€alumUa WLnihzvgit? \^>\^ in tfic Cit j> of iSeiu ^or fe c^d>\ College of ffl&psitwnsi anb burgeons Reference Htbrarp \ HANDBOOK OF ANATOMY BEING A COMPLETE COMPEND OF ANATOMY INCLUDING THE ANATOMY OF THE VISCERA, A CHAPTER ON DENTAL ANATOMY, NUMEROUS TABLES, AND INCORPO- RATING THE NEWER NOMENCLATURE ADOPTED BY THE GERMAN ANATOMICAL SOCIETY, GENERALLY DESIGNATED THE BASLE NOMENCLATURE, OR BNA. BY JAMES K. YOUNG, M.D., F.A.O.S. Professor of Orthopaedic Surgery, Philadelphia Polyclinic; Associate Professor of Orthopaedic Surgery, University of Pennsylvania; Orthopaedic Surgeon to the Philadelphia General Hospital: Fellow of the College of Physicians, of Philadelphia; Fellow of the Philadelphia Academy of Surgery; Fellow of the American Orthopaedic Association; Member of the American Medical Association, etc., etc FIFTH EDITION, REVISED AND ENLARGED WITH 154 ENGRAVINGS, SOME IN COLORS PHILADELPHIA A. DAVIS COMPANY, Publishers English Depot Stanley Phillips, London 1918 Copyright, 1889 Copyright, 1905 Copyright, 1909 Copyright, 1913 Copyright, 1917 BY F. A. DAVIS COMPANY Copyright, Great Britain. All Rights Reserved Philadelphia, Pa., U. S. A. Press of F. A. Davis Company 1914-1916 Cherry Street PREFACE TO FIFTH EDITION. The demand for a new edition of the author's Handbook of Anatomy has afforded him an opportunity to revise certain portions of the volume, made necessary by recent advances in anatomical studies since the publication of the fourth edition. In order to enhance still further any value that the work may possess, there have been some eliminations, many altera- tions and a large number of additions. These changes, together with the replacement of the older cuts in the chapter on Osteol- ogy by clearer and far superior illustrations, that comprised an osteologica] atlas in former editions, necessitated a recasting of the entire hook. A special "chapter devoted to Dental Anatomy, comprising the Anatomy of the Face. Teeth and daws, properly illustrated, lias been prepared by Joseph L. Appleton, dr.. B.S., D.D.S., of The Thomas W. Evans .Museum and Institute of Dentistry, I Fniversity of Pennsylvania. In order to make the work thoroughly accurate and mod- ern, it wa- deemed advisable to incorporate throughout the volume the Basle nomenclature or UNA as formulated and adopted by the German Anatomical Society. Each anatomical name has been critically examined and its UNA equivalent added. This is, perhaps, the only medium-sized anatomical work that fully include- the newer terminology. This arduous labor ha- been successfully prosecuted by Dr. Samuel Lewald, medical and historical writer, without whose valuable assistance this portion of the work could not haw been accomplished. The author trusts that these numerous changes will meet with the approval of professors and teachers in the leading medical and dental colleges, and that it may appeal to the -tudeiit body in general, for whom this labor has been under taken. d. K. Y. 222 S... Sixteenth Street. run PREFACE TO FOURTH EDITION. Although for over twenty years the book has been in con- stant use by large numbers of students and approved by professors and teachers in leading medical colleges and in Nurses' Training Schools generally, comparatively few errors or omissions of im- portance have been discovered; but in the present edition these have all been noted and corrected. The appreciative thanks of the author are extended to Dr. A. C. Pole, of Baltimore, for several pertinent suggestions in the preparation of this edition. J. K. Y. (iv) PREFACE TO THIRD EDITION. In the preparation of this new edition, the author has taken advantage of the opportunity to correct the few clerical errors which must necessarily lie round in a comprehensive though condensed work upon Anatomy, and to make such changes as have become necessary through the advances of the science since the publication of the last edition. In the section upon the Nervous System the greatest changes will be found, the first part having been rewritten. In the description of the Muscular System, the regional grouping has been retained, because it is more practical for the student. Attention is directed to the original colored plates of the Arterial System which offer the most complete classification of all the arteries in the smallest space, ami also to the original diagrams of the sensorj tracts from spinal cord to brain, and the motor tracts from brain to spinal cord. The author desires to express his gratitude to the student body of America tor their continued confidence in this early literary effort of his, and express the hope that the present edit ion may he as well received as the others. J. K. V. (v) PREFACE TO SECOND EDITION. The demand for a new edition has afforded an opportunity to thoroughly revise the text and illustrations, and to add such changes of description and nomenclature as have been intro- duced into modern methods of teaching. Whenever it seemed necessary or advisable sections have been entirely rewritten, but the same attention to conciseness and accuracy has been observed as in the former edition. The size of the pages has been considerably enlarged and increased in number, and the illustrations increased from 76 to 171. Many improved cuts replace those used in the former edi- tion and full-page engraved plates have been inserted in the text where they will be most serviceable. In its preparation, the last editions of both Morris and Gray have been freely consulted, so that it can be Used with either of these standard works. As in the former edition, although prepared particularly for students, sufficient descriptive matter has been added to render it valuable to the busy practitioner, particularly in the sections upon the Yiscera, Special Senses, Vascular System, the Nervous System, and Surgical Anatomy. Special attention has been given to the anatomy of the brain and nervous system, which will be found to compare favorably with the descriptions in larger works. The author has been much gratified to observe the tenacity with which students and physicians have retained the work for reference. The author would acknowledge his obligation to Dr. Eichard F. Gerlach, who has distinguished himself by his careful and accurate work upon Deaver's "Surgical Anatomy," for valuable assistance upon the revision. J. K. Y. (vi) PREFACE TO FIRST EDITION. The object of this little book is. as its title indicates, t<> furnish a concise though complete synopsis of human anatomy for the use of students of medicine and others. Appreciating, from a personal contact with students, the limited time at their disposal and the unlimited amount of medical material to be digested, the author has endeavored, by well-selected wood-cuts, typographical arrangement, and numer- ous tables, to facilitate the acquisition of a subject as difficult as it is essential, and elegance of diction has therefore been sacri- ficed to conciseness and accuracy. In its preparation the last edition of Gray's "Anatomy" [edited by Keen], as the work most largely employed, has been selected as the standard, but Leidy, Quain, Allen, Holden, Eenle, Schaffer, Klein, Brown Aids, and others, have been freely used, and on the special subjects Link, Spiegelberg, Savage, Sehroeder, Budin, Treve's "Surgical Applied Anatomy." and the "Ameri- can System of Dentistry" have been consulted. Although pre- pared particularly for students, sufficient descriptive matter has been added to render it valuable to the busy practitioner, par- ticularly the section- on the Viscera, Special Senses, Vascular System, ami Surgical Anatomy. The aim throughout has been to make it as thoroughly complete and accurate as possible, and at the same time readily accessible for reference or study. The writer would acknowledge hi- obligations to Messrs. S. X. Durborow and C. W. Holly, medical -Indents at the Uni- versity, for assistance in the correction of proof. .1. K. V. (vii) CONTENTS. PAGE T NTRODUCTORY 1 ( >STEOLOGY 3 The Frontal Bone 8 Parietals 10 The < tecipital Bone 12 Temporals 15 The Sphenoid 20 The Ethmoid 24 The Lachrymal 26 The Malar 27 The Superior Maxilla 29 Palate Bones •'■- The Inferior Turbinated Bonea ;{-l The Vomer 35 The Inferior .Maxilla :!:> The Sutures and Fontanelles 39 Fossae of skull 40 The Orbits 42 Foramina at the Base of the Skull 46 The Sternum •'- The Ribs •v- The Spine 5fl The Upper Extremity (i:i The Ann ~_[) The Forearm '- The Hand 7,i The Lower Extremity sl The Thirii 86 Th<- Patella sn The Log w The Fool n:< Table of Ossification " Abticulatioks uid Ligaments In| The Thors i "'!l The Pelvis "- x CONTENTS. PAGE The Muscular System 128 Muscles of the Head 129 Muscles of the Neck 133 Vertebral Region 136 Muscles of the Back 138 Muscles of the Abdomen 143 Muscles of the Upper Extremity 147 Muscles of the Forearm 140 Muscles of the Hand 153 Muscles of the Lower Extremity 155 Muscles of the Leg 160 Muscles of the Foot 163 The Fascias 165 The Heart and Vascular System 170 The Circulation of the Blood 176 The Arterial System 178 Table of the Arterial System 202 The Venous System 209 The Lymphatic System 218 The Alimentary Apparatus 223 The Teeth 224 The Pharynx 230 The (Esophagus 231 The Abdomen 232 The Peritoneum 235 The Stomach 238 The. Small Intestine 240 The Large Intestine 242 The Pancreas 245 The Liver 246 The Ductless Glands '. 250 Vocal and Respiratory Apparatus 254 The Larynx 254 The Trachea and Bronchi 259 The Lungs 261 The Pleura? 264 The Mediastinum 264 The Genito-Urinary' Apparatus 266 The Kidneys 266 The Bladder 270 Male Organs . . 273 Female Organs ; 283 The Mammary Gland 292 CONTENTS. \i PAGE The Nervous System 294 The Brain 298 The Cranial Nerves ;;_»:; The Spinal ( 'ord 332 The Spinal Nerves :;.!2 Table of Spinal Nerves :;4:: The Sympathetic Nervous System 350 Organs of Special Sense 355 The Nose 355 The Eye 356 Appendages <>i the Eye ■ 372 The Ear 375 Tlic Tongue •. 388 Tlic Skin and Appendag ■ • 389 Surgical Anatomy :>!>l Triangles <>f the Neck .' 391 Triangle in Front of Elbow-joint :!'.i4 Scarpa's Triangle 394 Axilla 395 Anatomy of I ternia 396 [schio-rectal Region and Perineum 4o-_! Anatomy of Lateral Lithotomy 407 Dental Anatomy 4i)'.) Index 42;; ILLUSTRATIONS. FIG. PAGE 1. Frontal bone, outer surface 7 2. Frontal bone, inner surface 9 3. Parietal bone, outer surface 10 -1. Parietal bone, inner surface 11 5. Occipital bone, outer surface 12 u. Occipital bone, inner surface 13 7. Left temporal bone, outer surface 15 8. Left temporal bone, inner surface 16 9. Left temporal bone, inferior surface of the petrous portion 17 10. Section of the temporal bone through the middle car, etc 19 11. Sphenoid bone, upper surface 21 12. Sphenoid bone, anterior surface 22 13. Ethmoid bone 25 14. Lachrymal bone 26 15. Xasal bone 27 16. Malar bones 2S 17. Superior maxillary bone 29 18. Superior maxillary, inner surface ' 30 19. Palate bone (viewed posteriorly) 32 20. Palate bone (inner surface) 32 21. Inferior turbinated bones 34 22. Vomer 35 23. Inferior maxillary bone 3G 24. Left half of inferior maxillary bone, inner surface 37 25. The hvoid bone 38 26. Fossae of skull 41 27. Facial portion of skull 43 28. Nasal meatuses 45 26 I'.ase of skull, external surface 47 30. Floor of the cranium 49 31. Sternum and costal cartilages 51 32. The peculiar ribs 53 33. The vertebral column, or spine 55 34. A dorsal vertebra 56 35. A cervical vertebra 57 36. The atlas 57 37. The axis 58 38. Seventh cervical, or vertebra prominens 58 39. Peculiar dorsal vertebra? 59 40. A lumbar vertebra , 60 41. The sacrum, anterior surface 61 42. The sacrum, posterior surface 62 43. The left clavicle, upper surface 64 41. The left clavicle, under surface 65 45. The scapula, anterior surface 66 46. The scapula, posterior surface 67 47. The humerus, anterior surface 69 is. The humerus, posterior surface 69 !!>. Flna and radius, anterior aspect 73 50. Radius and ulna, posterior aspect 75 51. Carpus, metacarpus, and phalanges, dorsal surface 77 Carpus, metacarpus, and phalanges, palmar surface 79 innominatum, outer -'nine. 82 54. Ok innominatum, Inner surface 83 ."..",. The femur, anterior surface 87 56. The femur, posterior surface 87 .".7. The tibia and fibula, anterior surfac 91 58. The tibia and fibula, posterior Burfaci 91 50. The tarsus, metatarsus, and phalanges. Mi Si lirfai 95 60. The tarsus, metatarsus, and phalanges, plantai surface !<7 (Xi i) x}v ILLUSTRATIONS. FIG. PAGE 61. The thorax Ill 62. The pelvis 112 63. The shoulder 115 64. The elbow 117 65. The knee 123 66. The knee 123 67. Muscles of the back 139 68. Muscles of the abdomen 144 69. The diaphragm 145 70. Muscles of the shoulder and arm 149 71. Triceps muscles 149 72. Posterior superficial muscles of the forearm 151 73. Muscles of the anterior femoral region 157 74. Muscles of the posterior femoral region 157 75. Anterior muscles of the leg 161 76. Posterior muscles of the leg 161 77. Right side of heart 171 78. Diagram of the foetal circulation 177 79. Arteries of the face 182 80. Internal maxillary artery 184 81. The arteries of the head and neck. (Colored) Pacing 184 82. The arteries of the upper extremity. (Colored) Facing 186 83. Abdominal aorta 195 84. The arteries of the trunk. (Colored) Facing 196 85. The arteries of the lower extremity. (Colored) Facing 198 86. Temporary teeth 226 87. Permanent teeth 227 88. Pharynx, laid open from behind 231 89. Regions of the abdomen 232 90. Abdominal viscera 235 91. The peritoneum 236 92. The liver 247 93. Transverse section of lobules of liver 24S 94. Vertical section of larynx •. 256 95. Heart and lungs 263 96. Longitudinal section of kidney : ■ 267 97. Diagram of the course of the uriniferous tubules 269 98. Male organs of generation 275 99. The testicles and epididymis 277 100. Vertical section of testicle 279 101. Internal female genitals 283 102. Female organs of generation ' 285 103. Section of an ovary 287 104. External female genitals 289 105. Surface anatomy of the myelencephalon 300 106. Left side of human cerebrum 303 107. Convolutions and fissures of the median and tentorial surfaces of the right cerebral hemisphere 305 108. Inferior aspect of cerebral hemisphere 307 ' 109. The lateral ventricles and choroid plexus 309 110. Mesial section of brain and brain stem 313 111. Interior of mesencephalon 317 112. Diagram of sensory tracts from spinal cord to brain 319 113. Diagram of motor tracts from brain to spinal cord 321 114. Scheme of the nuclei and root-fibers of the cranial nerves 323 115. Base of brain and cranial nerves 324 116. Optic, oculomotor, trachlear, and trifacial nerves Facing 324 117. Facial and auditory nerves 327 118. Glossopharyngeal, pneumogastric, spinal accessory, and hypoglossal nei-ves 329 119. Cervical plexus and branches 333 120. Section of the spinal cord and membranes .... 334 121. Brachial plexus and branches " 335 122. Lumbar plexus and branches Facing 338 123. Sacral plexus and branches Facing 342 124. The sympathetic system Facing 350 125. The olfactory nerves and nerves of common sensation to the nose 355 126. Nasal meatuses and accessory cavities 357 127. Inner wall of orbit and adjacent parts 358 128. Horizontal section of right orbit 359 129. Anteroposterior section of eyeball 360 130. Vessels of the eye 363 ILLUSTRATIONS. xv FIG. PACE 131. The structure of the human retina 365 132. Diagram of the retinal vessels 366 133. The orbital muscles :',Tn 1114. The ophthalmic artery and branches ::71 135. The lachrymal apparatus 374 136, External, middle, and internal ear 377 1'. The auditory ossicles 381 138. Organ of corti ::s) 139. Triangles of the neck 393 140. The axilla 395 141. Dissection of the inguinal canal ::;i7 142. Deep femoral region — the femoral vessels, etc. (McOrath) 399 143. Male perineum, superficial dissection 402 144. Male perineum, deep dissection in:! 1 15. Female perineum 405 146. Maxillary and mandibular process of first visceral arch and fronto- nasal process 409 147. Sutures of the hard palate 410 148. Four mandibles ranging from birth to eighteen months 411 149. Muscles of expression 412 150. Tooth development 413 151. Tooth development 414 152. Root growth 417 153. Occlusal surfaces of teeth 420 154. Cross section through roots of teeth in xitu 421 Chart of cranial nerves Facing 422 INTRODUCTORY. The term ''anatomy/' derived from the Greek dva, "through," and T€/Ave. cranium 8, face 14, ribs, sternum, and os hyoideum 26, upper extremity 64, lower extremity 62, ear ossicles 6. The hones of the head number 22, consisting of Eight cranial hours-, the frontal, 2 parietal, occipital, 2 temporal, the sphenoid and ethmoid hones; 14 facial bones, 2 lachrymal, 2 nasal, 2 malar. 2 superior maxilla'. 2 palate. 2 turbinal, 1 vomer, and 1 in- ferior maxilla. The hones of the trunk number 52, comprising Twenty-six vertebrae, 7 cervical, 12 dorsal. ."> lumbar, 1 sacrum and 1 coccyx. Twenty-six of the thorax, 7 pairs of true ribs, •"> pairs of false rilis. 2 pairs of floating ribs, the sternum and os hyoideum. The bones of the skull, face, thorax, and vertebral column arc known as the axial skeleton, and consist of 74 segments. The bonea of the npper extremity number 64, comprising on either side : — Two shoulder, clavicle and scapula; 1 arm. humerus; 2 forearm, radius and ulna: 8 wrist (carpus), scaphoid, semilunar, cuneiform, pisi- form, trapezium, trapezoid, os magnum and unciform; 1!) hand, 5 meta- carpal and 14 phalanges. The bones of the lower extremity number 62, comprising on either side : — One hip, divided into 3 portion-, the ilium, ischium and pubes; 1 thigh, the femur; 2 le<_r, tibia and fibula; 1 knee, the patella; 26 foot, 7 tarsal hone-, the astragalus, os calcis, scaphoid, cuboid, external. middle, and internal cuneiform; 5 metatarsal hones, and 14 phalanges. The hones of the shoulder-girdle, npper extremities, pelvic girdle, and lower extremities are known as the appendicular skeleton, and consist of 126 Fegments. They are divided into long, Bhort, llat or tabular and irregular. Long bones, smcli a femur, humerus, or radius, consisl of -hafi or diaphysis, and two extremities or epiphyses. (3) 4 HUMAN ANATOMY. The shaft or diaphysis is a cylindrical tube composed ex- ternally of compact tissue and internally of cancellous tissue, the center being hollowed out by the medullary canal. The epiphyses, or extremities, are expanded for articulation, and are composed of a mass of solid cancellous tissue with a thin coating of compact tissue. The short bones, asi tarsus or carpus, are small, compact, irregular cubes. Flat bones\, as those of skull and shoulder-blade, afford extensive surfaces for protection or muscular attachment. Under irregular or mixed bones are classed the vertebral, sphenoid, maxillary bones, and such that could not be placed under either of the other heads. The surfaces of the bones are marked by certain eminences and depressions, which have received the following names: — An apophysis is a prominent excrescence formed directly upon a bone, and is distinguished from An epiphysis, which has been formed from a distinct center and separated by cartilage, but afterward united to the bone. A head is a rounded, smooth, articular eminence for articulation. A condyle is an irregular prominence for muscular attachment. A trochanter is a large prominence for the attachment of rotator muscles. A tuberosity is a broad, uneven eminence. A tubercle is a small tuberosity. A spine is a sharp-pointed eminence. A line, or ridge, is a rough, narrow elevation, extending some distance. Others, from their fancied resemblance to ordinary objects, have received the following names: — Coronoid, like a crown; coracoid, like a crow's beak; unciform or hamular, hook-like; malleolar, like a mallet; mastoid, nipple-like; zygoma, yoke-like; pterygoid, wing-like; odontoid, tooth-like; spinous, thorn-like; styloid, pen-like; rostrum, a beak; vaginal, ensheathing; squamous, scaly; conoid, cone-like. The cavities of bones are divided into the articular and the non-articular. The articular are named acetabulum, measure-like ; glenoid, hollow; cotyloid, cup-like; facet, smooth; trochlear, pulley- like ; alveolar, socket-like. The non-articular cavities are named notches, fissures, grooves, furrows, fossae, hiatus, foramina, canals, sinuses, aque- ducts, cells, depressions, meatuses, etc. Composition of Bone. — Adult human bones have a specific gravity of 1.92, and are composed of about one-third (33.30) OSTEOLOGY. 5 organic or animal matter, principally gelatin, and two-thirds inorganic or mineral matter, as follows: — Gelatin and blood-vessels, 33.30; phosphate, carbonate, and fluoride of calcium, 64.34; soda, sodium chloride, and mag- nesium phosphate. 2.36 ; total, 100. Structure of Bone. — In structure they consist of an ivory- like compact substance (substantia compacta) inclosing a lamel- lar, recticular cancellous substance; also designated loose or spongy bone (substantia spongiosa) . They are covered with periosteum, and their cavities are lined with endostenm and filled with medulla, or marrow. Periosteum is a fibrovascular membrane, composed of two layers, the outer formed chiefly of connective tissue, the inner composed of several layers of elastic fibers, the deepest of which is the "osteogenic/' or bone-producing layer. The tendons and ligaments are attached to the periosteum by a mutual interlace- ment of the fibers. Endosteum, medullary membrane or internal periosteum, is a delicate, highly vascular membrane lining the cylindrical cavity of long bones. Medulla, or bone-marrow, is of two kinds, the red in the flat and irregular bones, and the yellow in the shafts of adult long bones. The red marrow (medulla ossium rubra) has few blood-vessels, but many corpuscular elements. These may be divided into three groups: (1) red, nucleated, true "marrow- cells," or myelocytes; (2) large, pinkish cells, "myeloplaques" of Robin, supposed to be the source of the red blood-corpuscles or erythroblasts ; and (3) giant cells, or "osteoclasts." The yellow marrow (medulla ossium ftavo) or myelocytes contains numerous blood-vessels, a few "marrow-cells," and a large quantity of fat which imparts to it its color. Blood-vessels of Bone. — The arteries consist of three sete: The arteries of the (1) compact and (2) cancellous tissue are derived chiefly from the periosteum, the latter being larger and Less numerous. The medullary canal of long bones has (3) one or more nutrient arteries, which penetrate tbe compaci tissue obliquely and divide into two branches, one passing upward, the other downward, in tbe canal. All the arteries anastomose freely with each other. The veins consist of three sets also: 1. Tbe veins of the compact substance are small and join the periosteum. 2. The veins of the cancellous 1 issue do not accompany the arteries, but in certain Localities form large, tortuous channels, as in the diploic jreins of ilie cranium. In the long bones they emerge .(3 HUMAN ANATOMY. at the ends, near the articular surfaces. 3. The nutrient arteries are accompanied by one or more large veins, which emerge at the nutrient foramen. Lymphatics and nerves are numerous in the periosteum and are also found in the substance of the bone. Microscopic Appearance. — Microscopically, bone-tissue con- sists of concentric layers or lamella, arranged, about the course of a vascular or Haversian canal /4eooo mcn i11 diameter. In and between these plates of bone-tissue are minute cavities, or lacuna?, each containing a bone-cell, or "osteoblast/' and from which diverge in every direction minute canals, or canaliculi, connecting the lacunas with each other and with the Haversian canals. Each canaliculus contains an artery, vein, and lym- phatic (Schaffer). In this manner every part of the osseous substance communicates. Development. — The development of bone is effected in two ways: (1) from cartilage, the intracartilaginous; (2) from membrane, the intramembranous ; the former occurring at the base of the skull for the protection of vital centers, or m the extremities to secure rigidity of the parts. In the intracartilaginous the parts are first formed in cartilage and then converted into bone. The process is as fol- lows: The cartilage cells at the "center of ossification" become enlarged and arranged in rows. The cartilaginous matrix also increases and separates the cells. Lime salts are deposited be- tween the rows of cells, inclosing them in oblong spaces called primary areola'. - Blood-vessels from the dee}} or osteogenic layer of the peri- osteum, carrying numerous osteoblasts (bone-forming cells) and osteoclasts (bone-absorbing cells) pass into the area of calcified cartilage (center of ossification). The osteoblasts re- place the calcified cartilage, which is absorbed, with new bone, which latter is absorbed in part (tunneled) by the osteoclasts to form the medullary spaces or marrow cavity. ■The walls of the spaces are gradually thickened by suc- cessive layers of osteoblasts, forming lamellae of bone, till noth- ing remains but a channel — the Haversian canal. The perichondrium having been in the mean time con- verted into periosteum, the osteogenic, or vascular, layer of this membrane furnishes a layer of osteoblasts that form enveloping, circumferential layers of bone. Thus, from the osteoblasts all the structures of bone are produced; the remains of a group of cells constitute an Haver- sian canal, the layers deposited by them and the adjacent cells, OSTEOLOGY. 7 the lamella1, and the isulated, persistent cells (osteoblasts) form the contents of the lacunae, or "true bone-cells/' with their nu- trient canaliculi diverging from them. In the intramembranous form the parts are first formed in membrane, and from one or more centers of ossification lime salts are deposited in radiating spiculae, or osteogenic fibers, in- closing the osteoblasts. From these fibers the process spreads, and vessels from the neighboring parts pass into it and form Haversian canals. Fig. L Frontal bone, outer surface. The formation of the lamella! and the lacunae is essentially the same as in the intracartilaginous, and in most bones both processes go on simultaneously. Ossification appears first in the clavicle and inferior maxilla (from fifth to seventh week) and last in the pisiform bone (twelfth year). The epiphyses ossify from birth and unite from the age of puberty on to maturity, in reverse order to the ap- pearance of ossification, being regulated, apparently, by the direction of the nutrient artery in the upper extremity toward the elbow and in the lower extremity from the knee. The only exception to this rule ie the lower end of the fibula, which ossifies and also becomee united before the upper (vide Table of u7ar fuasa Vagi naT jjroceti Styloid prueeet jStylo-'masroi-a7 foramen Jxiavlnr Si'rfaee Auricular future FIG. 9. Left temporal bone, inferior surface of the petrous portion. the digastric muscle, and still more internally the occipital groove (sulcus a. occipitalis) for the accommodation of the occipital artery. The internal surface is concave, and presents a groove, the fossa ngmoidea (sulcus si'jmoideus) for the lateral sinus, in the bottom of which is the opening of the mastoid foramen (foramen mastoideum ) . The petrous portion (pars petrosa pyramis) is pyramidal in shape, intensely hard, and contains the internal ear. It pro- 2 18 HUMAN ANATOMY. jects downward, forward and inward, and presents for exami- nation a superior, posterior and inferior surface. The superior or anterior surface (fades anterior pyramidis) is united to the squamous portion by the temporal suture and presents the following- : — An eminence (eminentia arcuata), marking the position of the superior semicircular canal; A depression, forming the roof of the tympanum; The hiatus Fallopii (hiatus canalis facialis), transmitting a branch of the middle meningeal artery and the petrosal branch of the A^idian nerve to the aqueduct of Fallopius; Foramen (apertura superior canaliculi tympanici) for the smaller petrosal nerve; Orifice of the carotid canal, transmitting the carotid plexus of the sympathetic and the internal carotid artery; and Depression (impressio trigemini) for Gasserian ganglion of the trifacial nerve. The posterior surface (fades posterior pyramidis) presents about its middle The meatus auditorius internus (meatus acusticus inter- ims), at the bottom of which is the lamina cribrosa, a perpendic- ular plate of bone, and numerous foramina for the exit of the auditory nerve and the auditory branch of the basilar artery, one of which is the commencement of the aquaeductus Fallopii (canalis facialis) for the facial nerve. Posteriorly to the internal meatus is the aquaductus ves- tibidi, lodging a process of dura mater, and transmitting an artery and vein, and between them another small depression for" the same purpose. The inferior or basilar surface (fades inferior pyramidis) presents, from within outward, the following : — A rough, quadrilateral surface, for attachment of the tensor tympani and levator palati muscles; The opening of the carotid canal (foramen caroticum ex- ternum), transmitting the carotid plexus of the sympathetic and the internal carotid artery; Aqiuzductus cochlece (apertura externa canaliculi cochlea), transmitting a vein to the internal jugular vein from the cochlea ; Jugular fossa (fossa jugularis), forming the jugular fora- men (foramen lacerum posterius) by articulation with the occipital bone, and transmitting the internal jugular vein, the glossopharyngeal, pneumogastric, and spinal accessory nerves. OSTEOID >GY. 19 and meningeal branches "J' the ascending pharyngeal and occip- ital aiter - . Foramen for Jacobsons nerve, the tympanic branch of the _ ssopharyngeal (ft. tympanicus) ; Foramen for entrance of Arnold's nerve, the auricular branch of the pneumogastric ; Jugular surface for articulation with jugular process on oc- cipital bone ; The vaginal process (vagina processus styloidea), extend- ing from the mastoid process to the carotid canal, and ensheath- ing Fig. 10. Section of the temporal bone (natural size) through the middle <-ar. Fallopian canal, mastoid antrum, and cells, showing dense bone between the antrum and cells, with no communication between them: 1, drumhead: L'. tip of the mallet handle; 3, anvil, showing the long crus at the right for articulation with the stirrup, and the short proc- t the left which serves the purpose of an anchor to the bone; 4, head of the mallet; '■>. tensor tympani muscle and tendon; 6, dense bone where pneumatic spaces are usually found; 7, pneumatic cells in the tip of the mastoid process: 8, Fallopian canal, for the facial nerve; 9, the Btirrup. At the right of 9 and at the left of the anvil is the aditus ad antrum, connecting the tympanum with the antrum. The styloid process (processus styloideus), affording at- lacluiiciit for the stylomaxillary and stylohyoid ligaments and bylohyoideus, stylopharyngeus and styloglossus muscles; The stylomastoid foramen (foramen stylomasioideum) , for exit of facial nerve; The auricular fissure (fissura tympanomastoidea) , for the exil of Arnold's aerve, the auricular branch of the pneumo- ric. The base of the petrous port ion presents The- external auditory meatus (meatus acusticus externus), • anal leading into the t) mpanum, and 20 HUMAN ANATOMY. The auditory process, a lamina of bone forming the greater part of its walls. Borders. — The superior border (angulus superior pyram- idis) affords attachment to the tentorium cerebelli, and has a groove for the superior petrosal sinus. The posterior border {angulus posterior pyramidis) forms part of the jugular fossa (fossa jugularis). The anterior border (angulus anterior py- ramidis) articulates with the spinous process of the sphenoid. Between the squamous and petrous portions may be seen the opening of the canalis musculotubarius; this canal is found either completely or partially divided by an osseous lamina, separated by the processus cochleariformis (septum canalis mus- cuiotuharii), for the tensor tympani muscle above and the Eustachian tube below. Muscular attachments are fifteen: — To the squamous portion, 2 — the temporal and masseter; to the mastoid portion, 6 — -sternomastoid, occipitofrontal!?, splenitis capitis, trachelornastoid, retrahens aurem, and digas- tricus; to the styloid portion, 3— stylohyoideus, stylopharyn- geus and styloglossus; and to the petrous portion, 4 — levator palati, tensor palati, tensor tympani and stapedius. Its ossific centers are ten (10), 1 for the squamous portion, including the zygoma, 1 for auditory process, 6 for' petrous and mastoid portions, and 2 for the styloid process. The sphenoid (os sphenoidale) is wedged in between the bones at the base of the skull anteriorly and resembles a bat with extended wings. It consists of A body, 2 Lesser wings, and 2 Greater wings, 2 Pterygoid processes. It presents 12 Foramina, 12 Pairs of muscles, and 12 Articulations, 14 Centers of ossification, and is in every respect the most important bone of the cranium, entering into the formation of 5 Cavities — (2) Zygomatic, (1) Anterior cerebral, (3) Sphenomaxillary, (2) Middle cerebral, (4) Nasal, and (3) Orbital, 3 Fissures — .(4) Pharyngeal, (1) Sphenomaxillary, (5) Nasal, (2)' Pterygomaxillary, and 4 Fossce — (3) Sphenoidal or foramen ( 1 ) Temporal, lacerum anterior. OSTEOLOCY. 21 The body (corpus) is a hollow cube and presents upper, lower, anterior and posterior surfaces. The upper surface (fades cerebralis) of the body presents from before backward the following points: — Ethmoidal spine (s/rina ethmoidalis) , for articulation with the cribriform plate of the ethmoid ; Optic groove (sulcus chiasmal is), lodging the optic com- missure and terminating in the optic foramen; Olivary process (iuuerculum sella'), a small, olive-like eminence behind the optic groove; Pituitary fossa or sella turcica (fossa hypohyseos), for the pituitary body; tliidlt Ck^ld prxcj P^/CTzrr CUiwutprocCM \ •M™^ •T/'W ~"-^_^^_^ ** \ , \ Groove jar Fig. 11. Sphenoid bone, upper surface. Middle clinoid processes (processus dinoidei medii), one on either side bounding the pituitary fossa; Dorsum sella, or dorsum ephippii, a quadrilateral plate of bone, bounding the pituitary fossa behind, and notched on either side for the passage of the sixth cranial nerves; Posterior clinoid processes (processus clinoideus posterior) are the terminations of the dorsum sella1 on either side and afford attachment to the tentorium cerebelli; Declivity, or clivus Blumenbachii (clivus), supporting the upper part of the pons Varolii; Carotid or cavernous groove (sulcus caroticus), lodging the cavernous -inns and the internal carotid artery; Lingula (lingtUa sphenoidalis) , a ridge of bone to the outer side of the cavernous sinus. 22 HUMAN ANATOMY. The lower surface presents : — Rostrum {rostrum sphenoidalis) , the continuation down- ward of the ethmoidal crest, for articulation with the ala? of the vomer; Vaginal processes, a thin plate of hone on either side artic- ulating with the edges of the vomer; and Groove (sulcus pterygopalatinus), converted into the pterygopalatine canal by articulation with the sphenoidal process of the palate bone for transmission of the pharyngeal nerve and pterygopalatine vessels. Jtery>joid R LAXATOR TYMPANI Internal Pterygoid, yilate HumuJxi.r Uroctst Fig. 12. Sphenoid bone, anterior surface. The anterior surface presents : — Ethmoidal crest or sphenoidal crest (crista sphenoidalis), a thin plate of bone articulating with the ethmoid; Openings of sphenoidal cells or sinuses (sinus sphenoidal 'es) partially closed by the sphenoidal turbinated bones or pyramids of Wistar (conchce sphenoid ales), two curved plates of bone ab- sent in infancy, but derived as a pair of cones from the ethmoid; The sphenoidal cells open into the superior meatus of the nose, but occasionally into the posterior ethmoidal sinuses. The posterior surface articulates with the basilar process of the occipital and becomes united to it between the eighteenth and twenty-fifth year. OSTEOLOGY. 23 GEEATEK OR TEMPORAL WINGS OF THE SPHENOID (AL.K MAiiX.E). The superior surfaces (facie* cerebralis) of the great wing, deeply concave, forms part of middle fossa of skull, and pre- sents the following : — Foramen rot and urn, for second division of fifth nerve; fora- men ocale, for third division of fifth, small meningeal artery, and small petrosal nerve; foramen Vesalii, for small vein, and foramen spinosum, for passage of middle meningeal artery. The exterior surface is convex and presents: — Pterygoid ridge or infratemporal crest (crista infratem- poral^), dividing the hone into two portions; Spinous process, for attachment of tensor palati muscle, and the internal lateral ligament of the lower jaw. The anterior or orbital surface (fades orbitalis) enters into the formation of the orbit and presents numerous external orbital foramina for passage of branches of the deep temporal arteries. The circumference of the greater wing forms part of the sphenoidal fissure and articulates with the frontal bone in front and laterally with the parietal, squamous and petrous portions of the temporal bone. The lesser or orbital wings (alee parvce), or processes of Ingrassias, project on either side from the superior surface of the body, to which they are attached by two roots. The upper surface forms part of the anterior fossa of the skull. The lower surface forms part of the orbit and the upper border of the sphenoidal fissure, or foramen lacerum anterius {fissiira orbitalis superior), transmitting the third", fourth and ophthalmic divisions of the fifth and the sixth cranial nerve, orbital branch of the middle meningeal artery, a recurrent branch of the lachrymal artery, the ophthalmic vein, and fila- ment- from the cavernous plexus. The anterior clinoid process (processus clinoideus anterior) i- formed by the inner extremity of the posterior border. Between the two roots is the optic foramen (foramen opticum), transmitting the optic nerve and ophthalmic artery. The pterygoid processes (processus pterygoidei) are two wing-like processes descending from the junction of the greater wings with the body. Each process presents the following: — An externa] pterygoid plate (luminu lateralis processus pterygoidei)} for attachment oi the external pterygoid muscle; g4 HUMAN ANATOMY. The pterygoid fossa (fossa pterygoidei) lies between the two plates and affords attachment for the internal pterygoid muscle. The internal pterygoid plate (lamina medialis processus pterygoidei), terminating below in a hook-like process — the hamular process (hamulus pterygoideus) — for the tendon of the tensor palati nrascle. The scaphoid fossa (fossa scaphoid ea) , at the base of the internal plate, affording origin for the tensor palati muscle and presenting just above it, the posterior opening of the Vidian canal (canalis pterygoideus [Vidii]) for the Vidian vessels and nerves. An angular interval between the two plates articulates with tuberosity or pterygoid process of the palate bone. The foramina are: Opticum, lacerum anterius, rotundum, Vesalii, ovale, spinosum, and two canals — Vidian and pterygo- palatine; articulates with twelve bones — two malar, two palate, vomer, and all of cranium. Its muscular attachments are eleven pairs — external and internal pterygoid, temporal, tensor palati, superior constrictor, levator palpebral, obliquus superior; superior, inferior, internal and external recti. Its ossific cen- ters are fourteen, as follows: Body, 2 anterior and 2 posterior; external pterygoid plates and greater wings, 2 ; internal ptery- goid plates, 2; lingula, 2; lesser Avings, 2; sphenoturbinals, 2. The ethmoid (os etlimoidale) is a spongy, cubical bone, consisting of a perpendicular lamella, a horizontal or cribri- form, plate, and two lateral masses. The horizontal or cribriform plate (lamina cribrosa) pre- sents from before backward the following points: — Two projections (processus alares) completing the foramen coBCum, the commencement of the longitudinal sinus; Crista galli, affording attachment to the falx cerebri; Foramina cribrosa, three rows, transmitting filaments of the olfactory body; Fissure for passage of the nasal branch of the ophthalmic nerve. The perpendicular plate (lamina perpendicularis, or mes- ethmoid) assists in forming the nasal septum, articulating in front with the crest of the nasal bones and the frontal bone and behind with the sphenoid and vomer. It is usually inclined to one side. The lateral masses — labyrinth (labyrinthus etlimoidalis) — are composed of the ethmoidal cells. OSTEOLOGY. The upper surface presents two notches, converted into fo- ramina by articulation with the frontal bone — The anterior ethmoidal foramen (caimUs ethmoidalis an- teriiis), transmitting the anterior ethmoidal artery and nasal nerve, and The posterior ethmoidal foramen (canalis ethmoidalis pos- terius), transmitting the posterior ethmoidal artery and vein. The outer surface presents The os phi mi tit (lamina papi/racea), a smooth plate of bone forming the inner wall of the orbit, and projecting downward from it is toitk mf.'turbinateJ 6. Fig. 13. Ethmoid bone. The unciform process (processus uncinatus), a hook-like plate of bone closing the upper part of the opening of the antrum. The inner surface forms part of the nasal fossa, and presents The superior turbinated bone (concha nasalis superior), forming the upper boundary of the superior meatus (meatus nasi superior), into which open the sphenoidal and posterior ethmoidal cell-, and the .Middle turbinated bone or the inferior ethmoidal turbinate bone (concha nasalis media), forming the upper boundary of the middle meatus of the nose, into which open the anterior ethmoidal cells, the frontal cells through the infundibulum (infundihuluni ethmoidale) and the antrum Eighmorianum | imi.s in n. i iHuris). 26 HUMAN ANATOMY. It articulates with fifteen bones — -two sphenoidal turbinated, sphenoid, frontal, and eleven of the face, the two nasal, two lachrymal, two superior maxilla?, two palate, two inferior turbi- nated and the vomer. It has no muscular attachments. Its ossific centers are three — one for the lamella and one for each lateral mass. BONES OF THE EACE (OSSA FACIEI). The bones of the face are fourteen in number — six pairs and two single bones: Two lachrymal, two nasal, two malar, two viihFrontdl Fig. 14. Lachrymal bone. superior maxillary, two palate, two turbinal, one vomer and one inferior maxillary. The lachrymal (os lacrimale) are two small, thin bones forming part of the inner wall of the orbit and the lachrymo- nasal duct, and named, from their resemblance to a finger-nail, the ossa" unguis. Each presents externally : — The lachrymal crest (crista lacrimdlis posterior) affords attachment for the tensor tarsi and separates the anterior sur- face from the groove; The groove forms part of the lachrymal groove and lodges the lachrymal sac; The hamular process (hamulus lacrimalis), or, when sep- arate, called the lesser lachrymal bone, projects downward and articulates with the lachrymal tubercle of the superior maxilla. OSTEOLOGY v; Internally : — A furrow, corresponding to the lachrymal crest on the ex- ternal surface ; The internal surface enters into the formation of the middle meatus, closing in the anterior ethmoidal cells. It articulates with four bones — the frontal, ethmoid, supe- rior maxilla and inferior turbinated. The only muscular attachment is the tensor tarsi. It is developed from one ossific center. The nasal hones (ossa nasalia) are two irregularly quad- rate bones forming together the bridge of the misc. The outer surface, covered by the compressor nasi and pyr- amidalis nasi muscles, presents A foramen (foramen nasale), transmitting a small vein. The inner surface presents u-iiA Frontal B. —Ojjpoiit* bone. (Outer surface.) Fig. 15. Nasal bones. with Tro ntaZ Szn7te. orest aZ JiroeesM. """fty Fig. 19. Palate bone (viewed posteriorly). HORIZONTAL f IAT! Fig. 20. Palate bone (inner surface). Each bone assists in forming three cavities — the outer wall and floor of the nose, the roof of the mouth and floor of the orbit; two fossae — the pterygoid (fossa pterygoid ea) and spheno- maxillary (fossa pterygopalatina) , and one fissure — the spheno- maxillary (fissura oroitalis inferior). The horizontal plate (pars liorizontalis) is quadrilateral. Its inferior surface (fades palatina) presents A ridge for the aponeurosis of the tensor palati muscle ; A groove (sulcus pterygopalatinus) entering into the for- mation of the posterior palatine canal; Accessory palatine foramina (canal es palatini) for poste- rior descending palatine nerves. Its posterior border (spina nasalis posterior) presents The posterior nasal spine (crista nasalis) for the azygos uvulae muscles. OSTEOLOGY. 33 The infernal border articulates with its fellow and presents A crest for the vomer. The vertical or perpendicular plate (.pars perpendicularis) projects upward and inward, terminating above in two well- marked processes — the orbital and sphenoidal processes. The internal or nasal surface (fades nasalis) forms part of the outer wall of the nasal fossa, and presents An inferior turbinated crest (crista conchalis) for the in- ferior turbinated bone; A superior turhinated crest (crista ethmoidalis) for the middle turbinated bone, separating the middle from the in- ferior meatus of the nose. The external or maxillary surface (fades maxillaris) forms part of the sphenomaxillary i'ossa and covers part of the orifice of the antrum. It presents at the posterior part • A groove, converted by the superior maxilla, into a canal, the posterior palatine, for the passage of the descending palatine vessels and the descending palatine branches from Meckel's gan- glion. The anterior border presents The maxillary process (processus maxillaris), closing part of the orifice of the antrum. The pterygoid process (processus pyramidalis), or tuberos- ity of the palate, at its lower part wedged in between the ptery- goid plates of the sphenoid, and giving origin to part of the superior constrictor and the internal pterygoid muscle. Foramina of accessory descending palatine canals, for pos- terior descending branches from Meckel's ganglion. The obbital process (processus orbitalis), a large, hollow, triangular process, is so called from its forming part of the orbit. It presents The anterior or maxillary articular surface for superior maxilla ; The- posterior or sphenoidal articular surface for sphenoid; The internal or ethmoidal articular surface, for ethmoid; A superior or orbital surface, a smooth portion entering into the format ion of the orbit ; An external or zygomatic surf ace], also smooth, forming part of the zygomatic fossa. Tin. sphenoidal pbocess (processus sphenoidalis) is a -mall, thin plate of hone which articulates by its upper surface with the -phenoidal turbinated hone; its inner surface forms :', 34 HUMAN ANATOMY. part of the nasal fossa, and its outer surface articulates with the pterygoid process of the sphenoid and forms part of the spheno- maxillary fossa. The anterior border forms a part of the sphenopalatine fora- men, which passes between the orbital process and the sphenoidal process. It articulates with six (6) bones. — ethmoid, sphenoid, supe- rior maxillary, vomer, inferior turbinated, and fellow of the opposite side. Its muscular attachments are azygos uvula?, tensor palati, internal and external pterygoid and superior constrictor. It has one ossific center at the junction of the plates. The inferior turbinated bones (concha nasalis inferior) are small, scroll-like bones, situated on the outer wall of the (Inner surface.) Fig. 21. Inferior turbinated bones. (Outer surface.) nasal fossa, separating the middle from the inferior meatus. Each presents two surfaces and three processes — the lachrymal, ethmoidal and maxillary. The internal surface is lined in the recent state with the mucous membrane of the nose. The external surface is attached to the superior maxilla and palate bone, and presents above the three processes. The lachrymal process (processus lacrimalis) forms part of the lachrymal canal, articulating with the superior maxilla and the lachrymal bone. The ethmoidal process (processus ethmoidal is) articulates with the hook-like process of the ethmoid bone. The maxillary process (processus maxillaris) projects out- ward and downward from the ethmoidal process, and narrows the orifice of the antrum. It articulates with four bones — ethmoid, palate, superior maxillary and lachrymal. Muscular attachments, none. It has one ossific center. OSTEOLOGY. 35 The vomer is a thin, triangular plate of bone, contributing to form part of the septum nasi. It is usually bent to one side, The superior border presents a groove with overlapping lamina?, or vaginal processes (alw vomeris) for articulation with the rostrum and vaginal processes of the sphenoi I. The inferior border articulates with the crests of the supe- rior maxilla? and palate-bones. The posterior border is smooth and free, forming the pos- terior nasal septum. The hi I end surfaces present Furrows foT blood-vessels, and Nasopalatine grooves for the nasopalatine nerves, which descend to the foramina of Scarpa. az* It articulates with six hones — the ethmoid, sphenoid, two palate and two superior maxilla?. It has no muscular attachments. It is developed from a single ossific center, although it begins as two cartilaginous laminae, which coalesce. The infebiob maxilla, or mandible (mandibula), is the largesl bone in the face, it consists of a body and two rami. The body (corpus mandibular) is the horizontal, horseshoe- like portion, containing the lower teeth. The external surface presents from before backward: — Symphysis, a vertical median line indicating the junction of the vaginal halves ; .Mental process ( prot uberan I in mentalis), a triangular prominence cons! ituting the chin ; Incisive fossa, for the origin of the Levator labii inferioris; Mental foramen (foramen mentale), for the exit of the menial nerve and artery ; 36 HUMAN ANATOMY. External oblique line (linea obliqua), running backward and upward from the mental process, for the attachment of the depressor anguli oris and depressor labii inferioris. The internal surface presents, from before backward : — Genial tubercles (spina? mentales), four in number, for at- tachment of the geniohyoglossi muscles above and the genio- hyoidei below; Sublingual fossa (fovea sublingualis), for the sublingual gland ; Internal oblique line or mylohyoid ridge (linea mylo- hyoidea), for the attachment of the superior constrictor, the mylohyoid muscles, and the pterygomaxillary ligament; i^v-4 Fig. 23. Inferior maxillary bone. Submaxillary fossa (fovea submaxillar is) , for the submax- illary gland. The superior or alveolar border (pars alveolaris) presents sixteen cavities (ten in childhood) for the teeth. The inferior border, or basilar portion (basis mandibular) is grooved posteriorly for the facial artery. The ramus, or perpendicular portion (rami mandibular), on either side is somewhat quadrilateral, and presents the fol- lowing points : — The internal surface of ramus; Foramen (foramen mandibulare) of inferior dental canal (canalis mandibular), for inferior dental vessels and nerve; Mylohyoidean groove (sulcus mylohyoideus) , described above ; OSTEOLOGY. 37 Spine, for attachment of the internal lateral ligament of the lower jaw. The upper border is surmounted by two processes, the coro- noid and the condyloid processes, separated by the sigmoid notch. The coronoid process (processus coronoideus) is in front, and affords attachment for the temporal muscle. The condyloid process (processus condyloid eus) articulates with the glenoid fossa of the temporal bone, being supported upon a constricted portion, the neck (collum- mandibular) , which receives the insertion of the external pterygoid muscle, and MIO'UVO-CLOSSUS CtNIO-HVOIDIU* jfyjcly'J R'fy B a dy Fig. 24. Left half of inferior maxillary bone, inner surface. presents to its outer side a tubercle for the external lateral ligament of the lower jaw. The sigmoid notch (incisura mandibula>) is crossed by the masseteric vessels and nerve. The lower border presents : — Angle of the jaw (angulus mandibulce) , the point of junc- tion of the ramus with the body ; Rough surface,, for attachment of the stylomaxillary liga- ment, and on its inner side the internal pterygoid muscle, and on its outer side the masseter. It articulates with the two temporal bones. The muscular attachments are fifteen pairs; to the external surface, sis — depressor anguli oris, depressor labii inferioris, levator labii inferioris, orbicularis oris, platysma myoides and buccinator; from the interna] Burface of the body, five — the 38 HUMAN ANATOMY. geniohyoglossus, geniohyoid, mylohyoid, digastric and' superior constrictor; and to the ramus, four — the masseter, internal and external pterygoids and temporal. It has two ossific centers, one for each lateral half, devel- oped partly from membrane, partly from cartilage, being pre- ceded in time only by one bone — the clavicle. Changes in Lower Jaw. — Certain changes occur in the size and shape of the lower jaw as age advances. At birth the body is shell-like, containing sockets for the ten temporary teeth, the mental foramen is large and opens beneath first molar, and the coronoid process is large and at right angles to base. Thyrohyoid* Slijlo-hyoid. ^Omo-hyoid, Mylo-hyoidi Sterno-hyoid: Fig. 25. The hyoid bone. In adults the base and alveolar process are equal in size, the mental foramen opens midway between upper and lower borders, and the rami are at nearly right angles to base. In advanced age the loss of the teeth and alveolar borders greatly reduces the body, the mental foramen opens near the alveolar border, and the rami are united at very obtuse angles with the base. The hyoid bone, or lingual bone (os liyoicleum) is a small, U-shaped bone' situated at the base of the tongue, consisting of a body ( corpus ossei hyoidei) ; two greater and two lesser cornua. It is supported by the stylohyoid ligament from the styloid processes of the temporal bones. The anterior surface is divided by a crucial ridge into four depressions for muscular attachments, and its center presents a tubercle. The posterior surf ace is in relation with the epiglottis, being separated by the thyrohyoid membrane. OSTEOLOGY. 39 The upper and hirer borders afford attachment to muscles, and the lateral surfaces are mounted with cartilage for articula- tion with the greater comua. The greater comua, or thyrohyals (comua majora), pro- ject backward, afford attachment to the hyoglossus, thyrohyoid, and middle constrictor of the pharynx, and terminate behind m a tubercle for the attachment of the thyrohyoid ligament. The lesser comua, or ceratohyals (comua minora), are two tuber-like projections attached at the junction of the body with the greater comua, and receiving the insertion of the stylohyoid ligaments. They do not articulate with any bone. Its muscular attachments are ten — the Hngualis, hyoglossus, geniohyoglossus, middle constrictor, stylo-, mylo-, genio-, sterno- thyro-, omo- hvoid; also aponeurosis of digastric and stylo- hyoid ligament. Os,sific centers, five — one for body and one for each horn. The Sutures and Fontanelles. — The sutures are divided into three sets, those of the vertex, side, and base of the skull. At the vertex of the skull there are three : — Sagittal, or interparietal (sutura sagittalis) ; Coronal, or frontoparietal (sutura coronalis) ; Lambdoid, or occipitoparietal (sutura lambdoid ea) . At the side of the skull are three : — Sphenoparietal (sutwa sphenoparietalis) ; Squamoparietal, or squamous {sutura squamosa); Mastoparietal (sutura parietomastoidea) . At the base of the skull are nine:— Basilar, in center (fissura spheno-occipitalis) , Petro-occipital ( fissura petro-occipitalis) , ~\ Maato-oecipital | sutura occipitomastoidea) , Lon either side. Petrosphenoidal (fissura sphenopetrosa) , Squamosphenoidal (sutura sphenosquamosa) , J The sutures of the face are very numerous, but the most important are : — Zygomatic, at tin- temporomalar junction; Transverse, posses from one external angular process to the other, and connects the frontal with the malar, ethmoid, sphenoid, lachrymal, superior maxillary and nasal bones; intermaxillary, the median suture between the superior maxillae; Symphysis, Hie remains of a fetal suture. The fontanelles (fontieuli) are six membri us intervals in the infant's skull corresponding to the four angles of the parietal bones. The anterior or bregmatic (fontwulus frontalis), larger 40 HUMAN ANATOMY. and lozenge-shaped, is at the junction of sagittal and coronal sutures; the posterior (fonticulis occipitalis) , smaller and tri- angular, is at the junction of sagittal and lambdoid sutures; and the four lateral are at anterior and posterior inferior angles of parietal bones (fonticulus sphenoidalis et fonticulus mastoideus) . The Wormian, supernumerary, sutural or epactal bones (ossa triquetra) are irregular bones interposed in intervals be- tween the cranial bones, chiefly in the course of the lambdoid suture. Congenital fissures from an arrest of ossification also occur, the most common being the parietal and sagittal fissures (Treves). FOSSAE OF SKULL. The fossa? at the lateral region of the skull are temporal, zygomatic, or infratemporal, and sphenomaxillary or pterygo- palatine fossa. The temporal fossa (fossa temporalis) is deeply concave in front, convex behind ; it is formed by parts of five bones — frontal, malar, sphenoid, temporal and parietal. It is bounded above and behind by temporal ridge; in front by the sphenoid, malar and frontal; below and externally by zygoma, and pterygoid ridge on greater wing of sphenoid. It is crossed by six sutures — sphenomalar, sphenoparietal, squamoparietal, squamosphenoidal, coronal and transverse facial. It opens below into the zygo- matic fossa, is filled by the temporal muscle, and lodges the deep temporal vessels. The zygomatic or infratemporal fossa (fossa infratem- poralis) is an irregular cavity containing parts of temporal, ex- ternal and internal pterygoid muscles, and is bounded above by pterygoid ridge of sphenoid and squamous portion of tem- poral, below by alveolar border of superior maxilla, in front by tuberosity of superior maxillary, behind by border of ptery- goid process, externally by zygoma and ramus of lower jaw and internally by external pterygoid plate. It is traversed by two fissures — sphenomaxillary and pterygomaxillary. The sphenomaxillary fissure (fissura orbitalis inferior) connects the zygomatic, temporal and sphenomaxillary fossa?, and transmits the infraorbital artery, ascending branches of Meckel's ganglion, the superior maxillary nerve and its orbital branch. It is bounded above by great wing of sphenoid, below by superior maxilla and part of palate bone; internally it opens into pterygomaxillary fissure and externally part of malar bone. OSTEOLOGY. 41 The pterygomaxillary fissure connects the zygomatic and sphenomaxillary fossa1, passing vertically between pterygoid process of sphenoid and superior maxillary bone, and transmits branches of internal maxillary artery. The sphenomaxillary or pterygopalatine fossa (fossa ptery- gopalatina) is a triangular cavity, bounded above by body of Fig. 26. 1. median suture; 2, anterior palatine foramen; 3, posterior pala- tine foranif-n: 4, posterior nasal spine; ;j, vomer; 6, hamular process; 7, pterygopalatine canal; 8, external plate; 9, scaphoid fossa; 10, foramen ovale; 11, foramen spinosum; 12, foramen lacerum medium; 13, zygoma; 14, basilar suture; 15, gienoid fossa; 16, external audi- tory meatus; 17, mastoid process; 18, styloid process; 1!», carotid foramen; 20, foramen lacerum posterius; 21, occipital condyles; 2'1. i- process; 23, foramen magnum; 24, occipital crest; 25, Inferior curved lines. sphenoid and orbital plate of palate-bone, within by vertical plate of palate, in front by superior maxillary, and behind by pterygoid process of Bphenoid. Jt lias communicating with it 42 HUMAN ANATOMY. Three f ossse — orbital, nasal and zygomatic ; Three fissures — sphenoidal, sphenomaxillary and pterygo- maxillary ; Two cavities — cranial and buccal; and TENDS OCULI Ant. Wdsal Spine IneLrive fossa,. Grew* far Taci'ai Cft Fig. 27. Facial portion of skull. . Five foramina — foramen rotundum, Vidian and pterygo- palatine posteriorly; sphenopalatine, on inner wall, and pos- terior palatine canal, and (sometimes) accessory posterior pala- tine canals below. It contains internal maxillary artery, superior maxillary nerve and Meckel's ganglion. The orbits are two pyramidal cavities, situated between the nose and external angular processes, the forehead and face, their bases outward and forward, their apices converging toward OSTEOLOGY. 43 the body of the sphenoid bone. They contain the eye and its appendages, and are each formed by seven bones — frontal, eth- moid, sphenoid (entering into both cavities), superior maxil- lary, lachrymal, malar and palate (separate in each). The loof (paries superior), concave, presents: — Externally, depression for lachrymal gland ; Internally, depression for pulley of superior oblique; Posteriorly, a suture between lesser wing and frontal bone. It is formed by the orbital plate of frontal and lesser wing of sphenoid. The floor (paries inferior) is short and flat, and presents: — Internally, a depression for inferior oblique muscle; Middle, the infraorbital groove for nerve and artery; Externally, suture of superior maxillary and malar ; Posteriorly, suture between palate and superior maxillary. It is formed by orbital processes of malar and superior maxilla and orbital surface of palate. Inner wall (paries mediaiis) presents crest of lachrymal bone, lachrymal groove, sutures between ethmoid and sphenoid and ethmoid and lachr}'mal. It is formed by Lachrymal, parts of ethmoid, and sphenoid and nasal process of superior maxilla. The outer wall (paries lateralis) presents suture between malar and sphenoid and orifices of malar canals. It is formed by orbital plate of sphenoid and orbital process of malar. The angles of the orbit are four — superior internal anil external, inferior internal and external. They present the following:- — Superior internal angle: Foramen ethmoidal anterior, transmitting anterior eth- moidal vessels and nasal nerve; Foramen ethmoidal posterior, transmitting posterior eth- moidal artery ami vein; Suture between frontal and ethmoid, and frontal and lach- rymal ; Superior external angle: Sphenoidal fissure, or foramen lacerum anterius, transmit- ting third, fourth, ophthalmic division of fifth and sixth nerves; orbital branch of middle meningeal artery, filaments of cavern- ous plexus of sympathetic, recurrenl branch id' lachrymal artery, and ophthalmic vein and process of dura mater. Su/urr between greal wing of sphenoid and malar bone, Inferior internal angle: 44 HUMAN ANATOMY. Suture between superior maxillary and palate bones, uniting with lachrymal and os planum of ethmoid; Inferior external angle: Sphenomaxillary fissure, for passage of superior maxillary nerve and its orbital branch, infraorbital vessels, and ascending branches from Meckel's ganglion. The circumference is bounded : — Inner side by nasal process of superior maxilla and internal angular process of frontal; Outer side by external angular process of frontal and malar bone; Above {mar go supraorbitalis) by supraorbital arch; Below {margo infraorbitalis) by lachrymal, superior max- illa and malar. It presents sutures malomaxillary below; Sutures frontomalar . to outer side ; Sutures frontomaxillary to inner side; Supraorbital notch or foramen above, for passage of supra- orbital nerve, artery and vein, and forms part of lachrymal groove. The apex presents : — • Optic foramen, for passage of optic nerve and ophthalmic artery. Each orbit presents nine (9) openings, as follows: — - 1. Supraorbital foramen for passage of supraorbital artery, vein and nerve; 2. Infraorbital canal for infraorbital nerve and artery; '3. Anterior ethmoidal foramen for anterior ethmoidal ar- tery, vein, and the nasal nerve; 4. Posterior ethmoidal foramen for posterior ethmoidal ar- tery and vein; 5. M'alar foramina, for malar and temporal branches of temporomalar nerve ; 6. Lachrymal canal, for lachrymonasal duct; 7. Sphenomaxillary fissure, for infraorbital vessels, supe- rior maxillary nerve and its orbital branches, and ascending branches from sphenopalatine (Meckel's) ganglion; 8. Foramen lacerum anterius (or sphenoidal fissure), for third, fourth, ophthalmic division of fifth and sixth nerves, oph- thalmic vein, branches of lachrymal and meningeal arteries, fila- ments of the sympathetic nerve and a process of dura mater for ensheathing of the nerve ; 9. Optic foramen, for optic nerve and ophthalmic artery. OSTEOLOGY. 45 The nasal fossae (cavum nasi) consist of two large cavi- ties separated from each other by the septum nasi (septum nasi osseum), and opening in front by the anterior nares (apertura piriformis), and behind into the pharynx by the posterior nares (choanal). They are formed by fourteen bones — the frontal, ethmoid, sphenoid and all the bones of the face except the lower jaw and malar. Aoof Ratal Jfatal .J/iiie of Frontal But* BcramulFUu •f£tiwtad SjJmmJ Prolt paticJ Amiga HatO'ltwhrymal Co-Mat MnmtU pmtttJi&tmjk /itfunJtla/um ■ Ouur Wall ■fawn.!, Pre.: trfSuptfa J.nrKrynuiL L',uifIt61tQLm Far- Lirrrurti medium Crifiot of Carried Canal vi:i;ti;i:i;.i-; (vertebra thoracales) have large, lie;irt-.dia|icd bodies, Hat above and below. Laterally have demi- articular facets (fovea costalis superior et inferior) for heads of ribs; long, strong, transverse processes, marked by Facel for tubercle of rib (fovea costalis vertebralis) ; vertical articular processes, superior backward, inferior forward; long, triangular, overlapping spinous proems; small, and nearly circular, spinal canal. 60 HUMAN ANATOMY. The peculiar dorsal vertebra? are : first, which resembles cervical, and has distinct facet for head of first rib, and half- one for half of second head; tenth, eleventh and twelfth; one facet for head of rib; and eleventh and twelfth, no facet on transverse processes for tubercle of rib. The lumbar vertebra (vertebra? lumbales) are the largest, and have transverse oval body, flat above and below; long, thin, transverse processes, representing ribs; vertical proc- esses interlocking with adjacent vertebrae; large, triangular spinal canal. The last lumbar is peculiar in having body thicker in front, shorter and stronger transverse process, and inferior articular processes wider apart, and directed forward. Super. Artie. Proc. Fig. 40. A lumbar vertebra. Muscular attachments to atlas, ten — rectus anticus minor, rectus lateralis, rectus posticus minor, obliquus superior and inferior, splenius colli, levator anguli scapulas, interspinous and intertransverse. To axis, ten — the last five of preceding, and obliquus inferior, rectus posticus major, semispinalis colli, mul- tifidus spinas, scalenus posticus and transversalis colli. To the remaining vertebras are attached anteriorly ten (10) muscles, and posteriorly twenty- two (22) [vide Spinal Muscles, p. 107]. Development. — The ossific centers for each vertebra are three primary, one for the body and one for each side, and five secondary epiphyses, as follows : one for tip of each spinous and transverse process, and the remaining two are thin plates, on articular surfaces. The exceptions to this rule are : atlas, with two primary centers and one epiphysis ; the axis, with three extra centers for OSTEOLOGY. 61 odontoid process; the seventh cervical, with one extra center for each transverse process, and the lumhar vertebras, two additional centers for tubercles. The sackum (os sacrum) is a large, wedge-shaped bone, base above, apex below, with expanded lateral masses and alse, composed of five consolidated sacral vertebra?, and forming the posterior wall of pelvis. The base (basis oss. sacri) articulates with the last lumbar vertebra. Its anterior edge forms the promontory or sdcrover- FromontoTy Fig. 41. The sacrum, anterior surface. tebral angle (promontoriam) on each side of which is a smooth triangular surface, the ala (ala sacral is). Behind the artic- ulating surface is the sacral canal (canalis sacfaUs), ami on either side are the superior processes. The apex (apex oss. sacri) is small and articulates with the coccyx. The anterior concave surface (fades pelvina) presents four ridges (lineaz transversa;) indicating the original separations, which terminate externally in eight anterior sacral foramina (foramina SOCralia anlcrinra ) . with wide, shallow i/ronrcs, for the exit of the anterior sacral nerves. To the outer side of these foramina i- the lateral mass (pars lateralis) for origin of pyri- formifl muscle. 62 HUMAN ANATOMY. The posterior surface (fades dor sails) presents rudimen- tary spinous articular and. transverse processes, the lamina? of the last two segments being deficient.1 On the other side of the spinous process is a broad, shallow concavity, the sacral groove, for the origin of the erector spinse muscle, and external to the articular processes are four posterior sacral foramina (foramina sacralia posteriora) for the exit of posterior sacral nerves. The lateral surfaces have each a rough articular surface for articulation, with ossa innominata, and below presents a deep Pig. 42. The sacrum, posterior surface. notch on each side of the apex, which is converted by articulation with the transverse process of the coccyx into the fifth sacral foramen, for exit of anterior division of the fifth sacral nerve. The sacral canal (canalis sacralis), continuous above with the spinal foramen of. the vertebrae, runs through the center. Large and triangular above, small and flattened below, its lower posterior wall is deficient (hiatus sacralis). It lodges the sacral nerves, and out of it pass the anterior and posterior sacral foramina. i The articular processes of the fourth and fifth unite to form the sacral cornua for articulation with the cornua of the coccyx. OSTEOLOGY. (53 It articulates with four bones, — two os,sa innominata, last lumbar vertebra and coccyx. Muscular attachments are seven — iliacus, piriformis, coc- cygeus, gluteus niaximus, latissimus dorsi, multifidus spinae and erector spina?. Its ossific centers are thirty-five, as follows: bodies, includ- ing interarticular plates, three each (15) ; arches, two each (10) ; lateral masses, six (6) ; epiphyseal plates of lateral surfaces, four (1). Total, 35. The coccyx (os coccygis) is a small' triangular bone, re- sembling a cuckoo's beak, and composed of four rudimentary vertebra? more or less co-ossified (vertebra coccygece). •The base articulates with the sacral apex, being prolonged upward into two cornua to complete the fifth posterior sacral foramen for exit of posterior Branch of fifth sacral nerve. Laterally the base is prolonged into rudimentary transverse processes, completing with the notch of the sacrum the fifth anterior sacral foramina for exit of the anterior branch of fifth sacral nerve. The anterior concave surface supports the rectum and gives attachment to levator ani muscle and anterior sacrococcygeal ligament. The posterior convex surface presents rudimentary, artic- ular processes and transverse grooves. The apex is roimded, occasionally bifid, or deviated to one side, and affords attachment to the levator ani and sphincter ani muscles. The muscular attachments are coccygei, gluteus maximus, extensor coccygeus (occasionally present), levator and sphincter ani. Its ossific centers are four — one for each segment. THE UPPER EXTREMITY. The upper extremity consists of the arm, forearm and hand. It is attached to the trunk by the shoulder-girdle formed by the clavicles, and scapulae and contains the following bones: Clavicle, scapula, humerus, radius, ulna, eight carpus, five meta- carpus and fourteen phalanges. Tin: CLAVICLE (clavicula) , or collar- or key- bone, is a long bone, curved horizontally like the italic letter /, extending almost horizontally between the sternum and acromial process of scap- ula. Its inner two-thirds are cylindrical and convex in front, its outer third flattened from above downward and concave in front. 64 HUMAN ANATOMY. It consists of a body, or shaft, and outer, or acromial, and inner, or sternal extremities. The shaft presents in its outer third the following points : — Superior surface, rough impression in front for deltoid mus- cle, rough impression behind for trapezius; Anterior border, occasionally deltoid tubercle about the center for attachment of deltoid; Inferior surface, conoid tubercle (tuberositas coracoidea) at the posterior border for conoid ligament; oblique line ex- tending outward and forward from this for attachment of trapezoid ligament. The inner two-thirds is prismatic and presents three sur- faces and three borders: — AeraLiiulUxtV Si 'e mih t Ej.-tr FIG. 43. The left clavicle, upper surface. Anterior surface, divided by line into upper rough surface for sternomastoid muscle and lower for pectoralis major; Posterior or cervical surface gives attachment to sterno- hyoid muscle and presents foramen (foramen nutricium) for nutrient artery. Inferior or subclavian surface presents : — Facet, continuous with sternal articular surface, for car- tilage of first rib ; Rhomboid impression (tuberositas costalis) for costoclavic- ular or rhomboid ligament; Subclavian groove for attachment of subclavian muscle; Longitudinal line, frequently dividing the groove into two parts, for intermuscular septum of the muscle. Anterior border separates the anterior and posterior sur- faces and limits the attachment of the pectoralis major. Superior border gives attachment to the sternomastoid muscle. Posterior or subclavian border is short and concave, extend- ing from the rhomboid impression to the conoid tubercle and giving attachment to the subclavian fascia. OSTEOLOGY. 65 The acromial extremity {ex/ rem Has acromialis) presents an oval facet (fades articularis acromialis) for articulation with the acromial process of the scapula and a rough circumference for acromioclavicular ligaments. The sternal extremity (extremitas sternalis) is triangular, and presents a facet (fades articulans sternalis) for cartilage of sternal articulation and rough circumference for sternoclavic- vdar ligaments. The clavicle articulates with three hones — the sternum, scapula, and first costal cartilage. The muscular attachments are seven — subclavius, platysma, sternohyoid, sternomastoid, trapezius, pectoralis major and del- toid. Fig. 44. The left clavicle, under surface. end. Its ossific centers are two — one each for bodv and sternal The scapula, or shoulder Hade, is a large, flat, triangular bone situated on the upper posterolateral aspect of the thorax from the second to seventh rib, inclusive. It consists of a body, head, neck, acromion and coracoid processes, and presents two surfaces, three borders, and three angles. The anterior surface, venter or costal surface (fades cos- tal is) presents: — Subscapular fossa (fossa subscapularis) , a concave tri- angular surface for attachment of subscapular muscles, marked by Oblique ridges (linen' musculares) for the tendinous inter- sections of the muscle ; Marginal warfare, for attachment of serratus magnus; Subscapular angle (angvlus subscapularis) , a transverse depression at the upper pari for the thickest pari <>r the muscle. The posterior surface, or dorsum (fades dorsalis), presents : — 5 66 HUMAN ANATOMY. Spine (spina scapulae), a bony ridge passing upward, for- ward, and outward, dividing it unequally into two fossae, afford- ing attachment to the trapezius muscle above and the deltoid below, and terminating above in the acromion process; Supraspinous fossa (fossa supraspinata) above the spine, for the origin of the supraspinatus muscle; Infraspinous fossa (fossa infraspinata) below, for the origin of the infraspinatus muscle; Fig. 45. The scapula, anterior surface. Nutrient foramen, near the upper part of the infraspinous fossa, for nutrient artery. The marginal surface extends from the glenoid cavity down- ward and backward to an inch above inferior angle, its inner side formed by An elevated ridge, for the attachment of the aponeurosis between the infraspinatus and two teres muscles; its outer side OSTEOLOGY. 67 by the axillary border. It is divided by an oblique line into two parts, the upper for the teres minor and the lower for the teres major muscles, and crossed by a groove near the upper part for the dorsalis scapulas vessels; >*'/ Co r a cot' & Fir,. 46. The scapula, posterior surface. A smooth, triangular surface at the root <>f the spine over which the trapezius glides. The superior border (marf/o superior), shortest, presents: — Suprascapular ootch (incisura scapula') at the base of the coracoid process, converted into a foramen [suprascapular] by the transverse ligament, for the passage of the suprascapular nerve ( the artery passing above) ; 08 HUMAN ANATOMY. Origin of omohyoid muscle from this border just internal to notch. External or axillary border (mar go axillaris), thickest, presents : — Eough impression, the infraglenoid tubercle (tuberositas infraglenoidalis), just below glenoid cavity, for origin of long head of triceps muscle, and below Longitudinal groove, for origin of part of subscapulars. Internal, veretebral border — base (mar go vertebralis) , long- est, presents : — Anterior lip, for insertion of serratus magnus; Posterior lip, for attachment above of supraspinatus, below infraspinatus. •The superior angle (angulus medialis) gives attachment to part of levator anguli scapulas; The inferior angle (angulus inferior), for attachment of teres major, and sometimes part of latissimus dorsi. The anterior or external angle (angulus lateralis) is thick- est, and forms the head. The head is connected to the body of the scapula by a con- stricted portion, the neck (collum scapulae), from the anterior surface of which arises the coracoid process. It presents The glenoid cavity or fossa (cavitas glenoidalis) , a shallow, pyriform, articular depression, deepened in the recent state by the glenoid ligament, for the head of the humerus. To its upper part or apex is attached the long head of the biceps muscle. The acromion process (acromion), forming the summit of the shoulder, is a triangular process, projecting forward, out- ward and upward, to overarch the glenoid cavity. It gives at- tachment by its upper surface to the platysma and deltoid and by its inner margin to the trapezius. It presents An apex, for attachment of coracoacromial ligament; Articular facet, just behind the apex for the clavicle. The coracoid process (processus coracoideus) — like a crow's beak — arches forward, upward, and inward above the glenoid cavity. It presents Attachment for pectoralis minor muscle from the anterior border near the tip; Apex, for attachment of coracobrachialis and short head of biceps ; Rough impression at the inner side of the root for the conoid ligament, and from it A ridge running outward and forward for the trapezoid ligament. OSTEOLOGY. 69 r--.a FIQ. 47. The humerus, anterior surface. Fig. 48. The humerus, posterior surface. 70 HUMAN ANATOMY. It articulates with the clavicle and humerus. Its muscular attachments are 18 : platysma, supra- and infra- spinatus, subscapular, deltoid, trapezius, omohyoid, ser- ratus magnus, levator anguli scapulge, rhomboideus major and minor, triceps, teres major and minor, biceps, coracobrachialis, pectoralis minor and latissimus dorsi. Its ossific centers are seven — one each for body, posterior border, and inferior angle, and two each for acromion and cora- coid processes. The humerus, or arm bone, the largest and longest bone of the upper extremity, consists of a shaft, head, neck, greater and lesser tuberosities and lower extremity. The shaft (corpus humeri), cylindrical above, flattened and prismoid below, becomes twisted in the middle, and presents A rough triangular surface about the middle of its outer surface (tuberositas deltoidea) for insertion of the deltoid muscle, and a Musculo spiral groove for the musculospiral nerve and supe- rior profunda artery, on each side of which arise the external and internal heads of the triceps muscle. The upper extremity presents The head (caput humeri) forming nearly a sphere, project- ing upward, backward and inward, articulating with the glenoid cavity ; The anatomical neck (collum anatomicum) , immediately beneath, is slightly grooved for the attachment of the capsular ligament ; Greater tuberosity (tuherculum ma-jus), external to the head and lesser tuberosity, with three facets from before back- ward for attachment of supraspinatus, infraspinatus and teres minor muscles; Lesser tuberosity (tuherculum minus), smaller but more prominent than greater, is anterior to head, for the subscajmlar muscle ; Bicipital groove (sulcus intertubercularis) , passes down- ward and inward between the two tuberosities and lodges the long tendon of biceps; The anterior bicipital ridge (crista tuberculi minoris), bounds the groove in front and receives insertion of pectoralis major muscle; The posterior bicipital ridge (crista tuberculi majoris), re- ceives the latissimus dorsi and teres major; The surgical neck (collum chirurgicum) , including the head, neck and both tuberosities; OSTEOLOGY. 71 A rough impression near the center of the inner border for the coracobrachial muscle ; Nutrient canal, below and directed toward the lower ex- tremity. The lower extremity presents from within outward the fol- lowing1 : — Internal condyloid ridge, extending upward from the con- dyle ; Internal condyle or epicondyle (epicondylus medialis), more prominent than external, gives origin to the flexors and pronator radii teres; EpitrocMea, an eminence separating the trochlea from the internal condyle; Trochlea {trochlea humeri) a pulley-like articulating sur- face for greater sigmoid cavity of ulna; Coronoid fossa (fossa coronoidea) a small depression bound- ing the trochlea in front, and receiving the coronoid of the ulna in flexion ; Olecranon fossa (fossa olecrani), a larger depression behind, and receiving the olecranon process of ulna in extension; Supratrochlear foramen, sometimes formed by perforation of one fossa into the other; Radial head, or eapitellum (capitulum humeri), a smooth, rounded eminence articulating with cup-like depression on head of radius : External condyle or epicondyle (epicondylitis lateralis), less prominent, gives origin to the extensors and supinators; External condyloid ridge, extending upward on the shaft from the condyle. It articulates with three bones — scapula, radius and ulna. Its muscular attachments are twenty-five — subscapulars, supra- and infra- spinatus, teres major and minor, pectoralis major, latis&imns dorsi, deltoid, coraeobrachialis, brachialis an- ticus, triceps, pronators and flexors to inner condyle, pronator radii teres., flexor carpi radialis, palmaris Longus, flexor sublimus digitorum, flexor carpi ulnaris, supinators and extensors to outer condyle, supinator Longus, extensor' carpi radialis longior and brevior, extensor communis digitorum, extensor minimi digiti. i Some confusion exists in anatomical works because of the loose. employment of the terms condyle, epicondyle, ami epitrochlear. tt should lie borne in mind, however, thai many anatomists use Hie terms external condyle and external epicondyle synonymously, while others designate the inner condyle ;i- the internal epicondyle or epitrochlea. 72 HUMAN ANATOMY. extensor carpi ulnaris, anconeus and subanconeus, supinator brevis. Its ossific centers are seven — one each for head, shaft, tu- berosities, condyles, radial head, and trochlear portion. The ulna, or elbow bone, is a long bone to the inner side of the forearm, and consists of a shaft and an upper and lower extremity. It forms the greater part of the articulation with the humerus, but does not enter into the formation of the wrist- joint, being excluded by the interarticular fibrocartilage. The shaft (corpus ulnce) is prismatic above, smooth and rounded below, and presents : — Anterior or palmar surface (fades volaris), gives attach- ment to the deep flexors and pronator quadratus; Nutrient foramen (foramen nutricium) on anterior sur- face, directed upward toward the elbow-joint; Posterior or dorsal surface (faeies dorsalis) marked above by an oblique line for part of supinator brevis, above which is smooth triangular surface for anconeus muscle, and the lower third for extensor muscles of the thumb; External or interosseous border (crista inter ossea) , sharp in middle two-thirds, for attachment of interosseus membrane. The upper extremity is large and irregular, and presents : — Olecranon process (olecranon — head of elbow), projects upward and forward, its apex being received into the olecranon fossa of the humerus in extension of the forearm; its upper border has rough impression for the triceps muscle; its lateral borders are grooved for external and internal lateral ligaments; Coronoid process (processus coronoicleus) , smaller than olecranon, projects forward from anterior surface, being re- ceived into coronoid fossa of humerus in flexion. Its upper surface forms part of the great sigmoid cavity. Its under surface has rough impression for insertion of brachialis anticus, and has, at its junction with the shaft, the tubercle of the ulna (tuberositas ulnce) for the oblique ligament. Its outer surface is the lesser sigmoid cavity. Its inner surface gives> attachment to the internal lateral ligament, and the flexor digitorum sublimis, flexor profundus digitorum, and one head of pronator radii teres. Greater sigmoid cavity (incisura semilunaris) is a large, semilunar depression between the olecranon and coronoid proc- esses, divided into two unequal lateral parts by an elevated ridge. It is continuous on the outer side with the lesser sig- moid cavity and articulates with the trochlear surface of the humerus. OSTEOLOGY. 73 tICTdJI DICITORU 1UBLIMII ccrnrioiutZ orig. Styloid freer u Stylotd£rct4St Fig. 49. Ulna and radius, anterior aspect. 74 HUMAN ANATOMY. Lesser sigmoid cavity (incisura radialis) is an oval, con- cave, articular depression, external to the coronoid process, for articulation with the head of the radius. Its prominent ex- tremities give attachment to the orbicular ligament. The lower extremity is small and cylindrical and presents : — Head (capitulum ulnce), an external, rounded, articular process, for the triangular flbrocartilage below and the sigmoid cavity of the radius externally ; Styloid process {processus styloideus) projects from the posterior and internal part of the extremity, its apex gives at- tachment to the internal lateral ligament of the wrist, and it is marked at its root by a depression between it and the head, for attachment of the fibrocartilage ; Groove, upon the posterior surface, for passage of extensor carpi ulnaris. It articulates with two bones — humerus and radius. Its muscular attachments are sixteen — brachialis anticus, triceps, anconeus, flexor and extensor carpi ulnaris, pronator radii teres, flexor sublimus and profundus digitorum, pronator quadratus, supinator breyis, extensor indicis, extensor ossis metacarpi, extensor secundi internodii pollicis and flexor longus pollicis. Its ossific centers are three — one each for shaft, olecranon and inferior extremity. The radius is a long bone, shorter than the ulna, situated on the outer side of the forearm, the upper end small, the shaft slightly curved, and the lower end expanded to form part of the wrist joint. It consists of shaft, upper and lower extremity. The shaft (corpus radii) is prismoid, slightly curved, and presents An internal or interosseous border (crista interossea) , sharp and prominent, for. interosseous membrane; An anterior or palmar border (mar go volaris) marked at its upper third by an oblique line, for attachment of flexor longus pollicis, supinator brevis and flexor sublimis digitorum; Anterior, palmar, or flexor surface (fades volaris), affords attachment above for flexor longus pollicis, below for pronator quadratus, and presents at the junction of middle and upper two-thirds a nutrient foramen directed upward; Posterior, dorsal, or extensor surface (fades dorsalis) gives attachment at upper third to supinator brevis, and at middle third to extensors of thumb. The upper extremity presents: — OSTEOLOGY. 75 ft* 1XT. CAJIPI IAD. l/T.CMH KAO 1ST. tUUMM MTtftN J|£: '.""« Bto. 50. Radius and ulna, posterior aspect. 76 HUMAN ANATOMY. Head (capitulum radii) — a cup-like cylindrical cavity, for articulation with capitellum of humerus, and on its side an ar- ticulating surface for lesser sigmoid, cavity of ulna and orbicular ligament, which nearly surrounds it; Neck (collum radii), the constricted portion below the head; Bicipital tuberosity (tuberositas radii), below and to inner side, divided by a vertical line into a rough surface posteriorly, for attachment of biceps tendon, and smooth surface anteriorly for bursa. The lower extremity, large, expanded and quadrilateral presents : — Carpal articular surface (fades articularis carpea), smooth, concave, triangular depression divided by an anteroposterior ridge into an outer facet for scaphoid bone and inner for semilunar ; Sigmoid cavity (incisura ulnaris), a shallow concavity at inner side of carpal end, for articulation with ulnar head ; Styloid process (processus styloideus), projects obliquely downward from the external surface, for attachment by its apex to external lateral ligament of wrist joint, and by its base to insertion of supinator longus muscle. Its outer surface is marked by two grooves for extensors of thumb ; The posterior surface of the lower extremity is also marked by three grooves from without inward for the following: ext. carpi radialis longior and brevior in first, ext. secundi internodii in second, and ext. indicis, ext. communis digitorum, and ext. minimi digiti in third innermost. This surface has also attach- ment of posterior ligament of wrist. It articulates with four bones — humerus, ulna, scaphoid and semilunar. Its muscular attachments are nine — biceps, supinator longus and brevis, flexor digitorum sublimis, flexor longus pollicis, pronator quadratus, extensor ossis metacarpi pollicis, ext. primi internodii pollicis and pronator radii teres. Its ossific centers are three — one each for shaft and each extremity. The Hand. — The bones of the hand consist of eight carpus, five metacarpus, and fourteen phalanges; total, twenty-seven bones. The carpus (ossa carpi) consist of eight small bones ar- ranged in two rows — first, or proximal row, from the radial side inward are scaphoid, semilunar, cuneiform and pisiform; the OSTEOLOGY. 77 second, or distal row, in same order are trapezium, trapezoid, os magnum and unciform. Scaphoid (os naviculare mantis — boat-shaped), largest of first row, convex above, concave below, articulates with five bones, Phal, Cafpni taearpa* 2* Aw ?Jtem Fig. 51. Carpus, metacarpus, and phalanges, dorsal surface. as follows: superior surface, with radius; inferior, with trapeziuln and trapezoid; internal, with semilunar above and os magnum beneath. The external surface has attachment of ex- ternal lateral ligament of wrist. 78 HUMAN ANATOMY. Semilunar (os hmatum — half-moon), crescentic in out- line, presents articulating surfaces for five bones, as follows: superior convex surface with radius, inferior concave facet with os magnum and unciform, quadrilateral internal facet with cuneiform, and external with scaphoid. Cuneiform (os triquetrum) is wedge-shaped, the base, di- rected outward, articulates with the semilunar, apex with the interarticular fibrocartilage of wrist joint, inferior surface with unciform, and anterior with pisiform ; in all, three bones. Pisiform (os pisiforme — pea-like) is a small, pea-shaped bone articulating with anterior surface of cuneiform, and afford- ing attachment to annular ligament and two muscles, flexor carpi ulnaris, abductor minimi digiti. Trapezium (os muXtangulum — a table), a very irregular bone, articulates above by concave surface with scaphoid ; below, by surface concave from side to side, convex from before back- ward, with first metacarpal ; by internal surface with trapezoid, inferior with second metacarpal. Its dorsal surface is rough; its palmar grooved by flexor carpi radialis tendon. Muscular attachments are three — flexor ossis metacarpi, flexor brevis pol- licis and abductor pollicis. Trapezoid (os multangulum minus), smallest of second row, is wedge-shaped, apex palmar ; articulates with four bones — superior surface with scaphoid, inferior with second metacarpal, external with trapezium, internal with os magnum and inter- osseous ligament. It has one muscular attachment for flexor brevis pollicis. Os magnum, or capitate (os capitatum), the largest car- pal bone, consists of a body, neck, and head. The latter projects from the superior surface and articulates with the scaphoid and semilunar. The inferior surface articulates with second, third, and fourth metacarpal bones; the external with trapezoid, and internal with unciform; in all seven bones. Muscular attachment for flexor brevis pollicis. Unciform (os hamatum — hook-like), a wedge-shaped bone, so named from the hook-like process (hamulus ossis hamati) projecting from its anterior surface, below its' "articu- lation with fourth and fifth metacarpals, and above its articula- tion with semilunar. This process gives attachment to annular ligament, flexor brevis minimi digiti, flexor ossis metacarpi minimi digiti, and is grooved for passage of other flexor ten- dons. External surface articulates with os magnum and inter- nal with cuneiform. OSTEOLOGY. 79 Muscular attachments are those to unciform process. Each carpal bone is developed from a single ossific center. Fig. 52. Carpus, metacarpus, and phalanges, palmar surface. Table of Articulations of Carpal Bones. — Scaphoid; 5; semilunar, 5 ; cuneiform, 3 ; pisiform, I ; trapezium, I ; trapezoid, I ; oa magnum, ', ; unciform, 5. 80 HUMAN ANATOMY. The metacaepus (ossa metacarpalia) are five long bones, resembling each other, and presenting a shaft (corpus) and two extremities. Anterior surface is concave, posterior convex; superior, carpal or proximal extremity or base (basis), irregular for articulation with carpal bones and with the adjoining bones; inferior, digital or distal extremity or head (capitulum) , pre- sents rounded head for articulation with phalanges. The first metacarpal (os metacarpale I), for thumb is supported on trapezium, articulates with first phalanx, and has four muscles attached— first dorsal interosseus, flexor, and extensor ossis metacarpi pollicis and the flexor brevis pollicis; the second metacarpal (os metacarpale II) by three, trapezium, trapezoid and os magnum, articulates also with third metacarpal and second phalanx, and has six muscles attached — first and second dorsal interosseus, first palmar interosseus,. flexor carpi radialis, and extensor carpi radialis longior and the adductor obliquus pollicis; the third metacarpal (os metacarpale III) supported by os magnum, articulates also with second and fourth metacarpal and third phalanx, and has six muscles attached — flexor carpi radialis, adductor obliquus pollicis, extensor carpi radialis brevior, adductor pollicis, and second and third dorsal inter- osseus. The fourth metacarpal (os metacarpale IV) is supported on os magnum and unciform, articulated with third and fifth metacarpal and fourth phalanx, and has three muscles attached — second palmar and third and fourth dorsal interosseus. The fifth metacarpal (os metacarpale V) supported on unciform, articulates with fourth metacarpal and fifth phalanx, and has five muscles attached — flexor ossis metacarpi minimi digiti, flexor and extensor carpi ulnaris, and fourth dorsal and third palmar interosseus. Ossific centers are two for each bone, one each for shaft and head, except thumb, which has one each for shaft and base, resembling the phalanges. Phalanges of the Hand (phalanges digitorum manus). — Each finger has three phalanges (intemodia), except the thumb, which has but two. Palmar surface concave, dorsal con- vex; the superior extremity articulates with head of metacarpal, and the inferior, concave from side to side, convex from before backward, articulates with second phalanx, which presents an opposite arrangement, except in the thumb, where it articulates with ungual or terminal phalanx. [Phalanges as a generic term is translated, intemodia; when applied to the phalanges of the hand, phalanges digitorum OSTEOLOGY. 81 manus, in contradistinction to the phalanges of the foot, pha- langes digitorum pedis.] The second phalangeal articulation is the same, and the ungual phalanx presents a rough, arrow-shaped extremity. The muscular insertions to the first row are : to thumb four, flexor brevis, abductor and adductor pollicis, and extensor primi internodii; to index two, first palmar and dorsal interosseus; to middle two, second and third dorsal interossei; to ring, two, second palmar and fourth dorsal interosseus; to little three, flexor brevis and abductor minimi digiti and third palmar inter- osseus. The insertions to second row are: extensor secundi inter- nodii and flexor longus pollicis; and to each of the others four, extensor communis digitorum, flexor sublimis digitorum, with the addition of extensor indicis to index and extensor minimi digiti to little. The insertions to third row are: extensor communis digi- torum and flexor profundus. Ossific centers are two for each bone, one each for shaft and base. THE LOWER EXTREMITY. The lower extremity consists of the thigh, leg, and foot. It is connected to the trunk by the pelvic girdle formed by the hip bones and sacrum, and contains the following bones : os innom- inatum, femur, patella, tibia, fibula, seven tarsus, five meta- tarsus and fourteen phalanges. The os inxominatum, or hip bone, is a large, irregular bone forming the lateral and anterior walls of the pelvis, and consisting of three bones, ilium, ischium and pubes, united about puberty. It presents The acetabulum or cotyloid cavity, a deep, cup-shaped cav- ity, for articulation of the head of the femur — the ischium forming a little more than two-fifths, the ilium a little less than two-fifths, and the pubes one-fifth. The bottom of the cavity presents a circular depression (fossa acetabuH), lodging a mass of Eat, and giving attachment by its edges to the ligamentum teres, and continuous below with a deep notch, the cotyloid aotch {inciswra acetabidi), for attachment of cotyloid and part of Ligamentum teres, converted into a foramen by the trans- verse ligament for (he entrance of the nutrient vessels and nerves. The margin of the acetabulum is deepened by a. fibro- cartilaginous ring. The obturator, or thyroid, foramen (foramen obiuratvm) is 82 HUMAN ANATOMY. an aperture large and ovoidal in the male, small and triangular in the female, situated on the anterior inferior surface between the ischium and pubes. It is closed, in the recent state, by a strong membrane attached to its margins except above exter- nally, where, a foramen exists for obturator vessels and nerve. OESCLUII WffUUBI Fig. 53. Os innominatum, outer surface. The ilium (os ilium), broad, flat, and triangular, forms the greater part of the bone, its base above, its apex at the acetabulum. The external surface, or dorsum, concave in front and be- hind, presents from below upward OSTEOLOGY. y;{ A groove, just above the acetabulum, for the reflected ten- don of the rectus femoris muscle; The inferior curved line, or inferior "luteal line (linea glutecB inferior), marking the lower border of the gluteus minimus ; The middle curved line, or anterior gluteal line (linea Fig. 54. Os innominatum, inner surface. glutem anterior), the longest of the three, marking the lower border of the gluteus medius ; and The superior curved line, or posterior gluteal line (linea glutecB posterior), the shortest, marks the lower anterior border of the glutens maximus. and from the Burface below which a i ise a few fibers of the pyriformis. The upper expanded border forms the- cresl of the ilium ta Uiaca \ , terminal ing in fronl in §4 HUMAN ANATOMY. The anterior superior spinous process {spina iliaca anterior superior);, giving origin to the sartorius and tensor vaginas femoris muscles and Poupart's ligament, below which is The anterior inferior spinous process (spina iliaca anterior inferior), for the iliofemoral ligament, and the straight tendon of the rectus femoris muscle; and behind in The posterior superior spinous process (spina iliaca pos- terior superior), for the oblique band of the sacroiliac ligament, and part of the multifidus spina? muscle, below which is The posterior inferior spinous process (spina iliaca pos- terior inferior) , for attachment of the great sacrosciatic ligament. Between the superior and inferior spinous processes, both anteriorly and posteriorly, is a notch, the former for partial at- tachment of the sartorius and passage of the external cutaneous nerve. The internal concave surface, or venter, presents The internal iliac fossa (fossa iliaca), lodging the iliacus muscle, and having a nutrient foramen at its lower part; Iliopectineal line (linea arcuata), limiting the fossa below, and separating the false from the true pelvis; Rough surface, divided into two parts — the posterior por- tion (tuberositas iliaca), for posterior sarcoiliac ligaments, and lower auricular or anterior surface (fades auricularis) for ar- ticulation with sacrum. The ischium (os ischii) forms the outer back part of pel- vis, and consists of a body, tuberosity, and ascending ramus. The external surface of the body forms a little more than two-fifths of the acetabulum, and presents A groove (sulcus obturatorius) below, for the tendon of the obturator externus. The internal surface is concave and smooth, and forms the lateral wall of the true pelvis. The posterior border presents The spine (spina ischiadica) of the ischium^ projecting downward, backward, and inward, from below the center for attachment of the gemellus superior, coccygeus, and levator ani muscles, and lesser sacrosciatic ligament; Great sacrosciatic notch (incisura ischiadica major), a deep notch above the spine converted into a foramen (foramen ischiadicum majus), by the lesser sacrosciatic ligament, trans- mitting the pyriformis muscle, superior gluteal nerve, gluteal vessels, sciatic vessels and nerves, and the internal pudic vessels and nerves ; OSTEOLOGY. 85 Lesser sacrosciatic notch (incisura ischiadica minor), be- low the spine, and between it and the tuberosity, converted into a foramen (foramen ischiadicum minus) by the great sacro- sciatic ligament, transmitting the obturator internus muscle and nerve, the internal pudic vessels and nerves. The lowest por- tion of the body presents Tuberosity (tuber ischiadicum), with Outer lip, for attachment of part of adductor magnus and quadriceps f emoris ; Inner tip, for part of great sacrosciatic ligament, erector penis, and transversus perinaei; Groove, on inner lip, for internal pudic vessels and nerve; Intermediate surface, for semimembranosus, semitendino- sus, biceps, adductor magnus, gemellus inferior and great sacro- sciatic ligament. The ascending or inferior ramus (ramus inferior ossis ischii) passes upward and inward from the tuberosity to join the descending ramus of the pubes, forming part of the inner margin of the obturator foramen. It gives attachment to gracilis, obturator externus, part of adductor magnus, erector penis and transversus perinaei; The pubes (os pubis) forms the anterior part of the pelvis and consists of a body, horizontal ramus, and descending ramus. The body (corpus ossis pubis) is quadrilateral, and presents An anterior surface, for attachment of adductor longus and brevis, and part of gracilis, adductor magnus and obturator ex- ternus ; Posterior surface, forming anterior wall of true pelvis and giving attachment to levator ani and part of obturator internus ; Spine (tuberculum pubicum) , upon the upper border, for Poupart's ligament and outer pillar of external abdominal ring; Iliopectineal line (linea arcuata), continuous with that on ilium ; Crest, along the upper border, internal to the spine; Angle, at the junction of the inner border with the crest and giving attachment to internal pillar of external ring; Symphysis, the internal oval border roughened by several ridges for articulation with opposite bone. The ascending or superior ramus (ramus superior ossis pubis) joins t lie ilium, forming the upper part of the obturator foramen, ami presents at its lower border a Qroove, for the obturator vessels and nerve. The descending or Inferior ramus (ramus inferior ossis gg HUMAN ANATOMY. pubis) is flat and thin, and joins the ascending ramus of the ischium. Each innominate bone articulates with three bones — the sa- crum, femur, and its fellow of the opposite side. Its muscular attachments are thirty-three — [ilium], latissi- mus dorsi, tensor vaginas femoris, obliquus externus, erector spinas, transversalis, quadratus lumborum, gluteus minimus, medius, and maximus, rectus, piriformis, iliacus, multifidus spinas, sartorius — [ischium], obturator externus and intern us, gemellus superior, coccj^geus, levator ani, semimembranosus and semitendinosus, biceps, quadratus femoris, adductor magnus, gemellus inferior, erector penis and transversus perinasi — ■ [pubes], pyramidalis, obliquus internus and externus, psoas par- vus, pectineus, adductor longus and brevis, gracilis and com- pressor urethras. Its ossific centers are eight — one primary for each division and five epiphyses, one each for crest of ilium, symphysis, tuber- osity of ischium, anterior spinous process, and acetabulum. The femur, or thigh bone, is the largest, longest and strongest bone in the body. In the erect position it inclines toward its fellow at the knee, being widely separated above, forming the sides of a triangle, the base of which isi greater in females from the greater breadth of the pelvis. It consists of a shaft, an upper and lower extremity. The shaft (corpus femoris) expanded above and below, curved and twisted, convex in front, concave behind, is nearly, cylindrical throughout, and presents Smooth anterior surface, for origin of crureus and sub- crureus muscles; Lateral surfaces, covered by the vasti externus and internus ; The posterior surface, rough and prominent, has its Nutrient foramen between the middle and lower two-thirds directed upward; Linea aspera, a rough, prominent, longitudinal crest de- scending from the trochanters along the middle third, bifur- cating and diverging at the inferior extremity to the condyles, inclosing The popliteal space (planum popliteum) , a smooth, trian- gular space, on which rests the popliteal artery, and which is Grooved at its inner margin by the femoral artery. The outer (labium laterale) and inner (labium mediate) lip of the linea aspera give attachment to the vasti externi and interni, three adductors, pectineus, biceps and gluteus maximus. The upper extremity presents the following: — OSTKOLOOY. 87 ifirUud^ Fio. 66. The femur, anterior surface. Fig. 56. The femur, posterior surface. gg HUMAN ANATOMY. The head {caput femoris), forming two-fifths of a sphere, articulates with the acetabulum, having a central oval depression (fovea capitis femoris) for the ligamentum teres; Neck (collum femoris), pyramidal, with excavated surfaces, connects the head with the shaft, the angle of its obliquity to the shaft varying much from puberty to old age, being, in the adult, about 130 degrees; The great trochanter (trochanter major), a large, rough, quadrilateral eminence, directed upward, outward, and back- ward, its external surface marked by a diagonal line for inser- tion of gluteus medius tendon, its outer surface smooth for the passage of the gluteus maximus tendon, separated by a bursa, and its superior surface marked by three impressions, from be- hind forward, for pyriformis, obturator internus and gemelli muscles. The gluteus minimus is attached to the anterior border ; Digital or trochanteric fossa (fossa trochanterics), to the inner side of the great trochanter, for insertion of obturator externus tendon; The lesser trochanter (trochanter minor), a small, conical projection at the base of the neck posteriorly, and giving attach- ment to the tendon of psoas magnus muscle, the iliacus being inserted below; Anterior intertrochanteric line (linea intertro chant erica) connects the trochanters in front and gives attachment at its upper part to the capsular ligament; Posterior intertrochanteric line (crista intertro chanterica) , a much more prominent ridge, connects them behind ; Tubercle of the femur, a prominence at the junction of the neck with the great trochanter, is the meeting-place of five mus- cles— two gemelli, obturator internus, gluteus, minimus and vastus externus; Tubercle of the quadratus, about the center of the posterior intertrochanter line, for the quadratus femoris; Linea quadrati, passes vertically downward from the mid- dle of the posterior line, and gives attachment to part of adductor magnus and quadratus femoris. The inferior extremity, large and cuboidal in form, pre- sents : — External condyle (condylus lateralis), shorter and broader than the internal, has behind its center the outer tuberosity for the external lateral ligaments of the knee, and a groove below the tuberosity for the tendon of the popliteus muscle. It also OSTEOLOGY. 89 gives origin to the outer head of the gastrocnemius, above which arises the plantaris muscle ; The internal condyle (condylus medialis), longer by half an inch and more prominent (so as to bring them on the same horizontal plane owing to the obliquity of the shaft), has on the inner surface the inner tuberosity for the internal lateral ligament; Adductor tubercle, at the summit of the internal condyle, marks the termination of the inner ridge of the linea aspera, and gives attachment to tendon of adductor magnus; Depression, above the articular surface of the internal con- dyle posteriorly, for the origin of the inner head of gastrocne- mius; Intercondyloid notch (fossa intercondyloidca) , separates the condyles behind, and lodges the crucial ligaments; Trochlear (fades patellaris) is a smooth surface between the condyles in front and articulates with the patella in front. The inferior surfaces of both condyles are smooth, continuous in front, covered with cartilage in the recent state, for articula- tion with the head of the tibia. The femur articulates with three bones — innominatum, tibia and patella. Its muscular attachments are twenty-three (23) — two vasti, three adductors, gluteus maximus, medius and minimus, pyri- formis, obturator externus and interims, two gemelli, quadratus femoris, psoas magnus, iliacus, biceps, pectineus, crureus, sub- crureus, gastrocnemius, plantaris and popliteal. its ossific centers are five — three primary, one each for shaft and each extremity, and one epiphysis for each trochanter. The patella is a flat, triangular bone, sesamoid in origin, developed in the tendon of the ^quadriceps extensor, forming the knee-cap, and entering into the formation of the knee-joint. The convex anterior surface is roughened by apertures for nutrient vessels. The posterior or internal surface (fades articularis) is divided by a vertical and transverse ridge into three surfaces — two smooth, articular facets for either condyle of femur above, the outer being deeper and broader, and a rough surface below, the apex (apex pateUce) for ligamentum patellae. The superior border (basis patella') gives attachment to the rectus and CTureus muscles, tbe internal and external lateral bor- ders to the vasti internus and externus. It articulates with the femur. [ta muscular attachments are four muscles — rectus Eemoris, crureus, vastus externus and vastus internus. 90 ' HUMAN ANATOMY. It is developed from a single ossific center. The tibia, or shin bone, extends on the internal aspect of the leg from the knee to ankle, and consists of a shaft, upper and lower extremity. The shaft (corpus tibia?) is triangular, prismoid, with the base above. Its anterior border and internal surface are subcutaneous, the former forming the shin or crest of the tibia (crista anterior) . The posterior surface (fades posterior) presents above an oblique line (linea popiitea) for the lower border of the pop- liteal muscle and fascia, and origin of the soleus, and parts of flexor longus digitorum and tibialis posticus muscles. It presents just below the oblique line a nutrient canal (foramen nutricium), the largest in the skeleton, directed downward. Its external border, or interosseous ridge (crista interossea) is thin, for attachment of interosseous membrane. The external surface and internal borders are covered by muscles. The superior expanded surface presents: — Head, consisting of two tuberosities, internal and external (condylus medialis and condylus lateralis), each having upon their upper surfaces a smooth, concave, ovoidal articulating facet (fades articularis superior) for the condyles of the femur, separated by The spinous process (eminentia intercondyloidea) of the tibia, in front and behind which is impression for crucial liga- ments, and laterally tubercles for the extremities of the semi- lunar cartilages; Tubercle (tuberositas tibial), on the anterior surface of the head, between the tuberosities, for the insertion of ligamentum patellae ; Popliteal notch (incisura popiitea), separating the tuber- osities posteriorly, for attachment of the posterior crucial ligament ; Transverse groove, on the posterior surface of the inner tuberosity, for insertion of semimembranosus tendon; Facet (fades articularis fibularis), upon the posterior sur- face of the outer tuberosity, for articulation of the head of the fibula. The inferior extremity, smaller than the superior, is quadri- lateral, and presents in front a smooth surface for extensor ten- dons ; behind, a groove for flexor longus pollicis tendon ; exter- OSTEOLOGY. 91 jtfro* ■^''"/^H. TrtTTB I H'U.~.t Fig. 57. The tibia and fibula, anterior surface. 'tyUiSfmem FIG. 58. The tibia and fibula, posterior surface. 92 HUMAN ANATOMY, nally, a rough, triangular, articular surface, for fibula, and in- ternally The internal malleolus (malleolus medialis), which projects downward, and articulates by its outer surface with the as- tragalus. Its inner surface is smooth and subcutaneous; its posterior border is grooved (sulcus malleolaris) for the tendons of the tibialis posticus and flexor longus digitorum muscles, and to its tip is attached the internal lateral ligament. It articulates with three bones — femur, fibula and as- tragalus. Its muscular attachments are twelve — ligamentum patellae, popliteus, soleus, flexor longus digitorum, tibialis posticus, semi- membranosus and tendinosus, gracilis, sartorius, tibialis anticus, extensor longus digitorum and biceps. It is developed from three ossific centers, one each for shaft, upper and lower extremities. The fibula, peroneal, or calf-bone, is a long, slender bone on the outer aspect of the leg, and consists of a shaft, upper and lower extremity. The shaft (corpus fibulce) is prismoidal, four-sided, twisted on itself, and arched backward. Its antero-internal border, or interosseous ridge (crista interossea), gives attachment to the interosseous membrane. The postero-internal border (crista medialis), called also oblique line, and the other surface and borders give attachment to all the muscles except the biceps. Nutrient foramen, about the middle of the anterior-internal surface, is directed downward. The superior extremity presents a neck, supporting a rounded, irregular head (capitulum fibula), which articulates by a flattened facet on its inner surface with the tibia, and is prolonged upward on its outer side by the styloid process (apex capituli fibula), giving attachment to the biceps tendon and the external lateral ligament of the knee. The inferior extremity expands into the external malleolus (malleolus lateralis), with its convex internal surface for ar- ticulation with astragalus, outer convex surface subcutaneous, and posterior border deeply grooved (sulcus malleolaris) for tendons of peroneus longus and brevis muscles. To the summit is attached the middle fasiculus of the ex- ternal lateral ligament, and to rough depressions in front and behind the anterior and posterior fasciculi. It articulates with two bones, tibia and astragalus. OSTEOLOGY. 93 Its muscular attachments are nine : soleus, biceps, three peronei, tibialis posticus, flexor longus ballucis, extensor longus digitorum and extensor longus hallucis. It is developed from three ossific centers, one each for shaft, head and malleolus. THE FOOT. The bones of the foot consist of seven tarsus, five metatar- sus, and fourteen phalanges. The tarsal bones (ossa tarsi) are: calcaneum, astragalus, cuboid, scaphoid, and internal, middle, and external cuneiform. They may be arranged into two series, anterior and poste- rior; the calcis and astragalus behind, and all the others in front of the calcaneocuboid and astragaloscaphoid joints [Cho- part's]. Calcaneum, or os calcis, the largest, is irregularly cuboidal. The superior surface presents two articular surfaces for the astragalus, separated by a groove for the calcaneoastragaloid ligament, and internally a projecting process, the sustentaculum tali, fur calcaneocuboid ligament. The inferior surface, rough and excavated, presents two tubercles, an outer and inner tubercle, for muscles and ligaments. Internal surface is concave, for passage of flexor longus and tibialis posticus tendons, and plantar vessels and nerves. External surface presents tubercle {processus trocldearis) for external lateral ligament of ankle and grooves for peroneal tendons. Posterior surface, projecting behind {tuber calcanei), pre- sents a smooth surface above for bursa and rough below for attachment of tendo Achillis. It articulates with astragalus and cuboid. Muscular attachments are eight: tendo Achillis, plantaris, tibialis posticus, abductor hallucis, a. minimi digiti, flexor brevis digitorum, accessorius and extensor brevis digitorum. Astragalus (tales) is an irregular, short bone, consisting of body (corpus tali), neck (collum tali), and head (caput tali). The quadrilateral body presents four articular surfaces, above for lower extremity of tibia and internal malleolus and external malleolus; below, two surfaces for os calcis, separated by deep groove for interosseous calcaneoastragaloid ligament; posteriorly it presents a deep groove (sulcus m. flexor hallucis longi) for flexor Longue ballucis tendon; and anteriorly, the rounded con- vex head supported on neck articulates with scaphoid, and rests upon calcaneoscaphoid ligament. 94 HUMAN ANATOMY. It articulates with four bones : tibia, fibula, scaphoid and os. calcis. It has no muscular attachments. Cuboid (os cuboideum—cuhe-like) is a small, pyramidal bone, between the os calcis and the fourth and fifth metatarsal bones on the outer side of the foot. * The upper or dorsal surface is rough for ligamentous at- tachment; the lower or plantar surface is grooved (sulcus m. peronwi longi) for tendon of peroneus longus, behind which is a ridge for the long calcaneocuboid ligament, terminating ex- ternally in the tuberosity of the cuboid (tuberositas ossis cuboidei). The external surface has a deep notch, the outer extremity of the peroneal groove. The posterior surface has triangular facet for os calcis, the anterior has two facets, separated by a ridge for the fourth and fifth metatarsals, and the internal surface has broad, square facet for external cuneiform, and sometimes a smaller facet for scaphoid. It articulates with four and occasionally with five bones. It has one muscular attachment — part of flexor brevis pol- licis. Scaphoid, or navicular bone (os naviculare pedis), is a boat-like bone placed between astragalus and three cuneiform bones. Its posterior concave surface articulates with head of as- tragalus, its anterior convex surface has three facets for cunei- form bones; its internal border presents the tuberosity of the scaphoid (tuberosit as ossis navicularis) for insertion of tibialis posticus, the only muscular attachment. Its other borders are roughened for ligamentous attachment. It articulates with four bones : astragalus and three cunei- forms. The cuneiform bones are named from their position the internal, middle and external. The internal cuneiform (os cuneiforme primum), the largest, has its base below, and articulates anteriorly with first metatarsal, posteriorly with scaphoid, and externally with second metatarsal and middle cuneiform. The plantar surface presents tuberosity for insertion of part of tibialis posticus and anticus tendons, the only muscular attachments. It articulates with four bones: scaphoid, middle cuneiform, first and second metatarsals. The middle cuneiform (os cuneiforme secundum), the smallest, has its base upward, and articulates posteriorly with OSTEOLOGY. 95 Tn„trmas$ UruL,n tf- S\% DICITORWtt Fig. 59. The tarsus, metatarsus, and phalanges, dorsal surface. gg HUMAN ANATOMY. scaphoid, anteriorly by a triangular facet with second metatar- sal, and laterally with internal and. external cuneiforms. It has no muscles attached. The external cuneiform (os cuneiforme iertium), in- termediate in size, and more regular, articulates posteriorly with scaphoid, anteriorly with third metatarsal, internally with middle cuneiform and second metatarsal, and externally with cuboid and fourth metatarsal. Muscular attachments are for flexor brevis pollicis and tibialis posticus. The metatarsal bones (ossa metatarsalia) have the same general form, as the metacarpal bones of hand, each consisting of shaft, head and base. The shaft (corpus) is prismoid and curved, with concavity below, convexity above. The head (capitulum) is rounded for articulation with phalanges, and has tubercles laterally for ligaments and a groove below for tendon of long flexor. The base (basis) is wedge-shaped for articulation with the tarsus and with each other. The first (os metatarsale I) and strongest articulates at base with internal cuneiform, at head with phalanx and second metatarsal, and has three muscular attachments : for tibialis anticus, peroneus longus and first dorsal interosseous. The second (os metatarsale II) is wedged in by base be- tween three cuneiform bones, articulates with second phalanx, first and third metatarsi, and has three muscular attachments : for adductor hallucis, first and second interosseous. The third (os metatarsale III) articulates with external cuneiform below and second and third metatarsi and third phalanx, and has four muscular attachments : for adductor hal- lucis, first plantar, and second and third dorsal interossei. The fourth (os metatarsale IV) articulates at base with ex- ternal cuneiform and cuboid and at extremity with 'third and fifth metatarsal and fourth phalanx, and has four muscular at- tachments: for adductor hallucis, third and fourth dorsal, and second plantar interossei. The fifth (os metatarsale V) has marked tubercle on outer side. It articulates with cuboid, fourth metatarsal, and fifth phalanx, and has five muscular attachments : for flexor brevis minimi digiti, peroneus brevis and tertius, fourth dorsal and third plantar interosseus. The phalanges of the foot (phalanges dig it o rum pedis) have same general characteristics as in hand, but are compressed OSTEOLOGY. 97 ku*a mi.) rtuiohf Fig. 60. The tarsus, metatarsus, and phalanges, plantar surface. 98 HUMAN ANATOMY. from side to side instead of from before backward ; and have the following nomenclature applied : first or proximal phalanx (phalanx prima) ; second phalanx (phalanx secunda) ; ungual or distal phalanx (phalanx tertia) . The muscular insertions to first row are : to great toe, transversus pedis, extensor brevis digitorum, adductor, abductor and flexor brevis hallucis; to second, first and second dorsal interosseus; to third, third dor- sal and first plantar interosseus; to fourth, fourth dorsal and second plantar interosseus; to fifth, abductor and flexor brevis minimi digiti and third plantar interosseus. To second row : to great toe, flexor and extensor longus hallucis; to remaining toes, extensor longus and brevis digitorum, flexor brevis digi- torum and lumbricales. To third row, or ungual phalanges: flexor longus and extensor longus and brevis digitorum. The ossific centers are one each for the tarsus, excepting the os calcis, which has an epiphysis for posterior part, and two each for metatarsals and phalanges. These latter are arranged in phalanges, one each for shaft and base, and in metatarsals, one each for shaft and head, excepting great one, which has one each for shaft and base. Sesamoid bones (ossa sesamoidea) are small, osseous masses, cartilaginous in early life, developed in the tendons to relieve pressure. They are of two kinds — -those over articular surfaces of joints, as patella; and those applied to the surfaces of bones, as in tendon of peroneus longus in the groove of cuboid bone. OSTEOLOGY. 99 Name. Yertebr.e. Atlas (3). Axis (6). Seventh Cervical. Lumbar Verte- brae (2). Sacrum (35). 15. 10. 6. 4. Coccyx (7). f'KANIAl, Bones. Occipital (7). Parietal (1), TABLE OF OSSIFICATION. Centers. Primary: — 1 for each lamina. 1 for each process. 1 for body. Secondary Centers: — 1 for each transverse process. 2 for end of spinous process. Additional Plates: — ■ 1 epiphyseal plate on upper and under surface of bod v. 1 for each lateral mass. 1 epiphysis for anterior arch. 1 for lower part of body. 1 for each lamina. 2 for lateral processes. 1 for apex of odontoid process. 1 for anterior and costal part of transverse process. 1 for each tubercle of superior articular process. 3 — 1 each for body and upper and lower epiphys- eal plate. 2 — 1 for each arch. 2 for eacli lateral mass of the fust three vertebrae. Each lateral surface has one epi- physeal articulation and ad- joining edge. 1 for each piece. 1. 2. 3. 4. 4 for tabular or epi-oceipital. 1 for each condylar portion. 1 for basilar portion. From membrane. ^iiiL'le center at parietal eminence Time of Appearance. 6th week. 8th week. 16th year. 21st year. All unite by 30th year. 8th week. 1st year. 6th month. 6th fetal month. 6th fetal month : joins 5th or 6th rear. 8th or 9th week. 16th year, and un'tes 18th or 20th vear. 6th or 8th month; joins 2d to 6th vear. 18t.li year, to 25th Birth. 5th to 10th year. 10th to 15th year. 15th to 20th vear. 8th fetal week. -1 tli in 6th year: joined In sphe- noid 18th to 25t'i year. 7th to 8th fetal week. 100 HUMAN ANATOMY. "Name. Frontal (2). Temporal (10). Sphenoid (14). Centers. 1 for each lateral portion. From membrane. 1 for squamous process. 1 for auditory process. 6 for petromastoid. 2 for styloid process. 8 for postsphenoid. 1 for each greater wing and ex- ternal pterygoid plate. 1 for each internal pterygoid plate. 2 for posterior part of body. 1 for each lingula. 6 for presphenoid: — 1 for each lesser wing. 2 for anterior portion of body. 1 for each sphenoturbinal. Ethmoid (3), 1 for each lateral mass. 1 for lamella. Face. Nasal. 1 for each bone. Superior Maxillae (4). 1 premaxilla. 1 prepalatine portion. Lachrymal. Malar. Palate. Inferior Turbi- nated. 1 maxillary portion. 1 malar portion. A single center. Single center. Single center at junction of plates. Single center. Vomer. Single center, but two laminae. Inferior Maxilla. Probably several centers; at birth consists of two halves. Body. Hyoid (5). 1 for body and for each cornu. Time of Appearance. 7th to 8th week. United by 4th year. 2d month. Later. 5th or 6th month. 1 for base ap- pears before birth and one after birth. Appear from 8th week to third year, and union of all parts is accom p 1 i s h e d by the 20th year, and with occipital, 18th to 25th year. 4th to 5th fetal month. Unite about 2d year. 6th week. Very early. Antrum appears about fourth fetal month. 7th week. 6th week. 2d fetal month. Middle of fetal life. 8th week. Very early, be- ing second in order. 3 months after birth. OSTEOLOGY. 101 Name. Sternum (6), Ribs (3). Upper Extremity. Clavicle (2). Scapula ( 7 ) . Humerus ( 7 ) . Ulna (3). Radius (3). Carpus. Centers. 1 for manubrium. 4 for gladiolus. 1 for ensiform appendix. 1 for each shaft, head, and tuber- cle, excepting the last two, which have but 2, the tuber- cles being absent. 1 for shaft. 1 for sternal extremity. 1 for body. 2 for coracoid process. 2 for acromion process. 1 for posterior border. 1 for inferior angle. 1 for shaft. 1 for head. 1 for tuberosities. 1 for radial head. 1 trochlear portion. 1 for each condyle. 1 for shaft. 1 for olecranon. 1 for lower extremity. 1 for shaft. 1 for upper extremity. 1 for lower extremity. Each lias single center: — I •- magnum. I 'nciforni. Cuneiform. Trapezium and semilunar. Scaphoid. Trapezoid. 1'isiform. Time of Appearance. Middle of fetal life. Before vertebrae. Epiphysis for tubercle ap- pears from 16th to 20th year, and unites about 25th. About 30th day. 2d month. Completed be- tween 20th and 25th year. !-'tli week. Upper extremity ossifies 2d to 4th year. Complete about 20th year. 8th week. 10th year. 4th year. Soon after hu- merus. 5th year; unites 17th to 18th year. 2d year; unites 20th year. Upper extremity joins 16th year; lower, 20 th year. 1st year. 1st year. 3d year. 5 tli year, fitll year. 8th year. 12th year. 102 HUMAN ANATOMY. Name. Metacarpus ( 2 ) . Phalanges ( 2 ) . Centers. Centers for each: — 1 for shaft and 1 for each distal extremity ex- cept thumb, which has one each for shaft and base. 1 each for shaft and base. Time of Appearance. 8th or 9th week. 3d year, and unites about 20th year. 8th week, and unites 18th to 20th year. Lower Extremity. Os Innominata (8). 3 Primary: — Ilium. Same as verte- • brse, 6th week. Ischium. 3d month. Pubes. 4th or oth month. 5 Secondary: — • 1 crest of ilium. Puberty. 1 anterior inferior spinous proc- ess. " 1 tuber ischii. it 1 symphysis pubis. " 1 acetabulum. 13th or 14th year; completed 25th year. Femur (5). 1 for shaft. 5th week. 1 for lesser trochanter. End of 1st year 1 for great trochanter. 9th fetal month. 1 for lower extremity. 4th year. 1 for upper extremity. 13th or 14th year. Inferior extrem- ity. The last is not united before the 20th year. Patella. Single center. 3d to 6th year; completed about puberty. Tibia (3). 1 for shaft. 7th week. 1 upper extremity. Birth ; unites 20th year. 1 • lower extremity. 2d year ; unites 18th year. Fibula (3). 1 for shaft. 8th fetal week. 1 upper extremity. 4th year. 1 lower extremity. 2d year ; the lower unites first — about 20th year. OSTEOLOGY. 103 Name. Tarsus. Metatarsus. Centers. os calcis: — 1 for body and 1 for posterior extremity. each for the following: — Cuboid. Astragalus. Scaphoid. Internal cuneiform. Middle cuneiform. External cuneiform. 1 each for shaft and digital extremity, except great toe, which is same as thumb. Phalanges (2). 1 each for shaft and proximal ex- tremity. Time of Appearance. 6th fetal month. 9th month. 7th month. 4th year. 3d year. 4th year. 1st year. 7 th week. 3d year. ARTICULATIONS AND LIGAMENTS. The bones of the skeleton are connected together by articu- lations or joints. These consist essentially of the expanded extremities of bones, covered with cartilage, often separated by interarticular fibrocartilage, held together by ligaments, and lined by synovial membrane. The bone entering into the articular lamella differs from ordinary bone by its extreme density, without Haversian canals, its lacunae being much larger, and without canaliculi. It is not perforated by blood-vessels. Cartilage is a whitish, highly elastic, non-vascular struc- ture, forming in the fetus the greater part of the skeleton, and found in the adult chiefly in the joints, the walls of the thorax, and certain orifices, as the nostrils, ears, etc. It is either temporary, becoming ossified later, or perma- nent, remaining unossified. The latter is divided into three varieties: articular., in joints covering the ends of the bones; costal, forming part of the thorax; and reticular, arranged in plates or lamellae to maintain the shape of parts. Fibrocartilage consists of a mixture of cartilaginous with white fibrous tissue. There are four varieties: — (a) Inter articular, interposed between the joint surfaces; (b) Connecting, binding bones together as in pubes; (c) Circumferential, deepening cavities, as glenoid cavity of shoulder; and (d) Stratiform, lining grooves for tendons. Synovial membranes are of three kinds: articular, lining the cavities of movable joints throughout except the surface of the cartilage; bursal, irregular cavities interposed at convenient positions to alleviate friction; from their contents they may be either mucous or synovial; and vaginal synovial membranes, or sheaths, surrounding tendons and diminishing friction. Synovia is a transparent, viscid liquid, albuminous in its nature. The articulations consist of three (3) classes: diarthrosis, movable; synarthrosis, immovable; and amphiarthrosis, mixed. (104) ARTICULATIONS AND LIGAMENTS. I(j5 1. The diarthrosis are subdivided into: — Arthrodia, gliding joint, as superior tibiofibular; Enarthrosis, ball-and-socket joint, as shoulder and hip; Ginglymus, hinge-joint, as knee and ankle; Troehoides, a ring surrounding a pivot, as atloaxoid joint and superior radioulnar. Condyloid, elliptical cavity receiving an ovoid head, as wrist joint. Reciprocal reception, a concavoconvex articulation, as carpometa- carpal joint of thumb. 8. Synarthrosis, surface immovably connected by fibrous membrane without synovial membrane. They are divided into: — Sutura, bones interlocking with one another; Schindylesis, a fissure in one bone receiving a plate of bones, as between vomer and sphenoid; Gomphosis, a socket with a pivot inserted, as in alveolar cavities for teeth. Synchondrosis, a temporary joint in which the connecting medium is cartilage. The sutura may be either true, sutura vera, or false, sutura notha, the former having three divisions : dentata, tooth-like processes, as interparietal suture; serrata, saw-like edges, as interfrontal suture; limbosa, dentated processes and beveled margins, as frontoparietal ; the latter two divisions : — Squamosa, overlapping beveled margins, and Harmonia, by union of roughened surfaces, as intermaxillary suture. 3. AmpliiartJirosis, bony surfaces connected by fibroearti- lage, with or without synovial membrane, as between vertebral bodies, and pubic symphysis. MOTIONS IN Joints. — Joints admit of four (4) distinct varieties of motion: — (a) Gliding movement, between contiguous surfaces; (6) Angular movement, as flexion, extension, adduction and ah duction; [c) ( ircumduction, as in true enarthrosis; (d) Rotation on its own axis, as between the atlas and axis. Ti;.m I'Oito.M axi LLARY, or 1 em | M n'oiiuiiid ibular articulation (articulatio mandibidaris) , is formed by condyle of lower jaw, below articulating with glenoid cavity of the temporal bone and eminentia articularis above. It is a double arthrodial joint. Ligaments. — External Lateral (ligamentum t'emporomm- dibulare). Origin, tubercle on outer edge of zygoma ; insertion, outer Bide of neck of condyle. 106 " HUMAN ANATOMY. Internal Lateral (ligamentum sphenomandibulare) . — - Origin, spinous process of sphenoid; insertion, lower circumfer- ence of inferior dental foramen. Capsular (capsula articularis) . — Origin, circumference of articulation; insertion, neck of condyle. Interarticular fibrocartilage (discus articularis) is within cavity; external pterygoid muscle attached to its inner edge. Synovial membrane is divided into two by cartilage. Intermaxillary ligament passes from external pterygoid process to coronoid. Styloniaxillary or Stylomandibular Ligament (ligamentum stylo mandibular e) . — Origin, styloid process; insertion, inner surface of angle of jaw. Nerves are derived from the auriculo- temporal and masseteric branches of the inferior maxillary. Articulations of Vertebral Column. — These are formed between the contiguous surfaces of the vertebral bodies and articular processes, inclosed in capsular ligaments, and con- nected with the following ligaments : — The anterior common ligament (ligamentum longitudinale anterius) extends from the front of the body of the axis down the anterior surface (ventral) of the spine to the sacrum, being expanded opposite, and attached to each intervertebral connect- ing fibrocartilage; The posterior common ligament (ligamentum longitudinale posterius) descends along posterior surfaces (dorsal) of bodies from axis to sacrum within spinal canal. It also expands opposite and adheres to intervertebral substance; Intervertebral substance (fibrocartilagines intervertebrales) , disks of varying size, twenty-seven in number, composed of decussating fibers of fibrocartilage, arranged in crescentic laminge with central, semipulpy substance, separate the verte- bras; Ligamenta subflava (ligamenta flava, ligamenta ihtercru- ralia) descend in pairs, twenty-three on either side, from one lamina to another, from axis to sacrum, inclosing spinal canal; Supraspinous ligament (ligamentum supraspinal^) strong cord descending from one vertebra to another, from vertebra prominens (seventh cervical) to sacrum. Its continuation in cervical region forms ligamentum nucha; Interspinous (ligamenta inters pinalia) stretch between spinous processes throughout dorsal and lumbar regions; Intertransverse (ligamenta mtertransversaria) , between transverse processes in lower dorsal and lumbar regions. ARTICULATIONS AND LIGAMENTS. 107 The arteries are derived from ascending cervical and verte- bral in cervical, intercostals in dorsal, and lumbars in lumbar regions. The nerves are from spinal nerves in each region. Atloaxoid or atlaxtoaxoideax articulation (articula- tio atlantoepistrophica) consists of of four joints, the two lateral joints being arthrodia, and that between the odontoid process and arch of atlas and transverse ligament — a diarthrosis rota- toria being double, one in front, atlo-odontoid, and one behind, bdontotransverse. The ligaments are: — Anterior atloaxoid, or anterior atlantoaxial, two — super- ficial and deep, connecting anterior borders together; Posterior atloaxoid, or posterior atlantoaxial, connecting the arches of the atlas and axis posteriorly, and pierced laterally by the second spinal nerve; Capsular (eapmlce articulares) , two, connecting the articu- lating processes laterally, and lined with synovial membrane; Transverse, or cruciform (ligamentum transversum atlan- tis), arises from the tubercle on one side of the lateral mass, passes across the back of odontoid process to be inserted into the opposite tubercle. The arteries are from vertebral, nerves from second cervical, or from loop between it and suboccipital. Occipitoatlotd (articulatio atlanto-Oi ipitalis). — Anterior occipitoatloid (membrana atlanto-occipitalis anterior) — two, superficial and deep, connecting the tubercle and upper border of anterior arch or atlas with basilar process and margin of foramen magnum of occiput. Posterior occipitoatloid, or occipitoatlantal ligament (mem- brana atlanto-occipitalis posterior — membranous), connects the upper border of posterior arch of atlas with posterior margin of foramen magnum. Lateral ligaments, two bands connecting transverse process of atlas with jugular process of occipital. Capsular ligaments (capsules articulares) connect the artic- ular processes of atlas with condyles of occiput. The arteries are from rertebral; nerves, from suboccipital. OCCIPITOAXOID. — Occipitoaxoid is a broad hand, the con- tinuation of posterior common ligament, extending from the body of the axis to basilar groove of occipital. odontoid, or check (ligamentum alaria), pass upward and outward from the summit of tin,' odontoid process, to he inserted into the inner side of the occipital condyles. 108 HUMAN ANATOMY. Ligamentum suspensorium, or central occipito-odontoid {ligamentum apices dentis), arises from the center of the trans- verse ligament at the apex of the odontoid process, and passes upward, to be inserted into the anterior margin of the foramen magnum. Occipitocervical, or cervicobasilaris {membrana tectoria, apparatus ligamentosus colli, posterior occipitoaxoid ligament), arises from the bodies of the third cervical and axis, and is inserted into the basilar groove of occipital. The arteries are from vertebral; the nerves from suboc- cipital. Capsular Ligaments. — Throughout the spine the articula- tions between the facets of the articular processes are surrounded by capsular ligaments, making twenty-five pairs in all. Costovertebral Articulations {articulationes costoverte- brales). — The costovertebral have each a double arthrodia be- tween the head of the rib and the bodies of the two adjacent vertebrae, except the first, tenth, eleventh and twelfth ribs, which have each but a single joint. Anterior Costovertebral, or Stellate Ligament {ligamentum capituli costm radiatum). — Origin, head of the rib; insertion, into body of vertebra above and below, and interarticular carti- lage between. Interarticular costovertebral {ligamentum capituli costw interarticulare) is within the capsular ligament, from the crest on the head of the rib to the intervertebral substance. Capsular ligament {capsula articularis) surrounds the artic- ular surfaces. Costotransverse articulations are united by three ligaments — anterior, middle, and posterior costotransverse ligaments — extending between the tubercles of the ribs to the transverse process below. Arteries, the intercostals ; nerves, anterior branches of spinal. Costosternal Articulations (articulationes sternocos- tales). — These are connected by arthrodia excepting the first, which is a synarthrodia. Anterior chondrosternal ligaments {ligamentum sternocos- tal radiatum) connect the chondral and sternal surface in front ; Posterior chondrosternal {ligamentum sternocostal^ radia- tum) connect them behind. Capsular ligaments {capsula articularis) surround all, and synovial membranes are present in all but the first, while the second has an interarticular cartilage interposed. ARTICULATIONS AND LIGAMENTS. 1Q9 The chondroxiphoid (Ugamentum costoxiphoidea) ligament connects the cartilage of the seventh and sometimes the sixth rib with the xiphoid appendix. Interchondral and Costochondral Articulations (articula- tiones interchondrales) . — The ribs are connected with their car- tilages by the periosteum covering them. The cartilages of the sixth, seventh, and eighth ribs, and sometimes the fifth and ninth, are connected by external and internal interchondral ligaments (ligamenta intercostalia externa et interna), held together by capsular ligaments and lined with synovial mem- brane. Ligaments of Sternum. — The first and second portions of the sternum are articulated by an amphiarthrodial joint, sometimes a diarthrodial joint, connected by the external inter- sternal and internal intersternal ligaments, and lined with synovial membrane. The third portion, or enisform cartilage, is united to the gladiolus by a synarthrodia! joint. THE THORAX. The thorax is a conical osteocartilaginous framework, formed by the dorsal vertebrae behind, the ribs, intercostal mus- cles, and costal cartilages laterally, and the sternum in front. The apex, or superior opening (apertura thoracis superior), is bounded behind by the first dorsal vertebra, laterally by the first rib, and in front by the npper border of the sternum. It transmits the following structures: — Esophagus, trachea, thoracic duct (on the left side), lymphatic vessels of right side of chest and of surface of liver, innominate artery (right side), left common carotid artery, left subclavian artery, right and left superior intercostal arteries, right and left internal mammary arteries, thyroidea ima artery (if present), right and left inferior thy- roid veins, right and left innominate veins, right and left phrenic nerves, right and [eft pneumogastric nerves and their cardiac branches, left recurrent laryngeal nerve, right and left first dorsal nerves, right and left sympathetic nerves and their cardiac branches, apices of lungs and pleurae, remains of thymus, and deep cervical fascia passing to peri- cardium. The longus colli, Bternohyoides and sternothyroides on both Bides also pass I trough it. The inferior opening {apertura thoracis inferior) is formed by the last dorsal vertebra behind, the last rib laterally, and the ensiform cartilage in front. It Is filled in by the diaphragm. The cavity (carum thoracis) of the thorav contains the following structures: the heart and great vessels, trachea, hron- 110 HUMAN ANATOMY. chi and lungs, azygos and bronchial veins, internal mammary arteries, pneumogastric, phrenic and splanchnic nerves, thoracic duct, esophagus, lymphatic vessels and glands (vide medias- tinum). Spine and Pelvis. — Sacrovertebral articulation. In addi- tion to those of the spine, given above, there are two ligaments connecting the last lumbar vertebra with the sacrum : — Lumbosacral ligament, arising from the transverse process of the last lumbar vertebra, and inserted into the base of the sacrum ; Iliolumbar ligament (lig amentum Violumbale), arising from the apex of the transverse process of the last lumbar verte- bra, and inserted into the crest of the ilium. The arteries are from last lumbar, iliolumbar and lateral sacral. The nerves are branches of fourth and fifth lumbar and sympathetic. Sacroiliac articulation (articulatio sacroiliaca) , between the articular surfaces of the sacrum and ilium, is an amphiarthrodial joint connected by The anterior sacroiliac ligament (ligamenia sacroiliaca on- teriora), crossing between anterior surfaces of the two bones; Posterior sacroiliac ligament (ligamentum sacroiliacum posterius) , the stronger, passes in different directions between the posterior surfaces of the two bones, firmly binding them together. One of these, stronger than the others, passes from the posterior superior spinous process to the third sacral trans- verse process. It is called The oblique sacroiliac ligament (ligamentum sacroiliacum posterius longum). The arteries are from gluteal, sacral, spinal and iliolum- bar; the nerves from lumbosacral and posterior sacral cords. Sacroiscliiatic Articulation. — -The os innominatum and sa- crum are united by two important ligaments — the great sacro- sciatic (posterior) ligament,^ and the lesser sacrosciatic (anter rior) ligament. The great sacrosciatic ligament (ligamentum' sacrotubero- sum) arises from the fourth and fifth transverse tubercles of the sacrum, from spine of ilium, sacrum and coccyx, and passes downward, outward and forward, to be inserted into the inner margin of the tuberosity of the ischium, being prolonged for- ward as the falciform ligament (processus fahiformis) protect- ing the internal pudic veins and nerves. It converts the sacrosciatic notch into the lesser sacro- sciatic foramen, transmitting the obturator internus muscle and nerve, the internal pudic vessels and nerves. ARTICULATIONS AM) LKiA.M K.X'I'S. Ill The lesser sacrosciatic ligament (ligamentum sacrospino- sum) arises from the lateral margin of the sacrum and coccyx, and is inserted into the spine of the ischium. It converts the sacrosciatic notch into the greater sacrosciatic foramen (foramen i xhiadicum majus), transmitting the pyriformis muscle, gluteal vessels, superior gluteal nerve, sciatic vessels and nerves, and the internal pudic vessels and nerves. Sacrococcygeal articulation (symphysis sacrococcygea) is an amphiarthrodial joint, resembling the vertebral, having an interarticular fibrocartilage, and connected together by Pig. 61. 1, manubrium; 2, gladiolus; ?,, xiphoid appendix; 4, first dorsal vertebra; 5, last dorsal vertebra; 6, first rib. An anterior sacrococcygeal ligament (ligamentum sacra- coccygeum anterius), and A posterior sacrococcygeal ligament (ligamentum sacro- coccygeum posterius), continuations respectively of the anterior and posterior common ligament. Lateral sacrococcygeal, intertransverse (ligamentum sacro- coccygeum laterale), and intercornual ligaments connecting the rudimentary tran verse processes and the cornua together. The arteries are from lateral and median sacral: Tin- nerve branches of coccygeal, fifth sacral, and posterior ,. Sacra media. 7. Ovarian. 4. Iliolumbar. 8. Pubic branches from external .">. Brancb.ee of vasa intestini ten- epigastric arteries. uia. 9. Lymphatic vessels. Nerves. 1. Sympathetic :5. Obturator. 1. Branches from hypogastric 4. Lumbosacral cord. pleXUS. The cavity of the pelvis is bounded behind by the sacrum and coccyx, in' fr-mf by the symphysis, and laterally by the body of the ischium, forming a curved canal, widest in the middle. 8 H4 HUMAN ANATOMY. It contains, in the recent state, the organs of generation, rectum, bladder, and their vessels, nerves, ducts, etc. Its axis corre- sponds to the concavity of the sacrum and coccyx. Its depth in front, at the symphysis, is about one and a half inches in the middle axial line, three and a half behind, in males four and a half to possibly five and a half. The outlet of the pelvis (apertura pelvis inferior) is bounded by three prominences, the tuber ischii laterally and the coccyx behind ; separated by three notches, the subpubic arch in front and the sacrosciatic notches posteriorly. Its principal diameters are : transverse, three and a half inches in male, four and three-fourths in female; anteropos- terior, three and three-fourths in male, four and one-half in female. In the erect position the pelvis is placed at an angle of from sixty to sixty-five degrees with the ground. Differences of Sex. — The male pelvis is characterized by the strength of the bones, by prominent muscular impressions, by the depth and narrowness of the cavity, the large obturator foramen, and the acute angle of the subpubic arch. The female pelvis is characterized "liytlfe lightness of the bones, the slight muscular impressions, by the shallow, expanded iliac fossae, the increased size of the cavity in every diameter, and the obtuse angle of the subpubic arch. ARTICULATIONS OF THE UPPER EXTREMITY. The sternoclavicular articulation (articulatio sternodavicu- laris) is an arthrodial joint, formed between the sternal end of the clavicle and the sternum and first rib, being divided into two cavities by an interarticular fibrocartilage. Its ligaments are: — Anterior sternoclavicular (ligementum sternoclavicular ~e) is a broad band passing between the articular margins, attached, to the interarticular cartilage and two synovial membranes, and forming the front part of the capsular ligament; Posterior sternoclavicular passes over the posterior aspect of the joint and corresponds to the former ; Interclavicular {ligamentum inter claviculare) passes from the sternal end of the clavicle on one side across the suprasternal notch to the clavicle of the opposite side ; Costoclavicular, or rhomboid (ligamentum costo clavicu- lare), passes from the rhomboid impression on the inferior sur- ARTICULATIONS AM) LIGAMENTS. 115 face of the clavicle to the upper surface of the first costal cartilage. The synovial membranes form two sacs separated by the interarticular cartilage. The arteries are from muscular branch of suprascapular and contiguous muscular branches; The nerves, from descendens noni. Acromioclavicular articulation {articulatio acromio- clavieularis) is an arthroidal joint formed between the acromial extremity of the clavicle and the acromial process of the scapula. Its ligaments are : — 1, superor acromioclavicular ligaments; 2, eoracoclavicular liga- ments; 3, coracoacromial ligaments; 4, transverse ligament; 5, cap- sular ligament; 6, coracohumeral ligament; 7, tendon of biceps. The superior acromioclavicular (ligamentum acromio- claviciUare) connect the adjoining parts of the superior surfaces of the clavicle and acromion process; Inferior acromioclavicular, connecting the inferior surfaces; Coracoclavicular (lijamentum coracoclaviculare) consists of two parts : — Trapezoid ( ligamen I h m trapezoideum) } broad, square, arises from upper surface of coracoid process, and is inserted into oblique line on under surface of clavicle; and Conoid {ligamentum conoideum), behind and internal, arises from base of coracoid process, and is inserted into conoid tubercle on under side of clavicle and a line infernal to it. The ligaments surrounding the joint form a capsule. 116 HUMAN ANATOMY. The arteries are from anterior circumflex, acromial thora- cic and suprascapular. The nerves are anterior circumflex and suprascapular. The scapula has, also, The coracoacromial (ligamentum coracoacromiale) , arising from the outer border of the coracoid process and inserted into .the summit of the acromion, and Transverse (lig amentum transversum scapula superius), a ligamentous band bridging the suprascapular notch, converting it into a foramen for suprascapular nerve. The si-iouldee joint (articulatio humeri) is an enar- throdial joint, formed above by the glenoid cavity of the scapula and below by the head of the humerus. Its ligaments are : glenoid, coracohumeral and capsular. The glenoid (labrum glenoidale) surrounds the edge, deep- ens the glenoid cavity, and is continuous above with the long head of the biceps tendon. The capsular ligament (capsula articularis) , extensive and loose, arises above it from circumference of glenoid cavity be- hind the ligament, is attached below to the anatomical neck of humerus, and is pierced by tendons of two or three muscles. The coracohumeral, or accesory (ligamentum coracoJiu- merale), is a fibrous band which extends obliquely downward and outward from the coracoid process to the anterior part of great tuberosity, strengthening the capsular ligament. The synovial membrane is extensive and reflected upon the tendon of biceps, which is inclosed in a tubular sheath of syno- vial membrane (vagina mucosa intertubercularis) , and upon the infraspinatus, and subscapularis muscles. Its arteries are derived from the subscapular, suprascapular, dorsalis scapulas, anterior and posterior circumflex; Its nerves, from the subscapular, suprascapular and cir- cumflex. The 'elbow joint (articulatio cubiti) is a ginglymoid articulation formed above by the lower extremity of humerus, below by upper extremities of ulna and radius. Its ligaments are external and internal lateral, anterior and posterior liga- ments. External lateral (ligamentum collaterale radiale) arises from external condyle of humerus and is inserted into outer margin of ulna. Internal lateral (ligamentum collaterale ul- nare), much stronger, consists of two portions; anterior arises from fore part of internal condyle to be inserted into coronoid ARTICULATIONS AXD LIGAMENTS. Ill process, and posterior from back part of condyle to inner mar- gin of olecranon. Anterior ligament arises above coronoid fossa, and is inserted into coronoid process of ulna and orbicular ligament. Posterior ligament, attached above olecranon fossa, and below to olecranon process of ulna. The anterior and posterior ligaments become continuous with the lateral to encircle the joint. The arteries are derived from superior and inferior pro- FlG. 64. 1, anterior ligament: 2 and 2, anterior and posterior portions of internal lateral; 3, orbicular ligament; 4, oblique; 5, interosseous membrane. funda. anaatomotiea magna, posterior interosseous recurrent, anterior and posterior ulnar recurrent and radial recurrent. The nerves, from median, ulnar, musculocutaneous and musculospiral. Tilt; RADIOULNAR ARTICULATIONS (articula I i ianeous. Brachialis Anticus (to. brachialis) . — Origin, inner and outer surfaces of the shaft of the humerus, embracing the inser- tion of the deltoid; insertion, into the anterior surface of the coronoid process of the ulna; action, flexes the forearm; nerve, \ the musculocutaneous. Triceps — Extensor Cubiti (to. triceps bracliii). — Origin, r by three heads — middle, or scapular head (caput longum), below the glenoid cavity of the scapula; external head (caput laterale), from the posterior aspect of the shaft of the humerus, between the upper part of the musculospiral groove and the insertion of the teres minor, and from the external intermuscular septum ; the internal head (caput, mediale), from the posterior aspect of the shaft of the humerus, below the insertion of the teres major, and below the groove for the musculospiral nerve; insertion, by a common head, into the posterior part of the under surface of THE MUSCULAR SYSTEM. 14D the olecranon process of the ulna; action, extends the forearm; nerve, the musculospiral. Subanconeus (m. anconceus) . — Origin, from the posterior surface of the humerus, above the olecranon fossa; insertion, into the posterior ligament of the elbow-joint; action, draws up the posterior ligament of the elbow-joint during extension of the forearm; nerve, the muscnlospiral. Fig. 70. Muscles of shoulder and arm; 4, subscapulars; 5, teres major; G, coracobrachial is; 7, biceps. Fig. 71. Triceps muscle: 1, external head; 2, scapular head; 3, internal head; 4, insertion. MUSCLES OF THE FOREARM. Anterior Superficial Layer. Phonatob Radii Teres (w. /initiator tens). — Origin, by two heads — the smaller from the coronoid process of the ulna, the ulnar head {caput ulnare), the larger from the humerus, the internal condyle, the humeral head (caput numerate), and from the common tendon of this group of muscles, and intermuscular Beptum; insertion, into the outer aspeel of the 150 HUMAN ANATOMY. shaft of the radius; action, pronates the hand; nerve, the median. "Flexor Carpi Radialis. — Origin, by the common tendon from the inner condyle, and from the intermuscular septum between it and the pronator teres; insertion, into the base of the metacarpal bone of the index finger ; action, flexes the wrist ; nerve, the median. Fig. 72. Muscles of forearm: 1, biceps; 2, brachialis anticus; supinator longus; 5, extensor carpi radialis longior; 6, radialis brevior; 8, extensor communis digitorum; 12, ulnaris; 13, 14, extensors of thumb. 3, triceps ; 4, extensor carpi extensor carpi Palmaris Longus. — Origin, from the inner condyle of the humerus by the common tendon and the intermuscular septum and the deep fascia; insertion, into the annular ligament, spreading out in the palmar fascia; action, renders tense the palmar fascia; nerve, the median. Till: MUSCULAR SYSTEM. i;,l Fuexob Caepi Ulnabis. — Origin, by two heads — one by the common tendon from the inner condyle of the humerus, the other from the inner margin of the olecranon, the intermuscular septum, between the ulna and the flexor suhlimis digitorum, and from the upper two-thirds of the posterior aspect of the ulna; insertion^, pisiform hone; action, flexes the wrist; nerve, the ulnar. Flexor Sublimis Digitorum — Perforates (m. flexor digitorum sublimis). — Origin, by three heads — one from the coronoid process of the ulna, the ulnar head (caput vlnare), above the pronator radii teres, another from the common tendon from the internal condyle of the humerus, the humeral head (caput humerale), and the third, the radial head (caput radiale) from the oblique line of the radius, from the insertion of the pronator radii teres, to the tubercle; each tendon divides at the base of the first phalanges (chiasma tendinum) to allow the passage of the tendon of the flexor profundus digitorum; inser- tion, into the lateral margins of the second phalanges by four tendons; action, flexes the second phalanges; nerve, median. Anterior Veep Layer. Flexor Profundus Digitorum — Perforaxs (m. flexor digitorum profundus). — Origin, from the inner side of the coronoid process, and from the upper two-thirds of the front ami inner aspect of the shaft of the ulna, between the brachialis anticus above and the pronator quadratus below; insertion, by four tendons into the bases of the last phalanges, perforating the tendons of the flexor sublimis; action. Hexes the phalanges; nerves, the anterior interosseous and the ulnar. Plexob Longus Pollicis (»'. flexor pollicis longus). — Origin, from the upper two-thirds of the shaft of the radius and from tin' interosseous membrane; insertion, into the base of the last phalanx of the thumb; action, ilexes the phalanges; nerve, the anterior interosseous. Pronator Quadbatus. — Origin, anterior border of the ulna and from the oblique line of the lower fourth of the anterior aspeci of the ulna; insert inn , into the lower fourth of the front and outer border of the shaft of the radius; action, pronates the hand ; nerve, anterior interosseous. Radial Region, - pii ltob Lovers (in. Iivaeliinniiliiitis) . Origin, from the upper two-thirds of the external condyloid ridge of the 152 HUMAN ANATOMY. humerus and the external intermuscular septum; insertion, into the base of the styloid process of the radius; action, supinates the hand ; nerve, musculospiral. Extensor Carpi Eadialis Longior (m. extensor carpi radialis long us). — Origin, from the lower third of the external condyloid ridge of the humerus, from the common tendon of the extensor muscles of the forearm from the external condyle, and the intermuscular septum; insertion, into the radial side of the base of the metacarpal bone of the index finger; action, extends the wrist; nerve, musculospiral. Extensor Carpi Eadialis Brevior (m. extensor carpi radialis brevis). — Origin, from the common tendon from the external condyle of the humerus, the external lateral ligament, and the intermuscular septum; insertion, into the base of the metacarpal bone of the middle finger on its radial side; action, extends the wrist; nerve, posterior interosseous. Posterior Superficial Layer. Extensor Communis Digitorum (m. extensor communis digitorum) . — Origin, from the external condyle of the humerus, the deep fascia, and the intermuscular septa; insertion, by four tendons into the second and third phalanges of all the fingers (at the first phalanx they receive the insertion of the lum- bricales and interossei) ; action, extends the fingers; nerve, pos- terior interosseous. Extensor Minimi Digiti (m. extensor digiti quinti pro- prius). — Origin, from the common tendon from the external condyle and the intermuscular septum; insertion, into the sec- ond and third phalanges of the little finger ; action, extends the little finger; nerve, posterior interosseous. Extensor Carpi Ulnaris. — Origin, from the middle third of the posterior border of the ulna, from the common tendon from the external condyle of the humerus, and from the fascia of the forearm; insertion, into the base of the metacarpal bone of the little finger, on the ulnar side; action, extends the wrist; nerve, posterior interosseous. Anconeus (m. anconams). — Origin, from the outer con- dyle of the humerus; insertion, into the upper fourth of the posterior aspect of the shaft of the ulna and the side of the olecranon; action, extends the forearm; nerve, musculospiral. THE MUSCULAE SYSTEM. i;,;; Posterior Deep Layer. Supinator Eadii Brevis (///. supinator). — Origin, from the external lateral Ligament of the elbow-joint, from the exter- nal condyle of the humerus, from the ulna below the lesser sig- moid cavity, and from the orbicular ligament of the radius; insertion, into the neck, bicipital tuberosity and oblique line of radius; the posterior interosseous nerve pierces this muscle; n< Hun, supinates the hand; nerve, posterior interosseous. Extensor Ossis Metacaepi Pollicis (m. abductor pollicis longus). — Origin, middle third of posterior surface of radius, posterior surface of lower three-fourths of shaft or ulna, ami interosseous ligament; insertion, into base of metacarpal of thumb; action, extends metacarpal of thumb; nerve, posterior interosseous. Extensor Longus Pollicis (rn. extensor poll icis longus). — Origin, from the interosseous membrane and from the pos- terior aspect of the shaft of the ulna ; insertion, into the base of the last or terminal phalanx of the thumb; action, extends the thumb; nerve, posterior interosseous. Extensor Brevis Pollicis (m. extensor pollicis brevis).— Origin, from the posterior aspect of the shaft of the radius and from the interosseous membrane; insertion,, into the base of the first phalanx of the thumb ; action, extends the thumb ; nerve, posterior interosseous. Extensor Indicis (m. extensor indicis proprius). — Origin, from the posterior aspect of the ulna and from the interosseous membrane; insertion, into the second and third phalanges of the index linger, along with the tendon of the extensor com- munis: action, extends the index finger; nerve, posterior inter- osseous. MUSCLES OF THE HAND. These are divided into three groups — the radial region, the ulnar region, and the palmar region. Radial Region. Abductob Pollicis (/». abductor pollicis brevis). — Origin. from the annular ligamenl and ridge of the trapezium; inser- tion, into base of the first phalanx of the thumb on its radial side; action, abducts the thumb from the median line; nerve, median. 0PPONEN8 I'oi.l.n is. Origin, from annular ligament and palmar surf ace of the trapezium; insertion, into metacarpal bone 154 HUMAN ANATOMY. of the thumb throughout the whole length of its radial side; action, flexes the first metacarpal bone ; nerve, median. Flexok Brevis Pollicis (m. flexor pollicis brevis) . — This muscle consists of two portions., divided by tendon of flexor longus pollicis; origin, superficial portion from outer two-thirds of annular ligament and trapezium, deeper portion (by some called adductor obliquus pollicis) from sheath of flexor carpi radialis, the trapezoid, os magnum, and base of first, second, and third metacarpal bones ; insertion, into either side of base of first phalanx of thumb, the inner portion joining the adduc- tor and the outer portion the abductor; each tendon has a sesa- moid bone developed in it; action, adducts thumb toward median line; nerves, ulnar and median. Adductor Pollicis (Adductor Transversus Pollicis). — Origin, from lower two-thirds palmar surface of middle meta- carpal bone; insertion, into ulnar side of base of first thumb phalanx and internal sesamoid bone ; action, adducts thumb ; nerve, ulnar. Ulnar Region. Palmaris Brevis. — Origin, from palmar fascia and annu- lar ligament; insertion, into skin of palm of hand; action, wrinkles skin of hand; nerve, ulnar. Abductor Minimi Digiti (m. abductor cligiti quinti). — Origin, from pisiform bone and from tendon of flexor carpi ulnaris; insertion, into base of first phalanx of little finger on its ulnar side; action, abducts little finger from median line; nerve, ulnar. Flexor Brevis Minimi Digiti (m. flexor digiti quinti brevis). — Origin, from annular ligament and tip of unciform process of unciform bone; insertion, into the first phalanx of little finger; action, flexes little finger; nerve, ulnar. Opponents Minimi Digiti (m. opponens digiti quinti). — Origin, from annular ligament and from unciform process of unciform bone; insertion, into ulnar border of whole length of metacarpal bone of little finger; action, flexes little finger; nerve, ulnar. Palmar Region. Lumbricales. — Origin, by four fleshy tendons from the tendons of the deep flexors — the first and second from the pal- mar surface and radial side of the tendons of the index and middle fingers, the third from the adjoining sides of the ten- dons of- the middle and ring fingers, and the fourth from the THE MUSCULAR SYSTEM. 155 adjoining sides of the tendons of the ring and little fingers ; insertion, on dorsal aspect of each finger into the expansion of the extensor communis digitorum ; action, abduct the fingers to either side of the median line; nerves, ulnar and median. Ij&EROSSEI -muscles consist of two groups, the dorsal and palmar. Dorsal Inteeossei (mm. interossei dorsales). — Origin, from two heads from the adjacent sides of the metacarpal bones, four in number; insertion, the first into the radial side of the base of the first phalanx of the index finger, the second into the radial side of the middle finger, the third into the ulnar side of the middle finger, and the fourth into the ulnar side of the ring finger, the middle finger having two, one 011 either side; action, abduct the fingers from the median line; nerve, ulnar. Palmar Interossei (mm. interossei volares). — Three in number. Origin, the first from the entire length of the meta- carpal bone of the index finger on its ulnar side, the second from the ring linger on its radial side, and the third from the little finger on its radial side; insertion, into the base of the first phalanx and into the expansion of the extensor communis ten- don of the same finger from which they arise; action, adduct the fingers toward the median line; nerve, ulnar. Muscles of the Lower Extremity, iliac region. Psoas Magnus (m. psoas major). — Origin, from the sides and anterior surfaces of the transverse processes and bodies of the last dorsal and all the lumbar vertebrae and the interver- tebral substances between them; insertion, into the lesser tro- chanter of the femur, unting with the tendon of the iliacus; this muscle is in relation behind with the capsular ligament of the nip, being separated from it by a synovial bursa; act inn. and rotates the femur inward, and also flexes the trunk and pelvis on the thigh; nerves, anterior branches of the lum- bar nerves. Psoas Pabvus (m. psoas minor). — Origin, from the lateral Burfacee of the bodies of the Las! dorsal and first Lumbar vertebrae and from the intervertebral substances between them; insertion. into tin* iliopectinea] eminence, joining the iliac fascia; action . assists the psoas magnue and renders tense the iliac fascia; nerves, anterior branches of the lumbar. [liacus. Origin, from the base of the sacrum and the ilio lumbar Ligament behind, from, the iliac fossa and inner margin 156 HUMAN ANATOMY. of the crest of the ilium and the anterior superior and! ante- rior inferior spinous processes of the ilium; insertion, into the oblique or intertrochanteric line of the femur to the outer side of the insertion of the psoas; action, flexes and rotates the femur inward and flexes the trunk and pelvis on the thigh; nerves, anterior crural and the anterior branches of the lumbar. FEMORAL AND GLUTEAL REGION. Tensor Vaginae FEmoris, or tensor fasciae femoris (m. tensor vagina? femoris, or m. tensor fascia? femoris) . — Origin, from the anterior superior spinous process between the sartorius and the gluteus medius, and from the fore part of the outer lip of the crest of the ilium; insertion, into the fascia lata, about the upper fourth of the outer side of the thigh ; action, renders tense the fascia lata; nerve, superior gluteal. Sartorius. — Origin, from the anterior superior spinous process of the ilium and the upper part of the notch below; insertion, into the inner and upper part of the shaft of the tibia; action, flexes the leg upon the thigh and the thigh upon the pelvis; nerve, branches of the anterior crural. Quadriceps Extensor (to. quadriceps femoris). — This extensive muscle covers the front and sides of the femur and consists of four portions — the rectus femoris, the vastus exter- nus, the vastus interims and the crureus. Rectus Femoris. — Origin, by two heads — one, the long head, from the groove above the brim of the acetabulum; the other, the short head, from the anterior inferior spinous process of the ilium ; insertion, by the common tendon into the patella ; action, extends the leg upon the thigh ; nerves, branches of the anterior crural. Vastus Externus (m. vastus lateralis). — Forms the greater part of the quadriceps extensor. Origin, by an exten- sive aponeurosis extending from the tubercle of the femur along the anterior border of the great trochanter and the whole length of the outer lip of the linea aspera; insertion, into the outer portion of the patella, joining the common tendon; action, extends the leg on the thigh; nerves, branches of the anterior crural. Vastus Internus and Crureus (m. vastus medialis el m. vastus intermedins) .■ — Origin, by a tendinous aponeurosis ex- tending from the neck of the femur throughout the whole length of the inner lip of the linea aspera ; insertion, into the common tendon, together with the rectus femoris and the vastus exter- THE MUSCULAR SYSTEM. 157 mis; action, extends the leg on the thigh; nerves, branches of the anterior crural. Subcrureus (m. articularis genu). — Origin, from the an- terior aspect of the lower part of the shaft of the femur ; inser- tion, into the synovial sac behind the patella. Fig. 73. Anterior femoral re- gion ; 4, tensor vaginae femoris; 5, sartorius; 6, rectus; 7, vastus exter- nus; 8, vastus internus; 10, iliacus; 11, psoas; 12, pi i t ineus; 13, adductor longus; 14, adductor mag- nus; 16, gracilis. Fig. 74. Posterior femoral re- gion; 1, gluteus medius; 2, gluteus rnaximus; 3, vastus externus; 4, bi- ceps; 0, semitendinosus; 7, semimembranosus; s, gracilis. GRACILIS. — Origin , from the inner margin of the ramus ->l the ischium and the pubes ; insertion, into the innei asped "I' the -haft <»f the tibia, below the tuberosity; action, flexes the 158 HUMAN ANATOMY. leg and draws it inward, assisting the sartorius; nerve, the obturator. Pectineus. — Origin, from the tendinous prolongation of Gimbernat's ligament and the linea iliopectinea ; insertion, into the rough line leading to the linea aspera from the lesser tro- chanter ; action, adducts the thigh ; nerves, obturator, the acces- sory obturator, and branches of the anterior crural. Adductor Longus. — Origin, from the front of the os pubis below the crest; insertion, into the middle third of the linea aspera, between the adductor magnus and the vastus internus ; action, adducts the thigh; nerve, obturator. Adductor Brevis. — Origin, from the outer aspect of the descending ramus of the pubes, between obturator externus and the gracilis; insertion, into the upper part of the linea aspera; action, adducts the thigh; nerves, branches of the obturator. Adductor Magnus. — Origin, from the tuberosity of the ischium, the ascending ramus of the ischium, and from the descending ramus of the pubes; insertion, into the rough line leading to the linea aspera from the great trochanter to the inner side of the gluteus maximus and into the adductor tuber- cle above the inner condyle of the femur {vide page 64) ; action, adducts the thigh; nerves, the obturator and a branch from the great sciatic. Gluteus Maximus (m. glutceus maximus). — Origin, from the superior curved line of the ilium, the posterior aspect of the last segment of the sacrum, the border of the coccyx, and the surface of the great sacrosciatic and posterior sacroiliac liga- ments ; insertion, into' the rough line leading to the linea aspera from the great trochanter, between the adductor magnus and the vastus externus, and into the fascia lata; action, it is a ten- sor of the fascia lata, and .an external rotator and extensor of the thigh; nerves, the inferior gluteal and a branch from the sacral plexus. Gluteus Medius (m. glutceus medius). — Origin, from the outer lip of the crest and the outer aspect of the ilium, between the middle and superior curved lines, and from the gluteal apo- neurosis; insertion, into the oblique line on the outer surface of the great trochanter; action, rotates the thigh outward; nerve, from the superior gluteal. Gluteus Minimus (m. glutceus minimus). — Origin, from the border of the great sacrosciatic notch and between the infe- rior and middle curved lines of the outer aspect of the ilium; insertion, into a depression on the front border of the great THE MUSCULAR SYSTEM. i.v.i trochanter; action, rotates the thigh inward; nerve, superior gluteal. PYEIFORMIS (in. piriformis). — Origin, from the anterior surface of the sacrum, between the first to the fourth anterior sacral foramen, and from the margin of the great sacrosciatic foramen, and from the anterior aspect of the great sacrosciatic ligament; insertion , into the upper border of the great tro- chanter, with the tendon of the obturator interims; action. rotates femur outward; nerve, obturator. Obturator Intbenus. — Origin, from the inner side of the obturator foramen and the internal surface of obturator mem- brane and the anterior and external wall of the pelvis; it passes out of the pelvis through the lesser sacrosciatic notch and receives the tendons of the gemelli muscles; insertion, into the upper border of the great trochanter in front of the pyriformis; action, rotates the thigh outward; nerves, branches from the sacral plexus. Gemelli consist of two muscles — gemellus superior and gemellus inferior. G-emellus Superior. — Origin, from the outer aspect of the spine of the ischium; insertion, into the tendon of the obturator interims. Gemellus Inferior. — Origin, from the outer border of tuberosity of the ischium; insertion, into the lower portion of the tendon of the obturator interims ; action, rotates the thigh outward; nerves, branches of the sacral plexus. Quadrattjs Femoris. — Origin, from the external border of the tuberosity of the ischium; insertion, into the upper part of the linea quadrati, on the posterior aspect of the trochanter major; action, rotates the thigh outward; nerves, branches of the sacral plexus. Obturator Exterxus. — Origin, from the inner two-thirds of the anterior aspect of the obturator membrane and the ten- dinons arch covering the canal for the obturator vessels and nerves, and the descending ramus of the pubis and the ascend- ing ramus of the ischium; insertion, into the digital fossa of the femur; action, rotates the thigh outward; nerve, the obturator. Bn EPS, or biceps flexoT cruris(wi. l>i<<-jis femoris). — Origin, arises by two heads — the long head [caput longum), from the lower and inner facet of the tuberosity of the ischium by a '•oiiiiiion tendon to the long head of the seinitendinostis ; the short bead (caput breve), from the outer lip of the linea aspera, tween the vastus externue ami the adductor magnus; insertion, outer ?-id lumbocostalis lateralis). The ligamentum arcuatum in- ternum (areas lumbocostalis medialis) extends from the body of the first or second lumbar vertebra to the transverse process, inclosing the psoas magnus muscle. Abdominal and Pelvic Fascia. — The transversalis fascia is a thin aponeurosis between the peritoneum and transversalis THE MUSCULAR SYSTEM. 167 muscle. It is attached below to the pubes, pectineal line, Pou- partfs Ligament and the femoral vessels, and becomes continu- ous with the iliac and pelvic fascia. The pelvic fascia (fascia pelvis) lines the pelvic cavity throughout, attached to the sym- physis pubis, margin of obturator foramen, and the sacrum and becomes continuous with the iliac and transversalis fascias. [Below it forms the arcus tendinece, for the origin of muscles and the division into the rectovesical and obturator fascias (vide Perineum). The iliac fascia (fascia iliaca) is a thin, fibrous mem- brane investing the psoas and iliacus muscles. Above it is attached to the Ligamentum arcuatum internum, laterally, with the bodies of the vertebras and sacrum, and below it is attached to Poupart's ligament, the femoral vessels, pectineal eminence, and the capsule of the hip-joint. Upper Extremity. — The superficial fascia is a thin mem- brane, inclosing the parts throughout, and adherent to the deep fascia beneath. The deep fascia is very dense and strong. It gives off numerous intermuscular septa, and is attached to the olecranon and back part of ulna, and becomes continuous below with the anterior and posterior annular ligaments of the wrist-joint, both of which it forms. The anterior annular ligament (ligamentum carpi trans- versum) is attached to the ridge of the trapezium, the tuberosity of the scaphoid, the unciform process of the unciform, and the pisiform hone, forming a strong, iibrous arch, under which the flexor tendons of the lingers pass. The posterior annular ligament (ligamentum- carpi dorsale) is attached to the radius, ulna, cuneiform and pisiform hones, and the palmar fascia, forming a fibrous arch for the passage of the extensors of the fingers. The palmar fascia (aponeurosis palmaris) is an exceedingly dense, triangular mass of adipose and fibrous tissue, attached behind to the anterior annular lip 'lit, and in front divides into tour fasiculi, each of which subdivides into two, inserted into the lateral ligaments of the metacarpophalangeal articula- tion-. These digitatione allow the passage of the flexor digi- toruni tendons, the digital vessels and nerves. It is attached above to the skin, and laterally gives off vertical septa, which separate the middle from the lateral groups of palmar muscles. 168 HUMAN ANATOMY. EASCTAS OF THE LOWER EXTREMITY. The superficial fascia- resembles that in other localities, except that, in the front of the thigh, it consists of two or more layers between which are found the superficial vessels, nerves and lymphatic glands. The under layer is attached to the margins of the saphenous opening, where it is perforated by numerous lymphatic vessels and blood-vessels ; hence its name, cribriform fascia {fascia cribrosa). The deep fascia of the thigh, or fascia lata, is a dense fibrous aponeurosis attached to the pubes, Poupart's ligament, crest of the ilium, sacrum and coccyx, inclosing the gluteus maximus and tensor vaginae f emoris between its layers, attached to the linea aspera, and below to all the bony points about the knee joint, being attached to the head of the fibula and tuber- osities of the tibia by the iliotibial band (trnctus ilioiibialis [M aissiati] ) . The saphenous opening (fossa ovalis) is a large opening in the fascia lata at its upper and inner part closed by the crib- riform fascia. Through it pass the internal or long saphenous vein, the superficial epigastric and external pudic arteries, and lymphatics. It is formed by the iliac and pubic portions of the fascia lata. The iliac portion becomes continuous at the pec- tineal line with Gimbernat's ligament, and its free border forms the falciform border (vide Femoral Hernia). The pubic portion is continuous behind the femoral vessels with the femoral sheath and the iliac and psoas fascia. At the lower border of the saphenous opening it is continuous with the iliac portion of the fascia lata. The deep fascia of the leg (fascia cruris) is attached above to the bony points about the knee, continuous with the fascia lata, invests the leg completely, except the inner surface of the tibia, sends in a deep transverse fascia between the superficial and deep muscles on the posterior aspect of the leg, and below becomes continuous with the annular ligament of the ankle- joint. The annular ligament of the ankle joint consists of three portions — the internal, external and anterior. It is attached to the bony points about the ankle joint, the external and inter- nal malleoli, surfaces of the os calcis, and allows the passage of the muscles to the foot. The dorsal aponeurosis of the foot (fascia dorsalis pedis) covers the back of the foot, uniting with the plantar fascia at THE MUSCULAR SYSTEM. 1G9 the sides, and terminating anteriorly at the heads of the meta- tarsal hones. The plantar aponeurosis (aponeurosis plantaris) consists of three portions, a central and two lateral portions. The middle portion commences at the inner tubercle of the os calcis and proceeds, gradually becoming broader, to the heads of the metatarsal bones, where it divides into four branches, each of which nearly surrounds the corresponding flexor tendon to which it gives passage, and is inserted into the edges of the dorsal expansion of the first phalanges. The inner portion arises from the internal annular liga- ment, covers the abductor pollicis muscle, and joins the dorsal aponeurosis internally and the internal septum externally. The outer portion, much stronger, arises from the os calcis, covers the abductor minimi digiti muscle, and joins the external plantar septum internally and dorsal aponeurosis externally, and is firmly attached to the base of the fifth metatarsal bone. Numerous septa pass from the upper surface of the plantar fascia between the plantar muscles and tendons to be inserted into the metatarsal and tarsal bones. THE HEART AND VASCULAR SYSTEM. The pericaedium is a conical, membranous, closed sac, surrounding the heart and the roots of the great vessels. It occupies the greater part of the middle mediastinum, its apex upward, its base downward and attached to the central tendon of the diaphragm. It is a fibroserous membrane, consisting of an outer fibrous coat and an inner serous coat. The latter is composed of a visceral layer (epicardvum) reflected over the heart and vessels, and a parietal layer lining the inner surface of the fibrous sac. It secretes a thin, serous fluid. The fibrous coat becomes continuous above with the deep layer of the cervical fascia, being prolonged upward on the outer surfaces of all the great vessels, except the inferior vena cava. The Heart (Cor). The heart is a hollow, muscular organ, placed obliquely within the chest, and inclosed within the pericardium. Its base (basis cordis) is directed upward and backward, opposite the fifth to eighth dorsal vertebrae; its apex (apex cordis) down- ward to the left, opposite the interspace between the fifth and sixth costal cartilages. Its weight is about ten to twelve ounces in male, eight to ten in female ;, length five inches, breadth three and a half inches, thickness two and a half inches. It is divided into four parts : — Right auricle, Left auricle, Right ventricle, Left ventricle, an auricle and ventricle on each side. This division is indicated on both surfaces of the organ by two grooves crossing each other and named, from their position, the longitudinal or the inter- ventricular (sulci longitudinalis) , the transverse or the auric- uloventricular (sulcus coronarius) grooves. Eight Auricle. — The right auricle (atrium dextrum) consists of two portions — a sinus or atrium and an appendix auriculce, which projects to the right side of the origin of the aorta. The walls at the fore and outer part and in the auricu- lar appendage are thickened by musculi pectinati. The mus- (170) THE HEART AND VASCULAR SYSTEM. 171 cular tissue of the auricles is connected with that of the ventricles by a small band of fibers, the auriculoventricular fasciculus, or "bundle of His." This smooth ridge is known inside the auricle as the crista terminate of His, and outside the groove is called the sulcus terminalis of His. Its interior presents the following parts:— Openings of the superior and inferior vena' cava', neither of which are protected by valves : Opening of coronary sinus {sinus coronarius), protected by a valve of two unequal segments— the coronary valve; Fig. 77. Right side of heart: a, apex; 6, right ventricle; d, pulmonary artery; e, f, chords tendineas; p, aorta; 3, superior vena cava; 5, inferior vena cava. Foramina Thebesii (foramina vcnarum minimarum), mi- nute foramina returning the blood from the heart-muscle; stachiap valve (valvula rcmr n,r,r info-ions [Eu- rtachW]), the remainfi of a fetal structure, extending from the right Of the orifice Of the inferior vena cava to the outer border of the oval foramen ; Fossa oralis, the obliterated foramen ovale of fetal lite: AnnuTui ovalis {limbus fossm oralis | VieussenU]) , the oval margin of the preceding structure; Tubercle of Lower (tuberculwn intervenosum \Lowen\),Q small rudimental projection on the right wall, directing the blood toward the annciilo\eiit ri.ular opening; 172 HUMAN ANATOMY. Musculi pectinati, elevated muscular columns before re- ferred to; Auriculoventricular opening (ostium venosum dextrum), an oval aperture about one inch in diameter, surrounded by a fibrous ring, and protected by the tricuspid valve. The Left Auricle (atrium sinistrum). — The left auricle is smaller, and its walls somewhat thicker than the right, and consists of a cuboidal sinus and an elongated appendix auriculae. Its interior surface is smooth, except the auricular appendage, which is provided with musculi pectinati, and presents the fol- lowing openings : — Pulmonary veins, four in number, two on either side; they are without valves; Auriculoventricular opening (ostium venosum ventriculi sinistri), an oval aperture, rather smaller than the right, and protected by the bicuspid or mitral valve. The Eight Ventricle (ventriculus dexter). — The right ventricle is triangular, with thick walls (one-quarter inch), and occupies the anterior part of the organ. Its capacity is about three fluidounces. It presents the following: — Infundibulum (conus arteriosus), a conical pouch, from which the pulmonary artery arises ; Columnse carneae (trabecular carnece), muscular columns projecting from the inner surface, three or four of which (mus- culi papillares) give attachment to the chordae tendineae; Chordce tendinew, delicate tendinous cords, connecting the margins and central portions of the tricuspid valve with the columns carneae; Tricuspid valve (valvula tricuspidalis) , consists of three segments or cusps (cuspides) which consist of triangular dupli- cations of endocardium, strengthened by fibrous tissue; they are attached by their bases to the auriculoventricular orifice (an- nulus fibrosis dexter) and their free margins give attachment to the chordae tendineae (ostium venosum ventriculi dextri). Semilunar valves (valvular sem.ilunares a. pvlmonalis) con- sist of three semicircular folds, guarding the orifice of the pul- monary artery; their free margins are thickened by nodules, the corpora Arantii (nodulus valvules semilunaris [Arantii~\) ; Opening of the pulmonary artery (ostium- arteriosum) , at the apex of the conus arteriosus, is circular in outline and pro- tected by the semilunar valves; Sinuses of Valsalva^ three pouches, situated one behind each semilunar valve. THE HEART AND VASCULAR SYSTEM. 173 The Left Ventricle (ventricutus sinister). — The left ventricle is longer, more conical, and its walls three times as thick a< the right. It forms the apex {apex cordis) and most of the posterior and inferior (facies diaphragmatica) portion of the heart. It presents the following parts in its interior :— Col urn iter carnece, and Chorda tendinece, much the same as the right; _ Uiriculoventricular opening {ostium venosum ventnculi sinistri), a little smaller than the corresponding orifice on the right side, similarly formed, and protected by the mitral valve; ' The mitral valve {valvula bicusptdalis) consists of two unequal segments, attached the same as the tricuspid, and also affording attachment to the chordae tendmese; Aortic opening {ostium arteriosum), a circular opening to the right of the auriculoventricular opening; The semilunar valves {valvule semilunares aorta'), three in number, surround the aorta, and are larger and stronger than those on the right side ; . Sinus aortici (sinuses of Valsalva) are depressions situated behind each valve. . „ Structure of the Heart. — The heart is composed 01 finely striated muscular walls, with external serous covering from the pericardium and internal serous lining the endo- cardium the latter continuous with that of blood vessels. The endocardium is a thin, translucent membrane, consisting ot endothelium and a fine basement membrane, beneath which is a fibrocystic layer. The doublings of these layers constitute the valve< The tricuspid and mitral valves, as well as the aortic and pulmonary, are surrounded by a fibroelastic ring {annulus fibrosa), which furnishes a "puctum fixuur for the varying musculature of the heart. . . The muscular structure consists of an intricate interlace- ment of fibrous Lands. Of these there are two groups— those of the auricles and those of the ventricles. The former consist of a superficial transverse set and the internal or deep set, of which there are the looped and the circular. The fibers of the ventricles consist of seven layers m a gen- era] oblique and circular manner, ami terminating in a whorl ,„. ,.,„•/,,,. a1 the apex, some of the abers fern, mating m the oolumnse carnese, musculi pectinati, while others ascend, torm- ina i„ their course a twisted loop like the figure s. The arteries of the Lean are i he ri-ht and left coronarv. The veins are the anterior or great, middle <>r posterior cardiac, the left auricular, 174 HUMAN ANATOMY. the right auricular and venge Thebesii. The lymphatics end in thoracic and right lymphatic ducts. The nerves are derived from the cardiac plexus of pneumogastric, spinal and great sympathetic. The Vascular System. The vascular system consists of four sets of vessels — ar- teries, capillaries, veins and lymphatics. Anastomoses are common. The vessels that convey blood to and from the tissues of the body generally constitute the general system. Those that carry blood to and from the lungs form the pulmonary system. The vessels passing to the liver form the portal system. The arteries, for the most part, are composed of three coats : — Tunica adventitia, or external coat, consists of fibrous tis- sue, thinnest on the largest trunks, and disappears in those which merge into capillaries. The tunica media,, or middle coat, is thickest in the large trunks, and consists mainly of elastic tissue, together with some unstriated muscle and some connective tissue; in smallest arter- ies it consists alone of muscular tissue. The tunica intima, or internal coat, thinnest, most trans- parent and elastic, is composed of lining endothelium, basement membrane, and layers of elastic tissue. The elastic tissue in layers forms nets, constituting so-called perforated or fenestrated membrane. It becomes continuous with capillaries. The walls of larger arteries are supplied by blood vessels from neighboring arteries. Nutrient arteries form a network of capillaries, and returning veins empty into contiguous veins. The nerves are both medullated and non-medullated and chiefly pass to the muscle-fibers. Arteries dilate and contract with the action of the heart, constituting the ptdse. The veins. — Larger, more numerous, and more capacious than the arteries, with one exception (the portal vein) they convey blood to the heart. The large veins emptying into the heart are : — Four pulmonary ; Superior and inferior vense cavse; Coronary veins from walls of the heart through the coronary sinus. THE HEART AND VASCULAR SYSTEM. 175 The larger arteries have, each, one companion vein, the medium arteries two. In some positions they form venous plexuses, corresponding to arterial anastomoses. Many large veins have valves, usually in pairs, attached by convex border. There are, however, no valves in the pulmonary, superior and inferior cavae, azygos, portal vein and branches, hepatic, renal, uterine, and spinal, and most of those of head or neck. In certain membranes and organs channels exist, lined by internal coat of blood vessel, termed venous sinuses, as in the dura mater, bones and uterus. The coats of the veins are similar to those of the arteries. and differ chiefly in the weakness of the middle or muscular coat, which allows them to remain open. The external fibrous coat has also longitudinal, unstnated muscular tissue in veins of abdominal cavity: — The middle, unstriated muscular and fibrous tissue; The internal, no fenestrated membrane. The walls have nutritive vessels, and are well supplied with non-medullated and medullated nerves, which pass to the mus- cular coat. _ , Both arteries and veins are insensitive m healthy Tin: capillaries communicate with the terminations of the arteries and veins, and do not communicate with tissue proper which is supplied by imbibition. They vary in size in different organs from y2ooo to Vsooo inch in diameter, but all permit the passage of blood corpuscles. Their form is depend- ent on the form of tissue supplied. Composition of the Blood. — Blood within the vessels is a perfect fluid, alkaline reaction, saline taste, average specific gravity 1060, and of feeble, peculiar odor. Within the arterial jselfl it is bright red in color, becoming in the veins (partic- ularly the venae portae and pulmonary arteries) a deep maroon or reddish-black color. It constitutes about Vi - or 1/u of the body-weight, an. I con- sists <-f the Liquor sanguinis and the corpuscles, red, white, etc. The red corpuscles or erythrocytes are elastic, specific gravity L088, Don-nucleated, and average about Hooo "M "' diameter. A cubic millimeter of blood contains, m the male. 5,000,000 red blood-cells, and, in the female, 4,500,000. On evaporation they become stellate, and swell up with water. They vary much in size and Bhape in differenl animals, and contain hemoglobin, which carries oxygen to the tissues an. I C02 to the bin. Clot. 176 HUMAN ANATOMY. The colorless corpuscles or leucocytes are free nucleated protoplasmic masses, capable of ameboid movement. They con- tain fat-corpuscles, myosin, cholesterin, protagon, glycogen and nuclein. There is about 1 white corpuscle to every 600 red corpuscles, or in 1 cubic millimeter of blood there are present about from 5000 to 10,000 white corpuscles, an average of 7500. According to Ehrlich and others, by means of a "tri-acid stain," five varieties of colorless cells may be distinguished as fol- lows : — 1. Small lymphocytes, 2. Large lymphocytes or mononuclear leucocytes, 3. Transitional leucocytes, 4. Polymorphonuclear leucocytes, and 5. Eosinophiles. The liquor sanguinis is a pale, amber-colored fluid, holding the corpuscles in suspension, and very prone to coagulate. The coagulation of the blood results in the formation of a clot, or crassamentum, and the serum, in the following manner : — C Cornuscles / White corPuscles corpuscles, j Red corpuacle8- Fluid blood J ,,, I P1fsma°r 1 Fibrin. ! eLl(luor \ Serum. J-Q i Sanguinis J berum. THE CIRCULATION OP THE BLOOD. Fetal Circulation. — The fresh arterial blood returning from the placenta through the umbilical vein enters the fetus at the umbilicus, passes along the suspensory ligament to the under surface of the liver, where a portion passes directly into the inferior vena cava by the junction of the ductus venosus with the left hepatic vein, a portion enters the liver and reaches the inferior vena cava through the hepatic veins, and the larger por- tion mixes with the portal venous blood before reaching the vena cava through the hepatic veins. It ascends in the inferior vena cava along with the venous blood from the trunk and lower extremities to enter the right auricle of the heart (Fig. 78). Here it is directed by the Eustachian valve through the foramen ovale into the left auricle, into the left ventricle, and so through the aorta, chiefly to the head and upper extremities, a small por- tion only entering the descending aorta. It is returned by the superior vena cava to the right auricle, passes over the Eu- stachian valve to the right ventricle, and so through the pulmo- nary artery toward the lungs, but (the lungs being almost THE HEART AND VASCULAR SYSTEM. 177 Fig. 78. Diagram of the fetal circulation: 1, umbilical cord, with veins and arteries; 2, placenta; :', divisions of umbilical vein; 4, hepatic branches; 5, ductus venosus; 6. inferior vena cava; 7, portal vein; 8, right auricle; 9, left auricle; 10, left ventricle; 11, arch of aorta; 14, superior vena cava; 15, right ventricle; 10, pulmonary artery; 17, ductus arteriosus; 18, descending aorta; 19, umbilical arteries. 12 178 HUMAN ANATOMY. impervious) most of it passes through the ductus arteriosus into the descending aorta (mixing with the small quantity from the left ventricle before mentioned) to supply the lower extremities, abdominal and pelvic viscera, and as venous blood to return through the umbilical arteries to the placenta. Circulation After Birth and in Adult. — The dark venous blood from the entire body is received through the superior and inferior venae cavse and coronary sinus into the right auricle, whence it passes into the right ventricle, to be sent through the pulmonary artery into the lungs. Here it becomes oxygen- ated, and as bright arterial blood it returns to the left auricle by the pulmonary veins, and passes into the left ventricle, whence it is distributed through the aorta and its branches to the entire body. AORTA (ARTERIA MAGNA ) . This is the main trunk of the systemic arteries. Com- mencing at the upper part of the left ventricle, it ascends for a short distance, arches backward over the right pulmonary artery, the root of the left lung, to the left side of the body of the fourth dorsal vertebra, where it passes downward through the diaphragm and becomes the abdominal aorta. It then descends to the lower border of the fourth lumbar vertebra, where it terminates by dividing into the left and right common iliac arteries. In its course it is divided into the arch, the thoracic aorta, and the abdominal aorta, the arch being sub- divided into the ascending, transverse and descending portion. Relations. — The ascending portion of the arch is in relation on the right side with the superior cava and right auricle; on the left side with the pulmonary artery; in front, with the pulmonary artery, the pericardium, the right appendix auriculae and the thymus gland; and behind, with the root of the right lung and the right pulmonary vessels. The transverse portion of the arch is in relation in front with the left lung and pleura, the left pneumogastric and phrenic nerves, the superficial cardiac nerves, the left superior intercostal vein, and the thymus gland; behind, with the trachea, the esophagus, the thoracic duct, the deep cardiac plexus, and the left recurrent nerve; above, with the arteria innominata, the left innominate vein, the left subclavian, and the left carotid; and below, with the left bronchus, the bifurcation of the pulmonary artery, the left recurrent nerve, and the remains of the ductus arteriosus. The descending portion of the arch is in relation in front with the root of the left lung and the pleura ; behind, with the left side of the body of the fifth dorsal vertebra; on the right side with the thoracic duct and esophagus; and on the left side with the pleura. THE HEART AND VASCULAE SYSTEM. 179 The branches of the aorta are, from the arch: — Two coronary, Left common carotid. Innominate, Left subclavian. The coronary arteries supply the heart and are two in number — the right and the left (a. coronaria cordis dextra et sinistra). The right arises from the aorta, above the right semi- lunar valve, and passes downward in a groove between the right auricle and ventricle on its posterior surface, dividing into two branches, one of which anastomoses with the left coronary. The left coronary arises above the left semilunar valve and descends in the anterior interventricular groove, where it divides into two branches, one of which anastomoses with the right coronary. Arteria Innominata (a. anonyma). — This is the largest branch from the arch of the aorta. It ascends to the upper border of the right sternoclavicular articulation, where it divides, into the right common carotid and the right subclavian arteries. It is a short vessel, from one and a half to two inches long. Relations. — It is in relation in front with the sternum, the sterno- hyoid and sternothyroid muscles, the left innominate and right inferior thyroid veins, the thymus gland, and the cardiac branch from the right pneumogastric ; behind, with the trachea; on the right side, with the pleura and right vena innominata and right pneumogastric nerve; on the left side, with the left carotid and the thymus gland. The Common Carotid Arteeies (a. carotis communis) . — The left common carotid, arising directly from the arch of the aorta, is longer and ascends more obliquely and is more deeply placed than the right. From the sternoclavicular articulation to their division at the upper border of the thyroid cartilage the common carotid on either side pursues the same course. At the lower part of the neck the two vessels are separated by a very short interval, which is occupied by the trachea, but at the upper part they diverge widely; the larynx, pharynx, thyroid body, and several muscles being interposed between them. It ascends the neck in a direction indicated by a line drawn from the sternoclavicular articulation to midway between the mastoid process and the angle of the lower jaw. Relations. —It is in relation in front with the sternomastoid, sternothyroid, sternohyoid and omohyoid scles, the anterior jugular and the superior ami middle thyroid veins, the sternohyoid artery, and the descendens and communicans noni nerves; behind ii rests upon the longus colli and rectus e;i|>iii> anticus major muscles and is in relation with the sympathetic nerve, the recurrent laryngeal nerve, and the inferior thyroid artery; on the Outer side, with the pneumogastric and 180 HUMAN ANATOMY. the internal jugular vein; to the inner side, the larynx, trachea, pharynx, thyroid gland, inferior thyroid artery and the recurrent laryngeal nerve. THE EXTERNAL CAROTID ARTERY (A. CAROTIS EXTERNA). From its commencement at the superior border of the thy- roid cartilage it passes upward and forward and then backward, beneath the anterior margin of the sternomastoid muscle, in a direction indicated by an imaginary line drawn from the supe- rior border of the thyroid cartilage to a point midway between the meatus and the neck of the condyle of the lower jaw. It ascends through the space known as the superior triangle of the neck. Relations. — It is in relation in front with the parotid gland and the facial nerve and tempo romaxillary vein passing through it, covered by the skin, superficial fascia, platysma, and deep fascia, and is crossed by the digastric and stylohyoid muscles, hypoglossal nerve, lingual and facial veins; behind it is separated from the internal carotid by the parotid gland, styloglossus and stylopharyngeus muscles, and the glosso- pharyngeal nerve, and at its lower part the superior laryngeal nerve ascends behind it; to its inner side it is in relation with the pharynx, hyoid bone, superior laryngeal nerve, and the ramus of the lower jaw, from which it is separated by the parotid gland. The branches of the external carotid are eight: — 1. Superior thyroid, 5. Posterior auricular, 2. Lingual, 6. Ascending pharyngeal, 3. Facial, 7. Temporal, 4. Occipital, 8. Internal maxillary. 1. Superior thyroid (a. thyreoidea superior) arises below the great cornu of the hyoid bone, passes beneath the sterno- hyoid, omohyoid and sternothyroid muscles to the upper part of the thyroid gland, where its terminal branches anastomose with its fellow of the opposite side. Its branches are : — (a) Hyoid (ramus hyoideus) , to the lower border of the hyoid bone ; (b) Superficial descending branch {ramus sternocleidomastoideus) , crosses the common carotid artery to supply the skin and sternomas- toid muscle; (c) Superior laryngeal (a. laryngea superior), passes beneath the thyrohyoid muscle with the superior laryngeal nerve; (d) Cricothyroid (ramus cricothyreoideus) , crosses the crico- thyroid membrane. 2. The lingual artery (a. lin gualis) arises from the ante- rior part of the external carotid, between the superior thyroid and the facial. It passes inward and upward to the greater THE HEART AND VASCULAR SYSTEM. 181 cornu of the hyoid bone, and, descending, reaches the under surface of the tongue and runs along its under surface as far as its tip, where it terminates as the ranine. In its course it passes through a triangle hounded on two sides below by the two bellies of the digastric, and above by the hypoglossal nerve, passing under the hyoglossus muscle. Its branches are : — Hyoid (ramus hyoideus) , to the hyoid bone; Dorsalis linguae (ramus darsalis Ungucc), supplies the epiglottis, soft palate, tonsil, and the mucous membrane of tongue; Sublingual (a. sublingualis), supplies mylohyoid and geniohyo- glossus muscles and sublingual gland; Ranine (a. profunda, lingua'), the terminal branch, ends at the tip of the tongue. 3. The facial artery (a. ma.villaris externa) arises from the anterior portion of the external carotid artery above the lingual. It ascends forward through the submaxillary gland, crosses the lower jaw at the anterior border of the masseter muscle, and crosses the cheek to the angle of the mouth and the side of the nose, where it terminates as the angular artery (a. angularis). Its branches in the neck are: — Inferior or ascending palatine (a. palatine ascendens) , supplies the soft palate, palatine glands, tonsil and Eustachian tube; Tonsillar (ramus tonsillaris), supplies the root of the tongue and tonsil; Submaxillary {rami glandular es) , supplies the submaxillary gland and neighboring lymphatics, muscles, and skin; Submental [a. submentalis ) , supplies the chin, lower lip and neighboring structures; Muscular, to the buccinator, masseter and internal pterygoid muscles. In the face: — Muscular, to the same muscles as given above; Inferior labial (a. labiaMs inferior), to the skin of the lower lip; Inferior coronary, to the mucous membrane, muscles and glands of the lower lip; Superior coronary (a. labialis superior), to the structures of the Upper lip and nasal septum; Lateralis nasi, to the skin of the nostrils, anastomoses with the nasal branch of ophthalmic; Angular I", angularis), supplies lachrymal sac and skin. All of these branches anastomose freely with the neighboring arteries. 4. The occipital artehy (a. arteria occipitalis) arises from the posterior portion of the external carotid, about tin' lower border of the digastric muscle. It passes beneath the stylohyoid muscle, the digastric muscle, and part of the parotid 182 HUMAN ANATOMY. gland, and, ascending, grooves the internal surface of the mas- toid portion of the temporal bone and distributes itself over the occiput as high as the vertex. Its branches are : — Muscular (rami musculares) , to splenius, digastric, stylohyoid, etc; Sternomastoid (a. stemocleidomastoidea) , crosses the hypoglossal to the sternomastoid muscle; Auricular (ramus auricularis) , is distributed to the back part of the concha; 12 10 Pig. 79. Arteries of face and head: 1, common carotid; 2, internal carotid; 3, external carotid; 4, occipital; 5, superior thyroid; 7, lingual; 9, facial; 10, temporal; 11, submental; 12, transverse facial; 13, infe- rior labial; 15, 17, inferior and superior coronary; 19, lateral nasal; 21, angular. Meningeal (ramus meningeus) , passes through the foramen lacerum posterius to the dura mater; Arteria princeps cervicis ( ramus descendens ) . Its superficial branch anastomoses with the superficial cervical, and its deep branch with the deep cervical branch from the superor intercostal. THE HEART AND VASCULAB SYSTEM. ^3 Cranial branches are distributed to the scalp over the occiput. 5. The posterior auricular artery (a. auric ularis pos- terior) arises from the posterior portion of the external carotid, on a level with the apex of the styloid process. It ascends beneath the parotid gland to a point between the mastoid process and the cartilage of the ear, where it divides into two branches, one going to the ear and the other to the occiput. Its branches are: — Stylomastoid (a. stylomasioidea) , supplies the mastoid cells, tympanum and semicircular canals, entering the cranium through the stylomastoid foramen; Auricular (ramus am inilaris ) , supplies the cartilage of the ear; Muscular, supply the sternomastoid, digastric and stylohyoid muscles; Glandular, to parotid gland. C. The ASCENDING PHARYNGEAL ARTERY (a. pharyngea OS- cendens) arises about the commencement of the external carotid artery and ascends to the base of the skull upon the rectus capitis antieus major. Its branches are: — Pharyngeal (rami pharyngei), supplies the three constrictors of the pharynx and the stylopharyngeus muscles; External, to the neighboring muscles, glands, and hypoglossal and pneumogastric nerves; Meningeal, enter the skull through the posterior and middle lacer- ated foramina and the anterior condyloid foramen. 7. The superficial temporal artery (a. temporalis su- perficialis) is the smaller of the two terminal branches. It arises in the parotid gland, crosses the root of the zvgoma, ascends forward a couple of inches, and divides into the anterior temporal and the posterior temporal. Its branches are: — Transverse facial (a. transversa faciei), supplies the masseter muscle, parotid gland and skin; Middle temporal (a. temporalis media), crosses the face to supply the temporal muscle and fascia; Anterior auricular (rami auriculwres anteriorcs) , supplies the lobule, external meatus and front part of pinna. The ENTEENAL maxillary (a. maxillaris in tenia) is the larger of the two terminal branches of the external carotid. It arises in the parotid gland about the level of the lower part of the lobe of the car. passing close to the inner side of the m of the condyle of the lower jaw to be distributed to the deep structures of the face. Its course is divided into three portions: the maxillary portion, passing inward and forward between the 184 HUMAN ANATOMY. internal lateral ligament and the ramus of the jaw; the ptery- goid portion, passing upward and forward upon the external pterygoid muscle; and the third part, or sphenomaxillary, enters the sphenomaxillary fossa. Its branches are, from the maxillary portion : — Tympanic (anterior) (a. tympanica anterior), supplies the tym- panum through the Glaserian fissure, forming a capillary plexus on the tympanic membrane; Middle meningeal ( a. meningea media ) , enters the cranium through the foramen spinosum, supplies the dura mater, and gives off Fig. 80. Internal maxillary artery: A, third portion; B, second portion; C, first portilon. a petrosal branch (ramus petrosus super ficialis) to facial nerve in hiatus Fallopii; Small meningeal (ramus meningeus accessorius) , or meningea parva, enters foramen ovale to supply dura mater and Gasserian ganglion ; Inferior dental (a. alveolaris inferior), accompanies the dental nerve through the inferior dental canal to the mental foramen, supply- ing the jawbone, teeth, chin and mouth. The pterygoid portion : — Deep temporal (aa. temporalis profunda anterior et posterior), supply the temporal muscle; Pterygoid (rami pterygoidei) , to pterygoid muscles; Masseteric (a. masseterica) , to the masseter muscle; Buccal (a. ljuccinatoria) , to the buccinator muscle. POSTEROMEDIAN GANGLIONIC POSTERIOR TEM1VRAI ANTERIOR TEMPORAL SUPERFICIAL TEMPORAL ANTERIOR AURICULAS MIDDLE TEMPORAL ORBITAL Po-TERIOR AURICULAR INTERNAL MAXILLARY ARTERU PRINCEPS CERV1CIS OCCIPITAL SUP BR. PRINCEPS CERVICIS' DEEP BR. PRINCEPS CERVICIS ASCENDING CERVICAL DEEP CERVICAL SUPERFICIAL CERVICAL SUPRASCAPULAR R. SUBCLAVIAN POST SCAPULAF SUP \ ENA CAVA B. PULMONARY A R I'l I.MONARV VEIN R AUBICLE • '.AKY INK VENA CAVA IHOBACII AORTA ANTERIOR _CI IMMUNICATING ANTERIOR CEREBRAL — MIDDLE CEREBRAL " OPHTHALMIC . TEMPORAL . POST COMMUNICATING POST CEREBRAL - SUPERIOB CEREBELLAR. SUP STERN0-MAST0ID. BASILAR - ANT. INFERIOR CEREBELLAR - L. INT CAROTID . FACIAL _ FIG. 81. 'I'lio art'Tiis o!' the head and neck, THE HEART AND VASCULAR SYSTEM. 185 Sphenomaxillary portion : — Alveolar or posterior dental branch (a. aheolaris superior poste- rior), supplies the upper molar and bicuspid teeth, antrum and gums; its principal branch is the superior dental; Infraorbital (a. infraorbitaUs) , passes through infraorbital canal to face: Posterior or descending palatine («. palatina descendens), descends posterior palatine canal and runs forward to foramen of St<'iison; Vidian (or. canalis pterygoidea) , accompanies the Vidian nerve; Pterygopalatine, supplies Eustachian tube and pharynx; Nasal, or sphenopalatine (a. sphenopalatine/,), descends through sphenopalatine foramen to antrum, sphenoidal and ethmoidal cells. THE INTERNAL CAROTID ARTERY (A. CAROTIS INTERNA) arises about the upper border of the thyroid cartilage from the bifurcation of the common carotid. It ascends in front of the transverse processes of the cervical vertebrae to the carotid fora- men in the petrous portion of the temporal bone to supply the anterior part of the brain, the eye with its appendages, and send branches to the nose and forehead. It gives no branches until it passes through the carotid canal, where, after piercing the dura mater at the anterior clinoid process, it divides into its terminal branches — the anterior and middle cerebral. Relations. — The internal carotid artery in the neck is in relation in front with the parotid gland, the glossopharyngeal nerve, the stylo- glossus and stylopharyngeal muscles, and is covered by the skin, super- ficial and deep fascia'; behind it rests upon the rectus capitis anticus major and has resting upon it the superior laryngeal nerve and the sympathetic; internally it is in relation with the pharynx, tonsil, superior laryngeal nerve and descending pharyngeal artery; externally ii is in relation with the internal jugular vein and the pneumogastric nerve. Its branches arc, from the petrous portion: — ■ Tympanic, internal or deep [ramus caroticotympamcus) , sup- plies tympanum. From the cavernous portion : — Artiriiv rrr-< [ttaciili, supply the Gasserian ganglion and pituitary body; Anterior meningeal ('/. menigea anterior), a branch from the pre- oeding v—el to the dura. Ophthalmic ('/. ophthalmica) arises about tbe position of the anterior clinoid process and passes through the optic fora- men to the outer side of tin- optic ner\e jo reach the orbit. At the inner angle of the eye it divides into its two terminal 186 HUMAN ANATOMY. branches — the frontal and the nasal. It supplies the muscles and the globe of the eye, and its branches may be divided into two groups — the orbital and the ocular. Orbital group : — Lachrymal (a. laohrimaUs) , supplies the lachrymal gland and gives off a malar branch which passes through the malar bone to the temporal fossa; Supraorbital (a. supraorbitals) , passes out through the supra- orbital foramen to supply the surrounding structures; Posterior ethmoidal (a. ethmoidalis jjosterior) , descends through the posterior ethmoidal foramen to the cells and adjacent parts; Anterior ethmoidal (a. ethmoidalis anterior) } through the ante- rior ethmoidal foramen to the nose; Internal palpebral (aa. palpebrales mediales) , supply the eyelids. Frontal (a,, frontalis), supplies the skin and muscles of the fore- head. Nasal (a. dorsalis nasi), supplies the lachrymal sac and bridge of the nose. Ocular group : — Muscular, supply the muscles of the eye; Anterior ciliary {aa. ciliares anteriores) , supply the iris, piercing the sclerotic near the cornea ; Short ciliary (aa. ciliaris posteriores breves), supply the choroid and ciliary processes, piercing the sclerotic near the optic nerve; Long ciliary (aa. ciliares posteriores longw), enter with the short ciliary, but run forward to the iris; Arteria centralis retinae, supplies the optic nerve and the retina. From the cerebral portion : — Anterior cerebral (a. cerebri anterior), supplies the ante- rior part of the cerebrum, the optic and olfactory nerves, and com- municates with its fellow by the anterior communicating; Middle cerebral, or Sylvian artery ( a. cerebri media ) , ascends in the Sylvian fissure, and supplies the middle lobes of the cerebrum; Posterior communicating (a. communicans posterior) , runs back- ward to join the posterior cerebral; Anterior choroid (a. choroidea) , supplies the choroid plexus. THE CIRCLE OF WILLIS (CIRCULUS ARTERIOSUS [wiLLISl] ) is an anastomotic circle formed at the base of the brain about the pituitary body, between the branches of the internal carotid and the vertebral arteries, for the purpose of equalizing the cerebral circulation. The anterior cerebrals, from the internal carotids, pass forward and are united in front by a short trunk, the anterior communicating. The posterior communicating unites the internal carotid with the posterior cerebral, the latter being the terminal branches of the basilar, which in its turn is formed by the union of the two vertebrals. Acromial thoracic M fl KJ"H I HoRai |c LONG THORACIC HAUIAI. ISDICIB nL^K PALMAH DIGITALS Fig. 82. The arteries of thr» upper extremity. THE HEART AND VASCULAR SYSTEM. 18? TIIK SUB< LAVIAX AKTERY (A. srBCLAVIA) on the right side arises from the arteria innominata, hehind the right sternoclavicular articulation, and on the left side directly from the arch of the aorta. They pass outward to the outer margin of the first rib, where they become the axillary, being divided in their course by the scalenus anticus muscle into three portions — the first portion to the inner side of the muscle, sec- ond portion behind it, and the third portion between its outer margin and the lower border of the first rib. The vessels differ only in their first portions, the left ascending more vertically. Relation The first portion of the right subclavian is in relation in front to the internal jugular and vertebral veins, the sternohyoid and sternothyroid and the clavicular portion of the sternomastoid muscles and the* pneumogastric, phrenic, and cardiac nerves; behind it is in relation with the longus colli muscle, the transverse process of the first dorsal or seventh cervical vertebrae, the sympathetic nerve, and the recurrent laryngeal nerve beneath with the pleura. The first portion of the left subclavian artery is in relation in front with the left internal jugular and innominate veins, the left carotid artery, the sternothyroid, sternohyoid, and sternomastoid muscles, the "pleura and left lung, the pneumogastric, phrenic and cardiac nerves; behind, with the vertebral column and longus colli muscle, the esophagus, thoracic duct, and the inferior cervical ganglion of the sympathetic; on the inner side with the esophagus, thoracic duct, and trachea, and on the outer side with the pleura. The second portion of the subclavian artery on either side is in relation with the subclavian vein, the scalenus anticus muscle and the phrenic nerve; behind with the middle scalenus muscle and pleura; above by brachial plexus, and below by the pleura. The third portion of the subclavian artery on either side is in relation in front with the exter- nal jugular vein and its branches, the clavicle, subclavius muscle and suprascapular artery, the descending branches of the cervical plexus and the cervical fascia; behind with the scalenus medius, above with the omohyoid muscle and the brachial plexus, and below with the first rib. [te branches are: — 1. Vertebra] i". oertebraUs) . 5. Transverse cervical, or 2. Thyroid axis [truncus thy- transversal is colli (a. 1 1 <>c< i rifiilis) . transversa colli). :;. Inferior thyroid (a. thy- 0. Internal mammary (a. /■<',»/< humeri posterior), en- circles the neck of the humerus to supply the shoulder joint and the deltoid muscle; Anterior circumflex (a. circumfiexa humeri anterior), de- scends beneath the short head of biceps and coracobrachial muscles to supply the shoulder joint. THE BRACHIAL ARTERY (A. BRACHIALIS) commences at the lower border of the latissimus dorsi and teres major muscles and descends on the front and inner aspect of the arm to half an inch below the middle of a line drawn through the condyles of the humerus, where it terminates by dividing into the radial and ulnar artery. Relations. — It is covered in front by the skin, superficial and deep fascia, and is crossed by the median nerve; behind it rests upon the triceps, brachialis anticus and coracobrachial muscles; And is in relation with the musculospiral nerve and the superior profunda artery ; to the outer side above it is in relation with the biceps and coracobrachialis muscles and the median nerve; and on the inner side with the internal cutaneous and ulnar nerve, and the basilic vein and median nerve below. At the bend of the elbow it is crossed by the bicipital fascia. Its branches are: — -Superior profunda (a. profunda brachii), accompanies the musculospiral nerve to supply the triceps, deltoid and coraco- brachialis muscles; Nutrient artery (a. nutricia humeri), supplies the shaft of the humerus; Inferior profunda {a. eollateralis ulnaris superior), accom- panies the ulnar nerve to supply the triceps muscle and struc- tures about the internal condyle ; Anastomotica magna (a. eollateralis ulnaris inferior), forme as anastomosis on the lower posterior aspect of the arm, with branches from the inferior profunda and recurrent ulnar -'.•Is. It supplies the triceps; Muscular (rami iinisiiilares), supply the muscles on the anterior aspect of the arm. THE BADIAL ARTERY (A. RADIALIS) :ds from the bifurcation of the brachial below the bend of the elbow to the wrist, where it winds around the carpus 192 HUMAN ANATOMY. beneath the extensor tendons of the thumb, passes between the two heads of the first dorsal interosseous muscle and into the palm of the hand, where it forms the deep palmar arch, anasto- mosing with the deep branch of the ulnar artery. Its branches are, in the forearm : — Radial recurrent (a. recurrens radialis) , supplies the braehialis anticus and supinator longus and brevis; Muscular {rami musculares) , supply the radial muscles; Superficialis volse (ramus volaris superficialis) , supplies the muscles of the thumb and anastomoses with the ulnar to form the superficial palmar arch; Anterior radial carpal (ramus carpeus volaris), supplies the wrist-joint. In the wrist: — Posterior radial carpal (ramus carpeus dorsalis) , supplies the wrist-joint, forms the posterior carpal arch, and gives off the dorsal interosseous arteries; Metacarpal, is called the first dorsal interosseous; it supplies the index and middle finger; Dorsales pollicis, supply the back of the thumb; Dorsalis indicis, supplies the outer and dorsal side of the index finger. In the hand: — Princeps pollicis (a. princeps pollicis), supplies the sides of the palmar aspect of the thumb; Radialis indicis (a. volaris indicis radialis), supplies the radial side of index finger; Perforating (rami perf or antes) , anastomose with dorsal interos- seous arteries; Interosseous (aa. metacarpew volares) , anastomose with digital branches of superficial palmar arch. THE ULNAR ARTERY (A. ULNARIS) descends from the bifurcation of the brachial below the bend of the elbow to the ulnar border of the wrist, passing beneath all the superficial flexors excepting the flexor carpi ulnaris, crosses over the annular ligament at the radial side of pisiform bone, and enters the palm to form the superficial palmar arch, anastomosing with the superficialis volse from the radial. Its branches are, in the forearm : — Anterior ulnar recurrent (a. recurrentes ulnaris anterior), sup- plies the pronator radii teres and braehialis anticus muscles; Posterior ulnar recurrent (a. recurrentes ulnaris posterior), sup- plies the joint and neighboring muscles, and anastomoses freely; Interosseous (a. interossea communis), passes backward to the upper border of the interosseous membrane, where it divides into THE HEART AND VASCULAR SYSTEM. 193 Anterior branch (a. mterossea volaris), descends on the front of the interosseous membrane; Posterior (a. intcrossea dorsalis) , descends on the posterior as- pect of the foramen and gives off a recurrent interosseous branch- Muscular {rami musculares), to muscles on ulnar side of forearm. In the wrist: — Anterior carpal (ramus carpeus volaris), supplies the front of wrist and anastomoses with the carpal branches of radial artery- Posterior carpal (ramus carpeus dorsalis), passes beneath the ten- don ot flexor carpi ulnaris and forms posterior carpal branch. In the hand : — Deep, or communicating branch (ramus volaris profundus) passes between the flexor and abductor minimi digiti to form part of deen palmar arch; * x «i l« Dlgfi1tal',.!"Pply- the Ul!\a,r Skle of the little finSer and adjoining sides of the little, ring, middle and index Angers. THE DESCENDING AORTA. The thoracic aorta (aorta thoracalis) extends from the .wer border of the fifth dorsal vertebra on the left side to about the last dorsal vertebra, where by passing through the aortic opening m the diaphragm it becomes the abdominal aorta. Relations— It is in relation in front with the left bronchus esoph- agus pericardium and left pulmonary artery; behind it rests uj'xuxe vertebral column and is ,n relation with the vena azygos minor; on the right side it is m relation with the thoracic duct; the esophLms at its upper part and the vena azygos major; on the left side with 'the left lung, the pleura, and the esophagus below. The branches of the thoracic aorta are : Pericardiac (rami pericardiaci) , supply the pericardium- Bronchial (aa. bronchioles), two or three in number, sup- ply the bronchi ; ' l Esophageal (aa. a> so phage®), supply the esophagus • Posterior mediastinal (rami mediastinales) , supply the neighboring pleura and lymphatic glands; Intercostal (aa. intercostal es), ten on each side, supply the intercostal spaces. They divide into— anterior branch, to inter- costal and pectoral muscles; posterior branch, to spinal column and dorsal muscles. THE ABDOMINAL AORTA (AORTA ABDOMINALIS) descends from the Inst dorsa] vertebra at the aortic opening of the diaphragm, to opposite the fourth lumbar vertebra, a little 194 HUMAN ANATOMY. to the left of the median line, where it divides into the two common iliac arteries. Relations. — Th" abdominal aorta is in relation in front with the pancreas, transverse duodenum, stomach, and lesser omentum, the mes- entery, splenic vein, left renal vein, celiac, solar and aortic plexuses; behind, it is in relation with the vertebral column, the thoracic duct, the left lumbar veins and the receptaculum chyli; on the right side with the inferior vena cava, vena azygos, thoracic duct, the right crus of the diaphragm above, and the right semilunar ganglion; on the left side with the left semilunar ganglion and the sympathetic nerve. The branches of the abdominal aorta are : — Phrenic, two in number, supply the diaphragm; Celiac axis (a. ccdiaca), divides into three large branches, the gastric, hepatic and splenic; 1. Gastric, or coronaria ventriculi (a. gastrica sinistra), is the smallest, and supplies the lesser curvature of the stomach ; 2. Hepatic (a. hepatica), enters the transverse fissure of the liver, and divides into two branches to supply the lobes of the liver. It gives off : — Pyloric (a. gastrica dextra), to lesser curvature of stomach; Gastroduodenalis, which descends behind the pylorus, and divides into Pancreaticoduodenalis superior branch to supply the pancreas and duodenum ; Gastroepiploica dextra, along the greater curvature of the stomach from right to left. Cystic ( a. cystica ) , supplies the gall-bladder. 3. Splenic (a. lienalis), the largest of the three, supplies the spleen and gives off the (a) Pancreatic (rami pancreatici) — to the pancreas; (b) Gastroepiploica sinistra, runs from left to right, along the greater curvature, to join the gastroepiploica dextra; (c) Gastric — vasa brevia (aa. gmtricw brevis), supply the fun- dus of the stomach. Superior mesenteric (a. mesenterica superior), supplies the small intestine (except first part of the duodenum) and the large intestine as far as the splenic flexure, and part of the descending colon. It gives off: — (a) Inferior pancreaticoduodenal (a. pancreaticoduodenalis infe- rior), supplies duodenum and pancreas; (b) Vasa intestini tenuis (aa. intestinales) , supply nearly the whole length of the small intestine (ileum and jejunum) ; (c) Ileocolic (a. ileocolica) , descends to the right iliac fossa, be- tween the layers of the mesentery, and divides into two branches; (d) Right colic artery (colica dextra), passes to the middle of the ascending colon, to divide into two branches; THE HEART AND VASCULAR SYSTEM. 195 («) Middle colic artery (colica media), ascends in the layers of the transverse mesocolon, to divide into two branches. Suprarenal (a. suparenalis media), supply the suprarenal bodies ; Renal (aa. renales), supply the kidneys, a large trunk on each side springing directly from the aorta; Spermatic (aa. spermatica} interna'), descend to supply the testicles, and in their passage form one of the constituents of the spermatic cord; Fig. 83. Abdominal aorta: 1, phrenic; 2, celiac axis; 3, superior mesen- t« ri<- ; 4, suprarenal: 5, renal; G, spermatic; 7, inferior mesenteric; 8, lumbar; 9, sacra media. Inferior mesenteric (aa. mesenterica inferior), gives off: — (a) l.cfi colic [coUca amisira), supplies the descending colon; 3igmoid (aa. aigmoidece), supplies the sigmoid flexure of the colon ; (c) Superior hemorrhoidal (». hrrh<>in it- posterior aspect. Lumbal branches («". Iwmbales), four or five on each side. tackward and outward and divide into the 196 HUMAN ANATOMY. Dorsal branches (rami dorsales), supply the spinal cord, muscles, and skin of back ; Spinal branches (rami spinales), divide within the spinal canal, ascend and descend, and form an arterial network throughout the whole length of the spinal canal; Abdominal branches (rami parietales), supply the abdom- inal walls; Middle sacral (a. sacralis media), is the continuation of the abdominal aorta, descending in the middle of the sacrum to the upper part of the coccyx. THE COMMON ILIAC ARTERIES (AA. ILIAC.E COMMUNES) descend from the bifurcation of the abdominal aorta, opposite the body of the fourth lumbar vertebra, to a point midway between the last lumbar vertebra and the sacrum, where they divide into the external and internal iliac arteries. The right common iliac is a little larger than the left, and passes more obliquely outward. Relations. — The right common iliac is in relation in front with the small intestine, the ureter, the peritoneum and the sympathetic nerves; behind, with the left and right common iliac veins, and to its outer side it rests upon the psoas musole, and is in relation with the vena cava and the right common iliac vein. The left common iliac is in relation in front with the ureter, the peritoneum, the superior hemor- rhoidal artery and the sympathetic nerves; behind, with the left com- mon iliac vein, to its outer side the psoas muscle, and to its inner side the left common iliac vein. The common iliac arteries give off a few unimportant branches to the ureters, peritoneum, etc. the internal iliac or hypogastric artery (a. hypogastrica) is about an inch and a half in length, extending from the bifur- cation of the common iliac, about the sacrolumbar junction, to the upper border of the great sacrosciatic foramen, where it divides into the anterior and posterior trunks. The branches from the anterior trunk are: — Superior vesical (a. vesiculis superior), is the pervious remains of the fetal hypogastric artery. It supplies the apex and body of bladder, ureter and vas deferens; Middle vesical (a. vesiculis medialis), supplies the base of the bladder and part of vesiculae seminales ; it may be given off from the superior vesical; r jit. suprarenal cystic R. MID SUPRARENAL. INK. PAXCREATICO DUODE.XAI GASTRO-DUODEN U. R. INF. SUPRARENAL SUP PAXCREATICO DUODENAL. R. GASTROEPIPLOIC R. RENAL MIDDLE COLIC RIGHT COLIC LUMBAR SPERMATIC ABDOMINAL AORTA INF. MESENTERIC MIDDLE SACRAL R. COMMON ILIAC SUP HEMORRHOIDAL ILIO LUMBAR R INT ILIAC R. EXT. ILIAC POST. TRUNK OF INT ILIAC ANT TRUNK OF INT. ILIAC. SUPERIOR VF-I. n OBLITERATED I) I OBTURATOR INFERIOR VESICAL MIDDLE VESICAL MID BiCMOKRHOTDAJ INTERNAL II HI SCIATIC MIDDLE VESICAL DEEP EriUASTRIC Fig. 84. The arteries of the trunk. THE HEART AND VASCULAR SYSTEM. 197 Inferior vesical (a. vesicalis inferior), supplies the prostate gland, base of bladder and vesiculae seminales ; Middle hemorrhoidal (a. hcemorrhoiddlis media), supplies part of the rectum ; Obturator (a. obiuratoria), passes along the side of the pelvis to the obturator foramen, through which it passes. It gives off within the pelvis the iliac (ramus ifiacus), vesical 1 ram us vesicalis), and pubic (ramus pttbicus) branch; Internal pudic (a. pudenda interna), supplies the external organs of generation. It passes out of the pelvis through the great sacrosciatic foramen, winds around the ischial spine, and re-enters the pelvis through the lesser sacrosciatic foramen. It gives off the following branches in the perineum : — (a) Inferior hemorrhoidal (a. hcBmorrhoidalis inferior), supplies the muscles and skin of the anus; (b) Superficial perineal (a. perinei) , supplies the erector penis and accelerator urinae muscles and the scrotum; (c) Transverse perineal, crosses the perineum on the transversus muscle, which, together with the structures between the bulb and anus, it supplies; (d) Bulbourethral, artery of the bulb (a. bulbi urethras), sup- plies the bulb of the spongy body and Cowper's gland; (e) Cavernous [a. profunda penis), supplies the corpus caver- nosum, and is one of the terminal branches: {f) Dorsal artery of penis, or clitoris (a. dorsalis penis), runs forward on the dorsum and supplies the glans and prepuce. In the female the internal pudic artery is of smaller size, but its course and distribution is quite analogous to its study in the male subject, i.e., the superficial perineal artery supplies the labia pudendi ; the artery of the bulb, the vestibule, and the vaginal erectile tissue, the artery of the corpus cavernosum, the clitoris; the arteria dorsalis clitoridis, the dorsal surface of the clitoris, being distributed to the membranous fold in the glans clitoris, the analogue of the male prepuce. Sciatic artery supplies the muscles at the back of the pelvis, coccygeug, pyriformifl and levator ani, and passes out of the greal sacrosciatic foramen to follow the course of the sciatic Its branches external to the pelvis are: — Coccygeal, supplies back part of coccyx; Inferior gluteal [a. gluteal inferior), to the gluteus maximus muscle : < omes nervi iachiadici ('/. lomilaiis ». ixchiadici), runs in the rabctance of the great sciatic nerve; Mii-enlar, to the back part of the hip; Articular, to supply the capsule of the hip-joint. 198 HUMAN ANATOMY. Uterine (a. uterina), is distributed to body of uterus,, giv- ing branches to the ureter and bladder ; Vaginal (a. vaginalis), supplies the mucous membrane of the vagina, giving branches to the rectum and neck of the blad- der. It is analogous to the inferior vesical, in the male. From the posterior trunk Iliolumbar (a. iliolumbalis) gives off two branches: — Lumbar {ramus lumbalis), supplies quadratus lumborum and psoas muscles; Iliac {ramus iliacus), supplies iliacus, gluteal and abdomi- nal muscles. Lateral sacral {a. sacralis lateralis), are two in number — the superior and inferior, supplying the contents of the sacral canal ; Gluteal {a. glutece superior), gives a superficial and deep branch to supply the gluteus maximus, medius and minimus muscles. THE EXTERNAL ILIAC ARTERY (A. ILIACA EXTERNA) extends from the bifurcation of the common iliac to Poupart's ligament, under which it passes to become the femoral. Relations. — It is in relation in front with the peritoneum and intestines, circumflex iliac vein, spermatic vessels, genitocrural nerve and the lymphatic vessels and nerves; behind, it rests upon the psoas muscle, and is in relation with the external iliac vein, which, together with the vas deferens, is also in relation to it, on its inner side, beneath Poupart's ligament. On its outer side it is in relation with the psoas inagnus and iliac fascia. Its branches are : — The deep epigastric {a. epigastrica inferior), arises above Poupart's ligament and ascends obliquely inward between the transversalis fascia and peritoneum to the rectus muscle, in which it ascends to anastomose with the termination of the internal mammary artery. It gives off: — Cremasteric {a. spermatica externa [male]; a. ligamenti teretis uteri [female] ) branch, to supply the cremasteric muscle; Pubic ( ramus pubicus ) , crossing Poupart's ligament to descend to the inner side of the femoral ring, and . Muscular, to supply abdominal muscles and the peritoneum. The deep circumflex iliac {a. circamflexa ilium profunda) ascends outwardly behind Poupart's ligament to the crest of the ilium, where it gives off : — Muscular branches, to supply the abdominal muscles. ILIOLl MPAR 1MTEKNAI Hue 01 m i mi bi 'i roi ]'•[<:. 86. The arteries of the lower extremity. THE HEART AND VASCULAR SYSTEM. 199 THE FEMORAL ARTERY (A. FEMORALIs) extends- from Pouparfs ligament down the inner side of the thigh to the opening in the adductor magnus — Hunter's canal ■ lis adductorius | llunteri]), where it becomes the popliteal. Its course corresponds to a line drawn from a point midway between the anterior superior spine of the ilium and the sym- physis pubis to the inner tuberosity of the internal condyle. In the upper part of its course it is superficial, where it passes through Scarpa's triangle (trigonum femorale), but in the lower part it passes backward and becomes very deep. Where it passes under Pouparfs ligament the femoral vein is to the inner side, and the anterior crural nerve to the outer side; thus, from within outward, Y. A. N. Relations. — It is in relation in front with, the sartorius, the long saphenous nerve, a branch of the anterior crural nerve, and is covered liv the fascia lata; behind, it is in relation with the psoas magnus, adductor longus, adductor magnus and pectineus and the profunda vein, and about the middle with the femoral vein; at its inner side it is in relation with the sartorius and adductor longus muscles, and at its upper part with femoral vein; on the outer side, with the vastus internus, and at its lower part, the femoral vein. The branches of the femoral artery are: — Superficial epigastric (a. epigastrica superficialis) , descends through the saphenous opening in the fascia lata, and ascends in abdomen, supplying the skin, inguinal glands and superficial fascia; Superficial circumflex iliac (a. circumflexa ilium superfi- cialis), passes outward to the skin of the groin and over crest of ilium ; Superficial external pudic (a. pudenda externa superfi- cialis), supplies the skin on the lower part of the abdomen; Deep external pudic (a. pudenda externa profunda), sup- plies the scrotum in the male, the labia pudendi in the female; Profunda femoris, arises about two inches below Pouparfs Hi, and passes beneath the adductor longus, giving off the following: — External circumflex («. circumflexa femoris lateralis), gives off ascending, transverse and descending branches, supplying the muscles on the front of the thigh as low as the knee; Internal circumflex (a. circumflexa femoris medialis), internally, supplying the adductor muscles and the hip joint; Thn-c perforating (ua. perforans prima, secunda, tertia), the adductor magnus and brevis, and supply the flexor 200 HUMAN ANATOMY. muscles of the thigh, and give off the medullary nutrient artery (a. nutricia femoris). The- vessel terminates by a fourth per- forating. Muscular (rami muscular es), vary in number, and supply the sartorius and vastus internus; Anastomotica magna (a. genu supremo), gives off two branches — superficial, accompanying the long saphenous nerve, and deep branch, to the inner side of knee, and supplies knee joint. THE POPLITEAL ARTERY (A. POPLITEA) descends! from the opening in the adductor magnus to the lower border of the popliteus muscle, passing behind the knee joint, where it divides into the anterior and posterior tibial arteries. Its branches are : — Muscular, gives off two principal branches : — Superior, to flexors of thigh and vastus externu's muscles ; Inferior, or sural (aa. swales), to heads of gastrocnemius and plantaris. Cutaneous, supplies the skin of the calf; Superior external articular (a. genu superior lateralis), supplies knee joint and vastus externus ; Superior internal articular (a. genu superior medialis), supplies knee joint and vastus) internus; Azygos articular (a. genu media), supplies the synovial membrane and ligaments of joints ; Inferior external articular (a. genu inferior lateralis), sup- plies knee joint and fibular head; Inferior internal articular (a. genu inferior medialis), sup- plies knee joint and head of tibia. THE ANTERIOR TIBIAL ARTERY (A. TIBIALIS ANTERIOR) descends from the bifurcation of the popliteal at the lower bor- der of the popliteus muscle ; passes over the upper border of the interosseous membrane, between the two heads of the tibialis posticus,- and descends on the anterior part of the interosseous membrane and lower part of the tibia to the ankle joint, where it terminates as the dorsalis pedis. In the upper third of its course it lies between the tibialis anticus and extensor longus digitorum, resting upon the interosseous membrane, in the middle third, between the tibialis anticus and extensor proprius pollicrs, and at the lower third it becomes more superficial, and lies between the extensor proprius and extensor longus digitorum tendons. THE HEART AND VASCULAR SYSTEM. 201 Its branches are : — Anterior recurrent tibial (a. recurrens tibialis anterior), ascends to supply the front and sides of the knee joint and anastomose with the anastomotica magna and popliteal; Posterior recurrent tibial (a. recurrens tibialis posterior), is an inconstant artery that is given off from the anterior tibial, before the latter passes through the interosseous space. It sup- plies the popliteus muscle, and anastomoses with some of the articular filaments of the popliteal artery; Muscular, supplying the muscles and skin of the neighbor- ing parts; Internal malleolar (a. malleolaris anterior medialis), arises two inches above the ankle joint, and supplies it and the struc- tures on its inner side; External malleolar (a. malleolaris anterior lateralis), sup- plies the outer side of the ankle. The doiisalis pedis artery (a. tlorsalis pedis) extends from the bend of the ankle, where it is continuous with the anterior tibial, to the first interosseous space, where it divides into the dorsalis hallucis and the communicating. Its branches are: — Tarsal (a. tarsea lateralis), supplies the tarsal articulations ami the extensor brevis digitorum muscle; .Metatarsal (a. arctiata), passes to the outer side of the foot, and gives off three interosseous branches to supply the adjacent sides of the toes; Dorsalis pollicis (a. dorsalis hallucis), passes along the outer border of the great toe, supplying it; Communicating (ramus plantaris profundus), descends be- tween the two heads of the first dorsal interosseous to complete, with the external plantar, the plantar arch; Interosseous, branches of the tarsal branch, three in num- ber. THE POSTERIOR TIBIAL ARTERY (a. TIBIALIS POSTERIOR) begins at the lower border of the popliteus muscle and descends along the posterior and tibial side of the leg to the space between the- heel and the inner ankle, where it passes beneath the abduc- tor pollicis and divides into the internal and external plantar arteries. Relatione. — In tin- upper |>art of its course it lies deeply, being covered by the gastrocnemius and aoleus muscle, but in the lower part it becomes more superficial, being covered only by the skin and the fascia. 202 HUMAN ANATOMY. Its branches are : — Peroneal (a. peroncea), descends the inner border of the fibula, supplying the muscles and skin of that region and the back of the ankle ; Anterior peroneal (ramus perforans), a branch of the pero- neal, pierces the lower part of the interosseous membrane to supply the dorsum and outer side of the tarsus; „ Muscular, supplies the posterior muscles; Nutrient (a. nwtritia fib idee), supplies the tibia, being the largest nutrient artery in the body; Communicating (ramus communicans) , passes to the pero- neal; Internal calcanean (rami calcanei mediates), supplies the inner side of the sole and heel. The internal plantar artery (a. plantaris medialis), the smaller of the two, passes along the inner side of the foot between the abductor pollicis and the flexor brevis digitorum to the inner border of the great toe, anastomosing with its digital branch. The external plantar artery (a. plantaris lateralis) passes across the foot to the base of the fifth metatarsal bone, where it turns inward and crosses the foot to the first inter- osseous space, where it anastomoses with the communicating branch of the dorsalis pedis to form the plantar arch. Its branches are : — Posterior perforating (rami perforant.es posteriores) , are three branches which pass between the heads of the dorsal interossei muscles; Digital branches (aa. metatarsece plantares) — these are four branches which supply the adjacent sides of the three outer toes, and the outer sides of the second and little toes. Arch of aorta, TABLE OF THE ARTERIAL SYSTEM. Aorta. Right coronary. Left coronary. Innominate, Left common carotid, Left subclavian. (Right common f External carotid, carotid, \ Internal carotid. Right subclavian. / External carotid, I Internal carotid. THE HEART AND VASCULAR SYSTEM. 203 r Ilyoid, Superiot i Superficial descending branch, thyroid, 1 Superior laryngeal, I Cricothyroid. External carotid, External carotid - continued, Lingual, Ilyoid, Dorsalis linguse, i Sublingual, I Ranine. Facial, In the aeckj On the face, Inferior or ascending palatine, tonsillar, submaxillary, sub- mental, muscular. Muscular, inferior labial, inferior coronary. superior coronary, lateralis nasi, angular. Occipital, J p Muscular. [ Sternomastoid, Auricular, Meningeal, Arteria princeps cervicis, Cranial branches. r Stylomastoid, Posterior j Auricular, auricular, 1 Muscular, I Glandular. Ascending 1 Pharyngeal, external meningeal, pharyngeal /•'"»' & Pi Temporal, Transverse facial, Middle temporal, Auricular, Anterior temporal, Posterior temporal. Internal maxillary, Maxillary portion, ' Tympanic ( anterior ) , Deep auricular, Middle meningeal, Small meningeal, Inferior dental. i Deep temporal, Pterygoid J Pterygoid, portion. 1 Masseteric, I Buccal. Spheno- maxillary pari ion. Alveolar, Infraorbital, . Tost, or ascending palatine. j Vidian, Pterygopalatine, N'asal or splicnopalatal. 204 HUMAN ANATOMY. i o -a X ortion9 } Tympanic -( internal, or deep). Cavern- ous portion, Arterise receptaculi, Anterior meningeal. Ophthalmic, Orbital group, Ocular group, Lachrymal, Supraorbital, Post, ethmoidal, Ant. ethmoidal, Palpebral, Frontal, Nasal. Muscular, Ant. ciliary, Short ciliary, Long ciliary, Arteria centralis retinae. Cerebral portion, Verte- bral, r Anterior cerebral, j Middle cerebral, I Anterior choroid, L Posterior communicating. Cervical Branches, Thyroid axis, Lateral spinal, Muscular. . Cranial Branches, , A . Posterior meningeal, Anterior spinal, _ Posterior spinal. Inferior thyroid, Superior scapular, ( Transversalis humeri ) , Transversalis colli, Transverse, Anterior inferior Basilar, Bifurcation. Recurrent tibial, Muscular, Internal malleolar, External malleolar. Dorsalis pedis ( continuation ) , Tarsal, 1 Three Metatarsal, J interosseous. Dorsalis pollicis or hallucis, Communicating, Interosseous. Posterior tibial, Peroneal, Muscular, Nutrient, Communicating, Internal calcanean, Anterior peroneal. Internal plantar, j Bifurcation. External plantar, J (fromexteraal 1 Three Posterior perforating, ( plantar) J F°Ur **** Pulmonary Artery (a. pulmonalis). — The pulmonary artery carries venous blood from the right ventricle to the lungs. It 'is about two inches in length, passes upward and backward to the left side to the under surface of the transverse portion of the arch of the aorta, where it divides into the right and left pulmonary arteries. It is attached to the under portion of the arch by a fibrous cord, the remains of the ductus arteriosus of fetal life (lig amentum arteriosum). The right pulmonary artery (ramus dexter a. pulmonalis) is larger and longer than the left (ramus sinister a. pulmonalis), and passes behind the ascending aorta and superior vena cava to the root of the right lung, where it divides into two branches. The left pulmonary artery passes in front of the descending aorta and left bronchus to the root of the left lung, where it divides into two branches. THE HEART AND VASCULAR SYSTEM. 209 THE VENOUS SYSTEM. The venous system, like the arterial, consists of two dis- tinct sets, the systemic and pulmonary. It is composed of seven { J ) main trunks and their branches : — Systemic. Pulmonary. 1. Coronary vein; Four pulmonary veins. 2. Superior vena cava; 3. Inferior vena cava. The systemic veins return the venous blood from the body, head and extremities to the right auricle. The portal vein, with its branches and capillaries, is an appendage to the systemic set, collecting the venous blood from the organs of digestion, i.e., stomach, intestines and the pan- creas ;*and also from the spleen, and carrying it to the liver, where it breaks into capillaries, and finally reaches the inferior vena cava by means of the hepatic veins. The pulmonary veins (v. pulmonales) are peculiar in carry- ing arterial blood from the lungs to the left auricle. Systemic Veins. — The coronary sinus (sinus coronarius) returns all the blood from the substance of the heart, except that returned directly from the walls of the right auricle by the vena Thehesii | venoe cordis minima). It is a dilatation of the great cardiac vein, about one inch in length, situated in the posterior part of the left auriculoventricular groove. It opens into the right auricle, its orifice being protected by the coronary valves, and receives the following : — Great cardiac, or left coronary Right cardiac or small coronary vein [v. cordis magna); vein {v. cordis parva.) ; Anterior cardiac (vv. cordis an- Left cardiac vein ( r. posterior ven- t, , lores) ; triculi simstri I ; Middle cardiac or posterior (v. Oblique vein of Marshall (v. ob- cordis media) ; /'7»« «*r" sinistn). The superior vena cava (v. cava superior) is a short trunk formed by the union of the right and Left innominate wins. It receives the vena azygos major (v. azygos), has no valves, and is smaller in size than the aorta. It ends in the right auricle, receiving the blood from the whole upper half of the body and the right Lymphatic and thoracic duets. The left innominate anonyma sinistra) passes to join the superior cava m front of the greal arteries of the arch. The inferior thyroid, the internal mammary and vertebra! veins follow closely the courses 0f the corresponding art. Tie., and terminate in the innominate Vein. u 210 HUMAN ANATOMY. The internal jugular vein (v. jugularis interna) is formed by the lateral and thq superior petrosal sinuses, descends at the outer side of the common carotid behind the anterior border of the sternomastoid muscle, and joins the subclavian vein to form the innominate. At its junction with the subclavian the left internal jugular vein receives the thoracic duct and the right internal jugular vein the right lymphatic duct. The Sinuses of the Dura Matek (sinus dwrce matris) . — These are venous channels analogous to veins between the layers of the dura mater. They are destitute of valves, follow no vessels, and their inner coat is continuous with the lining membrane of the veins. They are divided into two sets; those at the back and upper part of the skull are : — Superior longitudinal, Lateral sinuses (2), Inferior longitudinal, Occipital sinus. Straight sinus, And those of the base : — Cavernous (2), Superior petrosal (2), Circular, Transverse (anterior occipital, Inferior petrosal (2), Leidy ) . The superior longitudinal sinus (sinus sagittalis superior) arises at the foramen cecum, passes backward along the margin of the falx cerebri to the crucial ridge of the occipital bone, where it terminates in the torcular Herophili or confluence of the sinuses. The inferior longitudinal sinus (sinus sagittalis inferior) passes in the free margin of the falx cerebri. The straight sinus (sinus rectus) passes backward from the junction of the tentorium with the falx cerebri to enter the confluence of the sinuses. The lateral sinuses (sinus transversus) pass horizontally outward from the torcular Herophili, along the temporal bone to the jugular foramen, where they terminate in the internal jugular vein. The occipital sinus (sinus occipitalis), the smallest of all the sinuses, commences at the margin of the foramen magnum on either side and passes backward to the torcular Herophili. The cavernous sinuses (sinus cavernosus) pass from the sphenoidal fissure along either side of the sella turcica to the apex of the petrous portion of the temporal bone, where they join the petrosal sinuses. They are crossed by fibrous bands or offsets of the dura mater, and inclose the pathetic, motor oculi, THE HEART AND VASCULAR SYSTEM. gH abducens and ophthalmic nerves, and the internal carotid artery, from which they are separated by the lining membrane. The circular sinus (sinus circularis) is formed by two small vessels (sinus intercavernous anterior et sinus intercavernous posterior) passing in front of and behind the pituitary body, and connecting the cavernous sinuses. The inferior petrosal sinus (sinus petrosus inferior), on each side, runs in a groove between the petrous portion of the temporal bone and the basilar portion of the occipital, connect- ing the cavernous sinuses with the lateral sinuses. The superior petrosal sinus (sinus petrosus superior), on each side, passes along the upper border of the petrous portion of the temporal bone, connecting the cavernous with the lateral sinuses* above. The transverse or basilar (plexus basilaris) is a small, straight sinus, connecting the inferior petrosal and cavernous sinuses. Sphenoparietal sinus (sinus sphenoparietal) runs in a groove on the under surface of the lesser wing of the sphenoid. It takes origin from the middle meningeal veins, runs inward. passes through a fold of dura mater, and usually receives the anterior temporal veins1 from the diploe. Cerebral Yeixs (vence cerebri). — The cerebral veins con- sist of two sets — the superficial, on the surface, and the deep, within the substance. They include: — Superficial (vence cerebri externa) — Superior cerebral veins (vv. cere- Inferior cerebral veins (re. cere- bri superior es) , bri inferiores). Deep — ■ Ventricular veins, or venae Galeni Veni corporis striati and choroid i //•. (inh iii ; it. cert bri interna} | , vein (v. choroidca). The cerebellar veins consist of the superior superficial cere- bellar veins (vv. cerebelli superiores) , and the inferior super- ficial cerebellar veins (vv. cerebelli inferiores) ; blood is brought from the interior to these superficial veins through the deep cerebellar veins. Diploic and Meningeal Yeixs. — The diploic veins (vence diploica), five in number, — frontal (v. diploica frontalis), an- terior temporal (v. diploica temporalis anterior), posterior tem- poral (v. diploica temporalis posterior), occipital (v. diploica occipitalis superior) -communicate with the sinuses of the brain and with the vein- of the dura mater, the scalp and orbit. 212 HUMAN ANATOMY. The meningeal veins (vv. meningeal) follow the course of the corresjDonding arteries, two accompanying each vessel through its course, and open into the sinuses of the dura mater. The great meningeal veins terminate either in the cavern- ous sinuses or by emptying into the internal maxillary vein. The ophthalmic superior vein (v. ophthalmica superior) commences at the internal canthus of the eye in an anastomosis with the facial, passes backward along the inner part of the orbit, through the sphenoidal fissure, to empty into the cavernous sinuses. The inferior ophthalmic vein (v. ophtlialmica inferior) arises from the veins of the eyelids and the lachrymal sac. It runs backward along the floor of the orbit and joins the superior to form a common trunk, or else it enters the cavernous sinus by a single opening'. The external jugular vein (v. jugularis externa), smaller than the internal jugular, is formed by the union of the pos- terior auricular with the temporomaxillary veins; or it may be formed by union with the facial. It descends beneath the platysma muscle from the angle of the jaw to the middle of the clavicle, where it terminates by emptying into the subclavian. It receives the following veins : — Anterior jugular, Suprascapular, Posterior jugular, Transverse cervical. The anterior facial vein (v. facialis anterior) commences as the angular at the internal canthus of the eye, where it anas- tomoses with the ophthalmic vein, and receives the frontal vein. About the angle of the jaw it usually ends in the internal jugular, but sometimes empties into the external jugular or unites with the temporomaxillary (v. facialis posterior), to form the common facial vein. The common facial vein (v. facialis communis), formed by the union of anterior facial and the anterior division of the temporomaxillary vein, runs backward beneath the sternomas- toid muscle, crosses the external carotid artery, and empties into the internal jugular. It receives: — Supraorbital {v. supmorbitalis) Labial, and superior palpebral, Submental, Nasal veins, Submaxillary, Inferior palpebral, Palatine. Buccal and masseteric, The temporal vein, which is known, as far as the zygoma, as the superficial temporal vein {vv. temporaries superficiales) , is THE HEART AND VASCULAR SYSTEM. 213 formed by the anterior temporal, which anastomoses with the frontal, and the posterior temporal, which anastomoses with the occipital. It penetrates the parotid gland and forms the tem- poromaxillary vein by uniting with the internal maxillary. It receives : — Articular veins, Transverse facial, Anterior auricular, Parotid. Middle temporal [v. temporalis media ) , The internal maxillary vein follows the course of the corresponding artery, and receives veins corresponding to the branches of that vesssl. Some of these branches form the pterygoid plexus. It passes backward and unites with the tem- poral vein to form the temporomaxillary. The temporomaxillary vein (v. facialis posterior), formed by the junction of the internal maxillary in part or whole with the temporal, passes through the parotid gland and receives the posterior auricular, to form the external jugular vein. The occipital vein (v. occipitalis) follows the course of the artery, and terminates, usually, in the internal jugular; occa- sionally, in the external jugular. The mastoid vein, passing through a foramen in the mastoid portion, connects it with the lateral sinus. Veixs of the Tongue and Throat. — The veins of the tongue empty into the internal jugular. The lingual veins (vv. linguales) begin near the tip of the tongue under the name of the ranine; they commence on the dorsum, sides and under sur- face, and pass backward, receiving the dorsal lingual vein, and also the sublingual vein, as well as two small veins (vena' comites), and terminate in the internal jugular. The pharyngeal (vv. pharyngece) , commencing in the pharyngeal plexus (plexus pharyngeals), receives branches from the brain, and terminates about on a level with the hyoid bone in the internal jugular. The superior thyroid (v. thyreoidea superioris) conveys the Mood from the larynx, trachea and thyroid gland to the interna] jugular. Veins of the Upper Extremity. — Superficial and deep. The subclavian vein (v. subclavia), the continuation of the axillary, unites with the interna] jugular to form the innomi- nate. In its passage over the first rib it is separated from the artery by the Bcalenue anticufi muscle. 214 HUMAN ANATOMY. Deep Set. — The deep veins accompany the corresponding arteries and form the venae comites, one on either side, except the axillary, which has a single vein. The valves are more numerous in the deep set. About the middle of the arm one of the brachial veins receives the basilic vein, one of the larger veins of the superficial set. The two brachial veins unite with the basilic to form the axillary vein, which in turn becomes the subclavian vein, receiving in its course beneath the clavicle the cephalic vein, a branch of the superficial set. Superficial Set. — The anterior ulnar vein (v. ulnaris ante- rior) commences on the inner surface of the hand and wrist, and terminates by uniting with the posterior ulnar (v. ulnaris posterior) or the median vein. The posterior ulnar vein com- mences on the inner back portion of the hand, anastomosing with the radial cutaneous vein, and ascends to the bend of the elbow, where it becomes the basilic vein; The basilic vein (v. basilica), from its formation at the bend of the elbow, ascends the inner side of the biceps, pierces the fascia, to join the brachial vein. The radial (v. radialis) commences on the radial, dorsal aspect of the hand, and at the bend of the elbow unites with the median cephalic, to become the cephalic. The cephalic vein (v. cepnalica) ascends along the outer border of the biceps, and above in the groove between the deltoid and pectoralis major, and passes beneath the clavicle, to end in the axillary. The median vein (v. mediana cubiti) receives the blood from the palmar surface of the hand and the front of the fore- arm, and usually divides into two branches — the median ce- phalic, passing outward to join the cephalic, and the median basilic, passing inward to join the basilic. The median basilic vein, the larger, is the one usually selected for phlebotomy. Veins of the Trunk. — The vena azygos major (v. azygos) commences in the abdomen, opposite the first or second lumbar vertebra, as a continuation upward of the right ascending lum- bar vein. It communicates with the right renal and the inferior vena cava. Through the lumbar veins it establishes communi- cation with the right common iliac vein. It passes through the aortic opening of the diaphragm, or through an aperture in the right crus, ascends on the dorsal vertebras, arches over the root of the right lung, and empties into the superior vena cava. It is in relation on its left side with the aorta, thoracic duct and esophagus. It receives THE HEART AND VASCULAR SYSTEM/' 215 Vena azygos minor ( v. hernia- Lower 8 or 9 intercostal veins, zygos), Right superior intercostal vein. Left upper azygos vein ( vena liemi- Right subcostal vein, azygos accessoria), Esophageal (vv. nsophagea) , Lower end of left superior inter- Right bronchial, costal vein, The hemiazygos veins are two in number : — The vena azygos minor (v. hemiazygos) arises on the left side, similarly to the azygos major, and passes through the left eras of the diaphragm and about the eighth dorsal vertebra crosses beneath the aorta to join the vena azygos major; Left upper azygos vein (vena hemiazygos accessoria) com- municates with the upper left intercostal vein, and terminates either in the vena azygos minor or the azygos major. The ascending or inferior cava (v. cava inferior), formed by the junction of the common iliac veins, at the sides of the fourth lumbar vertebra, ascends on the right of the aorta, grooves the posterior border of the liver, and passes through the quadrate foramen in the central tendon of the diaphragm, and ends in the right auricle, where it is provided with the Eusta1 chian .valve (valvula v. cavce inferioris [Eastachii]) . It receives Middle sacral, Renal veins (vv. renales) — the left, Lumbar (vv. lumbales), the longer, crosses in front of Spermatic (vv. spermatica') , from the aorta, spermatic plexus (plexus pam- Suprarenal (vv. suprarenales) , piriformis), Phrenic (vv. phrenica') , Ovarian (vv. ovmica) , from Hepatic (2 or 3) (vv. hepaticrr) . ovarian plexus (plexus arteries ovariece) . The portal vein (vena porta?), about three inches in length, is formed by the inferior and superior mesenteric, the gastric, splenic and pancreatic veins. It ascends in the right border of the lesser omentum to the transverse fissure of the liver, where it divides into the right and left. Its blood is distributed through the liver, mixing with the arterial blood from the hepatic artery, to be returned to the inferior cava by the hepatic veins. It receives The superior mesenteric (v. mesen- Cystic (v. cystica), terioa superior), Inferior mesentery (v. mesenterica Splenic (v. Uenalis), inferior), Ctoronary (v. coronaria ventric- iliglit gastroepiploic. uli) . 216 HUMAN ANATOMY. The portal vein and its branches are destitute of valves. Veins op the Vertebral Column. — 1. The dorsi-spinal veins (plexus venosi vertebrates externi) commence in an intri- cate network surrounding the vertebral arches and their proc- esses. They communicate with the intercostal, vertebral, intra- spinal plexus, lumbar, sacral and the superficial veins of the back. 2. The intraspinal or meningorachidian veins (plexus ve- nosi vertebrates interni) form an intricate plexus between the vertebras and dura mater within the spinal canal. They coiit sist of four longitudinal veins — two in front and two behind. The anterior longitudinal spinal veins (sinus vertebrates long- tudinales), two in number, extend along the sides of the verte- bral bodies and opposite the bodies communicate by transverse trunks, which, in their passage beneath the vertebral ligament, receive the diploic veins, or venae basis vertebras (v. basiverte- bralis), from the interior of the body. The posterior longitu- dinal veins, also two in number, smaller than the anterior, extend down the vertebral arches and are connected by trans- verse branches opposite the latter. They receive the veins from the spinal cord and its mem- branes. 3. The venae basis vertebrae (v. basivertebral^) return the blood from the bodies of the vertebrae into the anterior intra- spinal plexuses. The venae medulli spinalis (vv. spindles) are the essential veins of the cord, situated between the arachnoid and pia mater. Common Iliac Veins (v. iliaca communis) . Formed by the internal and external iliac veins uniting opposite the sacroiliac articulation, pass beneath the right com- mon iliac artery to a point a little to the left of the body of the last lumbar vertebra, where they unite to form the inferior vena cava. The right is shorter and more vertical in its course, and both are without valves. The internal iliac or hypogastric vein (v. iliaca in- terna or v. hi/pogastrica) corresponds to the distribution of the corresponding artery. It receives the following venae comites: — Gluteal (vv. glutece), Lateral sacral (vv. sacrales lat- Sciatic, erales) and Obturator (v. obturatoriw) , Middle sacral; Internal pudic (vv. pudendce in- ternee), THE HEART AND VASCULAR SYSTEM. 217 and the following plexuses: — EZS2SS? f»—« EST- }.■»*-* The lateral and middle sacral form a small plexus — the plexus saeralis. The veins of the rectum, bladder, and generative organs anastomose freely and form three plexuses : — 1. Hemorrhoidal plexus (plexus hamorrhoidalis) encircles the lower part of the rectum, communicates with the sacral and prostatic plexuses, and veins from it join the inferior mesen- teric, internal iliac and. pudic veins. 2. Vesica prostatic plexus surrounds the membranous por- tion of the urethra, neck of bladder, prostate body and seminal vesicles. It communicates behind with the hemorrhoidal. 3. Uterovaginal plexuses (plexus utero vaginalis) . — These correspond in the female with the prostatic, and communicate with the vesical and hemorrhoidal plexuses, the ovarian, pudic, and. through the uterine veins join the internal iliac veins. During pregnancy these veins or plexuses become greatly distended and enlarged, forming the uterine sinuses, but retain a straight course. The pudic vein follows the same course as the artery, com- municates with the prostatic and hemorrhoidal plexuses, and terminates in the internal iliac vein. In the female it originates from the clitoris and perineum, and communicates with the vaginal plexus. The dorsal vein of the penis (v. dorsdlis penis) returns the blood from the body of that organ. It commences as two venae comites of the dorsal artery, which unite on the dorsal surface of the root of the penis, perforates the triangular ligament, and again divides to terminate in the prostatic plexus. The veins of the corpus cavemosum emerge at the lower groove, and turn round their outer side to join the dorsal vein. The dorsal vein of the clitoris has a corresponding origin and course, and empties into the vaginal plexus. The EXTERNAL iliac vein (v. iliaca externa,), the con- tinuation of the femoral, lies internal to the artery beneath Pouparfs Ligament, and joins the interna] iliac opposite the sacroiliac symphysis, to form the common iliac vein, it runs along the brim of the pelvis. It receives at its commencement the \ ni Cortical structure. (d) Curved tubule, J (j) Straight collecting tubule — descends through the medullary structure to open by an orifice in a renal papilla. The straight, collecting or receiving tubes converge as they descend, and unite and reunite until there are about one dozen, but they vary from eight to twenty, one opening on the summit of each papilla. In the cortical portion they form groups, or medullary rays, giving the appearance of conical masses in the cortical substance with their apices toward the periphery — the so-called pyramids of Ferrein. The tubules are composed of a basement membrane, lined with pavement epithelium. In the neck the epithelium becomes continuous with that of the Mal- pighian capsule, and at the glomerule the wall is reflected upon the inclosed tuft, or Malpighian corpuscle. In other situations the epithelium varies much in shape and size. The renal arter- ies subdivide at the hilus, cross the sinus, and enter the renal substance between the renal papillae. They terminate in the renal glomerules, or Malpighian corpuscles. Each one of these remarkable convolutions of capillaries is about one one-hun- dredth of an inch in diameter and inclosed in a pouch-like dilatation of the uriniferous tubules or Malpighian capsule, or capsule of Bowman. The efferent vessels form, together with others, a capillary network between and around the uriniferous tubules. They are made up of three sets: (a) the plexuses around the tubuli contorti; (&) the veins beneath the capsule; and (c) the plexuses about the apices of the pyramids of Mal- pighi. These form the vence interlobular 'es, which join the vence rectce, and together form the renal veins, vence propria? renales. Those in the sinus unite together to form the renal vein, and open into the inferior vena cava, the left crossing in front of the abdominal aorta. THE GENITOURINARY APPARATUS. 269 T and 8. Ascending limb of lleule B loop tuba. Subcapsular layer with- out Mai pigbian cor- puscles. 12. First part of col- lecting tube. 11. Distal convoluted tubule. A/ A. Cortex. 10. Irregular tubule. 3. Proximal convo- luted tubule. 9. Wavy part of as- cending limb. 2. Constriction or neck. 4. Spiral tubule. 1. Malpighian tuft surrounded by Bowman's capsule. 8. Spiral part of as- cending limb of Uenle's loop. B. Boundary Zone. 6. Descending limb of Uenle's loop tube. 15. Tubule of Bellini. 15 8. Honle s loop. ■ C. Papillary Zoijk. Diagram of urinlferous tubules: i. Malpighian capsule; 2, proxi- mal convoluted tubule; 2 b, distal convoluted tubule; '■'•. descending limii of Henle's loop; i, ascending limb of Uenle's loop; 5, irregular tubule; 6, collecting tube; a, apex of pyramid; 6, base of pyramid; C, cortical portion. 270 HUMAN ANATOMY. The ureter commences as a compressed pouch, the pelvis. Within the sinus the pelvis divides, and subdivides into several small funnels, or calices\, each one of which receives one or two projecting renal papilla?. The calices, pelvis and ureter are all of the same structure, consisting of a fibrous and unstriated muscular and a mucous coat. The fibrous coat (tunica adven- titia) becomes continuous with the capsule above and below in the fibrous structure of the bladder. The muscular coat (tunica muscularis) ceases at the base of the renal papillae, but the mucous membrane becomes continuous with that of the urinif- erous tubules. Its epithelium is of the transitional variety, consisting of several layers of cells of varying shapes, resting upon a basement membrane. The nerves are derived from the renal plexus of the sym- pathetic, formed by branches in the semilunar ganglion, the solar plexus and the lesser and smallest splanchnic nerve. The lymphatics communicate with the lumbar glands. The ureter proper is a musculomembranous tube, sixteen to eighteen inches in length, about the diameter of a goose quill, extending from its origin in the pelvis of the kidney to the basal or posterior angle of the vesical trigone, which it enters by passing obliquely through the muscular and mucous walls of the bladder. Course- — It descends upon the psoas magnus muscle, be- neath the peritoneum, being crossed by the spermatic vessels. About the first division of the sacrum it crosses the external or common iliac artery, passing behind the sigmoid flexure on the left and the ileum on the right side, to enter the pelvis, where it reaches the bladder within its posterior false ligament. In the male it passes behind the vas deferens. In the female it crosses the uterine artery one-third to one- half inch from the cervix uteri and passes along the side and upper part of the vagina to the bladder (vide Vagina). Muscles of the Ureters. — Two oblique muscles. Origin. behind the orifices of the ureters ; insertion, into the middle lobe of the prostate gland; action, they guard the orifices of the ureters, and prevent the reflux of urine. Suprarenal capsules (vide Ductless Glands). THE BLADDER. The bladder (vesica urinaria) is a musculomembranous sac, the reservoir for the urine, situated in the anterior part of the pelvis, behind the pubes, in front of the uterus and vagina in the female, and the rectum in the male. THE GENITOURINARY APPARATUS. •>] \ It measures, moderately distended, five inches in length, three in width, and holds about one pint. It consists of a body, summit. base and neck. The body {corpus vesica) is partially invested with peri- toneum behind, but in front it is wanting, the body heing in relation with the symphysis pubis, triangular ligament, and internal obturator muscles. The summit, or apex (vertex vesicce), is rounded, and directed upward and forward, being connected to the umbilicus by the urachus, the remains of the fetal allantois, and also by the obliterated hypogastric arteries, one on either side. The space hounded by the pubic surface and the sides of the bladder uncovered by peritoneum, filled by the rectovesical fascia, is known as the space of Eetzius. The base, or fundus (fundus vesicae), in the male is sit- uated upon a triangular space upon the second portion of the rectum, it- base formed behind by the rectovesical fold, its apex by the prostate gland, and its sides by the vas deferens and vesiculae scminales. In the female it is situated in contact with the cervix uteri and anterior wail of the vagina, adhering closely to the latter. It- upper portion has a peritoneal covering. The neck, or cervix (collum vesicce), is the contracted por- tion, coin unions with the urethra. It is encircled in the male by the prostate gland, and is directed obliquely when the in- dividual is in the erect posture. The ligaments of the bladder consist of five true ligaments derived from the pelvic fascia and the urachus; and five false ligaments derived from the peritoneum. The true ligaments are: — Two anterior (puboprostatic) (ligamenta puboprostatica) , two folds of rectovesical fascia passing from the pubic symphy- sie to the- cervix and upper surface of prostate gland; Two lateral folds of rectovesical fascia connecting the lat- eral surfaces of prostate gland with sides of base of bladder; The urachus {ligamentum umbilicale medium), an obliter- ated fetal structure, passing as a fibromuscular cord from the apei of tin; bladder to the umbilicus. The false ligaments are: — Two posterior ligaments (plicce rectovesicales) are peri- tonea] fold.- parsing between ^ides of rectum in male, sides of uterus in female, to posterolateral surface of bladder, inclosing obliterated hypogastric arteries, ureters, vessels and nerves; 272 HUMAN ANATOMY. Two laterals (Ugamenta umbilicale laterales) are folds of peritoneum passing from iliac fossge to sides of bladder; The superior (plica umbilicalis media) is a single peri- toneal fold inclosing the obliterated hypogastric arteries from apex of bladder to umbilicus. The structure consists of four coats, a serous, muscular, submucous and mucous. The serous coat (tunica serosa) is derived from the perito- neum. It covers the posterior surface from the ureters to the summit, and is reflected from the sides to the walls of the pelvis and abdomen. The muscular coat (tunica muscularis) is made up of unstriated fibers arranged in three layers: — (a) Longitudinal external layer (stratum externum), named the detrusor urines muscle; (b) Circular middle layer (stratum medium), forming at the neck the sphincter vesicae; (c) Longitudinal internal layer (stratum internum). The submucous or cellular coat (tela submucosa) , made up of fibrous and elastic tissue, supports the mucous coat and unites it with the muscular. . The mucous membrane (tunica mucosa) is smooth and of a pale rose color. Its epithelium is continuous with that of the ureters and pelvis of kidneys; its superficial layer, large, tessel- lated, polyhedral cells ; its deep layer of club-shaped and spindle- shaped cells. The inner surface of the base presents a triangular space, the vesical trigone, or trigonum vesica, apex in front, formed by the orifice of the urethra, its basal angles behind about two inches apart, and each about one and one-half inches behind the urethral orifice, formed by the orifices of the ureters: — U u u 2" The uvula vesica is an elevation of mucous membrane pro- jecting from the floor near the apex of the trigone into the orifice of the urethra. The arteries are the superior, middle and inferior vesical, with branches from the obturator and sciatic in the male, and branches from the vaginal and uterine in the female. The veins from the vesicoprostatic empty into the internal iliac vein. THE GENITOURINARY APPARATUS. 273 Lymphatics follow the course of the vessels and enter the lumbar glands. The nerves to the base and neck are from the third and fourth sacral, and to the summit, from the hypogastric plexus of the sympathetic. GENERATIVE APPARATUS. Male Organs. The male organs of generation consist of the testes, vasa deferentia. vesiculae seminales and penis. The testicles (testes) are two glandular bodies which secrete the spermatic fluid, and are suspended by the spermatic cords within the scrotum, the left a little larger and lower than the right. They are oval, compressed laterally, measure one and a half to two inches in length, one inch in breadth, one and one-fourth inches in their anteroposterior diameter, and weigh from six to eight drachms. They each consist of a body, or testicle proper, and an epididymis. The latter consists of a head or globus major {caput apididymidis) , body or central portion (corpus epidid- ymides) and tail or globus minor (cauda epididymidis) . The scrotum consists of the skin marked in the median line by the raphe (raphe scroti), formed by the union of the genital folds, and the dartos, a reddish, contractile tissue which sur- rounds the testes and extends from the raphe to the under sur- face of the penis, forming the septum scroti, which divides it into two cavities for the testes. The spermatic cord (funiculus spermaticus) is made up of the following structures: — Externa] spermatic, or intercolumnar fascia; Cremaster muscle; Internal spermatic fascia, or fascia propria; Yas deferens; Spermal ic artery •. Deferentia] artery from superior vesicle; Cremasteric artery, from epigastric; Vasa Bpermatica; Lymphatic vessels; Spermatic plexus of nerves [from renal and aortic plexuses]; Genital branch of genitocrural; Scrotal branch of the ilioinguinal, Tt commences al the internal abdominal ring, passes ob- liquely through the inguinal canal, emerges at the external abdominal ring, descends into the Bcrotum, and terminates at the posterior border of the testes. 18 274 HUMAN ANATOMY. The external spermatic or intercolirmnar fascia is derived from the aponeurosis of the external oblique muscle. This mus- cle arises within the inguinal canal from Poupart's ligament and pubic spine, and descends along the cord in loops. Its fibers are striated. The cremaster muscle, or cremasteric fascia {fascia cremas- terica) or middle spermatic fascia, is derived from the lower border of the internal oblique and transversalis muscle. The internal spermatic fascia, or fascia propria, is the infundibuliform process of the transversalis fascia. The proper coverings or tunics of the testicle are three — tunica vaginalis, tunica albuginea and tunica vasculosa : — The tunica vaginalis (tunica vaginalis propria testis) is a serous membrane surrounding the testes, and at the posterior portion is reflected on itself to form a sac. The outer portion of the latter is loosely attached, except at the lower part of the testicle, where the gubernaculum testis binds it down. The tunica albuginea is a dense, white fibrous structure, continuous at the upper part with a similar structure on the epididymis. At the back part it projects into the glandular substance of the testicle to form the mediastinum testis (corpus Tliglimori) , from which numerous imperfect septa, called trabecule (septula testis) diverge, and with similar cords from the tunica albuginea serve to maintain the shape of the gland and divide it into lobules. The tunica vasculosa (pia mater testis) lies within the tunica albuginea, and is the source and termination of the blood-vessels to the secretory substance of the gland, resembling the pia mater of the brain. The glandular structure of the testis is divided into several hundred (two hundred and fifty to four hundred) pyramidal lobules (lobuli testis), each one of which consists of from one to three tubuli seminiferi contorti. At the conical extremity of the lobules the tubules end in straight tubes, vasa recta, which enter the mediastinum to form the plexus retiformis. From the upper part of this the vessels unite into from twelve to twenty vasa efferentia, which pass out of the testicle to the epididymis. Within the epididymis these efferent canals form a series of spermatic cones, the coni vas- culosi. These end in a coarse, convoluted tube, about twenty feet in length, forming the body and tail of the epididymis, and ending in the spermatic duct, or vas deferens (ductus deferens). These tubes are lined with columnar ciliated epithelium. THE GENITOURINARY APPARATUS. 275 Fig. 98. on through bladder, urethra, and testicle: A, urethral orlfict /•' seminal vesicle; O, colliculus seminalis or verumontanum; I), prostatic utricle; /.', orifice of ejaculatory duet; F, suspensory ligament; ff, Cowper's gland; //. bulb of corpus spongiosum; /, corpus spongiosum; ./, urethra; K, seminal plexus; /-. testicle; l/, peritoneal fold; V space oi Retzius; O, corpus cavernosum; P, glans penis; (), prepuce. 276 HUMAN ANATOMY. The vas aberrans of Haller is a spermatic cone joining the epididymis, but unconnected with the testicle. The hydatids of Morgagni are one or two pedunculated bodies attached to upper part of testis or head of epididymis, and supposed to be the remains of Muller's duct. Structure of the Tubules. — The tubuli seminiferi are mi- nute convoluted tubules, closed at one extremity, consisting of a hyaline membrana propria lined with several layers of epithelial polyhedral cells, the seminal cells, from which the seminal or spermatic fluid is directly elaborated. These cells undergo a process of indirect division (Icaryo- hinesis) to form the spermatoblasts, which are arranged into bundles, and are converted into spermatozoids, the nuclei becom- ing the head, and the tail, or ciliary appendages, being after- ward developed. These cells are constantly cast off and replaced. Descent of the Testes. — In early fetal life the testes are placed at the back part of the abdomen, below and in front of the kidneys, and behind the peritoneum. About the third month a peculiar structure, the gubernacu- lum testis, appears, attached to the lower end of the epididymis, and extending as a cord to the bottom of the scrotum. It is supposed to cause the descent of the testicle. It reaches its full development between the fifth and sixth month, at which time the testicle reaches the iliac fossa. It enters the internal abdom- inal ring by seventh month, and the scrotum by the eighth month, carrying before it a fold of peritoneum, which is after- ward shut off, forming the tunica vaginalis testis. Other cover- ings of the testicles are also derived in this manner. In the female a structure similar to the gubernaculum forms the round ligament. The vas deferens has three coats — (1) an external fibrous coat {tunica adventitia), (2) an unstriated muscular coat (tunica muscularis), and (3) a lining membrane (tunica mu- cosa) of columnar epithelium. It is about one and one-half feet long and one line in diameter. From the tail of the epididymis it ascends in the cord, forming an important part, and lying behind the blood-vessels. It ascends through the inguinal canal, and at the internal ring descends on the bladder, crosses the ureter and obliterated hypogastric artery, and runs forward to form with the duct of the seminal vesicle the ejacu- latory duct. The walls of the vas are very thick and the canal very small. Its course for the most part is straight, but beneath the bladder THE GENITOURINARY APPARATUS. 277 it becomes enlarged, tortuous, arid more capacious, but again contracts near its termination. The arteries of the cord are: — The spermatic, to the testicle from the aorta: The vas deferens, or deferent artery, from the superior vesical; The cremasteric, from the deep epigastric. CREMRS.TER MUSCLE. Fig. 99. The testicle and epididymis: .4, testis; B, epididymis; 0, tunica vaginalis; D, vas deferens; /•;, spermatic artery and veins; F, artery of cord. The veins from the testis — spermatic reins — unite with branches from the epididymis to form the pampiniform plexus, which terminates on the Lefl side passing beneath the sigmoid re in the left renal vein, on the right side in the inferior Vlia I The lymphatics are numerous, and terminate in the lum- bar elands. 278 HUMAN ANATOMY. The nerves are from the sympathetic system, and form the spermatic plexus. The arteries of the scrotum are derived from the Cremasteric branch of epigastric; Superficial external pudic, from the femoral; Deep external pudic, from the femoral; Superficial perineal, branch of internal pudic, from internal iliac. The seminal vesicles (vesiculce seminales) are two mem- branous pouches, closely adhering to the under surface of the bladder. Their size varies, but they usually measure two and one- half inches in length, one-half inch in breadth, and two to three lines in thickness. Each consists of a tube closed at one end, about four to six inches long, the diameter of a quill, convoluted into a mass. Its structure is analogous to that of the ducts, but thinner. The ejaculatory ducts (ductus ejaculatorii) , two in num- ber, are formed by the union of the vasa deferentia with the duct of the vesiculge seminales. Each duct is three-fourths to one inch in length, and passes through the prostate gland to terminate at the margin of the sinus pocularis by a slit-like orifice. The arteries are derived from the inferior vesical and mid- dle hemorrhoidal. The veins and lymphatics correspond to, and accompany the arteries, and the nerves are from the hypogastric plexus of the sympathetic. The seminal vesicles are not only reservoirs, but secrete a fluid to dilute the spermatic liquid. The semen is a viscid, whitish liquid, composed of a color- less liquid, the liquor seminis, and the spermatozoa and seminal granules. The seminal granules are about one-four-thousandth of an inch in diameter. The spermatozoa are the essential elements, and consist of an oval head, a body or middle piece and a tail. The penis is composed of three columnar bodies, two cor- pora cavernosa and one corpus spongiosum, invested with skin, and filled with a peculiar, vascular, sponge-like structure. It is attached to the pubic arch and symphysis by its root (radix penis), has a free expanded extremity (glans penis) or head, and an intermediary portion, the body (corpus penis). The upper surface is the dorsum, or back. The glans is a blunt, cone-like body, expanded at its base, the corona, and attached by its cervix, or neck (collum glandis). THE GENITOURINARY APPARATUS. 379 The summit has a slit-like aperture, the meatus (orifnum urethra externum), or orifice of the urethra. The skin adheres loosely to the organ, and lias a loose, cup- like fold over the glans, called the prepuce {praspiitmm) , after which it is reflected into the cervix and glans, becoming con- tinuous with the mucous membrane of the urethra at the meatus, to which it is attached below by a hand or bridle, the franum (frenulum prceputii). Ttailem Vajl/tab* Tunic* Afluji a M Fig. 100. Vertical section of testicle. (After Gray.) The ddn covering the glans resembles mucous membrane, La very rascular and sensitive, hut about the cervix and corona ],;l_ Qumeroufl preputial glands— gUndula Tysom odonfen (glandndce Tysonii odoriferce), which Becrete the smegma. The superficial fascia is thin, free from adipose tissue, and continuous with that of the abdomen and scrotum. The suspensory Ligamenl (ligamentum suspensorvum perns) passes from the fronl of the pubic symphysis to the tunica al- Unnea of the corpora cavernosa. The fundiform ligament, or li^mentum fundiforme penis, was formerly described as the 280 HUMAN ANATOMY. suspensory ligament; this is not strictly correct, although the latter ligament is suspensory in character. The ligamentum fundiforme penis passes to the penis from the lower part of the linea alba, and at its penile attachment divides into two por- tions, between which pass the dorsal vessels and nerves. The corpora cavernosa (corpora cavernosa penis), or cav- ernous bodies, constitute more than two-thirds of the bulk of the organ. They spring as crura from the rami of the ischium and pubis, swell out into the bulb of the cavernous body, or corpus cavernosum, join each other in the median line, to ter- minate in a conical extremity, which receives the glans. They are grooved above for the dorsal vein, and below for the corpus spongiosum. They each have a strong outer fibrous membrane, which, on uniting, forms a thick partition, complete behind, but incom- plete and comb-like in front — septum pectvniforme. From this septum fibroelastic bands diverge in all direc- tions and form trabecules, which, with the blood-vessels occupy- ing their intervals, form the erectile tissue of the corpora cavernosa. The corpus spongiosum (corpus cavernosum urethrce) com- mences as. a bulb below the crura and in front of the triangular ligament. It expands anteriorly into the glans penis, which fits upon the conical termination of the corpora cavernosa. The urethra enters above and anteriorly to the bulb (bulbus urethra), and traverses its entire length to the summit of the glans, opening at the meatus. The corpus spongiosum consists of an external fibrous coat, thinner and more elastic than that of the corpora cavernosa, and an interior erectile tissue. A thin, muscular layer lines the external fibrous coat, and another is found beneath the mucous membrane of the urethra. The corpora cavernosa get their blood from the arteries of the corpora cavernosa, and branches from the dorsal artery of the penis, from the internal pudic. The corpus spongiosum is supplied by the artery of the bulb. The arteries of all three bodies terminate finally in the erectile tissue. Many arterial branches, especially at the root of the penis, form short convolutions — the helicine arteries (arteries helicince), which terminate in finer vessels, and sub- sequently open into the spaces of the erectile tissue. Others open directly. THE GENITOURINARY APPARATUS. 281 From these spaces the veins begin; some wind around the Bide of the organ to the dorsal vein, while others pass under the pubis to join the prostatic plexus. The lymphatics are numerous; the superficial join the inguinal glands, the deep join the lymphatic plexus about the prostate and other deep lymphatics of the pelvis. The male urethra (urethra virilis) is the common canal for the emission of semen and urine, extending from the neck of the bladder (orificium urethra internum) to the meatus urinarius (orificium urethral externum). It is from eight to nine inches in length, and consists of three portions — the pro- static, membranous and spongy. The prostatic portion (pars prostatica) extends from the neck of the bladder to the anterior border (apex) of the prostate gland. It is one and one-quarter inches in length, and is the widest and most dilatable part. Its floor is raised, and presents: — Yerumonatum, or caput gallinaginis (cotticulus seminalis), an elevated ridge; Prostatic sinus (utriculus prostaticus), on each side of the verumontanum ; Orifices of the prostatic ducts, in the floor of the prostatic sinuses; Sinus pocularis, a depression in the median line in front of the veruniontanum, presenting the slit-like openings of the ejaculatory ducts (ductus ejacututorii). This cul-de-sac is one- quarter of an inch in length. It is homologous with the uterus, and has received the name of utricle, or uterus masculinus. The membranous portion (pars membranacea) is about three-quarters of an inch in length, extending from the apex of the prostate gland to the corpus spongiosum above and in advance of the hull). It passes out of the pelvis beneath the symphysis pubis, traversing the triangular ligament. It is the least di lat ahle portion and has four coats: — Fibrous, continuous with both layers of the triangular liga- ment : erectile, continuous with that of the spongy body; an iinstriated muscular layer and a mucous coat, The spongy portion (pars cavernosa) extends from the membranous portion through the corpus spongiosum to the meatus urinarius on the summit of the glans. The portion within the hulh has received Ihe name of tudh- ous portion of the urethra. The spongy portion diminishes gradually in size to near the orifice, where ii suddenly dilates into the fossa navicularis (fossa navicularis urethra: \_Mor- 282 HUMAN ANATOMY. gagni] ) , contracting again at the meatus, the narrowest part of the urethra. The mucous membrane is provided with columnar epithe- lium, except near the meatus, where it is tessellated, a fibro- elastic submucous layer with unstriated muscular fibers, and numerous minute racemose glands, the glands of Littre (gland- ules urethrales) . These latter open, forward into the urethra by good-sized orifices, especially one in the upper part of the fossa navicularis, called the lacuna magna. Into the bulbous portion of the urethra the ducts of Cowper's glands open. The prostate gland (prostata) is a glandular body which resembles in size and form a chestnut, and surrounds the first portion of the urethra between the neck of the bladder and the triangular ligament. It measures one and one-half inches in length and breadth and three-quarters in depth. Its weight, about six drachms. Its flat under surface rests on the rectum. It has two lateral lobes (lobus dexter et sinister), and one middle lobe (lobus medius), which corresponds in position to the vesicle uvula, and is held in position by the anterior liga- ments of the bladder, by a portion of the deep perineal fascia, and of the levator ani muscle. It is perforated by the urethra and the common seminal ducts. Its structure consists of a mass of flbromuscular (unstri- ated) tissue with imbedded follicular pouches, the whole inclosed in a firm fibrous capsule, continuous in front with the trian- gular ligament, behind with the posterior layer of the deep perineal fascia. The muscular fibers are longitudinal and circular, the lat- ter surrounding the urethra, continuous behind with the blad- der, in front with the fibers about the membranous portion. The glands open into the floor of the prostatic sinuses by twelve to twenty ducts. The arteries are from the vesical, hemorrhoidal and inter- nal pudic. The veins enter into the formation of the prostatic plexus, receive the dorsal vein of the penis, and empty into the internal iliac vein. The nerves are from the hypogastric plexus. Cowpep/s glands, or suburethral glands (glandulce bulbo- uretlirales) , are two small lobular bodies, about one-quarter of an inch in diameter, inclosed between the two layers of the deep fascia, situated behind the bulb of the corpus spongiosum, below the membranous portion of the urethra. They are racemose THE GENITOURINARY APPARATUS. 283 gland- and empty their secretion by a long duct (ductus excre- iorms) into the bulbous portion of the urethra. Female Organs. The female organs of generation are divided into the exter- nal (partes genitales external muliebres), termed the vulva, or pudendum, consisting of the mons veneris, labia majora, minora, clitoris, meatus urinarius and orifice of the vagina ; and the internal (pars genitales internee muliebris) , consisting of Fig. 101. Internal female genitals. the uterus, ovaries, Fallopian tubes and vagina, with certain accessories. The uterus is a hollow, muscular organ for the reception of the fecundated ovum and the development and expulsion of the fetus. The virgin uterus holds an oblique anterior position in the pelvis, being supported by attachments to the vagina, rectum, bladder and sides of the pelvis. It is in contact with the bladder in front, the rectum behind, and above the small intestine. Its shape is pyriform, compressed from before backward, and measures three inches in length, two in breadth, one in thickness, and weighs from one to one and a hall' ounce-. J l consists of a nock, fundus and body. 284 HUMAN ANATOMY. The month, os uteri, or os tineas (orificium uteri exter- num), open into the vagina, being protected by two lips; an anterior (labium anterius) thick lip, and a posterior (labium posterius), long and narrow. The cavity of the uterus (cavum uteri) is triangular V from side to side, but a mere slit from before backward, and measures two and one-half inches in its longitudinal diameter. The two upper angles are prolonged to communicate with the Fallopian tubes ; the lower angle forms the ostium internum uteri (orificium internum uteri), communicating with the cav- ity of the cervix. Its structure consists of three coats : — Serous coat (tunica serosa), derived from the peritoneum and investing all but the lower anterior quarter ; Muscular coat (tunica muscularis), about one-half inch thick, composed of unstriated muscular fibers arranged into three layers; Mucous coat (tunica mucosa), has numerous tubular folli- cles, is lined with ciliated columnar epithelium, and has no sub- mucous coat to connect it with the muscular coat. The mucous membrane of the cervix is thrown into folds, or ruga}, which assume on the anterior and posterior walls a branched arrangement, or arbor viice uterina. It is lined by squamous epithelium, and presents numerous follicular glands — the ovula of Naboth, or glandulce Nabothi. The arteries are branches of the ovarian from the aorta, and the uterine from the internal iliac, remarkable for their tortuosity and anastomoses. The veins form plexuses or uterine sinuses, the branches of which correspond to the uterine arteries and terminate in the uterine plexuses. The lymphatics are very numerous and terminate in the lumbar and pelvic glands. -The nerves are from the ovarian and hypogastric plexus of the sympathetic. The ligaments of the uterus are foMs of peritoneum arranged into four pairs: — Two anterior, or vesicouterine, passing one on either side from the posterior surface to the cervix uteri ; Two posterior or rectouterine {plica rectouterinw) , passing be- tween the sides of the rectum and uterus, and inclosing a cul-de-sac, the rectovaginal pouch, or Douglas's pouch; A prolongation of this ligament upward to the second sacral verte- bra, with some unstriated muscular fibers derived from the uterus and vagina, forms the so-called uterosacral ligaments j THE GENITOURINARY APPARATUS. 285 Two lateral, or broad [Ugamentum latum uteri), extending from the uterus to the sides of the pelvis, dividing it into two portions, and inclosing the Fallopian tubes, ovary, ovarian ligament, uterine blood- vessels, lymphatics and nerves, and some unstriated muscular fibers; Two round ligaments [Ugamentum teres), are cords of unstriated muscular and fibrous tissue, extending from the side of the fundus uteri to the inguinal canal, where they are lost in the subcutaneous tissue of the pubes and labia majora. The canal of Nuck, a pouch of peritoneum, incloses the ligament in the young subject, but is usually obliterated later. „ UreMiro. Fig. 102. Female organs of generation. The ovaries correspond to the male testicles, and are sus- pended behind the broad ligament inclosed in its posterior layer. .They are largest from puberty to adult age, and measure one and one-half inches in length, three-quarters of an inch in width, and one-third of an inch in thickness, and weigh from one to two drachms. The inner border is attached to the fundus uteri by the ova/nan ligament, and its outer border to the fim- briated extremity of the Fallopian tube by a cord (tubo-ovarian ligament). 286 HUMAN ANATOMY. The structure of the ovary is made up of a reddish, spongy stroma, well supplied with blood-vessels, containing numerous ovisacs, or Graafian vesicles (folliculi obphori vesiculori [Graafi]), inclosed in a serous covering derived from the peritoneum. The serous coat differs from the peritoneum in having a single layer of columnar cells, the germinal epithelium of Waldeyer. The stroma of the ovary is a vascular tissue, composed of many spindle cells and a small portion of connective tissue. Upon the surface of the ovary, this tissue becomes niore firmly organized and tenacious; and was formerly, but erroneously, designated the tunica albuginea. The Graafian vesicles, or ovisacs, containing the human ova, vary in size from microscojfic bodies to one-quarter of an inch in diameter, and are most abundant in the perijDhery. In structure they consist of a fibrous coat — the ovicapsule — lined by a basement membrane — membrana propria — and a layer of cells, the membrana granulosa. The interior is filled with a transparent 'albuminous fluid, liquor folliculi. Tbe epithelial lining nearest the ovary presents an accumu- lation of cells — the germinal eminence, or discus proligerus, within which is the ovum or egg. Discharge. — The Graafian vesicles approach the surface of the ovary and burst, their contents passing into the opened aper- ture of the tube, the fimbriated extremity apparently applying it to the region of the bursting ovisac.1 The rupture occurs periodically and corresponds to the menstrual flow. The ovum is surrounded by the tunica vasculosa and some additional epithelial structures (retinacula). It is one-tenth line in diameter, and represents all the elements of an organized cell, being composed of a Cell-wall, or vitelline membrane, or zona pellucida; Cell-contents, or vitellus ; Nucleus, or germinal vesicle ; Nucleolus, or germinal spot. At the seat of the rupture the walls collapse, and the vesicle immediately becomes filled with a blood-tinged fluid, which, together with the hypertrophy of the walls, forms the l The fimbriated extremity contains no erectile tissue, and the ovum probably falls into the peritoneal cavity and is swept into the tube by the action of the cilia. (Spigelberg.) THE GENITOURINARY APPARATUS. 287 false corpus luteum, which remains and develops for two to three months and gradually disappears. Under the influence of pregnancy it enlarges for some time, forms the true corpus luteum of pregnancy, and disappears two lo three months after parturition. The arteries are the ovarian from the aorta. The veins form an intricate plexus from which emerge ves- sels corresponding to the* arteries; they form a plexus near the ovary — the pampiniform plexus — which communicates with the uterine and terminates as in the male. The lymphatics are numerous and of large size in the FIG. 103. Section of an ovary: e, germ epithelium; 1, large-sized follicles; 2, 2, smaller-sized follicles; 0, ovum within a Graafian follicle; r, r, blood-vessels of the stroma; (J, cells of the membrana granulosa. impregnated uterus, and terminate in the lumbar and pelvic glands. The nerves are from the ovarian and thoracic aortic plexuses of the sympathetic. The Parovarium, epo6phoron or organ of Etosenmuller, con- sists of a series of tortuous tubes extending from the ovary to a main transverse trunk, near the Fallopian tubes; and is the re- main- of the Wolffian body of embryonic Life, and corresponds to the origin of the epididymis in the male. Tin- Fallopian tubes, or oviducts {tuba ulcriiia FallopU), are the passageways for the ovum from (he ovaries to the uterus. 288 HUMAN ANATOMY. They are trumpet-shaped tubes about four inches long, with the largest extremity outward. Its ovarian extremity is fimbriated; hence its name, fim- briated extremity. It is also called morsus diaboli, from its supposed erectile action. One of these fimbriae extends along the border of the broad ligament to the outer extremity of the ovary, forming the tubo- ovarian ligament {fimbria ovarica). The ovarian orifice — ostium abdominale, or pavilion (os- tium abdominale tubce uterince) — is much larger than the uterine, or ostium internum (ostium uterinum tubce). The tube has three coats: — Serous (tunica serosa), derived from the peritoneum; Fibromuscular (tunica muscularis, stratum longitudinale et stratum circular e) , from the uterine walls; Mucous coat (tunica mucosa), with ciliated columnar epi- thelium, continuous with the uterine. The remains of the duct of Mtiller form the hydatid of Morgagni (appendices vesiculosi) , a small vesicle, attached by a long pedicle near the fimbriated, extremity. The arteries are from the ovarian, the veins follow the same course, and the lymphatics and nerves are the same as the ovarian and uterine. The vagina is a cylindrical membranous canal extending from the vulva to the uterus. It is formed by the coalescence of two symmetrical tubes in fetal life. Relations. — It is in relation in front with the base of the bladder and urethra, behind its upper fourth with Douglas's pouch, its lower three-fourths connected loosely with the rectum, and laterally with the broad ligaments, pelvic fascia and levator ani muscles. On the posterior wall just below the cervix uteri the -ureters approach each other, leaving a space of only three-quarters of an inch between them, an important factor in lithotomy. In the virgin adult it measures about four inches in length, one inch in width, somewhat larger in its middle. Its anterior (paries anterior) and posterior walls (paries posterior) are in contact. Its lower orifice, or entrance (orificium vagina), is con- stricted by a crescentic or circular fold, of mucous membrane, the hymen. The upper extremity, or fundus, receives the cervix uteri, extending higher up posteriorly (five to six inches), making the anterior lip apparently the longest. THE GENITOURINARY APPARATUS. 289 The mucous membrane has numerous transverse folds, or rugae (rugce vaginales), passing to either side of a median ridge, both anterior and posterior, the columnae vagina' (columnm rugdrum anterior et posterior), formed by the coalescence of the two tubes in fetal life. Some of these rugae present wart- MONS VENERIS fe? > J {Mesenceph- alon, or mid- brain, {Corpora quadrigemina, Crura cerebri, Aqueduct of Sylvius. THE NERVOUS SYSTEM. 299 Brain or Encephalon. [continued i Prosen- cephalon, fore-brain Thalamanceph- alon, or dien- cephalon, or J Optic thalami, Subthalamic regions, Pituitary and pineal bodies. Structures in interpe- duncular space. Optic nerve and retina. Hinder part of third ventricle. Cerebral hemispheres, Olfactory lobes, Lateral ventricles, Telencephalon < Foramina of Monro, j Anterior portion of i third ventricle. The medulla oblongata (myelencephalon) is the upper expanded portion of the spinal cord, extending between the lower border of the pons and the upper border of the atlas. It is divided by two fissures — the anterior (fissura mediana ante- rior) and posterior median (fissura mediana posterior) fissures — into two halves, each one of which is subdivided into four columns, from before backward, the following: — (a) Anterior pyramids (pyramis medulla? oblongata') , or corpora pyramidalia, are two pyramidal masses of white nervous matter, placed between the anterior median fissure and the olivary body, and continuous with the anterior columns of the cord below ; (b) Lateral tract and olivary body, are continuous with the lateral columns of the cord below; (c) Eestiform bodies (corpus restiforme) are continuous below with the posterior columns of the cord. They are com- posed of the fibers of the columns of Goll and Burdach and the direct cerebellar tract. They diverge, the interval between them being the lower portion of the fourth ventricle. The structure of the medulla oblongata consists of both white and gray matter, the former arranged into four columns, the latter contained in the interior. The gray matter of the medulla is partly arranged into - and partly continuous with the gray matter of the cord. The posterior horns are called here "the tubercles of Rolando" (tuberculum Eolandi). On the floor of the fourth ventricle {foi>t> of a number of thin plates, folia, arranged in a series of crescent ic curves, with the concavity forward. The cerebellum consists of a central lobe (vermis), and two lateral hemispheres (Itemisphcrria cerebelli). The latter are separated on the inferior surface of the cerebellum by a deep hollow, the valley or vallecula (vallecula, cerebelli), which Lodges the medulla oblongata. The floor of the vallecula is formed by the inferior portion of the vermis. The incisura semilunaris (incisura cerebelli anterior) sep- arates the hemispheres in front, and rests against the corpora quadrigemina; The incisura marsupialis (incisura cerebelli jiostrrior) separates the hemispheres behind, and receives the upper por- tion of the falx cerebelli ; The great horizontal fissure (sulcus horizontalis cerebelli) separates the cerebellum into a superior and an inferior surface. The upper surface (fades cerebelli superior) shows the superior vermis (vermis superior cerebelli), with its correspond- ing Lobes in the hemispheres. These latter are separated from each other by fissures. The following table gives the arrange- ment of the lobules : — G beat Horizontal ElSSIRE. Pre- Frsenulum Lingula. Frsenulum central fissure Post- Ala. rJbbus cenl ral is. Ala. Anterior Anterior crescentic Lobus culminis. crescent ic I'n- lobe. lobe. Posterior Posterior crescent i(- Lobus cljvi. crescenl ic lobe. lobe. I'o-I Posterior Posterior Great superior lobe Folium cacuminis. superior lobe. horizontal fissure. 302 HUMAN ANATOMY. The inferior surface (fades cerebelli inferior) of the cere- bellum presents the deep depression, the vallecula, which ren- ders the connection between the vermis and the hemispheres less intimate. The divisions of the vermis and their correspond- ing lobules in the hemispheres are as follows : — Great Post- Pre- Post- Great Slender lobe, post-gracile, posterior in- ferior lobule. Lobulus biventer. Amygdala. Flocculus. Tuber valvulse Pyramis. Uvula. Nodule. Posterior inferior lobule. Lobulus biventer. Amygdala. Flocculus. horizontal fissure. pyramidal fissure. pyramidal fissure. nodular fissure. horizontal fissure. The cerebellum is connected with the encephalon by the peduncles of the cerebellum,, three in number, from above down- ward, as follows : — Crura ad cerebrum, superior cerebellar, peduncles, or processus e cerebello ad testes (bracliia conjunctiva cerebelli), pass to the cerebrum ; Crura ad posterior, middle cerebellar peduncles, or proces- sus ad pontem (bracliia pontis), the transverse fibers of the pons Varolii, connect the hemispheres; Crura ad medullam, inferior cerebellar peduncles or proces- sus e cerebello ad medullam, the restiform bodies of the medulla oblongata (corpora restiformia), connect with medulla ob- longata. . The internal structure of the cerebellum consists of an arbor vitge arrangement of gray matter, inclosing a white mass. In center of the latter is found a grayish, dentated mass, the corpus dentatum (nucleus dentatus), an irregular capsule of gray matter opening anteriorly. TITE CEREBRUM consists of a large, ovoidal mass, divided into two lateral halves, or hemispheres (hemisphmria cerebri), by the great longitudinal THE NERVOUS SYSTEM. 303 fissure (fissura longitudinalis cerebri), connected by a white Transverse commissure — the corpus callosum. The surface is irregularly marked by convolutions, or gyri (gyri cerebri), separated from each other by irregular depressions, fissures ( tis.. The great transverse fissure of Bichal (fissura cerebri trans- betweeen the cerebellum mid the cerebrum, admitting the pia mater to form tin- velum interpositum. .;. The fissure of Sylvius -fissura cerebri lateralis [Sylvii]), be- ginning at tin- anterior perforated apace and ascending obliquely. 304 HUMAN ANATOMY. 4. The fissure of Rolando {sulcus centralis [Rolandi]), descends from near the middle of the great longitudinal fissure to join the fissure of Sylvius. 5. The parieto-occipital fissure {sulcus occipitoparietalis) , on the postero-Iateral aspect of the cerebrum. 6. The callosomarginal fissure {sulcus singulus) . 7. The collateral fissure {fissura collateralis) . 8. The limiting sulcus of Reil (sulcus circularis [Reili]). The principal lobes of the brain are six, as follows : — ■ 1. Frontal lobe- (lobus frontalis) , on the outer surface of the brain; it is bounded below by the fissure of Sylvius, and behind by the fissure of Rolando. On the mesial surface it is bounded by the callosomarginal fissure, and on the inferior surface it is bounded behind by the stem of the Sylvian fissure. On the outer surface it is divided into the (a) Ascending frontal convolution {gyrus frontalis as- cendens) ; (b) Superior frontal convolution (gyrus frontalis superior); (c) Middle frontal convolution (gyrus frontalis medius) ; (d) Inferior frontal convolution (gyrus frontalis inferior) ; On the mesial surface it is divided into (a) Marginal gyrus (gyrus marginalis) ; (b) Paracentral lobule (lobulus paracentralis) . On the orbital surface it is divided into (a) Internal orbital convolution (gyrus orbit alis internus) ; (b) Anterior orbital convolution (gyrus orbitalis anterior) ; (c) Posterior orbital convolution (gyrus orbitalis posterior) . 2. Parietal lobe (lobus parietalis) , lies between the fissure of Rolando, the parieto-occipital, and the fissure of Sylvius, and consists of five gyri : — (a) Ascending parietal (gyrus centralis posterior) ; (6) Superior parietal (lobulus parietalis superior) ; f Supra-marginal (gyrus supramar- (c) Inferior parietal (lobulus I ginalis) , parietalis inferior ) , '■ Angular (gyrus angularis), *- Post parietal; 3. Occipital lobe (lobus occipitalis) lies at the posterior aspect of the cerebrum and is divided into first, second and third occipital convolutions. 4. Temporosphenoidal lobe or temporal lobe (lobus temporalis) , occupies the middle fossa of the skull. 5. Island of Reil, or central lobe, lies within the fissure of Sylvius. It consists of six convolutions — the gyri operti. 6. The limbic lobe surrounds the corpus callosum. Its extremities are united by the roots of the olfactory tract. The inner or median surface of the hemispheres presents five fissures, as follows : — 1. Callosomarginal (sulcus cingulus) . 2. Parieto-occipital (sulcus occipitoparietalis). 3. Calcarine (fissura. calcarina) . 4. Occipitotemporal or collateral (fissura collateralis) . 5. Dentate fissure, or sulcus hippocampi (fissura hippocampi). THE NERVOUS SYSTEM. 305 The lobes on the internal surface are six in number, as follows : — 1. Callosal convolution {gyrus formcatus or gyrus cinguli), de- scends as the gyrus hippocampi and terminates as the uncinate gyrus. •1. Marginal (gyrus marginaUs), or first frontal convolutions. 3. Quadrate (projcuneus) . 4. Cuneus, or occipital lobule (lobus occipitalis). 5. Uncinate gyrus. 6. Temporosphenoidal lobe or temporal lobe (lobus temporalis). Sul C.C. Su?.ima>J- Sulsuhf' Sulsubp FlS.p-Q; ..sSulcalmiy fis. c&k. Fig. 107. Convolutions and ficsures of the median and tentorial surfaces of the right cerebral hemisphere. Fis. 8., fissure of Sylvius; Sul. cat. marg., callosomarginal sulcus; Sul. subf., subfrontal sulcus; Sul. C. 0., sulcus of corpus callosum; Sul. marg., marginal sulcus; Sul. subp., subparietal sulcus; Fis. p. o., parieto-occipital fissure: Fie. calc, cal- carine Bssure; Fis. den., dentate Assure; Fis. col., collateral fissure. {Whitehead, after Van Qehuchten.) The inferior surface (fades basalis encephali) of each. hemisphere lb divided into three lobes — the anterior, middle and posterior. The two former occupy the anterior and middle fossa of the skull and the posterior rests upon the cerebellum, separated from it by the tentorium. This surface presents for study from before backward the following points: — The longitudinal fissure (fissura longitudinalis cerebri), separates the two hemispheres; Corpus calloan in. the great transverse commissure of the cerebrum, extending bj means of its peduncles to near the Sylvian fissure; 20 306 HUMAN ANATOMY. [Aim ina eincrca. is ;\ thin, gray layer, forming the anterior part of the inferior boundary of the third ventricle: Olfactory nerve, with it* bulb; Fissure of Sylvius [fissura cerebri lateralis [Sylvii]), between the anterior and middle lobes of the cerebrum, and lodges the middle cere- bral artery: Anterior perforated space (locus perforatus anticus or substantia perforata anterior), transmits vessels to the corpus striatum: Optic commissure [chiasma optieum). is formed by the junction of the optic tracts; Tuber cinaeutn. is a gray eminence between the corpora albicantia and optic tracts, and forms part of the tloor of the third ventricle: Infiindibuluni, is a tube of gray matter connecting the pituitary body with the third ventricle: Pituitary body {hypopht/sis cerebri), is a small, vascular, bilobed body, connected by the infundibulum and occupying the sella turcica (for histology vide '•Ductless Glands"); Corpora albicantia. or mammillaria. are two white, rounded masses, formed by the folding of the anterior crura of the fornix, and are sometimes called the bulbs of the fornix: Posterior perforated space [locus perforatus posticus: or sub- stantia perforata posterior) . allows the passage of blood-vessels to the optic thalami; Crura cerebri (pedunculi cerebri), or cerebral peduncles, connect the cerebrum with the medulla, cerebellum, and spinal cord: they con- sist of the anterior portion, or crusta. and the posterior portion, or tegmentum, between which is a mass of gray matter — the locus niger: Pons Varolii, covers up the posterior portion of the cerebral lobes. Interior of the Cerebrum. — The interior of the cere- brum, viewed above the level of the corpus eallosum. presents a white surface — the centrum ovale minus, the margins of which are convoluted gray matter, and are called labia cerebri. It is studded throughout with minute blood vessels — puncta vaseulosa. The hemispheres, viewed on a level with the corpus eallo- sum. present a large white mass — the centrum ovale majus — in the center of which is the connecting band, the corpus eallosum. Corpus Calhmim. — This connecting band forms the roof of the lateral ventricles. It is about four inches in length and varies from an inch and a halt' to two inches in width, present- ing in front a bend, or genu (genu corporis caUosi). below which it terminates in the tuber einereum through the lamina cinerea. Posteriorly it forms a thick, rounded fold — the splenium (spleniutn corporis caUosi). or pad — which is continuous with the fornix. The peduncles of the corpus eallosum ( gyrus subcallosus or peduncular corporis caUosi) are two reflected bundles of white matter given off near the anterior termination of the corpus, THE NERVOUS SYSTEM. 307 and each passing backward across the anterior perforated space of its own side to the fissure of Sylvius. The superior surface of the corpus callosum shows a de- Julfitf. Fig. 108. Inferior aspect of cerebral hemisphrro. Bui. <>lf., olfactory sulcus; Bui. tim/i. inf.. Inferior temporal sulcus; Fis. col., collateral fissure. {Whitehead, after \'u each Bide by elevated bands, the stria longitudinales, or nerve- of Lancisi. Externa] to these are the stria longitudinaJes laterales. On either side of the 308 HUMAN ANATOMY. raphe are many transverse lines — the linear transversa?, which indicate the direction of the fibers of the corpus. The cerebral commissuees are connecting bands of gray and white matter, pursuing either a transverse or anteroposte- rior course. Transverse: — Anterior (commissura anterior cerebri), middle, gray, or soft com- missure (massa intermedia or commissura mollis), and posterior com- missure ; Corpus callosum; Optic chiasm (chiasma opticum) ; Fornix ; Pons Varolii; Posterior medullary velum (velum medullare posterius). A n tero posterior : — Corpus callosum (nerves of Lancisi) ; Fornix ; Fasciculus uncinatus (uncinate fasciculus) ; Taenia semicircularis (stria? terminalis) ; Callosal convolution (gyrus fornicatus) ; Inferior longitudinal fasciculus (fasciculus longitudinalis inferior) ; Olfactory tracts (tractus olfactorius) ; Crura cerebri (pedunculi cerebri) ; Peduncles of pineal gland (habenula) ; Processus e cerebello ad testes. VENTRICLES OF THE BRAIN. The interior of the brain contains . five distinct cavities, named the ventricles of the brain, situated as follows: Two lat- eral ventricles (ventriculus lateralis), in the upper part, within the substance of the hemispheres, the third ventricle (ventriculus tertius) between the optic thalami at the base of the brain, the fourth ventricle (ventriculus quartus) between the medulla oblongata and the cerebellum, and the fifth ventricle within the septum lucidum between the two lateral ventricles. The ventricles intercommunicate — the two lateral ventricles with the third by means of the foramen of Monro (foramen interventriculars) , the third with the fourth ventricle by means of the iter a tertio ad quartum ventriculum (aquceductus cere- bri), and, in the fetus, with the fifth, and through the infundib- ulum with the cavity of the pituitary body. The lateral ventricle (ventrv ulus lateralis) is bounded as follows : — The roof, the corpus callosum; the floor is formed by the follow- ing parts from before backward: corpus striatum, taenia semicircu- THE NERVOUS SYSTEM. 309 laris. optic thalamus, choroid plexus, corpus fimbriatum and fornix; internally, by tlie septum lucidum; externally, in front and behind by the brain-substance. Each lateral ventricle presents three cornua — the anterior cornu, posterior cornu, or digital comity, and the middle count. —-Antlcorr Nud cauef ton _— Post. cor. Fig. 109. The lateral ventricles and choroid plexus. St. term., stria termin- als; That., thalamus; Hipp., hippocampus; Fimb., fimbria; .1;// cor anterior cornu of lateral ventricle; Nucl. caud., nucleus caudatus; Col. for., columns of the fornix; Yd. int., velum interpositum; Ch. pi., choroid plexus; Cat wo., calcar avis; Post, cor., posterior cornu of lateral ventricle [Whitehead, after '/ray.) The anterior cornu (cornu anterius) curves outward and forward over the corpus striatum and into the anterior lobe. The middle cornu (cornu inferius) passes into the middle lobe, descending to the transverse fissure at the base of the brain. 310 HUMAN ANATOMY. Its course is backward', outward, downward, forward and inward (B., 0., D., F., I.). The posterior cornu (cornu posterius) runs backward into the posterior lobe, its course being backward, outward and inward (B., 0., I.). Parts of Lateral Ventricle — Corpus Callosum. — Described above. Septum Lucidum (septum p el 'lucid him) . — Forms the inter- nal boundary of the lateral ventricle. It consists of two layers of white and gray matter, and is attached above to the under surface of the corpus callosum. below to the fornix, and ante- riorly to the prolongation of the corpus callosum. Between the laminae forming the septum is a narrow inter- val— the fifth ventricle. The Corpus Striatum. — Situated in the lateral ventricle, its broad end directed forward into the fore part of the body and anterior cornu of the ventricle, its narrow end directed outward and backward, and separated from its fellow by the thalami optici. The intraventricular portion is called the caudate nucleus (nucleus caudatus), the extraventricular, the lenticular nucleus (nucleus lentiformis) , the two separated by the internal capsule. The internal capsule (capsula interna) is a large layer of white fibers, derived from the medulla and crura cerebri, sep- arating the lenticular nucleus from the caudate nucleus ante- riorly, and the lenticular nucleus from the optic thalamus posteriorly. The external capsule (capsula externa) is a small layer of white fibers on the outer surface of the corpus striatum, between the lenticular nucleus and the claustrum. The claustrum is a convoluted layer of gray fibers between the external capsule and the island of Reil. The Tcenia Semicircular is, or Horny Band of Tarinus stria terminalis). — A band of medullary substance in the fur- row between the corpus striatum and the optic thalamus. Its anterior portion descends with the anterior pillar of the fornix, its posterior portion passes into the descending horn. Beneath it is the vena corporis striati. The choroid plexus (plexus chorioideus) , a vascular mem- brane, occupying the margin of a fold of pia mater, known as the velum interpositum (tela chorioidea superior; or, tela chorioidea ventriculi tertii) . It runs across the floor of the lateral ventricle, and communicates with its fellow of the op- THE NERVOUS SYSTEM. 3-Q polite side through the foramen of Monro. Posteriorly it descends into the middle horn of the lateral ventricle. The corpus iimbriatum (tcenia hippocampi or tamia fim- bria), a narrow white hand behind the choroid plexus. It is the lateral edge of the posterior pillar of the fornix. The fornix, a lamella of white tibrous matter, beneath the corpus eallosum, continuous with it posteriorly, but separated from it anteriorly by the septum lucidum. It consists of two symmetrical halves which join to form the body (corpus forni- cis), each half having an anterior and posterior crus where they do not join. The anterior crura (columnar fornicis) curve down to the base of the brain, where each crus spreads out and curves upon itself to form the corpus albicans of that side. From this point it passes to the corresponding optic thalamus. The posterior crura (crura fornicis), at their commence- ment, are joined to the under surface of the corpus eallosum. They pass downward into the descending horns of the lateral ventricles, being continuous with the concave borders of the hippocampi majores. The lateral edge of the posterior crus is called the corpus fimbriatum. The lyra is a series of lines, some transverse, others longi- tudinal and oblique, on the under surface of the fornix, between the diverging posterior crura. Optic Thalamus (thalamus). — The thalami optici are two large ganglionic masses, situated between the diverging portions of the corpora striata. Each thalamus rests upon the eorre- sponding crus cerebri. The thalamus is bounded externally by the corpus striatum and taenia semicircularis. and internally forms the lateral boundary of the third ventricle. Its upper surface is partly covered by the fornix. Its under surface forms the roof of the descending horn of the lateral ventricle. It- posterior and inferior part exhibits two rounded emi- nences, the external (corpus geniculatum hiferale) and internal (corpus geniculatum mediate) geniculate bodies. Its anterior extremity form- the posterior boundary of the foramen of Monro, which foramen connects the two lateral ventricles with the third. Velum interpositiini (tela chorioidea ventriculi tertii), a vascular membrane, reflected from the pia mater into the inte- rior of the brain through the transverse fissure. It passes 312 HUMAN ANATOMY. beneath the posterior border of the corpus callosum and fornix, and above the corpora quadrigemina, the pineal gland and the optic thalami. It forms the roof of the third ventricle (plexus chorioidea ventriculi tertii). Its anterior extremity passes on each side into the corresponding lateral ventricle, forming the anterior extremity of the choroid plexus. The vascular fringes of the velum interpositum projecting into the third ventricle are called the choroid plexuses of the third ventricle. It has two veins, the venae G-aleni (w. cerebri internee), which run along its under surface and are formed by the veins of the choroid plexuses and the vena? corporis striata (v. cor- poris striata). The venge Galeni unite to form a single trunk ■ — vena magna Galeni — (v. cerebri magna) and empty into the straight sinus. The posterior cornu (cornu postering) of the lateral ven- tricle runs into the substance of the posterior lobe. On the floor of this horn is an eminence corresponding to a sulcus be- tween two convolutions, and called the hippocampus minor (calcar avis). Between the posterior and middle (cornu inferius) horns is another eminence — the eminentia collateralis, or pes accessorius. The hippocampus major, or cornu ammonis (hippocam- pus), a white eminence running the entire length of the floor of the middle horn. This eminence is the doubled-in surface of the gyrus fornicatus. The lower extremity of the hippocampus major is called the pes hippocampus (digitationes hippocampi). The fascia dentata (fascia dentata hippocampi) , the gray and serrated edge of the middle lobe. It is really external to the cavity of the middle cornu. The third ventricle (ventriciilus tertius) is a mere fissure in the median line of the cerebrum, situated between the optic thalami. It communicates with the lateral ventricle by the foramen of Monro and with the fourth ventricle by the iter a tertio ad quartum ventriculum. The cavity is crossed by three commissures — the anterior commissure, a white, rounded cord ; the middle or soft commissure, composed of gray matter ; and the posterior commissure, a white band connecting the two optic thalami. It is bounded by the following structures : — The roof, by the velum interpositum, suspending the choroid plex- uses of the third ventricle, and laterally the peduncles of the pineal gland; floor, by the parts inclosing the interpeduncular space at the base of the brain, viz. : the lamina cinerea, tuber einereum and infundib- ulum, corpora albican tia and the posterior perforated space; laterally, THE NERVOUS SYSTEM. 313 by the optic tlialami; m front, by the anterior commissure and anterior crura of the fornix; behind, the posterior commissure and the iter a tertio ad quartum ventricuhun. The fourth vextricle (ventriculus quartus) is a dia- mond-shaped cavity between the cerebellum behind and the posterior surface of the medulla oblongata and pons in front. It is inclosed behind by the pia mater, which contains an open- ing for the exit and entrance of the subarachnoid fluid from the subarachnoidean space of the brain and spinal cord, and a vascular fold of pia mater — the choroid plexus. Its lower Ch.ro i J /Pineil Stria ?^L /Kntal R«tess of W Vent. P'swA^, ■ I^IIS^ \ v? \W SM/ \"Su-P Clorp <^M.aJ. &-'t«ofR«f:P„f Space- ^••WjNV J "**"""— AF-Wexty- FIG. 110. Mesial section of brain and brain stem. 1, anterior commissure; 2, middle commissure; 3, posterior commissure; 4, pituitary body; IV, fourth ventricle. angle is continuous with tbe central canal of the spinal cord; and from the resemblance it bears to a writing-pen, the name calamus scHptorvus is applied. It communicates in front with the tbird ventricle by tbe iter a tertio ad quartum ventriculum. It is bounded as follows: — Tin- roof, valve of Vieussens (rchun mrihilhirr antrrius) and the cerebellum, containing in front the foramen of Magendie [apertura medialia ventricuM quorti), by which it communicates with the sub- arachnoidean -pace: the floor, of rhomboidal outline, is traversed by a vertical median fissure [sulcus longitudinalis fossce rhomboidece) con- tinuous with the central canal of the cord. At the broadest part of the ventricle are a series of transverse white lines or stria' medullares, de 314 HUMAN ANATOMY. rived from the cochlear root and nucleus. These striae divide the floor into two triangles, a superior and an inferior. The inferior triangle presents a groove, the fovea inferior, whose diverging limbs below form the so-called ala clnerea, which is a darker colored, triangular space. There are in this locality several eminences, corresponding with the nuclei of origin of the pneumogastric, glossopharyngeal and other cranial nerves. The superior triangle presents an elevation, produced by underlying white fibers, the fasiculus teres (colUculus facialis). Above and external to the fasciculus teres is a depression, the fovea superior; and just above this is a bluish area, the locus ccc-ruleus. This mottled appearance is caused by the presence of the substantia ferruginea, a peculiar pigment of some of the nerve cells and in them, one of the roots of the trifacial nerve terminates. Anteriorly, the pons Varolii and medulla oblongata; posteriorly, the cerebellum; laterally, the processus e cerebello ad testes, the restiform bodies, and posterior pyramids of the medulla. Its lining membrane is continuous with that of the third ventricle. The fifth ventricle (cavtim septi pellucidi) is a narrow fissure, formed within the two lamince of the septum lucidum, being originally a part of the great longitudinal fissure. It is bounded : — Above, by the under surface of the corpus callosum; Below, by the anterior part of the fornix; Laterally, by the lateral ventricles, from which it is separated by the septum lucidum. It is not lined with epithelium. The mesencephalon includes those portions of the brain substance which connect the cerebrum, cerebellum and medulla oblongata together, and comprises the following structures : — Crura cerebri, before described (ante, p. 306). The valve of Vieussens, or anterior medullary velum, is a thin layer of white matter stretched between the processes e cerebello ad testes, and forming the roof of the iter a tertio ad quartum ventriculum. It presents an elevated ridge descend- ing on its upper part from ' the corpora quadrigemina — the frenulum. Corpora or tubercula quadrigemina, or optic lobes, are four spherical eminences, placed in pairs above the valve of Vieus- sens, and behind the third ventricle, beneath the posterior bor- der of the corpus callosum. The two anterior are called the nates (colliculi superiores) , the two posterior the testes (colliculi inferiores). The brachia anterior — (brachium quadrigeminum superius) and posterior — brachium quadrigeminum inferius) are two white cords con- THE NERVOUS SYSTEM. 315 iiecting them with the optic thalamus and beginning of the optic Bracts. The processus e cerebello ad testes connect them with the cerebellum. To the outer side of the optic lobes are two small masses called the corpus geniculatum externum (corpus genie id atum Jul crate) and internum (corpus geniculatum mediate). The pineal gland, or epiphysis cerebri (corpus pineale), is a >mall reddish, conical body, resting upon and between the nates. It represents the rudiment of a median eye of certain extinct lizards and amphibia (Spencer). It is attached to the cerebrum by its two peduncles (habenula), and is held in posi- tion by a fold of the pia mater. It contains a cavity, tilled with a viscid fluid and secretory matter composed of phosphate and carbonate of lime, phosphate of magnesia, ammonia, and a little animal matter — -the acervulus cerebri. THE INTERNAL STRUCTURE OF THE MEDULLA OBLONGATA. The anterior and lateral pyramidal tracts of the cord are continued into the medulla oblongata as the pyramids. The lateral pyramidal tracts decussate in the lower part of the medulla. The anterior ground bundle is continued upward into the medulla as the posterior longitudinal bundle. It lies behind the pyramids in the medulla. The sensory columns of the cord (Goll and Burdach) are continued into the medulla oblongata and terminate in the funiculus gracilis and euneatis. They increase in size and each develops a nucleus: the clava and cuneate nucleus, respectively. These fibers partly form the restiform bodies. The decussation of the sensory fibers takes place at a higher plane than that of the motor fibers. It is also called the decus- sation of the fillet and it consists of the decussating fibers de- rived from the clava and the cuneate nucleus. These fibers are called deep arcuate fibers. After decussating the fibers are con- tinued upward behind the pyramids, displacing backward the posterior longitudinal bundle {fasciculus longitudinalis me- dialis). In the medulla on cross-section are seen the olivary bodies (nucleus olivaris inferior), which contain the dentate nuclei. The restiform bodies (corpus restiforme), ot inferior cere- bellar peduncles, are formed by the direct cerebellar tract, the superficial or externa] arcuate fibers (.fibres arcuatce external), the 316 HUMAN ANATOMY. internal arcuate fibers (fibrce arcuatw internee), and the cere- bello-olivary fibers. The formatio reticularis is seen behind the pyramids and the olivary bodies in the medulla oblongata. It is composed of the deep arcuate fibers, fibers of Gowers's tract and the antero- lateral ground-bundle. The gray matter of the cord is continued into the. medulla. The anterior horns are cut off and displaced by the decussation of the lateral pyramidal tract and the posterior horns are dis- placed outward by the increase in size of the posterior sensory tracts. The latter are known as the nucleus of Rolando and are capped by the substantia gelatinosa Rolandi. The central canal is expanded into the fourth ventricle. The gray matter forms nuclei for the cranial nerves in the floor of the fourth ventricle. THE INTERNAL STRUCTURE OF THE PONS VAROLII. On cross-section the pons is seen to consist of a dorsal tegmental part, and a ventral part, or crusta (pars basilaris pontis). The crusta is composed of transverse fibers (fibers from the cerebellum to the pons and from the nucleus pontis to the cerebellum), superficial (fibrce pontis superficiales) and middle, which go to form the middle cerebellar peduncles; longitudinal fibers, which belong to the pyramidal tracts, much scattered; and gray matter, which here forms a nucleus called the nucleus pontis. The tegmentum of the pons contains a thick layer of gray matter, which forms the floor of the fourth ventricle, and from which cranial nerves take their origin; formatio reticularis, which is the continuation upward of the same from' the medulla ; the superior olivary nucleus (nucleus olivaris superior) ; the posterior longitudinal bundle (fasciculus longitudinalis medi- alis) ; the fillet (lemniscus) ; the superior cerebellar peduncle (brachium conjunctivum) ; and the corpus trapezoides. The fillet occupies a position between the crusta and teg- mentum, and to differentiate it from a tract that makes its appearance above the nucleus of the third nerve is called the mesial fillet (lemniscus medialis). The other tract is named the lateral fillet. The lateral fillet (lemniscus lateralis) is composed of longitudinal fibers which take origin from the- nucleus of the eighth cranial nerve (cochlearis) of the same side, from that THE NERVOUS SYSTEM. ;;i; of the opposite side, and from the superior olive. They end in the inferior quadrigeminal body, the internal geniculate body and a few in the superior quadrigeminal body. Mtitcoti nuclcaurf. J.lint.cap. Ext. cap. ■Is/. ■MicL lent. '■■Claus. ■—Pl.intcap. .'Thai. ... C.j.m. ..T.nud.caud. ■Fimb ■-M pp. ..Post, con ule; Ext. cap., external capsule; Tat, Island of Retl; Vucl Umt Ducleus lentiformis; Clou., elaustrum; P. /. int. cap., posterior Hrnii o* internal capsule; Hipp hippocampus; Post, cor., „ st ,•],,•', -, . lateral ventricle-. (Whitehead, after Landois.) 'UOL,-"UI Lor,ul 01 The mesial fillet has been described. II lakes its origiu in the medulla i'mm the euneate and gracile nuclei of the opposite 318 HUMAN ANATOMY. side. Some of its fibers end in the superior quadrigeminal body, but the remainder pass through the subthalamic tegmental, region into the posterior part of the lateral nucleus of the optic thalamus. Some end here, while some are continued through the thalamus, enter the corona radiata, and pass to the posterior central gyrus of the Eolandic region. THE STRUCTURE OF THE CEREBELLUM. On section the gray matter of the cerebellum is found to occupy the cortex (substantia corticalis) ; to its arborescent appearance the term arbor vitce has been applied. The white matter, medullary body, of each hemisphere con- tains a nucleus of gray matter: the corpus dentatum (nucleus dentatus). The middle peduncles connect the cerebellum with the pons. The inferior peduncles, or restiform bodies, or crura ad medullam (corpora restiformia) , connect the medulla and cord with the cerebellum. The superior peduncles (brachia conjunctiva cerebelli) con- nect the cerebellum with the cerebral cortex. After decussating below the corpora quadrigemina some of the fibers pass to the opposite red nucleus (nucleus tegmenti) in the tegmentum of the crura cerebri passing through the optic thalamus to end in the Eolandic region, and others end in the optic thalamus. Each superior peduncle contains both afferent and efferent fibers. The cerebellar cortex consists of two layers: an outer, molecular layer, and an inner, granular layer. Between these two layers is a single layer of large cells, the corpuscles of Purkinje. THE STRUCTURE OF THE MID-BRAIN. The mid-brain consists of a dorsal part, the corpora quad- rigemina, and a ventral part, the crura cerebri. It is tunneled by the aqueduct of Sylvius, which connects the fourth ventricle with the third ventricle. The upper end of the crura cerebri is encircled by the optic tract. On section the crura cerebri show a ventral and lateral portion, — the tegmentum, — and a dorsal portion, the lamina quadrigemina. The ventral portion contains the substantia nigra. The fissure of Sylvius is surrounded by gray matter, from which the third and fourth cranial nerves take origin. THE NERVOUS SYSTEM. 319 SENSORY MOTOR AREA INTERNAL CAPSULE FIRST TEMPORAL CYBU8 QUADR10BM1NATB BODIES^- ' INFERIOR PEDUNCLE- OF CEREBELLUM, SENSORY DECUSSATION (DECUSSATION OF FILLET JWJCLEUS CUNEATIS OOWERSB TRACT EIGHTH NERVE NINTH NERVE TENTH NERVE NUCLEUS GRACILIS POSTEROMEDIAN COLUMN, POSTEROLATERAL COLUMN ■DIRECT CEREBEL1.AR TRACT POSTERIOR NERVE ROOT Wia. 112. Diagram of sensory tracts from spinal cord to brain (original). 320 HUMAN ANATOMY. The inferior or posterior quaclrigeminal bodies contain the fibers of the lateral fillet posteriorly and the brachium laterally. The superior or anterior quadrigeminal bodies consist of four strata: stratum zonale, stratum cinereum, stratum opti- cum and stratum lemnisci. The mesial and upper fillet in part end in the latter, which also contains large nerve cells. Some fibers from the occipital lobe and fibers from the retina, conveyed by the superior brachium, end in the superior quadrigeminal bodies. The superior cerebellar peduncles have been described ; they connect the cerebellum with the cerebral cortex. The red nucleus (nucleus tegmenti) is found in the teg- mentum. Some of the fibers of the superior cerebellar peduncles end there. The posterior longitudinal bundle lies in the tegmentum and is the continuation upward of the tract of the same name in the medulla and pons. It is connected with the nuclei of the motor nerves of the muscles of the eyeball. The lateral fillet is continued upward from the pons in the tegmentum. Its connections with the superior olivary nuclei and the nuclei of the eighth nerve have been described. The mesial fillet likewise is continued upward from the pons in the tegmentum of the crura. The course of the fibers has been described. The crusta of the crura contains the pyramidal fibers and the corticopontine fibers. THE STRUCTURE OF THE CEREBRUM. On cross-section the cut brain surface shows an outer gray cortex and the inner white matter and cavities (centrum ovale) , one to each hemisphere : the lateral ventricles. The white mat- ter contains certain masses of gray matter : the optic thalami and the corpora striata, which latter are on each side of the brain divided into the caudate and lenticular nuclei by the internal capsule. Between the optic thalamus and caudate nucleus on the inside and the lenticular nucleus on the outside passes the broad band of white fibers known as the internal cap- sule (capsula interna). As it nears the cortex the fibers spread out. To this is given the name corona radiata. Between the external capsule (capsula externa) and the cortex of the island of Eeil is seen a thin sheet of gray matter : the claustrum. The white matter between the claustrum and the lenticular nucleus is known as the external capsule. On horizontal section THE NERVOUS SYSTEM. 321 CORTEX OF THE. «8NS0RY MOTOR AREA CORONA RADIATA INTERNAL CAPSULE -V j fOURTH NERVE P0N8 •PINAL CORD DECUSSATION OF PYRAMIDS ANTERIOR PYRAMIDAL TRACT ANTERIOR COMMISSURE CAUDATE NUCLEUS OPTIC THALAMUS LENTICULAR NUCLEUS THIRD NERVR FIFTH NERVE — SIXTH NERVH SEVENTH NERVE NINTH NERVB TENTH NERVB ELEVENTH NERVB TWELFTH NERVB LATERAL PYRAMIDAL TRACT ANTERIOR NKRVK ROOT Fig. 113. Diagram of motor tracts from brain to spinal cord (original). 21 322 HUMAN ANATOMY. of the brain the internal capsule is seen to be bent upon itself, the interval between the optic thalamus and the caudate nucleus. This bend is called the genu (genu capsulce internee). One- third of the capsule lies in front of this bend, and is called the anterior limb (pars frontalis capsulce internee) ; the portion behind the genu is called the posterior limb (pars occipitalis capsular internee). The anterior limb contains (1) fibers that pass from the optic thalamus to the lenticular and caudate nucleus, (2) fibers that pass from the optic thalamus to the cortex of the frontal lobe, and (3) fibers that pass from the cortex of the frontal lobe to the nucleus pontis. The posterior limb contains (1) a continuation upward of a portion of the mesial fillet and the superior cerebellar pedun- cles, (2) the pyramidal tracts or motor fibers from the Eolandic area, (3) the fibers of the optic radiation (radiatio occipito- thalamica), (4) the fibers of the auditory radiation (radiatio temporothalamica) , and (5) the temporopontine tract. The opposite sides of the brain are connected by the com- missural fibers. Convolutions on the same side are connected by association fibers. Projection fibers are those which unite the cerebral cortex with nuclei in lower levels. They pass prin- cipally through the corona radiata. The cerebral cortex consists of five layers : the stratum zonale, the layer of small pyramidal cells, the layer of large pyramidal cells, the layer of polymorphous cells and a layer of fusiform cell-bodies. CRANIAL NERVES. The cranial nerves (nervi cerebral es) consist of twelve pairs, as follows : — 1. Olfactory, 8. Auditory (portio mollis), 2. Optic, 9. Glossopharyngeal, 3. Motores oculorum, 10. Pneumogastric (vagus, or par 4. Pathetici, vagum ) , 5. Trifacial, 11. Spinal accessory, 6. Abducentes, 12. Hypoglossal. 7. Facial (portio dura), 1. Olfactory nerve (n. olfactorius) , special nerve of smell. A number of nerves (20) arise from the olfactory bulb or lobe of the brain. Superficial origin of the tract by roots : internal from frontal lobe, middle or gray root from the olfac- tory tubercle (trigonum olfactorium) between the other roots THE NERVOUS SYSTEM. 323 of the tract, and external from the middle lobe; deep origin, from gyrus fornicatus, uncinate gyrus of limbic lobe; course, roots unite, pass forward, and form bulbus olfactorius (from this the olfactory nerves are given off) ; exit, foramina of crib- riform plate of ethmoid; distribution, by three groups to mu- FiG. 114. Scheme of the nuclei and root-fibers of the cranial nerves. (Whitehead, after Edinger.) ( Schneider ian) membrane of nares, inner to septum nasi, middle to roof of nasal fossa, outer to superior turbinated bone. 2. Optic nerve (n. opticus), special nerve of sight; super- ficial origin, optic chiasms or commissure formed by union of the optic tracts. The commissure and tracts contain intercere- 324 HUMAN ANATOMY. bral fibers, the cerebroretinal fibers of the same side, and the cerebroretinal fibers of opposite sides. Deep origin, the optic tracts arise from optic thalamus, the upper corpora quadri- gemina and corpora geniculati; course, diverge and pass for- FiG. 115. Base of brain and cranial nerves; 1, olfactory bulb; 2, optic nerves; 4, tractusi opticus; 5, crus cerebri; 6, third pair of nerves; 7, fourth pair of nerves; 8, fifth pair of nerves; 9, sixth pair of nerves; 10, pyramid; 11, olivary body; 22, pons Varolii; 24, seventh and eighth pairs of nerves; 25, ninth, tenth and eleventh pairs of nerves; 26, twelfth pair of nerves; 27, cerebellum. ward; exit, optic foramen; distribution, to ganglion cells in the retina. 3. Motor oculi (n. oculomotorius) , motor nerve; super- ficial origin, inner surface of crus cerebri ; deep origin, from the Forehtad ttud Stalp 'ouppa-Mytlidaad^arrh^ad la anJ Itn-krymiil >w [/al'tr'ilil PIG. 116. Optic, oculomotor, trochlear, and trifacial nerves, THE NERVOUS SYSTEM. 325 oculomotor nucleus in floor of aqueduct of Sylvius; course, from inner side of eras cerebri passes forward, descends along exter- nal wall of cavernous sinus; exit, sphenoidal fissure between two heads of external rectus muscle; distribution, by superior and inferior divisions, to all the ocular muscles (including the iris) except the external rectus and superior oblique. 4. Trochlear (to, trochlearis — pathetic), motor nerve; superficial origin, from valve of Yieussens, on outer side of crus cerebri; deep origin, from floor of aqueduct of Sylvius; course, outer side of crus cerebri, through outer wall of cavernous sinus ; exit, sphenoidal fissure; distribution, to superior oblique muscle. 5. Trifacial, or Trigeminus (to. trigeminus), common sensation, taste and motion ; superficial origin, by two roots, like a spinal nerve, from the side of the pons Varolii; deep origin, the motor root, from (1) a nucleus in the floor of the aqueduct of Sylvius (descending motor root), and (2) from a nucleus in the pons (these join to form the motor root) ; the sensory root ends in two terminal nuclei — (1) the sensory nucleus of the fifth nerve in the pons, and (2) the substantia gelatinosa Rolandi in the pons, medulla and the spinal cord as far down as the second cervical nerve; course, passes forward to apex of petrous portion of temporal bone, where the sensory root enters Gasserian ganglion, the motor passing beneath, and later joins a branch of the ganglion. It divides into three branches — ophthalmic, superior maxillary and inferior maxil- lary; exit, ophthalmic by sphenoidal fissure, superior maxillary by foramen rotundum, inferior maxillary by foramen ovale. Ophthalmic nerve (n. ophthalmicus), entirely sensory, sup- plies lachrymal gland, upper eyelid, skin and muscles of fore- head, eyebrow, aose, eyeball (ciliary muscle, iris, etc.), mucous membrane of eyelids and nose and the ciliary ganglion. Its branches are: — Lachrymal (n. lacrimalis) ; Frontal ( n. frontalis) ; Naaal (n. nasociliaris) . Superior maxillary nerve (to. maxillaris), entirely sensory also; supplies sensation to upper jaw, teeth, hard and soft palates, tonsils, gums, antrum of Eighmore, muscles, skin and mucous membrane of lower eyelid, muscles and skin of cheeks and upper lip, and mucous membrane of floor of nares. 326 HUMAN ANATOMY. Middle superior den- tal ( ramus alveo- la/ris superior me- dius ) , Anterior superior den- tal (ramus alveo- laris superior ante- riores ) , Palpebral ( rami pal- pebrales inferiores) , Nasal ( rami nasales interni) , Labial (rami labiates superiores ) . Its branches are :- Meningeal ( n. menin- geus medius) , Orbital ( n. zygoma- ticus ) , Spheno-palatine ( n. sphenopalatine ) , Posterior superior den- tal (rami alveolcures superiores posteri- or es ) , Inferior maxillary nerve (n. mandibularis) consists of two portions: anterior or motor, and posterior or sensory, having a threefold function, because one of its branches, the lingual, is joined by the chorda tympani nerve (given off from the facial nerve), which conveys gustatory fibers to the anterior two-thirds of the tongue. It supplies motion to all the muscles of masti- cation (except to buccinator), anterior belly of digastric and mylohyoideus ; sensation to skin of ear, lower part of face, lower lip and tongue/ Previous to its division, the primary trunk gives off the recurrent (n. spinosus) and the internal pterygoid (n. pterygoideus internus). Immediately below the base of the skull it divides into an anterior and a posterior portion. Its branches- are : — Posterior Portion. Auriculotemporal (n. auriculotem- poral ) , Inferior dental (n. alveolaris in- ferior ) , Lingual ( n. lingualis ) . Anterior Portion. Masseteric (n. mussetericus) , Deep temporal (nn. temporales profundi ) ( 2 ) , Buccal (n. buccinatorius) , Pterygoid (n. pterygoideus exter- nus ) , The fifth nerve has four ganglia connected with it: — (a) Ophthalmic, or lenticular (ganglion ciliare) ■ (b) Sphenopalatine, or Meckel's (ganglion sphenopalatinum) ; (c) Otic, or Arnold's (ganglion oticum) ; (d) Submaxillary (vide Sympathetic System) (ganglion sub- maxillar e) . 6. Abducens (n. abducens), motor; superficial origin, pyramidal body and pons Varolii; deep origin, floor of fourth ventricle; course, passes forward in cavernous sinus; exit, sphe- noidal fissure; distribution, to external rectus muscle. 7. Facial (n. facialis), motor nerve; superficial origin, lateral tract of medulla and pons Varolii; deep origin, from a nucleus in the pons, deeply placed, from which the fibers ascend THE NERVOUS SYSTEM. 337 k §.■§ &? ?s ill Fig. 117. 328 HUMAN ANATOMY. close to the floor of the fourth ventricle, where they form the eminentia teres, arch over the nucleus of the sixth nerve, and then emerge (this nerve carries some sensory fibers, probably gustatory fibers from the tongue, through the chorda tympani nerve to the pars intermedia [n. intermedins], near the nucleus of the ninth nerve) ; course, forward and outward, through in- ternal auditory meatus, aqueductus Fallopii and inner wall of tympanum; exit, stylomastoid foramen; distribution, to muscles of expression, buccinator, and platysma, posterior belly of digas- tric, stylohyoid attrahens and retrahens aurem muscles, to laxator tympani and stapedius through tympanic branch, dorsal surface of tongue through chorda tympani, and levator palati, and azygos uvulae muscles through Vidian. In its course through the temporal bone it communicates with many impor- tant nerves; in the internal auditory meatus, with auditory (portio mollis) ; in aqueductus Fallopii, with otic ganglion, by the small petrosal; Meckel's ganglion, by the large petrosal nerve ; the sympathetic of the great meningeal, by the external petrosal nerve; and at its exit from the stylomastoid foramen, with the glossopharyngeal, pneumogastric, auriculotemporal, auricularis magnus and the carotid plexus. Its branches are : — On the Face. {Temporal (rami temporales) . Malar (rami zygomatics). Infraorbital (rami buccales). In Aqueductus Fallopii. Tympanic ( n. stapedius ) . Chorda tympani. ^ r Buccal (rami buccales). r, . . . , j Supramaxillary (ramus mar- Cervicoiacial, < £• 7- •'-,-, v 7 > ' ', gnnahs mandioukc) . I Inframaxillary (ramus colli). Posterior auricular, Stylohyoid (ramus stylohyoideus) , Digastric (ramus digastricus) . 8. Auditory (n. acusticus), special nerve of hearing, con- sists of two portions — (1) cochlear, or auditory portion, and (2) vestibular, or fibers whose function is to localize position. The former pass from the cochlea to the (1) accessory auditory nucleus in the medulla, and (2) to the lateral acoustic tubercle in the medulla. These fibers, by means of the lateral fillet, communicate with the inferior corpora quadrigemina. The vestibular fibers (fibrce vestibularis) pass from the vestibule to the external and internal dorsal nuclei in the floor of the fourth ventricle, and by the sensory decussation to the nucleus cuneatus. THE NERVOUS SYSTEM. 329 Fiu. US. 330 HUMAN ANATOMY. Superficial origin, from groove between olivary and resti- form bodies; course, winds around restiform body, and passes forward to internal auditory meatus, with the facial; distribu- tion, to internal ear by two branches : — Vestibular, and Cochlear {vide Ear, p. 370). 9. Glossopharyngeal (n. glossopharyngeus) , or ninth, nerve of motion, common sensation, and taste ; superficial origin, from medulla oblongata, between olivary and restiform bodies ; deep origin^, from motor and sensory gray nuclei in floor of fourth ventricle; course, passes outward across flocculus; exit, jugular foramen, in separate sheath, in front of pneumogastric and spinal accessory, having two gangliform enlargements upon it — jugular, and the petrous or ganglion of Andersch; dis- tribution, to mucous membrane of fauces, tonsil, pharynx and middle ear, muscles of the pharynx, and special sense of taste to the base and sides of the tongue. Its branches are: — Tympanic ( Jacobson's ) , (n. tym- Pharyngeal branches (rami phar- pamcus ) , yngei ) , Carotid branches (n. caroticotym- Tonsillar branches (rami tonsil- panicus superior et n. carotico- lares), tympanicus inferior ) , Lingual branches ( rami linguales ) . Muscular branches (ramus stylo- pharyngeus ) , 10. Pneumogastric, vagus, or par vagum (n. vagus), tenth nerve, motor and sensory; superficial origin, from groove between olivary and restiform bodies by a dozen filaments ; deep origin, from nuclei in floor of fourth ventricle; course, passes outward across the flocculus; exit, jugular foramen in a com- mon sheath with the spinal accessory nerve; distribution (not definitely known), motor nerve to the organs of respiration and voice, and a motor and sensory nerve to heart, esophagus, pharynx and stomach. It branches are : — Meningeal (ramus meningeus) , Cervical cardiac (rami carcliaci Auricular (Arnold's) (ramus superior es) , auricularis), Thoracic cardiac (rami cardiaci Pharyngeal (ramus pharyngcus) , inferior -es) , Superior laryngeal, Anterior pulmonary, Recurrent laryngeal (n. laryngeus Posterior pulmonary, inferior), Esophageal (rami esophagei) , Gastric (rami gastrici) . It also communicates with the left hepatic sympathetic plexus. THE NERVOUS SYSTEM. 331 11. Spiral accessory (n. accessorvus) , eleventh, motor nerve; superficial origin, from lateral tract of medulla and spinal cord as low as sixth cervical nerve; deep origin, spinal portion from anterior horn of gray matter; accessory portion from gray nucleus in floor of fourth ventricle; course, the spinal portion enters the skull through the foramen magnum, and joins the accessory portion in the jugular foramen ; exit, jugular fora- men in sheath with the pneumogastric ; distribution, to sterno- cleidomastoid and trapezius muscles, communicating with the cervical plexus and pharyngeal and laryngeal branches of pneu- mogastric. 12. Hypoglossal (n. lnjpoglossus) , or twelfth nerve, motor nerve of tongue; superfii ial origin, from groove between olivary and pyramidal bodies by about a dozen filaments; deep origin, from gray nucleus at lowest part of floor of fourth ventricle; exit, anterior condyloid foramen; distribution, to the omohyoid (both bellies), sternohyoid, sternothyroid, thyrohyoid, and mus- cles of the tongue — styloglossus, hyoglossus, geniohyoid, genio- hvoglossus, communicating with the pneumogastric, sympa- thetic, first and second cervical and gustatory nerves. Its branches are: — Descendens hypoglossi (ramus descendens) , Muscular, Thyrohyoid '{ramus thyreohyoid-eus) , Meningeal. Membranes of the Cord.< — The membranes of the spinal cord are three — the dura mater, arachnoid and pia mater. Dura mater (dura mater spinalis) is> a loose fibrous sheath, continuous with the dura of the brain, and from which it differs in not inclosing the venous sinuses, not dipping into the fissures of the cord, and not being adherent to the bony canal. From the latter it is separated by the venous plexuses (plexus venosi vertebrates interni) and some connective tissue. It extends the whole length of the canal, from the foramen magnum (to which it is attached) to the top of the sacrum. Arachnoid (araclmoidea spinalis) is a delicate serous sac, continuous above with the cerebral arachnoid, inclosing the pia mater, from which it is separated by an interval — the sub- arachnoidean space (cavum subarachnoideale) . It is filled with the cerebrospinal fluid (liquor cerebrosjnnalis). The outer sur- face of the arachnoid is in contact with the dura, the space between them being called the subdural space (cavum subdurale). Pia mater (pia mater sjnnalis) is a fibrous membrane, closely adhering to the cord and forming its neurilemma. Over the anterior median fissure, if is strengthened by a fibrous hand 332 HUMAN ANATOMY. — the linea splendens — .and laterally has the lig amentum den- ticulatum. It terminates below the cord as the plum terminate — a slender filament. THE SPINAL CORD. The spinal cord {medulla spinalis) is the elongated portion of the cerebrospinal axis contained in the spinal canal. Its length is about sixteen to eighteen inches, extending from the medulla above to the lower border of the first lumbar vertebra below, where it terminates in the cauda equina by a slender pro- longation of gray substance, called the conus medullaris. It presents two enlargements, the upper or cervical (intu- mescentia cervicalis), extending from the third cervical to the second dorsal vertebra, and the lower or lumbar (intumescentia lumbalis), about the position of the second or third dorsal vertebra. It is divided into two lateral halves by the anterior (fissura mediana anterior) and posterior median fissures (sulcus medianus posterior), united in center by the commissure. The lateral portions are subdivided by anterolateral (sulcus lateralis anterior) and posterolateral fissures (sulcus lateralis posterior) into anterior lateral and posterior lateral columns, and poste- riorly a narrow fissure separates the posterior median column from the posterior median fissure. The gray substance occupies the center of the cord, and is arranged into two crescentic masses connected together by the gray commissure (commissura grisea). The posterior horn (columna grisea posterior) forms the apex cornu (apex columna grisea posterior), from which arises the posterior root of the spinal nerves. The anterior horn (columna grisea anterior) is thick and short, and affords origin to the anterior root of the nerve. The gray commissure contains throughout its whole length a minute canal — the central canal, or ventricle of the cord (canalis centralis), continuous above with the fourth ventricle. SPINAL NER.VES. The spinal nerves (nervi spinales) consist of thirty-one pairs, arranged in the following order: cervical, eight pairs; dorsal, twelve pairs; lumbar, five pairs; sacral, five pairs; coccygeal, one pair. Each of the spinal nerves arises by two roots, an anterior (radix anterior) or motor and a posterior (radix posterior) or sensory. The fibers of the anterior root arise from the antero- lateral columns, originating deeply in the gray mattei of the THE NERVOUS SYSTEM. 333 Fig. 119. 334 HUMAN ANATOMY. cord. The posterior roots arise at the posterolateral fissure, also originating deeply in the gray matter of the cord. The pos- terior roots have each a ganglion (ganglia spinales) developed upon it — except sometimes the first cervical. These roots unite and the nerve then subdivides into two branches, both having motor and sensory fibers. The posterior branches are small; they supply the skin and muscles of the back. The anterior branches supply the neck, front and sides of the trunk, and the extremities. Cervical Plexus (plexus cervicalis). — The cervical plexus is formed by the anterior divisions of the first to the fourth cervical nerves. It is covered by the sternomastoid muscle, and Fig. 120. Section of spinal cord and membranes: 1, dura mater; 2, arach- noid membrane; 3, ganglion on posterior root; 4, anterior root of spinal nerve; 5, 5, subarachnoid space; 6, posterior branch of spinal nerve; 7, anterior branch of spinal nerve. rests upon the scalenus medius and levator anguli scapulae mus- cle. It gives off: — Superficial. Superficialis colli (n. cutaneus Occipitalis minor (n. occipitalis colli), minor), Auricularis magnus (n. auricu- Supraclavicular (nn. supi'aclavic- laris magnus), ulares) . Deep. Deep (Internal Series) Deep (External Series) Communicating, Communicating, Muscular, Muscular. Phrenic ( n. phrenicus ) , Communicantes hypoglossi, The phrenic nerve,, or internal respiratory of Bell, is de- rived from the third and fourth cervical nerves, with a branch from the fifth. It descends into the chest between the sub- clavian artery and vein, between the pericardium and the pleura, to the diaphragm, to Avhich it is distributed (rami THE NERVOUS SYSTEM. 335 Fig. 121. 336 HUMAN ANATOMY. phrenicoab dominates) . The course of the two nerves differs in the thorax (vide Mediastinum). The beachial plexus (plexus brachialis) is formed by the anterior branches of the four lower cervical and the hrst upper dorsal nerves — the fifth, sixth and seventh forming one cord, and the eighth cervical and first dorsal another cord. Below the line of the clavicle (pars infracldvicularis) both these trunks divide, the adjacent cords of the two upper uniting to form the posterior (fasciculus posterior) , and the remaining cords forming the outer (fasciculus lateralis) and inner (fascic- ulus medius) cords respectively, receiving their names from their relative position to the subclavian artery. Each of these cords again bifurcates, the adjacent divisions at the outer end- ing of the cords, uniting over the artery to form the median nerve, the other divisions forming the musculocutaneous, ulnar, circumflex and musculospiral, the two latter being the divisions of the posterior cord. Its branches are : — Above the clavicle (pars supraclavicularis) : — Communicating, arises from the fifth cervical, and passes to the phrenic; Muscular (rami muscular es), supply the scaleni, rhom- boidii, longus colli and subclavius; Posterior thoracic (n. thoracalis longus), long thoracic, or external respiratory of Bell, arises by five roots from the fifth to seventh cervical nerves, which unite within the scalenus medius muscle and descend to supply the serratus magnus; Suprascapular (n. suprascapularis) , passes beneath the tra- pezius, through the suprascapular notch, to supply the shoulder- joint and supraspinatus muscle. Below the clavicle (pars subclavicularis) : — The anterior thoracic nerves (nn. thoracales anteriores) are two in number : an external branch from the outer cord and an internal from the inner cord. The former supplies the pec- toralis major and the latter the pectoralis minor, and sending branches to the pectoralis major. The subscapular nerves (nn. subscapulares) are three in number. The upper subscapular supplies the subscapular mus- cle, the lower subscapular nerve supplies the teres major and subscapularis, the middle or long subscapular supplies the latis- simus clorsi. The circumflex nerve (n. axillaris) accompanies the pos- terior circumflex artery, passing through the space formed be- tween the teres major, teres minor and long head of the triceps to supply the shoulder joint and skin over the shoulder and the THE NERVOUS SYSTEM. 337 neighboring muscles. It gives off two branches, an upper and a lower branch. The musculocutaneous nerve (n. musculocutaneus) pierces the coracobrachial muscle and descends the arm to supply the brachialis anticus. biceps, coracobrachial and the skin of the forearm. At the outer border of the tendon of the biceps, above the elbow, it becomes superficial and divides into two brandies — the anterior descending the radial side of the forearm to the wrist and supplying the skin of that part, the posterior branch descending the back part of the radial side of the forearm to the wrist. It supplies skin of lower third of forearm. The internal cutaneous nerve (n. cvianeus antibracliii me- dialis) descends the inner side of the arm together with the basilic vein to about its middle, where it becomes cutaneous, supplying the skin of this region. It has an anterior and a posterior branch. The lesser internal cutaneous nerve of Wrisberg (n. cuta- neus brachii medialis) is derived from the inner cord, and receives filaments from the eighth cervical, first dorsal and the intercostohumeral nerve, and is distributed to the skin on the inner side of the arm. The median nerve (n. medianus), formed by a root from the outer and inner cord of the brachial plexus, descends the arm, crossing the brachial artery from its outer to its inner side at the bend of the elbow. Its branches are : — - Muscular (rami muscular es) branches, supply all the superficial anterior muscles except the flexor carpi ulnaris; Anterior interosseous (n. intcrosseus antibracMi volaris), supplies all the 3eep anterior muscles except the inner half of the flexor pro- fundus digiforum; Palmar cutaneous (ramus cutan&us palmaris n. mediarri), crosses above the annular ligament, divides into two branches to supply tha ball <»f the thumb and the palmar surface of the hand; Branches 10 the muscles of the thumb (r< riniit icus extern us I . follows the sper- matic cord to supply the cremaster muscle; in female] supplies round ligamen! ; (0) Crural branch in. I umboi ngui nalis ) , descends m the sheath of the femoral vessels to supply the skin in front of the thigh. 1. External cutaneous (n. cutaneus femoris lateralis) emerges below the anterior superior spine of ilium and divides into: — ('/i Anterior braneli, i<> skin of outer and fronl aspect of thigh; (6) Posterior branch, to skin of outer and back aspect of thigh. 340 HUMAN ANATOMY. 5. Obturator (n. obturatorius) , follows the brim and outer wall of pelvis to foramen (obturator), which it pierces to enter thigh. (a) Anterior branch (ramus anterior), supplies the femoral artery; (b) Posterior branch (ram-us posterior), supplies the adductor muscles; (c) Articular branch, supplies the synovial membrane of knee joint. 6. Accessory obturator (n. obturatorius accessorius) , sends a branch to the hip joint and one to join the anterior branch of the obturator nerve. 7. Anterior crural (n. femoralis), the largest branch of the lumbar plexus, descends through the psoas muscle, beneath Poupart's ligament, into the thigh, where it divides into an anterior or cutaneous branch, and a posterior or muscular branch. It supplies all the muscles and the front of the thigh, excepting the tensor vagina? femoris, gives branches to the knee, and supplies the skin of the inner side in front of the thigh, and to the leg and foot. Its branches from the anterior division are : — (a) Middle cutaneous, to sartorius muscle and skin of front of thigh ; ( b ) Internal cutaneous, supplies the skin in the inner aspect of the leg; (c) Long saphenous (n. saphenus) , or internal saphenous, passes through Hunter's canal, accompanies the internal saphenous vein, to supply the skin of the inner side of the foot. From the posterior division : — (a) Muscular, to the muscles of the anterior and lateral aspects of the thigh; (b) Auricular, to the knee joint. The sacral nerves (nn. sacrales), five in number, divide into anterior and posterior nerves. The upper four sacral nerves, with the fifth lumbar, and a filament from the fourth, (the latter two forming the lumbosacral cord) together make the sacral plexus (plexus sacraUs). It lies upon the anterior sur- face of the pyriformis muscle, and is separated from the viscera, the sciatic and pudic branches of the internal iliac artery by the pelvic fascia. Its branches are: — 1. Muscular (rami muscular es) , supply the obturators, gemelli, quadratus femoris and pyriformis. 2. Superior gluteal (n. glutozus superior), supplies the glutei muscles and tensor vaginas femoris. THE NERVOUS SYSTEM. 341 3. Pudic (n. pudendus), accompanies the internal pudic artery, passing out of the great sacrosciatic foramen, around the spine of the ischium to re-enter the lesser sacrosciatic fora- men. It gives off: — (a) Inferior hemorrhoidal (n. hcemorrhoidalis inferior), supplies the skin of the anus ; (b) Perineal (n. perinei) , follows the course of the superficial perineal artery to supply the perineal structures generally; (c) Dorsal nerve of the penis (to. dorsalis penis), follows the course of the corresponding artery to supply the skin of the glans, prepuce and penis generally. 4. Small sciatic (n. cutaneus femoris posterior), supplies the skin of the thigh, leg, perineum and the gluteus maximus muscle. Its branches are: — (a) Inferior gluteal, to the gluteus maximus; (b) Internal cutaneous branches (toto. cluniuin inferiores laterales), to the skin of the inner and upper part of the thigh, and one branch, the inferior pudendal, supplies the skin of the scrotum in the male and labium in the female; (c) Ascending cutaneous branches supply the skin over the gluteus maximus. 5. The great sciatic nerve (n. isckiadicus) , the largest in the body, passes out of the great sacrosciatic foramen, and descends between the tuberosity of the ischium and the great trochanter to the lower third of the thigh, where it divides into the internal and external popliteal. It gives off: — (a) Muscular branches (rami musculares) , to the muscles on the inner and posterior aspect of the thigh ; (b) Articular branches, to the hip joint. The internal popliteal (n. tibialis), descends through the popliteal space to the arch of the soleus muscle, where it becomes the posterior tibial. It gives off: — (a) Articular branches (rami articulares) , to knee joint; (6) Muscular branches (rami musculares), to the muscles on the posterior aspect of the leg; (c) Kxicrnal, or short saphenous nerve (n. suraUs), passes be- tween the two heads of the gastrocnemius muscle, and descends the leg. receiving the comnxunicans peronei branch from the external popliteal, around the outer malleolus to supply the skin of the outer side of the foot. The 'posterior tibial nerve descends the leg in company with the posterior tioial vessels to below Hie inner ankle, where it divides into the external and internal plantar nerves. It gives oil':— 342 HUMAN ANATOMY. 1. Muscular branches (rami musculares) , to the deep muscles of the calf; 2. Internal cancaneal (rami aalctmei mediates), supplies the skin on the inner side of the sole and heel; 3. Articular branch, to ankle joint. The internal plantar (n. plantaris medialis) accompanies the corresponding artery to the inner side of the foot and gives off:— 1. Cutaneous branches; 2. Muscular branches; 3. Articular branches; 4. Four digital branches, supplying both sides of the first three toes and the inner side of the fourth. The external plantar (n. plantaris lateralis) supplies by a superficial branch the outer side of the fourth and both sides of the fifth toes, and gives off a deep, or muscular, branch. The external popliteal or peroneal nerve (n. peronceus com- munis) descends to the inner side of the biceps tendon, pierces the peroneus longus about one inch below the head, and divides into the anterior tibial and musculocutaneous. It gives off: — 1. Articular branches (rami articulares) , to knee joint; 2. Cutaneous branchesi (n. cutaneus surce lateralis), one of which is the communicans peronei (ramus anastomoticus peronceus or com- municans fibularis) , which joins the short saphenous. The anterior tibial (n. perona?us profundus) accompanies the corresponding artery on its outer side to the ankle, where, after aivingr off an articular branch, it divides into: — 1. External or tarsal branch, to the tarsal and metatarsal joints, and 2. Internal, to the dorsal adjoining sides of the great and second toes. The musculocutaneous (n. peronceus superficialis) gives off:— 1. Internal branches, (n. cutaneus dorsaUs medialis), supply skin of inner side of foot and ankle, and inner side of great toe; 2. External branch (n. cutaneus dorsalis intermedium) , to the dor- sum of the adjacent sides of the second, third, fourth and fifth toes. /* Fig. 123. Sacral plexus and branches. s First cervical < nerve, Second cervical nerve Third cervical - nerve, Fourth to eighth ■{ cervical nerves, THE XERVOIS SYSTEM. 343 Table of the Spinal Nerves. Complexus, recti and obliqui muscles. Posterior d his ion 1 suboccipital) Anterior division (occipital), Posterior division, Anterior division, Posterior divison, Anterior divison. Posterior divisions, Unites with the second cervical nerve, and supplies the anterior recti and rectus lateralis muscles. External "> Splenitis, complexus, tra- branch, / cheloinastoid muscles. , ,' ! ^Yith branch from third branch, i . . ,. ,. (great ' cervical, supplies the occipital), j skin of the scalP- f Small occipital, j Branch to superficial cervical, I Branch to great auricular, ^ Branch to eommunicans noni. f External ~] 0 , ] _ and [Splenius } internal < [_ branches, semispinals, complexus, trachelomas- toid muscles. Branches to form the great auricular, superficial cervical, and communican- tes minor. Internal branc 11a 1 \ Miisclea of the side of the 'foes, f neck. sternal ") 'anches, / External bm Skin and larger muscles of the neck. [ Branch to phrenic, Anterior division] Branches to the trapezius, scalenus j medius, levator anguli scapulae mus- I cles. of fourth, Anterior divisions, fifth to eighth, , , , ( Superficial Cervical plexus, 1 J. branches, Unite with the first dorsal nerve to form the brachial plexus. ( Superficialis colli, Ascending. - Auricularis magnus, ( < Occipitalis minor. Descending. Supraclavicular. 1 Permed by the anterior divisions of the first to the fourth cer- vical nerves, 344 HUMAN ANATOMY. Table of the Spinal Nerves.— (Continued.) Cervical plexus f ( continued ) , \ Deep branches, \ f Internal set, I Brachial plexus,1 < Above the clavicle, Below the clavicle, Three sub- scapular, Circum- flex, Musculo- cutaneous, External set, Communi- cating, Muscular, Posterior thoracic, Supra- scapular, Anterior thoracic, Communicating, Muscular, < , t muscles. Phrenic, Communicans noni. Muscular, Communicating. To phrenic. Scaleni, rhomboidei, lon- gus colli, subclavius. Serratus magnus. ( External respiratory nerve of Bell.) Shoulder joint and supra- spinatus muscle. External branch, Internal branch, Upper, Lower, Middle, Upper, Lower, Anterior branch, f Pectoralis \ major. f Pectoralis J minor, i, Pectoralis ( major. [ Subscapular \ muscle. Teres major. f Latissimus 1 dorsi. f Shoulder j joint, and \ muscles and [ skin about it. f Integument j of palmar sur- i, face of the j wrist. f Integument Posterior j of dorsal sur- branch, \ face of the | wrist. i Formed by the anterior branches of four lower cervical and first dorsal nerves. THE NERVOUS SYSTEM. 345 Table of the Spinal Nkkvks.— (Continued.) Brachial plexus, below the clavicle contirid, Internal f Anterior ] Integument of a nd internal j auu. inner side of cutaneous, 1 posterior f the arm. branches, J Lesser internal [integument of the inner cutaneous < sjcje 0f h,a arrrL (Wrisberg's), [ Median, - f Muscular, Anterior interosseous, Palmar cutaneous, Branch to the musclesi of the thumb, Digital branches, 1 ( Superficial anterior mus- J cles, except flexor carpi ( ulnaris. fDeep anterior muscles, ex- cept the inner half of flexor profundus digi- torum. J Ball of thumb and palmar \ surface of the hand. Opponens, outer head of small flexor and ab- ductor. fBoth sides of thumb, in- : dex and middle fingers i and radial side of ring [ finger. Articular, j (Elbow) Ulnar, in forearm,' f Inner half of deep flexors Muscular, j and flexor carpi ulnaris. Cutaneous, { Skin of the palm. (Both sides of little finger | and ulnar side of ring [ finger. Dorsal cutaneous, Articular, { (Wrist). I In. ir, in hand, Superficial palmar, Deep palmar, f Skin of both sides of little j linger, ulnar side of ring \ finger and palmaria brevia muscle. f luicrossei lumbricales, ad : ductor pollicis, inner \ head of flexor brevis pollicis. 34G HUMAN ANATOMY. Table of the Spinal Nerves. — (Continued.) Brachial plexus, below the clavicle, cont'd, Musculo- spiral, Dorsal nerves, Posterior divisions, Anterior divisons,i Lumbar nerves, Posterior divisions, Anterior _ divisions, \ f Supinator longus, extensor Muscular j carpi radialis longior, tri- branches, i ceps, anconeus, brachialis [ anticus. f Supplies adjoining sides of _ ,. , j thumb and index finger, Kadial, -j index and middlej middla [ and ring fingers. (Inner side of arm, elbow, radial side of forearm and wrist. f Posterior brachial and radial Posterior j muscles, except those sup- interosseous, i plied by ulnar (muscular [ branches ) . External branches, Internal branches, Upper six ( thoracic intercostals ) , ^Muscles and skin of the back. First intercostal crosses the axilla to join the lesser in- ternal cutaneous ( Wris- berg's ) . Lateral cutane- ■< ous, Ante- rior, Poste- rior, f Skin of ■j chest and (. mamma?. Skin over scapula and lower dorsal region. r. Lower six i The ]agt dorgal gWes off the (thoracico- ^ dorsilumbar to the quad- abdominal , ratus lumborum muscle, intercostals ) , [_ External branches, Internal branches, I Muscles of back. the skin of the f The four upper unite to form the lumbar plexus. The fifth, with a branch from the fourth, joins the sacral nerves to form the lumbosacral cord. I l To the walls of chest and abdomen. THE NERVOUS SYSTEM. 347 Table of the Spinal Nerves. — (Continued.) Ilio- hypogastric, Lumbar plexus. f Skin of gluteal region, ob- lliac, I lique muscles of abdomen. Hypo- I Skin of hypogastric region gastric, I and oblique muscles. Ilio- inguinal, Genito- crural, External cutaneous, ( Skin of inner and upper part of thigh, scro- ll turn, labium (in female). f Cremaster muscle, round liga- Genital, | ment (in femaie). Posterior, Anterior, Obturator, \ Posterior, Articular, Crural, { Skin of front of thigh. ( Skin of outer front aspect of Anterior, | thigh Skin of outer back aspect of / Skin of \ thigh. I Femoral artery. < Adductor muscles. Is" Synovial membrane of knee- joint. Accessory f Branch to hip- joint, branch to anterior obturator, \ branch of obturator nerve. f Sartorius Middle j muscle, cutaneous, \ skin of [_ thigh (ant,). f Skin of Internal j inner cutaneous, 1 aspect of leg. Anterior, division, Anterior crural, Skin of -, inner side of saphenous, j foot> Lone Posterior, division] f Muscles j of the an- _ _, . terior and f Muscular. J ];it|,n|| aspect of J [ thigh. [_ Articular, j Knee joint. 348 HUMAN ANATOMY. Table of the Spinal Nerves— (Continued. )_ Sacral nerves, ! Posterior I divisions, External and internal branches, Muscles and skin of the back. | Anterior f The four upper joining with the lumbosacral [ divisions, \ cord form the sacral plexus. Sacral plexus, Obturators, gemelli, quadratus femoris and Muscular, | piriformis. Superior j Qiu^ei muscles and tensor vaginae femoris. Inferior f gkin of ^ anug rhoidal, ( rineal structures. hemor Pudic, ■{ Perineal, < Peri < Skin of the penis. To hip joint. i Gluteus maximus. Dorsal nerve of the penis, Articular { Small sciatic, Inferior gluteal, Great sciatic. Internal cutaneous, Ascending cutaneous, Articular, Muscular, f Branch to skin of inner and upper part of thigh. H Inferior pudendal, skin of scrotum, labium in fe- ^ male. I Skin over gluteus maximus. -J Hip joint. / Muscles of inner and pos- \ terior part of thigh. Bifurcation, { ^Sal,' } PoPliteaL Internal popliteal, Articular branches, To knee joint. Muscular "1 To muscles of the posterior aspect of branches, J the leg. THE NERVOUS SYSTEM. 349 Table of the Spinal Nerves. — (Continued.) Conmmnicans \ skin of outer side of the foot poplitei, j Internal popliteal [continued) Posterior tibial (continuation), , ( Deep muscles of the Muscular, j c.{,f Plantai cutaneou Skin of inner side of sole and heel. :, { Articular, j To ankle joint. j Internal plantar, Bifurca- tion, 1 External plantar. Internal f Cutaneous, Muscular, plantar, \ Articular, '[_ Four digital. External plantar, Supplies the outer side of the fourth and both sides of the fifth toes. Mus- cular branch. External |H)pliteal * ( peroneal), Articular, j To knee joint. Two cutaneous, Anterior tibial, M usculo- cutaneous, Communieans peronei. External or tarsal branch, J Internal branch. Internal branches, External branch, 1 Tarsal and metatar- sal joints. f Inner and dorsal, ad- j joining sides of \ great and second I toes. < To skin of inner side \ of foot and ankle. f To dorsum of adja- J cent sides of third, fourth toes. and fifth 350 HUMAN ANATOMY. THE SYMPATHETIC NERVOUS SYSTEM. The sympathetic system (sympatheticus) , like the cerebro- spinal axis, is double, consisting of a gangliated cord {truncus sympatheticus) on each side, extending the entire length of the vertebral column, and numerous nerve fibers, both communicat- ing, by which it anastomoses with the cerebrospinal axis, and distributory, by which the blood vessels and viscera are supplied. Branches of the gangliated cords . ascend through the carotid canal to communicate with the ganglia of the fifth "cranial nerve, and also with each other through the ganglion of Ribes, situated upon the anterior communicating artery. They also communicate below in the ganglion impar (ganglion coccyg- eum impar), located in front of the coccyx. The ganglia of each cord correspond very nearly in posi- tion and number to the vertebra?, except in the cervical region, where there are but three : cervical three, dorsal twelve, lumbar four, sacral five. From the gangliated cords three important plexuses are given off — the cardiac, solar, and hypogastric — situated in the thoracic, abdominal and pelvic cavities, respectively. Cranial Ganglia — Ganglia Connected ivitli the Fifth Cranial Nerve. — Besides the Gasserian ganglion upon the root there are four — ophthalmic, sphenopalatine, otic and submaxillary — each of which has three branches of communication or roots — motor, sensory and sympathetic — besides several branches of distribu- tion. The ophthalmic, or ciliary ganglion (ganglion ciliare), is ' situated in the orbit between the external rectus muscle and I optic nerve upon the first division of the fifth cranial nerve. Its sensory root (radix longa ganglii ciliaris) is from the nasal branch of ophthalmic, motor root (radix brevis ganglii ciliaris) from the motor oculi or third, and sympathetic from the cavernous plexus. Its branches are the short ciliary nerves (vide Eye). Sphenopalatine, or Meckel's (ganglion sphenopalatinum) , the largest, is situated in the sphenomaxillary fossa upon the superior maxillary, or second division of the fifth. Its sensory root is derived from the superior maxillary, its motor root, from the facial, through means of the Vidian, and its sympathetic from the carotid plexus. Its branches are : — 1. Ascending (rami orbitales) ; } -Irterv To*UtJt*\ 7c TrTnptiftiehra/H'Ji ■jfolos. From l**£trrz£aLi \W*-* fn n J>t tm 3 ~?i'.r> .-. j/. I 'cm >. Ophthalmia -rtasit. It trre V^jryi Superficial flrf/wKz//rv/*J'asia/A*rrr i /reuniting fc-r*0m/4r&/ , .tt./ its 6rajt&M . ..!>,.; - RnmSfJlmtal Xern ^, Srvmf^Jij'.a/ A'tri't "~~^*J A^-yi.-reponyir,,! lr,iri,-Acl, .,r' .iW..*™*,^ ^ .^ »mVtlu,„!,„r.Vv, /'" JW^Kn £^& From. CoccfgealXerrx Fig. 124. The sympathetic system. THE NERVOUS SYSTEM. 35] f Anterior, or large palatine (». pilar Unas and rior 1 . 2. Descending, or pala- ! Middle, or external palatine (». pala- tine [mi. palatini \ , j /i'hi/s melius), \ Posterior, or small palatine (». pato- is tinus posterior) , f Pos { Na Posterior superior nasal (rami namks 3. Internal, -{ posteriores superiores), Fasopalatine (n. nasopaiat&nus) , 4. Posterior, Vidian (>i. canalis pterygoidea), Large superficial petrosal (n. petrosus supcrficialis major) , Deep petrosal (n. petrosus profundus), Pharyngeal branch. The Vidian nerve (n. canalis pterygoidea) passes from the baj?k part of Meckel's ganglion through the foramen laeerum medium, where it divides into the large superficial and deep petrosal nerves. Jt gives off the upper posterior nasal branches to the mucous membrane of septum, orifice of Eustachian tube. and roof of the nose. Or, more correctly, the Vidian is formed by the union of the great petrosal (large superficial petrosal) from the facial and the large deep petrosal from the carotid sympathetic, runs forward through the Vidian canal, and joins the sphenopalatine ganglion (Meckel's). In this description, the nerves given off to the nasal mucous membrane must be considered branches from the ganglion inclosed in the same sheath. The large superficial petrosal branch (n. petrosus super- firiahs major) enters the cranium through the foramen laeerum medium, passes beneath the dura mater and Gasserian ganglion, enters the hiatus Fallopii, receives a branch from the tympanic branch of the glossopharyngeal (Jacobson's), and through the aqueductus Fallopii to terminate in the geniculate ganglion of the facial nerve. The large deep petrosal branch (n. petrosus profundus) crosses the foramen Laeerum medium to the carotid canal, where it joins the carotid plexus of the sympathetic. The pharyngeal or pterygopalatine nerve descends from the back part of the ganglion through the pterygopalatine canal, to supply the upper part of the pharynx. Besides the Vidian and its branches there are two other petrosal nerves (vide Facial Sfvw), the small and external petrosaL 352 HUMAN ANATOMY. The small petrosal connects the geniculate ganglion of the facial;, within the aqueductus Fallopii, with the otic ganglion. The external petrosal connects the geniculate ganglion of the facial, within the same canal, with the sympathetic plexus of the middle meningeal plexus. The otic or Arnold's ganglion (ganglion oticum) is placed below the foramen ovale upon the inferior maxillary nerve, or third division of the fifth. Its sensory root is derived from the auriculotemporal branch of the inferior maxillary ; the motor root, from the inter- nal pterygoid branch of the same; the sympathetic root,, from the plexus on the middle meningeal artery. Branches are dis- tributed to the tensor palati (n. tensoris veil palatini) and ten- sor tympani (n. tensoris tympani) muscles. The submaxillary ganglion (ganglion submaxillare) is placed above the deep portion of the submaxillary gland. Its sensory root is derived from lingual branch of inferior maxillary; its motor root, from the facial nerve through a branch of the chorda tympani; and its sympathetic, from the plexus of the facial artery. Besides these ganglia, situated upon branches of the tri- facial nerve, the following are found within the cavity of the cranium : — Ganglion of Ribes, on the anterior communicating artery; Ganglion of Bidder, on the middle meningeal artery; Ganglion of Laumonier, on the internal carotid artery; Ganglion of Cloquet, on the nasopalatine nerve in the incisor fossa; Ganglion of Bochdaleck, near the sphenopalatine ganglion. Cervical Ganglia. — The cervical ganglia consist of three, — superior, middle, inferior, — of which the superior is the largest and the middle the smallest. These are connected above with the cranial ganglia, below with the thoracic ganglia, and com- municate with each other. The superior cervical ganglion (n, caroticus internus) lies opposite the second or third cervical vertebra, behind the inter- nal carotid artery. It is connected with the middle ganglion, the upper four cervical, and tenth and twelfth cranial nerves, distributes branches to the carotid (internal), cavernous (plexus cavernosus) and pharyngeal plexuses, and gives off the superior cardiac nerve to the cardiac plexus (plexus caroticus internus). The middle cervical ganglion (ganglion cervical medium) lies upon inferior thyroid artery, opposite fifth cervical vertebra. It is connected with the upper and lower ganglia and spinal THE NERVOUS SYSTEM. 353 nerves, and gives off the middle cardiac nerve (n. cardiacus medius) to cardiac plexus. The inferior cervical ganglion (ganglion cervicale inferius) lies internal to the superior intercostal artery, below the last cervical vertebra. It is connected to the middle ganglion, first thoracic, lower cervical nerves, forms the vertebral plexus, and gives off the inferior cardiac nerve (n. cardiacus inferior) to cardiac plexus. Thoracic ganglia (pars thoracalis) lie upon the heads of the ribs on each side of the vertebral column. They are con- nected with the inferior cervical ganglion above, the lumbar ganglion below, the dorsal spinal nerves behind, and give off internal branches divided into two sets of six each — -upper and lower. internal branches from upper set are distributed to the pulmonary and aortic plexuses (plexus aorticus thoracalis), and internal branches from lower set unite to form the three splanch- nic nerves — great splanchnic (n. splanchnicus major), lesser Bplanchnic (n. splanchnicus minor), renal splanchnic, — distrib- uted respectively to the semilunar ganglion, renal and supra- renal plexuses, and to renal and celiac plexuses. The lumbar ganglia (pars lumbalis) lie along inner mar- gin of the psoas muscle. They communicate above and below with the other ganglia and with the lumbar spinal nerves, and give off internal branches which form the hypogastric plexus. The sacral ganglia (pars sacralis) are situated internal to the anterior sacral foramina. They unite below in front of the coccyx iti the coccygeal ganglion, or ganglion impar. They communicate with the sacral nerves, join the pelvic plexus, and scud branches on the middle sacral artery. Cardiac Plexus (plexus cardiacus). — The cardiac plexus. formed from the superior, middle and inferior cardiac nerves from the cervical ganglia, and the cardiac branches from the pneumogastric and recurrent laryngeal, consists of two portions — the superficial and deep. The superficial cardiac plexus, situated beneath the arch of the aorta, is formed by the left superior cardiac nerve and the left inferior cervical cardiac nerve, and contains the cardiac ganglion of Wrisberg (ganglion cardiacum [Wrisbergi] ) . It form- part of the anterior coronary plexus, and sends branches to the left anterior pulmonary plexus. Tin' deep cardiac plexus lies between the arch of the aorta and trachea, and receives all the cardie nerves except the two mentioned above. It forms part of the anterior coronary 23 354 HUMAN ANATOMY. {plexus coronarius cordis anterior) and posterior coronary {plexus coronarius cordis posterior) plexuses. The solar plexus {plexus cceliacum), or "abdominal brain," consists of ganglia and a network of nerve branches, formed chiefly from the branches of the two great splanchnic nerves and branches from the right pneumogastrie. It is sit- uated between the aorta and the crura of the diaphragm and the stomach, and surrounds the superior mesenteric artery and celiac axis. Its ganglia are two crescentic ganglionic masses — the semi- lunar ganglia {ganglia coeliaca) — the largest in the body — sit- uated in front of the crura of the diaphragm. They are com- posed of small ganglia aggregated together. From the solar plexus and semilunar ganglion are derived numerous branches which are distributed as plexuses over all the abdominal arteries, as follows: — Cceliac {plexus cceliacus) , Spermatic (plexus spermaticus) , Gastric [plexus gastricus supe- Aortic (plexus (tortious abclomi- rior) , nalis) , Hepatic (plexus hepaticus) , Superior mesenteric (plexus mes- Splenic (plexus lienalis), entericus superior), Phrenic (plexus phrenicus), Inferior mesenteric (plexus rnes- Suprarenal (plexus suprarenalis) , entericus inferior), Renal (plexus renalis) , Ovarian (plexus arterke ovaricce), The hypogastric plexus {plexus hypogastricum) is sit- uated below the bifurcation of the aorta, in front of the sacrum, and is formed by branches from the lumbar ganglia and aortic plexus. It divides into two parts which, with branches from the sacral ganglia and nerves, become the pelvic plexuses. The inferior hypogastric or pelvic plexus on each side gives off the following branches : — Vaginal plexus, Inferior hemorrhoidal (plexus Tuemorrlioidalis inferior) , ( Small cavernous "| nerves (n. oavernosus j Prostatic plexus/ penis minor), \ (plexus prostaticus) ,\ Large cavernous j " ' nerves (n. oavernosus I (_ major ) , J Vesical plexus (plexus vesicalis), Uterine plexus. ORGANS OF SPECIAL SENSE. T 1 1 E NOSE. The nose (organon ol) 'actus), the special organ of smell, consist* of two parts, — the external prominence, or nose proper, and the internal cavities, or nasal fossae. fc&-"NASAL SEPTA OF S L branch) UPERIOR r ASAL 1 Fig. 125. The olfactory nerves and nerves of common sensation to the nose. (Ei-kley.) The nose proper is made up of a cartilago-osseons frame- work covered with muscles (vide Muscles) and skin, and lined with mucous membrane. The osseous portion is formed by the margins of the anterior meatus (vide Osteology). The cartilaginous portion consists of an upper (cartilago nasi lateralis) and lower lateral cartilage (cartilago alaris major) on either side, and the nasal septum dividing the nasal cavity into two nasal fossae durum nasi). Vrtenes are from nasal branch of ophthalmic and infra- orbital, nasal artery From superior coronary, and branches of the lateralis nasi. (355) 356 HUMAN ANATOMY. Veins empty into the ophthalmic and facial veins. Nerves from infratrochlear, infraorbital and facial. The nasal fossae (cavum nasi) are the two bony cavities already described (vide page 45). They are lined throughout with mucous membrane (membrana mucosa nasi) called Sclmeiclerian membrane, which is continuous with the antrum of Highmore, frontal, ethmoidal and sphenoidal sinuses, with the conjunctiva through the lachrymonasal duct, with the pharynx through the posterior nares (choanal), and with cavity of the tympanum through the Eustachian tube. Its epithelium is columnar, ciliated in the lower part and in the. sinuses, and is columnar, but not ciliated, in the membrane to which the olfac- tory nerve is distributed. Arteries are anterior and posterior ethmoidal from the ophthalmic to the roof, frontal and ethmoidal sinuses, spheno- palatine from the internal maxillary to the mucous membrane of the spongy bones, septum, and meatuses, and alveolar from the internal maxillary to the antrum. Veins empty into facial, ophthalmic, and beginning of great longitudinal sinus. Nerves are : — Olfactory, or first cranial, the special nerve of smell, is dis- tributed over the upper third of the septum and superior turbi- nated bones. Anterior dental branch of superior maxillary to the inferior turbinated bone and inferior meatus. Nasal branch of ophthalmic, to the outer walls and septum. Besides these, the sphenopalatine ganglion, the Vidian, the supe- rior nasal branch, nasopalatine and anterior palatine also send branches to the parts. THE EYE. The organs of sight comprise the eyeballs (bulbus oculi) and their appendages and the optic nerves. The eye is a spherical organ, situated in the anterior part of the skull, protected in front by several appendages, acted upon by muscles, and supplied by blood vessels and nerves. Within the orbit it rests upon a bed of fat, from which it is separated by a membranous sac — the capsule of Tenon (fascia bulbi [Tenoni]). It presents the following : — Diameters. — Anteroposterior, in the adult, .950 inch; transverse, .925; vertical, .915; oblique, .9-13. ORGANS OF SPECIAL SENSE. 357 Fig. 126. Nasal meatuses and accessory cavities. 1, right termination of the left frontal sinus; 2, right frontal sinus; 3, probe extending from the right frontal sinus through the infundibulum into the right nasal fossa; 4, ethmoid cells; 5, large opening into the maxillary sinus; 6, anterior antrum of the sphenoid bone; 7, posterior sphenoid antrum; v. middle nasal meatus; 9, Inferior meatus; i<», Inferior turbinated bone; 11, probe extending through the nasal dint. (Bishop.) 558 HUMAN ANATOMY. Anterior and posterior poles are the geometric centers of the cornea and fundus respectively. Optic axis (axis optica) is a straight line passing through the cornea and posterior pole of the eye. Line of vision, or visual axis (linea visus) , is an imaginary line which connects the point of fixation with the fovea cen- 81du8 front, yifttnltia papyr. oss. ethmoid. Foramina ethmoid, ?roc. orbit, oss. palat. Corpus oss. sphenoid. For. optic ffosura orbit, eup. Sella turcica \ Spina trochlear. j Bars orbit, oss. front. ! Fossa sacci lacrim. Oa nasale 'acies orbit. maxQl. Froo. uncin. oss. ethmoid. Froo. ethmoid, concha* nasal, inf. Proo. max ill. conchae nasal, inf. canal, palat. Sinus maxiliarial PaTs perpondic. oss. palat. Fig. 127. Inner wall of orbit and adjacent parts. (His.) (Ball.) tralis, through the nodal point, usually to the outer side of the center of the pupil. It forms with, the optic axis as it cuts the cornea, the visual angle — an angle of from 3° to 7°. Nodal point is an imaginary point — the center of curvature of the refracting media — where all the luminous rays pass with- out deviation. Equatorial plane, an imaginary plane passing through the center of the eyeball at right angles to the optic axis, dividing the globe into two hemispheres — the anterior and posterior. ORGANS OF SPECIAL SENSE. 359 Equator is the line upon the surface of the glohe where the equatorial plane cuts it. Meridional planes are imaginary anteroposterior planes co- inciding with the axis. Meridians are the lines upon the surface where these merid- ional planes cut it. The capsule of Tenon, tunica vaginalis oculi (fascia oculi [Tenoni]), is a fascia between the eyeball and the walls of the orbit, isolating the eyeball and allowing free movement. Fig. 128. Horizontal section of right orbit. 1, Horner's muscle; 2, septum or- bitale; '.',, fibers of the orbicularis palpebrarum muscle; 4, tarsal plate; 5, conjunctival sac; 6, outer palpebral ligament; 7, temporal muscle; S, wall of the orbit; 9, external rectus muscle; 10, orbital fat; 11, optic nerve; 12, internal rectus muscle; 13, inner check ligament; 14, inner wall of the orbit; 15, attachment of the capsule of Tenon (1G) to the conjunctiva; 17, Tenon's space; 18, outer check ligament; 19, lachrymal gland; 20, ciliary process; 21, iris. (Ball after Gerlach.) It i- continuous in front with the subconjunctival connective tiss iw. and behind with the dura mater, through the sphenoidal fissure and optic foramen, and consists of two layers — a visceral investing the posterior portion of the eyeball, and a parietal lining the cushion of fat on which the eye rests. The inner aspect is lined with flattened endothelial cells, and incloses a lymph Bpace communicating with subdural and subarachnoidean lymph spaces of the optic nerve sheath. It sup- 360 HUMAN ANATOMY. ports the lachrymal gland, is strengthened by numerous fibrous bands, and is pierced by the ocular muscles, inclosing them in imperfect sheaths. The capsule of Bonnet is the name given to the portion posterior to the passage of the tendons, and The capsule of Tenon is then applied to the anterior socket- like half. The eyeball (biilbus oculi) is composed of a large, opaque segment of a sphere, forming about five-sixths of the globe, for- the protection of its contents, and a smaller transparent seg- ment of a sphere, implanted upon and continuous with it in Canal of Schlemm. 8S~^*^^Canal of «s» Petit. SlIM'A INTERNAL ' RECTUS MUSCLE. Canal for central artery. Optic nerve. front. The optic nerves enter the eyeballs to their nasal side, in the direction of the axes of the orbit. It is composed of three tunics or coats : — 1. Sclerotic and cornea, 2. Choroid, iris and ciliary processes, 3. Retina; And three humors, -or refracting media : — 1. Aqueous humor, 2. Crystalline lens (and capsule), 3. Vitreous humor, or body. ORGANS OF SPECIAL SENSE. 331 The sclerotic coat, so called from its extreme hardness, is thicker behind (one twenty-tilth of an inch) than in front (one-sixtieth of an inch), and presents two surfaces for study — the external and internal. The external swrface is smooth and white, and has attached to it the various muscles of the eye. The internal surface is grooved for the passage of the ciliary nerves, and connected to the choroid by a fine areolar layer — the lamina fusca (laminae fascice sclerce). Behind, and a little to the nasal side, where the optic nerve enters, it presents a perforated appearance — the lamina cribrosa —the larger opening of which — the poms opticus — transmits the arteria centralis retinae; the others transmit the ciliary ves- sels and nerves. It is continuous in front with the cornea, over- lapping it a little on its outer margin. In structure it is made up of white, fibrous tissue, with a small quantity of elastic fibers and connective-tissue corpuscles. The arteries are from the anterior and short posterior ciliary. The veins — venous blood removed by venas vorticosse and the anterior ciliary. The nerves are from the ciliary. The cornea is the convex, transparent, nearly circular tissue forming the anterior one-sixth of the globe. It is from niie twenty-second to one thirty-second of an inch in thickness. Its thickness at the periphery is 1.12 millimeters, hence its pos- terior surface is more curved than the anterior. Its transverse diameter is a. little greater than the vertical, owing to the over- lapping of the sclerotic above and below. It is composed of four layers : — 1. Conjunctival epithelium, or anterior layer {epithelium cornea;), 2. Cornea proper, or sub- f Anterior elastic lamina (lamina stantaa propria, \ elastica anterior), 3. Posterior elastic lamina "| I lamina elastica paste- j ' ""' 1 1 ! Cornea proper; 4. Endothelial lining (endo- (Membrane of Deseemet. thelium camercB ante- | i ioris) , The conjunctival epithelium consists of several layers of cells (columnar, polyhedral and squamous) covering the ante- rior surface of the cornea, continuous wiih the conjunctiva. The anterior clastic lamina is the name given to the outer epithelial layer of the cornea proper. 362 HUMAN ANATOMY. The cornea proper is made up of a transparent fibrous structure, identical with the sclerotic, and consisting of about sixty layers or lamina, connected by a cement substance inclos- ing corneal spaces, each of which contains a corneal corpuscle. The posterior elastic lamina consists of an elastic homo- geneous membrane, internal to the proper structure of the cornea, and constituting, with the epithelial lining, the mem- brane of Descemet, or Demours. The structure of the cornea is non-vascular, being nour- ished by channels representing lymphatic vessels, and continu- ous with the corneal spaces. The nerves derived from the ciliary nerves are numerous, and form between the outer surface of the cornea proper and the epithelial covering the subepithelial plexus, from which is given off* the intraepithelial plexus. The second tunic consists of the choroid, lining the sclerotic coat throughout; the iris, the circular curtain suspended in the aqueous humor and perforated by the pupil; and the ciliary ligament and muscle at the junction of the two. The choroid (choroidea) is a dark, reddish-brown, highly vascular membrane, lining the sclerotic coat from the optic nerve to the ciliary ligament, where it projects backward in a number of processes, the ciliary processes. It is in contact externally with the sclerotic, to which it is attached by the membrana fusca, and internally with the retina. The structure consists chiefly of a dense capillary network. The choroid proper consists of an external and an internal layer, and is separated from the sclerotic by the lamina supra- choroidea, continuous with the lamina fusca of the sclerotic, . and from the pigmentary layer of the retina by the lamina vitrea. The external layer or the lamina vasculosa consists of a capillary network, derived from the larger branches of the short ciliary arteries, inclosing between the meshes large, star-shaped pigment cells, which are connected together by a delicate stroma. They, terminate in the venw vorticosce, which emerge through the sclerotic midway between the optic nerve and the margin of the cornea. The internal layer is also composed of a fine capillary net- work, derived from the short ciliary arteries> and continuous in front with the vessels of the ciliary processes. It is called the tunica Ruyschiana (lamina choriocapillaris) . The lamina basalis, or membrane of Bruch, is a thin, struc- tureless layer, separating the pigmentary layer of the retina from the stroma of the choroid. ORGANS OF SPECIAL SENSE. 363 The arteries are from posterior ciliary, short ciliary (prin- cipally in the external layer), and recurrent branches of long and anterior ciliary. The veins converge from the vena? vorticosae to form four or five trunks, which pierce the sclerotic midway between the optic nerve and corneal margin to join the cavernous sinus. The nerves are the long and short ciliary. Fig. ISO. Vessels of the eye. A, conjunctival vessels; B, anterior ciliary vessels; 0, vena vorticosa; D, posterior ciliary arteries; E, central retinal vessels. (Hall.) The ciliary processes (processus ciliares) are a series of sixty to eighty pigmented vascular processes arranged circularly around the lens behind the iris, and composed of the two inter- nal layers of the choroid folded inward. They are continuous in front with the ins, and are connected behind with the sus- pensory ligament of the lens. The structure is similar to that of the choroid. Tin: [BIS is a thin, circular curtain, suspended in the aqueous humor, between the lens and cornea, and perforated by a circular aperture — the pupil (pupUla). 364 HUMAN ANATOMY. Its circumference is connected with the choroid, and also by means of the ciliary ligament with the sclerotic and cornea. The sclerotic contains in this position, near its junction with the cornea, a circular canal (lymph canal, or venous sinus), the sinus circularis iridis, or canal of Schlemm (sinus venosus sclera?) . The circumference of the iris is also connected in front with the cornea by pectinate ligament (ligamentum pectinatum iridis), derived from the membrane of Descent et. The structure is composed of four elements : — . 1. A basement membrane and polyhedral cells, continuous with the membrane of Descemet; 2. Stroma (stroma iridis) consisting of bundles of fibrous tissue and cells; f Circular fibers (m. sphincter pupilkc) , ,r , „, ; forming the sphincter of the pupil; 6. Muscular fibers, ^ Radiating fibers (m. dilator pupillw) , [ forming the dilator of the pupil; 4. Pigment, consisting of polyhedral or round pigment cells, dis- tributed for the most part to the posterior surface, which, from its deep-purple tint, has been called the uvea. The arteries are from the long and anterior ciliary and from the ciliary processes (from the short ciliary). The veins join those of the ciliary processes and anterior ciliary veins. The nerves are the long ciliary from the nasal branch of the ophthalmic (first division of fifth), and ciliary branches from the lenticular or ophthalmic ganglion. The circular fibers are supplied by the fibers from the third or motor oculi, the radiating fibers are supplied by the sympathetic. The membrana pupiUaris is a delicate, vascular membrane, closing the pupil in the fetus, continuous at its margin with the iris. It usually disappears about the eighth month, but occasionally persists. The ciliary muscle (m. ciliaris) is a grayish, circular band of unstriated fibers, attached to the fore part of the choroid. It consists of two sets of fibers — the external radiating fibers (fibrce meridianales [Bruckei]), dilator pupillce — arising from the junction of the sclerotica and cornea, and is inserted into the choroid opposite the ciliary processes; the internal circular ORGANS OF SPECIAL SENSE. 365 Fig. 131. The structure of the human retina. I, pigment epithelium layer; II, rods and cones; III, granules of the visual cells; IV, outer plexi- form layer; V, layer of horizontal cells; VI, layer of bipolar cells; vil, layer of amacrine cells; VIII, inner plexiform layer; IX, gan- glion cell layer; X, lay«r ol' nerve fibers; 1, diffuse amncrin.' cell; 2, diffuse ganglion cell; 3, centrifugal nerve fiber; l, amacrine asso- • molecular lay< r i- composed of a- layer of glandular- like structure, forming a reticulum inclosing minute granules. 5. The vn/ner nuclear layer is composed of three sets of nuclear bodies, the first resembling bipolar nerve cells, the second without branches, and third continuous with the radiating fibers, or fibers of VI filler. G. The ouicr molecular layer resembles closely the inner molecular layer, from which it differs only by containing branched stellate cells. 7. The outer nuclear Un/er is composed of several layers of nuclear cells, separable into two kinds, the rod granules and cone granules, both continuous with the rods and cones of Jacob's membrane. s. The membrana Umitans externa is, like the internal limiting membrane, derived from the radiating fibers, or fibers of Mailer. '.i. Jacob's membrane, or the layer of rods and cones, consists of two distinct kinds of cells — the rods and cones, distributed alternately throughout this layer, the rods being much more numerous. the rod- are solid, stand perpendicularly to the surface, and con- -i-t of two portions — an outer striated and an inner granular. The cone- are flask-shaped, with their pointed extremities toward the choroid. Thej a No consist of two portion- — an outer striated and an inner granular. ID. The pigmentary layer was formerly described as a layer of the choroid. It has received the name of tapetum nigrum, ('insisting of a layer of pigmented hexagonal epithelial cells. In main of tin' lower animal-, this layer on the posterior surface i- destitute <\v less in the lower lid. are arranged ver- tically on the inner surface of the cartilages; they are straight, seba- ceous follicles, into which open a number of secondary follicles, termi- nating above in a blunt extremity, and opening below on the free margin of the lids by small foramina, corresponding to the number of tubules; The eyelashes (cilia), are arranged on the free border of the lids ill two or three rows. The CONJUNCTIVA is the mucous lining membrane of the front of the eye. It consists of two portions — the bulbar. reflected over the sclerotic and cornea, and the palpebral por- tion; lining the internal surface of the lids. The bulbar portion {tunica conjunctiva bulbi) is loosely connected with the sclerotic, hut over the cornea if becomes very thin, consisting only of the epithelial layer. The palpebral portion (I unit a conjunctiva palpebrarum) is thick, highly vascular, and contains many papilla'. At the inner angle of the eye it forme a semilunar fold, the plica semi- lnnari- [plica semilunaris conjunctiva?) - the rudiment of the nictitating membrane of birds, the membrana niclilans. 74 HUMAN ANATOMY. The point of reflection is called the fornix conjunctivae, and the reflected portions the superior and inferior palpebral folds. The lachrymal apparatus (apparatus lacrimalis) consists of the lachrymal gland, canals-, sac and nasal duct. The lachrymal gland (glandula lacrimalis) is an oval, glandular body, about the shape and size of an almond, sit- uated in a depression in the upper surface of the orbit near the external angular process. The anterior portion of the gland is sometimes described ducts' of/ lachrymal, GLAND CARUNCLE I CANALICULUS LACHRYMAL SAC NASAL DUCT Fig. 135. The lachrymal apparatus. (Gerrish, after Testut; EcMey.) as a separate lobe — the palpebral portion of the gland, or accessory gland of Eosenmiiller (glandula lacrimalis inferior) ; the back part of the gland is designated the superior lachrymal gland (glandula lacrimalis superior). The gland is attached to the bony roof of the orbit by the tarso-orbital fascia. The ducts, seven to ten in number, open upon the con- junctiva near its point of reflection by minute orifices arranged in a row. The laclirymal canals commence at the puncta lacrimalia, at the summits of the lachrymal papilla (carunculus lacrimalis), and descend by two canaliculi to empty into the lachrymal sac. The superior canal (ductus lacrimalis superior) descends obliquely inward and downward, while the inferior (ductus ORGANS OF SPECIAL SENSE. 375 lacrimalis inferior) descends at first and then passes nearh horizontally inward. The lachrymal sac (saccus lacrimalis) is the oval, dilated, upper portion of the nasal duet, lodged in a deep groove formed by the nasal process of the superior maxilla and the lachrymal bone. It is crossed by the tensor tarsi muscle, which acts as a compressor, and receives a fibrous expansion from the tendo oculi. Its structure is made up of a fibrous elastic coat, lined bv mucous membrane continuous with the nose and conjunctiva. The nasal duct {ductus nasolacrimalis) is a membranous tube about three-quarters of an inch in length, extending from the lachrymal sac to the inferior meatus of the nose, and lining the bony lachrymonasal canal. It passes backward, downward, and outward, and is protected at its inferior extremity bv a valve of mucous membrane — the valve of Hasner (plica lacri- malis \ Hasneri]). The mucous lining is continuous with that of the sac, hut instead of having squamous epithelium it is ciliated in the duct, THE EAR. The ear (organon a ud it us) — the organ of hearing — is contained in the petrous and mastoid portions of the temporal hone, and consists of three divisions, — external ear, middle ear or tympanum and internal ear or labyrinth: — or meatus; 1. External ear, ( Auricle or pinna. I. External auditory canal f Membrana tympani, 2. Middle ear o'r tympanum, J Tyn.pai.ic cavity. ' ', .Mastoid sinuses. (_ Eustachian tube; {Vestibule, Semicircular canals, Cochlea ; ■>. internal ear or j labyrinth. \ r TT, . , [ Utricle, I Membranous portion. \ Saccule, 1 semicircular canals. I Cochlea. The external ear includes two parts, the auricle and the external auditory canal. 376 HUMAN ANATOMY. AURICLE (AURICULA), OR PINNA. The external prominent portion is composed of cartilagin- ous segments, connected together by ligaments and muscles richly supplied with blood vessels, nerves, and lymphatics, and covered with skin. It presents the following elevations and depressions : — Fossa of the helix, a depression between the helix and anti- helix ; Fossa of the antibelix (fossa triangularis [auricula]), a depression above the bifurcation of the antihelix; Tragus, the prominence in front of the concha, usually cov- ered with hair; Antitragus, a small, conical eminence behind the tragus, from which it is separated by the incisura intertragica ; Lobule (lobulus auricula), the soft, rounded portion de- pending below the antitragus; Concha, a deep cavity, surrounded by the helix, leading into the external meatus. Muscles. — The muscles of the auricle include two sets, the extrinsic muscles, already described (vide Muscles), of which there are three — the attolens aurem, attrahens aurem and re- trahens aurem — and the intrinsic muscles, but slightly devel- oped, of which there are two sets, four on the anterior surface of the auricle-^-the tragicus (m. tragicus), antitragicus (m. antitragicus) , helicis major (m. heHcis major), and helicis minor (m. helicis minor) — and two on the posterior surface, transversus auriculae (m. transversus auricula3) and obliquus auriculae (m. obliquus auricula'). Arteries, auricular branch, from the occipital; posterior auricular, from the external carotid; and anterior auricular, from the temporal. The- veins correspond to the arteries. Nerves, auricular branch, from the pneumogastric ; auric- ulotemporal branch, from the inferior maxillary; occipitalis major and occipitalis minor; auricularis magnus, from the cervical plexus; and posterior auricular, from the facial. Auditory caxal, or meatus auditorius externus (meatus acusticus externus), is an osseocartilaginous canal, about one and one-fourth inches in length, extending from the concha to the tympanic membrane. The osseous portion (pars ossea tuba auditivce, or semi- canalis tuba, auditivce) forms about two-thirds of the passage, and consists in greater part (anterior and lower) of a curved plate of bone — the annulus tympanicus. ORGANS OF SPECIAL SENSE. 377 The anterior and upper part of the ring presents two spines — spina tympanica, major and minor — for the Ligaments of the malleus. The sulcus tympanicus is a furrow on the inner edge of the ring, for the attachment of the tympanic membrane. The cartilaginous portion (pars cartUaginea tuba' audi- tiva ) forms about one-third of the passage, and consists of the inverted cartilage of the tragus and concha, the upper and back part of which is deficient, the cleft being filled with fibrous tissue. Fig. 136. rnal, middle, and internal ear. 1, external ear; 2, middle ear: •",, internal ear: 4. pinna; 5. helix; 0, antihelix; 7 fossa naviculars; s. fossa innominata; 9, tragus; 10, antitragus: 11, concha; 12, lobe; I .;. meatus auditorius externus; 14, tympanic membrane; 15, promon- tory: in, foramen rotundum; 17, posterior wall of the tympanum; is, OBSlcula auditus; 19, Eustachian tube; 20, narrow canal; 21, vestibule: 22. semicircular canals, the superior, posterior, and horizontal; 23, ampullae, 21. cochlea; 26, prominence caused by the scala vestibuli; 20, scala tympani. (Boenning.) The canal, including the tympanic membrane, is lined throughout with skin, containing short hairs, vascular papilla', sebaceous and ceruminous glands (glandvXai ceruminosw) . Arteries — branches of the internal maxillary, posterior auricular and temporal branches of external carotid. Nerves from the auriculotemporal branch, the inferior maxillary and auricular branch of the pneumogastric. 378 HUMAN ANATOMY. MIDDLE EAR (aURIS MEDIA), OR TYMPANUM, is an irregular cavity (cavum tympani) , situated between the auditory canal and the labyrinth, communicating with the pharynx through the Eustachian tube, and also with the mas- toid cells. It contains a chain of movable bones, part of the chorda tympani nerve, and is filled with air. Its average diam- eters are about half an inch in height and width, and a line or two in depth from without inward. It is lined with mucous membrane, continuous with that of the Eustachian tube and mastoid cells, and which is reflected over all the tympanic con- tents. It is bounded by a roof, floor and four walls, which present the following points : — ■ Eoof (paries tegmentalis) , a very thin plate of bone {teg- men tympani), separates the tympanum from the cranial cav- ity, and corresponds to a depression on the anterior wall of the petrous bone; Floor (paries jngularis), forms the bottom of the jugular fossa, and presents: — The opening for Jacobson's nerve, the tympanic branch of the glossopharyngeal ( n. tympanicus ) . Outer wall, formed by the tympanic membrane, Shrapnell's membrane, and the annulus tympanicus surrounding it: — Notch of Rivinus (incisure/, tympanica [Rivini]), incomplete upper ring of bone; Iter chordae posterius (canaliculus chordce tympani), gives en- trance to chorda tympani nerve; Iter chorda* anterius, or canal of Huguier, gives exit to same; Glaserian fissure (fissura petrotympanica [Glaseri]), above and in front of annulus tympanicus, receives the anterior ligament of the mal- leus, the long process of the malleus, and the tympanic artery; Pouches of the membrama tympani, are two or three pockets formed by the doubling of the mucous membrane around the chorda tympani nerve. Inner wall (paries labyrinthica) , separates the tympanum from the labyrinth : — Fenestra ovalis (fenestra vestibuli) , an oval window, leading to the vestibule, closed by the membrane to which the base of the stapes is attached; Fenestra rotunda (fenestra cochlea?), .a round window, leading to the scala tympani, but closed also by a membrane — membrana tympani secundaria. Promontory (promontorium) , two grooves,, marking the first turn of the cochlea, and separating the fenestras ovalis and rotunda; ORGANS OF SPECIAL SENSE. 379 Rounded eminence of the aqueductus Fallopii (prominentia canaiis facialis), a small conical eminence above the fenestra ovalis, which con- tains within it the aqueductus Fallopii, the long- canal for the facial nerve. Posterior wall (paries mastoidea) : — ■ Openings of tfo mastoid cells, three or four in number, connect these sinuses with the tympanum. Anterior wall (paries carotica), is a thin plate of bone, which separates the tympanum from the carotid canal. Openings of the canal for tensor tympani (semicanalis m. Icnsoris tympani), lies above and parallel with the canal for the Eustachian tube, and transmits the tensor tympani muscle; Opening of Eustachian tube (tuba auditiva [Eustachii ] ) , the larger of the two, is separated from the former by a thin plate of bone, the processus cochleariformis (septum canaiis musculotubarii) . The tympanum is divided into two parts — the attic (re- cessus epitympanicus) of the tympanum and the atrium. The former is situated on a plane directly above the atrium, and is bounded above by the tegmen, externally by auditory plate, and internally by prominence of the tympanum. On the outer part of the attic is a smooth surface called the scute (Leidy). The Eustachian tube (tubce auditiva \ Eustachii]) is an osseocartilaginous canal, one and one-half to two inches in length, extending from the tympanum to the pharynx. It is about one-third osseous and two-thirds cartilaginous, and lined with ciliated epithelium continuous with the pharynx and tympanum. Its pharyngeal orifice is trumpet-shaped, and opens at the side of the pharynx, behind the inferior meatus. Muscles of the Eustachian tube: — Tensor palati (m. tensor veil palatini), the most important has been described [vide Palatal Region). Its action is to pull tin' anterior wall of the tube outward and downward, and widen the canal. Internal Pterygoid {vide Muscles). — Some fibers are in- serted into the floor of the tube. It net- as tensor of the fascia. lAgamenta salpingopharyngeal, arises from the superior and middle constrictors nf the pharynx, and is inserted by three to five tendinous cords into the pharyngeal orifice of the tube. Action, opens the tube. Arteries of Eustachian Tube. — Pharyngeal from the exter- nal carotid, middle meningeal from the internal maxillari ami -mall branches from the internal carotid. 380 HUMAN ANATOMY. Nerves, by branches from the glossopharyngeal, inferior maxillary, otic ganglion and facial nerve. Membrana tympani is a thin, parchment-like membrane, stretched upon the annulus tympanicus, forming the bottom of the external auditory canal, and separating it from the tym- panum. It is placed obliquely downward and inward at an angle of 45°. Structure. — Its three layers are :— External or cuticular, derived from the skin of the meatus; Middle or fibrous, consisting of two sets of fibers, radiating and circular, the latter forming a tendinous ring around the margin ; Internal or mucous, continuous with the mucous membrane of the tympanum. Its outer surface presents : — Manubrium of malleus, projecting downward and back- ward ; Processus gracilis of malleus, projecting outward above the manubrium ; Umbo (umbo membrance iympance), a depression of the membrane formed by the lower end of the manubrium; Yellow spot, the cartilaginous end of the manubrium; pyramid of light, a triangular cone of reflected light in the antero-inferior quadrant of the membrane. Inner surface (vide Outer Wall of Tympanum). Arteries, tympanic branch of inferior, branch from the internal carotid. Nerves, from the superficial temporal branch of the tri- facial and from the tympanic plexus. OSSICLES OF THE TYMPANUM (OSSICULA AUDITUS). The small bones of the tympanum are three in number, and connect the membrana tympani with the membrane closing the fenestra ovalis. They are the 1. Malleus, or hammer, consists of a head (capitulum mallei), neck (collum mallei), manubrium or handle, processus gracilis (processus anterior \Foli%\), and processus brevis (pro- cessus lateralis). The manubrium (manubrium mallei) is at- tached to the membrana tympani, and has the tendon of the tensor tympani attached. Processus gracilis is lodged in the Glaserian fissure. 2. Incus, or anvil, consists of a body (corpus incudis) and a long (eras longum) and short processes (crus breve). The ORGANS OF SPECIAL SENSE. 381 bodv articulates with the malleus and the long process by means of a rounded process. The os orbiculare (processus lenticularis) articulates with the head of the stapes. 3. Stapes, or stirrup, has a head (capitvlwm stapedis), neck, two branches or crura (crura anterius et crura poste- riiis). which unite into a cross-piece or bast' (basis stapedis) The neck has inserted into it the stapedius muscle, and the base is fitted to the fenestra ovalis. Fig. 137. The auditory ossicles. A. G., external meatus; M, membrana tym- pani ; n, handle of the malleus and its short process (p); h, head of the malleus; a, incus; K, its short process, with its ligament; I, long process; *, os orbiculare; »S', stapes; -lJ", Ax, the axis of rotation of the ossicles, shown in perspective; t, line of traction of the tensor tympani. The other arrows show the movements of the ossicles when the tensor contracts. (Boennhnj.) Ligaments of the ossicles (ligamenta ossiculorum audi- tus) : — Anterior ligament of the malleus (ligamentum mallei anterius), extends from the neck of the malleus to the outer wall of the tympanum; Superior Ligament of the malleus — ligamentum mallei sus|icn- sorium [ligamentum mallei superius), suspends the head of the malleus to tin- roof; Posterior ligament of the incus (ligamentum incudis posterius), attaches the shorl process to the posterior wall; Superior ligament of the incus (ligamentum incudis superius), ]- merely ■■> told of mucous membrane; while tin- inner surface ami cir- 382 * HUMAN ANATOMY. cuniference of the base of the stapes is connected to the margin of the oval window by means of the annular ligament of the stapes (ligamen- tum annulare baseos stapedis) . Muscles. — The tensor tympani and stapedius. Tensor t}rmpani (already described). Stapedius (m. stapedius) arises from the hollow of the pyramid on the inner wall, and its tendon is inserted into the neck of the stapes. It is the smallest muscle in the human body. Nerve from the tympanic branch of facial. Arteries: — Tympanic branch of internal maxillary; Stylomastoid branch of posterior auricular; Petrosal branch of middle meningeal; Tympanic branch from internal carotid; Branch from ascending pharyngeal. Veins accompany the corresponding arteries and empty into middle meningeal and pharyngeal. Nerves: — Chorda tympani, from the facial, enters the iter chorda? posterius, crosses the tympanum between the long process of the incus and the handle of malleus, and makes its exit at the iter chordae anterius; Tympanic branch of glossopharyngeal (Jacobson's nerve) enters the floor, supplies the fenestras and mucous membrane of tympanum and Eustachian tube ; Tympanic branch from facial to the stapedius; Branch from the otic ganglion to the tensor tympani. The tympanic plexus is formed upon the surface of the promontory, from the following nerves: — ■ Jacobson's nerve, tympanic branch of the glossopharyngeal; Branch of the superficial petrosal, from the facial; Branches from the carotid plexus of the sympathetic. INTERNAL EAR, OR LABYRINTH. The internal ear (auris interna), the essential part of the organ of hearing, consists of three complex cavities within the petrous portion of the temporal bone, rilled with fluid, the perilymph, and containing a membranous sac filled also with fluid, the endolymph, which receives the distributions of the auditory nerve. ORGANS OF SPECIAL SENSE. 383 Its divisions are : — (Vestibule, Semicircular canals, Cochlea ; f Utricle, , ,. i Saccule, Membranous portion, [ Cochlea. Vestibule (vestibulum) , is a common cavity of communi- cation between the bony parts of the internal ear. It is sit- uated between the tympanum without, the cochlea behind, and tli. semicircular canals in front, and measures one-fifth of an inch in its anteroposterior and vertical diameters, and less from without inward. It presents: — Fenestra ovalis (fenestra vestibuH) on its outer wall, closed by the stapes and its ligament; Fovea hemispherica (recessus spharicus) , a circular de- pression on its inner wall for the saccule, and perforated by the macula cribrosa (macula cribrosa media), for the vestibular filaments of the auditory nerve; Orifice of the aqucechictus vestibular, on the inner wall for the transmission of a small vein; Eminent ia pyramidalis, a vertical ridge on the inner wall separating the two fovea?. Fovea semielliptica (recessus ellipticus), in the roof lodges the utricle. SemicircuIiAB canals (canales semicirculares ossei) are three canals, forming each two-thirds of a circle, one-twentieth of an inch in diameter, and named, from their position, the superior, posterior and external. They are placed nearly at right angles to each other and open into the vestibule by live apertures — two extremities uniting to form one. The superior (canalis semicircularis superior) and poste- rior {canalis semicircularis posterior) arc both vertical, the former being more anterior. The externa] {canalis semicircularis lateralis) is placed horizontally, its arch directed backward. Ampulla, arc the dilated, flask-shaped extremities of the tubes, and aro about one-tenth of an inch in diameter. Tin: cochlea resembles closely a common snail's shell, placed with the base corresponding to the bottom of the meatus auditoriue internus, and its apes directed outward and forward. It consists of two parallel tubes one and one-half incbes in 384 HUMAN ANATOMY. length, one-tenth of an inch in. diameter, wound spirally for two and one-half turns around a central pillar — the modiolus. The modiolus, or columella (basis modioli), is broad at the base, but tapers gradually toward the apex, where it terminates within the last turn or cupola in the infundibulum, a funnel- shaped lamina of bone. Its interior is traversed by numerous canals for vessels and nerves, one of which — the canalis centralis modioli — runs nearly the whole length and transmits the arteria centralis modioli. ■ FIG. 138. Organ of Corti. AT, cochlear nerve; K, inner and, P, outer hair- cells; n, nerve fibrils terminating in P; a, a, supporting cells-; d, cells in sulcus spiralis; z, inner rod of Corti; Mb, Corti, membrane of Corti, or the memibrana tectoria; o, the membrana reticularis; H, O, cells filling up the space near the outer wall. (Bocnning.) Lamina spiralis, the partition between the two tubes, con- sists of two thin laminas of bone filled with cancellous tissue — lamina spiralis ossea — projecting from the modiolus half-way to the outer wall, the lamina being completed by the membrana basilaris. The osseous lamina terminates in the cupola in a hook-like process — the hamulus (hamulus lamince spiralis). The spiral canal, between the modiolus and the outer wall, is divided by the lamina spiralis ossea and membrana basilaris into two canals, or scalce — the upper scala vestibuli, and the lower scala tympani. ORGANS oF SPECIAL SENSE. 335 The first turn of the spiral canal presents three openings : — 1. Fenestra rotunda, a circular aperture communicating with the tympanum, and closed by the membrana tympanum secundaria. '2. Fenestra oralis, an oval opening, communicating with the vestibule. :'.. Aquaeductus cochlearis (apertura externa canaticuli cochlea), a Bmall foramen opening on the inferior surface of the petrous portion of the temporal bone, and transmitting a small vein from the cochlea to the jugular vein. 8cala vestibuli, begins at the fenestra ovalis, and ascends in a spiral course above the lamina spiralis and membrana basilaris, to communicate with the scala tympani, through the helicotrema within the vestibule. It contains perilymph. Scala tympani begins at the fenestra rotunda, and ascends, in a spiral course below the scala vestibuli, from which it is separated by the osseomembranous lamina. It also contains perilymph. 11 1'licotrema is an opening within the cupola, by which the two scalae communicate. It is formed by a deficiency of the osseous lamina. Canalis spiralis modioli is a small canal which winds around the modiolus at the attachment of the osseous laminae. Jt contains a gangliform swelling — ganglion spirale (ganglion spiral e cochlea) — from which nerves pass to the organ of Corti. The membranous labyrinth (labyrinthus membrana- n us) corresponds closely to the osseous, the vestibule consisting of two membranous sacs — the utricle and saccule — communicat- ing with each other, and the former receiving the five openings of the membranous semicircular canals, the latter communicat- ing with the membranous cochlea through the canalis reuniens (il ml us reuniens \IIenseni\). The membranous labyrinth is Burrounded everywhere with perilymph (perilympha) } filled with endolymph (endolympha | . and is composed of three coats — an outer fibrous layer, middle or tunica propria and inner epithe- lial Iay the utricle and ampullae of the superior find external semicircular canals; 388 HUMAN ANATOMY. Middle, enters the bottom of the fovea hemispherica, and is dis- tributed to the saccule; and the Inferior is distributed to the ampulla of the posterior simicircular canal. Cochlear nerve (n. cochlearis) , ascends in the canals in the modiolus, and divides into numerous branches which pass be- tween the plates of the lamina spiralis ossea, forming a plexus which contains the ganglia spirale (ganglion spirale), from which filaments are distributed to the outer and inner hair cells of the organ of Corti. Arteries, of Labyrinth: — Internal auditory, from the basilar, divides into vestibular and cochlear branches., which accompany the corresponding nerves ; Stylomastoid, from the posterior auricular, enters through the stylomastoid foramen. THE TONGUE.1 The tongue (lingua) is the organ of the sense of taste, and lies in the floor of the mouth. It is composed of a mass of muscles, the free surface of which is entirely invested with mucous membrane. It consists of three parts: — Base, or root (radix Ungues), attached to the hyoid bone, the epiglottis and the soft palate; Body (corpus linguae), the back or dorsum of which is marked by a median line or raphe, terminating behind in a cavity— the foramen cecum (foramen caecum lingua? [Mor- gagnii]). Apex (apex lingua?), or tip, narrow and pointed, directed forward. The mucous membrane (tunica mucosa Ungual) resembles skin, consisting of a corinm or mucosa, containing numerous papilla?, and covered with epithelium. The papilla? (papilla? lingua?) are most numerous over the anterior two-thirds of the dorsum, and consist of three varieties : — Circumvallate or papilla? maxima? (papilla? vallata?) , the largest, about ten in number, about one-twelfth inch wide, are arranged at the back part of the dorsum like the letter V with the apex directed backward; Fungiform or papilla? media? (papilla? fungiformes et papilla? lenticulares) , the medium, are scattered over the organ, especially at the tips and sides; 1 Vide Lingual Muscles, page 135. ORGANS OF SPECIAL SENSE. 389 Filiform, conical or papillae minima3 (papUlce conicce et papilla fUiformes), the smallest, are distributed over the ante- rior two-thirds of the dorsum. Mucous and serous glands and simple papilla?, such as are found in the skin, are also present. Taste buds, minute, flask-shaped bodies, about one-three- hundredths of an inch in length, are situated in the circumval- late and fungiform papilla. Arteries are branches of the lingual, facial and ascending pharyngeal. 1. Nerves. — Hypoglossal, the principal motor nerve. 2. Lingual branch of glossopharyngeal, the special nerve of taste. 3. Lingual branch of the trifacial, to sides and anterior part; nerve of common sensation. 4. Chorda tympani, probably nerve of special sense. 5. Superior laryngeal, filaments from its internal branch to the base. THE SKIN AND APPENDAGES. The skin (integumentum commune), besides being the spe- cial organ of the sense of touch, is an absorbing and excretory organ, and protects the underlying structures. It consists of two layers — the epidermis or cuticle, and the derma or true skin. The appendages are the nails, hair, sudorif- erous and sebaceous glands, and their ducts. The epidermis, or cuticle, is an epithelial structure con- sisting of four layers : — 1. Bete MalpigTm, the deepest layer, in contact, with the corium, is composed <>f several layers of round or polyhedral epithelial cells. This layer is pigmented in the negro and other dark races. 2. Granular layer, is a single layer of compressed, spindle-shaped cells. '■',. St rut am I uciilii m . is -till mure (-(impressed into scales. 4. Stratum corneum, several layers of horny scales. The derma, corium, or true skin, has two Layers, the reticular or deeper, and the papillary or superficial. Reticular layer {stratum reticulare) is composed of inter- lacing bands of white fibrous tissue, mixed with yellow elastic libers, and some iinstrialed muscular fibers where hairs are found, together with blood vessels, lymphatics and nerves. Papillary layer (stratum papUlare) is covered with minute, vascular, highly sensitive elevations- the papillae — from one- one-hundredth to one-two-hundred-and-fiftieth of an inch in 390 HUMAN ANATOMY. diameter. These papillae are in places arranged into parallel curved ridges, and have in the most sensitive situations some one of the peripheral end organs (already described) added to their structure. Nails (ungues) are horny plaques of modified epidermis, firmly adapted to the derma on the dorsal surfaces of the termiual phalanges. Each nail is firmly implanted by its root (radix unguis) into a fold of skin. The matrix (matrix unguis) is the derma beneath the body, which is vascular and highly sensitive. The lunula,, on account of its crescentic shape, is the white, less vascular portion of the matrix near the root. Haies (pili) are a cylindrical' modification of the epider- mis, and consist of a root imbedded in the skin, a shaft and a point. Eoot (radix pili) is lodged in a pouch-like involution of epidermis — the hair follicle (follicidus pili). This is composed of an inner or cuticular and an outer or dermic layer, and pre- sents at the bottom a vascular papilla, on which the hair rests, and from which it grows. Shaft (scapus pili) consists of a pith or medulla (sub- stantia medullaris pili) in the center, surrounded by a fibrous part containing pigment, and covered with a layer of scaly epithelium. Point contains the two latter elements, but has no medulla. Sebaceous glands (glandulce sebacece) are minute, gland- ular bodies, situated in the substance of the corium, and each opening by a single duct into a hair follicle, or upon the sur- face of the skin. They are most abundant where hairs are found. The Meibomian glands in the eyelids are the largest. Sudoriferous or sweat glands (glandulce sudoriferce) are minute, reddish, glandular bodies, situated usually in the sub- cutaneous connective tissue, each consisting of a single con- voluted tube. Each gland opens on the surface of the skin. SURGICAL ANATOMY. THE TRIANGLES OF THE NECK. The general outline of the neck is somewhat quadrilateral. Bounded in front by the median line, behind by the trapezius muscle, above by the lower border of the body of the maxilla and below by the clavicle. It is divided by the sternomastoid muscle running obliquely through it into two large triangles, an anterior and a posterior triangle. These are again sub- divided by the crossing of the omohyoid muscle into two smaller triangles each. The anterior into the superior and interior carotid triangles, and the posterior into the occipital and sub- clavian triangles. The digastric nmscle, at the upper part ot the anterior triangle, separates a triangular space called the submaxillary triangle. n„«™. The (interior triangle of the neck is bounded as follows.— Front, median line of the neck ; Behind, anterior border of the sternomastoid muscle ; Vl.uve. lower border of the body of the lower jaw and a line from its angle to the mastoid process, forming the base ot the triangle; . Below, the top of the sternum, forming the apex. The floor is formed by the sternohyoideus, sternothy- riodeus, thvrohvoideus, inferior and middle constrictors of the pharynx, the anterior belly of the digastricus, the mylohyoideus, stylonyoideus, and hyoglossus muscles. The roof is formed by the skin, superficial fascia, platysma myoides, and deep iascia. The inferior carotid triangle is the most inferior subdivi- sion of the anterior triangle, and is bounded as follows:— Front, median line of the neck; Behind, anterior border of the sternomastoideus ; Above, anterior belly of the omohyoideus; Below, the apex of "the anterior triangle. It contains the following structures:— Thyroid gland and lower part of the trachea and Larynx; Inferior thyroid and common carotid arteries; Inferior thyroid and internal jugular reins; Pneumogastric, recurrenl laryngeal, descendens noni, com- municans noni and sympathetic nerves. (391) 392 HUMAN ANATOMY. The superior carotid triangle is the middle subdivision of the anterior triangle of the neck, and has the following bound- aries : — Above, posterior belly of the digastricus; Below, anterior belly of the omohyoideus; Behind, anterior border of the sternomastoideus. It contains the following structures : — Termination of the common carotid; The internal carotid; The external carotid, and the following branches: — Superior thyroid, lingual, facial, ascending pharyngeal and occipital; The internal jugular vein, and the following tributaries : — Lingual, facial, superior thyroid, pharyngeal and occa- sionally the occipital; The pneumogastric, superior laryngeal, external laryngeal, descendens noni, spinal accessory, hypoglossal and sympathetic nerves. The submaxillary triangle is the most superior of the sub- divisions of the anterior triangle, and has the following bound- aries : — Above, the lower border of the body of the jaw and the imaginary line behind ; In front, the median line of the neck ; Behind, the posterior belly of the digastricus. It contains the following structures : — External carotid, internal carotid, facial, submental, mylo- hyoidean arteries; The internal jugular, the commencement of the external jugular, branches of the anterior jugular and the facial vein and its branches; The inframaxillary branches of the facial nerve, the ascend- iug branches of the superficial cervical nerve, and the pneumo- gastric and glossopharyngeal and mylohyoid nerves; Portions of the parotid and submaxillary glands, and also submaxillary lymphatic glands and vessels. The posterior triangle of the neck occupies the space be- hind the posterior border" of the sternomastoideus, and is bounded as follows : — Front, posterior border of the sternomastoideus; Behind, anterior border of the trapezius; Below, upper border of the middle third of the clavicle, forming its base; Above, the occiput, forming the apex. SURGICAL ANATOMY. 393 Its floor is formed from above downward, by the following muscles : — Splenius capitis, levator anguli scapuli, scalenus medius, scalenus posticus, and the upper portion of the serratus magnus. Its roof is formed by the superficial and deep fascia and below by the platysma myoides. FIG. 139. Triangles of the neck. (Bocnning.) It contains the following structures: — 'Jdic transversalis colli artery and vein; Spinal accessory and superficial plexus of the cervical nerve ; Lymphatic glands and vessels. The subclavian triangle, or the inferior division of the posterior triangle, is bounded — 394 HUMAN ANATOMY. In front by the margin of the sternomastoideus, behind by the posterior belly of the omohyoideus, below by the clavicle. It contains the following structures : — The subclavian, transversalis colli and transversalis humeri or suprascapular arteries; The transversalis colli, suprascapular and external jugular veins ; The descending branches of the superficial cervical plexus and the brachial plexus of nerves ; Lymphatic nerves and vessels. Triangle in Front of the Elhoiu-joint. Bounded — Externally, by supinator longus; Internally, by pronator radii teres; Base, above, by a line (imaginary) drawn through the con- dyles; Apex, below, by crossing of supinator longus and pronator radii teres. It is covered in by skin, superficial fascia and bicipital fascia; the floor is formed by oblique fibers of the supinator brevis and lower part of brachialis anticus muscles. It contains: — From within outward, median nerve, brachial artery, vense comites, biceps tendon and musculospiral nerve. Scarpa's Triangle (trigonum femorale) is a large triangular space situated in the upper part of the anterior surface of the thigh, through which the femoral vessels descend. It is bounded — Externally by sartorius, internally by adductor longus, above by Poupart's ligament; below, apex is formed by crossing of bounding muscles. It is covered in by skin, superficial fascia, cribriform fas-- cia and fascia lata, and its floor is formed by the following from without inward : iliacus, psoas, pectineus and portion of adduc- tor brevis muscles. It contains: — ■ The femoral artery (with its profunda and cutaneous branches), inclosed in the femoral sheath, femoral vein (joined by profunda and long saphenous veins), anterior crural nerve and its branches [from within outward being vein, artery and nerve], deep lymphatic glands and vessels and adipose tissue. SI RGICAL ANATOMY. 395 Axilla. This is a pyramidal space between the upper and lateral part of the chest and the inner side of the arm. It is bounded — In front, by the pectoralis major and minor muscles; be- FIG. 140. The axilla. 1, axillary artery; 2, brachial artery; 3, acromio- thoracic artery; 4, superior thoracic artery; 5, subscapular artery; 6, dorsalis scapulae artery; 7, posterior circumflex atery; 8, superior pro- funda artery; 9, posterior thoracic nerve; 10, long subscapular nerve; 11, median nerve; 12, cephalic vein; 13, musculocutaneous nerve; 14, teres major muscle. {Boenning.) hind, by latissimus dorsi, teres major und Bubscapularis; inter- nally, by the upper four ribe and intercostal and upper pari of serratlM magnilfl muscles, and ils apex i- directed upward, and 396 HUMAN ANATOMY. is formed by the space between the first rib, clavicle and upper border of the scapula, and its base by the skin and axillary fascia stretched across between the lower borders of the pec- toralis major and latissimus dorsi muscles. It contains: — The axillary artery and vein and their branches; the brachial plexus of nerves and the branches given off below the clavicle; branches of the intercostal nerves; ten or twelve lym- phatic glands and a quantity of loose adipose and areolar tissue. Popliteal Space. The popliteal space is a lozenge-shaped space situated at the back of the knee, and forms the ham. It is bounded — Externally, above the joint, by the biceps; below the joint, by outer head of gastrocnemius and plantaris; internally, above the joint, semitendinosus, semimembranosus, gracilis and sar- torius; below the joint, by inner head of the gastrocnemius. It is covered in by the skin, superficial fascia and fascia lata. Its floor is formed* by the lower part of the posterior sur- face of the femur, the ligamentum posticum Winslowi and the popliteus muscle and its fascia. It contains: — ■ The popliteal artery and branches; popliteal vein receiving the external saphenous vein (vein to the outer side) ; internal and external popliteal nerves and branches; articular branch of obturator nerve ; branch of small sciatic nerve ; four or six lymphatic glands, and a quantity of adipose and areolar tissue. ANATOMY OP HERNIA. Hernia is the protrusion of any part of a viscus from its natural cavity through the inclosing walls of the cavity. As usually understood, it refers to the protrusion of the intestine or mesentery, or both, from the abdominal cavity. The most common forms are : — External or oblique inguinal hernia; Internal or direct inguinal hernia; Femoral hernia; Umbilical hernia. 1. Oblique inguinal hernia escapes from abdominal cav- ity at the internal abdominal ring, carrying before it a pouch si RGICAL ANATOMY ;'.»; of peritoneum, descends along the inguinal canal in front of the cord, and emerges at the external opening. The anatomical parts concerned are the inguinal canal, with its interna] and external abdominal rings, the transversals fascia (vide Fascia), the peritoneum (vide Peritoneum) and Poupart's ligament. Fig. 141. Dissection of the inguinal canal: A, external oblique; B, li, in- ternal oblique; 0, transversalis; D, conjoined tendon; E, rectus ab- dominis with sheath opened; G, fascia transversalis; //, cremaster; /, infundibular fascia. The mguvnal or spermatic canal is about one and a half inches in length, extending from the internal abdominal ring to the externa] abdominal ring. It serves for the passage of the spermatic cord in the male, and the round ligament with essele in the female. Us boundaries are: — 398 HUMAN ANATOMY. In front, skin, superficial fascia, the aponeurosis of the external oblique muscle, the lower border of the internal oblique, and a small portion of the cremaster muscle; Behind, the fascia transversalis, the conjoined tendon of the transversalis and internal oblique muscles, the transversalis fascia, and subperitoneal fat and peritoneum; Above, by the arched border of the internal oblique and transversalis muscles; r~ Below, by the connection between Poupart's ligament and the transversalis fascia. The internal abdominal ring (annulus ingirinalis abdomi- nalis) is an oval opening in the transversalis fascia about half an inch above Poupart's ligament, midway between the sym- physis pubis and the anterior superior spinous process of the ilium. It is bounded — • Above and externally by the arched fibers of the transver- salis muscle ; below and internally by the deep epigastric vessels. It transmits a funnel-shaped fascia from its margins, the in fun dib uliform fas cia. The external abdominal ring (annulus inguinalis subcu- taneus) is a triangular opening between the two pillars in the aponeurosis of the external oblique muscle, to the outer side and just above the spine of the pubes. It is about half an inch wide and one inch long, and is bounded — Above, by the intercolumnar fascia; below, by the spine and crest of the os pubis, and on either side by the pillars or columns of the ring formed by the free margins of the aponeu- rosis of the external oblique. From the margins of the external abdominal ring arises the intercolumnar fascia. Poupart's ligament, or the femoral arch (ligamentum in- guinale), is the rounded lower fibrous margin of the external oblique muscle, extending between the anterior superior spine of the ilium and the spine of the pubes. The portion of the aponeurosis which is inserted into the pectineal line has received the name of Gimbernat's ligament (ligamentum lacunare). The triangular ligament or ligament of Colles (ligamen- tum inguinal reflexum) is the reflected inner portions of Gim- bernat's and Poupart's ligaments attached to the sheath of the rectus. The epigastric artery (a. epigastrica inferior) holds a very important anatomical relation to the inguinal canal and inter- nal abdominal ring. It ascends between the peritoneum and transversalis fascia to reach the sheath of the rectus muscle M RGICAL ANATOMY. 39U Fig. 142. Deep Femoral Region-the Femoral Vessels, etc., cut across as they emerge under Pouparfs ligament. AC, anttrior crural nerve; ( 7 .dge of the conjoined tendon; CK, crural ring; K, dotted line in- dicates the course of the deep epigastric artery; FB, femoral sheath; Q Gimbernafa ligament; TP, llio-pectineal ligament; P, Pouparfs ligament; 'PE, pectineus muscle. This muscle rests upon the pubic bone and is covered by its fascia— the pectineal fascia— which is some- what thickened Immediately beneath Pouparfs ligament, where it is known as the pubic ligament of Cooper. It will be noticed that the femoral sheath is divided into three compartments: the outer for the femoral artery; the middle for the femoral vein; the inner (OR) is1 ti,. . rural ring, the mouth of the crural canal. (McQrath.) 400 HUMAN ANATOMY. along the inner and lower margin of the internal ring and beneath the spermatic cord. The coverings of oblique inguinal hernia are : — (1) Skin; (2) superficial fascia ; (3) intercolumnar fascia; (4) cremaster muscle and fascia; (5) fascia transversalis, or infundibuliform fascia; (6) subserous connective tissue; (7) peritoneal sac. - 2. Direct inguinal hernia escapes through the abdom- inal wall, within Hesselbach's triangle, internal to the epigas- tric artery, and through the external abdominal ring, pushing before it a pouch of peritoneum. Hesselbach's triangle is a space situated at the lower ante- rior part of the abdominal wall on either side. It is bounded as follows: — Externally, epigastric artery; Internally, the outer margin of the rectus abdominis muscle ; Below, Poupart's ligament, forming its base. The struc- tures forming the abdominal wall at this cavity are, from with- out inward: — 1. Skin; 2. Superficial fascia; 3. Intercolumnar fascia; 4. Conjoined tendon of the transversalis and internal ob- lique ; 5. Fascia transversalis; 6. Subserous tissue; 7. Peritoneum. The anatomical parts concerned in direct inguinal hernia are the same as those in the indirect, already given. The cov- erings of direct inguinal hernia are those just enumerated. 3. Femoral, hernia escapes through the femoral ring, carrying before it a pouch of peritoneum, descends through the femoral canal and emerges through the saphenous opening in the fascia lata. The anatomical parts concerned in this hernia are: — The femoral ring, the femoral or crural canal, the septum crurale, crural sheath, deep crural arch and the saphenous opening. The femoral, or crural ring (annuhis femomlis) , is an oval opening, about an inch and a half in diameter, larger in the female and situated below Pou'part's ligament, between the inner side of the femoral vein and the margin of Gimbernat's ligament. It is the abdominal opening of the femoral canal SURGICAL ANATOMY. 401 (canalis femoralis), and it is closed in the recent state by the septum crurale and a small lymphatic gland. It is bounded — In front, by the deep femoral arch and Pouparf s ligament ; Behind, by the pubes, covered by the pubic portion of the fascia lata and the pectineus muscle ; Externally, by the femoral vein, from which it is sep- arated by a fibrous septum ; Internally, by the deep femoral arch, Gimbernat's ligament, the transversalis fascia and the conjoined tendon. The femoral canal (canalis femoralis) is a space from a quarter to half an inch in length, extending from Gimbernat's ligament to the upper margin of the saphenous opening. Bounded above by the femoral ring and below by the saphenous opening. It is the innermost compartment of the femoral sheath. Anterior wall — transversalis fascia, separated by the falci- form process (margo falciformis) of the fascia lata; Posterior wall — iliac fascia and pubic portion of the fascia lata; External wall — fibrous septum of the femoral vein; Internal wall — Gimbernat's ligament, deep crural arch and the junction of the iliac and tranversalis fascia. The septum crurale (septum femorale musculus) is a layer of dense areolar tissue, supporting small lymphatic glands and closing in the femoral ring. It is perforated by numerous lym- phatic vessels, and forms a barrier to the escape of hernia at this point. The crural or femoral sheath (fascia cruris) is a prolonga- tion downward of the fascia lining the abdomen (transversalis fascia in front, iliac fascia behind the vessels), closely adhering to the femoral vessels to about an inch below the saphenous opening (fossa ovalis). Its upper part is funnel-shaped and it- Lower part continuous with the sheath of the vessel. It is divided by septa into three compartments; in the outer is lodged the femoral artery; the middle is occupied by the femoral vein; and the innermost, the femoral canal, is empty, or occupied by a Lymphatic gland. Its outer border is pierced by the genito- c rural nerve, its inner by the internal saphenous vein. The deep femoral arch is a fibrous thickening of the trans- versalis fascia which forms the anterior Avail of the femoral gheath. It is* also known as the deep crural arch. The saphenous apenmg is an oval aperture of half an inch in width, an inch and a half in length in the upper and inner part of the fascia lata, between its two divisions, the iliac and 26 402 HUMAN ANATOMY. pubic portions. It is bounded externally by the falciform process of Burns, internally by the pubic portion of the fascia lata, which curves upward behind the saphenous vein. The opening is covered externally by the cribriform fascia (fascia cribrosa) and the skin. | The coverings of femoral hernia are: — , 1. Skin; 2. Superficial fascia; 3. Cribriform fascia ; Hnttrhrlayrref V,., r™j *"«■♦- ~» Fig. 143. Male perineum, superficial dissection. (Lydston.) 4. Femoral sheath, crural sheath, or fascia propria; 5. Septum crurale; 6. Subserous connective tissue; 7. Peritoneal sac. ISCHIORECTAL REGION AND PERINEUM. The outlet of the pelvis is a lozenge-shaped space divided by an imaginary line drawn in front of the anus, transversely between the anterior margin of the tuber ischii/into two parts, the ischiorectal region behind and the perineum in front. The ischiorectal region contains the external (m. sphincter ani externus) and internal sphincters (m. sphincter ani inter* SURGICAL ANATOMY. 403 mis), the corrugator cutis ani and the ischiorectal fossa (fossa ischwrectalis) . The latter is a pyramidal cavity, its apex reaching to the junction of the obturator and anal fascia and its l>ase formed by the skin. It is bounded—. Behind, by the edge of the great sacrosciatic ligament and gluteus maximus muscle; internally, by the levator ani, covered by the anal fascia, sphincter ani and coccygeus muscle; exter- nally, by the tuber ischii and obturator fascia. Artfrj of Cerpat Cavtnnmn 0*r*at Artorg j/* Pent* Aritry f/ Buli. Zmttmal PaJie Arte C*rp*r* dam FIG. 144. Male perineum, deep dissection. (Lytlston.) It t O/i hi ins; The internal pudic artery, nerve and veins inclosed in the fa-rial canal or canal of Alcock ; The inferior hemorrhoidal vessels and nerves; The perineal and fourth sacral nerves; and A mass of areolar tissue. PEEINEUM. The perineum is the triangular space between the amis and Bcrotum in the male, and between the aims and vulva in the female. The skin is dark, thin, and supplied with sebaceous and sudoriferous glands and is marked by middle Line or raphe. 404 HUMAN ANATOMY. Fascia of the Perineum. The pelvic fascia {fascia pelvis) at the brim of the pelvis is continuous with the transverse and iliac fascia; covers sac- rum, pyriformis muscle, external iliac artery and sacral nerves. It forms the lateral true ligaments of the bladder {ligamenta puboprostatica lateralia) in the male, and {ligamenta pubo- vesicalia lateralia) in the female. It also forms the anterior true ligament of the bladder {ligamentum puboprostaticum medium) in the male {ligamentum pubovesicale medium,) in the female. It forms arcus tendinege {arcus tendineus fascice pelvis), or white line, corresponding to division of pelvic fascia into .two layers, the obturator and rectovesical {fascia diaphragmatis pel- vis superior), extending from pubis to spine of ischium, from which originates the levator ani muscle, with rectovesical and ischiorectal or anal fascia? above and below the muscle re- spectively. The obturator fascia {fascia obturatoria) , the continuation of the pelvic fascia, incloses the pudic vessels and nerve in a sheath, covers the internal obturator muscle and forms the pos- terior layer of the triangular ligament. Bectovesical or visceral layer of the pelvic fascia, the com tinuation of the pelvic fascia, descends into the pelvis investing the prostate glaud and the vesicoprostatic plexus of veins, the bladder (forming the anterior and lateral true ligaments) and the rectum. It is perforated in the female by the vagina. Superficial perineal fascia {fascia superficialis perinei) consists of superficial fat layer and deep membranous layer, the fascia of Colles. The former is continuous with subcutaneous layer of buttocks, thigh, and labia, and posterior to anus be- come continuous with tissue of ischiorectal fossa. The superior laijer passes from rami of pubis and ischium to the tuberosity of ischia, covering ischiocavernous and bulbourethral muscles, and becomes continuous with the deep perineal fascia. The deep perineal fascia (triangular ligament of the per- ineum— trigonum urogenitale or diaphragma urogenitale) is a strong triangular membrane extending between the rami ischii et pubis, its apex attached to the under surface of the sym- physis. It is often described as consisting of two layers, the superior, deep or posterior layer {fascia trigoni urogenitalis superior), and inferior, anterior or superficial layer {fascia trigoni urogenitalis inferior) . It is pierced by the membranous urethra, and sends a fascia surrounding the urethral glands to the spongy body. In the female it is weaker, the urethra also pierces it, and it is continuous with the fascia of the vagina. si i;<;i< Al. ANATOMY. 405 Muscles of the Perineum. Ischiocaveknosl's (erector penis) arises in the inner sur- face of the tuberosity of the ischium, and is inserted into the side and under surface of the cms penis. In the female there is a similar insertion into the clitoris. It serves to maintain the organ erect. Nerve, perineal. Erector clitoridis (m. ischiocarernosus) corresponds to erector penis muscle in the male, but smaller. Female perineum: .4, anus; IS, bulbo-vaginal; C, coccyx; G, glu- teus maximus; P, perineal body; U, urethra; V, vagina; G, vulvo- vaginal gland; 1, clitoris; 2, its suspensory ligament; 3, crura clito- ridis; 4, erector clitoridis; 5, bulbo-cavernosus; 7, transversus perinaei; 8, sphincter ani; 9 and 10, levator ani; 11, coccygeus; 12, obturator externus. Accelerator ubism or ejaculatob beminis (m. bidbo- cavernosus) arises from central tendon of perineum and from mediaii raphe in front. It spreads out and is inserted from behind forward to anterior surface of triangular ligament, bull) and adjacent part of corpus spongiosum, to join fibers of opposite side, and to sides of corpora cavernosa ; anterior to erector penis and in fibrOUS expansion over dorsal Mood vessels of penis. It accelerates the How of urine and semen and contributes to erection of the penis. Nerve, perineal. 406 HUMAN ANATOMY. Vaginal constrictor or sphincter vagina (to. bulho- cavemosus) , analogous to foregoing muscle, surrounds the orifice of vagina. It arises from the centraL^endon and passes forward on either side of vagina to be inserted into the corpora caver- nosa and body of clitoris. It compresses the dorsal vein of the clitoris. Transversus perin^ei (superficialis) arises from inner part of ramus of ischium, and is inserted into perineal center in male, into the side of sphincter vaginae in the female. It steadies the perineal center. Nerve, perineal. Compressor urethrje (to. constrictor uretlirce) arises from the rami of pubis and ischium, passes inward and with its fellow of the other side, unites above and below the urethra, surrounding it from the bulbous portion to the prostate gland in the male. In the female it is inserted into the vaginal walls. External sphincter ani (to. sphincter ani externus) muscle arises from the apex of the coccyx and the superficial fascia, and is inserted into the perineal center, blending with the levator ani, accelerator urinae and transversus perinaei. It closes the anus. Nerve, fourth sacral. Internal sphincter ani (to. sphincter ani internus) is an aggregation of circular fibers of the intestine, forming a muscular ring one inch in breadth, surrounding the lower por- tion of the rectum. The external sphincter ani, the two bulbocavernosi and the two transversus perinsei unite one-half inch in front of the anus in the median line in the so-called central tendon of the per- ineum. Anal elevator (levator ani) arises in front from body and ramus of pubis and symphysis, posteriorly from spine of ischium, and on either side from angle -of arcus tendinew (white line). It is inserted by posterior fibers into coccyx, and ante- rior fibers into the muscle of opposite side in a median raphe extending from coccyx to anus ; middle fibers into rectum, join- ing with sphincter; anterior to prostate, blending with external sphincter and transverse fascia. In the female it is inserted into the vagina instead of the prostate. It helps with its fellow to form the floor of the pelvis, and supports the vagina, rectum and pelvic viscera. Nerves, infe- rior hemorrhoidal and fourth sacral. Cocctgeus muscle (coccygei) , from the spine of the ischium and lesser sacrosciatic ligament, and is inserted into the margin of coccyx and side of lower two segments of sacrum. SURGICAL ANATOMY. 407 It raises the coccyx and forms the posterior part of the pelvic floor. Nerve, anterior division of the fourth and fifth sacral. The muscles of the female perineum are : — Superficial Set. Sphincter vagina', or bulbocavernosus; Erector clitoridis ; Transversus perinaei superficial ; Sphincter ani ; Levator ani : Coccygeus. Deep Set. Transversus perinaei (profundus) ; ( SonstrictoT vagina? ; Compressor urethra1. The structures beneath the deep layer of the superficial fascia are : — Erector penis muscle ; Accelerator urinae muscle ; Transversus perinaei muscle and arteries; Superficial perineal vessels. The structures between the two layers of the deep perineal fascia are : — ( 'ompressor urethra? muscle; Membranous urethra; Subpubic ligament ; Dorsal vein of penis; Pudic vessels and nerves; Venous plexus; Cowper's glands and ducts; Arteries and nerves of bulb. ANATOMY OF LATERAL LITHOTOMY. The incision is made through a triangular interval formed by the transversus perinaei, accelerator urinae and erector penie muscles, and divides the following structures in the order named : — Skin and superficial fascia : Inferior hemorrhoidal vessels and nerves; Accelerator urinae muscle ; Superficial perineal vessels and nerve (sometimes); 408 HUMAN ANATOMY. Transversus perinsei artery and muscle; Deep perineal fascia; Anterior part of levator ani; Part of compressor urethras and accelerator urinse muscles; Membranous and prostatic portions of urethra; Left lobe of prostate gland. The structures to be avoided are : — In median line, rectum and bulb of corpus spongiosum ; Externally, internal pudic artery; Forward, artery of the bulb ; Backward, posterior part of prostatic gland and neck of the bladder. DENTAL ANATOMY. Embkyologically the oral cavity is formed by the first visceral and the frontonasal process. The first arch distad branches dichotomously. The superior arm is the maxillary process, the inferior arm the mandibular process. The latter joins with its fellow of the opposite side in the midline {sym- physis nuutdibularis) to form the lower jaw. Between the right Fig. 146. Maxillary and mandibular process of first visceral arch and frontonasal process, a, frontonasal process; b, superior arm (maxil- lary process) of first arch; C, inferior arm (mandibular process) of first arch; ii is 412 HUMAN ANATOMY. a delicate fold of tissue in the midline between the gum of the external alveolar plate and the upper lip. In artificial dentures room has to, be provided for its free movement. The body of the cheeks and lips is composed of the muscles of expression (Fig. 149). Internally they are covered by mucosa, externally by the skin. fig. 149. Muscles of expression. Muscles of the right side of the head and neck. 1, frontalis: 2, superior auricular; 3, posterior auricular; 4, orbicularis palpebarum; 5, pyramidalis nasi; 6, compressor naris; 7, levator labii superioris, alaeque nasi; 8, levator labii superioris; 9, zygomaticus major; 10, orbicularis oris; 11, depressor labii inferioris; 12, depressor anguli oris; 13, anterior belly of digestric; 14, mylo- hyoid; 15, hyoglossus; 16, stylohyoid; 17, posterior belly of digastric; 18, masseter; 19, sternohyoid; 20, anterior belly of omohyoid; 21, thyrohyoid; 22, 23, lower and middle constrictors of the pharynx; 24, sternomastoid ; 25, 26, splenius; 27, levator scapulae; 28, anterior scalenus; 29, posterior belly of omohyoid; 30, middle and posterior scalenus; 31, trapezius. ("Applied Anatomy and Oral Surgery," Robert H. Ivy.) The anterior orifice of the oral cavity (rima oris) is a trans- verse slit bounded by the projecting lips. The orbicularis oris (page 132) has no separate existence. The sphincter action of the lips is accomplished by a complicated interlacing of the muscle fibers from buccinator, depressor labii inferioris, depres- sor anguli oris, zygomaticus and risorius. The buccinator forms the muscular body of the cheek. Its attachment to the maxilla has to be considered in outlining upper artificial dentures. DENTAL ANATOMY. 413 The masseter, temporal, external and internal pterygoids all innervated by the fifth nerve, are the "muscles of mastica- tion" (pages 132 and 133). Between the right and left halves of the body of the mandible is the muscular floor of the mouth — digastric, geniohyoid and mylohyoid (.page 134). The floor of the oral cavity presents the tongue, with its frenum; on each side of this an elevation indicating the posi- tion of the sublingual glands, and anteriorly to the frenum, a papilla for the orifices of the sublingual and submaxillary ducts. For the description of the tongue see pages 135 and 388. For the salivary glands see pages 229-230. Tooth development. sr,ry \,, pharyngeal plexus on middle constrictor. Mucous membrane of interior of larynx and arytnioideus muscle. Exterior laryngeal i '"' " yroid muscles, branch pharyngeal plexus. ■thyroid. Two ficial card: To deep cardi sple: n left. Anterior pulmonary plexUS. Posterior pulmonary plexus. Plexus gulfe. arouud the ccsophagus. Stomach, communicating with solar, splenic, and hepatic plexuses. eating branches to pharyngeal and superior laryngeal branches of the pneuuio- id, trapezius, cervical plexus, occasionally great auricular nerve. Omo-byoid (both bellies), sternodiyoid, sterno-thyroid. Thyro hyoid muscle. Styloglossus, hyoglossus, geuio-hyoid, genio hyogloesua. With pm-uuiog's.-.trie, sympathetic, first and scpjji.l cervical and gll-talory I Termination 0 Functio Schneiderian muco brane of the nosi Nerve of special foramen. Nerve of special i In the superior obliqui (The trifacial is the large: ■gesk In muscles. Nerve of i Nerve of special Nerve of motion. Nerve of facial expression. Internal ear. Nerve of special si muscle. Nerve of special s( In muscles, mucous r brane, vessels, g (thyroid). Nerves of motion, si tion, and sympathy. In muscles. Nerve of tni INDEX. Abdomen. muscles of. 1-13 regions of, 2'M Acervulus cerebri, 315 Acetabulum, 81 Alveoli of lower jaw, 36 of upper jaw. 31 Amphiarthrosis. Anastomoses. 174 Anatomy, descriptive, 1 dental, 409 general. 1 of hernia. 396 of lateral lithotomy, 407 of teeth, 422 surgical, 391 Angle of jaw. :'.7 Anterior perforated space. 306 Annulus ovalis, 171 of subscapular fossa. 65 Antrum of Ilighmore. 29, 413 relation of roots of teeth to. 413 Anus, I'll Aorta, 17s, 193 Appendicies epiploic*, 244 Appendix auriculae, 170, 172 ensiform, '^ vermiformis, 243 Aponeuroses. 129 pharyng. al, 230 Apophysis, 4 Aqueduct of Svlvius (iter e tertio, etc.), 299 Aqueductus cochlea-, is fallopii. is. ::us vestibule, 18 Arachnoid, of brain. L"'7 of spinal cord, ::::! Arbor vita-. 290, 285 Arch, deep palmar, 192 plantar, 202 Areola of mammae, 292 Arm, 63 Arnold's ganglion. 352 propria- renales, _•;* Arterial anastomoses, 171 in, table of. 202 Arteries, belicine, 280 hepatic, 194, 249 lingual, pharyngeal sublingual, 180, isi Arteries of the ankle-joint, 125 auditory canal. 36i, 376 auricle, 376 bom choroid. 363 elbow |oint, 123 tcbian tube, .:7:i ::7I hip-joint, 120 Arteries of the iris, 364 kidney, 268 knee-joint. 121 labyrinth, 382 larynx, 259 lung, it,:: mamma-, 292 mambrana tympani, 380 nasal fossa?, 356 nose, 355 oesophagus, 193 ovaries, 284 pancreas, 246 penis, 280 pharynx, 230 retina, 36S shoulder-joint, 116 spleen, 251 stomach, 240 suprarenal capsules, 253 testicle, 278 thymus gland, 25L' thyroid gland, 252 tympanum, 380 uterus. :!xi wrist-joint, lis Artery, anastomotica magna, 191, 200 aorta, 178, 193 arteria reeeptaculi, is:, auricular, posterior, 183 axillary, 150 basilar, 188 brachial, 191 carotid, common. 17:i external, 180 internal, 185 carotid (external I, 180 (internal I. 185 cerebral, anterior. 186 middle, anterior, 186 posterior, 186 choroid, anterior, 186 circumflex, anterior. 189, 191 posterior, 191 communicating, anterior, 186 posterior, 186 coronary, 178, 179 dental, inferior. L84 digital, 193, 202 dorsalis, hallucls, 201 pedis, 201 i -trie, deep, 198 superficial, 199 I. 181 lc, 194 femoral, L99 gluteal, 198 irrhoidal, inferior, 197 middle. 197 , 195 patlc, 194 iliac, circumflex, deep, 198 I 123) 424 INDEX. Artery, iliac, common, 196 external, 106, 198 internal, 194, 196 superficial, 199 ilio-lumbar, 198 infraorbital, 185 innominate, 179 intercostal, 189 lingual, 180 lumbar, 195 mammary, internal, 189 maxillary, internal, 183 meningeal, anterior, 185 middle, 184 posterior, 188 small, 184 mesenteric, superior, 194 inferior, 195 obturator, 197 occipital, 181 ophthalmic, 185 palmar arches, 192 peroneal, 202 pharyngeal, ascending, 183 phrenic, 194 plantar, external, 202 internal, 202 popliteal, 200 profunda femoris, 199 inferior, 191 superior, 191 pudic, deep external, 199 superficial external, 199 internal, 197 pulmonary, 208 radial, 191 renal, 195, 268 sacral, lateral, 198 middle, 196 sciatic, 197 spermatic, 195 splenic, 194 subclavian, 187 subscapular, 190 suprarenal, 195 suprascapular, 189 temporal, 183 thoracic, acromial, 190 alar, 190 long, 189, 190 superior, 190 thyroid, inferior, 187 superior, 180 thyroidea ima, 252 tibial, anterior, 200 posterior, 201 transveralis colli, 189 tympanic, 184 ulnar, 192 uterine, 198 vaginal, 198 vertebral, 188 vesical, 196 Arthrodial joints, 105 Articulations, 104 acromioclavicular, 115 atloaxoid, 107 carpal, 119 carpometacarpal, 119 costosternal, 108 costotransverse, 108 costovertebral, 108 glenohumeral, 116 occipitoatloid, 107 Articulations, occipitoaxoid, 107 phalangeal, of foot, 127 of hand, 120 radioulnar, 117 sacrococcygeal, 111 sacroiliac, 110 sacroischiatic, 110 sacrovertebral, 110 sternoclavicular, 114 tarsal, 125 tarsometatarsal, 126 temporomaxillary, 105 tibiofibular, 123 vertebral, 106 Arytenoid cartilages, 254 Atlas, 58 Auricle, 376 Axilla, 395 Axis, 44, 58 cerebrospinal, 296 celiac, 194 optic, 358 thyroid, 188 visual, 358 Aaxis cylinder, 295 Azygos veins, 215 uvulaa, 136 Bartholin, duct of, 230 glands of, 292 Bauhin, valve of. 243 Bell, external nerve of, 336 internal nerve of, 336 Bladder, 270 Bodies, Malpighian, 268 Pacchionian, 297 geniculate, 311 restiform, 299 Body, ciliary, 363 olivary, 299 pituitary, 306 restiform, 299 Bone, astragalus, 93 calcaneum, 93 clavicle, 63 coccyx, 63 cuboid, 93 cuneiform, 78, 94 ethmoid, 24 femur, 86 fibula, 92 frontal, 8 humerus, 70 ilium, 82 incus, 380 hyoid, 38 innominate, 81 ischium, 84 lachrymal, 26 malar, 27 malleus, 380 maxillary, inferior, 35 superior, 29 nasal, 27 occipital, 12 os calcis, 93 os magnum, 78 palate, 32 parietal, 10 patella, 89 peroneal, 92 phalanges of foot, 96 of hand, 79 pisiform, 78 INDEX. 425 Bone, pubic, S5 radius. ~i sacrum. 61_ scaphoid, 77, 94 scapula, 65 semilunar, 78 sessamoid, 98 sphenoid, 20 stapes. 3S0 sternum, 52 structure, microscopic, 5 temporal, 15 tibia, 90 trapezium, 78 trapezoid. 78 turbinated, inferior, 34 middle, 25 superior, 25 ulna, 72 unciform. 78 vomer. 35 Bones of the body, 3 of the carpus, 76 of the cranium. 8 of the face, 26 of the foot, 93 of the hand, 76 of hard palate, 409 of the lower extremity, 81 of the metacarpus, 76 of the metatarsus, 96 of oral cavity, 409 of the tarsus, 93 of the trunk. 52 of the upper extremity, 63 Wormian, 40 Bonnet, capsule of. 360 Bowman, capsule of, 268 Brain, commissures of, 308 development, 298 divisions of, 298 ventricles, 308 weights, average, 29S Bronchi, 259 Brown stria of Retzius, 417 Brunner's gland, 242 Buccinator, 412 in artificial dentures, 412 Bulb of corpus cavernoflum, 280 of corpus spongiosum, 280 Bulbi vfstibuli, 291 Bulbs, olfactory, 323 Burns, process of, 402 Bursas of hip, 121 of knee, 122 Cecum, 243 f'alamus Bcriptorius, 313 Calices of kidney. 268 Canal, alimentary, 223 auditory external, 376 Internal, 18, 387 carotid, 18 central, of the cord, 332 crural. (00 I. 30 loral, io mastoid. 16 mi dium, 16 III. lit;, I. 36, 110 malar. II nutrient, .", obtui of Monro, 312 Foramen of Scarpa, 31 of Stenson, 31 of Winslow. 235 olfactory, 46 orbital. 24 ovale, 23 palatine, 31 posterius, 13 pterygopalatine, 22 rotundum, 23 sacral, 62 sphenopalatine, 32 spinal. 57 spinosum, 23 stylomastoid, 10 supraorbital, 8, 46 suprascapular, 67 thyroid, 81 Vesalii, 23 Foramina at the base of the skull 46 cf hard palate, 409 of mandible, 410 Thebisii, 171 Fort arm, 72, 14'.! Fcrmatio reticularis, 316 Fornix, 31] Fossa, canine, 20 caronoid, 71 digastric, 17 digital, 88 glenoid, 16, 68 hyaloid, 369 incisive, 20 infraspinous, 66 ischorectal, 402 jugular, 18 lachrymal, 0 navicularis of penis, 281 of vulvo, 289 olecranon, 71 ovalis, 171 pituitary, 21 pterygoid, 24 scaphoid, 24 sphenomaxillary, 44 sublingual, 36 submaxillary, 36 subscapular, 66 supraspinous, 66 temporal, 40 zygomatic, 40 Fossae, nasal, 45 of the skull, 40 Fourchette, 290 Fovea centralis. 367 hemispheric;!. 383 semielliptica. 383 Fr< nuluni, 31 1 Freiium lobii, II 1 in relation to artificial dentures, 412 preputii, 280 Call-bladder, 250 Call-duct, 250 Ganglia, 295 basic, 311 cervical, 3.",2 crania I I iO lumbar. :::>'■'. of fifth nerve of the sympathetic, 350 sacra I thoracic. 428 INDEX. Ganglion, Arnold's, 352 Gasserian, 350 irnpar, 350 Meckel's, 350 of Ribes, 350 ophthalmic, 350 otic, 352 sphenopalatine, 350 spirale, 385 submaxillary, 352 Gasser, ganglion of, 350 Geniculate bodies, 311 Ginglymus, 105 Gingival trough, 411 Gladiolus, 52 Gland, lachrymal, 374 mammary, 292 parotid, 229 pineal, 315 prostate, 282 sublingual, 230 submaxillary, 229 thymoid, 252 thymus, 252 Glandula coccygea, 253 Glandula intercarotica, 253 Glands, Brunner's, 242 Cowper's, 282 ductless, 250 gastric, 239 intestinal, 242 lachrymal, 373 lymphatic, 219 Meibomian, 373 of Bartholine, 292 of Tyson, 279, 292 peptic, 240 salivary, 229 sebaceous, 390 solitary, 242 sweat, 390 Glans clitoridis, 290 penis, 278 Glenoid, 16, 68 Glisson's capsule, 246 Glottis, 255 Gomphosis, 105, 414 Graafian vesicle, 286 Groove, bicipital, 70 carotid, 21 cavernous, 21 infraorbital, 31 musculospiral, 70 mylohyoid, 36 nasopalatine, 35 optic, 21 Gubernaculum testis, 274 Gums, 224, 411 Gyri of brain, 305 Hairs, 390 Hamstrings, 160 Hamular process, 26 Hand, 76 Hare-lip, 409 Hard palate, 409 sutures of, 409 foramina of, 409 bones of, 409 Haversian canals, 6 system, 6 Head of the femur, 86 of the humerus, 70 of the ulna, 72 Heart, 170 Helicotrema, 385 Helix, 376 Henle, tubes of, 268 Hernia, anatomy of, 396 femoral, 400 inguinal, 396 Hiatus Fallopii, 18 Hirlus of kidney, 266 Hippocampus major, 312; minor, 312 His, bundle of, 171 Histogenesis of dental tissues, 416 of enamel, 416 of dentin, 416 of cement, 416 Hasner, valve of, 375 Houston, valves of, 245 Huguier. canal of, 16 Humerus, 70 Humor, aqueous, 368 vitreous, 368 Humors of the eyeball, 368 Hunter's canal, 199 Hydatids of Morgagni, 276, 288 Hymen, 288 Hypochondrium, 234 Hypogastrium, 234 Ileum, 241 Ilium, 82 Incisura intertragica, 376 Incisal pad, 411 Infundibulum of brain, 306 Inguinal regions, 234 Intestine, large, 242 small, 240 Internal capsule, 310 Iris, 363 Ischium, 84 Iter chordae anterius, 378 posterius, 378 e tertio ad quartum ventriculum 313 Ivory, 226 Jacob's membrane, 368 Jacobson's nerve, 3,16, 336, 365 Jejunum, 241 Joint, ankle-, 124 elbow-, 116 hip-, 120 rotators of the, 159 knee-, 121 shoulder-, 114 wrist-, 118 Joints, motions in, 105 structures of, 104 Kidneys, 266 Labia majora, 290 minora, 290 Labium tympanicum, 386 vestibulare, 386 Labyrinth, membranous, 385 osseous, 383 Lacteals, 219 Lacuna magna, 282 Lacunae, 6 Lacus lachrymalis, 373 Lamellae, 6 Lamina cinerea, 306 cribrosa (of sclerotic), 361 (of temporal bone), 18 IXDEX. 429 Lamina fusca, 361 reticularis. 387 spiralis, 3S4 Lamina of cornea, 361 Lanoisi, nerves of, 308 Larynx, 254 Layer, dermoid, 389 ganglionic, 367 molecular. 367 Leg, 90 Lens, crystalline, 36S Lieberkiihn, crypts or follicles of. Ligament, annular, of foot. 16S of hand, 167 rhondroxiphoid. 109 conoid, 115 coronary, 121 cotyloid. 121 crucial, of knee-joint, 122 deltoid, 124 falciform, 401 Giiubernat's. 143, 398 400 glenoid, 116 iliofemoral or Y, 121 of Burns. 402 of Colles, 398 of Hey, 381 of Zina, 353,' 369 orbicular, 118 Poupart's. 143, 146. 398, 400 periodontal, 418 rhomboid, 114 round, of the liver, 249 of the uterus, 285 sacrosciatic, 110 stellate, 108 stylomaxillarv, 106 suspensory, of lens, 360 of liver, 249 of penis, 280 of spleen, 250 transverse, of hip-joint 121 trapezoid. 115 triangular, of perineum 104 Ligaments of the bladder, 271 broad, of uterus, 285 cheek, 107 of the knee-joint. 121 of the larynx, 255 of the liver, 248 of the ossicles, 381 of the ovaries, 285 of the sternum, 109 of the uterus, 284 peritoneal, 236-237 tarsal, 373 Ligamentum latum pulmonis 264 mucosum, 128 nuchas, 106 patella?, 121 pectlnatum iridis, 363 Buspensorium, 108 teres, 121 Winslowii, 121 Limbua lamina spiralis 385 Line, intertrochanteric, 88 Linea-aspera, 86 lliopectlnea, 81 quadrat! Llngula, i'i Liquor Uorgagni, 869 Lithotomy, structures affected 107 Liver 1M7 ligaments, 249 Liver, lobes of, 248 structure of, 246 vessels of, 249 Lobes of the cerebellum, 301 of the cerebrum. 302 of the liver, 248 Lobule of the ear, 375 Lobules of the liver, 246 of the lung, 262 Lobulus caudatus. 246 quadratus, 246 Spigelii, 246 Lower, tubercle of. 171 Lumbar regions. 234 Lung, broad ligament of 264 Lungs, 261 Lymphatics. 219 hepatic, 222, 249 of lung, 263 of penis, 281 of trachea and bronchi, 260 of uterus, 284 Lymph spaces of eyeball 360 Lyra, 311 Macula acoustica, 385 c.ribrosa, 383 lutea, 367 Malleolus, 92 Malpighi, bodies of, 251 pyramids of, 267 Malpighian corpuscles, 251. 268 tuft, 268 Mamma?, 292 Mandible, 410 Mandibular, canine, 421 fifth cuspid, 422 first incisor, 420 first molar, 421 first premolar, 421 fourth cuspid, 421 second incisor, 420 second molar. 421 second premolar, 421 third molar, 421 Manubrium of the malleus, 380 of the sternum. 52 Marrow of bone, 5 Masses, lateral, of ethmoid, 24 Mastoid process, 16 Maxillary, canine, 419 first, 419 first molar, 419 first premolar, 419 second, 419 second molar, 419 second premolar, 419 third molar, 420 Meatus auditorius externus 18 interims, 18, ::s; nasi. 1:, urinarius, 281, 291 Meckel's ganglion, 350 •Mediastinum, 264 testis, 274 Medullar, 5 oblongata, 299. 315 Medullary sheath, 294 substance, 266 Membrana basillaria 3S6 ' anuloi a, 286 prseformativa, 416 puplllariB, 364 Ruyschlana 430 INDEX. Menibrana tectoria, 387 tyrnpani, 380 secundaria, 362 Membrane, hyaloid, 369 interosseous, 119, 120, 124 Jacob's, 367 limiting, 367 of Descemet, 364 Nasrnyth's, 415, 417 of Reissner, 387 Schneiderian, 356 Shrapnell's, 378 vitelline, 286 Membranes of brain, 296 of the spinal cord, 332 synovial, 104 tarsal synovial, 126 wrist synovial, 120 Meridians of eyeball, 359 Mesencephalon, 314 Mesenteries, 236 Metacarpus, 80 Metatarsus, 96 Midbrain, 318 Modiolus, 384 Monro, foramen of, 312 Mons veneris, 290 Montgomery, glands of, 292 Morgagni, hydatids of, 276, 288 liquor, 369 Mouth, 223 Muller, duct of, 276, 288 Muscle, ciliary, 364 cremaster, 274 dilator pupillse, 364 levator glandulse thyroidea?, of the auricle, 376 sphincter pupillas, 365 Muscles of the abdomen, 143 of Hilton, 258 of Houston, 405 of the arm, 148 of the back, 138 of the ear (external), 129 of the epiglottis, 258 of expression, 412 of the Eustachian tube, 379 of the eyeball, 369 of the face, 130 of the forearm, 149 of the foot, 163 of the hand, 153 of the head, 129 of the hip, 155 of the larynx, 257 of the leg, 160 of mandible, 414 of mastication, 413 of the neck, 133 of the palate, 135 of the perineum, 407 of the pharynx, 135 of the shoulder, 147 * of the stomach, 239 of the thigh, 156 of the thorax, 145 of the tongue, 135 of the tympanum, 382 Muscular fiber, 128 floor of mouth, 413 Musculi pectinati, 172 Naboth ovules of, 284 Nasmyth's membrane, 415, 417 Nails, 390 Nares, 45 Nates of cerebrum, 314 Neck of femur, 88 of humerus, 70 muscles of, 133 triangles of, 391 Nerve, abducens, 326 Arnold's, 330 auditory, 328 Bell's external respiratory, 336 internal respiratory, 334 cells, 295 chorda tympani, 328 circumflex, 336 cochlear, 328 crural, anterior, 340 facial, 326 fibers, 295 genitocrural, 339 glossopharyngeal, 330 gluteal, superior, 340 hypoglossal, 331 iliohypogastric, 339 ilioinguinal, 339 intercostal, 339 intercostohumeral, 340 internal cutaneous, 337 interosseous, 337, 338 Jacobson's, 330, 382 laryngeal, 259, 330 maxillary inferior, 326 superior, 325 median, 337 motor oculi, 324 musculocutaneous, 337 musculospiral, 338 nonus or ninth pair, 330 obturator, 340 of Wrisburg, 337 olfactory, 323 ophthalmic division of fifth, 325 optic, 323 par vagum, 330 patheticus, 325 petrosal, external, 351 large deep, 351 large superficial, 351 small, 351 popliteal, 341 phrenic, 334 pneumogastric, 330 pterygopalatine, 351 pudic, 341 radial, 338 sciatic, great, 341 small, 341 spinal accessory, 331 subscapular, 336 suprascapular, 336 sympathetic, 350 tibial, anterior, 342 posterior, 341 thoracic, 336 trigeminus, 325 trochlear, 325 tympanic, 328 ulnar, 337 vestibular, 328 Vidian, 351 Nerve-fibers, optic, 323 tissue, 295 Nerves, 295 cranial, 323 INDEX. 431 Nerves, hepatic, 354 of ankle-joint, 125 of auditory canal, 377 of auricle. 370 of bladder. 273 of choroid, 363 of elbow-joint. 117 of Eustachian tube, 380 of eye, 372 of eyelids. 372 of eye-muscles, 363. 370, 371 of hip-joint. 121 of iris. 364 of Jacobson. 330 of kidney. 270 of knee-joint. 123 of Lancisi. 308 of larynx, 259, 330 of lung, 263 of membrana tymnani, 380 of nasal fossie, 356 of nose, 356 of pulp, 418 of ovaries, 287 of pancreas, 246 of pharynx, 230 of shoulder-joint, 115, 117 of spleen, 251 of stomach, 240 of suprarenal capsules, 253 of taste, 389 of testicles. 278 of thymus gland. 252 of tongue, 389 of trachea and bronchi, 260 of tympanum, 378 of uterus. _'M of wrist-joint, 119 petrosal. 351 popliteal. 341 sciatic. :;n spinal. 332 splanchnics, 353 table of, 343 thyroid, 252 Nervous system, 294 Newman, sheath of, 418 N« unlemma, 294 Neuroglia. 295 Neuroi Nipple. 292 Nose, 355 Notch, cotyloid. 81 Interclavicular, 41 intercondyloid, 71 popliteal. 60 Kivinus. 378 tic, 84, 85 old, 37 supraorbital, 8 suprascapular, 67 Nuck, canal of. 237 Nucleus, red, 318 Nymphs,