_Phytologia
vol, 2}
1966

MlesOUR!] BOTANICAL
GARDEN LIBRARY

PHYTOLOGIA

Designed to expedite botanical publication

Vol. 13 : February, 1966 No. 1
CONTENTS
MOLDENKE, : N., Materials toward a monograph of the genus
Lippia I Le eek Bee gh ae eee ee y ee ke 1
MOLDENKE A. t. Buak seu. 2 ee a ee 35
ELTEN; L. 1., Notes on Brazian Cyperdcede + 022. i ee 36
DeFILIPPS,-K. A., Distribution of Juacus tm Hlinois’. ceo. ee es 44

Published by Harold N. Moldenke and Alma L. Moldenke

—— Borxntce 303 Parkside Road
Plainfield, New. Jersey, ‘U.S.A.

WAR 28 1966 this nu

‘gence EO $6 at close of volume

aber, $1: per volume, $5.75, in advance,

MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. IX
Harold N. Moldenke

LIPPIA TAYACAJANA Moldenke
Additional bibliography: Moldenke, in ae 12: 506. 1966.
br:

Shrub, about 1 m. tall; branches slend tetragonal, ribbed,
densely short~pubescent_ bee young, abevesdnt in age and then’
with peeling shreddy bark, brown, somewhat twiggy below

j twigs
very short, leafy; nodes gis: anmlate; principal internodes much
abbreviated, 1—-3 cm. long; leaves decussate-oppos ite; petioles
slender, i2 mm. long and densely short—pubescent or obsolete;
leaf-blades firmly chartaceous, uniformly gray-green on both sur-
faces, oblanceolate-elliptic, 0.8-~1.9 cm. long, 3-9 mm. wide,
und

3
pressed~pubescent with antrorse canescent or y
h » about 5 mm. long and wide; bractlets cc ariparriy a '
bout mm. long and 1.5 mm. wide, sharply acu e at the apex,
densely _appregsed-strigose-pubescentj ae iypocrateriform,
white, PE airs its limb about 2 m,
he pecies was collected by Poereite Weberbauer

(no. 6510) ‘i the valley of the Mantaro, northeast of Pampas, in
the | province of Tayacaja, Huancavelica, Peru, at an altitude of
1800-—-1900 meters, in March, 1913, and is deposited in the United
States National Herbarium at Washington. The species is obviously

has been found growing

The Asher, Satie: & Cia. 17987, cited below, is placed here ten-
bse whi because. of its fragmentary nature
rbarium specimens, including the type, and 2 mounted
photographs page been examine
Citations: PERU: Huancavelica: a 6510 (N-~isotype, N—

— of oa. W--1,9540—type, Z—-photo of type). La Libertad:
Lépez Miranda & Sagdstegui 3358 (Z). Department undetermined:
Asher, Kates, & Cia. 17987 (Kr).

LIPPIA TEGULIFERA Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8:

309——310. 190). parvifolia Bri i
Synonyny : Lippia era var = Qe, ons
& Jard. Bot. Genev. 7-83 310. 190). ann eT var. tegu-

lifera [Troncoso], Darwiniana 12: 38078 & an ar wet tegu-
lifera Big. ex Voldenke, Résumé eer 10: 6, in syn. Lippia

2 PRITOLOGIS Vol. 13, no. 1

temulifera Briq. ex ee Résumé Suppl. 10: 6, in syn. 196).
Bibliography: , Ann. Conserv, & Jard. Bot. Genév. 7-8:
309——310, 190hs ‘aria, “in Chod. & Hassler, Pl. Hassler. 2 (11):
492. 190; Prain, Ind. Kew. Suppl. 3: 10. 1908; Moldenke, Lilloa
5: 429. 1940; Moldenke, Known Geogr. Distrib. Verbenae fed. 1)
3 :

se
‘ta1108 Uy: h6 & 7.
Alph. List Cit. 2: 3h2, 376, 4,0, 457, 400, & cer. C318), 33 ae
& 863 (199), and 4: 1165, i69, "& 12 — 1949; Mo Known
Geogr. Distrib. Ver benac., [ed. 2], 99, 105, & oe «19194 ee,
P 117, 126, 317, *& 463. ibe, Troncoso o, Bol. Soc

Bot. 9: 184. 1961; Troncoso, Darwiniana 12: 258 & 280—282, ae 33
k—o. 1961; Moldenke, Résumé Suppl. 10: & 6 (196) and 12: 11.

ustrations: Troncoso, Darwiniana 12: 281, fig. 12 ko.

Tr shrub or subshrub, gpreoray m, tall, with
the base; aerial stems erect, woody or eg simple
or seni internodes tetragonal, sade or very short, pubescent
throughout

ppro abundant, closely appressed to the branches in imbri-
cate fashion, sessile; leaf-blades elliptic tp elliptic-lanceolate,
8—-20 = - Lone, 5--8 mm. wide, shortly acuminate or acute
at the apex, more or less ney along the margins, more convex
below the siddle, regularly and abundantly crenate or crenulate,
cordate at the base, stiff, coriageous, rigid, gacda capi Phone,
scabrid above, sparsely strigose~pubescent with coarse hairs 0
both surfaces, » also appressed-pubescent beneath, ce eg
— nerved, lineate-rugose above; secondaries approximate » paral-
, impressed above; heads globose, medium-sized, opposite in
— leaf-axils, subsessile or very short-pedunculate, 5-8 mm
i and wide, cinereous-pubescent or incanous ; peduncles tetra-

ng,
nomal for the genus, included; fruit small, smooth, 1.5 mm. long,
virion densely villous like the fruiting-calyx, the "coced split-
ng apart on maturity.
7 he 190k) proposed this —— with ee vari-
eties, pre which I regard the first, var. olia, as the typi-

ype, then, of the species is that of a typical

he type
form, collected by Benedict Balansa (no. 1031, in part) on the

v

“-

1966 Moldenke, Monograph of Lippia 3

campos Caaguazu, Paraguay, on November 7, 187), and deposited
in the se Appenies Herbarium at pe pitoame eS et Jardin Botan-
iques at Geneva. Briquet describes this typical form as follows:
"Caules aerii 30-60 pilis brevibus adpressis et aliis longioribus
ascendenti-patulis nunc aliq. crispulis ferrugineis obtecti. Fo-
lia creberrima, parva, elliptico-lanceolata, utrinque pilis stri-

e ea subtu

conpit supra aspera, ferrugineo-viridia, guetta 0,8—2 x

0,3-—-0,8 cm. Capitula rest sect. long. 5—8 x 5—8 mm, subses-
silia vel brevissime pedunculata, pedunculi a ultra 5 m. al-
tis quam folia brevioribus." He. says of the species: "Cette re~
marquable espéce se place dans le groupe Axilliflorae de la sec-
tion Za a: elle ne peut 8tre étroitement rapprochée d' aucune
des especes eres son port, qui est celui des Oftia, lui
donne une place 4 part." He says that Balansa 1031 is ” actually
a mixture of the typical L. tegulifera, var. var. ovata, and var. pe~-
dunculata.

The second Briquet reference given ol the ee above
is cited at the Instituto Miguel Lillo "Plantae Hassl stoped

ae ab Ga have found this plant growing on wet or humid savan-
campos, high campos, and the banks of rivers, at 150 meters

aliscete, lowering from ae to May and in July, fruiting in

vember

ember, Rojas found it on a "campo pendiente Ze
loma"; See describes it as "rare" on campos, while
found it to be " a etd h uncultivated campos."

2 rig one

It should be noted that the University of ceggnaey specimen of
Ekman 201) is pee obey et mg the ps 8 form of the species, with
elliptic leaves, while the Britton Herbarium specimen of. the same
number is definitely var. ovata, with ovate leaves. Troncoso
Sot ge says: — a de la que su autor distingue

ariedades. La var a ha sido considerada por Moldenke

como la var. tipica. ry “oad el ejemplar Balansa 1031 pp
como tipo. El valor de las restantes: var. ovata, var . peduncula-
ta y var. grisea requeriré un minucioso estudio en base a observa-
clones de abundante material que permita apreciar la constancia de
sus caracteres paar igh pone a Por el momento me remito a citarlas

lectotype, Bm——-lectotype, Cb--lectotype, K-- P—lecto-
type, Ebr vaca tapelé Hassler 1079 (Cb, K, P), oC (Bm, Cb, K, P),
6108 (Bm, Cb, K, P, Si), 6106a (Cb); Jorgensen 37 3780 (Si); Montes
bo (94) 5:5 aes 3422 a 5151 ish (St), 6087 (Si). ARGENTINA:

rrientes: Burkart seat ore (ot a 3963 (M1). Misiones: Bur-
eats 14751 (Si), 15307 (Si); se wee B87 (i Ci). or
these she says that Ha aaa * 6x68 , K, P) and T. Rojas 6087 (Si)

are definitely staminate specimens; Daddies 6108 (Si), ensen
3780 (Si), and T. Rojas 3 sie (Si) are definitely pi istiiiste speei-

h PHYTOLOGIA Vol. 13, no. 1

mens; while Burkart 7861 (Si) is both staminate and pistillate.
Hassler 6108 is definitely var. ovata, 1079 was called var.
culata by Br. Briquet, and 6108a was called var. ovata by Briquet.

Material has been misidentified and distributed in eopimba as
var. ovata Briq. and as L. pumila Cham. In all, 36 herbari
specimens, including type material of all the Bese poabenicin and
6 mounted ‘photographs neve been examined ew Sie

876815, V), 9142 (Ca—929822, Cb, N, N—-photo, V, Z—photo); JUr-
gensen 3780 ee Osten 2223) ] (Du—197830, 5 Ue, W—~183632),
a [Herb. Mus. Argent, ee Nat. 28/2072 1 (w ); Kuntze s.n.

piateinae sa (N); T. Rojas as 3422 (Herb. Osten 18196] 6) (Ug),
. (Herb. Osten 15647] (Ug). ARGENTINA: Corrientes: Ibarrola
36) (N), 4012 (N, N, S); Ruiz Huidobro 2082 (S), 2169 (N, S, Vi).
Formosa: Venturi a (W-~1591)16). Misiones: Burkart W751. (N,
W—2056 385) ; Ekman 2011, in part (Mi); Montes 1715 (N); Quir Quiroga
49 [Herb. Osten 7682) (Ug), 432 [Herb. Osten 7682] (Ug), 7682
(Go), s sen. (Herb. Osten 7682) (S (S); D. Rodriguez 697 [Herb. Inst.
Miguel Lillo 32236] (Ca—3372, N, W—1802580, W—1858291).

LIPPIA TEGULIFERA var. GRISEA Briq., Ann. Conserv. & Jard. Bot.
Gensv, 7-8: 310——311. 190).
Bibliography: Briq., Ann, Conserv. & Jard. Bot. Genév. 7-8:
310—311. 190); Moldenke, Known Geogr. Distrib. Verbenac., [ed
1), 41 & 96 (1942) and [ed. 2], 99 & 191. 19h9;3 Noldenke, *nésumé
Lis niana de

griseous with short appressed hairs and longer ascendi ones;

leaf~blades elliptic-lanceolate, medium in size, radiior: dense,

very acute at the apex, roughened above and pubescent with small
ttered hairs, densely and rather softly pubescent beneath,

griseous, 2—3 cm, 1 » 1 cm. wide; heads griseous-canescent

to 1 cm. lo ong and 8 mm. wide, short—pedunc ulate; peduncles shorter

beac baad subtending leaves, 1--1.5 cm. long; bractlets ovate-

vein tote of this variety was probably also collected by Bene-
dict et Me hough Briquet dates no collectar or number,
ures on plain of Pirayu-bi, Par. on sbesirg 3: 1978
and is probably deposited in the Delessert Herbarium at
servatoire et Jardin Botaniques at G Geneva. Nothing is iret og
me of this taxon except what is stated in the literature.

LIPPIA he ee 9 ioc toe Briq., Ann. Conserv. & Jard. Bot.

fuerte: Briq., Ann. Conserv. & Jard. Bot. Gendv. 7-8:
a6 se Moldenke, Lilloa 5: },29 (1940) and 8: 426. 192; Mol-

1966 Moldenke, Mpnograph of Lippia S

denke, Known Geogr, Distrib. Verbenac., [ed. 1], h1 & 96. 19h2;
Moldenke, Alph. List Cit. 1: 26, 263, 26h, & 266. 19,6; Moldenke,
ya Age eal 2: 386. 1947; Molde enke, Lilloa 1): 6 : wee 1948; Mol-

nown Geogr. Distrib. Verbenac., [ed. 2], 105, & 191.
ohse erie Alph, List Cit. 3: 690 (199) and St: 1257. ge
pres résumé 117, 126, & 463. 1959; Troncoso,

This varie differs from the typical form of the ee in
having its leaves less maar l -riggead crowded, often somewhat larger,
to 2 cm. long and 1.5 cm. wide, ovate, rather acutish at the apex,
the margins below the middle el Phi hen convex, the base con-
spicuously corda

The type of this variety was collected by Benedict Balansa (no.
1031, in part) on the campos at Caaguazu, Paraguay, on November 7, FR

din ques a ng Briquet
(1904) Balansa 1031 is a mixture of typical L. tegulifera Brigq.,
var. pedunculata, a and var. ovata. Similarly, - the specimen of
Hassler 6108 Babee by Macbride in the herbarium of the
Botanisches Museum at n as his type ph nage “aprep’ 17551
was originally Siteled’ 6108", but Briquet avers t this nun-
ber consists of a mixture, t t00, and he cites the var. ovata por-
tion as "6108a", Troncoso (1961) regards the true 6108 ma’ terial
as var. pedunculata Briq.

The variety ovata is described as having yellow flowers, like
the typical form, grows 20—-80 cm. tall, on campos and rocky cam-
pos, flowering in September and November . The Britton Herbarium
specimen of Ekman oo is definitely var. ovata, with ovate sears,

e the University o: f Michigan specimen of ihe same number i
definitely the typical form of the species, with elliptic poe a

Material has been misidentified and distributed in herbaria as
L. tegulifera Briq. and as L. pumila Cham. In ‘ erbari
specimens, including type material, and 11 mounted photographs
have been examined by me.

Citations: PARAGUAY: Balansa 1031, . monde saeco tae ~
photo of isotype, N--iso N—photo
isotype, Z—photo of sai Hassler 1958 rea é108 ( ahem °
Mi, N, N, S), 6108a (Macbride photos 17551] (It—photo, Kr—photo,

hoto, N--photo, W-—photo), 9906 (N, N--photo, V, Z—photo);
Hauthal 23 (N). ARGENTINA: Misiones: Elman 201, (N, S); A. G.
Schulz - ‘7110 (Sz).

LIPPIA TEG » PEDUNCULATA Briq., Ann. Conserv. & Jard.
2th i ag 7-8: “310. 190).
ane graphy: Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8:
0. 190h3 i Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], Pigg
& ry (1942) and ieee 21, 99& “gt Tos Moldenke, Résumé 11
463. 1959; Trone , Darwiniana 280--282, fig. 12 a--j. me
oth apa peupeak. an 12: 281, fig. 12 a—j. 1961.

6 Toes VEU ol se Vol. 13, no. 1

This variety differs from the typical form of the species in

lanceolate, very acute at the apex, the crenate-dentate teeth

more acute, more robust, 2--2.5 cm. long, 6--8 mm. wide, the in-

dumentum of the stem and leaves as in var. ovata; heads longer~

pedunculate, the peduncles equaling the leaves or sometimes sur-
ets

The +
1031b) in clearings in the forests, Cordillera de Peribebui, Para-
guay, on January 11, 1877, and is deposited in the Delessert Her-
barium at the Conservatoire et Jardin Botaniques at Geneva. Ba-
lansa 1031 is, according to Briquet (190), a mixture of typical
L. era, var. ovata, and var. unculata. This portion
was apparently collected at a different locality and on a dif-
ferent date than the other two portions previously discussed. It
is said to have had a heavy woody rootstock. Troncoso (1961)
states that Briquet regarded Hassler 1079 as this variety; she
regards Hassler 6108 as this variety, too, but I cite this number
pao ha ovata. Nothing further is known to me of var. pedun-

LIPPIA TEPICANA Moldenke, Phytologia 1: )5),—-55. 190.
Synonymy: Lippia mexicana Moldenke, Alph. List Invalid Names
Suppl. 1: 1), in syn. 197. Lantana canescens Benth. ex Molden-

ke, Résumé 30h, in syn. 1959 [not L. canescens H.B.K., 1817].
Bibliography: Moldenke, Phytologia 1: l5y--h55 (1940) and 1:
50h. 1941; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 17
& 96. 1942; Moldenke, Alph. List Cit. 1: 233. 1946; E. J. Salisb.,
oo “et > eet wag Moldenke, Alph. List pry om :
ods Ik. 3 Moldenke, Alph. List Cit. 2: 47h, 477
& 595 (1948) and 3: 788. 19h9; Moldenke, Known Geogr. Distrib. —
+, [ed. 2), 31 & 191. 1949; Moldenke, Résumé 37, 30h, 31h,
& 163. 1959; Langman, Select. Guide Lit. Flow. Pl. Mex. S15 &
1010. 1964; Moldenke, Phytologia 12: 108, lll, 112, & 363. 1965.
rowned shrub or tree, 1.2-—-1.8 m, tall; branches and
branchlets rather slender, acutely or obtusely tetr, onal, so

1966 Moldenke, Monograph of Lippia 7

der, mostly impressed above (or appearing as though raised by
being more densely pilose), rounded-prominent beneath; secondaries
r side

sharply prominent beneath; inflorescence axillary » paniculate,
one panicle and sometimes also 1 or 2 sidetahenies peduncles is-
suing from each axil or sometimes only several single-headed pe-
duncles; the simple inflorescences _ cm. long, the panicles to
9 cm. long; peduncles and infl e-branches very slender, 1—
4.5 cm. long, densely puberulent, hae obs scent, or strigose
with cinereous hairs; panicles with 1-- --3 whorls of ca: capitate bran-
ches; heads hemispheric during anthesis, oblong in fruit, densely
many~-flowered, 3--1 mm. long, 5--10 m. wide; bractlets large,
ranks istent, dry and scari-

1969) at Tepic, Nayarit, Mexico, between January 5 and February
6, 1892, and is deposited in the Britton Herbarium at the New
York Botanical Garden. Lantana canescens Benth. is based on Ar-
séne 32),3 from Michoacén, Mexico, deposited in the United States
National Herbarium at Washington. Humboldt, Bonpland, and Kunth's
species of the same name is a valid species "in Lantana
Standley identified the type of Lippia tepicana as | as Lantana in-

volucrata L., noting that "the Mexican form differs somewhat from
the West Indian, but not consistently". To this Britton has ap-
pended his comment "But this is ae the West Indian plant!"

Li tepicana has been found on grassy slopes with scattered
shrubbery on hills, at altitudes of 1325 to ~~ meters, flower-
ing in Jamary, February, May, and November, fruiting in Febru-

May. Gentry & Gilly found ey "in highly

el eri
wisidentitied ant dtsteiveta in herbaria under the names L. cal-
licarpaefolia H.B.K., L. pringlei Briq., L. umbellata Cav., Lanta-
na involucrata L., and even en Melampodium sp. in the Carduaceae!
In all, 19 herbarium specimens, es the types of all the
names involved, have been examined
Citations: MEXICO: Jalisco: H. S. Geukty 5353 vt N). Micho-
acén: Arséne 323 (W--1000868) . Nayarit: DeLeon 1618 (Mi), 1629
(Mi); Gentry & Gilly 108 (Rf); M. E. Jones Spat g ae Se
Po--15392h), 23116 (Gg—172647, N, N, Po—191958), 23417 (pace:
191959); Edw. - Palmer 1969 (ca—770321—i sotype, Cp—isot, type, Fs—
isotype, Mi—isotype, N—type, S—isotype).
LIPPIA THYMOIDES Mart. & Schau. ex Schau. in A. IC., Prodr, li:

Lippia ocymoides Mart. & Schau. ex Briq. in Engl. &

8 Pat t.0L 0G & Vol. 13, now 1

Prantl, Pflanzenfam. ) (3a): 152, sine es

sitiiograpty? Schau. in A. DC., Profr cath 1847; 3
chau, in Mart ae Bras. 9: 20. 1851; sada: in Hook. f.
pert Ind. K 2: 96. 18943; Briq. in Engl. & Pr antl, Nat.
Pflanze (3a): 152. 1895; Solered., ek Herb. Boiss. sér.
pe 628. 1898; Moldenke, Known Geogr. Distrib. Verb , fed.
1], 38 & bag oldenke, Alph. List Cit, 2: 429 (198) and 3:
691 & 711. 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed.
2], Se 191 5 1b19; Moldenke, Résumé 9) & ney ree Moldenke,
Résuné s Suppl. 8: 5. 18643; Moldenke, Phytolog : 2). 1965.

shrub, 1--1.3 m. tall; branches c ceoieted: ah reading, virgate,

Hstetm-ateatres: with abbreviated axillary branchlets; leaves

decussate-opposi mm. long, 2 mm. wid
cuneate, rather acute or obtuse at the apex, entire and revolute
along th » i-veined, scabrous above, gl ru-

sterile branchlets; heads solitary in the upper leaf-axils, very
short~pedunculate, subglobose or oval, com ompact, about 8 mn » long;
bractlets subherbaceous, imbricate, ovate-lanceolate, acute at
the apex, 3-veined, glandular-pulverulent on the back, ciliate

eum at Munich and photographed there by Macbride as his type pho-
tograph number 20335. Schauer classified the species in his sec-
tion Za apania, subsection Axilliflorae. It is hus
from the original collections. in all, mounted phototypes have
been tna oe me.

Citations: BRAZIL: Bahia: Martius 1955 [Herb. Monac. 137; Mac-
bride Sinton 20335] (It--photo of type, Kr—photo of type, N=
photo of type, W—photo of se

LIPPIA TORRESII Standl., Field Mus. Nat. Hist. Publ. Bot. 18:
1010—1011. 1938.
nyny: Lippia worress ae n herb,

Bibliography: P - Standl., a sto; Nat. Hist. oo Bot.
18: 1010—~1011. 1938; “Moldenke? Ga. List Common Names 19395
Moldenke, Suppl. List sig ommon Names 5. 1940; Moldenke in sep &
Schery, ae Mo. Bot, om 335-336, too Moldenke

&

rib. V 1
Phytologia 2: 107. 19,5; Moldenke, Alph. List Cit. 1: 7. 196; E.
J. Salisb., Ind, Kew. Suppl. 10: 13h. 1947; lioldenke, Phytologia

denke, ’
999-1001. 1949; meee Known Geogr. Distrib. Verbenac.,

2], 39, Wl, & Mp ba 949; Dayton, Phytologia h: 259. 1953; Bris-
icky, Stern, & Chambers, Trop. Woods 109: 78. 1958; Holdenke, Ré-

1966 Moldenke, Monograph of Lippia 9

sumé 7, 48, & 463. 19595 Moldenke, Résumé Suppl. 8: 1. 196h;
Moldenke, Phytolo gia 50k. 1966.

Large forest tree, pas - tall; crown broad, flat or rounded;
trunk 15—90 cm. in diameter at tevene at height; bark neutral gray,
brownish, or light-brown to gray-brow, neutral-brown, or dark-

i se

adrate or square, satiealy rs cnet with

short sordid hairs; leaves decussate-opposite, » With a
mint-like odor, rather large, petiolate; peti nleu al slendér, 1.5—
2.5 cm. long, densely hispidulous; leaf-blades thick and fleshy

when fresh, membranous when dry, barely or slightly silttenal or
"a little stiffened but wine bright~green or dark emerald-green

above, ligh r pale-green to gray-green beneath, not shi

po ie 5c ‘ ficistie 1221 cm. long, 6—9 cm. wide, acute or
subacuminate at the apex, sharply crenulate along the margins,

rather abruptly cuneate-attenuate at the base, scabrous and ru-

e
touch above and more or less bullate with sharply reticulate
veinlets, much paler beneath and densely velutinous-pilose with
straight spreading but more or less entwined hairs throug
midrib slender, prominent beneath; secondaries about 10 per side,
arcuate~ascending with a rather broad angle; peduncles axillary,
fasciculate, very slender, about 1.5 cm. long, pu

attracting many insects; or flattened, globose
cm, wide when in fruit; bractlets light- or

violet-purple on t andular, slightly glutinous, membran-

ous and accrescent in the frui tag hen

the wind, venose, ciliolate on the margins, otherwise —
broadly rounded a at the apex; calyx almos st 2 mn, long, dens
villosulous; corolla hypocrateriform, yellow, pal aed ping or
light primuline yellow to primrose, greenish-yellow, or lemon-

and maturing cream-color, almost completely glabrous on the outer
surface, the tube slender, 2 mm. long, the lobes or 5, expanded

he type of this handsome species was collected by Aus
eat th (no. H.436) at tatesso de Zarcero, at an altitude of f 1960
Me, Alajuela, Costa Rica, in March, 1938, and is deposited in the

in honor of Rubén Torres Bases Costa Rican naturalist

ens
herbaria and that the flower-heads have the "appearance of roset-
tes resembling Lan Lantana or a tiny artichoke". The flowers are de-
scribed as "yellow" on P. H. Allen 1481 and Skutch 191, "greenish-
yellow" on P. H. Allen 1565, "pale-yellow” on Skutch 5167, "light
primuline-yellow" on A. F. . F. Smith H.187, white” on A A.F. .F. Smith 83

10 FRETTOLOGT<a Vol. 13, now 1

and Stork 183, "primrose to pale-yellow or cream-color" on A. F.
Smith 1569, "cream" on Lems 5049, and "lemon-chrome maturing to
cream—color" on A. F, Smith P.227h. Wood samples of Stork 4590

chigan he: rbaria
and a pollen sample taken from P, H. Allen 1565 is represented in
the slide collection of the Harvard Univ versity | herbariun,
The species has been found in forests and at the edge of for-
ests, in potreros and gti » in heavy loam or in rich loam and
mould, in semi-shade, on brushy slopes, and in the fog — influ-
= "at altitudes of 1130 to 2300 meters, flowering in
ebruary, A » June, October, and November, fruiting im 4 January,
—— and — A. Pr. Smith found it at the "edge of woodland,
sure in heavy clay-loam soil at upper limit of
an tt Pacific watershed.....fairly c
pec

+ in "upper montane forest with Quercus copeyensis, Q. costari-
censis, and abundant epiphytes."
~~ Vernacular names recorded for it are "caragra", "caragra negra!’
and "caragre", Material has been misidentified and alae eraetten
in herbaria as L. substrigosa Turez.

n the Harvard University herbarium there are deposited the
tutecictins specimens identified as L. torre torresii, but not as yet
checked by me: COSTA RICA: San José: E. L. little 6015 ee S.

- Agr. Forest Serv. 95036] (A). PANAMA: Chiriqui: M. E. Da-

Widson 793 (A), 1356 (A). Standley (1938) also cites the Fae
; ake oe eee en ae COSTA RICA: Alajuela: A. F. Smith H
187 (F), H F—type); Standley & Torres Rojas as 47937 (). | Fe
José: P. C. St Standley 13519 (F).

In os aay 26 herbarium specimens have been examined by

Citations: COSTA RICA: Alajuela: A. F. Smith 83 eis 183]
(Mi), 1569 (N), H.187 (N), P.227h (A, Ca—12727). Cartago: Lems
5049 (N); Tonduz 1876 (Br). Heredia: Skutch S67 (W—2351972).
San José: Skutch Tas 91 (A, N, S)s Stork 590 (La, Mi); Tonduz 1700
(Br, Br, N), 7843 (Br (Br), sen. {Herbd. Inst. P’ Physico-geogr. Nac. __
Costaric. 1781] (Br, N). PANAMA: Chiriguf: < H, Allen 1481 (G,
N), 1565 (G, N, N); Stern & Chambers 78 (Bs,

LIPPIA TRACHYPHYLLA hie Ann, Conserv. & Jard. Bot. Genév. 7-8:
: Lippia trac’ lla Mold. ex Stellfeld, Trib. Farmac.
19 (20): conte sphaim.
raphy: Briq. im. Conserv. & Jard. Bot. aye bd
naomi 190k; Prain, Ind, Kew. Suppl. 33 dol. 2

1966 Moldenke, Monograph of Lippia 1

and hk: 1293. 19493; Moldenke, Known Geogr. aga Verbenac., [ed.
2], 81, 97, 99, 105, & en ‘29195 Stellfeld, . Farmac.

(10): 169. 1951; Angely, Fl - Paran. 7: 12. 1957; Moldenke, Résumé
9h, 11h, 117, 126, 317, & 463. 19593 Angely, Fl. Paran. 16:
(1960) and 17: 6. 1961; Troncoso, Darwiniana 12: 258, 282, & 283,
fig. 13. 1961; Troncoso, Bol. Soc. Argent. Bot. 9: 18h. 13
Moldenke, ee 12: 2h, 90, & 290. 1965; Moldenke, Résumé

Ss x

Illustrations: Troncoso, Darwiniana 12: 283, fig. 13. 1961.
Dioecious decumbent herb; stems procumbent, issuing from a
hard thick rootstock, diffusely branched; branches dark-green
— ly strigose above with appressed antrorse hairs; middle "in
ternodes 2-—5 cm. long; leaves decussate-opposite, medium in size;
ad

petioles 3 ong; leaf-blades rather thick, rigid, dark-
en on both surfaces, ovate or 3—l,.5 cm. long,
2—2.5 cm. » acute or subobt + more convex along

de use at the apex,

below the middle and conspicuously crenate-serrate
(the teeth Ri a mm, long and 2—5 m. piers dly rounded:
cuneate at the base and prolonged into the petiole, pilose above
with small ox ae hairs that are thickened at their base, con-
spicuously scabrous, appressed-strigose beneath especially on the
venation, sparsely so on the lamina; venation prominently penni-
nerved, the midrib lineate above, seamen beneath, the ) or 5
secondaries anastomosing toward the margins, lineate above, pro-
minulous beneath; inflorescence axillary in the upper leaf
pedunculate; peduncles shorter than the subtending leaves, 1—1l 5
em. long, appressed strigose-pubescent, the hairs beneath the
head mixed with stipitate glands; heads subspheric, about the
size of a large pea (Pisum sativum), to 8 m. long and 10 m.
wide, grayish; lower bractlets ovate-acuminate, to 5 mm. long,
sometime es sing the head, densely strigose-canescent on the
e than the flowers

tire
iform, yellow or ge age ye exserted, surpassing the calyx-
mouth by 3 mm., the t e glabrescen ent below, pubescent above, the
limb pubescent. on the eae the lobes 4, short and ovate, the
anterior one longer, to 0.5 mn. long; stamens and pistil normal
ture

“4

(no. 4626) on the sandy borders of the Y'-acan-guazu near Valen-
zuela, Paraguay, in November, 1882, and is deposited in the De-
lessert Herbarium at the Conservatoire et Jardin Botaniques at
Geneva, where it was photographed by Macbride as his type photo-
graph rumber 24,672.

et (190) has this to say about this plant: "Espéce trés
caractérisée, voisine du L. Cham, dont elle différe par

qu
capitules, ses corolles d'un rose-violacé (jaunes dans le V. as-

12 rat CLOG TsS Vol. 13, now 1

errima) etc." In speaking of his L. modesta, he says "Cette es-

e est a peu prés intermédiaire entre le te asperrima Cham. et
le L. trachyphylla Briq. ci-apres décrit. Elle est caracterisée
par une hétérophyllie assez marquée, des feuilles a a peine eet et
non pas vraiment scabres et bien moins rugueuses que
deux espéces. L'indument des pédoncules ressemble a walet au L.
trachyphylla, mais ceux-ci sont plus allongés et plus tenus. Au-
tant qu'on gi en juger sur le sec, la corolle paraft &tre jaune
comme dans le L. asperrima." If it isr really true that L. trachy-

lia wept ta its relatives in having rese-violet flowers
tas stated on the type collection), then the Fries 616, cited be-
low, which is said to have had yellow flowers should b be examined
again, as it may not be L. trachyphylla after all,

iitophs trachyphylla has been found growing on campos and open

campos and along river borders, at altitudes of 2385 to 3000
meters, blooming in January, Mare h, October, and Rete ay Mater—
ial has been misidentified and distributed in herbaria as L. tur-
nerifolia Cham. as

In all, - herbarium pi al and cients mounted phototypes have

d by me
Citations: BOLIVIA: Cochabamba: Kuntze s.n. [Tunari, ner (N).
Province undetermined: Cuming 150 (V). PARAGUAY: alana
Le cae photos eke Gt—rhote of type, Kr——photo of =
to of type, W--photo of type). ARGENTINA: Salta: R. E. a a
616 (S); T. Meyer er 1610 (Ganeiay, eee

STIS Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér.
2, be 1161. 190).
nymy: Lippia a tristis var. normalis Briq. in Chod. & Hass-
ler, Bull. Herb. Boiss., sér. 2, kz 1261. 190. Lippia angusti-
folia Chod. ex Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér.
2, h: 1159, in syn. 190 (not L. angustifolia Cham., 1832, nor
pif & Spoons 1947, nor H.B.K., 19h7, nor es 190, nor
Schau., 1959]. Lippia turneraefolia Chod. ex Briq. in Chod. &
Hassler, Bull. Herb. Boiss., sér. ee hae in eke not
ie pcentells Cham., 1851], oon ceo eg i ee z
Briq. in Chod. & Se Bull. Herb. Boiss., Sg » hl: “use, in
syn. 190 [not L. turnerifolia Cham., 1832). Lippia a tristis var.
tristis Troncoso, Darwiniana 12: 28). 1961,
ography: "Brig. in Chod. & Hassler, Bull. Herb. Boiss., sér.
She a rg Briq. in Chod. & Hassler, Pl. Hassler. 2

Prain, Ind. Kew.
denke, Lilloa 5: 29. 19),0; Moldenke, Suppl. List Invalid names 6.
, Lilloa 8: 26, 19 ; d

8
aa
ef
2

Names 32. 192; Voldenke, Known Geogr. Distrib. Verbenac., [ed. 1],

1966 Moldenke, Monograph of Lippia 13

41 & 96. 1942; Moldenke, Alph. List Cit. 1: 263 (1946) and 2: 600.
1948; Moldenke, Lilloa 1: 47. 1948; Moldenke, Alph. List Cit. 3:
6936 1949; Mold enke, easki'G ans Distrib. Verbenac . » (ed. 2], 99

: Bt ool eldenke, B Résumé 117, 318, & 63. 19593; Troncoso,
niana 12: 258, maa a asl con & 287, fig. 1h. 1961;
pectisag ge Soc . Bot. 9: 184. 1961; Moldenke, Phytolo-

gia 12: "3 & 139. issee “Moldeckos, Résumé Suppl. 12: 10 oie 1965.
ustrations: Tronc oso, Darwiniana ana 12: 28), fig. 1). 1961.
0 ;

longer rite hairs interspersed among (sometimes
sometimes ws leaves decussate-opposite, dimorphic, ee or
qabeeiaize, lower peeiat atte green, oval to ovate or ovate-

sub

ed-extemate at the base, strigose—pilose
or substrigose on both surfaces with small antrorse hairs » some=
times roughened, with longer spreading and glandulose hairs usu-
ally ae aonet © pao ng (especially above) re drying, 3-
or 5=plinerv he base, sometimes rugose above because of the
bea potas venation which is conspicuous
beneath, not puncta ; inflorescence pedunculate;

rs and often also glandular—pilosulous; heads ovate to ovoid-
base: about 5 mm. long and wide during anthesis, 10 mm. long
and wide in fruit; exterior bractlets green, elliptic-lanceolate
and 3 mn. long by as mm, Wide during anthesis, broadly ovate and
5——6 mm. long by 3 mm, wide after anthesis, acuminate at the apex,
hr

r 0, ong, pubescent on the outside, obscurely sub
bilablate, "hs it90 subentire; corolla hypocrateriform, yellow or
lemon-yellow, small, densely puberulent on the outside, surpassing

c t

segments, about 1.6 mm. long and 1.5 mm, wide, smooth, the inner

This species was based on two collections made in Paraguay by
Emil Hassler —- the first (ans wpa) 2 September in the campo near
Carimbatay, and mr other (no. n part, or 546la) in November
in the campo near Igatimi. “The fon tenner of these has been designated
as lectotype of nia spattien by Troncoso and was photographed by
Macbride in the Delessert Herbarium at the Conservatoire et Jardin

14 PHYTOLOGIA Vol. 13, no. 1

otaniques at Geneva as his type photograph number 267).
Briquet (190) says: oa — remarquable espéce se place

a :
stifolia, il appartient 2 un groupe tout différent a capitu-
les dispoete. en racine terminal, alors que le L. tristis appar
tient au groupe des Euzapania 2 capitules axillaires." For his
var. normalis he says: "Folia caulinaria superiora lanceolata vel

ences for this species and its +t: al synonym are
"Plantae Hasslerianae XI (190) ae & a.

The Sk ee growing in fields and on campos,
flowering in September and November. aeniied (1961) says: "Es-
pecie basada en los sintipos Hassler 4554 y Su6la. Debe consid-
erarse como lectotipo Hassler 4554, por or figurar en la serie de

Pete del ejemplar Hassler 455k pero el mismo ejemplar posee

smo. La b sw
tea fructffera dibujada, anchamente oval, y la Piers y
a (los dos ejemplares del centro son Q, los de los pemer
dos 3 ) me permiten afirmarlo, El sintipo Hassler 56la es un in-
dividuo 3 - Sus Png son tempranamente vaducas | por or falta de fe-

var, aberrans pasa a ser una especie afin

3
beet
eile

@ perenne dioico, de 30—1,0 cm de alt., rizoma tuber-
oso-lefioso, » gree, que te numerosos tallos aéreos, erectos,
delgados, subcilfndricos, striados, verdes, que al secarse
ennegrecen en _ &pice, con largos pelos extendidos y pelos estri-
goso mezclados con pelitos glandulosos, breves, m4s abundantes
hacia el ye de los tallos. Hojas a ng peer las in-
ee pequefias, ovales a oval- » de

1966 Moldenke, Monograph of ee 15

das, atemadas hacia ja bas
de lat. y long. respect. 2 enteras 0 n 1-2 dientes en el &pice,
de borde recurvo, ennegreciendo al socarse, principalmente hacia

diente hacia el 4pice, las superiores mayores,
e,

long. y lat., de igual longitud que las flores o poco m&s breves,
hfspidas y glanduloso-pilosas hacia el 4pice; cAliz dimimto, di-
filo, a caduco, membrandceo, glabro o ligeramente pubescente;
corola citrina o amarilla (segin el autor), infundibulariforme,

posterior m4s pequefio, tubo breve, * pubescente exteriormente; es-
tambres ), didfnamos, subsesiles, inclusos, insertos en la porci-
6n superior del tubo corolar. Gineceo de la flor 3 tee nte

co, separable espontdneamente en 2 meri rto e
Zz acrescente, densamente albo-tomentoso, de superficie dorsal
lisa, convexa erna excavada." She cites Hassler 554

y cara int
(Cb—cotype) and 5i6la (X—cotype).

It should be noted here that the names L. angustifolia Cham, &
Schlecht., L. angustifolia H.P.K., and L. angustifolia Schau. are
all myms of the valid L. angustifo olia Cham., while L. angusti-
folia Morong is a synonym of the valid L. morongii Kuntze; L.
turneraefolia Cham. is a synonym of the y valid L. turnerifolia Cham.

all, herbarium specimens and ), mounted photographs, in-
cluding type material of most of the names involved, have been ex-
amined
pre fet PARAGUAY: Hassler 1554 [Macbride photos 24674] (Ca—
94h4354--cotype, It——photo of cotype, Kr—photo of cotype, N—co-
type, N—cotype, N--photo of cotype, S--cotype, W--photo of cotype).

LIPPIA TRISTIS var. ABERRANS Briq. in Chod. & Hassler, Bull. Herb.
Boiss., Piss 2, hz: 1161. 190).
Synonyny a aberrans (Briq.) Troncoso, Darwiniana 12: 258
& 267-268, — - 1961.
bliography: Briq. in Chod. & Hassler, Bull. Herb. Boiss.,
sér. 2, h: 1161. 190k; Briq. in gonad & aoparears Fi. —— 2

cub
117 & 463.1 1969; oeg o, “Darwiniana 12: 258, 259, 267—268, fig.
5, & 285. 1961; Troncoso, Bol. Soc. Argent. Bot. 9: 18h. 1961; Mol-

16 Peeey TO B04: 1k Vol. 13, now. 1
denke, Résumé Suppl. 5: 7. 1962; Moldenke, Phytologia 12: 2h.
1965.

une
?

leaves, 7--l12 cm. long, and the leaf-blades narrow~lan ceolate,
pellucid~punctate on the lower surface, with the lateral veins
hardly visible. It is said to hohe w 50. cm. tall, to be dioecious,

ic reference is cited as "Plantae Hasslerianae XI (190) 96."
type of the variety was —— by Emil Hassler (no.
5461) on the campo near Igatimi, Paraguay, in November, 1898, and
— deposited in the —— i at the Conservatoire et
n Botaniques at
tne (1961) ae sented cunt identific6 el ejemplar Hassler
5461 pp. como L. tristis var. aberrans, El estudio del ejemplar
tipo del Herbario de Ginebra me permite afirmar que se trata de
una pry once bien diferente de L. tristis. Difiere de asta es-
pecie por sus largufsimos pedinculos que que sobrepasan 2 a 3 veces
la longitud de las ho sa” (7-12 cm de long.), por sus hojas an-
gostamente lanceoladas » con pete laterales poco visibles y con
puntos pelticidos en el envés de la lémina. L. tristis posee di-

bf ere!
exclusivamente estrigosa en tallos, hojas y pediinculos en L. ab-
errans; pelos subestrigosos, pelos largos extendidos y pelitos
reves, glandulosos en L. tristis (ver fig. 14). Por todos estos
caracteres diferenciales aeons Ags la variedad aberrans debe
cia a constituir una mueva es

"L. aberrans es una especie nepal consistuyendo el ejemplar
Hassler 5461 del herbario del Ginebra. el individuo femenino (ver
figs.). Rojas 5268, aunque algo mayor en el porte, representarfa
el individuo masculino." She cites from Paraguay: Hassler 56
Mi oe 9117 (Bm, Cb); T. Rojas 5268 (Si), not as yet seen by

‘In all, 5 herbarium specimens, includi ng type material of both
names invo lived, and 2 mounted phot ographs a Pes been examined

Citations: PARAGUAY : Hassler 5161 (N—isotype) ; J8rgensen
4583 (Cp, N, N, N--photo, 8, Z--photo).

LIPPIA TROLLII Moldenke, Pigsaderis Ge PratPaat ties

Bibliography: —— Phytologi S hbbtit a9 Moldenke,
Biol, Abstr. 33: 1862. 1929; Malate. Fa umé 25 & 163. 1959;
gree niu rant 1: 7. 1959; Hocking, Excerpt. Bot. heb:

Shrub; branches rather slender, gray, decidedly tetragonal, gla-

1966 Moldenke, Monograph of Lippia 17

psbaee me te 3 branchlets and twigs sed slender, tetragonal,
very de ense ly short-pubescent, often more or less ac eate with
scattered n Bein nodes not anmlat e; erinoteat internodes 1—~2.5
t

r
1—3 mm. long, ee, whi tecpubeaonet; leaf-blades thin-c

ovate, 2-——3.8 cm. tone. ee cm. wide, sore a. subacute at
the apex and base, regularly serrate from near the base to the a-
pex with appressed antrors e teeth, rather comab a pubescent a-
bove with somewhat butboie banat ed hairs » very densely pubescent
nul

s
ieee impressed above, slightly prominulous beneath; veinlet
reticulation indiscernible above, eae or indiscernible be-
neath; inflorescence terminal, forming a simple or compound ra-
ceme $9 em. long and 2--l cm. eae, peduncles slender, tetrag-
onal, about 2 cm. long, densely short-pubescent like the twigs;
phe ica slender, tetragonal, densely short-pubescent; bracts foli-
ous, similar to the uppermost leaves but smaller, narrow-

with whitish hairs, one pair a at each node of the rachis; individ-
heads 2 or 3 per node, many~flowered, very dense, 7--9 mn.
long, about 7 mm. wide, on very slender sec ondary peduncles 5-6
mm. long and densely whi tish-villosulous; bractlets elliptic or
elliptic-lanceolate, about mm. long, 1.5—2 mm, wide, acute at
the apex, usually more or less falcate and conduplicate, scatter-
ed=pilosulous with rather stiff whitish hairs, more densely so on
the margins; calyx mimte, densely villous; corolla hypocrateri-
form, its slender tube about ) mm. long, the limb 2 mm. wide,
glabrous on the outside, barbulate in the throat within; fruit
not fas a
e type of this distinctive species was collected by Karl

troli (no. 1207) — 5 whose honor it was named — at Vallegrande,
Santa Cruz » Bolivi an altitude of 1900 meters, on January 28,
1928, and is skied As the Sictoec te of the Botanisches Museum
at Berlin. The species nen Pere found at altitudes of 1500
1900 meters, flowering amuary.

In re 3 owning pes iauaer including the type, have been

e d by me
Citations: BOLIVIA: Santa Cruz: Troll 1054 (B), 1207 (B-type,
Z——isotype).

LIPPIA TURBINATA seghtg » Abhand. Kaiser. Gesell. Wiss. GUtting.
19: 243. 187
nymy : Lipps a disepala R. A. Phil., Anal. Univ. Chile 90:

syn. 190. Lippia poleo Lillo ex Moldenke, Suppl. List Invalid

18 PRET ULOGI Ss Vol. 13, no. 1

Names 6, in syn. 1941. Lippia poleo var. stifolia Lillo ex
Moldenke, Résumé Suppl. 10: 6, in syn. 196).

Bibliography: Griseb., Abhand, Kaiser. Gesell. Wiss, GOtting.
19: 243. 18743; Lorentz, Veg. Nordeste Prov. Entre Rios, ed. ae
150. 1878; Griseb. Abhand. Kaiser, Gesell. Wiss. GUtting.

(Symb. Fl. Argent] } 278. 8. 18795 ‘illo, Fl. Tucunan 9h. 1888; ee
in Hook. f. & J : 96. 1894; Re As Phil., Anal.
Univ. Chile 90: “62 “tasgs) ast ‘oti 415. 1896; Kuntze, Rev. Gen.
Pl. 3 (2): 252 & oo 1898; Durand & Jacks., Ind. Kew. Suppl. 1:
pct 1903; Hayek in Engl., Bot. Jahrb. 2: 166. 19085 Tavares,

prope del = oad 106. 1919; Sanzin, Anal. Soc. Cient. Argent.
Buenos Aires 88 (i—iv): 102. 1919; Doming., Mat. Med. ee 1928;
Seckt, Fl. Cordob, }18, 1929-1930; Stapf, Ind. Lond.. }:

1930; ’seckt, Rev. Univ. Nac. Cordoba 17: 90. 1930; ose: Gesdha.
Pl. Amer. Sud 350, fig. 814-818 b—-f. 1933; Parodi, Rev. "argent.
Agron. 1: 201. 193k; Latzina, Trab. Inst, Bot. & Farm, Buenos Ai-
res oa ahi 112. 1935; Troncoso, *Darwiniana 3: 52. 19373 Latzina,
Liles 1: 169. (29375 Monticelli, Liles 3: 357. 1930; Moldenke,

nvali
graph. Distrib, Verbenac., [ed. 1], Ka he iB, 13, 73, 95, & 96.

19hes. Sree: Sage List Invalid Names 192; hepa get

: 426—, Bed and 10: 33h & © 380. 19

P satit sag 86. 195; gee Acufia, Catalog Ee Ps Cata-
marq. 17. 1945; Moldenke, Phytologia 2: cz : at 1945; Moldenke,
Alph, List Cit. 1: 73, 95, 96, 12h, 21, 1946; Moldenke,
= : 148 & 149. 1916; Moldenke, ater: gn Invalid Names

Suppl. 1: lj. 1947; Lorentz, te gt Prov. os,

2, 150. 1917; Cozzo, Revist. Me 1h: 252. 19473 Yol-
denke Phytologia 2: 338 (ag) se "Os ag 1948; Moldenke, Lil-
loa 1): 78. 1948; Moldenke, Alph. List Cit. 2: 355, 368, 369,

denke, Known e Di °
73, 101, 105, 162, 189, & 1. 1993 E - N. & A. L. Moldenke, Anal.
Inst. Biol. Mex. 20: 1 1949; Moldenke, Phytologia 3: 30 & 305.
1950; Acevedo de feria Bol. Mus, Nac. Hist. Nat, Chile 25: h5.
1951; Troncoso, Darwiniana 10: 80-83. 1952; 225 Moldenke, Phytologia
5: 96. 195h; Acevedo de Vargas, Biol. Abstr. 28: 90.'195h; Hock-
, Dict, Terms Pharmacog. 165. ee woldenke, Résumé 8), 121,

126, 220, 310, 312, a, a & 463. 1959; Mufloz Pizarro, Espec.

nt. Deser. Philippi 110, 1960; omwied § atau Suppl - 10: §&
6 (1964), 11: 5&7 (196 él). and 12: Se 1965; Moldenke, Biol.
Abstr. 16: 326. eos Moldenke, Phytologia 12: 26, 47, 61, 63,
196, 218, 219, & 288. 1965,

Ts mows : Tavares, Broter. Zool. 13: 97-98, pl. 3, Pe

—5. 1915; Ss , Anal, Soc « Cient. Argent. Buenos Aires 88 (i-—-
nh 102, isi9; oe. Zoocéd. Pl. Amer, Sud 350, fig. si--818

1966 Moldenke, Monograph of Lippia 19

be-f. 1933.

Shrub, 0,l--2 m. tall, canescent or glabrous, much-branched,
very aromatic; branches erect, virgate, terete or obscurely tet-
ragonal, 23--30 cm. long, often about diameter

leaves mostly ternate, sometimes decussate-opposite

anceolate or oblanceolate, 1.8--2 cm, ; « Wide, acute
at the apex, appressed crenate-serrate al margins, some-
times ent. low and with about 3 teeth per side above, narrow-
ed and attenuate into the petiole at the base, trig-
illose, the se rs and on the marg with a tubercle

hi a
lar-roughened beneath, the venation prominent or very prominent
beneath, impressed (often deeply) above; inflorescence

rt, a » Single in each axil, subequaling or slightly
shorter than the subtending leaves or only 1/) to 1/3 as long,
subinvolucrate; peduncles equaling the heads in length, very
8 appressed-pubescent; heads subglobose, 5—7 mm. wide;
bractlets rhombic-subrotund or broadly ovate to obovate, flat,

lower ones 3-1); mm. long and 2 mm. wide; flowers very fragrant;
calyx diphyllous, the sepals ovate-lanceolate, hispid with spread-
ing setae, keeled, the keel lanate-ciliate; corolla small, hypo-
crateriform, violet, lilac, or wine-color to purplish-white,
white-lilac, or white, about 5 mm. long, the tube about twice as
long as the calyx, the limb regular, wide

The type

as sheet number 5/813 in the herbarium of the National Museum of
ve

Chile. The of: 1, disepala was collected by August Figueroa
in the Andes of Jorquera, Atacama, Chile, sent to Philipp: Man-

along the Rfo de la Tala, Catamarca, Argentina, on February 3,
1873, and is deposited in the Britton Herbarium at the New York
tani

Bo en.
Philippi (1895) says for his L. disepala: "Parece que las dos
Cocas del fruto no se separan en la madurez, pero no tanto frutos

bien maduros. — ?Seria acaso la L. as folia Rich., planta
comun cerca de Cérdoba, i en varias parted del mundo? La mestra

20 Pay TO L6G Ik Vol. 13, now. 1

difiere de la descripcion de esta especie dada por Schauer....por
sus tallo sos, no erizados, los pediinculos bastante gruesos,
las

traza de querer florecer.

The length of pistil i in relation to the size ne the pollen in
this species is discussed by Covas & Schnack (195). H. H. Bart-
lett 20181 has its leaves quite narrow and may actually represent
f. angustifolia Osten. The flowers are described as "violet" on
Venturi 609 & 1292; "lilac" on Cuezzo 25; “purplish-white" on T.
Meyer 37355 twhite-Lilac" on Cuezzo 978; "?hite and wine-color"
on Monetti 143; and "white" on Brizuela 581, I. M. Johnston 4987,

T. Meyer 1190 & 4235, Rufz Huidobro 3043, Schreiter s.n., A.G.

Schulz 2891, Sparre 1198, Terribile 6h7, and Venturi 336 pet & 8120.

oods along rivers, at the edges of rive streams and
roadsides, at altitudes of from 70 to 2500 meters, flowering from
October to June, rui rom

to it as "abundant" in Catamarca, calls :

in valley" in the same province. Villafafie refers to it as

busto coposo" in Cérdoba, while Johnston says "local in dryish

meadows" in Atacama. Vernacular names reported for it are "Argen-
gt tt

pe .
pais", and "té del pafs", Di Lullo and Meyer both aver that it
is a medicinal plant. Parodi (1934) says "Arbusto achaparrado, de
icas y

J aromfticas y caducas, originario de las sierras de Cér-
doba. Su f. medicinal y la facilidad de multiplicarse
por gajos y semillas permitido la difusién de su cultivo, aun-

en escasa proporcién, por los amante la medicina casera.”
Troncoso (1937) "La he visto escasamente cultivada en s
‘ de sus hojas utilizada medic

mn dae ente.
Originaria de las sierras de C6érdopa. Bs. Aires: no. 83 (ex. cult.
in Hort. Bot. Fac. Agr. Bs, As.) ." a (2938) 8a says "se la
cultiva en muchas partes, aun en Bueno:
In the Instituto Miguel spend the 2 original ce Pedant refer-
ences for L. aprica and L. dis are given as "Philippi, Pl

The La
"Latzina, Index II (1937) 189", the Sanzin reference as "Sanzin,
Verbendceae (1919) 102", and the Cozzo reference as "Cozzo, Raices

1966 Moldenke, Monograph of Lippia 21

gemiferas 2 ee 252."

Material has been misidentified and distributed in herbaria
under the names L. alba wo) N. E. Br., L. asperifolia Rich.,
L. geminata Kunth, L. geminata var, microphylla Griseb., and L.
nodiflora Mic hx.

On the other hand, the Galander s.n. [Cérdoba, 2.1 -1880], Hier-
ous sen. [Rfo Cérdoba, 12.X1.1877], Lorentz s.n. [Cérdoba, 2/

80], T. Meyer zr 13015, 1322, 13436, & 13 2, Ruiz Huidobro s.n.
[Barrio Mirador, 19-IV-1 » Scala | ala 201, and Villafafie 681, dis—
tributed as L. nae are orm ie f, sassvisnlia Osten,

LEE St gg ip rl in

655, 2429, 2683, 3799, 4288, 8115, 8120,

In all, 195 herbarium specimens and 5 mounted photographs, in-
cluding type or ilgstd oe material of all the names involved, have
been examin as

Citations: CHI son. (Herb. Mus. Nac. Chile
54814] reteset I. op Johns ae et 7 (S, W-1497728); Ricardi &
Marticorena 37/132 (Ac); Werdermann am 7 (Ca—278556, Gg—123382,
S, W—170923). Tacna: Allende s.n._ Sen. (Herb. Mus. Nac. Chile 5813]
(Hetphiote); ARGENTINA: — Aires: Boman om [Sierra de la
Ventana, Avril 1908) (Br). Catamarea: b (N); ed
1g (N), 382 (N), L521 (N, S), 528 (N), da ti (N, i eh, 581 (N),
(au—121218, N), 780 O (N, Ok), 860 (N, Ur), 975 (N), 987 (n), eae
(N, S), 1083 (N, S), 1220 (N), 1252 (N); Castellanos s.n. (Herb.
Mus. Argent. Cienc. Nat. fat. 30/604) (N (N); Hauman s.n. (Herb. Mus. in
gent. Cienc. Nat. 23860] (N); JUrgensen n 102) [Herb. Inst. Miguel
Lillo 32080; Herb. Osten 12h] (Ca——20 (Ca—202220, N, N, Ug, W—70750),

[Andal, ndalgald, N v. 1916; Herb. Fac. . Med. Buen
Teak} (Sp—-25782) ; Hocetta & Hieronymus 373. (N); a ee & wol-
denke 19732 (Es, Lg, N); Pierotti s.n. [18/2/hh] (Ca), s.n. [12-
IIT-19hh] (W—-193173h) 3 Quintero intero 65 (N); Villafafie 1147 (N) (N). Cha-
co: T. Meyer 8759 (N); Ragonese vagonese 8 (Herb. Mus. Arce: Argent. Cienc. Nat.
13345) (N); A. Ge G. Schulz 1188 z 188 (N), 6389 (Z). Cérdoba: Bale 961
(N); H. H. Bartlett (a= in eet epee 2018) (it, pal
190876); Burkart 20 ans 3 Castellanos s.n. [Herb. Mus.
Argent. Sain, We Nat. 31197] (N); Job 55u (N, N); Kuntze s.n. [Cér-
ng ce cigs ras; 2 Loronts tom pide mip Tract

isotype, of t hoto o.
ree); Nicora 578 Os otDeneht "33" (Ny, iis70 UN (N, Vi); 0 0'Donel &
Rodriguez Vaquero 66091); Pierotti 5121 (N); Rodrigo odrigo 278
(N, S), 22 218], (N, Wie psta 783 ( (Au—-12121h), 3166 (Gg—I10923);

22 Pee T: 6 L.0-0:1-8 Vol. 13, no. 1

arre 1379 (S); Terribile 647 (N), 692 (N, S); Varela 08 (W—
EES 7)3 V Villafafie L (N, N, i St), 69 (N). Entre Rfos: Lorentz Se

ne [Concepcion del Uruguay , 1880} (W--12336h2). La Rioja: Cas-
tellanos s.n. [Olta, Feb. eb. 25, 19L0; Herb. Mus. Argent. Cienc. Nat.
33698] (N); Cuezzo 978 (N, Ok, Rf); Gémez s.n. [Herb. Mus. Argent.

Cienc. Nat. 28/775) (N); T. Neyer 4190 (Ca—7718), Ca—16h777),
4235 (Ca—77199, N, N); Varela 769 (Ca, N). Mendoza: Covas 150h6
Ws A. Rufz s.n. tHerb. . Mus. Argent. Cienc, Nat. 23928] (N); Ru- Ru-

z Leal 1142 (N), oe (N), L490 (NW), 6754 (i); Sanzin 1832 [Herb,
ee en 12799) (Ug). alta: Castellanos Son. [Herb, Mus. Argent.
Cienc. Nat. 181] 5 Correa 5 [Herb. Inst. Miguel Lillo 3701]
(Au, Ca—155116, N); Garolera | & Romero s.n, [Cuesta del Obispo,
21-I-19h7] (N); Hunziker 978 (W—180]058); Lorentz & Hieronyms
373 (W--2815hh); Lourtei Lourteig 1031 (N); Malvarez 176 (N); T. Meyer
o (N), 3447 (Herb. Inst. Miguel Lillo 17335] (N), 348 [Herb.

. Migiel L: Lillo 35570] (En), 3453 (Herb. Inst. Miguel -
36871) (N), 3726 [Herb. Inst. Miguel Lillo 35473] (N), 3735
(Herb. Inst. Miguel Lillo 35)82] “(ca-—882510, N), 1209 (N, S);
Moldenke & Moldenke 19738 (Es, Lg, N, Sm); O'Donel) 251) (Ca—
165621), 25L8 (Ca—16813); Rufz Huidobro 513 (W—193)085); A. Ge
Schulz 1449 (N), 2891 (N); Sota 162 (N); S Sparre 1198 (S, S); Wall
& Sparre s.n. [near Salta, 4/12/,6] (Ew, Ew, Bw). 5 San Juan: H.
ce Bartlett 19559 (ca—772029, Mi, N, W—190)563) ; Castellanos
05 (W—21982)3) ; Cuezzo 1845 (N) . San Iufs: Bruch & Carette

Son. [Alto Pencoso, 1.191] (N, N, N); Digilio & Grassi 2017 (N);
Rutz Leal 910 (N); Varela 75 (N) 5 Yepes s.n. (Herb. Mus. Argent.
Cienc. Nat. ;» 29/90] (N). Santiago del Leamige Bale (Ca-~
165673, N); Di Lullo 74 (Herb. Inst. qual 1atie ieee ) (N, a“
1934387); P. Garcia 958 (Ca); Lillo ta [Herb. Inst. Migwel
1o 32088] (N); F. Luna 20 (N), 31 (S)3 Pierottd 8 (N); Rutz Pe
dobro 3043 (N, S). Tucumén: Cuezzo 25 (S); Lillo 18826 [Herb.
Inst. Miguel Lillo 32077] (N); T. Meyer 3093 (N); Noose 143
(Herb. Osten 10875] (Ug), 1949 [Herb. Inst. Miguel Lillo 32090]
(N, Vi); Peirano s.n. [El Nio, April 29, 1937; Herb. a Miguel

Lillo 32616] (N), s.n. [Amaicha, Dec. 1917; Herb. Osten 12190]
(Ug), sen. ee == ee Dec. 1917; Herb. Osten 12199]
(Ug); Sparre 78h (S); Vv 609 (W—1591226), 103k (N), 2361
[Herb. Osten 17265] (N, te 8 aieo2ss). 4292 (N, N, S, Ug, W—
1591439), sen. (Herb. Mus. Argent. Cienc. Nat. 2 3813] (N); E. E.
Wall 10 (Ew); Wall & S s.n. [Trancas, 19/11/6] (Bw, Ew, Ew,
Ew, Ew, oi Ew). CULTIVATED: Argentina: santraneh i OH, N)-

1966 Moldenke, Monograph of Lippia 23
LIPPIA erry f. ANGUSTIFOLIA Osten ex Moldenke, Lilloa 5:
31 29,

Bibliography: Moldenke, Lilloa 5: 31 (190) and 8: ng 192;
Moldenke, Known Geogr, Distrib. Verbenac., [ed. 1], 43 & 96.
1942; Moldenke, Lilloa 10: 3h. 19h; Moldenke, Phytolo gia 2: 82

ph.

ke, Lilloa en wis ae Moldenke, ey 481.
denke, Alph t Cit. 2: 377, Ubl, & 537 *Ga5l8), 3: 673, ius,
909, & 913 {asls), and h: 129i. 199; Moldenke, Phytologia 3: 76.
1949; Moldenke, Known G Geogr. Distrib. Ver rbenace, See . gab ae -
191. "19495 H. N. & A. L. Moldenke, Anal. Inst. Biol.
194.9; ee Revist. Invest, Agric. 5: "22, 83, 36, & age Fos ‘ige)
and 6: 70, 75, & 83. 1951; pea » Inform. "Mold. Set 51 Spec.
3. 19564 “tolaeneh Résumé 126, re & stg 1959; Moldenke, Résu-

suppl. 1s 7. (1959) and 1: 5&

This form differs from t ee form of the species in its
uniformly very narrow leaves, which, including the petiole, are 1—
3 cm. long and only 1—l; mm. wide.

The type of the form was “gollected by Augusto Gustavo Schulz
(no. 1489) in cultivation at Colonia Benftez, Chaco, Argentina,
in 0 October, 1933, and is deposited in the Britton nectar at

New Yo rk Botanic al Garden. The collector notes that
plant is medicinal, used in the treatment of stomach troubles and

to aid digestion,

Ragonese (1951) describes it as follows: "Arbusto, de hojas
opuestas, lanceoladas, es has, de 2 a 5 mm de ancho, con el
borde aserrado, cubiertas por pelos cortos y rigidos; flores

» pe en inflorescenc
infusién de las hojas y ramitas de esta especie se le atribuyen

as
propiedades estimulantes y ténicas, siendo muy utilizada en medi-
cina popular. Es originaria de las llamuras y de la regién ser-
rana cordobesa." Collectors also describe it as a low shrub or
depressed aromatic shrublet or a tree-like ssh 4 to 1.5 m. tall,
with white flowers, inhabiting ay fields, streambanks, the
edges of arroyos, and fencerows at 390 to 1000 meters altitude,
flowering from October to May, and fruiting in December and Janu-
ary. Common names reported for it are "poleo" and "poleo crespo".

t

"htufig bei Cérdoba". Meyer reports it as adventive in alfalfa
fields, and notes that it is cultivated for its stomachic ee
ties in the Chaco. The corollas are described as "white"
Cuezzo 930, Lossen 221, Luna 31, Ragonese 621, and Villafafte 181,
189, é1i, 7 & cc ye

Material has been misidentified and distributed in herbaria as
typical L. turbinata Griseb. and under the name Lippia nodiflora

In all, 76 herbarium specimens, including the type, and 2 moun-
ted photographs have been ed me.

Citations: CHILE: Atacama: Ricardi & Marticorena 3615/33 (Ac);
Werdermann 1,8) (N). ARGENTINA: Chaco: T. Meyer 677 [Herb. Osten
227057 (Ug). cérdoba: Balegno 330 (Ca—166170), 1122 (Au—121213,

2h PHYTOLOGIA Vol. 13, no. 1

N); H. H. Bartlett 19237, in part (Mi), 20112 (MMi); Capitanelli
S.n. . [Rutz Leal 15 15972] (Ss); Castellanos s.n. Neat Brochero,
Nov. 12, 1927; Herb. Mus. Argent. Cienc. Nat. 11672] (N) 3 Cups °
930 (N); G. Dawson 78 (N, N); Escalente 37 (W--198797L); Galander
s.n. [Cérdo oba, 2.11880] (Br); Gillies s.n. [Province of Cordova]
(Lu); Herb. A. Gray s.n. (Cordova, 1872] (C); Herb. Mus. Hist.
Nat. Montev. 960), (Ug); 5 Hieronymus sen. [Rfo Cérdoba; 12.x1.1877]
(Br, N—photo, Sp—-25786, Sp, W-—2815)3, W—117797L, Z—photo);
J. H, Hunziker 1291 (N); * Kuntze sn. [Cérdoba, XII .91] (N)5; Lor-
ne Son. jen. [Cérdoba, 2.1.80] (N, W—~-701568); iveaies 221 (Ba, Vi);
Meyer 13015 (s), 13329 (S), 13436 (S), 13549 (S); M Moldenke &
aieeae 19714 (Es, N); O'Donell 1 1956 (N); D'Donell & Rodriguez
Vaquero 09 (Ca—166101); Osten 13106 (Ug); Pontusi Sen. n. [Rutz
Teal 12 16131] (Ss); Ragonese 62l1 [Herb. Inst. Bot. Minist. Agr.
Nac. 68447] ( sh pagonese & Piccinini 6148 [Herb. Inst. Bot. Min-
ist. Agr. Nac 75) (N); M. Rufz 421 (N); Rufz Huidobro 255
(Ca, N), 452 fe tosiioa): San. [foledo, 17-I¥-1 OLY (N), sen.
[Barrio Mirador, 19-1V-19hh] (W—-2019637) ; Rufz Leal 12218 (Z);
Scala 201 (W--1691708), 2001 (N); Villafafie 181 (N), 266 (N),

300 (N), 489 (N), 622 (N), Soh (N), 717 (N), 770 (N); Wall &

Sparre s.n. [La Falda, 15/1276] (Ew); La Rioja: Rufz Leal 16513
(Rl). Mendoza: Rufz Leal 7088 (N), 8507 (N), 907 (N). San
Luis: Kurtz 8523 (N); Butz Leal 10438 (N). Santa Fé: Balegno 616
. Santiago del Retatos Cuezzo 2L55 (N, S); M. J. de Luna 31

ect ent mene remanence

palegno
1227 (N, S); O'Donell 4316 (N); Schreiter 7962 (Herb, Inst. Miguel
ad 32616] (N). CULTIVATED: Argentina: A. G G. Schulz 1489 (N—
type

LIPPIA TURBINATA f. MAGNIFOLIA Moldenke, Résumé Suppl. 10: , nom.
nud, Ms ; els Phytologia 10: lie. June 18. 196k.

bra oe ippia fissicalyx Troncos Stal wc 10: 83—86,
fig. 1952.

Bibliography: Troncoso, Darwiniana 10: 83—86, e 6. 19523 Ge
Taylor, Ind. Kew. ato hs, 82. 1959; Moldenke, ‘agents: Suppl. 7:
6 (1963) and 10: hk. 19643; Moldenke, Phytologia lo: 172. 1964; Mol-
a Biol. Abstr. 46: 326. 1965; Moldenke, Résumé Suppl. 12: 5.

Illustrations: Troncoso, Darwiniana 10: 85, fig. 6. 1952.

This form differs from the typical form of the species in hav-

p
and the peduncles usually 2 per axil. Collectors describe it as a
shrub, to 1.5 m. tall, with corolla varying from white to flesh-
color, oo "Lilac » rose, violet, or violet-white.
The type of t e form was collected by Santiago Venturi (no
8120) at an erie de of 1300 meters, Campo Quijaro, Rosaste da Ler-

1966 Moldenke, Monograph of Lippia 25

Salta, Argentina, in January, 1929, and is deposited in the
United States National Herbarium at Washington. The type of L.
fissicalyx was collected by Venturi (no, 8115) at the same local-
ity on January 19, 1929, and is deposited in the herbarium of the
Instituto Botanico Darwinion at San Isidro » Argentina

in December and from March to May. The only common name recorded
for it is "poleo". The flowers are described as "violet" on Ven-
turi 655; "rose" on Venturi 4288; "violet-white” on Venturi 2129;
"lilac" on Herrero 72 and V and Mebors 8115; "flesh-color" or "pinkish-
white" on Lorentz | & Hieronymus 236; an and "white" on H. H. Bartlett
19778, T. Meyer Meyer 3195, and Venturi 2683, 3799, & 8518.

tall, branched; branches tragonal — sh=b. » Ssubgla
rous; branc Fs onal, » pubescent; middl
internodes 3—l cm. long, on floriferous “aoa to 5.5 cm. long;
ves decussate-opposite, rarely val or sometimes o
va 1—),.5 em. long (including the eter wh. 0.5--1.7 cm.
oe Dper acute at the apex, cuneate a e t bicolor-
ed, rly dentate on the upper 2/3, aul at the

the lamina (the hairs callose at the base, especially those

ward the leaf-margins), peg igri and finely reticulate
beneath with the hairs larger along the venation, densely glandular
punctate among the pubescence; venation deeply impressed above,
especially toward the margins and in the teeth, prominent beneath;
peduncles 1 or more usually 2 per 3. 3-6 mm. long, pubescent;
heads hemispheric, 5--6 m. wide; bractlets in many series, the
lower ones larger, firm, oval-lanceolate, sieges mm. long, 1—1.8
mm. wide, 3-veined, pubescent, the upper ones oval, concave, 1.8—
2mm. long, 1,7—-2 m. wide, Ascii: sprig pubescent, ciliate-
margined, glandular—punctate; calyx in bud 2-Lobed, a ag anterior

two p. te

the keel; corolla small, tubular, 3.5--l; m. long, bilabiate, the

lower lip 3-1 obed, the middle lobe Sintabpaace the upper reduced

and 2-lobed, the tube about 3 mm. long, gently dilated toward the
and ar-punctate

di b
cluded; ovary oval, glabrous; stigma ae aba eg: fruit dry, ob-
Gris., la especie or sus hojas cara ir ae vez —

m&s affn, po
das, por sus capftules generalmente de a 2 por axila;
adem4s semiesféricos, no o muy poco alargados en la ee al

26 PHITTODOGIA Vol. 13, no. 1

munca turbinados, y por sus hojas m4s anchas y meno y
rigidas. Sin embargo, la existencia de formas de hojas anchas en
L. turbinata hace a veces diffcil la separacién y sugiere la idea
de la existencia de hfbridos,
"Por inscripciones en ejemplares del Museo de Cérdoba, parece
evidente que Grisebach consideré esta nueva especie como Lippia
+ microphylla, pero también aplicé ese nombre a ejem-
plares eee ae de Lippia turbinata. Es poco probable, sin em-
bargo, que se trata de dicha variedad por haberse descrito ésta
siete’ aeael nte de Cuba
Ind. Isl., p. Wg? (2864; — el mismo autor m4s tarde la cita
para Tucum4n y +, Symb., p. 278, 1879). Lippia fissi-
calyx, en tod so = sine de Lippia turbinata y no de Lippia
geminata HBK. Por otre parte para la nomenclatura, si se confirma
esta sin sinonomie, la variedad no tiene mayor interés; como ya existe
una L. microphylla Cham, (1832), no era — sre —
de de Grisebach al rango de especie." She
Lorentz & Hieronymus 236 (Cd) & s.n. es riwey a trakel. (ca) and
Vattuone | 29 (Si), from Jujuy Burkart & Troncose sen, [Jujuy, 15.
II.1940] (Si--11097), and from Tucum4n Lorentz & Hieronymus 111)
(Cd), D. Rodriguez 212 (Si), and Venturi i Bee ts Si) & & 2Le9 (s acca
Material has been misidentified and d 5 ee in herbaria
under the names L, geminata var. microphylla Griseb., L. globi-
flora var. geminata f, glabriuscula Kuntze, L. poleo Lillo, and
L. turbinata Griseb. Venturi 8518 was mistakenly cited by me as
Ls turbinata in Lilloa 1}: 47 and his 655 & 8120 as L. turbinata
in ny Alph. List cit.
In all, “2 herbarium specimens, — type material of both
names involved, have been examined
Citations: ARGENTINA: Catamarca: tetsaiis 237 (N), 810 (N),
1217 (W—2123672); Luna Risso 196 (N). La Rioja: T. Meyer 3495
(N, N, W—1909097). Salta: H ae Be H. Bartlett 19678 (Mi, W—190)616) ;
Herrero 72 (W—20h9728); Lorentz & Hieronyms 23% (N, Weslo )s
Venturi 3799 (W--1591280), 8115 8115 (Gg—16665), W—1L)3377, W
1591398), 8120 (Gg—-166670-——isotype, N-~isotype, ¥——1,3380--type),
$.n. [Agua Cal. Caliente, 2.I1V.1925; Herb. Mus. Argent. Cienc. Nat.
orem (™). Santiago del Estero: H. H. Bartlett 19778 (Mi, W—
: Monetti s.n. [Leal, Las Lecinas, 25.V.1913;
—s Miguel Lillo 32086] (N, Ug—1193h, W--1802569) ; De
Rodriguez 212 [Herb. Mus. Argent. Cienc. Nat. 23810; Herb. Inst.
Miguel Lillo 3209;] (N, N); Venturi 655 [Herb. Inst. Miguel Lillo
32092] (N, W--1591227), 229 [Herb. Osten 17238] (Gg—-1518h3, Ug,
W—13h3280, W--1591256), 2683 (W—-1591263), 1268 (N, W—1591435),
8518 (E—96196h, N, S, W--1591)06) .

PPIA TURNERIFOLIA Cham., Linnaea 7: 217. 1832 [not L. turneri-
folia Chod., 190k, nor Hieron., 1917].

1966 Moldenke, Monograph of Lippia 27

Synonymy: Lippia turneraefolia Cham. apud Morong, Britton, &
Vail, Ann, N. Y. Acad. Sci. 7: 196. 1893 [not L. turneraefolia
Chod., 190]. Lippia turnerifolia a normalis Ki Kuntze, Rev. Gen.
Pl. 3 (2): 253. 1898. -=2 turneriifolia Cham. ex Rosengurtt,
Estud. Prad. Nat. Urug. 3: 235, sphalm . 1943. Lantana turnerae-
folia Cham. ex Moldenke, Alph. List Invalid Names Suppl. 1: 13,
in syn. 1947. Lippia turnerifolia Cham. & Schlecht. ex ‘Maiden,
Alph, List Invalid Names Suppl. 1: 15, in syn. 1947. Lippia tan
nerfolia Cham. ex Moldenke, Résumé Suppl. 12: 11, in syn. 1965.
Lippia t peels var, normalis Kuntze ex lioldenke, Résumé
Suppl. 12: rg nny 1965,

Eieltceranhos Cham., Linnaea 7: 217. 1832; Steud., Nom. Bot.,
ed. 2, 2: 5h. 180; D. ’pietr., Syn. Pl. a Rue ot Walp.,
Repert. Bot. Syst. \: heed 1845; Schau.

in Mart.,

a ra: 1851; ee tz,
Veg. Nordeste Prov. En tre Rfos, ed. oe 122 & 160. 1878; Griseb.,
Abhand. Kaiser. Gesell. Wiss. GUtting. 2h: [Symb. Fl. Argent.

277. 1879; Lillo, Fl. Tucuman 9. 18 8; Morong, Britton, ; oer

1
Boiss., sér. 2, k: 1158 & 1159. 190; Briq. in Chod. & Hassler,
Ag Pog es 2° (11): 493 & 49h. 190h; Briq., Ann. Conserv. &
ard. Bot. Genév. ae 311. 190); Glaz., Bull. Soc. Bot. Franc
8, Mém. 3 Ba eony Herter, Florul. Urug. 105. 1930; Moldenke
in’ Fedde, R ov. Sp. 39: 146. 1936; Herter, Revist. Sudam.
Bot. h: Seer 186, 19373 Gallinal, Campal, Aragone, &

1939; Moldenke » & 431-432. 190; Ragonese,
arwiniana 5: 19121; ‘inidecke, * Known Geogr. Distrib. Verben-
ac., [ed. 11, 38, "yO, bi, 43, & 96. 1942; Moldenke, Lilloa 8:
427. grey odi, Darwiniana 6: 175. 19135 Rosengurtt om

Prad. a 3: 235. 1943; Moldenke, Lilloa 10: & 381
(19h) we 12: 129. 196; Moldenke, Alph. list Cit. 1: 28 & 39.
196; eae Fl. Rio Grande do Sul 221, 223, & 234—-235, fig.
Be osha teelcia or Ok Sa Ett) ke
& 150 19h? eee 1 $ - ; . 3 Mol-
denke, a List Invalid Name es Su - = aS & 15. 1947; Molden-
ke, Lilloa 14: hO—1, 4h, & 48. Alph. List Cit.
: 358, 368, 426, bh8, 457, Es 5. a & ‘e628 ob), 33 1 8
19h TT,

98
1251, & 1256. 1949; Moldenke, Known Geo . Distro: Tervenasy
2

] 99, logia
3: 287. 1950; Stellfeld, "trib. Farmac. 9¢ 10): 169. "1951; a
Fl. Paran. 7: 12. 1957; Moldenke, R Sok, = okt, 219, Lem,
308, 318, & 163. 1959; racer Résumé Suppl. 1: 7. 19595 —
Fl. Paran. 16: 60 (1 and 17: 6. 1961; Troncoso, Bol.
Argent. Bot. 9: 18h. 1961; Sane Résumé Suppl. lo: 6 (396h),

hk: 1

28 Peat O L064 Ts Vol. 13, no. 1

ll: 7 (196), and 12: 11. 1965; Moldenke, Phytologia 12: 2h, 26
305 Bs 131, 294, 347, & 356. 1965; Rambo, Pesquis. Bot. 21:
23 & [59
Illustrations: Augusto, Fl. Rio Grande do Sul fig. eek 196.
Dioecious dense bushy shrublets, usually 0.5--0.8 m
sometimes to 1.3 m. tall, witha creeping rhizome stens bec -
ceous or suffruticose Mes erect, simple, rather tere ie sute
white rigid spreading hai , finally calvescent and \ soidtiatal be-
low; leaves Fa pipe rg varying in size and shape with the
stature of the entire plant; petioles short; leaf~blades lanceo-
late or oblong to obovate, 2--5 cm. long, usually about 1 cm.
wide or wider, acute at the apex, crenate-serrate along
gins, entire and cuneate at the base and there narrowed into the
sekinla: * ev gshr aca he apart above, strigose above with bulbous-
based hairs and hars rough, hirsute on the venation beneath
but finally merely strigose there; inflorescence axillary and ter-
3 peduncles filiform, sometimes short and iach y 1/3 as
long as the subtending leaves » sometimes elonga’ surpassing
the leaves and stems, hirsute; heads rounded or ‘onsidieas: dense,
to 1.3 cm. wide, involucrate; bractlets ovate, imbricate, soma
ate at the apex, equaling the corolla-tube; flower ery
corolla hypocrateriform, yellow or light-yellow to deaporaiioe:
sometimes reddish inside.

ected by
Friedrich Sellow in dry grassy places on dry hillsides near the
b

fields on mountainsides, and at the edges of woods, at altitudes

of 400 to 2000 meters, flowering from August to liarch. Balls

found it on "steep slopes in heavy clay soil amongst undergrowth

and trees, usually on be edge of thickets". Rosengurtt calls it

"frequent" in Uruguay, Hunziker refers to it as "rare" in Salta

peg Parodi (19)3) tice eeu en praderas humédas" in Corrien—
Se.

Briquet (190) says "Cette —_ est extrémement rapprochée
de la eetnetants [L. asperrima], dont elle se distingue cependant
nae ses pedoncules p. plus courts, nullement glandul eux-visqueux,

s hirsutes per de longs poils raides étalés ou subas-
ohitante trés denses."

The flowers are described as "light-yellow" on T. Rojas 377,
"deep-yellow" on Balls 5931, "yellow, reddish inside" on Tt. , Rojas

is ea, kale

23037, 23346, & 36735, F. C. Hoehne §.n., Hoehn

ae T2h7, Job 15)3, T. Meyer 3452, Oste “a 5 : ‘She abe
261,36, T. Rojas Wi, A. G. Schulz 6 6983, Sleumer & Vervoorst 2778,

Stellfeld 1039, and Venturi 7359.

1966 Moldenke, Monograph of Lippia 29
It should be noted here that the = turnerifolia of Hieronymus,

referred to in the synonymy above a synonym of Lantana aris-
tata var. angustifolia (Kuntze) oitaeke and that of Chodat is
Lippia tristis Briq., L. turnerifolia v amporum Griseb. is L.

sconpprooenapiaenedenas: © a aeteaeemmaees
contermina Bri Briq., and Le turneraefolia cual is L. tristis Briq.

Subsection Axilliflorae, Material has been misidentified and
distributed in herbaria under the names L. asperrima Cham., L,
trachyphylla Brigq., L. turnerifolia var, , angusta Kuntze, and L Lan-
tana sp. On the other hand, the Herb. Inst. Miguel Lillo 32176,
35569, & 37994, Herb. oe oi Argent, ee rate 32, Herb. Os-
ten 17237 & 22239, Job 1 3, JUrgensen 377 euntae son. [Jujuy,
Oct. 1892], Walme s.n. Tias Palnas almaso, “RO73/903 ], T. Meyer 262,
Monetti i 1238, Te T. Roig 15427, Rosengurtt B.3002, stitem

» Dec. 16, 1923) & s.n. [Cerro San Lorenzo, Oct. 8, 1 “T0251,
and Venter bee 21132, 2627, & 3614, distributed as L. turnerifo~
lia, are L. asperrima Cham.; Gallinal, Aragone, Bergalli, Cam
&R Rosengurtt 1616, Herb. Herter 50765, Herb, Osten 22703, Herter

‘sperrima var.

nen me a te nce a te

3OW48 a and Ar Archer & & Gehrt 91 ar gl are L. el rst Siahiies Herb.
Herter 8259), Herter r 989, Malme 2791, and Morong 575 are L. mo-
lia

rongii Kuntze; Kuntze sen, [Tunari, V.92] is L. trac

Regnell
1088 and Widgren 1088 are Lantana glaziovii Moldenke. The Jtns-
son 35a cited by me in Phytologia 12: 90 (1965) as L. asperrima
is” actually L. turnerifolia.

Schauer (1951) cites a Riedel s.n. from Rio Grande do Sul,
while Ragonese (1941) cites his no. 2808 from Santa Fé. Glaziou
(1911) cites his no. 21890, but this is actually Casselia glazi-
ovii (Briq. & Moldenke) Moldenke.

ee all, 5) herbarium specimens and 5 mounted photographs, in-
cluding type material of all the names involved, have been exam-
ined

= Eber BRAZIL: Paran&: Dombrowski, Saito, & Pereira 97/
771/290 (Ac); Dusén 2306 (Ja—3)918), s.n. “Tl duguacialyva, 24.10.
1910] (S), son. [Tamandua, 2h.11.1910] (S); H. M. Filho 50 [Herb.

Sp-~23037), s.n. [Vila Velha, Ponta Grossa, Nov. 3, 1928; Herb.
Inst. Bot. S. Paulo 233h6, in part] (N); Jtnsson 3a (Ca—50167h,
Ca~—533217, Mp, N, S, S, W—1481949); Stellfeld 26 (Mp--1395),
1039 [Herb. Mus. paren. *2019] (N); Tessmann s.n. [17-10-l,7; Herb.
Mus. Paran, 2508] (N). Rio Grande do Sul: do Sul: Rambo 21436 (Rb); Sel-

30 PHYTOLOGIA Vol. 13, no. 1

low s.n. [Brasilia, 17.9.1828; Macbride photos 3333] (Br—iso-
type, It--photo of type, Kr—photo of types ee =
of type, N—photo of isotype, Vt--isotype, Z--photo type).
Santa Catarina: Castellanos 21,72) (Ac); Dusén 17928 (3); “Smith &
Klein 11592 (W—22h9378), 11599 ( _(H—229 379) 5 aii Reitz, &
Caldato 9535 (Ok). S&o Paulo: Hoehne & Gehrt s san. [Ipe rs, Nov.
10, 1936] (Sp--36735). SORE Chuquisaca: earit 60 (B). PARA-
GUAY: Hassler 9312 (N); Osten 9181 (S, Ug); T. Rojas is 280a [Herb.
Osten 7909] (N, Ug), 377 [Herb. Osten 7901] (Ug), ll [Herb. Os-
ten ao (Ug). URUGUAY: Arechavaleta s.n. [vicinity of Salto,
Nov. 1898; Herb. Osten 3755] (Ug); Herb, Herter 83831 (N); A.
Lutz 1653 (Lz); Osten 0 (Ug), 1358 (Ug); Schrtéder s.n. [Paysan-
du, Nov. | et oat Osten 16350] (Ug). ARGENTINA: Corrientes:
Parodi 12 86 (N). Jujuy: Balls 5931 (W—1777760). Misiones:
Bertoni ers 3 (N)5 Ekman 1998 (S); Rosa Medina 55 (N, S); A. G.
Schulz 6983 (Sz). Salta: Hunziker 127 (W--180h110); Sleumer . &
Vervoorst st 2778 (W—-2173213). Tucum4n: Venturi 7359 (E—96238h) .

TURNERIFOLIA var, ANGUSTA Kuntze, Rev. Gen. Pl. 3 (2): 253

[as "P angusta"]. 1898.

Bibliography: Kuntze, Rev. Gen. Pl, 3 2 253. 1898; Briq.,

. Conserv, & Jard. Bot. Genav, 7-8: —— 190h; Briq. in
Ched. & Hassler, Bull. Herb. Boiss., pa : 1158. 190k; Briq.
in Chod. & Hassler, Pl. Hassler. 2 (11): or “pols Moldenke,
Known Geogr. Distrib. Verbenac., [ed. 1], 38, 9
Moldenke, Alph. List Cit. 1: 262 (agli) and 2: so, » 553 & 628.
1948; Moldenke, Known Geogr. Distrib. Verbenac., 2), 81, 99,
& 191, =) Moldenke, Phytologia * “87, 1950; Series Résumé
117 & 463. tego Moldenke, Phytologia 12: 29. 1965; Rambo, Pes-
quis. Bee 21: 28. 1965.

This alate me fers from the typical form of the species in
ae its leaf-blades 3-5 cm. long, but only 1 cm. or less in

dth

The type of the variety was collected by Carl Ernst Otto Kuntze
in southern Paraguay in September, 1892, and is deposited in the
Britton Herbarium at the New York Botanical al Garden. It is nota
well-marked variety. Briquet (190) in speaking of his L. moron-
gii says "Le L. Morongii ressemble beaucoup dans ses formes ré-
duites au L. Tammaeliotie ® angusta, quoique le plus souvent son
port plus robuste et ses feuilles moins rugueuses le fassent
facilement reconnaftre. Il s'en distingué tte pe agra par
ar ae ee de poils allongés et + —— » exclusivement remplacés
par ils strigueux ascendants et courts: en wetes; les
bractéoles extérieures des capitules sont largement ovées-acumin-

collected on September 9, 187L, with yellow flowers. The Hoehne
& Gehrt s.n, [Iperé, Nov. 10, 1936; Herb. Tt Bot. S. Paulo
36735], distributed as this variety, seems to represent the typi-
cal form of L. turnerifolia ins

1966 Moldenke, Monograph of Lippia 31

all, 3 herbarium snippit f, including the type, of this

caeaaee have been examined

Citations: PARAGUAY: Hassler 7 308 (Bm, N); Kuntze s.n. [Std
Paraguay, IX.92] (N-type).

LIPPIA TURNERIFOLIA var. SESSILIFOLIA Moldenke, Phytologia 3: 376,
nom. nud. 1950; Lloydia 13: 22h [as "turnerfolia var. ses-
silifolia"), Hare!

Bibliograpk graphy: Molde Phytologia 3: “dy 1950; Moldenke,
Lloydia 13: oe ag. Sections, Résumé 9), & 463. 1959; Rambo,
Pesquis. Bot Se

This vari sa are s from the typical form of the species in
having its leaves pce oblong-elliptic, 2-- cm. long, and 6—
9 mn. wide.

The type of the variety was collected by Padre Balduino Rambo
(no. 28180) in shrubby fields at Nonoai on the central Rio Uru-
guay, rande do Sul, Brazil, on March 3, 195, and is sae
in PE ge herbarive of the "colegio. Anchieta at P

© (1965) cites his collections as Herb, Anchieta "28180 & £ 281556
por Rio Grande do Sul. He describes the plant as a "Small shrub,
50 cm. high", growing on "Dry, more or less open or gate Campos"
— known to to him o only from the one locality, Nonoai on t ruguay
river. He gives the general distribution of the species, however,
as "From Parand to RGS; f araguay northern Argentina and
Uruguay". The taxon is known to me thus far only from = oo
collection, of which I have examined the type and ) p

Citations: BRAZIL: Rio Grande do Sul: Rambo 26180" igimmtate "of
bles % F—-photo of type, N--photo re type, Rb—type, Z—photo of

LIPPIA UKAMBENSIS Vatke, Linnaea 3: 528-529 [as "L.? ukambensis"].

Bibliography: Vatke, Linnaea 3: page Raping Jacks, in
Hook, f. & Jac cks., Ind. Kew. 2: 96. 189); in Engl., Pfl. Ost-
Afr. C: 338. al Je LES Baker in iitsett ayer, Fl. Trop. Afr. 5:
281. 1900; Moldenk Geogr. Distrib. Verbenac., [ed. 1], 50
. 96 (1942) and ae i, 118 & wags megs Moldenke, Résumé - 146,

& 463. 1927} ee ae Phytologia 12: 42, 106, 230, 307, 352, & Nera
(1965) and 12: 483. 1966.

Ss or ae 0.6--3 m. tall; stems pithy, branched; sap
colorless; branchlets hispid, villous upwards; leaves decussate-
pposite, stif

eolate, to 5 cm. long, crenate along the margins, attenuate at

the vere hispid on both surfaces, rough; heads solitary in the leaf-

axils, short—pedunculate, subglobose to finally oblong, about aus cm.
long; "bractlets oblong, longer than the corolla-tube, —
the apex; flowers with the odor of carnation (Dianthus c
L.); calyx green; cree’ hypocrateriform, white or P oeaectorel
to lemon~yellow with a darker center; anthers yellow,

he type of this species was collected by Johann Maria Hildebrandt
(no. 2739) at Kitui, Ukamba, Kenya. The flowers are described as

32 PEt Tt 0 G:0'o Ta ; Vol. 13, now. 1

"white" on A, Peter 12280 & 16706, "lemon-yellow with darker cen-
ter" on Tanner R.T.89, and "calyx green, corolla cream, anthers
yellow" on Drummond & Hemsley 22h5.

The species ies has been found growing in secondary bushland, in
grass on hillsides, and on grass steppes, at altitudes of 800 to
1880 meters, flowering in January, February, April, July, and Au-
gust. Material has been misidentified and distributed in herbaria
under the name L. asperifolia A. Rich.

The Greenway 3116 and A. Peter 112), distributed as L. ukam-
bensis, appear “appear to be L. schliebeni Moldenke. The G.F. Bs Elliot
6331 Seliection, judging from the photograph cited b below, | greatly
resembles L. schliebent:, It is possible that L. schliebeni and L.
ukambensis are actually conspecific. A. Peter 1715, 4,3360a, 143525,

& 16706 ha have very scabrous leaves, te his his 16807 has some of the
leaves scabrous and some not scabrous

In all, 12 herbarium specimens a 1 mounted photograph have
been examined by me

Citations: TANGANYIKA: Drummom& Hemsley 22))5 (B,

Peter 1682 [0. I.W1] (B), 1715 [0.1.42] (B) rato ir.3} hep
1,3360a [V.306] (B), 43525 [V.313] (B), 16087 {¥.139] (B,

46706 [V.283] (B); Tanner R.T 849 (Ca—182991). KENYA: P. Fe Se
Elliot 6331 (N-~photo).

LIPPIA UMBELLATA ssi Icon. 2: 75, pl. 19h. 1793.
My My 3 scabra Sessé & Moc. ex Moldenke, Prelim.
Alph. List Invalid N ic 29, in syn. 190. Lippia parviflora Ses-
sé & Moc. ex Moldenke, Prelim. Alph. List Invalid Names 32, 3m
syn. 1940 [not L « parviflora Prenat 1947]
Bibliography: | Cave; Icon. 2: 75, pl. 19h. gif S- ~~ in nike de

180; D. Die 1. 3: 599 Walp. td Bs Bete po
hs fers 181,53 Schau. in A. DC., Prodr. : S91. 187; Jacks. in
Hook. f. & Jacks., Ind. Kew. 2: 96. 189k; Briq. in Engl. & Prantl,

Nat. Pflanzenfam. 4 (3a): 151. 1895; Greenm., Field Mus. Publ. Bot.
re 341. 1912; Gandoger, Bull. Soc. Bot. France 65: 63. 1918; P. C.

pl.
1926; A. B. Seymour, Ind. Fungi N - 588. 1929; P. C. Standl.,
Field Mus. Publ. Bot. ae “woos. 1930; “Sisot, Ind. Lond. : 125.
1930; H. B. Davis, Life & Works Pringle 1936; P. C. Standl.,
Field Mus. Publ. Bo Bot. 18: 1010. 1938; clasiie, Alph, List Common
Names 11. 1939; Moldenke, Prelim, Alph. List Invalid Names 29 & 32.
190; Wors rsdell, Ind, Lond. Suppl. 2: 58. 191; Calderén & Standl.,
Fl, te 2a se 2, 238. nn ‘Moldenke, Known Geogr, Distrib. ver-

19 - Bol, 6: 2h. 19h85
Moldenke, Alph. List Cit, 2: 353, ele és, “170, io "339-543, 595»

1966 Moldenke, Monograph of Lippia 33

& 632 (1948), 3: 685, 769, 788, 829, 830, 833, 878, 885, 925,

926 (1949), and ): a 1075, “10%, 1180, & 1303. "1993 H. Bs &

A. L. Moldenke, Anal. Ins O: 10. 1949; Moldenke,
own Geogr. Distrib. vores, “ted ah: 31 & 191. 1949; Molden-

ke, Spec. Subsp. Cont. Mold. Set 45 p. [ 2). 1951; ont age In-

form. Mold. Set 46 Spec. 3 (1951), WB ‘Spec. (3 [3] ao ), and 49

ec. ;

o
©
&
Wh
B
HE
e |
ae
°
BS
* #
~
ER
@ mi
3
BB
ae

g
Terms Pharmacog. 128. 1955; Moldenke, Eta pty Set 51 Spec
3. 1956; Moldenke, Résumé 37, 307, 316, & 163. 1959; Moldenke, Ré-
sumé Suppl. 3: 9. 1962; N. i H. Krauss, as Bull. 18: 93.
196); Moldenke, Phytologia 12: 2h, 115, g. Aa, Wis, Ing,
168, Ais AT, 389. ee 223, 263, am 337, 62 & 363 (1965)

0
_ Illustrations: Cav., Icon. 2: pl. 19h. gtk Anon., Salvador
Minist. Instrucc. Publ, Fl. Salvad. 2: pl. 76. 192 “eer

Bush, loose shrub, or small weak round-headed tree, 1--12 mn.
tall, pleasantly aromatic with the odor of menthol; jean ¢ Sd

trag posi
fragrant, stidrtpattc sates patie scarcely 2.5 cm. long, tere te,
connate; eit nm es dark-green above piace 6 or whitish beneath,
narrow- late to ovate or broadly o 6—-20 cm. long, o
tuse to astenniats at the apex, acute to eon at the base, fine-
ly or coarsely crenate or Se along the margins, penninerved,
rugose, scabrous or scaberulous above, subtomentose or woolly=
tomentose beneath; inflorescences umbellate, in loose rounded
cymes to 15 cm. wide, the umbels many, axillary at the apex of the
branchlets; primary peduncles almost ), cm. long, with about 5 rays

to an acumen a globo

semibifid, the hemispheric segments persistent; corolla hypocrater-
iforn, varying from green- or greenish-yellow, yellow, pale-
yellow, deep-yellow, or rich-yellow to dull yellow-cream, fawn
white, sometimes yellowish and turning red, or deep-yellow with g
whitish eye, 2-lipped, the tube conic, as as lo ong as the calyx, the
throat open, the limb )-parted, the lobes rounded, subequal, rev-
olute; stamens ), short, 2 slightly longer than the others; fila-
ments capillary; anthers reddish, pip ovary ovate; style sin-
ple; stigma globose, subterminal, not emarginate, tomentose;

t a small capsule, glabrous, covered by the mature calyx-
Segments, 2-celled, each cell 1-seeded.

The type of this species was collected by Antonio José Cavanil-
les from cultivated material growing in the botanical garden at
Madrid, Spain, flowering and fruiting there at the end of December,
grown from seed brought from Mexico two years previous. Cavanilles
notes "Linnaeus de Lippia disserens calicem dixit superum, quem
Gaertner novissime inferum delineavit tabula 56 ubi Lippiae ameri-

3h PEITO0L.0G.2-A Vol. 13, no. 1

canae fructum descripsit, qui plantae nostrae fructui quadrat,
seminibus exceptis quae minime nuda sunt sed capsula biloculari
tenta. Linnae

nomine, quas Gaertner reduxit ad Daleam et Lippiam; ex Lippia
ovata illud, istud ex Americana. Huic =~ pists sai adiungo
Phrymae nowine in laudatum hortum introduc

It should be noted here that the L. spebegee of of Gardner,
mentioned in the synonymy above, is a synonym of L, gracilis
Schau. The Langlassé 7, cited below from Guerrero, may have
been stoma in Michoacén -- the locality given on the label is

reno

Sueeas (1847) places L. umbellata in his Section Rh al
Subsection A; Briquet ( 1895) classifies it in Subgenus Za a,
Section Rhodolippia. Walpers (1845) reduces L. eellicerenetotia
H.B.K. to this species, but this is entirely unjustified.

i ta has been found growing on hills and Pg

foothills and dry sunny hillsides, in rich loam soil, woods,
oak woods and Fenbiiba = » oak & pine or pine-oak-fir ieeatns pine &

ed.
of cultivated land, and on pine-oak-madrofio openly forested slopes,
at altitudes of £ 400 to 2850 meters, flowering from September —
March and in June, and fruiting from October to March. Calde
& Standley (91) describe it as "common" in El Salvador; MeVaugh
says of it in Jalisco "abundant here and at lower elevations",
“frequent on steep mountainsides in pine-oak forests (zone o. of.
broad-leaved trees in barranc ", and "occasional on steep slopes

Koelz found it on precipitous rocky and ravi
of gypsum and slate in or ae woodland with Juliana, Bursera,
Cephalocereus, Cryptoc » and Cassia,

Standl ey (192h 2h) and and Hocking g (1955) i the —_e used in
the trectaant of colic Oe s (196) notes that the

leaves are mined by the ste otis poubet pace s Guérin, but
probably he is referring here to L, hypoleia, since his no. 858,
identified by him as L. umbellata, is actually L. SO ng oe
mour (1929) reports that L. umbellata is attacked by

Puccinia elatipes Arth, & Holw. and Prospodium aigptae Somes )
Arth. [Puccinia lippiae Speg.]. Common names reported for it are

de monte", "nacare", "oreganillo", "oregano montes", "orégano

lilo®
however, is also applied to L. cardiostegia Benth. Le gaye
H.B.K., and L. ocephala Schlecht. & Cham
The flowers are described as "green-yellow or white" on Hinton
11910, “rich-yellow or fawn color" on Hinton 13550, "greenish- __
yellow" on Hinton.15706, "dull yellow-cream" on Balls 589, "pale-

yellow" on “ant foarte 12H, S banca on Hinton 1279 12791, Me’ McVaugh &
Koelz 1069,

1966 Moldenke, Monograph of Lippia 35

Standley (1930) reduces L. albicaulis Greenmm. and L.
Loes. to synonymy under this : species, the latter with a Par pl
He says "I have not seen the type of L. yucatana, collected at
rege Itz4 by Seler (no. 4918), and it may be referable rather

to L. myriocephala Schlecht. & Cham." In = 192) publication he
reduces L. albicaulis Greemm., L. chiapasensis Loes., L. pringlei
Briq., and L. substrigosa Turez. to L. caberiata, and says "The
numerous specimens at hand show considerable variation, especially
in pubescence and size of heads, but there do not appear to be any
well-marked dacatredag te by which’ it is possible to distinguish the
proposed segregates,

Material has been ‘hoarse and distributed in herbaria

under the names L. callicarpaefolia H.B.K., L. cacarpeer
Kunth, L. durangensis Moldenke, and L. involucrata Sessé & Moc,

BOOK REVIEW
Alma L. Moldenke
"The Chemistry of Plant Processes", by C. P. Whittinghan,
llustr. Philosophical Library, Ince, “New York 16, a gg
$7.50
The author's words of intention -- "to introduce students who

plant metabolism" —- c t study of this
concise book. The firs deals in a quite effective and up-
a guia with cellular metabolism — structure, function,

enzymes, fermentation, Ds etelaaie biological syntheses, photo-
synthesis, giyoiee n metabolism. The second half deals in a more
dated manner with plant processes -- osmosis, water relations,
translocation, growth.

At the end of chapters there are short, pertinent biblio-
graphies. At the end of the book there is a well selected gener—
al bibliography which should be useful to students because it is
not indiscriminately and discouragingly 1 The work is
marred unnecessarily by a few typographic errors » as in
spelling of Nicotiana tabacum on Plate 3 and in names of research-
ers in the bibliography, and a few errors of content, as
statement that only far-red absorbing phytochrome causes floral
induction. The inclusion of generic names would have been
helpful in the index.

s book would serve as a good text for the topic in a gen-

a beta course or as a basic one augmented by more detailed
and more modern material in a plant physiology course.

NOTES ON BRAZILIAN CYPERACEAE — I
Liens Teixeira Eiten
Instituto de Botanica, Sao Paulo

1. Cyperus pelophilus Ridley

Among specimens of Cyperus sent to me for determination by
the Instituto Agronomico do Norte, Recife, were encomtered three

TrO~

th collection numbers. These three, oe another speci.

fron

ara), seemed to be all the same species. In KHkenthal's mono~
's sul

ct. Sulcatae from the other sections in the subgenus, is nei-

Under sect. Sulcatae the material fell into & rivularis Kunth,
the only possibility in the section. This species is cited for

S described as obovate, compressed-biconvex and short-
ly apiculate. (I had no material at hand of Cc. rivularis for
examination

the other sections, the four collections fell direct-
ly into sect, tachyi. Three species of this section
the material at hand, are described as having sulcate achenes, C.
atribulbus Kik., C. pelophilus Ridley and C, sulsinux C.B. Clarke.

Southern Africa; it is described as perennial, the culms with
bases. The material at hand is anmal, the base of the
Culms is not bulbous, and, in general, the spikelets are mich
longer. From ¢, Sulsimx the material differs in fundamental
characters, but exactly fits C. pelophilus in these same charac-
ters, as shown in Table I.
Alth the other species in the section did not fit the
material, a special effort was made to see whether it was not
a form of C, Rolystachyos Rottb., a very polymorphic species
th non-sulcate achenes, occurring in both the Old and New
World Tropics. Only the varieties + and
(Torr.) Fern. (var, ¢ Boeck.) were considered, since
the other varieties recognized by Kikenthal seem to be merely
variations of these two. Table II compares the material with
36

1966 Eiten, Brazilian Cyperaceae

and with the two varieties of polystachyos in those

pelophilus.
characters that most distinguish these taxa.

Table I

Character Cc. sulsinux* material
number of always 1 always 2
stamens
number of 3 5
nerves on
‘lume
anther linear short-oblong
shape
number of 34, 6, &7, 4-6, 5
involucral

cts
achene oblong or obovate, apex
shape oblong-elliptic, truncate,

apex acute, apiculate

apiculate

short-oblong

5-6

t
apiculate

37

obovate, apex
runcate,

* I was not able to see any material of this species; the hes

mation was taken from the text and illustrations in Kikent:
(1936) and Kern (1954).
Table II
Character Material pelophilus polystachyos
var. var.
polystachyos.
underground fibrous fibrous rhizomes (anmmal )
organs roots roots (perennial)
(ammal) (annual)
achene sulcate sulcate non-suleate non-sulcate
face
achene obovate, obovate, oblong or obovate, apex
shape apex apex subovate— truncate or
truncate truncate oblong, apex rounded-
subtrumcate truncate
&chene 1.1 -1.3 not given 0.75-1.0 (0.8) - 1.0
length mm mm im
glune 1.6-1.75 not given 1.5 - 2.0 1.5 = 2.3
length 1.5-1.8
mm 1.75
a.
for the 4
collections

1.5-1.8

38 PHYTOLOGIA Vol. 13, no. 1

The material differs from polystachyos particularly in
characters of the achene, an organ of highest importance in the
ee It is concluded that the material is C. .

Since C. pelophilus is cited only for Africa and Madeges-
car, this represents, so far as I know, the first citation of
this species for the New World. It is not possible eaPisae tell if
the species is —— or introduced into northeastern Brazil.
The Pernambuco pecimens are weeds, but no habitat is given for
the Ceara stechaen. omits was collected more than 50 years ago.

2. Cyperus capillifolius A. Richerd

Examining three collections from the Rio de Janeiro Botani=
cal Garden, G. Black 2162 B, L. O. William & S. V. Assiz 6799,
and I concluded that mae were C. capillifolius.

by M. Barros in 1955,

These three collections are neither C, rivularis Kunth var

nor C. Brae tte (Steud.) Kik.; they differ in

several importan t characters, especially in the bracts, glumes
nd achenes. Not ha al at hand of C. or
other well-identified material of os the comperi-
SON Wes pony Bos the d Bees te and illustrations of these
species by Kikenthal (1936), Fernald (1950), Mah lanteeh
(1960), ae (1352) and Barros (1947). The following t
hows the compa

Table III
Character Tivularis material ¢.
var. var. a = Brade 17597 ius.
lagunetto b = Black 2162 B
= &
Assiz 6799
number of (2) 34% OO} 23 a 2 2
bracts be 2
C= 2
glume 2 o0~2.5 2.1% am 1.25 1.25
Imm Cm 1,25
achene 1.0-1.5 1.0-1.1 ap 0.75 0. 65*#
length be 0.70-0.90

1966 Eiten, Brazilian Cyperaceae
Table III (contimed)

Character rivularis rivlaris material § capili-
folius

* Calculated from drawing a Barros (1947).

*# Calculated from drawing and from glume length given in text

in Ktkkenthal (1936). The fa.» given in Kilkenthal's genes

does not correspond to the amplification actually us

ney To nt the achene profiles more easily, all nl drawn
the s

The material evidently conforms O papenli tol ius and not
to The two varieties of the atter species have

achense Glin soundad or with shortascumiziete apices,
achenes of and of the three gol lectdons at hand

from an African specimen illustrated by Kifken-
thal, does not show this depression. If the depression really
is lacking in in all Africen material and occurs in all Americen

39

Fe) PHYTOLOGIA Vol. 13, no. 1

material, it could at most be a basis for separating
dota th — in all other characteristics tha bias Seca from
both continents re essentially equal,

pe the literature, Cyperus rivularis var. riy-
ularis is restrleted to North America, while the subspocies

tg occu re and also in Central America and the
Andine repick ae South America.
tha

rid, Risde) from
Serra da Lapa, Brazil. According to Riedel 's itinerary giv-
en in the Flor ate Mebiiiasic. this cot fers to the Serra da
Lapa near Belo Horizonte, Minas Gergis, not the moumtain
range of the same nam near Bom Jesus da Lapa in Bahia,

Minas and all grow at high altitudes:

George Black 2162 B, 26 Feb 1944, Catas Altas. The
city of Catas Altas, 60 ion ESE of Belo Horizonte, lies at
an altitude of about 300 meters, but the mountainous coun-
try around it reaches 2000 meters

Louis &S. VY. tests 6799, 5 5 May meet
Serra de Monjolo, 18 km WNW of ay of Serro, Municipio de

Tro. The map gives an altitude of 1063 maters along a
Toad near the mountain range of tie at de Monjolo, but the
rangs itself reaches mch greate r heights.
oh Brade 17597 & A, Barbosa, Po April 1945, S4o Sebasti-
Go do Parafso, Morro do Bau. paretein, » Morro do Bau.

me

The fact that the Riedel collection etal by Kikenthal
and these three further collections all come from the same
general region and have not come to light anywhere else in
Perse so far makes it probable that the species is native

this region, If it were an introduced weed, o
se Ra it rather scattered around larger areas of the
country in semi-dry mountainons areas, Unfortunate no

oe it would be a further example of the floristic we.
onship between Africa and Central Brazil.

Bibliography
Barros, M. 1947. Cyperaceas, in Descole, Genera et species
antarum argentinarum. Buenos Aires.
Fernald, M. L. 1950, Gray's Manual o Botany. New York.
Gleason, : sag 1952. Britton & Roi Tllustrated Flora.
ew

Kern, J. A. 1954. Reinwardtia 3: 4
Kikenthal, G, 1936, Pflanzenreich Iv-20,
Mohlenbrock, R. H, 1960, Amer, Midland tat 4g Sah:

1966 Eiten, Brazilian raceae

Fig. 1. Habit of Cyperus pelophilus Ridley (Loefgren 860,

Cearé), X 1/2,
Fig. 2. Cyperus pelophilus Ridley (Loefgren 860).
A. Spikelet, X 10.
B. Rachilla of lower part of spikelet, X 25.
C, Flexnous, winged rachilla of upper part of
spikelet, X 25.
D. Glume containing achene in natural position,
E. Upper part of a glumes showing micro, X 40.
Fe Flower, X 65,

X 25.

G. Mature achene, X 40, showing one of the sulcate

faces.

H, Transverse section of achene at the level indicated
by the dashed line in G, showing the two sulcate
faces. The depression is separated from the rest

of the face by a sharp border.

PHYTOLOGIA Vol. 13, no. 1

FIG.1

DISTRIBUTION OF JUNCUS IN ILLINOIS

%
R. A. DeFilipps

This article is — ed use as a supplement to A Taxon=
omic Study of Juncus in Illinois. Am. Midl. Nat. 71 (2): 296-
319. 196. I based that — cy oe in 1961-1962) on par pee
from the herbaria of the following institutions: Chicago Museum
of Natural History (F), Illinois State Maseum, Springfield om

Illinois Natural History Survey, Urbana (ILLS is}, U niversity of
Illinois, Urbana (ILL), 1 University of Tilipois, ise o (CHT),
Southern Illinois University, Carbondale (SIU),

In October-December 1965, the above herbaria were searched
for additional material, and the collections of the Missouri

of all the herbaria for the opportunity to see their collec~
tions.

During the course of these studies, specimens were encou
tered which sometimes alter the distetbetion of Juncus in 7
inois as shown on the maps included by G.N.Jones & Geb.
in Vascular Plants of Illinois 1955, and ce i as,

— eee, Soe, SAINOLS

1. Juncus secundus Beauv., in Poiret, Encycl., Method. Bot.
Suppl. 3: i20- 1813. (Map 1)

aor ples Specimens: GALLATIN CO.: SW, of Gibsonia,
a 33509 9 (ILLS), HARDIN CO,: E. of Lamb, Evers 1150 (ILLS) «
ON CO.: Fern yet Bollwinkel & Windler FC37 ). POPE

Cont "1 mis SW, of crag g aA E SATNE CO,: O
Indian Stonefort, Bailey + rains 166E I (StU). UNION CO,: Pan-

ther's Den, MacMahon ges ¢

2. Juncus dudleyi Wieg. in Bull. Torrey Club 27: 52h. 1900.

“(ilap 2

Representative oe ALEXANDER CO,: Cache, Cran-
will s.n. (ILL). BOONE CO.: E, of Irene, Fell 5781 Oe .
CARROLL CO.: oo sepia Wunderlin 2099(WIU ).
CO.: Bear McDonald s.n. (ILL), CHAMPAIGN CO.: aang
Jones 17902 “aby CLAY CO.: Larkingsburg Township, Evers
23815 (ILLS), COLES CO.: 31 mi. E. of Charleston, Evers 3502

TLLS). COOK CO.: Morton Grove, Glassman 139 (CHT). DE KALB
CO.: We of ea Ever's 33682 THs). Pace CO.: We of

— ° ’ ern nois Univ., e

1966 DeFilipps, Juncus in Illinois 5

GRUNDY 0O.: S. of Morris, Evers ae (ILLS), HENDERSON CO.:

Pee moat Patterson Sone CO.: Geneseo, Dobbs s.n.
(ILLS). es CO.: Fountain ay S. of Gorham, Evers "36430

(ILLS). TASP ER CO.: W. of sie Ozment & DeFilipps 1736A _
(SIU), JO DAVIESS CO.: ) m. N. ee “Evers 30027 (ILLS).
KANE CO.: N. of Elgin, Steyerma te 68356 (F). KANKAKES CO.: SE,
St. Anne, Fuller 8931, (ISH). RNCK CD. : 1 mi. S. of Maquon,
Evers 221) (ILES), LAKE CO.: Wauconda, Evers 556 (ILLS),
LIVINGSTON CO.: 3 mi. W. of ares Fuller 6537 (ISM). t
2 mi. E. of Kingdom, Keit TIS). LOGAN 0O.: Near Lincoln
Fuller S76y rah "uG BONO CO. We of Macomb Myers 52h (ISM).
- aT eee eros Vase oat (ILL). i.) oe ~ ie
» Evers “198 ie lS) ON CO.: oH. 0 of ese
23786 (ILLS). CO.: Se of Oakford, Rexroat 839 (r es
MERCER CO.: 9 mi. or of New Boston, Evers 2 24311 (ILLS). MORGAN
CO.: 8 mi. SW. of Arenzville, Rexroat 2737 fai). OGLE CO.:Kill-
buck reeks ye 369 (ISM), PLATT CO.: E. of Lodge, Evers
38% (ILLS), SCOTT CO.: Near Oxville, Rexroat 3738 (ISM).
ST. CLAIR Bs Cahokia, collector unknown (MO). STARK CO.: Near
Nady Petra, Chase 256 (IIL). TAZEWELL [ELL CO.: Spring Lake, Chase
ie VERMILION CO.: E. of Muncie, Evers 38610 (ILLS).
WABASH Aa Near Patton, Schneck sen. (ILL). WILL CO.: Romeo,
steyermar k 6586 (F). WILLIAMSON CO.: Devil's Kitchen Lake,
eo 8 3 (SIU). WINNEBAGO CO.: Rockford, Fuller *&
fae 10 TU). WOODFORD CO.: S. of Spring Bay, Chase 9692

LEE CO.:

3. Juncus interior Wieg. in Bull. Torrey Club 27: 516.

presentative Specimens: ADAMS 0O.: S. of Lima, Evers
38377 (THiS) BOND CO.: SW. of Mulberry Grove, seb 23882
WN CO.: E. of Siloam Springs State Par Evers
35216 (ILLS), CALHOUN 00.: Brussels, Muller eo (ETS), css
ot eee (ISM). “cmegy oiea chai oo ma isher,

Evers 39 TLLS), CHRISTIAN C Thcin hon Ake a
CIA cee of Marshall, 8 Sees ‘ase “a Beton a

kingsburg, Evers 3719) (ILLS), CLINTON 00.: N. of are
Byers 18017 Je S Oss : SW. gi Mattoon, Evers 38166
{HILS) COOK CO.: Skokie asa Glencoe, Sherff sen. (F).
CRAWFORD CO.: Robi seein Tehon een ILLS). RLAND CO.:
N. of Neoga, Evers 38216 Rpt E WITT CO.: NE. of nO
Evers 60100 (ILLS). DOUGLAS CO.: * of Galton, Evers 309
ULES). Do eae E CO.: We “at Wheaton, Moffatt 769 (ILL). FAY-
BITE 00.3 Ne of Farina, Ever s 34120 (ILLS). FORD CO.: Ne of
Sibley, Evers eri (TuS}-F FRANKLIN CO.: S. of Benton, Evers
65231 (ILLS). FULTON CO.: N. of Anderson Lake, Rexroat 1,919
SM). GALLATIN 0O.: Patonar 4 sen. (ILLS), GREENE CO.:
; mis Ne of Eldred, Evers 3327) (ILLS). HAMILTON CO.: S. of
n, Evers 37359 (ILLS). HANCOCK CO.: E, of Warsaw, Evers
3626. (ILLS), HARDIN CO.: N. of Elizabethtown, Evers 30376
(ILS). HENDERSON CO.: Oquawka, Patterson sen. (F). HENRY CO.:

L6 PHYTOLOGIA Vol. 13, no.

3-1 mie N. of Annawan, Dobbs son. (F). IROQUOIS CO.: 2 mi. E, of
Watseka, ace 71127 (ILLS), cgay eer Finney, Mohlenbrock

5306 (SIU). JASPER CO.: Berto n, veo . JEFFERSON COe:
SE. of Opdyke, Evers 37326 (ILL sy “JERSEY C O.: We. of Chautauqua,
Evers 2504) (I Peale JOHNSON CO.: peeppare Bluff, Bailey &

2166 ( » KANKA e: Kankakee, SeDe (F)> KNOX Csr
Near Sera Chase 1859 (ILL) < + SALLE CO.: Starved Rock
State Park, Ries & Werner sen. (ILLS), LAWRENCE CO.: 2 mi. E. o
Lawrenceville, Sivert sen. (ILL), LEE CO.: Green River Area, W-
kins & Anderson s.ne ). LIVINGSTON CO.: Dwight, Evers 37935
TIEs je 2 cecal O.: C. & D. Heitmann 87h (ISM), MC DONOUGH CO.:

SE. andon, Evers 13822 (ILLS), MC HENRY CO.: NE, of Harvard,
Piel ee (ILLS). MACON CO.: Elm Ridge, Cloke asa (F), MAC
OUPIN CO.: E. of Car ma Evers 3639 (TLLS), MADI ae CO.:
Godfrey, "Winterrin er 17048 (ISM). MARION CO.: SW. of K mundy
Evers 37269 (IIDS). iaSoW Co W. of f snence he Rexroat 1,928 (ISM).
MENARD Athens, Hall s.n. (F). : N. of Keithsburg,
Evers 2279 g (ris). MONROE CO.: SE. mre erloo, Evers 31,002
{IIs )- ~ “Oy? of Nokomis, teers 65898 eas MOUL=
TRIE CO,: is Oe ai ae “Evers 12862 (TEES). OGIE CO.: N. o
Davis ee Fell 53495 (ISM). PEORIA Bs xexice, Chase 13469
(SIU). PERRY CO.: Tamaroa, Stout sen. (F). PIATT 0O.: 2 mie W. of
Nant Fs Evers 53908 (ILLS). PIKE CO.: Near Barry, Campbell &
Alexopoulos Sane ILLS). PULASKI CO.: S. of Ullin, Evers 33152
TNAM

ILLS). PU CO.: NW. of eee Evers 306)\2 (ILES). RANDOLPH
CO.: N. of Evansville, aes h (I CHLAND

co

Township, Rid, 300 (F), R Helin COs Near Silvis, Fell &
Fell F #us-a51, CT) ST. cfaTR Cons NW. of Marissa, Boewe Seno
(ILLS N

q

of tee wits State Park, Bell & Br
yee a Mt. Carme . 2 Schneck ee ve :
° vers 1179 Les W N COs: E, of Ven-
edys Evers 1790 ” (TELS). “WAYNE CO,: of i City, Evers

90 (ILLS 11995" CO.: Ne of Bene. Ahles 1,671 (IIT).
— CO.: 2 mi. W. of Erie, Fell & Fell » WIL
TAMSON CO.: S, of Cambria, Evers BM ae _ (TEES). WINNEBAGO CO.:
Near South Beloit, Fuller 13773

he Juncus temis Wi

lid. Sp. Pl. 2: 214. 1799. (Map h)
eS ars - edeengg ADAMS C

: Quincy, Brinker 1201
(isu). ALEKANDER C obee i Cranwill s gut CL). BOND CO.: Ne
amalco, Evers Si TLLS sed Ly f Belvidere Fell
Suess (1 su}. BROT CO. i

o3 3 mi oP Cie aS teat (IL).
REAU CO.: E, o kilwa, Ska a 77 '‘ALHOUN CO.: So
of Hamburg y Trice | 972 (ILLS), C, O.: ge Rg all Palisades

1966 DeFilipps, Juncus in Illinois 7

State Park, mare 29900 Sige he CASS CO.:N, of Virginia, Rexroat

3159 (ISM). ATGN CO.: Near Urbana, Jones Jones 13662 (ILL). CHRIS~
TIAN CO.: iaytore tis Said 2217 (ILL). CLARK CO.: Lincoln

Trail Stat Ozment & De s ee (51). clay CO.: N. of

Louieva Lie} teens Lae 1377 01377 (TL ae

— (ILLS we Re piel go 7: SW, of Big Bers "38186 (ILL

COOK Co. n,; pee: 1373 (F). CO.: S. of panne?
ites ivers "6798 CHES, isrLand co, CO.: ile el Dell Ar rea, Spoo-
ner sone (ILLS). DE KALB CO.: Ne airdale, Fell 53153 (1s)
DE WITT CO.: NW. of Kenney, vere oF 90 js Lis). DOUGLAS -

of oe Evers 38119 (ILLS). DU PAGE CO.: Wheaton, "dice 1681
a DGAR CO.: SE. of er Evers sua (ILLS). EDWARDS CO.:
ra et Evers )1,866 (ILTS). EFFINGHAM CO, . of on
Beane “Bees (ILLS). FAYETTE CO.: N. of Laclede, Evere 13030 aS,
: N. of Zeigler, Evers 756 (ILL ILS}, “FULTON CO.

of Banner, fvene 17020 (ILLS). Bet one Pounds Hollow akg:
pc re AB 30315 a LLS). GREENE COs: Barrow, Evers 11947 (iis).

Os: of Mazon, Evers 17365 “(r LLS), HAMILTON CO.: Ss. of
ii eee. er 11123 (ISM). HANCOCK CO.: Carthage, Kirchherr
100 (ISM). HARDIN CO.: Elizabethtown, Evers 23538 (ILLS), HEN=
DERSON Co.: Oquawka, Patterson Son. we HENRY CO.: Geneseo

of Newto Opdyke ,
Evers 34156 (11LS). JERSEY CO.: Near Otterville, Winterringer

25203 Tish
J C

15292 (ISM). JO DAVIESS CO.: S. of Rice, Evers
SO

CO.: E, of Dahinda, tne 3966 (ri
tag ‘Grahaa9 9391 (ILL), LA SALL LLE CO.: Starved mak State
k, Werner s.n. (ILLS), LAWRENCE CO.: Birds, Boewe s.n. (ILLS).
un GOs Amboy Long 2h ate Specie CO.: 3 m. pa of
Rowe, Fuller 8537 a TEST. : of Lake Fork
76233 pee So onctoe ob : 2 Macon, Myers 309 ( ISM). TE Ry

. “S eheat Nason fat" ; oe sear 2130 erie:
vers 6879 ( ie ON ah Elm rem, oke

MACOUPIN CO.: hipman, Evers 308 in (TILES

ON CO,: “Codie, gfe er 6 Bee aged =e : ‘Salen,
Bebb s. e3

ipps TIO3A (SIU). MASON CO,: ES ae To pokes » ales pikes 21 a5 Tit.

MASSAC CO.: E. of Joppa, Evers ueTOe (ILL > Rex-
roat 3753 (ISM), MERCER CO.: W. of Eliza, gix Ege ” UIs),
MONROE CO.: 1.mi. S. of Fults, Ouiene SoM. '» MONTGOMERY

llivan, 0
gon, Evers 37612 (ILD os
8949 (ISM). PERRY aoe Se of Pyatts, Sears 1872
eas. CO.: Near Monticello, Jones 178 800 (TLL. cn coe Pitt=
ld, Boewe sen. (ILLS). POPE CO.: Ne of Herod, Ahles 2726
(tin) PULASKI CO.: 1 mi. Ne of America, Evers 11522 (ILLS).

48 PRY T040:G.2-a Vol. 13, no. 1

PUTNAM CO.: mi. E. of Henry, Cament. & DeFilipps 1181 (SIU).
RANDOLPH CO.: Near Modoc, a cak 216R ani PcuTAND CO.: Ol-
ney y Rigg ay 120 (F), ROCK ISLAND CO.: Port Byron, Fuller 13788
oie LAIR oh Dutch sania, Weill “858 (ISM). SALINE CO.:
1 mi. om, “ot Carel r Mills, Evers 2769 (ILLS). hel CO.:
Salisbury T enstics Fuller 7943 (ITLS). SCHUYLER CO.: NE. Be Lit-
tleton, ayers 3 Tits). ScOTT CO.:.2 mi. N. of goa! r,
TSM). SHELBY CO.: Sigel, Evers 21088 (TELS), STARK
CO.: 2: = NW. of Wady Petra, Chase 64 (F). TAZEWELL CO.: N.
of East Peoria, Evers 39457 (ILTS). UNION Co.: = of Larue, Evers
23935 (ILLS), VERMILION Be, — of Oakwood, Evers 38646 (I IELS)e
WABASH CO.: Bellmont, Boewe sen. (ILLS). WARREN CO.: Ne of Mon-
mouth, Ozment & DeFilipps 1060 (SIU)« WASHINGTON CO.: Vene
Evers 33949 (ILLS). WAYNE CO.: E. of Sims, Evers }),286 (ILLS).
WHITES CO.: Dogtown Hills, S. of Maunie, Evers L876 (ILLS). WHITE=
SIDE CO.: E. of Fulton, Ozment & DeFilipps 517 (SIU). StS ce
Romeo, Umbach sene (F). WILLIAMSON CO.: = of Cartervi
1862), (ILLS). WINNEBAGO CO.: Sa ae, Fell & Fell 7B=360 TLL e.
WOODFORD CO.: S. of Spring Bay, Chase 9006 (ILL

5. Juncus vaseyi Engelm. in Trans. Acad. St. Louis 2: 48.1866.
(Map

Pigg oe ome Specimens: COOK CO.: Cleaver's Woods, Bebb
942 (ILL). MC HENRY CO.: Near Ringwood, Vasey s.n. (MO). WINNEDA-
GOCO.: 2 mi. We of South Beloit, Fell & Fell 17176 (ILL).

6. a eenei Oakes & Tuckerm. in Tuckerm. in Am. Jour.

¢ 45: 37-6 1843. (Nap 6)

entative Bag hae: grt ya CO.: Washington Heights,
Bebb ou? (F). _zROgiOT S CO.: $ mi. N. of Pittwood, Evers 19032
{IEEs). KANKA : Kankakee, Seti 80 (ILL), LAKE 00.: Tilinois
Beach State Pats en 202 (I LLY. DEE CO.: Green River Area, Bell~

rose sen. (ILLS : 2 mi. SE. of Custer Park, Steyermark
@UB5u (F). WINNEBAGO CO.: Near Shirland, Fell 53870 sn

7. Juncus gerardi Loisel. in Jour. Bot. Desvaux 2: 28. 1809.
(lap 7

Representative a COOK CO.: Chicago, 92nd. & Stony
Island, Glassman 3788 (CHI).

8. Juncus bufonius L. Sp. Pl. 328. 1753. var. bufonius (Map 8a)

Presentative Specimens: CARROLL “ee Ne. of Savanna, Ahles
4336 ae CHAMPAIGN CO,: Urbana, Ahles 6326 (ILL). COOK Otic
ees Vasey sen. (F). DU PAGE CO.: We of Wheaton, Moffatt 168
(ILL). HENDERSON CO.: Near Biggsville, Patterson Soe
a E. of Beach, Gates 2782 @). MC HENRY CO.: Ringwood, ase s
Sen. (ILL). WABASH COs: Shearer sen. (ILL). WASHINGTON CO.: Irv=
ington, French s.n. (SIU), WILL 00.: Drummond, Swink 6-A (SIU).

1966 DeFilipps, Juncus in Illinois hg
Juncus bufonius L. var. congestus Wahlb. Flora Gothoburgensis

38 1820. (Map 8b)

ms Vie’ Specimen: COOK COQ.: South Chicago, Hill 25.
ALLL

twee

De Juncus effusus L. Sp. Pl. 326. 1753 var. solutus Fern. & Wieg.
n Rhodora 12: ae 1910, (Map 9)

pian ene Specimens: ALEXANDER CO.: SE, of Olive Branch,
avers 44486 (ILLS). po ieee CO: Taylorville, Suciewes Son. (ILL).

eee SeNe (ILL). FRANKLI xe : Near Christonber, Jones i
Lig) s HAMTLTO TON CO.: 2 me E. = _Fiopolis, Evers 503 (IL

2.
peter ¢ belt “kt Alexopoulos Sene (ILLS), JOHNSON CO.: Near
Ter 165 SAL

Vienna, Fuller 6 (ISM). LE CO.: Starved Rock State Park,
Lans )o LAWRENCE CO.: 1 mi. SE. neeville, Siv-
ert Sone ). LEE CO,: Amboy, Long 762 (ILL). MARION CO.: SW.
of Alma, Ahles 5377 (ILL). MASSAC CO.: NW. of sticss ae aynse

(ILLS), Ferg CO.: Ne of ee Bailey & Sw

PERRY CO.: NW. of Bepine Nao peers oto TTS) eo

Lake Glendale, Bailey & SIU). PULASKI CO.: Ne eg

Olmstead, Ahles 2500 (T GOR TSLAND oe : Near Silvis, Chase

16999 (ILL). SALINE CO.: 1 mi. SW. sag Carrier Mills, Evers 2808

ee): UNION CO.: Pine Hills, Ha 9 (SW). WASHINGTON CO
Med oe pemnch Arenoh 82. (SIU). WHIT or a of Norris yy bee

23900 s). WIDLTAMSON CO.: SW. of C rville, Evers
oe GO CO.: 2 mi. W. of South Beloit; Fell & Fell Fl7-
160 (ru). ee

10. Juncus balticus scare n Mag. Ges. Naturf. Freunde Berlin
3: 298. 1809. - dittoralis Engelm. in Trans. Acad. St.
Louis 2: 2. 1866. LO

Representative Specimens: COOK CO.: Chicago, Vas
(ILL). JO DAVIESS CO.: E. of Woodbine, Ahles )15 (ILis aE
CO.: Waukegan, Fuller 7768 (ISM). MC HENRY CO.: A Aigonquin,
Nason Sene (ILL).

ik Juncus biflorus Ell. . S,. C. and Ga. 1:07. 1817.

oF erie les tha

e au (tuts), oh Specimens: Aah a 3 : 1 mi cas aa
vers 39318 (I CLAY 0O.: Greendale, Evers °
CLINTO f Posey, Evers 3 31326 (I 50058 ( CO.: We 0
“ened aa ‘8h, (ILL). CUMBERLAND CO.: 1 mie S. of Neoga,

Winterringer 69h, ILL). EDWARDS CO.: 2 mi. W. of Samsville, Evers
70577 (TUES), EF EFFINGHAM CO.: NE. of Mason, et 3082 (ILLS).

ae arent

50 PAY TOLOGIA Vol. 13, no. 1

FAYETTE CO.: E, of Vandalia, O'Dell 543 (ILL). FRANKLIN 00.: S.
of Benton, Evers 18615 (ILLS). GALLATIN 00.: Rt. iS at buates Rde,y
Ozment & DeFili Uyil (S10), HAMILTON ©O.: S. of Dale, Winter-
ringer 11]22 (ISM). HARDIN CO.: Near Lamb, Buser 2856 (ILLS

: eo of Su
Salem, Ahles 205 rn): PERRY CO.? Se of Sunt Sunfield, "Byers 30565
(ILLS},- POPE Belle Smith Springs, ol t 731 AG df Reka
CO: We of “Pulasit, Mohlenbrock & oa jag ta LAND CO.:

4 mie SW. of Olney, Scherer & Shaw 396 set TSM). pt ate Oe
Brendel pies el SALINE CO.: W. of Harrisburg, Evers s 17523

CO.: Sigel, Evers 3407) (SIU). UNION CO.: French
2975 (SIU). eg CO.: S. of Keensburg, Ahles 1656 (ILL).
WASHINGTON CO.: Irvington, French sen. (SIU). WAYNE CO.: E.o
Sims, Evers 681 (ILLS), WHITE COv: S. of Norris City, Evers
17785 ELE oe ce WILLIAMSON ©0.: Crab Orchard Lake, Sands & Garr-
ison 33

s biflorus Ell. f. adims Fern. & Grisc., in Rhodora
(Map 11b)

Juncu
312 157s 19456

Representative Bee Saees: ALEXANDER CO.: McClure, Cranwill
CLARK Ci

Sens (ILL). : 3 mi. N. of Marshall, teen er Iho?
(ISM). WABASH Co,: pra Mt. Carmel, Schnec Schneck sn, TILL),

12. Juncus marginatus Rostk. Monog. Junc. 38,pl. 2, f.3. 1801.
=e 13 , ’

Representative Specimens: ADAMS CO.: Burton Cave » Evers
Jones, & Jones 27h (ILL), CASS CO.: Beards stown, enks 3919 g ).
COOK Cov: “Thorn ton, Fuller 1699 (ISU). Pores CO.: N. of Thom—
eee bese re 4193 CTLLS). GALLA —) S. of Kedron, Evers
5, 26753 (I LLS). CO: Augusta SeNo ns TROQUOIS Ce?
Near Be eceretiia, ans 18817 (113. SSNS 05 : E. of Vienna,
euterns er 8998 (ISM). KANKAKEE CO.: Near Ste ‘kane’, Jones
il ~ LAWRENCE CO.: E. of Billett, Byers pee (ILLS ak
LEE to's Amboy, Long 598 (ILL), ig = f Manito, Chase
5260 (ILL). MENARD Go= Athens, "@. MORGAN CO. Tr
of Jacksonville, Rexroat 1,058 (usmy, REGALAND HLAND CO.: 33 mic No of

Ellery, Boewe sen. (ILLS), WHITE (O.: Norris City, Ahles 1676
(ILL), WILL 0.: 2 mi. SE. of Custer Park, Steyermark 602

(F). WILLIAMSON 00,: Devil's Kitchen Lake, Stookey & Fore 517
(SIU). WINNEBAGO ©O.: Campbell Bog, Fell 51895 (TSM. ae

13. Juncus nodosus L, Sp. Pl, ed. 2: 1,66. 1762, (Map 13)

Representative Specimens: COOK CO.: Chica g 688
CO.: go pansin
DU PAGE 0O.: Bense enville, Bennett s.n. (ILLS), KANE - oF

1966 DeFilipps, Juncus in Illinois 51

Elgin, Steyermark 68353 Sa KANKAKEE (O.: Rock gr Hill 317.=-
1873 se IDL} TARE 00 E. of ee aya 167 (ILL). LEE We: a
boy, Long 798 (ILL), MC HENRY ~ Ringwood, ase pr (a.
OGLE CO.: Byron, Blount 196 cs PEOATA 00.7 eoria, Brendel
SNe (ILL), WASHINGTON CO.: rvington, French Sen. (S

COs: Joliet, Gaels 420 (F), "wEaNesaGe 0.: Rockford, Fuller &
Haime 2131H (ISM). WOODFORD CO.: Adams Mill, eee Sen. (ILL),

1y. Juncus torreyi Coville, in Bull. Torrey Club 22: 303,
(Map il).

“BI.

presentative Specimens: ADAMS 0O.: 1 mi. N. of Ursa,
Brinker ors (ILL), ALEXANDER CO.: We of Cairo, Winterringer 3122
(ISM). NE CO.: Near apo rol 54863 (ISM). BROWN Co.:N.
of tis. peed 2951 (ISM), OLL CO.: lississippi Palisades,
Wanderlin TOT (Wu), CASS CO.: oe Ashland, Chase 11292 (Il).
GN i

Andrews bo
ann S26 Gig). COOK 00.: ft oes Grove, Glassman 3735 (CHI).
CRAW O.: 1 mi. S. of York, Ozment & DeFilipps 1250

25 Cre ‘

=a: la,
: NE. of Wheaton, Moffatt ay (iy. EDGAR oa, of
Chrisman, Winterringer 1156 ‘a ou oe a Mie 4 ae Bro=

FORD CO.: Roberts, Ahles 317 (ru). or RANKLIN OO. “Taton Lake,
Sands sen. (SIU), FULTON CO.: - S. of Vermont, Evers 30935
(IIs), panei 0.28.08 Cee eg “Evers 17769 (ILLS),

CO.: SW, rdner, Evers 21683 ( reap HANCOCK CO.: E. of Ni-
ota, pets is erst (ILLS). HENDERSON CO.: Near Oquawka, Patterson
Sen. (F), HENRY CO.: 1 mi. E. of povetbhes Dobbs s.n.

TROQUOIS CO.: Sand region, Tehae s.n. (ILL). JACKSON 00.: Bos-
quent Cranwill s.n. (ILL). JEFFERSON CO.: E. of Opdyke, Evers

os apes DAVIESS CO.: mi. S. of Galena, Ozment &
i

: gi e

Eas Anne, Graham 9332 (ISM). KNOX 00.: Altona,

ae terrhigor "1317 Tiel) LAKE 00.: Beach, Fuller 9390 (ILL).
Rock State Park, - rner sen SNe search ogg
RENCE CO.: E, x F Biiate, Evers 371) ~ LEE 00.:
Long 201 (ILL). LIVINGSTON CO.: NW. of Sis reun, Evers “5s (ius),
LOGAN CO.: Near lincoln, Fuller 7182 (ILL). MC DONOUGH OO: N.
of Bushnell, Myers 1166 (ISM). MC HENRY CO.: Algo aati Nason
° Nomar’ st r 180 (ISM). — ae a of

Robert—
Son sen. (ILLS). MARION 00.: NE. of Alma, Peale roe (irs).
“Ree -: W. of Varna, Evers 109 7 (ILLS), MASON CO.:
Mason ae Evers £881 (ILLS). MENARD CO.: S. of Oakford, !
roat 516), ( ae "MONTGOMERY CO.: Litchfield, Jones 2206 (rir).
Nose apis :

seorda, sate SeNe ee PERRY CO.: S. of Sunfield, ele

30560 (IILS). PLATT CO.: NW. of Lodge, Beare 46885 (ILLS}.

52 Poy 7.0L OG TA Vol. 13, no. 1
PUTNAM CO.: S. of Swan Lake, Ozment & ein 1:5 (SIU), RAN-

DOLPH CO.: 5 mi. S. of Sparta, Winterringer SM). RICHLAND
CO.: mi. N. ee Olney, Scherer & Shaw 60> (iT 15 "ROCK ISLAND

=

CO.: Near P Byron, Fuller der 13787 (Th), Ske IR CO.: Cahokia,

E 13926, ay SALINE CO.: 3 mie SW. of Eldo Sethe Evers 51586.

TES. ~~ SANGAM — ed E. of Buffalo Hart, Evers 12037 (ILLS).

SCHUYLER CO.: S, f Frederick, Evers 5910 ” (TEES). aes oss Mer~

ritt, Rexroat 3797 (ISM), SHELBY CO.: 2 mi. E, Towe

Boewe Sen. (ILLS). STARK K CO,: SE, of Wyoming, Evers en (ru),

TAZEWELLCO.: E. of Mackinaw, ae 18218 (ILLS), VERMI

Muncie, Bell 231 (ILL). == ge a — Srearer aca. (i "Hh,
LL CO.? 1 mi f Brai

Mle ° of (CHI °.
Kent Creek, Fell S1250" (ist) angen re. of Minonk, Evers

S610 (ILLS),

15. Juncus brachycarpus Engelm. in Trans. Acad. St. Louis 2:
LST. 1868. “(Map 15)

: Representative Specimens: ALEXANDER CO.: Near Diswood,
winters r 10 (ISM), BOND 00. Eo of Greenville, Winterrin-
et 2 5 ST

CO.: Andrews sen. (I
CLAY CO.: S, of “Flora; Winter r 8206 (ISM), CILNTON Co.:
of Posey, Evers = 21328 (TS ae COOK CO.: Thornton, rae a “(r3u).
CRAWFORD CO.: Between Oblong and Stoy, Ahles °
LAND CO.: S, - Greenup, Winterringer 1290 (ISM). snl rn
2 mi. W. of Samsville, Evers 70576 (ILLS). EFFINGHAM CO.: NE, of
6 (ILLS). FAYETTE

HAR
DIN 0O.: Near Rosiclare, Winterringer 2h ). JACKSON CO,:
a me Mohlenbrock a SIU), JASPER CO.: SE, of Newton,
vers LLS), FERSON CO.: SE. of Opdyke, Evers SHES
(TELS). KANKAKEE CO,: SE, of St, Anne, Fuller 893 eg ery
RENCE CO.: E. of Billett, Evers 29612 (TEES). WACO! 1m. N.
of Decatur, Shouse Sen. (ITD). ACE + Canddawinie, ew
son sen. (I DISON CO.: Near Canteen ‘Lake, Eggert s.n. (MO).
MARION co.: Salen, Ahles 2886 Ci), “uC HENRY 00.7 Winewocd
Vasey son. (ILL), MENARD CO.: Athens, Hall s.n. (F). MONROE
COL: 10 = SE. of Waterloo, Winterringer’ 6666 (ISM). MONTGOMERY
sib S. of Litchfield, Winterri 3897 (ISM). PEORIA ge age
ay Brendel Sen. (ILL LL) O.: 3m. N. of Tamaroa
6601 LL TT CO.: 2 mi. W. of Monticello, Evers "33310 o¢rtts
(0.: Herod, Schopf s.n. (ILLS). PULASKI G0
Winte inger 11128 (ISM), RANDOLPH CO.: Red Bud, Pome!

Sen. (MO), RI CO.: 3 mi. Ne of Olney, Scherer ILL).
ST. R CO.: Brendel s.n. (ILL), e: Near Texas City,

Buser 6251 (ILLS), SANGAMON 0O.: : Between Bivartin & Illiopolis,
Winterringer 1 17220 (ISM), UNION 0O.: Panther's Den, MacMahon 68
e WABASH H CO.: Near Mt. Carmel, Schneck SeoNe (ILE). WASHING~
TON CO.: cpg. French sen. (SIU), WHITE CO.: E, of Norris
eae saise SEES) (TLDS), WILL CO.: 1 mis SW. of Braidwood, Evers

AMSON CO.: S, of Cambria, Evers 37439 (ILLS).

1966 DeFilipps, Juncus in Illinois 53

162 Juncus scirpoides Lam, Encycl. Meth. Bot. 3: 267, 1789,
ars actrpoldes” (Map 16)

ae arr SS 0O.:
3920 (F). yma CO,: f Billett, Evers 3707, (113) ae
WARD CO.: S. of niles ‘Reka 5515" i. § pears

17. Juncus diffusissims Buckl. in Proc, Acad. Phila. 1): 9.

presentative Specimens: JACKSON CO.: Giant City, Mohl-
gnemock 25 ge ). JEFFERSON CO.: Mt. Seo City Reservoir, Fore

5 (SIU). LAWRENCE CO.: 1 mi. NE. of Sunade: Sivert
Bde va MASSAC CO.: W. of Midway, fond "67062 (ILLS).

18. Juncus nodatus Cov. in Britt. & rie Illustr. Fl. N.
Une Ste, ed.2, 1: 82, 1913. (Map 18)

Representative Specimens: BOND CO.: Near reais.

Pepoon & Barrett 5167 Spee CHRISTE AN CO.: Lake P x sea c
Sen. (ISM). CLAY CO.: S, o f Flora, Ahles 2672 aoe
Con: page pc Evers 30850 (ILLS). oe :
=013 ISM). FRANKLIN CO.: Benton Lake, Fore Stookey iy 68 (St A Taty,

HANCOCK CO.: Augusta, Mead Sene (MD). + ACRSON CO.: Crab Or
ioe Abney & Dillard 113 (SIU). pote IN rei No of Mt. = ntl
ore & Stookey 74h (SIU). on ei CO.: E. of Sumer, Sivert son.
Wat % Py TWACOUPIN-GO : Carlinvill sant Raeting Sen. (

. ILLS).
CO.: Salem, Ahles 2887 Les WONT NTGOMERY CO.: SW. of Litehtiel4,
Evers 2078 (ILLS). POPE CO.: Near Bell Smith Sprin ty He Bail
Sw 302 028 See a ST. CLAIR CO.: Brendel s.n. (ILL ‘AS a
Oat tin Erenes srengh SDs (SIU CO.: Fairfield Reservoir,
re & Siete 727 (SI WHITE CO.: Norris City, Buser 2589

of Scher 27 CO.: Cambria, Bell s.n. (SIU),

19. Juncus acuminatus Michx. Fl. Bor. Am. 1: 192. 1803. (Map 19)

Representative Specimens? ALEXANDER CO,: Cache, Cranwill

Sone (ILL), BOND CO.: E. of Smithsboro, Evers 18053 Sg CASS
Col: Blurr Springs, Rexroat 3785 (ISM). CHRIST

ville, Andrews s.n (Tn. or CO.: Fqagedin Hill 36. isan fii),

CRAWFORD CO.: on. S. of Heat. ville 7; Evers 4089

sey CO.: Ne of Neoga, Evers 38221 { lS EDHARDS CO.:? 3 Mi Ne
bion, Fore & Stookey 716 HAM CO.: S, of Mont

ses Evers 1448 (1 ° FAYETTE CO.: 2 mi. N. of Hagerstom,

Evers 23866 bt FRANKLIN CO.: Ben (SIU).

(ILLS), HANCOCK 00.7 Augusta, Mead sen. (M0). HARDIN CO.: NE. of
rar ieg prone ee 346 (ILLS). "HENDERSON CO.: Oquawka, Patterson

12 (F Ow: 3= mi. N. of Annawan, Dobbs sen.
TROQUOIS ie Seca region, Tehon seme (ILLS). JACKSON CO.: Giant
a "es 4 (SIU), JEFFERSON CO.: NE. of Mt. Vernon, Boewe

SON CO.: Busby Hollow, Voigt s Sele (SIU), KAN-
i Pt Hill 51,1871 (ILL), LAWRENCE CO.: E, of

5h PHYTOLOGIA Vol. 13, no. 1
Billett, Piece 29611 (ILLS), LEE CO.: Amboy, Lon eee (ILL), MC-

ef wood, Vasey sen. (F). MACOUPIN ee
Andrews SeNo (HLL) » MADTSON CD CO.: H. Eggert sen. eg
CO.: Bebb S +n (F), MASON CO.: W. of Saidora, yf aa we
MASSAC CO.: NE. of “gf pate ope aeennaer 1s919 (ISM). MENARD
GOs: Athens; Hall 1 ps (F), MO COs: of go ead Bollwin-
kel 253 (SIU), PEORIA CO.: aovncleys Son. "(F). PERRY CO.: S. of =
atts, Evers 18722 (ILLS), foie 0O.: N. of bea
657 (I ; ad NE. o ambersburg, Evers 60233 (ILL i
POPE CO.: E, of Di n Springs, pent 23552 (ILLS). PULASKI CO,:
N. of Pulaski, Evers héib6 CIS). RANDOPPH 0O.: Coulterville
Reservoir, Fore & Stookey 645 (stu), ag Saar CO.: Decker eee
ship, Rid 2hi (F). we ISLAND CO.: Port abe: Fuller 13789
(isu) "Ss Chale 00. W. Edge pet ‘Neill ou :
orado, ecaee erig (IL LS). STARK ear Wady Petra,
ied 678 (IIL). TAZEWELL CO.: Pekin, Chase 1061.8 (ILL), UNION
st S. 0 Pita, Pepoon & Mohr 307 (ILLS). WABASH CO.: Shearer
farm, Shearer SeMe (TEL seme cllatall CO.: Venedy coe, Evers
33940 (ILLS). WHITE CO.: S. of Nor s City, Evers 17782 ( oy LES).
WHITESIDE CO.: 1 mi. We of Erie, Puller peeve (ye WILL CO.:

2 mi. SE. of ste! Eark, Steyermark ILL). WILLIAMSON CO.:
Carterville, Bell sen. (ar0) NNEBAI come 3 mio We. of Shir-
land, Fell & Fell FL6L01 (ISM).
20. Juncus canadensis J. Gay e in Mem. Soc. Histe
Nats Paria Tt Ts. 12S. (ilep 20)
sentative Specimens: CASS CO.: W. o
Rexroat 7 waee > (TSU). COOK CO.: Thornton, Fuller ia Ge th Hal

e? Augusta, Mead sen. (MO). HENRY CO.: 2 mie
daker's, Do Ne 3 f gets Tee,
Evers e710 (T TLLS), KANKAKEE CO.: Kankakee, Hill. 91.1973 (ILL).
LA O.: Zion, Evers 0109 (ILLS), LEE CO, ae ae Lon oh
HENRY CO. Ringwood, ool SoM. (F).

Athens, Hall s.n. (F). OGLE CO.? Pine Rock Bog, en ongberg 63/158
(ISM). WABASH CO.: About Mt. Gormel. Schneck son.

Wt
CO.: SW. of Wilmington, Pepoon & Barrett 536) (ILLS). WINNEBAGO
CO.: NW. of Rockton, Evers 10226 (ILLS).

él. Juncus brachycephalus (Engelm.) Buch. in Bot. Jahrb. 12:
268. 1890. Tip 21)

Representative Specimens: COOK CO.: Englewood, Hill 114
LL). KANE CO ae ets ae B ace (F). KANKAKEE CO.: Kankakee,

an 107 107 (TLL) cone ts), Peter Benke re 933 2 CF). MC HENRY CO.:
of Hebron —— Brere Ol ILLS ae Brendel

ane (LiL), TA é5 (Ct ig Ree OTF

I

N, of ity aie iver rs 21739
wn a CO.: oh Mills, McDonald sen. (FT.

22. Juncus alpims Vill. Histoire des Plantes de Dauphine 2!
233. 1787. vare rariflorus Hart tme, Skand. Fl., ede 7;

240. 1858, (Map 22a)

1966 DeFilipps, Juncus in Illinois 55

Representative Specimens: COOK CO.: South Chicago, Hill
3521875 (ILL). KANE CO.: Elgin, Vasey Sen. (F). LAKE CO.: Wau-
kegan, Gates 2923 (ILL), MC HENRY CO.: Ringwood, Vasey sen.

(F 67500)

Juncus alpims Vill. var. fuscescens Fern. in Rhodora 10:
1B. 190s ~ (Map 22b) a

Representative Specimens: COOK CO.: Chicago, Lansing 257
(F). LAKE 00.: Wankegan, Gates 2859 (F).

Map 1 Map 2 ae
Juncus secundus Beauv. Juncus dudleyi Wieg.

VIDOTOLAHd

7 7on “Ct *ToO,

Map
Juncus buf

8a Ww
onius L. var.
bufonius

996T

STOUTTTI uy snoune ‘sddtttjeq

Ls

gS

VIOOTOLAHG

Map 8b Map 9 Map 10
Juncus bufoni Juncus effusus L. var, Juncus balticus Willd,

nius L,
vare congestus Wahlb, solutus Fern. & Wieg. var. Jittoralis Engelm.

Ton “CT “Tos

7
@\@

e 6ie

ele bad

L) € eé@

ele \e\e

\ele

Map lla Map 1 Map 12

Juncus biflorus Ell. Juncus biflorus Ell, Juncus marginatus Rostk,
f. biflorus f. adinus Fern, & Grisc.

996T

STOUFTII ut snoune ‘sddtrtgeq

6

09

YTUO TOA Ss

7 *ou ‘CI ‘toa

996T

STOUTTTI ut snoune ‘sddt Taq

Map 19
Juncus acuminatus Michx. —

VIODOTOLAHG

Ttou *€T. *ToA

996T
a

dt {tae

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i

STOUTTT

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FRITOLCGGIA

Vol. Ady beso 1

JO DAVIESS STEPHENSON WINNEBAGO [BOONE | McHENRY LAKE
CARROLL OGLE
DeKALB KANE COOK
DuPAGE
WHITESIDE nT
EE
KENDALL WiLL
LaSALLE
BUREAU
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UNDY
ROCK ISLAND 8
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STARK PUTNAM
KNOX
=] MARSHALL LIVINGSTON
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c=] IROQUOIS
= PEORIA
8 WOODFORD
—) MoLEAN
AZEWELL
HANCOCK MeDONOUGH FORD
VERMILION
‘MASON
CHAMPAIGN
SCHUYLER wae ti DewitT
ADAMS
PIATT
BROWN MENARD
Sai
MORGAN NGAMON ‘DOUGLAS EDGAR
PIKE
scorr
MOULTRIE
HRISTIAN
COLES
SHELBY
GREENE MACOUPIN
CLARK
own — CUMBERLANO
FAYETTE EFFINGHAM
JERSEY JASPER RAWF OR
MADISON BOND
CLAY
wRi
aannon RICHLAND La ENCE
CLINTON
‘SAINT CLAIR
= EOWARDS WABASH
|ASHINGTON JEFFERSON
MONROE
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FRANKLIN
JACKSON
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ILLIAMSON SALINE GALLATIN
UNION JOHNSON POPE JARDIN
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PULASKI

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7

PHY TOLOGIA

Designed to expedite botanical publication

Vol. 13 March, 1966 No. 2 ©
CONTENTS
WURDACK, J. J., Certamen Melastomataceis X hi ee ee ee a 65
MOLDENKE, A. L:, Book zeviews. — 3 623 3 81
SMITH, L. B., Notes on Bromeliaceae, XXIII . 84
MOLDENKE, H. N. , Materials toward a Bes Ge a the genus
Lippia. x Me es is os . 162

(| 0 Published by Harold N. Moldenke and Alma L. Moldenke

303 Parkside Road
ce eeaben New Ages U.S.A.

Price of this. number, $1; per volume, $5. 75, in advance,

of volume
Wresoun Botanica or $6 at close

APR 11 1966

GARDEN Lisrary

2562 {
CERTAMEN MELASTOMATACEIS X.

John J. Wurdack
Dept. of Botany, U. S. National Museum

a botanist whose ultimate aim is taxonomic orderliness,

continuation of my velitation account non-Ecuadorian
novelties collected by the Maguires during their 1959 Andean
trip, miscellany discovered during as iW. Be

cuadorian colle tions, and asticcio from u. Also _

mino y
Melastomataceae for the Flora de Venezuela so as to ba with
my crotchets.

MACATREA LASIOPHYLLA (Benth.) Wurdack, comb. nov.
Shactogastra lasiophylla Benth., Hook. pay Bot. 225-2914
1840
erolepis lasiophylla (Benth.) Triana, Trans. Linn. Soc.
Bot. os 40. 1871
Tibouchina lasiophylla (Benth.) Cogn., Mart. Fl. Bras. 14
(3): 297 85.
Actémitheta erecta Gleason, Bull. Torrey Club 58: AISI 21931:
Acisanthera lasiophylla (Benth. ) Gleason, Phytologia 3:
243. 1950.
Sephantheropsis williamii Brade, Inst. Nac. Pesqu. Amaz.
Publ,<Bot./ 8-6,
One of the Savaenata circumstances for scientists in
botanical taxonomy is that plants, unlike wives and some other
fa

above-cited Se Semis cages e pubescence and
floral features indica see nL. tadaantiviil represents a reduced
evolutionary offshoot ss opie to M. multinervia Benth., wit

stamens really quite pmo to other species in the genus.
R.I.P.

ae ee Wurdack, nov

anther ae (bent: ) Cogn. affinis, se
chlvcia. male eeee glabris —_ “ apicibus ultimis apares ee
glanduliferis ornat

Ramuli teretes mises rotundato-quadrangulati sicut inflo-

rescentia sepala extus hypanthiaque modice vel den — pilis
erectis gracilibus laevibus glanduliferis 1-2(3.5) mm longis
setosi, ramis infra pilis non glanduliferis. Petioli . 3-1 cm
dense breviterque setosi pilis gees non glandulosis;

66 PHYTOLOGIA Vol. 13, no. 2

lamina 3-5.5 X 2-3.5 cm firme membranacea integra vel obscure
erenulato-serrulata late ovata vel elliptico-ovata apice late
hebeti-acuto vel acuto basi rotundata vel paulo (ad 4 mm) cordata

superficieque modice vel sparse setulosa. Panicula terminalis

ciliis glanduliferis 0.2-0.3 mm vel apicem versus usque ad 0.7
mm longis. Stamina dimorphica glabra; filamenta 12-15 mm vel
8.5-10 mm; antherarum thecae 9-12.5 mm vel 6.5-8 mm subulatae,
connectivo basi 3.5-5 mm vel 1-1.5 mm prolongato ventraliter
1.2-1.3 mm incrassato et bilobulato. Stylus 16 X 1 mm glaber;
s atum non expansum; rium apice densiuscule
striguloso pilis non glanduliferis, lobis apicalibus 2 mm altis.

Type Collection: J. J. Wurdack 632 (holotype US 2404208;
isotype USM; 12 additional isotypes to be distributed),

Molina 5

collected in Quebrada km below Chachapoyas,
oh chapoyas, Depto. zonas, Peru, elev. 2200 m, 30 May 1962.
Lax s -~4 m, locally frequent in moist ravine at forest

edge. Petals magenta; anthers purple with yellow connective
prolongation."
Paratypes (all Peru): Cajamarca: Cerros de Cutervo,
Ferreyra 814; Cutervo, Ferreyra & Acleto 15374. Amazonas:
uma-urcu, Soukup 4048; Pomacocha, Ferreyra & Acleto 15171;
co

Cdclic, Lépez Miranda, Sagd4stegui, & Collantes

Calla-Calla, Hutchison & Wright 5649.
The epithet chosen commemorates my botanical artist and

plant nurse, © grew the species from seed to flowering size.

has

obtained a mitotic chromosome n r (to be published elsewhere) -
Through F. G. seeds w rious
botanic gardens. Tibouchina mariae is perhaps the most abundant
melastome in the Pp
is readily propagated by cuttings. While the flowers are ve
attractive, the sprawling habit, with rooting at the nodes,
detracts from possible use as a house plant.

cal element in T. laxa has roughened cauline and

photograph 36166). After examining a type sprig (BR) of

1966 Wurdack, Certamen Melastomataceis 67

i. cymosa Cogn. and also the holotype (US) of T I. asperipilis
2 i can see no reason for any separation from typical

ty
Azuay, and Loja in Ecuador to Piura and Cajamarca (but not
Amazonas) in Peru.
There are tw poss not too clearly defined variants of

laxa; one Sa ooth (or nearly so) but eglandular hairs
peop Canar: Biblian, Asplund 17644; Asorgues, Pauls 7083.

Azuay: Susudal, Asplund 17814. Peru: Cajamarca: Colasay

Woy thaalenied 6908), while the other has roughened and reed vices

hypanthial hairs (Ecuador: Guayas: 118 km east of Guayaquil,

Maguire & Maguire 44266. Loja: San Pedro, Towns Townsend Al05. Peru:

Piura: Ayabaca, Soukup 4299; Abra de Eebeullay Bare Ferreyra 13752.
The

Cajamarca: Hualgayoc, Soukup 3853). o forms are by no no
means as sharply delimited from the Bie 85 pat mind oft

s e
species as these specimen citations would indicate, there i
Miers in abundance of eteadubantce and degree of trichom

roughening.
Apart from the collections cited above for T T. maria
specimens from Huambos, Cajamarca, Peru, seem best Soaks hy cna

species, although one (Soukup 4479) lacks gland-tipped hairs and
the other (Ferreyra 8436) has very sparse glandular hairs on the
pedicels and hypanthia. The Ecuadorian T. pendula Cogn. resembles
pe tg

with very short connective prolongation and hypanthial hairs very
slightly roughened and eglandular.

TIBOUCHINA piste aia Cog
After comparison with ean ae 5002 (BR), a series of
recent pn collections have been referred to T. weberbaueri:

Vargas 3378. d by Macbride in the Flora of Peru, Mexia -
8056 is also this sa Two features not noted in the original
description are th ominance of 4-merous flower d the

b
haire,-< Both ‘t, decora Cleasen and T. tet Eapet ala

t orm
5-merous flowers, while the pear i ang 2423» a has Senitais
barbellate (rather than smooth) cauline hair

68 PHYTOLOGIA Vol. 13, no. 2

ERNESTIA KARURUANA Wurdack, sp.
E. tenellae (Bonpl.) DC. affinis, indumento hispido in

glanduliferis in foliis plerumque non glandulosis. Petioli ca.
0.2 cm; lamina 2-2.5(-3) X 1-2 cm ovata vel oblongo-ovata apice
paulo acuminato basi cordulata 5-7-nervata fragilis setoso-
serrulata. Paniculae 6-13 cm longae multiflorae; pedicelli

2.5-3 mm longi; fl 4- Hypanthium (ad torum) 2.5 X 1.7
modice pilis glanduliferis gracilibus 1.5-2 mm longis obsitum;
calycis tubus 0. tus, lobis 4.3-4.5 X 0.7 mm anguste

lanceatis apice setiferis. Petala 9.5-10.5 X 6-8 mm asymmetrice
elliptica quaeque apice setula unica armata. Stamina dimorphica
glabra, filamentis 5.5 mm vel 3.5 mm longis, antheris poro

ventralibus -3 mm longis basi 0.3 mm coalitis simplicibus
vel interdum unibarbellatis. Stamina minora: the ongae,
connectivo infra thecas usque ad calcar dorsale 0.5 mm longato,

per 0.5 mm paulo incrassatis simplicibus usque ad basim divisis.
Stylus 8.5 X 0.2 mm glaber; stigma truncatum; ovarium 3-loculare
pa eg paucis (ca. 8) glanduliferis 0.4-1.3 mm longis

te Collection: R. E. Schultes & 1. Cabrera 19176 (holo-
type GH; isotypes NY, us) , collected in Savannah Goo-ran-hoo-d4,
Mesa de Yambi, R{fo Karurd, 1920' N, 71°20' W, Vaupés , Colombia,
elev. 290-300 m, 15-16 April 1953. "Flowers white

Ernestia tenella a the sists species with pubieruiows
ovaries (E. ovata Cogn., qua eta Berg ex Triana
adenotricha Uribe) have oS apne hairs less Gin - 1 mm long,
foliar hairs less than 2 mm long, and distinctly developed

petioles 0.5-2.5 F previously indicated (Mem. N. Y.-

t ef aati ae ST BREE » at least part of E. tenella
shows predominantly 3-locular ovaries. ong hairs of
E. karuruana suffice as a species distinction from all ¢ yer i ha
All of the two flower ovaries and six fruit capsules visible 0

Schultes & Cabrera 19176 showed 3 cells

MONOCHAETUM HUMBOLDTIANUM (Kunth & Bouché) Kunth ex Walpers var-
HIRTUM (Karst.) Wurdack, comb.

Grischowia hirta Karst. , Auew: Neue Gew. 16, pl. 5. 1848.

Monochaetum hirtum (Karst. ) Triana, Trans. Linn. Soc. Bot.
28: 64. 1871.

Monochaetum brevifolium Gleason, Am. Journ. Bot. 16: 522.
1929.

1966 Wurdack, Certamen Melastomataceis 69

No specific distinctions based on pubescence posture or
density are possible in the abundantly collected species of the
Venezuelan Coastal Cordillera. If the Pico Naiguata sampl

plants have denser and erect stem hairs. In addition t
collections cited in 1929 for M. hirtum and M. brevifolium, var.
hirtum is also now represented by Steyermark 62961 from Naiguata
and Killip & Rohl 37161 and 37204 from between El Junquito and
Agua Negra (D.F.), as well as the following originally cited by

Rancho de Boqueron) varies slightly from var. humboldtianum

toward var. hirtum,. Steyermark 61589 from Anzoategui (cited in

Fieldiana Bot. 28: 1037. 1957 as M. hirtum) actually represents
ciduous in

and foliar pubescence.

MONOCHAETUM HUMBOLDTIANUM (Kunth & Bouche) Kunth ex Walpers var.
CHARDONII Wurdack, var. nov.

A var. hirto differt foliis supra densiuscule aequaliterque
brevistrigosis pilis gracilibus 12-14/mm quad.

Type Collection: C. E. Chardon 201 (holotype US 1801626),
collected along the highway to Choroni, Edo. Aragua, Venezuela,
elev. 1200 m, 26 Oct. 1940.

type: G. Ferrari 72 (Herb. Maracay), from "cumbre de

Para
la carretera de Choroni," Edo. Aragua, Venezuela, elev. 1500 m.

MONOCHAETUM GLEASONIANUM Wurdack, sp. nov.
S Grischowia. M. lindeniano Naud. affinis, sed ramubrum

ect,
pubescentia parciore hypanthiis sparse gracili-strigulosis
tami re) bu.

Frut
petioli foliorum venae primariae subtus sparse vel modice laxo-
Strigulosi pilis gracilibus laevibus, in nodis gracili-setosi.
Petioli 0.5-1 em; lamina 2-4 X 1-2 cm ovato-elliptica apice
acuto vel paulo acuminato basi late acuta, breviter 5-plinervia
Pare interiore 1-3 mm supra basim inserto nervis secundariis
Supfa occultis, supra in lineis inter nervos sparse gracili-
i e

laxostrigulosa. Flores in dichasiis terminalibus 1-3-floris
dispositi; pedicelli 0.5-1 cm longi sicut sepala extus hypanthi-
aque sparse strigulosi pilis gracilibus 0.3-0.5 mm longis.

70 PHYTOLOGIA Vol. 13, mo. 2

Hypanthium (ad torum) 6 mm longum extus in tori —— ape
pilis suberectis gracilibus 1-2 mm longis armatum; sep a 6-7 X
anthesim decidua lanceata acuta ciliolata. cael non
en mm

visa. Stamina glabra, filamentis 5 ongis. Stamina maiora:
antherarum thecae 4 mm longae, appendice dorsali 3.5-4 X 0.5 X
0.7 apice rotundat ere minora: antherarum thecae 4 mm
longae, appendice dorsali 3. .3 mm circum medium 1 mm lata
et paulo auriculata apice “armen eet X 0.4 mm
gneser' ovarii apex — = - glanduliferis modice armatus.

e Collec Reed (Syracuse University Andean

Typ on:
Expedition) 32 ors is e US 1618468), collected in terrace
alluvium at Chac chapo, Edo. Mérida, Venezuela, elev. 2620 m,

Paratype: Reed 48, from the same locality.
Monochaetum lindenianum has Saas spreading cauline pubes-

morphic stamens. The other species of Gleason
P

s.
tions as Sharp undescribed and had noted the affinity with
M. lindenian

AXINAEA TOVARII Wurdack, sp. nov.
A. sclerophyllae Triana et A. tomentosae Cogn. affinis, sed
foliis proportional iter angustioribus subtus demum glabratis.
Ramuli primum quadrisulcati demum rotundato-quadrangulati

stout pe petioli : alae venae primariae hypanthia (basim versus)

inflorescentiarum rami pedicellique primum dense pilis barbellatis
0.4-0.7 (-1) mm longis erectis armati demum maha Petioli
3.5-5.5 cm; lamina 11.5-21.5 KX 3.5-8.5 cm subcoriacea basim
versus integra apicem versus inco Fi serrulata (dentibus 0.2-
0.3 mm altis et 5-9 mm inter se distantibus) elliptica apice per
1-2 cm gradatim hebeti-acuminato basi sie (basi ipsa paulo

glabra, subtus in nervulis fea ip te pe primum sparse puberula

Panicula 7-10 X 7-9 cm remis chic 4 deflexis, pedicellis 2-5 X
1 mm, bracteolis 0.5-0.7 X 0.3 mm oblongis caducis ob pilos fere
occultis; flores 5-meri. Hypanthium (ad torum) 5.5 X 7.5 mm;

1966 Wurdack, Certamen Melastomataceis 71

calyx 2 mm altus ek Se i limbo integro, dentibus exterioribus
inconspicuis 0.5 mm longis hebetibus vix elevatis. Petala 16-

et inconspicue retuso. Filamenta 8-9 mm labra; antherarum
thecae 4.5-5.5 mm longae oblongo-subulatae poro minuto dorsaliter
inclinato, appendice dorsali inflata 3-4 X 2-2.5 mm apice hebeti.
Stylus 22 X 1.7-0.8 mm in ovarii ast 2 1.5 mm immersus; apg

-8 mm latum; ovarium 5- Gc ibeitare glabrum apice 5-lobulat
lobulis 0.7-1 mm 1baets et 0.3-0.6 mm retusis

Type Collection: Oscar Tovar 4753 (holotype US 2446955

isotype USM), collected in ce ja ja de montana at Marcavalle, se
Huachocolpa and Tinta » Prov. Taya acaja, Depto. spegyr etre
Peru, elev. 2700 m, 20 April 1964. "Arbol 7-10 m. Flor
purptreas."

Both suggested relatives have leaf blades with length/width
ratio mostly 1.5-2 (rather than 2. 5-3.2), as well as persistently
puberulous lower surfaces. Axinaea sclerophylla has considerably
larger hypanthia and A. tomentosa shows slightly deflexed leaf

base auricles. Both A pennellii Gleason and A. weberb
Cogn., with leaf blades distinctly serrulate an sally > appa

a
are more distant relatives; the former species has essentially
glabrous hypanthia, while the latter has quite small hypanthia.

MERIANIA BRITTONIANA Wurdack, nom
Axinaea speciosa Britton, Bull. oe: Club 17: 57. 1890,
non M. speci speciosa (Bonpl.) N
MERIANIA CUZCOANA Wurdack, sp. nov gAet
M. brittonianae Wurdack affinis, sed ramulis novellis inflo-
escentiisque glaberrimis foliis subtus non furfuraceis trirenatis.
uli primum indistincte qiabanguiacs demum hyenas a sicut
folia inflorescentiae hypanthiaque glabri. Petioli 2-2.5
graciles; lamina 5.5-9.5 X 2.5-3.2 cm tenuiter coriacea ihteers
elliptica vel paulo oblongo-elliptica apice acuto basi obtusa
ple a urvata

puncticulata punctulis primum nigro-rubris demum nigris. Panicula
8- cm longa submultiflora; flores 5-meri, pedicellis 1. 5-2 cm
longis ad medium vel paulo supra articulatis, bracteolis dela il

aia sivbats 3.5 mm longo (distaliter per 0.5 mm subabrupte
hebeti-caudato) et dorsaliter dente ascendente hebeti crassius-—
culo 0.5 mm libero armato Stylus 11-15 X 0.9-0.5 mm glaber in

72 PHYTOLOGIA Vol. 13, no. 2

ovarii apice 0.7-1 mm immersus; stigma truncatum non expansum;
ovarium 5-loculare glabrum oie: lobulis 5 breviter emarginatis
0.7 mm Sea srmero :
Type Collec ‘ Vargas C. 5107 (haloes US 2438729;
isotype CUZ), pang in "laderas boscosas de Pillahuata

rov. Paucartambo, Depto. Cuzco, Peru, elev. 2000 m, 9-10 Ma ay

pri
blade base) and pay stellulate- or pinoid-furfuraceous
young branchlet inflorescences. From the general facies,
ag a eat probably be aligned with the 6-merous
exame The sectional de-

rg
‘al
ish)
wn
~
val
>
ae
¢
i)
5 A
KSIES
[>
e
a
be
» Io
5
n
lo
n
=
(=|
n
ian
i‘)
ie)
tom

Britton's epithet is unavailable in Meriania, having been pre-
occupied in M. speciosa (Bonpl.) Naud

ne! MERIANIA VARGASII Wurdack,
In an thadatterteal M. _Salophyllae (Cham.) Triana et

M. arboreae (Naud. ) Triana affinis, sed foliis tenuioribus late
oblanceolatis 5-plinervatis hypanthii pilis longioribus differt.

Ramuli um sulcato-quadrangulati demum subquadrangulati
sicut petioli inflorescentiaque pilis 0.5-1 (in nodis usque ad
2) mm longis basim versus paulo barbellatis interdum minute
caduceque glanduliferis armati tarde glabrati. Petioli ca. 1.5
em; lamina 16-20 X 5-7 cm membranacea apicem versus serrulata
(dentibus 3-4 mm inter se distantibus et ca. 1 m altis) late
oblanceatis apice paulo hebeti-acuminato basi attenuata (basi ipsa

us
tubus 2.5-3 mm altus, lobis interioribus membranaceis 7-7.5 mm

inclinato. Stamina maiora: antherarum thecae mm longae, cornu
basali 3 mm longo, appendice dorsali ascendente 1.5 mm longa.
Stamina minora: antherarum thecae 9 mm longae, cornu basali

1966 Wurdack, Certamen Melastomataceis 73

2.5 mm longo, appendice dorsali ascendente 0.5 mm longa.
Stylus 15 X 0.8-0.6 mm glaber in ovarii apice 1.4 mm immersus;
stigma truncatum 0.8 mm latum; ovarium glabrum apice paulo

b

ype Collection: C, Vargas C. 13240 (holotype us 2438703;
isotype CUZ), collected at Hacienda Guayanay, Prov. Convencién,
Depto. Cuzco, Peru, elev. 1800 m, 15 May 1960. "Arbusto 3 m in

Te |

ile the ascending dorsal connective appendages would place
M. vargasii in Sect. Adelbertia Triana, probably genetically
closer relatives are to be found in Sect. Umbella a

ther connective. e ivian species differs fr M. vargasii
in the shorter Sparser pubescence and elliptic markedly pl
nerved leave he inner pair of primary lateral veins div ging

n
elliptic leaves; the ormer species has reticulate-bullate upper
leaf surfaces, while the latter shows very short hypanthial

nce

Vargas 10644 (US), from Tunquimaya, Prov. Convencién, Cuzco,
is certainly conspecific with M. vargasii and, in vegetative
feat ow y

external calyx teeth are not emergent (about 1 mm r than
the calyx lobes); the 1 rge anthers have a distinct ascending
dorsal toot i connectives, while the small s have

only the barest suggestion of such a tooth. I hesitate to
formally describe a variety without further collections.

MICONIA LITTLEI Wurdack, sp. nov.
ect. Tamonea. M. longicaudatae Cogn. et M. leucanthae
Gleason affinis, sed floribus minoribus.
6-15 m alta; ramuli primum sulcato-quadrangulati demum

oO
es
n
n
@
:
a,
ty
Le |
=
ad
a”
g
na
os
ie)
Ln}
n
@
n
ct
o
ram
a
c
e
i*})
rt
i
=
o
(0)
4
¢
~
&
=|
°
*
a9
ey
o
a
fo
rt
~

in superficie glabra. Panicula 7-10 (vel in statu fructifero
usque ad 15) cm longa pyramidata multiflora; flores 5-meri,

Th PHYTOLOGIA Vol. 13, no. 2

pedicellis ca. 1.5 mm longis plerumque ad medium articulatis,

bracteolis 0.5-0.6 X 0.1 mm linearibus valde caducis. Hypanthium

(ad torum) 1.9 mm longum extus densiuscule minutissimeque

stellulato-puberulum; calycis tubus 0.3 mm altus, pebits interiori-
al ibus

apice asymmetrice rotundato. Filamenta 2.3-2.6 m —s a
glanduliferis 0.1 mm longis ornata; antherarum thec e 2.3-2.5 X
0.4 mm oblongo-subulatae minute porosae, connectivo usaiuee non
prolongato. Stylus 3.5 X 0.45 m basim v ersus densiu

ar

1/3-inferum superne costatum in angulis apicem versus modice

Ainge pilis 0.05 mm longis non glandulosis; fructus sphaeroi-
us 3-3.5 mm diam. paulo 10-costatus.

Type Collection: E. L. Little 6177 (holotype US 1877635),
collected in wet tropical forest 12 km west of Santo Domingo de
los Colorados on the trail to Rancho Gr se Prov. Pichincha,
Ecuador, 5 April 1943. "Small tree 20 ft., DBH 3 in. Flowers

le

Ve ame: Co
Paratypes (all Fuchs Prov. gneraldes; Ecuador): 4 km w
of Borbén, iasete 6380 (US); across Rfo Quinindé from peau
elev. 65 m, Little 6249 (US); ant shore of Rfo Esmeraldas
opposite Quinindé (Rosa Zarate), As splund 16338 (S); "Carretera
anto Domingo de los Colorados-Quinindé, km 171-188, hasta el
Rfo Cocola, alt. 100 m.s.m.," Acosta aolis 16219 (F).
Bot

th suggested — have hypanth: ia plus calyx lobes
3.5-4 mm long and petals 4-6 mm long; is porated and Costa
Rican i gra ihe are nas distinct from one another. The
Peruvian M. doneana differs from M. littlei in the canal larger
prominently econ flowers; a recent collection of M. doneana

is Schunke A118 (US), from Schunke Hacienda above San Ramén,
Junin. Miconia caudata (Bonpl.) DC. has persistently furfur

lower leaf surfaces and slightly larger flowers with oblate calyx
lobes only 0.3-0.5 mm long

— oo Wurdack, sp.
- Tamonea. A aaa 22 25 et 36 Monographiae
Copan foliis supra strigulosis differt.

Ramuli primum sulcato-quadrangulati demum teretes sicut
petioli foliorum venae primariae subtus inflorescentia hypanthi-
aque modice furfuracei pilis stellulatis vel paulo pinoideis 0.1
(-0.2) mm longis. Petioli 3.5-6.2 cm; lamina (acumine excluso)

obtusa, 5-nervata (pare inframarginali debili anean nervis
secundariis 3-5 mm ite se distantibus supra planis subtus
creberrime elevatis nervulis subtus planis et reticulatis areolis
0.2-0.3 mm ee supra sparse setulosa pilis simplicibus ca.

1 mm longis et 1/mm quad. laxis, subtus in nervulis minute granu-
losa in Se ee glabra. Panicula 15 X 10 cm pyramidata

1966 Wurdack, Certamen Melastomataceis 75

submultiflora; flores een: pedicellis 1(-2) mm longis robustis,
bracteolis 1. es 1.7 X 0.3-0.5 mm caducis. Hypanthium (ad torum)

bus vaehtle 0.1-0.15 mm eminentibus. Petala 5 X 3-3.5 mm oblongo-
obovata apice rotundato et asymmetrice retuso, extus centraliter
modice stellulato-puberula intus minutissime granulosa. Stamina

glabra in dimensionibus paulo anisomorphica; filame 5~5.8 :
antherarum thecae 5.4 X 0.9 F y gin
convoluto apice minute (0.2 mm) uniporoso, connectivo dorsaliter
-3-0.4 vel prolongato truncato 12

in ovarii apice ersus; stigma capitellatum
diam.; ovarium 3-loculare 1/6-inferum apicem versus minutissine
granulosum.

ype Collection: E. Asplund 19526 Fah 5 S), collected
in rastrojo at Shell Mera, Prov. Napo-Pastaza, Ecuador, elev.

1000 m, 1 Mar. 1956. "Shrub about 3 m high, calyx asin green,

petals white, anthers pink."
All e species of Miconia in Sect. Tamonea with caudate-

oft
acuminate leaf blades have the upper leaf surfaces glabrous.

dua
cordulate leaves, as well as (ex descr.) the | Apia ° Eluniaiaa
filaments and style. rene recent Peruvian collections (US
unfortunately not in flower, of M. ciliaris are Killip & Smith
24739 (San Ramén, Junin) po 24959 (Colonia Perene, Junin). _

MICONTA MITUANA Wurdack, sp. nov.
systema Cogniauxii M. aureoidei Cogn. affinis, sed
petiolis foliis juvenilibus : supra inflorescentiisque sparse
setulosis foliis 7-nervatis bracteis maioribus.
uli primum quadrisulcati demum teretes sicut petioli

10 . -11 cm tenuiter coriacea indistincte ecient
ulata ovato-elliptica apice per ca. l subabrupte hebeti-
a et mucro l- ongo terminato basi rotundata vel

albidis immersis obsita. Panicula 9-13 cm longa
in nodis 4-6-verticillatis, ramulis ultimis plerumque
Cillatis; flores 6-meri, pedicellis ad anthesim 8-10 mm longis;

a)
49

76 PHYTOLOGIA Vol. 13, no. 2

bracteae duae ad hypanthii basim insertae 9-10 X 6 mm obova
ellipticae ad anthesim caducae intus sparse vel ares inane?
puberulae anthium (ad torum) 5. calye -3.

xtus

modice et intus sparse stellulato-puberulus, dentibus exteriori-

bus caeeanad torus pilis stellulato-fasciculatis 0.3-0.5 mm

lon orna : recat 8.5 X 4 mm apicem versus graciliter

prin on alioqui glabra obovato-oblonga apice ono et paulo
retuso. tamina isomorphica glabra; Fi lansiit ta 9-10 mm; antlerarum
thecae 11.5-12 mm longae subulatae, connectivo dered’ ee basim

ce in

collo argenteo-strigilloso ef Ales alto eases stylum protracto.
Collection: Bassett Maguire, Celia K. Maguire,

Alvaro son uaa 44089 Golotyee US fig Asotypes COL, NY),

‘collected at woodland edge, Cerro Miti, Vaupés, Colombia, elev.

400 m, 4 Sep. 1959. "Tree t 70 m; petals pink or orangesred at

base; stamens wholly yetiae 2" Occasional."

Miconia aureoides has stem, foliar, and inflorescenc
pubescence of short thick pinoid hairs, 3- or faintly S-plinerved
leaves, ovate bracts only about 2 mm long, and much shor
anthers. An isotype (NY) of M. aureoides shows both FLanents
and style puberulous va the stamen connect ve basall

calyx, probably M. sa diidulageie fey and M rey Triana are

aspect of M. dodecandra (Desr.) Cogn. (as I hade er and

pa a amed the Mitu colbectteny; but that wt item ad
pecies has a regularly lobed calyx and (usually) 5-nerved leaves

ee are glabrous above.

MICONIA CARPISHANA Wurdack, sp.

nov

Sect. Amblyarrhena. M. cen trophorae Naud. in systema
Cogniauxii affinis sed foliis nem
+

Panicula 4- em multiflora; flores 5-meri conferti 0.5-
1.2 mm pedicellati, bracteolis 0.8-1.5 X 0.1-0.5 furfuraceis
valde caducis. Hypanth (ad torum) 1.6 .8 mm sparse (vel

stellulatis; calycis tubus 0.2 mm altus, lobis interioribus

1966 Wurdack, Certamen Melastomataceis 77

-4 mm altis rotundatis, dentibus exterioribus callosis non
eminentibus. Petala 1.2-1.3 X 1.2-1.3 mm extus el ie intus
minutissime papillosa suborbicularia apice rotundat Stamina
isomorphica glabra; filamenta 2 mm ihiogay mbbeeeren? ehacad 1.4 Xx

fe)
versus paullulo clavatus in ovarii apice 0.5 mm immersus; stigma
truncatum 0.45 mm diam.; ovarium 2(?)-loculare 1/2-inferum apice

babies smrgorde minute (0.2 mm) denticulato.

Type Collection: Bassett Maguire & Celia K. Maguire 44436
(holotype US 2444821; isotype NY), collected on che west side of
poor Pass, Depto. Huan sooninaeh bh Peru, elev.» 2850 m, 22° Oct. 1959:
"Tree to : m. Flowers white.

hess ia centrophora has denticulate leaf blades with
eS ain secondary veins, straight stiff trichomes mixed with
e

and s sparsely glandular-puberulous filaments and style.
ignaria te ex Naud., keyed by Cogniaux to the species grouping
around M. centrophora, actually is, because of its f ss
foliar pubescence, closely related to M. pubescens Soo y DC;
In leaf texture, M. carpishana resembles M. obovata Triana and
M. spatellophora Gleason. The former species sieises at least in
leaf shape and (if my tentative identifications of Pennell 3152
from Tolima and Bro. Tomas 2391 from Caldas are correct) the
obovate broadly pored anthers (of Sect. Cremanium, rather than
Sect. Anblyarrhena) ; the latter has oblong narrower leaves and
shorter sep

MICONIA BENEOLENS Wurdack, sp. nov.

Sect. Amblyarrhena. M. tererae Naud. et M. cookii Gleason
affinis, sed foliis minoribus inflorescentiae ramis ramulisque
amorpho-furfuraceis.

uli primum hebeti-quadrangulati demum teretes, primum ;
Tat

Petioli 1-2 cm; lamina 4.5-7.5 X 2-4 cm subrigida integerrima

elliptica vel paulo obovato-elliptica apice hebeti-acuto

Mebeseaens (usque ad 0.5 cm) lateque hebeti-acuminato basi late
ervis se i

ren se distantibus supra pau pressis subtus inconspicue
elevatis n 1 astomosantibus areoli 5 atis, supra
glabra et nitidula, subtus is primariis sonar oe

0.1-0.15 mm in pedicelli medio
vel paulo supra insertis valde pot ety Hypanthium (ad torum)

78 PHYTOLOGIA Vol. 13, no. 2

1.8-2 mm altum glabrum vel primum tenuiter amorpho-furfurace
mox glabratum; calycis tubus 0.3-0.4 mm altus, lobis interioribus

granulosa suborbicularia paulo apice asymmetriceque retu
Stamina isomorphica glabra; filamenta 1.7-2.1 mm; antherarm
thecae 1.4-1.6 X 0.5-0.6 X 0.5-0.6 mm oblongae apice unipor
(poro 0.1-0.15 mm diam. paulo dorsaliter inclinato), caeantase
nec prolongato nec appendiculato. Stylus 4 X 0.3-0.4 mm glaber
rs sparsissime spenaebpec pupereret in ovarii apice ™mm
mmersus; stigma truncatum 0.5 mm diam.; ovarium A nese apice
aoehee ae 6 mm a glandulis 5 et 0.05 mm longis coronato
e Collection: J. A. Steyermark 54789 (holotype
US isstiaa: isotypes F, NY), collected on the western slopes
the Cordillera Condor and northwest slopes of Nudo de eerie
d Tambo

calyx pale green; anthers dull yellow; leaves subcoriaceous,
deep green above, pale parents seen petiole red; in Eitest with
Cinchona Ee eka hoja de i

Paratype: Reinaldo Espinosa E-1078 soe £21078 (F, NY), from Namanda,
Loja, Ecuador, elev. 1500-1600 m, 24 Nov. 1946. "Arbusto coposo-
Hojas coridceas, brillantes, claras en el envés. Racimos flordes
muy apretados. Flores color blanco, caliz amarillento. Crece in
bosque achaparrado.

Both suggested relatives have leaf blades 10-22 cm Siceng as

ly as
M. c cookii; and soa taae 18365). " Wiconda cookii has laxer leaf
venule areoles (ca. 1 mm across) than M. beneolens. Steyermark

is not at all closely related to that species. I was not able to
determine the ovary-cell number from the available collections
of M. beneolens.

)
£ OMIA MOLINOPAMPANA Wurdack, sp. nov

t. Cremanium. M. glabulifiorad. (Rich.) Triana in foliorum
forma similis, sed ramulis primum rotundato-quadrangulatis,
pedicellorum bracteis minoribus antheris biporosis — ert

Frutex 0.3-0.7 m altus glaber. Ramuli primum obsc

quadrangulati demum teretes. Petioli 0.4-1.2 cm longi graciles;
lamina 2.5-5 X 1.4-2 cm subcoriacea distincte ciliolato-serru
lata ciliis — crassis ca. 0.5 mm longis et ca. 1-1.5 m
inter se distantibus elliptica vel oblongo-elliptica apice acuto
vel paulo (per 0.3-0.5 cm) hebeti-acuminato basi late acuta vel
obtusa, trinervata (nervis primariis lateralibus interdum subtus

1966 Wurdack, Certamen Melastomataceis 79

basim versus per 1-3 mm costa coalitis) nervis secundariis
gracilibus supra et subtus planis vel paulo impressis 1.5-2 mm

Type Collection: J. J. Wurdack 1389A (holotype US 2404293;
isotype USM; 7 additional isotypes to be distributed), collected
in the jalca zone 1-5 km west of Molinopampa, Prov. Chachapoyas,
Depto. Amazonas, Peru, elev. 2400-2450 m, 18 July 1962. "Flowers
white. Occasional.'

a globuli has ‘ah rply 4-angle branches, ee
persistent pedicellar bracteoles 1.5-2 -4- thers
with t ding nearly to the the pect apex. While

resemble M,. molinopampana that Central Mmaticen species has the

pe tals and anthers; as suggested by Macbride and confirmed by
Gleason's notes from Berlin, "he Peruvian record of this species
is surely based on a misidentification, Weberbauer 4105 having

m the lower leaf surfaces. e e collection of
M. molinopampana is female, while Wurdack 1389B is e.
Wurdack 1428, also assigned to M. m a, W Hected
in a sheltered ravine on Cerro Malcabal, above Molinopampa; this
collection has larger leaves (to 10 X 4.5 cm the extensiv

The only other sympatric species of the small-flowered group in
that species is also
apparently dioecious, with Wurdack 1433 eaten Malcaba

Amazonas) being male while Weberbauer 4389 (BR) is fenaia. A

n
co)
°
ct
oO
al
om
B
5
Sioa
5
a
Qu
r
ct
°
)
09
=

80 PHYTOLOGIA Vol. 13, fio. 2

recent male collection from Carpish, Hugnuco (Ferreyra 2376, US,
USM) also seems referable to M. dumetosa, although having 3 Lightly
longer cauline pubescence and thicker petioles. Miconia

but s ored anth and hermaphroditic flowers. Miconia
crassistigma Cogn. (Heberbauer 3407, BR) is certainly specif-
ically distinct from M. dumeto a (although also from

Hudnuco) ;
only female flowers are Siebacri y known and the species. sieges be
evaluated in connection with M. galactantha Naud. when more
material is collected.

HENRIETTELLA MAGUIREI Wurdack, sp. nov

H. membranifoliae Cogn. affinis, aed foliis crassioribus
supra | glabris sclerideis occultis differ

Ramuli robusti obtuse agearangelarty. sicut petioli et folia
subtus primum densiuscule pilis erectis 0.2-0.5 mm longis dense
asperis armati tarde glabrati. Petioli 1-1.5 em longi robusti;
lamina 13-18 X 8.5-13 cm rigide coriacea integra late elliptica
apice late acuto basi obtusa, breviter (5-7 mm) triplinervata
(pare inframarginali debili neglecto) nervis secundariis 6-8 mm
inter se distantibus supra impressis subtus prominenter elevatis
nervulis supra obscuris subtus elevato-reticulatis, supra glabra.
Flores in ramulis infra folia fasciculati plerumque 4-meri;
pedicelli ca. 5 X 0.3 mm ebracteolati pilis clavatis 0.05 mm

longis appressis modice ati. Hypanthium (ad torum) 1.6 X

1.9-2 mm glabrum; calycis tubus 0.3-0.4 mm altus undulate (0.1-
0.3 mm) lobatus lobis remotis. Petala 2 X 2 mm mboidea

apice late acuto. na isomorphica glabra; filamenta 1.8 3
antherarum thecae 1.5 X 0.6 pice paulo emarginato poro
0.4 mm lato dorsaliter inclinato, connectivo dorsaliter ad bas
dente hebeti 0.2 lon ato lu xX 0. 4 glaber

in ovarii apice immersus; stigma truncatum 0.45 mm diam.; ovarium
4(2)-loculare 2/3-inferum apice conico 8-sulcato 0.5 mm alto
glabro.

Type Collection: Bassett rege & Celia K. Maguire 44011
(holotype NA 2444809; isotypes COL NY), collected on steep
slopes in scrub forest 30 km west of Cajamarca, Depto. Tolima
Colombia, elev. 3200 m, 18 Aug. 1959. "Stems short. vraquesti

Henriettella membranifolia has virtua leaf blades which
are sparsely strigulose above and wi h obvious linear erp’

5-merous South American analogue, H. sylvestris Gleason, ipschaaee
much thinner leaf blades sparsely strigulose above and with meso-
phyll inclusions, as well as petals 2.5-4 mm long. Crystal

inclusions in the leaves of H. maguirei certainly cannot be seen

superficially. Some of the material of the type collection ma
have been distributed under the collection number 44010 which is
actually Tibouchina paleacea (Triana) gn. Some doub fe)
exists ° correctness of the type locality; t

Cajamarca; Padre Uribe is attempting to recollect the species.

BOOK REVIEWS
Alma L, Moldenke

"In Gardens of Hawaii", by Marie C, Neal, new and revised edition,
xix+92h pp., illustr. Special Bulletin 50, Bishop Museum
Press, Honolulu, Hawaii. 1965. $15 .00

Ever since the author of this book joined the staff of the
Bishop Museum in 1920 she meticulously collected data on the orna-
mentals, fruits, vegetables, and weeds in local gardens, publish-
ing first in book form "In Honolulu Gardens" in 1928, revising
this in 1929, producing under the present title the expanded 1948
edition, and seeing on her deathbed the first copy of the present
work which has over a thousand more species described and over a
hundred more illustrations. This careful study covers 3,000
species in 1,20 genera and 20) families. Of these, 700 are il-
lustrated,

The plant descriptions make excellent word pictures, rendering
recognition and identification easy for both the educated amateur
and the professional botanist. Hawaiian lore and native names en-
rich the text and charm the reader. The scientific mames are

8
the work of the e eifted author herself. The 53-
page 3—columned index is replete

Since so few of the Hawaiian garden plants are native to the
islands, the value of the book is enhanced by giving the geograph-
many

ic origin range of these introductions from the food
plants brought in the long canoes from Tahiti for the original
Polynesian settlers to the pres and
hidden seeds on G, I. boo

This book forms a tion to Miss Neal's profession-

a superb
Wigabhcete etit Sank an a Fithlae wuektad te ter ites

The trustees of the Juliette M. Atherton Trust are to be con-
gratulated for their financial support in the printing of this

"Monograph of Cyrtandra (Gesneriaceae) on Oahu, Hawaiian Islands,"
by Harold a oh John, 465 pp., illustr. Museum Bulletin 229,
Bishop M eum Press, Honolulu, Hawaii. 1966. $12.00

This is a thorough 59h 4 by a recognized eacip road on a genus

that Stes Reompianene. 4) ement in the wet forests of Hawaiian

mountains, yet has proven difficult to collect ore to preserve,
81

82 PHYTOLOGIA Vol. 13, no. 2

The Ts covers 126 taxa, representing 11) 138 species, 7 vari-

a There are, "an addition, 2 putative etckae for
cohen The genus enus ranges from the Hawaiian an Islands through Malay-
sia.

This study was initiated in 1935 and 19% when the author sur
pin be the types preserved in European and Uni States

was terminated share after years of field study by the
pos cng his students

There are beautifully clear drawings and photographs of flower-
ing twigs and of herbarium sheets, presen nting ample details for

ably. The
since that of Hillebrand in 1888

"An ees ie ya Biology", by Dale S. Braungart and Ross
H. Arnett, second revised edition, xiv ; “\20 pp., illustr.
C. Vv. se Sg Siena St. Louis, Missouri. 1965. $8 250

It is satisfying to welcome this second edition to the college
classroom because it shows considerable improvement over the

and ()) the aera arranged by chapter hic Sones at the end
x ae is and carefully selected for both quantity

The most important new feature in the preparation of this
edition is the careful comparison of the subject matter eee the
culum Study te

1966 Moldenke, Book reviews 83

"Plant anatomy", by Katherine Esau, second revised edition, x *
767 pp., iliustr. John Wiley & Sons, Inc., New York i New
York, 1965. $14.50

Since 1g? the first edition of this magnificent work has been
available virtually the world over in botanical and university
libraries -- in seriously decreasing numbers — — mae gor and
professional workers as a reliable and thorough s k of

information, illustration and taniigNian sigs devel-

plant struc-

ture and structure range. This work was a pen and ink "crystal-
lization" of years of teaching and jintanehine this topic.

Since last year this second edition has been preg pr ts
too is magnificent. It covers the following topics in o
ell

3
truc
periderm, stem, leaf, root, flower, fruit, and seed. At the end
of each chapter there are listed detailed and updated references.
The indexes are full and carefully compiled. Many new and ex-
en

emp.
terms, especially in reference to the protoplast, cell wall
» and the i trnatienienaiiad: relationship in apical meri-

Because enthusiasm about the new discoveries on the molecular
ts newer workers to ignore the

author wisely cautions "A biologist, regardless of

specialization, cannot afford to lose

if his goal is the understanding of the organic world, Knowledg
of the grosser aspects of structure is tal for effective
teac and research in the more specialized areas of biology.
Moreover, the trend er a reduction of emphasis on factual in-
formation in modern teaching makes a readily accessible deposi-
tory of the basic fr ges on ak structure doubly important

rg jacket on "Plant Anatomy" claims correctly that it
"gives the student a spirit of discovery in his study of this
active field as well as continuing to provide a well documented
reference for plant scientists." Magnificent.

NOTES ON BROMELIACEAE, XXIII

Lyman B. Smith

VRIESEA

pb po Lindley, Botanical Register 29: pl. 10. 1843. Name con-
rved as "Vriesea" in the Incernational Code of Botanical
neliea iedick cf. Taxon 8: 232. 1959. pe: V. psittacina
Hooker) Lindley (Ti andsia 1 os .W. J. Hook-
° ‘Not Vriesia Hassk.
Hexalepis Raf. Fl. Tellur. “i 24. 1838.
cophyllum sensu Mez, Bull. Herb. Boiss. ser. II. 3: 131.
=1903, in greater part, not André 1889; cf. Smith & Pitten-
igh. Journ. Washington Acad. Sci. 43240121953.
Vriesea subgenus Alcantarea E. Morren ex Mez in Mart. Fl.
Brase%S, pt.°S2 516se1 .
Cipuropsis Ule, Verhandl. Bot. Ver. Brandenburg 48: 148. 1907;
mith & Pittendrigh; Journ. Washington Acad. Sci. 43: 40 04.

Neovriesia Britton in Britton & Wilson, Sci. Survey Porto Rico
5 35
Alcantarea (E. Morren ex Mez) Harms, Notizblatt 10: 802. 1929.

LEAVES always entire, rosulate; blades predominantly che add
and inconspicuously lepidote, often banded or ct ter SCAP
usually well developed. INFLORESCENCE simple or compoun und; ee
usually piriereghs tie phar often secund-flowe fe as well. FLO-
BRACTS conspicuous, enfolding or concealing the bases of the
flowers. FLOWERS mostly subsessile or with short thick eos
PETALS free or very short-connate, ligulate, bearin ng two scales
Mez's report of a single scale each in triligu tetute ta Mez

included or exserted. OVARY nearly or piece superior;
style Pangaee. OVULES many, usually Soe oe FRUIT capsular.
SEEDS fusiform with a long straight basal c
Mexico and the West Indies to Peru, Félivia, Paraguay, and
northern Argentina, with the greatest concentration of species
in eastern Brazil and the next largest in Costa Rica.

e present synopsis is written as much for the amateur as for

ey would seem self-evident to the latter. For the same reason,
arge number of old species are illustrated to facilitate the
Lapattet Wetted of those in cultivation
As indicated in the peptic sda above; there has been consi-
the cor

cd dropped the final "e" from the name of the Dutch botanist,
r. W. de Vriese, and gave it a Latin form by adding "ia" to the
8h

1966 Smith, Notes on Bromeliaceae 85

end. Later botanists, beginning with Beer, considered this gram-
matically incorrect a sed de Vriese's full family name plus a
inal "a". In 1906 in the first list of conserved generic names

ion. Each spelling has precedents t be interpreted in
its support or the matter would h been settled long ago, so
the question was raised again and the committee on th serva-—

tion of generic names gave a ruling in favor of the second spell-
ing, "Vriesea", with a definite clarification of its use. So
this nagging irritation is laid to rest and we can proceed to
more important matters.

e confusion about the limits of Vriesea is a much more seri-

botanists have tried desperately to find some other characters
that will correlate with the basic one o tal d An
h

Bh

tus as closely allied to Vriesea. In practice, the characte
aborted branches breaks down, and of André's two original spec
Thecophyllum wittmackii proves to be a Guzmania and T. poort

Oo
ies

petal-appendages to be a different kind of struct ose
of typi Vriesea, that is laterally produced auricles rather
than termi roduced scales. I ha tried to follow him in

s
cultivation, and Neovriesia un a ub-
lished, as He alepis was already available to Britton when he
followed eri Code in search of strict priority. Both
cases are technicalities that t amateur can ha ignore.

ea t a separate genus of Alcantarea on t -
Sis of apical as well as basal coma on the se and flaccid

petals is more logical than the previous examples, but pending

86 PHYITOLOGIA Vol. 13, no. 2

more evide it seems best not to change Mez's system unless a

guish species. The subdivisions of Vriesea may be summarized as

i. ene with the apical appendage minute and undivided; Jee
ixrm and remaining more or less erect after anthesi
sichieeeti " VRIESEA
2. oe exserted; . mostly —* colored with red,
ange, or yel , SI see SSE steht VRIESEA
ae avueatia tiotadeds eciete aaetly ‘ake green or —
ction IPE
1. Seed with the apical appendage divided into a ane
petals soon flaccid and drooping....----+ . Subgenus ALCANTAREA

far as information permits, the text of this paper is or-
dered in accordance with the above classification, but the posi-
i The key to

lated species. Cross references are given to several species of
Tillandsia whose ligulate leaf-blades tend to make them confused
with Vriesea when they lack petals.

ARTIFICIAL KEY TO SPECIES

1. Flowers in exactly 2 ranks but sometimes twisted until secund.
2. Flowers not secund except sometimes ——T as the petals are
exserted from the bracts on one s
3. Leaf-blades narrowly triangular, eriakiy eS apes
bke

4, Inflorescence compound......sssseeeeeecercrrecs ey I

4. Inflorescence simple. oe .....Subkey II

3. a ligulate ka sedans wet: tapering: ev weeds from
base to apex, usually glabrous or obscurely lepidote.

5s 1aslorescencd compound.
6. Upper scape-bracts equaling or exceeding the internodes.
Subkey II1
6. Upper scape-bracts shorter than the aa dans eee IV
5. Inflorescence simple
7. Scape straight or ascending: inflorescence erect.

1966 Smith, Notes on Bromeliaceae

87

8. Flowers oe erect, either imbricate or appressed to the
rhac ‘Ake seme oe Subkey V
8. Flowers sg hieek thes loueat OE not iebeteete. spread-
ng or divergent with evident space between
ie

Subkey VI
Scape decurved; inflorescence pendulous.........Subkey VII
2. Flowers secund at anthesis and after; floral bracts often se-
cund a 3; stamens almost always included.
9. A he — WMO Vikan ecw eee
9. Inflorescence simple
1. Flowers in mor

Subkey VIII
seeseessSubkey IX

than 2 ig gy BT ng or so few that

their relative position is not evident.
10. Inflorescence compound; stamens included...... ...Subkey X
10. Inflorescence simple; stamens mostly exserted......Subkey XI
SUBKEY I
1. Flowers imbric
2

Spikes much pent than the primary bracts; sepals not over
25 mm eee: leaf-sheaths dark castaneous or ferruginous.
3. Floral bra
4. Sepals ee the floral bracts Ecuador, Peru.
1. V. bitchcockians
4. Sepals equaling or shorter than the floral brac
5. Spikes

b
subterete, very narrow; floral bracts 27 mm long.
ae ee ee Oe eru..

< ¥. reuhit
as Sethan strongly chbiniatat ‘£tpral bracts 35- “50 mm
6. Floral bracts 35 mm long, n

long.
erve eru,..3..¥. cereicola
6. reat bracts 40-50 mm long, nearly even.

Venezuela,
Ombi as... os sikae nataneviets. Vourobusta
S. Floral eRe carina
7. Sepals 23 mm

long; | SERIE pendent. Pl. I, figs.
ss POPUL... Seis eis eee ene + . V. olmosana
7. Sepals 8-
8. Spikes panes goad sxcandlnn ih digitate; petals denticu-
ate.
9. Scape little if any ti than the spikes; pete pa
green. Nicarag To

tee yee OA. Vs Seis
9. Scape much Lnaee than i meine petals wh te
Colombi ,65.--V- erenulipetala
8. Spikes nearly all ger pay PA “spreading abruptly from
on

the
erect sterile base; infloresce
10. Inflorescence tripinnate;
f

spikes on hg = the «
of the branches. Ecuador.........66. V. chi aut ame
10. Inflorescence not more than bipin

pinnate.
11. Floral bracts distinctly and densely lepidote; spikes
oblong o

many-f lowered
Venezuela to Bolivia.
67. V. heterandra
» 7 cm long; spikes feu-ficwaced
naked sterile bases. Venezuela...68. V.

ax
bracteate sterile bases.

12. Inflorescence dense
with

lacera

88 PHYTOLOGIA Vol. 13, no. 2

Et: aan sa nies obscurely lepidote, appearing glabrous;
spikes lanceolate; pe neo ett early or quite entire
fi 5

lilac c or white. Pl. ig. 5. Mexico, West Indies,
nezuela, Colombia re 69. V. didistichoides
the

2. Neate fe least the lowest) less ae twice as long as
imary bracts; sepals 8-48 mm lon
13; Sepala 8-10 mm long. Pl. I, fig. a Colombia, Ecuador
70. V. fragrans
13. Sepals 22-48 mm long
4. Floral bracts mm ceene: ee gia ee densely 4-5-
OPS vr tis betes tee eee web es 71. V. drewii

15. Inflorescence erec
16. Floral bracts nearly all convex and ecarinate, 24 mm
g 1. I, fig. 7. Colombia, Ecua dor.
6. V. cylindrica
16. Floral bracts carinate.
17. Spikes many-flowered, much exserted above the primary
bracts; floral og 30-40 mm long, diffusely lepi-
dote. I, PETU.cccccecees V. harmsiana
17. ne ne fev-ftovered, almost wholly Severed by the pri-
mary bracts; floral bracts 23 mm lon ~ densely lepi-
dot PL. Ls igh 9. Peru......8. V. tillandsioides
15. Inf iprastehbe pendulous.
To: hae 1 bracts much exceeding the sepals, 40 mm long.
UT: ts) geeerer a: © ore ne ea arr ia V. arpocalyx
18. Floral bracts equaling or " shorter than the ; gepale; 22 mm
ong. Venezuela to Peru......+++-+-> 10. V. tequendamae
1. Flowers lax
19. Floral bikeote much poroanpR i the piesa 35 mm long.
i.

20. i phe wach shorter than the spreading branches of
the broad lax inflorescence; floral i x 18-25 mm
Ly 1

long,
about equaling the sepals. Pl.

12. V. lubbersii

—

20. Primary bracts much exceeding the short strict branches of

the slenderly cylindric inflorescence; floral bracts 8-10
mm g, one-half or one-third as ern as sas sepals.
PL SCE Pie Bri eis ....72, V. drepanocarpa

SUBKEY II

1. Floral bracts much shorter than the sepals, a
suborbicular; inflorescence lax. Brazil..73. V. goniorachis
1. Floral bracts equaling to much exceeding the gupals:
cence dense or subdense.
2. Leaf-blades with brown sSrisaeetcagsnty inflorescence 3-flowered.
4.

inflores-

Chath Mie ok sek nh nce esse ees 74. V. jimenezii
2. Leaf-blades dobedier rous 3 tht Toresewice ‘rexcnany-flovered
3. Leaf-blades green, Obecuiilly lepidote

1966

Smith, Notes on Bromeliaceae 89
4. sg ie onsig to 50 mm ind much exceeding the sepal
MG LA vre-wrttwerstelecialew am bs ae Oa biRGle cee © ere - V. sulcata
4, fiseal "i Padte 25. mm. n long, Moth E equaling the se
Brazi

pals
sisal’ Nie Slate, ole Guat, SMS asf crtieekants
as ime 26 1ades! ease eee from lete covering of scal

a es
a5 ee broadly winged and shea lading the bases of the

e. Gir ivancanes complanate, 35 mm wide.

Ecuador
14. v. barclayana
6. Inflorescence terete, 7 mm in diameter. Peru.
3. V. cereicola
5. Rhachis not more than

angled.
T ed ae bracts convex and ecarinate, 45 mm long; stamens
erted. Ecuador, Peru... eooeelLd. Ve. patula
i Floral bracts carinate, to 16 mm long; ct wes included.
Nicaragua to Peru....

cescewseee- 64. V. chontalensis
SUBKEY III

1. Flowers all erect and contiguous at anthesis

vie Sevin prette black, fleshy; sterile ace oF the branche
mot Brazil. wveccccccvevcsecsvecessssece V. funebris

rap Floral be ler

3. Sepals exserted ahs the floral bracts

by the primary br fact’ sepals 10-16 mm

11 others oa sod te ieing the ortnary
racts; sepals ca. 40 mm zil.
Fath ate sterile bases of
t ranches; floral bracts errs carinate, 35 m
lo AES eaNeen . V. gravisiana
5. Primary bract équaling ¢ he short “sterile base of the late-
ral branch; floral bracts ecarinate, less than 25 mm
long. ee faa sincorana
3. Sepals cour By the Stovet lgvdecs
6. Scape very short and hidden by tie leaves; spikes linear,
45-60 cm long. British Guiana..... . V. soderstromii
6. Scape well developed.
7. Leaf-blades marked with dark wavy cross bands or spots.
8. Floral bracts 30 mm long, lustrous (cf. Tillandsia

oezlii).
8. Floral bracts 55-60 mm long, subchartaceous and nerved
when dry. Pl. I, figs. 13, 14. Trini

78. V. glutinosa
7. Leaf-blades concolorous

9. Inflorescence eripinnate at least at bas
10. Floral bracts broadly convex toward sbi and weakly if
at all carinate; sepals 9-40 mm long (cf. Tillandsia
boliviana, denudata, excelsa, fendleri, guatema-
platyphylla, rubella, selleana,

10. risa bracts sharply carinate toward apex.

PHYTOLOGIA Vol. 13, no, 2

Lhe viene} — to 33 mm long, rapidly decreasing in
siz ard (cf. Tillandsia stenoura

1hi a heat ae 17-25 mm long, gradually dacveas ing in
size upward.

12. Posterior sepals carinate (cf.
uidae, krukoffiana
32% peanemict and anterior ‘eqpale all ecarinate.
b

.79. V. elata
$5 Jat lorescence ir more _ bipinnate.
13. Floral
1

Tillandsia acuminata,

Colom-

oral bracts ecarina

Inflorescence oe or subdigitate from a few
spikes.

15. Leaf-sheaths densely ferruginous-lepidote, strongly
mtrasting with the blades; sepals to 15 mm long.
yD Pe Giese a Bo 7a So eialaal Colombia, Peru
80. V. chrysostachys
15. Leaf-sheaths — or purplish, not pene es
much with the blades; sepals 24-40 m ge (ck,
Tillandsia fendleri a ), grandis, 1 pote
14. Inflorescence 1
16. Sepals 12-17 mm

g
17. Floral bracts prominently nerved,
Bees oe

rather thin; se-
long (cf. Tillandsia rubella,

a ae tional eee even, coriaceous; epee to 17 mm
long. Costa Rica....+.++++e+s ge camptoclada
er Sepals 24-40 mm ee
8. Floral bracts sig gp broadly subtruncate,
ong, narrow and exposing most of the a -
ie Pade sie ere na nee ae ame .82. V. hodgei
18. Floral egy elliptic or ovate, broadest at. or. be=
ee middle (cf. =e
» grandis, rauhii).
13. ploxek os " sharply pac ate toward a
19. Primary bracts equaling or exceeding he lower spikes
(cf. Tillandsia spp. of West Indies, Mexico, Central
America, and Andean South Ameri ca).
ry bracts much shorter than all of the spikes.
20. Leaf-blades densely and conspicuously cinereous—

andsia carnosa, denudata,

19. Prima

Tillandsia arguta,
us s, kuntzeana, oerstediana - cf.
n "Mi

areas
20. Leaf-blades obscurely lepidote to glabr

21. Leaf-blades attenuate at apex but hiveiee ligulate
22. Spikes reflexed (cf. Tillandsia somnians somnians) .
22. Spikes erect or subere
23. Floral bracts 40-45 mm pg spikes long-stipitate
(cf. Tillandsia deppean aM:
23. Floral bracts 20-2

lon
24. Firs bracts oblong, ceva (cf,

Tillandsia
24. er bracts elliptic, even. Peru.

1966 Smith, Notes on Bromeliaceae 91
83

- V. subandina
21. Leaf-blades acute or rounded and apicula

25. Floral eae 40-50 mm long (cf. Tillandsia hetero-
h

phyll ralensis, Sigmoidea).
25. Floral rere ong.

26. Floral eter narrow eas revealing the alate rha-

his at anthes

27. a opoee 7 cm lon ig #loral bracts nerved (cf.
Tillandsia zamorensi

27. Spikes to 25 cm

cm long; Clee bracts even. Trini-
orthern South America.

Fs 84. rubra
26. Floral: eects broad, but someting shares and
revealing the rhachis
28. Floral bracts nortecsaas even (cf. Tillandsia
aequatorialis, denudata, pyramidata, reversa,
perba).

on no sterile
base; floral bracts apace De lepi-
an toward apex, 27-32 mm rg fe Pa.

ide BON CEs ve ted Vv. Siciiveins
30. pe 6- Sf lowered with 1-5 sterile bracts at

ase; floral bracts sparsely lepidote
hr

oughout, 35 mm long........19. V. pinottii
1. Flowers (or at least the lowest) lax, more or less spreading.
Jig Percise slender, 1-2 cm

ong, exceeding the floral bracts.

esd

° . Ne otece bats
i mededate stout, Séubily dab Less than 1 cm long, alway
much shorter than the floral brac

32. Primary bracts equaling or exceedi se "*ebe lowest branches.
33. Sepals acute, all ecarinate, maximally 20-

on,
Come ene ebeesoteereserecuvetesss V. peeckrun
33. ote obtuse to gi otragee or vetuss: neekinate (poste-
» maximally 9-20 mm
34. Sepals’ pean eter witase leptdote with rather coarse
free-margined scales Mela. coo0ss. 87s (V. egregia
34. Sepals cbeGad or sub

ae isteinad or with obscure
tiform scales (cf. Tillandsia biflora, ionochroma,
st take} wangerinii).

32. Primary bracts all shorter than eee spyesecs branches.
o lax

‘:
oe pits bracts exceeding the sepals (cf. Tillandsia
thyrsigera).
Shs nies bracts shorter than the eure Piet,
Bra CEP S CE Bee eee Bes is 1

36. Seni ins not over 24 mm long.

38. Inflorescence beiaiim nate; floral bracts nerved.

39. Floral bracts much shorter than the sepals, ecarinate.

1
haces diets

92 PHYTOLOGIA Vol. 13, no. 2

OMDLA. sce e cece serene rvcecessecess V. hospitalis
39. ade bracts about equaling to aia chster ly - exceeding
epals, carinate.
40. Spikes meat ovate, subdense. Colombia.
. V. magdalenae

40. Spikes elongate, very lax (cf. Tillandsia | delicatula).
38. Shas a not more than bipinnate.
41. Sepals 9-10 mm long; floral agentes strongly nerved (cf.

Tillandsia singularis, trunc

41. gig be 16-24 mm long; floral aaa faintly nerved to

42. siakal bracts about 5 times as long as the internodes,
conduplicate, sharply carinate (cf. Tillandsia

carrierei) .
42. Floral bracts only 2-3 times as long as inter-
nodes, eee faintly if at all carin
ong, about equaling the oblong-elliptic
oral bracts; pedicels to 8 mm long. Cost

XV.
43. Sepals 17 mm long, much exceeding the muborbicular
floral bracts; pedicels short.
91. V. thyrsoidea
35. Inflorescence broad, lax
44, at al bracts 7- 17. mm Long, ovate to reniform, mostly
overing only about half of each sepa
45. poral bracts ovate or elliptic, dtetinctly longer than
Ww cf. Tillandsia asplundii, laxissima, pinnata
venusta).
45. mae eh suborbicular to reniform. i.
46. Scape mesh rved; inflorescence niger: floral bra
to Bicede cr ceve vans .21. V. billbergioides
46. Scape ry inflsetanctace erect.
47. paypricpeots es tripinnate with its lower branches divi-
and recurving; floral bracts 7 mm lon
2. Y. leptanthe
47. Inflorescence bipinnate; floral bracts 10-17 mm long
sharply carinate. Pl. I, fig ‘

23. V. rodigasiana
44. Floral bracts 20-50 mm lon

48. Rhachis of the spike elender. about 2 mm in diameter.

ong; lower primary bracts subfoliaceous.
Brazel 6355. Gk. ee ced. CEC ORL es MERE S 86. V. sceptrum
SO. Sepals abortet than — floral bracts; primary bracts
ot at all foliaceo
SLs Prawits sessile (cf. reitandeda dyeriana, hanaleana) -
mm

seeeeveeeeeeee

single bract. Pl. I, f

schwackeana

schwackes*=

52. Sterile bases of the branches elongate with several

Smith, Notes on Bromeliaceae 93

Brazil. 24, , Brusquensis
49. fat "baa nearly o or natin ‘straight ae apex,
often ecarinate.
53. Leaf-blades acumina
54. Floral bracts suberect, about 3 times the aeecir ab
exceeding the sepals; sepals 17 mm long, short
e (cf. Tillandsia maculata).
i 1-2 times the paint
shorter than the sepals; sepals 26-29 mm long.
55. Scape-bracts apiculate, the upper ones much reduced;
amens equaling or exceeding the — Brazil.
2 neoglutinosa
55. Scape-bracts acuminate, all large et subfoliaceous;
Stamens included. Pl. I, fig. 1

Vv. “altodaserrae

53. Leaf-blades more or less rounded and ipicdhar

6. Sepals acute; leaf-sheaths purple. Brazil.
57. Sterile bases of the branches bracteate.

- V. muelleri

57. Sterile bases of the branches naked
V. haematina
56. Sepals obtuse; leaf-sheaths pale green throughout or
castaneous toward base, sometimes red-spott
58. Leaf-blades densely cinereous- -lepidote, nator tted.
figs. 20, 21. Brazil.....27. V. saundersii

their
senaeoe about 4 times as — as the internodes.
ceeeeee94, V. monacorum
59% a brace ‘slightly £$: Me all carinate, mostly
much less than 4 times as long as the internodes.
r

fig. 17. Brazil, Paraguay, Argentina
2

0. V. friburgensis
60. ee bracts shorter than the long bracteate
rile bases of the branches. Pl. I, fig. 22.

ad, Guiana, Brazil..........28. V. procera

nid
48. eats oe 9 spike stout, much more than 2 mm in

er.

61. Floral bracts tee ee acute to obtuse or truncate,

y ecarina

62. Bhachis aerate: geen bracts about as long as the

rnodes (cf. Tel andsin rariflora).

62. Rhachis flexuous to geniculate; floral bracts much

r than the interno ae
63. Bhachets broadly alate with relatively short inter-
is).

64. Sepals less than 20 mm long.
65. hapa bracts and sepals coptoneind nerved; leaf-apex
ft ce. Tiliandeia platyphylla).

94 PHYTOLOGIA Vol. 13, no. 2

65. Floral bracts and sepals even; leaf ending wi
hard acuminate cusp. Colombia..95. V. pipiens
64. Sepals 30-50 mm long.
66. Floral bracts suborbicular, strongly gibbou
67. Flowers to 8 cm long; rhachis genicu ulate. Brazil.
218. V. extensa
67. Flowers to 12.5 cm long; rhachis unknown. Pls ks
fi Panama, Jamaica (?).......219. V. alta
66. Wipeal Sena narrower, mostly eceakoht along the
apical half.
68. Rhachis geniculate. Pl. I, fig. 2%. Brazil.
220. V. geniculata

68. Rhachis merely flexuous.
69. Floral bracts about as as as the aoa -
pull. & ts razil..96. gigantea
69. Floral “bracts 2- Bit get in arian Fg "glaucous.
(cf. seyeaelye viridiflora).

ex
61. shapes bracts. acute. carinate; petals flaccid after
is. zil.

Bra
70. Rhachis ieee 1fatelL eerie floral bracts
not more than twice as long as the internodes at
esis. Pl. Il, fig. 1..--... ers V. regina

70. Rhachis strongly geniculate; floral bracts goes ose
ice as long as the internodes. Fi. fig.
222. V annir

SUBKEY IV

1. Inflorescence subcorymbose, only 45 mm long, its primary
“ ag involucrate below it; sepals 22 mm long, much exceed-

e floral bracts. Brazil.....-++++++++- .97. V. paradoxa
L. ip opstiieoas elongate, much cahbaiing: its oacgea saa
2. Upper scape-bracts imbricate, only the lower
than the iuneennes tcf: Tillandsia ferertty preesere

laxissima, trunc
2. Upper scape-bracts oe than the internodes and usually
e others also.
3. Branches of the inflorescence 2-3 cm long. Costa Ric
4. Sepals 21 mm long; pedicels 6 mm long......98. V. Si dieeing
4. Sepals 10 mm long; pedicels 11 mm long....-- 99. V. turbinata
. Branches of the inflorescence much more than 3 cm long.
5. Scape decurved; inflorescence pendulous; floral bracts
suborbicular to broadly o — much shorter than th
sepals, membranaceous. azi
6. Axis of the inflorescence “geniculate; ere rounded.
D5 Ne billbergioides
6. Axis of the inflorescence nearly serait ee
SubOD CUBE: kbc ab none woes n wes pr ae i. . languida
5. Scape straight or gacund ings iitinceacence.&
7. Primary bracts much exceeding the short aa “naked
bases of the branches
8. Floral bracts crap equaling to exceeding the sepals.

1966 Smith, Notes on Bromeliaceae 95

9. Inflorescence 3- ae ee sepals 17 mm long (cf. Til-
andsia maculata).
9 Litaarescaed ope eee sepals 24 mm long. Brazil.
101. V. a
8. Floral bracts shorter than the sepals.
20. tah ae dense, its branches erect; bracts and
als castaneous. Brazi oc, | Se ene. V. maculosa
10. Inflorescence lax, its branches en ge to . spreading;
bracts and sepals yellow or greeni
a eS Floral. bracts elliptic, ecarinate, Sut little shorter
than the sepals (cf. Tillandsia venusta).
1l. Floral bracts suborbicular, carinate, about half as
mg as the sepals. Pl. I, fig. 18. Brazil.
23. V. rodigasiana
7. Primary bracts shorter than ig long, usually bracteate,
sterile bases of the branch
12. Sepals 35-50 mm long; flo a bracts ame below the
apex, strongly rugose on dryin zit;

18. V. extensa
12. Sepals not more than 29 mm lon
if, seth bracts about equaling the internodes or slightly
more; branches usually spreading
14. Leaves acuminate; stamens Gesectads Brazil.
5. V. neoglutinosa
14, bank gee broadly eg or rounded and apiculate; stamens
ncludéd, Pl. 1, fig: 2225 Trinidad, :Guiana, Brazil
28. V. procera
13. Floral bracts much exceeding the internodes; branches
erect or ascending. Braz
15. Floral bracts tuthedis avbhdte: branches subequal.
. V, stricta
15. Floral bracts ecarinate; the terminal branch much
larger and with a longer sterile base than ch
lateral Ones fii ceie cee oe eevccteceee IO4,. V. minarum

SUBKEY V
1. 2 << the highest or lowest) exceeding the broad floral
bra

2. Fidzai | bracts coriaceous, even, white-lepidote (cf. Tilland-

sia excavata).
2. Floral ceares rather thin, nerved, nearly or quite glabrous.
3. Inflorescence many-flowered, to em long, exceeding the
mitoe floral bracts vane or not at all carinate.

azil.. ; 29. V. gradata
3. erstualecs ta-fto meted ‘6-1 12 cm long, about equaling the
leaves; floral bracts sharply carinate. Brazil

30. V. modesta

1. Sepals equaling the floral bracts or short

4. Floral bracts carinate so far as visible, uit or brightly
colored, green, yellow, or red.

te teehee Rae or kOe SER MOMES Es oe web CR eee bee eee e ee

96 PHYTOLOGIA Vol. 13, no. 2
5. Apical half of the floral bract spreading and not imbricate
even before anthesis; keel of the floral bracts stra ight
meave toward the base, convex toward the apex; inflo-
rescence broadly rounded at apex
6. Floral bracts acuminate, 60-70 mm. long.

eeeee

bas
raz RS Series Ch as ara «as cxythrodactyion
6. Floral rapes beneiiy acute, abate 45 mm long. i Be
= _ oa mtral America and Colombia to aps a
5, eet yee
im

as ; (mettaontbides
(as well as rhe einindas) of aa floral bract
ibricate unt after anthesis; keel of the floral bracts
y convex; inflorescence usually acute.
7. Floral bracts densely lepidote
8. Indument of the floral bracts red-brown; floral bracts so
ee as to be nearly horizontal.
azil, ("Ecuador

Pls Eigcf£te.5.
SPSL Ere D Sse ie ee 32. V. barilletii
8. ae of the floral branded vingwaness a bracts
much less crowded, suberect.

Brazil..33. V. vulpinoidea
7. Floral —— glabrous or very sparsely and FP y
lepidot

9. Margins of the floral bracts with a strong almost semicir-
cular curve.
10; infderacseucie several in each rosette, lateral; scape-
racts distichous; posterior sepals carinate (cf. Til-
a multicaulis).
10. Inflorescence single, ate scape-bracts polysti-
chous; sepals ecarinate.

i

11. Floral bracts with obtusely siaded prince bright red
with broad yellow margins, 45-60 mm
cence ee: to linear,

inflores-
15-40 cm tong Pl. TT; "fig.
ee re rere ek eee ee . V. incurvata
Lb Bloral. bracts swith ev week curved margin -

=z
»

mm
scence usually uencent ata? or elliptic.
very d st ack floral bract more than
covered by the e below it, red with yy ta or
ree margins, its hat slightly curved.
Cite Leg oteork.: dienes. TOs SHELLS. wiles She. Be "inflata
12. Inflorescence cdenke: each floral bract less than

half covered by the one below it, its keel strongly
curved.
13. Sepale 38 mm long. Pl. 11; fig. 8.

36. V. petropolitana
gale 27 wi Mondor Sioee cease sere. — V. pinottii
9. sss rat the Sinker bracts Aightiy curv’

14. goo eho 6-13 cm

ong, 2-3 times as aoe as the se-
holly green or

red.
5; siarst orans attenuate, straight, 11-13 cm long; pos
erior

eeenee

1966 Smith, Notes on Bromeliaceae 97

14, Floral bracts not over 5 cm lo ong, about twice as long as
he carinate sepals at most, mostly bicolorous.
16. Inflorescence subquadrate, almost as broad as cone
scape very slender. Pl. II, fig. 11.
38. V. carinatn
16. Inflorescence much longer than broad; scape ease
mm wide

17. Side of the floral bract about 3 mm long,

17. Side of the floral bract 10-15 mm wide, 40-50 mm long.

18. Scape elongate, exceeding the lea aves; plant flowering
5-8 dm high; floral bracts coriaceous (cf. Tilland-
Sia heterophylla).

18. Scape not exceeding the leaves; plant flowering 2.5-5
dm high.

g
19. Inflorescence much exceeding the leaves, 16-26 cm
ong, much more an 3 times as long as broad;
floral bracts 45-50 mm long; sepals narrowly el-
liptic, obtuse or emarginate. Pl. II S. tigtei2.
39. ‘V. duvaliana
19. Inflorescence barely or not exceeding the leaves, 11
ng, much less than 3 times as long as broad;
floral bracts 40 mm long; sepals lance-ovate,
acuteseoPly Ils figs hemes peetinicl paraibica
4. Floral bracts ecarinate or nearly s
20. es bracts bicolorous, dark see seach and coriaceous
cept for the pale soft apex, 3-8 cm long. Brazil
06. V. longicaulis
20. Floral bracts uniform or near y so
21. Floral bracts appearing rugose, at least under a lens.
22. Floral bracts thin or subpapyraceous when dry.
23. asepyrbeacts broadly ovate; sepals ere elliptic, 24
ong. Brazil. 1. V. rhodostachys
23% i ak ot Peasy + c subfoliaceous; | sepals broadly
oblong-elliptic, 18 mm long. emala.
107. V. pycnantha
22. Floral bracts thick, rigid, coriaceous.
24, Leaves broadly acute or obtuse, apiculate.
25. Floral bracts barely imbricate. Costa Rica.
V. rugosa
25. Floral bracts 3 to 4 times as long as the internodes.
l. Il, fig. 14. Guatemala to Colombia
109. V. gladioliflora
24. Leaves acute, not apiculate. Costa Rica.
26. Floral bracts broadly acute, much exceeding the sepals.
110. V. pachyspatha
26. Floral bracts obtuse, barely exceeding the sepals
ill. V. macrantha
21. Floral bracts even or longitudinally striate, not rugose.
27. Flowers reflexed at anthesis but the floral bracts often
still imbricate.. Pl. II, fig. 15. Mexico.
112. V. ovandensis
27. Flowers not reflexed.

98 PHYTOLOGIA Vol. 13, m.
28. —— bracts densely imbricate and concealing the
anid o

no. haat 10-15 mm long; scapes very slender, numerous,
ateral (cf. Tillandsia eiskp haiti) 2

29. pusett oe 40 mm long; scapes rather stout, single,
cen
a0. Seaperbracts peoreceneh lowest floral bracts long-

acuminate. Lé8e 40s cae ..--113. V. acuminata

30. seape-bracts Salis iform; of the floral bracts

nate (cf. vitthatite opeaatant? glossophylla,

eri).
28 wiauel are inrolled re the sepals and exposing the
rhachis or barely imbricat
long; anos hinted very narrowly line-
bia 114.

bd
r, acuminate. Colombia......--- V. schultes*an
31. Sepals 15-35 mm long; leaf-blades ligulate, acute
subobtuse and apiculate.

32. Sepals 15 mm lo — sis bracts 22 ie long; leaves

acute. Costa Rica.......s+se0s : . V. brachyphylla
32. Sepals to 35 mm oad floral bracts Hee 55 mm long;

aves broadly acute or subobtuse and apic ulate.
33. Floral bracts even; sepals erent broad
acute. Costa Rica... ..eeeseececes .116. V. brunei
33. — bracts Se 5 nerveds sepals 3655

sili 42. V

ng, rounded

Poe ere wey ye ea Ee . . pauciflora
SUBKEY VI
de _ merely divergent at anthesis
2. Sepa Sire es long; leaf-blades narrowly linear, acumin
Voces Seer eees cece eessoues +114. V. sineltec tant
2. Sepals 1s Aa! te

. Scape-bracts oa equaling or exceeding the internodes.
4. Sepals 22-28 mm

5. Floral bracts eee the sepals
6. 9 Sarees ovate; floral Beats ecarinate. P

dacItl
4,2.) Costa Rica. . os sees yeudsLb? (We apiculate
6. Senkts "g mm wide; floral peacts carinate. Brazil.
Vy. biguassuensis
5. Floral bracts shorter than the sepals. Venezuela.
44. V. laxa
4. Sepals 35-40 mm lon

g-
7. Floral bracts quore than the sepals, ecarinate (cf.

Costa
cide kud nenbs bs ese et eeune see rugosa
8. BE mes bracts faintly ferved; “about equaling the . sepals.
Bolivia, Argéntinas 69s. cs.sisvces ..---45. V. maxoniana
Ki moos SP ate: sho

rter thas: the upper ssh te floral
much shorter than the sepals.

Brazil
9 Pediat slender, 8 mm long; sepals subacute, "25- 30 mm long

1966 Smith, Notes on Bromeliaceae 99
Ba Sy Ween ap eee 2c Weer Nae ---46. V. amethystina
ng -

acs
9. Pedicels very short or sates poten: 3 gos ~22 mm lon
10. Leaf-blades ET eRe, ope almost the same lengt
the sheaths, 35 mm w 18. V. lancifolia
10. Leaf-blades linear, several tinas ‘Sacer than the sheaths
mm wide. .... +. seeeeeeell9, V. parviflora
1. Flowers spreading oe maf bened a santhanab
11. Floral bracts with truncate bases, usually bright red or
yellow, sometimes pale green.
n Bias saree pages from about the same length as the internodes
to as long, from slightly shorter than the sepals
to erty them; rhachis slender, flexuous, its spe ie
nodes more or less curved; inflorescence few-flowered.
Brazil.
Sepals 35-40 mm long; floral bracts about twice as a as
the internodes; scape-bracts all imbri icate; stam

=
lo

-
eo

te]
ternodes in ae variety with a simple pat Aor arene
Stamens included. «iss ss. sia) nies 00's - procera
12. Floral bracts mostly 3 times as long as the. interapdes: > or
Fe rhachis stout, straight or geniculate with straight
nternodes; inflorescence usually many-flower
14, et bracts recurved toward the apex and with undulate
n to 55 mm long, enfolding and exceeding
the sepals. Pl. III, fig. 5. Brazil....48. V. recurvata
14. Floral bracts straight or incurved toward apex
15. Floral bracts sharply carinate, the keel incurved toward
the apex, nerved; sepals 20-30 mm long.
16. Rhachis straight or nearly so; flowers sessile (cf.
Tillandsia dyeriana).
16. Rhachis more or less geniculate; flowers on short stout

oral bracts cuborbicular, rather
thin, only the lowest spreading and not imbricate.

29. V. gradata
17. Sepals included; floral bracts narrower, coriaceous,
Spreading with age. .Pis.1Il, fig..7.
50. V. ensiformis
1l. Floral bracts with decurrent auricles at base, usually dull
een; eae included so far as known.
19 cy eee BER ene floral bracts about equaling the
pals. Cuba, Puerto Rico, Venezuela
1 V. macrostachya
19. Leaf-blades marked with spots or lines or both to —

ral bracts much shorter than the sepals.
9. Brazil sseesi2Zl. Y. fenestralis
long

figs
18, hesite 17- 35 oe

100 PHYTOLOGIA Vol. 13, no. 2

20. Floral bracts smooth and even when dry, coriaceous, not at
fleshy.

21. Floral sna dark castaneous with pale margins, from

s tly shorter ‘s gine ie i than the sepals.

RP ote or hie] be ‘Y. atra

2. Sepals very broad,

ce,
acute or subacute. Pl. fig. 1. Trittidad, Brazil
23. V. jonghei
23. ps0 titan shorter than the upper internodes; leaves

ersely marked with purple lines;

flowers re-
fle nets at

anthesis ; sepals broadly rhs Aal obtuse.
Wa. 2 3. Brasiis 6. eee VN fosteriana
22.6 Sepals nserowly elliptic or suboblong, much "longer t
b about half exserted; flowers usually yorthory
eae ee oe

= oR est ae eas-

vee - platynema

e or nerved whist a fleshy

or subch e

24. san bracts fleshy, becoming sabia when dry; rhachis
sto

2% sepals acute, to 35 mm long; floral bracts obtuse

ely
Ble. Brass, 4 se ce ee se ws V. wawranea
20% Riser bey subobtuse, 25-44 mm long; Flovel Meats convex,

ecarinate.

26. Inflorescence covered with a strongly cin he mate-
61; ¢

° te before
satet usually with dark nag es ah tel TY, fig. 6.
2nele, Bragile: vias ees bi tuminoss
26, Inflorescence dry; floral bracts eee nev
br » concolorous. A354 126. Ve tei ellii
24. Floral bracts Coteus
coriac

oming nerved Paks: dry, subchartaceous oF

27. Sepals covered ed the floral bracts; floral bracts about
4 times ong as the

internodes. Pl. te #¢ 15.
Mexico. cee phe ovandensis
275 — exeerté ed for geet. halt th air length Man the
abs

sriempa Fagen bracts only about 2 times as

odes
25. Scene bracts aeiaets imbricate; Beige stout; sepals
: mm long. Pl. West Indies,
ern h America --.-125. V. platynema
28. Scape-bracts slightly shovter:’ thai the n
nodes; pedicels rather slender; rir to 22 mm long.
hat. . Cea cee as es che eccs caus. 4. Bae

———

CHOCO HOEHSEHEHOHERESH HHH HEE HHS HOHH HHO SEH ESE HERES HEHBEORESDEEHEEHEH®F

1966 Smith, Notes on Bromeliacese 101

SUBKEY VII

1. Floral bracts densely verrucose. Cost MeSH ai 5 Vv. nutans
1. Floral bracts even or nerved but not verrucos
2. Leaf-blades marked on the upper or both s ides "ad ar
spots, broadly rounded; floral bracts farinaceous, a 40 mm
long. Brazi
3. Floral bracts abhi okies and concealing the rhachis, broader
than long, equaling the sepals. Pl. e 8.
sd ares, pardalina
3. Floral bracts ee the spreading flowers and thus ex
posing the rhachis, broadl ly ovate, tome or shorter
Vv s. 9

an the sepals. Pl. IV, ; oe 52. V. guttata
2. Leaf-blades concolorous, pa
4. Floral bracts er oe

ing
5. Scapes Many, lateral: floral bracts 15-25 mm long (cf. Til-
landsia compl
5 — ome anh floral bracts 50-55 m oa
Be are Oe te -33. V. obliqua
4. Etiral salones spreatiting’ and exposing this ht at anthesis
6. Floral bracts laterally compressed, sh ly carinate,
i o 40 mm

sepals. Pl. IV, fig. 11. Brazil...... -54. V. retroflexa
6. = bracts not compressed, slightly if at all carin ate,
usually much shorter than the sepals; pedicels slender,
8-20 mm long.
7. Floral bracts much more than half as wide as long, 35-45
mm long, their margins much overlapping behind the se-
9 9 PLeTV, C18 12 THa idad, Venezuela, eee

e114.<2 ear ig oe as Me camden

7. Floral beacie® nde? more er half as bios as ‘ie ot over
5 mm long, their margins miveirr toa J little if a all.

PL. DV, eiget 13 18S he ---.96. V. scalaris

SUBKEY VIII

2. Scape-bracts shorter than the upper internodes (unknown in y.
imii

3. Meliovicesohes dative or subdense; branches erect or suberect;
a al bracts mostly more than ‘tw ice as long as the inter-

S; sepals 22-24 mm long. Brazil.

vine ie dark renee | Bs ae half as long as

e internodes. seeeeeeee +130. V. crassa

4, hier care aistovddes ‘Sag a little ghavter than
the internode Seeeeecescceceeveevenseesel3l, V. densiflora

4. Lear

PHYTOLOGIA Vol. 13, no. 2

3. Inflorescence lax; branches more or less spreadin
5. Floral bracts imbricate; scape unknown (cf. ritigaiiie

cucullata).
5. Floral bracts less than twice as long as the internodes.
6. Sepals equaling the floral bracts,

to 23 mm long. ba.

132. V. dissitiflora
6. Sepals exceeding the floral bra
7. Pedicels 1-2 cm long, Sadie we were bracts, very
slender; sepals 7-11 mm long. Costa Rica
. V. pedicellata
qT. pestis shorter and stouter, much BE ee by the floral
bra

ranean 20-flowered, to 55 cm long; sepals 15-20 mm
long. Pl. IV, figs. 15-17. Hispan niola.

3. V. tuerckheimii
8. Branches 2- oe sometts to 18 cm long with the sterile
part much the lon
9. Sterile — of he es bracteate; sepals 1
long. Zit. isco 14s: Ve ae cei tiore
9. Sterile phe: oF the ‘Lcanckan naked: sepals 9 mm long.
Costa Rica. a me ctor v. triflora
2. Scape-bracts all imbricate (un known i t uerckhed rhe we
10. pice mee flowered; floral eae Braap ree ig ads than
s long as the internodes.

ae Pedicels, 1-2 cm thes Miaka or exceeding the floral
bracts. Costa
sas Floral bracts fie. ‘coriaceous; pedigeit stout,21
Senne 4d Veeaa « 136. V. dieccle

a2} Floral Adele ‘Geubconacnous: pedicels: oats slender, 1-2

.85. V. pedicellata
il. pegieais sister” ge Fh a e fs the floral bracts.
er, at most RAEOAY exceeding 2 mm in
ameter; floral bracts mostly thin

14. Leaf-blades narrowly triangular, 15 mm wat

Brazil.
V. lubbersii
14. Leaf-blades ligulate or linea
15. Sterile bases of the haa ‘mack longer than the few-
overt: arte part; nepale narErery oblong,
Jamaica. ..+++es dhe Vv. swartzit
je sterile “phees of ue Sraaston cere ghorter than the
fertile part. Brazi
16. Sy pry bic Granted: plant 35 cm high; sepals
23 mm long, about equaling the floral bracts
Lees rae
16. pce much branched; plant 6-30 dm high
Lic Se yon See mm ars branches 8-12 cm 5 a taf lor es
part Oe arent
iT, Savile é 25 8 mm oo. pa ce 50 cm g roy inflo-
rescenc ‘
18. Upper Prago-be dite and lower primary bracts folia-

ceous; floral bracts broadly ovate, acute, mu uch
shorter than the sepals.

1966 Smith, Notes on Bromeliaceae 103

Lae wT ee 5-8 cm wide; plant lacking rhizo

fig. 18...........58. V. ‘phi Lippocoburgs
19; Leaf-blades 2- “4 em wide; plant bearing elong

Je etUsVENE Yaieesi Se. ORE v. aa
18. Uoper seape-bract rad lower primary bracts - lanceo-
la flor 2

sence or obtuse, fe ye shorter than or equaling
the sepals i visas sch. e8ie. 64 ‘ -28. V. procera
13. Branches stout, eget: more hae 2 mm or dimisters. floral
bracts mostly coriaceous.
20. Floral bracts much shorter than the sepals.
re oo ree bracts exceeding the sterile bases of the
anches; floral bracts 25-30 mm long. Jamaica
23.°V. ‘gibba
21. Primary bracts much shorter than the sterile bases of
the branches.
22. Floral bracts 20-25 mm long; sepals narrowly obovate,
mm long (cf. Tillandsia mima
22. vii bracts 7-15 mm long; sepals e elliptic: 15-20 mm
long. Pl. IV, figs. 15-17. Hispanio
3. V. tuerckheimii

ed
23. Rhachis verrucose just below each node; floral bracts
broadly ovate, dtcathane: PIS IV, figs 19.:- Costa
o Colombia, West Indies.........138. V. ringens
23. Rhachis not at all verrucose.
24. Sepals acute. Brazil
25. Primary bracts with distinct elongate blades; inflo-
rescence much branched.
26. Floral bracts ovate, loosely and incompletely
sheathing the base of the flower, becoming slight-

thin margin; sepals oblong, to 37 mm long, about 3
times as long as broad. Pl. I, fig. 25.

6. V. gigantea

26. Floral bracts egos ci tightly — cage ata
sheathing the bas he flower, becoming secund,

even; sepals Lance-ovate, 30 mm iets about twice

as lon d.
23 teats bracts bladele ess; ‘pibonbpiebous; inflores-
e few-branched; sepals 26 mm long.
140. V. ruschii
24. Sepals obtu
27% roe biaits 32 mm long, about twice as long as the
ternodes; sepals elliptic. Brazil.
141. V. hoehneana
27. Floral bracts 25 mm long, about equaling the inter-
des; sepals broadly elliptic. Trinidad, Guiana,
azonian Brazil... <.isss ee see% 6.2142. V. lamazonica
10. Branches densely flowered; floral ‘bracts twice as long as
the internodes or longe

OOOO eee H ESHA HEHHE CECE OEE HCC OEE HHS EE HH HEH EOE HEL EBEOCE®S

10h PHYTOLOGIA Vol. 13, no. 2
28. Floral bracts drying dark castaneous except for the apex
or the margins. Brazil.

295 — acute; floral bracts broadly ovate, ot pale
slightly rugulose
L

near the wens “4 8).¢
ly much exceeding the sepa
29.5 Sepals obtuse; floral bracts
7

>
si tp with narrow
margins, 2-3 cm i usually shorter than t

pals... PL. V i; seg hes V> iatiatae

28. eicesi bracts usually ereses or preaaltihs sometimes red o

pale brown, but always pale on dryin

30. Floral bracts equaling or exceeding she sepa

east the lowest flowers.

31. Sepals twice as long

;. fERy

ls of at
as ope or narrower;
narrowly obtuse to acumin

floral bracts
32. Rhachis verrucose below . “nodes floral bracts acumi-
mate oP1.1V, fie. 19. a Rica to Colombia, West
Ea a not = i verrucose;
nar

Vv. ringens
Adored binede acute or
34. Fisted aun qn te egy sharply ee Brazil.
V. brusquensis
33. Floral bracts broadly convex, w r not at all
carinate (cf. Tillandsia aoe etc secun
31. Sepals but slightly longer than wide; floral -ge ob-
tuse, dull.

34. Floral brac convex, ecarinate, 22-30 mm long.
35. Spikes many with naked sterile bases. = Rica.
81. V. camptoctads
35. Spikes few with long many-bracted sterile ‘pases
e6ur2, Colombia.....ese--
30% Floral br

. «144.°V. confusa
34. Florat brsuta akeee toward apex, 35- mn lo
alvador ies Vey es .145. V. cornus-cervi
aéts talwaye dterancnly: shorter than the sepals.
36. Flowers upwardly secund; branches spreading, many; se-
pals 35-40 mm long; petals flaccid. Bra
37. Rhachis strongly eS only a few flowers secund.
Ply sil;fie? 224

222. V. brasiliana
nd.
fee. 4, 34 cy iit ee Ges 284 Y- imperialis
36. Flowers downwardly secund,
38. Leaves ornamented with dark purple irregular transverse
ands; floral bracts broadly ovate, carinate;
narrowly elliptic,

37. Rhachis merely Pininones side of the; flowers secu
iV, -£

rinate; sepals
25 mm long, much one rtedi: Pl. Vs
tT il...........+-146. V. hieroglyphica
38. Leaves without purple bands but ares with narrow
dark green lines.
39. Sepals 20 mm long; branches 4-8-flowered, short, very
slender. Brazil.

40. gece bracts nearly oe the sepals; branches
ith naked sterile base
147.
40. Floral bracts only about half as long as

V. pe enduliflora
the sepals;

1966 Smith, Notes on Bromeliaceae 105

branches with bracteate sterile bases
- V. peneda as-viannae
339; ee much re _ 20 mm long or the branches many-
ered and s
41. Laseb ates see in scape-bracts age ae or acute
with a stiff Lpklatecuneaiee terminal c
42. Sepals cag equaling the 3 cm long floral “tracts.
a Ric

fo)
43. Floral pants obtusely carinate, more or less se-
with the flowers. Pl. V, fig. 8. Mexi ico,

emala also. Fe pana went 9. V. werckleana
43. victt praads etutinnte: er at Pall se as
O. V. kupperiana
42. Sepals distinctly iets ach the floral bracts
44. Floral bracts abruptly acute, the keel peels

curved from base ye apex. Wie Ve Le. Se GOBCH
Rica to Colombia, Greater Antilles.

151. V. sanguinolenta
44. Floral bracts eases _gibbous about one-third
Jam

r
subrounded with a flat soft apiculus. gang
45. Inflorescence few-branched, the terminal branch with
ongate sterile base almost cutitets covered
with bracts.
46. Sepals to 35 mm long; floral bracts barely = as
long as the Aue ta atat plant generally 2m
t he arabes 141. V. St ae
46. Sepals 17237 teal ak tone: floral bracts from barely im-
ricate to 3.5 times the internodes; plant 6-12
ta
47. Leaf- biades glabrous above; floral bracts to 35 mm
ong, 2.5-3.5 times the intern

152. V. longiscapa
47. Leaf-blades covered on both sides ta appressed
cretaceous brown-centered scales; floral bracts
10 mm long, scarcely more than twice the inter-
nodes. aso $6 b Swe 5 153. V. cearensis
45. Inflorescence may Be entichsd he terminal branch not
h different from the others
48. Floral bracts 3 to 4 times as long as the inter-
odes, remaining imbricate, obtusely carinate;
-blades concolorous........154. V. hydrophora

leaf-blades marked with dark green cross-lines.
155. V. pastuchoffiana

SUBKEY IX

1. Floral bracts acuminate, sharply carinate toward apex; rhachis
verrucose below each node. Pl. IV, fig. 19. Costa Rica to

106 PHYTOLOGIA Vol. 13, no. 2

Colombia, West Indie ; 138. V. ringens
g ae bracts acute or gfhg or af “rarely. the. lowest acumi-
nate then the rhachis not verrucose
2. Inflorescence dense, the floral Ste at least twice as long
as e internodes
3. Scape-bracts ahortes than the internodes
4. Floral bracts acute, green; leaves Soonlaaee. Guatemala.
156. V. pectinata
4, Stet bracts obtuse, castaneous; leaves pee! rounded or
truncate. Brazil weeeeeel5/. V. clausseniana
a: Kaecbrac cts all exceeding the inte rnodes
5. Floral bracts rugulose or verrucose Se reashout when dry.
6. pe .pyerts sorts contracted to a distinct triangular
sepals 158. V. woodsoniana

haan ae

ama.
6. Floral ‘caste Pepe abrupt distinction between base

ay Leaf bighue broadly acute or rounded and apiculate
8. Floral bracts becoming secund with the flow

owers; sepsis
twice as long as broad. British Guiana.

. V. pachychlamys
sepals tase peel twice as
Guat eet
gladioliflora
Te iae-b taded Gecatnates “Horel “beacts Diemis more than
s long as the internodes.
, green and obscurely lepidote on
floral bracts rugulose,
thick, coriaceous. Costa Rica......160. V. tonduziana
g. Leaf-blades pale and densely appressed-lepidote beneath;
epals not over 25 mm long; floral bracts rather thin,
verrucose. Pl. V, figs. 10, 11. Me meee

8. Floral bracts remaining erect;
long as Ae: baa... Pl. tl, £18

4. Ve vanhyningii
5. Floral bracts even or nerved but neither rugulose nor

xcept for the pale soft apex, 3-8 cm long.

Brazil.
106. V. longicaulis
10. Floral bracts uniform or near

ly s
11. Floral bracts remaining erect, Bae becoming secund with
the flowers.
12. Floral bracts ecarinat
13. Floral bracts oie wake twice as long as the inter-
nodes. Braz eee LY ee unilateralis
13. Floral bracts Gs when dry, densely imbricate.

a
14. Leaf-blades acuminate, concolorous, subglabr
114

ous.
as, & acuminata
14. Leaf-blades subrounded and gaara densely white
lepidote beneath, subglabrous abov

“163. V. bicolor
12. Floral bracts carinate at least toward the ap

1966 Smith, Notes on Bromeliaceae 107
15. Leaf-blades acuminate; floral bracts up to 5 cm long,
much exceeding the sepals. Lesser Antilles
164. V. guadelupensis
15. Leaf-blades rounded and apiculate. Brazil.
16. Floral bracts greenish brown, obtusely ee
Ee istine
16. Floral bracts bright red except for the —_ apex
and margins, sharply carinate.
QO. V. interrogatoria
11. Floral bracts becoming secund with the flowers.

O mm in diameter, dark, its

spotted;. plant 1-2; m high. ..Pl., V,.-fig. 9.
Rica to Colombia, Greater Antilles.
151. V. sanguinolenta
17. Rhachis terete or angled with its internodes subcylin-
rical, rarely more i 5 mm in diamet
18. ae biades less than 2 cm wide, acuminate: plants not
dm high; ae alee few-flowered. Costa
Seis ster eeoltaes Supsecunda
puis: pad 5-20 dm high; inflo-
escence usually many-flowe
19. Floral bracts strikingly ie with contrasting
margins.

Rica, Busens
18. Leaf-blades 3-10 cm w

20. oe bracts bright red with green apex and margins
Pi ews eee pene ees N. interrogatoria
20. Ponat bracts drying dark. castaneous with a row
pale margin, even, lustrous, bro adly convex
21. Leaf-blades very obscurely Cenchulatenlantdobe be-
neath; floral bracts broadly ovate. Costa Rica
166

gi. pittiers
21. Leaf-blades covered beneath with white scales;
fil

oral bracts broader than long. Costa Rica.

167. V. ampla
19. Floral bracts green, dull buff, or pale brown.

23. Sepals 15-20 mm long; leaf-blades 2.5-4 cm
24. Leaf-blades concolorous; floral bracts equaling
the sepals. Pl. V, fig. 12. Cent tral America,
British Guiana, Vedsiueta, Colombia
Vv. viridetiora
24. pa cep a with fine longitudinal r
floral bracts much exceeded by i se-
cates "Pl. vy, gos 13. Costa Rica.
69. V. marnier-lapostollei
23. Sepals 35 mm long; leaf-blades 6 cm wide. Cuba.
0. V. haplostachya

22. Floral bracts carinate.

26. Sepals up to 34 mm long; floral bracts barely
twice as long as the internodes; plant usually

108 PHYTOLOGIA Vol. 13, no. 2

2m oor higher. Brazil........-. 141. V. hoehneana
26. Sepals 20-27 mm toa floral bracts two and a half
three times as long as the i aris ierad plant
6-12 dm high. Brazil......... Ns Pe longiscapa
2. Inflorescence lax, the floral bracts distinctly fod than
twice as long as the internodes.
27. Leaf-blades linear, 4 dm long and not over 5 mm wide; in-
lorescence typically aint floral bracts
mbranaceous. Costa Rica.......... sLioe Ve arama
27. Leaf-blades narrowly értaigaiat or ligulate, 15-80 mm wide
inflorescence normally much more than 3-flowered; ftere®
bracts firmer.
203 perce or at least the upper ones, shorter than the
rmodes; sepals much exceeding the floral bracts
29. Leaf-blades narrowly triangular, acuminate; floral bracts
ob ecarina
30. peptone bracts and sdpals thick, ee Brazil.
173. V. brassicoides
30. Floral bracts and sepals ee didtracedua.
Venezuela joie ded i OSI Saisie es seeuee 174. V. fibrosa
29. pagoutiheng wis ligulate, ee or broadly rounded ‘and apicu-
la floral bracts acute.
SV Floral bracts ecarinate; sepals suboblong, to 40 mm
CO LOMB disses o-s:cete's oteis'e's wletees's Feel fia BM, oc gpa

i Oe Floral ‘bracts avateee “gepaid elliptic, 25-28 mm lon
17

ee ue swe PO ees sell Oe ve Pppieat
28. aciieaaiy all imbricate

32. Floral bracts exceeding the sepals.

33. Rhachis verrucose below each node; floral Mactan rugu-
lose; leaf-blades concolorous, green wide.
ai so ke HOS CEC eee bas de MOST e es et T. a Carte

33. Rhachis not Gericuae floral ions even; leaf-blades
banded or spotted, 30-35 mm

34. andere in Os acute or sidsinava sepals 17 mm long.
ritiel Homdurag is ii ee se a oo 178. V. schippii

34. Hr gore bracts vei, ‘sepals 26 mm long. Costa Rica.

179. Y. macrochienys

32. Floral bracts distinctly exceeded by the sepa

35% ae narrowly triangular, Peal contnate. not

wi

3
36. Floral bracts fleshy-coriaceous, coarsely lepidote ex-
— n age; flowers vba geo sg ge 27-30
- 180.

mm long razil...... V. oligantha
36. visas bracts eis Na A spars sely and minutel
lep te; flowers spreading; sepals Scaggs elliptic,

18 mm long. Pl. V, figs. 14, 15. Trinidad.
181. V. broadwayi
35. Leaf-blades ligulate, acute or rounded and apiculate or
etimes the apex acuminate for a short distance,

15-8 0 mm wide.
37. Plants less than 1 m high; scape slender; floral bracts
ecarinate.

1966 Smith, Notes on Bromeliaceae 109

38. Maven bracts little more than once as long as the in-
nodes; leaf-blades eeaige Se Pi:
figs, 16. 1), .Bragél« -182. V. racinae
a8, raed bracts nearly twice as wpeis as Sie internodes;
leaf-blades concolorou
39. Tanta ci secs broadly scien and nedabacaagrig floral
bracts remaining erect or nearly azil.
1 Vv. rs
39. me “paneen acuminate; floral bracts babceiad secund
the flowers. Colombia.......183. V. vexillata
= 9 ge A 4 2 m high or more; seta stout; floral bracts
obtusely carinate toward a
40. Be bas: narrowly elliptic; tous bracts about twice a
ong as the internodes. Brazil....141. V. enhenala
40. Sepals very broadly elliptic: floral bracts about as
long as the internodes. Trinidad, Guiana, Brazil.
142. V. amazonica

SUBKEY X
1. Branches of - inflorescence well developed, at least below
he fl

2. Flowers vier eames (pedicels at most 4 mm long and very

stout
3. Rhachis weld developed, the flowers racemos
4. Primary brac covering the branches (cf. Tillandeta

ts
biflora, stipitata
4. Primary Rees ts much eh than the branches, or if
longer than the lowest ones (V. Splitgerbert) then narrow
and spreading and not covering them
5. Flowers erect, densely imbricate, numerous. Pl. V, figs.
18, 19. Costa we Trinidad, Guiana, Venezuela
Colombia.... : p wemep ee ete splitgerberi
5. Flowers divergent. to centendines 1 :
6. Floral bracts 30 mm long, onnmaiiien the get Brazil.
N. corcovadensis
6. Floral bracts 4 mm long, much shorter than the
C a ware tbe. cine sce eo v. fee
3. Rhachis aborted, the cioumen. nccraie
7. Branches short, the lower ones 10-15- Sintered. Pl. V, fig.
20. West Indies, Venezuela, Colombia, Ecuador
oV cepitulicers
7. Branches elongate, the lower ones 2-flowered. Costa Rica.
87. V. crassiflora
2. Flowers distinctly pedicellate. Costa Rica.
GS. Pedicels 11. mm Jong seissceuphoreerassa0anees 99. VY. turbineta

10. Sepals symmetrical, narrowly obovate, broadly rounded, 14
mm long. vps oe bre ean ss sseaeesaeOe V. cylindracea

10. daeais strongly. asymmetric, slightly emarginate, 10 mm
long (undeveloped) ........+seee+0++++-+189. V. leptopoda

110 PHYTOLOGIA Vol. 13, no. 2

9. spree very stout; leaves green gS with violet be-
Nees wo ee ee ees ee se V. latissima
de sical “7 the inflorescence very sho a or “aborted with the
flowers fascicled in the axils of the primary bracts, or
else the perfect flowers ei Seber a on each branch (the genus
hyllum as used largely by M
Ae press 3 or more in the axil of ie Yeast each lower primary
act.

2s vient bracts exceeding the eu this leaves concolorou

Costa Rica... . reese sus Be bracteosa
12, rr ne much shorter’ ee the i gipats: leaves cros
bar

13% Bapeiles: “priees large, en thiet floral bracts broadl
reniform. Costa Rica... er LS ee cashed aks
13. Primary bracts saat tibbee’ petal bracts suborbicular.
Guatemala, Honduras...... Seccev sive. Ve montana
11. Flowers 1 or 2 in the axil 3 cach orinaty bract
14. Perfect flowers 2 in the axil of each primary beack:
15. Pedicels slender, 1-2 cm long. Costa Rica.
193. V. violascens
15. Pedicels stout and short or lacking.
16. Floral bracts exceeding the s

sepal
Les — = the leaves and scape-bracts covered with pale
sed scales. Costa Rica
18. iatassacne uy ie ferttie throughout.
19. Leaf- Le ae Posckcate angular, attenuate, 2

5 mm
wide imary ste ey ae adinig...1194 5°V.s leucophy1la
ag, beaf-biades ligulate, abruptly acute, 35 mm wide

ary bracts erect.....cscecees en aes spectabilis

18. Toftereacanse sagen with a coma of ‘sterile bracts
at apexes’. .196. V. comata

17. Blades of the tanves aor scapesbrdcts glabrous or sub-
la

glabrous
20. Sepals 7- Sa long.
21. gee bracts erect; leaf-blades acute, 30-40
de. Lesser Antilles............. a at
21. Primary bracts igeuieed coverd a apex; leaf-blades acu-
sep » 25 mm wide. Trinidad, eee (Isla Mar-
gar a) secceesseeteneseenessteeesDD . V. johnstonii
= per i 10-15 mm
. Leaf-blades ale throughout with fine irregular
cro oe oe bapa densely capita 14
—cylind Couta Rica.....0. 2.2199. Ve notats
22, leat Finlee sg Maes or slightly marked at
23 meget ligulate, acute to rounded and gor eeaaes!
inflorescence — sepals 15-17 mm long.
24. Sepals Ret: rbicu Costa Rica...190. V. bracteos*
24. Sepals elliptic. Tcolbubia. <<. (208: V. hygrometrica
23% gaged oe digg ane inflorescence lax at ieeat
d bas

e: 0

owar Ecua iViGecese V. paupera
16. Floral ieee shorter thea? the ‘epee’
25. Leaves with fine wavy cross-lines.

1966 Smith, Notes on Bromeliaceae ni

26. Floral bracts ovate, acute; primary bracts suberect;
leaf- 5s acuminate, 40 mm wide; sepals 16 mm lo

Costa Ric .
26. Floral Mas suborbicular to aes tame rounded o

t cate; primary bracts spreading to reflexed.
27. Leaf-blades acuminate, 20 mm wide; sepals 8 mm lon
slightly asymmetric. Costa Rica..... 203. V. lineata

28. Scape erect; sepals broadly elliptic, 12-20 mm long.
29. Inflorescence few-flowered, subdense, 5 cm long;
sepals 12-15 mm long. Guatemala, Honduras.
192. V. montana
29. Sra eres many-flowered, slenderly cylindric,
5-30 cm long; sepals 20 mm long. Costa Rica.
V. irazuensis
28. Scape — sepals broadly ehcerath 20 mm long.
Costa 205 oriensis
254 vanvee unmarked or sometimes with straight g Peron race
str

30. istcniecnes Pach sane or short-cylindric, not over 5
cm long. Costa Rica

O13 Seton 8 mm — stout: leaf-blades 4-5 cm wide.
206. V. capitata
31. Pedicels lacking; leaf-blades 6-15 mm wide.
32. Floral bracts broadly elliptic, to 12 mm long.
207. V. attenuata
32. Floral bracts squamiform, 2.5 mm long.
208. V. stenophylla
30. Inflorescence cylindric, 7-27 cm long.
33. Leaf-blades and scape-bracts acuminate, pale-lepidote;
primary bracts reflexed. Costa Rica.
34. Sepals 9 mm long, orbicular.......209. V. balanophora

loral bracts 8 mm long; sepals 17 mm long.
210. V. viridis
35. Inflorescence few-flowered, pics Lax, / cm . long;
floral bracts 16 mm long; sepals 20 mm long
11. V. hainesiorum
33. Leaf-blades (unknown in V. spectabilis) and scape-
bracts acute or rounded and apiculate.
36, Primary and scape-bracts marked with fine purple lon-
gitudinal stripes like the leaves. Costa Rica.
212. V. williamsii
36. Primary and scape-bracts unmarked.
mF oF ante bracts erect or iti so, not recurving
owar a Rica » Ve spectabi is
a7. oianes bracts passe to urecucted, at lea
toward apex.
38. Primary bracts densely ee

39. Floral bracts acute, 12 mm
213. V. standleyi

Costa Rica.

112 PRESTO Gis Vol. 13, no. 2

39. Floral bracts rounded, 3-8 mm long.
188. V. cylindracea
38. Primary bracts glabrous and even or verrucose.
40. Inflorescence erect; sepals obovate or narrowly
elliptic, twice as long as wide. Puerto Rico.
214. V. sintenisii

40. Inflorescence al pend, sepals very broa dly
obovate. Costa Rica.......... 205. V. ororiensis
14. ag diode flowers single in the axil of each primary bract,

e presence of a floral bract Prod He a compound
entire escence. Costa Rica
41. Flowers (branches) secund...........+, 215. V. singulit tors
41. Flowers (branches) remaining polystichous.. 216. V. vittata

SUBKEY XI

hee eiaawaeet Gal ein ce only at yee aa elon-
A UVEPTCly) seers oes se ses 125. V. platynema
1a Ee a oaks iy polystichous-— ae uwers a.
2. Leaf- Bee ligulate, acute, 4-5 cm wide. Pl. VI, figs. 1,
Pe Pe Ree ie ray err ers tae 217. V. malzinei
rp tet bisaes a ertengutar, acuminate, razil.
3. weed s all bearing recurved oe blades; inflores-
ense, few-flowered, 4-9 cm long....62. V. poenulata
3. Seape-bracts beartng erect blades, those of the upper ones

very short.
4. Inflorescence dense except at base; flowers in about 4
mostly suberect; floral bracts inflated. neh VI,
ee Pe ee er -63. £lammea

igs
4. Tat iovek coun lax ficwers in slightly 1 more “than ] “tanks,
all divergent or spreading at anthesis; prion bracts
closely enfolding the sepals......... 6i. V. corcovadensis
Subgenus VRIESEA
Section VRIESEA

1. V. HITCHCOCKIANA (L. B. Smith) L. B. Smith, Contr. U. S.
Nat. Herb. 29:446. 1951. 5 ie hitchenekiana L. B. Smith,
Contr. Gray Herb. 89:10 4, figs 1930. Ecuador, Peru.

2. V. RAUHII L. B. Sm Ae pykctaeb is pi. 1, figs. 55-6
1958. Stamens probably eubectud, Pl. I, fig. 1: Spike x 1/4.

3. V. CEREICOLA (Mez) L. B. Smith, Phytologia 6:194. 1958; 8:
504. 1963. Tillandsia cereicola Mez, Fedde Rep. Nov. Spec. 3:34.
1906. Vriesea ea lopezii L. B. Smith, PHytologia 4:218, pl. 2,
figs. 8-10. 1953. Peru.

4, V. ROBUSTA (Griseb.) L. B. Smith, Phytologia 7:4. 1959.

figs. 9-12. 1955. Stamens (immature 2) about equaling the
re Mrvancerene Ae Colombia

1966 Smith, Notes on Bromeliaceae 13
- V. OLMOSANA L. B. Smith, sp. nov. AV. cereicola (Mez) L.
B. Smith, cui valde affinis, bracteis florigeris carinatis
differt.
T stemless, flowering to 6 dm long with the inflorescence
extended. A

e
numerous, 5-6 dm long, completely covered wi

nerved and reddish toward apex, yellowish

green below; pedicels obscure. SEP free, lanceolate, broadly
acute and apiculate, 23 mm long, ecarinate, subcoriaceous; petals
linear, 35 mm

long, bearing 2 large linear-lanceolate acute
scales at base, the blades deep blue-violet with a narrow white
margin; stamens exserted. Pl. I, fig. 2: Leaf x 1/4; fig. 3:
Spike x 1/4; fig. 4: Sepal x 1.

fig. 46. 1951. Pl. 1, fig. 7: Spike x 1/4. Colombia, Ecuador.

7. Vv. A (L. B. Smith) L. B. Smith, Contr. U. S. Nat.
Herb. 29:447. 1951. Tillandsia harmsiana L. B. Smith, Contr.
Gray Herb. 98:16, pl. 4, figs. 12-15. 1932. Pl. I, fig. 8: Spike
Bh eru.

8. V. TILLANDSIOIDES L. B. Smith, Phytologia 9:256, pl. 4,
figs. 6, 7. 1963. Pl. 1, fig. 9: Spike x 1/4. Peru.

TeV, OCALYX (André) L. B. Smith, Contr. U. S. Nat. Herb.
29:445. 1951. Tillandsia arpocalyx André, Enum. Bromél. 7. Dec.
13, 1888; Rev. Hort. 60:567. Dec. 16, 1888. a. :

° « Nat
Tillandsia tequendamae Andre’, Enum. Bromél.
- Dec. 13, 1888; Rev. Hort. 60:568. Dec. 16, 1888. Venezuela to
Peru,

11. V. PEREZIANA (André) L. B. Smith, Contr. U. S. Nat. Herb.
29:444, 1951. Tillandsia pereziana André, Enum. Bromél. 7. Dec.
13, 1888; Rev. Hort. 160:567. Dec. 16, 1888. Colombia,

- V. LUBBERSII (Baker) E. Morr. ex Mez in Mart. Fl. Bras. ay
Pt. 3:533, pl. 99. 1894. Tillandsia lubbersii Baker, Handb.
Bromel. 219. 1889. Pl. I, fig. 10: Spike x 1/4.

13. V. TRIANGULARIS Reitz, Anais Bot. Herb. Barbosa Rodrigues
42155: pls 4.1952, Brasil.

44, V. BARCLAYANA (Baker) Lv B.. Smith, Contr. U. S. Nat: ig
29:517, fig. 75. 1951, °Tillandsia barclayana Baker, Journ. Bot.
25:239. 1887, T. lateritia André, Enum. Bromél. 6. Dec. 13,

ny PHYTOLOGIA Vol. 13, no. 2

1888; Rev. Hort. 60: 566. Dec. 16, 1888. Ecuado
15. V. PATULA (Mez) L. B. Smith, oe 5: 1288. 19552 2ii-
landsia patula Mez, Fedde Rep. Nov. 23562906. Ecuador, _
Peru.
16. V. GRAVISIANA Wittm. Gartenfl. 39:494, figs. 81, 82. —
V- lubbersiana hort. ex Wittm. loc. cit., not E. Morr. Bi:
fig. 12: Terminal spike x 1/6. i.
17. V. SINCORANA Mez, Fedde Rep. Spec. Nov. 12:418. 1913.
Brazil.
Vv. SCHWACKEANA Mez _ DC. Mon. Phan. 9:590. 1896 2: Pix T,
fig. 16% Spike x 1/5. Braz
19. V. PINOTTII Reitz, pn Bot. Herb. Barbosa Rodrigues 4:
12, pl. 3. 1952. Brazil.
~ VY. FRIBURGENSIS Mez in Mart. Fl. Bras. 3, pt.33;537. —
URGENSIS. Vriesea tweedieana sensu F. Mue Pe
Seer 42:738. 1893, in part. Inflorescence distinctly longer
than broad; branches erect or suberect; floral bracts all ecari-
nate, to 35 mm long; flowers all spreading. Pl. I, fig. ee

Var. b. PALUDOSA (L. B. Smith) L. Smith, Anais Bot. Herb.
Barbosa Rodrigues 4:68. 1952. se at " paludosa Ls! Be Smithy-Ar=
quiv. Bot. Estado S. Paulo n. ser. 1:119, pl. 128. 1943. V.
saundersii sensu L. B. Smith, Anais Bot. Herb. Barbosa Rodrigues
2:26, 54. 1950, not E. Morr. Upper floral bracts carinate, about

Var.
bosa Rodrigues 4:68. 1952. Vriesea tucumanensis Mez in DC. Mon.
Phan. 9:585. 1896. V. caldasiana Mez, op. cit. 607. V. argen-
tinensis Spegazzini, Physis 3:158. 1917. V. lutinosa Wawra var-
viridis Hassler, Ann. Conserv. & Jard. Bot. Geneve 20:319. 1919.
Branches spreading to arching-recutved, lax, flowers spreading.
Brazil, Paraguay, Argentina

21. V. BILLBERGIOIDES E. ‘err. ex Mez in Mart. Fl. Bras. 3,
pt. 3:534, pl. 100. 1894. Brazil.

Var. a. BILLBERGIOIDES. Scape-bracts imbricate; inflorescence
bipinnate.

Var. b. SUBNUDA L. B. Smith, Arquiv. Jard. Bot. Rio de Janeiro
alc 1951. Scape-bracts lax.

. AMPLA L. B. Smith, Arquiv. Jard. Bot. Rio de Janeiro

15< 330, | oan r. 1958. Inflorescence amply tripinnate

23%
ridlandsia rodigasiana Baker, Journ. Bot. 26:138. 1888. T. twee-
die loc. cit. T. citrina Baker, Handb. Bromel. 224.
1889, See citrina E. Morr. ex Baker, loc. cit. nomen. V.
tweedieana F. Mueller, Gartenfl. 42:738. 1893; Mez in Mart. Fl.
Bras. 3, pt»: 33561, 1894, as to basonym. V. vitellina F. Muel-
ler, loc. cit. Pl. I, fig» 18. Spike x 1/4. >

24. V. BRUSQUENSIS Reitz, Anais Bot. Herb. Barbosa Rodrigues
4:10; pls’ 2. 1952.: (Brazil.

5. V. NEOGLUTINOSA Mez, Engl. Pflanzenreich IV. 32:636. 1935.
Tillandsia glutinosa Mart. ex Schult. in R. & S. Syst. 7:1225.

1966 Smith, Notes on Bromeliaceae 5

1830. Vriesea glutinosa Wawra, It. Sax.-Cob. 167. 1883, not
Lindl. 1856.

26. V. MUELLERI Mez, Bot. Jahrb. 30, Beibl. 67:7. 1901.
Brazil

27. Yy. SAUNDERSII (Carr.) E. Morr. ex Mez in Mart. Fl. Bras.
3 Ci 4240s: Louw. Enchloricn saundersit = i Rev. Hort. 44:
300. 1872. Tillandsia saun Sem. Hort. Berol.
1873, App. 4:6. 1874. Vriesea Pivaievis we in Mart .-¥1.
Bras... 3, pt: 32536, -1894,- Pie 1, flee. 20: Leaf ¢ 1/3; “fig. 212
Spike x 1/3. Brazil

28. V. PROCERA Care, ex Schult.) Wittm. Bot. Jahrb. 13,
Beibl = 29:27. I

Vat," a. CERA. Tillandsia procera Mart. ex Schult. in R. &
Vriesea gracilis Gaud. Atl. Voy. Bonite

ts.

V. procera var. gracilis Mez in Mart. Fl. Bras.
Tillandsia viscidula Britton, Bull. Torrey Club 48:

gr 2 5x boy g. 22: Spike x 1/4. Trinidad, Guiana, Brazil.
RUBRA L. B. Smith, Arquiv Sct Estado S.
ser. 22197. 1952: “Bratte- red goa yellow apices. Braz
° . B. Smith, Arquiv. Bot. Estado S. Paulo n

lo
ser. 1:121, 1943. Tillandsia gracilis Ule, Bericht. Deu tsch.
Bot. Gesellsch. 18:325, pl. 10, figs. 7-11. 1900. T. ermestii
Mez, Engl. piairenetel EV. 324636. 1935. Branches ar Yipee to
recurving, often twisted so that the flowers are s floral
d

re se
bracts membranaceous, nerved, much shorter than the sepa sis.
Brazil.

Va EBILIS Mez in Mart. Fl. Bras. 3, pt. 3:540. 1894.
Scape-bracts nearly all shorter than the internodes; inflores-
cence simple or subsimple; some of the floral bracts incurved.
Brazil.

29. V. GRADATA (Baker) Mez in Mart. Fl. Bras. 3, pt. St523.
1894, Tillandsia gradata Baker, Journ. Bot. 26: 105. 1888.

Brazil

30. V. MODESTA Mez, Bot. Jahrb. 30, Beibl. 67:7. 1901. Brazil

31. V. ERYTHRODACTYLON E. Morr. ex Mez in DC. Mon. Phan. 9:
569. 1896. Vriesea psittacina var. erythrodactylon E. Morr.
Belg. Hort. 32:287. 1882. Tillandsia carinata sensu Baker,
Handb. Bromel. 212. 1889, in part, not as to type. Vriesea du-
valiana sensu Wittm. Bot. Jahrb. 13, Beibl. 29:21. 1891, not ge
Morr. V. carinata sensu Mez in Mart. Fl. Bras. 3, pt. 32516.

1894, Po part, not as to type. Pl. II, fig. 3: Spike x 1/4.
Brazi

32. v. BARILLETII E. Morr. Belg. Hort. 33: ee pl. aS aia i:
B. Smith, Phytologia 8:507. 1963. vrieses eumorpha
Carr. Rev. Hort. 55:394. 1883, nomen. Tillandsia Sookie ts
Baker, Journ. Bot. 26:79. 1888. Vriesea epee hort. ex

116 P £.¥.7 06,.1.06.2 4 Vol. 13, no. 2

Baker, Handb. Bromel. 215. 1889, nomen. Pi... L1;. figs. 5s; Spikex
1/4. Brazil.

. V. VULPINOIDEA L. B. Smith, Arquiv. Bot. Estado S. Paulo
n. ser. 1:122, pl. 134. 1943. Brazil.

34. V. INCURVATA Gaud. Atl. Voy. Bonite pl. 68. 1843. Vriesea
psittacina var. truffautiana Andre, Journ. Soc. Hort. France Li 3
3:89. 1881. V. inflata sensu Andre, Rev. Hort. 59:44. 1887, as
Tillandsia, not Wawra. Tillandsia incurvata Baker, J
26:49. 1888, not C. Wright, 1871. Vriesea truffautiana hort. ex
Baker, Handb. Bromel. 213. 1889, nomen. Vriesea rostrum-—aquilae

Mez in Mart. Fl. Bras. 3, pt. 3:518, pl. 107. 1894. V. duvaliana
sensu Alexander, Addisonia 19:47, pl. 632. 1936, not E. Morr.
1, EL, £ie5..62 Sp 1/4 2m

V. INFLATA (Wawra) Wawra, It. Sax.-Cob. 161. 1883. Vrie-
sea carinata var. inflata Wawra, Oesterr. Bot. Zeitschr. 30:183.
1880. Tillandsia inflata Baker, Bot. Mag. pl. 6882. 1886, as to
bas . Vriesea incurvata var. inflata Mez in Mart. Fl. Bras.

but not as to basonym of Wawra. Pl. II, fig. 8: Spike x 1/4.
Brazil.

37. V. SPLENDENS (Brongn.) Lem. Fl. Serres 6, Misc.:162, fig.
1850-51. oma

Var. a. SPLENDENS. Tillandsia splendens Brongn. Ann. Fl.: POz
mone 18:311. 1845. Vriesea speciosa Hook. Bot. Mag. 74:pl. 4382.
1848. Tillandsia vittata Rich. ex Baker, Handb. Bromel. 215.
1889, nomen. T. picta hort. ex Baker,
brina hort. ex Baker, loc. cit., nomen, not op. cit. 1/. Leaf
blades with broad irregular cross-bands. Pl. II, fig. 9: Leaf
blade x 1/4; fig. 10: Spike x 1/4. Guiana, Venezuela.

Var. b. LONGIBRACTEATA (Baker) L. B. Smith, Smithsonian Misc.
Coll. 126:36. 1955. Tillandsia longibracteata Baker, Journ. Bot.
26:81. 1888. Vriesea splendens var. formosa Witte, Semperv. 18:
361. 1889. Tillandsia splendens var. formosa Suringar ex Witte,

1889. Vriesea longibracteata Mez in DC. Mon. Phan. 9:568. 1896

V. splendens major hort. Duval, R

V, longibracteata wartel
songibracteata wartetit

Leaf-blades concolorous. Trinidad, Guiana, Venezuela.

38. V. CARINATA Wawra, Oesterr. Bot. Zeitschr. 12:349. 1862.
Vriesea brachystachys Regel, Gartenfl. 15:258, pl. 518. 1866. V-
psittacina var. brachystachys E. Morr. Belg. Hort. 20:161. 1870.

rr .

49. 1888. T. psittacina sensu E. Morton, Brazil Fl. 2:pl. 43.
1893. Pl. 11, £ie.. li: Spike. x 1/4. Brasil,

. DUVALIANA E. Morr. Belg. Hort. 34:105, pls. 7, 8. 1884.
Vriesea psittacina var. duvaliana Andre, Rev. Hort. 56:559. 1884.
Tillandsia duvaliana Baker, Journ. Bot. 26:48. 1888. Pl. II,

1966 Smith, Notes on Bromeliaceae 117
fig. 12: Spike x 1/4. Brazil.

40. V. PARATBICA Siecan It. Sax.-Cob. 160, pl. 33, :figs:B,-pl.
36, fig. B. 1883. Vriesea carinata var. c
terr. Bot. Zeitschr. 30:183. 1880.
Journ. Bot. 26:82. 1888, as "

: Baker, loc. cit., nomen. Pl. II, fig. 13: Seika | = on
Brazil.

41. V. RHODOSTACHYS L. B. Smith, Arquiv. Bot. Estado S. Paulo
n. ser. 1:121, pl. 132. 1943. Brazil.

oe, V. PAUCIFLORA Mez, Fedde Rep. Spec. Nov. 16:72. 1919.
Brazi

rey Vv. eee eee Reitz, Anais Bot. Herb. Barbosa Rodrigues
4:6, ol... 1.1952. ats 3

44. V. LAXA Mez in a DC. Mon. Phan. 9:578. 1896, emend. L. B.
Smith, Phytologia 7:4. 1959. Ti llandsia laxa Griseb. Nachr. Ges.
Wiss. Goett. 1864:18. 1865, not Griseb. Fl. Brit. W. Ind. 596.
18

Titieweae maxoniana L. B. Smith, Donte, Gray Herb. 124:
pl. 2, figs. 1-3. 1939. Vriesea acterion Castellanos, Lilloa
11: 150. 1945; Gen. & Sp. Pl. Argent. 3:249, pl. 70. 1945.

Bolivia, Aceettine:
46. V. AMETHYSTINA E. Morr. Belg. Hort. 34:330, pls. 15,
1884. «Sil anata amethystina Baker, Journ. Bot. 26:104. tg

Pl. III, fig. 3: Spike x 1/4. Brazil
47

. v. PSITTACINA (Hook.) Lindl. Bot. Reg. 29:pl. 10. 1843.

a. PSITTACINA. Tillandsia psittacina Hook. Bot. Mag. 55:
Floral bracts red with yellowish apices, ecari-
“a III, fig. 4: Spike x 1/4. Brazil.

Var... b. RUBRO-BRACTEATA Hook. Bot. Mag. 85:pl. 5108. 1859.
Vriesea petttacina var. bracteis omnino coccineis E. Morr. Belg.
Hort. 32:287 V. krameri E. Morr. op. cit. 34:369. 1884.

azil (7).
c. DECOLOR Wawra, Oesterr. Bot. Zeitschr. 30:183. 1880.
Floral bracts Ati green, some of them more or less carinate.
Brazil, Paragua
oo RECURVATA Gaud. Atl. Voy. Bonite pl. 69. 1843. Til-

iandsia recurvata Baker, Journ. Bot. 26:106. 1888, not L. 1762.
T. decurvata Baker, Handb. Bromel. 216. 1889. Pl. III, fig. 5:
Spike x azil.

49. yee oo hort. ex E. Morr. Belg. Hort. 32:289. 1882.
Tehhies poittacina x brachystachys E. Morr. Belg. Hort. 29 :300
1879.

50. V. ENSIFORMIS (Vell.) Beer, Bromel. 92. 1857. Brazil.
Var. a. ENSIFORMIS. Tillandsia ensiformis Vell. Fl. Flun.
; Icon. 3:pl. 129. 1835. Vriesea conferta Gaud. a
Voy. Bonite pl. 65. 1843. V. conferta var. recu

rvata Wawr
Oesterr. Bot. Zeitschr. 30:184. 1880, in part as to po oa

118 PHYTOLOGIA Vol. 13, no. 2

tions. Tillands2 | selloana Baker, Journ. Bot. 26: 104. 1888. T.
heterostachye ean op. .cf@,- 106; 2: epdaeddia hort. Makoy,
60:20 “1890. Vriesea crousseana Rev. Hort. Belg. 19%

258. 003i V. ais oana Mez in Mart. Fl. Bras. 3, pt. 3:547.
1894. Leaf-blades concolorous; floral bracts coriaceous through-
out, bright red. Pl. III, fig. 7: Spike x 1/4.
r. b. STRIATA Alvim Seidel, Brom. Soc. Bull. 13:89, fig.
1963. gg striped with creamy yellow
V . WARMIN (E. Morr.) L. B. Smith, equiv: Bot. Estado
Ss paaia n. ser. 1:116. 1943. Vriesea Waratneit E. Morr. Belg.
Hort. 34:260, pls. 12, 13. 1884. Tillandsia warmingii Baker,
Journ. Bot. 26: 104. 1888.
are d. BICOLOR L. B. Smith, Arquiv. Bot. Estado S. Paulo n.
ger, i: a wakes
sip ALINA Mez in Mart. Fl. Bras. 3, pt. 3: 523. 1894.
rillandsia guttats Baker, Journ. Bot. 26:108. 1888, in part, Bey
type. Pl. IV, fig. 7: Leaf-apex x 1/4; fig. 8: Spike

a. fesail,

. GUTTATA Linden & Andre, Ill. Hort. 22:43, pl. 200.
1858 Tillandsia guttata Baker, Journ. Bot. 26:108. 1888. IT.
duvaliana Baker, Handb. erriee 212. re. in tt Pig 1¥,
fig. 9: Leaf-apex x 1/4; fig. 10: Spike x 1/4. 2 aie

53. V. OBLIQUA Quintus ex Wittm. Gartekit. 41: 201, pl. 1369.
1892. Brazil.

54. V. RETROFLEXA E. Morr. Belg. Hort. 34:185, pl. 10. 1884.
Vriesea _psittacina x simplex Mez in Mart. Fl. Bras. Soot ¢ Sir52a-
1 z . psittacina x scalaris E. Morr. ex Mez in DC. Mon. Phan.
fig. 11: Spike x mos Brazil.

simplex Vell. Fl. Flum. 133. 1
sea scalaris sensu Antoine,
Morr. 1879. V. mesiana Hort. Vs
nomen. Pl. IV, fig. 12: Flower x l.
emer Brazil.

56.-V

Trinidad, Venezuela,

‘ genes E. Morr. Belg. Hort. 29:301. ie
Var IS. Tillandsia scalaris Baker, Journ. Bot. 26:
108. 1888. yicrat bracts red with yellow apices. Pl. IV, fig.
13: Flower x 1; fig. 14: Spike x 1/4. Brazil.
r. Di DIS Mez in DC. Mon. Phan. 9:578. 1896. Floral
and the remainder of the inflorescence green. Brazil (7).
L. B. Smith, Arquiv. Bot. Estado S. Paulo n-
sefe 2258, ~ 200 194k

=r
58. V. PHILIPPOCOBURGII Wawra, Oesterr. Bot. zeitschr- 30:219.
1880. Tillandsia philippocoburgii Baker, Journ. :138

59. nov. Vriese
philippocoburgit | bak vagans L. B. Smith, Arquiv. Bot. Estado g.
Paulo n. ser. 1:1 pL. ESL. 1949," Be duit.
NOTE: Mulford Footer, Padre Raulino Reitz, and Mrs. Ada Abend-
a above a distinct species because of their
egpsees with it “9 the field and in cultivation. Accordingly
am following this unity of opinion and publishing the necessary

1966 Smith, Notes on Bromeliaceae 1g

combination to elevate the group to specific rank.
60. V. INTERROGATORIA L. B. Smith, Arquiv. poe Estado S.
Brazil

Paulo n. ser.. 1:117, . 424. 1943,

61. V. CORCOVADENSIS (Britten) Mez in Mart. Fl. Bras. 3, pt.
3:532. 1894. Tillandsia ventricosa Wawra, Oesterr. Bot it
schr. 30:222. 1880, not ses 1 - T. corcovadensis Britten,
Journ. Bot. 26:172. 1888. . oligantha | Baker, Handb. Bromel

215. 1889, in part, not as oe type. Vriesea riubide E. Morr. ex
Mez in Mart. Fl. Bras. 3, pt. 3:574. 1894. V. ventricosa Mez in

62. V. POENULATA (Baker) E. Morr. ex Mez in Mart. Fl. Bras.
35 BGe 1325/3, p91. 106. 1894. pounds ta glaziovii E. Morr. ex
Baker, Handb. Bromel. 229. 1889. + poenulata Baker, op. cit.
230» eae
FLAMMEA B. Smith, Arquiv. Bot. Estado S. a n.
ser. 1: sa. Dl, a0 9 pis not hort. Duval Rog nomen. Vv
fig. 3: Leaf x 1/4; fig. 4:Spike x 1/4. Brazil

Section XIPHION

64. V. CHONTALENSIS (Baker) L. B. Smith, Contr. U. S. Nat.
Herb.. 29:518, fig. 76. aa Tillandsia chontalensis Baker,
Journ. Bot. 25:237. 1887. spuria Mez & Werckle4, Fedde Rep.
Spec. Nov. 16:74. 1919. sicarapua to Peru

65. V. CRENULIPETALA (Mez) L. B. Smith, Phytologia 5:288.
1955. Tillandsia crenulipetala Mez, Fedde Rep. Spec. Nov. 16:76.
1919. T. attenuata Rusby, Descr. So. Am. Pl. 4. 1920. Colombia.

66. V. APPENDICULATA (L. B. Smith) L. B. Smith, Contr. U. S.
Nat. Herb. 29:445. 1951. Tillandsia appendiculata L. B. Smith,
Lloydia 11:307, fig. 6. 1949. Ecuador.

. HETERANDRA (André) L. B. Smith, Contr. U. S. Nat. Herb.
29: 443, 1951. Tillandsia heterandra André, é, Enum. Bromél. 7. Dec.
13, 1888; Rev. Hort. 60:567. Dec. 16, 1888. Venezuela to Bolivia

68. V. LACERA (L. B. Smith) L. B. Smith, Fieldiana Bot. 28:
sea 1957. Tillandsia lacera L. B. Smith, op. cit. 28:150, fig.

D=F,..1

69. Vv. DIDISTICHOIDES (Mez) L. B. Smith, Phytologia 5:281.
1955. Tillandsia didistichoides Mez in DC. Mon. Phan. 9:778.
1896. Pl. I, fig. 5: Spike x 1/4. Mexico, West Indies, Venez-

a
70. V. FRAGRANS (André) L. B. Smith, Contr. U. S, Nat, Herb.
29:443. 1951. Tillandsia fragrans Andre, Enum. Bromél. 7. Dec
13, 1888; Rev. Hort. 60:567. Dec. 16, 1888. Pl. I, fig. 6: Spike
x 1/4. Colombia: Ecuador
V. DREWII L. B. Smith, Phytologia 5:401, pl. 1, figs. 4-7.

REPANOCARPA (Baker) Mez In DC. Mon. Phan. 9: 581. 1896.
Tillasdeie deepancuarpa Baker, Journ. Bot. 26:41. 1888. Vriesea
dusenii L, B, mith, Contr. Gray Herb. 98:17, pl. 5, figs. 3, 4.
Li ky aa oe Es Ais ll: Spike x 1/2. Brazil.
73. V. GONIORACHIS (Baker) ene in Mart. Fl. Bras. 3, pt. 3:
545. 1894. Tillandsia goniorachis Baker, Journ. abe: 25:303.

120 PHYTOLOGIA Vol. 13, no. 2

1887. Brazil

74. Ve JIMENEZII Mez & Tonduz, Fedde Rep. Spec. Nov. 14:246.

1916. Costa Rica.
. V. SULCATA L. B. Smith, Mem. New York Bot. Gard. 9:315,
fig. ~ 1957. Venezuela.
FUNEBRIS L. B. Smith, Arquiv. Bot. Estado S. Paulo n.
ser. tr oe pl. 122. 1943. Brazi ki

77. V. SODERSTROMII L. B. Smith, Bromeliad Soc. Bull «3252;
fig. 1963. British Guia

78. V. GLUTINOSA ei Bye t Chron. 388. 1856. Tillandsia
glutinosa gg Griseb. Fl. Brit. W. Ind 9 4
Schult. 1830. stenostachya Baker, Siten, Bot. 26:109. 1888.
Vriesea pen Mez In DC. Mon. Phan. 9:564. 18962 F147;
fig. 13: Spike x 1/8; fig. 14: Sepal x ei Trinidad.

79. V. ELATA (Baker) L. B. Smith, Phytologia re i 1955.
Tillandsis elata Baker, Journ. Bot. 26:46. 1888. T. excelsa Mez,
wro Ping eR to Wittm., in DC. Mon. Phan. 9: 766. 1896. T.
eiait ata Se loc. cit. Colombia.

80. V. CHRYSOSTACHYS E. Morr. Belg. Hort. 31:87. 1881.

Var. a. CHRYSOSTACHYS. Tillandsia chrysostachys Baker, Bot.
Mag. 112:pl. 6906. 1886. T. trinitensis Baker, H Handb. Bromel.
e

Vu c c ah ker m:

and apiculate or broadly acute, rs 5 em wide; inflorescence simple
or rarely digitate; petals yellow. Pl. I, fig. 15: Inflorescence
x 1/8. Trinidad, Colombia, Peru.

Var. b. STENOPHYLLA L. B. Smith, Bot. Mus. Leafl. Harvard 17:
71. 1955. Leaves acuminate, ae 1 cm wide; inflorescence
simple; petals white. Colom

Vv. CAMPTOCLADA Mez & oak. Fedde Rep. Spec. Nov. 14:
247. 1916. Costa Rica
82. V. HODGEI L. B. “Suith, Caldasia 5:14, fig. 1948. Colom-

83. V. SUBANDINA (Ule) Sm. & Pitt. Journ. Wash. Acad. Sci. 43:
404. 1953. Cipuropsis subandina Ule, Verhandl. Bot. Ver. Bran-
denb. 48:148. 1907. Tillandsia subandina Mez ex L. B. Smith,
Contr. yee Herb. 98:16. 1932. Peru

84. V. RUBRA (R. & P.) Beer, Seca: 98. 1857. Tillandsia ru™
bra R. & he Fl. Per. 3:40, pl. 266. 1802. Vriesea albiflora Ule,
Verhandl. Bot. Ver. Brandenb. 48:141. 1907. Tillandsia rhododac”
tyla Mez, Fedde Rep. Spec. Nov. 16:76. 1919. Trinidad, northern
South America.

85. V. PEDICELLATA (Mez & Wercklé) Sm. & Pitt. Journ. Wash.
Acad. Sci. 43:403. 1953. Thecophyllum pedicellatum Mez & Wer-
ck1é, Bull. Herb. Boiss. IL. 3:136. 1903. Costa Rica.

86. V. SCEPTRUM Mez in DC. Mon. Phan. 9:606. 1896. Brazil.

87. V. EGREGIA L. B. Smith, Phytologia 4:355, pl. l, figs.

4-6. 1953. Venezuela.

88. V. HOSPITALIS (L. B. Smith) L. B. Smith, Phytologia 5:397.
1956. Tillandsia hospitalis L. B. Smith, Caldasia §216,*21¢8
1948. Colombia.

89. V. MAGDALENAE L. B. Smith, Phytologia 8:500, pl. 2, fig-

1966 Smith, Notes on Bromeliaceae 121

6-8. 1963. Colombia.
90. V. RANIFERA L. B. Smith, Phytologia 9:242, pl. 1, figs. 1
2. 1963. Costa Rica.
91. V. THYRSOIDEA Mez in Mart. Fl. Bras. dy Phe 355505. 1896,
Brazil.
2. V. ALTODASERRAE L. B. Smith, Contr. Gray Herb. 98:16, pl.
os figs, 1, 2. 1932. Pl. I, fig. 19: Spike x 1/4. Brazil.
ae V. HAEMATINA L. B. Smith, Arquiv. ee Estado S. Paulo n.
ser. 1:117, pl. 123. 1943. Brazil.
We Vv. MONACORUM L. B. Smith, Arquiv. Bot. Estado $. Paulo n.
ser. 1:119, pl. 127. 1943. Brazil
95527: KILLIPIANA oD OM Ci, Peo lésls Diaieatie« dy Tae.
1-3. 1954. Colombia

ands

sellata Linden, Catal. 9. 1873. Vriesea tessellata E. Morr.
Belg. Hort. 32:381, pls. 14-16. 1882. Tillandsia reticulata Ba-
ker, Gard. Chron. III. 1:140. 1887. Vriesea reticulata Mez in
Mart, on: Brad, '3) pts 23 1557; 1894: V. mosenii Mez, op. cit.
558. V. alexandrae hort. Sander, Gard. Chron, 245. 1903. Pliet,
fig. 25: Section of spike x 1/2. Brazil

97. V. PARADOXA Mez in DC. Mon. Phan. 9: 604. 1896. pEaeets

1

99, V. TURBINATA (ies 5 ret S te. & Pitt. Vedi sh.
Acad. Sci. 43:403. 1953. Thecophyllum turbinatum Mez & aay
Bull. Herb. Boiss. II. 4:1122. 1904. Costa R

100. v. erie a L. B. Smith, Arquiv. Bot. Speer S. Paulo n.
ser. 1:118, pl. 125, fig. 1. 1943. Brazil.

Ol. Vv. TRILIGULATA Mez in Mart. Fl. Bras. 3, pt. 3:541. 1894.
Brazil.

102. V. MACULOSA Mez, Fedde Rep. Spec. Nov. 12:418. 1913.
Brazil.

oF V. STRICTA L. B. Smith, Arquiv. Bot. Estado S. Paulo n.
SOR. Ailes, pl. 133. 1965, Brasil.

ii: V. MINARUM L. B. Smith, Arquiv. Bot. Estado S. Paulo n.
Ser. 1:118, pl. 126. 1943. Brazil.

5. V. HELICONIOIDES (HBK.) Hook. ex Walp. Ann. Bot. 3:623.

Re « 731226.
1830, in Synonymy, not Renealmia disticha L. 1759. Vriesea fal-
kenbergii hort. Bull, Gard. Chron. 1:759. 1880. V. bellula hort.
Linden, Cat. no. 109:7. 1883. V. disticha ee as Rev. Gen. 3,

Pt. 2:304. 1898. Guzmania obtusa Rusby, Mem. N. Bot. se = i
212. 1927. Wriesea bullata hort. ex C. aoe ee ‘Bull.
Nat.Hort. Fran 3 1990; Pi, Il, fig. 4: Spike x Mi.

106. V. LONGICAULIS (Baker) Mez in Mart. Fl. Bras. 3, pt. 3
542. 1894, Vriesea longhes sensu Wawra, Oe sterr. Bot. Zeitschr.
30:220, 1880, not E. Morr. V. jonghei forma "capsulis secundis",

122 PHYTOLOGIA Vol, 13, mo. 2

It. Sax.-Cob. 168. 1883. Tillandsia Longicauls Baker, Journ
Bot. 26:80. 1888. Vriesea longicauli secunda Mez in Mart.
Fl. icone Rpts 32543. 543. 1894. Braz a.
107. PYCNANTHA L. B. Smith, Contr. Gray Herb. 154:38, pl. 4
figs. io-ia. 1945. oe
108... ee Mez & Setckic; Bull. Herb. Boiss. II. 4:866.
ager Costa R
V. pees LIFLORA (Wendl.) Ant. Wiener I1l. Gart. 5: 98.
aie “Tillandsia gladioliflora Wendl. Hamb. Gartenz 19:31
1863. Vriesea princeps hort. Linden ex Batal, Bertaaet, 26: :158.
1877; E. Morr. Belg. Hort. 30:216. 1880, nomen. BRE & OOS eee
Spike x 1/4. Guatemala to Colombia
110. V. PACHYSPATHA Mez & Wéxtkic, Bull. Herb. Boiss. II. 4:
867. 1904. Costa Rica
111. V. MACRANTHA hes & Werckle, Bull. Herb. Boiss. II. 4: 867.
ae Costa Rica
cee Ve OVANDENSIS Matuda, Cact. & Sucul. Mex. 2:78, fig- 51.
1957... Pls: 1, fie. 35 Spikes 1/4. Mexico
115>-¥. ACUMINATA Mez & Wercklé, Bull. Herb; Boiss. II. 4:868.
1904. Costa Rica.
114. V. SCHULTESIANA L. B. Smith, Bot. Mus. Leafl. Harvard 16:
197, ag gis: Colombia
1 CHYPHYLLA Mez & Wercklée, Bull. Herb. Boiss. II. 4:
869. 104. "desta Rica.
V. BRUNEI Mez & Werckle, Bull. Herb. Boiss. II. 4:865.

ae

17. V. APICULATA L. B. Smith, Phytologia 8:221, pl. 2, figs-
1-3. 1962. Pl. III, fig. 1: Leaf-apex x 1/4; fig. 2: Spike x
1/4. Costa Rica

18;. Vi LANCIFOLIA (Baker) L. B. Smith, Lilloa 6:386, pl. 2;
figs. 3, 4. 3941. Sue lancifolia Baker, Handb. Bromel.
202. 1889. Vriesea platen nii Mez in Mart. Fl. Bras. 3, pt-

546. 1894, in part, not as "Ee type. Brazil.

119.7 ¥. PARVIFLORA L. B. Smith, Arquiv. Bot. Estado S. Paulo
Sees E219, 910. 125, tte. ie. 1943. Brazil.

120. ¥. MACROSTACHYA (Bello) Mez in DC. Mon. Phan. 9:601.
1896. Caraguata macrostachya Bello, Anal. Soc. Esp. Hist. Nat.
12:122. 1883. Neovriesia macrostachya Britton in Britton & Wil-
son, Sci. Surv. Porto Rico 5:142. 1923. Cuba, Puerto Rico,
Venezuela.

121. V. FENESTRALIS Linden & André, I1l. Hort. 22:124, og
215. 1875. Tillandsia fenestralis Hook. f. Bot. Mag. 112:p
6898. 1886. Vriesea fenstrata hort. Rev. Hort. 66:285. tans no-
men. V. hamata L. B. Smith, Arquiv. Bot. Estado S. Paulo n. ser-
1:59, pl. 77. “77. 1941. Pl. III, fig. 8: Leaf-apex x 1/4; fig. 9:
Spike x 1/4. Brazil.

. ATRA Mez in Mart. Fl. Bras. 3, pt. 3:543, pl. 101.

ONGHEI (K. Koch) E. Morr. Belg. Hort. 28:257. 1878.
Encholirion jonghii Libon ex K. Koch, Allg. Gartenz. 22. sods
nomen. Tillandsia jonghei K. Koch, Wochensbhr. 11:91. 1868.
Vriesea xiphion Platzmann ex E. Morr. Belg. Hort. 24:291. 1874,

1966 Smith, Notes on Bromeliaceae 123

nomen. Encholirion yqenit May, Rev. Hort. 50:140. 1878, probably
an error for "jonghe Vriesea gamba F. Mueller, P-gp 83:460.
250)... Pl. LV. tte, . Gea x-1/4.

Trinidad, Bra
AL. B. Smith, Arquiv. Bot. Estado S. Paulo
: ‘ Pl. IV, figs. 2: Leaf-apex x 1/4;
fig. 3: Section of spike x 1/4. Brazi
125. V. PLATYNEMA Gaud. Atl. Voy. Bonite pl. 66. 1843.
1. Flowers distichous throughout; sepals all exceeding the
floral bracts
2. Leaf-blades éssentiaily concolorous.
d and apiculate.

ieee saa tee eis eee ts ay
5. Scape slender; sere ‘scarcely more ‘phen 2 mm in
GLAMCEEY Cis eset eee ar. c. gracilior
4 cog acute .wn. Th Ness owes ets Re ea Seka Var. d. libonii
acuminate oi. bet oe Pee Pee re ONS acta VOU. ey ‘Tosea
2 “bee-biades not “concolorous

7. Leaf-blades pale-stri

pevecveseeeeecebewss Var. f. striata
7. Leaf te pare pee victor hackatt: gieen above, pale-striate
near > ee ees eee SERS O86 oie eek a ° vartenats
1. Flowers se fore of inflorescence polystichous with : epals
shorter than the floral bracts...... site ae a Var. h. wrightii
Var . Tillandsia platynema Griseb. Nachr. Ges
Wiss, beece 1864:19.

1865, as to basonym. Encholirion coxulli-
num J, Linden, Catal. 19:27. 1865. Vriesea corallina Regel, Gar-
tenfl. 19: 354, pl. 671. 1870. Encholirium corallinum Linden ex
André, I11. Hort. 18: 136, pl. 70. 1871. Tillandsia corallina K.
Koch, Ind. Sem. Hort. Berol. 1873, App. 4:5. 1874. Tillandsia
bicolor sensu Niederlein, Bull. Mens. Prod. Arg. 3 557 1890.
Pl. IV, fig. 4: Leaf-apex x 1/4; spe 5: Section of spike x 1/4.
West err eastern South Americ

- b. FLAVA Reitz, Anais hot: ‘ek: Barbosa Rodrigues 4:15.
1952. Beast.

ar. c. GRACILIOR L. B. Smith, Arquiv. Bot. Estado S. Paulo n.
im ny 121. 1943. Brazil.
d. LIBONII Mez in Mart. Fl. Bras. 3, pt. 3:553. 1894.
Beeler a liboni hort. ex Baker, Beirne Bromel. 220. 1889, no-
men. Bra azil (7).

ar. e. ROSEA (hort. ex Antoine) Mez in Mart. Fl. Bras.
3:552. 1894. Vriesea rosea E. Morr. ex Mez, loc. cit., nomen.
ar. f£. STRIATA (Wittm.) Wittm. ex Mez in Mart. Fl. Bras. <
pt. 3:553. 1894. Vriesea corallina var. striata Wittm. Bot.
ae 13, Beibl. 29: $91." Brazil.
- VARIEGATA (Guillon) Reitz, Anais Bot. Herb. Barbosa
Rodrigues 4:15. 1952. Encholirion roseum variegatum Guillon,
Rev. Hort. 55:470. 1883. Brazil.

ar. h. WRIGHTII (L. B. Smith) L. B. Smith, Phytologia 7:174.

35° pt.

12h PHYTOLOGIA Vol. 13, mo. 2

2 Guzmania wrightii L. Smith, Contr. Gray Herb. 117: Lbs
1937. Vriesea wrightii gma Mem. Soc. Cub. Hist. Nat. 15:
oe ae ub
V. WAWRANEA Ant. Phyto-Icon. 1, pls. 1, 2. 4864, Tite
ore wawranea Baker, Journ. Bot. 26:105. 1888. Brazil.

27. V. BITUMINOSA Wawra, Oesterr. Bot. Zeitschr. 42254705
1862. Tillandsia platynema sensu Baker, Journ. Bot. 26:106.
1888, in part. Vriesea platynema Wittm. Bot. Jahrb... 13
Beibl. 29:21. 1891. Pl. IV, fig. 6: Base of spike x 1/4. Venez-
uela, Brazil.

128. V. REGNELLII Mez in Mart. Fl. Bras. 3, pt. 3:548, pl.
102. 1894. Brazil.

129. V. NUTANS iL. B. Smith, Phytologia 7:175, pl. 2, figs. 65
7. 1960. Costa

130. V. CRASSA te in Mart. Fl. Bras. 3, pt. 3:566, pl. 104.
1894. Brazil.

131. V. DENSIFLORA Mez in Mart. Fl. Bras. 3, pt. 3:567. 1894.
Brazil

132. V. DISSITIFLORA (C. Wright) Mez In DC. Mon. Phan. 9: 604.
1896. Tillandsia dissitiflora C. Wright in Sauvalle, Anal. Acad.
Ci. Habana 8:54. 1872. Vriesea erici Harms, Notizblatt 12: 533.
1935. Cuba.

133. V. TUERCKHEIMII (Mez) L. B. Smith, Contr. Gray Herb. 17:
33. 1937. Tillandsia tuerckheimii Mez in Urb. Symb. Ant. 7:174.
1912. Pl. IV, fig. 15: Section of branch x 1/4; fig. 16: Sepal x
12 ftocr 7% Petal 2 ae ke ape iola.

134. V. SPARSIFLORA L. B. Smith, Contr. Gray Herb. 95:48, pl.
10, —- do 241934) «Bracis.

135. V. TRIFLORA Sm. & Pitt. Journ. Wash. Acad. Sci. - 403.
1953. Thecophyllum panniculatum Mez & Werckle, Bull.

a IL. 4:1123. 1904, not Vriesea paniculata Mez Nae “Costa
ca.

136. V. DISCOLOR (Mez & Wercklé) Sm. & Pitt. Journ. Wash.
Acad. Sci. 43:402. 1953. Thecophyllum discolor Mez & Werckle,
Fedde Rep. Spec. Nov. 14:246. 1916. Costa Rica.

137. V. SWARTZIL (Beker) Mez, Engl. Pflanzenreich IV. 32:400.
1935. Tillandsia swartzii Baker, Journ. Bot. 26:12. 1888. . Ti2-
landsia j Dank Le ta Sw. ex Baker, loc. cit. as synonym, not he ss

762. Jamaica
138. 4, RINCENS (Griseb.) Harms, Notizblatt 10:801. 1929
1866

Tillandsia ringens Griseb. Cat. Pl. Cub. 255. ee chagresi-
ana Baker, Journ. Bot. 26:1 88. a veitchii Baker, Handb.

Ss.
V. MORRENII Wawra, Oesterr. Bot. Zeitschr. 30:219. 1880-
Vr ee morrenii var. disticha Wawra, op. cit. 220. Tillandsia
morrenii Baker, Journ. Bot. 26:139. 1888. Brazi
140. V. RUSCHII L. B. Smith, Arquiv. Bot. Estado S$. Paulo n-
ser. .1:59, pl. 78. 1941. Brazil.
141. V. HOEHNEANA L. B. Smith, Proc. Am. Acad. 68:150, pl- 1»

Smith, Notes on Bromeliaceae 125

1966
figs. 11-13. 1933. Vriesea Eweedteana a? in DC. Mon. Phan. 9:
608. 1896, in Sa (?), not as to bas Vite pan listé sensu

Bot. Mus. Puiises 07, eg: 1925, not E. Morr.

AMAZONICA (Baker) Mez in Mart. Fl. Bras. 2 Bo. 3 ooAy
Mart. ex Schult. in R. & S. Syst. 7:
Journ. Bot. 26:108. 1888. Vrie-
sea gigantea Mez in Mart. Fl. Bras. 3, pt. 3:566. 1894, not Gaud.
» Guiana, Amazonian Brazil.
143. V. ITATIAIAE Wawra, Oesterr. Bot. Zeitschr.

Hoehne, Album Sec.
Brazil.

30:221. 1880.

a
Tillandsia itatiaiae Baker, Journ. Bot. 26:110. 1888. Vriesea
ttm.” Bot.” Janurb. 13, Beibl. 29:20. levi. * El. Vv,

schenckiana Wi
fig. 1: Branch x 1/4. Brazil.
th, sp. nov. * camptociada Mez &
Wercklé, < cui affinis, inflorescentia sales SaaS eriret basi-
bus Balt ahd elongatis multibracteatisque differt
LANT flowering 9-10 dm high. VES rosulate, ee 8 = long,
diitety and obscurely lepidote throughout; ehe aths broadly el-
liptic, ca. 15 cm long, nearly black seam Pica anaes,
broadly acute with a stout involute te gat '< p, 7-8 cm
flat. SCAPE erect, rather slender; aoa “b Les strict, imbri-
others broadly ovate with a

obconic, to 10 mm long in fruit.
long, ecarinate, coriaceous, even; petals
Stamens, white, musky and ee at night PCL
cylindric, acute, beaked, 3 cm long. Pl. V, Fig. 2: Inflores-
cence x 1/6; fig. 3: Sepal x

LOMBIA: Narifo: Epiphytic, Barbacoas, 17 November 1946, M.
B. Foster 2193 (US,

145. V V. CORNUS- pet ackacbaer Univ. Hamburg, Abh. Geb. Aus-
landsk. 61, Reihe C, Naturwiss. 18:82, figs. 79, 89. 1956.
Salvado
514, 146. v. HIEROGLYPHICA (Carr.) E. Morr. I11l. Hort. a u: pl.
1884. Massangea hieroglyphica Carr. Rev. Hort. 50:175,

: Tillandsia hieroglyphica Baker, Journ. Bot.
er, H

mel. 149. 1889, nomen. M. ti
nomen. Pl. V, fig. 6: Leaf-apex x 1/4; fig. 7: Branch x 1/4.
Brazil,
147. V, PENDULIFLORA L. B. Smith, Arquiv. Bot. Estado S. Paulo
Re ser. 1:120, pl. 129. 1943. Brazi

148. V. SEGADAS-VIANNAE L. B. Smith, Smithsonian Misc. Coll.
126:35, 120, fig. 40. 1955. Brazil.

149. V. WERCKLEANA Mez, Bull. Herb. Boiss. II. 3:136. 1903.

126 PHYTOLOGIA Vol. 13, no. 2

Pl. V, fig. 8: Apex of branch x 1/4. Mexico, Guatemala, Costa
Rica.
150 . We Apter eney Suessenguth, Bot. Arkiv 39:384, fig. l.
1939. Costa

Lis Ve SANGUINOLENTA Cogn. & March. Pl. Ornem, pl. 52. 1874.
Tillandsia sanguinolenta Baker, Handb. Bromel. 226. 1889... Vrie-
sea alfarovii Mez, Fedde Rep. Spec. Nov. 14:247. 1916. Encholi-

France V. 3:349. 1930. Vriesea urbaniana Harms, Notizblatt 12
647 1995; “Pi. V, figa 9: Branch x 1/5. Costa Rica to Colombia,
Greater Antilles.

152. V. LONGISCAPA Ule, Bericht. Deutsch. Bot. Gesellsch. 18:
323. 1900. Brazil.

153. V. CEARENSIS L. B. Smith, Phytoligia 7: 255, pie 1, fige-
11, 12. 1960. Brazil.

154. V. HYDROPHORA Ule, Arch. Mus. Nac. Rio de Janeiro 10:189.
1899; Bericht. Deutsch. Bot. Gesellsch. 17:2. 1899. Brazil.

5. V. PASTUCHOFFIANA Glaziou ex Mez in Mart. ¥1, Bras. 35
pt. 3:564. 1894. Tillandsia pastuchof* tana hort. ex Baker,
Handb. Bromel. 226. 1889, nomen. Brazi

156. V. PECTINATA L. B. Smith, Lilloa 6:387, 393, pl. 1, figs-
10-13. 1941. Guatema

157. V. CLAUSSENIANA (Baker) Mez in Mart. Fl. Bras 3. pte
545. 1894. Tillandsia clausseniana Baker, Handb. ‘ironed. 313.
1889. Brazi
1S8;v. WOODSONIANA L. B. Smith, Ann. Missouri Bot. Gard. 26:
275, pl. 20. 1939. Panama

Vv. PACHYCHLAMYS Mez in DC. Mon. Phan. 9:594. 1896, . Tab
landsia pachychlamys Baker ex Mez, loc. cit., nomen. British
Guiana.

160. V. TONDUZIANA L. B. Smith, No. Am. Fl. 19:166. 1938;
Lilloa 6:397, pl. 2, figs. 18, 19. 1941. Costa Rica
161. V. VANHYNINGII L. B. Smith, Phytologia 7: 173, eg
figs. 1-3. 1960. Pl. V, fig..10: Leaf-apex x 1/4; i. af Spike
x 1/4. Mexico.
162. V. UNILATERALIS (Baker) Mez in Mart. Fl. Bras. 3, pt. 3:
545. 1894. Tillandsia unilateralis Baker, Journ. Bot. 26:105.

Le

16%. ¥, none AS B. Smith, Phytologia 7:174, pl. 2, figs. 4»
5. 3960. Costa

164; Vv, eters censts (Baker) Mez in DC. Mon. Phan. 9:596.
1896. riiessts guadelupensis Baker, Handb. Bromel. 213. 1889.
Neovriesia guadalupensis Britton ex Stehlé, Fl. sate ead 1:212.
1936, nomen. Lesser Antilles.

165. V. oh peste Wittm. Bot. Jahrb. 11:69. 1889. Tilisesees
subsecunda Baker andb. Bromel. 217. 1889. Vriesea diminu
Mez & = ry Herb. Boiss. IL. 4:869. 1904. Costa tte
Panam

166. V. PITTIERI Mez, Bull. Herb. Boiss. II. 3:135. 1903.
Costa Rica.

1 . AMPLA L. B. Smith, Phytologia 8:497, pl. 1, figs. 4-®
1963. eoaté Rica.

1966 Smith, Notes on Bromeliaceae 127
68. V. VIRIDIFLORA (Regel) Wittm. ex Mez,

1 Engl. Se
RVG et aOl. 1995,” Pita Sem. Hor

cairnia viridiflora Regel, Ind.

Tillandsia viridiflora Regel, how. fee

nomen. Vriesea viminalis E. Morr.

eos 20ee, “PEL Vy “Pie; 32

British Guiana, Venezuela, Colomb
LOS sV MARNIER-LAPOSTOLLEI Gy as) Smith, Bromel. Soc. Bull.

14:108, figs. 1964. Pl. V, fig. 13: Spike-apex x 1/4. Costa
Rica.

e 1/4. Central America,

170. V. HAPLOSTACHYA pi Wright) L. B. Smith, Contr.
Herb. 114: 9, pl. 2, fig. 1936. Tillandsia Sasruaramee Cc
Wright in Seinduiie’ eat: ie: Ci. Habana 8:73. 1871. Cuba.

171. V. CHIAPENSIS Matuda, Anal. Inst. Biol. Mexico 23:130.
1953. Mexico

7a Vy GRAMINIFOLIA Mez & Wercklé, Bull. Herb. Boiss. II. 4:
868. capa Costa Rica.
3. V. BRASSICOIDES (Baker) Mez in DC. Mon. Phan. 9:598.

Tillandsia brassicoides Baker, Journ. Bot. 26:12. 1888.

iif

174, V. FIBROSA L. B. Smith, Mem. New York Bot. Gard. 9:315,
fig. 59. 1957. Venezuela.
17 S L. B. Smith, Bot. Mus. Leafl. Harvard 12:121,
fig. 1946. Colombia.
- V. PLATZMANNII E. Morr. Belg. Hort. 25:349, pl. 23. 1875.
Tillandsia platzmannii Baker, Journ. Bot. 26:104. 1888. Brazil.
177 - VERRUCOSA L. B. Smith, Phytologia 5:44, pl. 5, figs.
4, 5. 1954. Colombia
178. V. SCHIPPII e B. Smith, Contr. Gray Herb. 98:18, pl. 5,
figs. 5, 6. 1932. British Honduras.
179. V. MACROCHL re Mez & Wercklé, Bull. Herb. Boiss. II. 4:

ANTHA Cones) Mez in Mart. Fl. Bras. 3, pt. 3:544
dsia oligantha Baker, Journ. Bot. 25:345. 1887.

18
1894, Tillan
Brazil,

181. V. BROADWAYI L. B. Smith, Proc. Am. Acad. Arts & Sci. 68:
149, pl. 1, figs. 9, 10. 1933. Pl. V, fig. 14: Leaf-apex x 1/4;
fig. 15: Spike x a}h. Trinidad.

182. V. RACINAE L. B. Smith, Lilloa 6:387, 413, pl. 3, figs.

>
5, 6. 1941. Pl. V, fig. 16: Leaf x 1/4; fig. 17: Spike x 1/4.
Brazil,
183. V. VEXILLATA L. B. Smith, Bot. Mus. Leafl. Harvard 17:72,

Pl. 21, figs. 1, 2. 1955. Colombia

184, = SPLITGERBERI (Mez) Sm. & ‘Pitt. Journ. Wash. Acad. Sci.
43:403. 1953,
he precophyllum spi splitgerberi pois ak Evolution 2: 60:

: ’ DIFFUSA Sik. & pice Journ. Wash. ae Wel. 43:402.
Thecophyllum 1 axum Mez & Werckle, Bull. Herb. Boiss. Il.
4: 1123. 1904, not Vriesea laxa Mez, 1896. Costa Rica
186. V. CAPITULIGERA (Griseb.) Sm. & Pitt. Journ. Weel Aced.

128 PHYTOLOGIA Vol. 13, no. 2

Sci. 43:402. 1953. Tilia ae a agrien Cat. P1.. Cub.
254. 1866. T. fastuo a Andr Brome ey ekooes
68.

mn. Mez, lo cit.
a chachartenie capituligera Harms, Notizblatt 10:804. 1929. _The-
cophyllum capituligerum L. B. Smith, Contr. Gray Herb. 98:14.
1932. T. fastuosum Mez, Engl. Bera oadal ot Iv. 32:423, fig. 90.
1935. Pl. V, fig. 20: Spike x 1/4. West Indies, Venezuela,
Colombia, Ecuador.
. V. CRASSIFLORA (Mez & Werckle) Sih. PLtts Toute. Wash.
Acad. Sci. 43:402. 1953. Thecophyllum crassiflorum Mez & Wer-
cklé, Bull. Herb. Boiss. II. 3:1 138. 1903. Costa Rica
188. V. CYLINDRACEA (Suessenguth & Goeppinger) Sm. "’ Pitt

Journ. Wash. Acad. Sci. 43:402. 1953. Thecophy cylindraceum
Suessenguth & nits age WR Bot. Jahrb. 72:292. 1942. Costa Rica.
i Pi

953. Thecophyllum rubrum ec ei Mez é Werckle, Bull. Herb. Boiss. Il.
4:878. 1904, not Vries2a rubra Beer, 1857. Costa Rica.

190. V. BRACTEOSA (Mez & Wercklé) Sm. & Pitt. Journ. Wash.
Acad. Sci. 43:402. 1953. Thecophyllum bracteosum Mez & Werckle,
Fedde Rep. Spec. Nov. 14:246. 1916. Costa Rica.

191. V. NEPHROLEPIS Sm. & Pitt. Journ. Wash. Acad. Sci. 43: :403

139. 1903, not Vriesea werckleana Mez, 1903. Costa Rica
¥.. T (L. B. Smith) Sm. & Pitt. Journ. Wash.
Sci. 43:69. 1953. Thecophyllum montanum la 8. Smith, ar sce
Pub. Bot. 17:319, pl. 7. 1938. cuekene a Honduras.
193. V. VIOLASCENS (Mez & Wercklé) Sm. & Pitt. Journ.

Was
Acad. Sci. 43:403. 1953. Thecophyllum violascens Mez & eg:
Bull. Herb. Boiss. II. 4:877. 1904.

194. V. LEUCOPHYLLA L. B. Smith, bei Ek “32498, pl. 2;
figs. 1, 2. ieee sta Rica

195. V. SPECTABILIS (Mez & “Werckis) Sm. & Pitt. Journ. Wash.)
Acad. Sci. 43:403. 1953. Thecophyl lum spectabils Mez & Werckle,
Bull. Herb. Boiss. Il. 4:873. 1904. Costa Ric

v. COMATA (Mez & Wercklé) Sm. & Pitt. fe Wa sh. Acad.

Sci. 43:402. 1953. Thecophyllum comatum Mez & Wercklé, Bull.
Herb. Boiss. II. 4:871. 1904. Costa Rica.

197. V. ANTILLANA Sm. & Pitt. Journ. Wash. Acad. Sci. 43:403.
1953. Guzmania urbaniana Mez in DC. Mon. Phan. 9:920.
Thecophyllum urbanianum Mez, Bull. Herb. Boiss. iC B25 1903,

not Vriesea urbaniana Ha 1935. Lesser Anti
‘ JOHNSTONIT corte = & Pitt. Journ. Wash. Acad. Sci-
43:402. 1953. Thecophyllum johnstonei Mez, Bull. Herb. Boiss-

Il. 4:872. 1904. Trinidad, pa Ps (Isla Margarita).

99. V. NOTATA Sm. & Pitt. Journ. Wash. Acad. Sci. 43:403.
1953. Thecophyllum pittieri Mez, Bull. Herb. oe TY." 4s tons
1903, not Vriesea pittieri Mez, 1903. Costa Ric

200. V. HYGROMETRICA (André) Sm. & Pitt. oe Wash. Acad.
Sci. 43:402. 1953.

Var. a. HYGROMETRICA. Caraguata hygrometrica Andre, Enum.
Bromél. 6. Dec. 13, 1888; Rev. Hort. 60: 566. Dec. 16, 1888.

1966 Smith, Notes on Bromeliaceae 129

Guzmania hygrometrica André ex Mez in DC. Mon. Phan. 9: oa, prem
Thecophy1lum hygrometricum Mez, Bull. Herb. Boiss. II.
1903. Colombia

Varo bs ANGUSTIFOLIA L. B. Smith, Phytologia 8:222. 1962.
Costa Rica

201, Vs " PAUPERA (Mez & Sodiro) Sm. & Pitt. Journ. Wash. Acad.
Sci. 43:403. 1953. Thecophyllum pauperum Mez & Sodiro, Bull.
Herb. Boiss. II. 4:876. 1904. jecuador.

202. V. PICTA (Mez & Wercklé) Sm. & Pitt. Journ. Wash. Acad.
Sci. 43:403. 1953. Thecophyllum pictum Mez & Werckle, Bull.
Herb. Boiss. IL. 4:874. 1904, Costa Rica

203. V LINEATA (Mez & Wercklé) Sm. & Pitt. Journ. Wash. Acad.
Sci. 43:402. 1953. Thecophyllum lineatum Mez & Werckle, Bull.
Herb. Boiss. II. 4:875. 1904. T. balanophorum var. subpictum
Suesseng. Bot. Jahrb. 72:291. 1942 (?). Costa Rica

204. V. IRAZUENSIS (Mez & Wercklé) Sm. & Pitt. iad,
Acad. Sci. 43:402. 1953. Thecophyllum irazuense Mez & Marcia
Bull. Herb. Boiss. II. 3:138. 1903. Costa Rica

0534. Magy ei tee (Mez) Sm. & Pitt. Journ. Wash ar Sci.

5 i i :

43:403. 1953. zgmania ororiensis Mez in DC. Mon. Phan 7917.
1896. Thecoph Tie ororiense Mez, Bull. Herb. Boiss. II. 3:131
1903. T. kupperi Suesseng. & Goeppinger, Bot. Jahrb. 72 292
1942. Vriesea ku Sm. itt. Journ. Wash. Acad. Sci. 43:

206. V. CAPITATA (Mez & peek Sm. & Pitt. Journ. Was
Acad. Sci. 43:402. 1953. hecophyllum capitatum Mez & meted
Bull. Herb. Boiss. II. 4: oe 1904. Costa Rica

07. V. ATTENUATA Sm. & Pitt. Journ. Wash. hae. Sci. 43:401.
1959s. The ecophyllum acuminatum L. B. Smith, Contr. Gray Herb.
117: 30, pl. 2, figs. 28, 29. 1937, not Vriesea acuminata Mez &
Werckle, 1904. Costa Rica

08. V. STENOPHYLLA (Mez & Wercklé) Sm. & Pitt. Journ. Wash.
Acad. Sci. 43:403. 1953. Thecophyllum stenophyllum Mez &

43:402. 1953. Guzmania balanophora Mez in DC. Mon. Phan. 9:918.
1896, Thecophyllum balanophorum Mez, Bull. Herb. Boiss TL. 3
131. 1903. Costa Rica
210. V. VIRIDIS (Mez & Wercklé) Sm. & Pitt. Journ. Wash. Acad.

Sci. 43:403. 1953. Thecophyllum viride - & Werckle, Bull.
Herb. Boiss. IL. 4:872. 1904. Costa Ric

211. V. HAINESIORUM L. B. Smith, ‘eknieela 8:498, pl. 1, fig.
7-10. 1963,

212. Vv. WILLEAISIE 1. B. Smith, Phytologia 6:193, pl. 1, figs.
1, sae te Costa Rica

13
Sci. 43: 403. 1953. Thecophy11um iat L. B. Smith, Contr
ie aT

Thess ae sintenisii Mez, Bull. Herb. Boiss. II. 3:131. 1903.

130 PHYTOLOGIA Vol. 13, no. 2

Puerto Rico

2I5 N: SINGULIFLORA (Mez & Werck1é) Sm. & Pitt. Journ. Wash.
Acad. Sci. 43:403. 1953. Thecophyllum singuliflorum Mez & Wer-
ck1¢é, Bull. Herb. Boiss. II. 4:870. 1904. Costa Rica.
& Wercklé) my & Pitt. Journ. Wash. Acad.
Sci. 43:403. 1953. Thecophyllum vittatum Mez & Wercklé, Bull.
Herb. Boiss. II. 4:871. 1904. Costa Rica.

217. V. MALZINEL E. Morr. Belg. Hort. 24:313, pl. 14. 1874.
Tillandsia malzinei Baker, Rep. Kew Gard. 1878:59. 1879. Pl. VI,
fig. 1: Leaf x 1/4; fig. 2: Spike x 1/4. Mexico.

nN
<
<
be
tar
>
iz
Ld
=
@

N

Subgenus ALCANTAREA

218. V. EXTENSA L. B. Smith, Arquiv. Bot. Estado S. Paulo
ser. 1:116, pl. 120. 1943. Vriesea regina sensu Wittm. cevtiatt:
Brazil

9. Vv. ALTA (Baker) E. Morr. ex Mez in DC. Mon. Phan. 9:617.
1896. Tillandsia alta Baker, Handb. Bromel. pag 4 pies 1 gee a ie 2
fig. 23: Flower and bract x 1/5. Panama, Jamaica (7).

220. V. GENICULATA (Wawra) Wawra, It. Max. 156, pl. 25 (except
the serrate leaves). 1866. Piatysta chys geniculate Wawra, Oes-

terr. Bot. Zeitschr. 12:345. Vriesea gigantea sensu Lem.
TIL: Bort. “4751. 5 1867, ee as to qercneeee. V. glaziou-
ana Len. I 4, Misc.:43, fig. 2. 1867 gigantea sen-
su Regel, Gartenfl. 16:385. 1867. V. regina sensu on.

. ser. 3:234, fig. 41. 1875. V. regina var. glaziouana Wawra,
Oesterr. Bot. Zeitschr. 30:218. 1880. V. gi a sensu Carr.
Rev. Hort. 57:387. 1885 (7). Tillandsia regin a sensu Baker,
Journ. Bot. 26:139. 1888, in part. Vriesea vasta Mez in Mart.

Fl. Bras. 3, pt. 33572. 1094. Pl. T, ee ¥¥ Spike x 1/4.
Brazil.

221. V. REGINA (Vell.) Beer, Bromel. 97. 1857. Tillandsia re~
gina Vell. Fl. Flum. 136. 1825; Icon. 3:pl. 142. 1835. T. blokii

nfam. ed
blokii Mez, Engl. Pflanzenreich IV. 32:405. 1935. Pl. II, fig.
1: Section of nda -. bgp a
222. V. BRASILIANA L. m ith, Arquiv. Bot. dager ’ Paulo

n. BOP. T3315, Pr. uy. isis Vriesea regina Mez in Ma Fl.
Bras. 3, pt: 3: 569. 1894, in part. Pi. Il, fig. 2: nie ny spike
x sk Brazil.

223. V. GIBBA L. 4 Smith, Contr. U. S. Nat. Herb. 29:524,
fig. 80. 1954. Jamaica.

224.° ¥. ovine Carr. Rev. Hort. 60:58. 1888. Vriesea gi-
caeas sensu Lem. I1l. Hort. 14:sub pl. 516. 1867, as to descrip-

Tillandsia regina sensu Baker, Handb. Bromel. 227. 1889, in part.
Alcantarea imperialis Harms, Engl. & Prantl, Pflanzenfam. ed. 2.
15a:120. 1990, Fl. Vs chee @ lasowae x 1/43 fig. 5: Branch x
1/4. Brazil.

1966 Smith, Notes on Bromeliaceae 131

».4
(by species numbers of Vriesea)

TAREA imperialis 224; regina 221.
cheatuAtA hygrometrica 200; macrostachya 120; sintenisii 214.
CIPUROPSIS subandina 83.

ENCHOLIRION soealt tien 125; liboni 125d; roseum variegatum
125g; sanguinolentum 1
ENCHOLIRIUM corallinum 125.
ZMANIA balanophora 209; capituligera 186; fastuosa 186;
grometrica 200; ororiensis e003 sintenisii 214; splitgerberi re
banian

GUZVRIESEA magnifica - see appendix under Vriesea magnifica.

MASSANGEA hieroglyphica 146; Hegre pt is 146; tigrina 146

NEOVRIESIA guadalupensis 164; macrostachya 120.

PITCAIRNIA viridiflora 168.

PLATYSTACHYS geniculata 220.

SCHLUMBERGERIA capituligera 186

THECOPHYLLUM acuminatum 207; balanophorum 209; balanophorum
var. subpictum 203; bracteosum 190; capitatum 206; capituligerum

sco

um ye ororiense 20 53 panniculatum hale pauperum 201; pedicella-

um 85; pictum 202; pittieri 199; rubrum 189; singuliflorum 215;
sintenisii 214; epactabile 1933 ise, 184; standleyi 213;
stenophyllum 208; turbinatum 99; urbanianum 197; violascens 193;
viride 210; vittatum 216; werckleanum 191.

_TULLANDSTA ase 219; amazonica 142; amethystina oe appendicu-

a 66; appuniana 37b; arpocalyx 9; attenuata 65; barclayana 14;

spy rs M, 32: ‘piel 125; brassicoides 173; capituligera 186;
carinata 31, 38; var. constricta 40; cereicola a chontalensis
64; chrysostachys 80; clausseniana 157; corallina 125; corcova-
densis 61; cre nulipetala 65; crousseana 50; , pil 48; didis-

gracilis Griseb. 28a; gracilis
EYE uadelupensis 164; incurvata 34; inflata 35, 36; itatiaiae
143; jong 3; lacera 68; lancifolia 118; lateritia 14; laxa

127; ay AREY 176; poenula ; procera 28 ne Pp tt 38,

47; recurvata Baker 48 gina 220, 221, 224; reticulata 96; rho-
dodactyla 84; ringens 138; robusta 4; rodigasiana 23; rubra 84
Sanguinolenta 151; saundersii 27; scalaris 56; sation 5

132 PHYTOLOGIA Vol. 13, no. 2

simplex 55; splendens 37; var. formosa 37b; spuria 64; stenosta-

vittata 37; warmingii 50c; wawranea 126; zebrina 37.

VRIESEA acuminata 113; albiflora 84; alexandrae 96; alfarovii
151; alta 219; altodaserrae 92; amazonica 142; amethystina 46;
ampla 167; antillana 197; apiculata 117; axpendi culata: argentin-
ensis 20c; arpocalyx 9; atra 122; attenuata 207; aurea

a coe 115; brachystachys 38; bracteosa 190; brasilia
22; brassicoides 173; broadwayi 181; brunei 116; aul 24;
rat A 105%
a na 20c; camptoclada 81; capitata 206; capituligera
; 38; var. constricta 40; var. inflata 35; catha-

80b; citrina Baker 23;

50a; var. recurvata 50a; confusa 144; corallina 125a; var. stti-=
ata 125f; corcovadensis 61; cornus-cervi 14

cryptantha 80; cylindracea 188; cylindri

Glatieiedataes 69; diffusa 183;
diminuta 165; discolor 136; dissitiflora 132: Wate riche oe

s 50; var. d. color
a 50b; var. warmingii 50c: beti : 132: segthtcdacteion
31; eumo oepb 32; extensa 218.
Ryperamog 105; fenestralis 121; fenestrata 121; fibrosa
174; flamm . Smith 63; fosteriana 124; fragrans 70; fri-
burgensis 20; var. paludosa 20b; var. tucumanensis 20c; funebris

hillegeeriana 224; hitchcockiana 1; hodgei 82; hoehneana 141;
hospitalis 88; hydrophora 154; hygrometrica 200; var. angustifo-
lia ow
terica 45; imperialis 224; incurvata 34, 36; var. inflat

Bhi pee 35; interrogatoria 60; irazuensis 204; itatiaiae 143.

jimenezii 743 johnstonii 198; jonghei 106, 123; forma "capsu-

W

5: " krame ri 47b; kupperi 205; kupperiana 150
68; gece riggs languida 100; latissima i axa
44; leptantha lep a 189; leucophylla 194; lineata 2
peel gm nr -apntieligned Be wartelii 37b; eae ne 106;

1966 Smith, Notes on Bromeliaceae 133
VRIESEA (continued)

rae eects 106; longiscapa 152; lopezii 3; lubbersiana 16;
nen P12,

macrantha 111; macrochlamys 179; macropoda 35; macrostachya
120; maculosa 102; magdalenae 89; malzinei 217; marnier-lapostol-
lei 169; maxon igha 45; mesiana 55; minarum 104; modesta 30; mona-
corum 94; montana 192; morreniana 49; morrenii 139; var. disticha

neoglutinosa 25; nephrolepis 191; notata 199; nutans 129.

obliqua ma oligantha 180; olmosana 5; ororiensis 205;
ovandensis 112

pach ychlamys 159; pachyspatha 110; paludosa 20b; ey gees?
138; paraibica 40; paradoxa 97; pardalina 51; parviflora 1

debilis 28d; var. gracilis 28a; var. rubra 28b; var. tenuis 28c;
psittacina 47; var. sbdorge. bg rere 303 Var bracte is omnino cocci-
neis 47b; var. carinata 38; var. de color 47c; var. duvaliana 39;

var. erythrodactylon a: var hcl eletiiaes 49; "var. rubro-bracteata
7

racinae 182; ranifera 90; rauhii 2; reeurvare 48; regina 218,
220, 221, 222; var. glaziouana 220; regnellii 128; reticulata 96;
retroflexa 54; rhodostachys; ringens 138; robusta 4; rodigasiana
23; rosea 125e; rostrum-aquilae 34; rubida 61; rubra 84; rugosa
108; ruschii i 140.

Sanguinolenta 151; saundersii 20b, 27; scalaris 55, 56;
viridis 56b; sceptrum 86; Seuanetiecg 143; schippii 178; : Sohal
na 3; se

sc

simplex 55; sincorana a7 pe Be rp a 215; sintenisii 214; soci-
alis 175; soderstromii 77; sbatattioes 134; speciosa 37a; specta-
bilis 195; splendens 37; splendens major 37b; splendens var. for-
mosa 37b; var. longibracteata 37b; splitgerberi 184; squamosa 40;
Standleyi 213; stenophylla 208; stenostachya 78; stricta 103;
Subandina 83; subsecunda 165; suleata 75; swartzii 137

tequendamae 10; tessellata 96, 141; thyrsoidea 91; tillandsi-
Oides 8; tonduziana 160; triangularis 13; triflora 135; triligu-
lata 101; truffautiana a tucumanensis 20c; tuerckheimii 133;
turbinata 99; tweedieana ee tweedieana F. Mueller 20a, 23.

vagans 59; vanhyningii 1 MORE se 220; veitchii 138; ventrico-
Sa 61: verrucosa 177; vexillata 183; viainblis 168; violascens
493; viridiflora 168; viridis 210; vitellins 23: vittata 216;
Bc .
rmingii 50c; wawranea 126; werckleana 149; williamsii 212;
wonOncadans 158; wrightii 125h.

xiphion 123,

13h PHYTOLOGIA Vol. 13, no. 2

APPENDIX
Additions, seeping oa saqhndne taxa,
amed hybri
NOTE: Unless otherwise indicated, information about hander is
bf lepzenseich” Iv. 32:406-410. 5: BSB

from Mez
Bromeliad rn Bulletin. = A. B. Graf, Exotica ed. 3.
1963. = W. Richter, tes von heute und m

sabes pnt 1962.
names of hybrids and in the
Correct specific names can

use of specific names of
be found in the index above.

Vriesea aeris-incola Mez in AEE « Fl. Bras. 3, pt. 3:555. 1894
= TILLANDSIA AERIS-INCOLA sieges
x Africain C. Chevalier = V. at UBBERSII x MEPHISTO, BSB 1:21.
André, Rev. Hort. 41:300, pl. 1899 = INCURVATA x
(PSITTACINA x CARINATA).

albescens Alv. Silveira, Fl. Mons iae YM 32he 131; fig»: 2> 1931,
nomen, apparently an error for his glauca.

albo-nitens ted x Kew. Suppl. 1: 1906 (error) = VITIS
ALBO-NITENS hor
aloifolia seas ) Beer, Bromel. 95. 1857 = TILLANDSIA FLEXUOSA

we
anceps Lem. Fl. Serres 5:sub pl. 432. 1849 = TILLANDSIA ANCEPS

Lod
dreana hort. Duval, Rev. Hort. 66:135. 1894 = PSITTACINA x

LONGIBRACTEATA.

o-purpurea Alv. Silveira, Fl. Montium 2:Add. 3, pl. L3ky
a ~— 1931. Identity not resolve

aan Rh Sst Opoix, Rev. Hort. 71: 583. 1889 = (x REX) x
(x POEL
ort. Liege, Gartenfl. 42:340. 1893 = (CARINATA x

parecer: x ENSIFORMIS.
x aurora major hort. Liége ex Wittm. Gartenfl. 42:340. 1893;\*
PSITTACINA ; x ENSIFORMIS.
Belgica C. Chevalier, BSB 1:21. Identity?
x bijou hort. Duval, Rev. Hort. 65:213. 1893 - parents?
billbergia Lem. I1l. Hort. 16, Misc.:91. 1869 = TILLANDSIA
VIRIDIFLORA (Beer) agree
x Boetscheria, BSB 4:58 - parents un
botteri hort. ex ii Gartenfl. 33: 293. "1884 = TILLANDSIA
igo eR & Cham
brachystachys major tusets Liése, Gartenfl. 42:339. 1893 =
(CARINATA x BARILLETII) x CARINAT
x brachystachys splendens keay uval, Rev. Hort. 64:551.
1992 = CARINATA x PSITTACINA var. Searenaruank
bracteosa Beer, Bromel. 263. 1857, nomen = TILLANDSIA
CIRCINNATA Schlecht.
caespitosa E. Morr. ex mers Handb. Bromel. 213. 1889 =
TILLANDSIA MULTICAULIS Steu
x candelabrum, BSB 4:55 - ‘ parents unknown.

1966 Smith, Notes on Bromeliaceae 135

x rappel hort. pores? Rev. Hort. 66:285. 1894 =
ardinalis hort. Duval, Rev. Hort. 62:358. ge CARINATA x

Setreacraa var. RUBROBRACTEATA.

carinata var. wawra, BSB 5:43, nomen = ?

x Chantrieri Chantrier, R 279 = ? Ekeaves dark-banded.
x citrina hort. Opoix, Rev. Hort. 77:127..1905 = ?

x Closoniana hort. Liege, Gartenfl. 42:431. 1893 = BARILLETII
x (CARINATA x PSITTACINA).

x Closoniana brachystachys major, BSB 2:41 = (x MORRENIANA) x
BARILLETII

x Croix d'Honneur C. Chevalier, BSB 1:21 = ? Branched.
“pipet la
sec iat aie F. Mueller, Gartenfl. 42:737. 1893 = ?
inadequate.
Devansayana hort. Duval, Rev. Hort. 65:202. 1903 = CARINATA x
ier var. RUBROBRACTE

ATA.
cteur Lebel hort. Duval, Rev. Hort. 70:539. 1898 =
CONFERTA x ConkER?, X).

Description

x Donneai hort. Makoy ex Witte, Hybr. Brom. 8. 1894 =
SES x aed
Duchart 4 hort. Duval, Rev. Hort. 66:135. 1894 = ((CARINATA
x PSITTACTHA) x BARILLETIIL) x SPLENDIDA.

x Ducretii hort. Duval, Rev. Hort. 68:251. 1896 = ?
x Duvalii, BSB 1:21 = error for V. duvaliana E. Morr.?
x Duvalii major hort. Duval, Rev. Hort. 66:135. 1894 =
DUVALIANA x (x FULGIDA).
elegans hort. Duval, Journ. Soc. Nat. Hort. France 1892:706
= (BARILLETII x (CARINATA x PSITTACINA) ) x DUVALIANA.
elegans E rr. Rev. Hort. 64:551. 1892 = (BARILLETII x
(CARINATA x PSITTACINA)) x (x FULGIDA
a hort. et Rev. Hort. 69: 307. 1897 = ?

eros hort. Duval, 4 AS Hort. 69:140. 1897 = CORALLINA var.
ROSEAT = x ((CARINATA x PSITTACINA) x BARILLETII).
throdactylon var. striata Seidel, G 1734, nomen. Leaves
pale-stripe d.
Xx esperanza hort. nisi Rev. Hort. 69:140. 1897 =
(x KITTELIANA) x SAUNDERS
Espirito Santo Santo, G 173 ee
x Favorite , R >» R 279'= ie ected x ? Inflorescence simple.
Floral bracts keeled.
x fenestralis var. mortfontanensis Chantrier, BSB 13:99 =
TRALIS x TESSELLATA.
x Flaming | Sword, BSB 10:39 = SPLENDENS var. MAJOR x SPLENDENS
var. LONGIBRACTEATA.
x Flamme Richter, BSB 10:39; R 280, 282 (B), 304 = (x VIGIERI)
x ee TIL.

x flammea hort. Duval, Rev. Hort. 74:27. 1902 = (x VANGEERTII)
x aes

ammendes Schwert Richter, BSB 10:39; R 280 = SPLENDENS
ace aoe x SPLENDENS var. LONGIBRACTEATA.
X fulgida hort. Duval, Ill. Hort. 60:87, pl. 67. 1888 =

136 PHYTOLOGIA Vol. 13, mo. 2

at eee x DUVALIANA.
fulgida hort. Makoy, Witte, Hybr. Brom. 8. 1894 = INCURVATA
x ee MORRENIANA) .
. Devansaye, Rev. Hort. 69:163. 1897 =
deena ay x ENSIFORMIS.
x gemm val, Gartenfl. 42:340. 1893 = (BARILLETII x
CoaRINATA x PSITTACINA)) x (DUVALIANA x INCURVATA,
erbe de feu C. Cheva lier, BSB 1:21 = ? Tall. scape.
branched £ inflorescence. Red bracts
x Gi t Richter, BSB 10:39; R 280 = (x POELMANII) x (VRIESEA
Sr. = pai eracima) aoe VERSATLLENSIS) .
x ginoti Chantrier, BSB 13:99 = FENESTRALIS x ENCHOLIRION

glauca Alv. Silveira, Fl. Montium 2: Ada 3, 1931. Description
inadequate for classificat ion
glaucophylla Hook. Bot. Mag. 74:p1.4415. 1848 = TILLANDSIA

glaucophylla hort. ex Mez, Engl. ee EVio 322209:
1935 = HECHTIA TILLANDSIOIDES (André) L. Be Sait
* ploriosa hort. Duval, Rev. Hort. 66:135. en BARILLETII x
INCURVA'
geass Richter, BSB 10:39; R 280 = CORCOVADENSIS x

AMETHYSTINA x PSITT “s
x grandis Maron, Rev. Hort. 60:580 1888
x griesseniana h Rev. Hort. 70:231. ite

% eri

x "Heinrich Schmidt" Mez in DC. Mon. Phan. vi at "1896 =
(DUVALIANA x SOCMEALACELAa) x PSITTACINA.

HELICONIOIDES (105):

Var. b. POLYSTICHA L. B. Smith, Phytologia 8: aeet Costa Rica.

x Henrici hort. Duval, Rev. Hort. 66:239 ?
x "Henri Ducret" hort. Duval, evs Hort. 70:231. 1898 = ?
nieebalyohiens" marginata", G 1735, nomen. Leaves pale-banded.
HIEROGLYPHICA (146): V. hieroglyphica var. zebrina R Ruschi,
Bull. Mus. Nation. Hist. —: TL. 263547," fig. 1954.

x hoelscheriana, R 270 = Inflorescence simple, lax,
decurved.

x illustris Richter, R 280 = (x FLAMMENDES SCHWERT) x
SPLENDENS.

imbricata (Vell.) soa gnats 94. 1857 = ? Based on Til-
landsia imbricata Vell. almost certainly a Vriesea.- Not
collected again so far ab “Labed

x imperialis sag Duval, Rev: Hort. 73:53. 1901 =

(x oe x (Gc REX).
s hort. Leiden, Gartenfl. 41:1, p1l.1362. 1892)

Inspektor Kolb" — in DC. Mon. Phan. 9:571. 1896 =
PSITTACINA var. RUBROBRACTEATA x (ROSTRUM-AQULLAE x DUVALIANA) -
x "Inspektor estar Mez in DC. Mon. Phan.9:567. 1896 =

BARILLETII x GUTTATA.

Smith, Notes on Bromeliaceae 137

Gartenfl. 42:340. 1893 = BARILLETII

1966
x intermedia hort. Liege,

x FENEST ee
i hort. ex Focke, Pflanzenmischl. 530. 1881 =

CARINATA x x ea comat CINA
Kitteliana Wittm. Gartenfl. 39:326, fig. 62, 63. 1890 =
SIL.

BARTLLETTT x SAUNDERS
x Komet Richter, BSB 10:39; R 280 = CORCOVADENSIS x (x SCEPTRE
D'OR).
Soc. Nat. Hort. France

x Kram ro-fulgida hort. Duval, Journ
1893: 501 = = PSITTACINA var. RUBROBRACTEATA x (ROSTRUM-AQUILAE x
IANA) .

DUVALI
ni eee tteana hort., BSB 15:46. = AECHMEA RECURVATA (Kl.) L. B.

Liége ex Wittm. Gartenfl. 42:340. 1893 =

att odiensis hort.
habredeer *°¢G iacaone, x PSITTACINA).
Leonii hort. Duval, Rev. Hort. 74:27. 1902 = (x REX) x
(x MIRABILIS).
the following

x Leopoldi Witte, Gartenfl. 40:476. 1891 =
x Leopoldiana Wittm. Gartenfl. 42:340. 1893; 54:281, pl. 1539.

1905 = SPLENDENS x MALZINEI.
jindentt Lem, Til. Hort. 16:pl. 610. 1869 = TILLANDSIA CYANEA
Linden ex K. Koch.
Fi. Bras. 3, pt. 22395, nin 10s,

luschnathii Mez in Mart.
1894 = TILLANDSIA SPICULOSA Griseb.
Hort. 77:127. 1905 = ?

x luxemburgensis Opoix eae
- 69:336. 1897 = TILLANDSIA

macropetala hort. Rens "Hoe
VIRIDIFLORA (Beer) Baker.
maculata (R. & P. ) seein Bromel. 98.1857 = TILLANDSIA MACULA-

TAR.

usanne de Smet" hort. Gartenfl. 58:200. 1909 =

(x MTAABTLTS x (x VANGEERTII).

x magnifica hort. Makoy, Rev. Hort. Belg. 14:186. 1888 =
VRIESEA SPLENDENS x GUZMANIA ZAHNII = GUZVRIESEA MAGNIFICA M. B.
Foster, BSB 13:8

x Magnisiana Kittel & Wittm. Gartenfl. 38: 343, fig. 1889 =

BARILLETII x FENESTRALIS.
x Magnusiana Ind. Kew. suppl. 1:458. 1906 = the ore
Rev. Hort. 77:127. 1905 =

* majestica hort. Duval,
x Marechaliana hort. Makoy ex Witte, Hybr. Bromel. :. 1894 =
TNCURVATA x (x MORRENIANA) .
x Mariae André, Journ. Soc. Nat. Hort. France 1889:577; Rev.
Hort. 61: 300, pl. 1889 = CARINATA x BARILLETIL.

x Marie (error) = the above.
x "memoria Moensi" hort. Begs thd — Hort. 69:307. 1897 =

x Menelik hort. Liége, BSB 1:2
x = mephisto hort ..; BSB 1:2) = 4
minima hort. Duval ex Witte, Hybr. Bromel. 9. 1894 =

(x lieth x DUVALIANA.
Xx Mirabilis hort. Duval, Rev. Hort. 74:27. 1902 = VRIESEA
ENDENS x GUZMANIA ZAHNII = GUZVRIESEA MAGNIFICA M. “4 Foster.

x Moensiana hort. Makoy, Rev. cnet 70:246. 1898 =
x Morreniana, cf. species 49 abov

SPL

138 PHYTOLOGIA Vol. 13, n0.

morreno-barilletiana hort. Duval, 111. Hort. 36:103, pl. 91.
1688 = (x MORRENIANA) x BARI LLETII.
= Boren ee Mare hort., BSB 2:40 = the abov

ae ware

Cogn. arch. in =: Pl. Orn anne. 2:39. 1874 =
égemineth <FUSATCA Res & André

x nigricans hort. Duval, Rev. doce. 77:254. 1905 = ROSTRUM-
AQUILAE x ENSIFORMIS.

x oblige cf. species 53 ab

Papa Chevalier C; “Chevalier, “BSB 1:21 = (x MEPHISTO) x
PASTUCHOFETANA.

x perfecta hort., BSB 4:38; R 272, 280 = ?
dense. Floral bracts re

x Petersiana Wittm. Gartenf1. 44:456, fig. 92. 1895 =
BARILLETII x GUTTATA.

x Pleurvicii hort. Duval, Rev. Hort. vo 76: p. ? 1894 = ?
ort. Duval, Rev. Hort. 69: Bei 1897 =(x GLORLOSA)

Branched. Spikes

<< (s VANGEERTII).
x Poelmanii superba hort. Gartenfl. 58:200. ta = ae above.
x Polonia C. Chevalier, BSB 1:21; 15:69 (pl.

)>
x Pommerescheana Wittm. Gartenfl. 42:129, pl. ake 1893 “
SPLENDENS x (x MORRENI

ANA) .
x "President ee hort. Duval, Rev. Hort.

73:54. 1901 =
(x REX) x PLATYNEMA.
x President 0. Lamarche C. Chevalier, BSB 2:41 = INCURVATA X
BARILLETII.

x Prince Charles C. Chevalier, BSB. 432e*
pill bracts.

2? Branched. Salmon-
x Prince Leopold C. Chevalier, BSB 1:21 =?
—s. washed with yellow.

x procera bicolor Dutrie, Bull. Hort. 117. 1 ne 1947; BSB
11:62 = - ErTTEL LIANA) x (x BRA RACHYSTACHYS MAJOR

> psittacina hybrida Richter, BSB 10:38; R sabe = PSITTACINA

Branched. Bracts

x psittacina picta — ie ex Witte, Hybr. Brom. 10. 1894
= (x oe a BARIL

Ni ueeattake BSB 1:21 = error for var. RUBRO-
BRACTEATA probably.

PSITTACINA ve Vriesea psittacina var.

exilis Neum Ann. Soc.
Hort. Paris

pls
x_ps ei caese Sead hort. Duval, Journ. Soc. Nat. Hort
France 1893:30 (DUVALIANA x ROSTRUM-AQUILAE) x PSITTACINA-
pulverulento-lineata Carr. Rev. Hort. fer 89, fig. 18 8
(sterile) = VRIESEA subgenus ALCANTAREA
purpurascens hort. Bull, Gard. Chron.
NIDULARIUM INNOCENTII Lem.?

x Quintusiana hort. cope & ie a Tuinb. 1900:305, p1- 53
Rev. Hort. Belg. 66: 1894 =
ramosa

109%, 12413."

265. ey. nomen = TILLANDSIA UTRICULATA-

x rex hort. Duval, Rev. Hort. sia 1894:217, pl.; Rev- Hork®
79:570, 0; oie 1907 = (BARILLETII x (X MORRENIANA)) x (CARINATA x
PSITTACINA var. RUBROBRACTEATA).

1966 Smith, Notes on Bromeliaceae 139

x rex candelabra Chantrier, BSB 1:33 = (x REX) x ?
x rex major hort. Duval, Rev. Hort. 67:582. 1895 = (x REX).
x rex rubis hort. Duval, Rev. Hort. 70: se ae =
X rex superba hort. dts age Clic e

rnse

x
roezlii hort. Linden ex bekiee, reais Bromel. 207. 1889,
nomen = = Reptdigg ROEZLII E. Morr
oseo-lineata hort. Makoy, Rev. —_ 66:306. 1894 = ?
x sikre Richter, BSB 10:39; R 304 =
x St. Joseph, BSB 4:9 = ?
Sanctae-crucis S, ee Trans. Linn. Soc. II. 4:491. 1895 =
TILLANDSTA PARAENSIS
anderiana vittn. ehcweite! 46:377. 1897 = BARILLETII x
(PSITTACINA x CARINA
"sceptre d'or" pede, Gartenfl. 58:200. 1909 = SAUNDERSII x
(x GLORIOSA).
x sceptrum hort. (not Mez), Rev. Hort. 70:246. 1898 = above.
schlechtendalii Wittm. and var. alba Wittm. Bot. Jahrb. 11:69.
1889 = TILLANDSIA ANCEPS Lodd.
Setacea Hook. Bot. Mag. 60:pl. 3275. ae TILLANDSIA
tyre saieed TA Sw. var. VENOSISPICA Mez = T. COMPRESSA Bert. ex R.

ebertiana hort. Journ. Soc. Nat. Hort. France 1899:665 =
TILLANDSTA LEIBOLDIANA Schlecht.?
xX souvenir de Jos. Mawet C. Chevalier, BSB 1:24 =
x sphinx hort. Duval ex Witte, Hybr. Brom. 10. aan = FENES-
TRALIS x LONGIBRACTE
x splendens illustris hort., BSB 11:7 = SPLENDENS x
SCHWERT) .

(x FLAMMENDES
x splendens var. cayenne Lecoufle, BSB 13:105 (leaves cy ?
splendens var. major hort., BSB 11:7; R 269, 277 = horticul
tical selected clone.
SPLENDENS (37):
ar. c. STRIATIFOLIA M. B. Foster, Bromel. Soc. Bull. 5:92.
aede Venezuela
| x x splendida tionk. Duval, Rev. Hort. 64:551. 1892 = CARINATA x
* A).
etre E. Morr. Cat. Brom. Liége, 17. 1873 = TILLANDSIA
Whip mister Scheidw
x superba hort. Duval, Rev. Hort. 71:169, 511. 1899 =
(x KITTELI LIANAL x (x RE
fenuifolia Beer, fac 96. 1857 = ce gear er FLEXUOSA Sw. as
to description and T. TENUIFOLIA L. as to onym.
x tenuis hort. Edinburgh ex si "png Pt aceecoutcls TV¥c 32:
409, 1935 = = PER-PSITTACINA x CARIN
fessellata Sanderae hort. eae ony Hort. 65:202. 1893 =
GIGANTEA, arama

tessellata variegata hort. Gartenfl. 42:305. 1893 = eeaeigy

rete var. roseo-pictahort., R 277. Leaves with lar
T0Se€-red spots.

140 PHYTOLOGIA Vol. 13, no. 2

tricolor hort. ex Baker, Handb. Bromel. 109. 1889 =

PITCAIRNIA

Fl. Montium 2:pl. oH 1931 = error
for AECHMEA TRICOLOR Alv. Silveira, op. cit. 3

x trifurcata hort. Duval, Rev. ort 65221 ve

utriculata Regel, “Ind. Sem. Hort. Petrop. a,
UTRICULATA L

x Van Ackeri hort., BSB 11:61 = (x POELMANII) x

x Van Geertii hort. Duval, “ee Os 68:251. 1896; 1r61

= INCURVAT ATA x

= maker ae Truff. Ill. Hort. 36:73, pl. 87. 1889 =
DUVALIANA x CARINATA.

x Vigeri hort. Duval, Rev. Hort. 70:395. 1898 =

REX) .

TILLANOSEA

RODIGASIANA x

(x
x Bates erecta hort., we Li22ee oe saint eae Kot
x viminalis-rex hort., BSB 1:62 =
Morr. ex Baker, es Heke "304. 1889, nomen =

TILLANDSIA SPICULOSA Griseb.
x Hai tonis hort. ae BSB 1:21
Weyr ingersan ana ba ttm. eit.
BARILLETIT x
x Wioteana pony ie ex Wittm. Gartenfl. 42:340. 1893 =

BARILLETII x PSITTACINA.
a hort. ex Mez, Engl. Pflanzenreich IV. 32:667.. 19337

a 7, ¢1gc 1. ee =

Wirtean
error for the abov
x Witteana A. "Goval. Rev. Hort. 66:435. 1894 = DUVALIANA x

(x MORRENIANA
x "Witte genice?! Mez In DC. Mon. Phan. 9:572. 1896 = CARINATA

x ENSIFORMIS.
x Wittmackiana Kittel, Gartenfl. 37:553, pl. 1283. 1888 =
BARILLETIT x (x MORRENIANA
XX Veieslandsia Prof. Boul Lene C. Chevalier, BSB 1:23 = VRI

SEA ee SOUVENIR DE JOS. MAWET) x TILLANDSIA CAESPITOSA ate tas

& Cham. (= T. SaCEREAULTS

TE: The above compilation is doubtless far from compl ety and
should be greatly enlarged when the Bromeliad Society's list of
hybrids is published. However, it is cog that it will prove
useful as a general reference until then.

MISCELLANEOUS NOTES
TROPICAL AMERICA

BROMELIA REDOUTEL (Baker) L. B. Smith, comb. nov.

Bromelia karatas var. caulescens Redoute, Lil. 8:pl. 457. 1816.
Karatas redoutei Baker, Handb. Bromel. 3. 1

Bromelia caulescens Kuntze, Rev. Gen. 3,

889.
pt 6:2:202. 1898, nomen
ee

web Hort. Redoute (Herb.? basis of Redoute, Lil. plate

457 in enti
So

far, no sg ers specimen of Redouté's plant has been

1966 Smith, Notes on Bromeliaceae 1

found, so the type must be the plate and description for the pre-
kno

sent. If it were to be placed with any Known species, it would go
with B. serra Br ised « rather than with B. balansae Mez as former-
1 H wever, it fers from B. serra in its nearly naked sca

best to consider it one more of the many species described from
cultivation and never recollected in the wild.

COSTA RICA, PANAMA

TILLANDSIA OERSTEDIANA L. B. Smith, sp. nov.
Tillandsia , pantculata Cham. & Schlecht. var. costaricensis Mez in
DC. Mon. Phan. 9:703. 1896.

wilisidsis rubra R. & P. var. costaricensis (Mez) Mez, Pflanzen-
Se

reich 4
Tillandsia deppeana Steud. var. costaricensis (Mez) L. B. Smith,
Phytologia 5: aoe 1956.
deppeana Steud., cui affinis, foliorum laminis sans cg
bus subtus dense cinereo-lepidotis; spicis patentibus brevite
Stipitatis, floribus minoribus differt.

LANT flowering 6-15 dm high. LEAVES rosulate, many, 4-8 dm
long, bicolorous, covered beneath with persistent appressed cine-
reous scales; sheaths subelliptic, large; blades subligulate,
attenuate, to 4 cm wide. SCAPE erect; scape-bracts erect, sub-
foliaceous, dense ely imbricate. INFLORESCENCE laxly bipinnate;
primary bracts exceeding the sterile mostly bracteate aoa of

o
ia]
om
Q
rt
n
a
la]
=
Q
wf
ct
0)
a
Q
°
5
re)
@
»
iia
=]
aq
rt
a.
0)
va]
+
6
=
on
n
is)
ina
ct
a
(1)
n
Cy

long, bee ee the sepals, coriaceous, even, ick ocitae: Suectusiaaials
flowers subsessile. SEPALS lanceolate, acute, 25 mm long, cari-
Leaf-

te i
apex x 1/4; fig. 6: Spike x 1/4; fig. 7: Posterior sepals

COSTA RICA: Lankester 1526 (GH) ; Echeverria 221. (GH). Oriri,
s. n. (GH); alt. 1200

—— S- 2a tc. ype).

- PANAMA: Chiriquf: “Boquete, alt. 1150 m, June 1938, Davidson
S25 (F).

ECUADOR, PERU

TILLANDSIA LATIFOLIA Meyen, Reise 2:45. 1835; Obs. Bot. Itinera
in Nov. Act. Nat. Cur. 19, Suppl. 1:437. 184
Tillandsia kunthiana Gaud. Atl. Vos. Banite pl. 53. 1842; 1. 3.

mith, Phytologia 7:108. 1959.
In 1959, I ado pted Tillandsia kunthiana Gaud. as the correct
ame for this species because I failed to discover the earliest
pantieaca of T. latifolia Meyen. I now return the species to
ee ae pote along with var. major Mez and var. divaricata
enth.) Mez
e Ven ties are typical of the arid coastal region,

but var, divaricata is distinctly more northern than the others.

142 PHYTOLOGIA Vol. 13, no. 2
PERU

PITCAIRNIA PUNGENS H.B.K. var. FLAVA L. eg Smith, var. nov.

A var. PUNGENS petalis flavis differ

PERU: Cuzco: Prov. Ur ub amba: Chiff aoe Inca road 1 km east
of Machu Picchu, Valley of Rfo Urubamba, ca. 80 km WNW of Cuzco,
alt. 2550-2800 m, 3 January 1963, H. H. & C. M. Iltis & D. & V.
Ugent 1128 (US, type; WIS, iuntyoes.

PUYA aera L. B. Smith, sp. nov.
. angulonis L. B. Smith, cui affinis, bracteis stnlsie of
quam SMa longioribus sed reflexis, sepalis apiculat
differt.
PLANT flowering about 1m high. LEAVES ca. 4 dm long; sheaths
suborbicular, 55 mm wide, entire, lepidote beneath toward apex,

u

covered on both sides with appressed cinereous scales but more or

less glabrous above with age and thus distinctly vemee. "50 lax-
SCAPE

ly serrate with brown mostly recurved 3 mm long spines.

erect, 15 mm in diameter, very finely Hoslipap APPT becom
glabrous; scap racts erect, densely imbricate, the r folia-
ceous, th per broadly ovate with linear blades, nearly gla-
brous, pectinate-serrate. N RESCENCE simple, strobilate, CY
lindric, 2 dm g, 6 cm in diameter; — gist ovate, acumi-
nate, pungent, 6 cm long, exceeding the flowers but recurved at

gen
anthesis, pectinate-serrate, coriaceous but Seeetkc eS? nerved,
nearly glabrous, sublustrous; pedicels obconic, 5 mm long, dense-
ly white-stellate. SEPALS lance-elliptic, broadly acute and api-
culate, 21 mm long, coriaceous, coarsely and ao nerved,
slightly white-ste ellate at base; petals about 4e ie en
green (! Hutchison), barely era a the stamens; str
twisted after anthesis. Pl. , fig. 8: Floral bract si Flower

he
PERU: La Libertad: Prov. Huamachuco: 40 km above Pullac cross”
ing over Rio Chuagon Valley between Rfo Mardtion and Buldibuyo,
alt. 3640 m, 7 Aug. 1964, PB. C. Hutchison, J. K- Wright & R- M-

Straw 6170 (US, type; UC isot type).

PUYA GLANDULOSA L. B. Smith, sp-

AP. meziana Wittm. atque S eatubas aliis generis inflorescen—
tiae glandulis sessilibus differt.

PLANT known only from fragments, probably flowering ove r2m
high. LEAVES over 12 dm long; blades very na rrowly tri ngs
over 4 cm wide, obscurely lepidote between the nerves beneath,
glabrous above, axly serrate wit broad castaneous antrorse 12
mm long spines. SCAPE gros INFLORE branching
CG euccneuay’ primary bra unknown; branches cylindric, rather
dense but some of the aaa visible at anthesis, except for

to
SEPALS linear-lanceolate, obtuse, 37 mm long, ecarinate, thin;

1966 Smith, Notes on Bromeliaceae 143

petals ca. 7 cm long, exceeding the stamens, green (! rh a
naked, twisted together after anthesis. Pl. VE, fig. 40: Flora

x.)
PERU: La Libertad: 11 km above Samne, alt. 2290 m 5 Aug
964, B. C. Hutchison, J. K. Wright & R. M. Straw 6123 (us. “Sypes
UC isotype).

eee ee ee L. B. Smith, sp. nov
auhii in systema mea proxima sad scapi bracteis mney
Libus ced nigris nec fragilis, floribus minoribus differ
own anth

mn lon
spines. SCAPE straight, 12 mm in diameter, densely and Rtaeae:
white-stellate, soon glabrous; scape-bracts erect, the lowest un-
known, the others linear-lanceolate, the median serr lan he
upper entire. INFLORESCENCE simple, cylindric, 4-5 du long, ra-
ther lax at least toward base, except for the petals covered with

lowest owers, elsewhere distinctly shorter, coriaceous; pedi
cels slenderly obconic, 12 mm long. SEPALS linear-lanceolate,
broadly acute and apiculate, 30 mm long, ecarinate, coriaceous;
petals 5 cm long, about equaling the stamens, naked, metallic
blue (! Hutchison). Pl. VI, fig. 12: Scape-bract x 1/2; fig. 13:
Floral bract and flower = 1/25 fig. 14: Sepal x

PERU: Ancash: 1/2 km above Chancos on chi an. to Vicos, alt.
2875 m, 14 March 1964, P. af Hutchison & J. K. Wright 4380 (US,
type; UC isotype).

PUYA ILTISIANA L. B. Smith, sp. n

In systema mea P. gummifera Mer % Sodiro atque P. asplundii L.
B. Smith affinis, a priore bracteis florigeris homomor phis, flo-
ribus es a pets ppd indumento albido, a ambobus se-
palis s haud obovatis differ

LANT obi only from ecco ‘but probably less than 1 m

ry bra the upper sc
Spikes but their blades sometimes sli ghtly eg PR ey! spikes
i rag sessile, densely few-flowered, the er o 4 cm
1 the terminal 8 cm; floral bracts broadly Site : Sina:
ea exceeded by the sepals; flowers sessile. SEPALS narrowly

bh PHYTOLOGIA Vol. 13, no. 2

lanceolate, acute, 14 mm long. CAPSULES subglobose, 12 mm long.
Pl. VI, fig. 15: Spike x 1/2; fig. 16: Sepal x l.

PERU: Apurimac: Prov. Aymaraes: Talus slope with bromeliads,
valley of Rfo Challhuanca, ca. 18 km north of Challhuanca on road
to Abancay at km 104 from Abancay, alt. 2700 m, 17 Dec. 1962, H.
H. & C. Iltis & D. & V. Ugent 613 (US, type: WIS, isotype).

PUYA MACROPODA L. B. Smith, sp- nov.

P. sanctae-martae L. B. Smith in systema mea proxima sed pedi-
cellis fructiferis plus minusve applanatis multo majoribus, sepa-
lis valde angustioribus acuminatisque differt.

PLANT flowering ca. 2 m high. LEAVES Unknown but presumably
spinose-serrate judging by the scape-bracts. SCAPE straight,
slightly compressed, to 3 cm wide, finely white-stellate

S, s a s, nerv
when dry; petals 5 cm long, yellow-green (! Hutchison) , twisted
together after anthesis. CAPSULE very broadly ellipsoid, 25 mm
long. Pl. VI, fig. 17: Floral bract and old flower x 1/2; fig-
18: Fruiting sepal x l.

Lg Amazonas: Prov. Chachapoyas: Dry slopes overlooking the
Rfo Utcubamba, 11 km west of Chachapoyas on the road to Caclic,
alt. 1930 m, 26 June 1964, P. C. Hutchison & J. K. Wright 5855,
(US, type; UC, USM, isotypes).

PUYA PONDEROSA L. B. Smith, sp. nov.
AP. herzogii Wittm., cui affinis, bracteis primariis florige-
risque ovatis ex sicco stramineis vel pallide brunneis, sepalis

rt.
ANT flowering somewhat over 1m high. LEAVES 6 dm or long-

t ery short internodes, the lowest subfo fe) xr
own. I CENCE densely cylindric, 40 cm lon

diameter, bipinnate but appearing simple, at first whitish-tomen—

tose except the petals; primary bracts ova e, about ic

s ovate, acute, about equaling the sepals;
thin; pedicels stoutly obconic, 10 mm long. SEPALS lanceolate,

1966 Smith, Notes on Bromeliaceae 145

acute, 25 mm long, thin-coriaceous, drying stramineous, their
tomentum persistent; petals ca. cm long, exceeding the stamens.
Pl. VI, fig. 19: Floral bract and flower x 1/2; fig. 20: Sepal

1

x
ERU: Ayacucho: Prov. Lucanas: Rocky slopes in arid basaltic
Tola heath (Lepidophyl1lum quadrangulare), descent to Lucanas
(Puquio) from Paso de Kondor sencha, on Nazca-Puquio Road,
Galera", alt. 3800 m, 13 Dec. 1962, H. H. & C. Iltis & D.

Ugent 445 (US, type; WIS, isotype).

&

"Pampa
v.

PUYA WRIGHTII L. B. Smith, sp. nov.
P. rauhii L. B. Smith in Systema mea proxima sed foliorum

T flowering over 2 m high. LEAVES to 13 dm long; sheaths
suborbicular, glabrous toward base; blades very narrowly triangu-

Ose, 2 cm long; seeds reticulate, the wing circumferential, cu-

PERU: Cajamarca: Prov. Jaen: Cliff faces and steep rocky
Slopes, a little north of Pucard, alt. 1000 m, 12 Jan. 1964, P.
&. Hutchison & J. K. Wright 3560 (US, type; UC, isotype).

TILLANDSTIA HUTCHISONII L. B. Smith, Sp. nov.

AT. marantoidea Rusby, c erisimiliter affinis, omnibus
PE rg valde majoribus, inflorescentia ramosissima, spicarum
Thachi geniculata differt.

PLANT flowering 1.5-2 m high (! Hutchison & Wright). oe
over 8 dm long; sheaths broadly ovate, densely punctulate-lepi-
dote, dark Castaneous beneath; blades ligulate, broadly subacute

w
beneath, glabrous above. SCAP own. CENCE amply
3-4-pinnate, glabrous, red except the petals; primary
broadly Ovate, acumina 10-11 cm long, exceeding the bracteate

1h6 PHYITOLOGIA Vol. 13, no. 2

flattened, flexuous; secondary bracts scarcely larger than the
floral bracts, excee ag the bracteate sterile bases of the axil-

i:

PERU: Amazonas: Prov. Bongara: On road to Rioja, 5 km north of
the siorth end of Laguna Pomacocha, alt. 2000 m, 8 Oct. 1964, P.
C. Hutchison & J. K. Wright 6801 (US, type: UC, isotype).

pr sagen PIURENSIS L. B. Smith, sp

T. rubella Baker atque T Serban affinis, a priore foliorum
laminis haud ligulatis sed | ‘iccte triangularibus, bracteis 7
rigeris laevibus, a posteriore floribus multo major ribus, a amb

ng urely p
broadly si Paine aeaeehis distinct, ca. 10 cm long, aia casta-
neous above, concolorous with the blades beneath; blades narrowly
triangular, long-attenuate, 5 cm wide, pale green striated or
marmorated dark purple-black (! Hutchison). SCAPE slightly
curved but probably ascending, 6 mm in diameter, glabrous; scape~
bracts suberect, subfoliaceous, their bases exceeding the inter-
nodes but not altogether covering them. INFLORESCENCE bipinnate,

planate, 7-8 cm long, 2 cm wide (fruting), 8-9-flowered; floral
bracts imbricate and concealing the rhachis, elliptic, acute
carinate and en —_ toward apex, 25 mm long, eq ualing
the sepals, coriaceous, » glabrous; flowers subsessile.
SEPALS Lidasecohbenbeer eee eesti acute, 24 mm long, thin,
nerved, obscurely lepidote, the anterior free, the posterior
connate for 6 m, ee: Pl. VI, fig. 25: Spike x 1/2; fig-
26: Posterior sepa

PERU: Piura: bet aasakebaabal Cliff faces, Rfo Huancabamba
gorge 3-4 km below Huancabamba on the road to Sondor, alt. 2000
m, 16 Sept. 1964, P. C. Hutchison & J. K. Wright 6616 (UC, type)-

rag 1-2 } sae et PLATYPHYLLA Mez, Fedde Rep. Nov. Spec. 3:
ot 6; L. B. Smith in Macbride Fl. Peru, Bot.Se ries
aH bee Nat. Hist. 13:558. 1936, emend. 1966.
rorteesenien ee ample tripinnatim sigan Se ramis spicas ad 8
eee petalis nudis; staminibus exsertis
Cajamarca: Prov. Hualgayoc: Between Sen Pablo and San

1966 Smith, Notes on Bromeliaceae 147

Miguel, alt. 1700 m, 2 May 1904, Weberbauer 3888 (B, type;

phot. 11521). Prov. Jaen: Km 113 betwee n Acapulco and pacers on

ae Rfo Huancabamba, alt. 1000 m, 10 Jan. 1964, P. C. Hutchison &
- Wright 3516 (UC, US).

pv “TILLaNDSTA “ace -CATOPSIS) TETRANTHA R. & P. var. RAMOSIOR L.
Sm ar. nov.
ar. ee a et varietatibus alteris inflorescentia tri-

nnata differt. A T. bakeri L. B. Smith bracteis primariis

RU: Amazonas: Prov. Chachapoyas: Cerros Calla Calla, 18 km
above Leimebamba, km 410 on vbr to Balsas, alt. 3100 m, 14 June

— ES Butchtson & J. K. Wright 5669 (US, type; UC, bel
NOTE: In its small oh ti bracts and large floral bract
rilandsia tetrantha var. ramosior is closer to var. seit ace
than the typical variety
BOLIVIA

PUYA UGENTIANA L. B. Smith,

n systema mea P. canbe Sera atte, sed foliorum laminis
supra dense lepidotis, inflorescentiae indumento persistente,
floribus longe pedicellatis differt.

PLANT flowering over 2 m high. LEAVES over 1 m long; blades

ry narrowly triangular, r ; a » obscurely lepidote
between the nerves b h, covered w appressed cinereous
scales above, laxly serrate with castaneous antrorse spines 3 mm
long. CAPE straight, cm t b soo brous ;

bracts lan olate, acute, n long and about equaling the
sepals; pedicels slenderly rd Re: 15 mm long. SEPALS lan-
ceolate, harrowly obtuse, 30 mm long; petals linear, stellate at
least toward base. CAPSULES subglobose, 2 cm long. Vit.
fig. 1: Floral bract and flower x 1/2: fig. 2: Sepal x

BOLIVIA: Chuquisaca: aia Ago se Steep rocky slope
Puyas and pentechia, 29 km ENE of Sucre on the Sucre-Aiguile ae
(15 km ENE of Sucre by ma ates ce},alt. ca. 1950 m, 13 Apr

1963, D. Ugent 4998 (US, cise WIS, isotype
BRAZIL

AECHMEA CALATHEOIDES L. B. Smith, sp. nov.
A Ae. basi-lateralis (Lem.) L. B. Smith, cui valde affinis,
sepalorum partibus liberis serrulatis, pedicellis longis differt.
LANT caulescent; rhizome 15 mm in diameter. LEAVES f

148 PHYTOLOGIA Vol. 13, no. 2

rosulat 7 dm long, densely white-lepidote beneath, glabrous

liptic, sparsely serrulate. INFLORESCENCES simple, strobilate,
1 ;

lepidote; pedicels slender, 5 mm long. SEPALS 22 mm long, con-
nate for 15 mm, the free apices broadly spatulate, apiculate,
serrulate. FRUIT aera 14 mm in diameter. Pl. VII, fig. 3:
Inflorescence x 22; fie. 4s" Fruit: x: 1;

BRAZIL: Ama apa: tn shade, forest swamp, wet ec lay soil, immedi-
mod east of Colonia Agricola de Oiapoque, 3” 43" N, 51° 55' W

t 4 km north of mouth of Crici’ River, 14 Aug. 1960, H. S.

el W. A. “tales & L. Y. Th. Westra 47527 (NY, type).

AECHMEA shes: (Regel) - B. Smith, comb. nov.

AECHMEA FARINOSA var A

Lamprecoccus iv4npeus Rocce Ind. Sem. Hort. Petrop. 1868:79.
869.

aociiadll farinosa hort. Makoy ex Regel,loc. cit., nomen in synon.

Aechmea c cong teerats var. farinosa (Rege1) Baker, Handb. Bromel.
als 89; L. B. Smith, Bromel. Brazil, Smithsonian Misc.
made 126:204. 1955

Aechmea glomerata var. ¥ortnoes (Regel) Mez in Mart. Fl. Bras. 3,
pt 892.

$7 32316.
onym for Aechmea farinosa was antedated by that for Ae-
glomerata Mez, but the later name was invalid because of Hooker's
earlier use of it for another species. Then ig substitute name,
Ae. conglomerata, was later than the basonym of Ae. farinosa, 80

that it becomes nec essary to make new combinations for all three
varieties of the species

oe pnp ing ad? L. B. Smith var. CONGLOMERATA (Baker) L-
Smi com!
Lanprocaecus ga ee Beats Bromel. 105. 1857.
Aechmea ees erata gest ex Baker, Handb. Bromel. 52. 1889; L-
=e mith, Bromel. Brazil, Smithsonian Misc. Coll. 126: 1204.
an

i 4
te a omerata are gay in Mart. Fl. Bras. 3, pt...3:315;
» hot Hook.

eine fe eg ay apsits L. B. Smith var. DISCOLOR (Beer ex
er) L. B, Smt comb. nov

Lanprocaccus ee sad ig var. dibeuior Beer, Bromel. 105. 1857,

Acchnea ‘glonerats discolor hort. ex Beer, Bromel. 105. 1857,

hechnes congloserata = discolor Beer ex Baker, Handb. area
mith, Bromel. Brazil, Smithsonian Mis

1966 Smith, Notes on Bromeliaceae 149
Coll. 126:205. 1955.

BILLBERGIA BRACHYSIPHON L. B. Smith var. PARAENSIS L. B. Smith,
var. nov
- brachysiphon sepalis inaequalibus, apice latioribus
sistent. Pl. VII, fig. 5: Sepals: x 1.
L: Pard: Epiphytic, braver 25 May 1958, P. B. Cavalcante
4g3- tee 23049, type).

BROMELIA CACHIMBENSIS L. B. Smit p- nov.

AB alansae Mez, cui affinis, ramis infimis basi longe ste-
rilibus, pedicellis gracilibus longioribusque, sepalis majoribus
diffe

pues known only from So Ser irtien 3 shoot, over 4 dm high.

SCAPE ca. 1 cm in diameter t apex, covered with tomentellous
pale brownish indument. INFLORESCENCE subcylindric, dense, 35 cm
long, 12 in diameter, the axes and ovaries densely tomentel-

slender, in fruit. SEPALS oblong, cucullate n
duplicate, 26 mm mg, free, oS aa soon glabrous; petals 33
long, elliptic; act aee connate at base for 10 mm; ovary subcy-
lindric, 18 mm long, e pigynous bite narrowly funnelform, 5 mm

high. FRUIT biti ellipsoid, 25 mm long. Pl. VII, fig. 6:
a

RAZIL: Para: Cachimbo, 9% 20' Ss, 54° 53' W, 9 Nov. 1955, W.
prise in 252 (IAN 111630, type).

BROMELIA OLIVEIRAE L. B. Smith nov.

Habitu eitieausr & agavif Siiae ® Brongn. ex Houllet valde simu-
lans sed pedicellis fracciferis elongatis, sepalis fere duplo
majoribus differt.

own only from fragments. LEAF (only one) with linear

bel mm, ered beneath with fine appressed wh

Scales, = above, laxly serrate with distally antrorse
basally retrorse black curv long spines. SC stout, to
1 ng, densely lanate with fine pale brown scales; scape-
bracts densely imbricate, elliptic, thin, serrulate, mi-
gh and inconspicuously lepidote, nearly white when d

FLORE subglobose, ca. ter, dense, anes:

cels slender, to 12 mm mm long. SEPALS like the floral bracts but
Pah pe 25-30 mm mm long, glabrous; petals imperfectly known, line-
rT, glabrous; Stamens with filaments connate for 6 mm; ovary with
narrowly funnelform epigynous tube 5 mm long. FRUIT ellipsoid,
at least 3 cm long (young). Plus Villy figs 2:.Flower x-.1

150 PHYTOLOGIA Vol. 13, no. 2

BRAZIL: Parad: Upland rain forest, km 147 of the Belém - Brasf-
lia Highway, 17 Feb. 1960, E. Oliveira 518 (IAN 105255, type).

DYCKIA IRMGARDIAE L. B. Smith, sp. no
abitu D. reitzii L. B. Smith a bat similis sed sepalis poste-
rioribus Eiceec amet, filamentis ultra tubum petaleo-stamine-
r

PLANT ne to 1m high. LEAVES very numerous ina

with black antrorse curved 2 mm long spines. SCAPE erect, 5 mm

in diameter, densely ferruginous-lanate; scape-bracts densely im-

bricate, narrowly triangular from a broadly ovate base, ferrugi-

nous-lanate. INFLORESCENCE simple, densely many-flowered, ca. 3

dm long, densely ferruginous-lanate; floral bracts like the upper

scape-bracts, mostly exceeding the flowers; pedicels stout, com
n

the petals; stigmas subsessile. Pl. VIL,fig. 8: Floral bract and
flower x 1; fig. 9: Sepal x
BRAZIL: Rio Grande ii Sul: —— by top of aoa ee 20
Nov. 1964, L. B. & R. C. Smith, L. Schultz & Oyara Ss,
_—
e species is named in honor of our wee Dra.
ie Btn of the Universidade do Rio Grande do Sul.

DYCKIA MARITIMA Baker, Handb. Bromel. 136. 1889.
Dyckia tomentosa Mez in DC. Mon. Phan. 9:515. 1896.

IL: Rio aes do Sul: Tweedie s. n. (K, t 3; GH phot
Sellow Bromel 73 (P, type of Dyckia tomentosa ens * ol i photo 3003)
Térres, Reitz Reitz 4427 (HBR); Smith & Reitz 5824 (Ry > 20S

TE: Dyckia ma maritima Baker is abundant and pan rt on ae
rocky headlands at rend and both Tweedie and Sellow must hav
passed very close this place. It is more than likely that
Térres is the type hacataee for both species.

DYCKIA PARAENSIS L. B. Smith, sp.
Habitu D. duckei L. B. Smith jalde: similis sed bracteis flori-
geris quam . pedicellis robustis longioribus, cs mtis ultra

r
entire. INFLORESCENCE simple, very lax, r 2 dm long, gla-
brous; floral bracts like the upper scapeieekees to 5 mm long;

1966 Smith, Notes on Bromeliaceae 151
pedicels spreading, rather stout, 3 mm long. SEPALS elliptic-
oblong, broadly rounded, 6 mm long, ecarinate, rather thin;
petals 10 mm long, about equaling the stamens, the blades very
broadly rounded, carinate; stamens with filaments short-connate
above the junction with the petals; stigmas subsessile. Pl. VII,
fig. 10: Floral bract and flower x 1; fig. 11: Sepal x 1: sig.
eet Pietil & 1.

: Pard: On sandstone ledges, Santa Cruz dos Martfrios,
campos of Andorinhas, region of Aragudia, alt. 600 m, 15 June
1953, R. L. Frées 30030 (IAN 79853, type).

DYCKIA PUMILA L. B, Smith, sp. nov.

D. duckei L. B. Smith, cui affinis, pedicellis brevibus
crasse obconicis differt,

PLANT nearly stemless, flowering 18 cm high
Spreading rosette, 13 cm

LEAVES many in a

ee
linear-triangular, pungent, 7 mm wide, t
Spreading slender brown spines 2 mm long, covered with
Ee

u

>
shorter than the internodes, ovate, attenuate, obscurely serru-
d, 2-5 cm long, very

13 mm long, dark castaneous. Pl. VII,

RAZ talus, at base of cliff face, ca.
50 km south of Caiaponia on road to Jataf, Serra de Caiapd, 17
12" S, 51° 47' W, alt. 800-1000 m, 25 Oct. 1964, H. S. Irwin & T.
R. Soderstrom 7365 (US, type;NY).

DYCKIA SILVAE L. B. Smith, sp. nov.
BD. tuberosa (Vell.) Beer in systema mea affinis sed petalorum
laminis magnis valde patentibus, stylo nullo differt.

Frown, nearly glabrous; blades linear-
wide, thick, laxly serrate with spreading slender brown spines 3

ly ovate, attenuate, slightly
RESCENCE s le, subdensely many-flower very
Sparsely white-lepidote; floral bracts like the highest scape-
bracts, Serrulate, the lowest about equaling the sepals; pedicels
Subcylindric, thick, 3 mm long, strongly constricted at base and

152 PRET OPC GT & Vol. 13, no. 2

appearing F ieee cel SEPALS ovate, obtuse with a wide thin mar-
7

RAZIL: Pard: On rocks in campo, region of Garotire, Mun.
Altamira, 8 Aug. 1962, N. T. Silva 784 (IAN 114554, type).

ENCHOLIRIUM GLAZIOVII Mez in Mart. Fl. Bras. 3, pt. 3:505. 1894,
emend. L. B. Smith i 6.
Eneholawn subsecundum sensu L. B. Smith, Smithsonian Misc.
Mile b26543. 1955; dined B. Smith 7036, non Mez 1896.

be ad 9 dm longis; vaginis wenkiovatiee. ad 9 cm latis,
atro-castaneis, apice dense brunneo-lepidotis, alibi glabris;
laminis anguste triangularibus, 5 cm latis, basi spinis latis

late ellipticis, apice attenuatis, ae ue serrulatis. sPl. VEIL,
fiz. 16: plas bract and flower x

BRAZIL as Gerais: Without Aieeude data, Glaziou 19918 pe
type, F photo 11456). Chapeu do Sol, Serra do Cipo 19° 40
43 57' W, alt. 1000 m, 29 April 1952, L. B. Smith & Mus. R. 7036

S).
NOTE: It could well be that L. B. Smith 7036 is topotypic.

Glaziou collected in company with Schwacke and on occasion also
appropriated the collections of others as his own (S er, Bot
Jahrb. 76:153. 1954), and Schwacke is have collected on

the Serra do Cipd. L. B. Smith 7036 agrees closely with such
parts as are represented in the rather incomplete type.

ENCHOLIRIUM MAGALHAESII L. B. Smith, sp. nov.
A E. subsecundo (Baker) Mez, cui affinis, inflorescentia flo-
ribusque valde minoribus differt.

tric, suboblong, subtruncate, 7 mm long; gutels narrow, barel
exceeding the sepals; stamens exserted; ovary narrowly pyramidal,
long. CAPSULE ovoid, dark castaneous; var ee ee ular-
falcate. PI. VII, fig. 17: Floral bract and flower x 1.
BRAZIL: Minas Gerais: Campo, slopes of Serra de ‘headin eee
July 1960, M. Magalh@es 18056 (IAN 107941, type).

eee eseseeeesesesaeseeeseeeeseeeeeneeeeseeesesessesees eee eee oo 8 eS

1966 Smith, Notes on Bromeliaceae 153

GRAVISIA RODRIGUESIANA L. B. Smith, sp. nov.
omnibus speciebus generis adhuc cognitis inflorescentia e
spicis elongatis dense digitata differt.
PLANT flowering over 6 dm high. LEAVES 14 dm long; sheaths
broadly elliptic, distinct, 20 cm long, dark castaneous, minutely

appressed-lepidote; blades ery narrowly triangular, e,
coriaceous, soon glabrous, sparsely serrate with slender spread—-
ing dar mm lo spines CAPE ight, stout, appressed-

lepidote; scape-bracts strict, large and very densely imbricate,
drying dark castaneous, the lower subfoliaceous, the upper with

gues), drying to dark castaneous, obtusely carinate, punctulate-
lepidote; spikes subcylindric, 10 cm long, 3 cm e nsely

broadly elliptic, rounded and mucronate, 30 mm long, equaling the
e ari (! Rodri-
gues), drying stramineous; flowers subsessile. SEPALS asymme-
ric, 19 mm long, short-connate; epigynous tube infundibuliforn,

mm high; petals and stamens unknown. P1. VII, fig. 18: Spike x
1/2; fig. 19: Sepal x 1.

RAZIL: Amazonas: Mata de baixio, Bosque de Palmeiras, Reserva

Florestal Ducke, Manaus, W. Rodrigues 5399 (US, type; INPA 14068,
isotype), ,

ORTHOPHYTUM LEPROSUM (Mez) Mez in DC. Mon. Phan. 9:117. 1896; L.
B. Smith, Brom. Brazil, Smithsonian Misc. Coll. 126:80.
1955.
*rantleia leprosa Mez in Mart. Fl. Bras. 3, pt. 3:259, pl. 58,
ig. 2. 1891.
IL: Minas Gerais: Cachoeira do Inferno, Rio Jequetinhonha,

ery
caatinga on rock outcrop, between Itaobim and Joaima, ca. 1
5, 41° 15' W, Nov, 1959, M. Magalh@es 17598 (IAN 106716).
NOTE: The Magalh&es collection must be from very near the type
locality and, except for a loss of indument due to wear, it ap-
Pears to be a close match. The type locality has been given er-

roneously as in Goids when actually is in northeastern Minas
Gerais near the remainder of the genus which is limited to north-
fastern Brazil. er supposedly western locality for the

been a label mixture in 0. foliosum and that his 1079 of Mato
Grosso should have been 881 of Espfrito Santo.

PITCAIRNIA IRWINIANA L. B. Smith, sp. nov.

Petalis ignotis, P. ensifolia Mez vel P. karwinskyana Beer in
Systema mea affinis sed sepalis omnibus late alato-carinatis
differt,

T nearly stemless, flowering about 3 dm high. LEAVES
many in a fasciculate rosette, the larger ones over 2 dm long;
sheaths broadly ovate, 2 cm long, imperfectly known because of

15h PHYTOLOGIA Vol. 13, noe 2

fire damage; blades dimorphic, some reduced to subulate dark
spinose-serrate spines, others foliaceous, linear, subulate-
attenuate, scarcely if at all narrowed at base, 10 mm wide

covered white appressed scales but becoming glabrous above,

deciduous along a straight transverse line, entire excep

short persistent spinose-serrate base. CAPE erect, 5 inudi-
ter, te-flocculose; scape-bracts all exceeding the inter-

nodes, the lower foliaceous with deciduous blades, the upper much

smaller and ovate. INFLORESCENCE simple, dens -6 ;

floral bracts like the upper scape-bracts, 2 cm long, exceeding
the slender 13 mm long pedicels. SEPALS oblong-elliptic, acute,
15 mm long, all broadly alate-carinate; petals and stamens un-
known; ovary about 2/3 superior. Pl. VII, fig. 20: Old flower
(type) x 1; fig 21: Young inflorescence Coatatyon) x13. fig. (22s

cis

IL: Goias: Rooted among boulders, mesa summit, near eS-
carpment, ca. 50 km south of Caiaponia on road to Sates, serra do
Caiaps, 179 12' s, 51° 47' W, alt. 800-1000 m, 25 Oct. 1964,

S. Irwin & T. R. Soderstrom 7347 (US, type; NY). Same, base -
escarpment, 26 Oct. 1964, Irwin & Soderstrom 7370 (NY, US).

PITCAIRNIA LIMAE L. B. Smith, Phytologia 7:254, pl. 1, figs. 9;
10. 1960, emend. 1966.
Florifera ad 75 cm alta; foliis multis, homomorphis; vaginis
late ovatis, 3 cm longis, atro-castaneis; laminis persistentibus,
8

latis, subtus densissime cinereo-lepidotis, supra mox glabris,
basi laxe spinoso-serrulatis, alibi integris; petalis nudis vel
basi ligula auctis.

BRAZIL: Cearé: Serra de Maranguape, cume da Rajada, 14 Sept.
1908, A. Ducke s. n. (MG 1661, topotype).

Plate I

Fig. 1: Vriesea rauhii. 2-4: V. olmosana. 5: V. didistichoides-
6: V. fragrans. 7: V. cylindrica. 8: VY. harmsiana.
9: V. tillandsioides. 10: V. lubbersii. 11: V. drepanocarpa-

19: V. altodaserrae. 20, 21: V. saundersii. 22: V. procera.
23: V. alta. 24: V. geniculata. 25: V. gigantea.

155

1966

Smith, Notes on Bromeliaceae

Plate I

(For explanation see preceding page)

156 PHYTOLOGIA Vol. 13, no. 2

Plate II

: Vriesea regina. 2: V. brasiliana. 3: V. erythrodactylon-
4: V. heliconioides. 5: V. barilletii. 6: V. incurvata. 7: V-

gladioliflora. 15: V. ovandensis.

157

Smith, Notes on Bromeliaceae

1966

Plate III

piculata. 3: V. amethystina. 4: V.
8, 9: V. fenestralis.

5: V. recurvata. 6: V. morreniana.

7: V. ensiformis.

Fig. 1, 2: Vriesea a
Psittacina.

158 PHYTOLOGIA Vol. 13, no, 2

Plate IV

Fig. 1: Vriesea jonghei. 2, 3: V. fosteriana. 4, 5: V-
platynema. 6: V. bituminosa. , 8: V. pardalina
9, 10: V. guttata. Tl: Ve retroflexa. 12: -V. eimplex-
13, 14: V. scalaris. 15-17: V. tuerckheimii. 18: V.
philippocoburgii. 19: V. ringens. -

Fig.
imperialis, 6,
Sanguinolenta.
aE oy;

Tacinae,

Smith, Notes on Bromeliaceae

Plate V

: Bo 345 °5t-V
1: Vriesea itatiaiae. 2, 3: V. confus ;
7

: V. hieroglyphica. 8: V. werckleana

10

11: V. vanhyningii. 12: V. cee ee
lei. : V. broadwayi. > ae
tet, se Ce 20: V. capituligera.

159

160 PHYTOLOGIA Vol. 13, no. 2

Plate VI

|
|

Fig. 1, 2: Vriesea malzinei. 3, 4: V. flammea. 5-%. Tillandsia
oerstediana. 8, 9: Puya coriacea. 10, 11: P. glandulosa.
12-14: P, hutchisonii. 15, 16: P. iltisiana. 17, 18: P
macropoda. 19, 20: P. ponderosa. 21, 22: P. wrightii.

23, 24: Tillandsia hutchisonii. 25, 26: T. piurensis.

Shien

1966 Smith, Notes on Bromeliaceae 161

Plate VII

-
AV)

i
i
i

a: 1, 2: Puya ugentiana. 3, 4: Aechmea calatheoides. 5:
bergia brachysiphon var. paraensis. 6: Bromelia cachimbensis.
* B. oliveirae. 8, 9: Dyckia irmgardiae. 10-12: D. para¥nsis.
13: D. pumila. 14, 15: D. silvae. 16: Encholirium glaziovii.
Av: E. Magalhaesii. 18, 19: Gravisia rodriguesiana.
20-22: Pitcairnia irwiniana.

MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA,. X&
Harold N. Moldenke

PPIA UMBELLATA Cav.
Additional bibliography: Moldenke, Phytologia 13: 7 & 32——35.

“< the other hand, the Hinton 129h5 and S. S. White 5271, dis-
tributed as L. umbellata, are actually L. callicarpaefolia lia H.B.K.;
P, C, Standley 20803 is L. cardiostegia Benth.; Heyde & Lux 4387
is Le controversa sa Moldenke; Holway 307 is L. costaricensis Mol-—
denke; Edw. Palmer 79 (april to Nov. 1896] is the type collection
of L. curtisiana Moldenke; Paray 3123 is L. durangensis Moldenke;
Burd a 67 is L. gentryi Standl.; Balls . B.5489, N. L. H. Krauss 858,
and Rzedowski 6906 are L. hypoleia Briq.; H. S. Gen 2 and
Mexia 1636 are bins.” jaliscana Moldenke; H. Pittier 5387 and R. S.
Williams ms 151 are L. oxyphyllaria (Donn, . Sm.) Standl.; Arséne 5293,
Breedlove 1572, D. S. Correll 11,396, Galeotti 787, He S. Gentry

2037, J. Gonzalez Ortega ga $707, Binto Hinton 11717, 12949, & 13614, Mexia

ee 1/2 & 135, Edw. Palmer 479 [Oct. 189 — March 1895], | 2088

& Rose . 14798, Je 1 de Rzedowsli 15630 & 19280, and Cc. L

Saith are L. pringlei Briq.; Ghiesbreght 63h, Heyde & & Lux 1,389,

Ttirckheim (in trigosa

Turez.; DeLeon 1618 & & 1629, H. S. Ge entry 3393 | and Edw. Palmer

1969 are L. tepicana Moldenke (the latter e type collection);
Gaumer & sons 23627 bis is L. Laces die a Be
In Harvard University herbarium the foll ime:

me: MEXICO: Guerrero: E. W. Nelson 2285 (G), Se (G).- Hidalgo:
H. E. Moore 2000 (G). Michoacdn: E. W. Nelson 6951 (G). Morelos:
& undetermined: Berlandier 1028 [Cor
a da Guchilapa] (G); Halbi 53 [Balsas] (G). GUATEMALA:
hmetenango: Skutch 1080 (A). EL SALVADOR: San Salvador: Cal-
er 0s P. C. Standley 19395 (G). Sonsonate: P. C. standley

In all, AP eaeeten eons bs ebsites re oe

Citations MEXICO: Colima: McVaugh & Koelz 1069 (Mi). Federal
District: Bourgeau 968 (Br, S); J. N. Couch M.126 (Hi—11509)5 x
onnet 157 (N); H. N. Moldenke 198 19853 (B, F, Mg, Mr, No); Moldenke =
eo ke 19853 (Bu, Bs, Es, Fy, Hk, Hw, Le, Lg, Lm, Mm, N, Ok, at,

Rs, S, Sm, Sm, Ss, Ss, 25 Paray 2188 Cp) Russel & Sourtron 18
priest ton be N). ‘Reareeto 2 L . Jalisco:

Cb). Ja
Vaugh 10070 (Mi), 117h6 (ui), Se ie (Mi). entre ee is Be
~ 362

1966 Moldenke, Monograph of Lippia 163

Good: 2178 (Ca—130059, En—6170, I, N, S); Hinton 393 (N, N),
2099 (N, N), | 2574 (K), 2918 (N, N), 7237 (K, N), 11910 (La),
13550 (La); M. E. Jones 7h7 (Po--68896); Matuda 18690 (N), 25831
(cb); Paray 1349 (Ip); | Pringle 11089 (Fs, of rtigne ¥. & 3
s, vt); Pas s Purpus 169i (Ca—139739, N, Po—64317). Michoacén:

séno 8.n. [Morelia, -12-1910] (B, yw, N); Hinton 11910 uate Tt
Mi, | N, Rf, Ur), 12791 (Mi, N, < nies 13550 (Au, It, Mi, N, Rf,
Ur), 15706 (N, N, Rf), 15707 (La); King & Soderstrom 5161 (ca);
E. W. Nels Nelson S56 (N); aot ens (Br, C, Ca—25123, Io. eo OB nied,
Me, Me, Me, xi, Me—15U0T, Ob ecemcd ed Pa, P1—2259h,

8, Vt). Morelos: Froders Hultén 308 (S); E. a oe
(W—~176023) 28 (iO) Pringle 771; (I, Vt), ne (La,
Vt), 11090 (Cm, It, N); Rowntree 300 (Fs); W. Trelease (Ur).
Nayarit: Edw ioe Sen. [Tepic, Ja Jan.—-Feb, 1892] (Cp as Be Oaxaca:
Galeotti 795j (N, V); C.. C.L. Smith 225 (Vi). Puebla: C. C. Deam
135 (@), Ba | [Ixtaccihautl, Jan. 5, 1899] (Mi). Sinaloa: H. S.

Cp). Vera Cruz: Balls 5489 (W—-1793856). State undetermined:
Haenke 1588 (N); Quarles van Ufford 105 (Ut); Sessé, Macifio Cas-
tillo, & Maldonado 2123 (Q), 2203 (Q), 2218 (@), 2219 (Q, Q

LIPPIA VELUTINA Schau, in A. » Prodr. 11: 576. 187.

mymy: Lippia mollis ee ae ex Moldenke, Suppl. List Invalid
Names 6, in syn. 19
¥ Bibli Lography: Schau, in A. DC., Prodr, 11: 576. 18)7 spree

Fl. Bras. 9: 226. 1851; Bocq., Adansonia 3 - 1863;

Jacks, in Hook. ws & J » Ind, Kew, 2: 96. 189k; hae Bull.
Bot, Franc e 58, os 33 50. 1911; st

valid Names 6, 1911; Moldenke, Known Geogr. > Verbenac.,
fed. 1], 38 & 96. 1b u25 | Moldenke, Alph. List Invalid Names 31.
1942; Mold enke, rasa List Cit. 2: 413 Ene). 2 ca os 693, 710,
~ 93¢ (1949), and h: 1300, 1949; Moldenke gr. Distrib.

Py ed. 2), 81 & 191. 19h9 selaeahe,. #1 P ss ot

es § raat
1950, me Résumé 9, 314, & 463. 1959; Troncoso
t WSS. 196 1965. z 1961; Moldenke, Phytologia 12: 23, 7, 152, "TT, 310,

Shrub, to 2 m, tall; branchlets subtetragonal, subpulverulent~
ar and appressed-tomentose; leaves decussate~opposite;
petioles 6—8 mm, long; leaf=blades ovate-oblong or lanceolate,

acute and not at all attenmate at the a very saga y apicu

late at the tip, te secattl: acu the base

and contracted into the petiole, penninerved, flat, piston an
» densely very short—tamentose with appressed —

cent the lower ones on the branchlets 5-—-7.5 cm. long

and 1.8—3 om. wide ng in size “ up=

= only 2.5 cm. long; secondaries ‘

’
Sed
ro above, prominent beneath, the areoles not truly bullate; in-
rescence subpulverulent-glandular and appressed-tomentose; pe-

16h PHYTOLOGIA " Vol. 13, nooa

duncles axillary, aggregate, flaccid, 812 m. eee longer than
the minaret a petioles; heads tetraquetrous, subglobose, about 6

drying
its tube ) m. long, slightly curvate, ventricose and pubescent

below the mouth, the limb pubescent on the back, its lobes rounded,

apex
invested by the persistent bat not diphyllous calyx, ellipsoid,

at Morro do Ernesto, near Cujab4, Matto Grosso, Brazil, and on
Po. a

Emam
Corallingo, Goifs, Brazil, deposited in the Vienna, Munich, and
DeCandolle herbaria. The ’ first~mentioned of these is also the
type collection of L. mollis and a specimen so labeled in the
Delessert Herbarium at the Conservatoire et Jardin Botaniques at

him as his photograph number 216
Schauer (187) places the species in his Section Goniostachy~
wa and says "Proxime affinis L. Salviaefoliae, sed distincta
statura validiora, foliis figura alienis, magis elongatis, utrin-
que acutis, haud rugosis, indumento brevissimo appres
laxo vestitis, ramealibus et subfloralibus majoribus." The flow
e

"yellowish", Material has been misidentified and distributed in
herbaria as L. origanoides H.B.K. On the other hand, the Glaziou
uae , distributed as L. velutina, is actually L. salvisefolia

ae 7 herbarium specimens, including type material of both
— involved, and 11 mounted ohstupeailie have been examined by

Ca eka: BRAZIL: Matto Grosso: Silva-Manso s.n. [Morro do
Ernesto; Herb. Martius 102k; Macbride photos 7861 & 2676) (Br--
» F--photo of cotype, Kr—photo of cotype, Er

Se ie
W—photo of cotype, W--photo ag cotype,
nas Gerais: ‘a P. Duarte 7580 [Herb. Brad. fad. Fre 0) s
uagalioe 29 THerb. Jard. Bot. Belo . 45152] (N), sues (N)-
ater Martius 1037 (M).

wy

1966 Moldenke, Monograph of Lippia 165

LIPPIA seit Cham., Linnaea 7: 232--23). 1832.

mymy: Lippia attemata Mart. » Flora 21, Beibl. 2: 59. 1838.
Aloysia attemata Walp., Repert. Bot. Syst. : 42. 1845. Lippia
hirsuta Si yornontoides (Cham.) Kuntze, Rev. Gen. Pl. 3 (2): 252.

1598. iippia caniflora Mart, ex Moldenke, Alph. List Invalid
Names aon 1: 13, in syn. 1947. Lippia a ireste & vernonioides

(Cham.) Kuntze ex Moldenke, Résumé Suppl. Pe in syn. 196.

Lippia hirsuta var, vernonioides Prsnsg ) ake ex Moidenke, Ré-

sumé Suppl. 12: 10, in syn. 1965. Lippia tarweta f, latifolia
sumé

hirsuta f., lo olia Kuntze ex Moldenke, Résumé ‘Suppl. 12:
J ae

2l, bl. 2: a 1838; Steud. Bot., e > a? Sh. 1840;
-» Syn. Pl. 3: 600. ey em Walp., Repert. Bot. Syst.
W2 & 54. 18h5; Benth., Pl. Dis” isles Schau. in A. DC,

e Li: 589. 1847; Schau. in sy Ble o 9: hh. 1851;
Hiern, Vidensk. M 4 - 1877—1878: 99. 1877; Jacks. in
Hook. f, & Jacks., Ind. Kew. 2: 95. 1893 S. Moore, ‘

- Bot., s 2, hs 1895; Kuntze, Rev. Gen
(2): 252. 1898; H. HL x Bot. Club 27:
Ze, P t. France ém. 3: 542. 1911; T. Her-
herb. Leiden 29: hh. 1916; Moldenke,

zog, Meded. Rijks ; $
ise 1903 moniacht Sweet «: List Invalid Names 1 & 5. “iat Mol~

7 tes
9), and h: 112& 1292. 19h9; Moldenke,
gr. Distrib. Verbenac., [ed. 2], 81, 97, & 191. ae F Be
C. Hoehne, Ind, Bibl. & Num. Pl. Col. Com, Rondon 349. 1951;
denke, Phytologia h: 70. 1952; Moldenke, Résumé 9), 11h, 233, “DD,
311, 315, & 463. 1959; Moldenke, Phytologia 12: 24, 102, 207,
965; Moldenke, Résumé Supp 5.
or shrublet, 0.8-~1.8 m, tall; stems erect, 0.30.6 m.
tall, simple or forked, tetragonal, hirsute; leaves decussate-
opposite or ternate, variable in size and shape; petioles short,
te; leaf-blades rigid, ovate or elliptic to oblong, 7. 10
oes lone, 2.5—- om. wide, acute at the apex, crenate-serrate a-

166 PHYTOLOGIA Vol. 13, no. 2

me te A i en globose or ovoid, many-flower ed,
6—8 mm, wide, finally becoming subcylindric and almost 1.5 cm.
long, the inaces ones sessile; bractlets imbricate, scale-like,
obovate, subrhomboid-rounded toward the apex, mucronate a + the a~
pex, incano-holosericeous on the back, cuneate~ iculate and

rose center, slightly more than 3 m. pi villous-pubescent on
b

limb short, "the lowest expanded and pena toates fruit splitting
peg 2 seeds, typical of the genus.

he type of this species was collected by Friedrich Sellow in
SZo aids Brazil, and was deposited in the herbarium of the

tograph h number
Chamisso, in his original description (1832), says merely "E
Brasilia misit Sellow." The type of be 3 attenuata was collected

by Carl Friedrich Philipp von Martius ¢ 224) near Cujaba, Mat-
to Grosso May, 1833, and is mie at tad in the herbarium
of the Botanisches Museum in M he of L. hirsuta f.

00 me seid at Yapacani, Santa Cruz, Bolivia, in 1892, and is de-
posited in the Britton Herbarium at the New York Botanical Garden.
Sotestien (1847) places L. vernonioides in his Section Zapania,
Subsection Paniculatae. Moore (1095) compares it with his L. jar

gadensis, to which he points ‘anil that it is closely rela
Steudel pnt and Walpers (1815) ee both L. Rarer
and L. a as valid spec

The Tan Fas ad found meget on campos, hilly or dry campos,
and on terraf. among rocks on high campos, in open erases soil,
sandy places at the edge of woods, dryish open grassy places on
the ascent of mountains, cerrados near roadsides, dry gras

t

tubercula lefiosa, que crece hasta el tamano de una manzana" .
The flowers are described as "yellow-white" on Mendes Magalbles
19025; "white" on be ae: poexbe 778, G. Gehrt s.n., Hoehne & G

BeDe,
with yellow et on ae ee 148] and Malme 34338; "whi white with
yellowish center" on Staines ch 1 70275 toes “con centro réseo @
amarelo" on Macedo 2288; and * Wprancas com centro amarelo" on Mace
do 2292.
In the Instituto Miguel Lillo the Herzog reference
bibliography above is cited as "Herzog, Bolivia III (1916) “ae sn

1966 Moldenke, Monograph of Lippia 167

the Rusby reference as "Rusby, Bolivia Bulletin XXVII (1900) 80,"
Material has been misidentified and distributed in herbaria as
Lantana sp. On the other hand, the P. Clausen s.n. [Minas rissa

cides is actually Hyptis microphylla Pohl in the Lamiaceae.

i ceeieeeaemcnelliameeeioed path 9 cose tinacteg gratia ad

Cenam) from Minas Gerais, and L. Riedel s.n, and Silva Manso SNe
(Herb, Martius 22h] from Matto Grosso.

In all, 79 herbarium specimens, including type material of all
the names involved, and 9 mounted photographs have been examined

me,

Citations: BRAZIL: Goids: G, Gardner 3946 (Cb); Murga Pires &
Black 2361 (Z). Maranh4o: Murga Pires & Black 2146 (N), 2319 (N).
Matto Grosso: Herb. Rio de Janeiro 16804 (Ja); F.C. Hoehne Com.
Rondon 19h (Sp), 130h (1); Malme 1692 (S), 3433 (S), 3u33a (8);
Martius 22 22 [87] (Br, F~-photo, N—photo, Si—photo, Z—photo);
Pilger 571

" Jan. 5883] (N); Hoehne & Gehrt s.n, , April 2h, 1926]
N Pie Macedo 2268 (N), 2292 2 (i, ta ; Mendes
19025 | Brad. 20163) (N); losén 1993 (Ss) vue Ge ore S)3 Reg-
nell - ea [Caldas, 18,5] (S), 111.9 (Caldas, 30/1/1865] (s,
¥—132300 » DI.9hk (6/11/1870) (e-271925), TI1.9iha (S)3 A.
Saim-Baite p Tyereiowd Widgren s.n, [1045] (S). Pard:
Murga Pires Wurga Pires S351 Gy Paulo: Edwall s.n. en. (Morro Pellado,
June 190k) (N, § cosa G. Gehrt s.n. ; 8en. [Itarapina, April 30
1923] (N, Sp—-8351); F.C. Hoehne s.n. (Mogy-mirin, ah we 1927)
(Ny N, Sp—20h11)5 W. H Hoehne 2709 (Bh, It, M1, N, Qu, J
Kiehl S.n, (Herb, Inst. Agron. Est. 3764] (N, *sp-=11982); ; peter
& Lund 207 2073 (N); Sellow s.n. (Brasilia; Macbride photos 17552]
(It=photo of type, Kr-—photo of type, Ne-photo of type, N—photo
of type, a oie of Sieh State undetermined: J. E. Pohl s.n.
( - Mus. Caes. Vindob. 1839] (Br). BOLIVIA: El Be-
ti: Fletactmecn’ 306 (S); H. H, Rusby 927 (C, Pa, Pa, Pr, Pr, W—

7553). Ua Pas La a.Fem Ki: C4rdenas 1935 (N); H. H. Rusby 926 (C). Santa
Cruz: My ; 2028 (N); T. Herzog 1484 (S); Kuntze s.n, [Prov.

bree wr Gr N, ns Sen. [Yapacani, 1892] (N);

os Arroyitos, 6.XII.1916) (N, S); Steinbach 207)
et ES {Herb. + Ae 17922] (N, Ug), 6306 (A), 7027
(Ca W, W, 8S, Ut—91356).

a Kuntze, Rev. Gen. Pl. 2 (3): a Bi 3836.
Synonymy: Lippia floridana mutes ex Moldenke, Suppl. List
valid Names » in syn. 19h1. a verticillata Legrand ex or
Genke, Résumé" 318, in syn. 19 = Lippia villafloridans Kuntze ex
Me, thed Hh ine. iC

168 PHYTOLOGIA Vol. 13, no. 2

Bibliography: Kuntze, Rev. Gen. Pl, 2 (3): 253--25h. 1898;
Thiselt .= 9 Ind. Kew, Suppl. 23 106. 190k; Moldenke
List I

3h: 9. 1933 Moldenke, Suppl. __ Names 5. 1941; Molden
e, Known Geogr. Dis rib. ye Soe , 1 & 96. play | Mol-
Alph. Invalid Names 30. 1582; nae :-

denke, List
427 ee and 1): 8. gst worden, og es afte Cit. 2: 627
1 <

» 119, 22, 328, & — 9594, eres
. 93 18h. 1961; Rambo, te 21: 28—
ny & me eter ‘aaeae Phytologia 12: bo & & “6 3965) and 13:
29. 1

Dioecious perennial shrub or subshrub, very small, 10-15 cm.
tall, few-branched, only the lowest portion woody; branches her~

tube conic, the limb oblique, l-lobed, one of the lobes larger
than the others; stigma obliquely bilobed; fruit pubescent but
scarcely glandulose as the outside, splitting subspontaneously
into 2 cocci on ma

The type of this stench was collected by Carl Ernst Otto
Kuntze at Villa Florida, Paraguay, on October 2, 1892, and is de-
ar=-

lin was photographed the:

mee it mumber 17583, but is now destroyed. The type of L.
verticillata was collected by Carlos Marfa Diego Enrique Legrand
(no. 52h) in rocky ground at Rfo Yi, department of Florida, Uru-
guay, on November 2, 1916, and is deposited in the herbarium of
the aed Nacional at Montevideo.

The species has been found growing in meadows and quebradas,
on rocky and dry grassy campos, between arroyos, and on rocky

in January November, fruit~

ground, flowering J » March, April, and ’

ing in January. The flowers are described as "yellow" o =
nal, Aragone, Campal, & Rose t PE.5SL71, Rambo Hitz
and Rosengurtt B.4501, and as Bee, on 52h.
Material has be ed and distributed in herbaria under
the names L. imbricata Kuntze and L. olia °

Kuntze (1998) says "Der Kelch ist esos flach und encore
schweisst gez¥hnt, sodass di Material zur Tre .
Lantana Camara durchgreifend ist." Rambo (1965) vie his See"
tion in Herb. Anchieta 3033 from Rio Grande do Sul, Brazile

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PHY TOLOGIA

Designed to expedite botanical publication

Vol. 13 April, 1966 No. 3
CONTENTS
MOLDENKE, H.N., Materials toward a CE ae of the genus
Rey NE ee 169
MOLDENKE, H. N., Additional notes on the genus Verbena. 1. . . 179
MOLDENKE, H.N., Notes on new and noteworthy plants. XLIII . . 218
MOLDENKE, A.L., Book reviews. ........ ‘ ees |
ALLEN, C. K., Notes on Lauraceae of tropical America I.
The generic status of Nectandra, Ocotea and Pleurothyrium zat
: _ ALLEN, C. K., Notes on vecset American Lauraceae Il.
i Costa Rica OSE gms ee ein = 232

Published by Harold N. Moldenke and Alma L. Moldenke

303 Parkside Road
Plainfield, New sungihe U.S.A.

Price of this number, $1; per volume, $5. 75, in salvanee,
. 'Boranieas or $6 at close of volume

MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. 1
Harold N, Moldenke

TA VILLAFLORIDANA K
Additional bibli pelican ‘Holdenke, ae cea 13: gy 1966.
Rambo (1965) describes this plant as a "Small plant, 1
almost rosette like, stalk of the ‘ates escence Barve Arnon
longer than the sten", growing on "Dry, grassy Campos" in the
"Northwestern Highlands only" in Rio riding ao Sul, with a general
distribution of "Paraguay, ROS".
In all, 10 herbarium specimens, including t, material of
most of the pane involved, and S: mounted shokeainatta have been
examined
Sitatlonss “BRAZIL: Rio Grande do Sul: Castellanos s.n. [Santa
do Livramento; Herb. Inst. Miguel Lillo 15763] (N)j Rambo
3033 (Rb). PARAGUAY: Kuntze s.n. [Villa Florida, 2/X/92; Macbride

Kr-—photo
of Sap dad N= to. of isotype, W-—-photo of iso-
type}. URUGUAY: astellanos we {cerce e Arroyo Catalancito; Herb.
Inst. Miguel its 15764) (N), 8 sen. [Rfo Yi; ie Inst. Miguel
Lillo 15773] (N); Gallinal, A & Ros

PE.S171 ( (N); OS or (N, Ug); 5 es 1. a

LIPPIA VOLKII Moldenke, Phytologia 5: 226-227. 1955.
Bibliography: Moldenke, Phytologia 5: 226--227. 19955 | sagen
Résumé 152 & 463. 1959; G. Taylor, Ind. Kew. Suppl. 2. 1959;

Moldenke, Biol. awe 30: 3591 . 1956; Anon., Assoc. cp ‘A.

or less Seaettemanhies striate-sulcate, glabrous or mimtely pu-
berulous in the sulcations on the younger parts, with rather few
Short side-branches; nodes annulate; principal internodes 3-3

» uniformly grayish-green on both surfaces,
oblong or sui ong-aehient aba, rier cm. long, 4-10 mm, wide, acute
or subacute at the apex, shallowly crenate or subentire, acute at
the base, glabrous and dens profane les on both surfaces;
midrib very slender, impressed above, prominulous beneath; secon-
daries very slender, 5 or 6 per side, arcuate-as perigee ye not anss-
Yowosing at the somewhat impressed above and promimul
beneath; veinlet reticulation sparse, inoonapdouems-on both sur-

ous and
tate, 3--15 mm. long; heads oblong, 5-10 m. m. 10ng,
oT mm, wide, densely many-flowered, selpars.” not elongating af
169

170 PHYTOLOGIA Vol. 13, no. 3

at
punctate, ciliolate along the margins; corolla-tube
long, densely puberulent outside, the limb about 2 m. wide,
densely puberulent on the outer face.
The type of this species was collected by Otto Heinrich Volk
_ ere in Southwest Africa

been examined by me.
Citations: SOUTHWEST AFRICA: Volk sen. (Mu--type, Z--isotype)-

PIA WHYTEI Moldenke, Phytologia 1: 428429. 190.
Bibliography: Moldenke, Phytologia 1: 428—-h29 (1940) and 1:
50h. 191; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1] 51

« Kew. Su He. Ne. & Ae Le Molde
2: 89. 198; Moldenke, Alph. List Cit. 2: 585 (1948

}, and h: 1032, 109, 1060, 1123, & 12h8
Phytologia 3: 138. 1949; Moldenke, Geo

short~pubescent; leaf-blades firmly chartaceous, dark-green above,
lighter beneath, lanceolate or elliptic, 2.5-—- cm. long, §—18
hort-acuminate at the apex, regularly crenu~

toward the tips of the branches, a pair in each axil; spikes ses”

sile or subsessile, subcapitate, to about 12 mm. long and 10 mie

wide, very densely many-flowered; bractlets ovate, about 2 mm.

long, sharply acute or short-acuminate at the apex, strigose with
hardly s

has been found growing in sandy soil,
soil at river edges, at altitudes of 1330 to 2500 meters, flower
ing in January, February, May, and August to October, fruiting =

1966 Moldenke, Monograph of Lippia 171

March, May, September, and October. Dttmmer refers to it as
"locally frequent" in’ Uganda. A vernacular name recorded for it
is "kengekioudet", The flowers are described as "white" on Dt
mer 3736 and Garcia 315, "green" on T. H. E. Jackson 423, and
"purple" on Borle 3

Brenan (195)) | a L. whytei to synonymy under L. ened
(Burm, nie acy saying "this plant appears to me to be scarce-
ly m row-Leaved, = ta ile-flowered form of L. -_

vanica."

~~ Material has been misidentified and distributed in herbaria
under the names L. adotnsis Hochst., L. africana Moldenke, L. as-
perifolia A. Rich., "Le “WL. asperifolia Rich. vel forma", L. sis
(Thunb, ) cones os L. javanica (Burm. f.) Spreng., L. oatesii
Rolfe, and Lantana salviaefolia Jacq. In all, 31 herbarium um spec=-
imens, ee ee Se the type, and mounted photographs have been

me,
a itatiate: CONGO LEOPOLDVILLE: Quarré 87 (N), 1146 (Br), =
(Br), 486 Be (Br, N), 52hO (Br, N), 530 (Br); "J. F. W." es (Br).
ee Dttimmer 37% (W—1172961). TANGANYIKA: Piemeisel &
hart 513 (W—1373312); Rounce 231 (K); Schlieben 319 m 319 (Br). KENYA:

assa: R, Monteiro 3h (U1). Zambezia: Torre 3637 (Ul).
LIPPIA a, H. H. W. Pearson in Thiselt.-. » Fl. Cap. 5:

196. 1
gynorgay: Liscin abvtsaie africana Moldenke, Phytologia 2: 169. 198.

Lantana achroleuca Dinter ex Moldenke, Résumé 306, in syn. 1959.
__ Bibliography: H. H. W. Pearson in Thisel.-Dyer, Fl. Cap. 5:

196. 1901; A. W. HILL, Ind. Kew. Suppl. 61 117. 1926; Moldenke,

122, 189, & i91 1849 M 3) wala’ eee & 293 2950)
e oldenke
and’ 3: 56—L58. 2 ; 1951; Wild, Westocss Pal ste Handb, 158, 9535 E
« ° Ind, Kew, "suppl. li: °
5: 98, 195k; Moldenke, Inform. Mold. Set 19 apse. 2 niaete and ,
Vora

Pars ; UiB150, 263, 25h, 306,
(seas, 12595 Molden, dawns Suppl, 1s 10 (2559), 3: 38 & 3

3 3 ?
172, a2, 209, 231, 250, 368, Uab, LSB, 133, ln, & b62. 1965)

172 PHYTOLOGIA Vol. 13, now 3

Wild, Kirkia 5: h. 1965.
Suffrutescent herb, undershrub, subs ti
shrub, 0.5—1 m, tall; rootstosk perennial; stens terete or sub-

angular

rather obtusely strigose with whitish antrorse
and rather densely 3 nodes anmilate; principal
5.8 cm, long; leaves decussate-opposite, swee
when crushed, he odor of 1 > th
a cluster of smaller ones on much abbreviated twigs in their

axils; petioles very slender, 1-~3 m. long, antrorsely strigose
and resinous-granular like the branches; leaf-blades chartaceous,
bri

re

serrate along the margins, acute at the base and cuneately nar—-
rowed into the petiole, rugose, scabrid-pubescent or strigose a~-
bove with bulbous—based antrorse hairs and su » densely
short~pubescent and densely resinous-granular beneath or scabrid-

the
glandular or densely strigose and
the outer ones 8 mm, long and 2.5 mm. wide,
ers

, the anterior one larger;
ovary and style together 3—3.5 mm. long; fruit shortly oblong, .
mm. long, 1—1.5 m. — plano-convex, flattened at the comis~
sure, smooth or very delicately sculptured, glabrous.
The type of this perplexing species was collected by Frederick
t+ was named near ’

Erik in the Transvaal, on
posited in the herbarium of the Naturhistoriska

1966 Moldenke, Monograph of Lippia 173

Stockholm, The type of Lantana ochroleuca was gathered by Kurt
Dinter (no. 582) in Jarmary in Southwest Africa and is deposited

in the herbarium of the Botanisches Museum at Berlin.
The species has been f. bankenveld and bush on

ound
quartzite ridges, herbaceous savannas, gallery forests, Acacia
savannas, subtropical gum
grassveld

The flowers are described as "violet" on
on Ne Ce Chase 1885 and Eyles 8770, "pale yellowish-white” on

Meeuse 9029, and "white" on Acock 8373, L. E. Codd 490k, Eyles
2228, Li benberg 2762, H. M. Lloyd 6, and Meeuse 9236.

South African ones as

1. Calyx distinctly 2-lobed.
2. Bractlets more than mm. long and 3 m, wide, surpassing
fl L,. scabe

the flowers Le
2a. Bractlets less than | mm. long and 3 mm. wide, not sur-
Passing the flowers L. javanica.
la. Calyx truncate, subtruncate, or obscurely lobed.
3. Leaf-blades serrate or crenate-serrate.

4. Bractlets more than ), m. long; calyx truncate or
a obscurely }\-toothed ; L. wilmsii,
» Bractlets less than mm, long; calyx obscurely
Sched. co on L. rehmanni .
3a. Leaf-blades crenate -L. pretoriensis.

Meeuse, in a letter to me dated December 9, 1953, says: "The
on's spe-

Lippia material is extremely variable and some

a

nt

Says: "I am inclined to batlave that Pearson's L. re
a L, bazeiana and L. pretoriensis can hardly be differe
Le asperifolia’ («+ javanica)*.
ae J. L. Warren, (rhe letter to me dated July 5, 1961, says:
Lippia are found in the area, Lippia javanica
(ara, f.) Spreng., Lippia scaberrima Sond., and Lippia wilmsii
haces The first two are easily distinguishable using bract
characteristics as in Flora Capensis but I do not consider the

17h PHI-T01LO06:748 Vol. 13, no. 3

criteria used in Flora Capensis to distinguish Lippia wilmsii
Pearson infallible, I have studied the records of Lippia pia in the
National Herbarium, Pretoria, amongst which are specimens person-
ally identified by you, but I am still not satisfied as to the
characteristics which have been used to separate Lippia wilmsii
Pearson from Lippia javanica (Burm. f.) Spreng."

Material of L. wilmsii has been misidentified and distributed

in herbaria under the nam olia Hochst., L. asperifolia
Rich., L. bazeiana H, H. W. Pearson, L. H. He We Pear-

son, L. scaberrima Sond., and Lantana salvifolia Jacq. On the
other hand, the Troupin 601 & 11.032, distributed as L. wilmsii,

are var. villosa (Moldenke) | Moldenke, Troupin II.670 is ee grandi~

folia Hochst., and J. Buchanan 1381 and Shantz 267 are L. woodii

Moldenke .

Pearson (1901) cites the following: KENYA: Scott-Elliot 68h.
ZAMBIA: Cecil 219. MALAWI: J, Buchanan 1381; Myte Sens son. [Mount
Zomba, 000--6000 ft.]. SOUTH AFRICA: Transvaal: W. ims 1180.

In all, 67 herbarium specimens, Seep el type sctaztal of all
hag a invo ved, and 10 mounted photographs have been examined

Citations: RUANDA & URONDI: a 5702 wm, oe os a
(Hi—206785). TANGANYIKA: H. M. h6 (KX)

ZAMBIA: C. E. F. Allen 32) (af, K » Rh); Herb. — Se
Stat. Mazabuke 186 (Af, Cb), 218 (af). SOUTHERN RHODESIA: N. Ce
Chase 678 (Govt. Herb. Salisbury 20186] (N), 1885 (Rh—-27086);
F. Eyles 5228 cin), = Ae Je Ce Hopleins = yen. [15/11/37] (Rb

6632); 0 p Ae de Tl 3 J. A. G. Walker
227 (Rh); 0. West yo git tr Wild ess “Ga iT580).
MALAWI: J. Buchanan 136 (N), 765 (eae, W-—-80710). PORTU-
GUESE EAST AFRICA: Mosambique: A. Peter 51166 [S.66] (B); Rattray
1103 (Rh—~-195)9). SOUTHWEST AFRICA: Ee 250 (F—photo, K, N, ¥~
photo, Sg—photo, Z—~photo); Dinter 582 (Af, B, Br, N, S). SOUTH
AFRICA: aa of edhey es Hope: J.P. He P. He a, 8373 (N, 3). Transvaal:
Breyer s ene - 18325) (Af); Le B. Codd 3372
(Ss), "iS (605 Ee. E.G 578, in part (Af), 911 (Af)s Bap-
Acock 1321 (s), 8, 8); Kassner 139 (Br); H. Lang 8+
n. Sie eameraal wis 32208] (Tm); Leendertz 1322 (Herb. Trans~
vaal Mus, 5587] (Af), 2202 [Herb, Transvaal Mus. 7326] (Af, Cb),
Sen, sans [20-%-1909) (cb) Les. Te ts Ce hs any 262 (at), 3187 (Af)s
Marlo a (Af); Meeuse 9029 j Mes s.n. Ns [106
ra (cb); Noge 14129 (ar ay Pott "S648 F. A, Rogers 25077
. Transvaal Mus. 20966] (Af); Schweickerdt 1299 fir Cb) 5 Se
he ; sil Th (Af); E. Wall ois L/TO/A938T 3), 5M. may!
F—photo, K--photo, S, Si--photo, Z—-photo), 8:
n. [20/10/938)" (x, 8)5 une 3380 fue rb. Ganttens igus 588 94
Tif—isotype, N—photo of type

1966 Moldenke, Monograph of Lippia 175

PPIA ay var. SCABERRIMA (Moldenke) Moldenke, Phytologia }:
179. 1953.
may my . pia africana var, scaberrima Moldenke, Phytologia

3: 38-39. 1
Bi acrestg: Moldenke, Pe oes cod 3s Psi (1948) and 3: 77
& 138. 1949; Moldenke, Known Geogr. Verbenac., [ed. 2],
120, 122, & 189. 1949; Moldenke, Phytologis 3: 58 ( 1951) and toe
179. 1953; veepes Biol. Abstr. 27: 1887 & 2026. 1953; Hauman
Assoc. Etud, . Afr. Trop. Index 1954; Moldenke, Résumé 149,
15h, & 463. 1959) Sodierie, —_ Suppl. 3: 16 & 33. 1962.
This fers from t un aaa Mecaace Getsaah

Rodin as a bush, 1 m, tall, with white flowers and purple fruit.
The type of the variety was collected by P. Krook [Penther
1765] along the Tsitsa River, Griqualand East, South Africa, on
rte 22, 1895, and is deposited in the herbarium of the Natur-
ska Riksmis

Lantana salvifolia Jacq. In all, 5 herbarium specimens, including
bs material of both nanos involved, and 4 mounted photographs

“Citations, SOUTHERN RHODESIA: R. G. N. Young 882 (Rh). MALAWI:
Whyte s.n. [Nyika Plateau] (K). SOUTH AFRICA: Cape of Good Hope:
Krook s.n. Jide Sai. (F--photo of type, Ne-isotype, N--photo

of type, S—type hote of type, Z—-photo of type). Trans-
Vaal: Rodin Rodin 3871 on 02252). ;
PIA WILMSII var, VILLOSA (Moldenke) Moldenke, Phytologia h:
180, i

a africana var. villosa Moldenke, Phytologia
e agape c - Lippia © wilmsii var. var. tomentosa Moldenke, Résu-

3, nom. md. 1959.
Bibliography: Moldenke, Phytologia 2: 469--70 & 1,82 (1918)
and 32 3: 13t. 19 Molden, Known Geogr. Distrib. Verbena

» [ed
al, +
te 199; Moldenke, Phytologia 3: 292° On ‘
its 58 asa,” aa om 180. 19533 uoldenke, os agi

1 By + 1953; Hauman, Assoc. ae 2 . Fi. Afr. Trop.

tpt, Sige sad Fa 142, 143, 145, 146 nih8, ake. “ey 463. 3 39595
1966, 2» Résumé Suppl. 3:°16 & 33 (1962) and 12: 6, 7, & 11.

This variety differs from the + f th ies in
ypical form o. e@ spec
the villous pubescence on its branches, twigs, petioles, leaf-

176 PHYTOLOGIA Vol. 13, no. 3

blades, peduncles, and bractlets, and in its larger flowering~-
heads, the heads being to 13 mn, "wide and the bractlets to 7 mm.
long. It is described i ty ma as a suffrutescent perennial
herb, 30-60 cm. tall, t, many-stemmed, with a taproot and
thick rootstock; sap riterd anes flowers aromatic; corollas white
or ninenk eames to pale-yellow, yellow, yellowish-browm, or

ng ri
long fringe of trees, in grasslands and coppices, on yon open
burned raha | ste, forests, and in gravel and stone
on hillsides, "at altitudes of 100 to 2200 meters, flowering in
March, May, and September to are Bec refers seo it as

poussant
de végéta," Vernacular names recorded for it are

is 22 3 a — misidentified and distribu in herbaria
names Lippia africana Moldenke, L. coparieuita Riche,
and L. wilmsii H. H. W. Pearson.

The flowers are described as "yellow" on Troupin 6014, "pale-
yellow" on Troupin II.832, "yellowish-brown" on Tanner er 1692,
"white" on J. C. Hopkins s.n., "yellowish-white” on Maas—Geester-
anus 786, and "primrose-yellow" on edhead 1155

In all, 19 herbarium specimens, including type matartan of all
the names involved, and 5 mounted photographs have been examined

me,
Citations: CONGO LEOPOLDVILLE: Cabu 66 (Br); Quarré 3
Br, N). RUANDA: Troupin 60Ll dag 11.833 3 2

¥—-pito. of of type, S—type of type, Z

TANGANYIKA: Tynes B13 D.15 ots, Dae Bs Tamer 1692 (Ca—183151)

KENYA: Maas—Gees 86 (Ca—92086, 5)j Thorold 2733 (K)«
7

E. Milne-Redhead
3.0, [15.12 peed DEE Bee

LIPPIA WOODII M » Phytologia 2: 318—319. 19h7.
Bibitserapigt over ig Phytologia 2: ms n9 & ion 19473
Moldenke, Known Geogr. Distrib. » [ede 2], 120, 122, &
191. 19493 Moldenke, Alph, Liet Cit. 38 oa (299) and a hs 1028,
nee ert a: 19h9s Molde ah
3 E. J. Kew. Hel 1953 ifolde ,
néouné 149, 25 & “ts3. 3 cia ecm 3B. 1953
stems subterete toward the base, subtet~

1966 Moldenke, Monograph of Lippia 177

ragonal toward the apex, mostly unbranched, densely puberulent,
less densely so in age; nodes not annulate; principal internodes
314.5 cm. long; leaves decussate-opposite, few, very fragrant,
usual a

equaling the subtending leaves; peduncles slender, 2—--.5 cm.
long, densely short-pubescent with brown hairs; spikes capitate,
ne cm. long, 1.2——1.l, cm. wide, densely many-flowered;

surpassing the
flowers; corolla NIDECTE MEAL Sime 3--, mm. long, densely short-
pubescent with whitish hairs on the outer surface, the limb 1—
de,

The type of this species was onliasted by John Buchanan [J.
x. wees 6937] at Blantyre, Malawi, and is deposited in the her-

of th the Chicago Natural His at sheet mumber
83373. It was named in honor of John Medley Wood who distributed

H. W. )
per sk closely beoddbiae and to. Soak “at is obviously closely

In all, 7 herbarium specimens, including the type, and 5
mounted p photographs have been examined by me.
: Citations: MALAWI: J. Buchanan 1361 (K, W—807314), 8 Sate (J.
+ Wood 6937] (F—83373— Pa ey rit of ire. | pbs
type, N—photo of type, Poy bint BPS Piss
RHODESIA: J. C. F. Hopkins he Pr nae) a, hee
7497). SOUTH AFRICA: Transvaal: S Shants 267 (W—1595613) .

LIPPIA YUCATANA Loes, in Fedde, Repert. Nov. i 9: 36h. 1911.
Synonymy : [ae eas caulis Greem., Field Mus. Publ. Bot.
rg BOS. 19,
Biblio wag Loes. in Fedde, Nov. Sp. Abe 364. 1911;

Ograp Repert.
Greem., Field Mus, Publ. Bot. 2: 340-31. 1912; Prain, Ind.

40;
Carnegie Inst. W Wash Publ, 522: 16h, 167-168, 219, 220, & 222.

178 PHYTOLOGIA Vol, 13, no. 3

peas | ton macmeet Fo Known Geogr. Distrib. — [ed. 1], 17 &
96. 1 Alph, List Invalid 29. 1942; Moldenke,
Piytologds ai : 107, "5; Moldenke maaan) “Lis t Cit. 1: 227-229,

300, 316, 319 (1946), 2: 328 ey 3: wens . 919
(slo), and kk: 1239. “13193 Moldenke, Known Geogr. Distrib. Verben-
ac. » 31 & 191. 19h93 oldenke, Résumé 37, 310, & 463.
1959; Sehdanhe, Phytologia 13: 35 &

Shrub or tree, to 3m. tall; araiee me bbls white ype

Ww

shortly and very densely hispidulous throughout above
bulbous-based hairs (the bases peietatanty: rather Herigend or
more or less densely puberulent or appressed-puberulent to hir-

tellous beneath, especially on the venation, rather more rvenntias
ous re young; peduncles axillary, 2--5 in each _
(rarely solitary), slender, often nutant, 0.8—~3 cm, long,

enc a ae YS or substrigillose; heads subglobose, co tS
mm, long, 6-~10 mm. wide; bractlets loosely eniner das *nembran-

ous, ovate or subreniforn, —7 um. long, h—-9 mm, wide,
acute or shorty subacuminate at the apex, cordate at the base,
us— ent and g

mm. wide, subvillous, its lobes obtuse and ciliolate; corolla

cles was collected by Georg

hes Museum in Berlin, where t by Mac~
bride as his type photograph mumber 175 a is now destroyed.
The type of L. albicaulis was collected by G rge Franklin Gau-

white stems, Material has been misidentified and distributed in
as L. pabeliste Cav., a mich-misunderstood epecdahe. Ft The

more striking c distinguishing atana from L. ui
bellata and the pins other species of ait go mediate group are

1966 Moldenke, Monograph of Li Lippia 179

ee smooth ashy-white stems and br. ranches, puberulent branchlets,
and dark-green elliptic-lanceolate mostly obtuse leaves which are
only short—pubescent on both surfaces and often only very slight-
80.
The species has been collected in a in Jamary and
March, in fruit in March. Gaumer 23627 & 2))11 have the younger
Lédives more densely velutinous benea eath than i is seen on any of the
other collections. Standley (192h, 1930) erroneously reduces
this species to synonymy under L. umbella liata and records the ver—
nacular name "salvia poblana", as Well as "xoltemuuc" and
"xoltémuc"

In all, 25 herbarium specimens, including type material of all
the names "involved, and 17 mounted photographs have been examined

ne

Citations: MEXICO: Quintana Roo: Gaumer & sons 1475 (B, F—
58273, S). Yucat4n: G. F, Gaumer 871 (F3667h, a), 9 971 ou a Ca—
pie De 1660, F—37319, F—photo, » N—photo

to, S——photo, Si—photo, Z—~pho oto, $Pnots). *2109 * (p= 8907,

c, N), 23627 (r-useoes, N), 23627 bis (F—6602h), 24176 (F—
552180, NS N, S), 2uh12 (F—5520h); Hunnewell 18293 (os Seler &
Seler 4918 (Macbride photos 1755)] ~ (F—€63083—photo mel —
isotype, G. G—photo of type, Kr--photo of type, Harpe d
photo of type, N-~photo of type, N—photo of type, N——photo of
isotype, S—photo of type, Z—-photo of type, Z—photo of isotype).

ADDITIONAL NOTES ON THE GENUS VERBENA. I
Harold N, Moldenke

VERBENA [Dorst.] L.
Additional synonymy: Glanduria Troncoso, Biol. Abstr. 6 (21):

tional bibliography: N. J. Anderss., G Veg
’ N, J. Anderss., af Akad, Handl. rectbe ss 185 7
18555 H. B. Davis, Works Pringle © 2, 49, 20k, 130, 132, 152,
169, & 193. 1936; ‘Christie, . 21 :
rt Phytopath e Ke) Th te “19505 Moldenke, Biol, Abstr.
i Bog (1959) and 35: uo (1): 7. Deitschman & Pruett, Proc. 17th

t. Weed Control Conf. aoe se a Pl. Hunt, Andes 2h6.
136 $61; Reitz, Sellowia 13: 67& iio. Feaanrested chats, oss Herbs & Spices

153 & 232. 1961 it. Fils, 3-23. 3961; Mol-
» Biol. incl oe 79, hha, & * 1036 (i961) aa ete a
2155. 19625 2} Brilmayer, ALL abo about, Vines 350» tg Toca dig

Pruett 1962; Hocking, Exce: nts » Aol:
292 2 (ah 13s 2 & 5 (062) ant A.6: Remar ib 8 tiakett,

180 PHYTOLOGIA Vol. 13, no. 3

ae
i8--L85, fig. 1-6. 1964; Soukup, Biota 5: 19h. 196h; A. Webster,
arib, Gard. 8, 3 129, & 138. 196; Altman Dit rs o
Data Book 49. 1964; Ma » Amat, Herb. 25 (3-): 42. 196h5
Baker, Castanea 29: 183. 196h; e

Cambridg. 179. 1964; R. Pearson, An. & Pl. Cenozoic Era 58 & 72.
1964; Loewenfeld, Herb. Gardeni ing egy 1964; Reitz & Klein,
Sellowia 16: 73 & 117. 196k; Angely, Fl. . 322 69. 1965

_—s Fis ge 45: bore “156, 377 2 7, 6658, 7026, & 8025
(1964) and 3616, & pu (1968), 6’ (10):'B.130 (1965),
L6 ais B.129 (i985), and U6 (15): B.127. 1965; Moldenke,

logia 11: 4,35—509 (1 965) and 12: 6, a, ry 48-50, 55, 57, 63,
8h, 87, 118, 205, 207, 211, ed 226, 229 230, Bee 5, & 288.
ch.

19653 Anon., Ind, Sem. Sti .
1965; Anon., Delect, Sem. oa ere ‘hay. t 3 356
1965; Cody, Ind. Sem. Pl. Res. Inst. Ot 1965: 31. 1965; Ray-
m ° art if ooo Jard « Mo e
19653 Florists’ Rev. A 9655 Recon Biol. Bags hé (9):
hort 1965; M pteesat tia Anon., Bi
tes W6 (Lh): 130" 6s), ar ie i: ‘yaa8 9es$ and (aay:
ars & B.128. 1965; M Résumé Suppl. eg ‘ 2, 3 7
» 12, & 13. 19653 hoon, eae Etad. I

. wre
1964: 56. 1965; Rambo, Pesquis. Bot. 21: 3-6, 58—[60] . 19655
Goodies Loveless, * soe Fl. Barbados 36, & 485. 19653 Neal,
. Hawaii, new rev, ed., 720-722. 1965; Moldenke, Phytolo-
gia te: “79-80. 1966; L. M.’Reid, Park Managenent Ser. Bull.
&

In all, 39,033 herbarium specimens and mounted illustrations,
as well as 1750 literature references, are cited in the main
body of this work, of which this is the first supplement.

Addi tio: abbreviations for the names of herbaria whose ma~
hensaege is cited in this wark are:

arhus University, Aarhus, Denmark
Av = Facultad de Agronomia y Veterinaria, Universidad de Buenos
Aires

» Buenos Aires, few
Ib = Institute de Botanica Agricola del I. N. T. A-, Buenos
Aires, Argentina
Ip = Escuela’ Nacional de Ciencias tena Instituto Politec-

co Nacional, Mexico aa adie
Brilmayer (1962) speaks o ee ee "Yerbens as follows, but

Nutt., and V. peruviana (L.) Britton: "Tender perennials, hardy
in Southern g grown in cooler climates.
Wh growing seasons are short, they should be s

1966 Moldenke, Notes on Verbena 181

VERBENA ABRAMSI Moldenke

Additional heme Moldenke, Biol. Abstr. 37: 2h55.
1962; Hocking, » Bot. A.6: 53h. 1963; Moldenke, Biol. Ab-
str. ss 6658. 196k) Mcldenie, Phytologia 11: 437. 1965.

xVERBENA ADULTERINA Hausskn.
Additional bibliography: Hocking, Excerpt. Bot. A.6: 53h.
19633 Moldenke, Phytologia 11: 437. "1965.

ALATA
dditional bibliography: Hocking, Excerpt. Bot. ge ege
1963 serene Biol, Abstr. 5: 3136. 1 eee tere
: bs. Pesquis, Bot. = jl—32 & [59]. 19s.
‘a 396) cites Herb. Anchieta 6 838 [Bruxel s.n.] and 3159
sees Santa Catarina, 113k, 13h, 28735 [Bmrich s.n.], 29983 [Barich 8. 8.
fe), 32302 323025 35416, 1,083, 083, W585, 4819, Ugly 1629, MEO
9 9760, 520 52606 [Ear [Emri. h sn), 53933, 39 7.
5735, S80h6, 56353, 6198, 60516 [Sacco s.n.], and 64063 [Sacco
] from Rio Grande do Sul, and 35956 [Hatschbach 8h] fron Pare-
hims

RGS"
Additional citations: BRAZIL: Sbate undetermined: Sellow Bete
[Macbride photos 17399] (W—photo).
VERBENA pase « ALBA Molde: pao BS
Addi ti mal bibliography: Mol i 8: 148—1h9.
1961; Hocking, Excerpt. Bot. La fet

Addi ta Sere Biol. Abstr. 37: 2455.
onal bibliography: M oldenke, Biol.

1962; Hocking, Excerpt, Bot. LoS: ul (2962) and A.6: 534. 19635
Koldenke, Phytologia 11: 437138, bhO, bhB, 488, 490, & h91.

Rzedowski 366, distributed as V. ambrosifolia, is ac~
V. andrieuxii Schau., E. M. Marsh 21,8 and C. . Ro

vy. Tipianats tide Nutt., and &, G, Marsh 1188 & 152) are

Vv
Z:
Is

I

oe
‘a

tional ci apa pes : C, M, Rogers
On) 188 (Mi), eons. (), 6033 33 (lS), x), 60M “ony. OKLAHOMA :
2123 (Mi TEXAS: Br r Co.: Rose-
doon 1373 (ta) ; B. H. Warnock 20121 (14),
56 OB} og Loan 3250) « a" MEXICO: Union Co.: C. M.
ers 5956 (Mi : Coahuila: E.G. Marsh 707, in part (Au),
<8. (Loma Aaa ts eg

# fi
ce
rela
i
nk
Fe

i?)
o
©

182 PHYTOLOGIA Vol. 13, no. 3

AMBROSIFOLIA f. pee berry

Additional bibliography: - Bot. A rag : hh. 1962;
Moldenke, Phytologia 10: 92 Cash)” and rete teks 3

Additional nal citations: ARIZONA: Cochise Co.: Deaver 6579 (Ac).
MEXICO: Guanajuato: J. Rzedowski 9542 (Ip).

VERBENA AMOENA Paxt.
Additional bibliography: Moldenke, Phytologia 10: 92. 196.
— collectors have found this * plant in flower and frui
in June and July.
Additional citations: MEXICO: Federal District: Paray 27> 2928
(Ip). México: D4vila s.n. [23.01 .1963] (Ip). CULTIVATED: India:
Khoshoo 1 (Z).

VERBENA ANDRIEUXII S
Additional ne Moldenke, a Sonn 10: 92. 196k.
edowski ki found this plant growing a the edge of an arroyo at
2250 meters altitude, with purple flowe: Loox: in flower and fruit in
September. Material has been misidentified and distributed in
herbaria as V. ambrosifo. lia Rydb.
Additional citations: MEXICO: San Luis Potosi: J. Rzedowskt
4,366 (Ip).

VERBENA ARAUCANA R. A. Phil
Additional bibliography: “Moldenke, By Sage gem 11: 38. 1965-
Additional citations: CHILE: Linares: . Philippi sen, [Lin
ares; Macbride photos 1700] (W-—photo). a Stanesr ago: Macbride
photos 343 (W--photo).

VERBENA gan Moldenke
Additional ography: , Excerpt. Bot. A.5: 45. 1962)
oes tg oa ge 8: 379. eoeze Moldenke, Biol, Abstr. 37:

xVERBENA ARGENTINA Moldenke
Additional bibliography: , Biol. Abstr. 37: eed ee
— Excerpt. Bot. A.5: ie. 6 1962; Moldenke, Phytologia 10

""tatteat uths _Ptaogis 3: tot 36
bibliography: i Moldenke, logia 11: o 17026
Additional citations: BRAZIL: Matto mouee. Moore 1083 [Mac~
bride photos 1703] (W—photo of type).

VERBENA ATACAMENSIS R

Addi tional Mbticmoone. Moldenke, Phytologia 11: 138. 1965

Recent collectors have found this plant in flower in November
and in fruit a October and November. The flowers on Ricardi Ci 24,70
are described as having been "white".

Additional citations: CHILE: Antofagasta: Ricardi 2470 (2),
3569 (Ac). Maule: Pfister s.n, [Termas de Canquenes, 3-11-1952)

1966 Moldenke, Notes on Verbena 183

(Ac).

VERBENA BALANSAE Bri
Additional bibli Mogranty: Moldenke, Phytologia 11: 438, 1965;
Rambo, Pesquis. Bot. 21: 58. 1965.

VERBENA BERTERII (Meisn.) Schau.

Additional bibliography: Moldenke, Phytologia 11: 438-39.
1965; Moldenke, Résumé Suppl. 12: 10. 1965.

Additional citations: CHILE: Cole chagua: Bertero 7); [Macbride
photos 7853, in part] (W--photo of isotype). Valparaiso: Bertero
1389 [Macbride photos 7853, in part] (W—photo).

VERBENA BIPINNATIFIDA Nutt.
Additional biblio ography: Loes., Verh. Bob. Ver. Brand, 53: 75
[Abhand1, 2,0]. 1912; Brilmayer, All about Vines 350. 1962; Hock-
Excerpt. Bot, Abs 53h. 1963; Moldenke, parang ll: 439——

LLL NY cee

respecti
Brilmayer (1962) says at this plant "tender trailing perennial
grown as an : leaves and clusters of lilac
flowers. Native of some aidedataia states." McCart found the
cal

Over limestone of the Cretaceous Eagle Ford Group. The E. G.
wash 152h 152h, distributed as V. bipinnatifida, is actually V ve , cili-
ata Benth,

‘Addi tional citations: OKLAHOMA: Cimarron Co.: C. M. Rogers
L705 (Mi). TEXAS: Bee Co.t R. L. Crockett 7965 (Id). Bexar Co.:

lucas, & Barkley ~ Kerr Cos: R. L. Crockett Ld).
Kinble 6 ese Demaree 8513 5 i) Kinney C Co. Strother 263 (Ip).
‘cltullen Co,: Alvarez, , Salazar, & See
222199) ; Sele TF Ue ee. Tarrant Co.: Mecart 60920 (Au—
236817). “Tom Green Co.: Ballinger s.n. [april 9, 1961) (Au

(Ld). MEXICO: Coahuila: E. M. Mare (an—213884). Durango:
Powell & Edmondson 961 (Mi). ae Wate Par Paray 1201 (Ip).

VERBENA BIPINNATIFIDA var. LATILOBATA Perry
wa te) gga bibliograpty: Moldenke, Phytologia 11: 137, ho,

18h PHYTOLOGIA Vol. 13, no. 3
xVERBENA HARDI Moldenke

Additional bibliography: Moldenke, Phytologia 11: hkl, 448, &
462. 1965.

BONARIENSIS L.
Additional bibliography: Hocking, Excerpt. a a $ 2

Moldenke, evils a ll: a—b2 & 470 (1965) and
1965; wag gL 6 Suppl. 12: 1. 1965; Rambo, sab ag
21... Jee & (39 ]. 19

5.
in found this species growing on boggy river margins,
in gallery Fouriee and ruderal in Santa Catarina, Brazil. The
Pedersen 614, distributed as ¥. bonariensis, actually represents
vars agers Briq. The R. ee Crockett 825, cited below,
may actually not have been collected in the county from which it

ter 233) at the British
Rambo (1965) cites awe peers 3521, augis [Reitz hows
and 61150 [Mattos s.n.] from Santa . Santa Catarina 603, |, 832; y, 3036,

DA, 2724, FO See ees 2s 2 2

25693, 26266, 28486, 30933, 32965 [Streider s.n.], 35458 (Ritter
s.n.J, 36u12, 3754 (Emrich s.n.J, 36179, 38325, 36108, 1,08, 38916,
38973, 3981, wus, ols, Lota, h07h, 11219, 43510, 1,3906,
, & g281 , 19696, 50690 [S einen L155],

1003 (Sehnen SESE. 51361 Tbbh, $330 » 51305, 50 59120 [Pivetta
983), 59122 [Pivetta aay as [Sacco 630), 60929 [Camargo
1625], 61150 [Sacco 666], 62709 [Cc © 2363), 63285 [Sacco 705 705],
63818 (Sacco 39], and 640k (Sacco 410] from Rio Grande do Sul,

Brazil -- all his own collections except where otherwise ‘indica~
ted. He indicates, however, that numbers 3558 & 38,08 have
otherwise been identified as var. conglomerata Briq., while
41219 has been identified as V, inamoena Briq. He comments that

a "Y, inamoena,.....4s at best a variety of V. bonariensis L." He
describes the species as eres stiff, psveereys
branched, up to 1 m high", growing on "Dry to somewhat moist,

bare or more or less vegetated Gihoe |: cs emeee over the
whole of the Campos area” in Rio Grande do Sul, with a general
distribution "From southern USA through the whole of tropical ant
subtropical America, with several varieties."
Additional citations: LOUISIANA: Was
50721 we TEXAS: Jefferson Co.: R. L. prin et has tia):
uanabara: M. C. Vianna 23 [Herb. Cent. Pesq. sq. Florest.

1673] a Rio de Janeiro: Castellanos 256k [Herb. Cent. Pes
Florest. 3179] (Ac). oa Catarina: Smith & Klein 13968 (Ac)>

:. Buenos : Boffa s.n. (Berisso, UL m1/19 Mi) 5
Hicken s.n. [rar 1906]. (K)5 Rufz Huidobro 1666 (Mi), 1695 (i),

1966 Moldenke, Notes on Verbena 185
771 (Mi). PORTUGUESE EAST AFRICA: Lourengo Marques: F, A. Men-

x
donca 2973 (U1); Torre 2239 (U1), 728 (U1).

BONARIENSIS var. CONGLOMERATA
Additional bibliography: Rambo, pean Bot, 21: 33. 1965;
Moldenke, Phytologia 11: 2. 1965.
Pedersen found this plant growing i t ground.
Addi tional citations: ARGENTINA: PAB Pedersen 61) (S).

VERBENA BRACTEATA Lag.

Additional bibiogrepiy & Rickett, New Field Book Am. Wild Fis.
280, fig. 98. 1963; Ho king, Excerpt. Bot. A.6: 53h. 1963; Sharp
& Baker , Castanea 29: 183. 1965 Moldenke, Phytologia 11: 40,

» LLB, pitt, & 176. 19653 Cody, Ind. Sem. Pl. Res. Inst.
Ottawa 1965:
tional re Reged te Rickett, New Field Book Am. Wild Fls.
fig. 98. 1963.
Sharp & Baker (196) state that this esa "seems restricted

to the western tier of counties" in Tenness but a glance at my
distribution map, published in Phytologia, volume 8 (1962), shows
this to be incorrect. Muhlenbach 1525 was originally identified

by the collector as "V. simplex x agen but I do not think
it represents a hybrid, at least insofar as can be seen from
Specimen cited below
Additional citations: "VERMONT : Chittenden Co,: A. R. Moldenke
2133 (Ac). MASSACHUSETTS: Bristol Co.: W. R. Taylo rE oe (i).
INDIANA: Saint mt: Josmh Co.: W. We n Jr. 5.0. x 30)
(Mi). Saint Louis: Muhl ae ie), 3h Se 312
(Ac), 3525 aoe 1625 (Ac). UTAH: Rich Cow: M. Ee Jones sn. 1. [Al-
ta, Aug. 2, 1879) (i (Mi). NEVADA: Clark Co.: Clok ay $096 8096 (ii). CO-
LORADO: Las Animas Co.: C. M. Rogers 435 (ii), 36 (Mi). OKLA~
i Cimarron Co.: C. M. Rogers 1,668 1,668 (Mi); rater 3139 (N)-
TEXAS: Dawson Co.: Lundell & daniel a ty (Id). Ochiltree Co.:
Correll & Ogden 28352 (Ld). Go.: De Se teers 30290
(ld). ARIZONA: Coconino Co: gree $87 (ac).

BRACTEATA f. ALBIFLORA (Cockerell) Moldenk
Additional bibliography: Etige bis Excerpt. Bot. ri bi 53h. 19633
Moldenke, Phytologia 10: 96. 196l1.

VERBENA SRASTLTENSIS Vell
Additional b: bliography: Hocking, Excerpt. Bot. A.6: 53h. 19635
es Piytalogia 11: bh2, bhS—bk6, & 502. 1965; Moldenke,
sumé Suppl. - 1965.
Pedersen tones sis plant growing in sandy rather moist ground
formerly plowed and much trampled
Additional citations: ALABAMA: Shelby Co.: Demaree 50643 (Ac).
IANA: DeSoto Par.: Demaree 50829 (Ac). Plaquemines Pa
Raree 50745 (ac). Saint Tammany Par.: Demaree 1999) (Ac), 5076

186 PHYTOLOGIA Vol. 13, no. 3
(Ac). Washington Par.: Demaree 50723 (Ac), 5073h (Ac). BRAZIL:

(Ac), 23599 [Herb. “Gent. Pesq. Florest. 2005] (Ac). Séo Paulo:
G. Eiten en 1595 (W—2369602). CHILE: Linares: Ricardi 2826 (Ac).

VERBENA Moldenke

Additional synonymy: Verbena cabrera Moldenke ex Hocking, Ex-
cerpt. Bot. A.5: 45, et Loe

Additional bibliogran hy: H , Excerpt. Bot. Aces Bs 1968)

12.

VERBENA CALLIANTHA Briq.
Additional bibliography: Hocking, Excerpt. Bot. A.5: 5. 1962;
Moldenke, Phytologia 11: h6. 1965.

VERBENA CAMERONENSIS L. Davis
Additional “yee a Moldenke, Phytologia 11: Wh6 » 1965.
e flowers of this plant are posted as "purple" on R. Me
3983 and J. Ruedowski 7360 and as “lavender" on Re M. King
42k9. “This same collector describes the plant as locally abun-
dant along eet road in partial shade" and as “abundant weeds"
in "open sun" in San Luis cere Be Mexico. Recent collectors
have found it in fruit in March and April. Material has been
misidentified and distributed in herbaria as V. delticola Small
and V.

Additional citations: MEXICO: San Luis Potosi: R. M. Bs
3983 (Mi), L2k9 (Mi, W-2365016); J. Ruedowski 7309 9 (Ip, Ip
7360 (Ip), 3871 (Ip). Tamaulipas: J. A. de la Cruz sen, [12.1¥
1961

VERBENA CAMPESTRIS Moldenke
ae bibliography: Mo p, Hiielesta Li: bh6. 19655
Pesquis. Bot. 21: 3h hey & t {8 Ppt
were (1965) cites “be Anchieta lisa fet ts 3348) and 48330
(Reitz 3486] from San rina, Brazil. He descri ribes

far, SC only*.

CANADENSIS (L.) Britton

Additional bibliography: Loes., Verh. Bot. Ver. Brand. 53: 79
[Abhandl. 240]. 1912; Hocking, Excerpt. Bot. A.5: bh (1962) and
ahs oe. 19635 sRickett, New Field Eee Am, Wild Fls. ai ae

3 Mo. e, Phytologia 11: 440, 4hl, 4h3, bh6-—

455, 485, 86, 188, & 90. 5965. :

Additional illustrations: Rickett, New Field Book Am. Wild
Fils. fie. 98. 1963.

tional citations: TEXAS: Chambers Co.: R. L. Crockett 8310

1966 Moldenke, Notes on Verbena 187
(Ld).

VERBENA CANADENSIS f. CANDIDISSIMA (Haage & Schmidt) Palmer &
sogitttonal bibliography: Moldenke, Phytologia 11: 48 & 450.

VERBENA CANESCENS H ote
Additional nymy: Verbena canes H.B.K., in herb,

Additional tbl segranlyt te oes., Verh. — Ver. Brand, “yet 74

{Abhandl, 239]. 1912; H. B. Davis, Life & Works Pringle 21 & 130.
1936; Moldenke, Biol. Abstr. 5: 7026. 196145 gore ee
Be 450-451, 502, & 503. 1965; Moldenke, Résumé Suppl. 12:

nguez and McCart found this species growing in hard
silt in Tamaulipas. Graham and Johnston found it at

sandy 1
500 feet altitude in the same state, while Rzedowski encountered
Poto;

4251 are described as eres but on Gonzalez Medrano s.

Re as "blue". Rzedowski found it growing in . abandoned fields.
The R. L. Crockett a6, cited below as from Blanco County,
Texas, may actua ome from Travis County —- its

mally ha ts la
3 sage merely "Johnson pepe eg while 8077, cited from Nu-
Se may actually be from the Federal District, Mexico —
its label reads "Monterrey—Mexico r
tributed in herbaria

Material has been misidentified and aa
under var, roemeriana (Scheele) Perry. On the

ist Small. The 0, M. M. Clark 7059 Parson & B Kley 1

G.L. Smith 221, in part, cited by me in Figtchogia 8: Li? aes 13963)
aa to be V. ¥. pinetorum Moldenke.

Additional citations: ee Blanco Co.: R. L. Crockett 8626
{ia). a MEXICO: Nuevo Leén: R. Le Crockett Sort Cd ak en

si: J. Rzedowski 251 = > 5278 (IP), 20} OP) Ip),
§3h7 (Ip), 9571 (Ip). oh (tp) : — Dominguez & ] art 8227 (Au
2222 2208, Au—236267) ; Gonzalez Medrano $.n- Res a Mezquite,
18 IIT 1963} (Ip); Graham & Johnston 1,707 707 (Mi).

CANESCENS f. RA Moldenke
= Additional bibliography: ging nar Abstr. 45: 7026.
964; Moldenke, Phytologia 11: 51.

VERBENA CANESCENS var. ROEMERLANA bas Perry
“ongecteare reeset Verbena canes var. roemeriana (Scheele)

dit thorn Phytologia 11: 451, 502,
mal nt °
bibliography: Moldenke, gi ’

188 PHYTOLOGIA Vol. 13, no. 3

Gould found this plant growing "in hehe dark loamy soil on
rocky knoll" in Coahuila, while Salgado encountered it in
"sandy soil associated with Larrea, name teloua, and Flourencia."
The J. Rzedowski 251, 5278, 78, 6165, & 63h7,. originally distri-
buted as this variety, are are actually all typ typical V. canescens H.

Additional citations: TEXAS: Hidalgo Co.: M. L. Walker 66 (Ld).
Kinney Co.: Strother 26, (Ip). MEXICO: Coahuila: Fuentes te Sn.
bees wa (Ip); F. We W. Gould 10656 (Mi); Salgado s.n. och Se

LINA L.
Additional bibliography: Hook., Trans.
20: 195. 1847; Loes., Verh. gee Ver. Brand. are “mh need
239]. 1912; in B. Davis, Life & Worka Pringle 132. 1936; Moldenke,
Biol. Abstr. 36: 036. 1961; ooking’ Excerpt. Bot. A.5: hh.

escribed as "blue" on Eesdlars 8677 & 8699.
ede Ler hia ekee raat the es ad on steep slopes Pe a
in Durango. Material has been Sigldehtified and and dis-
tributed in herbaria as V. officinalis L. On the other hand, the
Scora & Scora 252, distributed as V. carolina, is actually V.
litoralis H.B os
Andersson (1854) speaks of a "Verbena caroliniana var. foliis
incisis, laciniis grosse serratis Hook." fae, however (1647),
refers to the pla plant as "VY. polyst: polystachya var. foliis incisis seer
tis grossé serratis"], based on a Darwin collection fran Jam

(Mi). Federal District: J. Bandon 2 2078 Tp) México: M.
Abarca 13 (Ip); Cota 45 (Ip); aray 2h)9 3 Paray 2uh9 (Ip); J. Rzedowski 15832
(Ip). Sonora: Gentry, sca & Arguelles 19303 3 (W—2366189)- °
GUATEMALA: Huehuetenango: Breedlove 8677 (Ac), 5699 (Ac).

VERBENA CAROLINA f. ALBIFLORA Moldenke
Additional bibliography: Hocking, Excerpt. Bot. A.5: li. 19625
—, Phytologia 10: 101. 196k.
dditional citations: EL SALVADOR: Santa Ana: Tucker 1307 (iit).

VERBENA CHEITMANIANA Molde
BO oon Mihai weekee ke Moldenke, Phytologia 11: 52 & h69-

VERBENA CHILENSIS Moldenke
Additional bibliography: Moldenke, Phytologia 11: l52. 1965
Additional citations: ARGENTINA: Mendoza: R. A. Philippi 5-0:

1966 Moldenke, Notes on Verbena 189

Deettiona:; Herb. Mus. Nac. Hist. Nat. Chile h2hh6 & 5790; Mac-
ride photos 1717] (W--photo of isotype).

Ls spe peenrets Benth.
tional synonymy: Verbena cililata Benth. ex Moldenke, Ré-

sad Be Suppl. 12: sts in syn. 1965. Verbena ciliata var. ciliata
Wiggins, in her

Addi tional saNiaabwaaie. Loes., Moan bats Ver. Brand. 53: 75
[Abhandl., 2,0]. pa ie s prt — Works Pringle ve 1936;
a? Résumé S ee 3 Moldenke, Phytologia

: hhO, 4h8, \S2—-L53, 190, 1st, t 193 (1965) and’13: 181 & 183.

Johnston found this species growing in pale clay loam at the
edge of a field in Coahuila, and Gould found it in Gry. soil
of an open field with mesquite cover in Chihuahua. The flowers
on S. 5. Mite White 2018 are described as "blue", while those on J.
Ruedowski 3051 were "purple". The R. L. Crockett 8091 & 8138,

1201,
distributed as V. ciliata, are are scene ee W, Eipinatifia it Tutt;
c. Cantu 69, Claflin 41, Montemayor and Ramirez, ae & Mc
Cart 876 8763 are V. ciliata var. Seeger nk Perry; N4& Sone
(gosto 195] is V. eleg elegans H.B.K.; and Wiggins & Wiggine 16073 16073

SV. gooddi Brig.

Additional arhebbiones ARIZONA: Coconino Co.: Cureton i); (Ac).
MEXICO: Chihuahua: F, W. Gould 8755 (Mi); S.S. White 2018 (Mi).
Goamuila: Re L. Crockett 6138 (Id); M. C. Johnston 2600 (Au—
251801, Mi); E.G. Marsh 1186 (Au—212966), 152 (Au—213393).
Federal District: P Paray 1211 (Ip). Hidalgo: J. “J. L. Castro 17
Ps Galvan sen, [28.¥1I.1963] (Ip); Gonzdlez Quintero 349 (Ip);

J. Rz

Crockett 8091 (Id). Puebla: C. Galderén sone [19 IV.196h) (Ip).

San Inis Potosf: Je Rzedowski 3051 (Ip, Ip), 43h (Ip), 8801 (Ip).
a: Galicia s.n. [27.¥.1962] (Ip).

VERBENA CILIATA var, LONGIDENTATA P

Additional bibliography: Moldenke, ey Piytologts 11: 40, 448,
\s2, ate! <M . ihe 1965; Moldenke, Résumé Suppl. 12: 1. 1965

8 variety has been "misidentified and distri

ted t in tethers J the misspelled name V, cililata Benth. ond
species has been f ound by recent collectors growing rowing in black
Clay, red or black soil, marl, sandy silt, Pleistocene gravel,
and hard~packed dark gray soil. The Flores call it "abundant
red soil", The toh ees 64, distributed as this variety,
18 actually V, bipimatifida Nutt.

Additio Co.: C. Cantu 69 (40235231).

mal c itations ae Bee
Dimmit Co,; R. Acosta 80 (Au—232739) ; Chaves, Flores, & Hein 7h

190 PHITTPOLOGIA Vol. 13, no. 3

(Au— — Duval Co.: R, Acosta 9 (Au--232661); P. Cabrera
56 (Au——233116) ; Noyola 21 yal ~(au--233118). Hidalgo Co.: Be er Croc-
ett 8177 (La); Gonzalez & & Quesada 33 (Au--23311;). Jim m Hogg Co.
Raquel Bruno yy (ku——23231)). Jim Wells Co.: Rebecca Bruno
Coa s Gamboa & Dohnke 151 (Au--233117). LaSalle Co.1—
Flores & Flores 126 (Ku——232912) 5 A. Sanchez 100 Die Shas °
Maverick Co.: Te Brant 12 (Au—~232631). Medina C
Laurel 110 (Au——232656). Nueces Co.: Claflin yh le
Webb Co.: ye: Cisneros 23 (Au—-233108); Ramirez, Lopez, & & McCart 8763
(Au—-233107). MEX . MEXICO: Tamaulipas: Montemayor 31 (Au--235227)«

VERBENA CILIATA var. PUBERA ( eene) Perry
hs ea dd bibliography: Moldenke, Phytologia 11: 453, 490, &
91.1 e

VERBENA CLAVATA Rufz & Pav.
Additional bibliography: Moldenke, Phytologia 11: 53. 1965.
Additional citations: PERU: Ancash: Weberbauer 2768 {Macbride
photos 171,16] (W—photo), Arequipa: Meyen She [Pampa Grande;
Macbride photos 17,09] (W--photo). ys Rutz s.n. [Macbride
photos 1708] (W--photo of cotype).

VERBENA CLOVERAE Moldenke

Additional synonymy: Verbena cloveri var. eilacina Moldenke,
Résumé Suppl. 12: a in syn. 1965. er SS nd

lilaciana ae , Résumé Suppl. 12: os

ry ma. ub ography — enke, Pigtologie ae en & 502.
1965; J olsieite Résumé Suppl. 12: 2 & 12.

e flowers are dsacttioad as "blue" on B. e Turner 4978 and
bhace on Alvarez, Guajardo, Salazar, & McCart , T1836 “The spe-
cies has been found in fruit also in February and November and in
flower in November. Recent collectors have found it oie in
sand, red loam, sand with small gravel, deep sandy s
sandy marl silt, orange-colored sandy loam, loose fete ee
sand, loose or very loose light-tan sand, and deep a all
loose sand. The Flores call it "abundant =

: Atas

(an-229140) Brooks Co.: Painter & “parley 1318 ‘yiaie (ia) heat
Cos: Dickey 61 (Au—233352). Duval Co.: Alvarez, Guajardo, Sa~
lazar, & McCart 7657 (Au—-222268, Au—-222269). Frio Co.: Painter,
Lucas, & Barkley 1230 (Ld). Jim Hogg Co.: Alvarez, Guajardo,
Salazar, & McCart 7783 (Au—222267); Rebecca Bruno }1 Lilie
232671); P P. Cabrera 57 (Au—-232672). “LaSalle Co: Alvarez

ardo, Salazar, & McCart 761 (Au—222263); Flores & & Flores ad
Ku—222h69) 5 K. Sanchez 7 (Au—-233h31); M. de J. Solis lie 122

—- ———_—

(Au~-233351). Starr Co.: R. Acosta 7h eae ae P. Cabrera

1966 Moldenke, Notes on Verbena 191

22226). Zapata Co.: Raquel Bruno 31 (Au—232753); Ramos, Sando-
val, Salazar, & McCart 78)6 (Au—222266) ; A. Sanchez 87 (Au
233106); Ke M. de J. Solis 79 Pe neon MEXICO: Ta reeendisies Do-

VERBENA CORYMBOSA Ruf.

Additional eetareresie: "wold enke, Phytologia 11: 5h. 1965;
Rambo, Pesquis., Bot. 21: 35 & [59]. 1965.

Smith & Klein found this species growing ruderal, in oe fields,
aoa gallery forests, and by small rivers in Santa Catarina,
bears

Rambo (1965) cites Herb. Anchieta 50793 [Reitz 3770] & 54307
(Reitz 1970] from Santa Catarina and 3472) & 50505 from Rio Grande

im high", growing in "Rather moist places, densely vegetated" in
the "Northeastern Highlands only" in Rio Grande do Sul, with a gen-
eral distribution of "Peru, Chile, » RGS, SC."

ditional citations: BRAZIL: Santa Catarina: Smith & Klein
22998 (Ac), 13026 (W—-251610), 13460 (Ac).

VERBENA CUMINGII Moldenke
Additional bibliography: Moldenke, Phytologia 9: es 1963.
The species has been collected in flower in Octobe
OP hoa citations: CHILE: Coquimbo: pacticicha | & Matthei
Zz).

VERBENA CUNEIFOLIA Rufz & Pav.
: ogiditional bibliography: Moldenke, Phytologia 11: 5) & 48h.
Saunders describes this plant as very free-flowering, to four
Bote tall, with a strong sweet scent and bluish-white flowers.
: PERU:

G. E. S
berartaant undetermined: Dombey s.n. [Macbride photos 17111}

VERBENA DEAMII Moldenke
Additional bibliography: Moldenke, Phytologia 11: hh8, 454—L55,
& h62. 1965,
VERBENA DELTICOLA Small
960s tonal bibliography: Moldenke, Phytologia 11: h48 & 455.
The Ry L. Crockett 8043, cited below, is inscribed "Mercedes—
Monterrey Road", so the exact locality of collection is not known.
© Species has been found by recent collectors in gravel and on
@d sandy hillsides. Rzedowski a 2 was encountered at 2750
neters sititude and had "purple*
Material has been misidentified and distributed in herbaria as
Ye timidula Perry. The Rzedowski 7309 & 7360, distributed as V.

192 PHYTOLOGIA Vol. 13, no. 3

delticola, are actually V. cameronensis L. I. Davis.

Additional citations: TEXAS: Hidalgo Co.: R. L. Crockett 820
(Ld), 867 (Ld, Ld), 869 (Id), 8185 (1d). MEXICO: Coahuila: E.G Ge
Marsh 1822 (Au—-219621) . Durango: Paray 3243 (Ip). Nuevo Leén:
R. L. Crockett 8043 (Ld); Diaz Luna sen, 126.1x.1959] (Ip); Do- Do

soviet San Luis Potosf: Paray 565 Sta J. Rzedowski 7101 (Ip),
10026 . preg eons

VERBENA DISSECTA Willd.
Additional bibliogra aphy: Troncoso, Darwiniana 13: 68, 70,
ee Sy Moldenke, Phytolopia "ua: Mss. 19653 Moldenke, fog.
é Suppl. 12: 5. 1965; "Rambo, Resquis., Bot, 21: 35—37 & [59].

Cabrera sone is said to have had "rose" flowe
Rambo cites Herb. Anchieta 9, 1837, 303h, 3038, 3043,
3939, 2h2, "1307, 1,362, Te ee, oe 3587, 8890, BE

Shs Tiere ak OE ees Ct ote Pace

27686 [Henz

Santee! Dimers. Saetegs? Seek, | Meee oe eee

Ob, 480 4807) [Sehnem 31661, 48478 [Sehnen 2128), 51378, 78, 51652,
36208, 56225, 61723 [Camargo 2567], 6212h [Cam [Camargo 2022 2022], 62518
(Cama Camargo 2126], & 62995 [Schlichting s.n.] from Rio Grande do
Sul, and 54316 (Reitz 993] from Santa Catarina, Brazil — all
collected by himself except where otherwise indicated. He says
that the plant is "Prastrate to ascending, branches abundant,

ore or less wet places, preferably sandy meadows
and rocky outcrops.....0ver the whole of the Campos area, espec-
ially on the northeastern Highlands" in Rio Grande do Sul, with
a general distribution "From S, Paulo to Uruguay and central Ar-
gentina to Patagonia",

Troncoso (196) says: "Gl. dissecta ha sido citada originari-
amente de Chile en base a material de Luis Née, recogido a fines
del siglo XVIII, durante la expedicién Malaspina. Todas las ci-
tas pos steriores. para Chile son pie ala recoleccién de Née.
Moldenke en su monografia del género Verbena.....la cita de Bra-
sil, Bolivia, Uruguay y Argentina, pero no de Chile. i "al

material estudiado por vat seflala un frea limitada a Buenos Aires,
Uruguay y sul de Brasil. Es muy probable pues, qe cono'en sae
casos ya verificados, el ejemplar de Née, ere de Chile, haya
sufrid ntina o Uruguay-

st
2]
3

me
bot4nicos de la Peninsula hispano-lusitana: 183. t8e8) . Verbena
oe f, latilobata.......basada en un solo ejemplar de le Mon
tevideo, Uruguay (Berro 751), presenta m4s similitud con Gl.
pessenre: que con Gl. pu pulchella, debiendo incluirse en la primera

1966 Moldenke, Notes on Verbena 193

Difiere notablemente de Gl. pulchella, no sélo por la forma y el
tamafio de los segmentos pn = 3-——-7 mm lat. y 7 mine obtuso),
sino por sus caracteres florales y tipo de pubesc El ta-
mafio de la flor, del cAliz y de sus dientes, el Pree y forma de
las bracteas, la pubescencia del pelitos extendidos, no aplicados
(no estrigosos) Son caracteres coincidentes con Gl. dissecta. El
ejemplar tipo de la f, latilobata, que me fue facilitado por el
New York Botanical Garden, se diferencia sin embargo de Gl. dis-
secta tipica por las lacinias foliares m4s obtusas, la inflores-
Cencia menos ramificada y 1a pubescencia m4s rala. Considero Lae
esta diferencias pueden aceptarse como variaciones normales den-
tro de la especie." She cites from Brasil Tweedie s.n. (ier
Hooker] (K), from Uruguay Herter 96556 (Si), Fruchard 8.n,

Brava, Soe, Ls Commerson s.n. [Montevideo V-1767] (: te) -
Berro N), and from Buenos Aires, Argentina, Nicora a (33),
beige 259 oe (Si), Martinez Crovetto 43 (Si), and Boelcke 6342

Si SOR

Argen-
tine species of this complex rag Troncoso (196), quoted above, the
Specimens cited by me under V. dissecta in previous installments
of these notes should be re-examined. The New Yerk Botanical Gar-
den specimens of Rodrigo 2339 and Herter 96556 — collections

dissecta, seems to be better placed as V. microphylla H.B.K.; _
verbs Inst. Miguel Lillo 31745, Herb. Osten 10676, and Jorgensen
1613 & a.n. 8.n. [26.III.917) are V. parodii (Covas & eek senpen®) Molden-
ke; Villafatie 52l, is V, pogsetons Klotssch; Boffa 330, T. Meyer
12290, and Rambo 1891], are V V. pulchella Sweet; Chicchi Be a
part, ‘o Aragone Bergalli Campal, & Rosengurtt 1133
1821, Herb. Herter 60852, Herbs Jussieu Suns, sod Rosengurtt B
ot are hiner’ selloi Spreng.; Rambo 10103 and Rosengurtt PE.5262 PE 5260 are
ana (Troncoso) Shinners; and Herb. Inst. el Lillo
$a se son, [Patagon. 1882/4], Schreiter 190k, Tweedie s.n.,
and Wilkes AYE are V. ten enera Spreng.

The Ven Ventart 10301 I, originally distributed as V. dissecta is V.
Perakii (Covas & Schnack) Moldenke; Legrand 2717 is V. selloi
Spreng.; and Brade 7000, Gallinal, Aragone, Bergalli, Campal, &
oe PE.5262, Herb. Com, Googe & Geol, 3685, Herb. Inst.

cenit emer etcetera
Se nema oman ermtin

It is very mbste seaae that most af the Uruguayan material which
e

Seems to me to be far rTaasap ig poapenllcy peter aa
Shella and so I contime to regard it as V. pulchella var. lati-

19h PHYTOLOGIA Vol. 13, no. 3

lobata Moldenke.

~~ The Frenguelli 105 105 & 137, cited below, were erroneously cited
by me previously as sv. pule chella, while the Boffa 25 and Commerson
8.n. were previously - cited by me as V. tenera.

Additional citations: BOLIVIA: Province undetermined: Kuntze
s.n, [Macbride photos 172h] (W—photo). URUGUAY: Coumerson s.n.
| ceescmpodl photos 39502, in part] (W—photo). CHILE: Province un-

: Née s.n. (Herb. Willdenow 11139; Macbride photos
a7h60) Ow (W—photo ) of type). ARGENTINA: Buehos Aires: Boffa 25 (N);
Commerson s.n. [Buenos Ayres et Monte Video] (N), s.n. . [Macbride
photos photos 39502, in part] (W—-photo). Entre Rfos: Cabrera 10792 (W—
2370642). Santiago del Estero: Frenguelli 105 (N N), 137 (N).

VERBENA DISSECTA f. ALBA Moldenke
Additional biblio Fie ef Bares Moldenke, Phytologia 9: 30 1963.
The Herter 1805 [Herb. Herter 965861, cited by me in previous
installments of this work as V. dissecta f. alba, seems definite-
ly to represent typical V. pulchella Sweet instead.

VERBENA DUSENII Moldenke

Additional bibliography: Moldenke, Phytologia 10: het A 196).

” vor citations: BRAZIL: Paranf: Saito & Pereira 16 9/66
C)e

IANA Schau.
Additional bibliography: H orks Pringl
49. 1936; Moldenke, Phytologia Ms is MSs & S02. 1965.
J. Rzedowski 5324 & & 8058 are said to have had "purple" flo-

‘3 and were collected alo along ponte: blooming in September
and Poe grrpaeed
citations: MEXICO: San Luis Potos{: J, Rzedowski
$32h (1 (Ip), 80 8058 (Ip).

RLRUNS H.B.K

Additional bibliography: s Pringle
— - 19365 Moldenke, Phrtalogia in: 8 & a (965) "and 13:

” ccmnins found this plant growing in Abies religiosa and
Pers ceppeans woods. The Paray 565 and J. Rsedowslt 710
& 10026, originally distributed as V. elegans, are actually V.
deiticcle Small.

Additional citations: MEXICO: Hidalgo: Gonz4lez Quintero 317
(Ip, Ip); Riviello s.n, [Marzo 196] (ip) s Ze Bae’ Je oe aeiceskt 17002
(tp) » 18192 (Ip). Wuevo Leén: Manning & Manning 53254 (ku

7. erdab et San Luis Potos{: J. Rzedowski “8130 (ip aa Ty ', Tamaulipas:
artin & Sarabia 1181 (Ip). Vera Crus: Népoles s.n. [Agosto

TET Ge ) Pas ees

1966 Moldenke, Notes on Verbena 195

ELEGANS var, ASPERATA Perry
Additional bibliography: Moldenke, Phytologia 11: 56 & 91.

The P aray 323, distributed as V. elegans var. asperata, is
actually V. delticola Small, while M Martin é Sarabia 1081 T and Je
Raedowski 6. 8130 are typical ve elegans ans H.B
~~ Additional | citations: MEXICO: Sepre Fuentes C, sen. [20.1X.
1959] (Ip). Federal District: La Lagiinez 55 (Ip). San Luis Potosi:
J. Raedowski 8817 (Ip), 9685 (Ip). Tamaulipas: R. M. King 1513
a a

(va

cuat Wet Moldenke
Additional bibliography: pcan. Pals egears li: 456-157 &
467. 1965; Moldenke, Résumé Suppl. 12 1965.
One of the A. R. "yo idenke 821 i fds cited by me previously

as in my personal h herbarium is now in the herbarium of Aarims Uni-
versity.

VERBENA EPHEDROIDES Cham
Additional bibliography: ee eho a ll: 457 & 481.
aoe Rambo, ee ot Bot. 21: 3 e [59].
he Eye erdam 1 Seishin Vat iiuaes as v. ephedroides,
is ae five scoparia (oil. & eer Moldenke.
Rambo (1965) cites Herb. Anchieta 3093h from Rio Grande do
sul, Brazil, collected by h by himself. He says of this plant "Habit:
etly the same as V, alata Cham.; I cannot see any reasonable
difference between the two 5: species." He gives its habitat as
"Densely vegetated swamps" in the "Northeastern Highlands (in
Company of V. alata ({)" and the general distribution as "From
Minas Gerais to Uruguay and Misiones.”
Additional citations: BRAZIL: Sfo Paulo: N. Santos 5743 [239
es (Ja). State undetermined: Sellow s.n. [Brasilia meridionali;
acbride photos 17h15] (W—photo of type).

xVERBENA FECUNDA Moldenk
octet tional Mb lipassniees Moldenke, Biol. Abstr, 33: 1215. Pe}
a » Excerpt. Bot, A.5S: kh. 1962; Moldenke, Phytologia 10: 106

denke
Additional bibliography: M » Biol. Abstr. 33: ag 19595
penne Excerpt. Bot, A.5: Me 1968 0 Moldenke, Phytologia 11: 57.

flats bt roads, in campos » low woods, low pastured woods, and on

196 PHYTOLOGIA Vol. 13, no. 3

rock outcrops in Santa Catarina. Rambo (1965) aries Mant «il Pe
Bras. 9: 192 as "1951" instead of mete and cites Herb.
eta 2526, 4825, 8955, 11550 [Buck sen}, 11579 Het vaste), 3)
36113, 51000 [Sehnem S131], 5U5TL, SW642, 50373, 59237, and 28 Boe
(Camargo 3118 3118] from Rio Grande do Sul, Brazil, wil, all coll
himself except where otherwise indicated. He es en the plant
as "Weak erect, herbaceous, up to 20 cm high", growing in "Yoist
places in grassy Campos", and notes that mall peters ps from
the northeastern Highlands” 0 of Rio Grande do Sul, a a general
rector "From Minas Gerais to RGS and Misione

citations: BRAZIL: Minas Gerais: 61 20 [Macbride
photos 17439) (W—-photo). Santa Catarina: Smith & Klein 13333
(Ac), 13390 (Ac), 13740 ai a 13870 (Ac); Sm Smith & Reitz tz 12182
(Ac), 12831 (Ac), 14370 (Ac

VERBENA GLANDULIFERA M
eine bibliography: snare Phytologia 11: 58. S90 2
tio citations: ARGENTINA: Santiago del Estero: Piero
"h" ve et 3/1¥/uh; Herd. Inst. yams Lillo 9960] (Mi).

GLUTINOSA Kuntze
Additional bibliography: sieaee Siveslogss 1: Pg 1965.6
Additional citations: ARGENTINA: Mendoza: Kuntze 2 (Macbride
photos 171,18] (W—photo. of ou

tame GONZALEZT CC é

tional bibliography: » Excerpt. Bot. A.6: 535.
19635 aldenke, Phytologia rf ag: "1965.

paki Uibliccre) 458—L5
re) nf Moldenke, P| logia 11: —— 59
igo, & lot (1965) and 33189. Sekccads oon
f this species has been misidentified and distribu-
ironies "V. ciliata var. ciliata". On the he other hand,
the R. L. a 820, 867, 869, & 8185, distributed as V.
, are actually V. Vv. delticola Small.
ey oeege citations: ARIZONA: Coconino Co.: Cureton 21 (Ac);
Deaver 5822 (Ac); K. Greenwell 12 (Ac); P. Vogt 21 (Ac). Mohave
ees Dearer £135, in part (Ac); Eley 37 (Ac). Yavapai Co.: Ge Me
errard 5 (Ac). MEXICO: Baja California: ns & Wi
12073 ti}. aj Wiggins & Wiggins

GOODDINGII var. NEPETIFOLIA Tidestr,
Additional bibliography: Moldenke, Phytologia 11: 59 & Lgl.
Straw & Forman found this growing o ny ee
grazed hillsides and washes in sandy-clay stig
scribe the corollas as "near deep purplish-pink, OS RP 6/0 aa
the Munsell system per Nickerson color fan".

1966 Moldenke, Notes on Verbena 197

Additional citations: ARIZONA: Coconino Co.: R. Pardy 32 (Ac).
Mohave Co.: Deaver 6135, in part (Ac). Pima Co.: Hess & Procter
son, (September 5, 1938] (Z). MEXICO: Chihuahua: Straw | & Forman
61h (MA).

XVERBENA GOODMANI Moldenke
Tes peagewe bibliography: Moldenke, Biol. Abstr. 33: os 19995
ie gk ae Bot. A.5: lh. 1962; Moldenke, Phytologi

9.19

VERBENA GRACILESCENS (Cham.) Herter
Srereiaeee ages tag rer wel ee 11: 459. 1965;
Rambo, Pesquis, Bot. 21: 38 & [60].
cite (1965) cites Herb, Anchieta “hes, 9176, Zee. 32310,
and 59154 [Pivetta 966] fram Rio Grande do Sul, and 54326 [Reita
5006] from Santa Catarina, Brazil — all collected by himself

« Paulo araguay to Pa a",
Additional citations: ARGENTINA: Pohl ors Se Lossen 466 (G).

GRACILIS Dest.
Additional bibliography: H. B. Davis, Life & Works Pringle 152.
19363 Moldenke, Phytologia 11: 59. 1965.
ould found this pastures along a moist

ant open
draw in Chihualma. The flowers are described as "purple" on J.
Ruedowsaki 3338, 20082, & 20219.
tional | ei tationnt MEXICO: Chihualma: F. W. 871
(Mi). Federal District: J. Rzedowski 20082 (Ip). eo Je
Reedowski 20219 (Ip). San Luis Potosf: J. | J. Rzedowski 3338 (Ip),
3BUB (Ip), 7998 7995 (Ip).

Addi ton logia 11: 459. 1965.
tional titdicer : Moldenke, sha he a ll: °

Recent collectors grapiys Me ‘ound this plan + growing at 2100 to 3450
poe altitude, flowering from February to April and in September,

and September
juan citations: CHILE: Tarapacd: Marticorena, Matthei, &
Quezada 2 (Ac); R. A. Philippi s.n, [Sibayaj Macbride photos
179] (W—photo); 1 3 Ricardi & Marticorena = (Ac); Ricardi,
Marticorena, & Matthed 8 (Ac), 174 (Ac), 307 (2)-

oF crc » 12: 1. 1965;
onal bibli Moldenke, 3 de i
ii eabs, onks, Phytologia ll, Vos so, MiB, SOL, & 302 (2968) an 13:

Recent collectors have found this species growing in loose
» Sandy or loose red silt, dark or black soil, red-brown or

198 PHYTOLOGIA Vol. 13, no. 3

black clay, red or brick-red loose sand, deep orange-colored
loose sand, light tan clay, very pale tan loam, hard sandy marl-

brush, and in wet places at water's e edge. Turner found it grow-
ing along with V. cloverae Moldenke in Frio County.
The R. Me King 294, originally distributed asV, halei, is ac-

tually Centrostachys aspera (L.) Standl. in the Amaranthaceae.
3

urticifolia", but I do not think that it is a hybrid. | On Parks,
, & Turner the flowers are described as "small yellow’
{adi tional al citations: MISSISSIPPI: Forrest Co.: Re Le crate
8671 (Ld). LOUISIANA: Cameron Par.: R. L. Crockett , 8398 (Ld).

TEXAS: Aransas Co.: D. S. Correll 28920 (Id). Bast Bastrop Co.: B.
R ez 72 * (an~-200864) Bexar Co.: Chavez, Flores, & Hein
2). Brooks Co.: Diana Rodriguez 0 mez 0 (Ku—233350); Vela Vela

(Au—23
12 (Au--233357). Callahan Co.: N.C. Henderson 63-197 (Au—
56). Comanche Co.: N. C. Henderson 63-600 (Au—227282) « Dal=
las Co.: 0, Sanders s.n. (Irving, July 1, 1940] (Id). DeWitt
Co.: Blanch 17 (Au--222252). Dimmit Co.: R. Acosta 81 (Au--
233359); Al Alvarez, Guajardo, Salazar, & McCart 7619 (Au—-222265) ;
Chavez, Flores, ¢ & Hein 76 (Au—233418) 3 Fowler & Vargara 112 (Au-
2326U8) 3 HE. es 76 (ku--235232). Duval Co.: Alvarez, Gua jar
do, Salazar, & McCart 7671 (Au—-22225)); Flores, Chavez, & Hein
13 (Au—233425); Gamboa & Dohnke 130 (Au—232668); R. Garza 72
‘(Au--222253); A. Trevino 5 (Au—222260, Ip). Fayette Co.: Co.: Ripple
a2 (Au--22688]). Frio Co.: Painter, Lucas, & Barkley lu2co 14226
Ld); B. L. Turner 990 (Au--229),09) . Gonzales Coe: M. de la Le
Hernandez 58 (Au--222250). Hardin Co.: R. L. Crockett 1157 (1d).
Hidalgo Co.: Painter & Barkl ey 14128 (Ld). Jim Wells Co.: es Pe
Cabrera 101 (Au--233347); Gamboa & Dohnke 1h0 (Au—233)21).
Jones Co.: N. C. Henderson 1 63-170 + i eee) . LaSalle Co.: Bo-
ate es (Au--222257); Wine wineh 2 Gs—222280) Lavaca Co.: one
(Ip). McMullen Co. : Alvarez, Guajardo,
en Teak (Au—222256); Minerva Martinez 70 (hu—233356) « * bine
Co.: Demaree 738 (Ac), “FOgOT (Ac). San Gry vedo ~ Le
Crockett 8226 (Ld, Ld). Starr Co.: M. de
233419) « Ran ie Co.: Ne ea Henderson | rr Puls FeF68) . val
Verde Co.: Parks, Warnock, & Turner 25) (Mi). Webb Co.: Cisneros
2h thn ose B, Gutie pes 37 Sager ROE soto, Herrera, &
McCart 8101 (au—222262). Zapata Co.: Guerra 0 (Au--232633) «
High = R. L. Crockett 8356 (Ld). Mustang Island: B. H. Wat
0999 (Ld). MEXICO: Chilmahua: Pringle s.n. (Valley, Ortds,
poe i, Ti, 1887] (mi). Tamaulipas: Carazos 50 (Au—235228) ; Domin-
M. & McCart 8183 (Ip), 82h9 (Au—222258); M. Leal 72 (Au~
222 OST: Ip)$ Pauli Paulita M Martinez z 60 (Au—233420) ; Reséndez 9 (Au

1966 Moldenke, Notes on Verbena 199
2222)8); Zertuche 36 (Au—222255) .

VERBENA HASSLERANA Briq.
Vepaniars bibliography: Moldenke, Biol. Abstr. 33: ans ; 019995
em ie Excerpt. Bot. A.5: kh. 1962} Moldenke, Phytolo
0. 1965

Additional citations: PARAGUAY: Hassler 646), [Macbride photos
2h685] (Mi—cotype, W—photo of cotype).

VERBENA HASSLERANA var eh Mol
Additional repped: Molde: oa 1. Abétr. 33: a "i
Excerpt. Bot. A.5: hh. 7963; Moldenke, Phytolo;
178-179. 1963.

VERBENA HASTATA L.

Enended synonymy: Verbena lanceolata Boldo ex Moldenke, Alph.
List Invalid Names Suppl. 1: 25, in syn. 197 [not V. lanceolata
Beck, 1828, nor Willd., 1825).

Additional bibliography: Deitschman & Pruett, Proc. 17th N.
Cent. Weed Control Conf. 1960; Deitschman & Pruett , Weed Abstr.
11 (3): 211. 1962; Rickett, New Field Book Am. Wild Fls. 280, pl.
ees Moldenke, Biol. Abstr. 5: 3136. 1964; Moldenke, Phyto-

ce

denke, Biol, Abstr. 6: 3616. 1965; Moldenke, Résuné Suppl. 12: 1.
1965; L. M. Reid, Park Management Serv. Bull. 5: 5 & 8. n.d.

: ted a illustrations: Rickett, New Field Book Am, Wild Fls.
P. 1963.

Deitschman & Pruett (1960) report eradicating this plant by
use of 2,)D and cultivating between rows of Pims, Juglans, and
Popalus,” MacKeever reports this his no. MV.400 had open
Spikes with dark-blue flowers, while MV.l402a, found at the edge of
@ brackish marsh, had the flower-spikes dense at the apex and the

corollas

purple
The M. E. Jones 1187, distributed originally as V. hastata, is
actually var. ‘scabra Moldenke, while Heidenreich 298 > (71 7130) rs xVe
r Mol

Additional citations: MAINE: Franklin Co.: A. R. Moldenke 1125
(Ac). Somerset Co.: A. Re oe ii 112k (Ac). NEW HAMPSHIREs
Carroll Co.: A, R. Moldenke 1127 (Ac). Merrimack Co.: A. R. Mol-
Senke 1129 (Ac). VERMONT: Bennington Co.: A. R. Moldenke 1135
(Ac). “Chittenden co.: B, Be ee 1138 (Ac). Essex Co.: As R.
wokdenke 113k (Ac). Franklin Co.: A. R. Moldenke 1080 (Ac).

moille Co.: A. R. Moldenke 1077 (Ac), 1239 (Ac). Orleans oe Ae

R. Moldenke 1136 (Ac). Rutland Co.: A. R. Moldenke 1137 (Ac).

Washington Con A. R. Moldenke 1076 (Ac). Windsor Co.: A. Re ag

ioe ty hoe ay warns 4s ; H. N +s Msckoaver sae

N). 4 to Oet He Ne
Ue). toapeins Sep yea tionision g [Slaterville, 7/8/39] (Ip).

200 PHETTOLOGIA Vol. 13, no. 3

Washington Co.: A. R. Moldenke 1071 (Ac). ‘INDIANA: Porter Co.:

M. W. lyon Jr. s.n. [Mineral Springs, VIII.3.192h] (Mi). KENTUCKY:

Bath Co,: Plymale s.n. [July 16, 1939] (Mi). Powell Co.: F. T.

wrens Y10 (Mi). MISSOURI: Saint Louis: Muhlenbachs 1035 (Ac),
307 (Ac), 2402 (Ac). COLORADO: Las Animas Co.: C. M. Rogers

To8l (a).

HASTATA f. CAERULEA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 465. 1965;
Moldenke a Biol. Abstr. 6: 3616.
It is very possible that MacKeever “yv.1,00, cited previously
under V. hastata, represents this form since the collector notes
that its flowers were "dark-blue" rather than purple.

‘A HASTAT. te f. ROSEA Cheney
ditional bibliography: Moldenke, Phytologia 11: )65--l)66.
19655 Moldenke, Résumé Suppl. 12: 1. 1965.

VERBENA HASTATA var. SCABRA Moldenke
si heathen bibliography: Moldenke, Phytologia 11: 62 & 1,66.

Additional citations: UTAH: Utah Co.: M. E. Jones 1487 (Mi).
TEXAS: Hemphill Co.: Correll & Correll 3003h (Ld).

VERBENA HAYEKII Moldenk

— bil cp peemer ie Moldenke, Phytologia 11: 66. 1965.

dditional citations: PERU: Junin: Weberbauer 2573 [Macbride
piers 17442] (W--photo of type). rE Pa

» growing on "Dry, grassy C
ad seg catta in that Rey with a general distri-
seetae = "RCS and
eo onal BP ah (hog BRAZIL: Santa Catarina: Smith & Reits

barsarrey pm Spreng. 965
onal bibliography: M li: 466. 19653
Rambo, Pesquis. Bot. 21: 3639, Sh, ahs ‘& Teo]. 2 6s

Rambo (1965) cites Herb 661h, soise [Reits 10h}
32221, 36421, 45518 SEL: ShgpIe e t $6783 rem Rio Grande do

and 36896 [Hatechbach ” Soo] from » Brazil - all col-

pa od trike himself except where arr indicated. He admite,
however, that no. 36421 has been identified as var. gracilis

ee, = An pieneueereen

—-

a aa a

1966 Moldenke, Notes on Verbena 201

Dusén, He describes the species as a "Small shrub, up to 30 om
high, more or less hairy » growing "In more or less oe be
Campos" in the "Northeastern Highlands only" in Rio Grande do Sul,
with a general distribution "From Bahia to RGS and Misiones",
Additional citations: BRAZIL: Minas Gerais: N. Santos 570k
(Ja). Parand: Dombrowski, Saito, & Pereira 808/598/192 (Ac);
Reiss l6a (Mi); Smith & Klein 11936 (Ac); Smith, Klein, & Hatsch-
bach 1562 (W-~-21,51599). Santa Catarina: Snith. & Klein 1302,
(ae), 1373 13733 (W-—2151607), 13885 (Ac). Sto Paulo: Leite 3373 (G).
CULTIVATED: * Germany: Otto s.n. [Macbride photos 1720] (W—photo).

VERBENA HIRTA var. GRAC én
Additional bibisenrephes Moldenke, Phytologia 10: 110. 196);
Rambo, Pesquis., Bot. 21: 39. 1965.

Sud th and his associates Dag sigismey se this plant in gallery
forests and bogs, as well as on dry sunny banks, flowering fruit-
ing in December, The altitude given on the label of one of the
Smith & Klein ee tions is "12000 m.", obviously a typographic

Additional matttlincs BRAZIL: Paran&: Dombrowski 1252 [Herb.
Inst.Def. Pat, iene 9446] (Ac), 1356 [Herb. Inst. Def. Pat. Nat.
Hares (Ac); Dombrowski, Saito, & Pereira 91:5/770/289 (Ac). San-

: Smith & Klein in 1339) (W—251609), 1355 (Ac).

VERBENA HISPIDA Rufz & Pav
soit tional bittdogranhy: Moldenke, Phytologia 11: 66 & 8h.

Additional citations: PERU: Hudmuco: H. Rufz 1/3 [Macbride
Photos 29686] (W—photo of type), sn. [Peruvia et Chili; Mac-

8.n. [Termas » 12-I-1950] (Ac).
VERBENA HOOKERIANA (Covas & Sehuack) Moldenke
Additional bibliography: M » Phytologia 11: tg rigs °

The Chicchi 264 Pie Poa = a pope por ure of this spec
as selloi Spreng., not V. dissecta as stated by me mateo
(1 iad tonal citations: ARGENTINA: La Rioja: G. Dawson 3363

VERBENA HUMIFUSA Chan,
Additional biblio. graphy: Moldenke picoere li: 466. 1965;
Rambo, Pesquis. Bot. 21: 39, 58, & (60).
(1965) cites Herb, 1, 90633, sere {Henz s.n.],
43261, ioe, jeeth, Bol, il, 1a, 61272 [Emrich s.n.],
ande do Brazil — collected by

1 roo to 4O cm
rot ", Growing in "Dry, grassy Campos". He notes "All collections
the northeastern Lowlands and sea coast in that state,

202 PHYTOLOGIA Vol. 13, no. 3

gives the general distribution as "From Paraguay and Paran& to
RGS and Uru

Additional citations: BRAZIL: State undetermined: Sellow s.n,
(Campos de Brasil meridional; Macbride photos 17,21] (W-—photo of
cot;

XVERBENA HYBRIDA Voss
Additional synonymy: Verbena hybrida L. apud Rambo, Pesquis.
halm. 1

sp

Additional bibliography: Brilmayer, All About Vines pir ds 1962;

A. ee Weaver, Carib, Gard. 18, 76, 129, & 138. 196k; Al & Ditt-

Biol. Data Book 9h. 196; Moldenke, Biol. Abstr. Bert “s37t.

196h Woldenke, Phytologia 11: 1156 & l66—L67. 1965; Rambo, Pes-
quis. Konink, Rotte

, Ann ai
Montréal hs te 19655" Florists! Rev. April 15. 1965; Neal,
In Gard, Hawaii, new rev. ed., 722. 1965; Moldenke, Phytologia 13:
180. 1966.

Altman & Letom oats report that this plant is attacked by
the chrysanthemum tode (Aphelenchoides ritzema-bosi) in both
North America and ane This nematode attacks about thirty
rbssae: kinds of of Rais, including chrysanthemum, larkspur, phlox,

rry, and ginnia. It is a vagrant = sa page of the buds
nied foliage. Symptoms are crumpled distorted leaves eaf-
spots. Control is effected by the hotwater treatment ae the dor-
mant plants and by parathion sprays.

r (1962) says: *hortensis — verbena hybrids -- The
well-known atin verbenas, with trailing stems, soft hairy leaves,
and clusters of vivid flowers sometimes with sparkling white eye.
Seeds are usually listed as 'trailing v rari! in contrast with
dwarf or bushy types, or as hybrida eigen — i a selection of

red te nk almon shad or mixture
Raymond (1965) lists "Candy Stripe", "Dark. fae", spr
Blue", and "Mid-Blue" as Dwa e-pink

many v: varieties offered to the trade Buckley & Son and
Schlosser & Sons [both of Springfield, Illinois], Wamego Floral
Company [of Wamego, Kansas], ehm Greenhouses [of Huntington,

In « lettes to me from Jerome H, Kantor, Director of Research
for tne W. Atlee Burpee Company, Philadelphia, Pennsylvania, he
says: "We are glad to know of your interest in verbenas are
asking Mr. Ellwood Pickering, HP is doing our flower research 4b

1966 Moldenke, Notes on Verbena 203

true breeding strain of this color by inbreeding, The ewe is

a heterozygous form, the dominat being purple and the recessive
ng white. Theoretically, it should be possible t

good lavender by making , hybrid, but so far we 6 been un-

fact, a lavender mixture can be made by planting these two, Laven-
der Glory and Floradale Beauty, as it includes practically all of
the verbena colors with the exception of blue."

Mr. Pickering, in his letter to me of March 11, 1965, says:
"There is little that I can add to what Mr. Kantor has told you..
about Miss Susie Verbena. It was as he said a mutant in a trial
row of Floradale Beauty. I might add that Miss Susie has a pur

Bea i

of n the same color. The extra petal ssive
when crossed wi egular type. Only one or two will appear
in a population of sever. F/2 plants. It is also inter-

colors in this type on the mar
1.196h] Gane citations: eSpetoer ig poe Madrigal s.n. (11.

XVERBENA ILLICITA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 67. 1965.
One each of the A. R. Moldenke 833 & 975 collections cited

ty me previously as in my personal herbarium are now in the her-
barium of Aarhus University.
VERBENA I

Briq.
Additional bibliograp’ Moldenke, Phytologia 10: 111. 196)
pres Pesquis, Bot. 21 iat 1965; Moldenke, Phytologia 13: 18h.

1
Rambo (1965) cites Herb. Anchieta 41219, collected by himself
in Rio Grande do sul, Brasil, * as having been identified as this

Species, but comments tha t "V. inamoena,.....is at best a variety

of V. bonariensis L." and so he cites it under the latter species.

Addi tional bibl neha EON
onal biblicera Moldenke, tt anee a ll:
a Anon., Ind. Sem. oo h, Konink. Dierg. (6). 1965 ;
isidenke, Résumé Suppl. 12: 5. 19653 Sarees nig et Bot. 21: 58,

1965,
Additional citations: BRAZIL: Rio de Janeiro: Strang 580 [Herb.
Cent. Pesq. Florest. 3336] (Ac).

XVERBENA INTERCEDENS Briq.

20h, PHYTOLOGIA Vol. 13, no. 3

Additional bibliography: Moldenke, Phytologia 11: 68, 1965.
Additional citations: PARAGUAY: Hassler 6149 [Macbride photos
1722] (W--photo of cotype).

VERBENA INTERMEDIA Gill. & Hook.
Additional bibliography: Moldenke, Phytologia 11: 68. 1965;

ae & Reitz pn t ruderal and in dry woods, at
300 meters a
et 3 (1965) ee Herb. Anchieta 1,386 & 34723 from Rio Grane
do Sul, Brazil, collected b; ‘himself, “but comments

He describes the plant as "Erect, avitr, up to SO cm high", grow-
ay. in "More or less wet, grassy Campos" in the "Northeastern
" of that state, with a general distribution of "RGS,
n

entina a",
Additional citations: BRAZIL: Santa Catarina: Smith &R Reitz

1h330 (Ac). ARGENTINA: Mendoza: Wilczek 56 [Macbride photos
214,683] (W--photo) .

VERBENA JORDANENSIS Moldenke
Additional bibliography: Moldenke, Phytologia 9: 385—386.
-

Smith & Reitz found this species ruderal in dry open flats by
roads, at 700--900 wnt altitude, Ab ectoeoiy in October.
dditional citatiions: BRAZIL: Santa Catarina: Smith & ‘Reita
1279 (Ac). S&o panaee ‘Leite 37 ddabteee) ‘

VERBENA LACINIATA (L.) Bri
Additional bibliogra : Yoldenke, Biol. past. 35: 1688. an
Hi 3 Tronc arwiniana

225 & Rambo, toeatis Bot.
to the recent fine pageorene pel “work on some of the
gentine members of this species complex done by Troncoso (1965),

it appears that the Herb, Herter Tt, pee, & 96600, Herter
18lce & 181i, and Herter & Strahl i161, riginally distributed 4s
ve. V. erinoides Lam. and cited by me Faeekacmey as V. tenera Sprenés
are actually V. selloi Spreng., while the Herb. Inst. %. Miguel
Lillo 32759, Schreiter 190k, Tweedie gen., and Wilkes ee Sy
also originally so y so distributed but cited | by me as V. ecta
Willd., are actually V. tenera Spreng.

rpghtiitional citations: ECUADOR: Chimborazo: Fagerlind & Wibos

VERBENA LASIOSTACHYS Link 7006+
dditional bibliography: Moldenke on ey Abstr. 45: 5377 &
1964; Moldenke, Phytologia 11: 69.

LILACINA Greene .
Additional bibliography: Moldenke, Phytologia 11: 169. 1965

many

1966 Moldenke, Notes on Verbena 205

Moldenke, Résumé Suppl. 12: 12. 1965.
Additd onal citations: CEDROS ISLAND: Haines & Hale s.n. [9
March 1939] (Mi); Re. Moran 10698 (Mi).

VERBENA LINDMANII Briq.
Rey iikiindeaes Moldenke, Pees li: 469. 1965;
Pesquis. Bot, 21: hl—2 & [60

lm 8 Oe ee ee Fee eee oe

31y 3u722,
37777, 39352, 39h, 39621, 39697, 10398, 11059, 51h13, 51557,
53037, 57071, 59160 [Pivetta 972], ‘© 6158) [Camargo 2683) from

bach 1181] from Paranf, Brazil — all collected by himself ex-
cept where otherwise init: He describes the plant as eben
» more or less leafless, a few branches from a woody ro
stock, up to 75 cm cm high", growing on "Very Budd open Cisse <5
Apparently over the whole of the Cam ampos area” in Rio Grande do
2 with a general distribution "Fron Parnd to RGS and Misio-
s

erin LIPOZYGIOIDES W
Additional Ribliograni Mold Biol. Abstr. 45: 7026.
1964; Moldenke, ib aie gat a 93 ryy & 501—502. 196h.
Additional c HILE: vince undetermined: Besser
136 [Macbride Abra nen) (W--photo of logotype).

LITORALIS H.B.K.

Enended Synonymy: Verbena lanceolata Willd. ex Spreng. in L.,
Syst. Veg., ed. 16, 2: 78. 1825 [not V. lanceolata Beck, 1828,
nor Boldo, 1947]. Verbena madiflora Nutt. ex Turez., Bull. fino.
Nat. Mosc. £5 (2): 195-196. 1863 [not V. mudiflora L., 1808].

Additional & emended bibliography: N. J. Anderss., Galap.
Veg. 200, "185h5 N N. J. Anderss., Vet. Akad. Handl. Stockh. 1853:
a 1855; Turez., Bull. Soc. Nat. Mose. 36 (2): 195-196. 1863;

e 132

- B. Davis, Life & Works Sen pe - 1936; Hocking, Excerpt.
Bot. Aw: 592, 19623 Soukup, Biota 5: 19). 196h; a | Biol.
Abstr. 15: 6658, 7026, & 8025. ds. 196hy Neal, in feed, new
rev. ed., 721—— 722. 1 1965; & [60].

1965; Moldenke, Piytalogia 11 li: preg rs (e970, ‘k “son “G368)s 12:
205 (1965), and 13: 188,

Recent collectors rie ite this plant in much dis-

woods of Quercus, Liquidambar, and Clethra and abundant

on ©xposed lava on Quaternary continental deposits. Raven &
Teeealove found it growing on slopes with Quercus, Pimus, and
Liquidambar, at 5100 feet altitude. They describe the flowers
ot neir no. 19907 as "pale-blue"; Tucker 290 had a
flowers, while his his 1308 had "corolla-throat pale-
white, lobes lavender"; Ricardi & Marticorena OAT bs ? bad hive!

206 PHYTOLOGIA Vol. 13, no. 3

flowers. A common name for the plant in British Honduras is

"verbena de montana", while Neal (1965) records "ha‘uowi", "ha‘uoi}
"oi", and "owi" from H Hawaii and Israel records "puringa" from

Easter Island.

The aang 6417, originally distributed as V. litoralis, is

actually V. y V. brasiliensis Vell. Andersson ays cites a C. Darwin

s.n. from Chatham Island in the Galapagos. o (1965) cites

cee eres me enema” eee ce eee ee we nee Seperate

Prastnatnctn 3669 1 TFri eer mem te ee sen emer ial

ies , oe ; 1:8067, 14,8657, 19506, 19723, mes ere SeNely
5055h (Sennen LL70), 51bL9, 53120, 55075 [Camargo 90), 59429 [Ca~
margo 959], 6006 Raa ar ee 61623 [Camargo 2727],
62112 [Camargo 2014], 6 2500 [Camargo rgo 21h5], 6 6325 63255 [S [Spies sen],
63272 [Sacco 827], 63275 [s Sacco 855] , 057 TSacco 259) from rom Rio
Grande do Sul, 53631 from Paran4, and 3116, 60152, 61117 [Mat-
tos s.n.J, & élik2 Dh [Mattos s.n.] from Santa. Catarina, Brazil —
all collected by himself e except where otherwise indicated. He
describes the plant as "Herbaceous, more or less lignified at
the base, moderately branched, up to 1 m high.....Apparently not

the cultivated area" in Rio Grande do Sul, with a general distri-
er "From soutlinens USA to Uruguay, central Argentina, Chile

Fernandes

plantdeuat se MEXICO: Chiapas: Raven & Breedlove
19907 (Ac). México: Lagfinez 87 (Ip). ico fe Hinton & al.
13956 (Mi). San Luis Potosi: R. M. King 432 (Mi); J. Rzedowski
8137 (Ip). Vera Cruz: J. Rzedowski 18943 (Ip); Scora a & Scora
2452 (Ip). GUATEMALA: El Petén: Contreras 2635 6 (Ss). BRITISH
HONDURAS: Gentle 7119 (S$). EL SALVADOR: Chalatenango: Tucker 990
(Mi). Santa Ana: Tucker 1308 (Mi). BRAZIL: Rio de Janeiro: eiro: Cas
tellanos 2h51) [Herb. Cent. Pesq. Florest. 3206] (Ac), 2h562
[Hers. Cent. Pesq. Florest. 3177] (Ac). CHILE: Atacama: 3 Ricardt
& Marticorena 3629/17 (Ac). Colchagua: Ricardi s.n. [Cerros d¢
Nancagua, li-I-1951] (Ac). Tarapacé: Ricardi cardi 3352 (Ac). EASTER
ISLAND: Israel s.n, [24.V.1956] (Ac).

bees orice se Sbene rentaggiene Moldenke 16
fe) ography: Moldenke, P logia 10: 75--70-
1964; Soukup, Biota 5: 19h. 196. emer

VERBENA LOBATA Vell. 9653
Additional bibliography: Moldenke, Phytologia 11: 470. 19075
Rambo, Pesquis. Bot. 21: ee & [60]. 1965.
Smith & Reitz found this plant growing on cliffs in Santa C@

1966 Moldenke, Notes on Verbena 207

tarina,
or Bea (19685 cites Herb, Anchieta 2293, 2782, 2816, 1295, 479,
4482, 4505, 8663, 8881, B96l,, 29963, 29983, 30977, 32116, 32118, 32 32302,
32303, Bie, 2 32939 9 [Friderichs Seno), 34726, 35193, 35725 (Hen: [Hens
$.N. 8.n.], 36418 387L2, 4.0268, 4,832, 1988, LS 8, Le 45640, Ls 45682, _
Li72h a 3958), L808, 18588 (Sehnem Sene}, 50185 [Emrich Se
Be}, 51130, 5105, bih6e, ¢ 51912, 519]9, 5208h, 52017 [Burich 5.D.],

22h0e, 24712, 519K9, 52084, 52817 [Emrich s.n.

margo "20971, "62709" icanan a 2332) : fee Rio Grande do | Sul, » 31861

pence 222 225), 49570, 49605, 50968 [Sehnem 5449], 60070, & 61145

ape from Santa Catarina, and and 35960 [Hatschbach Lo 403] &

saa [hiatschbach 4306] from Paran4, Brazil -- all collected by
himself except wh xcept where otherwise teghiniad: He describes the

plant as "Ascending to erect and leahing to the neighbors, up to
€ e

"Northeastern quadrant only, very common on the Highlands" in Rio
Grande do Sul, wish a general distribution "From Minas Gerais to
RGS and Misiones."

Additional citations: BRAZIL: Parané: Saito & Pereira 506/103
(Ac). Santa Catarina: Smith & Reitz 11178 (W—2)51602). State
poets Sellow s.n. (Brasilia; Macbride photos 34348] (W—
Pp

LOBATA var. GLABRATA Moldenke
Additional pete : Moldenke, Phytologia 10: 82. 196h;

» Pesquis. ve
Rambo (1965) pei Herb. “Anchieta 2816, collected by himself
in Rio Grande do Sul, Brazil, as Ve eo ta Vell., but admits
that it has been identified as var. ta.

LONGIFOLIA Mart. & Gal.
tional bibliography: Turez., Bull. Soc. Nat. Mosc. a ay

With white corollas, growing in semi-shade on rocky clay loam, and
ae ne in June. ‘Rzedowski encountered it at 100 meters alti-
tude lowering and fruiting in February.

Addi tional citations: MEXICO: Guerrero: Floyed & Ryan 8, (Mi).
San Luis Potosf: J. Rzedowski 975) (Ip).

VERBENA LONGIFOLIA f. ALBIFLORA Moldenke
Addi tional aestyoresren Moldenke, Biol. Abstr. 36: 036. 1961;
veking, Excerpt. Bot. A.5: lh. 1962; Moldenke, Phytologia 10: 85.

LUCANENSIS Moldenke
ddi tional bibliography: Soukup, Biota 5: 19. 1964; Moldenke,
Plytologis 10: 85—~86. 196).

208 PHYTOLOGIA Vol. 13, no. 3

VERBENA MACDOUGALII Heller

Additional bibliography: Moldenke, Biol. Abstr. 5: 6658
8025. 196; Moldenke, Phytologia 11: h62 & 470—l71. 1965; stab
ésumé S

ui? (Ae) «

MALMII Moldenke
Additional bibliography: Moldenke, Phytologia 10: 121--122.
1964; Rambo, Pesquis. Bot. 21: 58. 1965.

VERBENA MARITIMA Small
Additional bibliography: Moldenke, Phytologia 11: 471. 1965.
Demaree found this plant on sandy c
Additional citations: FLORIDA: Brevard Co.: Demaree 501708
(Ac). Indian River Co.: Demaree 922 (Ac).

VERBENA MARRUBIOIDES Cham.

Additional om genie a a 11: 71. 19653
Rambo, Pesquis. Bot. 21: 6 a is

The Reiss l6a, originally di hake cy as V. marrubioides, is

reng.
Rambo ise 65) cites Herb. Anchieta 51, 27065, 27293, 29028,

29105, 30629, 37698, 43689, L363, Wuls6, 18879, 57298, & & 62095
[Camargo 1636] from Rio Grande do Sul, and 31723, 31902 [Reits
1281), & 21906 [Reitz 1297] from Santa Catarina, Brazil — all
collected by himself except where otherwise indicated. He de-
scribes the plant as "Prostrate, branches radiating from a woody

the northeastern Lowlands and sea coast" in that ete and gives
the general distribution of the species "From Paran4 to Uruguay}

Additional citations: ar, Paran4: Dombrowski 683 (Z)3 Dom
browski, Saito, & Pereira 815/605/199 (Ac). State undet ermined:
Sellow s.n, ay (nent merids merid.; Macbride photos 17429] (W--photo of
type) .

ENA CA Spr 2.
Additional bibliography: Rambo, Pesquis. Bot. 21: 6—hi7-
1965; Moldenke, Phytologia 11: Whe 1965.
Rambo (1965$ places V. Biog! Dore Cham. in the synonymy of
this species on page 6 of his work, but keeps it as distinct 0”
49-50. He cites for V. megapotamica Herb. Anchieta 133,

4561, 8513, 8781, 8999, 9388, 9767, 11535 [Buck Sn], 30980,
34729, ae, 318, Slot, $6659, S5127 (Pivetta 97], & 592hh
from Ri , and 60153 from Santa ta Catarina, Brazil —
all eotlactat 5 by thdesls except where otherwise indicated. He de-
scribes the plant as "Herbaceous, vigorous, erect or recumbent,

1966 Moldenke, Notes on Verbena 209

with large, and showy flowers in dense in s", growing

in "Wet to swampy ee an the Campos pers of Rio Grande do

par specifically "Northern ¢ especially in the Northeast",
and

ampos,
th a eenee) Sieinibattos "From Minas Gerais to Uruguay
pl Aires.
Additdonns citations: BRAZIL: Rio Grande do Sul: a A
pera phaser 17438, in part] (W--photo of type). te unde-
3 8.n, {Brasil meridionali; Macbride ee 17438,
by part] ee

VERBENA MENDOCINA R
Additional abiteureae: Koldenke, Phytologia li: Se 1965.
Additional citations: ARGENTINA: Mendoza : R. A. Philippi s.n.
(1868; Macbride photos 174,30] cecatae of " isotype)

ope grer ery Bent! oe
tional bibliography: 2, Bull. Soc. Nat. Mosc. 3% (2):
ie 163} Figg hata ia rtelonia. ll: 471—l72 & 502, 1965.
& Edmo

eS ae Recent collectors have found it growing in secondary

or xerophi lous matorral, ruderal, and in cultivated soil.
Additional citations: MEXICO: Federal District: S. Espinosa s.

n. [Ciudad Universitaria, Ree and (Ip). Hidalgo: I. M. C. F

Sen. [h.VIIT.1963] (Ip); L. G. Quintero s.n. (71x63) (a “Tee

ico: Galicia s oeBe _(22.virr 1952) (Ip); Lachica s.n. [23.V1.1963]

(Ip); Madri 5 (Ip); Paray 2956 (Ip); Villamar C. s.n. [6.
ae aa ‘ ee Powell & Edmondson 91) (Mi). Vera Cruz:

“ate
Additional ibliogra Biota e lh. eng § Moldenke,
Prytologia 11 aus ire, » ll, & b ABs. (1 (2965) and 88. 1965.
T o the r sionary na =r by Troncoso
(2965) on penaecs aon Ase: came of this species complex, it

6i3, originally distributed as V. microphylla and cited by me
Previously as V. tenera, are actually V. selloi Spreng., while the

Carette s.n. [Monte Hermoso, 19161, Was a me as V. microphylla,
is actually V. tenera Spreng. The O'Donell 1600, cited below, was

Previously erro: rroneously cited stg me as V, dissecta Willd., while A.
zs Hunziker 671 and T. Meyer 209 were previously cited by me as

Vi tues Spreng.

The pce on T, Moree, 2408 2409 are described as "purple", while

Hunzike were "violet", Hunziker found the
species “Biatione alow ‘ada in Cérdoba.
Additional citati ARGENTINA: Buenos Aires: T. Meyer 2,09

6 Cérdoba: A. 7. ‘ienativee 6471 (N). Tucumdn: O'Donel1 1600

210 PHYTOLOGIA Vol. 13, no. 3

VERBENA MINUTIFLORA Briq.

eorge ie bibliography: Moldenke, Biol. Abstr. 35: 1688. 1960;

Excerpt. Bot. A: 592. 1962; Angely, Fl. Paran. 32: 69.

196 3 Moldenke, Phytologia 11: 472. 1965; Moldenke, Résumé Suppl.
Let: Se

Leite 37h is said to have had "violet" flowers, blooming in
November. Smith & Klein found the species growing in bogs and
ruderal rivers.

Additional citations: BRAZIL: Parand4: Saito & Pereira wef
(Ac). Santa Catarina: Smith & Klein 13169 (Ac). Sao Paul
Leite 37h) (Ac).
XVERBENA MOECHINA Moldenke

Additional synonymy: Verbena lanceolata Beck, Am. Journ. Sci.
& Arts 14: 118. 1828 [not V. la V. lanceolata Boldo, 197, nor wis

Weeks
ditional bibliography: Beck, . Sci. & Arts 1):
1828; j Holden, Phytologia 11: 472. ni96e; i iipichlly spain Me
1

e V. lanceolata of Boldo is V. hastata L., while that of
Willdenow is V. litoralis H.B.K.

I have not seen the actual type of Beck's Verbena lanceolata,
but from his deseription and _ locality of collection it wan

Stems simple, two or three feet high, sti erect, hairy.

thr nches long, about an inch in breadth, ed to-
omewhat acute, coriaceous, much attermated at base, sub

sessile. Spike simple, nal, dense flowered, imbricate.

F » blue. Bracts narrow-lanceolate , acute, longer

than the calyx. Resembles V. Stricta in meee The name, 4l~

though definitely proposed as a new one and accompanied by 4
— description, has been ignored by the tier’ of the Index

MONTEVIDENSIS Spreng
Additional bibliography: Hocking, Excerpt. Bot. Ae: 592+ 19625
Reitz & Klein, Sellowia 16: 73 & 117. 196k; Moldenke, Biol. Abst
45: 6658 (1964) and 46: 32h6. 1965; Moldenke, Phytologia 11: 473.
1965 Zi: hO—b1 & [60

woods in S. Bra:
cites Herb. anchista ys, 1129, ‘Si, , 518; pai 1020) ee

52083 ye, » tel, i, li, igs, pa “Sa791 [camarge
» 24995, 9, 56655, 59873 (Camargo 1192], 62791

2480), ats (Camargo 3125], & 63416 [Sacco 1006] fron Rio Grane
do Sul, and $1,308 [Reitz l L971], 54372 [Reite 5092], 58407 [Reits

|

1966 Moldenke, Notes on Verbena 211

8374), 60193, 6110 [Mattos s.n.], oie} [Mattos s.n.J], & 64283
[Mattos Sen.) from Santa Ca atarina, Brazil — all collected by h by him-
self ex except | where otherwise pees He describes the plant as

Gerais to RGS and northeastern Ar °

Additional citations: BRAZIL: Parand: Dombroski 103) [Saito
851; Herb. Inst. Def. Pat. Nac. 93] (Ac); Dombro Dombroski & & Saito
973/795 (Ac); Lourdes Pereira 96 [Herb. Inst, Def. Pat. Nac. 945]
(Ac); Reiss ve a) 3 (Mi); Saito 69 [Herb. Inst. Def. Pat. Nac. 9h2]
Ac). Santa Catarina: Cas tellanos 2766 [Herb. Cent. Pesq. Flo-
rest. 3377] (Ac); Smith & Klein 1102 ite poe Smith & Reitz
14332 (W—-2)51613). URUGUAY: H. He Bartlett 21301 (G); Sellow s,
Rh, (Montevideo; Macbride photos 3 17432) (Wp (W—-photo of type).

VERBENA MULTI IGLANDULOSA Moldenk
Bibliography: es shaeioie 12: 79-180. 1966.
ac ns: CHILE: Coquimbo: Marticorena & Matthei 315 (Z—

NANA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 73. 1965.

VERBENA NEOMEXICANA (A. Gray) Small

Soy Stee Giuseaeening Moldenke, Phytologia 11: 473, 490,
& 502. 1965,

Weber & Charette found this plant growing on Acacia flats and

low rocky knolls in Chihuahua, Mexico. The Straw & Forman 1380

istributed as this species is actually V. perennis var. johnstoni
oldenke, Pi

Additional citations : TEXAS: Brewster Co.: E. Ge Marsh Jr. s.
ne om Fra bic (Au--2261:82) . Travis Co.: R. B. Payton 26 (Id).
MEXICO: Chihuahua: Weber & Charette 11660 (Mi).

VERBENA NEOMEXICANA var. HIRTELLA Perry
1965s ttonad bibliography: Moldenke, Phytologia 11: 73 & 502.
The Salgado s.n. [17.V.1961], originally distributed as this
variety, is ac is actually V. canescens var. roemeriana (Scheele) Perry.
Additional citations: MEXICO: Puebla: F. M. Leal s.n. [Teplaca,
2h.VII.1955] (Ip).

NEOMEXICAN XYLOPODA
is dditional perry Mae es solace. ig ee 11: h73 & 502.

VERBENA NIVEA Moldenke

212 PHYTOLOGIA Vol. 13, no. 3

Additional synonymy: Verbena nicea Moldenke ex Hocking, Ex-
cerpt. Bot. A.5: 45, sphalm. alm, 1962.
coin wee raphy: Moldenke, Biol. Abstr. ih 1062. :
rpt. A.5: 45. 19625 ne Phytologia
abe 15683 3 ooh ‘ rciees a 12: 12. 1965.

VERBENA NIVEA f. ROSEA Moldenke
Additional synonymy: Verbena nicea f. rosea Moldenke ex Hocking,
Excerpt. Bot. A.5: 45, sphalm. yphalm, 1962.
at nal bitiiaceshies Moldenke, Biol. Abstr. 37: 1062. 1962;
erpt. Bot. A.5: 5. 19625 Moldenke, Phytologia 11: uth.
1965; ltonke. Résumé Suppl. 12: 12. 1965.

Additional bibhsoerartiy’ N. J. Anderss., Galap. Veg. 199. 185k;
N. J. erss., Vet. Akad. Handl. Stockh. "1853: 199. 18995 Loese,
Verh, Bot. Ver. Brand, 53: 7 [Abhandl. 239]. 1912; H. Be "Davis,
T,

; Loewenfeld . Gardening 21),-——- 5
str. h5: 7066 (1964), 5 (20): ae (1964), and h6: 32h6. 19653
Moldenke, Résumé Suppl. 12: 8 & aes pati Phytologia ll:
» 1966.

Bohm (1964) states that this is an ran transitional host
for Aphis frangulae (A. gossypii), which attacks Cucurbitaceae and
Rhamnaceae in

eastern Austria in time of depression in midsummer
and late summer, and is an essential limb in the secondary host
plant range which concludes with the Cucurbitaceae in autumn.
a pres says that * flower which possessed all desirable
magical si vervain or ho

3
g

n the rt
Druid prtente ihe to make a aproveene ph to earth one they cut
a twig n or mistletoe. In German the plant, and indeed
ths whole Tatennoshe, is called iron-herb, and Hieronymus Bock,
reproving the women of the time for their foolish caparathtiiil
goes on to say 'Thus the vervain is called iron-herb with us, be-
cause we use it to harden iron’. Poachers in Bohemia rubbed their

port
throughout the county Saleh deaetil day ] on roadsides and
c

ac
been found at the climatic optimum in the Sa pay interglacial
period in association with Hedera and Buxus pollen
Clair (en ——- the following nedtevel. couplets:
th Vervin or Dill
ag oy bold ¢ ae sleep on every hill"

1966 Moldenke, Notes on Verbena 213

"Trefoil, Vervain, John's Wort, Dill
Hinder witches of their will"

"Here holy vervayne and here dill
'Gainst witchcraft much availing"

The same author adds the vernacular name "vervin". The M. Abarca
13, originally distributed as ¥. officinalis, is actually V.
carolina L.

Additional citations: ISRAEL: Beach 5407 (W—-233667L). AFGHAN-
ISTAN: Beach 5110 (W—2336478). KOREA: Yongsok 8347 (Mi). HAI-
NAN ISLAND: Lei 231 (Mi).

VERBENA ORIGENES R. A.

Additional et faaiais a Moldenke » Phytologia 11: 475. 1965.

S species has been found growing at 2900 meters altitude in

Atacama

Addi tional citations: CHILE: Atacama: C. Castro s.n. [La Puer-
ta, 20.X1I.1963] (Ac); R. A. Philippi s.n. [Pajonal; 11.185h;
Macbride photos 17434] (W—photo); Ricardi, Marticorena, & Matthei
630 (Ac). Coquimbo: Ricardi, Marticorena, & Matthei 690 (Z).

VERBENA OVATA Cham.

Addi tional be pa ag Be get: Bs poser 10: 289—292.
are Rambo, Pesquis. 2 7 &

ambo (1965) cites paith ge hieta 567 & 61356 [Pivetta 1125]

tron Rio Grande do Sul, the former peed a by himself. He de-
anes the ee as "Erect, very c , leaves A ae and
broad", growing in "Swampy thickets" 4 rh the "Northwestern High-
lands only" of that state, with a general distribution of "Misi-
ones, western RGS, Uru ruguay".

Additional citations : BRAZIL: State undetermined: Sellow 3671
(Macbride photos 171,35] (W--photo of type) .

VERBENA P PARANENSIS Moldenke

ich bibliography: Moldenke, Biol. Abstr. 33: 1215.
19595 king, Excerpt. Bot. A.5: hhe 1962; Moldenke, Phytologia
10: 292-293" a50h,

VERBENA PARODII I (Covas & Schnack) Moldenke
Additional bibliography: Moldenke, cree a pony Rabe
ecent collectors have encountered thi s specie fruit
and ¥. rete vaio

saben 21623 1613 & Sn. (a6. is Herb. Inst. Miguel Lillo Lillo
31745, and Her! Herb. Osten 0676, ted below, were previously i incor-
rectly cited by me as “rine Fevecta Willd., while JUrgensen 1737,
Herb. Inst. Miguel Lillo 3172, and Rufz Leal 7092 were previous-

21h PHYTOLOGIA Vol. 13, no. 3

ly incorrectly cited as V. tenera Spreng.

Additional citations: “ARGENT GENTINA: Catamarca : Jorgensen 1613
(Herb. Inst. Miguel Lillo 31745] (N), 1737 (Herd. Inst. Miguel
Lillo 31472] (N), 8.D. {[26.III .917; Herb. rb. Osten 19676] (N). Men-
dora: Rutz Leal 7092 (N).

VERBENA PARVULA Hayek
enh premio Moldenke, Biol. - 35: 1688.
1960; Hocking, Exc to Als eee 1962; gonad Phytologia

10: 296-297 {196h) ae th 485.

VERBENA PARVULA var. GIGAS Moldenke

hdd? tone) bibliography: elas, Biol. Abstr. 35: 1688.
1960; Hocking, Excerpt. Bot. A.l: 592. 1962; Moldenke, Phytologia
103 bon. 196k. :

VERBENA PERAKII (Covas & Bac nrenstlh Moldenke
ac ae Pe fe ope tngk aria ee a & Schnack ex
Hocking, Exc oes 335, 3 aig BBS:
Additio ok & fee pombe a aa "Bot. A.6: 335.
1963; Moldenke, og gia ui: She 1968) aside” Résumé
1

found this species growing in sandy soil at 750 meters
altitude, with blue flowers (Venturi 10301), in anthesis
March. This collection was previously erroneously cited by me
as V. dissecta Willd.

Additional citations: ARGENTINA: La Rioja: G. Dawson 3380 (Ip)-
Salta: Venturi 10301 (N).

VERBENA PERENNIS Woo
Adat tional eieliceregiy VMoldenke, Phytologia 11: 475, 76, &

502. 1
ts citations: TEXAS: Brewster Co.: B. H. Warnock
20437 (Ld). ee

VERBENA PERENNIS var. JOHNSTONI Moldenke
Y gakeaes bibliography: Moldenke, Phytologia 10: 305-306.
Straw & Forman describe this plant as an erect perennial herb,
common but scattered throughout woods in open pine-Yucca-scrub
rea community on all slopes, with light-violet gee ers f

to white in the throat, flowering and fruiting in July. 0
asttonal citations: MEXICO: Nuevo Leén: Straw & Forman 1360

same cient ag,
tional biblio loeees Moldenke Lae go — 2 ’
467, 2 hT5, & 16. 1965; Moldenke, Résumé Suppl. 12: 1. 1965+
describes the specimen cited below as ; mplant. Re
erty in seratae a yard round, only the flowers ascending".

1966 Moldenke, Notes on Verbena 215

Additional citations: ALABAMA: Co ounty undetermined: Winchell
207 (W—73533). INDIANA: Saint Joseph Co.: Nieuwland s.n. [St.
Mary's, Sept. 15, '2h] (Mi).

VERBENA PERTURBATA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 176. 1965;
Moldenke, Biol. Abstr. 6: 32h6. 1965.

VERBENA PERUVIANA (L.) Britton
Additional bibliography: Moldenke, Biol. Abstr. 35: 1688 (1960)

and 36: 719. 1961; Brilmayer, All About oe 350. 1962; Hocking,

Excerpt. Bot. A.l: 592 (1962) and A.5: bls. 19625 Moldenke, Biol.

Abstr. h6: 632, 32h6, & 3618. 1965; Rambo, esquis. : 7

49 & [60]. 1965; Moldenke, Phytologia 11: "ihe, 470, & b une-LT?

1 a! 180. 1966.

r (1962) calls this plant "Peruvian flame" and de-
scribes it as "Evergreen trailer offered by house and greenhouse
growers for hanging gare and summer containers. Its 2 inch
crinkly leaves are suenly toothed, the flowers a glowing crimson.
A oben 'Chiquita' has flowers striped with gay red and white,

ppermint
Ranbe (1965) prea Herb. Anchieta 450, 1137, 3042, 7850, 8990,
= aos 9958, 11252 [Heidler s.n g-n.], 25190 25190 izesteees a2,
5213 [Theissen 8en.], 25970 (Henz s.n.], 26767 (Friderichs s.n.],
28, 29550 50, 32016, 32539 [Henz Sal, 33438 38 [Henz 2-1) 2 36L17,
030 [Spies s.n.], 37347, 37498, 38753, 39592, 39810
12781 eo , 13282 , Lapel Wats, 16395, 782 17621 ‘eta 3

Camargo 33 at, 59156 [Pi a vetta aay & 60L56 [Camar amargo 502] from Rio
Grande do Sul, and 55318 [Reitz 4703] from Santa Catarina, Brazil

owing in

ampos......Over the whole of the Campos area, especially
in the northeastern Lowlands and near the sea coast" in Rio Grande
8 Sul, oe Ae general distribution of "From Peru, Bolivia and

rthern Argentina, RGS and Uruguay".

Additional citations: BRAZIL: State undetermined: ig gm
Macbride patee 17407] (W--photo). URUGUAY: Comerecr 72 [Her
Jussieu 5141, in part; Macbride photos 39503, in part] premtote)
ARGENTINA: Buenos Aires: Commerson 71 [Herb. aah p98 fea el).
Part; Macbride photos 39503, in part] (W--photo

PERUVIANA f. ALBA Moldenke
Additional bibliography: Moldenke, Biol. Abstr. 36: 719. 1961;
lig. nee Bert Bot. A.5: hl. 1962; Moldenke, Phytologia 10:

PERUVIANA var. GLABRIUSCULA Kuntz
Additional bibliography: Moldenke, Phytologia 10: )90—91.

216 PHYTOLOGIA Vol. 13, no. 3

196; Moldenke, Biol. Abstr. 6: 3618. 1965.

VERBENA PHLOGIFLORA Cham
Additional bibliography: Rambo, Pesquis. Bot. 21: 46, h9--50,
& [60]. 1965; Moldenke, Phytologia 11: 467 & 477. 1965.
Recent collectors have found this plant cos: > pte eens
stream beds, on rocky banks, and ruderal = Santa C
Rambo } cites Herb. Anchieta huh, 1120, 296h [Ba gli estes s.n.],
4273, 4488, 4508, 4775, 78L8 [Thiessen ce, “T1605 [Buck Bell,
25258 Tiheissen sane, 26992, 26503, 30011 [Burich s.n.], 32617,
33279 (Emrich s.n.J, 35460 [Ritter s.n.], 35908, 36419, 37693,
38555, 39020, ; Bae, we Bite L020, L165, L908h, 50183 [Er
rich s.n.], 369, S40 59, & 55939 from Rio Grande do Sul,
os6é Traits aay } cic 0 TReits ooh 5090], & 55301 [Retz 472)
— bee Onion, “and 3982 from Uruguay -- all ~ collected
except wi otherwise noted. He admits, however, that
ose [Reitz 5073] —_ been identified as f. alba Moldenke. He
describes V. phlogiflora as "Herbaceous, weak, erect or leaning
to the neighbors, up to 1 m high", found in tte +t to swampy
thickets in the Campos area.....O0ver the whole area of the Cam-
pos" in Rio Grande do Sul, tye with a general distribution
"From Cuba to Uruguay and no rn Argen
Additional citations: BRAZIL: P Paran4: Dombrowski 528 (Ac).
Catarina: A. Castellanos 21,780 [Herb. Cent. Pesq. Florest.
3564] (Ac); Smith & Klein 13223 (Ac); Smith & Reitz 12592 (Ac).
PARAGUAY: Hassler 4585

ar IFLORA f. ALBA ee
Additional bibliography: , Pesquis. Bot. 21: 50. 19653
Moldenke, Pay locte li: Wie 19 1962.

VERBENA PINETORUM Moldenke £02
Additional ais arenes: Moldenke, Phytologia 11: h77 &
(1965) and 13: 187. 1966; Moldenke, Résumé Suppl. 13+ Bs i
Recent collectors have found this species gr rocky
soil and subsea: pratttte in August. The 0. M. he 10595
sn: , cited be-

vious installments of these notes.

Additional citations: MEXICO: Hidalgo: 0. M. Clark 7059 (N)s
Galvén s.n. [4.VIII.1963] (Ip). México: Paxson & & Barkley 1Gle77 1641839
(N). Oaxaca: C, L. Smith 221, in part (N).

NA PLATENSIS Spr é0]> |
Additional peceet rae AA Rambo, Pesquis. Bot. 21: 50—51 & [
1965; Moldenke, Phytologia 11: 177. 1965; Moldenke, Biol. Abstt»
6: 3618. 1965,
Rambo (1965) cites Herb. Anchieta 27997 [Rosengurtt B.610) &

32950 [Rosengurtt PE PE.S056] from Urug rom Uruguay and $3156 [Spies s-Me 5 ole |

1966 Moldenke, Notes on Verbena 217

Rio Grande do Sul, Brazil. He describes the pant as "Prostrate,
rooting at the lower joints, branches radiating, up to rocks cn
rl growing in "Dry, grassy Campos" in the "Southwestern plains
in Rio Grande do Sul, with a general distribution of "Chile,

thas Argentina , Uruguay, w westernmost RGS, Paragua:

Additional attaldages BRAZIL: State undetermined: Sellow SDs
(Wacbride photos 3,356] (W--photo). URUGUAY: Sellow 8.n, [Monte-
video; itnabiride” saan 17452] (W—photo of type).

PLICATA Greene das He
dditional mek Moldenke, Phytologia 11: 11~19 .
‘Q964) Poy li: & aT, & 502. 1965; Moldenke, Résumé Suppl.

ak saakke on Shinners 30033 are described as having been
"violet-blue". Recent collectors have found the species growing

oadsi "abundant
Webb County, Texas. The pt lg i. & metas 8183, 0 sayin
distributed as V. plicata, is actually | V. tmlei Small.

Additional citations: OKLAHOMA: pee ios C. M. Rogers 5437
(Mi), 5990 (Mi). TEXAS: Andrews Co.
(Au--221199) . ei 2 Coes Pe cabrere, ae 253852) ; N. Castillo
§ (du—-23342h); Flores, Chavez, & Hein h2 (Au--233433). Frio Co.:

Painter, & Bark Raak 7 14227 (Ld); J. R. Rodriguez 2 (Au—
235209); Be Le B. L. Turner 49 4989 . (Au—-229812). Jim Hogg Co.: | D. Montes
Sh (4u--232735). LaSalle Co.: Winch 6 (Au—2222h0), Starr Co.:
R, Acosta 73 (Au—232669); Alvarez, Guajardo, ee & =
523 (an (au—22221,3), BOL. (Au-222194) j Banos, Sandoval , Salazar,
7867 (Au--2222h,). Stonewall Co.: Shinners Sap “Ge =
208). Webb Co,: Arzola 143 remrsiotonl Flores & Flores 117
(Au~-23337h); Gamboa & Dohnke 10h (an--23275); Jorge paces 2 39
(Au—2321,68) ; Sate, ote, Herrere, & WeCart 8070 (A 2008). mate

Co.: Herrera MoCart 7698 (Au—222238, Ip 1p); Ramos,
laser, & McG. Hetart ELT 7 (Au—2222h2); M, de J. Solis if 1 = 55H).
MEXICO: Guerrer 3 Be de la Garza 67 gto oe

POGOSTOMA Klotzsch
ang te onal bibliography: Moldenke, Phytologia 11: 19—20 (196)
$ - 1965.
Villafafie 52) is described as having had mly eitel by a
Jamuary. It was previously erroneous me
Ys dissecta Willd. ‘The Hunziker collection cited below was pre-
viously cited by me, in error, as V. tenera Spreng. The flowers on
sane are described as "violet®, blooming in March at 2500
Additional citations: ARGENTINA: Cérdoba: Villafafie 52 (N). La

218 PHYTOLOGIA Vol. 13, no. 3

Rioja: A. T. Hunziker 5160 (N). CULTIVATED: oe Herb. Hort.
Bot, Berol. son. [Oct. 1839; Maebride photos 1740, in part] (f#—
photo of cotype), s.n. [Aug. 1843; Macbride 1740, in eet (Ww
photo of cotype) -

NOTES ON NEW AND NOTEWORTHY PLANTS. XLIII

Harold N. Moldenke

socio KEGELIANA (KOrn.) Moldenke, comb
Sos kegelianus Ktrn. in Mart., fl. B Bras. 3 (1): 438.
186

te VIVIPARA var. LONGIPILOSA Moldenke, var. nov
aec varietas a forma typica speciei pedune culis densissime
seta willosis, pilis patentissimis,

This variety differs from the typical preee of the species in
having the peduncles very densely white-villous, with long rather
stiff hairs spreading at right angles.

e type of the variety was collected by Rolla Milton Tryon, Jt»
and Alice F. Tryon (nee 6784) in dryish soil under rocks on
sedg

3 from Bell
zonte, Minas Gerais, Brazil, on November 21, 1965, 8 and is deposited
in my personal herbarium at "Plainfield, New Jersey

LIPPIA RECOLLETAE var. PICKELII (Moldenke ) Moldenke, comb. nov.
Lippia pickelii Moldenke, Lilloa $: l2h—-l25. 19h0.

LIPPIA TAYACAJANA var. SESSILIFLORA Moldenke, var.
etas a forma typica speciei totzscunceanitey sessil-

ibus
This variety differs from the typical form of the species in
having its inflorescences sessile or subsessile.

The type of the variety was collected by Paul C. Hutchison and
J. Kenneth Wright (eee 7025) in the —- of the oes erate 2e
im, above Balsas on the road to Cetendin, l; km. east of Hacienda
Laer, at an ping on of 1725 Drip ed caeine of Celendin, Caja
» Peru, on er 17, 1 and is sited in my pers
berbariom at Plainfield, New Jersey. igs ep dexortt this
plant as nearly or completely leafless stride te The
local fee several km,, the flowers th yellow Sonar
Mo goergs & tea being made apa te the ype = by the nar

~n a retaecnermneeeneteneronnetam

BOOK REVIEWS
Alma L. Moldenke

"The Great American Forest", as Rutherford Platt, xii + 271 p
. illustr. Prentice-Hall Series in Nature and N Natural ee
Prentice-Hall Publishers, Inc., Englewood Cliffs, New Jer-
sey 07632. 1965. $6.95

~ "If only people would catch a vision of our fabulous f rapietet
their ancient heritage, their beauty and beneficence, their m
ing for our lives toda y- - - before it is too late", So —

The book has several of the author's artistically exquisite
Photographs. There is a useful index. Scientific oar

rstandable manne
the book deserves wide circulation) not familiar with our common

names for trees, the scientific names would have been a helpful
ted in the text or in a separate listing. The type is

grade school instructors could be much enriched by the
k.

"The wildness of our gis age is a quality of life for all
Americans whether or not they personally see it. The mere idea

eyes ~- the knowledge that such things exist,
there is a North Woods, an unspoiled area in the deciduous forest,
219

220 PHYTOLOGIA Vol. 13, no. 3

a deep silent valley in the Rocky ate aempomag enhances the value
of every county park, hedgerow and woodlot."

"Advances in re Pee 2", edited by J. B. Cragg, xi
264 pp., illus ademic Press, London & New York City,
New York. os sap 6d. or $9.50.

Continuing the intention expressed in the first volume "to
present comprehensive accounts of selected topics of ecological

cology, as well as specialists in ecology tain a balanced
picture of ng place", this volume presents four
uable papers in quite thorough form.

The first is on population dynamics, is written by M. E. Solo-
mon of the Pest Infestation Laboratory of Slough in is
entitled "Analysis of Processes Involved oe the Natural Control
of Insects", and is concerned with expounding the types of pro-
cesses influencing abundance, the roles of different factors and
processes in natural control, the density relationships in the
action of premtars and parasites, and an inquiry into parasite~

The second aS entitled "The Use of Statistics in Phytosocio-
logy", ie Pepin kg by J. M. Lambert and B. M. Dale of the Botany
Departmen of the Wie vaceity of Southampton in England, is de-
veloped sph some ideas and techniques differing from those
presented in the earlier volume by Dr. M. E. D. Poore, and is
effective in demonstrating the value of computers in various
melee of analysis of phytosociological da

e third is "Litter Production in Piette “of the World" by

sis of litter » a survey of factors effecting the
fall of litter, the seaso hanges in ude
leaf crops, a study of leaf litter index to net pro~

assume additional significance - as a source of food protein by

chemical extraction for the world's e tions and 86
a site for cultivating Imge crops of edible fungi, while waiting
for rhead forests to mature.

The
Harrison of the Department of Botany of the University of
ham in s work deals with genecological categories, t
technique and differentiation, with the evolutionary aspects

he
genecology is a distinct synthetic discipline culled from gene por
taxo and plant physiology that should not be assimilated

NOTES ON LAURACEAE OF TROPICAL AMERICA
I, THE GENERIC STATUS OF
NECTANDRA, OCOTEA AND PLE UROTHYRIUM

Caroline K. Allen

Recent collections from the tropics, particularly from So-
uth America, have made available materials that in the course of
Study contributed much towards clarification of our concept of the
genera of Lauraceae. It seems appropriate at this point to dis-

hemisphere, represent approximately half of the known species
of this region; Pleurothyrium Nees ex Lindley (1836, p. 442)* is
@ small genus of about twenty species, occurring in northern So-
uth America, predominantly in Peru.

A study of the available types in conjunction with pertinent
literature shows that taxonomists in general have had a more or
less clear concept of Nectandra and Ocotea, Pleurothyrium, how-

ver, has been a problem. Gareful consideration of the material

dra and Pleurothyrium apart from Ocotea. It is the comparative-
ly little-known Pleurothyrium that I wish to discuss here. Recent
Studies of the genus have shed light on staminal structures which,

nd | ellia. The members of the genus are mentioned as
trees or shrubs, with sparse, alternate, evergreen, coriaceous,

etait
Pleuroth rium, from the Greek words meaning "'side'' and "little
h

°Penings" because of the small, lateral valves of the anthers ob-
served Particularly in stamens of the first two series.
221

222 PHYTOLOGIA Vol. 13, no, 3

penninerved leaves, often ferruginous beneath. The inflores-
cences have been variously described as axillary or terminal,
paniculate or thyrsoid, with deciduous bracts; the flowers herm-
aphrodite, with a 6-parted perianth, coriaceous or fleshy, the
lobes (deciduous in fruit) more or less spreading, oblong, the
tube obconic; the fruit a berry, immersed, at least in the young
state, in the ee tube of the perianth, with spreading-
crenate margin.

Nees has described the perianth of the genus as having 6
persistent, glandulose lobules" alternating with the lobes of the
perianth; 9 perfect stamens, similar structurally, the anthers
Sukcuhiat 4-loculed, all of the locules lateral, parallel, ovate,
some retrorse, others introrse, the filaments short, the stamin-
odia short, tooth- Pager biglandular, the glands originating at the
base from the back.

e diagnosis of his first-published species of the genus,
a coe Nees elaborates further his description
of the staminal cycles. He notes 6 glands, alternating with the
— at io height of the stamens, thick, fleshy, glabrous, yel-
eae from the wide base into an obtuse acumen twice

se width of the base, without (in the dried state) showing an in-

terminate crease, within furnished with broad, obtuse keels,
stretching from the base into the bottom of the flowers; 9 stamen®
clustered, short, small compared with the flower, glabrous, the
6 exterior (ser I & II) opposite the lobes of the perianth, the fila
ments narrow, shorter than the anthers, the anthers oT
in the transverse diameter slightly larger, obtuse, the apex bent
forward, a little more extrorse, the interior face a little more
swollen; lateral locules in pairs, ovate, parallel, subequal, pro"
duced from the base almost to the apex, dehiscent by valves from
the bottom to the top, the exterior ippnies of each pair slightly
lower and turned about neeeenany: + e interior, however, turned
introrsely; the anthers of the third or inner cycle (ser IIL) of st#
mens conform to those of “— vraag the filaments slightly shorte’
and broader than those of the latter, the glands at the base of
these are obsolete or, rather, entirely confluent with the gland-
ular base of the flower.

* 2 ae
I have not studied fruiting material of this species; nor can I
visualize the structure of the fruiting cup here described.

eK. ; é ial
Biglandular staminodia have not been noted in our materia

1966 Allen, Lauraceae 223

Actually, Nees described the floral structure as it appear-
edtohim. The 6 glands alternating with the 6 lobes (the term

be so fused as to lose their identity and thus appear to be lacking
altogether. Nees does not elucidate further here on the locules
of the anthers.

the lateral pairs almost contiguous at the apex, one locule intr-
Orse, the other conversely more retrorse. (I take this to mean
that, seen laterally, the 2 locules appear one above the other,

but ina different vertical plane, the upper introrse, the other ex-
trorse.) Because Nees has not taken into consideration the ori-
gin of the tissues of the so-called "disk", his description does not
give an accurate picture of the structure or origin of the staminal
cycles,

Shortly after Nees! introduction of Pleurothyrium, Endlich-
€r in his Genera Plantarum (1836-40, p. 320) published a diagno-
Sis of the genus essentially the same as that of Nees, except for
his interpretation of the staminodia as a fourth staminal cycle.
Dietrich (1840, p. 1332) apparently copied Nees; Spach (1841, p.
472), merely assigns the genus to the Tribe Dicypellia.

M
hypogynous disk adnate to the tube of the calyx, the free apex of
the disk inflexed above the ovary, forming a narrow ring bearing
Stamens and glands. The 9 fertile stamens inserted in the ori-

fice on the epigynous ring are free, subequal, and much shorter

the anthers are cubical-oblong, truncate,

*
This is inaccurate; the glands arise at the base of the filament.

22h PHYTOLOGIA Vol. 13, no. 3

the four locules oval, placed in a nearly horizontal series, the
outer facing the sides, the intermediate being slightly higher,
facing introrsely in the outer series of stamens, extrorsely in the
inner series; staminodia 3, small, triangular, eglandular; the
ovary included entirely in tube, free, ovoid, the style equalling
the stamens; the stigma depressed-subcapitate; fruit unknown,
undoubtedly surrounded by a truncate cupule provided with a du-
plicate margin. sry

In his treatment of the Lauraceae in Martius’ Flora Bras-
iliensis, Meissner is still preoccupied with the structure of the
hypogynous disk in the flower of Pleurothyrium, but at the same
time distinguishes the stamens of the third cycle with their pairs
sessile glands arising at the base of their filaments. (In some
instances, one is able to separate the inner cycle easily from the

ands of the latter.) Meissner's lengthy description of the anthers
does not give a clear image of their structure. He speaks of the
four locules being ina nearly horizontal plane but does not men-
tion the parallel vertical planes in this connection.

Baillon (1870, p. 438, 439; 478) is the first author to report
the occurrence of a thick obconical receptacle, clothed (with pub-
escence) within a thick disk which more or less curves at the ape
(presumably the glands first mentioned by Nees.) He describes 9
fextile stamens with the perigynous perianth inserted outside the
margin of the disk; the 6 outer alternating with the lobes of thes®
the 3 inner (ser III) biglandular at the base; the filaments thickish,
the anthers thick, cubical-oblong, their four locules placed in
series at length almost horizontal; the exterior sublateral, the in-
termediate (locules) situated slightly higher facing, in the 6 outer

es, introrsely, in the interior (ser III), extrorsely, the 3
sterile staminodia (ser IV), small or none; ovary free, included
in the receptacle; the berry surrounded by the receptacle, devel-
oped in fruit, truncate at its apex. Baillon believes that Piew
thyrium differs from Nectandra in minor or secondary character®
only. Eee oe eRrar

ee : : ‘ g
Meissner'’s is the first mention of the staminodia being egland
ular.
3K

The suburceolate calyx surrounding the young fruit freq
bears the remains of the lobes and staminal cycles; this ©?
might give rise to the impression that the margin of the mature
cupule would be double.

vently
nditio

1966 Allen, Lauraceae 225

tioned by Nees in his description of P, bifidum, are relics indic-
ating the points of separation of the once discrete glands. In ad-
dition, there is the convex pubescent projection (the obconical
tube mentioned by previous author s) on which seemingly the ovary,
the three cycles of stamens with their associated glands and the
staminodia, all of which cycles actually arise from the "epigyn-
ous" ring of Meissner which, if it is not a true hypanthium, is
analogous to the latter structure.

Bentham and Hooker (1880, p. 159) accept Meissner's inter-
pretation of the narrow ring formed by the hypogynous disk.

Mez (1889, p. 468) seemingly was the first student of the
Lauraceae to understand the structure and origin of the "disk"
mentioned by previous authors. In his key, he separates Ocotea
and Nectandra from Pleurothyrium by the presence in the latter

"berry" in the early stages of development.

Mez observed that the singular genus, closely associated
to none, is easily and naturally separated by nature of the glands
and anthers. He notes three species which seem to show affilia -

mens of ser I & II; and Ocotea bahiensis which, provided with
ighteen glands, shows an entirely different form of anthers.

Pax (1889, p. 116), on the other hand, still adheres to the
theory of the fleshy disk surrounding the filaments at their base
and spea rated into six teeth alternating with the stamens.

5 “pecies were described by O,C, Schmidt (1928, p. 235) with
all stamens Surrounded at base by two large, confluent glands,
the anthers Subrectangular...."'

226 PHYTOLOGIA Vol. 13, no. 3

Macbride (1938, pp. 929-931) in his Flora of Peru gives
only a basic generic description which differs from previous trea-
ments in stating that the perianth tube is rarely well defined. Ap
parently he has not seen the fruit described by Nees. Since Nees!
publication, Macbride seems to be the first to question the valid-
ity of the genus Pleurothyrium in relation to Ocotea.

Lasser (1942, pp. 118-19) published descriptions of two
species from Venezuela in which he describes the anthers as sub-
globose. An illustration of Pleurothyrium reflexum shows a lat-
eral (or profile) view of a stamen of the outer series--the first
depicting the actual structure of the stamens of the genus.

The important contributions of Kostermans (1952, 1957) are
discussed at the end of this paper.

Hutchinson (1964, p. 138) separates Pleurothyrium from the
rest of the tribe Cinnamomi by: Flowers bisexual; staminodia of
the 4 cycle present, thickened at the apex; disk well-developed,
6-lobed. His description goes back to Nees! diagnosis of P. bi-
fidum.

Of the early students of the Lauraceae, only Nees and Meis-
sner made any attempt to describe the differences that they ob-
served that prompted them to recognize Pleurothyrium. I have
gone into the history of their treatments at such great length ber
cause, inspite of the lack of clarity of their descriptions, and the
lack of any supporting illustrations, both of these taxonomists
recognized an entity distinct and separable from Nectandra and
Ocotea. The latter genera are, for the most part, as I hope t0
demonstrate in a forth-coming paper, distinct from each other.

In his Historical Survey of the Lauraceae, Kostermans
(1952, pp. 22(121), 23(122)) brought the history of Pleurothyrium
up to date. Because he considered the character (the number of
glands) used by Mez to delimit Pleurothyrium to be of no generic
value, he has reduced the latter to Ocotea. Mez and Kosterman®
agree that the position of the anther locules, likewise, is of n°
generic value. But, both failed to take into account the strongest
character brought out by Nees--the consistently distinctive shape
of the anthers which separates the two genera.

F m-*

Subsequently, Kostermans (1957, p. 39) has seen fit to c? :

bine all three genera, Ocotea, Nectandra and Pleurothyrium sc
Ocotea. If we accept this reduction, the entire taxonomic fabr

nee;

1966 Allen, Lauraceae 227

of the Lauraceae will be further imperiled; for elsewhere in the
family there are several instances where reasons equally

To summarize: I continue to maintain the three genera Oco-
tea, Nectandra and Pleurothyrium, The accompanying figures
show the essential differences in the staminal cycles of the three
taxa. The characters that I believe to be useful and valuable ap-
pear in the following key. Until the fruit of Pleurothyrium be-

228 PHYT0d06.7-4 Vol. 13, no. 3

KEY TO THE GENERA

NECTANDRA, OCOTEA AND PLEUROTHYRIUM

1. Stamens (with one known exception)

of ser I & II without glands
anthers of ser I & II when viewed
ey
locules occurring on
the introrse surface.
2. Qne pair of locules in the upper

half of the anther, the other

pair in the lower (introrse

view Peace bhecbeis tsa. eee

eel

ae

are formation, upper pair lat-
eral, lower pair extrorse
(extrorse View: ser it] oo. ..cc ees: NEC I Raee

EE
3. Anthers ovate, fleshy, pap oo

lose, attentuate; connecti

tissue abundant.... Sec nara pomeein

3. Anthers usually broader than

locules occu-
pying entire anthers

ereree ee ene

rors ss : Section Porostema

Stamens (with two known exceptions)
of ser I, II & III with glands;
anthers of ser I & II inclined

but viewed laterally
one of each pair visible ....... PLEUROTHYRIUM

1966 Allen, Lauraceae 229

LITERATURE CITED
Baillon, Hist. Pl. 3: 439, 478. 1870.
Bentham & Hooker, Gen. Pl . 3(1): 159. 1880.
Dietrich, Syn Pl. 2: 1332. 1840.
Endlicher, Gen. Pl. 1: 320. 1836-40.
Hutchinson, Gen. Fl. Pl, 1: 138, 1964,

Kostermans, Jour, Sci, Res, Indonesia 1(5): 22(121), ae a
Comm, For, Res, Inst, Indonesia 57: 39-4

Lasser, Bol, Tecn, 3: :18-19,' 1942.
Lindley, Nat, Syst, Bot, ed, 2, 442, 1836,
Macbride, Field Mus, Publ, Bot, 13(2): 928-931. 1938,

Meissner, DC, Prodr, 15(1): 168. 1864;
in Martius, Flora Brasiliensis, 5(2): 279-280. 1866.

Mez, Lauraceae Americanae 468-472, 1889,

Nees, Syst. Laurin. 23, 24; 349-352. 1836.

Pax, Engler & Prantl, Pflanzenfam, 3(2): 113, 116. 1889.
Schmidt, Notizbl, Bot, Gart, Berlin 10: 235, 1928,

Spach, Hist. Nat. Veg, 10: 472, 1841,

230

Fig.

PHYTOLOGIA

EXPLANATION OF PLATE
I. Pleurothyrium

Ze, stamens, ser I & II, ventral.
Ze', stamens, serI& II, dorsal.
Ze", stamens, ser I & II, lateral.

stamens, ser III, ventral.
zit, stamens, ser III, dorsal.
Zfi"', stamens, ser III, lateral.

; 4d .cores

Ae, stamens, serI & II, ventral.
Af, stamens, ser III, ventral.
Af', stamens, ser III, dorsal.
Af", stamens, ser III, lateral.

. Ill. Nectandra Section Pomatia

Ze, stamens, ser I & II, ventral.

eral.
Zg, staminodium, ser IV, ventral,

. IV. Nectandra Section Porostema

Ze, stamens, serI & II, ventral.

er
Zg, staminodium, ser IV, ventral,

Vol. 13, no. 3

1966 Allen, Lauraceae 231

iti oa
tee.

Fie I
Ze’ Ze y da Zg’

an PB i

zg a

NOTES ON TROPICAL AMERICAN LAURACEAE
Il. COSTA RICA

Caroline K. Allen

Ocotea skutchii C,K, Allen, Jour. Arnold Arboretum 26: 352.1945

Seotea williamsii P,H, Allen, Rainforests of Golfo Dulce
» pl. 27 27. 1956.

Distribution: Known only from Costa Rica.
me. Be Rica: San José: Vicinity of El General, Skutch 3062

(f1., type, GH: isotype, ane Puntarenas: forested hills near
Golfito de Golfo Dulce, Paul H. Allen 5983 (fl., fr. isotype (US).

The distinguishing features by which Paul Allen recognized

his new species were the extremely large, oblong or ellipsoid
fruits and the axillary tufts of hair that occurred abundantly on
the lower leaf surface. Only the cupule of Ocotea skutchii was

available at the time of publication, but according to the descrip-

tion, it is identical with that of O. williamsii. The tufts of hair
on the lower surface of the leaves do not appear onclose examin-
ation to be the usual type of pubescence found in the glands of

many species of Lauraceae. Possibly they represent an epider- —

mal growth induced by insect attack near the primary and fre-
quently the secondary nerves.

The flowers available from the type number proved ”
abnormal in structure, a condition which could also be the re
of insect infestation,

However, the general habit of growth of the branchlets,
twigs and the leaves with long-decurrent blades simulating of
petiole are the same in both types. The greenish yellow St:
the specimens in the dried state presents a striking background .
for the bright yellow venation, another character in commom
Both are trees over eighty feet in height

Research for these studies was carried out at the New
Botanical Garden with the assistance of the National Scienc®
dation (Grant No GB-3048), whose generous support made if
publication possible

232

ool

the
a long

york
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| PHYTOLOGIA

Designed to expedite botanical publication

Vol. 13 July, 1966 No. 4 ~

CONTENTS

5 _ FOSBERG, F. R., Systematic notes on Micronesian
plants. 2 . Po ee 233

. OLDENKE. H. N., Notes on new and pened
Urs Xv : peek Gee i ee a RAR

MOLDENKE. H.N., Additional notes on re genus
Sees I 2 aS pee nee . 243

| MOLDENKE, HN: oe notes on the genus
Citharexylum.

_ Published by Harold N. Moldenke and Alma L. Moldenke

303 Parkside Road =
Plainfield, new Jersey, US.A.

Wine of this number, $1; per chtuine: $5.75, in n advance,
: ne or aes at clone of volume ,

- ime As number Bes was issued a Apri 6, 1966

SYSTEMATIC NOTES ON MICRONESIAN PLANTS, 2.

by F. R. Fosberg

a - (For No. 1 see
Phytologia 5; 289-292, 1955). The present number contains notes
and new combinations in the Piperaceae,

PIPERACEAE
e family Piperaceae is inherently a very difficult one,

Th
aS are most families with reduced or simplified floral structure
and many species, There is no doubt that a large number of

Even k
Species that he had studied in the field, had great compunction
in reducing those that he only knew from dried material and es-

to

s
his predecessors, such as slight differences in leaf shape,
Ora of Micronesia, the trivial nature of some

t

° e
Cures used to Separate them became very evident, and attempts to
fonstruct a usable key were frustrating.

Some of these problems are more or less alleviated by com-
bining Species where they do not seem represent morphological-
mu distinct populations. To extent this is done below.

ver, even this leaves certain species very difficult to sep-
arate in the field. Further field study may eventually indicate
233

23h PHYTOLOGIA Vol. 13, no.

that ae omia glassmanii, P. kusaiensis and P. breviramula
should be combined, and that Peperomia el and P. krae-
meri are cospecific with P. mariannensis. I hesitate, however,
to make these reductions without an opportunity to look again
at the plants in the field, now that I have be ae understand-
ing of the characters that purport to separate t them

e reductions made below are discussed ei The

eas status of Piper methysticum For £. is: aleo
considered, The species of Piper recognized pis Micronesia in

the present treatment are mostly very distinct and easily sep-
arated if fertile material is ce ottaule

PIPER L.

PIPER BETLE L., Sp. Pl. 28, 1753.

its range shows that it is a complex of forms which seem to hav
little significance, although they are by no means all found
throughout the range of the species. To disentangle these des

treatment of the Micronesian Piperaceae (Occ. Pap. Bishop Mus.
22: 83-108, 1959) admitted only two, and those at the varietal
level.

e the ett texmnces are only trivial ones, mainly in
leaf out iide and prese absence of puberulence on the
nerves, and since thers is no clear geographical separation, I
prefer to treat the variations as formae. The necessary combin-
oo synonymy, and distinguishing characteristics are as fol-

PIPER BETLE L. f. BETLE
Piper betle L., Sp..:Pl, 28, 1753.

striate-angulate, leaf blades eee nt base
tlieda 4 subcordate or only slightly cordate, glabrous

eems to be the plant indicated by Linnaeus in Flora
aay bites i- 12, 1747, which L take to be the main basis for
the Species Plantarum. treatment, Therefore the type locality
- pomersclags ae Ceylon, in spite of being indicated by Linnaeus
ni

In Micronesia this form is found in the Marianas and i
known from Agrigan, Alamagan, Saipan, Rota and Guam islands.

1966 Fosberg, Micronesian Plants 235
PIPER BETLE f£. MARIANUM (Opiz) Fosberg, stat. nov.

Piper marianum Opiz in Presl, Rel, Haenk. 1: 159, 1828,

Paper betle var. mariannum [sic/ C. DC. in DC., Prodr,
16(1): 360, 1869,

Piper Tee om Opiz in Presl, Rel. Haenk,

1: . 156,

Stems terete, leaf blades glabrous beneath, ee ovate-
cordate, basal sinus deep, base not conspicuously obligq
apex notably acuminate,

iz' description of P. potamogetonifolium fits this form,

Opi
The basis of the statement by Safford (Contr. U.S.N.H. 9: 355,
1905) that this plant is an "undershrub" is unknown, as Opiz
gives no information as to habit, Piper betle is a on imber,.

n Micronesia £. marianum is known only from Saipan and Guan,
in a: Marianas,

PIPER BETLE £, DENSUM (Bl.) Fosberg, stat. nov.

Piper densum Bl., Verh, der Bat. Genoots. 11: 193, 1826.

Piper betle var. densum (Bl.) C. DC. in DC., Prodr, 16(1);
>» 1869

Stems terete, leaf blades puberulent beneath, especially on
the nerves, broadly, somewhat irregularly, ovate cordate, with
deep basal Sinus, not or slightly oblique.

Yuncker, op. cit, p- 90, has referred the puberulent Micro-
nesian specimens here. We have not seen Blume's type, which
“ame from Java, A specimen collected in Barrigada Village,Guam
(Stone 5126) is almost imperceptibly puberulent, or papillate,

Nn Micronesia f, densum is known from Alamagan and Guam in
as lacianas, and Palau, Yap and probably Ponape in the Caro-

PIPER GUAHAMENSE C. DC. in DC., Prodr. 16(1): 336, 1869.

__. Typically this species, in Guam, is somewhat puberulent or
hirtellous on the under sides of the leaves and on younger parts.
= me islands ike rebe.e north--Rota, Tinian, and Agrigan--it is
tabrous, Ag Ss plant has also been collected on Guam,
Tuncker (1959) igbnwisies the glabrous form as var. glabrum,

2% PHYTOLOGIA Vol. 13, no. &

Although the ranges of the puberulent and glabrous plants
ture of the difference

are partially distinct, the trivial natu
fo

and wa

sheet is the only basis for ascribing the glabrous form to Guam,
However, there seems to be no reason to suspect an erroneous
locality for the sheet.

PIPER GUAHAMENSE C.DC, f£. GUAHAMENSE

Known only from Guam.

PIPER GUAHAMENSE £. GLABRUM (Yuncker) Fosberg, stat. MOV.

Piper guahamense var. glabrum Yuncker, Occ. Pap. Bishop
Mus. 22: 87, 1959.

Known from Guam and the Marianas Islands north of Guam.

PIPER LATIFOLIUM L, f., Suppl. Pl. 91, 1781.

4
Piper micronesiacum Hosokawa, Trans. Nat. Hist. SOC.
Formosa 32: 387, 1942.

Kusaie, Mt. Keies, Mallens, Hosokawa 9498 (A,US, isotype)

This collection, i.e., type of P. micronesiacum Hos. was
referred to P, guahamense by Yuncker, and tentatively by mee It
differs in having several spikes in an axil and in being less

prominently puberulent,

Smith's excellent discussion of P. latifolium and its

A. Ce
allies (Jour. Arn. Arb. 24: 349-351, 1943) Suggested looking at
he Marquesas, Society»

severa xil.
rather variable, from glabrous to puberulent on petioles and

Herbarium, from ti, Rarotonga, Rurutu, and t
mostly studied either by h or by Yuncker, illustrates bea
well this variation, The Hosokawa 9498 collection fa :

within the range of variation of this material, though i
hard to compare in leaf shape, as the leaves are folded
ly pressed, It is puberulent, though less so than Grant_214

t is
and bad-

1966 Fosberg, Micronesian Plants 237

from Tahiti, The leaf blades seem slightly narrower than those
of some of the South Pacific specimens, and more strong
e from

from other
parts of its range, it will Wi plage on possible to separate
varieties or forms in this specie

PIPER HOSOKAWAE Fosberg, nom, nov.

Piper ae va Be Hosokawa, Trans, Nat. Hist.
oc a 28; “153 1938 (non Piper sa tansoae Hoso-
kawa 193

PB decumanum, so not much was recorded. These sheets are not
cited below,

e leaves are broadly ip yeadebeotta, tending slightly to
iol; to at least 27 x 15 cm, deeply cordate, oblique, basal
lobes sometimes overlapping, petiole about 3 cm long, winged part
way up; pistillate spike elongate, about 6 cm thick, ovaries con-
Spicuously drawn out to a neck at summit, stigmas 3, tomentulose,
Spreading, pointed

Endemic to the Palau Islands, so far as known,

ine Islands, Palau: Peliliu, Hosokawa 9226 (A, BISH,
isotype); i tpt ped Hosokawa 7098 (A, BISH).

Named for Prof, T. Hosokawa, of Kyushu —!" who has
devoted much attention to Micronesian Piperac

238 PHYTOLOGIA Vol. 13, no. &
PIPER METHYSTICUM Forst. f£.,De Pl. Esc. Oc. Austr. 76: 1786.

he name Piper oes has been almost universally
— to the widespread Pacific shrub with leaf-opposed

variously kava (Polynesia), yangona (Fiji) or oakaa gee
The root is also a pharmaceutical item in some western coun-

e existence of an earlier effectively Rte Piper
ue

the legitimacy of Foster's nam Linnaeus filius ake d
P. methysticum with a description, applying clearly to
species with axillary spikes. at the end of

the volume he says, "Pag. 91. Piper methysticum lege Piper
latifolium,'

Je W. Moore (Occ. Pap. sua i Mus, 10(19); 3-4, 1934)

dealt with this question. He to accept an author' s right
to correct his own errors and pa er regarding P. methysticum
L.f., thereby, as never having been published at t all, leaving
us free to use P, methysticum Forst, f

th (Jour, Arn. Arb. 24: 349, 1943) and Merrill

A. Smi
(Bot. rie Voy. 357, 1954) following him, considered that
Pp methysticum Lef. was in synonymy, therefore not validly pub-
li

This is certainly a practical and in this case, at leas
an acceptable solution. It is necessary, however, to pe "the
case in terms o

sticum Forst,

he earlier of the two names is regarded as vel
well as effectively, published, then ance r not the author “ts

tion, hence | non-acceptance of t atter Therefore, it i il
not validly published and cannot be einer as published at 4

1966 Fosberg, Micronesian Plants 239

in the sense of the Code. Hence it is not an earlier homonym

of P. methysticum Forst. f., which we may continue to use in

its traditional sense

PEPEROMIA R, & P.

PEPEROMIA MARIANNENSIS C.DC,, DC. Prodr. 16(1): 442, 1869.
Peperomia guamana C.DC., Phil. Jour. Sci.Bot. 9: 72, 1914,
Although there is a or ecsdcanrt range of variation i

number of characters in the Peperomia population of the Marianas,

b

there seems to be little Pipher bent between these variations,

n
on n the basis of opposite, sities leaves in the latter, vs. al-
ternate acute ones in P, guamana. Howe » the isotype sheet
£ P. guama u

2. mariannensis though Yuncker called it P. guamana, Stone 4713
has Opposite, mostly acute leaves

The conclusion, after examination of a good series of spe-
cimens, is that o only one species is represented na the Marianas,
Its correct name is P. mariannensis C.DC. Dr. B. Stone (ms,
1966) has quite independently machi at the dadetascen that
these two species should be combin

s Yuncker has pointed out, plants of this species from the
northern Marianas tend to be larger in stature than those from

Guam, ev
nothing else, I agree with Yuncker in not regarding it as merit-
ing taxonomic recognition,

Distinction from P kraemeri C.DC. is a more difficult prob-

cies. Glabrous plants from Palau referred by Yuncke

nha seem in P, kraemeri with no difficulty and are so
fisposed of here, Part of Fosberg 25869, from Urukthapel, Palau,
is appressed hirtellous and seems to belong to P. batedensts
the glabrous r

remainder to P, kraemeri.
PEPEROMIA MARIANNENSIS C. DC. £. MARIANNENSIS

Peperomia mariannensis C, DC., sensu stricto.

form is known from Agrigan, a Alamagan, Sarigan,

This
Saipan, Tinian, Agiguan, Rota, and Gu

20 PRI.TOL,0GC.1.4 Vol. 13, no. 4
PEPEROMIA MARIANNENSIS f£. SAIPANA (C. DC.) Fosberg, comb, nov.
Peperomia saipana C. DC.in Merr.,Phil.Jour.Sci.Bot.9: 72,1914.

Peperomia guamana var, saipana (C. DC.) Yuncker, Occ. Pap.
Bishop Mus. 14(2): 15, 1938.

A form with hirtellous stems occurs on Saipan and possibly
Tinian (ine oken wa 8023, cited by Yuncker as from Pagan, is really

has no discreet i range, as f, mariannensis also
occurs on eeipan and Tinia

C. de Candolle, in his original descriptions of P. saipana,
said soactii lis that it is glabrous, £ were the case,

presumably destroyed. | since he then used the agen pape!
for the hirtellous plant, we may, I think, safely assume that the
Fritz specimen that de Candolle Sokecthad was Spo g par?

PEPEROMIA PONAPENSIS C, DC., Bot. Jahrb. 56: 504, 1921.

Peperomia volkensii C. DC., op. cit. p. 503.

Peperomia gibbonsii C. DC., ope cite p. 504.

have been regarded as three species, all glabrous, have
been recognized from low elevations in the eastern Caroline am nd

ied specimens of P. ponapensis and B dolkensi differ only
in that those of P, vonuenett have the lea predom nantly

1966 Fosberg, Micronesian Plants 21

xamination of Yuncker's descriptions of the two show
that yt are essentially identical except that P. volkensii ho
the leaves predominantly opposite while P. ponapensis has them
predominantly alternate,

e gibbonsii has not been recollected since the original
“nace ma The characters cited as differentiating it from
ssi

S occur not infrequently among other ee but this char-
Bn is especially noticeable in the type speci of this spe-
cies." Unfortunately this specimen was probably ae with the
Berlin Herbarium during World War II.

ames n
simultaneously and, because P. ponapensis has been the most wide-
ly used name, it is here chosen for the combined species.

PEPEROMIA PONAPENSIS C,.DC, var. PONAPENSIS

Peperomia ponapensis C.DC. sensu stricto

Found in the eastern Carolines (Ponape, Pingelap and Kusaie)
and central to southern Marshalls (Lae, Ailinglapalap, i Pe
Jaluit and Ebon atolls). The records from the Marianas (Sa
gan and oe admitted by Yuncker are here referred to hey very

Similar P, mariannensis.

PEPEROMIA PONAPENSIS var. TRUKENSIS (Yuncker) Fosb. comb. nov.

Peperomia erukensis Yuncker, Occ. Pap. Bishop Museum 14:
23-24, 1938,

“ay around, It is here regarded as a variety of P. ponapensis.

NOTES ON NEW AND NOTEWORTHY PLANTS. XLIV
Harold N. Moldenke

BOUCHEA BOYACANA var. GLABRATA Moldenke, var. nov.

Haec varietas a forma typica speciei ramis ramulisque mimtis-
sime puberulis vel glabratis inflorescentiis minutissime puberulis
recedit,

This variety differs from the typical form of the species in
having its branches, branchlets, and leaves only most minutely
and obscurely nicroscopically pube erulent or glabrate one its ra-
chis and calyxes very minutely mesh er acne ae pube

The type of the variety was collected by Paul C. utoticechl ‘sink
J. Kenneth Wright (no. 6592) along roadsides on desertic steep
slopes, at an altitude of 2050 meters, in the vicinity of paar
heeoe between there and Sondorillo, in the provinc

Piura n r 196, and
the H. N. Moldenke herbarium at Plainfield, New Jersey. The col-
lectors describe the plant as an herb to 3/) of a meter tall,
with blue flowers

bee omen JAPONICA f. ALBIFLORA Moldenke, f.
aec forma a forma typica speciei corollis albis recedit.
this form differs from the typical form of the species in
having white coro.

The type of the form was collected by Ernest Henry Wilson
(no. 1044) at Kongo-san, in the province of Kogen, Korea, at 330
meters alt: altitude, on July 5, 1918, and is deposited in the United
7 National. Herbarium at Was ashington. The collector notes

one 8

in eee FRUTICOSUM f. BAHAMENSE (Millsp.) Moldenke, stat.

Cithare ion bahamense Millsp. in Britton, Bull. N. Y. Bot.
Gard, 3: » 1905.

LANTANA MINASENSIS var. LONGIBRACTEOLATA Moldenke,
Haec varietas a forma typica speciei bracteolis Mest ad z Me
elongatis recedit.
S variety differs from the typical form of the species in
its elongate. Hirde ts, the outermost being to 1 am. in length.
t f the variety was collected by Alberto Castella
’

as deposited in the H. N. Moldenke herbarium at Plainfield, New
ersey

e

STACHYTARPHETA CAJAMARCENSIS Nodderike, SP. nov.

Suffrutex, ramis acute tetragonis perspicue alato-marginatis

dense griseo-pubescentibus, foliis oppositis ellipticis vel 1am
22

it —ccsmennncintiiniitiniseanediiaianailtai ccs

1966 Moldenke, New and Noteworthy Plants 2h3

ceolato-ellipticis acutis, ad basin in petiolum longe acuminatis,
utrinque dense puberulis; inflorescentiis terminalibus solitariis
usque ad 15 cm. longis densissime miltifloris percrassis robustis

eae; stems and branches acutely tetragonal, conspicuously
alate-margined, densely grayish- or browmish-pubescent throughout;
nodes abbreviated, 1~-3 cm. long, the lowest to 7 cm.; leaves de-

na
short petiole, densely seca peas Bh 6 bi surfaces; mid~
rib slender, flat above, slightly eath; secondar-
ies slender, numerous, close, 7-~10 per aa inotae ans ’
ominul 3 inflores

before and during anthesis, less so after anthesis, pA
Sutulous; calyx aylintels, about t 15 mm. long, densely puberulent,
hireutulous on the 5 ribs, the rim sharply 5-toothed; corolla
ae rateriform, its tube ” about 2 cm. long, the limb * about 1 ca.

Hutchison and J, Kenneth weight tak 3645) in . oh at the base
of a granite cliff, Rio Chamaya, 166 km. east of Olmos, at an al-
titude of 700 meters, in the province of Jaen, Cajamarca, Peru,
on January 19, 196), and is deposited in the herbarium of the
niversity of California at Berkeley.

ADDITIONAL NOTES ON THE GENUS VERBENA. II
Harold N. Moldenke

VERBENA [Dorst.] L
Addi ti Shot Ry emended biblio ography: A. ‘eg Spt om parle
Narr. Exped. China Seas & Japan 2: 316. 18565 Bot aor
ke 53h, 1879; Aitch,, Journ. Linn. Soc. Lond. Bot. 18: "ae. 18 18805"
N. L. Britton, N. J. Geol. Surv Final Po State Geol. 2 Oye
194, 612+ 18895 "urs , How to 41d Fls.,
314-316, ic in: : oe pl. 135. 1896; Stack, Wild aFis.

2hy PHYTOLOGIA Vol. 13, no.

39351, 00, & 410. 1909; W. J. Beal, Mich. Agr. Coll. Exp.
. 260: 153, fig. 153. 1910; Je Matsumura, Ind. Pl. Jap.
2 2 (aye 53 53h. 1912; W. J. Beal, Mich. Agr. Coll. Exp. Sta. Bull.

19395
bert, How to Know Spring Fls. “10, Megas ‘on & 298. anty “Sonik.
Wild Flow. Dossier 1: 20, fig. 3. 1 951g Gorsch., Fl. URSS 19:
694. 1953; Kitamura, Fl. "afghan. 327. 560; Dyer, Verdoorn, Codd,

9
o>, Pleo & 210 king, Ex-

cerpt. Bot. A8: 226-227 (1965) and A9: 364—~-367. 1965; Buchhein,
Huntia 2: 38. 1965; Moldenke, Phytologia 13: 179-218. 1966;
Shimers, Sida 2: 266. 1966; Moldenke, Résumé Suppl. 13: 1—3, 6,
& 7. 1966; Cody, Ind. Sem. Bot. Gard. Ottawa 1966: 19. 1966;

ig, Taxon 15: 30. 1966; Gaiser, Surv. Vasc. Pl. Laabto
Co. 100-101. 1966.

VERBENA ABRAMSI Moldenke
Additional bibliography: Hocking, Excerpt. Bot. A9: 367. 19655
Moldenke, Phytologia 13: 181. 1966,

VERBENA AMBROSIFOLIA

Additional bibliography: Mold enke, Piytoloeia 13: 181—182.
1966; Moldenke, Résumé Suppl. 13: 2.

Crutchfield Soa this plant Seiki in "sandy gravelly soil
at edge of road" in New Mexico.

Additional. sae, NEW MEXICO: Chaves Co.: J. Rs Crutch-
field 87 (Ld). Pigg

AMBROSIFOLIA f. EGLANDULOSA Perry
dditional babbsegeigars Moldenke, Pertrebth 13: 182. 1966;
Notdenke, Résumé Suppl. 13: 2. 1966.

VERBENA AMO) Paxt. 82. 19663
Additional bibliography: Moldenke, Phytologia 13: 182.
Moldenke, Résumé Suppl. 13: 2 & 6, 1966.

ANDRIEUXII S
we bahiieeregees Moldenke, Phytologia 13: 181 & 182.

Addit aut orcs 182. 19665
biblio : Moldenke, Phytolo 13: °
Moldenke, Résumé Logrnty 3. 1966. e

VERBENA BIPINNATIFIDA Nutt,
193i titional bibliograpty: Uoldenke, Phytologia 13: 180, 161,
Recent collectors have found this plant growing in red or in

1966 Moldenke, Notes on Verbena 2h5

loose black soil, loose sand, sandy silt, clay.

Additional citations: TEXAS: Bee ne ’Rios & Cavazos 206 (Ld).
Brown Co.: L. Horner 20 (ld). Coryell Cos: Baise 13 (Id). Duval
Co.: Gonzales-Arro (Ld) ; Fe majarde 62 ( aa, 2 E. F. Gutdier-

rez 56 (Ld); ‘Tlaguno zat BI (14). Cost ee & Dohnke 151
tai La Satie Ose Co.: =p A. ea 6 (ld). Live Oak Co.: Rios
& Cavazos 222 (Ld). Medina Co.: Barrera & Laurel 110 (1d).

VERBENA BONARIENSIS L L.
i eee bibliography: Moldenke, Phytologia 13: 18h—185,

Reed found this plant fruiting in Novesber.
SS a citations: ALABAMA: Crenshaw Co.: C. T. Reed 2073
Ld

VERBENA BRACTEATA Lag. & Rodr.

Additional bibliography: Muenscher, Cornell Agr. Exp. Sta.
Bull. 635 [Weeds N. Y.], pr. 2, 10 & 387. 19995 Hocking, Excerpt.
Bot. A9: 367. 1965; Moldenke, Phytologia 13: 185
Résumé Suppl, 13: 1, 1966; Cody, Ind. Sem. Bot. Gard,

1966: 19. 1966; Gaiser, Surv, Vasc. Pl, Lambton Co. 100, 1966.

Recent collectors have found this plant growing in brownish

=]

1 on dry and sunny prairies near the edge
of a natural lake in rolling grasslands in New Mexico. The
flowers on J. R. Crutchfield 423 are described as "

Additional oi citations: NEW MEXICO: Colfax Co.: Lucas 164 (Ld).
San Miguel Co,: J. Re. Crutchfield 423 (1d).

mer Vell.
Additional b iography: Moldenke, Phytologia 13: 185-~186.
19665 Moldenke, "ae Suppl. 13: 3. "1966.
VERBENA CAMERONENSIS L. I. Davis
1 96g citional bibliography: Moldenke, Phytologia 13: 186 & 192.

VERBENA CANADENSIS (L.) Britton
Additional bibliography: Quick, Wild Fls. cette States 373 &
518. 19395 M. J. Cuthbert , How to Know Sp: fig.

ring Fls
2976. 19432 Hoe ocking, Excerpt. Bot. A9: 367. 1965; Wolaeake,
Phytologia'13: 186~2187, aye
Addi ti illustrations: M. J. Cuthbert, How to Know Spring
Fls. fig, 297, 19)3,

Buchoz (1783) colored illustration of this species,
Cited in my Ses Peg is unnumbered in the New York Botani-
3

cal Garden lib of this work. The Co 2
Sona) Crutchtiers 30903 are described as “lavenier® and the
lant is Said to ha dy bank from a woody

Additional citations: KANSAS: Labette Co.: Lerew Kan.2-180
(ld), TEXAS: Henderson Co.: Correll, Correll, & “& Crutchfield

2h6 PHYTOLOGIA Vol. 13, no.
30903 (1d).

0B XK
Additional biblio graphy: Moldenke, heer oe 13: 187-188 &
216. 1966; Moldenke, Résumé Suppl. 13: 1966.

e ALBIFLORA Moldenke
phy: Hocking, Excerpt. Bot. A9: 367. 1965;

gomctem rt
Additio. bibli Liogra
Moldenke, eniberts 13: 187. 1966.

VERBENA ENS var Sor mgmt (Scheele) Perry
Additional bibliography: M ldenke, Phytologia 13: 187-188.
1966; Moldenke, Résumé Suppl. ris} 7. 1966.

VERBENA CANIUENS
Additional said unerabtips Moldenke, Phytologia 13: 451. 1965;
Angely, Bibl. Veg. Paran. 196. 196).

LINA L.
Additional bibliography: Moldenke, Phytologia 13: 188. 1966.
ae citations: HONDURAS: Cop4n: Molina R. 12892 (W—

VERBENA CILIATA Benth, 89. 1966
Additional biblic graphy: Moldenke, Phytologia 13: 189. ;
Moldenke, Résumé Suppl. 13: axe
The 81 and Rios & Cavazos 206 me 222, distributed as Ve
ciliata, are actually V. bipinnatifida Nutt., Anda 8 is V. ceili
ata var. longidentata Perry, and L. R. Sele 25 is V. tenuisects
Briq.

Le)
ge

VERBENA CILIATA var. LONGIDENTATA P
Additional synonymy: Verbena ciliata var. longedentata Perry,

qeittitional bibliography: Moldenke, Phytologia 13: 189--190-
Recent pelle have found this plant growing in pe —

sandy or fine sandy silt, and caliche, Material has been mi as
identified and distributed in herbaria under the name Vv. clover

— orvecanne ais the Barrera & Laurel 110, Gam Ganbet eres
ohnke 151, Gonzalez—Arroyo 99, F. Guajardo 62, Ee F, Gutserr=
%, and I, A» Martines rtines 66, distributed as V. ciliata 2 var. longi
Gentata, actually V. bipinnatifida Nutt., while 7. Brunt 12
is V. ciliata var, pubera (Greene) Perry.

Additional citations: TEXAS: Dimmit Co.: Anda 8 (1d)j Chavees
Flores, & Hein 7 (Ld); Guerra, Garcia, & Garcia 32 (Id).

Wells Co.: Sees 3. Ld). Webb Co.: Dickey 137 “137 (Ld). Zapata 00!
F, Guajardo ardo 2 (Id gp : (

1966 Moldenke, Notes on Verbena 2h7

VERBENA CILIATA var. PUBERA (Greene) Perry

Additional bibliography: Moldenke, Phytologia atte 1s ee

Recent collectors have found this plant growing in 1 oie
loose sand. Material has ree Ee hiigetts” and Ss aeettated | in
herbaria as V. qua

Additional citations: Serie Jin ‘bie Co.: L. Moran 10 (Ld).
Maverick Co,: T. Bruni 12 (Ld). ce me

CLOVERAE Moldenke
Additional synonymy: Verbena cloveri var. cloveri Moldenke,
in herb. Verbena cloveri var. iilacima Moldenke, in herb
Additional bib: bibliography: Moldenke, P ‘ Phytologia 13: 190191.

Recent collectors have found this t growing in gravel or
in loose red sand, Material has epee misidentified and distribu-
ted in in herbaria as V. halei Small. On the other hand, the F.
Guajardo 42, distributed as V. cloverae, is actually V. ciliata
var. lon longidentata Perry, while R. A. Leal 30 is Vv. licata Greene.

Additional citations: TEXAS: Duval Co. Co.: Cc. Cantu 55 (Ld). Jim
Hoge Co.: R. Bruno )1 41 (Id); Sosa 32 (Ld). Kleberg Co: Ri Rios &
Cavazos (Id). La Salle Coe: A A. Castro 55 (1d); M. de J. Solis
122 (Ld). Starr Co,: R. Me Navarro 98 (Id). Webb Co.: Arzola ola 143
via) ol snore Garcia, & Garcia 2h (Id). Zapata Co.: Flores &

lie (14); F. Guajardo 3 Bi (Ld).

CORYMBOSA Rufz
Additional bith iograrhy: “Wondene, Phytologia 13: 191. 1966;
Moldenke, Résumé Suppl. 13: 3. 1966,

VERBENA DEAMIT Mol
Additional Bibliogrartys Bolderts ators 13: 191. 1966;
Gaiser, Surv. Vasc. Pl. Lambton Co. 400,

Addit ne 13: 186 & 191
onal bibliogra Moldenke, lee i sd
192. 1966; Moldenke, sic Suppl. 13: 2. 1966

VERBENA DEMISSA Mol,

apne bibliography: Moldenke, Phytologia 12: wa « 1965.
Recen sparsely

hillsides, at 3300 meters altitude, in flower and fruit in a

sapguct tonal citations: ECUADOR: Caflar: J&tiva & Epling 260

pore ey DISSECT Willd,
tional bibliography: Moldenke, Phytologia 13: 192--19k, 213,
Tr & 217, 1966; Moldenke, Résumé Suppl. 13: 3. 1966.
Ballets 231 [Herb. Inst, Migoel Lillo 31690] cited by me
ye vious ly as this species seams, to be V. selloi ro hee
ts leaves do resemble saes of Eger, te dissecta otas but its calyx

21,8 PHYTOLOGIA Vol. 13, no.

and bractlets more closely resemble those of V. selloi. Possibly
4+ may be a hybrid between the two species.

VERBENA DISSECTA f. ALBA Moldenke
Additional bibliography: Moldenke, Phytologia 13: 19h. 1966;
Moldenke, Résumé Suppl. 13: 3. 1966.

VERBENA ELEGANS H.B.K
Additional bib ibliography: mangenie, Phytologia 13: 19l--195.
1966; Moldenke, Résumé Suppl. 13: 2. 1966.

VERBENA ELEGANS var. ASPERATA Perry
Additional bibliography: Moldenke, Phytologia 13: 195. 1966;
Moldenke, Résumé Suppl. 13: 2. 1966.

XVERBENA ENGELMANNII Mo noe
ea bibliography - Britton, N. J. Geo
tate Geol. 2 @)s ate 1889; lioldenke, pion t = 13:
195. = 3966} Gaiser, Surv. Vasc. Pl. Lambton Co. 100. 1
rid is recorded ty "Britton (1889) from “Near Riegels~
wuie® in Warren ove New Jersey. Gaiser (1966) says “only om
Walpole rey oe in two locations.......Both popes siesta are &-

he island", Lambton County, Ontario

camabeaeaae mere sera 96-197

0! bibliography: Moldenke, Pde ag a ph ee: 916

eee Moldenke, grr or 232 2. i

ies The flowers on Crutchfiel d 243 are described as "light-blue"
plant was font growing on a dry rocky hillside.

Py ripe citations: ARIZONA: Mohave Co.: J. Re Crutchfield

VERBENA GOODDINGII var. NEPETIFOLIA Ti
Additional bibliography: — Petiscks 13: 196-—197-
1966; Moldenke, Résumé Suppl. 13: 2. 1966.

VERBENA HALEI Small

sAAdies com bibliography: Moldenke, Phytologia 13: 197--199 &
Recent collectors report this plant growing in sandy silt,
sandy loam silt, black clay, fine sandy silt of the Claiborne
Group, and in open fields. Material has been misi ited ie

other hand, the R. Bruno )] and Sosa yg
are actually V. cloverae Moldenke. Malacara P. & Gutierrez Fe Fe =
is a mixture with ¥. plicata Greene,

Additional citations: MISSOURI: Saint Louis: Mublenbachs ie
(Ac), a3 (Ac), 1607 (Ac), TEXAS: Bee Co.: Bice & cavaz0s 202

1d). “Bexar Co.1 Che cheves, Heese, & Hein 96 lage rg immit Cos?
3787 (1d). ary Ae pis bom & Cavazos 181 om tas" Jim pe Coe!

1966 Moldenke, Notes on Verbena 2h9
P. & Gutierrez F. 21, in part (ld). Jim Wells Co,: P.

Malacara P
Cabrera - 101 (1a). mir Oak beck Rios & od 221 (1d).

ae Be Ee Gut Co.: Rios & Cavazos 165 (Ld). Webb Co.: Cisneros

2h (Id); J. Gonzalez li (1d); J. M. M. Hughes lj) (Ld); S. Webber 12

(Id). Zapata Co.: F. Guajardo 3 39 (Ld). Zavala Co.: Moore, Car-
).

Fasco, Gongora, & McCart 5997 (Id

HASTATA L.
Additional biblio ography: N. L. Britton, N. J. Geol. Surv.
Final Rep. State Geol. 2 (1): 19h. 1889; Mrs. W. S. Dana, How to
Know Wild Fls., rev, ed., eee 365, & 371, pl. 135. aes
Stack, Wild Fls. 349--351, 1,00, . 1909; W. J.
Agr. Coll. Exp. Sta, Bull. 260: 153, fie 183 (1920) and 251, "ed.

2, 115, fig. 1% 1915; Sievers, U. S. Dept. Agr. Misc. Publ. 77:
15, fig. 2h. 1930; Quick, Wild As. ue States 5TIn= 312, aucay
& 518, fig. 190. 19395 Muenscher, Cornell A r, Exp. Sta.

635 [Weeds N. Y.], pr. 2, 10 & 387, 1939; Oswald, Wild FL

Dossier 1: 20, fi » Pr. 1951; Hoc , Exce erpt. Bot. AS: 206 “(2965)

and A9: 367. 1965; F. H, Montgomery, Native Wild Pl., pr. 2,

fig. 209. 1965; Cody, Ind. Sem, Bot. Gard, Ottawa 1966: 19. "19663

Moldenke, Phytologia 13: 199-200. 1966; Gaiser, Surv. Vasc. Pl.
on Co. 100. 1966.

Additional illustrations: - Dana, How to Know Wild
Fis., rev. ed., 315, pl. 1g. 1 3965 v. "J. Beal, Mich. Paha Coll.
Exp, Sta, Bull. 260: fig. 153 (1910) and a ed. 2 fig. 17h.
19155 Sievers, U.S, Dept. Agr. Misc. Publ. 77: fig. 2h. 1930;

Fils.

» Wild North. States 375 fi. 130, ety Oswald, Wild
Flow. Dossier 1: 20, fig. 3. 1951; F. H. Montgomery, Native Wild
Ple, pr. 2, fig. 209. 1965.

VERBENA HATSCHBACHT Mol
Additional biksegrante | Moldenke, Phytologia 9: 284-285.
19635 Hocking, Excerpt. Bot. A8: 226. 1965.

VERBENA HAYEXIT vf
Addi tional Gindieetankhs 3 Hoes Excerpt. Bot. AS: 226. 19653;
Moldenke, Phytologia 13: 200, 1966.

HERTERI Moldenke
Additional bibliography: Hocking, Excerpt. Bot. A8: 226. 1965;
mee Pigisiesla tar ios 1966; ; Moldenke, Résumé Suppl. 13: 3.

VERBENA HIRTA Spre:
Additional nitdingrentels Hocking, Excerpt. Bot. AB: 226. 1965;

Moldenke, Phytologia 13: 200—201. 1966.
Smith’ and his seaccietns found this plant growing in thickets,
— low woods, forests, gallery forests, dry fields, and dry

°PeN ground, on Sieg along boggy brooks, and ruderal., Smith &

250 PHYTOLOGIA Vol. 13, no.

Klein 14936 represents the very hairy form of the species. The
Tryons ; found the species in shrubby grassland and sedgeland with
local elfin-woods dominated by granitic rocks. aa: & = Bye
6713 had "blue-purple" flowers and was "woody at bas
Additional citations: BRAZIL: Rio de Janeiro: 14 107 1 Tipe
Cent. Pesq. Florest. 345] (Ac); Tryon & Tryon 6713 aD) :
on A. Castellanos 2675 (Herb. © Cent. Pesq. oe horest. an
Ac).

var. S Dusén
Additional cranes: Hocking, Excerpt. Bot. A8: 226. 19655
Moldenke, Phytologia 13: 201. 1966.

VERBENA HISPIDA Rufz & P
sAditional b babldibgiaieys Hocking, Excerpt. Bot. A8: 226. 1965;
Moldenke, Phytologia 13: 201. 1966.

VERBENA HOOKERIANA (Covas & Schnack) Moldenke
Additional bibliography: Hocking, Excerpt. Bot. A8: 226. 1965;
Mold enke, Phytologia 13: 201. 1966.

EIMIFUSA Cham,
Additional bibliography: Hocking, Excerpt. Bot. AB: 226. 19655
Moldenke, Phytologia 133 ol 202. 1966.

XVERBENA HYBRIDA Vos

Additional patthoipheatiy: McCurdy, Book Gard. Fls. 72--73, 30,
& 311. 1931; Hocking, Excerpt. Bot. AS: 226. 1965; Moldenke, Phy-
tologia 13: 202-203, 1966; Moldenke, Résumé Suppl. 13: 7. 1966+

xVERBENA ILLICITA Moldenke 3 7. 19653
Additional bibliography: Hocking, Excerpt. Bot. AS: 227.
Moldenke, Phytologia 13: 203. 1966.

VERBENA INCISA H
Addi tional, herpes Angely, Bibl. Veg. Paran. 136. og Fe
=, Excerpt, Bot. ey oer oT 18s; Moldenke, Phytologia 13: 207
INTERCEDEN,
pleat 2 19653

Additional bibliography: Hocking, Excerpt. Bot. A8: 227.
Moldenke, Phytologia 13: 20320). 1966.

Addi Loney Hocking, Excerpt. Bohs 2 :
woldenke, go epee 13: 20h, 1966; Moldenke, Résum Suppl. 134

eters (L.) Brig. A: 227. 1965
Additional liegraphy: Hocking, Excerpt. Bot. :
Moldenke, Phytologia 13: 204 & 213. 1966.

1966 Moldenke, Notes on Verbena 251

The Bailetti 231 (Herb. Inst. Miguel Lillo 31690), distributed
as V. erinoides Lam. and c ited by me previously as V. diss dissecta
mid. » 18 probably ¥V. selloi Spreng.

VERBENA LANDBECKI R.
Additional bibliography | lordenke, Bi tse snes li: Pe 1965.
Additional citations: CHILE: Aco : Landbeck s.n. [Choapa;

Macbride photos 17125; Herb. Mus. tis. Hist. Nat. Guile 54,672]
(W——photo of isotype). Coquimbo: Marticorena & Matthei 409 (Z).
VERBENA LASIOSTACHYS Link

Additional bibliography: Hocking, Bot. A8: 227 (1965)
and A9: 364, 1965; Moldenke, Eeeadeies 13: “Sok. 1966.
VERBENA LASIOSTACHYS f, ALBIFLORA

Additional bibliography: etic, “iytalogia 9: 65-166.
196k; Hocking, Excerpt, Bot. A9: 364. 1965.

VERBENA LIPOZYGIOIDES Wal

» Phyto
: the 1965 reference given ab ove this species is referred to

lipozygioides Walp." through a typographic error

anal LITORALIS H
Additional Saki bemioig ine ocking, Excerpt. Bot. A9: a. 1965;
Tha a pivtologia 13: te 1966; Moldenke, Résumé Suppl.
Jétiva & Epling found this plant growing in secondary forests,
Material has been mini danti flag and dis tributed in herbaria as V,
caro L. and even as S a nigriflora Epl. in the cea
Turezaninow (1863) has this to bo

exaspe
panveclato-subulatis calycem aequantibus; nuculis striatis laevi-
insula Sandwicensi Wahoo lecta. Species V. angustifoliae

errini
instructa: Specimen a Galipan, prov. Caracasanae Funk No. 325, e
Prope Cumanacoa Funk No. Moz Sh ot ¢ et Galeoties td Nos 35695 fuculae
SJusdem magnitudinis, apice par iosae, folia latiora et an-
Sex 1or@+ Haec forte ad V. 1 Stags teaaea Tear. ak Gel. apecten’...-
Sed specimina Funkiana e Caripe sub No, 637 et Lindeniamm e vic-

252 PHYTOLOGIA Vol. 13, no. §

iniis Meridae panicula gulto mimus ramosa et oligostachya gauden-
tia, fere medium tenent inter hanc et V. fl omnes ad
speciem unicam reducendae. Planta in schedula dicitur annua.
Stirps Novo~Grenadensis e prov. Ocana, Schlim, No. 564 speciem
valde propinquam indicat at caule e een hedula fruticoso et foliis

pagina superiore integer: S pauc npnadoope hk receden-
te." He places the discage in the fositn Leptosta
Additional citations: COLOMBIA: Valle del ts Eguet

ras 8219 (W--251148), 8478 (W—-21;511)5) ECUADOR: tot
Jativa & Epling 55 (La—l9643).

VERBENA LOBATA Vell.
Additional bibliography: peleings patente Bot. A9: 365. 1965;
Moldenke, cin gers oe = 0620704
tio : BRAZIL: Rio de Janeiro: Herb. Inst. 0s-
waldo Cruz 651 “u-216b227).

VERBENA LONGIFOLIA Mart. & opener
Additional bibliogra xcerpt. Bot. aoe — en
a ery .. Phytologia 13 13: 207. » 19660 Pes yal Résumé

VERBENA INA R. A. Phil.
Additional bibliography: Hocking, Excerpt. Bot. A9: 365+ 1965}
pein Role deen J 13: 209. 1
The collection cited below was previously cited by me as V.
about in error,
~~ Additdonal citations: URUGUAY: A. Castellanos s.n. [23. I.
1947; Herb, Inst. Miguel Lillo 15126) (N).

965
Additional tabi ihgreptg Hocking, Excerpt. Bot. A9: 365. 19635
Moldenke, Phytologia 13: 209. 1966.

‘A MICROPHYLLA H.B.K.
ditional bibliography: Hocking, Excerpt. Bot. A9: 365. 1965

: 209. 1966.
en "t737 cited by me as V. microphylla in 19L8 anc
V. parodii (Covas

as V, tenera in 196] is actually V & Schnack)
Moldenke,
VERBENA MINUTIFLORA Briq.

Additional bibliography: , Bibl. Veg. Paran. 196. 196k)

Moldenke, Phytologia 13: 210. 1966.

VERBENA MONTEVIDENSIS Spreng.

Additional bibliography: Hocking, Excerpt. Bot. A9: 365+ 19653
Moldenke, Phytologia 13: 210—211. 1966.

Additional citations: BRAZIL: Santa Catarina: A. Castellanos
24880 [Herb. Cent. Pesq. Florest. 3516] (Ac).

1966 Moldenke, Notes on Verbena 253

MULTIGLANDULOSA Mold
Additional banhtoreniy Moldenke, Phytologia 13: 211, 1966;
Moldenke, Résumé Suppl. 13: 3. 1966.

VERBENA OFFICINALIS L,
Additional and emended bibliography: A. Gray in M. C.
J 18

7 Y
Excerpt. Bot. A9: 365, 19653 Moldenke, -Phyeetoct - 22-213,

agititional citations: GERMANY: ener & Degener ener 2921 (Ac).
er & Degener

RETE: Degener & Degener 29389 (Ac FORMOSA: Degener
20585 (Ac).

VERBENA PARODII (Covas & Schnack) Mold
: oeeetional bibliography: Moldenke, ay eaiogin 13: 213-21).

The JUrgensen 1737, cited below, was erroneously cited by me
as V. microphylla H.B.K. in my Alph. List Cit. 2: 599 (1948) and
as V, tenera S Spreng. in Phytalogta 10: 148 (1964) and li: 275
(2965), whi while Venturi 5267 was incorrectly cited as V. tenera ver.

iflora Kuntze in P logia 11: 276 (1965).

The flowers are de avila "white" on Venturi 5267 and as
"rose=blue" on JUrgensen 1737. Venturi found the species in
flower in September at 700 meters altitude,

Additional Split gee Seerigre aehemense en 1737
pert Inst, Miguel Lillo 31472; Herb. 0 11009) (, (N, Ug, W—
74768) . Nee Venturd 5267 (be nisss18, F159 167) «

VERBENA PERENNIS
Additonal bibliography: eset Phytologia 13: 21). 1966.
The flowers on Correll rrell 30882 are described as

"purple" and the iar wae ton growing in rocky soil on small
un

ogititiona citations: TEXAS: Pecos Co.: Correll & Correll

Addit rt 13: 21y--215
Additional bibLiogrephys Moldenke, eae 8 215.
1966; Gaiser, Surv. Vasc. Pl. Lambton 1966.

PHLOGIFLORA
Additional Ciwiacereeias J. Matsumra, Ind, Pl. Jap. 2 (2): 534.
1912; Moldenke, Phytologia 13: 216. 1966.
by The recone of this species being cultivated _ sg as given
atsumura (1912) probably refers to xV. rida Voss.
probably, amietoratne

Sta ned: Sellow 8.Ne
(Maebride photos 34351] (W—photo of type).

25h PHYTOLOGIA Vol. 13, no. k

PLATENSIS Spre
tional bibliogr: Excerpt. Bot. A9: 366. 1965;

Addi ata ocking,
Moldenke, Phytologia 1 13: fait 1966.

VERBENA PLICATA G:
Additional eB adeno Verbena placata Greene, in
Additional bibliography: Hocking, A ack. Bot. oa gre 1965;
Moldenke, fa betes 13: 217. 1966.
Recent collectors have found this plant growing in loose sand,
%

a transposition of labels) as Larrea tridentata Coville in the

Z llaceae., On the other hand, the Rios & Gavexce . 28 & ca
uted as V. plicata, are V. halei Small.

Gutierres F. ala and Rios & Cavazos 233 | are mixtures aoe 7 ieee

Additional citations: TEXAS: Duval Co.: N. Castillo 6 (Id);
aes ot & Gutierrez F. 35 (Ld). Hidalgo Co.: R. As]

Ld). Jim Hogg Co.: Malacara P. & Gutierrez F. 21, in part art (Ld);
a & pele 102 (Id). Jim Wale Coa We castitio 27 (La, 1A).
Ta Salle Co.: Gonzalez & Quesada 4 (Ld). McMullen Co.: fies &
Cavazos 233, in part (Id). Starr Co.: Rios & Cavazos 52
Webb Co.: J. tesa os 26 (Ld); Guerra, Garcia, & Garcia 23 Mid);
Rios & Cavazos 33 (Id); S. Webber 20 (Ld). oaks Co.: M. de Je
Solis 106 (Id). MEXICO: Tamaulipas: F. A. Barkley 1432) (Id).

VERBENA PORRIGENS R,
Additional bi sbhacientins Woldenke, Phytologia 11: 77. 1965-
set a citations: CHILE: Aconcagua: Landbeck s.n, [near

Quilimar, Dec. 1861; Herb. Mus. Nac. Hist. Nat. Chile 5729;

Macbride bared 1741] (W--photo of isotype).

VERBENA PULCHELLA Sweet
Additional synonymy: Glandularia 2 pulchel checagrre' Troncoso,
Darwiniana 13: 73. 196k Glandularia ella var. p ma

Additi ography: Troncoso eon
473-76, fig. 2. 1964; Moldenke, Phytologia 11: 77 ti965) and
13: 192. 1966; Shinners, Sida 2: 266. 1

Additional illustrations: Troncoso, Darwiniana 13: h7h, fige

Troncoso (1964) describes this species as follows: "Planta
sufruticosa en la base, de tallos generalmente muy @x~

eo
inferior sobre el borde y el nervio m epi
» hipofilo comfinmente con puntos sekiiediibs ieee “Tente),

1966 Moldenke, Notes on Verbena 255

margen revoluto (principalmente en material de herbario), In-
florescencia solitaria en ramitas terminales ergui

~negruzca, reticulado-estriada, la ventral, blanco verrucosa
— figura), pericarpio prolongado y ensanchado en la base, ésta
wera,"

She notes "Habita en campos altos y arenosos del S de Brasil,
Y, centro y litoral argentino. Es usa especie muy variable!
She cites from Buenos Aires Burkart 23908 (Si) and 8490 (Si) and
cultivated in Buenos Aires Herb. Bentham s.n. [Soc. Hort. London,
20-VII-1829] (K), and from Uruguay Felippone 3147 (Si).

1. Plants ascending or prostrate, rooting only at the base.

2. Bractlets lanceolate-subulate, strigose-hirsutulous; calyx-
teeth subulate; anther-appendages cylindric, slightly ex-
serted from the corolla=-throat Ve pulchella.

2a. Bractlets oval, "pestafiosas" » subglabrous; calyx-teeth a-
cute, apiculate, short; anther-appendages club-shaped,
Plainly exserted fromthe corolla-throat

+e Se ees puichella var. clavellata.
la. Plants creeping, in general low, muc shorter and more slen-
der V. pulc

$366 Nb aed hi var ilior.
ver, after examining an isotype of her var. gracilior, I
find this to be typical V, selloi Spreng. Herter 1 °

f. alba Moldenke but seems definitely to represent ¥. pio
instead. The H. M, Smith 5, cited below, was mene cited by
me as V. tenera Spreng. in Phytologia 11: 275 (1965) and was
originally distributed as V. tenuisecta Briq. Troncoso (1964)
cites Rodrigo 2339 as V. dissecta, but the New York Botanical
Garden specimen of this collection, at least, is definitely v.
chella, Rosengurtt B.992, which I previously cited a V. pul-
chella, Proves to be the type collection of V. subincana (Tron-~
Shinn

ers,
Rambo 4891), is described by the collector as prostrate, with

violet flowers, growing in sandy places, Herter encountered the

Species at 2 meters altitude. The Boffa 330, T. Meyer 12290, and

256 PHYTOLOGIA Vol. 13, no.

Rambo bo 48914, cited below, were previously serspernal cited by me
as as Ve Gissecta Willd. On the other hand, 11i 105 &
137, ). previously cited by me as v. prichella, ns to be Ve V. dis-
secta instead. Similarly, Herb. Herter 83911, Herter 181g, | and
. 1. Lima s.n. [II .19h6; ee Rio Jan. 55h66), previously cited
me subincana, but Herb. Herter

der V, dissecta and V. selloi have the name V. pulchella given as
a of V. tenera on tosis labels ina very old handwriting,

but I do not think tha that this reduction is valid.

Additional citations: BRAZIL: Rio Grande do Sul: Rambo )891)
(N). URUGUAY: Herter 1805 [Herb. Herter 96556] (N, Z)3 He es
Smith 5 (W- 1193661). ARGENTINA: Buenos Aires: Boffa 330 ( (N).
Salta: | T. Meyer 12290 (¥).

HELLA f. LATILOBATA ee

Additional bibliography: Moldenke logia 11: 26. 1963
Troncoso, Darwiniana 13: é8, 470, & * Be LBe. 1964; Moldenke,
Phytologia 13: 192. 1

Troncoso (196)) redibes this taxon to synonymy under V. dis-
secta Willd., but I do not agree. Its characters are much
closer to those of V. pulchella if I am interpreting this spe-
cies and V, dissecta correctly.

VERBENA PULCHRA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 478. 19655
Pesquis. Bot. 21: 51 & [60]. 1965.
nate (1965) cites Herb. Anchieta 29682 [Henz s.n.) from Rio
Grande do Sul. He describes the plant as "Very vigo rous, erect,
ntly more than 1 m high", wert hy in "Wet, densely and
vegetated Campos". He notes "Only one Hindplace in the
northeastern Lowlands" of Rio Grande do Sul, and gives th
eral distribution of the species as "S, Paulo to Misiones and
Uruguay".

VERBENA PUMILA Rydb.

Additional bibliography: Moldenke, eons Abstr. 45: dig

1965 Melgenke, esp Suppl. 12: 1. 1 Hocking, Excerp
Bot. A9: 366. 1965; Molderke, Phytologia 41: 478, igo,

(1965) and 13: 186. 1966, Moldenke, Résumé Suppl. 13: 2. 1966.
Bruni this Dp. rowing in rocky me in Nuevo
Leén; others have found it in red loamy soil, "Jos,

nock, & Turner refer to it as a "low infrequent annual” with
"whitish flowers" (their collection may, eehee. represent ++
albida Moldenke; the base of the cocci in this collection is
wide-flering, @ puzzling character occasionally seen in speci-

1966 Moldenke, Notes on Verbena 257

mens tentatively regarded and cited as v. punils) The J. A. de
ja Crug s.n. [12.IV.1961], distributed as V V. pumila, is actually
vy. cameronensis L. I. Davis.

(Id); MeCart 6079 (14). Frio Co.: Lucas, Painter, & Bark)
14213 (Ld); Painter, Lucas, & Barkley ey 14201 il (Id). La Salle Co,:

A. = n ape Ueret2B9el. Llano Co.: Rose-Innes & Warnock 793
Ta). : Barrera & Laurel 11] (1d). Val Verde Coe:

Barkley 11,261 (Id); Bruni & P Pena 38 (Au--232622, Id); Oatman &
Rowlett ote af Feb, 196) (Au—227525). San Luis Potosf: a
7536 (Ip).

VERBENA PUMILA f. ALBIDA Moldenke

Additional bibli cevanias "eeldaiie: Biol. Abstr. 45: 2772. 196k;
Moldenke, Phytologia 11: ne 1965.

Possibly Parks, Warnock, & Turner 236, from Val Verde County,
Texas, and described as ha having Wwhitish flowers", may belo
here. Its cocci are wide-flaring at the base, and it is cited
herein tentatively as Y. pumila Rydb.

VERBENA QUADRANGULATA Hi
Additional “epee Moldenke, Phytologia 11: 478. 1965.
species has been collected in fruit also in February.

tan fine s
grayish calcareous silt light grayish-tan sandy loam, light-tan
or loose light-tan sand’ » Pale-tan loose sand, very loose hia

in parral the
water's edge. The Flores call it "abundant in caliche", while
Gamez describes it y "common in fine sandy silt". The I. Moran

20, distributed as V. quadrangulata, is actually V. ciliata var.
pibers (Greene) Perry, while Barrera & Laure] 111 is V. pumila
nal citations: TEXAS: Duval Co.: P. Cabrera 53 (Au—

23096, ‘ta) 3 Ne Castillo sen. [10 oaareh 63h i556) Aa
Rirez, Lopes, & McCart 5 u-—232663); Ramos, S
Gart 8007 (hu—2220h8) ag Co.t Rs Le Crockett 013 (Id).
vim Hogg Co.: Alvarez » Guajarte, saniesets “& MeCart 7784 (Au—
22226)5 Sanchez, Rodrigues, & McCart 6311 (Au—202217, Ip). Kle-
berg Co.: Villarreal & Earles ™ 102 (Au—232962, Id). La Salle Co.:
wtelio lg (Au-~222218). McMullen Co.: Alvarez, Guajardo, Sala~
bw) mar, & McCart art 7723 (Au—222225), Starr Co.: Alvarez , Suajerdo,
paasar, & Wo & WeCart 792) (au—222220), 8042 (Au——222216)5 Me

> 9 (id); Ramos, Sandoval, Salazar, & McCart 7879 i
anes Ip). Webb Co,: Alvarez, @ Guajardo, jardo, Salazar, & MoCart 7632
(4u 2222169) Cardenas 57 (Id); Flores & Flores "138 (Au—

258 PHYTOLOGIA Vol. 13, no.

Sandoval, Salazar, & ‘Vetart 7848 (Au—22 = To99 Au——22221: 9).
CO: Tamaulipas: pas: Dominguez & & McCart 8182 (Au—232959) 5 Escalante
36 (Ip).

VERBENA RAI
Adaisiona i binlioer Serie? ,» Excerpt. Bot. AS: lh. 19625
Moldenke, P 78, lia, 190, & 97 1206

te ei : B. H. Warnock 48
(Ld), 20105 (Ld).

VERBENA RADICATA Moldenke
Additional bibliography: Moldenke, Phytologia 11: 78. 19653
Moldenke, Résumé Suppl. 12: 13. 1965.

VERBENA RADICATA var. GLABRA (Hicken) Moldenke
Additional bibliographys Moldenke, Phytologia 11: 478. 1965;
Moldenke, Résumé Suppl. 12: 13. 1965.

VERBENA RAMBOI Moldenke
Additional bibliography: Kelldenks, Ab peter Li: 52—53. 196h;
_—e sorter Bot. 21: 51 & [60].
965) cites Herb. Anchieta Serie and 25787 from Rio
para oft er both collected by him by himself. He suggests that the
be co ec with V. stellarioides Cham., and de=
scribes it as "Erect, stiff, coarse, about 30 cm high", growing
in See shrubby coast! on the "Southwestern plains only"
in Rio Grande do Sul. He gives its general distribution as "S0
far, it [Rio Grande do Sul

VERBENA RECTA H.B.K.
Additional bibliography: Moldenke, Phytologia 11: 78. 1965+
Recent collectors have found this plant growing in Abies for-

ests, Abies religiosa woods, and Pinus hartwegii woods, at , at altd-
tudes of 3000 to 3200 meters, flowering in July. The Paray 2uh9,
distributed as V. recta, is actually Vv. carolina L

Additional citations: MEXICO: Federal ere aray 56h 26h * (Ip)+
Hidalgo: Galvan s.n, [.VIII.1963] (Ip). México: J. Rzedows=
18h09 (Ip), son. [h.VIII.1962] (Ip).

REITZII Moldenke
PP go og ine sighing gpetesins:t Phytologia 11: 58--59-
ai weer giass be 1
Rambo (1965) c 3 Herb, Anchieta 50761 (Reitz 3663] and
50794 (Reits 3726) pee Santa Catarina, He describes the Pal
uffruticose pere — erect, apparently about 1m ee
in "Wet, apparently densely * vegetated Campos" and no

1966 Moldenke, Notes on Verbena 259

"To be found along the northeastern frontier" of Rio Grande d
Sul, but notes that its general distribution is "So far, SC ae
Catarina ] only",

VERBENA REITZII var. CASTRENSIS Moldenke, var. nov.

varietas a forma typica speciei recedit ramis ramulisque
foliisque antrorse strigosis, foliis 1—1.5 cm. longis ad cm
acutis sessilibus, inflores centiis spicatis usque ad 2,8 cm
elongatis.

This variety differs from the typical form of the species in
having the stems, branches, branchlets » leaves, and peduncles
appressed-strigose with antrorse hairs » the — only 1--1.5
Cm» long and acute at the sessile (not amplexicaul) base,
inflorescences bec to 2.8 cm. long airing a and ae ras:

» Brazil, on
N. Moldenke herbarium at Mai atiota’ New Jersey.
i a BRAZIL: Parané: Smith, Klein, & Hatschbach 11526

VERBENA RIBIFOLIA Walp.
Additional bibliography: Moldenke, Phytologia 11: 59—62.

196).
Additional citations: pyres: Concepcién: Besser 12 [Macbride
photos 17443] (W—~photo of type).

VERBENA RIBIFOLIA f, Acevedo de V
Additional Etnies Moldenke, Phytol ogia 11: 60. 196k.
Ricardi found this plant growing at rs altitude.
Additional citations: CHILE: Colchagua: Ricardi 3175 (2).

VERBENA RIGIDA Spreng,

aoshtditional bibliography: Moldenke, "19635" ae 33: —
Hocking, t. Bot. . 1962

Bains as, Shah cebactaa teen tered Ree, Sea Tee

Phytologia i: 478—-1,79.° 19655 “ilieks. Biol. pe “hes ne

19655 Moldenke, Résuné gat 12: 3 5. 1965; Neal,

von eVe @d., 722. 1965; Rambo, Pesquis. Bot. 2i: 553 ‘& “(oo)”

Sidey 3509 is said to have had "mauve" flowers and was found
rie See Kaen = oath es Klein fou found the species in for-
a. sat) cite tt, anita i 1, Bob, eb, i
= (Heidler r 8en.], 11227 7 [Friderichs fs sul, 3910 (rerio
Undo) Gat’ Gwe’ Gee, SA. Ue, RL Le, ees,

260 PHEYTOLOGTIA Vol. 13, no.

= 58857 oon te 173], 62057 [Camargo 19h], 63055 [Schlich-
ting s.n. ], 63278 [Sacco 853 J, 63786 [Sacco s.n.], and 63861
[Sacco 82] from Rio Grande do Sul, 47913 [Sehnem 3036] from Santa
Catarina, and 1,052 and 080 from Uruguay, by himself
— where otherwise indicated, He describes the species as
"Erect, stiff, coarse, up to 0 cm high", growing in "Dry, more or
less shrubby Campos, shows weedy tendencies......0ver the oe

re
(Ld). BRAZIL: Minas Gerais: N, Santos 592) (Ac). Santa Catarina:
A. Castellanos 2689 [Herb. Cent. Pesq. Florest. 3317] (Pf);
Smith & Klein 13813 (W--251612). ARGENTINA: Buenos Aires: Gillies
sen, (Macbride photos 3439] (W--photo). SOUTH AFRICA: Transvaal:
Sidey 3509 (W--2377223).

VERBENA RIGIDA f. GLANDULIFERA Moldenke
ee bibliography: Moilavce, Biol. Abstr. 33: ge 19595
erpt. Bot. A5: hh. 1962; Moldenke, Phytologia ll: 95—

%, seeks ts liesti, Bibl. Veg. Paran. 196. 196;

RIGIDA var. OBOVATA (Hayek) Moldenke
Additional bibliography: Moldenke, Phytologia 11: 479. 19655
Moldenke, Résumé Suppl. 12: 5.1
dditional citations: BRAZIL: Parand: Dombrowski & Saito 1308/
212k (Ac). PARAGUAY: Pedersen 5205 (W—2],32935) «

VERBENA RINCONENS
Additional betiéertines oldente, Phytologia ee N79 1965
Dodds & Simpson found this plant Powe ing
Additional citations: MEXICO: México: Dodds & Simpson 9 "9 (ua).

Mery RUNYONI Mo as
Additional bibliography: Mold » Phytologia 11: 109——
(1964) and 11: 460 & 502. 1965; Melsene, Résumé Suppl. 12: 1.

Additional citations: TEXAS: Cameron Co.: Fleetwood 8001 (Au
230978). Hidalgo Co.: R. L. Crockett 8032 (1d).

— RYDBERGII Moldenke 162 & W79
tional bibliography: Moldenke, Phytologia 11: Tl

sscte'y Moldenke, Résumé Su 3363. Gaiser, Surv. Vas¢-

Lambton Co. 100. RUS DO teks SPL Ae

966.

One each of the A. R. Moldenke 819 and 942 specimens cited
previously as being deposited in my own herbarium is now yt now in the
ing 4 in an open field in Ontario. Gaiser (1966) says "only &t

int Edward, V. hastata was on lower land under the Ambassador

Bridge, ve. stricta ons on sandy esta of St. Clair R.; mimerous Hy"
brids were along the roadway between the parents and intersing “

1966 Moldenke, Notes on Verbena 261

sis a joes. "
onal citations: ONTARIO: Peterborough Co,: Heidenreich
292° [7138) (Mi).

VERBENA SANTIAGUENS nke
Additional Bibliography: Js $o~ eats Bot. Ab: 535. 1963;

VERBENA SCABRA V
$0 Additional bibliography: Moldenke, Phytologia 11: 479, 501, &
“an

Proctor found this plant growing at the edge of swamps in
Jamaic

Additional citations: JAMAICA: G. R. Proctor 2322 cee HIS-
PANIOLA: Dominican Republic: Fuertes 392 (Mi )S 1758 (Ma

VERBENA SCROBICULATA Griseb,
Additional bibliography: Moldenke, Phytologia 11: 137—1,0.
196h; Rambo, Pesquis. Bot. 21: 50, 1965.

VERBENA S SELLOI Spreng.
ig fates selloi (Spreng.) Troncoso, Darwiniana 13:
1. 196). ‘Glandularia pulchella var, gracilior Troncoso, Darwin-
dana 13: 476. 196). Verbena pulchella var, gracilior (Troncoso)
> 266. 1966. me

ag : Spreng. Vege, ed. 16, 2: 750. 1825;
nf » Prodr, a: “Sho "a8, Schau. in Mart., Fl. Bras.
9: 19h, 1864; 5 Briq., Ann. Conserv, & Jard, Bot. oabe "2B: 296—
297. 190k; Hayek Sobel Kaiser. Akad, Wissensch. ssi el
79 (1)? 294. 1908; Moldenke, Lilloa 6: 333. 193k;
winiana 3; 5), 19375 Moldenke, Phytologia 9: 389, 2h, a "336. 19635
Troncoso, Darwini + L681 70 & 8-43, fig. 6 - 1964; Mo
denke, R as Sex Suppl. ie
Tiust trations: rhiaabo o, 0 OR rr gat a a 6. poe
© (1 in resurrecting thi:

cites the Peied rete eh Neen peer 2553 3 (Bi)5 00 cd

Sctor undesi ted 8 - (Carras

(Si), 22h0 ( Si); 27 Gh), o7k2. ah arti (, ae an (9, 3356

(Si); Herb a, Horven 33 87430 (Si); Herter 66161 SoD.

ey 11.1927] (P); Osten 5642 (Si); H. v9 [earn 8.n, (Car
©, 23.%.19);2] (si—25668); Rosengurtt 100) (Si), 2116 (Si),

Bal ae 1); Rosengurtt & al. 1133 Gi).

lek sl os fo céspedes extensos

floriferas o ean, belies “th a violdceos, lisos,
Glabrescentes o con pubescencia hfspida, extendida o

rllos y pediinculos, mas de a extreme de los mismos,
Entremdos breves, de 0,5--2 cm long. alarg4ndose hacia el 4pice
de las em long. Hojas pinnatisectas, de 0,7—~2 cm

Pramas
(aro mayores) long. x 0 »5—1,5 cm lat., de 4mbito triangular,

262 PHYTOLOGIA Vol. 13, no. }

enango
cidas o breves ramitas 6 died axilas; lacinias breves, subagudas

hirsu
dos (visibles sub pe iy nervio medio impreso en el epifilo,
ente en 6] hipofilo. Inflorescencia en 2 0 3 cabezuelas

terminales rara vez solitarias, espiciformes en la fructifa-
cacion hasta de ), cm long., pedunc ulos laterales,
generalmen largos que el central, con pelitos hispidos ex-

tendidos o retrorsos. Brdcteas lanceoladas, subuladas, de
(=-6) mm long., pestafiosas, el resto glabras. CAliz cilindrico,
mm long

cos que superan notablemente las tecas, exertos
ta corolar. Gineceo tipico, estilo filifore de 0, "él cm
long. Mericarpios cilfndricos, parduscos, de 2,8-—-3 m long.,
deados y alveolados en el *&pice, truncados en la base, cara
dorsal alveolado-sureada, cara ventral blanco-verrucosa, pores
carpio levemente prolongado en la base, pero no ensanchado.
She notes that the species is "Abunda en praderas

pu gen

ar "Si bien la descripcién de Sprengel es muy breve, e5 °D
tante coincidente, por lo cual creo conveniente rehabili-

tar este Best pr pues de no hacerlo deberia describirse

” GdiLeatoee ecnen this plant as 1— . tall, the corollas
varying from blue, bluish, or corak Hel fing to violet, lilac,
purple, bright purple-lilac, or reddish. They ha found it
ng in dry sand

April. The cammon names "margarita morada" and myer be meona" are
recorded by Herter in Uruguay. Swallen describes it as ore agrees
it .

8 c , While Smith r

Material of V. selloi has been misidentified and distributed

. lactate er the names V. dissecta Willd., V. erinoides Lae,
; ata (L. *) Briq., VY. microphylla H.B.K., ¥, pulchells

et, V. tenera Spreng., and V, temisecta Brig, The Chicclt 2 wh

eclisettieg cited below, is a mixture with V. Se a (Covas
Schnack) Moldenke. The label on the R Rossagetts Fe specinel |
also cited below, originally was inscribed "B-2116 oP, ae Pt |
fraction was apparently later crossed out |

1966 Moldenke, Notes on Verbena 263

The flowers are described as "blue" on Berro 134, 4753, L75u,
6720, & 8219; "blue or violet" on Berro 47525 " purplish-blue" on
Berro 8205; "bluish" on Berro 8390; "violet" on Berro 4323, Kra-
povickas 2752, and Burkart 22380; "reddish" on Schreiter 6613;
"lilac" on Herter 181i; “purpler on Berro 8701 & & 8775; a and
"bright purple-lilac" on Herb. Herter 50082.

a Chicchi o in part, Gallinal, ee Be

tr bel te eer reviousl? errone.

were previously cited by me as vy. tanehs ee These changes
in determination are due, in large part, splendid revisi-
onary work recently published nif Troncoso “(a96h) on certain
gentine members of this species complex. It is probable that
most of the material cited by me previously under V. tenera will
prove, upon re-examination, to be V. selloi. The Bailetti letti 231,
cited below, is placed here tentatively. It was originally dit dis-
tributed as rte erinoides Lam., then cited by me as V. dissecta
Willd. Its leaves have the exact appearance of those of V. dis-
Secta, but its ceri _ those of ve 3s genet It may
possibly oper oa a hybri tween the two 8]

Troncos eee says tou wee V. pulchella me "i or: "A

ap

ure Burk no. 25360} at Villa Gesell, Gral. Madariaga, Buenos
Aires, Argentina, on March 5, 1961, and is deposited in the herbar-
ium of the Instituto Botanico Darwinion at San Isidro. She c
also from Buenos Aires Hicken 8.n. forkl; Pinto, 10-I-1898] (si),

Sn. [Olivos, barrancas, 30-11-1929] (Si), and Nicora 201) (Si

T have, however, exam ined an isotype of this taxon in the he United
States National Herbarium and find it to be identical = ae
V. selloi. Shinners (1966) maintains san it is the co
o J

as ecta Ge
Citations: BRAZIL: Rio Grande di =r Rambo 34719 (N); Swallen
eh erg eget URDGUAY: He H, Bartlett Boe haga
3h (N), 4323 (N), 4752 (N), 4753 (N), 475k (N), 6720 (N),
op On 8205 (KN), 8219 (N), 8390 (N), 8701 (N), free (N), 8775
b. inst. el Lillo

STAT » Sen. [26,X1I.1946; Herb. Inst. Miguel caias 11755] (N,
N), 8.n, (18.1.19h7; Herb, Inst. Miguel Iillo 15138] (N, N), son.
Mores, : TI-1947; Herb. Inst. Miguel Lillo 11768] (N), gape, (9-X1E IRs
erb. Inst. Mi Lillo 11766] (N); Felippone 5915 (N); Gallinal,
Aragone, Ber ail, ¢ ran Hi ser EE, a Ths (i, i821 (NY, 2299 =e
W173 750}. Herb. Herter 50862 (N); Herter 18lc THerb. Herte
sel (N), ist Th (Herb. Herter 96600] (N); Herter & Strahl 181

erb. Herter 79 79174) (N); Legrand 2717 io Moldenke & Legrand s.n.
(Herb. Legrand 2791); Moldenke & Moldenke 19692 692 (N), 19696 (N)5

26, PHYTOLOGIA Vol. 13, no.

Rosengurtt B.2116 (N); A. Saint-Hilaire c?.1882 (N)5; H. M. Smith
14,0 (W--1193 3793); T 3 Teague 8.n. @ 8M. (22.11.1947; Herb. Inst. ;, Migue elo
Lillo dee (N, N). ARGENTINA: Buenos Aires: Burkart 22380 (W—

lata Sees ee Page] (W—227527). La bite; Chicchi 26), in
Salta: Garolera & Romero S.n. {Cuesta del Obispo, 8.I.

Tucum4n: batlettd 231 (Herb. Inst. Miguel Lillo 31690] (N). cuL-
TIVATED: Russia: Herb. Hort. Bot. Petrop. s.n. (W-—-71959) « LOCAL-
ITY OF COLLECTION N UNDETERMINED: Herb. Jussieu | 5.n. (N).

ESSILIS (Cham,) Kuntze
Additional pen somenty: gees oe Excerpt. Bot. a9: 366. 19655
sects Phytologia 11: 79. 1965.
onal citations: LS State undetermined: Sellow 1563
bigebtae photos 177] (a—photo of type).

pe Arak Lehn.
onal bibliography: Turez., Bull. Soc. Nat. Mosc. 3% (2):
195. . N. L. Britton, N. J. Geol. Surv. Final Rep. State Geol.
2 (1): iy? h. 18895 Quick, Wild Fls. North. States 372-2373. 19393
Mid rt, How to Know Spring Fls. 110, sgl fiom 19435
solace py Abstr. 46: 32h6. 19655, Ho cking
Bot. AJ? 366. 19653 yoldinke, Phytolo oils “486 (1965) ‘and 13:
185. py sha Gaiser, Surv. Vasc. Pl. Sato - 100.
nal illustrations: “ie J. sett ix to Know "spring
Fls. pet oe
Sddi tional ee cetiisl ALABAMA: DeKalb Co.: Demaree 50551 ase
sient Saint Louis: Muhlenbach 182 (Ac), 1208 (Ac), 1u6D (#

VERBENA SIMPLEX f. ALBIFLORA Molde
Additional iblingrastar | Macs recog Phytologia 11. 224 tess, 19655
Sree Biol. Abstr. 6: 326. 1965; Hocking, Excerpt. Bot. A?

XVERBENA SOLBRIGII Mol
Additional bib liogrepy Moldenke, Biol. Abstr. 35! weer 1960}
iB wwe Bot. Al: 592. 1962; Moldenke, Phytologia ll:

ENA STELLARIOIDES Cham
Additional bibliography: Rambo Pesquis. Bot. 21: 51, 53, &
[60]. 1965; Moldenke, Libemeeseg ja: 480. 1
Rambo (1965) s that V. ramboi Moldenke may be conspe-
cific with V. ih sorte phony Under the latter, he cites Herbe
Anchieta 9555, 9756, 10156, and 58698 [Camargo 51] from Rio

narrow and long", growing in "Dry, grassy Campos"

1966 Moldenke, Notes on Verbena 265

Campos only" in Rio Grande do Sul, with a ye distribution of
"Paraguay, westernmost RGS [Rio Grande do Sul],

Additional citations: BRAZIL: Rio Grande do pos Sellow s8.n,
{Macbride photos 17))6] (W——photo of type).

VERBENA STOREOCLADA Bri
Additional UibiSogranie Moldenke, Phytologia 11: 188--190,
1964; Rambo, Pesquis. Bot. 21: 58. 1965.

VERBENA STRICTA Vent.
Additional biblio ography: N. L. Britton, N. J. Geol. Surv.
Rep. State Geol. 2 (1): 19h. "1889; . fe Beal, Mich, Agr. Coll.
¢ Sta. Bull. 267, ete: 2, JS, fig. 27D 1915; Muenscher, Cor~
nell Agr. Exp, Sta’ 635 [Weeds N. Y.J, pr. 2, 10 & 387.
1939; Rickett, New pat Book Am. Wild Fls. 280, fig. 98. 1963;
Y. Matsumura 6

» 500
1965; F. H. mg prea Oe Native Wild Pl., p LZh, fig. 10,

5; Moldenke, Résum wpl. 12: 1 & 13 (1965 and 13: 1. 1966;
» Surv. Vasc. ik Lamb - 100. 1966,

Additional illustratio: aes d a ‘Beal, Mich. Agr. Coll. Exp.
Sta. Bull. 267, ed. 2, fig. 175. 1915; Rickett, New Field Book
Am. Wild Fls, fig. 98. 1963; Fe He Montgomery, Native Wild Pl.,

- 1965.

Muhlenbach 133 1335 was identified by the collector as "V Ve lex
x stricta", but I do not think pooh is a hybrid. One. — of
the A. R, Molideniee 92h and H. N. Moldenke 21909 specimens cited
previously by me a: by me as being deposited in my personal herbarium is
now in the sacelbacet a of Aarhus University. C.M. Rogers 6111 is
Said to have had "vio Late flowers,

Additional citations: INDIANA: Porter Co.: M. W. Lyon Jr. s.n.
(Baileytown, IX.13.1925] (Mi). KENTUCKY: aaa Co.: Shacklette
506 (Mi). KANSAS: Crawford Co.: H. F. Becker s.n, (Pittsburgh, 3

June 196) (Mi). MISSOURI: Saint Louis: Muhlenbach us (Ac), 1335
an tins COLORADO: Las Animas Co.: C. M. Rogers 4967 (Mi), peers Mi),

» 6956 (Mi). TEXAS: Hemphill Co.: er cea & Ogden 28302
(Ia), 28301 7)

VERBENA STRICTA - ALBIFLORA Wadmond

Additional mibliceceae: Moldenke, Phytologia 11: 480. 1965;
Moldenke, Résumé Suppl. 12: 1. 1965.
@ 2) Aidt tional citations: MISSOURI: Saint Louis: Muhlenbach 35

tao, c ters hytologia 11: 238—2)
ibli Moldenke, P’ :
498, & 500, 1965; pao waa Bot ci 55h & [60]. 1968.
spsiatee hte Cites Herb. Anchieta 2817, 8492, aon Te,

» 32h50, 34728, 32, lg3i9, 5 is "50999, 519h0,
27h, 56605, all from 0 Grande do Sul, and 55298 [Reits L7h6]

266 PHYTOLOGIA Vol. 13, no. k
32000 — 4742], 55303 [Reitz 4739], and 56868 [Reits s.n.]
anta

‘ ” Braz ‘ collected by yy himself except where
pty Gated The Santa Cata he says, were iden-
tified by me as V. preng. He describes V. strigosa as a

"Low , branche 3
st high", found in "More or less wet, peaty, grassy Campos" on the
ghlands only, especially the northeastern sector" of Rio Grande
do Sul, with a general distribution of "From S, Paulo to RGS [Rio
Grande do Sul]".
Additional citations: BRAZIL: S¥o Paulo: Sellow s.n. [Brasilia;
Macbride photos 17,8] (W--photo of isotype).

VERBENA SUBINCANA (Troncoso) Shinners, Sida 2: 266. 1966.
Synonymy: Glandularia subincana Troncoso, Darwiniana 13: ,78-—
81, se? 5.1
graphy: Tro. roncoso, Darwiniana 13: aa & 478-181,
mice ae + 96h Shinners, oe : 266. 1

tions: Tronc Datwitt ana 13: 1,80, nied 5. 196.

beet circ. 25 cm alt., caulibus ad basim prediredth interdum
» demum ascendentibus, subtetragonis; pubescentia a~-
lous versus ’ siibincana, ad caules Setrebdo-biapit , ad bracteas
nervis rpacia calycique glandulis patelliformibus intermixta;
internodiis 1--5 cm longis; foliis Beem ago 1--),5 cm lon-

subincurvato apicem versus incrassato; antheris
dricis elongatis, exsertis, munitis; stylo 8,5--9 mm longo; fruc~
tu 2,5-—-2,8 mm lo ongo, mericarpiis obtusis , dorso-reticulato-

ta sa

sulcatis......P nne, séfruticosa en la base, con tallos

extendidos, rastreros o radicantes en la base, luego erguidos,
sin formar c subtetr4gonas con oubescencia hfspida,
endida o retrorsa sobre los 4ngulos, el resto cano-hispidulo,
m4s densa hacia el 4pice d4ndole un aspecto ecino. Entre~

nudos breves, de 1—3 cm long, m4s largos hacia el extremo de
» hasta de 5 cm long. Hojas pinnatisectas, c fascic-

ulo de hojitas en axilas, de 1-3 (—l,5) om long,
tenuadas en breve pecfolo, ldmina de 4mbito oval-triangulars | :
’

hirsutas en el epifilo con pelitos rfgidos, callosos en 1a
densamente en el hipofilo sobre los nervios y el margen sub-

revoluto Bio en material seco), nervio medio impreso eae

suo

proto desert nr otbprer ager rants nfspid nispia-
cacién 7 cm Long pediinculo hfispido y cano~
ulo con pubescencia retrorsa. Brécteas lanceoladas, subcdncavas

1966 Moldenke, Notes on Verbena 267

en la base, de —1,5 mm long, generalmente superando la 1/2 del
cdliz, nervio medio “manifiesto, pestafiosas, elresto incano-

te en la 1/2’ inferior = con “peliial m4s rigidos y oblicuos hacia
el 4pice de la costillas y los dientes » ciliolado en el borde,

cm

long., brevemente encorvado y ampliado en la parte superior. Es-
tambres t{picos, los superiores con apéndices anterales cilfndri-
oO in-

cos, super: ecas omando en la garganta corolar, 4G:
eceo tipico, estilo de 8,5—-9 mm long. ppg pad de 2,5--2,8

ong-, recubiertos por el c4&liz persistente madure se
hiende lateralmen > cos, se@ “te

resto surcado, cara ventral blanco-verrucoso (sublente), pericar-
ie prolongado por debajo de la parte fértil, no ensanc
Habita lade

Sp: br&cteas, cfliz y ulos, a “ corola glabra
exteriormente y en general por ser toda ella ho m4s hirsuta y

Pubescencia, no estrigosa y por los mericarpios menores y redon-

deados Phas el 4pice, no rostrados." She cites from Entre Rfos,

‘rgentina, Burkart 25425 (Si) and from Uruguay Rosengurtt & al.

492 (at) and PE.5262 (Si). ;
type of the s, species was collected by Bernardo Rosengurt

Qurvich (no. B B.992), common in riverside thickets at Arroyo Val-
» Rfo A.

and low or dry ro rocky sunny sandy camp
slopes, at 150 meters altitude, flowering in September, October,
and F

Sul,
Brazil. The © flowers are described as "lilac" on Rose + B.992,
"violet" on Herter 181g, and "purple" on Rosengurtt B.3756 and
aro em, = Eee, —_ & Rosengurtt mete:

Fee ne, Bergalli, Campal, & Rosen
ett ¥B.2262, gare 5.n. ee ee 1887; Herb. Com. Geogr. e Geol.

268 PHYTOLOGIA Vol. 13, no. k

3685; Herb. Inst. Bot. S. Paulo 15725], Rambo 10103, and Rosen-

t B.3756, cited below, were previously cited erroneously by
me as Ve V. dissecta Willd., while J. T. Lima s.n. [II.19)6; Herb.
Rio de Jan. 55466] was ee TOES, cited by me as V. pulchella

Sweet.

Citations: BRAZIL: Rio Grande do Sul: Rambo 10103 (N). S&o
Paulo: Brade 7000 [Herb. Inst. Bot. S. Paulo 6728] (N, N); J. Te
Lima s.n. [II 196; Herb. Rio de Jan. 551,66] (N); Léfgren s.n-

{2 [22=-9-1887; Herb. Com. Geogr. & Geol. 3685; Herb. Inst. Bot. te Se
Paulo 15728] (Nt). URUGUAY: Gallinal, Aragone, Bergalli, Campal, &
Rosengurtt PE.5262 (N, N); Herter erter 16lg {Herb. Herter 83911] (N);

Rosengurtt B.992 (N--isotype), B.3756 (N, N).

VERBENA SULPHUREA D. Don

Additional bibliograp ra ae Excerpt. Bot. A9: 366. 1965;
Moldenke, Phytologia 11: 80-h81 "(1968) and 13: 213. 1966; Mol-
denke, Résumé Suppl. 13: 3. 1966.

The Herb. Inst. Miguel Lillo 31745 and JUrgens snsen 1a dis-
tributed as V. sulphurea, are actually V. parodii (C
Schnack) Moldenke, not V. dissecta Willd. as stated aR me yar

viously.
Additional citations: CHILE: Coquimbo: Marticorena & Matthei
7h (Ac), 105 (Ac), 230 (Ac); Ricardi 2028 ven Ricardi & & Marti-
corena 236/621 (Ac). Valparaiso: Behn . [Vifla del | del Mar, 19.
XI .1922] (Ac); ee sen. (Quinteros, 5. 9H ci9h2] (Ac).

rysend tikes nocaranye “Yoldei Phytologia 11: 480. 19033
Moldenke, Résumé Suppl. 13: 3. 1966. we
Additional citations: anise: Valparaiso: F, Torres 8.N-

{Quintero, 19.1X.1958] (Z).

ey aasiaacad Denitees desig rahioges 11: 481. 19655

8: Moldenke, ps : e
Moldenke, Résumé ain aa 3. 1966 fa Sts

Recent collectors have found this plant in flower and fruit
in October,

Additional citations: CHILE: Antofagasta: Ricardi 2701 (Ac),
3090 (Ac). Coquimbo: Marticorena & Matthei 350 (Z)6

UPINA L.
Additional bibliography: Hocking, Excerpt. Bot. A9: 366. 19655
Moldenke, Phytologia 11: 251--261 & 90. 1965.

VERBENA TENERA S :
Additional bibLiogranky: Troncoso pra ee 13% £ M6B—LT3» 6!
196; Hocking, t. A9: 366. 19655 Rambo, Pesquis. & 185+ 96

S155 & & Jol. 1965) Soldenes. Phytologia 11: 458, 468, 48

rations: Troncoso, Darwiniana 13: 472, fig- 1

1966 Moldenke, Notes on Verbena 269
1964.
Smith and his collaborators found this plant growing dry
Open ground and on rock outcrops, as well as in bogs, in Santa
C.

Rambo (1965) cites Herb. Anchieta 9389, 9913, 10005, 10103,

1155], 61443 (Sacco 646], 61444 [Sacco 647], 63207 [Spies s.n.],
and 632h), [Spies s.n.] from Rio Grande do Sul, Brazil ol-

g

radiating, up to 0 om long", growing in "Dry, more or less open
to grassy Campos" in the "Western to central Campos area, absent
from the northeastern
eral distribution "From Rio de Janeiro to Uruguay and northern
Argentina",

Troncoso (1964) describes the species as follows: "Herb&cea
perenne, cosa en la base, rastrera, muy extendi

ta, cm
cfeulo de hojitas menores o ramitas abreviadas en las >
lacinias filiformes de 0,5--1 mm didm., por lo general enteras,
8 » Subrectas (incurvadas en el material seco), glabres-
centes o con pelitos estrigosos ralos en ambas caras, ne
medio unico inconspicuo en el epifilo, bordes extendidos, a veces
en el material de herbario parecen subrecurvos. Inflorescencia
®n cabezuelas terminales solitarias, pedunculadas, especiformes,

lanceol —3,5 mm long., con nervio medio visible, subcén-
Cavas en la osas. eilfindrico, i.
long. (ineluidos los dientes), densamente estrigoso-pubescente en

Br&écteas y g
Corola glabra exteriormente, tubo de 12—13 mm long. Estambres
ces anterales c cos
Sobrepasando las tecas y exertos. Pistilo tipico, estilo fili-
forme, de 10,5—~11 mm long, Mericarpios cilfndricos, de 4,7—5,5
°

hon notes "Habita principalmente en terrenos pte: hapecie
vecina de Gl. pulchella var. pulchella, pero ésta se diferen-
Go* Por las lacinias de las hojas mis anchas, toda la planta més

Tadicantes sélo en la base.....Destaco que Grisebach en Symb. ad

Uren ptteent. p. 277. 1879, al citar la especie para E. Rios,

(Ve ones seflala: 'verae speciei folii segmenta angustissima
/3 lin. fere lata), glabriuscula, corolla longius exserta et

270 Pe 20k OcOTas Vol. 13, no.

caulis suffruticosus". Los ejemplares uruguayos estudiadosse
hallan determinados por el Dr. C. Osten y coinciden exactamente
con el concepto de Grisebach." She cites: ARGENTINA: Entre Rfos:
Burkart 2309) (Si), 3616 (Si); Herb. Inst. Darwin. 1217 (Si);
Lorentz 183 (Bf); Ragon nese & Martinez Crovetto retto 9 (Ib, Si), 5006
(Si); Rivas s.n. (Holt, 15-x1-192),) (I1b—77616). URUGUAY: Berro
se [Mercedes] (Ug); Millot s.n. [Herb. Rosengurtt]; Osten 1656
Ug

She ape 10% = following key to distinguish this species
from its relative

1. Patents seecsanin rala o densa en tallos, hojas, cAliz
teas; pedtinculos con pelos splicados, acroscépicos; ode
liz estrigoso o cano-estrigoso en toda su superfice.

2. Dientes del cAliz subulados o sipheltlatoe? ensanchados en la
base, eo o subrectos; apéndices anterales exertos,
su Ss tecas; mericarpios de 4pice rostrado o sub-
pesos y base dorsal del pericarpio ensanchada.

3. Segmentos de las hojas filiformes, de 0,5--1 m lat., sub-
glabros, bordes extendidos; brdcteas ovales u oval-
lanceoladas, de 1/3—-1/ la longitud del cAliz, rara vez
meerer) nericarpio os de ,5—-5,5 (—-6,5) mm mee i

ventral

3a. seeiuciaea de nee i linear-lanceolados, de is =
lat., m4s densamente estrigosos, borde revoluto o —
revoluto; aptote teas leececkadde. 2 do o
1/2 del oflis; mericarpios de 3,5—-l mm 18) brevemen-

rostrad ulchella.
2a. Dientes del oftis setoso-filiformes, recurvos, Acre
en 7 apéndices anterales inclusos, no rts fsb -

”
la. Pubescencia h{spida o hirsuta en tallos, hojas, c vhs y
brdcteas; Minculos con pelos extendidos o mds cominmente
retrorsos, basiscépicos; c4liz hirsuto sobre las costillas,
el rest © subglabro o cano-hispfdulo.
h. Nit Toreaceaeta en cabezguela solitaria; tubo corolar glabro
xteriormente; céliz y br4cteas cano-hispfdulas, con peli-
tos m&s rigidos sobre las costillas del cAliz a —
- Inflorescencia en 2 6 3 cabezuelas terminales
beg Pee aia pubescente BRO y es +5 F 5 eAlis J

brécteas :
Ss Coemenes 2 6 3 terminales, rara vez solitarias, alargan-
e después de la antesis; brécteas lanceo » large”

aN ae age eeeee seca eV S@LL04 selloi.
5a. Cabezuelas cimoso-panicul . ‘Snflorescencia 2 & 3 veces veces
ba oma, corimbiforme; gel im no alargéndose des~

by me previously as V, tenera actually represents V. selloi 3¢

cording to this revision made by Troncoso. Burkart 22300, isch :

1966 Moldenke, Notes on Verbena 271

cited previously as V. tenera, is the type collection of ¥. pul-

chella var. gracilior (Troncoso) Shinners, but this I regard as a

synonym of V. selloi Spreng. Burkart 3616, which she cites as Vv.
| have ve previously cited as V. vy. tenuis ecta Briq.

“The. flowers are described as "blue" on Schreiter 1904. The
Moreno s.n. [Patagon. 1882/4], Schreiter 190, Tweedie s 8.n., and
Wilkes s Exped, S.n., cited below, were previously erroneously ci-

ted by me as V. di: dissecta Willd., while Carette s.n, [Monte Hermo-
so, 1916] was | previously cited by me in error as sv. microphylla

BK.

On the other hand, the Boffa 25 and Commerson s.n. aiakegoerd
Ayres et Monte Video}, previously cited by me as ¥. ti tenera
actually V. dissecta Willd.; A. Castellanos S.n. (stash: ier.
Inst. Miguel Lillo 15120] is V. mendocina R. A. Phil.; A. T. Hun
ziker 6,71 and T. Meyer 2409 are V. microphylla H.B.K.; Heres
atts Miguel Lillo 31472, J& JOrgensen 1737, and Rufz Leal 7092

V. parodii odii (Covas & Schnack) et A. a ae 5160 is V

ostoma Klotzsch; H. M. Smi hella Sweet; a8 | Pas
Bartlett 20821, Berro © 13h, aE j te us, 75h, 6720,

Berne by eos ATG2, M33, Ay SIO,

ne ee

lanos $.n,. (1 .XI 1916], 8.0. locate isle 8.n. ae (6 1a9t" 5.ne
(7.X1I.1947], & fete [9.XII .1947], Felippone 5915, Gallinal,
gone, Bergalli ,& Rosengurtt 2299, Garolera & Romero ,
ontag del = I.19h7], Herb. Inst. Miguel Lillo 11755,

» 11766, 11768, 15138, 157%, & 32795, Herb. Herter 7917h,
Beer, & 96600, Her Herb. Hort. Bot. Petrop. s.n., Herter 1éle & 1614,
Herter & Strahl 161, | Krapovickas 2752, Legrand 2717, he &
Legrand s.n. [Herb. iearenal 2791), Moldenke & Moldenke 19692 &
19696, Edw. Palmer s.n, [La Plata Exped. Capt. mein Rambo 34,719,

Addi tional citations: BRAZIL: Santa Catarina: Reitz & Klein
Bis (W--2h03363); Smith & Klein 13388 (Ac), 13730 O (W—21S1590)5

Smith & Reitz 1,372 (W—2),51603). URUGUAY: Sellow 99 (Macbride
Photos 17h50] (W-=photo of type). ARGENTINA: Buenos Aires: pero
Tiitikes {Monte Hermoso, a) a a Negro: a

Lillo 3 32759) (N). Ror re Saati Sitgie sn. [Patagon.
athe (N). LOCALITY OF COLLECTION UNDETERMINED: Tweedie s.n.

272 P BE 820 4:0:6'T A Vol. 13, no. 4

VERBENA TENUISECTA Briq.
Additional synonymy: Verbena tenuisecta Brig., in herb. Ver-
bena teniusecta Brig., in her
Be hime bibliography: Mol denke, Biol. Abstr. 36: 719. 1961;
ont BY Bot. A5: ll. 1962; Sharp & Baker, Castanea
183. 1963 Troncoso, Darwiniana 13: heeerts & \71-\78, ee <
ie Moldenke, Phytologia li: 448, 481, 485, & 486 1965; ; Neal,
Ra he oe rev. ed., 722. 1965; Hocking, Excerpt. te:
19: 366. 1965; Pesquis. Bot. 21: 58. 1965; Moldenke, Résu-
ms Bee 12: iF EB; rt 13 (1965) and 13: 6. 1966; Shinners, Sida
2 2
. Aaa dona illustrations: Troncoso, Darwiniana 13: 477, fig.
- 19
Recent collectors have found this plant growing in black clay,
loam, calcareous marl, and in fine sandy silt. Godfrey re-
ports it "abundantly naturalised along roadsides" in North Caro-

er acts ae (196)) tah gnc Bec species as follows: "Hierba
renne, rastrera, de xtendidas, radicantes, las extremi-
dades florfferas pp a tes ra ga Shite a subtetr4gonas, glab-
rescentes o pilositsculas, abs densamente pubescentes hacia el
&pice. Entrenudos breves, de 1,5--3 cm long. Hojas Peper
tas, con fascfculos de hojas menores en las axilas, ldmina de
pene » atenuada en sagan, de 2—~3 cm long., lacini-
su ectas, enteras o den margen ex-
tendido a sveusiaeh (en eavaxtas ney Pies en el epifilo,
con un nervio central prominente en el hipofilo, estrigosas en a.

4 °
@ 3--3,5 mm long., estrigosas, C4liz cilfndrico, de 8—-9 mm
pte (iaeintaos los dientes), densamente cano-estrigoso, con po~
cas gldndulas pateliformes, subsésiles; hesigreag” aa ee
setoso-filiformes, desiguales, los mayores de 2--3 mm long.,
generalmente recurvos, Corola glabra, tubo ae 11--12 mm long.,

garganta c bres t{picos, los superiores con apén-
dices anterales reducidos, no o las tecas ni la garganta
corolar. Gineceo tipico, de 0,91 cm ou ios pard
negruzcos, ar nengh , de 2,5, andy redondeados en el
&pice, truncados © apenas ensanchados en la base, 4pice y
dorso retioulade” Hy te mitad mS el perccs es stefan 4 cara
n o-verrucosa o sublisa. a: Para y ME arge®

tino......Especie may vecina por su ndbi to de G. pulchella y Gl:
tenera, de las que se diferencia por los dientes del c4liz, seto-
s0-filiforme rmes, de base eo enapea y por los mericarpios m&s pe-
queflos, de 4pice redondeado rostrado." She cites from Pare
ra sass 160) 1603 (Si), JOrgens nes 3771, and T, Rojas 2528 (si) ‘
Shinners (1966) maintains that the correct name for this

is V. pulchella var. gracilior (Troncoso) Shinners, but this —

1966 Moldenke, Notes on Verbena 273

belongs in the Synonymy of V., selloi Spreng.
The Burkart 3616, cited by me previously as V. tenuisecta, is

regarded by Tr by Troncoso (1964) as representing V. tenera Spreng. in-
s

Recent collectors have found V. termisecta growing in dry or
Somewhat moist te rather open | places in Rio Grande do Sul,

il. The flowers are described as "violet" on A. T. Hunziker
3333. This ehiieht tic and Malme 31,8, sree below, were previous-
ly e: erroneously cited by me as ve. tene a Spreng. Material of V.
tenuisecta has been misidentified aie distributed in herbaria as
V. ciliata Benth,

Additional citati das SOUTH CAROLINA: Dillon Co.: R. K. God-
frey SC.149027 (Ld). ALABAMA: Crenshaw Co.: C. T. Reed 2025
(ld). LOUISIANA: Jeff Davis Par.: R. L. Crockett 8313 (14).
TEXAS: Jasper Co.: D. S. Correll 2727 (ld). Jefferson Cos: Re
L. Crockett 8418 (Ld). Nueces Co.: B. E. Campos 1-A (Au—

23 232,72) « Starr Co.: P. Cabrera 87 (Au--232h74). Tyler Co.: R.
L. Crockett 6707 (Ld). Webb Co.: | L. R. Cuesta 28 aes Zapata
Co.: Balleza .&Y. Valdez 169 (Ld); Flores & Powell 106 (Ld). BRA-
ZIL: Rio Grande do Sul: Malme 31,8 (N). PARAGUAY: Hassler 61h
[Macbride photos 17451) (W--photo). ARGENTINA: Buenos Aires: A.
1. Hunziker 3333 (N). Cérdoba: Hieronymus s.n. (Cérdoba, 14/11/
1876] (Mi). “Formosa: Morel 530 { ua). CULTIVATED: India: Kho-
Shoo 2 ( td 3 (as), aon, Thier ocal Botanic Gardens, Lucknow
( Texas: Y, Villarreal 20 (Au—232h)6), 253 (Ld).

bee at TENOIS ECTA var. ALBA poscees . aa he ei
onal biblio Moldenke 0. °

1965; Moldenke, » Rbown Supp 12:1 D (1965) a and 13: a - 19562

Recent collec rs have found this plant growing in

silt, rig Fe ruary.

Additional citations: LOUISIANA: Sabine Par.: Demaree 18315

(Ac). MEXICO: Tamaulipas: E. Villameva 39 (Au--232li2). CULTI-

VATED: India: Khoshoo (Ac),

VERBENA TENUISECTA var. GLABRATA Moldenke
plgeuen bibliography: Moldenke, Biol. Abstr. 36: 719. 1961;
oe 5 Bot. A5: hk. 1962; Moldenke, Phytologia 11:

039%.

VERBENA TEUCRIIFOLIA Mart. & Gal.
Synonymy: Verbena teucrifolia Mart. & Gal. ex Molden-
a Soong Suppl. 7: 10, in syn. 1963; Hocking, Excerpt. Bot. A9:
Additional bibliogra: Moldenke, Phytologia 11: 305—
30: 1965; Beamann, Bol, gee . Bot. Mex. 29: (71] & 7h. 19685
Hocking , Excerpt. Bot. ior "66. 1 1965.
Recent colle ctors have found this plant growing in Pinus pat-

27h PHYTOLOGIA Vol. 13, no.

ula woods, and material has been misidentified and distributed in
herbaria as Rubiaceae.

Additional citations: MEXICO: Federal District: Powell & Ed-
mondson 746 (Mi). Tlaxcala: V. Aguilar 8-A-21 (Ip). Vera Cruz:
Dodds 99 (Mi); Vela G.1227 (Ip).

THYMOIDES Cham.
Additional bibliography: Pesquis. Bot. 21: 5h--55 &
[60]. dase “pet s geetogle i: $11-=31), 1965.
n found thi t growing in bogs and on river-

ra re tarina, Bras °
cong (1965) cites Herb. Anchieta 2790, 4538, 9407, 11850 fAu-
Sen.], 29122, 31077, 3132, 34725, | 36415, 37491, 37827,
Loney: 1356h, 1g091, 18873 , 57608 [Pivetta 629], 56722 [Camargo
35], 58730 [1 0 [Camargo 216], | 58767 [' (Camargo 105], & 61302 [Pivetta
1235} from Rio do Sul, and an, 35979 (Hatschbach 500],
and 9579 from Santa Catarina, all collected by himself except

Paraguay through South Brasil to

Additional citations: BRAZIL: Santa basiaaahi Sellow 1182 [Mac-
bride photos 17454] (W—photo of cotype); Smith & Klein 13610
(W—-21,51608) . ;

meres
ibeioes ography: Moldenke, Phytologia 11: 325--326
965), ent 13 13: : 19. 1 eta
h 1822 distributed as V. tumidula is actually
Vv. deities —

VERBENA URTICIFOLIA

Additional bibliography: N. L. Bri kitten, N. J. Geol. Surv. Final
Rep. State Geol. 2 (D: tL. pet Sigh Ww. S. sae Boe How Pian
Wild Fls., rev. ed., 122, 365, & 372. 1896; W he Agre
Coll. Exp. Sta. Bull, 260: 153, te. “154 (1910) a a 2st, fe 2,
116, fig. ee 1915; Muenscher, Cornell Agr. a # Sta » 635

eeds N. Y.J, pr. 2, 10 x 387. ih Quick, Wild Fls. North
States 372 & o1B 1939; Moldenke, Biol. ines. "36: 719. 19615

rations: Agr. EXxP«
Sta. Bull. 260, fig. 15) (1910) Hn "er Fe a fig. Oe. 19155
Rickett, New Field Book Am, Wild Fis. fig. 98. 1963.

&

1966 Moldenke, Notes on Verbena 275

Additional citations: VERMONT: Windsor Cogs A. R. Moldenke
1130 (Ac), 1132 (Ac). INDIANA: Porter Co.: M. W. n Jr. 8D.
Keiser, VIII.10.192)] (Mi). aerate Cheatham Co.: Demaree
51102 (Ac) - MISSOURI: Saint Louis: Muhlenbach 993 (Ac), 10h
(Ac), 1286 (Ac), 1293 tga Se (Ac), 2135 (Ac). TEXAS: Bowi
Co.: D. S. Correll 31288 (Ld).

URTICIFOLIA var. INCARNATA (Raf.) Moldenke
eae sepa ate oy A wes —wival, Biol. Abstr. 36: Pig ony
ocKing, Excerpt. Bot. A 1962; Moldenke, Phytolo
011. 1965. :

ee URUGUAYENSIS Moldenke ae
tional bibliography: Hocking Berne Bot. A9: 367. H
Moldenke, Phytologia 11: 20—~I21, 1965

xVERBENA VAGA Moldenke
Additional syno P echete xVerbena vaca Moldenke ex Hocking, Ex-
eerpt. Bot. A9: 367, om
Additional ablation ‘Moldenke, Phytologia 11: 421. 1965;
» Excerpt. Bot. A9: 367. 1965.

VERBENA VALERIANOIDES H.BeK.

on. bibliography: Moldenke, Phytologia 11: 481-82.
Additional citations: COLOMBIA: Cundinamarca: Humboldt & Bon-

Pland s.n. [Macbride photos 39)76] (W--photo of type).

RET olde
Additional biblio eo Hocking, Excerpt. Bot. A9: 367. 1965;
Noldenke, Phytologia 11: 82. 1965.

VERBENA VENTURII Moldenke
Aiditional bibliography: Hocking, Exerot. Bot. A9: 367. 1965;
Moldenke, Phytologia 11: 182-183. 1965.

VERBENA VI VILLIFOLIA H
Additional bibliogra she phy: king, Excerpt. Bot. A9: 367. 1965;

Moldenke, Piytclogia 11 ll: 2 185-18 1965.

Notes collectors have found this plant in flower and fruit in

Additional citations: PERU: Junfn: Hutchison & Tovar 209 (Ac);
Weberbauer 2582 [Macbride photos 171,58] (W—-photo of type).

VERBENA WEBERBAUERI Hayek
Additional bibliography: Hocking, Excerpt. Bot. A9: 367. 1965;
Moldenke, Phytologia 11: 185. 1965.
Additional citations: PERU: Puno: Weberbauer }0 [Macbride
Photos 17159] (W--photo of type).

XVERBENA WINGEI Moldenke

276 PHYTOLOGIA Vol. 13, now 4

Additional bibliography: Moldenke, Phytologia 11: 4,85--1,86.
1965; Hocking, Excerpt. Bot. A9: 367. 1965.

ENA WRIGHTII A. Gray

Additional bibliography: Hocking, Excerpt. Bot. as 367. 1965;
os Phytologia 11: 437, 440, is2, & 186-197. 1965.

In s

ot
Porn narl, seh Warnock says that it is "frequent" in

Te)
107 & 319 are described as "pink", On Beasley & Finzel 005 they
were "violet-pink" and on Crutchfield 63 ee Te ae

Additional citations: TEXAS: Brewster Co.: E. Marsh 247 (Au--
214398); B. H. Warnock 20022 (Ld). El Paso Co.: Lundell & & Lun
dell 16941 (Id). Presidio Co.: B. H. Warnock 900), lb (Mi). NEW MEX-
ICO: Otero Co.: J. R. Crutchfield d 63 (Ld). Socorro Co.: J. Re
Crutchfield 107 (Id). ARIZONA: Apache Co.: Deaver 5599 (Ac Ac),
6483 (Ac). Gila Co.: Beasley & Finzel 805 (Id). Navajo Co.: a.
R. Crutchfield 319 (Ld). MEXICO: Nuevo Leén: May, Carrosco, I-
barra, & MeCart 6902 (Au—-232372).

VERBENA WRIGHTII f. ALBIFLORA rene denke
Additional bibliography: Hoc Excerpt. Bot. A9: 367. 19655
Moldenke, Phytologia 11: 191 & 19 7eu98, 1965.

VERBENA cose easton

Enen : Verbena matthesii Turcz., Bull. Soc. Nat.
eet 336 (2) 1 16157. oe. t
ed bibliography: ‘Tares , Bull. Soc. Nat.
Mosc, 36 (2): sen iag 1863; Hoc Hock Aya Excerpt . te A9: 367.
1965; Moldenke, Phytologia 1i: he & “f,98--507. 19

Turczanonow ” (1863) describes his ¥. peree lw as follows: "V.
caule fruticoso tetragono, paniculato-ramoso, pilis adpressis aut

© Oppo

tubo corollae calycem parun a hmaoN. nuculis transverse rugo-
sis. Ad V. menthaefoliam Bth., wees a descriptione, videtur acm
bracteae

cedere; at | non parvae, calyce paulo breviores, CO~
rolla calyce paulo nec triplo tajor "
Additional citations: TEXAS: Bras : Demaree 50859 (Ac)o

Fayette Co.: Matthes 13 [Macbride un 34303) (W--photo) .
Galveston Co.: R. L. Crockett 8462 (Ld).

ADDITIONAL NOTES ON THE GENUS CITHAREXYLUM. II
Harold N. Moldenke

B, Juss,

Additional & emended synonymy: Citharexylum apud A. L.
Juss., Gen. Pl., ed. 1, 108. 1789. Cithar, et oe L. apud L. C.
Rich., Act. Soc. Hist. Nat. Paris 1: lll. 1792. Citharexilum L.
apud M. Kunz, Anatom. Untersuch. Verb. 70, sphalm. 1911.
Citharexilon. Harvey ex Moldenke, Alph. List Invalid Names Suppl.
1: 4, in syn. 1947; Soukup, Biota kz: 260. 1963. Hadongi Gagnep.
in Humbert, Not. Syst. lh: 30. 1950. Catharexylum Greenm. ex
Moldenke, Résumé Suppl. 3: 30, in syn. 1962. SSE n Bark~
ley ex Moldenke, Résomé Suppl. 3: 30, in syn. 1962. Citharaxilium
gan bécand Suppl. 13: 6, in syn. 1966. Ci Sates aT Molden-

, Résumé § Suppl. 13: 6, in syn. 1966.

appt & emended bibliography: A. L. Juss., Gen. Pl., ed.

and

> Soc.
3
& 340. 1878; Solered., Bull. Herb. Boiss +s ioe % 6: 628. 1898;
An-
atom, Untersuch. Verb. 69-70 & & 78, fig.

8.
Contrib. U. S. Nat. Herb. 23: 1235, eit, & 122° 92h; Britton &
P, Wils., Scient. Surv. Porto Rico 6: 137 & lky—1h6. 192 25; Pater-

- i Ca é- & Cientif. Pl. Mex., [ed. 2], 339.
1937; Troncoso, Darwiniana : 50, 51, & 56. 1937; ees "Lecce.
Bot. 2: 97 & Lo8. 190; ‘eal, In Gard. Hawaii 635, 639, & 640.

1948; Gagnep. in Humbert, Not. Syst. ll: 30. 1950; "aa t,

Bot. 2: 28 & 37, 1950; Moldenke, Biol, Abstr. 27: 2026 & 3121.

ioe Pe joalis sbe, Ind. Kew. Suppl. 11: 55 & 106, es} Kg a

Lit Dept. Ree pe Serv. Agric. 110—1ll &

i 1353; peer ahtie of lava & Descrip. Fam gre z. 192,
me 258, » 270, & 2b, 2g5hs Moldenk Verb. 19-20. 1955;

ik 1. ar Moldenke, Biol. Anstr. ea
2353" 1968; Voldenke, *p » 2%

hale fg ei
: Stor 1 (1958), 6% 48-505 (a hy and 7: 7—77.

19595
owla 11: 77 & 97, 1959; Moldenke, Biol. Abstr. 33: ieee (1989),
as i 0933), and 35: 1h64—1h65 & 1689. B89. 19605 Ang ely, Liv. Gen.
. 1960 Angely, Fl. 15) 12" oh. 1960;
Seal, Biota 3: 30. 1960; Mufloz Pizarro >, hee. ae Descr.

« 1960; Anon., Assoc. Etud. Tax. Fl. « Trop. Ind.
19993 53. ae Troncoso, Darwiniana 12: 260. 1364; Moldenke, Biol.

1962; Angely .
22: 39. 1962; Soukup, Biota h: 172, 189, & 192, a ie enisate

2; Mo umé S : 3.
962; Hocking, Bxverpt. Bot. A5# beke (ioksy ant ASS 3s. tees,
Shelford, Ecol, N. . kes, met a 557. 1963; T. Mey., Op. Lill. 10:

278 PHYTOLOGIA Vol. 13, no.

50. 1963; Soukup, Biota : 260 & 312 Os) and 5: 35, 37, 117,

123, . ae 1964; Moldenke in Shreve & ag Ve sig & Fl. Son.
1259. 196h; Little & Wadsworth, U. S. Agr. Forest

ees seria ° or 9: 7879, fig. 226. 96h; Menninger Sea-

side Pl. 193 & 2 1965 Langman, . Guide Lit. Flow. Pl.

Mex. 160, 2h9, 356, 515-517, 63 33, 25, & 1010, 1964; Ang ely,

Bibl. Veg. Paran, 12 128 & 195. 196k; Moldenke, Résumé re 10: 5

3

(1964), 11: 5 & 6 (1964), and 12: 3 & 9. 1965; Moldenke, Phytol
gia 12: 6, 21, & 27. 1965; Go vel & vr, Fl. Bar-
bados 35, 356, & 468-69. 1965; Neal, In Gard. new rev.
ed » 721, & 726. 1965; Rambo, Pesquis. Bot. 21: ry ss &

«,» 720 P
[59]. 1965; Moldenke, Résumé Suppl. 13: 1, 2, & 6. 19

Little & Wadsworth (196) note that "The generic name (Cithar-
exylum) is Greek for fiddlewood. The English name fiddlewood rood. and
similar ones in French and Spanish for eur baa species refer to
the use for violins, guitars, and other m instruments. The
Spanish nam e péndula, meaning pendent, Pree ase the long slender
drooping 0. * angie Clusters of flowers and fruits

Aristeguieta (195) records the sant and verna pees es

Rincon 3 "palo guitarro", "pendare", "péndula", "totumilla",

, and "totumillo blanco", while Soukup (1962, 1563 —.
196h) sae "“choloquillo cimarrén", "mullahuayo", "quinielli
blanco" [with a question], "tasta", "tosta", and "turucassa" —
of these, "tasta" and * tosta" are applied also to Escallonia,
while "turucassa" is applied also to Porlieria and Rauwolfia.

It should be noted here that Hadongia Gagnep. was or
proposed by Gagnepain as a gems in the Bignoniaceae. The H.
Pittier 9967, originally distributed to herbaria as a species of
Citharexylum, is actually Guettarda discolor Rusby in the Rubia~
ceae.

breviations used on rs sy dates Sie pees of this paper

for — names of herbaria in ns are ac agr 2%
not accounted for = n Phytalogi 5: “inaise. (1955) and
(1958), will be found in the introduction to my paper

entitled Materials Govan a senflgaticoe of the genus Callic ‘
I" which follows the present one in the present jou pr maage

eral AFFINE D, Don akal
onal & emended bibliography: Sessé & Moc., Pl. Nou. HisPss
ed. 2, 96, pl. 417. 1893; M. Kunz, Anatom. Untersuch. Verb. 69—
& ar fig. 8. bes r. C. Standl., Co mee ip U. S. Nat. Herb.
23: 1238 & 121. 192k; H. B. Life & Works Pringle 158.
motels ee phytatogia 6: "7162282. 1958; Hocking, Excerpt.
aitons: "sessé & Moc., Pl. Nou. Hisp., ed. 2, ple 17.

it? MK » Anatom. Untersuch. Verb. "fig. 8

once sens te 17 in Sessé & Mocino's work (1893) Dr-
meas McV. in a letter to me dated October 11, 1963,
"Inclosed are two photographs that I took his summer,
of original plates made in Mexico probably in 1 by the artists

791,
of the Royal Botanical Expedition.....plate ml? of the Plantae No-

1966 Moldenke, Notes on Citharexylum 279

Hispaniae (cited on p. 96, ed. 2), with type-locality
Patiblines and/or Coahuayana, This is presumably the same species
as C. sessei D. Don, of which I saw a specimen at Oxford this
sumer." As yet I have not been able to examine the Oxford spec-
imen oes to here, My letter to Oxford offering to annotate
the ma

et
Recent collectors describe C. affine as a tree or as tree-like,
5—10 m. tall; trunk 20-25 om. in diameter at breast t height;
corolla lavender or pale-lavender to pacing: or purple, pale-
hairy at the orifice; fruit orange-brown when immature ure, b1ue-biack
ha

vannas &
orange" or "turning orange-red and finally blue-black when aikerten
The corolla is denaciben as "purple" on Alava & Cook 118) and Mc
Vaugh 14898, "lavender" on McVaugh 15697, "pale-lavender" on
Gentry & Gilly 10705, "violet" on Wilbur 2121, and "deep-blue" on
& Gilly 10826 26. Alava & Cook describe the fruits as "ber-
ries", but they are actually drupes. The plant has been found in
flowe: ower also in the month of March.
isotype, Andrieux 135, in the Delessert Herbarium at the
Conservatoire et Jardin Botaniques at Geneva was photographed
there by Macbride as his type photograph number 7877.
The Gentry, Barclay, & Arguelles 20252, distributed as C. af-
fine, is actually c. gleasonianum Moldenke.
~ Addi tioned citations: MEXICO: Colima: R. McVaugh 15697 (Mi).
Jalisco: Wilbur 2121 (Mi). México: Andrieux 135 [Macbride photos
7877] (N—photo of isotype); Hinton 9011 (Rf). Nayarit: Feddema
AAS (Mi); Gentry & Gilly 10705 (RF), 10626 (Rf); Re lcVaugh 111698
08); P Paray 2687 (Ip). Sinaloa: Alava & Cook 148 (iii); J. Gon-
zalez Ortega 53 5430 (Ip).

CIT

Addi
one s} ee at,
a 233 1237 & Liogenty 192h3 "Totdeckos, is Pipiologia ie "381-286.
Re Be rag ci describe this apg as a slender shrub, to 12
fect ¢ tall, white flowers, blooming in July, growing in are
5 at alti: of 1,00 to 1600 meters, fruiting also in Septem

Additional citations: MEXICO: Hidalgo: C. L. Lundell 12558 (Rf);
Moore & Wood 3845 (Ba). Querétaro: Paray 2169 (Ip).

TTHAREXYLUM AMAZONICUM Moldenke
Additional tithese ae Moldenke, Phytologia 6: 286—-287. 1958.

280 PATTOLOGIA Vol. 13, no. 4

An isotype, Spruce 211), deposited in the herbarium of the
Naturhistorisches Museum in n Vienna, was eo pent there by
Macbride as his type photograph number 3,3

Additional citations: BRAZIL: Amazonas: Spru ce 211) [Macbride
photos 34318] (W—photo of isotype).

CITHAREXYLUM AMBIGUUM Moldenke
Additional bibliography: Moldenke, Phytologia 6: 287--288.
1958; Langman, Select. Guide Lit. Flow. Pl. Mex. 515. 196k.

igen p peo
2 Cyppehsoowaia Moldenke, Phytologia 6: 288--289
sss). ead 7 73. 1959.
teinbach describes this plant as a agen 1--2 m, tall,
at white tar 6019) flowers, ends + 2000 neture altitude,

aii
Additional citations: BOLIVIA: Cochabamba: Steinbach 6019 (A)-
La Pag: Mandon 1,93 [Macbride ase 28393] (W--1610705—--isotype).
Santa Cruz: Steinbach 8572 (G).

CITHAREXYLUM ARGUTEDENTATUM Moldenke
Additional synonymy: Citharexylum argutidentatum Moldenke, Ré-
13: 6 7

1 ae ;
ditional bibliography: Mold denke, Phytologia 6: 289--290.
19685 Soukup, Biota : 172. 1962; Moldenke, Résumé Suppl. 13: 6
Vargas Calderon found this re ee growing at 3000 to 3650
meters altitude, blooming in January and February. The CArdenas
5h6h, distributed as Cc. S cetonmiaawan: is actually C. punctatum
Greemm,.
Additional citations: PERU: Cuzco: Vargas Calderon 207 (W—
1995350), 7684 (W—1987877) .

ITHAREXYLUM BERLANDIERI B. L. Robinson
opie ge bibliography: M. Kunz, Anatom. Untersuch. Verb. 70.
1911; P. C, Standl., Contrib. U. S. Nat. Herb. 23: 1238--1210.

icine U. S. Dept. . Agric. Handb. 41:
33 Moldenke, Wav titbese ‘ee 2 ‘ lL ya Résume
Supple ise 1. tee gia "290-291. 958; Moldenke,
A tional vernacular names recorded for this plant are
"encorba gallina" and "orcajuela". The plant has been found by
duous tropi

ridges, among brush on soil derived i resi igneous (basalt)
rock near arroyos, in badly disturbed brush on black clay soil

1966 Moldenke, Notes on Citharexylun 281

derived from basalt, and in soft shale at the mouths of canyons,
owering in n Septemher and October, Kenoyer & Crum call the
species a "forest tree", but Correll refers to it asa whee rte
shrub" and Gentry a "slender shrub", The corollas on C. L. Lun-
dell 10715 are described as "white", and the fruit on Correll &
Johnston 19774 as "orange-red". Readcnenict found the plant gr grow
re at ing a& 1350 a re altitude in San Luis Potosf{. Crutchfield &
"abundan’ Tamaulipas, but Graham & Johnston

14770 (Ip). Sinaloa: N&poles 10 (Ip). Tamaulipas: Crutchfield &
Johnston 5699 (Mi); H. S. aes it 6701 (Ca—-946182, Du-—356150) ;
oe Medrano 5 5 (Ip); Graham & Johnston 4082 (uit), 4398 (MA) a
44OL (Mi); Kenoyer & Crum | 3365 (Mi); Manning & 1 & Manning 53385 (Ip);
J. Tosynatt 86 Gol (Mi). :

CITHAREXYLUM BOURGEAUIANUM G: em.
Additional bibliography: . Standl., Contrib. U.
1s. 23: 1238 & 120. i925 ioldenke, ereboleeia 6: teal

ITHAREXYLUM BRA (A. Gray) A. Gray
Additional & emended synonymy: Citharexylum brac nan Gray
ex Moldenke, Suppl. List Invalid Names 2, in syn. 19. Cithar-
Sxylun brachyanyiun Gray ex Moldenke, Phytologia 6: 295, in syn.
Sphalm. 1958,

"pee he gg bibliography: M. Kunz, Anatom. Untersuch. Verb, 70.
1911; zee Standl., 2 contrib. Us S. "Nat. Herb. 23: 1237 & 1238.
192h5 He Works Pringle 87. 1936; Moldenke, Phy-
tolopia " Bem 19595 Moldenke, Résumé Suppl. 13: 6. 1966.

ent collectors describe this plant as a bush or shrub,
bri, °

ee Bs 5
Scarlet fruit, 6 mm. in diameter fruiting in October and Novem-
pe The Lundells refer to the fruits as "berries", but they

are actually drupes, The flowers are described as "white" on C.
ea Muller 3006. The plant has song Seer growing on limestone
Slopes and on thin limestone soil, in matorral vegetation, in al-
luvial ground with Larrea, and in’ savostieiea with Agave lechegu-
sila and grasses. The leaves on M. T. Edwards 292 are extra

erts open
on Larrea-Flonrensia deserts in Chihuahua. He calls it

282 PHYTOLOGIA Vol. 13, no. h

"common in Larrea-Flourensia desert", but "infrequent in desert
scrub on protruding basalt dikes" in Coahuila. Crutchfield &
Johnston — to it as "infrequent in desert scrub on limestone
hillsides."

Additional citations: TEXAS: Zapata Co.: Lundell & Lundell
we (Rf). MEXICO: Chihuahua: C. H. Muller 3333 (ca 7o1s0L,

Mi, Rf). Coahuila: Crutchfield & & Johnston 5871 (Rf); I. M. John
pe, 7048 (Ca-——7h126)); M. C. Johnston 5871 (Mi); C. H. [. Muller
2008 (Ca—71961, Mi, RE), 30 3033 GH, Rf); Salgado s.n. (18.1.
1961] (Ip); & Mueller 9 9 (Mi, st). Durango: Correll & John-
ston 20217 (Rf). Nuevo Leén: M. T, Edwards 292 oeaahtls

e

5608 (Ip), 7636 (Ip). Zacatecas: See agd Xolocotas es an
J. Reotonntcl 9323 i 9323 (Ip, Ip); Shreve 8590 (Ca—731757), 9203 (Ca—
731694) .

CITHAREXYLUM CAUDATUM L.

Additional synonymy: Citharaexylum caudatum L. apud L. C. Rich,
Act. Soc. Hist, Nat. Paris 1: 111. 1792. Cytharexylon caudatum
L. apud Macfadyen in Hook., Bot. Misc. 2: ae 1831. cutharen =

caudatum L, apud M. Kunz, Anatom. Untersuch, Verb. 70, sp

Additional & emended bibliography: L. C. Rich., Act. Soc. Hist.
Nat. eete 13; 17925 hoger Sg in Hook., Bot. Misc. 2: 115.

Jahrb. 2: 170. 1908
are Verb. 70. 1911; Britton & P. Wils., Sci
pom iy ae ee 1925; Robledo, Lecce. Bot. 2: 498. 1910; Neal
‘- Phytologia 7: The

. e es Sicae. Handb

1964; Moldenke, Résumé Suppl. 11: 6. 1964; Neal, In Gard. Hawail,

new rev. ed., 726, 1965, Moldenke, Résumé Suppl. 13: 6. 1966.
Additional illustrations: Little & Beer E U. S. Dept. Agr
Recent collectors describe this plant as having a trunk to 25

Bartlett & Lasser as "orange, le". Allen refers to
e Er

Moldenke. Steyermark 38920, from Guatemala, does not appear t?
be typical either -- the collector ‘Aeeuetae its leaves as mrich-
green above with orange-brom midrib, age n beneath with

light-brown mazid, ib, frett spray pendent, fruit shiny". P. H. Ab
len 3597, from » has very broad leaves. Fee & Miller 2

has wood =i sanples mounted on the herbarium sheet.

1966 Moldenke, Notes on Citharexylum 283

Material of C. caudatum has been misidentified and distributed
in herbaria as C. ilicifolium H.B.K., C. schottii Greenm., C.
spinosum L., Byrsonima crassifolia = (Le) H.B.K., and Picramnia an-
tidesma Sw. On the other hand, the Shank & Molina R. 270, dis-
tributed as Citharexylum caudatum States is actually C. | c. hexangulare

Little & Wadsworth (1964) describe C, caudatum, as it grows in
Puerto Rico, as follows: "Shrub or tree of upper | upper montane forests

hit
unequal spreading corolla lobes and borne on short stalks 1/16
inch long; gad (3) shiny pear-shaped or elliptic fruits nearly 1/2
h long gis from green to orange brown or black, with cup-
like caly at b

‘g h

inner bark light brown and bitter. The ong narrow twigs are light
gray brown, with pr

"Petioles are 1/--3/) inc Blades pred inches long
and 3/4--2 1/4, inches wide, orien tiisbeniel a and with edges
Sometimes turned under a gery slightly shiny green above and
dull r beneath. On the lower surface are many minute in-
conspicuous pa visible with a lens

wi ae clusters penoomes) are 1 1 1/23 pebes 2908, terminal

e

and lateral, bearing lightly fragrant flowers.
shaped calyx as about 1/8 4 8 inch long and broad, minutely S-toothed;
the white f corolla with tube nearly 1/4 inch long and

hairy

5 lobes 1/8 inch Trig; stamens nearly 1/8 inch long inserted on
Corolla tube; and greenish pistil 3/16 inch long with 2-celled o-
vary, slender style, and slightly 2-lobed stigma.

he fleshy fruit ts (drupes) slightly 2-lobed have cuplike calyx
1/8 inch lo at base. They contain 2 agen ce yon A
brown nutlets ef ee tee. each l-seeded. Flowering and
ing throug

how
"The light peeves wood is used for posts and fuel. Also a
honey plant,

"Upper mountain forests and dwarf forests of mountain peaks in
Puerto goes
T ord the additional common names "café cimarrén", "café
roars ? “Wfiddleweod", "péndula de sierra", and "racemose fiddle-

dditional citations: rege Chiapas: Matuda 3771 (Mi, Rf),
316 (Mi). Tabasco: Matuda 3066 (Mi). GUATEMALA: Izabal: Steyer-
mark 38920 (W—19)958). BRITISH HONDURAS: Gentle 1793 (Ld, 8),

28h, PHYTOLOGIA Vol. 13, no. h

8079 (Id), 8488 (Ld, S). PANAMA: Canal Zone: P. H. Allen 1836
(i); Bartlett & Lasser 16308 (Ii), 1631h (Mi). Colén: P. H. Al-
len 3597 (Ca— (Ca—790799, W-—1976267). Panam4: Duke 1598 (ca), 4599
(Ca). PEARL ISLANDS: San José: C. 0. Erlanson on 23 ; (Sa
Mi), 310 (Mi, N); Harlow 5 (W—~2398301); 1. M. Johnston
202),005). CUBA: Las Villas: R. A. Howard 520), ee BY; etn Leén
& Jack 1395) (W—~228943). Oriente: Linden 1789 (P); C. Wright —
T3e [1860] (Cca-936807). JAMAICA: Bancroft & Miller 3 (W—
1555633) 3 Harris & Lawrence C.15508A (Ca——98h476)5 P Proctor &
Stearn 1149 (W--2351899); Wolle s.n. (Tl). HISPANIOLA: Haitd:
- H. Bartlett 17363 (Mi). “PUERTO RICO: Alain 10350 (G, 1, N,
rar 10733 0733 (G, N, N, §); 3 Webster & Miller 8713 13 (8S). PO POLYNESIA
TAN Is alm: A, R. Cooke s.n. 1. {Manoa trail, 9/26/SU
(St); Pearsall s.n. [Pauoa Flats, Aug. gs 1, 1953] (BL). CULTIVATED:
Hawaiian Islands: © Caum s.n. (dca 5, 1938] (Bi); Degener & Mura-~
a 20075 (W--1991209); F. E. Egler 37-90 (Bi); Potter & Miya-
sen. [11/26/57] (Bi). Peru: Klug g 355 (P); Martinet , 693
ye a tP), Geis (P, PB; Fos

CITHAREXYLUM CHARTACEUM Moldenke
— bibliography: Moldenke, Phytologia 6: 310-~312.

enberg found this plant growing in a pe ropobad at 580
meters altitude, but misidentified it as C. quitense Spreng.

Additional citations: PERU: Province ni teataet Ellenberg
1489 [Huasimo ntrdl. Suyo] (Ut—163209b) .

ITHAREXYLUM DAWEI Moldenke
Weteemes bibliography: Moldenke, Phytologia 6: 317-~319-

Lasser & Foldats describe this plant as a tree 5 m. tall, with
"frutas maduros, drupas, morado oscuro", fruiting in February -
Material has been misidentified and distributed in herbaria as Ce
decorum Moldenke. On the other hand, the Aristeguieta 793 193, ¢ dis-
tributed as "Citharexylum cf, dawei Moldenke", is actually Co
spe ecen eum Pittier.

dditional citations: VENEZUELA: Falc6n: Lasser & Foldate 3087
(ve=36553). et

CITHAREXYLUM DECORUM Moldenke
Additional bibliography: Moldenke, Phytologia 7: The 3 1999 °
Brito describes a cos -patty as a thee, 12 m.

branches for 9 m., fl in June, and called a mpause” “rhe

Bernardi rnardi 6739, cited ‘ag me in the id Bisco given pe Bag proves,
on closer exam examination, to be C. poeppigii Walp. aa ?
Lasser & Foldats 3087, distributed as C. decorum, is actually iyo
dawei Mold enke, while Steyermark 86556 i is C. See coeum vars

1966 Moldenke, Notes on Citharexylum 285

brittonii Moldenke,
Additional citations: VENEZUELA: Vargas: Brito 4 (Ve).

ITHAREXYLUM DENTATUM D.
Additional bibliography: "olde enke, Phytologia 6: 332—~333.
1958; Hocking, Excerpt. Bot. A5: 43. 1962.
An isotype, Rufz s.n., deposited in the herbarium of the Botan-
isches Museum at Ber rlin, was photographed there by Macbride as
his type photograph number 17592, but is now most unfortunate
dest Hbenee

Additi citations: PERU: Ancash: Rufz s.n. [Maccos; Macbride
photos 17527 (W—~photo of isotype).

ITHAREXYLUM DISCOLOR Tures.
Additional synonymy: Cithar exylum discolor discolor Urb. & Ekm, ex Mol-
denke, Résumé Suppl. 10: 5, in syn. 196k.
Additional Siasscareniees Moldenke Jor epuepins 6: 334-336.
1958; Moldenke, Résumé Suppl. 10: 5. 196k.
ent collectors describe this ethos as a shrub, 3—5 mn.

wi
green » growing from 630 to 1700 meters altitude, flowering
- in May, and fruiting page August.
Additional citations: HISPANIOLA: Dominican Republic: R. A
Howard 12285 (ii, N, S), 1230h (Mi); Jiménez 1002 (W—2350)01).
Haiti: Leonard & Leonard 15066 (W-—1h53385, W—1453386) .

CITHAREXYLUM DONNELL~SMITHII Greenm

Additional ae octinceiai soem Greenm. ex
Moldenke, Résumé Suppl. 3: 30, in sy
i ofdéttional Mahila eenipiags jusiaetis, Prpahogia 7: 7h. 1959; Mol~
e Résumé Suppl. 3: 30. 1962.

Recent ser seisibe this plant as a tree, 20-60 feet

j trunk erect, to 0 cm. wide at the base, 36.5 cm. wide at
breast he height; bark dark-gray or dark gray-brown, shaggy; leaves
enamel eeeely thickened and stiffened, dark opaque-green,
wi

rowing in
Scil along open roadsides in the upper limit of the tropical zone
er in gray loam of pastures, at altitudes of yet to 6600 feet,
in January in February and April, knom as
"chorrito"., The Stendliiy & «Pasi V.266h, cited in Phytologia 7:
74 (1959) as from Sonsonate, is actually from Almachapén, El Sal-
vador, The Stork 3283, Mieisitiees as C, donnell-smithii, is ac-

Lae eraentun Green. xander 1217 (W,
dditional citations: MEXICO: Chiapas: E. J. Ale: r

N,N, Z). GUATEMALA: Sacatepéquez: P. C. Standley 64953 (Mi). EL
SALVADOR: La Libertad: M. C. Carlson 228 (Ca--703661). Santa Ana:
Allen & Armour 6798 (Mi). COSTA RICA? Alajuela: A. Smith P.2337

286 PHYTOLOGIA Vol. 13, no.

(Ca—12726); A. F. Smith 26 [Stork 126] (Ld, Mi). San José:
Tonduz 1419 (G).

HAREXYLUM DRYANDERAE Moldenke
Additional bibliography: Moldenke, Phytologia 6: 30--3)1.

958.
Additional citations: COLOMBIA: Valle del Cauca: Dryander 2362
(Bm--isotype) .

CITHAREXYLUK ELLIPTICUM Sessé & eee

Additional bibliography: M. , Anatom. Untersuch. Verb. 71.
1911; P. C. Standl., Contrib. » Bs "Nat. Herb. 23: 1238 & 12h0.
192h; Moldenke, Phytologia 6: 342—345. 1958.

Recent collectors describe this plant as a shrub or low shrub,
1-~3 m. tall, with white flowers and orange or orange-red fruit,

11K3 (MA), 2706 (N, 2); Paray 1938 (Ip), 2869 (Ip); Paxson, Web-
ster, & Barkley ey 17M620 (Au—~123187, Au--169676) ; Sauer & Gade
3110 (Mi). CUBA: Matanzas: Leén 13905 (W--2289)h2) .

CITHAREXYLUM FLABELLIFOLIUM ee as
Additional bibliography: Standl., . U. S. Nat.
ecah 23: 1237 & 1239. I9eh; Micldenke, cian i The 19593
Moldenke in Shreve & Wiggins, Veg. & Fl. Son. Des. 2: 1259 196h.
ren agate collectors describe this plant as a shrub or seit

it er
ation with the following dominants: Pachy Hato pring i (Se “ta

——e & Rose, Bursera spp., Jatro iia sop., Fours
s Nash, Fore hammerta watsoni Rose, and Machaericereus ee
Geel } Britton & Rose eet ey
citations: MEXICO: Baja California: J. H. Thomas Si2t
cuieshaorAh: Sonora: H. S. Gentry 721 (Mi); Shreve “ais 0a Mi)-

MONSERRATE ISLAND: R. V. Moran 3886 (Ca- —— Ly) .

HAREXYLUM FLEXUOSUM Join & pert D. Do
Additional synonymy: Cithare ao entice um Humb,. & Kunth ex

denke, aphy: Bot. Voy. Sulphur 154. 18h65
ae Phytologia 6: 319-353. "1968; Soukup, Biota h: 172+

This re has been found growing at ),00 meters pel 2 |
August. raga (1962) records the common
‘ohtlowdan Cimarron", An isotype of C. spinosum, Humbolc® Humboldt & & Bor

1966 Moldenke, Notes on Citharexylum 287

pland 3687, oo in the herbarium of the Botanisches Museum
at Be rin, was photographed there id Macbride as his type photo-
_ number 17601, but is now destroyed. The type of Rauwolfia
» Rufz & Pavon 88, reste in the Madrid herbarium,
was as istered there by Mi Mac ride as his type photograph mt

29685. The Soukw ukup 498, distributed as C. flexuosum, is ac
Duranta skottsbergiana Moldenke,
” Additional citations: PERU: Arequi nberg 2691 (Ut—

163206b) ; Rose & Rose 18988 (Wao 962572) ‘ ee Humboldt &
Bonpland 3687 (Herb. U. S. Nat. Arb. 196719; Macbride photos
17601] (W-=photo). eitiniec Rufz & Pavon 88 (7/78; Macbride
Photos 29685] (W-~photo).

ITHAREXYLUM FRUTICOSUM x

Additional synonymy: tharoxylon fruticosum L. ex Moldenke,
Résume Suppl. 3: 30. 196 >.

Additional bibliography: Griseb., Cat. Pl. Cub. 216. 1866; M.
Kunz, Anatom. Untersuch. vores Tl. 1911; E. L. Little, U. S. Dept.
Agr. Forest Serv. Agric. Handb. 1: 111. 1953; Moldenke, ee: 20-
el. 19555 Moldenke, Phytoloris 7: 75. 1959; Jimenez, List
erna: 61; Menninger, Flow. Trees World 292, wert Hocking

C)
og in herbaria as C. spinosum L. Britton & Wilson (1925)
a hes to synonymy under C. fruticosum, call-

ing ms a a Saventin
Recent So arg desaeiie C. fruticosum as a spindly sparsely
anched shrub, 1 m. tall, or a a tree, 2.7--9 m. tall, the petioles

orange-col lored, the ‘oordlias white, and the trait reddis h-orange

sbabd ting coastal limestone bluffs, and found at altitudes to “to 600

appears this statement: "Gifford says that rp is the true nene
wood, It — not agree with Small", There are a few hairs in
= f this specimen. Webster refers sks ins species as “a
Common" os Orie nte, Cuba,
Little & Wadsworth (1964) describe the plant as follows: "This
Small tree is easily recognized by: (1) o nae mostly elliptic,
or

er small
5 slightly unequal seeteing corolla lobes, borne <a t
talkless on are ing axes; (3) nearly round, shiny,
reddish-brown or black fruits 3/8 *inch or less in diameter; and
(b) rough, Meht. brown bark, becoming much fissured and s

288 PHYTOLOGIA Vol. 13, no.

in long narrow
"Evergreen shed or small slender tree 10--,0 feet in height
and to 1 foot in trunk diameter, The inner bark is whitish and

nodes, slender, light brown or green, bec oming gray and finely
fissured. Petioles are 1/2~—1 inch long. Leaf blades are 3—7
inches long and 1—2 1/2 inches broad, short- or donee at
both ends or sometimes rounded or notched at apex, t es

ns pist
celled green ovary, slender style, and slightly 2=lobed stigma.
The drooping clusters of fleshy fruits (drupes), sometimes abun-
dant, are reported to be sweet and edible. Each has a stone
pcnegh iy palseudt into 2 2=seeded nutlets. Flowering and fruiting
through the

"The sareibed is light brown ss and the heartwood is
reddish. The wood is very hard, heavy 7 {epocttie gravity 0 Onli
and strong. Considered a good fenc epost i to Rico
wood of this and related species ne been ‘ed louitali for
furniture, construction, and violins, guitars, and other msical
instruments

"Sometimes planted along highways and fences and in g
an ornamental. Also a honey plant. This Mabe fe is yan gaat gti

dditional
2h, os (Hi—-18)16)). Dade Co.: Tidestrom 698 (ca--8 ony
Totten & Gifford s.n. [December 30, 1940] (Hi--22775)- Ble

Lépez Figueiras 2952 (W—-2228995) ; Morton & Alain 9194 (W—
278); G. L. Webster 1071 (ud). “Province undetermined:
383 (P). JAMAICA: West & Arnold 910 (N). HISPANIOLA: Doninioal

1966 Moldenke, Notes on Citharexylum 289

Republic: B. Augusto 1061 (N); Howard & Howard 9757 (B). PUERTO
RICO: Re a Finch 70 70 (Mi); A. A. Heller 4421 (P); Ss: Sintenis 720b
(P), 1915 (1 (P), 20% 2085 (P). WINDWARD ISLANDS: M Martinique: He: Herb. M Mus.
Paris s.n. (P). Ronde: Re A. Howard 10733 (N). St. =r Web-
ster & Miller 9356 (Mi). TOBAGO: Pu Purseglove P.6363 (

229296). VE! VENEZUELA: Sucre: Ari Aristeguieta & | & Agostini "bes (N).
BRITISH GUIANA: E. L. Little 16710 (Z).

CITHAREXYLUM FRUTICOSUM f. BAHAMENSE (Millsp.) Moldenke
Synonymy: Citharexylon bahamense Millsp. in Britton, Bull. N.

apud Prain, Ind. Kew. Suppl. 3: 3. re

Bibliography: pee Bull. N. Y. Bot. : 450—L51.
1905; Prain, Ind. ag 1. 3: 43. 1908; oidéake, Prelim
List Invalid Name < 19; i Moldenke, Alp t Invalid Names

ae cae Hoddentes, Résumé 253 & 255, "19555 “woldenice, Phytologia
19

his ce pubescent form of the species was hitherto
Tegarded by me as var. villosum (Jacq.) 0. E. Schulz, but I feel

tified and distributed i n herbaria as C. fruticosum L. and as ce.
Sriticosum var. wtlicgts Gee) ne E. Scmlz.

The type of the form was c ted by Nathaniel Lord Britton
and Lewis Jones Knight ese Cao 233) in a coppice on Farrington
Road, New Provi lands, on August 2, 190, and is

nce, Is
deposited in the Britton Herbarium at the New York Botanical Gar
den, The form has been found growing in coppices, copses, and
hillside thickets » on hills, and along roadsides, flowering in
rhegibaee f August, October, and November, fruiting in August and
ctober,
Citations: BAHAMA ISLANDS: New Providence: Britton & Brace 233

ton whi
» Z——photo of type). VIRGIN ISLANDS: ste John: Britton &
Shafer 21,9 (MN, ¥—756261). Tortola: Fishlock 198 (A, G, K, N, W—
MOP) Wein Gocta: Fishlock 73 (N).

CITHAREXYLUM FRUTICOSUM var p SEE TORE Moldenk
Additional bibli raphy: i Verb. a1. 1955; Moldenke,
Phy tologia 6: 366-— 8 (1958) airs 13: 28—285. 1966.
n Trinidad this plant is said to be "widely distributed in
ies districts, but ner Senin: Endemic", Actually it is
ndemic to

rinidad, occ °
sauela, Cowan & Forster describe it as an "infrequent tree with
Pendent, branches", growing at 100 meters altitude, flowering in
t - Delgado calis it a shrub, 5m. tall, while Steyermark re-
8s to it as a tree, 10 m. tall, with subcoriaceous leaves, deep-

290 PHYTOLOGIA Vol. 13, no.

green above, paler-green beneath, the calyx and rachis pale-green,
known as "totomillo sabafiero" in Venezuela. Material has been
misidentified and distributed in herbaria as C, decorum Moldenke
and as C. spinosum L. =

amg citations: TRINIDAD: Cowan & Forster 1278 (N)-

VELA: Bolfvar: Steyermark 86556 86556 (Ve), 88236 88236 (N), 1), 88829 (N);

Delgad o 421 (Ve).
CITHAREXYLUM FRUTICOSUM « SMALLII Moldenk

Additional aikiievertets “Moldenke, Phytologia 6: 368 & 383.
1958; Hocking, Excerpt. Bot. A5: 43. 1962.

CITHAREXYLUM FRUTICOSUM var. SUBSERRATUM (Sw.) Moldenke
Additional bibliography: age age Pl. Cub. 216, 1866;
Moldenke, Phytologia 6: 383--386.
citations: HISPANI rand ws ae Republic: Vélez
3738 (W-1957415) .

IT ICOSUM var. SUBVILLOSUM Moldenke
Additional aaaay ography: Moldenke, Verb. 21—22. 1955; Molden-
ke, Poi be gia 6: 386--388. 1958.
ditional poet omen FLORIDA: Big Pine Key: Killip 40702 (Ca--
S377) VIRGIN ISLANDS: St. Thomas: Eggers 90 (W——195920 207); De
Hummel s.n. [1.1I.1958] (Go, S).

CITHAREXYLUM FRUTICOSUM var. VILLOSUM (Jacq.) 0. E. Sch
le are iae O) See ; Cat. Pl. Cub. 216. "18664 Ae
0. 1878; Moldenke, Phytologia 7:

15 (i969) and 13) 289. 2 ; ass ps

Recent collectors describe this plant as a small tree, to 15
or 20 feet tall, with white fragrant or very fragrant flowers, and
"leaves shining tear Apo below", inhabiting rocky hill-
slopes, rocky or oe, limestone hillsides, to 3000 reat altitude,

S

re
The very narrow-leaved form of this variety, repteoe by
Millspaugh's C, bahamense, seems to me now to be worthy of nomen
clatural recognition. Therefore , the Britton & Brace 233, Britten Britto’
& Shafer 2h9, and Fishlock 73 & 198, cited by me in my “monograph
as var. villosum, are now regarded as f. bahamense (Mills Millsp.) Mol-
denke instead — the first-mentioned being the type collection.
The flowers on R. A. Howard 12068 noth described as "white" and

an
(Mi, N). Haitd: Holdridge 11,28 Mi); G. R. Proctor 106%
2225297). CULTIVATED: Senegal: an Mim).

ITHAREXYLUM FULGIDUM Moldenke 4
Additional bibliography: Moldenke, Phytologia 6: 394--395+ 19%

1966 Moldenke, Notes on Citharexylum 291

6k.

Moore describes this plant as a shrub, to 10 feet tall, with
red to black fruit, found along roadsides in pine woods, at 2000
meters altitude, fruiting in Oskeban,

Po pga citations: MEXICO: Hidalgo: H. E. Moore 5308 (Ca——
918892

CITHAREXYLUM GLABRUM (S. Wats.) Greenm reget
Additional synonymy: Citharexylun giabrum Greenm, ex Moldenke,
Résumé Suppl. 13: 6, in syn. 19

Additional bibliography: M. Kunz, Anatom. Untersuch. Verb. 71-
t2. 19113 P. C. Then Contrib. U. S. Nat. Herb. 23: 1238 &
12h0, 192); H. B. Davis, Life & Works P e 72, 668, & 669.
1936; Moldenke, me igs 6: rian Fe: 1988; Moldenke, Résumé
Suppl. ey Bes.

Diquet [whose mare is pga 2 a * — herbarium
labele] records the vernacular "chilcahua s plant.
Paray found the species ts sage at "9000 a eters ae fan

Additional citations: MEXICO: Guerrero: Hinton ae (Rf). Jal-
per Diquet s.n. [Decembre 1912, Huejotitan] (Mi). Na

Paray 3395 (Ip, Mi).

ITHAREXYLUM GLAZIOVII Moldenke
Additional bibliography: Moldenke, Phytologia 6: 398—100.

Ducke describes the fruit of this haere as "frutos maduros
Yolen da and collected it in fruit i ruary.
Additional citations: BRAZIL: Cear4: "pucke 2532 (W—23L37h2) .

tional army persia "Woh denke Phytologia .
1958; Langman, Select. G Tian: Pls tex, 50E: 1964; Mol-

or montane shrub, with white flowers, growing on limestone.
Additional citations: MEXICO: Puebla: Gen , Barclay, -
elles 20252 (W—2151070); J. Rzedowski 1888 (Ip).
CITHAREXYLUM HERRERAE M
Additional aceente: “Moldenke, Phytologia 6: 01—103.
1958; Soukup, Biota l: 172. 1962.
Soukup (1962) records the vernacular name "huairuro"., An iso-
ype, Herrera 677, Nate ee in the herbarium of the Botanisches
Museum at Ber Berlin, was photographed there by Macbride as his type
nee number 1759 , but is now destro:
citations: U: Cuzco: Herrera “677 [Macbride
Photos 17598] (W—photo of isotype).

ITHAREXYLUM HEXANGULARE Greenm.
Additional bibliography: M. Kunz, Anatom, Untersuch. Verb. 72.

292 PHYTOLOGIA Oe

1911; P. C. Standl., Contrib. U. S. Nat. Herb. 23: 1238
192); Moldenke, Phytolo gia 6: 403-106 (1958) and 13: hat es.
Recent co

ectors describe this species as a s m, tall,
or a tree, ° with horizontal whorled branches » Leaves
aay membranous, deep-green above, dull-green beneath, flowe
fruit r on limestone rock a-

» very grant, ’
bey riversides , on steep mountainsides from the oak zone to
of the pines, and in Rhiscphors peeite from 1 to 2250 meters
altitude, flowering in July and November, fruiting in October and
December . McVaugh describes it = "occasional along old roads"
in Jalisco. Additional vernacular names are "frutillo" and
- ae gon paves ae gage age ks Ceca distributed in

r a as C. ca un L. .

Additional citations: MEXICO: rer 5 3861 “(d).
Chiapas: Gilly & Hernandez Xolocotzi 193 (Mi). Jalisco: R. Mc
Vaugh 20632 (Mi). Oaxaca: J. V. Santos os 3618 (Mi). Quintana Roo:
Lundell & Lundell 7679 (Rf). Tabasco: Matuda 3562 (Mi). Vere
Cruz: Dodds 110 (Mi); J. Ruedowski 17175 (Ip). GUATEMALA:

Petén: Contreras 1831 (Ld, S); Steyermark 5851 (W~19l,9698) «
BRITISH HONDURAS: Gentle 4,858 (La). COST. COSTA RICA: Limén: Shank &

Molina R. 4270 (W--2085192).

HAREXYLUM HIDALGENSE Moldenke

parece, synonymy: Citharaxilium hidalgense Moldenke, Résumé
Suppl. 13: 6, in syn. 1966. Cit lum hidalgense Moldenke,
Résumé Suppl. 13: 6, in syn.

Additional bibliography: Moldenke, Phytologia 7: 75. 19593 Mol-
denke, Résumé Suppl. 13: 6. 1966.

The stems on Hinton 15606. are pubescent and the plant was in
flower when collected; on C. L. Lundell 12642, collected while in
fruit, the stems are glabrous. Moore found the plant growing on
limestone ledges and in streamside thickets, at 1800—2000 meters

or lavender flowers, and fee. fruit, flowering and fruiting in A~

ee citations: mextc CO: Hidalgo: M. Martinez s.n. [Zacu-

tipén, May-—June 1940] (G); H. E. Moore 2631 1 (Ca——919 332) 5
eich 8969 (Ut--28)21a—isotype); Vela G. 569 (Ip). Méxicot
ater zac ee ao eee Hinton 15606 (Rf). Puebla: C. Le

CITHAREXYLUM HINTONI Moldenke Be Og
Additional bibliography: Moldenke, Phytologia 6: 08—
19585 5 Langnan, Select. Guide Lit. Flow. Fl. Mex. 515. 196h.
dditional citations: MEXICO: México: Hinton 751) (Rf).

xCIT

EAREXTLIM HYBRIDUM Mo er
Additional bi abbtaprioter Woldenke, spat 6: Wu. 1
Additional citations: TOBAGO: W. E. Broadway 1,064 (P—isotyPes

1966 Moldenke, Notes on Citharexylum 293
P—isotype) .

CITHAREXYLUM ILICIFOLIUM H.B.K
Additional bibliography: Schwencke » Zytol. Untersuch. Verbenac
17 . Oe 1931; lo ava shh Zytol. serves 6 & Psp b 19355"
- Dahlgre: ve t. Tidsk. 32: 231. 1938;
ota 3 1: "80. 1956; tanks, Phytologia 7: 75 (1959) is Cy) P55,

Recent collectors describe this plant as a shrub or small tree,
past r 5m, tall, with white flowers, and dark cherry red-black

cate » fruiting in February and November. Ewan encountered it
in cultivation by a lake. A specimen of Herb. Humboldt s.n.

number 17593, but is now destroyed. The Caum s.n. [June 5, 1938],
distributed as C. ilicifoliun, is actually C. Ce. caudatum L.

Additional citations: ECUADOR: Aguay: Gilar 6 (N). Pichincha:
Barclay, Juajibioy, & Tinajero 7821 (Mi); Herb. Humboldt Sen.
(Chillo; Macbride photos 17593] (W—photo of ames de qi: Eeingres
951 (Gg—35486,). CULTIVATED: Ecuador: Ewan 16,13

xCI aa JAMAICENSE Moldenke
ional bibliography: Moldenke, Phytologia 6: 16~-117
958). an tos 282. 1966,

- Bartlett 17363, cited under C. caudatum L., is not
at he wpa of that species. Its leaves are too pointed and
small, and the venation is too much impressed above and prominent
beneath, It may well represent the present hybrid and therefore
represent its first record from Haiti.

CITHAREXYLUM JORGENSENII (Lillo) Moldenke
Additional synonymy: Citharexylon jo rgens enii (Lillo) Moldenke

Spud T. Mey., Op. Lill. 10: 126. 1963.
Addi tional bibliography: Moldenke, Phytologia 6: i718.
me sis Op. in. : 50 & 126. 1963; Moldenke,

specimen of Schuel 191, deposited in the herbarium of the Natur-
‘istorisches Museum at Vienna, was photographed there by Macbride
as his type photograph number 31,315.

Additional citations: ARGENTINA: Jujuy: Schuel 197 pie) «
159127h) Jorgensen 1,7 en 47 (W——92165l,--isotype) ; Venturi i 2961

CIT

EAREXYLUM JURGENSENI Briq.
. Addi tional bibliography: Moldenke, Phytologia 6: )18—20.
9585 Langman, Select, Guide Lit. Flow. yl. Mex, oa & 1010. 196).

294 : PHYTOLOGIA Vol. 13, no.

McVaugh describes this plant as a tree, 10 m. tall, the trunk
15 cm. in diameter at breast height, the inflorescence
calyx-tube red-lavender distally, and corolla blue~lavender, the’
thr

mally,
2% of moist tropical forests, with oak forests on the ridges
with sree, Inga, and Sideroxylon in the barrancas, at alti-
<a . 800 1200 meters, gp in September.
The hep hanno Jurgensen en 259, sited in the pelea
a. at the Conservatoire et oe) aie D cotet cian
pres photographed there by Macbride as his type waategrenl bitte Fe
So faye citations: MEXICO: Nayarit: R. McVaugh 16920 (Mi).
Oaxaca: Jurgensen 259 [Macbride photos 21623] (N—photo of type).

Mag prose UM KARSTENI Moldenke
Addi ti a ea Citharexylon karsteni Moldenke, Résumé
Suppl. 13: 6, 66.

syn.
Addi tional halcel Beaaair Moldenke, = Na ans 6: 420-22.
ee vient Résumé Suppl. 13: 6.

sotype, Triana 2071, deposited oe the ~seagerns* Herbarium

ue te ervatoire et Jardin Botaniques was photo-

pei there by Macbride as his type photograph aban 28395.
Additional citations: COLOMBIA: Cundinamarca: Triana 2071

[Macbride photos 28395] (W—photo of isotype).

eed Fen tae . LANCEOLATUM Moldenke nee
° ee Citharexylon ee var. lanceo.
ea oer 13: 6, in syn.

Additional Dinhograpt | Moldenke Piytoogia 6: 22——h23.
weg —— Résumé S eS 6. 1

reports "fruit ee “ ous be first orange, na

ack b otivon by 2-seeded, resembling Rhamms" and found the plant
growing on a dissected plateau (badlands).

Additional citations: COLOMBIA: Huila: H. L. Mason 13799 (Mi)-

CITHAREXYLUM KERBERI G
Additional bibliogr ster Contrib. U. S. Nat.
ines 23: 1237 & 1239. 192h; "Yoldenke, sar Papeckopt 6: 23—h2h.

CITHAREXYLUM KOBUSKIANUK Moldenke
sheet tional bibliography: Moldenke, Phytologia 6: 424—li25+
z describes the corolla of this plant as white, and io

the srestee along roadsides at 1750 meters altitude, flowering +"
June, and called "veneno de burro". Material has been misidenti-
fied’ and distributed in herbaria as Duranta mutisii

Additional citations: PERU: La Libertad: A. Lopez 1520 2 1520 (2)5
Vargas S. 39 (W—2253623).

1966 Moldenke, Notes on Citharexylum 295

ITHAREXYLUM KUNTHIANUM Moldenke
Additional biblio pg Moldenke, Phytologia 7: 75. 199) «
An isotype, Humboldt & Bonpland 5898, deposited in the

stroyed. Material has been misidentified and distributed in her-
baria as Cordia sp. and as Croton SP.

Additional citations: COLOMBIA: Cauca: Alston 8300 (S)5 Snei-
dern 2816 (W--2103916). Valle del Cauca: Humboldt & | & Bonpland —
5898 (Herb. Willdenow 11476; Macbride photos 17602; Herb. U. S.
Nat. Arb. 197895] (W—photo of isotype); Killip 1126) (G).

CITHAREXYLUM LAETUM Hiern
°F igen bibliography: Paran. 7: 11. 1957; Mol-
Phytologia 6: SL opeksee wi9365 A Aree , Fl. Paran. 16: )5
(19603 and 17: 6. 1961; Angely, Bibl. Veg. Paran. 128. 196).

TTHAREXYLUM pettus Moldenke
Additional bibliography: Moldenke, igbeans com et ac ore
and 6: ))8. 1959; Woldenke, Biol. Abstr. 33: 186

a ee
tall, the trunk erect, 50 cm. wide at the base or os 60 cm
at breast height, "The * bark light-brom, forming long elevated
Strips, th the

Side, at aatAtadae of 2100 to 2700 meters, flower sigesbys,
and fr in February. In oveier: 1953, Steyermark noted on
a sheet of Tucker fon "probably new species s but flowers necessary
for determination tion

Addi tional citations EL yan ge Bay same Tucker 1049
eee pt Ca~~388 Rf). COSTA RICA: Alajuela: A. Smith P.2397
(Ca—12715) , piacd ey Skutch 51,87 (#2351973)

CITHAREXYLUM LAURIFOLIUM H. Hayek
Additional bi eae age Nessie. Biota 1: 180. 1956; Moldenke,
Mhytologia 62 lil,9—-50.
t;

Botanisches Museum at Berlin, was photographed ae oui by Macbride

*s his type photograph number 17594, but is now destroyed.
Additional citations: PERU: Puno: Weberbauer 873 [Macbride

Photos 1759),] Sccacts! of type).

CIT RINUM Van Hutte

HAREXYLUM LIGUST
Additional bibliography: P. C. Standl., Contrib. U. S. Nat.
Herb. 23: 1238 & 1241. 192h; H. si Davis, Life & Works Pringle 217.

296 PHYTOLOGIA Vol. 13, no. 4

1936; Moldenke, page ates 6: pag Tsar 19995, ne, Taxon 9:
1960; elect. Gui . Flow Mex. 715 & 1010.
spk Woldenke, a Wer Suppl. 3: Pat 1966.
plant has been collected in anthesis in June. Material

has been misidentified and distributed in herbaria under the
name C, lucidum Schlecht.

Additional citations: MEXICO: Federal District: Paray 549 (Ip).
Hidalgo: Pringle 15608 (Ca—1682h0, Gg-—)21297, Mi, ce 4

agen LUCIDUM Schlecht. & Cham.

Additional iggy aes lucidum Schlecht. ex Molden-
ke, Résumé Suppl. = _ 1

Additional bibliog so P.C . Standl., Contrib. U. S. Nat.
Herb. 23: 1238 & 12h0. | Woldenke, Pavtologia 6: LSh--456-
1959; Moldenke, oes a hoe 6. 1966.

Rzedows untered ot A pe growing in Liquidambar for-
ests, teestdig 4 as May. The Paray 549, distributed as C. lucidum,
is actually C. ligustrimm Van Houtte

Additional citations: MEXICO: San cia Potos{: J. Rzedowski
10502 (Ip). Vera Cruz: Liebmann 11365 (Bi), 15486 (Bi).

UM LYCIOIDES D. ag:

Additional bibliography: Contrib. U. S. Nat.
Herb, 23: 121. 192h; penis itera 6: pte 19296
The Lundells describe this species as a slender shrub, 5 feet

tall, with o grange ig [they are Sinece drupes),
one hills, fruiting in August. Material has
been etary bien: and pe eb in herbaria as C. tetramerun
- S. Brandeg. See further notes of possible apeieation to
the present species under C. rosei var. e Moldenke.
Additional citations: MEXICO: Hidalgo: taal & Lundell
12332 (Mi, N, Rf).

CITHAREXYLUM MACRADENIUM
itio:

Addi
a me 13: 285. 1966; Uodéente, ,_Résuné Suppl. 13: 2. 1966.
t+ growing a + 6800 feet al-
tatnde. “flowering in July; Stork Soond it along *Badaiaen at
the edge of potreros
Additional eitahisoes COSTA RICA: Cartago: Stork 3283 (JM, ¥).
San José: Chrysler & Roever 5127 (Mi).

CITHAREXYLUM MACROPHYLL
Additional bibliograpty: nteidenke, Paytologis 6: 462--h6h.

1959; Moldenke, Résumé Suppl. 12: 3. 1
i describes this apetias ta as "Arbol, 2h a altura bape :
cm, Haz verde obscuro e. 3 =

oliares: 2, ligeramente rosadas.
res: blancas, fragrantes. Corola blanca; filamentos blancos-

1966 Moldenke, Notes on Citharexylum 297

—— marrén-cremosas. CAliz verdoso. Estilo verdoso. Estig-
a amarillento*, wien ni§ says "tree 20 m. tall; leaves sub-
sp

0 eums.
M en widely re eg at a distributed in her-
baria under the names C, poeppigii Wal C. poeppiggii Walp.,
igui Walp. In fact, the Aristaguiete Bh UA

by me in Phytologia 6: $00 2959) as C. s C. poeppigii, ar pall C.

word "Vaupés" and he phrase "galhos novos 8cos" on its
The Mell & Mell 1 2u7, cited by me in Phytologia 6: 163 (2959) as
c. macrophyllum, ap appears to be C. spinosum L

Additional citations: COLOMBIA: Valle del Cauca: Lehmann BT.
= a {4}. gt One Fact Aristeguieta 2787 (N). Bolfvar:

ta Amacuro: L. M. Berti 166 (N, Z); J» A.
Sager Fee. wer BRAZIL: Amazonas: G. A. Black )8-2491 (N,
Murca Pires 528 (N, N,N). Pard: G. A. Black 50-9001 (N);

ta Pe @ 3175 (N), 3209 (N, Z), S187 (N); N. 7. Silva 57802

CITHAREXYLUM oe Moldenke

Bibliography: Moldenke, Phytologia 9: 98-99. 1963; Koldenke
Biol. Abstr, 1,3: a3. 19635 Ho cking, Excerpt. Bot. A7: Sh. 196k.
Matuda found t & svoing in advanced forests, at 2786
neters altitude, fakin n May.
Pah an MEXICO: a oo Matuda 5467 (Ld—-type, Z~-isotype),

a a MEXICAN Molde 6: hob—L65.
onal bibliography: Moldenke, Phytologia =
19593 Langman, Select. Guide Lit, Flow. Pl. Mex. S15. 196h.

UM MIRIFOLIUM Moldenke
Additional bibliography: Moldenke, Phytologia i & 1959.
Cuatrecasas & Castaneda describe this species a tree, 8 m.
tall, with erect branches ; the leaves coriaceous, yea sh-
green, very g shiny above, the fruit blackish-brown, growing at al-

298 Putt 021.06 Ta Vol. 13, no. 4

titudes of 2700--2800 er carh fruiting in November
Additional citations: COLOMBIA: Magdalena: Cuatrecasas & Cas-

taneda 21,683 (n—23251,08), 25221 (2).

TTHAREXYL INNI D, Don
Additional bibliography: P. C. ontrib. U. S. Nat.
= fo 1237 & 1239. 192k sidan: ae Bitictouis 7 6. "1959.
jonal citations: MEXICO: Chiapas: Matuda 188), (Du—
Pie 4537 (Rf). uéxico: Hinton 5393 (Rf), S7ly (Ca—939137,
, Ur). Vera Cruz: Liebmann 15489 (Bi).

ITHAREXYLUM MOCINNI var. LONGIBRACTEOLATUM
Additional Macrae: Moldenke, Saye 6: Br TUT.

1959.
One branch of Matuda 5014, in the Lundell Herbarium, has in-

conspicuous bractlets, while the other branch has the typical
elongated bractlets. *vatuda 408, in the same herbarium, is in-
scribed "type collection" in error -- the type is Matuda s.181.

Additional citations: MEXICO: Chiapas: Matuda 08 08 (Ld, Mi),
2663 (Ca—91.3669, Du-—369921), 3923 (RE), Soh (1a)

CITHAREXYIUM MONTANUM var, CHIMBORAZENSE Moldenke
Additional bibliography: Moldenke, Biol. Abstr. 27: 2026.
1953; Moldenke, Phytologia 6: 74. 1959.

CITHAREXYLUM MONTEVIDENSE (Spreng.) Moldenke
Additional synonymy: Citharexylum montevidensis (Spreng.) Mol-
denke, Phytologia 7: 30, sphalm. 1959. Citharexylum barbinerva
Cham. ex Moldenke, Résumé Suppl. 3: 30, in syn. 1962. Citharexy-
lum montevidense Spreng. ex Moldenke, Résumé Suppl. 9: 3, in Sy?»

A ddi tional a Arechavaleta, Anal. Mus. Nac. Montev
: 149-~152, pl. 1. 19025 Herter, Estud. Bot. Reg. Urug. Bb: 20.

19334 Angely, aran. 7: 11, 1957; Moldenke, Phytologia 7: 76-
1959; Angely, Fl. Paran. 16: 5. 05 A. tz, Inst. Tec.
Rio Grande do Sul Publ. 30: 15. 1960; Rambo, Pesqis. Bot. ue

(1960) = 12: 21. 1961; Troncoso, Bol. S00. Argent. Bot. 9:
hagely, 7 ~ foment oe top he Arbustil. Pas. Publ. 313- ee
- Paran. 17: « 1961 1. Veg. Paran

; Rambo Peaguts. Bot. 21: Herd hee 1965; ous aia
Suppl. 13: 6. 1
Recent feet describe this plant as a shrub, 3 m- tall,
a small tree, —-5 m. tall, odo. ghee f the buds carrot-red
(612, Roy. Hort. Soc. Colour Chart 1938-42], the flowers fragrant,
the corolla yellow [Pedersen 5161) or Chinese-yellow (606/2, Rove
Hort. Soc. Colour Chart 1938-2; Bracelin 1327], and the fruit

streams, on banks of arroyos, and in valleys with outcrops © k-
coarse calcareous sandstone. Pedersen calls it "common in in thic

1966 Moldenke, Notes on Citharexylum 299

ets on moist ground" in Paraguay. Troncoso (1937) says "Lo he
visto cultivado en algunas calles de Martinez (Bs. Aires) 3 ademds,
segin el ingeniero L. R. Parodi existe en el parque 3 de Febrero,
una avenida arbolada con esta especie." She cites her no. 189

AL

Rambo (1965) cites Herb. Anchieta 8313, 9368 [Mmrich s.n.],
9348, 26234, 29666 (Henz sono), 34379 [Emrich s.n.], 44323, 58790
[Camargo 228], 60702 [Machado s.n.], and 63935 [Sacco 62] fran

Gerais and Paraguay through South
Brazil to Uruguay, and through Argentine Mesopotamia to B. Aires!!
7 a Schauer's work in Mart., Fl. Bras. as "1951" instead of

Additional citations: BRAZIL: Santa Catarina: Sellow 1879 [Mac-
bride photos 17591, in part] (W--photo), s.n. [Brasilia; Macbride
photos 17591, in part] (W—photo), PARAGUAY: Pedersen 5161 (Ss,
W--2,32653). URUGUAY: Arechavaleta 2 (Mi); H. H. Bartlett

» Mi). : Buenos Aires: Bri-
gerolli 4032 (W—-1987946). Corrientes: Pedersen 5379 (S, W—
2432715). CULTIVATED: California: Bracelin 1327 (Gg--339915,
Go); Chisholm s.n, [3 June 1952] (Gg—h12080) «

CITHAREXYLUM MYRIANTHUM Cham.
Additional synonymy : harexylon myrianthum Cham. ex Moldenke,
Résumé Suppl. 3: 31, in syns 1962.
Additional bibliography: Morong, Britton, & Vail, Ann. N. Y.
Acad. Sci, 7: 198, 1893; Reitz, Anais Bot. 2: 28 & 37. 1950; Ange-
P :h a ll: 7. 1959;

196 > Paran and 17: 46.
a pad Rambo, Pesquis, Bot. 12: 21 (1961) and 1h: 33. 1961; Veloso
lose 2 Sellowia 15: 102. 1963; Moldenke, Résumé Suppl. 12: 3.

55 Rambo, Pesquis., Bot. 21: 1j—15 & [59]. 2965.
2 Recent collectors describe this plant as a tree, 5--10 m. tall,
* & very tall tree, with white flowers » blooming from October to

300 PHYTOLOGIA Vol. 13, no. 4

ember, growing in bn a a at altitudes of 95 to 500 meters.
proves tes ecorded for it are "café do mato"
"pau de tuc

», and *

A cotype, mae 650, uascived in the herbarium of the Botan-
isches Museum at Be rlin, was photographed there by Macbride as
his type photograph number 17597, but is now destroyed. Similarly,
the type of C. macranthum, Fiebrig 596 596, in the same herbarium,
photographed b by him as his type photograph number 17596, is also
now destroyed. The United States National Herbarium specimen of
Rojas 11536 looks almost exactly like material now passing as C.
rigidum (Briq.) Moldenke. These two een pa more intensive
— . to determine if they are really di

o (1965) cites hos inckivia 1079, 2153, 11264 [Emrich s.
n.], fsa 266 (Emrich s.n.], 26659, 26946, 30606, 32681 681 [Henz s.n-],
32737 [Henz s.n.], 33757 [Bmrich s.n.], 45136, 59803 [Camargo

Grande do Sul, 50926 [Hatschbach 1653] from Parand, and 148431
Waite. 460], 1 [Reitz 61], and 55296 [Reitz 479] from
by hi

the northeastern quadrant except for the Highlands proper" in
Rio Grande do Sul. He gives its general eistribution as "Bahia
to RGS [Rio Grande do Sul] ; Paraguay to Misiones".

Additional citations: BRAZIL: Bahia: Sellow 650 [Macbride
photos 17597] Ce chaks: of cotype). Guanabara: ake Castellanos
23440 [Herb. Cent. Pesq. Florest. 1700] (Ac); Lanna Sobrinho 527
[Herb. Cent. Pesq. Florest. 2202] (Ac); B. Lutz a 1201 (Ja—1iA0,
Ja, Ja, Ja, Ja, Ja), 1410 (Ja—-11,099, Ja); Palacio ’
Cuezzo 4017 (Je— sill, Ja). Rio de Janeiro: rome 7817 (Ja

(N), 2325 (Gg); Reitz 2237 (W—21046h7), 8237 (Ca—li7509); Reits
& Klein 7607 (Mm), 80 8043 (Mm), 9280 (Gg). ~ PARAGUAY: ties 398
[Macbride photos 17596] (W--photo); 3 Hassler 557 (G)5 Ped

3263 (W—2169410), 42h3 (W--2283735); Rojas 1536 (W—- 21981 ui).

CITHAREXYLUM OLEINUM (Benth.) Moldenke
Additional rea Citharexylum oleinum Moldenke, Résumé

ins

dditional bibli : P. C. Standl., Contri 3
Herb. 23: 1252, ae croiy: Po Phytologia 6: iBT. "19595
L » Select. Gui de Lit. Flow . Pl. Mex. 515 & 1010, 196h%
Moldenke, Résumé Suppl. 13: 6.

McVaugh describes this Ae as a shrub, 1-1. 3 m, tall, with
white scentless flowers, flowering in April, and ab sae
north slopes of barrancas, at an altitude of 2700 ateres acai

1966 Moldenke, Notes on Citharexylum 301

owski found it growing at 1850—-1900 meters altitude, in chapar-
His om 7287, distributed as this species, is probably not
verbenace
sAdtttonal citations: MEXICO: Hidalgo: Paray 3200 (Ip). Quer-
taro: R. McVaugh 10360 (Mi). San Luis Potosf: J. R Rzedowski 6030
(Ip), 6655 (ip).

ee OVATIFOLIUM Greenm
tional bibliography: P.C ° ios » Contrib. U. S. Nat.
Herb. 23: 1238 & 12,0. 192h; H. B. D ‘Lite & M oes Pringle
153. 1936; Moldenke, Phytologia 6: 1 ne
The Paray 226), distributed as c. eeatAroLita “is actually C.
hidalgense Moldenke .

CITHAREXYLUM PACHYPHYLLUM Moldenke
Additional bibliography: Moldenke, Phytologia 6: l91--\92.

Additional citations: PERU: Lima: Asplund 11196 (W—222)),30) .

CITHAREXYLUM PENTANDRUM Moab
Additional bibliography: M Anatom. Untersuch. Verb. 72.

1911; Moldenke, Phytologia 6: ee. 1959.

xCITHAREXYLUM PERKINSI Moldenke
Additional TRENT? Citharexylum perkinsii Moldenke, Résumé
~~ 10: 5, °
tional tht iwameitann Moldenke, ES eruee oe 6: 96—h97.
1959; Moldenke, Résumé Suppl. 10: Sy i
describes this plant as a shrub, "2 me tall, growing at
1300 meters altitude.
Additional citations: PUERTO RICO: Alain 9465 (N, N, N).

ear pene Be UM POEPPIGII Walp. gis aagis
onal synonymy: Citharexy um poeppigui Walp. ex Mo

Résumé Suppl, 12: 9, in syn. gt ere lum poeppiggii Walp.
ex Moldenke, Résumé Suppl. 13: 6,

a P-green a 5 er~green

ent, very showy, the rachis pale buff-green, the calyx pale-

green, and the fruit round or ro und-oblong, at first green, then

5 range, fi vivid-red or scarlet and shiny, very showy,

6¢ to Bane and 12 m., wide, growing in forests : friting pot
meters, f1 ruary. and ng

March and er. mre rps ae marrén

ri cure ays "Tro: ;

saint ? § madera marrén rosada; hojas i Haas flores cu-

6 6 = lertas amarillas 3 hojas verticiladas [but they are decus—
Pposite on all the specimens cited below], glandulas escu-

302 Pete OO Tf Vol. 13, no.

teliformes abe los pecfolos."

Additional vernacular names Reve for the eg are
"cerezillo", tuatahnayo o", "quineillia blanco", totumillo
negro". Mater ial has been saat dant’ tied and dines ee in her-
baria as C. decorum Moldenke and Guettarda sp., while Cardona
1421 was first identified as something in in the Bignoniaceae, ceae, then
in the Combretaceae! The Cuatrecasas 1599 & 17606, cited by me
in Phytologia 6: 500 (1959) as C. poeppigii, prove on sheet ex-
amination to be C. po eppigii var. calvescens Moldenke ,

Aristeguieta 2787, Black 48-2491 & 50-9081, and Murga "Pires “528,
3175, & 3209 are C. macrophyllum Poir. The L. M. Berti 166 and
Steyermark rk 87,72, “distributed as C. poepp igii, are actually Cc.
macrophyllum, Ownbey 2715 is C. poeppigii var. margaritaceum
Poepp. & Moldenke, and R. E. Schultes 3381 is C. ulei Moldenke.

Additional citations: VENEZUELA: Amazonas: Cardona 121 (Ve).
Bolivar: Bernardi 6739 (N), 7190 itn ad “Delta Amacuro:
Steyermark 87472 (N). PERU: Loreto: 365 (W--1459252).
artin: Woytkowski 7095 aa Ti 6 (W-~2)53515) «

ITHAREXYLUM POEPPIGII f. ANOMALUM Moldenke
Biblio cera Moldenke, Phytologia 8: 59. 1963; Moldenke,
Biol. Abstr. 1,3: 1278. 1963; Hocking, Excerpt. Bot. A7: liSh.
Little describes this plant as a tree, 8 m. tall, with
trunk diameter of 15 cm. at breast height, growing in the wood
in a llano, at 100 meters altitude,
Citations: VENEZUELA: Barinas: E. L. Little 15087 (Ve).
Bolfvar: Bernardi 2142 (Ve—type) .

ay ore LUM POEPPIGII var. CALVESCENS Mol
ography: Moldenke, Phytologia 8: 189. » 19635 Moldenke,
Biol. Abstr. 3: 1278. 1963; Hocking, Excerpt. Bot. A7: li5h.
trecasas describes this plant as a tree or "arbolito; tal-
lo 8 cm. did4m., blanquecino con estribos basilares; corteza

b 9° ocrécea; ranosa verde claro; c verde pdlido;
corola blanco verdosa, tubo de la a blanco verdos °
blanco" and found it growing in woods, oe Pei

crt POEPPIGII var. MARGARITACEUM Poepp. . Moldenke
tional bibliography: Moldenke, Phytologia 6: 500-501.
19595 Moldenke, Résumé Suppl. 12: 3.1965.
escribes this as a small tree, with white flowers,
evtinns in potreros, and A yttctivern in December.

1966 Moldenke, Notes on Citharexylun 303

Additional citations: ECUADOR: Napo-Pastaza: Harling 3650 (S);
Ownbey 2715 [Herb. U. S. Nat. Arb. 217809] (W). PERU: Loreto:
Kug 3943 (W--1458369).

CITHAREXYLUM PTEROCLADUM Donn, Sm
Additional Baird barack: Yoldenke, Phytologia 7: 77 (1959) and
13: 292, 1966.
The flowers on Hinton 13738 are described as "blue", while

those on Skutch 2021 were "pale lavender with dee eper pure streaks
on lower paeen Steyermark 5851, distributed as C. pterocladum,
is actually C are Greenn,.

Additional Tieton eopron, Chiapas: Matuda 188) (Ca—
903787). Guerrero: Hinton 1,082 (Mi, Rf). Michoacdn: Hinton
12631 (Mi, Rf), 13738 (Rf, 3 Ur, W--18),3285), 13739 (Rf, Rf, Ur, W--
rte 286), 15852 er GUATEMALA: Quezaltenango: Skutch 2021 (W—

388, "W—=18L4389 oe

ITHAREXYLUM PUNCTATUM Gres.

Additional synonymy: Aegi phila punctatum Greenm. ex Moldenke,
Résumé Suppl. 12: ah Gr sig rs
Additional bibliography: » Biota 1: 180. 1956; ‘eer

Phytologia 6: S0li——Soe : 19595 eae Biol. Abstr. 33: 1
1959; Moldenke, Résume Suppl. 12: 9. 1965; Moldenke, Phytologia
13: 280. 1966.

aterial has been misidentified and distributed in herbaria
as C. argutedentatum Moldenke and as Terminalia

Additional citations: BOLIVIA: Cochabamba: M. Gardena 46h,
(W—2250912 ); Collector undesignated 5 (W--2320699 MF bers 70
(Ko). La Paz: H. H. Rusby 2058 (W--32353). Department undeter-

_—_—_ —_—

mined; Solleskor undesignated 186 [Liriuni] comp) .

ITHAREXYLUM QUERCIFOLIUM H Hayek

Additional bibliography: pied Piytologia 7+ 7-0. 19595
Moldenke, Biol. Abstr. 3h: 481. eit 4. a ae agjgndesvecanasiveeyds
43. 1962; Moldenke, Résumé Suppl. 12: 3.

The type s imen, Weberbauer “L2h8, deposited pt waptheaase tei er

an ional citations: PERU: Cajamarca: Weberbauer 248 [Mac-

Ca —,
ie ghies ay aa La Libertad: Ellenber
rH 7 naib x say (W--photo of type). g

CITHAREXYLUM QUITENSE Spreng. aes
Additional sahitarverers h 170.
19085 Mol ldenke, Phytologia a a mages) eo “ak 1966.
The type specimen of C. molle, Bonpland 3817, deposited in the

304 PHYTOLOGIA Vol. 13, no. h

Bonpland Herbarium at the Muséum National d' Histoire Naturelle at
Paris, was photographed there by Macbride as his type photograph
number 39481. The Ellenberg 1,89, stributed as C. quitense, is

dditional citations: ECUADOR: Guayas: Asplund 5170 (Gg--
ity W—-2223701); Bonpland 3817 3817 [Macbride photos 39481] (w—
photo).

CITHAREXYLUM RACEMOSUM Sessé & Moc
Additional bibliography: Woldenke, Phytologia 7: 10—1l. mk
4 Illustrations: Sessé & Moc., Fl. Nouv. Hisp., ed. 2, pl.
1893.
Dr. Rogers McVaugh, in a letter to me dated October ll, 1963,
sent me a photograph of the original Sessé & Mocino plate “dophet-
ing this plant, with the comment: "Plate 418 of the Plan

thing else." Recent sian pga a the plant o$ a shrub,
2.5 m. tall, with orange-red fruit, growing in gray or brom
gravelly soils in tie ie atts association, at 1000 meters
altitude, and fruiting in July.
ditional citations: MEXICO: Puebla: Smith, Peterson, & Teje-
da 4121 (W--2398123, Z).

CITHAREXYLUM pede eee Moldenke
dditional bibliography: Angely, Fl. Paran. 7: 6. 1957; Mol-
denke, Phytologia 7: "12-13. 1959; Rambo, Pesquis. Bot. al: 15 &
[59]. 1965.
Reitz & Klein describe this species as a . tall,
immature fruit green, growing at an aittade: of e380 neters, howe
ering and fruiting in December
Rambo (1965) cites Herb. Anchieta 3052 [Reits get. sre
ta Catarina, Brazil, He describes the species as a "Small tree”
growing in "Half-high woodlet" in "Southernmost SC “(santa oat

easternmost c
gives its general distribution as "SC".
8022 (2) citations: BRAZIL: Santa Catarina: Reits & Klein

CITHAREXYLUM RETICULATUM H.B.K.
Additional bibliography: Moldenke, Piytologia 7: as 1959+
specimen, Bonpland 3119, deposited in th Bonp.
pa iad at the Muséum National d'Histoire ai at Paris,
we there by Macbride as his type photograph number
Additional citations: ECUADOR: Loja: Bonpland 319 [Macbride
photos 39179] (W——photo of type).

oe
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PHY TOLOGIA

Designed to expedite botanical publication

gt

Vol. 13 August, 1966 No. 5

CONTENTS

e _ HAWKES, A. D., Grafia, a new genus of orchids 2. 2 oS ee 905

MOLDENKE, H.N., Notes on new and noteworthy plants. XLV . . . 306

BNC eS A D...A problem in Deudrobamc

on the genus Aegiphila.

on the genus Lippia.I1. . . . 343

+303 Parkside ead”
Plainfield, New Etsy U oe

GRAFIA, A NEW GENUS OF ORCHIDS
Alex D. Hawkes

The genus Phalaenopsis of Blume has long been a popular

th orchid collectors, and has in the past received “Sebidee able
nie attention from students, serious and casual, of the fa-
mily. But it is only in recent years that we have seriously con-
sidered the components of this group, which dates from the year
1825,

As is well known, in the early days of botany, a truly diverse
array of orchids was named, by different authors, as Phalae-
nopsis— a pow ee which also prevailed in Cymbidium, Den-
drobium , Epiden etc. Now, segregates from Phalaenop-

sis Bl. have — established (or re-evaluated) and accepted,

these Doritis Ldl. Kingie ella Rolfe, and Paraphalaenopsis A, D.
Hawke

es.
Considerable additional critical study of the orchids we still
consider as Phalaenopsis remains to be done. The genus, as it
is presently delimited, is still a motley poisecmabinnh of plants,
several of which should be ahaccaas as distinct t
Reichenbach's plant named Phalaenopsis Bicianiit in 1865 is
One of these, one which now requires our serious consideration.
This had been collected in Moulmein, cae by the Rev
Parish, and introduced into England by Messrs. Low and Co.the
Previous year. It has, for more than a century, been in cultiva-
tion, yet though its aberrancy in its genus has been commented
Pon in the literature, no orchidologist has resolved the problem.
Iam convinced that this orchid constitutes a genus of a single
Somewhat variable and rather widespread species, amply distinct
from Phalaenopsis of Blume. I therefore propose the following
new genus to accommodate at

GRAFIA A.D. Hawkes, gen. nov
Differt ab Phalaenopsis Pines in labellum libere mobilum,

crista semi-lunata valde LSoeeleee fa antice appendix angusta,
4-divisa angustissima or

A genus of a sin ingle species, ranging from the Himalayas to
Burma. It differs from Phalaenopsis Bl. in its freely mobile
labe which bears a semi-lunate crest, this strongly fim-
briate; this is followed, anteriorly down the segment, by a for-
ward-thrust, narrow appendage, this divided almost to its base
into a ceara of slender filaments.

306 PHYTOLOGIA Vol. 13, no. 5

I would temporarily place the new genus in somewhat closer
proximity to Ornithochilus Wall. than to Phalaenopsis Bl. itself,
but believe that additional study of the entire alliance may alter
this opinion. Since Grafia is present in choice contemporary
collections, further fresh materials of the flowers will be wel-
come from my correspondents for study

I wish to propose the new genus to baer Alfred Byrd Graf,
editor of the remarkable and internationally valued volumes en-
titled Exotica. As seems customary with such serious work-
ers, Mr. Graf's efforts have been largely unacknowledged, and
I am sure that all of us who value his labors will consider the
epithet of the new genus singularly appropriate.

GRAFIA PARISHI (Rchb.f.) A.D.Hawkes, comb. nov.
Phalaenopsis Parishii Rchb.f. in Bot. Zeit. 146. 1865. Pha-
laenopsis Parishii Rchb.f. var. Lobbii Rchb.f. in Saund. ,
Refug. Bot. 85.
Distribution: Sikkim , Bhutan, and Assam to Moulmein and
Sperergicen districts in Burma.

e plant is frequently deciduous in the wild, rarely
so under a eT The Himalayan phase, described as a dis-
tinct variety by Reichenbach, does not differ save in coloration,
hence is here considered a synonym of the typical species.

Grafia Parishii is taken up in greater detail in The Orchid
Weekly, V Vol. 5, 1966, to which the interested reader is refer-
red. It is described as Phalae nopsis Parishii Rch hb; £,:2n ey
Encyclopaedia of Cultivated Orchids 368. 1965, and is illustra-
cS as Phalaenopsis Parishii Lobbii in Graf's Exotica 3: 1305.
This is a very attractive, profusely-flowered orchid when well
grown, and in future our avid breeders are certain to take it under
Serious consideration as a parent; we already have casual indica-
tion (see Orch, Weekly 4: 462. 1964) that some efforts have been
made in this respect.

NOTES ON NEW AND NOTEWORTHY PLANTS. XLV
Harold N,. Moldenke
CLERODENDRUM GLABRUM var. VAGUM (Hiern) regonanrey vias Alph.

List Invalid Names 0, hyponym. 1940; comb.
Siphonantims ra var, vaga Hiern, rity Afr. "Pl. Welw. 1: 8h2-

snip amma ttt i a a a aa 8 I ng

1966 Moldenke, New & noteworthy plants 307

STACHYTARPHETA SCABERRIMA var. PILOSA Moldenke,

Haec varietas a forma typica speciei ramis Lr iateecinti tints
dense piloso-pubescentibus recedit.

This variety differs from the typical form of the species in
having its stems, branches Mera rachids, bractlets, and

blue flowe

—— LONGIFOLIA var. PUBESCENS Moldenke, var. nov.
Haec varietas a forma typica speciei ubique perspicue breviter
pubescens rec

This variety dif fers from the typical form of the species in
being cheng grress ee ae throughout.
The type of the variety collected by D. E. Breedlove (no.

12292) on a steep slope with © Guise and Pinus, at an ela of
7000 feet, 19 km. — of Sola de Vega, Se

Puerto Esco do, Oaxaca, Mexico, on August 1965, and is de-
posited in the H. N. Moldenke herbarium at Hatitiast: alg asain,
The flowers are described as blue

A PROBLEM IN DENDROBIUM
Alex D. Hawkes

In an article ol scsi not long ago (in Orch. Rev. 73: 87 - 88.
1965.), R. E. Holttum has pointed out that the name of the com-
monly-cultivated orchid Dendrobium a ageregatum Roxb, is a ho-
monymic one, this in reference to a paper of mine in which cer-
tain such cases were taken up (l.c., 21 - 22. 1965.). Dr. Holt-
tum, unfo ortunately, did not take time to discuss the correct epi-
thet of this Dendrobium, and in none of his published works, nor
his book, has he done so. This seems regrettable, since he is so
vehement about my failure to attend to this matter.

It is indeed puzzling that none of us who have worked with the
huge and com plex Dendrobium alliance has attended to this rath-
€r obvious situation, one which I herewith rectify.

Three totally distinct orchids have been described as Dendro-
bium aggregatum. In this case, the species in common and wide-
Spread cultivation under this name dates from 1832, dean the
epithet used by Roxburgh in 1814 was validated. In the interim,

308 PHYTOLOGIA Vol. 13, no. 5

however, in 1816, Humboldt, Bonpland, and von Kunth, following
their explorations in the Americas, published an orchid which
they named Dendrobium aggregatum. Since no true Dendrobiums
occur in the Western Hemisphere, this — been shown to be ref-
ee to Maxillaria aggregata (HBK) Ld
e third orchid named Dendrobium eee gatum dates from

Pie by Nathanael Wallich; this is now referred to D. Griffith-
ianum

In my recent Encyclopaedia of Cultivated Orchids (Faber, Lon-
don, 1965), I called this plant Dendrobium aggregatum Roxb., and
also considered a variety amply distinctive. These erroneous de-
cisions on my part, following Holttum and all previous writers on
the subject, are hereby corrected.

The next available name for the plant so long known as Dendro-

bium aggregatum Roxb. is
DENDROBIUM te yenea Steud. , Nom., ed.2, 1: 490. 1840.

Dendrobium a atum Roxb., ore Beng. 63. 1814; and Fl.
Ind. 3: 477. tea not HBK, . Gen. et Sp. 1: 358. 1816;

r. Wall, , Catal. no. 7411B. si 28.
pas AR poe South China to Burma, Thailand, and Laos.

- var. JENKINSID (Wall.) A.D. Hawkes, comb. no
Dendrobium Jenkinsii Wall. in Ldl., Bot. Reg., = 37. 1839.
D robium a tum Roxb. var. Jenkinsii Ldl.
Distribution: ge repre Burma.

Amore lengthy, pertinent discussion of the problem is being
published in The Orchid Weekly, Vol. 5, 1966, to which the inte-
rested student is referred.

NOTES ON BULBOPHYLLUM — 1
Alex D, Hawkes

g the preparation of a check-list of the orchid subtribe
Bulbopiyilina, for publication in my continuing 'Index of Orchi
daceous Plants," a number of taxonomic alterations have PF
necessary,

The first of this series of notes considers a quartet of species
of Bulbophyllum Thou. whose names require clarification.

oe

1966 Hawkes, Notes on Bulbophyllum 309

BULBOPHYLLUM FLICKINGERIANUM A. D. Hawkes, nom.nov.

Bulbophyllum fimbriatum H.Perr. in Not. Syst., ed. Humbert,

9: 145. 1941, not Rchb.f. in Walp., Ann. 6: 260. 1861;nor
H. Perr. in Not. Syst. 14: 1953. 1951.

Reichenbach's Indian Bulbophyllum fimbriatum, based on Lind-
ley's Cirrhopetalum fimbriatum, predates both of Perrier de la
Bathie's Malagasy plants by many years. I wish to propose the
above new name to accommodate this species described from Ma-
dagascar, to recognize the considerable orchidological accomplish-
ments of my colleague, Edward A, Flickinger.

BULBOPHYLLUM MAUDEAE A. D, Hawkes, nom.nov.
Bulbophyllum nigrilabium H.Perr. in Not. Syst. 14: 150.1951,
not Schltr. in Fedde, Repert. Spec. Nov., Beih. 1: 800. 1913.
Schlechter's Bulbophyllum nigrilabium, from New Guinea, pre-
dates that of Perrier de la Bathie by many years. The new name
is proposed to recognize the indefatigable assistance of my general-
ly unsung collaborator, Miss Maude Drum.

BULBOPHYLLUM MAYAE A.D.Hawkes, nom.nov.
Bulbophyllum fimbriatum H. Perr. in Not. Syst. 14: 1953. 1951;
not Rchb.f. 1861; nor H. Perr. 1941.

As noted under Bulbophyllum Flickingerianum A.D. Hawkes, two
of Perrier de la Bathie's Malagasy species named B. fimbriatum
are homonymic because of the long-prior Bulbophyllum fimbriatum
of Reichenbach filius. This new name is given to honor Mrs. May
Moir, who has long afforded unique assistance to orchidists and or-
chidologists everywhere, and who delighted in the flora of the island
of Madagascar.

BULBOPHYLLUM MOLDENKEANUM A, D. Hawkes, nom. nov.
Bulbophyllum microglossum H,Perr. in Not. Syst. 14: -151.
1951, not Ridl. in Jour. Linn. Soc. 38: 325. 1908.
Perrier de la Bathie's Bulbophyllum microglossum is an unten-
able name, being predated by that of Ridley for a completely differ -
ent Malayan plant. I wish to propose the new epithet to honor, ina
‘mall degree, the efforts of Dr. Harold N. Moldenke, co-editor and
Publisher of Phytologia, a periodical which has done so much to fa-
Cilitate contemporary botanical publication.

ADDITIONAL NOTES ON THE GENUS CITHAREXYLUM. III

Harold N. Moldenke

UM B, Juss,
Additional bibliography: Moldenke, Phytologia 13: 2h2 & 277—
30h. 1966; G. Taylor, Ind. Kew. Suppl. 13: 31. 1966.

CITHAREXYLUM BRACHYANTHUM (A. Gray) A. Gray
coping synonymy: Citharexylum b rachyanthum (A. Gray 6x
Hemsl.) A. Gray apud G. Taylor, Ind. Kew. Suppl. 13: 31. 1966.
Saaeisianek Hy, Fis ppl Pe Moldenke, Phytologia 13: 281—262.
1966; G. Taylor, Ind. Kew. Suppl. 13: 1.

ITHAREXYLUM FLABELLIFOLIUM S, Wat
Additional bibliography: duke. payvatoeia 13: 286. 1966;
G, Taylor, Ind. Kew. Suppl. 13: 31. 1966

CITHAREXYLUM HIRTELLUM Standl,
EE shia oe bibliography: Moldenke, Phytologia 6: 09-~I11.
Breedlove & Raven refer to the fruit of this plant as “ber-
ries", but they are goccumen eS — collectors eg!
the plant growing on steep heavily wooded slopes, fruiting in
October.
Additional citations: MEXICO: Chiapas: Breedlove & Raven

23625 (2).

xCITHAREXYLIM JAMAICENSE Moldenke
Additional bibliography: G. Taylor, Ind. Kew. Suppl. 13: 3l-
1966; Moldenke, Phytologia 13: 293. 1966.

Additional iitasaareses ue Moldenke, Phytologia 6: h72—h7h
(1959) and 13: 298. 1966.
—— ; Cxtharexyiun perkinsii Moldenke, Résuné
Suppl. 10: 5, in syn; 196h3 G. Taylor, Ind. Kew. Suppl. 13+ 3l-

Additional bibliography: Moldenke, Phytologia 13: 301. 19663
G, Taylor, Ind. Kew, Suppl. 13: 31. 1066,

UM RIGIDUM (Briq.) Mold

CITHAREXYL
Additional bibliography: Bn png Phytologia 7: 15--17 (1959)

and 13: 300, 1966,

CITHAREXYL' votre.
Additional bibliography: P. C. Standl., Contrib. U. S- Nate
310

Fae Ee NO RENE MEE

1966 Moldenke, Notes on Citharexylum 311

he 23: 1237 & 1239. 192h; Moldenke, Phytologia 7: 18—~20. 1959;
denke, Biol. Abstr. 37: 1062. 1962; Hocking, Excerpt. Bot. A.5:
iS 1962; Moldenke, Phytologia 13: 296. 1966.
Recent collectors describe this plant as an "abundant treelike
pee heres 10 cm, in os antes ace a

stream ky oak-

covered Mantel netas. or with uyrtillocsctus and piliberny tia in mator-
ral, at 2000—-2100 matine altitude, flowering in September, and
fruit ting in November, H. S. Gentry 856) 856 was collected in the
ene se Sierra Papantén near highway on Durango-Zacatecas

, therefore, have actually come fram either of
onag sta tas.

Additional citations: MEXICO: Durango: H. S. Gentry 856)

805272, Mi). San Luis Potosf: J. Rzedowski i sheh (Ip). poet aT
R. MeVaugh 17670 17670 (Mi, Mi, Z).

ITHAREXYLUM ROSEI var. DURANGENSE Moldenke
Additional & emended synonymy: Rauwolfia oides Lag. & Rodr.,
Anal. Cienc, Nat. 5: 69—~70. 1802. Rauwolfia lycioides Cav. apud
Jacks. in Hook. f. & Jacks., Ind, Kew. 2: 693. 1895.
In a series of very interesting commnications from Dr. Rogers
MeVaugh, dated October 11, November 5, and December 2, 1963, I
c

original type specimen of Rauwolfia lycioides Lag. & Rodr. in th

and Rauwolfia oe were ante on different specimens from the
Same source......Don's specimen came from Pavén, and according to
Don it was thought by "Pas to 3% a new species of Rauwolfia.....
Rauwolfia lycioides 'Cav.' was actually published by Lag. & Rodr.
in a signed article; probably Cav. gets the credit, as in Index
Kew., because he was the professor."

Recent collectors describe this plant as a small, spreading
shrub, about 6 feet tall, with the fruit red when ripe, growing on
low limestone

Conglomerate, at altitudes of 1 rs, fruiting in July,

September, tober. = on == who refers to the fruits as

"berries" [they are act s) — ese the ges "scarce",
e try describes it as "anon

Additional citations: MEXICO : pibendol Correll & & Johnston 20
(RE); H. S. Gentry 6286 (Mi); M. C. Johnston 262k (Au--12320h).
CULTIVATED: Spain: Herb. Hort. Bot. Matrit. son. [Herb. Univ. Mich.
neg- 1700) (Z—-photo).

UM ROSEI var. PILOSUM Moldenke
Bibliography: Moldenke, Phytologia 8: 1h. 1961; Moldenke, Biol.
Abstr. 37: 1062, 1962; Hocking, Excerpt. Bot. A.5: 45. 1962.

312 PHYTOLOGIA Vol. 13, no. 5

Citations: MEXICO: Jalisco: R. McVaugh 17152 (Mi-—-type, Z--
isotype).

ROXANAE Molde
Bibliography: elses i ctiake T2: pa Ay 1961; Moldenke,
Biol. Abstr. 36! 1961; Hocking, Exp . Bot. AS: bh. 1962.
Carter des ni fey plant having Pn nee be stri-

ges 'ttarbert, Hyptis emoryi, and Aloysia barba
aibatoesr: MEXICO: Baja California: A. Carter 123 om
Gg); Carter & Ferris Ferris 4,000 (Ca~~199586—type, Z——isotype) «

CITHAREXYLUM SC
cr P. a Standl., Contrib. U. S.
Herb, 23: 23 r Tos 192h; yelsecks, betel Se aio. i oe “t9595
é 33 &
names found this plant growing on grassy limest e hill-
sides in Sinaloa

Additional peel pee MEXICO: Sinaloa: Edw. Palmer 1523 (Ca--
770322); Waterfall 16266 (Z).

CITHAREXYLUM SCHOTTII Gre
Additional bibliogr: ee Standl., Contrib. U.

Beat oats & 12hi. 1935, oldenke, peetipglt 7: 55226 “(959)
ch centie | 4793, distributed as C. schottii, is actually &- Cs
caudatun L. L.

~~“{aditional citations: MEXICO: Quintana Réo: Lundell & Lundell
7781 (Du--3619)6, Rf). Yucat4n: Lundell & Lundell ell 7078 a
36305, Rf), 8132 (Du-360781, Rf, Rf).

CITHAREXYLUM SCHULZII Urb. & Ekm. 69
Additional bibliography: Moldenke, Phytologia 7: Been t 1997+
Holdridge describes this plant as a alti-

leaves rigid, the fruit _ globose, ene at "1150 50 ters

tude, fruiting in October )
Additional citations: HISPANIOLA: Hafti: Holdridge if fo @ 1775 ()»

ITHAREXYLUM SESSAEI D. Don re
Additional bibliography: J = Untersuch. Verb. ee
73. 1911; H. B. Davis, Life & veanine —e 19363 = —
Phytologia 7: 27-28. 1959; en Résumé Suppl. 13:
Moldenke, Phytologia 13: 279. 1966. Boe
Hgnton — ths plant as 3 m. tall, with white fl ’
at 2200 eee altitude, flowering in Novem
ber; he notes that he found only " plants".

a TBE AN Peers a ONES

1966 Moldenke, Notes on Citharexylun 313
Additional citations: MEXICO: Michoac&én: Hinton 15678 (N, N,
CITHAREXYLUM SHREVEI Mold
Additional bibLiograpiy: Moldenke, Phytologia 7: 28—29. 19593
Moldenke in Shreve & Wiggins, Fl. Son. Des. 2: 1259. 196k.

CITHAREXYLUM SOLANACEUM Cham
“cca & ago: bibliograpty: Ranbo, Sellowia 7: 260 &

288. 1956; Angely, Fl. Paran. BN “ann ¥ oldenke, Pitan ogia
73 pe Mata ingely, FL ° 16: 45 (1960) and 17: h6.
19613 » Pesquis. Bot. 21: 1-26 & [59]. 1965; | sane
Rpisiogia. 13: ae 1966.

Smith & Klein Ec this species growing in forests and at
forest margins. Rambo (1965) cite Herb. hithdate a $229,
oe? ae ee mye: 54255 (Emrich s.ne], &

Rio Gr: do Sul, and 5092 Oe et 1591] from aa

omiages all collected by meal? except where otherwise stated.
He calls it a "Small to medium ae up to 10 m. high",

in "Araucaria forests, secondary growth....All collections from
the northeastern quadrant, especially the Highlands; only 6559

dditional citations: BRAZIL: Parand: Reitz & Klein 17750
(Ac). Rio Grande do Sul: Sehnem 5071 (B, B). Santa Catarina:
Smith & Klein 13219 (Ac). S&o Paulo: Puiggari 3330 3330 (P).

CITHAREXYLUM SOLANACEUM var. MACROCALYX Moldenke
Additional bibliography: “Woldenke, Phytologia 7: 31--32. 1959.
The type specimen, Sellow s.n., deposited in the herbarium
of the Botanisches Museum at Berlin, was photo t by
Macbride as photograph 17600, but is now destroy~

his
» The Brescia & Borsant 3361, cited below, bears a notation
"Corresponde a mestra No. [pee", but I have not as yet seen this
latter collection,
Additional citations: BRAZIL: Rio Grande do Sul: Brescia &
Borsani 3361 (Z). State undetermined: Sellow s-n. ("varietas";
lacbride, pho photos 17600; Herb. U. S. Nat. Arb. 197511] (W--photo of

CITHAREXYLUM SPATHULATUM Moldenke &
ner gees bibliography: poner ley Pivtaloeia 7, 73 gimecpates 1959.6
rell & Johnston describe this as a sh
on aes J ——

branched
ional citations: TEXAS: Hidalgo Co.: Lundell & & Lundell
9953 (Mi-type), 12689 (Re) , 12774 (Rf). Starr rr Co.: Correll &
Johnston 18059 (Rf); Lundell & Lundell 12676 (Rf).

CITHAREXYLUM SPINOSUM L.

314 PHYTOLOGIA Vol. 13, no. 5

Additional synonymy: Hadongia eberhardtii Gagnep. in Humbert,
Not. Syst. 1h: 30. 1950. Cyt harexylon spinosum L. ex Moldenke,
Résumé Suppl. 13: 6, in syn. 19

dditi emended bibliography: Griseb., Cat. Pl. Cub. 216-
1866; Hayek in Engl., Bot. Jahrb. 2: 169. 1908; Britton & P.
Wils., Scient. Surv. Porto Rico 6: 145 & 146. 1925; Robledo, Lecc.
Bot. 2: 498. 1940; Neal, In Gard. Hawaii, ed. 1, 639. 1948; Gag-
nep, in Humbert, Not. Syst. 1h: 30. 1950; E. J. Salisb., Ind.Kew.
Suppl. ll: 106. 19533; Moldenke, Verb. 22—~23. 1955; Moldenke,
1 jad e Gard. Luc

Phytologia 7: 77. 1959; Nair an, k-
now 76: 10 & 23, pl. 1, fig. os text-fig. 16. 1962; Little & Wads-
worth, U.S. Dept. Agr. Forest Serv. Agric. Handb. 29: poe 196h5

Moldenke, Résumé Suppl. 11: 5 (1964) and io: 9. 1965; Gooding,
Loveless, & Proctor, Fl. Barbados 356, 68, & 69. aks Poa In

Additional illustrations: Nair & Rehman, Bull, Nat. Bot. Gard.
Lucknow mt pl. 1, fig. 6, text-fig. 16. 1962; Anon., Short Guide
Bermudas 16 [in color]. ned.

The type of Hadongia eberhardtii was collected by Philippe
Albert Eberhardt (no. Anos: Sa0), probably from cultivated material,
at Hadong, Tonkin, ochina, and is deposited in the herbarium
of the Muséum fey d'Histoire Naturelle at Paris. Ww
originally proposed in the Bignoniaceae, as the type species of
a monotypic genus

Wilder dance ban C.s sum as a large spreading tree. Box
affirms that on Antigua it is "occasional to frequent locally in
woodlands, chiefly on xerophytic hillsides near the sea". W. Re
Taylor 49-1155 represents a vigorous shoot neve its leaves are

flowers are described as "white" on G. Le
Webster 1582, "cream" on Clemens & Clemens 3755, and "cream
colored" on “on 0. E. White 18. Whi White comments that girls in Burma
wear its flowers in their hair.
The Mell & Mell 247, cited below, was erroneously cited by me
‘Tium Poir. in Phytologia 6: 63 (3959) The A. Re
Cooke s.n. [Manoa trail, 9/26/sl,] and Degener & Murashige 20073,
igs as C. Sipown, are actually C, caudatum pa while
R. A. Howard 10733 and Purseglove P.6363 are C, fruticosum L.,
and Gi Cowan & Forster 1278 fat least insofar as the Britton Herbar~
ium is concerned] is C. fruticosum var. brittonii Mol-

denke.
Nair & Rehman (1962) describe the pollen-grains of C. spinos™

as follows: 3-zonicolporate, spheroidal (diameter 36 3 range 32~
39 ¥); ectocolpium faint, ends pointed, crassimarginate; endocol-
%

(faint 10), based on Herb. Nat. Bot. Gard. Lucknow mere, Palide
2621, not seen as yet by me.

eearienemmneneecuaatmmemeemeraieneneeetinitit

1966 Moldenke, Notes on Citharexylum 315

Little & Wadsworth (196) Beg "A related species of péndula
(Citharexylum s spinosum L.), © known as susana, is wild and
planted in St. Croix and St. poe tee and ranges southward to
northern South America. It has elliptic leaves 3—8 inches Aen,
ins when

ct
pe
5
E
rt)
a
Qa.
La)

Additional citations: BERMUDA ISLANDS: tn W. R. Taylor
49-1155 (Mi). Main: Bailey, Bailey, Whetsel, Degener, & McCallan
s.n, [Experiment Station, Sept. %. 10, 1921] (B: (Bi). Saint erie:

W. R. Taylor 49-1114 (ut) LEEWARD ISLANDS: Antigua:
(i). Guadeloupe: Questel 627 (W--245370h), 4663 (eas),
oho (W--21,53677), 5066 5066 (W—2453703). WINDWARD ISLANDS: Martin-
ique: Sieber Fl. Mart. 156 (P); ae & Stehlé 505 ey eg
6819 (W--2453705). TOBAGO: W. E. Broadway y 3621 (P). BRITISH
GUIANA: Irwin 510 (W—22128)6); Mell a 4. Mell 27 (N). FRENCH
GUIANA: Martin s.: sen. [Cainne] (P). INDIA: Mysore: Ramaswany 642
(Lw). INDOCHINA: Tonkin: Eberhardt 3ha7 7 (P). POLYNESIA: HAWAII-
AN ISLANDS: pats G. L. Webster 1582 (Mi ie W—-2132h72). CULTIVA-
TED: Burma: . White 18 (W--2073086) . Curagao: Arnoldo 2089
2110507) a Islands: G. P. Wilder 1234 (Bi, Ca-—948871).
Hawaiian Islands: C. M. Cooke sen. [May 10, , 1941] (Bi); A.F
gudd sen, (Hakipmu, July 3, 1939] (Bi, Bi); H. Ne Moldenke 85h
(QG); Teixiera s.n. (May 27, 1937] (Bi). Indochina: Clemens &
Clemens 3755 (W—1128066). Java: Herb. Hort. Bogor. X1.G.17a 1.G.17a_
(N). Senegal: Adam 16948 (Mm), 16949 (Mm).

I var. MEXICANUM M

CITHAREXYLUM STANDLEY
Additional bibliography: Moldenke, Peeainets 7: 7. 19
McV te ni describes this plant a s tree-lik e, hm. tall,
fl

erassy. slope of mountain summit in deciduous woodland dominated
a legumes, Bursera, and Cnidoscolus, at 500 meters al~-
ude. ee ges sree IRE
Additional citations: MEXICO: Colima: R. McVaugh 15551 (Mi).

CITHAREXYLUM STEYERMARKII Moldenke

Additional bibliography: Moldenke, Phytologia 7: 48--l9. 19595
Woldenke, Biol. Abstr. 3h: uf sf 1959} and 35: 1465. 1960; Hock-
ing, Excerpt. Bot. A.5: 43.

AdSitionar seal lg—50 3989
ety Sop ype: ae: “Yold denke, Phytologia 7: spe ° 5
ee Biota 3: 30. 1960; G. Taylor, Ind. Kew. Suppl.

CITHAREXYLUM gpm cenarag Foes
Additional synonymy: terexylen subflavescens Blake ex Molden-
ke, Résumé Suppl. 6: 9, = syn. 1963.

316 PHYTOLOGIA Vol. 13, no. 5

Additional bibliography: Moldenke, Phytologia 7: 50--53. 1959;
Moldenke, Résumé Suppl. ees - 1963.
Bernardi says of this p. ant "madera resistente a la intemperie
-——- bayas sat -- Me aparentenente restringida a la zone
Hu

selva lada'" and records the common name teelvio"
Prem it at 3000 meters altitude. ane esis describes it as a
tree, 8--10 m, tall, the calyx olive- , corolla white, rachis

ve~green
green-buff, leaves dark-green above, olive-green beneath, and
petioles olive-green, growing in cloud forest at 2200 meters al-

titude, blooming in July.

Additional citations: VENEZUELA: Aragua: H. Pittier 9333 at
J. A. Steyermark 9298 (Z). wera Bernardi 2069 (Ve—76
TAchira: Hueck s.n. [Paramo la Negra, 10.).1958] 8] (Ve—l2762), rh

n. [Paramo la Negra, 2725.58] (Ve—-l2762, Ve).

THAREXYLUM SUBTHYRSO: Pittier
Additional vinitueecres Moldenke, Phytologia 7: 53--55 (1959)
and 13: 284. 1966.

Steyermark describes this plant as a vining shrub, the leaves
firmly membranous, dull-green on both surfaces, and the flowers
greenish-yellow, inhabiting dry reforested slopes, flowering in
June. Material has been misidentified and distributed in herbar-
ia as C, dawei Moldenke.

Additional citations: VENEZUELA: Federal District: ristegu-
ieta 793 (Ve--32606); Lasser 742 (W--187959h)5 J. A. toana
86308 (WF (W—-2h30158). Lar&: Tamayo 3757 (Ve=-30045) -

CITHAREXYLUM SULCATUM Mo’
Additional bibhiograply: MoLdenke, Ph Phytologia 7: 7: hos 1959»

Recent collectors de: e this plant as a , OMe
tall, trunk 7 cm, in Faitien at Wears height, gant ey, slight-
ly fissured, leaves coriaceous, deep-green, calyx green, and

fruits shiny-green, becoming violaceous, ” cone tee | canyon etre
pole A meters altitude, flowering in June, and fruiting in

A specimen of Triana 2072, deposited in the Delasoan’ ange
at the Conservatoire et Jardin Bo otaniques at Geneva t
eesent there a Macbride as his type photograph number or 28398, , bu
Ss no
Additional citations: COLOMBIA: Cundinamarca: Cuatre sare
millo, & Huertas 25833 (Z); Kgie = 5067 (W--22535u41); Little rattle % ’ Little
9201 9201 (N); Triana 2072 [Macbride photos 28398] (W--photo).

CITHAREXYLUM TETRAMERUM T. S. Brandeg.
Additional bibliography: +3 C. Standl., Contrib. U.
poli 23: xe & 1238. 1924; Moldenke, Phytologia 7: 63 3 (3999)
he Lundell & Lundell 12332, distributed as C. tetramerm, is
actually C. ly CG. lycioides D. on.

1966 Moldenke, Notes on Citharexylum 317

CITHAREXYLUM TEISTACHE OM Turez
Additional synonymy Githarenylin tristachuym Turcz., in herb.
Additional bibliography: Kunz, Anatom. Untersuch. Teak 73.
1911; Moldenke, Phytologia 7: 63-66. 1959.
Recent collectors describe this plant as a large bush or shrub,
rowing in

» August,
ecenber, and mga in November . Morton 4083 exhibits some
leaves smooth and some hairy.
Additional pe pe pa CUBA: Las V: : Re A. Howard 5650 (W—
somata 6629 (W—1959175) ; as 4083 ne et AS) . Oriente:
Lépez Figueiras 7173 (Bm); Ekman 1509 (W—-2113h55), 8019
iia Lépez Figueiras Figueiras er (W——2226623).

ITHAREXYLUM ULEI Molde:
Additional lavatnevcpae? Moldenke, Phytologia 7: 66--68 (1959)
and eas 302. 1966.

deposited in the Delessert aaa at the Conservatoire et Jar-
din Botaniques at Geneva, was photographed there by Macbride as
his type photograph number 28399, but is not a eae

Additi citations: COLOMBIA: Putumayo: R. E. Schultes 3361
a: BRAZIL: Amazonas: Ule 5501 [Macbride pe ; photos 2399)

ITHAREXYLUM ULEI var. CALVESCENS Moldenke
Adtitional. ba bibliography: Moldenke, Phytologia “fh 68. 1959.
i Additional citations: BRAZIL: VMaranhfo: Frées 1719 (Mi—

UM WEBERBAUERI Hayek
Additional bibliography: Moldenke, Phytologia 7: 71-73. 1959;
Soukup, Biota 3: 30. 1960; Moldenke, Plytologia 13: 303. 1966.
a rec

the Botanisches Museum at Berlin, was photographed there by Mac
bride as his 7 photograph mumber 17603, but is now destroyed.
The Ellenberg 19k6 oh6, distributed as C. weberbaueri, is actually
Cc

G+ quercifolium Hayek.

Additional citations: PERU: Hudmmco: Weberbauer 3731 [Macbride
igen 17603; Herb. U. S. Nat. Arb. 197893] (W--photo of type).
“ —_

rtad: Angulo 1267 (2)

ADDITIONAL NOTES ON THE GENUS ABGIPHILA. XVII
Harold N. Moldenke

A complete list of the abbreviations employed by me for the
names of herbaria in which cited oF or on are deposited will be
published shortly in this sen in my work on the gems Calli-
. Meanwhile, most of them will be found explaine prereptetce
gia 5: 154159 (1955), és “22 (1958), 7: 91-92 (1959), 7
125-1 ( $96) 7: 343 (1961), 8: 95 (1961), 9: 191 (1963), “and

AEGIPHILA Jacq.

Additional & emended synonymy: Aegiphila L. ex A. L. JUSS «,
Gen. Pl., ed. ‘a 107. 1789. Aegiphill. a Briq. ex Moldenke, Résumé
ae ote in syn. 1966. Ageiphila oe in herb.

nded b bliography: Aw L. JUSS ey Gene Pl., ed.
a 107, (2789) rte ed. 2, 120. 1791; L. C. Rich., Act. Soc. Hist.
Nat. Paris 1: 106. 1792 ; Link, Enum. Pl. Berol. Alt. 1: 124. 1821;

° Ze 2h6 65 Schum, in ’ *
Jahresber. 3 (1): 495 & 496. 19023 Loes., Verh. Bot. Ver. Brand.
53: 81 [Abhandl. 246]. 1912; P. C. Standl., Contrib. U. S. Nat.

].

Herb, 23: one & 1253—-125), 1924; Robledo, Lecc. Bot. 2: 499.
190; C. L. Lundell, Contrib. Univ. Mich. Herb. 8: 60, 75, So
& 89, 1942; Moldenke, Biol. Abstr. 27: 2026. 19533 uoldenke, ¥
benac. 2h, 1955; Angely, Cat. Estat. Gen. Bot. Fan. a dbet ios6,
Moldenke, Biol. Abstr. 30: Rar 1956; Angely, aT h&
= 295%) Reitz ee : 49, 66, & . 1959; “Voidenke, Biol.

tr. 33: 1215°(1959) and 2 217.1960; Angely, Liv. Cop. _
er. 35 & 37. "19605 surge Bol. Soc. Argent. Bot. 9: 182

185. 1961; i 260. 1961; Moldenke, Bil.
Abstr. %: 36 (19 21) and and 37: 215 & 1062. 1962; Hocking, Excerpt.
Bot. A.k: ms & 593 (1962) and A.5: hh & 537. 1962;

_ aes tee ag ule, ta k: 81 & 191. 19625 Mol-

’ ° q . 9 3 Vege 7

1964; Reitz & Klein, S 16: hh, BY 6h, 103, & 108. 196k;
Langman, Select. Guide Mis iam. sorter nn ,» 748, & 1010. 196h5
Gooding, ss aeaan & Proc 3-355 & 463- 19655

O, 8. Bot. 21: aa 3, & (el. 1965; Moldenke, P
toga’ 325 : er rere Résumé i: 12, i ook (1965)
Soukup’ (1 i962) 5 records the vernacular name "chirapa~sacha", 4

tT Acgiphi"*
sp. by C. Weurman at Leiden, is a species of Pisonia in the T°t=
iaceae, while the Cuatrecasas 23088 is Petitia nsis Jacde
Recently Hatschbach has collected a strange plant in southern
318

1966 Moldenke, Notes on Aegiphila 319

Brazil which the late N. Y. Sandwith, at the Royal Botanic

dens, Kew, felt is an Aegiphila. He. says, in a letter to me dated
August 3, 196: "Today I examined it and in every flower the gy-
noecium was different, but the tiny, reduced tetramerous corolla
with ); stamens inserted near the top of the tube was alwa,

tip, or it was a long, spirally coiled, hairy organ ending in an
empty inflated ovary terminated by a subulate undivided stylet I
cannot help feeling that this plant may, after all, be a new
‘lusus' of an Aegiphila in the group of Ae Sellowiana and A. ver-
ticillata: the foliage and indumentum are iy nti arb ah indeed
te) ese, & they are so common in S. Brazil. e inflorescence

oe SOUEIFER Moldenke
bibliography: Moldenke, logia 7: 456 (1961) and
€ Bs. 96h; | Moldenke, Biol. Abstr. 36: 036 (1961) and 37: 1062.

on f this plant has eth misidentified and distributed
in herbaria as = odontophylla . Sm., a species to which it
is certainly very closely ited.
Additional ¢ oh Birger COSTA RICA: Alajuela: A. F
[Stork 4210] (Mi). COLOMBIA: Cauca: Sneidern ser (2103 e

AEGIPHILA ALBA Moldenke
Additional “bibLiogrepty: Moldenke, Phytologia 7: eeu 1961.
Cazalet & Pennington describe this species as a tree, 40 feet
tall, 8 inches in diameter at breast height, hilous, "the

female flowers in axillary clusters; fruit a green acorn", They
found it in a seasonal rainforest at 1000 feet altitude and iden-
tified it as "Aegiphila aff, valerii Standl." Eggers also de-
Scribes it as a tree 3 feet tall, flowering in February. J&tiva

ling call it a "rounded tree", with "white cauline flowers",
growing in forests and virgin forests at 700 meters altitude,
Tlowe July. Material of this species has been misidentified
and distributed in herbaria as A. monstrosa Moldenke.

fig wane Dieter ECUADOR: Guayas: Eggers 1448 (A). Pich~

t & Pe ton 5090 (N); J&tiva & Epling 35h (La—

iseis) See a Than ies La--53199) «
AEGIPHILA ANOMALA Pi

Additional walnguss “Hegiphila anonala anomala Pittier ex F. Miranda,

Bol. Soc. Bot, Mex. 26: 160, sphalm. 1961.
Additional “bib bliography: F. Miranda, Bol. Soc. Bot. Mex. 26:

320 PHYTOLOGIA Vol. 13, no. 5
160 & 169. 1961; Moldenke, Phytologia 7: 457. 1961.

AEGIPHILA AUSTRALIS &M
Additional am cremaae “Yoldenke, Phytologia 7: 458 (1961)
and 8: of 1961; Rambo, Pesquis. Bot. 12: 21 (1961) and 21: 6 &
[59]. 196

Rambo (i968) cites Herb. Anchieta 333 [Sehnem sen.) & 61186

Grande do Sul, the one not credited to someone else having been
collected by himself. He calls the plant a "Small tree, up to 7
m high", growing in "Half-high woodlets, secondary growth" in
the Sehinestens Highlands only" in Rio Grande do Sul, and gives

its general distribution as "SC [Santa Catarina] and RGS [Rio
Grande do Sul]".

IPHILA BARBADENSIS Mi

Additional + Steite f "waide » Phytologia 7: 458. 1961;

» Loveless, & Proctor, Fl. Petes 35h & 463. 1965.

AEGIPHILA BOGOTENSIS (Spreng.) Moldenke

Additional synonymy: Aegiphila bogotensis (H.B.K.) Moldenke,

lim. Alph. List Invalid Names 1, in syn. 1940.

Additional bibliography: Moldenke, Phytologia 7: 458—-h59
(1961 a i 8: 16. 1961.

casas and his associates describe this plant as a tree,

825 1 z dau, the young branches ochraceous-ferruginous,
leaves flexible, rather dark and subcoriaceous or coriaceous and
yellowish- or dark-green, shiny above, lighter beneath, the calyx
greenish-ochraceous or ochraceous-ferruginous, the corolla

ruary, growing for ng bus
and on disturbed slopes, at 2700-3140 meters eltitade, steyar-
mark records the vernacular name 1

A

1142916). Cauca: Cuatrecasas & Willard 26166 (Fg). Cundinamarca?
Garcia-Barriga 1331 (Ve--30779, W—1907600). Magdalena: Cuatre-
casas & Castafieda 25219 (Z). Valle del Cauca: Cuatrecasas, , Lopes
Figueiras, & Willard 26817 (W--2455933). VENEZUELA: Lard: J. Ae
pee oe 50265 (Ve--28826). Mérida: Garcia-Barriga 13294 (Ve)-
Tachira: Aristeguieta 2568 (Ve—39795, W—2248871) «

AEGIPHILA BOGOTENSIS AEQUINOCTIALIS Moldenke cs,
Additional bibli hse eee Biol. Abstr. 27: 2026. 19535
Moldenke, Phytologia 7: 159. 1

PHILA BRACHIATA Vell,
pect egiiors bibliography: Herter, Florul. Urug. 10h. 1930; #
Ne eat Sudan. Bot. liz 185. 19375 Ranbo, Sellowia ie “a

* ‘dle 195
1961} Rambo hana t Bot. 12: 21 (1361) and 21: CoP 196s

ic ld>~scttsaridhcseesticaieiahidiinandaabiamiinameoaianndipaaiie

1966 Moldenke, Notes on Aegiphila 321

Bormmtfller 581 is described as having had yellow flowers. The
Species has been found ast at altitudes of 50 to 900 meters,
in woods, forests, forest margins, and ruderal. Material has
been misidentifie a and distributed in herbaria as A. sellowiana
Cham. and as Acnistus sp. The Sellow s.n., photographed by Mac-
bride as his type photograph mmber 17590, was deposited in the
oe of the Botanisches Museum at Berlin, but is now destroy-

Rambo (1965) cites Herb. Anchieta 3979 from Uruguay,
6687, 9217, 26483 [Hens s.n.], J, 26508 [Henz s.n.], 26211, ee
32309 [Buck s.n.], 326)0, 32928 [Friderichs ate) 32936 [Frider-
ichs s.n.], 35579 [Vier sen.], 37330, hh610, 48h13 [Sehnem 225)],
48423 [Sehnen 2149], 50618 [Sehnem nem 3919], 87579 [Pivetta 657],
6205 [Camargo 1977], 62529 [Camargo 228h], 62539 [Camargo 216h],
625u1 (Camargo 2088], 6256 [Camargo 2077], and 62565 [Camargo
aa] from Rio Grade do Sul, Brazil, all collected ‘ted by himself
where otherwise designated. He describes the plant as
"meceaiert up to 10 m high", growing at "Edge of the rain forest,
secondary forest, Campos woodlets"...."through the whole of the
forested area" in Rio Grande do Sul, with a general distribution
an Rio de Janeiro and Paraguay down to RGS [Rio Grande do Sul]
@

Sess citations: BRAZIL: Paran4: Curial 1052 (W~—1951772);
Hatschbach 6327 (W—-2320786). Rio Grande do Sul: Bornmtfiler 345
(G), 581 (G); Sehmem 2149 (B). Santa Catarina: J. F. T. Midler
Son, [Rio Claro] (P); Smith & Klein 13233 (Ac); Smith & Reitz
12h51 (Ac), 12759 (Ac). State undetermined: Sellow s.n- ~[iacbride
Photos 17590] (w- (W——photo).

AEGIPHILA oldenke
oh aig pec stae Moldenke, Phytologia 7: 459--1)60. 1961.
nal citations: BRAZIL: Amazonas: Ducke II hkl (W—

1875380);, Krukoff 5060 (W162 821).

AEGIPHILA CANDELABRUM B
ues enhances: Moldenke, Phytologia 7: 60--461 (1961)
The — 8120, photographed by Macbride as his type photo-
phe oa deposited in the Delessert Herbarium at the
Conservatoire et Jardin Botaniques at Geneva. Material of this
Species has been misidentified and distributed in herbaria unde
the name A. cuspidata Mart. & Schau. On the other hand, the Wool-
ene 1OL6, “aistributed as A. candelabrum, is actually A. lanceola~
Moldenk eee
Additional citations: PARAGUAY: Hassler 7974 (Mi), 8120 [Mac-
bride photos 21,621] (W—photo of type).

AEGIPHILA CAUCENSIS Mol
Additional weeavuerangr Moldenke, Phytologia 7: 61. 1961.

322 PHYTOLOGIA Vol. 13, no. 5

records the vernac oes name "rupifias" for it and notes, “compare
no. 5496", a collection which > were not as yet s
Additional citations: PERU: Junin: Woytkows ia 6432 (2).

AEGIPHILA CEPHALOPHORA S
here bibliography: Sr ddcais: Phytologia 7: 461--l62.
19 Me
nal citations: PANAMA: Barro Colorado Island: Shattuck
988" (t-2366733) .

ABGIPHILA CHRYSANTHA Hayek
Additional bibliography: Moldenke, Phytologia 7: bé62. 1961.
The Eggers 14348 photographed by Macbride as his type photo-

wee mumber 20349 is pa Pay in the herbarium of = ho Botaniachs
eum in Munich, while the Poeppig 231) photographed by him

os s type photograph number 3313 is deposited in the herbarium ra

the Naturhistorisches Museum at Vienna. Straw describes the

plant as a small tree, 3 meters tall, with greenish Doses ("but

past flowering”), growing in partial shade near a river on open

Additional citations: ECUADOR: Guayas: 14348 [Macbride
photos 20349] (N--photo). Manabi: greed Ly a (W—-1618703, W--
1618777). PERU: Loreto: Poeppig 2314 [Macbride photos 34313]
(W—-photo); Straw 2426 (W—2h51475). MOUNTED CLIPPINGS:

Engl., Bot. Jahrb, 2: 171--172. 1909 (W).

AEGIPHILA CONTURBATA Moldenk
kati Saal. bibliography: Molde e, Phytologia 7: 62. 1961.
The 8.n, specimen wee reetiat by Macbride as his type
Bane Sn umber 28377 is deposited in the i sere "Narbart i
at the Co: nose ntciice ae Jardin Botaniques in Geneva
Additional cita : BRAZIL: Maranh%o: Newman s.ne [Macbride
photos 28377] femchete. of type).

ote CORDATA Ws
tional tinue a : 46263.
(a 61) ave hs 9 i phy: Moldenke, Phytologia 7: 6
he Poeppig 2158 specimen photographed by Macbride as his type
thik raph number 34312 is deposited in the herbarium of the Nat-
urhistorisches Museum at sae

tional citations: PERU: Junin: Woytkowsks 5977 (W—-2h5309)-
Loreto: Poeppig 2158 [Macbride photos 34312] (W—-photo of type)

anorearla weet (Ruts & Pav.) Mat ldenke of abo

tS) : Spreng. in L., Syst. Veg- Mid

Wor (1825) and 5: 126. 1828; Sldenke, - Phytologia 73 b3. 2 1961
The specimen of Ruiz & Pavon s.n. photographed by M

his type photograph number 28378 is deposited in the ee,

<7 TS.

1966 Moldenke, Notes on Aegiphila 323

Herbarium at the Conservatoire et Jardin Botaniques in Geneva.
Additional citations: PERU: Hudmuco: Rufz & Pavon s.n, [Mifia;
Macbride photos 28378] (W-~photo of isotype).

AEGIPHILA COSTARICENSIS Moldenke
onal bibliography: m7 eH = 7: 463-64.
1961; 2s pagan Résumé Suppl. 13: 2.
Matuda foun acres species in i psotmry: in "November. Steyermark
describes it as a tree, 5 meters tall, with membranous leaves
that are Foca above and mer flowers, ecm in rheng ye

along rocky strean- t 750--900 me
Additional cite tosis MEXICO: “Chlapas: “ers ae. (a), 6152
nts VENEZUELA: Carabobo: J. A. Steyermark 94313 (Z), 9

AEGIPHILA CRENATA Moldenk.

Additional bibliogra meee - Paran. 7: 6 & 11. 1957;
Moldenke, Phytologia 7: jis (5st) oo "8: 16. 1961.

The en of Dusén 1051, photographed by Macbride as his

Additional citations: BRAZIL: Parané: bake iosla [i (Macbride
Photos 30182] (W-—photo of isotype).

ABGIPHILA CUNEATA Moldenke
Additional bibliography: M Phytologia 7: 46) & 467.
ig. Moldenke, Biol. Abstr. ae "host (1961) and 37: 215. 1962;
ened, Résumé Suppl. 10: 2. 196).
and

mM. ‘ flattish samt the yo aig
Pices of the branches pile eS with mauve hairs, the flow
white, blooming in September, and growing at 1450 "tet Staats.

Additional citations: ECUADOR: Napo-Pastaza: Grubb, Lloyd,
Pennington, & Whitmore 1658 (N, N).

AEGIPHILA CUNEATA var. HIRSUTISSIMA Moldenke
Additional bibliography: Moldenke, Biol. Abstr. 27: 2026. 1953;
Moldenke, Phytologia 7: 67. 1961.
main cr penne Vv Brand. 53: 81
tional bibliography: Loes., Verh. Bot. Ver. e
[Abhandl, 246]. 1912; a A Standl., Contrib. U. S. Nat. Herb. 23:

tional mam Aegophila elata Sw, ex Hocking, Dict.
halm, 1955.

Pharmacog, 6, sphalm -
Additional bibliography: C. L. Lundell, Contrib. Univ. Mich.

32h, PHYTOLOGIA Vol. 13, no. 5

Herd. 8: 60. 192; ae eine _— ireargees 6. 1955; Mol-
pre Phytologia 7: 68-469. 1961; Gooding, Loveless, & Proctor,
Fl. Barbetes 35h, 55, & 63. “19655 "Woldenke, Résumé Suppl. 13:

2. 19
Recent collectors describe this plant as a shrub or arching
shrub, 2 m. tall, a small tree, or a woody vine or liana, of-
ten somi-climbing, the stems 1 inch in diameter; leaves papery or
subcoriaceous, dull or — glossy and medium— or deep-green
above, dull and paler beneath; calyx green; flowers cream-color
creamish, aeons greenis ieyelion, or —— fruit obovoid,
vermillion-o » 1.2--1.5 cm. long, about 8 mm, wide. It has
been found growi “4 along roadsides in secondary f pages and on
broken ridges, in thickets on “te hillsides, ait Si econdary
growth on banks of small rivers, at altitudes of 350 on 1250 m.,
in

"creamish" on Gentle 457, "light-yellow" on on J. V. Santos 292,

"yellow" on m Aristeguieta & Pannier 1856. “Tundell ell (1942)
cites Matuda 3031, 3081, & 3406

dditional cit wh prema MEXICO: Oaxaca: J. V. Santos 29% (ii).
Vera Cruz: J, V. Santos 2315 (Mi). GUATEMALA: El Petén: in: Contre=
ras 3259 es 337% 74 (Ld, S). BRITISH HONDURAS: Gentle 456 56 (Rf),
UL57 (RE, Rf), 6752 (Ld, S), 7726 (1d), 8009 (Id, S). JAMAICA:
G. R. Proctor 23913 (Mi). VENEZUELA: Barinas: Breteler 3735 (¥)-

Bolfvar: J. A. Stey Ss 2728 (Ve). Monagas: Lasser “Tasser & . Vareschi
as ee. » Yaracuy: Aristeguieta & Pannier 1856 —a
9).

ae ELEGANS Moldenke 961
onal bibliography: Moldenke, Phytologia 7: 469. 19015
Moldenke, Résumé Suppl. 12: 3. 1965.

Asplund 8935 seems to be a mixture with Cornutia odorata
(Poepp. & Endl.) Poepp. —- the Stockholm herbarium sheet heet is Cor-
nutia, but the United States National Herbarium sheet is Aegiph-
ila, although distributed as Cornutia. ass

Additional citations: ECUADOR: Napo-Pastaza: Asplund 8935,
part (W--222),015).

AEGIPHILA FARINOSA Moldenke
Additional bibliography: Moldenke, Phytologia pene 470. 1961-
The species has been collected in * fruit in Jan
Additional citations: COLOMBIA: Valle del Cauca: at Lépes Figue

eiras 8307 (W--2455215).

AEGIPHILA FENDLERI M 961
Additional bibliog sane Moldenke, Phytologia 7: 71. 190+»
Steyermark describes this mas t as "vining"; leaves subcoria~

ceous, deep dull-green above, vous-green beneath; calyx ere° waa
ish; Corolla srenecauiing: oone at 1300—-1,00 m. altitude,

1966 Moldenke, Notes on Aegiphila 325

flowering in October.

Additional citations: VENEZUELA: Aragua: H. Pittier 14982 (W—
1832690); J. A Steyermark 89931 (Z); Tamayo . 1643 (W—-1833098) .
BRAZIL: Amapé: | Murca farca Pires, Rodrigues, & Irvine 5039 (N).

AEGIPHILA FERRUGINEA Hayek & Spruce
Additional cea ea Moldenke, Phytologia 7: 471. 1961.
The specimen of Spruce 5473, photographed by Macbride as his
type photograph number 330, is deposited in the herbarium of

ie e
psc ea rane citations: ECHADOR: Chimborazo: Rimbach 616 [Herb.
« Yale School Forest. 3199] (W--17)2057). Pichincha: Aspl
sigs 2223162) Spruce 5473 [Macbride photos 34311] (W-—photo

AEGIPHILA FILIPES Mart. & ees
Additional bibliography: Schum, in Just, Bot. Jahresber.
is sg Se 495. oe Moldenke, ee sea 7: int make 1961; Soukup,
io
Soukup aes — for this plant the vernacular
"c etanabepisache a", a name, however, which is also applied ‘
species of Psychotria, Vernonia, and Wulffia. Material has been
misidentified and distributed in herbaria as A. martinicensis
Jacq. The specimen of Martius s.n., photographed by Macbride as
his type photograph naar mber 20350, ie Moe endo in the herbarium
of the Botanisches Mus
Additional rept sey Preaeen agdalena: H. H. Smith 1831
(Mi). BRAZIL: Amazonas: Krukoff s28 ¢ (W662). Pard Paré: Martius
8.n. (Macbride photos 20350] (N— (N—photo of cotype).

IPHILA FLORIBUNDA Moritz & peel
Additional bibliography: Moldenke, Phyto seg ag r lite he
The specimen of Moritz 1765, Ssotseienked by Mac s his

type photograph number 31,310, is deposited in the Biative of the
Witevliutorte hie — t Vienna.

Additional citations: VENEZUELA: Aragua: Moritz 1765 [Macbride
Photos 34310] rls

AEGIPHILA FLUMINENSIS Vell.

te, Piytalogs no yr gh Angely, Fl. Paran. 7: 11. 1957; Molden-

logia 8: 16, 196
ror are pia as "cream" on Hatschbach 6616 and as

enti tds on Glaziou 3067. The plant has been found growing "em
terrenos baldios" by Hatschbach. It has been collected in anthesis
in reeves and Dec salel; The peri se name is sometimes uppercased.
‘ dditdonal citations: BRAZIL: Paran4: Hatschbach 6616 (W
32165), Rio de Janeiro: eink 3067 (Ja—11193).

IPHILA GLABRATA Moldenke
Additional bibliography: Moldenke, Phytologia 7: 473. 1961.

326 PHYTOLOGIA Vol. 13, no. 5

Wurdack describes this plant as a woody vine, climbi ed Te m.,
th white flowers, occasional in rainforest we ‘niddle n
slopes, at 550--750 m. altitude, blo in 0

oming
Additional citations: PERU: Loreto: Wurdack 2390 * (W——21,03697) «

AEGIPHILA
ger bibliography: Moldenke, Phytologia 7: 73. 106s
his plant has been found growing at 1200 m. altitude.
Lasser 308, distributed as this species, is actually A. gan

H.B.K.
Additdonal citations: VENEZUELA steguieta & Fol-
dats 1451 (Ve—33507). BRAZIL: Sends ne - Bo (W—21,39039) «

AEGIPHILA GLANDULIFERA var. PARAENSIS M
= eee cong bibliography: Moldenke, ie pa 7: 473—-L7h.
9
Additional citations: BRAZIL: Amap&: Murga Pires, Rodrigues, &
Irvine 51052 (N).

GLOMERATA Benth
tional & emended bibliography: Benth., Bot. Voy. Sulphur
ashe “Tabs Moldenke, Phytologia 7: h74—h75. 1961.

AEGIPHILA GRANDIS Moldenke
= D srerinn nas bibliography: Moldenke, Phytologia 7: 75-76.
Lasser describes this ere as a tree, 10 m, tall, with a
columnar trunk, irregular crown, and very fragrant + flowers. He
xeeretn the vernacular name "oreja de burro". The specimen of
riana 2080 photographed by Macbride as his type photograph num
oS ey a laos tage ed in the Mie ua Herbarium at the Conser-
ques at Gen
ides citations: COLOMBIA: aabaenesat Triana 2080 [Mac-
pie photos 28379] (W--photo). VENEZUELA: Bolfvar: Lasser 1167 ser 1167
e).

AEGI

PHILA GRAVEO
Additional balntecmetty ‘ian. Phytologia 1 hie tego
The specimen of Lund 796 photographed by Mac

fied and distributed in herbaria as So Sp.

Additional citations: BRAZIL: Bahia: Duarte 6847 (Herb. Brad.
24404] (W#—2436357). So Paulo: Lund 796 [Macbride photos 7880]
(N--photo of type, W--photo of type).

AEGIPHILA GUIANENSIS Moldenke
Additional bibliography: Moldenke, Phytologia 7 Ts & rohit 1961.
The specimen of Triana 208) photographed by Mac on: Ae

type photograph mmber 25300 is deposited in the naar Herbar-

1966 Moldenke, Notes on Aegiphila 327

ium at the Conservatoire et Jardin Botaniques in Geneva

Additional citations: COLOMBIA: Cundinamarca: Triana "208 [Mac-
bride photos 28380] (W—-photo). VENEZUELA: Gudrico: Aristeguieta
4215 (Ve--l6119).

LERI Briq.
Additional synonymy: Aegiphila hassleri Chod. ex Moldenke, Ré-
tor ae Suppl. 3: 29, in syn. 1962; Rambo, Pesquis. Bot. 21: 7 &
9
eaters bibliography: Rambo, B seye pies 7: 207. 1956; Angely,
Fl. Paran. 7: 11. 19573; Moldenke, Phytologia 8: 16. 1961; Rambo
Pesquis. Bot. 21: 7 & ts9]. 19653 foiincks, Résumé Suppl. 13: 3

The specimens of Hassler 3193 photographed by Macbride as his

under the name Citharexylon sp. The Del Puerto 323k, cited below,
bears a ——- Os Pag ce a muestra No. 4166", a specimen not
as yet se
Rambo (1968) ¢ cites Herb. Anchieta 990 & 9952 from Santa Cate-
rina, 11251, Loos, 45761, 49h79, 54254 [Emrich corte goths [Ca-
nargo 1138], & & 62037 ’ [Camargo 2037] from Rio Grande , Brazil,
all collected by him himself except breig otherwise Ries ct. He
calls the plant a "Small tree, up to 7 m high", at the
of the forest, secondary growth", is Rio Grande do Sul, "Ap-
parently mextonre forested area only", "with a general distribution
"From P ar through South Brazil and Argentina Meso-
potamia to RGS [Rio Grande do oe and Uruguay".
Additional citations: BRAZIL: Santa Catarina: Smith & Reitz
22694 (Ac), 12751 (Ac). PARAGUAY: Hassler 3193 [Wacbride photos
263) (W—-photo of cotype). URUGUAY: Del Puerto 3234 (Z). AR-
ENTINA: Misiones: Montes 121 (W--20)9836)-

AEGIPHILA HERZOGII Mold.
Additional biblio ea Moldenke, Phytologia 7: 477. 1961.
The specimen of Herzog 1369 photographed by Macbride as his
aig airioaeaph number 28381 is deposited - the Delessert Herbar-
t the Conservatoire et Jardin Geneva
meen onal citations: BOLIVIA: Santa ret Herzog 1369 [Mac-
bride photos 28381] (W--photo of isotype).

AEGIPHILA HIRSUTISSIMA Moldenke
Additional bibliography: Moldenke, Phytologia 7: 4,78. 1961.
us Burkart 17014, distributed as this species, is actually A.
= 8 oB eK

a Aadh tonal citations: COLOMBIA: Bolfvar: F. W. Pennell 4563 (W—

328 PSYT-OLOG IA Vol. 13, no. 5

AEGIPHILA INTEGRIFOLIA ner Jacks
Additional synonymy: Callicarpa foliis foliis ovatis acutis integer-
5 densis axillaribus

densis axiitaribus ot verm*

pat a fe ° .
Spreng. in _ Syst. ‘a — 16, 1: 419 & 421. 1825; J. Aw & de
H. Schultes, Mant. 3: "3 & 51. 1827; Spreng. in L., Syst. Vege,
ed. 16, 5: 126. 1828; Stapf, Ind, Lond. 1: 526. 1929; Moldenke,
Phytologia 7: 479—L80. 1961; Tamayo, Bol. Soc. Venez. Cienc.
& 165. 1961.

nal illustrations: Jacq., Sel. Stirp. Amer. Hist. pl.
pas & fig. “ 7. 1%633 TG Sel. Stirp. Amer. Hist. Picta pl. 259,
Me. 6 [in color]. 1780.

McCarroll describes the fruit of this species as red. The
plant has been found in anthesis in January and in fruit in April.
An additional common name is "ganzbluttrige Schtnbeere". The
Borsotti ss and il. Williams 13401, distributed as this species,
are actually A. parviflora Mi Moidenke, Lasser 1167 is A. sessili-
flora Moldenke, and Sehnem 3781 is not , verbenaceou

pata (1825) and Schultes (1827) place this toute! in i
synonymy 0. e North American Callicarpa americana L., a P

pabie > Seay

Additional citations: COLOMBIA: Boyac4
232h023). VENEZUELA: Amazonas: Ll, Williams ess é€ Fn 1Seh616),
15080 (W—1877h12). Anzoftegui: Aristeguieta & Agostini 4714
(N). Aragua: Agostini & Farifias 107 (B). Bolfvar: H. M. ~ Curran
26 (Ve—h1363); Lasser 1291 (Ve), 1604 (Ve); Tamayo s.n. . (Sta.
Elena, Marzo 1946] (Ve). Mérida: E. E, Le Little 16209 (Ve). ona-
gas: Aris eta 3916 (Ve-562). State te undetermined: Tamayo
2982 slate e—27077). BRITISH GUIANA: Tutin 59 (W—-1743381). BCUA~
? Napo-Pastaza: Asplund 8868 (W-—2224005), 10166 (W—222h168)5
Mexia 7173 (W—-1663739), 7187 (W—16637h7), 7289 (W--1663799) -
PERU: Cuzco: V. argas Calderén 7385 (W—-1995360). Loreto: 0. Ve
Nufiez 2449 (W—-2121593); Schunke 343 (W-—-1459230). Puno: “MeCar-
roll | Zou 2h Ot (Mi), 132 (Mi). BRAZIL: Amap4: Irwin, Egler, & uurga

1 (N).

EEE

IPHILA INTERMEDIA Moldenke

Additional begga 14 Neldeniet, Phytologia 7: 480. 19615
Moldenke, Résumé Suppl. 12: 2. 1965.

It should be noted vas ale some of the leaves on Ll. Willi- B
ams 1317), cited below, greatly resemble those of A. integrifolia
(Jaeq.) Jacks. The specimen of Herb. Gen. Mus. pore, 2200 sii
graphed by Macbride as his t hotograph number 28382 1s
posited in the Delessert 2 Sim the Conservatoire et Jardin

Botaniques in Gen

1966 Moldenke, Notes on Aegiphila 329

Additional citations: VENEZUELA: El Ratén Island annem
Ll, Williams 1317 (W-—-228869). BRAZIL: Maranh%o: Herb. Gen.
Mus . Para. 2: 2270 [Macbride photos 23382] (W--photo of isot; carroe

AEGIPHILA LAETA H.B.K.
Additional bibliography: Moldenke, Phytologia 7: 481. 1961.
Be sate of this species has been misidentified and distributed
herbaria as Callicarpa sp. On the other hand, the Duke 1555,
dstrtata as pe laeta, is not verbenaceous.
Additional citations: PANAMA: Darien: Duke 4116 (Z). VENEZUELA:
Zulia: Mocquerys rys 1018 (W238 3053) «

ae A ovghag (Aubl.) Gmel.

few meters tall; leaves chartaceous, bearing green “glands beneath
near the base of the lamina; corolla infundibuliform with patent
limb, very pale greenish yellow, almost white, very fragrant;
style concolorous » Llong-exserted, furcate; anthers brown", flow-
ering in January.

he Peddena 2001, distributed as A. laevis, seems to be a spe-
cles of Pisoni: Pisonia or or some related genus

Additional citations: SURINAM: Kraner & Hekking 2746 (N).

AEGIPHILA LANATA Moldenke
Additional bibliography: Moldenke, Phytologia 7: 82. 1961.
The specimen of Glaziou 21917 photographed by Macbride as his
type photograph number 20383 is deposited in we — Her-
barium at the Conservatoire et J otaniqu
Additional citations: BRAZIL: Brasilia: ane epi ngs
7823 (N). Goids: Cobra & Oliveira 2h (Z); Cobra & Sucre 30
(ae); Glaziou 21917 [Macbride photos 28383] (W—photo of oe ian).

ag si omgeterie Moldenke
bibliography: Moldenke, Phytologia 7: 482—L83
asa) ie ts 13: 321. 1966.
Colston describes this plant as follows: "Woody climber, 1
n. 10 cm. diam, at base, with fairly clos e branches at teen
ties; hse pale era h-brown with reticulate £ fine lines; on trees
in wer rolla yellow, quickly falling; filaments
white; ches frelas style white; calyx & peduncles pale yellowy-
reen; leaves dark yellow-green, with slight sheen; midrib and
lower surface very pale ; branchlets pale yellow-green”. matersa3.
wee pect penye misidentified and distributed in T parbario as A, candela-
"Additional citations: BRAZIL: Minas Gerais: Heringer 6371 (2).
PARAGUAY: Woolston 1046 (W—228165)) .

AEGIPHTLA LAXICUPULIS Moldenke

330 PHYTOLOGIA Vol. 13, no. 5

Additional bibliography: Moldenke, Phytologia 7: 83. 1961.
This species has been found growing at 600 feet altitude by
Allen & Severen, misidentified and distributed in herbaria as A.

martinicensis Jacq.
Additional citations: EL SALVADOR: San Miguel: Allen & Severen

AEGIPHILA LAXIFLORA Be
Additional biblio cera: Moldenke, Verbenac. 29-30. 1955;
1.

specimen of M, R. Schombur 772 photographed by Macbride
as his type photograph number 2036] is deposited in the Delessert
Herbarium at the Conservatoire et Jardin Botani met ” Geneva.

Additional citations: eee Bolivar: J ge A. teyermark
57719 (Ve—26393). Monagas: J. A. Steyermark Eis (Ve~—36230) «
BRITISH GUIANA: M. R. Schambur ae Hie t12 [Macbride photos 28384] (W—
photo of isotype).

AEGIPHILA LEHMANNII Moldenke
as “peas bibliography: Moldenke, Phytologia 7: 483—l8h.

The specimen of Triana 2083 photographed by Macbride as his
type photograph number 26365 is deposited in the Del geees Herbar-
nave at the Conservatoire et Jardin Botahiques in Genev:

tional citations: COLOMBIA: Boyac4: Lawrance 156 (—
22388). Cauca: Sneidern 1660 (W--2103662). Chocé: Triana 2083,
in part (Macbride photos Sictos SteT (W—photo).

AEGIPHILA maroon Mol
Additional bibli areca Moldenke, Pn kg in ble UB te
Lge fruit of this species is decided] ; » and
not at all wrinkled, tetragonal, or rinaty | when pirrte
Ao tian citations: VENEZUELA: Carabobo: J. Ae Steyermark

ABE LHOTZKIANA C os ueader
Additional biblio : C) Fl. Paran. 7: 11. 19573
ke, Phytologia 8: Locrpi scan
The specimen of Casaretto 2022 photographed by Macbride as his

type photograph number 2161) $e coponisos in the Delessert ener
ium at the Conservatoire et Jardin Botaniques in Geneva, beg
Sellow s.n, [Brasilia] photographed by him as his type Snotograph
mmber 17585 was deposited in the herbarium of the Boteni solr yu-

eum in Berlin, bat is now destroyed. The Eitens found this oe
cles in "degraded cerrado; young barbatimf%o, trees and shoots dem
m, tall, spaced 2—5 m, apart; ground recently burned; shrub 4s
2m, tall, thin, erect, unbranched until the top", flowering in

Additional citations: BRAZIL: Amapé: Murga Pires , Rodrigues, &
Irvine 51116 (N). Bahia: Casaretto 2002 [Macbride Tide photos 2yeus)

1966 Moldenke, Notes on Aegiphila 331

(W--photo). So Paulo: Eiten & Eiten 24,35 [from plant 6] (N),

2435 [from plant 7] (W—-2371219); Mat’ Mattos & & Mattos 8418 (W--245192);
Mimura 23 (N), 49 (N), 216 (W—2hl5188). State undetermined:
Sellow s.n. (Brasilia; Macbride photos 17585] (W-—-photo of type).

ae Dot upipenis Turez
mal & emended bibl iography: - Soc. Nat. Mosc.
% (2): (218-219. "28635 Moldenke, "rigtologie 73 = ia 1961.
imen a hlim 688 photographed by Macbride as his
se > pacers: ae er 24616 is i? in the “opi Herbar-
ium at the Conservatoire et J n Botaniques in Geneva.
Turezaninow se 3) describes this plant as follows: "Ae, ramis,
petiolis, paginae inferioris foliorum costa atque panicula paten-
tim pilosis; foliis breviter petiolatis basi obtusis in petiolum
brevissime decurrentibus, oblongo-lanceolatis longe ac
integerrimis, same fere glabris; paniculae terminalis, folia pa-
rum superantis pedunculo ad medium refracto; cymis oppositis numer
osis, erg deflexis; bracteis ovatis vel bang sian acum-

dditional citations: COLOMBIA: Santander: Schlim 688 [Mac-
bride photos 2,616] (W--photo of isotype).

AEGIPHILA LUSCHNATHI S

Additional bibLograpiy: omapers Phytologia 7: 485. 1961.

The specimen of Luschna . (Herb. Martius 10,0) photographed
by Macbride as his type pho Soro number 7879 is deposited in the
Delessert Herbarium at the Conservatoire et Jardin Botaniqu

Additional citations: BRAZIL: Rio de Janeiro: Luschnath s.n.
(Herb. Martius 100; Macbride photos 7879] (W--photo of cotype) 5
Miers "3712 (W—-2383081)

AEGIPHILA MACRANTHA Duck

Additional hibiidesantes Moldenke, Phytologia 7: oolong nin

ermark describes this species as a woody vine,

or high-climbing liana, the leaves subcoriaceous, ee on,
pale-green beneath with raised veins, corolla buff-cream, frui
salys ~green, urning yellow, growing at altitudes
of 250 to 800 meters, flowering in August, fruiting in December,
Material has been misidentifed and distributed in herbaria as A.

ional stations Pisses ey rag ro Je Ae ermark
86660 (Ve), 88157 (N). Miranda: J. A. Steyermark 89365 t sosee CY
BRITISH GUIANA: Sandwith 1202 Fie. 2 BRAZIL: Par&: Ducke
843 (W—1875670).

AEGIPHILA MARTINICENS
IS Jaca.
Additional synonymy : Aegyphile martinicensis DC. ex Moldenke, Ré~

332 PHYTOLOGIA Vol. 13, no. 5

sumé Suppl. 13: 6, in syn. 1966. Aegyphila martinicensis Jacq.
ex Moldenke, Résum né Suppl. 13: 6, in syn. 19
Additio

; Voldenke 8, piotcon ogia 13: es & 330. + 1966
Recent collectors describe this plant as sin arching
shrub, or small scraggly tree, 5--15 feet tall, h white or
whitish flows and orange fruit, growing in ond or, stone
walls, in mossy forests on northern ridges, or "common" in wood-

Allen & Severen 689), distributed as A. martinicensis, is actual-
ly 4. A. lTaxicupulis Moldenke, while H. He Smith 668 is A. mollis H.
B.K, and H. H. Smith 1831 is A. filipes Mart. & S % Schau. The flow-
ers on Webster, “Ellis, & Miller a are described as “whitish",
but on R. A. Howard 11158 8 as "Ww mw,

Additional citations: PANAMA: Canal Zone: Duke 4685 (Ca). Da
rien: Duke 411) (E). PUERTO RICO: Alain soe (G, oe ee! S3j)5
Otero 270 (Mi), 308 _ vane aL : Dominica: W
1320 (G), 3093 (G), 3824 (G) & Hodge 1258 (G), eg “G )s
1631 (G), 2073 (G), 2572 (ay. BOE wre 2 = 29s y(G), 3077
G)3 StehlS & Stehié 6329 (W245 36 stehlé
2813 (W—2)53662); Stehlé & Stehlé 7019 pee a 3 rs
Ellis, & Miller 9110 (Mi). WINDWAHD ISLANDS: Grenada: Hunmnewell
19279 (Rf). Martinique: H. Stehlé 2393 (W—2453694); Stehlé &
Stehlé ee (W—2153699); Webster, Ellis, & Miller ae 8 (ii) «
Saint | : R. S. Cowan 1571 (N). Saint Vincent: R. A. Howard

—

22158 ney VENEZUELA: Zulia: Moc ocquerys 910 (W—2383052)«

IPHILA MARTINICENSIS var. OLIGONEURA (Urb.) Moldenk
Additional epepoenndterd Yoldenke, Phytologia 7: 1,88. 1961.
W. T. Stearn regards this s a valid species. He 8

that W. Harris 11716, identified =e me as A, swartziana Urb., is
amen representative of the female-predominant form of A. oligom

IPHILA MEDITERRANEA V yh
Additional bibLiogeapty: “Moldenke, a ologia 18 org ier ‘
Glaziou 1546 is said to have been oe mn from

"whitish" corollas. stein 457 ual valyateeth pe are ~ at

1966 Moldenke, Notes on Aegiphila 333

long-cuspidate nor setaceous-tipped like those on most of the Rio
de Janeiro specimens — it was misidentified as Cordia sellowiana
Cham. and distributed as such in herbaria, An isotype of A. ces-
trifolia Gardn., deposited in the herbarium of the Naturhistor—
isches Museum at Vienna, was photographed there by Macbride as his
type photograph number 3);358.

Additional citations: BRAZIL: Rio de Janeiro: G. Gardner 100
(Macbride photos 34358] (W-—photo); Glaziou 1546 (Ja—lll92).
State undetermined: Ramiz Galvao sn. [Rio Comprido] (P). ARGEN-
ag Misiones: Rodriguez 457 (Herb. Inst. Miguel Lillo 57251]

HILA MEMBRANACEA Turcz.
Additional bibliography: Moldenke, Phytologia 8: 16. 1961.
An isotype of this species, Hostmann 89, deposited in the De-
lessert Herbarium at the Conservatoire et Jardin Botani
Geneva, was photo ed by Macbride as his type nad ceh te mum-

Additional citations: : Bolfvar: Steyermark & Nilsson
193 (N). SURINAM: Hostmann 89 [Macbride photos 2617] (W--photo
of isotype). aa

AEGIPHILA MOLLIS H.B.K.

Additional synonymy: Aegiphila mollis Humb. & Bonpl. ex Steud.,
Nom. Bot., ed. 1, 1: 16. 1821. Aegiphila mtisii Humb. & Bonpl.
ex Steud., Nom. Bot., ed. 1, 1: 16, 1621. Aegiphila salutaris
Humb. & Bonpl. ex Steud., Nom. Bot., ed. 1, 1: 16, 1821. Aegiph-

mollis Bonpl. ex Spreng. in L., Syst. Veg., ed. 16, ue

a
Syst. Veg., ed. 16, 1: 422. 1825. Ae salutaris Humb. &
Kunth +, Ann, Nat, Hist. 2: 49. 1639. Aegiphila mtisii

‘ris Kunth ex Schau. in A. DC., Prodr. 11: 65h. 1917. Aegiphyla
mollis H.B.K. ex Moldenke, Résumé Suppl. 6: 9, in syn. 1

Museum at Ber de as his ¢
was photographed there by Macbride as his type
Photograph number 17588, but is now destroyed; similarly, the type

334 PHYTOLOGIA Vol. 13, no. 5

of aaa: velutina, in the same herbarium, was photographed by
type photograph number 17568 and is now also destroyed.
It is seenivad that the Humboldt XII, with no definite loc
of collection indicated, and also - depoaited in the Berlin herbar-
dum and photographed there by Macbride as his type photograph
number 17586, may have been the actual type of A. mollis, but it
is also now dest: stroyed
Material of A. mollis has been misidentified and distributed
in herbaria as A. glandulifera Moldenke, A. hirsutissima Moldenke,
. martinicensis Jacq., and A. mollis var . intermedia Moldenke.
oa the other hand, the Elias 1458 and Haught 3736, distributed as
A. mollis, are actually var. var. intermedia.

Additional citations: COLOMBIA: Cauca: F, C. Lehmann 6692 [Mac-
bride photos 17568] (W--photo). Magdalena: H. iW. Smith 868 (Mi),
870 (Mi), 1860 (Mi). VENEZUELA: Amazonas: Humboldt 983 ett
Willdenow 2834; Macbride photos 17588] (F--photo);. Fann Pannier
Schwabe s.n. [Ledio y Alto Orinoco] (Ve). Aragua : Burkart Tous
(Ve); L Lasser 905 (W-1879928) . Gudrico: cetatmeniata ta 42h9 (ve—
46833), 616 (Z); Aristeguieta & Hern4ndez 2307 (Ve—37286) . Mér-
ida: J. A. Steyermark bean Seah (Ve—25827). Yaracuy: Aristeguieta &
Pannier 1803 (Ve-—34309). State undetermined: Humboldt X11 (Herb.
Willdenow 2838; Macbride photos 17586] (W--photo).

AEGIPHILA MOLLIS var. INTERMEDIA Moldenke
Additional bibliography: Moldenke, Phytologia 7: we: 1961.
Material of this variety has been misidentified distributed

in herbaria as typical A. mollis H.B.K. On the other pg the H.
H. Smith 870, distributed in some toy as A. mollis var. inter-
media, is actually typical A, mollis. ‘ited

Additional citations: COLOMBIA: Atldntico: Elias 1458 (W—
1594641). Magdalena: sie 3738 (W—-1708553).
AEGIPHILA MONSTROSA Molde

Additional bibliograp’ "Yoldenke Phytologia 7: 490 (1962) and
13: 319. 1966. 7 e =

k refers to this plant as a small tree, 25 feet tall,
with pale olive-green leaves; Contreras found it in ramonal border”
ae a lake; Lundell calls it an ps ADS shrub or small tree, 25

t tall, the trunk 3 inche eter, growing

Co:

Additional citations: GUATEMALA: El eal 1656 Ga, :
ar C. L. Lundell one (Ld, S), 1564 (1d), 1 Ld, oa” Sh6- Izabal:
A. Ste eee ll (Rf). ae TISH HONDURAS: —_ 5078 (14),

5500 (Id), Toe Cay ‘902 (La

1966 Moldenke, Notes on Aegiphila 335

AEGIPHILA MULTIFLORA Rufz & pines Ht

Additional bibliography: K. Schum. in Just, Bot. Jahresber. 28
(1): 496. at Woldenke, Solabergta “hs 148. i9bs; Perea Biota
1: 180--181, 1956; Moldenke, Phytologia 7: 491. 1961

A Kieren of Rutz 187, } riginally deposited in the herbarium
of the Botanisches Museum at Berlin, but now dest: Neersee ™ photo-
graphed there by Macbride as his type photograph number 17587

Additional citations: PERU: Hudmuco: Asplund 12580 pea
Department undetermined: Dombey s.n. [Peru] (W--1 706192) j Rufz 187
[Macbride photos 17587] (W—-photo). BOLIVIA: El Beni: H. H. Rusby
2619 (W—33109) .

AEGIPHILA NOVOFRIBURGENSIS Moldenke
Additional bibliography: Moldenke, He dee san 7: 4gl. hee

isotype of this species, P. Clausen 19, deposited in th
um of the Conservatoi Sot iota sqes at ae was
photographed there by Macbride as his type photograph ‘

tional citations: BRAZIL: Rio de Janeiro: P, Clausen 19)
(Macbride photos 28386] (W-=photo of isotype).

AEGIPHILA OBDUCTA Vell.

Additional bibliography: Angely, Fl. Paran. 7: ll. 19575 Mol-
denke, Phytologia 8: 17. 1961; Reitz & Klein, Sellowia 16: 55 &
108. “Sees Rambo, Pesquis. Bot. 21: 8 & [59].

Rambo (196 Herb. Anchieta 3258, 31676, 50296, 50333,
5035h, Skate. Ss Sion 61h] and 55600 [Reitz 5170] from Santa mta Ca-
tarina, Brazil, all collected by himself except where otherwise
designa: th

ae eseeeejust reaching the coast
(31676)" of Rio Grande do Sul, with a general @istribution “Fron
Onia down to RGS [Rio Grende i ul)".
Additional ehuatsie BRAZIL: Parand: Hatschbach 4986 (A).
atarina: Gaudichaud 173 ( sy; Hatschbach 1927 (W—2027120) ;
Reitz 357 (W—2027679); Reitz & Klein 1864 (N).

Additional biblices aphy: Robledo, Lecce. Bot. 2: 499. 1940; Mol-
denke, Phytologia 7: *92—193. 1961.
Cowan & ero describe this yaaa as a tree, 4 m, tall, with
Pendent branche yellow in primary f
hed and on nets cultivated slopes, at omat 150-600 meters altitude,
looming in April.
Additionay citations: TOBAGO: Cowan & Proctor 1455 (N).

AEGIPHILA ODONTOPHYLLA D.
1 peer hibitcerapie: *ucLienke, Phytologia 7: 493 (1961) and
: 1966,
The H. F. Smith 110 and Stork 4210, distributed as this species
in some herbaria, are re actually ¢ the related A. aculeifera Moldenke.

336 PEI TOLoOG rT. Vol. 13, no. 5

ABGIPHILA PANAMENSIS M
Additional nice. aaoneelg » Phytologia 7: 93--li9h.

Brenes found this plant growing at 1100 meters altitude, flow-
in June, The Feddema 1887, distributed in some herbaria a as
A. panamensis, is actually a species of Schlegelia.
~~ Additional citations: COSTA RICA: Alajuela: Brenes s.n. [Herb.
Inst. Physico-geogr. Nat. Costaric. 148] i eae

AEGIPHILA PANICULATA Moldenke

Additional bibliography: Moldenke, Phytologia 7: 9h. 1961.

Duke refers to this plant as a vine or shrub, with greenish
flowers, blooming in September.

Additional citations: PANAMA: Darien: Duke 4095 (Ca), 4109 (2).
Panam4: Duke 3805 (Ca).

AEGIPHILA P,

Additional ealieeramr ts Pia Fl. Paran. 7: ll. 1957; Mol-
denke, Phytologia 8: 17. 1961.

A imen of Hassler 98, sapere 2 = the herbarium of the
Conservatoire et Jardin Botaniques at Ge was photographed
there by Macbride as his type photograph wrx a 24,618.

cent collectors have found this species gr g in cam
cerrado (widely nape ba scrub) in a at 950 oe alti-
tude, fruiting in January. og: found it flowering in October.

Additional itasones PAZ Goids: i. T. Silva = 57721 (N)-
Paran4: Hatschbach 7687 (Ca), "3523 (Ca), ~ 852 (Ca); Smith, Klein,
& Hatschbach 14729 (Ac) . S#o Paulo: Lanna Sobrinho =a 136 (Herb.
Cent. Pesq. Florest. 1222] (Ac); Mimura 160 tH alhe190 PARA~
GUAY: Hassler 98 [Macbride photos 24618] (W—~photo 3 om
T. Rojas s.n. [Hassler 10852] (A).

ABGIPEILA PARVIFLORA Moldenke
Additional bibliography: Moldenke, Phytologia 7: 495 (1961) and
13: Pb 1966.
ecent collectors describe this plant as a tree, 10 m, tall, oF
. seal shrub, with an irregular crom, the trunk to 0 cm. in di-

eadows, called "papagaio"
in June. Material has been misidentified and distrib:
uted in herbaria under the names A. integrifolia (Jacq.) Jacks.
Aegyphylla sp,
A specimen of Spruce 589, an isotype of the species, deposited
in the herbarium of the @ Conservatoire et Jardin Botaniques at Gene~
va, was photographed there by Macbride as his type photograph nua”

Additional citations: VENEZUELA: Apure: Borsotti 3 (Ve), 2
(Ve--39518); Ll. Williams 131,01 (w-~1802208 . BRAZIL: Minas Ger~
ais: Heringer er 5802 (B). | (B). Paré: | Spruce 589 [Macbride photos 28387]
(W—photo of isotype).

1966 Moldenke, Notes on Aegiphila 337

AEGIPHILA PENDULA Moldenk
Additional <a geeen ans Moldenke, Phytologia 7: 96. 1961;
Moldenke, Résumé S Supel. 12: 3. 1965.

Additional citations: VENEZUELA: Amazonas: Ewel 199 (2).

AEGIPHILA PERUVIANA Turez,.
bs Ege synonymy: Aegiphila peruana Turcz. ex Soukup, Biota
a e 1962.

Additional airy fT ieee Phytologia 7: )96—197.
1961; beak, Biota h: 81. 1962.

An isotype, Spruce 275, deposited in the herbarium of the
Dsl Cire et Jardin Botaniques in Geneva, was photographed
there by Macbride as his type photograph number 2/619. Soukup
collected the species in anthesis in August. Wi ows
scribes the plant e a shrub, 2.5 meters tall, with yellow fruit
in March, growing in secondary forests at 500° meters altitude.

Additional citations: PERU: San Martfn: Spruce 275 [Macbride
photos 24619] (W--photo of isotype); W ki 5413 [wood vouch~
er 15513] (W—2)13510), 7136 (W—2h53190) . SE undeter=
mined: Soukup 4371 [villarica] (W--2199967)

AECGIPHILA PLATYPHYLLA Briq.
Additional synonymy: Aegiphilla platyphylla Briq. ex Moldenke,
es Suppl. 13: 6, in syn. 1966.
Additional bibliography: Moldenke, Phytologia 7: 97. 1961;
Moldenke, Résumé Suppl. 13: 6.
type specimen, Hassler 8056, “deposited in the herbarium of
ardin B

IPHILA PURPURASCENS M
Additional ter paaalnnceny ete Biol. Abstr. 27: 2026. 1953;
Moldenke, Phytologia 7: 198. 1961.

AEGIPHILA QUINDUENSIS (H.B.K.) Moldenke
13: ‘gale cpa ire Phytologia 7: 498 (1961) and

Recent collectors call this a treelet, to 5m. tall, not much

» growing at 900 meters altitude, fruiting from September
‘o November. Material has been mis identified and distributed in
ee as A, membranac
Julian Ri tee. in etter to me dated November 8,

1963, “says: "I was not able to S tise my specimen (no, 91800) with
seule in A. quinduensis, as there was some question in my mind

ss to whether it might not be A. martinicensis. We have no eet
ial of the latter mews for comparison. The difference Sones

oe °

teg to separa a ¥ an pve with 'blades

338 PHYTOLOGIA Vol. 13, no. 5

etc.' from A. quinduensis with 'blades thin-chartaceous or submem-
branous' is not altogether easy to apply in this present case.
Also, some of the calyx-rims in my specimen appear subtruncate or
repand and could be placed in the section of your key on p. 269."
Additional citations: VENEZUELA: Aragua: Arist eta 51
(Ve); J. Garcia 53 (Ve—30983); Lasser 2058 (Ve—25129); H. Pit-
tier 1547% (Ve—26967), 15481 (ver=26896), 35048 (Wo 26959)5
pater & or & Nakichenevick © 15550 (Ve~~-29312); Je oraark 2
5aso7 (B, ¥_—2430130) . - iirande 7: Aristeguieta 485k Bs fe ghee eg Von-
agas: J. A. Steyermark 620h6 r= hy

AEGIPHILA RACEMOSA Vi
Additional ian har Moldenke, Phytologia 7: 498-99.

Archer and Silva both describe this plant as a vine. Kecent
collectors have found it on terrafirma, blooming ae January and
March, The flowers are described as "yellow® on Archer 8278 and
A. a 93, and "green" on Archer 8120.

tional citations: VENEZUELA: Bolfvar: Lasser 1983 (Ve); J+
A. pe 8841 (N). BRAZIL: Amazonas: G. P. = Fe Cooper : SNe
[Lower Rio Madeira, March 20--2h, 1945] (W—-213907h), son. [Ita-
pumua] (N). Par&: Archer 8120 (N), 81,1 (W—2),39075), 8278 (N)5
Murca Pires 8227 (N); A. Silva 93 (Ni, [, W--2),39609). MOUNTED ILLUS-
TRATIONS: Moldenke, Phytologia 2: kl, fig. h. 1948 (N).

AEGIPHILA RIEDELIANA Sc
Addi tional. Grhiacesacter? Moldenke, Phytologia 7: 499. 1961
Rambo, Pesquis. Bot. 21: 8 & [59]. 1965.

A cotype, J. E, Pohl 4392, deposited in the herbarium of the
Naturhistorisches Museum in V Vienna, was Ea there by
Macbride as his type tograph number

Rambo (1965) cites Herb. inchieta EBL06 [Reitz 8383] from San-
ta Catarina and 0320 and 59493 [Camargo 681] from Rio Grande do
Sul, the second being his own collection. He describes the plant
as a "Small tree, up to 5 m high", growing in "Half-high wod,
secondary growth", and found "So far, northeastern Highlands —
in Rio Grande do Sul, with a general distribution son, Sen Bahia
RGS [Rio Grande do Sul]",

Additional citations: BRAZIL: Bahia: J. E. Pohl 4392 [Macbride
photos 34309] (W-—photo of cotype).

AEGIPHILA RORAIMENSIS Molde
Additional bibliography: Tohono, Phytologia th 500. 1961-

Steyermark
leaves membranous, pot cee ve both 5 tens > and the pubescence
on the stems, peduncles, and calyx fulvous tawny-brom, growing

1966 Moldenke, Notes on Aegiphila 339
A. Steyermark 60812 (Ve—35968), 93834 (Lw), 94085 (Z).

AEGIPHILA SALTICOLA Moldenke
Additional bibliography: Moldenke, Phytologia 7: is 94 aoe

Additional citations: BRAZIL: So Luiz Island: 11856
(Mi, W—1830837
AEGIPHILA SELLOWIANA
rage SOA ceniotonsaun ely, Fl. Paran. 7: 11. 1997 Reitz,
66, & 8h. 19585 Rambo, Pesquis.

: Lo,
136 oldenke, ’Phytologia 73 501. 1961; Reitz & Kein, ,Sellowis

sr
6: la, 64, & 103. 1964; Rambo, Pesquis. Bot. 21: 8-9 & [59].

fv collectors have found this plant flowering in May and
ting in May and August. The leaves on Belém 1459 are up to

and 16 cm, wide!
Reitz (1959) records the vernacular names "gaileira",
"gaioleira", “pau-de=gaiola", "pau gaiola", “tamanqueira", and
eiro", Angeli records _ — molulo". Castellanos
The Sehnem 2149, dist tributed as Ve “sellowiana, is actually A.
brachiata Vel. Vell., while Hatschbach 26h and Machado de Campos ua
are A. verticillata Vell. ~——S
Rambo (1965) cites Herb. Anchieta 31530 and 58429 from Santa
aha and 1216, 3762, a= [Theissen s.n.], 11822 en Ss.
n.], 27529 (Heng ; Mes, 2625. [Henz s.n.], 35671 [Henz s.n.],
38582, 36835, 38896, a ae 3 39062, hO7Th, L85k0 [Sehnem 3739],
Wé2a1, 51009 (Sehnen $606], £2931, = [Pivetta 647], 6178h
(Camargo 2864], and seats [camargo 17ihy, nese Rio Grande do Sul,

"Edge of the f ndary growth, Campos woodlots" in the

te Read heb Rhee Gees of the Highlands only" in Rio

— do Sul, with a general distribution of "Fron Amazonia to
RGS [Rio Grande do Sul]",

Additional citations: BRAZIL: Bahia: & Magahlaes 1091
(Ac). Espirito Santo: Belém 1459 (Ac). ae ire Ae —
hos 2348) [Herb. Cent. Pesq. Florest. 1714) (Pf); Lanna Sobrinho
is [Herb, Cent. Pesq. rt iy 1159] (Ac). Parand: Hatschbach
2615 (W#—2057308) 7709 (Ca). Rio de Janeiro: Angeli 100 [Herb.
Cent, Pesq. Florest. 333] (Ac), 101 [Herb. Cent. Pesq. , Florest.
335] (Ac); 0. M. Barth I.71 (Herb, Inst. 0. Cruz 98] (W—2342877).

© Grande do Sul: Ram Rambo 52931 (B); Setmem 3739 (B), 5606 (B, B).
Santa Catarina: Kle Klein 132 (W—2120120); Reitz 3379 (W——-20278)6) 5
Reitz & Klein 7259 (N), 8 8257 (MN), 17357 (ac).

ABGIPHILA SESSILIFLORA Moldenke

Additional biblio Moldenke, Phytologia 7: 502. 1961; Mol-
fenke, RSsuné Suppl. capt: 2 & 3. 1965; Moldenke, Phytologia 13: 328.

3h0 PHYTOLOGIA Vol. 13, no. 5

966.

Recent collectors describe this plant as a tree, 6—-10 m. tall,
with an irregular crown and green fruit, growing at 1200 to “1,00
m,. altitude, and flowering in er and August. ave refers to
it as "rare in this locality" in Antioquia. eeeeee has —
misidentified and distributed in herbaria as A. integrifo
(Jacq.) Jacks. and as Lucuma sp.

Additional citations: 3: COLOMBIA: Antioquia: Archer 589 (W—
en . Cundinamarca: Fern4ndez & Mora 1,02 (W—21L1)385). VEN-

: Trujillo: Lasser 1167 1167 (W—1908975).

ARGIPHILA SESSILIFLORA var. CUATRECASASI Moidenke
Additional bibliography: vie ldenke, Phytologia 8: 385.
Hocking, Excerpt. Bot. A.6: 455. 1963; Moldenke, Biol. inners ‘as

f oo COLOMBIA: Magdalena: Cuatrecasas & Castafieda 25h0k
type).

AEGIPHILA SKUTCHII Moldenk
Additional Aikinamephes Molden » Fiytologis 7s Red 1961.
Breedlove describes this nie as as 10 fee white
flowers, growing on a steep moist slope at €100 feet Merrie
fl in August.
Additional citations: MEXICO: Chiapas: Breedlove 11978 (Z)3
Matuda 190 (Rf) ie CGE

AEGIPHILA SMITHII Moldenk
Additional nied topeacies Moldenke, Phytologia 7: 502--503.

Additional citations: PERU: Loreto: Asplund 14048 (W—222h811)5
Mexia 6499 (W--1691256). San Martin: KI ae 3854, (W—158301)

AEGIPHILA SPICATA (Rusby) Moldenke
Additional bibliography: K. Schum, in Just, Bot. reese ie"
28 (1): 295. 1902; Moldenke, Phytologia 7: 503. 1961

IPHILA S
Additional i bbls ae: se sie Be ieee 7: ene sohtes
The type this species, J. 1022, deposited in

he

graphed there by meornaoe as his type photograph number
Additional cita BRAZIL: Minas Gerais: J. B. Pobl 1022

[Macbride photos 31,308) (W——photo of type).

AEGIPHILA SPRUCEANA Moldenke

Additional bibliography: Moldenke, Phytologia 7: 503-—50l.

An isotype of this rece: Spruce 2296, deposited in the ails

barium of the Conservatoire et Jardin Botaniques at Geneva
i pag there by err as his type photograph number

1966 Moldenke, Notes on Aegiphila 341

dditional citations: BRAZIL: Amazonas: Spruce 2296 [Macbride
photos 28388] (W—photo of isotype).

AEGIPHILA STEYERMARKII Moldenke

Additional as senaraeel Moldenke, Biol. Abstr. 35: 2177. 1960;
eee, Phytologia 7: 50). 1961; Hocking, Excerpt. Bot. A.l: 593.
962

Steyermark & Aristeguieta describe this plant as a shrub, 1.5
m. tall, the leaves subcoriaceous, deep dull-green ahr dull-
green beneath, the nervation raised, the sereabe oT green, the

corolla creamy-white, the filaments whitish, and the sta Bi buff-
creamy, growing o n forested slopes at 150 ws Ae altitude, flower-
ing in March,

Additional citations: VENEZUELA: Bolfvar: S Steyermark & Aristeg-
uieta 79 (Z).

AEGIPHILA STEYE I var, MACROPHYLLA Moldenke
Additional bibliography: Moldenke, Phytologia 7: 50l-~-505.
1961; Hocking, Excerpt. Bot. A.l: 593. 1962.

AEGIPHILA TRUNCATA Moldenke

Additional bibliography: Moldenke, Phytologia 8: 17. 1961;
Moldenke, Biol. Abstr. 37: 215. 1962.

Additional citations: COLOMBIA: Antioquia: Metcalf & Cuatre-
gasas 30072 (W—1833329).

ARGIPHTLA Precip rh Moldenke

Additi ibliography: Moldenke, Biol. Abstr. 30: 3551. 1956;
Woldenke, phytic 8: 17. 1961.
AEGIPHILA V.

Addi tional “rhoppes sat Jos Moldenke, Phytologia 8: 17 (1961) and
13: 319. 1966.

Matuda describes this plant as a tree, 20 meters tall.
Cazalet & Pennington 5090, distributed ag A, valerii, 1s aie

A. alba Moldenke.
Additional citations: MEXICO: Chiapas: Matuda 1876 (Rf).

PHILA VENEZUELENSIS Moldenke
Additional nal biblioerapiy: Moldenke, Biol. Abstr. 33: 1215. 19593
Inldenke, Phytologia 8: 18. 1961; Hocking, Excerpt. Bot. A.5: hh.

Steyermark and his associates describe this plant as a shrub or
hi ay shrub, 2 feet tall, the Tr prt psa or membran-
ous, green or -
ig flowers pendent ie gon ore along petit a at iihodes
el oT Sol peters, flowe owering in May.
tations: VENEZUELA: Bolfvar: J. A. Steyermark 60263
ease), (S5ibe (Ve); 3 Steyermark, Dunsterville, & Dunsterville

3h2 PHYTOLOGIA Vol. 13, no. 5

IPHILA VENEZUELENSIS var. SERRATA M

Additional bibliography: “Moldenke, ey = Abstr 33: 1215. 1959;
Moldenke, Phytologia 8: 18. 1961; Hocking, Excerpt. Bot. A.5: uh.
1962.

AEGIPHILA VERRUCOSA Schau.
Additional bibliography: Moldenke, signage — ie 19. a.

Recent collectors sabe ike this plan a 10 meter
tall, the leaves subcoriaceous, Sentient above, "Ett area <n
neath, the flower-buds and calyx green, the corolla creany,

at 1700 to 1800 meters altitude , flowering in June and
October. An isotype, Moritz 897, deposited in the herbarium 0
the Conservatoire et Jardin Botaniques at Geneva, was photographed
there by Macbride as his type photograph number 26389.

Additional citations: VENEZUELA: Aragua: Aristeguieta & Medina
2964 (Ve--2095), 2972 (Ve——l210k) ; Moritz 897 oot net
28389] (W—photo of isotype) e Allen 90886 Fed

eral District: Lasser 978 (We RE} ot 979 (¥ oe EO a
Be Maguire 39137 (N).

AEGIPHILA VERTICILIATA Vell.
Additional bibliography: Angely, Fl. Paran. 7: 11. 19573 Mol~-
denke, woth dbentacee 8: 19-20. 1961.
collected this species in a savanna of cerado veget
tina, The pooh type specimen of A. tomentosa Cham., Sellow “5122,
deposited in the herbarium of the Botanisches Museum at , Berlin,
was photographed there by Macbride as his type photograph number
17589, but is now destroyed. A specimen of Casaretto © 262h, in the
pecs gh Herbarium at the Conservatoire et Jardin botaniques at
eneva, was photographed by him as his type photograph number
261s and may be regarded as the type of A. lanata Casar.

Recent collectors have found A, verticillata growing on

Ss and sandstone ing at stream mar , and in bogs, for
ests, ‘ied Deas fields, at 950 meters altitude, flowering in Jan-
uary. Material has been misidentified and distributed in herbar-
ia as A, sellowiana Cham,

Additional citations: BRAZIL: Minas Gerais: Casaretto oe
[Macbride photos 2615] (W--photo). Paran&: Hatsc
2057321); Hatschbach & Pereira 7668 [Herb. * Sabah Tod ach LODE] ay
Reitz & Klein 17562 (Ac), 17876 (Ac); Smith, Klein, & Hatschbach
14508 (Ac). Slo Paulo: Machado de Campos 12 Dos id (N, W—2369519) 5
Mimura 23 (W—2)01,883), Lo (i= (W—-21,01,900) ; Sellow 5122 [Macbride
photos 17589] (W--photo).

AEGIPHILA VILLOSA eg ) Gmel.

pieyei one Aone nym * Regiphite wee vilosa (Aubl.) Gmel. ex Molden~
ke, Résumé scone n syn.

” additional biblio cena: woldenke, Phytologia 8: 20. 1961; Mol-
denke, Résumé Suppl. 7: 7. 1963.

1966 Moldenke, Notes on Aegiphila 343

Recent sn have found this plant growing in full expo-
sure to the on disturbed areas, flowering in July, and de-
scribe it as a spies tree, with white aoe

Additional citations: BRAZIL: Amap4: W We. Ae “Egler Ea vagalee
Mus. Par, Emil. Goeldi 24634] (N); Frées & & Black 27ly
2248464); Murca Pires, Rodrigues, & Irvine 50327 - he

AEGIPHILA VITELLINIFLORA Klotzsch
tional synonymy: Aegiphil a vittelliniflora Klotzsch ex
Rambo, Pesquis. Bot. 21: 58, sphalm. 1965.

Additional Teg crecinay Hayek in Engl., Bot. Jahrb. 42: 172.
1908; Mo ¢, Phytologia 8: 20--21. 1961; Rambo, Pesquis. Bot.
a to 1365) Moldenke, Résumé Suppl. 13: > 1966} Moldenke, Phy-
Bat i

records the vernacular name "marmelinho" for this

Bt ae ional citations: BRAZIL: Rio de Janeiro: liar s.n.
[2-12-1922] (Ja—11)10h, Ja); Sampaio 8199 (Ja——lh835).
AEGIPHILA VITELLINIFLORA var. EGLERI M

Bibliography: Moldenke, Phytologia a “Ses. 1962; Moldenke,
ae Abstr. 42: 1517. 1963; Hocking, Excerpt. Bot. A.6: 455.

Citations: BRAZIL: Amap4: Irwin, Egler, & Murga Pires 47162
(N-type) . ee? ne eee eee
AEGIPHILA WIGANDIOIDES Lundell

Additional bibliography: Moldenke, Phytologia ): 38. 1953;
Moldenke, Biol. Abstr. 37: 1062. 1962,

: a tional citations: MEXICO: Chiapas: Matuda 4253 (Ld—iso-
ype

ADDITIONAL NOTES ON THE GENUS LIPPIA. I
Harold N. Moldenke

Houst.
Additional & emended bibliography: Robins. es bie
Gray, New Man, Bot., ed. 7, 688-690 & 906, fie. oh & 32. 1908
Loes., Verh, Bot, Ver. Brand. 53: 43 & 75-78 {Abhandl. "2G 2h3).
. pg

L036 eet af 37: 7 1062, 1962; Hocking, Excerpt. Bot. A.5: bh

34h PEYTOL0G I-A Vol. 13, no. 5

& 45. 1962; Dyer, Verdoorn, Codd, & Letty, Wild Fils. Transv. 281,
pl. a tig. Sais Rickett, New Field Book Am, Wild Fis. 280.
1963; T cena 28: 170. 1963; scone — ma.

Gooding. Loveless, & Proctor, ser! 353, 36 32, & ibe 19655

Gard. & 7 965;
19): B.72 (1 46 fon} 53 : oetiae (ges), “and 46 (22):
le do): 2: Pa Be 3 a Cs)" eats, Bot. 21: 2h—-29, 58, & [59].
Ag

) « 19655
a Biol. Abstr. 46: 32h6 & 3616 (1965) and 6 (23): B.72&
B.9h. 19653 Moldenke, Résumé Suppl. 12: 1—7 & 9--12. 19655 Mol-
el Phytologia 12: 6—-71, 73--120, 130-181, Sg oe ee
pe

33 1966
A. Jere, T axon ie, 161. 1366; Anon., Biol. peta og ‘a7 (8): ‘Se 9.

In the main body of this work, to which this is the first
plement, 16,427 herbarium n specimens and 854 mounted seek eiaidee
2 other illustrations ited.

anonymous editor of tie pinaica Abstracts" [6 (23): B.72
& 39h) places the gems Lippia + in the family Phytolaccaceae for
some reason entirely unknown to me!

Several additional stvroriatiehe for the names of herbaria in
which cited material is deposited used in the following notes

will be explained in my Callicarpa paper which will appear short-
~o on these pages.

LIPPIA ABYSSINICA (Otto & Dietr.) cae
Additional bibliography: recat » Abstr. 37: 1062. 19625
re ae Résumé Bd ia = ie 0. $9665 aasas. Phytologia
39--hh, 176, 30. 298, 305, 307, 350—352, 431,
bit, “igs, & 459 (965), tor ub3 (1966),’ and 13 171. 19665 Molden
ke, Résumé S Suppl. 13: 966.
sonpla “this plant may attain a height of 3 me-
ters; his 11.635 is described as having had white corollas, flow-
ering in March, He records the vernacular name "umuzirarorwe".
The A. Peter 11351, distributed as L. adotnsis yng is actu-
ally L L. schliebeni Moldenke.
Additional citations: ERITREA: Pappi 5602 (W—-1969226). ETHI-
OPIA: qmper 376 (W--80646). RUANDA: “Troupin 11.635 (w#—
237511 ANGOLA: Province undetermined: A. Rocha 60 a 60 [Malange

INICA var. PUBESCENS (Moldenke) molienks si
Additional bibliography: Moldenke, Biol. Abstr. 37: 1062. ob
a, Phytologia 12: h3—Wh. 1968; nit Résumé Suppl. 47°

LIPPIA ACUTIDENS Mart. & Schau.
Additional bibliography: Moldenke, Phytologia 12: 2h & bl--li5-

1966 Moldenke, Notes on Lippia 345

1965.
Additional citations: BRAZIL: Minas Gerais: Martius s.n. [Yha;
Macbride photos 20319] (W--photo of type).

LIPPIA AFFINIS Schau
Additional bibliography: Moldenke, Phytologia 12: 23, 26
ne 5 162, 164, 170, 189, 196, 219, 283, 332, 339, 346, & ie.

ello Barreto aren cited below was originally iden-
tified and distributed as L. origanoides H.B.K.
a citations: BRAZIL: Minas Gerais: Mello Barreto 1080)
J

LIPPIA ALBA (Mill.) N. EB. Br.
Additional & emended synonymy: Lippia panamensis Turcz., Bull.
Soc, Nat. Mosc. 36 (2): 201--202. 1863. lLippia havannensis Turez.,

Bull. Soc. Nat. Mosc. 36 (2): 202203. 1863. Lippia melissae-
odora C, A, Mey. ex Moldenke, Résumé Suppl. 12: 10, in syn. 1965.
Lippia melissacodora C. A. Mey. ex Moldenke, Résumé Suppl. 12: 10,

syn, 1965.

Additional bibliography: Loes., Verh. Bot. Ver. Brand. 53: 75
[Abhand.. a) 1912; H. B. Davis, Life & Works Pringle 52. 1936;
Moldenke, Biol. evi 36: 4036. 1961; Angely, Bibl. og Paran.

S

loldens, ey tol ogia 12: 2h, h7-—7, 76, » hy CN 19,

oe Sale ano, 196, 217 219 22 ——

20, 260, 2627 26h’ 38 310; 358; 3k; 450, 19 (1368), 122 487
& 505 (1366), "and 13: 17, 19, 21, 26, 1966.

The name, "Li ia melissaeodora, tains “to be based on the Herb.
Hort. Bot. Petrop. s.n. deposited in the Leningrad herbarium, a
duplicate of which is in the United States National Herbarium at
Washington,

The Eitens record the species as escaped from cultivation in
razil, where it is used to make a tea in the treatment

are mS Tastenes ene Torr
Turezaninow (1863) eae his L. havannensis as oni
"L, caule tereti fruticoso miltirameo, ramisque pube adpres

heal in Gunhdian Cetenutéa crenatis, longitudine petiolorum su-
Pra rugosis, utrinque subtus densius tomentosis; pedunculis

llis solitariis folia aequantibus; ns, pg hemisphaericis;
bracteis lanceolatis acutiusculis glandulosis margine ciliatis,
tubum corollae tegentibus; calycibus pubescentibus carina dense

346 PHYTOLOGIA Vol. 13, no. 5

ciliatis. Flores e schedula albi." He describes his L. panam
sis as follows: "L. ramis teretiusculis aut superne * Steurag tats,

glabriusculis, subtus pubescentibus; pedunculis in axillis soli-
tariis petiolum duplo superantibus foliis brevioribus; pa dee to
late rhombeis in acumen breve producti, calycibus que dors

guste alatis, totis pilosis; tubo corollae calycem duplo eeelatas
coccis maturitate secedentibus. Habitus fere L. geminatae

folia duplo minora, in pagina superiore rugosa . glabra OF es.
tenuiter sed manifeste alati." He places it in Section Dipte

(1965) cites Schauer's work in eg Fl. Bras
"1951" instead of 1851, doubtless through an hographic ‘or =
phic error. He cites Herb, Anchieta 43h, 3a, ae {Ea-

pogra)

rich s.n.], 11545 [Buck ey 261,30 [Buck “sen.), 27560 60 [Henz s.
n.J, 28261, 29629 [Emrich s.n.], 35551 [Henz s.n.], > 30012, 39489,
143206, 7332 [Spies s.n.], 47815 [Sehnem 3823], 969k, i, 50,97 7 [Seh-
nem nen 3872], 59158 [Pivetta 970], and 61654 [Camargo go 2785) from Rio
Grande do Sul, a all collected by hims himself, many from cultivated ma~
terial, except where otherwise designated. He describes pe
plant as SS "Shrub = long, os more or less recumbent

ranches", growing "apparently on sandy, Mheicgrss AMPOS sees
often Sarkieated for medicinal saan ithe apparently ear
the sea coast" in Rio Grande do Sul, with a saviors distribution

"From southern USA to central Argentina and Uruguay.

Additional citations: TEXAS: Hidalgo Co.: M. L. sake 34, (Ud).
MEXICO: San Luis Potosf: Crutchfield & Johnston 5135 . Tamau-
lipas: R. M. King 035 (Mi). WINDWARD ISLANDS: eee a Re As
Howard 718 ee VENEZUELA: Apuré: Humboldt & Bonpland 5 sD.

[Macbride photos 39485] (W--photo). ECUADOR: Guayas: F erlind &
Wibom 138 (Mi). BRAZIL: Ceard&: Drouet 2561 eas fers PARA
ee Hassler 7397 (Mi) 5 T. Rojas 228 (A). CULTIVATED: Brazil:

n & Se E (W—2)],5201). Russia: Herb. Hort. Bote Petrop.

IEPA AL vas Rare aac aie 965; Mol
bliograp Mohiathes Phytologia 12: 71. 19655 ™0*

denke, Biol. Abstr, 36: 136. 2 1961. #

LIPPIA

ALNIFOLIA
. ee opep races Moldenke, Phytologia 12: 2h, TL, & 73~

dditional citations: BRAZIL: Minas Gerais: Martius s-Ne (ac~
oils photos 20321] (W--photo of cotype).

LIPPIA AMERICANA L,.

1966 Moldenke, Notes on Lippia 347

Additional bibliography: Moldenke, nape ge ogia 12:-20,;23, 63,
mé Suppl. 13:

Additional Ly epeet COLOMBIA ena: H.
ae erga” do Norte: Schlim ‘om [Macbride ue Fotos ate
W--phot

LIPPIA AMERICANA f. eee aitat (Benth.) Moldenke

Additional synonymy: Lippia hyptodes Benth. ex Kuntze, Rev.
Gen. Pl. 3 (2): see seme Sm OF

Additional bibliography: “Yoldenke, pubes ey 12: 63, 77, &
79-81. 1965; ac preci Résumé Suppl. 13: 7

LIPPIA ANGUSTIFOLIA Cham.

Additional bibliography: Moldenke, Résumé Suppl. 12: 10. 19655
eae Pesquis. Bot. 21: 25 & [59]. 1965; Moldenke ds boas

: ah, 82-85, 102, 205, 259, 293, 294, 347, & Wel, (1965)
ae a &15. 1966.
should Pe noted here that Le. angustifolia Chod. is a syno-

nym oe L. tri de

Rambo (1 1968) cites Herb. Anchieta 25780 from Rio Grande do Sul

and 36858 [Hatschbach 661 661] from Paran4, the former collected by
himself. He describes the plant as a "Slender, woody >
up to 1 m high", growing in "Swampy thickets" on the

branched,
"Southwestern plains only" in Rio Grande do Sul, with a general
ye agit of "Minas Gerais to RGS [Rio Grande do Sul]; Para-
guay to U ay."

Laditionst citations: BRAZIL: Minas Gerais: Sellow s.n. [Bra-
Silia merid.; Macbride photos 1795] (W--photo of of cotype). PARA~
QUAY: Hassler 7971 (A).

LIPPIA ARECHAVALETAE Moldenk
Additional Phe eee ‘Bano , Pesquis, Bot. 21: 25 & [59].

wig pelceake hytologia 12: 86—-87 & 366 (1965) and 43%: 3s Yi ee:
19653 cites Herb, Anchieta 2751, 3920, & 28181
eat oe first as "var. micro, 3720, atl ecl-

lected by himself. He describes the plant as a "Slender, woody
1 m high” on "V dry Cam

area
general distribution "From Paran4 through 1aa-ecaaaeaad Mesopotamia to
Western RGS [Rio Grande do Sul] and Uruguay."

TA ARECHAVALETAE var. MICROPHYLLA Molde
Additional bibliography: Rambo, Pesquis. ge 21: 25. 1965; Mol-
denke, Phytologia 12: 87. 1965.

LIPPIA ASPERRIMA Cham.
Additional ae oenety Pesquis. Bot. 21: 25-26 & [59].
1965; Moldenke logia 12: 2: 2h, 63, 88--9, 135, 136, 232, 290,

347, 429, & 43f, (1965) and 13: 11, 12, 28, & 29.
The Sasson os sd Be by me in * Phytologia 12: 90 (1965) is actu-

348 PHYTOLOGIA Vol. 13, no. 5

ally L. t turnerifolia Cham. cS ath 1as9 in.
Rambo (1965) cites Herb. Anchieta 3035, 3040, E
26114, ei {rate S.n.J, QoOu1, 5201 1’ (Hatschbach 1006],
53011, 53437, 59123 [Pivetta 961], 60323, 63153 [Spies s.n.], &
63196 [Spies s.n. KE from Rio Grande do Sul, “all collected by him-
self | except where otherwise noted, He describes the plant as a
"Small, woody shrub 10--50 cm high", growing on "Very dry, open,
ea

]; Paraguay to northe aoe
Agi tioaat citations: BRAZIL: Parana: Dombrowski 68 (Ac);
Dombrowski, Saito, & Pereira 757/550/1ny (Ac), a aaTor 272 pees
ARGENTINA: Salta: T, Meyer 3452 (Herb. Inst. Miguel Lillo 3 69]
(G).
LIPPIA ASPERRIMA var. LONGIPEDUNCULATA Moldenk

Additional bibliography: Moldenke, phoke logis 12: 91--93 & 29h
(1965) and 13: 29. 1966.

PIA SAE Briq.
eras bibliography: Moidenke, Phytologia 12: 2h, 63, 94-—
97

; saute citations: PARAGUAY: Balansa 1033 [Macbride photos
24645] (W--photo of type), 1033a (N); Hass’ Hassler 8653 (A).
LIPPIA BAUMII Gttrk

Additional bibliography: Moldenke, Phytologia 12: 48, 56, 97—
98, 225, & 230. 1965.

LIPPIA BELLATULA Moldenke

Additional bibliography: Hocking, t. Bot. A.9: 367+ 19695
Moldenke, Phytologia 12: 2h : "98-99. "Tee, Moldenke, Biol. Abstr.
6: 32h6. 1965.

tional citations: BRAZIL: Bahia: Martius 1983 [Macbride
photos 20322] (W-~photo of type).
LIPPIA BOLIVIANA

Rusby
ai. bibliography: Moldenke, Phytologia 12: 99—-LOL & 217+
1965.

Additional citations: BOLIVIA: Cochabamba: Byerdam 24885 (G)-

BOTHRIOURA Bri sik:
cee ibiepraphys Moldenke, Phytologia 12: 64 & 101—
19

citations: PARAGUAY: Hassler 771 [Macbride photos
ouéhe} aotasanees W—photo of type). —S™”
LIPPIA BRACTEATA

Additional abo itis Moldenke, Phytologia 12: 103, 137, 100s
& 265. 1965.

1966 Moldenke, Notes on Lippia 349

LIPPIA BRACTEOSA (Mart. & Gal.) Moldenke

Additional bibliography: Greermm., Field Mus. Publ. Bot. 2: 31.
1912; H. B. Davis, Life & Works Pringle 1. 1936 ; Moldenke, Phy-

tologia 12: 102-—~10 105, 2 tt & 268 (1965) and 12: 502. 1966;
Moldenke, am Suppl. 13:

flowers this aries - described as "yellow" on Breed~

dove 1419) and on Sacitave & Raven 13178. These collectors re-_
a ae is 3—l feet tall, flowering in October and
December ing on steep slopes with Quercus, at 2500--5100
feet SH orgy “The Rzedowski 17649, distributed as L. bracteosa,
is actually L. inopinata Moldenke.

Additional | citations: MEXICO: Chiapas: Breedlove 119k (Z);
Breedlove & Raven 13178 (Ac). Michoacdn: Diguet s.1 s.n. [Barranca
de Yungapeo, Octobre 1907] (Mi).

bea oe etapa J. G. Baker
dditional bibliography: re Phytologia 12: 42, 105-~106,
2a, 232, 307, 352, & 431. 1965

LIPPIA Sond.
sggittttional repcppeaetich Moldenke, Phytologia 12: 107 & 232.

LIPPIA CALLICARPAEFOLIA H

Additional bibli pene ra set Verh. a Ver. Brand. 53: 78
[Abhandl., 23]. 1912; H. B. Davis, Life & Works Pringle 143 & 30.
1936; Moldenke, Résuné § Suppl. 12: "2 & 10. 1965; Moldenke, ea
60, les - 98, ew aR 137, ey ee 268,

= city : Simpson describe this plant as a shrubby climber and
found i on the bottom of a barranca, fruiting in gro
The soetins has also been collected in fruit in January, and
barium material has been misidentified and distributed in iebtarcia
ascea sp. The Mercado s.n. [Mayo ae distributed as L.

the Harvard University herbaria, but have not been verified :
y ut have no

MEXICO: México: Seler & Seler [NAW (G). Morelos: Seler & Seler 376

(G). Vera cruz: “Ervendber rg 278 erg 278 (G); Kerber s.n. [Prope Cordoba,

1X.821) (A). The Herd. Colo. A. & M. A. Co “A. Coll. s SNe, eet Xo-

a are re actually L. substrigosa Turcz., while Hinton ae is
umbellata Cav.
=: Ubellata
ten ttonal citations: MEXICO: México: Gilly & Simpson 23 pt
Faray 548 (Ip), 3340 (Ip). GUATEMALA: Amatitlan: Kellerman 6379
Tones 3056). Sacatepéquez: S. S. White 5271 (Ip).

350 PHYTOLOGIA Vol. 13, no. 5

LIPPIA CANDICANS Hayek
Additional bibliography: Moldenke, Phytologia 12: 113-114.
1965; Moldenke, Résumé Suppl. 12: hl. 1965.
This species has been found ae in chapada of tall trees
on lh of a low plateau at an altitude of 300 meters.
tional citations: BRAZIL: Goids: E. Y. Dawson

(2
Praaye G. Gardner 3942 {Macbride photos 31,329) (W—photo of type)
3: Eiten & Eiten 4162a (W—2))5195) .

TA ent east Benth.

Additional bibliography: Moldenke, plenary 12: 63, 104, DL
116, 137, 189, 301, & a eee * and ag 3h & 162. 1966.

Breedlove 12098 is yellow" fife This col-
lector encountered the ae th on Saw and on gradual heavily
wooded slopes.

In the Harvard University herbaria there are also the following
specimens filed under this species but not as yet verified by me:
HOND! anta : Molina (G

Island: C. L. Smith s.n. 71893) (G). Department undetermined: C-
Wright s.n. [1853-56] (G). COSTA RICA: Alajuela: Hunnewell 16720
(G). Heredia: P. H, Allen 565 (A).
Additional citations: MEXICO: Chiapas: Breedlove 12098 (Rf),
13921 (Ac). Se eee

LIPPIA CARVIODORA Meikle
ERT Ha bibliography: Moldenke, Phytologia 12: 116—-118 &

IA CHACENSIS Moldenke
Epi naan bibliography: Moldenke, Phytologia 12: 63 & 119.

LIPPIA CHEVALIERII Moldenk
Additional bibLiograpiys Moldenke, Phytologia 12: 2, 43, 19—
120, & 130—131. 1965.

LIPPIA CHIAPASENSIS Loes.

Additional synonymy: Lippia chiapensis Loes. ex P. C. Standle,

Contrib, «Ue Ss. Kat. Herb. 23: 126, sphalm. 192h.

bibliography: Loes e9 Verh. we Vere Br —
53: 78-19 [Abhandl . 2h3—2hh]. 1912; P. C. Standl., Contrib. J. 8
Nat. Herb. 23: 12h6. 192h; Moldenke, Phytologia 12: pam "(1965)
12: ae en 5 and —o. 6.2 Aly wo

found ae plant growing on heav

hopes sate on Sones with eases and ay 1,800——8200 a al-
titude, and describe it as a shrub, 8 feet tall, flowering in No-

- Standley reduces the species to synonymy under Le We —
ta Cav., but to this I cannot agree. The flowers are described a5
eee on Breedlove 14073 & 1095.

In the Harvard University herbaria there re the following two

1966 Moldenke, Notes on Lippia 351

a identified as L. chiapasensis, but not verified as yet
: MEXICO: Chiapas: anne 13 (G); E. W. Nelson 3482 (G).
= hisses bears an annotation "aff. L L. graveolens". The Ghies-
breght 634, distributed as chiapasensis, is actually L. 7. sub~
strigosa Turez
Additional citations: MEXICO: Chiapas: Breedlove 11,073 (Rf),
14095 (Ac); Breedlove & Raven 8331 (Ac); Raven & Breedlove 20076
(ac). c

LIPPIA CHRYSANTHA G

Additional bibliogr repiy: Loes., Verh. Bot. Ver. Brand, 53:
[Abhand], 21,3]. 1912; H . Davis, Life & Works Pringle 20). 1955
Voldenke, Phytologia. 12: Aeni33. oe

arvard University herbaria there is a specimen of Se-

ler & Seler 2023 (G) deg jaca. identified as this species, but

not as as yet verified by me. The Paray 301), distributed as L.
chrysantha, is actually L. pringlei Briq.

LIPPIA CIPOENSIS Moldenk

Additional biblio arasnie Moldenke, Biol. Abstr. 33: 1862. 1959;
Moldenke, Phytologia 12: 133. 1965.
LIPPIA CONTERMINA Briq

Addit ional bibtograpty: scones Phytologia 12: 13)—1% &
290 (1965) and 13: 29.

ag bo ire ee Molde

ional Santina? ie enke, Phytologia 12: 136-138
0965), rp 50, (1966), and 13: eases 1968.
Breedlove & Raven encountered this species in dense woodland

With Quereus and Pinus, at $300 feet altitude, flowering and
fruiting in | Jamary; their no, 8381 is described as having had
"yellow" flowers. Rzedowski found the plant in oak and pine
Woods at 1600 meters altitude.

Additional citations: MEXICO: Chiapas: Breedlove & Raven 831
(2); J. Rzedowski 663 (Ip). HONDURAS: Morazdn: N. L. H. Krauss
98 (Ac). Tegucigalpa: N. L. H. Krauss 991 (Ac).

LIPPIA “csmayeete riq.
Additional ibliography: Moldenke, Phytologia 12: 138——1,0
(1965) ana 13: 1h. 1566.

TA CORYMBOSA Cham.
Sr toe al bibliography: Moldenke, Phytologia 12: 140—-1)1 &

LIPPIA COSTARICENSIS Mold
302 mitional babbbogranhy ; Molden, Phytologia 12: 141—1h2 &
302 (1965) and 13: 162,

352 PHYTOLOGIA Vol. 13, no. 5

LIPPIA CURTISIANA Molde
Additional bib ograpt Moldenke, Phytologia 12: 142—1h3
(1965) and 13: 162. 6.

LIPPIA D IS Glaz.
Additional bibliography: Moldenke, Phytologia 12: lub-1h6,
& - 1965.

LIPPIA DRACOCEPHALOIDES Turcez.
Additional bibliography: Moldenke, Phytologia 12: 14y6--147.
1965.
Turezaninow (1863) places this species in Section Rhodoli
Additional citations: BRAZIL: Brasilia: Heringer 8939/1133 Gh.

LIPPIA DUARTEI Moldenke

Additional Mplisectohy!-Y king, Excerpt. Bot. A.9: 367.
19655 Moldenke, Phytologi 147-148. 1965; Moldenke, Biol.
- h6: 3246. 1965.

LIPPIA DUMETORUM Herzog
tren bibliography: Moldenke, Phytologia 12: 148, 170, &
203. 19:

TA DURANGENSIS Mold
iedisiocel bibhiograniy: Moldenke, Phytologia 12: 149-150
(1965) and 13: 35 & 162, 1966.
aray found this anes flowering and fruiting in in evens
Material = Tass misidentified and distributed in herbaria as
umbellata C
iadittoosl. citations: MEXICO: Durango: Paray 3123 (Ip)-

PPIA EKMANI Moldenke
Additional bibiiegreptys | melgente marbekogie 12: 150--151.
1965 Ranbo, Fesquis. Bot. 21: 26 & {591 :
Rambo (1965) cites st. Ane 3041 : 569 [Pivetta 1157]
o Grande do Sul, the a aitearted | by himself. He
scritinn we plant as an "Elegant, woody shrub, up to 75 cm high",
C

tat
Campo 2 only" in that state, with a general distribution of mi si-
ones; western RGS [Ri o Grande do Sul)".

PIA ELEGANS Cham.

Additional bibliography: woldenke, Phytologia 12: 151—152, 17%
wa “ b6 (1965) and 12: 488. 1

flowers on Jesus 85 are setuid as "white". The F. Ce

Been: s.n. (Sta, Barbara, 10-1-1921], cited by me as this species
in Phytologia 12: 152 (1965), proves to be L. rubiginosa Schau. ine
stead, The Guedes 4,7 distributed as L. elegans is actually Le
schomburgkiana jana Schau.

Additional citations: BRAZIL: Minas Gerais: Jesus 85 (Z).

1966 Moldenke, Notes on Lippia 353

LIPPIA ELLIPTICA Schau.
Additional bibliography: Moldenke, Phytologia 12: 153 & 37.
1965.

LIPPIA FELIPPEI Moldenke
Additional bibliography: » Excerpt. Bot. A.9: 367. 1965;
ut ia Phytologia 12: Parry 1965; Moldenke, Biol. Abstr.
965 «

IA FERRUGINEA H.B.K
Additional bibliography: Moldenke, Phytologia 12: 155-~157
(1965) and 13: 1. 1966.

LIPPIA FLORIDA C
wrap esas cibatcenaniys Moldenke, Phytologia 12: 159--161 &
1965
The Tryons describe this plant as a few-stemmed shrub, “3 2 me
tall, with lavender bracts and flowers that are yellowis sh t
white, growing in grassland and sedgeland with shrubby are
with sandstone and quartzite rocks and derived soils, at 1120 Me
altitude, flowering in November
had Additional citations: BRAZIL: Minas Gerais: Tryon & Tryon 6835

LIPPIA FORMOSA T. S.
Additional bibLiceraphy: epee Phytologia 12: 161-162, 16h,
189, 283, 332, & 339. 1965.
Rece nt collectors have found this plant fruiting January.
n a aaa citations: MEXICO: Baja California: gown Huerta H.

LIPPIA FRAGRANS Turez
Se ditional bibliography: Moldenke, Phytologia 12: 162-164, 189,
’ sd & 339. Se
Rec b collectors have found this plant growing at 1250 meters
altitude, owering in September.
Additional citations: CHILE: Coquimbo: Ricardi & Marticorena

4521/906 (z).

LIPPIA FRANCENSIS Molde
; ig eer came Moldenke, Phytologia 12: 16l--165 &
+ 196

GARDNERIANA S
Additional bib liees oa Turez. ate ey 3% (2):
205. 1863; Moldenke, Sonptaldeta 12: is, 165-165, & 00. 1965.
Turezaninow (1863), in speaking of his L. co i lacunosa

Similis, at folia petiolo etiamsi brevissimo, tamen manifesto in-
Sident, obtusa nec acuta, bracteae praeter inferiores obtusae,"

354 PHYTOLOGIA Vol. 13, no. 5

LIPPIA GEHRTII Mol
Additional Lye youd Moldenke, Phytologia 12: 166--167 &
268. 1965.

LIPPIA GENTRYI cp
Additional b an ee lees Phytologia 12: 167--169 &
363 (1965) and 13: 162.

NDULOSA Schau.
Additional bibliogr. mea: price, Phytologia 12: 169--171,
222, any pines 965) and 1 : 487. 1966.
The rs on B Belén 1156 are naar as “cream-colored” and
the a as in anthesis in Jun
afaitional citations: BRAZIL: ae Belém 1156 (Ac).

eee stouys Fn
Liography: Moldenke, ear ve 170, 172--173,
& is (2965), 12: 488 (1966) , and 13: 32 & 34. 1
Lbfgren 586, distri buted as L. gracilis, i: actually Le

schiabicieetons Sabai

LIPPIA GRANDIFLORA Mart. & Schau.
Pa 02 nace bibliography: Moldenke, Phytologia 12: 174-175 &
280. 1965.

LIPPIA nppeiecahse evi
ional eases Moldenke, Phytologia 12: 175-~178 &
ne — and 13: 17h. 1966.
found this species growing in wooded savannas} ae =
S00 Go gosh te tora ded “eiae te flowers. The Troupin 11.635,
iributed as L. grandifolia, is actually L. abyssinica (Otto & u Distr)

Additional citations: RUANDA: Troupin 11.670 (W--237511).
LIPPIA GRATA Schau.

Aitttioens bibliography: Moldenke, Phytologia 12: 170, 178-179,
&

LIPPIA GRAVEOLENS H.B.K
Additional synonymy: Lantana graveolens Crutchfield & Johnston,
in herb, Pee ee

Additional bibliography: Loes., Verh. Bot. Ver. a 53: 76
77 (abhandl. 21 & 2.2]. 1912; H.'B. Davis, jiite & Pringle, is

3, 301, 310, 332, 333 339 sto, z 363 0 a oo soy
Se tints 13: 3h. 1966; Moldenke, Résumé Suppl. 12: 1 (1965) and

ent collectors describe this <n abe = shrub, bush, or —
brittle twiggy perennial s—12 dm, tall, or a shrub 3-6 greeny
aronatic, with light-yell ow calyxes, ercunj crean-white
lemon-yellow c orollas, and yellow anthers, growing on rocky —

1966 Moldenke, Notes on Lippia 355

banks or rocky clay roadsides, hills, brushy slopes, or em 58%
among boulders, and on slopes or ete slopes — Quer at
altitudes of 1000--1850 meters, flowering in J tiga Septen-
ber, and December. Breedlove 11777 & 13177 ary anton, Barclay,
& Aveuskion 20221 are said to have had “white” flow owers, while on
Herald & Clark rk 429 they are described as "cream", on De S. Cor-
rell rell 31565 "er "cream-white", on Correll & Wasabminen 27707 "lemon-
yellow", a and on Feddema 2288 & 2433 ‘white with a yellow throat".
Rzedowski found the spe species growing in association with Helietta
parvifolia,

In the Harvard University herbaria there are specimens of Wig-
gins 11379 (G) from Baja California and of Nelson & Goldman 7220
(G), identified as L. palmeri S. Wats., which seem (to be L. grav-
eolens "ach although other material of the former collection
is cit as L. palmeri, The Ghiesbreght Sai, distributed
as L. Biimokena: - actually L. substrigosa T Turez

Additional citations: TEXAS: Brewster Co.: De Se Correll 31565
(Ld); E. Marsh 8. rset Webb Co.: Correll & Correll
32139 (Id). Zapata Co.: Correll & Wasshausen 27707 (Ld). MEXICO:
Chiapas: Breedlove 11777 (Rf), 13177 (Ac). Coammila: J. A. de la
Crus s.n. [17.V.1961] (Ip); Marin s.n. [17-I-1961] (Ip, Mi); E.
Marsh 542 542 (Au—~-21249h); E. G. Marsh 1832 (Au—213612); Salgado s.
R, [26.¥.1961] (Ip). Guerrero: H Herald & Clark 429 (Mi); Lord &-
WcComas 621 (Mi). Jalisco: Feddema jema 2268 (Mi), 2433 (Mi); J. 1 Je Rze=

dowsid 17557 (Mi). Morelos: Paray | 3015 (Ip). Puebla: Gen entry, _
Barela ee 2 ER Arguelles 20221 (W—2 Son J. Rzedowski 9010 (Ip).
Jue: 2148 (Ip). San Luis Potosf: J. Rzedowski 1533

(Ip, 2), 3 Bae Tis y. olae (Ip). Tamaulipas: Crutchfield & John-
ston } 4986 (Mi 3 Gonzales } Medrano 108 (Ip).

ek ce GRISEBACHIANA Mol
Additional ericcaien: Uoldenke, Phytologia 12: 19-197,
230, 236, & 240. 1965.

HASSLERIANA Chod.
Additional bibliography: esquis. Bot. 21: 26 & [59].
ot ake, Mebpiarta 12: or 197-199 & 211 (1965) and 12: 190

—

Rambo (1965) cites Herb. Anchieta 1000, from Rio Grande do Sul,
collected by himself. ~ He “ie deseribes scribes the species as a "Small s
", growing on "Dry, shrubby Campos" and notes that —
in Rio Grande do Sul "One ais findplace". He gives the general
distribution of the species as "Paraguay, RGS [Rio Grande do Sul]¥
L Hassler Sed distributed as L. hassleriana, is actually
L. sclerophylla Briq Fe eee a

LIPPIA aa Mart. & Schau,

386 PHYTOLOGIA Vol. 13, no. 5

tional bibliography: Mansf., Notizbl. Bot. Gart. Berlin 9:
155 292g Moldenke, Phytologia 12: 199--201, 27h, & 437 (1965)
1966.

LIPPIA HICKENII Troncos
Additional masonite; Moldenke, Phytologia 12: 202--203
(1968) and 12: 196. 1966.

LIPPIA HIERACIFOLIA C
Additional btbLiograniy: Rambo page Bot. 21: 26-—-27 & [59].
> Moldenke, Phytologia 12: 203--205.

© (1968) cites Herb. Anchieta 1:38, eae: 1802, 4867, 6735,
soos (Rarich s.n.], 8362 (Hurich s.n.], 9105, 9456, 9953, 10059,
10077, 25547 [Heidler 3. S.n.], 25614, 25782, 2 27010, 28682 682 [Emrich s.
n.J, 2939, ae — woe 38138, 3 » 38681, 39170, 70, 39467, 395u2,

Semaine?

do Sul, all col ollected by
designated. He describes the plant as "Herbaceous, from a
rootstock, up to 1m high", growing on "Dry, grassy or more or less
shrubby Campos......Frequent all over the Campos area" in that
state, with a general distribution of "Argentine Mesopotamia, RGS
(Rio Grande do Sul], Uruguay".

LIPPIA HIRSUTA L.

Additional ry igg th Koldacie, Biol. Abstr. 6: it
Moldenke, Résumé $ ; Moldenke, Phytol fos a iz: *
205—-209, 292, & 363 F i968), ork nt & 86 (1966), and 13: 165.

Additional citations: VENEZUELA: Mérida: Bernardi 3135 (N).

LIPPIA HIRTA (Cham.) Meisn
Additional biblio graphy: ag eg Pere ae 12: 198, 199, &
ie (1965) and 12: 489 & 90.
lant has been found pede = “900 gcseteck rene wa The
PP ic. on Reitz & Klein 17411 are ee eae
Additional citations: BRAZIL: Parand&: N77 (Herb.
Inst. Def. pearance Nat. 9449] (Ac); Reitz & ein Tt (Ac); Smith &

Klein 14922

eae HYPOLEIA Brigq. 90, 213-216,
Additional bibliography: Moldenke, Phytologua 12: 190, <47~
& 302 (1965) and 13: 31, & 162. 1966; Moldenke, Résumé Suppl. 13+ 3+

Raven & Breedlove describe this species as a shrub, te feet
» Srowing on steep heavily wooded slopes with Quercus, Fim’
Liquidambar , and tree legumes, at altitudes of 2500--5500 feet.
They describe the flowers as "white" on their nos. 19516 ad
The flowers are also said to have been white on Breedlove 1
Molina R. 11277 & 11330, "cream color" on Breedlove & Raven 2 Ey

1966 Moldenke, Notes on Lippia 357

and "pale-yellow" on Breedlove & Gregory 14215 and Dressler &
Jones 190. Breedlove & Gregor Gregory | found the plant growing on steep
northwest facing slopes with Quercus and Arbutus. Rzedowski found
it in much disturbed woods of Quercus, Liquidambar, and Clethra;
Molina R. calls it "frequent in mixed wet woods" in Honduras. It
has been coliected in anthesis and fruit in October.

Additional citations: MEXICO: Chiapas: Breedlove 13968 (Rf);
Breedlove & Raven 13603 (Ac), 13615a (Ac); Raven & Breedlove 19816
(Ac), ue (Ac). Nayarit: Breedlove & Gregory 14215 Re. Puebla:
Paray 2301 (Ip). Vera Cruz: Dressler & Jones 190 y (Mi); aray 2892

Paray 2001

(Ip); J. Rzedowski 18956 (Mi). HONDURAS: El Parafso: ae E,

24330 (W--2)5228). Morazdn: Molina R. 11277 (W-~2hL5592).

LIPPIA INOPINATA Moldenke

Additional bibliography: Hocking, Excerpt. Bot. A.8: 226. 1965;
Moldenke, Phytologia “ae Eig 218. 1965; "Weldarke, Résumé Suppl.
12: 2 (1965) and 13: 1. 1966.

Rze — describes this species as a rare shrub, 2 5m, tall,

growing at 1700 m,. altitude in "ladera fgnea con vegetacién de
se ay ges deciduo" in Jalisco. pereene found it in xeric

@ gentle south slope with ia, Bursera fagaroides,
sii pthisee intrapilosa at an altitude of ite meters, fruiting in
sheen he MeVaugh reports that it is "abundant" and e perens it
a t m, tall, with conspicuous greenish-white

growing on steep hills in the deciduous forest with Sepeerti, Cor-
dia, and various legumes, at 700 meters altitude, flowering in
Varch, and also in deciduous forest in wooded Tt the forest
composed mostly of Acacia, Bursera, and Ipomoea . It has also been
found in flower and fruit. in November. cate has been misiden-
tified and distributed in herbaria as L. bracteosa (Mart. & Gal.)
Moldenke , Tw Aaeere Ce

Additional citations: MEXICO: Aguascalientes: Detling 8739 (Mi).
Jalisco: R, McVaugh 22888 (Mi); J. Rzedowski 17649 (Au—238081,

Ip, Mi). Puebla: R R. -MeVaugh 22493 291,93 (Mi).

LIPPIA INTEGRIFOLIA (Griseb,) Hieron.
sg eerses bibliography: Moldenke, Phytologia 12: 196 & 218-——-220.

Additional citations: ARGENTINA: Catamarca: Cristébal 482 (Ca—
1181622) ,

LIPPIA INTERMEDIA Cham.
pont pasion) bibliography: Moldenke, Phytologia 12: 220--221 &
LIPPIA Iopop

PHYLLA S
‘ ES Te pone Moldenke, Phytologia 12: 221—~222 &

Additional citations: BRAZIL: Rio de Janeiro: Castellanos 23221

358 PHYTOLOGIA Vol. 13, no. 5
(Herb. Cent. Pesq. Florest. 1068] (2).

ager ape i Molde
ional hour Moldenke, Phytologia 12: 222-~223
(1968) Sen 13: 162. 1966.

LIPPIA JANGADENSIS S. Moore
Additional bibliography: Moldenke, Résumé Suppl. cee es 195
Moldenke, Phytologia 12: 223——225 (1965) and 13: 166.

LIPPIA JANGADENSIS var. EITENORUM Moldenke
Additional bibliography: Moldenke, Phytologia 12: 22h--225.
1965; Moldenke, Résumé Suppl. 12: . 1965.

LIPPIA JAVANICA (Burm. f.) Spre

Additional bibliography: Wild. ‘Kirkia 5: h. 19655 poldenitae es
sumé Suppl. 12: 10. 1965; Moldenke, Phytologia 12: 225235, 2
264, 307, 343, 3h, 352, 358, L32, Lk, 436, 450, 459, & LOL (ages),
12: 483 (1966), and 13: 171, 173, 17h, & 176. 1966; Moldenke, Ré&-
sumé Suppl. 13: 4. 1966

Recent collectors describe this plant as an "annual herb os the
savannas" (Torre 7696 . 7706) ora pipe 1——3 meters tall,
aromatic leaves used for making te i anon in ruderal fers
secondary woods, the te of afte and open aa of Pari-
nari sp., at 500—800 meters altitude, fhommesre in spi ways

1973 & 2367, "whitish" on Torre 2168 65, and "cream-white" on es,

Sousa 157. The vernacular names §Ldnaficarntie a” and "m!fungofana"

are recorded from Portuguese East Africa, Mendonca 737 is said to
h Stuhlmann 1.168, Sousa 158 matclies Jackson 22, “and Mendonca

1S matches Schilectrtent 12230 an and Wilms 1182 in the * British Museum

“the ie "Peter 2056 & 263, distributed as this species, are actur
ally L. 8 schliebeni Moldenke, Garcia 315 and Torre 3637 are Le
whytei | Moldenke, while Baum "250 and and Leendertz sn. [20.x.1909] “

Ne
~ Additional citations: SOUTHERN RHODESIA: E. A. Robinson 6170 ae (H).
PORTUGUESE EAST AFRICA: Cabo Delgado: Torre ee Paiva 12020 (Gi).
Inhambane: Torre 2768 (UL). Limpopo: Torre 2367 (U1). J, Eourengo
Marques: F. A. Mendonga 1645 (U1), 3013 3 (Ul); Ee 4 Sousa 157 (U1),
158 (U1); T Torre 1973 1973 (U1), 7 7696 (U1), 7 7706 (U1). “Vanica e Sofala:
147 47 (01), 1: 12kh (U1). 1 Mozambique: F. A. Mendonga 120° 1201 (Ul,
Ul); Torre 899 (UL), 1201 (U1), 1383 (U1). Miassa: F. A. Mendongs
737 (U1); Torre 43 (U1, U1); Torre & Paiva 10920 (U1). Zanbezia!
Torre 1513 (U1), 5266 (U1).

LIPPIA JUNELLIANA (Moldenke) Troncoso

i

1966 Moldenke, Notes on Lippia 359

Additional bibliography: Moldenke, Phytologia 12: 196, 235--
237, 240, 357, & 358. 1965.

LIPPIA eee

a
mal bibli
us’ (19 (965) and 13: 363,
ow (1863) pre his L. cordata as follows: "L.
ie eyes tetragono, apice ramoso viscido-pubescente; foliis
oppositis brevissime periolatis, cordato-oblongis, supra lucidis
bullatis parce piliferis, subtus reticulato-rugosis densius vesti-
is

grenty: Hordenke, Phytologia 12: 238--239 &

culatis, folia non superantibus. Capitula primo minora, postea
excrescint ad magnitudinem L. lupulinae. Bracteae in sicco fus-
cescentes ; in capitulis junioribus floribus breviores, dein illos
entes. L, Gardnerianae, e descriptione hujus, valde
similis, at folia petiolo etiamsi ’previssimo, tamen manifesto in-
Sident, obtusa nec acuta, bracteae praeter Ste Seon! eae "
Heringer reports the leaves and flowers to

be
tional citations: BRAZIL: Brasilia: H catnine °9195/389 9 (N).

LIPPIA LANTANIFOLIA F. Muell.
gg am bibliography: Moldenke, Phytologia 12: 194 & 2h0.

LIPPIA LASIOCALYCINA Cham.

Additional bibliography: Moldenke, Phytologia 12: 2h0-—-2h2 &
349. 1965; Moldenke, Résumé peg a h. 1965.

Recent collectors refer to this plant as a shrub, 2 m. tall,
with fragrant flowers and pale-purple corollas, flowering in
August, and growing on sandstone. Castellanos * 24347 is said to
have had "rose" corollas

Additional citations: BRAZIL: Bahia: Castellanos 23 (Ac,
Ac). Goids: Prance & Silva 58522 (N). Maranh&o: N. T. Tf. Silva
27775 (N (N).

LIPPIA LASIOCALYCINA var. SAINTHILAIREI Moldenke
oo rasp pelted age Moldenke, Phytologia 12: 22, 252, &
- 19

ite aa LL ERTENSIS Mol
nal bi fitiagrartys es Phytologia 12: 254-—255.
19655 1 pert Résumé Suppl. 12: 2. 1965.
LIPPIA LINDMANII Briq
Scoethmeay tibliography’ Moldenke, Phytologia 12: 255-—256 &

LIPPIA LONGEPEDUNCULATA
( Additional bibl tccrapees per Phytologia 12: 257-~260
1965) and 13: 1h & 29. 1966.

360 PHYTOLOGIA Vol. 13, no. 5

Additional citations: PARAGUAY: T. Rojas s.n. [Hassler 2592]
(ca--950hN)0)

LIPPIA LUCENS Standl.

Additional bibliography: Moldenke, Résumé Suppl. ge pet 19653
Moldenke, Phytologia 12: eae (1965) and = 50h.

Additional citations INDURAS: Comayagua: Yuncker ea &
Youse 6003 (Dp, Mi), 6387 ( (Bt, Dp, Mi).

— oe to ULIFORMIS Moldenke
dditional bibliography: Moldenke, Phytologia 12: 264--265.
1965; Moldenke, Résumé Suppl. 12: 6. 1965.

LIPPIA LUPULINA Cham
onal & emended bibliography: Turcz., Bull. Soc. Nat.
Mosc. 36 (2): 20h & 205. 1863; Greenm., Field Mus. Publ. B Bot. 2%
341. 1912; Moldenke, Résumé Suppl. 12: th. = & 10. 1965; Moldenke,
Phytologia 12: 256, ” 265—=272, & 360 (1965) and 12: 502. 1966.
Turezaninow (1863) describes his L. clausseni as follows: "Le
caule fruticoso tetragono, molliter piloso; foliis oppositis
ee petiolatis ot Han obtusis grosse crenatis, in utraque
ensiuscule pubescentibus; Sy axillaribus solitariis
folio fraser pedunculo) —— ribus. Bracteae magnae roseae, capit-
ula illis L. 1 e Pad
Additional citations: wants Brasilia: Heringer 7873/67 (N)-
Parand: Reiss 12 (Mi); Reitz & Klein 17516 (Ac).

LIPPIA LUPULINA var. PARAGUARIENSIS Chod
Emended eo tr Lippia ae Turez., Bull. Soc. Nat.
rig emended bi biisiierts:
Mosc. 36 (2): 20h—-205. 1863; ietdenie, Ph rigneiegti “so 12. "$965;
Moldenke, Résumé Suppl. 12: hie 1965.
Turezaninow (1863) oe is i "renifolia as follows: "L.
— fruticoso teretiusculo cano-pubescente; foliis oppositis

essilibus, subrotundo-reniformibus obtusis, sinu profundo, super

eve pedunculatis subterminalibus. C apitala rigs Er
in L. lupulina, in statu sicco bracteis fiscescentibus obvallata.
He classifies it in Section Rhodolippia.

LIPPIA MARRUBIIFOLIA Reichardt
Additional bibliography: Moldenke, es 123 273--2% ©
437. 19655 Moldenke, Résumé Suppl. 12: h. 1965.

LIPPIA MARTIANA S ae

Additional Ag said Moldenke, Résumé Syppl. 12: 4

Melanie Phytologia 12: 27h--275, 310, 355, & 55 (1965) “and ibs
The Regnell IIT .942 [26/10/1848] distributed as L. martiana 18

)

1966 Moldenke, Notes on Lippia 361

actually L. sidoides Ch
Additional citations: BRAZIL: Brasilia: De Jesus s.n. (Campus
de U.N.B.] (N).

LIPPIA MATTOGROSSENSIS Moldenke
Additional bibliography: Moldenke, Phytologia 12: 276--277.

oe citations: BRAZIL: Goids: E. Y. Dawson 1486 (Z--
hoto).

LIPPIA MEGAPOTAMICA Spre
Additional pibliosrente: iter age Phytologia 12: 277-278.
1965; Rambo, Pesquis. Bot. 21: 58. 1965.

LIPPIA MICROCEPHALA C

Additional biblio eee Moldenke, Résumé Suppl. 12: 4. 1965;
Moldenke, Phytologia 12: 278-281 & 363. 1

Additional citations: BRAZIL: Brasilia: Sucre 284, (2).

IPPIA MICROMERA S

Additional bibLiograniy’ 72h.
1965; Moldenke, Phytologia 12: rabim231, 3 532, “33 é “339 19653
Moldenke, Résuné Suppl. “2: 2 & 3. 1965.

LIPPIA MICROMERA var. HELLERI Britton) Moldenke
Additional my ne aeterens Moldenke, Phytologia 12: 283--286,
332, & 339. 1965.

LIPPIA MICROMERA var. PALUDICOLA Moldenke
sph tonal bibliography: Moldenke, Phytologia 12: 286-287.

i = ip mea describes the plant as aromatic, with white flowers,
Coming i

Additional citations: BRAZIL: Bahia: Castellanos 25101 [Herb.
Cent. Pesq. Florest. 3870] (Z).

oer eet: é 1. 12: 10. 1965;
tional bibliography: Moldenke, Résumé Suppl. .
cee Phy tologia 12: 287-289, 365, & lO (1965), 12: 486 & L867
and 13: 26,1

Adaitional” citations: "BRAZIL: Bahia: Castellanos 25108 [Herb.
Cent, Pesq. Florest. 3877] (Ac). Maranh#o: Castellanos 25356 [Herb.
Cent. Pesq, Florest, 4125] (Ac).
LIPPIA MODESTA Bri

Briq.
Additional eae at Moldenke, Phytologia 12: 289-291
(1965) and 13: 12. 1966 :

LIPPIA MORITZII Turez.
Additional bibliogr aphy: Moldenke, Phytologia 12: 291-292 (1965)

%2 PHYTOLOGIA Vol. 13, no. 5
Turczaninow (1863) classifies this species in Section Zapania.

LIPPIA MORONGII Kuntze
Additional bibliography: Rambo, Pesquis. Bot. 21: 28 & [59].
1965; Moldenke, ee Ss 12: 293-~297, fig. 1-12, 347, & 43h
(1989) and 13: 15, 29, & 966.
ereunaa? *voldenke, “phytologia 12: 295-296, fig. 1—12.

The L. angustifolia of Chodat, mentioned in is ie of this
species, }, belongs in the synonymy of L. tristis B

collected by himself. He “He describes the plant as a sega up to
on "Dry, mo

growing ’ eg .*
One only findplace on the southwestern plains" in that state. He
gives the general distribution of the species "From aes pee
through Paraguay and northeastern Argentina to western RGS [
Grande do Sul], and Uruguay

LIPPIA MULTIFLORA Moldenke
Additional bibliography: Moldenke, Phytologia 12: 297-300, U8,
458, & 459. 1965.

MULTIFLORA var. PUBESCENS Moldenke
This taxon is now reduced to synonymy under L. savoryi Meikle.

OCEPHALA Schlecht. & Cham.
Additional bibliography: P. C. Standl., Field ep — Pm
3: 403. 1930; H. B. Davis, life & Works Pringle 81, 66 66),
eg lvoe. Phytologia 12: 278 & 300-303 OS (1968), obi Age (19
& ° e

i ove found this plant growing on slopes with an

referable here as a synonym, but I regard it as a distinct species:
Lippia sestosechise has been found in fruit in October. The
Dressler & Jones 190, Molina R. 11277 & 11330, and J. Rzedowsid
18956, distributed as L. myriocephala, are actually rie ~ hypoeia
Briq.
Additional citations: MEXICO: Chiapas: Raven & Breedlove 20118
(Ac). San Iuis Potosi: J, Rzedowski 835 (Ip, Ip)-

LIPPIA MYRIOCEPHALA var. INTEGRIFOLIA L
were piel iserenhe: Moldenke, Mertelcgla 12: 302 & 303-
Tucker describes this plant as a spreading tre + he wee tall,
with white corollas, the throat yellow within, flow ering in Jay
, and inhabiting the north slopes of the "highest peak" at 13
1380 meters altitude.
Additional citations: EL SALVADOR: Moraz4n: Tucker 793 (Mi)+

LIPPIA NIGERIENSIS Mold 306
Additional Sthleeresie? Moldenke, Phytologia 12: 298, 304—-"""

= |

1966 Moldenke, Notes on Lippia %3
458, & 459. 1965; Moldenke, Résumé Suppl. 12: 5 & 6. 1965.

LIPPIA NIGERIENSIS var. BREVIPEDUNCULATA Moldenke
This taxon is now reduced to synonymy under L, savoryi Meikle.

LIPPIA OATESII Rolfe
eens coed Mercagt beh Wild, Kirkia 5: 4. 1965; Moldenke, on
ppl. 12: 7. 1965; Moldenke, Phytologia 12: 306--307, po a
ee (1965) and 13: 171. 1966.

Sree plone & woe
tional bibliography: , Life & Works Pringle 136,
187, “668, & 669. 19363 SS aaa ge het So 12: 307-308. 1965.
LIPPIA OBSCURA Br

Additional sieLoiAisteri Moldenke, Phytologia 12: 275 & 309—

Let
Recent’ collectors — Reap esi this species in secondary

™ » pastures, and cerrado (open scrub), on rock righ oo and
ruderal, at 81,0--860 aetere aivaege : hagas uary.
Addi tional citations: BRAZIL: Parand: Smith Sxisia, & Hatsch-

bach 160) (Ac), 1637. (he). PARAGUAY: eae 5016 (ifi-——isotype).

LIPPIA ORIGANOIDES H.B.K

Additional bibliogr apty: Moldenke, Résumé Suppl. 12: 10. 1965;
Moldenke, Fevtslonia 1 12: 283, 286, 310-312, 331--334, 339, 3h6,
bl6, 452; & Sh (1965), 12: “tse (1966), and ‘138 16h. 1966.
Breteler ribes this plant as a shrub, 1

brown ‘ake
ellow-orange in the throat, growing with other shrubs on
Southern slopes in poor rocky soil, at 1150 m. altitude, flower-
ing in October. Cuatrecasas & Jaramillo refer to it as a ——
with spreading branches, grayish lea vr and corolla and
» Srowing on arid clayish hills, om 2550 to 2700 fie
altitude, The Martius 1037, aieeutael « as L. origanoides, is ac-
ally L. velutina Sch: Schau
Additional citations: COLOMBIA: Cundinamarca: Cuatrecasas & Ja-
Famillo 26688 (W—-2)55980) . Magdalena: H. H. Smith 587 (ii).
VENEZUELA: MG Mérida: Breteler 3205 (N).
LIPPIA OXYCNEMIS Schau.
oo bibliography: Moldenke, Phytologia 12: 334--335.
The flowers on Sucre 750 are said to have been "lilac".
Additional citations: BRAZIL: Brasilia: Sucre 750 (Ac).

LIPPIA 0 OXYPHYLLARIA jog
Additional bibliogr: pati ge es Suppl. 12: 2. 1965;
ars j ziviologia 1 12: 436-337 oT 368), ei 503 & 50h (1966),

364 PHYTOLOGIA Vol. 13, no. 5

Ss rsa cepbeme
ddi tional liography: Moldenke, Résumé Suppl. 12:
19655 Moldenke, gent 4 12: 332 & 337—-3h2 (1965) nee 3: 35 é

Recent collectors have found this plant growing in ee desert
washes, at 225 feet altitude. The Guzm4n Huerta H.331 andI. L.
Wiggins 5539, distributed as L. palmer, are perhaps epe re-
garded as representing var. spicata Ro
516 Raps gina citations: MEXICO: Baja sabieubital Wiggins & Ernst

76 (Mai

LIPPIA PALMERI var. SPI
Additional itp en 55 jr 5 ee 12: 330 J
ae 1965; Moldenke, Résumé Suppl. 12: 2&10. 1965.

s describes the flowers of this plant as “in small stout
ie and found it growing along dry washes, blooming in April.
Guzmén Huerta found it in fruit in January and identified it as L.
palmeri S. Wats

Addi tional thaliokad MEXICO: Baja California: Guzm4n Huerta H.
331 (Ip); I. L. Wiggins 5539 (Mi).

LIPPIA PEARSONI Molde:
Additional Prepac Moldenke, Phytologia 12: 342--3h3.
1965; Moldenke, Résumé Syppl. 12: 6. 1965.

PIA PEARSONI var. SESSILIS Moldenke p
Additional bibliography: Moldenke, Résumé Suppl. 12: 6. 19655
Moldenke, Phytologia 12: 343. 1965.

are pesca te ase! H.W.
graphy: Sepher ad Phytologia 12: 343-34,
368, & go "(965) 8 a a 13 :
The £ Galpin 5%, Rigor distributed as L. pedunculata,
is pio, oy wilmsii H. H. W. Pearson

LIPPIA PEND
Additional bibliography: Moldenke, Résumé Suppl. 12: 5 19655
Moldenke, Phytologia 12: 333 & 345-346 (1965) and 12: 505. 1966

LIPPIA PHRYXOCALYX Brig.
dditional bibliography: Moldenke, Prrvelsets 12: 347-—349-
1965; Moldenke, Résumé Suppl. 12: 4. 1

LIPPIA PICKELII Moldenke pyit
This taxon is regarded Miss me as L, recolletae var- pic elii
(Moldenke) Moldenke, which se nr

TA PLICATA J.

Additional pibic inte Yoldenke, R&sum$ Suppl. 12: 57s 10s,
& 11. 1965; Moldenke, Phytologia 12: + 350-354 & & 31 (1965) and ub
483. 1966; Moldenke, Résumé Suppl. 13:

1966 Moldenke, Notes on Lippia 365

LIPPIA PLICATA var. ACUMINATA (Moldenke) Moldenk

tional bibliography: Moldenke, Phytologia “ho: 352-353.
ick Moldenke, Résumé Suppl. 12: S—7 & 11 (1965) and 13: h.
1966

Recent collectors wp tigi: this plant as a shrub or subshrub,
2--3 m. tall, growing on rocky deserts among granite boulders and
near rivers » at 1450 nm. asst: flowering in March, fruiting in
September. The corollas on Torre & Paiva 11072 are described
having been "white", Material has been misidentified and distri-
buted in herbaria as Lantana sp.

Additional citations: PORTUGUESE EAST AFRICA: Niassa: R. Mon-
teiro 35 (UL). Tete: Torre & Paiva 11072 (U1).

LIPPIA erty op eee var. PARVIFOLIA Seeger Moldenke
Additional bibliography: Moldenke, ete 12: 353—-35h.
1965; Moldenke, Résumé Suppl. 12: 5 6 - 1965.
— POHLIANA Schau.
Additional bibLiograpiy: Moldenke, Phytologia 12: 35h4--355
(1965) and 12: 9h.

LIPPIA rae purer -_ H. W.

Additional b graphy: maidecies, Phytologia 12: 3lh, 357--359,
436, & 461 (1965) a hoe 13: 173. 1966.
al a?

tana? tain ésumé Suppl. 12: h & 5.
ional tin foamereyi Moldenke, Résumé Sup
19685 i Moldenke, Phytologia 12: 359-360. 1965.

LIPPIA PRINGLET Briq.

Additional bibliography: P. C. Standl., ntrib. U. 8. Nate
Herb. 23: 1246. 192); Moldenke, "ales ag "12: 10. 1965} Mo
denke, Phytologia 12s 60-368" (1965) and 13: 7, 35, & 162. 1966;
Moldenke, Résumé $ eid! 2s ae

Standley (192k) reduces this species to synonymy under L. umbel-
ata Cav., but I feel that the two taxa are abundantly distinct. ~
Recent collectors have found L. pringlei in mesophytic woods with

S, in secondary matorrales, and on the banks of large perman-
Snt creeks, at 250--1900 meters altitude, describing it as a shrub
2.5--3 m » or a tree, 15--20 feet ema refers to it
’s an "occasional tree, along pa area" on karstic limestone
eurero ° dland at 2000 tu Rzedowski & McVaugh
describe the bracts as "amarillento-verdoso p4lido" and found the

pag ng on "calizas k4rsticas con vegetacién de matorral

kar
Secundario , VeVangh found it on steep rocky hills with Acacia,
Guazuma, and Bursera at m0) Be SHE, the "bracts conspicuous,
Sreenish yellow; corolla yel
L. Material has been nepaee bap pe and distributed in herbaria as
=: chrysantha Gree

Additional rprepoes MEXICO: Guerrero: Feddema 2868 (Mi); Rze-

366 PHYTOLOGIA Vol. 13, no. 5

donsic & McVaugh 307 (Mi). Jalisco: Diguet 457 (Mi). Morelos:

3014 (Ip). “Nayarit: J. Raedowski ii, 15630 (Ip, Mi). Oaxaca:

a nek coe 22282 (Mi); J Je Rzedowski 19280 (Mi). Puebla: Mercado
3.n. (Mayo 1945) (Ip). “Sinaloa: Breedlove 1572 (Mi).

LIPPIA PUMILA C

Additional bibliography: Moldenke, Résumé Suppl. 12: h. 19655
Moldenke, Phytologia 12: 367, 4291130, : fd (1965) and 13: 4 &
5 3 Mol 3

Recent collectors have found this pl se tins on pre in
low or low pastured woods, and in bogs, at 000 meters alti-
tude, flowering from October to December. Smith & Klein 2 13343
and Smith & Reitz 12853 are said to have had d yellow ; flowers. Ma-
terial has been misidentified and distributed in herbaria as
Verbena sp.

dditional citations: BRAZIL: Santa Catarina: Smith &
1338 (W--2451601), 13786 (W—-2h5160); Smith & Reitz eitz 12853 33 O—

ayer RAMBOI Molde
ditional i taetecty gk Bot. 21: 28 & [59].
19685 Ble Moldenke, Phytologia 12: 2s lsh-iae.

o (1965) cites Herb. Anchieta 1869 Tou sn], 46306,
cage 54583 from Rio Grande do edema, all one hee by bine
except where ere otherwise designated. He describes the plant 4s
"Woody s ,» 2m high", growing in "Dense ieee forest in
moist places.....Along the southern rim of the northeastern

only" in that state, godt a general. distribution of
"So far, RGS [Rio Grande do Sul] o

LIPPIA RECOLLETAE Moro
Additional Seiaerened Voldenke, Phytologia 12: 432—3k
a and 12: 505. 1966; Moldenke, Résumé Suppl. 13: 3 & 7-

LIPPIA RECOLLETAE var. PICKELII se pee ae
— Lippia pickelii eer a 5: h2k--h25~ 190-
Additional bibliography: Mold Hong a 12: 349-350 1966
peed nes ook Range 19665 Minprirens “theual’tappl: 13) 3 & 1
: Hassler 12040 (ca--950509--isotype, N=
girs, Fig. > ae photo of type, N--photo of isotype) -

LIPPIA REHMANNI H. H. W. Pearson ae
Additional synonymy: Lippia rehmanni Gtirke ex Moldenke, 8
a 316, Rea 1959. Lippia rehmannii rehmannii Gttrke ex Moldenke, Résu-
316, syn. 1959.
wheal ttonal er. Verdoorn, yon & Letty, ag?
Transv. 281, pl. 140, . 1962; Moldenke, Phytologi@
13-36 & Pes (ises) and 3 "i713, & 17h. Te

1966 Moldenke, Notes on Lippia 367

Additional illustrations: Dyer, Verdoorn, Codd, & Letty, Wild
Fls, Transv. pl. 140, fig. 4. 1962.

LIPPIA RZEDOWSKII Moldenke

Bibliography: Moldenke, Phytologia 12: 478--479. 1966; Molden-
ke, Roepe Suppl. 13: 1. 1966.

ema describes this plant as a shrub, to 1.5m. tall, the

—— ee the throat of the corolla. yellow, blooming in
ovember, nd growing in disturbed grassland with shrubs in gul-
lies, at "1800 meters altitude.

Citatio ons: MEXICO: Zacatecas: Feddema 2396 (Mi); J. Rzedowski
17635 (Ip--type, Mi—isotype, Z--isotype). __

od Rr Se IAEPOLIA:C
dditional bibli crete Moldenke, ans 12: 452—-l56
(1965), 12: 496 & 497 (1966), and 13: 16h.
LIPPIA SAVORYI Meikle
Additional bibliography: Moldenke, Phytologia 12: 458-160 &
462 (1965) and 13: 362 & 363. 1966.
Peet — Sond,
onal bibli os aig son ag Phytologia 12: 36 & 459—
se (1965) and 13: 173 & 174. 1966.
SCHLIEBENI Moldenk
Additional bdEioaresis Moldenke, Phytologia 12: 48248)
(1966) and 13: 32. 1966.
PIA SCHLIMII var. GLABRESCENS (Moldenke) Moldenke
Additional Dibli graphy: Hocking, Excerpt. Bot. A.9: 367. 1965;
Moldenke, Phytologie 1 12: 48h--l86. 1966.

SCLEROPHYLLA B
Additional eit oscar Moldenke, Phytologia 12: 488--l.92
(1966) and 13: 355. 1966.
LIPPIA SERICEA Cham,
Additional bibliography: Moldenke, Phytologia 12: 492—l9h.

The flowers on Sucre 713 are described as having been yellow.
Additional citations: BRAZIL: Brasilia: Sucre 713 (Ac).

LIPPIA SIDOIDES Cham.

Additional bibliography: , Pesquis. Bot, 21: Be 1965; Mol-
denke, Phytologia 12: oe et 7 (1366) and 13: 361. 1966
LIPPTA

SUBSTRIGOSA Ture
Additional eit agnor rg ager oe 12: 501-505
(1966) and 13: 10, 36, 162, & 355.
Recent collector have ee this alt growing in pinewoods and

368 PHYTOLOGIA Vol. 13, no. 5

on steep moist slopes with Quercus and Pinus, fruiting in August,
at altitudes of 1000 to 3065 meters, and describe it as a shrub,
1--5 meters tall or a tree 2 2—6 meters tall, with viscid foliage,
"frequent" in Guatemala and shrank: on hills and in matorral"
in Honduras, flowering and fruiting in Marc h.

he flowers are described as "cream" on Molina R. & Molina
12116 & 12389, cose Teh on Molina R. 11327, and yoni on
Molina R. 11290 & 1

Additional Sivktfons: GUATEMALA: Alta Verapaz: Molina R. & Mo-
lina 12116 (W—2hy542h). Guatemala: Molina R. & yekines folina 12389 (W—
245425). Huehuetenango: Breedlove 11,86 (A Ac). HONDURAS: S$:
parafso: Molina R. 11327 (W—-2hy5621), 11365 ee 128h)5

(W--2h)5652). Inbtibucd: Molina R. & Molina Tina 13931 (W--2))5359) «

— — eS

Moraz4n: Molina R. 11290 (W=—2h),5586) «

LIPPIA TAYACAJANA Moldenk
Additional ap peered Moldenke, Phytologia 12: 78 90h
and 13: 1 & 218. 1966; Moldenke, ome Suppl. 13: 3-

LIPPIA TAYACAJANA var. SESSILIFLORA Moldenke
Bibliography: mol genke, Phytologia 13: 218. 1966; Moldenke, RE
sumé Suppl. 13: 3.
itations: PERU: pe Hutchison & Wright 7025 (z—type) «

LIPPIA TEGULIFERA v Briq.
Additional bab ography: alias: Phytologia 13: 3--6- 1966+
Additional c ARAGUAY: T. Rojas s.n- [Hassler 9906]
(Ca—950h,31). bce

LIPPIA TEPICANA Moldenke
Pee os bibliography: Moldenke, Phytologia 13: 6-7 & 162.

LIPPIA TORRESII Standl.
Add&tional bibliography: Moldenke, Résumé Suppl. 13: 7 19665
Moldenke, Phytologia 13: 8--10. 1966.

LIPPLA TURNERIFOLLIA C

nett adi tional teccanyi Moldenke, Phytologia 13: 12, ll, 1%
>

vaso citations: BRAZIL: Parané: Reitz & Klein 17722 (#—

LIPPIA UMBELLATA 8
Additional Penltncaree Moldenke, Phytologia 13: 162--103, ’
& 179. 1966. : au
King & Soderstrom found this plant Dagrret on pinewood
er = egg at al yer sero of 8 8500-9000 feet and describe it as #
» to - tall, ed areas, the Paget yellow an the
bracts Tight pried ‘< ocd sample accompanies their nO» 5161.

BOOK REVIEW

Otto Degener

In 1959 G. C. Ruhle published a ae "Haleakala Guide",
With a colored photograph of the silversword (Argyroxiphium sand~
Wicense DC.) native to the Island va Maui. Now appears a compan-
ion boo: booklet of 72 pages, ener ® Canyon and Kokee, A Nature Guide"
(Kauai Publishing Co , Lite, awaii, $1.50), with a color photo-
graph of the Kauai pe ee (Wilk esia gynnoxi phium A. Gray).
The author is Thelma A. Hadley, and her sponsor the "Hui O Laka".

and, above all » Plants. There are l2 haif-tones,
that ae _ drifts at Kalalau Lookout being sar tliel iets lovely.
© many publications, all typographical and other errors

"clutches" » aS appears on page 60. What raises the a a of an
old ig ogist ae the reviewer, however, is the word
used on page 3, "berry" used for ase a on page 39,

a
ae
f:
i
6%
B
&
@
a
B,
a8 i
8
ue)
oa

im;

Qpuntia megacantha Salm-Dyck, a plant with tiny, caducous, awl-
shaped leaves and a broadened stem. The pukiawe (Styphelia
—weiameiae (Cham, & ees Rs Muell. & S. douglasii (A. Gray)
Skottsb.) belongs to the Epacris Family, while the ukiuki (Di-
nella Sp.) belongs to the Lily Eten The silversword is not

linited to to Maui. David Douglas, before his murder on the slopes
of Mauna Kea, Island of Hawaii, by the escaped convict Ned Gurney;
used its dry stalks as firewood. Not six native lobelias are pe-

A. Gray rather than to the Dracaena, better called Pleomele.
Zingiber (not Z Zingeber) zerumbet is a ginger thought to be of
ancient introduction by Vhs Pollynes Polynesians from the South Seas, and is
hepa about as native to the Hawaiian Islands as are the Hawaii-

themselves, The idan rent the Kokee Museum is Santalum pyru-
satin A. Gray, with claret flowers; pense S. ellipticun vam Gaud., with

sFeenish-yellow flowers, is mostly a coastal plant of other islands.
Om the late Earl E. Sherff's i Raat re stay of the lapalapa

ne species — iat. lium (H. & A.) Seem., ‘ati endemis to Cam
One. All are characterized by leaves bearing many leaflets that
369

370 PHYTOLOGIA Vol. 13, no. 5

tremble in the slightest breeze. The koa of Kauai is not Acacia
koa A. Gray but the distinctly Kauai species called Acacia kauai~
ensis by Hillebrand in 1888 and subsequent botanists.

The author's account of man's stewartship of this wonde rful
region is most disheartening. With man's silly introduction ree
the passionflower (Tacsonia mollissima HBK) that smothers na
trees, the raspberry (Rubus penetrans L. H. Bailey) and Likouiaan
(Tibouchina semidecandra Cogn.) that crowd out native shrubs and
herbs, the barn owl which a fat ate flying mongoose, bes Bons oat

man proudly slashes a road with his new toy through the endemic
jungle from the end of the Kalalau Parking Area around the ge
of Kalalau Valley. The road leads practically from nowhere t
nowhere, and is par excellence the ideal method for recente
the beauty of this ~— area by erosion -- in fact, such ero-
sion may promote an eventual break-through between Kalalau bet
and the Waimea Canyon a ein. May the Lord have mercy on
poor Souls who have contributed to the rape of Kauai's natural
resources and beauty!

The present state of Kauai has stimlated in some quarters the
current demand that the most interesting and scenic areas renain-

Having
Waimea Canyon and Kokee since 1922 during protracted b botanical
expeditions, the reviewer feels it is too late to expect the
ederal Government to pull the chestmts out of the fire 1° for
citizens of Hawaii Nei. Without a king's ransom, Kauai can never
be brought back to a natural state within the standard required
of a National Park. If the Departm of the Interior has
Fosppete funds eee for obey one of a National Park
s

lible readers.

The above review was prepared by Dr. Otto Degener, dis
author of the monumental "Flora Hawaiiensis” 4
student of the Hawaiian flora.

distinguished
nd a lifelong

BOOK REVIEW
Alex D. Hawkes

Venezuelan Orchids Tilustrated, Volumes 3 and 4, by G.C.K.
Dunsterville & L.A.Garay; 348 pp. + 344 pp., each with 150 pls.
plus maps + photographs; 1965 and 1966; Andre Deutsch, Lon-
don; distributed in USA by Museum Books, Inc., 48 East 43 Street,
New York 10017. — $20 per volume

In time, a great many books dealing with the Orchidaceae
have been published. Those under the authorship of Mr. Dunster-
ville, long a resident of Caracas, dealing with the species of his
adopted land, are justifiably among the most famous and widely
purchased by connoisseurs.

To date, Mr. Dunsterville has, in his amazingly accurate and
truly magnificent plates, taken up 650 kinds of orchids known to
inhabit Venezuela. We are assured that an additional two volumes
yet remain to be printed, and all of us active in the field look for-
ward to these with great anticipation.

The work of Mr. Dunsterville has received international ac-
claim since his meticulous drawings and comprehensive descrip-
tions and comments first commenced appearance some years ago.
Kudos for his ability is assuredly more than necessary, since he
must be counted among the finest critical orchid illustrators of
Our or any time.

Regrettably, the international opinions regarding his collabo-
rator's ability fall far short of this high opinion. Mr. Garay,
Some of whose first articles in this country I published in a neces-
Sary sort of "translation"' from his version as submitted to me,
Seems to have adopted only a totally autocratic attitude towards
his readers who might dare to question his implied total command
of the Orchidaceae. His command of English has improved vastly,
at least in print, since I received his garbled manuscripts, for he

of whom know him primarily through the otherwise invaluable Dun-
Sterville volumes — agree with his taxonomic decisions. his
case, in almost every instance, we cannot argue, since he does
not deign to explain why his opinions must be accepted by all of us.
This is indeed unfortunate, since a number of the orchids which
have come to his attention are more or less frequent in cultiva-
tion, and indeed are often parents of horticulturally popular, in-
‘ernationally registered hybrids.

All of this is blithely explained away in the Preface to Volume
3 of the present work, where the following truly incredible state-

371

372 PHYTOLOGIA Vol. 13, no. 5

ments, obviously from Garay's hand, which must be unique in
our time, appear, with reference to the "unexplained" (tke quotes
are from the book) synonyms listed: "It is not customary to re-
duce various specific epithets to synonymy without explanation,
but a glance will show that so many are these synonyms that dis-
cussion and explanation of each action would require far more
space than could possibly be allotted in these pages. We would
like to assure our readers, however, that our judgement in re-
ducing a species to synonym status is in every case based on ac-
tual examination of the type specimens."

Venezuelan Orchids Illustrated is a memorable series of vol-
umes, artistically without peer in our time. It seems most un-
fortunate, though, that Mr. Dunsterville had not allied himself
with an orchidologist, rather than with a person who creates his
own unique regulations as he goes along.

BOOK REVIEWS
Alma L. Moldenke

"Native Wild Plants of the Northeastern United States and Canada”,

by H. F. Montgomery, xxxvi & 193 pp., illustr. Ryerson Press
of Toronto and Frederick Warne & Company of New York. 1965.
$3.95

This pocket-size book can be a helpful companion out on the
trail or on the library shelf. It epee et succinctly and illus-
trates by clear line drawings and two dozen beautiful color photo~
graphs almost a tenth of the indigenous plants of the area. T
commonest and the showiest among the herbs are chosen. The time

of blooming is often noted, but the ranges of the species are me
mentio:

The plants are arranged into families te a Gray's Man-
ual. A fifteen page simple and obvious key directs es untrained
plant hunters to the proper sections of the boo on ss

The printing, paper, drawings and photographs are all fine -=
though the word "aquatic" slipped through cieecehiad on p. xxi.
There is a simple glossary: nothing more is needed since the

This book is an effectively popularized path to one phase %
the out-of-doors written by an excellent botanist with years ie
field studies, nature photographing and sketching to his credit.
wesis se the Northern United States and Cenatiats by F. H. Monte”

» xxviii & 226 pp., illustr. Ryers ss of Toronto
Aederiak Warne & Company of New gy "965. $3.95

1966 Moldenke, Book reviews 373

This book a a pani fine companion volume to the o
viewed t therefore belongs in ote pock-
et of the sane field sac or on the same library shelf.

The forma ar - simple language, elementary key, fam-
ily a cane mea a recognizable pastebin dagisaes of 365 of the most conm-
mon and/or the most conspicuous ted" plants, clearcut line

s of almost 300 of them, frequent mention of origins and

distribution, helpful glossary.

While the bo ok is enriched with many French-Canadian common
names, many from the standardized American list are ignored.

"The ged Dictionary of Science", edited by E. C. Graham, xvi &

pp., illustr. Evans Brothers, Limited, of London and
Macmillan Company of New York. 1965. $5.95

more than 25,000 words and word-greups taken from all branches of

ology, and , as as from important fields of sci-
énce in industry, such as Engineering, Aeronautics, and Metallurgy"
in Basic sh and such additional t

t 1
People reading in the various sciences will find this book
helpful, quite w p-to-date, provided with both American and Engli
spellings wherever they differ, ee much more efficient to handle
than any huge general dictionary eople with training in the
Classical languages and linguistics ett probably be distressed

t given.
The botanical sciences seem to be somewhat slighted. As an ex-
rise - Pe ~ “crepuscular" is defined only zoologically, where
f flower opening might also have been mentioned -- but per=
a the definition is Kes! enough so that its botanical applica-
tion could be correctly

"Drawings of British Plants", Part XXII Serapinilariecene (1), by
Stella osacrete, 39 plates & index. G. Bell & Sons, London.
1966. lls 6d

y this Bane es
covered volume. Besides whole plant iuinstratione » flower,
fruit, root and other parts are show separate ly with their eagah-
tic features and accurate magnificatio
May this’ project continue to be i with the same or simi-
lar scientiric accuracy and artistic beauty until all the know
f this area are so trea
"Flora of Turkey and the Aegean Islands", Volume I, edited by P. H.
Davis, 567 pp., illustr. University Press, Edinburgh, and
Aldine Publishing Company, Chicago. 1966. $27.50

This critical, thorough work is the first of eight planned vol~-

374 PHYTOLOGIA Vol. 13, no. 5

umes. It covers the pteridophytes, gymnosperms, Ranunculaceae,
Paeoniaceae, Nymphaeaceae, Berberidaceae, Papaveraceae, Crucifer-
ae, Capparidaceae, Resedaceae, Cistaceae, Violaceae, olygal-
aceae, with carefully constructed keys, full diagnostic descrip-
tions, synonymy, literature references, habitats, distribution
records by word and by many maps, type citations, taxonomic re-
lationships, and some drawings.

This excellent project represents needed scholarship on 902
species, of which 20 percent are endemic since the last publish-
ed flora was

Irano-Tur , Mediterranean), and s
homeland (gene pool) of several of today's western cultivated
plants and weeds.

"Chromosomes, Giant Molecules, and Evolution", by Bruce Wallace,
xi &171 pp., illustr. W. W. Norton & Company, New York.
1966. $1.95

The author offers this paperback book as his sword in a duel
with the anti-evolutionists; he wants to put it into the hands of

and paren
these people to read at home or to study in adult education
groups, seminars, courses in state teachers’ colleges, or biol-

emphasize, contrary to the author's clain, that such classical
evidences for evolution from comparative anatomy, © ryology,
paleontology, plant and animal breeding, etc. have been effec-
tively convincing. Witness the changes in and the increase of
biology textst This book just adds more modern, convincing evi-
dence in fine forn.

inducing polyploidy; they appear in newsprint and popular scien-
— articles. The 39 charts and their explanations are y
effective.

"Taxonomic Terminology of the Higher Plants", by H. Ie resorts
= Hafner Publishing Company, London & New York-
Pr4

About half of this book consists of a very well defined 10% |

sary of a representative, but not sufficient, number of poner"
botanical taxonomy, plant distribution and spel

cation. About a quarter of the book follows with an interesting

a ee ee ee

1966 Moldenke, Book reviews 375

subject classification with repeated definitions under such topics
vation, corolla, -o » Stamens, t terminology.
Then there are listed almost 2,000 specific epithets with gender
endings and their meanings. Eighteen pages of Greek and Latin
components with their meanings and cognates can be partic eek fe

er of this review copy is poor and very dry. The prin-
ting is clear and a free of errors, although "original" is
misspelled on p.

e@ book would be "really ne only to beginning or transient
batenks in the field of taxonomic botany.

ignabe Biology", by Mary S. Percival, xv & 23) pp., illustr.
Pergamon Press, Oxford, London, Edinburgh, New York, Paris,
. 1965. $3.85

This paperback book is a delightful and thoroughly scientific
of the science of flower life which lasts from

pollen dehiscence of the first ripe or the incep-
tion of pollen receptivity by the first stamen until the end of
these activities. It the literature from Sprengel'

mark publication of 1793 up to current research. It covers for

owers - sex, structure, isolation mechanisms, use as animal food,
and pollination of the following types - anemophilous, hydrophil-
ous, ornithophilous, cheiropterophilous, entomophilous. There are

two records of rain pollination and the mention of doubtful

malacophily, Finally, there is a short chapter on practical work
in floral biology. The paper, printing, illustrations and indexes
are excellent,

This book is a gem for the general reader of scientific —
ests and for biology course enrichment on both the secondary a
university levels.

"Laboratory Manual for Plant Anatomy", by Richard A. Popham, vii &
= 966 $h..95 illustr. C. V. Mosby Company, St. Louis, Missouri.

So many of the printed and mimeographed lab manuals in use over
the last ‘hundred —_ on our campuses have been worse than wor

for if it is not there" - ang so orients instructors and students
om a more truly saa een approach,

Proper vocabulary is stressed at the start of each topic, with
the students hunting a meanings or depending upon class lectures.

376 PHYETtORLOGd A Vol. 13, no. 5

Fine photomicrographs and drawings are given for student labeling
in a variety of ways on their own study time: but one good draw-
ing should be required. The many questions asked are thought=-
provoking, organizing, exploring. The topics are covered develop-
mentally and include roots, stems, leaves, and germinating seeds.
Outside of a flower bud section, nothing is shown on flower or

re. ?

It is g to note that species names are given along with the
generic ones under the illustrations. The instructor should surely
let the students know what these plants are since several may be
passed

ie botanist check on the validity of the names used, for then
certainly "Clerodendrum" would have been employed instead of
"Clerodendron", "Pinus nigra var. poiretiana" would have been

used instead of "Pinus laricio", etc. A much wider range of plants
is shown here than usually appears in such a text or manual.
References are well arranged according to topics, but no mention
of Nehemiah Grew's classic work is given.

“The Plant Kingdom, a Laboratory Manual", edition 2, by Paul C.
Lem tr. C. Ve

on & Norman H. Russell, viii & 215 pp., illus
by Company, St. Louis, Missouri, 1965. $3.90

ax
uses didactically a not very modern system of plant classification.
Hopefully the class lectures will to the students the con~
cepts system and systems, and the fact that the latter are not
as s Cc

outline then.

Squeezed in at the end of the book are a few lab exercises on
photosynthesis and fermentation, as well as some field exercises
in ecology, all similar to those done with the brighter classes
our present-day high schools.

Reference sources given at the end of each exercise unit are
well chosen, but there is no mention of Lyman Benson's excellent
books on the subject of plant classification.

"Nitrogen Fixation in Plants, by William D. P. Stewart, 168 PP»
illustr. Athlone Press in London & Oxford Press in Toronto
& New York. 1966. $.00

This compact outgrowth of university lectures explains with ex-
cellent clarity the nature, physiology, and biochemistry of bes:

utilization of molecular nitrogen for cell protein and protoplas®

with bluegreen algae as green manure on rice fields, for
There is a fine bibliography.

PHYTOLOGIA is financed entirely by its contributors, each one Bes,
in advance for the entire cost of printing, binding, and distributing his
tribution. All money received from subscribers, after the expenses of a
lections have been deducted, will be distributed among i contributors upon
the completion of a volume, in proportion to the space which they have used.

Each contributor is therefore a shareholder in the magazine, assuming his
part of the expenses and sharing in the profits, if any accrue.

Each number consists of not less than 32 pages. All manuscript accepted
will be published in the next issue, so that the size of numbers may vary
greatly. A volume will contain about 32 aa 512 pages, or a smaller
number of with an equivalent number of plates. This plan insures
immediate publication of all accepted onc

Illustrations will be published according to the desires of the authors. No ee
extra charge is made for line drawings, such as are ordinarily reproduced peat re
zinc, or for diagrams, tables, or charts, provided they oo to certain :
limitations of size and proportion. An extra charge will be made for half- —
tones, depending on their size, as fixed by the engraver, a a minimum of
about $2.25.

Articles dealing with research in all lines of botany, in any reasonable
length, rate eae sketches, and critical reviews and summaries of liter-

amateur or so-called popular type, and polemics will not be published. Advice
on the suitability of manuscripts will be solicited, if necessary, from quali-
fied botanists. :

Under the present cost of printing, the basic rate for a page OF fraction
thereof is approximately $3.00 for an edition of 250 copies. This price is subject
to change without notice, since it depends entirely on the prices ee
in the printing industry.

_ Reprints will be farsiahed at cost. A proportionate fraction of the ction
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2 ee Cuan ian the editors will send detailed instructions ms concerning
preparation of manuscript or further information about the —
ee ee

PHYTOLOGIA

Designed to expedite botanical publication

Vol. 13 September, 1966 No. 6

CONTENTS

_ DUGAND, A., & JOHNSTON, M. C., A new Karwinskia
(Rhamnaceae) from northern Colombia ........... 377

DUGAND, A., Notas sobre la flora de Colombia y paises vecinos ., 379

“MOLDENKE, H. N., Additional materials toward a monograph of the

genus Cullicarhd. Pra ee ee ee ee a a ea 401
MOLDENKE, H. N., Two new Satie’ OF EMO 6 ink ca 445
. MOLDENKE, is tsi, ROOOR VOU ee ee tee a 446 7

Wrsvoun: Boranrcat
OCT 3~ 1966

GARDEN LIBRARY

Published by Hanae N. Moldenke and Alma L. Moldenke

303 Parkside Road
Plainfield, New Jersey, U.S.A.

"Price of thin. deaber: $1; per volume, $5 75, in » advance, : i
or $6 at Se of volume —
Volume 13, number 5, was issued — 24, 1966

A NEW KARWINSKIA (RHAMNACEAE) FROM NORTHERN COLOMBIA,

Armando Dugand
Barranquilla, Colombia
Marshall C. Johnston
University of Texas Herbarium, Austin, Texas

€ following is part of a taxonomic study of the flora of
the semi-arid areas on the Caribbean Coast and the lower Magda-
lena Valley in northern Colombia, South America, in which the
senior author has been engaged for many years

uring the last ten months he has had the opportunity of
critically studying his own botanical gatherings as well as
those made by several other collectors and botanists who have

northern Venezue s of the study has been
made possible through a Fellowship grant from the John n
Guggenheim Memorial Foundation, to the Directors and Trustees

» the Associate Curators and other
staff members of the United States National Herbarium, Smithson-
ian Institution, Washington, D.C.

The junior author acknowledges the assistance of a grant
from the Michaux Fund of the American Philosophical Society in
his study of Karwinskia and other Rhamnaceae.

WN; (ee) va -
Arbuscula usque ad 5 m. alta, trunco basi usque ad 27 cm. diam.
ramu UL

rum
hornotini oppositi, graciles, fusci, propter ipsorum
intorsionem ¢ fere plano jacentes, tempore florendi flagel-
- 1m. crassi, internodiis 3 ad 5
Praediti, sparsim minuteque hirtelli aut inferioris internodiis
Glabrati. Petioli ob torsionem internodiorum eodem fere plano

0.5

q
sunt, 35-62 mm, longa, 17-28 mm. lata apice acutissima vel brevi-
ter acuminata vel acute apiculata; omnia supra atroviridia in
377

378 PHYTOLOGIA Vol. 13, no. 6

sicco saturate olivacea vel fuscescentia, subtus valde palli-
diora subglauca nervis pinnatis conspicuis at vix prominulis

sub angulo PO-45° divergentibus strictis vel leniter arcuatis et
an

mm. longae, caducae.

Cymae umbelliformes axillares 2-h-florae, pedunculo 1-3 1/2
mm. longo sparse minuteque hirtello, pedicellis ad anthesin
2.3 mm. longis, postea ad 4-5 mm. protractis, sparse minute hir-
tellis denique glabratis. Cupula floralis 2-3 m. lata, pilo-
sula; sepala deltoidea 1-1 1/2 mm. longa; p
mn. longa. Ovarium ut videtur biloculatum, loculo utroque
uniovulato.

so Collection: COLOMBIA. Departamento del Magdalena:
alle del Ri (parte occidental), al oeste de Los Venados
(10° N., 73° 42! W.)3; llanuras subxeroffticas de sabana abierte

°o

arbustos y bejucos), 60 m. sobre el nivel del mar, Abril 23,
1963, A. Dugand 6326 (holotype US; isotypes: COL and others to
be distributed).

nly other collection seen by us is COLOMBIA, Depart-

s Venados (10° lat. N., 73° 45" long. W.), 70 m. alte,
Abril 21, 1963, A. Dugand 6294 (COL, US, etc.

This is a small tree with declining branches found rather
scatteringly in the small, isolated patches of woodland (locally
wn as"matas de monte”) that dot the otherwise open savannas
on the semi-arid plains west of the Rio Cesare, 4 tributary of

the Magdalena River in northern Colombia. The flowers when
collected were green color, with the petals paler, almos
whitish.

In most species of Karwinskia, the whiplike early-season
flowering branches proceed to elongate, producing a dozen or
more sterile nodes. This occurs while the drupes at the early
nodes are maturing. The leaves at these sterile nodes are more
remote, longer, proportionately narrower, firmer and less
branous than those of the flowering nodes. Unfortunately we d
not yet have of this new species, K. colombiana, material show-

Fa =

long and 10-23 mm. broad, of the middle nodes 33 4 mm.
13-24 mm. broad, and of the fourth of fifth node 39-80 mm. long

ee eg Pn ae ee

sree te

1966 Dugand & Johnston, A new Karwinskia 379

and 16-26 mm. broad; the blades of the middle and distal nodes
show more (usually 9 to 12 pairs) lateral nerves than those of
K. colombiana and the nerves have a greater tendency to exhibit

c teristic of the Mexican Karwi as. The petioles are 8.
mm ng, us longer than those of K. ene. Each e

has 8 to 15 flowers, versus the 2 to ine

cyme of K. colombiana. With all these ddetinetions, K. colom-

biana is an adequate ly distinct species, at least as distinct as

Various ee which have Horie Sousidered't o be

species in M

Not o eal = ‘eis colombiana the first species of this
genus to be recorded from Colombia; it is also the first record
of tS api for South America. A report of the genus from
the sony Chaco was based on a collection of Rhamnidium
elaeocarp

NOTAS SOBRE LA FLORA DE COLOMBIA Y PAISES VECINOS

ando Dugan
prs eae ae

La serie de artfculos que inicio con estas notas ha de con-
tener algunas de las observaciones mas ortantes que he hecho
:

tg
z
:
‘
Bo
a:
5

Os
nomenclatura de las especies que componen la ae "gel ane
cAlido de Colombia, con especialidad la de la Costa del Caribe
y la correspondiente llanura infero-magdal mica. En gage
casos estas observaciones atafien tambien a la flora de
Tegiones inmediatamente vecinas en el litoral caribe (Venezuela
y Panamé) y aun la de pafses m4s lejanos del continente suramer
icano o de la América Central.

e un ~~ dio sash ae hace muchos afios

rman parte d
intennnnta: en demasiadas ocasiones por circunstancias diver-
Sas, que pipe no so la taxonomia de ae especies, sino
también el a specto sinecial de ellas en cuanto sean componentes

de oplectiniaenes definidas (sinecias: formaciones, asocia-

clones) en el territorio que ha sido teatro principal de mis

ectividaies, Dicho territorio haéllase comprendido grosso modo
Mar Caribe

Limi
de ihe Antillas, y que se extie nde desde el litoral hasta aprox-
imadamente 9° 30' de latdond septentrional, desde la gg nor-
tela del Departamento de Bolivar hasta la peninsula de
Guajira, pans at las Takes que bordean el Bajo Magdstene
y

hasta su des embocadura, y las que se dilatan entre este rfo
€l Cesare,

€ studio peeaty: de mis colecciones ha recibido impulso
extraordinar » en muchisimos casos definitivo, gracias a mi

€stada de mds de un afio en el Instituto Smithsoniano, de aaa,
ington, E.U.A., en cuyo herbario (United States National

380 PHYTOLOGIA Vol. 13, no. 6

Herbarium) héllanse depositades numerosas e important{simas
colecciones de la regidn neotropical, y cuya pibliohess botan-
ica es de las mas completas en este ramo. Me ha sido gue
llevar a buen cabo esta temporada de estudios merced a
Fellowship que me fue gentilmente concedido por la John ppcet
Guggenheim Memorial Foundation, a cuyo Presidente, Administra-

tambie
rere pone pre. “gel -oiebrrmaetp Smithsoniano, particular
mente al Director del Museo de Historia Natural, al Presidente
del Departamento de Botanica, al Curador Encargado la
Division de Fanerdgamas, a los Curadores Asociados del Herbario
al d

estas entidades que de un modo u otro estado su ayuda
muy valiosa facilitando grandemente la epi de mi
rabajo.

CACTACEAE

ACANTHOCEREUS SICARIGUENSIS Croizat & Tamayo, Bol. Soc. Venez.

Cienc. Nati 11: 75. 1947 fYeneavslec Estado hare Aiaarkas)
especie de cactaécea, sin

pitanayo comin de tres filos (A. ies (L. a perp

ase con m ia en los bosques 4 de la costa
payed pa del Caribe desde la Guajira (donde los indi
llaman kachésura De to gntico, donde se
conoce como "carddon de perro”. lares se identifican

Mis ejemp
satisfactoriamente con los on Venezuela en el Instituto Botdnico
de Caracas.
Los tallos o cladodios son primeramente erguidos, luego
menos agin es pha © también reclinados sobre

y 1m. 60 de nates san: nt cierto modo heteromorfos, @
saber: los tallos inferiores (de 3 a 4 cm. de ancho) tienen 4 0
5 filos (a veces By ligeramente ondeados o festoneados; 10s
siguientes (de 2 a 2.5 cm. de lat. y largos de menos de un

tro) tienen 3, 4 0 5 alas longitudinales 4d em, de
alto, ramente ondeadas en el filo (el cilindro lefioso in-
terior tiene .8 cm didmetro); de estos c s salen
otr g ge cs al ert fe o

uni

cho, otras, ten con sdélo 3 filos y anchos de 2 Y)
estos brotan finalmente artejos achatados y anc{pites “(ae eer
dos filos) mcho més cortos (4 a 15 cm. de longitud), pero

en ios

El tallo basilar viejo y sublefioso de estas plantas pre=
senta a manudo mayor numero de filos o "costillas” (6 a 8),
pero es casi siempre muy corto.

Las pees | flores que he visto, ya my marchitas y en mal
estado (al parecer son nocturnas), eran de color psn reise
ZCO3 su deuetan d total es de 15-17 cm., el tubo es wis
(0.5 cm, diem.) y cil{ndrico desde la base hasta unos 11 @ a
cm, y luego expandese en forma de emb

See

re

1966 Dugand, La Flora de Colombia 381
N e ha descrito el fruto de Acanthocereus sicariguensis;
o he visto muy s. Uno madurar que coleccioné arriba de

diametro, con 9 costillas longitudinales poco prominentes, cada
costilla con aréolas flocosas blancuzcas, globulosas, de unos
2mm. de diam., distantes unas de otras ioe mm., y que llevan

s flores y los frutos brotan perio net los artejos cortos
y sceipives como ve los ree at y 4-5-2

te)
la planta crece en lugares Aescebiektse asf asoleados, y verdos
palido cuando se halla a la sombra. En muchas ocasiones, pines,
he observado plantas a la sombra o al sol cuyos artejos ostenta-
ban ambos colores o matices intermedios.

Esta especie, muy bien definida por Croizat y Tamayo en su
protélogo, tiene cierta afinidad con el A. brasiliensis Br. &
Rose, del cual distinguese a simple vista por tener las aréolas

2

espin{feras mucho ma rea unas de otras oo. a
2 » a veces 2 1/ yooh raramente 3 cms.). En A. brasilien-

sis nhl tanee a mds de 3 cms. fratesicemente a oe o 5 cms.
COLoME MBIA. - Atlantico: aitite de Salgar, 40 m., matorrales

spinosos, Feb. 1964, Dugand 6706 (US) y 6706-B (US); - Poco
winded tess al S. de caseayal:; camino a Lefia, Feb. 1964, Bigand
6694 téaic en US). Bolivar: Hacienda de Coloncito, near Turbaco,
200-300 m., Nov. 1926, Killip & A.C. Smith 14339 (US). Guajira:
al S.E. de Riohacha, Kms. res a 12 de la carretera a Maicao, =ences
de 20 m., Jan. 1964, Dugand 6654, — a ria : pte ura"
Tee tena Ber del rio peed al o Venados
(10° la 42 ' long. W.), anes “bir Catto forthe 60 m.
Feb, 1961, yitine 5587 (COL).

ACANTHOCEREUS TETRAGONUS (L.) Hummelinck, Succ. 20: 11, 165.
948; Backeberg, Cact. 4: 1933. 1960); Arnoldo, Zakflora
(Curagao) ed. 2: 99, figs 68, 69, 70. 196
Cactus tetragonus L. Sp. Pl. 466. 1763 (Curagao); non
S-_tetragonus Willd. 1797.
eus tetragonus Miller, Gard. Dict. ed. 8:
non cereus ‘ae agonus Saasorth 1812 es Cereus sieieicans sensu
Br. & Ros
Ac a pentagonus sensu Br. & Rose, Cact.
1920; Hummelinck Med. Bot. Mus. Herb. Utr. 35: 42. 1938 atque
auct, gs
athocerens colombianus Br. & Rose, Cact. 2: 122. 1920

(Coeabi: Calamar and Bonda); Backeberg, Cact. 4: 1932. 1960.

hocereus Pitajaya Dugand ex Croizat in Caldasia 7: 135.
1943; "8 non ‘actu s Pitajaya Jacquin, Enum.23. 1760; Sel. Stirp
Amer. ee

ai as colombianus de Britton y Rose, cuyo Tipo

emsans” ;: rab sby 23, NY) fue coleccionado cerca de Calamar,

382 PHYTOLOGIA Vol. 13, no. 6

Depto. de Bolivar, en la regidn costanera del Caribe que estoy
estudiando, no se puede distinguir de los ejemplares comunes de

minales cuya anchura excede la de 4 a 6 cms. que se observa
usualmente en esta especie selepnner® desecados), pues llega a
1/2 cms. segtn los que he visto. or lo tanto no dudo que
algunos ~chenaia los 9 sau del Tipo de colombianu
En todo caso, la flor de H.H. Smith 2423 , procedente He

mn y Rose citan c
re colombianus, esidéntica a las de Elias 306 y 1089 (U8), ge
and Jaramillo 3413 (US), 3415 (COL) y_Dugand 6794 (US) il
artejos corresponden bien a 1a los dee ejemplares sin flor
Dugand & Jaramillo 3436 (US), Romero-Castaiieda 2074 Poet - otros.
El ntime el tamafio de las espinas radiales y de la unica

tetrdgonos. a los he visto pentdgonos, excepto raras veces
en la base misma de la planta, donde el tallo principal forma
tronco lefioso.

especto al Cactus Pitajaya de Jacquin, dicho autor dice en
el protdélogo que el fruto esta adornado por fuera con "folio
aliquot calycinis praemorsis Pereskiae adinstar' y tiene la pulpa
blanca; lo cual no corresponde de ningiin modo al Acanthocereus
tetragonus cuyo fruto no presenta tales "hojue elas" (escamas
folidceas * sino aréolas espinulfferas, y ademds s mS puree: es de
co

ficie d e.
i ie gamos por las tales "hojuelas calicinas" o escama
a

tallos triangulares y cuyo fruto, rojo carmesi por fuera, —.
blanca la pulpa, y ademds este provisto de escamas foliadceas como
las que describe Jacquin,

Seguin dicho tokiaice, la epoca de pianten = Cactus Pita

e julio a septiembre. Coincide asf c a del Acantho~ |

cereus tetragonus, que. @in duda alguna es el Sse a de pitahay@
silvestre mas comin y conocido en el litoral nortefio de Colombia.
En muchos lugares dridos de la llanura costera abunda este
cardén, que otros llaman simplemente "pitahayo", formando grupo®

eee

——)

1966 Dugand, La Flora de Colambia 383

pequefios y gregales caracterizados por numerosos tallos y artejos
muy alargados, ora mds o menos erguidos o inclinados, ora arque-

del Norte de Santander, valle del rio Peralonso, 120 m, (Araque-
Molina y Barklay 18 NS 129). Hallase igualmente en la costa de
Venezuela y por supuesto en la isla de Curaz zao, su patria
lasica.

ACANTHOCEREUS TETRAGONUS (L.) Hummlk. var. MICRACANTHUS Dugand,

atque centrali = cm. gis, centrali interdum usque a
m. longa, omnis albidis apice autem infuscatis. Areolae
spiniferae inter se 4-5 cm. distantes, ca. 0.7 cm. diam., lanu-

COLOMBIA. - Atldntico: bosque semidrido junto al arroyo de Juan
de Acosta; alt. 15 m.; 27 Jan. 1962, Dugand 6018 (herb. Dugand)
vulg. "eardén de pitahaya'

1962 plantée jun artese de micracanthus en el jardfn de mi
Casa; a poco arraigd y ha crecido muy lentamente produciendo

sea Ys iy distinta, cercana quizas al A. subinermis Br. & Rose,
de Méx

servado la mismfsima variedad en (distintos Oe del
icictebesan ts ——— y en las cercanfas de Los Venados,
Depto. del Magd

PILOSOCEREUS LANUGINOSUS (L.) Byl. & Rowl. in Cact. & Succ. J.
Gr. Brit. 19: 67.
cactus jamtiet needs Ly Bee Pl. ed. I: 467. 1753 (Curacao).
e Cephalocereus bang i nosis (L.) Britton & Rose, Contr. U.S.
at. Herb. 12: 417. 1909.

Pilocereus eas (L.) Backeberg, Kakt. -ABC 330.1935
Pilocereus colombianus Backeberg, Kakt. -ABC 330. 193
(quoad planta littoris septentrionalis Colombiae; non Cephaloce-

reus colombianus Rose 1909).
“sus _sofoubianus

Pilos socereus colombianus Backeberg, Die Cact. 4: 2433. 1960
(quoad pl. litt. septentr. Colomb.; non Cephalocereus colombia-
Rus Rose 190 09)

38h, PHYTOLOGIA Vol. 13, no. 6

rueba evidente de la wconfusidn que solemos padecer a menu-
s

arios o incompletos, es la siguiente: Britton y Rose al
describir a Cephalocereus colombianus (cuyo Tipo cincgvedes del
valle del Dagua, en el occidente de Colombia) citaron como de

n la cos aribe; pero m4s tarde (en The Cactaceae 2:
55-56. 1920) enmendaron esta identificacion diciendo que Los
jemplares del n de Colombia, citados en la descripcion

e are or

original de Cephalocereus colombianus, "we now include in Cepha-

locereus russelianus". En realidad las dos especies son muy
tin !

erg, en su reciente obra (Die Cactaceae 4: 2433-2434,
2449-2450. 1960) tampoco parece estar bien seguro de la identi-

ficacion de estas cactdceas puesto que a Pilosocereus
Puerto Colombia los cita como P. co Lonbianus (p. 2434) y a los

de la Guajira, como P. lanuginosus (p. 2450). En realidad los
de la costa colombiana del Caribe (Puerto Colombia, Sabanilla,
Santa Marta y - mire son identicos a los de Curazao, ti{pi-
cos de lanugin

Bac

“ner se refiere a la localidad de colombianus asi:
"yon NW-Kolumbien (Venticas uct ——, situando erréneamente el
page a rio _— el n ste de Colombia, cuando lo cierto

ntra a pil ya de este nda en las faldas de
is Gai dieera: Occidental que miran hacia el Océano Pacifico.
Puerto Colombia y la peninsula de la — hdllanse en la costa

tienen 25 aguijones o mds en cada areola, y ciertament “i

los que menciono bajo dicho nombre, en el supuesto se dudoso
de que sean realmente especie distinta.

PILOSOCEREUS COLOMBIANUS (Br. & Rose) Byl. & Rowl., Cact- &
wee, J. Cr. Beit. 19+. 3,;66,-1957..5 Backeberg, Cact. 43
2433. 1960 (quoad planta vallis fluminis Daguae in Colomb.
Pie 8 tit
Cephalocereus colombianus Britton & Rose, Contr. U- p.-Hah>
Herb. 12: 416. 1909 (Colombia: Depto. del a Cordillera

——

1966 Dugand, La Flora de Colombia 385

Sete gine. Venticas del "Dugua" [=Dagua], 600-700 m., Feb.
1906, H. Pittier, Tipo).

En la i ae original se sittia la localidad- eB en el
Departamente del Cauca. Para beneficio de los que no tien
mayor informaciodn acerca de la geografia con pobre ik fe) sé sus
divisiones paidepdecktagadaehaeses as, aclaro que hace mas de
cincuenta afios pertenece al Dégerodueats del Valle, opugeegeds
del Cauc

Couits orme lo advirtieron los mismos autores (en The
ceae 2: 55-56.1920) el ejemplar W.R. Maxon 3845, colecctonade en

Cc ece

tipica de pidmauiecews colombianus: COLOMBIA. - Valle: Dagua
Valley, open limestone valley below Dagua, 700- 900 m., me, May 1922,
Pennell 5412 (US), 5413(US); - Dagua Valley, Espinal, below
on 600-900 m., open arid slo opes, Sept. 1922, Killip & Hazen

7 (US). Hay en el herbario US cre ectoyraf ise de
prec altima planta en su nébitad natura

EUPHORBIACEAE

CHAMAESYCE MESEMBRYANTHEMIFOLIA (Jacq.) Dugand, comb. nov.
Euphorbia mesembrianthemifolia Jacquin, Enum. 22. 1760, et
Sel. Stirp. Amer. 150. 1763 ("Carthagenae in rupestribus mariti-
mis"); Gooding et al. Fl. Barbados 250. Fi
spporbia a teres Jacquin, vi Stirp. Amer.
ed. : 189, 0; - Dugand, Caldasia 2 (8): 295. 1943.

Eup foectreg buxifolia Lamarck, Enc. Meth. A271 tI86s =
Boissier in DC. Prodr. 15 (2): 15. 1862; Fawc. & Rendle Fl. Jam.
4: 336. 1920.

. Euphorbia litoralis Kunth in H. et B. Nov. Gen. & Sp. 2: 54.
817.
Chamaesyce buxifolia (Lam.) Small, Fl. SE. U.S. 711. 1903.

Hace mas de veinte anos (en Caldasia 2: 295. 1943) adelante
la tésis fae que el epfteto legftimo para esta espe
byanthentfolia = Jacquin por cuanto antecede a buxifolia de

que ambos se i ;

epee por © Botssier en 1862, aunque entonces no hizo v
la Prioridad del primero.
cien rrensaces en su Flora of Barbados (1965, 50),
Gooding, re y Proctor reconocen la sol tee Ae ‘al epiteto
me

de Jac
Sigu iendo yo el criterio de varios botanicos en lo que
respecta al genero Chamaesyce Rafin. 1817 - que otros no

386 PHYTOLOGIA Vol. 13, no. 6

reconocen, o tratan como subgenero de Euphorbia - propongo la
combinacidn nomenclatural que encabeza este articulo. Si bien

su obra, publicada aproximadamente en 1780. En efecto, derivase
de Mesembryanthemum L., que Linneo escribid cee Serene con

(aunque desde el punto de vista filoldgico lo correcto serfa con
i) y continuo escribiendo con y en sus obras posteriores
COLOMBIA. - Magdalena: Santa Mart re, H.H. Smith 1920
(US); San andeds (Isla): Dec. 1961, ‘Daniel 5634 (US
CURAZAO. - r Willemstadt, coral sea beach, March 1913, Brite
ton & Shafer “2933 (US); Rif, March 1952, Arnoldo 2075 (US

Adem: e visto numerosos ejemplares de la costa de Venezue-
la y de tae resw gee
meres es Kunth in H. et B. Nov. Gen. & Sp. 2: 75. 1817.

Hels usby, Descr. "300 N. Sp. S. Amer. Pl. 44.

1920 (plains a malay ast of Cienaga, Sept., H.H. Smith 368).

A tassicne de L. Croizat al pie del ee cae
de C. _clenagensis (herb. NY) (sefiala que el Tipo de C. fragil

en Paris tien puis mucho més ees efias. En nae caso - dic
Croizat - est timo es un ejemplar muy pobr

Vista la esate? & del Tipo de paste. " coleccionado por
Humboldt y Bonpland en Cumand, Venezuela, puedo equipararlo fac-
ilmente con ejemplares de idéntico a (Dugand 5660), de las
a semidridas al oeste del R —— cuyas hojas tienen

P e longitud y 1 a 2 cms. de ancho; igualmente
Haught 4155 de las acemiada de ie “Maeve en el Depto. del Mag-
dalena, y varios de la costa de Ven mados por
Foster Smith (No. 251) y el Hermano es Cons 2967, 2980,
3333). Los hay también intermedios como mi No. 5664 de la misma

Santa Marta, y finalmente los hay semejantes al Tipo de ciena~
gensis, tales como Dugand 5820 y 6323, de la misma localidad que
mi 5664. El 5820 de mi Seas es notablemente idéntico al
Tipo cienagensis

xtremo _90e de la variacidn en el tamafio foliar
pS en mi No. 6285, de la regidn del Cesare, Cafio seyaniets

es mi No. 5660 de una localidad muy cercana a la anterior;
este, las hojas no exceden de 2 a 3 cms. de Pacicee por ee de
a 1 cm

Los ejemplares de hoja pequefia de mi colecetén proceden
todos de plantas que crecfan al descubierto, en lugares csidas
lantas m

por la _— sombra que reina en el sotobosque subxerofiti
s los ejemplares mencionados peste hdllanse el herbariD
US y po de ellos en el COL.

CROTON GUILDINGII Grisebach, Fl. Brit. W. Ind. 38. 1859
roton bondaensis Rusby, Descr. 300 N. Sp. S. Amer. Pi. 43.

1920.

1966 Dugand, La Flora de Colombia 387

COLOMBIA. - Magdalena: 6 miles north of Bonda, 100 m., July,
H.H. Smith 1467 (ejemplar en flor, Tipo de C. bondaensis Rusby,
- También citado como Tipo hay un ejemplar con frutos, H.H.
Smith 1467, de Masinga, coleccionado en el mes de noviembre
(NY, US). Los dos ejemplares citados llevan el mismo ntmero

Santa te y muy cerca entre sf.

Otros ejemplares examinados: Atlantico: region de Bar-
ranquilla, llanada de Juanm mina, 10 m. alt., bosques subxero-
fiticos, Jul. 1962, Dugand 6101 (COL, US); - Alrededores de
Gala ~ y seacmers teks 1943, Dugand & Jaramillo 3292 (COL,US).

mi en tender no hay Silencers significante entre los
sjenplares antillanos (guildingii) y los de la costa colombiana
del Caribe (bondaensis) . Eggers 6953 de la isla St. Vincent, en
las pate Britdnicas, patria cldsica de C. guildingii
Griseb., tiene las St tio del apice peciolar sésiles (como
las describe Grisebach, loc. cit.) o muy brev a cet ip dermal
i fe) tamano de

Siendo estos tltimos los de hoja m4s grande. En la regién cos-
oe RRMA Ana este es un arbusto de 1.50 a 1.80 m.
altur

Pa pala NOBILIS L.f. var. HYPOMALACA (Standl.) Dugand,
comb.
Phyl lente nobilis var. hypomalacus rd Carnegie
Inst. Wash. Phub. 461: 68. 1936 (Tux pefa, Campeche, Mexico
llanthus antillanus var. Sprite ati (Stand1. ) aide :
pataioste Bs 3337-0-1939¢
Phyllanthus ote penorehhs Rusby, Descr. 300 N. Sp. S. Amer.
+ 42, ip (quoa one fructif. saltem, H.H. Smith 410).
De acuerdo con rady L. Webster (Journ. Arnold Arbor. 39:
208, 1958). has asin antillanus (A. Juss.) Muell.-Arg. (Cica
antillana A. Juss. 1824) y Ph. nobilis (L.f.) Muell.-Arg. (Mar-
n

Tario, no se abre en "cocas separadas, la cubierta seminal es
farnosa por fuera, cartilagfnea o huesuda por dentro, y el hilo
€s profundamente excavado.

e visto un isdétipo de la variedad Phyllanthus_ nobilis
hypomal cus de Fe cadiew, oriundo de Tuxpefia, Campeche, Mexico
Gundeli 897, Us), luego traneferido por Lundell a la especie
antillanus; y he examinado los ejemplares citados por Rusby en el

388 PHYTOLOGIA Vol. 13, no. 6

ree de je anthus heteromorpha. Uno de éstos (H.H. Smith
mn i es

la aa EE original es de miles south of Mamatoco, abou
400 feet, Apri s egun la queta en el ejemplar mismo

(Herb. de "dry thickets near Masinga, 250 feet, May 30"
Los dos lugares hallanse basta cerca uno del otro al pie de

designado como Tipo por Rusby. otro ejemplar citado en el
protdlogo (H.H. Smith 410), procedente de Masinga, Ju
tiene frutos y hojas adultas; de e 31 dice Rusby que "cree" que

a.
En rigor, el Tipo de este concepto es H. Smith 1716 por
En cuanto Bae ro (H.H. Smith 410)
aunq

os.
os ejemplares que he coleccionado en el norte de Colombia

son todos fructiferos, excepto uno estéril, e idénticos al citado
No. 410 fruct{fero d de H.H. Smith. Por la pubescencia menuda y

plares de Honduras, Dept. de a, colecc ados po .
Molina, cuyo No. 7014 y 7033 son masculino y de hoja menuda-
mente pubescente, mientras que 7035, femenino, es casi tan
glabro como los ejemplares genuinos de nobilis. Intermedio oe
este concepto es el Tipo de Phyllanthus heterot heteromorpha Rusby, re se
H.H. Smith 1716, aunque por su escasa pubescencia en el envés rk

(US); - Edges of dry forest, 6
co), about 400 feet, April, H.H. woecery oy toner de “Phyllanths
heteromorpha Rusby, DS). meac.ven flor;- Bosque sombreado marginal
del Arroyo Caraballo, al 0. de Fundacion, cerca de Retén, Mayo
1935, Dugand 822 en flor; - Valle del Rio Cesare (parte occiden-
tal), al oeste de Los Venados (10° N., wig W.), llanuras sub-
xerofiticas, Abril 1963, Dugand 6320 (COL, US) en frute.
a colombian del Caribe a arbolito se conoce

con ios nombres de pi py arito y perla. El Ultimo débese al
aspecto del conjunto seminal, que por su forma esferoide, y &
lustre y color blancuzco azulino de su cubierta exterior, tiene
alguna semejanza con una perla. Por cierto que Linneo hijo, al
describir a Margaritaria nobilis (el nombre del género se deriva

1966 Dugand, La Flora de Colombia 389

del latin Margarita, perla), hizo referencia al mismo hecho ("nu-
cleus... nitidissimus margaritae colore omnium in tota natura").

agape cece NOBILIS L. f£. var. NOBI
argar varia 2 nobilis bets; eae ye 428. 1781 (quoad pl.
foem. a
omparada con su variedad pypowslaca (ver el articulo an-
terior), hon ojas presentan en el enves una ténue eae
a

P
tacto, la variedad cite de esta especi
e hojas completamente glabras.

VENEZUELA. - Carabobo: between Nevedn and Urama, 1920, Pittier
8925 (US); Sucre: rn slopes of Cerro Imposible, 140-200

ferible a nobilis. Dicho ejemplar consiste en dos ramulos separ-
ados: uno con flores femeninas, otro con frutos.

iivag oa ELSIAE Urban, Repert. Sp. Nov. 15: 405. ws penne
on

COLOMBIA. - Atldntico: entre Palmar de Varela y Pon nei “ep
del rio Mag » Marzo 1932, Dugand of ve isma loca

Marzo 1933, Dugan (Y); - Confluenc e los tonto ns
Led

Ho
on, bosque marginal, Dic. 1963, D nea “6585 (US); misma locali-
zo S

(U:
a parte interior de la cccebacd de este arbol es de subido
Color rosado

LEGUMINOSAE-MIMOSOIDEAE

ALBIZIA GUACHAPELE (Kunth in H. & B.) Dugand, comb. nov
es Acacia guachapele Kunth in H. et B. Nov. Gen. & Sp. GO: 261.
23

Sa, Zathecollobium pret Bo Pittier, Contr .:0.S. Nat ; Hard
2464,

n US
Senanea_sananigua Pittier, Arb. y Arbust. peal 4-5 Decad:
94. 1925; Trab. Mus. Comerc. Venez. 90. 1
s+ it ae hee Britton & Rose ex Record in Trop. Woods
11: 14.. 1927s. Woodson. & Schery Fl. Panama in Ann. Mo Bot. Gard.

390 PHYTOLOGIA Vol. 13, no. 6

ays oe 1950.
seudosamanea guachapele Harms, Notizbl. 11: 54. 1930;
Hh [ pois p, Ann. N.Y. Acad. Sci. 35: 130. 1936.
C rando la fotografia del Tipo de Acacia guachapele
(Humb. & sotahs 3805, Herb. Mus. Paris) y un ejemplar de los al-
rededores de Guayaquil, tipico de marae especie (L. Mille 33, (US),
ademds de uno de la Provincia de — Rios, Quevedo revi cate'! 6537,

una séla margen, en vez de ambas como ocurre en las cites Al-
bizias, no me parece suficientemente importante en este grupo
para brs a guachapele ¢ como género distinto. Puede constituir
cuando mds una Secc
lo sefalo i. pet Little (Amer. Midland Nat. 33: 510.
1945) la graffa original emple da por Durazzini (Mag. Toscan. a.
ar de

tr i
Z para "tavietaatl digdmoslo asi, el apellido Albizzi. En este
caso es preciso conservar la grafia original Albizia conforme al
Art. 73 del Cédigo, pues no se trata de un simple error ortogré-

consideracion fin e suma importan , a mi modo de ver.

1 geet conveniente Gapens para el Cédigo de No-
menclatura una regla que ertos casos permita estabilizar
oficialmente la graffa “a gieotie nombres de género bien i
dos, consagrando la co br mn inveterada de escribirlos,
sobre todo si ella es correcta ortogrdficamente, y te endo en
cuenta el método actual a formacidn de los nombres de gén-

Pp
ero (ver Art. 73, Recomendacién 73-B, p4rrafo (a) para el caso
de aia vs. Albizia
a Albizia auachepele héllase distribufda por las regiones
Pe prota ice ate desde Chiapas, en México (Matuda 4021,
ic aate 6008) y la Sleenas Central hasta la parte nortefia de Sur-
américa (Colombia, Venezuela, occidente del Ecuador) y se encu”

L
que se anotan en Colombia son los siguientes: "tabaca" y "sana

1966 Dugand, La Flora de Colombia 391

guaro" en el Atldntico y norte de Boltvar; “"masaguaro" en Arauca;
igu

e
amarillo", corruptela uatcnde ten de "igud amarillo'

CHLOROLEUCON MANGENSE (Jacq.) Britton & Rose, N. Am. Fl. 23: 38.
gga

Sa mangensis Jacquin, Enum. 34. 1760; Sel. Stirp. 267.
1763 i cng Manga, Cartagena, Colombia
e Sp

SAG1 1829
Sachacsinbick mangense (sabe: ) iaebeiae: Contr. Gray Herb.
‘ 3. 1919; - Woodson et Shery, Ann. Mo. Bot. Gard. 37: 241.
Albizzia marthae Britton & Killip, Ann. N.Y. Acad. Sci.
as (Colombia, Magdalena: Santa Marta, H.H. Smith 296, Tipo
nm US).

A la lista de sindénimos ya enumerados por Britton y Killip

nada difiere de los ejemplares florecido e hojas tiernas de
Chloroleucon mangense (por ejemplo, Dugand 4620 y 5004, Elia
186, Haught 4117 y H.H mith : del norte de Colombia).

ro s.n., Santa Marta, Colombia), cuya fotograffa he visto. Es
a pesar de es iguales los epftetos respectivos,

@
HH
t=]
»
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COLOMBIA. Magdalena: Bavders s/n., 1822 (foto 6971 Herb. De-
lessert); Santa Marta, H.H. Smith 2064 y 2065 (US); Dry forest
Rear Fonseca, 175 m., April 1944, Haught 4117 (US) Atldntico:
Vicinity of Barranquilla, Mayo 1927, Elias 186 (US); - Manatf,
Dugand isn (US); entre Palmar de Varela y Ponedera, Dugan 7a
4182, 4438, 4620 (US) ; Malambo, Dugand 5004 (US); carretera
Puerto Colombia, kildmetros 5 a 6, Dugand 6569, 6793 y 6938 (sy.
Bolfvar: north of Arjona, Nov. 1926, Killip & A.C. Smith 14494
(US); Cerro de la Popa, Cartagena, Barkley & Araque Molina 19
Bo-044 (US).

CUADOR: El Oro, 6 nn aa eo of Arenillas, June ge,

- Numerosos ejenplares de Chloroleucon mangense he ex-
aminado en el herbario US, idénticos a los de Colombia. En su

flora of Panama,Woods on y Schery sdlo citan uno de aquel pais
(Veraguas: Santiago, ich 1082).

El nombre vulgar més usual para este drb Shep eas nt en
el norte de Colombia (Deptos. del Atlantico, Naga na y norte

de Bolfvar). Tambien lo llaman "hoyo de zorra" en spr lu-
8ares del Atldntico.

392 PHYTOLOGIA Vol. 13, no. 6

que dich lldmase en realidad Manga. H411 en 5 a

interior de Cartagena, y un angosto canal la separa la abe

firme, a la cual esta unida por varios puentes. Hoy totalmente
la

urbanizada constituye un importante barrio renidenciad de
ciudad de Cartagena.

ENTEROLOBIUM CYCLOCARPUM (Jacq.) Grisebach, Fl. Brit. W. Ind.
226. 1860.
ee as Jacquin, Fragm. Bot. 30. 1801.
2? i Britton & Killip. Ann. N.Y. Acad. Sci.

35: ar “1936 Site vicinity of we aayilia, Atldntico,
Colombia, Elias 499 en

amen del Tipo. de Albizzia longipes me ha convencido de
que no se trata de nada distinto a la conocidfsima especie cuyo
nombre encabeza este art{fculo y que es tan comtin an las regiones
cdlidas neoknontas} 8 desde el sur de México hasta el norte de
auEane a.

s vulgares:"carito" y "orejero"” en la costa colombi-
ana me Goctbe y el Bajo Magdalena.

SENEGALIA ELIASIANA Britton & Killip, Ann. N.Y. Acad. Aci. 35:
145. 1936 (Colombia. - Atldntico: Baranoa, Elias 602, Tipo en
Us).

Acacia Eliasana (Br. & Kill.) Standley, Trop. Woods 52: 2).

193
En clave que Britton y Killip (op. cit. pp. 141-142) dan
para las ak de Senegalia en Colombia, clave que por ciert

adolece de algunas deficiencias, colocan a S. affinis y S. elia
siana una al lado de la otra (Nos. 11 y 12) aneaceanaen eee por
‘el ancho de los folfolos ("about 1 1/2 mm." en affinis; “about
2 ae mm." en eliasiana) y por el caracter de la pubescencie®
el envés de los mismos ("loosely pilose" en affinis; "appresse
pubescent” en eliasiana).

He examinado el tipo de eliasiana (herb. US) y un fragment
eodeataun del de affinis en el mismo herbario, compar4n dolos aa

en
ed-

abra.
Falta pany’ mejores ejemplares de la supuesta especie at”

lados de la Zona Bananera pues fue en Guamachito donde el

lo entonces podra definirse si es sindnima de S. eliasiana-

1966 Dugand, La Flora de Colombia 393

COLOMBIA. - ALeatice: Baranoa, Nov. 1928, Elias 602 (Tipo de

S. eliasiana, US); - Baranoa, en la carrete era a Usiacurf, 130 Ms,
Oct 19.36; ear 1055 (US); misma region, 100 m., Sept. 1937,
Dugand 1122 (US); - Alrededores de Galapa y Baranoa, Jul. 1943,

and & Jaramillo 3441 (COL, US); - E aranoa y Galapa, 80-
100 m., Dic. 0, Dugand 5400 (US); misma localidad, 100 m.
Abril 1961, Dugand 5646 650 (US); misma loc m., Sept

Ene. 1962, Dugand 6048 y 6053 (US); cerca de Ponedera, Sept.
1963, Dugand 6456 (US).

Ademas, en mi herbario particular: Atlantico: Entre Bara-
hoa y Galapa, 80-100 m., Dic. 1960, Dugand 5401, 5402, 5403.

didas o arqueadas hacia abajo y recués stanse a menudo sobre otros
le

omplem ar la des cripcidn original un tanto defici-
ente “6 STE arse iana diré que el pecfolo presenta una o
dos glandulas, y ademd4s hay otras gldndulas mas o menos conspi-

sdlo de 5 7 (Dugand 5646, 5940) o ia 12 a 14 (Dugand 3441,
5780, 6053) ; los folfolos son de 15 a 40 en cada lado de la ra-

La legumbre madura, ya al punto nage Re erpgang suele alcanzar a
veces de 15 a 18 cm. por 2.2 a 2.4 cm. (Dugand d 5403, 5963, 6048,

6053) y puede ser, como ya lo dije antes, mds o menos oubdérula °
totalmente glabra

THEOPHRASTACEAE

JACQUINIA L., Diss. Fl. Jam. App. 27. 1759 (Jaquinia) ;

Acad. 5: 388. 1760 (Jaquinia); Sp. Pl. ed. 2: 271. oie sii)
Jacquinia Jacquin, Enum. Pl. Carib. 2 et 15. 1760; Sel. Stirp.
Amer. Hist. 53. 1763; Linn. Gen. Pl. ed. 6: 101. 1764; Syst. Nat.
ed. 12: 175. 1767.

a lo senald E. L. Little (Amer. Midl. Nat. 33: 510

1945) la graffa original de este re es Jaquinia. Asi aparece,
en ecto, en d s disertaciones de las enidades Aca

u
Yam raeside D.D. Car. Linneo roposuit Carolus Gust. Sand-

394 PHYTOLOGIA Vol. 13, no. 6

mark, Yermelandus", publicada en Upsala el 13 de diciembre de
1759. La Ultima pdgina de dicha disertacion. o sea la p. 388 del

de Jacquin. Por lo tanto e oO.
omenzando por Jacquin (Enum. Pl. Carib. 15. 1760) ha sido
e a

a a mencionada Flora Jamaicensis. azon ha sido expli-
cada por William Thomas Stearn en 1 rte introductoria de
reimpresion facsimil del Species Plantarum d mneo, ed.

y no a los estudiantes o graduandos que
parecen como autores de ellas. En efecto - sigue explicando el

responsables de lo que en ellas se trataba, siempre que actuaran
omo presidentes en el debate piblico que se acostumb cele-
brar en las universidades para presentarlas. actuacion de

sis a La
graduando se limitapa a pagar de su peculio la impresidn de la

debate, Para mayores detalles constiltese a Stearn. loc. cit.

Amoenitates Academicae
vol. 5, anuncia q voltmen est4n reunidas varias di-
sertaciones sobre Fisica, Medicina y Botdnica publicadas separa
ente con anterioridad, Flora Jamaicensis sal a la lu
piblica el 13 de diciembre de 1759, fecha que figura en su propia
pagina titular, mientr ue el mencionado voltimen no f 1

cado antes del 4 de agosto de 1760, pues ésta es la fecha que
aparece en la pdgina dedicatoria.

Con respecto a la ortograffa de este nombre genérico, si se
aplica sin reservas el Art. 73 del Cdédigo de Nomenclatura debie-
ra escribirse Jaquinia y no Jacquinia, pues la primera eS 1

orma que aparece en la publicacidén original. Linneo también
escribid Jaquinia en la segunda edicién de su Species Plantaru™
- 1762); empero - de suma importancia - en 1

La
o>

1966 Dugand, La Flora de Colombia 395

ona pagal por los botdnicos, con rar{fsimas excepciones sg

ejem los autores de la reciente Flora of Barbados, p. 320
1965, ceoplean Jaquinia).
Como ya 1

manifesté algunas paginas atrds, en ocasidn de

acqui Jaquini Al contrario
del caso Albizia sineeaiminie: PE en aot de Jaquinia
trdtase, a mi modo de ver, de un simple error ortogrdfico.
Prueba de ello es que Linneo mismo enmendé la graffa adoptando
la forma Jacquinia desde 1764 en adelante; lo cual equivale a
una enmienda ortogra4fica de las que permite el Art. 73 en su
primer es rafo.

eable y de esperarse que en un prdéximo Congreso In-
ae Botdnica se resuelve modificar el Art. 73 del

encia sy todos los botdnicos y mayor estabilidad de la nomen-
clatu

r lo pronto, funddndome en que la graffa —* Jaqui-
nia no i ae a una latinizacidén intencional no error
ie deh continio empleando la forma pradteisald peusitise

a por Jacquin (1760 y 1763) y aceptada por Linneo
(1764 en adelante).

JACQUINIA ARISTATA seewarern Fragm. Bot. 64, t. 95, f. 2. 1809
(Venezuela:

uinia caracasana Kunth in H. & B. Nov. Gen. Sp. 3:
251. 1818 neon litoral drido entre La Guayra ; Cabo
— Humb. & 1., Tipo en herb. Mus. Paris; foto en US).

acquinia ici ien ch Jacq., Mez in Engl. Pflanzenreich 4,
236-43 40. 1903

Jacquinia genioulate Mez, loc. cit. 38 ("Venezuela"sphalm.
= Colombia: an der Miindung des Rio Magdalena bei Sabanilla; -
Karsten, Herb. n).

Jacquinia gracilis Mez, loc. cit. 39 (Colombia: "in der
Sierra de Sta. Marta" (2?) - Funck No. 334, Herb. Deless.; foto
del tipo en U ——

Jesuints oreleroite Mez, loc. cit. 41 (Venezuela: La
a, “ = sten, Herb. Wien
nia aciculata Mez din Urban Sumb. Antill. 2: 450.
1901 Tie classicus certus haud manifestus; cum magna autem

396 PHYTOLOGIA Vol. 13, no. 6

probabilitate in litore colombiensi maris Caribaei situs).
Jacquinia aurantiaca auct. plur.

Que yo sepa, hasta ahora no se habia aclarado - inexplica-
blemente por cierto - cual es la patria cldsica de la Jacquinia

claramente que fue trafda de Caracas ! onviene aclarar de una
vez que esta indicacién de localidad no significa tanto la ciudad
como la antigua provincia de Caracas, que en aquel tiempo (antes
de la independencia de Venezuela) comprendfa por lo menos los
Estados actuales de Aragua y Miranda adema's del Distrito Feder-
al, cuya cabecera es la ciudad de Caracas.

La descripcidn de Jacquin, ademés de la excelente ilustra-
cién en colores naturales (plancha No. 95) que la acompana, no
ofrece duda acerca de que la Jacquinia aristata es la misma

Pittier 12425 y Curran & Haman 935, coleccionados muy cerca
localidad-tipo de J. ovalifolia; y una buena parte de las de

e

mm. ) vez muy jévenes, son del Estado Lara, en Venezual

(Saer 66 y Ginés 4400, US), pero sus flores son jdénticas a las

de todos los demds de ese pais y de Colombia, inclusive Elias

1163, de Puerto Colombia, que es el de hojas mayores que he visto

(hasta 80 mm. x 25 mm.).

J. aristata la inflorescencia es fundamentalmente rac
¢ nd

i
m
mosa, pero a veces tiene aspecto de o los pedicelos
y estos
{qu

tH
=|

los en longitud. Y el estilo es siempre evidente, corto
tivamente alargado, pese a que Mez lo vio muy breve en aciculat®

1966 Dugand, La Flora de Colombia 397

y casi nulo en caracasana.
La flor antes de abrirse tiene los pétalos verdoso-amaril-
lentos; al desplegarse la corola y el conjunto de estaminodios

i aesc
agradables que conozco destdcase el suave olor de las flores de

Jacquinia aristata.

Con relacidén a la J. geniculata Mez, no he visto el tt ae ni
foto de #1, pero he observado numerosos ejemplares Si vaake
(y coleccionado algunos) en la propia localidad-tipo eebitala por

quis de la inflorescencia de su J. geniculata no n comin
sino rarisf{fmo en los ejemplares de la regidn de Sabanilla, Sal-
ar, Puerto Colombia, Barranquilla » © sea la parte septen-

fio Por jlo t
del Pin de la inflorescencia es cardcter cuando md4s puramente
SpE hae

que atane a la J. gracilis a! el ejemplar represen-
t hah en ats "fotografia del Tipo (Bune 334, herb. pereonise) es,
para todo objeto prdctico, igua los d eada de J. aristata. La

e

"Caracas", 0 détulo adherido a la cartulina en 190

"Caracas (Amérique Méridionale)". Sa s que re los muchos
lugares visitados por Nicolds Funck en Venezuela y b

embarcdse hacia Hambu urgo a fines del mismo affo. Es probable que
llevara consigo los ejemplares de sus colecciones de fines de

398 PHYTOLOGIA Vol. 13, no. 6

1843, que entrego en Europa en enero de 1844. Esto refuerza mi
tesis de que el Tipo de gracilis procede de Colombia y no de

Ahora bien, si Funck colecciond su ejemplar en la region de

Sierra Nevada.
No es posible apreciar en la foto del Tipo de J. gracilis el
anteras que, se un Mez, tienen el conectivo pro-

caracter de las an

longado en acumen m la de los ldculos o tecas, diferencian-
dose asi - siemp gun Mez - del grupo en q cho or
coloca a J. aristata, ovalifolia, aciculata,caracasana y_aura
tiaca, las cuale en "acuminado" el conectivo, o lo tienen
escasam Be 4" , pues, mas cuantitativa que cualitativa.
El dpice del conectivo en los numerosos ejemplares (vivos y de
herbario) de J. aristata que he examinado es casi s re agud

y proléngase un poco m4s alld de las tecas; pero en muchas flores
de edad algo avanzada aparece mds o menos tronchado, como mordi-
do, quizas por insectos.

Con respecto a la J. aciculata Mez, la fotografia del Tipo

alguna, o que atane a l e inflorescencias,

ejemplares de Colombia y Venezuela que aqui adscri J. a
tata Jacq. La patria cldsica de esta aciculata es también dudo-

: lo nada d y sélo sabemos Julius von Rohr

colecciond plantas para Martin Vahl en el afio de 1786 en Jamaica,
Puer e Ant s menores, asi n las is-
las subcontinentales de Trinidad y Curazao junto a la costa de la
Ameri ; este continente en Su , Cayena, Santa
Marta y Ca na. Por lo tanto, vista la similitud del Tipo de

timos colombianos subrayados arriba. En ambas localidades se
encuentra la J. aristata, aunque mds abundante y f4cil de encon-

26 J. aciculata. Este fue coleccionado seguramente POF |
Carlo Giuseppe Bertero en el litoral norteno de Colombia, quizas
en Santa Marta pues él visitdé esa ciu en diciembre de 1820.

Me an ostener esta tesis el hecho de que un nimero cercano
de su coleccidn, el 2612 (Serjania curassavica, citado por Rad1-
kofer en Pflanzenr. 98a, IV. 165, b 1: 203. 1933), es de Bar-
ranquilla, en cuyos alrededores colecciond Bertero en octub
1820. All{ se encuentra tambien la J. aristata Jacq.

Jacquinia aurantiaca auct. - Algunos ejemplares de Colombia y
Venezuela, de los que examin¢é con motivo de este estudio en el

1966 Dugand, La Flora de Colombia 399

herbario US, llevan rétulo de J. aurantiaca Aiton. Gran numero

del litoral colombo-venezolano; pero el tiempo me falto
estudiarlos a espacio.

4

son En cuanto
J. aurantiaca se dio originainente como “ipxae Sandwich" (=Ha-
waii), lo cual es sin duda errdne

acquinia aristata Jacq. es por lo general un arbusto is ar>
Seas de 2 a 3m. de altura, a veces d4rbol pequefio de 4 a 6 nm.

ne
40 cms. de aidnetic en los brioies viejos, pero por lo general
no excede de unos 20 cm 8.3 y cuando es ar wini = = = abies
cipales tienen cuando mas unos 10 c
muy osa; las ramas rfgidas. Es “earacterfatico ce 7 taccion
na

ta, en las llanuras del Rio Cesare.

Nombres vulgares. - En la costa de Colombia: "barbasco",
"barbasco e pia", "sarnfcalo", "sarniclo", "sarnisclo". En
Venezuela: "trompito" y "trompillo"

VENEZUELA. - Anzodtegui: valley of Puerto La Cruz, Pittier 9204
(US), Aragua: Ocumare de la Costa, Ll. Williams 12185 (US). Ca-
rabobo; Puerto Cabello, Rose 21857 (US). Distrito Federal: Ca-
bo Blanco, Curran & Haman 935 (US); Pittier 12425 y 14411 (US);
between Caracas and La Guayra, Rose 21721 (US). Falcén: Para-
guand, Adfcor ra, Tamayo 928 (US). Lara: Bobare, 600 m., Zahn 185
(US); Bar uisimeto, Saer 66 (US )3 between Rio Tocuyo jae Carora,
Pittier 12611 (US); Carora, Ginés 4400 (US); sin loc. Delgado 95
US). Sucre: Cumanacoa, Netting 22 (US). Zulia: Isl Sa
Carlos, Curran & Haman 786 (US). Estado ?: sin loc. Curran &
Haman 421, 455, 470 (US).
COLOMBIA. - Atldntico: Barranquilla, Elias 945 y 975 (US); Puer-
to Colombia, Elias 1051, 1162, 1163 (US); Sabanilla, Dugand 5213

oe
ya, Cuatrecasas 34298 (US); entre Barranquilla y Galapa, Dugand
135 (Y); Santo Tomds, Paul B-8 (US); entre Palmar de Varela y
Ponedera, Bar ah nd 352 (Y¥), 4603 (COL, US). Bolfvar: vicinity of
Cartagena, Heriberto 279 (US). Guajira: “Riohacha, Sinclair s.n.
(US); Cuatrecasas 25453 y 25460 (US); Fonseca, Haught 3991 “ieee
Magdalena: Santa Marta, H.H. Smith 413 (US); Dawe 487 (US

4,00 PHYTOLOGIA Vol. 13, no. 6

Curran 384 (GH, US); Killip & A.C. Smith eggs —_— Sp st
Molina & Barkley 18-Mag 090 (US); playa de Con a de San
ae Dugand 5894 (US); Donjaca, east of Sa oh bi aad om

,» GH); valle del Rio Cesare, Los aE wees 5586 (US);
ae Marthe 4 Cerro San Antonio, col. ?

zco 0s estimado — grater Aristeguieta,
Encargado del Instituto a s de Venezuela, en acas, la
da muy valiosa resto

J. aciculata y J. geniculata,

adscribirse a Jacquinia aristata Jac

Notas sobre la flora de Colombia y paises vecinos.

English Summary

rst rib a series of short papers concerning main-
ly littl e known, new or otherwise interesting elements
of the flora of the Caribbean coastal plain and the
lower Magdalena valley in northern Colombia. In seven
ral instances species from other sections of Colombia
and neighboring areas in Venezuela and Panama, as well
as other countries in South and inchs tee America and
the Antillean islands are mentio

Ee hes dirs ~e agate: th. yoBy ls micracsn viet oscce-
reus mare ianus eck "B & Rowl.); EUPHOR- :

BIACEAE (Chamaesyce nielabe cea ao (Jacq.) Duges
orotee “tre ilis tan th; = oton SE Ti Griseb.; Mar-

era eee a Le et omalaca

H SU INUSARCRINOROED Albizia

guschapele (k (kunt Dugand; Chloroleucon mangense (J~
Jacq.) Br. & Rose; Enterolobium cyc locarpum (racer?
Griseb.; Senegalia eliwat ite Br. & Riiiip), and THEO-
PHRASTACEAE (a discussion of the correct spelling of

Jacquinia vs. Jaquinia, and an evaluation of the lit-
le known J. : Ta

=a stata Jacq. 1809, of which several sy~
nonyms are

A. Dugand

ADDITIONAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS CALLICARPA. I
Harold N,. Moldenke

This is the thirtieth in my series of works of monographic nature
on the genera of Verbenaceae, Avicenniaceae, Stilbaceae, Chloantha-
ceae, and Symphoremaceae. Previous genera so treated by me are
Acantholippia Griseb., Aegiphila Jacq., Amasonia L. f., Avicennia
L., Baillonia Bocq., Bouchea Cham., Casselia Nees & Mart., Castelia
Cav., Chascanum E. Mey., Citharexylum B. Juss., Cornutia Plum., Di-
ostea Miers, Dipyrena Hook., Hierobotana Briq., Lippia Houst.,
Parodi anthus Troncoso, Petitia Jacq., Petrea Houst., Priva Adans.,
Pseudocarpidium Milisp. : , Recordia Moldenke, | Rehdera Moldenke ‘.
Rhaphithamnus Miers, Stylodon Raf Raf., Svensonia Moldenke, Tectona L.
f., Verbena [Dorst. ] L., and Vitex Tourn. The N ew World and cul-
* vated members of the genus Callicarpa were also treated by me in

his fashion in Fedde, Repert. Spec. Nov. 39: 288--317 and 40: 38—
1 a £1996) In order to conserve space, material recorded in this

evious work is not repeated here except where it needs addition

Pas emendation

The list ot abbreviations employed by me in the citation o

as given in Phytologia 5: 15l--159 (1955) and added to
he

a
eleven years. My reasons for employing these short abbreviations,
rather oe the often very much longer ones recommended by the
international committee, are stated in the 1955 work and in pre-
Vious publica secmae As of this ccmniai. formal-
ly eg aah 3 herbarium specimens from "28h institutional and

ie | es merger Herbarium, Harvard University, Cambridge,
Massachusetts

Aa = H. Ahles Herbarium, University of North Carolina, Chapel Hill,
North Carolina

Ac = Botanisk Institut, Aarhus Universitet, Aarhus, Denmark

Ad = Universidad de los Andes, Mérida, Venezuela

Af = National Herbarium, Pretoria, Transvaal, $ outh Africa

Ag = Carlos Mufios Herbariun, Ministerio de Agricultura, Santiago,
Chile

Ah = J.C, arthur Herbarium, Purdue University, Lafayette, Indiana
Ak = Bit = Hancock Foundation, University of Southern California,
a. s Angeles, California ks
New York State Museum, Albany,
Am = Southwest Research Station, larscid Museum of Natural History,
Portal, Arizona
An = Institut Francais d'Afrique oe Dakar, Sene
Ar = United States es oo Arboretum, Washington, D. C.
As = C, F, Asenjo Herbarium, University of Wisconsin, Madison, Wis-
4,01

onsin
Au = Gniversicy of Texas, Austin, Texas
Av = Facultad de Agron nomfa ay Vet eterinaria, Universidad de Buenos
tina

Aire
B = Botanisches Museum und Garten, Berlin-Dahlem, German

dos

Be = Barnard College Herbarium, New York Botanical yt ~~
York City

Bd = Herbarium Bradeanum, Rio de Janeiro, Brazil

Be = Instituto Agronomico de Norte, Bel Braz

Bf = Instituto de Bot4nica y Farmacologia, Buenos Aires, Argen-
tina

Bg = Bergens Museum, Bergen » Norway
Bh = Instituto Agronomico do Servico Publica de Estado, Belo Hor-
zonte, il
Bi = Bernice P, Bishop Museun, ag “70-0
ig 32 Bajo? TePBicing » Jamaica
Fores partmen = ance Thatta
= = University of Colorado, Bouldes, Colorado
Bm = British Museum of N Natural History, London, England
Bn = Central College, reage ore, Mysore, India
= = Instituto de la Sall , Bogoté, Colombia

ndiana
Bu = W. M, Buowel] Herbarium, ahvanedtey sof stsent:, sored: Gabitins

By = Bureau of Plant Industry Station, wey Maryland
By = ie ag uaaeey Brooklyn, New York Cit;

Bz = Herbarium Bogoriense, Buitenzo org, Java, Todonetis
c= — University Herbarium, New York Botanical Garden,
ew York C

Ca = betounctte ae California, Berkeley, California
Cb = apap ay Herbarium, Conservatoire et Jardin Botaniques, Gen-
‘va, Switzer]
Ce = inteteds Se shag Colorado Acti s Colorado
Cd = Museo de Cérdoba » Cérdoba, Argen
Ch = Carey Herbarium, "Royal Botanic Bitos: Kew, England
Ci = Escuela Lgl Agricultura Tropical, "cali, Colombia
Cl = Royal Bo tanic Gardens, Calcutta, India
t

» Ohio
Co = North Appalachian Experimental Watershed, Coshocton, Ohio
Cp = Universitetets —— rote, Copenhagen, Denmark

Sansalito, Califo

rnia
Cs = Department of Biology, Lotte teed and Mechanical College
of Texas, College § tation, Texas
Ct = "Mine 9 Herbarium, University of Capetown, Capetown, South

» Cambridge University pomeeee England
oe i Gent's Zone Biological Area, Barro Colorado Island, Canal Zone

1966 Moldenke, Monograph of Callicarpa 403

D = Academy of Natural at nt Philade use nb ys Sg
Da = United States Department of Agriculture » Washington, D.C.
De = De Candolle Herbarium, Conservatoire et jardin A th
Geneva, Switzerland
= Botanic Gardens » Dehra Dun, Uttar Pradesh, India
= Delzie Demaree Herbarium, Monticello, Arkansas
Ot

WORER SH
cs
p
oo
i
Na}
F
Lax)
g
:

Germany
Dt = eee Herbarium, Dartmouth College, Hanover, New Hampshire
ey Herbariun, Stanford University, Stanford, California
Botany Department, College of Agriculture, Davis , California
= Missouri Botanic. al G. Garden, Saint Louis, Missouri
= Earlham College, Earlham, Indi
= Universidad Nacional de Loja, Loja, Ecuador
= Ro c Garden, » Scotland
= José 088 Bugenio Leite Herbarium, Novo 6 iste Rio de Janeiro,

mak Herbarium, New rpg lind Louisiana
Er = Palynologiska Laboratoriet, Bromma, Sweden
Es = pi sage ete ntal Agronomica, Santiago de las Vegas, Ha-

= Brik Wall # arti heng Stockholm, Sweden
r = Chicago Natural History Museum, Chicago, Illinois
Fe = Colorado Agricultural and Mechanical College, Fort Collins,
Colorado
Fg = Arizona State College, Flagstaff, Arizona
Fj = Fred B. Jones Herbarium, Corpus Christi, Texas
Fl = University of Florida, Gainesville, Florida
iy tad Nacional de Agronomia, Universidad Nacional, Medel-

lin, Colombia
Fo = Instituto Botanico della Universita, Florence, Italy
Fs = — Shreve Herbariun, University of Arizona, Tucson, Aré-

Fx = baeetta E. Fox Herbarium, Reto neess Louisiana
Fy = _Untversity of Arkansas, Fayetteville, Arkansas
G = Gray Herbarium, Harvard University, Cambridge, Massachusetts
Ga = iaahien Agricultural Experiment station, Experiment » Georgia
Ge = H. S. Gentry Herbarium, Tucson, Arizo
Gg = California Academy of Sciences, San nee ES California
Gl = Museu Goeldi, Belém, Par4,
h

, GUteborg, Sweden
Gp = Ontario Agricultural College, Guelph, Ontario, Canada
Gt = Botanische Anstalten
Gu = University of Georgia, Athens, gn
H = Duke University, Durham, Seah came
Ha = Colegio de la Salle, Vedado, , Cuba
HD = H, Bassler Herbarium, New oa ahtenionl Garden, New York
City

40h PHYTOLOGIA Vol. 13, no. 6

He = W. G. Herter Herbarium, Montevideo, Uruguay

Hi = University of North Carolina, Chapel Hill, North Carolina

Hk = University of Helsinki, Hels

Hn = Herbario Nacional Co olombiano, Instituto Ciencias Naturales,
gates Colombia

Hp = H, Hapeman Herbarium, Minden, Nebrask

Hq = Tasti tuto Botanico Dr. Julio eewtepins , Coimbra, Portugal

Hr = H, Httrlimann Herbarium, CIBA, Basel, Switzer.

Hs = Crispus Attucks High School, Indianapolis,

Hu = Sam Houston State Teachers College a Huntsville , ‘eves

Hv = Academia de Ciencias, Havana, nae

Hw = Howard University, Was n, D

FS aes Herbarium, Catholic Gavaratty of America, Washing~

» DC.
Ib = Firs Pate de Botanica e Agricola del I. N. T. Ae, Buenos
tina

Aires
Id = University of Idaho, Southern Branch, Pocatello, Idaho
Ih = Instituto de Historia Natural, Curitiba, Parand, Brazil

Je = Museu Nacional, Rio de Janeiro, Guanabara, Brazil
Jc = J. Cuatrecasas’ Herbarium, Bensenville, Illinois
Je = William Jewell College, Liberty, Miss
Jn = Aage Bohus-Jensen Herbarium » Tymebye, Denmark
Jr = Hebrew University, Je erusalem, Isra
Jz = J. de J, Jiminéz Herbarium, bedetaco de los Caballeros,
Dominican Republic
K = Royal Botanic Gardens
Ka = Kansas State College, pet Mery , Kans
Ke = Kern pile ha Pe ennsylvania State acieee, State College,

Kg = Desc aeits of Agriculture, Kagoshima University, Kagoshima,
Ki = te Bis c Herbarium, University of Rochester, Rochester,
rk

Ko = Botanisches bgrtwedt Universit#t Kiln, Kin, Germany
Kr = B. A. Krukof. ff Herbarium, Smi Smithtown, New York

Ku = Eberhard Kausel Herbarium, Santiago, Chile

Ky = University of Kentucky, Lexington, Kentucky

L = Jardin Botanique Principal, Leningrad, U.S. S.R

La = University of California at L. A., Los Angeles, California
Lb = University of Missouri, Columbia, Missouri

ld =C.L. 11 Herbariun, apes T

1966 Moldenke, Monograph of Callicarpa 40s

New York City
li = Colegio Salesiano, Lima, Peru
Ll = Lloyd Li pow h Cineinna nnati, Ohio
Im = Los Angeles County Museum, Los Angeles, California
Lo = Gaulterio Looser Herbarium, Santiago, Chil
= Linnean Herbarium, Linnean Society, London, England
Lu = Botanisk Museum, University of Lund, Lund, Sweden
Iw = University of Kansas, ence, Kansas
Lz = A. Lutz Herbarium, Universidade do Brasil, Rio de Janeiro,
Guanabara, Brazil
M = Meisner Herbarium, New York Botanical Garden, New York City
Ma = Forest De Department of Malaya, Kepong, Selangor, Malaya
= Melbourne Botanic Garden, Melbourne, Victoria, Australia
Mc = J, ea hire Herbarium, Sebring, Flori

= University of Maryland, College Park, is a
Me = Engtitute de Bielogia, Universidad natonad de México, Mexi-
co — —_

College, Bangalore,
Mg = a Botanical Garden, Mo ntrenl, aliases Canada
Mh = Matuda Herbarium, Mexico City, M
Mi = ~ University of Michigan ic chigan
- C. MacKeever Herbarium, Mount, Vernon, New York
0 = = Instituto Miguel Lillo, Tucum4n, Argen entina
Mm = McGill University, se sig Quebec, Canada
Mn = University of Montana, Missoula, Montana
Mo = Morris Arboretum, Philadelphia, * Pennsylvania
Mp = Museo Paranatnse, Curitiba, P , Braz
Mr = Morehead State College, Morehead, Kentucky
Ms = University of Massachusetts, Amherst, Massachusetts
Mt = ee Thais Herbarium, St. Mary's High School, Perth Amboy,
ew J

ersey
Mu = ae ches Museum, Munich, Germany
Mv = Marie-Victorin Herbariun, Yontreal Botanical Garden, Montreal,

Que
N = Britton Herbarium, New York Botanical Garden, New York City
Na = Natal Government Herbarium, Durban, Natal, h Africa
Nb = Stephen F. Austin State College, Nacogdoches, peas
Ne = State Normal College, Natchitoches, Louisiana
~s ~ Notre Dame —— Notre crs Indiana

ew G

O- = Unbtersity. of Tennessee puesta Tennessee
Oa = Oakes Ames Economic Herbariun, Botanical Museum, Harvard Uni-

06 PHYTOLOGIA Vol. 13, no. 6

Or = Oregon State College, Corvallis, Oregon
Os = Osborn Botanical Laboratory, Yale University, New Haven,
ecticut

Ot = National Herbarium of cheers Ottawa, Ontario, Canada

Ox = Oxford University, 0 ;

P = Muséum National + <A anete Naturelle, Paris, France

Pa = College of Pharmacy Herbarium, New York Botanical Garden,
New York City

Pb = R. Probst Herbarium, Langendorf, Switzerland

Pe = Philip Cheitman Herbarium, New York City

Pe = Museo de Historia Natural, Universidad Nacional Mayor de San
Marcos, Lima, Peru

Pf = Centro de Pesquisas Florestais e Conservac&o de Natureza,
ijuca, Rio de Janeiro

Ph = Philippine Bureau of Science, Manila, Philippine Islands

Pi = Polytechnic Institute of Puerto Ric co, San Germ4n, Puerto Rico

Pm = Pharma Herbarium, CIBA, Basel, Switzerland

Pn = Parque Nacional de Serra dos orgios, Terezopolis, Rio de Jan-
eiro, Brazil

Po = Ponond College, Claremont, Califo

Pr = Princeton University Herbarium, New ew York Botanical Garden,
New York City

Pu = Purdue Unive sete Lafayette, Indiana

Q = Jardin Botanico, M

C en Queensland, stews
R = Trinidad & Tobago Botanical Garden, ical beans Trinidad
Ra = Museo Nacional de Historia Natural, Buenos Aires, Argentina

Rd = Herbario Barbosa Rodrigues, Itajai, Santa Catarina, Brazil

Re = R. E. Schultes Herbarium, Cambridge, Mas Massachusetts

Rf = Texas Research Foundation, Renner,

Rg = J. T. Roig Herbarium, Estacion cieitineetved Agronomica, San
de las Vegas, Havana

Rh = a Herbarium, Causomy, Salisbury, Rhodesia

Rl = Herbario Ruiz » Godoy C Mendoza, Argentina

Ro = = Sexts Rosa pay College, santa Rosa, California

Rr = Robert Runyon Herbarium, Brownsville

S = Naturhistoriska Riksmuseum, Stockholm, en
Sa = Henry R. Carter Memorial Laboratory, * savannah, Georgia
Se = Colegio Notra Sefiora de la Caridad, Santiago, "cuba
Sd = San Diego Society of Natural Histo ry, San Diego, California
Se = University of Washington, Seattle, Washington

do E S

he Sherwood
Si = Instituto Darwinion » San Isidro, ent:
Sj = University of Puerto Rico, San Suan, Puerto Rico

1966 Moldenke, Monograph of Callicarpa 407

Sk = University of Saskatchewan, Saskatoon, Saskatchewan, Canada
Sm = Southern Methodist University, Dallas, Texas
Sp = Jardim Botanico, So Paulo, Brazil
ed = E, R. Squibb and Sons, New York 5
= Sul Ross Teachers College, gare
3s = Science Service, Department of Tories: Ottawa, Ontario,

Sz = A, G, Schulz Herbarium, Colonia Siete, Chaco, Argentina

T = Torrey He erbarium, New York Botanical Garden, New York City

Te = Torrey Botanical Club Herbarium, New York Botanical Garden,
New York Cit

Th = Thunberg H erbarium, Botaniska Institutionen, Uppsala,

Tj = mt $y of Santo Domingo, Santo Domingo, Dominican a

Tl = Tulane University, New Orleans, Louisi

T™m = Transvaal Museum, Pretoria, Trans vaal, ott err

= = United States Field Station, Sacaton, "arizona

= S. M. Tracy Herbarium, Texas exas Agricultur. al Experiment Station,
College Station, Texas

Tu = University of Arizona, Tucson, Arizona

U = Jenman Herbarium , Botanic Garden, “pee, British Guiana
Utah

on, Portu

Um = University of Montreal, Montreal, Quebec, Canada

Up = University of Pennsylvania, Philadelphia, Pennsylvania
ni is

N
: = Museo Comercial de Venezuela, Caracas, Venezuela
= Marie-Victorin Herbarium , University of Sareal. Montreal,

ec, Canada
Vl = Facultad de Agronomia del Valle, Valle del Cauca, Colombia
Vt = University of Vermont, Burlington, Vermont
Yu = Botanisches Institut der Universitut, Vienna, Aus
Vx = Teettalogi sis Institutionen, Uppsala bivaritiess, Sebuie:
W = United States National Museum, Smithsonian Institution, Wash-
on * *
Wo = Wilson Brown Herbarium, Jesuit Tertianship, Auriesville, New
¥

~s = West musisie University, Morgantown, West Vir,

Faculdade de Farmacia e Odontologia, Universidad de Sfo Paulo,
SHo P. fens, Brazil
Wi = Witte Memorial Museum, San Antonio, Texas

Wl = William Lucian Herbarium, Waterbury, Connecticut
Wp = University of Manitoba, Winnipeg, Manitoba,

4,08 PHYTOLOGIA Vol. 13, no. 6

Ws = University of Wisconsin, Madison, Wisconsin
Ww = Rob and Bessie Welder wildlife Foundation, Corpus Christi,

T
Wx = Max Gordon Herbarium, New York City
X= coe Boissier, Conservatoire et Jardin Botaniques, Geneva,

Xa = Blatter Herbarium, Saint gt College, Bombay, India
Y = Yale School of Forestry, New Hav Connecticut

Yu = T. G. Yuncker Herbarium, Erivelicastie » indiana

Z = H. N. Moldenke Herbarium, Plainfield, New Jersey

— A L., Act. Soc. Reg. Sci. Ups. (1741); Sp. Pl., ed. 1,
— css a Gen. Pl., ed. 5, 50. 175h.

adits nded synonymy: Tomex L., Nov. Pl. Gen. Diss.
Dassow 5, erie “Tila Herm. ex L., ae Pl. Gen. Diss. Dassow 5.
1747. Spondylococcos Mitch., Act. Phys.-med. Acad. Caes. Leopold.-
Carol. Nat. Cur. 8 (app.): 218. 1748. Johnsonia T. Dale ex P.
Mill., Gard. Dict., ed. fs — 75 (1752), ed. 7. 1759. soe
iococcos Mitch. apud L., Sp. Pl., ed. 1, 1: ~ in syn. 1753.

L., Sp. ct ed, =o 1s hs in syn. 1762. Tomox L. apud Adans.,

Gen. Pl. 637, in syn. 1838. Callic . apud Hassk., Cat. Pl.

Ind. Bat. 2: 884, in syn. 1857. Spondylococca Mitch. apud Benth.
in Benth. & Hook. f., Gen. Pl. 2 (2): 1150, in syn. 1876. Caly-
carpa L. apud Featherman, Rep. Bot. Surv. South Cent. La. 99+ 1891.
Burchardia (Heist.) Duham. ex Jacks. in Hook. f. & Jacks., Ind.
rit) 1: 386, in syn. 1893. Tomea L. apud Jacks. in Hook. f. &
acks., Ind, Kew. 1: 386, in syn. 1893. Callicapra Rama Rao,
Flower. Pl. Travancore 31h, sphalm, 1914. Burcardia (Heist.) Du-
ham. ex Bakh., Bull. Jard. Bot. Buitenz., ser. 3, er. 3, 32 9, in syne
1921. Burcardia Mill. ex Moldenke in Fedde, Repert. Spec. Nov.

1966 Moldenke, Monograph of Callicarpa 409

39: 291, in syn. 19%, Spondylococcum Mitch. ex Moldenke in Fed-
de, Repert. Spec. Nov. 39: 291, in syn. 1936. Cali L. ex
Moldenke in Fedde, Repert. Spec. Nov. 39: 291, in syn. 1936.
Calicoc L. ex Moldenke in Fedde » Repert. Spec. Nov. 39: evi,
in syn. 1936. Callicarpha L. ex Moldenke in Fedde, Repert. Spec.
Nov. 39: 291, in syn, 1936. Callocarpa L. ex Moldenke in Fedde,
Repert. Spec. Nov. 39: 291, in syn. 1936. Callycarpa L. ex Mol~
denke in Fedde, Repert. Spec. Nov. 39: 291, in syn. 1936. Catli-

Schau. apud Moldenke in Fedde, Repert. Spec. Nov. 2: 2h2,
sphalm. 1937. Caliocarpa L. ex Moldenke, Prelim, Alph. List In-
valid Names 9, in syn. 1940. Callicarpia L. ex Moldenke, Pre-
lim. Alph. List Invalid Names 13, in syn. 19)0. Porphyria Lour.
ex Moldenke, Prelim. Alph. List Invalid Names 37, in syn. 19)0.
Spondylococca L. ex Moldenke, Prelim. Alph. List Invalid Names 0,
in syn. 1940. Calicarpus L. ex Moldenke, Suppl. List Invalid
Nanes 1, in syn. 191. Calleocarpa L. ex Moldenke, Suppl. List
Invalid Names 1, in syn. 19).

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410 PHYTOLOGIA Vol. 13, no. 6

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ed. 2 [Willd.], 1: 87—-89. 1793; Vent., Tabl. Rég. ne 318—~
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o
b
HA
a
5
aa
x
as

1966 Moldenke, Monograph of Callicarpa 411

—— Gewtichsk. —-199, & 77h. a i
in L., Syst. Were, o7 ‘ks, aren (2): ar : 388 (1827) and 5:
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12 PHYTOLOGIA Vol. 13, no. 6

299. 1858; Van Houtte, Fl. Serres, ser. 2, 13: 127-~128, Bis ah
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99 ’
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1966 Moldenke, Monograph of Callicarpa 413
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iy PHYTOLOGIA Vol. 13, no. 6

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R

456, & 539. 1901; Garcia Caflizares, Fl. Cub. 69. pe Barn Durand &
acks., Ind. Kew. Suppl. 1: 3. 1901; Gattinger, Fl. Tenn. 143.
1901; N. L. Britton, Man. Fl. North. U. S., ed. 1, 779+ ~ 90h v9

Britten in Banks & Solander, m1. Austr. Pl. Endeavour 1: 7

1966 Moldenke, Monograph of Callicarpa 41s

(1901) and 2: a. 8 3% 1901; Diels in Engl., Bot. Jahrb. 29: 5h:7.
1901; Banks & So. Bot. Cook's Voy. 2: pl. 237. 1901; F. Ms
Bailey, Queensl, 48 sites 7h. 1901; J. J. Wood, Rec. Bot.
aes" a eee Pe & 129, 19025 By P, Wright in Cassell, Dict. sant
» 1: 156. 1902; K. Schum. Just, Bot. Jahresber.
28 B (a5: 55 1902; Verrill, Trans. Conn. Acad. Arts & Sci. 1 (2):
a6 & ree: 1902; F. S. Earle, Bull. Ala. Agr. Exp. Sta. 119: 101.
bactreen Shrubs Vines NE. U. S. 273. 1903; J. fo
Fl. east. U. S., et adi 1015. 19035 Peet, Trees
Shrubs Sete Park }. 1903; » 2h: 633. 1 9035 Ce K.
Schneid., Dendrol. Winterstud. "188, 198, & "367. 1903; Prain, Beng.
Pl. 827. 1903; E. D. Merr., Philip. Bur. Forest. Bull, 1
1903; C. B. Clarke in Schmidt, Bot. Tidsskr. 26: 1m—172. 1905
Mak., Bot. Mag. Tokyo 18: be. Fuse Thiselt -~Dyer, Ind. Kew.
Suppl. i 32. 190k; eis & Loes, in Engl., Bot. Jahrb, 3h, Beibl.
75: 61. 190h; F. M. Bailey in ag Exped. Bell.-Ker. Parliam.

ws
H

27. 1 Robinson & Fern. in A. Gray, New Man
9 *

oly. 1908; C. B. Robinson, Philip. Journ. Sci. Bot. 3: 215. on
E. D. Merr., Philip. Journ. Sci. Bot. 3: 262-263. 1908; King

Gamble, Journ. Asiat. Soc. Bengal 7h (2), page ams 794 & 308
808. 1908; Milisp., Field Columb. Mus, Publ. Bot. 2: 312—313.
1909; Urb. in Fedde, Repert. Spec. Nov. 7: ig pe E. D. Merr.,
Philip. Journ. Sci. Bot. 3: 430-31. 1909; King & Gamble, Mat.
Fl. Malay, Penins. 21: emia 19095 H. Ne Ridl., Journ. Fed.
Malay St. Mus. hh: 56. 135. 1909; Apgar,
Ornament. Shrubs U. S. tToeteoo, Pa “05.2505. 1910;

List Pl. Formosa 83-3. 1910; P. Henderson, Handb. Pl., new ed.,

416 PHYTOLOGIA Vol. 13, no. 6

65. 1910; Elm., Leafl. Philip. Bot. 3: 861--866. 1910; Mak., Bot.
Mag, Tokyo 2h: 28--29, 1910; Merr. & Merritt, Philip. rhe ee Sei.
Bot. 5: 380--381. 1910; Burkill, Rec. Bot. Surv. Ind. }:

1910; C. K. Schneid., Tlust. Handb. Laubholzk. 2: sir—s9h. "19;

° Nov. 10:
Hayata, Veget. Mt. Fuji 67. 1911; Willis, Rev. Cat. Indig.
Pl. Ceylon 69, 1911; Urb., Symb, Antil. 7: 355-~357. 1911; Hayata,
Journ. Coll. Sci. Imp. Univ. Tokyo 30 (1): [Mater. Fl. Formos.]
218--223. 1911; Craib, Kew Bull. Misc. Inf. 9: bie. 1911; Hoaasus,
Beih. Bot. Centralbl. 28 (2): 429. 1911; Gerth van Wijk,
Plantnames 217. 1911; Talbot, Forest Fl. Bombay 2: 3h5. 19113 ‘Co-
ker, Pl. Life Hartsville 31, "3h, & 100. ut Hesse, Mitteil.

b. Fl. S
Leid. 12: 15, 1912; Koord., Exkursionsfl. Java 3: 13h. 1
Loes., Verh. Bot. Ver. Brand. 53 82 fabhendt puis). isis pum &
Tutcher, Kew 10: —-203 &

mr rs sett Be at ed. 2, 3: 99, mes pee 1913; Lévl. in
pec. Nov. 2 182, 1913; , Bot a
2s 3 -eTh. “3913; F. M. Bailey, Pr ten i tak. Queensl. Pl. 362—
386. 386. 19135 Rehd. in C. S. Sarg., Pl. Wils. 1: 310. 1913; J. K.
: eat 159 &

9 Small, Fl. Southeas
pod 1913; ae Oesterr, Gart.-Zeit. 8: onacart| She, "15.
1913; McAtee, Torreya a 13: 234. 1913; Ramo Rao Flower. Pl. Tra-
. 19 rd

e e oy, Stand. Ld 2 29+
191); Fedde, Repert. Spec. Nov. Gesamtverz. 58. 191); ats Pittier,
Contrib. U. S. Nat. Herb. 18: 77. 19k; R. Ms Harper,
ria. State Geol. Surv. 6: [Geogr. & Veget. North. Fla.] 1 138, 213,

20, 2h5, 261, “oth, B93, & + 310. 191k; Koidz., Bot. Mag. Tokyo 28:
1st 152. 191h; Mak lag. Tokyo 28: 181—182. 191k; E. De

1966 Moldenke, Monograph of Callicarpa 417

Trees 383 & 385. 1915; N « Quelp. Isls. 76. 1915;
Fl. Mt. Chirisan 3. bones. tiv. Fl. Kouy-Tchéou })),0. none, 1 ; Lévl é,
- Yunnan. 2. 1916; E. D. Merr a Philip. Journ. Sci. Bot. 11:
309. 1916; ae in Curtis Bot. - 1y2: pl. 8682. 1916; Hayata,
Icon. Pl. Formos. 6: er 38. 19165 Kade. Bot. Mag. Tokyo 30:
326. 1916; Rehd. ta em Sarg., Pl, Wils. 3 366—~371. 1916; Je
Ke Small, Journ. N. Y. Bot. Gard. 18: 99-—100. od E. D. Merr.,
Philip. Journ, Sci. Bot. 12: 108 & 298-301. 1917; E. D. Merr.,
aad beng Rumph. Herb. Amboin. 4819, 526, & be 1917; Ewart
& Davies, Fl. North. Terr. 237. 1917; E. D. Merr., Journ. Straits
Ne ter,

New Internat. Dict. Eng. Lang. 312 & 2278. ri ae, Ct. Pl.
Spgs 277. 1917; Kanehira, Formos. Trees, ed. 1, 39h. 1917;

» Nutt. Pl. Nederl. Ind. oo fe 1917; Wc Rep. Veg.
ad Mts. 183. oe N. L. Bri Fl. Be sai 318--319. 1918;
Basu, Ind. Medic. Pl od at a pl. neveitetey 1918; Ww. Treleas
Wint. Bot., ed. 1, 331. 1918; Fuller, Propag. Pl. 258 & 33. 1918;
Messel, Gard. Fl. 26. 1918; Koidz., Bot. Mag. Tokyo 321 56 & 13%6—

Journ

.
o

oni 5 J
seabed Malay. Arch. 45—-92, [345]--347, 349, & (361)—362.
Bot. 1y: 249 & kS1-—

1920; Bakh., Bull. Jard.
Buitenz., ser. 3, 3: 1 3, 81, & [106 ]—108 . 1921; R. M. Harper,
Ann. Rep. Fla, State Geol. Surv. 13: [Geogr. Cent. Fla.] 100, 106,

1923; H. N. Ridl. . Br. Roy. Asiat. Soc. 1: (Mal

For. Trees] 83--8); - “LI, med “Vottet, Rev. Hort. 94: 390—391.
1923; H. J. Lam in Lauterb., Engl. Bot. Jahrb. 59: 2h—25 & 88--
90. aot. pao in belts & Lam, Nov. G Rloed Uj, Bot. 1: 167~168.
+ ontrib. U. S. Nat. Herb. 23: [Trees & Shrubs
Mex<4 3 1332 e "1253. aly H. ie Chung, Mem om. Sci. Soc OCs China i (1):
225226. 192k; Urb. in Fedde, Repert. Spec. Nov. 20: 345-36.
192h; Olmstead, aoonae, & Kelsey, Stand. Pl. Names, ed. 1, 61.

418 PHYTOLOGIA Vol. 13, no. 6

192h; W. J. s Amateurs 26 & 2. 192); L. H. Bailey,
Man. ak Pl. 1, 653. aval Mak., Ill, Fl. Jap. [89h]. 1923 Gamble,
Madras set hoo i BENS Nakai, Bot. Mag . Tokyo 383

Clute, Amer, Bot. 31: 29. 1925; H. J oJ, .

592 25. 1925; Stapp, Gartenwelt 29: ions 1928; pee Rev. Hort.

97: 416. 1925; W. J. Bean, Trees & Shrubs Hardy Brit. Isls., ed.
Bot. 30: 86~--87.

euwen, ds Plant. Bot. Gard. Buitenz. Zooced, Netherl. East
Ind. LB3--L86. 1926; M. F. Baker, Fla. Wild Fls. 191—-192. 1926;
arayanaswand, & Ramaswami, Rec. Bot. Surv. Ind. 11:
1926; -. D. Merr., Disc. & Bibl. Philip. Flow. Pl. 62, 88, i 100.
1926; J. Hutch., Fam, Flow. Pl. 1: 309. ee 4 ba Britton, Bull.
Torrey Bot, Clud 53: 63. i526; A. A. W. Hill ew. Sopris 6:
34. 1926; Rehd., Journ, Arnold Arb. 8: 16-217, “oemish, & _
1927; R. M. Harper, Ann. Rep, Fla. State Geol. Surv. 18: [Nat

akai,
19275, Yanagita, Shinrin paimacions no hibyé Dsusetsu [Illustr.
Seedlings

Re onom. Pl. Ald. 2: 311. 1928; Domin, Bibl. fe
22 (i) 7 fig. ayy 1928; Hottes, Book Shrubs, ed. l,
1471) wet oh & King, Proc. Iowa Acad. Sci. ze
185 & eae pe oe hve Fe » Bot. Mag. a 36: 22.

r. W. i Encyel, es ed. 19, 70. 1930; P. "co. standl.,
,

Field Mus. 1 t.9 1930; Uphof in Karst. & Sc
Vegetationsb. 21 (1-2): n.p. "1930; Moldenke, List Spec. Mold.
ou - U.S.A, Set 10, 1930; P. Cc. +, Field Mus. Publ

ander: cl
TL. 1931; P'ei, Sinensia 2 Ay 65—66. 1931;

tersuch. Verbenac. 17--31 & 3, fig. ee 1931 Hottes Book
Shrubs, ed. 2, 167--168 & 430. a9 5 2 Be shea Ga. State Coll.
Agr. Ext. Div. Bull. 402, , he (38). 1931; C. wa “Caras

Enum. Pl. Austr. Occid. 3: eee "1931; Kanehira, Bot. Tokyo

1966 Moldenke, Monograph of Callicarpa hig

95: [346]. 1931; Metcalf, Lignan Sci. Journ. 11: 06. 19323; C. T.
Simpson, Fla. Wild Life i82. wend Pei, age Sci. Soc. China 1
(3): [Verbenac. China] 13--56, pl. 7. 1932 3; P. Dop, - Soc.

« Nat. Toulouse 64: 97—513. 1932; inlets Brittonia 1:
185. . 1932; cet Aristocrats for the ge ot. 1932; K. R.
oynton, N. Y. Herald Trib., editor, sec , November 13. 19323
Woldenke, Bull. Torrey Bot. Club 60: rie 733; Svenson, Brook-
lyn Bot. "Gard, oon — * 1933; Kanehira, Fl. Micrones. 6——
oh 1933; H. Lund, - Y. Bot, Gard. 3h: 37. mf a. We

Spec. Nov, 33: 113 & ae: 143. 1933; Vilm., Bull. Soc. °
France 87: 82. 19333 J. K. Small, Man. Southeast. Fl. 1143 & 1518.
1933; ties aki, Nippon Shokubutsu Zufu [Jap. Bot. Illustr. Album]
1593 & 159}. 1933; Grey & Hubbard, List Pl. Grow. Bot. Gard. At-

kins Inst. 38 & - 19333 Mak., Gensyoku Yagai-Shokubutu [Nature-
Col. Wild Pl.] h: 281, 296, & 309. 1933; Bot. Sci. Select. es

th ed, Encycl. Brit. 175. —— Crevost & Pételot, Bull.
Indochine 37: 1289—1291. 193; Elm., Leafl. Philip. Bot. 6 °3222.
193k; E. J. Ale exander, Journ. N. Ye Bot. Gard. 35: 133. 193h3

Moldenke, Torreya 3h: 8. 193k; Moldenke, Brittonia 1: 253, bh,
260-261, 337, 396, ls, & lig Soar last cover page os Teel.
a ande

C Sy: Journ,
320—32h. 193); Paterm., Beitr. Zytol. ote ae gt 8, 27——
[55], & [56], pl. 3, fig. 2227, & pl. 5, fig. Poll 19.354
& E. Z, Bailey, Hortus ipatioe 670. 1935; L. H. Bailey
Florists Handl. Verbenac. mss. 1935; E. D. Merr ge hag
Philos, Soc., a ser., “3h (2): [Comm. Lour.] 3S & 332. 19355 C.
M

Coa.

Hist ae 7
(2): 95, 19355; P. Dep in Leconte, Fl. Gén. Indo-Chine : 787—799.
1935; Suzuki, Trans, Nat. Hist. Soc. Formos. 25: 130. 19353 Mol-
denke, gn ly ae Bot. “club 62: [368]. 1935; Arthur & Cummins,
Philip, - Sci. Bot. 61: 80. 1936; Beer & Lam, Blumea 2:
221-232, 19365 Sinisa Formos. Trees, ed. Pa 642—647, fig.
599-60. 19 936; Macself, Sanders Encycl. Gard., ed. 22, 71. 1936;
Moldenke in Fedde, Repert. Spec. Nov. 39: 288—317 (1936) ane h0:
38—1 21 163, 166, & 196. 1936; Masamune, Short Fl. Formos. 179-—-
180. 19365 loldenke, Decorat. Pl. Wint. Months [1]. 1936; Thelin,
Gard. Chron. Am. lil: 70. 1937; C. L. Lundell, Carnegie Inst. Wash.

Publ. 178s [Veg. Peten] 7, 75, 109, 113 138, & ney 1937;
Moldenke, Chron. Bot. 3: 311. 1937; say Journ. - 18:
Bet Cory, Tex. Agr. Exp. Sta. Bull. 550: 88. 1937} Kolden-

Phytologia 1: 191, 192, 197, 223, 227, 232, & 23h. 19375
te, Book Shrubs, ed 7 & {30. 1937; Moldenke,

Sudan, Bot. h: 17. 19373; Moldenke, “Seat Course sys st. Pag Lect.
1: 5 (1937) and 9: 2. 1337; Moldenke in Fedde, Repert. Spec. Nov.

420 PHYTOLOGIA Vol. 13, no. 6

42: 235, 238, 2h2, & 23. 1937; Moldenke, Brief Course Syst. Bot.,
rev. ed., Lect. 1: 7. 1938; Woldenke, Syst. Bot. Lect. 22: (1).
1938; H . State Coll. Ag Bull. 402, ed.
= 9385 Moldenke, Cult. Pl. 35. 19385 K. V. ;
Svensk Bot. Tidsk. 231. 1938; Macself, Sanders Encycl.

ed. 21, pr. 3, 71. 1938; A.W. Ind. Kew. Suppl. 9: 5. 1938;
PF, dl 1. Bot. 18: 999. 1938; Bullo

Icon. Essenc. Forest, Jap. 2: fig.

Icon. Eesenc. Forest. Jap. 2: fig. 28. 1938; Nakai,

Bot. 14: 61. 1938; Anon., N. Y. World-Teleg. 17, Seas ae 15395

Fedde & Schuster in see Bot. Jahresber. 59 (2): 416. 19395 H.

Pittier, Gen. Sgt Venez. Clav. Anal. 2 e

4: 317, 332, & 333. 19393 Moldenke, Annot. & Classif.

118. 1939; Moldenke, Alph. List Common Vern. Names eee 10-31,

& 33. 19395 Moldenks, Geogr. Distrib. Avicenn. 2—8, 13, 15—17,

one ae 35, & 36. 1939; Moldenke, Brief Course ee Bote,
ta & 97. 19393 Moldenke in Fedde, Repert. Spec. Nov.

A. Taylor, Pl. Used as Curative:
List Common Vern, Hanes 1j—12,
a Carnegie Inst. Wash. Publ. bay 9 & es 7-200. 190; Rehd.,

- Cult. Trees, ed. 2, pr. 1, 803-80 & 932. 19h0; penne
ormo .

Trans.
rubs, ed, se he Pa & 30. 19h2; aon N. ve Her’
9 3. 43: 36 & 2. 19h25

pr. 71 : ,
22, 2h, 25, 21, 23, 96, St, ia, bh & Sr one “"Molderke "in Lun-
dell, Fl. Tex. 3 (1): ih &'77--80.'19)2; Kanehira & Hatusima, Bot.

1966 Moldenke, Monograph of Callicarpa 421

t 8h & 229. 1943; H. F. MacMillan, Trop. Plant. & Gard., ed.
5, 104. Bo ra Moldenke, agicee Geogr. Distrib. Verbenac. Suppl. 1:
Veh. 19433 F. C. Hoelne, & t. Anual Inst, Bot. 19h: 0. 19hh;
» Journ, Avtold « 253 roped ae E. L. D. Seymour, N
Gard. Encycl. 117. macht S. A. Cain, Found. Geogr. 25h. 19hh;
Betts, Jefferson's Gard. Book 30, 139, 140, & 672. 1944; Murrill,
Guide Fla. Pl. 19 & 21. anak Roig y Mesa, Seiatc Medic! 7
ton & J

7 8

112—1ih, 116-"120) 123-135, 128--130, 135, 135—141, ih3, 1b,
147, 148, 63, 1572-162, 166, 169, 170, 17)--179, 181, 183-188,
190, 192-195; 197—200, 202—-205, 207--211, 216, 217, 220, 221,
22l,--229, 231, 232, 23h, 235, 239, 2h0, 2h2, 243, 2h5—2h9, 253—
262, 265-278, 280-286, 288, 291-299, 301—316, & 318—320.
196; Falc&o, "Guia Visitant. Jard, Bot. Rio de Jan. 42. 1947; K.
W. Hunt, Am. Midl. Nat. 37: 689 & 77. Be gal ee Sm., N.Y

5 by 35-39, id, 13, LSU, 50-52, Sin256,
3861, 63-66 TL, 5-77, 79, 81, 69— , oh, rT—idk, iot—toy,

11: 37, 39,
Woldenke, Phytologia 2: 255, 310, 326, 327, 33h, 342—3U6, &
387 (1947) and 2: 479, 483, & 506. 19h8; Serger Notice Phyt.

es [2]. 1948; Moldenke, ye al: aie 19 omnia
astanea e 120, & 121. 1948; H. Ne. & A. Le 7 Pi.
Life 2: 31, 23, 2h 28, 32, Be esa at a, oh

18,
72, 7h, ~ {ant 31. 83, & - Sperm.
1: 182-185. 19485” Erickson, ik 6s: str. "1916; 8 iH. ‘st. Souls: Pat
ific Sci. 2: 273. ‘ola; WN , 640. 19h85
Rieti BotsGinde beed took. List 1948: 2. 1918; maaeeke, Aiph.

27, 329-331, 333-33 oo
349-351, 354—356, 358, 359, 389—395, 397——L1l, LIb— Lis, Lis,
429, 432—h37, bh3, bh7——b52, L5u-—-L59, 462, 463, » 469,
470, 472--h7h, 477; 478, 480-182, 4B, 486—191, Weae-hoe, 199,
503, 50h, S062- ? 512-51, 518--522, 52h—531, 53h, 536,
5uS; 548-550, ooh, 556--570, or eh 576, S77, 579—~—581, eae.

422 Pal J T'07G O:Grick Vol. 13, no. 6
585, 588, 592, 59, 598, 601—609, 61-619, 621, 625, 626, 631,

3h, t oli —652 (29l8),” 3: 695-660, 663-666, 673, 67h,
Gea 82’ 2° 683, 695--700, 702, 706, 708, 709, 712——71k, 717—
732, 736-738, 7 Tie O-—7hh, 77, nes 767-171, 773-776, 778—
80, 782--795, 197--802, 805—~807, 809-811, 820, 522,

8, 1191, 1192, 1202,
21h, 1216—1218, 1220, 122, 1226—1228, 1230—~1232, 123h——12h3,
12h5, 12h6, 12h8, 1251-1255, 1259, 1260, 1283, 1288, saa 129i
1293, 1296-1299, & 1303. 1949; J. L. Hoffm., The S 25, 9

3 E. D. Merr., Ind.
Berg. 15 (2): 2, 199; R. F, Thorne, Castanea 1): 91. 19495 Ber-
ger, Determinat. Houts. Mal. Fam. 39° & 72. 199; Mes rt
2 ’ £

- & A. L. Moldenke, Anal. Inst. Biol. Mex. 20: 3h. 19
ssadenke, Bol. Soc. Venez. Cienc. Nat. 11: 291. 1950; 2
77

pa 31: 2l,. 19805 i Moldenke, Bevist. = compa - 8s Jeg, 1?
172. 1950; eee, Lloydia 13: 205, 1950; T. a Everett, Ne ce
Bot. Gard. cet ng list ae a & 5. 1950; MoBacharns List

e Nov. _ :
Mold. Set 6 Spec. 2. 1951; woldenke, Biol. Abstr. 25: 3050. 19515
N. Y. Bot. Gard. Seed Exch. List 1952: 2 & 5. 1951; Keevar, 008
ting, & Anderson, Bull. rabbi tite Club 78: 09 & 420. 19513 ad
put, N. Y. Herald Trib., sect. h, 17, September 6, 19915 Sage
Ill. Fl. Jap., rev. ed. 1951; Chang, Acta Phytotar. 1: 27

22 ie) Wea 195 = si hg——hot
1; Moldenke 33 ge ti 07,
Né1—l6h, él, 187188, & Sol einer eta Ca aa

1966 Moldenke, Monograph of Callicarpa 423

& Shrubs East. N. as 16, 103, & 329. 1982; Anon., Biol. Abstr.
26: 3446. 1952; E. He Walker & al » Fl. Okinawa mong aad Razi,
Poona Univ. Journ. 1 (2): \7. 1952} 3 Santapau Cavanil-~
les 11: 310. 1952; Moldenke, Biol. Abstr. 26: 63 & iT (1952)
and 27: 98 & 3121. 1953; Roig y Mesa, Dicc. Bot. 2: 27h, 389, 390,
& 996. di sere de Leon Fao Soc. Cub. Bot. 10: 23. 1953; Je
A. So he oe i 238 19533 yer E —— Botan, Staats-
. Mtinchen ey 225. 19533 ogia 4: 191, 2, 393, | oe
205-287, 3925 393, U50, & ‘yer (19533, liz 506 (19 95h), and
28—— , 100, 1954; Moldenke, Inform. Mold. Set 5 oe
i. phi Molden, mrt Calif, Hort. Soc. 15: 80, 81, & 85—
1954; W. ©. Klein View Guinea 2: 241. 1954; Laszlo & Hen-
ee Science 119: 628. 195k; Re - Fe Thome, An. Midl. Hat. 52:
ty "195h; Teks . rooks, Proc. Fla. State Hort. Soc. 67: 81-82.
Menninger, 195), P Price List t ta) 1954; Anon., Biol. Abstr.
28, hose (196h)” ps 27: 3764. 1955; Santapau, Joum. Bombay Nat.
Hist. Soc. 53: 27. 1955; Mizushima, Misc. Rep. Research Inst. Nat.
Res. 38: 123. 223-1858 Benning Dict. Terms enemies - 29, 88, & 89.
55; Masamune, Enum, Trache Isls. 7 6. * 9555 Mol-
denke, ar Abstr. sii 2935. 1955; er apy Fam
1956; Moldenke in ¢ esman, Fl. Trin, & Tobago 2 (6): ies. 19 19565
Moldenke, Inform. weld. Seb 51 Spec. a 1956; Moldenke
Cat. Estat, Gen. Bot. Fanerog. 17: 3. 1956; Moldenke in Peers
Fl. Madag. 174: hh—8 & 265, fig. vil & 2. 1956; Alain, Rev.
Soc. Cub. Bot. 13: 37—38. 1956; Kasapligil, Pl. Jordan 102. 1956;
Putz, N. Y. Herald Trib. October 7, 1956; Spector, age Biolog.
Data’ pe 1956; Alain in Leén & Alain, Fl. Cub. h: & 304--309,
fig. 131. 1957; Anon., Biol. Abstr. 29: 3165. 1957; eta.
1957 Price List [2]. 1957; Vah Steenis, Fl. Males. Bull. 380, 453,
xxix. 1957; Monachino, Rhodora 59: 19. 19873 Le Benson, Pl.

Pd
Soe
8
Ses
RES
ge
3
a
ru
ee

Biol. Abstr. 30: sites 19985 “Shanners Spring Fl. Dallas 328.
1958; Conde, Hist. Bot. Cuba . 1958; deRoon, Internat. Direct.

2

& Mori, Misc. Rep. Res. er akg Resource es 48:

1958; J, & 1; _Bush-Brom, Am, Gard. em rev. ed., 269. 19583
Nut tut, tree Bore , Spring 'S9 Price gr "19595 Krtissm., Handb.
1: 253-285. 1959; Hara, Outline Phytogeogr. Japan 7, 3h,

55 5, & "8. 1959; Kitamra, Fauna & Fl. Nepal 208-209. 1959; Hatu-
ints & Sci. Div. Ryukyu Univ. Math. & Sci. 3: 107.

1594 #50; ot. 2 1h. 1959; Rai, Rec.
+ Surv. Ind. 18: 9. 1959; Moldenke 8 12 -
23, 25, 27, 3h, h0, bi, 43, bh, 6, b7, 49, 50, 53, 5k, 56, 58,
59, , 82, 113, 155, 157 160, 16 165--168, 171--175

,

» 70,
202-— oe.
211 2137 2il,, 229° 230, 23K, 2h0--2k8, 260° 29k, 295, 2987 30L,
319, 336, 337, 3h5, 6° 35h, 355, 379, 396, 399, 408, hog, its,

42h PHYTOLOGIA Vol. 13, no. 6

ara; 423, 2h, h27, & Lb3—bb5. 1959; Subramanyan,

urve India 1:1 133. 1959; Moldenke, Résumé Suppl. 1
16, & 2h. 19593 Moldenke, Phytologia 6: 509 (1959) De t "21, "0, fee
& 91 (1959), and 7: 123, 2hh, 293, 300, & 305. 1960
ésumé suppl. 2s a5 - 1960; Menninger , 1960 Pri ce List ee rei
[2]. 1960; Angely, Liv. Gen. Bot. Bras. 35 & 0. 1960; Moldenke,
Biol. Abstr. 35: 1687--1688. 1960; Rehd., Man. Cult. Trees, ed.
2, “gk ae ee . 932. 1960; T. H. Everett, New Illustr. Encycl.

- & Ferns Tex. coals Bend 142. 1961; W. H. ey Southw. Nat.
: (1): 45—h8. 196k "Werden, Biol. Abstr. 36: 036 Fo and
7: 1062. 1961; M , Phytologia 7: 321, 326, 382,
igi, & 508 (1961), "8: e7'& 9 (A9e1), and BY 176, 273, & a obs.
1962; Hatusima, Mem. South. Indust, Sci. Inst. Kagoshima U Univ. 3

5 (1 ) and A.5: & hy ° _cteeyes

861. 1962; amt “CD! "302-303.
1962; Keever, Ecol. Monog. 32: 160. 1962; Nair &
Rehm., Bull. ard. Lucknow & 23, fige 8 &
19. 1962; Woldenke, Biol. Abstr. 373 1062 1962) and. 39: 61h.
1962; Mold Suppl. 3: 2—6, 16—2h, & 30
(1968), he 1, 3, & 7-11 (1962), ay 1, 2, & i 2 ta568), 62 8
(1963), and 7: 1, 2, os helford, Ecol. N. Am. 6

,& ee 1963; Hocking, Excerpt. Bot. A.6:
las & 535. "19635 "H. L. Hoffm., Cas bis tanea 23: 170. 19633 Je As
erm., Fl. Mo. 1257, 1262—126h, & 1267, ee 1849, pl. 303,
tie, h. 1963; li, Wood, Pl. Taiwan 17, 816, 81 Beebe, * 94h, fig.
Lie 1963; Anon., Biol. Abstr. he (3): B.17- a6)
Dansk Bot. Arkiv 23: 87. 1963; Li, Morris Arb. Bull KE (aj &
3—-7, fig. 1—7. 1963; Van Ste sass Fl. Males. Bull. 18: 1069.
1963; Van Campo & Planchais, Pollen & Sp. Bibl. 5: 472. 19635
Moldenke, RE Abstr. 2: 1517. 1963; Bhatnagar, Journ. Ind.
Bot. sere (3): 374. 1963; Bush-Bromm, Shrubs & Trees Home 4
A on pee Sharma & ecient Journ. Genet. 53:

» Fl. Java 2: 595 & & 600—601. 6 jhe sae .
sere Stich. Konink, eh oat Dierg. [6]. 19655 Neal, Neal, In Gard.
Vv. ed., 721, 726, & 727. 1965; San oe

Arb, Bull. 16: S1--£3, 1965; Moldenke, Prytologia 12: a 12: 6 (1965)s

1966 Moldenke, Monograph of Callicarpa 425

$.143. A900 Be my i r Herald Trib. ey sect. 2, 20, February 6.
1966; G. Taylor, Ind. Kew. Suppl. 13: 21. 1966; Balakrishnan, Bi-
olog - Abstr. 7? 4615. 1966; J. % Clark, Card Ind. Gen. Sp. Pl.
Nee
This is a genus of about 188 species and subspecific taxa and
hybrids, widely distributed in subtropical and tropical Asia and
Oceanica, to a lesser extent in the Americas, a few species ex-
te into the temperate portions of Asia and North America
south to Australia and west to the Seychelles and Madagascar; * many
are cultivated for their handsome fruit. The type species is Cc.
americana L, The generic name is derived from the Greek words,
KZNKOS, beauty, and KEPTS fruit — the first letter of the
first Greek word is given erroneously as "X" in my earlier work
this ge h a printer's typographic error which I was
not permitted to correct.

Bentham & Mueller (1870) ahaa to "a few African and American
species", but I know of no ae een of this genus native to conti-~
nental Africa, They say that "T pial is it readily distin-
guished from Premna by the Tatiaveateies and by the flowers more
regular with isomerous stamens. The differences in the fruit may
not be constant.

It is of interest to note how authors in the past have dif-
fered in their estimates of the size of this genus. Bentham (1876)

and

Although Loureiro proposed the name Porphyra in 1790 and Agardh
did not propose his algal gems of the same name until 1822, the
latter has un fortunately been conserved and is valid for the algal
genus under the present edition of the International Rules of Bo-
tanic Nomenclature!

Referring to Hasskarl's "correction" of the spelling of the
name Ci carpa to "Callicarpus", it is pointed out by * allo
(1950) et under these same present rules the name mst remain

orig proposed by

It should be ater here that Illa aor "included * the gener-

© synonymy above, being a mononomial, actually also a part o
the synonymy of the species Callicarpa ae (L.) s mathe The

aurchardia of Robert Brom, referred to above, is a genus in the
Liliaceae.

Geunsia Blume is included in the synonymy of Callicarpa

Schauer (187), Miquel (1857), Lemaire & eh =< Ao), and

26 PHYTOLOGIA Vol. 13, no. 6

Bakhuizen van den Brink (1921), but I agree with Dr. Lam in re-
garding it as a distinct and valid genus, differing in having its
gynoecium usually 5~ (rarely -) haope its stamens ae
5 in mumber, and its leaves u ee dae sophyllous. Aganon Raf.
is included in the synonymy of Callicarpa by Jackson (1893) and
by Bakhuizen we den Brink het as well as by myself in my 1936
work on the genus, but, according to Merrill's monumental commen-

ome Gmnencemenmee” codes Guniemeniniemas | ammmimanan

allicarpa is p ,
Verbeneae, by Reichenbach (1827, 1828, 1833), with "Spondylococcus
Mitch. non W." as a synonym. This same author regarded Po
ee as a valid and distinct genus. Dahlgren (1938) also places
allicarpa in the Labiatae,

additional German common name for the genus as a whole, not
in

s members of the group are called "beautyberry",
"beautyberries", or sbasute-bereies". DeLaszlo & Henshaw cha
quote H. A. Marshall (1922 species

regarding a
called "“argerarger" in the Torres Straits, whose young asaves,
well chewed by a woman until her body its saturated with
swallowed juice, will cause permanent Roepe Jones (1798) re-
cords the vernacular name "m4s

© gynoecium morphology of the genus is discussed by Jun
(193k). Roxburgh (1820) refers to the fruit as a l-seeded sane,
with the seg erect, and ge d with a perisperm". Actually,

mental shrubs grown for the abe fls. and handsome
frs. late in Pheer rather tender ed back, the
ene shoots usu. usually flower and fruit the sie season,"

the Linnean Herbarium there are three specimens preserved
site genus 136, Callicarpa, The first of these is labeled "l.
americana" in Linnaeus’ own handwriting, plus a label of Gronovius
and the notation "vera". The second specimen is labeled "2. to-
mentosa", with the added notation "cana ?" by J. E. Smith {accor
ding to B. D. Jackson]. The specimen is neither C. tomentosa (Le)
Murr, nor C. candicans (Burm. f.) Hochr. [formerly know as C.
cana L.], but is typical C. nudiflora Hook. & Arn. The third
specimen is big pe So, “labeled merely "3" and "Ind. or.", —
is plainly C. candicans
Cain (19])) reminds us that Callicarpa is one of the consider
able mumber of genera listed by Asa owe as being represented
Japan and pillpesrasecin {eastern Asia] and in eastern North America,
but not in Europe or western North America

1966 Moldenke, Monograph of Callicarpa 427

Reatarte apes records the fungi Meliola cookeana (black mil-
dew) from s gemus in Florida and Louisiana and Cercospora cal-
licarpae ‘last rat) from South Carolina to Texas. It is most
probable that she refers here to C. americana L. as the host,
Arthur & Cummins (1936) record the fungus Uredo callic Cc
from an unidentified species of Callicarpa represented by Clemens

5029.
a dentified species of this genus from the Philippines is
illustrated by Ettingshausen (1861). The Frutex ceramicus of
rw pais Ih, plate 60, is an as yet unidentified species of
Merrill (1917) says "This was described from mater-
a. Sierebine in Ceram and Banda and is undoubtedly a species
of Callic - Hasskarl, Neue Schilttssel (1866) 84, suggested
that it might be Grewia inae s Blume, but the drawing certain-
ly represents no Grewia, while the description seems to conform to
Callic « Field work is necessary in Ceram and Banda before the
position of Frutex ceramicus can be definitely settled. Hasskarl
ee+eeAlso suggests that Feilielus alter silvestris may be a spe-
cies of Callicarpa, but the description is too incomplete to war-
rant a definite prrervere of this to any ptigets ext genus." I may
mention here that, of the present writing, I know only C.
longifolia Lam. Trial. cenmaae a
The Matheus 1210, distributed in some herbaria as a Callicarpa,
is actually a species of Siparuna in the Monimiaceae; Clenens &
Clemens 959 is Buddleia racemosa Torr. in the Loganiaceae. Wiss-
mann 3, i in the Vienna herbariw rbarium, is determined as Callic on
its | label, but consists of only two leaves which I do not think are

A sheet from Australia labeled "Burchardia ia umbellata aR. Bre" is in
the Callicarpa folder in the herbarium of McGill (Gill University, but
ds not verbenaceous — it represents a valid genus and species in
he

eae,
The Uphof (1930) reference in the bibliography: above is some~
times oe epee: dated "1935--1936" in ature; the Lam & Mee-

ct
a
2
ry
8
ct
@
E
‘
8
a
ie
i
~~
ke
Oo
~~
ise)
ww

error, r

is often misdated "189", but the section on Gamopetalae was actu-

ally issued in 1892. Siebold, Jaarboek van de Koninklijke Neder-
van

che Maatschappij tot oedig en Tuinbouw 1845: 71,
Pl. 5 & 6 is often cited in litera as "Ann. Hort. Pays—Bas pl.
5 & 6", an alt tle. The Buchoz (1783) reference is some-

labelled "xoxvi" in the New York Botanical Garden library copy
reference "Journ. Malay. Br. Roy. Asiat. Soc. 2: 615. 1923" is is.

428 PHYTOLOGIA Vol. 13, no. 6

sometimes rine but on oe is no ~~ page mumber in the stated
volume; the same is true of the "Journ. Roy. Asiat. Soc. Bengal
73 (3): 1012. & 1018. 1909",

imens from the herbaria of the University of California

3
B
g
i]
be
Eg
Q
mw
gees
FE
:
E
3
Ba ee
2
H
&

1936 work on this gemus without herbarium sheet numbers, are re~-
peated in the present work with the proper sheet numbers added.

An alphabetic list of names applying to taxa excluded from the

genus Callicarpa:

Callicarpa acuminatissima Sg ae & Binn. = Geunsia acuminatissima
(Teijsm. & Bimn) H. J. Lam

Callicarpa affinis zim, = Pee farinosa Blume

Callicarpa apotnsis Elm. = - Geunsia . apotnsis Laan Moldenke

Callicarpa cavaleriei Lévl. = Ilex sp., llicaceae

Callicarpa cinerea A. Rich. = Petitia pian s Jacq

Callicarpa cordifolia Rufz & Pav. = av. = Aegiphila mri eat (Rufz &
Pav.) Moldenke

Galle dearpe cumingiana Schau. = Geunsia cumingiana (Schau. ) Rolfe
sestys, Suatete dentata Pav. = Cormtia grandifolia (Schlecht. & Cham. )

ae
callicarpa dentate dentata Sessé & Moc. = Cornutia grandifolia (Schlecht.
& Cham.) Schau. ieee as 3 ‘e
Callicarpa discolor Willd. = egiphila integrifolia Jacq.) acks»
Le epiphytica Elm, = Geunsia fla da (Eim.) H
Callicarpa esquirolii Lévl. [Esquirol mre ert Cibettias

» Hydrangeaceae
catticarpe’ sequirolit meets [Esquirol 754] = Caryopteris panicu~
pe Ns Ce Be
Callicarpa iiadat bask. & Quisumb, = Geunsia pene
(Schau.) Rolfe
Callicarpa flavida Elm. = Geunsia flavida (Elm. ) He Je
eee globiflora Rufz & Pav. = Aegiphila antagiitalth (Jaca)

acks,
callicerpa hexandra Teijsm. & Bimn. = Geunsia hexandra (Teijam. &

Binn.) Koord.

Callicarpa hexandria Teijsm. & Binn, = Geunsia hexandra (Teijm™-
& Binn.) Koord. Koord

Callicarpa integrifolia Jacq. = Aegiphila mies (Jacqe)
Jacks,

oacacarpa janate ianata Lam, = Premna tomentosa W.

SE leveilleana Fedde = | = Dichroa Feit poe Lour., Hydranges-

caliieerpe martini martini Lévl. = Caryopteris paniculata C. Be Clarke
Callicarpa martinii Lévl. - Caryopteris paniculata C. B. Clarke
Cailicarpa mekongensis W. W. Sm. = Premna mekongensis W. W. Sm

1966 Moldenke, Monograph of Callicarpa 429

Callicarpa paloensis Elm, = Geunsia palo Satis ibe) ease
Callic ae ganieulata (Kurz) Clarke = Caryopteris paniculata C.

B.
camsearpe ps pasiciatel Lam. = Chilianthus arboreus (L. f.) Benth.,
eas

Sage pentandr Roxb, = tandra (Roxb.) Merr.

Sens pentandr. “ay farinosa aa St eey Bar Bakh. = Geunsia farinosa
Blum

Git 14oerpa pentandra f. floccosa Bakh. = Geunsia hexandra (Teijsm.
& Binn.) Koord.

Callicarpa pentandra f, glabra Bakh. = Geunsia pentandra (Roxb.)
Merr.

Callicarpa pentandra f. glabrescens Bakh, = Geunsia pentandra
(Roxb. ) i

Callicarpa pentandra f. pubescens Bakh. = Geunsia pentandra (Roxb.)

Merr,

Callicarpa pentandra var. cumingiana (Schau.) Bakh. = Geunsia
c ana (Schau.) Rolfe

Callicarpa ee var. cumingiana f. dentata Bakh. = Geunsia
cumingiana var. dentata (Bakh.) Moldenke

Callicarpa pentandra var. cumingiana f. genuina Bakh, = Geunsia
cumingiana (Schau.) ‘Rolfe
Callicarpa pentandra var. cumingiana f. pentamera (H. J. Lam)

Callicarpa pentandra umingiana f. typica Bakh. = Geunsia

cumingiana i 7a ey
Gallicarpa pentandra var . paloensis (Elm.) Bakh. = Geunsia palo~

Lam
Callicarpa pentandra var. paloensis f. apoensis Bakh. = Geunsia
apotnsis beri ) woldonke
“Scarpa B pentan . paloensis f. apoensis (Elm.) Bakh.

iets banat “(zln. ) Moldenke
isc pentandrs var. paloensis f. celebica (Koord.) Bakh. =

Geunsia palotnsis var. celebice (Koord.) Moldenke
Callicarpa pentandra var. paloens oensis f. furfuracea Bakh. = Geunsia
furfuracea (Bakh.) Mol a

Callicarpa pentandra var. paloensis f, gemmina Bakh. = Geunsia
o8nsis (Elm.) H. J.
Callicarpa pentandra var pees is f. typica Bakh. = Geunsia
palo§nsis (Elm.) Bees
Callicarpa tandra poceeocal Bakh, = Geunsia cumingiana

Callicarpa pentandra var. repleta f. celebica (Koord.) Bakh.
Geunsia paloUnsis var. celebica (Koord. Molde:

Gallicarpa p tandra = Neg replete f. furfuracea Bakh, = Geunsia
furfuracea (Bakh.) Moldenk

430 PHYTOLOGIA Vol. 13, no. 6

Callicarpa pentandra « typica (Schau.) Bakh. = Geunsia pentan-
dra (Roxb.) sia
Callicarpa pentandra Mak typica f. farinosa (Blume) Bakh. = Ge-

pentandra

unsia farinosa | Blum
Callicarpa pentandra var. pice a f. flavida (Elm.) Bakh. = Geunsia
flavida (Elm.) H. J. rage mee

Callicarpa p tandra var. "pps a f. genuina Bakh. = Geunsia pen-
tandra (Roxb.) Merr S a
Callivarpa pentandra var. typica f. hexandra Bakh. = Geunsia hex-

Callicarpa pentandra var, typica f. hexandra (Teijsm. & Binn.)
Bakh, = Geunsia hexandra (Teijem, & Binn.) Koord.
Callicarpa pentandra var, var. typica f. pubescens Bakh, = Geunsia
cumingiana (Schau.) Rolfe
Callicarpa prismatica Robledo = Bouchea prismatica sag : Kuntze
Callicarpa reticulata Poepp. = Petitia do omingensis J
Callicarpa sorsogonensis Elm. = Geunsia palotnsis enh H. J. Lam
Callicarpa spinosa Harshberger = Aralia See nosa L., Araliaceae
08

Callicarpa umbellata Lour. = ? =? Premna sp.
Callicarpa eee Diels = Viburnum r rhyrtd dophy1L.um Hemsl.,
oliac

Callicarpa Hii Presl = Siparuna velutina (Presl) Moldenke,
Monimiaceae site OE,

CALLICARPA ACULEOLATA Schau. in A. DC., Prodr. 11: 642. 16h7.
Emended synonymy: Callicarpa verticillata Hort. ex Moldenke in
Fedde, Repert. Spec. Nov. 0: 79, in syn. 1936.
Bibliography: Schau. in A. DC., Prodr, 11: 642. 1BL75 Grisebe,
Abhandl. Ktnig. Gesell. Wissen. G&8tting. 7: 256. 1857; Jacks. in

Fie!
May
g@
3
$
E
3
S
t
ba
rf
Ny
ke

535 78-82, 120, & 121. 1936; Mo

vicenn. 7. 1939; Moldenke, P - Alph. List Invalid vanes ets

19405 eed Known Geogr. Distrib. Verbenac., ed. 1, 26 & 86.

19,2; Moldenke? Alph, List Invalid Names 11. 1942; Moldenke, Alp».

List Git. 1s 33 (2946) 23 ait, 121, & 432 (1916), and 3: 699+,
own Geogr. Di . Verbenac., ed. 2, 17 & 170+

Moldenke
199} Moldenke, Rbeuné 56, 2h7, 7 t li. 1959.
In all, 10 he , including type material of all
the — geek xainy and 7 mounted photographs have heen examined

vphsttonan citations: HISPANIOLA: Dominican Republis: Bertero
9 [Macbride photos 33931] (De—type, E-—1192hh—isotype, 7 Fo
Bie ae of type, Kr—-photo of type, Mu--9l1--isotype, N--

1966 Moldenke, Monograph of Callicarpa 431

— of type, N--photo of type, Z—-photo of type); Lavastre 1718
N, e

CALLICARPA ACUMINATA H.B.K., Nov. Gen. & Sp. Pl. 2: 252. 1818
t C. acuminata Roxb., 1814, hyponym].

Additional & emended synonymy: Callicarpa subintegerrima H.B.K.,
Nov. Gen, & Sp. Pl. 2: 252-253. 1818. Galdscares acuminata Humb.
& Bonpl. apud Steud., Nom. Bot., ed. 1, 137. 1621. Callicarpa
subintegerrima Humb. & Bonpl. apud Steud., Nom, Bot., ed. 1, 137.

Kunth apud D. Dietr., Syn. Pl. 1: 428. 1839. Callicarpa mollis
Req. ex D. Dietr., Syn. Pl. 1: 28. 1839 [not C. mollis Koord.,
in herb., nor Matsumura, 1922, nor Shirasawa, 199, nor Sieb. &
Zuce., 184h) + Callicarpa subintegerrima Humb. & Kunth apud D.
Dietr., Syn. Pl. 1: 428. 1839. Callicarpa mollis Willd. ex
Steud., Nom. Bot., ed. 2, 257, in syn. 1840. Callicarpus acumin-
ata Humb. apud Hassk., Cat. Pl. Hort. Bot. Bogor. Alt. 136. 1844.
Callicarpa acuminata Kunth apud Schau. in A. DC., Prodr. li: 6L4.
1847. Calli schlimii Turcz., Bull. Soc. Imp. Nat. Mosc. 36
(2): 217--218. 1863. Callicarpa acuminatta H.B.K. apud Goyena,
Fl. Nicarag. 2: 566, sphalm. 1911. Callicarpa minutiflora Rusby,

37. 1821;
st. Veg., ed.

cer hanes 10. 1814;

H.BoK., Nov. Gen. &
ed. 1, 1

Alligem. Medi
D. Dietr., Syn. Pl. 1: 428. 1839; Steud., Nom. Bot., ed. 2, 257.
180; Hassk., Cat. Pl. Hort. Bot. Bogor. 136. 1844; Walp., Repert.
aero 3 Rosenth.,

tze ft Be Hemsl., sien ae cas are = Pl
> Vv. Gen. Pl s623 03. 913 Sess Oe, bay
18. 1893; Jacks. in Hook. f. & Jacks., Ind. Kew. 1: 386. 18935

432 PHYTOLOGIA Vol. 13, no. 6

— & Moc., Fl. Mex., ed. 2, he 189h; — Field Columb.

- Publ. Bot. 1: 42. 1895; . in Engl. & Prantl, Nat. Pflan-
ceuren: k Gas 166. 18954 He aX came, Mem. Torrey Bot. Club 6:
106. 1896; Bull. Herb, Boiss. l: 3h5. 1896; Just, Bot.
Jahresber . 23 5 (aS: 76 & Sil. 1897; Goyena, Fl. Miearag, "2: 5
1911; Loes., Verh. Bot. Ver . Brand. 53: 81 [Abhandl. ee 1912;

, Bull. Jard. Bot. Buitenz., ser. 8 3: 22, 1921; P.
Standl., Contrib, U. S. Nat. pin Ea 23: 1253. 192k; N. L. Britton,
- Torrey Bot. Club 53: 463. 1926; H. H. Rusby, Mem. N. Y.
1927; P. C. Standl., Field Mus. Publ. Bot. 10:
To

’
5, 18, & 2h. 190; Moldenke, Prelim. Alph. List Invalid Names
9, & 1113. 1940; Moldenke, mae ae Inst. Berg Publ. 522: er age
200. 19,0; Woldenke, Known Geo gr. Distrib. Verbenac., ed. l,

» 30, 31, 3h, LO, 71, & 86. 19h2; cidenke, Alph. List In-
valid Names 2 & 8—-11, 1 1942; C. L. Lundell, Contrib. Univ. Mich.

S
oy
=
Re
.
¥

2h, 35°
» 507, 526 529, 549, 557, 566, 569, 573, 603, 607,
679 me 709, ae 7 hy e "1 768, ; “teste
? ? 730, 740, 7 . (os 39 779,
es e16--820 8 Be Br, is 900, 9 » 903,
925, 918, 959, 360° 962, 96, $65 970, a3, &
: 98 1 00%, 1006, 1013

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8s
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£
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9 20 Ns te 230, 21, shh, 245, 2h7, & us.
19593 Woldenke, Résumé’ Suppl. “Ae 19623 J. A. Clark, Card Ind.
Gen. Sp. Pl. n.d.

Recent collectors describe this species as a woody shrub, 1-~
20 feet tall, wide-spreading, scrambling, or even scandent, ‘or 8

1966 Moldenke, Monograph of Callicarpa 433
small, riigerpatie or bushy tree, 4--10 m. tall, the trunk 3/l to 3
e —c

in
dense _clusters. Steyermark refers to the fruit as "berries", but
e drupes. The corolla is described as “whitish" on Mar-
paedaldettn 195, "creamy-white" on R. McVaugh 104,89 and i.
Williams 981k, "lilac and yellow" on Klug 3879, "violet and | yel-
low" on Klug ug Libs, and ia tant on P, H. Allen 2188 & 528, Be
aert 26, Breedlove 6488, Cooper 220, Cuatrecasas 962, Haught
2127 & & 1,868 » Re M. King 3 om Leavenworth 226, Lundell & Lundell
7108, 7519, 7810, & 16402, Steere 100k, Steverwaste ark 56771, and
Tuncker r 4883. B Bequaert ert 26 is very densely pubescent and may pos-
sibly represent C. pringlei Bri
Common and vernacular names recorded for C. acuminata are
, "aurora", "axin", "blackberry", "Bonpland's Wirbelbeere}
"ceniciento", "filigrana de pifla", "flor de chichalaque", "fructa
*patzahumac .

’ "sac pukim",
"uva", "uvilla", "vara de alcalde", "vara del alcalde", "x puc

carpa acuminata Roxb. It is most probable that he is referring
here to C, americana L. or to C. roigii a both of which are
found in Pinar del eee while Soe te =“ Ce. ac is unknown

from the island. The e C, acuminata Ro = Se in the syn-
onymy of C. nudiflora Hook. ~& Arn., an Old World species.

The type of C, subintegerrima was probably collected somewhere
in Colombia [roreesdt in Regno Novo-Granatensis?", according to
oo original description] and is deposited in the Bonpland

a!

The C, americana Sessé & Moc. included by me in the synonymy of
C. acuminata in my 1936 work belongs actually in the synonymy of C.
Pringlei Briq. instead and the discussion of this name on p. hl of
my work should be so emended. The name, C, lancifolia Merr., men-
tioned in the synonymy above, is a synonym of C. i merrillii Molden-
ke, but C. lancifolia Millsp. is a valid species. The C. mollis of
SPR rg is actually C. caudata Maxim., that of Matsamura is Pia Ge
okinawensis Nakai, that of Shirasawa is xC. s Mak.,
While that of Siebold & Zuccarini_is a valid species. Hasskarl re-
cords (184) a "Callicarpus acuminata Hmb. Sprg." from Japan, but
this is doubtless an error for or Callicarpa nudiflora Hook. & Arn.
(c, crenata Roxb. oe:

plant has been found growing in forests or mountain forests,

rg fevests: light or tropical forests, thickets or open thickets,

43h PHYTOLOGIA Vol. 13, no. 6

old or occupied clearings, acahual, llanos, the alluvial soil in
secondary matorral, secondary growth, lowland jungles, second

ests on — hills, alo: ong semidesert roadsides, on slopes and
ides, at the edge of brush, and along village borders,
a“ t altstndes | of 10 to 1800 mete rs, flowe owering from March to Decem-
g in February, April to September, and November.
eed Me King describes it as "locally abundant in open sun" in Alta

in San Martin, Peru; while McVaugh says "occasional in oak forest
on limestone” San Luis Pitan and Crutchfield & Johnston found

"occasional in swamps and thickets along the coast; r oer G to Pa-
and Mexico", while later (1938) he says "occas in
thickets of the tierra caliente, Mexico to Bolivia",

e of the specimens of Gouin s.n., cited below, has its fruit
modified into pubescent galls, The Tonduz s.n. (Herb. Inst.
Physico-geogr. Nat. Costaric. 1882] cited by me on p. 2 of
1936 work as from "Province undetermined", is actually from Linén,
Costa Rica. The Holton 772 from "Las Caflas", Colombia, is said by
Dugand to be from Valle del Cauca. The Seas sn. from "Coyaje-

Honduras, cited on p. 2 as deposited in the United States Nation-
al Museum at Washington, is now in the Marie-Victorin Herbari

at the University of Montreal. The Gaumer 1688, cited on p- h2
as from Yucatén, is actually from Quintana Roo, ; Mexico. E. P
Killip changed the number on ctenicans "9,02" in the United
States National Herbarium to read "962" on specific instructions
from the collector.

Material of C. acuminata has been misidentified and distribu-
ted in herbaria u under the names C. pringlei Briq., C. pringleii
Briq., C. subpubescens Hook. & Arn., Clibadium sp., Psychotria
hebeclada Dc. » and Viburnum americanum Mill.

The H.B.K. reference given in the bibliography of this eo
a — cited a a MELTA. bat apparently ag tually be "18184

by (189 6). reference is cited as "Rusby, pi ne Memoirs VI
(2896). "106" at the Instituto Miguel o in
» 52 specimens of C. rec ate including type
ee of almost all the nasies involved, and 56 mounted photo-
graphs a been or by me
& emended citations: MEXICO: Campeche . Landelt
a8 (cal86976, Du-—222626, E—1017115, F—70052h, Weigle)
2006 (ca--U86898, Du—22h,768, E--1017030, F--700607, arrepiig

Chiapas: Breedlove 6286 (Ac), 6488 (Z); Gilly & Hernandez Xolocotss

a Gus), = a ay; iat Matuda 1012 an i nae 3641 (F—1026375, Mb,

Mi, . Hidalgo: O. M. “Clark THOS (ok--18723) 5, sehnooberges ,
aoh7 oe © Jalbedss Mexia a 1108 (Ca—350067, Du—17 3

—e

1966 Moldenke, Monograph of Callicarpa 435

F—689780, Gg——154936, La, W-~1317854). Nuevo Leén: O. M. Clark
6814 (Ok-~18717); Graham & Johnston 1640 (Mi). Oaxaca: Martinez-
Calderén 195 (Ca~936137, Me, Rf); Nelson 2718 (W—228713); Or-
cutt 5221 (E (E—711862); J. V. Santos ; 262 (M4) 5 Ll. Williams 981)
(F—898110, N). Quintana Roo: G. F G. F. ~Gaumer 1688 8 (B); Lundell & &
Lundell 7810 (Mi, N, Rf). San Luis Potosi: M. T. Edwards 1,55 (au,
Du--278627, | 7915215, N), 609 (Du—276821, F—915189); G. L.
Fisher 45168 (Du--3)9953), 115169 (Ew); Hitchcock & Stanford | 696
(W-~1806795); Kenoyer 02 (Mi), 4.608 (F—1000786), A.631 (F—
100003), s.n. [Tamazunchale, 8-29-38] (Fs); W. C. Leavenworth
226 (Mi, N, Ur Ur, Ur); LeSueur 25 (F-~1003633); | ; Lundell & Lundell
7108 (Mh, i, N); R. McVaugh 10489 (Mi); J. Rzedowski 7827 (Ip,
Ip). Tanaulipas: Barkley, Webster, & Paxson 664 (N); V. H. Chase

7587 (F—1001717, Fs, Mi, Ur); 0. M. Clark 661h (F1--228);
Crutchfield & Johnston 5799 (is) 5 Dressler I9hla (Mi); Hitchcock
& Stanford 6868 (Ca-—710768, Du—315601, N, “Or—5L912, P1——130110,
Po-=266550, ~Se——58670, Ur); Kenoyer & Crum 3292 (Mi), 3316 (Mi);
Meyer & Rogers 21,87 (N); Edw. Palmer “il, (Ca--717602, E—T7775h6,
F—136173, W—ls63050), 174 (Ca—153287, E—118813, F—217514, W—
57204), 388 (Ca—14569h, hh, E—118816, F—217702, W-572619), 495
(E—778585, | F—)36h,29, W--1163)06) ; Rutten & Rutten-Peke elharing
Ugh (Ut); J. Rzedowski 10345 (Mi); Wooton s.n. (Buena Vista] (Mi—
Photo, W-~98973). Veracruz: Ervendberg 212 dberg 212 (D-~612060, P);
Fournier s.n. (Laguna de Tampico, 1838] (P)j Goldman 710 (W—
397080); Gouin s.n. [1867] (P, P)} Liebmann 11165 (W—1315035),
a Ne Tiree 1301 (W—-131508)); Edw fen, Eper 6 2s ve =
iF aray 3159 (Ip); Purpus 437 (Ca—143278, E119
119225, F-=3LL052- We-814833), 8070 (Ca—218881, Du—20699l,, ‘Po—
aphex, 3 . V. Santos 2101 (Mi), > 2651 (Mi). Yucatdn: ad 26

(T0785. W—-1,77671), 80 (I--photo); G. F. Gaumer 68ha (E—
951705), 870 (E—119268, F—36673, Us, W--571806), 1688 (E-—
852861, F——58)86, W--1267)0h), 24071 (E—-948114, F—552075, Po—
171965, W—-1268166), s.n. [Buena Vista, 1899] (F—187286, Ge—-
160139, Us, V); Langman an 332 (w--2022085) 5 Linden Linden s.n. [Juillet]
(P); Lundell & Lundell 7519 (Du—363000, |, Mi, N)j Steere 1004
(E—108722L, F--€68]93 Ch TIT (Me); S as rda 15> (F—890h93).
State undetermined: Houstoun s.n. (P); Sessé, Mocifio, Castillo, &
Waldonado 517 (F—850927, Q). GUATEMALA: Alta Verapaz: R. M.

3354 (Mi, N); Tejeda 128 (W--862087); Ttrckheim sieaiteee a (E-=119269,

meme W—398505, W—13232h7). Chiquimla: Ae
31737 (F~103),965) « El Petén: Aguilar Hi z (ou-—239585, E—
1067855, Poriia vo) ; Contreras 2213 (Ld, S), 2705 (Ld, S), 3339

» S)3 C. L. Lundell 2088 (F (685323), asi ~ (F—693905), 3027
(P=26959585, 3635 (ii), 3755 (F-68972, Gg——2h8095, Mi, S), » LOw02

436 PHYTOLOGIA Vol. 13, no. 6
(Ld, S). Izabal: H. Pittier 8622 (W—-1013513); P. C. Standl

24399 (W—115001). Zacapa: C. C. Deam 6300 (E—119218, Ed, F—
285137, Vt, W--579609); Record | & Kuylen G.130 (W—1315339, Y—-
10081). BRITISH pal Chanek 198 (F—7019h9); Gentle 25 (F—
696464), 199 (Mi, N); C. L. Lundell ie (F—683522); Winzerling
III.3 (W--1266653). ane Atlantida: E.R. Mitchell 70 (F—
580597); P. C. Standley 55185 masters: W—-108711). Comayagua:
J.P. Edwards P.616 (B, Ca—522750, F—688148, W—1588697). Cor-
tés: Yuncker uncker 883 (: (Dp, E--1087012, F-79089) « Santa Barbara:
Thieme 5507 (W—3550)2, W--1323251). BAY ISLANDS: sae G. Fe
Gaumer 6h, (Sg—1605),, W-56137) . COSTA RICA: Cartago: Leén
1597 [Herb. Inst. Interamer. Turrialba 596] Asset oth os 1685
[Herb. Inst. Interamer., Turrialba 871] (W—-2021)91); H. Pittier
3173 (Br, Br); Standley & Valerio 46750 (W--125483h) . ~~ Tanén:
Kuntze 1968 (F—297690); 1 Morley 809 (Ca—756765) ; H. Pittier 425k
(W—1323253), s sen. [Herb. Inst. Physico-geogr. Nat. Costaric. __
9741] (Br, Br, W—1323255), s.n. [Herb. Inst. peyalae- eee Nat.
Costaric, 12688] (W--1323252); J. D. Smith hoe (W—13232h9)5 Fe
L. Stevens 732 (W—~1166552). Puntarenas: P. H. Allen 528) (N)j_
Brenes 12267 (F—855893), 12339 [218 & 818] (F--856001); Cufodon-
tis 1 117 (N) 3 Tonduz 6819 (Br, W—-1323250); M. Valerio 22 in
668656). San José: Tonduz 779 (W~355566, W—13232h8), sete
(Herb. Inst. Physico-geogr. Nat. Costaric. 12788] (W—354272
35876). PANAMA: Bocas del Toro: G. P. Cooper 555 (pe 196s8,
1521698, Y—12188); H. von Wedel 2666 (N). Canal Zone: Pi
5u57 (W—-1166031, W—1166032). Coclé: P. He Allen 2488 (N)- Co-
1én: G. P, Cooper 220 (Ca--5),9869, F—-765683, H--38652, N), a
(F—76568h, N), 654 (F—579218, W—-1521215, Y--12287). PanamA: He
et 5520 (W—715771). PEARL ISLANDS: San José: Cc. 0. Erlanson
: Antioquia: Brahe , Araque Molina, g Barkley
ec 70 (N); Haught 4868 (W—1709525); Toro Toro 1293 (Fn——1666)-
Cauca: Triana 3712 By: 186] (Je). Cundinamarca: Mutis 707 (Cb,
F—712886, W--1561452). Norte de Santander: Schlim 79 (B—-photo,
Br, S—photo, cies 68). Tolima: Bonpland 169) [Macbride photos
39492] (F—~1038391—photo of type & isotype, Kr--photo of type
isotype, N-photo of type, N--photo of type & isotype, P-—iso
S--photo of type, Z--photo of type), sn. (Quindiu] (P); Cuatre-
casas 91462 eee) (#1850801) Hartweg 1315 (Lu); Haught 24127
(N). Valle del Cauca: Cuatrecasas 22883 (F—-1341700); Fe Ce 3
mann 7703 (r—sT6035, W--938508), K.286 (F--578236). Department
undetermined: 8.n, (E~photo, N—photo, P, P, Z—photo)j
Linden 1196 aoe VENEZUELA: Lard: Badillo 72 (ve) . Trujillo:
J. A. Steyermark 56771 (N). ECUADOR: Los Rfos: Asplund 5410 (S, *
1930520). PERU: San Mart{n: Klug 3879 (E—110h815, F--766387, 08-
248620, I, N, S, W—1)58287), 44) (Ca--710140, E—11108)0, F—

1966 Moldenke, Monograph of Callicarpa 437

853163, Gg--2)7856, I, N, S, W--1458558); Ll. Williams 6817 (N);
Woytkowski 51:58 [wood voucher 15524] (W--2hi3i7h), 7283 (W--
2453294). Department undetermined: Herb. Pavon s.n. [Peruvia]
(X). BOLIVIA: La Paz: M. Bang 1669 (E—~119256, Ed, F—-77781, Mu-
1730, Pa, Vu, W—-56135, W—56136); M. CArdenas 1768 (W~-1232)16).
CULTIVATED: knatehae Herb. Hort. Schtnbrunn s «1 s.n. fe (V)3 H Herb. Hort.
Vindob. s.n. [9/1803] CUyo? 43

CALLICARPA ACUTIDENS Schau, in hb DC., Prodr. 11: 65. 187.
Bibliography: Schau, in weg roe Sep — 1875 Jacks.
in Hook. f. & Jacks., Ind. ew, 1: 386. « Soc
di Nat. Toulouse by: 198-501, "sesso, a i. 19325 X Moldenke,
Geogr, Distrib. Verbenac., » 58 & 86 (1942) and ed. 2,
138 & & ak 1949; Moldenke, Résumé eto & 3. 1959.
ets densely grayish floccose-tomentose; leaves lanceo-
lato-oblong, about ire cme long and | cm. wide, short-petiolate,
de:

margins render the species very distinct.
Dop (1932) cites Eberhardt 2621 and Gaudichaud 125 [probably
type] from Annam, Indochina, and comments: "Cette = est
heii du C, macrophylla Vahl. Em particulier la présence sur
les deux faces de la feuille de poils étoilés rapproche eo
les deux espéces. Dans la revision Bakhuizen van der Brink C.
u rang de synonyme de C. cuspidata Roxb. Or
celle derniér espéce est définie par cet auteur comme ayant des

C. macrophylla. Ilya = une erreur évidente C. acutidens ne
tre r me uspldate, mais placé a c8te de C, macro-

Phylla, dont la ai s feuilles et leur découpure

ale la séparent nettement. et est & remarquer que dans la forme

CALLICARPA ACUTIFOLIA Chang, Acta Phytotax. Sin. 1: 28). 1951.
Bibliography: Chang, Acta Phytotax. Sin. 1: 28h. 1951; G. Tay-
lor, Ind. Kew. Suppl. 13: 21. 1966.

4,38 PHYTOLOGIA Vol. 13, no. 6

othing is known to me about this taxon except that it is sup-
sa to be native to Kwangtung, China. The eT publication
has not as yet been available to me for consultation

CALLICARPA ALBIDO-TOMENTELLA Merr., Philip. Journ. Sci. Bot. 12:

1917.
Bibliography: E. D. Merr., Philip. Journ. Sci. Bot. 12: 300-—-
301. 1917; E. D. Merr., am. Philip. fags et que A.W. Hill,

Su
Verbenac., ed. 1, 61 & 86 (2912) and ai — rae wat a 19493 “Yol-
denke, Résumé 182 & 13. 1959.
+ 2m. tall; branches terete, grayish, glabrous;
ranchlets very densely and minutely grayish- or whitish-puberv-
Tent with stellate indumentum; leaves “decussate-opposite; petioles
ane b ran-

dens
ous, lanceolate, 7--11 cm. long, 1.5--2.5 cm. wide, gradually nar-
rowed upwards to the slender 8 acuminate apex, entire,
narrowed below to es obtuse base, the base of some leaves slight-

ly inequilateral, the upper surface brownish-olivaceous, glabrous,
entirely

scured by the indumentum beneath; inflorescence axillary, 1.5--2
cm. long, slender, pedunculate, dic hotomous, few-flowered, densely
stellate-puberulent, the peduncles surpassing the primary branches

with very short white hair, the rim truncate or very obscurely l-
denticulate; fruit subglobose, purplish when fresh, about 2.5 mm.
de, glabrous.

The type of this little-known species was collected ers Maximo
Ramos [Herb. Philip. Bur. Sci: 26976] pepe a small stream in the
of ab

the province of Abra, Luzon, Philippine Islands, on paras 5D»

1917, and was deposited in the herbarium of the Phili

of Science at Manila, but is now des . Merrill (isi?) = ied

that this is "a most characteristic species, distinguish

white or grayish-white, dense indumentum, ms or
’

ves whi en wt
the upper surface. It somewhat resembles C. angusta Schauer,

is not ogee allied to that species." Nothing is knom to m™
this except what is given in the literature.

ieee ALONGENSIS Dop, Bull. Soc. Hist, Nat. Toulouse 64: 509+

932.
Bibliography: P. Dop . Soc. Hist. Nat. Toulouse 64: 500,
501, 509, sli,” i Siz. 13325 A A. W. Hill, Ind. Kew. Semple oe
19385 Moldenke, Known Geogr. Distrib. Ver t 86
(1942) and ed.’2, 135 & 196, 1949; Moldenke, ideas ‘8 & Lh3.
"Shrub; ranchlets subtetragonal, lightly fulvous-pubescent;

1966 Moldenke, Monograph of Callicarpa 439

leaves decussate-opposite; petioles 6--10 mm. long; leaf-blades
chartaceous, rigid, 10—13 cm. long, 2--3 cm. wide, acute and
uminate-caudate at the apex, denticulate along the margins ex-
cept at the base, attemmate and acute at the base, stellate-
e W.

pubescent abo en ure, but glabrous and brunneous when
mature, lightly short-tomentose th whitish stella
hairs; mi P ent beneath; secondaries er, 20--22, as-

many-flowered, 2--2.5 cm. long, 1.5--2 cm. wide, dichotomous,
fulvous stellate-pubescent; bractlets minute, subulate; flowers
very small; calyx conic, 1 mm. long, stellate-tomentose, the rim

.
me 3 style equaling the stamens; st
pe tM Late-pubesvont and glandular; drupes black, 1 m. a
Bvitte aes
The type ns ot pagreyte was collected by 0. Debeaux (no. 300)

ses fleurs de moitié plus peretees On peut la considérer comme
une espéce géographique dérivée du C. acuta." 7 species to
eas: he refers here is altiet Cc. , angusta Scha

I know nothing of C. alo ensis | except what te cgtven in the
literature quoted above ee Rarer

CALLICARPA AMERICANA L., Sp. Pl., ed. 1, 1: 111. 1753 [not S: amer-
icana Blanco, 188), nor Hort., 1936, nor Lour., 179h,
Sessé & Moc., 1893].

Additional & emended synonymy: Anonymos baccifera verticillata,
folio molli et incano, ex inavtns tg Almagest. Bot. 33, pl.
136, fig. 3. 1696. Frutex baccif baccifer verticillatus, foliis scabris
latis dentatis et conjugatis 3 Catesb., Carol. 2: 47. 1733. ” Frutex
foliis amplis subrotundis acuminatis ex adverso binis, viminibus |
lentis infirmis, quasi le quasi levi canitie tectis Gronov., Fl. Virg. , Virg. 138.
1739. Johnsonia floribus verticillatus sessilibus T. Dale ex P.
Mill., Gard. Dict., ed. 7. 1759. Burchardia callicarpa Crantz,
Inst. Rei Herb. 2: 360. 1766. Callic fol. serratis subtus to-
mentosus L., Mant. 2: 33, 198, & 331. Let Johnsonia americana
Mill., Gard. Dict., ed. 8. 1768. Frutex baccifer verticillatus,
foliis scabris latis dentatis & conjugatis Catesb. apud Houtt.,
lin. Pfl. Syst. 3: 101, in syn. 1778. Callicarpa foliis serratis,
subtus tomentoeis apud Houtt., Lin. Pfl. Syst. 3: 101, “in syn.
1778. Anonymos baccifera verticillata, folio molli & incano, ex

eae sa ae” ed

AR) PHYTOLOGIA Vol. 13, no. 6

America Pluk. apud Houtt., Lin. Pfl. Syst. 3: 101, in syn. 1778.
Frutex . foliis amplis subrotundis acuminatis ex adverso binis, vi-
minibus lentis infirmis quasi leni canitie tectis Gronov. apud
Houtt., Lin. Pfl. Syst. Syst. 3: 101, in syn. syn. 1778. ca Callicarpa foliis
aaa acutis serratis, subtus subtomentosis, baccis glomeratis
Lan., Lam., Encycl. Méth. Bot. 1: vs 5630 1783. Callicarpa foliis serratis
subtus tomentosis L. apud J. A. Murr. in L., Syst. Veg., ed. lk,
153, in syn. 178). Anonymus &c. Pluk. apud Vitm., Sum. Pl. 1:
307, in syn. 1789. Frutex &c. . Catesb. apud Vitm., Sum. Pl. 1:
307, in syn. 1789. Anonymus is baccifera, verticillata, folio molli,
& incano Pluk. apud Lour., Fl. Cochinch., ed. 1, 1: 70, in syn.
1790. Anonymos baccifera verticillata, folio molli & incane Pluk.
apud Gaertn., Fruct. & S qm. Pl. 2: 60, in syn. 1791.
baccifer verticillatus foliis scabris latis dentatis & en
Catesb. apud Gaertn., Lh pe & Sem. Pl. 2: 80, in syn. 1791.
Anonymus baccifera, verticillata, folio molli et incano Pluk. a-
pud Lour., Fl. Cochinch., ed. 2, 1: 88, in syn. 1793. Callicarpa
serrata Moench, Meth. 168. 1794. Callicarpa viburnifolia Salisb.,
Prodr, Stirp. Chap. Allert. 53. 1796. Johnsonia (americana)
floribus verticillatis sessilibus, foliis ovato-lanceolatis op-
positis, caule fruticoso Mill. apud Willd Willd., Linn. Sp. Pl. 1: 619,
in syn. 1797. Anonymus baccifera verticillata, folio molli “
incano, ex America Pluk. apud Willd., Linn. Sp. Pl. 1: 619, in
syn. 1797. Frutex baccifer verticillatus, foliis scabris lati
dentatis et coniugatis Catesb. apud Willd., Linn. Sp. Pl. 1 est 19;
in syn. 1797. Frutex foliis amplis subrotundis acuminatis ex ad-
verso binis, viminibus lentis infirmis quasi leni canitie tectis
Gronov. apud Willd., Linn. Sp Sp. Pl. 1: 619, in syn. 1797. Calli-
carpa americana; foliis ovatis acuminatis inaequaliter obtuse obtuse
dentatis basi cuneato-attenuatis integerrimis subtus r ramulisque
leviter tomentosis, calycibus nudiusculis Willd. apud ere &
Semult., Syst. Veg. 3: 93, in syn. 1618. Callicarpa ramis
pruinoso-tomentosis, foliis lato~ovalibus, utrinqu eae , den-
tatis subtus subtomentosis, cymis sessilibus petiolo “previoribus
Michx. apud Roem, & Schult., Syst. Veg. 3: 93, in syn. 1618. Cal-
licarpa americana Willd, ex Roxb., Fl. Ind., ed. 1 (Carey & Wall.],
1: 407, in obs. 1820. Johnsonia americana Moench ex Steud., Nom.
Bot., ed. 1, 137, in syn. 1821. Anonymos baccifera etc. Pluk. 4
pud Walp., Repert. hk: 127, in syn. Te Frutex baccifer verticil-
latus ete. Catesb. apud Walp., Repert. lj: — in syn. 16 ayn. BLS. Oe

Jacks., Ind. Kew. 1: 361, iw syn. lees, Callicarpa amerikana L.
ex Moldenke in Fedde, Repert. Spec. Nov. 39: 30, in syn. 1936. =
Callicarpa villosa Baldw.. a ‘Moldenke in Fedde, Repert. Spec. Nov-

1966 Moldenke, Monograph of Callicarpa 4h

39: 305, in syn. 1936 [not C. villosa Roxb., 181}, Vahl,
1794). Callicarpa americana var. ar. purpurea ea F. J. rammed sat Mol-
denke in Fedde, Repert. Spec. Nov. 39: 305, in syn. 1936. Calli-
carpia americana L. ex Moldenke, Prelim. Alph. List Invalid Names
13, in syn. 1940. Callicarpa virginiana L. ex Moldenke, Suppl.
List Invalid Names 2, in syn. 19). Gal Licarpus americana L. ex
Moldenke, Suppl. List Invalid Names 2, in syn. 191. SEE
americana L. ex Moldenke, Suppl. List Invalid Names 1, in syn
19hl. Ca Calicarpus americana L. ex Moldenke, Suppl. List Invalid
Names 1, in syn. , in syn. 1941. C Callicarpa americana Roxb. ex Roig y Mesa,
Plant, Vedic. 775, sphalm. 1945. Calicarpe americans alicarpa americana L., in
herb. Callicarpa americana Lam., in herb

Bibliography: Pluk., Alnegeets Bot. 33, ple ae Poy 3. 1696;
Catesb., Nat. Hist. Carol., ed. 1, 2: Wa, *pl. 7. 3 Je &
Gronov., Fl. Virg., ed. 1, pr. 1, 138. 1739; L., Et =
Sei. Ups. 80. 17h; J. F. Gronov., Fl. Virg., ed. 1, pr. 2, 138.

Donald [R. W. Dickson], Comp. Dict. Pract. rn ey
Willd., Enum. Hort. Berol. 1: 157~- 1809; W. T. Ait., Hort. Kew.,

hh2 PHYTOLOGIA Vol. 13, no. 6

ed. 2, 1: 2h6—-27. — Poir., Encycl. Méth. Suppl. 2: one neg
Pursh, Fl. Amer. Sept. : 96. 161k; Raf., Fl. Ludovic.
Schul =a st. Veg. 3: 93. 1818; H.B.K., Nov. 2
& Sp. Pl. 2: 252. 1818; Sims in Curtis, Bot. lag. 7: pl. 2107,
in obs. 1819; Michx., Fl. Amer. Bor., ed. 2, 1: 95. 1820; Roxb.,
hos. 1820; Steud., Nom. Bot., ed. qi 137. 1821; Link,

-, ed. 16, 419. 3 J. A. & Je He Schult., Mant. 3: 50 & a
1827; Reichenb. in Mvssler, Handb. G k., ed. 199.
1827; Spreng. in L . Veg., ed. 16, 5: 126. 1828; Croom &

ng. in L., Syst 16 °
, Cat. Pl. Neighb. New Bern , 1833; Nutt., Trans. Am. Phil-
osoph. Soc., new ser., 5: 185. 183); Bac iman, Cat. Phaenog. Pl. &
Ferns 5. 1834; Kostel., Allgem. Mediz .-pharn. Fl. 3: 828. 183h;
Barton, Elem, Bot., ed. 2, 222, pl. 10, fig. 3. 1836; a Comp.
t. Mag. 33 Bb 1836; Croom, cat. Pl. Native or pe a byes
Bern 25. 1837; Endl., Gen. Pl. 637. 1838; Hook. &
Beechey ig “hs. 1838} D. Dietr., Syn. Pl. 1: 28. 18595 Patel,
Cod. Bot. Linn. Ind. Alph. 3. 1810; Steud., Nom. Bot., ;
oh 18105 praritione "Birds of Amer. 2: pl. 86. 18); porais “sp. Pl.
1: 342. 18h2; Walp., Repert. h: 126--127. 1845; West. Journ. Med.
& Surg., new ser., 3 (6): 1). 1815; Jacques & Hérincq, Man. Gén.
Fl. Arb. & Arbust. [Fl. Jard. Europe] 3: 502. eee Lindl.,
Veg. Kingd. 663. 1846; Schau. in A. DC., Prodr hh.
1847; Porcher, Med.—bot. Cat. oy & Ferns 12. 1 Leh7; White, sercerae

. Dia

Ler Verbenac.} 129 & 192. 1863; A. W. Chapm., Fl. South. U. S.,
ed. 1, pr. 2, 309. 1865; Miq., Am. Mus. Bot. Lugd. ‘Bat. 2: 98-

1865; Griseb., Cat. Pl. Cub. 216. 1866} "E. Pritz., Icon. Bot.

Ind. 1: 188, 1866; M. ne Curtis, Geol. & Nat. Hist. Surv. N. 0. 3

LO. 1867; Sauvalle, Fl, Cub. 12. “6s, A. Gray, School & Field

29. AO P. Herderson, Handb. Pl. 3. 1881; A. Gray, School

Field Book Bot., pr. 5, 23. 1881; Nichols., Haustr. Dict. Gard.
° ham, F tere South. Us a ° 2, pre 1, re

188); Wood & McCarthy, W. m Fl. 36. 1 886; Maxin., aélang

Biol. 12: 507 & 508. 1806; Ge qettiokes, Tenn. Fl. 71. 1887; A+ are,

1966 Moldenke, Monograph of Callicarpa 43

South. U. S., ed. . * 1887; gatadhtranse gr EE

ac 18. 1888; F. J. faite . Gard. 10: 463. 1889; A. Wood, New
Am, Bot. & Flor., ed. Willis, 237. 1889; A. W. Chapm., Fl. Sou
U. S., ed. 2, pr. 3, 309. 1889; P. Hende Handb. Pl.

in Hook, f. & Jacks., Ind. Kew. 1: 386. 1893; Sess6 & Moc., Pl.
Nou. Hisp. 2: 18. 1893; Bois, Dict. Hort. 232. 18931899; Sessé
Moc., Fl. Mex., ed. 2, 17. 1894; Briq. in Engl. & Prantl, Nat.

164: [Fl. N. Garol.] 332. 1899; Ae S. Hitchc., Trans. Kans. Acad.
Sci. 16: 147. 1899; Kearney, Contrib. U. S. Nat. Herb. S: 317.
1900; Re. M. Harper, Bull. Torrey Bot. Club 27: 420. 1900; Kearney,
Science, new ser., 12: 832, 834, 835, & 840. 1900; Diels in Engl.,
hg Jahrb. 29: 51,8. 1900; Kearney, Contrib. U. S. Nat. Herb. Se

» 380, 456, & 539. 1901; Sabhinee Fl. Tenn. & Philos. Bot.
3. 1901; C. — Contrib. Us S Nat. Herb. 6: 695. 19015 N. Le
Britton, Man. Fl. North. States, ed. 1, 779. 1901; mag Earle

1

° Ala Agr °
Dict. Pract. Gard., ed. 1, 1: 156. 19023 J. K.

OCs
Suppl. it 12: J. Matsumura, Ind. Pl. 7g 4 2:
525-529, mares J at. saat, : ri. aes Ue S., ed. 2, 1015.
1913; Britton & Br., Illustr. Fl., ed. 2, 3: 99, fig. hs. 1913;
J. K. Small, Fl. Miami rug 1913; ‘ds Ke oe Small, Shrubs Fla.
1913; Hickel, Bull. Soc. Dendrol. France 28: jio—111, fig. lise.
19135 iol. Soc. Wash. e262 9» 19133 134 uehtee, Torreva

tie. Fla.
Geol. Surv. 6: [Geogr. & Veget. North. ar2.1 198, 213, 220, 245,
261, 2, 33, & ho. 191k; L. Hs Bailey, Stand. Cycl. Hort. 2: ” 628.

hhh PHYTOLOGIA Vol. 13, no. 6

1914; W. J. Bean, Trees & Shrubs Hardy Brit. Isles, ed. 1, 2: 280
& 103. Io F. S. Mathews, Field Book Am. reas 383 & 385. 1915;
R. . Rep. Fla. State Geol. Surv. 7: 169, 171, 17h

& 1%. 19155 Pollard in W ebster, New Internat. Dict. Eng. Lang.

312. 1917; J. ; « Y. Bot, Gard. 18: 99-~100. 1917;
Free, Fls, Winter 1. 1917; G. V. N Addisonia 3: 5, pl. 103.
1918; W. Trelease, Wint. Bot., ed. 1, 331. 1918; N. L. Britton,
Fl. Bermuda 318 & 319. 1918; R. M. .; gy 1: 202. 1920;

c
M. S. Young, Univ. Tex. Bull. 2065: [Seed Pl. Austin] 66. 1920;
R. M. Harper, Bull. Torrey Bot. Club 7: 305 & 309. a Britton
& Millsp., Bahama Fl. 373. 1920; R. M. Harper, Ann.
aye Geol. tei 13: [Geogr. Cent. Fla.] 100, 106, & "16. 1921;
> Bui. « State Geol. Surv. 17: 237. 1921; *z. D. Schulz,
500 Wild Fls. cent or Olmstead, Coville, & Kelsey, Stand. Pl.
Names, ed. 1, 61. 192; P. C. Standl., Contrib. U. S. Nat. He .
23: 1253. 192k; W. paces, Wint. Bot., ed. 2, Rea 19253 age
Bean, Trees & Shrubs Hardy Brit. Isles, ed. h, 1: 280. 19253 B
ker, Fla. Wild Fls. 192. 1926; i Man. sip Wiegs: ed. te
775 & 776. 1927; R. M. Harper, Ann. Rep. Fla. State Geol. Surv.
18: [Nat. Res. South. Fla.] 10h & 108. 1927; Walcott, N. Am. Wild
ig 3: pl. 210. weed Hottes, Book Shrubs, ed. 1, 147. 1928; E.
Schulz, Tex. Wild Fls. 3h]. 1928; R. M.

1929; W. J. Bean, Trees & Shrubs sles, ed. ‘leg J : 280.
1929} Stapf, Ind. Lond. 1: 525. A929 3 Behe , List S hao
east. U.S. A. Set 10. 1930 Bailey,
iil, 1930; Me? in Karst. & hort ag Vegetationsbild. a1 a2):
nD. 1930; Macself, Sanders Encycl. Gard., ed. 21, pr. 1, 7i-
1931; poo Book’ ane ed. 2, 168. 1931; P'ei, Verbenac. China
1h & 36. 19323 P. Dop, Bul . Soc. Hist. Nat. Toulouse 6): 198.

1932; Grey & Hubbard, ist Pl. Atkins Inst. 33. 1933; J. Ma
Man. Southeast. 4a 13. 119335 Macself, Sanders Encycl. Gard., ed.
1, pr. 2, 71. 193k; J. A. Harris Physico-chem. Prop. P.: PL. Sap
1,0, 175, & 193. 2955 "crevost & Pételot, Bull. Econ. Indochine
37: 1289. 193); . Alexander, Journ. N. Y. Bot. Gard. 35: 133.
1934; L. H. Satie, i ports $s Handl. Verbenac. n.p., ms3.
19353 C. A. o, La. Cons 28 bir h: 35. 1935; T- W. Sanders, En-
cycl. Gard. "1. 19% ogee Seb rad Bg rt. Spec.
Nov. 39: 296-258. ry Bg eet! 7 lag36) and ea My tes 92, ae
117-- S, ede I

eer ae » & 29. 1940; Rehd., Man. Cult. Trees, ed. 2, PY
” sok; me 932. 29105 Moldenke, Carnegie Inst. Wash. Publ.
sees “Be. 190; M enke, Phytologia 1: 497. 191; mae Suppl +
nvalid ye & 2. 1941; Geiser, Field & Lab. 9: bh. 19l-

TWO NEW SPECIES OF LIPPIA

Harold N. Moldenke

LIPPIA CULMENICOLA Moldenke, sp. nov.

Arbor; foliis inter anthesin parvissimis ca. 1 cm. longis m.
latis ellipticis utrinque pilosulis ad apicem et bas in acutis,
supra mediam pauciserratis; capitulis numerosis breviss e ped -
culatis; pedunculis filiformibus 4--8 mm. longis puberulis; capit-
ulis parvis 5--10 m, latis, 5 m. longis; bracteolis purpure
pilosulis.

Tree, to 7m. tall, practically leafless during anthesis;
branches numerous, very twiggy, ue Aah glabrous; twigs nu-
merous, slender, gray, & glabrous, somewhat tetragonal and sulcate;
nodes annulate; principal i shabtoded abbreviated, 1—3.5 cm.
long; leaves decussate-opposite, very small and pro ly immature
i lepine :

pedunc.

h—-8 um. long, puberulent; heads small, subglobose or flattened,
about 5 mm. long, here mm. wide during anthesis; bractlets ovate,
Md m. long, about 3 mm, wide, subacute at the apex, purplish-

m when fresh, decidedly purple when dry, pilosulous; flowers
ihontded Aa. the bractlets.

of this distinctive species was collected by geet

MeV augh 23036) on summits of the driest hills near the oc
between Barra de Navidad and Playa de Cuastecomate, Jalisco,
ico, in a tropical deciduous forest with Forchammeria, Astronim,
and. Bursera, at an elevation of 50 meters abover sea-level, on
March 16, pasa sia i. deposited in the herbarium of the Univer-
ay of Michigan e collector notes that the species was "oc~
i

LIPPIA MICHOACANA Moldenke, sp. m

Arborescens; foliis inter pind Be parvis usque ad a cm.
longis 1 em. latis obovate-subrotundatis supra sparsis o-
sis, subtus in venis venulisque principalibus dense pilosis apicem
versus paucidentatis; inflorescen
pitulis breviter pedunculatis subglobosis 1
teolis viridulo-albis pilosis cdliolst tis.

Treelike, to 6 m. tall; bran hes and branchlets very twiggy,
glabrate, gray; an sondage more or less Deru pllnes
with elongate hairs; leaves decussate-opposite , small at time of
anthesis, very ssa OM or subsessil ; petiol es oe
to 1 mm. long, ely hirsutulous; peared cp Re obovate-subro
uring anthesis, rather Coens

1.5—2 cm. latis; brac-

dens
to 1.5 om. long and 1 cm. wide

46 PHYTOLOGIA Vol. 13, no. 6

seen on both surfaces, mostly rounded or retuse at the apex, a-
bacute at the base, usually with a very few rounded and

hirsute with long antrorsely spreading hairs; heads subglobose, a-
te

lose and jy gat, flowers included, inconspicuous; corolla green-
ish-white

The type of this species was collected by Rogers McVaugh (no.
22869) on hills in a deciduous forest ("now nearly leafless") a-

Zi
Mexico, with Bursera, Ipomoea, and various legumes, at 350--l5
meters altitude, on March 7, plaka and is deposited in the herbar-
ium of the University of Michigan

BOOK REVIEWS
Alma L. Moldenke

"Organization and Evolution in Plants by C. W. Wardlaw, xiii &
iPad illus. Longmans, Green & Company, Ltd., London W.l
9 sh.

Thi a very thoughtful, logical treatment of these important
and interne lated topics in plants In fact, the author treats
evolution as "essentially a progressive elaboration of organiza-
tion" and primary organization in any vascular plant as the "inte-
grated action of three main groups of factors": (1) those determin-
ing the inception of primary organogenic patterns in meristems,
(2) those contributing to the elaboration of maturing regions, (3)
specific genetical and ae Coahars determining the sequenti
syntheses of enzymes, horm etc. He describes fully and crit-
dolly’ such Vadtous thetites’ of organization as: holistic, morpho-
logical, physiological, physical and mathematical, protoplasmic,
genetical, and integrative

Listing here the ten principles of organization that the author
develops will indicate the approach of this book:
1. Organization involves energy and substances so interrelated and
1

1966 Moldenke, Book reviews Lh?

4. Biological patterns an expression of the attainment of dy-
nami ibrium in scion pie mone gp eg systems,
described as organismal reaction sy:

5. The way in which a particular oath caoiaie differentiated is

ni
teristically determined as a whole.
6. However essential, or limiting, in morphogenesis and organiza-
tion a particular a e may be, it can only produce its ef-

tem in an emb eg or meristematic cell or region.
7. Differently constituted reaction systems may yield comparable
primary patterns; and the same reaction syaten may yield differ-

- The progressive organization which becomes mani during on-
oocrarpoes = the resul = many interrelated serial processes;
d environn are involved in

ical, organismal en
the "infuction, and sere dp of the successive phases of develop-

9. the physiological reonre ty which determine reciprocal relation-
ships of various kinds contribute significantly to the organiza-
wal ol

10. Integration is an essential condition for the endurance of
smal reaction systems and for the development of anne
features,
There is much more of value in the book. The porn
are a definite asset. The most impressive one to me is yah fe 402
sh the long, convoluted, filamentous suspensor of Se

with the legend "It is hard to accept that the embryogenesis 0 ae

symnosperms i Been either precise adaptation or the principle
of economy". There is a useful index. The copy is neat and ac~
curate except fon t € slip. - "referred back" - on n-

- 303. My
ly regret is that the writing style is not conducive to compelled,
continued reading, to the probable loss to students assigned this
valuable reading.

"Botany: A Laboratory Manual" by Shirley R. Sparing, Walter H.
Muller, and William L. Theobald, viii & 181 pp., illus.
Macmillan Company, New York, & Gollier-Macmillan, Ltd.

London. 1966. $3.50

This looseleaf laboratory companion to the text "Botany: A
inctional Approach", by Walter R. Muller, one of the above listed

authors, is richly planned for a semester course. It can also
eas

tention to important items, the use of really good illustrations,
the — provoking questions, the reasonable limitation of
craw and the sending of the students outside the lab
for aut a materials are gp etongoen IG [Some instructors forget that

48 PHYTOLOGIA Vol. 13, no. 6

that is where plants live].
few suggestions for improvement would be: requesting sketches
on p. 5 to establish a visual image of assorted leaf types,

surely as to plant source on pp. 13, 36, 37, and 38, and adding
more modern and detailed experimental material on photosynthesis
and respiration.

"Primary Photoprocesses = Biology" or J. B. _—— vi & 323 pp.,
illus., North-Holland Publishing Company,
John Wiley & Sons., Inc. - ca cracmanee ees pec Division -
New York, 1965. $11.50

This book presents at first a general view of the biological,
chemical and physical uy yteen. and secondly and comparative-
ly individual considerations of the primary photoreactions of sev-
eral visible light-dependent ate Se through the formation
of products functioning only as initial reactants in the chain of
chemical "dark" reactions. Vital Suiskepheitinthe start with the
consistent act of light absorption and then follow an amazing nun-
ber of divergent processes. There is an excellent survey of the

ee
which the visible light emitted directly or rdeagrinoaingad is a
product rather than an agent in revious c

This book is well planned for the graduate or aavaneed undergrad-
uate level for ute in biophysics, biochemistry and related courses:
It is well indexed. The copy is neat except for the misspelling
of toe (ore 7 "ed on p. vi and the garbling of sentence struc-

PHYTOLOGIA is financed entirely by its contributors, each one paying
in advance for the entire cost of printing, binding, and distributing his con-
tribution. All money received from subscribers, after the expenses of col-
lections have been deducted, will be distributed among the contributors upon
the completion of a volume, in proportion to the space which they have used.
Each contributor is therefore a shareholder in the magazine, assuming his
part of the expenses and sharing in the profits, if any accrue.

Each number consists of not less than 32 pages. All manuscript accepted
will be published in the next issue, so that the size of numbers may vary
greatly. A volume will contain about 32 signatures, 512 pages, or a smaller
number of pages with an equivalent number of plates. This plan insures
immediate publication of all accepted manuscript.

port
tones, depending on their size, as fixed by the engraver, with a minimum of
about $2.25. .

Articles dealing with research in all lines of botany, in any reasonable

amateur or so-called popular type, and polemics will not be published. Advice
on the suitability of manuscripts will be solicited, if necessary, from quali-
fied botanists.

Under the present cost of printing, the hasic rate for a page or fraction
thereof is approximately $3.00 for an edition of 250 copies. This price is subject
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Reprints will be furnished at cost. A proportionate fraction of the edition

of 250 copies is also furnished gratis to contributors.

-quiries may be addressed to the magazine or to either editor.

PHYTOLOGIA

Designed to expedite botanical publication

Vol. 13 October, 1966 No. 7
CONTENTS

DEGENER, O., & DEGENER, I., Yes, thank you; we love ferns . . 449
MOLDENKE, A. L., Book reviews . - 452
SMITH, L. B., Notes on Bromeliaceae, XXIV. . -. «+ + +++ 454
MOLDENKE, H. N., Additional materials toward a agus

of the genus Galticaena: De es a ae pi. « 406
Index to authors in Volume Thirteen .. . 507
Index to supra-specific scientific names in Volume Thirteen 507
Publication dates of Volume Thirteen ... +. ++ +++ s+ se ee

yal ; Nn bet er

Published by Harold N. Moldenke and Alma L. Moldenke

0

303 Parkside Road
Plainfield, New Jersey, U.S.A.

Price of this —_— $1; per volume, $5.75, in advance,
; cme or $6 at close of volume .

or Ld : ;
. Volume 13, number 6, was issued September 19, 1966

YES, THANK YOU; WE LOVE FERNS
Otte Degener & Isa Degener

Dr. W. H. Wagner, Junior's article on "Pteridology in "
the Newsletter of the Hawaiian Botanical Society 2:

8 expressed to which we personally do
scribe. First, to quote: "I do not agree with oma teat
the Hawaiian flo ora will soon disappear and that 'all is lost'."
We believe ephemeral malihinis tend to lack the vision former and
present kamaainas gradually acquire. Dr. J. F. Rock, not long

we many
Plant friends had disappeared. How rapid Dr. Wagner's "soon" is
for definite areas is debatable. "Soon," for example, caught up
years ago with the dry native forest and i plains ef
West Molokai where the kane writer gathered such plants as Can-
thium, Gardenia, Nestegis, and Sesbania in 1928, all now repre-

sented in he one ep as =o to give | give an inkling as to what the
flora there was like. in 1928, he failed to unearth t
West Molokai Kokia pee ts at that time already extinct in the

wild state. "Soon" is ju: just aro around the corner for the Kanepun

forest, overrun with axis deer, of Lanai; the belt of forest

southeast of Ulupalakua, Maui; and the Sesbania colony of Kaena

Point, Oam. "Soon" is somewhat deferred for the flora peculiar
Kaala

engulf
by the planting of sugarcane; and for square miles of native for-
est on the same island which are being bulldozed for the planting

of Fraxinus uhdei. To dub as "propaganda" our efforts in behalf

is almost all
Possible ae these, as we mentioned
ra Hewaiiensis "fies" bon *is/63, as a distinct species
is not Salt ae a t at all.
The third statement that ee is now nigh for collecting

450 PatTOLOGIA Vol. 13, no. 7

by specialists, rather than crude gathering or "hay—baling' of
all plants in sight," we consider a half-truth. We workers today
feel frustrated when investigating a species to find that the

World. Our aim, living in a fast-vanishing endemic flora, is to
while th shines, we advise our amateur and pro-
ompetincy: colleagues to do likewise. Is it wise to lull them in
© complacency?

Were our public herbaria deprived of the riches collected by
such "amateurs" as the horticulturist Douglas; Chaplain Diell;
planter Baldwin; Director Brigham and the tubercular college
youth Horace Mann, Jr.; physician Hillebrand; rancher Munro; ac-
C Topping; and the youngster Rock who studied to enter the
priesthood and never took a formal college course in Botany; what

d

to Guire, Oba . and many
As it is impossible to publish authoritatively about any group
of plants whatsoever without familiarity with their literature,
we herewith add a Supplement to "Some Recent References on
Pteridology" appearing on page 123 of Dr. Wagner's article in the
November Newsletter. As our work is incamplete, we hope our
readers further supplements with the aim not only
of getting but for evermore keeping Dr. Merrill's famous biblio-
graphy of a
far as the Hawaiian Islands are concerned. To do so, the species
in the Merrill

posing in the Marie C. Neal Herbarium of the B. P. Bishop Museum.

Supplement 1 to Some Recent References to Hawaiian Pteridology

Caum, E. L., Notes Fl. & Fauna Lelma & Kaula Isl. Occas. Pap. B.
P. Bish. Mus. 11 (21): 1-17. 1936. [About Doryopteris decip-

1966 O. & I. Degener, Ferns 451

iens (Hook.) J. Smith]
eee C- a The genus Cyrtomium. Am. Fern Journ. 20 (2):
-52

Wi Fite hotetne ;” Caum, Vasc. Pl. - Bull. B. P. Bish.
Mus. er oe 7 1-1. ion. Uhtout Peete
Chrysler - Struct. Leaf. Gleich. Am. Journ, Bot. 30

(9); eth3. 1 1913.
Clausen, R. T., Monogr. Ophioglossaceae., Mem. Torr. Bot. Club 19
(23: "he “177. 1938.
Degener, 0., . lye. - Haw. Bot, Gaz. 80 (1): 26-7. gi
Degener & Shisahocerose: “71. Haw. Books 1-7. Fam. 1-21.
1932 - — Bete 19 966.
Degener, 0., & rn Gemus Unrec. Haw,
Phyto asa 9 a): "$586. shes 4 res Pa [See Ching, R. C.,
Acta Bot. Sinica 10 (3): 256. [Sept.] 1962]
age . Zie ee Beg Bearb. Spora. Leptosp. Farne Neur. Meth.
959
aor roy Mitchell, Checkl. Pl. Haw. Nat. Park. Nat. Hist. Bull.
93 1-75. 19h.
at F, R., Miscel. Notes Haw. Plants. Occas. Pap. B. P. Bish.
Mus. 16 (15): ge ipparie: bogar [About eris odus
(Schkuhr) oa Eas —_ - Notes Haw. Plants. ees - 2):
29=hh. 1962. [ bout Ophio ines pendul um var. falcatum
Presl
oer LF te « Pac. Isl. III. Bull. Torr. Bot. Club
+ 386-397, i9i3..
moabsay Os > Immi ants Haw. Isl. II. » Pap. Univ.
Haw. ies "f-37. 198. tibet Asolle isiicner Lam. ]
Fosberg Pe R., Upper Limits Veg. Mauna Loa. Haw. Ecol. 40 (1):
oe ah
Fosberg & Hosaka An Open Bog on Oahu. Occas. Pap. B. P. Bish.
1s (1): 2: 1938.
Fries, ay Am, och Krit. Geol. Ftren, FUrhandl. 71 (2): 347-355.

Hartt & Neal, Plant Ecol. Mauna Kea. Haw. Ecol, 21 (2): 237-266.

Hatheway, W. H., Comp. Cert. Nat. Dry For.: Mokuleia, Oahu, T. H.
Reol. Monogr. 22: 153-168. 1952.

Hawkes & Degener, The Upright Ps flotum. Am. Fern Journ, )0 (3):

Mery E. Y., Heol. & Flor. Stud. Kipspa Guleh, Oahu. Oceas. Pap.
. Bi

Neal, ve We List Vas . Mauna Kea, Aug. 1
Acad. Sc. B. P. Bish. ~ pec. Publ. 3h: 13. 1939
Ruble, G, Ce, Guide Sect., Isl. Maui, Haw. 1-9). 1959.

» Ow um Haw. Isl. Occas. Pap. B. P. Bish.

Aspleni
Mus. 19 (6): 171-18h. 1918.
’ ee pics mete Haw. Veg. Bot. Tidskrift 45 (1): 12=

Skotteberg, C., Remarks Fl. High Haw. Vole. Acta Horti Gotob. 63
47-65. 1930. —- Vasc. Plants Haw. Isl. Il. ibid. 10: 97-193.

452 PHYTOLOGIA Vol. 13, no. 7

1936.

Skottsberg r. Sven Berggren's Coll. Haw. Vasc. Pl. Bot.
Not. (iand) 19): state. 193.

Smith, F. G., Diellia and i s Variations. Occas. Pap. B. P. Bish.

St. John, oglos Occas. Pap. B. P. Bish.
Mus. 2 (28): *351-359. “sho. - sat ea Comb. Gleich.- - -, x
ibid. 17: 79-8). 192. - Ophioglossum Isl. Pac. ibid.
17 (13): 177-182. ee

St. John, H. » Dryopte: --. Bull. Torr. Bot. Club 72: 22=30.

Tryon, 2 M., Rev. Genus Doryopteris. Contr. Gray Herb. 143:1-80,
1942.

BOOK REVIERS
Alma L. Moldenke
apm oii pe ht in Higher Plants" edited by J. B. Pridham
2 pp., illus., Academic Press, London &
oon rates 75 sh. se $12.00
e valuable papers are the proceedings of the 196), Plant

studies.
Because the level of approach is both advanced and well ex-

plained this book can serve as a really useful source of informa-
rs all related

elds. Fine copy, diagrams, illustrations and indexes add to
the value of the book.
“Comparative Phytochemistry" edited by T. Swain, xiii & 360 ppe,

fahewe’ Academic Press, London & New York. 93 sh. or

at @ symposium by the Phytochemi al Group ia Gennetiee
in 1965 and represents thé organizational vos of one of the same
editors. Since it describes the of research of the most

progress
recent years of this newly established branch of science and the

1966 Moldenke, Book reviews 453

ications that may be dram, it will serve as a useful source
book to both botanists and chemists. Heywood's "Phytochemistry

d Taxonomy", Mentzer's "Biogenetic Classification of Plant Con-
stituents", and Alston's "Chemotaxonomy or Biochemical Sys t=
ics" are the introductory papers. They are followed by detailed
ones on the distribution and synthesis of alkanes, natural acety-
lenes, terpenoids, carotenoids, polysaccharides, asperulosides,
aucubins, ranunculin, sulphur compounds, amino acids, alkaloids
and th their phytochemistry, betacyanins, betaxanthins, hydroxy-
quinones, flavonoids, and dihydrochalones. The results of the

the above-mentioned
an error in sentence structure on p. 53), the diagrams
dexes helpful.

"The Romance of the Fungus World" by R. T. & F. W. Rolfe, xx &
- Johnson Reprint Corp., London W.1 & New York

309 pp., illus
10003. (original 1925) 1966.

Way back in my high school student days I was completely fas-
cinated with the first edition of this book and was so sure for a
few months at least that I would just have to become an amateur
Pie professional mycologist. The Rolfes, who set as their goal in

o sO

its charm, its ait enh eek accuracy, and its historical and literary

materia jals.

The chapters deal veh a i in mytholo Meher aap. and es
tive

h or changes

rain fall", and the fascinating hobbies of their rma study
their influence in history. On p. 76 "vitamins" are omitted
the list of essential foods even tho rngh they were known at the time

sonous and nonpoisonous mushrooms by decanting the fluid in which

ing by removing the noxious alkaloids, The many photographs do not
have the clarity of those in the original edition, but they are
still fairly good.

NOTES ON BROMELIACEAE, XXIV

Lyman B. Smith

UNITED STATES

TILLANDSIA BARTRAMII E11. Bot. S. C. & Ga. 1: 379. 1817.
Tangs Juncea | sensu LeConte, Ann. Lyc. N. Y. 2: 130. 1826;
Mon. 9: 676. 1896, wuoed pl. Florid., non Poir.
? Tillandsia eis LeConte, sa LYGeNewke £5 coke AAU,
Tillandsia tenuifolia sensu Mez in DC. Mon. 9: 674. 1896; Small,

ee Uso. 245. °1903; tit syn. Tillandsia bartramii

E
Tillandsia simulata Small, Man. Se. Fl. 270, 1503. 1933.
Tillandsia myriophylla Small, Man. Se. Fl. 270, 1503. 1933.
UNITED STATES: GEORGIA: On old trees in the Mortar Swamp,
Liberty County, Lewis LeConte s. n

Lewis LeConte s. n. (CHARL, holotype, phot. GH;
NY, isotype).

FLORIDA: Manatee River (Manatee County), June 1878, A. P.
Garber s. n. (NY, lectotype of Tillandsia simulata Small; US,

isotype).

setacea Sw. (T. tenuifolia
the remainder of this complex on its leaves

auctt., non L.) from

least 1 mm in diameter at mid-point (Pl. I, fig. 2: LeConte). I-
setacea is RT to the southern half of Florida, while the re-

mainder of the complex is rare there but common in the northern
half and gasdese Georgia

but so far in vain. The oldest
bartramii Ell. and judging from the descriptions, T-

pinifolia af aoe belongs here, while his T.

pears to synonym of T, ta Pp

species each been located as indicated above, but T. hystricina

remains a mystery with oo ae of relationship . elsewhere be-

1966 Smith, Notes on Bromeliaceae 455

cause of the leaves with "stiff nearly Beret te
s

use T. setacea and T. bartramii for the southern and northern
species respectively.

TILLANDSIA FASCICULATA Sw. var. CLAVISPICA Mez in DC. Mon. Phan.
9: 682. 6
Tillandsia bracteata Chapm. Fl. So. U. S. 471. 1860, non Vell.
1825.
UNITED STATES: FLORIDA: "South Florida", Chapman s. n. (NY,

lectotype of Tillandsia bracteata Chapm.).
Chapman's description of his Tillandsia bracteata is so ambi-

guous that is impossible to place it among the few pet 9: of
Florida without material, and that is poorly labelled. ever,
all material so labelled, whether by Chapman or later cue,
s T. fasciculata § In Florida, the common variety «tea
ciculata and undoubtedly the major element Chapman's species
is var. den Mez, but is interesting to note the occur-
rence of common Cuban var. clavispica Mez in southern Florida

all, bracteata Chapman is a later homonym and consequently its
typification is of minimal importance.

TILLANDSIA INCURVA Griseb. Nachr. Ges. Wiss. Goett. "1864": 15.
18

lorida record for Tillandsia incurva Griseb. (North Ameri-

can Flora 19: 148. 1938) was based upon an unn ed specimen
collected by Blodgett at Key West (NY). Although the form of its
lorescence very closely resembles that of _T. incurva, re-

leaf-sh I now believe that the specimen is a hybr a
do ant T. valenzuelana A. Rich. with some larger species suc
as T. fasciculata Sw. that wml account for the much larger flo-

ral bracts and sepals.

In any event, T. incurva should be removed from the list of
Florida species. Whether there will ever be an authentic record
is dubious, for although Cuban species like T. pruinosa ar -
nh pag in southern Florida from time to ime T. incurv
cies of mountain rainforest and could scarcely find a

ielesnae Saeed there.
MEXICO

PITCAIRNIA scored & L. B. Smith, sp.
jimenezii L. B. Smith atque P. a SEES Mart. in systema
mea come sed sepalis posterioribus alato-carinatis differt.
PLAN ith a short erect stem covered with old leaf- grin,
ig a ages: cm ag og LEAVES all alike, fasciculate at
of the stem, to 5 dm long, entire; sheaths si rt Ea

456 PHYTOLOGIA Vol. 13, no. 7

15-20 mm long, dark castaneous, glabrous at least with age;
blades linear, slightly contracted at base, filiform-acuminate,

at base. STAMENS included. OVARY 3/5 superior. SEEDS shortly
bicaudate. Pl. I, fig. 3: Flower x 1; fig. 4: Anterior sepal x
1; fig. 5: Posterior sepal x 1,

joc, alt. 1050 m, 30 October 1965, D. E. Breedlove 13991 (US,
type; DS). Chiapa, alt. 1350 m, M. B. Foster 2963 (US, paratype)

SALVADOR

GREIGIA ROHWEDERI L. B. Smith, sp. nov.

G. van~hyningii L. B. Smith, cujus bracteas valde imitans,
foliorum vaginis inconspicuis, bracteis primariis pallidioribus
serratisque differt.

PLANT known only from fragments. LEAVES over 2 meters long,
covered beneath with coarse pale appressed scales, soon glabrous
above; sheaths inconspicuous, pale brown toward base; blades
linear, acuminate, somewhat narrowed toward base, to 38 mm wide,

acuminate, pungent, 5-8 cm long pale c ou s
lepidote, entire, INFLORESCENCES complanate-capitate, com
t x

ral bracts narrowly lance-triangular, 33 mm long, thin except for
the pungent castaneous apex, entire, sparsely pale-lepidote.
SEPALS like the floral bracts, 23 mm iong. Pl. 1, fig :
Primary bract x ae fin" 3 Sepal x 1.

R: Santa Ana: Epiphytic and terrestrial in cloud forest
Hacienda Montecristo, north of Metapan, alt. 2200 m, 27 August

ous mostly entire primary bracts, and thus the characters taken
hw seem reasonably secure.

SS RO OS Oe ae oe aed SeCeeeCeoneeece

eee
eee ee ee eee ser eereeeneeeeeeesee

1966 Smith, Notes on Bromeliaceae 457
VENEZUELA

sme aoe ALTENSTEINII (Lk., Kl. & Otto) Lem. var. MINOR L. B.
» var. nov
A var. ARS OL scapi bracteis superioribus quam intern
“ids brevioribus, inflorescentia brevi, basi latibracteata ee Ton
SCAPE-BRACTS shorter than the ares gee ap eine INFLORESCENCE
short-cylindric or subglobose, to 11 ca long; basal floral bracts
bro net ee I, fig. 8: Inflorescence x ere
UELA: Aragua: Alto de Choroni, alt. 800-1400 m, V. M. Ba-
a. Are (Universidad Central de Venezuela). Same, alt. 800-
1300 1 m, 8 May 1949, Badillo 1928 (UC VEN). Same, 12 June 1952,
Schnee 953 CicvEN EN). Same, terrestrial in rich soil in full sun,
alt m, 5-6 April 1962, T. R. Soderstrom 975 (US, type).

PUYA FLOCCOSA (Linden) E. Morr. var. COMPACTA Lc Bu; Smithy) vars

A var. ges inflorescentiae ramis brevibus dense florige-
ris bracteas primarias subduplo pees es dif

INFLORESCENCE with branches 2-3 c m long i including the naked
Sterile base, about twice as long as the serrulate rose primary
bracts. Pl. I, fig. 9: Branch x 1/2.

VENEZUELA: Mérida: On rocks, July 1846, Funck & Schlim 1055
(P, type).

COLOMBIA

GUZMANIA BICOLOR L. B. Smith, sp. nov.

G. graciliore (André) Mez, cui affinis, foliorum vaginis om-
nino castaneis haud striatis, laminis poprene angustioribus,
scapi bracteis omnibus longe laminatis differ

LANT caulescent; stem at least 35 cm ae: elowsine shoot 20

beneath. § E erect, very slender; scape-bracts erect, densely

long. Pl. I, fig. 10: Branch x 1/2; fig, 11: Sepals x 1
COLOMBIA: Valle: Rocky hillside near quebrada, in guess near

km 58 on the Big eaT: between Cali and Buenaventura, 12 Aug

1965, F. A. Barkley & L. E. Willard 35472 (US, type).

eee eeee
Catt Me ee ee ee ee Re ee ee ee ae ee ee ee ae eee eee

458 PHYTOLOGIA Vol. 13, no. 7
BRAZIL

AECHMEA BAHTANA L. B. Smith, sp. nov.

Ae._tomentosa Mez atque Ae. stelligera L. B. Smith affinis, a
priore bracteis primariis angustioribus, axibus gracilibus, a
posteriore bracteis primariis omnibus magnis et ramos multo supe-
rantibus, a ambobus sepalis utrinque late alatis differt.

LANT flowering over 1 m high. LEAVES 7 dm long; sheaths el-

liptic ple, a ong, castaneous beneath toward base, sub-
densely vestite with appressed brown-centered scales; blades li-
ulate, ac ‘ cm wide, densely and finely purple-spotted,
subdensely lepidote beneath, becoming glabrous above bu sen-
tially concolorous, subde sely serrate, the al spin re-
curve on E erect, slender, densely vestite with

S ca. 3 cm long, bearing 2 short dentate scales at
base; blades elliptic, obtuse. STAMENS included; pollen nearly
spherical, ca. 50 u in diameter, reticulate, at least 6-porate.
OVARY cylindric, 10 mm long; epigynous tube cylindric, 2.5 mm
long. Pl. I, fig. 12: Branch x 1; fig. 13: Sepal x 1; fig. 14:
Pollen x ca. 500.

BRAZIL: Bahia: Rio de Contas, Bom Jesus, August 1913, Luetzel-
burg 300 (M, type; phot. US).

BROMELIA EITENORUM L. B. Smith, sp. nov.

5. antiacantha Bertol. atque B. balansae Mez affinis, a priore
bracteis florigeris sepalisque longis angustisque, a posteriore
bracteis florigeris sepalisque haud carinatis, a ambobus habitu
minore distinguenda.

PLANT stemless, flowering 8 dm high. LEAVES many in a bulbous
rosette, 75 cm long; sheaths ample, 4 cm long, dark castaneous,

also serrulate; blades ear, ac » pungent, 40 mm ’
white-lepidote beneath expecially between the nerves, soon gla-
brous above, laxly ser wit der uncinate brown 7 mm long

iptic, entire, the lowest short-
laminate, the upper bladeless, about half as long as the few-

1966 Smith, Notes on Bromeliaceae 459

flowered fascicles, subchartaceous, soon glabrous; floral bracts
oblong, obtuse, 2 cm long, about equaling the ovaries, thin
ecarinate, white, soon glabrous; flowers subsessile. SEPALS like
the floral bracts, 15 mm long. PETALS erect, 23 mm long, gla-
brous, white at base, dark dull purple above. TAMENS included;
filament-tube 8 mm long. OVARY slenderly ellipsoid, 2 cm long,
white-flocculose. FRUIT broadly ellipsoid to 4 cm long, nearly
even. Pl... fig.:15: Branch Pei 1/2; fig. 16: Sepal x 1.

IL; Maxanhfo. Mun. Lor@to: Edge of brook in flat "chapada"
or semi-closed woodland, "Ilha de Balsas" region, between the
Rios Balsas and Parnaiba, about 35 km south of Loréto, ca. Fe a
S, 45° 4' W, alt. 300 m, 15 April 1962, George & Liene T. Eiten
4312 (US, type; flowering) ; same, 3 April 1962, Eiten iten 3974 (US,
fruit).

ORTHOPHYTUM Beer, Flora 37: 347. 1854; L. B. Smith, Brom. Brazil
in Smithsonian Misc. 9 1262 179.3955.
Prantleia Mez in Mart. Fl. Pe Pt. Bt 25e6 ABO).
sincoraca Ule, Bot. Jahrb. re “i912. 1908.
Cryptanthopsis Ule, Bot. Jahrb. 42: 193. 1908.
Six additional species of Orthophytum have been discovered

t
resting to note that the aesctet es of citations from Goids and
Mato Grosso now limits the range of the genus to a s ies from
Minas Gerais and Espirito Pe to Bahia and Parafba

1. Inflorescence sunk in the center of the leaf-rosette or of
tminal leaves, — scape lacking or short and ale:
by the leaf-sheat
2. Plant caulescent, stent hing; oe seule triangular,
5- 0.

mm wide. —_— San QO. vagans
2. Plant begin pa s, but ——- per ae Nag or
3. L een tsa es linear, "30 cm long, the anbiee mostly antrorse;
rrowly triangular, 30 mm long.....2. 0. navioides

3. aah ee a narrowly triangular, 3-6 cm long, maximally 5-15
mm wide, the spines recurved; sepals 14 mm long.
4, Leaf-blades only 5 mm wide, the spines 1 mm long; flowers
alaetrpedtceLinles sepals lanceolate, acute.
oenum
4. =n 11-15 mm wide, the spines 2-3 mm Tosca ica
sile; sepals narrowly triangular, et
O. saxicola
1. cane either raised above the leaves on a ‘distin net
scape or itself elongate
5. Inflorescence short and pata
6. Spines of the leaf-blade eeeucced. 2-3 mm long; leaf-blade
3-6 cm long; inflorescence simple; floral bracts much
exceeding the flowers, recurved. Bahia.....4. 0. saxicola

oeeee
Cee eee eHee ee ees eeeeeeaegeeeeeere eee eeee

4,60 PHEYITOLOGIA Vol. 13, no. 7
ede perees of the leaf-blade predominantly antrorse; leaf-blades
h large

y= cade ‘oblong, 17-20 mm long, densely gh flowe

fasciculate, inflorescence Snaonn subglobose. Mine
RP ORT niet ate’ aie sive mello-barretoi

7. Sepals narrowly erienpulat, “12- 1s. mm Site 5 nck strongly
vestite.

8. Floral bracts straight, about equaling the flowers;
flowers spicate; inflorescence digitate. Bahia.

6. O. rubrum
8. Floral bracts recurved, the lower ones distinctly longer
tha e flowers.

a. Cael aitberade simple, spicate. Espirito Santo.

- O. fosterianum
9. a compound, densely appdata r Minas
ra Sig Ws bie Ww o's bbb’ o ov bce ote O. compactum
oe Setiseecanes elon ate, lax at Teaae eebiard
10. Upper

ee ase,

primary bracts Fient ike Many times exceeding the

fascicles of flowers

1l. Scape-bracts and p
ruptly a ees into very narrowly triangular

blades ; inflorescence dense toward apex; scape elongate.

E 9

biecy bracts with broadly ovate sheaths

irito Santo.) -2. seeessee 9. 0, foliosum

il. Seape-bracts and primary hoiets aeLehoue distinct
ate or linear, acuminate;

wip lax; scape short.

sheaths,

inflorescence almost

Minas Gerais, Espirito Santo.

10. O. duartei

10. Upper primary bracts short, scarcely more than twice as
long as the fascicles of flowers.

12. Scape-bracts lanceolate, abruptly acuminate; sepals 10 mm
long. Minas Gerais.

13. Leaf-spines 8 mm long; inflorescence lanate especially on
the axis ....11. 0. leprosum

io ps Laaf=spines 2 mm a dona taf loréecénte glabrous.

tp O. glabrum
12. Scape-bracts Pap eo long-caudat
44, ee simple, the bracts : suerne, but the
Espirito Santo. -7. O. fosterianum
14, Inflorescence compound, the larger henutis with fascicles
spikes of axillary flowers.
aD: Scales of the

st remote.

leaves wholly appressed, suborbicular,
entire; sepals 14-15 mm lon ng.
16. Inflorescence dense for about half of its length.
vole - O. maracasense
16. Inflorescence lax except ton: the: extreme apex
Bepitito Santee: ocr isicice i so.: mee | 0. sanctum
15. Scales of the “es more or less s apieadinl ga and lacerate
17, ei maximally 8-11 mm long; leaf-scales coarsely
erate, a deciduous above argon ie
17. Sepals maximally 14-15 mm long; leaf-scales finely and
eeply lacerate, Sightteneainas a on both sides.
Minas Gerais 16. 0

ere P magalhaesii

1966 Smith, Notes on Bromeliaceae 461

1. O. VAGANS M. B. Foster, Bromel. Soc. Bull. 10: OY, fies.

1960.

BRAZ

Janeiro, R. G. Wilson 578 (US, type).
Smith, Brom. Brazil in

Zz ‘
Smithsonian — Colle 1263 °34,.17961955, Cryptanthopsis navi-
mith, Contr. Gray Herb. 129: 31, pl. 3, figs. 4-6.

IL: Espirito Santo: Source unknown, cultivated in Rio de
)

9

BRAZIL: Bahia: On perpendicular rocks above stream in isolated
ravine, Jacobina, alt. 500 m, 16 June 1939, Foster 90 (GH, type;
ReelS)
3. 0. AMOENUM (Ule) L. B. Smith, Brom. Brazil in Smithsonian
Misc. Coll. 126: 33, 179. 1955. Sincoraea amoena Ule, Bot.
Jahrb. 42: 191, fig. 1 A-F. 1908; Mez, Pflanzenreich IV. 32: 9,
fig. 3. 1934. Pl. I, fig. 17: Flower x 1; fig. 18: Petal and
—- x if (after Ule).

ahia: On rocks, Serra do Sincora, alt. 1400 m,
Paced. 1906), Ule 7106 (B, type).
SAXICOLA (U: (Ule) L. B. Smith, Brom. Brazil in Smithsonian
Misc. Coll. 126: 34, 179. 1955. Cryptanthopsis saxicola Ule,
Bot. Jahrb. 42: 193, fig. 1 G-K. 1908. Pl. I, fig. 19: Flower x
a poet Ule).
RAZIL: Bahia: On rocks, near Maracas, alt. 1000 m, September

1906. Ule 7031 (B, type). heii 21 October 1948, Foster 2471-A
(US). “Very dr dry table rock area between Bahia and Milagres, alt alt.
450 m, 18 October 1948, tikes “2641 (US).

action in the key to genera in h
I followed this system in describing Cevptanthopais agviniiss
of

in 1940, but in 1948 Mulford Foster collected materia

raea Cryptanthopsis to Orthophytum is based then on the unre-

Root of the pollen character which has failed to correlate
with other floral characters in a n ases, as well as on
the positive characters of similarities of other Fevet structure
and of geographical distribution.

5. 0. MELLO-B I L. B. Smith, Bol. Mus. a Rio de

Janeiro nov. ser. no. 15: 2, figs. c-e.
IL: Minas Gerais: Mun. pes raage ubas: Palacio, Serra d
>

Saree do Cipo, Foster 631 (GH);

Pires & Black 2719 (IAN).
UBRUM L. B. Smith, Brom. Brazil in Smithsonian Misc.

462 PHYTOLOGIA Vol. 13, no. 7

— oars 34) 200, 7t te. .835-.1955.,
RAZIL: Bahia: Table Rock near Maracas, 1948, Foster 2444 (US,
ag
7. O. FOSTERIANUM L. B. Smith, Bromel. Soc. Bull. 8: 24, fig.
1958.
IL: Espirito Santo: Near Santa Teresa, 26 October 1948,
Foster 2487-A (US, type).
5 O. COMPACTUM L. B. Smith, sp.
foliosum L. B. Smith valde sonic, sed pes arate om-
compacta Sia interrupta, bracteis haud recurvato-serratis,
sepalis minoribus differt.
es; flowering shoot 2 dm h
red beneath with finely gpa eer
scales, soon glabrous above; sheaths small and inconspicuous;
te, 23 mm wide, laxly serrate with an-

tish; flowers sessile.
13 mm long, nerved,
PETALS easy bearing 2 highly adnate
scales, the blades elliptic, obtuse. NS included. Y
Narrowed at base, 8 mm high, strongly steed and alate with
the i Pls os fig. 20: Flower x l.
RAZIL: Minas Gerais: Terrestrial on rocky slopes by the high-

way from Nanuque . Teofilo Otoni, 14 August 1965, R. P. Belem
1626 (US, type; hb. Univ. Brasilia

« 0. F ge L. B. Smith, deck Bot. Estado SA Paulo nov.
SQLs bs. 58, 74. 1941.
BRAZIL: rerio Santo: On side
su

of 2 — in semi-moist condi-
tion but in full

gave a very iy ap idea of the geographic
diet efockia. of oethophy tum foliosum

hia By —

age

r, acuminate, laxly capemminernes with spreading oF
antrorse broad flat spines 0.5-1.5 mm long. SCAPE erect, slender

1966 Smith, Notes on Bromeliaceae 463

very short, naked or with a few foliaceous bracts. oie a ica

cicles; primary bracts foliaceous ostly large but the upper re-
duced; floral bracts straight, narrowly triangular, 10 ong,
exceeded by the sepals, entire, thin, rv pale-lep

owers sessile EPALS like the floral bracts but lanceolate,

mucronate, 10 mm long, the posterior ones alate. PETALS imper-
fectly known, slightly exceeding the sepals. Pl. I, fig. 21:
Apex of inflorescence x 1/2; fig. 22: Flower x l.

Minas Gerais- Espirito Santo: In colonies in fields

Duarte s. n. (RB 94376). Same, cultivated, 7 November eid AY
B Duarte 3909 (RB, form with longer narrower leaves, possibly
due to cultivation

11. 0. LEPROSUM (Mez) Mez in DC. Mon. Phan. 9: 117. 1896.
prantiela sggproes Mez: in Mart. Fl. Bras. .3;* pt. 32 259, pl, 58;
fig.

eases fice locality, Glaziou 14035 (K, US phot. 4184).
Minas Gerais: Cachoeira do Inferno, Rio Jequetinhonha, (Septem-
ber 1820), Pohl 5229 (W, type, formerly cited as from Goids).

ss)

0. GLABRUM (Mez) Mez in DC. Mon. Phan. 9: 117. 1896.
ppantteta glabra Mez in Mart. Fl. Bras. 3, pt. 3: 258, pl. 58,
fig. 1. 1891; Pflanzenreich IV. 32: 72, fig; 34.

IL: Minas Gerais: S%o Miguel, Rio Jequetinhonha, (August
1820), Pohl 3436 (BR, type; GH phot. 2792

13. 0. MARACASENSE L. B. Smit h, Brom. Brasil in Smithsonian

Mise. <Coll, 126: 33¢ 180. fig. 842 1955.

ZIL: Bahia: Table Rock area near Ma aracas, alt. 900 m, 21
October 1948, Foster 2471 (US, type; US phot. 4245
TUM L. B. Smith, Bromel. Soc. Bull. 12: 52; 4198.

salice
L: Espirito Santo: Near Santa Teresa, alt. 900 m, 7
reg ree Foster 846 (US, type).
15. 0. DISJUNCTUM L. B. Sm ith, Brom. Brazil in Smithsonian
Misc. Coll; 126:°33, 160; fie. 85.. 49
15a. Var. DISJUNCTUM. Flowering 35-50 em high. Sepals 11 mm

BRAZIL: Paraiba: On rocks at Gusianda, between Campina Grande
and nisin J Gn Pernambuco), alt. 450 m, 11 October 1948, Foster
2419 (US, type). Pernambuco: Mun. Quipapa: Engenheiro Pelada, ada, 12

RB, US).
15b. Var. MINOR L. B. we Phytologia 7: 255. 1960. Flower-
ing 15 cm nt i a Sepals 9 mm long.
Without locality, Instituto Biologico da Bahia 1083
(RB 976079, Pernambuco: Mun. Quipapa: On granatic outcrop of
mountain, Uzina Agua Branca, Fazenda Pelada 12 July 1950, D. A.
sama 50-592 (IPA, type; US a phoks 5700

L6h, PHYTOLOGIA Vol. 13, no. 7

o O. MAGALHAESII L. - Smith, sp. nov.
disjuncto L. B. Smith, cui affinis, foliis lepidibus pa-
tentibus profunde apelin utrinque vestitis, sepalis majoribus
diffe
ee flowering over 55 cm high, its base unknown, LEAVES (or
basal scape-bracts) 50 cm long, covered throughout with a dense

trorse spines. SCAPE erect, 5 mm in diameter, densely tomentose-
lepidote; scape-bracts reduced sae but little changed in

shape. INFLORESCENCE laxly compound, tomentose-lepidote, becom-
ing glabrous with age; primary bracts subfoliaceous, all but the

t
PETALS imperfectly known. Pl. Se fig. 23: Section of inflores-
x

: Minas Gerais: Mun. It caedae uri: Ouro Verde, 9 ow 1958,

Mendes Magalhacs 19187 (Hb. Bradeanum 17950, type). Mun. Ata-
eia: In co on rock outcrop, Ouro Verde, 8 Satenie® viet

Mendes ey eed 12035 (RB 111107).

Plate I

Fig. 1: Tillandsia setacea (Small & & Mosier 5506), leaf-section
x 25. 2: T. bartramii CeConte) , leaf-section x 7:
Greigia rohwederi. 8: Pitcairn
Buya floccosa var. ae 20 52%
Aechmea bahiana 5

amoenum. 19: O. saxicola. 20: 0. compactum. 21, 22: 0.
duartei. 23, 24: 0. magalhaesii.

1966

Smith, Notes on Bromeliaceae

ADDITIONAL MATERIALS TOWARD A MONOGRAPH OF THE GENUS
CALLICARPA. II

Harold N. Moldenke

CALLICARPA L.
Additional & ee bibliography: Decne., Nouv. Ann, Mus.

1
Maatsch. sere t Sgemaos (Ann. Hort. a lid se 3 185:
7-72, <5 & 6. 18453 Pate Prol. Fl. Jap Pe
Arb. & esis 179. 1877; D. Thompson, The EE: eer Si 518-—-
519. 1880; Franch., ude.’ aa Sci. Nat. Cherbourg 2: 240. 1882;
3

19; Nakai 2
1: 450-64, fig. 213--219. 1927; S. Sasaki, List Pl. Formos. 350.
“ibd Van elle, Journ. N. Y. Bot. Gard. 43: 36 & h2. 1942; T. Ae
t. Surv. Ind. 1: 11h. 1959; Nath, Bot. Surv. So. Shan

s lsh. "19805 Legris, Trav. Sect. ot nst. ¢- Pond.

a 18h, og 39h, 501, 513, & 558. 1963; - Hawkes, Guide Pl.

Everglades Park {3, 45—-h7, & 9. 1565; ; Woldenke, Phytologia
Phot $65 Aaa: Gen, Costa Ric.

Four=-
, Imp. ep Fam. Costa Ric. 13. 1966; Moldenke, Soak Sail
ang 10, 1

The L., Mant. Pl, Alt. 198 (1771) reference given in the bib-

erroneous "1767" for it. The "Journ. Roy. Asi
Son Bengal B M3) “082 & 1018. 1909" reference sometimes cited
is an error for King pd rom Mat, Ri Malay. —_ (Ql: 1012—-
1018 (1909), which is a reprint from Journ. Roy. Asiats Soc.
Bengal 7 (2), extra no., 802—808 (1908). The Decal ne (1835)
reference in the bibliography above is sometimes

uae’ as, for instance, by Lam (1919), or oleed as "5:

CALLICARPA ACUMINATA H.B.K,
Addi

tional bibliography: Résumé Suppl. 1h: 5 & 7.
19663; Moldenke, Phytologia 13: 3t kot 1966. eg

Se ne
Additional & emended b ography: Bakh., Bull. Jard.
— ser. 3, 3: 23. 192; Moldenke, Phytologia 13: 17

Bakhuizen van den Brink wei? places this species in the sym

1966 Moldenke, Monograph of Callicarpa 467
onyuy of what he calls C. cuspidata Roxb. [=C. peduncuilata R.Br.].

CALLICARPA ACUTIFOLIA Chang
Additional bibliography: Moldenke, Résumé Suppl. 1h: 3. 19663
Moldenke, Phytologia 13: 37-38. 1966.

CALLICARPA AMERI

Additional & emended
(1862) and 3: 3/2, Eg Bocas Rev. Verbenac. 129 & 192. x82 635
Gerth van Wijk, Dict. oer at 2a (1911) and 2: 132 & 512.
1916; W. H. Harvey, Univ. Ga. Agric. Ext. Serv. Bull. 02, ed. ea
20 & 31. 19h]; F. E. Selon Check List Ferns & Flow. Pl. Seas
State Park 50. 1942; Moldenke, Known Geo - Distrib. fouuse
2h, 71, 80, & 86. 192; pac aap Alph.
List Invalid Nanes’7, 8, 11, 2h, fa 192; Moldenke in’ Lundell,

Texas 3 (1): 7780. 1942; Van Dersal,

0, 259, & 260. 192; peda sap ‘nook, Shrubs, ed. h, 168. i9h23 ae
self, Sanders Encycl. + Pts Ths i9h2; R. nll
Ecol. Monog. 13: 130 136, é 169. “13055 ze ’G. Yuncker in
Fl. Texas 3: 139. 1943; J. J. Drayton, Carol. Flor. 15- 19135 5 EL al
Davis, Fla. Geol. Surv. Bull. 25: 100 & aa 1943; Moldenke,
Known Geogr. Distrib. Verbenac. Age 1: 1943; Betts, Jeffer-
& 672. iy "P. Cy Hoehne, Relat.

artin, Am. Midl. Nat. 3%: 608-609, pl. 50. 9h eee
Alph. List Cit. 1: 3, 4h, 6--l1, 13-15, 17—20, 22, 23, 25,

29, 30, 32--3h, 37, ae 41, 43, 45-7, 51, 52, at 59 63, a.
71, 76 ; 9h, 97—101, 104, 107, 112—~

118, 120, 123-125, ey 137—1l, 143, ibs, 153, 157—159,
162, 166, 169, 175-1 77, 181, 183, 186, 190, aoe 193, 195, 197,
i i ie 1, ’ 23h, 235,

99 203 207, 209:
239; a 2h2, 2i5~-29, 253, 256—-262, 265, 367-2 77, 280, 21,

Bol.
Soc. Venez. Cienc. Nat. 11: 39, ki, h2, & 51. sg K. W. Hunt,
Am. Midl, Nat. 37: 689 & 77. 1917; Tatnall, Fl. Del. 218. 1917;
hytologia 2: 326, 327, & 3h6 (1947) and 2: 9. 1948;
1948; M anea 13: 121

; ard.
a : : 3,
343, 35-356, 390, 391, 393-298 Spt—tot, 103, 405, 40709,
452; 45h. | 45S, 458, \s9, 69. ee 472, 7 tae » bE, 481, 482,
48); 4,87 , 489—h91, » 495, 99, 503, 50h, 507—509, 5.
518-522 ; 52h, 96598 530, 5%, 543-515, zug—80" 555, 556,

1,68 PRY.TOLOGLIA Vol. 13, no. 7

558--560, 562, 56h, 566--568, 570, 572, 57h, 576, 577, 579, 581,
583--585, 588, 592, 598, 601, 60h, 606, 609, 61k, 615, 618, 621,
626, 640, 643, 6b, 646, 649, & 651 (198)
663-66 » 678-680, 68

47, 749-752, 754, 756, 759761, 763, 767, 769-771, 773--776,

778, 2 5, 797-802, 806, 816, 822, 825, 827,

828, 831, 832, 835, 8h1, 842, 8k, 847, 848, 850--853, 855, 857,

859, 868, , 878, 880, 882, 8 86, 888, 891, 89h, 895,
9

002—-1005, 1012, 1014, 1016—-1018, 1020, 1022-1025
1032, 103h—1042, 1046, 1052, 1054, 1058, 1060, 1062, 106h, 1066,
1070, 1071, 108); 1086, 1 a

220,
123k, 1238, 122, 12h6, 128” 1252, 1255, 3283, 1291, 1292, 1296,
12° & 289-19 191,95 Moldenke, Phytologia 3: 72°& 131. 19495 Mol-

°B
1: 280. 1950; H. N. & A. L. Moldenke, inet Inst. Biol. Mex. 20:
a, ea Testovit, Pl. Disease Handb. 451. 1950; G. T. Moore,

1

3 » Phyto a
- & 304 (1350) and 3: Lh9—51. 1951; Keever, *Oosting, & Ander-
m, Bull. Torrey Bot. Club 78: 1,09. 1951; W. J. Bean in Chitten-

8:
New Britton & Br. Ill. Fl., pr. 1, 3: 137 & ie 19523 or
Phytologia h: 88. 1952; Ponce de Leon, di v. Soc. Cub. Bot. 10: 23.
1953; J. A. Steyerm., Rhodora 55: 238--2h1. 1953; rad y Std
Dice. Bot. 2: eid & goer 1953; Thorne, Am, Midl. Nat. 52

2
& Br. Tl. Fl., — :. 5 "137 & gi ae? Anon., Bi-
ol. Abstr. 30: 3826. 1958; Krtissmann, wt 1? 25h.
1959; Moldenke, Résumé 8-12, 1h, 18, 21--23 ee. or 3h, h9, 50,
53, 213, 23h, 2u0, 2h1, 2u7, 248, 29,, 301, hohe bh3. 1959 9} Mol-
denke, Résumé Supp 293

Fiamma &: Kes 28’ ( - Yonog. & 5 h
Suppl. 3: 2-6 & 28 (1962), hz 1, 2, & 10 (1962), 5: 1, 2, &
(1962), and 7: 1 & 2. 19633 Shelford, Ecol. N. Am. 67, 77, 82, &

1966 Moldenke, Monograph of Callicarpa 69

582, fig. 3A. 1963; Li, Morris Arb. > ant tal, 3a & Ms fag

ba? 1963; Gleason & Cron nquist, Man Ud Pl. 582. 19633 J. A.

Steyerm., Fl. Mo. 1262-126 & "1267, map 189, pl. 303, fig. i

1963; Lakela, Sida 1: 6. 196k; H. ae Hoffm., Castanea 29: 31 & 0.
196k; Re le Taylor » Plants Colo on. D ’

keer Résumé Suppl. 8: 1 (196k), 10: 2 (19645, and 12: 1. 1965;

In Gard. Hawaii, ed. 2, 726

cud de Pl. Everglades Nat. Park 43, 45--l\7, &
Morris Arb. Bull. 16: 52. 1965; Moldenke » Phytologia 13: 328,

gre ete 33, & 439—bhh. 1966; Moldenke, Résumé Suppl. 14: 1 &

59. 19

Dlustrations: Pluk., -irkag whe Bot. pl. 136, fig. 3. 16%;
Duhan., & Arbust. 1: 112%, pl. hh. 555 Kniphof, Bot.
Orig. Herb. Viv. 1: ai" 196 ie color} (1758) and ; oF.
1758; » Abhandl. grr: Page & Strttuch. 1: ae 1763; Buch-
oz, hast Nat. Rated Vég. 6 » Ple et 1775; Buchoz, Herb.

Tabl.
Méth. [(I11 eee Gen.] 1: pl. 69, fig. 1. Ahooas Gaertn., Fruct. &
H

. 6 : 1, fi i
Ornament. Shrubs U. 8. 289, 290, fig. 503 & 50h. 19105 Co Ke
Schneid., Ill. Handb. Laubholzk. 2: 592, fig. 38h f—g. 19
Hickel, Bull. Soc. Dendr. France 28: 16, . ie. "15133 Britton

Am. 67, fig. 3A. 1963; Li, Morris Arb. Bull. 14: [1] &h fig.
1-1, 1963; Je A. Steyern iy Bik Mo. eS 303, fig. kes 1963; R Ss
1

round c ve imple
or with a few long branches, sometimes trailing over other vegeta-
tion, the branches whip-like, the leaves soft-green, long-petio~
late, usually very coarsely and more gee less irregularly toothed

teeth, usually
stellate-tomentose or densely rusty-downy beneath, the petioles
1—-3 cm. long or about 1/3 to 1/5 as long as the leaf-blade
peduncles 310 mm. long, as long as or shorter than the petiole;
calyx-rim subtruncate, i Sorte or very shortly triangulate-

toothed; corolla from bluish or lavender to pale pinkish-
lavender, lavender-pink, pink, pinkish, light~pink, pale~ .
rer ient te, or even pegten anthers yellow; fruit

470 rere OLOGiIa Vol. 13, no. 7

from bright-blue or pink to magenta, violet, shining-violet, red-
violet, reddish-violet, rose-violet, purple, dark red-purple,
cerise-purple, or brilliant wine-purple when mature.
flowers are described as "pink" on G. J. Goodman 567h,

Lundell & Lundell 9653, and G. T. Robbins 2648; "pinkish" on Lun-
dell & Lundell 7048 & 12939 and H. N. Moldenke 296; “light-pink"
on G. T. Robbins 2660, Thieret 15945, and Whitehouse 16158;
"pale-pink" on Thieret 1593h; "pale pinkish-lavender" on Lundell
& Lundell 9615; "pinkish-white" on Tharp, Turner, & Johnston
547135 "lavender" on C. L. Lundell 13930; "lavender-pink" on Shin-
ners 20235; "pink-purple” on W. H. Duncan 20155; and "white" on
G. He Mueller 8026. woes

The leaves are ternate on Cuthbert s.n, [June 13, 1902]. The
fruit is referred to as a "berry" by Cronquist, the Lundells, and
Whitehouse, but is actually drupaceous. On one sheet of O. E.
Jennings s.n. {Fort Meade, late Dec. 1916] in the Carnegie Museum
herbarium the fruit looks white when dry — this sheet 8
sibly represent var. lactea F. J. Muller. Hottes (192) calls

"one of c fruit, but it is more tend
than the Asiatic species. In its bitat, it is found in rich
woods and thickets from Florida to Texas and, Ten-
nessee, eceeethe er illustration is
taken from its natural habitat in Virginia. The plant fl

from to bears ripened frui yun tober." Santa-
mour (1965) found the chromo number 2n = 3%. He con-

some t
cludes, ver, that “an accurate appraisal of the cytological
situation in Callicarpa will result only from a critical cyto-
taxonomic study or a large mmber of species."

Collectors have found the species growing in loose or Carrizo
ere Susquehanna oam

swamp jungles, woods rich lowland woods, low open or sandy oak
woods, low moist or low moist open woods, shady or low moist flood-

woods, sandy pine-oak or low maple woods, low or dry sandy
woods, rich or rocky woodlands, dry woodlands, forests and
forests, maritime and pine forests » Swamp and pine-oak-hickory for-
ests, in open places in woods and somewhat marshy places in for-
ests, hammocks and high hammocks, oak hammocks and sandy marl
mocks, pastures and cow pastures, dense sandy woods and live oak
woods. It has been found in pinelands, barren pine woods, decidu-

1966 Moldenke, Monograph of Callicarpa 471

ous forests, thickets, roadside or woodland thickets, sandy pine-
lands, pine-oak-hickory woo woods, alluvial woods, swamps and live
oak forests, brackish marshes, valleys, salt marshes,

creek bottoms, bottomland pastures, roadside ditches, rich river-

exposure,

Collectors have encountered it along country roads and fence-
rows, along ricers and small streams, and on stream banks, at the
edge of low or moist woods, at the edge of swamps and creeks, in
the borders of thick woods, at the edge of swampy or of oak=p

and

woods r wood] Ss, at the base of bluffs, close
to creeks, among te boulders, at the e of =
ets, in pond or clay pond margins, at the edges of granite ou
c * oak on mountain slopes, in

rders, at the edges of lakes and of upland woods, at the edge of
deciduous of cypress swamps, as well as

river bluffs, stream and creek banks, savannas and pine savannas

low or dry hillsides, mountainsides, wooded slopes and wet slopes,

and on granite outcrops. It seems to thrive in gravelly clay pine
bo ks

the wooded slopes of streams, open dry oak and wet Limestone

oak-hickory=pine woods, ~hickory Ss

ds, bo ood=pine forests, openi

dlands, cutover pine and awe depressions,
and low eral. It has been seen in woodlands on sand
and woodlands on upper river terraces, in mountain ravines, barren

ew 7 © rests, pin » ravines in elands, the

and in thickets at the edge of deciduous woods
It has been collected in anthesis from April to

December and
fruit in every month of the year, growing at altitudes of ag to
1500 rail Plée we a a4 ccancans en a eagsnte et dans la Penn-
sylvanie"; Demaree + “very common" in ety County,
Aniansas; Thorne 2 (2951) grdet by it as "abun ", Ewan says

ry abundant h ak=-pine wae dy Riley refers
or it as rare ‘woods transitional between pine and deciduous

forest" in St. Tammany Parish, Louisiana, while Totten, Raynal,
Hoffman, & party call it tyvery abundant" in Alexander County,
North oF cont ta found it "frequent in oak forests" in
Marshall Coun klahoma

Lakela (19613 pare that © allicarpa americana is "occasional"
in Pineola Grotto, Citrus County, Florida — the "plants rooted in

inata & Q. myrtifolia, Carya floridana, Fraxims, & Pinus cli clausa"

472 Peet OL OG1 A Vol. 13, no. 7

in Hillsborough County, and Godfrey states that it is "abundant

in thickets at edge of deciduous woods" in Leon County of the

same state. In lundell (192) its Texas distribution is given as
follows: "In woods and moist thickets, wet slopes, low rich bottom-

L. C. Taylor notes that it is "a favorite food of cardinals :

Bermuda. It is widely cultivated, although not as much so as the

Asiatic species, C. dichotoma (Lour.) K. Koch and C. japonica

Thunb, Tatnall (1947) records it from Wicomico County, Maryland.

Harper (1928) records it from Butler, Chilton, Coffee, Colbert,

Covington, Fr » Greene, 8, ©, Monroe, Morgan, Pick-
es, Alabama, "and

f places fire is
rare, but fire is not necessarily fatal to it. The slope of a
ravine on the University campus where Callic is abundant was

ot. ear Cc.
the competition of other plants which are less sensitive."

1945) describes our plant as common in the Gainesville
area of Florida, the corolla bluish or pink, and the fruit violet
or magenta. J.H. Davis (1943) describes it as a characteristic
plant of the Parkwood loamy sand soil of southern Florida. Matsu-

1966 Moldenke, Monograph of Callicarpa 473
mura (1912) eerie it as cultivated in Japan. My son and I have

seen it c in the Foster Gardens at Honolulu, Hawaii,
in 1961. McVaugh h (193) records it from the shrub layer where
the shad wi copallimm. He also gives

rocks" along with Forestiera ligustrina, Gelsemium s
Quercus $8 georgiana, and Vitis rotundifolia, He describes it as a“
“common shrub throughout thi: this region, and apparently quite toler-
ant of soil acidity, as it is one of the chief shrubby invaders
of vegetation mats on the flat rocks, where it is associated with
Rhus c mum, and, in the mature stages of the association,
with Juniperus virginiana, Vaccinium Vaccinium arboreum, and Pimms taeda.
The violet—purple fruit res s abundantly produce: ebaded and very very ornamental,
and the plant is richly deserving of cultivation on its account."
Van Dersal (1942) comments that Callicarpa americana and Hy-
ea ifolia "which do not occur naturally in th in the northern
part of region 25 [of Mumford's Plant Growth Regions], have long
been enn grown there. Neither of them, however, can be
gromm in region 27." He describes the beautyberry as follows:
"Found in the Southeast is a lovely purple-fruited ei known
locally as French miberry. The plant is no mulberry and it cer-
tainly name is American

June to August. The fruits are small and berry-like, shining
violet or red-violet in color, and are carried in clusters along
the whip-like branches much like beads on a string. The whole
tone of the shrub after the leaves have fallen is a sage or

poor ° s
to Texas and Florida, in plant growth regions 17, 20, 25, 28, 29,
and 30. It is not hardy in Massaclmsetts or in England except in
the milder parts of that country. Sand, clay, loam, and gravel

es. The s is v tt to b , 12 species of which

are known to feed upon the fruite; ied inelude cardinals, thrushes,
» mockingb and towhees

Steyermark (1953) gives a very interesting account of the al
ical history of this plant in iadecss "The French Mulberry o
Beauty=berry......, one of the most beautiful of native pee pk
shrubs f in a wild sta Missouri. E. J.
Palmer, Bush, Letterman, Eggert, and Kellogg were well acquainted
with if they collected it from various lo-
calities, many as-Missouri border, but, al~

L7h PHYTOLOGIA Vol. 13, mo. 7

proved to be based upon misidentifications., In Dr. Moldenke's
Monograph.......the Missouri records are likewise shown to be
based upon unauthenticated reports.

Mat the New York B hiedoak Ol Garden in the spring of 199 I con-
ferred with Dr. H. N. Moldenke about the distribution of the Ver-
benaceae in Missouri, and was surprised to learn that he had ex-
amined a specimen of ee americana supposedly collected in

County
of terrain for Callicarpa. If it occurred anywhere in the state,
it should have been found in extreme southern Missouri in an un-
glaciated section,

nT same spring, I wrote Mrs. Young and expressed my inter-
est in learning the exact station where she had collected the

am normally a very careful worker and I hada large collectian of
Chariton Co., Mo., also of Logan Co., Ark. In working these over
—- with both places in my mind — I simply wrote the one state
when it nee have been the other. I take great pride in seldom
% this was a bad one. So sorryi' As of the
to

"It was while botanizing on September "29 1949, along a stretch

of White River in Taney County by the Missouri-Arkans as line, just

within Missouri territory, that I made a real discovery. From a
Ri known

place on the White River as Brown's Ferry
north and east C Hol along as rome feo
stone, wooded slopes that faced to the south into That
n not found hing especially exciting; then $
denly I spotted some clusters of small ts

of an

of purple (between a doge purple [color 732, p. 96, vol. 1. 1938]

and beetroot purple ae sn we ee nd 2. 1939] according

to the colour chart of the rticultural Society). Nearing
and examining then, se vubieetans at once that they were

indeed those pf the ae hunted Callicarpa. Not one bush, but
encountered in

searc
erates. that = high rocky slopes to the f limestone
8 hest point reached b the water
1 ae gus Steen po: eing > or feet dis e
"I collected many i marl specimens for distribution to vari-
herbaria and was realize that here was a new no
western limit of Batcinetias for this shrub, At the same time,

1966 Moldenke, Monograph of Callicarpa 75

however, my pleasure in seeing this remarkably beautiful shrub
was dispelled by the thought that it would not be long before
pees plants and the species, at the only known station in the

become exterminated. For, although the altitudinal
distribution of Callicarpa was found as high as sixty feet above
the pec water level, the future impoundment of the Bull

would not only ¢ completely cover this sixty feet, but

fifty Sax sixty additional vertical feet. This would bring the
level of the waters well up on the high south-facing
limestone bluffs that towered far above the last upper stand of
Calli bushes.

"On May 5, 1951, I re-visited this spot and found the plants
cult

faded in
The same author (965) says a only — known in Mis-

SOULS Ci acece en exterminated the impounded waters of
Table Rock ae isa nabs Mees the seopppedaeds 3 of
Illinois Herbarium appears a specimen of this labelled '

albidus, Jackson County, May 29, 188', collected by Bush. The
label i i from a Mo. State University sheet (no. 17619) rene obvious-
ly is a mixed label.

It should be recorded here that Smith & Stewart S.103 and Rad-
ford & Stewart S.103, cited below from rom cultivation, were grown
from seeds obtained in Cumberland County, North Carolina, while
Radford & Stewart S.56 is from seed collected in Alexander County
of the same state

The specific name of our plant is very often upper-c ased,
practice now frowned upon by followers of the present ntaee of
the International Rules of Sohasraet Nomenclature.

Under Genus 136, Callic » in the Linnean ee in London,

nyms and are so listed by me in the present work.
It should be noted that the C. americana of Blanco, mentioned in

476 PHYTOLOGIA Vol. 13, no. 7

in my 1936 work, unless one does not wish to recognize the valid-
ity of C. ei as a distinct taxon].

Sprengel (1825) and Schultes (1827) place Callic ‘lobi-
flora Rufz & Pav. in the synonymy of C. americana L., But it actu-
ally is a synonym of Aegiphila integrifolia (Jacq.) Jacks. Be-
cause of this faulty synonymy, Sprengel gives the distribution of
C. americana as "Carolina. Virginia. Amer. austr." Martin (1916)
discusses the endosperm morphol

The sheet of W. C. Coker s.n. [Arboretum, October 21, 19h)
has a label inscribed "White-berried form and purple-berried form
from the same root", but the specimen comprises just two tips of
branches, the ae ip on both of which have dried exactly the
same as is seen in the normal purple-fruited

m and ve ar names recorded %

can beautyberry", "American c carpa", “americanische Wirbel-

’ he Schinbeere", "ame: irbelbeere",
“beauty berry", "beauty-berry", "beautyberry" » "beauty-fruit",
"B "Be. mulberry”, "bird-eye", “bunchberry§
"burcardie d'Amerique", "callicarpa améri ®, "cakii da!
Amérique" shrubby callicarpa", "commode mul
"common American beautyberry", "filigrana", "f e masorcay
"filigr » "filigrana fructo morado", "filigrana fruto
morado", *f morada", "foxberry", "French berry", "French-
lace", "Frenc "8 h-mlberry", "inkberry", "Mexican-
mulberry", "purple beauty-berry", "sour=bush", "

"Spani
spice". It should be noted, however, that the name "bunchberry"
is also applied more commonly to Chamaepericlymenun canadense (L.)
Asch. & Graebn. in the Cornaceae, "foxberry" to Mitchella repens
L. in the Rubiaceae and Uva-ursi Uva-urei procumbens Moench in the Vaccini-
aceae, and PROP ertsn to Llex gl Tiex glabra (L.) A. Gray in the Diicacese
—e americana L, in the Phytolaccaceae,
Te ahey Peninia us that the common name "mulberry"

applied to this and related species because the fruits

of C, americana, type species of the genus, are borne so close ee

gether in the fruiting cyme as to appear like one compound frui
Seymour (1929) records often following fungi as attacking Galli-
arr americana: Atractilina calli Dearness & Barthol.,
ryosphaeria callic Cke., Cercospora callicarpae Cke., C-
pulvinulata Sacc. & Wint., Coniothyrium callic Cke., Diatrype
aaars B. & Rav., Meliola amphitricha Rav., M. cookeana Sacc.

& & Spee, - inermis Kalchbr. & Cke., and Melogramma callicarpae
Cke. Westcott (1950) says that it is attacked by black mildew
Meliola cookeana) in Florida and Louisiana and by leafspot (Cer-
sospers callicarpee) from South Carolina to Texas. Brooks (195i)

that this species is one of the hosts for Radopholus sim-
tis Florida, Coker reports "Infested with soft scale insects
belonging to a group closely related to that of cochineal. It is

1966 Moldenke, Monograph of Callicarpa 477

large, measuring when fresh up to 1 cm. wide and shrinking very

little ris drying. It has a soft spongy covering, almost white as

chalk, The insect itself is full of a deep red thickish liquid,
etcalf of know

as in the cochineal. Dr. M of State College did not
t us or ies. The ins fairly common o
Mulberry here [Bull's Island, South Carolina] oc less

carpa americana is parasitized by Cuscuta coryli Engelm. in Texas.
According to Rehder ae) Cataonepe americana was introduced
into cultivation in 170, but actually it was first sent to Eng-
land for ae in bok [see Bean (195i). It was growm
both by J omas Jeffers
gardens. sain (1935) avers oe: teat it is retbice to the harticul-
e now Howell

e Sarcoxie Nurseries in Sarcoxie, Missouri. Mattoon
(1988) iavte eleven horticultural ac pores
he physico-chemical properties of the sap in relation to phy-
togeography are discussed by J. A. aces (193k). He based his
work on specimens collected in the Royal State Park, Florida, as
well rub-covered

poset reer ye isa ear otk, Fe Tp gpa barca compa ris
munity of pine and hardwoods in a deep hammock near Pablo,
Johns County, Florida. He also used material from a wooded bluff
satanealay ser shate 0h 8b ane a ANS Ses See © above St.
ary's, Georgia, in low flat sandy oak woods 1 li gang goa
Fort Oglethorpe, in flat pine-cabbage palmetto woods at the head
of navigation on Turtle River, and in moist almost ea geste
at Cabin Bluff Wharf, at the head of the Cumberland River in the
rth Caro

swamps at South Mills, Camden County, swampy depressions and
wooded co ei a Swansboro, and the wooded aang side of
sand dune ck Counties ye a
" ieavton G93). report that it "Blossoms in Julys grows
eful in dropsical complaints." Rosenthal
ter sind e

Wassersucht gebrittuchlich, die Frtichte rothe F. " Itis
also reported to be useful as a fish-poison. L. A. eggs (19h0)

Amerinds use it against dysentery, colic, and dizziness. "The
roots are boiled with those of Rubus sp. The patient dicks the
decoction to cure dysentery....A “A decoction made from roots
berries is drunk Angered colic,.....The roots are boiled and the
decoction is drunk during the attacks of eral The Koasati

She makes the comment that "The sweating of the patient during .
fever attack would be beneficial regardless of the plants used.

478 PHYTOLOGIA Vol. 13, no. 7

notation "drawn for book." Murray (1774, 1784) cites the Linnean
Mantissa synonym as page "126", but it was actually published on
page

The Buckley s.n. (July, 1820] specimen cited on page 313 of my
1936 work is actually from Wilcox County, Alabama, the G. M. Mer-
rill 21 is from Pope County, Arkansas, and the Gattinger s.n. 6 othe
Teast o: of Cleveland] is from Bradley Count: y, Termessee —- the coun-
be names in all these cases having been omitted by the printer

when making corrections in the page-proof. The Andrews specimen

from "Lyon's View", cited Ay the same page, is from Knox County,
Tennessee. On page 314 the Edwards collection cited from Fort
Towson, "Arkansas", is actually from Fort Towson in Choctaw County,
Oklahoma. Likewise, the Trelease collection from "Wagner Cove" is
in reality from Magnet Cove, Hot Spring County, Arkansas.

perdi 1067 is labeled "Comanche Spring: New Braunfels,
pies o", sO may actually have been gathered in Comal, Comanche, or

ome other county of Texas. C, C. Perry s.n. [En route from San
ints si P see to San Antonio, — — 1878] may have been col-
ected in Mexico or several othe counties rather
sae County as here cited, The “abel of M. Morgan 91 is inscribed
ee

Ace cording » the M
ne [Rebel Retreat, June 1879] oan “have been collected i "ts iedell
or McDowell. Counties, North Carolina. The Collector undesignated
s.n. [Herb. Iowa State Coll. 108282] is labeled "probably Wichita,
Kans.", but this is doubtless an error because the species is not
from that state -- its labels appear ve
one in longhand on the same sheet is inscribed "C Caprifoliaceae 457
B) ric s s*,

The Avery, Jefferson, and Weeks Islands, cited below, are in I-
beria Parish, Louisiana; Key bare) is in Monroe County, Florida;
and C Island ri in Dare County and Smith Island in Bruns-
wick County, North Carolina,

The C. Hartmann s

- (Huntsville, Summer 1912] specimen in the
University of Cali arets herbarium has the collector's surname mis-
spelled "Hartman",

Material of this species has been misidentified and distributed

in herbaria under the names C. cana Spreng., C. JUSS +,

and Maclura aurantiaca Nutt. On the other hand, the Rob. Burton

s.n. [October 19h6], Radford 1,849, and W. H. Rhoades s.n. [North
y actually

1966 Moldenke, Monograph of Callicarpa 79

n. from Arkansas is Symphoricarpos orbiculatus Moench in the C Cap-
Tifoliaceas; and Pammel & Pammel s.n. [Miami, Jan. 15, 1928] is
Trema floridana Britton in the U Ulmaceae e

~~ Hooker (1836) cites T. Drumond ond 25lbis & sen. [1833] from
ouisiana

In all, 2558 herbarium specimens and 59 mounted photographs and
illustrations = been examined by me, including type material of
most of the name ved.

Additional & pile se citations: MARYLAND: Wicomico Co.: W W. M.

Canby 72h (W--1323259). VIRGINIA: Elizabeth City Co.: T. W. Chick-
ering s.n. [Fortress Monroe] (F-14957); Churchill s.n- 1. [July 22,

Co.: Sargent & Smith s.n. [vicinity of Gloucester] (D—702382).
Isle of Wight Co.: A. R. | Moldenke 1,76 (Fg). James City Co.: R. W.
Menzel 71 (Mi); Sargent, Thayer, & Sm Smith s.n. [near Williamsburg]
(D—70203). Mathews Co.: Rothrock s.n. [Aug. 5, 1879] (E—
119321), sen. [Aug. 10, 1879] (F-32232). Nansomond Co.: Britton
Britton, & Vail s.n. [Suffolk] (F—39771); A. R. Moldenke ]77
(Fg). Norfolk Co.: Blanchard s.n. [Dewall's Point] (W—309653);
Britton, Britton, & Vail s.n. [July 2d, 1892] (Ca—-5062), Cm);
Curtiss s.n. [June 27, 7, 1872) (a1), son. (F—149781); E. Durand s.
ne (F——725399); J. A. Harris C.1933) (F--901598, H—-13009); Hul Hub-
richt B.2559 (Au); a: E. Hyams s.n. [near Sewall's Point, Nov. 7,
1890] (Dt); Kea 296 1296 (W—35601h) 1498 (W--356192); A. R. Mol-
denke 479 (Fg); N ma 3.0. (aly 10, 1897] (Hp); Rugel s.n. (near
Portsmouth] (Ed); A. B. Seymour 16 { 91718] (E--981763, Sw. Feat
358636, Lb—-16203). Northampton Co.: W. M. Canby s.n. (Rastvill

Oct. 1867] (Pa), s.n. [Cape Charles City, Aug. 189k] (2 (p1—22686),,"
8.n. [eastern shore of Md. & Va.] (F--315218), aes (Dt); Canby &

——— a

Pe Ook irginia Beach

& Long s.n. [Sep. 15, » ; 1907] (Up—Li6 7k) 5 Bright 1196 1196 “ms F. E.

er 10-269 (N); bie 9773 (Up); Hars er 8.n. n. (Virginia Beach]
pA. eer Elise = 107 Mapes ~ E--119297, feral

[Virginia masks a (Cm); re s.n. [Inlet] 8s.

[Virginia Beach] (W—5929)6). Poros asf Cos: a & — ong 6363
(N, Up); Tidestrom 68) -geeiprmags W—-693896). Surry Co.: A. R.
Moldenke 75 (Fg). War : Swinerton s.n. (Newport News] (Mu—

8580); A. R. Moldenke 470 (Fg). Bladen Co.: Ahles & Haesloop
29127 (Hi—105610); Biltmore Herb. 1720c (Al, E--119153, Fe, Io—
TL, Na—10852, S, Wu, W—33209)); F. E. Boynton s.n. [Feb.
1896] (E~119320); A. A. Heller 14041 (Du-—157515, E--932538, F-—

689834, Se-—85971). Brunswick Co.: Fox & Wells 501 (No—8577);
R. K. Godfrey 6381 (No—858h) ; Reed & Stites 3k (Dt), 10h (Dt);
B. W. Wells s.n. [7/3/19)0] (No—-8578). Camden Co.: D. S. Cor=
Fell 2071 (Bt—33196, H——-32131); J. A. Harris 17733 (W—90h925)5
A. R. Moldenke 1,81 (Fg). Carteret Co.: Byster 265 (Ur); Fox &
Whitford 1108 (No--11379); R. Gray s.n. [Shackleford Banks, 6/28/
38] (H—-50202); B. W. Wells s.n. [6/28/1930] (No—-8583). Chatham
Co.s Radford 6288 (Hi—59)13). Chowan Co.: Ahles & Duke 50922
(Hi--106785); D. S. Correll 2033 (H—32133); A. R. Moldenke 169
(Fg). Columbus Co. Ahles 475 (Ur); Blomquist & Correll 4899 (=
)3 Re K. Godfrey 6279 (Hi—-10563); P. 0. Schallert 64, in
part (Ur); Fisgand & Meaning 2689 (Po--21h782). Craven Co.: Rad-
ford 37377 (Hi--105609). Cumberland Co.: Radford & Stewart 8.n.
{oct.”8, 1939] (Hi—~13123). Currituck Co.: Bartley & Pontius 129
(N); F. Blanchard s.n. (Snowden, Nov. 9 190] Coe) Dare Co.: F.

ne (Au, Ca~-882,39, Du-369LL9), son. (7/1h/h1) ();
Radford, Haesloop, & Miller 7553 (Hi )- Duplin Co,: Aliles
35720 (Hi—105605). Durham Co.: Bl 4.703 (H-—-20hy1). Edge~

comb Co.: Bridgers s.n, [Tarboro, Oct. 1916] (Hi—S9414). Frank-
lin Co.: J. S. Holmes [Sept. 29, '1h] (Hi—59)05). Gaston
Cos: Oosting 1637 (H—L2785). Gates Co.: Ahles & Duke hi595 (Hi-
106786), 51468 (Hi—106787). Graham Co,: Radford 15910 (Hi—
92919). Greene Co.: A. R. Moldenke 65 (Fg); Radford 0343 (Hi—
104895). Halifax Co,: Bartram s.n. [Weldon] (D—536223); Re Ke
Godfrey 5230 (H—L70kk); H. R. Totten s.n. [Oct. 28, 193k] (Hi—
5911). Harnett Co.: Foust & Foust s.n. [6/29/38] (N)5 Radford
6278 (Hi--54202); Radford & Stewart 630 (Hi . Henderson
Co.: Small & Huger s.n. (Chimney Rock to Hendersonville] (F—-
401309). Hertford Co.: D. S. Correll 2298 (H—-32256); Woods &

1966 Moldenke, Monograph of Callicarpa 481

Moreland BWO.43 (Hi--3,131, No—13877). Hoke Co.: Bell 185-552
(Hi—33633). Hyde Co.: Radford 39034 (Hi—104896). Iredell Co.:
M. E. Hyams s.n, [Rebel Retreat, June 1879] (Ka). Johnston ae
Deans s.n, (Selma, Oct. 22, 1937] (Hi--59401); Fox, Whitford,
Campana 1227 (No-~12638); B. E. Smith s.n. [Sept. Us, 1932) | oe
59410). Jones Co.: Beaven 22 (32498) 5 Corbin & Wyatt 6-137
(Hi~-57633); J. S. Holmes s.n. (Maysville, July 12, 1919] (Hi--
590) ; McCarthy s $.N. ante River] (W— 787638) . ae Co.: S.
Stewart 818 (Hi—106788). Lenoir Co.: D. S. Correll 142) (E—
11150, H--32257); Radford 28990 tiers He Re j H. R, Totten s.n.
fAug. 8, 1931] (Hi—59)08). Martin Co.: R R. K. Godfrey 7028 (H—
50271, No—-8582); A. R. Moldenke 471 (Fg); Radford 3965 (Hi—
104422) , 41927 (Hi=-10]423). Montgomery Co.: Oosting 1858 (H—
49628); Radford 19366 (Hi—9292h). Nash Co.: Ahles & Leisner
21279 (Hi~9342h). New Hanover Co.: MacElwee s.n. [Wilmington]
(D—517h09); Macfarlane s.n. [June 16—-22, 189k] (Up—36437), 8.
n. (June 26, 1908] (Up—-39738); Mae taatiane & Davis s.n. cp: 18,
189h) (Up); Uhler 68 (W—-1h67250); Wells & Shunk Sen. [May lh

1937] (No—8579). Northampton Co.: Ahles & Haesloop 52388 (Him
106789). Onslow Co.: R. K. Godfrey 6396 (H--55308, Hi—10l93),
6455 (Hi—10327); J. A. Harris C.19570 (P1—8h073); C. E. Wood
6463 3 (Hi--51185). Orange Co.: T. Brown s.n. [Oct. 18, 1948) (Hi-
35042). Pamlico Co.: Davis & Davis 10510 (We); Godfrey & White
6801 (F1--123)3, No-~8581) ; Oosting 33212 33212 (H—~17907); Radford
5821 (Hi--10454)). Pasquotank Co.: Ahles & Duke 48078 (Hi—

8, Hi--104h19); D. S. Correll 2097 (H—-32132); A. R. Molden-
= 472. (Fg). Pender Co.: Ahles 36182 82 (Hi--104565) ; “i, E. Hyams
S.n, (Burgaw, July 1880] (Dt, Dt, Po—85770), s.n. [Bengaw] (E—-

Tipis6, W-~1,7596). Perquimans Co.: A. R. Moldenke 1,67 (Fg).

Polk Co,: E. C. Townsend Sen. [Chimney Rock, July , duly 16, 1897] (Pl—
86885). Richmond Co.: Radford 14482 (Hi—92926) Robeson Co.:
Ahles 37281 (Hi—10)543). Rowan Co.: Biltmore Herb. 1720e (B, By
Dr, F——806690, Na—10867, Vu, Vu, W—-332095); A. A. Heller 140
[June 28] (E—119298, F—313137), 140 75 by J (Ca——2509h, “Ca—
25095, E--119299, Ed, F—396189, Up-—17065, Up—17066); J. Be Hor-
ton 557 (Hi—93425); J. K. Small s.n. [Dunn's Mtn.] (F—179893).
Rutherford Co.: Biltmore , Herd. 17200 [July 22] (Go, Lu, S); C. H.
Hall s.n, (Shaking Bald ng Bald Mt.) (Vt); B Rugel s.n. [inter Asheville et
Rutherfordtown, Aug. 1842] (Ed, Go); E. C. Townsend oes [Chimney
Rock] (W—3h152h). Sampson Co.: Ahles & , Haesloop 29966 (Hi—
104542). Scotland Co.: Ahles & Haesloop 26699 (Hi—1OU5U1); R. K.
Godfrey 5066 Lpeineg . Stanly Co.: Ahles & Radford 11933 (Hi—
92927). Tyrrell Co.: D. S. Correll 1899 (H—-3213h, N). Wake Co.:
Blomquist & lucent "20Th (H—32362); Buell & Godfrey 1713 (H—

1,82 PRITOCLOGT «A Vol. 13, no. 7

Paris & Daniel s.n. [10/31/5h] (Hi—58139). Roanoke Island: Rad-
ford & Stewart 826 (Hi—-15067, Ur). Smith Island: Blomquist s.n.
(Sept. 3, '32] (Ob—68638)5 Fox & Wells 501 (Ca—7)3928)3 Je A.

M. G. McCarthy s.n. (July 1885] (W—1h7599), son. [Oriente Caroling
Sept., Julii 1885] (Hi--5906), s.n. [eastern N. C.] (Ed, F
349111, W--227156). SOUTH CAROLINA: Abbeville Co.: Radford 25956
(Hi—93388). Aiken Co.: Radford 543 (Hi—16750). Allendale Co.:

Cow: We H. Davis 9372 (E—~899622); H. R. Totten s.n. [6-29-1935]
(Hi—5916). Bamberg Co.: Ahles & Haesloop 30571 (Hi—105611).
Co.: A. R. Moldenke 2h (Fg). Beaufort Co.: Ahles & Bell

Barmre &
15551 (Hi—93398); C. P. Alexander 258 (W--910h55); C. R. Bell
4789 (Him-92921). Berkeley Co.: Ahles 35503 (Hi—105612); J. C.
Barber s.n. [Aug. 21, 1949] (N); Revenel 3712 (Ed). Calhoun Co.:
Ahles 35252 (Hi--105607); Ahles & Haesloop 30187 (Hi—105606) .
Charleston Co,: Harshberger s.n. [August 8, 1911] (Up—633k4)5 Ke
We Hunt 2690 (It), 2691 (Hi—2762l,); Stewart s.n. [near Charleston]
(F—115750). Cherokee Co.: Ahles 3,395 (Hi--105608). Chester-

field Co.: Radford 15726 (Hi~-92922). Clarendon Co.: Radford
2487 (Hi—93392). “Colleton Co.: C. R. Bell 3622 (St). Darling-
ton Co.: M. A. Curtis s.n, {Society Hill, June 1852] (Pa); Je Be
Norton sone [July 8, 1920] (Hi—59396); B. E. Smith 372 (Hi—
20487), 1304 [June 1940] (Hi—26486), 130; [Aug. 3, 190] (Hi—
26486), 1456 (Hi—2648h), s.n. [Hertsville, 10/20/31] (Hi--59392)«
Dillon Co.: Ahles 37160 (S); Ahles & Haesloop 27826 (Hi-~10h89h) 5
Ae Re Moldenke 151 (Fg). Dorchester Co.: Ahles & Haesloop 37809
Hi~121516). Edgefield Co.: Radford 2639) (Hi--93393). Fairfield
Cow Ce Re Bell 9452 (Hi—9339h); House 2h7h (W-—514145). Florence
Co.: C. R. Bell 13378 (Hi—121517); Wiegand & Manning 2690 (Po—
214783). Georgetown Co.: D. S. Correll 5292 (H—h2513); Godfrey &

Tryon 126 (Ca--956875, Gg—290707, H—58063, It, N). Greenville
Coot Oe Me Freeman 57521 (Hi—93416); Thomason 61 (Gg—310909)5

1966 Moldenke, Monograph of Callicarpa 483

Wherry s.n. [September 20, 1934] (Up). Greenwood Co.: W. He. Da
vis vis 9128 (E--898877) ; Radford 26737 (Hi—93415). Hampton Co.: | C.
es: Bell | 3913 (Hi-92920). Horry Cos: Coker & Totten s.n. ey
13, 132) (HL (Hi—-59399)5 A. R. Moldenke 156 (Fg). Jasper Co.: Ahles
& Bell 1566), (Hi--93h18) . Kershaw © Cow: | Britt 8h e-TeOo
Lancaster Co.: Ahles & Haesloop 2757 (Hi—1OMh25) ; Huntley 139
(H=-54983). Laurens Co.: C. Re Bell 10277 (Hi-—-93421); 0. ae
Freeman 561,96 (Hi—92923). Lee Coe: A. R. Moldenke 1,38 (Fg)3
Radford 27319 (Hi—93420). Lexington Co.: Hexamer Hexamer 36) (Us), sen.
(Lexington, 1 1, June 1855] (Br); Hexamer & Maier 8. 8.n. 's [June

'55] (Cm); E. A. McGregor 208 (Du—75709), 3.0. [vicinity of
Batesburg] (W--982620) ; ; Radford 27152 (Hi--93u19). Marion Co.:
C. Re Bell 13661 (Hi—-1a 1518) 5 A. R. Moldenke 459 (Fg). Marlboro
Co. | Radford 190 19030 (Hi—92925). ~ MeCormick Coe: : Radford A
(Hi—93)22), 39828 (Hi—10N42h). Newberry Co.: C. R. Bell 9191
(Hi—9323). Oconee Co.: A. P. Anderson 1455 (Io—17l37), sen.
(Walhalla, October 1897] (Io—21)09). Orangeburg Co.: Ahles & &
Leisner 31515 (Hi~-104421); A. R. Moldenke 1,60 Oe: Pickens
Co.: Bannister & Anderson 575 (Hi--104420). R: Co.: C

Bell 1600 (Hi—18005)); Crawford s.n. [Columbia] Lae
Seeman sen. (Columbia, Oct » 1932] (H—5h21); 3 Ke A Taylor 91
(Ur), Sn. en. (Columbia, June 192) (Dt). Saluda Co.: Radford 26853
(Hi-—-93)27). Sampson Co.: A. R. Moldenke 6) ret Sumter Co.:
Radford 27522 (Hi--93426). Union Co.: 0. Mi. | Freeman 56667 (Hi—
92928). Williamsburg Co.: Radford 2),6h) (Hi--93396). York Co.:
Ahles 365 (Hi-—10539); Ahles & Haesloop 31128 (Hi—~10h538).
Bull's Island: W. C. Coker s.n, Tchristmas 1945] (Hi—28149). Isle
of Palms: T. G. "Harbison 12 (Hi--8)18). Pawley's Island: Radford
24915 (Hi—93L17). St. Helena Island: Cuthbert s.n. [June 13,
1902] (Fl—20934). Sull Island: W. Palmer s.n. (W—513617). With-
erspoon Island: B. E. Smith 145 (Hi—26)85). County undetermined:
Curtiss s.n. [S. C., 1852] (Us); J. A. Harris C.19701 (W—1135755).
GEORGIA: Bacon Co.: A. Re Moldenke 343 (Fg). Baldwin Co.: H. Ne
Moldenke 23037 (Ac). Bibb Co.: Mattams s.n. [Macon] (Pa); Re Re
Moldenke 368 (Fg); W. H. Rhoades s.n. sen. [near Macon] (Hs). Bleckley
Cost &: | A. R. . Moldenke 375 (Fg) Brooks Co.: A. R. Moldenke 311 (Fg).
Burke Co.: A. R. Moldenke 08 (Fg). Charlton Co.: : Strausbaugh & &
Core s.n. [June 5, 195k] (We). Chatham Co.: Crawford s.n. [Thun-
derbolt] (D—602h),3); Miller & Maguire 1252 (Ta—16568) ; Oemler s.
n. [1837] (Lu). Chattooga Coe: Freeman & ’& Freeman s.n. [Blowing
Spring near Chels sea, June 9, 1921] 1) (Ar—-19782). Clarke Co.: J. He
Miller 106 (F—-556659), E.103) (Up). Clayton Co.: E. B. Travis s.n.
TRiverdale, 10-8-30] (Ga). Colquitt Co.: Kirkpatrick s.n. [Doerun
8-130] (Ga). Columbia Co.: Pyron & MeVaugh 10h3 (Up). Dade Coe:

48h PreITOLOGIA Vol. 13, no. 7

Graves 1054 (E—975)90). Decatur Co.: A. R. Moldenke 295 (Fg).
DeKalb Co.: Alexander, Everett, & Pearson s.n. [Stone itn., Oct.
2 a as We Me ees se by UT (W—-1323260), 6457 ee Se
n. [Stone Mtn.] (F—351669); D. S. Correll 6579 (H--55781)
Davis 3121 rau. 5689 Pe Eggert s.n. SD. [Little Stone Mtn.
st R. M. Harper 187 (E—1191,9, © Ed, F—10636, W--38),819) ;
h & Pyron 373 (0)3 F. W. Pennell 4027 (Up—62323)3 Pyron &
ioe E.7480 (Up); J. K. Small s.n. (base of Little Stone Mtn.]
(F—180766, W—561)0); J. D. Smith s.n. [Stone Mtn.] (W—1323257);
Whitaker 101 (Ca—882l)0); Wurdack s.n. (Stone Mtn., 9-26-13]
(Cm). Dodge Co.: A. R. Moldenke 385 (Fg). Dougherty Co.: J. M.
Gillespie s.n. [Oct. 6, 1925] (Du--153267). Floyd Co.: M. G. Mc
Carthy s.n. [Rome] (Na—2223). Fulton Cos: P. 0. Schallert 6h,
in part (S). Glynn Co.: Pyron & McVaugh E.7478 (1 (Up); Van Halfern
s.n. (Brunswick, 6-23-27] (Ga, Ga). Grady Co.: A. R. Moldenke
30h (Fg, S). Jasper Co.: Porter s.n. [Monticello, 15)6] te
Jeff Davis Co.: A. R. Moldenke lenke 352 (F (Fg). Jefferson Co.: A. R
Moldenke 03 (Fg). Liberty Co.: Grimm sen. [6-27-)3] (Cm). Town-
des Co.: Quarterman 1266 (Va). McDuffie Co.: H. H. Bartlett 950
(Im, Po—6]818, Ur). McIntosh Co.: D. S. Correll 54h2 J,2 (No—18320,
We); H. H. Smith 2256 (F-27902, 01). Miller Co.: Duncan &
Hatcher 1707 (Ky). Monroe Co.: A. R. Moldenke 382 (Fg). Newton
Co.: Du Dunham s sen. [Nov. 2, 1928] (F—861059) . Oglethorpe Co.: Dun-
can 1426 (B (Bt——55291). haben Co.: Coken, Totten, & Harbison =
[Aug. 29, 1932] (Hi—-59397, Hi~59398) . palnend 1 Co.: Cuthbert
sen [July 15, 1899] (F1—20935), s.n. [July 20, 1899] a,
sen. [July 17, 190k] (Fl—20933). Rockilale Co.: Cron nquist 1,850
(Ca—789000, N, Tl), 5323 (N, Tl). Screven Co.: Ahles uses 9 (Hi—
201966); Boole "988 (Hi—183817) ; A. R. Moldenke 25 (Fg). Telfair
Co.: A. R. Moldenke 360 (Fg). Thomas Co.: A. R. Moldenke 3 (Fe)$
Mrs. A. P. Taylor 8 sen. (Thomasville, June 12, 1903] (Oa), Sone
[Thomasville, Aug. 28, 1903] (Oa), sen. [Thomasville] (Oa). Twiges
Cost A. Re Moldenke 38] 38h (Fg). Walker Co.: Ruth 229 (Ed, Ed).
Washington Co.: A. R. Moldenke 399 (Fg). Wayne Co.: W. H. Duncan
7655 (Ca--771503, No—1369). Sapelo Island: W. H. Duncan
(Hi—106314, Lb— Lb——l12769). Tybee Island: Grosvenor s.n. n. (F—-33316)+
County undetermined: Feay s.n. (Pa); J. A. Harris 0.19847 [Turtle
River] (Ca—610125); Keen & Keen s.n. [s. Ga.) (Pa). FLORIDA: Al-
achua Co.: Arnold s.n. [6/1731] (Fl—-20938); Collector undesigna-
ted s.n. (Io—127901); Gillman s.n. [Waldo] abaere al Kea s.Ne
1576733) (F1--20937) ; Loucks & & West s.n, (Gainesville, 6-27] (-
209h0); C. D. Mell s.n, . (Gainesville, Dec. 23, 1906] ty. 158h890))
Stuhr s.n, [Gaines ville, Nov. 1925] (Or—33128); A. D. Welch s-Me
(Gainesville, april 19,'1927] (Io—-127331). Bradfard Co. West &

1966 Moldenke, Monograph of Callicarpa 485

Arnold s.n. [Hampton, 15 May 1940] (Fl-—-32357). Brevard Co.: A.
Ae Baldwin | 13 (D--504178, Up--17067), 42 (Up—1706k); Collector
undesignated 226 d 226 (Gg—-208927) ; P. 0. Schallert 64, in part (Ur).
Calhoun Co.: A. R. Moldenke 273 (Fg). Clay Co.: E. L. Braun s.n.
ie Cove Springs] (Cn). Collier Co.: c.c. Deam 65825 (No—
87); Muenscher & Muenscher 14211 (N); Scrull s.n sen. (Naples, 5/
poy (F1--274,76). Columbia Co.: A. S. Hitchcock s.n. ie
City] (F--234221); McCulloch 0 (F1—209)11, W—1l0438),
[Lake City, June 17, 1901] (Ar—7506, Io—107150); Rolfs ay (Ga,
Ga, Ur), son. [Lake City, 9-17-93] (Ga). Dade Co.: SNe
[Miami, Aug. 1916] (H--26376, Up--70511); Buswell s.n. = Thais oF 22,
1936] (Bu); Cooper s.n. [Ft. Dallas, April 1559] (Ms); H. fe
Cowles 2-8 [188] (Bt--39883, Ob), s.n. (Miami, Apr. 'O] (Ur
Gratty s.n. (Miami, Jan, 29, 1933] (Io--10216); Duckett co
[Brickell Hammock] (W—1)85633); Elder 323 (H--73168); A. P. Gar-

cember 1911] (Up--63380); Hawkins s.n. [1-25-28] (Fl—20939); A.
S. Hitchcock s.n. (Miami, June—-July 1898] (F--231737); Howell . 872
(W—893578); Janish & Janish 552 (Du—3927l9); Macfarlane's party
sen. [Dec. 1901] (Up, Up); B. McAllister 92 (H—1L71); Meredith
8.n. [Miami] (D—62)967); H. N. Moldenke 296 (Go, S, Up, Ur),
3728 (N); Mosier & Safford 3 303 3 (Ar=-19781); | Edw. Palmer 02 (E-—
11918); Safford s.n. [Simpson's Hammock] (W—111687)) Safford &
Mosier 21 (W—1035857), 303 (W—1036142); J. K. Small 732) 2h (N),
7376 (N)3 Small & Carter 91 (D—-57091) , 651 (D--569957); Tides-
trom 6959 (Ar—6959, Ar——10065, Ca—882h5, H—67898, Mi-—photo,
W—751531). DeSoto Co.: Harshberger s.n. [June 3, 1912] (Up—
63010), sen. [Ft. Ogden] (D—-560758); E. West s.n. [Lacy, 23 Sept.
1938] (F1—153). Duval Co.: Curtiss 1971 [fls., June] (Mi, “a
Mu--1648, Or--83)0, P, Vi), 1971 [frs., Sept.] (Ca—25093, D—
550100 Meg ee * g—119305 “F—=148426, wie Mn—15350, Mu—1648,
— B. V, vt), 1387 (ca Risers “909, E—766880, W—
218281), * 41935 (fis., June 6) (Al, 101990, 14,7392, Io—
3876, Mm—15353, W—22hh9h), Loss (eve:} faeces), 6189 [June
12, 1898] (Fl—209l2, Hi—5939h, Ur, Vt, Yu), 6189 [Aug. 30, 1898]
F1—209)2, Hi--5939h, Ur, Vt, Va); Fredholm am 1h, (Pe—85766), 5294
(E—818111, W—717178); A. S. "Hitchcock s.n. 3.0. [Jacksonville] ifn
231890) ; Keeler sn, (vicinity of Mayport and Jacksonville] (01);
Lighthipe 105 (Ob—50623), sen. [South Jacksonville, June 17, '97]
(Hi—-59393), 8 8.n. [South Jacksonville, Sept. 15, '97] (Hi—59393).
ane Co.: “Brinker 12 (I). Flagler Co.: West & Arnold s.n.
a ler Beach, 15 Apr. 190] pocreadeie grag , 10/10/
] (Fl 3284) « Franklin Co tmore Herb. 1720b
332093) , 17208 (E-11915h, ota | is, W--335089); A. W. Chapman 0

4,86 Pt 2 ODLODG IA Vol. 13, no. 7

alt Spee son. (E--119233, W—-56142); Herb. Chapman
5980 310); McAtee 1764 (W--51798); Saurman 62 (W—
e588), "69 (E—119153), s.n. [Apalachicola, 1567] (Pa), s.n.
[Apalachicola] (Up--17062). Gadsden Co.: H. Foster 11 n—
20931). Glades Co.: Lovett 201 (H—~3864). Hamilton Co.: West &
Arnold s.n. (White Springs, 11 Apr. 190] (Fl--32027). Hernando
Co.: R. N. Jones s.n, [vicinity of Brooksville] (W--1086820).
Highlands Co.: Hunnewell 10483 (Ua--36989). Hillsborough Co.: F.
S. Blanton 6662 (Du--2011)8, En, H—22121, Hi--591,00, Po--220816,
S); S. M. Deam 2 2738 (Dm, Ok, Po--26768h) ; ee P. Garber s.n. (Tam-
pa] (F—139684, Us, W—265077) ; Ray, Lakela, & Patman 10133 (Hi—
201,00). Jefferson Co.: Herb. Exploration Party 1939 sen. [Wa-
cissa, 16 Mar. 1939] (F1—-30168); A. S. Hiteheork oa [June-July
1898] (E—-119181, F——-234220). Lake Cos: A. Se Hiv S.H cheock s.n.
[vicinity of Eustis, June & July 189] (Ka), s.n. ai stis, 1609)]
(F1—-209h4), s.n. [Eustis] (E—11923), F—~233197); Nash 725 (Ca=-
104989, E--119236, Ed, F-~38)16, Mm--15352, Vu, W—228226), 215
(W—252136, W=-3096)0). Lee Co.: A. S. Hitchcock 272 (E~1191uh,
F—101341, W—-387306), s.n. (Myers, July--Aug. 1900] (Ka); C.
Skottsberg s.n. [Fort Myers, 20.5.1935] (Go); J. P. Standley 192
(E--819012, F-73957, Po-~17h969, Ur, W--910889). Leon Cos: Re
K. Godfrey 53359 (Hi--193406, N). Levy Co.: A. P. Garber s.n.
[Nov. 1877] (Io, Vt); Janish & Janish 339 (Du--392743, N). Liber-
ty Co.: West & Arnold s.n. [Torreya State Park, 22 July '40] (Fl—
32355). Manatee Co.: J. H. Simpson 40 (D~526671, F--231129, Ge—-
163107, W—334922, W—33h93h); Ss. MT. Tracy 7531 (Up--50863), 753
(cax-181832, Cm ae B-11932h, B-119 325, E~-119 326, Ed,
o=-{\8621 , Tr, W— yy 2191). Monroe Co.: A. De "Welch
1s oe Sosa. se ke 3. eee s.n. [Fernandina] (E—- _
Orange Co,: Barrows (Winter Park, 189] (Dt); Mac
sia 8.n. [Sanford] (D— a Aare 405 (H—S12659)5 Vester-
lund s.n. [June 1889] (Us, Us, Us). Osceola Co.: Mearns 6 (W—
391101); Singletary 151 (H-1917). Palm Beach Co.: G. L. Bates
s.n. (Jupiter, 1089] ~(Fe=~7573), 8 sen. [Jupiter] (W—5611)1); W.
B. Fox s.n, [April ls, 1945] (We); A. A. S. Hitchcock 1489 (F--
233566); E. West sen. [Lake Worth, 6 Sept. 1939] (F1--30748). Pas-
co Co.: Arnold s.n. n. [New Port Richey, 23 Sept. 1939] (Fl--31073);
Carpenter s.n. [New Port Richey, October 22, 1936] (Vt); O'Neill
oa (St. Leo] (E—953153). Pinellas Co.: Se M. Deam 1958 3 (i
)j Tidestrom 7014 (W--751546); Me H, Williams s.n. [Pine
Point] (D—6 ee Polk Co.: 0. E. Jennings s.n. [Fort Meade,
te Dec. 19 Cm, Cm, Cm); Jennings, Jennings, & Jennings 5-M-
(Dec. 27, 1916] (cm); McFarlin 2m (i) 5573 (Mi); 0 Ohlinger 109
(E—119152, F—230)69, F1—209h3, Ga, Ur), sen. [May 17, 1894] (Ga)s
J. De Smith s 8.n. [Peace e Creek] ea sessed): Putnam Co.: West &

1966 Moldenke, Monograph of Callicarpa ),87

Arnold s.n. [East Palatka, 7 June 1910) (Fl~-32351). eg Johns
Cos: L. C. Reynolds 311 (W--1323263); M. C. Reynolds s.n. (Dry
sandy places) (Pr), s.n. (Up—17068); aP J. Dd. Smith 2 423 jWe-as2326}
West & Arnold s.n. [Crescent Beach, 7 June 19]0] (Fl--32356).
Saint Lucie Co.: A. C. Beal s.n. Con Walton, Jan. Fe (Ur);

ea 2ande 2192 (Z). Seminole Co.: D. S. Correll 6367 (H--
55787); P. O. Schallert 64, in part (B, B, Hi--54927, Ok, S, S).
Suwannee Co.: Herb. Exploration Part: 1939 sen. (Toktucknee.
Springs, 17 Mar. 1939] (Fl--30167). Volusia Co.: H. C. Beardslee
81 (O0b--97271); Hood s.n. [Orange City] (E--850163); B. H. Patter-
son s.n. (Nov. 27, '17] (Cm), s.n. [Nov. 26, 1919] (Cm). Walton

: Ashe sen. (Mossyhead, 7/31/26] (Hi); E. G. Hume s.n. [6/2/

38] ‘(FI=12388); 3 Tyson 642 (Ur). Key Largo: Me Bush sen. [Key
Largo, Feb. 29, 1936] (Cm); Janish & Janish 495 [Key Largo] (Du--
392747); H. R. Totten S.n. (December 28, 190] (Hi—22705). Coun
ty undetermined: G. H. Bates s.n. [1889] (Bl—-4.2206, Ka, W--561),1);
A. W. Chapman [South Florida] (E—119319), s.n. [Florida] (Gg—
31966, “0k); C 3 Collector undesignated § sen. (Ky); “Curtiss s.n. [1875]
(F—31,827, F—~259936, F—-259980); T & Gray sen. y sen. (Us). ALABA-
MA: Baldwin Co.: Demaree 35926 aay, ete % Co.: Eg,

[Blount Springs] (E—11916h); J Je D. Smith 153 (W-1323256). Chero-
kee Co.: G. F. Freeman s.n. a out Mtn., July 14, 1905] (Ka—

67033). Cullman Co.: M. [, Morgan a 4 sii Co.: Koepper &
Justice Son. en Millbrook, Fine 20, 1943] (Dp--38370 Etowah

t s {[Blaikireek Falls] (E—119163) ; G. N. vase SN.

[oadedee i878} (Pr, Vt). Jackson Co.: A. R. Moldenke 77h (S).
Jefferson Co.: A. S. Hitchcock s.n. [Birmingham] (F—23h 792). Law-
rence Co.: A. R, Moldenke 767 (S). Lee Co.: C. F. Baker 8433 (E—
119176), s.n. pk 9-20-96] (Po--6,809) ; He Re Baker s.n. [Au-
burn, 21 August 1929] (H—-107723); Earle & Baker s.n. {Auburn, 7/
6/1897) (E--119177, Ka, Ur), sen. [TIeABSTT Cb b=-50622), son.
|Auburn, 9/20/1897} (Ka); L. He H. Pammel sen, [Auburn, June 3, , 1921)
(Io==1020k) ; Pollard & ee a (W--4271L). Madison Co.: Howell
505 (W--99630); W. H. Rh sen. [near New Hope] (Hs); Shimek
5.0. fineaan i Ge “y 78721) . Marshall Co.: A. R. Mol-
denke 772 (S). Mobile Co.: Graves 649 (W—-1085022); Cc. ., Mohr s s.
ne ee, July] (Du--90915), sen. [Mobile] (W—-771910); | F. H.
Sargent s.n. [June 18, 1950] (Ok). Montgomery Co.: Koepper & & ; Jus-
tice s.n. [near Montgomery, June 27, 1943] (Dp--38438); L. He Pam-
mel | 8.n. sn. (Montgomery, May 30, 1921] (Io—102655). Tuscaloosa Co.:
W. H. Rhoades s s.n. (near Tuscaloosa, July 1926] (Bt-—30605), s.n.
[near Tuscaloosa, | Aug. 1927] (N). Wilcox Co.: Buckley s.n. (July
180] (Br). Cousity undetermined: Buckley s.n. [July 1840] (Br),
Sn. (Ca-—379992); Peters s.n. [July h, 1867] (Pa); G. N. Vasey s.
n. [1878] ee SOT? Fo—-502, Ka); Winchell s.n. (Ms.) MISSISSIP-

4,88 PHYTOLOGIA Vol. 13, no. 7

PI: Adams Co.: A, R, Moldenke 743 (B); Shimek s.n. [Natchez] (E—
119190, F—78633). Claiborne Co.: A. R. Moldenke mke 71 (B); J. M.
White s.n, [Martin] (Na—5711). Forrest Co.: F. A. B Barkley 1 a3.
(Ok—18135). Hancock Co.: Bonck sen. [June 9, 1940] (Tl). Har-
rison Co.: L. H. Pammel s.n, 1. (Gulfport, Jan. 192h] (Io—110972) ;
Pollard 1143 (E—11919h, F—-171238, W—-271618) ; S. Me Pee 4982 4982
(Dm, E--119158, Ea, F—81041, Hi—S9l,02, W~3)1065); Trécul 97
(P, P, P). Jackson Co.: Demaree 3 32091 (Ok); Diener ‘- & (T1); i Mac-
farlane & Goertz s.n. [June 12, 1905] 905] (Up—39350) » SNe [Ocean
Springs] (Vu); J. Skehan to (Ur), 22420 (Ur), s.n. [Seymour &
Earle 111] (E—~9]2930, 364183, Hi—59395, Lb—16h21, Ur),
s.n. [Ocean Springs, 6/7/98] © e- 219282), s.n. (Ocean Springs, 8/
15/95) (E--119193). Lafayette Co.: A. R. Moldenke 766 (S). Lau-
e Co.: Schuchert s.n. [Meridian] (W--200833). Lowndes Co.:
- R, Moldenke 763 (B). Noxubee Co.: A. R. Moldenke 761 (S). Pa-
nola Co.: Eggert s sen. [16 Sept. 1896] (E—119162). Pontotoc Co.
A. R. Moldenke 765 (B). Scott Co.: W. H. Rhoades s.n. (Lake, July
1927] (N). Sh Sharkey Co.: A. R. Moldenke . 757 (B). (B). Simpson Co.:
Holt 3 (W--587203). Warren Co.: W. H. | W. H. Rhoades sen. (Vicksburg,
July 1927; H. N. Moldenke 17079] (Bt, N, St). Winston Co.: A. Re
Moldenke 756 (B). County undetermined: Gerhart 59 (N)5 Montgomery
sen. (Nov. 1 1920] (E-~859168). TENNESSEE: Anderson Co.: Wilkens
3713 3 (Up). Blount Co.: Jennison 37 (0, 0), 503 (0). Coffee Co.t
C. B Black s.n. (Manchester, June 20, 1930] (Go). Franklin Co.: Eg-
gert s.n. [Cumberland Mtn.] (E—119166); G Gattinger s.n, [Cowan]
(0). Grundy Co.: W. H. Rhoades s.n, (Mt. Eagle] (Hs). Hamilton
Co.: Anderson & Jennison 1329 (G), sen. [Signal Mtn.] (0); Jenni-
son & Anderson | s.n. [Chattanooga] (E— 978480) 5 Underwood & Sharp
2383 (0). Knox Coe: G. Andrews s.n, [Lyon's View] (E—770172)}
Ruth 193 (E~119161), 498 (W—3 W—SLB6TS), 515 (E~-119147), s.n. [Aug.
25—Sept. 10, 1898] (Ur). Lauderdale Con Sharp, Fairchild,
Clebsch, & Clebsch 7954 (No—16)9)). Lawrence Co.: Clebsch,
& Clebsch’ 20 (N). Marion Co.: A. R. Moldenke 776 (S). Morgan Co.
Coffman 91 . (W—178876), s Son, [Oct. 1889] (Ww—! (W—-,78875). Shelby Co.:
C. E. Moore 79 (W—~1)66116). MISSOURI: Taney Co.: J, A. Steyermark

69453 (8, Ur). ARKANSAS: Arkansas Co.: Kellogg sei Ler
Post] (E—119322, E—879215). — Co.: Demaree 23903 (Ca—
882,37, N). Baxter Co.: BE. J. 477 (E71 ae 71, E—717372) «

Bradley Co.: Demaree 13565 (iy, om “T6305 (N), 19535 (N). Cleburne
Cow: E. J. Palmer 69) 6981 (E— (E——752702, E—787155 2 Crawford Co.: De-
maree 1502b (E—960171), 15277 (N, Se—55736)} MeKinney 91 (AU);

E. J. Palmer 22199 (W-—~121118); Shearer s.n. (Aug. 28, 1919] (U4).
Desha Go.: Howell mell 613 (¥-~615530). tse Drew Co.: Demaree 13bL3 (N).
Faulkner Co.: E. S. Ford 98 (Hi—203728, It), 119 (It). Garland
Co.: V. H. Chase 9753 (Ur); Demaree 15361 (N, Ok); Earll s.n. [Hot

1966 Moldenke, Monograph of Callicarpa 489

Springs] se fegly Letterman s.n. [Hot Springs] (D—563760, E-
119167, E1191 Va ae 965733) Meek s.n. [Hot Springs,
8-11-89] Gennes H, Rhoades s.n. [Hot Springs] (Hs); R.
Runyon 1203 (N), 1461 (W=1567991); Se cully 982 (Sd—~2372h); Tre-
lease s.n. [Hot Springs] (E—119179) ; Tweedy 1,88 ly 488 (Au). Hempstead
Co.: Bush 3837 (E—119316); Demaree 21506 (N). Hot Spring Co.:
Demaree 18883 (Bt——55321, Io—15222), Ok), Ok); Trelease s.n. [Magnet
Cove] (E—-119178). Howard Co.: Demaree 9742 (Po--2 (Po--2091)2). Izard
Co.: Demaree 23395 (N); Hubricht B.2277 B.2277 (N). Jefferson Co.: De-
maree 13937 (N). Johnson Co.: Demaree 19977 (N). Lee Cos: Dema-
ree 12973 ree 12973 Sepia kia, N). Logan Co.: Welch 119) (Dp, Lu); Mrs
H. 8. You Young s.n. iy fe 21, 1925] (Cm). Marion Co.: E. J. malnar
Seay 1 7553, _W-580519) Miller Co.: 0. M. Clark 3979 (E—
2h) ; 2h491 (N); Heller & Heller ]158 (E—119180, w#—

a, ae 8.n. Senge ‘(D=-563883, E—~770)85, W—
985731); Pringle s.n. n. [Texa » 30 July 1886] (Vt). Montgomery
Co.: Demaree 9603 jcognns 0), 39945 (Lb——53420). Nevada Co,:
Hollister 28 28 (W—587722). Phillips Co.: Demaree 19760 (Ca--882),38,
Ge—316191, | N). Pike Co.: L. H. Pammel s.n. [6-10-1911] (Io—
81687), s.n. [6/11/1911] (Io—55068). Pope Co.: G. M. Merrill 21
(Cp, Du—2)6556, E-103000h). Pulaski Co.: R. J.B Bailey s.n. SoM.
(Oct. 17, 1935] (cm), s.n. [Oct. 2h, 1936] (Cm), : S.n. [Nov. 1936]
Ng! Coville 13 (W--210608); Demaree 37717 (Ib—67h7); Eggert s.

- [7-3-9h] (E—119159, E~119160, E—-119170, E~119191, Po—
i808 08); Engelmann 205 (E—~119303, E—11930h); Hasse 2063 (D—
586827), Sone [Little Rock, June 1886] (Al, Ca—192607, Du—-90913);
Letterman s.n. [Little Rock] (E--770252, W-~985732). Saline Co.:
J. M. Gre M. Greerman 4301 (Herb. Drushel 5311) (E--902885, Ur). Sebas-
tian Co.: Bige Bigelow s sen. [Fort Smith] (W--56129). Sevier Co.:
Brinkley 38 (F—8964)2), 338 (Au, Au, F—899057, N); Fassett 27677
(Vi). Washington Co.: P, Me Beck 85 (Ok). Yell Co.: Demaree
18197 (N). County undetermined: | Buchholz sen. {central Arkansas]
(E—892615). LOUISIANA: Allen Par.: r.t A. R. Moldenke 711 (S). Avo-
yelles Par.: A, R. Moldenke 727 (S). “Beauregard Par.: | A. R. Mol-
denke 710 (S). Bienville Par.: A. R. Moldenke 68) (B). Bossier
Par,: eg R. Moldenke 680 (B); Penn s.n. 1. (Aug. | 26, 6, 1941] (Tl);
Shoop s.n. (Benton, July 20, 1935] (F—902h19); Trelease sn.
[Aldenbridge] (E—119189). "Caddo Par.: Moldenke & Woods (s)
Calcasieu Par.: Allison 67 (W—514026), 319 (E--119323, F—28651,
W—-514121); A. Chase 6095 (Mi, W-—503419); Gander 7601 (Sd~25188);
K. K. Mackenzie 531 (Dt, E--119157, E—119329, Ka); Solereder s.n.
[Lake Charles, Sept. 1833) (Mu—l111). Catahoula Par.: A. R. Mol-
denke Thy (B) . DeSoto Par.: A. R. Moldenke 692 (S). East Baton
Rouge Par.: S. M. Lewis s.n. Touly 15, 1951) (Ur); Romm s.n. [July

490 FRes.eULbOGiIa Vol. 13, no. 7

15, 1951] (Ur). Evangeline Par.: Ewan 19342 (Tl); A. R. Moldenke
713 (S). Grant Par.: Ewan 17495 (T1); A. R. Moldenke 725 (B).
Iberville Par.: Bush 362 (E—119156, E-—-129945, W--318509). Jef-
ferson Par.: H. P. Riley s.n. (Grand Isle, May 30, 1938] (Se—

47504). La Salle Par.: A. R. Moldenke 745 (S). Livingston Par.:
Correll & Correll 9167 (H—-75520, N). Natchitoches Par.: L. E.

ex L06 (Ne), Lil (Fx); A. R. Moldenke 695 (S); E. J. Palmer 792h

D—-69 35,8 ; E—~793709, Gg—~162157). Orleans Par.: T. Drummond
254 bis (Lu), 254 ter (Ed, Us), s.n. [vicin, New Orleans, 1832]
(Lu), s.n. [New Orleans, 1833] (Ed, Ed, Lu). Rapides Par.: C. R.
Ball 5h1 (E--119145, F-93548, W—382819); A. R. Moldenke 726 (S).
Red River Par.: A. R. Moldenke 691 (S). Sabine Par.: L. E, Fox
4LO (Fx); A. R. Moldenke 696 (S); Shinners 20235 (Ca—l6300).
aint Par.: Ewan 18869 (Tl). Saint Tammany Par.: Anect 67
(W--1087035); Arséne 11376 (W—1031498); Bomhard s.n. (Mandeville,
Dec. 8, 1928] (Cm); Ewan 17458 (T1), 17681 (T1); A. R. Moldenke
2h6 (Fg, 2); H. P. Riley s.n. (Mandeville, Oct. 25, 1936] (Se—
Tgho1). Union Par.: R, Gaines s.n. [6-23-50] (Ca—86357). Vernon
Par. A. R. Moldenke 702 (S). Washington Par.: D. V. r 8.0.
(Bogalusa, 1931] (Mi); A. G. Watkins s.n. [6-)-33] (Tl). Webster
Pars: A. R. Moldenke 683 (B). West Carroll Par.: Moseley s.n.
(Holly Ridge] (F—282161). West Feliciana Par.: Ewan 187h6 (Tl).
Avery Island: Thieret 16087 (W--2)22076). Jefferson Island: Thie-
ret 15945 (W--2422031), Weeks Island: Thieret 1593) (W—2h22028) .
Parish undetermined: T. L. Andrews 49 (Io—107277)} Barbe son.
(Basse Louisiana, 1839] (P); Collector undesignated s.n. [Herb.
Jussieu 1061] (P, P); Prat s.n. (Louisiana; Herb. Jussieu 5061]
(P); Teinturier s.n. [1836] (De). OKLAHOMA: Bryan Co.: W. S.
Blain 285 (W—1623326), s.n. (Durant, July 8, 1935] (Bt--29669)3
Bragg 95 (St~-15607); E. J. Palmer 10h)1 (Du—205596, E—80)539,
Gg—-31971). Choctaw Co.: 0. F. Hay 103 (Ok); Kilgore 1h7 (St—
13738); E. J. Palmer 8345 (E—76)183, E~793710, Gg——31967, Po—
220832). Cleveland Co.: D. Simpson s.n. [6-23-38] (Ok). Latimer
Coe: Me Cox 62 (St--17514); G. J. Goodman 20h7 (E—~1087898, N, Ok—
10236); A. Hill 83 (St); C. W, Hodges 61 (St--17513); Wallis 7530-
1 (St). LeFlore Co.: 0. W. Blakley 1,98 (E~-782299), son. [Ste-
vens 1498] (Ok); Demaree 15723 (N), 16036 (N, Ok); Ortenburger s-
n, (Kiamitchi River] (W—13650)2); E. J. Palmer 20550 (W--1119229),

4LO86 (N); G. W. Stevens 2688 (Du—67961, E—781217, Ok, St~-9178,
Ur, W--910034); Stratton 1832 (St—26146, St--261)7). Love Co.?
Goodman & Rice 5678 (Ok). Marshall Co.: Alder s.n. [July 2, 1950]
(Ok); Basler s.n. [Island no. 2, July 2, 1950] (St); G. J. Goodman
iil2 (S), 5674 (Ca—81900, Ok, Ur), 6015 (Ok); G. Morgan 6 (5);

Starks sen. [25 July 1950] (Ok); Waterfall 11856 (St), 12299 (St).

1966 Moldenke, Monograph of Callicarpa 91

McCurtain Co.: Estes 101 (St--17488); M. Hopkins 2883 (Ok, St—
14799); Hopkins & Cross 2338 (Ok); Jamison 62 (St—-17879); Little
& Olmsted 11), (0k--10232), 1507 (Ok--10233), s. s.n. [June 28, 1930)
(Ok--1023h); Neese 102 (St-=18301) ; Nelson, Nelson, & Goodman
5ho2 nee aah a2 © (Ok); G. T. pees 2648 (Ok), 2660 (Ok),
2826 (Ca 78); C. C, Smith 908 (Ba, Ok); Stratton 573 (E—
916679); W Pe sen. [July 9, 1955] (Ok). Pushmataha Co.: Herb.
C. Thomas sen. [6-15-1937] (0k); Hopkins & Cross 2218 (0k); Water-
fall 302 (St--9176, St--9177, St). Sequoyah Co.: Wallis 6140 (St),
1996 (St); Waterfall 9533 (st). Tulsa Co.: pe Bebb s.n. [bank
of Salt Creek] (Ok). County undetermined: C. C. Parry sen. [In-
dian Territory, 1878] (Io). TEXAS: Angelina Co.: oe sen. [8-7-
34] (Au), sen. [8-10-1934] (Au), s.n. [Aug. 12, 1934) | (Au, Au, Au), 8.
n. (Sept. 1934] (Au, Au). Aransas Co.: Cory us7h2 (Au); F. Be
Jones 4073 (Ww). Austin Co.: H. Wurzlow s.n. [Industry, 1891]
(Ur). Bastrop Co.: C.C.C. Plant Project s.n. [Bastrop —

Tharp 253 (Au, ui), Bene saaeaoee State Park, Fall 1936] ia), 8.
n. n. [6-15-37] (Mi). Bexar Co.: Berlandier 1678 os Biltmore Bae
17203 j (Dm); Bush 818 (E--119148); Clare 273 3 (Mi);
1832 (Ca~~10)988, E—-119235, Ed, Mm—15351, ee ast),
ce G. Jermy 121 (W—315720) 5 J. Jermy 8 (501895); Mets 273 (Ms,
Me Bee 270) 8 sen, (San Antonio, Aug. 28, 1931] (Dp); C. C. Par-
sen. [En route te from San Luis Potosi to San Antonio, Texas, “Au
gust 1978] (Io); Wilkinson 12 (E—-119146), 119 (E--119187). Blanco
Cost E. J. Palmer 12862 (Ca--25599, E--827554, Gg--31968). Bowie
Co.: Milligan sen. [Dalby Springs] (W--503756); Ward s.n. [west
of Texarkana] (W--1l7597). Brazos Co.: Bain 19 (0); W. S. Jen-
nings 71 (0k); L. H. Pammel s.n. [College St Satter July Ts88) (I0—
1201), s SN. [College Station, Jul. 26, 1921] (Io—102010), s 8.N.
(College Station] (E--118818, B--119300) ; Reeves 188 (N, N). Camp
Co.: Moldenke & Woods 559 (S). Cass Co.: Cheng s.n. (Atlanta,
14-26] (Au). Chambers Co.: G. L. Fisher 7 (F-67719). Cherokee
Co.: Trelease s.n. [Forest] (E—119231). Colorado Co.: E. J. Pal-
mer 6592 (E--755442, W—-588658). Comal Co.: Lindheimer 297 (E—
pleas E-——119309, Ka), 1067 (Au, Br, agSoyasan: D—535535, Ee=
Ed, F--218267, Me, Me, Mu—l056, Na--29097, Ok--18037,
Tp hB38e *"w—5S02L40) « Dallas Co.: Dappri Boots 7176 (sm); C. L. Lun
dell 13930 (N); Lundell & Lundell 7048 (Au, Mh, Mi, N, N)3 Rever—
chon 745 (E—119150, E—119330), 3905 (E—119172, F-11933, ¥—
501158), s.n. [Feb. 1872] (E-—119315), s.n. (Dallas, 1876] (F--
81519h); Ruth 1333 (Sm); Shinners 8482 (Ca—733262, N, Sm); Ster
henson s.n. [June 19, 1928] (W—132573). DeWitt Co.: C. Be
Mueller 5028 (N, Sm, St); M. Riedel s.n. [6-3-2] (Au). “Ellis Co.:

492 PHYTOLOGIA Vol. 13, no. 7

Moldenke & Woods 576 (S); Tharp s.n. [Onion Creek, 6-18-32] (Au,
Au). Fayette Co.: Forshey s.n. [Rutersville, 1857] (Ka—95509);
Matthes 20 (Dr, 01). Freestone Co.: Lundell & Lundell 12939 (N).
Goliad Co.: C. B. Williams 50 (D--6289)8, N). Gonzales Co.: Mc
Bryde M.517 (Nt), s.n. [6/29/31] (Nt); Normand s.n. [5-31-28]

(Au, Au); E. D. Schulz 2666 (La), s.n. [Ottine, Aug. 192] (Wi),
Son. [Gonzales] (Wi, Wi); Tharp son. [Palmetto State Park, 6-30-
39] (N). Grayson Co.: C, Richardson 15 (Au), son. (Denison, Oct.
17, 1933] (E—1072509); Whitehouse 17187 (Mi, N). Gregg Co.: C.
L. York s.n. [Fall 1937] (Au). Guadalupe Co.: Groth 1g2 (Dt, F--
181881, Gg—31970, Io—75833, W--717779). Hardin Co.: Lundell &
Lundell 11537 (N). Harris Co.: Armer s.n. {Houston, July 1927]
(Sm); Boon 3h (Au), 378 (Au, N); 0. Degener 5233 (Ms); G. L.
Fisher 88 (W--503159), sn. (Herb. Banker 2307] (N), sn. [Houston
June 2, 1912] (Ca—8824)8, Ew), Sen. [June 9, 1912] (Vi), s.n.
(Houston, Aug. 1, 1912] (Ew), sn. (Houston, Aug. 5, 1912) (Vu),
sn, [Aug. 30, 1912] (Vi), s.n. (Houston, June 2, 1913] (Fs), sn.
(Houston, June 2, 1914] (Vu), sen. [Aug. 15, 1915] (Gg—-200881),
s-n. [Houston, Sept. 14, 1930] (Bt--37637), 8.n. (Houston, Aug. h,
1936] (Fs), s.n. [Houston, June 14, 1947] (Go); O. Mueller s.n.
(Rose Hill, 8/28/1929] (Wi); L. H. Pammel s.n. { Houston, Jul. 27,
1921] (Io—~10)611); Tharp sen. (9-1)-33] (Au, Au); Tweedy 118 (W—
147598). Harrison Co.: A. R. Moldenke 676 (S). Henderson Co.:
B. B. Harris s.n. [7-10-27] (Nt), sen. [8-6-27] (Nt); A. R. Mol-
denke 667 (S). Hill Co.: Shinners 10537 (Tl). Hopkins Co.: Mol-
denke & Woods 566 (S). Jackson Co.: A. R. Moldenke 20) (Fg, S).
Jasper Co,: Correll & Correll 12536 (Sm); Cory 49833 (N, Sm); E.
R. H. s.n. [Kirbyville, Dec. 21, 1902] (Io--82905). Jefferson
Co.: Hook son. [Beaumont, Sept. 10, 1932] (Au, Au); A. R. Molden-
ke 225 (B, Fg, S). Liberty Co.: V. Lehmann s.n, (Cleveland, 6-2-
3h) (Au). Marion Co.: Moldenke & Woods 554 (S). McLennan Co.:
Pace 2h (E—119312); Posey 10 (Nt); L. D. Smith 615 (St). Mont~
gamery Co.: Dixon 27 (F—238186); Warner 1 (Hu). Morris Cos:
Moldenke & Woods 556 (S) » Nacogdoches Co.: Crausley sen, [July 18,
igh] (Au); Lundell & Lundell 9615 (Ld, Mi, N), 9663 (id, Mi, N)-

ange Co.: Kirn s.n. [Orange, 10-18-22] (Wi), sen. (Orange, 10-27-
22] (Wi), son. [Orange, Oct. 27, 1927] (Wi); M. S. Young s.n. [0-
range, 4/19/18] (Au). Polk Co.: Girvin 101 (Au); Tharp, Turner, &
Johnston 54713 (St). Raines Co.: A, R. Moldenke 660 (S). Red Ri-
ver Cos: E, Adams 19 (Nt). Robertson Co.: F. A. Barkley 13590 (au,
Ca~—-882lh, N, Ok, St). San Augustine Co.: Crocket sel. (W—
500016). San Patricio Co.: F. B. Jones 2139 (Ww). Shelby Co.: H.
B. Parks Jr. 2310 (Au). Stephens Co.: M. L. Grant 8193 (Ok). Tar-
rant Coo: Ruth 198 (au), 1333 (La, Nt, 0, W—-1260308), s.n. [Hand-

1966 Moldenke, Monograph of Callicarpa 493

ley, Sept. 12, '10] (Au), sen. [July 20, 1930] (0); Whitehouse
16158 (Mi, N). Travis Co. Bo, Bogusch 69 (Po--161327); Carener &
Studhalter 4613 (Au); Krodel 35 (Au, Au); McKee & Wesley 3860
(Au); R. H R. H. Pati Painter s.n. [Austin, 8/6/23] (Ka--70282) ; H. B.
Parks sen. (Aug. 20, 1929] (Wi, Wi); R. 3. Payton 54, (Au); Rip-
perton & B. & Barkley 1452) 152k (Au, Ca—882)46 }3 ee D. ,D. Schulz s.n. [Aus-
tin, 7-11-1920] (Wi), sn. [Hamilton Pool, 7-11-1920] (Wi), son.
. 0 mi, s. of Austin, 8-13-1921] (Wi), s.n. (Austin, August 1 1921]
Wi, Wi); Strandtman. sen. [July 17, 1941] (Au); L. C. Taylor 3038
(ca—697811); 7 Tharp 4UL16 (Au) , sn. (Austin, 7-21-0] (Ca—
882443, N, P1—1e1ehh,, Se-—7561h, Vi Vi), sen. (Austin, 8-31-)1]
(Ca —882))1, N, Sm), s.n. [Onion Cr., 10-30-)1] (Ca—882))2) ;
York & Wolf 46. (Au). Trinity Co.: deodrm $ s.n. [June 1936) (Au).
Upshur Co: re R. Moldenke 674 (B, Fg). n Zandt Co.: Bass 18
(Nt); A. R. Moldenke 671 (S); Sides 7 ler Whitehouse 16152 (Mi,
N). Victoria Co.: Ferris & Duncan 3258 258 (Du--125410, E--8934%,
Gg--31969). Walker Co.: Dixon a3 on 32 (F=-237968); Cc, Hartmann s.n.
(Huntsville, summer 1912] (Au, Ca~-882447). Waller Co.: B. Hall
438 (g—-119182, F~,54359, Pa, Po--85767, W--56131). Washington
Con: C. C. Albers 32022 (Au); Brackett 253 (Au), s.n. [7/1/39]
(Au). Wilson Co.: Edw. Palmer 1054 (Pa, W—56130); Parks & Cory
15121 (Tr), 15123 ‘(i). 15124 (Tr). Wood Co.: A. R. Moldenke 669
(S). County undetermined: Collector undetermined Th (E—-119169) 5
Eggert sen. (E—-119168); Gaut 228 (W--50)928); Lindheimer 13h (E-
119302, E--119306, E--119307); Schlottmann s.n. [1886] (Ah);
Tharp s.n. (To—-10),905) 5 M. J. White xix [Chronister] (Au).
BERMUDA ISLANDS: Main: Brown & Britton 222 (Ca—-370289, D—511260,
F—203802, Lu, Up--55080, W—-52)873); 0. y. Degener sen. [Paget
Marsh, Aug. 13, 1921] (Ms). CUBA: Pinar del Rfo: Acufla 10715
(Es); Britton, Britton, Earle, & Gager 676) (W--6970)8); Earle
751 (F—-285879); Ekman 10786 (N); Leén 400 (Ha), 5106 (Ha), 17897
(Ha); a & Shafer ‘er 13607 [Shafer 3215] (Ha); Marie-Victorin ae
3 (Vi, Vi); Vi); O'Donovan 5306 (P o~~64,806); Palmer & Riley 547 547
Tt), Shafer 1090) (E—118017, F—-325328, W—699179) ; Van
Hermann 86), (F—155) ag ws W--845265) , 65 (Ca~10133,
F—20072h) 5 Zarragoitia 5306 (F—185737, W—8)5510). Province un-
determined: C, Wright 231 1/2 (Herb. Sauvalle 1772] (Hv), 3168
[1860—-186; Herb. Sauvalle 1772] (Hv), 3168 [1865; Herb. Sauvalle
1772] (Hv, Hv), 3168 [Pulneios] (Pa), 3168 (E—119130, W--56138,
W--1361168). ISLA DE PINOS: Britton, Britton, & Wilson 1500) (ca,
F-59550, W—7932h2); Leén 17897 (N); Palmer & Riley 1060 (W—
372982); A. A. Taylor 230 (F—-120571, W--426935). CULTIVATED:
Alabama: Justice ‘3.n, (Montgomery, June 26, 1942] (Ba). Belgium:
Herb. Martius s.n. (Br)s M. Martens s.n. [h. b. Lov. 1838] (Br);

Lok PHYTOLOGIA Vol. 13, no. 7

Lejeune s.n. [h. Leod.) (Br). California: Bellue s.n. [Sacramen-
to, Jan. 1942) (Gg--293960). Cuba: J. G. Jack 8664 (Gg—237835).
District of Columbia: Vasey s.n. [Agricultural grounds, Washing-
ton) (W-~56133). Florida: Wedding 115 (Ar--19783). France: Herb.
Harvey 8.n. e R. P. 1819) (Du--166513); Herb. Hort. Paris. s.n.
(B); Herb. M msp. sn. [1807] (Dc). Georgia: Tidestrom 112 (Ar—
19784), A nite (Ar—715823). Germany: Herb. Btttner s.n. (B); Herb.
Hort. Acad. H ad. Helmstedt gen. -[Aug. 1767] (Ed); Herb. Hort. Bot. Mo
nac, 8.. sens (Mu——1h26) 5 Herb. Hort. Sprengel s.n. (B); Herb. A. ¥.
Roth s.n, [hort. Bremen] (B); Her Herb. Schwigrichen sen. (Mu--142h).
Ttaly: Herb. Prager 18668 (Gg-—3196)) ; Turra s.n. [Vicenza] (S).
Java: Herb, Hort. Bot. Bogor. XV.JA.XXIX.5 (Bz--2636)). Kentucky:
Short s.n. [Aug. 1835] (Ky--1773). Missouri: Herb. Ames s.n.
[Saint Louis] (Oa). North Carolina: W. C. Coker s.n. a

October 21, 1944) (Hi-~27h72); V. Matthews s.n. (Arboretum, July
a 1934) 1 (8) Radford & Stewart S.56 (Hi--3)969), S 3.103 3 (—
34982); Smith & Stewart S 0103 ave Oklahoma: R. E. Miller 52M.

[Stratton isloT (St—-261)8). Spain: Collector undesignated ae
(Q), 21 (Q). Texas: C, M. Parker 792 (Au); Studhalter 193 (Au).
LOCALITY OF COLLECTION UNDETERMINED: Banks s.n. (U.S.A.] (8);
Bigelow s.n. [Camp no. 6, July 30, '53] (W—-56132); Collector un-
designated 2 (Au), s.n. [probably Wichita, Kans.] (Io—108282),
s.n. (Au, Au, De, De, De De, Du-~166514, Mu-~1685, Pr); Curtiss s.n-
(Southern States, 1875] (Ka, Ky); Dean s.n. (E=-119296); Duf Dufour
Sen. (Mu--9)6); Preder sen. [Caroli ina] (De); Herb. Bernhardi 8. Boe
(E--119242); Herb. Buckley s.n. sen. (E--119301); Herb. Cha Batis’
[Southern Flora] (Ms); Herb. Ho Hort. Bella s.n. (B); Herb. Jesup S- sup Ss.
n, [Southern States] (Dt); Herb. L. Lamarck sen. (P); Herb. Link s.
n. (B, N—photo, S—-photo, Z—-photo); Herb. Riehl s.n. (E--119315)5
Herb. Schimper s.n. (Mu—125); Herb. Schmiedelian s.n. (Mu-——9l3)5
Herb. Schreber s.n. (Mu--94, Mu--9)5); Herb. Zuccarini s.n. (Mu—
1686); Joor s.n. (E--119188); Leconte s.n. (Amer. sept.) (P)j
Lindley s.n. [U. States] (P); L. H. Pammel s.n. (seedlings, March
1927] (Io—~126838); Plée 365 (P); Tharp s.n. (Au). MOUNTED ILLUS-
TRATIONS: Anon., color plate (N); Buchhoz, Herb. Color. Amer. pl.
se (Gb) 5 Buchoz, Hist. Nat. Reg. Vég. 6, dec. 10, pl. 5.

CALLICARPA AMERICANA var. LACTEA - Muller, Am. Gard. 10: 463
fas "Cc, americana lactea"). 1889; Moldenke in Fedde, Repert.
Poe . Nov.

dditional & pide sy synonymy: Callicarpa lactea F. J. Muller,

Am. renry 10: 463. 1889. Callicarpa americana var. alba Hort. ex

Rehd, in L. H, Bailey, Stand, Cycl. Hort. 2: 628. 191. Callicarpa

americana var. alba Clute, Amer, Bot. 31: 29. 1925. Callicarpa

1966 Moldenke, Monograph of Callicarpa 495

americana var. alba Rehd., Man. Cult. Trees, ed. 1, 776. 1927.
Callicarpa americana var. alba Berckmanns ex Moldenke in Fedde,
Repert. Spec. Nov. 39: ont tag in syn. 1936; piaber ore Alph. list In-
Callic

valid Names 9, in syn. 1940. carpa americana f, leucocarpa
Standl. ex Moldenke, Suppl. List Invalid ae 1, in syn. 19,1.
Callicarpa americana var. albocarpon Fox ex Moldenke, Alph. List

Invalid Names Suppl. 1: 3, in syn. 1947. Callic americana f.
lactes (F. J. Muller) Rehd., Bibl. Cult. Trees Le 1949. Calli-
americana var. leucocarpa Standl. ex Moldenke, Résumé 2i1,

Inv.
= 1940; Moldenke, Suppl. List Invalid Names 1. 1941; Moldenke,
own Geogr. Distrib. Verbenac., [ed. 1], 4, 5, 12, 71, & 86.
19123 Moldenke, Alph. List Invalid Names "Sg & 9. 1942; Moldenke in
ell, Fl. Texa s 3 is if & 79-——80. 1942; Moldenke, Known Geogr.
istrib. V Mac « : 1. 1943; Moldenke, Phytologia 2: 73,
88, & 93 (95) = os ss. "196; Moldenke, Alph. List Cit. 1: 19,
» 37, 158, 175, 202, 203, 209, 305, & 309. 1946; Moldenke, Phy-
tologia 2: 327. Poe "woLdenke, Alph. List Invalid Names Suppl. 1:
Soc.

sman, Handb 9
12, lb, 22, 23, 27, 213, 2h, 2hh, & Lh3. 19595 sya Man, Cult.
al, & 932. 13605 Yoldenke, Résumé Suppl. 3:
Morris Arb. 1h: Be

1963; Moldenke, R - 14: 1&6.1
t collectors describe this plant as a shrub, 1m. tall,
fl white, and the » whi snowy~

and fruits are rea His var. alba is based on an unnumbered col-
lection made by H. A. Rankin in Columbus County, North Carolina.
Matthews also avers Aunt the flowers are "pure-white" as rene as
the fruit. Earle (1902) says "a form with white fruit occurs" in
Adal A note on W, C. Coker sn. |Arboretun, October 21, “19bh]

496 PHYTOLOGIA Vol. 13, no. 7

makes the following amazing claim: "White-berried form and purple-

berried form from same root" — the specimen consists of just two

branchlet tips, the fruits on both of which have dried identical-
e

ly as those of the normal form. Ewan notes "growing with th
usual red-purple shrubs" in Evangeline Parish, Louisiana.

Callicarpa americana alba Berc F. S.
Earle s.n., collected at Auburn, Lee County, Alabama, on September
25, 1897, and deposited in the herbarium of Kansas State College;
var. albocarpon Fox is based on L. E. Fox 2226 from Natchitoches
Parish, Louisiana, deposited in the Fox herbarium; and f. leuco-
carpa is based on G. L. Fisher 5.n., collected at Anahuac, Cham-
bers County, Texas, on September 18, 1931, and deposited in the
herbarium of the Chicago Natural History Museum.

variety was collected 5 miles south of Silsbee, Hardin

P.
on September 30, 1926, by S. N. Crisp; and in the Glen Saint Mary
Nursery, Baker County, Florida, on August 20, 1935, by L. H. Bai-
ley and by H. H. Hume.

Dr. - Wherry, in a letter to me dated January 9, 1937,
states that in about 1920 he visited the Berckmanns place in Au-
gust, Georgia, and that the widow of the founder pointed out C.
americana var. lactea as one of the most interesting plants there.
On September 11, 1936, he collected the variety about 3 miles
northwest of Robert Post Office, which is just east of Hammond ,
Tangipahoa Parish, Louisiana. It was growing with the typical
form of the species, although the bush of albino nature was obvi-
ously weaker, lower, and paler,

Collectors have found the plant growing in old fields and on

dry hills, at 550 feet altitude, flowe in July, and fruiting
from August to October. Bailey (1935) states that it is offered
to the horticultural trade Sanford. Common names recor

15 herbarium specimens, including type material of most of the
names involved, and 6 mounted photographs have been examined by me.
Additional & emended citations: NORTH CAROLINA: Orange Co.: Le
Clair s.n, (Mason Farm, July 15, 1937] (Hi—6300), s.n. [Mason
Farm, July 1937] (Hi—8985). Pitt Co.: Crisp s.n. (Greenville,
Sept. 30, 1926] (Hi—59407). SOUTH CAROLINA: Beaufort Co. Melli-
champ s.n. [Bluffton] (E—119216). FLORIDA: Baker Co.: H. H, Hume
Sn. [Glen St. Mary, 20 aug. 1935] (Fl~209%). Columbia Co.!
Rolfs 183 (F—228793), 18h (E~—119151, F—22879, Fl1—209L5, Ur).

1966 Moldenke, Monograph of Callicarpa 497

a sey Co.: McFarlin 6468 (Mi), 9746 (N). ALABAMA: Lee Co.:

- S. Earle s.n. [Auburn, Nov. 1878] (Gm) » Son. [Auburn, 9-25-
a (Ka), sen. {Auburn, Nov. 1898] (F—-170730, W—3528). AR-
KANSAS: Hot Spring Co.: Demaree 1653 (N, Ok, Se—55743). LOUIS-
IANA: Evangeline Par.: Ewan wan 19360 (Tl). tins oln Par.: Cocks s.n.
[Rustin, Sept. 1914] (Tl). Natchitoches Par.: L. E. Fox 2226
(Fx). ‘EXAS: Chambers Co.: G. L. Fisher 6 (F—67696), sn. [A-

namac, Sept. 18, 1931] (Du-~283952, Ew, Wi). Hardin Co.: Parks &
Cory 19988 (Tr), "19989 (Tr). Jasper Co.: C. L. Lundell 11890 (Sm).
San Jacinto Co.: Parks & Cory 19650 (Tr). Upshur ne = J. Pal-
mer 31738 (E—956795). CULTIVATED: Florida: L. H. Bailey s.n.
{Glen st. Mary Nursery, Aug. 20, 1935] (Ba). North ere v.
Matthews s.n. [July 11, 193)] (Ei—-59)12) . MOUNTED CLIPPINGS:
Clute, Amer. Bot. 31: 29. 1925 (Up).

CALLICARPA AMPLA Schau,

Synonymy: Catlicarpa ampla Schau. ex Moldenke in Fedde, Repert.
Spec. Nov. 2: 22, sphalm. 1937.

Bibliography: Schau. in A. DC. e lls 642. 1873 —

ok. f. & Jacks., Ind. Kew. 11 3h6. 1 3893} Brig. in Engl. & Prantl,
Nat. Pegs lls (3a): 166. gett pe Urb. in Fedde, Repe 4: Spec.

oh . + Gomion Vern ee Om Moldenke, Geogr.
Distrib, ony 7&8. 19393 oldenks Suppl. list Common Vern.
Names 5, 190; Moldenke, Phytologia 1: 497. 191; Moldenke, Know
Geogr. Distrib Terbenas. =~ 27 & 86. 19h2; ‘ectaarke, Phyt o-

69, Alph.
117, 179, & 301 (a), ah 47, 403, h20, 436, 528, 55h, & 6ho
(1918), 3: 695, 822, 853, 85h 856, 26, 9329 938, & 958 (2919),
and : 1026, 1058, isho; Known Geogr. Distri
Ms pee ed, 2, 5. 4 e et % te; Moidenke, Résumé 58, 59, "260,
&
Recent collectors describe this plant as a small tree, 15 feet

Ww v
lar name is "capa rosa". Jackson (1893) ascribes the species to
Jamaica in error. Urban (1922) compares C. oblanceolata Urb. with
C. ampla as follows: "Ex antillanis maxime accedit ad C. amplam
Schauer (e Portorico), quae foliis ovatis v. ellipticis, margine
supero crenatis v. crenulatis, subtus ji paca a ap mailto
tenuioribus, fructibus minoribus facile discernenda est

Material” has been misidentified ~ er in herbaria un-
der the name Egiphylla macrophylla

In all, 60 herbarium specimens, rotten type material of all
the names * involved, and 21 mounted photographs have been examined

498 PHYTOLOGIA Vol. 13, no. 7

"= Additional & emended citations: PUERTO RICO: Baudin 208 [Herb.
Jussieu 5071] (P); Eggers 1297 (W—132325\); Holdridge 2b5 (MN);

Ledru 148 (P), sen. [Herb. Jussieu 5071] (P); Sintenis Sintents Bes S (i
1323265), 2165 (Lu, Mu--3839, W—132326h), 4530 (Be, L

403410), 4883 (E—~119132, F-79945, Lu » Mu--3768, “AB tydler r 283

ph
photo of Aaiaiday N—-photo of ny N--photo of isotype, Ada
of type, S—photo of Pores dee Z--photo of type) « VIRGIN ISLANDS:
St. Thomas: Riédié 2 (P), Sn. (P). LOCALITY OF COLLECTION UN-
DETERMINED: Baudin Sen. (P); Pl Plée s.n, (P, P)3 Riédlé s.n. (P).

CALLICARPA apiett Schau. in A. DC., Prodr. 11: 642. 187.

Synonymy: Call icarpa angusta var. typica H. J. Lam, Verbenac.
Malay. Arch. 66. 1919. Callicarpa acuta Schau. ex P. Dop, Bull.
Soc. Hist. Nat. Toulouse 6): 509 & 512, in obs. 1932. Callicarpa

acuta var. typica H. J. Lam ex P. Dop, Bull. Soc. Hist. Nat. Tou-
louse a 509 & 512, in obs. 1932.

Bibliography: Schau. in = DC., Prodr. 11: 642. 1847; Miq.
Ind. Bat. 2: 889. 1 1856; Vidal, Revis. Pl. Vasc. Filip. 208. Bees

e Acad. Sci. git econ (M61. Biol.] 12: 506. 1886;

Jacks. in Hook. f. & Jacks., Ind. Kew, 1: 386. 1893; E. D. Merre,
Philip. Bur. Forest. Bull. 1: 51. 190 King & Gamble, Journ. Roy.
Asiat. Soc. Bengal 7h (2), extra no., “hos. 1908; King & Gamble,

Mi 5.
Sci. Bot. 7: 312 (1912) and 1a 299 & 301. 1917; H. J. Lam, Ver-
pes pa Malay. Arch. 8, 58, & 67. 19195 ges Bull, Jard. Bot.
Buitenz., ser. =? 3: 10% don 1921; E. Ds Enum, ois Pl.
3: 382. 1923; P. Dop, Bull. Soc. Hist. Nat. eeisnas. Bh: 509 &
512. 1932; Woldenke, Alph. List Common Vern. Names 10. 1939; Mol-
denke, Prelim. Alph. List Invalid Names a 1940; Moldenke, Known
Geogr. Distrib. Verbenac +, ed. 1, 61 & 86, 19425 ; Moldenke, Alph.
List Invalid Names 8, 1942; Moldenke, Phytologia 2: 93. 1945;
Moldenke, Alph. List Cit. 1: 5 (196) and 2: 163 & h70. 19h, Mol-
denke, Known Geogr. Distrib. Verbenac.. ed. 2, 1,0 & 176. 19193
reps Résumé 182, 241, & 43. 1959; Moldenke, Phytologia 13:
ésumé S 5. 1966

Praha about 2 m. tall; stems about ee in abel pot

tomentose, scarcely many-flowered; peduncles as long as or almost
as long as the petioles; flowers small; calyx 2 mm. long, the rim

1966 Moldenke, Monograph of Callicarpa 499

obscurely -toothed or very minutely Piggy 7 relay Pe
twice as long as the calyx, ) mm. long, tomentose
outside; stamens and style long-exserted; mature err weit, ;
ess than 2 mm, in diameter, not depressed at the apex, with
poaktoped stellate te hairs.

The type of this species was collected by Hugh Cuming (no.

forests, on dry soil, in open forests, and al

forest veo hare » at altitudes of 1000 to 2000 feet, flowering in
October, fruiting in February, Marc h, and October.

The ener are described as "white" on Herb. Philip. Bur. Sci.

46685, while it is the fruits that are said to be white on Herb.

Philip. Bur. Sci. 467h1 & 46829. A vernacular name recorded for

the species is "dirik-dirik".

Maximowicz (1886) compares his C. pilosissima with C. angusta
as follows: "Similis Callicarpa angustae Schauer (Cuming! n. 1425)

e Philippinis, quae tamen differt % pube, foliis subtus incanis
basi cuneatis, corolla extus tomentosa; nec non Callicarpa cauda-
tae m., cujus " ddagnosim inserere liceat...."

Lam (1919) cites a "Com. d. 1. Fl. for. d. Fil. no. 25" from
Luzon as the typical form of C, ta. By this curious citation
he probably means 1425, the type collection. His var.
longifolia, said to © differ in its larger leaves, 9--18 cm. long,
1.7--l cm, wide, the petioles 8--18 mm. long, and the venation a-
bundant but obscure on both leaf-surfaces , is C. rivularis vc
He also cites under C. blancoi Rolfe [=C. form osana Rolfe], wi
@ question, a Teijsmann s.n. 1 s.n. (Herb. Lugd.—Bat. 508 2265-307} ace
termined as C, angusta by Ha Hasskarl.

Material of Cc. angu angusta has been misidentified and distributed
in herbaria under the names C. bicolor Juss., C. cana L., and C.
lo songifolia Lam. On the other hand, the Loher 12589, « Shatributed

+ angusta, is actually Cc. longipetiolata var. glabrescens ens Mol=-
asia “Loher 12312 and Ramos . & Edaflo s.n. (Herb. Philip. Bur. Sci.
48530] are C. subintegra Merr.; and Ramos & Edafio s.n. [Herb.
aan Bur. Sci. 33636] is Geunsia ramosi “Moldenke.

23 herbarium specimens, mens, including type material of all
tg se * involved, and 1 mounted photograph, have been examined
me.
Citations: WESTERN PACIFIC ISLANDS: PHILIPPINE ISLANDS: Luz
Ahern's collector kee a sen, [Herb. Philip. Forest. Bur. 1888] (Bz—

172h7, | N, Po—6811); Cuming 1h25 (N--isotype, Z—photo of isotype);
M, Ramos "1958 (Bz—17273), sn. n. [Herb. Philip. Bur. Sci. 1030]
(Bz—17248, N N); Ramos & Edatio s s.n. (Herb. Philip. Bur. Sci. 26367]

(Bz—17250), s.n. (Herb. P hilip. Bur. Sci. 34051] (N), s ae (Herb.
Philip. Bur. Sei. 16685] (Ca--309226, N), s.n. (Herb. Philip. Bur.

500 Fear2cOLbogctls Vol. 13, no. 7

Sei. 46741] (Ca--309183, N), sen. (Herb. Philip. Bur. Sei. 6829]
(Ca—309331). Mindoro: Merritt 5 s.n. (Herb, Philip. Forest. Bur.
8788] (N); M. Ramos s.n. [Herb. Philip. Bur. Sci. 39455] (Bz—
172h9, N), son. [Herb. Philip. Bur. Sci. 39497) (Bz—-172h5, Bz—
1726). Palawan: Foxwo xworthy s.n. (Herb. Philip. Bur. Sci. ]
(Bz—17595, N), s.n. [Herb. Philip. Bur. Sci. 719] (Bz—-1759h, N).

ent > egecatagian King & Gamble, Kew Bull. Misc. Inf. 1908:
pk

Bibliography: King & Journ. Roy. Asiat. Soc. Bengal
7h (2), ¢ extra no., 802 a ag 1908; King & Gamble, Kew Bull.
Misc, - 1908: 106. 1908; King & dasble, at Fl. Malay. Penins.
21: jot “1015. 1909; Prain, Ind. Kew. Suppl. 4: 34. 1913; H. J.
Lam, Verbenac. Malay. Arch. "YB & & 66-67. 1 19195 Bakh., Bull. Jard.
Bot. Buitenz., ser. 3, 3: 10 & 19. hat . N, Ridl., Fl. Mal.
Penins. 2: 61h & 616, 1923; Calder, N
Rec. Bot. Surv. Ind. 11: 2h. 1926; P. Dop, Bull. Soc. Hist. Nat.
— - 502, 503, 511, & 512. 1932; loldenke, Known Geogr.

Distrib. Verbenac., “ = 59, 60, 63, 66, & 86 (1942) and ed. 2,
137, 138, 143, 147, S Sakesdiciseice Phytologia 3: 2
(1950) and h: 122 & 123. “ty Moldenke, Résumé 177, 179, 187,
196, & 4h3. 1959.

Low, slender, twiggy — to 4 m. tall; branchlets obtusely
or obscurely quadrangular, rayish or Serviginois with an appres-
sed tomentum = stellate had} leaves decussate-opposite, their

silvery, or tawny pubescence; midrib stout, impressed above, prom
ent beneath; secondaries slender, 9--15 per side, eleva
more or less irregular, issuing at 60--70° from the midrib and at

long, stellate-pubescent outside, the obese erect, rounded; sta-
mens inserted at or very near the base of the corolla-tube, hardly
exserted; filaments | mm Gre slender, scarcely or not at all
exserted; anthers oblong, 1.5 mm. long, much glandular-dotted on

1966 Moldenke, Monograph of Callicarpa 501

both sides; style slender, exserted; stigma age ovary vil-
lous; fruit drupaceous, agar te -5 mm. in diameter, de-
Pones I at the apex, black or dark-blue when "ripe, glabrous; py-

pecies was based on a collection of Henry Nicholas Rid-
ley Gee 8330) from Lankawi Island, one of Charles Curtis (no.
3197) fr engl bag ae one of Benedetto ep mem, ee 1596)
Bat » and two made by Ulu Buboug es ; collec
tor nos. "7036 & a 8036) from the top of limestone ve 200-=300
meters rs altitude, in Perak, Malaya. Ridley 92?) ae a Ridley
specimen from Selangor r and a Curtis collection a ge Island.
ios species has been found growing on docrlvibig limes
limestone cliffs, and the cleared tops of limestone hills,
at “506 & to 1300 feet altitude, flowering in October and November,
fruiting in February. The flowers are described as "violet"
x ge tndrenic ty 1906 and as “white” on Gusdorf 96. The leaves on
Nur 34,369 are especially small. King & Gamble (1908) note that
"The Kedah specimens [by which they mean those from Lankawi Is-
land] have narrower leaves than the rest, with more slender
branchlets. The species comes near to Cc. angusta, Schauer, from
the Philippines, but differs in the almost quite entire leaves
and larger flowers and fruit." Lam (1919) says "The ea has
close affinity with C. angusta var. @ [=C. rivularis M rr.) and
perhaps it will appear that it is synonimous with it. ope it
seems to differ from it by the pubescent corolla and the hardly

» 21 herbarium specimens have been examined by
Citations: THAILAND: Khantchai 1087 [Herb. Roy. cara Dept.
24217) (Z). MALAYA: Kelantan: M. R. R. Henderson 19597 (Bz—-17260,
Ca—-32713); Nur & Foxworthy 1211) (Bz—-17259, Ca——346272). Pa-
hang: M. R. Henderson 22265 (Bz--17258, N), 25049 (N). Perak: M.
R. Henderson 23754 (B2—-17261) , 23805 (N).- Selangor: Nur 34369
(Ca--3100, S)> MALAYAN ISLANDS: Langkawi: M. R. Henderson 29152
Aa ean “INDONESIA: GREATER SUNDA ISLANDS: Banka: Btnnemeijer
1906 (Bz—-17252, Bz--17253); Teijsmann s.n. (Bz=-17254, Bz--17255,
Bz—-17256, Ca—265958, Ut—-53171).

aay ipa dl.
Bibli Moldenke, Known Geogr. Distrib. Verbenac., ed. 1
ee & 86 Coie) ir ed. 2, "15 & 176. 1949; Moldenke, Résumé 192 e

It ota: not seem that this binomial was ever validly published
with a Sencar of any sort. This waetar 7 said to be endem-
ic to Sarawak, but whatever the original source of this information

auk ek I published it in 192 has since been lost

aac ARBOREA Ro pee 1814; Fl. cam
ed. 1 [¢ sarge gti wosos vabzo not C. arborea
1940, nor Miq., 1885, nor Wall., 1

502 PATS OLOGIA Vol. 13, no. 7

Additional & emended synonymy: icarpa villosa Roxb., Hort.
Beng. [10], hyponym. 161 [not C. ee Baldw., dw., 1936, nor Vahl,
1794]. Callicarpa tectonaefolia Wall., Numer. List 50, no. 1827.
1829. Callicarpa arborea var. villosa (Roxb.) King & Gamble,
Journ, Asiat. Soc. Bengal 7 (2), extra no., 803 [as "var. villosa
Roxb.]. 1908. Callicarpa lanata Hosséus apud Craib, Contrib. Fl.
Siam Dicot. 163, in syn. 1912 [not C. lanata Geable. 1878, nor L.,
1771, nor H. J. Lam, 1940, nor Lam., , 1821, nor orp 1893, nor
Vahl, 1856, nor Wali. , 19h0, nor Zipp., 1641). Calli arborea
var. villosa King & Gamble ex Craib, Contrib. Fl. saz Dicot. 163.
1912. Calli Callicarpa tectoniolia Wall. ex Bakh., Bull. Jard. Bot.

Buitenz., ser. 3, 3: 21, sphalm. 1921. Callicarpa arborea arborea var.
villosa Gamble ex H. N. Ridl., Fl. Malay. Penins. 2: 615. 1923.
Callicarpa tomentosa Bakh. (in part) apud P'ei, Verbenac. China
21, in syn. 1932 [not C. tomentosa Hook, & Arn., 1918, nor KUnig,
1893, nor L., re nor (L.) Murr., 1774, nor Lam., 1783, nor
Murr., 177), nor Thunb., 1959, nor Vahl, 1791, nor mila., 1808].

carpa cehowia a var. villosa Fletcher ex Moldenke,
Suppl. 1h: 6, in syn. eis celsicarpa tomentosa f. feces:
(Roxb.) Bakh. ex Moldenke, Résumé Suppl. 1h: 7, in syn. 1966.
Callicarpa tomentosa f. ville sa the ) Clarke ex Moldenke, Résu-
mé Suppl. 14: 7, in syn. 1966. Callicarpa tomentosa var. arborea
(Roxb.) Bakh. ex Moldenke, Résumé Suppl. 1): 7, in syn. 1
Callicarpa tomentosa f, villosa (Roxb.) King & Gamble ex Voldenke,
Résumé Suppl. 1): 7, in rae syn. 1966.

Bibliography: Murr. in L., Syst. Veg., ed. 13, 130. 1773 as
Hort. Beng. [10]. 1h Wall. in Roxb. » Fl. Ind., ed. 1 [Carey &

Roxb., Fl. In
Pl. 1s = 1839; “oe & Hérinc neq, Man. Gén, pl Arb. & Arbust.
Prodr. 11:

Koord. & Valet., Bijér. Kenn. Booms. Java 7: 175. 1900; J. J. Wood
Rec. Bot. Surv, Ind. 2: 20. 1902; E. D. Merr., Philip. us Forest.

1966 Moldenke, Monograph of Callicarpa 503

Bull. 1: 51. Be Prain, Beng. Pl. 827. 1903; Strachey, Cat. Pl.
Kumaon 136. 1906; King & Gamble, Journ. Roy. Asiat. Soc. Bengal
7h (2), tg no., 802 & 803. 1908; King & Gamble, Kew Bull. Misc.
Inf. 1908: 106. 1908; King & meray Mat. Fl. Malay. Penins. 21:
802. 1909; Elm., Leafl. Philip. e 3: 866. 1910; + ieee Monga
Bot. Centralbl. 28 (2): 29. oui kab Kew Bull. Misc. Inf. 9:
442. 1911; Craib, Contrib. Fl. Siam Dicot. 163. 1912; E. De Merr.,
Philip. Journ. Sei. Bot. 12: 298. 1917; Heyne, Nutt. Plant. Ned-
erl. Ind. 4: 107. 1917; Basu, Ind. Medic. Pl. 3: 3, pl. 732a

1918; H. J. Lam, Verbenac. Malay. Arch. 80 & 82. 1919; H. oe "Ridl.,
Journ. Fed, Malay States Mus. 10: 110. 1920; Bakh., Bull. J

Bot. Buitenz., ser. 3, 3: 21 & 30. 19213 H. N. Ridl., Fl. inlay.

66. 1931; P'ei, Verbenac. China 1) & 21—22. 1932; P. op,
Soc. Hist. Nat. Toulouse 6): 600, 503—-50h, 511, & 512. 1932;
pig: in Page Pest Spec. Nov. 39: 302 (1936) and 40: 98,
106, ——113, 12h, & 130. 1936; A. W. Hill, Ind . Kew.
sant: is LS. 1938; eoataihe? Geogr. Distrib. Avicenn. 35. il
Moldenke, Suppl. List Common Vern. Names <i dky dee aby IOs
oh, = ete ped, Moldenke, Prelim. Alph. List Invalid Names 9,
ag ee 190; Biswas, Ind. Forest Rec. Bot., new ser., 3: 1.
9 ; Worsdell, Ind. Lond. Suppl. 1: 160. 191; Moldenke, Known
Rie te

10
and l:'1102, 1103, & 1110. ne Moldenke, Phytolo
139. 19h9; Moldenk e, Known Geogr. Distrib. Verbenac., art “i 12 2h,
125, 125, 130, 1%, "137-10, 143, 148, 156, ae 176. 1949; Molden~

213

, Bot. Surv. So. Shan States ly. 1960; N. R
Nat. Bot. ‘aes rt Rass tela
ke, Résumé Suppl. 3: 30 (1962) and 4: 7. 3 Legris, Trav.
Sect. Scient. Inst. Frang. Pond. 6: 213, 50%, 513, & sabe 19633
Moldenke, Dansk Bot. Arkiv 23: 87. 1963; Bhatnagar,
Bot. Soc. 42: 37h. 1963; Neal, In Gard. Hawaii, ed. 2, TT « 1965;
Moldenke, Résumé Suppl. 1): 3, 4, 6, & "te 966 «

Illustrations: Basu, Ind. Medic. Pl. 3: pl. 732a. 1918; H. N.

Ridl., Fl. Malay Penins. 2: 615, fig. 128. 1923; Nair & Rehman
Lucknow 76 2, fig. 62.

Bull. Nat. Bot. Gard. : pl. ap
——5 m. tall, small to medium or even tall
stout, branched, deciduous tree, to 25 m. tall, usually moderate-

50h, PHYTOLOGIA Vol. 13, no. 7

lowish mealy-tomentose; trunk stout, paeely erect, to 6 cm. in
diameter at een height and 85 cm girth, the bole 2 Eee tall;
wood light in weight, rather soft or aievataly hard

ed,
Visible; head round; bark gray or grayish to pale gray-brown or
b h, smooth or rough, longitudinally fissured, tawny-gray on
shoots, the inner bark pal e-brown; branches stout, spreading or
long-strag gling, gg tga pew. “branchlets thic k or stout, ob-
furac 8

tusely tetragonal, densely cinereous- or flavescent-fur
or covered with senee gray-tawny ae F dich een forked stellate
or mealy tomentum on the er parts, less so in age; nodes an-

young
nullate; principal Guteroodas” 3-5—5.8 cm. long; leaf=scars sun-
ken; leaves decussate-opposite; petioles stout or very stout, 2—~
7 cm. long, above, many-furrowed in drying, dens sely
cinereous~furfuraceous or mealy-tomentose; leaf-blades chartace-

ous or coriaceous, firm, wee or wee eee and shiny above,
white beneath, eis , broadly ell tic, or elliptic-ovate to ob-
long, oblong-ovate or aiiditis len sective » ventricose, 11-3)
em. long, : - Wide, conspicuously tapering
both ends or acuminate or c nate to sho
cuminate or bluntish-acuminate at the apex, entire or sometimes
re seit iculate » abruptly acute -acute or speiags
the or evenc centrally decurrent-acute in a rounded or sub-

cordate base, smooth and glabrous or glabrescent (except for the
midrib) above when sya very densely furfuraceous-tomen ous,
eae napa od gray-t mentose, or closely stellate-tomentose be-
with very short, widte, closely appressed tomentum, very
scene yee ge like the branchlets on both surfaces when inma-
ture and unexpanded; midrib stout or very stout, flat above, very
prominent beneath; secondaries slender or stout, 7-1, per side,
issuing at an angle of about 550 from the midrib and gently arcu-
the

mature
e branchlets and the divisions ee the nde eytnag bracts
rous

ous; flowers small, very preertinnd almost seni, ee or
rather offensive-sce ented; pedicels very short; buds lilac, puberv-
see at first on the outside, later glabrous; calyx campanulate
bular, pale~green, ‘Too ¢ mm. long, 1.5——2 mm. wide,
aay or rather densely pulverulent- “~puberulent on the outside,

1966 Moldenke, Monograph of Callicarpa 505

glabrous on the inner surface, the rim truncate and obscurely =
repand-dentic atta or obscurely and mimutely l-toothed; corolla
infundibular, small, varying from blue, light-blue, mauve, or li-
lac to deep~lilac ; liiac-purple, purple, pale-purple, lively
purple, or rose~purple, sometimes violet, pale-violet, red,
red, pink, pale pinkish, rose-pink, old-rose, rose-lavender, or
y ube

even ™ ellow", exserted, its t usually much longer th
“cy. » 1.5-—3 mm. long, slightly widened at the
ape or very t ent-puberulent on the

side, one smaller than the dant; stamens , inserted 0.5—0.7 mm.
ove the base of ot corolla-tube, long-exserted; filaments slen-
der, mauve, 3.5—l.5 mm. long or twice as long as the coro a,
ro y own

ends, the apocolpium 7p in di-

endine te, more or less areolate, the columella

closely packed, most of the g fol ape :
pistil exserted; style capillary, mauve, 4.5——5 mm. long, glab-
rous; st: t capitate or peltate, about 0.5 m. e
later broadly b ovary superior, globose, about 1 m.
long and wide, densely puberulent, 1-cell ed

c patelliform, glabr paceous, small
round or subglobose to globular, succulent, green or greenish
when young, later pinkish, magenta le, vio-

» 2-—6 mm. long and wide, glabrous, smooth or

por ee n drying; seeds ) or less, convex on the mutside, with
2 flat faces on the inner surface, rough, the interior integument

hard like the shell of a nut; perisperm oval ft, daline;
embryo straight, erect, — as long as the perisperm; cotyle-

c erenc Razi
(1946) describes it as a microphanerophyte according to sconces s
classification pd life-forms and records it from Mysore.

rs scribed as "pink" on Brass 526 Carr 11535, M.
S. Clemens 10870, Khantohad 1080, and = ng 23914; "rose-lavender"
bs Koelz 2053h; "lilac" on Btnnemeijer 28; "dull-red" on ae

pee aes 22196, Koelz 22718, Sindhiponse 45, and Yates sa es
"light—blue" on Asdat 2; "blue" on Yates 1078; mauve" on “22E
3905; "rose-purple" on Koelz 30119; "violet" on Bunkird s.n.
Told-roset on Koelz 254,78; “lilac or rose-pink" on on Haniff f 249; and

506 P84 %:0 LOG27.4 Vol. 13, now 7

"yellow" on Khantchai 350.

The species has been found growing in forests » marshy or decid~
uous forests, dipterocarp or mixed evergreen forests, ri
ns

ore open open y places, the ins of mixe
thickets, sec er dry situations, near plantations, at
the base of limestone cliffs, in open woods, by ; k-
ets, and amongst scrub, sea~level to 8000 feet altitude,
and fruiting to October. Roxburgh states

that it "Flowers at various times through the year" at Chittagong.
Tsang refers to it as "scattered shrubs, climber, 9 feet tall" in
Kwangsi. Brass calls it "common in rainforest association"
in Papua; Yates says "in open grassland to second growth forest"
in Sumatra. Kingdon-Ward describes it as "very common in second

shrubs in deciduous and in mature forests on hills", In Kwangsi
— described by Ching as "a fairly common tree on cutover hill-
sides",

Watt (1889) says "Met with in Kumaon, Oudh, eastern Bengal,
and Burma; chiefly in second-growth forests". He also reports
that the bark is aromatic and bitter, and is applied in decoction
in the treatment of cutaneous diseases. He quotes a surgeon, W.

requ deserted hill to
elevation." Balfour (1885) calls it "a stout, tall tree of Nepal,
ss onte, the Morung mountains, Goalpara, Chittagong, and

"India, Pakistan, Nepal, southern China, Indo-china, and Thail

to Malaya, the Philippines , Sumatra, Mantawi Islands, and New

Guinea." I have, however, seen no material of the typical form

pl aa apne: ste the Philippines and feel now that all records
8 from th oc

i. wee e Philippines refer to var. psilocalyx

Bhatnagar (1963) calls C. arborea a "rare evergreen tree"
ms other writers refer to it as deciduous] in the Hollong Nahor

tic formation Occupying flat land and low broken hills with lux-
uriant vegetation. Ridley (1923) avers that it grows in "Woods
ow » no e" in Malaya. He cites a Cantley collec-

from Negri Sembilan, a Wry collection from Perak, and a Cur
tis collection from Penang. He records it also from Kelantan.

| LIBRARIANS -- Please paste this over the top portion of Vol. 13, page 507

——

Wurdack, Jede 5

1966 Index

Index to Authors in Volume Thirteen
Allen, C, K., 221, 232
DeFilipps, R. A., hh
Degener, 0.,
Degener, 0., & Degener, I., 49
Dugand, A., 379
» Ae, & Johnston, M. C., 377

» 207, = 371
Moldenke, A, La, 38, 81, s2

507

72, Lk6é, 452
Moldenke, . N., 1, 162, 16, 179, 218, 242, 243, 277, 306, 310,

318, 343, hou, Lbs,
Smith, L. B., ee ish”

1966

Index

507

Index to Authors in Volume Thirteen

Allen, C. K., 221, 232
DeFilipps, Re Sig

Degener, 6., & Degener, I., h9
Dugand, A., 379

and, A., & Johnston, M. C., 377

Dug

Eiten, is Tey
Fosberg, F. R.,
Moldenke, A. ae ,
Moldenke

66°
Smith, L. ie as 45k
Wurdack, Je , 65

81, 219, bs,

-— = =—=& -— e = eo

452
* H. Ne, 1, 162, 169, 179, 218, 2h2, 2h3, 277, hOl, bbs,

Index to Supra-specific Scientific Names in Volume Thirteen

Abies, 19, 258
Acacia, 173, 211, 311, 357, 365,

f
é
Acisanthera, BK:
j
]

Aechmea, ry a 147, 148, 161,

Aegiphi Lis 303, 318-343, 401, 427,
bake

Ageiphila, 318
Albizia, 389, 390, 395, 00
Albizzia, 389-392, 395
cantarea, 8)-86, 130, 131, 138
2

wphelenchoides, 202

TE tee F See Pte I tee | be

64
jaceae, 81, 87, 89, 91,
93, 95, 97, 9%, 101, 103, 105,

508 PHYTOLOGIA Vol. 13, no. 7

107 lll, 113, 115, 117, Castelia, 01

ag; A iy, ta, Se or

; i 497

a, us’ ny’ ig 3B agg, Seen, U0,

Pag ga ae ee
Buddieia, 437, 478” Cephalocereus, 3h, 383-385
ulbophyllinae, 308 Cercospora, 427, 476

hyllum, 308, 309 Cereus, 381
Bureardia, 08 Chaetogastra, 65

aes, 408, 425, 427, 439,

0

Bursera, 34, 386, 312, 315, 357,
365, 4hS, Lh6

Capparidaceae,
Caprifoliaceae, 430, 478, 479
Caraguata, 122, 128, 129, 131

Chamaepericlymenum, ),76
Chamaesyce, 385, 00
Chascanum, 0
Cheirodendron, 369
Chilianthus, 29
Chloanthaceae, 1,01
Chloroleucon, 391, 100
Cinchona,

Cc
Cinnamomi
Ciparopets, Bh, Le

27
Citharexilum, 277, 282
Citharexylon, 22, 286, 289,
meet 294, 327
tierenyime, 242, 277-304, 310-

Samatn, "30h, 334, 401, 428

1966

Cremanium,
Croton, vs nee. 386, 387, 400
Cruciferae Pee ff

Grptanthnoeis, 459, 461
3h

sspar ae

Encholirion, 122, 123, 126, 131,

Encholirium, 123, 121, 152, 161
Enterolobium, 392,

Euphorbia, "385, 386
ad 385, 400

Euvriesea
Bugapania,

Index

Fascicularia, 61
2

Forestiera, 1,73
Fouquiera, 286
Fraxinus, 449, 471
Frutex, Frutex, 27, 439, lho
Gamopetalae, 27

Gesneriaceae, 81
Geunsia, 25, 428-130
Glandularia, 192, 193, 21h, 25h,

Gravisia, 153, 161
Greigia, 456, L6h
Grewia, 427

Grisehowia, 68, 69

Guazuma
Gattis, 278
Guamania, 85, 121, 12h, 127-129,

Hadongia, 277, 278, 314
Hechtia, 136
Hedera, 212

Hylocereus, 382

Hyptis, 167, 312

Dex, 428, 476

Tlicaceae, 428, 476
425

Tia,
Inga, 29h, 391

510

Ipomoea, 357, hhé
Jacquinia, 393-400
Jaquinia, 393-395, 400

ae oe: 439, 440, 475
99

duglans, | ; 1

Kingiella, 305
9

tea? ey
&
F
Oo

~

ae

.

27, 29, 32,
367, 168, 110, 171, 173-175,

Larrea, 188, 254, 281, 282
La 453

>
Lauraceae, 221, 223, 225, 227 :
229, on, 232

Lippia, 1, 3-21, 23-38, 162-179,

Li 218, 39-368, Lon 56, us h
qui ar 2 a5:8t
poeansacee, » Wet, 29, 478
Lophocere

Machaericereus, 286
Maclura, L478
Margaritaria,

passanges, a
Maxill

Setaatias et

387-389, 00
131

FertTOLoura
eliola, 427, 476
Melogramma, 1,76

Meriania, 71-73

esembryanthemum, 386
2sua,

iconia, 73-80
imosa, 312, 391, 392

yrtillocactus, 311

ectandra, 221, 22-228, 230

Neovriesia, 8h, 85, 122, 126,

Nestegis, )9

374
Ocotea, goed: 230, 23
Oc Octotoma 3h
Opbioglossacese » 4d1
Ophioglossum, 51, 452

hilus, 306
Orthophytum, 153, 459-h6h

Oxybaphus,
Pachycereus, 286

Paeoniaceae, 37)
e

a rey t

Panicum, 16

apaveraceae, 37)
Paraphalaenopsis, 305
peesneet, 338

Parodianthus, 1,01

Peperomia, 2 , 23, 23h, 239-2h1
Pereskia

Perlarius us, ve

Petitia, 318, 401, 28, 430
Petrea, 01

Phalaenopsis, 305, 306
Phyllanthus, 387, 388
Phytolacca, 76
Phytolaccaceae, 3h, 476

1966

Pilosocereus, 383-385, 00
Pinus, 199, 205, 258, 273, 350,
31, rad es 3%, 471-73
Piper, 233-
Piperaceae, ge
Pisonia, 318
Pisoniaceae, 318
Pisum, 11
Pitcairnia, 127,

» 140,
153, Th, 161, iss, (ist, “Mal
Pithecollobiun, 389 7.
Pithecolobium, 39
Platystachys, "130, 131
Pleome le, 369
Pleurothyrium, 221-228, 230
Polygalaceae, 37

orlieria, on
: orostema, .
: Srp, as ee 26

09
er ee ned 463
Premna, pe”

Prionophy2ivm, see

Priva, 01
Prosp

spodi un » 3h
Pseudo idice, 401

Pseudo-catopsis, 17

Psychotria, 318, 325, h3h
Pterolepis, 65
Pucc

a, 3k
Puya, 12-115, 147, 160, 161,
L6),

wuereus, 10, 205, 312, 349-351,
357, 268, 456, 471, 473
adoehane. "76

Ranunculaceae, 37h

Index

Rauwolfia, Pe 287, 311
Recordia,

Rehdera, oe

enealmia, 121
Resedaceae, 37)
Rhamnaceae, 212, 377
Rhamnidium, 379

Rubiaceae, 274, 278, 476
Rubus, 370, 477
Salvi:

Schlegelia, 336

Schlumbergeria, 128, 131

Scrophulariaceae, 373

Senegalia, 392, 393, 00
7

Stylodon, 4,01
Styphelia, 369

512 PAT TOLOGTA

Subpilocereus, 385

Reon eae eae, 01

ea, 8h, % 110, 120,
121, 12h, 127-1
Theophrastaceae, 393, 4,00
Tibouchina, 65-67, 80, 370
Tillandsia, 8-86, 89-99, 101-
~ 10h, 109, 122-128, 130, 131,
13h, 1%-141 147, "160,
WSh, 455, vikg
408

Vol. 13, no. 7

Vacciniaceae, 176
Vaccinium, 473
Verbena, 179-217, 23-276, 307,

Verbenaceae, 212, h01, 7k
Verbeneae, 26

Vernonia, 318, 325
Viburnum, 430, 43k

Vicia, Vicia, 453

Violaceae, 37h

Vitis, 134, 473
Vriesea, 8, e234, 136-138,
160

Wulffia, 318, 325

Xiphion, 86

Yucca, 21)

Zapania, 3, 8, 29, 34, 166, 362
Zingeber, 369

ber, 369
Zygophyllaceae, 25)

Publication dates of Volume Thirteen

No. 1 — March 7, 1966

ugust 2 Poel
No. 6 — September 19, 1966
No. 7 —— October 14 , 1966

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