a ee See enn a este Set 2 —— ANS I) s i “J ial en a one rae! 2 thn a che A eneas THE MAMMALIAN FAUNA OF THE EDINBURGH DISTRICT Y Only 200 copies printed. THE MAMMALIAN FAUNA OF THE EDINBURGH DISTRICT, WITH RECORDS OF OCCURRENCES OF THE RARER SPECIES THROUGHOUT THE SOUTH-EAST OF SCOTLAND GENERALLY BY WILLIAM EVANS, F, B.S. E. SECRETARY TO THE ROYAL PHYSICAL SOCIETY ae - ton Inst EDINBURGH: M‘FARLANE & ERSKINE 1892 ALLHSON Tap SIMTHSON A } y, FESIC® PREFACE THE present volume is substantially a reprint of a paper forming part of the “Proceedings” of the Royal Physical Society for the Session 1890-91 (vol. xi., pp. 85-171). The paper was read at the meeting of the Society on 15th April 1891; and printed in December, a few additional records and observations being first added to bring it down to date. The reproduction of the memoir in its present form is largely due to the representations of a number of friends, who have been good enough to say that it supplies a long felt want, and to suggest that, if issued as a separate volume, its usefulness would be materially increased. Having per- sonally experienced the want of a treatise on the Mammalia of the country surrounding the Scottish metropolis, I have the less hesitation in giving effect to their suggestion. For the permission to reprint the paper, I take this opportunity of expressing my sincere thanks to the Council of the Royal Physical Society. My best thanks are also due to many friends and correspondents—Mr Eagle Clarke in particular—for valuable assistance rendered in a variety of ways. The information I owe to each has been acknow- ledged, as far as possible, in the text. Walk: EDINBURGH, April 1892. eer SUE ey : ; «< Hah JANOVI AN) ‘Dy, 14 Pie ee - 2 THE MAMMALIAN FAUNA OF THE EDINBURGH DISTRICT INTRODUCTORY REMARKS THE late Mr E. R. Alston, in the closing sentences of his Catalogue of the Mammalia of Scotland,! pointed out to Scottish naturalists that the distribution of the mammalian- life of the country was much in need of revision, and cited the Shrews, Mice, and Voles as especially deserving of attention. Impressed with the truth of his remarks, I have been endeavouring during the last few years to work out in some degree of detail the distribution in our own neigh- bourhood of the above-mentioned groups, and also of the Bats, which had been equally neglected. My original intention was to communicate to the Royal Physical Society the results of my observations on these groups alone. Being, however, also in possession of a mass of data bearing on the past and present distribution of the other recent animals of the class Mammalia occurring in the district, I ultimately decided to combine the two sets of notes, and lay them before the Society in the present form. The “Edinburgh District,’ as here understood, embraces the valley of the Forth, and such parts of the adjoining areas ? Published in 1880 by the Glasgow Natural History Society as part of its ‘Fauna of Scotland, with special reference to Clydesdale and the western district,” 8 INTRODUCTORY —Tay and Tweed—as lie within a moderate distance, say twenty to thirty miles, of the city, the whole being capable - of investigation in the course of a series of easy excursions, seldom requiring more than a day for their accomplishment. It is, in fact, practically the same area as that adopted by Balfour and Sadler in their “Flora of Edinburgh,” and shown in the map which accompanies both editions of that work,—a section of country presenting a combination of physical features peculiarly rich and varied. The counties embraced are—on the south, East Lothian, Midlothian, West Lothian, and Peebles, with parts of the adjoining counties of Berwick, Roxburgh, Selkirk, Lanark, and Stirling; and on the north, Fife, Kinross, Clackmannan, and a portion of Perth. Through the centre—from west to east —winds the Forth with its estuary and broad firth, into which innumerable tributary streams from secondary valleys empty their waters. Fresh-water lochs and ponds also abound. The upper part of the main valley, penetrating as it does the south-western section of the Perthshire Grampians, is thoroughly wild and alpine in character. From the rugged mountains of this north-west corner, a series of sub-alpine ranges—the Campsie Fells, the Pent- lands, the Moorfoots, and the Lammermoors—with their connecting moorlands, constitute the watershed on the south ; while the almost alpine Ochils, the Cleish Hills and the Lomonds, mark it on the north. Between this rampart of hills and the shores of the Forth, every variety of lowland country is to be found—fertile lands and barren commons, green meadows and furze-clad hills, breezy heights and secluded dells, with woods and plantations of deciduous trees and pines on every side. The part of the Tweed area of which we take cognisance lies largely in the pastoral county of Peebles, and consists for the most part of grassy INTRODUCTORY i) and heather-clad hills, intersected by a multitude of glens dear to the angler. The section of the Tay area falling within our limits is mainly lowland towards the east, and highland in the west. It will thus be seen that the district, whether we contemplate it as in the natural garb of former times, or as now changed in outward aspect by the hand of man, is well fitted to be the home of a mammalian fauna rich both in species and in individuals. In the case of the rarer and more interesting species, occurrences throughout the south-east of Scotland generally will be alluded to. From the earliest times man has ever exercised a modify- ing influence on mammalian-faunas, adversely affecting some species either by direct persecution or by rendering the country unsuitable to their habits, and directly or indirectly fostering the increase of others. He has, moreover, long been in the habit of importing certain species from one country or district to another, so that it is not always easy to separate the indigenous from the introduced. The more populous a district becomes, and the more its agricultural industries are developed, the greater will be the changes on its fauna. Add to these factors the existence for many centuries of a large and influential class of landowners holding strong _ views regarding the preservation of game, and it will readily be understood that the district around Edinburgh was prob- ably the first in Scotland to witness a radical change in the character of its mammalian-life within historic times. The larger predaceous animals, such as the Wolf and the Bear, which carried destruction among the flocks, and even threatened the life of the herdsman himself, would be among the first to succumb. Many species would be hunted for their skins or their flesh; others mainly for sport. The smaller Carnivora would receive further attention on account of their visits to the poultry-yard, and Hares and Rabbits b 10 INTRODUCTORY because of injury to the crops. Then came the game laws—another interference with the balance of nature— accelerating the destruction of the predatory animals, and facilitating the increase of the rodents. The inordinate increase of the Rabbit led in its turn to a universal system of trapping to keep it in check, and from that day the fate of most of the remaining terrestrial Carnivora was sealed. Boece and Sibbald have put us in possession of much valu- able information regarding the fauna of Scotland in the sixteenth and seventeenth centuries, but as a rule their statements are too general to be of direct interest in the present inquiry. But from the “Old Statistical Account” of the parishes we get some excellent glimpses into the state of our fauna a century ago, Even then the predatory animals had been in great measure banished to the outlying parishes, where, however, they were still not uncommon, as the following extracts clearly testify :— CAMPSIE (STIRLINGSHIRE).—After mentioning the Badger and the Fox and their varieties, the writer of the article con- tinues: “There are likewise (on the Campsie Fells) weasels, otters, polecats, hedgehogs, wild cats; and, of late, several martins have been seen among the rocks. . . . It may be observed, that beasts of prey are every day becoming scarcer. Till within these two years, we had a regular bred huntsman who hunted this district ; his salary was paid by the tenants, at so much per plough, which huntsman and dogs were kept and fed by each tenant in his turn” (vol. xv., p. 323). CALLANDER (PERTH).—“ Red deer come here for food and shelter in severe winters. oes breed in our woods. Hares, rabbits, foxes, wild cats, badgers, otters, moles, polecats, weasels, and black martins, are also to be found here” (vol. x1:, p. 598). DouneE (Soura-West Perrn).—, Yrattus, ;, sylvaticus, : Muscardinus avellanarius, Mustela erminea, 5 martes, », putorius, », Vulgaris, Narwhal, Nomenclature followed, Orea gladiator, Otter, Ox, Great ene Horne’: INDEX PAGE PAGE 25 | Phoca greelandica, 13, 56 103 », vitulina, ; : 57 103 Phoceena communis, 106 103 | Physeter macrocephalus, . 12 Pilot Whale, 109 13, 25 Pipistrelle, 19 Plecotus auritus, yf 5 Polecat, 49 mi Porpoise, 106 saa Publications eonenltedt 117 ae a Rabbit, 84 80 Rat, Black, 74 8] », Brown or Norway, Ue 99 Razorback, 99 40 Reindeer, 5 Cl Rodentia, 13, 60 ane Rorqual, Common, 99 es 5 Lesser, 102 i a Rudolphi’s, 101 a: Pa Sibbald’s, . 98 42 105 | Sciurus vulgaris, 60 3] Seal, Common, : 5 Bil 12 », Greenland or ee 13, 56 78 », Grey, 55 76 », Hooded or Bladder nose, 59 77 Shrew, Common, 26 75 », Lesser, 20, 2 73 », Water, od z8 Sorex minutus, 27, (2 76 » pygmeus, 27, 72 74 | Se temitenre 29, 30 . 97 | » vulgaris, 26 13, 63 Squirrel, . . = . 60 53 Statistical Account (Old), Ex- Aq tracts from, . 10, 11 49 | Stoat, : 53 52 Talpa europea, 31 12 | Tay Whale, ; 96 15 | Trap for Micro-Mammals, 16 Tursiops tursio, . : 114 107 40 | Ungulata, 13, 87 12 Urus, 12 INDEX PAGE Vespertilio daubentoni, . >» 20) Whale; Blue; “5 mystacinus, . 13, 23 », Caaing, re nattereri, F Ss 22 »» Hump-backed, Vesperugo pipistrellus, . 5 OY oe edlotie Vole, Bank, . : : 6 fe »» Sowerby’s, pee lel dees 9. : : 7 65 ye perm; ae Water. : ; 5 ae! 5, White, $5 », black variety, . 65 | White Cattle, . Z Wild Cat, Weasel, . : . : 52 | Wolf, Whale, Beaked or Bottle-nosed, 103 Printed by M‘FARLANE & EnskineE, Edinburgh. nga hae ee vinuia may F ea The main n fauna of the Edinburgh dis