Hoyas in the Wild This is a slide from March 1986. Near village of Ghara above Tatopangi Central Nepal, 5800' elevation. Tree overhanging Kali Ghandaki River gorge. Photographer unknown Oh There it is ! 1 A pdf publication devoted to the Genus Hoya ISSN 2329-7336 Editor: Dale Kloppenburg Volume I Issue 2 December 2013 Contents This second issue will be devoted to the publication of some more new hoya species from the Philippines and one from Samoa. Contributors: Ulysses Ferrer as, botanist, plant collector, plant taxonomist. Dale Kloppenburg retired: Graduate of UC Berkeley, plant genetics, Lt. USNR, Plant Breeder and Research Agronomist, now taxonomist of Genus Hoya. George Mendoza, Landscape Artist, Nurseryman and Plant Explorer from Quezon City, Philippines, visit his web site www.forest-treasures.com. Simeona V. Siar former University Former Researcher -Crop Science Cluster, Institute of Plant Breeding. College of Agriculture, University of the Philippines. Now deceased. The following new species are presented in PDF format with ISSN number. 1. Hoya apoensis subsp. sagittaria Kloppenburg, Siar & Ferreras 2. Hoya davaoensis Kloppenburg 3. Hoya pubicorolla Kloppenburg, Mendoza & Ferreras 4. Hoya pubicorolla subsp. anthracina Klopp., Mendoza & Ferreras 5. Hoya tiatuilaensis Kloppenburg 6. Hoya taytayensis Kloppenburg & Siar NOTE: please see the Website publication of these species at ‘Vww.rare- hoyas.com”. Go to end and click on “publication” to access my HOYA NEW publications. 2 Hoya apoensis subsp. sagittaria Kloppenburg, Siar & Ferreras ISSN 2329-7336 Specimen collected by Ulysses Ferreras on Mindanao, South Cotabato Province, Tampakan Municipality, near Sagittarius Mining Inc, Base Camp. N 06° 28’29.4” E. 125° 03 ’29.0”. Altitude 1.395 mas! Habitat: found growing in a low montane residual forest at the top of a hill. Forest patch less than half a hectare with understory planted with coffee. Intermediate adjacent area already converted to cornfield. Collection number UF 737. Via Dr. Simeona Siar in small poly bag, 1/22/10. 4 flowers on pedicels. Hoya apoensis subsp. sagittaria Kloppenburg, Siar & Ferreras subspecies nova, Holotypus 71722 (CAHUP) hie designatus. Section Acanthostemma (Blume) Kloppenburg, similis Hoya apoensis Kloppenburg & Siar sed differt in parvis modo. Corollae color roseus non citreus; flos parvior 1.08 cm diametro applanatus contrstre 1.26 cm; coronae lobus brevior 0.18 cm contrstre 0.22 cm, differt. This new subspecies is similar to Hoya apoensis but slightly different, the color of the corolla is rose red not bright yellow. The corolla flattened is slightly smaller, the coronal lobes apex - apex are shorter and there are some minor differences in the pollinarium. Photomicrographs of flower parts with measurements: Pedicel: slightly curved, 1.5 cm long, terete, glabrous, 0.10 cm in diameter, slightly bulbous at the calyx. Enlarged about 25X. Picture enlarge about IIX, bottom of calyx bulbous. 3 Calyx: small triangular sepals, triangular to sub-obtuse at apex. 0.11 em long, 0.09 em widest at base, glabrous no overlap, ligules. Enlarged about 12X. Ovaries: dome shape tapering to apex, glabrous, 0.15 em long, 0.10 em wide at the base. Outside view of the flower showing pedieel and ealyx attaehed. Corolla: rotate ends reeurved to revolute, glabrous outside, sliek, inside pubeseent with prominent eells pointing eentrally, apex inside glabrous, inside at eentral area slight pentagonal distinetion with slight less puberulanee. Enlarged about 12X. Sinus - sinus 0.30 em Sinus - eenter 0.26 em Sinus - apex 0.39 em Apex - eenter 0.54 em Widest 0.35 em Above: photo outside surfaee of the eorolla. Eeft: inside surfaee of the eorolla enlarged ea. 12X. 4 Corona: bilobed, inner lobe narrow spatulate, outer lobe obtuse, bilobes flat near lobes and bending inward and touehing at outer apex with rounded ends, suleate. Channeled below narrow with large open eenter and thiekening of short eentral eolumn, hair eells Enlarged about 13X. around eolumn Apex - apex 0.18 em Apex - eenter 0.20 em Center - bilobes 0.23 em Bilobe extensions 0.06 em Ret. - ret. 0.06 em Ret. - eenter 0.06 em Aw. - aw. 0.16 em Aw. - eenter 0.15 em 5 Leaves glabrous, pinnate nervation ea. 38° to midrib, euneate base, elliptie, apex aeute to sub apieulate. Leaf measurements at the end taken from herbarium sheet by Dr. Simeona Siar. 6 Pollinarium enlarged ea. 80x. Pollinium length 0.35 mm widest 0.12 mm Retinaculum length 0.15 mm shoulder 0.05 mm waist 0.04 mm hip 0.06 mm ext. 0.03 mm Translator length 0.15 mm depth 0.02 mm Caudicle bulb diam. 0.05 mm Translator/caudicle type: fb/ew 7 X. CAHUP 71722 (Philippines UF 737) Leaf sample Length (cm) Width (cm) 1 7.60 2.70 2 7.60 2.60 3 9.30 2.40 4 9.60 2.50 5 8.10 3.20 6 7.30 2.30 7 8.90 2.80 8 7.10 2.00 9 6.30 2.50 10 5.90 1.80 Mean 7.77 2.48 Range 5 . 90 - 9.60 1 . 80 - 3.20 8 Hoya davaoensis Kloppenburg ISSN 2329-7336 Hoya davaoensis Kloppenburg sp. nova, holoypus 49272 (UC) hie designatus, similis Hoya pubicalyx Merrill, Philippine Journal of SeieneelS, C5:1918, 131, eum foliis similis sed petioli 2X longiores; pedieello brevior 1.67 em eontrastre 1.80 em; eorona parvior 0.72 em diam. eontrastre 1.00-1.20 em This new speeies has foliage similar to Hoya pubicalyx Merrill but the petioles are two times longer and the pedieels are shorter 1.67 em vs. 1.80 em, and the eorona is smaller in diameter 0.72 em vs. 1.00-1.20 em Measurements and observations from the UC Type sheet: Stems: very thiek 0.5 em in diameter, nodes enlarged 1.0 -1.2 em in diameter 17 em long. Blade: oval 8-11 em long x 5 em wide, apex short apieulate rounded, base rounded to sub aeute, glabrous above and below, obseure pinnate nerves, margins reeurved. Petiole: striet 2 em long, ehanneled above, glabrous. Peduncle: 3 em long, glabrous, striet, 0.1 em in diameter. Raehis enlarged 3-4 em long. Pedicel: strict, filiform, glabrous, 2.0 cm long. 12 open, 15 unopened buds, not open all at once. View of the pedicel, calyx and corolla outside. Pedicel is glabrous strict, terete. Sepals: ovate, 0.37 long and 0.27 at the widest, edge entire overlapped ca. 1/2. Corolla: outside glabrous, inside papillose. Strong collar 0.07 cm tall and 0.14 cm at opening, (had been eaten by bupestids). Sinus - sinus 0.57 cm Sinus - apex 0.55 cm Apex - center 0.83 cm 9 Inside surface of the corona, inner lobe almost touch in center, dorsal nearly horizontal rising slightly to outer apex, cupped with umbo near outer rounded apex and reflexed. Surface glabrous, sulcate. Scales are channeled below. Inner apex dentate. Apex - apex 0.35 cm Apex - center 0.36 cm Widest 0.20 cm Pollinarium Pollinium length 0.80 mm widest 0.25 mm Retinaculum length 0.20 mm shoulder 0.15 mm waist 0.10 mm hip 0.12 mm ext. 0.03 mm Translators length 0.15 mm depth 0.04 mm Caudicle bulb diam. Translator/caudicle type: d/o 10 Collected by Ramos & Edano at Mati, Davao, Mindanao, Philippines, March -April 1927, UC #49272 also at (B) Flower red salmon color in open places in forest stream, low altitude. This is not Hoya meliflua Merrill. 11 Sheet at (B) with same number labeled Hoya r I ,-A i.»j Of. THf t>-Ui WH f I )#, jf ri ■’ y -5 ji ■ 12 Hoya pubicorolla Kloppenburg, Mendoza & Ferreras ISSN #2329-7336 Hoya pubicorolla Kloppenburg, Mendoza & Fereras sp. nov. Holotypus Edano #3235 (PNH) hie designatus. Frutex scandens, inflorescentiis exceptis glaber, ramis teretibus, circiter 0.3 mm diametro; foliis carnosis, ellipticus, viilgo ad 13.8 cm longis et 4.5 cm latus, pallidis, nitidis, basicuneatus, apice acuminatis, pinnatinerviis, nervis utrinque circiter 6, utrinque valde obscuris; petiola ca. 1.2 cm longis; pedicellis tenuibus circiter 3.3 cm longLis leviter pubescentibus; floribus rotatis, insiccitate brunneis, circiter 2.2 cm diametro complinatus; sepalis ovatis, subacutis, extus leviter pubescentibus, 0.48 cm longis; corollae lobis late triangularibus, acuminatis, circiter 0.64 cm longis, indus dense pubescentibus, apice leviter reflexis, glabrous; coronae lobis coriaceis, oblongo-ovatis, 0.56 cm longis, acuminatis vel acutis, patentibus, apice leviter emarginatus. A scandent shrub, entirely glabrous except the inflorescence, the branches terete, rather pale when dry, about 0.3 cm in diameter. Leaves opposite, fleshy, elliptic generally 13.8 cm long, 4.5 cm wide, base cuneate, apex distincth' acuminate; lateral nerves pinnately arranged, indistinct, about 6-8 on each side of the midrib, ascending, scarcely more evident than are the lax, indistinct reticulations; petioles about 1.5 cm long, the pedicels spreading, slender, about 3.3 cm long, sparingly appressed-pubescent. Cal\Tc-segments ovate, subacute, externally somewhat pubescent, about 0.48 cm long. Corolla-lobes broadly triangular, about 0.64 cm long, spreading, rotate, externally glabrous, internally densely pubescent the apex somewhat recurved and glabrous. Corona lobes spreading, coriaceous, oblong-ovate to ovate-lanceolate, acuminate, 0.56 cm long, the external tip emarginate, the internal one ascending, the upper surface plano-convex and somewhat keeled in the median portion. The anthers greatly exceed the inner coronal lobes. This species for years has been misidentified as Hoya pnbicalyx Merrill and widely distributed under this name in the trade. This new species leaves are elliptic to ob- eli iptic, with a cuneate base not oblong to oblong-obtuse with an obtuse base; average mature leaf is ca. 15.0 cm long and 4.3 cm at the widest the corolla inner surface of this (new) species is densely pubescent not papillose and much larger, lobes here 0.86 cm long versus 0.60 cm long, flattened this new species is 2.20 cm in diameter; the corona lobes just exceed the corolla sinuses with a diameter of the corona of 1.26 cm Note: most of the clones of this species, but not all, have silvering on the leaf upper blade often with some pinkish tones added in. The petiole is grooved above, slightly curved, often corky, 1.5 cm long, 0.4 cm in diameter the sap is also clear not noted in the Hoya pnbicalyx Merrill type description. Some background on the Philippine Herbaria : (PNH) the Philippine National Herbarium at Manila was destroyed at the end on world wai* 11, The original Type sheet ‘‘Luzon, Tayabas Province, Mauban, Bur. Sci. 19484 Ramos & Edano, January^ 24, 1913, on trees along streams in damp forests” was destroyed. All subsequent new herbarium sheets bearing the name Hoya pnbicalyx are misidentified in my opinion, the first of these sheets 13 were added in (PNH) by Edano #3235 1948. I have been unable to loeate any leetotypes of this speeies in any herbaria. Next (PUH) herbarium at Quezon City was also destroyed in World War 11 it was reestablished in 1946 about the same time as the (PNH) Herbarium. Definitions: Papillatus = having papillae, nipple-like surfaee. Pubescens = hairy as opposed to glabrous. Puberulous = minutely pubeseent, downy with soft short hairs. Photomicrographs and measurements of floral parts follow: Pedieel enlarged about 8X. Surfaee with seattered pubeseenee, terete straight. This is from a non-silvered leaf form sent from the Philippines by Dexter Heusehkel. Pedieels appear the same but the sepal lobes differ from those “Red Buttons” whieh has long narrow sepals. Pedicel: 3.3 cm long x 0.13 cm in diameter, surface reddish with whitish lenticels. Calyx enlarged about 8X, same as above. Outside pubescent, ciliate; inside glabrous, overlap about 1/3. Calyx: apex reaches the corolla sinus. Sepals 0.48 cm long and 0.18 cm at the widest. Diameter about 1.0 cm flattened. 14 Calyx enlarged about 8X. Inside glabrous, bases of sepals overlap, ligules present. Ligules in this speeies, variants are very variable, from long slender exeeeding the sinuses to short as here and in this ease with 2 missing. Edges eiliate apiees narrowly obtuse. Ovaries: dome shaped with separation on the inside surfaees near the apex. 0.25 em tall and base pair 0.20 em Ventral view of eorolla lobe enlarged about 8X. Surfaee is glabrous, almost erystalline as is the inner surfaee at the apex, whieh here turned under as are the lobe edges. Sinus - sinus 0.55 em Sinus - eenter 0.46 em Sinus - apex 1 . 1 0 em Apex - eenter 1 .32 em Dorsal (top) view of a eorolla lobe and part of the eorona enlarged about 8X. Inside surfaee very dense pubeseenee. Coronal lobes exeeed eorolla sinuses. Ventral view of the eorona enlarged about 8X, seales are ehanneled and edges very finely suleate; with long under eurve below anther wings. Central eolumn thiekened. 0.10 em tall. 15 Top view of the eorona enlarged about 8X. Inner lobes do not reaeh the eenter where the long anthers rise above the eenter, whieh are 0.13 em long. Inner lobe keeled on top and spatulate. Seales broaden outwardly, then tapering to an emarginate apex, dorsal surfaee slightly sunken and with sharp edges above and slightly raised down the eenter. Apex - apex 0.56 em Apex - eenter 0.63 em Widest 0.33 em Ret. - ret. 0.14 em Aw. - aw. 0.35 em Aw. - ret. 0.17 em View of the pentagonal stylar erown with most eoronal seales intaet enlarged about 8X. The stylar apex is raised, somewhat eolumnar with a mealy texture. 16 Pollinarium enlarged about 165X. Measurements for this elone Pollinia length 0.72 mm widest 0.23 mm Retinaculum length 0.27 mm shoulder 0.22 mm waist 0.09 mm hips 0.10 mm extensions 0.08 mm Translators length 0.15 mm depth 0.02 mm Caudicle bulb diam. 0.1 1 mm Translator/caudicle type: Is/o Leaves: opposite, glabrous, petiolate, lateral nerves pinnately arranged, indistinet, elliptie, base euneate, apex aeute, dorsal surfaee often blotehed (silvered) but some elones not. Leaf measurements: in eentimeters. length width petiole length 1) 15.0 4.3 1.8 2) 13.0 3.8 2.4 3) 12.7 4.3 2.6 4) 11.5 3.9 2.1 5) 8.7 3.3 1.8 Range: 00 o 3.3- 4.3 1.8-2.6 17 A typical plant of this species photo by Ann Wayman in Central Point Oregon, USA. Previously thought to be Hoya pubicalyx Merrill. Below a typieal Hoya pubicorolla flowering plant below blooming in Fresno, California, USA. Misidentified as Hoya pubicalyx Merrill. 18 A list of herbarium sheets labeled as Hoya pubicalyx Merrill, all ineorreetly labeled as near as I ean determine. Herbarium Sheets: Edano #3235 1948 tPNHh Kondo and Edano #36884 1948 tPNHh #36790 1957 tPNHh Britton #19486 1953 tENHE Mandoza #97463 1967 tENHE Fox # 9226 1949 tENHE ? #13972 1951 tENHE Velaseo #9135 1954 tCAHUPE #9136 1954 tCAHUPE Kienholtz #15460 1924 rUQ. 19 Hoya pubicorolla subsp. anthracina Kloppenburg, Ferreras & Mendoza ISSN #2329-7336 Hoya pubicorolla subsp. anthracina Kloppenburg, Ferreras & Mendoza subsp. nova holotypus 14597 (PUH) hie designatus. Similis speeie, sed pedieilis longior 3.8 em eontrstre 3.3 em; sapalis longior 0.20 em eontrstre 0.18 em et eorollae lobis brevior 0.55 em longis eontrstre 0.77 em; similes Hoyas pubiealyx Merrill, Philippine Journal of Seienee 13 (1908) 221. sed eorolla intus non dense papillosis sed dense pubeseentibus, differt. This new subspeeies differs from the speeies in that the pedieels of this sub- speeies are longer 3.8 em eontrstre 3.3 em and the sepals are longer 0.20 em versus 0.18 em and the eorolla lobes are shorter 0.55 em long versus 0.77 em, also similar to. H. pubiealyx but the inner (dorsal) eorolla surfaee here is very pubeseent not densely papillose, among other differenees. Photos of flower parts and details: Pedicel: section enlarged ca. 27X, strict, terete, glabrous, 3.8 cm long and 0.12 cm in diameter. Calyx: side view enlarged ca. 15X, the base is pubescent, as is the sepal outer surfaces, also ciliate. Calyx top view enlarged ca. 12X, the inner surface is slick glabrous, sepals 0.20 cm long and 0.20 cm at the widest, ligules are at the sinuses 1-11-2-2 pattern Ovaries: are columnar, glabrous, 0.24 cm tall and base pair 0. 15 cm wide. 20 Corolla: ventral lobe area enlarged ea. 13X the surfaee is glabrous, sinuses eonduplieate, apex and edges turn under and apex on dorsal side is also glabrous. Sinus - sinus 0.55 em Sinus - eenter 0.50 em Sinus - apex 0.65 em Apex - eenter 1.05 em Widest 0.70 em Corolla dorsal surfaee, here pubeseent; diminishing in eenter of lobes and also nearer the eenter, here there are stellate hair eells pointing inward toward the eenter. Corona: ventral surfaee enlarged ea. 8X, the lobes are ehanneled with edges round and rolled under, the surfaee is smooth only under high magnifieation are slightly suleations visible, eentral eolumn 0.07 em tall, surrounding at base elear but granulate. Corona dorsal surfaee enlarged as above. Again glabrous surfaees Inner lobe is raised, sub terete with anthers greatly exeeeding by 0.07 em, dorsal eonvex exeept along the side shelves whieh extend about half way toward the outer apex shelves sharp. Apex - apex Apex - eenter Widest Ret. - ret. Ret. - eenter Aw. - aw. Aw. - eenter that is sub aeute, edges of 0.52 em 0.59 em 0.29 em 0.12 em 0.15 em 0.32 em 0.32 em 21 Corona scale side view enlarged ca. 9X, surfaces are smooth, glabrous, anthers greatly exceeding inner lobe, outer apex end slightly rolled down but sub acute. Pollinarium enlarged ca. 180X. Pollinium length 0.83 mm widest 0.20 mm Retinaculum length 0.26 mm shoulder 0.20 mm waist 0.10 mm hip 0.16 mm ext. 0.05 mm Translator length 0.10 mm depth 0.03 mm Caudicle bulb diam. 0.08 mm Translator/Caudicle Type: d/o Ratios: p/w 4.2 p/r 5.2 Flowers in globose clusters of ea. 28 flowers. 22 Photos of flowers of this species sent by George Mendoza Vile #14 DSC 2391 2392 2395 GM 23 Notations: H. pubicalyx pedicels pubescent 3.5 cm long. Merrill said inside corolla densely papillose this one is definitely pubescent ! ! ! Inner lobes not as long and are not terete but dentate. Corona lobe dorsal more keeled. Coronal lobes a little longer 0.55 cm versus 0.50 cm (Corona smaller on this one). 24 Hoya tiatuilaensis Kloppenburg ISSN 2329-7336 Hoya tiatuilaensis Kloppenburg sp. nova Type 9089 HAW hie designatus similis Hoya samoensis Seemann sed partes omnes parvi, folia 5.5 -6.5 em x 3 - 3.5 em eontrstre 7 - 10 em x 3.5 - 5.8 em; peduneuli 4.5 em longi eontrstre 5.5 - 6.5 em; eoronae lobis 0.20 em longi x 0.10 em, lati eontrstre 0.35 - 0.40 em x 0.15 - 0.20 em, differt. Description of the Herbarium sheet 9089 HAW: 9089 as Hoya pottsii Traill, 5 Jan, 1993 Tiatuila, Samoa. Vine with eream eolored flowers, sap not notieeably milky, oeeasional, but only one seen in flower, in the ridge forest on Tiataula ridge east of Vatia. Elevation 180 m. 2 stems 5 leaves, 1 flower globose eluster of ea. 34 flowers. 1 stem with adventitious roots all along. Leaves glabrous, broadly ovate-laneeolate, shortly attenuate, apex aeute, base rounded but not eordate, 5.5 - 6.5 em long 3 - 3.5 em widest nearer the base, nervation quintuplinervis 5 distinet on both surfaees, same eolor as the leaf, seeondary nerves some looping near the margins, primary nerves extend to near the apex. Petiole glabrous, not thin 1.5 em long probably grooved above, same eolor as the stem, eireular gland at the attaehment above. Internodes glabrous, mostly 8 em long, terete, 0.2 em in diameter, roots up to 3 em long, fine, nodes not mueh enlarged. Pedunele glabrous, straight, 4.5 em long, terete raehis a little longer, faseieles. Pedieels. glabrous, very fine, terete, 34 flowers. 2 em long, light eolored. Note: not H. pottsii, leaf quintuplionervis not palmate and base not eordate. Flower relatively small. Calyx large, reaehes sinus of eorolla. Micro Photos of flower from sheet 9089: Outside view of the flower enlarged about 8X. Pedieel is glabrous, terete, short, 1.05 em long 0.05 em in diameter. Calyx is very large, outside granulose, inside smooth, sepal apex reaehes the eorolla sinuses. Flower not yet opened. 25 Calyx: Left outside view, right inside view enlarged about 8X., glabrous. Sepals 0.13 em long, 0.13 em at the widest, near the base. One ligule seen. Outside view of the eorolla enlarged about 8X. This surfaee is glabrous, eentral ring raised (eonvex). widest out from the sinus area. Sinus - sinus 0.25 em Sinus - eenter 0.21 em Sinus - apex 0.38 em Widest 0.33 em Apex - eenter 0.55 em So the flower diameter flattened is 1.10 em Lobes Inside view of the flower enlarged about 8X. Surfaee is finely and evenly pubeseent. Coronal apex is aeute. Inner lobe long apieulate, dentate, do not reaeh the eenter. Inner lobe ean be seen as yellow linear line near the eenter Top view of the eorona enlarged about 8X. Outer apexes seem to be emarginate. Lower surfaee ehanneled. Apex - apex Apex - eenter Widest Anther wing - aw. Retinaeulum - ret. Anther wings are very bulbous Aw. - eenter Ret. - eenter 0.20 em 0.23 em 0.10 em near the inner lobe. 0.15 em 0.08 em 0.13 em 0.05 em 26 Several retinaeula enlarged about 82X. Head is long with shoulders well down, waist only slightly narrower, and hips a little extended. length 0.20 mm shoulder 0.13 mm waist 0.09 mm hip 0.10 mm extensions 0.08 mm Redueed pieture of the type sheet, to the right. Pollinia here still in the anther saek enlarged about 82 X. The length is ea. 0.55 mm long and 0.20 mm widest. 27 Hoya taytayensis Kloppenburg & Siar Hoya taytayensis Kloppenburg & Siar sp. nova, typus 9385 (BO) hie designatus. Similis Hoya crassicaulis Elmer ex Kloppenburg Fraterna 8/3:1995,10 sed differt, eoronae lobi extus obtusus non aeutus; folia parvior sed similis, pollinium multo longior 0.70 mm eontrastre 0.50 mm. This new speeies is similar to Hoya crassicaulis but the outer lobes of the eorona are obtuse not aeute, the foliage is similar but smaller and the pollinium are longer among other differenees. The speeies was eolleeted by E. D. Merrill at Taytay, Palawan, Philippines in 1913. Worked 11 Nov. 1993. Stems: heavy 0.4 em + in diameter, nodes larger, internodes 0.5- 1.0 em long, glabrous. Leaf blade: ovate to ovate elliptie, base obtuse, apex sub aeute, 6-9 em long and 3.5-6 em wide, glabrous on both surfaees, nerves pinnate 4 pair, edges rolled under. Petiole: 0.25 em long dark eolor, glabrous. Peduncle: short 1.5 em long. Rachis: Broad rounded. Eeft: inside view of the ealyx, ovaries dark near eenter. Right: bottom view of the ealyx. Pedicel: 1-1.5 cm long, straight, glabrous. Calyx: sepals are 0.20 cm long and 0.17 cm wide at the base. Ovate with a few cilia on edges, overlap about 1/3 at base. Ovaries: 0.13 cm tall and 0.09 cm wide at base pair. Very short domed, glabrous. 28 Corona: saddle shaped, outer apex obtuse, inner lobe raised with ends turned in on drying and do not reaeh the eenter. Dorsal with eentral ridge and sunken in eentral portion. Channeled below View of the flower enlarged about 8X. Corolla outside glabrous, inside pubeseent. Apex inside glabrous. Sinus - sinus Sinus - eenter Sinus - apex Apex - eenter Widest 0.50 em 0.35 em 0.39 em 0.60 em 0.45 em Apex - apex Apex - eenter Widest Ret - ret. Aw. - aw. 0.18 em 0.23 em 0.16 em 0.10 em 0.15 em I had a diffieult finding any pollinaria sinee I had only one flower from the herbarium sheet. I managed to reeover 2 retinaeula and a few separated pollinia. Below is the photo. Pollinium length 0.70 mm widest 0.23 mm Retinaculum length 0.21 mm shoulder 0.15 mm waist 0.10 mm hip 0.13 mm ext. 0.02 mm 29 I I I I Poor scan of the herbarium sheet (BO) # 9385 collected by E. D. Merrill at Taytay, Palawan in May 1913. About ^2 real size. 30