire medal, Officer of the Netherlands Order of Oranje Nassau, and so forth, our dedication
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FLORA MALESIANA
is only a very small one. Still, I feel certain that it would have ranked very high with him, because it lies precisely in the field of botany in which he spent probably the best years of his life, and which intrigued him most of all.
He had never any intention to join in our work by personal contributions. Apart from the fact that the Flora started when he was already well in his seventies, he had used his capacities primarily in making tools for botanical workers, such as his critical evaluations of basic early works (BLANCO, RUMPHIUS, BURMAN, LOuREIRO, and in his last years ROXBURGH), enumerations of the flora of local areas (the Philippines, Borneo, Hainan, Guam, Banguey I.), the elaboration of large single collections from various parts of the East (Burma, Sumatra, NE. Borneo, New Guinea, together with Miss Dr. L. M. Perry) and in compiling valuable bibliographies (the Pacific, East Asia, the Philippines, Borneo). Through his immense knowledge of plants and books he could produce such works at a speed at which normally large teams of workers would have been necessary.
By necessity there is a certain restriction of purpose and objects in making such tools, several were also called ‘bibliographical enumerations’, indicating that they were not critical and that he was not responsible for the correctness of the names. To expect or request this is of course completely unfair and unjust; they were intended as tools, and this purpose was accomplished.
There was a similar restriction in his systematical work, as he confined his revisions mostly to limited areas, many in the Philippines (Syzygium, grasses, Leguminosae, etc.), Borneo or New Guinea. Nevertheless it was an immense achievement, but at a certain level, as good as it could be under the circumstances, but largely floristic and not critical. He did not contribute essays on theoretical systematics.
Quoting RosBIns, ‘it has been said that MERRILL seldom went far below the surface, and that he was content in most cases to classify the plants with which he dealt’. Ropsins defended MERRILL in commenting: ‘This type of research was proper for the region he studied and was the only procedure which permitted him to do what he did in his lifetime. It is exactly this characteristic ability to deal superficially with extraordinarily large numbers of plants that makes so apt the epithet ‘the American Linnaeus’ which has been applied to him. Actually, however, MERRILL did go below the surface. He recognized that classification was prerequisite for other investigations and the magnitude of the job he set for himself left little time to pursue anything else. But his studies of the floristic and faunistic relationships of the Philippines to other Malesian areas, of the significance of vernacular plant names, and of the origin of cultivated plants are examples of ‘below the surface’ investigations carried out by MERRILL.’ I may add his excellent synthesis of the distribution of the Dipterocarpaceae which induced him to make a rough outline of the phytogeography of Malesia, and especially its relation to the Formosan flora, in the light of correlating biological distribution with geological history.
To be fair, we should always keep in mind that during the period in which he achieved his great contributions to Malesian botany, that is 1902-1923, he started without a predecessor from absolute scratch, without personnel, without a book or collection, in an almost unexplored very rich archipelago covered largely by primary forest. Later too he had to work under scientific vacuum conditions in that he had almost no colleagues around him in Manila, there were hardly any botanists in the whole of Malesia — except for VALETON and J. J. SmirH in Bogor and RIDLEY in Singapore — while the floras of the islands surrounding the Philippines (Borneo, Ce- lebes, the Moluccas and New Guinea) were botanically only known in the most fragmentary way.
If he had aimed at a critical systematical study of the Philippine flora in 1902, he would have had to wait for half a century for publication and could never have composed the major tools which now belong to our standard bibliographical equipment.
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Dedication
Even in the Netherlands Indies, which had an infinitely better botanical basis, TREUB had at the turn of the century concluded that a composition of a comprehensive Flora would be entirely premature because of the primitive state of exploration and publication and that only the framing of the very local ‘Flore de Buitenzorg’ (from Batavia to the peak of Mt Gedeh, W. Java) could be realized as a cautious approach to later projects of large size. Even for this very restricted Flora the volume on Phanerogams never appeared, except for the Orchids. And that was for Java, botanically the best known island in the whole archipelago with a proportionally poor flora compared with the true Malesian element in the Larger Sunda islands. Even a dozen years later BACKER spent three years in the field to get an overall coverage of the Javanese flora!
This illustrates the desperate position MERRILL had to face, a position that contemporary botanists working on the Malesian flora do not always realize. I have sometimes traced arro- gance in the rejection by a few contemporary, ‘angry’ youngsters of MERRILL’s floristic methods and premature publication of novelties. It also is for their education that I have in some detail accounted for MERRILL’s work and life, projected on the background of the state of Malesian botany in the first decades of this century, with full exposure of the then prevailing conditions in the hope that they will reach a better understanding of the level at which MERRILL had to work by necessity. I have also pointed out what our present-day knowledge owes to his collecting drive and to the pioneer works, bibliographies and other tools which are in our constant use and which he had created from scratch. Possibly they may ask themselves what they would have achieved had they stood in MERRILL’s shoes in 1902.
Naturally it is a blessing to be able to work now in well-equipped centres provided with ample facilities, under social security and pension conditions, at leisure on a regional-monographic basis. But let us remember that we harvest what others have sown during the past seven decades.
Among those who paved the way MERRILL was the outstanding figure, a man of boundless energy and vision, a great organisator and a great botanist. It is for these reasons that we dedicate this volume with due respect to the memory of this prominent American scientist.
BIOGRAPHICAL SOURCES
BREMEKAMP, C. E. B.: Levensbericht, E. D. Merrill. Jaarboek Kon. Ned. Ak. Wet. (1955/56) 1-7.
BurKILL, I. H.: Prof. E. D. Merrill. Nature 177 (1956) 687-688.
—: Elmer Drew Merrill. Proc. Linn. Soc. Lond. 168 (1957) 51-56.
Ewan, J.: E. D. Merrill. J. Wash. Ac. Sci. 46 (1956) 267-268.
Howarp, R. A.: Elmer Drew Merrill, 1876-1956. J. Arn. Arb. 37 (1956) 197-216, port.
Lam, H. J.: A Dutch tribute to Merrill’s work. Natuurwet. Tijdschr. Ned. Ind. 102 (1946) 153- 154.
LANJouw, J.: Elmer D. Merrill, 1876-1956. Taxon 5 (1956) 21.
MERRILL, E. D.: Autobiographical: Early years, the Philippines, California. Asa Gray Bull. n.s. 2 (1953) 335-370, 3 ports.
QUISUMBING, E.: Elmer Drew Merrill. Philip. J. Sc. 85 (1956) 181-188.
RicketTT, H. W.: Elmer Drew Merrill, 1876-1956. J. N.Y. Bot. Gard. 6 (1956) 84, 90.
RossBins, W. J.: Elmer Drew Merrill (1876-1956). Yearb. Amer. Philos. Soc. (1956) 117-119.
—: Elmer Drew Merrill, 1876-1956. A Biographical Memoir. Biograph. Memoirs 32 (1958) 273-333.
ROLLINs, R. C.: Elmer Drew Merrill, Administrator and Botanist. Science, n.s. 123 (1956) 831- 832.
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FLORA MALESIANA
ScHuLtTes, R. E.: Elmer Drew Merrill. An Appreciation. Taxon 6 (1957) 89-101, 2 ports. and a caricature.
VeRDOORN, F. (ed.): Merrilleana. A selection of the general writings of Elmer Drew Merrill, Sc.D., LL.D. Chronica Botanica 10 (1946) 129-393.
WaLkKeR, E. H.: Dr. Elmer D. Merrill, 1876-1956. Pacific Sci. 11 (1957) 135-136.
C. G. G. J. VAN STEENIS
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ABBREVIATIONS AND SIGNS
acc. = according
Ak. Bis. = Aklan Bisaya (Philip. language) Alf. Cel. = Alfurese Celebes (language) alt. = altitude
Anat. = Anatomy
Ap. = Apayao (Philip. language)
app. = appendix, appendices
appr. = approximate
Apr. = April Arch. = Archipelago atl. = atlas
auct. div. = auctores diversi; various authors
auct(t). mal. = auctores malayenses; authors dealing with Malesian flora
auct(t). plur. = auctores plures; several authors
Aug. = August
Bag. = Bagobo (Philip. language)
basionym = original name of the type specimen; its epithet remains permanently attached to the taxon which is typified by it provided it is of the same rank
Bg. = Buginese (language)
Bik. = Bikol (Philip. language)
Bil. = Bila-an (Philip. language)
Bill. = Billiton
Bis. = Bisdya (Philip. language)
Bon. = Bontok (Philip. language)
Born. = Borneo
Bt = Bukit; mountain
Bug. = Buginese (language)
Buk. = Bukidnon (Philip. language)
. = circiter; about
C. Bis. = Cebu Bisaya (Philip. language)
cf. = confer; compare
Chab. = Chabecano (Philip. language)
citations = see references
cm = centimetre
c.n. = see comb. nov.
comb. nov. = combinatio nova; new combination
c.S. = cum suis; with collaborators
cum fig. = including the figure
cur. = curante; edited by
D (after a vernacular name) = Dutch
Daj. = Dyak (language)
Dec. = December
D.E.I. = Dutch East Indies
descr. added behind a reference = means that this contains a valid description
diam. = diameter
Distr. (as an item) = Distribution
Distr. (with a geographical name) = District
ditto = the same, see do
Div. = Division, or Divide
div. = diversus (masc.); various
do = ditto (Ital.); the same
Dum. = Dumagat (Philip. language)
dupl. = duplicate
E = east (after degrees: eastern longitude)
E (after a vernacular name) = English
Ecol. = Ecology
ed. = edited; edition; editor
e.g. = exempli gratia; for example
elab. = elaboravit; revised
em(end). = emendavit; emended
em(erg). ed. = emergency edition
Engl. = English
etc., &c. = et cetera; and (the) other things
ex auctt. = ex auctores; according to authors
excl. = exclusus (masc.); excluding, exclusive of
ex descr. = known to the author only from the description
f. (before a plant name) = forma; form
f. (after a personal name) = filius; the son
f. (in citations) = figure
fam. = family
Feb(r). = February
fide = according to
fig. = figure
fl. = flore, floret (floruit) ; (with) flower, flowering
For. Serv. = Forest Service
Sr. = fructu, fructescit; (with) fruit, fruiting
Fr. (after a vernacular name) = French
G. = Gunung (Malay); mountain
Gad. = Gaddang (Philip. language)
gen. = genus; genus
genus delendum = genus to be rejected
Germ. = German
geront. = Old World
haud = not, not at all
holotype = the specimen on which the original description was actually based or so designated by the original author
homonym = a name which duplicates the name of an earlier described taxon (of the same rank) but which is based on a different type species or type specimen; all later homonyms are nomencla- turally illegitimate, unless conserved
I. = Island
ib (id). = ibidem; the same, in the same place
Ibn. = Ibanag (Philip. language)
ic. = icon, icones; plate, plates
ic. inedit. = icon ineditum, icones inedita; inedited plate(s)
id. = idem; the same
i.e. = id est; that is
If. = Ifugao (Philip. language)
Ig. = Igorot (Philip. language)
Ilg. = Llongot (Philip. language)
Ik. = Lloko (Philip. language)
in adnot. = in adnotatione; in note, in annotation
incl. = inclusus (masc.); including, inclusive(ly)
indet. = indetermined
Indr. = Indragiri (in Central Sumatra)
inedit. = ineditus (masc.); inedited
in herb. = in herbario; in the herbarium
in litt. = in litteris; communicated by letter
in sched. = in schedula; on a herbarium sheet
in sicc. = in sicco; in a dried state
in syn. = in synonymis; in synonymy
Is. = Islands
Is. (after a vernacular name) = language)
Ism. = Isamal (Philip. language)
isotype = a duplicate of the holotype; in arboreous plants isotypes have often been collected from a single tree, shrub, or liana from which the holotype was also derived
Iv. = Ivatan (Philip. language)
Isindi (Philip.
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FLORA MALESIANA
J(av). = Javanese (language)
Jan. = January
Jr = Junior
Klg. = Kalinga (Philip. language)
Kul. = Kulaman (Philip. language)
Kuy. = Kuyonon (Philip. language)
Lamp. = Lampong Districts (in S. Sumatra)
Lan. = Lanao (Philip. language)
lang. = language
l.c. = loco citato; compare reference
lectotype = the specimen selected a posteriori from the authentic elements on which the taxon was based when no holotype was designated or when the holotype is lost
livr. = livraison, part
(heea— e-x(plurs)
m = metre
M = Malay (language)
Mag. = Magindando (Philip. language)
Mak. = Makassar, Macassar (in SW. Celebes)
Mal. = Malay(an)
Mal. Pen. = Malay Peninsula
Mand. = Mandaya (Philip. language)
Mang. = Mangyan (Philip. language)
Mar. = March
Mbo = Manobo (Philip. language)
Md. = Madurese (language)
Minangk. = Minangkabau (a Sumatran language)
min. part. = pro minore parte; for the smaller part
mm = millimetre
Mng. = Mangguangan (Philip. language)
Morph. = Morphology
ms(c), MS(S) = manuscript(s)
Mt(s) = Mount(ains)
n. = numero; number
N = north (after degrees: northern latitude); or New (e.g. in N. Guinea)
NE. = northeast
nec = not
neerl, = Netherlands, Netherlands edition
Neg. = Negrito (Philip. language)
N.E.I. = Netherlands East Indies
neotype = the specimen designated to serve as nomenclatural type when no authentic speci- mens have existed or when they have been lost; a neotype retains its status as the new type as long as no authentic elements are recovered and as long as it can be shown to be satisfactory in accordance with the original description or figure of the taxon
N.G. = New Guinea
N.I. = Netherlands Indies
no = numero; number
nom. = nomen; name (only) = nomen nudum
nom. al. = nomen aliorum; name used by other authors
nom. alt(ern). = nomen alternativum; alternative name
nom. cons(ery). = nomen conservandum, nomina
conservanda; generic name(s) conserved by the International Rules of Botanical Nomenclature
nom. fam. cons. = nomen familiarum conservan- dum; conserved family name
nom. gen. cons. = see nomen conservandum
nom. gen. cons. prop. = nomen genericum conser-
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vandum propositum; generic name proposed for conservation
nom. illeg(it). = nomen illegitimum; illegitimate name nom. leg(it). = nomen legitimum; legitimate name
nom. nov. = nomen novum; new name
nom. nud. = nomen nudum; name published with- out description and without reference to pre- vious publications
nom. rej(ic). = nomen rejiciendum; name rejected by the International Rules of Botanical No- menclature
nom. seminudum = a name which is provided with some unessential notes or details which cannot be considered to represent a sufficient descrip- tion which is, according to the International Rules of Botanical Nomenclature, compulsory for valid publication of the name of a taxon
nom. subnudum = nomen seminudum
nom. superfl. = a name superfluous when it was published; in most cases it is a name based on the same type as an other earlier specific name
non followed by author’s name and year, not placed in parentheses, and put at the end of a citation = means that this author has published the same name mentioned in the citation in- dependently. These names (combinations) are therefore homonyms. Compare p. 111 under Wahlenbergia lines 6 & 7 where there appear to be three different genera all called Lightfootia by three different authors, and belonging to three different families. The same can happen to taxa of lower ranks, for example species; compare p. 1294 lines 7 & 8 under Lobelia heyniana, where it appears that there are two different species of Lobelia named L. decurrens, viz by ROTH and by CAVANILLES in which the latter antedates the former
(non followed by abbreviation of author’s name) before a reference (citation) headed by an other author’s name = means that the second author has misinterpreted the taxon of the first author. Compare for example p. 1265 under Lobelia alsinoides lines 7 & 8 the synonym name L. filiformis; CAVANILLES misapplied in his de- scription and figure the name LAMARCK had given to another species through an erroneous identification. The sense CAVANILLES gave to the Lamarckian plant name does not invalidate the latter: CAVANILLES’s use of the name also does not represent a proper synonym; his name has no status and its mention serves only to indicate the identity of his text and plate
non al. = non aliorum; not of other authors
non vidi = not seen by the author
noy. = nova (femin.); new (species, variety, e/c.)
Nov. = November
n.s. = new series
Nn. Sp. = nova species; new species
n. (sp.) prov. = nomen (specificum) provisorium; provisional new (specific) name
n.v. = non vidi; not seen
NW. = northwest
Oct. = October
op. cit. = opere citato; in the work cited
Abbreviations and signs
p. = pagina; page
P. = Pulau, Pulu (in Malay); Island
Pal(emb). = Palembang
Pamp. = Pampangan (Philip. language)
Pang. = Pangasinan (Philip. language)
paratype = a specimen cited with the original description other than the holotype
part. alt. = for the other part
P. Bis. = Panay Bisaya (Philip. language)
P.[. = Philippine Islands
pl. = plate
plurim, = plurimus; most
p.p. = pro parte; partly
pr. max. p. = pro maxima parte; for the greater part
pro = as far as is concerned
prob. = probabiliter; probably
prop. = propositus; proposed
Prov. = Province
pr.p. = pro parte; partly
pt = part
quae est = which is
quoad basionym, syn., specimina, etc. = as far as the basionym, synonym((s), specimen(s), efc. are concerned
references = see for abbreviations the list in vol. 5, pp. cxly-clxv
Res. = Residency
resp. = respective(ly)
S = south (after degrees: southern latitude)
S (after a vernacular name) = Sundanese (lan- guage)
Sbl. = Sambali (Philip. language)
SE. = southeast
sec. = secus; according to
sect. = sectio; section
sens. ampl. (ampliss.) = sensu amplo (amplissimo) ; in a wider sense, in the widest sense
sens. lat. = sensu lato; in a wide sense
sens. str. (strictiss.) = sensu stricto (strictissimo) ; in the narrow sense, in the narrowest sense
Sept. = September
seq., Seqq. = sequens, sequentia; the following
ser. = series
s.l. = sensu lato; in a wide sense
S.-L. Bis. = Samar-Leyte Bisaya (Philip. language)
Sml. = Samal (Philip. language)
5.n. = sine numero; (specimen) without the col- lector’s number
Sp. = Spanish (language)
sp(ec). = species; species
specim. = specimen(s)
sphalm. = sphalmate; by error, erroneous
Spp. = species; species (plural)
Sr — Senior
5.5. = see sens. str.
ssp. = subspecies; subspecies
Susirs — SEG sens. str:
Stat. nov. = status nova; proposed in a new rank
Sub. = Subanum (Philip. language)
subg(en). = subgenus; subgenus
subsect. = subsectio; subsection
subsp. = subspecies; subspecies
Sul. = Sulu (Philip. language)
Sum. E.C. = Sumatra East Coast
Sum. W.C. = Sumatra West Coast
Suppl. = Supplement
SW. = southwest
syn. = synonymum; synonym
synonyms = the names of taxa which have been referred to an earlier described taxon of the same rank and with which they have been united on taxonomical grounds or which are bound together nomenclaturally
syntypes = the specimens used by the original author when no holotype was designed or more specimens were simultaneously designated as type
t. = tabula; plate
Tag. = Tagalog (Philip. language)
Tagb. = Tagbanua (Philip. language)
Tagk. = Tagaka-olo (Philip. language)
Tapan. = Tapanuli (in NW. Sumatra)
taxon = each entity throughout the hierarchic ranks of the plant kingdom which can be described and discriminated from other taxa of the same rank
Taxon. = Taxonomy
Tg = Tandjung (Malay); cape
Ting. = Tinggian (Philip. language)
Tir. = Tirurai (Philip. language)
transl. = translated
type = each taxon above the rank of a species is typified by a type belonging to a lower rank, for instance a family by a genus, a genus in its turn by a species; a species or infraspecific taxon is typified by a specimen. The name of a taxon is nomenclaturally permanently attached to its type; from this it cannot be inferred that the type always represents botanically the most typical or average structure found in the cir- cumscription of the taxon
type specimen = the specimen or other element to which the name of a species or infraspecific taxon is (nomenclaturally) permanently at- tached; botanically a type specimen is a random specimen on which the name was based by de- scription. Therefore, it does not need to repre- sent the average or most typical representative of a population. See holotype, isotype, lectotype, syntype, paratype, and neotype
typ. excl. = typo excluso; type exluded
typ. incl. = typo incluso; type included
typus = see type and type specimen
var. = varietas; variety
var. nov. = varietas nova; new variety Vern. = Vernacular vide = see
viz = videlicet; namely
vol. = volume
W = west (after degrees: western longitude) Yak. = Yakan (Philip. language)
@ = diameter
3 = male (flower, etc.)
2 = female (flower, etc.)
3, 9 = bisexual (flower)
) (2) = dioecious with unisexual flowers ©) = monoecious with unisexual flowers
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FLORA MALESIANA
(3%) = polygamous
< = less than (size, number, efc.) (2°) = polygamous x 2/5 = 2/5 of natural size oo = many montana = means that the epithet montana is ~ = more than (in size, number, etc.)
that of a hybrid
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ADDENDA, CORRIGENDA ET EMENDANDA
As was done in the preceding volumes, it seemed useful to correct some errors which have crept into the text of volumes 4, 5 & 6 as well as to add some additional data, new records, and new species or other taxa which came to our knowledge and are worth recording.
Additions of the Amaranthaceae | owe to Dr. R. C. BAKHUIZEN VAN DEN BRINK f. and Mr. J. F. VELDKAMP, of the Alismataceae and Hydrocharitaceae to Dr. C. DEN HARTOG, of the Celastraceae and Thymelaeaceae p.p. to Dr. DinG Hou, of the Malpighiaceae to Dr. M. Jacoss, of the Burseraceae p.p. to Dr. C. KALKMAN, of the Caprifoliaceae to Dr. J. H. KERN, of the Burseraceae p.p., Connaraceae, Dichapetalaceae, Goodeniaceae and Loganiaceae to Dr. P. W. LEENHOUTS, of the Gnetaceae to Dr. F. MARKGRAF, Of the Simaroubaceae to Mr. H. P. Nootesoom, of the Convolvulaceae to Dr. S. J. VAN OosTsTROoM, of the Thymelaeaceae p.p. to Mr. H. K. Airy SHAW, of the Ericaceae and Flacourtiaceae to Dr. H. SLEUMER.
Printing errors have only been corrected if they might give rise to confusion.
Volume and page number are separated by a colon. Page numbers provided with either a or b denote respectively the left and right columns of a page.
Aceraceae 4556, cultivated, a recent introduction; Morobe Distr., Bulolo, NGF 7384). All 4: 3-4, Acer laurinum Hassk. identifications by SAUER, 1971. 592ab Add to Distr.: Rare in Borneo and the 4: 79b Change the name of species 8 into: few localities at unusual lowland altitude; 8. Amaranthus hybridus L. ssp. incurvatus now also found on Mt Kinabalu at c. (GREN. & GODR.) BRENAN var. panicula- 1000 m (SAN 38438); probably also in the tus (L.) MANsr. See MANSFELD, Die mountains of N. Thailand, as E. MURRAY Kulturpfl. Beih. 2 (1959) 54, and BRENAN, merged (in sched.) A. garrettii CRAIB, Watsonia 4 (1961) 268. Kew Bull. (1920) 301, with it. 4: 91 Emend the key to the species of Al/ter- nanthera as follows, second entry of fork Aizoaceae under 3: 3. Three outer tepals in their lower 4: 269a Glinus lotoides L. 1/31/59 distinctly 3-nerved, their bases Add to synonymy: Holosteum hirsutum at least indurate. Anthers 5. Pseudo- L. Sp. Pl. (1753) 88. Holotype from staminodes well-developed, wider than India. Cf. STEEN. Blumea 13 (1965) 167. long to ligular, lobed or fimbriate. 4: 274a Trianthema triquetra ROTTL. ex WILLD. Hairs dentate. Add to Distr.: Philippines (Mindanao). 3a. Leaves mucronate, often coloured. Cf. STEEN. Blumea 12 (1964) 320. Anthers linear, °/4-1 mm long; pseudo-staminodes ligular, apex Alismataceae (DEN HARTOG) fimbriate, as long as to longer than the stamens. Pistil bottle-shaped, 5: 327 Bottomline, replace ‘6. S. sagittifolia ssp. apex not emarginate. Apparently leucopetala’ by: 6. S. trifolia. never setting seed in Malesia. 5: 332b Replace the name Sagittaria sagittifolia 3. A. ficoidea ssp. leucopetala (MiQ.) Hartoc by: 3a. Leaves acute, not mucronate (in 6. Sagittaria trifolia LINNE, Sp. Pl. 2 Malesia). Anthers reniform to (1753) 933, and add the subspecific name ellipsoid, !/3—?/5 mm long; pseudo- to its synonymy. There is no change in the staminodes wider than long, with text. 3-4 + triangular lobes, reaching 5: 333b In text line 3 from bottom, replace ‘S. the base of the anthers or shorter. sagittifolia ssp. leucopetala’ by: S. trifolia. Pistil broadly obcordate. Seeds 5: 334b Add to the Notes: This taxon cannot be usually developed, (broadly) obcor- maintained as a subspecies of S. sagitti- date, narrowly winged. folia, as the sepals in the mature 9 flowers 3a. A. paronychioides are reflexed. For this reason it belongs to 4: 93a, Alternanthera ficoidea (L.) R.BR. ex another species group within the genus. 594b R. & S. 1819, non A. ficoides P. BEAUVY. Fl. Oware & Benin 1818. As these are Amaranthaceae homonyms (Code Art. 75) this specific (BAKHUIZEN VAN DEN BRINK, name is illegitimate. VAN STEENIS & VELDKAMP) We are, however, not certain of its full synonymy and for this reason tentatively 4: 79b Amaranthus dubius Marv. refrain from making new combinations Add to Distr.: Central Java (Mt Lawu, both for the species and the variety. It Temanggung: LORZING 346), Lesser may be that Telanthera manillensis WALP. Sunda Is. (Alor: JAAG 413), New Guinea 1843 contains the oldest basionym.
(West: Star Mts, Sibil Valley, KALKMAN To avoid this difficulty we accept tenta-
FLORA MALESIANA
[ser. I, vol. 68
Fig. 1. Alternanthera bettzickiana (REGEL) NICHOLS. /. Flower, abaxial, without bract and bracteoles
< 12, 2. ditto, without abaxial tepals, »
12, 3. staminal tube, « 24, 4. young fruit, x 24. — A. parony-
chioides St. Hit. var. paronychioides. 5. Flower, abaxial, without bract, « 12, 6. ditto, without bracteoleS
and abaxial tepals,
4: 93b
12, 7. staminal tube, x
tively specific status for this taxon in Flora Malesiana:
Alternanthera bettzickiana (REGEL) NI- CHOLS. Ill. Dict. Gard. 1 (1884) 59 (‘bettzichiana’); Voss in Vilmorin’s Blu- mengartn. ed. 3 (by SIEBERT & Voss), 1 (1895) 69 (non vidi, ex Ind. Kew.); ASCHERS. & GRAEBN. Synopsis 5, 1 (1914) 365. — Telanthera bettzickiana REGEL, Gartenflora 11 (1862) 178, descr.; Ind. Sem. Hort. Petrop. (1862) 28 (non vidi). — Fig. 1.
VELDKAMP (Blumea 19, 1971, 169) assumes the addition by NICHOLSON “Brasilia, 1862’ to be a very indirect reference to REGEL’s basionym with which I can agree. VELDKAMP, /. c., also indicated that var. versicolor (REGEL) BACKER, sometimes regarded as a sepa- rate species, must be regarded as a synonym of A. bettzickiana.
Add the following species:
3a. Alternanthera paronychioides St. Hiv. Voy. Brésil 2, 2 (1833) 43; DuLTA & Mitra, Ind. Forester 87 (1961) 304, f. 2; PEDERSEN, Darwiniana 14 (1967) 437; VELDKAMP, Blumea 19 (1971) 167, f. 6-8.
24, 8. young fruit (J-4 Cayosa 75, 5-8 CAyosa 104).
var. paronychioides — Fig. 1.
Prostrate, branched annual. Branches up to 70 cm, rooting at the nodes, angular and villose at apex, becoming terete and glabrous at base, indument only persis- tent at the nodes. Hairs dentate. Leaves spathulate-oblong to -lanceolate, up to 7 by 2 cm, acute, never mucronate, base cuneate, narrowed into the petiole, moderately appressed-pubescent to glab- rous, usually tufted under the inflores- cences. Spikes sessile, apical on short axillary branches, subglobose to shortly cylindric, up to 2 by 1 cm. Bracts, bracteoles scarious, white. Bracts ovate- oblong, 21/4-3 by 11/4-11/2 mm, acu- minate, glabrous, -+_ convex, 2/3—3/4 times as long as tepals, longer than brac- teoles. Bracteoles ovate-oblong, 13/4—21/2 by 3/4 mm, acute to acuminate, glabrous, + falcately folded along midrib. Tepals oblong to lanceolate, acute to mucronu- late, in lower half 3-nerved and sparsely strigose, upper half stiff, but not coria- ceous. Adaxial tepal 3—4 by 4/s—11/4 mm, flat; abaxials 31/44 by 1-11/4 mm, flat; laterals 21/4—31/4 by 1—11/4 mm, folded
September 1972]
Addenda, corrigenda et emendanda
917
4: 964, 594b
5: 415a
5: 207b
along midrib. Stamens 5, all fertile (in the material seen); filaments at base united into a c. '/4 mm high cup, free parts c. 1/2 mm long, filiform; anthers reniform to ellipsoid, 4/3—!/4 mm long, yellow; pseudo-staminodes wider than long, with 3-4 + triangular lobes, reach- ing the base of the anther or shorter. Pistil broadly obcordate with a short, stout style in the notch; stigma capitate, papillose. Utricle broadly obcordate to obcordate, up to 13/4 by 11/4 mm, narrowly winged, brown; the plants seem to fructify during drying; so young fruits are usually present, contrary to the situa- tion in A. ficoidea ssp. bettzickiana, where they have never been observed in Malesia.
Distr. Native of tropical America, introduced as an alien in other countries, including Europe and _ Indo-Malesia, India, Thailand; in Malesia: West-Java, Philippines (Luzon: Manila, Quezon City; Guimaras I.; Mindanao).
Ecol. Disturbed places, railway tracks, banks of rivers and lakes, up to 250 m.
Vern. & Uses. Simsim (Bis.), used for feeding hogs (Mindanao).
Note. By BACKER confused with A. ficoidea ssp. bettzickiana and in habit difficult to discern from it, but distinctly different in the structure of the filaments, pseudo-staminodes, and anthers. Gomphrena celosioides MART.
Add to references: STEEN. Nova Guinea, Bot. 23 (1965) 495.
The spreading of this weed continues steadily eastwards and has now reached Timor, Papua, and Micronesia. It was also found in Goodenough I., off NE. Papua, along an old military road (Brass 24432).
Batidaceae
Batis argillicola VAN ROYEN.
Add to Distr.: It also occurs in Northern Queensland. Cf. S. T. BLAKE, Proc. R. Soc. Queensl. for 1961, 73 (1963) 61. It occurs there in Burke Distr., near Karumba, at the mouth of the Norman R., on clay-pans adjacent to the shore, in exactly the same habitat as near Merauke.
Betulaceae
The basionym of Alnus maritima (Marsu.) NuTTALL is Betula-Alnus mari- tima MARSHALL, Arb. Am. (1785) 20. Though not noted, I had not seen this work and depended on _ authority. Dr. BAKHUIZEN VAN DEN BRINK Saw it at Paris, in 1959, and reported that the only description or distinction mentioned by MARSHALL is ‘leaves long and narrow’. This, I believe, cannot be accepted even
5: 208
5) 210
Se li
5: 214a
Se021Sa
5: 218a
5: 220
as a specific diagnosis, and the name I regard as an invalid nomen seminudum. The proper name for this species is: Alnus japonica (THUNB.) STEUD.
Legend fig. 1: Change Alnus maritima (MarsH.) Nutt. into Alnus japonica (THUNB.) STEUD.
Burseraceae
(LEENHOUTS, Dacryodes and Santiria in co-operation with KALKMAN)
Add to Dispersal, 2nd paragraph: In
Java, fruits of Canarium are occasionally
eaten by bats (see VAN DER PiL, Acta
Bot. Neerl. 6, 1957, 299).
Add to Wood anatomy: BurGEss, Tim-
bers of Sabah (1966) 60-70, f. 11.
Insert after the section on
anatomy:
Phytochemistry. See HEGNAUER, Che-
motaxonomie 3 (1964) 310-318, 647, 669.
Add to Morphology, 3rd paragraph: See
for a more detailed discussion on the
stipules of the Burseraceae and some related families WEBERLING & LEEN-
HoutTs, Abh. Ak. Wiss. Lit. Mainz M.-N.
Kl. 1965, n. 10 (1966) 495-584.
Protium javanicum Buro. f.
Add to description under Branches,
before ‘spines’: sometimes branched.
Add to Distr.: Flores.
Protium macgregorii
LEENH.
In description delete after
(2 unknown). Insert after
3 flowers.
Add to Distr.: Also in Normanby I.
Add to Ecol.: also in swamps. Change
highest altitude into 1100 m.
Garuga floribunda DECNE.
Add to description, Ist sentence: decid-
uous.
Add to Distr.: Malay Peninsula.
Dacryodes VAHL.
Add to Distr.: According to NoRMAND,
Comptes Rendus [Ve Réun. A.E.T.F.A.T.
(1962) 291, the number of African species
is about 10.
Replace the Key to the species by the
following one:
1. Leaf-bearing branchlets c. 2 cm thick. Leaves 6-8-jugate; petiole c. 20 cm long; leaflets 20-60 cm long, with 18-38 pairs of nerves. Inflorescence 35-120 cm long, the main branches up to 55 cm long 5. D. kingii
1. Leaf-bearing branchlets up to 11/2 cm thick. Leaves rarely more than 4-jugate; petiole mostly up to 15 cm long; leaflets mostly less than 20, rarely up to c. 40 cm long, with usually less than 15, rarely up to 25 pairs of nerves. Inflorescences
Wood
(Be Me “BAi)
‘Flowers’: SPistiles aan
918
FLORA MALESIANA
[ser. I, vol. 68
mostly shorter, their main branches up to 30 (2) or 40 (4) cm long.
2. Lowermost pair of leaflets more or less stipule-like,. often caducous. Inflorescences 10-140 cm long, their main branches up to 30 (2) or 40 (3) Cmiblong.)) ue ete a) laxa
2. Lowermost pair of leaflets not much different from the others, not cadu- cous. Inflorescences mostly less than 30, rarely up to 60 cm long, their main branches up to c. 25 cm long.
3. Indumentum at least partly consist- ing of minute, stellate hairs. 16. D. nervosa 3. Indumentum consisting of simple hairs only. 4. Nodes of leaf-rachis distinctly swollen.
5. Leaflets when dried greenish above. Inflorescences mostly axil- lary, together sometimes pseudo- terminal, rarely terminal. Calyx sparsely hairy, corolla (sub)glab- rous. Fruits distinctly bulging on one side . . 1. D. rugosa
5. Leaflets when dried brown above. Inflorescences terminal. Flowers densely tomentose. Fruits not
bulging . . 7. D. rubiginosa 4. Nodes of leaf-rachis not or hardly swollen.
6. Inflorescences terminal (vegeta- tive terminal bud absent), more- over sometimes in the upper leaf-axils.
7. Pith of branchlets without vascu- lar strands.
8. Inflorescences densely woolly; calyx slightly pubescent, corol- la (sub)glabrous.
2. D. costata
8. Inflorescences densely minutely
tomentose, including the
flowers . . 6. D. incurvata
7. Pith of branchlets with vascular strands.
9. Leaf-bearing branchlets !/4-!/2 cm thick, pith with some to many vascular strands. Petiole 3-15 cm long, pith with some to many vascular strands; leaflets glabrous, with 6-18 pairs of nerves. Peduncle 0-6 cm long.
Fruits 11/2-3 by 3/4-13/4 cm. 6. D. incurvata
9. Leaf-bearing branchlets c. 1/2 cm thick, pith with some vascu- lar strands. Petiole 3-9 cm long, pith with some vascular strands; leaflets beneath on midrib and nerves densely pubescent, with 7-13 pairs of nerves. Inflorescences branched
from the base. Fruits 2—21/2 by 1-11/4 cm. 7. D. rubiginosa
9. Leaf-bearing branchlets c. 1 cm
thick, pith with many vascular strands. Petiole 9-15 cm long, pith with many vascular strands; leaflets glabrous, with 13-18 pairs of nerves. Peduncle 3/42 cm. Fruits 4-43/4 by 221\/2cem . . 8. D. elmeri
6. Inflorescences axillary (vegetative
terminal bud present), sometimes together pseudoterminal.
10. Leaflets pubescent beneath.
11. Leaves 1-4-jugate. Inflores- cences short-peduncled.
3. D. puberula
11. Leaves 7—9-jugate. Inflores- cences long-peduncled.
9. D. rostrata f. pubescens
10. Leaflets glabrous or only hairy
on the midrib beneath.
12. Pith of branchlets with some to many, pith of petiole with several vascular strands.
13. Branchlets densely minutely villous. Leaves 3-4-jugate; petiole strongly flattened at base, 9-15 cm long; leaf- lets 12-22 cm long, brown above when dry, shortly acuminate, with 13-18 pairs of nerves, reticulation incon- spicuous above. Inflores- cences 17-45 cm long, pe- duncle 3/4-2 cm. Stamens unknown. Fruit 4-4/4 by 2-2!/2cm . 8. D. elmeri
13. Branchlets glabrous except the tip. Leaves 2—10-jugate; petiole terete to strongly flattened at base, 3-26 cm long; leaflets 3-25 cm long, brownish to greenish above when dry, mostly rather long and slender acuminate, with 5-20 pairs of nerves, reti- culation manifest above. In- florescences 5-35 cm long, peduncle 0-15 cm. Stamens adnate to the disk. Fruit 13/44 by 3/4-21/4 cm.
9. D. rostrata
13. Branchlets glabrous except the tip. Leaves 3-4-jugate; petiole strongly flattened at base, 41/2-10 cm long; leaf- lets 5121/2 cm long, green- ish above when dry, shortly acuminate, with 10-13 pairs of nerves, reticulation mani- fest above. Inflorescences 71/2-10 cm long, peduncle 31/2-5 cm. Stamens free
September 1972]
2216
fe 220 222b
mn
5: 224a
5: 2246
Sie225a
Addenda, corrigenda et emendanda
919
from the disk. Fruit un- known 10. D. crassipes 12. Pith of branchlets without,
pith of petiole with few vascular strands. 14. Petiole terete to slightly
flattened at base. Wall of fruit kernel thick and hard. 12. D. expansa 14. Petiole strongly flattened to channelled at base. Wall of fruit kernel very thin.
15. Leaflets 3-6 times as long as wide; leaves 3—15-jugate, the petiole 2—20 cm long, leaflets 6-42 cm long.
14. D. longifolia
15. Leaflets up to 3 times as long as wide; leaves up to 5-jugate, the petiole up to 8 cm long, leaflets up to 17 cm long.
16. Base of leaflets often oblique. Peduncle 0-4 cm long. Calyx 3 mm high. Fruit 21/4-31/2 by 11/4-2
cm 11. D. macrocarpa 16. Base of leaflets not oblique. Peduncle 1/2-1
cm long. Calyx smaller. Fruit 16-18 by 9-11 mm. 15. D. breviracemosa Dacryodes rugosa (BL.) H. J. LAM. Change in description: Branchlets ...; pith without or with many peripheral vascular strands ... Petioles terete to distinctly flattened at base. Add to Ecol.: Also in secondary forests. Dacryodes costata (BENN.) H. J. LAM. Change in description: Leaves ...., glabrous to densely pubescent ...; nerves up to 17 or 18 pairs. Fruits sometimes ellipsoid. Add to Ecol.: Also in secondary forests. Dacryodes laxa (BENN.) H. J. LAM. Description: add to Ist sentence: small buttresses sometimes present. Branchlets . .; pith sometimes with a closed cylinder of many small vascular strands. Dacryodes kingii (ENGL.) KALKMAN. Change in description, Ist sentence ‘12 m’ into: 20 m. Branchlets ...; pith with some peripherally arranged to several, partly peripheral, partly scattered small vascular strands. Petiole up to 28 cm. Leaflets ...; apex up to 2 cm blunt- acuminate. Fruits oblong or ovoid, ..., more or less oblique. Dacryodes incurvata (ENGL.) H. J. LAM. Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 164. Description: Change greatest height of tree into 40 m. Leaves 1-—5-jugate. 3 Panicles may be up to 30 cm long. Fruits ovoid to ellipsoid, up to 31/2 cm
5: 226a
5: 226b
22a
S22 1D 5: 228a
5: 2285
5: 229a
53 2296
long, said to be orange when ripe.
Add to Ecol.: SAN 25326 and 29004 are collected at 1400 and 1500 m altitude resp. Fi. Jan.Aug.
Add: Uses. Fruits edible.
Dacryodes rostrata (BL.) H. J. LAM. Description: Change greatest height of tree into 45 m. Branchlets exceptionally up to 15mm &. Petioles terete to strongly flattened at base, . . . Leaflets up to 25 cm long. Sometimes all nerves looped and joined near the margin.
Add after Fruits: yellow to purple when ripe.
Add to Uses: In Brunei cultivated by the Kedayans for the fruits, the pulp of which is eaten.
Dacryodes macrocarpa (K1NG) H. J. LAM. Change in description, 3rd line, ‘glabrous’ into: hairy at the tip only.
Line 4 from top, add after ‘Fruits ovoid’: to ellipsoid.
var. macrocarpa.
Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 164.
Ecol.: Change highest altitude into 1400 m.
var. kostermansii (KALKM.) KALKMAN. Add to Distr.: N. Borneo.
Dacryodes expansa (RIDL.) H. J. LAM. Add to literature: SMyTHIES, Common Sarawak Trees (1965) t. 8.
Change in description line 4: Petioles 21/2-91/2 cm. Change in line 9 ‘not arching’ into: arching or not. Change dimensions of fruits into: 31/2—-S by 2-31/2 by 21/2-3 cm; add: rosy apple-red when ripe.
Change Uses: Only the pulp of the fruits is eaten.
13. Dacryodes papuana Husson.
This turned out to be synonymous with Scutinanthe brevisepala_ LEENH.; cf. LeEENH. Blumea 12 (1964) 19.
Dacryodes longifolia (KING) H. J. LAM. Description: Add after tree: 10 m by 30 cm. Line 2, after ‘glabrous’ add: apart from the tip. Add to fruits: red when ripe. Add to Distr.: Philippines (Mindanao). var. longifolia.
Change in description: 4-15-jugate. Add to Distr.: Philippines.
In Ecol. change as follows: Fr. Nov. (Mal. Pen.), May (Philippines).
Add after 15. Dacryodes breviracemosa:
16. Dacryodes nervosa (H. J. LAm) LEENH. Blumea 12 (1964) 19. — Santiria nervosa H. J. LAM, Ann. Jard. Bot. Btzg 42 (1932) 206, t. 11 f. 65; Bull. Jard. BoteBize sil, 1251932) S87sk. 6 t 29; KALKMAN, Blumea 7 (1954) 539; LEENH. Fl. Mal. I, 5 (1956) 233.
Tree, 12-30 m by 20-100 cm, with up to 21/2 m high buttresses which are 11/2 m
920
FLORA MALESIANA
[ser. I, vol. 66
spreading. Branchlets 4-8 mm thick, the tips ferrugineous-tomentose; terminal bud 1/2-1 cm long; pith without vascular strands. Leaves 1—4-jugate. Petioles 4—9 (-14) cm, slightly to strongly flattened at the base; pith with few vascular strands. Leaflets oblong to ovate (to suborbicular), 51/2-17 (-22) by 3—71/2 (-101/2) cm, upper surface greenish when dried, lower surface pubescent to glabrous, indumen- tum partly or entirely consisting of minute stellate hairs; base broadly cuneate to rounded; apex subabruptly, bluntly acuminate; nervation § rather prominent beneath; nerves 10-15 (—18) per side, more or less curved, mostly not distinctly looped and joined except towards base and apex; reticulations minute, hardly or not conspicuous above. Panicles axillary, often on short, leafless lateral shoots with a terminal bud, narrow, 11/2—51/2 (—20) cm long, ferrugi- neous-tomentose; peduncle up to 10 cm long. Flowers c. 2 mm long, sessile to shortly pedicelled, glabrous or stellate- tomentose and glabrescent. Calyx 11/2 mm. Petals outside glabrous or hairy. Stamens free from the disk. Disk thick- annular or (2) cupular, radially furrowed and with undulate rim. Pistil in 3 flowers moderately reduced. /nfructescences up to c. 5 cm long, ferrugineous-tomentose, with a few fruits. Fruits ellipsoid, immature ones c. 11/2 by 0,9 cm.
Distr. Malesia: Sumatra, Banka, Malay Peninsula, and Borneo.
Ecol. Primary and secondary forests, up to 60 (-750) m. F/. Febr.—March, June, Oct.
Vern. Bantan burung, kedondong tund- juk, sisip baniéng, Sum., asam-asam, Banka, kedudong, Mal. Pen., engai, mendjelih, selada, Borneo.
Note. Originally, KALKMAN already inclined towards inclusion of this species in the genus Dacryodes, as appears from his identification labels, but finally he decided to leave it in Santiria by lack of evidence. Only when fruiting material became available the generic identity became sufficiently clear.
Santiria Bu. Replace the Key to the species by the following one:
1. Petiolules 3-3/4 cm long.6. S. ridleyi 1. Petiolules up to 3 cm long.
2. Bract-like cataphylls between the leaves present. Leaves (60—)80—135 cm long . . 10. S. megaphylla
2. No bract-like cataphylls between the leaves. Leaves rarely more than 60 (up to 85) cm long.
3. Anthers adnate (mostly also visible under the fruit).
4. Pith of branchlets with vascular
strands. Flowers 4-10 mm long, calyx 3-7 mm high, ¢ flower with 6 fertile stamens. Stigma on fruit c. 90° excentric . 11. S. griffithii . Pith of branchlets without vascu- lar strands. Flowers 2-3 mm long, calyx 1-21/2 mm high, ¢ flower with 3 fertile stamens and some- times up to 3 staminodes. Stigma
on fruit less than 90° excentric. 12. S. rubiginosa
3. Anthers basi- to dorsifixed. 5. Calyx at anthesis 1!/2-3 mm high.
6. Pith of branchlets with rather many vascular strands. 2. S. mollis 6. Pith of branchlets without vascu- lar strands.
7. Terminal bud 2-3 cm _ long. Flowers 4-4!/2 mm long, calyx 21/2-3 mm high.
3. S. grandiflora
7. Terminal bud up to 2 cm long. Flowers 2-4 mm, calyx 1-2 mm high.
8. Terminal bud 1-2 cm long. Stigma on fruit less than 90° excentric . 5. S. oblongifolia
8. Terminal bud 14/2-1 cm iong. Stigma on fruit usually more then 90° excentric.
9. S. apiculata
5. Calyx at anthesis up to 11/2 mm
high.
9. Pith of branchlets with vascular strands.
10. Flowers (3) 3-4 mm, calyx 11/2-21/2 mm high. 2. S. mollis
10. Flowers 2 mm, calyx 1/2—3/4 mmhigh . . 4. S. laevigata
9. Pith of branchlets without vascu- lar strands.
11. Mature leaves beneath hairy at least on midrib and nerves, mostly also on the veins.
12. Calyx in anthesis 1/2-1 mm high. Stigma on fruits less than 90° excentric.
1. S. tomentosa
12. Calyx in anthesis 1-11/2 mm high. Stigma on fruits 90° or more excentric.
9. S. apiculata
11. Mature leaves beneath glab- rous or only hairy on midrib.
13. Calyx during anthesis 1/2—3/4
mm high. 14. Stigma on the fruits up to 90° excentric . 4. S. laevigata
14. Stigma on fruits more than 90° excentric . 7. S. conferta
13. Calyx during anthesis 1-2 mm high.
15. Terminal bud 1-2 cm long.
September 1972]
Addenda, corrigenda et emendanda
O20
So231la
Si 23 5223240
: 232b 2334
nn
530233b
5: 234b
s256a
5: 236b
Stigma on fruit less than 90° excentric. 5. S. oblongifolia 15. Terminal bud !/2-1 cm long. Stigma on fruit usually more than 90° excentric. 9. S. apiculata Santiria tomentosa BL. Add to literature: ANDERSON, Bull. Sing. 20 (1963) 165. In description, line 4, add after ‘pubes- cent’: to tomentose. Add under petioles, after ‘flattened’: or channelled. Add to Distr.: Philippines (Mindanao). Add to Ecol.: also in secondary forests. Santiria grandiflora KALKMAN. Add to Distr.: several new collections from Brunei. Santiria laevigata BL. Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 164. Add to description: Branchlets thin-hairy when young. Leaflets beneath sometimes sparsely hairy on midrib and nerves; base not rarely somewhat oblique; apex acutely to bluntly short-acuminate.
Gard.
Add to Distr.: Philippines (Mindanao). .
Santiria oblongifolia Bu.
Add to description: Buttresses up to 2m high. Petioles sometimes narrowly grooved at base. Fruits red when ripe.
Santiria conferta BENN.
Change in description: Tree, 4-35 m by up to 70 cm @&, withup to 1!/2 m high buttresses. Infructescences to 30 cm long. Fruits to 13/4 cm long, stigma lateral to near the pedicel.
Add to Distr.: N. Borneo, at 1500-1800 m alt.
Delete 8. Santiria nervosa H. J. Lam. Santiria apiculata BENN.
In description add at end of sentence on calyx: to sepals less than 1/2 connate. Petals inside glabrous or sparsely hairy. Add under fruits: red when ripe.
Insert before the Key to the varieties: Note. A great part of the material can easily be subdivided into the following three varieties; however, some specimens show characters of more than one variety. var. apiculata.
Change Ecol. highest alt. into 1500 m. var. rubra (RIDL.) KALKMAN.
Add. to Ecol.: Primary and secondary forests up to 1600 m.
Santiria megaphylla KALKMAN.
In description, add at end of Ist sentence: by 45 cm @. Change in line 2 ‘11/2’ into: 1, and in line 4 ‘5S’ into: 8.
Line 8, after ‘part’ add: to all looped and joined. Delete after panicles “(3 un- known)’, the same after flowers. Insert before ‘Infructescences’: Pistillode in 3 flowers minute.
Add to Distr.: Brunei.
5: 236a 5: 236b 59 2376 5: 246 58 WA 5: 247a 5: 2476 5: 2476, 567ab
SeeZoil
Add to Ecol.: In primary Dipterocarp forest on damp to swampy, shallow clay soil, 0-150 m.
Santiria griffithti (HOOK. f.) ENGL.
Add to synonymy: Amoora aphanamixis Auct. non R. & S.: Mia. Sum. (1861) 196. Change in description: Leaves excep- tionally to 15-jugate. Petioles sometimes strongly flattened at base. Leaflets exceptionally also hairy on midrib above and on veins beneath.
Calyx (in vivo) olive to red. Petals (id.) yellowish-white.
Change in Ecol. highest alt. into 700 m. Add to Notes: The collections For. Dept. Sarawak 12745 and 15613 repre- sent a strongly pubescent form with flattened petioles.
Santiria rubiginosa BL.
Change in description: Tree not always buttressed. Branchlets mostly glabrous. Add to Distr., under New Guinea: Vogelkop Peninsula.
var. rubiginosa.
Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 164.
Scutinanthe THw.
Line 2 from bottom, delete ‘pilose’. Key: A further difference between the two species is in the fruits, these being densely pubescent in S. brunnea, glabrous in S. brevisepala.
Scutinanthe brunnea THW.
Add to literature: WyYATT-SMITH & KOcHUMMEN, Mal. For. Rec. 17, rev. ed. (1965) 348.
In description, add: Sometimes the leaves are fully glabrous. Change: Fruits finally glabrescent, yellow.
Add to Ecol.: In Sarawak at c. 800 m, in Sabah at c. 1200 m alt.
Scutinanthe brevisepala LEENH.
Add to literature and synonymy: LEENH. Blumea 12 (1964) 19. Dacryodes papuana Husson, Blumea 7 (1952) 167, f. 1; Leenu. FI. Mal. I, 5 (1956) 228. Add to description: Tree up to 35 m high, 43-55 cm @, sometimes buttressed up to 3 m. Branchlets not always con- spicuously lenticellate. Leaflets lanceolate to broad-elliptic or subovate, up to 10 cm wide, coriaceous or pergamentaceous to chartaceous, the base broadly cuneate to subcordate, mostly slightly oblique. Inflorescences axillary on short axillary shoots the vegetative terminal bud of which usually develops later on. Ovary (2-)3-celled. Mature fruits — slightly oblique, ovoid to ellipsoid, constricted or shortly stalked at the base, pointed at the apex, 21/4-3 by 1!/2-13/4 cm, glabrous. Seed 1.
Canarium STICKM.
Add to Distr.: The genus can be sub- divided into 3 subgenera: subg. Canarium,
922
FLORA MALESIANA
Sie2525 SS,
comprising the sections Canarium and
Pimela; subg. Africanarium LEENH. nov.
stat. (Canarium sect. Africanarium LEENH.
Blumea 13, 1966, 396), monotypic,
W. Africa; subg. Canariellum.
In Key to the species, add to couplet 5,
2nd lead: (in C. album sometimes
papillose, then stamens connate halfway up or more, pistil pilose, and fruits white when ripe).
Replace couplets 14 to 19 incl. by the
following:
14. Stipules fugacious, present only in the terminal bud, even the scars nearly invisible.
15. Leaflets equal-sided at base. Pith of branchlets always with central vascular strands. Fruit very peculiar (see fig. 21k, in vol. 5), 4-41/2 by 21/9-23/4 by c. 11/2 cm.
48. C. cestracion
15. Leaflets oblique at base. Pith of branchlets mostly without central vascular strands. Fruit ovoid to spindle-shaped, round in cross- section, 21/2-31/2 by 11/2-2 cm.
53. C. album
14. Stipules persistent to caducous, scars well visible.
16. Infructescences (sub)spicate, often with many fruits; fruits ovoid to subglobose, rather small (9-14 by 4-11 mm) 51. C. asperum
16. Infructescences racemoid to thyr- soid, mostly with few fruits; fruits mostly relatively longer and always bigger.
17. Vascular strands in pith of branch- lets all peripherally arranged; twigs long remaining densely hairy (rarely, only one of these charac- ters holds good).
18. Indumentum pilose. Flowers long and slender, corolla more than two times as long as the calyx; filaments in 2 flowers nearly completely connate. Fruits usual- ly prismatic, blunt 3-angular in cross-section, at apex mostly truncate and ‘shouldered’, (sub)- glabrous.
33. C. pilosum ssp. pilosum
18. Indumentum tomentose to velve- ty. Corolla less than two times as long as the calyx; filaments in © flowers free. Fruits ellipsoid, usually velvety.
49. C. vrieseanum
17. Vascular strands in pith of branch- lets only partly peripherally ar- ranged; twigs soon glabrescent.
19. Stipules inserted on the petiole up to 3 cm from its base. Fruits fusiform, 4 by 2 cm.
52. C. vitiense
[ser. I, vol. 6®
19. Stipules inserted on the twig at the base of the petiole. Fruits ellipsoid, up to 31/2 by 2 cm.
19A. Leaflets rounded at base.
Fruits 3-31/2 by 114/2-2 cm,
pyrene smooth, sterile cells moderately reduced.
44. C. macadamii
19A. Leaflets cuneate at base. Fruits
2 by 3/4 cm, pyrene irregularly
grooved, sterile cells nearly completely reduced.
45. C. chinare
5: 254, Replace couplets 44 to 50 incl. by the 255 following:
44. Inflorescences axillary. 45. Stipules rather caducous, roundish, + herbaceous 17. C. luzonicum 45. Stipules persistent, linguiform, stiff- coriaceous 18. C. ovatum 44. Inflorescences terminal (lower branches often in the upper leaf axils). 46. ¢ Flowers with 3 stamens.
47. Vascular strands in pith of branch- lets all peripheral. Stipules cadu- cous, c.2mm @.
5. C. caudatum f. caudatum
47. Vascular strands in pith of branch- lets partly central. Stipules sub- persistent, c. 15 by 10 mm.
54. C. reniforme 46. 3 Flowers with 6 stamens.
48. Stipules (rather) persistent, attach- ed on the petiole 1/2-11/2 cm from its base . : . «) 10. €. lamii
48. Stipules caducous, mostly attach- ed on the twig at or partly, excep- tionally fully, on the base of the petiole.
49. Stipules oblong, 1-5 by !/2-12/4 cm; scar linear, c. 1/2 cm long.
14. C. vulgare
49. Stipules reniformous, much smal-
ler; scar elliptic to drop-shaped,
1—2 mm long.
50. 3 Inflorescences very lax, bran-
ches long and patent. J Flowers
13 mm long. 6. C. divergens
50. 3 Inflorescences not very lax.
3 Flowers c. 8 mm long.
50A. Veins and reticulations more
or less prominulous and
well-visible on the under
surface of the leaflet; nerves
11-15 pairs, mutual distance
along the midrib usually less
than 1 cm 1. C. littorale
50A. Veins and reticulations nearly
invisible in dried specimens;
nerves 5-10-15 pairs, mutual
distance along midrib 1-11/e
cm . 4. C. patentinervium
5: 255 Replace 63 Ist lead and couplet 64 by the
following:
September 1972]
Addenda, corrigenda et emendanda
a 250d
5: 258a
5: 258b
5: 259a
Be 2590
5: 260a
5: 260b
63. Filaments at least nearly halfway connate.
64. Branchlets long remaining densely ferruginous-woolly. Leaves up to 8-jugate; nerves 14-17 pairs. Fruit- ing calyx c.5mm @. 40. C. kipella Branchlets glabrous or puberulous at the tip only. Leaves up to 6-jugate; nerves mostly less than 14 pairs. Fruiting calyx 8-11 mm @. 64A. Leaflets lanceolate. Pith of branchlets always with central vascular strands.
39. C. intermedium Leaflets (oblong to) broad-ellip- tic (to suborbicular). Pith of branchlets mostly without central vascular strands . 55. C. pimela Canarium littorale Bv. Add to literature: LEENH. Blumea 9 (1959) 337; Back. & BAKH. f. Fl. Java 2 (1965) 115; Meer, Bot. Bull. Herb. Sandakan 11 (1968) plate between p. 111 and 112 (seedling). Add in Notes to the area of f. pruinosum (ENGL.) LEENH.: Brunei and Sabah. Canarium latistipulatum RIDL. Add to literature: LEENH. Blumea 9
64.
64A.
(1959) 341. Add to description: Fruits spindle- shaped, -+ round in_ cross-section,
7 by 24/4 cm, glabrous; pyrene smooth, rounded triangular in cross-section, the lids intruded, lids c. 3-4 mm thick. Seeds 2, sterile cell moderately reduced. Canarium perlisanum LEENH.
Add to literature: LEENH. Blumea 9 (1959) 342.
Canarium patentinervium MIQ.
Add to literature: LEENH. Blumea 9 (1959) 342, f. 15.
Add to description: buttressed. /nfructescences glabrescent.
Add to Ecol.: also in secondary and swamp forests.
Tree sometimes sometimes
Canarium caudatum KING.
Add to literature: LEENH. Blumea 9 (1959) 343.
Add to Notes sub f. auriculiferum
LEENH.: also known from the Malay Peninsula.
Canarium divergens ENGL.
Add to literature: LEENH. Blumea 9 (1959) 346.
Canarium kinabaluense LEENH.
Add to literature: LEENH. Blumea 9 (1959) 346.
Add to description: Jnflorescences termi- nal, laxly thyrsoid, c. 4-6 cm long, few-flowered, minutely tomentose, gla- brescent. Flowers (2) 1 cm long. Calyx 5 mm, minutely tomentose. Stamens 6, glabrous, inserted on the rim of the disk. Disk glabrous, adnate to the receptacle.
Si
5; 2615;
an
26la
567a
: 262a : 262b
: 263a
923 Pistil glabrous. Add to Distr.: East Borneo. Add to Ecol.: down to 450 m. Canarium maluense LAUT. Add to literature: LEENH. Blumea 9
(1959) 347.
ssp. maluense.
Add to description, Ist sentence: up to 60 m high, with up to 11/2 m high buttresses.
Add to Distr.: Louisiade Arch.
Add: Uses. Timber for construction. Canarium megacarpum LEENH. Add to literature: LEENH. (1959) 351.
Canarium lamii LEENH.
Add to literature: LEENH. Blumea 9 (1959) 351.
The following changes should be made in the description: Tree up to 42 m by 66 cm, sometimes with buttresses and stilt- roots. Leaves 3-4-jugate. Stipules sub- persistent or more or less caducous, inserted 1/2-11/2 cm from the base of the petiole. Leaflets up to 22 by 10 cm, base rounded to cordate; nerves 10—15 pairs. Inflorescences (2) remotely spicate, densely tomentose. Flowers: old 2 known. Calyx 7 mm high, the lobes 11/2 mm; outside tomentose, inside appressed short-hairy, densest near base and margin. Corolla: petals 71/2 by 5 mm, outside densely appressed short-hairy in the upper half mainly along the midrib, inside glabrous, in vivo orange to red. Staminodes 6, inserted on the disk, c. 31/24 mm long, glabrous. Disk adnate to the hollowed receptacle, free rim 1/5-3/4 mm _ high, fleshy, glabrous. Pistil glabrous; ovary 3 mm; style 11/4 mm, thick; stigma globular, 1 mm @. Infructescences with 1 or few fruits, rusty tomentose; calyx to 2!/2 cm @, densely rusty tomentose when young. Add: Ecol. Primary and _ secondary forests up to c. 1250 m.
Canarium sylvestre GAERTN.
Add to literature: LEENH. Blumea 9 (1959) 352.
Add to description, Ist sentence: some- times with stiltroots.
Add to Ecol.: also in secondary forests. Canarium piloso-sylvestre LEENH.
Add to literature: LEENH. Blumea 9 (1959) 353, f. 16.
Add to description: Acumen of leaflets blunt to acute. Calyx 21/24 mm high, outside sometimes slightly pubescent towards the apex. Disk 3—6-lobed.
Add to Ecol.: In seasonally inundated primary forest.
Canarium salomonense B. L. BURTT. Add to literature: LEENH. Blumea 9 (1959) 353.
Change in description: Tree up to c.
Blumea 9
924 FLORA MALESIANA [ser. I, vol. 6° 40 m by 80 cm, mostly with buttresses. LANESSAN, PI. Utiles Col. Frang. (1886) Canarium vulgare LEENH. 309, nom. nud.
Add to literature: LEENH. Blumea 9 Add to description: Pith of branchlets (1959) 358; Back. & BAKH. f. Fl. Java 2 dark- to light-brown, either with peri- (1965) 115. pherally arranged, or with scattered
5: 265a Add to Distr.: New Guinea. vascular strands.
5: 265b Add to Ecol.: Alt. up to 1200 m. Add to Distr.: Borneo, Sabah (Beaufort
5: 2665 Canarium indicum L. Distr., Ulu Lumat, SAN 44543).
Add to literature: LEENH. Blumea 9 Add to Ecol.: Primary forest. (1959) 359; Back. & BAKH. f. Fl. Java 2 Canarium apertum H. J. LAM. (1965) 115. Add to literature: LEENH. Blumea 9
5: 269b Canarium kaniense LAUT. (1959) 386.
Add to literature: LEENH. Blumea 9 : 2756 Incertae sedis: The systematic position of (1959) 362. C. pseudodecumanum and C. decumanum
5: 270b Canarium luzonicum (BL.) A. GRAY. remains uncertain, even with growing Add to literature: LEENH. Blumea 9 knowledge. The blastogeny is in full (1959) 363. accordance with sect. Canarium, the Add to Distr.: Mindanao. ontogeny of the stipules of C. decumanum
5: 27la Canarium ovatum ENGL. with sect. Pimela, however.
Add to literature: LEENH. Blumea 9 : 276a Canarium pseudodecumanum HOcnrR. (1959) 364. Add to literature: LEENH. Blumea 9
5: 2716 Canarium odontophyllum Mia. (1959) 388: Meter, Bot. Bull. Herb. Add to literature: LEENH. Blumea 9 Sandakan 11 (1968) plate between p. 111 (1959) 365. and 112 (seedling).
5: 272a Add to Uses: In Sarawak grown for its : 277a Canarium decumanum GAERTN. fruits. Add to literature: DouGLas & BAAS Canarium denticulatum BL. BECKING, Bull. Bot. Gard. Btzg III, 17 Add to literature: LEENH. Blumea 9 (1947) 295-296, t. 11; LEENH. Blumea 9 (1959) 367; BAck. & BAKH. f. FI. Java 2 (1959) 389: BAcK. & BAKH. f. FI. Java 2 (1965) 115. (1965) 114; Meer, Bot. Bull. Herb.
5: 274a ssp. kostermansii LEENH. Sandakan 10 (1968) plate between p. 138 Add to Distr.: Sabah. and 139.
Canarium karoense H. J. LAM. Add to Distr. under New Guinea: Add to literature: LEENH. Blumea 9 Japen I.
(1959) 370. : 278a Canarium oleosum (LAMK) ENGL. Canarium megalanthum MERR. Add to literature: LEENH. Blumea 9 Add to literature: LEENH. Blumea 9 (1959) 391.
(1959) 370, f. 17; Meer, Bot. Bull. Add to description, Ist sentence: Tree Herb. Sandakan 11 (1968) plate between up to 50 m high.
p. 111 and 112 (seedling). : 27865 Add to Distr. under Lesser Sunda Is.: Add to description: Sometimes with up Sumbawa.
to 1 m high buttresses. Stipules sometimes : 279a Canarium balsamiferum WILLD. caducous, sometimes inserted on the base Add to literature: LEENH. Blumea 9 of the petiole or even slightly on the twig. (1959) 392.
Leaflets often chartaceous, base some- : 279b Add to Distr.: Lesser Sunda Is. (Sum- times nearly equal-sided, margin some- bawa).
times serrulate near the apex. : 280a Canarium trigonum H. J. LAM.
5: 274b Canarium pseudopatentinervium H. J. Add to literature: LEENH. Blumea 9
LAM. (1959) 393.
Add to literature: LEENH. Blumea 9 Canarium euryphyllum PERK.
(1959) 385. Add to literature: LEENH. Blumea 9
Add to description: Buttresses sometimes (1959) 394.
present. Leaflets: apex shortly blunt- to : 280b Add to description: Corolla reported
acute-acuminate; nerves not to distinctly to be yellowish red.
looped and joined. 3 Inflorescences with : 28la, Canarium kostermansii LEENH.
up to 10 cm long lower branches, these 567b Add to literature: LEENH. Blumea 9
as well as the main axis laxly set with (1959) 398, f. 23.
subsessile glomerules of flowers. 3 Flowers Add to description: Leaflets: acumen
3-4 mm long pedicelled. Disk in 3 acute to bluntish; nerves 20-25 pairs.
flowers cushion-shaped, c. 1 mm high, Add: Ecol. Primary forest.
densely hairy, without rudimentary : 2816 Canarium pilosum BENN.
pistil. Add to literature: LEENH. Blumea 9 5: 275a Canarium grandifolium (RIDL.) H. J. LAM. (1959) 398.
Add to literature: LEENH. Blumea 9 (1959) 386, f. 21, non BAILLON ex DE
: 282a
ssp. borneensis LEENH. Add to Distr.: Sarawak (G. LAMBIR).
September 1972]
Addenda, corrigenda et emendanda
5: 282b
5: 283a
5: 2836
5: 284b
5: 285a
Si 285b
5: 286a
5: 286b
5: 287a
5: 289a
5: 289b
Add to Ecol.: from the lowland up to c. 1500 m. Canarium merrillii H. J. LAM.
Add to literature: LEENH. Blumea 9 (1959) 402.
Canarium gracile ENGL.
Add to literature: LEENH. Blumea 9
(1959) 396, f. 22.
Add to description: The flowers are said to be purplish, the fruits first red, when ripe black.
Canarium dichotomum (BL.) Mia.
Add to literature: LEENH. Blumea 9 (1959) 423.
Add to description: Leaflets sometimes beneath rather densely appressed short- hairy; acumen sometimes slender and acute. 2 Inflorescences up to 35 cm long. Add to Ecol.: Sometimes also in secon- dary forests. Alt. up to 1000 m. Canarium fusco-calycinum RIDL.
Add to literature: LEENH. Blumea 9 (1959) 424.
Add to description: Nerves geniculate or looping near the margin.
Add to Ecol.: Primary lowland Diptero- carp forest at 100 m.
Canarium australianum F.v.M.
Add to literature: LEENH. Blumea 9 (1959) 421; SpecHT, Rec. Am. Austr. Exp. Arnhem Land 3 (1958) 460.
Add to description: Leaflets sometimes oblong.
Add to Distr. under Australia: northern part of W. Australia. Canarium intermedium H. J. LAM. Add to literature: LEENH. Blumea 9 (1959) 419, f. 29.
Canarium kipella (BL.) Mia.
Add to literature: LEENH. Blumea 9 (1959) 419; Back. & BAKH. f. FI. Java 2 (1965) 115.
Canarium pseudosumatranum LEENH. Add to literature: LEENH. Blumea 9 (1959) 431.
Change in description: Leaves 6—9-juga- te; leaflets 7-20 by 3-6 cm. /nflorescences 20-30 cm long.
Canarium sumatranum BOERL. & KoorD. Add to literature: LEENH. Blumea 9 (1959) 432, f. 31.
Add to description: Leaflets up to 23 cm long, rough above; nerves up to more than 30 pairs. 2 Inflorescences \ike the 3 ones. 2 Flowers 6 mm long, the calyx 21/2 mm.
Canarium hirsutum WILLD.
Add to literature: LEENH. Blumea 9 (1959) 424, f. 30; Back. & BAKH. f. Fl. Java 2 (1965) 115.
the
Canarium hirsutum WILLD. var. hirsu- tum.
Add to Distr.: Solomon Is.
Canarium macadamii LEENH. — Fig. 2.
Add to literature: LEENH. Blumea 9
Fig. 2. Canarium macadamii LEENH. a. Staminodes and disk of & flower from outisde, b. & c. stamens and disk of 3 flower from outisde (6) and on longitudinal section (c: mind the tiny pistillode),
all »
5: 290a
5: 2906
6 (a NGF 21682, b-c MACADAM 206).
(1959) 448.
Add to description: Tree 30-40 by 3/,-1 m. Leaflets 10-20 cm _ long. 2 Inflorescences 8-10 cm. ¥% Flowers
7 mm; calyx 4 mm; petals outside sparsely hairy; staminodes confluent with the disk; disk glabrous, protruded into 6 tongue-shaped, fleshy, 2 mm long lobes alternating with and nearly equal- ling staminodes. Infructescences 8-15 cm long; calyx 5-7 mm @.
Add to Ecol.: Alt. 1100-1500 m. FI. Febr.
Add: Uses. Seeds edible.
Canarium chinare GRUTTERINK & H. J.
LAM. Add to literature: LEENH. Blumea 9 (1959) 449.
Canarium rigidum (BL.) Mia.
Add to literature and synonymy: LEENH. Blumea 9 (1959) 434, f. 32. — C. poly- phyllum K. Scu. in K. Sch. & Hollr. Fl. Kais. Wilh. Land (1889) 63; LEENH. Fl. Mal. I, 5 (1956) 290, f. 20 d; Blumea 9 (1959) 434, f. 33.
Add to description: Tree up to 30 m by 40-45 cm. 2 Inflorescences 8-9 cm long, the partial inflorescences c. 2 cm long, with 1 or 2 flowers. 2 Flowers 71/2 mm long, the calyx 4!/2 mm high, outside densely appressed pubescent; staminodes free outside the disk, 2 mm long; disk faintly lobed, c. 1 mm high, long-hairy towards the margin; pistil glabrous. Add: Uses. Reported to be used for buildings proas.
Canarium polyphyllum K. SCH.
With new collections that became avail- able it became more and more clear that the present species and C. rigidum repre- sented only extremes as to hairiness, leaf ratio, and sculpture of fruit kernel, but could not well be demarcated. Hence, C. polyphyllum had to be reduced to C. rigidum.
Canarium cestracion LEENH.
Add to literature: LEENH. Blumea 9 (1959) 436.
Change in description: Vascular strands in pith of branchlets may also be mainly peripherally arranged. Infructescences
926
FLORA MALESIANA
[ser. I, vol. 6%
Se 29a
55 29 0D
293a
5: 295b
5: 296a
5: 2965
5: 2966
3-20 cm long. Fruits, if 1-seeded, much flattened in cross-section.
Add to Distr.: A second collection (NGF 25654) from Morobe Distr., at 30 m alt.
Canarium vrieseanum ENGL.
Add to literature: LEENH. Blumea 9 (1959) 437.
Canarium acutifolium (DC.) MERR.
Add to literature: LEENH. Blumea 9 (1959) 445.
var. acutifolium.
Change in description: Tree 13-45 m. Leaflets up to 40 by 15 cm. Fruiting calyx up to 74/2 mm @. Fruits up to 171/2 by 121/2 (1-seeded) or 171/2 (2-seeded) mm. Seeds exceptionally 2.
Add to Distr.: New Britain.
Canarium asperum BENTH.
Add to literature: LEENH. Blumea 9 (1959) 439; Back. & BAKH. /. Fl. Java 2 (1965) 115.
SSP. aSperum var. asperum.
Add to Uses: On wood see SCHNEIDER, Bull. Bur. For. Philip. 14 (1916) 129. Replace the heading of 52. Canarium schlechteri Laut. by the following:
52. Canarium vitiense A. Gray, U.S. Expl. Exp. Bot. 1 (1854) 373; LEENH. Blumea 9 (1959) 451; Blumea 13 (1965) 166. — C. samoense ENGL. in DC. Mon. Phan. 4 (1883) 134; LEENH. Blumea 9 (1959) 452. — C. schlechteri LAvuT. Bot. Jahrb. 56 (1920) 328; LEENH. FI. Mal. I, 5 (1956) 296, f. 20 e; Blumea 9 (1959) 444. — C. smithii LEENH. Bish. Mus. Bull. 216 (1955) 12, f. 6; Blumea 9 (1959) 450. — C. hacciferum LEENH. Bish. Mus. Bull. 216 (1955) 19, f. 9. Add to de- scription: 2 Inflorescences racemose, 6—9 cm long, tomentose. 2 Flowers 1 cm long, less slender than ¢ ones, subsessile; calyx 4 mm; staminodes 3 mm, connate for less than 1 mm; disk annular, low, long-ciliate; pistil 4 mm, glabrous, ovary tapering into the style.
Replace Distr. by: Malesia: eastern half of New Guinea, Louisiade Arch., Admi- ralty Is., Bismarck Arch., Solomons, Fiji, Samoa, and Tonga.
After species 52 the following species should be added:
53. Canarium album (Lour.) RAEUSCH. Nomencl. ed. 3 (1797) 287; Hance, J. Bot. 9 (1871) 38; GUILLAUMIN, Bull. Soc. Bot. Fr. 55 (1908) 617, t. 19 f. 1; HAYATA, J. Coll. Sc. Imp. Univ. Tokyo 30 (1911) 52; Merr. Int. Rumph. (1917) 304; WALKER, Imp. Trees Ryukyu (1954) 148, f. 82; LEENH. Blumea 9 (1959) 402, f. 24; BARANOV, Quart. J. Taiw. Mus. 20 (1967) 367, cum fig.; non BLCo, FI. Filip. (1837) 793 (= C. luzonicum A. GRAY). — ? C. sinense Cana Rumen. Herb. Amb. 2
(1741) 154. — Pimela alba Lour. FI. Coch. (1790) 408.
Tree, up to 30m by 1!/2 m &. Branch- lets 5-6 mm @&, the young parts fulvous tomentose, older parts sometimes gnarly by the strongly prominent scars of leaves and inflorescences; pith with a peripheral cylinder of small vascular strands, rarely moreover some strands in the central part. Leaves 3—6-jugate. Stipules present in the bud only, inserted on the twig next to the petiole, even the scar usually nearly invisible. Leaflets usually distinctly sinu- ous, especially the basal pairs, lanceolate or elliptic (to ovate), 61/2-14 by 2-5!/s cm, chartaceous to coriaceous, glabrous or with some scattered bristles on the nerves below, lower side often minutely verrucose; base oblique, cuneate to rounded; margin entire; apex tapering to subabruptly acuminate, acumen up to 2 cm long, blunt; nerves 12-16 pairs, angle to midrib 65-75”, straight to faintly curved in the broader, strongly curved in the narrower side of the leaflet, more or less distinctly looped and joined near the margin; intercalary veins sometimes distinctly developed; reticulation dense. Inflorescences axillary, tomentellous to glabrous, 3 thyrsoid, 15-30 cm long, many-flowered, 2 racemoid, 3-6 cm long, with up to 12 flowers. Flowers sparsely tomentose to glabrous, 3 51/2-8 mm long, 2 c. 7 mm. Calyx 21/2-3 mm, in 2 flowers subtruncate. Stamens 6, glabrous, the filaments more than halfway (in 2 flowers up to nearly completely) connate. Disk 3 globular to cylindrical, 1-11/2 mm high, slightly 6-lobed, solid or with a central canal, the upper side with some bristles; in 2 flowers annular, faintly 3-lobed, 1 mm high, thick and fleshy, pilose on the inner surface. Pistil densely short-pilose, in flowers minute or none. Infructescences up to 15 cm long with up to 6 fruits; calyx flat, 3-lobed, 1/5 cm @, the lobes recurved. Fruits ovoid to spindle-shaped, round in cross-section, 21/2-31/2 by 11/2-2 cm, glabrous, in vivo white when ripe; pericarp rather thick; pyrene acuminate, rounded (to 6-angular) in cross-section, with a distinct groove between the blunt angle-ribs and the lids, the latter with a faint median rib, surface furthermore slightly undulated; lids 11/2-2 (3) mm thick. Seeds 1 or 2, the sterile cell(s) slightly reduced, round in cross-section.
Distr. Annam (from about 16° N northwards), Tonkin, southern China (up to about 27° N), and Hainan; as thespecies is much cultivated, mainly in the same region, it is sometimes difficult to decide whether in some part of the area it is wild or naturalized, or even planted. In
September 1972]
Addenda, corrigenda et emendanda 927
Malesia: Sumatra, East Coast (Medan, planted in and naturalized near a park: LORZING 16519, 17240).
Ecol. In light to dense forests on dry to moist soils, usually at medium altitude (400-1200 m).
Uses. Especially in SE. China common- ly planted for ornament and as a fruit tree. The fruits, of which the pulp as well as the seeds are eaten, prepared in several ways, are highly esteemed by the Chinese. Furthermore, the wood and the resin are sometimes used.
Note. C.albumbelongs to sect. Pimela, to the relationship of C. pilosum, and seems to be nearest to C. pimela.
54. Canarium reniforme KOCHUMMEN & Wuitmore, Gard. Bull. Sing. 24 (1969) 2.
Tree, up to 18 m by 30cm @. Branchlets 5-7 mm @, long remaining fulvous- tomentellous; pith with many vascular strands, partly peripheral. Leaves 2-4-ju- gate. Stipules attached on the base of the petiole, mostly partly on the twig, oblong to reniform, 15-25 by c. 10 mm, rather stiff, persistent. Leaflets ovate to ovate- oblong or elliptic, 31/2-16 by 11/2-10 cm, stiff-coriaceous, beneath thinly puberu- lous, further glabrous; base of laterals slightly oblique, cuneate to rounded, margin entire, apex rather abruptly, bluntly to acutely acuminate; nerves 7-14 per side, angle to midrib c. 60-70°, nearly straight to curved, looped and joined near the margin, veins and vein- lets much more slender, but well-visible on both sides. Inflorescences terminal, c. 25 cm long. Flowers: 2 unknown. Calyx 5 mm high, outside puberulous, inside glabrous. Petals outside hairy in the central part. Stamens 3, adnate to the disk. Disk solid, globular, tapering into a ‘style’, glabrous. J/nfructescence c. 10 cm long, glabrous; calyx flat, orbic- ular, 1-11/2 cm ©, with inside an annu- lar, not-lobed, glabrous disk. Fruits ovoid to ellipsoid, 5—51/2 by c. 23/4 cm, in cross-section blunt-triangular; pyrene rather smooth; lids 3-4 mm thick. Seed 1, the fertile cell orbicular in cross-section, the sterile ones nearly completely redu- ced.
Distr. Malesia: the Malay Peninsula (Perak).
Ecol. Understorey of primary hill- forest at c. 300 m alt. Fr. June.
Note. C. reniforme belongs to sect. Canarium and is closely allied to C. patentinervium and C. caudatum. With the latter it shares the number of 3 stamens, with the former the general habit, apart from the peculiar stipules.
55. Canarium pimela LEENH. Blumea 9 (1959) 406, f. 25. — ? C. sinense Tsjacana
RumpuH. Herb. Amb. 2 (1741) 154. — Pimela nigra Lour. Fl. Coch. (1790) 407. — C. pimela Koen. Ann. Bot. 1 (1805) 361, t. 7, f. 1, nom. illeg.; HANCE, J. Bot. 9 (1871) 38; Fors. & HEMSL. J. Linn. Soc. Bot. 23 (1886) 113; Merr. Int. Rumph. (1917) 304; Merr. & CHUN, Sunyatsenia 2 (1935) 253; Merr. Comm. Lour. (1935) 227; non BL. Bijdr. (1826) 1162 (= C. kipella), nec SPAN. Hook. Comp. Bot. Mag. 1 (1835) 346 (= C. oleosum), nec Bico, FI. Filip. (1845) 545 (= C. asperum).
Tree, up to 30m by 11/2m @. Branch- lets 7-10 mm 2, glabrous; pith with a peripheral cylinder of vascular strands and sometimes a few in the central part. Leaves 4-6-jugate, glabrous. No stipules. Leaflets oblique, often distinctly falcate, broad-elliptic (to ovate or suborbicular, rarely oblong), 6-17 by 2—71/2 cm, charta- ceous to coriaceous; base acute, often decurrent; margin entire; apex rather abruptly acuminate, acumen _ short, broad, and blunt; nerves (8—) 11 (—15) pairs, angle to midrib 70—75°, straight to faintly curved, looped and + joined near the margin; veins and veinlets coarsely reticulate. Inflorescences axillary, glabrous, laxly thyrsoid (¢) to race- moid (2), 15-40 cm long, ¢ many-, © few-flowered. Flowers (sub)glabrous,
3S 7 mm long, slender, 2 9 mm long.
Calyx in 3 flowers 21/2 mm, distinctly lobed, in 2 flowers 31/24 mm, subtrun- cate. Corolla in 2 buds characteristic- ally conical. Stamens 6, glabrous (except 2 rows of bristles on the anthers in 2 flowers), in dj flowers nearly halfway, in 2 flowers slightly more than halfway connate. Disk annular, 1/2-1 mm high, fimbriate, in 3 flowers thick-fleshy with a narrow central canal, in 2 flowers thin, slightly 6-lobed. Pistil glabrous, in ¢ flowers absent. Infructescences 8-35 cm long, lax, with 1-4 long-stalked fruits; calyx nearly flat, faintly triangular to suborbicular, 8-10 mm @. Fruits narrowly ovoid, 34 by 11/4-2 cm, round to slightly triangular in cross-section; pericarp thin; pyrene smooth or with a faint median rib on the lids; lids c. 3 mm thick. Seeds 1 or 2; fertile cell(s) usually with a distinct adaxial rib, sterile cells moderately reduced.
Distr. SE. China (from about 27° N southwards), Hainan, and Indo-China; as the species is much cultivated, espe- cially in southern China and Tonkin, it is not well possible to draw the natural limits of its present area of distribution. In Malesia: Borneo (Sarawak, near Belaga, JAcoss 5239).
Ecol. Indense to open forests, usually at medium altitude; in Malesia in primary
928
FLORA MALESIANA
[ser. I, vol. 68
358296
6:
6:
121b
122
hill-forest below 500 m. Fi. Aug.
Uses. Planted as an ornamental and especially as a fruit tree. The fruits are highly esteemed among the Chinese; they are comfited or pickled. Only the pulp is eaten. The wood and resin are some- times used, but are of no great value.
Notes. C. pimela belongs to sect. Pime- la, to the relationship of C. pilosum, with C. album and C. parvum LEENH. (N. Viet- nam) as its possibly nearest allies.
For the complicated nomenclature of the present species see LEENH. /.c.
Add to Excluded genera: Nothoprotium Mia. Sum. (1861) 527 = Pentaspadon (Anacardiaceae); reduction already made by MARCHAND, Reéy. Anacard. (1869) 90, 183.
Campanulaceae
Sphenoclea zeylanica GAERTN.
Add. to Distr.: Northern Territory of Australia. Cf. BLAKE, Austr. J. Bot. 2 (1954) 137.
Codonopsis lancifolia (ROXB.) MOELIONO ssp. lancifolia.
Add to synonymy: Canarina moluccana Roxs. [Hort. Beng. 87] Fl. Ind. ed. Carey 2 (1832) 173; ed. Clarke (1872) 298. Cf. STEEN. Nova Guinea, Bot. 12 (1963) 191.
No specimens or drawing could be traced, but the brief description is clear. ROXBURGH had this species twice in his Flora, but the Moluccan one had 6-mer- ous flowers and was therefore arranged in another Linnean class.
In his monograph of Canarina, HED- BERG erroneously dismissed the Rox- BURGH name as a nomen nudum (Svensk Bot. Tidskr. 55, 1961, 19).
Add to the species of Lobelia:
Lobelia donanensis VAN ROYEN, Kew Bull. 20 (1966) 305, f. 1.
Small herb, witha glabrous, up to 12mm long stem. Leaves alternate, 11/2-21/2 cm long; limb ovate or ovate-elliptic, 7-12 by 5-7 mm, rounded or broadly cuneate at base, wavy along margins and with distinct, pear-shaped glands, obtuse or rounded at apex, coriaceous when alive, glabrous on either side, pilose along margin; petiole up to 11 mm, pilose in apical part only. Flowers choco- late-coloured, axillary and _ terminal; pedicels up to 6 mm, glabrous. Calyx 21/23 mm long; lobes lanceolate-linear, 1—11/2 mm long, usually with 3 glands on the teeth along the margin, pilose on outside. Corolla 5-6 mm long; tube 2-21/2 mm, pilose on outside, dorsally split to 1 mm from the base; free part of all lobes c. 3 by 3 mm, acutely acuminate;
Gemlssy Geas7 6: 141
ventral lobes on inside with dark purple papillae and near the throat also with 3 or 4 longitudinal crests. Stamens c. 3 mm, entirely connate; filaments glab- rous; anterior anthers 11/2 mm long, the posterior ones c. 1 mm; all connectives papillate, not barbate at tips. Ovary glob- ose, c. | mm @, glabrous; style glab- rous; stigmas dark purple, papillate along the margins. Capsule globose, 2!/2- 3 mm @, glabrous, thin-walled. Seeds obovoid, c. 0.7 mm long, subtriangular in cross-section, smooth.
Distr. Malesia: East New Guinea (Milne Bay Distr., Maneau Range, Mt Donana), one collection.
Ecol. Between dead moss in open grassland on limestone, 2250 m. Fi. fr. Aug.
Notes. According to the author allied to L. archboldiana, but differing in the gland-bearing leaf margin, the corolla lobes which are papillose inside and with longitudinal crests near the tube, the papillose connective, and glabrous ovary.
At Leiden we have no material of either species and these data are not checked. Lobelia borneensis (HEMSL.) MOELIONO. Add to Distr.: Also in Flores (Lesser Sunda Is.). Cf. STEEN. Blumea 15 (1967) 153.
Replace Phyllocharis DieELs, 1917, non FéE, 1824, by: Ruthiella STEEN. Blumea 13 (1965) 127, and the species names of the four species of this Papuan genus on p. 137-139 by:
1. Ruthiella oblongifolia (DrieELS) STEEN. Ee
2. Ruthiella schlechteri (DIELS) STEEN. /.c. 3. Ruthiella subcordata (MERR. & PERRY) STEEN. /.c.
4. Ruthiella saxicola (VAN ROYEN) STEEN. hes
Bottom line: Replace Legousia speculum- veneris (L.) Fiscu. by: Specularia specu- lum-veneris (L.) CARUEL (1888).
The generic name Specularia HEIST. ex Fase. Enum. Pl. Hort. Helmst. (1759) 151, nom. valid.; ed. 2 (1763) 225, descr.,
has. distinct priority over Legousia DuRAND, Fl. Bourg. 1 (1782) 37; 2 (1782) 26.
Caprifoliaceae (VAN STEENIS & KERN)
4: 175
4: 194
In this family we have omitted to mention any cultivated species, as none were known outside scientific botanic gardens, as far as collections were made. It was overlooked that in the former century KorTHALs had collected one which was described by MIQUEL as a new species from Java. Add to:
Formerly cultivated
September 1972]
4: 178a
4: 181
4: 182
Weigelia coraeensis THUNB. Trans. Linn.
Soc. 2 (1794) 331; Hara, En. Sperm.
Jap. 2 (1952) 63; STEEN. Blumea 13
(1965) 167. — Weigelia fallax Mia.
Fl. Ind. Bat. 2 (1856) 128. — Diervilla
fallax (Miq.) BorrL. Handl. FI. Ned.
Ind. 2 (1891) 6.
Note. Native in Kyushyu I., Japan, but rare in cultivation. This may well have been an original import by the East India Company, via the Dutch Settle- ment in Deshima, in VON SIEBOLD’s time. In the Catalogue of TEYSMANN & BINNEN- DIJK (1866) two Diervillas are mention- ed to be cultivated, but this was appar- ently unsuccessful and the genus is ab- sent in the Catalogue of the Bogor Bota- nic Gardens of 1930.
Lonicera malayana HENDERSON.
Of this extremely rare species two new
collections have come in, both collected
on the Selangor/Pahang Gentinh new
access road, on steep hillside, 1000 m
(FRI 3882 & 4519 WuitmMore, ff. and fr.
respectively).
Add to description: Big woody climber,
stem 2!/2 cm @, reaching to 20 m up a
tree; leafy branches scattered all the way
up. Fruit globular, c. S-8 mm @, black, juicy. Seeds 1-4, c.4 mm long, + convex, surface wrinkled.
Note. In both specimens the condensed inflorescence is terminal, without axillary stalks.
Line 35 from bottom. Delete from the
species of which the flowers are unknown
V. amplificatum and V. clemensae.
Replace lines 5-26 and complete the key
for flowering specimens as follows:
11. Corolla not distinctly tubular, tube less than 3 times as long as lobes, the latter often more than 1 mm long.
14. Inflorescence shortly pyramidal, paniculate. Filaments 1-3 mm long. Tube of corolla 1/4-1 mm long.
15. Leaves entire, glandular-pitted beneath in the axils of the primary and secondary side-nerves.
14. V. clemensae
15. Leaves crenate-dentate.
16. Leaves thinly coriaceous. Corolla 2-2!/2 mm long. Filaments in- serted at the base of the corolla. (Between 500 and 1500 m altitude, sometimes up to 2300 m.)
10. V. lutescens
16. Leaves manifestly coriaceous. Corolla 3 mm long. Filaments adnate to the corolla !/2-1 mm above the base. Without fruits hardly distinguishable from V. lutescens. (Altitude at least 2300 m.) . . . 21. V. junghuhnii
14. Inflorescence corymbiform. Fila- ments at least 6 mm long. Tube of
Addenda, corrigenda et emendanda 929
the corolla usually exceeding 1 mm. 17. Filaments in bud with inflexed top, 6 (-7) mm long.
18. Corolla broadly tubular, obovoid in bud, tube about 21/2 mm, lobes about 1!/2 mm. Leaves obtuse or shortly and bluntly acuminate.
3. V. glaberrimum
18. Corolla shortly tubular-turbinate, globular in bud, tube about 2 mm, lobes 1!/2-2 mm. Leaves gradu- ally long-acuminate.
4. V. platyphyllum 17. Filaments serpentine in bud, (7-) 8-10 mm long.
19. Underside of leaves with distinct glandular pits at the base on both sides of the midrib and often smaller ones in higher nerve-axils. Corolla turbinate, tube 21/2-3 mm, lobes 11/2-2 mm.
9. V. vernicosum
19. Leaves without glandular pits. Corolla rotate-cupular, tube 1 (-11/2) mm.
20. Leaves up to 26 by 14 cm, glabrous to softly villous be- neath. Corolla lobes 11/2 mm long . . 12. V. amplificatum
20. Leaves 12-17 by 8-9 mm, hispidulous beneath. Corolla lobes 2—2!/2 mm long.
8. V. hispidulum
4: 1885 12. Viburnum amplificatum Kern, Reinw.
1951) 1508. 8 BIS Mailed: 4: (1951) 188; Sarawak Mus. J. 9 (1960) 679, f. 1. — Descr. emend. — Fig. 3.
Small tree up to 15m, sometimes shrub- like. Leaves more or less coriaceous, dull, dark olivaceous above, brownish green or brown beneath, glabrous above, glabrous to softly villous with simple, forked, and stellate hairs beneath, neither glandular-pitted at the base nor bearded in the nerve-axils, elliptic-oblong, ovate or obovate, up to 26 by 14 cm; apex abruptly short-acuminate (acumen rather blunt, 1/2-1 cm); base cuneate to some- what rounded, slightly decurrent on the petiole; margin entire, sometimes remote- ly and obscurely undulate; nervation indistinct above, prominent beneath; primary nerves 4~7 on either side of the midrib, anastomosing; petioles 2-4 cm. Inflorescence umbellate, corymbiform, up to 20 cm across; axes stellately pubes- cent to subglabrous; peduncle stout, up to 10 cm; primary rays up to 8, up to 10 cm. Flowers small, c. 4 mm wide. Calyx-limb distinctly 5-lobed; lobes triangular, glandular-ciliolate, c. 1 mm. Corolla globular in bud, rotate-cupulate, glabrous, white; tube 1 mm, lobes ovate, c. 11/2 mm. Stamens inserted at the base
FLORA MALESIANA
[ser. I, vol. 68
Fig. 3. Viburnum amplificatum KERN. a. Flower bud, x5, b. stamen in bud, 13, c. expanding
flower,
x 61/2, d. calyx, e. glands on margin of
calyx lobe, f. part of corolla with stamen, x5, g. anther, x 13, A. fruit, 2, i. ditto, in cross sec-
6: 230
tion (a—i KOSTERMANS 13864).
of the corolla, much exserted; filaments serpentine in bud, 7-8 mm; anthers oblong, 1 mm. Ovary cylindrical, glabrous, 11/2 mm. Drupe oblong, sometimes very slightly dilated upwards, much flattened, with a distinct groove on both sides, at first green, twining black, 15-16 by 6-7 mm. Endocarp with a broad longitudinal groove on the dorsal side, the incurved edges forming a deep, broad, in cross- section bilobed furrow on the ventral side.
Distr. Malesia: N. and E. Borneo.
Ecol. Primary forests, up to 600 m.
Celastraceae (DING Hou)
Add before Taxonomy:
Palynology. An important study of the pollen of seven genera of Malesian Celastraceae was published by DING Hou, Blumea 17 (1969) 97-112, 1 fig., 8 pl., of which the main results with relation to affinity and generic distinc- tion are the following:
(i) The distinction of Kokoona and Lophopetalum is corroborated by the difference in pollen types.
(ii) In Lophopetalum at least four pollen subtypes can be distinguished.
(iii) Pollen of Sarawakodendron, de-
6:
25ie
232
: 2416
: 243a
: 256
260, 264
: 2665
: 2685
31391
: 392
39392
scribed below, shows a great resemblance to that of the related genera Kokoona and Xylonymus.
(iv) Pollen of Hedraianthera and Bras- siantha resemble that of the three latter genera but shows also resemblance to that of the African genus Salacighia. Unfortunately the numbering of the genera is erroneous; in the sequence of the first key it must be:
9. Bhesa, 1. Celastrus, 2. Maytenus, 3. Xylonymus, 12. Perrottetia, 8. Micro- tropis, 4. Euonymus, 5. Glyptopetalum, 6. Kokoona, 7. Lophopetalum, 10. Cas- sine, 11. Pleurostylia.
Maytenus emarginata (WILLD.) DING Hou.
Add to Distr.: Lesser Sunda Is. (Flores). Maytenus diversifolia (MaAxImM.) DING Hou.
Add to Distr.: Flores.
: 248ab Euonymus cochinchinensis PIERRE.
Add to Distr.: Burma, Formosa, and Flores. Glyptopetalum Taw. Add to Distr. (in China); Kweichow. Cf. Dinc Hou, Blumea 12 (1963) 59. BALAN MENON (Mal. For. 27, 1964, 18-21) confirmed the generic difference between Lophopetalum and Kokoona on their wood anatomy. Lophopetalum WIGHT ex ARN. Add to Distr.: A new species of the genus, the first from Australia, has been discovered by Mr. ByRNEs in the Kimber- ley District; it is related to the West Malesian species. Lophopetalum floribundum WIGHT. Add to Distr.: Two additional collections from Johore (FRI 8843) and Pahang (FRI 8147). Lophopetalum macranthum (LoEs.) DING Hou. Add to Distr.: A good flowering speci- men collected from East New Guinea (Morobe Distr.: NGF 37402). The new genus Sarawakodendron (re- cently found in Borneo) keys out for flowering material to Salacia. For fruiting material Sarawakodendron keys out at fork 8, which can be improved as follows: 8c. Fruits spindle-shaped, 3-angled, 3-cel- led, 3-valved. Seeds with a caruncle- like aril surrounded by many filamentous, fringed appendages. 3a. Sarawakodendron
3a. SARAWAKODENDRON
Dinc Hou, Blumea 15 (1967) 141. — Fig. 4.
Small tree, containing kautchuk, Leaves alternate. Stipules small, caducous. Jn- florescences solitary, axillary, simple, pe-
September 1972] Addenda, corrigenda et emendanda 931
Fig. 4. Sarawakodendron filamentosum DiNG Hou. a. Habit, x 2/3, b. inflorescence, <4, c. flower, <2, d. flower with petals removed, 3, e. stamen, x 8, f. longitudinal section of flower shown in d, x3, g. longitudinal section of ovary, <8, A. cross section of ovary with position of stamens indicated, x 8, i. fruit x 2/3, 7. dehiscent fruit with one valve showing scars of seed attachments, x 2/3, k. one fruit valve with attached descendent seeds, 2/s, /. seed, <1, m. basal part of seed seen from beneath, x 2, 7. co- tyledons, <2 (a-h DinG Hou 333, i JUGAH ANAK Kupt S 24897, j-n JUGAH ANAK Kupi S 24898).
932
6: 397b
6: 404
6: 410a
6: 4105
6: 4135
FLORA MALESIANA
[ser. I, vol. 68
duncled, few-flowered, the axis of the racemes densely set with imbricate, decus- sate, persistent bracts. Flowers bisexual, solitary, pedicelled. Sepals 5, imbricate. Petals ditto. Disk extra-staminal, fleshy, flattened. Stamens 3, extrorse, transver- sely dehiscing. Ovary partly immersed in the disk, 3-celled, with 3 stigmatic lobes; no style; ovules c. 8 in each cell. Capsule ellipsoid, pointed to both ends, hard, 3-gonous, with 3 thick valves, dehiscing to the base. Seeds descending, narrowly elliptic, with a fleshy orbicular aril provided with numerous filiform, fringy appendages.
Distr. Malesia: Monotypic.
Note. Allied to Salacia, Lophopetalum, Kokoona, and Polycardia of Madagascar.
Borneo: Sarawak.
1. Sarawakodendron filamentosa DING Hou, Blumea 15 (1967) 141, f. 1. — Fig. 4.
Tree, 7-12 m, 10-15cm @, the vege- tative parts containing yellow kautchuk particles and resinous threads. Stipules 1/> mm, -+ erose. Leaves oblong to lanceolate, acuminate, 10-25 by 4-10 cm, subentire; nerves 6—7 pairs; petiole 2/3-1 cm. Inflorescences 1-2/2 cm; pedicels articulated near base, 11/2—-2 cm. Sepals 11/2-2 by 1 mm, semi-orbicular. Petals c. 5 mm @, pale-orange. Disk c. 4 mm @&. Stamens reflexed at anthesis. Capsule 6-81/2 by 2-3 cm. Seeds 2-21/ by 1/2 cm, albuminous; aril 5-7 mm @, the chalazal filaments 11/2-2 cm long; embryo narrow-lanceolate, 18 by 4 mm; cotyledons foliaceous.
Distr. Malesia: Borneo (Sarawak).
Ecol. Understorey tree in lowland kérangas forest.
Note. Its position exactly links the families Celastraceae and Hippocrateaceae in their former circumscription. Siphonodon peltatus DING Hou.
Add to Distr.: A second collection is HARTLEY 13179 with flowers in anthesis. Salacia L.
Line 1 literature: replace (1767) by (1771). Add to synonymy: Annulodiscus TARDIEU Biot, Bull. Soc. Bot. Fr. 95 (1948) 264; FI. Gén. L.-C. Suppl. 1 (1948) 812. Salacia sororia Mia.
Add to Distr.: Now also found in the Solomons (Guadalcanal, BSIP 9152). Salacia forsteniana Mia.
Add to Distr.: Now also found in the Solomons (Wagnia & SE. New Georgia Is.: BSIP 5436, 5985).
Salacia grandiflora Kurz.
Add to synonymy following NG, Blumea 18 (1970) 412: Maba hierniana K. & G. J. As. Soc. Beng. 74, ii (1905) 203. — Diospyros hierniana (K. & G.)
6: 415a
6: 419a
4: 595a
4: 533
4: 569a
4: 584b
BAKH. Gard. Bull. S. S. 7 (1933) 173. Salacia verrucosa WIGHT.
Add to synonymy: Annulodiscus nigri- cans TARDIEU BLOT, Bull. Soc. Bot. Fr. 95 (1948) 264; Fl. Gén. I.-C. Suppl. 1 (1948) 812.
Salacia chinensis L.
Line 1 literature: replace (1767) by (1771).
Chenopodiaceae
Chenopodium pumilio R. Br. Prod. (1810) 407; BLAck, FI. S. Austr. 2 (1948) 289; AELLEN in Hegi, Ill. Fl. Mitt.-Eur. 3, 2 (pt 2) (1960) 578, 597, f. 255 D-E; STEEN. Blumea 15 (1967) 154.
Small, soft, prostrate aromatic herb, with green flowers. Easily distinct from C. carinatum R. Br. by narrow, not crested perianth segments, the nut being discernible between them.
Distr. New Zealand and Australia, apparently recently introduced in Male- sia: New Guinea: Morobe Distr. (B.G.D. Leron Cattle Station: E. E. HENtry NGF 16668, Aug. 1, 1963).
Ecol. Bare patch in browsed grassland, at 150 m.
Combretaceae
Due to the interest and activity of the Division of Botany, Lae, many additions must be made to the Combretaceae of New Guinea, two papers on which ap- peared by M. J. E. Coope, in the ‘Manual of the Forest Trees of Papua and New Guinea’, Port Moresby 1964, n. 1, 45 pp., 25 pl., and in a much revised edition of this, 1969, 86 pp., 32 pl., 8 maps. Herein one named and three unnamed new New Guinean species are distinguished, besides some of the Solomons and New Ireland and full keys are given. A formal treatment will soon appear. : It has also been found that within Terminalia occur at least four distinct types of germination, a character which may add to the systematy of the genus. Coope’s paper also adds considerably to our understanding of the ecology of the species and provided numerous field notes. The reader is referred to COODE’s paper for details. Terminalia crassifolia EXELL. Add to Distr.: This Papuan endemic now also recorded from Australia: Northern Territory, Fletcher Creek, Wearyan R.; cf. Muelleria 2 (1971) 135. Add to Insufficiently known species:
Terminalia macrantha Rojo, Blumea 17 (1969) 93, f. 1. Tree, c. 18-20 m, 40cm @. Indumentum
September 1972]
5: 495
5: 496
of simple, brown, sericeous hairs. Young branchlets c. 1-2!/2 cm @, glabrescent. Leaves densely crowded at the very ends of branchlets, chartaceous, rather laxly hairy, above shiny, verruculose, glabres- cent except on the main nerves, beneath persistently hairy, obovate-oblong, 12-31 by 5-12 cm, generally widest at about 3/4 of the length, top rounded or sometimes emarginate, base narrowly cuneate or sometimes decurrent, glandless; midrib prominent on both sides, densely hairy; nerves on both sides rather widely spaced, hairy, the upper 3 or 4 pairs arcuating towards the top but not anastomosing at the margin, connected by thin veins in a more or less scalariform pattern with some interconnections between them, domatia none; petiole 8-24 mm, densely to sparsely hairy, without glands. Spike axillary, c. 13-25 cm, lower c. 7-8 cm flowerless; densely hairy. Bracts + obovate-oblong to + linear, c. 3 mm. Flowers sessile, densely hairy, greenish yellow, fragrant. Part of the flowers, scattered in the inflorescence, with rudi- mentary style and somewhat smaller. In the bisexual ones, the lower receptacle (ovary) 3-6 mm; upper receptacle funnel- shaped, 2-3 by 31/2 mm. Calyx lobes deltoid, 2 mm, sparsely hairy inside. Filaments 9-11 mm, glabrous; anthers 2/3 mm. Disk sparsely set with rather lax hairs. Style simple, terete, 9-10 mm, glabrous. Ovules 2, pendulous.
Distr. Malesia: Philippines (Samar: Mt Calbiga, Wright), once collected (PNH 6409).
Ecol. On top of flat ridge, 300 m, lo- cally common.
Vern. Bongoran, S. L. Bis.
Note. For lack of fruit its affinity can- not well be established. There is some re- semblance to 7. zollingeri, which has coriaceous leaves, more nerves, a smaller upper receptacle, and shorter filaments, and also with T. darlingii, which has also more nerves, subopposite glands at the base or top of the lamina, larger bracts, a shorter lower receptacle, and longer filaments.
Connaraceae (LEENHOUTS)
Add to Ecology: Cf. H. G. BAKER, Bot. Gaz. 123 (1962) 206-211, on heterostyly and pollination; he also confirms the seed dispersal by birds.
Add to Morphology, paragraph on the arilloid: Cf. CORNER, Phytomorphology 3 (1953) 471, for a different opinion as to the morphological nature of the aril-like structures.
After Morphology add: Phytochemistry. Cf. HEGNAUER, Che-
Addenda, corrigenda et emendanda
: 499a
= 505
: 506a
: 507b
933
motaxonomie 3 (1964) 545-546, 673. Add to Taxonomy, paragraph on affinity with other families: On embryological grounds, MAURITZON, Act. Un. Lund N.S. 35, n. 2 (1939) 13 & 39, points to a possible relationship with the Cunonia- ceae; GUTZ-WILLER, Bot. Jahrb. 81 (1961) 38-39, gives a not convincing argumentation for the inclusion of the family in the Sapindales; HUTCHINSON, Gen. Pl. 1 (1964) 162, derives the family from the Dilleniaceae; HEGNAUER, Che- motaxonomie 3 (1964) 546, finds some phytochemical support for a position near the Leguminosae; LEINFELLNER, Oest. Bot. Z. 118 (1970) 542-559, from a study of the gynoecium, also con- cluded to a closer relationship with the Leguminosae.
Cnestis palala (LoUR.) MERR.
Add to literature: VipAL, Fl. Camb. Laos & Vietn. 2 (1962) 13, t. 1 f. 1-9; Corner, Life of Plants (1964) t. 23. Add to Distr. of ssp. diffusa: Erroneously cited by VIDAL, /. c., from Sumatra, the Malay Peninsula, and Borneo.
Agelaea macrophylla (ZOLL.) LEENH. Add to synonymy: Myristica laurina (non BL.) Hocure. Candollea 6 (1936) 459. Agelaea trinervis (LLANOS) MERR.
Add to literature: VIDAL, Fl. Camb. Laos & Vietn. 2 (1962) 18, t. 1 f. 10-20. Add to Uses: In Vietnam, an oil from the seeds is used for lighting (VIDAL, /.c.). Agelaea borneensis (HooK. f.) MERR. Add to Ecol. as exceptional highest altitude: (-1300) m.
Agelaea insignis (SCHELLENB.) LEENH. Change description as follows: after Branchlets, ‘probably’ should be omitted. Leaflets, Sth line, add after tomentose: above glabrescent. Include before Fruits: Infructescences solitary or fascicled, cymose, up to 5 cm long, fulvous- velutinous. Add at the end of the de- scription of the fruits: via greenish-choco- late to red.
Roureopsis PLANCH.
Change the date of publication of B. & H. Gen. Pl. 1 (sub Taeniochlaena) into: 1862.
Roureopsis asplenifolia SCHELLENB.
Add to literature: CorNER, Life of Plants
(1964) t. 23. Insert after 1. Roureopsis asplenifolia SCHELLENB.: la. Roureopsis stenopetala (GRIFF.) SCHELLENB. Kew Bull. (1927) 375; Cras, Fl. Siam. En. 1 (1928) 362;
SCHELLENB. Pfl. R. Heft 103 (1938) 110; VIDAL, Fl. Camb. Laos & Vietn. 2 (1962)
23, t. 2 f. 1-6. — Cnestis stenopetala GriFF. Notul. 4 (1854) 433, t. 611 f. 2 (‘steriopetala’). — R. incurva PIERRE,
Fl. Coch. 5 (1898) t. 379 A.
934
FLORA MALESIANA
[ser. I, vol. 6%
5: 508a
S13 S10)
5: 509a
5: 5095
De a10
Liana. Branchlets fulyvous-tomentose when young, glabrescent. Leaves 5—10-ju- gate, leaflets mostly opposite or nearly so, the petiole and the rachis tomentose. Leaflets subsessile, the lateral ones (lower- and uppermost excepted) distinct- ly asymetric, 11/2-3 by 1/4—-1!/2 cm, basal ones broad-ovate, up to 13/4 by 11/4 cm, terminal leaflet elliptic or oblong, 2—31/a by 1!/4-12/4 cm, all stiff-chartaceous to subcoriaceous, midrib puberulous at both sides, furthermore glabrous; base rounded to (terminal) cuneate; apex blunt to rounded, emarginate; nerves 4-5 pairs, the lowermost at the broader acroscopic side ascending, all looped and joined, veins about as strongly developed as the nerves. IJnflorescences racemose, umbelliform, with up to c. 6 flowers, c. 11/2-2 cm long, bracts tomentellous, furthermore glabrous; rachis c. 4 mm, pedicels slender, c. | cm long or more. Flowers 5-merous. Sepals linear-lanceo- late, 3-5 by 1 mm, blunt, glabrous but for the tomentose outer side of the tip. Petals linear, c. 8 mm long, puberulous at the tip. Stamens confluent at base. Fruits 1 (?) per flower, c. 11/2 cm long, glabrous.
Distr. Burma, Thailand, Cambodia, Laos, and Malesia: Malay Peninsula (Ranong, Phangnga).
Ecol. Primary and secondary forests at low altitude. F/. Febr.
Note. Nearest to R. asplenifolia SCHELLENB. as already mentioned under that species (5: 5075). The main differ- ences are the usually smaller number of slightly bigger and stiffer leaflets which are not hairy along the margin and have the base mostly rounded, the much longer pedicels, and especially the long and narrow sepals in the present species. Roureopsis emarginata (JACK) MERR. Add to literature: VipAL, Fl. Camb. Laos & Vietn. 2 (1962) 24, f.2 f. 7 & 8. Add to Distr.: Laos.
Sect. Taeniochlaena LEENH.
Change the year of publication of B. & H. Gen. Pl. 1 into: 1862.
Roureopsis acutipetala (M1Q.) LEENH. Add to literature: VipAL, Fl. Camb. Laos & Vietn. 2 (1962) 26, t. 2 f. 9-11. Line 4 literature: change 1865 into: 1862. ssp. borneensis (SCHELLENB.) LEENH. Add to Distr.: According to VIDAL, /.c., also in S. Vietnam.
Rourea AUBL.
The first citation for Jaundea should be changed as follows: Jaundea GILG in E. & P. Nat. Pfl. Fam. 3, 3 (1894) 388. Add to Sect. Palliatus: Cf. LEENH. in Steen. Pac. Pl. Areas 1 (1963) 278, map 7h
Rourea oligophlebia MERR.
5: 5l4a
5 ibd
5: 5155
Silda
SHAS TB
525192
3115204
Add to literature: VmDAL, FI. Camb. Laos & Vietn. 2 (1962) 31, t. 3 f. 1-4. Rourea minor (GAERTN.) LEENH.
Add to literature: VipAL, Fl. Camb. Laos & Vietn. 2 (1962) 34, t. 4 f. 1-10; W. R. Sykes, Fl. Niue (1970) 70, f. 4. Add to literature sub Rourea acropetala PIERRE: VIDAL, Fl. Camb. Laos & Vietn. 2 (1962) 32, t. 3 f. 5-7.
Add to synonymy, after Santaloides cordatum: Santaloides ovale SCHELLENB. Bot. Jahrb. 59 (1924) Beibl. . 131, p. 29. Add to synonymy, after Santaloides luzoniensis SCHELLENB.: Connaropsis ru- bescens RIDL. J. Bot. 62 (1924) 295; cf. LEENH. Blumea 12 (1963) 20.
Add to synonymy, after Santaloides elmeri SCHELLENB.: Rourea ovale LEENH. Fl. Mal. I, 5 (1958) 520; cf. LEENH. Blumea 12 (1963) 21.
Add to Distr.: Solomon Is., Tonga, Niue I.
VIDAL, /.c., distinguished the three groups cited as subspecies as follows: ssp. microphylla (Hook. & ARN.) VIDAL for the small-leaved category, ssp. minor for the intermediates, and ssp. monadelpha (RoxsB.) VIDAL for the group with few, relatively large leaflets. I had refrained from giving such a subdivision as these groups are neither morphologically, nor geographically sharply delimited. Fur- thermore, ViDAL excluded R. acropetala PIERRE from the synonymy. That species should mainly differ from R. minor by the reduction of the terminal leaflet and by the distinctly mucronate apex of the leaflets.
Add to the 4th paragraph, on more or less intermediate forms: R. ovale (Bor- neo).
Rourea mimosoides (VAHL) PLANCH. Add to literature: VIDAL, Fl. Camb. Laos & Vietn. 2 (1962) 41, t. 3 f. 8-12. Add to Distr.: Vietnam.
Add to Ecol.: peat-swamp.
f. mimosoides. Add to literature: ANDER-
son, Gard. Bull. Sing. 20 (1963) 172. Omit under Distr.: Sumatra excepted. Rourea prainiana TALBOT.
Sometimes a big climber.
Add to Distr.: Ceylon (acc. to TALBOT). Add after Malay Peninsula: also Pahang (G. Benom).
Under Incertae sedis: Drop R. ovale (SCHELLENB.) LEENH. as ripe fruits showed that this is identical with R. minor. Add under Incertae sedis:
Rourea pinnata (MERR.) VELDKAMP, Blumea 15 (1967) 543. — Sarcotheca pinnata Mere. J. Str. Br. R. As. Soc. 86 (1922) 314; KNutnH, Pfl. R. IV, 130 (1930) 426.
Liana. Leaves
Twigs subglabrous.
September 1972]
Addenda, corrigenda et emendanda
o55
5: 521a
52 5216 23a
5: 523b
5: 5246
39/526
3-4-jugate; lateral petiolules 4-5 (-7) mm long. Leaflets elliptic to oblong, 12-27 by 4!/2-11 cm, thin chartaceous, shining above, rather dull beneath, minutely hairy on midrib and nerves beneath; base equilateral, in lateral leaflets round- ed (to subcordate), in the terminal leaf- let broadly cuneate; apex tapering acu- minate, acumen short, broad, and blunt; nerves 10-12(-15) per side, pinnate, ascending, slightly curved, only the uppermost ones joined, intermediate veins often nearly as strongly developed as the nerves, veins and veinlets scalari- form, beneath much better visible than above. /nflorescences axillary, soli- tary or 2 collateral ones together, up to 7 cm long, sparsely branched, rather densely minutely hairy. Calyx 21/2-23/4 mm high, outside densely appressed hairy, margin densely ciliolate, inside glabrous. Coro/la (not yet fully developed) 33/4 mm long. Ovary and lower half of the style sparsely woolly. Fruit unknown.
Distr. Malesia: North Borneo (once collected at Batu Lima near Sandakan).
Ecol. On dry forested slopes at low altitudes. Fl. Oct.
Note. As the fruit is still unknown it is not well possible to include this species into one of the subgenera or sections. The key in Fl. Mal. I, 5 (1958) 513, leads to R. minor. It is distinctly different from that species, however, in the following characters: greater number of nerves, nervation open and — by the intermediate veins — seemingly very dense, venation distinctly scalariform (in R. minor mostly laxly reticulate), inflorescences puberulous, calyx outside densely hairy.
Ellipanthus tomentosus KURZ.
Add to literature: VmpAL, FI. Camb. Laos & Vietn. 2 (1962) 46, t. 4 f. 11-14. Add to first sentence of description: up to 40 m high, sometimes buttressed.
ssp. tomentosus. Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 172.
var. tomentosus.
Add to Uses: According to VIDAL, /.c., the wood is soft and not durable. Ellipanthus beccarii PYeRRE.
Add to literature: Meter, Bot. Bull. Herb. Sandakan 10 (1968) pl. between p. 104 and 105.
Add to Ecol.: peat swamp forests.
Key to the species.
Lead 10(b), add to length of petals: (—5) mm.
Lead 16(a), insert between ‘dry’ and ‘blackish-verrucose’: mostly.
Connarus euphlebius MERR.
var. bullatus LEENH. Replace ‘in the press” by: 106.
Sisto,
SIC Sys ys 17)
5: 533b
5: 534a
5: 535a
329'535D
5: 536a
5: 538a
5: 5386
5: 539a 5: 5395
ssp. moluccanus LEENH. Replace ‘in the press’ by: 106.
Connarus villosus JACK.
Add to Distr.: N. Borneo (P. Gaya). Add to Ecol.: Altitude up to 500 m. Connarus culionensis MERR.
Add to literature: VIpAL, Fl. Camb. Laos & Vietn. 2 (1962) 54, t. 5 f. 13 & 14. Add to Distr.: Sarawak, Sabah, S. Viet- nam (VIDAL).
var. culionensis.
Add to Distr.: According to VIDAL, /.c., also in S. Vietnam.
var. stellatus (MERR.) LEENH.
Delete in description the sentence ‘Fruit unknown.’ (The fruit is not differ- ent from that of the type variety.) Connarus odoratus Hook. f.
Add to Note: The delimitation of C. villosus, culionensis, and odoratus has to be studied anew when more material will be available. Especially in northern Borneo, where the three species overlap, the identification is often uncertain. Connarus paniculatus RoxB.
Add to literature: VmaAL, FI. Camb. Laos & Vietn. 2 (1962) 55, t. 6 f. 1-10; ANON. Ic. Roxb. (1970) t. 28.
Add to Distr.: Burma.
Add to Notes: VIDAL, /.c., separates the species into two varieties: var. panicu- latus and var. hainanensis (MERR.) VipaL. Among the Indo-Chinese mate- rial of the former of these he distinguishes 3 formae. According to him, the material of the Malay Peninsula belongs to var. paniculatus.
Connarus semidecandrus JACK.
Add to literature: Vmar, FI. Camb. Laos & Vietn. 2 (1962) 51, t. 5 f. 1-12; ANDERSON, Gard. Bull. Sing. 20 (1963) 172.
Add to Distr.: Sumbawa, Flores; according to Hosokawa, J. Jap. Bot. 13 (1937) 275, also Truk I.
VIDAL, /.c., distinguishes among the Indo-Chinese material, here all included in group 8, 3 forms, exclusively on leaf characters.
Connarus cochinchinensis (BAILL.) PIERRE. Add to literature: VmaL, FI. Camb. Laos & Vietn. 2 (1962) 58, t. 6 f. 11-18. Connarus lamii LEENH.
Replace ‘in the press’ by: 106.
Connarus monocarpus L.
Add to literature: CORNER, Life of Plants (1964) t. 23.
Description of the fruit, 3rd line, read: in the latter case usually distinctly shortly stipitate.
Connarus lucens SCHELLENB.
Start description with: Liana, up to 10 m. Add to Distr.: Now collected in some parts of Sarawak.
Add to Ecol.: Clayey river bank, peri-
936
FLORA MALESIANA
[ser. I, vol. 68
5: 540a
5: 5406
5: 5406
5: 54la
4: 390
odically flooded, up to 250 m.
Connarus winkleri SCHELLENB. ssp. philip- pinensis LEENH.
Replace ‘in the press’ by: 106.
Connarus schumannianus GILG.
The head should be changed as follows:
18. Connarus conchocarpus F. v. M. Fragm. 5 (1866) 105; SCHELLENB. Pfl. R. Heft 103 (1938) 228.
ssp. schumannianus (GILG) LEENH., nov. stat. — C. schumannianus GiLG in K. Sch. & Laut. Fl. Schutzgeb. (1900) 341; SCHELLENB. Pfl. R. Heft 103 (1938) 253; LEENH. FI. Mal. I, 5 (1958) 540, f. 11 j. Add to Notes: Ssp. conchocarpus from NE. Queensland differs from ssp. schu- mannianus in the following points: branches more densely lenticellate; leaves more often 2-jugate; leaflets thicker, mostly at base more acute and _ less peltate, midrib often above flat rather than sunken, nervation and venation more coarse and more prominent be- neath; inflorescences apparently usually axillary; fruits hardly curved, smaller (21/4 by 13/4 cm).
The relationship is doubtless with C. pickeringii A. GRAY (Fiji, Solomon Is.) and with C. salomoniensis.
Connarus salomoniensis SCHELLENB.
Add to description of Petals, after ‘21/2 by 11/2 mm’: or linear-lanceolate, 5 by 1 mm.
Replace the Ist paragraph of the Notes by:
The present species is doubtless related to C. conchocarpus and to C. pickeringii A. GRaAy from Fiji and the eastern Solomon Is. (San Christobal, Vanikoro I.). From the former it is distinctly different in its flowers and fruits, but hardly so in the leaves. C. pickeringii differs constantly by the on both surfaces hairy petals and the smaller fruits (up to c. 3 by 21/2 cm); moreover, the leaflets of the latter species are as a whole narrower (up to 71/2 cm) and hardly, if all, ovate. I should not be surprised, however, if these differences should fade away when more material from the Solomon Is. comes available. The only reason that I not yet reduce C. salomon- iensis to C. pickeringii is that the charac- ter ‘glabrous petals’ respectively ‘hairy petals’ seems as a whole to be important in this genus.
If C. salomoniensis and C. pickeringii should be combined, no reason would be left to keep C. peekelii upright as a species.
Convolvulaceae (VAN OosTSTROOM)
In Key line 18 from top read instead of “6. Styles 2, free or united near the base’:
4: 392
4: 398
6. Styles 2 or 1, in the latter case 2-bran- ched with the branches distinctly visible.
Replace forks 7 and 8 bij the following:
7. Style 1, with 2 branches.
7a. Flowers in a long racemose inflores- cence, each flower inserted on a large, elliptic, white bract. Corolla to 11/2 cm long. Ovary glabrous; stigmas horse-shoe-shaped. 5a. Neuropeltopsis Flowers in an umbelliform cyme, not inserted on the bracts. Corolla 3-5 cm long. Ovary hairy; stigmas globose-peltate 4. Bonamia 7. Styles 2, free.
8. Each style forked and with 2 filiform or slightly clavate stigmas. Corolla- limb nearly entire. Small herbaceous PlantSiemery | eee 3. Evolvulus
8. Styles not forked; stigmas capitate or kidney-shaped. Corolla-limb dis- tinctly lobed.
8a. Large woody twiners. Bracts much enlarged in fruit, elliptic, scarious. Stigmas kidney-shaped. 5. Neuropeltis 8a. Low perennials. Bracts unaltered in fruit. Stigmas capitate. 3a. Cressa Line 1 from top: the correct name for 1. Subgenus Grammica is: 1. Subgenus Cuscuta. Insert above 4. Bonamia:
3a. CRESSA
LINNE, Sp. PI. ed. 1 (1753) 223; Gen. PI. ed. 5 (1754) 104; R. Br. Prod. 1 (1810) 489; Bru. Fl. Austr. 4 (1869) 437; F. M. BatLey, Queensl. FI. 4 (1901) 1074. — Fig. 5.
Low, much-branched, grey-pilose, pe- rennial plants, often suffrutescent at the base. Leaves small, sessile, entire. Flowers small, shortly pedicelled or subsessile in the upper leaf-axils, in terminal clusters or spikes; bracteoles 2, small, at the base of the calyx. Sepals 5, subequal, obovate, imbricate, about as long as the corolla- tube, not enlarged in fruit. Corolla regular, funnel-shaped, white or rose; lobes 5, spreading or reflexed, about as long as the tube, pilose outside. Stamens 5, exserted; filaments adnate to the corolla-tube, filiform, glabrous; pollen globular, smooth. Ovary hairy, 2-celled, 4-ovuled; styles 2, exserted, free, filiform, each with a capitate stigma. Capsule exceeding the calyx, opening by valves, usually 1-seeded; seed smooth, glabrous.
Distr. A genus with a small number of closely related species, considered by some authors as a single one (for example Cuotsy and BENTHAM), occurring in the warm temperate and tropical regions of both hemispheres in sandy, maritime
Ta.
September 1972]
Addenda, corrigenda et emendanda
ox"
Fig. 5. Cressa cretica L. a. Habit, x4, c. calyx, supported by 2 bracteoles,
or saline circumstances; in Malesia: Lesser Sunda Is. (Timor). Ecol. Generally in sandy, maritime
or saline terrain.
1. Cressa cretica LINNE, Sp. Pl. (1753) 223; Cuotsy in DC. Prod. 9 (1845) 440; Bru. Fl. Austr. 4 (1869) 437; F. M. BAILEY, Queensl. Fl. 4 (1901) 1074. — C. australis R. Br. Prod. (1810) 490. —
x4, x6 (a-d CINATTI 339, Port. Timor).
An erect or diffuse, much-branched perennial, sometimes almost woody at the base, c. 20 cm, strigose appressed- hairy all over. Leaves subsessile to shortly petioled, elliptic-oblong or slightly ob- long-obovate, acutish, 5-9 by 3-4 mm. Bracteoles ovate-oblong. Flowers sessile, in terminal heads. Sepals broadly obova- te, obtuse, ciliate, 3!/2 mm long. Corolla shortly exceeding the calyx, hairy out- side, 5!/2-6 mm long. Anthers oblong, c. 14/3-11/2 mm long.
Distr. As the genus; in Malesia: Lesser Sunda Is. (Portuguese Timor: Batugadé, lagoon of Bé-Malai, CrNnaAtTTI 339), once collected.
4: 402
Ecol. At 100 m from the sea in a pure stand in the shade of the mangrove tree Avicennia marina. Fl. Aug. 1962.
Note. The Timor material exactly matches specimens from tropical Austra- lia, which form BENTHAM referred to C. cretica sens. lat., observing that the flowers of tropical Australian specimens are larger than those in the European typical form. In the latter the bracteoles are mostly linear, the sepals c. 3 mm long, the corolla c. 4 mm, the anthers 1 mm. Insert above 6. Porana:
5a. NEUROPELTOPSIS
Ooststr. Blumea 12 (1964) 365, f. 1. — Fig. 6.
Differs from Neuropeltis by the longer and less dense racemose inflorescences, the already at flowering time large, white bracts, the broader glabrous wings at both sides of the hairy midpetaline areas of the corolla, the filaments almost wholly adnate to the corolla, and the style with 2 short branches, each bearing a horse-shoe-shaped stigma.
Distr. Monotypic, endemic in Malesia.
1. Neuropeltopsis alba Ooststr. Blumea 12 (1964) 365, f. 1. — Fig. 6.
A woody twiner. Young branches tomentellous, adult ones glabrescent or glabrous. Leaves elliptic, c. 8-12 by 4-7 cm, rounded at the base or very shortly attenuate into the petiole, abruptly acuminate and mucronulate at the apex, glabrous or nearly so above, rather densely pilose beneath; lateral nerves 5-6 on either side of the midrib; petiole c. 2-3 cm long. Jnflorescences from the leaf-axils, exceeding the leaves, c. 18-20 cm long; pedicels 2-8 mm long. Bracts white, elliptic to broadly elliptic, at the base very shortly attenuate into the petiole, abruptly acuminate and mucro- nulate at the apex, bearing the flower a _ little below their centre, c. 4 by 2-21/2 cm, shortly pilose, mainly on the midrib and at the margins. Sepals c. 3 mm long, two outer ones elliptic, obtuse, three inner ones broader than long, c. 4 mm broad. Corolla funnel- shaped, c. 12-13 mm long, 5-lobed, the lobes a little shorter than the glabrous tube, broadly ovate, the midpetaline areas pilose outside, the glabrous wings irregularly dentate. Filaments pilose at the base. Ovary glabrous; style 1, with 2 short branches; stigmas horse-shoe- shaped, papillose.
Distr. Malesia: E. Borneo (Sangkuli- rang Distr., Karangan R., NW. of Sangkulirang, KosTERMANS 13588).
Ecol. On sandstone, at c. 20 m alti- tude. F/. Aug.
FLORA MALESIANA [ser. I, vol. 6®
AK
Fig. 6. Neuropeltopsis alba OoststR. a. Flowering branch, 2/3, b1~®. sepals 1-5, x6, c. corolla with stamens, ™ 3, d. pistil, « 6 (a-d KOSTERMANS 13588).
September 1972]
4: 431
4: 435a
4: 440
4: 441
4: 444a
4: 4455
4: 446b
Addenda, corrigenda et emendanda
939
Jacquemontia CHOISY. Line 5 from bottom, after ‘lanceolate’ add: to linear. Jacquemontia browniana OOSTSTR. Line 2 from top, after ‘with’ add: greyish, pale brown or. Line 4 from top, after ‘lanceolate’ add: to linear. Line 4 & 5 from top, alter the sizes as follows: 21/2-7 cm by 2-10 mm. Merremia DENNSTEDT. Change in the Key forks 13 & 14 (lines 20-22 & 51 from top) as follows: 13. Sepals partly or all attenuate-acu- minate towards the acute apex. 13a. Leaves entire. Sepals 12-15 mm long, the outer ones verruculose and sparsely hairy. Ovary hairy. 6a. M. aniseiifolia 13a. Leaves generally dentate near the base. Longest sepals to 7 mm, rarely to 10 mm long, smooth and glabrous. Ovary glabrous. 6. M. tridentata 13. Sepals obtuse or rounded at apex. 14. Sepals 10 mm long or mostly shorter. 15. Twining or prostrate herbs .. . ec. 14. Sepals longer than 10 mm.
20. Sepals at most 12 mm long... efc. Line 1 from top: the correct name for 1. Section Eu-Merremia is: 1. Section Merremia.
Merremia emarginata (BuRM. f.) HALLIER
As already stated in my monograph of the genus Evolvulus (Thesis Utrecht, 1934, 245; Med. Bot. Mus. Herb. Utrecht 14, 1934, 245) Evolvulus gange- ticus)(.) LL: Sp! Pl. ed. 2 (1762) 391 (= Convolvulus gangeticus L. Amoen. Acad. 4, 1756, 306) belongs to the Menis- permaceous Cocculus hirsutus (L.) DIELS (= Cocculus villosus DC.), such in agreement with HALLIER f. (Bull. Herb. Boiss. 6, 1898, 720, 723). VERDCOURT (in Hubbard & Milne-Redhead, ed., FI. Trop. East Afr., Convolv., 1963, 55) who could study the type in the Linnean Herbarium, also arrived at this conclu- sion. The statement by CUFODONTIS (Bull. Jard. Bot. Brux. 31, Suppl., 1961, 743) that the correct name for M. emarginata should be M. gangetica (L.) CuF. (also based on Convolvulus gange- ticus L.) is therefore untenable.
Merremia tridentata (L.) HALLIER f. Line 4 from top: the correct name is: ssp. hastata Ooststr. Blumea 3 (1939) 317, f. 2-0, because the synonym Convolyulus hastatus Drsr. in LAMK, Enc. 3 (1789) 542, non Forsk. 1775, is nom. illeg. This is in accordance with Art. 72, note, of the International Code. Insert before 7. Merremia quinquefolia:
4: 447a
4: 451b
4: 452b
4: 457b
4: 458 4: 459
6a. Merremia aniseiifolia Ooststr. Blu- mea 12 (1964) 363, f. 1. — Fig. 7.
A herbaceous twiner (or prostrate 7). Stems slender, sparsely pilose, glabres- cent. Leaves sparsely pilose, lanceolate to linear-lanceolate or sometimes oblong, (21/2-) 5-7 cm by 6-10 mm, rounded at the base, acute and mucronulate at the apex, entire, adpressed-pilose at the margins, otherwise sparsely pilose to glabrous; lateral nerves 4-6 on either side of the midrib; petiole 3-5 mm long. Flowers in 1-flowered cymes. Peduncles axillary, 2-4 (-6) cm long, slender, sparsely pilose to glabrous; pedicels thickened towards their top, minutely warty, 6-10 mm long. Bracts minute, subulate. Flower-buds conical, acute. Sepals equal in length or the inner ones a little shorter, 12-15 mm long, the two outer ones rather thick, ovate-lanceolate to narrowly ovate, attenuate or acumi- nate towards the acute apex, minutely warty and sparsely short-pilose, the three inner ones membranaceous, oblong, cuspidate, smooth and glabrous or only the third sepal warty at the base. Corolla funnel-shaped, probably c. 2-21!/2 cm long, glabrous, yellow. Stamens included; filaments inserted c. 21/2 mm above the corolla-base, 6-7 mm long, shortly hairy at the base. Ovary pilose; style included, c. 8-10 mm long, glabrous.
Distr. Malesia: West (BW 9511 Mort).
Ecol. In grassland, at c. 600 m altitude. The correct reference for species 8 is: 8. Merremia quinata (R. Br.) OosTsTR. J. Arn. Arb. 29 (1948) 417, not Nova Guinea n.s. 5: 22. This combination antedates that made by Kerr, FI. Siam. En. 3, 2 (1954) 106.
Merremia borneensis MERR.
Add to Distr.: E. Borneo (KOSTERMANS 10035).
Merremia peltata (L.) MERR.
Line 15 from bottom, after ‘petioles’ add: rarely on the whole surface.
Operculina riedeliana (OLIV.) OOSTSTR. A specimen from the Malay Peninsula (Negri Sembilan), taken by RIDLEY (Fl. Mal. Pen. 2, 1923, 459) for Merremia crispatula PRAIN, belongs to O. riedelia- na. RIDLEY misquoted both collecting site and collector. Mr. H. M. BuRKILL (in litt.) was so kind to indicate the right collecting site as Bukit Dusun Paya and the collector as ALvmns (7. 1181). See my paper in Blumea 3 (1939) 368, line 1 from top.
Line 12 from bottom, replace ‘J. ochro- leuca’ by: I. ochracea.
Line 13 from bottom, replace congesta’ by: 6. I. acuminata. Bottomline, replace ‘3la. J. * sloteri’ by:
New Guinea
One
940
FLORA MALESIANA
[ser. I, vol. 66
Fig. 7. Merremia aniseiifolia Ooststr. a. Flowering stem, (a-c MOLL BW 9511).
4: 460
4: 461
4: 462
3la. I. x multifida.
Line 9 from top, replace ‘32. J. digitata’ by: 32. I. mauritiana.
Line 21 from bottom, replace ‘18. /. maxima by: 18. I. sepiaria.
Line 7 from top, replace ‘14. J. gracilis’ by: 14. I. littoralis.
Line 13 from bottom, replace ‘35. J. riparia’ by: 35. 1. rubens.
Line 2 from top, replace ‘6. /. congesta’ by: 6. I. acuminata.
x 2/3, b1-5, sepals 1-5, x3, c. pistil, x4
4: 464a Ipomoea plebeia R. Br.
4: 464
Add to Distr.: Also in tropical Africa (ssp. africana MEEUSE) and India (ssp. indica VERDC.). The specimens from Malesia and Australia belong to ssp. plebeia. See VERDCOURT in Hubbard & Milne-Redhead (ed.), Fl. Trop. East Afr., Convolv. (1963) 94.
Line 14 from bottom: the correct name for 2. Section Pharbitis is: 2. Section Ipomoea.
September 1972] Addenda, corrigenda et emendanda
94]
4: 465a The correct name for species 4 is:
ns
: 465b
: 470a
: 472a, : 56la
: 472b, : 561b
: 472b
: 473b
: 475b
: 483a
: 4845
4. Ipomoea purpurea Rotu, Bot. Abh. (1787) 27, because the synonym Con- volvulus purpureus L. Sp. Pl. ed. 2, 1 (1762) 219 is nom. illeg. This is in ac- cordance with Art. 72, note, of the Inter- national Code.
The correct name for species 6. [pomoea congesta R. BR. is:
6. Ipomoea acuminata (VAHL) R. & SCH. Syst. 4 (1819) 228, based on Convolvulus acuminatus VAHL, Symb. Bot. 3 (1794) 26. I. acuminata Ruiz & Pav. FI. Peruv. 2 (1799) 11 & pl. 120 f. b is not valid, because it was published as a superfluous name for /. angulata ORTE- GA, Hort. Matr. Dec. 7 (1798) 83.
The correct name for species 14. [pomoea gracilis R. BR. is:
14. Ipomoea littoralis BL. Bijdr. (1826) 713. I. gracilis R. BR. appears to be a distinct species, apparently rare and local, and confined to the north coast of Australia (STONE, Micronesica 1, 1964, 126; FosBerG, ibid. 2, 1966, 151-152). The correct name for species 16. [pomoea ochroleuca SPANOGHE is:
16. Ipomoea ochracea (LINDL.) G. Don, Gen. Syst. 4 (1837) 270, based on Convol- vulus ochraceus LINDL. in Edwards, Bot. Reg. 13 (1827) t. 1060.
Add to Distr.: Also in tropical Africa and perhaps in tropical America. See VERDCOURT in Hubbard & Milne- Redhead (ed.), Fl. Trop. East Afr., Convolv. (1963) 115-116.
The correct name for species 18 is:
18. Ipomoea sepiaria KOEN. ex ROxs. Fl. Ind. ed. Carey & Wall. 2 (1824) 90. I. maxima (L. f.) DON ex SWEET (Con- volvulus maximus L. f.) is a distinct species. For the typification of the latter, see VERDCOURT, Kew Bull. 15 (1961) 7. Add to Distr.: Also in tropical Africa. The correct reference for species 20 is: 20. Ipomoea pes-caprae (L.) R. Br. in Tuckey, Narr. Exp. Zaire (March 1818) 477, non (L.) Sweet, Hort. Suburb. Lond. (July 1818) 35. See STEARN, Taxon 10 (1961) 237-238.
The correct name for the hybrid 31a is: 31a. Ipomoea = multifida (RAFIN.) SHIN- NERS, Sida 2 (1966) 265.
The correct name for species 32 is:
32. Ipomoea mauritiana JAcQ. Collect. 4 (1791) 216; Pl. Rar. Hort. Schoenbr. 2 (1797) 39, t. 200. J. digitata L. is a distinct species, endemic in the West Indies, the name of which was long used for J. mauritiana.
The correct name for species 35. /po- moea riparia G. DON is:
35. Ipomoea rubens CHotsy, Mém. Soc. Phys. Genéve 6 (1834) 463. See VERD- COURT, Webbia 13 (1958) 324 and do
4: 487a
4: 488a
4: 495 4: 502a
4: 510a
: 2646, 2 Spiho)
nas
S205
SRO307
S30307D
5: 309a
in Hubbard & Milne-Redhead (ed.), FI. Trop. East Afr., Convolv. (1963) 134. Ipomoea tuba (SCHLECHTEND.) G. Don. Add to Distr.: Thailand (Pur 4326). Line 24 from top: the correct name for Ipomoea dasysperma JACQ. Eclog. Pl. 1 (Aug. 1816) 132, t. 89 is:
Ipomoea tuberculata KeR-GAWL. in Ed- wards, Bot. Reg. 1 (Febr. 1816) t. 86. See VERDCOURT in Hubbard & Milne-Red- head (ed.), Fl. Trop. East Afr., Convolv. (1963) 123.
Line 16 from bottom, replace ‘17. A. capitata’ by: 17. A. capitiformis.
The correct name for species 17. Argyreia capitata (VAHL) CuHotsy (1833), based on Convolvulus capitatus VAHL (1794), non Desv. 1792, nec Cav. 1793, is:
17. Argyreia capitiformis (PoiR.) Oost- STR. nov. comb., based on Convolvulus capitiformis Pork. in LAMK, Encycl. Suppl. 3 (1814) 469.
Argyreia congesta OOSTSTR.
Add to description: Fruit ellipsoid, 10-12 mm long, red (MUKMIN AMIR in herb. Sandakan, n. 35633).
Corynocarpaceae
Corynocarpus cribbianus (F. M. BAILEY) L. S. SMITH.
Add to Distr.: This species is also found in several of the Solomon Is.
Dichapetalaceae (LEENHOUTS)
Dichapetalum THOU.
Change the year of publication of THouars into 1806. Add to literature: HuTCHINSON, Gen. Pl. 1 (1964) 216. Add to genus diagnosis: The fruits of at least some species are apparently dehiscent (D. papuanum, D. helferianum, D. gelonioides, cf. KANNLAL & Das, FI. Assam 1, 1937, 246), exposing the orange- coloured to scarlet, thin mesocarp. Include after Wood anatomy: Phytochemistry. See HEGNAUER, Che- motaxonomie 4 (1966) 14-17.
Insert in the Key after fork 8 2nd lead: 8A. Inflorescences scorpioid.
16. D. scorpioideum 8A. Inflorescences not scorpioid.
Insert in the Key after fork 15 2nd lead: 15A. Fruits 3 cm long.
17. D. grandifolium 15A. Fruits up to c. 11/2 cm long. Dichapetalum timoriense (DC.) BOERL. Omit the questionmark before D. peekelii KRAUSE.
Add to Distr.: New Britain, Solomon Is. Add to Ecol.: Stems sometimes hollow and inhabited by ants.
Notes, 3rd paragraph: From the addi- tional notes in PEEKEL’s MS flora of the
942
FLORA MALESIANA
5: 3096
5: 310a
5: 310b
Se Silas) 52 3124
S125
Bismarck Archipelago it is clear that D. peekelii is synonymous with D. timoriense.
Dichapetalum papuanum (BEcc.) BOERL. Omit from synonymy: D. grandifolium RIDL.
Omit the headline of ssp. papuanum.
The entry on D. papuanum ssp. borneense LEENH. should be replaced by the follow- ing:
17. Dichapetalum grandifolium RIDL. Kew Bull. (1930) 373; LeENH. Blumea 12 (1963) 21. D. papuanum (BECC.) BOERL. ssp. borneense LEENH. Reinw. 4 (1956) 81; Fl. Mal. I, 5 (1957) 310.
Dioecious liana. Branchlets glabrous, purple-brown, later on greyish. Petioles 'lo-11/2 cm long, glabrous. Leaves elliptic to oblong, 13-25 by 6-12 cm, chartaceous, glabrous; glands few, small, scattered all over the lower surface of the leaf; base acute, slightly decurrent; apex more or less abruptly acuminate, acumen short and broad, blunt to acute, mucronulate; nerves 7-10 pairs, curved, most of them distinctly looped and joined. Inflorescences (Q unknown) 1-4 cm long, distinctly stalked, repeatedly branched, with several flowers. Flowers (3) 5'/2 mm long. Petals obovate, halfway incised, glabrous. Disk lobes c. 1/5-3/4 mm, 2-lobed, glabrous. Pistil- lode densely tomentose. Infructescences small, short-stalked, with 1-2 fruits. Fruits 2—3-lobed, c. 3 cm long, 214/23 cm wide, smooth, shortly and densely fulvous-tomentose, with distinct, narrow sutures.
Distr. Malesia: Borneo (Sarawak and North Borneo).
Ecol. In primary forests at low alt. Fi. June, July, Nov., fr. Nov.
Note. Possibly nearest allied to D. papuanum which it distinctly resembles especially vegetatively and in the flowers (apart from the larger dimensions). Well characterized by the exceptionally large fruits.
Dichapetalum gelonioides (ROxB.) ENGL. Add to literature: KANJILAL & Das, FI. Assam 1 (1937) 245, and sub Chailletia sumatrana MiqQ.: TALBOT, For. FI. Bombay Pres. 1 (1909) 253, f. 151.
ssp. tuberculatum LEENH.
Add to Distr.: Sumatra.
ssp. pilosum LEENH.
Add to Distr.: Malay Peninsula (Selan- gor).
Add to Notes: The fruits are sometimes reported as being poisonous. Dichapetalum tricapsulare (BLCO) MERR. After the entry on D. glabrum Exo. add: non D. glabrum (VAHL) PRANCE (1968), nom. illeg.
5
[ser. I, vol. 66
: 313a
: 313b
: 314ab Dichapetalum
: 3145
oloa
e) SSD
Dichapetalum griffithii (HooK f.) ENGL. Change in description the minimum number of nerves into 8.
Dichapetalum setosum LEENH.
Add to Ecol.: Altitude up to 360 m. Dichapetalum steenisii LEENH.
Add to description: Sometimes a shrub. Leaf base acute to blunt.
Add to Distr.: E. Borneo (ssp. steenisit). longipetalum (TURCZ.) ENGL.
Add to literature: CHUN & CHANG, Fl. Hainan 2 (1965) 203, f. 407, and sub D. hainanense ENGL.: CHUN & How, Act. Phytotax. Sin. 7 (1958) 16.
Change in description: Branchlets some- times nearly terete. Leaves ovate- to obovate-elliptic to -oblong, above some- times glabrous.
Line 15 from top, replace ‘1 (—2) —lobed’ by: 1 (3) -lobed.
Add to Distr.: S. China, SE. Thailand. Dichapetalum helferianum (KURZ) PIERRE. Add to Distr. after Malay Peninsula: Peninsular Thailand.
Dichapetalum laurocerasus (Hook. f.) ENGL.
Change in description: Leaves up to 15 cm long, thin-coriaceous to charta- ceous; acumen short to rather long; nervation mostly inconspicuous. Dichapetalum sessiliflorum LEENH. Replace the description by the follow- ing: Dioecious liana, shrub, or up to c. 5 m high treelet. Branches densely fulvous-tomentose to -velutinous when young, glabrescent, greyish to purplish brown. Leaves elliptic to lanceolate, 9-28 by 3-11 cm, pergamentaceous to chartaceous, above glabrous to hairy on midrib and nerves, beneath sparsely to densely appressed-pilose at least on midrib and nerves; glands few, on the lower side, mainly near the base; base acute to rounded; margin minutely crenulate to entire; apex acuminate, acumen short, broad, and blunt to long, slender, and acute; nerves 7—16 pairs, slightly to strongly curved, at least the upper ones distinctly looped and joined. Flowers (only 3 buds known) axillary, 1 or 2, subsessile, or in short-stalked, few- to several-flowered glomerules, in vivo pale pink. Calyx densely ferruginous- tomentose. Petals ovate, slightly emar- ginate, outside (margin excepted) and inside at the base long-pilose. Disk annular, adnate to the stamens. Pistillode patently stiff-pilose. Fruits solitary, short- stalked, triangular-ovoid, c. 3 by 21/2-3 cm, smooth to tuberculate, densely and shortly ferruginous-tomentose, glabres- cent, without sutures, in vivo orange when ripe, 3 (-1) -celled; stones free, woody, strongly corrugated.
September 1972]
Addenda, corrigenda et emendanda
943
5: 3l6a
4: 379a
Add to Distr.: Vogelkop Peninsula, New Britain, Solomon Is. (Three Sisters L., BSIP 17224).
Change in Ecol. altitude: from sea-level to 1800 m. Add: Fi. Febr., Oct.-Nov.,
fr. March-April, Oct.-Nov.
Add: Uses. The leaves are medicinally used (‘chewed and the extract spat onto wounds to relieve soreness’; FRODIN, New Britain); the fruits are edible. Dichapetalum tenerum LEENH. Add to Ecol.: Fr. July-Aug. Add after 15. Dichapetalum LEENH. :
tenerum
16. Dichapetalum scorpioideum LEENH. Blumea 13 (1965) 162.
Dioecious (?) liana. Branchlets densely fulvous-tomentose, glabrescent, purple- brown, sparsely lenticellate. Leaves c. 1 cm long petioled, blade elliptic, 16-20 by 9-11 cm, chartaceous, when young fulvous-hairy mainly on midrib and nerves, later subglabrous, lower surface with scattered small glands mainly in the basal part; base rounded, slightly atten- uate; apex rather gradually, shortly, broadly, and bluntly acuminate; nerves 6-8 per side, curved, only the upper 2 or 3 more or less distinctly looped and join- ed near the margin. J/nflorescences shortly (2!/2-5 mm) and thickly pedun- cled, with 2 spirally recurved, c. 11/2 cm long, densely and shortly fulvous-hairy cincinni which are densely covered on one side with two rows of alternating flowers, on the other side with the bracts. Flowers seen in bud only, 3 unknown. Petals rather deeply bifid, outside sparsely appressed-hairy. Disk lobes low, broad, slightly bi- to trifid, long woolly- ciliate. Ovary densely fulvous woolly, 2-celled; style 1, cylindrical, short, with 2 spreading stigmas. Fruits about semi-elliptic, flattened, 20 by 13 by 9 mm, densely fulvous-velvety; pericarp with broad suture; 1-seeded.
Distr. Solomon Is. (Treasury Group, Mono I.).
Ecol. Secondary forest sandstone slope. Fi. fr. April.
Note. The relationships of this species are uncertain. The kind of inflores- cence, its most distinctive character, is unique among the Asiatic and Pacific spe- cies, but is known from some African species.
on rocky
Droseraceae
Drosera petiolaris R. Br.
Of this rare species a second collection was made in the Western District, T.N.G., in an open grass-sedge plain, growing on sand over clay, rosettes over 14 cm @,
locally common c. 1 mile south of Morehead Patrol Post, along track to Tonda, Aug. 8, 1967, R. PULLEN 7139.
4: 379ab Drosera spathulata LABILL.
6: 423 6: 426b
6: 469
6: 474
6: 474
6: 480
6: 661
6: 662 6: 663a
This species has for the second time been found at low altitude in Malesia, viz in Sarawak, near Telok Asam, Bako National Park, 120 m, by Prof. Purse- GLOVE, on mud by sides of streams. Bako National Park is situated on generally poor, podsolized soils carrying heath forest. The size of the flower dissected is slightly smaller than described in FI. Mal.: sepals 2 mm, petals pink, nearly 2 mm; but otherwise the specimen shows no deviations.
Epacridaceae
Line 8 from top, replace ‘R. Br.’ by ‘(RA BR) I SPRee
Styphelia malayana (JACK) Spr.
Add to synonymy: Leucopogon ophirensis GriFF. J. As. Soc. Beng. 23, ii (1854) 638.
Ericaceae (SLEUMER)
Line 13 from bottom replace ‘ Wirtgenia’ by: Andresia.
In Key to the genera (line 9) replace ‘2. Wirtgenia’ by: 2. Andresia. Rhododendron L.
It has appeared that there are still new species of Rhododendron, amongst others from Borneo and New Guinea. I am planning to give a supplement in future but not of other Ericaceous genera.
Line 2 from top replace ‘3. Subg. Pentanthera sect. Pentanthera by: 3. SuBG. ANTHODENDRON sect. Antho- dendron.
Line 16 from top, omit ‘(BL.)’.
Line 18 from top, after ( Vireya’), insert: non RAFIN. 1814.
Line 18 from top, omit ‘(BL.)’.
Replace the infrageneric epithet 3. Sub- genus Pentanthera by: 3. Subgenus An- thodendron (RcuB.) ENDL. ex WILS. & REHDER, Monogr. Azaleas (1921) 115. Replace the infrageneric epithet 1. Sec- tion Pentanthera by: 1. Section Anthoden- dron (RcHB.) ENDL. Gen. (1839) 759.
In Key to the species line 8 read: 283. R. macrosepalum.
Replace the name 283. Rhododendron linearifolium S. & Z., non Por. in LAMK, Encycl. 6 (1804) 267, by: 283. Rhodo- dendron macrosepalum Maxim.
Flacourtiaceae (SLEUMER)
Scolopia SCHREB.
I am engaged in a new revision of this genus. It has appeared that the new record of S. kermodei C.E.C. FISCHER
944 FLORA MALESIANA [ser. I, vol. 68 (hitherto known from Burma and Anda- finely reticulate, crenate, 6-11 by 4-8 cm, mans) from Malaya (cf. Blumea 17, with 1 or 2 small glands at base near the 1969, 270), has proved to belong to an apex of the petiole, the latter slender, undescribed species. 1-1!/2 cm. Racemes 4—8-flowered, rachis
5: 14a Paropsia vareciformis (GRIFF.) MAST. (5 mm or less) and pedicels (11/2-2 mm) Add to synonymy: Alsodeia chrysodasys puberulent. Perianth segments 4, sub- Mia. Fl. Ind. Bat. Suppl. 1 (1861) 390; orbicular-ovate, puberulent outside, ci- Ann. Mus. Bot. Lugd.-Bat. 5 (1869) 215. liate, c. 2 mm. Disk shortly 8-lobed. Cf. Jacoss, Blumea 15 (1967) 137; 3 Flowers: stamens c. 25, exserted; SLeum. Bull. Jard. Bot. Brux. 40 (1970) filaments unequal, 2-2!/2 mm; ovary 67, f. 5 (distr.). rudimentary, 0.7 mm. 2 Flowers: ovary
5: 35b, Scaphocalyx spathacea RIDL. ovoid, attenuate at apex, glabrous, 2-3
565a Add to Distr.: E. Borneo (Berao). mm; stigmas 2, sessile. Fruit not known.
5: 39a Trichadenia philippinensis MERR. Distr. Malesia: Philippines (Palawan), Add to synonymy: The plant distributed twice found. under the name Neotrewia arborea ELM. Ecol. On limestone hill. nomen in sched., which was referred by Note. X. palawanense is distinguished MERRILL to Neotrewia cumingii (M.A.) from X. luzonense by the broadly ovate, P. & H. (cf. MerRR. En. Philip. 2, 1923, at base truncate, rounded or slightly 437), has appeared to belong to 7. cordate leaves, which bear 2 basal pairs philippinensis. of lateral nerves.
5: 51 Line 17 from bottom replace ‘Sect. 5: 80 Under Fig. 35d read 3 instead of 9. Pythagorea’ by: SECT. BLACKWELLIA. 5: 95a Casearia grewiaefolia VENT.
5: 52 Line 14 from bottom replace ‘Subgenus Replace the epithet var. deglabrata Pythagorea (LoUR.) SLEUM.’ by: Sub- K. & V. by: genus Blackwellia (LAMK) WARB. var. gelonioides (BL.) SLEUM. comb. noy. Line 8 from bottom replace ‘Section 5: 985 Casearia pallida CRA‘.
Pythagorea (Lour.) O.K. *by: Section Add to Distr.: Malay Peninsula (Se- Blackwellia BENTH. langor).
5: 65b Bennettia papuana GiLG is reduced to 5: 105 Under Excluded: Dovyalis macrodendron Blumeodendron papuanum P. & H. GILG, which was already tentatively (Euphorbiaceae). Cf. Airy SHAW, Kew removed from Flacourtiaceae, has ap- Bull. 16 (1963) 349. peared to belong to Suregada ROTTL. (Ge-
5: 65b, Hemiscolopia trimera (BOERL.) SLOOT. lonium) of the Euphorbiaceae. Cf.
566a Add to Distr.: Peninsular Thailand STEEN. Nova Guinea, Bot. n. 12 (1963) (Nakon Srithamarat). 190. 5: 685 Add after 3. Xylosma luzonense (PRESL) CLOs, etc.: Gnetaceae (MARKGRAF, Zurich) 3a. Xylosma palawanense MENDOZA, Phi- lip. J. Sc. 93 (1964) 514, f. 2. 4: 337 Add before Vern.:
Shrub or tree, up to 7 m, glabrous, Palynology. ERDTMAN (Bot. Notis. whether or not armed. Leaves broadly 1954, 80) has found that the subdivisions ovate, apex subacuminate, base truncate, of the genus have separate pollen types. rounded or slightly cordate, coriaceous, 4: 338 Replace the keys by the following:
KEY TO MALE PLANTS
1. Trees or shrubs, only occasionally and partly climbing. Leaves thin, yellowish when dried. Inflorescence yellowish; collars flat, almost always conspicuously distant from each other.
Sect. Gnetum subsect. Eugnemones.
NN
neither velvety nor whitish
Trees. Sterile 2 flowers ovate, long-beaked; beak finely velvety, whitish. 2. G. costatum . Trees or shrubs. Sterile 2 flowers sas pe (only in var. griffithii beaked), the tip
1. G. gnemon
3. Shrub. Inflorescence simple, slender, its | axis scarcely Ven mm , thick, flower clusters up to
2mm @
var. tenerum
3. Trees or shrubs. Inflorescence simple or ‘branched, thick, its a axis dn mm \ thick, flower clusters
up to5mm @.
4. Trees. Inflorescences almost always branched, all collars distant
var. gnemon
4. Shrubs. Inflorescences always simple, often only the lowermost collars distant (inflores-
cence unknown in var. gracile).
5. All collars distant. Sterile 2 flowers beaked (oblong, glabrous)
var. griffithii
5. Collars at least partly contiguous. Sterile 9 flowers shortly acuminate. 6. Only the uppermost collars contiguous. Sterile 2 flowers globose with short tip.
var. brunonianum
September 1972] Addenda, corrigenda et emendanda 945
6. All collars contiguous or only the two lowermost distant. Sterile Y flowers tapering. var. ovalifolium 1. Lianas. Leaves brown or black when dry, coriaceous (thin only in 6. G. neglectum and 3. G. tenuifolium). Inflorescence not yellowish, its collars always fairly approaching each other (the axis never visible between them), their edges bent upward!. Sect. Cylindrostachys MARKGR. 7. Collars of inflorescence dish-like, the flowers coming out freely. 8. Leaves with numerous spicular cells parallel to the secondary nerves, therefore silky above when dry. Stamen with one microsporangium only . . . . . 1. G. gnemonoides 8. Leaves with few or no spicular cells. Stamen with two microsporangia. 9. Leaves thin, green when dry, large, elliptic. Inflorescence simple (rarely once branched), slender (3 mm broad), spike itself 2cmlong .... . . . . 3. G. tenuifolium 9. Leaves coriaceous. Inflorescence always branched.
10. Leaves small, obovate and tailed, distinctly reticulate, black when dry, striate above by spicular cells. Inflorescence once branched, slender (3 mm broad), spike itself 1-1+/2 cm long) = . . 5. G.arboreum
10. Leaves often large and broad, not striate, "mostly black when dry and inconspicuously reticulate. Inflorescence branched several times, thicker (4 mm broad), spike itself up
to 4 cm long : . . 4. G. latifolium 11. Leaves elliptic, brown when dry, distinctly reticulate, secondary nerves distinctly
joined : . . var. funiculare 11. Dry leaves nigrescent, “inconspicuously reticulate, secondary nerves ending open. 12 seaves almost! orbiculanm ents) lesen ee Ge ee. oo Vaseelaxiintescens 12. Leaves elliptic.
13. Leaves small, up to9 cm. Spike short,11/2cmlong . . . . . . . var. minus
13. Leaves large, up to 20 cm. Spike 2-4cmlong . .. . . . ._ var. latifolium
7. Collars of inflorescence cylindric, keeping the flowers enclosed. 14. Inflorescence branched, mostly large, with often very long stalks. Exserted part of the stamen longer than the perianth. 15. Dried leaves black, with narrowed base, relatively rich in fibres. Inflorescence 15—20 cm long, catkins twice the length of their stalks. Sterile 2 flowers obliquely conical. 9. G. ridleyi 15. Dried leaves brown, with rounded base, without conspicuous fibres. Inflorescence 30 cm long (in var. abbreviatum 4-6 cm), catkins as long as their stalks. Sterile 2 flowers ovate, straight . ire) eh ee oe al Galeptostachyum 16. Catkins at most 34 mm broad and 4 cm long. 17. Leaves large, up to 30 by 12 cm. Inflorescence 30 cm long, catkins 3-4 cm long. var. leptostachyum 17. Leaves small, not more than 12 by 6 cm. Inflorescence 4—6 cm long, catkins 1!/2cm Jong. var. abbreviatum 16. Catkins at least 4mm broad and upto6cmlong . . . . var. robustum 14. Inflorescence unbranched (once branched in 6. G. neglectum), often cauline. Exserted part of the stamen shorter than the perianth. 18. Leaves thin, tapering at both ends. Inflorescence slender (3 mm). 6. G.neglectum 18. Leaves coriaceous. Inflorescence thick (4-5 mm). 19. Inflorescences 6 cm long, drooping (unknown in 8. G. k/ossii from Borneo, but its rough @ one is of this type). Flowers numerous, imbedded in many hairs. 7.G.cuspidatum 19. Inflorescence short, mostly erect (2-3 cm). 20. Inflorescence very thick (7 mm). Leaves large. 21. Leaves oblong-obovate, somewhat silky above by spicular cells. Flowers immersed between few hairs. S02) 22 10 Gi loerzineh 21. Leaves elliptic, not silky. "Flowers. immersed ‘between numerous long hairs. 13. G. macrostachyum 20. Inflorescence moderately thick (4 mm), shortly stalked, mostly erect. Leaves small, up to 15 cm long. 22. Leaves firm (not fleshy), with distinct nervation, not glaucous, not cuneate. In- florescence 3 cmlong . . oy 2G: tla 22. Leaves fleshy, with indistinct nervation, “more or ‘less glaucous. Inflorescence 11/2—2 cm long. 23. Leaf base mostly acute, leaves often lanceolate. Collars of inflorescence with angular lower edge. Fruit short-acuminate . . . . . . . . . 14. G.microcarpum
(1) 3 Inflorescence unknown in 10a. G. raya MARKGR., 1la. G.globosum MARKGR., and 7a. G. acutum MARKGR.
946 FLORA MALESIANA [ser. I, vol. 68
23. Leaf base mostly rounded. Collars of inflorescence with vaulted lower edge. Fruit loMe-ACUMIMALE ps GE 8 ee ae Gage, 6 ee
KEY TO FEMALE PLANTS
1. Trees and shrubs, only occasionally or partly climbing. Leaves thin, yellowish when dry. Inflorescence yellowish, collars flat. Fruit almost velvety. Sect. Gnetum subsect. Eugnemones.
2. Tree. Flowers ovate, long-beaked; beak finely velvety, whitish . . . . 2. G.costatum 2. Tree or shrub. Flowers globose, shortly lipped: ae in var. hac tii beaked), the tip not velvety nor whitish . . ; . . . 1. G. gnemon
3. Collars of inflorescence remote. 4. Flowers globose. Fruit ovate, obtuse. . Tree. Inflorescence mostly branched, all its internodes long (*/2-1 cm). Fruit large (2 cm long) Poti . . var. gnemon 5. Shrub. Inflorescence simple, ‘at least its two lowermost ‘internodes long, all others short and hidden. Fruit small (1 cm long), inserted on a thickened rachis. var. brunonianum 4. Flowers oblong, beaked (unkown in var. gracile). Fruit acute. 6. Fruit oblong. Axis of inflorescence thick (1 mm); internodes 1/2 cm long. var. gracile 6. Fruit ovate. Axis of inflorescence slender (1/2 mm); internodes 11/2 cm long. var. tenerum 3. Collars of inflorescence contiguous. Inflorescence short. Flowers acute. 7. Flowers globose, beaked. Fruit globose .. . a ges. ome. voRentirchin 7. Flowers ovate, acuminate. Fruit ovate, long- acute i . . var. oyalifolium . Lianas. Leaves coriaceous, brown or black when dry (thin only in 6. G. neglectum and 3. G. tenuifolium). Inflorescence not yellowish. Collars dish-like. Fruit smooth or warty, not velvety. Sect. Cylindrostachys MARKGR. 8. Inflorescence branched (unknown in 5. G. arboreum, but the 3 one branched, small). 9. Leaves obovate-cuneate, tailed, small, distinctly nerved below, densely striate by spicular cells above. Fruit long- stalked ew . . 5. G.arboreum 9. Leaves broadest in or below the middle, mostly large, ‘not densely striate above. 10. Leaves brown when dry, secondary nerves distinctly joining. 11. Nerves all remote, arcuate. Inflorescence rich and spreading, often 30 cm long, in var. abbreviatum much shorter. Flowers globose, shortly tipped embedded in dense hairs.
Fruit sessile. . . 16. G. leptostachyum 12. Leaves large, up to 30 by I2¢ cm. ‘Infructescence 20-40 cm long, its internodes 8-12 mm long.
13. Infructescence 20-25 cm long. Fruit ellipsoidal, 2 by 11/2 cm. Internodes 12 mm long. var. leptostachyum 13. Infructescence 30-40 cm long. Fruit parade eis 1.8 2 1.3 cm. Internodes 8
mmlong . . . . var. robustum 12. Leaves small, not 1 more than 12 by 6 cm. Infructescence 10 cm 1 long, its internodes 5 mm long. Fruit 21/2 by 11/2 cm. . . . var, abbreviatum
11. Secondary nerves at the leaf base approaching each other, all with a straight lower part. Inflorescence not so rich, 15 cm abs Flowers obliquely beaked, conical, embedded in few hairs. Fruit stalked aebets . . 4. G. latifolium var. funiculare
10. Leaves black when dry, secondary nerves ’ indistinctly joining. 14. Leaves with conspicuous spicular cells, though not striate. Flowers embedded in
numerous hairs. Fruit very large (6 cm), obtusely turbinate, sessile . . 9. G. ridleyi 14. Leaves without conspicuous spicular cells. Flowers not embedded in numerous hairs. Fruit elliptic, up to 21/2 cm long, stalked. . . ‘ . . 4 G. latifolium
15. Leaves almost or quite orbicular. Fruit oblong- ‘obovate, rather long-stalked. var. laxifrutescens 15. Leaves elliptic. Fruit stalk thick. 16. Leaves small (not longer than 9 cm). Fruit ovate, small (11/2 cm long) var. minus 16. Leaves large. Fruit large, 2-21/2 cm, broadly ovate, long- or short-stalked. var. latifolium 8. Inflorescence simple (exceptionally once branched in 6. G. neglectum). 17. Leaves thin, tapering at both ends. Secondary nerves straight and broken. 18. Secondary nerves remote but not extremely so. Spike short (4 cm). Collars contiguous. Fruit 2 cm long, longitudinally furrowed, acute, with a long, slender stalk. 3. G. tenuifolium 18. Secondary nerves extremely remote (up to 3 cm). Spike long, 8 cm. Collars not contigu- ous. Fruit small, brownish yellow, sessile, smooth, obtuse, 11/2 cm long. 6. G. neglectum 17. Leaves coriaceous, secondary nerves bent, not broken. 19. Leaves silky above by numerous parallel spicular cells. Flowers obtuse. Fruit obtuse, large, Warty.). 30. kw wl os we ww mtn vk ce
September 1972]
4: 340 4: 343a
4: 343a
4: 344a
Addenda, corrigenda et emendanda
947
20. Tertiary nervation of leaves indistinct. Fruit smooth.
21. Leaves elliptic, 15 by 7 cm. Fruit ellipsoidal, 5 by 3 cm
. 10a. G. raya
21. Leaves lanceolate, 8-9 by 2!/2 cm. Fruit globose, 41/2 ‘cm @. (la. G. globosum 20. Tertiary nervation of leaves distinct below. Fruit verrucose, S-6 by 2-3 cm.
11. G. gnemonoides
19. Leaves not silky by spicular cells. Flowers acuminate. Most inflorescences cauline. 22. Leaves fleshy, with indistinct nervation, more or less glaucous.
23. Flowers ovate. Fruit more or less obtuse, yellow, 2 cm long. 14. G. microcarpum
23. Flowers oblong-conical. Fruit long-acuminate, 21/2 cm long, pink. 15. G. oxycarpum 22. Leaves firm, not fleshy, not glaucous, with distinct nervation.
24. Leaves obovate-cuneate, striate above by spicular cells. Flowers obtuse, not embedded
in thick hair masses. Fruit obtuse, 4 cm long
10. G. loerzingii
24. Leaves elliptic, not striate above. Flowers embedded i in thick hair masses.
25. Leaves broad-elliptic. Fruit large, acute, rough
8. G. klossii
25. Leaves twice as long as broad. Fruit not rough. 26. Collars contiguous, their hair tufts enormous. Fruit small, up to 2 cm long, almost
globose
13. G. macrostachyum
26. Collars remote, hair tufts large, but not enormous. 27. Leaves large. Inflorescence elongate, fruiting 9-15 cm long. Fruit 2!/2-31/2cm long.
28. Fruit shining, broad-ovate, obtuse, 21/2 by 11/2 cm
. 7. G. cuspidatum
28. Fruit opaque, fleshy, acute, slightly bent upward, 31/2 by 1.8 cm. 7a. G. acutum
27. Leaves small. Inflorescence short. Fruit small, elliptic,
Line 13 from top Gnemonomorphi’ by:
Gnetum neglectum BL. Add to literature: MARKGR. Blumea 10 (1960) 431; ibid. 19 (1971) 108.
Add to description: ¢ Inflorescence often cauliflorous, branched once, its branches slender, 2-4 cm by 1 mm; catkins 3-5 cm by 3 mm. Collars cylindric-infundibuli- form, 3 mm high. ¢ Flowers numerous, obconical, 2 mm high; stamen bilocular, cells white, splitting on their top. Sterile 2 flowers 4-6, ellipsoidal, short-acute, 11/2 mm long; involucre chartaceous; ovule obliquely ovoid, gradually acumin- ate, 1 mm high. Add to Distr.:
Sarawak, S_ coll. 28459).
Gnetum cuspidatum BL. — Fig. 8.
Add to literature: MOLESWORTH ALLEN, Mal. Nat. J. 18 (1964) 168-169, 3 photogr. (finely illustrated description of 3 in- florescence).
replace ‘Section Section Gnetum.
Borneo (Brunei, S 5752; 202; Sabah, SAN
7a. Gnetum acutum MARKGR. 0m. nov. — G. acutatum MARKGR. Blumea 13 (1966) 404, non Mia. 1860.
Large liana, 25 m by 2!/2cm @; stems sulcate and densely lenticellate; twigs terete, smooth. Leaves glabrous, elliptic, acuminate, 11-17 by 6-7/2 cm, firmly chartaceous; nerves 9 pairs, arcuate, united before the margin, prominent beneath; petiole 12 mm. 2 /nflorescence caulifiorous, not branched, in fruiting state 9 cm, rachis 5 mm @; peduncle 8 mm; collars initially infundibuliform, later 1 cm wide. 3 Flowers unknown. © Flowers immersed in a dense hair-cush- ion, cream flushed with pink, acute-
4: 344a
4: 3445
11/2 by 0.8 cm. 12. G. diminutum
ovoid, apex upcurved; external involucre fleshy, 4 by 2 mm, !/2 mm thick, median one conical, thin, 2 by 1 mm, internal one chartaceous, ampullaceous, 11/2 by 0.8 mm, lengthened into a thin apical tube 2 mm exserted, fid at apex. Fruit acute-ovoid, slightly curved, 31/2 by 13/4 cm; external involucre fleshy, 1 mm thick, median one hard, glossy and fibrous, internal one chartaceous, glossy; nucellus with embryo 21/2 by 3/4 cm, acute-ovoid.
Distr. Malesia: West Borneo: Sara- wak (Rejang delta, ASHTON S 17804).
Ecol. Alluvial forest.
Vern. Layah, Iban.
Note. Allied to G. cuspidatum by the simple, cauliflorous inflorescence with thick axis and densely hairy pulvini, fruit-size and leaf-shape. Differs by the acute fruit with dull, fleshy external involucre; also by more acute @ flowers, farther exserted micropylar tube, and obviously thinner leaves.
Gnetum ridleyi GAMBLE ex MARKGR. Add to literature: Blumea 12 (1963) 78. Add to description: 3 Inflorescence 15 cm (or longer), widely divaricate-branched; fertile parts 3-5 cm, twice as long as peduncle; collars numerous, 2 by 4 mm, cylindric. Flowers interspersed with numerous hairs; ¢ flowers obconical, much narrowed to base, 11/2 by 3/4 mm; sporophyll thin, 2 mm exserted; sporan- gia 2, ovoid. Sterile 2° flowers 6, obliquely conical, 11/2 by 1 mm, external and inter- nal involucre strongly fibrous (E. Malay Peninsula, P. Tioman, KADIM & Noor 644).
10a. Gnetum raya Markar. Blumea 14 (1966) 284.
Fig. 8. Gnetum cuspidatum BL. Abundant setting of seed, in Sarawak, near Bintulu; * '/10 (DING Hou 321).
September 1972]
Addenda, corrigenda et emendanda
949
4: 3445
Large, glabrous liana, 20 m; twigs terete, smooth, lenticellate below the thicker nodes. Leaves elliptic, coriaceous, at base narrowed into petiole 1 cm, at apex short-acuminate, up to 15 by 7 cm, with parallel fibres visible on upper sur-
face, grey-shining; nerves 6-8 pairs, arcuate-joining 1 cm from margin, prominent beneath, veins indistinctly
reticulate. ¢ Unknown. Fruiting 2 in- florescences often several at one node, 3-4 cm by 4-5 mm, collars dish-like, 6-8 mm ©. Fruit immersed into cushions of short hairs, ellipsoidal, obtuse, 5 by 3 cm, attenuated at base into a hollow pseudostipe 5-10 by 8 mm, opaque, finely punctulate, grey-green, inside with a sour smell. Outer involucre fleshy, 2 mm thick, fibrous at the inner surface, middle one coriaceous, longitudinally sulcate, inner one chartaceous. Seed (unripe) oblong, 30 by 8 mm.
Distr. Malesia: Central Borneo (Up- per Kapuas, Bt Raya, Sibu, S 23801); Sarawak (Kuching, near Matang, 500 m, mixed Dipterocarp forest, S 25646).
Ecol. On sandy clay, 200 m.
Vern. Akar téngang, Iban.
Notes. Allied to the Sumatran G. loerzingii MARKGR., with which it shares a large-fruited group with fibrous leaves and fruits (G. klossii MErRR., G. ridleyi GAMBLE, G. gnemonoides BRONGN., G. cuspidatum BL.). The peculiar smell of the seed is also found in G. loerzingii. This species differs by obovate-cuneate leaves and by a rough fruit surface.
lla. Gnetum globosum MarkGr. Blu- mea 19 (1971) 108.
Liana with terete, smooth twigs. Leaves coriaceous, light brown when dry, lanceolate, 8-9 by 2-2!/2 cm; nerves straight, at an angle of 30° with the midrib, arcuate-joining !/2 cm before the margin, hidden among numerous parallel fibres on either surface; petiole !/2 mm. 3 Inflorescence unknown. Fruiting ° inflorescence axillary, simple, peduncle 7 mm, rachis 13-23 by 3 mm, with 6-8 mm long joints; collars dish-like, 1 by 4 mm. Fruit globose, smooth, grey-green, 41/2 cm, contrasted at base; outer invo- lucre rather fleshy, 2 mm thick, fibrous inside; middle one 1/2 cm thick, outside fibrous, inside smooth; inner one chartaceous, outside sparsely fibrous, inside smooth. Seed globular, 21/2cm ©.
Distr. Malesia: Malay Peninsula (Pahang: Ulu Sat, FRI 15262), one collection.
Ecol. Riverside forest, 120 m. Fr.
July. Note. Approaching G. ridleyi GAMBLE
4: 346b
52335
from Pahang by its rather large fruit and fibrous leaves; G. ridleyi differs by much larger and less fibrous leaves, by ramified inflorescences and larger, spindle-shaped fruits. G. gnemonoides BRONGN., of the same group, has lenticellate fruits and leaves that beneath are not fibrous but reticulate.
Gnetum leptostachyum BL.
Add the following variety:
var. abbreviatum MARKGR. Reinwardtia 1 (1952) 462.
Leaves up to 12 by 6 cm, coriaceous, beneath distinctly reticulate-veined. g Inflorescence 4-6 cm, branched. ¢ Cat- kins 11/2 cm by 3 mm. Infructescence up to 10 cm (internodes 5 mm). Fruit large, 21/2 by 11/2 cm.
Distr. Malesia: N. Borneo, Mt Kinabalu, 1200-1800 m (CLEMENs 32990, type; 32276, 32475, 32488, 32601, 32698, 32991).
Note. Some inadequate specimens I have identified formerly as G. diminutum MarkGr.; the branched inflorescences, however, prove them to belong to G. leptostachyum, \t is a mountain variety with reduced size of leaves and inflores- cences, combining the narrow, short 3 catkins of the lowland variety /eptosta- chyum with the short-jointed, large- fruited 2 catkins of the lowland variety robustum.
Goodeniaceae (LEENHOUTS)
Add to family diagnosis: Leaves some- times verticillate.
Under Distribution the number of Malesian species of Scaevola, apart from the littoral S. taccada, should be changed into 5.
Add to Pollination: See also CAROLIN, Proc. Linn. Soc. N.S.W. 85 (1960) 197-207. G. W. GILLETT, in a letter dated 1-9-1964, on Hawaiian Scaevola informed me: ‘Incidentally the S. taccada populations around here are strictly inbreeders, the indusium being filled with pollen before the corolla opens, after which the indusium closes over the pollen-filled stigma so that there is no chance that foreign pollen could ever make contact with the stigmatic surface. I have checked this in a very large num- ber of flowers in widely separated populations. Our Hawaiian endemics do not behave this way.’
Add to Anatomy: CARLQuisT, Ann. Mo. Bot. Gard. 56 (1970) 358-390 (moreover a very interesting general paper).
Add after Anatomy:
Phytochemistry. See HEGNAUER, Che- motaxonomie 4 (1966) 212-215.
950
357506
5: 336a
5: 336
5: 337b
Add to the introduction to the family:
FLORA MALESIANA
[ser. I, vol. 66
Morphology. CAROLIN, Proc. Linn. Soc. N.S.W. 84 (1959) 242-255, on the anat- omy and especially vasculation of the flowers. CAROLIN, ibid. 91 (1966) 58-83, on the morphology of fruit and seed as well as on the systematics of the family in general.
Cytology. See PEACOCK, Proc. Linn. Soc. N.S.W. 88 (1963) 8-27.
Velleia J. E. SMITH.
Add to literature: CAROLIN, Proc. Linn. Soc. N.S.W. 92 (1967) 51.
Velleia spathulata R. BROWN.
Add to literature: CAROLIN, Proc. Linn. Soc. N.S.W. 92 (1967) 51.
Description, line 3, replace ‘herbaceous’ by: fleshy.
Goodenia J. E. SMITH.
Distr., change to: three species known from outside Australia/Tasmania.
Add the following key:
KEY TO THE SPECIES
1. Plant 20-60 cm high. Leaves linear- lanceolate, up to 22 cm long. Inflores- cences many-flowered.
3. G. purpurascens
1. Plant up to 10 cm high. Leaves ovate or obovate, up to 6!/2 cm long. Flowers solitary.
2. Plant glabrous. Leaves coarsely dentate 1. G. koningsbergeri 2. Plant densely hairy. Leaves entire. 2. G. pumilio After Goodenia koningsbergeri add:
2. Goodenia pumilio R. Brown, Prod. (1810) 579; BANKs & SOLAND. Bot. Cook’s Voy. 2 (1901) t. 177 f. A; Krause, Pfi. R. Heft 54 (1912) 93; RipsDALE, Trans. Papua N.G. Sci. Soc. 9 (1968) 18. — Fig. 9.
Erect or prostrate, fairly densely stel- late-hairy, annual or perennial herb. Leaves radical-rosulate; (broad-)ovate to spathulate, up to 6 by 3 cm but usually much smaller, herbaceous, attenuate at base; margin entire; apex blunt. Flowers solitary, axillary, 4-5 mm long, hairy; pedicel c. 1 cm long; bracteoles 0. Corolla c. 3 mm long, dark reddish. Capsules (ovoid-)oblong, 3-4 mm long. Seeds many, orbicular, small.
Distr. Australia (Northern Terr., Queensland) and Malesia: New Guinea (Papua, Western Distr., near Weam, RIDSDALE & GALORE NGF 33733; Bula village, PULLEN 7009).
Ecol. Savannah grassland, c. 10-35 m. Fi. fr. Aug.
Note. We owe the identification of this and the following species to Prof. R. C. CAROLIN, Sydney.
Fig. 9. Goodenia pumilio R.Br. a. Habit, 2, b-c.
x10, d. corolla, from above, x10, e.
style and stigma, 20, f. calyx with fruit, «10, g.
seeds, 8 (a—g PULLEN 7009).
3. Goodenia purpurascens R. BROWN, Prod. (1810) 578; Krause, Pfl. R. Heft 54 (1912) 91; F. M. BAILEY, Compr. Cat. Queensl. PI. (1913) f. 254.
Erect, sparsely hairy to subglabrous, annual or perennial herb. Leaves mainly radical rosulate; linear-lanceolate, up to 22 by 1/4-13/4 cm (the few cauline ones much shorter), somewhat fleshy, sessile with a slightly narrowed base; margin subentire, with a few minute teeth; apex acute. /nflorescences terminal, up to 25 by c. 5 cm, thyrsoid, repeatedly laxly branched and many-flowered. Flowers 10-15 mm long, thin-hairy to nearly glabrous; pedicel c. °/4 cm _ long; bracteoles 0. Corolla c. 3/4-11/4 cm long, pink to purple. Capsules ovoid, 3-4 mm long. Seeds many, lenticular, minute.
Distr. Australia (Northern Terr.,
September 1972]
an
: 3385
: 3385
: 3395 = 839
Addenda, corrigenda et emendanda
eI
Queensland) and Malesia: New Guinea (Papua, Western Distr., Wassi Kussa R., Henty & KATIK NGF 38744).
Ecol. Savannah, in shallow
alt. a few metres. F/. July. Calogyne pilosa R. BROWN. Add to Distr.: Now known from several localities in both eastern and western parts of New Guinea. Add to Ecology: Also collected in marshy places. Change highest altitude to 250 m. Lechenaultia filiformis R. BROWN. Add to Ist sentence of description: sometimes woody at base, hence appar- ently perennial, up to c. 50 cm high (NGF 38778). Line 1, replace ‘650’ by : 850. Scaevola L. Add: nom. cons. Description line 1, after ‘opposite’ add: or whorled. Distr., last line: the number of Malesian species “1° should be changed into 3. Replace the Key to the species by the following one: 1. Leaves opposite, not tufted. Scrambl- ing or climbing shrubs. Flowers yellow. 3. S. oppositifolia 1. Leaves either spirally arranged, or verticillate, usually tufted at the ends of the branches. Erect shrubs. Flowers not yellow. 2. Leaves in whorls of four. S. verticillata 2. Leaves spirally arranged.
3. Calyx-lobes usually 2!/2 mm or more. Flowers 2-21/2 cm _ long. Plant from the beach 1. S. taccada
3. Calyx-lobes 1 mm long or calyx not distinctly lobed. Flowers 1—-11/2 cm long. Mountain plants.
4. Leaves petioled, 31/2—-20 by 11/2-8 cm, flat. Flowers arranged in inflorescences, 1 cm long; calyx distinctly lobed . 2. S. micrantha
4. Leaves subsessile, 21!/2-41/2 by 1/5—-3/4 cm, with revolute margins. Flowers solitary, 11/2 cm long; ca- lyx not distinctly lobed. New Guinea (Fl. Mal. 5: 567) S. pauciflora
water ;
5: 339a Replace the name Scaevola sericea VAHL
by:
1. Scaevola taccada (GAERTN.) ROXB. Hort. Beng. (1814) 15, based upon Lobelia taccada GAERTN. Fruct. 1 (1788) 119, t. 25 f. 5. The vernacular name taccada, used as specific epithet, must be accepted as an indirect reference from ROXBURGH to GAERTNER’S name.
This is strengthened by the direct reference of both to Béla-Modagam RHEEDE, Hort. Malab. 4 (1673) 119, t. 59, as well as by the reference in Roxs. FI. Ind. 2 (1824) 146 to GAERTNER. See H. St. JOHN, Taxon 9 (1960) 200-208;
an
: 340a
: 341la : 341b
: 342a
: 342b
FosBerG, Taxon 10 (1961) 225-226. The synonym Scaevola lobelia Murr. Syst. Veg. ed. 13 (1774) 178 is illegitimate by the citation of Lobelia plumierii L. Sp. Pl. (1753) 929 as a synonym. Moreover, Scaevola plumierii as well as S. taccada are included, hence it has partly to be excluded from the Malesian flora. Line 3 from top replace ‘12-26 by 5-10’ by: 39 by 13. Add to Distr.: E. Africa (Kenya). Add to Ist paragraph: For dispersal see also G. L. Lesko & R. B. WALKER, Ecology 50 (1969) 730. According to these authors the fruits, which remain viable for a long time, float in sea-water but germinate in fresh water only, that means after having been drifted ashore and after rain. Scaevola micrantha PRESL. Add to Ist sentence of description: or treelet to 10 m by 5 cm. Add to Distr. after Borneo: Mt Meliau, Mt Tavau. Add at the end of Distr.: Cited from Botel Tobago (near Taiwan) by HatusimA, Mem. Fac. Agr. Kagosh. Un. 7 (1970) 327. After Scaevola micrantha PResL add:
Scaevola verticillata LEENH. Blumea 12 (1964) 317, f. 1. — Fig. 10.
Shrub. Branchlets + terete, woolly tomentose; leaf-axils provided with a tuft of c. 11/2 cm long, isabelline, sericeous hairs, in older leaves forming a kind of papyraceous ligule. Leaves whorled in fours, + tufted at the end of the branches; petiole c. 3/4-1 cm long, grooved above, densely woolly; blade obovate-oblong, 5!/2-7 by 21!/2-3 cm, chartaceous, apparently convex with recurved margins, fairly densely shortly tomentose above, glabrescent, densely woolly tomentose beneath; base cuneate, slightly attenuate; margin minutely serrate towards the apex; apex acute; midrib slightly sunken above, prominent beneath; nerves c. 10-12 pairs, promi- nulous above, invisible beneath. /nflores- cences condensed, few-flowered, peduncle short; bracts relatively large. Flowers 14 mm long. Calyx lobes different, dorsal one narrowly deltoid, acute, 2!/2 mm, the others ovate, rounded, ventral one 1 mm, laterals 3/4 mm, all outside (as well as the ovary) fairly densely, inside sparsely woolly. Corolla tube inside densely woolly, more sparsely so in the basal 1/4 of the lobes, outdise densely woolly-tomentose but for the basal half of the tube; membranous margins of the lobes broad and crispy in the upper half, in the lower half narrowed and with some long and stiff marginal
952
FLORA
MALESIANA
[ser. I, vol. 68
Fig. 10. Scaevola verticillata LEENH. a. Flower, 4, b. calyx, <4 (note the three different kinds of lobes!), c. & d. anther from in- and outside resp., < 8, e. indusium, = 8 (showing the spreading hairs at its base) (a-e W. MEWER SAN 28818).
5: 3426
5: 344a
5: 369
35) 370
hairs and 2 or 3 lengthened teeth crowned with a brush-like bundle of hairs. Style with a few woolly hairs in the basal half and a collar of long, stiff, spreading hairs just under the indusium. Fruits unknown.
Distr. Malesia: Borneo (Mt Tambuyo- kon near Mt Kinabalu).
Ecol. Subalpine vegetation, on ser- pentine, 2500 m. Fi. July.
Note. Nearest allied to S. micrantha and S. pauciflora. Especially charac- terized by the whorled leaves and the heterosepalous calyx.
Scaevola oppositifolia R. BROWN.
Line 21 from bottom, add after ‘acute’: or sometimes rounded, truncate, or even subcordate.
Line 7 from bottom, add after ‘Style glabrous’: to sparsely pilose.
Add to Distr.: Ferguson I., Normanby I., and Goodenough I.
Hamamelidaceae
Distylium Step. & Zucc.
The genus Distylium was also recorded from Central America with 2-3 spp. Recently ENpREss (Bot. Jahrb. 89, 1969, 355) has referred these American species to a separate genus Molinadendron ENpRESS, which he even says stands very remote and does not belong in the same tribe. The arguments for this seem to me, however, to be of more trivial nature and I do not agree with their inflation to generic rank.
Sycopsis OLIVER.
Recently ENpREss (Bot. Jahrb. 90, 1970, 30) has from the genus Sycopsis split off
323710
55/381
5: 393a
5: 402a
4 Sino-Malesian spp. to represent a separate genus Distyliopsis ENDRESS, which the author maintains close to Sycopsis. To this also belongs the single Malesian species S. dunnii HEMSL. The arguments on which this distinction is based are partly due to a theoretical explanation of inflorescences, partly certainly not of generic rank (degree of connation of sepals) and partly of no value whatsoever (almost replacing ranges). I am not prepared to accept this as a distinct genus.
Sycopsis dunnii HEMSL.
Add to Distr.: Now also found in North Borneo (Mt Kinabalu, Mesilau R., RSNB 7022) (add to fig. 5).
Hydrocaryaceae, see Trapaceae Hydrocharitaceae (DEN HARTOG)
A monograph has appeared by DEN HartoG, The Seagrasses of the World, Verh. Kon. Ned. Akad. Wet. A’dam, afd. Natuurk. ser. 2, 591 (1970) 1-275, 63 fig., 31 pl., and furthermore world maps of distribution of seagrasses in Pacific Plant Areas 2 (Blumea, Suppl. 5) (1966) 208-219. Both contain a wealth of new data of various kind.
Blyxa novoguineensis HARTOG.
Add to Distr.: It has probably also been found in the Philippines (Luzon) by Jacoss, but unfortunately the specimen is sterile.
The authority of Enhalus acoroides is: (L. f.) Royre Ill. (1840) 453. This publication antedates STEUDEL’s Nom.
September 1972]
5: 408 5: 410a
5: 4lla
5: 412a
6: 153a
4: 211b
4°) 215b
Bot. ed. 2, 1 (1840) 554 by a few months. In Key, line 7, replace ‘2. H. minor’ by: 2. H. ovata.
The correct name of 2. Halophila minor (ZoLL.) HartoG is: 2. Halophila ovata GAupD. in Freyc. Voy. Bot. (1827) t. 40, f. 1. In the latter published text of the same work, /.c. 430, GAUDICHAUD added as a synonym Caulinia ovalis R.BR., which would make his own name illegit- imate; but his earlier published plate is validly published. Cf. HARTOG (1970) 251. Halophila decipiens var. pubescens HAR- TOG is no longer recognized. Cf. HARTOG (1970) 254.
Halophila beccarii ASCHERS.
Add to Distr.: Found abundantly in the Malay Peninsula. Cf. HARTOG (1970) 262.
Juglandaceae
Engelhardia spicata LECHEN. ex BL. Replace the name var. colebrookeana (LINDL. ex WALL.) O.K. by:
var. integra (KURZ) MANNING, comb. nov. — E. villosa Kurz var. integra KURZ, For. Fl. Burma 2 (1877) 492.
Juncaceae
Juncus effusus L.
Add to Distr.: Malay Peninsula (Pahang, Cameron Highlands, G. Batu Brinchang, 2 clumps along roadside, c. 1 mile from summit, c. 2000 m, J. SINCLAIR 9956 (SING), Nov. 4, 1958).
Widely distributed in Malesia. In the Malayan specimen the inflorescence is up to 18 cm, due to elongated branchings. The fact that this species was found so late in Malaya and along a roadside in the newly opened Cameron Highlands is no proof at all that it is not native. In these forested mountains Juncus could original- ly equally well have been very scarce because of its heliophilous ecology, but extended after man artificially opened the forest. This happened also in Came- ron Highlands with Juncus prismatocar- pus R.Br., first collected in 1958. Cf KeRN, Gard. Bull. Sing. 17 (1958) 91-92. The like happens frequently; for example on Mt Diéng, in Java, Gentiana quadrifaria is a common weed on paths, though doubtless native and_ scarce before this mountain was deforested. Add to Excluded:
Luzula silvatica (HuDSs.) GAUDIN, Agrost. Helvet. 2 (1811) 240; Bucn. Pfl. R. Heft 25 (1906) 55. — Juncodes silvaticum (Hups.) O.K. Rev. Gen. PI. 2 (1891) 725.
This was recorded by O. KUNTZE from Central Java: Diéng Mt (O. KUNTZE 5715b, Bucn. /.c.). Though I have not seen the specimen there is presumably not
Addenda, corrigenda et emendanda
6: 296
953
the slightest doubt about BUCHENAU’s identification. Dr. BAssetTr MAGUIRE wrote me (26-2-60) that is it not in the New York Bot. Gard. Herbarium with KUNTZE’s collection. The species is not known to occur outside Europe. It is also strange that it is a b-number, which might suggest that it was mixed with something else. Could it have happened that KUNTZE brought along old drying paper which he had used in Europe and in this way transported a European herbarium specimen to Java? This has occurred before, and by miracle also with a plant from Mt Diéng, viz Luronium natans (L.) RAFIN., cf. Fl. Mal. I, 5 (1957) 3346. This mislocation could be proved by means of using diatoms adher- ing to the plant as tracers (cf. STEEN. Taxon 5, 1956, 157-158) by VAN DER WeRrFFE (Blumea 7, 1954, 599-601). If we had KUNTZE’s specimen we _ could probably apply this method with success. Another solution to the mystification could be that a European collection got mixed with Javanese collections after KuNTZE had returned to Europe; a similar thing happened with RIDLEyY’s Javanese collections which contain a few European Carices.
It is said that L. silvatica is sometimes dispersed outside its natural habitat with grass-seed in Holland, but this cannot have happened in KUNTZE’s time to Java. It certainly does and did not occur in Java.
Loganiaceae (LEENHOUTS)
Add to Phytochemistry: Cf. HEGNAUER, Chemotaxonomie 3 (1964) 307-310, 647. Insert before Delimitation and subdivi- sion:
Cytology. Cf. GaADELLA, Act. Neerl. 11 (1962) 51-S5. Palynology. Cf. PUNT & LEENHOUTS, Grana Palynologica 7 (1967) 469-516. Add to the discussion on the position of the Buddlejeae: W. TROLL, Jahrb. 1965 Akad. Wissensch. Lit. Mainz (1966) 128, on arguments derived from the morphol- ogy of the inflorescence, also concludes to a closer relationship with the Loga- niaceae.
Add to (4) Peltanthera: Revision: LEEUWENBERG, Act. Bot. Neerl. 16 (1967) 143-146.
Add to (8) Gomphostigma: Revision: VERDOORN, FI. S. Afr. 26 (1963) 168-171. Add to (10) Emorya: See: NORMAN &
Bot.
Moore, Southwestern Naturalist 13 (1968) 137-142. Add to (11) Adenoplusia and (12)
Adenoplea: Both reduced to Buddleja by LEEUWENBERG, Act. Bot. Neer]. 16 (1967) 143.
954
FLORA MALESIANA
(ser. I, vol. 6®
6:5 297
6: 300
6: 302
6: 303a
6: 304b
6: 308a
6: 3085
6: 3lla
6: 315a
6: 3166
6: 317b
6: 318a 6: 320a
Add to (13) Bonyunia: 4 or 5 spp. 6: 320b Fagraea ridleyi K. & G.
Revision: LEEUWENBERG, Act. Bot. Neer]. 18 (1969) 152-158.
Add to (16) Usteria: Revision: LEEUWEN- BERG, Act. Bot. Neerl. 12 (1963) 112-118. Insert before E. Strychneae: D’. Ploco- spermeae.
(18a) Plocosperma. Monotypic; Central America. Revision: LEEUWENBERG, Act. Bot. Neerl. 16 (1967) 56-61.
Add to Desfontainea: Revised and placed in a separate tribe Desfontaineae by LEEUWENBERG, Act. Bot. Neer]. 18 (1969) 669-679.
Add to Retzia: Revised and placed in a separate tribe Retzieae by LEEUWENBERG, Act. Bot. Neer]. 13 (1964) 333-339. Add to Dispersal, 1stjsentence, after ‘bats’: and by Viverridae (‘luwaks’), cf. DOCT. v. LEEUWEN, Trop. Natuur 21 (1932) 142. Add to Anat.: See SminGH, Gard. Bull. Sing. 22 (1967) 193-212, for sclereids; BurGEss, Timbers of Sabah (1966) 385- 388, for wood-anatomy.
Add to 22, 2nd lead: (Confusion is possible with specimens of F. auriculata ssp. borneensis with very faint auricles and 1-3-flowered inflorescences.) Fagraea elliptica Roxs.
Add to literature: BACK. & BAKH. f. Fl. Java 2 (1965) 211.
Fagraea fragrans Rox.
Add to literature: Troup, Silvic. Ind. Trees 2 (1921) 676; MITCHELL, Mal. For. 27 (1964) 127; BAck. & BAKH. /. FI. Java 2 (1965) 211; SmytTHiEs, Common Sarawak Trees (1965) 87; KENG, Ma- layan Seed Plants (1969) f. 140-141. Add to Distr. 2nd paragraph: MIQUEL’s citation from the Moluccas is apparently based upon De Fretes 5742 from Ambon U)
Insert under Uses before ‘A decoction. .’: see SCHNEIDER, Bull. Bur. For. Philip. 14 (1916) 202, f. 71.
Fagraea racemosa JACK ex WALL.
Add to literature: ANDERSON, Gard. Bull. Sing. 20 (1963) 192; Back. & BAKH. f. Fl. Java 2 (1965) 211.
Fagraea ceilanica THUNB.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 211.
Add (sub F. litoralis Bu.): Doct. v. LEEUWEN, Trop. Natuur 21 (1932) 142; ANDERSON, Gard. Bull. Sing. 20 (1963) 192.
Add to Distr.: Bougainville (NGF 31290, ‘ternatana’), Solomon Is. (New Georgia: WHITMORE R.S.S. 6361, ‘ternatana’; Guadalcanal: R.S.S. 86, ‘/itoralis’, R.S.S. 98, ‘ternatana’).
3rd paragraph, ‘F. oblonga’, distribution: Borneo (Sarawak). Fagraea acuminatissima MERR. Add to Ecol.: also on coastal rocks.
add to
6: 323a
6: 324a
6: 327b
6: 328a 6: 328b
6: 332b
6: 333a 6: 335a
6: 335b
6: 336
6: 337b
6: 340a
6: 343
6: 345 6: 346
Add to description, Ist sentence: treelet (5 m).
Add to Ecol. 3rd line, after ‘sandstone’: and limestone.
Fagraea blumei G. Don.
Add to literature: BAck. & BAKH. /. Fl. Java 2 (1965) 211.
ssp. blumei.
Add to Ecol., after altitude: Kinabalu up to 3000 m. Fagraea fastigiata BL.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 211.
Fagraea auriculata JACK.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 211.
Add to Distr.: Sumbawa, Flores.
ssp. auriculata.
Add to Distr.: Sumbawa, Flores.
ssp. borneensis (SCHEFF.) LEENH.
Read Inflorescences: (1—)3-7— flowered. Fagraea gracilipes A. GRAY.
Add to literature: PARHAM, Pl. Fiji Is. (1964) 176, f. 64.
Fagraea gracilipes A. GRAY.
Add to Distr. sub Solomon Is.: Choiseul. Fagraea berteriana A. GRAY ex BTH. Add to literature (sub F. schlechteri GILG & BENED.): GUILL., THORNE & VIROT, Un. Iowa Stud. Nat. Hist. 20, 7 (1965) f. 15 (p. 55).
Ecol.: Change highest altitude into 1700 m. Add: Flowers open in the morning, drop off in mid-afternoon.
Buddleja Houst. ex LINNE.
Replace ‘Spelled Buddleia by most au- thors’ by: Incorrectly spelled Buddleia by most authors; cf. Int. Code of Bot. Nomencl. (1961) Appendix II, Nomina Familiarum conservanda, sub Buddle- jJaceae.
Add to synonymy: Toxina Norona, Verh. Bat. Gen. 5 (1791) 4, nom. nud. Buddleja asiatica Lour.
Add to literature: BAcK. & BAKH. f,. Fl. Java 2 (1965) 212.
Buddleja davidii FRANCH.
Add to literature: BACK. & BAKH. f. Fl. Java 2 (1965) 212.
Gelsemium JuSSIEU.
Add to literature: DUNCAN & DE JONG, Sida 1 (1964) 346-357; ORNDuFF, J. Arn. Arb. 51 (1970) 1-17.
Add to description: Seeds not winged in G. rankinii.
Strychnos L.
Add to literature: KRUKOFF, Mem. N.Y. Bot. Gard. 12, 2 (1965) 1-94; LEEUWEN- BERG, Med. Landb. Hogesch. Wagenin- gen 69 (1969) 1-316.
Add to Distr.: Cf. VAN BALGooy, Blumea Suppl. 5 (1966) 256, map 141. Add to Taxonomy: A new subdivision has been proposed by LEEUWENBERG, /.c.,
or
on Mt
September 1972]
6: 347
Addenda, corrigenda et emendanda 955
who distinguishes between 12 sections.
As far as the Malesian species are con-
cerned, these are the following:
Sect. Strychnos: S. angustiflora, S.
ignatii, S. kerrii, S. lucida, and S. nux-
vomica.
Sect. Penicillatae: S. axillaris, S. ridleyi,
and possibly S. me/lanocarpa.
Sect. Brevitubae: S. flavescens, S.
luzonensis, and S. vanprukii.
Sect. Lanigerae: S. borneensis, S. curtisit,
S. lanata, S. lanceolaris, S. ledermannii,
S. maingayi, S. minor, S. oleifolia, S.
ovata, S. polytrichantha, S._ thorelii,
S. villosa, and possibly S. rufa.
Key to the species:
Replace couplet 3, as follows:
3. Petiole 2-4 mm. Inflorescences few- flowered. Style glabrous. Fruits 2—21/2 cm @, pedicels only slightly thickened.
3. S. lucida
3. Petiole 4-11 mm. Inflorescences many- flowered. Style thin-woolly. Fruits 3—6 cm @, pedicels thickened.
3A. Leaves elliptic-oblong, 2—3 times as long as wide, shining green when dry. Calyx glabrous; corolla 14-15 mm long, inside woolly at the mouth; anthers 1 mm. Fruit thick- walled. Paired tendrils frequent (always a liana) 23. S. kerrii 3A. Leaves broad-ovate to elliptic, c. 11/4-11/2 times as long as wide, dull green when dry. Calyx outside pubescent; corolla 10-12 mm long, inside woolly in the lower half of the tube; anthers 13/4 mm. Fruit thin-walled. No tendrils (mostly a tree) . . . . 2. S.nux-vomica Insert between 6 and 7: 6A. Twigs sharply quadrangular, branch- es often rounded quandrangular. Dried leaves beneath mostly yellow- ish brown to copper-red. Anthers glabrous 12. S. vanprukii
6A. Twigs and branches terete. Dried leaves beneath either dark brown or greenish. Anthers hairy.
Add to 8, 2nd lead, at end of Ist sentence:
(exceptionally glabrous).
Replace couplet 12, as follows:
12. Corolla 9-10 mm long.
12A. Leaves elliptic, c. 21/2 times as long as wide, base cuneate, apex mostly distinctly acuminate. Calyx outside glabrous, sepals rounded
to nearly truncate; corolla inside densely woolly from slightly above
the base to halfway the lobes; pistil fairly densely hairy from about halfway the ovary upwards.
13. S. borneensis
12A. Leaves mostly ovate to broad- ovate, 1-2 times as long as wide,
base (obtuse to) rounded to
cordate, apex not to. shortly acuminate. Calyx outside puber- ulous, sepals acute; corolla inside woolly in the mouth and on the lower part of the lobes; pistil glabrous . 22. S. angustiflora 12. Corolla up to 61/2 mm long. Replace in 14, Ist lead, ‘S. colubrina’ by: S. minor. Add to 14, 2nd lead, after ‘Anthers subsessile’: or filaments short.
15, 2nd lead, 2nd sentence, read: Leaves 3—5—plinerved. Insert after the key to the species the following: With the key as published originally in the Flora Malesiana, only flowering material could be identified. Four species the flowers of which were known too incompletely or not at all (S. melanocarpa, S. quandrangularis, S. rufa, and S. thorelii) could then not be included. Out of these four, one (S. quadrangularis, now S. vanprukii) could be added in the above Addenda to the key as in the mean time the flowers became known. But fruiting and sterile material, more than half the collections coming in, could still not be named. For that reason I have compiled the following synoptic key to all Malesian species, making use of all main characters. This is a multiple entry key, which means that one can start with every character, and can make use of all characters available in the material. The numbers after each lead represent the species showing that character state: species in which that character is still unknown are included with a question mark. The best way to work with this key is first to try whether the specimen to be identified shows one or more ‘rare’ characters, to note down all numbers of species that may show that character (including those with a question mark), and, with each following character, to narrow down this series of possibilities. Finally, this will lead to one or a few species; comparison with the descrip- tions and with material will mostly be conclusive. If identification in this way does not lead to any species, you may try what species shows the smallest number of differences: it may be that your specimen represents a species not included in the key, but it is also possible that it shows a character state not yet known from one of the species. The numbers of the species are in accordance with the following enumeration.
1. Twigs
aliglabrousAds 3a4+54) oe okON EI.
122 1S 14S ANG ANSA22° 235-24.
b. hairy (mostly slightly and early glabrescent): 2. 3. 5. 6. 14. 16. 17. 18.
956
FLORA MALESIANA
[ser. I, vol. 68
OS
co)
LOX 2OR215 22% Thorns or spines presents 24.35 lsa22.
. absent: all species.
Leaves, shape (irrespective of place of greatest width) rhomboid: 18.
. suborbicular (about as long as wide):
DeSe WE Sy22:
elliptic (+ 11/2 times as long as Wide) cele sn4. Sami ecrelile toes: 14. 16. 18. 20. 22. 23. 24.
oblong (+ 21/2 times as long as Wide) aula TOs ss OMAOM 1213: [AS Gi Sen ORO 23 245 lanceolate (3 or more times as long as wide): 1. 4. 8.9. 11. 12. 15. 16. 18. Di 2D
Leaves, place of greatest width about the middle: 1. 2. 3. 4.5. 6.7.8. LOMAS Ae Sel Ga ison 20! Mle pss 38).
distinctly below the middle: 1. 2. 3. SHON OO MOA Gy 17. 18! 19: 2ORAII22F 23824"
distinctly above the middle: 11. Length of leaves
Up toMlOicnil 2s S44 onGs 08. 110: eee al Saati Saal Oates 9) 20) De 22 23924)
1O=Sveme le 2 Ay SaGass 9) 10) 11: P25 AG eS20221. 22823) 24: 15-20 cm: 1. 6. 10. 11. 12. 18. 20. more than 20 cm: 1. 11.
Width of leaves
up to 5 cm: all species. S—OlemiPile2esn4s 52608. 10s 12: 1ANIGI1 8520222573424"
more than 10 cm: 1. 2. 11. 12. Leaves
glabrous: 1. 2. 3. 4. 5. 6. 7. 8. 9. 10. LD AS 4a Sreliowlica ol 22. 24.
. not fully glabrous: 6. 17. 18. 19. 20. Dit,
Lower side of leaf
warty(-papillose): 3. 11. 18. 20. SMOODNMl AS Ones oO lO ndiie Ll SAS 6s lalSealOs 21) 22) 2504)
Leaf base
anpilarswle 234445: 6s. 9, LON 11 17.3: 14 AS Ges MOND 14 22) 23. 24.
rounded: tine as a4 aS NOne SH lO iie 2A aGs 1 Sel OMORe 1 22" 23) 24.
(sub)cordate: 2. 3. 6. 11. 18. 20. 21. aby
Leaf apex
acuminate: 1.2.4.5. 6.7.8.9. 10. 11. IDRIS 14 iSaOmiia dss 19420) 2i- DO OBO A
not acuminate: 1. 2. 3. 5. 10. 11. 12. GPUS 922923:
11.
Leaf apex or acumen
ay acute: 14223574: 5: GalOmiieae rs:
14. . terminal (either on a main branch
LAS TSS Lor 17s l8 19 A2ORele 22503" 24.
blunt to rounded: 1. 2. 3. 5. 7. 8. 9. LOM 22.
emarginate: 3. 18.
. Number of main nerves
32.1..2. 324.5 -9R1E eS eS. 16.17.18. 19:20) 219228235245 52:3. 526.758) 9 gM es. 16:1718:.20. 21.2258:
Te Mdia:
. Petiole, length
up tor4 mmia3, Sale 2 Se Ges: 20. 24.
4-71/2 mm: 1. 2. 4. 5.6. 7. 8. 9. 10. 10512513) 14 1S AG ae Oe oe 223i
more than 71/2 mm: 1. 2. 5. 6. 9. 10. 11512516; 18: 19921 eas Inflorescences
or on a short axillary shoot with at least one pair of leaves; sometimes with a pair of basal branches in the upper leaf axils): 1. 2. 3.4. 5. 6.7. 8. Salil, 14: 167; ls2iee2 28h eAs
. axillary, sometimes ramiflorous or
together pseudoterminal: 1. 4. 5. 6. 78.10) 115 12: ABA IS GateealS: 20. 21. 24.
Inflorescences, length
up tos cmitle 2345557 cement. NUSy5 MIC a IMR iss, TIS P10), Ail P22. 228), 24.
5=10' cms 1.54.75 2168 8) SOM 2. 1B), Moy 9/5 Il, 728),
more than 10 cm: 6. 11.
. Calyx outside
hairy? 1. 2: 35:4: SaGas,Oniiealeelio: 16M SP20R21 R22
glabrous: 4. 6. 7. 10. 11. 12. 13. 14. 18. 19. 21?. 23. 24.
Calyx inside
hairy 74: 6:12. 139195 202222 glabrous: 1. 2. 3. 4. 5. 7. 8.9. 10. 11. 12) 13.9145 IS oy se ZO ree) Dye), OEE
Sepals
acutes 12 2532556: 102 aleater 18519920?) 217522824"
. blunt, rounded, or truncate: 1. 2. 4.
SH6D 78h 9 AO ey 12S ealeree eels 18.202:212. 23:24,
Corolla, total length
up to 5mm: 4. 5. 6. 11. 12. 14. 15. 16. 17. 18. 19. 20. 21?. 24.
2 S=10imms 7. 899. LOPE Aeie.
22»
10 mm or more; 1. 2. 3. 21?. 22. 23. Corolla tube
longer than limb: 1. 2. 3. 18. 20?. 217. wey
. equal to limb: 11. 13. 14. 15. 16. 18.
September 1972]
Addenda, corrigenda et emendanda
o57
one
22.
19; 200; 217.122: 24:
. shorter than limb: 4. 5. 6. 7. 8. 9. 10.
27. 2025217.
. Mature corolla outside
hairy: 5. 6. 9. 15. 16, 18. 20?. 217. fully glabrous: 1. 2. 3. 4. 5. 7. 8. 10. ie 2a 47 182195 2009 20?. 22. 23. 24.
Corolla inside
glabrous: 3. 16. 20?. 21?.
with a whorl of bristle-like hairs in the mouth or up to halfway on the lobes: 17. 18. 19. 20?. 21?. 24?. woolly in the tube, sometimes ex- tending to the lobes: 1. 2. 3. 6. 8. 9. TOM MR1S 32.07.21):
only woolly about the mouth: 4. 5. 214s 116.202, 219222. 235247: woolly on the lobes, sometimes from the mouth upwards: 7. 11. 12. 15. 16. 19. 20?. 217. 247.
. Insertion of stamens
inthe mouth: 1)..2) 3.4. 5s 6. 7.8. 9: LOM IOS 1AM Selo 202752 7. 22N23)
in the tube: 12. 14. 18. 19. 20?. 21?. 2223. 24:
Filament, length
up to 1/2 mm: 1. 2. 3. 5.6. 11. 14. 15. 16. 17. 18. 19. 20?. 21?. 23. 24. ol foomment. sa SnonlOwli. Wa lst LSS U8s 202. 217:
11/2-21/2 mm: 4. 5. 7. 9. 12. 13. 20?. PAE
21/2 mm or more: 4. 8. 12. 20?. 217. DD:
. Anther
OVvater 4) SO Ul 12s US16. Wie US: 195202, 21. 24:
elliptic? 25 5, 9F 1 1213145207: DIED 3:
oblong: 1 37457510! MA12? 132207: P22 623:
lanceolate: 4. 7. 8. 10. 11. 13. 207. 21?.
. Anther, length
up to 1 mm: 4. 5. 6. 7. 11. 12. 14. 15. GHG UseOe 2075217. 23" 24: more than 1 mm: 1. 2. 3. 5. 8. 9. 10. MINIS 202 217h 22) 23%
. Anther
hairy: 4. 5. 6.8. 10. 11. 13.15. 16. 17. 18. 19. 20?. 217. 24.
elabrous:le2:3. 457. cool 2 13: 1451162202221. 22.23.
. Pistil, length
up to 4mm: 4. 5.6. 11. 12. 14. 15. 16. 17. 18. 19. 20?. 21?. 24. more than 4 mm: 1. 2. 3. 7. 8. 9. 10. AIS 2.024219 22523"
. Ovary
Hain yA KOM erOwo Ons 135 15. 17. 20?. 21?. 24.
Slabrouse le 20344. SaGai2. 14516. 18. 19. 20?. 212. 22.§23.
: 347a
: 349b
: 350a
: 350b >) SI
2 shlsye
30. Style
ay hairy 2 AS NGe Tash. 1113: (4. 15206217 72075:217, 23. 247.
b: @elabrouss1) 3943'5.6)-11..12. 16. 18. 19. 20?. 21?. 22. 247.
31. Pedicels in fruit
a. much thickened: 1. 2. 7?. 9?. 107. 13. 14?. 15. 17?. 19?. 20. 23. 247.
b. only slightly thickened: 3. 4. 5. 6. 7?. SiO? sOP aie 2er4? Gh 722 18: 19?. 21. 22. 24?.
32. Fruits
a: Vglobulan?1) 253545556777; 897. 107. PQS: 4 eS Pom? Lee 9 2: 20. 22. 23. 24.
b. ellipsoid or ovoid: 1. 5. 6. 7?. 9?. 10?. LAP AUS Pa aS 2
33. Fruits, diameter or length
a.) upito 24/5 emir 3h4: 5165772977; 102. IE 2S 1421S oaks. 197. Pile Q2ADA
b. more than 21/2 cm: 1. 2. 7?. 8. 97. ODA SP ate 2a 20) 212. 23:
34. Fruits
a. thick-walled: 1. 7?. 9?. 10?. 14?. 15?. 172 ASP 23H 24?
b. thin-walled: 2. 3. 4. 5. 6. 7?. 8. 92. LO? UID. 1S 4 ispalGs 1772 1S: 19?. 20. 21. 22. 24?.
35. Number of seeds
aot 16) 4S SG ia oe el Orentem 2. 13% L416 Ba99821. 225 247
bs more thani219243562 775 8. 9% 107: LTS PAA LS Oi Oe 2022 223: 24?.
1. Strychnos ignatii BERG.
Add to literature: BACK. & BAKH. /. FI. Java 2 (1965) 210; TrrEL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 59, t. 4 f. 6-8.
Change S. philippinensis BLco into S. philippensis BLco.
2. Strychnos nux-vomica L.
Add to literature: Troup, Silvic. Ind. Trees 2 (1921) 673; PETELoT, PI. Médic. Cambodge, Laos & Viet-Nam 2 (1953) 169; BAcK. & BAKH. f. Fl. Java 2 (1965) 210; TirEL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 71.
Add to Distr.: Burma.
3. Strychnos lucida R.Br.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 210.
4. Strychnos maingayi CLARKE.
5. Strychnos ovata HILL.
Add to literature: TrREL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 55, tshiail=se
Add to synonymy: Strychnos sp. MERR. Pl. Elm. Born. (1929) 252.
Add to description: Young twigs some- times sparsely minutely hairy. Leaves up to 11 cm long.
Insert after the paragraph on Ecology:
958 FLORA MALESIANA [ser. I, vol. 66 Uses. In Sarawak, Ukit nomads pre- Distr. S. and NE. India, Thailand, pare one of the elements of their dart poi- Laos, Vietnam, and Malesia: Malay son from this species. Peninsula (Perak, Selangor), Borneo
6: 3515 6. Strychnos villosa HILL. (Sarawak, Central E. Borneo).
Add to literature: BACK. & BAKH. f. FI. Ecol. Dense jungle, from the lowland Java 2 (1965) 209. to above 1000 m. F/. Aug., Nov. fr. Sept.
6: 353a 7. Strychnos curtisii K. & G. Uses. The bark of the roots is used for 8. Strychnos polytrichantha GILG. arrow-poison. See BURKILL, Dict. (1935)
6: 353b 9. Strychnos oleifolia HILL. 2099.
6: 355a 10. Strychnos lanata HILL. Vern. Akar ipoh, Mal. Pen.
The correct name for species I1 is: 13. Strychnos borneensis LEENH.
11. Strychnos minor DeNNst. Schluess. 6: 357a 14. Strychnos ledermannii GILG & BENED. Hort. Malab. (1818) 33; Busser & Add to description: Leaves sometimes PuiLtcox, Taxon 20 (1971) 537-543. — S. with 3 main nerves.
colubrina Auct. non L.: Hitt, Kew Bull. 6: 3575 15. Strychnos lanceolaris Miaq.
(1917) 157 et seg.; LEENH. FI. Mal. I, 6 16. Strychnos flavescens K. & G.
(1962) 355; BAcK. & BAKH. f. FI. Java 2 Add to description: Leaves in a specimen (1965) 209; TireEL-RoubeT, Logan. Cam- from Borneo (Sarawak For. Dept. bodge, Laos & Vietnam (1970) 50, t. 13460) 5-plinerved, the outer nerves INE Sh. diverging at, the inner distinctly above Add to description: Leaves sometimes the base. Corolla of the same specimen not acuminate at apex; lower side mostly woolly inside all over the inner surface of smooth to rarely minutely papillose. the lobes; stamens with short filaments.
6: 356a Add to Distr.: Lesser Sunda Is. (Flores). 6: 358a Add to Distr.: Borneo (Sarawak, Sabah).
6: 356b Replace species 12 as follows: Add to Ecol.: Altitude up to 1350 m. F7/. 12. Strychnos vanprukii Crais, Kew Bull. from March onwards, fr. Febr., Sept. (1911) 421; Hitt, Kew Bull. (1917) 139; 17. Strychnos luzonensis ELM. in Craib, Fl. Siam. En. 3 (1951) 62; 18. Strychnos axillaris COLEBR. TiREL-RoubeET, Logan. Cambodge, Laos Add to literature: BACK. & BAKH. f. FI. & Vietnam (1970) 48, t. 2 f. 1-7. — S. Java 2 (1965) 209; TireEL-RoubeT, Logan. aenea Hitt, Kew Bull. (1917) 138, cum Cambodge, Laos & Vietnam (1970) 37, fig., incl. also var. acuminata. — S. t. 12 f. 1-3. quadrangularis Hitt, Kew Bull. (1917) 6: 359a Add to Distr.: Lesser Sunda Is. (Flores: 205; LEENH. FI. Mal. I, 6 (1962) 356. — S. KOSTERMANS 22156). maingayi CLARKE ssp. borneensis LEENH. 6: 3605 19. Strychnos ridleyi K. & G.
Blumea 14 (1966) 230. 6: 36la 20. Strychnos rufa CLARKE.
Climbing shrub or liana, provided with 21. Strychnos thorelii PIERRE ex Dop. double tendrils. Twigs slender, mostly Add to literature: TrREL-RoubDET, Logan. sharply quandrangular, glabrous; branch- Cambodge, Laos & Vietnam (1970) 82, es rounded quadrangular. Leaves 71+/2- iia UPA I Bh.
20 by 21/2-11 cm, ratio c. 2-3, widest 6: 361b 22. Strychnos angustiflora Bru. J. Proc.
about or sometimes below the middle, thin-chartaceous to papyraceous, be- neath copper-red to yellowish brown or sometimes green when dried, smooth and glabrous; base cuneate to rounded; apex short- to caudate-acuminate, acute; strongly 3—S-plinerved; petiole 4-5 mm. Inflorescences axillary, 11/2-6 cm long, lax, few- to many-flowered, puberulous or glabrous. Flowers 5-merous. Sepals c. 1 mm, acute or blunt, outside glabrous, inside glabrous or nearly so. Corolla 3-4 mm long, the tube 1/4-11/4 mm, outside glabrous, inside woolly about the mouth up to halfway the lobes and papillose hairy at the tips. Stamens inserted at or slightly below the mouth, filament 1-3 mm long, anther ovate, elliptic, or oblong, 1/5-3/,4 mm long, glabrous. Pistil 3/4— 23/4 mm long, glabrous. Pedicels in fruit hardly thickened. Fruits globular, c. 2 cm @, thin-walled, said to be white. Seeds 1, semiglobular, c. 1 em @ and 8 mm thick, glabrous.
Linn. Soc. Bot. 1 (1856) 102; Fl. Hong- kong (1861) 232; Hit, Kew Bull. (1917) 182 cum fig.; MERR. Lingn. Sc. J. 5 (1927) 148; Herktots, Hongkong Natur. 4 (1934) 108, f. 3; TrreL-RoubDeET, Logan. Cambodge, Laos & Vietnam (1970) 62, t. 5 f. 1-5. — §. nux-vomica Auct. non L.: LEENH. FI. Mal. I, 6 (1962) 349 pro specim. Philipp.
Liana, provided with simple tendrils and spines. Twigs puberulous, early glabrescent. Leaves 3-12 by 11/2-7 cm, ratio 1-2, widest below, sometimes about the middle, chartaceous to thin-coria- ceous, smooth and glabrous; base rounded to cordate, exceptionally obtuse; apex not or tapering short-acuminate, acute to blunt, mucronate; 3—5-plinerved; petiole 4-10 mm. Inflorescences terminal on short axillary shoots, rather lax and fairly many-flowered, 2-4 cm_ long. Flowers 5-merous. Calyx outside puber- ulous, inside glabrous, sepals acute, 1—-11/» mm long. Corolla c. 1 cm long, the tube
September 1972]
Addenda, corrigenda et emendanda
about as long as the lobes, outside papillose but not hairy, inside woolly in the mouth and on the lower part of the lobes. Stamens inserted about the mouth, filament 3-4 mm long, anther oblong, 13/4-2 mm long, glabrous, blunt. Pistil c. 1 cm long, glabrous. Pedicels in fruit only slightly thickened. Fruits globular, 2-4cm @®, thin-walled. Seeds 1 or 2 (rarely more), disk-shaped, c. 11/2-1?/4 cm @, sericeous.
Distr. China (Kwantung), Hainan, Hong-Kong, Thailand, S. Vietnam, and Malesia: Philippines (Mindoro, Oriental Prov., Puerto Galera Bay, once collected).
Ecol. Collected on the shore above tide level. Fr. April.
Note. S. angustiflora is rather closely al- lied to S. nux-vomica and, whereas the dif- ferences in the flowers are clear, vegetativ- ely and in fruit they look much alike. Asa whole the leaves are broader, more ovate, and at base deeper cordate than in S. nux- vomica, but less so than in the also closely allied S. nux-blanda. It was Dr. N. G. Bisset, London, who, on account of the alkaloids found in the seeds, first expressed his doubt as to the identifica- tion of the Philippine specimen.
23. Strychnos kerrii HitL, Kew Bull. (1925) 426; in Craib, Fl. Siam. En. 3 (1951) 58; TrrEL-ROoUDET, Logan. Cam- bodge, Laos & Vietnam (1970) 78, t. 5 f. 6-9.
Liana, provided with double tendrils. Twigs glabrous, sparsely lenticellate. Leaves 7!/2-14 by 4-5 cm, ratio c. 2-3, widest about or sometimes below the middle, papyraceous to chartaceous, smooth and glabrous; base cuneate to rounded; apex acute to abruptly acute- acuminate; 3 (rarely 5) main nerves; petiole 5-10 mm. /nflorescences terminal, dense and many-flowered, 4-6 cm long. Flowers 5-merous. Calyx glabrous on both sides, sepals rounded, | mm long. Corolla 10-15 mm long, the tube 4-5 times as long as the limb, outside glab- rous, inside densely woolly in the mouth. Stamens inserted in or slightly below the mouth, filament very short, anther elliptic or oblong, 1-1/2 mm long, glabrous. Pistil c. 12-13 mm long, ovary glabrous, style hairy in the lower half. Pedicels in fruit strongly thickened. Fruits globular, 3-5 cm @, the wall c. 1/2 cm thick. Seeds unknown.
Distr. Assam, Burma, Thailand, Indo- China, and Malesia: probably in the Malay Peninsula.
Ecol. Lowland rain-forest.
Note. Dr. N. G. Bisset, London, in- formed me that he had seen sterile material of this species collected in the
6:
: 363a
365
: 365b
: 366a
: 3665
: 367a
Salta
S/2a
—3575a
ons
= 35a
1a
Malay Peninsula.
24. Strychnos melanocarpa GILG & BE- NED.
Gardneria ovata WALL.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 210.
Neuburgia BL.
Add to Notes: Some species of Psychotria (Rubiaceae-Coffeoideae) show a_ great resemblance to Neuburgia. They differ mainly by the connate stipules not adnate to the petioles, by the cupular 5-toothed calyx, the glabrous anthers, and the triangular style with a truncate stigma; the ovary is for the main part superior but the fruits are inferior.
Neuburgia corynocarpa (A. GRAY) LEENH. Add to Uses: In NE. New Guinea, Finisterre Mts, the wood was formerly used for making bowls and plates.
Add to Notes: For an opinion as to specific delimitation in the Pacific contrary to the one expressed here see A. C. SMITH, Pac. Sci. 23 (1969) 387. Neuburgia sarcantha (GILG & BENED.) LEENH.
Add to description: Leaves from 7 by 51/2 cm, nerves 5-8 pairs.
Neuburgia kochii (VAL.) LEENH.
Add to description, Ist sentence: ap- parently sometimes a creeper (NGF 14779). Add to leaves, entry on the base: rarely rounded to cordate.
Neuburgia rumphiana LEENH.
Add to Distr.: Terr. of New Guinea (Sepik Distr.).
Neuburgia celebica (KOORD.) LEENH. Add to Distr.: New Britain, Solomon Is. Geniostoma rupestre FORST.
Add to literature: BACK. & BAKH. /f. FI. Java 2 (1965) 207.
Add to Notes, at the end of the Ist paragraph: An example of such a local subdivision has been given by BACKER & BAKHUIZEN f., J/.c., who distinguish among the material from Java between 3 “microspecies’.
Geniostoma arfakense KAN. & HAT. Add to description, under fruits: purple when ripe.
Cynoctonum GMEL.
The correct name and citation for the genus is Mitreola LINNE, Opera Varia (1758) 214, validated by indirect refer- ence to Gen. PI. ed. 1 (1737) 377. Cynoctonum mitreola (L.) BRITT.
The correct name is: Mitreola petiolata (GMEL.) Torr. & GRAY, FI. N. Am. 2 (1841) 45.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 208; TrReEL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 132, t. 9.
Add to Distr.: W. Africa (possibly in- troduced, cf. Herne, Kew Bull. 17, 1963,
6: 378a
6: 378b
6: 380b 6: 381b
6: 382a
6: 383a
6: 384a
FLORA MALESIANA
[ser. I, vol. 66
17D) Cynoctonum sphaerocarpum LEENH. The name should be corrected as follows: Mitreola sphaerocarpa (LEENH.) LEENH., nov. comb. Add to Note, distribution of C. pedicel- latum: Assam (Khasi Hills). A third species of the same Asian relationship is Mitreola reticulata TirEL, Adansonia II, 9 (1969) 119; Logan. Cambodge, Laos & Vietnam (1970) 136, t. 10, only known from N. Vietnam. Spigelia anthelmia L. Add to literature: BACK. & BAKH. f/f. FI. Java 2 (1965) 207. Add to Distr.: Introduced in Java 1845, cf. Back. Trop. Natuur Jubileumuitg. (1936) 54. Mitrasacme elata R.BR. Add to Distr.: Flores. var. brevicalyx LEENH. Add to Distr.: Flores, SE. New Guinea. Add to Ecol.: from c. 30 m onwards. Mitrasacme pygmaea R.BR. Add to literature: BACK. & BAKH. /. FI. Java 2 (1965) 208; TiREL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 148, t. 11 f. 10-16. (sub M. nudicaulis Bru.): De Voocp, Trop. Natuur 30 (1941) 103, f. 4. var. pygmaea. Add to Distr.: Sumbawa. Drop: Mt Merapi, Bali, Lombok, Timor. Insert after 2. Mitrasacme pygmaea R.BR.: 2a. Mitrasacme erophila Le&ENH. Bull. Jard. Bot. Brux. 32 (1962) 446, f. 72; TIREL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 143, t. 11 f. 1-5. — M. pygmaea R.Br. var. pygmaea LEENH. FI. Mal. I, 6 (1962) 383 p.p., incl. f. 43. Erect, annual herb, up to 15 cm high, simple or branched at base. Leaves rosulate, (oblong-)lanceolate, 3-6 by 3/4—2 mm, herbaceous, puberulous above, glabrous beneath, base cuneate, apex acute, 1-nerved. Jnflorescences laxly umbellate, few-flowered; peduncle 4-11 cm long, terete, puberulous at base, with some scattered pairsof scale-like, 1—11/2 mm long empty bracts; pedicels !/2—-21/e cm long. Ca/yx campanulate, 2 mm long, glabrous or puberulous, the lobes trian- gular, acute. Corolla widely infundibu- liform, 31/2-4 mm long, white, hairy in the throat, the lobes c. 11/2 mm long, oblong or obovate, rounded. Stamens inserted in the tube at c. 2/3 of the height; filament 1/2 mm long; anther orbicular, 1/7 mm &, cleft at base, dehiscing all around, ciliate along the margin. Pistil 2'/4 mm high, styles nearly completely connate, stigma deeply bilobed. Capsule globular, 11/2-2 mm @, styles short, free. Seed angular-ovoid, brown, reticulate.
6: 385a
6: 386a
6: 386b
Sea27b
4: 45a
5: 345
Distr. Assam and Malesia: Central Java (Merbabu-Merapi), Lesser Sunda Is. (Bali, Lombok, Alor, Timor).
Ecol. Grassland, Eucalyptus-forest etc., locally often common, from 650- 2000 m, on Bali, Mt Agung, near solfata- ra, even at 3150 mm. Fi. fr. March-June.
Note. Originally, when revising the genus Mitrasacme for the Flora Malesia- na, I did not recognize this material as different from M. pygmaea var. pygmaea, even though later onI described the pres- ent species as new from Assam. For differences with M. pygmaea see there. Mitrasacme indica WIGHT.
Add to literature: BACK. & Baku. f. FI. Java 2 (1965) 208; TiREL-ROUDET, Logan. Cambodge, Laos & Vietnam (1970) 145. Add to synonymy: Non RIDSDALE, Trans. Papua N. G. Sci. Soc. 9 (1968) 18 (=Lindernia subulata R.BR.). Mitrasacme saxatilis BACK.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 208.
Mitrasacme neglecta LEENH.
Add to literature and synonymy: BACK. & BAKH. f. Fl. Java 2 (1965) 209. — Evolvulus sp. RIDSDALE, Trans. Papua N. G. Sci. Soc. 9 (1968) 17.
Add to Distr.: SE. New Guinea. Mitrasacme bogoriensis LEENH.
Add to literature: BACK. & BAKH. f. FI. Java 2 (1965) 209.
Add to Excluded:
Logania dentata (ELM.) HAYATA, J. Coll. Sc. Imp. Un. Tokyo 25, art. 19 (1908) 162, t. 28. — Nertera dentata ELM. Leafl. Philip. Bot. 1 (1906) 15 = Hemiphragma heterophylla WA. (Scrophulariaceae). Cf. Merr. in Hayata, /c.; HALL. f/f. Beih. Bot. Centralbl. 39, 2 (1923) 161.
Malpighiaceae (JACOBS)
Aspidopterys elliptica (BL.) Juss.
Add to Distr.: Lesser Sunda Is. (Bali, KOSTERMANS 272, distributed as Tristel- lateia; Flores, cf. STEEN. Blumea 15, 1967, 153).
Moringaceae
Moringa oleifera LAMK. Add to synonymy: M. domestica HAM. ex HENSCHEL, Clavis Rumph. (1833) 44.
Pittosporaceae
In the revision of Pittosporum only 3 endemic species were known from New Guinea which stands in contrast with the large number of endemic species in New Caledonia and also the much higher number in Australia.
It has appeared, however, that further
September 1972]
5: 348
5: 348
exploration has yielded more novelties in
the last decade, partly described by Dr.
BAKKER, partly by Dr. ScHopbe. But I
hesitate to share the view of the latter that
East New Guinea would represent a
‘centre of diversity’ for the genus. It may
later appear that some novelties are
possibly marginal extremes of other spe- cies and I feel that hybridization is also not excluded. | cannot recognize two of
SCHODDE’s taxa at specific level.
In the Key to the species (flowering
material), replace the first entry of fork 5
by:
5. Flowers always axillary, solitary.
5a. Leaves mostly small, obovate, c. 2-4(-81/2) by 1-3(-4) cm, abruptly shortly acute-acuminate, coriaceous. Reticulation conspicuous, prom- inent on both sides. Twigs glab- rous. Flowers 12 mm. Ovary glab- rous. Fruit 12 by 8 mm.
5. P. berberidoides
Sa. Leaves small, obovate to spathulate, 3-25 by 2-10 mm, glabrous, herb- aceous; apex broadly cuneate, with a blunt tip; nerves not prominent, few. Twigs covered by a persistent indument of short, thickish ferrug- ineous hairs. Flowers 8 mm long. Ovary with a few long hairs at the base . . . P. inopinatum
In the Key to the species (dry fruiting
material), replace forks 5 and 6 by:
5. Nerves and larger veins distinctly prominent on both leaf surfaces. Leaves obovate, + pointed, c. 2-4 by 1-3 cm.
5a. Main nerves c. 10-15 pairs, close. Leaf apex suddenly acute-acumina- tely contracted. Young twigs glab- rous 5. P. berberidoides
5a. Main nerves 4-6 pairs. Leaves with rather bluntish apex, in pseudo- verticils. Young twigs densely fer- rugineous-hairy 5b. P. pumilum
5. Nerves and larger veins not prominent above.
6. Fruits longer than broad (not includ-
ing the stipe and apical mucro).
6a. Twigs densely persistently set with dark-brown thickish hairs. Leaves small, 3-25 by 2-10 mm, spathula- te, herbaceous, spirally arranged, with blunt tip; nerves 3-4 pairs, rather indistinct above (neither impressed nor prominent). Fruit valves thickish. 5a. P. inopinatum
6a. Otherwise.
7. Remains unaltered.
7. Remains unaltered.
6. Fruits as long as broad or slightly
broader than long. 7a. Fruits globular, 11-13 mm @, with thick valves. Leaves not in
Addenda, corrigenda et emendanda
961
pseudo-verticils, very thick, small, + bullate, 6-14 by 3-7 mm, with 3-4 pairs of nerves which are im- pressed above. 6a. P. pullifolium var. globosa 7a. Leaves much larger with many more pairs of nerves which are not impressed above.
5: 349a Pittosporum sinuatum BL.
5: 349b
5: 350b
33-3516
D324
5: 353a
Add to description: A specimen from East New Guinea (Brass 30801) has entirely glabrous, slender ovaries in pre- dominantly ¢ flowers; all other material has pubescent ovaries.
Add after 1. Pittosporum sinuatum the following variety:
la. var. efuniculare STEEN. var. nov. — P. tenuivalve SCHODDE, Blumea 15 (1967) AVipaiee:
Funiculus brevipes. Typus ROBBINS 888.
Falls within the variable P. sinuatum, differs obviously only by the short fu- nicles.
Distr. Malesia: East New Guinea (Madang Distr., Kubor Range: Ros- BINS 888, 1121), not seen.
Ecol. Montane forest, 2100-2300 m.
Note. I agree with Dr. SCHODDE’s tentative suggestion that this represents perhaps only a mountain form of P. sinuatum.
Pittosporum ramiflorum (ZoLL. & Mor.) ZOLL. ex MiqQ.
Add to Distr.: Lesser Sunda Is. (Sum- bawa). Cf. BAKKER, Blumea 11 (1962) 426.
Line 6 from top, add after P. ramiflorum f. macrocarpum: Another specimen, again from NW. New Guinea, Vogelkop Peninsula, was collected (BW 6880). It can easily be distinguished from the equally large-fruited Philippine P. resini- ferum by the absence of the large resini- ferous cavities in the fruit valves. Cf. BAKKER, Blumea 11 (1962) 426. Pittosporum resiniferum HEMSL.
Omit in key and description that fruit is compressed; this is only so in immature ones and caused by pressure. It is really very broad-ellipsoid, tending to globular; occasionally it is 3-valved.
Add to Distr.: Solomon Is. (Bougainvil- le: SCHODDE 3767), climbing tree, in fr., 800 m. Hitherto only known from N. Borneo and Philippines.
Pittosporum berberidoides BURKILL.
Add to description: Tree up to 14 m, stem to 20 cm @. Leaves up to 12 by 41/5 cm; petiole to 2 cm. Cf. also BAKKER, Nova Guinea n.s. 9 (1958) 339; Blumea 11 (4962) 426.
Add to Distr.: Now found in many other places (Mt Wilhelm; Chimbu; Sepik- Wangi Divide; Goroka, efc.) and as low
962
FLORA MALESIANA
[ser. I, vol. 66
Fig. 11. Pittosporum inopinatum BAKKER. a. Habit, nat. size, 6. flower, <3, c. petal, «5, d. style, 7, f. fruit, nat. size, g. seeds, <2 (a-g ROBBINS 829),
5: 353a
down as 2400 m. Add after 5. Pittosporum berberidoides BURKILL:
5a. Pittosporum inopinatum BAKKER, No- va Guinea n.s. 9 (1958) 339, f. 1. — Fig. 11.
Much-branched shrub or dwarf tree, 11/2-21/2 m. Twigs covered with a persistent indument of short, thickish ferrugineous hairs; internodes up to c. 11/2 cm. Leaves spirally arranged, herbaceous, obovate to spathulate, entire, 3-25 by 2-10 mm, glabrous, apex broadly cuneate with a blunt tip, base decurrent, cuneate; nerves 3-4 pairs, indistinct above, not prominent on either surface; petiole c. 1-2 mm below the decurrent leaf base. Flowers solitary, axillary, pendent, c. 8 mm. Pedicels ferrugineous-hairy, c. 4 mm. Bracts triangular, very narrow, acute, c. 2 mm. Sepals free, narrow-oblong, acute, c. 2 by ?/4-1 mm, yellowish green with a purple top. Petals free though cohering, ligulate, narrowing towards the rounded tip, c. 7-8 by 2 mm, pink or purplish especially at the top. Stamens sagittiform, filaments narrowing towards the anthers, c. 21/2 mm; anthers c. 1 by 1/2 mm. Ovary depressed-ellipsoid, glabrous, c. 3 by 2 mm; style glabrous, c. 1 mm; stigma capitate. Fruit 2-valved, ellipsoid, c. 11/2 by 1 cm; valves coriaceous, orange-yellow to brown, rather thin; placentas slightly raised; funicles c. 10-12
on each placenta, inserted up to ec. 2/3 et the length of the valves, in ripe seeds up to c. 3 mm long, very dark violet. Seeds c. 4-6 on each placenta, irregular- globose, dark violet, c.4 mm @.
Distr. Malesia: East New Guinea (Mt Otto, 3 collections; Mt Piora, NGF 16535).
Ecol. Mountain forests of Podocarpus- Libocedrus, 2550-2860 m, said to be common. Fi. fr. Sept.
5b. Pittosporum pumilum SCHODDE, Blu- mea 15 (1967) 406, f. 1.
Slender shrub, 1 m. Twigs densely ferrugineous-hairy; internodes with a few tiny cataphylls. Leaves mostly in pseudo-verticils, spathulate, with rather rounded apex, glabrous, thinly coria- ceous, 11/2-41/2 by 1/2-2 cm; nerves 4-6 pairs, on both surfaces prominent; base attenuate, a petiole hardly discernible. Fruit solitary, pseudo-terminal, later subterminal, on a rather stout glabrescent peduncle 5-8 mm long, ellipsoid, when young shortly stipitate and rostellate, 21/2-3 by 11/2 cm, glabrescent, red (drying orange-yellow); valves outside rugose, inside without transverse ribs and set with funicles all along their length. Seeds c. 8, semi-reniform, 5mm, on rather long funicles, reddish brown, drying black.
Distr. Malesia: East New Guinea (Morobe Distr.: Mt Shungol), one col- lection.
September 1972]
Addenda, corrigenda et emendanda
963
5: 3536
5: 354a
5: 3554
5: 356b
5: 3605
Ecol. Lower mossy forest, 2100 m. Fr. Dec;
Note. According to SCHODDE most related to P. sinuatum, but to my opin- ion most allied to P. berberidoides through the remarkably prominent veins. Pittosporum pullifolium BURKILL.
Add to description: A specimen with young fruit had the _ infructescences axillary on the twigs instead of terminal. Cf. BAKKER, Blumea 11 (1962) 426.
Add after 6. Pittosporum pullifolium the following variety:
6a. var. globosum STEEN. var. nov. — P. nubicola SCHODDE, Blumea 15 (1967) 411, f. 3-4.
Differt ab species: Capsula_ globosa, I]-Il3 mm 2.
Shrub 1-2 m. Leaves thick, bullate by recurved margins, c. 10 by 5-6 mm; midrib and c. 3 pairs of nerves impressed above. Infructescence of c. 5 fruits apical; capsules globose, 11-13 mm @.
Distr. Malesia: East New Guinea (Kubor Range: Mt Kinkain), one collec- tion.
Ecol. Border of alpine shrubbery adjacent to peaty grassland, at 3600 m. Fr. July.
Note. As SCHODDE already remarked close to P. pullifolium, of which the alpine dwarfed forms have similarly small leaves, and in fact grew to near 100 m from this form which differs by the globo- se fruit.
Pittosporum pentandrum (BLANCO) MERR. Add to Distr.: N. Borneo. Cf. BAKKER, Blumea 11 (1962) 426.
Pittosporum moluccanum (LAMK) Mia. Add to synonymy: Vareca moluccana Roxs. FI. Ind. ed. Carey 1 (1832) 647.
In Fl. Mal. I, 5 (1954) 33, this was excluded from Hydnocarpus and _ tenta- tively assigned to Rinorea. ROXBURGH’S type (BM) was kindly identified by Mr. H. K. Atry SHAW (in /itt. to Dr. JACoBs, cf. Blumea 15, 1967, 138). No change of epithet is necessary.
Add to Excluded:
Pittosporum serrulatum JACK ex ROXB. Fl. Ind. 2 (1824) 401; ex Grirr. Calc. J. Nat. Hist. 4 (1843) 195; cf. Merr. J. Arn. Arb. 33 (1952) 240= Rinorea lanceolata (Roxs.) O.K. (Violaceae).
Podostemaceae
Add to Distr.: The knowledge of the distribution of the family in Indo- Australia is still expanding. A new spe- cies, of Indotristicha, has been found in Malaya. TUYAMA & Hara (J. Jap. Bot. 39, 1964, 185-188) mapped Asian local- ities, which comprise amongst others
localities in the East Himalaya, the
Chinese mainland in Fukien, Kwantung,
Kanton (TUYAMA & Hara, l/c. 185;
CHAO, Contr. Inst. Bot. Nat. Peiping 6,
1948) and Hainan (CHUN, FI. Hainanica,
1964, 373), Thailand (VAN ROoyEN,
Blumea 10, 1960, 141; Dansk Bot. Ark.
23, 1965, 185) and the Ryu Kyu Is. Prob-
ably the range of the family extends by
several genera all over SE. and E. Asia
(tropical to warm-temperate), but the
small plants are evasive to collectors.
Recently, an undoubted member has also
been found by Mr. Byrnes in the Kim-
berley District, NW. Australia; this is distinctly different from Torrenticola from Queensland. It bears resemblance to
Indotristicha malayana, but has only 2
stamens. Miss ASHTON, Melbourne, has
tentatively referred it to Tristicha trifaria
(Bory ex WILLD.) SPRENG., a species
known from the tropics of America and
Africa. Replace the Key to the genera
into the following:
1. Leafy (flowering) stems very short, hardly 10 mm long, with scattered, imbricate, 3—7-segmented leaves. Spa- thella oval, nippled, usually dehiscing irregularly. Tepals 2, narrow. Stamen 1. Stigma 2 avis, like Gladopus
1. Leafy (flowering) stems 25-100 mm, often branched. Leaves distichous, laterally compressed, the lower entire, the upper 3(-4)-dentate. Spathella oval, tipped, irregularly circumscissile-de- hiscent. Tepals 2, narrow. Stamen |. Stigmas 2. . . 2. Torrenticola
1. Leafy shoots up to 20 mm, rarely branched, with imbricate triangular leaves in 3 ranks, those of 2 ranks 11/2 by !/2 mm, those of the 3rd rank 1 by 3/4 mm. No spathella. Tepals 3, half- way connate. Stamens 3. Ovary 9- veined. Stigmas 3. 3. Indotristicha
Cladopus H. MOLL.
Add to Distr.: SE. Asia (Siam, Hainan,
Kwantung, Kanton, Fukien) and Ryu
Kyu Is. Possibly monotypic.
Cladopus nymani H. MOLL.
Add to Distr.: Possibly all the names and
localities in SE. & E. Asia of Cladopus
refer only to one species; see VAN ROYEN,
Dansk Bot. Ark. 23 (1965) 185.
Add after 2. Torrenticola etc.:
3. INDOTRISTICHA
VAN RoyeEN, Acta Bot. Neerl. 8 (1959) 474; Baku. f. Taxon 18 (1969) 598. — Dalzellia (non WiGHT) ENGL. Nat. Pf. Fam. Nachtr. 3 (1908) 135-136; ibid. ed. 2, 18a (1930) 33, f. 24.
See for the main characters in the key.
Distr. Species 2, one in India (W. Ghats and S. Canara to Travancore), one in Malaya.
FLORA MALESIANA
[ser. I, vol. 66
Fig. 12. Indotristicha malayana DRANSF. & WHITMORE. a. Habit, sterile, < 3, b. ditto, fertile, = 3, c. tepals,
<7, d-g, four stages in the development from flower to fruit, » 14.
1. Indotristicha malayana DRANSF. & WHITMORE, Blumea 18 (1970) 154, pl. 1, f. 1. — Fig. 12.
Rhizome creeping, flattened, thalloid, closely adpressed to the substratum, 1/2 mm wide, occasionally to 1 mm, of indefinite length, branching irregularly. Leafy shoots rarely flowering, borne irregularly, usually closely, to 2 cm long, rarely branching, usually curved, pale green and attenuate in shade, tinged red and shorter in full sun, irridescent. Stem triangular in section. Leaves sessile in very close whorls of 3 unequal ranks, the bases touching, increasing in size to about two thirds of the length from the rhizome, and thence decreasing slightly; two ranks long triangular to 11/2 by '/o mm, outline slightly asymmetric recurved, convex towards the third rank which is broadly triangular to 1 by 0.7 mm. Also with flower bearing leafy shoots, often clustered, with smaller rather distant leaves. Flowers protogyn- ous, lateral, solitary; peduncle slender, stiff, erect, to 5 mm long at anthesis, subtended by two unequal, concave, sometimes mucronate, chartaceous im- bricate bracts to 1 by 1/2 mm. Tepals 3, valvate, obovate, 1 by 0.8 mm, slightly concave, chartaceous, translucent. Sta- mens 3, at first hidden by tepals, later at anthesis longly exserted to 1 mm on stiff, erect, pink filaments, later shrivelling and becoming thread-like and twisted ; anthers ovate-oblong, 0.4 mm long, cream. Ovary pale straw-coloured, obovoid, becoming narrowly obovoid by anthesis to 1.6 by 1 mm with 9 rib-like veins; styles short, pink, 3, flattened in a vertical plane, with a deeply divided outer margin; placenta- tion free central, ovules numerous, tiny, cylindrical. Capsule narrowly obovoid,
4: 258b
6: 180a
2 by 1.3 mm, light brown, crowned by remains of styles.
Distr. Malesia: Malay Peninsula (Pahang, Trengganu), at least 3 localities.
Pontederiaceae
Monochoria hastata (L.) SOLMS.
Add to Distr.: Now also recorded from Australia: Northern Territory, Litch- field Homestead. Cf. Muelleria 2 (1971) 134.
Primulaceae
Lysimachia capillipes HEMSL.
Add to Distr.: East New Guinea (Moro- be Distr.: Bulolo, NGF 30727, Wau, TGH 11596, NGF 35800; Eastern Highl. Distr.: NGF 27158), 100-1700 m. Fi. fr. Jan.-July.
Note. The Papuan material is homo- geneous and is tentatively best classified with the Chinese-Luzon distributed L. capillipes. It must be remarked that the anthers have no permanent apical pore; they seem to dehisce with an apical slit- like pore which later extends downwards to a completely lengthwise splitting of the anther-cells. The calyx is narrow in all specimens (in contrast with that of L. laxa BAUDO) but its length seems to vary in proportion to the length of the capsule. It seems to be not always exceeding the capsule, but this could not well be checked as most capsules were dehisced and the more or less flattened valves are of course much longer than the undehisc- ed capsule. The Papuan specimens are more-stemmed but rather erect. More and better preserved material is desirable for a definite conclusion. — L. R6st, Oct. 1971.
September 1972]
Addenda, corrigenda et emendanda
965
6: 1855
6: 192
SAD 5: 192a
Sie 4b
5: 444
Lysimachia decurrens Forsv. f.
Add to Distr. and map (fig. 10): SW. Central Celebes (Latimodjong Ra., KJELL- BERG 1425). Cf. BENTVELZEN, Blumea 13 (1965) 140.
Add to the excluded names:
Hottonia sessiliflora VAHL, Symb. 2 (1791) 36, the type of which was describ- ed from ‘India orientalis’ (C), from BURMAN’s herbarium; it belongs accord- ing to VAN DER MEWDEN (Blumea 17, 1969, 311) to a species of Limnophila (Scrophulariaceae). The epithet is al- ready occupied in that genus.
Proteaceae (SLEUMER)
Helicia rufescens PRAIN.
Add to Distr.: Borneo (Brunei). Heliciopsis rufidula SLEUM.
Add to Distr.: Borneo (Sarawak).
Rhizophoraceae
Range maps of many species were given by Dinc Hou in Sreen. Pacific Plant Areas | (1963) maps 2-10, 20, 23. Distribution past and present. The second paragraph refers to the peculiar distribu- tion of mangrove species, Rhizophoraceae and otherwise, of which no species is in common to those in the Atlantic and In- dian Oceans. As a matter of fact this holds also for marine Phanerogams. On the other hand it has appeared that at least of the genus Rhizophora the three species of the Caribbean occur also on the west coast of Africa and that at least two of these also occur on the Pacific coast of tropical America. Cf. Dinc Hou, Blumea 10 (1960) 625-634. This means that there must have been in the past an open seaway between the Americas and this is corroborated by the geology. Africa seems to have been a distinct bar- rier between the Indian and Atlantic Oceans.
I have discussed these major phytogzo- graphical features in 1962 in a pader (Proc. Kon. Akad. Wet. A’dam ser. C, 65: 164-169) in which it was also shown that the marine Rhizophoraceae seem to have originated in the Indo-Malesian tropics where all 4 genera are centred, the Americas having only 3 out of 7 species of the one genus Rhizophora.
As to the African barrier, it seemed to me that whereas even today species of the shore plants of the pes-caprae and Barringtonia formations cannot round the Cape of Good Hope because of temper- ature conditions and oceanic circulation system, the temperature must have been the chief factor that the dispersal of more megatherm mangrove species — of
5: 445
5: 448
which dispersal is far more restricted by the viviparous drifting embryos which can only disperse in non-turbulent water and near-shore shallow muddy coasts — could not be effective even under the more ameliorated Tertiary climatic con- ditions.
Mr. MuLLER (Review Palaeobot. Palyn. 6, 1968, 281-282) correctly stated that there was a northern open seaway, the Tethys, between the Indian and At- lantic Oceans and that this must have had an impact on diffusion of seaborne dispersed marine Phanerogams. He derives that the replacement of Atlantic and Indian Ocean species — if we pre- sume that the Tethys was an effective dispersal route indeed (including suitable ecological shore conditions!) — must be ascribed to the assumption that these Phanerogams had not yet evolved (or were at least not yet occurring in the Indian Ocean — y.St.) in the Early Tertiary and that when they came there the Tethys had by that time lost its capacity for their dispersal. That the Tethys must have had this function seems to be proved by the recent find of fossil Nypa pollen in West Africa (Upper Cretaceous to Upper Eocene), a genus which obviously got extinct there and in the Caribbean simultaneously for un- known reasons.
Unfortunately we have no clear view of the course of the sea current regime at the time of the Tethys Sea.
In addition we should mention that Rhi- zophoraceous pollen is only found as early as the Upper Eocene, but not (yet) in Paleo- cene or Upper Cretaceous sediments Cf. MULLER, Biol. Rev. 45 (1970) 434, f. 5.
One important conclusion can, I be- lieve, not be doubted, except by super- stitious diffusionists, namely that dispers- al of both mangroves and marine sea- grasses can only take place at short distances and their occurrence is testi- mony of near-landmasses in the past, in contrast to most beach and Barringtonia formation species which are indeed frequently equipped to perform long- distance waterborne dispersal.
Add to Taxonomy: According to Mr. MULLER (in Jitt.) the pollen of Carallia is similar to that of marine genera (Rhizo- phora, Bruguiera) and does not warrant a separation into a distinct family. Rhizophora L.
Add to Distr.: DING Hou has revised the entire genus Rhizophora after the account in Fl. Mal. was written. Though this has brought no changes in taxonomy or distribution, some ranges in the West Pacific appear to be more extensive: R. mucronata LAMK extends to Tonga,
966
FLORA MALESIANA
[ser. I, vol. 68
5: 453b
5 457
5: 46la
5: 464a
5: 468b
5: 474
5: 477a
5: 477a
5: 480
5: 483
R. apiculata Bui. to the Solomons and New Hebrides, R. stylosa GRIFF. is also in Micronesia (Guam & Marshalls). Cf. DinGc Hou, Blumea 10 (1960) 625-634, map. The proper references of Rhizophora mucronata seem to be: Porr. in LAMK, Tabl. Encycl. Méth. Bot. (text) 2 (1794) 517; LAMK, Tabl. 1 (1797) t. 396 f. 2; Porr. in LAMK, Encycl. 6 (1804) 189; DC. etc. Bruguiera LAMK. The date of publication of the genus is: 1797. Bruguiera gymnorrhiza (L.) LAMK. In references line 2 the year of publication is: 1798. SAVIGNY seems to be responsible for the treatment of the genus in LaA- MARCK’S work. Bruguiera exaristata DING Hou. Add to synonymy: B. eriopetala var. exsetata VALETON, Bull. Dép. Agr. Ind. Néerl. 10 (1907) 38 (type: Kocu s.n., L). Bruguiera hainesii C. G. ROGERS. Add to Distr.: Also found in the Sundri- bans. Cf. S. K. MUKERJEE, Bull. Bot. Surv. India 8 (1966) 357. Anisophyllea R.BR. ex SABINE. Add to description of embryo: This is solid and indeed consists of an almost undifferentiated hypocotyl which con- tains the reserve food, similar as in several other tropical tree genera e.g. Barring- tonia and Bertholletia. From this ‘pre- adapted’ structure one can imagine the origin of the further differentiated em- bryogeny of the marine species.
Anisophyllea beccariana BAILLON.
Add to Distr.: Central East Sumatra
(Tenajan R., SOEPADMO 37).
Anisophyllea ferruginea DING Hou.
Add to description: Mature leaves gla-
brous. Inflorescences up to 4cm. Fruits
1(-2)-seeded, 8 by 4 cm, glabrous; pericarp 8-10 mm @, exocarp soft. Seeds very hard, solid and large, 51/2 by 2 cm.
The two additional specimens (ASHTON
BRUN 580, S 7867) have mature in-
florescences and fruit. In mature state the
rusty tomentum disappears.
Combretocarpus Hook. f. and C. rotun-
datus (MIQ.) DANSER.
Add to Distr.: Malay Peninsula (Johore),
in peat swamp forest of Ayer Hitam
South For. Res., occupying c. 30% of
the big trees in the Reserve. Cf. F.S.P.
Na, Mal. For. 29 (1966) 32-33, 1 fig.
Carallia Roxs.
Replace in the Key forks 5 & 6 into the
following:
5. Petals persistent. Calyx lobes hairy inside or at least at base or margin. Seed obovoid or ellipsoid. Embryo curved.
6. Leaf margin distinctly fimbriate.
5: 484a
5: 485a
Disk slightly crenulate. Fruit more
than twice as long as wide, c. 15 by
6mm 6. C. suffruticosa 6. Leaf margin entire or almost so, at most very short-dentate or crenulate.
Disk distinctly lobed halfway. Fruit
less than twice as long as wide. 6a. Cymes only in the axils of the upper pair of leaves of each shoot. Stip- ules up to 16 mm long. Calyx lobes 3-4 mm. Petals 3-4 mm. Seeds c. 9 by 6 mm. 5. C. borneensis Cymes in the axils of successive leaf pairs of each shoot. Stipules c. 5 mm long. Calyx lobes c. 11/2 mm long. Petals c. 21/4 by 1 mm. Seeds c. 8 by 3 mm. 5a. C. longipes 5. Petals caducous. Calyx lobes glabrous.
Seed reniform or curved. Embryo
curved.
Carallia eugenioidea KING.
Add to Distr.: Indo-China (Vietnam). Cf. VU VON CUONG, FI. Camb. Laos & Vietn. 4 (1965) 172.
Add the following species:
5a. Carallia longipes DING Hou, Nova Guinea, Bot. n. 4 (1960) 21-23, f. 4. — Fig. 13.
Shrub, 3-5 m. Leaves chartaceous to thin-coriaceous, elliptic-oblong or ellip- tic, sometimes ovate, 31/2-61/2 by 11/2- 31/2 cm, base cuneate rarely rounded, apex acute, margin obscurely serrulate in upper half; nerves 5—7 pairs, + elevated on both surfaces, veins reticulate, + elevated above, visible or obscure be- neath; petiole 5-10 mm; 2-4 small, with conical to subulate appendages at the base on each side, persistent but breaking off easily. Stipules c. 5 mm. Inflorescences up to 7 cm long, axillary, solitary, on several nodes along the young branches, usually few-flowered cincinnal cymes; internodes distinct, up to 11/2 cm, very rarely with a few short internodes between the long ones; peduncles 1!/2-4 cm. Flowers sessile, not resinous, slightly obovoid-oblong, c. 6 by 3 mm. Calyx lobes 5, ovate, c. 1.6 by 1 mm, acuminate, sparsely puberulous on the margins and towards the base inside. Petals persistent, obovate-spathulate, c. 21/4 by 1 mm, unguiculate, lamina short-fimbriate. Fi/- aments 31/2-41/2 mm, the lower 2/5 united into a tube, free parts fleshy; anthers small, c. 0.4 by !/4 mm, obtuse. Disk fleshy, distinctly 10-lobed. Ovary l-celled, 10-ovuled; style cylindric, c. 31/2 mm, slightly and gradually thickened towards the base; stigma obscurely capitate. Fruits dark-red, broadly el- lipsoid or subglobose, 13-15 by 10-13 mm. Seed 1, oblong-obovoid, slightly ribbed, c. 8 by 3 mm, reddish to dark brown. Embryo straight.
6a.
September 1972]
Fig. 13. Carallia longipes DING Hou. a. Habit, 2/3, b. apex of branchlet showing appendages outside
the base of the petioles and stipule,
<2, c. inside view of stipule showing colleters, «7, d. node of
branchlet, <3, e. flower, <3, f. longitudinal section of flower (style removed), «7, g. petal, «13, A.
stamen, x 13, 7. style and stigma, <7, /. cross-section of ovary, <7, k. yGung fruit,
5: 4885
5: 491b
SPIE eee Se
(a & e-k BW 4980, b-d & | BW 7371).
Distr. Malesia: West New Guinea (Vogelkop Peninsula).
Ecol. Lowland forest, on limestone, 275 m.
Note. Closely related to C. papuana DING Hou.
Gynotroches axillaris Bu.
Omit under Distr. ‘Australia’; cf. DING Hou & STEEN. Pac. Pl. Areas 1 (1963) 284, map 20.
Though the distribution was correctly mentioned under the genus, Australia was by error mentioned in the distribution of the species, which induced Miss BURBIDGE to enter this generic record in the Dict. Austr. Pl. Gen. (1963) 141. It is of course not excluded that this may be found in future in the N. Queensland rain-forest, as it is abundant in New Guinea and the Solomons, but at present there is no proof for this.
Pellacalyx saccardianus SCORT. Add to Distr.: Also in Peninsular Thai- land (near Trang, RFD 35161).
Sarcospermataceae
4: 3465 Sarcosperma uittienti H. J. LAM.
S297.
Add to Distr.: In addition to the 2 spe- cimens mentioned by H. J. LAM & VAROSSIEAU, Blumea 3 (1938) 194: Sumatra (Eastcoast, Wampa Valley, 550 m, GALOENGI 432; Karo country, Biang Valley, 800 m, LORZING 14457) and Malay Peninsula (Selangor, Ulu Gom- bak, 600 m, SF 34191 Mow. Nur, FRI 1950 FRANCIS NG).
Scyphostegiaceae
Scyphostegia STAPF and S. borneensis STAPF, — Fig. 14.
At my suggestion Dr. W. A. VAN HEEL made a detailed anatomical-morphologi- cal investigation because Dr. J. Hut- CHINSON in the 2nd edition of his ‘The Families of Flowering Plants’ 1 (1959) 326-329, f. 187 a—b, did still accept a disharmony between the 3 and ° flowers,
968 FLORA MALESIANA [ser. I, vol. 68
Fig. 14. Scyphostegia borneensis STAPF. a. Ls. of the ovule at the stage of the first formation of nuclear endosperm, the funicle shows a con- striction just below the meristem (indicated by a cross) of the later formed arilloid, x20, b. l.s. of young seed, with the arilloid developed above the constriction, embryo still small, 3, c. fully mature, dehisced fruit, the fleshy pericarp valves each with a part of the stigma on top, the bunch of dark, erect seeds attached to the base of the fruit, at base each surrounded by the (pale) arilloid, x 4/3 (after VAN HEEL).
the latter being accepted as a fleshy receptacle containing numerous achenes, thus remaining at variance with SwAmMy (Proc. Nat. Inst. India 19, 2, 1953, 127-142) and myself (Fl. Mal. I, 5, 1957, 297-299), who interpreted the fruits (‘achenes’) of HUTCHINSON as ovules. Furthermore, HUTCHINSON placed the family in Celastrales, near Siphonodon, probably because of the similarity of the
6: 193
thickened stigma.
VAN HEEL has found (Blumea 15, 1967, 107-125, 13 fig.) that the reproductive units are doubtless seeds, that Hut- CHINSON’S ‘receptacle’ is a pericarp, and that his assumed ‘tepals’ in the 2 flower are really an aril(loid). Besides, the ovule shows some remarkable char- acters, namely a pedestal funicle with a constriction, a protuberance on the chalazal side, and a 5-lobed exostome.
In my description of the fruit I had already observed that in later stages it was breaking up at the mouth. Dr. W. MEIER, to whom we are extremely obliged for the very complete pickled material, succeeded in finding in Sabah the final fruiting stage in which the pericarp is split to the base into 9-12 valves; it is really a fleshy capsule, but is probably swallowed by animals in im- mature state as happens with so many fleshy fruits in Malesia. Fig. 14.
Our suggestion of affinity with Fla- courtiaceae seems to be not unreasonable to VAN HEEL, from the morphological viewpoint the only discrepancy being that the placentation in the latter family is as far as known lateral-parietal and never basal as in Scyphostegia. But the pla- centation in Tamaricaceae, also listed in Parietales, seems intermediate, according to VAN HEEL, who will pursue further studies in Flacourtiaceae for checking this.
It is curious to find that HUTCHINSON even in his latest work ‘Evolution and Phylogeny of Flowering Plants’ (1969: 360, f. 310 A, B) maintains his view that the ovary of Scyphostegia represents a disk concealing free carpels, completely neglecting the detailed morphological work by VAN HEEL which I had personal- ly brought to his attention in 1967. A remarkable case of prejudice leading to quixotic stubbornness which as far as I know finds no parallel in botanical science.
Simaroubaceae (NOOTEBOOM)
Line 5 from top, replace ‘Samadera’ by: Quassia.
Line 6 from top, after ‘Irvingia’ add: Allantospermum.
Line 2 from bottom, replace ‘AUBL’ by: L.
Line 6 from bottom, after ‘Klainedoxa (erron. Klaineodoxa)y add: Allantosper- mum.
Add to the phytochemistry:
H. P. Nootesoom (Blumea 14, 1966, 309-315) made an additional study on phenolic compounds in the family of 13 Malesian species of 9 genera. This con-
September 1972]
6: 196
6: 196 6: 198
6: 201b 6: 2065 6: 218a 6: 220a
firms the suggestions that Simaroubaceae are fairly closely related to Rutaceae.
The genus /rvingia seems to fit well with Simaroubaceae.
But Suriana deviates distinctly from all others which would sustain the findings of GUTZWILLER (Bot. Jahrb. 81, 1961, 1-49) and others who regard this as a separate family.
Change in the Key to the genera:
1. Leaves simple.
2. Leaf-buds enclosed by caducous intra-petiolar stipules. Carpels con- nate. Stamens 10, without an adaxial scale. Plant not bitter.
2a. Stipules leaving conspicuous an-
nular scars. Ovary 2-celled. Fruit drupaceous . ... 9. Irvingia
2a. Stipular scars obscure. Ovary 5-
celled. Fruit a capsule. 10. Allantospermum
2. Stipules absent. Carpels free or con- nate. Stamens 5, or 10 and then with an adaxial scale.
Be SEO HE In Key fork 4 second lead line 2, replace ‘without’ by: with.
Quassia L. Add to references: NOOTEBOOM, Blumea
11 (15 Dec. 1962) 509-528, gave a new
subdivision of Quassia.
In caption fig. 4 replace ‘Q. harmandiana’ by: Eurycoma harmandiana.
Eurycoma harmandiana PIERRE.
Add: Distr. Fig. 4.
Lines 10 & 11 from top, replace “(K. & V.) Koorb.” by: (non K. & V.) Koorb. Ailanthus fordii NOOTEBOOM, FI. Mal. I, 6 (1962) 220. — Fig. 15.
In my world revision of the genus Ailanthus in Fl. Mal. (/.c. 215-220, f. 17-18) I described a new species, known only from one old collection from Hong Kong made by Forpb, a. 1884- 1886, and named this A. fordii, represent- ed in the Kew Herbarium and that of the British Museum (Nat. Hist.), London. Forp collected this in SE. Hong Kong I. ‘near Cape d’Aguilar, as a small but conspicuous emergent tree from the shrubberies, the bare trunk being sur- mounted with foliage like a palm, the leaves being c. 40 cm long.’
Thanks to the kind intermediary of Mr. D. R. W. ALEXANDER, Director of Urban Services, where the Hong Kong Herbarium is housed, the Forester of the Urban Services Department, Mr. Lau YUNG-SuM, who acts as a collector of plants and seeds, has found that in the original locality which is still fully intact, viz on the Cape d’Aguilar headland, 10 trees could be located. In Nov. 1968 only one tree was in flower and that proved to be male. But its identification
Addenda, corrigenda et emendanda 969
Fig. 15. Ailanthus fordii NOOTEBOOM. a. Fruit, b-d. germlings, the pericarp sometimes remaining at the base of the hypocotyl, sometimes elevated with the cotyledons, plumule central and germina- tion distinctly epigeal, seed germinated imme- diately after receipt, b—d after 2 weeks, e. germling with first leaves, 2 months later; all in greenhouse at Leyden. All x 3/7.
is fully certain, as the species differs from all others in both flowers and in fruit structure and could be easily checked on a duplicate forwarded to me and de- posited in the Rijksherbarium.
Mr. ALEXANDER added that ‘the flowering tree is c. 30 feet tall, growing between granite rocks in a steep ravine where it faces the westerly afternoon sun but is sheltered from the strong north- east wind. All ten trees have the same characteristics of a grey trunk, with surprisingly large bole at the _ base, tapering sharply upwards. Most of the trees are forked about two feet from the ground and continue upwards with two main stems. The wood appears brittle; the lower trunk carries numerous scars and callouses where the wood has grown over the sites of the broken-off branches. It appears likely that the tree is slow- growing. It is possible that more trees may be found on this headland which is about 2 km long and 1 km broad, rising at its highest point to 325 m. There are numerous ravines with big granite boul- ders. The undergrowth in parts is very thick and it is by no means easy to tra- verse the headland looking for Ailan- thus.
In examining the Hong Kong Herba- rium I found 4 Ailanthus sheets of A. fordii, all wrongly assigned to A. mala- barica.
FLORA MALESIANA
[ser. I, vol. 66
It is interesting to enumerate the specimens now known, and their locali- ties, herbarium specimens as well as living trees:
Hong Kong: Cape d’ Aguilar, 1884-1886 Cu. Forpb s.n. (K, BM); ditto, Y.S. Lau (Mr. LAU YUNG-SuM) 1438, Nov. 1968, tree 30 ft, in rocky ravine. Mt Gough, a spur of the Peak area, south, leg. W.J.T. (UTCHER), 24 Oct. 1905, Gard. Dept. Hong Kong Herb. No. 666, in flower; ditto, No. 667 leg. Mr. Lo (Quai), 16 Jan. in fruit; beside the Peak Tram track near its upper terminus, c. 400 m, a well-grown tree c. 10 m high (ALEXANDER, in /itt.).
New Territories: Sha Tin, leg.? 24 Jan. 1905, Bot. & For. Dept. No. 4715, in fruit; in Jan._March 1969 16 living trees located on hillsides (ALEXANDER, in litt.).
Cultivated: In N.B.G. (New Botanic Gardens) 93, leg. H. C. TANG, Gard. Dept. No. 12830, in flower, tall decidu- ous tree, bark grey, smooth, 16 March 1949. Mr. ALEXANDER reported that this cultivated tree is still alive. It is 12 m high, with a single trunk bare of branches for its first 5 m. It shows no thickened stem-base. It flowered Nov. 1968 and is male.
Notes. It is of course a pleasure to observe that this unique endemic of Hong Kong is still present in native vegetation and at least in its /Jocus classicus in such rough terrain that it will probably remain there for a long time to come. A Nature Reserve is, however, badly needed for it and we hope that the Hong Kong Government will be sym- pathetic towards this. Because the other places are partially residential areas, no tree could be found any more on Mt Gough and only one at another place on the Peak. Also at Sha Tin there is con- siderable development and though still 17 trees could be discovered this coastal area, adjacent to the big inlet in the east central part of the N. T., is a threatened area for native trees.
Early March 1969 Mr. ALEXANDER sent a parcel of fruits to the Rijksher- barium and these were distributed to various botanical gardens. Germination was at Leyden perfect and provided the material for the figure here reproduced. Fruiting is obviously in Jan.-Febr.
The fruits collected from the single tree in the Hong Kong Botanic Garden, which can therefore not be male, as the herbarium suggested. Hitherto the genus was said to be dioecious. A local study of the flower morphology and biology seems desirable to solve this discrepancy.
6: 220b Add at base:
Doubtful
Ailanthus esquirolii LEVEILLE, Fl. Kouy- Tcheou (1914-15) 404, nomen; Le Monde des Plantes 17 (1915) 23, descr. — The diagnosis is too short for identification; the specimen could not be located and probably does not belong to Ailanthus according to REHDeER (J. Arn. Arb. 14, 1933, 227).
Add above Excluded:
10. ALLANTOS PERMUM
FORMAN, Kew Bull. 19 (1965) 517, t. 1; WEBERLING & LEENHOUTS, Abh. Akad. Wiss. Lit. Mainz, M.-N. KI. 2. 10 (1965) 544; Nootesoom, Adansonia 7 (1967) 161-168. — Cleistanthopsis CAPURON, Adansonia 5 (1965) 213, t. 1. — Fig. 16.
Trees, glabrous. Leaves simple, pen- ninerved, entire; beneath the upper epidermis a hypodermal layer which contains mucilage. Stipules intra-petiolar, soon caducous. Inflorescence a panicle. Flowers bisexual. Sepals 5, imbricate, connate towards the base. Petals 5, free, imbricate. Stamens 10, free, sigmoid- folded in bud. Disk intrastaminal, 10- lobed. Ovary 5-celled with 1 pendent anatropous-epitropous ovule centrally attached near the apex of each cell; style terminal, sigmoid-folded in bud. Fruit a_ septicidal and incompletely loculicidal capsule, the valve-halves becoming slightly twisted after dehiscen- ce, detaching, leaving a central columella; the (often aborted) seeds are attached to an enlarged placenta towards the apex of the columella. In A. multicaule the young fruit is + drupaceous. Seeds shining and waxy, cotyledons planoconvex, radicle pointing upwards; plumule very small.
Distr. 2 spp., 1 in Malesia (Malaya and Borneo) and | in Madagascar (A. multicaule (CAPURON) Noor., /.c. 163).
Notes. This genus has a remarkable distribution and an equally interesting taxonomy. It was only recently in- dependently described by CAPURON who arranged it in /rvingioideae and by For- MAN who removed I/rvingioideae from Simaroubaceae and Ixonanthoideae from Linaceae, joining them as distinct sub- families into a new family /xonanthaceae. Allantospermum he arranged in subfamily Ixonanthoideae.
Independently I came to the same conclusion as CAPURON, viz that A/lan- tospermum belongs to Simaroubaceae- Irvingioideae. The morphological char- acters are in favour of this and this disposition is also sustained on chemo- taxonomical grounds; see NOOTEBOOM, Adansonia 7 (1967) 161-168.
September 1972] Addenda, corrigenda et emendanda 971
Fig. 16. Allantospermum borneense FORMAN. a. Habit, x 2/3, b. flower, 4, c. ditto, floral parts removed
except ovary, <8, d. ditto, lengthwise section, e. fruit, x 2/3, f-e, fruit valves, x 2/3, h. seed, » 2/3, 7.
columella, (7. placental outgrowth, 2. sterile ovule), + x2. — ssp. rostratum NOoTEBOOM, j. Columella, x2, k. young fruit, x 2/3 (a-d S 3364, e-h S 15166, i S 15162, 7 SAN 36068, k SAN 36015).
The controversy about the systematical affinity of A//antospermum has also been tried to solve by means of anatomical and palynological comparison.
Wood Anatomy. Rojo (Adansonia 8, 1968, 73-83) examined its wood anatomy and found this distinctly different from that in Jrvingioideae and possibly more approaching, but still different from Ixonanthes and Ochthocosmus (Ixonan-
thaceae sens. str.). Wood anatomically Irvingioideae constitute a distinct group within Simaroubaceae. For its wood anat- omy Allantospermum could be another group of the same rank as the other distinct wood anatomical types /rvingioi- deae, Kirkioideae, Picramnioideae and Alvaradoideae (cf. WEBBER, Am. J. Bot. 23, 1936, 577-587).
Leaf Anatomy. METCALFE, LESCOT &
FLORA MALESIANA
[ser. I, vol. 6®
LosBreEAU (Adansonia 8, 1968, 337-351) reported the differences between Al/lanto- spermum and Irvingioideae, mainly being the absence of mucilage cavities in the former and their presence in the latter and other Simaroubaceae. Remarkable, however, is the common presence of cristarque cells in both Allantospermum, Klainedoxa and Irvingia. The absence of mucilage canals is a negative character shared with [xonanthaceae sens. str.
Palynology. In her comparison with Irvingia, LOBREAU, /.c., stresses differ- ences more than the undoubted similarities and is apparently unaware that the general Simaroubaceae pollen type is close to that of A/lantospermum.
In comparing with /xonanthaceae she does the opposite and stresses similarities more than the differences and strangely enough does not comment on the striking difference in exine structure between Allantospermum and Ixonanthes. Her conclusions rest therefore only on a comparison with Ochthocosmus. Here she states that in both genera the polar triangle is small, but omits to mention that this is only true for A. multicaule. In A. borneense the polar triangle is large. The endexinous thickenings which form the other support for her thesis are in all types concerned weakly developed and should not carry weight.
LopreAu’s idea that Allantospermum pollen is more similar to /xonanthaceae pollen than to Irvingiaceae would there- fore appear to be weakly founded. The striking similarity of Allantospermum pollen to other Simaroubaceous pollen and the undoubted resemblance to Irvingia pollen support, in my opinion, NOoTEBOOM’s idea to include A//anto- spermum in Simaroubaceae subfam. Ir- vingioideae. — J. MULLER.
Concluding, the evidence is first that the genus stands apart in various aspects. Second that the morphological, chemo- taxonomical and palynological characters all point to inclusion with Irvingioideae in Simaroubaceae, that the balance in leaf anatomy is 50: 50, and that the wood anatomy would be in favour of inclusion in [xonanthaceae sens. str.
1. Allantospermum borneense FORMAN, Kew Bull. 19 (1965) 517, t. 1; NooTe- BooM, Adansonia 7 (1967) 162, pl. 1 f. a—b. — Fig. 16 a-i.
Tree, up to 30m by 50cm @ ; buttresses short to steep, up to 75 cm high, some- times with warts. Stipules only present in leaf-buds, narrowly elliptic-oblong, point- ed, c. 1 cm (ForMAN, /.c.). Leaves glossy above, dull beneath, elliptic with cuneate to broadly rounded base and acuminate,
blunt apex, 7-16 by 3-8 cm; nerves 5-10 pairs, arching upwards and meeting in a not very conspicuous intramarginal vein; veins + transverse, netted; midrib and nerves prominent, veins slightly promi- nent on both surfaces; petiole dark-col- oured, grooved above, c. 1 cm. Panicles lax, up to 7 cm. Bracts early caducous, leaving an obvious scar. Pedicels 7-9 mm. Sepals boat-shaped, reflexed at anthesis, 3-4 by 2-3 mm, caducous when the fruit matures. Petals elliptic to obovate, 4-5 by 21/2-3 mm, membranous, veined, reflexed at anthesis, caducous. Stamens up to 6 mm long, with versatile, c. 1 mm long latrorse-introrse anthers. Disk c. 11/2 mm @ and !/2 mm thick. Ovary 5-lobed, c. 11/2 by 2 mm;; style filiform, c. 3{-4) mm; stigma capitellate. Capsule broadly ellipsoidal, 5-lobed, 21/2-5 cm long, abruptly (up to 3 mm) acuminate at apex. Seeds cylindrical, often slightly curved, 2—2!/5 by 4-6 mm.
Distr. Malesia: Malaya (Penang, Perak, Trengganu), Borneo (Sarawak, Brunei).
Ecol. Obviously not rare in primary lowland rain-forest.
Notes. In my paper (/.c. 164) I wrote: ‘I never saw a lobed structure as depicted in Forman’s paper’. In fact, the placental outgrowths are somewhat lobed, and the lowermost part sometimes tears from the columella (probably due to desiccation in the herbarium). In some cases the seeds leave some material when tearing off from the placenta.
A. multicaule from Madagascar differs from A. borneense in the following char- acters: Tree with many trunks. Stipules narrowly lanceolate, acute, 2—2!/2 mm long. Leaves (narrowly) ovate, with rounded base and long-acuminate, acute apex, 31/2-10 by 11/2-31/2 cm; nerves 3-5 pairs. Fruit c. 2 by 1.7—2 cm. Seeds 10-14 mm long, possessing a_horse- shoe-shaped arillodium directly above the hilum.
ssp. rostratum NOOTEBOOM, ssp. nov. — Fig. 16 j-k.
Inflorescentia praesertim basi race- mosae_ plusminus condensae. Capsula apice longiter acuminato. — Typus: SAN 36068 (L).
Distr. Malesia: Sabah (Sandakan, Lahad Datu and P. Sakar, SAN A 4162, 17478, 26030, 36015, 36068, 37479, 37931, 38777).
Note. Different from the type sub- species in having a shorter, more con- densed_ inflorescence predominantly branched towards the base and up to 2 cm long; acumen on the fruit 6-8 mm long.
September 1972]
Addenda, corrigenda et emendanda 973
4: 281
4:
282
Sonneratiaceae
Sonneratia L. f.
It has appeared that S. caseolaris (L.) ENGL. can grow in fully fresh water without any connection with brackish water. It has been found along the shore of the Sentani Lake, at 75 m altitude, on sandy clay or peaty soil, and also along the Obehfareh R., near Hollandia, in the North of West New Guinea. Other plants occurring along the lake-shore are Crinum asiaticum L. and Pongamia pinnata (L.) Merr. which are otherwise largely con- fined to the sea-shore. This lake contains also several marine animals. It is accepted that it was, even in historical time, a lagoon in open connection with the sea which has become an inland lake by the rapid upheaval to which the northcoast of New Guinea is subjected. It is most curious that the species has been able to maintain itself. It produces the normal aerophores.
Shortly afterwards we received material of the same species from East New Guinea, again from the northcoast, where Mr. J. S. WOMERSLEY collected it at 150 m altitude, in Morobe Distr., 10 miles from Lae, as a 9 m tall tree along the inner margin of a sago swamp, near Oomsis on Wau road (NGF 15307), Aug. 15, 1962.
Though it can be cultivated in fresh water, e.g. at Bogor Botanic Gardens, at 250 m alt., this is the first time it was found in fresh water under purely natural conditions. Obviously, if conditions are favourable and upheaval rapid, marine plants may maintain themselves for some time. The trees in question are not thick enough to have survived since the be- ginning of the upheaval which started some 4-5 centuries ago and must date from later generations. Also at Bogor regeneration takes place under fresh- water conditions. Cf. STEEN. Nova Guinea, Bot. n. 12 (1963) 189.
A distribution map of the genus is published in Pac. Pl. Areas 2 (1966) 248-249,
The species of Sonneratia can mostly be distinguished also in the sterile state, except for S. alba and S. caseolaris. Mr. J. MULLER has found a most easy differentiating character in the leaf tip which bears a thickened pitted gland (? hydathode) peculiar to S. caseolaris; in S. alba instead the leaf margin at and near the ending of the midrib is thinner and finely recurved without such thicken- ed tip. This character is possibly not absolute, but in any case most helpful. Bic lize Add the following new paragraphs:
Anatomy. It is to be regretted that Dr. C. A. STACE in his work on epidermal characters on mangroves (New Phyto- logist 66, 1966, 304-318) omitted to examine Sonneratia. Dr. DinG Hou found that the stomata of Sonneratia
® : a
Fig. 17. Leaf tips of Sonneratia. — a—d. S. caseolaris (L.) ENGL., <3. — e-g. S. alba J.SM., <3. The difference is not completely exclusive and some intermediates occur (a Koorpers 4442, b6 NGF 5026, c BW 3198, d Koorpers 14219, e KARSTEN s.n., f COERT 1456, g BORSSUM WAALKES 272).
974 FLORA MALESIANA [ser. I, vol. 6
Fig. 18. A. Sonneratia ovata, B. S. alba * ovata, C. S. alba, D. S. alba x caseolaris, E. S. caseolaris, all x 1/2 (A MULLER 5, Labuan, B MULLER 8, Brunei, C MULLER s.n., 28-12-63, Labuan, D MULLER 11, Brunei, E MULLER s.n., 30—5—64, Brunei).
agree in structure very well with those of Rhizophoraceae.
He found, however, that they differ in the species, those of 8. alba and SS. ovata being much larger than those in S. caseolaris. Moreover, the two former species have a peculiar type of glands (?) in the leaves, of the size of stomata and equalling the latter in number. They are surrounded by a fairly large number of radially arranged cells. In section there appears to be a large sac-like hollow (cell or intercellular space ?) below them. These glands or whatever they may be are not found in S. caseolaris.
Hybridisation & Chromosomes. J. Mutter & Mrs. S. Y. Hovu-Liu (Blumea 14, 1966, 337-343) have together worked on the identity, the pollen and the chromosomes of pickled material of Sonneratia collected by MULLER and by Mr. J. P. VAN NIEL, in Seria, Brunei. This was initiated through Mr. MULLER who in his pollen studies had found discrep- ancies in the pollen when in Brunei.
It has been shown that in material collected by MULLER S. alba may form hybrids with S. caseolaris and with S. ovata, the characters of which have been tabulated by Mutter (/.c. tab. 1) as to morphological characters, amongst them the fruits (fig. 18). This is the first time that the occurrence of hybrids in man- grove trees was reported (it was recently suggested by BretTeLreR in Acta Bot. Neerl. 18, 1969, 434-444, that Rhizo-
Phora harrisonii is also a hybrid).
Mr. MULLER says that the hybridisa- tion is due to the rather unusual situation that the the three parent species grow side by side in the Brunei River estuary; usually they are ecologically separated, S. alba being the more marine, S. ovata the least marine, while S. caseolaris is ecologically intermediate. The very nar- row mangrove belt along the rather steep banks of the Brunei R. has telescoped the various ecological zones and increased chances of contact. Another factor pos- sibly promoting hybridisation may have been the scarcity of S. ovata which thus stands a large chance of being cross- fertilized with S. alba pollen. This is also suggested by the obvious concentration of alba * ovata hybrids around the single observed locality of S. ovata.
The reduced fertility of the hybrids indicates that the three are good species, but the sterility barrier between S. ovata and S. alba appears lower as compared with that between S. alba and S. caseo- laris.
Mrs. Hou-Liu recorded her experience with the chromosomes which appeared very difficult to count; all species have i1— ile
MuLLteR & VAN STEENIS (North
* Queensl. Natur. 35, 1968, n. 147, 6-8, 1
map) revised the genus Sonneratia for Australia and mapped its localities in Northern Australia and Queensland where only S. alba and S. caseolaris
September 1972] Addenda, corrigenda et emendanda 975
occur, but also indubitable hybrids be- tweenthem,as proved by fruit anomalies and pollen sterility.
Palynology. J. MULLER (Pollen et Spores 11, 1969, 223-298, 15 fig., 15 pl.) has made an extensive examination of pollen of Sonneratia and compared it with that of Duabanga, and various Lythraceae. In S. alba and S. caseolaris he found a distinct intraspecific varia- bility whichis geographical and which is assumed to be genotypic. These were charted. A key was given to the 5 species on pollen characters.
Fossil pollen. J. MULLER (in Cran- well, ed., Ancient Pacific Floras, Univ. Hawaii Press, 1964, 33-42, 2 fig., 1 pl.) studied fossil pollen of Borneo. In NW. Borneo pollen of the S. caseolaris type is oldest and found onwards the transition between Oligocene and Miocene; the S. alba type is younger, from the start of the Pliocene. This is not reflected in the pre- sent range, S. alba is the wider distributed species. S. ovata was not charted; this
4: 283a
4: 288
is said to be closest related to S. alba.
MULLER refers also to papers of Indian palaeobotanists who have recorded Pliocene fossil wood of Sonneratia-like structure and a flower and fruit from the Eocene.
A tentative picture of the evolution of Sonneratia pollen was presented by GERMERAAD, HOPPING & MULLER (Rev. Palaeobot. Polynol. 6, 1968, 189-348) in which is discussed a Lythraceous Tertiary pollen type, Florschuetzia trilobata, which is presumed to be ancestral to Sonneratia, and pointed to the remarkable resem- blance of the latter to the pollen of the Lythraceous genus Lagerstroemia.
Affinity. Pollen types closely com- parable to those of Sonneratia are found in Duabanga and furthermore in various Lythraceae (MULLER, /.c. 1969, 291-292), notably the New World subtribe Dipluso- dontinae (Diplusodon, Lafoénsia) and the Old World subtribe Lagerstroemiineae). Palynologically there would be no objec- tion to merge Sonneratiaceae with Lythraceae.
Sonneratia acida L. f. var. mucronata MiqQ. was reduced here to S. caseolaris (L.) ENGL. MIQUEL’s variety was based on two sheets, one of HORSFIELD and one of REINWARDT. The first is in U and appears to belong to S. alba J. Situ. I could not find at U or L the REINWARDT sheet ticketed by MIQUEL.
Duabanga Bucu.-Ham. — Fig. 19.
Add under the genus the following para- graphs:
Blastogeny. Seedlings have been de- scribed and depicted by Troup (Silv. Ind. Trees 2, 1921, f. 228) and JAYAWEERA &
Howard (Baileya 10, 1962, 8-13, 2 fig.).
Hybridisation & Chromosomes. JAYAWEERA (J. Arn. Arb. 48, 1967, 89- 100, 8 fig., 1 tab.) has found that four large trees growing in the Botanic Garden at Peradeniya and raised from seed received from the Botanic Gardens, Bogor about 1853, are distinctly different from the two known species. He described this as a new species, D. taylorii JAYA- WEERA. Fig. 19 b—b’.
It struck us that in a number of char- acters the new species is intermediate between the continental SE. Asian spe- cies D. grandiflora and the Malesian D. moluccana, which have replacing ranges. At our suggestion Mr. GEESINK has studied the new species and satisfactorily shown that it must be a primary hybrid between the two known species. See his study in Blumea 18 (1970) 453-456, | fig.
Naturally this hybrid could not have been found in the wild, because of the replacing ranges of the parents. However, in the Botanic Gardens at Bogor, where cultivated species are grown in systematic arrangement, the two species grow side by side in the same compartment parcel VII. D, so that hybridisation can easily occur. See TEYSMANN & BINNENDIJK, Cat. Hort. Bog. (1866) 241 and Dakkus, Cat. Bot. Gard. (1927) 119.
A further communication from Dr. JAYAWEERA iS most interesting, viz that the seed of D. x taylorii is for 95% fer- tile, from which it should consequently be deduced that D. grandiflora and D. moluccana are rather subspecies (replac-
976
FLORA MALESIANA
[ser. I, vol. 66
ing races) of one species only than species in their own right.
A further study of chromosomes seems desirable; this has only been observed for D. grandiflora by J. L. THOMAS (Baileya 10, 1962, 13), n=24.
Sphenocleaceae, see Campanulaceae
4: 530b
4: 53la
4: 532a, 5: 564a
42 352, 353
Stylidiaceae
Stylidium inconspicuum SLOOT.
Add to literature: ERICKSON in Specht, Rec. Am.-Austr. Exp. Arnhem Land 3 (1958) 312; Triggerplants (1958) 197-198. Add to Distr.: North Australia, Arnhem Land (Groote Eylandt: Hemple Bay). Note. Mrs. ERICKSON refers this with some doubt to the species which was first described from Java. In our expe- rience Malesian representatives of genera which are characteristic of Australia, especially herbaceous ones, are almost never endemic in Malesia but almost always also present in Australia (as for example in Goodeniaceae, Amaranthaceae, etc.); the existence in Australia of this hitherto endemic species of Java would be no great surprise.
Stylidium javanicum SLoot.
Add to literature: ERICKSON, Trigger- plants (1958) 196.
Note. At some time it was assumed that also this Malesian endemic (Indramaju; Sumba) occurred in NW. Australia, but on closer examination Mrs. ERICKSON and Mr. WiILuLis found the Australian specimens allied but specifically different and have described them as S. fluminense ERICKSON & WILLIS, Vict. Natur. 83 (1966) 108, pl. 2, f. 1-6.
Stylidium pedunculatum R.BR.
Add to literature: ERICKSON, Trigger- plants (1958) 190, pl. 55, f. 5-9. Note. I have reduced to this S. erickso- nae WILLIS, Vict. Natur. 73 (1956) 43; ERICKSON, Triggerplants (1958) 189, pl. 55, f. 10-17. — syn. S. androsaceum O. SCHWARZ in Fedde, Rep. 24(1927) 105, non LINDL., nec DC.
Both Mrs. ERICKSON and Mr. WILLIS believe these two species to be different
49
50a
Thymelaeaceae
and distinguish them as follows:
1. Leaves shortly ciliate on the margins only, terminating into a long hair-like point. Peduncles 2-6 cm. Small petals notched, corolla throat quite naked.
S. pedunculatum
1. Leaves bristly-hairy all over, blunt to acuminate but not hair-pointed. Pe- duncles 6-11 cm. Smaller petals entire, corolla throat with 4 obscure ap- pendavesimn sna S. ericksonae I must remark that the Aru specimens
are almost glabrous and that I am not particularly impressed by the amount of hairs on the leaves. A very hairy specimen of the North. Terr., viz S. T. BLAKE 16371, was by him identified as S. pe- dunculatum, but must then belong to S. ericksonae, but the smaller petals are not notched as far as I can observe. And also I am not impressed by the notching of the smaller petals if I see the plate; it may be that two taxa are involved but the scant material at present available does seem to indicate the desirability of future research on delimitation and status.
Styracaceae
Bruinsmia BOERL. & KoorD.
Change in the description: Leaves almost entire to coarsely crenate. Calyx truncate or 5-lobed. Corolla dull greenish or white. Ovary 3-5(-6)-locular. Fruit oblong to globular. Pericarps pulpy or drupaceous. Seeds l-co per cell. Cf. STEEN. Bot. Jahrb. 86 (1967) 390.
Change last line on Distr. as follows: Distr. Two species, one in Malesia, the other one in Burma, Assam, and Thai- land.
Bruinsmia styracoides BOERL. & KoorD. Add to Distr.: Philippines, Mindanao (PNH 36146). Cf. STEEN. Philip. J. Sc. 88 (1959) 121.
By this discovery also this genus can be removed from the few which cross Ma- kassar Straits without intermediary stations either in the Philippines or Lesser Sunda Islands. Also Bromheadia and Pericopsis are now removed from this list, published by me in Bull. Jard. Bot. Btzg III, 12 (1932) 259.
Gonystyloideae (AiRY SHAW)
Gonystylus TEYSM. & BINN.
Replace the Key to the species by the following:
REVISED KEY TO THE SPECIES
1. Leaves very long, 40-50 cm, venation bullately impressed above, very prominent below. Inflorescence robust, with a very thick rachis, up to 5mm @. Flowers large, with c. 40 disk- lobes and 80 stamens. Style robust, with large stigma, and 4—7 small clavate ‘parastyles’ around
September 1972] Addenda, » corrigenda et emendanda 977
the base. . . 2. G. areolatus
1. Leaves shorter, "3-40(-43) ¢ cm, ‘venation rarely bullately impressed (cf. 26. G. nervosus). Inflo- rescence much less robust. Flowers smaller, disk-lobes and stamens 10-40. Style slender; stigma small; ‘parastyles’ absent (rarely present: cf. 6. G. xylocarpus and 18. G. micranthus), but sometimes represented by 3-4 small rounded humps.
2. Inflorescence-branches elongating almost indefinitely, forming long crowded cicatricose racemes with a few flowers at the apex at the time of flowering. Leaves 10-29 by 5-14 cm. Disk-lobes (where known) 15-16, densely retrorse-setulose. Capsule + lanceolate, 3- or 6-ribbed (Sect. Auxanthus Airy SHAW).
3. Leaves up to 29 cm long, with a dull, ‘shagreened’ surface, the lower surface always bearing minute adpressed hairs, lying parallel to the nerves, often difficult to see (W. Indonesian
Borneo; SW. Sarawak) . . . . 1. G. augescens 3. Leaves up to 20 cm long, with a very smooth ‘and somewhat shining surface, the lower side entirely devoid of minute hairs (NE. Sarawak; Brunei) . . . . . 25. G.lucidulus
2. Inflorescence-branches not elongating, bearing fascicles or short irregular racemes of flowers on short nodulose side-branches. Disk-lobes glabrous or occasionally setulose. Capsule + globose, not or sometimes weakly ribbed.
4. Leaves gradually narrowed at the apex into a relatively long slender acumen, cuneate at the base, up to 20 by 51/2 cm, almost glabrous, drying chestnut-brown. Inflorescence ferrugineous-tomentellous, branches shortly racemiform. Flowers truncate at the base.
11. G. acuminatus
4. Leaves not gradually narrowed into a long acumen, usually rather suddenly narrowed into
a relatively short acumen or cusp, sometimes rounded or even retuse. 5. Pedicels 2-3 cm. Leaves usually large (up to 40 by 15 cm).
6. Midrib distinctly raised above; leaves up to 43 by 12cm, drying olivaceous-brown, almost glabrous; petiole 11/2-2 cm. Inflorescence rather robust, to 17 cm; disk-lobes 30-35, glabrous) “29 207): pcan Oe Hol. er i) 2G. See 22s Gyeostalis
6. Midrib impressed above.
7. Petioles usually or sometimes exceeding 2 cm. 8. Leaves, especially midrib and nerves, ochraceous-tomentellous beneath; petioles and inflorescence-rachis strongly angled, tawny-tomentellous; disk-lobes 40-45. 28. G. spectabilis 8. Leaves glabrous beneath; petioles and inflorescence-axis terete or less strongly angled, cinereous-pubescent; disk-lobes 20-40. 9. Petiole up to 21/2 cm; leaves drying some shade of ochraceous brown. Inflorescence
elongate, to 22 cm : . . 5. G.macrophyllus 9. Petiole 3-4 cm, 4-5 mm @; ': leaves drying pale greenish above, with a narrow purple margin. Inflorescence to 11 cm = RUB Ae NEF ee... 27. Ganopilis
7. Petioles not exceeding 2 cm. 10. Leaves glabrous or almost so. 11. Leaves drying a dark purplish-leaden colour; base of calyx narrowing gradually
into pedicel; disk-lobes 20-30 . . . b pasher oer ure calophyllus 11. Leaves drying a light brownish colour; base of calyx broadly truncate, passing abruptly into the pedicel; disk-lobesc.50 . . . . . . 20. G. calophylloides
10. Leaves thinly pilose or pubescent beneath, at least on the midrib. 12. Leaves drying a light green with a narrow dark brown margin; disk-lobes c. 40. 4. G. reticulatus 12. Leaves greyish-green above, ochraceous-brown beneath; disk-lobes c. 30. 21. G. consanguincus 5. Pedicels !/2-2 cm. Leaves small or medium.
13. Calyx-segments strongly reflexed or revolute at anthesis; disk-lobes 7-12. Inflorescence usually considerably branched. Flowers small. Leaves small, up to 13 by 5 cm, dis- tinctly shagreened.
14. Disk-lobes tomentellous throughout and setulose within. Leaves elliptic or almost rhomboid, glabrous or almost so, usually ochraceous when dry; nerves rather steeply ascending : . . 14. G. forbesii
14. Disk-lobes glabrous. “Leaves elliptic to oblong but 1 never subrhomboid, drying brown- ish or greenish; nerves rather widely spreading.
15. Young parts and inflorescence densely fulvo-velutinous. Leaves usually -- pubescent
below, glossy above (even when dry), rather variable in shape, often cuneate at base.
15. G. velutinus
15. Young parts and inflorescence tomentellous. Leaves glabrous or almost so below, dull above when dry, regularly elliptic-oblong, mostly rounded at base.
16. G. maingayi
13. Calyx-segments not or scarcely reflexed at anthesis.
978 FLORA MALESIANA [ser. I, vol. 6%
16. Disk-lobes 10-12, very slender, pustulate toward apex. Flowers very small (4-6 mm). 3 filiform, clavate parastyles present. Leaves drying a dull purplish-leaden colour, up: to5iby Gicmiy i. at . bee fos solseiGeimicranthus
16. Disk-lobes 13-40. Flowers larger.
17. Leaves drying a purplish-leaden colour; midrib flat or slightly raised above. 12. G. confusus 17. Leaves not drying purplish-leaden. 18. Disk-lobes retrorse-setulose within.
19. Leaves drying greyish-green above with a narrow brown border, pinkish-brown below, chartaceous; nervation lax, the primary nerves being distinctly differentiated from the secondaries: disk=lobest20=22.5) (ae tee . . 9. G. keithii
19. Leaves drying brownish or ochraceous, or greenish without a brown border, + coriaceous; venation dense, without distinctly differentiated primary nerves.
20. Leaves drying chestnut-brown throughout; disk-lobes 25-30. 10. G. brunnescens
20. Leaves drying ochraceous or greenish; disk-lobes 35-40 . . 23. G. decipiens
18. Disk-lobes glabrous.
21. Midrib distinctly prominent above. Sepals narrowly triangular-lanceolate. Leaves yellow-ochraceous when dry : . . . 7. G. stenosepalus
21. Midrib flat or channelled above. Sepals ovate- deltoid.
22. Leaves small, 4-15 by 2-7 cm, coriaceous, often + conduplicate, drying dull purplish-red below and chestnut above; nervation relatively inconspicuous. 19. G. bancanus
22. Leaves various, but not conduplicate, nor drying as above; nervation more
conspicuous.
23. Indumentum of inflorescence tomentose or tomentellous, 7.e. with short spreading hairs, usually + fulvous. 24. Leaf-surface not dull and ‘shagreened’ (though not shining); pedicels 11/2-21/2 cm; calyx 5-8 mm long; disk-lobes c.30 . . . . . 21. S. consanguineus 24. Leaf-surface dull and ‘shagreened’, slightly glaucescent below; pedicels 0.8—2 cm; calyx 5-6 mm long; disk-lobes 20-30.
25. Leaves long and narrow, up to 27 by 7 cm, at least 4 times as long as broad, glabrous; inflorescence up to 28 cmlong . . . . . 24. G.glaucescens
25. Leaves less elongate, up to 17 by 81/2 cm, only 2-3 times as long as broad, often tomentellous below; inflorescence up to 12 cm long. 13. G. affinis
23. Indumentum of inflorescence thinly adpressedly grey-pubescent. 26. Leaves large, glabrous, up to 34 by 12 cm.
27. Leaves coriaceous, drying pale green above with a narrow brown edge (as in 9. G. keithii and 4. G. reticulatus); nerves not bullately impressed; petiole robust, terete, up to4cmlong . . ohn | 2iatGenobilis
27. Leaves chartaceous, not drying pale green with a brown edge; primary nerves + bullately impressed; petiole much less robust, up to 2 cm long.
26. G. nervosus 26. Leaves small or medium, 9-24 by 3-71/2 cm, glabrous or shortly adpressed- pubescent below.
28. Leaves rigidly coriaceous, broadly elliptic, up to 17 by 9 cm; inflorescence robust, up to 18 cm; disk-lobes 35-40; parastyles 2—3; fruit massive, pericarp up to 2 cm thick, almost StOUYs sane . . 6. G.xylocarpus
28. Leaves thinly to firmly chartaceous; inflorescence rather slender; disk-lobes 25-30; parastyles absent; fruit with moderately thick and woody pericarp.
29. Leaves 12-24 by 41/2-71/2 cm; inflorescence 10-20 cm. 8. G. borneénsis 29. Leaves 9-11 by 3-4 cm; inflorescence 2-3 cm, few-flowered. 17. G. pendulus
4: 35Sa Gonystylus xylocarpus Airy SHAW. parastyles 2-3, flattened-clavate, 11/2 Add to literature: Fl. Mal. I, 4 (1953) 355; mm. Fruit dehiscing by 4 valves. Kew Bull. 17 (1964) 450. Add to Ecol.: Dominant tree in kerangas Add to description: Tree up to 36 m. on flat ridge, 30-150 m altitude. FI. Inflorescence robust, up to 18 cm long. March, fr. March, June, July. Pedicels up to 1.8 cm, grey-sericeous. Add to Vern.: Ramin batu, garu mélitan, Calyx 7 mm long, 7-8 mm @, thickened Sarawak. and truncate at the base, very shortly Replace the Note by: Noteworthy as adpressed-sericeous; segments + deltoid, one of the very few species of the genus in somewhat obtuse and very shortly re- which ‘parastyles’ are developed. curved at the apex. Disk-lobes 35-40, 4: 359a Gonystylus maingayi Hook. f. narrowly subulate, 3-4 mm long, glab- Add to literature: Airy SHAW, FI. Mal.
rous, epustulate. Style pilose below; I, 4 (1953) 359; Kew Bull. 17 (1964) 456.
September 1972]
4: 359b
4: 36la
4: 361b
Add to Distr.: Borneo (Sarawak, Brunei, Sabah).
Add to Ecol.: Common dominant tree in primary peat-swamps, up to 15 m. F7. Febr., Oct.-Nov., fr. Febr., April.
Add: Uses. Wood used for planks and boards for domestic buildings (Sabah). Add to Vern.: Ramin batu, Sarawak, bidaru, Sabah (from Brunei informant). Add to Notes: This species apparently occupies a similar peat-swamp habitat to 19. G. bancanus, but is evidently far more local. The fruit develops into a curiously asymmetrical, bean-like form, with one of the three valves almost abortive. Gonystylus micranthus AiRY SHAW.
Add to literature: Fl. Mal. I, 4 (1953) 361; Kew Bull. 17 (1964) 457.
Add to description: Tree to 15-20 m. Leaves with drip-tip up to 2!/2 cm. Calyx ovoid, up to 6 mm. Disk-lobes 10-12, very slender, pustulate towards apex, glabrous. Style surrounded at the base by 3 conspicuous, filiform, clavate- capitate parastyles, 11/2 mm long.
Add to Distr.: S. Indonesian Borneo. Add to Ecol.: Primary lowland forest or marshy forest, on sandy loam soil, 20-90 m. Fl. Febr., March, July, Aug.
Add to Notes: The fruit of this very distinct species is still a desideratum. After 19. Gonystylus bancanus (MiIQ.) Kurz add the following species:
20. Gonystylus calophylloides Airy SHAW, Kew Bull. 17 (1964) 448.
Small tree to 6 m. Leaves oblong or slightly oblanceolate-oblong, 30-37 by 11-12 cm, slightly cordate at base, round- ed and abruptly shortly caudate at apex (cauda 1.2-1.8 cm by 2-4 mm), chartaceo- coriaceous, glabrous, pale ochraceous- brownish (greenish tinged above) when dry; midrib moderately robust, very prominent and subcylindric beneath, flat or scarcely prominulous above; nerves very slender, c. 35 pairs, distinctly prominulous on both surfaces; petiole 1-11/2 by 4 mm, very rugose. Jnflores- cence very abbreviated, 21/2 cm, few- flowered, sericeous. Pedicels 1.2—2.5 cm, densely ochraceous-sericeous. Calyx broadly ovoid, truncate at base, 7-8 cm @: sepals broadly deltoid, 7-8 by 4-6 mm, slightly recurved and sometimes with a slight thickening at the apex, dorsally ochraceous-sericeous. Disk-lobes c. 50, robust, laterally flattened, 6 mm long, glabrous, epustulate. Style elongate, long-hairy, surrounded at the base by 4-5 small obtuse ‘parastyles’. Fruit subglob- ose, 31/2 cm @%, apparently 5-valved, rugulose. obscurely puberulous, subtend- ed by the 10 mm long sepals and borne on a striate pedicel 2!/2 cm by 3-4 mm.
Addenda, corrigenda et emendanda 979
Seed apparently solitary, large.
Distr. Malesia: Borneo (NE. Sara- wak).
Ecol. Banks of rocky stream at 210 m. Fi. fr. July.
Vern. Ramin, Sarawak.
Note. Closely related to 3. G. calophyl- lus of SW. Sarawak, differing in the pale ochraceous-brown colour on drying, and in the broadly ovoid shape of the calyx, the truncate base of which passes abrupt- ly into the pedicel.
21. Gonystylus consanguineus Airy SHAW, Kew Bull. 17 (1964) 454.
Tree, 16-40 m. Leaves elliptic to oblong, 10-25 by 5-9 cm, broadly cuneate or rarely rounded at base, rather abruptly caudate-acuminate at apex (cauda 1/2-21/2 cm), chartaceous to coriaceous, mostly grey-green above when dry, ochraceous-brown beneath, upper surface dull but scarcely ‘shagreen- ed’ and glabrous or very sparsely pilose towards the base, lower surface also dull, very shortly tomentellous or spreading- puberulous, at least along the midrib; midrib moderate, cylindric-prominent beneath, not deeply impressed above; primary nerves 15—25 pairs, together with the numerous minor nerves conspicuous- ly prominulous, especially beneath; petiole 7-12 by 1-2 mm, shortly fulvo- tomentellous or rarely glabrescent. Jn- florescence 10-13 cm, sparingly and very shortly branched, sparsely fulvo-to- mentellous. Pedicels 11/2—21/2 cm, dense- ly fulvo-tomentellous. Sepals 5-8 by 3-4 mm, externally densely fulvo- tomentellous. Disk-lobes c. 30, glabrous, epustulate, irregularly connate. Style glabrous.
Distr. Malesia: Borneo (W. & E. Indonesian Borneo, Sarawak, Sabah).
Ecol. Very varied: primary forest or disturbed forest on black or brown soil, once over sandstone, up to 60 m (Sabah); primary lowland Dipterocarp forest (Sarawak); loam soil and coral limestone, or sand and limestone, at 50-400 m (E. Borneo); old secondary forest, in bog on clay submerged during Westmonsoon, at 40 m (W. Borneo; identity not quite certain). F/. June, Nov., fr. July, Sept., Dec.
Vern. Bidaru, Sabah, ramin bukit, Sarawak, ngalin, njoelir (nyulir), tempé- éng, E. Borneo, mélingkat pépah, W. Borneo (?).
Note. This species resembles 8. G. borneénsis in its conspicuous raised parallel nervation and large flowers, and 13. G. affinis RADLK. in its more stiffly coriaceous leaves and subtomentellous indumentum, but the inflorescence and
FLORA MALESIANA
[ser. I, vol. 68
flowers are larger than in either. It seems to be remarkably indifferent in its ecology.
22. Gonystylus costalis Airy SHAW, Kew Bull. 23 (1969) 269.
Small tree, 41/2 m. Leaves large, oblong- elliptic, 25-421/2 by 8-12 cm, base shortly cuneate or subrotundate, apex subrotun- date and shertly (11/2 cm) apiculate- caudate, margin conspicuously revolute, coriaceous, glabrous or with a few lax hairs beneath near the base of the midrib, brownish when dry, or greenish above, dull, under a lens densely minutely puncticulate above, very minutely granu- lose below; midrib moderately robust, prominent and 2—2!/2 mm thick beneath, broadly (2 mm) but shallowly elevate above and bordered on each side by a groove which is often indistinct or obso- lete on account of the oblique nerve-bases traversing it; principal nerves c. 30 pairs, scarcely distinguishable from the minor ones, prominulous on both surfaces, occasionally subbullately impressed, con- spicuously anastomosing near the margin and conspicuously abruptly decurving below into the midrib; petiole 11/2-2 cm by 4-6 mm, rugulose. /nflorescence robust up to 17 cm, with an occasional subbasal branch to 7 cm, rachis compressed, very shortly fulvous-tomentellous. Pedicels up to 2.3 cm, tomentellous. Calyx 7-9 mm long, externally sericeous. Disk-lobes 30-35, subulate, glabrous, epustulate. Style 7-8 mm, glabrous.
Distr. Malesia: Borneo (Central Sarawak).
Ecol. Presumably evergreen rain-for- est, on ridge on sandy clay soil at 210 m. Fl. Oct.
Note. Readily distinguished from all other species except 7. G. stenosepalus by the raised midrib on the upper surface of the leaves. From that species it differs in the much greater size of the leaves, the broader elevation of the midrib, the denser arrangement and _ occasionally bullate impression of the nerves, which are more or less decurrent on to the midrib, and by the much less conspicuous glandular puncticulation.
23. Gonystylus decipiens AIRY SHAW, Kew Bull. 17 (1964) 454.
Tree, 25-30 m. Leaves elliptic or almost oblong or oblanceolate, 15-20 by 6—7!/2 cm, cuneate or somewhat rounded at base, shortly caudate-acuminate at apex, cauda obtuse, chartaceo-coriaceous, en- tirely glabrous except for the midrib, not or scarcely shining, greenish or fuscous above when dry, subochraceous below; midrib moderate, prominent and rather long-adpressed-pilose below, narrowly
impressed and glabrous above; nerves densely parallel and prominulous, the primaries hardly distinct from the re- mainder; petiole 11-14 cm by 2-21/2 mm, striate, sparsely long-pilose or glabres- cent. Inflorescence 10-15 cm, rather many- flowered, with numerous abbreviated branches, very shortly grey-subsericeous. Pedicels 8-12 mm, sericeous. Calyx 5-7 mm long and wide, sepals subobtuse. Disk-lobes 35-40, subulate, 3-4 mm, sparsely retrorse-setulose, epustulate. Style glabrous, surrounded at the base by 4-5 very small subglobose parastyles.
Distr. Malesia: Borneo (Central Sa- rawak).
Ecol. Primary rain-forest on sand- stone, below 500 m. F/. Sept.
Notes. Among the rather few species with setulose disk-lobes, this is perhaps nearest to 9. G. keithii, from which it is at once distinguished by the dense promi- nent venation with scarcely differentiated primary nerves, and by the quite different colour assumed by the leaves on drying, especially lacking the narrow brown margin of G. keithii. From the somewhat similar 8. G. borneénsis and 21. G. consanguineus it differs in the setulose disk-lobes and adpressedly long-pilose, rather than shortly tomentellous, midrib beneath.
24. Gonystylus glaucescens Airy SHAW, Kew Bull. 17 (1964) 451.
Tree, 10 m; branchlets conspicuously elevate-lenticellate. Leaves oblong-oblan- ceolate, 21-27 by 5-7 cm, + rounded- cuneate at base, narrowed and shortly (5-10 mm) acuminate-caudate at apex (acumen obtuse), margin distinctly thick- ened, rigidly coriaceous, quite glabrous, greenish and dull-shagreened above when dry, or scarcely shining, greyish-ochra- ceous beneath and very dull, as though glaucescent; midrib moderately robust, prominent and subcylindric beneath, slightly impressed or flat above; nerves numerous, widely spreading, the smaller veins clearly reticulate above but lax and rather indistinct below; petiole 1—-11/2 cm by 3-4 mm, rugose, glabrous. Jnflores- cence terminal, elongate, robust, 28 cm long, rachis 3-5 mm thick, conspicuously fuscous-lenticellate, in the fruiting stage finely fulvous-puberulous, branches ab- breviated, nodose, densely fulvous-pu- bescent. Flowers unknown, but (from the fruits) sepals 5-6 by 2-4 mm, subobtuse, ochraceous-sericeous; disk-lobes 25-30, glabrous, epustulate. Fruit (immature) obtusely tetragonous-globose, 3!/2-4 cm @, clearly 4-valved, pedicel 11/2-2 cm by 3-4 mm, fulvous-puberulous.
Distr. Malesia: E. Indonesian Borneo.
Septem ber 1972]
Addenda, corrigenda et emendanda 981
Ecol. Sandstone ridge at 400 m alt. Fr. Sept.
Note. The long, narrow, _ stiffly coriaceous leaves, more than 4 times as long as broad, with their dull, slightly shagreened, almost glaucescent surface, and the robust, elongate inflorescence, are highly distinctive.
25. Gonystylus (§ Auxanthus) lucidulus Airy SHAW, Kew Bull. 17 (1964) 447.
Tree to 36 m. Leaves oblong, elliptic- oblong or slightly lanceolate-oblong, 10-20 by 5—7%/4 cm, base rounded (rarely slightly cuneate or very slightly cordate), apex narrowed or rounded, shortly abruptly caudate (cauda 5-15 mm), subacute or obtuse, moderately coria- ceous, greenish above when dry, grey- brown to purplish beneath, very smooth on both surfaces (except for the nerves) and manifestly somewhat shining, quite glab- rous, under a lens + clearly dark- puncticulate; midrib moderately robust, almost cylindric beneath, deeply impres- sed above; nerves slender, spreading, sharply prominulous on both surfaces; petiole 1.2-1.8 cm by 2-3 mm, grooved above, sparsely fulvous-puberulous. /n- florescence very similar to that of G. augescens, but the rachis often thicker and more nodose, and the bracts much broader, suborbicular, 7-8 mm broad, dorsally carinate. Immature fruit asym- metrically lanceolate-ovoid, 3.5 by 1.7 by 1.5 cm, obliquely acuminate or sub- rostrate, 3-ridged, the ridges often marked with a slender groove, and alternating with 3 humps between the ridges towards the apex, very finely + stellate-ochra- ceous-puberulous, subtended by the persistent 5-6 mm long densely sericeous sepals and borne ona5 mm long densely grey-sericeous pedicel.
Distr. Malesia: Borneo (NE. Sara- wak; Brunei).
Ecol. Primary lowland Dipterocarp forest, 30-270 m, on yellow sandy clay. Fl. April-June, young fr. June.
Vern. Ramin, Sarawak.
Note. Closely related to 1. G. augescens, of SW. Sarawak, and adjacent W. Indonesian Borneo, but differing in the smaller and narrower, smooth and shining leaves, and in the total lack of the minute adpressed hairs, lying parallel to the nerves on the undersurface, which are always found (by careful searching!) in that species.
26. Gonystylus nervosus Airy SHAW, Kew Bull. 17 (1964) 452.
Small tree, 4-5 m, Leaves elongate- oblanceolate, rarely oblong, 17-34 by 5-9 cm, cuneate at base, narrowed to
subrotundate at apex and abruptly narrowly caudate, cauda 1-4 cm, acute, chartaceous, glabrous, not shining, ob- scurely grey-brown above when dry, similar or subcastaneous below; midrib relatively slender, prominent below, impressed above; primary nerves 25-30 pairs, strictly parallel, sharply raised beneath, subbullately impressed above, conspicuously arcuate-anastomosing to- ward the margin; petiole 1-11/2 cm by 2-4 mm, fulvous-pubescent at first, finally glabrescent. /nflorescence 2-4 cm, few-flowered, grey-sericeous. Flowers on- ly known in bud stage: disk-lobes about 30, glabrous, epustulate; style glabrous. Fruit ellipsoid, 31/2 cm, 3-valved, subtend- ed by 5-6 mm long sepals and borne on a 2 cm long peduncle. Seeds 2, flattened hemi-ellipsoid, 2.2 by 1.6 cm, shining, light chestnut.
Distr. Malesia: Borneo (SW. & NE. Sarawak).
Ecol. In primary rain-forest, appar- ently always on or near limestone hills, 60-270 m. F/. June, July, fr. June, August.
Note. The elongate leaves, with their somewhat bullately impressed main nerves, recall those of 2. G. areolatus, but G. nervosus is probably most closely related to 8. G. borneénsis. The latter species, however, seems almost always to occur on more or less acid soils (basalt, kerangas, efc.); I have only seen one collection from limestone.
27. Gonystylus nobilis Airy SHAW, Kew Bull. 23 (1969) 271.
Tree to 24 m; branches robust, almost 1 cm @. Leaves large, elliptic to sub- oblong, 23-34 by 9-12 cm, base broadly cuneate to rounded, apex narrowed to somewhat rounded, very shortly, broadly and not abruptly acuminate (acumen acute, | cm), firmly coriaceous, quite glabrous, dull and grey-green above when dry, with a very narrow purplish margin, smooth beneath, not shining, distinctly pale when dry; midrib robust, subcy- lindrically prominent beneath, deeply and narrowly impressed above, primary nerves c. 20 pairs, lax, conspicuously anastomosing near the margin, obscurely prominulous above, sharply prominent beneath; petiole very robust, 3-4 cm by 4-5 mm, terete, sparsely adpressed- pubescent. Inflorescence 11 cm, simple, rachis terete, 3-4 mm @, adpressed- pubescent, flower-nodules very short. Fruit transversely subellipsoid, 61/2 by 51/2 cm, conspicuously obtusely 4-ridged, strongly rugulose.
Distr. Malesia: Borneo (W. Central Sarawak).
Ecol. Primary lowland Dipterocarp
982
FLORA MALESIANA
[ser. I, vol. 68
6:
forest, altitude and soil unknown.
Vern. Ramin. Sarawak.
Note. Notable for its large glabrous leaves and especially for the remarkably long, robust, terete petiole. The nervation and the colouring of the leaves on drying somewhat recalls 9. G. keithii on a larger scale.
28. Gonystylus spectabilis AIRY SHAW, Kew Bull. 23 (1969) 269.
Tree, 24 m; branches robust, ridged, tomentellous. Leaves broadly elliptic- oblong, up to 32 by 14!/2 cm, slightly but distinctly cordate (rarely rounded) at base, rounded and shortly (to 2 cm) abruptly cuspidate-caudate, chartaceous to thinly coriaceous, glabrous above, finely velutinous beneath with short white hairs, bright green (especially be- low) when dry, slightly shining above; midrib strong, very prominent below, 3-4 mm thick, densely fulvo-tomen- tellous, narrowly impressed above; main nerves (scarcely distinguishable from the minor ones) 15-20 pairs, slender, sharply prominulous below, scarcely so above; minor nerves very numerous; petiole robust, 11/2-21/2 cm by 45 mm, longitudinally rugose, often almost tetra- gonous, densely tomentellous. /nflores- cence robust, terminal, to 22 cm, little branched; rachis compressed, up to 6 mm thick, strongly grooved, fulvo-tomentel- lous, branches to 7 cm, widely spreading. Pedicels to 2!/2 cm by 21/2 mm, tomentel- lous. Calyx 1-1.2 cm, truncate at base, externally tomentellous. Disk-lobes 40- 45, narrowly subulate, glabrous, epustul- ate. Style 8-9 mm, glabrous.
Distr. Malesia: Borneo Sarawak).
Ecol. Presumably rain-forest, on ridge at 195 m. Fi. Oct.
Note. Related to 4. G. reticulatus, 3. G. calophyllus and 20. G. calophylloides, but differing in its dense fulvous-tomen- tellous indumentum and in its petioles sometimes reaching 21/2 cm; further from G. reticulatus in the absence of a narrow brown border to the leaves when dry, and from G. calophyllus in the 40-45 rather than 20-30 disk-lobes.
(Central
Aquilarioideae & Thymelaeoideae (DiING Hov)
9a Aquilaria malaccensis LAMK.
23
Add to synonymy: A. agallocha Roxs.: cf. DinG Hou, Blumea 12 (1964) 286. Enkleia GRirF. Add to references: NEVLING, J. Arn. Arb. 42 (1961) 373-396, 9 fig., map.
He gave an extensive, detailed treat- ment of the morphology and anatomy, including a systematical revision. He
27b
35a
added one new species from SE. Asia. See the remark below on the generic difference with Linostoma. E. malaccensis seems to be restricted to Malesia. Linostoma WALL. ex ENDL. Add to references: NEVLING, J. Arn. Arb. 42 (1961) 295-320. NEVLING does not distinguish sections within the genus to which I agree. Furthermore he remarked correctly that the generic difference between Linostoma and Enkleia tends to disappear as the main differential character : stamens in One or two series respectively is rather breaking down in one species which is almost intermediate in this respect. Linostoma pauciflorum GRIFF. Add to Distr.: NEVLING, who first hesitated to accept Psilaea dalbergioides Mia. from Sumatra as a synonym (1/.c.), later traced its type in CAL and found (J. Arn. Arb. 43, 1962, 221) it correctly reduced by DinG Hou. Thus there are 2 collections from Sumatra: mainland, pr. Sibolga and Simalur I. Linostoma longiflorum HAL. f. is by NEVLING (J. Arn. Arb. 42, 1961, 313) reduced to L. pauciflorum; he showed that the size and shape of the bracts is variable, but he found no such graded variability in the flower length. Line 14 from top, replace ‘375’ by: 357.
Trapaceae
The family name Hydrocaryaceae must give way to the now conserved name Trapaceae.
Typhaceae
Typha L.
Miss B. G. BricGcs and Dr. L. A. S. JOHNSON (Contr. N.S.W. Nat. Herb. 4, 1968, 57-69, 2 fig., 1 tab.) have published a detail account of Typha in Australia in which they also examined Malesian material. They maintain that what has been called in Malesia and Australia 7. angustifolia L. sens. lat. is not that species which to them is confined to the northern hemisphere; however, they provide for this no further arguments. The Malesian and Australian material they bring to two species, 7. domingensis PERS. 1807 and 7. orientalis PR. 1852 (type from the Philippines). On the basis of the Austra- lian material it was established that two taxa are concerned, which differ in chromosome number, JT. domingensis 2n=—30 and T. orientalis 2n=60. They differ also in minute other characters. The authors state that they are often found in mixed stands but do not hybrid-
September 1972]
Addenda, corrigenda et emendanda
983
ize; this is not so astonishing, as hybrids
between 7. angustifolia and T. latifolia
(2n also 30) are in Europe also very rare. The two taxa, which both occur in
Malesia, can be distinguished by some
minute morphological details. To my
view they are merely cyto-subspecies.
For those who want to do further work
on this in Malesia I have copied the key
given:
1. Bracts in the 2 inflorescence numerous, broadly spathulate (usually 4—8 cells across the lamina). Stigmas linear. Mature 2 spikes !/2-2 cm @, length 6-20(—30) times @, cinnamon-brown (due to the numerous pale ends of the bracts interspersed among the darker stigmas and the carpodia). 3 and & spikes separated by (4/2-)2-5 cm. Sheaths of the upper leaves not auric- ulate or only the uppermost 1-2
Mature & spikes 1-3 cm @, length 5—10(-18) times @, chestnut-brown (due to the great predominance of the brown stigmas at the surface). J and & spikes contiguous or separated by up to 2(-6) cm. Sheaths of the 2-4 uppermost leaves usually distinctly auriculate T. orientalis
4: 243a Typha angustifolia L.
4: 244b
4:
117
Add to synonymy: 7. angustata BoRY & CHAUBARD, Exp. Sc. Morée 2, | (1832) 3385; HENDERSON, Mal. Wild Flow., Monoc. (1954) 211 f. 126.
Add to Distr.: Malay Peninsula: Kuala Selangor; Kedah: Kuala Muda; Lang- kawi: Tasek Bayang Bunting.
Add to Vern.: Bulrush, E, banat, M. Note. By a curious oversight this was omitted from RIDLEY’s Materials and his Flora. Near Kuala Selangor it is found in ditches along the roadside in association with Acrostichum aureum and Pluchea, both typical for brackish water.
Umbelliferae
Trachymene RUDGE.
Fig. 20. Trachymene tripartita HOOGL. a. Habit, 1, b. leaf-blade, < 10, c. leaf-sheath, = 10, d. involucral bract, x10, e. & f. flower with only one petal still present, = 15, g. fruit, x 15 (a-g HOOGLAND & PULLEN 5965).
984
4: 118
FLORA MALESIANA
Many new collections of the genus have come in from the Papuan highlands and a revised account has become most desir- able. A newly described species is to be added:
Trachymene tripartita HooGL. Blumea Suppl. 4 (1958) 231. — Fig. 20. Glabrous perennial. Stem little branch- ed, bearing rosettes. Leaves: sheath 4-8 by 2 mm, attenuate into petiole; petiole 7-30 mm, exceeding the blade; blade 3-partite, 5-8 by 5-8 mm, broad-cuneate at the base, lobes almost equal, obtuse, mucronate. Umbel 1 from a rosette, peduncle + as long as the leaves or + shorter, in fruit twice as long, striate or subsulcate; involucral bracts 5—7, linear- lanceolate, 3-4 mm by 3/4 mm, obtuse, subappressed. Flowers 5—10, in 1(—2) rows.
[ser. I, vol. 68
5: 556b
Pedicels in fl. 1-2 mm, in fr. to 7 mm, incurved. Calyx lobes minute. Petals rounded, c. 1 by 0.8 mm. Filaments linear, widened to base, 0.4 mm; anther- cells 0.3 by 0.15 mm. Ovary 0.7 by 1 mm; style c. 0.4 mm. Mericarps 2 by 1.7 mm, equal; carpophore undivided, 4-apicul- ate, sulcate.
Distr. Malesia: East New Guinea (Mt Wilhelm), open rocky slopes and tussock grasslands, 3600-3750 m.
Note. Closest related to 7. novo- guineensis (DOMIN) Buw., clearly char- acterized by the 3-partite leaves with undivided lobes.
Daucus glochidiatus (LABILL.) FISCHER. Add to Distr.: East New Guinea: Foramburo, 2500 m, 28 Oct. 1960, E. BORGMANN 340.
INDEX TO SCIENTIFIC PLANT NAMES
compiled by
M. J. VAN STEENIS- KRUSEMAN
Families and higher taxa have been entered under their name. Names of families which have been revised in volumes 4, 5, and 6 have been entered and are printed in bold type, so that as far as this is concerned this index is complete for all preceding volumes as well. Suprageneric epithets have been entered under the family name to which they belong preceded by the indication of their rank (subfamilies, tribes, efc.). Infrageneric epithets have been entered immediately under the generic name to which they belong preceded by the indication of their rank (subgenera, sections, series, e/c.). Infraspecific epithets have been entered under the specific name to which they belong preceded by the indication of their rank (subspecies, variety, forma, efc.). New names and new combinations have been printed in bold type, synonyms in italics. ‘Map’ printed behind a page number denotes that a map of the concerned taxon is present on that
page.
An asterisk behind a page number denotes the presence of a figure of the concerned taxon.
Page numbers in bold type denote main treatment.
Some minor printing errors in plant names have been corrected.
Of synonyms with a double authority, the latter has not always been cited in full. See for example under Dortmann(i)a Adans. which was accepted by O. Kuntze to replace Lobelia. The full authority can
easily be derived from the text.
Acer garrettii Craib 915 laurinum Hassk. 915 Aceraceae 4: 3—4, 592; 6: 915 Actinidiaceae s. str. 4: 37-39 Adenanthera triphysa Dennst. 219 Adenoplea Radlk. 295, 296, 337, 953 madagascariensis Eastw. 340 Adenoplusia Radlk. 295, 296, 337, 953 Adnaria /anceolata (Bl.) O.K. 761 Aegiphila viburnifolia Juss. 248 Aegle marmelos (L.) Correa 69 Aeschrion Velloso 212 excelsa (Sw.) O.K. 193 Aétoxylon 3, 5, 6 Agallochum Rumph. ex Lamk 6 coinamense Rumph. 9 malaccense O.K. 9 malaicense Rumph. 9 secundarium Rumph. 9 Agapetes D.Don 469, 471, 473, 474, 746, 878, 879 map sect. Pseudagapetes Airy Shaw 878
subg. Dimorphanthera (F.v.M.) Drude 885 acuminatissima (Miq.) Niedenzu 761 alberti-eduardi Sleum. 879, 882*, 883 amblyornidis Becc. 892 beccariana Koord. 904 brassii Sleum. 879, 881, 882* brevicuspis Sleum. 884 carrii Sleum. 879, 881, 882*
coriacea (Bl.) G.Don 828 costata C.H.Wright 879,882*, 883 cuneifolia (Bl.) G.Don 874 elliptica (Bl.) G.Don 873 filicicola Sleum. 883 floribunda (Bl.) G.Don 828, 872 forbesii F.v.M. 904 griffithii (non Clarke in Hook.) K. & G. 753 helenae (F.v.M.) F.v.M. ex Sleum. 878, 879, 882* lanceolata (Bl\.) Niedenzu 761 laurifolia (Bl.) G.Don 872 leptantha (Miq.) Niedenzu 764 lucida (B\.) G.Don 814 meliphagidum Becc. 906 micrantha Ridl. 753 microphylla Jungh. 814 moorhousiana Lane-Poole 892 moorhousiana F.v.M. 893 myrtoidea = myrtoides 812 myrtoides (Bl.) G.Don 812 myzomelae Becc. 913 obtusata Sleum. 884 parviflora Ridl. 753 perakensis Ridl. 753 polyantha (‘Miq.’) Niedenzu 872 prainiana Koord. 911 pubescens Ridl. 753 rubrocalyx Sleum. 879, 881 var. pilicalyx Sleum. 881 var. rubrocalyx 881 sclerophylla 879, 880, 882* scortechinii (K. & G.) Sleum. 878, 879, 880*, 882*
stenantha (Schltr) Sleum. 879, 884* varingiaefolia (Bl.) G.Don 815 viridiflora (Schltr) Sleum. 879, 883 vitis-idaea Sleum. 879, 881, 882* vonroemeri Koord. 911 vulgaris Jungh. 815 wrayi Ridl. 753 wrightiana Koord. 914 Agapetes (non D.Don) Koord. 885 Agelaea borneensis (Hook. f.) Merr. 933 insignis (Schellenb.) Leenh. 933 macrophylla (Zoll.) Leenh. 933 trinervis (Llanos) Merr. 933 Aglaia sapindina (F.v.M.) Harms 237 Aglaiopsis glaucescens Mig. 237 miquelii Merr. 237 Aidia cochinchinensis Lour. 309 Ailanthopsis poilanei Gagnep. 226 Ailanthus Desf. 193, 194, 195, 196, 215, 216, 969 altissima (Mill.) Swingle 193, 215,216; 2075. 220 blancoi Merr. 218 cacodendron L’Herit. 220 calycina Pierre 218 erythrocarpa Carr. 220 esquirolii Léveillé 970 excelsa Roxb. 216, 217*, 219 fauveliana Pierre 219
986
FLORA MALESIANA
[ser. I, vol. 66
fordii Nooteboom 216, 217*, 220, 969* giraldii Dode 220 glandulosa Desf. 220 gracilis Salisb. 211 grandis Prain 218 imberbiflora F.v.M. 219 integrifolia Lamk 193, 206*, 216, 217*, 218 map, 220 ssp. calycina (Pierre) Nooteboom 218 map ssp. integrifolia 218 map var. B Lamk 219 Kurzii Prain 219 macrophylla Hort. 220 matirei Gagnep. 220 malabarica DC. 218, 219, 969 var. mollis K. & V. 219 pro parte malabarica (non DC.) F.-Vill. 218 var. mollis K. & V. 218 pro parte mascula Hort. 220 moluccana DC. 218 var. mollis (non K. & V.) Koord. 218 peekelii Melch. 218 var. glabrata C.T.White 218 peregrina F.A.Barkl. 220 Philippinensis Merr. 219 pongelion Gmel. 218, 220 procera Salisb. 220 punctata F.v.M. 220 rhodoptera F.v.M. 220 rubra Hort. 220 scripta Gagnep. 220 sutchuensis Dode 220 triphysa (Dennst.) Alston 193, 217*, 219 map, 220 vilmoriniana Dode 220 Ailantus = Ailanthus 215 Aizoaceae 4: 267-275; 6: 384, 450, 915 Albonia Buchoz 215 peregrina Buchoz 220 Alfaroa 143, 146, 148 Alismataceae 5: 317-334; 6: 158, 915 Allanthospermum Forman 968, 969, 970, 971*, 972 borneense Forman 971*, 972 ssp. rostratum Nooteboom 971*, 972 multicaule (Capuron) Nooteboom 970, 972* Allospondias lakonensis (Pierre) Stapf 226 Alnus japonica (Thunb.) Steud. 917 maritima (Marsh.) Nuttall 917 Aloe 31 Alphonsea arborea (Blco) Merr. 421 Alsodeia chrysodasys Mig. 944
glabra Burgersd. 235 Alternanthera Forsk. 915 bettzichiana = bettzickiana 916 bettzickiana (Regel) Nichols. 916* ficoidea (L.) R.Br. ex R. & S. 915 ssp. bettzickiana (Nichols.) Backer 917 var. versicolor (Regel) Backer 916 ficoides P.Beauv. 915 paronychioides St.Hill 915, 916 var. paronychioides 916* Alytostylis Hook. f. 95 Alyxia spanogheana Mig. 336 tetragona R.Br. 336 Amaracarpus 387 Amaranthaceae 4: 69-98, 593; 5: 554; 6: 450, 915-917 Amaranthus dubius Mart. 915 hybridus L. ssp. incurvatus (Gren. & Godr.) Brenan var. paniculatus (L.) Mansf. 915 Amaroria A.Gray 221 soulameoides A.Gray 221 Amentiferae 143 Amoora aphanamixis Auct. non Re 8S. 921 Amphicalyx heterophyllus (Bl\.) Hassk. 738 latifolius Bl. ex Hassk. 738 pilosa Bl. 729 Amyxa cf. Airy Shaw 3, 5, 6, 47 pluricornis (Radlk.) Domke 47* Anacardiaceae 194, 226, 291, 928 Anacolosa frutescens BI. 420 Anacyclodon Jungh. 424, 425 pungens Jungh. 431 Anagallis L. 173, 174, 175, 176 subg. Anagallis 176 subg. Centunculus (L.) P.Taylor 176 subg. Jirasekia (Schmidt) P. Taylor 176 arvensis L. 176 nana Schinz 176 pumila Sw. 176* var. pumila 176 Anasser Juss. 369 laniti Blco 373 moluccana Lamk 373 Anassera rumphii Span. 373 Ancistrocladaceae 4: 8-10; 5: 553 Andrachne cavalerieri Léveillé 179 Andresia Sleum. 669, 943 malayana (Scort. in Hook. f.)
1O5SE220)
Sleum. 669* Andreya 295, 296 Andromeda 472 ovalifolia Wall. 675 Androsace Tourn. 173, 175, 186 rotundifolia Hardw. 192 saxifragifolia Bunge 191, 192 tonkinensis Bonati 382, 383 umbellata Merr. 191, 192 Anhella tristis 873 Anisophyllea R.Br. ex Sabine 966 beccariana Baill 966 ferruginea Ding Hou 966 grandis (Bth.) Burk. 361 Annamocarya Chev. 143 Annonaceae 421 Annulodiscus Tardieu Blot 932 nigricans Tardieu Blot 932 Anthocleista 293, 295 Anthodendron Rchb. 661 Antirrhoea hexasperma (Roxb.) Merr. 38 Antonia Pohl 295, 296 griffithii Wight 341 Antoniaceae Hutch. 296 Apocynaceae 93, 294, 297, 315, 318, 363, 368, 369 tribe Tabernaemontanineae 297 Apodostigma 227, 390 Aponogetonaceae 4: 11-12; 5S: 553 Aquilaria Lamk 2, 3, 4, 5, 6, 31 sect. Agallochum Hall. f. 6 sect. Amphinoman Hall. f. 6 sect. Brachythalamus Hall. f. 39 sect. Gyrinops Hall. f. 39 sect. Gyrinopsis Hall. f. 6 sect. Lachnolepsis Hall. f. 39 acuminata Quis. 11 agallocha Roxb. 982 apiculata Merr. 2, 8, 11 bancana Miq. 15 beccariana v. Tiegh. 7*, 8, 13, 41 borneensis Gilg 9 brachyantha (Merr.) Hall. f. 7* 2810812 citrinaecarpa (Elm.) Hall. f. 8, 10 cumingiana (Decne) Ridl. 1, 4.6. 7*. 85 la sks var. parviflora Airy Shaw 13 filaria (Oken) Merr. 8, 11 map, 12 grandifolia Domke 13 hirta Ridl. 7*, 8, 12 macrophylla Miq. 15 malaccensis Lamk 7*, 8, 9, 982 microcarpa Baill. 7*, 8, 9 moluccana Hall. f. 39 moszkowskii Gilg 12
Sept. 1972]
Index to scientific names
987
ovata Cav. 9 parvifolia (Quis.) Quis. 8, 12 pentandra Blco 15 podocarpus Hall. f. 42 rostrata Ridl. 8, 13 secundaria DC. 9 tomentosa Gilg 11 urdanetensis (Elm.) Hall. f. 6, 8, 10 versteegii Hall. f. 42 Aquilariella v.Tiegh. 6 borneensis v.Tiegh. 9 malaccensis v.Tiegh. 9 microcarpa v.Tiegh. 9 Araliaceae 194 Araucaria 356 Arbor coeli sive Caju langit Rumph. 218 Arbutus 472 coriacea Bl. 828 Arctostaphylos 472 Ardisia ochracea Elm. 260 Argyreia capitata (Vahl) Choisy 941 capitiformis (Poir.) Ooststr. 941 congesta Ooststr. 941 Arthrosolen 4 Aruba Aubl. 202 Arytera litoralis Bl. 226 Asperula 294 Aspidopterys elliptica (BI.) Juss. 960 Astrephia chinensis Dufr. 455 Astroloma R.Br. 424 Azalea (non L. sensu Salisb.) Desv. 661 subg. Pentanthera (G.Don) K.Koch 661 subg. Tsutsutsi (G.Don) K.Koch 661 brookeana (Low ex Lindl.) O.K. 655 citrina Hassk. 579 indica L. 663 indica (non L.) Sims 665 var. angustifolia Bl. 663 var. floribunda Bl. 663 var. spathulata Bl. 663 var. variegata Bl. 663 Jasminiflora (Hook.) O.K. 565 javanica (Bl.) O.K. 624 lamponga (Miq.) O.K. 544 ledifolia Hook. 662 malayana (Jack) O.K. 532 mollis Bl. 661 moulmainensis (Hook.) O.K. 666 mucronata Bl. 662 multicolor (Miq.) O.K. 621 retusa (Bl.) O.K. 482 rosmarinifolia Burm. f. 662 sinensis Lodd. 661 subsessilis (Rendle) Copel. f. 664
teysmannii (Miq.) O.K. 626 tubiflora Bl. ex DC. 532
Banksia Forst. 44 musculiformis Gaertn. 367, 368 Basellaceae 5: 300-304; 6: 450 Batidaceae 5: 414-415; 6: 917 Batis argillicola v.Royen 917 Béla-Moédagam Rheede 951 Bennettia papuana Gilg 944 Bernadina Baudo 177, 184 laurina Baudo var. stenia Baudo 184 mauritiensis Baudo 184 parviflora Baudo 185 Bertuchia Dennst. 299, 301 | Betula-Alnus maritima Marsh. 917 Betulaceae 5: 207-208; 6: 143, 472, 917 Bhesa Ham. ex Arn. 227, 228, 230, 231, 232, 280, 283 map, 390, 391, 930 archboldiana (Merr. & Perry) Ding Hou 281%, 282, 284 indica (Bedd.) Ding Hou 281*, 282, 283 moja Ham. ex Arn. 283 paniculata Arn. 229, 231, 280, 281*, 282, 283 robusta (Roxb.) Ding Hou 280, 281*, 282, 283 Biporeia Petit-Thouars 199 Bischofia javanica Bl. 53 Bixaceae s. str. 4: 239-241 Blumeodendron papuanum P. & H. 944 Blyxa novoguineensis Hartog 952 Boerhaavia = Boerhavia 452 Boerhavea = Boerhavia 454 Boerhavia L. 450, 451, 452 sect. Adenophorae Heimer! 452 acutifolia S.Moore 454 chinensis (L.) Aschers. & Schweinf. 450, 452, 453%, 454, 455 map diandra L. 454 diffusa L. 453*, 454 f. B paniculata (Rich.) O.K. 454 f. y repens (L.) O.K. 454 var. acutifolia DC. 454 var. obtusifolia DC. 454 var. pubescens DC. 454 erecta (non L.) Burm. f. 454 erecta L. 452, 453*, 454 glabrata Bl. 454 glutinosa Vahl 454 helenae R. & S. 455 hirsuta L. 454 mutabilis R.Br. 454 var. pubescens (R.Br.)
Choisy 454 procumbens Banks ex Roxb. 454 pubescens R.Br. 454 repanda Willd. 455 repens L. 454 var. diffusa (L.) Boiss. 454 var. procumbens (Roxb.) Hook. 454 scandens L. var. chinensis (L.) O.K. 455 tetrandra Forst. 454 Bonamia Thou. 936 Bonyunia Schomb. 295, 296, 954 Boscia 61 Bougainvillea Comm. ex Juss. 450, 451, 455 sect. Tricycla 456 < buttiana Holtt. & Stand. 456 glabra Choisy 451, 457 peruviana Humb. & Bonpl. 456, 457 spectabilis Willd. 456, 457 var. glabra (Choisy) Hook. 457 Brachythalamus Gilg 39 caudatus Gilg 42 podocarpus Gilg 42 versteegii Gilg 42 Brassiantha A.C.Smith 227, 230, 389, 390, 391, 392, 393, 394, 930 pentamera A.C.Smith 393*, 394 Bromheadia Lindl. 976 Brossaea bandongensis (Zoll. ex Miq.) O.K. 695 fragrantissima (non Wall.) O.K. 688 leucocarpa Bl. 692 nummularioides (D.Don) O.K. 684 a. normalis O.K. 684 B glauca O.K. 684 Brownlowia tersa (L.) Kosterm. 203 Brucea J.F.Mill. 193, 194, 196, 209 acuminata Li 212 amarissima Desy. ex Gomes 211 dubia Steud. 214 glabrata Decne 211 gracilis DC. 211 javanica (L.) Merr. 195, 209, 210*, 211 map, 212 luzonensis Vidal 212 macrobotrys Merr. 212 membranacea Merr. 212 mollis Wall. ex-Kurz 193, 210*, 211, 212 map quercifolia Seem. 212 stenophylla Merr. 212 sumatrana Roxb. 211
988
FLORA MALESIANA
[ser. I, vol. 68
sumatrensis Spreng. 211 Bruguiera Lamk 965, 966 eriopetala W. & A. ex Arn. var. exsetata Val. 966 exaristata Ding Hou 966 gymnorrhiza (L.) Lamk 966 hainesii C.G.Rogers 966 Bruinsmia Boerl. & Koord. 976 styracoides Boerl. & Koord. 976 Buddleia, see Buddleja 336 Buddleja Houst. ex L. 293, 294, 295, 296, 297, 336, 953, 954 subg. Buddleja 336, 337 sect. Alternifolia Marquand 339 subg. Nicodemia Leenh. 336, 337, 340 acuminatissima Bl). 337 amentacea Kranzl. 339 arfakensis Kaneh. & Hatus. 339 asiatica Lour. 293, 336, 337, 338*, 339%: 954 var. brevispica Val. 337 var. densiflora (Bl.) K. & V.
337
var. salicina (Bl.) K. & V. 337
var. sundaica (Bl.) K. & V. S57
brasiliensis Jacq. f. 340 crispa Bth. 337 curviflora H. & A. 337 davidii Franchet 336, 340, 954 densiflora Bl. 339 diversifolia Vahl 340 gynandra Marquand 336 indica Lamk 340 japonica Linden 337 lindleyana Fortune 337 madagascariensis Lamk 337, 340 neemda Buch.Ham. ex Roxb. 337 var. philippinensis Cham. & Schlechtend. 337 officinalis Maxim. 336 otophylla Hassk. 340 paniculata Wall. 337 salicina Lamk 337 sundaica Bl. 339 venenifera Makino 337 virgata L. f. 339 Buddlejaceae 295, 954 Buginvillaea = Bougainvillea 455 Buginvillia racemosa Blco 460 Bulbophyllum 109, 113 Burmanniaceae 4: 13-26, 592; 5: 553 Burseraceae 5: 209-296, 567; 6: 194, 195, 917-928 Busbeckea Endl. 69 Butomaceae 5: 118-120, 566
Cactaceae 450 Cadaba Forsk. 61, 62, 93, 97 map sect. Cadaba 93 sect. Eu-Cadaba Endl. 93 capparoides DC. 62, 93, 94*, 95*, 97 map Cadellia 194 Callitrichaceae 4: 251-252 Calogyne pilosa R.Br. 951 Calpidia Thouars 457, 459 brubobiana (Endl.) Heimerl 461 cauliflora (Scheff.) Heimer] 462 corniculata Heimer! 464 cuspidata Heimer! 462 excelsa Heimer] 461 grandifolia Heimer! 463 lauterbachii Warb. ex Heimer] 463 longirostris Heimer! 463 miilleriana (Warb.) Heimer] 462 pancheriana 459 rostrata Heimerl 463 spathiphylla Heimer! 463 Camotain Blco 355 Campanopsis (R.Br.) O.K. 111 marginata O.K. 116 var. rigida O.K. 116 Campanula 107, 108, 109, 112 sect. Campanopsis R.Br. 111 agrestis Wall. 115 carnosa Wall. 111 ceylanica Seba 129 circaeoides Fr.Schmidt ex Mig. 111 dehiscens Roxb. ex Wall. 115 gracilis Forst. 115 gracilis (non Forst.) Sims 118 indica Dietr. 115 lancifolia Roxb. 120 lavandulaefolia Reinw. ex BI. ii 15) littoralis Labill. 115 marginata Thunb. 115 medium L. 141 quadrifida R.Br. 115 rapunculoides L. 141 rotundifolia 110 sieberi Dietr. 115 vincaeflora Vent. 118 Campanulaceae 107-141, 928 subfam. Campanuloideae 108 tribe Campanuleae Bth. 109 tribe Pentaphragmeae Schonl. 109 tribe Sphenocleae Schon. 09
subfam. Lobelioideae 108, 122 tribe Lobelieae Bth. 109, 140 Campanulopsis Zoll. & Mor. 111 cyanea Zoll. & Morr. 115
Campanumoea BI. 109, 118, 119 axillaris Oliv. 120 celebica Bl. 120, 121 cordata Miq. 120 Japonica Maxim. 120 javanica Bl. 119 var. japonica (Maxim.) Ma- kino 120 lancifolia Merr. 119, 121 maximowiczii Honda 120 truncata Diels 120 Campylostemon Welw. 389, 390 Canarina moluccana Roxb. 928 Canarium Stickm. 917, 921 subg. Africanarium Leenh. 922 sect. Africanarium Leenh. 922 subg. Canariellum 922 subg. Canarium 921 sect. Canarium 922, 924, 927 sect. Pimela 922, 924, 927, 928 acutifolium (DC.) Merr. 926 var. acutifolium 926 album (Lour.) Raeusch. 922, 926, 928 apertum H.J.Lam 924 asperum Bth. 922, 926, 927 ssp. asperum var. asperum 926 australianum F.v.M. 925 bacciferum Leenh. 926 balsamiferum Willd. 924 caudatum King 923, 927 f. auriculiferum Leenh. 923 f. caudatum 922 cestracion Leenh. 922, 925 chinare Grutterink & HJ. Lam 922, 925 decumanum Gaertn. 924 denticulatum Bl. 924 ssp. kostermansil 924 dichotomum (BI.) Miq. 925 divergens Engl. 922, 923 euryphyllum Perk. 924 fusco-calycinum Ridl. 925 gracile Engl. 925 grandifolium (Ridl.) H.J.Lam 924 hirsutum Willd. 925 var. hirsutum 925 indicum L. 924 intermedium H.J.Lam 923, 925 kaniense Laut. 924 karoense H.J.Lam 924 kinabaluense Leenh. 923 kipella (BI.) Mig. 923, 925, 927 kostermansii Leenh. 924 lamii Leenh. 922, 923 latistipulatum Ridl. 923
Leenh.
Sept. 1972]
Index to scientific names
989
littorale Bl. 922, 923 f. pruinosum (Engl.) Leenh. 923 luzonicum (BI.) A.Gray 922, 924, 926 macadamii Leenh. 922, 925* maluense Laut. 923 ssp. maluense 923 megacarpum Leenh. 923 megalanthum Merr. 924 merrillii H.J.Lam 925 odontophyllum Migq. 924 oleosum (Lamk) Engl. 924, 927 ovatum Engl. 922, 924 parvum Leenh. 928 patentinervium Miq. 922, 923, 927 perlisanum Leenh. 923 piloso-sylvestre Leenh. 923 pilosum Benn. 924, 927, 928 ssp. borneensis Leenh. 924 ssp. pilosum 922 pimela Koen. 927 pimela Leenh. 923, 927 polyphyllum K.Sch. 925 pseudodecumanum Hochr. 924 pseudopatentinervium H.J.Lam 924 pseudosumatranum Leenh. 925 reniforme Kochummen & Whitmore 922, 927 rigidum (BI.) Mig. 925 salomonense B.L.Burtt 923 samoense Engl. 926 schlechteri Laut. 926 sinense Cana Rumph. 926 sinense Tsjacana Rumph. 927 smithii Leenh. 926 sumatranum Boerl. & Koord. 925 sylvestre Gaertn. 923 trigonum H.J.Lam 924 vitiense A.Gray 922, 926 vrieseanum Engl. 922, 926 vulgare Leenh. 922, 924 Cankrienia de Vriese 186 chrysantha de Vriese 190 farinosa Zoll. 190
Cannabinaceae 4: 222-223
Capparidaceae 61-105 subfam. Capparidoideae 62 subfam. Cleomoideae 62 Capparidales 63 Capparis Tourn. ex L. 61, 62, 63, 69, 73 map, 89, 91 sect. Busbeckea (Endl.) B. & H. 69, 70, 92 sect. Capparis 70 subsect. Corymbosae DC. 70
subsect. Pedicellares DC. 70 subsect. Seriales DC. 70
sect. Eucapparis Plum. ex DC. 70 sect. Monostichocalyx Radlk. 70, 86 acuminata Willd. 84 acuta (sphalm.) Koord. 92 affinis Merr. 79 andamanica King 78 aurantioides Pres] 87 baducca (non L.) Blco 90 billardierii DC. 86 borneensis Merr. 83 brachybotrya Hall. f. 71, 73 f. angustifolia (Hall. f.) Ja- cobs 75* f. brachybotrya 75 var. angustifolia Hall. f. 75 brachyscias Turez. 83 brevispina DC. 87 breynia Jacq. 70 buwaldae Jacobs 71, 85 callophylla BI. 71, 81, 82 map callophylla (non Bl.) Mig. 77 callosa Bl. 86 canescens Banks ex DC. 92 cantoniensis Lour. 70, 71, 76 map, 77, 81, 93 carandas Burm. f. 93 carolinensis Kaneh. 82 celebica Miq. 76 cerasifolia A.Gray 83, 84 copelandii Elm. 77, 78 cordifolia Lamk 89 corymbosa Lamk 81 crassifolia (sphalm.) A.Gray 84 crassifolia (sphalm.) C.Muell. 83, 84 cucurbitina King 71, 85 cumingii Merr. & Rolfe 81 dahlii Gilg & K.Sch. 82 dasypetala Turcz. 84 dealbata (non DC.) Back. 83 dealbata DC. 87 diffusa Ridl. 71, 81 discolor (non Donn.Smith) Standl. 93 elliptica Span. 77 emarginata Pres] 79 erycibe Hall. f. 70, 71, 74*, 75 erythrodasys Miq. 87 finlaysoniana Wall. ex Hook. f. & Th. 86 flexuosa Bl. 86 floribunda Wight 70, 71, 78 map f. floribunda 78 map f. induta Jacobs 70, 78 map foetida Bl. 84 forsteniana Miq. 86 galeata Fresen. 90 grandiflora Wall. ex Hook. f. & Th. 84 hasseltiana Miq. 76 horrida L. f. 87
var. B erythrodasys Miq. 87 horrida (non L. f.) Mig. 84 var. a Miq. 84 ilocana Merr. 79 korthalsiana Miq. 86 kunstleri King 73 lanceolaris DC. 70, 71, 76, 77 map, 78, 79, 82 larutensis King 70, 89 lasiopoda Turcz. 83 linearis (non Jacq.) Blco 87 littoralis Merr. 79 lobbiana Turcz. 70, 71, 79, 80* loheri Merr. 79 longestipitata Heine 70, 71, 77 longipes Merr. 73, 93 longipes (non Merr.) Stand. 93 lucida (Banks ex DC.) Bth. 62, 70, 92 map luzonensis Turcz. 78, 79 var. ampla Merr. 78, 79 mariana Jacq. 89 micracantha (non DC.) Blco 87 micracantha DC. 62, 70, 71, 75, 85, 86, 89 ssp. korthalsiana (Miq.) Ja- cobs 85, 86 map ssp. micracantha 85, 86 map var. callosa (Bl.) Hall. f. 86 micrantha (sphalm.) (non A. Rich.) Spreng. 85 mitchellii Lindl. 92 mucronata Elmer 81 myrioneura Hall. f. 83, 86 var. latifolia Hall. f. 86, 87 nemorosa (non Jacq.) Blco 87 nigricans Span. 83 nobilis (non Bth.) F.v.M. 92 nummularia DC. 91 oblongata Merr. 77, 78 octandra Jacq. 78 odorata Blco 86 oligostema Hayata 78 ovalifolia Zipp. ex Miq. 86, 87 oxyphylla Miq. 84 palawanensis Merr. 79 paniculata Ridl. 75, 76 perakensis (Scort. ex King) Rid]. 83 platyacantha Turcz. 77 pubiflora DC. 62, 70, 71, 82, 83*, 84 map, 85, 93 var. moluccana Miq. 83 var. perakensis Scort. 83 var. sumatrana Miq. 83 pumila Champ. 76 pyrifolia Lamk 70, 71, 84 map, 89 quiniflora DC. 70, 71, 73 map, 81, 88*, 89 renominata Jacobs 93 retusella Thw. 79
990
FLORA MALESIANA
[ser. I, vol. 66
richii A.Gray 89 robusta Heine 81 roxburghii (non DC.) Span. 77 roydsiaefolia Kurz 86 rufescens Turcz. 87 salaccensis Bl. 76, 93 var. celebica Miq. 76, 77 sandwichiana DC. 90 scortechinii King 62, 70, 71 sepiaria L. 71, 76, 79, 81 map, 92 var. acuta Vidal 79 var. B glabrata DC. 79 var. grandifolia Kurz & Prain 79 var. retusella Thw. 79 var. trichopetala Val. 79 var. vulgaris Hook. f. & Th. 79 spinosa L. 62, 70, 89, 91, 92 var. mariana (Jacq.) K.Sch. & Laut. 70, 89, 90*, 91* var. nummularia (DC.) F. M.Bailey 91 subacuta Miq. 79, 92 subcordata Span. 89 subspinosa Roxb. 77 tomentosa Lamk 81 torricellensis Laut. 77 trapeziflora Span. 89 trichopetala Val. 79 trinervia Hook. f. & Th. 70, 71, 73 map turczaninowli Elmer 81 tylophylla Spreng. 81 umbellata R.Br. ex DC. 79, 81 venosa Merr. 86 versicolor Griff. 93 viminea (non Hook. f. & Th.) Fern.-Vill. 87 viridis Elmer 77, 78 zeylanica (non L.) DC. 84 zeylanica L. 70, 71, 72*, 86, 87 map zeylanica (non L.) Roxb. 87 zippeliana Miq. 71, 78, 82, 86 map var. novobritannica Laut. 82 var. novohibernica Laut. 82 Caprifoliaceae 4: 175-194, 598: 6: 928-930 Capura L. 28 Capusia Lecomte 394 Carallia Roxb. 965, 966 borneensis Oliv. 966 eugenioidea King 966 longipes Ding Hou 966, 967* papuana Ding Hou 967 suffruticosa Rid]. 966 Cardiocarpus Reinw. 221 amarus Reinw. 221 Cardiophora Bth. 221 hindsii Bth. 221 Carices 953 Carissa carandas L. 93
grandis Bert. ex Guillemin 335 Carya 143 illinoensis 143 Caryo juglans Kirchh. 143 Caryophyllaceae 450 Caryospermum BI. 230, 288 alpestre O.K. 290 arborescens F.v.M. 291 moluccanum Bl. 291 Philippinense Vidal 291 serrulatum Migq. 290 Casearia grewiaefolia Vent. 944 var. deglabrata K. & V. 944 var. gelonioides (BI.) Sleum. 9
(Wang) K.Koch
pallida Craib 944 velutina BI. 38 Cassine L. 231, 232, 284, 391, 392, 930 subg. Elaeodendron Loes. 285 sect. Cassine Loes. 285 sect. Elaeodendron Loes. 285 australe (Vent.) O.K. 287 discolor Wall. in Roxb. 275 elliptica O.K. 286 fortunei O.K. 252 glauca (Rottb.) O.K. 228, 284, 286 var. cochinchinensis Pierre 285*, 286 map viburnifolia (Juss.) Ding Hou 228, 284, 285*, 286 map Cassiope 472 Casuarina 152, 221 junghuhniana Miq. 153, 187 Casuarinaceae 143 Catanthera F.v.M. 914 Catha Forsk. 229 edulis Forsk. 230 fasciculata Tul. 51 montana G.Don 241 vitiensis A.Gray 242 Caulinia Willd. 153, 164 indica Willd. 164, 166 ovalis R.Br. 953 Cedrota guianensis Blco 461 Celastraceae 49, 227-291, 389- 421, 930-932 tribe Hippocrateaceae Hook. f. 389 Celastrales 53, 390, 968 Celastrineae R.Br. 389 Celastrus L. 228, 229, 230, 231, 232, 233, 238, 240, 391, 392, 930 sect. Eucelastrus W. & A. 233 sect. Gymnosporia W. & A. 238 subg. Celastrus 234 ser. Axillares Rehd. & Wils. 234 ser. Paniculati Wils. 234
Rehd. &
alpestris Bl. 290 apoensis Elmer 236 australis Harv. & F.v.M. 235 axillaris Ridl. 236 bivalvis Jack 276, 277 championii (non Bth.) King 236 dispermus 230 diversifolius Hemsl. 242 emarginatus Willd. 241, 242 franchetiana Loes. 236 glaucus Vahl 286 hindsii Bth. 234, 235, 236 jackianus Steud. 237 lucida Wall. 237 lucidus L. 237 malayensis Ridl. 236 marianensis Koidz. 236 micrantha Roxb., nomen 237 monospermoides Loes. 234, 235, 236, 237, 243 monospermus Roxb. 234 montana = montanus 241 montanus Roth 241, 242 montanus Roxb. 242 multiflorus Roxb., nomen 235 novoguineensis Merr. & Perry 233722345255 nutans Roxb., nomen 235 obtusifolia Roth 237 oppositus Wall. in Roxb. 288 orbiculatus 228 paniculatus Willd. 229, 231, 234, 235 ssp. multiflorus Ding Hou 235 ssp. paniculatus Ding Hou 235 ssp. serratus Ding Hou 235 var. balansae Loes. 235 var. poilanei Tardieu 235 var. venulosoides Kanj. & Das 235 papuana Warb. 235 pauciflora Wall. 237 polybotrys Turcz. 235 racemosa Turcz. 235, 290 racemulosa Franch. 236 racemulosus Hassk. 236 repandus Bl. 237 robustus Roxb. 283 scandens L. 228, 230 semiarillata Turez. 241 senegalensis Lam. 242 stylosus Wall. 234, 235*, 237, (sphalm. Willd.) 420 ssp. glaber Ding Hou 237 ssp. stylosus Ding Hou 237 subspicatus Hook. 235 tonkinensis Pitard 236 trigyna Roxb. 237, 238 wallichii G.Don 237 Centrolepidaceae 5: 421-427 Centropogon lucyanus Schonl. 108
Sept. 1972]
Index to scientific names
Centrospermae v.Wettst. 174, 450, 451 Centunculus L. 174, 175, 176 indicus Royle 176 pentandrus R.Br. 176 pumilus O.K. 176 tenellus Duby 176 Ceodes J. & G.Forst. 457, 459 brunoniana Skottsb. 461 corniculata Merr. & Perry 464 corniculata (non Barg.-Petr.) Merr. & Perry 462 excelsa Skottsb. 461 longirostris Merr. & Perry 463 umbellata = umbellifera 460 umbellifera J. & G.Forst. 460, (sphalm. Skottsb.) 461 urocarpa Merr. & Perry 463 Cephaelis 336 Cephalostigma DC. 12 113 erectum (Roth) Vatke 113 hirsuta Edgew. 113 hookeri Clarke 114 paniculatum DC. 114 paniculatum (non DC.) Hos- seus 114 schimperi Hochst. ex Rich. 113
LOO 10 1,
Ceratophyllaceae 4: 41-42 Ceratopteris thalictroides Brongn. 171 Cerbera 368 musculiformis Lamk 367, 368 Cerium Lour. 177 spicatum Lour. 185 Cervicina Delile 111 gracilis Britt. 116 Chailletia sumatrana Miq. 942 Chamaedaphne 472 Cheiloclinium 227, 389 Cheilotheca Hook. f. 670 Cheilotheca (non Hook. f.) Prain 669 malayana Scort. 669 Chenopodiaceae 4: 99-106, 594; 6: 450, 932 Chenopodium carinatum R.Br. 932 pumilio R.Br. 932 Chilianthus 296 Cladopus H.MOIl. 963 nymani H.MOll. 963 Cleidion spiciflorum (Burm. f.) Merr. 203 Cleisocratera Korth. 387 Cleistanthopsis Capuron 970 Cleome L. 61, 62, 63, 99, 100 sect. Cleomes (DC.) Schult. 100 subsect. Pedicellaria (DC.) Schult. 100 subsect. Siliquaria (Forsk.) Schult. 100 sect. Corynandra(e) (Schrad.)
Schult. 100
sect. Gymnogonia R.Br. 100
sect. Gynandropses (DC.) Schult. 100
sect. Polanisiae (DC.) Schult. 100
sect. Ranmanissa (Endl.) Gri- seb. 100 sect. Rutidosperma [Jtis 100 sect. Tarenaya (Raf.) IItis 100 aculeata L. 100, 104, 105* acutifolia Elmer 103 affinis (non DC.) Spreng. 101 alliacea Blco 101 alliodora Blico 101 aspera Koen. ex DC. 105* blumeana D.Dietr. 101 blumeana Schult. 101 chelidonii (non L. f.) Burk. 103 chelidonii L. f. 62, 100, 102, 103*, 104 ciliata Schum. & Thonn. 104 gigantea (non L.) Blco 101 gynandra L. 62, 100, 101 houtteana Schlechtend. 102 hulletii King 104 icosandra L. 103 natalensis 62 pentaphylla L. 101 rutidosperma DC. 98*, 100, 104, 105* sandwicensis A.Gray 102 speciosa Raf. 62, 100, 101 f. alba 101 speciosissima Deppe ex Lindl. 101 spinosa Jacq. 62, 100, 102 triphylla L. 101 viscosa L. 62, 100, 103*, 104 f. deglabrata (Back.) Jacobs 104 f. viscosa 103 Cnestis palala (Lour.) Merr. 933 ssp. diffusa (Blco) Andreas 933 stenopetala Griff. 933 steriopetala = stenopetala 933 Cocculus hirsutus (L.) Diels 939 villosus DC. 939 Cochlospermaceae 4: 61-63 Codonopsis Wall. 107, 109, 110, 118, 119* albiflora Griff. 120 celebica (Bl.) Miq. 119, 120, 121 cordata Hassk. 120 cordifolia Komarov 120 javanica (Bl.) Hook. f. 108, 110, 119, 120* lancifolia (Roxb.) Moeliono 110, 120, 928 ssp. celebica (BI.) Moeliono 121 ssp lancifolia 121, 928
100,
99]
leucocarpa Miq. 120, 121 parviflora 119* purpurea Wall. in Roxb. 119 truncata Wall. ex DC. 120 viridis 119 Coinochlamys 296 Combretaceae 4: 533-589; 5: 564; 6: 932-933 Combretocarpus Hook. f. 966 rotundatus (Miq.) Danser 966 Commicarpus Standl. 452 chinensis Heimerl 455 plumbaginea 453 Comocladia serrata Blco 419, 420 Compositae 108, 109 subfam. Cichorieae 108 Connaraceae 5: 495-541; 6: 933-936 Connaropsis rubescens Rid\. 934 Connarus cochinchinensis (Baill.) Pierre 935 conchocarpus F.v.M. 936 ssp. conchocarpus 936 ssp. schumannianus (Gilg) Leenh. 936 culionensis Merr. 935 var. culionensis 935 var. stellatus (Merr.) Leenh. 935 euphlebius Merr. 935 ssp. euphlebius var. bullatus Leenh. 935 ssp. moluccanus Leenh. 935 lamii Leenh. 935 lucens Schellenb. 935 monocarpus L. 935 odoratus Hook. f. 935 paniculatus Roxb. 935 var. hainensis (Merr.) Vidal 935 var. paniculatus 935 peekelii Schellenb. 936 pickeringii A.Gray 936 salomoniensis Schellenb. 936 schumannianus Gilg 936 semidecandrus Jack 935 villosus Jack 935 winkleri Schellenb. ssp. philippinensis Leenh. 936
Conopharyngia 294 Contortae 294, 297 Convolvulaceae 4: 388-512, 599: 5: 558; 6: 390, 936-941 Convolvulus acuminatus Vahl 941 capitatus Vahl 941 capitiformis Poir. 941 gangeticus L. 939 hastatus Desr., non Forsk. 939 maximus L. f. 941 ochraceus Lindl. 941 purpureus L. 941 Cordia clitoria Bleo 464
992
Coris 174 Cortex filarius Rumph. 11, 12 foetidus Rumph. 12 Cortusa 174 matthioli L. 174 Corynocarpaceae 4: 262-264; 5: 557; 6: 941 Corynocarpus cribbianus (F.M. Bailey) L.S.Smith 941 Costera J.J.S. 469, 470, 471, 473, 474, 740, 742 map, 746 borneensis J.J.S. 741, 744 cyclophylla (Airy Shaw) J.J.S. & Airy Shaw 741, 742, 743* elegans J.J.S. ex Dunselman 745 endertii J.J.S. 741, 745 lanaensis (Merr.) Airy Shaw 741, 745 loheri (Merr.) Airy Shaw 741, 744* lucida (Merr.) Airy Shaw 741, 742 ovalifolia J.J.S. 741, 745 sumatrana J.J.S. 741, 744 tetramera Sleum. 741, 744* Couthovia A.Gray 296, 363, 365 alata A.C.Smith 366 astyla Gilg & Bened. 365, 366 brachyura Gilg & Bened. 365 brassii S.Moore 365 calophylla Gilg & Bened. 367 celebica Koord. 366, 367 collina A.C.Smith 365, 366 corynocarpa A.Gray 365 densiflora K.Sch. 365 kochii Val. 366 leucocarpa Merr. & Perry 365 macrocarpa A.C.Smith 366 macroloba A.C.Smith 366 macrophylla Merr. & Perry 367 neo-ebudica Guill. 366 novo-britannica Kaneh. & Ha- tus. 365 novo-caledonica Gilg & Bened. 366 nymanii Gilg & Bened. 365 pachyantha A.C.Smith 366 pachypoda Gilg & Bened. 365 rhynchocarpa Gilg & Bened. 365, 366 sarcantha (non Gilg & Bened.) Cammerl. 367 sarcantha Gilg & Bened. 366 seemanni A.Gray 365 terminalioides Gilg & Bened. 365 toua Kaneh., 367 undulatifolia Kaneh. & Hatus. 366 urophylla Gilg & Bened. 366 yunzaingensis Merr. & Perry 365
742,
FLORA MALESIANA
[ser. I, vol. 66
Covilhamia Korth. 95 ovata Korth. 97 Coxia Endl. 177 Crassulaceae 4: 197-202; 6: 173, 195 Crataeva L. 63 Nirvala Ham. 68 octandra Blco 78 religiosa Forst. f. 65 Crateriphytum Scheff. ex Koord. 296, 363, 365 moluccanum Scheff. ex Boerl. 367, 368 Crateva L. 61, 63, 69 A Corner 68 adansonii DC. 68 axillaris Presl 66 B Corner 65 hansemannii K.Sch. 65 hygrophila Kurz 65, 69 lophosperma Kurz 68, 69 macrocarpa Kurz 65, 66 magna (Lour.) DC. 68 marmelos (L.) Correa 69 membranifolia Miq. 65 nurvala Ham. 62, 68 var. nurvala 64*, 65*, 68* odora Ham. 66 f. axillaris (Presl) Jacobs 65*, 66, 67* religiosa (non Forst. f.) Blco 68 religiosa (non Forst. f.) Bl. 68 var. nurvala (Ham.) Hook. f. & Th. 68 religiosa Forst. f. 65*, 66, 67, 69 speciosa Volkens 65 tapia (non L.) Bl. 66 tumulorum Miq. 66 unilocularis Ham. 69 Cressa L. 936 australis R.Br. 937 cretica L. 937* Crinum asiaticum L. 973 Cruciferae 62, 63 Cuervea 390 Cunoniaceae 49, 933 Cupania spinosa Blco 241, 242 Cuscuta L. subg. Cuscuta 936 subg. Grammica (Lour.) Yun- cker 936 Cyathodes Lab. 424, 433 colensoi (Hook. f.) Hook. f. 428, 431 laurina R.Br. ex Drude 437 Cyclamen 173, 175 persicum L. 175 Cyclocodon Griff. 118, 119 adnatus Griff. 120 distans Griff. 119 lancifolium Kurz 120 truncatum Hook. f. & Th. 120 Cymothoe Airy Shaw 740
cyclophylla Airy Shaw 742 Cynoctonum J.F.Gmel. 293, 295, 296, 297, 375, 959 mitreola (L.) Britt. 293, 374*, 375, 959 var. intermedia Hochr. 377 var. lilacina (Back.) Bakh. f. 375 var. orthocarpa Hochr. 375 paniculatum (Wall.) B.L.Rob. 375 pedicellatum (Bth.) B.L.Rob. 377, 960 petiolatum Gmel. 375 sphaerocarpum Leenh. 374*, 375, 377, 960 Cyrtophyllum Reinw. ex Bl. 299, 303 caudatum (Ridl.) Ridl. 307 fragrans (Roxb.) A.DC, 307 giganteum (Ridl.) Ridl. 307 lanceolatum DC. 307, 309 peregrinum Reinw. ex BI. 307 speciosum Bl. 303 var. montanum Ridl. 303 wallichii (Bth.) Ridl. 307
Dacryodes Vahl 917 breviracemosa Kalkman 919 costata (Benn.) H.J.Lam 918,
919 crassipes Kalkman 919 elmeri H.J.Lam 918 expansa (Ridl.) H.J.Lam 919 incurvata (Engl.) H.J.Lam 918, 919 kingii (Engl.) Kalkman 917, 919 laxa (Benn.) H.J.Lam 918, 919 longifolia (King) H.J.Lam 919 var. longifolia 919 macrocarpa (King) H.J.Lam 919 var. kostermansii (Kalk- man) Kalkman 919 var. macrocarpa 919 nervosa (H.J.Lam) Leenh. 918, 919 papuana Husson 919, 921 puberula (Benn.) H.J.Lam 918 rostrata (Bl.) H.J.Lam 918, 919 f. pubescens 918 rubiginosa H.J. Lam 918 rugosa (BI.) H.J.Lam 918, 919
Dais (non L.) auct. 15 coccinia Gaudich. 18, 21 dubiosa Bl. 20 dubiosa (non BI.) Decne 17 laurifolia (non Jacq.) Blco 18 octandra L. 17, 18
Dalrympelea Roxb. 51 pomifera Roxb. 58
Dalrympelia = Dalrympelea 51,
Sept. 1972]
58 Dalzellia (non Wight) Engl. 963 Daphne L. 2, 3, 4, 5, 6, 34, 35 sect. Daphnanthoides Gilg 37 sect. Eriosolena (BI.) Meisn. S18); Big) aquilaria Blco 31, 35 arisanensis Hayata 38 cannabina (non Lour.) Schau- er 33 cannabina (non Lour.) Wall. 37 composita (L. f.) Gilg 2, 36*, 37, 38 var. montana Hochr. 37 f. macrophylla Hochr. 37 decandra Bl. 38 foetida (non L.) Blco 32 indica (non L.) Blco 32 indica L. 35 javanica Thunb. 38 kiusiana Miq. 38 luzonica C.B.Rob. 2, 36*, 37, 38 montana Meisn. 38 odora Thunb. 38 papyracea Wall. ex Steud. 37 pendula Sm. 38 var. y concolor Meisn. 38 var. B montana Meisn. 38 Phaetida 32 sp. Steen 35 Daphnobryon Meisn. 43 ericoides Meisn. 44 Datiscaceae 4: 382-387 Datura 253 Daucus glochidiatus Fischer 984 Decaisnella O.K. 6 cumingiana O.K. 15 Decaspora laurina O.K. 437 Decatoca F.v.M. 423, 424, 434, 436 map spenceri F.v.M. 435* Deltaria 3 Denhamia Meisn. 240, 243 parvifolia L.S.Smith 240 pittosporoides F.v.M. 230, 240 Dentella 253 erecta Roth ex R. & S. 113 perotifolia Willd. ex R. & S. tS Desfontainea Kunth 295, 297, 954 Dichapetalaceae 5: 567; 6: 941-943 Dichapetalum Thou. 238, 941 gelonioides (Roxb.) Engl. 941, 942
(Labill.)
305-316,
ssp. pilosum Leenh. 942 ssp. tuberculatum Leenh. 9
glabrum (Vahl) Prance 942 grandifolium Ridl. 941, 942
Index to scientific names
griffithii (Hook. f.) Engl. 942 hainanense Eng}. 942 helferianum (Kurz) 941, 942 laurocerasus (Hook. f.) Engl. 942 longipetalum (Turcz.) Engl. 942 papuanum (Becc.) Boerl. 941, 942 ssp. borneense Leenh. 942 ssp. papuanum 942 peekelii Krause 941, 942 scorpioideum Leenh. 941, 943 sessiliflorum Leenh. 942 setosum Leenh. 942 steenisii Leenh. 942 ssp. steenisii 942 tenerum Leenh. 943 timoriense (DC.) Boer]. 941, 942 tricapsulare (Blco) Merr. 942 Diervilla fallax (Miq.) Boerl. 929 Dillenia 49 Dilleniaceae 4: 141-174; 5: 557; 6: 933 Dimorphanthera F.v.M. 469, 470, 471, 473, 474, 746, 885, 886 map sect. Brachychone Schltr 890 sect. Cyclosiphon Wernh. 890 sect. Dimorphanthera Sleum. 886, 890 subsect. Brachychone (Schltr) Sleum. 890 sect. Pteridosiphon Wernh. 886, 889 sect. Trochilanthe Schltr 887, 896 subsect. Trochilanthe (Schltr) Sleum. 896 alba J.J.S. 904 albiflora Schltr 887, 892 alpina J.J.S. 888, 905 var. alpina 905 var. pubigera Sleum. 905 alpivaga Sleum. 886, 889, 912 amblyornidis (Becc.) F.v.M. 892 var. amblyornidis 887, 892, 893*, 896 var. moorhousiana (F.v.M.) Sleum. 887, 893, 894* amoena Sleum. 886, 887, 896 anchorifera J.J.S. 887, 888, 902 apoana (Merr.) Schltr 887, 895 arfakensis J.J.S. 892 beccariana (Koord.) J.J.S. 888, 904 brachyantha Sleum. 887, 899 brassii Sleum. 887, 888, 897, 898*
Pierre
993
breviflos Sleum. 886, 891 brevipes Schltr 887, 888, 900 calodon Sleum. 887, 889, 910 chlorocarpa Sleum. 895 clemensiae Sleum. 888, 902, 903 * collinsii Sleum. 908 var. collinsii 888, 908 var. montis-wilhelmi Sleum. 888, 908, 909* cornuta J.J.S. 887, 899 var. cornuta 899 var. tenuiflora Sleum. 899 crassifolia Sleum. 888, 901 darmandvillei J.J.S. 914 declinata Sleum. 886, 890 dekockii J.J.S. 887, 893 var. chlorocarpa (Sleum.) Sleum. 894, 895 var. dekockii 893, 894 var. pubiflora Sleum. 887, 894, 895 denticulifera Sleum. 905 var. denticulifera 888, 889, 905 var. pubens Sleum. 888, 905 dielsiana J.J.S. 891 doctorsii J.J.S. 887, 888, 906 dryophila Sleum. 889, 914 var. dryophila 914 var. trichoclada Sleum. 914 elegantissima K.Sch. 888, 908 eymae Sleum. 887, 898 forbesii (F.v.M.) F.v.M. 888, 904 forbesii [non (F.v.M.) F.v.M.] Warb. 908 gracilis Sleum. 887, 900 hirsutiflora Sleum. 887, 897 intermedia (non J.J.S.) Diels 891 intermedia J.J.S. 886, 892 kalkmanii Sleum. 888, 908 kaniensis Schltr 908 kempteriana Schltr 886, 891 lancifolia Sleum. 889, 910 latifolia Schltr 887, 897 leucostoma Sleum. 888, 889, 905 longifolia Kaneh. & Hatus. 888, 907 macleaniaefolia = macleanii- folia 889 macleaniifolia Wernh. 886, 889 magnifica Sleum. 886, 891 megacalyx Sleum. 889, 910 meliphagidium = meliphagi- dum 906 meliphagidum (Becc.) F.v.M. 888, 906 microphylla Sleum. 889, 913 militaris J.J.S. 889, 910 mindanaensis Merr. 887, 892 moluccana J.J.S. 896
994
FLORA MALESIANA
[ser. I, vol. 66
moorhousiana F.v.M, 893 moorhousiana (non F.v.M. s. str.) Schltr 892 myzomelae (Becc.) J.J.S. 889, 913 nigropunctata Sleum. 888, 901 obovata J.J.S. 736, 911, 912 obtusifolia Sleum. 889, 913 ovatifolia Sleum. 888, 907 parviflora J.J.S. 889, 912 parvifolia J.J.S. 889, 912 peekelii Sleum. 888, 907 prainiana (Koord.) J.J.S. 889, 911* profusa J.J.S. 891 pulchra J.J.S. 887, 896 racemosa Schltr 887, 898 robbinsii Sleum. 888, 902 splendens Sleum. 888, 904 steinii Sleum. 887, 895 thibaudifolia Sleum. 888, 901 torricellensis Schltr 887, 899 tridens J.J.S. 886, 890 umbellata Wernh. 886, 890 vaccinioides Sleum. 889, 912 velutina Schltr 887, 896 vestita Sleum. 887, 900 vonroemeri (Koord.) J.J.S. 889, 911 wollastonii Wernh. 887, 898 womersleyi Sleum. 888, 906 wrightiana (Koord.) J.J.S. 889, 914 Dioscoreaceae 4: 293-335; 5: 557 Diosma serrata Blco 235 Diospyros hierniana (K. & G.) Bakh. 932 Diotocranus 297 Diplomorpha Meisn. 28 Diplusodon Pohl 975 Diplycosia Bl. 469, 471, 473, 474, 677, 696 map, 912 abscondita Sleum. 697, 703 acuminata Becc. 700, 722, 132% amboinensis Becc. 699, 716, 132% aperta J.J.S. 697, 706 apiculifera J.J.S. 701, 731 apoensis Elm. 699, 716 atjehensis Sleum. 697, 701, 707 aurea Sleum. 697, 704 haclay(a)ensis Elm. 740 barbigera Sleum. 697, 703 bartlettii Merr. 738 brachyantha Sleum. 698, 699, 701, 713 var. brachyantha 713 var. parvula Sleum. 713 breviflora Ridl. 676 calelanensis Elm. 720 capitata Sleum. 699, 702, 721 carrii Sleum. 697, 702
caryophylloides J.J.S. 699, 721 var. caryophylloides 721 var. longipes Sleum. 721 caudatifolia Sleum. 700, 729 celebensis J.J.S. 700, 730 chrysothrix Stapf 697, 702 ciliolata Hook. f. 698, 708, (Bis cinnabarina Sleum. 700, 724, (32% cinnamomifolia Stapf 702, 735*, 736 clementium Sleum. 697, 704, 105% commutata Sleum. 701, 734 consobrina Becc. 700, 724 cordifolia Ridl. 723 coriifolia Sleuam. 734 crassiramea Sleum. 699, 701, 719 crenulata Sleum. 700, 723 edulis Schltr 699, 701, 735 elliptica Rid]. 700, 723, 732*, 778 endertii J.J.S. 740 ensifolia Merr. 702, 735*, 737 erythrina (Hook.) K. & G. 815 fasciculiflora Merr. 720 filipes Sleum. 698, 708, 732* fimbriata Sleum. 699, 722 glabra Merr. 720 glauciflora Sleum. 698. 712 gracilipes J.J.S. 700, 730 haemantha Sleum. 700, 730 heterophylla Bl. 702, 738 var. heterophylla 738 var. latifolia (Bl.) Sleum. 701, 738, 739*, 740* var. obovata J.J.S. 738 heterophylla (non BI. s. str.) K. & G. 740 hirsuta Sleum. 698, 709 kalmiifolia Sleum. 699, 717 kemulensis J.J.S. 701, 731 kerintjensis J.J.S. 740 var. elliptica J.J.S. 740 kinabaluensis Stapf 699, 701, MAT 35% kingii Merr. 723 kjellbergii J.J.S. 698, 709 kosteri Sleum. 698, 710 kostermansii Sleum. 700, 724 lamii J.J.S. 699, 719 lancifolia Ridl. 700, 728 var. calvescens Sleum. 729 var. lancifolia 728 latifolia Bl. 738 latifolia (non B.) Hall. f. 738 lavandulifolia Sleum. 700, 727 ledermannii Schltr 698, 700, 713 lilianae J.J.S. 697, 706 loheri Merr. 697, 706 lorentzii Koord. 701, 736
lucida Merr. 742 luzonica (A.Gray) Merr. 699, 701, 720 var. calelanensis Sleum. 720 var. luzonica 720 var. merrittii (Merr.) Sleum. 720, 721 var. pubens Sleum.720 lysolepis Sleum. 701, 734 macrophylla Becc. 740 memecyloides (non Merr. 717 memecyloides Stapf 700, 729 merrittii Merr. 721 microphylla Becc. 700, 723, (32 microphylla (non Becc.) Clar- ke 723 minutiflora Sleum. 701, 731 morobeensis Sleum. 698, 713 var. morobeensis Sleum. 698, 713 var. ovatifolia Sleum. 714 mundula (F.v.M.) Schltr 684 muscicola Sleum. 699, 716 myrtillus Stapf 700, 728 opaca C.B.Robins. 721 orophila Sleum. 699, 702, 720 parvifolia Merr. 698, 711 pendens Sleum. 698, 699, 714 penduliflora Stapf 697, 701, 707, 735* piceifolia Sleum. 700, 726 pilosa Bl. 700, 729 pinifolia Stapf 700, 725*, 726", Que pittosporifolia J.J.S. 732 var. pittosporifolia 701, 732 var. punctiloba Sleum. 699, 733 pokapindjangensis J.J.S. 706 pseudorufescens Sleum. 700, 711 var. elliptifolia Sleum. 698, 700, 711 var. pseudorufescens 698, 711 pubivertex Sleum. 697, 707 punctulata Stapf 701, 736 retusa Sleum. 701, 733 rosea Sleum. 698, 709, 710* rosmarinifolia Sleum. 701, Wil Bye rubella Sleum. 700, 728 rubidiflora J.J.S. 701, 733 rufa (non Stapf) Merr. 704 rufa Stapf 697, 702 rufescens Schltr 698, 708 rupicola Sleum. 698, 714, 732* sagittanthera J.J.S. 700, 722 salicifolia Sleum. 702, 737 sanguinolenta Sleum. 701, 733 saurauioides J.J.S. 697, 704 scabrida Becc. 699, 721
(Elm.)
Stapf)
Sept. 1972]
scandens Merr. 720 schramii Sleum. 697, 703 schultzei Schltr 699, 714 setiloba Sleum. 700, 724 setosa J.J.S. 698, 710 soror Becc. 698, 699, 712 var. nuda Sleum. 701, 712 var. soror 712 sphenophylla Sleum. 132 e133 stellaris Sleum. 699, 715 stenophylla Sleum. 700, 727, 132* subglobularis Sleum. 699, 715 sumatrensis Merr. 699, 718* tetramera Sleum. 697, 698, TA lat triangulanthera J.J.S. 698, 708 trinervia Elm. 699, 719 var. trinervia 719 var. urdanetensis Sleum. 719 undata (non J.J.S.) H.J.Lam 706 undata J.J.S. 699, 715 urceolata (non Stapf) Ridl. var. 740 urceolata (non Stapf) Sleum. 734 urceolata Stapf 701, 734, 735* urdanetensis Elm. 719 varians Sleum. 699, 715 viridiflora Sleum. 702, 737 var. megalantha Sleum. 738 var. viridiflora 737 Dipsacaceae 4: 290-292; 5: 557 Dipterocarpaceae 154 Disiphon Schltr 785 papuanum Schltr 786 Distyliopsis Endress 952 Distylium Sieb. & Zucc. 952 Dodecatheon 174 Dodonaea 152 Dolianthus C.H.Wright 387 Dortmanna Adans. 121 Dortmannia (sphalm.) O.K. = Dortmanna Adans. 121 alsinoides O.K. 127 campanuloides O.K. 131 chinensis O.K. 131 colorata O.K. 124 erecta O.K. 124 griffithii O.K. 127 haenkeana O.K. 136 inconspicua O.K. 129 leschenaultiana O.K. 123 nicotianaefolia O.K. 124 pyramidalis O.K. 124 radicans O.K. 131 reinwardtiana O.K. 129 rosea O.K. 124 subcuneata O.K. 128 succulenta O.K. 128 trialata O.K. 129 trigona O.K. 127
701,
(Elm.)
Index to scientific names
var. affinis O.K. 128 var. microcarpa O.K. 127 var. terminalis O.K. 127 zeylanica O.K. 128, 129 Dovyalis macrodendron Gilg 944 Drapetes Banks ex Lamk 2, 4, 5, 6, 43 sect. Daphnobryon Boer). 43 dieffenbachii Hook. 43 ericoides Hook. f. 2, 43, 44* map muscosus Lamk 43 tasmanicus Hook. f. 43 Drimyspermum Reinw. 15, 16 ambiguum Meisn. 17 blumei Decne 20 blumei (non Decne) Hassk. 17 burmanni Decne 17, 18 cauliflorum Thw. 20 coccineum Becc. 18, 21 cumingti Meisn. 21 laurifolium Decne 17 laurifolium (non Decne) Hassk. 20 longifolium Miq. 17 macrocarpum Scheff. 22, 23 perrottetianum Decne 18 phaleria Meisn. 20 revolutum T. & B. 21 urens Reinw. 20 urens (non Reinw.) Scheff. 18 Drosera petiolaris R.Br. 943 spathulata Labill. 943 umbellata Lour. 191, 192 Droseraceae 4: 377-381; 5: 557; 6: 943 Drymispermum = mum 15, 16 Duabanga Buch.-Ham. 975* grandiflora (Roxb. ex DC.) Walp. 975*, 976 moluccana BI. 975* < taylorii Jayaweera 975* Durio 70
Drimysper-
Dysoxylum dasyphyllum Mia. 218 quercifolium (Seem.) A.C.
Smith 212
Ebelingia Rchb. 297 brownii Steud. 2)7 paucijuga O.K. 208 Edgeworthia 3 Elaeocarpaceae 420, 914 Elaeocarpus oxypyren K. & V. 420 Elaeodendron Jacq. f. ex Jacq. 228, 284, 285 australe Vent. 287 ellipticum Decne 286, 287 var. glauca Pierre 287 var. montana (Thw.) Pierre 287 var. roxburghii (W. & A.) Pierre 287
995
fortunei Turez. 252, 253 glaucum (Rottb.) Pers. 286 var. macrocarpa K. & V. 286 horizontale Turcz. 241 Javanicum Turez. 252, 253 microcarpum White & Francis 288 mindanaense Merr. 288 pauciflorum Tulasne 287 roxburghii W. & A. 286, 287 subrotundum King 228, 286 viburnifolium Merr. 286 Elaeodendrum = Elaeodendron 284 Elatinaceae 4: 203-206 Eleocharis 165 Ellipanthus beccarii Pierre 935 tomentosus Kurz 935 ssp. tomentosus 935 var. tomentosus 935 Emorya Torr. 295, 296, 953 Enchysia Presl 140 Engelhardia Lechen. ex BI. 143, 145, 146 map, 147, 148
sect. Engelhardia 145, 147, 148
sect. Oreomunnea_ (Oerst.) C.DC. 148
sect. Psilocarpeae Nagel emend. Leroy 145, 148
sect. Prerilema (Reinw.) C.DC. 148
sect. Trichotocarpeae Nagel 148
aceriflora Bl. 152 acerifolia (sphalm.) Hook. f. 152 apoensis Elm. ex Nagel 148, 151 chrysolepis Hance 154 colebrookeana Lindl. ex Wall. 153 esquirolii Lév. 153 fenzelti Merr. 154 formosana Hayata 143, 154 lepidota Schltr 148 mollis Hu 148 nucifera (Ludwig) Madler 147 nudiflora Hook. f. 150 var. crenata Hook. f. ex Gamble 150 palembanica Miq. 150 parvifolia C.DC. 146, 150 permicrophylla Elm. 150 philippinensis C.DC. 152 polystachya Radlk. 143, 154 pterococca O.K. 154 var. a roxburghiana O.K. 154 rigida BI. 144*, 145, 148, 149* roxburghiana Lindl. ex Wall. 152 roxburghiana Wall. 148, 149*, 154
996
FLORA MALESIANA
[ser. I, vol. 6®
selanica Bl. 154 serrata Bl. 144*, 146, 148, 149, 150* sp. Corner 151 spicata Lechen. ex BI. 143, 144*, 145, 146, 147*, 148, 151%) 152*) 153815451953 var. a genuina K. & V. 151 var. B aceriflora K. & V. Syl var. y colebrookiana K. & V. 151, 154 var. colebrookeana (Lindl. ex Wall.) O.K. 151, 153, 953 var. formosana Hayata 154 var. integra (Kurz) Man- ning 953 var. spicata 153 subsimplicifolia Merr. 148 villosa Kurz 153 var. integra Kurz 153, 953 wallichiana (Lindl. ex Wall.) C.DC. 154 B chrysolepis C.DC. 154 zambalensis Elm. 148, 149 Engelhardtia = Engelhardia 145 Englerodaphne Gilg 79 Enhalus acoroides (L. f.) Royle 952 Enkleia Griff. 2, 4, 5, 6, 23, 982 coriacea Hall. f. 25 malaccensis Griff. 24*, 25, 982 malayana Griff. 25 paniculata (Merr.) Hall. f.24*, 25 riouwensis Hall. f. 25 zippeliana Hall. f. 25, 26 Epacridaceae 50, 422 map-444, 943 subfam. Epacrideae 423 tribe Epacrideae 423 subfam. Styphelieae 423 tribe Styphelieae 423 Epacris 423 Epigynium coriaceum (Bl.) KI. 828 ellipticum (Bl.) K1. 873 floribundum (Bl.) K1. 872 laurifolium (Bl.) K1. 872 lucidum (Bl.) KI. 814 varingiaefolium (Bl.) Kl. 815 Ericaceae 49, 423, 424, 469-914, 943 subfam. Arbutoideae 473, 474 tribe Andromedeae 474 tribe Gaultherieae 474 tribe Pyroleae 474 subfam. Ericoideae 473 subfam. Monotropoideae 469, 473 subfam. Rhododendroideae 473 tribe Rhododendreae 474
subfam. Vaccinoideae 472, 474 Ericales 473, 669 Eriosolena Bl. 35 affinis Zoll. 38 composita v.Tiegh. 38 montana Bl. 38 var. @ macrophylla Hassk. 38 var. & minor Hassk. 38 pendula Bl. ex Lecomte 38 viridiflora Zoll. & Mor. 33 Erodium L’Heérit. 445, 448 cicutarium (L.) L’Heérit. ex Ait. 445, 448 Ervatamia sphaerocarpa Burk. 347 Erycibe 390 Erythroxylaceae 5: 543-552 Eucommiaceae 229 Euonymus Tournef. ex L. 227, D282 955 OMS eRe S252 43. 245, 253, 254, 256, 264, 274, 391, 392, 930 subg. Euonymus 247 subg. Kalonymus Beck 247 acuminifolius Blakelock 247, 251 var. borneensis 251 alatus {non (Thunb.) Sieb.] Elm. 249 alatus Sieb. 230 atropurpureus Jacq. 229, 230 bancanus Miq. 249 benguetensis Merr. 248, 253 castaneifolius Ridl. 246*, 247, 251 cochinchinensis 248, 286, 930 congolensis Wilczek 245 coriaceus Ridl. 249 decora W.W.Sm. 245 elmeri Merr. 249 europaeus L. 229, 230 f. genuinus Back. 248 fengii Chun & How 256 fimbriatus Baill. ex Laness. 267 fortunei (Turez.) Hand.-Mazz. DID e253 var. radicans (Miq.) Rehd. DD, fungosus Ohwi 252, 253 glaber Roxb. 247, 250 glandulosus (Merr.) Ding Hou 248, 251 grandiflorus Wall. 264 horsfieldii Turcz. 249 impressus Blakelock 247, 250 japonicus (Thunb.) L. f. 252, 253 japonicus Thunb. 246*, 248, 252 map var. radicans Miq. 252
(Bl.)
Blakelock
Pierre) 247,
javanicus Bl. 231, 245, 247, 248, 250 f. genuinus 248 var. conocarpus Hassk, 249 var. coriaceus 248 var. elmeri 248 var. genuina K. & V. 248 var. genuinus K. & V. 248 var. horsfieldii Turcz. 248 var. sphaerocarpus Hassk. 248, 249 var. talungensis Pierre 248 var. timorensis 248 latifolia (L.) Mill. 245, 246 lichiangensis W.W.Sm. 245 linearifolia Franch. 245 marivelensis Elm. 258 micropetalus Ridl. 249 moluccensis Blakelock 248, 254 moultoni Ridl. 251 nana M.B. 243 oliganthus Merr. 248 omeiensis Fang 245 pahangensis Ridl. 248 philippinensis Merr. 248 platyclinis Ohwi 243 radicans (Miq.) Sieb. ex Miq. 25D) recurvans Miq. 247, 250 rufulus Rid]. 250 sumatranus Miq. 248 timorensis (non Zipp. ex Span.) Turez. 248 timorensis Zipp. ex Span. 248, 249 tonkinensis Loes. 247 viburnifolius Merr. 248, 286 wrayi King 247, 250, 258 yunnanensis Franch. 245 Eupatorium inulifolium 145 Euphorbiaceae 4, 203, 420, 944 Eurycoma Jack 193, 196, 203 apiculata Benn. 205, 206 dubia Elmer 206 eglandulosa Merr. 206 harmandiana Pierre 205, 206, 969 latifolia (sphalm.) Rid]. 205
longifolia Jack 204*, 205, 206* map ssp. eglandulosa (Merr.)
Nooteboom 206 map ssp. longifolia 206 map var. cochinchinensis Pierre
205 var. merguensis Pierre 205
merguensis Planch. 205 tavoayana Wall. 205 Euscaphis Sieb. & Zucc. 49, 51 Evodia 226 meliaefolia (Hance) Bth. 206 parviflora Craib 56 triphylla DC. 214 Evolvulus L. 936
Sept. 1972]
Index to scientific names
997
gangeticus (L.) L. 939
sp. Ridsdale 960 Evonymus Auct. 245 Exobasidium 172, 471, 659, 747
Fagaceae 143, 145, 472 Fagaceae-Castaneae 143 Fagara 207, 209 piperita (non L.) Naves 208 Fagraea Thunb. 293, 294, 295, 297, 298*, 299, 301*, 303 map, 312, 318 sect. Corymbosae Bth. 315 sect. Cyrtophyllum (Reinw.) BI. 300, 303, 309 sect. Eufagraea Bl. 315 sect. Fagraea 300, 315 sect. Fagraeae verae Bl. 315 sect. Parviflorae Bth. 303 sect. Pseudocorymbosae Soler. 315 sect. Pseudoracemosae Soler. 311 sect. Racemosae Bth. 300, 309 subg. Cyrtophyllum (Bl.) Miq. 303 subg. Eufagraea Miq. 315 acuminatissima Merr. 302, 319, 320, 954 affinis S.Moore 335 alteniana F.v.M., nomen 335 amabilis S.Moore 332 amboinensis Bl. 316, 318 ampla S.Moore 333 angiensis Kaneh. & Hatus. Siveslseslo* annulata Hiern 300, 302, 303, 318, 333, 334, 335 anthocleistifolia Gilg & Bened. 326 appendiculata Bl. 311 archboldiana Merr. & Perry 317, 318 aurantiodora §.Moore 304 auricularia = auriculata 327 auriculata Jack 300, 301*, 302, 326, 329*, 954 ssp. auriculata 328, 954 ssp. borneensis (Scheff.) Leenh. 328, 329, 954 ssp. parviflora Leenh. 303, 328 berteriana A.Gray 301, 302, S116. 3175 31.9, 331,,333;335, 954 birmanica Gandoger 317 blumei G.Don 293, 299, 300, SOI 302, 315; 3116; 317, 318, 320, 322*, 324, 325, 954 ssp. blumei 323, 324, 954 ssp. plumeriaeflora (DC.) Leenh. 323, 325, 326 blumei Steud. 320 blumii = blumei 329 bodenii Wernh. 302, 333, 335,
336 borneensis Scheff. 327, 328 bracteosa Cammerl. 328 calcarea M.R.Hend. 303, 332 calophylloides Gilg & Bened. 335 cambagei Domin 300, 332, 333 cardinalicarpa Elm. 336 carnosa Jack 302, 316, 331 carstensensis Wernh. 302, 326, 327* caudata Ridl. 307, 309 ceilanica Thunb. 301, 302, BOS NS1SeS16s) 31828319. 320, 323, 954 f. khasiana 303 var. brevituba Clarke 315 celebica Bl. 316, 318 chinensis Merr. 317 coarctata Bl. 311 var. ligustrina (Bl.) Miq. 311 cochinchinensis (Lour.) A.Chev. 307, 309 congesta Bl. 316 congestiflora Elm. 311 cordifolia Bl. 311 coromandelina Wight 316 crassifolia Bl. 316 crassifolia Wall., non Bl. 320 crassipes Bth. 311 crenulata Maing. ex C.B.Clar- ke 293, 299, 300, 301, 324* cuernosensis Elm. 322, 323 currant Merr. 327, 328, 329 curtisii K. & G. 303, 331, 332 cuspidata Bl. 311 cymosa Merr. 311 dasyantha Gilg & Bened. 335 dolichopoda Gilg & Bened. 332) dubia Wall. 336 elata Merr. & Perry 300, 332 elliptica Roxb. 293, 300, 301, 303, 304*, 309, 315, 954 epiphytica Elm. 327, 328 eucalyptifolia Cammerl. 311, SSS 15 euneura Scheff. 327, 328 eymae Back. 301*, 303, 333 fagraeacea Druce 332 fastigiata Bl. 301*, 323, 324, 325, 954 fastigiata (non Bl.) Cammerl. 325, 328 fastigiata (non BI.) Ridl. 324 flavidula Ridl. 331 forstenii Bl. 316, 318 forstenii (non Bl.) Koord. 325 fragrans Roxb. 293, 300, 301, 303, 304, 305*, 306*, 307*, 308*, 309, 323, 954 fuscescens Bl. 316 galilai Gilg & Bened. 335 gardeniaeflora Wernh. 317, 318
gardenioides Ridl. 300, 302, 331, 332 ssp. borneensis Leenh. 332 ssp. gardenioides 331
garneri Thw. 317
gigantea Ridl. 307, 309
gitingensis Elm. 302, 317, 334, 335
gracilipes A.Gray 300, 301, 303, 309, 332, 334, 335, 954
gracilis Cammerl. 311
grandifolia Merr. 311
grandis Panch. & Sébert 335
heterophylla Bl. 316
imperialis Miq. 327, 328, 329*
intermedia Ridl. 322, 323
involucrata Merr. 301, 328, 3297755054 sole oor var. Jlongipetiolata
329, 330
Jackii Elm. 327, 328
jJasminodora Gilg & Bened. 333
Javanensis Bakh. f. 304
javanica (Reinw. ex) Bl. 327, 328
khasiana Bth. 316
kimanga = kimangu 303
kimangu Bl. 303
ksid Gilg & Bened. 335
kusaiana Hosok. 335
lanceolata Bl. 316
lanceolata (non Bl.) Hend. 331
lanceolata Schnizl. 309
lanceolata Wall., non Bl. 307
latifolia Mig. 311
ledermannii Gilg & Bened. 335
leschenaultii B\. 320, 323
ligustrina Bl. 311, 312*, 315 var. brachystachya Bth. 311 var. disparifolia Bl. 311
litoralis Bl. 316, 317, 318, 323, 954 var. amboinensis Bl. 316 var. forstenii (Bl.) Mig. 316 var. moluccana Miq,. 316
littoralis = litoralis 316
loheri Merr. 317, 318
longicuspis Gandoger 319
longiflora Merr. 302, 326
lutea Cammerl. 317, 318
lutea (non Cammerl.) Merr. 322.1323
macgregorii Merr. 326
macrodendron Gilg & Bened. 31
macroscypha Baker 302, 328, 329*, 330, 331
macroscypha (non Baker) Hei- ne 329
maingayi C.B.Clarke 311, 315
malabarica Bl. 316
malabarica Wight, non Bl. 316
malayana Mart. 311, 315
Merr.
998
melanochlora Gilg & Bened. 333, 335
minor (non BI.) Heine 311
minor Reinw. ex BI. 316, 318
monantha Migq. 331
336
morindaefolia (Reinw.) BI. 293, 311 var. robusta BI. 311 morindifolia = morindaefolia 311
muelleri Bth. 300, 332, 333 negrosensis Elm. 322, 323 nonok Elm. 327, 328 novae-guineae Cammerl. 335 nov. sp. aff. longiflora 325 oblonga K. & G. 317, 318, 954 obovata Bl., non Wall. 320, 323, 324 obovata (non Wall.) King 319, 320, 323 obovata (non Wall.) Miq. 321, 323 var. brevicalyx Bakh. f. 322, 323 var. latifolia Mig. 321, 323 obovata Wall. 315, 317 var. brevicalyx Bakh. f. 316 var. gardneri Clarke 315 var. latifolia Miq. 315 var. papuana F.M.Bailey 316, 335 obovato-javana Bl. 293, 320, 323 var. bebeak Bl. 320, 323 var. latifolia Mig. 320, 323 obtusifolia Merr. & Perry 332 oxyphylla Mig. 316, 318 pachyclados K.Sch. 336 pachypoda Gilg & Bened. 335 papuana Merr. & Perry 333 pauciflora (K. & G.) Ridl. 311, 315 peekelii Gilg & Bened. 335 pendula Merr. 310*, 311, 315 peregrina (Reinw.) Bl. 307 picrophloea Bl. 303 plumeriaeflora DC. 320, 323 plumeriaeflora (non DC.) Koord. 325 plumeriaefolia = plumeriaeflo- ra 320, 325 pluvialis S.Moore 335 prainii Gandoger 317 pseudoelliptica Kaneh. & Ha- tus. 304 pusilliflora Bakh. f. 304 pyriformis 8.Moore 317, 318 racemosa Jack ex Wall. 293, S005. 302, 3107 Siiee3i12=, 313*, 314*, 316, 954 var. coarctata (Bl.) Miq. 311 var. grandis Wall. ex A.DC. 311
FLORA MALESIANA
[ser. I, vol. 6&
var. pauciflora K. & G. 311
rahmatii Merr. 317
resinosa Leenh. 302, 331
ridleyi Gandoger (non K. & G.) 307
ridleyi K. & G. 303, 316, 320, 321*, 954
robusta Bl. 311
rodatzii K.Sch. & Laut. 311
rosenstromii C.T.White 335
rostrata Bl. 316
rotundifolia Ridl. 331
sair Gilg & Bened. 335
salomonensis Gilg & Bened. 335
salticola Leenh. 834453354550
samoensis Gilg & Bened. 335
sasakii Hayata 317, 318
schlechteri Gilg & Bened. 335, 954
scholaris Blco 311
sororia J.J.S. 307
sp. Endert 307
sp. Kaneh. & Hatus. 332
sparei M.R.Hend. 317, 318
spatiosa S.Moore 322, 323
speciosa (Bl.) Bl. 303
speciosa (non Bl.) Ridl. 307
spicata Baker 310*, 311, 315
splendens Bl. 316
stenophylla Becc. 311, 315
suaveolens Cammer]. 333
subreticulata Bl. 311
sumatrana Miq. 304
tacapala Leenh. 301*, 323, 325, 326 ssp. ceramensis Leenh. 325,
326
ssp. gracilis Leenh. 325 ssp. tacapala 325
ternatana (non Miq.) Holth. & Lam 324
ternatana Miq. 317, 318
tetragona Span. 336
teysmannii Cammerl. BIS
thwaitesii F.v.M. 311
truncata Bl. 302, 316, 320, 325
tubulosa Bl. 302, 320
umbelliflora Gilg & Bened. 301, 303, 309, 333
uniflora (non Merr.) Heine 329
uniflora Merr. 331
vaginata K. & G. 322, 323
valida Miq. 304
vitiensis Gilg & Bened., non Seem. 335
vitiensis Seem. 336
volubilis Wall. 311
wallichiana Bth. 307, 309
woodiana F.v.M. 301, 325, 326
zeylanica = ceilanica 315
802% 333;
311,
Fagus 49
Feroniella pubescens Tanaka 208
Flacourtiaceae 5: 1-106, 565; 6: 4, 943-944, 968
Flagellariaceae 4: 245-250; 5: 557
Flemingia Hunter 299, 301, 303 fragrans Hunter 307, 309
Florschuetzia trilobata 975
Fli(e)ggea serrata Miq. 236
Forsellesia 227
Fructus musculiformis Rumph. 367, 368
Fumariaceae 62, 63
Funis toaccae Rumph. 318
Gaertnera Lamk 387 Galium 253, 294 Gardneria Wall. 295, 296, 297, 361 fagraeacea F.v.M. 332, 333 ovata Wall. 362*, 363, 959 wallichii Wight ex Wall. 363 Garuga floribunda Decne 917 Gaultheria Kalm ex L. 470, 471, 472, 473, 474, 677, 678 map, 696, 697 sect. Amphicalyx Endl. 696 abbreviata J.J.S. 678, 682*, 683 acroleia Sleum. 678, 681, 682* arfakana Sleum. 679, 689 atjehensis J.J.S. 678, 680*, 681 bandongensis Zoll. 694, 695 barbulata Sleum. 678, 682 benguetensis Copel. f. 679 berberidifolia Sleum. 679, 686, 687* blumei F.v.M. 778 borneensis Stapf 678, 679, 680* calyculata Wernh. 691 celebica J.J.S. 679, 686 var. celebica 686 var. petiolata J.J.S. 686 ciliolata (Hook. f.) F.v.M. 708 crenulata Kurz 691 crenulata (non Kurz) J.J.S. 694 cumingiana (non Vidal) Merr. 695 cumingiana Vidal 694 dialypetala Sleum. 678, 680*, 681 fragrantissima (non Wall.) K. & G. 690 fragrantissima Mor. 688 var. punctata (Bl.) J.J.S. 688 fragrantissima Wall. 688 var. papuana J.J.S. 691 gracilescens Sleum. 679, 687 heterophylla (Bl.) Endl. ex Hassk. 738 hirta Ridl. 679, 691
(non Wall.)
Sept. 1972]
intermedia J.J.S. 679, 690 itoana Hayata 679, 680 kemiriensis Sleum. 678, 682*, 683 latifolia (B\.) Endl. ex Hassk. 738 latifola Endl. ex Zoll. 738 laxiflora Diels 694 leiotheca Sleum. 692 leucocarpa BI. 470, 677, 679, 690, 692, 693* f. glabra J.J.S. 692 f. pubescens J.J.S. 695 var. hirta Val. ex J.J.S. 679, 692, 695 var. leucocarpa 692, 694* f. cumingiana (Vidal) Sleum. 692, 694 f. leucocarpa 692 f. melanocarpa J.J.S. ex Amsh. 692, 695 f. scandens Hochr. 692,
694 var. melanocarpa J.J.S. ex Steen. 695 var. melanocephala Steen. 695
var. papuana Becc. 689 var. psilocarpa (Copel.) Sleum. 692, 695 var. seminuda J.J.S. 694 losirensis Sleum. 678, 682%, 683 luzonica A.Gray 720 malayana Airy Shaw 679, 682*, 690 mundula F.v.M. 470, 678, 680*, 684, 778 var. mundula 684, 685%, 686 var. setifolia Sleum. 684, 686 var. tanythrix Sleum. 684, 686 notabilis Anthony 681 novaguineensis J.J.S. 679, 680*, 695 var. novaguineensis 695 var. pascua Sleum. 695 nummularioides D.Don 677, 678, 680*, 684 pernettyoides Sleum. 680*, 695 pilosa (Bl.) Zoll. 729 psilocarpa Copel. f. 695 pullei J.J.S. 679, 691 var. leiotheca Sleum. 692 var. pullei 691 punctata BI. 677, 679, 688, 689*, 694 repens Bl., non Raf. 684 solitaria Sleum. 678, 680*, 681 sp. Vidal 782 tanythrix Sleum. 686
(Sleum.) (Sleum.)
679,
(Sleum.)
Index to scientific names
var. setifolia Sleum. 686 trichophylla (non Royle in DC.) Hassk. 684 viridiflora Sleum. 679, 691 yunnanensis (Franch.) Rehd. 694 (Gay) Lussacia 761 Gaylussacia Mor. 873 lanceolata Bl. 761 Gelonium Roxb. ex Willd. 944 Gelsemium Juss. 293, 294, 295, 296, 297, 343, 954 elegans (Gardn. & Champ.) Bth. 343, 344* rankinii 954 sempervirens 343 sumatrana (Bl.) Boerl. 343 Geniostoma J.R. & G.Forst. 293, 295, 296, 297, 369 acuminatissima Merr., non Gilg & Bened. 373 acuminatissimum Gilg & Be- ned. 371 acuminatum Wall. 373 acutifolium Hiern 372 antherotrichum Gilg & Bened. Sil arboreum O.K. 371 var. laevigatum(BI.)O.K. 371 archboldianum Merr. & Perry 373 arfakense Kaneh. & Hatus. 371, 373, 959 australianum F.v.M. 371 avene Val. 371, 372 batanense Mertr. 371 brevipes Merr. 371 caulocarpum K.Sch. 371 celebicum Val. 371 crassifolium Bth. 373 cumingianum Bth. 371, 372 dallmannense Kaneh. & Ha- tus. 371 dasyneurum Gilg & Bened. 371 fasciculata Quis. & Merr. 371 gilgii Merr. & Perry 373 haemospermum Steud. 293, Sil. ove var. angustifolia Bl. 371 var. elongata BI. 371 var. erosa Bl. 371 var. laevigata Bl. 371 var. rugulosa Bl. 371 lanceolatum (non Bojer) Z. & M. ex Mig. 371, 372 lancilimbum Merr. 371 lasiostemon Bl. 371 var. moluccanum Bl. 371 laxa Elm. 371 longipes Merr. 371 micranthum DC. 371 mindanaense Elm. 371
Gaylussacia
elliptica Zoll. &
999 miquelianum K. & V. ex Val. 293, 371 moluccanum Val. 371 montanum Zoll. & Mor. 371, 372 nigrescens (Blco) Merr. 371, B75 oblongifolium K. & V. ex Val. 293, 371 obtusum Merr. & Perry 372 oleifolium S.Moore 373 pachyphyllum Merr. 371 philippinense Merr. 371, 373 psychotrioides Gilg & Bened. 371 pulgarense Elm. 371, 372, 373 pullei Cammerl. 371, 373 ramosii Merr. 371 randianum Merr. 36973715 372 reticulata Bl. 371 rupestre Forst. 293, 369%, 370, 371, 373, 959 schlechteri Gilg & Bened. 371 stenophyllum Gilg & Bened., non Merr. 373 stenophyllum Merr. 371 weinlandii K.Sch. 371, 372 Gentiana quadrifaria 953 Gentianaceae 294 Gentianales 297 Geraniaceae 195, 445-449 Geraniales 195 Geranium L. 445 sect. Australiensia 445 sect. Chilensia 445 sect. Columbinum 445 sect. Striata 445 affine (non W. & A.) Britten 447 ardjunense Zoll. & Mor. 447 australe (non Nees) Allan 448 carolinianum 445 clemensiae Knuth 446 dissectum (non L.) Hemsl. 446 var. glabratum Hook. f. 448 glabratum (Hook. f.) Small ex Hanks & Small 448 homeanum Turcz. 446, 448 macrorrhizum L. 445 microphyllum Hook. f. 446 monticola Ridl. 446 nepalense (non Sweet) Backer 448 nepalense (non Sweet) Doct.v. Leeuwen 447 nepalense Sweet 448 papuanum Ridl. 446 var. alpestris Ridl. 446 parviflorum Willd. 448 philonothum DC. 446 pilosum [non (Sol.) Forst.] F.v.M. 446 potentilloides L’Hérit. ex DC. 446, 447*
& Perry
1000
FLORA MALESIANA
var. ardjunense (Zoll. & Mor.) Carolin 447 var. potentilloides 447 sarawaketense Knuth 446 Girocarpus (sphalm.) = Gyro- carpus 152 Glaphyria nitida Jack 814 Glaux 173 Glinus lotoides L. 915 Glochidion 420 Glosocomia D.Don 118 Glyptopetalum Thw. 228, 232, 252, 254, 264, 391, 392, 421, 930 acuminatissimum Merr. 256, 257 calocarpum 256 calyptratum 256 euonymoides Merr. 256 euphlebium (Merr.) 256, 257, 258 fengii (Chun & How) Ding Hou 256 glandulosum Merr. 251
Merr.
loheri Merr. 256, 257 marivelense (Elm.) 255*, 256, 258, 421 var. euphlebium Merr. 258 palawanense Merr. 256, 257 quadrangulare Prain ex King DIAS 259%, LIOVLOI ZO remotinervium Merr. 258 reticulatum Merr. 258 scortechinii King 248 zeylanicum Thw. 256, 257 var. brevipedicellatum Ding Hou 255*, 256, 257 var. zeylanicum 258 Gnetaceae 4: 336-347; 6: 944— 949
Merr.
Gnetum L.
sect. Cylindrostachys Mef 945, 946 sect. Gnemomorphi Mgf 947 sect. Gnetum 944, 946, 947 subsect. Eugnemones Megf 944, 946 acutatum Mgf 947 acutum Mef 945, 947 arboreum Foxw. 945, 946 costatum K.Sch. 944, 946 cuspidatum BI. 945, 947, 948*, 949 diminutum Mef 945, 947, 949 globosum Mef 945, 947, 949 gnemon L. 944, 946 var. brunonianum (Griff.) Mef 944, 946 var. gnemon 944, 946 var. gracile Mgf 944, 946 var. griffithii (Parl.) Mef 944, 946 var. ovalifolium (Poir.) BI. 945, 946 var. tenerum Mef 944, 946
[ser. I, vol. 66
gnemonoides Brongn. 945, 946, 947, 949 klossit Merr. 945, 947, 949 latifolium BI. 945, 946 var. funiculare (BI.) Mef 945, 946 var. latifolium 945, 946 var. laxifrutescens (Elm.)
Mef 945, 946 var. minus (Foxw.) Megf 945, 946
leptostachyum BI. 945,946,949 var. abbreviatum Mef 945, 946, 949 var. leptostachyum 945, 946 var. robustum Megf 945, 946, 949 loerzingii Mgf 945, 947, 949 macrostachyum Hook. f. 945, 947 microcarpum BI. 945, 947 neglectum BI. 945, 946, 947 oxycarpum Ridl. 946, 947 raya Mef 945, 947 ridleyi Gamble ex Mgf 945, 946, 947, 949 tenuifolium Ridl. 945, 946
Gnidia 3, 4, 30
oppositifolia (non L.) Blco 48 Philippinica Meisn. 48
lucidulus Airy Shaw 977, 981
macrophyllus (Miq.) Airy Shaw 15, 977
maingayi Hook. f. 977, 978
micranthus Airy Shaw 977, 978, 979
nervosus Airy Shaw 977, 978, 981
nobilis Airy Shaw 977, 978, 981
pendulus Airy Shaw 978
reticulatus (Elm.) Merr. 977, 978, 982
spectabilis Airy Shaw 977, 982
stenosepalus Airy Shaw 978, 980
velutinus Airy Shaw 977
xylocarpus Airy Shaw 977, 978
Goodenia J.E.Smith 950
koningsbergeri (Back.) Back. ex Bold. 950
pumilio R.Br. 950*
purpurascens R.Br. 950
Goodeniaceae 5: 335-344; 6: 108, 109, 949-952
Guioa koelreuteria (Blco) Merr. 226
Guttiferae 294, 336
Gymnosporia Hook. f. 229, 238,
Gomphostigma Turcz. 295, 296, 953 Gomphrena celosioides Mart. 917 Gonus Lour. 209 amarissimus Lour. 211 Gonystylaceae, see Thymelaea- ceae Gonystylus Teysm. & Binn. 1, 3, 5, 6, 976-982 sect. Auxanthus Airy Shaw 977, 981 acuminatus Airy Shaw 977 affinis Radlk. 978, 979 areolatus Domke ex Airy Shaw 977, 981 augescens Ridl. 977, 981 bancanus (Miq.) Kurz 2, 5, 15, 978, 979 borneénsis (v.Tiegh.) Gilg 978, 979, 980, 981 brunnescens Airy Shaw 978 calophylloides Airy Shaw 977, 979, 982 calophyllus Gilg 977, 979, 982 confusus Airy Shaw 978 consanguineus Airy Shaw 977, 978, 979, 980 costalis Airy Shaw 977, 980 decipiens Airy Shaw 978, 980 forbesii Gilg 977 glaucescens Airy Shaw 978, 980 keithii Airy Shaw 987, 980, 982
240, 241 ambigua Vidal 241, 242 curtisii King 240 diversifolia Maxim. 241, 242 emarginata Thw. 241 inermis Merr. & Perry 241 montana Bth. 241 var. /ittoralis Back. 241 var. parvifolia Pitard 242 neglecta Wall. ex Laws. 237, 420 nitida Merr. 242 senegalensis (Lamk) Loes. 241 spinosa (Blco) Merr. & Rolfe 241 var. parva Merr. & Rolfe 241 trigyna 238
Gynandropsis DC. 99, 100
affinis Bl. 101
gynandra Briq. 101
pentaphylla (nor. DC.) Blco 101
pentaphylla DC. 101
speciosa DC. 101
Gynotroches axillaris Bl. 967 Gyrinops Gaertn. 2, 3, 4, 5, 39,
41 map
caudata (Gilg) Domke 39, 40*, 42
decipiens Ding Hou 4, 39, 40*, 41
ledermannii Domke 39, 41
ledermannii (non Domke) Merr. & Perry 42
Sept. 1972]
moluccana (Miq.) Baill. 4, 39, 40*, 41
podocarpus (Gilg) Domke 39, 40*, 42
salicifolia Ridl. 39, 40*, 41
sp. Hall. f. 42
versteegii (Gilg) Domke 2, 39, 40*, 42
walla Gaertn. 39
walla (non Gaertn.) Koord. 42
Gyrinopsis Decne 6, 12
acuminata Merr. 11
brachyantha Merr. 10, 11, 12
brachyantha (non Merr. 1912) Merr. 11
citrinaecarpa Elm. 10
cumingiana Decne 15 var. pubescens Elm. 15
decemcostata Hall. f. 15
grandifolia Quis. 13
parvifolia Quis. 12
pubifolia Quis. 15
salicifolia Quis. 41
urdanetense Elm. 10
Gyrocarpus pendulus Blco 152
Haematosperma arborescens Hassk. 371 Haemodoraceae 5: 111-113 Haemospermum Reinw. 369 arboreum Reinw. 371, 372 Halophila 171 beccarii Aschers. 953 decipiens Ostenf. var. pubescens Hartog 953 minor (Zoll.) Hartog 953 ovata Gaud. 953 Hamamelidaceae 5: 363-379; 6: 145, 195, 472, 952 Hannoa Planch. 198, 202 Harrisonia R.Br. 193, 195, 196, 207 bennettii Benn. 208 var. multijuga F.-Vill. 208 var. paucijuga F.-Vill. 208
brownii A.Juss. 207, 208* map
citrinaecarpa Elm. 208
paucijuga Oliv. 208
perforata (Blco) Merr. 207,
208*, 209 map Hasskarlia Meisn. 51 Hebonga Radlk. 215 mollis Radlk. 219 obliqua Radlk. 219 siamensis Radlk. ex Craib 219 Hedraianthera F.v.M. 390, 393, 394, 930 porphyropetala F.v.M. 393 Hedyotis 379 Heimerlia Skottsb. 457, 459 brunoniana Skottsb. 461 Heimerliodendron Skottsb. 457, 459 brunonianum Skottsb. 461
Index to scientific names Helicia rufescens Prain 965 Heliciopsis rufidula Sleum. 965 Helobieae 157 Hemiphragma heterophylla
Wall. 960 Hemiscolopia Sloot. 944
Herbaceae Hutch. 62, 451 Hermicium (sphalm.) = Hermi- dium Hermidium alipes S.Wats. 450 Hexaspora C.T.White 240 pubescens C.T.White 229 Heynea Roxb. 226 Hibbertia 49 Hippobroma G.Don 139, 140 longiflora G.Don 140 Hippocratea 231, 389, 397 arborea Roxb. 420 beccarii Tuyn 402 bojeri Tul. 420 brachystachys Ridl. 400 cassinoides DC. 402 cumingti Laws. 399, 400 ellipticarpa Merr. 402 ferruginea King 402 glaga Korth. 402 grahami Wight 402 hasseltiana Miq. 398 indica (non Willd.) Span. 402 indica Willd. 401, 402 f. timorensis Miq. 401 var. evonymoides Bl. 401 lawsonii Elm. 400, 417
trimera (Boerl.)
macrantha auct. non Korth. 400
macrantha (non Korth.) King 400
macrantha Korth. 398 macrantha (non Korth.) Rolfe 399 maingayi Laws. 262 maingayi (non Laws.) Vidal 269 megalocarpa Merr. 402 nigricaulis Ridl. 400 obtusa Ridl. 406 obtusifolia (non Koord.-Schum. 400 obtusifolia (non Roxb.) Merr. 406 pauciflora DC. 400 f. minor Miq. 400 f. novoguineensis Miq. 400 rigida Span. 400 salacioides Korth. 402, 404 sogerensis Baker f. 398 sp. Burk. & Haniff 406, 407 timorensis Span. 421 trichopetala Merr. 399 trilobulata Ridl. 398 volubilis (non L.) Blco 401, 402 zippeliana Miq. 402 Hippocrateaceae 228, 229, 230,
Roxb.)
1001 231, 389, 390, 932 Hippocraticeae Juss. 389 Hiptage /awsonii Elm. 417 Holosteum hirsutum L. 915 Homalanthus giganteus 152 Homalium Jacq.
subg. Blackwellia (Lamk) Warb. 944
sect. Blackwellia Bth. 944
subg. Pythagorea (Lour.) Sleum. 944
sect. Pythagorea (Lour.) O.K. 944
Hornera Jungh. 48 glomerata Jungh. 48 umbellata Jungh. 48 Hottonia sessiliflora Vahl 965 Huertea 49 Hydnocarpus 963 Hydrilla verticillata 164 Hydrocaryaceae 4: 43-44; 6: 982; see Trapaceae Hydrocharitaceae 5: 5695163) 15Se 1752 Hydrophyllaceae 4: 207-209 Hymenanthes Bl. 656 Hypericineae 49 Hypsagine Jack ex Burk. 421 Hyptiandra Hook. f. 202
381-413,
laera Copel. f. 740 lanaensis (Merr.) Copel. f. 745 loheri (Merr.) Copel. f. 741 lucida (Merr.) Copel. f. 742 Ignatia amara L. f. 347, 349 Ignatiana philippinica Lour. 347, 349 Ilex racemosa Oliv. 290 Indotristicha v.Royen 963 malayana Dransf. & Whitmo- re 963, 964* Inocarpus 201, 226 Ipomoea sect. Ipomoea 940 sect. Pharbitis (Choisy) Gri- seb. 940 acuminata Ruiz & Pay. 941 acuminata (Vahl) R. & Sch. 939, 940, 941 angulata Ortega 941 congesta R.Br. 939, 940, 941 dasysperma Jacq. 941 digitata L. 940, 941 gracilis R.Br. 940, 941 littoralis Bl. 940, 941 mauritiana Jacq. 940, 941 maxima (L. f.) Don ex Sweet 940, 941 < multifida (Raf.) 940, 941 ochracea (Lindl.) G.Don 939, 941 ochroleuca 939, 941 pes-caprae (L.) R.Br. 941 plebeia R.Br. 940
Shinners
(sphalm.) Span.
1002
FLORA MALESIANA
[ser. I, vol. 68
ssp. africana Meeuse 940 ssp. indica Verdc. 940 ssp. plebeia 940 purpurea Roth 941 riparia G.Don 940, 941 rubens Choisy 940, 941 sepiaria Koen. ex Roxb. 940, 941 < sloteri 939 tuba (Schlechtend.) 941 tuberculata Ker-Gawl 941 Irvingella v.Tiegh. 223 harmandiana v.Tiegh. 225 malayana v.Tiegh. 225 oliveri v.Tiegh. 225 Irvingia Hook. f. 193, 194, 195, 196, 201, 223, 969, 972 harmandiana Pierre ex Lecom- te 225 longipedicellata Gagnep. 225 malayana Oliv. ex Benn. 223, 224*, 225*, 226 map oliveri Pierre 225 Irvingiaceae 195, 972 Iso Kuroggi Kaempf. 253 Tsolobus DC. 121 caespitosus Hassk. 130 campanuloides DC. 130 kerii DC. 130 radicans DC. 130 roxburghianus DC. 130 Tsotoma Lindl. 108, 139, 140, 141 longiflora Presl 140 runcinata Hassk. 140 Itea 280 Ixonanthaceae 970, 972 Ixonanthes Jack 971, 972
G.Don
Jacquemontia Choisy 939 browniana Ooststr. 939
Jasione 108
Jaundea Gilg 934
Johnia Roxb. 404 sumatrana Jack 421
Juglandaceae 143-154, 953
Juglandicarya Reid & Chandler 143
Juglans 143 plerococca = pterococca 154 pterococca Roxb. 152, 154 regia 145 rupestris Engelm. 143 sieboldiana 143
Juncaceae 4: 210-215; 5: 557; 6: 953
Juncaginaceae 4: 56-57; 5: 554
Juncodes — silvaticum (Huds.) O.K. 953
Juncus effusus L. 953 prismatocarpus R.Br. 953
Kaernbachia Schltr 49, 51 brachypetala Schltr 59 pentandra Schltr 59
Kalmia 472 Karin-Njoti Rheede 199 Kelleria Endl. 43 papuana Domke 44 Kentia Steud. 299, 301, 309 morindaefolia Steud. 311, 315 Kippistia Miers 389 Klainedoxa 194, 195, 972 Kokoona Thw. 228, 231, 232, 258, 260 map, 264, 389, 391, 392, 930, 932 coriacea King 260, 261 lanceolata Ridl. 261 littoralis Laws. 231, 260, 261 luzoniensis Merr. 262, 402 ochracea (Elm.) Merr. 259%, 260 ovatolanceolata 259*, 260, 261 reflexa (Laws.) Ding Hou 231, 259*, 260, 262 scortechinii King 261 scortechinii (non King) Steen. 261 sessilis Ding Hou 260, 261 zeylanica 260 Kokoonia = Kokoona 260, 402 Kuhlia H.B.K. 301 Kuhlia Reinw. ex BL, H.B.K. 299, 301, 309 morindaefolia Reinw. ex BI. Sia sis Kurrimia Wall. ex Arn. 230, 280 archboldiana Merr. & Perry 284 bipartita Laws. 283 calophylla Wall. 283 gracilis Vidal 284 indica Gamble 283 luzonica Vidal 282 maingayi Laws. 283 minor Rid]. 282 paniculata Wall. ex Arn. 282 pulcherrima (non Wall.) Baker f. 420 pulcherrima Wall. ex Laws. 283 robusta Kurz 282, 283 var. roxburghii Pierre 283 var. thorelii Pierre 283 Kurrimia Wall. ex Meisn. 280 Kurrimiaceae 230
Ruidly228,
non
Labordia Gaud. 295, 296, 370 Lachnolepis Miq. 39 moluccana Mig. 39 Lachnopylis 296 Lafoénsia Vand. 975 Lagansa alba Rumph. 103 rubra Rumph. 101 Lagerstroemia L. 975 Lasianthus furcatus (Miq.) Bre- mek. 373 Lasiolepis Benn. 207 bennettii Planch. 208
var. @ paucijuga (Benn.) Planch. 208 var. 2B multijuga (Benn.) Planch. 208 paucijuga Benn. 208 Lasiosiphon 4 scandens Endl. 25 Launaea pinnatifida 223 Lauraceae 194 Laurentia Mich. ex Adans. 109, 110, 139, 141 sect. Isotoma (R.Br.) Endl. 107, 140 subg. Jsotoma Peterm. 140 gasparrinii (Tineo) Strobl. 140 longiflora (L.) Peterm. 108, 138*, 140 var. runcinata (Hassk.) Wimmer 140 Lechenaultia _ filiformis 951 Ledum 472 Leea 52 Legousia Durand 928 speculum-veneris (L.) Fisch. 141, 928 Leguminosae 216, 226, 933 Lemnopsis Zoll. 171 Leptopteris Bl. 343 sumatrana Bl. 343 Leucopogon R.Br. 424, 425 sect. Anacyclodon (Jungh.) Miq. 425 sect. Stypheliopsis Miq. 425 abnormis Sond. 433 acuminatus R.Br. 432 acuminatus (non R.Br.) Du- perrey 433 colensoi Hook. f. 428 hookeri Sond. 428 javanicus de Vriese 431 lancifolius Hook. f. 433 malayanus Jack 426 var. o Kurz 426 var. moluccanus (non Scheff.) Kurz 426, 433 moluccanus Scheff. 433 obovatus Fawe. 429, 432 obtusatus Hook. f. 428 ophirensis Griff. 943 papuanus C.H.Wright 443 Philippinensis (Merr.) Hoso- kawa 429 suaveolens Hook. f. 428 Leucosmia Bth. 15 Leucothoé 472 Libocedrus 333 Lightfootia L’Hérit., non Sw. 109, 111; 1123918 gracilis Miq. 115 var. lavandulaefolia Miq. ib iey Lightfootia Sw. 113 Ligia Fasano 28 Lignosae 62
R.Br.
Sept. 1972]
Index to scientific names
Lignum colubrinum _ timorense Rumph. 350 Limnophila sp. 965 Limonia pubescens Hook. f. 208 Linaceae 970 subfam. Ixonanthoideae 970 tribe Erythroxyleae 195 Linaria 294 Lindernia subulata R.Br. 960 Linociera sp. 280 vitiensis A.C.Smith 336 Linostoma Wall. ex Endl. 2, 4, 5, 26, 982 sect. Eulinostoma Meisn. 27 sect. Psilaea (Miq.) Hall. f. 27 subg. Linostoma Kurz 23 subg. Nectandra {(non Berg.) Roxb.] Kurz 27 leucodipterum Hall. f. 27 longiflorum Hall. f. 2, 26*, 27, 982 pauciflorum Griff. 2, 26*, 27, 982 scandens Kurz 25 Lissanthe R.Br. 424 montana R.Br. 429 Lobelia L. 107, 108, 109, 110, MOM 1 122)) 129591355137, 140, 928 sect. /sotoma R.Br. 140 affinis Wall. ex G.Don 128, 129 var. lobbiana Clarke 128 alata Labill. 136 aligera Haines 129 alsinoides Lamk 107, 122, 123, 12659127amap; 128; 130, 1527 anceps L. f. 129 anceps Thunb., non L. f. 136 angulata Forst. 108, 110, 123, 1SS51347°1135* archboldiana (Merr. & Perry) Moeliono 123, 131*, 928 arenarioides DC. 129 arfakensis Gibbs 134 aromatica Moon ex Wight, nomen 123 barbata Warb. 128 beddomeana Wimmer 124 hegonifolia Wall. 133 bialata Merr. 129, 130 borneensis (Hemsl.) Moeliono 122, 133 map, 928 brachyantha Merr. & Perry Ay I1si: 182* caespitosa Bl. 130 campanuloides Thunb. 130 camptodon Wimmer 124 chevalieri Danguy 127 chinensis (non Lour.) Hance 127 chinensis Lour. 108, 123, 130* cliffortiana L. 136
Wall. ex
colorata Wall. 123, 126 conferta Merr. & Perry 123, 185" cordigera Cav. 136 decurrens Roth 129 dichotoma Migq. 129 var. aligera Wimmer 129 var. pilosella Wimmer 129 donanensis v.Royen 928 doniana Skottsb. 124 dopatrioides Kurz 127, 128 dubia de Vriese 115 epilobioides Wimmer 124 var. sarasinorum Wimmer 124 erecta Hook. f. & Th. 123 erinoides (non L.) Thunb. 130 erinus L. 136 erinus (non L.) Thunb. 130 eryliae Fischer 124 eurostos Voigt 123 excelsa Lesch. ex Roxb. 123 filiformis (non Lamk) Cav. 126 var. /uzoniensis Pers. 126 fossarum Wimmer 124 graminea Lamk 136 griffithii Hook. f. & Th. 127 var. dopatrioides 127 var. genuina 127 haenkeana DC. 136 hainanensis Wimmer 127 hederacea Cham. 134 heyneana = heyniana 127, 129 heyniana R. & S. 123, 129, 130 map, 132*, 176 horsfieldiana Mig. 134 hosseusii Wimmer 127 var. villosa 127 inconspicua Rich. 129 inflata L. 108 javanica Thunb. 133 /aurentia L. 140 laxiflora H.B.K. 136 var. nelsonii McVaugh 136 leucanthera Kerr 124 linnaeoides (Hook. f.). Petrie 134, 135 var. brevipilis 134 littoralis R.Cunn. ex A.Cunn. 134 lobbiana Hook. f. & Th. 128 longifiora L. 140 longifolia DC. 136 luzoniensis (Pers.) Merr. 127, 128 micrantha Hook. 129 microcarpa Clarke 127 montana Reinw. ex BI. 107, 108, 122, 132*, 133 map nelsonii Fernald 136 nicotianaefolia Roth ex R. & SHO, (NO 22.9123: 124". 125* map, 126
126,
1003 var. bibarbata Wimmer 123 var. macrostemon Skottsb. 123 var. mollis Elm. 123 var. nicotianaefolia mer 123 var. trichandra Clarke 123 nummularia Lamk 133 obliqua Ham. ex D.Don 133 palustris Kerr 124 paradoxa Wimmer 134, 135 philippensis Skottsb. 124 plumierii L. 951 pratiana Gaudich. 133 pumila Burm. f. 136
Wim-
purpurescens Wall., nomen 123
pyramidalis Wall. 123 var. B 123
radicans (non Thunb.) Hos- seus 127 radicans Thunb. 130 reinwardtiana DC. 129 reniformis Cham. 134 robusta Wall. ex Voigt 123 rosea Wall. 123 rotundifolia Banks & Sol. ex Hook. f. 134 rugulosa Graham 134 sebae DC. 129 seguinii Léveillé & Van 124 sinaloae 140 sp: Gril 127 sp. Steen. 126 stimulans Ham. ex D.Don 123 stipularis Roth 127, 128 subcuneata Miq. 128 subincisa Wall. ex DC. 129 subracemosa Miq. 129 var. rigidior 129 succulenta Bl. 128, 129 f. glabra 128 var. lobbiana 128 sumatrana Merr. 122, 126* syphilitica L. 136 taccada Gaertn. 951 terminalis Clarke 127, 128 tetragona Bl. 136 thorelii Wimmer 127 trialata Ham. ex D.Don 126, 127, 129 var. lamiifolia 129 triangulata Roxb., nomen 127 trichandra Wight 123 trigona (non Roxb.) Hook. f. & Th. 129 trigona Roxb., nomen 126 wallichiana Hook. f. & Th. 123 wallichii Steud. 123 zeylanica L. 122, 123, 128, 129 var. affinis 128 zeylanica (non L.) Moon 129 var. aligera Haines 129 var. parviflora Danguy 129
1004
FLORA MALESIANA
[ser. I, vol. 68
var. walkeri Clarke 129 Lobeliaceae 108, 109 Locandi Adans. 199 Locandia O.K. 201
glandulifera Pierre 201
indica O.K. 201
madagascariensis O.K. 201
mekongensis Pierre 201
merguensis Pierre 201
pendula Pierre 201 Loeseneriella A.C.Smith 262,
390, 391, 397
cumingii (Laws.) Ding Hou
397, 399
macrantha (Korth.)
Smith 397, 398 map
pauciflora (DC.) A.C.Smith
399*, 400, 417
sogerensis (Baker
Smith 397, 398
zippeliana A.C.Smith 402 Logania 295, 296 dentata (Elm.) Hayata 960 Loganiaceae 298-387, 953-960 subfam. Buddlejoideae 294 subfam. Loganioideae 294 tribe Antonieae 294, 295, 296, 297 tribe Buddlejeae 294, 295, 296, 297, 953 tribe Desfontainea Leeuwenb. 954 tribe Euloganieae 295 subtribe Buddlejeae 295 tribe Gelsemieae 294, 295, 296, 297 tribe Loganieae 295, 296 tribe Plocospermeae 954 tribe Potalieae 294, 295, 296, 297 tribe Retzieae Leeuwenb. 954 tribe Spigelieae 295, 296, 297 tribe Strychneae 294, 295, 296 Loganiales 294, 297 Lonicera malayana Hend. 929 Lophopetalum Wight ex Arn. DDD Bae 22930 Me Sila 32. 260, 262, 266 map, 391, 392, 930, 932 subg. Solenospermum (Zoll.) Val. 262 beccarianum Pierre 263*, 264, 266 celastroides Laws. 269 celebicus Koord. 270 coriacea Ridl. 261 curtisii King 268, 269 dubium Laws. 261 fimbriatum (non Wight) F.- Vill. 269 fimbriatum Wight 267, 268 floribundum Wight 229, 265, 266, 930 fuscescens Kurz 269 glabrum Ding Hou 265, 266
A.C.
ED ALC:
grandiflorum Arn. 264
havilandii Ridl. 266
intermedium Ridl. 270
javanicum (Zoll.) Turez. 228, 265, 269
javanum = javanicum 269
ledermannii (Loes.) Ding Hou
265, 271 littoralis Ridl. 261 macranthum (Loes.) Ding
Hou 263*, 265, 268, 930 maingayi Ridl. 261 micranthum Loes. 265, 268 multinervium Ridl. 227, 228,
232655 2702 Tlie oblongifolium King 270 oblongum King 269 pachyphyllum King 265, 267 pallidum Laws. 231, 263%,
264, 265, 268
var. curtisii (King) Ridl. 268 paucinervium Merr. 270 penduculatum Ridl. 250 reflexum (non Laws.) King
266 reflexum Laws. 262 rigidum Ridl. 228, 263*, 265,
267 scortechinii King 266 sessilifolium Ridl. 228, 262,
264, 265 sp. Thorenaar 269*, 270 subobovatum King 265, 272 subsessile Ridl. 267 torricellense Loes. 265, 271 toxicum Loher 230, 270 wallichii Kurz 269 wightianum Arn. 231, 264,
265, 267
var. macrocarpum
267 winkleri Loes. 267 Lubinia Vent. 177, 184 lineariloba Pax 184 lubinoides Pax 184 mauritiana Spreng. 184 spathulata Vent. 184 Lunasia 221 Luronium natans (L.) Raf. 953 Lussa Rumph. 209 radja Rumph. 211 (Gay)Lussacia = Gaylussacia 761 Luzula silvatica (Huds.) Gaudin 953 Lyonia Nutt. 469, 472, 474, 674 ovalifolia (Wall.) Drude 673*, 674*, 675 map Lysimachia Tourn. ex L. 173, (WEE IB 17/7/ subg. Lysimachia (£u-lysima- chia) 177 sect. Alternifoliae Knuth 180 sect. Apodanthera 177
Pierre
subsect. Ramosae 177 ser. Evalves 177 ser. Valvatae 177 sect. Nummularia 177, 183 ser. Debiles 177, 183 ser. Japonicae 177, 183 subg. Palladia (Moench) H.- M. 177, 184 acroadenia Maxim. 185, 186 ardisioides Masam. 178, 179 capillipes Hemsl. 177, 178, 179, 180 map. 181, 182, 964 chapaensis Merr. 181 consobrina Hance 185 cuspidata BI. 180, 181 var. glabra Mor. 181 var. glabrescens Knuth 181 var. hispida Knuth 181 cuspidata (non BI.) Hand.- Mazz. 179 debilis Wall. 182 var. minor Baudo 182 var. vulgaris Baudo 182 decurrens Forst. f. 177, 178, 184, 185 map, 965 var. acroadenia Makino 185 var. recurvata Mats. 185 deltoides Wight 183 floribunda Zoll. & Mor. 180, 182 fragrans Hayata 178, 179 garrettii Fletch. 178 glaucophylla Hook. & Arn. 184, 185 japonica Thunb. 178, 181, 182 var. japonica 177, 182, 183 map var. minutissima Masam. 183 var. papuana S.Moore 177, 183 map var. subsessilis Hara 182 var. thunbergiana Hara 182 javanica Bl. 185 keiskeana Miq. 185, 186 klattiana Hance 181 laxa Baudo: 17 eigeceio =: 180 map, 182, 964 lineariloba Hook. & Arn. 184 lobelioides Wall. 186 lubinoides Sieb. & Zucc. 184 maculata R.Br. 182 mauritiana Lamk 177, 178, 183, 184 map montana (Reinw.) Bakh. f. 177, 178, 180, 181 map var. montana 181 var. platyphylla Bentvelzen 181 microphylla Merr. 182, 183 minutissima (Masam.) Ma- sam. 182, 183 multiflora Wall. ex Duby 185 nebeliana Gilg 184 obovata Ham. ex Hook. f. 186
(Merr.)
Sept. 1972]
Index to scientific names
peduncularis Wall. ex Kurz 177, 178, 182 map
pierrei Petitmengin 182
platyphylla Merr. 181
procumbens Baudo 183
ramosa Wall. ex Duby 180, 181, 182 var. grandiflora Franch. 180 var. typica Knuth 178, 180 var. zeylanica Hook. f. 180
rapensis F.B.H.Brown 185
recurvata (Mats.) Masam. 185, 186
rubida Koidz. 184
siamensis Bonato 182, 183
sikokiana Miq. 177, 178, 179* map, 180
simulans Hemsl. 178
sinica Merr. 185
solanoides H.-M. 174
spathulata Schouw 184
spatulata = spathulata 184
suborbicularis Went 182, 183
trichopoda Franch. 181
uliginosa Bl. 181, 182
uliginosa (non Bl.) Klatt 181
vulgaris L. 174
Lythraceae 198, 975
subtribe Diplusodontinae 975
subtribe Lagerstroemiineae 975
Maba hierniana K. & G. 932
Macanea arborea Blco 421
Macgregorianthus Merr. 23 paniculatus Merr. 25
Maerua 61
Maesa membranifolia Mez 237 perakensis Rid]. 290 ramentacea (Roxb.) A.DC.
237
Magnolia 228
Magnoliaceae 228
Malpighiaceae 5: 125-145, 566; 6: 960
Malpighiales 195
Mangifera L. 231 glauca Rottb. 286
Mannia Hook. f. 202
Manotes asiatica Gagnep. 205
Manungala Blico 199 pendula Blco 201
Mastixiodendron — pachyclados (K.Sch.) Melch. 336
Mauduita Comm. ex DC. 199 penduliflora Comm. ex DC.
200
Maurocenia sect. Triceros O.K. 51 pomifera O.K. 58 zollingeri O.K. 56
Maytenus Molina 227, 228, 229, 2305 231-,)232/9238, 240) 243; 391, 392, 930 acuminatus 228
bilocularis (F.v.M.) Loes. 243
boaria Molina 240
crassa Ding Hou 240, 242
cupularis Ding Hou 239*, 240 243
curtisii (King) Ding Hou 239*, 240 dispermus 230 diversifolia (Maxim.) Ding
Hou 231, 232, 240, 241, 242, 391, 930 emarginata (Willd.) Ding Hou 229, 239*, 240, 241, 930 ilicifolia Mart. 229 magnifolia 228 nitida Mart. 242 rapakir Loes. 242 senegalensis (Lamk) Exell 240, 241 spinosa (Gris.) Lourt. & O’- Donell 229 vitiensis (A.Gray) Ding Hou 242 Mazus pumilus (Burm. f.) Steen. 136 Medicia Gardn. & Champ. 343 elegans Gardn. & Champ. 343 Melastomataceae 914 Meliaceae 194, 212, 220 Melichrus R.Br. 424 Menyanthes trifoliata 294 Merremia Dennstedt 939 sect. Eu-Merremia 939 sect. Merremia 939 aniselifolia Ooststr. 939, 940* borneensis Merr. 939 crispatula Prain 939 emarginata (Burm. f.) Hall. f. 939 gangetica (L..) Cufod. 939 peltata (L.) Merr. 939 quinata (R.Br.) Ooststr. 939 tridentata (L.) Hall. 939 ssp. hastata Ooststr. 939 Messerschmidia argentea 197 Micropyxis Duby 176 pumila Duby 176 tenella Wight 176 Microsemma 3 Microtropia Rchb. 272 Microtropis Wall. ex Meisn. DI TEDISS 2295230) 2325272. 275 map, 391, 392, 930 basilanensis Merr. & Freem. 279 bicolor Merr. & Freem. 276 bivalvis (non Wall.) Koord. 277 bivalvis (Jack) Wall. 272, 274, 276 borneensis Metr. 279 chartacea Merr. & Freem. 279 coriacea Wall. ex Ettingsh. 280, 412
& Freem.
1005 curranii Merr. 273, 274, 278 var. obovata Merr. & Freem. 278 var. zambalesensis Merr. & Freem. 278 discolor (Wall.) Wall. 272, 274, 275
elliptica King 273*, 274, 277 fasciculata Quis. & Merr. 279 filiformis King 232, 276, 277, 391 javanica Merr. & Freem. 277 kinabaluensis Merr. & Freem. 274, 277 var. acuminata 277, 278 lanceolata Boerl. & Koord. 280 longifolia Wall. 274 longirostris Merr. & Freem. 277 ophirensis Ridl. 277 ovata Merr. & Freem. 274, 275, 279 pachyphylla Merr. & Freem. 274 pauciflora Boerl. ex Merr. & Freem. 276 peduncularis Rid). 276, 277 Philippinensis Merr. 279 platyphylla Merr. 274, 275, 279 var. ellipticifolia Merr. & Freem. 279 ramiflora (non Wight) Thw. 279 rigida Ridl. 275, 278 rostrata Merr. 279 rubra Elm. ex Merr. & Freem. 279 sterrophylla Merr. & Freem. 277, 278 suborbiculata Merr. & Freem. 279 sumatrana Merr. 275, 278 tenuis Symington 272, 274, 275, 276 tetrameris Ding Hou DBF DTA 215 valida Ridl. 274, 276 vinculans Boerl. & Koord. 277 wallichiana Wight ex Thw. 215, 279 zeylanica Merr. & Freem. 279 Mirabilis L. 450, 451 jalapa L. 450, 451 Mirabilis Rumph. 451 longiflora (non L.) Blco 451 Mischopleura Wernh. 914
272,
Mitrasacme Labill. 293, 295, 296, 297, 375, 378 albomarginata Leenh. 379,
385* alsinoides R.Br. 379, 384 alsinoides (non R.Br.) Clarke 384
FLORA MALESIANA
[ser. I, vol. 66
var. indica (Wight) Hara
384 alsinoides (non R.Br.) Merr. 382 bogoriensis Leenh. 379, 386*, 960 capillaris Wall. 382, 383, 384 chinensis Griseb. 382 connata R.Br. 381 crystallina Griff. 384 elata R.Br. 293, 379, 380, 381* map, 960 var. brevicalyx Leenh. 380*, 381, 960 var. elata 380*, 381 erophila Leenh. 380, 384, 960 exserta F.v.M. 382 indica Wight 379, 384 map, 386, 960 longiflora F.v.M. ex Bth. 380, 381 malaccensis Wight 382, 383 micrantha Domin 382, 383, 384 neglecta Leenh. 379, 386, 960 nudicaulis (non Bl.) Bth. 382, 383, 960 nudicaulis Reinw. ex Bl. 380, 381, 382 paludosa R.Br. 385 polymorpha R.Br. 379, 384 polymorpha (non R.Br.) Clar- ke 382, 383, 384 var. grandiflora Hemsl. 382 var. parishii Clarke 382 pusilla Dalz. 384, 385 pygmaea R.Br. 379, 382, 960 var. grandiflora (Hemsl.) Leenh. 383 var. malaccensis (Wight) Hara 380, 382*, 383, 384 var. parishii (Clarke) Leenh. 383 var. pygmaea 380, 382*, 383*, 384, 960 saxatilis Back. 379, 385, 960 setosa Hance 382 setosa (non Hance) Masam. 382 tenuiflora Bth. 379 trinervis Span. 380, 381, 382, 383 Mitrasacmopsis Jovet 297 Mitreola Boehm. 375 Mitreola L. 296, 375, 959 inconspicua Zoll. & Mor. 375 oldenlandioides (Wall.) G.Don YS, SMT) paniculata Wall. ex G.Don 375 var. lilacina Back. 375, 377 perriert Jovet 377 petiolata (Gmel.) Torr. & Gray 375, 959 reticulata Tirel 960
sphaerocarpa (Leenh.) Leenh. 960 turgida Jovet 377 Mitreola Schaeff. 375 Modagam Rheede 315 Molinadendron Endress 952 Mollugo nudicaulis Lamk 384 Monocelastrus Wang & Tang 231233234 monosperma (Roxb.) Wang. & Tang 234 virens Wang & Tang 234 Monochoria hastata (L.) Solms 964 Monopsis simplex (L.) Wimmer 129 Monotropa L. 472 humilis D.Don 670 uniflora L. 670 Monotropaceae 473 Monotropanthum Andres 670 Monotropastrum Andres 469, 470, 474, 670 humile (D.Don) Hara 670, 671* Moringa domestica Ham. ex Henschel 960 oleifera Lamk 960 Moringaceae 4: 45-46; 5: 554; 6: 62, 960 Mostuea 294, 295, 296 Moya Griseb. 238 spinosa Griseb. 229 Muenteria Walp. 212 Myoporaceae 4: 265-266 Myricaceae 4: 276-279; 6: 143, 146 Myristica Jaurina (non BI.) Hochr. 933 Myristicaceae 194 Myrsinaceae 173, 174, 237 Mystroxylon 229, 284
Najadaceae 157-171 Najadales 157 Najadeae 171 Najas L. 157, 158 subg. Caulinia Aschers. ex Rendle 161, 164 sect. Caulinia A. Braun 161, 164 subg. Eunajas Aschers. ex Rendle 162 sect. Eunajas A.Braun 162 sect. Najas 161, 162 bengalensis Horn af Rantzien 169 brevistyla Rendle 167 browniana Rendle 158, 160*, 161, 163 map, 165 celebica Koord. 167, 168 falciculata A.Braun 165, 166 falciculata’ (non A.Braun) Coert 168 foveolata A.Braun ex Rendle
166 gracillima (A.Braun) Magnus 165, 166 graminea Del. 158, 161, 169, 170 map var. angustifolia Rendle 169 var. delilei Magnus 169 var. graminea 158, 160*, 170 var. minor Rendle 169 var. robusta de Wilde 170 var. (B) ‘enuifolia (R.Br.) A.Braun 167, 168, 169 graminea (non Del.) Mig. 168 graminea (non Del.) Ridl. 169 hagerupi Horn af Rantzien 165 horrida 170 indica (Willd.) Cham. 158, 160*, 161, 164, 166, 167 map, 170 var. gracillima A.Braun ex Engelm. 165 var. macrodictya A.Braun 166 var. rigida A.Braun 166 intermedia Gorski 162 kingii Rendle 166 kurziana Rendle 160*, 161, 163 map, 165 lacerata Rendle 166 lobata Blco 166 major All. 162 B angustifolia A.Braun 162 ¢ intermedia A.Braun 162 malesiana de Wilde 158, 160*, 161, 169 map marina L. 158, 159, 160, 161, 162, 170 var. B 162 var. y 162 var. angustifolia Rendle 162 var. intermedia Rendle 162 var. marina 160*, 162, 163 map var. sumatrana de Wilde 160*, 162, 163 map var. zollingeri Rendle 158, 160*, 162, 163* minor All. 161, 164 var. indica A.Braun 166 minor (non All.) Hook. f. 166 obyvoluta Blco 171 oguraensis Miki 164 palustris Blco 166 pseudograminea W.Koch 167, 168 seminuda Griff. ex Voigt 169 tenuifolia (non R.Br.) auct. 168 tenuifolia R.Br. 158, 161, 167, 168 ssp. pseudograminea (W. Koch) de Wilde 167, 168 var. celebica (Koord.) de
Sept. 1972]
Index to scientific names
Wilde 158, 160*, 168
var. pseudograminea 159*, 160*, 168 map ssp. tenuifolia 167, 168 tenuifolia (non R.Br.) Miq. 166 Naumburgia Moench 174, 175, iL 7/7 Nectandra (non Berg.) Roxb. 26 Neerija dichotoma Roxb. 286, 287 Neojunghuhnia Koord. 785 insignis Koord. 787 Neotrewia arborea Elm. 944 cumingii (M.A.) P. & H. 944 Nephelium 283 Nertera dentata Elm. 960 Neuburgia BI. 293, 295, 296, 297, 363, 959 celebica (Koord.) lLeenh. 364*, 365, 367, 368, 959 corynocarpa (A.Gray) Leenh. 364*, 365, 366, 959 kochii (Val.) Leenh. 365, 366, 959 moluccana (Boerl.) Leenh. 364*, 365, 367, 368*, 369 musculiformis (Lamk) Miq. 367, 368 rumphiana Leenh. 364*, 365, 367, 959 sarcantha (Gilg & Bened.) Leenh. 363, 365, 366, 959 sumatrana (Miq.) Boerl. 369 tuberculata Bl. 365, 367, 368 tubiflora Bl. 365, 367, 368 Neuropeltis Wall. 936, 937 Neuropeltopsis Ooststr. 936, 937 alba Ooststr. 937, 938* Nicodemia Tenore 296, 336, 340 diversifolia Tenore 340 madagascariensis R.N. Parker 340 philippinensis Elm. 341 Nima Ham. ex Juss. 212 Niota Lamk 199 commersonii Pers. 200 globosa Blico 203 lamarckiana Bl}. 200 lucida Wall. 200 pentapetala Poir. 199 polyandra Buch.Ham. ex W. & A. 203 tetrapetala Poir. 200 Norrisia Gardn. 293, 295, 296, 297, 341 maior Soler. 341, 342* malaccensis Gardn. 341, 342*, 373 var. major (Soler.) Ridl. 342 var. malaccense Ridl. 341 var. pubescens K. & G. 341 malaccensis (non Gardn.) Merr. 341 Philippinensis Elm. 341
Nothocnestis Miq. 280 sumatrana Miq. 283 Nothofagus 49 Nothoprotium Miq. 928 Niarvala Rheede 68 Nuxia 294, 295, 296 Nyctaginaceae 450-468 Nypa 965 Nyssaceae 4: 29-31
Ochranthe Lindl. 49, 51 Ochthocosmus Bth. 971, 972 Odyendyea (Pierre) Engl. 198, 202 Olacaceae 420 Oldenlandia brachyphylla Merr. 384 Oleaceae 280, 296, 297, 336 Olus album Rumph. 464 album insulare Rumph. 464 Operculina riedeliana (Oliv.) Ooststr. 939 Ophiorhiza L. 375 mitreola L. 375 mungos L. 375 Ophispermum Lour. 6 Orchipeda sumatrana Miq. 369 Oreamunoa = Oreomunnea 148 Oreomunnea Oerst. 145, 148 Orescia Reinw. 177 montana Reinw. 181 Orobanchaceae 294 Orobanche 294 Oxybaphus 451
Pachistima 228 Paliurus dubius Blco 208 _ perforatus Blco 208 Palmerella 140 Papaveraceae 5: 114-117; 6: 62, 63 Paphia Seem. 878 helenae (F.v.M.) Schltr 879 stenantha Schltr 884 viridiflora Schltr 883 Paracelastrus Miq. 272, 277 bivalvis Miq. 276 wallichianus F.N.Williams 279 Parietales 968 Parophiorrhiza Clarke 375 Paropsia vareciformis (Griff.) Mast. 944 Passerina javanica Thunb. 48 Pedaliaceae 4: 216-221; 5: 557 Pedicellaria Schrank 99 pentaphylla Schrank 101 Pelargonium L’Herit. 445, 448, 449 <asperum Ehrh. ex Willd. 449 crispum 449 graveolens Thunb. 449 radens H.E.Moore 449 Pellacalyx saccardianus Scort. 967
1007
Pelletiera 173
Peltanthera Bth. 294, 295, 296, 953
Pemphis acidula Forst. 198
Pentaceras australis Hook. f. 220
Pentachondra javanica Zoll. 431
Pentaphragma 107, 108, 109, 110
Pentaphragmataceae 4: 517-528 Pentaphylacaceae 5: 121-124, 566 Pentapterygium Klotzsch 878 scortechinii K. & G. 879 Pentaspadon Hook. f. 928 Peracarpa Hook. f. & Th. 107, 109, 110 carnosa (Wall. in Hook. f. &. Th. 110 var. circaeoides 111 circaeoides Feer 111 luzonica Rolfe 111 Pericopsis Thw. 976 Pernettya 472 repens (BI.) Zoll. 684 Pernettyopsis K. & G. 469, 474, 675 breviflora (Ridl.) Ridl. 676 malayana K. & G. 676* subglabra K. & G. 676 Perriera 194 Perrottetia” HUBiK., non’ DC. 228-229) 1230.0 231g 252-0240: 288, 390, 391, 392, 930
Roxb.)
alpestris {non (BI.) Loes.] Koord. 291
alpestris (BI.) Loes. 229, 290, 291 map
ssp. alpestris 289*, 290 ssp. moluccana (BI.) Ding Hou 289*, 290, 291 ssp. philippinensis (Vidal) Ding Hou 289*, 290, 291 var. philippinensis Stapf 291 arborescens (F.v.M.) Loes. 29] arisanensis Hayata 290 caudata Ridl. 291 grandifolia Rid]. 291 lauterbachiana Loes. 291 f. B macrophylla Loes. 291 moluccana (Bl.) Loes. 291 nervosa Ridl. 291 philippinensis Loes. 291 racemosa (Oliv.) Loes. 290 schlechteri Loes. 291 traumatophylla Merr. & Perry 291 Phaleria’ Jack.2; 3,.5,'15; 19* ambigua Hook. f. 17 amboinensis Mertr. 21 axillaris Elm. 23 blumei (non Bth.) Hemsl. 18 calantha Gilg 22, 23 capitata Jack 2, 14*, 16, 20*,
1008
FLORA MALESIANA
[ser. I, vol. 66
21 map cauliflora Bedd. 20 coccinea (Gaudich.) 16, 21 cumingit F.-Vill. 20 dubiosa Zoll. 20 elegans Perry 14*, 16, 17 laurifolia Hook. f. 2, 17, 18 var. javanica Val. 17 longifolia Boer|. 17 macrocarpa (Scheff.) Boerl. 2, 144516522" neumanni F.v.M. 2 nisidai Kaneh. 16, 19 octandra (L.) Baill. 2, 14*, 16, 17, 18 map var. /aurifolia Warb. ex von Malm 17 octandra [non (L.) K.Sch. & Hollr. 22 papuana Warb. ex K.Sch. & Laut. 22 parvifolia Back. 17 pentecostalis Léandri 4, 16 perrottetiana (Decne) F.-Vill. 5, 16, 18 map, 19 platyphylla Mertr. 21 revoluta Boerl. 21 sogerensis S.Moore 16, 21 sp. Gilg 22 splendida Val. 18 subcaudata Merr. & Perry 21 urens Koord. 20 vriesii Baill. 21 wichmannii Val. 22, 23 zippelti Baill. 21 Phanrangia Tardieu 231 Philagonia Bl. 226 Phylidraceae 4: 5-7 Phyllocharis Diels 107, 109, 110, 137, 928 lamiifolia Wimmer 139 oblongifolia Diels 137, 139* saxicola v.Royen 137, 139*, schlechteri Diels 137, 139%, subcordata Merr. & Perry 137, 139*, Phyteuma 108 Phytolaccaceae 4: 228-232; 5: 557, 6: 109, 451 Picraena Lindl. 212 Picramnia 194 Picrasma Bl. 193, 195, 196, 212 andamanica Kurz ex Benn. 214 denhamii Seem, 214 javanica Bl. 213*, 214 map nepalensis Benn. 214 philippinensis Elm. 214 quassioides (D.Don) 193, 194, 214 Picroderma laotica Thorel ex Gagnep. 226 Picrolemma pseudocoffea Ducke 194
F.v.M.
Baill. ]
Benn.
Picrophloeus Bl. 299, 303 javanensis Bl). 303, 304 Picroxylon Warb. 203 siamense Warb. 205 Piddingtonia DC. 121 cyanocarpa Hassk. 132 montana Mig. 132 nummularia DC. 134 palliardii Lehm. 134 patens Miq. 132 Pieris (non D.Don 1834) Clarke 472, 674 ovalifolia (Wall.) D.Don 675 Pierreodendron Engl. 198, 202 Pimela alba Lour. 926 nigra Lour. 927 Pimelea Banks & Soland. ex Gaertn. 2, 3, 4, 5, 6, 43, 44 subg. Thecanthes (Wikstr.) Gilg 46 brevituba Fawc. 46 concreta F.v.M. 45*, 46* map cornucopiae Vahl 45*, 46 map philippinensis C.B.Rob. 46 sp. Dammerm. 46 Piper 468 Pirola = Pyrola Pisonia Plum. ex L. 450, 451, 457, 462 map aculeata L. 450, 457, 459, 460, 463*, 467 map alba Span. 464, 466* ‘anisophylla Hassk. 467 artensis (Montr.) Barg.-Petr. 459, 460 aruensis Barg.-Petr. 461 beccariana Barg.-Petr. 463 brunoniana Endl. 460 cauliflora Scheff. 460, 462 map corniculata Barg.-Petr. 460, 464 diandra Pulle 458*, 459, 460, 463 excelsa Bl. 460 excelsa (non BI.) Corner 459, 464, 465 fragrans 459 gammillii Merr. 461 grandifolia Warb. 463 grandis R.Br. 197, 223, 459, 460, 464, 465* map, 466* var. sylvestris (T. & B.) Heimer! 464 inermis (non Jacq.) Forst. ex Seem. 464, 466, 467 inermis Jacq. 466, 468 limonella B\. 467 lineatipilum C.DC. 468 longirostris Teysm. & Binn. 459, 460, 463*, 464 map macrophylla (Bojer) Choisy 464 major Baill. 462 membranacea K.Sch. & Hollr. 468
micrantha Val. 463 mitis L. 467 mooriana F.v.M. 461 morindifolia R.Br. ex Wight 464 miilleriana Warb. 459, 460, 462 map, 464 nishimura Koidz. 461 procera Bertero ex Guill. 464, 467 rostrata Warb. 463 spathiphylla K.Sch. & Laut. 463 sylvestris Teysm. & Binn. 464, 467 triandra Barg.-Petr. 463 umbelliflora (Forst.) Seem. 460, 461* map villosa Poir. 467 Pittosporaceae 5: 345-362; 6: 373, 960-963 Pittosporales 63 Pittosporum Banks ex Gaertn. 152, 960 berberidoides Burkill 961, 963 ferrugineum 12 var. filarium DC. 11 filarium Oken 11, 12 inopinatum Bakker 961, 962* moluccanum (Lamk) Miq. 373, 963 nubicola Schodde 963 pentandrum (Blco) Merr. 15, 963 pullifolium Burkill 963 var. globosum Steen. 961, 963 pumilum Schodde 961, 962 ramiflorum (Z. & M.) Zoll. ex Mid. 961 f. macrocarpum Bakker 961 resiniferum Hemsl. 961 serrulatum Jack ex Roxb. 963 sinuatum BI. 961, 963 var. efuniculare Steen. 961 tenuivalye Schodde 961 Pityopus Small 670 Platycarya 143 strobilacea 143 Platycodon grandiflorum (Jacq.) DC. 108, 141 Pleurostylia W. & A. 227, 228, 232, 287, 391, 392, 930 heynei W. & A. 288 opposita (Wall.) Alston 287*, 288
wightii W. & A. 288 var. neocaledonica Loes. 288
Pleurostylis = Pleurostylia 288 Plocosperma Bth. 954 Plumbaginaceae 4: 107-112; 6:
173, 174, 451 Plumbaginales 451 Plumbago 451 Podandrogyne 100
Sept. 1972]
Podocarpus 333 Podostemaceae 4: 65-68; 6: 963-964 Polanisia (non Raf.) DC. 62, 99, 100 angulata DC. 102 chelidonii DC. 102, 103 dodecandra (L.) DC. 100 icosandra W. & A. 103 f. deglabrata Back. 104 f. typica Back. 103 viscosa DC. 103, 104 f. typica Back. 103 var. @ Back. 103 var. deglabrata Back. 104 Polanisia Raf. 62, 99, 100 graveolens Raf. 99, 100 Polemoniaceae 4: 195-196 Polycardia Juss. 932 Polygalales 63 Polypremum 293, 295, 296 Pongamia pinnata (L.) Merr. 973 Pongelion Adans. 215, 216 sect. Ailanthus Pierre 215 sect. Euailanthus Engl. 215 sect. Eupongelion Rumph. ex Pierre 215 cacodendron Farwell 220 calycinum Pierre 218 excelsum Pierre 219 fauvelianum Pierre 219 grandis v.Tiegh. 218 imberbiflora Pierre 219 malabaricum Pierre 219 wightii v.Tiegh. 219, 220 Pongelion Rheede 219 Pongelium glandulosum 220 vilmorinianum v.Tiegh. 220 Pontederiaceae 4: 255-261; 5: 557; 6: 964 Posoqueria longiflora Aubl. 349 Potalia 295 Potaliaceae Hutch. 297 Potamogetonaceae 157, 171 Pragmatropa Pierre 245 Pragmotessara Pierre 245 japonica Pierre 252 Pratia Gaudich. 107, 109, 119, APM 22. 135 angulata (Forst.) Hook. f. 122, 134, 135 var. arenaria 134 archboldiana Merr. & Perry 131 arenaria Hook. f. 134 arenosa = arenaria 134
Pierre
begoniaefolia = _ begonifolia 134
begonifolia (non Lindl.) Hos- seus 128
begonifolia Lindl. 134 begoniifolia = begonifolia 134 borneensis Hemsl. 133
Index to scientific names
var. grandiflora Stapf 133 conferta Wimmer 135 hederacea G.Don 134
var. elliptica 134 linnaeoides Hook. f. 134 montana Hassk. 132
f. variegata Hochr. 132
var. cyanocarpa Wimmer 132 nummularia A.Br. & Aschers. 134, 135
ovata Elm. 128
papuana §.Moore 134, 135
podenzanae S.Moore 134
radicans G.Don 131
reniformis Kanitz 134
repens Gaudich. 134
serpyllacea Presl. 134
thunbergii G.Don 131
torricellensis K.Sch. & Laut. 128
wollastonii S.Moore 134, 135
zeylanica Hassk. 134
Primula Tourn. ex L. 173, 174,
175, 186, 188
sect. Callianthae Pax 191
sect. Candelabra Balf. f. 186, 191
sect. Cankrienia (de Vriese) Pax 186
sect. Proliferae Pax 186, 188, 191
auricula L. 175
cortusioides L. 174
elatior (L.) Hill 173
farinosa L. 186, 189
helodoxa Balf. f. 190, 191
hirsuta All. 188
hortensis 188
imperialis Jungh. 189, 190, 191 var. gracilis Pax 189
kewensis W.Wats. 175
khasiana Balf. f. 190, 191
kuhlii Bl. 190
malacoides Franch. 192
minutiflora Forrest 191
mollis Hook. 174
obconica Hance 174, 192
praenitens Ker-Gawl 175, 188, 192
prolifera Wall. 173, 186, 187*, 189*, 190* map
saxifragifolia O.K. 191
sieboldii Morren 174
sinensis Lindl., non Lour. 175, 188, 192
smithiana Craib 190, 191
sumatrana Merr. 190, 191
umbellata (Lour.) Bentv. 186, 189, 191*, 192 map
veris L. 173, 174 var. elatior L. 192
Primulaceae 173-192, 964 subfam. Androsaceae 174 subfam. Corideae 174
1009
subfam. Cyclamineae 174 subfam. Lysimachieae 174 subfam. Samoleae 174 Pristimera grahamii A.C.Smith 402 indica A.C.Smith 401 Proteaceae 5: 147-206, 566; 6: 368, 965 Protium javanicum Burm. f. 917 macgregorii (F.M.Bailey) Leenh. 917 Pseudais Decne 15 coccinea Decne 21 Pseudogardneria Racib. 296 Pseudospigelia 296 Psilaea Miq. 27 dalbergioides Miq. 27, 982 Psychotria L. 387, 959 Pteridophyta 171 Pterilema Reinw. 145 aceriflorum Reinw. 152 Pteris wallichiana Agardh 187* Pterocarya 143 Punicaceae 4: 226-227 Pyrola Tourn. ex L. 469, 472, 474, 671 Japonica Klenze ex Alefeld 673 ssp. coreana Andres 673 Japonica (non Klenze ex Ale- feld) Steen. 671 sumatrana Andres 671, 672* Pyrolaceae 472, 473 Pyroleae 472 Pyrospermum Miq. 280 calophyllum Miq. 282 Pyrus granulosa Bertol. 878
Quassia L. 968, 969 harmandiana (Pierre) 969
Noot.
Randia cochinchinensis (Lour.) Merr. 309
Rapuntium Miller 121 affine Pres] 128 alsinoides Pres] 127 angulatum Pres] 134 arenarioides Pres] 129 caespitosum Presl 130 campanuloides Pres 130 chinense Pres| 130 coloratum Pres! 123 haenkeanum Pres| 136 leschenaultianum Pres! 123 longiflorum Miller 140 longifolium Presl 136 nicotianaefolium Pres| 123 nummularium Pres| 134 pyramidale Pres} 123 radicans Pres| 130 reinwardtianum Presl 129 reniforme Pres] 134 roseum Pres| 123 succulentum Presl 128
FLORA MALESIANA
[ser. I, vol. 68
1010 trialatum Pres] 129 wallichianum Presl 123 zeylanicum Presl 128 Reissantia Hallé 390, 391, 400 cassinoides (DC.) Ding Hou 402
ferruginea (King) Ding Hou 402
grahamii (Wight) Ding Hou 402, 403*
indica (Willd.) Hallé 401
Resedaceae 62, 63
Restionaceae 5: 416-420, 569
Retzia Thunb. 295, 297, 954
Rex amaroris Rumph. 221
Rhamnaceae 49, 280, 284
Rhamphocarya Kuang 143
Rhesa = Bhesa 282, 283
Rhizophora L. 965 apiculata Bl. 966 harrisonii Leechm. 974 mucronata Lamk 965, 966 stylosa Griff. 966
Rhizophoraceae 5: 429-493; 6: 361, 965-967, 974
Rhododendron L. 469, 470, 471, 472, 473, 474, 475 map subg. Anthodendron (Rchb.)
Endl. ex Wils. & Rehd. 661, 943 sect. Anthodendron (Rchb.) Endl. 661, 943 subg. Anthodendron (non Wils. & Rehd.) Sleum. 661, 662 sect. Anthodendron Sleum. 662 subg. Azaleastrum Planch. 480, 665 sect. Choniastrum Franch. 480, 665, 666 map subg. Eurhododendron (Endl.) Koehne 656 sect. Eurhododendron Endl. 657 subg. Hymenanthes (BI.) Koch 480, 656 sect. Hymenanthes 480, 657 subsect. Irrorata Sleum. 480, 657 map subg. Hymenanthus = Hyme- nanthes 656 subg. Lepidorrhodion Koehne 480
subg. Pentanthera (G.Don) Pojarkova 480, 661, 943 sect. Pentanthera 480, 661, 943 subg. Pseudovireya Clarke 480 subg. Rhododendron 479, 480, 943 sect. Discovireya Sleum. 480 sect. Eurhododendron Endl. 480
sect. Hadranthe Schltr 497, 501, 547
sect. Hapalanthe Schltr 606
sect. Linnaeopsis Schltr 568
sect. Schistanthe Schltr 501, 606
sect. Vireya (Bl.) Copel. f. 475, 477*, 478*, 479, 480, 943 subsect. Albovireya
Sleum. 479, 537, 538
map subsect. Astrovireya Sleum. 580
subsect. Euvireya Copel. 1, ss, ASS SPS De 567 map, 606 ser. Buxifolia Sleum. 538, 568, 580 ser. Citrina Sleum. 568, 579 ser. Javanica Sleum. 536, 568, 606 ser. Linnaeoidea Sleum. 567, 568 ser. Saxifragoidea Sleum. 567, 574 ser. Stenophylla Sleum. 568, 576 ser. Taxifolia Sleum. 568, 575 subsect. Leiovireya Copel. f. 606 subsect. Linearanthera Copel. f. 580 subsect. Linnaeopsis (Schltr) Sleum. 568 subsect. Malayovireya Sleum. 477*, 478*, 479, 527 map subsect. Malesia Copel. f. 580 subsect. Phaeovireya Sleum. 478*, 479, 501, 511, 527 map subsect. Pseudovireya (Clarke) Sleum. 476, 478*, 479, 480 map, 527, 536 subsect. Schizovireya Sleum. 606 subsect. Siphonovireya Sleum. 476, 479, 497, 527 map subsect. Solenovireya Copel. f. 476, 479, 527, 547 map sect. Zygomorphanthe Schltr 480, 501, 537, 576, 580, 606 ser. Malayanum Copel. ii SPT) subg. Tsutsutsi (G.Don) Po- jarkova 480, 661 sect. Tsutsia — Tsutsutsi
662
sect. Tsutsugi = Tsutsutsi 662
sect. Tsutsusi = Tsutsutsi 662
sect. Tsutsutsi 480, 657 map, 662
subg. Vireya (Bl.) Clarke 480
abietifolium Sleum. 581, 590, 591
acrocline Sleum. 652
acrophilum Merr. & Quis. 581, 595
acuminatum Hook. f. 528, 534*
adinophyllum Merr. 481, 490*
aequabile J.J.S. 538, 539
agathodaemonis J.J.S. 497, 499, 505
agathodaemonis (non J.J.S. 1913) J.J.S. 1936 500
album BI. 538, 539, 626
album (non BI.) Ridl. 539
album (non BI.) Zoll. 540
alternans Sleum. 581, 592
alticolum Sleum. 582, 597
amabile Sleum. 548, 559
anagalliflorum Wernh. 568
andersonii Rid. 536
angiense J.J.S. 652
angulatum J.J.S. 607, 616
apoanum Stein 528, 531
araiophyllum Balf. f. & W.W. Sm. 661
archboldianum Sleum. 548, 555
arenicolum Sleum. 539, 546
arfakianum Becc. 581, 609, 610, 648
armitii F.M.Bailey 548, 557
asparagoides Wernh. 643
asperrimum Sleum. 509
asperum J.J.S. 476, 478, 517, Ryle
astrapiae Foerster 515
atjehense Sleum. 472, 657, 658*, 659*
atropurpureum Sleum. 570, 582, 602*, 603
aurigeranum Sleum. 609, 642
baenitzianum Laut. 608, 627
bagobonum Copel. f. 580, 586
banghamiorum (J.J.S.) Sleum. 581, 596
basirotundatum J.J.S. 626
beccarii Sleum. 607, 622, 626
beyerinckianum Koord. 477*, 478, 511 var. longipetiolatum J.J.S.
511
bloembergenii Sleum. 608, 633
bodenii Wernh. 497
brachygynum Copel. f. 609, 610, 650
brachypodarium Sleum. 549,
Sept. 1972]
Index to scientific names
561, 562* brassii Sleum. 470, 582, 600 brevipes Sleum. 609, 642 brevitubum J.J.S. 607, 622 breynii Planch. 663 brookeanum Low ex Lindl. 475, 537, 610, 655 var. brookeanum 655, 656 var. cladotrichum Sleum. 655, 656 var. extraneum Sleum. 655, 656 var. gracile (Low ex Lindl.) Sleum. 655, 656 brookeanum (non Low ex Lindl.) Stapf 653 bryophilum Sleum. 477*, 509 bullifolium Sleum. 522 burmanni G.Don 662 buruense J.J.S. 608, 632 buxifolium Low ex Hook. f. 582, 603 var. buxifolium 603, 604 var. robustum Sleum. 591, 604, 612 caespitosum Sleum. 568, 569*, ai /i be calceolarioides Wernh. 634 callichilioides Wernh. 628 var. minor Wernh. 628 calodon Ridl. 591 calosanthes Sleum. 582, 605 candidapiculatum Wernh, 572 carrii Sleum. 548, 557 carringtoniae (non F.v.M.) Lane-Poole 500 carringtoniae F.v.M. 548, 552, 558 var. maius J.J.S. 554 carstensense Wernh. 548, 553 caryophyllum Hayata 664 celebicum (Bl.) DC. 606, 615 cernuum Sleum. 538, 545 cerochitum Balf. f. & Forr. 661 chamaepitys Sleum. 548, 552 chevalieri Dop 475 christi Foerster 606, 612, 614 var. loniceroides Schltr 612 christianae Sleum. 471, 608, 637 chrysopeplon Sleum. 512 ciliilobum Sleum. 482, 494 cinchoniflorum Sleum. 497, 498* cinerascens Sleum. 540, 561 citrinum (Hassk.) Hassk. 579 var. albiflorum Miq. 579 var. citrinum 579 var. discoloratum Sleum. 580 citrinum (non Hassk.) Miq. 580 clementis Merr. 625 coelorum Wernh. 568, 573
coenenii J.J.S. 652 commonae Foerster 581, 587, 588*, 589* commutatum Sleum. 608, 629 comparabile Sleum. 608, 636 comptum C.H.Wright 543 var. comptum 538, 543, 544* var. trichodes 538, 544 coniferum Wernh. 543 convexum Sleum. 651 copelandii Merr. 567 coriifolium Sleum. 612 cornu-bovis Sleum. 580, 585 correoides J.J.S. 538, 543 corruscum Ridl. 659 crassifolium Stapf 607, 626 crassineryium Ridl. 626 cruttwellii Sleum. 548, 554, 555+ culminicolum F.v.M. 580, 581, 608, 610, 651 var. angiense (J.J.S.) Sleum. 476, 524, 651, 652 var. culminicolum 597, 651, 652 var. nubicola (Wernh.) Sleum. 580, 581, 651, 652 cuneifolium (non Stapf) Rendle 495 cuneifolium (non Stapf) Ridl. 496 cuneifolium Stapf 496 var. subspathulatum (non Ridl.) Merr. 496 var. subspathulatum Rid. 586 curranit Merr. 583 curviflorum J.J.S. 606, 613 cuspidellum Sleum. 608, 610, 629 cyatheicolum Sleum. 520 cyrtophyllum Wernh. 481, 491 dasylepis Schltr 512 delicatulum Sleum. 501, 507 var. delicatulum 507 var. lanceolatoides Sleum. 509 devrieseanum Koord. 515 ssp. astrapiae Foerster 515 devrieseanum vel aff. Stonor 505 devriesianum = devrieseanum 515 dianthosmum Sleum. 477*, 526 dielsianum Schltr 510, 511 var. dielsianum 510 var. stylotrichum Sleum. 510 discolor Warb. 615 disterigmoides Sleum. 568, 569*, 573 doctersii J.J.S. 643 dubium K. & G. 659
1011
durionifolium Becc. 528, 529,
531 durionifolium (non Becc.) Stapf 531 edanoi Merr. & Quis. 549, 564 elegans Ridl. 591 elongatum BI. 565 englerianum Koord. 608, 609, 628 ericoides (non Low ex Hook. f.) Burtt 496 ericoides Low ex Hook. f. 481, 491 var. ericoides 491* var. silvicolum Sleum. 492 erosipetalum J.J.S. 481, 485 extrorsum J.J.S. 501, 504 eymae Sleum. 501, 503 fallacinum Sleum. 528, 530*, 531 filamentosum Wernh. 560 flavoviride J.J.S. 608, 637 fortunans J.J.S. 528, 535 franssenianum J.J.S. 614 frey-wysslingii J.J.S. 580, 582 fuchsii Sleum. 609, 648 fuchsioides Schltr 486 fuchsioides aff. Stonor 575 fuscum Bl. 532 galioides J.J.S. 586 gardenia Schltr 514 gaultheriifolium J.J.S. 477%, 481, 484 var. expositum Sleum. 485 var. gaultheriifolium 484 gibbsiae J.J.S. 652 gilliardii Sleum. 522 giulianettii Laut. 538, 546 glabrifilum J.J.S. 635 glabriflorum J.J.S. 606, 614 goodenoughii Sleum. 549, 561 gorumense Schltr 634 gracile (non Low ex Lindl.) Becc. 629 gracile Low ex Lindl. 656 gracilentum F.v.M. 568, 570* gregarium Sleum. 651 habbemae Koord. 497 habbemai = habbemae 497 haematophthalmum Sleum. 522, 524 hameliiflorum Wernh. 481, 486 hansemannii Warb. 633 hatamense Becc. 580, 584, 606, 608 hatamense (non Becc.) Sleum. 652 helwigii (non Warb.) Koord. 499 hellwigii Warb. 501, 504 helodes Sleum. 581, 594 herzogii Warb. 470, 497, 500* himantodes Sleum. 528 var. himantodes 528
1012
FLORA MALESIANA
var. lavandulifolium Sleum. 529 hirtolepidotum J.J.S. 607, 618 hooglandii Sleum. 501, 507, 576* hybridogenum Sleum. 528, 537 impositum J.J.S. 609, 649 impressopunctatum J.J.S. 607, 617 incommodum Sleum. 497, 500 inconspicuum J.J.S. 470, 477*, 538, 582, 598 inconspicuum aff. Stonor 569 indicum (L.) Sweet 662, 663 f. variegatum (BI.) DC. 663 intranervatum Sleum. 610, 654 inundatum Sleum. 497, 499 invasorium Sleum. 481, 488 jasminifiorum (non Hook.) F.-Vill. 531 jasminiflorum Hook. 476, 478, 537, 548, 549, 565 var. copelandii (Merr.) Sleum. 566, 567 var. heusseri (J.J.S.) Sleum. 565, 566 var. jasminiflorum 566 var. maculatum Ridl. 566 var. oblongifolium Sleum. 566, 567 var. punctatum Ridl. 528, 537, 566 jasminiflorum (non Hook.) Koord. 579 jasminiflorum (non Hook.) Merr. 567 jasminiflorum (non Hook.) Ridl. 563, 566 Jasminiflorum (non Hook.) Sarasin 560 javanicum (BI.) Benn. 471, 478*, 537, 607, 610, 624* var. citrinum Hassk. 624 var. gymnocarpum Hochr. 624 var. javanicum 625, 626 var. schadenbergii (Warb.) Sleum. 625 var. teysmannii (Miq.) K. & G. 528, 537, 625 var. tubiflorum Hook. 640 jJavanicum (non Benn.) Clarke 620 jJavanicum (non Benn.) F.-Vill. 641 jJavanicum (non Benn.) Koord. 615 javanicum (non Benn.) Malm 650 jJavanicum (non Benn.) Steen. 621 keditii Sleum. 606, 612 kemulense J.J.S. 607, 623
keysseri Foerster 651 kjellbergii J.J.S. 630 klossii Rid]. 666 map, 667 kochii Stein 609, 641 konori Becc. 477*, 478*, 515, SUG7, 51873 S19N652 korthalsii Miq. 657, 660 lacteum Stapf 551 laetum J.J.S. 476, 478, 517, 521, 608, 609, 638* laetum (non J.J.S. 1914) J.J.S. 1936 643 lagunculicarpum J.J.S. 538, 542 lami J.J.S. 582, 599 lampongum Mig. 538, 544 lanceolatum Ridl. 609, 610, 647 langbianense Chey. ex Dop 659 laoticum Dop 667 lauterbachianum Foerster 633 ledifolium (Hook.) G.Don 662 leptanthum F.v.M. 478, 512, Sls leptobrachion Sleum. 608, 631 leptomorphum Sleum. 581, 593 leptopeplum Balf. f. & Forr. 661 leucobotrys Rid]. 666 leucogigas Sleum. 477*, 478, 608, 632 leytense Merr. 609, 649 var. leytense 649 var. loheri (Copel. f.) Sleum. 649 lindaueanum Koord. 481, 486 var. bantaengense J.J.S. 486 var. cyclopicum Sleum. 486 var. latifolium J.J.S. 486 var. lindaueanum 486 var. psilacrum Sleum. 486 lineare Merr. 528, 535 linearifolium S. & Z., non Poir. 943 var. macrosepalum (Maxim.) Makino 662, 663 linnaeoides Schltr 568 lobbii Veitch 640 loboense Copel. f. 607, 619 lochae F.v.M. 475, 636 loerzingii J.J.S. 607, 616 loheri Copel. f. 649 lompohense J.J.S. 608, 632 var. grandifolium J.J.S. 633 longiflorum Lindl. 475, 478, 537, 566, 608, 609, 640 var. bancanum Sleum. 641 var. heusseri J.J.S. 566 var. longiflorum 640 loranthiflorum Sleum. 549, 564 lowei = lowii 654
[ser. I, vol. 68
lowii Hook. f. 610, 654
lowii (non Hook. f.) F.v.M. 628 lukiangense Franch. 661 luraluense Sleum. 580, 608, 636 lussoniense Rendle 583 luteosquamatum Sleum. 582, 598, 599 maboroense Schltr 627 macgregoriae F.v.M. 478, 522, 608, 609, 610, 633, 634*, 652 var. glabrifilum (J.J.S.) Sleum. 634, 635 var. macgregoriae 634, 635 var. mayrii (J.J.S.) Sleum. 634, 635 macrosepalum Maxim. 662, 663, 943 macrosiphon Sleum. 548, 552 magnificum Sleum. 515 maius (J.J.S.) Sleum. 548, 554 malayanum Jack 477*, 478*, 528, 532, 586,193 p97, 017 var. axillare J.J.S. 533 f. axillare 532, 533 f. latifolium Sleum. 532, 533 f. ovatum Sleum. 532, 533 var. infrapilosum Sleum. 532, 533 var. malayanum 528, 532 var. pilosifilum Sleum. 528, 532, 539 var. pubens Sleum. 528, 5325559 malayanum (non Jack) Koord. 540 malindangense Merr. 496 maxwellii Gibbs 609, 644 mayrii J.J.S. 635 megalostigma F.v.M. 628 melantherum Schltr 520 meliphagidum J.J.S. 482, 493 microphyllum J.J.S. 568, 571 mindanaense Merr. 610, 650 minimifolium Wernh. 483, 571 mjoébergii Merr. 529 molle (Bl.) G.Don 661 mollianum Koord. 610, 652 moszkowskii Schltr 643 moulmainense Hook. 666 map, 667*, 668 moultonii Ridl. 607, 623 mucronatum (BI.) G.Don 662 multicolor Miq. 607, 621 var. curtisii Henslow 621 multicolor (non Miq.) S.Moo- re 580 multinervium 549, 550* murudense Merr. 626 murudense (non Merr.) J.J.S.
Sleum. 547,
Sept. 1972]
Index to scientific names
618 muscicola J.J.S. 568, 572 myrsinites Sleum. 576, 578 nanophyton Sleum. 492 var. nanophyton 481, 492 var. petrophilum Sleum. 482, 493 natalicium Sleum. 547, 549 neriifolium Schltr 501, 506 nervulosum Sleum. 646 var. exuberans Sleum. 610, 646 var. nervulosum 609, 646* nieuwenhuisii J.J.S. 580, 586 ningyuenense Hand.-Mazz. 659 nitens Sleum. 593 nodosum C.H.Wright 651 nortoniae Merr. 528, 533 nubicola Wernh. 652 nummatum J.J.S. 481, 484 obscurinervium Merr. 656 obscurum Sleum. 528, 535 oliganthum Sleum. 548, 556 ombrochares Balf. f. & Ward 661 opulentum Sleum. 519 oranum J.J.S. 643 orbiculatum Rid]. 549, 563, 608, 609 oreadum Wernh. 548, 560 oreites Sleum. 481, 485 var. chlorops Sleum. 485 var. oreites 485 orion Ridl. 490 var. aurantiacum Ridl. 640 oxycoccoides Sleum. 568, 369s 57/3 oxyphyllum Franch. 666 pachycarpon Sleum. 606, 614 pachystigma Sleum. 607, 616 papuanum Becc. 581, 594, 5955 papuanum (non Becc.) C.H. Wright 546 partitum J.J.S. 647 parvulum Sleum. 568, 569*, 573 pauciflorum K. & G. 581, 591 var. calodon (Ridl.) Sleum. 591 var. pauciflorum 591 pennivenium Balf. f. & Forr. 660 perakense K. & G. 481, 488, 536 perplexum Sleum. 607, 626 phaeochitum F.v.M. 511 Phaeochitum (non F.v.M.) Wright 526 phaeochristum Sleum. 476, 478, 523 phaeopeplum Sleum. 476, 478, 517, 519, 644 phaeops Sleum. 525
planecostatum Sleum. 580, 586
pleianthum Sleum. 548, 556
pneumonanthum Sleum. 549, 563
podocarpoides Schltr 579
polyanthemum Sleum. 609, 641
poremense J.J.S. 610, 652
porphyranthes Sleum. 582, 599
prainianum Koord. 501, 505
proliferum Sleum. 538, 540
protandrum Sleum. 497, 498
psammogenes Sleum. 581, 591
pseudobuxifolium Sleum. 581, 587
pseudomurudense Sleum. 607, 618
pseudonitens Sleum. 581, 593
pseudotrichanthum Sleum. 548, 559
psilanthum Sleum. 511
pubigermen J.J.S. 580, 583, 597 var. banghamiorum J.J.S.
596
pubitubum Sleum. 548, 558
pudorinum Sleum. 538, 545
pulleanum J.J.S. 483 var. maiusculum Sleum.
481, 484
var. pulleanum 481, 483
purpureiflorum J.J.S. 576, 578
pusillum J.J.S. 568, 572
pyrrhophorum Sleum. 527, 582, 597
quadrasianum Vidal 482, 494
banahaoense Copel. f. 495
davaoense Copel. f. 495
halconense Copel. f. 495
lutea H.J.Lam 496
marivelesense Copel. f.
496
monodii H.J.Lam 496
negrosense Copel. f. 495
pulogense Copel. f. 495
f. pulogense (non Copel. f.) H.J.Lam 496
f. rubra H.J.Lam 496
var. angustissimum Sleum. 494, 496
var. borneense J.J.S. 494, 497
var. cuneifolium (Stapf) Copel. f. 494, 496, 478*
var. davaoense (Copel. f.) Sleum. 494, 495
var. intermedium Merr. p.p. 495
var. intermedium Merr. s. str. 494, 496
var. malindangense (Merr.) Copel. f. 494, 496
1013
var. marivelesense (Copel. f.) Sleum. 494, 496 var. quadrasianum 494, 495 var. rosmarinifolium (Vi- dal) Copel. f. 494, 495 var. selebicum J.J.S. 494, 496 var. villosum J.J.S. 494, 496 radians J.J.S. 548, 560 var. minahasae Sleum. 560 var. radians 560 rappardii Sleum. 519, 525 rarelepidotum J.J.S. 620 var. ootrichum Sleum. 609, 621 var. rarelepidotum 607, 620 rarum Schltr 501, 507, 508* renschianum Sleum. 610, 650 retivenium Sleum. 607, 610, 653 retrorsipilum Sleum. 549, 565 retusum (BIl.) Benn. 470, 471, 477*, 481, 820* f. angustifolia Miq. 482 var. epilosum Sleum. 483 var. macranthum Hochr. 482 var. retusum 482 var. trichostylum Sleum. 483 retusum (non Benn.) F.-Vill. 494 retusum (non Benn.) Steen. 566 retusum (non Benn.) Wernh. 486 revolutum Sleum. 501, 503 rhodochroum Sleum. 521 rhodoleucum Sleum. 548, 558 rhodopus Sleum. 608, 627 rhodosalpinx Sleum. 548, 551 rhodostomum Sleum. 581, 592 ripense Makino 663 ripleyi Merr. 581, 596 var. basitrichum Sleum. 581, 596, 597 var. cryptogonium Sleum. 527, 596, 597 var. ripleyi 596 robinsonti Rid]. 607, 620, 621 rosendahlii Sleum. 608, 635 rosmarinifolium Vidal 495 rubellum Sleum. 526 rubrobracteatum Sleum. 582, 605 rubropilosum Hayata 664, 665 rugosum Low ex Hook. f. 590, 606, 611, 612 var. coriifolium (Sleum.) Sleum. 581, 590, 612 var. rugosum 611 ruttenti J.J.S. 547, 550 saavedranum Diels 512
salicifolium Becc., non BI. 609, 646 salicifolium Bl. 621 sarasinorum Warb. 625 saruwagedicum Foerster 481, 487* var. alpinum Sleum. 487 saxifragoides J.J.S. 574*, 575, 589* sayeri Sleum. 606, 615 scabridibracteum Sleum. 608, 609, 639* scrabrum G.Don 662 scarlatinum Sleum. 582, 601 schadenbergii (non Warb.) Merr. 619, 641 schadenbergti Warb. 625 schizostigma Sleum. 482, 493 schlechteri Laut. 608, 631, 632 schoddei Sleum. 523 schultzei Schltr 512 scortechinii K. & G. 481, 490, 536 seimundii J.J.S. 481, 490 seranense = seranicum 619 seranicum J.J.S. 607, 617, 619 sessilifolium J.J.S. 607, 622 sheilae Sleum. 581, 590 siamense Diels 666 simsii Planch. 662, 663, 665 simulans Sleum. 582, 600 sinense (Lodd.) Sweet 661 solitarium Sleum. 523 spathulatum Ridl. 481, 487, 536 spectabile Merr. 625 spondylophyllum F.v.M. 520 stapfianum Hemsl. ex Prain 548, 551 stelligerum Sleum. 501, 506 stenaulum Balf. f. & Forr. 666 stenophyllum Hook. f. 576*, Sif var. angustifolium J.J.S. 577 stolleanum Schltr 510 stonori Sleum. 587 stresemannii J.J.S. 610, 653 suaveolens Sleum. 563 subcordatum Becc. 608, 630 subcrenulatum Sleum. 582, 605 subpacificum Sleum. 549, 565 subsessile Rendle 662, 663, 664*, 665* var. baucoense Copel. f. 664, 665 subuliferum Sleum. 582, 601 subulosum Sleum. 576, 578 sumatranum Merr. 609, 647 superbum Hort. ex Lavallée 505 superbum Sleum. 501, 505 syringoideum Sleum. 548, 549, 553 taxifolium Merr. 575
FLORA MALESIANA
[ser. I, vol. 68
taxoides J.J.S. 481, 483
teysmannii (non Miq.) Hend. 620
teysmannii Miq. 626
thaumasianthum Sleum. 515
torricellense Schltr 635
toverenae F.v.M. 515
toxopei J.J.S. 610, 655
trichanthum Sleum. 559
triumphans Yersin & Cheval. 475
truncicolum Sleum. 525
tuba Sleum. 548, 557
tuberculiferum J.J.S. 509
tubiflorum Low ex Lindl. 640
tubiflorum (non Bl.) Mor. 540
tubiflorum Reinw. in Bl. 532
tubulosum Reinw. 615
uliginosum J.J.S. 638
ultimum Wernh. 582, 600
undulaticalyx J.J.S. 648
vanderbiltianum Merr. 481, 488*
vandeursenii Sleum. 580, 581, 585
vanvuurenii J.J.S. 471, 608, 630
variolosum Becc. 536 var. andersonii
Sleum. 528, 536
var. variolosum 528, 536
velutinum Becc. 611
versteegiil J.J.S. 539, 546
verticillatum (non Low ex Lindl.) Becc. 567
(RidI.)
verticillatum (non Low) Koord. 560 verticillatum (non Low ex
Lindl.) Low ex Hook. f. 604 verticillatum Low ex Lindl. 606, 610 f. velutinum (Becc.) Sleum. 606, 607, 611 f. verticillatum 609, 610 verticillatum (non Low ex Lindl.) Vidal 583 vidalii Rolfe 580, 583, 584* villosulum J.J.S. 606, 613 vinicolor Sleum. 528, 529 vinkii Sleum. 482, 493 vitis-idaea Sleum. 581, 592 vonroemeri Koord. 633 warianum Schltr 514 wentianum Koord. 608, 610, 628 wentianum all. Stonor 612 westlandii Hems]. 666 whiteheadii Rendle 583 wilhelminae Hochr. 528, 536 williamsii Merr. ex Copel. f. 607, 619 wollastonii Wernh. 628 womersleyi Sleum. 603, 568, 569* wrayi K. & G. 657, 659, 660*
var. elliptica Ridl. 659 var. minor Ridl. 659 wrightianum Koord. 478, 582, 604 var. cyclopense J.J.S. 604 var. insulare Sleum. 604 var. ovalifolium J.J.S. 600, 604 var. piliferum J.J.S. 594 var. wrightianum 604 xanthopetalum Merr. 607, 617 yelliotii Warb. 538, 542, 599 yelliottii = yelliotii 542 zippelti Bl. 579 zoelleri Warb. 476, 478, 519, 609, 635, 639, 643, 644*, 645*, 652 zollingeri J.J.S. 538, 540, 541 * var. latifolium J.J.S. 540 zollingerianum = zollingeri 540 Rhoeadales 62 Rhoipteleaceae 143 Rhus cacodendron Ehrh. 220 javanica L. 211 cf. linguata W.Slis 291 vernicifera DC. 220 Rigiolepis Hook. f. 754 bigibba (J.J.S.) J.J.S. 761 borneensis Hook. f. 756 caudatifolia (Merr.) J.J.S. 765 endertii J.J.S. 758 filiformis J.J.S. 758 korthalsii J.J.S. 763 lanceifolia Ridl. 755 lanceolata (Bl.) J.J.S. 761, 762 f. elliptica J.J.S. 762 f. marapiensis J.J.S. 762 f. sumatrana (Miq.) J.J.S. 762 lancifolia = lanceifolia 755 leptantha (Miq.) J.J.S. 764 f. elliptica J.J.S. 764 lobbii Ridl. 755 macrophylla J.J.S. 757 moultonii (Merr.) J.J.S. 763 salicifolia J.J.S. 757 sulcata (Ridl.) J.J.S. 758 uniflora (J.J.S.) J.J.S. 757 Rinorea 963 glabra O.K. 235 lanceolata (Roxb.) O.K. 237, 963 Rockia Heimer] 457, 459 Rosaceae 472, 878 Rosales 49 Rourea Aubl. 934 subg. Palliatus Leenh. sect. Palliatus 934 acropetala Pierre 934 mimosoides (Vahl) Planch. 934 f. mimosoides 934 minor (Gaertn.) Leenh. 934, 935
Sept. 1972]
Index to scientific names
ssp. microphylla (Hook. &
Arn.) Vidal 934 ssp. minor 934 ssp. monadelpha Vidal 934 oligophlebia Merr. 934 ovale (Schellenb.) Leenh. 934 pinnata (Merr.) Veldkamp 934 prainiana Talbot 934 Roureopsis Planch. 933 sect. Taeniochlaena Leenh. 934 acutipetala (Miq.) Leenh. 934 ssp. borneensis (Schellenb.) Leenh. 934 asplenifolia Schellenb. 933, 934 emarginata (Jack) Merr. 934 incurva Pierre 933 stenopetala (Griff.) Schellenb. 933 Roydsia Roxb. 95, 97 floribunda Planch. ex Hook. f. 97 parviflora (non Griff.) King 99
(Roxb.)
philippinensis Turcz. 97 scortechinii King 99
Rubiaceae 107, 195, 294, 297, 309, 336, 349, 373, 379, 387, 401 subfam. Coffeoideae 959 tribe Hedyotideae 297 tribe Psychotrieae 387
Rubiales 294, 297
Ruppia 171
Ruppiaceae 171
Rutaceae 69, 194, 195, 206, 214, 220, 221, 226, 969
Ruthiella Steen. 928 oblongifolia (Diels) Steen. 928 saxicola (v.Royen) Steen. 928 schlechteri (Diels) Steen. 928 subcordata (Merr. & Perry)
Steen. 928
Sagittaria sagittifolia L. ssp. /eucopetala (Miq.) Har- tog 915 trifolia L.915 Salacia L. 231, 390, 391, 392, 404 map, 421, 930, 932 alternifolia Scort. 421 amentacea Ridl. 408 amplifolia Merr. ex Chen & How 412, 413 hartletti Ridl. 420 beccarii Ridl. 417 blepharophora Ding Hou 405, 408 buddinghii Scheff. 412 campanuloidea King 408 castaneifolia Ridl. 406, 413 celebica Bl. 412
cerasiformis Teysm. & Binn. |
chinensis L. 392, 406, 419 map, 421, 932
coromandeliana Binn. 421
cymosa Elm. 405, 407, 408, 416*
diandra Miq. 410 f. lanceolata Miq. 411
disepala (C.T.White) Hou 420
erythrocarpa K.Sch. 404, 405, 411
euphlebia Merr. 406, 418
evonymiflora Zipp. ex Mid. 419
exsculpta Korth. 406, 417
flavescens Kurz 412 var. dumosa King 412
forsteniana Miq. 404, 405, 410, 932
grandiflora Kurz 406, 413, 932 var. longifolia (Hook. f.)
King 413
griffithi Laws. 400
integrifolia Merr. 415
intermedia Ding Hou 411*
jJavanensis Bl. 400
kalahiensis Korth. 406, 420
kamputensis Pierre 412
klossii Ridl. 408
korthalsiana Miq. 405, 406, 407 map, 420
kraemeri Ding Hou 410
lanceolata Teysm. & Binn. 408
latifolia Wall. ex Laws. 419
laurifolia Stapf 405, 406, 417
lawsoni King 412
ledermannii (Loes. ex Harms) Ding Hou 405, 410
leucoclada Ridl. 405, 406, 415
litseifolia Ridl. 415
littoralis Back. 419
lobbii Laws. 417
longifolia Hook. f. ex Laws. 413
longipedicellata) Ding Hou 406, 413
macrocarpa Korth. 412
macrophylla BI. 405, 412 map, 416* var. angustifolia Miq. 412
maingayi Laws. 406, 416%, 417
marginata Ding Hou 405, 406, 413
megasperma Ridl. 417
melitocarpa Bl. 408
minutiflora Rid]. 420
naumanni Engl. 419
nitidissima Merr. 406, 418
oblonga (non Wall.) Rendle
Teysm. &
Ding
405,
412 oblonga Wall. 420 oblongifolia Bl. 405, 408 f. latior Miq. 408 ovalis (non Korth.) Koord.- Schum. 419 ovalis Korth. 406, 415, 416* ovalis Laws., non Korth. 412 papuana (Loes.) Ding Hou 405, 408, 409* patens Decne 419, 420 perakensis King 400 Philippinensis Merr. 406 polyantha Korth. 414 prinoides (Willd.) DC. 419, 420, 421 var. macrophylla (Bl.) King 412, 419 var. timorensis Span. 419 radula (non G.Don) Dietr. ex Hassk. 406, 407 radula G.Don 407 roxburghii (non Wall.) Vidal 415 roxburghii Wall. ex Laws. 420 rubra Laws. 417, 418 scortechinii King 413, 414 sinensis (non L.) Bleco 406 sinensis Gmelin 418, 420 socia Craib 419 sororia Mig. 405, 409*, 420, 932 sp. 280 subalternifolia Merr. & Perry 405, 407, 416* subscandens Elm. 420 triplinervis Llanos 421 venosa Ding Hou 406, 417 verrucosa Wight 406, 414%, 932 vimeria Wall. 392, 406, 418, 421 viridis Craib 408 wenzelii Merr. 405, 411, 416* wrayi King 418 Salacicratea Loes. 404 australis Loes. 420 brassti A.C.Smith 409 diandra (Miq.) A.C.Smith 410 glandulosa A.C.Smith 405, 409 kraemeri (non Loes.) Kaneh. 419, 420 kraemeri Loes. 410 ledermannii Loes. 410 papuana Loes. 408 sarasinorum Harms 410 sororia A.C.Smith 409 Salacighia Loes. 930 Salicaceae 5: 107-110 Salvadoraceae 4: 224-225 Samadera Gaertn. 193, 194, 198, 199, 226, 968 brevipetala Scheff. 201 glandulifera Presl 201 indica Gaertn. 199
1016
FLORA MALESIANA
[ser. I, vol. 66
var. brevipetala_ (Scheff.) Back. 199 var. papuana Laut. 199 madagascariensis Juss. 200 mekongensis Engl. 201 pentapetala G.Don 201 tetrapetala Wall. 200 Samandura L. ex Baill. 199 indica Baill. 201 madagascariensis Perr. de la Bathie 201 mekongensis Pierre 201 Sambucus 51 Samolus 173, 174 valerandi L. 173, 192 Sanango G.S.Bunting & J.A. Duke 295, 296 Santaloides ovale Schellenb. 934 Santiria Bl. 917, 920 apiculata Benn. 920, 921 var. apiculata 921 var. rubra (Ridl.) Kalkman 921 conferta Benn. 920, 921 grandiflora Kalkman 920, 921 griffithii (Hook. f.) Engl. 920, 921 laevigata Bl. 920, 921 megaphylla Kalkman 920, 921 mollis Engl. 920 nervosa H.J.Lam 919, 921 oblongifolia Bl. 920, 921 ridleyi H.J.Lam 920 rubiginosa BI. 920, 921 var. rubiginosa 921 tomentosa BI. 920 Sapindaceae 49, 194, 226 Sapindales 49, 53, 933 Sapindus saponaria L. 226 Sapotaceae 229 Saprosma 387 Sarawakodendron 930 filamentosa Ding Hou 931*, 932 Sarcocaulon 445 ’ rigidum Schinz 445 Sarcosperma uittienii H.J.Lam 967 Sarcospermataceae 4: 32-34; 6: 967 Sarcotheca pinnata Merr. 934 Saururaceae 4: 47-48 Saxifragaceae 173, 174, 280, 472 Scaevola L. 949, 951 lobelia Murr. 951 micrantha Pres! 951, 952 oppositifolia R.Br. 951, 952 pauciflora Leenh. 951, 952 plumierii 223 sericea Vahl 951 taccada (Gaertn.) Roxb. 949, 951
Ding Hou
verticillata Leenh. 951, 952* Scaphocalyx spathacea Ridl. 944
Schmidelia conferta Blco 282 Schmidellia = Schmidelia 282 Scolopia Schreb. 943 kermodei C.E.C.Fischer 943 Scopolia (non Jacq.) L. f. 35, 37 composita L. f. 38 Scrophulariaceae 136, 294, 295, 296, 297, 960, 965 Scrophulariales 297 Scutinanthe Thw. 921 brevisepala Leenh. 919, 921 brunnea Thw. 921 Scyphostegia Stapf 967, 968* borneensis Stapf 967, 968* Scyphostegiaceae 5: 297-299; 6: 967 Scyphostrychnos 296 Selenocera Zipp. ex Span. 375 secundiflora Zipp. ex Span. 375 Sericolea Schltr 914 Shorea selanica BI. 154 Simaba Aubl. 198, 202 Simarouba 194, 198 cedron Planch. 194 glauca 194 Simaroubaceae 193-226, 968— 972 subfam. Alvaradoideae 195, 971 subfam. Irvingioideae 193, 194, 195, 970, 971, 972 subfam. Kirkioideae 195, 971 subfam. Picramnioideae 195, 971 subfam. Simarouboideae 195 subfam. Surianioideae 193, 194 tribe Simaroubeae 198 subtribe Eurycominae 205 subtribe Simaroubinae 195, 205 Simarubopsis Engl. 202 Simirestis 390 Siphonodon Griff. 227, 228, 229, 230, 231, 390, 391, 392, 394, 968 australe Bth. 230 celastrineus Griff. 229, 231, 394, 395*, 396* map var. acuminatissima Merr. 395 var. integrifolia Tardieu 412 var. subglobosa Merr. 395 peltatus Ding Hou 395*, 397, 932 pyriformis Merr. 395, 396 var. parvifolius Merr. 395 Sirioides Rumph. 355 alter Rumph. 355 Solanaceae 297 Solenopsis 140 Solenospermum Zoll. 231, 262, 264 apiculatum Ridl. 272
aquatile Ridl. 270, 271 Javanicum Zoll. 269 ledermannii Loes. 271 littorale Loes. 261 macranthum Loes. 264, 268 micranthum Loes. 268 oblongifolium Loes. 270 pallidum Loes. 268 paucinervium Loes. 270 torricellense Loes. 271 var. opacum Loes. 271 toxicum Loes. 270 Solmsia 3 Sonneratia L.f. 973 acida L. f. var. mucronata Miq. 975 alba J.Smith 973*, 974*, 975
alba * ovata 974* caseolaris) (.) Engel )973*, 974*, 975
ovata Backer 974*, 975 Sonneratiaceae 4: 280-289; 5: 557; 6: 973 Soulamea Lamk 193, 196, 221 amara Lamk 193, 197, 221, 222*, 223 map soulameoides (A.Gray) Noote- boom 221 terminalioides Baker 221 Sparganiaceae 4: 233-234 Specularia Heist. ex Fabr. 928 speculum-veneris (L.) Caruel 928 Speirema Hook. f. & Th. 121 montanum Hook. f. & Th. 132 Sphaerodiscus Nakai 245 cochinchinensis Nakai 248 Sphenoclea 107, 108, 109, 110 zeylanica Gaertn. 928 Sphenocleaceae 4: 27-28. See Campanulaceae Spigelia L. 295, 296, 297, 377 anthelmia L. 376*, 378, 960 Spilanthes urens 223 Spinifex 221 Spiraeaceae 195 Stackhousiaceae 4: 35-36 Staphylea 49, 52 Staphyleaceae 49-59, 214, 230 Steironema Raf. 177 Stellera L. 28 chamaejasme L. 28 passerina L. 28 Stenanthera R.Br. 424 Stenocarpus moorei F.v.M. 428 Stixis Lour. 61, 63, 95, 97 map fasciculata var. borneensis Heine 97 floribunda (Planch. ex Hook. f.) Pierre 97 obtusifolia (Hook. f. & Th.) Pierre 97 ovata (Korth.) Hall. f. 97 ssp. fasciculata (King) Ja- cobs 97 map
Sept: 1972]
ssp. ovata (Korth.) Hall. f. 96*, 97 map
philippinensis (Turcz.) Merr. 95, 97 map
scortechinii (King) Jacobs 97 map, 99
suaveolens (Roxb.) Pierre 95, 97
Struthiola 4
Strychnos L. 205, 293, 294, 295,
296, 297, 343, 361, 954
sect. Booneae Duvign. 346
sect. Breviflorae Progel 345
sect. Brevitubae A.W.Hill 345, 346, 955
sect. Floribundae Duvign. 346
sect. Intermediae Progel 345
sect. Lanigerae A.W.Hill 345, 346, 955
sect. Ligustroides Duvign. 346
sect. Longiflorae Progel 345
sect. Penicillatae A.W.Hill 34529346, 352*, 357,955
sect. Sambae Duvign. 346
sect. Strychnos 955
sect. Tubiflorae A.W.Hill 345, 346
acuminata Wall. ex DC. 355
aenea A.W.Hill 958 var. acuminata A.W.Hill 958
andamanensis A.W.Hill 351
angustiflora Bth. 345, 955, 958, 959
arborea A.W.Hill 358, 360
armata A.W.Hill 358, 359
axillaris Colebr. 346, 347, 351, 352%, 958.) 360)" 3615955, 958
balansae A.W.Hill 347, 349
bancroftiana F.M.Bailey 355, 356
barbata A.W.Hill 355
beccariti Gilg 347, 349
borneensis Leenh. 346, 347, 355, 356, 357*, 955, 958
celebica Koord. 355
cenabrei Merr. 358
chloropetala A.W.Hill 358, 359
cinnamophylla Gilg & Bened. 355
colubrina Auct. non L. 346, BAPE SOs 55941 3945.11355, 356, 358, 361, 955, 958
confertiflora Merr. & Chun 351
curtisii King & Gamble 345, 346, 351, 353, 355, 955, 958
cuspidata A.W.Hill 347, 349
dubia A.W.Hill 355
flavescens K. & G. 346, 347, 357, 955, 958
forbesii A.W.Hill 354*, 355, 356
grandis Wall. 361
hirsutiflora A.W.Hill 351
horsfieldiana (non Miq.) Koord.-Schum. 351
horsfieldiana Miq. 358, 359
hypogyna C.B.Clarke 355
ignatii Bergius 297, 345, 346, 347, 348*, 351, 361, 955, 957
ignatii (non Berg.) Vidal 355
impressinervis A.W.Hill 351, 358, 359
kawbet A.W.Hill 358, 359
kerrii Hill 955, 959
kerstingti Gilg & K.Sch. 355
krabiensis A.W.Hill 347, 349
lanata A.W.Hill 345, 346, 355, 955, 958
lanceolaris Miq. 346, 347, 357, 955, 958
laurina DC. 350
laurina (non DC.) Heine 356
laurina (non DC.) K. & V. 351
laurina Wall. ex DC. 355 var. thorelii A.W.Hill 355
ledermannii Gilg & Bened. 346, 347, 357, 955, 958
leuconeura Gilg & Bened. 355
ligustrina Bl. 350
lucida R.Br. 293, 346, 349, 350 map, 355, 955, 957
luzonensis Elm. 346, 347, 355, 358, 955, 958
maingayi C.B.Clarke 346, 350, 351, 955, 957, 958 ssp. borneensis Leenh. 958 var. fructuosa Clarke 347,
350
malaccensis Bth. 358, 360
melanocarpa Gilg & Bened. 361, 955, 959
merrillii A.W.Hill 355, 356
minahassae Koord. ex Boerl. 355
minor Dennst. 955, 958
monosperma Miq. 358
mucronata A.W.Hill 358
multiflora Bth. 355, 356
muricata Kosteletzky 350
myriantha Gilg & Bened. 355
nux-blanda A.W.Hill 349, 959
nux-vomica Auct. non L. 958
nux-vomica L. 297, 345, 346, 348*, 349, 955, 957 f. depauperata Miq. 349
oleifolia A.W.Hill 346, 347, 353, 355, 955, 958
oophylla Gilg & Bened. 358, 360
ovalifolia Wall. 347, 349, 351
ovata A.W.Hill 346, 351, 352 SS gS OW,
ovata (non Hill) Merr. 358
palembanica Miq. 358, 359
panayensis A.W.Hill 351
penicillata A.W.Hill 358, 360,
Stylidium
1017
361 philippensis Blco 957 philippinensis Blco 347, 349,
957 pilgeriana Gilg 358 plumosa A.W.Hill 358, 359 polytoma Gilg & Bened. 358,
360 polytoma (non Gilg & Bened.)
Kaneh. & Hatus. 355 polytrichantha Gilg 345, 346,
353, 357, 955, 958 potatorum (non L. f.) F.-Vill.
355
var. multiflora Vidal 355 pseudotieuté A.W.Hill 347 psilosperma F.v.M., 358, 360 pubescens C.B.Clarke 358, 360
var. scortechinii K. & G.
358 pycnoneura Gilg & Bened. 355 quadrangularis A.W.Hill 346,
347, 356, 955, 958 quintuplinervis A.W.Hill 358,
359 ridleyi K. & G. 346, 347, 360,
955, 958 robinsonti A.W.Hill 358, 360 roborans A.W.Hill 350 rufa C.B.Clarke 358, 361, 955,
958 schmidtii Gilg 358, 359 scortechinii A.W.Hill 358,
359 septemnervis C.B.Clarke 355
var. imberbis A.W.Hill 355 silvicola A.W.Hill 355 similis A.W.Hiull 355 sp. Hill 358 sp. Merr. 957 sp. 1 Hill 351 sp. a & B Koord.-Schum. 351 spireana Dop 349 tesseroidea A.W.Hill 358, 359 thorelii Pierre ex Dop 361,
955, 958 tieute Leschen. 347, 349 vanprukii Craib 357, 955, 958 villosa A.W.Hill 346, 351, 355,
955, 958 viridiflora A.W.Hill 358 wenzelii Merr. 358, 360
Stylidiaceae 4: 529-532; 5: 564;
6: 108, 109, 976
androsaceum O.
Schwarz 976
ericksonae Willis 976
fluminense Erickson & Willis 976
inconspicuum Sloot. 976
javanicum Sloot. 976
pedunculatum R.Br. 976
Styphelia J.E.Smith 423, 424
subg. Cyathodes (Lab.) Drude 425, 426 map, 433
1018
FLORA MALESIANA
[ser. I, vol. 68
subg. Leucopogon (R.Br.) Drude 423, 425, 426 map sect. Styphelia 423
‘abnormis (Sond.) J.J.S. 425, 433
abscondita J.J.S. 425
acuminata (R.Br.) Spr. 423, 425, 432
arfakensis Kaneh. & Hatus. 443
brassii Sleum. 425, 433, 434*
brevistyla Moore 434
carstensensis Wernh. 441
celebica J.J.S. 439
cornifolia Rudge 437
culminis Wernh. 440
cuspidata (R.Br.) Spr. 432
cymbulae (Lab.) Spr. 427
dekockii J.J.S. 440
forbesii Sleum. 425, 432
gjellerupti J.J.S. 438
hookeri (Sond.) J.J.S. 429
javanica (de Vriese) J.J.S. 423, 425, 430*, 431*, 432
juniperina (Forst.) Pers. 434
lamii J.J.S. 443
lancifolia J.5.S. 433
learmonthiana Gibbs 439
leptospermoides (R.Br.) Spr. 433
malaica Jack 426
malayana (Jack) Spr. 425, 426, 943
423,
var. malayana 425, 426, 427* var. novoguineensis 425,
427, 428*, 429* moluccana J.J.S. 433 montana F.v.M. 429
var. hookeri 429 nesophila (DC.) Sleum. 432 nubicola Wernh. 442 nutans J.J.S. 441 var. arfakensis J.J.S. 441 obovata J.J.S. 432 obovata Malm 429 obtusifolia J.J.S. 429 var. hypoleuca J.J.S. 429 oxycedrus Lab. 434 papuana (C.H.Wright) Koord. 443 Philippinensis Merr. 429 pungens Koord. 431 rapae Sleum. 434 spicata J.J.S. 429 suaveolens (Hook. f.) Warb. 423, 425, 428, 429* trilocularis J.J.S. 429 var. guinquelocularis J.J.S.
429 trochocarpoides F.v.M. 437 vandewateri Wernh. 429 vannouhuysii J.J.S. 440 wetarensis J.J.S. 432
Styracaceae 4: 49-56; 6: 976
Sulamea = Soulamea 221
Suregada Rottl. 944
Suriana L. 193, 194, 195, 196, 969 maritima L.
map, 223
Surianaceae 195, 196
Sycopsis Oliver 952 dunnii Hemsl. 952
193 aL Gaal 97
Tabernaemontana corymbosa Roxb. 315 Taeniochlaena Hook. f. 933 Talauma 228 Tamaricaceae 968 Tamaricales 63 Tapiscia 49 Tayotum nigrescens Blco 373 Telanthera bettzickiana Regel 916 manillensis Walp. 915 Terminalia L. 932 catappa 325 crassifolia Exell 922 darlingii Merr. 933 macrantha Rojo 932 polyantha Pr. 48 zollingeri Exell 933 Ternstroemia megacarpa Elm. 336 penangiana Choisy 336 Tetramyxis Gagnep. 226 Theaceae 472 Thecanthes Wikstr. 44 cornucopiae Wikstr. 46 Thibaudia coriacea (Bl.) BI. 828 cuneifolia Bl. 847 elliptica Bl. 873 floribunda Bl. 872 laurifolia Bl. 871 lucida Bl. 814 myrtoides Bl. 812 rosea Jungh. 872 singalensis Korth. ex Boerl. 762 varingiaefolia Bl. 815 Thylachium lucidum Banks ex DC. 92 Thymelaea 28 Thymelaeaceae 4: 349-365 (Go- nystyloideae); 6: 1-48, 976 subfam. Aquilarioideae 1, 3, 982 tribe Microsemmatidae 3 subfam. Gilgiodaphnoideae 3 subfam. Gonystyloideae 1, 2, 3, 976-982 subfam. Thymelaeoideae 3, 982 Thymelaeales 451 Tiliaceae 4, 203 Timeroyea Montrouz. 457 Tonsella chinensis (L.) Spreng. 419 prinoides Willd. 418
Toona sinensis (Juss.) Roem. 220 Torrenticola Domin 963 Tovariaceae 62 Toxicodendron altissimum Mill. 220 Toxina Norona 954 Trachymene Rudge 983 novoguineensis (Domin) Buw. 984 tripartita Hoogl. 983*, 984 Trapaceae 98? Trianthema triquetra Rottl. ex Willd. 915 Tricalysia tinagaoensis Elm. 23 Triceros Lour. 51 cochinchinensis (non Lour.) Mor. 56 Trichadenia 39 philippinensis Merr. 944 Trichilia 194 connaroides (W. & A.) Bent- velzen f. connaroides 226 f. glabra Bentvelzen 226 Trientalis 174 Trigoniaceae 4: 58-60 Tripterygium Hook. f. 229, 230, 231 wilfordii Hook. f. 229, 230 Triscaphis Gagnep. 49, 212, 214 kerrii Gagnep. 214 Tristellateia Thouars 960 Tristicha trifaria (Bory ex Willd.) Spreng. 963 Triumfetta 223 Trochisandra Bedd. 280 indica Bedd. 283 Trochocarpa R.Br. 423, 424, 434, 436 map subg. Pseudocyathodes Sleum. 437, 443 subg. Trochocarpa 436, 437 arfakensis (Kaneh. & Hatus.) Sleum. 443, 444* bellendenkerensis Domin 437 celebica (J.J.S.) Steen. 439 dekockii (J.J.S.) H.J.Lam 435*, 440* dispersa Sleum. 442 gjellerupii (J.J.S.) H.J.Lam 438, 439* lamii H.J.Lam 443 laurina (R.Br. ex Drude) R. Br. 423, 437, 438* learmonthiana (Gibbs) H.J. Lam 439
nubicola (Wernh.) Sleum. 442, 443
nutans (J.J.S.) H.J.Lam 435*, 441*, 442*, 443
papuana (Wright) Sleum. 443 vannouhuysii H.J.Lam 441 Tubiflorae 294, 297 Turneraceae 4: 235-238
Sept. 1972]
Index to scientific names
Turpinia Vent. 49, 51, 56 map arguta 55 borneensis (Merr. & Perry) v. d.Linden 50*, 53, 54,55, 56 brachypetala (Schltr) v.d. Linden 52, 54, 59 cochinchinensis (Lour.) Merr. 58 formosana Nakai 55 glaberrima Merr. 56 gracilis Nakai 56 grandis v.d.Linden 52, 53, 54, 55, 58 indochinensis Merr. 55 laxiflora Ridl. 53, 54, 55, 57 lucida Nakai 58 malabarica Gamble 59 montana (BI.) Kurz 50*, 52, 994, 55256 a@ genuina Kurz 55 f. arborescens Hochr. 56 f. scandens Hochr. 56 var. borneensis Merr. & Perry 56 nepalensis 52, 55, 56, 57, 58, 59 nitida Merr. & Perry 52, 54, 58 ovalifolia Elm. 50*, 53, 54, 58 pachyphylla Merr. 58 papuana Harms 59 papuana Merr. & Perry 59 parva K. & V. 56 parviflora Craib 53, 56 pentandra (Schltr) v.d.Linden 52, 54, 55, 59 pomifera (Roxb.) DC. 50*, 52, 54, 58 var. sphaerocarpa (non Hassk.) King 57, 58 robusta Craib 58 simplicifolia Merr. 52, 53, 54, 55 sp. Merr. 58 sphaerocarpa Hassk. 50*, 52, 53; 54, 57, 58, 59 var. pubescens v.d.Linden 57 stipulacea v.d.Linden 50%, SIP 52953254: 55 trifoliata Ridl. 58 unifoliata Merr. & Chun 53, 5
versteeghii Merr. 59
Typha L. 982
angustata Bory & Chaubard 983
angustifolia L. s.]. 982, 983
domingensis Pers. 982, 983
latifolia L. 983
orientalis Pr. 982, 983
Typhaceae 4: 242-244; 6: 982
Umbelliferae 4: 113-140, 595; 5: 555; 6: 194, 983-984
1019
Urophyllum streptopodium Wall. 373
Usteria 295, 296, 954
Utania G.Don 299, 301, 309 morindaefolia G.Don 311
Vacciniaceae 473 Vaccinium L. 469, 470, 471, 472, 473, 474, 741, 746, 878 sect. Bracteata Nakai 747, 791, 792 map sect. Euepigynium Schltr 791 sect. Galeopetalum (J.J.S.) Sleum. 747, 748 map, 753 sect. Neojunghuhniana (Koord.) Sleum. 747, 753 map, 785 sect. Nesococcus Copel. f. 791 sect. Oarianthe Schltr 747, 765 map sect. Pachyanthum Sieum. 747, 748 map sect. Rigiolepis (Hook. f.) J.J.S. 747, 753 map, 754 subg. Galeopetalum J.J.S. 753 absconditum J.J.S. 767, 775 acrobracteatum K.Sch. 470, 793, 796, 800, 868* acuminatissimum (non Miq.) Merr. 756 acuminatissimum Miq. 755, 761 f. acuminatissimum 761 f. ellipticum (J.J.S.) Sleum. 761, 762 f. javanicum Miq. 762 f. leptantha = leptanthum f. leptanthum (Miq.) Vuyck 764 f. marapiense (J.J.S.) Sleum. 761, 762 f. sumatranum Miq. 762 var. singalense J.J.S. 762 acuminatissimum (non Mig. s. str.) Ridl. 762 acutissimum F.v.M. 831 var. acutissimum 795, 831 var. pilosistylum Sleum. 796, 831 acutissimum (non Warb. 830 adenanthum Sleum. 868 adenopodum Sleum. 755, 765 adenotrichum Sleum. 796, 846 adenurum C.E.C.Fischer 840 agusanense Elm. 793, 807 aitapense Sleum. 795, 800, 833 albicans Sleum. 795, 832 var. albicans 797, 832 var. pseudopsammogenes Sleum. 832, 833 var. pubens Sleum. 832, 833 alvarezii Merr. 798, 800, 857 var. alvarezii 857
F.v.M.)
var. moisense Copel. f. 858 ambivalens Sleum. 796, 799, 801, 847 amblyandrum F.v.M. 768, 783 var. amblyandrum 766, 783, 784 var. maiusculum Sleum. 783 var. pungens Sleum. 783, 784* ambyandrum = amblyandrum 783 amphoterum Sleum. 793, 797, 798, 848 amplexicaule J.J.S. 794, 799, 861 amplifolium F.v.M. 748, 751 var. amplifolium 751, 752, 753 var. giganteum Sleum. 751, 752 var. oblongum Sleum. 751, 752 var. stabilipes Sleum. 751, 752 ampullaceum Sleum. 792, 804 andersonii Sleum. 755, 763 angiense Kaneh. & Hatus. 798, 800, 856 angulatum J.J.S. 798, 799, 856 angustilimbum Merr. 870 apiculatum Sleum. 800, 869 apoanum Merr. 895 apophysatum Sleum. 797, 850 appendiculatum Schltr 794, 826, 861 ardisiflora = ardisiiflorum 843 ardisiiflorum Ridl. 843 ardisioides Ridl. 843 ardisioides Wernh., non Ridl. 863 arenarium Sleum. 784 artum J.J.S. 798, 854 atrescens Sleum. 753 aucupis Sleum. 794, 822 auriculifolium Sleum. 792, 805 bancanum (non Mig. s.str.) Clarke 842 bancanum (non Mig. s:.str.) K. & G. 842 bancanum (non Miq.) Koord. 847 bancanum Midq. 796, 803, 840, 841* var. bancanum 841, 842%, 843 var. kemulense J.J.S. ex Sleum. 841, 843 var. kunstleri (K. & G.) Sleum. 841, 842 var. tenuinervium 841, 842, 843 bancanum (non Mig.) Ridl. 854 banksii Merr. 797, 849
JS.
1020
FLORA MALESIANA
[ser [fyolie®
banksii (non Merr. 1906) Merr. 850 barandanum Vidal 793, 806 var. barandanum 807 var. cagayanense Copel. f. 807
var. hutchinsonii (Merr.) Copel. f. 807 barbatum J.J.S. 768, 781 bartlettii Merr. 800, 867 benguetense Vidal 797, 801, 875 besagiense J.J.S. 796, 838 bigibbum J.J.S. 754, 761 blepharocalyx (non Schltr) Lane-Poole 876 blepharocalyx Schltr 801, 876 blumeanum Niedenzu 872 bodenii Wernh. 766, 767, 774 borneense (non W.W.Sm.) Anderson 763 borneense W.W.Sm. 754, 756 var. borneense 756 var. poianum (J.J.S.) Sleum. 756 brachycladum Sleum. 766, 770 brachygyne J.J.S. 794, 824 brachytrichum Sleum.796, 847 bracteatum Thunb. 747, 791, 793, 801 brassii Sleum. 799, 864 var. brassii 864, 866 var. madarum Sleum. 866 breviflos Ridl. 842 brevipedunculatum J.J.S. 799, 863 buxifolium Hook. f. 839 buxoides J.J.S. 777 calelanum Elm. 895 camiguinense Merr. 795, 836 capillatum Sleum. 794, 799, 866 capillipes Sleum. 754, 757 cardiophorum Sleum. 786, 789 carneolum Sleum. 831 var. carneolum 795, 831 var. nesopbilum Sleum. 795, 832 caudatifolium Merr. 765 caudatum Warb. 797, 801, 875 cavendishioides Sleum. 798, 800, 801, 858 centrocelebicum Sleum. 792, 803 var. centrocelebicum 803 var. maius Sleum. 803 ceramense Sleum. 767, 775 cercidifolium J.J.S. 794, 827 chionostomum Sleum. 776 ciliatipetalum J.J.S. 777 claoxylon J.J.S. 798, 859 clementis Merr. 795, 796, 836* coelorum Wernh. 768, 784
collivagum Sleum. 772 contractum Sleum. 794, 821 convallariiflorum J.J.S. 794, 826, 861 convexifolium J.J.S. 766, 772 cordifolium Stapf 794, $27 coriaceum Hook. f. 792, 795, 796, 838 coriaceum (Bl.) Miq. 828 cornigerum Sleum. 796, 837 corymbiferum Miq. 878 costerifolium Sleum. 796, 801, 845 costeroides Merr. 742 crassiflorum J.J.S. 767, 777 crassiflorum (non J.J.S. 1912 & 1914) J.J.S. 774 crassifolium Andr. 746 crassistylum Sleum. 799, 863 crenatifolium Sleum. 797, 849 cruentum Sleum. 798, 831, 857, 877 cryptodon Sleum. 776 culminicolum Wernh. 766, 767, 774 cumingianum Vidal 797, 798, 850 var. cumingianum 850, 851 var. igorotorum Copel. f. 851 var. marivelesense Copel. f. 851 var. pyriforme (Merr.) Co- pel. f. 851 var. tayabasense Copel. f. 850 cuneifolium (BI.) Mig. 796, 847 var. acutum Miq. 847 cyclopense J.J.S. 773 f. cyclopense 767, 773 f. glabrum Sleum. 768, 773 var. arfakense J.J.S. 773 cyrtodon Miq. 872, 873 daphniphyllum Schltr 794, 826, 861 debilescens Sleum. 797, 852 decorum Ridl. 852 decumbens J.J.S. 767, 776 dempoense Fawc. 810 dempoense (non Sp.Moore) de Voogd 811 densifolium J.J.S. 766, 769 dialypetalum J.J.S. 752*, 753 dictyoneuron Sleum. 798, 800, 858 var. dictyoneuron 798, 858 var. koébrense Sleum. 859 var. oreophilum Sleum. 859 dipladenium Sleum. 755, 763 disterigmoides Sleum. 777 dominans Sleum. 471, 793, 809 dubiosum J.J.S. 747, 793, 808 eburneum Ridl. 843
elegans Elm. 795, 829
ellipticum (Bl.) Miq. 815, 873 var. macrocalyx J.J.S. 873
elliptifolium Merr. 795, 835
endertii (J.J.S.) Masam. 758
endertii J.J.S. 795, 835
epiphythicum Merr. 793, 806
erythrinum Hook. 815
euanthum Bl. 872
eugenoides Sp.Moore 811
evanidinervium Sleum. 768, 784
eymae Sleum. 786, 790, 791*
fastigiatum Sp.Moore 872
filiforme (J.J.S.) Sleum. 754, 758
filipes Schltr 796, 844
finisterrae Schltr 766, 769
fissiflorum Sleum. 747, 748
flagellatifolium Copel. f. 755, 765
floribundum (BI.) Miq. 872
forbesii Fawe. 810
foxworthii Copel. f. 837, 847
fraternum Sleum. 793, 798, 859
gitingense Elm. 792, 803
gjellerupii J.J.S. 798, 801, 871
glabrescens K. & G. 792, 804
glandellatum Sleum. 794, 813
globosum J.J.S. 768, 778, 781 var. latifolium J.J.S. 779
goodenoughii Sleum. 795, 829
gracile J.J.S. 797, 799, 851
gracilipes Sleum. 801, 874
gracillimum J.J.S. 801, 877
grandibracteatum Schltr 793, 809
habbemae Koord. 798, 855 var. habbemae 855 var. parvifolium J.J.S. 855 var. pluriglandulosum J.J.S.
798, 855
haemanthum Sleum. 769
haematochroum Sleum. 766, 767, 770
halconense Merr. 798, 799, 860
hasseltii Miq. 854 var. sabuletorum Ridl. 854
hatamense Becc. 766, 770
hatamense (non Becc.) Koord. 782
helenae F.v.M. 879
hellwigianum Sleum. 795, 830
henrici Sleum. 754, 759
hispidulissimum Wernh. 786, 789
hooglandii Sleum. 793, 799, 862
horizontale Sleum. 796, 839, 840*
hosei Merr. 873
hutchinsonii Merr. 807
igneum J.J.S. 767, 775
Sept. 1972]
Index to scientific names
1021
igorotorumCopel.f. ex Elm.851 ilocanum Merr. 810 imbricans J.J.S. 786, 789 inconspicuum J.J.S. 768, 781 indutum Vidal 792, 793, 804 ingens Sleum. 747, 748, 749* insigne (Koord.) J.J.S. 786, 787 irigaense Merr. 798, 801, 859 jagori Warb. 800, 870 Javanicum Hook. 873 kemulense Sleum. 754, 758 keysseri Schltr ex Diels 748, 750* var. acutatum Sleum. 751 var. keysseri 750 kjellbergii J.J.S. 791, 793, 802 korinchense Ridl. 794, 811 var. korinchense 792, 811* var. losirense Sleum. 812 korthalsii (J.J.S.) Masam. 763 korthalsii Mig. 795, 828 kostermansii Sleum. 786 kunstleri K. & G. 842 kunstleri (non K. & G.) Ridl. 843 lageniforme (non J.J.S.) Ka- neh. & Hatus. 868 lageniforme J.J.S. 799, 800, 862 lanaense Merr. 745 lanceifolium (Ridl.) Sleum. 754, 755 lanceolatum (B\.) J.J.S. 761 f. ellipticum Sleum. 762 f. marapiense (J.J.S.) Sleum. 762 f. sumatrana (Mig.) Sleum. 762 lancifolium = lanceifolium ies) latissimum J.J.S. 794, 796, 821 laurifolium (BI.) Mig. 471, 746, 801, 871*, 878 var. arborescens O.K. 873 var. ellipticum (Bl.) Sleum. 872, 873 var. glanduligerum Sleum. 798, 872, 874 var. laurifolium 872, 873, 874 var. pensile (Sp.Moore) Sleum. 872, 873 var. robustum Sleum. 872, 873 var. sarawakense (Merr.) Sleum. 872, 873 var. trichodes Sleum. 872, 873 ledermannii Schitr 773 leptanthum Mig. 755, 764 f. ellipticum (J.J.S.) Sleum. 764 f. leptanthum 764 f. malayanum Sleum. 764
(Ridl.)
var. ellipticum (J.J.S.) Masam. 764 leptocladum Sleum. 799, 861 leptomorphum Sleum. 799, 866 leptospermoides J.J.S. 779, 780 f. glabrum J.J.S. 767, 768, 780 f. leptospermoides 767, 768, 779*, 780* ligustrifolium J.J.S. 800, 870 littoreum Miq. 747, 797, 854 lobbii (Ridl.) Sleum. 754, 755 loheri Merr. 741 longepedicellatum Sleum. 799, 863 longibracteatum Ridl. 804 longilingua Sleum. 753 longipes Sleum. 753 longiporum (non Schltr) Diels 876 longiporum Schltr 795, 831 longisepalum J.J.S. 786, 788 loranthifolium Ridl. 797, 853 lorentzii Koord. 782 f. lorentzii 768, 782 fi, puberultiny Jasna 768; 782 lucidum (BI.) Mig. 746, 794, 814* f. epiphyticum O.K. 814 f. terrestre O.K. 814 var. lucidum 799, 800, 814 var. micranthum Hochr. 814 var. orientale Hochr. 820 var. pumilum J.J.S. 867 var. roseitinctum Sleum. 815 var. typicum Hochr. 814 luridum Sleum. 777 luzoniense Vidal 796, 843 macbainii F.v.M. 747, 748 macgillivrayi Seem. 791 macgregorii Merr. 804 macrophyllum (J.J.S.) Sleum. 757 malacca = malaccense 801 malaccense Wight 801 var. bancanum Miq. 801 var. celebense J.J.S. 802 malacothrix Sleum. 786, 787 marginellum Sleum. 778 mearnsii Elm. 870 medinilloides Elm. 745 megalophyes Sleum. 747, 794, 825 megaphyllum Sleum. 754, 757 var. adenophorum Sleum. 757 var. megaphyllum 757 micrantherum Stapf 843 micranthum = micrantherum 843 microphyllum Reinw. ex BI.
742, 766, 768, 778 microphyllum (non Reinw. ex Bl.) F.-Vill. 812, 850 microphyllum (non Bl.) K. & G23
microphyllum (non BI.) J.J.S. 775 mindorense Rendle 778 minimiflorum Sleum. 754, 759 minusculum Sleum. 797, 852 minuticalcaratum J.J.S. 794, 797, 822 f. capillatum Sleum. 866 f. glabrum J.J.S. 822, 823* f. latifolium J.J.S. 822 f. minuticalcaratum 822, 823 miquelii Boerl. 470, 794, 810* var. atjehense Sleum. 811 var. miquelii 793, 810 mjoebergii J.J.S. 795, 834 molle J.J.S. 796, 844 var. molle 844, 845 var. mollissimum (Sleum.) Sleum. 792, 845 mollissimum Sleum. 845 monanthum Ridl. 754, 756 montis-ericae Sleum. 793, 809 moultonii Merr. 755, 763 muriculatum J.J.S. 795, 829 var. albidum J.J.S. 829 myrianthum Sleum. 765 myrsinoides Schltr 766, 772 myrtoides (Bl.) Miq. 746, 747, 793-0 1944 S12 813754821 var. B 810 var. celebicum J.J.S. 812 nitens Sleum. 800, 869 nitidum Andr. 814 nyctericoides = nycteroides 823 nycteroides Wernh. 823 oblongum Wright 752 obovalifolium Sleum. 824 obversum Miq. 815 oranjense J.J.S. 777, 778 var. marginellum (Sleum.) Sleum. 767, 778 var. oranjense 767, 777, 778 oreites Sleum. 786, 788 oreomyrtus Sleum. 767, 777 otophyllum Sleum. 794, 827, 861 pachydermum Stapf 795, 834* palawanense Merr. 795, 837 var. foxworthii (Copel. f.) Sleum. 837 var. palawanense 837 paludicolum Sleum. 794, 814 papuanum J.J.S. 868 paradisearum Becc. 794, 824* parvibaccatum J.J.S. 811 parvulifolium (non F.v.M.) Diels 769 parvulifolium F.v.M. 766, 771
1022
FLORA MALESIANA
(ser. I, vol. 68
pauciflorum Fletch. 753 pensile Sp. Moore 873 perakense Ridl. 852 perrigidum Elm. 798, 855 philippinense Warb. 793, 808 phillyreoides Sleum. 799, 861 piceifolium Wernh. 767, 774 pilosiflorum J.J.S. 768 pilosilobum J.J.S. 474, 799, 867 piperifolium Sleum. 755, 762 piroliflorum J.J.S. 826 platyphyllum Merr. 793, 810 polvanthum O.K. 872 var. bicolor O.K. 872 var. viridiflorum O.K. 872 profusum J.J.S. 824 prostratum Sleum. 767, 768, 785 psammogenes Sleum. 795, 830 pseudocaudatum Sleum. 797, 801, 874 psittacobium Sleum. 824 pubicarpum Ridl. 801 pugionifolium Sleum. 767, 774 pullei J.J.S. 766, 768, 782 pyriforme Merr. 851 quinquefidum J.J.S. 794, 823 var. oranjense J.J.S. 824 var. quinquefidum 823, 824 rariflorum Schltr 773 retevenium = retivenium 856 reticulato-venosum Sleum. 799, 866 retivenium Sleum. 798, 856 retusifolium J.J.S. 799, 864 ridleyi Sleum. 753 rigidifolium Sleum. 796, 799, 846 rizalense Merr. 810 robustum Ridl. 873 rollinsoni Hook. 814 roseiflorum J.J.S. 798, 799, 860 rubroviolaceum Sleum. 801, 877 sabuletorum Ridl. 854 sabuletrum = sabuletorum 854 salicifolium (J.J.S.) Masam. 757 sanquineum Schltr 768, 781 sarawakense Merr. 873 scandens Schltr 797, 853 schimperi Koord. 815, 820 schlechterianum Sleum. 785, 786, 787 schoddei Sleum. 799, 862 schultzei Schltr 767, 773 sclerophyllum Sleum. 799, 867 scortechinii K. & G. 793, 807 scyphocalyx Sleum. 766, 771 sessiliflorum Schltr 769 simulans Sleum. 795, 833
var. leptopodum Sleum: 834 var. simulans 833 sororium J.J.S. 767, 776 sorsogonense Elm. 8C4 sp. Koord. 774, 815 sp. Merr. 844 sp. Rappard 874 sp. Stapf 859, 861 sp. Vidal 812, 850, 875 spaniotrichum Sleum. 786, 790 sparsicapillum Sleum. 766, 769 sparsum Sleum. 801, 876 stabilipes Sleum. 752 stapfianum Sleum. 793, 796, 838, 839 var. minus Sleum. 839 var. stapfianum 839 steinii Sleum. 801, 876 stellae-montis Sleum. 799, 800, 862 stenanthum Sleum. 796, 843 stenolobum Schltr 794, 825 striicaule Sleum. 471, 876 var. adenodes Sleum. 798, 877 var. pubiflorum Sleum. 796, 877 var. striicaule 801, 876, 877 subobovatum Fletcher 852 subulisepalum J.J.S. 786, 790 sulcatum Ridl. 754, 758 suluense Copel. f. 840 summifaucis Sleum. 796, 845 sumatranum Jack 877 sylvaticum Elm. 800, 870 taxifolium Sleum. 766, 772 tenerellum Sleum. 754, 760 tentaculatum J.J.S. 795, 828 tenuipes Merr. 796, 844 teysmannii {non (Bl.) Miq.] Koord. 873 teysmannil Miq. 872 teysmannii (non Miq.) Rid. 852 thibaudifolium Wernh. 793, 806 tiariforme J.J.S. 868 timonioides Wernh. 786, 788 timorense Fawc. 792, 793, 796, 797, 802 var. denticulatum Fawc. 802 tomicipes J.J.S. 796, 800, 869 torricellense Schltr 868 trichocarpum Sleum. 792, 805 tubiflorum J.J.S. 797, 851 turbinatum Merr. 857 turfosum Sleum. 799, 801, 874 uniflorum J.J.S. 754, 757 urnigerum Sleum. 779 uroglossum Sleum. 754, 759, 760* urophyllum Merr. 753 varingiaefolium (BI.) Miq.
470, 471, 746, 794, 802, 815, 816*, 817*, 818, 819*, 820* f. parvifolia Miq. 815 f. sublanceolata Miq. 815 var. angustifolium O.K. 815 var. calcaratum Sleum. 793, 815, 820 var. erythrinum (Hook.) O.K. 815 var. orientale Sleum. 815, 820 var. pilosiusculum Hochr. 815 var. racemosum Hochr. 815 var. typicum Hochr. 815 var. varingiaefolium 815, 816, 821 varingiaefolium [non (BI.) Miq.] Miq. 845 varingiaefolium [non (BI.) Miq.] Vidal 812 varingifolium = varingiaefo- lium 815 versteegil Koord. 766, 771 vidalii (non Merr. & Rolfe) H.J.Lam & Holth. 850 vidalii Merr. & Rolfe 797, 848 villarit Vidal 812 villosiflorum J.J.S. 766, 768 viridiflorum J.J.S. 795, 833 viscifolium K. & G. 800, 852, 853 var. bicalcaratum Sleum. 853 var. minus K. & G. 852 var. viscifolium 797, 853 vonroemeri Koord. 768, 780 warburgii Sleum. 794, 821 whiteanum Sleum. 767, 775, 778 whitfordii Merr. 768, 782 wisselianum Sleum. 792, 805 wollastonii Wernh. 768, 785 wondiwoiense J.J.S. 767, 772 woodianum Copel. f. 797, 850 wrayi Ridl. 843 xerampelinum Sleum. 799, 864, 865* zollingeri Miq. 872
(Hochr.)
Valeriana chinensis L. 455 Valerianaceae 4: 253-254 Valli-Modagam Rheede 326, 328 Vareca moluccana Roxb. 963 Velleia J.E.Smith 950
spathulata R.Br. 950
Ventilago 280
dichotoma (Blco) Merr. 284
Vernonia arborea 152 Vertifolia rubra Rumph. 291 Viburnum amplificatum Kern
929, 930*
clemensae Kern 929 glaberrimum Merr. 929 hispidulum Kern 929 junghuhnii Miq. 929
Sept. 1972]
lutescens BI. 929
platyphyllum Merr. 929
vernicosum Gibbs 929 Vieillardia Brongn. & Gris 457 Viereya = Vireya 480 Violaceae 237, 963 Violales 63 Vireya Bl. 480
alba Bl. 539
celebica Bl. 615
javanica Bl. 624
retusa Bl. 482
tubiflora Bl. 532 Vitmannia Vahl 199
elliptica Vahl 199
lucida Steud. 201
polyandra Steud. 203 Voacanga sp. 369
Wahlenbergia Schrad. ex Roth 1072 109) 110; WI 125 3: 128 agrestis DC. 115 bivalvis Merr. 117, 118 candollei Tuyn 114 confusa Merr. & Perry 107,
mise i4. 115* consimilis Lothian 118 dehiscens (Roxb.) DC. 115 erecta (Roth ex R. & S.) Tuyn
113, 114* eurycarpa Domin 115 gloriosa Lothian 118 gracilenta Lothian 116, 118 gracilis (Schrad.) DC. 115,
116, 118
var. hirsuta Jungh. 115
var. vincaeflora DC. 118 gracilis E.Mey., nomen 117 hirsuta Steud., nomen 113 hirsuta (Edgew.) Tuyn 113 hookeri (Clarke) Tuyn 107,
113, 114* indica DC. 115 lavandulaefolia DC. 115 marginata (Thunb.) DC. 107,
MMOs iss 10S; 1l6*, 117*
map
var. grandiflora Tuyn 113,
118 subvar. trichogyna (Stearn) Tuyn 118
multicaulis Bth. 115
paniculata (Thunb.) DC. 114
perotifolia W. & A. 113
quadrifida (R.Br.) DC. 115, 118
sieberi DC. 115 simplicicaulis de Vriese 115 trichogyna Stearn 118 vincaeflora (Vent.) Decne 118 Weigelia coraeensis Thunb. 929 fallax Mig. 929 Wendlandia 152 Wickstroemia = 28 Wikstroemia Endl. 2, 3, 4, 5, 28 subg. Chamaejasme (Amman) Domke 28 subg. (Eu)wikstroemia 28 sect. Euwikstroemia Meisn.
Wikstroemia
acuminata Merr. 31
amplifolia (Schltr) Domke 35
androsaemifolia Decne 2, 29*, 30) 325338
androsaemifolia Hand.-Mazz. 34
angustissima Mertr. 30
aurantiaca 4
australis Endl. 28
brachyantha Merr. 2, 29*, 30, 31
calva Back. 32, 33
candolleana Meisn. 33
candolleana (non Ridl. 32
candollet = candolleana 32
chamaejasme (L.) Domke 28
clementis Merr. 31
crassifolia Merr. ex Domke 31
fenicis Merr. 33
indica (L.) C.A.Mey. 2, 3, 4, 30, 34 var. viridiflora Hook. f. 35
junghuhniana = junghuhnii 32
Junghuhnii (non Miq.) K. & V. 32
Junghuhnii Migq. 32, 33, 34
lanceolata Merr. 30
linearifolia Elm. 35
longifolia Lecomte 33
meyeniana Warb. 30, 33
nutans Champ. 33
Meisn.)
Index to scientific names
1023 ovata C.A.Mey. ex Meisn. 30, 31, 32 map ovata (non C.A.Mey.) Vidal 35 pachyphylla Merr. 35 polyantha Merr. 30, 32 pulgarensis Elm. 35 ridleyi Gamble 30, 34 ridleyi (non Gamble) Gibbs 32 spanoghii Decne 33, 34 subcoriacea Merr. 35 tenuiramis Miq. 29*, 30, 31 venosa Merr. & Perry 30, 33 viridiflora Meisn. 35 Willughbeia auriculata Spreng. 327, 328 elliptica Spreng. 303 fragrans Spreng. 307 racemosa Spreng. 311 volubilis Spreng. 311 Wimmeria Schlechtend. 228, 229, 231 Wirtgenia Andres 474, 669, 943 malayana (Scort. in Hook. f.) Andres 669 Wrightia laniti (Blco) Merr. 373
Xanthophyllum subglobosum Elm. 395 var. longifolium Elm. 395 Xanthostemon brassii Merr. & Perry 428 Xanthoxylum = 56 Xolisma Raf. 674 ovalifolia (Wall.) Rehd. 675 Xylonymus Kalkman 231, 232, 243, 391, 392, 930 versteeghii Kalkman 245 Xylosma luzonense (Presl) Clos 944 palawanense Mendoza 944 Xyridaceae 4: 366-376, 598; 5: 557
Zannichellia 157 Zannichelliaceae 157 Zanthoxylum 207, 209 montanum BI. 56 serrulatum Bl. 56 Zygophyllaceae 4: 64
469, 470,
Zanthoxylum
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: bine! ! | ce ns *: ye Volume 1. Cyclo 1 collector , Collec ?: 0. pp. cl i 639 at : oe (in preparation) ‘ Volume 3. ian plant geo; | : “(In pr eparation) ak Volume 4: General chapters and revisions. 1948-1954 . pp. ccix + 63] Volume 5. Bibliography, specific delimitation, & re visions. 1955-1958. pp. cccxlii + 596 | Volume 6. Systematic revisions. 6 parts, 1960-1972. pp. 20 +- 1023. Volume 7. Systematic revisions. Part 1. 1971. pp. 263
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