open if eet Rotor ne parernne Ved G4 s, THE JO; Uy Ry Nea OF » THE LINNEAN SOCIETY. ZOOLOGY. (Gay ie ING SL \ Noe 5.053 VOL. XI. Qs mal pane? Sey ee ~ LEPo s yY LONDON: SOLD AT THE SOCIETY’S APARTMENTS, BURLINGTON HOUSE AND BY LONGMANS, GREEN, READER, AND DYER, AND WILLIAMS AND NORGATE. 1873. PRINTED BY TAYLOR AND FRANCIS, RED LION COURT, FLEET STREET. LIST OF PAPERS. Auuis, THomAS, F.L.S. On the Skeleton of the ApteryX 2.2... eee ee eee e cree rennes 523 Barn, W., M.D., F.R.S., &e. Description of some new Species of Annelida and Gephyrea in the collection of the British Museum ............-+++-065 94 Burmeister, Dr. Hermann, F.M.L.S. Observations on a Light-giving Coleopterous Larva .......... 416 CamBnrinGE, Rev. O. P., M.A. On some new Species of European Spiders. (Plates XIV. & MAE ooecncoso boos ondogeenn vo po0cdspodGD Odo RODD HO. 530 CoLtiinawoop, CuTHBERT, M.A., F.L.S., &c. On a new Form of Cephalopodous Ova. (Plate Dasher’ oh 90 Day, Surgeon-Major Francis, F.LS., &c. On some new Fishes of India .........se cess eeee se neereee 524 Garner, Rosert, F.LS. On the Formation of British Pearls and their possible Improve- TON wogoebodnobeob06 Jed Op ORD oopD noo abee Ooo snp uL Oa Ot .. 426 Guuicx, Rev. Joun T. On Diversity of Evolution under one set of External Conditions. 496 Kirsy, W. F., Assistant in the Museum of the Royal Dublin Society. On the Geographical Distribution of the Diurnal Lepidoptera as compared with that of the Birds .......++sssseesesereeee 431 Luxxock, Sir Joun, Bart., F.R.S., F.LS. On the Origin of Insects......... ss see e rere rete tener eee enes 422 M‘LacuLan, RoBERT, F.L.S., Sec. Ent. Soc. On new Forms, &c., of Extra-European Trichopterous Insects. (Plates 11., TIL., & TV.) ... cs cee eee reese eens aE acate 98 An Attempt towards the Systematic Classification of the Family Ascalaphid@.........seeec essen eee Leos PHUG OOM OOO 219 Mutter, ALBERT, F.L.S. Note on a Chinese Artichoke-Gall (mentioned and figured in Dr. Hance’s paper “ On Silkworm-Oaks ”) allied to the Eu- ropean Artichoke-Gall of Aphilothrix gemma, Linn, ........ 428 iv Page Monir, James, M.D., F.L.S., &c. Notes on the White-beaked Bottlenose, Lagenorhynchus albi- Prosi Ere (CRED Wo) GeaccngoondbondoosoboccKoseu0s 141 Murray, ANDREW, F.L.S. On the Geographical Relations of the Chief Coleopterous Faune. 1 OrmeERoD, Miss ELEanon A. Observations on the Cutaneous Exudation of the Triton cristatus, on Greatuwater= Newt woul orto sictereiievehorercrerarey ctor aicrs ratele tee 493 Pascogr, Francis P., F.L.S., late Pres. Ent. Soc. Contributions towards a Knowledge of the Curculionide.— Lepren Ul (etry WAGs AOL AUIS We ID.) s650s560cc0cc8 50 154 Contributions towards a Knowledge of the Curculionide.— LetigSUBG | (eet 0-605 01 UI GUL) pooanncsgasooebad. 440 Ports, Tuomas H., Esq., F.L.S. Notes on Keropia crassirostris, Gml. (“‘ Piopio”) ............ 505 SaunpErs, Hpwanp, I’.L.8. Descriptions of Buprestide collected in Japan by Groner Lewis, Hsq....... lao qenondcdeaes DRE onde Bibs a.6u mde 509 Smitu, F. Esq. A Catalogue of the Aculeate Hymenoptera and Ichneumonide of India and the Eastern Archipelago, with Introductory Re- marks by A. R. Watiace. (Communicated by W. W. SVMs JONG) 6 AGS dodood co UnoocenduodoGDOGodCOnCOaS 285 TrimMEN, Rowand, F.L.S., F.Z.S., M.E.S. Note on a Paper by ANDREW Murray, F.L.S., “On the Geo- graphical Relations of the Chief Coleopterous Faune” .... 276 STUN TDR Xai a alle la a ere hace eM age ail amauta tani mee cer amaielis tec erere 549 THE JOURNAL OF THE LINNEAN SOCIETY. On the Geographical Relations of the Chief Coleopterous Faune. By Anprew Murray. [Read December 17, 1868.] * Juparne simply from their structure, habits, and economy, there are reasons why Beetles ought to excel every other class of organ- ized beings as exponents of the past geography of the globe. I say so after having turned over in my mind and contrasted every class of animals and plants with each other with the view of de- termining for myself which would be most likely, through the study of its geographical distribution, to throw light upon the past history of the earth. I can think of none so likely to do so as insects, and of insects as Beetles. Over all marine animals they have the insuperable advantage of inhabiting the enclosed instead of the enclosing spaces, of living on dry land and not being able to go beyond it. Over plants, with which their distribution in many respects accords, they have © the advantage of being more difficult of dissemination, for neither their eggs nor themselves are endowed with the dormant vitality of seeds, nor with that endurance of exposure to different condi- * This paper was read on December 17, 1868, but by permission of the Council I have brought it down to the state of our knowledge at the date of publication. —A. M. LINN. PROC.—ZOOLOGY, VOL. XI. 1 2 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF tions which may, and occasionally does, enable seeds to be carried in the stomachs of birds or floated across wide oceans to distant lands *. They have also the advantage over the larger and more highly organized animals in that they can survive and find food where the latter could not. Their food is so various that nothing but a total extinction of all other life could wipe them off from the face of a country—a partial submergence of land for even a short period might destroy every mammal upon it, but so long as a tree-top is above the flood or an uncovered rock remains on which they can take refuge, the life of the Beetle class is safe when the waters abate. A succession of cold seasons in which no plant can bloom might destroy those kinds of animals for which, like the bee, flowers and honey are necessaries of life, some bee- tles might indeed then go ; but there are plenty that feed on leaves or stems to preserve the Beetle type in the frozen land. Their numbers, too, multiply the chances of escape in the case of dis- aster, and their powers of flight enable them to take advantage of such as occur. Further, the powers of flight, although sufficient for a moderate distance, are not like those of birds, so great as to carry them to new lands at great distances and so to risk the dis- turbance of faunas which such powers, if possessed by such mul- titudes, might possibly produce. In many respects, too, they are as much adstricti glebe as plants themselves, for a vast host are limited each to one particular plant for food. As in plants, in- deed, there are some kinds of Beetles more open than others to the suspicion of having been introduced from one isolated land to another, as, for example, the timber-borers or Longicorns. But there are others, as the hunting or carnivorous species, the apte- rous species, the blind insects, and others of less specialized struc- ture, whose presence in discontiguous countries seems to bid defi- ance to any explanation other than that of former continuity of soil. In Madeira, for instance, where the number of admittedly introduced species is very great, there is not one introduction be- longing to the hunting families; and if this is the case there, not- * It is a digression, but it is worth making one, to point out that if plants can be disseminated in the way supposed, and Beetles, or certain families of Beetles, can not, the attempt to explain the distribution of the former as due either solely or mainly to these means must be abandoned in every case where their distribution corresponds with that of the latter. The common effect must have been produced by a common cause. And it so happens that this correspondence exists in all the more important and puzzling facts of distribution. TILE CHIEF COLEOPTEROUS FAUN®. 3 withstanding the accessibility of Madeira to the introducing agencies of man (to which most of the other introductions are referable), it becomes still more difficult to conceive of the dis- semination of that kind of beetles by agencies independent of man. But besides the advantages which the structure, habits, and economy of beetles give for the interpretation of their geographi- eal distribution, there is another important speciality inherent in them which I shall amply illustrate in the course of this paper, and which renders them peculiarly available for the study of its problems, viz. a long-enduring persistency of form by which the same type has been preserved through diverse modifications during many geological epochs. This peculiarity is shared by all other insects, as well as in different degrees by all beings of inferior organization ; and the consequence is that in trying to make out the past history of a country through its fauna and flora, we must take each class of beings by itself and study its relations separately, or we shall run the risk of confounding events belong- ing to different dates. To do otherwise would be like attempting to compile a history of England by combining the political history of one age with the ecclesiastical of another and the scientific of a third. The mammalian fauna took its present form long after the insects had received theirs, and these earlier-dated forms should therefore be able to tell of events long antecedent to what the mammals could speak of. The relations of each must there- fore be studied independently, and it is only after all shall have been separately deciphered that the conclusions respectively drawn from each can be brought together and some common general result arrived at. In the mean time, by endeavouring to ascertain the relative date of appearance of insects of various types in dif- ferent countries, we may be able to assign the order of precedence of a succession of events whose occurrence we can scarcely doubt, but whose order of date we could not otherwise guess at. My purpose in the present paper is to submit some infer- ences of this nature from a general view of the geographical dis- tribution of the Coleoptera, indicating a somewhat different ar- rangement of land and water in ancient times from that which is usually supposed, and to strengthen these inferences by references to what seem to me corresponding facts in other branches of na- tural history. The first point to which 1 shall direct attention is the very in- 1* 4 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF teresting problem of the occurrence of similar forms in the tem- perate and cold regions of both hemispheres. Hitherto the hypotheses by which this has been attempted to be explained have, I believe, either been: 1st, by accidental introduction ; 2nd, by the supposition that the glacial epoch had so modified the climate of the globe as to allow an extension or interchange of faunas lying on different sides of the equator; 3rd, by the suppo- sition that one general fauna had formerly extended over the whole world, and that the similarities which we find in antipodal countries are’relics of this general fauna; and 4th, that a former geographical connexion with identity of fauna and flora must have subsisted between the two regions. There is another hypothesis to the effect that the similar species are representative species, meaning by representative something different from derivative and independent of affinity. This latter proposition, I frankly confess, is beyond my conception. I cannot conceive of any other kind of representation in species than that arising from derivation. The other hypotheses are entitled to more consideration, and I shall briefly state my opinion upon each. As to similarity being due to the introduction of species by accidental dispersal, it is to be noted that this cause must always be in its very nature exceptional and isolated, and cannot be expected to make its impress on a whole fauna. The Atlantic islands, for example, which were, and perhaps still are, very generally cited as an instance of colonization by acci- dental introduction, have been shown by Mr. Wollaston to have all one coleopterous subfauna, and that one peculiar to themselves. The general basis is European, but overlying that is something else, a number of species of special type found in all and found no- where else. Now if these islands, as is maintained by some, have been peopled by chance visitors from Europe, how have they all gut in addition this special type? and why should there be, as in Trophonius Cave, “ vestigia nulla retrorsum”’? Why have none of these special forms ever wandered to Europe? Why should things only come from Europe and nothing ever go back in re- turn? There is no law against reciprocity here, and yet it is rigorously excluded. Moreover the explanation, if true, should apply to every part of the globe, and illustrations of its existence should be in greatest force in the lands which are nearest to each other and which have most intercourse with each other. And this is notoriously not the case. Australia, although so much TUE CHIEF COLEOPTHROUS FAUND. i) nearer India than Chili, has more affinity with the latter than the former. The Cape-Verde Islands, although so near Cape Verde, have their affinity not with Africa, but with Europe and the other Atlantic Islands. Even in the introductions by man it does not apply. Hear what DeCandolle says of plants. “TI am surprised that the commerce of the United States, with Brazil, Chili, New Zealand, the Sandwich Islands, and China, a commerce which has been carried on with great activity for upwards of thirty years, has not yet brought about the naturalization of species from these regions. Up to the present time there is no appearance of it. The Rubieva multifida, which has begun to appear at New York, and of which the naturalization is not yet consolidated by the proof oftime, is the only plant perhaps which has come in this manner. In future some will arrive, without doubt.” [Why so? Surely not from what has happened in the past?] “ They may compen- sate to some extent perhaps the probable diminution of those which will come from Europe’’*. Facts are accumulating upon us to show that diffusion of plants and animals by accidental circumstances beyond physical barriers, such as seas or impassable mountains or deserts, bears no import- ant part in the establishment of any definite fauna or flora. They bear a part, although asmall one, in the introduction of occasional new clements into a fauna or flora; but these remain like lumps of stone lying on a soil with which they can neither become in- corporated nor harmonize, usually readily distinguishable and re- ferable to the mountains or strata more or less distant from which they have come. Actual continuity of soil aud non-interruption by barriers is, I believe, the only cause by which any fauna with a definite character (and no true fauna is without one) has been produced, and subsequent isolation, at least so far as regards phy- sical conditions, that by which it has been preserved. The coral islands of the Pacific are a case in point. They have been sup- plied both with a fauna and flora entirely from without and by chance dispersal ; and they furnish an admirable example of the kind and amount of inhabitants that is to be got by such intro- ductions, even under the most favourable circumstances of tran- quil seas, warm climate, and favouring currents ; and allowance to such an extent I am always ready to make in examining the ele- ments of any fauna or flora. The details of such a fauna and flora will be given further on when I come to discuss the fauna of * DeCandolle, Géogr. Botanique raisonnée, p. 755 (1855). 6 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF the Pacific islands; in the meantime I may briefly characterize it as meagre to the last degree, most unequal in its proportions, and all traceable to the shores of the nearest lands from which the currents set. The supposition that the existence of the similarity in ques- tion is due to the facilities for migration to or from the northern or southern hemispheres afforded by the low temperature of the glacial epoch is open to various answers. But it is unnecessary to discuss them at all; for I shall presently show that the re- semblances to which I have to refer were already in existence before the glacial epoch commenced, consequently could not have been caused by it. The hypothesis that similar forms occurring at distant places are the remains of a general fauna (or, at least, of a more general fauna than now exists), which had in former times ex- tended over the whole or the greater part of the world, is more attractive or more formidable. I used to think that in that hypothesis I had a satisfactory explanation of all such anomalies as I speak of. Like Shak- speare’s barber’s chair, it fitted all comers. If the similarity was widely spread, it was due to universal prevalence in former times. If found only in one or two isolated spots, then there were solitary relics of a once universally distributed type! But I confess that my faith in my specific has latterly been a good deal shaken. It costs me nothing to say so, for consistency is a vice to which I have never been addicted. I believe it still to be probably the true explanation of those cases (as in Ferns, for example) where the same type is very widely and gene- rally distributed; but I have abandoned it for most isolated instances, aud for all specially localized faunas. In the first place, although I do not dispute that in the earlier stages of the history of our planet there was a greater homogeneity of type than there is at present, it seems pretty well established now, that there have been geographical regions with faunas and floras differing from each other, not indeed to the same degree as now, but to some extent, from the very earliest times of which we have any fossil record ; and in the next place, although it is not impos- sible that a universally distributed form may have died out every- where but in one or more specified spot or spots, the doctrine of chances prevents us accepting the hypothesis whenever such relics cease to be solitary. Species No. 1 may be a relic left at THE CHIEF COLEOPTEROUS FAUNZ. 7 spots A and B, and nowhere else; but the moment we find another supposed relic, species No. 2, also left in A and B, and nowhere else, doubt assails us, and increases in an inverse ratio with the occurrence of every additional relic. The fourth supposition is, I think, the true one, namely, continuity of soil at some former period; and upon that as a basis I rest the propositions Iam about to submit. Upon it, I think, I can explain satisfactorily many of the remarkable in- stances of peculiar geographical distribution which have hitherto defied the ingenuity of naturalists to solve, and notably that which I have first sct before mo, viz. the resemblance which species from the temperate regions of the northern hemisphere bear to those from similar latitudes in the southern hemisphere. With the help of the above postulate I can trace the links all the way from the one to the other plainly in insects, plants, and land-shells, and more imperfectly in the higher animals; but also in them, if allowance be made for the greater variability in form in the higher animals under change of condition of life, and their distribution be examined in relation to the geographical epochs in which the different forms respectively came into being and most prevailed. The absence of particular mammals in a particular land cannot vitiate my theory, if the distribution of animals in it had been completed before the mammals appeared. For the better understanding of my argument I shall first state the results at which I have arrived. The position I am about to maintain then is, that, subject to modifications to be afterwards mentioned, all the Coleoptera in the world aro referable to one or other of threo great stirpes. These three no doubt originally sprung from one stirps, and ac- quired their distinguishing features by long-continued isolation from each other, combined with changes in their conditions of life. But now we have three, and only three, great strains, sometimes intermingling with each other, sometimes underlying or overlying each other, and sometimes developed into new forms, but always distinguishable and traceable to one or other of the three sources.. These are—l, the Indo-African stirps ; 2, the Brazilian stirps ; and 3, what, for want of a better name, I shall call the micro- typal stirps, in allusion to the general run of the species com- posing it being of a smaller size, or, more strictly speaking, not containing such large or conspicuous insects as the others. It 8 MR. A. MURRAY ON TIE GEOGRAPHICAL RELATIONS OF is not altogether a satisfactory name, because the stirps does con- tain some large species, and it is not peculiar to it to abound in -small ones. But, taken asa whole, its ingredients are smailer and more modest in appearance than those of the others. The fauna and flora of our own land may be taken as its type and standard. A like tripartite basis may be traced in every class of beings. It may happen, indeed, that one or other of them, as the Bra- zilian stirps in mammals (Edentata &c., for example), may have almost died out; in others some former stirps, extinct in all the rest, may have survived in some isolated part of the world (as plants in Australia) ; but, subject to such exceptional modifica- tions, the leading features of my proposition will be found gene- rally applicable to all. It does not come within the scope of my present paper to show more than its application to Coleoptera ; but I do not mean to deprive myself of the aid to be derived from the occurrence of a similar arrangement in other classcs of organized beings, whenever I find that my position needs strength- ening. In many points our materials for working out the sub- ject are so meagre that they require every collateral aid, and it is obvious that the more widely I can show the arrangement to apply, the more will my conclusions, as to their occurrence in the Coleoptera, be strengthened. The Indo-African stirps, as its name implies, inhabits Africa south of the Sahara, and India and China south of the Hima- layas, also the Malayan district, the Indian archipelago, and the New Guinea group. This range is less modified by the general introduction of foreign elements than that of the next stirps. The Brazilian stirps inhabits South and Central America east of the Andes, and north of the River Plate, and furnishes, moreover, a large share in the constitution of North America, but has algo received in return a very perceptible tinge from the microtypal stirps. In the microtypal stirps I include the fauna of Europe, Asia north of the Himalayas, astern North America, so far as not modified by the Brazilian element; and, what has less of this strain, the whole of North-west America, California, part of the Mexican fauna, Peru, Chili, the Argentine Republic south of ucuman, Patagonia, Tierra del Fuego, Polynesia, New Zealand, and Australia. When I first broached this view to one of my friends, I was THE CHIEF COLEOPTEROUS FAUNZ. 9 met by the exclamation, “ What! Australia and Europe the same! Nonsense: Australia, of all places, is the least like Europe; when you go there you pass into a wholly new country: everything is reversed there; the very leaves grow upside down; it is like visiting some great city of the dead !”’ I grant it in some things, but not in all. Before I have done, I trust to prove that, in conformity with the principle I started with (that we must not judge of the fauna of one class by the fauna of another), it is not so in Beetles. It must be remem- bered that the present flora of Australia once flourished in Europe. Professor Unger may have occasionally allowed his imagination too free a rein, and the determinations of many of the fossil species by him and Professor Heer on which he based his conclusions in his ‘New Holland in Europe’ may be in- sufficient or erroneous, but the fact will not be disputed that the Eocene Flora of Europe has many points of correlation with the present flora of Australia. The resemblance no longer exists in the living floras of the two countries; in Australia alone has the old flora survived. As regards insects, on the other hand, we know, from the re- searches of Heer and other naturalists, that the Beetle-fauna of Europe in the Miocene time was of the same type as the present Beetle-fauna of Europe and Asia. There are, unfortunately, no similar materials applicable to the Eocene epoch, nor has any one utilized the lesser materials that exist as Heer has done for the Miocene epoch; but from the fact that the insects derived from the still older beds of the Stonefield slate belong to the same stirps (I say so on my own authority and from personal examination), and that the whole of the Miocene materials yet made public, although drawn from several places and beds of dif- ferent age, all belong to one fauna, it seems probable that the Entomological Fauna of Europe in the Eocene age was the same as in the Miocene. It is an assumption, but not wholly without warrant; and starting from it, my hypotheses is that, like the Eocene Flora in Australia, it has survived in its Eocene form down to the present day; only it has done so more perfectly in England than Australia, while the flora has only done so at all in the latter. In short, I should hold that if the researches of Sir Charles Lyell should end in carrying back the antiquity of man to the Eocene time, and if the ghost of an Eocene naturalist were to be allowed to revisit the glimpses of the moon, he would 10 MR. A. MURRAY ON THD GEOGRAPHICAL RELATIONS OF find in Australia the type of both the plants and beetles very much as he left them. In Europe he would find only the beetles. Indeed I am strongly disposed to claim even a greater antiquity for our present Coleopterous fauna. Some may remember that when insect remains were first found in the coal-formations, the surprise was general among naturalists at finding them so small in size and so little different from those of the present day. They expected that they should have been as much beyond the existing type in size and splendour as the Megalicthys ex- eceds a Herring. Nature, according to the notions of those days, was in her youth in the Carboniferous epoch, and they ex- pected something of the extravagance of youth in her proceedings. It now seems more probable that the Coleopterous fauna there was the same in type then as now, and that it has continued so in the region I speak of for all the intervening period, in accord- ance with the rule already referred to, that the lower we descend in the scale of organization, the more persistent is the general character of the forms of which life is composed. It is not a reply to say that the Eocene flora, which has changed in Hurope, being lower in the scale of life than the fauna, should have been equally persistent. It is not lower in the scale of life than insects. They are notin the same scale at all. They are on two distinct and separate ladders; and the Hocene plants, which have changed, were high up on their ladder (the very mammals of vegetable life), while the Eocene Coleopterous fauna was low down on its. It is to be borne in mind, too, that we have every reason to believe that the changes in condition of life since the Eocene epoch have been much greater and more frequent in Europe than in Australia; and if the plants are ac- cepted as being more likely to change than the insects under altered conditions in life, it is in Europe rather than in Australia that a change in them was to be expected. Of course, in what I have been saying, and shall further say on this subject, I speak of the Coleopterous fauna of Australia as a whole. In one sense it cannot be disputed that it is dif- ferent from that of Hurope. The species are not the same, and there are a multitude of peculiar forms; but the type, especially of what I regard as the more important test-groups, such as the hunting unintroduceable species, is the same. The peculiar forms can almost always be traced back to enlargement or deve- lopment of some microtypal form. Putting aside such exceptions, THE CHIEF COLEOPTEROUS FAUNA. 11 the general facies is the same, and a large proportion of the genera are the same, and it will be still greater when we get rid of the feeling that the genera must necessarily be different, because they come from such a distant country. Some of the species are scarcely distinguishable from our own, and even the relative proportions of numbers of species and genera in different groups are the same. My conviction is, that there has been certainly one, possibly two, great continental routes of communication between the northern and southern hemispheres, both now lying buried in the ocean,—the one at the bottom of the Atlantic, the other in the depths of the Pacific; and I hope, from an examination of the traces left on the ruined piers which mark the course of these ancient viaducts, to show the course that they took and the inhabitants that used them. If any one, following in the steps of Sir Charles Lyell, ob- jects to such a wholesale erection of continents on the ground of their magnitude, I have only to remind them of the vast extent of land which has appeared above water since the Tertiary epoch. Some drying up of the ocean during that period no doubt has taken place, but nothing sufficient to account for the immense tracts of country which have become dry land; and it ‘is not a matter open to argument or discussion, but a mathema- tical necessity, that if land, previously below the water, comes above it, a corresponding quantity of land which was previously above it must then go below it. Let us now turn to the three great stirpes, and pass each of them in review, trace their course, and determine their limits. I shall begin with the microtypal stirps (with which we are most familiar). It is the most extensive of the whole, being distributed over the whole world, with the exception of the In- dian, African, and Brazilian regions; and even they, from va- rious exceptional causes, have a greater or less tinge of it in their faunas. It contains some minor faunas, and these, again, a number of subfaunas. The Europeo-Asiatic region is one of these minor faunas, and of it the Atlantic islands, the Me- diterranean, and the Mongolian are subfaunas. Taken as one fauna, the Europeo-Asiatic extends from the Azores east to Japan, the whole of that vast space being inhabited entirely by the same type and, for the most part, by the same species, a few only dropping off here and there, and being replaced by 12 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF others of the same general character. As to the Atlantic islands, the task is easy to decipher their relations; Mr. Wollaston has done it ready to my hand in his various admirable researches on their Coleoptera. It would be idle to vaunt the merits of his works to Fellows of the Linnean Society. Mr. Wollaston is one of our number, and we are entitled to regard his honours as gems in our own chaplet, if not laurels of our own growth. In interpreting the faunas of these islands, I have only to recapitu- late the results of his researches; on almost every point I arrive at the same conclusions that he has done. He has removed all possibility of doubt as to the general identity of the faunas of the northern groups with that of Europe, and notably with the Me- diterranean section of that fauna, or as to their individual identity with each other as members of one and the same subfauna. In the Madeiran group (see ‘Insecta Maderensia’ and ‘ Catalogue of Madeiran Coleoptera’) he showed that out of 580 species, 314 are species already known on the Continent of Europe; true, he considers (in which he goes further than I would) that so many as 120 of these had been imported by man, or otherwise found their way to the islands ; but, even after deducting these, he leaves 194 known European species aboriginally present, as against 266 endemic species. These endemic species, again, are all akin to the European forms, fit easily into their places among them, and all possess the facies of that fauna. I have already alluded to the want of reciprocity between Madeira and Europe in regard to any specialities they possess, and shall merely illus- trate that remark by noting the fact that, although Mr. Wol- laston credits Europe with a recent remittance of nearly the half of the European species, he acknowledges that no repayment in kind: has ever been made by Madeira, not a single example of any of its peculiar species having ever found its way to Hurope, ex- cept in an entomologist’s box ; and this, be it remembered, although the means of introduction have been at least as open on the return as on the outward voyage *. In the Canary Islands (see ‘ Catalogue of Canarian Coleoptera’) * I know it may be replied to this that an unusual proportion of the Madeiran endemic species are apterous; but this, even although it were a good answer, would only account for the deficiency of a proportion equivalent to the relative number of apterous, as against winged species; but it is not a good answer even as regards them ; for no one supposes that the introduction of species from the continent to Madeira has been by actual flight. It is floating wood and birds that are usually referred to as the vehicle or mode of transmission. TILE CHIEF COLEOPTEROUS FAUNZ. 13 Mr. Wollaston next found that out of a total of 930 species, 224 are identical with Madeiran species, and, notably, that the same peculiar types which gave to Madeira the character of a subfauna, are also present there in force. The Cape-Verde Islands tell the same tale. Previous to the appearance of Mr. Wollaston’s ‘Colcoptera Hespcridum,’ the usual belief among entomologists was, that the fauna of the Cape-Verde Islands partook more of that of the coast of Africa, nearest which they lic, than that of any other country. Mr. Wollaston has shown that this is a mistake. In his introductory remarks he says, “ Our recent explorations in the Cape Verdes have shown their Coleopterous population to be so far more than I had anticipated on the Canarian and Madeiran type, that I am anything but certain that it would not be more natural to regard the whole of these Atlantic islands as characterized by a single fauna—un- mistakably the same, even whilst necessarily differing as to many of its exact details (and through the fact of mere distance) in the more widely separated groups.” From my own materials I rather inclined to the more general notion, and I therefore care- fully tested Mr. Wollaston’s conclusions by his data, and the result fully corroborated his view. Out of 275 Cape-Verde species, 91 were common to the Canaries, and 81 to the Ma- deiran group. The African element proved slight, as Wollaston said, and such as might fairly enough be referred to chance in- troductions from the opposite coast of Africa. The European element continues, as before, the staple, and a new phase of the peculiar endemic subfauna of Madeira is also a characteristic element of its fauna. In support of the above statements, I shall merely specify one or two of the most striking of the types which are present in all the Atlantic-island groups under the same or similar forms. In Madeira the Heteromera are characterized by the presence of the endemic genera Hadrus and Hegeter, Hadrus having three spe- cies, Hegeter only one. In the Canaries, Hadrus has disappeared, but Hegeter has nineteen species, and in the Cape Verdes He- geter is reduced to one, but a new form, Oxycara, has taken its place with ten specics. In Madeira, the Curculionide are dis- tinguished by a profusion of Cossonide containing new genera and new species in a marked degree. The same prevails in all the islands ; so with Acalles, a small genus with few species in Europe, but with an especial redundancy in all the islands. -Ad/antus or 14 MQ. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF Laparocerus is another special new development confined to them, but present in most of them in greater or lesser numbers ; thus in Madeira there are thirteen species, in the Canaries thirty- five species, none in the Cape Verdes, where, however, Dinas, a new Brachyderidous insect, similar to it in appearance, comes either to take its place or that of Brachyderes, which is also found in some of the Atlantic groups. In the Clavicorns, the remarkable genus Tarphius, a consideration of whose relations would require space which cannot be given here, characterizes the Canaries and Madeira, as Attalus does in the Malacodermata. As to the Azores, Mr. Crotch has completed Wollaston’s work for him there. Asa matter of sentiment, one would have liked to have seen the whole finished by Mr. Wollaston himself, as he had done so much and so well; but the naturalist is rather un- grateful in this respect, and cares little how he gets his know- ledge, provided he does get it. Mr. Crotch’s contribution there- fore (Proc. Zool. Soc. 1867) is a welcome, as it is a trustworthy and careful, record of the Coleoptera of the Azores. His mate- rials are, indeed, far less complete than Wollaston’s in the other islands ; but although imperfect as regards proportions, they suf- ficiently reveal the character of the fauna. Mr. Crotch records - 213 species, of which 160 are European; and among those not European, he describes a Tarphius, a Laparocerus, an Attalus, an Acalles, and a new member of the Cossonide—all sufficient indications of the Azores being a member of the same system as the other Atlantic islands. How the European character of this general fauna is to be accounted for, except on the supposition of a former connexion of them all with Europe, and how the presence, of these special forms of the same subfauna in all the islands, and nowhere else, is to be accounted for except on the supposition that, after they were disunited from Europe, they were still united among themselves, it is for those who advocate the theory of dispersal by chance introductions to say. The Azores seem to occupy nearly the western extremity of this ancient land; not far beyond them a deop valley, the deepest part of the Atlantic, intervenes between them aud the coast of America. Up this the Gulf-stream scours, as it probably has done from early days far back in geological time ; and if there is any place in the world to which we might reasonably expect a few waifs and strays to be brought by currents, it would be the Azores ; and yet there are only three in this position, all Brazilian THE CHIEF COLEOPTEROUS FAUN. 15 and all insects which probably pass their larval state in timber ; they are an Elaterid (a species of Holus, 4. melliculus), which is found all along the coast of South America from Rio to Demerara, Monocrepidius posticus, another Elater which is otherwise confined to the La-Plata district in South America, and Teniotes scalaris, a Longicorn. In the other Atlantic islands there is only one such introduction, a North-American Longicorn (Clytus erythrocepha- lus), which has been found on the Salvages. The supposed in- troductions to the Azores from the European side of the Atlantic are, according to Mr. Crotch, much more numerous. He divides the 170 European species of the Azores into “two groups (70 possibly indigenous and 101 almost certainly introduced by colo- nists) ;’? and the mode of introduction assigned by him is (I know not on what grounds or with what primd facie probability) their importation in earth at the roots of garden-plants. Certainly in this case the operations of the chance-introduction theory (three species coming with the current against 101 against it) would seem to call for some apology or explanation ; but those who, like myself, reject that theory as capable of doing any thing more than furnishing accidental exceptions, will only see in the 101 supposed introductions (probably, but not necessarily, under deduction of a few cosmopolitan species) 101 natural denizens belonging to the microtypal stirps, and present in their natural capacity of legiti- mate descendants of the aboriginal heirs of the soil. The only remaining vestiges which may be supposed to have formed part of this ancient Atlantis are Ascension Island, St. Paul’s, St. Helena, and Tristan d’Acunha. Of these, St. Paul’s is, I believe, beyond its limits, and belongs to another fauna and another stirps, its fauna, so far as I know, being Brazilian ; but more information is still wanted regarding it. Ascension Island is a barren rock of recent formation, said to be almost without any fauna but what has been introduced by man within a hundred years or so; but it has never been tho- roughly examined by any competent naturalist. What we do know of it has been picked'up during brief flying visits by natu- ralists who, like Mr. Darwin, touched at its port, and did what they could in a limited time. The only animals recorded, so far as I know, are one Slug (Limas ascensionis), and, if we go by that, we must put it down as microtypal, and two Sea-shells (Zz- torina milaris and Nerita ascensionis), found by Mr. Cumming -on its shores. past 16 MN. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF St. Helena, that great puzzle of naturalists, is a crucial test to my hypothesis of a communication. between the northern and southern hemispheres by an Atlantic continent; if that link snaps, the whole chain will fall to the ground. It will, of course, not touch the evidence for a communication between the northern and southern hemispheres by the Pacific; but a microtypal St.- Helena fauna is vital to an Atlantic communication. I say that its fauna is certainly microtypal, and if so, almost necessarily a branch of the Atlantic type of that stirps; there is nothing else microtypal within reach for it to be attached to. Some three years ago Dr. Hooker gave an admirable lecture on oceanic islands *, in which he discussed the origin of the flora of St. He- lena, and on the whole seemed inclined to refer it to Africa. More in the spirit of “audi alteram partem” than from any settled conviction of my own, I wrote a reply +, in which I gave some reasons for thinking that it might more probabiy have been originally connected with and peopled from Europe, although also possibly connected at some period with Africa. More ma- ture consideration and subsequent researches have confirmed my opinion; and the following examination of the character of its plants and animals will show the grounds on which I rest it. In mammals, of course, nothing is to be expected. The only allusions to them that I can find is the statement { that in cutting away the lava at Ladder Hill, many feet below the surface, small bones have been found, apparently about the size of those of a rat, and more particularly a small rib-bone entirely covered with an incrustation of stalactite. In what manner these have originally come there must ever remain a mystery: there is but one probable mode of accounting for it, on the supposition that the animal might have crept into a crevice of the rock and there died ; for if a bed of lava in its liquid state had flowed over them, they would probably have been consumed, and would not have been found incrusted by stalactite. I find it also recorded in ‘Baynes’s Tour through St. Helena,’ p. 119 (1817), that at the beginning of this century the “ Manati or Manatee, Sea-cow or Sea-lion”’ existed in such numbers as to furnish employment for a fishery on it; and of course if the Manatee did exist there, it * Published in ‘ Gardeners’ Chronicle,’ January 1867. + Published in ‘ Gardeners’ Chronicle,’ February 1867. t See ‘ Proceed. Agri. and Hort. Soc. of St. Helena,’ 1826, p. 30. TILE CMWIEF COLEOPTEROUS FAUNZ, 17 would be almost proof positive of former continuity of land with some country where ‘the Manatee lived; for it is a herbivorous animal, and could neither have crossed from South America or Africa (where different species of Manatee still live) to St. He- lena as it now stands. But itis not a Manatee; the observer from whom Baynes quotes says it is undoubtedly the Sea-lion of Anson, and gives a description of it, which shows that it must have been a species of Scal, doubtless that which he supposes (the Sea-lion, or Phoca leonina), which also occurs at Tristan d’Acunha, and yields plenty of oil, Carmichael mentioning that one animal there will give 70 gallons. In ornithology there is, I believe, only one undoubted aboriginal land-bird, the Charadrius pecuarius of Temminck, a small Plover, named the Wire-bird (probably so called from its wire-like legs). It is so exceedingly close to the Cape C. Kitélitzii, that it has been confounded with it by ornithologists ; until lately it was shown by Mr. Layard to be distinct. The Plover (and like the others this Cape species) is a migratory bird, consequently it is not difficult to suppose that it might, in the course of its migrations, have been blown off from the coast of Africa to St. Helena. But after arriving there it must have become modified by the altered conditions of life into the C. pecuarius, and, among other modifi- cations, ceased to be migratory, for that bird is a constant resident in St. Helena all the year round. Baynes, in his ‘St. Helena,’ speaks of the Grenadier Grossbeak (Lovia orix) as an inhabitant, and says it is locally called the Wire-bird. That it is so called is certainly a mistake ; but if really an inhabitant of St. Helena, it does not seem a likely one to have been introduced ; and if not introduced, then it certainly is micro- typal, all the species of the genus being confined to the northern hemisphere. If the Canary has not been introduced, it would be another microtypal species, and more than that, a species belong- ing to the Atlantic subfauna. It is, however, said to have been introduced either intentionally or involuntarily by man. It is suited to the climate, and being a universal favourite of man, nothing seems more likely than that it should have been intro- duced by the escape of cage-birds ; but I cannot learn that the belicf rests on anything more than presumption and probability ; and it may be said, on the other side, that if St. elena was once a member of the Atlantic fauna, it is natural that it should occur there, and that, although so great 4 favourite and universal a com- LINN. PROC.—ZOOLOGY, von. XI. 2 18 MR. A. MURRAY ON THE GEOGRAPHIOAL RELATIONS OF panion of man, it is not found naturalized in any other islands, notwithstanding that hundreds exist equally well adapted to it and equally inhabited by canary-loving settlers ; a sufficient reply to which may be that there is no island exactly similarly situated in regard to man’s arrangements, and that other birds of a similar nature have actually been naturalized there, the Java Sparrow and Indian Haverdavats being known to have been introduced, and the fact and date of their introduction being on record. They have thriven as well as the Canary, all three being as common as Spar- rows. All, however, that I wish to do is to enter a caveat against taking the introduction of every microtypal species for granted. My own belief is that there are no aboriginal birds in St. THelena: perhaps its isolation was perfected before birds appeared in the lands with which it had previously been connected. Besides the above, the Common Fowl, the Guinea-fowl, the Pheasant, the Red-legged Partridge, the Peacock, and the Pigeon have all been intentionally introduced by man. Governor Beatson gives the names of a very few of the sea- birds found on or about the island; but their range is so wide that they can scarcely be cited as bearing on this inquiry. The Turtle is the only reptile mentioned as found at St. He- lena, but no freshwater reptiles or fishes are known; at the same time it must not be assumed that none can exist. Although there are no streams, there is water, and there are terrestrial and marsh-shells (Swecineas) ; and if the advocates of chance dispersal are correct in their reasoning, small fishes from distant lands might now and then be introduced by sea-birds. It is against their theory that they have not. Governor Beatson (loc. cit.) also gives a list of seventy marine fishes taken at St. Helena; but as they are all designated by their local names, it is of no scientific value. Passing it, we have a thoroughly scientific and dependable, although smaller, list in two parts by Dr. Giinther, in the ‘ Proc. Zool. Soc.’ 1868, p. 225, and 1869, p. 238, made from a collection sent by Mr. J. C. Melliss, a resident in St. Helena, to whom naturalists owe more than to any previous observer for information as to ils zoology. 1 will presently be seen that, besides the fishes, his collection of spiders and beetles supply the most important part of our material in these classes of animals. A copy of Dr. Giinther’s list will be found in the Appendix. Itis not to be expected that the marine fauna can be applied in the same way as the terrestrial fauna to THE CHIEF COLEOPLEROUS FAUNZ. 19 the elucidation of the distribution of animal life, nor does it fol- low that because we see certain great divisions in terrestrial dis- tribution, the same number and the same local distribution is to be found in them also; greater latitude and extent of range must be allowed to marine animals, and especially to fishes (the birds of the sea), than to land animals. The difference in their conditions of life in the sea is less than on land. Geological changes, such as the opening of the Isthmus of Panama and the Isthmus of Suez, have a more important bearing upon their distribution than upon those of land animals, inasmuch as the opening of a door to admit a new element is more important than shutting it after it has been already admitted. The knowledge that such events have taken place, however, enables us to reconcile the occurrence of marine animals in places otherwise difficult of explanation, as, for example, Saurus atlanticus, both at Madeiraand Zanzibar. With the help of such aids I by no means despair of being able to show that a similar distribution, in the main, exists in marine ani- mals to that in terrestrial ; not exactly placed alike, but proceeding from the same causes, and the deviations traceable to the different treatment, conditions and events to which they have been sub- jected. Their distribution must be studied (and happily we have the means of doing so) more in connexion with their geological history and the fossil remains of their ancestors. It is not my present business to attempt to do this; and I shall not do more than indicate the line of argument which such considerations, at first sight, seem. likely to lead to. Take the Sea-perches, the Percide (not merely the genus Serranus as now understood, but the group of allied genera of which it may be said to be the type), a group containing the first dozen species in Dr. Giinther’s list. Beginning in the Chalk with genera which are now all extinct, increasing in the Hocene, so that half of the genera now survive and are established in the Newer Tertiary, so that all the genera now existing were then present in England, that type would ap- pear to be properly microtypal. The sea-shells being for the most part dependent on the lands on whose shores they live, and therefore bound to them, are safer and more direct indications of the character of these lands than the fishes ; and their own stirps generally corresponds with that of the terrestrial inhabitants, although, from the causes already alluded to, they are sometimes exposed to diverging influences from which the latter are free. We have as yet, so far as I know, no list of Q* 20 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF the marine mollusca of St. Helena. Mr. Woodward in his manual mentions that Mr. Cuming collected sixteen species of sea-shells, _ of which seven are new. I cannot find that he has published these ; probably Mr. Lovell Reeve may have done so in his ‘ Conchologia Iconica,’ but I have not found them. Mr. Cuming’s collection, however, is now in the British Museum; and besides there are in it a number of other shells collected at St. Helena, amounting in all, with his, to about fifty species. I have gone hastily over these under the kind and able guidance of Mr. Baird ; and although I should be sorry to attempt anything like the determination of the species on the strength of such a hasty inspection, yet I think I may venture to give a list of the genera to which they belong, es- pecially as E had Mr. Baird at my elbow to advise me when I was making my notes upon them. I accordingly give a copy of my memoranda regarding them in the Appendix. On looking at this, I think conchologists cannot fail to be struck with the correspon- dence of the distribution of the species found there with the Co- leoptera belonging to my microtypal stirps. The range of many of them is put down in our books as world-wide, just in the same way as many of my microtypal genera of Coleoptera stand as cosmopolitan, merely because they are found at distant points of the microtypal range ; thus Zwcina is world-wide because it is found on the coasts of Europe, North America, the West Indies (a debateable frontier in all classes of animals), St. Helena, Tierra. del Fuego, New Zealand, and Japan (all microtypal), and its fossil distribution corresponds so far as we know it. So Mytilus is world-wide. Mr. Woodward's localities are “ world-wide— Ochotsk, Behring Sea, Russian Ice meer, Black Sea, Cape Horn, Cape, New Zealand.” Others, such as Venus, Venerupis, Corbula, &c., have the same microtypal habitats, with the addition of the Indian Ocean, whieh may have been reached through the Red Sea when the ports of the Isthmus of Suez were open. The Patellide, the Rissoide, Lntorina, Cecum, Cerithium, Chemnitzia, EHulima, Nassa, all occupy microtypal ground. It seems to me, too, that the others, which are more widely distributed, will be found to be of older geological date. The land mollusks are of course better authorities as to the character of the fauna of the island. We have, however, no list of them, although Mr. Benson has described some of the living and Edward Forbes some of the fossil species, and also made a few remarks on them in the Geological Society’s Journal, 1852, p. 197, THE CHIEF COLEOPTEROUS FAUNA. os 21: and elsewhere. I have therefore combined these, and shall ven- ture to add similar notes as to the genera anda few of the species. from the British-Museum collection to those which I have given’ of the marine species. With one exception, they are all Euro- pean-looking Helices, Bulimi, Pups, Succinee, and similar forms; . the exception is a large semifossil Bulimus (B. auris-vulpina, Reeve) which looks recent, but of which the animal has never been found; the nearest affinities of this species have been thought by some to be with Polynesia. The affinities of the other species have been thought by conchologists, I believe, to lean most to Chili; but this I apprehend to have arisen rather from a reluctance to look for this relationship in our own land. Divested of prejudice, it is difficult to conceive anything more close in appearance to the British species without being actually iden- tical than they are, and any greater resemblance to the Chilian species I believe to be impossible ; and if it did exist, it could not go for much, for the land-shells of Chili are microtypal too, many of the Helices and Bulimi being exceeding like those of Eurepe; and we all know that two things which are each equal to a third are equal to each other. Others have sought to ac- count for this close resemblance to our own species by supposing them to be modifications of species brought in the earth at the roots of plants from Britain. The occurrence, however, of so many other species in other classes like our aoe species seems fatal to this view. Of other sea animals, I have to mention two species of an An- nelid (Ditrupa), also of a northern type. There are four Crustaceans mentioned by Governor Beatson —Shrimps, Crawfish, Stumps, and Long-legs—which by their names and the character ascribed to them by Governor Beatson, viz. that they resemble our lobsters in taste and colour, suggest our northern species ; but in ignorance of what they really are, we must pass them by. The Rev. O. P. Cambridge has lately reported on a small col- lection of spiders made by Mr. T. J. Melliss, and described the new species in the Zoological Society’s ‘ Proceedings ; and he says that so far as so small a number of species (only twenty- two), of which nine were new, may justify a general remark upon the character of the Araneidea of St. Helena, it appears to bear a thoroughly European stamp, one alone belonging to any genus not indigenous to Kurope. Four, if not five of them have been 22 MR. A. MURRAY ON THE GEOGRAPHIOAL RELATIONS OF recorded as indigenous to Great Britain, three to Algeria, and three to Egypt. Among the new species Mr. Cambridge found “but little to denote a locality so near the tropics.”’ (See extract from Mr. Cambridge’s paper in the Appendix.) Mr. Cambridge also records two Scorpions from St. Helena in the same collection, Lychas maculatus, Koch, and L. americanus, Koch (American but easily introduced). The butterflies seem as badly represented as the birds; and I would recommend to the consideration of the advocates of intro- duction by chance dispersal the fact that the two classes of ani- mals best provided with means of dispersal are precisely those which, along with the mammals, are least represented. I can find no published notice of any Lepidoptera in St. Helena. No spe- cimens of any exist in the British Museum; and the solitary species that I can learn by inquiry to have been met with is the Cynthia Cardui*. Cynthia Oardui, I need scarcely say, is what is usually called a cosmopolitan species; but in very many instances it will now be found that what have been called cosmopolitan forms are only microtypal, that is, found in every part of the world but those parts of India, Africa, and Brazil to which the microtypal stirps had not had access. Until lately, our knowledge of the Beetles of St. Helena was limited to some twenty species or so. Mr. Wollaston has re- cently, however, considerably extended it, mainly through the researches of Mr. Melliss and Mr. Bewicke, and'has published a catalogue (see ‘Annals of Natural History,’ 1869 and 1870) in which seventy-five species are enumerated. His observation upon these is as follows. “If we exclude from consideration the twenty-six species (above alluded to) which have wnquestionably been brought into the island through the medium of commerce, and which enter into the fauna of nearly every civilized country, I need scarcely add that the St.-Helena list, as hitherto made known, * Prof. Westwood is my authority for this, and for the sake of preserving the information he gives, I quote what he says: “ As to the insects of St. Helena, I ‘am sorry to say that I can give you scarcely any information. In one of Dr. Burchell’s cabinets was a drawer filled with insects from that island, but it unfor- tunately had no door and had been left neglected. After Dr. Burchell’s death some wretched moths got into that particular drawer and devoured nearly everything. I kept all the fragments possible, and can determine some fourteen or fifteen spe- cies of common forms, Coccinella, Sepidiwm, Necrobia, Cynthia Cardui. It for- tunately happened that the type specimen of the curious Aplothorax Burchelli remained intact.” THE CHIEF COLEOPTEROUS FAUNZ. 23 possesses nothing whatever in common with those of the three sub-African archipelagos which lie further to the north—though the great development of the Curculionideous subfamily Cosso- nides is a remarkable fact whichis more or less conspicuous throughout the whole of them.’ In this judgment I cannot concur; the list seems to me brist- ling with Atlantic affinities and points of correspondence. It is one of the very few instances in which I do not go entirely along with my friend Mr. Wollaston’s conclusions ; and I believe the difference on this occasion arises chiefly from our looking at the subject from opposite stand-points. T am looking at it as part of a larger stirps, he as an independent object. I am anxious that he and the large number of readers whio, relying on his well-known judgment and acumen, will naturally accept his con- clusions as their own in fide parentum, should see that I have strong grounds for dissenting from him; and I have therefore given in the Appendix a copy of the list of species recorded by him distributed into the stirps and faunas to which I think they belong, with full notes containing my reasons for placing them as I have done whenever any doubt seems likely to exist about the matter. On referring to this, it will be seen that, according to my view, the seventy-six species as yet recorded as inhabiting St. Helena (whether by Mr. Wollaston or others) are to be accounted for as follows. There are :— 1. Of doubtful identity and uncertain eee 9 through the fault of the original describers ... 9. New endemic species which I have not seen either in nature or figured, and as to the afli- nities of which I am thus unable to form an "@)TEO)1), pejece Noda cadanoaepsngaebe seo! ede dou soupedeGe 3. Cosmopolitan or introduced, which, with two 3 exceptions, belong to the microtypal stirps ... } ; 17 Deducting these 17 from the total 76, there remain 59 belonging to the different stirps as follows :— South-American stirpS ......-..-2+-s1+6 1 Tndo-African stirpS ......-..ssseeereeees 2 Microtypal stirps, European branch... .56 — 59 24 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF Of 56 members of the European branch of the microtypal stirps, I find 12 which have no greater affinity for one part of the European fauna than another, 8 which are new, but whose general affinity lies with species characteristic of the Mediter- ranean subfauna, 40 whose affinity is nearest to the fauna of the other Atlantic islands, and 1 (Pristonychus complanatus) which is generally distributed in microtypal countries, and which has hence been supposed cosmopolitan, but which in reality has not been found out of the microtypal bounds. Much of the weight to be given to this apportionment of the elements of the isle must depend on the value of my reasons which are given in the notes to the list of species in the Appendix, and to these I must refer the entomological reader. I may only say here that the instances which have had most weight on my own mind, are, lst, the occurrence of a large Carabus (a hunting carnivo- rous genus limited to microtypal countries, which it would seem impossible to introduce except by continuity of dry land), which, according to the high authority of Prof. Lacordaire, has most affinity with species found in Syria (7. e. in the Mediter- ranean district with which the Atlantic Islands are otherwise most connected); 2nd, but of still more importance, the presence of species of genera which are particularly prominent or abundant in the other islands, as Calosoma, Bembidium, Lemophelus, Anobium, Opatrum, &c.; and 3rd, and of most importance of all, the pre- sence of new forms allied to species already known as character- istic of or confined to the other islands of the Atlantic, as dd- - croxylobius, Nesiotes, and Notioxenus, representing respectively the prevailing element of Cossonus, Acalles, and Atlantis in them. That a particular genus is represented (however critical this genus may be), when the representation is only by a single species, is not nearly so strong evidence of common origin as common exu- berance of some particular form in both faunas under comparison ; for the occurrence of a single species may be explained away when the presence of many defies dispute. And I rest as much on the occurrence of the typical character of facies as on actual identity of genus or species ; for in the development of new forms Nature often refuses to go by our generic characters, and produces some- thing exactly similar in appearance but with some deviation in what the systematist chooses to call important organs, a deviation which to his mind is fatal to generic identity, but to mine insignifi- cant in the face of persistence of facies. TIE CHIEF COLEOPTEROUS FAUNA. 25 So far as the fauna goes, therefore, I have little doubt that the majority of zoologists will agree with me in referring it to the Atlantic subfauna of the microtypal stirps. But when we come to the flora, we have new light thrown upon the subject. It is said to be rapidly losing its original features; when Burchell visited the island, it was still nearly in its natural condition, and out of 169 plants collected by him, 40 were endemic and very peculiar, and of the remainder, a considerable proportion seem to have been of European type. Dr. Hooker, in his lecture on Oceanic Floras, says of this, “ Dr. Burchell’s collection includes 169 flowering plants, but most unhappily he has not indicated which are bond fide natives and which have followed the track of man and animals introduced by him, and which have become quasi-indigenous or naturalized. Some years after Dr. Burchell’s visit, however, an eminent Indian botanist, Dr. Roxburgh, visited St. Helena, and drew up a catalogue of the indigenous, naturalized, and cultivated plants then existing, carefully indicating the truly indigenous ones that were then surviving.” This flora of Dr. Roxburgh’s, however, is imperfect, some of Dr. Burchell’s species (now in the Herbarium at Kew) not being included in it, pro- bably having become extinct in the interval between his and Bur- chell’s visit; and a strong desire is felt by those interested in the subject that a fresh flora of St. Helena should be published by some competent botanist. Dr. Hooker’s talents, position, and acquaintance with the subject point him out as the most fitting person to do so: and I trust the general wish that he may undertake it will lead to its own fulfilment. As our knowledge of the flora stands, however, I believe the actual facts which have been ascertained regarding it are that it contains, Ist, a considerable number of plants known to have been introduced from various countries, but chiefly from Europe ; 2nd, a considerable number of European species or genera which are not known to have been introduced, but which are taken for © granted to have been so on account of their European habitat ; 3rd, a small proportion (but still too large a proportion to be ac- counted for by chance dispersal), the affinities of which are clearly with the Cape flora. Dr. Hooker’s conclusion to this effect is thus stated in his lecture above referred to:—“ From such frag- mentary data it is difficult to form any exact conclusions as to the affinities of this flora; but I think it may be safely regarded as an African one, and characteristic of Southern extra-tropical Africa. 26 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF The genera Phylica, Pelargonium, Mesembryanthemum, Osteosper- mum, and Wahlenbergia are eminently characteristic of Southern extra-tropical Africa ; and I do not find amongst the others any in- dication of an American origin, except a plant referred to Physalis. The Ferns tell the same tale ; of twenty-six species, ten are abso- lutely peculiar ; all the rest are African, although some are also Indian and American.”’ On this sentence, while I implicitly accept its conclusions, I shall only remark :—1, that Mr. Baker, in his admirable paper on the geographical distribution of Ferns, seems to me to bea little more favourably disposed to America in his estimate of their relationship ; 2, that some of the African spe- cies, as Banksia and Protea, may have an Australian significance as well as an African—not that 1 think that either touches Dr. Hooker’s conclusion, but in trying to sum up impartially I do not wish to overlook any point; and 38, that there is besides what may be called an under layer peculiar to the island itself, and found nowhere else on the face of the globe, such as arboreal Composite (tree-daisies, as it were). Dr. Hooker regards these as too abnormal to have their affinities with the plants of neigh- bouring continents made out. I cannot think so if he will lend himself seriously to the work. The general result which I draw from the whole flora is, that we have here a compound flora certainly two deep, possibly three deep. We have, in the first place, I believe, a genuine natural Atlantic, that is, European flora; for in the face of the decided testimony to that effect given by the fauna I cannot accept Dr. Roxburgh’s conclusions as to the supposed introduction and natu- ralization of every species having a European habitat. If they can be proved to have been introduced, good and well; but I object to take the thing to be proved as part of the proof. And, in the next place, I believe we have the traces of an older African flora (why I call it older I will explain when I come to speak of an ancient connexion between Patagonia and South Africa) ; and I believe that both are due to actual continuity, however circuitous or interrupted, with the respective countries the impress of whose floras they bear. Before leaving St. Helena, I have just one other argument to adduce in support of its former connexion with the other Atlantic islands, and that is the fact, which has only recently been ascer- tained, or, at any rate, only recently laid down in our maps, that there is a long band of elevated submarine bottom running north THE CHIEF COLEOPTEROUS FAUNA. 27 from St. Helena to the Cape-Verde Islands, and embracing in its course Ascension Island and the shoal ground on the equator. The next trace of a microtypal element in the southern Atlantic is the island of Tristan d’Acunha; and in obedience to the natural train of thought, I shall begin with its flora, as I have just left that of St. Helena. I should have liked to have given in the Appendix a copy of the ‘ Flora St. Helenica,’ partly in illustration of what I have said regarding it, and also for the purpose of contrasting it with a similar list of the flora of Tristan d’Acunha given by Capt. Carmichael in the 12th volume of the ‘Transactions’ of this Society; for we find the same elements in both,—a mixture of European and African types in nearly the same proportion. The St.-Helena list is rather long, and I hope may soon be supplied by a better; the Tristan d’Acunha list is short, and an abstract of it may be convenient, and one is therefore given. The only shrubby plants in the island (trees there are none) are Phylica arborea, and either one or two species of Hmpetrum. Phylica is an African genus represented by two species in St. Helena; and its occurrence in both St. Helena and Tristan d’Acunha fur- nishes at least a presumption in favour of the two islands having once been in communication with each other and with the African continent. Empetrum (our Crowberry), on the other hand, is, as every one knows, a most characteristic type of the Scandinavian flora, and not less so of the Magellanic and Ant- arctic Flora generally. So is the genus Chenopodium, wild spe- cies of which also occur in both St. Helena and Tristan d’ Acunha. The genus Pelargonium has also species in both islands: it, not- withstanding the presence of a straggler in Syria, is unquestion- ably African, its species in that continent being numbered by hun- dreds. It seems of no consequence that, as Dr. Hooker informs me, the Tristan-d’Acunha species belongs to a different section from the St.-Helena species. We should have expected that they would be different; the greater the deviation the longer the probable period since they started from common parents, and the stronger the presumption in favour of my view of the connexion with Africa being very ancient. But what must strike every one most in running their eye over Capt. Car- michael’s list is the resemblance to our own flora. We there see Ranunculus, Rumex, Cardamine, Atriplex, Gnaphalium, oe Carex, and similar genera. 28 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF _ Capt. Carmichael does not say much about the insects, but what he does say tells the same tale as the plants,—“ Three small species of Curculio.”’ Thus we have, again, small Curculios, proba- bly similar to those which have given a character to the Coleoptera of the other islands ; with them four Phalenas, the old genus for the typical British Moths—a Hippobosca (qu’ allait-il faire dans cette galére, where were neither horses nor other land quadru- peds for them to feed on ?),—two species of Musca, and a Tipula. Of Crustaceans, an Oniseus, an Astacus, and a Cancer, all cha- racteristic types of the European fauna. Of the land-shells we may say the same; we know only two, both species of the genus Balea, a genus allied to Pupa, of which species have nowhere been met with elsewhere, except in Hungary, Norway, Porto Santo (one of the Madeiran group), and New Granada. The Norway species has also been found on the highest peak of Porto Santo, The only locality not entirely microtypal is New Granada; and of it the mountainous part is microtypal, the plain Brazilian. In which of these the Balea occurs I do not know; but the probability is in favour of the mountains, because the climate of the lower parts is so dangerous that it is almost entirely in the mountains that collecting has chiefly taken place. The sea- shells and other marine objects recorded by Capt. Carmichael all have the same microtypal tinge. Chzton, Cardiwm, Patella, Buccinum, Sepia, Echinus, and comillinee sound mar- vellously like the contents of one’s basket after a rummage along the coast in our own country. Have we now reached the southern limit of the ancient At- lantis? Is Tristan d’Acunha its outmost cape? Has it stretched for interminable space to the South Pole without leaving an in- dication of its existence? or has it trended off to the Falkland Islands and South Shetlands, and joined Tierra del Fuego, and possibly Patagonia? If there were no other way of account- ing for the microtypal character of the fauna and flora of South America south of the Plata, all to the south of it bemg micro- typal, one might feel disposed to assume that it did; and had we only the flora to go by, I should probably adopt that view, for we have in Tierra del Fuego and the other antarctic islands the very types of European plants that we have noted in Tristan a’ Acunha—Empetrums, Ranunculus, Cardamine, Wild Celery, &c. &e. But I shall presently show that there was another route by THE CHIEF COLEOPTEROUS FAUNA, ; 29: which a communication between the arctic and antarctic hemi- spheres was effected, and that the affinities of the Coleoptera of Tierra del Fuego and Patagonia rather point to that being the channel of communication so far as they were concerned. It may. probably have been the case that there was interrupted commu- nication between Tristan d’Acunha and these antarctic islands, which in their turn had interrupted communication between Cape Horn, New Zealand, and Australia. Leaving this question for future solution, I shall now revert to the European fauna, of which we have only touched on the most western limit, and trace it eastward. I have already said that the whole fauna from the Azores to Japan was one and the same. No better proof of this can be given than a comparison of the list of species from one end of the continent to the other. We have no complete lists of the Coleoptera all over the country, our lists of the east of Asia being comparatively imperfect, but they are still sufficient to illustrate the identity I desire to point out. We have a list of those found by Schrenck in Amourland and Eastern Siberia, made up by Motschoulsky, and published in Schrenck’s ‘ Reisen im Amurlande’*. We have also some simi- lar data regarding the Coleopterous fauna of South-east Siberia, collected by Raddé in his explorations; but this is very imperfect, and relates more to genera than species. Motschoulsky’s list of species found both in Amour and Eastern Siberia contains 810 species. The portion of these found in Hastern Siberia is not, however, so applicable to my present comparison as the list of species found in Amour, which extends to the extremest limit of Asia. The number of species from it, enumerated in the list above referred to, was 340; butafuller list was published afterwards by him f, which contained 564 species; and I have made it the basis of a Table, which will be found in the Appendix, from which the range of the species composing it can be ascertained. As it does not forward this inquiry to know what particular species are limited to Amour, I have left out all in that position, except when they represent a genus not otherwise present, when I give * T may here say, parenthentically, that Count Motschoulsky’s tendency was certainly not to diminish the number of new species, but rather to increase them, so that any insect that he admitted to be the same as one previously de- scribed may, without much doubt, be accepted as really such. + ‘Catalogue des Insectes rapportés des environs du Fl. Amour, depuis la Schitka jusqu’é Nikolaévsk, examinés et énumérés par V. Motschoulsky,’ Moscow, 1860. 30 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF one species of the genus to show that it is present. I have also’ added a few on the authority of Raddé and others from South- east Siberia. The total number in my list thus purged and augmented is 382; but adding the number of the endemical species, of which I have not given the names, we start with 608 species known to inhabit the extreme east of Siberia. Of these the numbers are, in Amour EO NER ce Se Mem oars a RU EY A MRL ate Sa lh 297 Western Siberia, or the districts of the Ural Mountains... 227 North and Mid-Europe, as distinguished from East or West Muroperi tie. sbi cote: ills aie une cme cre nn Meee 213 East Europe as represented by France and Belgium ...... 184, Site ds ON so ae ae, Real eel oe Le eR er eG area 133 he Mia deiras iwi ise noMane wants ek GuaLes arene ier Tere eee 10 Whe Azores ites ieee sete. Bene rst laine aide eet eat 7 To which I may add, in anticipation of what I must pre- sently say in speaking of North America,— On the western side of North America ...............-.20.0.0s 8 On the eastern side of North America .....................+5- 23 The details will be found in Table VII. in the Appendix. The diminution in identity of species as we go further from our starting-point (wherever we begin) and their replacement by new strains is, it will be seen, exceedingly gradual and equal, and the proportion of identical species persisting through the immense stretch of country embracing Asia and Europe very remarkable. But what is of still more importance in this inquiry is the identity of the genera. Using the word in its large sense, the same genera are spread over the whole region in question ; used in the more restricted sense, adopted by modern naturalists, a similar replacement of one form by another allied one, which we have seen occur in species, takes place also in these groups of species. Thus, in recording the species from the Amour, Motschoulsky has thought it necessary to propose a number of new genera for the new forms; and the proportion of these to the old genera found there was about a fifth, He records 239 genera, of which 85 are new. And, curiously enough, this is very close upon the numbers which Wollaston has turned out in his work at the other end of the string on the Coleoptera of Madeira; he records 286 genera, of which 44 are new. THE OHIEF COLEOPTEROUS FAUNZ. 3L The same fauna goes southwards through Mantchouria and Korea into China; and about Shanghai we get to the line where it meets the Indo-Malayan fauna. We have a tolerably fair (although far from complete) notion of the Coleopterous fauna of that part of China. Mr. J. C. Bowring procured important material from that quarter. Mr. W. W. Saunders has also made some of its species known ; and latterly Mr. C. W. Goodwin, Assistant Judge of the Consular Court of Shanghai, has sent some important collections made in the im- mediate vicinity of that city to one of our London entomologists, M. de Rivas, who, I trust, will ere long give to entomologists a catalogue of the species. In the meantime these materials (which I have had the advantage of studying) show that the Coleopterous fauna of Shanghai is a mixture of a few Indo- Malayan types (such as Copris molossus, Euchlora viridis, Cero- sterna punctata variety) with a mass of smaller species mainly belonging to the Europeo-Asiatic fauna; some identical with European species, the majority new species of the same type. A small collection of Coleoptera made by Dr. Collingwood at For- mosa, which he has been kind enough to show me, exhibits the same mixed fauna, and of nearly the same kind and proportions. An exactly similar intermixture occurs on the opposite coast of Japan; but what is most remarkable is, that although it occurs in the Beetles, Butterflies, Bugs, &c., it does not occur in the Hymenoptera. The great majority of the Beetles are of the Europeo-Asiatic type, and a certain proportion (as in Amour) are identical with, or only very slightly different from British species; the minority consists of species of the Indo-Malayan type, and indeed of the identical species which occur at Shanghai (Copris molossus and a variety of Cerosterna punctata, being two of the most prominent insects in both). But the Hymenopterous fauna is not of this mixed character; it is entirely Chinese. Mr. Frederick Smith, our first authority on the Hymenoptera, and who, from his position in the British Museum, has unusual opportunities of observing collections from all quarters, tells me that he has never seen a Hymenopterous insect from Japan of other than the Chinese type. It is the only class of insect, so far as I know, in which this deviation from the typical character ob- served in others occurs. Why should this be? Is there any peculiarity in the life of the Hymenoptera which can account for - it? The only one I know of is, that one large section of them 32 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF (the Bees) are dependent on flowers for subsistence in the larva state; and supposing the cold of the glacial epoch to stop the flowering of plants in Japan without killing the plants, the bees might be exterminated while the other classes still survived. Were Japan, therefore, separated from the mainland, so that on the restoration of a milder climate no fresh supply of species could be received from the north, and united to Southern China, so that it received its new inhabitants from it, and then finally separated from it as it now stands, we should perhaps have an explanation of the actual phenomena as regards bees; but there are other Hymenoptera to which this explanation will not apply, and further research may show that the exclusion of northern types is not so rigorous as at present appears. At any rate, it seems to me thatif the whole earth might be replenished by chance colonization, then the presence and absence of particular classes of insects in Japan is without explanation. I presume it will not be necessary for me to show that the same distribution prevails throughout Europe and Asia in every class of animals ; Dr. Sclater was the first to do so in the birds, I have elsewhere done the same for the mammals. Dr. Ginther has done it for reptiles, Gabriel Koch has done it for the Lepi- doptera, Meyen and Hooker for plants. In fact every person is at one upon it, each in his own speciality. The Europeo-Asiatic Beetle-fauna* does not stop even at Japan ; it passes over into North America by Behring’s Straits, or rather, I should say, it is found in North America on the other side of Behring’s Straits. In Russian America we have a fresh crop of Europeo-Asiatic forms, genera and species ; and here another noteworthy circumstance presents itself. It is generally taken for granted that there is a uniform homogeneous arctic fauna which extends all round the arctic circle. It is so, and it is not so. It is so on the large scale, but not so on the small. The arctic fauna is subject to the laws of spreading by continuity and stoppage by barriers just the same as any other fauna. I have elsewhere endeavoured to show that the mamma- lian fauna of Greenland is Europeo-arctic as distinguished from Americano-arctic. I maintain that the homogeneity of a fauna * T was unable, in my ‘Geographical Distribution of Mammals,’ to adopt Dr. Sclater’s terminology of Palwarctic, Neoarctic, &c., because we did not agree in the extent and limits of our regions; and now, of course, in this paper I can still Jess do so, as a principal effect of my hypothesis, if it be sound, must be to still further break down their limits and destroy their solidity. THE CHIEF COLEOPTEROUS FAUNR. 33 depends on other causes than uniformity of condition of life within its limits. I cannot doubt that if there had been an isolated communication between the Indo-African districts and the North Pole, we should there have had a fauna related to and developed out of that fauna, and wholly distinct from the other faunas of the arctic regions. It is continuity of soil or freedom of intercommunication which has produced the present uniformity of fauna in the arctic regions; but where minor inter- ruptions exist, or old barriers or conditions equivalent to a bar- rier formerly existed, there are also subdivisions in the character of the fauna, and in the position of these minor divisions we see the operation of these laws and are able to trace the existence and former position of the barriers. Thus we find two minor subfaunas in Arctic America, an eastern and a western one. Two causes may have produced these. One of these may have been the sea which, it can scarcely be doubted, formerly existed be- tween the Gulf of Mexico and the Polar Sea, in the line of the Missouri and Mackenzie rivers ; another may have been that the ground now occupied, by one of these subfaunas was under water at a later period than the other, so that it was peopled at a dif- ferent date from it. Probably both contributed to produce the present arrangement of the subfaunas to the east and west of the Mackenzie River. That there was a barrier there, and that that side was still supplied with the same general type (though with minor deviations), is to be explained by their having re- ceived their species from the same general stock, but coming to it from different directions, the one from the east, the other from the west. That the minor differences to which I allude are, in the case of North America, to be referred to this cause, and not to mere gradual increase of variation arising from in- crease of distance, seems to be a legitimate inference from the fact that while the whole of the north of North America, without exception, belongs to the Europeo-Asiatic type, there are a number of European genera which occur in North-east America, and not in North-west, and a few which occur in North-west, and not in North-east America. In the Appendix I have given a list of genera of Coleoptera which inhabit both sides of the Pacific, and do not occur in the Atlantic States of the American Continent, and also of a list of some species of other genera, similarly distributed. These are almost literally taken from my friend Dr. Leconte’s Reports in LINN. PROC.—ZOOLOGY, VOL. XI. 3 34 Mk, A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF the ‘ Pacific Railroad Reports, 47th parallel, and on the Co- leoptera of Kansas and Eastern New Mexico, as verbally cor- rected by him for me down to the most recent date, and only one or two being added by myself. Some of the genera or species in these lists may yet be met with in Eastern America; but after making allowance for this, enough would seem still to remain to warrant us in holding that a certain proportion of these must have reached America vid Siberia, and that, in like manner, most of those in the Eastern North Atlantic States have probably originally come vid Europe and Greenland. North America has no special fauna or flora of its own. That which it has is a mixture of the microtypal and Brazilian stirpes intermingled with fresh importations of different dates, and mo- dified by the advance and retreat of the glacial epoch; but, on the whole, the preponderating element in its fauna is the micro- typal. What I am now pointing out with regard to Beetles may be traced to a greater or less extent in every branch of zoo- logy and botany. I could go over each, pointing this out; but I will wait until the fact is disputed. Its origin is of very old date, the elements now respectively found in Europe and America haying been already settled in each country before the Miocene time. Professor Heer’s admirable papers on European fossil Tertiary insects give us the means of inferring this, and at the same time furnish arguments against his and Professor Unger’s scheme of the Miocene Atlantis, which they held to have united Europe to America in the line of the Azores, and which, they think, served as a, bridge for the intercommunication of the plants and animals in the two continents. That there was formerly a continent in the Atlantic is, I think, proved to de- monstration by the facts already mentioned regarding the faunas of the Atlantic Islands. But that it reached America is gain- said not only by the facts adduced in Professor Oliver’s able paper on the subject, published in the ‘ Natural History Review,’ and by those of other able naturalists, a réswmé of which I have already given elsewhere (‘ Geographical Distribution of Mam- mals’), but by the examination of Heer’s lists of species, to which I am about to refer.. If the reader will turn to the Ap- pendix, he will find in one of the Tables a list of all the genera of Professor Heer’s ‘Miocene European Coleoptera,’ with the ex- ception of a very few, which he could not refer to known genera, and which I have omitted. In that list I have noted in columns TIE CUIEF COLEOPTEROUS FAUNZ. 35 . opposite to each genus the different countries in which they are now found; and the result shows, first, that all these Mio- cene genera, excepting such as are universally distributed, are now confined to my microtypal regions; and, secondly, as re- gards Europe and America, that among them are plenty of genera which now inhabit both Europe and America, but not one that now inhabits America and does not inhabit Europe, while there are a few well marked and characterized forms, as, for example, Pelobius, Capnodis, Microzowm, and some others of less marked distinction, as Perotis and Hurythyrea, which now inhabit Europe and do not inhabit America. This is as strong evidence in kind (I do not dispute that it might be stronger in quantity, that is in number of forms), but it is as strong in kind as a fossil collection from one country alone could give, that the same distribution which prevails in these two continents at the present time, prevailed already in the Miocene epoch. The genera which are now peculiar to Europe were then peculiar to it, and, consequently, the inference is strong that no communi- cation between the two countries has ever existed since the days when these fossil insects were in life. If we had an American collection of the same age in which types now peculiar to America were found, the evidence would of course be still stronger, but it would be repetition of what we have already observed in Europe. The same relations between the Ameri- can and European Miocene species are to be found in other classes. I give a somewhat less elaborate (as regards num- ber of regions) list of the distribution of all the other existing genera of insects recorded by Heer and Krantz, not only for its bearing on this inquiry, but in the expectation that it may be convenient and useful to other students of geographical distribu- tion to have such a list at hand. The Hemiptera have been gone over for me by Mr. Dallas, our first authority on that branch, and the table and notes embrace the information derived from him, although he is not responsible for it all, his attention having been only specially requested to Europe and America. In that table it will be seen that the Miocene genera Prostemma, Nepa, and Diplonychus, and in the Homoptera the genus Tettigometra, are now found in Europe and not in America, and that one or two instances where the reverse seems to be the case are due to modern alterations on the genera used by Heer. In the Diptera it will be seen that all the genera are both American and’ European. 3% 36. MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF The above are not: the only points in which Professor Heer’s Miocene species lend important help to the student of geogra- phical distribution.. They are of essential assistance in deter- mining the southern limit of the microtypal stirps in Europe and Asia, and whether some of the forms which are found in the southern part of the European range really belong to it, or are immigrants from the African or Indian region lying to its south. The South-European fauna is composed partly of the same species as that of the districts more to the north, partly of distinct species of the same genera, and partly of what may be regarded as modified forms of the same general stock, but having a consider- able effect in altering the facies of the fauna. Besides these, there are a few (perhaps in all not more than ten or twelve) species which have probably sprung from the African stirps, and established themselves in Europe by immigration. The southern limits of the fauna of this region, which extends along the bed of the Sahara onwards to the Caspian and Mongolian Steppes, are the deserts of the Sahara, which cut it off from Europe, and the Himalayan range, which divides it from India and China. As regards the Sahara, it is its southern border which is the limit. Its bed seems to have been raised by a force operating from the north. The strata, abutting on the Atlas mountain-range, rest inclined on its flanks as if tilted up by it. The effect of this elevating force operating in the north would, of course, be to raise the part of the Sahara nearest it first out of the water; the last vestige of the sea would be at the south, consequently the bed to the north would be first colonized, and it could only be so from the north. The facts of geographical distribution quite cor- respond with this view. The fauna and flora of the desert is Mediterranean, not Senegalese. Returning to the Asiatic terminus of the microtypal stirps, let us now endeavour to trace its further course. The genus Blaps, which is a characteristic feature in the Coleopterous fauna of Central Asia, will furnish us with the means. It may be taken as a representative case applicable to other species also, although it certainly is the most striking instance which occurs to me. Up- wards of 100 different species of Blaps, out of a total of about 150, have been described as inhabiting the country between Southern Russia, Mongolia, and Mantchouria. Now if we cross to Cali- fornia in continuation of the same line we have not Blaps, but we have Blaps’s brother, and he has been a twin. We have THE CHIEF COLEOPTEROUS FAUNZ. 37 Lleodes, its perfect counterpart and representative ; and it is to be observed that while the facies of the species actually inhabit- ing California is entirely that of Blaps, a number of species which are found in Kansas and on the eastern flanks of the Rocky Mountains have a somewhat different facies; and I should add that the supposition that these are stragglers from the Californian shores is strengthened by the fact that the genus does not occur to the east of the Missouri: other Hete- romerous forms, reminding us of Mediterranean and Asiatic species, occur in California, and the whole of the north-west of America has a greater prepouderance of the microtypal stirps than perhaps occurs east of the Rocky Mountains. The Brazilian ele- ment is less sensibly present, such Brazilian genera as Passalus, Dynastes, Monocrepidius, Macrodactylus, Dichelonycha, Phaneus, Gymnetis, &c. being absent in California, although present in the Eastern States. M. Candéze, in his work on the Elaterids, notices that Meristhus scrobinula is found both in Mexico and China, and adds that he has found other species common to these countries, notwithstanding their distance from each other (Candéze, ‘ Elateride,’ i. 165). Other facts in other branches of natural history lend strength to the idea of a former communica- tion having existed between Asia and California. For example, in Mammals, there is a peculiar genus of Moles, Urotrichus, which has not been met with anywhere but in Japan and Cali- fornia. In plants, the botanist will remember that the Co- niferous subgenus Pseudostrobus, so abundant in Mexico and California, in the Oid World reappears in Japan, and only there. ‘The Menzies and Douglas type of Spruces does the same, species almost identical with them occurring in Japan. The Chamecy- paris of California is only another name for the Retinospora of Japan; and among herbaceous plants similar relations can be pointed out. In the Sandwich Islands, again (so far as we know, which is not so much as we wish), which from their position may probably have been part of any northern land which for- merly existed in the Pacific, as well as, at some period antecedent or subsequent, a part of Polynesia, the same character of fauna is present. Among the Lepidoptera we have Sphinx cingulata, Linn., or what is scarcely distinguishable from it, it in its turn being the scarcely distinguishable American representative of our own British Sphinx convolvuli. 'The only Coleopterous genera which I know from them are Anchomenus, Colymbetes, Agabus, Hydrobius, Hete- 38 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF rophaga, and Dryophthorus (Calandra), all of which are certainly microty pal. Next step to the south of California comes Mexico. It also is largely supplied with Hleodes; and although some of the finest and showiest non-microtypal Coleoptera in the whole world come from Mexico, they have no bearing on this part of my inquiry ; for they come from parts of Mexico which are in direct communi- cation with another stirps, the rich Coleopterous fauna of Brazil and Venezuela; and the vast multitude of small European-looking species which occur on the high lands and western side is quite sufficient for my purpose. The collections made by Truqui in Mexico show this thoroughly microtypal character im a very marked way, Staphylinidous genera, such as Falagria, Homa- lota, &c., abounding. Mexico, being a sort of halfway house be- tween Europe and Australia, might be expected to contain species both from the north and the south which have got thus far. Eleodes is an instance of this from the north, Philonthis another ; both reach as far as Chili, but not into Australia. Zopherus, on the other hand, is an instance of a species which occurs in Australia, and runs up into Mexico, where it is in strength, and goes even alittle further. Mexico may, indeed, have been its start- ing-point, but the connexions and relations of it and the allied genus Vosodendron decidedly indicate a separation between the eastern and western type of both; and the western type extends into Australia and New Caledonia. Between Mexico and Peru, west of the Andes, there is a con- siderable space, as to which more information must be obtained in every branch of natural history before we can satisfactorily dispose of this question. There have been many collectors in it, but they have usually hurried to the interior and across to Columbia and New Granada; and I have seen no coast-collection of Coleoptera, nor do I know of any published lists. From the mountains them- selves we have, however, received very considerable collections. Thanks to Professor Jameson, of Quito, we have a fair knowledge of the Coleopterous fauna of its neighbourhood. That of Bogota also is pretty well known. J'rom these I can say that they con- sist of a mixture of microtypal with Columbian forms, in which the Columbian predominate; but the microtypal is represented by undoubted members of that stirps, such as Graptodera, Philonthus, small Marpalide, &c. Of other classes of animals the birds are best known, through the exertions of Messrs. Fraser and Salvin ; THE CHIEF COLEOPTEROUS FAUNZ. 39 and, as 1t seems to me, there is a similar mixture of stirps in them. In such an inquiry as this, however, birds would require a special examination for themselves, their power of flight, and, still more, their migratory instincts, complicate their distribution so seriously. To do so fairly, the main distribution would require to be taken, in all doubtful cases, as the test of the stirps, leaving exceptional deviations out of view, whether they can be accounted for by exceptional causes or not. . Passing southwards to Peru and Chili, the number of Europco- Asiatic genera diminishes, but the general facies still remains. The Chilian species in many cases belong to European genera, and the general facies is of the same character. -Blaps still shows itself, only it has now passed out of the form of Eleodes into that of the smaller Nycterinus. The genus Carabus, which was lost in Mexico, has here retained its footing ; it is found in great beauty in the Chilian Andes, although very limited in number of species. Carabus is a genus almost entirely confined to Europe, Asia, and North America. Africa proper has it not; India has it not ; and, although it goes against my argument, I must in honesty add Mexico appears not to have it. St. Helena, the Chilian Andes, and Australia are the only places in the southern hemisphere where it occurs. In Australia the genus has undergone some mo- dification (into Pamborus), and in St. Helena (nto Haplothoraz), but still true scions of the Carabi, and bearing all their facies. The Feroniade, too, which form a very characteristic element in the European and American faunas, are fairly represented in Chili, strong in Australia, and absent from Brazil, Africa, and India, except in places which of themselves suggest that they are emigrants from over the border. Such are the species in South America from the mountainous parts of Columbia, or in India from both sides of the Himalayas; Pristonychus complanatus, a European species, seems to beat all others in the possession of “an undergoing stomach to endure whatever may ensue.” It occurs in Chili and also in the Canaries and in St. Helena. An- other somewhat remarkable form is the genus Thalassobius, bee- tles which live under high-water mark; it belongs to the Tre- chide, which seem peculiarly adapted for trying strange modes of life, and peculiarly open to the impression of altered circum- stances in them, turning into Anophthalmi of various kinds in dark caverns, into Apus and Thalassobius under the sea. pus is the form they have taken under high-water mark on the coasts 40 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF of Europe, Thalassophilus at Madeira, Thalassobius at Chili; and the late Mr. Wm. Sharpe Macleay informed me, in a letter written not long before his death, that he had found a similar species on the shores of Australia. Not then having my eyes open to the true significance of the occurrence of those species in these localities, I considered that probably they would be found on all coasts. Ido not expect thisnow. I imagine they will be found confined to the coasts of the lands to which my microtypal stirps extends; and, in point of fact, they have not as yet been found anywhere else. The distribution of the blind-cave Coleoptera is very remark- able. In the caves where they occur in Europe (chiefly in Car- niola, Hungary, Corsica, and the Pyrenees) almost every new cave produces a new species closely allied to, but distinct from, those in the nearest caves; but more remarkable still, the Mam- moth Cave of Kentucky produces a species of Anophthalmus so close to the Carniolan species that it is only on examination that one sees they are distinct. The Anophthalmi and their allies are carnivorous, hunting beetles, and, as I have just said, their parent type seems to be Zrechus; but the same thing occurs with another totally different type, Adelops, a clavicorn allied to Catops. Not only in the different caverns and also under moss and in dark places do different species of this occur, but again in the Mam- moth Cave of Kentucky it reappears side by side with Anophthal- mus in an all but identical form there. And here, while upon the cave-insects, I may remind the reader of the blind Reptilia and Crustacea of which allied forms occur in the European and American caves; and I would also draw their attention to a lately described form of cave-locustrian which has a distribution still more in accordance with the range of my microtypal stirps. One species occurs in caves in Europe, another in America, and a third in a limestone cave at Colling- wood, Middle Island, New Zealand. They were at first described under different generic names, it being supposed, probably from the distance of their localities, that they must be distinct ; but Mr. Scudder,the eminent American orthopterist,has shown that all three belong to one genus, which he hasnamed Hadenecus. Although they inhabit the deepest parts of the caves, they are not blind, but have the long legs which seem characteristic of the Anoph- thalmi and Cave-Araneide. In the Elateride the characters are slender and often artificial, and so not well adapted for the elimination of questions of geo- VTE CHIEF COLEOPTEROUS FAUNZ. 41. graphical distribution ; but even in them we can satisfactorily es- tablish the presence of the microtypal element in microtypal lands | and its absence elsewhere. Take the genus Elater proper. In it, of fifty-three species, twenty-three occur in the Europeo-Asiatic district, twenty-five in North America east of the Rocky Moun- . tams, two in New Holland*, and if we unite to it the genus Grammophorus, which has quite the facies of Hlater and stands next to it, we must add four from Chili. In the Buprestide the genus Stigmodera is often quoted as a striking illustration of affinity of animal life between Chili and Australia. It is impossible to dispute the absolute identity of their type; they do not, however, pass further to the north than Peru. Anthaxia is another type whose distribution corroborates my hypothesis. It is all but absent from Africa, India, and Brazil, or only very sparingly, and not very characteristically, represented by one or two species at the Cape of Good Hope or in the Malayan region; but in Chili it is so identical in appearance with our European species, that I remember when I first got some Chilian species I put them aside as obviously ticketed with an erroneous habitat. They also occur in Australia, although the species there are not so absolutely European in appearance. A not less striking resemblance between Chilian and European species occurs i a heteromerous genus from Mendoza, at the eastern foot of the Andes (Cacicus americanus), which is so exactly a large counterpart of Hlenophorus collaris from the Mediterra- nean, that I hold it to be perfectly certain that if both had been found in the same locality, only one genus would have been made for both. It is an out-of-the-way-looking genus, and no other example of the form occurs anywhere else on the face of the earth, so far as is yet known. The Scauride present similar South-of- Europe resemblances. The Gallerucide, a family which is represented by different forms in the different regions where it occurs, are represented in the Huropeo-Asiatic regions by the Halticidw. These are very numerous also in Chili. The genus Lithonoma, of which only two species have hitherto been described (one from Spain), re- appears in Chili, from whence I have received a species not yet described. In Cryptocephalus again, although the type leaves very * One is recorded as having come from the Kast Indies, without more precise indication ; but as that word generally includes the Himalayas, which are half Kuropeo-Asiatic, the locality cannot be counted either way. 42, MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF little room for change in appearance, there isa certain difference be- tween the Indo-African species and the microtypal. Brazil has only two or three of its own, and they have, to all appearance, been derived from North America or the west of the Andes. And there is again a difference between the European microtypal and the North-American microtypal, the latter having a facies of their own, which is shared by the Chilian and Peruvian species, and also in a less degree by the Australian. In the Casside we have the well-marked North-American genus Porphyraspis running down into Chili. In the Coccinellide, the Hippodamias (with the exception of one straggler in the Brazi- lian region, and one or two on its borders near Quito and Bogota) are entirely confined to the microtypal range, Chili being its southern limit, but it has not been met with in Australia. Coc- cinella proper, however, which has a similar range to Hippodamia, occurs there, and one or two stragglers have also found their way to the Cape, and one (C. transversalis) to the Malayan region. The microtypal stirps in the southern extremity of South Ame- rica is divided into two subfaunas by the Andes: that on their western flank is merely a continuation of the fauna of Western Peru; that on the eastern flank is cut abruptly off on its northern margin by the river Plata, where it meets the Brazilian type. The demonstrated history of this country sufficiently explains this distribution. Mr. Darwin in a few lines tells it thus: “ The landscape has one character from the Strait of Magellan along the whole eastern coast of Patagonia to the Rio Colorado ; and it appears that the same kind of country extends northerly in a sweeping line as far as San Luis, and perhaps even further. To the eastward of this line lies the basin of the comparatively damp and green plains of Buenos Ayres. The former country, in- cluding the sterile traversia of Mendoza and Patagonia, consists of a bed of shingle worn smooth and accumulated by the waves of a former sea; while the formation of the Pampas (plains covered by thistles, clover, and grass) is due to the estuary mud of the Plata deposited under a different condition of circumstances.” (Darwin, Journal, p. 402.) In those days the water came quite up to the mountains on the western as well as the eastern side; for we learn from the same source that “the valleys in the Cor- dillera are filled with an immense thickness of stratified allu- vium, which in all probability was accumulated at the bottoms of deep arms of the sea, which, running from the inland basin, pene- THE OHTIEF COLEOPTEROUS FAUNZ. - 43 trated to the axis of the Cordillera in a similar manner to what now happens in the southern part of the same great range.” There seems no reason to doubt that Patagonia and Chili were both supplied with their present faunas and floras from the main microtypal stock on the Andes. It is natural that from them the newly exposed sea-bottoms should have been peopled as they ap- peared, and quite in accordance with this that the stock flowing off to the right hand and the left should, while retaining a com- mon character, have each respectively minor peculiarities. This is what we should expect, and this is what we find. In both we find the same forms of microtypal Carabide, Cnemacanthus, Har- palus, Antarctia, &c., the same modifications of Heteromera, as Nyctelia, Cardiagenius, Praocis, &c.; and in both the fauna, as a whole, is remarkably scanty. In Patagonia, however, there occur one or two forms whose presence it is not easy to account for. The genus Eucranium occurs, not on the desert-plains between the mountains and the sea, but at Mendoza, at the foot of the eastern flank of the Andes, where the plains begin to rise into the desert base of the moun- tains; for notwithstanding the-advantage of water from their snowy peaks, the coarser shingle at the base of the mountains maintains its sterility equally with that of the less-watered finer shingle at a greater distance from them. Now Eucranium is un- doubtedly the representative of the Caffrarian genus Pachysoma, which is one of the Ateuchida, or pill-rolling beetles of Africa and India, the ancient Scarabeus of the Egyptians; and if this were a solitary case, I might perhaps have tried to get over it by arguing that although the Scarabcus is certainly an Indo-African form (being found both in India and Africa, and in prepondera- ting numbers in Africa and all over Africa), it might yet have originally been microtypal because it is found in the Mediterra- nean district, not only in Egypt, Algeria, and Barbary, but also in Italy, Greece, &c., and from thence might have extended into Africa. But against this is the fact that Ateuchus is not found in Heer’s lists of Miocene species, although Gymnopleurus, another pill-rolling beetle, whose distribution is similar to that of Ateuchus, is recorded there. The latter fact may be only an earlier instance of what has taken place in Ateuchus, or it may refer to some more ancient state of things; for all Coleoptera have no doubt been originally connected: but the connexion of Ateuchus with Europe is not the immediate point ; it is the connexion between one of its A+: MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF South-African peculiar forms and a closely allied form in Patagonia. Any doubt, however, that I might feel vanishes when I find other African forms or their representatives in Patagonia. In the Straits of Magellan has been found MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF dopterous insect in existence). It is an unusual thing at any time to meet with a gay-coloured moth ; but one with metallic brilli- ancy is still rarer; Urania exceeds any thing I know in this respect ; and it stands per se, nothing else like it in any other genus. Ofthis genus there are six species—one in Madagascar (Urania Ripheus), and the other five within the range of the Brazilian fauna, viz. two in Brazil (on the Amazons), another in Venezuela, one in Cuba, and one in Jamaica. Stelidota octomaculata is an example of affinity with North America. The mammal Solenodon of the West Indies has been claimed as allied to the Madagascar Gentetes (this, I think, on insufficient grounds); but Madagascar is the only place where (with one exception) Iguanoid lizards oc- cur out of America, and the only place in the Old World which furnishes examples of the American Colubrine forms Xiphosoma and Heteroda. The Lemurs, too, as I have elsewhere argued, are perhaps more nearly related to the Opossums or Squirrels than to the Monkeys; and if it be to the Opossums, that would be a link the more with Brazil. It is to be observed that all these afti- nities are confined to Madagasear and do not touch South Africa. According to my views, they are insoluble except by the supposi- tion of a dry-land communication between Madagascar and South America. My conjecture is that when the communication be- tween Patagonia (the penultimate) and the Cape was interrupted by the sinking of the land, all the land did not sink. The ground now occupied by Patagonia did sink; the land next Africa also sunk, but a mountain-range survived running from Cape Frio (Rio Janeiro) obliquely across the Atlantic to a point a little to the south of the Cape of Good Hope, there rounding the Cape and running up to Madagascar exactly in the shortest line that a ship could sail directly from Rio Janeiro to Madagascar. It may seem too child-like and direct to the purpose to propose such a route; I felt it so until I studied the sea-bottom, when I found that there a broad raised ridge does run along the bottom of the sea exactly in the direction [ have laid it down. I was not aware of this until I saw it so mapped in a map of the bed of the Atlantic in Mr. Keith’s Johnston’s New Physical Atlas. No one will dispute the importance of the configuration of the bottom of the sea as an indication of the line of ancient conti- nents; and on the strength of this and of the fauna of Madagascar, I think I have very fair grounds on which to base my hypothesis. Being a ridge, it would continue so when above water, and not THE CHIEF COLEOPTEROUS FAUNA. 69 appear as a plain like the desert Patagonian junction, but a moun- tainous or hilly country which would be watered by streams, clothed with forests and fitted for the habitation of such sparkling creatures as the Urania; for we see that nature always assimilates the aspect of the inhabitants of a country more or less to its pre- vailing hue; and where forests and flowers and dewdrops abound, there she clothes them in her most gorgeous robes. It may be that the last scene of all of this strange eventful history, prior to the appearance of the land as it now stands, was the extension of the microtypal regions of Cape Horn out to Kerguelenland, whereby the antarctic islands had already re- ceived their present flora, an extension which must have sub- sisted until a comparatively recent period, at least subsequent to the glacial epoch; otherwise I do not see how these islands could have been redintegrated in the possession of their flora after the retreat of the ice. As regards the bridge or range to Madagascar, that must have been its last scene; for otherwise we should not have it preserving its position and contour at the bottom of the sea. Africa itself is not difficult to read. Subject to the modi- fications of which I have been speaking, the whole of the eastern half of the continent is one broad band composed of one fauna. Lay a parallel ruler on the map, with one limb along the east coast and the other limb drawn back as far west as Congo on the west coast, and you have the region I refer to pretty fairly marked out. It includes Abyssinia, Somali-land, Mozam- bique, Natal, the Cape, Namaqua-land, and Angola. A suc- cession of great lakes and deserts is known to mark out part of the western margin of this region; to the west of it, or rather of the barrier so composed, we have what I may call an island sur- rounded by a nearly dry ditch, viz. the unknown region between Gaboon and the Congo on the south, a moat only partially sup- plied with water, the Albert-Nyanza line of lakes on the east, the Sahara on the north, and the Atlantic on the west. The countries of which this island is composed are, Senegal, Guinea, Old Calabar, and Gaboon. While it has a large share of the general African element of the eastern side of the continent, it has also specialities of its own, and superinduced upon it the very per- ceptible flavour of Brazil of which I have already spoken. In fixing the southern limits of this South-American element in West Africa, I have been guided partly by the descriptions 70 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF of the Coleoptera of Angola, published by Erichson in Wieg- mann’s ‘Archiv,’ in 1848, as corrected by Wollaston’s removal of the Cape-Verd species, which the death of the collector, Grossbentner, had occasioned to be confounded with the ge- neral Angolan collection, and partly by the collections made by Dr. Welwitsch. The Fellows of the Linnean Society know Dr. Welwitsch chiefly as a botanist, or, perhaps, through his and M. Morelet’s recent work ‘On the Mollusca of Angola,’ as a conchologist. But his entomological collections are not less admirable in every respect than his botanical; and through his liberality and kindness I have had the advantage of studying in them an amount of material greatly exceeding in extent the collection described by Erichson. From it I am enabled to say that the Brazilian element does not come south into Angola, The type of the Angolan Coleopterous fauna is Caffrarian beyond any question. The Brazilian stirps should alone now remain to be treated of ; but in speaking of other regions I have already said by anti- cipation every thing that I have to state regarding it. There is one point, however, on which I have a general remark to make, which is also specially applicable to it. In Columbia and some of the border-lands nearest the microtypal stream in the Andes we see many fine, rich, glowing metallic species, which, from their size and beauty, we are naturally led to refer to the Brazilian stirps, but which in reality belong to genera which, without doubt, are naturally microtypal, as Harpalus, Philonthus, Xantholinus, Staphylinus, &e. When we have to determine to which stirps such species belong, we must discriminate between the natural brilliant elements of the Brazilian fauna and the su- peradded brilliancy developed upon microtypal forms by the special conditions of the locality. Columbia and Ecuador abut on the eastern margin of the microtypal range in Equatorial America, and a modification of the stirps is to be expected there. Mexico, Central America, the West Indies, New Granada, Colum- bia and Ecuador, Cayenne, and in some cases even Venezuela are to be regarded as debatable lands, in which the true character of the stirps to which the species inhabiting them belong is to be determined, not by the place in which they are found, but by a sound consideration of their affinities and distribution elsewhere. THE CHIEF COLEOPTEROUS FAUNZ. 71 APPENDIX. Tasiz I.—Showing the actual present distribution of the following existing genera recorded to have been found in the Miocene beds of Oeningen, Radoboj, Rott, Aix in Provence, and Iceland. _ _Exranation.—A single asterisk means that the genus occurs in the countries indicated by it. A double asterisk that it occurs in force. Where numbers are given, they indicate approximately the number of species known to occur in the country in question (note 1). 1. Coleoptera. “| Jndo- Microtypal stirps. Devalable §| African (note 2) Sau ci geticpss £ [es Genera. g ig :|3 S| a] dd s FI es 84 atlatts, : o| 4 ai 2 le oy 2 : 3 54 slslslBal ole Slal [slale 3 41S Saale aie ols] (Hiss 2 she kel BIN EIS SR Sta] sales S| sl lle || Sal Shasta yis). BlaPrsleiste| S| 2] BISSICRE BIO m! 2) 4) aisle hale Cymnidis ....... supsoobe vee... (O0/36/34! 41313 ]...]... Py NOR ea aca igs 2 Brachinus ...............00e08- Sd cd baked cal eal Gel Gel Be £ |boooas * |*| 1x] x Anchomenus (note 3) ,...../42/35/61/13/12)3)3)4) 5 |...f.{Lal...1 3/3 Feronia (Argutor)............ weelexex] x [peepee] Ke PL. ee] P |e |e Hanpalug. (iii ss descncbsencar exlexfee] effete tet oe fox fo. x|?Pl ele] Bembidium .................. ed a Bee ee 2 ale Palwobiusyassascecsee nee * Agabus ...........00 G000s00000 Seda eae ee) eed 2 2 x Colymbetes ...............65. we xe feel LH [HL HLH] OH LeeLee deeedeee Dytiscus....... poeb0000000000000 Se ea ad el Soe a a Bee Bee ee a ee Oe Hydrophilus .................. cae a ea ee ee Carr Boao Mapaue x % Hydrous ..... gea6dq000000 coool 1%] He] He] KILL. E} looocedbooouc Pa pee ee Hydrobius..................05 ¥ |e] x] % * & |lbaoo0d|poooodl oqo oa9| bod locd lane Philhydrus ............. coef #] [HLH] HK]. §5) |codd0d bosood loodune| dod oe * Laccobius. ...............0e000- *|#] | x A08 ood Ochthebius ...... dosoacon0od [UT al! 2311 63 bac}osellooallood accocdl Loorsdl oonbod lone Loelloe Tachyporus (note 4)......... x%! x [xs]. ..] x Philonthus..................06. wxlexiex! ee [ele tae] % fo... * | % * Staphylinus .................. 24/15)8)5)}2?|/2)1)1) 8 | 1 |......158 1 Xantholinus ...............06- #11]... <) || beo|| peel boeaae * |* * Lithocharis ...............06 #1 21H 1...] # fowefeeelee. 2 | 6poboe basaee % Jefe. SUBMIS) Gondosooqcagoaso00000 veel [eee] M]eeelese[ee loool #1 SlOHUS Ne cscs nes odocaganaNda ee al ee ea Bel | {} |Ibeadoo| booed wa eee eee * Oxy POmitay ie semeceareece ences %1...| % Anthophagus.......c.sse.00 ee ea Omalium ..........cecesceeeee wee] 4 19% P| Bosoad boasad eecllona das 72 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF 1. Coleoptera (continued). | Indo- Microtypal stirps. Peobetable a! African j r| stirps. : (e3) Genera. SI hualince : SB) dh SAH ES ro) 18 Bla eSlsg e i | : 41S 4/0 q|* os d SISIE GIeolE sla g 2 O15) EES ols rs = Ni sia sie sia ela a0 ‘S) ay A le S) s F Ele| S| cal ie ce | Aly| Al 231159 [lao lboo baal locdllaod|eodbod|becooolloabocd fon * Oreina ........... 00 090000000 * ]*]...[eL i Gonioctend..........cccscereees % [%]*]..fee. Coccinella (note 27)......... x feel] He | WASIA ses sestsicoccvoce delessilecieca} ie [Ge Notes to 1. Coleoptera. Note 1.—This Table is made up to show the distribution of par- ticular forms. On this point, the relative number of species is a secondary consideration ; and even were it not so, we could not use it; for the statistics we have are, for any purpose requiring accu- racy, wholly unreliable, from insufficient exploration and imper- fect knowledge of the contents of different countries, from unna- tural dismemberment of forms into new genera, and from the inaccuracy or carelessness of describers, who, besides describing, as new, species which have been already described, have multi- plied or diminished the number of species, each according to his .own notions of what constitutes a species. To say nothing of the number, however, might sometimes lead to an undue idea of the strength of representation of the same genera in different countries; I have therefore occasionally added the approximate numbers of the species known in each district, when it could be done without much trouble and with some approach to accuracy. Where I have not: given the numbers I have marked the dis- THE CHIEF COLEOPLEROUS FAUNA. y 75 tricts where the genus is strong by a double asterisk, where weak by only one. My numbers, where given, have no pretence to more than a general approximation: for my purpose this is enough; and I believe they will be found sufficiently near to answer it. Note 2.—The geographical regions in this Table are :— 1. Europe, including the Mediterranean district, North Africa as far south as the southern margin of the Sahara, Syria, Asia Minor, and the shores of the Black Sea. 2. Asia, north of the Himalayan range, from the Ural moun- tains and Caucasus eastward to the Pacific. 3. North America east of the Rocky mountains. 4. North-west America from Behring’s Straits to Mexico. 5. The Chilian region, including the whole of South America to the west of the Andes, the part of Peru in the Andes, part of the South of Bolivia, and the western and southern part of Para- guay, also La Plata, Patagonia, and Terra del Fuego. 6. Australia, including Van Diemen’s Land. I should have wished to divide Australia into the northern and the southern halves ; but as yet our materials are insufficient to allow this to be done. More attention is now bestowed on localities; and I trust it will not be long before we can fairly allot the different portions of the Australian fauna. 7. New Zealand. 8. Polynesia. In this I include the islands between Australia and Chili, except New Zealand. 9. Debatable land between North and South America, in- cluding Florida, New Mexico, Mexico, Central America, and part of Columbia, New Granada, and Cayenne. I have added this and the two following columns of debatable land for the pur- pose of as much as possible keeping the elements of the faunas on either side of it free from what may have been a foreign in- fluence. The reader can carry it to the credit of either as seems to him agreeable, or omit it altogether. 10. Debatable land between the East Indies and Asia, in- cluding the Himalayas, Nepal, Silhet, the Burmese mountains, part of China, &c. : 11. Debatable land between North Africa and the northern frontier of Senegal. 12. Brazilian region, the parts of South America lying east of the Andes and not above disposed of. 76 MR. A. MURRAY ON THE GEOGRAPHIOAL RELATIONS OF 18. The Indian region, including the East Indies, the Ma- layan Peninsula, the Indian Archipelago, Siam, Cochin China, the south of China, the Philippine Islands, and New Guinea. In any list for more general purposes, New Guinea and the Philippine Islands should be kept separate; but, looking at the genera I have to deal with, this is not necessary here. 14, The West-African region, consisting of the country from Senegal to Gaboon inclusive, and eastward until it meets the East African region, wherever that may be (probably in the line of the lakes). 15. South Africa, containing Angola, Caffraria, Natal, Mo- zambique, and northward to Somali-land. The south of Arabia also, I believe, properly belongs to this province ; but it does not happen to come into question here. _.16. Madagascar. I do not include in it the neighbouring islands of Bourbon, Mauritius, Rodriguez, &c., although they do belong partly to it and partly to India; but the points in which they correspond with it are those in which Madagascar also coincides with Africa; and my object in keeping Madagascar separate is not to show its relation to Africa or India, but its more unexpected relations with South America. When I have to deal with species from the smaller islands (Bourbon, &c.), I carry them to the credit of India or Africa, according to which the affinities of the species indicate. Note 3.—Anchomenus. Some of these supposed Brazilian Anchomeni are, doubtless, as suggested by Lacordaire, Dyscoli ; and of others the habitat is possibly erroneous. Several of the citations are of old date, when every thing from South America was ticketed Brazil; still there are undoubted species from Brazil. Note 4.—Tachyporus. (Out of more than 250) only one occurs in Nepaul, doubtless a straggler, one in Bengal (which may have been derived from an Himalayan straggler), one in Java, and one in New Guinea. Note 5—Byrrhus. I include in this the allied genera, Curi- mus, Cytilus, Morychus, and Amphicyrtus. Note 6.—Onitis. Tolerably numerous in Europe and Asia, but only one in North America, one in India, and one in Au- stralia. . Note '7.—Aphodius. Only one recorded from Brazil, out of an THE OMIEF COLEOPTEROUS FAUND. =i wa immense number which, with that exception, are found in the microtypal and Indo-African regions. Note 8.—Anoplognathus. 1 have included under this head: Cotalpa (North American) and Brachysternus &c. (Chilian). Note 9.—Pentodon. Although not present as a genus in Ame- rica, it may be present under some one or other of the American’ types of the Pentodontide, as Podalgus, Heteronychus, Bothy- nus, &e. Note 10.—Ancylocheira. Inthis I elute Bulis and sthaeaias Note 11.—Ampedus. In this I include Elater and Grammo- phorus. Note 12.—Ischnodes. By Teehnoudes I suppose Heer to mean’ the second section of Candéze’s Anchastus. The first section consists of three Brazilian species, which I omit. The second is distributed as in the Table, and, besides, contains one St. Helena species, Anchastus atlanticus. Note 13.—Cardiophorus. Only one in Australia out of about 150 species. Note 14.—Telephorus. I include Podabrus under Telephorus, not only on account of the indistinctness of the fossil specimens, but from the closeness of their natural relationship. Note 15.—Clerus. I am not sure in what sense Heer intends Clerus to be used; and his figure scarcely helps us. I have ~ taken it in the narrower sense in which it is now used. _ Note 16.—Gonocephalum. In this I include Opatrum. Note 17.—Boletophagus, including Ulodes and Eledona. I can find only one species recorded from India, without special au- thority. If from the Himalayas, it doubtless is.a straggler from the north. Note 18.—Uloma. The common species .is introduced with cereals into all lands—which, as Lacordaire says, leaves its real native country in uncertainty; but, from its occurring in the Miocene fauna of Europe, it probably ought to be referred to Europe. Note 19.—Apion. The debatable-land species are chiefly from Columbia, which is half microtypal. Note 20.—Brachycerus. In this I include all the Brrciy conde. . It may be a question whether this form was originally African, and from Africa passed into South Europe previously to the Mio- cene times, or, being European, it subsequently found its way | into Africa, which is now its head quarters. 78 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF Note 21.—Hipporhinus. Similar remarks apply to this. Note 22.—Acalles. Largely represented both in itself and by allied genera in the Atlantic Islands. Note 23.—Cossonus. Ditto. Note 24.—Lamia. The genus Lamia is too large to know what type of it Heer referred his fossil to. Iam in doubt and have had to leave this blank. Note 25.—Dorcadion. I include the Mexican form Monilema as part of Dorcadion in this inquiry, not as otherwise a bad genus. Note 26.—Donacia. Lacordaire, in speaking of the Indian and African species of Donacia, cites, as an interesting fact in re- gard to their distribution, that they have more analogy with the species of North America than with those of Europe. I cannot, however, see it. Note 27.—Coccinella. The special genus or subgenus of a fossil Coccinella can scarcely be distinguished. I have therefore included several allied genera besides Coccinella, which makes it cosmopolitan. Strictly confined to the modern subgenus, it has been met with everywhere but in Polynesia and Madagascar. The vast numbers in which it has often been met with, and the appearance of flights of them migrating to other quarters, have probably something to do with its wide distribution. 2. Orthoptera. of|/_d|/ a] g ene Genera. Sd = 7 £ 2 x 8 |'2 s Bsolsa| #/ a| S| 2 lee E Aid| 4 io | 4 | 4 |= Phaneroptera ............ * * o00)) {| 06g x ee * Gryllacris .........+......6. dog |tsoca 300 ae * o0c * Cidipoda .............0.0. % * * x * Gomphocerus (note 1) ...| * * g06 x soc % IWIGRONHIS) Geacogboocesoncdocnac * * * * * # * IBY EN poconsceasosqop noDGCOdGe * * * * * x * Note 1.— Gomphocerus. This genus is divided into two sections, - of which the first is confined to Europe. THE OHIEF COLEOPTEROUS FAUNZE. 79 3. Neuroptera. : aloge 2.9 Sees i |e bie Genera. BS Ee] £ i 3 S SE i) o Rel@aia| se | 8| 3/6 Termes) (7201e")))yaseee nee oe * * * |x New Zeal.) x * x PAG ION Ee eect nate cise * * * |* Polynesia.) x * * WAGSCHNA fk asesscematnec se * x * |x Polynesia.) * * * Corduliay sds esas x * * |* New Zeal.| * |S.Afr.] x Tiibellula yeas senses snes * # * |» Polynesia. x * * Phryganea(note 2) ...... * ¥ 0060 500 30 || 600 ? BNGtACUSH MMe edean ee oles * x * * * * * Miyrrrieleor tease suascsae ss * * * * % # * Thewctran: Pac sesccseats * * Tetimuss eee ca. see 290) |b sooo Illy ous 260 * * Note 1.—Termes. Almost entirely tropical now; but two or three small species still survive in Europe (France, &c.). — Note 2.—Phryganea. Used in the sense of Phryganide; Phry- gene itself only occurs in the Europeo-Asiatic and North-Ame- rican districts. 4, IZymenoptera. a ; a a a. SS § a ; of s | Genera. S) 5 # on Ss 8 SE, Biel 0S | i) Gel as Els Bsital aio} £€] 216% Xylocopa ........s.seeeeee * | # * ® % * * Osmia aie see saan caine cies % 900 ate se ... |S.Afr. IBomlbusisss. secsese scenes * * ane * * sep Anthophora ............... % * * * x |S.Afr.] x LN OTM een Sdanndeeeasnecicene % 6 % * MESPAa) esictosscaotateanes raise * * Sap * see IHORMIGAeseneee eee * * x * % ¥ ? POnera ieee nieces * nee * sae * % ¥ Wily ABeaieEy || Goobenencapoedcdee * * x * * * * Pompilus ...............06 * * * 600 * 200 * Ichneumon ..........0.06. * * * * * * * Anomalon ...:........0.-00: * * * NG B50 * * OLA 7 DEG) cocodsnocnoo sesccndee * * * * * * * ACORNItES............ ccc eeees * * Hemiteles ...............00- ¥ % ¥ Tenthredo ............e.0008 * x ; * * * 80 MR. A. MURRAY ON THE GEOGRAPHIOAL RELATIONS OF 5. Lepidoptera. Oo 8 a q ae g| 4 ; |.Sa Genera. os 25 & pl S 4 & Ae@s|/ 4/6] 4] 3/6" Vanessa (note 1).........0+ % % * * % Psyche (note 2) ...-....+06- * * x x |S.Afr) x There are also four new fossil genera, which Heer calls Pie- rites, Bombycites, Noctuites, and Phalenites, which, he says, were respectively allied to Pieris, Bombyx, and the old genera Noctua and Phalena, which are generally distributed. Note 1.—Also found in the Navigator Islands. Note 2.—Psyche. Used in the sense of Psychidew. The recorded species of the restricted genus Psyche are confined to Europe and Ceylon. Be lgé| a 8 n "3 f Genera. Bx ¢ B £ iS S| § ag\|az& a| |e) ee a Neh} a ee Alel2/S] Jelelg Bam eoallees HIN |; : 2}/_/ S's |. ‘ i | 8 S/ ZEB BlSIEIE |S i312 AlO|Ala|o/4 [zim ja ls IP Lochionussacnccceeeeeeeee maceeeeeees * | Bieta niee ek oes eachia caremeeuamsiedeane lheae x | x | * | * x | x | * Chleenius ...... BRR Ae AAP HOGOGE M.|... fea | toate est ake *x | * Harpalusiccitesccs: costeseadcnsses Melee * | x | | * tale: AnchomenusS.........seecceseeeeeees WEA soo * | * | x Bembidium ....... aE aoa senat alse M.)... *x | x | x 506 |}-d00 || TF Colymbbetes|hi 2.2. -rsceste-s ete esieiee J oe ee ied | ea We EN Bot coal ly tt JMSEASYUS)rnc9001900800000050008000000000 M.]... * | ok | x 0 || c00 |}-0001] UF (ERATE cocandeaococsoondeoseocenGs0000N4) coc * | kK} |] x |... | # | | % Bolitocharay) -peccssseseeeeesteesseecees|| see * 84 MR. A. MURRAY ON TIE GEOGRAPHICAL RELATIONS OF Tae II. (continued). Microtypal stirps. Genera. ; & ||. g\4/8 s £\g|e a a\3|a/4 B Se eer ea les =| Bilo|s ): E ol1g|/2)] 46/4] 6 AIO IAIAICI14 Placusa ........ Hootionooaeds go06n00008000 ves * | Staphylinus ......... sdosenooococonodild | sos * | x | * | x Philonthus .....ccccseseeesceeseoes M,|.-- *% | ox | %* | x SIMMS socqdbonsedebdndasdasaabono5gs M)... * | x Zirophorus.............5.¢ Sgoo050808004| 959 36% || C Lispinus........... ShosgogoodoooaSscnncca}| 208 os Tsomalus ............ pogdoadsoosennaoee ses Chrysodema ............eececeeeeeeeee 206 |} 600 |) bod Hurythyrea .......... S0ecaadedooees MHI) one * JMTTPNTEY geseasqcndcosios onbaqoosodéec M. rs Photophorus (peculiar ‘to Poly- nesia, but allied to Pyrophorus, which is Brazilian) .........0...6. teed 900 || coc Dicrepidius (doubtful if really this genus, see Candéze) ............... . # |... Monocrepidius .. ...........eeeeeeee. . x | * | x Simodactylus, allied to Eudac- tylus ...... sedans Pu(eioislslsilate’sseieiislorsre ls|| slew earl Rtsoulltcereen [teete Hae Oophorus(the old genus of this name had no good characters; and as Candéze has broken it up and partitioned its contents, it is im- possible to say where Fairmaire’s insects should go; Candé:e him- self does not say, but suggests Heteroderes) ............ poossban. ond) 2 IGlellazaan Nee dea dara selena palege seperate eras eserauil rsa | ese pe sea | ae Cylidrus...............0+ soncodandoconsel] cas 965 ||‘c00 |] coo ll ose |) <4 Tarsostenus ..........s.ssseeeees s00gdl| 902 ooo || £2 |] 2 |\ coc Tillus (Clerus is the Miocene ge- nus, but it may be held to include TRUS) caonasensosctedda995900%000 M.. * | # | | x Corynetes ....... Sere eet orale ee cant ae ao ae x | x | * | Carpophilus ...... cooxssanodaconscqsceel) 006 alae |*], Nitidula ...........,. dcaQa0n0008 «x | x | * Hpurea ............. égpdacdedes .. pM x | x Stelidota ............00e- Keveaebe es aese|) Gone Anthrenus ...........46 HoaDD nC HOCeCLUGAA soe * | x % Wermestes)y mass escrectcsce reer ac M.... fa || ae |i se |p ee Platysoma........... peqcdeds920000006¢el| o00 * | x x I PEVROTNTD WS ood oneconndcadeanoaaascqaccobel| aoc oe || | New Zealand. Fos ee a | Brazilian region. | Brazilian stirps. * Indo-A frican stirps. : yt | India. 3% VHE CHIEF COLEOPLEROUS FAUNS. Tanne IL. (continued). Microtypal stirps. Genera. | Brazilian region. | Brazilian stirps. | Cosmopolitan. | New Zealand. | Doubtful. Tlydrobius .......- .....eeeeee ee M,)| ... Philhydrus ......-..seeeeeseee eens M, ... Cyclonotumr ............seeeeeeeee ee mck Aphodius ........-seeeeeeeee ec ees M. \ OXYOMUS ...ceesceeeseeeeeeene ree ens OVryctes .....cscscereseeeer eee seeeeeeees ie Figulus .........scscceesssereeeeceeeeees an Dorcus (Alcimus) ............00000200- Ns Passalluisitiscs uenedosen cones doumecenemans me) Opatrinus ? (real genus doubtful) MWerchemtmbpeneccesscscsectcceceeeeeerctct ae Boletophagus.... .....cseceeeeeeeeece es ve Diphyrynchus (probably allied to Uloma, fide Lacordaire) ......+.: Heterophaga (the species cited is cosmopolitan) .........s+.+eseeeee de Wikre, _{secosesasansuoosadseunba0000 M. x | New Holland. yes | North America. * ¥ ADO HR HHH | Europe and Asia. * * eo: + eH * * * * (CERONGIMNY, 5... cogcbasoacosds0db00000000 oll gue Amarygmus ........ OE yaar nEMBON aaaseo| Wie Olistlinemayswacsaesesccelesw ak osemel eee ee ANTUNES” cosdooocnoosnendooood oodgonooal| Gob Moordell ay eases ioasistccacewsljacw eles celal ste ZoNitid ......0s...e 00 acos06 asegeadoReoel Vinod IMOOREGE “sdonococcoqaco0b0cds8000000008] 9x0 ae 000 Selenopalpus.........c.cceeeeeseseneeee| one soa'|} coo |}'a90 | 000] ooo |} 12 IDTIVSLNEY. cso nnoonsceonadoGooandoddasacoonal| aon Rhinobrachys (supposed by Lacor- ‘| daire to be near Protedus, Pasc., a New-Guinea form) ............ Mropidenesimsveesske ster s meee ee (ees sailboat seal G2 al Gb lbooed fay ( The Otiorynchidee of Polynesia & the Indian archipelag. The rest of the fa- mily are Kuropxo- Asiatic & Brazilian. x *¥e* : wR DS * x XH 9 OK x ac) Spheerorhinus Trigonops ... Celeuthetes ... Psomeles...... Elytrurus INGLE) ooodrons docs baanebacnreousnbns tes 360 |] 200)|| 000 || oc {f-aoo:t} cool ode Cryptorhynchus ...............+6. M.)...)...] * | # |] # | # |---| x Tylodes (doubtful, fide Lacor-| ~ daire; probably should be ALGHNES)) secacoocsodcoxescexdcs0 bad M | Indo-African | India. Kok Ok * + | Africa. % # HS * * # * +++ tC —+ *« 85 stirps. * 86 MR. A. MURRAY ON TILE GEOGRAPITIOAL RELATIONS OF TasuxE IT. (continued). Pa El og Microtypal a 8 stirps. Ee % B. S| sa | 4 Genera. . 5 E d i; {4 ].2 anos Sele) Elle «|e S\3 elelels|lelsl4l ale B\ElEl= AlE|Ela|S 8 216 ° o|o] s a AJOIAl4|o/2\42/a}8\< ANGGINES pconooopseob0nbacdes gaosn6000 WY Paes |W ose x | t | t | oo Sitophilus ............ godendeeescnadenod|| 039 * rg Wallin rat ee ee Pelinades ono secelssnen| ees ong | E3 * il S Dryophthorus .......5.cscecceusreeees| v0 * | * ‘ COBBONUS ......ceceersceesecereecees M.|... # | x Phy ail ANNOY A NCLTALEY Qosagonopaobosdoonvabuce) 606 |) bon || Boe |] G05 |} Goo || o0el| bos 900 * TEIREEAUITMS, Goadonpecosn9q00n0050090090095)| 000 £3 |] doel| 600 * op Catolethrus ...........cccecscseceseres| cee | coe | cee | cee | cee | eeed one * Proéces ....+......- Doondncdesooa00NN8)| coc ae p90 =| Io AACCUDssoonn05. dansqooosonetiooaanDodl) 30 * | * | |.) ... x | x | * Psepholax (resemble Scolytidze). ANEW) Gooac090000000090990009000000000 % Platypus .....020....0ccsascececereree-| oe woe] #1] * | % * | x | * Temnorhopalon (position and afli- nities not known, supposed near Cerylon). Cicones .......... pansoons d0Nd00Nb000D000)] 000 % | % a DitOM A) (o.- eee ceececensserssceceessacisel cle: * | x ? © S oS CWenyloneeenetareceneteseteaeree meter liar * | * |... S Hmmagleus ...............c0ceecees see| a Silvanus....... Te a Sana ee ed * | * x Tt Rhizophagus ..........cecsceceseersceee| vee % | * PURO OSUD,” bcoosacccopadnnon.00K9Ge0d0009)| onc * | * |. * Laemophlous..............s0seeeee weealieee eo eA hetie lisol aes wR DendrophaguS .........0..s.seseeeceee| oe * | * *1 * | 3 Mallodon (belongs to the Rempha- AY NES) oogusconsocdodendosgonSAonooonasod|)/ab WP |lcoo | 3 oof |) 3 () OBIT Gongesdano95009000n05000000 ooadod|| StromatiumM .......sceecseeescneeecees * Steirastoma .........ccseeesescnsceseee| one 200 |] 000 |] ca] coc * TAR FEIOOES socodoooecoonoso0D0nan00009004)| 000 % Lagocheirus ............ Sqoavnnadesou04l| oo0 600'|| 86 |ligoal) coo * ONAN ocoosgabccsconnonsse0000s9008008 M)...]...] # | * | % | % * | * | x Hesperophanes ............ceeceeseeee-| oo we | ¥ 1% | % Oopsis. Saperda ....... Kagodsscansoceesn000008 M)... x | * * * | x Promecotheca ...........cseceeeeneee| one 506 |] 090 |} 290 || 000 | 2a9]| coc * Raphidopalpay suc-s.ssasccenscee secs] ce 580 t x | x Aulacophora ......... HERS HEGRR Aan RG ie oe ita t | #1 % Cryptocephalus...............002se000-] oo we low | |e]. | ok | | x Coccinella ............s..s0sceseeees M)... x |x [ox | kl | xe | | x TIE CHIEF COLEOPTEROUS FAUN®. 87 Taste IJI.—Showing relations of genera found in New Cale- donia. (From Montrouzier’s Essay in Ann. 8. Ent. 1860.) Microtypal. 8 : I -, | Uni- Genera. se eg Be 4 Brazil. versal, Eda] 2 | 3 Adz) 4a | 4 Cicindela ...............06. * * pe * * Distipsidera ............-+- aie * Cymindis ...............-6- * Tricothorax, n. gen. Scarites: qacigieosivocapeoace x * * Es * Chlaenius ........0.....000. * Eg ES * * Lissauchenius .....: Pesoeal bees * Oplonusy trecesscecenen cog|f £2 Rembus eoc6iibecacssccseenss 500 * Catascopus...............0+ 306 * Feromiay ee accesses * bax acca Gcase eee * Catadromus ..............- boo * Bembidium ............00- * Cybister ..........00 ooago0ee # * * * * Colymbetes ............05. * Copelatusiieeccss-nee ssc 0 * * Pachytes, n. gen. Dineutes.........0.cscceeeees %* * * * * Ochthebius.................- Stagnicola, n. g. Hydrobius .................- * Hydrophilus ............... * x * * ® Buprestis (Cyria?)......... * Abrobapta (Melobasis)... * Diphucrania ............ cod] oec % Agrypnus .........0.06 coed! 23 * * * * Athous? acosecsacc-ausccuees * Ludius ........... seseaaaee * later we ee soe * 57 Nycterilampus. Laius. Lomcechusa ? ......... APeos Wane Cylidrus.......... govendeen0s * * * * x @lerus iii vce esscssces BaceSces * * * * * TNI@OPALH) Gonodadosooscedconose Ace * Necrobia...........sccescsee: % x * ¥ IP GIM UBER eke eee pe 2k * * * x Tips sciico) Mitaeicesaeascectncs * Mycetophagus ............ Ok Dermestes ...............0+- ES * * * * Trinodes...........e0s00e008- * Macrosternus............++. Saprinus.........- Be aeucnoe * 88 MR. A. MURRAY ON THE GEOGRAPHICAL RELATIONS OF TaBLE IIT. (continued). Microtypal. : : Genera. se] a2) Fad Seer | ca i) aes BSE ! rt o E.S<| @ £ S EE AG Si) eI 2 Fy 5 Asa] A =) Onthobium (near Tessa- (REGLOD)) oceiaconescas00e09806 ae x Aphodius .............060. * * * * * Spheeridium ............... 500 000 % Rhizotrogus ............00: * * Ee aul alae * Cyclocephala....... pose6eas 066 og a0 * Hexodon .......... cocooseel} 900 * Scarabeeus ............ apoode % * * * * Ceratophyus .......... steal BE 1] ORVEIES. soacoonsoensa0ngc0000 900 Ne & * Megalemus, n. gen. Lucan .....000ceereeeeeee es * * * * # Ryssonotus............ coceaal| coc * TEMG) 5 .coconconacosabdase 300 * Passalus.......00..00++-+00 S06 Be * Opatrum .........c00ee ee, * Toxicum ............60605- Acanthosternus ............ 008 * Neomida....... Eonosagsoaccon * Diaperis............... poe * Leptomorpha...... seems eraes ae Pachycorus, n. gon. Tenebrio...... podpuoascbabo8 * * * * * Uloma ...... paodngGan000ee * * * * x Tribolium .................. * Po coplbinemetccaeteecen cee * * * * * Megapalpus, n. gen. Isopus, n. gen. TENNIS, cooooonqoanancooce cool} <3 FIGEIMWUN GoopaconocpeKoH0oe Bor * Ditylusieeacceelse cert so00e||, i Tagria............ godansoneccd ti) 3 * * * * Mordella ............-.508 * Telephorus............... | * Nacerdes............cessoeeee| * List of non-microtypal Species in the Coleoptera of Polynesia, and their sources (all the rest are microtypal). I. North America.—Clytus erythrocephalus and Ptychodes vit- tatus, Longicorns (timber-borers). II. Brazil.—Lagocheirus araneiformis and Steirastoma stellio, Lon- gicorns (timber-borers); Brenthus bidentatus, a Brenthid (tim- ber-species). THE CHIEF COLEOPTEROUS FAUNA. 89 II. Hast Indies and Philippine Islands.—Chlenius guttatus, a Carabid (carnivorous hunting insect) ; Hesperophanes luzo- nicus, Longicorn (timber-borer); Figulus fissicollis (Lucanid), Apate religiosa (Xylophage), and Eurythyrea scutellaris (Bu- prestid)—timber-insects. IV. New Holland.—Oopsis nutator, Longicorn (timber-borer) ; Amarygmus hydrophiloides, Heteromere, and Apate pusilla, Xy- lophage (timber-insect) ; Nacerdes bivittata, Heteromere, and Staphylinus erythrocephalus, Staphylinid (microtypal). V. New Zealand.—Dendrophagus suturalis, Cucujid (bark-in- sect); Staphylinus oculatus, Staphylinid (microtypal). VI. Cosmopolitan, origin doubtful.— Plochionus bonfilsit, Ca- rabid. ° List of Genera and Species found in the Europzo-Asiatic regions, and also in North-west America, but not in the eastern side of North America. I. Genera.—Callisthenes, Miscodera, Leistus, Trachypachys, Pe- lophila, Anillus, Necrophilus, Pteroloma, Lyrosoma, Spherites, Bolitochara, Syntomium, Phlaoneus, Arpedium, Deliphrum, Malachius, Calcar, Rosalia, Ergates, Mesosa, Timarcha. II. Species belonging to other genera than the above.—Platynus Bogemanni, Carabus Vietinghovii, Colymbetes dolabratus, Necro- phorus mortuorum, Olistherus megacephalus, Klater nigrinus, Corymbites confluens, Helodes variabilis, Dinoderus substriatus, Serropalpus striatus, Ohrysomela lapponica, C. viminalis. List of Genera of Coleoptera of Old Calabar cither Brazilian or with Brazilian affinities, taken from my “Coleoptera of Old Calabar ” (so far as published) in the ‘ Annals and Mag. of Nat. IList.’ Galerita, Lia, Goniotropis, Hypolithus, Celina, Contipus, Axyra, Taracta, Platychora, Melittoma, Ptilodactyla, Dilobotarsus, Belionota, Parandra, Dorycera, Callichroma. Brazilian section.— ime, @idenoderus, Lrachelophanes, Distenia, Smodicum, Stenochia. Note.—The other Tables referred to in the body of this paper, and which accompanied it, relate to matters which are more ge- nerally admitted ; and it has therefore not been thought necessary to print them. LINN. JOURN.—ZooLoGy, VOU. XI. 7 90 DR. 0. COLLINGWOOD ON A NEW FORM On a new Form of Cephalopodous Ova. By Curunrrr Continewoop, M.A., F.LS., &e. (Puate I.) [Read February 3, 1870.]} Tur large grape-like masses which constitute the ova of the common Cuttlefish (Sepia), are of so remarkable a form, and so commonly met with, that they attracted attention very long since. Aristotle, whose acquantance with the reproductive bodies of the Tetrabranchiates was not far behind that of the present day, was no stranger to these large and singularly formed bodies; and they are commonly taken as the type of the spawn of Cepha- lopods. But the ova of this group differ considerably in size and appearance, as well as in the numbers produced by a single individual. In the case of Sepia, nature seems to have taken special care to preserve these important bodies, having encased them in a flexible horny covering, prolonged at one extremity into a kind of tendril or filament, which entwines round some fixed object which serves an anchorage. In the Poulp (Octopus), Aristotle informs us that a shell, or some such convenient nidus, receives the eggs, which adhere to it and are thus in some de- gree, at least, protected from injury. In Loligo, &c., great num- bers of ova are produced: cylindrical sheaths of a gelatinous con- sistence are formed, each about 4inches long and about 7 inch in diameter, and tapering at the free ends, the opposite ends being all attached to some foreign body by filamentary processes from 3 an inch to an inch in length. In each of these radiating bodies there may be 200 ¢apsules, each of which contaims from 30 to 40 minute spherical ova. In Sepioteuthis there appears to exist an intermediate form of ova, which connects the ra- diating sheaths of Zoligo with the large capsular ova of Sepia. The ova are ‘(as in Loligo) spherical, and enveloped in sheaths ; but, as in Sepia, these are fewer and longer; while in the Di- branchiates the ova occupy a considerable space at the bottom of the shell, as, for example, in Argonauta. In none of these, however, which represent the characters of of the Cephalopodous ova, as far as known, is there any approach to the characters of a remarkable body which I recently dis-. covered in the Atlantic Ocean, the nature, however, of which was incontestable. We were becalmed in lat. 37° N. and OF CEPHALOPODOUUS OVA. 9) long. 28° W., and the sea was swarming with beautiful objects, which I was watching from the chains, and making attempts to capture therefrom, when I saw an object which at once withdrew my attention from all the rest (the most familiar illustration I can give of its appearance would be to liken it to one of those cylindrical knitted comforters worn by ladies)—about 2 fect long and about 4 or 5 inches in diameter, closed at both ends, and floating expanded upon the surface of the water. The folds of the web were of a dark colour; and the web itself looked ex- tremely delicate, so much so that, except at these apparent folds, it could scarcely be distinguished at all. At the same time I thought I should be able to hook it up entire, and fetched a grapnel for that purpose. I should have ill succeeded in the attempt, however, as it turned out; but the ship being at this juncture getting up steam and nearly ready to move, the first Lieutenant, Mr. Stewart, kindly lowered a boat for me as it drifted past. From the boat it could scarcely be perceived; and when, by directions from the deck, it was ultimately found, some difficulty was experienced in getting it into the boat; for, although it had appeared very solid and distinct in the water, it proved impossible to drag it up, and at the first attempt the mass slid away from the bucket placed under it, and, being so evenly balanced, disappeared, and could not be found again immediately. I was sadly afraid it would be lost; for our vessel was only waiting for the boat’s return to steam away. Presently, however, it was found again; but in attempting to push it into the bucket, it broke in halves. Ultimately one of the halves was secured ; and this was quite sufficient for the purpose. On close exami- nation I was suprised to find that it consisted of a large mass of semisolid, perfectly transparent jelly ; and what appeared to be the dark folds of the web were rows or clusters of round black spots, each of the size of a large pin’s head, arranged in single rows along the outer part of the cylindrical mass of jelly, the rows not being regularly distributed, but running partially round its circumference, some for a longer distance than others. In some cases, two or three rows were placed close together side by side, and were separated by an interval from the next series, which might contain two or three rows or only one row of spots. The spots appeared to be most thickly clustered about the edges of the body as it lay in the bucket, and least numerous Vhs 92 DR. C. COLLINGWOOD ON A NEW FORM upon the central parts—an effect more apparent than real, and depending upon the cylindrical form of the mass. Although only half of the original mass was secured, the soft jelly having divided in getting it into the bucket, it had the appearance of an entire body, the fracture of the gelatinous mass not interfering with its symmetry. Turning my attention now to the black spots, I at once saw, even without the aid of a lens, that they were egg-sacs containing young Cephalopods. These were extremely active, moving freely in the sacs and contracting their bell-shaped bodies as they leaped about in their narrow chambers. Each egg-sac was per- fectly spherical and transparent, the circumference alone being visible, and was imbedded in the soft gelatinous transparent mass just as is the case with the spawn of the frog. The dark- coloured spots were entirely due to the coloured bodies of the embryo animals, which, in most cases, appeared to be just ready to be extruded. On placing the embryos under the microscope, I found that some of them were almost transparent, and exhibited their internal organization. The external surface of the bell was covered with epithelium of columnar form; and the same struc- ture also extended over the arms, The bell was covered with dark-coloured spots—which in the most immature specimens were mere minute round specks, becoming in a further advanced condition irregular and angular as well as of a larger size. The eyes were large and prominent, and seated upon short and thick footstalks, and their dark pigmentary substance was distinctly visible through the transparent bell of the younger individuals. The arms were short, covered with epithelium upon the convex side, and having a few rudimentary acetabula upon their con- cave surfaces. Upon the upper part of the bell, on either side, was a small fin-like projection, visible even in the least-mature specimens. Having secured this curious body, and examined its general form and appearance, and placed some of the embryos under the microscope, I was under the necessity of leaving it in a bucket of sea-water for two or three hours. When I returned to it at the expiration of that time, it appeared to have vanished. In some astonishment, I put my hand into the water, and found therein a large mass of soft transparent jelly, entirely invisible in the water. On closer inspection, I discovered that every one of the young embryos had been discharged from its sac, and that they OF CEPHALOPODOUS OVA. 93 were lying in little heaps at the bottom of the bueket, either dead or dying. They had entirely lost the active movements which had at first distinguished them ; and an occasional contraction of the bell was the only sign of life which any of them exhibited. Those which I had myself separated from the mass, and previously placed in a tumbler of water, were by far the most lively; and from these the accompanying figures were made. On no other occasion did I meet with a body of this nature ; and the only thing I ever saw approaching to it in form was in the Indian Ocean, north of the Equator, when I one day ob- served something of the kind pass by, which had been a puzzle to me ever since ; for the rate at which we were steaming (ten knots) rendered it impossible to take any accurate note of it. Nor should I have been able to guess the character of the body I have here described, had I not been so fortunate as to secure it for closer examination. The very great contrast which this body offers to the known forms of the spawn of Cephalopods in general is very remark- able; and its singular resemblance to the spawn of the Am- phibia is no less worthy of attention. What this may signify is a matter of interesting consideration. The embryo stages of this animal (of which I have preserved a few) will, of course, offer some, though a very imperfect, clue to its adult form, and to the determination of its genus. The presence of fin-like projections upon the upper portion of the bell seems to point out its separa- tion from the genera Eledone, Octopus, Tremoctopus, and Ar- gonauta, though to which of the pinnated genera (Histioteuthis, Sepiola, Rossia, Sepia, Sepioteuthis, Verania, Onychoteuthis, Enoplo- teuthis, Loligo, and Loligopsis) it may belong, or whether to some new genus, cannot now be determined. The body was evidently perfect in itself, and perfectly symmetrical ; and itis curious to ob- serve so large a mass, and such a vast quantity of animals as the product of a single individual. Probably in it, as in the Frog during the breeding-season, the ovaries occupy the greater part of the body; and probably, also, as is the case with the Frog, when the ova are deposited in the water, the jelly-lke sub- stance in which they are enveloped absorbs a large quantity of the fluid, so that the whole mass rapidly increases in volume until it becomes many times as large as the animal from which it was expelled. 94, DR. W. BAIRD ON NEW ANNELIDA AND GEPHYREANS. These interesting questions may, it is hoped, yet be eluci- dated, and the affinities of the animal determined. In the mean time I have thought it best to bring forward the fact for the information of zoologists and physiologists. DESCRIPTION OF THE PLATE. A, natural size of ova. B, C, young cuttles under a 2-in. object-glass (24 diam.). D, an embryo, as seen under a 1-in. glass. 4H, F, arms ( in.), showing the rudimentary acetabula. Description of some new Species of Annelida and Gephyrea in the Collection of the British Museum. By W. Bainp, M.D., E.BS., &e. ; [Read April 7, 1870.] ANNELIDA. 1. Nevutiys Maoanprewi, Baird. Body elongate, tapering towards the inferior extremity, which terminates in one rather long seta. The sides containing the dorsal feet strongly ridged across. Proboscis rather short and rounded. Setx of upper lobe of feet few in number and serrated near the tip ; rather shorter and broader than those of ventral lobe, which are numerous and not serrated on the edges. Colour of the dorsal region, in the centre, of a pinkish hue. Length about 6 inches. Hab. Coruna, &. M‘ Andrew and H. Woodward, Esqs. 2. Nerurnys mmpressa, Baird. Body of a yellowish colour. Dorsal and ventral regions smooth, of a pearly, somewhat iridescent hue. Ventral surface marked with a bluish impressed line in the centre. Ilead small; antenne indistinct. Proboscis rather long, cylindrical. Papille on the summit of it, round the mouth, rather large and fleshy, disposed in a series of 12 on each side. Feet on upper part of body small and close-set, becoming larger and more separate as they de- scend. Lamellx ovate. Setigerous lobe rather large. Superior branchial process involute, large, twisted once and a half round. Setw of setigerous lobe of three kinds :—one, short, curved at the tip and beautifully and minutely jointed; a second, simple, long, and slightly serrated on the outer edge; and the third, long, compound, the edges of the appendage minutely toothed on the edge, as is also the top of the shaft. This specics resembles very much the Mephthys longisetosa, DR. W. BAIRD ON NEW ANNELIDA AND GEPHYREANS. 95 the chief differences being in the ventral cirrus or branchial process, the more decidedly serrated sete, and the habitat. Length about 4 inches. Hab. Loto, coast of Patagonia, Dr. Cunningham. 3. Neruritys LuTREA, Baird. This species is considerably smaller than the preceding, but resembles it in most respects. The sete of the feet are long; but instead of being serrated on one edge, they are divided across in numerous small joints or articulations. In length it is only 2 inches. Hab. Otter islands, coast of Patagonia, Dr. Cunningham. 4, CLYMENE GRossa, Baird. Body of a straw-yellow colour, much wrinkled on the surface, and thick. Head-lobe of considerable size and much wrinkled. Cephalic plate large and crenate on the upper edge; crenations about 12 in number, each crenation again having two slight cre- nations on the summit. First segment of body without seti- gerous feet. Three following segments with a fascicle of setw only. The middle ones with a fascicle of sete, and a lobe possessing nu- merous very short set on it. Unfortunately the two Museum specimens are imperfect at the inferior portion. Hab, Straits of Magellan, Dr. Cunningham. 5. CuyMENE INSIGNIS, Baird. Body elongate ; thickest in the middle, which exhibits a sort of sheath or tube in which the worm lives. Cephalic lamina very small, entire. Posterior extremity obliquely truncate, with no infundibuliform appendage. Segments of body very indistinct ; one or two, of the anterior portion, without set; the other segments possess two rami. The anterior half of the body, exhibiting these sete, is very large, and the sete are very long and filiform. Posterior portion of body has the fasciculi of sete small. Hab. G. SrIPHONOSTOMA ANTARCTICUM, Baird. Sete: surrounding the head numerous, very short and fine. Branchio short, numerous. Ifcad withdrawn. Body covered with an enveloping substance like that of most of the known species. Seta of the inferior ramus of fect single, crooked or ? Taken during the Congo Expedition. 96 DR. W. BAIRD ON NEW ANNELIDA AND GEPHYREANS, hooked at the point, and of nearly a black colour. Colour of body varying from a very dark to a light brown, and of a trans- parent look. Length of body in longest specimen nearly 3 inches. Hab. New Zealand, Dr. A. Sinclair. 7. Magascotex (PeRicu#Ta) ANTARCTICA, Baird. Body consisting of about 180 rings. Setw, surrounding the body, short, black, rather distant. Rings not keeled; larger and more distinct at the anterior extremity, closer at the poste- rior end, and all smooth. Length 7 inches. Hab. New Zealand. 8. Mr@asconex (Pertonmta) Sanctm-Hetenm, Baird. Body consisting of about 86 rings, which are more distinct at the two extremities than in the centre. The 11 or 12 rings at each end, have an acute ridge or keel in the centre; those of the middle portion of the body have the keel flattened. The body of the rings is finely striated. Sete short, of a dark colour at the posterior extremity, rather distant from each other. In the centre of the body and at the anterior extremity they appear (in the specimen from which this description was drawn up) retracted, leaving only a mark where they are situated. The first 7 or 8 rings, at the anterior extremity, are strongly rugose or wrinkled. ) Length from 1 inch and 9 lines to 3 inches. Hab. High ground at St. Helena, J. C. Melliss, Esq. 9. LuMBRIoUS JULIFoRMIS, Baird. Body of about 120 rings. Of a nearly black colour with me- tallic reflections. Rings smooth, narrow, close-set, slightly keeled in the centre. Setz in four double rows, two ventral and two dorsal. Body of about equal size at each extremity. Lower extremity conical, pointed. The 10 or 11 anterior rings are the largest. Altogether this worm resembles very much in appearance a species of Julus. Length of medium-sized specimen 23 inches. Hab. ? Collected during the Antarctic Expedition. 10. Lumsricus Guiipinet, Baird. Body consisting of about 160 rings, narrow and close set DR. W. BAIRD ON NEW ANNELIDA AND GEPHYREANS. 97 together. Sets in four double rows on the back, each row very much approximated. No set# on ventral surface. Colour of a pale straw hue. Rings have the surface corrugated ; and the anterior ones are each slightly keeled in the centre. Length 2 inches and 3 lines. Hab. Island of St. Vincent, West Indies, Rev. Lansdown Guilding’s Collection. 11. LumMbBriovus rusro-Fasciatus, Baird. Body of a dirty yellow colour, banded across the back with a broad fascia of a red hue. The ventral surface is yellow. The red band extends across the centre of the segments. Anterior and posterior extremities both obtuse. Length between 2 and 3 inches. Hab. St. Helena, J. C. Melliss, Hsq. GEPHYREA. 1. AsPprposiPpHon JuKEstI, Baird. Body nearly smooth, of a light straw-colour. Anterior shield dark, slightly granular, more slender than the posterior, which is of a lighter hue, and radiately granular. Granules very small. Length about 3 an inch. Circumference about 8 lines. Hab. feibeudeds in a piece of coral from Lee Sandbanks, dredged in 14 fathoms, J. B. Jukes, Esq. 2. Houiunus FAROIMEN, Baird. This is a very large species, the middle-sized ones resembling in general appearance a large sausage. The two spines on the anterior portion are large and well developed. The hinder portion exhibits only one row of spines, instead of two as in most of the known species. The skin is leathery and smooth; the two ex- tremities are bluntly pointed. The longest specimen we possess is about 16 inches long, the shortest fully 7 inches in circum- ference. Hab. We possess five specimens of this species, all from Punta _Arenas, on the coast of Patagonia. Collected by Dr. Cunning- ham, of the Surveying Expedition to the Straits of Magellan, to whom we are indebted for several species of Annelides above _ described. 98) > MR. R. M‘LACHLAN ON NEW FORMS, ETC., On new Forms, &c., of extra-European Trichopterous Insects. By Rosrert M‘Lacutray, F.L.S. (Puarss II., IIT. & IV.) [Read June 2, 1870.] Tue present paper may be regarded as a continuation of se- veral memoirs by me on exotie Zrichoptera, published in the ‘Transactions of the Entomological Society of London’ (Trans. Ent. Soc. ser. 8, vol. i. pp. 801-812, 492-496, vol. v. pp. 247- 278). Many of the insects here noticed I owe to the liberality of my valued correspondent Mr. Henry Edwards, of San Fran- cisco, from whom I had already, during his residence in New Zealand, received such substantial evidence of his desire to assist me by collecting these neglected insects, and who, since he has made Western America his home, has continued to help me. I have not, however, confined myself here solely to Californian species, but have added several remarkable forms from other parts of America, and also from the Old World. No doubt it is always advisable to restrict general papers of this nature within geogra- phical limits; but this applies most forcibly to families which have already been made the subjects of general study. To fol- low this plan in exotic Zrichoptera would be almost impossible, inasmuch as, though occasionally a considerable number of spe- cies may be collected in one locality by an entomologist who attends to other insects besides the almost hackneyed Butterflies and Beetles, many interesting forms must remain unnoticed in collections for years, because they are the results of only desultory observation on the part of collectors. This, there- fore, must be my excuse for the scattered nature of the materials in this paper. When the day shall arrive when Neuropterists may be as plentiful as Lepidopterists, Coleopterists, and even Hymenopterists now are, it will then be absolutely necessary that workers should confine themselves, in each paper, within limits, either of locality, or family, or genus; to do that now would put a stop to all work, because, by the omission of any notice, collectors would fail to bestow any attention whatever on these insects, and the evil would be increased rather than mitigated. As in previous papers, I have endeavoured to illustrate by means - of outline figures those intricate points of neuration and se- condary sexual characters which form so essential a part in the OF EXTRA-EUROPEAN TRICHOPTERA. 99 study of Zrichoptera, and which can often be explained intel-— ligibly by a few strokes of the pencil, however inartistic these may be, when words fail to illustrate the meaning. It may not be out of place here to say a few words on the systematic position of the Zrichoptera. The remarks that fol- low have, to a certain extent, been excited by a recently published American work, by Dr. J. S. Packard, jun., entitled a ‘ Guide to the Study of Insects,’ a work strikingly original in its concep- tion, and oue which will doubtless do much towards furthering the already rapidly increasing taste for entomological studies in the United States. But it is necessary, first of all, just to glance at the position generally accorded to the Neuroptera. It has long been seen that the order, as defined by Linné, is composed of most incongruous materials; and Erichson attempted an ame- lioration of this condition by grafting all those families with in- complete metamorphosis upon the Orthoptera, still maintaining the two orders in juxtaposition. Since his time various authors have made this division, termed pseudo-Neuroptera, a veritable re- fuge for the destitute. To it have been added, from time to time, Mallophaga, Thysanura, Thysanoptera, and even the Strep- siptera, for no other reason, so far as I can see, than that they would not fit in satisfactorily elsewhere; and the characters of the order being so elastic, it was easy to find some peculiarities which gave these outlying families admission therein. That the Linnean families grouped now with Orthoptera have more affinity thereto than to the Neuroptera as usually constituted, is evident ; yet I see no reason whatever why the Odonata should not form an order apart, possessing, as they do, characters absolutely sw generis. The admission of them into Orthoptera renders an already heterogeneous order an absolute chaos. For my part, I have been content to consider the Newroptera as an order, in the Lin- nean sense, divisible into three great divisions, pseudo-Neuroptera, Planipennia, and Trichoptera,—but this only as matter of con- -venience; for I am convinced that contained therein are consti- tuents of several orders, each of equal value with such as Lepz- doptera and Coleoptera, and that the day will arrive when, from an increase of knowledge in embryology and anatomy, the order Neuroptera, as constituted by Linné, will be scattered widely—a dismemberment that would have occurred long since, only that there still exists a lingering disinclination to thoroughly upset the Linnean system. ~ 100 MR. R. M‘LAOHLAN ON NEW FORMS, ETC., Dr. Packard’s arrangement is founded on the idea that in insects, as in all other divisions of the animal kingdom, there are certain groups more elevated, others more “ degraded,” than the rest. Acting upon this, he places the Hymenoptera as struc- turally and psychically, if I may use the term, superior to all other insects. Then follow Lepidoptera, Diptera, Coleoptera, Hemiptera, Orthoptera, and, last of all, the Newroptera, in the Linnean sense (but including Thysanura), an order which, ac- cording to him, “mimics every suborder of insects,’ being “comprehensive or synthetic types, combining the structure of all the other suborders.’’ I would here particularly call atten- tion to the relative positions occupied by Lepidoptera and Tri- choptera, the latter forming nearly the last division of Wew- roptera. I emphatically enter my protest against such a wide separation of the two groups, considering, as I do, that, whatever may be the condition of the Trichoptera with regard to others of the Linnean groups of Neuroptera, their relationship to the Le- pidoptera is close, and that an attempt to thus widely separate them is an outrage on both. In metamorphosis the resemblance is nearly complete, the fact of the pupal limbs not being en- closed within a common integument not availing much when their condition in certain micro-Lepidoptera is taken into con- sideration: the possession of mandibles by the Trichopterous nymph is not of much importance, inasmuch as these organs bear no relationship to the aborted mandibles of the imago; they simply replace the acid or mechanical means by which a Lepi- dopterous imago frees itself from its cocoon. The imago in Lepidoptera is almost constantly furnished with scales on the wings and body, scales of a peculiar nature, the analogues of which are seen only in Lepisma; but many Trichopterous insects have, in the male, a modification of these scales in the form of short inflated hairs, generally intermingled with ordinary hairs ; and in some genera this tendency towards a scaly clothing is as marked as is its absence in some Lepidoptera. The neural arrange- ment is not at all incompatible with a close relationship ; nor are the parts of the mouth, excepting the absence of a developed haustellum ; yet many of the larger Zrichoptera frequent flowers for the purpose of extracting the nectar; and though I am un- able to say by what means this is effected, it seems probable that it is done by prolongation, at will, of the upper portion of the cesophagus into a sort of false haustellum. Perhaps the OF EXTRA-EUROPEAN TRICHOPTERA. . 101 strongest mark of demarcation is the presence, in most Lepi- dopterous imagos, of a spine-like process near the base of the costa of the hind wings, wanting in all Trichoptera. That this process is a modification of a vein is almost certain; and I ap- prehend that, when the homologies of neuration are better under- stood, this negative character in Zrichoptera will not be found of much importance. My own inclination tends strongly towards maintaining Trichoptera as a separate order in juxtaposition with Lepidoptera; and I am thus content to share the pity bestowed by the reviewer of Huxley’s ‘Introduction to the Classification of Animals,’ in the ‘American Naturalist’ (a journal receiving Dr. Packard’s inspiration) for November 1869, by whom we are told that (p. 545), “ the strangest, and, humanely speaking, sad- dest feature of this classification, is recognizing the Neuropterous family Phryganeide as a distinct order (Zrichoptera).” In a divi- sion of insects such as the Linnean Newroptera, which is so tho- roughly heterogeneous, much allowance should be made for dif- ferences of opinion, and it is scarcely fair to bestow such dog- matic censure upon any system, however opposed it may be to individual convictions. Family PHRYGANEIDZ. The following is an attempt at a systematic and synonymic catalogue of all the described species of this family, taken in its limited sense. The genera are not well-defined, notwithstanding the size of the insects, the neural characters not being suffi- ciently stable, or rather, perhaps, the materials at present in hand being too meagre, to enable me to draw lines of demarca- cation absolutely satisfactory. A few notes on the general characters are here given. Colpomera, M‘Lachlan, which I was inclined to place as a sec- tion of Phryganea in its limited sense, on account of the strong facial resemblance of the type to P. japonica; is evidently a good genus. The general characters are as in Phryganea; but the anterior wings are narrower, the apex being falcate, the apical. margin strongly excised. The apex of the abdomen of the female (which sex I have only recently seen) is produced into a telescopic tube, indicating some peculiar mode of life, and quite different from the blunt apex of Phryganea. The neuration differs in the sexes, as in P. grandis and allies. 102 _ MR. 8. M'LACHLAN ON NEW FORMS, ETC., Phryganea, Linné (as restricted), has moderately narrow ante- rior wings, the apex of which is rounded, oblique, or slightly sinuate. In the typical species there is an additional apical cel- lule in the 9 in all the wings; but in a section of the genus the neuration is similar in both sexes, or as in the ¢ of the typical species (Zrichostegia, Hag., Brauer) ; and in another section the anterior wings have the like neuration in both sexes, but the posterior wings possess an additional fork. The discoidal cell of the anterior wings is elongate in all. Holostomis, Mannuh., differs from Phryganea in its very broad anterior wings. In the typical species, the neurativn of the an- terior wings is alike in both sexes (similar to the typical forms of Phryganea 8), but the posterior wings of the 9 have an addi- tional fork; H. Maclachlani, White, has the additional fork in all the wings of the 9; and on this account I transferred it to Phryganea; but, in its form, it is evidently better placed here. Neuronia, Leach, is scarcely to be separated from Lolostomis : the species are, as a rule, smaller, with the discoidal cell shorter ; but possibly the two genera should be united under Neuronia, which is the older name. The neuration of the anterior wings is alike in both sexes; but the posterior wings of the 2 have an additional fork. Agrypnia, Curtis, is distinguished by the narrow, Limnophili- form anterior wings, the neuration alike in both sexes, the spines of the tibie and tarsi few in number,—at present one of the best- defined genera. Corromera, A‘ Lachlan. 1. C. stnrensis, M‘Lach. Trans. Ent. Soc. Lond. ser. 3, vol. i. p. 302. Hab. North China. Purya@anea, Linné (restricted). = Trichostegia, Kolenati. A. Ale antice et postice faminis furca apicali addita instructe (=Phryganea, Hag., Brauer). 2, P. saponica, M‘Lach. Trans. Ent. Soc. Lond. ser. 3, vol. v. p- 248. Hab. Japan. 3. P. cranpis, Linn. F.S. 379; Hag. Linn. Ent. vol. v. p. 363.— OF EXTRA-EUROPEAN TRICHOPTERA. 103 Trichostegia grandis, Kol. Gen. et Sp. Trichop. pt. 1, p. 84.—P. atomaria, Steph. Ii. vol. vi. p. 206. Hab. Europe. 4. P. srriata, Linn. F. S. 378; Hag. Linn. Ent. vol. v. p. 363.—P. Beckwithi, Steph. Iii. vol. vi. p. 206.—P. fulvipes, Burm. Handb. vol. ii. p. 934. Hab. Europe. 5. P. cinEREA, Walk. Cat. Brit. Mus. Neurop. pt. 1, p.4; Hag. Neurop. N. Amer. p. 252.—P. divulsa, Walk. Trans. Ent. Soc. Lond. ser. 2, vol. v. p. 176 (locality erroneous). Hab. North America. 6. P. varia, Fab. Ent. Syst. p.77; Pict. Recherch. p. 160, pl. xi. f. 1.—Trichostegia varia, Kol. Gen. et Sp. Trichop. pt. 1, p. 86.—P. annularis, Oliv. Encye. vol. vi. p. 558.—P. variegata, Humm. Ess. li. p. 23. Hab. Europe. 7. P. sorpipa, M‘Lach. post. p. 106. Hab. Japan. 8. P. oBSOLETA (Hagen), M‘Bach. Trans. Ent. Soc. Lond. ser. 3, vol. v. p. 16. Hab. North and Central Europe. B. Ale postice feminis furca apicali addita instructe. 9. P. vestira, Walk.—Neuronia vestita, Walk. Cat. Brit. Mus. Neurop. pt. 1, p. 10, 9.—P. vestita, Hag. Neurop. N. Amer. p.253. —N. commixta, Walk. 1. c. 3 .—P. commixta, Hag. I. c. Hab. United States. C. Vene alarum ant. et post. in utroque sexu ut in mare divisionis A. (=Trichostegia, Hag., Brauer). 10. P. minor, Curt. Phil. Mag. 1834, p. 125; B. EL. pl. exii.—Tri- chostegia minor, Kol. Gen. et Sp. Trichop. pt. 1, p. 87.—P. mixta, Burm. Handb. vol. i. p. 934.—P. tortriceana, Ramb. Neurop. p. 471. Hab. North and Central Europe. Hotostromis, Mannerheim. A. Vene alarum ant. et post. ut in divisione A. Phryganee. 11. H. Mactacuuant, White, Trans. Ent. Soc. Lond. ser. 3, vol. i. Proc. p. 26.—P. Maclachlani, M‘Lach. Trans. Ent. Soc. Lond. ser. 3, vol. v. p. 249, pl. xvii. fig. 1. Hab. North India. Var. REGINA, mihi (an sp. distincta ?). Hab. Japan. 104 MR. R. M‘LACHLAN ON NEW FORMS, ETC., B. Ale postice feminis furca apicali addita instructe; vene alar. antic. in utroque sexu ut in mare divisionis A. Phryganee. 12. H. pHauamnorpes, Linn.—P. phalenoides, Linn. F. S. p. 378.— H. phalenoides, Kol. Gen. et Sp. Trichop. pt. 1, p. 82.—P. speciosa, Lat. H. N. vol. xiii. p. 86.—P. daurica, Fisch. Ent. Russ. p. 52, pl. (Neurop.) ii. f. 1. Hab. North Europe. 13. H. arrata, Lepchn.—P. atrata, Lepchn. Iter Sibir. vol. 11. pl. x. f. 9.—H. atrata, Hag. Ent. Ann. 1859, p. 70.—P. altaica, Fisch. Ent. Russ. p. 52, pl. (Neurop.) i. f. 2. Hab. Lapland; Russia; Finland. 14. H. meLaueuca, M‘Lach. post p. 106. Hab. Japan. Note.—The variety of H. Maclachlani, from Japan, indicated under the name of regina, is perhaps a distinct species. I have only seen one female example, lent to me by Baron De Selys Longchamps. It differs from the Indian specimens in the an- terior wings being much narrower, elliptical at the apex; the markings of these wings darker, the apical portion of each wing being black, with few yellow irrorations ; the costal spots elon- gate and not divided; the basal portion of the hind wings much darker, blue-black, this colour extending further along the costal margin; the apical band broader; hence the yellow band is narrower, and there are no spots on the costal portion of this band: beneath, the dark portion of these wings is intensely blue-black. Nevnronra, Leach. 15. N. parDa.is, Walk. Cat. Brit. Mus. Neurop. pt. 1, p.7; Hag. Neurop. N. Amer. p. 250. Hab. Nova Scotia. 16. N. SEMIFASCIATA, Say.—P. semifasciata, Say, West. Quart. Rep. ii. p. 161; Amer. Ent. vol. ii. p. 97, pl. xliv.—N. semifasciata, Hag. Neurop. N. Amer. p. 250.—N. fusca, Walk. Cat. Brit. Mus. Neurop. pt. 1, p. 9.—Ptilostomis Kovalevskii, Kol. Gen. et Sp. Trichop. pt. 2, PeelLoSeplentaetew le Hab. North America. 17. N. postica, Walk. Cat. Brit. Mus. Neurop. pt. 1, p: 8; Hag. Neurop. N. Amer. p. 251. Hab. North America. OF EXTRA-EUROPEAN TRICHOPTERA. 105 18. N. oceLuirera, Walk. Cat. Brit. Mus. Neurop. pt. 1, p. 8; Hag. Neurop. N. Amer. p. 252. Hab. North America. 19. N. concatenata, Walk. Cat. Brit. Mus. Neurop. pt. 1, p. 8.— N. irrorata, Hag. Neurop. N. Amer. p. 249 (nee Fab.). Hab. North America. 20. N. upapponica, Hag.—P. reticulata, var., Zett. Ins. Lapp. col. 1061 (nee Linn.). Hab. Lapiand; Island of Oesel. 21. N. revicutata, Linn.—P. reticulata, Linn. F. S. p. 378.—Ohi- gostomis reticulata, Kol. Gen. et Sp. Trichop. pt. 1, p. 81, pt. 2, pl. v. f. 57.—N. reticulata, Brauer, N. A. p. 44. Hab. North and Central Europe. 22. N. cuaTHRATA, Kol.—O. clathrata, Kol. Gen. et Sp. Trichop. pt. J, p. 82.—H. clathrata, Hag. Ent. Ann. 1859, p. 69. Hab. North and Central Europe. 23. N. oceLuicERA, Walk. Cat. Brit. Mus. Neurop. pt. 1, p. 83 Hag. Neurop. N. Amer. p. 259. Hab. Nova Scotia. 24, N. Srauit, M‘Lach. Trans. Ent. Soc. Lond. 1868, p. 289. Hab. Sweden. 25. N. ruricrus, Scop.—P. ruficrus, Scop. Ent. Carn. p. 690.—N. ruficrus, Brauer, N. A. p. 68.—N. fusea, Steph. Ill. M. vi. p. 234, pl. xxxiv. f. 2.—P. striata, Burm. Hand. vol. 1. p. 935.—Oligotri- cha chloroneura, Ramb. Névrop. p. 473.—Anabolia analis, Kol. Gen. et Sp. Trichop. pt. 1, p. 80. Hab. Europe. Note.—Walker’s three species, NV. fusca, postica, and ocelli- fera, are doubtfully distinct according to the types; the names are here used in accordance with the sense in which Hagen has ap- plied them in his ‘ Neurop. N. America.’ IV. concatenata is very closely allied to VV. lapponica; N. ocelli- gera to N. clathrata and N. reticulata. AGRYPNIA, Curtis. 26. A. picra, Kol. Gen. et Sp. Trichop. pt. 1. p. 79. Hab. North Europe. 27. A. PaceTana, Curt. B. E. pl. dxl.—P. wxgrota, Burm. Handb. vol. ii. p. 935.— Oligotricha strigosa, Ramb. Névrop. p. 473. Hab. North and Central Europe. Note.—Two as yet undescribed species of Agrypnia in Hagen’s LINN, JOURN.—ZOOLOGY, VOL, XI. 8 106 MR. R. M‘LACHLAN ON NEW FORMS, ETC., collection are noticed by name only, viz. A. glacialis, ag., from North America, and A. islandica, Wag., from Iceland. PuRyGaNnea, Linné. PHRYGANEA SORDIDA, nov. sp. P. varie affinis, sed alis anticis latioribus, fusco-griseis fusco plus nebulosis; alee posticz ad apicem anguste fusco-limbate ( ). Long. corp. 73 lin. (=15 mill.); exp. alar. 183 lin. (=39 mill.). Hab. Japonia (in Mus. auct.). Evidently allied to P. varia, and perhaps scarcely more than a form of that species. The insect, however, is more robust and rather larger; the anterior wings broader, more clouded with fuscous, especially in the basal half, which is almost entirely fuscous ; the ground-colour brownish grey, instead of the whitish grey of varia; the hind wings with a narrow, smoky-fuscous, apical margin. ,_The anal parts are similar to those of varia, only that the lateral lobes seem to be larger and more quadrate. I have one female example, from TWakodadi. Honostomis, Mannerheim. HoLosToMIs MELALEUCA, n. sp. H. atra, nitida. Pedes abdomen- que sordide nigri. Alz antice pallide straminez, punctis nigris sat dense conspersz ; posticz albze, maculis duabus costalibus ante apicem ornate, late fusco-limbate (g). Long. corp. 73 lin. (=15 mill.); exp. alar, 24 lin. (=50 mill.). Hab. Japonia (in Mus. Brit.). Head and thorax deep shining black (antennz broken): palpi and legs dull black with a greyish tinge. Abdomen dull black: a long trian- gular superior median lobe, shining black, directed strongly down. wards, notched at the acuminate apex, and bearing, before the apex, a needle-shaped process on either side: penis long, flattened, awl- shaped, testaceous (there are also two small testaceous processes which apparently belong to the app. sup.). Anterior wings very pale straw-colour, rather densely irrorated with small black spots, some of which are confluent and form reticulations ; two larger costal spots near the apex, some larger spots towards the inner margin; the apical margin regularly spotted; veins pale, except where they traverse the black markings. Posterior wings white, subopaque ; a large wedge-shaped black spot on the costal margin above the discoidal cell, the point nearly reaching it; beyond this, nearly at the apex, a second large, irregular, black spot; one or two small black dots near the middle of the costa ; apex and apical mar- gin broadly fuscous, with a semilunate pale straw-coloured mark on the extreme margin in each apical cellule; veins pale.‘ - OF EXTRA-EUROPEAN TRIOHOPTERA. 107 There is one g example in the British Museum, from Hakodadi. The species is evidently allied to H. atrata, Lepchn. (altaica, Fischer), but differs in its black legs, and in the complete, broad, fuscous margin of the hind wings. Fam. LIMNOPHILIDA. GrammMatauLius, Kolenati. GRAMMATAULIUS BREVILINEA, n. sp. G. fusco-niger, subtus griseo- ochraceus ; capite, prothorace, mesothoraceque in medio lurido-rufis. Pedes griseo-flavi; tibiis tarsisque nigro-spinosis. Alze anticw angu- state, elongati, ad apicem vix dilatatze; margine apicali obliquo, paullo exciso; testacese, rufo-brunneo nebulosx, pterostigmate, area suturali cellulaque apicali tertia fuscis, lineis duabus brevibus in area inter- clavali nigris ; postice albids, hyalinz, ad apicem flavescentes ; cellula apicali tertia pallide fuscescente (Q). Long. corp. 8 lin. (=16 mill.); exp. alar. 20 lin. (=43 mill.). Hab. Japonia (in Mus. auct.). Head above lurid reddish, suffused with fuscous in the middle, quite flat, triangularly produced in front, truncate behind; face and palpi testaceous ; eyes black, reticulated with grey, Pronotum large, trans- versely quadrangular, divided in the middle by a longitudinal line, reddish. Mesonotum broadly black at the sides, and with a broad longitudinal reddish middle band. Metanotum black, somewhat piceous. The whole under-surface of the body greyish ochreous. Legs greyish yellow, tibize and tarsi with numerous black spines, anterior femora sometimes fuscescent internally. Abdomen fuscous above, greyish ochreous beneath: in the female are two long, cylindrical, testaceous, divergent, finger-shaped appendices; bencath these a short, broad, up-directed plate, which is deeply excised at the apex, and two large, oval, obtuse, lateral valves (or inferior appendices). (Pl. II. fig. 1.) Anterior wings long and narrow, the costal inner margins nearly parallel, the apex slightly dilated, the apical margin oblique, excised at the sixth apical cell: colour dull testaceous, suffused with pale reddish brown, the apical portion with paler irrorations; pterostigma fuscous, third apical cell fuscous with some pale dots, sutural area fuscous, but leav- ing the extreme inner margin pale; area interclavalis with two short longitudinal black lines; veins testaceous. Posterior wings broad, subhyaline, the apex and pterostigmatical region yellowish ; third apical cellule suffused with pale fuscous; radius crossing the first apical sector at its extremity, forming a fork. (‘The neuration in each of my two examples is irregular: in one the third apical sector in both an- terior wings, and in the right posterior wing, is furcate at its extre- 8* 108 MR. RB. M‘LACHLAN ON NEW FORMS, ETC., mity; in the other this sector divides from, or soon after, its com- mencement, and joims again before the extremity in all the wings, forming along loop). - I have two females from Japan. It is a true Grammataulius, and a very strongly marked species. SrEnoriyLax, (Kolenatt. STENOPHYLAX GENTILIS, noy. sp. S. pallide testaceus. Antenne pedesque testacci; tibiis tarsisque nigro-spinosis. Abdomen supra nigro-terminatum ; appendicibus superioribus parvis, brevibus, fimbri- atis, flavis ; app. inf. sursum directis, fimbriatis, flavis, ad apicem nigro- truncatis, dentatis. Ale antics elongate, gradatim dilatate, pallide flavee, immaculatze, subnitide ; venis flavis; anastomosibus fuscis ; margine apicali anguste obscuriore ; postice pallidiores (¢). Long. corp. 53 lin. (=11 mill.); exp. alar. 17 lin. (=36 mill.). Hab. America boreali (in Mus. auct.). The whole body, including antenne, palpi, and legs, testaceous; tibize and tarsi with black spines; eyes black. The last dorsal segment of the abdomen is conically produced at its apex, which is black and scabrous; app. sup. small, rounded, concave internally, yellow, and fringed with yellow hairs; app. intermed, black, truncate (?) ; app. inf. directed upwards, yellow, fringed externally with long yellow hairs, the apex black and truncate, furnished with small teeth. Anterior wings elongate, broad, the apex parabolic, nearly uniformly pale yellow, almost nude, and shining, the membrane finely rugulose ; inner margin (area suturalis) deeper yellow; apical margin narrowly obscure ; veins yellow, the anastomoses fuscescent ; a whitish dot at the thyridium, and another at the areulus. Posterior wings hyaline, tinged with yellow; anterior margin deeper yellow. T have one male, from the White Mountains of New Hampshire, sent by Mr. H. Edwards, of San Francisco. The species is allied to the European S. hieroglyphicus, striatus, &c., in which the wings are elongate, and the first apical cell in the anterior pair scarcely longer than the succeeding cells. S. LIMBATUS, nov. sp. S. rufo-testaceus. Antenne testacee, fusco- cingulate. Pedes flavi. Abdomen supra fuscum, infra ochraceum ; .segmento ultimo lateraliter productum; app. sup. parvis, subqua- dratis, flavo-fimbriatis ; app. inf. sursum directis, ad apicem truncatis, extus fimbriis longis instructis ; app. intermed. elongatis, spiniformi- bus, rectis, ad apicem abrupte uncinatis. Als anticz breves, late, ad apicem valde obtuse, testacez ; nebulain cellula thyridi (puncto albo ad thyridium incluso) maculis duabus (una ad basin cellule apicalis OF EXTRA-EUROPEAN TRICHOPTERA. 109 secunde, altera quarte) limboque apicali intus dentato pallide brun- neis: posticx hyaline (¢). Long. corp. 43 lin. (=9 mill.) ; exp. alar. 123 lin. (=26 mill.). Hab. Terra Nova (in Mus. auct.). Iead and thorax reddish testaceous, with sparse reddish hairs; antennze testaceous, with fuscous rings; palpi yellowish; eyes black. Legs yellow, tibiz and tarsi with short black spines, a black point on each trochanter, internally. Abdomen fuscous above, ochreous beneath ; margin of last dorsal segment regularly concave in front, produced at the sides into a triangular tooth, the upper edge of which is excised and beset with numerous very short black spiny hairs; app. sup. small, yellow, subquadrate, truncate, fringed with yellow hairs; app. intermed. long, in the form of two closely applied straight spines, the tips of which are suddenly curved downwards; app. inf. directed up- wards, projecting beyond the lateral production of the segment, yellow, truncate at the apex, and fringed externally with long yellow hairs. Anterior wings short and broad, much dilated at the apex; the apical margin oblique, pale testaceous, the membrane finely rugulose, nearly nude, and shining ; a cloud in the cellula thyridii extending also above it, and there enclosing a white dot at the thyridium ; two irregular spots, one placed at the base of the second, the other in a similar po- sition in the fourth, apical cells, and a broad apical margin which is dentate internally (being produced into an acute triangle along each apical cell) pale brown; ramus clavalis margined beneath with brown ; veins testaceous, with short concolorous hairs; first apical cell longer than the second, but not inordinately so. Posterior wings hyaline, whitish, slightly yellowish at the apex; veins pale yellowish; fifth apical cell scarcely reaching the anastomosis. (PI. II. fig. 2.) I have two males, taken at St. John’s, Newfoundland, by Mr. G. F. Mathew. In the form of the wings the species approaches S. dubius, punctatissimus, &c.; but the first apical cell in the an- terior wing is much shorter than in those species. PLATYPHYLAX, nov. gen. Characteres ut in Stenophylact (sensu stricto), sed calcarium formula 1, 2, 2. Agreeing in almost every respect with the typical forms of Stenophylax (e. g. hieroglyphicus, striatus, &c.), but with only 1, 2, 2 spurs instead of 1, 3, 4. _ I form this genus for the reception of some insects that have been placed in Hnecyla on account of their spur-formula being identical (7. ¢. so faras the winged male of Znacyla is concerned), but which are evidently very closely allied to Stenophylax and s 110 MR. R. M‘LACHLAN ON NEW FORMS, ETC., should be placed next thereto. Ihave already (Stettiner ento- mologische Zeitung, 1867, p. 54) separated certain forms with the same number of spurs into a distinct genus under the term Pota- morites ; but these, in the narrower form, and pouched hind wings of the male, come near Drusus. Platyphylax is really so near Stenophylax that, without examining the spurs, the species might pardonably be supposed to pertain to the latter. In Platyphylaxz should be placed the European £. Frauenfeldit, Brauer and H. Kolenatii, Kol. (= Frauenfeldii $?), the North- American £. subfasciata, Say, EL. designata, Walker, and LE. lepida, Hagen, and the Chinese species described below as P. lanuginosus. E.. irrorata, F. (=tntercisa, Walk., Hag.), and EH. preterita, Walk., probably form another genus. #. areolata, Walk., is probably a true Hnecyla; but it is desirable to see the female. PLATYPHYLAX LANUGINOSUS, nov. sp. P. fuscus, abdomine ochraceo. Antenne palpique fusco-nigri. Pedes flavi, tibiis tarsisque fusco- nigris. Altee antics late, testaceo-fuliginose, dense et breviter te- staceo-hirsute ; venz pilis erectis fuscis fimbriatz : postice fuliginoso- subhyalinz, margine costali apicem versus flavido ( @ ). Long. corp. 7 lin. (=15 mill.); exp. alar. 18 lin. (=37 mill.). Hab. Shanghai (in Mus. auct.). Head fuscous above, posterior margin and a small tubercle on each side close to the eyes testaceous ; ocelli white; antennx blackish, the basal joint with blackish hairs, a few testaceous ones being mtermin- gled; face ochraceous; palpiblackish. Thorax fuscous above, ochra- ceous beneath ; posterior half of metanotum yellowish. Legs: cox, trochanters, and femora testaceous ; tibize and tarsi fuscous, armed with numerous short blackish spines. Abdomen ochraceous ; at the apex are two short and obtuse appendices (my individual carries at the ex- tremity of its abdomen a dried mass of gelatinous matter, such as en- velopes the eggs). Anterior wings broad, the apical margin oblique, somewhat sinuate, and narrowly darker; the colouris smoky with a testaceous tinge; and there is a uniformly dense, almost woolly clothing of short procum- bent testaceous hairs, intermingled with which are short, erect, blackish hairs; and on the veins, especially on the cubitus, are longer, erect, blackish hairs ; a white dot at the thyridium, and another at the ar- culus ; veins pale fuscous ; first to fourth apical cells all more or less truncate at the base, fifth acute, scarcely reaching the anastomosis, furnished with a short footstalk ; a black horny dot at the base of the third apical cell. Posterior wings smoky subhyaline; veims blackish- fuscous ; apical portion of costal margin, and the subcosta and radius at that portion yeliowish. T have one female, from Shanghai, taken by Mr. W. B. Pryer. OF EXTRA-EUROPEAN TRICHOPTERA. 111 NEOPHYLAX, gen. nov. Calcaria 1, 2,4. Als antics dense pubescentes, apicem versus gradatim dilatate, margine apicali sinuato ; cellula discoidali elongata : postice cellulis apicalibus 5 instructe. Abdomen infra apicem versus dentibus duobus instructum (3). Head. Antenne about the length of the wings, moderately short, the basal joint longer than the head. Eyes large. Ocelli present. Max- wlary palpi with short and oval basal Joint; second joint long, gradually thickened; third joint rather shorter than the second, cy- lindrical: labial palpi with two short and thick basal joints, and a longer, slender, and cylindrical terminal joint. Thorax short. Ab- domen slender: penultimate and antepenultimate seginents each fur- nished with a tooth beneath: appendices little prominent. Legs mo- derately long; tibize and tarsi with few spines: spurs 1, 2, 4; the inner subapical spur on the posterior tibia very small, scarcely more than a tooth-like tubercle; the other pairs subequal. Anterior wings clothed with dense short pubescence, and with short fringes ; narrow at the base, gradually widened to the apex ; apical margin oblique, slightly emarginate in the middle of the margin of the fourth apical cell, elevated at the point of termination of upper branch of the fork of the ramus thyrifer, and afterwards gradually emarginate to the anal angle, which is rounded; discoidal cell very long and narrow, closed ; apical cells long and narrow, the first, third, and fifth acute, or subacute, at the base, and longer than the second and fourth ; radius strongly bent before its termination. Posterior wings broad, the fringes long at the anal angle; subcosta and radius run- ning very close together for more than half their length, then becom- ing confluent, or nearly so, afterwards disuniting, the radius then curved ; ramus subdiscoidalis simple; hence there are only five apical cells (three apical and two subapical, according to the nomenclature of Kolenati); discoidal cell broad, closed (¢). A singular genus, which should probably be placed near Apa- tania, with which it agrees in its spur-formula and densely pubes- cent anterior wings. The shape of the anterior wings is peculiar, and the neuration of the posterior wings very remarkable in the small number of apical cells, in this respect unique in the family Limnophilide. NEOPHYLAX CONCINNUS, nov. sp. N. testaceus. Pedes nigro-spi- nosi. Ale: antice fulve, fusco-pubescentes, punctis albidis obsoletis irrorate; margo dorsalis maculis tribus flavis ornatus ; ciliis apica- libus fuscis, albido-interruptis ; posticae fumato-subhyalinte. Long. corp. 33 lin.(=7 mill.); exp. alar. 93 lin. (=20 mill.), Hab. America boreali (in Mus. auct.). 412 MR. BR. MSLACHLAN ON NEW FORMS, ETC., Head (with the antennz, palpi) and thorax testaceous; occiput, basal joint of antennz, and prothorax clothed with testaceous, with an ad- mixture of fuscous, hairs; and there is a fringe of similar hairs on the facial margin of the eye-sockets; eyes dark coppery. Legs testa- ceous, the posterior tibie paler; tibize and tarsi with few, short, black spines ; spurs reddish-testaceous. Abdomen pale whitish testa- ceous; on the antepenultimate ventral segments is a very small, reddish-testaceous, triangular tooth, and on the penultimate segment a much Jarger tooth ; appendices testaceous ; app. sup. small, rounded, and ear-shaped, extending little beyond the cavity of the last seg- meni; app. intermed. placed close together, proceeding from under the middle of the upper margin of the segment, nearly straight, and flattened laterally; when viewed from the side each appendage is seen to be dilated at the base, then with the upper margin excised to the apex, which is obtuse; app. inf. inserted close together on the ventral margin, band-like, curved strongly inwards, forming a deep incision when viewed from beneath, the apex obtuse. Anterior wings fulvous, thickly clothed with short, procumbent, fuscous pubescence, the apical half irrorated with many small and indistinct whitish dots; inner margm with three yellow spots, viz. an elongate one at the base, a long triangular one about the middle, and a small” one before the anal angle; the pubescence in the spaces between these spots is darker, almost blackish fuscous ; apical fringe alter- nately fuscous and whitish; veins testaceous, the costal margin at the base, and the basal portion of the radius, ciliated with fuscous. Posterior wings subhyaline, slightly smoky ; the fringes at the anal angle very long, silky, and whitish. (PI. II. fig. 3, details.) T received one male example from Mr. J. Angus, of the State of New York. fam. SERICOSTOMATID A. Norrposta, Stephens. NoTipoBIA GRISEOLA, nov. sp. WN. nigro-fusca. Caput protho- raxque cinereo-hirsuta: antenna palpique fusci. Pedes flavescentes, antici omnino, femoribusque intermediis posterioribusque mterdum _ fuscis. Abdomen fuscum, cinereo-hirsutum, linea utrinque pallida : appendices inferiores ¢ magna, truncata, supra in dentem uncina- tum incurvatum intus producti. Ala anticw grisew, dense cinereo- pubescentes : postice pallidiores (3, 9). Long. corp. d 23 lin. (=6 mill.), Q 27-4 lin. (=63-83 mill.) ; exp. alar. 9-123 lin. (=19-263 mill.). Hab. California (in Mus. auct.). Biackish fuscous. Head and prothorax clothed with whitish ashy-grey hairs, changing to fuscous on the face; antenne fuscous, paler and OF EXTRA-EUROPEAN TRIOHOPTERA. 113 somewhat yellowish in some individuals; palpi fuscous, the maxillary pair in the 3d very small and applied against the face ; hinder mar- gins of meso- and metanota yellowish. Legs yellowish; the ante- rior pair altogether fuscous, which colour sometimes pervades also the intermediate and posterior femora; coxe blackish-fuscous. Ab- domen fuscous, sometimes yellowish, with a pale line along each side, and clothed with ashy-grey hairs: appendices of 3 yellow; app. inf. large, longer than broad, the apex truncate, the superior edge rounded, the apical margin furnished with an acute claw- shaped spine, which is turned inwards, these appendices fringed with yellowish-grey hairs; penis slender obtuse, straight, notched at the apex above. In the Q the abdomen is depressed, and at the apex is a large oval pouch, which is usually filled hy a dark olive-green mass of eggs; but when empty the upper portion is seen to be furnished with a broad median lobe, on each side of which is a somewhat tri- angular valve. Anterior wings grey, densely clothed with ashy-grey pubescence, mingled with fuscous; in the ¢ there are two or three small elongate spaces of white pubescence on the inner margin, and an indication of whitish dots in the discal and apical portion of the wing; fringes grey. Posterior wings paler grey, with long grey fringes at the anal margin. (Pl. IL. fig. 4, details. ) I possess two males and five females, from California, sent by Mr. Henry Edwards; the females vary very much in size and comparative robustness, the larger individuals having the legs and antenne darker; but all seem to pertain to one species. It is is a true Wotidobia, as is the following species, and in structure is quite identical with the typical WV. ciliaris of Europe. _ NoTIpOBIA NIGRICULA, noy. sp. N. nigra. Caput thoraxque aureo- hirsuta; antennz palpique fusci. Pedes flavo-fuscescentes, anteriores obscuriores. Abdomen fusco-nigrum, nigro-hirsutum : appendices inferiores magni, late, excise, supra in dentem incurvatum intus producte. Als fuliginoss, sparse brunneo-pubescentes : anticarum pterostigma flavum (¢). Long. corp. 3 lin. (=63 mill.); exp. alar. 93 lin. (=20 mill.). Hab. California (in Mus. auct.). Dull black. Head and prothorax above clothed with golden-yellow hairs; on the face the hairs are mostly blackish; antennze fuscous ; palpi fuscous, clothed with fuscous hairs, the maxillary pair small, curved upwards, and closely applied against the face ; eyes brown, somewhat coppery. Hinder margin of the meso- and metanota yel- lowish and shining. Legs obscure yellowish, with fuscous pubescence ; all the coxe blackish, and the anterior femora, tibia, and tarsi dark fuscous; spurs yellow, the pair on the anterior tibie fuscous. 114 MR, R. M‘LACHLAN ON NEW FORMS, ETO., Abdomen blackish fuscous, clearer beneath, clothed with black hairs: last segment above fringed with long, blackish, curved hairs: inferior appendices very large, longer than broad, yellow, concave internally, the superior margin rounded, apical margin deeply excised, superior angle produced into an incurvated tooth turned inwards, these ap- pendices clothed externally and fringed with long blackish hairs ; penis subobtuse, the point visible below the app. inf. (Pl. II. fig. 5.) Anterior and posterior wings uniformly fuliginous, subdiaphanous, - clothed, but not densely, with short brownish pubescence, which becomes somewhat golden on the costal margin of the anterior wings ; and in these wings the pterostigma is indicated by a narrow yellow space; fringes brownish-grey, becoming pale grey towards the anal angle of the posterior; veins fuscous. I have one male, from California, sent by Mr. Henry Edwards. It differs from WV. griseola by the uniform smoky colour of the wings, and in the form of the appendices. Nosoprvs, gen. nov. Calcaria 1, 4,4. Antennarum articulus basalis elongatus, hir- sutus. Palpi maxillares parvi, ad frontem arcte applicati: labiales valde elongati, compresso-dilatati, squamati ; articulo basali parvo, 2° elongato, dilatato, 3° ad apicem acuminato. Pedes antici tibia brevissima calcare singulo uncinato -in- structa, tarsorum articulus basalis valde dilatatus, intus sul- catus, infra dense cerato-squamatus: intermedii posticique graciles. Als antic ovales, hirsute; cellula discoidali oc- clusa, angustata, cellula thyridii perelongata; cellulis sep- tem apicalibus: postice in medio dilatate; cellula discoi- dali parva, occlusa (¢ ). Head deusely clothed with long hairs; antenne not so long as the wings, moderately stout, the apical half subserrate imternally, basal joint nearly twice the length of the head, strong, hirsute, the suc- ceeding joints short and transverse ; eyes small and round; maxil- lary palpi very small, somewhat clavate, directed upwards and lying closely applied against the face, clothed externally with long and strong hairs; labial palpi very large and long, densely clothed with scales, the basal joint short, second very long, compressed and di- lated, third about as long as the second, and equally broad at the base, but gradually acuminate to the apex. Legs: anterior pair ab- normally constructed as follows :—the coxa elongate, and ordinary ; the trochanter small and cup-shaped ; femur long, moderately slender, gradually diminishing from base to apex; tibia very short, sub- OF EXTRA-EUROPEAN TRICHOPTERA. 115 ovate, truncate, and dilated, slightly scaly, armed with one stout, claw-shaped spur ; first joint of tarsi enormously dilated, twice the length of the tibia, sulcate internally, the lower surface densely fur- nished with waxy-looking scales; succeeding tarsal joints short and small, gradually diminishing in length and thickness; intermediate and posterior legs slender, and of the ordinary form, each tibia furnished with an apical and subapical pair of long and equal spurs. Abdomen short and somewhat stout; inferior appendices short, curved. Anterior wings oval, rather densely clothed with short hairs, the fringes somewhat long; subcosta and radius nearly straight, parallel; dis- coidal cell narrow, closed by a straight veinlet; cellula thyridii very long, extending nearly to the base, and reaching to the middle of the discoidal cell, closed by a straight veinlet ; a veinlet unites the lower fork of the ramus discoidalis with the ramus thyrifer, placed level with that closing the discoidal cell; an oblique veinlet beneath the middle of the cellula thyridii unites this with the cubitus anticus; seven apical cellules, the first extending along one-third of the upper edge of the discoidal cell, third shorter than the first, but longer than the second, fourth equal to the second, fifth longer than the first, extending to a level with the middle of the discoidal cell. Posterior wings moderately long, gradually dilated to beyond the middle, apex parabolic, costal margin with a short inturned fringe, anal portion with very long fringes; subcosta and radius united for some dis- tance, afterwards separating and diverging; discoidal cell small, sub- triangular, closed by a straight veinlet ; a second veinlet unites the lower edge of the discoidal cell to the ramus subdiscoidalis ; lower branch of the ramus discoidalis simple; ramus subdiscoidalis simply and longly furcate. A genus abundantly distinct by the enormous labial palpi, and very abnormal structure of the anterior legs, the aborted tibia and enormous first tarsal joint in these legs being very re- markable ; the mass of scales on the surface of this strange tarsal joint has, at first sight, the appearance of a waxy secretion, but resolves itself into waxy-looking scales under a high power. The genus is evidently a near ally of Mormonia; and nature would seem to have selected this group as one in which she can best display her wealth of forms. In this group is also exhibited a more or less constant tendency to substitute a scale-like clothing for hairs in the male sex. In the typical species of Mormonia ' (M. hirta) this clothing pervades almost the entire insect; in Nosopus it is concentrated, so to speak, upon the labial palpi and the abnormal tarsal joint. It is possible, nay, almost certain, 116 MR. R. M‘LACHLAN ON NEW FORMS, ETO., that the female will be found to have ordinary palpi, and the usual slender anterior legs; and, in all probability, 2, 4, 4 spurs ; for one spur may be reasonably supposed to be aborted in the anterior male tibia. Nosopus PODAGER, nov. sp. N. fuscus. Caput griseo-hirsutum. Antenne flavide, fusco-annulate, articulo basali supra griseo, infra nigro-hirsuto. Palpi maxillares nigro-hirsuti; labiales rufo-squamati. Pedes antici rufo-fusci, tarsorum articulo 1° infra rufo-squamato : in- termedii posticique testacei. Abdomen fuscum. Ale griseo-fusce, subhyaline, griseo-hirsute : antice ad costam marginisque interioris basin breviter nigro-fimbriatz (¢). Long. corp. 3 lin. (=6 mill.) ; exp. alar. 9 lin. (=19 mill.). Hab. California (in Mus. auct.). Dark fuscous. Head above, and basal joint of antenne, clothed with grey hairs, face and maxillary palpi with black hairs; antennie (ex- cept the basal jot) yellowish, with narrow fuscous rings; labial palpi densely clothed with reddish scales. Anterior legs reddish fuscous, the lower and outer side of the first tarsal joint with dense waxy-looking reddish scales; intermediate and posterior legs testa- ceous, the coxze fuscous. Abdomen fuscous, the margins of the seg- ments greyish: from beneath the upper margin of the last dorsal segment proceeds a shost, broad lobe, which ends im two updirected triangular pointed branches; app. inf. short, band-like, curved im- wards, the apex toothed. Wings greyish-fuscous, sparingly clothed with dark grey hairs: in the anterior wings the pterostigmatical region with denser hairs; apical fringes grey; costal margin, and inner margin at the base, with short blackish fringes : posterior wings with grey fringes, becoming blackish at the base of the costa: veins pale grey in all the wings. (PI. IT. fiz. 6, details.) I have one male, sent by Mr. H. Edwards. DINARTHEUM, gen. noy. Calcaria 2,4,4. Antennarum articulus basalis rectus, longissi- mus, corporis longitudini «#qualis, irregulariter compressus, utrinque fimbriis longis, necnon ad basin spina robusta instructus; articuli ceteri breves, graciles, basalis semel sumpti longitudinem haud superantes. Palpi maxillares elongati, porrecti, plumosi, 2-articulati; articulo 1° modice robusto, 2° gracili, curvato: labiales pergraciles, elongati ; ar- ticulo 1° brevissimo, 2° elongato recto, 3° 2° «quali, curvato. Pedes graciles. Al antice ovales, squaiate hirsutaque, sulco longitudinali elongato angustato mediano instructs; mar- OF EXTRA-EUROPEAN TRICHOPTERA. 117 gine costali intus fimbriato, margine apicali fimbriis longis in- structo; cellula discoidali occlusa, venis irregularibus (¢ ). Head very small: eyes small and round: first joint of the antenne extraordinarily long, equal to, or exceeding, the length of the whole body, standing out straight from the head, strong, compressed, but irregular in its breadth, arising from a prominence on the head, fur- nished internally at the base with a strong curved spine, which is widened and truncate at the apex, the basal portion clothed with stout hairs, afterwards fringed with two rows of very long slender hairs ; the rest of the antenne not exceeding the length of this first joint, articulated to it almost.at a right angle, strongly curved, com- posed of numerous short and slender joints: maxillary palpi appa- rently 2-jointed (or there is perhaps a short, but invisible, basal joint), long, extending far beyond the head ; first joint long, porrect, band-like, almost geniculate at the base, afterwards bent, strongly plumose and scaly; second joint equalling the first in length, but slender and curved: labial palpi very slender ; first jomt very short, scarcely visible ; second joint long, cylindrical, straight ; third joint equal to the second, curved. Thorax small. Legs slender, spurs 2, 4, 4, moderately long and subequal, hairy, those on the anterior tibia less equal and more hairy than the others; tarsal joints long and slender. Abdomen short; inferior appendices long. Anterior wings oval, scarcely dilated, the upper half thinly clothed with scales intermingled with hairs, the lower half with hairs only : these two divisions are separated by a deep, narrow, longitudinal groove, extending from near the base almost to the apex, containing more closely placed scales; the groove is placed between the ramus thyrifer (which it nearly obliterates) and the ramus clavalis, this latter - being very strong and furnished with an updirected fringe of long seale-like hairs, which form a cover over the groove; subcosta and radius straight and subparallel; discoidal cell short, narrow, closed ; no apparent cellula thyridii; the neuration in the lower half of the wing (below the groove) irregular, forming five large irregular cells: costal margin with a long inturned fringe in its basal half; apical margin with very long fringe. Posterior wings short, scarcely so broad in their widest part (which is beyond the middle) as the antcrior, the apex elliptical; hairy and with a few scales on the costal margin ; fringes very long: neuration regular; subcosta and radius confluent for the greater part of their length, afterwards separating and di- vergent, forming a long apical fork ; discoidal cell very similar to that in the anterior wings, closed: lower branch of ramus discoidalis simple ; ramus subdiscoidalis simply forked, connected with the ramus discoidalis by an oblique veinlet beneath the discoidal cell. Like the last genus, allied to Mormonia, or perhaps more nearly 118 MR. R. M‘LACHLAN ON NEW FORMS, ETC., to Lasiocephala, with which it has some affinity in the maxillary palpi; but the extraordinary form of the antennex is without parallel anywhere in the Trichoptera. In Wosopus the anterior legs were the members in the construction of which nature had departed from her usual routine; here the legs present no special characters, but every thing is thrown into the development of the antenne, with a result which, to say the least, is bizarre. The groove or pouch of the anterior wings is not of so great signi- ficance, as modifications of this already exist in many genera, and where such a groove is present, it often, as in the present instance, causes irregularity in the neural arrangement. The female will probably be found to have ordinary antenne, and to resemble that of Mormonia. DINARTHRUM FEROX, sp. nov. JD. fusco-testaceum. Antennarum articulus basalis fuscus, fimbria grisea, ad apicem flavescente ; articuli cxteri pallide flavi, fusco-cingulati. Palpi flavi, maxillares griseo- hirsuti nigroque squamati. Pedes pallide flavi. Abdomen supra fuscescens, infra ochraceum: appendices inferiores ad apicem longe dentate. Alse pallide grisez, griseo-fimbriatz, nigroque squamatie : antics margine costali basin versus sulcoque nigro-fimbriatis (¢). Long. antenn. artic. primi 27 lin. (=6 mill.); long. corp. 23 lin. (=6 mill.); exp. alar. 10 lin. (=2I1 mill.). Hab. in India septentrionali (in Mus. auct.). Fuscous, or yellowish-fuscous, all the under parts of the thorax yellow. Head fuscous above, yellow beneath, clothed with greyish fuscous hairs: basal joint of antennz fuscous, becoming yellowish towards the apex, the basal portion of the tooth almost black and somewhat shining, fringes dark grey, blackish at the basal portion, and yel- lowish at the apical; rest of the antenne pale yellow, with brown rings: maxillary palpi yellow, clothed with long dark grey or blackish hairs, intermingled with a few black scales; labial palpi pale yellow. Legs yellow, with yellow spurs. Abdomen somewhat fuscous above, the margins of the segments broadly darker, under surface ochra- ceous: the margin of the last dorsal segment is produced into a tri- angular prolongation in the middle; from beneath this prolongation proceeds a yellow, shining, triangular lamina, which is deeply grooved in the centre, the sides sloping obliquely upwards, having the appearance of two valves soldered together; app. sup. yellow, short and broad, subquadrate, proceeding from beneath each side of the prolongation of the last dorsal segment; app. inf. long, yellow, directed up- wards, the apex furnished with two long spines or teeth, each of which is as long as the simple basal portion, the appendices are hairy ; interiorly, viewed from beneath, between the app. inf. are OF EXTRA-EUROPEAN TRICHOPTERA. 119 seen two small yellow appendices, which are probably the app. intermed. Wings pale grey, with long grey fringes. Anterior wings clothed with golden-grey hairs, the upper portion (above the groove) with nu- merous, slightly attached, black scales; these scales are absent in the lower portion; costal margin with a strong inturned fringe of blackish scale-like hairs at the base, and a fringe of similar hairs on the ramus clavalis closing over the groove ; veins yellow, especially those in the lower portion, the apical ones fuscescent. Posterior wings clothed with grey hairs, and with scattered black scales on the anterior margin; veins fuscous. (PI. II. fig. 7, details.) I have two males of this extraordinary creature, which were given to me by Capt. A. M. Lang, R.E., by whom they were captured in North India. Fam. LEPTOCERIDA. PERISSONEURA, gen. nov. Calcaria 2, 4,4, pubescentia. Caput transversum, inter oculos excayatum ; ocelli desunt: antenne graciles, articulo basali bulboso: palpi maxillares elongati, hirsuti, articulo basali brevi, ceteris elongatis inter se longitudine fere equalibus : labiales parvi. Thorax robustus. Pedes graciles. Abdomen robustum. Alx ample, latw, ad apicem obtuse, pubescentes; venis ro- bustis: anticarum radio cum sectore apicali 1° juncto, venulis transversalibus in area costali plurimis, quarum una furcata ; anastomosis ante medium sita; cellula discoidali elongata, occlusa, ante apicem venula transversa insititia instructa ; cel- lulis apicalibus decem, elongatis: posticw anticis haud la- tiores (¢). Head transverse, polished, excavated between the antennz ; no ocelli; eyes moderately prominent ; antenne not longer(?) than the wings, slender, the basal joint bulbous; maxillary palpi long, hairy, ascend- ing, basal joint very short, second and third joints long, equal, cylin- drical, stout, fourth and fifth slightly shorter and thinner, the latter obtuse at the apex; labial palpi small, hairy, first joint very short, second and third longer, equal. Prothorax small and transverse, hairy. Mesothorax robust, convex above, polished. Legs slender, pubescent ; spurs 2, 4, 4, pubescent; anterior tibia with a pair of moderately long and subequal spurs; intermediate and posterior tibixe each with two pairs of long and subequal spurs. Abdomen very stout, long; the apex beneath forms a polished, flattened sur- face, on which is seen an oval scale on each sideof the vulva, and beyond these two acute valves. 120 MR. R. MSLACHLAN ON NEW FORMS, ETC., Wings broad, clothed with moderately dense, short pubescence; the veins very strong and much elevated. Anterior wings much dilated at the apex, which is obtuse, the apical margin obliquely rounded, the inner margin very concave; subcosta straight, running into the costal margin; radius parallel with the subcosta, running into the first apical sector near the apex, joined to the subcosta by a short transverse veinlet; costal area with the usual basal veinlet, followed by a broadly furcate veinlet, and after this by 4-5 oblique veinlets, all strong and well marked; first apical sector near its base joined to the radius by a veinlet; discoidal cell long, its apical quarter nar- rowed after the points of departure of the first apical sector, a trans- verse veinlet at about the point of departure of this sector, and another beyond, at the point of furcation of the lower branch of the ramus discoidalis; cellula thyridii very long, extending nearly to the base, gradually dilated to the point where it is closed by a transverse veinlet ; anastomosis complete, situated before the middle of the wing; apical cells ten in number, very long, the fifth and seventh not reaching the anastomosis, and acute at the base. Posterior wings scarcely so broad as the apical portion of the anterior, obtuse, the apical and inner margins gently rounded, costal margin folded nar- rowly inwards for the greater part of its length; subcosta and radius separated only at the base and apex; discoidal cell shorter than in the anterior, similarly formed, but without the supplementary inner veinlet ; forks one, two, three, and five all present; costule nume- rous; cubitus furnished with a fringe of strong oblique hairs, which lie close to the membrane beneath it: marginal fringes scarcely pre- sent in either pair of wings (@ ). In no other genus am I aware of the existence of the numerous strong costal veinlets here present. It is true that the species of the anomalous family Céistropside (Polymorphanisus &c.) present an analogous character ; but in them these veinlets are ill-developed, and have been aptly termed by Brauer, the founder of the family, “false veinlets.” Neither am I aware of the exist- ence of a supplementary veinlet in the discoidal cell in other genera. Although I place the genus in the Leptoceridex, I am by no means sure of its position, which can only be decided by the discovery of the male. In fact, several points of structure rather indicate that its true location would be in the Seri- costomatide, in the vicinity of Barypenthus and Musarna. ‘The form of the maxillary palpi of the female is not inconsistent with its position in either Leptoceride or Sericostomatide. PERISSONEURA PARADOXA, n. sp. P. atra, capite thoraceque ni- tidis. Pedes picei, genibus calcaribusque testaceis. Abdomen sor- OF EXTRA-EUROPEAN TRICHOPTERA. 121 dide nigrum ; linea utrinque laterali ochracea ; segmentis apicalibus ventralibus flavo-marginatis. Al fusca, subnitide, nigro-pube- scentes ; venis piceis (2 ). Long. corp. 73 lin. (=15 mill.) ; exp. alar. 22 lin. (=46 mill.). Hab. Japonia (in Mus. Dom. De Selys). Head and thorax deep shining black; eyes greyish ; antenne blackish; palpi yellowish, clothed with blackish hairs; prothorax above clothed with black hairs, beneath forming a pale yellow space extending between the anterior coxz, and on each side of these. Legs pitchy, the anterior pair paler, somewhat testaceous; the knees and spurs testaceous. Abdomen dull black, an ochreous line along each side, and the margins of the terminal ventral segments yellow; terminal portion beneath flattened, smooth and yellow: vulvar scales large, oval, piceous, lying closely applied to the surface ; beyond these are two acute triangular yellow valves, the tips blackish and extending somewhat beyond the apex of the abdomen. Wings uniformly fuscous, somewhat shining, clothed with blackish pubescence, which is more dense on the posterior wings and on the costal portion of the anterior; in the anterior pair, beyond the ana- stomosis, is an appearance of an indistinct whitish discal space; the veins all piceous. (PI. II. fig. 8, details.) For the opportunity of examining and describing this curious insect I am indebted to the courtesy of my friend the Baron de Selys Longchamps, the learned monographer of the Odonata. AscararnomErvs, Walk. (Cat. Brit. Neurop. pt. 1, p. 79). Calearia 2, 4,4. Caput: § oculi permagni, supra fere con- nexi; Q oculi parvi, distantes ; vertex transversus, cirris in- structus : ocelli desunt: palpi maxillares teniiformes, valde hirsuti, articulo basali brevi, 3° valde elongato, 2°, 4°, et 5° brevioribus ; labiales parvi, graciles: antenns valde elongati, in 2 breviores, cylindrici, graciles, vel interdum in ¢ pra- datim crassiores, articulo basali brevi. Corpus robustum, breve. Pedes graciles. Ale fere nude : antice elongate, api- cem versus dilatate, margine apicali obliquo; ven» robustze, radio cum sectore primo paullo ante apicem conjuncto (et in al. post); cellula discoidali brevi, occlusa; cellula thyridii permagna, ante ale medium sita ; cellulis apicalibus elongatis, angustatis: postice elongato-ovales, anticis fere dimidio bre- viores. Head of the 3 occupied almost entirely by the eyes, which are very large, and nearly confluent above: that of the 9? with the eyes small; the vertex transverse, furnished with large tubercles, whence LINN. JOURN.—ZOOLOGY, VOL. XI. 9 122 MR. R. M‘LAOHLAN ON NEW FORMS, ETC., arise tufts of long hairs: ocelli absent: antenne very long and slender, but sometimes gradually thickening to the apex ; basal joint short, succeeding joints (after the second) long, but those of the apex becoming gradually shorter and almost transverse: maxillary palpi ascending, the joints band-shaped and very hairy; first jot short, third very long, second, fourth, and fifth each shorter than the third, but the second longer than the fourth or fifth: labial palpi very small and slender, the two end joints elongate. Mesothorax very robust, long-oval, nude, but with a tubercle near the point of con- nexion of each anterior wing, whence arise long hairs similar to those on the vertex. Abdomen short, very robust in the 2. Legs slender, pilose; spurs 2,4, 4, the pairs subequal. Wings nearly nude and shining, the veins very conspicuous and strong, alike in both sexes. Anterior wings elongate, dilated towards the apex, which is considerably produced; apical margin very oblique ; inner margin concave; radius becoming confluent with the first apical sector a little before the apex, but sending a short branch to the costal margin; discoidal cell closed, short, elongately triangular, no veinlet between this cell and the radius; cellula mediana longer than the discoidal, equal to it at its extremity, but extending further inwards at its base; cellula thyridii very broad, commencing near the base, but not extending to the middle of the wing (ending before the commencement of the discoidal cell), hence the sixth to ninth apical cells extend far into the wing; all the apical cclls narrow and very long. Posterior wings elongately oval, much shorter than the ante- rior wings, and scarcely broader: radius confluent with the first sector, as in the anterior, and the subcosta also appears to be con- fluent with the radius at its apex; the cubitus is furnished with a fringe of long hairs towards the base, and the veins of the anal angle are similarly fringed. The appendices of the male are complicated: the app. sup. rather long, narrow at the base, but gradually dilating into a spoon-shaped club; the app. inf. two-jointed, the apical joint being short and ovate; between the app. sup. are two large blades, nearly uniting in the middle, but with the obtuse points divergent ; and between and below these there is the penis, only the apex of which is visible. In the female the extremity of the abdomen is broad, forming a large open pouch with two broad side valves and a median prolongation of the last dorsal segment ; the apical ventral segments are narrow and transverse. The figures on Pl. III. (fig. 9) are taken from A. finitimus, M‘Lachlan. The original species, A. humeralis, Walker, is larger and darker, and, with the appendices somewhat different, though OF EXTRA-EUROPEAN TRICHOPTERA. 123 formed after the same plan. In both, the antennz of the female are much shorter than those of the male, in humeralis remarkably so. In finitimus these organs are slender throughout in the ¢, whereas in humeralis they are gradually incrassate in the apical portion. A completely analogous formation of the eyes in both male and female is to be found in the micro-lepidopterous genus Adela. I have diagnosed and described this extraordinary genus anew, because the description given by Mr. Walker is insufficient. It appears to me that, having regard to the homologies of the arrangement of the nervures in Trichoptera, an error has been frequently committed, of which I have myself been guilty in some previous descriptions in former papers. I allude especially to the area or cell which has been called the “cellula thyridii ” in the anterior wing. To Kolenati we are indebted for a very lucid explanation of the different veins, areas, and cellules; and in his index wing (Gen. et sp. Trichop. pt. 1, tab. i. f. 1), taken from Glyphotelius, the “ cellula thyridii ” is the area between the two veins which he terms “ radii ramus thyrifer,” and “radii ramus clavalis,” and which I have called (Trichop. Britannica) the “ su- perior and inferior branches of the ramus thyrifer.”” But in almost all genera of Hydropsychide, and in several genera of Lepto- ceride, e. g. Ascalaphomerus, Anisocentropus, Ganonema, Asoto- cerus, Calamoceras, Heteroplectron, &., the superior branch of the ramus thyrifer (“ radii ramus thyrifer”” of Kolenati) furcates near the middle of the wing; and this furcation is generally closed by a transverse veinlet placed nearly on a level with that closing the discoidal cell, and forming a cellule, which has been usually termed the “cellula thyridii,” though the true cellula that should be so called, equivalent to that in the Limnophilids, &e., lies beneath it, extending to near the base, and usually ending soon after the commencement of the cellule formed by the connected ramules of the superior branch. This latter cellule, then, I propose to call the “cellula mediana.” A reference to the outline figures of the wings of any of the above-named genera will more fully explain my meaning. In Mr. Walker’s description of the genus Ascalaphomerus the words “cell of the thyridium” should then read “median cel- lule ;”” and his “ interclaval areolet ” is in reality the true “ cel- lula thyridii.” Hrrrroprecrron, gen. noy. Calearia 2, 4,2, 3; 2,4, 4, 2. Maris tibie postice fimbriis 9* 124 MR. BR. M‘'LACHLAN ON NEW FORMS, ETC., longis extus instructw. Antenne alis paullo longiores, intus subserrate. Palpi maxillarese longati; articulo 1° brevi, 2° et 3° elongatis, 4° et 5° brevioribus. Als antice ample, elongato-triangulares: pube brevi dense vestite; margine apicali obliquo ; venis ¢ 2 xqualibus; cellula discoidali oc- clusa, elongata, angustata; cellulis apicalibus novem angu- statis, quarum 1* ad basin cellule discoidalis fere extensa: postice breves, late; radio cum sectore primo ad apicem conjuncto (3 2). Head. Antennz stout, tapering to the apex, rather longer than the wings; the basal joint short, subglobose ; third and succeeding joints long (the third longer than the others); each, at its apex internally, carries one or two short spines or bristles, causing the antenn to appear somewhat serrated, but these are absent in the extreme apical portion. Eyes small. Maxillary palpi long, stout, hairy: basal joint short; second very long; third slightly shorter than the second, and thinner; fourth and fifth each about one-third shorter than the third. Labial palpi small: basal joint very short; second long, third still longer than the second. Prothorax very small; on each side beneath is a semicircular lobe. Mesothorax oval, elongate. Metathorax cordate. Abdomen short, moderately stout; ¢ superior and inferior appendices present; penis (or upper penis-cover ?) long, triangular; 9, apex of abdomen oblique, forming a shallow cavity beneath. Legs: anterior pair short; intermediate and posterior very long; posterior tibiz of the dg fringed externally with very long silky hairs; tarsal joints long ; in the ¢ the first two joints have, ex- ternally, long bristle-like hairs, which, becoming agglutinated, have the appearance of long spines. Spurs: ¢, 2,4, 2, the pairs sub- equal, those on the anterior tibiz shorter; 9, 2,4, 4. Wings. Anterior wings ample, elongately triangular, the apical mar- gin oblique; densely clothed with short pubescence; frmges very short : subcosta and radius regular, straight, connected towards the apex by a transverse veinlet; discoidal cell very long and narrow, closed ; median cell longer than the discoidal, and broader; cellula thyridii commencing near the base and extending to the middle of the median cell; nine apical cellules, all of which are long, the first extending to two-thirds the length of the discoidal cell, first, third, fifth, seventh, and ninth acute at the base, second, fourth, sixth, and eighth truncate. Posterior wings short and broad, the costal margin rounded up to near the apex, the apical portion being slightly excised ; apex obtuse; radius becoming confluent with the first apical sector before the apex; neuration otherwise much as in the anterior wings, allowing for the usual differences; the discoidal cell is perhaps open in these wings (the possible position of a trans- OF EXTRA-EUROPEAN TRICHOPTERA. 125 verse veinlet closing it, is marked in the figure by a dotted line; but I could not clearly determine it). It is an almost invariable rule in Trichoptera that the number of spurs on the posterior tibiw shall equal, or exceed, that of the intermediate. However, in 1863 (Trans. Ent. Soc. Lond. 8rd series, vol. i. p. 492) I described a genus (Anisocentropus) in which the spur-formula was 2, 4, 3, in both sexes, and another (Nesopsyche) in 1866 (1. c. vol. v. p. 268), with the formula 3, 4, 3. And in JZeteroplectron we have a still more remarkable case, the formula being 2, 4,2 for the ¢, though regular (2, 4, 4) for the 2. The genus is probably allied to Anisocentropus, somewhat resembling its general form, but differing (besides in the spurs) by the shorter, stouter, and subserrate antenne, different pro- portions of the joints of the palpi, minor characters of neuration, and the long-fringed posterior tibie of the ¢. But it has little affinity with any cther described genus of Leptoceride. HETERKOPLECTRON CALIFORNICUM, nov. sp. H. fusco-nigrum. Caput thoraxque aureo-hirsuta. Antenne fusce. Pedes flavi; maris tibize posticee extus fimbriis aureo-griseis longis instructz. Alz anticex fuscse, aureo-pubescentes; venis venulisque fusco-nigris: posticae fusco-nigree (5 @). : Long. corp. 43-53 lin. (=11-123 mill.); exp. alar. 14-16 lin. (=29- 34 mill.). Hab. California (in Mus. auct.). Blackish fuscous. Head clothed with golden hairs, springing from tubercles on the crown; thus there is a rounded tubercle on each side close to the eyes, another in front between the basal joints of the antennz (these simulate ocelli), and an elongate one on each side on the posterior margin (when the hairs are removed, these tubercles are seen to be paler brown than the ground-colour) : antennz fuscous or black, somewhat yellowish in some ¢ individuals, with obsolete darker annulations; eyes dark coppery; palpi clothed with fuscous hairs. Prothoraz clothed with golden hairs; and there is a broad line of similarly coloured hairs down the middle of the mesothorax. Legs testaceous, darker in the 2 ; in the ¢ the outer side of the tibize is fringed with very long greyish-golden, silky hairs, and the tarsal joints have long spine-like tufts of greyish or fuscous hairs. Abdomen blackish ; appendices of the ¢ somewhat testaceous; app. sup. elongately triangular, pointed, arising from each side of the middle of the margin of the last dorsal segment, fringed with long golden hairs; app. inf. thick, cylindrical, curved upwards, fuscous at the base, then testaceous, and black at the tips, fringed with long golden hairs; from the interior of the cavity of the last segment 126 MR. R. M‘LACHLAN ON NEW FORMS, ETC., arises the long, triangular, yellow penis (or perhaps it is rather only the penis-cover), the apex of which is somewhat produced and notched, this member is concave beneath, and extends beyond the appendices. Anterior. wings varying from dark golden brown to blackish fuscous, uniformly of one tint without markings (the 9 always the darker), clothed with golden or fuscous pubescence (when the pubescence is removed, the membrane appears to be sprinkled with somewhat numerous, but indistinct, pale dots); fringes fuscous; veins brown, costa, subcosta, and radius darker, because thicker ; upper branch of the ramus thyrifer, in that portion of it that forms the upper boundary of the cellula thyridii, whitish, semitransparent. Pos- terior wings smoky blackish, the veins darker; fringes blackish. (Pl. III. fig. 10, details.) I have examined six males and two females, sent to me by Mr. Henry Edwards, of San Francisco. ’ Ganonema, MU‘Lachlan (Tr. Ent. Soc. Lond. ser. 3, vol. v. p. 253). In this genus should be placed Hydropsycha vicaria, Walker, Cat. Brit. Mus. Neurop, pt. 1, p. 114, from Venezuela, the type of which is a single unexpanded example with broken antenne. I have received a second individual from the same quarter, from which I have drawn up the following description. A second species is also from Venezuela. These do not differ sufficiently in structure from the Malayan G. pallicorne to necessitate the formation of a genus for their reception, notwithstanding the wide difference in locality. I still think that the suspicion ex- pressed by me (J, c. p. 255), that Asotocerus and Ganonema may be identical, is well-founded, especially as the neural differences in the fore wings are more apparent than real, inasmuch as the lower branch of the ramus discoidalis is really only simply fur- cate in G. pallicorne, the supposed additional sector belonging to the ramus thyrifer; hence there are the same number of sec- tors in both genera. The neuration of the hind wings of both the Venezuelan species is like that in Asotocerus, both being males ; thus it is very pfobable that the differences are sexual, as I sus- pected, In the form of the wings the South-American specics agree with Ganonema*, * A very closely allicd genus is Calamocerus, Brauer, as would seem to have been since recognized by its describer (Verh. Zool. Bot. Gesell. Wien, 1868, p- 406). I cannot help thinking that the locality, “ Gibraltar,” given for C. marsupus, has arisen from an error in labelling, and that the insect is really exotic, OF EXTRA-EUROPEAN TRICHOPIERA. 127 GANONEMA vicarium. (Hydropsyche vicaria, Walk. j.c.) G. ferrugi- neum, mesothoracis lateribus nigricantibus. Antenne flavee, nigro- annulate, articulis singulatim spina brevi intus ad apicem instructis. Pedes flavi, tibiis tarsisque intermediis extus obscurioribus. Ale anticz griseo-fulvze, nigro-pubescentes et fimbriate, maculis aureis plurimis indistinete irrorate ; margine apicali obliquo, paullo rotundato ; cellula apicali 1" anastomosim attingente: postice grisco-subhyaline, griseo- fimbriate (¢). Long. corp. 4 lin. (=9 mill.) ; long. antenn. 15 lin.(=32 mill.); exp. alar. 12-13 lin. (=25-27 mill.). Hab. Venezuela (Dyson; Goring; in Mus. Brit. et auct.). Ferruginous (or reddish-testaceous). Head clothed with scattered yellowish hairs upon the vertex, and with a few distant black hairs at the margins of the orbits: antenne yellow ; each joint, after the basal, conspicuously black in its apical half, the apex of each bearing a short black spine internally (in the last third the annulations be- come indistinct and brownish, and finally disappear): eyes coppery : maxillary palpi yellow, clothed with long black hairs, intermingled with some yellowish ones: labial palpi with yellow hairs. Meso- notum bearing a broad black stripe along each side. Legs yellow, with yellow pubescence; but the intermediate tibia and tarsi are rendered blackish externally owing to the presence of sparse blackish pubescence. Abdomen yellowish, the apex obscure: app. sup. long, flattened, somewhat lanceolate, yellowish, the points ap- proximating ; app. inf. long, thinner than the app. sup., directed up- ward, with a tuft of blackish hairs at the tips; between the app. inf. is seen the short, thick, yellow penis (or cover?). (Pl. LI. fig. 11.) Anterior wings obliquely rounded at the apical margin ; grey or greyish- fulvous, densely clothed with blackish pubescence, especially at the apical portion, and with numerous but ill-defined spots formed by golden-yellow pubescence ; apical fringe short, blackish, golden at the extreme base ; first apical cellule reaching the anastomosis, as long as the third, impinging only slightly upon the discoidal cell. Pos- terior wings pale grey, subhyaline, iridescent ; fringes grey; veins brownish. The pubescence of the anterior wings is only lightly attached ; the golden markings are more conspicuous when the wings are closed. GANONEMA MOLLICULUM, n. sp. G. flavo-testaceum. Antenne flavee, brunneo-annulatx. Pedes flavi. Ala anticx subhyalinie, aureo-pubescentes, brunnescenti-fimbriate ; margine apicali oblique truncato; cellula apicali 1° petiolata: postice albido-subhyalinz, albido-fimbriatz (¢ ). 128 MR. R. MSLACHLAN ON NEW FORMS, ETC., Long. corp. 3} lin. (=7 mill.); long. antenn. 16 lin. (=34 mill.); exp. alar. 12 lin. (=25 mill.). Hab. Venezuela (Goring), in Mus. auct. Testaceous yellow. Head and palpi clothed with yellowish hairs; antennz pale yellow, the apical half of each joint pale brownish, not toothed internally. Legs yellow, with yellow pubescence.