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RSE S TET vv eeu ‘ the od gcceS 6s PAK - = i eh Fy Some t : ] ‘ ' i Siew Tap ee q . > ‘ re ok uty ; = \ . re Lage | P ees A -_ ] is : ¥ Ail, 2 a ~ 1 4 Y_ © Var. ¥ . é Fad I EM) eh IN : >. RP ars — i . UJ = is —_ a par a® fi ~ PN Raa ees aS ¢. aAna® A Ac amaranas AAR Y aN em a 36 1 La be React pay ialay.Y wee . vi : a=: mamaMa@e RaMeanaar i an ~ , ™ rE NA} 3 VINA alee! lV AP Pn PYAY aR ~~ Dem Nala ASA A OO lo OW Y “Ae x - 3—~ Ly WN \ — 7. a a ro e | Cots <>) nage ma a « Cc ZO G ; : 4 q ac OCG i m ¢ ( OS a @C CLG - Gi Yeas LaQuGa @ es / ex. (an rae @ a < me (A me We (CE . a ( c | r r 4 (4 Zan an "4 t C a q « re -, Cre a An Ce CE 7 on i -a® A ar HITHERTO UNPUBLISHED PLATES OF TERTIARY MAMMALIA AND - PERMIAN VERTEBRATA Prepared under the Direction of EDWARD DRINKER COPE for the di U. S. Geological Survey of the Territories with Descriptions of Plates by WILLIAM DILLER MATTHEW PUBLISHED AND DISTRIBUTED WITH THE COOPERATION OF T UNITED STATES GEOLOGICAL SURVEY BY THE AMERICAN MUSEUM OF NATURAL HISTORY et ASF. MONOGRAPH SERIES NUMBER 2 gS 1915 | vy PREFATORY NOTE. The accompanying plates were prepared and printed by the United States Govern- ment for the Report on Tertiary Vertebrata by E. D. Cope, forming Volumes III and IV of the final report of the Geological and Geographical Survey of the Territories, F. Y. Hayden in charge. The first part of this Report (Volume III), treating of the Eocene and a part of the Oligocene vertebrates was completed and published in 1885. The second part, treating of the remainder of the Oligocene and all the Miocene vertebrates, was never completed. A large number, but not all, of the plates for this part were pre- pared and printed, together with seven plates for a contemplated report on Permian Vertebrata. The specimens illustrated are contained in the Cope Collection, purchased by the American Museum of Natural History. The ownership of certain parts of the collec- tion having been satisfactorily adjusted it seemed proper that the plates should be distributed for the benefit of palzeontologists, jointly by the United States Geological Survey and the American Museum of Natural History. The plate descriptions were prepared and the plates bound and distributed by the latter institution. Illustrations of many of the specimens here figured, copied from these plates, have been published by Professor Cope, chiefly in the American Naturalist of 1880-1888. The nomenclature and identification of the material has been materially changed since 1880-1883 when the illustrations were made, and it has been thought advisable to give in the plate descriptions both Cope’s identifications and the revision now current (1915). No text is known to have been prepared by the author, but the substance of his views and interpretations of the later Tertiary vertebrates of Western America may be found in his numerous publications dealing with fossil mammals, chiefly in the American Naturalist, and Proceedings of the American Philosophical Society, between 1880 and 1892, also in Bulletins and Annual Reports of the U. S. Geol. Geog. Sur. Territories, 1872-1882 and the Texas Geological Survey Reports, 1891-1893. A number of plates illustrating the Brontotheriide of the Oligocene of Colorado are included in the Monograph of that family by Henry Fairfield Osborn, and are omitted from the present series. SCHEDULE OF PLATES. Permian Vertebrata. Twelve plates; Nos. 1, 3, 7,8, 10, 11, 12, 13, 16, 16a, 19, 22. Oligocene Ungulata. Fifteen plates; Nos. 102, 104, 105, 108, 109, 109a, 110, 110a, ik, Ths ahiB, bist, wali, Waleyy eG. Miocene Carnivora and Rodentia. Seven plates; Nos. 118, 118a, 118b, 119, 119a, 119b, 119¢e. Miocene Proboscidea. Five plates; Nos. 120, 121, 122, 123, 124. Miocene Rhinoceroses. Twenty plates; Nos. 125, 126, 127, 129, 150, 131, 132, 133, 134, 135, 136, 137, 138, 139, 140, 141, 142, 143, 144, 144b. Miocene Equide. Seven plates; Nos. 145, 147, 148, 148a, 149, 150, 151. Miocene Camelide. Two plates; Nos. 153, 154, w it PEA BE 1. PERMIAN VERTEBRATA. MEGALICHTHYS Nomenclature Cope 1880-3 Revised 1915 Ectosteorhachis nitidus Megalichthys (= Palzobatrachus) nitidus lias. 1-38. Megalichthys nitidus (Cope 1880). Type specimen, two-thirds natural size. Amer. Mus. No. 7239, Cope Coll. Wichita basin, Texas. Fig. 1, ventral view of head and fore part of trunk; fig. 2, 2a, 2b, dorsal, lateral and anterior views of head. (Fig. 4 is not identified). [LIN SIHOVHYOHLSOLOG - Oo OO Fie le ik PERMIAN VERTEBRATA VERTEBRATE REMAINS FROM VERMILLION Co., ILLINOIS Nomenclature Cope, 1880-3 Revised, 1915 Orthacanthus quadriseriatus Pleuracanthus quadriseriatus Diplodus compressus Diacranodus texensis Ctenodus pusillus Gnathorhiza pusilla a gurleyanus Sagenodus fossatus + fossatus i. fossatus Ceratodus paucicristatus : paucicristatus “_-vinslovii a vinslovii Peplorhina arctata Spherolepis arctata Janassa strigilina Janassa strigilina “ gurleiana “ gurleiana Archeobelus vellicatus Archeobelus vellicatus Cricotus heteroclitus » Cricotus heteroclitus Lysorophus tricarinatus Lysorophus tricarinatus Diplocaulus salamandroides Diplocaulus salamandroides Fies. 1-50. Vertebrate remains from the Permian bone bed of Vermillion County Illinois. This fauna described by Professor Cope in 1877 was the first Permian fauna found in North America, and contains the genotypes of many of the genera later described from better material in Texas and New Mexico. The collection is the property of the Walker Museum of the University of Chicago, and has been redescribed by Dr. E. C. Case (1900) in the Journal of Geology, Vol. VIII, pp. 698-729, with five plates of figures. The revised nomenclature is auct. Case and Hussakof 1911. Fias. 1, la. Plewracanthus quadriseriatus (Cope, 1877). Fragment of head spine, X 2. Type specimen, No. 6502, Walker Mus. (Case, 1900; pl. i, fig. 3). Frias. 2-4. Diacranodus texensis (Cope, 1890). Teeth, No. 6504, Walker Mus., referred by Cope to Diplodus (Didymodus) compressus Newberry. (Case, 1900, pl. i, fig. 5; I am unable to state the enlargement, as Dr. Case does not give it in his figure, nor have measurements been published). Fic. 5. Spherolepis arctata (Cope, 1877). Dentigerous plate, x 3. Type specimen No. 6511, Walker Museum. (Hussakof, 1911, pl. xxxi, fig. 2). G. 6, 6a, 6b, 6c. Gnaithorhiza pusilla (Cope, 1877). Right palatal plate, x 3, type specimen, No. 6508, Walker Mus. (Case, 1900, pl. i, fig. 9; Hussakof, 1911, pl. xxvii, fig. 10). Fia. 7, 7a, 7b. Sagenodus fossatus (Cope, 1877). Upper dental plate, natural size, No. 6509, Walker Mus., type of Ctenodus gurleyanus Cope, 1877. (Case, 1900, pl. i, fig. 8; Hussakof, 1911, pl. xxvi, fig. 9). Fia. 8, 8a, 8b, 8c. Sagenodus fossatus (Cope, 1877). Imperfect left mandibular plate, immature, about $ natural size. Type specimen, No. 6506, Walker Museum. (Case, 1900, pl. i, fig. 10; Hussakof, 1911, pl. xxvi, fig. 8). Fic. 9, 9a, 9b, 9c. Sagenodus pancicristatus Cope, 1877. Imperfect dental plate, natural size, Type specimen, No. 6505 Walker Mus. (Case, 1900, pl. i, fig. 11; Hussakof, 1911, pl. xxvii, fig. 4). FE I Kia. Fia. Fia. Hie. Fig. Fia. Fics. lies. Kies. Fic. / . 10, 10a, 10b, 10e. Sagenodus vinslovii (Cope, 1875). Incomplete left palatal plate, natural size, Type specimen, No. 6507 Walker Mus. (Case, 1900, pl. i, fig. 6; Hussakof, 1911, pl. xxvii, fig. 7). 3. 11, lla, 1lb. Janassa strigilina Cope, 1881. Tooth probably of median series, twice natural size. Type specimen, No. 6500 Walker Mus. (Case, 1900, pl. 1, fig. 1; Hussakof, 1911, pl. xxvi, fig. 1). 12, 12a, 12b. Janassa gurleiana Cope, 1878. Small tooth, lacking the root, twice natural size. Type specimen, No. 6501, Walker Mus. (Case, 1900, pl. i, fig. 2; Hussakof, 1911, pl. xxvi, fig. 2). ;. 18, 18a, 13b, 18¢e. Arch@obelus vellicatus Cope, 1877. Jaw fragment with tooth, about natural size. Type specimen, No. 6524, Walker Mus. . 14, 14a. Undetermined jaw fragment, ‘Species two” of Cope, 1877, auct. Case (agrees better with “species one” of Cope’s description), natural size and enlarged to 7. No. 6567, Walker Mus. (Case, 1900, pl. v, fig. 12). Fig. 14b, tooth, undet. . 15, 16. Cricotus heteroclitus Cope, 1875. Two intercentra, ? types of C. discophorus Cope, 1877, natural size, Walker Mus.,? No. 6518. (Case, 1900, pl.i, figs. 13-14). . 17, 17a, 17b. Cricotus gibsoni Cope, 1877. Supposed caudal vertebra, natural size. Type specimen, No. 6521, Walker Mus. Case, 1900, pl. i, fig. 15. 18. ? Diplocaulus, fragment of sculptured plate. 19-20. Lysorophus tricarinatus Cope, 1877. Two vertebral centra, twice natural size. Typical specimens. No. 6526-8, Walker Mus. (Case, 1900, pl. ii, fig. 12). 21-23. Diplocaulus salamandroides Cope, 1877. Vertebre, about 3 natural size. Typical specimens, Nos. 6513-6, Walker Mus. (Case, 1900, pl. v, fig. 17). i, 24. Unidentified jaw fragment “Species one” of Cope, 1877, auct. Case (agrees better with “species two” of Cope’s description). No. 6567, Walker Mus. (Case, 1900, ‘pl. v, fig. 12). Probably enlarged, but as Cope gives no measurements and Case does specify the enlargement on his figure I can not determine how much. 25. Unidentified. 26, 26a, 26b. Clepsydrops ? pedunculatus Cope, 1877. Humerus, lacking distalend. Half natural size. No. 6542, Walker Mus. (Case, 1900, pl. iii, fig. 5; reference auct. Case). 27. Unidentified. 28. Unidentified vertebra. 29. Hi intercentrum. 30-35. Pelycosaurian carpal and tarsal bones, unidentified, probably Clepsydrops. No. 6559, Walker Mus. (Case, 1900, pl. v, figs. 18-21.) Enlargement not determined. 37-47. Clepsydrops metapodials and phalanges. Enlargement not determined. Nos. 6563-5, Walker Mus. (Case 1900, pl. v, figs. 3-10). 48-49. ? Podial bones of Pelycosaurians. 50. Coprolite. No. 6574, Walker Mus. co . ¢ om j-4, SELACHII 6-10. DIPNO!I li—-1l2. JANASSA 13. ARCHAEBOBELUS 19-20. LYSOROPHUS 21-23 DIPLOCAULUS 2 4-46 =... tte ry - 8 PLATE VIL PERMIAN VERTEBRATES. 'TRIMERORHACHIS Fia. 1, la. Trimerorhachis insignis Cope, 1878. Skull, superior and inferior views, natural size. Metatype, Amer. Mus. No. 4557, Cope Coll., Permian, Wichita basin, Texas, Fias. 2, 2a, 3. ?Trimerorhachis, not identified. ¥,. .CHIS AC ORH: R a leer = - oe. a = a ay abe are oe : er Paes a 7 : care & Spee eeeme Ti fydtws- 4 hie 7 - it a =m, Af ee a ee : ag Y. - PLATE VIIL PERMIAN VERTEBRATA. 'TRIMERORHACHIS Fries. 1-11. Trimerorhachis insignis Cope, 1878. Occiput, jaws, vertebre and limb bones, natural size. Type specimen, Amer. Mus., No. 4565, Cope Coll. Lower Per- mian, Wichita Basin, Texas. * TS Oo } a Ge : 7 ioe 3 st ee ay | =e, cans i iS ae a aa ray “a b a 7 Ww ey =F Syne, Pe eee ee) oe ha + ox” ele a ed a] aye PLATE CXXXIV. MioceNE RHINOCEROSES. SKULLS OF APHELOPS AND TELEOCERAS Fig. 1. Teleoceras (= Husyodon) fossiger (Cope, 1878). Occipital view of skull. Amer. Mus. No. 8383, figured on pll. CX XXI-CXXXII. Fia. 2. Teleoceras (= Eusyodon) fossiger (Cope, 1878). Top view of skull. Amer. Mus. Cope Coll. Republican River beds, Decatur Co., Kansas. Fic. 3. Aphelops megalodus Cope, 1878. Occipital view of incomplete skull. Amer. Mus. No. 8383, Pawnee Creek beds, near Pawnee Buttes, Colorado. Specimen figured on pl. CXXIX and CXXX. All figures two-fifths natural size. Fo —, =—> a - — a 7 7 as 2 7 7 a - — v7 > _ f _ = 7 - a = Tt 7 - _ _ ia _ 7 - = as = = a » 7 : Th =e: - cm on. - - rie 7 7 = to Js : 7 of =: ~ _ a eh = ON 40 : fi ae oan) me a4 = Even cj ira 7 -- axel >>. ¥ , S t ee . - _— ae co -_ _ x — . _ ~ a - = — =_ a7 ? .- = a fens ~~ ~~“ a — < - . - 7 - a 7 PLATE (Qo. MroceNr RHINOCEROSES. SKULLS OF APHELOPS AND TELEOCERAS Fig. 1. Aphelops megalodus Cope, 1873. Skull and lower jaws, occipital view, one-third natural size. Type specimen, Amer. Mus. No. 8292, Cope Coll., figured on pl. CX XV- CXXX. Pawnee Creek beds, near Pawnee Buttes, Colorado. Fic. 2. Teleoceras (= Eusyodon) fossiger (Cope, 1878). Skull, occipital view, one-third natural size. Type specimen, Amer. Mus. No. 8390, Cope Coll., figured on pl. CX XXIII. PLATERCXX XV: Miocene RHINOCEROSES. VERTEBRA OF TRELEOCERAS ~ Nomenclature Fries. 1-6. Teleoceras (= Eusyodon) fossiger (Cope, 1878). .Cervical and dorsal vertebre, two-fifths natural size. Amer. Mus. Cope Coll. Republican River beds, De- eatur Co., Kansas. PLATE SOX VI Miocene Ruinoceroses. Limp anp Foot Bones or TELEOCERAS Nomenclature Cope 1880-3 Revised 1915 Aphelops (in part) Teleoceras Fies. 1-7. Pelvis and foot bones chiefly of T'eleoceras fossiger. Amer. Mus. Cope Coll. Re- publican River beds, Decatur Co., Kansas. ene ae n 7 iff = oa iW ap Ae Miocent RHINOCEROSES. Fries. 1-9. PLATE CXXXVIII. Limb and foot-bones chiefly of Teleoceras fossiger. publican River beds, Decatur Co., Kansas. Amer. Mus. Cope Coll. Lims anp Foot Bones oF TELEOCERAS Re- PLAWE CXS Miocene RHINOCEROSES. SKULL OF APHELOPS MALACORHINUS Aphelops malacorhinus Cope, 1879. Side view of skull one-half natural size. Type specimen, Amer. Mus. No. 8381, Cope Coll. Republican River beds, Hitchcock Co., Nebraska. Specimen figured on pll. CXL-CXLI. US GEOLOGICAL SURVEY OF THE TERRITORE ee + = PLATE Cx Miocene RHINOCEROSES. SKULL OF APHELOPS MALACORHINUS Aphelops malacorhinus Cope, 1878. Superior view of skull, two-fifths natural size. Type speci- men, Amer. Mus. No. 8381, Cope Coll. Specimen figured on pl. CXLI. Republican River beds, Nebraska. Ls] ao] ce) Son Lith clair & § fALAC ORHINUS d APHELOPS PEATE CX: Miocene RHINOCEROSES. SKULL OF APHELOPS MALACORHINUS Aphelops malacorhinus Cope, 1878. Palatal view of skull, two-fifths natural size. Type speci- men, Amer. Mus. No. 8381, Cope Coll., from Republican River beds of Nebraska. Specimen figured on pl. CXL. PLATES. @xiih Miocenr Ruroceroses. Bones or APHELOPS MALACORHINUS Fias. 1-6. Aphelops malacorhinus Cope, 1878. Skeleton bones referred to this species by Professor Cope. Amer. Mus. Cope Coll., from Republican River beds, Nebraska. Fig. 1, 1a, 1b, pelvis, anterior, posterior and external views, one-fourth natural size; fig. 2, 2a, tibia, anterior and distal views, two-fifths nature; figs. 3-6, dorsal and a cervical vertebra, two-fifths nature. These specimens were presumably referred to this species on account of their being found associated with the type or some referred skull material of A. mala- corhinus; but the record of such association has been lost, and a search through the published references, field records and other possible sources of information has failed to recover any conclusive evidence upon it. The association of skull and skeletal parts in the related species A. ceratorhinus Douglass indicates how- ever that these bones are correctly referred to A. malacorhinus. (W.D.M.). WE aLv"Tel SHMOLMMAL FAL JO AMAMINS TvoIDO OH Sn PEATE CEH MiocENE RHINOCEROSES SKULL AND SKELETAL Parts oF APHELOPS ? MALACORHINUS Fics. 1-7. Remains of Rhinoceroses, part or all referable to Aphelops malacorhinus. Amer. Mus. Cope Coll. Republican River Bed, Kansas or Nebraska. Fig. 1. Humerus, anterior view. Fig. 2. Part of maxilla. Fig, 3. Aphelops malacorhinus, occipital view of skull. -Paratype, Amer. Mus. No. 8439, Cope Coll. Fig. 4, 4a. Cervical vertebra, lateral and inferior views. Fig. 5, 5a, 5b. Ulna, probably A. malacorhinus, external, anterior and distal views. No. 8382. Fig. 6, 6a. Proximal and distal ends of femur. Fig. 7, 7a, 7b. Astragalus, dorsal, plantar and distal views. PLATE CX MiocENr RHINOCEROSES. SKULL OF PERACERAS Peraceras superciliosus Cope, 1880. Side view of skull, one-half natural size. Type specimen, Amer. Mus. No. 8380. Republican River beds, Nebraska. Specimen figured on pl. CXLIVb. PLATE C rt PLATE CXLIVb. Miocene R#HINOCEROSES. SKULL OF PERACERAS ‘Peraceras superciliosus Cope, 1880. Top view of skull, two-fifths natural size. Type specimen, No. 8380, figured on pl. CXLIV. U.S. GEOLOGICAL SURVEY CO THE TERRITORIES PLATESGXEY. Miocene Equip. SKULL OF MrRYCHIPPUS Nomenclature Cope 1880-3 Revised 1915 Hippotherium speciosum (in part) Merychippus sp. Fras. 1-8. Merychippus sp. Skull, lateral, palatal and superior views, one-half natural size. Amer. Mus. No. 8347, Cope Coll., from Republican River beds, Driftwood Creek, Hitchcock Co., Nebraska. Pia. 4. Merychippus sp. Fourth premolar and first molar of No. 8347, natural size, crown view. PLATE CXLVIL Miocene Equip®. SKULL AND Bones or MERYCHIPPUS Nomenclature Cope 1880-3 Revised 1915 Hippotherium seversum (in part) Merychippus isonesus Fias. 1-8. Merychippus isonesus (Cope, 1889). Skull and parts of skeleton, figured also on pl. CXLVIII. All one-half natural size except fig. 16. Type specimen, Amer. Mus. No. 8175, Cope Coll. Mascall formation, Cottonwood Creek, John Day Valley, Oregon. Fig. 1, 1a, skull, lateral and palatal views; fig. 1b, two upper molars, m!~? of the right side, natural size; fig. 2, 2a, atlas, superior and posterior views; fig. 3, sacrum, inferior view; fig. 4, pelvis, inferior view; figs. 5, 6, proximal and distal views of femur; figs. 7, 8, proximal and distal views of tibia; fig. 9, inferior view of astragalus. PLATE -€xX iby alr Miocenrg Equip#®. SKELETON OF MERYCHIPPUS Nomenclature Cope 1880-3 Revised 1915 Hippotherium seversum (in part) Merychippus isonesus Hippotherium sphenodus Merychippus sphenodus Fies. 1-8. Merychippus isonesus Cope, 1889. Parts of skeleton with skull figured on pl. CXLVI1. One-half natural size. Type specimen, Amer. Mus. No. 8175, Cope Coll. Mas- call formation, Cottonwood Creek, John Day Valley, Oregon. Fig. 1. Pelvis, external view; fig. 2, 2a, femur, anterior and posterior views; fig. 3, 3a, tibia, anterior and posterior views; fig. 4, hind foot, dorsal view; fig. 5, 5a, internal and external views; figs. 6, 6a, navicular, proximal and distal views; fig. 7, 7a, ectocuneiform, proximal and distal views; fig. 8, metatarsals and phalanges, internal view; 8a, 8b, proximal and distal views of metatarsus. Fras. 9-10. Merychippus sphenodus Cope, 1889. Upper premolars, crown view, natural size. Type specimen, Amer. Mus. No. 8281, Cope Coll., Pawnee Creek beds, Pawnee Buttes, Colorado. EVER = Cc RIUM 7 J Ei © MMEs PLATE CXLVIIla. Miocenr Equip#. TrmnTH AND Brain Cast OF MERYCHIPPUS, ETC. Fic. Fic. HirGeame Fia. Fias. 5-7. Fig. 8, 8a. Fias. 9-11. Nomenclature Cope 1880-3 Revised 1915 Hippotherium sphenodus Merychippus sphenodus Protohippus (in part) Protohippus sp., aff. perditus Protohippus perditus (in part) Merychippus sejunctus Protohippus insignis (Cope, non Leidy) Merychippus labrosus ete. Protohippus sejunctus (in part) Merychippus isonesus Hippidium spectans Pliohippus spectans Merychippus ? sphenodus (Cope, 1889). Upper premolars; paratype, natural size, Amer. Mus. No. 828la, Cope Coll., Pawnee Creek beds, Colorado. ? Merychippus, unidentified specimen, upper molars. Merychippus sejunctus. Upper teeth, natural size, crown view. Amer. Mus. No. 8254, Cope Coll., recorded as ‘‘East of Laramie Peak, Wyoming.” Protohippus sp., aff. perditus. Upper teeth, crown view, natural size. Am. Mus. No. 8340, Cope Coll., Republican River Bed, Nebraska. Hind foot of this individual figured on pl. CXLIX. Type specimens of Protohippus labrosus Cope, Amer. Mus. No. 8266, Pawnee Creek Bed, Pawnee Buttes, Colorado. Fig. 5, upper teeth, crown view, natural size; fig. 6, 6a, lower jaw external and superior views, half natural size; fig. 7, upper jaw, half natural size, external view. Part of this specimen is now missing, only the lower jaw (fig. 6) being identifiable. This lower jaw is a composite of two individuals, the symphysis belonging certainly to a different individual and probably to a distinct species from the remainder of the jaw, which along with the upper teeth figured here (fig. 5) but now lost, agrees very nearly with Merychippus sejunctus. The symphysis, however, which in conformity with the author’s description and evident intent must be regarded as the primary basis of the species and selected as lectotype, is clearly distinct from M. sejunctus and cannot be correlated with any of the better known species. W.D.M., 1913. Merychippus isonesus, natural braincast, lateral and superior views. From the specimen figured on pl. CXLIX. Amer. Mus. No. 8105, Cope Coll., Deep River beds, Deep River (Smith Creek), Montana. Pliohippus spectans (Cope, 1880). Upper teeth, natural size, type specimen, Amer. Mus. No. 8183, Cope Coll., Maseall formation, Cottonwood Creek, John Day Basin, Oregon. Fig. 9, 9a, first or second, upper molar, crown and anterior views; fig. 10, second upper premolar crown view; fig. 11, upper incisor, crown view. I L-S NLIGHUHdI d ¢€ SNddIHOLOUd # ‘Z 1a SHIYMOLMYEAL HHL 10 AGASNS TVOIDO'TORD Sn PATE CXL Limp AND Foot Bones or MiocENnE EQuips® Nomenclature Cope 1880-3 Revised 1915 Protohippus sejunctus (in part) Merychippus isonesus Protohippus perditus Protohippus sp. aff. perditus Hippidium ? pernix ? Hipparion sp. Fics. 14. Merychippus ? isonesus (Cope, 1889). Limb and foot bones with skull. Amer. Mus. No. 8105, Cope Coll., Deep River beds, Deep River (Smith Creek), Montana. Fig. 1, la, ulno-radius, anterior and external views, one-half natural size; fig. 2, 2a, 2b, 2c, fore foot, anterior and external views, with proximal and distal views of metatarsus half natural size; fig. 3, proximal view of magnum and unciform with part of metacarpal III, two-thirds natural size; fig. 4, proximal view of tibia, one-half nature. See also pl. CXLVIIIa. Vic. 5-6. Protohippus sp. aff. perditus. Parts of skeleton associated with upper jaw figured on pl. CXLVITIa, Am. Mus. No. 8340, Cope Coll., from Republican River beds, of Nebraska. Fig. 5, pelvis, inferior view; fig. 6, 6a, 6b, anterior, internal, proximal views of median metatarsal; both one-half natural size. Fic. 7. Atlas, unidentified (not Equid). Fic. 8. ? Hipparion sp. Metatarsal III and proximal phalanx, anterior and external views and proximal view of metatarsal, one-half natural size. Amer. Mus. No. 8357, Cope Coll. Republican R. beds, Nebraska. (The specimen includes remains of more than one individual and no teeth are associated, so that the reference is wholly doubtful. It has the proportions of the American species of Hipparion but is exceptional among them in the tarso-metatarsal articulation). i : § i ‘} = ges SSE Ne oy ’ ‘ ’ A an fs . a % ¢ N * p - ’ iS : a ~ ra : PLATE CL: Miocenre Equipa. SKULL OF MERYCHIPPUS Nomenclature Cope 1880-8 Revised 1915 Protohippus sejunctus Merychippus sejunctus Fries. 1-3. Merychippus sejunctus (Cope, 1874). Skull of type specimen, lateral, palatal and 3 superior views, one-half natural size. Amer. Mus. No. 8291, Cope Coll., Pawnee Creek beds, near Pawnee Buttes, Weld Co., Colorado. Skeleton parts figured on pl. CLI. Ay PEA TE -Ciek Miocene Equip®. SKELETON Parts oF MrERYCHIPPUS Fias. 1-14. Merychippus sejunctus Cope, 1874. Skeletal parts of type specimen, skull figured on pl. CL. All one-half natural size, Amer. Mus. No. 8291, Pawnee Creek beds, near Pawnee Buttes, Colorado. Fig. 1, lower jaw, superior view; fig. 2, 2a, atlas, superior and inferior views; fig. odontoid 2b, process of axis; fig. 3, 3a, third cervical vertebra, inferior and posterior views; fig. 4, 4a, seventh cervical, anterior and inferior views; fig. 5, 5a, first dorsal, anterior and lateral views; 6, 6a, 6b, 6c, humerus, anterior, external, proximal and distal views; 7, 7a, 7b, tibia, anterior, external and distal views; figs. 8-14, hind foot, anterior and external views, 8b, inferior view of astragalus, 8c, 9b, of caleaneum and astragalus, 10b, proximal view of navicular and cuboid, 11b, 11c, proximal and distal views of metatarsus, 12b, proximal view of proximal phalanx; 13b, of second phalanx, 14b, 14c, proximal and inferior views of ungual phalanx. The humerus and tibia here figured were found in the Cope Collection with No. 8251, a specimen which Cope referred to Protohippus perditus, and which is now (1913) referred to Merychippus ? sphenodus. It appears probable that parts of the type skeleton of No. 8291 had become mixed with No. 8251, but the two individuals are §0 closely allied specifically that I am unable to clear up the confusion, et Sb fe en, Post Als PROTO HI o . PEATE. CLI MiocENE CAMELID Nomenclature Cope 1880-3 Revised 1915 Protolabis transmontanus Miolabis transmontanus Protolabis heterodontus Protolabis heterodontus Fics. 1, 2. Miolabis transmontanus (Cope, 1879). Skull, lateral and palatal views, two-thirds natural size. Type specimen, Amer. Mus. No. 8196, Cope Coll., from Mascall formation of Cottonwood Creek, John Day Valley, Oregon. Fias. 3, 4. Prololabis heterodontus (Cope, 1874). Upper and lower jaws, side’ view, and palatal view of upper jaw, two-thirds natural size. Type specimen, Amer. Mus. No. 8296, Cope Coll., from Pawnee Creek beds, near Pawnee Buttes, Weld Co., Colorado. SOY Snr SSS; I2, PROTALABIS SIREN, PLATE Clie SKULL AND Jaws oF Miocene CAMELS Nomenclature Cope 1880-3 Revised 1915 Procamelus angustidens Cope 1872 = Protolabis angustidens Cope “Procamelus angustidens” Cope 1881 = Procamelus inzequidens sp. nov. Fic. 1. Procamelus inequidens sp.nov. Side view of skull and jaws, No. 8446, type, from Upper Miocene, Republican R. beds, Driftwood Creek, Hitchcock Co., Nebraska. This skull was referred by Cope to P. angustidens, but differs from topotypes of that species in the absence of the upper incisors and other characters of less moment. Fias. 2, 3. Inferior and superior views of same skull. Fia. 4. Protolabis angustidens Cope, 1872. Lower jaws, type specimen, superior view. From Middle Miocene (Pawnee Creek beds) near Pawnee Buttes, Colorado. Cope Coll., Am. Mus. No. 8294. PROCAMELUS ANGUSTIDEN, jOPE. + . ; 5 rer . F 1 7 ! + ¥ ' ‘ r 1 ob i vet v L! 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