Broad tree lined avenues 1981, Picture by Dale Kloppenburg Bogor Botanical Garden, Indonesia What you may find when your out hunting for hoyas, yes and we found hoyas. A pdf publication devoted to the Genus Hoya ISSN 2329-7336 1 Volume 4 Issue 1 January 2015 Editor: Dale Kloppenburg Contents When a species is collected from the wild, I feel it is wise to identify it, propagate it and name it. In this way it will eventually get it into commercial channels, be distributed to all those interested in this genus and thus be preserved. If in the future the species is lost through natural causes or forest destruction it will still be here on earth in your collection. The following new species are presented in PDF format with ISSN number. 1 . Hoya histora Kloppenburg 2. Hoya palawanensis subsp. minora Kloppenburg 3. Hoya velasioii sbubp. grandiora Kloppenburg 4. Hoya columna Kloppenburg 5. Hoya maximowayettii Kloppenburg 6. Hoya wibergiae subsp. alba Kloppenburg 7. Hoya vicencioana subsp. quezonensis Kloppenburg 8. Hoya velasioii Kloppenburg 9. Hoya capotoanensis Kloppenburg 10. Hoya galeraensis Kloppenburg 11. Hoya sulitii Kloppenburg NOTE: please see the Website publication of these species at ‘‘www.rare- hoyas.com”. Go to end and click on “publication” to access new species publications. 2 Hoya histora Kloppenburg ISSN 2329-7336 Hoya histora Kloppenburg sp. nova, holotypus 5292 (CAHUP) hie designatus non similes ulluis Philippina hoya sp. folio pinnatinervis, eoronae lobus apex - apex 0.25 em longus, eorolla diametro eomplanatus 0.90 em et pollinarium translatoribus typus Is/o, differo (id est) eeterus. This speeies has an unusual eombination of eharaeters that distinguishes it from other Philippine hoya speeies in that the foliage is pinnately nerved, lobes of the eorona from apex - apex are 0.25 em long, the eorolla diameter flattened is 0.90 em and the pollinarium translators are Is/o type (in addition the pollinaria is 0.40 mm long). Named for the Natural History Museum at Univ. Philippines at Los Banos. Pedicel: enlarged about 16x. Surfaee is slightly pubeseent (a few short stiff hair eells). 1.5 em long, terete, 0.04 em in diameter. There was no ealyx on the seleeted flower from the herbarium sheet. Outside surfaee of the eorolla enlarged about 16x, glabrous but granulate. Sinus - sinus 0. 17 em Sinus - eenter 0.15 em Sinus - apex 0.32 em Apex - eenter 0.45 em Widest 0.25 em Inside view of the flower, the eorolla surfaee appears to be glabrous but granulate. Corona lobes are horizontal with eenter raised slightly, inner lobes are short spatulate and nearly toueh in the eenter, retinaeula and anther wings were eovered, dorsal a little eoneave with a linear low keel and a boss near the inner lobe base. Outer lobes are narrowly rounded and exeeed the eorolla sinuses. 3 Top view of the eorona enlarged about 16x. See keel and boss on dorsal surfaee of the eoronal lobe. Apex - apex 0.25 em Apex - eenter 0.27 em Widest 0.16 em Ret. - ret 0.06 em Aw. - aw. 0.13 em Bottom view of the eorona enlarged about 16x. Lobes are ehanneled from outer apex to about even with the sinuses, eentral eolumn appears short. Surfaees of the lobe forming ehannel are diagonally suleate and dorsal surfaee is longitudinally suleate. Side view of one seale greatly enlarged, aetually the lobes are horizontal with the eenter raised a little here on drying slightly the outer lobe is turning up. Inner spatulate lobe diffieult to diseern. 4 Above pollinarium greatly enlarged. See seale for measurements. Blaek arrow head is 0.1 mm long base 0.05 mm wide; main markings on stem also 0.5 em intervals . Pollinarium length 0.40 mm widest 0.15 mm Retinaculum length 0.19 mm shoulder 0.06 mm waist 0.04 mm hip 0.08 mm ext. 0.02 mm 5 Translator length 0.15 mm depth 0.02 mm Caudicle bulb diam. 0.07 mm Translator/caudicle type: Is/o Pollinia inner end type: R Measurements of herbarium sheet below: Stems glabrous, terete ea. 0.27 em in diameter, internodes 8.9 - 10.7 em long, nodes enlarged. Petiole: +/- 1.8 em long, glabrous eurved, possibly grooved above. Leaf: opposite, petiolate, ovate, shortly aeute apieulate, glabrous, 8.9 - 10.7 em long x 3.6 - 5.4 cm, widest near the middle. Pinnate nervation. Peduncle: appears short, flower in globose cluster. 6 Reduced Copy of the Type Sheet CAHUP #5292 7 Hoya palawanensis subsp. minora Kloppenburg ISSN 2329-7336 Hoya palawanensis subsp. minora Kloppenburg, holotypus CAHUP #5297 hie designatus, similes sp, sed sepala non triangularis, et 0.13 em x 0.10 em eontrastre 0.25 em X 0.15 em et apex sub-rotundus et ovaria brevior; eorolla diametro eomplanatus 1.10 em eontrastre 1.56 em, aliter similes. This subspeeies is similar to the speeies namely in the pollinaria and anther wing type but in most parts a little smaller, the sepals have rounded apiees not triangular in shape and are o.l3 x 0.10 em versus 0.25 x 0.15 em and the eorolla diameter flattened is 1.10 em versus 1.56 em mueh smaller otherwise the parts are similar. Details follow: Pedieel enlarged about 16x. Curved, terete, glabrous 1.8 em long, 0.04 em in diameter. Outside of flower enlarged about 8x. Sepals do not reaeh the eorolla sinuses, here dried and flared out from eorolla surfaee whieh is glabrous. 8 Pedicel calyx and ovaries side view enlarged about 8x. Ovaries are narrow domed shape, 0.11 cm tall and 0.08 cm wide at abase pair, glabrous. Top view of the calyx and ovaries enlarged about 8x. Sepals are glabrous inside and out, 0.13 cm long and 0.10 cm at the widest, overlap at the base about 1/4. There are faint ligules, sepal apex narrowly rounded Corolla outer surface, glabrous, cut more than !/2 way, central collar thickened and convex. Sinus - sinus 0.29 cm Sinus - apex 0.37 cm Sinus - center 0.22 cm Apex - center 0.55 cm Widest 0.33 cm Inside surface of the corolla enlarged about 8x. This surface is finely pubescent, lobe apex is acute and broadest in the middle. 9 Inside view of the flower enlarged about 8x. Coronal outer lobe exeeeds the eorolla sinus by a eonsiderable amount, is narrowly rounded an elliptie shape, dorsal a little eoneave with a raised keep from apex to apex. Inner lobe is blunt dentate and does not reaeh the eenter. Anther wings at apex are rounded and protrude a little. Lower surfaee of the eorona, glabrous, ehanneled all the way to the eentral eolumn, anther wings protruding, lower outer edges of seales are thin and sharp edged. Apex - apex 0.28 em Apex - eenter 0.33 em Widest 0.13 em Ret - ret. 0.09 em Ret - eenter 0.08 em Aw. - aw. 0.15 em Aw. - eenter 0.15 em Ret. Pol ratio 1:4.3 Pollinarium enlarged about I55x. Pollinium length 0.47 mm widest 0.20 mm Retinaculum length 0.12 mm shoulder 0.1 1 mm waist 0.06 mm hip 0.08 mm ext. 0.06 mm Translators length 0.12 mm depth 0.01 mm Caudicle bulb diam. 0.08 mm Translator/caudicle type: d/o Pollinia inner end type: RT 10 Internodes: 6.5-13.4 cm long, some adventitious rooting. Nodes only slightly enlarged. Glabrous. Leaf blades: opposite, petiolate, elliptic acuminate, 8.1 - 12.9 cm long x 3.3 - 4.2 at the widest, both sides glabrous, plinerved. Petiole: 0.87 - 1.75 em long twisted. Glabrous. Rachis: 2.19 em long. The following sheet was labeled Hoya macgregorii Schlechter but it is not that species. Outer lobes of corona on that species are not “breviter exeises” briefly excised. Also the corolla is much different. 11 Reduced copy if the type sheet cahup #5297 Misidentified as Hoya mcgregorii Schlechter 12 Hoya velasioii subsp. grandiora Kloppenburg ISSN 2329-7336 Hoya velasioii sbubp. grandiora Kloppenburg, holotypus #18682 (CAHUP) hie designatus, similes sp. sed eorolla diamentro eomplanatus grandior 1.04 em eontrastre 0.90 em et pollinia longior 0.45 mm eontrastre 0.43 mm et retinaeulum longior 0.20 mm eontrastre 0.15 mm, differt. This subspeeies was eolleeted by the late Professor Juan Paneho it is similar to the speeies but different. Many parts are slightly larger, the eorolla diameter flattened is 1.04 em vs. 0.90 em and the pollinia are also longer 0.45 mm versus 0.43 mm and the retinaeulum is also longer 0.20 mm versus 0.15 mm among other differenees. Flower from Herbarium sheet for Identifieation. Pedieel and ealyx enlarged about 8x. The pedieel is 0.06 em in diameter, broken off so do not have the length. The surfaee is very finely erystalline puberulous. Sepals very small and ovate, 0.07 em long and 0.07 em at the widest with ea. % overlap at the base, ligules present, outer surfaee as with the pedieel very finely puberulous and dense, inside glabrous and shiny, membranous. Ovaries, here dark 0.09 em tall and base pair 0.09 em wide. glabrous. Outside surfaee of the flower enlarged about 8x. Sepals are small and do not eome near the eorolla sinuses. Corolla outer surfaee is finely granulate, glabrous. Sinus - sinus 0.21 em Sinus - eenter 0.15 em Sinus - apex 0.40 em Apex - eenter 0.52 em Widest 0.30 em 13 Inside view of the flower enlarged about 8x. Corolla inside is pubeseent, eoronal lobes exeeed the sinuses of the eorolla, dorsal is eoneave with a keel all the way down eenter with a small umbo at the widest portion. Corona is glabrous, suleate, horizontal, inner lobe short spatulate nearly touehing in eenter. Bottom view of the eorona enlarged about 8x, lobes are ehanneled nearly to the thiekened eentral eolumn, surfaees suleate, glabrous. Upper surfaee of eorona (with eorolla). Apex - apex Apex - eenter Widest Ret. - ret. Aw. - aw. 0.30 em 0.31 em 0.13 em 0.07 em 0.15 em 14 Pollinarium enlarged about 165x. Pollinia are laying slanted so full extent of breadth ean not be measured in this photo. Pollinium length 0.45 mm widest 0.13 mm Retinaculum length 0.20 mm shoulder 0.08 mm waist 0.04 mm hip 0.08 mm. ext. 0.03 mm Translator length 0.07 mm depth 0.01 mm Caudicle bulb is here oval 0.03 X 0.06 mm Translator/caudicle type: d/o Measurements from the herbarium sheet below: Pollinia inner end type: RT Leaves: opposite, petiolate, broadly ovate short apieulate, glabrous, base obtuse, pinnate nervation, mostly 9-15.6 em long x 4. 8-6. 6 em broad. Petioles round 1.0 - 1.8 em long. 0.2 em 0.3 em in diameter. Stems: petiolate, 0.03 - 0.4 em in diameter, terete, glabrous, nodes slightly enlarged, 3- 14 em long. Pedunele ea. 5 em long, raehis fine, pedieels 2.0 em long, flower elusters globose. 15 A reduced copy of the type sheet CAHUP # 1 8682 1 V k. * * I It ^ • py J-.'* V > 'r ■‘H- ,j._i «• ■■ >1 # I « ^ t -ifi- r -f’’ ' ' . •■ , ? 'rf' f?* ■ ■ • -> ^Lat‘^’ *r^-m ' »• ar...^a-L i : J^-i I itarr^ WAl . '”■■ ■p.M. * i . ■ ^ ? . ^ . PIVF4ml|. Ji iiaJic*!: 11- *' ■■‘jr^ .. ..M»ti-|V, ■“.V-j-* ■ *,• . b. •* n|j| ( |. j ||,i ■ KE ■■ I ■ - P I* ■■• ■■ I 1 U^a*! n:"” <»iiei"Tr*- “ MM u.m^.Lci «|6>r 1 i JP ■ 4 [« 4 T ki r. y **4 C-Afi. ' ( ‘p| f IMl IMV-- M r -bl 16 Hoya columna Kloppenburg ISSN 2329-7336 Hoya columna Kloppenburg sp. nova, holotypus #41193 (CAHUP) hie designatus. This is the first Philippine hoya speeies with a pollinia 0.38 mm long, it also does not fit any present Philippine hoya with a pollinia 0.16 mm wide, in addition this speeies has an unusually long thin eoronal eolumn 0.07 em long, the pollinarium is also unique in several ways. Is it does not eonform to any other Philippine hoys speeies at this time. So it is unique in a number of ways. It has been named for the unusual eoronal eolumn. Determination: possible new speeies. Pedieel enlarged about 8x. em long, 0.05 em in diameter. Terete, glabrous eurved, 1.5 Side view of the ealyx and ovaries enlarged about 8x. Calyx lobes do not reaeh the eorolla sinuses are glabrous on both surfaees. Sepals are 0.10 em long and 0.10 em wide at the base, no ligules observed. Ovaries: short, glabrous 0.10 em tall and base pair 0.05 em wide. 17 Top view of a calyx with some sepals missing enlarged about 8x, they are thin, glabrous, triangular with broad base and rounded apex. Closed flower outside surface with pedicel attached enlarged about 8x. Calyx lobes have turned up from the surface on drying. Outer surface of corolla is glabrous and granulose. Note the sepals do not reach the corolla sinuses and that the coronal lobe outer apex exceeds the sinus (lower right hand corner). Corolla outer surface, flower closed, corolla is thin, deeply cut and wide above the sinuses. Sinus - sinus 0.15 cm Sinus - center 0.10 cm Sinus - apex 0. 30 cm Apex - center 0.40 cm Widest 0.30 cm 18 Inner surface of the corolla enlarged about 8x. This surface is finely puberulent except the raised collar and the edges of the lobes. Collar is a little thickened and 0.05 cm tall. Top view of a flower showing that the coronal lobes exceed the corolla sinuses. Corona is glabrous and finely sulcate on surfaces. Bottom side of the corona enlarged about 16x, the lobes have acute outer apices and channeled to near the central column, with the sides diagonally sulcate. Anther wings are acute and project slightly beyond the corona sinuses. Another view of the corona bottom showing the long central column, it is 0.07 cm tall. Outer coronal lobes are raised with acute apices. 19 Top view of the eorona. Inner lobes are aeute sharply raised, spatulate and do not eover the eenter. Dorsal surfaee is a little eoneave. Apex - apex 0.20 em Apex - eenter 0.22 em Widest 0.10 em Ret - ret. 0.05 em Ret - eenter 0.05 em Aw. - aw. 0.10 em Aw. - eenter 0.08 em Measurements from the herbarium sheet. Foliage: opposite, petiolate, ovate apieulate, thin, glabrous, quintuplinerved, size very uniform, 7.2-8. 1 em long 3.2 - 5.1 em widest near middle. Petioles: 1.6 em long and 0.35 em in diameter, round. Peduncles: variable in length 2.2, 6.4; 10.1 em smaller diameter than most stems. Internodes: mostly 9.5 em long up to 0.7 em in diameter many smaller. Pollinarium below enlarged ea. 25 Ox 20 Pollinarium length 0.38 mm widest 0.13-16 mm Retinaculum length 0.09 mm shoulder 0.10 mm waist 0.04 mm hip 0.06 mm Translator length 0.08 mm depth 0.02 mm Caudicle bulb diam. 0.04 mm oval. Pollinia inner end type: R Translator/caudicle type: p/o Ret:/Pol. ratio: 1:2.25 21 Reduced copy of the type sheet CAHUP #41913 r. r , *^r *■ * i/'Lll =1 ? i *:£* 22 Hoya maximowayetii Kloppenburg ISSN 2329-7336 Hoya maximowayetii Kloppenburg sp. nova, holotypus 5271 (CAHUP) hie designatus, non similes alius hoya speeies, flos laeteus, eorollae lobus diametro eomplanatus 0.90 em, extus glabra, intus eonfertus pubeseentia et eoronae lobus exterior obtusa; pollinia 0.38 mm longus. This hoya speeies is not similar to any other hoyas, flowers are whitish (as few are) and the eorolla flattened has a diameter of 0.90 em with the ventral surfaee glabrous and the dorsal surfaee densely pubeseent, the eoronal outer lobes are obtuse, the pollinia are 0.38 mm long. See the following detailed measurements of parts. This new speeies is named for my old friend, Maximo Wayet who worked for years for Professor Juan Paneho and eolleeted many speeies in the northern Luzon areas. Colleeted by Maximo Wayet at Benguet Village, Querino, Luzon, Philippines Photos 19 April 2006. Determination: near 5269 & 5293 - Stems: terete, glabrous. Peduncles: nill - 3.9 em long on sheet below. Foliage: opposite, glabrous, oblaneeolate, triplinerved apex aeute base euneate 9.3-12.1 em. long x 2.3 - 3.9 em at widest. Petioles short ea. 1 em. Pedicel: enlarged about 8x. Curved, terete, glabrous, purplish, 1.8 em long 0.05 em in diameter. Flowers whitish. Calyx: enlarged about 8x. Sepals nearly reaeh the eorolla sinuses, slight basal overlap; I did not see any ligules, edges entire, thin. 0.12 em long 0.10 em at the widest. Ovaries: short dome, glabrous, 0.07 em long and base pair 0.05 em wide. Flower outside showing a dark ealyx and eorolla lobes eupped up in drying, eorolla outside glabrous, apex aeute. 23 Corolla: outside surface enlarged about 8x, lobes are deeply cut and surface glabrous with a central collar a little thickened. Sinus - sinus Sinus - center Sinus - apex Apex - center Widest 0.14 cm 0.13 cm 0.28 cm 0.45 cm 0.29 cm Inside surface of the corolla enlarged about 8x, this surface is densely pubescent except near the center collar area. Jfil r ■ ' « M W P '■.‘A f ? j ** • - j •- A V . 4 Corona: under surface enlarged about 16x. Lobes channeled below to near the thickened central column that is ca. 0.05 cm tall, surfaces are glabrous and lobes are finely sulcate. 24 Top surface of the corona enlarged as above. Surface is glabrous. Inner lobe raised a little and does not reach the center, outer lobe obtuse, dorsal slight concave with a low ridge all the way. Apex - apex 0.20 cm Apex center 0.22 cm Ret. - ret. 0.06 cm Aw. -aw. 0.10 cm 25 Pollinaria both with one pollinium missing and retinaculum twisted a little on both. Head of arrow is 0. 1 mm long. Pollinium length 0.38 mm widest 0.15 mm Retinaculum length 0.16 mm shoulder 0.13 mm hip 0.04 mm waist 0.05 mm ext. 0.04 mm Measurements of the retinacula are close approximations. Can not determine length of the translators, although they are short, nor the caudicle bulb diameter. Translator/caudicle type: Is/o Pollinia inner end type: T Retinacula to Pollinium Ratio 1:1.7 26 Reduced copy of the type sheet, CAHUP #5271 27 Hoya wibergiae subsp. alba Kloppenburg ISSN 2329-7336 Hoya wibergiae subsp. alba Kloppenburg subsp. nova, holotypus 14203 PNH, hie designatus. Similis sp. sed flos eolor differt et eorolla diametro eomplanatus brevior 1.00 em eontrastre 1.60 em et eoronae lobus brevior et angustior 0.30 em x 0.32 em eontrastre 0.40 em x 0.48 em, aliter similes. This subspeeies has a pollinarium similar to the speeies but the flower eolor is different, and the eorolla flattened has a smaller diameter 1.00 em versus 1.60 em and also the eoromnal lobes here are shorter and more narrow, 0.30 em x 0.32 em versus 0.40 em x 0.48 em, otherwise they are similar. Colleeted by G. E. Edano Vietoria Mts. Palawan 18 Mareh 1951 growing in moss labeled H. gracilis Sehleehter ineorreetly, it is not in the Seetion Aeanthostemma (Bl.) Kloppenburg. White flower, 600m elevation. This appears to be elose to Hoya wibergiae Kloppenburg. Roll 137 Drawing 245. Stems flexible, glabrous, rooting below nodes. Leaf blades: 6-9 em long x 1.5 - 3 em widest, glabrous. Petiole 0.4 - 1 em ehanneled above. Peduncle: very short, 0.4 em pubeseent. ■ Pedieel & ealyx enlarged about 8x. Pedicel: 1.2 cm long strict, finely pubescent, with e vascular bundles, ca. 0.5 cm in diameter enlarging below the calyx. Calyx: 0.13 cm long x 0.1 1 at widest, broad ciliate Outside of flower enlarged about 8x. Sepals reach about y4 way to the corolla sinuses, is deeply cut. Eobe apex acute widest above the sinuses. 28 Flower inside enlarged about 8x. Corolla is pubeseent. Coronal lobes exeeed the eorolla sinuses. Sinus - sinus 0.23 em Sinus - apex 0.37 em Sinus - eenter 0.19 em Apex - eenter 0.50 em Widest 0.30 em Bottom view of the eorona enlarged about 8x. Lobes are ehanneled nearly to the thiekened eolumn, apex sub aeute. lobes are wide in the eenter above the sinuses, anther wings are wide and protrude beyond the sinuses. Top view of the eorona enlarged about 8x. Seale dorsal has a large keel extending down eenter longitudinally, surfaees glabrous. Inner lobes dentate. Apex - apex Apex - eenter Widest Ret. - ret. Aw. - aw. 0.30 em 0.32 em 0.17 em 0.10 em 0.23 em 29 / • Pollinaria enlarged about 85x. Pollinium: length 0.60 mm widest 0.25 mm Retinaculum: length 0.25 mm, shoulder 0.10 mm; waist 0.03 mm; hip 0.06 mm, ext. 0.01 mm Translator: length 0.07 mm, depth 0.02 mm Caudicle: bulb diam. 0.04 mm Translator/caudicle type: p/o Pollinia inner end type: RT 30 Hoya vicencioana subsp. quezonensis Kloppenburg ISSN 2329-7336 Hoya vicencionana subsp. quezonensis Kloppenburg sp. nova, holotypus 5294 (CAHUP) hie designatus. Similis sp. sed folia breviore 7.8-12.6 x 3. 5-5. 4 eontrastre 10.5- 17.6 X 6.0- 8.2 et eorolla diametro eomplanatus breviore 0.90 em eontrastre 1.22 em, differt. The subspeeies is similar to the speeies but the foliage is smaller and the leaf bases are not as eordate and the eorolla flattened is also smaller 0.90 em in diameter versus 1.22 em, among other differenees. Colleeted by Blass Hemaez 17 Mareh 1990, UP Landgrant, Quezon, Philippines. Roll 242. Details follow. Internodes: 5.5 - 10 em long Foliage: opposite, petiolate, elliptie with an apieulate apex aeute, base slightly rounded, glabrous, pinnately nerved, blade 7.8-12.6 x 3.5 - 5.4 em; petiole 2.7 em long, glabrous. Peduncle: extremely short here with raehis faseieulate. Photo and data 3/29/99. Pedicel: filiform, terete, glabrous 1.0 em long 0.05 em diam. Calyx: short, broad, triangular sepals, eiliate, outside surfaee granulate, glabrous. Ovaries: dome shaped 0.07 em tall and base pair 0.05 em wide, glabrous. Flower, eorolla dark, ventral glabrous, dorsal pubeseent, with eorona in eenter sitting atop a eolumn, flowers very small. Outer lobes raised above the eenter, dorsal surfaee sway baeked with eenter raised, outer apex rounded, thin, prominent anther wings. Inner lobes round (dentate?) sloping outward, dorsal with eentral ridge, same on underside. Sinus - sinus 0. 1 8 em Sinus - eenter 0. 1 5 em Sinus - apex 0.30 em Apex - eenter 0.45 em Widest 0.25 em 31 Another view of the eorona. Photos do not show mueh detail and are of little value. Retinaeulum and extensions well developed, inner apex rounded, with translator on left side attaehed well down on side. I ean not determine measurements as I have no seale showing in photo. Pollinium has germinated, apex rounded inner apex narrowing and also obtuse. It is 0.42 mm long and ea. 0.15 mm at the widest. 32 Another photo of a pollinarium. Here the pollinium seems to measure 0.40 mm long and 0.16 mm at the widest. Retinaeulum turned on its side is ea. 0.20 mm. long. Pollinia inner end type: T Translator/caudicle type: d/o ? 33 Reduced copy of the type sheet CAHUP #5294 N i rLJ» V#^rr 34 Hoya velasioii Kloppenburg ISSN 2329-7336 Hoya velasioii Kloppenburg, Holotype 9136 (CAHUP) hie designatus. Similes Hoya afuangae Kloppenburg & Cajano 2015 sed eorolla diametro eomplanatus brevior 0.90 em eontrastre 1.20 em et pollinia 0.43 mm longa eontrastre 0.39 mm, differt. This speeies is similar to Hoya afuangae Kloppenburg & Cajano 2015 but differs in the pedieel being slightly puberulous, the eorolla flattened 0.90 em vs. 1.20 em, the pollinia lengths longer 0.43 mm vs. 0.39 mm and the leaves are here pinnately nerved not plinerved, among other differenees. Colleeted by J. R.Velasio. Photos 3/2/92 Roll 103-14 Drawing 182. Pedicel: enlarged about 16x. 1.5 em long 0.04 em in diameter, slightly puberulous, terete, eurved. Baek side of the ealyx enlarged about 8x. Sepals are 0.12 em long and 0.07 em wide at the widest near base, thin, apex rounded. Corolla outside view lobes folder in over eenter enlarged about 8x, surfaee is finely granulose, glabrous. Central eollar thiekened protruding toward the ealyx, opening 0.10 em x 0.07 em 0.02 em tall. Coronal outer lobes exeeed slightly the eorolla sinuses; are ehanneled all the way to the eentral eolumn, surfaees are suleate Sinus - sinus 0.25 em Sinus - eenter 0.26 em Sinus - apex 0.30 em Apex - eenter 0.45 em Widest 0.28 em 35 Corolla lobe enlarged about 8x, apex is aeute. Corolla inner surfaee is finely puberulous. Bottom view of the eorona enlarged about 8x. Lobes are ehanneled all the way to the long eentral eolumn, apiees are aeute. Central eolumn is 0.04 em tall, half with eorolla and half on eorona 0.012 em in diameter. Top view of the eorona enlarged about 8x. Lobes are eoneave with a keel running the entire length, inner lobes are elavate and raised a little, broadest at the anther wings. Anther wings are thiek, apiees rounded. Apex - apex 0.30 em Widest 0.13 em Ret. - ret. 0.05 em Ret. - eenter 0.06 em Aw. - aw. 0.15 em Aw- eenter 0.12 em Side view of a eoronal seale enlarged about 16x. Seales are horizontal with inner apex raised equal to the anthers that are still exposed in the eenter, lobes thin. 36 Pollinarium enlarged about 165x. Pollinium length 0.43 mm widest 0.14 mm Retinaculum length 0.15 mm shoulder 0.08 mm waist 0.07 mm hip 0.08 mm ext 0.06 mm Translators length 0.12 mm Translator/caudicle type: p/o Pollinia inner end type: RT Caudicle. bulb diam 0.05 mm Ratio: r/p 1.8 depth 0.03 mm Leaves: opposite, petiolate, glabrous, elliptie-laneeolate, base narrowly obtuse, apex aeute, pinnate nervation, midrib prominent below and raised. 37 Holotype copy here reduced CAHUP 9136 V- ^ r ^ ■ .« 1 ^ X tmltt iM cAl> * ■■ -*** 38 Hoya capotoanensis Kloppenburg ISSN 2329-7336 Hoya capotoanensis Kloppenburg sp. nova, holotypus 15642 (PNH) hie designatus. There is no present hoya speeies with pollinia this short (0.22 mm) with translator type Is/o nor any with a pollinia this short and with eoronal lobes this size (0.25 em.). It is unique and different in these respeets. This new speeies was eolleeted by G. E. Edano at Mt. Capotoan, Samar 10 Jan. 1952 along a forested ridge, 180 m. elevation, flower greenish-yellow. Stems: beeoming large, woody, nodes enlarged, glabrous. Blade: 7.5 - 12 em x 3.5 - 4.9 em, glabrous, leaf base sub-euneate to sub-obtuse, apex aeuminate, nervation 3-5 triplinerved. Petiole 1-2 em long, glabrous ehanneled above, deeiduous, eireular sears, eurved. Rachis: domed, short. Pedieel and ealyx side view enlarged about 8x. Pedieel is 2.5 em long, terete, glabrous; ealyx base is tapering. Calyx: sepals 0.12 em x 0.09 em, triangular, edges membranous, 10 % overlap, apex sub aeute. Ovaries: 0.6 em x 0.05 em small domed. Bottom view of the eorolla enlarged about 8x, outside surfaee is glabrous. Sinus - sinus Sinus - eenter Sinus - apex Apex - eenter Widest 0.20 em 0.20 em 0.30 em 0.45 em 0.26 em 39 Top view of the eorona enlarged about 16x, lobes exeeed the eorolla sinuses, anthers exeeed the inner lobe apex. Dorsal is broad with a slight domed eentral ridge. Apex - apex Apex - eenter Widest Ret.- ret. AW. - aw. 0.25 em 0.28 em 0.14 em 0.08 em very far in toward eenter. 0. 16 em very prominent extended, rounded ends. Pollinarium enlarged about 165x. Pollinium length 0.22 mm widest 0.05 mm Retinaculum length 0.15 mm 40 shoulder waist hip ext 0.10 mm 0.06 mm 0.08 mm 0.03 mm Translators length 0.07 mm ea. Translator/caudicle type: Is/o Pollinia inner end type: R 41 ifL % m hifu !Hi I ■ - ^ ^ f^tjcjiLih. Of TTfF tnittii'PntfEs Reduced copy "of the* herbarium sheet. 1 CTl? ,v ^ v e T ^ I tfcr M ‘ IVi-mti ft iff^ar^r^. I FTt ^ ^ ■Zj^ ^Uf V p ■ .jr ■, P^H» ■'* * ^ \mlTii i f ^^Jh■l r • ij r; h ?■■•■■“• p 1*4 V 42 Hoya galeraensis Kloppenburg ISSN 2329-7336 Hoya galeraensis Klopenburg sp. nova, holotypus 3312 (CAHUP) hie designatus. No other Philippine hoya speeies has a white flower with palmate nerved foliage. There are only about 5 speeies with white flowers and only a few speeies with palmately nerved leaves. This new speeies also has an unusual domed eoronal eenter that is rare. This unique speeies was eolleeted by the late professor Juan at Puerto Galera, Palawan, Philippines in 1954. 1 first examined it in 3/29/99. Film roll 244, The ealyx base and pedieel 0.06 em in diameter. Corolla outside enlarged about 16x. Flower is labeled as white, eorolla is refiexed with thiek ealyx base and pedieel, surfaee is glabrous, lobes turned under along sides. Apex aeute. Sinus - sinus Sinus - eenter Sinus - apex Apex - eenter Widest 0.17 em 0.17 em 0.32 em 0.46 em 0.28 em V 43 Inside (dorsal) surface of the corolla enlarged about 12x. This surface is granulose- glabrous with central collar thickened and raised a little. Bottom view of the corona. Scales are channeled below nearly to the center with overlapping sulcate sides, apex acute. Channel lobes fold out to form a “V” channel centrally. Center is a short thickened walled column. Anther wing apices rounded and not protruding from the sinuses. 44 Top view of the flower with eorona, the lobes do not reaeh the eorolla sinuses. Lobes raiser over dome like eenter outer apex rather obtuse, inner lobe spatulate nearly eovering the eenter. Dorsal surfaee keeled part way Apex - apex Apex - eenter Widest 0.15 em 0.17 em 0.15 em Leaves: opposite, petiolate, glabrous, elliptie-ovate with base obtuse, apex aeuminate, palmate nerved. 45 Reduced copy of the type sheet, CAHUP 3312 Loci L£ty_ lalanj or DcfSr ILIPPIWES HB^rlun. Collogfl i£rle(»ltur4 IftlVaralty of tlV« PhlTlpplfleo S<;iennfiq n coiin»jli nutiife ^ / TTUaLH^tr] Aftituda above a.av'al Trca: ahrub; JmaTir vine: hiarb Z/i:.^ Hel£h!; ■ Dtamecer, hr^nf st [OnOr, cDtgr ■ gtg , > ceTor 7 "ctg^ Speelaf note uaaa CoLl^ bo ■ -jd T Uat c ~" i i i^j ^ JJLOHA OF THE pKll-U’PtNEa 46 Hoya sulitii Kloppenburg ISSN 2329-7336 Collected by M. D. Sulit, (PNH) 10147, 15 April 1949, Mt. Katanglad, Bukidnion Prov. Mindoro, Philippines. Flower creamy, collected at 1800 m asl at forest edge. This new species has some characters near Hoya cardiophylla most similar in foliage but here the corolla is densely pubescent inside, which negates it being that species. The pollinium is similar however but smaller 0.48 x 0.19 mm versus 0.50 x 0.17 mm and the retinaculum is longer 0.26 mm versus 0.20 mm. Pedicel: section with the calyx attached, enlarged about 8x, they are 2.5 cm long, terete, strict, finely puberulous 0.13 cm in diameter. Calyx: outside granulate, inside slick glabrous. Sepals 0.28 cm long and 0.25 cm at the widest. Ovaries: are dome shaped, glabrous 0.18 cm long and the base pair is 0.25 cm wide. Top view of a flower enlarged about 8x. The crown is relatively horizontal, although on some dried flowers the outer lobes of the corona turn up as in the next picture. These flowers were soaked in Kew Solution and then photographed. 47 Another side view of the flower enlarged showing the densely pubeseent inner eorolla surfaee, espeeially dense inward from the eorolla lobes, whieh are here reflexed. The eolumn is also relatively long, so the eorona sits well above the eorolla surfaee. Lobes are ehanneled below, with suleate surfaees, anther wing apiees protruding slightly and triplex. Side view of a eoronal seale enlarged about 16x. that would appear to indieate that the inner lobe exeeeds the anther and is narrowly dentate (whieh is different then H. cardiophylla) and that the outer lobe turns up, possibly from drying, also not like in the other speeies in whieh the eorona is horizontal. Dorsal is eoneave, anther wings deeply seythe shaped. Pollinarium enlarged about 165x. The retinaeulum is similar to that of Hoya cardiophylla but slightly different, also the pollinia are mueh shorter and the translators although attaehed well down, are here eurved inwardly. Pollinia length 0.48 mm. widest 0.19 mm. Retinaculum length 0.26 mm. shoulder 0.12 mm waist 0.05 mm. hip 0.10 mm. Translator length 0.17 mm. depth 0.03 mm. Caudicle bulb diameter 0.05 mm. Translator/caudicle type: Is/o 48 Leaves: opposite, petiolate, glabrous, ovate with eordate base and aeute apiees, pinnate nervation, with mostly 6 pairs of nerves sub-anastomosing, margins a little revolute, midrib plainly visible below. Redueed eopy of the type sheet: 49