FLORA AN ANNOTATED LIST OF THE VASCULAR PLANTS OF THE COUNTY OF HALIBURTON, ONTARIO ELEANOR G. SKELTON AND EMERSON W. SKELTON HALIBURTON FLORA AN ANNOTATED LIST OF THE VASCULAR PLANTS OF THE COUNTY OF HALIBURTON, ONTARIO ELEANOR G. SKELTON AND EMERSON W. SKELTON ROM LIFE SCIENCES MISCELLANEOUS PUBLICATIONS ROYAL ONTARIO MUSEUM TORONTO ROYAL ONTARIO MUSEUM PUBLICATIONS IN LIFE SCIENCES The Royal Ontario Museum publishes three series in the Life Sciences: contributions: a numbered series of original scientific publications. occasional papers: a numbered series of original scientific publications, primarily short and of taxonomic significance. miscellaneous publications: an unnumbered series on a variety of subjects. All manuscripts considered for publication are subject to the scrutiny and editorial policies of the Life Sciences Editorial Board, and to independent refereeing by two or more persons, other than Museum staff, who are authorities in the particular field involved. LIFE SCIENCES EDITORIAL BOARD Senior editor: J. H. McAndrews Editor: M. D. Engstrom Editor: G. B. Wiggins External editor: C. S. Churcher Manuscript editor: C. S. Churcher Production editor: J. E. Hawken The Royal Ontario Museum is an agency of the Ontario Ministry of Culture and Communications. Canadian Cataloguing in Publication Data Skelton, Eleanor G., 1908-1989 Haliburton Flora : an annotated list of the vascular plants of the county of Haliburton, Ontario cs<\ (Life Sciences miscellaneous publications, ISSN 0082-5093) Includes bibliographical references. ISBN 0-88854-353-0 1. Botany - Ontario - Haliburton (County). I. Skelton, Emerson W., 1907- . II. Royal Ontario Museum. III. Title. IV. Series. QK203.05S57 1990 581.971361 C90-093701-7 Publication date: 1 February 1991 ISBN 0-88854-353-0 ISSN 0082-5093 © Royal Ontario Museum, 1991 100 Queen's Park, Toronto, Canada M5S 2C6 Typesetting by Trigraph Inc. Printed and bound in Hong Kong by C & C Offset Printing Company Limited Colour separation by Goody Color Separation (Scanner) Limited Cover: Yellow water-lilies, Nuphar variegata, Haliburton, 1977 (Photo: Emerson W. Skelton) CONTENTS Foreword 5 Introduction 9 Features of Haliburton Significant to the Flora 9 Topography, Drainage, Roads 9 Rocks and Glacial Deposits 10 Soils 14 Climate 15 Forests 15 Wetlands 16 Open-water Habitats 19 Other Habitats 19 Methods 20 Collecting and Collections 20 Territory Covered 20 Identification and Nomenclature 20 Voucher Specimens 21 Abundance Ratings 21 Acronyms, Abbreviations, and Symbols 22 Herbarium Acronyms 22 Township Abbreviations 22 Other Abbreviations and Symbols 22 Maps Used in Establishing Locations 23 Annotated List of the Vascular Plants of the County of Haliburton Fern Allies 25 Ferns 27 Conifers 31 Flowering Plants 32 Monocotyledons 32 Dicotyledons 61 Acknowledgements 121 Literature Cited 123 Additional Publications Consulted in Identification 125 Index to Common Names 127 Index to Scientific Names 137 25 LIST OF FIGURES Fig. 1 Location of the county of Haliburton in southern Ontario, and township abbreviations used in the Annotated List 8 Fig. 2 Main highways and larger lakes of the county of Haliburton 1 1 Fig. 3 Postglacial topography of the county of Haliburton 13 FOREWORD It was with surprise and sorrow that I learned of the passing of Mrs. Eleanor Skelton on 1 October 1989. My memories of Eleanor will be of an extraordi- nary woman, an impressive botanist, a devoted wife, and a good friend. Born and raised in Toronto, she undoubtedly developed a love of nature and the outdoors during her childhood and youth when she spent her summers at the family cottage on Gull Lake in Haliburton. She pursued her education through the university level and graduated from the University of Toronto with a General Arts degree with emphasis on biology. Eleanor was ready to accept any challenge. Having decided that biology, especially the study offish, sparked her interest, Eleanor made inroads into this male-dominated field by gaining employment with the Ontario Fisheries Research Lab from 1929 to 1932. In 1931 she deposited her first plant collec- tion in the herbarium at the University of Toronto (TRT). It is difficult to talk about Eleanor without mentioning her husband Emerson. They met while they were students at the university and were mar- ried in 1931. Their mutual interest in natural history, and in fact in all facets of science, and their enjoyment of the outdoors continued to be important throughout their married life. During the years of Emerson's career as a chemical engineer he became an accomplished photographer of wildflowers. Emerson retired in 1975 and the couple embarked on a special project. I first met Eleanor and Emerson in 1976 when they were auditing a course in plant taxonomy at the University of Toronto Botany Department and were bringing in plant specimens to identify from their summer holiday in Hali- burton. With the enthusiastic support of Dale Hoy, at the time curatorial assistant at TRT, they began their project of documenting the plants of the county of Haliburton. For eight years, from 1976 to 1983, the Skeltons rented a cottage in Haliburton, sometimes departing from the city as early as April and returning to Toronto at Thanksgiving. They conducted field excursions, collecting and carefully pressing plant specimens throughout the growing sea- sons. During the winter they identified and painstakingly mounted the collec- tions, providing TRT with some of that herbarium's finest specimens. They would mark particular trees or shrubs— in order to collect flowering, fruiting, and mature leaf material from the same individual— and mount all the stages on a single sheet, sometimes even including a close-up photograph of the flower or the habitat. In all they donated more than 2100 herbarium sheets representing more than 930 taxa to the TRT. The neatness and completeness of their specimens are outstanding. The collections are a real asset to the her- barium and are often consulted by artists and researchers; this shows that nonprofessional or "amateur" botanists can make very worthwhile contribu- tions to science. Emerson became the grass and sedge expert of the team, while Eleanor was the aquatics specialist. Aquatic plant communities always commanded a very special place in Eleanor's heart, and she spent innumerable hours in the canoe with Emerson and investigated the submerged, floating, and emergent vegetation of the many rivers and lakes in Haliburton. She made some out- standing discoveries, such as locating a colony of the provincially rare blad- derwort Utricularia geminiscapa and an especially large colony of another bladderwort U. gibba. Her interest in the effects of acidic precipitation on the vegetation of Haliburton— in particular on the aquatic systems— led her to confer with the Ontario Ministry of Environment staff at Dorset, Ontario, and eventually to contribute lists of plants present at specific lakes in the region to the Acidic Precipitation in Ontario Study. During the years the Skeltons spent collecting their specimens from Hali- burton, Emerson pursued another project. He was involved in annotating and recording specimens of the James H. White Collection, which had been acquired from the University of Toronto Faculty of Forestry. These speci- mens, from the early 1900's, were donated to TRT in 1976 when their histori- cal significance became apparent. Once the Skeltons' own collections from Haliburton were completed to their satisfaction, from 1983 to 1986 they took on the mammoth task of writ- ing up their findings and incorporating geologic, topographic, and soil infor- mation. This phase also included checking the herbaria in Ottawa and in the Algonquin Park Museum for additional Haliburton records. Their intention was to produce a scientifically valuable work that would be straightforward enough to be used by naturalists. Because of their contributions to the herbar- ium, the Skeltons became field associates in the Department of Botany of the Royal Ontario Museum in early 1987. During the final writing stages, Emerson's health began failing, leaving the onus to fall on Eleanor. She rose to the occasion and completed the man- uscript, which was then submitted to the Museum for publication. The review process, both internal and external, resulted in further changes that Eleanor ably dealt with right up until the summer of 1989, with help from the Museum's Department of Botany. In the fall of 1988 the Skeltons had moved to Owen Sound. In the follow- ing spring they began exploring their new surroundings and, when I visited that August, even sent specimens back with me to the herbarium. During my visit, Eleanor joined me on a brief trip to collect fruiting specimens of Crataegus douglasii south of Wiarton. Although obviously in pain, she showed undaunted interest as she searched the roadsides for this unusual black-fruited hawthorn. It would have been very satisfying for Eleanor to have been able to browse through Haliburton Flora. Even though this cannot be so, her work in print will be a fitting tribute to this remarkable person and to her husband, both of whom I am very proud to have known. Their retirement project has proved to be a very valuable scientific contribution. Sheila McKay-Kuja Department of Botany University of Toronto December 1989 LondonV/ \ ■ ^2 \ /^^J] f."\ \JK Windsor >' )^^\r ?^L**r LAKE ^,« ^''"ewe / \