Phytologia vol, 72 1965-1966 F PHY TOLOGIA Designed to expedite botanical publication Vol. 12 April, 1965 No. 1 fi = CONTENTS 1 LAUGHLIN, K., Some dendrologic comments on Steyermark’s CAPE OF MESSOMFE OU O0os eis sine cao su Coes be eee meee e es 1 MOLDENKE, H. N., Materials toward a monograph of the genus Et PN Fae aia ie oo eo Ss Ee eee oe ea ees 6 WAS E, Bic hin DOGR VOUIEW og oO 5 ek hn nk eed eH a ee eee 42 Published ey Harold N. Moldenke and Alma : Moldenke 15 Glenbrook Avenue Yonkers : New York, USA. Price of this number, $1; per volume, $5. 75, in n advance, . : or $6 at close of volume of i Volume M, ‘No. A was issued March 09, 1965 SOME DENDROLOGIC COMMENTS ON STEYERMARK'S "FLORA OF MISSOURI" Kendall Laughlin 165 Pine Ave., Chicago, Ill. This magnum opus of 1800 pages, enbellsentg by hundreds of excellent drawings, is the result of thir- ty years of work by Julian Alfred Staveraerk Ph.D. Most of the descriptions do not state whether the leaves are alternate or opposite. This information can apparently be gleaned from the General Key, but only after a long search for the family. A single line beneath the name of the genus "Monoecious--dioe- Okie She ee ee ee --leaves" would have been worth while 530--Fagus grandifolia. Steyermark does not recognize the type species of the Beech in Missouri. There are nevertheless trees in southeastern Missouri with cuneate leaves like the type species. There is Such a native tree 4 feet 9 inches in circumference between the driveway and a side street beside Marvin Lepchenske's home at 124 S. Locust Street, Dexter. 544 Steyermark does not recognize the vari- ety RRB as distinct from Quercus falcata var. pagodaefolia. It seems to me that much of the confu- Sion between these two varieties has come about be- cause Ashe's description of leucophylla lire that the undersurface of the leaves was whitish (according to Deam, Trees of Indiana, p. 163). (I confess that I have had to change some of my ideas as the result of like leucophylla is brownish or light grayish-russet turning brown in age. The leaves of leucophylla have fewer, broader lobes than pagodaefolia. While the two varieties often intergrade, prefer to recognize them as distinct, as Moore has done in his Trees of Arkan- sas (p. 53). Some trees of leucophylla have bark like the White Oak. There are two very typical trees o this variety on the west side of Selma Street 175 feet South of Main Street in Poplar Bluff and near the Southwest corner of the town hall square in Dexter. P, 547. Steyermark does not mention the concen- tric circles around the apex of the acorn-nut of Quer- £us coccinea, which I think is the most distinctive 1 2 PHYTOLOGIA Vol, 12, no. 1 character of the species. Steyermark recognizes Q. coccinea var. tubercula- he had ta, but expresses a little skepticism, as if he late acorn-cups 25-28 mm. wide. There should be no doubt about the distinctiveness of tuberculata in Mis- souri, Illinois, etc. I think that the occurrence of Quercus Nuttallii (p. 551) in Missouri should be investigated. 1 ex- I found it well-nigh impossible to convince Stey- ermark of anything about a hybrid Oak. I gave him a photograph, leaves and acorns from a very good speci- men : o Bs lxxxii, but he blandly states on Pp. 542 that this hy- jab is "known definitely only from Cape Girardeau O- I gave him leaves and acorns from five specimens of Q. Xmutabilis in Miquoria. He shows its occurrence in Mississippi Co. on b. name, See PHYTOLOGIA 6:374-378. : new gee in rahe Park, Kansas City, Jackson O., W a circumference of 5 fee i height of 66 feet, Lone ee neeh AES S n Map 766 on p. 540 the "x" for Quercus XEggle- Stonii should be Spotted in Jackson Co. instead of Bates Co, ince this book was compiled, I have described in S PHYTOLOGIA Q. Xdiscreta (Shumardii X velutina), Q. 1965 Laughlin, Dendrologic comments 3 Xmegaleia (lyrata X macrocarpa) and Q. Xriparia (rubra X eoveercas Schneckii) from Missouri. I recentl - scribed in PHYTOLOGIA 9:488-495 Q. Xcolumnaris (palu- stris X mates toes tree now in Chicago; on p. 547 Steyermark mentions this A Bhd as having been found in Osage and Benton Cos As Steyermark states on p. Er bl Celtis is greatly in need of a revision in Missou The species occidentalis and. laevigata can be distinguished in the field by their bark. The bark of occidentalis is almost entirely covered with excres- cences; the bark of laevigata is lighter colored, gti? F and only partly covered with excrescences. n Miquoria there is a kind of Celtis that is distinctly shrubby and with Cornus Drummondi, Rhus ra- dicans and_Arundinaria gigantea it forms a dense un- derstory. The specimen that I have has leaves more than twice as long as wide reaching a length of 7 cm., rounded and nearly symmetrical at the base, acuminate at the apex, 3-nerved, dull yellow green on both sur- faces, scabrous above, quite thin, definitely toothed above the middle with short narrow teeth smaller than in occidentalis, the petioles, 5-7 mm. long, and the hackberry and it.is certainly on important constituent tion of tenuifolia. While the vegetative characters of my plant fit his general description of laevigata except for its small Size, it does not fit any of his descriptions of the varieties, so that there seems to be doubt about its identity. Pages 796 to 799--PYRUS--MALUS ermark uses the generic name Pyrus for the Apples and Crab Apples. Malus is the name approved in the International Code of Botanical Nomenclature, Pp. 195. Steyermark shows that the anthers of ioensis and coronaria are red. This is not correct according to my observation. The anthers of all specimens of ioen- Sis that I have observed were moderate orange yellow. There are twenty sheets of specimens of ioensis from Missouri in the herbarium of the Chicago Natural His- tory Museum but not a single sheet shows the color of the anthers. This is a very important character that k PHYTOLOGIA Vol. 12, no. 1 has been generally overlooked by most taxonomists. 2 or of the anthers of coronaria is also or- ange yellow. But the color of the anthers of lancifo- lia is deep pink or reddish pink. Steyermark shows that coronaria var. coronaria is found in Clay, Jackson, Saline, Texas, Madison and Butler Counties. is a specimen from Jackson Co. labeled coronaria col- lected by B. F. Bush at Independence in 1899, his No. is inconceivable that anyone making a careful survey of Clay Co. could have listed coronaria without list- ing lancifolia. eel certain that coronaria var. coronaria is not found in Jackson and Clay Cos. Perhaps it is found rarely in uplands in southeastern Missouri. It is an Eastern species, The reader is referred to my monograph on lanci- folia in PHYTOLOGIA 9:108-112. 1 have broadened three of the characters of the Key shown on p. 111 of that article to read as shown below, based on Study of ad- ditional material: Sena CHARACTER}MALUS LANCIFOLIA MALUS CORONARIA 5 Averaging 49% to |Two-thirds to four- 60% as wide as ifths as wide as long. long. ; Moderate yellow |Dark greenish yel- green or olive green. low or dark yellow. Deep pink Orange yellow. dish pink ee or red- 1965 Laughlin, Dendrologic comments 5 Being a splitter on Crataegus, I rejoice in the 50 species that E. J. Palmer recognizes on pp. 2 822, mostly in southern Missouri, tho I know nothing about them. ut in working out this genus for Glea- son and Cronquist's "Manual of Vascular Plants" the same man Seip ape only 21 species worthy of descrip- tions east the Missouri River. This condensed treatment far too crude and inaccurate for the Chi- cago region, ere there are at least 26 species. incorrect. There is typically a deep narrow sinus above the lower lobe, as shown in Gray's Manual and I am glad that Steyermark differentiates Shes lanata (=mexicana) from americana on pp. 856-858. have observed for decades that the fruit of lanata re bluish--perhaps purplish blue--and different from the red fruit of americana. This difference has been ig- nored and misstated in many books. In describing the Pumpkin Ash on p. 1179 Steyer- mark does not mention the length of the calyx on the samaras, which is 6 mm. in Miquoria (when they can be found) , contrasted with 1 mm. for the Green Ash. There seems to be an introgressive hybridization of the Pumpkin Ash into the Green Ash in Miquoria. In 1957 I found a Pumpkin Ash 90 feet tall near the entrance with samaras on one side of the tree like the Pumpkin Ash and on the other side like the Green Ash. Where xenogamy takes place in the Melioides Subsection of Fraxinus, the fruit is not very reliable. MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. I Harold N. Moldenke This is the twenty-ninth in my series of works of monographic nature on the genera of Verbenaceae, Avicenniaceae, Stilbaceae, Chloanthaceae, and Symphoremaceae. Previous genera so treated by me are Acantholippia Griseb., Aegiphila Jacq., Amasonia L. f., Avicennia L., Baillonia Bocq., Bouchea Cham., Casselia Nees & Mart., Castelia Cav., Chascanum E. Mey., Citharexylum B. Juss., Cornutia Plum., Diostea Miers, Dipyrena Hook., Hierobotana Brigq., Parodianthus Troncoso, Petitia Jacq., Petrea Houst. » Priva Adans., Pseudocarpidium Millsp., Recordia Moldenke, Rehdera Moldenke, Rhaphithamnus Miers, Stylodon Raf., Svensonia Moldenke, Tectona L. £., Verbena [Dorst.] L., Vitex Tourn., and the New World and cultivated members of Callicarpa L. Full explanation of the abbreviations employed herein for the names of the 276 herbaria whose material, in whole or in part, was the preparation of these works will be fo in Phy- tologia 5: 154-159 (1955), 6: 242 (1958), 7: 91-~92 (1959), 7: 123-12 (1960), 7: 343 (1961), 8: 95 (1961), and 9: 191 (1963), with the following additions: Bn « Central College, ore, My~ ; Cd = Museo de Cérdoba, Cérdoba, Argentina; Fj = Fred B. 8 barium, Corpus Christi, Texas; Ij = Science Museum, ca on Missouri, Columbia, Missouri; $j = University of Puerto Rico, San Juan, Puerto Rico; and Ww = Rob & Bessie Welder Wildlife Federa- tion, Corpus Christi, Texas. die ge L-, Sp. Pl., ed. 1, 633 (1753), Gen. Pl., ed. 5, * Lippia L. ex A. L. Juss., Gen, Pl., ed. 1, 109. 1789. Lippi Kunth ex Spreng. in L., Syst. Veg., ed. 16, 2: 930. 1825. 7: 2h. 1832. Juss. ex Steud., Nom. Bot., ed. 2, 23 5h, in syn. 18h1 (not Zapania Lam., 1791, nor Scop., 1806]. a Schau. apud R.A. Phil.. Anal. Univ. o~ Stachyum (Schau.) Small, Fl. Southeast. J. S., ed. 1, 1012 & 1337. 1903, Lippia [Houst.] L. ex Robinson & Fern, in A. Gray, New Man. Moldenke, Alph, List Invalid Names Suppl. 1: 10, in syn. 1947. Lippi H.B.K. ex Moldenke, Castanea 13; 116, sphalm. 1948. Lipps L. ex Troncoso, Darwiniana 12; on sphalm. 1961. 1965 Moldenke, Monograph of Lippia 7 . Dict, ed. 5, “tipeid. 17635 Ls, Gen. Pl., ed. 6, 1764; 'P. Mill., Gard. Dict. - 7, 1: Lippia. 1764; Crants, Inst. 1: Sh6. 1766; L., Mant. 1: “bos 3 1767; P. Mill., Gard. Dict., ed. Nov. 1: > “a dc on Bee, Delic. Fl. & Faun. Insub. 1: & 66. 1790; Gil & Xuares, Osserv. pl. 5. 1790; Poir. Tabl. Encycl, Méth. Bot. [Illustr. Gen.] 1: 58--59, pl. 17. 171, Ae he Juss., Gen. Pl. » ed. 2, 122. 17913 Le Ce Rich., ct. Soc. Hist. Nat. Paris 1: 105. 1792; Cav., Icon. 2: 75, pl. 19h. 17933 Willd., ba ger 1: 116--117. 1797; Curtis, Bot. Mag. ll: pl. %7. 1797 33 - in La, ° Méth. Bot. [Illustr. Gen.] 5S & 9 pl. 539, fig. NE & 2. 17973 Jacq., Hort. eae 59, pl. re 1798 a P e 3 7 & 22, pl. 32, US Nat, Hist. Paris 7: 10--75. 1806; Pers. Syn. Pl. 2; 139--110. 1806; Raf., Med. Repos. NY 5: 352. 1808; Poir. 39 1 « LO es Bot. Cult., ed. 2, 7: = 12 ° ° Sept. 2: 417-18. 181); R. Br, in Sal e 1614; H.B.K., Nov. Gen. & Sp. Pl., ed. fol. pict., 216. 1817; J. Sm, ry Cycl. 3: 3& hk. opel ae Nov. Gen. & Sp. 1818 + quart., 2: 260—~269. a) ed, 1, 233, 303, & "898. is, “Kunth, Syn. Pl. 2: 52th. 1 1823; Nees, Nov. Act. Acad. Caes. Leopold.-Carol. Nat. Cur. 11 Be » 1823 235 Than Fl. Cap., ed. Schult., 7. 1823; Choisy, - Nat. omer. 2: 98. 1823; Mart., — he Med. 103--10h, "Bos A re St.-Hil., Plant. Usuel. Bras. pl. pp. 1-3. 182-1828; Spreng. in L., Syst. Veg., ed. 16, 2: aie 763, = ee 8 PHYTOLOGIA Vol. 12, no. 1 — 1825 Act, Acad. ee eee -~Carol. Nat. ys 39 eichenb., —_ con. Bot. Exot, ; pl. 169. 1830; Hook., Bot. Misc. 23 2%, 1831; Cham., Linnaea 7: 213--2)3 & 375-377, pl. 7, fig. C & D. 1832; Scrak in Mtssler, Handb, Gewichsk., ed. 3, 1: lxxv. 1833; Schrad., Ind. Sem. Hort. Gvtting. 183: nep. 18345 He Be » © - +t OW » Gard. ser, a 35 & 103. 1836; Hook. & Arn., Bot. Beech. Voy « bis2. “peg Bojer, Hort, og 25h. 18373 uM Cag 4 we 2, Sod 2: 59 Gen. .e a: 633." 1838; in Sweet, Brit. Flow pat . ° 3 $ » 208, (1840) and 2: 51, 5h, 750, 751, & 797. 3 Hochst., poe 2h, ° ba Span., Linnaes 15: 330. 18h13 Otto & Dietr., Gartenseit. 9: 379, ish; SHY Useful & goons Pl. In- dia 31. 1841; Scheele, Linnaea 17: 351, 184,33 D » Syn. Pl. 3: « 18),3; Benth., 9 Hartw, 122. tah, Walp., Nov. Act. Acad. Caes, fees e-Carol, Nat. Cur, 19, Suppl. 1: 377. Comment. 242, 18, Bull. seats Sci. Brux. 11 (2): 319 & 326~-327. 1844; Kunth & Bouché, Ind. Sem. Hort. Berol. 185: 10, 1853; F. Krauss, Flora 28: 68. 1845; Walp., Repert. Bot. Syst. hi: 33, 41-57, 6h, & 13h. 18)5; G. G Gardn. in pba Lond, os Bot, he 133-13). 1845; Presl, Wseobecny Rostlinopsis 2: 1203. 18h6; 2ucc., Del. Sem, Hort. Monac whe 186; Benth., Bot. Voy. Sul 153--15h,. 1846; Benth., Pl. Hartw, 25-216, 186; Schm,. in A. DC., Prodr, 11: he & ‘572-594. 1847; Kunth & Bouché, Ind, Sem. Hort. Berol, aps 12. 18473 Clos, ann. Sei. Nat., ser. 3, 10: 378— Por 18185 : Ce Gay, ok Fis. Chile Bot, 5: 27--28, 1895 Lindl., 1965 Moldenke, Monograph of Lippia 9 Surv. 2: 126—127. 18583 Griseb., Fl. Brit. W. Ind. 495. 1861 Bocq., Adansonia 2: 89, 109, 112 Gros 1864; Vilm., Fleurs Pl. Terre, ed. 1, 53 Re Ae Phil., Anal, Univ. Chile 27: 350. », Cat. Pl. Cub. 215~ "1866; ‘ ent. Therap. o 38. 1 Phil., Anal. Univ. Chile 35: 192-193 (1870) “and 36: rs i i870; Re A. Phil., Sert. Mendos, Alt. 34. 1870; Delchev., Rev. Hortic. . ennamen, ed. 1, 128. 1872; Gibert, Enum. Pl. Montev. hh. 1873; Griseb., bhand, 2 * 187k; Gri- Abhand, Kaiser. G Wiss. GUtting. 19: 22-2) seb., Pl. Lorentz. 19h-~196. 1874; D. Oliv., Trans. * ° « Bot. 293 132. 18 a Ulric. , I ° » ed. 2, 128, 18753 Benth. in Benth. & Hook, f., Gen. Pl. 2 (2)s 1133 & hnie—3. 18763 rage Fl. des Pigg _ Eis 3 J. G. Baker, Fl. Maurit. h. 252. noid See » Vid~ pe Meddel. Kjgbenh. 1877-1878: "36-100. tS Loren’ Nei = Nordeste Prov. Entre Rios, ed. 1, 15, 19, 87, 122, 150, 16 : ’ & App. iv. 1878; A. Gray, Syn. Fi. N- Am. 2 (1): 333 & 338-2339. 1878; Griseb., Abhand. Kaiser. Gesell. Wiss. GUtting. ° Fl. Arg ent.] 271219 18793 Boiss., Fl. Orient. hi: 530-533. 1879; 6 Ste Lag., Ann e Soc. Bot. Lyon 73 129. 1880; Hieron., . Acad c Dict. Eng. sei Pl. is8. = iB8hs A. We “Chapm., Fl. South. U. S., ed, 25 pre 4, 308 & 68). 188); "wots, Verh. gg Boog ricer . Linn, 21: 230. 188); Je Ge Baker, J . lam, ie: Lond. Bot, ai “ishs 1885; A. W. Chapm., Fl. South. U. 1887; Moloney, Sketch Forest. W. Afr foi : "583: ier 5874 Lillo, Fl. Tucuman 9). 1888; S. Wats., 89; A. We c +, Bak th. U. *s ed, , a 308 Z we "1889; Rolfe in Oates, — Land, ed. 2, 407. P1889; c ser. 2, 2: 196 (1889) and 3: 163. 1891; Baill., eee 11: 81. 1891; R. Ae P °9 e e oie Bs - 3. an Morong, Britton, & Vail, 3 nn N. Y. Acad. 6-197. 1892; Landb. Journ. Uitg. « Landb, puptaecis: 1892; = M. Wood, Ann. Rep. Col. Herb. $8025 5 Rio de la Loza, Farmacia 10 PHYTOLOGIA Vol. 12, no. 1 ed, 2 atelier’ 189k; Robinson & Greenn » Proc. An. ag 292 31. 189h3 Briq. in Engl. d Prantl, Nat. Pflansenfan, , (3a): 150--152. 18953 Jacks. in Hook, f & Jacks., Ind. Kew. 2: 28 ttre Gr 35 ° 3 — 3B. 1895; S. Moo: P ° Soc, er, 2, 435-37. 1895; H. H. Rusby, M eM» Torrey Bot. Club hs 3-2 18953 Rob e, Am, Journ, Sci. 150 [ser. 3, 50]: 1895; L. H. Bailey in A, Paiva a * & ° ° ws & 513. e Phil., Anal, 62h, pl. 1, fig. anne (i896). ha od sber, 2 (1): k 97. 1902; Bettfreund Fl. Are gent. 3: pl. 156. 1902; Fiori & on. e 360. 19023 Chod., Pl. ler. 1 a 198-200. 1902; Chod., Bull. Herb. Boiss., sér. 2, 2: 81 ol ° Bot. Club 29: 597. 1902; Wi ambes + 350. 1903; Greem., Proc cad. Sci, 1903; J. K. Small, Fl. Southeast. U. S., ed. 1, 1007, 1012, & 1337 1903; » Trans. - Acad. Sci. 18 « 1903; acks., Ind. Kew. Suppl. 1: 250. 1903; N. L. Britton, Torreya 3: 105. 1903; Briq. in Chod. & Hassler, Bull, Herb, iss., sér, 2, h: 340, 1067—1068, & 1155—-116,, 190); Thiselt.-Dyer Kew. Suppl. 2: 106. 1 3 » Emm. Vasc. Pl. Surinam, Jacks., Ind, Kew, hg 1: 507. 1906; Hayek in Fedde, Repert. Nov. Sp. 2: 1965 Moldenke, Monograph of Lippia 1 He Rusby, « 8: 116. 19123; Loes., Verh. Bot. Ver, Brand. ary 7-78 & & “52283. 1912; Blin, Jardin 26: 262. 19123 Rio de la Losa, Bol. Cienc. Méd. 3: 21-25, 1912 Krinzl., Bull. Jard, Bot, — 13: 90-9. 1913; A. Chev., Etud. Fl. Afr. Cent, Frang¢. 1: 22. 19133 Fe M. Bailey, Compreh,. Cat. « Pls 382. 19133 Je XK. Small, Tis Southeast, Ue Se, ed. 2, 1007 & do12. a3 Britton & Br., Tlustr, Fl., ee Fi 3: Prain, Ind. Kew. Suppl. is 142. 1 ; Buysman, Flora 207: 355. aos i Holland, Kew Bull. age th jin “i 517-518. 1915; Shreve, Carnegie Inst. Wash. Publ. Se Presid. S. Moore, Journ. Bot. 57: 247. 1 Argent. Buenos Aires 88 (imiv): go doors 1919; Alvares, on e « Nov. Expl. - Afr. ; Frang. "hs sO3e. 1920; Urb 59a 595. 1921; posta Ind. Kew. Suppl. 5: %, zat, & 153. 19213 & 1100. 19223 Urb. in » Repert. Nov. “Bp. 18: 19 195. 19223 ncporen ages cm Estud. pot. *wordéste 38 22h. 1923; M. Martinez, Cat. Alfabet. . Vulg. Cientif. Pl. 12 PHYTOLOGIA Vol. 12, now. 1 Mex., [ed. 1]. 1923; Molfino, Not. Bot. 2: 103. 1923 os cam Standl, Contrib. U. S. Nat. Herb, 233 1235 & 1243-129. 19 E Repert. Nov. Sp. 20: 3. 1924; Mansf. intial Bot. Gart. Berlin 9: 155. 192h3 Mak., Ill. Fl. Jap. (78). 2 Johnst., Proc. Calif. Acad. Sci., ser. h, 12 (ys ian a4 & 1926; Sade Minist. Instrucc. Publ. ° 23 pl. "6. 19263 Hegi, Illustr. Fl. Mittel-Eur. 5 (3): As & za. 19273; Bonnier, Fl. Compl. France Suisse & Belg. 9: pl. 97. 1 273 H. He Rusby, Mem. N. xe Bot. Gard. 73 33. 1927; Gonzalez, week Lombardo, Pl, Diaph, Fl. Urug. 1 fAnal, Univ. Montevid, 137): 33 1928; Do- ape Mg Med. se 1928; A. B. Seymour, Host Ind. N. Am 588. 1929; Massart nt Mission Belg. Brésil 1922-23 1: fig. 432. 19235. rs W. Hill, Ind. Kew. Suppl. 7: 139. 1929; Seckt, Fl. Cor- » 416-118 & 623. 1 1929-193, Baeza, Nomb. Wulg. Pl. Silv. ’ > Fis Bull. Torrey aks Club ub Bs ety 1931; Chiov., Fl. Somala 2: 359. 1932; Rydb., Fl. Prairies & Plains 679 & 9119. 19323 > song Pl. Amer » W. Hill, o Sud 3h) & 349-350. 1933; A. W er — uppl. 8: 137. — Dinsmore in Post, Fl. Syria, ed. 321 ‘ "B91. a Je Ke Southeast, Fi. 1140 & 15332 1933; Bain nes, Cat = Gen, Pl, Cult. dard, Prado : ° 19333 *» vil Y. e Nippon Shokubutsu Zufu (ep. Bot. Tllustr. Album] 717, 19335 Dop in Le~ moo Fl, cet ndo-chine hj: sie: & 761. 193k « Standl., sah | oa, wage Le Coint e, Amaz Py asa III arv. & ° . e, Syete Bot. 62. 193k; Moldenke ~~ Piytologia a 95 . 98 (93h) and « Inst. Bot - Buenos Aire oa Fedde, ° (1936) and i; 10° él, ak 134, & — 71936) T. Meyer, Rev. Argent. Agron, 23 cx 1936; o Os Hoehne, Resen,. Hist. Comm. 1965 Moldenke, Monograph of Lippia 13 Viges. Sec. Bot, Agron. Inst. Biol. S. eet ne, & 161. 19373 P. C. Standl., Field Mus, Publ. Bot. 17: 207--208. 1937; C. L. Lun- omb. Vulg. & Cientif. Pl. Mex., [ed. 2], 339--3h0. tad “nieens Revist. Sudam, Bot. }: otc 19373 ei bet Ind. enman Herb. 46), mss. 1937; Tron Darwiniana 3: 50-~53. 1937; Mol- denke, Revist. Sudam. Bot ds (1937) - bi 2. 1937; Cory, Texas e Exp. Sta, Bull. Be 88. 1937; B. Sm. Lilloa i: ° Mus. Bot. 18: 1008--1011. 1938; Moldenke, mile Bot. Lect. 22: (1). peel Moldenke in Fedde, Repert. Nov. Sp. 3: 12 (1938), 45: 126 1938), and 6: 201. 1939; Pittier, Supl. » Venez 55, 62, 97, & 119. 1939; respond Beih, Bot. Centralbl. 59: 275. 1939; Herter, Revist. t. 6: 97. 1939; Pittier, Gen. Pl. Venez. C e 23. 9393 Moldenke, Alph. List mn Names 2, 4, 5, 7, 8, 10—-1h, 17, 19=-21, 23, 25--30, 32, & 3h. 1939; Rylander, World F. a & 708. 19395 Moldenk®, Brief Course Syst 19395 Moldenk : 286, 291 291, “2 259" (1939) at “4 570, 371, 37h, 376, 38 ¥580--386, 403, LOk 408, 413, b15, & 118-432. 1940; Robledo, Lecc. Bot. 2: 498. 19h0; Moldenke in Ridin He 6G t 3$--3%. — B. H. - Davis, Mycologia 38 170. 1940; a ce Carn ig Prelin. - List 2h 45--)8, & 6. 1p Bl; Mol- denke in Pulle, Fl. Surinam, k (2): 260 267—-270, & 27h. 190 nke, Phytologia 1: 412, 423--429, : r 55, & L66—— 9 19. eo saet Darwiniana 5: 13. 19); Hutchins. & Bruce e, é 191: 178. 19h1; Sperry, Sul Ross State Teach. Coll. Bull. 22: hi. 19hl; 0. B. win. ie Leeag uid-Afric. Bosbouver. 6: 92 & 7. 191; Rosengurtt Gurvish, Chron. Bot. 6: 10-11. 191; Herrera, Sinop. Fl. Cuzco 1: 351-—352. Be yicay. Calderén & Standl., Fl. Sal- vad., ed. 2, 237--238. 1941; Biswas, Ind. Forest. Rec. Bot., new ser., 3: 2. 191; Oppenheimer & Bah genic Bull. Soc, Bot. Genev. 31: 363. 19la; Moldenke, Suppl. List Invalid Names 1—7, 9, 1, & hin: 191; Moldenke, Darwiniana 5: 168, 170, & 171. 191; Worsdell, Ve ser - Chil. Moldenke, Lilloa 6: toe 301° (19h) and 8: li—lak, 20, mete & 423-427. 19h2s A. M. T. Davis, Study Boscaje Palma 33 & é2, the- sis. aie ole tz-Gerald, Herbarist 8: 36--37. 1942; Moldenke in 1, PHYTOLOGIA Vol. 12, no. 1 i oO peri? Ps Pa sirens Babs APURS ports, Metorta & & Le6n, Contrib. » & 381. 1942; Meeuse, Blu- kong gy “iskzy cet C. as Lundell, , Contrib, Univ. Mich, Herb. 8: 1. & 92. 192; Moldenke, Known Geogr. Distrib. Verbenac ay oe ob 12 17, 20--35, ale 38, 710-13, las, Sh, 43% a 95, & igh2 Moldenke, Alph. List Invalid Names h, Wap B35 20% 27-33, 3%, fo, 46--L9, 57, & 58. 19,2; stelifeld, Os ieee Mic ° 43; Parodi, Darwi udam . 73 Nat. Urug. hi 2354 1933 Niemeyer & Stellfeld, Arquiv. Mus. Paran. 3: 18. 1 1943; Raim pep a Bol. es Hist. Nat. Jav. Prado 7: 2h1--2)2. 1943; Eo L. D. S » New « Encycl. 1279. 19kh; L. H. Bailey, Man, Cult. Pl. Epa & 825, ole 1 lioldenke, Bot. Gaz. 106: 162. 3 Prof. A. M. B 19 () Lilloa 10: 337-340, 3h2——3hh, 365—~367, & 375--381 (194) and 1s ° 3 Be. P. Reko, Mitobot. Zapot. 7, & 1 945; ana 7: 86, 1945; Dar & J mosome Atl, 2ihé 19453 P. A. Acuffa, Ls Cc 19h5; M. M z, Bol. » Bot, Mex. 2: 2 & 13. 1953 Le Cointe, . 289. 19453 Roig y Mesa, Plant. M 793, & 79h. 19h53 Mo ogia 2: 65, 67, 69—7h, 76, 78— mtrib - Hist. Nat. + 19463 Augusto, Fl. Rio Grande 5 Sul 217, 221, 223 = 25-235, fig. 105d. 1946; Moldenke, Alph. List cit? 1 a. As si, se? 22-2, 26 or E 2 .— a 3, sh 3 s $7, 69, 73 ey hae? ag Nh 94--99, 106, 07, 1h, hho, » nf 20, 421,’12h-~128, 131—135, 137° 1h0, 145 1 16h, 179, 188--190, 192, 19h-~196, 201, 203, 205, 22° 215-2 2h, cae 233, 238, P 2h2, 26, 249-251, 255, 259, 262—~26L,, 266, , 300 318, 319, & 323-325. #5. bs idenk: 563 Mold a i 307, Sie 1 enke, gia 2: 156 (196) and 2: 226, 233, 2h, 310, 3th——319, 330, a om SuL, 3u7, 363, 365--366, & 383-387, 1947; Moldenke, Co Syst. Bot. 93. 197; itz, Veg. No: tre Rios, ed. 2, 15, 19, 87, 122, 150, 165, 172, & App. iv. 19473 Daniel Cent. Antioq. h. 19473 Falefo, Guia Jard Bot. Rio Jan, h2. 19h:7 « Acad ace Cienc. Nat. 21, fig. 3. 19h7 Moldenke, Alph. List det Invalid, ser, : 180 & ET 19473 Moldenke, “Lille 13: 9. 19475 Moldenke at a Venez. Ciene, Nat. 11: 37, 38, L2--l6, 18) & ie _19h7; W. H. Hodge, Re- vist. Fac. Nat. Agron. 7: 313. "1975 c » Revist. ecare ent. Agron. 1h: 252. 1975 Glover, Checklist Brit. & ftal. Trees 268. 1947; Robyns, Fl. Sperm, Parc Nat. Albert. 23 138139" & 616. 1947; 1965 Moldenke, Monograph of ae 15 638, fig. 2hhb. 1948; H. N. & A. L. Moldenke, Pl. Lite 2: 16, 18, 20--23, 30, 19—53, et Bly 68, 69, 71, 73--79, 81, 82, & 86—~ 90. 198; Moldenke, Castanea 13: 113--118 & 120. Toss Moldenke, 2 70 705, 709=-712, Tih —716 35 725, 7 26, 729--7 3, 738, 70, Th5-— 752, 75, 755, 757, 759, 62" a 770, 772-775, 780--783, 785——- 788, 7 798-802, 80), 806-821, 813--8 2h 903, 905-917, 996, 998—-1001, 152 971s & 7m oben 8 1025 se BE 108, 1046, 109, 1060, 1062, 106k, 1067, 1068, 1070—108h, ee 1099; 1100 1110 112 901192 1197, 1201-120); 1208-1211, 1213-1221; 1223, 1227; 1228, 1233— 122, Labb—1251, 125-1261, 1286, 1289, & 2B0— 13h. 1949; idenke, Known Geo ed. 2), 2 > wer es 31, ; TT —50, 53 53, Sh, $638, 60 72, 73, 80, 81, 97, 99-101, 10h, 105, 1, “te, 153, 161, 162, 189—191,"& 2ih. 1949; J. L. Ho 11 & 99. 19493 Westcott, Pl. ay go Handb. 56, 88, 5148, 7120, . nald in A. Gray, Man. Bot., ed. 8, 1212 Be bn an Fam. Dicot. 200 & "23h 1950; Matuda, Am. Midl. wat : S76. 19805 16 PHYTOLOGIA Vol. 12, now 1 F. A. Barkley, Determinac, Ejemp. Herb. Fac. Nac. Agron. Medellin 2 (1): 11 & 1h. 1950; Basi Ecology 31: 28h. 1950; Langman, Sel- de M 223-22. 1950; Moldenke, R Sudam. Bot. 8: 167 & 173. 1950; Moldenke, Venez - Nat. 11: 289. 1950; Moldenke, Phytologia 3: nites ~-273, 285, 287, 289-293, 30h, 305, 375-~378, & 1 (1950) and 3: h19—20, 58, 60, 466, 485, 487, & 508. 1951; Steliteld “trib. Farmaceut. 19°(10); 16hi't & 1668-165 « 1951; F. C. Hoehne, Ind, Bibl. Num. Pl, Col. Com. Ro 348--3L9. 1951; Moldenke, Am. Journ, Bot. 38: 326~-327. 1951; Shreve & We h. 1951; Ragones 45—52, Sh, Ae 59, é1m-6h, 72, 2, 83, 85, } and 624548, 50, 51,57, 62, » tig. nity 1951; A ra de. Vargas, Bol. Hist. Nat. (Santiago, Chile] of 45. 19513 Moldenke, Spec. Subsp. Cont, Mold. Set 45 [2]. 1951; Moldenke, Inform. Mold. Set Me Spec. 3. 1951; Moldenke, Biol. Abstr. 25: 3051 accor Sg 26: 1471. 1952; F.C. Hoehne, Relat. Anual Inst. Bot. » Paulo 1951: 139, oe Je He Hunziker, Revist. Invest. Agric. rf (2): 17h--1 175» ae 8 92. 1952; Howell, Wasmann Journ. Biol. pity 1952; F. Miranda, Veg. Chiapas 1: 287—-288. 1952; Mol- ; New gsr sgh 69==89 19523 P gia 70, 73, & 12h (1 $2) and hs: 19— 180, 187, 189, 190, 199, rhe ge 267, & 292. 1983" Moldenke, Biol. Abstr. 27: 1887, 2026, & 3124, 1983 mild, Victoria Falls # San 9 2: 265 ogia lh: 259. 19533 E. J. Salisb., Ind. Kew. Suppl, 132 & 138. 1985 Stancati, Biol. abstr, 25: 4060 (195) and 2B: 3299 ‘ 3533. i9ct; Calpouzos, Econ. Bot. 8 223-——228, 230, & 232. 1 3 Lombardo, Invent. Pl, Cult. Montevid. 145, 248, 265 ; ta Asoc. so T ‘ P Index n.p. 1954; M. C. Carls Club 81: 391 & M , Bull. Torrey Bot. 396. 195k; Rambo, Sellowia 6; 60, 8h, pare: 53. 1 95h Barton & De Mayo, Journ. Chem. Soc. Lond, 1954 By (3): 887-900. 1954; wild, South, pom Bot, Dict. 101. 5 Brenan, Mem. N. Y. Bot. "card. 9: 36. 195k; Moldenke, Inform, ag Set 8 Spec. [3] (195) and bd Spec, 2, 195); ete, — ore Hort. Soc. 15: 80. 195k; argas gery 90h. 1954; Moldenke, Phytolo- gia hs 509 (95h), 52 2h, 26, & 9599 (195h)) and $1 206-227 pi ow. Trin. & Tob. 6 » Dict. erms Pharmacog. 128 ae & e253. eos Seadoo Ciencia 15; Lille. 2 19553 Calpeeee, ol. Abstr. 29; 19 cr ae ‘ Mendes 3 pe 8 7: 3766. 19555 2 Afr. 5 ter, Revist. Sudam. Bot. 10: 259, 1956; Barton & De M Biol. Abstr. 30: 066. 1956; Raedowski : Inst. Biol. ws i 273 193. 1956; Rambo, Sellowia 7: 260, 281, & 288, 1956; Moldenke, Biol. 1965 Moldenke, Monograph of pane 17 Abstr. 30: 3551. 19563 Angely, Cat, - Fanerog. 17: 1956; Specter, Handb,. Biol. Data ny : or “1586; Moldenke in (Whe Madag. 17: 13--15 & 270, fig. i3; 1956; Molden- ke, Inform, Mold. Set 51 see! pec. 3. 19565 oldenke in Dawson, Los Angeles Co. Mus. Contrib. Sci. 7: oto, he: 2 & 3. 19573 Mol- el 1958; De Roon, In Direct. Spec. 229. 1958; Hoida, Biol. Abstr. 30: 4066. 1958; Strausbaugh & Core, Fl. We Va. 3 776 1958; B. H. on bee. W. Journ. Biol. 16: 1958; Moldenke New a & Br. Somat Fl., pr. 2, 3: 123 & 135. 1958; ‘Bristaiy, 8 & Chamber ee Woods vr 78. 1958; Angely, Fl. P waz 27 17. Hes Yoldenke, Am ° Nat. 59: 333. 1958; fast 2 cone: - India 18: 31& a. 1959; hie Taylor, Ind. Kew, Sup 909 roa t eee ae 6: b 7, 450 40, h2--L5, 47, 48, 2-9, Tlinetr, An i 13: 206. 1960; ae Pizarro, Espec. Plant. Costa Garci 3 307, 310, Rll, 3h, Angely, Fl. Paran 12 & 2h (29605, 16: $9-260 (1960), and 17; Ub. yea; Tamayo, Bol. Soc, Venez. Clenc. Nat. 22: 128 & 173. 1961 pevagyres. ae Soc. Bot. Mex. 26: 143 & 148. 1961; Troncoso, Bol. sareeeee ot. 9: 181—~185. 1961; pene Rhod. Agric. Journ. 58 (3)s 1 73-177. 1961; o, Arbus aie 3 “eave til. Pas. Publ. fete} Sear, Darwiniana 12: 266-298, —l, fig. 1~18. 1 sea, Phytologia 7: So "150, ‘ 516 (1961) and 8: 8, 130, 13 in; & 133~~137. 1961; Anon., Assoc. Etud. Tax. vo 1960+ 60" (age 2 ‘ne ;menoos = reas 3 Cockbill, Weed Abstr. in’ ‘Be is8. 19025 od bed se aoe 32: Suppl. 790=~792. ate e, Senck, Biol. Me 282 & = 19623; Maria, Pl. Vall. - 38. 1962; Fra~ dusco, Revist. Inst. Munic. Carlos Thays 2: 67. vigees Hocking, io as = 190. 196 - South ss Sia Sci. Inst. rset 3 te 31. & er (2): 232. 19625 a Th — eur: Fs a 1962; Moldenke, soe ogia 8: best och, “1, 388 (1962), 18 PHYTOLOGIA Vol. 12, no. 1 8: 518 (1963), and 9: 7-8, 99, 187, 350, 389, & 393. 1963; Sou- kup, Biota i:'261, 288, 305, 3% ; ole Am. 370, 491, & 576. 1963; Hocking, Excerpt. Bot. A.6: 45S. 1963; Oo, Biol, Abstr. 42: 909. 19633; Anon., Biol. Abstr. 2: 1517, B.57, & B.108. 1963; Steyerm., Fl. Mo. 1257 & 1262—~1263. 3 Journ. Genet. 58: 358 & 377. 19633; 643. 1963; Meikle, Kew Bull. 17: 173. Tax. Fl T : ’ Fam. F «Pl. S. Afrs 128 & 129. 1963; Raven, ¢ Rev. Biol. 38: 163. 1963; Moldenke, Résumé Suppl. 6: (1963), 7+ b-—-8 & 10 (1963), 8: 1, 2, & b—6 (1964), 10: 2-6 Sreaxt and 11: 4-8. 1964; Moldenke, Phytologia 9: 503 & 509 (1964) and 10: 59, 60, 91, & 170—172. 19643 As L. Moldenke, Phy- 10 » 1964s Soukup, Biota 5: 16 & 62. 196h3 S Guide Lit, Flow. Pl. Mex. 160, 177, 208, 263, 121, LO, S15, S16, 588, 596, 628, 78, & 1010. 196k Holdenke, Biol. Abstr. 4S: 2772, 5019, 6657, & 7026. 1964; Moldenke, Phytologia ll: 510. 1965. Erect. bushes, shrubs, unders 8, or subshrubs, rarely trees, brous or variously pubescent with simple hairs, often hirsute or tomentose, sometimes functionally dioecious, in some grassland regions with herbaceous branches issuing from a basal or subter- ranean ys xylopodium; leaves opposite or ternate, rarely alter- nate or in 8a »s i » vary: tire to variously toothed or lobed, exstipulate, peti olate or sessile, flat or rugose above, va from thin~membran- ous to vy-coriaceous, mostly penninerved; inflorescence indeter- ite, centripetal, — or capitate, solitary or fascicled in rminal staminate plants have flowers with well-developed anthers and a- , or compressed and o arinate or 2-alate, sometimes 2-lipped, its rim 2- or l-fid or h-dentate; corolla white or variously col- ored, hypocrateriform or infundi » gamopetalous, inferior, zy- gomorphic, its tube cylindric, straight or incurved, very slender, slightly exserted from the calyx or rarely elongate, equal in di- 1965 Moldenke, Monograph of Lippia 19 ulatiaea a at lipped, h-part the lobes A tg oft retuse at thes apex, the posterior one entire, emarginate, or even bifid to about the middle, the lateral ones exterior, the anter- ior one often 1 Fy ens » inserted at about the middle of the corolla-tube, included or slight ’ sometimes only 1 or sent or completely absent on pistillate plants; anthers ovate pendaged, the thecae allel, often unap par aborted or even absent on pistillate plants; pistil one, "often a- borted or non-functional on staminate plants; ovary superior, globose, compound, 2-celled, each cell. l-ovulate; style single, often short; stigma single, rather incrassate or capitate, oblique or recurved; ovules basal and erect or affixed laterally near the base; fruit sm small, dry, ovoid, included by the fruiting-calyx and sometimes adnate to it, dividing into 2 pyrenes or nutlets at maturity, the pericarp papery and hard, the exocarp membranous and rarely ee from the pyrenes; seeds without endosperm; cotyledons eee ( 1923) gives the sporophytic chromosome number for ppia as 32 and 36, but he includes the genus Phyla in his con- cept of L tippta so it is not clear which of these mmbers actual- ly applies to Lippia. Troncoso (1961) reports that in various species of the Section Euza a (which she calls Section Lippia) "there exist feminine divi s whose flowers lack stamens, but which nevertheless develop fruits and seeds. These meh evidently receive pollen from other individuals or are apomictic. a nh a with appar- ently hermaphroditic flowers, on the contrary, ne fruit and their flowers fall very quickly. From the large pind of speci- are masculin sm ina group of species of Lippia, it was necessary to oe ror specimens of the other sex and this was done in a number of cases. two species, can now be interpreted as secondary sexual differences." She says "El género Lippia.......siempre ha sido considerado de feet hermafroditas, En realidad la mayorfa de las especies lo son, pero en el Silindooss Lippia, Secc. Lippia......he descub- pect un grupo francamente dioico, aunque a primera vista no lo - Las flores funcionalmente masculinas conservan un gineceo abortivo, pero en cambio las flores otalmen en el género andino de Malvdceas Nototriche, donde Krapovickas (1957) habla de 'androdioecia morfolégica' ne 'dioecia funcio- Sh yr Sse as follows: "I. Plantas tandadinn:... caste tulos pequefios. Flores por lo general sin rastros de e ap a tigry o estan- 20 PHY 7050 G6:3<4 Vol. 12, no. 1 inodios, muy rara vez 1 6 2 estambres abortados. Corola m4s re- ducida. Tubo corolar angosto, cilfndrico. CAliz bien desarrol- lado, conspfcuo, acrescente en el fruto. Corola persistente, atin en el fruto en de esarrollo, Gineceo normal; estilo largo, casi hasta la ae aie del tubo corolar. Fructificaci6n normal, II. Plantas masculinas......Capf{tulos mauores. Flores con } estambres t should be noted here that the entire rete by Junell (1931) of the gynoecium morphology of Lippia is based on species which are now excluded from the genus (Aloysia, Diostea, and Phyla and is therefore valueless here, The genus is mentioned by Rosen- gurtt Gurvich (1941) as one of the genera in Uruguay with most lefi sas" Lippia is a genus of about 252 species, "varieties, and named forms, widely distributed in subtro opical and tropical America, a fow also in tropical portions of the Old World. The genera Acan- tholippia Griseb. (5 species), Aloysia Ortega (51 species and var- ieties), Burroughsia Moldenke (2 species), Nashia Millsp. (7 spe- cies), Phyla Lour. (20 species, varictie es, and ani forms), and a por- tion of Diostea Miers (3 species) are often included in on Spe’ but are kept separate by me. Cryptocalyx Benth., = alone more piles, in that of Phyla Lour. and i — iden. ed in honor of and is dedicated to A Auguste Lippi (1678-170). ~ Italian explorer and naturalist in Abyssin- he was red at the age of 26 by natives. Souku (1963) refers to him ‘a a French physician and botanist and gives his first name as "Augustin", while Acufla (1945) oe . of c - The a. of the genus is Tippia americana L., based on specimen no. 1 under genus 801 in mn Herbarium at i in the Linnean + London = s of interest “ei ace, bas Bentham (1876) regarded the enus, in its broad e, on 90 species, Baker (1877) gon & — later (1900) says ines 60", and Pearson (1901) says The genus was plac ela (1744) in his group Alatae, by Linnaeus (1753, 175) in his Didynamia Angiosperma and later in his Stellata, by Adanson (1763) in his Verheneae, and by Rtfling (1774) in his Lonicerae. In 1789 A. L. Jussieu classified it in what he called the Vitices, section II "Flores spicati, in spicis 1965 Moldenke, Monograph of Lippia 21 alterni", along with Petrea Houst., Citharexylum B. Juss., Duranta L., Lantana L., Tali alea Aubl. {now known as Amasonia L. Hi Tamonea Aubl. [=Ghinia Schreb. ], Verbena {Dorst.] L., and the non- verbenaceous genera Spielmannia Medic. [=Oftia Adans., in the Myoporaceae] and Perama Aubl. [in the Rubiaceae]. In 1806 the same authority placed Lippia in "les Verbénacées". In 1790 it was placed by Necker in his P. ° "p " according to Steudel, 1841], along with Amasonia, Cith- arexyium, Duranta, Lantana, Petrea, Clerodendrun Burm., D ee Neck. [=Amasonia] Ovieda | Le [-Clerodendrun], Premna L., Volkameria L. [=Clerodendrun], as well as the nae betawoous Spielmannia [-oftia, Myopora Myoporaceae], Selago L. [in the eee and 35 other genera, of which 12 are now placed in the Scrophul iaceae, 5 in the Acanthaceae, 3 each in the Pedaliaceae, oes. ae, and Bignoniaceae, 2 each in the Gentianaceae and Gesneriaceae, and 1 each in the Caprifoliaceae, Martyniaceae, Loganiaceae, Oro= banchaceae, and Polemoniaceae In 1805 J. H. Jaume Saint-Hilaire first proposed the name which we now employ for the family, Verbenaceae, and placed Lippia in his Section 2 "Fleurs disposées en épis iitacuse” along with Cith- Zapania Juss., and the non carbonanacun Perama and Spielmannia. Sprengel (1825) placed Lippia in the Section tion Verbeneae and End- licher (1838) in the Tribe Lippieae of this family. Reichenbach, however, classified (1827, ee 1833) Lippia in his Section Ver- ly Labiata beneae of the seed (1847, 1851 y di ies the genus Lippia into 5 sections and 3 subsections, as follows: Section 1. Al Aloysia. Racemi vel spicae laxae. Calyx inaequaliter h-fidus. Section 2. Goniolippia. Capitula densa tetraquetra, pedunculata, urima ht gemina-=-p ae Section 3. Dipteroc e = a densa plurifariam imbricata, 8 squarrosa. Calyx compressus, bicarinatus vel biala- longe ciliatus, Sieriiees bifidus. Sects wiih Zapania. Capitula densa plurifariam imbricata. nae neque. post anthesin auctae. Calyx eus,. Subsection 1. Axillifloras. Capitula axillaria. Subsection 2. Paniculatag. Capitula densa terminal i-paniculata. agente at Corymbosae. Capitula oe subspicata vel su » 1axa, corymboso-panicul Section eo Rhido olippia. Capita tula leone. Os orator aequaliter imbricata. Bracteae membranaceae, latae, petaloideae, pur- purascentes, post anthesin demum ¢ onspicue auctis. 22 PHYTOLOGIA Vol. 12, now 1 Bentham (1876) says "Genus a Schauero in sectiones 5 imprimis ad inflorescentiam dividitur, dein ad variationes calycis et fruc- tinguendas, etsi characteres florales fere eidem." He divides gracilibus laxis, bracteis parvis angustis, calycibus in pleris- que dense hirsutis", and Zapania, with "spicis densis cylindraceis Vv. globosis, bracteis latis obtusis v. brevissime cuspidatis rarius acutiusculis nunc herbaceis imbricatis » nunc coloratis in prov. Salta, semina tamen in specimine nostro nondum perfecte matura. Albumen parvum verisimiliter adest in speciebus pluribus Lippiae, et Acantholippia facile pro sectione Lippiae habenda, calycibus dense hirsutis floribusque Aloysiae, sed habitu fruticuloso saepe spinescente, foliis parvis oppositis v. alternis saepius lobatis et marginibus valde recurvis subtus quasi can iculatis, spicis saepius brevibus densis. Species adsunt 3 v. h, omnes ex America aus Lan similior, (1891) duo caeterum : Briquet (189) divides the genus Lippia into 10 W char- acterized as follows : —* Subgenus ke Aloysia (Ort.) Schau, Verlungerte Trauben oder » terminal, oder in Rispen gruppiert. Blumen oft in Scheinquirle z usammengesogen. Bracteen klein, oft abfallig. Kelch fast gleich h-spaltig. Ausschliessiich Cc. * Subgenus 2. Zapania (Scop,) Benth. Kurtze, gewthnlich zu Kupf- mi lungernd, mit d eesin ver t decussierten, gefa en dachziegelig sich deckenden Bracteen, axiller, sc atialt: ! Blumen sehr klein. Kelch unbefltgelt. Zahlreiche sttdameri- kanische, besonders brasilianische Straucher. Section 2. Acantholippia {Griseb.) Briq. Ktpfchen etwas thrig, sich ver. mnd, mit dachziegelig sich deckenden Brac- teen. Kelch unbefltgelt,. Kleine, dornige Straucher mit kleinen, rtfc gebogenen e Section 3, Dipterocalyx (Cham.) Schau. KUpfchen klein, mit mehreren Reihen dachziegelig sich deckenden Bracteen, axillar, oft mit den reduzierten Blutter art Rispe bildend, Kelch zusammengedrttckt, beiderseits mit einen schmalen, be- 1965 Moldenke, Monograph of Lippia haarten Fltigel versehen, kurz 2-teilig. Frucht im Kelch eingeschlossen. Tropisch-amerikanische Stra#ucher. Section }. Buzapania Briq. Ktpfchen mit mehreren Reihen dach- ziegelig sich deckenden Bracteen, wa&hrend der Bltttezeit sich + verlimgernd. Bracteen concav oder flach, vias Kelch deck- end. Kelch kurz rthrig, hin und wieder zusammengedrtickt, aber nicht fltgelig. Subsection 1. Axilliflorae (Schau.) Briq. [not "Axillifolorae" as written by Augusto (191,6)]. Reafoken axilltr, + gestielt. sahivetiche tropisch-amerikanische Arten von sehr Ra enem Aussehen Subsection 2. Paniculatas (Schau.) Briq. K¢pfchen halbkugelig, mit sehr gedraingten Blumen, gestielt, einen + rispigen oder cymUs-rispigen, terminalen B ltttenstand bildend. 8--9 brasil- ianische Arten. Subsection 3. Corymbosae (Schau.) Briq. Ausgebildete Kepfchen nftrmig verltngert, in rispigen Dolden gruppiert. 6--7 brasilianische Arten. Section 5. Rhodolippia (Schau.) Briq. KUpfchen zu Anfang der Bltttezeit halbkugelig, mit gleichen, dachziegelig sich i enden Bracteen, rh einen Involucrum verse racteen des » dlass, lila oder rosafarbig, Involucrmms bre nach der Bltttezeit pore wachsend, endlich sehr gross, aderig. Subsection . KOpfchenstiele zu mehreren in den Blattachseln. Mexikanische en, Subsection S. KOpfchenstiele einseln oder zu 2 in den Blatt- achseln, e sche Strttucher mit prtichtig lila- oder rosafarbig pesgebataeten: Bracteen des Involucrum. Of these groups, his Subgenus 1, Aloysia, is now regarded as a separate gems, A ia Ortega; his Section 2, 2 acemolppee, is now, er » the separate genus Acantholippia Griseb.; and his Subsection te oy is now, in its herbaceous or subherbaceous part, the separate genus Phyla Lour. With the re- moval +. a peed ‘velongin to these segregated genera, there er Section 2, Spinulosae Moldenke, nom. nov. Foliis spinulis oientas. 1; alae tees, Section 3. Dipte — L. adotnsis Hochst. [=L. abyssinica (Otto & Dietr.) Cuf.], L. americana L., L. hemi rica Jacq. (=L. americana], L. hirsuta L. f., L. hirta (Cham.) Schau., 2h PHYTOLOGIA Vol. 12, no. 1 L. ocephala Schlecht. & Cham, Subsection 1. Axilliflorae — L. alnifolia Schau., L. argyro- phylla Schau. [=Lantana aristata (Schau.) Briq. Np L. aris aristata ee {=Lantana aristata (Schau.) Brigq.], L. asperrima Cham., rifolia L. C. Rich. [=L. Javanica (Burm. f.) Spreng. ip : a Mart. & Schau, [=L.1 bellatula Moldenke], L. ferru- ginea H.B.! HBK., L. filifolia Mart. & Schau & Schau., L. ta HeB.Ke (=L. alba (Mill.) N. E. Br.], L. hermannioides Cham. [=L. mic- rocephala], L. L~ iodophylla Schau., L. linearis H.B.K., L. ,. mic- rocephala Cham., L. micromera Schau., L. nana Schau., Le pseudo-thea (A. St.-Hil.) Schm., L. mila Cham., L. riedel- jana. Schau., L. erceitilons Mart. & Schau., L. thymoides Mart. & Schau., L. turnerifolia C Subsection 2. Seditinias = S- ceseetass Cham., L. herbacea Mart., 2. hieracifolia Cham., L. intermedia Cham., | L. oxycne- mis Schau, » L. vernonioides Cham. Subsection 3. _Senyabos ae — L. acutidens Mart. & a sq L. corym= bosa Cham., L. grandiflora Mart., L. lacunosa Mart. & Schm., Le 2 Tasioaiyotnn Cham., L. rotundifolia Cham., L. Di cisstans Section c Rhodolip Subsection ]}. peg Moldenke, nom. nov. Pedunculis in axil- lis bebigteaies mmerosis — L. cad tommaatolis H.B.K., L. um- Cav. Subsection 5. Brasilianae Moldenke, nom. nov. Pedunculis in ax- illis foliorum solitariis vel a L. elliptica Schau., L. eupatorium Schau., L. florida Cham., L > ee gardneriana Schau., L. hederaefolia Mart. & Schau., % L. lupulina Cham., L. rhodoc- nen nenis Mart. & Schm, Troncoso (1961) adds to Section Goni ostachyun, L. obscura sriq.; to Section Dipterocalyx, L. hassleriana Chod., L. Ce 2 Briq.; to Section Euza {which she calls Sect. aj, section pani te e she calls "Series", i ee (Brig.) Troncoso [=L. tristis var. aberrans Briq.], L. balansae Briq., pry contermina Brig, L | cordacea Briq., L. longepedunculata Kuntze, L. modesta modesta Brigq., L. L. phaeocephal a Rrig., L. polytricha Briq., L. tegulifera Briq., L. trachyphylla Brigq., L. tristis Briq.; to Sub- agers Paniculatae [which she calls "Series" ], L. sca - re laxis , pedunculatis axillaribus, insertione see a peti- olo distanti (supra-axi Nari}, bracteis lanceolatis, pro rata mag- nis, coloratis, floribus breviter pedicellatis, calyce hirto, bi- labiato-bifido, h-dentato. -- 1, Briq. It should be mentioned here that some o the references given in 1965 Moldenke, Monograph of Lippia 25 ps em geet of this genus on previous pages are cited differ- ly (and sometimes incorrectly) in other works eo the wes US. po eon Apex “ag ntioned the foll s Je Linn. Soc. Lond. Bot. 20: 225. 1883 [often cited as S waekaniy Blue, Bijdr. 1: 321-822. 1826 [often cited as "Blume, Bijdr. 811"); Cham., Linnaea 7: 275 [this is apparently an error or seme ease aes er is no mention of Lippia on that page]; Coult., Contrib. at. Herb. 2: 328-329. 1892 [often cited as "1891" or as nid3ie) 3 Delchev., Rev. Hortic. hl: 316-317. 1872 [often cited "Carr." as author, but the article is definitely signed by belgian not Carriére]; Desv., Journ. Bot. k: 177. {apparent- nay an error of some sort, since there is no rma ; Lippia on page); Emory, Rep. Ue S. & Mex. Bound, 127. 1888 [often cited as "Torr., Bot. Mex. eA? "27, i889"); Hook., Bot. aaa ae > Masel 1829 fenge ee cited as "Bot. Misc. 1: Med. Paris"); pee 1851 rl ci Fl. Bras. 10: 2i9")}; Woldenke, Holmbergia ];: “cae My “ole E [sometimes wise trae cite "1944"]; Moldenke, Phytologia 2: 65--125 5 [often ceils cited as "1! 19hhn};” Morong, Britton, & vail, ye N. Y. Acad. Sci. 7s 196-197. 1892 [sometimes cited as "Morong, N ot. Exo 1 {sometimes cited as "Reichenb., Hort Bot. ecg Pree we of as " " ; Thiselt.-Dyer, Fl. Cap..5 (1) 189-197. 1901 Faden cited as "1910" or mgion]; Walp., Nov. Act. Acad. Caes. Leopold fee Nat. Cur. 19: suppl. 1: 377. 1843 [sometimes cited as "17, 1 377"]3 J. K. Small, Fl. Southeast. U. S., ed. 1, 1012 (1903) "hia ed. 2, 1012. 1913 f sometimes cited as "101)"]. Zohary (1962) describes an ecologic association in Palestine called the Lippieto-Trifolietum fragiferi association, but it is Phyla nodiflora (L.) Greene to which he refers here. te a few vernacular or common names have been recorded by authors in the past for the genus as a whole. Mostly, > they refer only to species native or cultivated in the area with which the particular writer is acquainted; er they refer to to an unidentified plant thought by the wri a member of this genus; often er pecies now pla in segregated era. Neck 90) "lippée"; Gerth van Wijk aie Westcott (1950) lists _ttog-truite and "lemon-verbena"; Lombard (1954) records "salvia trepadora"; and Soukup ce 1963, 196h) 26 PHI TOLOG] & Vol. 12, no. 1 lists "aimara", peadpepalia®, "cedro pashaco", "chichara caspi", "cidra", "huari pancara , "micho caspi", rile "orégano", "pampa orégano®, and —ty sone Eyies and others record "zezuru', "zimbani", and "zumb ani" from Rhodesia. Actually, the name fog- fruit applies only to the genus ays Lour., not to Lippia, while "Zitronenlippe" applies to Aloysia t riphylla (L'Hér.) Britton. The name "cariaquito negro" recorded for Lippia by Ll. Williams on his collection no. 12877 actually belongs to Varronia globosa Jacq., while the "escoba ne; negra" recorded by Hjalmarson pes Ae to Cordia cana Mart, & Gal., both members of the Ehretiaceae ~~ Razd (1950) avers that dissemination of members of ie genus Lippia is endozoic. Seymour (1929) lists Meliola cookeana Speg. and M. lippiae Maubel. as infesting an unidentified species of Lippia. Houard (1933) calls attention to the awcal that Lippia species are infested with insect galls in Argentina (he probably refers here to L. turbinata Griseb.). Woatenee. (1950) records the following animal and fungal pests of Lippia: Spot Anthracnose (Sphaceloma oma lippiae) in Indiana; Black Mildew (Meliola lippiae) in Florida [both of these probably are pests of P Phyla, not dippials Southern Blight (Sclerotiun rolfsii) in Califomia [this is Aloysia triphylla, not Lippiat}; Leaf Spot Ba eae lippiae ae widespread; Cylindrosporium lippiae in wees seg Knot Nematode (Heterodera marioni); and Root Rot (P hymatotrichum omnivorum) in Members of the genus are said to be used as a tea and as a scent in Rhodesia, also to supply of this herb comes from Europe, These latter plants belong to the genus Origamm of the family Lablatas. Species in other genera are also called ‘ore regano', or some variation of this nam find in many parts of the world, The term ‘preganc} should not be used ular her Le Cointe (1934, 197) speaks of < Lippia commonly aga Hees in S8 Paulo, Brazil -~ this is probably L. alba (Mill.) N. E. Br. T ha nn ine as ga ens (as far as ~ 9 an — at Fv ta pabeioes to ay rete authors. 1965 Moldenke, Monograph of Lippia 27 Material has been identified by various botanical workers as Lippia in the past which actually belongs in such other verbena- ceous genera as scan oe, Sees Burroughsia, Citharexylun, Diostea, Junellia, Lanta Nashia, Neosparton, Phyla, Stachytar- Lantana, Nashia, Neosparton, Phyla, Acrocephalus (Lamiaceae), Avicennia (Avicenniaceae), Buddleia (Loganiaceae), Machaoni: Machaonia (Rubiaceae), Microdon (Selaginaceae), Rhaphiodon (Lamiaceae), and Valeriana (Valerianaceae). ran Herbarium 608297, distributed as Lippia, are “are actually Acrocephalus villosus Benth. (Lamiaceae), of which the first-mentioned is the type collection; Galeotti 795b is Batis maritima L. (Batidaceae); M. E. Jones 370 is Borreria podocephala Benth. (Rubiaceae); M. E. Jones 366 is Buddleia marrubiifolia Benth. ( Loganiaceae); Bernardi s.n. and Linden 121) are Cordia SP. (Bhretiacons)5 Hjalmarson s.n. [Hon Honduras] is Cordia cana Mart. & Gal.; M 1585 is Eryngium prostratum Nutt. (Ammiaceae) ; Glaziou ST is Hyptis longipes St.-Hil. (Lamiaceae), Hassler 11062 is Hyptis sp. aff. H. lutes- cens Pohl, Hjalmarson s.n. [Honduras] is H. urticoides Kunth, and Pittier 1,945 is Ryptis sp.; JUrgensen 2179 is Lantana glutinosa Poepp.; Swartz s.n. [Jamaica] is the type collection of Machaonia Varronia g globosa gg list of spe species and of names proposed in this genus, but now excluded therefrom, follow Goniostachyum citrosum Snall = Lantana microcephala A h. Lippea hirsuta hirsuta var. purpurea Hort. = Lantana Dr baaariane H.B.K. Lippea juncea | Gay = Diostea juncea (Gill. & ill. & Hook.) Miers Lippea lycioides (Cham.) Steud. = Aloysia gratissima (Gill. & ok.) Troncoso Lippia achyranthifolia Desf. = Lantana achyranthifolia Desf. Lippia aculeata L. = Phyla pene EEE) Crome Lippia aegyptiaca Carr. = Phyla nodiflora var. reptans (H.B.K.) Moldenke Lippia semptiece Delchev. = Phyla nodiflora var. reptans (H.B aK) Mo Lippia affinis Briq. = Aloysia sellowii (Briq.) Moldenke Lippia aloysioides Loes. = Aloysia aloysioides Loes. & Moldenke Lippia aphylia R. A. Phil. = = Diostea scoparia (Gill. & Hook.) oe appencuate Robinson & Greenm. = Burroughsia appendicu- ta (Robinson & Greemm.) Moldenke ore arguta Mart. = Lantana aristata (Schau.) Briq. Lippia argyrophylla Schau. = | Lantana aristata (Schau.) Briq. 28 PHYTOLOGIA Vol, 12, no. 1 Lippia aristata Schau. = Lantana aristata (Schau.) Briq. Lippia aristata aristata f. plu ripedunculata Kuntze = Lantana aristata var. angustifolia (Kuntze) Moldenke Lippia aristata var. angustifolia Kuntze = Lantana aristata var. tifolia (Kuntze) Moldenke sippls 2 aristata var. pluripedunculata Kuntze = Lantana aristata ¢ angustifolia (Kuntze) Moldenke Sonia asaate abate Urb, = Nashia armata (Urb.) Moldenke Lippia aspenfolia Mettam = Lantana scabrifolia Moldenke Lippia asperifolia Benth. = Phyla scaberrima (A. L. Juss.) Moldenke Lippia asperifolia Reichenb. = Ph: Phyla scaberrima (A. L. Juss.) Mol- Lippia barbata T. S. Brandeg, = Aloysia barbata (T. S. Brandeg.) ~~ Woldenke denke Lippia betulaefolia Humb. & Bonpl, = Phyla betulaefolia (H.B.K.) ene Lippia wotuinefolis Humb. & Kunth = Phyla betulaefolia (H.B.K.) ~~ Green Lippia tabaleetolth i H.B.K. = Phyla betulaefolia (H.B.K.) Greene Lippia betujaefolia Kunth = Phyla betulaefolia (H.B.K.) Greene Lippia betulifolia H.B.K. «= Phyla betulaefolia (H.B.K.) Greene Lippia betulifolia Kunth = Phyla betulaefolia (H.B.K.) Greene Lippia caespitosa Rusby = Phyla caespitosa (Rusby) Moldenke Lippia canescens Humb. = Phyla 1 & nodiflora var, canescens (H.B.K.) oldenke Lippia canescens Humb. & Bonpl, = Phyla nodiflora var. canescens 7 Moldenke Lippia canescens ae & Kunth = = Phyla nodiflora var. canescens (H.B.K.) Moldenk Lippia $e H.B.K. = Phyla nodiflora var. canescens (H.B.K.) Lippia gprs Se Kunth = = Phyla nodiflora var. canescens (H.B.K.) oldenke oe ee aterm Lippia canescens Rich. = Phyla nodiflora var. reptans (H.B.K.) Moldenke Tipp canescens Robinson = Phyla nodiflora var. reptans (H.B.K.) enke i teccemmenarmenel Ligsta capitulis ovatis, foliis linearibus antogerrinis L. = inace ~~ Mierodon ovatus (L (L. ) Choisy, § ania cayensis Urb. Urb. = Nashia cayensis Seite Lippia chamaedrifolia Steud. = ' Moysia chamaedryfolia Cham. Lippia chamaedrifotia Steud, = Aloysia olia Cham. Lippia chamaedrioides Steud. = Aloysia chamaedryfolia Cham. Lippia c = Aloysia chamaedryf Cham, olia L, olia —- chamaedryoides Steud. = Aloys chamaedryfolia Cham, Lippia chamissonis Benth, & Hook. = Lantana chamissonis (D. Dietr.) 1965 Moldenke, Monograph of Lippia 29 Benth. Lippia chamissonis D. Dietr. = Lantana chamissonis (D. Dietr.) enth. Lippia chamissonis Schau. = Lantana chamissonis (D. Dietr.) Benth. Lippia chamssonis D. Dietr. = Lantana chamissonis (D. Dietr.) Benth. Lippia chilensis Schau. = Aloysia salviaefolia (Hook. & Arn.) Moldenke Lippia cilindrica Scheele = Stachytarpheta cayennensis (L. C. Rich.) Vahl Lippia citrata Schlecht. = Aloysia triphylla (L'Hér.) Britton Lippia citriodora Humb. & Kunth = Aloysia triphylla (L'Hér.) ri io] nm Lippia citriodora H.B.K. = Aloysia triphylla (L'Hér.) Britton Lippia citriodora Kunth = Aloysia triphylla (L'Hér.) Britton Lippia citriodora (Lam.) H.B.K. = Aloysia triphylla (L'Hér.) n Lippia citriodora (Lan.) Kunth = Aloysia triphylla (L'Hér.) ton Lippia citrodora H.B.K. = Aloysia triphylla (L'Hér.) Britton Lippia citrodora Kunth = = Aloysia - triphylla riphylla (L'Hér.) Britton Lippia citroidora H.B.K. = = Aloysia triphylla (L'Hér.) Britton Lippia cujabensis Mart, = Lantana brasiliensis Lippia ouneafolia (Torr.) Steud. = Phyla cuneifolia (Torr.) Gre Lippia Sabatcita Torr. = Phyla cuneifolia ( Torr.) Greene Lippia cuneifolia Steud. = = Phyla cuneifolia (Torr.) Greene Lippia cuneifolia Torr. = Phyla cuneifolia (Torr.) Greene Lippia cuneifolia (Torr.) Greene = Phyla cuneifolia (Torr.) reene Lippia cuneifolia (Torr.) Steud. = Phyla cuneifolia (Torr.) reene Lippia cuneifolia Torr. & Steud. = Phyla cuneifolia (Torr.) = reene Lippia cuneifolia Zipp. = Phyla nodiflora (L.) Greene Lippia cuneifolia var. angustissima Gr. = Phyla cuneifolia (Torr.) Greene Lippia cuneifolia var. incisa (Small) Blankinship = Phyla incisa Small Lippia cuneifolia var. incisa (Small) Lindheimer = Phyla incisa Lippia = operas Scheele = Stachytarpheta cayennensis (L. C. eli cymosa Sw. = Machaonia cymosa (Sw.) Griseb., Rubiaceae Lippia darwinii Speg. = Neosparton ton darwinii Benth. Lippia densispicata Kunth & Bou ché, Ind. Sem. Hort. Berol. 12. 8L7 = Aloysia densispicata a (Kunth & Bouché) Moldenke, comb. 30 PHYTOLOGIA Vol, 12, no. 1 nov. oe deserticola F. Phil. = Acantholippia deserticola (F. Phil.) nei deserticola R. A. Phil. = Acantholippia deserticola (F. ~~ Phil.) Moldenke Lippia dinteri Moldenke = Lantana dinteri Moldenke Lippis dulcis culeis Sessé & Moc, = = Phyla scaberrima (A. L. Juss.) Mol- Liseie antita aati Trev. = Phyla scaberrima (A. L. Juss.) Moldenke Lippia dulcis var. mexicana Welmer = Phyla scaberrima (A. L. Juss.) Moldenke Lippia echinus Spreng. = Rhaphiodon echinus (Spreng.) Schau., Lan- liaceae Lippia fastigiata T. S. Brandeg. = Burroughsia fastigiata (T. S. Brandeg.) Moldenke Lippia fiebrigii Hayek = Aloysia fiebrigii (Hayek) Moldenke Jippis filiformis Schau, = Phyla nodiflora var. rosea (D. Don) Moldenke iippta a filiformis Schrad. = Phyla nodiflora var. rosea (D. Don) Moldenke Lippia fimbriata Rusby = Lantana achyranthifolia Desf. Lippia floribunda Hort. = Lantana achyranthifolia Desf. Lippia floribunda R. A. Phil. = Al oysia reichii Moldenke Lippia foliolosa R. A. Phil. = Acantholippia seriphioides (A. Gray) Moldenke Lippia foliosa Phil. = Acantholippia seriphioides (A. Gray) Molden- ke Lippia foncki R. A. Phil. Aloysia foncki (R. A. ard ) Moldenke Lippia geminata Millsp, = i microcephala A. Ri Lippia mute ene Griseb, = Lantana ical Lute eesinate 6 sais Griseb. = Lantana lockhartii (Griseb.) opps age Rose = Avicennia germinans (L.) L Lippia gracilis R. A. Phil. = Acantholippia trifida trifida (C. Gay) Mol- Lippia grandiceps Taylor = Aoysis virgata (Ruiz & Pav.) A. Le Juss. io eesoctecitt Hieron. = Aloysia a gratissima (Gill. & Hook.) Lippe grissbachit Lorentz & Hieron, = Aloysia sellowii (Briq.) Lappe reeen (Griseb.) Hieron, = Acantholippia hastulata Griseb. Lippia hastatula Hieron. = Acantholippia hastulata Griseb. Lippia ia hastulata Griseb, = Acantholippia hastulata Griseb. Tippia hastulata (Griseb.) Hieron. = Acantholippia meres Griseb. Lippia pee hispida Gay = Acantholippia trifida (C. Gay) Moldenke Lippia ilan ilan Bailes = Aloysia looseri looseri | Moldenke 1965 Moldenke, Monograph of Lippia 31 Lippia ilan ilan Baines = Aloysia looseri Moldenke Lippia imbricata Kuntze = Lantana achyranthifolia Desf. Lippia inaguensis (Millsp. ) Urb. = Nashia inaguensis Millsp. Lippia incisa incisa (Small) Tidestr. = P hyla incisa Small Lippia incisa Tidestr. = Phyla incisa oe Lippia involucrata Cooper = "= Lantana involuc Lippia —esenoiies Griseb. = - Buddleia Teainstiee (irinibs ) Hoss., Saas 7 ae pny Hort. = Citharexylum ligustrinum Van Houtte uncea juncea Gay = Diostea juncea (Gill. & Hook.) Miers a, dances Gill. & Hook. = Diostea juncea (Gill. & Hook. ) Lippia pas (Gill, & Hook.) Schau, = Diostea juncea (Gill. & Hook.) Miers SEEMS aes (Miers) Gill. & Hook. = Diostea juncea (Gill. & ok.) Miers Lippia = ancea Schau. = Diostea juncea (Gill. & Hook.) Miers pp sumees var. © Gill. =| Diostea juncea (Gill. & Hook.) Lippia ge A. Rich. = Lantana kisi A. Ri Lippia lagustrina Britton = Al Aloysia ieee fs (Gill. & Hook.) ~~ ~‘Proncoso Lippia lanceolata Michx, = Phyla lanceolata (Michx.) Greene sepeS aancoolate (Wiche. ) Greene = Phyla lanceolata (Michx.) Lippia e lanceclate (Pursh) Michx. = Phyla lanceolata (Michx.) ene Lippia Tenseclté Rich. = Phyla lanceolata (Michx.) Greene Lippia seaecuste Rose = Phyla nodiflora var. reptans (H.B.K. ) meses Py ae larceslata var. recognise Fern. & Grisc. = Phyla lanceo- ta (Michx.) Green Lippta te leptophylla tone = Aloysi tophylla Loes. & Moldenke Lippia eerie G. Thuret = Atha arexylum ligustrinum Van Houtte Lippia ligustrina Auct. Amer. = Aloysia gratissima (Gill. & Hook.) Troncoso =e Lignetrine Britton = Aloysia gratissima (Gill. & Hook.) Lippia 3 pela Kuntze = Aloysia gratissima (Gill. & Hook.) Troncoso Lippia ligustrina Lag. = Junellia ligustrina (Lag-) Moldenke Lippia ligustrina (Lag.) Britton = Junellia ligustrina (Lag.) Moldenke 32 PHYTOLOGIA Vol. 12, no. 1 Lippia Ligustrine (Lag.) Kuntze = Junellia ligustrina (Lag.) Mol- Lippia a Ligustrin Nutt. = Aloysia gratissima (Gill. & Hook.) Lippia Tigeeiran G. Thuret = Citharexylum ligustrinum Van Houtte Lippia ligustrina var. casadensis Hassler = = Aloysia casadensis ui iiges ompinony Bri Aloysia gratissima var Lippia ligustrina var. lasiodonta qe = ° paraguariensis (oie) sien Lippia = See er + paraguariensis Briq. = Aloysia gratissima guariensis (Briq.) Moldenke Lippia 1 a Ligustrina var. schulzii Standl. = Aloysia gratissima var. Standl.) Moldenke isppta Ug igs z rinia 0. K. = Aloysia gratissima (Gill. & Hook.) Lippia Tlareeasd Schrad. = Phyla nodiflora var. rosea (D. Don) enke pps uP oides (Cham.) D. Dietr. = Lantana chamissonis (D. Lippia rasitiennee (Cham. ) Rusby = Lantana chamissonis (D. Dietr.) eee nth. Lippia Lippioides Rusby = Lantana chamissonis (D. Dietr.) Benth. Lippia lithosperma Mart. = Lantana brasiliensis Link Lippia litoralis Kunth = Verbena litoralis H.B.K. appre ees R. A. Phil. = Phyla nodiflora var. rosea (D. Mo Lipa litorlis Kunth = = Verbena litoralis H.B.K. Lippia littoralis Phil. = = Phyla nodiflora var. rosea (D. Don) ldenke ppt ae Ts Se Bradeg. = Aloysia barbata ft, S. Bradeg.) Lippia loschartd4 Lockhart Grosourdy = Lantana lockhartii (Griseb.) G. Don Lippia longifolia Sessé & Moc. = Phyl. Phyla stoechadifolia (L.) Small Lippia iooseri (Moldenke) Looser = = Aloysia looseri looseri Moldenke aoe ieciies Steud. = Aloysia gratissima (Gill (Gill. & Hook.) Lippia a icoldes Steud. = Aloysia gratissima (Gill. & Hook.) Tron- Lippe macrastaciys (Torr.) Wats, = Aloysia macrostachya (Torr.) ut stestisios (Torr.) Wats, = Aloysia macrostachya (Torr.) ldenk Lippta macrophylla Cham, = Lantana macrophylla (Cham.) Schau. eis macrophylla hs A. Phil. = A Acantholippia deserticola (F. Phil.) Moldenk Lippia macropoda Torr, = Lantana macr ropoda To Lippia saercekaaien Torr, = Aloysia aczahaniors (Torr.) Moldenke 1965 Moldenke, Monograph of Lippia 33 Lippia macrostachya (Torr.) Moldenke = Aloysia macrostachya (Torr.) nk arse macrostachya S. Wats. = Aloysia macrostachya (Torr.) Mol- Lippia macrostachys Watson = Aloysia mac necrostachys ( crow») Moldenke Lippia maldonadoi Moldenke = Lantana xenica Mol Lippia maritima Kearney = Phyla lanceolata er "Gbbian Lippia mexicana Hocking = Phyla scaberrima (A. L. Juss.) Moldenke Lippia micrantha Briq. = Lantana micrantha Briq. Lippia microphylla R. A. Phil. = Acantholippia a deserticola (F. “Phas } Moldenke Lippia modiflora L, = Phyla nodiflora (L.) Greene Lippia modiflora (L.) Michx. = Phyla nodiflora (L.) Greene Lippia montana T. S. Brandeg. = Aloysia barbata (T. S. Bramdeg.) Moldenke Lippia montevidensis Spreng. = Lantana montevidensis (Spreng.) aes. Lippia myrtifolia Griseb. = Nashia eS, 5 Griseb.) guescxi Lippia nahuire Get Gentry = Aloysia nahuire Gentry & Moldenk Lippia ripensis Urb. = Nashia n: nipensis seaate (te) as Lippia nodiflora Birch = Phyla nodiflora (L.) Greene Lippia nodiflora Cham. = Phyla nodiflora var. rosea (D. Don) Mol- enke Lippia pedi fore Eggers = Phyla nodiflora var. reptans (H,B. K.-) ~~ Moldenk Lippia nodiflora H.B.K. = Phyla nodiflora (L.) Greene Lippia nodiflora L. = Phyla nodiflora (L.) Greene = ets nodiflora (L.) Boiss. = Phyla nodiflora (L.) Greene Lippia nodiflora (L.) DC. = Phyla nodiflora (L.) Gree Tete nodiflora (L.) Eich..= Phyla nodiflora (L.) Greene ia nodiflora (L.) Michx. = Phyla nodiflora (L.) Greene Lippia nodiflora (L.) Rich, = Phyla nodiflora (L.) Greene Lippia nodiflora Lam. = Phyla nodiflora (L.) Greene Lippia nodiflora Michx. = Phyla nodiflora (L.) Greene 1959; Cuf., Bull. e Bote B $e : Suppl. 790. 1962; Hocking, Excerpt. Bot. aed si 45. 1962} “Cuf., Senck. Biol. 43: 282 & 329. 1962; Anon., Assoc, Etud. Tax. Fi. Afr. Trop. Index 1961: 60 5(962) oe 1962: 63. 1963; uoldenke, Ré- Woody o woody tos wn 2 i ‘ats r bush to “undershrub, or in 's, sometimes even more in a whorl, sessile or subsessile; leaf-blades subcoriaceous, oblong or oblong~lanceolate or oblan- ceolate-oblong, acuminate at the apex, attemate at the base, ser- rate from the middle upwards or obscurely crenate, revolute along , lineate-wrinkled and subrugose as well as rather scabrous or * hispid-scabrous and shiny on the upper surface, — ly pubescent or setose and glandular-punctate as well as pale beneath, the lower ones sometimes 10--12.5 cm. long eg 2.5 vat apex, twice as long as the c and subequaling the corolla, Squarrose, rigid, canescent— » the outer ones 3 mn. ; fl odorous or with very little scent; calyx about 1.5 mm. patente"]; corolla hypocrateriform rm, white or whitish to cream or e-yell pium arborescens) with a yellow center, subequaling the sub- tending br. ranis 3 foliis verticillatis ternis, brevissime cetinlxla lanceolatis, acutis, basi attenuatis » Supra rugosis scabris tus subvillosis; capitulis s axillatibus, verticillatis, breviter ee ee subglobosis; bracteis ex. ovata basi longe acuminatis, tubum corollae aequantibus. Sehr wahrscheinlich ist dies die L. polyoepiala Rob. Br., die dieser gelehrte Botaniker in dem Ver- 1965 Moldenke, Monograph of Lippia kl Lippia oder Zapania zu gehtren, was tfhringens auch ohne Ansicht der Frttchte nicht zu bestimmen ist. — Sie wurde aus Samen von Herrn Schimper unter Nr. 305 eingeftthrt, und uns vom Herrn Garten- Inspector Held in Karisruh m mitgetheilt. Die Pflanze ist halb strauchartig, mit langen krautartigen Aesten, die unbewaffnet, und mit kurzen anliegenden Haaren besetzt und dadurch scharf sind. Die Blatter stehen quirlftrmig an drei um den Stengel, sind ganz drei Zoll und dartber lang, einen Zoll breit, lanzettfurmmig, spitz, an der Basis in den Blattstiel verschmi#lert, ’ sehr dicht und conn De auf der Oberfluche runzelig, scharf und zerstreut auf der Unterflache aimee patchee weicher und karssottig. ots Bittthenkepte stehen in Pers? Blattquirlachsel einen an der Basis breit und eirund, verschm#lern sich dann aber plttz- lich in eine lange schmale spitee, die jedoch die Blumenkronen nicht therragt; sie sind zottig und bewimpert. Die Blumenkronen sind weisz, mit einer schwach ins Graue gehenden Feérbung. Die Blttthen haben einen starken Citronengeruch." They say af it — h. Karlsruhe e seminibus ex Abyssinia, Sc r 305. Richard (1851) says of L. adoénsis: "C'est une ae dce exces- sivement polymorphe: tantét — tomenteuse et l'un gris cendré, tan brunftre, et a ae écartés et trés-rudes. Ses feuilles sont ou ovales ongées aigués, ou étroites et a ore , nea ae mar= ph nat de dentelures fines dans leur contour ept la partie inféri de leurs bords qui est pl ey ag les rp re pc seat tre longuement pédonculés, groupés au nombre de trois - aisselle ge _feuilles supérieures. La surface supérieure des est comme bulleuse et c chagrin The t¢ of "Tippin adoénsis was nettactes by Georg Heinrich Wilhelm Schimper (no, I II 1079) on mountainsides close to deep val- worth noting here that Lippia schimperi Hochst. is a synonym of Lantana viburnoides (Forsk.) Vi ccemmnmnimmmteeteed Schauer (1647) and Schumann (1902) regard L. grandite olia Hochst. as a syno of the species here under | greet ae but in given by Richard (1851) separating seen type material of both L. grandifolia [Schimper II.7 734] and of ue adoénsis (Schimper II.1079]. Hutchinson goes so far as to add ven L. rug rugosa A. @ A. Chev. to this synonymy. If this were true, then the statements made by some authors that the inflorescence of L. abyssinica is “very variable, the very numerous heads sometimes 2 PHYTOLOGIA Vol. 12, no. 1 forming a dense terminal panicle" would have justification. Cer- tain authors also make the assertion that the leaves "barely ex- ceed the peduncles in length in the inflorescence"; actually I have often seen them as large there as farther dowm the stem in the true L. abyssinica The species = a aie’ of Section Dipterocalyx, and has been found growing on savannas or Combretum savannas, in grasslands, a- mong underb aah and at the edge of water, as well as on steep mountainsides adjacent to deep valleys, at altitudes of 1000 to 2500 meters, flowering in May, August, October, and November, and fruiting in’ May and August. A notation on A. Chevalier 10655 avers that it is employed in internal medicine in Ubangi. ee (1915) reports its use aecepgerees HI pedi — natives of Nigeria n regard to the color of the : they are @ described as Mmhite" on A. Chevalier 5395 & 5396) iehdtdsh™ by J. G. Baker; "cream-color" on on Be M. ( M. Gardner 37273 "pale~yellow" on "ase 109; and "the color of the | heliotrope of of gardens" on J. 0. Cooper s.n- Common and vernacular names recorded for the species include gu "guilel", "kani ba", "kas{", "kimbo", "kingkilli ba", sosliieie®, t", "mbormbor", "ngZsu", ngtsurut, "nySna", "saa-nunum" and "sisiling hyamo", : Material of this species has been misidentified and distribu- ted in herbaria under the nam e Lantana vib ides Vahl. the other hand, the Dawe 30, esti as L. adoensis, is actually L. chevalieri Moldenke, Humbert 8182 is L L. grandifolia Hochst., Schlieben 319 is L. a (Burm. f.) Spreng., and Le Lely 115 and Somerset aire, are L. nigeriensis Moldenke. Lath distinguishes: bg tropical African species of Lippia 5 heen o him as 1. Bractlets omehontan: roret ‘at the apex, 2..Peduncles very s short... ...0 -L. oatesii Rolfe. 2a. Peduncles elongate ean J. G. Baker. la. Bractlets not orbicular, not — at SA ae regress ao at "the a a $e inat-blitia orbicular to obovate....L. somalensis Vatke. 5a. Leaf-blades oblong, rugose -L. javanica (Burm 7) aa ha. Leaves por oe oblong or oblanceolate-obl wee +L. abyssinica (Otto & Distr. cuf. 3a. Bractlets “not cuspidate,. “ys By 6. Bractlets acuminate at the apex, 7.. Leaves decussate o+eL. ukambensis Vatke. 7a. Leaves ternate 3 burtonii J. G. Baker. 6a. Bractlets merely acute at the apex.L. plicata plicata J. G. Baker. 1965 Moldenke, Monograph of Lippia 43 Chevalier (1920) cites his no. 2771 and two s.n. collections from Senegal, his 12948 and Caille 14772 & 18059 fram French Gui- nea, and his 67, 307, 2772, and three s.n. collections from French Soudan, I regard Ch Chevalier 67 and Caille 11772 as representing L. chevalierii Moldenke. De Wildeman (1911) cites Thonner 23) from Bangala & Ubangi. Baker (1900) cites the following: MALI: Senegambia: Thierry 227. ABYSSINIA: Plowden s.n.; Quartin-Dillon s.n. [Maigouagoua]; Schim- per 42, 734, & 1079. SENEGAL: Heudelot 103. GAMBIA: Brown-Lester L& 28; Ingram s.n. SIERRA LEONE: Scott-Elliot 1,262, 1265, & 11878. NIGERIA: Baikie 11; Barter 768; Millen 143. CONGO LEOPOLD- VILLE: Burton 8eN.3 Dupuis s.n. [Bingila]; Smith | 8.0.5; Soyaux 67. UGANDA: Wilson 130. KENYA: Petherick SNe [Neangara]; Schwein- furth 2230 & 2687; Speke & Grant s.n. [Unyoro]. ANGOLA: Johnston S.n.3 Monteiro s.n.; Welwitsch 5627 & 27i1. Linen walier (1913) cites his nos. 5395, 5 5396, 7483, & 10655 from Ubangi-Chari in the Central African Republic. Hutchinson & Dal- “s (1931) say of this species that it inhabits chiefly the grass Savannas and is found in "Senegal and French Sudan to Northern Ni- geria and Leite Colony, extends to eastern and southern tropical Africa." They cite Baikie 11; Barter 768; Browm-Lester | & 28; Chevalier 20h10; Dalziel 177; Heudelot 1 1033 Ingram s.n.3 Johnson 5h2 & 58h; Mildbraed aed 7238; M Millen len 143; Scott Elliot 4262, — 1878; and Thierry 227. H However, I regard Brown-Lester |, & . 26, Ingram 8.n., and Thierry 227 as L. chevalierii Moldenke. Not having as yet seen any of “the other collections cited, I am not at all certain that all (or any) of them really represent L. abyssinica, In a 16 herbarium specimens, including the type collections of at least some of the names involved, and 12 mounted photographs of L. abyssinica have been examined by m Citations: ERITREA: Pappi 2176 (S), $602 (Ca--99),3)7). ETHIO- PIA: J. 0. Cooper s.n. [Djem-Djem Forest, t, IX.X.26] (K); Quartin- Dillion s.n. [1862] (F-~photo, K, N--photo, Sg--photo, Z—photo photo); Schimper 2 (F—photo, K, N—photo, Sg--photo, Z-—photo), 46 (B), 11.1079 (B, E—116703), IIT .1L52 (E—11670h, F--photo, N, N——photo, 8, si— ate pet Z——photo). CONGO LEOPOLDVILLE: Lebrun 588 (K). : Lugard 109 (K, N). KENYA: V. Bogden VB.555 (Ca-—98L106) ; H. Me Gardner 3727 (K). LIPPIA ABYSSINICA var. PUBESCENS (Moldenke) Moldenke, comb. nov. Synonymy: Lippia adoénsis var. pubescens Moldenke, Phytologia 8: 58. 1961. pega pf Moldenke, Phytologia 8: 58. 19613 Anon., Assoc. Etud. Tax. Fi. Afr. Trop. Index 1961: 60. 1962; Moldenke, Résumé Suppl, 3: “sb. "1962; Hocking, Excerpt. Bot. A.5: 5. 1962. bh PHYTOLOGIA Vol. 12, no. 1 This variety differs from the typical form of the species in having its stems and branches, as — as the under surface of the yt densely brown-pubescen @ type of the variety was ee by B. B. Drummond and J. H. Hemsley (no. 4496) on a grassy hillside with scattered shrubs and shrub thickets » Subject to burning, at Katera, at = ss of 1200 meters, Maskara district, Uganda, on October 1, 1953, is deposited if the herbarium of the Botanisches aa at sean: In all, only se aes specimens, including the type, have been examined Citations: DOANDA Drummond & Hemsley 1,96 (B-type, Z-~isotype). LIPPIA ACUMINATA C. Wright ex Griseb., Cat. Pl. Cuba 215. 1866. ae tet Cat. Pl. Cuba 215, gee. Jacks. in Hook. £7 eT Kew, 2: 95. 189; Moldenke, Known Geogr. Distrib. shane 1], 25 & 95. 19h2; ‘woldenke, Phytologia 2: 36 197; Mol ldenke, Known genet. sire erben aus ieee 189.° 19h9; Voldenke, Alph. Li Zs 1258. 19h9; Alain in rae in, Fl. Cuba h: 288 & 289. 19575 Molderive, "tent 52 & 160 brac tlots subrotund, cuspidate at the apex, subglabro sube ing the corollas; calyx very minute, subcylindric, Setansly bi- cae many times shorter than the corolla~tube; corolla white. The type of this rare species was collected by Charles Wright (no. 3162) somewhere in western Cuba {"Cuba occ."] between 1860 and 1864, and is ia, deposited in the Sauvalle Herbarium at the Aca- peices de’ ak in Havana, Cuba. The species has been collected rocky p mgt the base of cliffs, and on mountaintops, at Rae ond rg pe to 1000 meters, in flowe rin July, and in fruit in on eh March. herbarium specimens, including the t; and 8 moun- ted agree have rll apa me, ee ra; Citations: CUBA: Las Villas: Alain 637 6373 (Z)3 Ekman 162h3 (F— photo, N, N—photo, S, Si-photo, 7— Z—photo), 18959 (S). Province undetermined: C. Wright 3162 [Cuba occ.; Herb. Sauvalle 1758] (F—photo of type, Hv--t ype, Hv—~isot » N--photo of type, Si-- photo of type, Z--photo of type). ACUTIDENS Mart. & Schau. ex Schau. in A. IC., Prodr. ll: 590591 . 18h7. : ography: — Page : 590-591. 1817; Schau. in anh Fl. - 9 262-oh7. isei; Jacks, in Hook. f. & Jacks., Ind. Kew. Py °s. 894; Moldenke, Known Geogr. Distrib. Verbenac . » fed. 1], 37& 55. +: Bp Moldenke, Alph, List Cit. 1: 1965 Moldenke, Monograph of Lippia Ls 223 (1946) and 3: 691 & 712. 1919; Moldenke, Known Geogr. Distrib. erbe [ed. 2], 80 & 189. 199; Moldenke, Résumé 92 & 160. 1959 » to about 2m, tall; stems erect, subsimple, tetragonal, smooth; branches tetragonal, pubescent; leaves decussate-opposite, rather heavy or coriaceous, ovat short—petio- de- pedunculate, eventually subspicate , solitary in the leaf-axils, subcorymbose-congested at the apex of the branches; bractlets mm. long, equali point, the limb bent downwards » velutinous on the upper surface; fruit adhering to the calyx, recondite, oblong, one of the py- renes often tabescent. e type of this apparently rare species was collected by Carl Friedrich Philipp von Martius in fields at Yha, Minas Gerais, Brazil, adjoining the boundary of Gofas, in September, 1818, and is deposited in the Martius Herbarium at the Botanisches Museum Munich. The species is a member of the Section Euzapania, Subsection Corymbosae. It has been collected in flower and fruit May, July, and September. In all, herbarium specimens and ); phototypes have been exan- ined by me. Citations: BRAZIL: Maranh%o: Murca Pires & Black 162ha (Be-- 50206), 2342 (Be—58767). Minas Gerais: Martius s.n. [Yha; Mac- bride photos 20319] (Kr--photo of type, N—-photo of type). Piaui: G. Gardner 290 (M, N). TA AD Bibliography: Hayek in Fedde, Repert. Prain, Ind. Kew. Suppl. : 142. 1913; Moldenke, Known Geogr e 95 (19 189. 1949; Moldenke, Alph. List Cit. 3: args 1 aia Résumé 92 & 460. 1959; Moldenke, Résumé S 2 = 9 * em. tall; branches few, elongate, appres- shrub, about 50 4 Sed-pubescent; leaves decussate-opposite, sessile, imbricate, about 1 cm. long, ovate, deeply cordate at the base, subacute at th he ; e stem flat, the apex, smooth above, glistening ser toward Sericeous-tomentose benea L6 PHYTOLOGIA Vol. 12, no. 1 re species was collected by George Gardner te 030) rts rootas, "Brasll, and is deposited in the herbarium of the Naturhistorisches Museum in Vienna, where it was pean ps by Macbride as his type photograph no. 3328. Hayek comm "Habitu Lippiae teguliferae Briq......non dissimilis, poy autem omnino planis nequaquam nervoso-rugosis, indumento alieno capitulisque longius petiolatis valde diversa." e orig description gives Gardner 2933 as the type collection, but this appears to be a typographic error for "3933" as is edadinly indi- cated on the Vienna holotype. hus far, yo : herbarium specimens and 3 phototypes have been examined Citations: BRAZIL: Brasilia: Murga Pires, Silva, & Souza 9669 (Lw, Z). Gofas: G. Gardner 3933 [Macbride photos 3320) (1t— photo of type, Kr--photo of type, N--photo of type). LIPPIA AFFINIS Schau. in A, DC., Prodr. 11: 576. 1847 [not L. affinis hag se. ie hau. , Prodr. 11: 576. nat Schau. 228, i851;" Rae " Vidensk del xsdeeon.? 1877-1878; 7. 1877; Jacks. in Hook. f. "k sae Ind. Kew. 2: 95.1 Mold , Ab tee al: 167. 1935; Moldenke, & 10 3s Mold (1946), 2: 362—36, 413, S51, & enke, Al (pu8), xt 730 “casus), and = ee! be - 1949; Moldenke, Known Geogr. 9 Venez. Cienc. Nat. 11: 289, 1950; Calpouzos, Eaonom. Bot. 8: 228. 1951 Yoldenke, Tafora. old. Set LB Spec. [3]. 19543 Mendes Anais V Anual Soc. Bot. Bras. 254--255 & 272—— 275. 1956; Moldenke, "hbswas 92, athe & 460. 1959; Renné, Levant. erb. - Agron. Minas 150. i960 Shrub or bush, 0.8-~2 m, tall; branchle ts subterete, tomentose; leaves decussate-oppos site; petioles very slender, about 1 el cm long; leaf-blades ovate, to 3.3 cm. long and 1.6 cm. wide, acute at the apex, rounded or subcordate at the base, crenate along the margins, penninerved, rugose and beautifully holosericeous above, incano-tanentose beneath; peduncles axillary, aggregate, flaccid, s incan: g pep in the h ertaiten of se i opp Museum at Berlin a member of the Sec- F bl po Ha ; 1965 Moldenke, Monograph of Lippia 47 graveolens H.3.K., L. origanoides H.B.K., L. glandulosa Schau., L. elegans Cham., Liv gracilis Schau., L. martiana Schau., L. ob-= scura ra Brigq., L. pendula Rusby, L. hibigiines Schau., L. salviaefo- iia Chan., L. 8 schomburgkiana Schau., L. sidoides Cham., L. velutina Schau., and others. Schauer (1851) says "Lippiae microphyllae accedit capitulis et fere ind dumento, recedit tamen ab ea foliorum figura et magnitudino L. origan anoidi primo ridge similis apparet, atis wh in the treatment of dysentery. The corolla is reported as white on F. C. Hoehne s.n. (Herb. Inst. Bot. S. Paulo “278h & 5217], Sampaio 6829, Mexia 5625, and Williams & ; Assis 5809, and as yel- lowish on ——s Magalhfes 2507. Common names reported for the species ar ", "orégano", "oregano di burro", and "poleo". m "oregano", "oregeno", "origano", or eerneene along with L. formosa T. S. Brandeg., L. fragrans Turez., L. graveolens H.B. K., L. micromera Schau., L L. micromera var. helleri (Britton) Mol- denke, L. origanoides H.B.K., Le “L. palmeri S. - Wats., L. palmeri var. spicata R Rose, L. pendula Rusby, - ‘Lantana - involucrata L ) ee amg L. achy- ranthifolia Desfa; Ia te L. trifolia L., ,., Coleus amboinicus Lour., Hede- gma floribunda Standl., H. patens Jones, Hyptis albida H.B.K., He americana (Aubl.) Urb., He capitata Jacq., H. suaveolens (L.) Poit., Limnophila stolonifera (Blanco) Merr., Monarda austroman- tana Epl., Ocimum basilicum L., Origanun vulgare L., | L., O. majorana p Aes Poliomintha 1. longiflora Gray, and Sal: Salvia sp. The name "poleo" is applied also to L. alba (Mill.) N. E. “E. Brs, L. grisebachiana Moldenke, L. integrifolia (Griseb.) Hieron., Lo turbinata Griseb. and its varieties, and Mentha pulegina L. The L Lippia affinis of Brig Briquet is a synonym of Aloysia sellowii (Briq.) Moldenke. The Kuntze 8.n. [Velasco, 200 n., ORG. ks ee cited hereinafter under L. salviaefolia Cham., may prove to repre- sent L. affinis instead. Material of L. affinis has been misidentified and distributed in herbaria under the names L. berterii ii ae and L. eee H.B.K. On the other hand, the Curran & H carts Barriga 1,875, Lelmann B.T. 815 gee he Fister BOR, » Tl, & 10512, Saer 62 , H, H, Smith 587, and Tamayo 3% & om go as L. affinis, are all L. origanoides H.B.K. tt ee 21 herbarium specimens of L. affinis have been exam- d eC. oltationa: BRAZIL: Minas Gerais: Black & Mendes alhfes 51~- 11776 (Be--6960)) ; F. C. Hoehne s.n. (N, N, Sp, abe Mendes 18 PHYTOLOGIA Vol. 12, no. 1 Magalh&es 2507 [Herb. Jard. Bot. Belo Horiz. 3709] (N); Mexia 5625 (Gg, Go, N, N, S); Sampaio 6829 (Herb. Jard. Bot. Belo lo Horiz. 12307] (N); L. B. Smith 7069 (N, ~Z); Williams & Assis 5809 (G, N). SXo' Paulo: Vie; = & & Viegas s s.n. {Herb. Inst. Agron. Est. S. eae 3980] (W--1775602). “BOLIVIA: Santa Cruz: Kuntze s.n. (Sta. 1000 m., V.92] (N, W--701990), s.n. (Sierra, 2000 m., ¢ 1892} “(W). LOCALITY OF COLLECTION UNDETERMINED: Herb. Martius 12008 (Br). LIPPIA ALBA (Mill.) N. . Br. ex Britton & Wils., Scient. Surv. Porto ti 6: 141. 1925, Nepeta flore albo, spica habitiori Sloane, Cat. Pl. mis . Jamaic. 1: 65. 1696. Lantana alba Mill., Gard. Dict., ed. 8, Lippia no, 8. 1768. Verbena globiflora L'iér., Stirp. Nov. 1: 22-23, pl. 12. 1786. Zappania odoratissima Scop., Delic. Fl. & Fam. Insub. 1: 3--35, pl. 15. 1786. Zapania asperifolia A. Rich. ex Marthe, Cat. Pl. Jard. Méd. Paris 67. 1801 [not L. seperifolia Benth., 1947, nor Hochst., 1947, nor Reichenb., 1828, nor L. C. Rich., 1906]. Verbena globifera L'Hér. ex A. Rich, in Marthe, get Pl. Jard. Méd. Paris 67, in syn. 1801. come globiflora ra a. L L. Se, Ann. Mus. Hist. Nat. Paris 7s 72. Zapania odorata mig Syn. Pl. 2: 140. 1806. Lippia cape H.B.K., Nov. Gen. & Sp. Pl. 2: 266. 1818 [not L. geminata Milisp., 1930, nor Schl., 1964]. Verbena odorata Pers. ex Steud., Nom. Bot., ed. 1, 873 & 898. 1821 [not V. odorata Desf., 1841, nor Meyen, 183), nor jer: 196]. Zapania o ocoeesionine sine Scope og teud., Nom. Bot., ed. 1, 898, in syn. 1821 (H.B.K.) Spreng. in L., Syst. Veg., ed. 16, "2s eT 182 — ta lantanoides Willd. ex Spreng. in L., Syst. Veg., ed. 16, 2: ey se as 2 ee jippte geminata Kunth apud Spreng. in ie, Syst. : 763, in syn. 1825; Schau. in A. DC., Prodr. 11: oB8. *sh7. taco globulifera era L'Hér. apud Spreng. in L., Syst. Veg., ed. 16, 2: 751, in syn. 1825. Lantana mollissima Desf., Cat. Hort. Par., ed. 3, 393. 1829. Lippia citrata Willd. ex Cham., Linnaea 7: 215, in syn. 1832. Lippia asperifolia Poepp. ex Cham., Linnaea 7: 215, in syn. 1832, Lantana odorata Weigelt ex Cham., Linnaea 7:215, in syn. 1832 [not L. odorata Ait., 1847, nor L., 1767]. Lippia citrata Cham., Linnaea 7: 21 7: 214. 1832. Lantana lippioides Hook, & Arn., Bot. Beech. Voy. 305. 1837. Lantana geminata Spreng. apud Steud., Nom. Bot., ed. 2, 2: 8. 18ll. | Ver- bena globulifera Auct. ex Steud., Nom. Bot., “a. 2, 2: Sh & 750, in & syn. 1841. Lippia ia & geminata Humb, & Bonpl. ex steud., Nom, Bote, ed. 2, 2: 5h, con syn. 1841. Zappania odorata Pers, ex Steud., Nom. Bot., ed. 2, 2: 797, in syn. ° -. gappania lantanoides "Lam. 1965 Moldenke, Monograph of Lippia hg Steud., Nom. Bot., ed. 2, 2: 797. 181. Lantana cuneatifolia Klotzsch ex Walp., Repert. Bot. Syst. 4: 45, in syn. 1645. Nepeta maxima flore albo, spica habiliori Sloane ex Walp., Repert. Bot. Syst. lh: 47, in syn. 1645. Lippia geminata Humb. & Kunth ex Benth., Bot. Voy. Sulphur 153. 1846. Lantana geminata Spreng. ex Morren, Belg. Hort. 1: 134, in syn. 1851. Verbena lantanoYdes Auct. ex Morren, Belg. Hort. 1: 134, in syn. 1651. Lantana lippioides Hook. & Arn. ex Morren, Belg. Hort. 1: 13h, in syn. 1851. Zapania globiflora Poir. apud Schau. in Mart., Fl. Bras. 9: 236, in syn. 1851, Lippia geminata var. microphylla Griseb., Abhand. Ktnig. Gesell. Wissen. Gttting. 7: 255. 1857. Lippia panamensis Turcz., Bull. Soc. Nat. Mosc. 3% (2): 201. 1863. Lippia havannensis Turcz., Bull. Soc. Nat. Mosc. 3% (2): 202. 1863. Lippia asperifolia H.B.K. ex Bocq., Adansonia 3: 2h). 1863. Zapania geminata (H.B.K.) Gibert, Enum. Pl. Montev. hl. 1873. Lippia lantanoides Coult., Contrib, U. S. Nat. Herb. 2: 328--329. 1892, Lippia crenata Sessé & Moc., Fl. Mex., ed. 2, 140. 189) [not L. crenata (Griseb.) Kuntze, 1898, nor Kuntze, 190), nor Pearson, 1959]. Lippia havanensis Turcz. apud Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 95. 189). Lippia globiflora (L'Hér.) Kuntze, Rev. Gen. Pl. 3 (2): 251. 1898. Lippia globiflora albiflora Kuntze, Rev. Gen. Pl. 3 (2): 2 » hom. nud. 1898. Lippia globiflora f. pubescens Kuntze, Rev. Gen. Pl. 3 (2): 252, nom. nud. 1898. Lippia globiflora & normalis Kuntze, Rev. Gen. rmalis f, lilacina e Pl. 3 (2): 251. 1898. Lippia globiflora 4 no Kuntze, Rev. Gen. Pl. 3 (2): 251. 1096. Lippia globiflora geminata (H.B.K.) Kuntze, Rev. Gen. Pl. 3 OF 251-252. 1098. Lippia globiflora } geminata f. glabriuscula Kuntze, Rev. Gen. Pl. 3 (2): 252. 1898. Lippia globiflora 6 microphylla Griseb. ex Kuntze, Rev. Gen. Pl. 3 (2): 252. 1898. Lippia globiflora 5 microphylla f. glabriuscula Kuntze, Rev. Gen. Pl. 3 (2): 252. 1898. Zapania lantanodes Lam. ex Kuntze, Rev. Gen. Pl. 3 (2): 251, in syn. 1898. Lippia globiflora Kuntze apud Thiselt.-Dyer, Ind. Kew. Suppl. 2: 106, in syn. 190). Lippia globiflora var. normalis Kuntze apud Briq., Ann. Conserv. & Jard. Bot. Genév. 78: 315. 190). Verbena globulifera Spreng. apud H. Verbenac. Malay. Arch. 369, in syn. 1919. Lippia globiflora var. geuinata (Kunth) Kuntze apud Seckt, Rev. Univ. Nac. Cordoba 17: ee 1930. Lippia lantanoides (Lam.) Herter, Revist. Sudam, Bot. : 185. 1937. Zapania globiflora (L'Hér.) Willd., Poir., & Juss. ex Moldenke, Lilloa \: nate oe 1939. Zapania globiflora (L'Hér.) Juss. ex Moldenke, Prelim. Alph. List Invalid Names 5h, in syn. 19,0, Zapania globiflora (L'Hér.) Poir. ex Moldenke, Prelim. Alph, List Invalid Names 5h, in ayn. 1940. Lippia 50 PHYTOLOGIA Vol. 12, no. 1 globiflora var. normalis f, lilacina Kuntze ex Moldenke. Lilloa 5: 4i9, in syn. 19h0. Lippia globiflora var. geminata (H.B.K.) Kuntze ex Moldenke, Carnegie Inst. Wash. Publ. 522: 165, in syn. 1940. Lippia globiflora var. geminata f, glabriuscula Kuntze ex Moldenke, Lilloa 5: 419, in syn. 1940. Lippia globiflora var. microphylla Griseb. ex Moldenke, Prelim. Alph. List Invalid Names 31, in syn. 1940. Lippia trifolia Sessé & Moc. ex Moldenke, Pre- lim. Alph. List Invalid Names 32, in syn. 190. Lippia virgata Sessé & Moc. ex Moldenke, Prelim. Alph. List Invalid Names 32, in syn. 1940 [not L. virgata Steud., 181, nor (Rufz & Pav.) Steud., 191]. Verbena odorata L'Hér. ex Moldenke, Suppl. List Invalid Names 9, in syn. 19hi. Lippia alba H.B.K. ex Moldenke, Suppl. List Invalid Names 5, in syn. 191. Lippia germinata H.B.K. ex Raimondi, Bol. Mus. Hist. Nat. Jav. Prado 7: 2hl. 1943. Lippia alba (Mill.) P. E. Brown apud Roig, Plant. Medic. Cuba 79, sphalm I9h5. Verbena globuliflora L'kér. ex Augusto, Fl. Rio Graride do Sul 235, sphalm. 1946. Lippia alba N. E. Br. ex G. L. Fisher, Am. Bot. Exchange List. 196. Lippia alba (Mill.) Britt. ex Moldenke, Alph. List Invalid Names Suppl. 1: 13, in syn. 197. Lippia citrata Cham. & Schlecht. ex Moldenke, Alph. List Invalid Names Suppl. 1: 14, in syn. 19h7. Lippia balsamea Mart. ex Moldenke, Alph. List Invalid Names Suppl. 1: 13, in syn. 1947. Leonotis myrtifolia C. Wright ex Moldenke, Alph. List Invalid Names Suppl. 1: 13, in syn. 1947. Lantana geminata var. microphylla Griseb. ex Moldenke, Résumé Suppl. 2: 9, in syn. 1960. Lantana trifolia Sw. ex Moldenke, Résumé Suppl, 2: 9, in syn. 1960 [not L. trifolia Ait., 1947, nor Briq., 1963, nor Cham., 18)7, nor A. Chev., 19h2, nor H.B.K., 1963, nor L., 1753, nor Sessé & Moc., 19h0]. Lippia alba (L.) N. E. Br. ex Moldenke, Résumé Suppl. 2: 9, in syn. 1960. Lippia alba (Mill.) N. E. Brawn, apud Renné, Levant. Herb, Inst. Agron. Minas 150, sphalm. 1960. Lippia alba (Millspaugh) N. E. Br. apud Angely, Fl. Paran. 16: 59, sphalm. 1960. Lippia alba Muell.) N. E. Br. apud Tamayo, Bol. Soc. Venez. Cienc. Nat. 22: 128, sphalm. 1961, Phyla geminata H.B.K. ex Nair & Rehman, . Nat. Bot. Gard. Lucknow 8: pl. 1, fig. 3, text-fig. 13--16. 1962. Lippia citrata var. triphylla Kappler ex Moldenke, Résumé Suppl. 9: l, in syn. 196h. “Lippia lantanoides (L.) Coult. ex Moldenke, Résumé Suppl. 9: 5, in syn. 196). Lippia alba (Mill.) N. E. Britt. ex Moldenke, Résumé Suppl. 9: , in syn. 196). Phryma caprtata Sessé & Moc. ex Moldenke, Résumé Supp}. 9: 5, in syn. mi =e glabriflora Kuntze ex Moldenke, Résumé Suppl. 10: 5, in syn. 1964. Zappania globiflora Poir. ex Moldenke, Résumé Suppl. li: 8, in syn. 196). Bibliography: Sloane, Cat. Pl. Ins, é ‘ oane Voy. Jam. 1: 173, pl. 108, fig. 1. ge ach “i a pina 1965 Moldenke, Monograph of Lippia 51 60, pl. 71, fig. 2. 1755; Mill., Gard. Dict., ed. 8, Lippia no. 8, 1768 ; L'Hér., Stirp. Nov, 1: 22-=23, OL. i2: 1786; Scop., Delic. ar. F e Cat. Pl. Jard. wéd. Paris 67, 1801; A. L. Juss., Ann. Mus. Hist, Nat. Paris 7: 72. 1806; hei ea 2: 140. *1806; Poir. in Lan», Bneyel. Uéth. Bot. : 80. 1808; Dun. Cours., Bot. Cult., ed. 2, 2: 6%. 1811; J. sa. i oe 5 es 36: no. 3. 1817; H.B.K., in Bot. Exot. 23 Si. 1828; Desf., soe Hort. on; ed. a 393. 1829; 2: pl. 169. 1830; Hook., Bot. Misc. 2: 2%. 1831; Cham., Linnaea 73 By 1832; Hook. & Arn., B ot. Beech. Voy. 35. 1837; Steud., Nom. Bot., ed. 2 - B ns —)5, . Voy. 8 ciple 153. 186; Schau. in ae I., Prodr. ll: 382-285, 1817; Schau. in Mart., Fl. Bras. 9: 235—236. 1851; Gesell. Wissen. Gutting. 7: 255. 1857; J. Torr, in Emory, Rep. U. S. & Mex. Bound. Surv. 2: 127. 1858; Griseb., Fl. Brit. West Ind. : 336. 19, 87, 122, 150, 172, & A P. iv. 1878; Griseb., Abhand. Ktnig. Gesell, Wissen. Gvtting. : [Symb. Fl. —- 3 3 oes te Fl. Orient. : 532. 1879; ree er, - Linn. Bot. 20: 225, 1883; Coult., “rs a Ue S. 3. Mat ot. - roa 329. 1892; Jacks. in Bate t.s ve. » Ind. Kew. 2: 95. 189k; Saas & Moc. fs Mex 140. 189 in 7 ’ ; ‘am. ny "(a): 46 & "152, tig. = a c & d. 189); Jacks. in Kk. rae & J e, Ind. Kew . 2: 12h8. 1895; Just, Bot. Jahresber. 23. (2): 77. 1897; Kuntze, Rev. Gen. Pl. 3 (2): 251-252. 1898; Millsp., Field opts oe anes 2: 91. 1900; ve G. Baker in Thiselte Dyer Fi. oan 3 1900; H. H. W. Pearson in Thiselt.- Dyer, Fl. here ease “1901; Briq. in Chod., Bull. Herb. Boiss., a 2, 2: . & . 1902; Briq. in Chod., Pl. Hassler. 1 (9): 198 & 200. 19023 J. K. small, ms Southeast. U. S., ed. 1, 1012. 1903; Durand & Pitas Ind. Kew. Suppl. 1: 250. 1903; Thiselt.-Dyer, Ind. Kew. Suppl. 2: m0. 190k; Millsp. & Loes. in Engl., bey Jahrb 26: Beibl. 80: 25. 1905; Briq., Ann. rie & Jard., B enév. 7-8: 315. 190); Briq. in Chod. & Hassler, pe hag 2 (ai): 493. "50h; Hicken, Chlor. Plat. Argent. 7 ag 1910; 52 PHYTOLOGIA Vol, 12, no. 1 Buchtien, Contrib. Fl. Bolivia 1: 166. nt ae Gerth van Wijk, Dict. Plantnames 1: 767. 1911; Glaz., « Soc. Bot. France 58, Mém. 3: 541. 1911; Urb., Symb. Ant iui. a 532 (1911) and 7: 353. 19123 J. K. small, Fl. Southeast. U. S., ed. 2, 1012. 1913; F. M. Bailey, Compreh. Gat. Queensl. Pl. 362. 1913; Gerth van Wijk, Dict. Plantnames 2: 15). 1916; T. Herzog, Meded. Rijksherb. Lei- den 29: 4. 1916; H. J. Lam, Verbenac. Malay. Arch. 369. 1919; An Host 8 Fungi N. Am. 588. 1929; A. W. Hill, Ind. Kew. Suppl. 7: 139. 19293 Seckt, Fl. Cordob. ia? 1929--19304 ierver bev ee 105. Uni as ° Ss 10 . C. Standl., Field Mus. Publ. Bot. 18: 1008 & 1009. 1938; Mol- denke, Lilloa h: 293--295, 19393 B el *pittier, Supl. Plant. Usual. Venez. 97 & — 1939; Moldenke, t ames 4, 5, » 275:29;5' 2D 5 ee & 26. gra Moléenke, ane Inst. Wash. » Known G D b. Verbenac., (ed. aids 12, 17, 20-22, 25--35, 37, 0, wo 83 73, & 95. 1925 1942; Moldenke in Iundell, Fl. Texas 3 (3: 53h. 19125 Marie~ | Inst. Bot : 307. 1943; Herter, tpy eyed + og 7: 225, 19433 cians, Lilloa & 380, Roig y re ong e= Cu 477-78, 793, ‘ toh. 9s, ee Cointe, 289. 1945; Moldenke, Phytologia 2: 70, 7h, 78, 81, 85-85, 207, & ety 1953 Moldenke, Castanea 10: hl. 1: OLS; Rosengurtt, . Prad. Nat. Urug. 5: hi. 1946; Augusto, Fl. Rio Grande d Paghar "223 & 235. 19h6; G. L. Fisher, Am. Bot. Exchange List. 196; Moldenke, Alph. List Cit. 1: 15, 22, 27, 39, h6, 50, 58, 62, 63, 67, 69, 83, 92, 99, 111, 112, 11h, 120, 126, 127, 131, 1,0, 187 165, 171, 172, 1965 Moldenke, Monograph of Lippia 53 1947; Le Cointe, Amaz. Brasil. III Arv. & Pl. Uteis, ed. 2, [Bras- iliana, ser. 5, 251:] 180. 1947; Daniel, Verb. Cent. Antiog. h. 19h73 E. J. Salisb., Ind. Kew. Suppl. 10: eae rly eb. a9 Moldenke, ; . Known . Distri b. benac. [ed.2], 22, 30, 35, 36, 38, 39, bh, — 53, 5h, 56-58, 60, 63, 66--68, 70, 72, 80, 97, 99, 100, 10h, 127, 161; 189, & 190. 1919; Roig'y Mesa, Lilloa 18: 193. 1949; Rosengurtt, Lilloa 20; 17h. 199; H. Ne & Ae Le Moldenke, Anal. Inst. Biol. Mex. 20: 9. 1949; Moldenke, Alph. List Cit. 3: 65h, 658, 673, 67h, 676-678, 680, 696, 70, 705, 716, 723, 726, 730, ct i, fi, 1595 76h, 766, 5 J Piso & Marcer. Bot. Bras. 73 & 106. 199; Matuda, Am. Midl. Nat. lk: 576. 1950; Moldenke, sate esa Bot. 8: 173. 1950; Fe A Barkley, Determinac. Ejemp. Herb. Fac. Nac. Agron. Medellin 2 (i): 14. 1950; Moldenke, Prytologia 3: 286 "& 381 (1950) and 3: 452, 453, 460, & 487. 1951; F.C. Hoehne, Ind. Bibl. & Num. Pl. Col. Com. Rondon 318. 1951; Ste liteld, Trib. Farmac. 19 (10): 168, 1951; Moldenke, Phytologia : 69° (1952) and hj: acide 19535 F. Mir- Veg . Chiapas 2 2: 265 & 363. 1953; Roig y Mesa, Dicc. Bot. lL: 96, 501, “$00, 509, 8L8, 890, & 943 (2953) pd 2: shy tab 195k; Lom= bard rdo, inyent > Cult. Mf Montevid. 145 gee lowia™ 6: au, ms 153. 195h; Moldenke, ee Calif 15: 80, grok Woldenke, Fl. Trin. 2 (6): 0-92. ~ 19555, Moldenke, Fam. 2 Verbenac. 9 1985; “Hooking, Bist Pharmacog. 128, 1955; P. ekee ral Leon, Revist. Soc. Se hs 12: 51. 1955; Moldenke, Inform. Mold. Set 51 rik 3. 1956; Alain hs 288-—289. 1957; Angely, Fl. Paran. 73 12. 1957; Voldenke, Résumé 27, 36, ae eure 45, 47, 52, 55--60, 62—6h, 67, 71, 75, 77, 78, 80, ns , 116, 119, 125, 163, Sh Poe: Yi T:0 10:6: DA Vol. 12, nosed 209, 220, 301, » 309-313, 316--318, 361, 36, 368, 371, 393--395, 5, li2s, 60, : oe 17 Moldenke, Résumé $ Suppl. 1: 3& (1959) and 2: 7, 9, & 11. 1960 ; Rend, Levant. Herb, Masta Agron. Minas 150. 1960; Angely Fl. Parans ee 5 i 960) and 17: 6. 1961; Tamayo, Bol. Soc. Venez, : 128 & 173. 1961; Moldenke, “Pigtologia 8: 13635 ‘ aT “as6a), and 8: 257. 1962; Hocking, Exc rpt. Bot. A.S: kh. Rehman, Bull. Nat. Bot. Gard. Lackionk 76: 8 & 9, pl. 1, aR “a text-fig. 13--16. 1962; Moldenke, Résumé wet “b 1, 9, 12, 13, 26, & 33 (1962), ki: 7 (1962), 6: 3 (1963), and 7: l, 1, & 8. 1963; Moldenke, Phyto- te. 389 & 393. Ar Soukup, Biota ): 288, Ate ee ea ustrations: Sloane, ay gots 7 pl. "108, fig. 1. 1707; Nair & Rehman, Bull. Nat. Bot. Gard. Lucknow 96: $s 1; fig. An erect perennial herb or mee, eared: densely puberu- lent, low, coarse, erect shrub or subs » to2m a strong lemon (Citrus limonia), lime (C. beast ttoiiay: mint (Men- tha) or sage (Salvia) odor, odor, sometimes s: scrambling, procumbent or semi-proc umbent, or more or less decumbent t, rarely creeping or scandent, usually much branched from the base and with long root- gga at . J g iu 5 ee BE & 8 3 z » trai s opposite or ternate, thickish, very aromatic- -fragrant; petioles slender, 3--8 mn. long, cinereous-pubescent; leaf-blades ovate or oblong, 2-—7 cm. long » usually 1.2--2,3 cm . wide, dark-green a- bove, acute or obtuse at the apex, conspicuously’ serrate or ser- rulate to crenate along the margins (except at the very base) ° hirtellous or puberulent and more or less rugose (when = bove, densely short-pubescent or soft—velvety to tomentose b neath with cinereous 8; fe) ence axillary, capitate, usu- ally much shorter than — rhs bib leaves or only ie petiole, soli oe ie he upper le corolla, not accrescent; flowers fragrant or non-odorous; calyx 2-toothed; corolla hypocrateriform, varying from pinkish—-blue, light violet-blue, pink, light-pink, pinkish, rose, pale-rose, or ht-lilac pale- ,» purple- nt, the endocolpium clear lalongate, 3.5 x 19.2 the lateral ends pointed, the exine oa 8» thick, "the ectine atans 1965 Moldenke, Monograph of Lippia 55 granulate; chromosomes small, their number: 2n = 0 (1,27). The type of this common and widely distributed, apparently sented by the L. H. Bailey Hortorium negative 5057. N.E. Brown says that the Columbia University Herbarium specimen of Pringle 1960 is an exact match for Houstoun's specimen, except that the elsewhere. The specific epithet is sometimes uppercased for no valid reason, The plant belongs in the Section Zapania, Subsection The so-called type of Lippia asperifolia A. Rich. was collected from a cultivated plant in the Jardin des Plantes in Paris, a dup- licate being deposited in the DeCandolle Herbarium at Geneva, this name is actually a nomen illegitimum based on Verbena globi- flora L'Hér. The type of Lippia asperifolia Poepp. was collected by Edouard Friedrich Poeppig somewhere in Cuba. The name Lippia asperifolia H.B.K. is based on a Bonpland collection from Cumana, Venezuela, deposited in the Bonpland Herbarium at the Muséum Nat- i d'Histoire Naturelle at Paris, but H.B.K. (1818) plainly say "Lippia asperifolia Rich." on page 265 of their work, giving Willdenow 1112),, deposited in the herbarium of the Botanisches Museum at Berlin; that of Verbena globiflora L'Hér. is a specimen collected by Alexander Cruckshanks at Lima, Peru. The type of Lip- pia balsamea was collected by Joannesa de Santa Barbara in Rio Grande do Sul, Brazil, in 1836, and is deposited in the herbarium of the Jardin Botanique de l'Etat at Brussels; that of L. citrata is Sieber s.n. from Paré, Brazil, sent by Graf Johann Centurius von Hoffmannsegg in 1830, deposited in the same herbarium; that of L. citrata var. triphylla was gathered by August Kappler (no, 1860) from cultivated plants in Surinam and is deposited in the herbarium of the Naturhistoriska Riksmuseum at Stockholm; that of Lantana mollissima is a specimen cultivated in the Jardin des Plantes and deposited in the Paris herbarium, a duplicate collected in 1825 de- posited in the DeCandolle Herbarium at Geneva. The type of L. havanensis was collected by Jean Jules Linden (no. 72 bis) in the TT province of Havana, Cuba The type of Lippia geminata is a collection made by Friedrich Heinrich Alexander von Humboldt and Aimé Jacques Alexandre Bonpland at Rio Apure, Venezuela, deposited in the Bonpland Herbarium at Paris and there photographed by Macbride under his type photograph no. 39485; that of L. geminata var. microphylla was gathered 56 PHYTOLOGIA Vol. 12, no. 1 Edouard Placide Duchassaing de Fontbressin on the island of Guade- loupe and is said to have been a shrub 1--2 feet tall, with small leaves 1.6 cm. long and 8 mm. wide, subequaling the peduncles, and violet-purple corollas. Lantana trifolia Sw. is based on Duss 4552, cultivated on Guadeloupe island and deposited in the Britton Herbarium at the New York Botanical Garden; Lippia glabriflora is based on Hassler 11112 in the United States National Herbarium. The type of Lippia crenata was collected by Martin Sessé y La- casta, José Mariano Mocino, Juan Diego del Gastillo, and José Mal- donado (no, 2215) somewhere in Mexico, deposited at Madrid; that of L. panamensis is Fendler 220 from Chagres s, Panama, deposited at Leningrad; and t and that of Leonotis myrtifolia was aalleeted by Charles Wright (no, 3157) in Cuba, deposited at Stockholm. Lorentz & Hi- eronymus 1174 is a cotype collection of L. globiflora @ geminata 2. emenenls while L. se a& normalis f. lilacina is n Otto Kuntze collections om (a) Concepcion, northern ifagtay and (b) Matto Grosso, rector July 1892, daposhted in the Britton Herbarium It is worth noting here that the Lippia asperifolia of L. C. Richard, referred to in the synonymy - above » belongs in the synonymy of L. javanica (Burm. f.) Spreng., that of Hochstetter belongs to L. baumii baumii Gurk Gtirke, and that of Bentham and of Reichenbach belong to Phyla scal soaberrina (A. L. Juss.) Moldenke; the Lantana trifolia of Aiton, of Chamisso, and of H.B.K. belong in the synonymy of L. tri- folia L., that of Brides belongs to L. micrantha Briq., that i ae A. Chevalier belo: ongs to L. mearnsii Moldenke, and that of Sessé & Mocino belongs to L. hispida H.B.K. lLippia geminata Schl. is a syn- onym of L. graveolens H.B.K., while L, geminata Millsp. is Lantana microcephala A. Rich. The L. crenata @ of (Griseb.) Kuntze and nd of Kuntze are L. junelliana (Moldenke) Troncoso, while that of Pearson is L. Ee H. H. W. Pearson, r (1883) states that his Lippia oligophylla is "allied to lL. sdsisehick and L. geminata", but , but actually it is not even ver- benaceous -- it is Acrocephalus villosus Benth. in the Lamiaceae. — scaberrima Pers. is included in the synonymy of what we ppia élba by Sprengel (1825), Lippia dulcis Trev. is inaliated by Steudel (181), and Lantana lavandulacea is included by stead in the synonymy of Phyla scaberrima (A. L. Juss.) rie, Ha pane (1939) reduces Lippia origanoides H.B.K. to synonymy under but it is a separate and distinct species. Boissier (1879) placed Lantana indica @ Roxb., L. dubia Royle, and L. collina Decne. in the synonymy of what he called Lantana alba Mill., , but in my estimation Lantana indica is a distinct and valid species, with +; contin piacere as a synon, synonym, while L. dubia is a synonym of L. trifo- 1965 Moldenke, Monograph of Lippia 57 The Lantana inermis, foliis oppositis, ovatis dentatis, flori- bus capitato-umbellatis, folioso-involucratis J. Burm. in Plum., Py Pl. Amer. 60 (1755) and Lantana inermis, foliis obpositis, ovatis J. Burn, Plum., Pl. Amer, mer. pl. qiy fig. 2 (1755) are sometimes cited as belonging to Lippia alba, but I regard them as belonging to the synonymy of Lantana lanaiouneda Britton instead. Verbena capensis Thunb., Lippia capensis (L.) Spreng., L. ca ensis Spreng., L. capensis (Thunb.) Spreng., and L. scabra Hochst, are often giv en in the synonymy of L. alba by various authors, but belong definitely in the synonymy of | L. javanica (Burn. f. ) gpiteng e being based on African material. t is currently being called Lippia alba is apparently a very polymorphic taxon. pirnaniye > should be divided into two species or vardahiue, typified by the L. geminata H.B.K. and the L. globi- flora (L'Hér-.) Kuntze forms. 3 these were to be kept separate ‘ then the name Lippia lantanoides Coult. would certainly belong in the synonymy of the former for which it was only a new name when published, while Zapania lantanoides Lam. would go to the synonymy of the latter for which it was also merely a new name when pub- lished. Schauer (1851) actually keeps the two apart and te to i nc Lantana lippioides Hook. & Arn., Le mollissima Desf., L. odorata Weigelt, Lippia asperifolia Poepp., L. citrata citrata Cham., and Verbena lantanoides Willd. (2) L. asperifolia A. i. Rich. — including Lanta- na lavandulacea Willd., Lippia capensis capensis Spreng., L. scabra Hochst., Verbena globiflora L'Hér., Zapania nia globiflora Poir., Z. lantanoides Lam., Z. odorata Pers., and Z. odoratissima Scop. He says of L. geminata: "Planta habitu semper insignis, neque cum ulla alia con- fundenda. L. citrata et L. geminata omnino non differunt. Capit- - Ita P revs Seéieton videbatur. -- AL. asperifolia Rich. abhorret; toto habitu, foliis supra wisine strigoso~scaberrimis sed mollibus et aetate demum scabridis, praecipue vero — spay bona Guitar alienis." For L. asperifolia his commen "Rigoroso ex- amine discrimen inter r plantan Americanam et sade allud dete- gere non oe nisi quod in Africana bracteae essent magis acu- minatae, flores paullo majores." Morren (1851) also keeps the two parate, Briquet oO says [p. 251] "Nous partageons entiérement 1'opin- ion de M. O. Kuntze que le diastaction des L. geminata et L. asperi~ a theres que Schauer l'a admise dans le Prodromus....ne a me. C'est avec raison que l'auteur a rétabli tpi collective la peroate nore [L. “pinkfiera); con- formément aux lois de la Nomencl. . 57." + Kuntze MSstadt endl says "L. globiflora OK. (Verbena yr: L' fer. 178) = Zapania odoratis- Sima Scop. 1766 = Zap. lantanodes Lam. 1791 = Lippia asperifolia 58 PHYTOLOGIA Vol. 12, no. 1 Rich, 18., = L. geminata HBK. 1817) “ normalis 0. Ktze. Frutex pubescens minus ramosus foliis majoribus (--8 cm longis, ——5 om latis) pedunculis plerumque solitariis; f. lilacina 0. Ktze. Mat- togrosso. Paraguay. Hierzu auch L. citrata Cham. © geminata 0. Ktze. (HBK.) Frutex ramosior foliis duplo minoribus [p. 252] pe- dunculis saepius geminatis floribus albis vel lilacinis 3 f. glab- riuscula 0. Ktze. Caulibus foliisqe glabriusculis. Argentina: Salta, Tucuman (L. & H.). ‘lanceolata Gris. Foliis angustis (1: 4--6). f. incana 0. Ktze. Folia utrinque incana. Argentina: Oran (Hieronymus). é microphylla Gris. Folia minima 1-~1 1/2 cm longa 5-~8 mm lata. f. glabriuscula 0. Ktze. Argentina: Catamar- ca (Lorentz & Hieronymus). Ich kann Lippia asperifolia, fur die der altere Name globiflora einzutreten hat, nicht von L. geminata unterscheiden; weder die Blttthenfarbe (albiflora, lilacina) noch die Behaarung (f. pubescens, glabriuscula, incana) dieser weitver- breiteten Art, noch der Habitus bietet einem Anhalt dazu. grvsser, robuster und verzweigter der Strauch wird, um so kleiner egen die Blatter zm sein, Die grvsstblattrigen Formen wachsen an Flussufern und feuchten Stellen, zeigen aber geringe Verzweigung (jttngere Exemplare?)." Mocquerys 851 was determined by Troncoso as "Lippia alba (Mill.) ppar represent the typical form of the species and deserves nomenclatur- gnition. A notation on Hering s.n. in the Leiden herbarium says "Lantana canescens Kunth Spr. 2: 763; Lantana mollissima, breve pedunculata; affinis iconi Burm. Amer. tab. Tl, 2.2, 9008 The "Lippia asperifolia L. C. Rich." of Pulle, Enum. Pl. Suri- ) appears to be an error for L. asperifolia A. Rich.; the "Lantana sp.?" of Sessé & Mocino, based on their NO. 2195, is plainly Lippia alba, but the "Lippia alba" cited by Lun- dell (1937) is Lippia graveolens H.B.K. marshes and swamps, s or in general, low d lagoons, fields Sunny or moist fields, hedgerows, open or waste places,wet or campos, mois ery Gry savannas, as well as on road shoulders in the full open sun, at altitudes from sea-level to 2500 meters. rowing i It has been found g in clay or wet alluyial clay soil, clay- 1965 Moldenke, Monograph of Lippia 59 loam or moist loam, dry or dry sandy soil, dry exposed soil, deep loose or wet sand, dry alluvial gravel, and dry periodically in- undated soil, blooming in every month of the year and fruiting from February to December. a few pl g an old estate pond at Coconut Hill, in the driest part of the limestone region. Evidently very rare and local." Schipp describes the species as a ra n secondary ng stem diameter of half an inch, pendulous branches, and heliotrope- colored flowers. Standley (1938) reports it from "Thickets of the Pacific tierra caliente, ascending to the region of San José; th gh campos in the Chaco. Parodi (193) says "Arbusto de hojas caducas, originario del Brasil, Uruguay, Paraguay y Noreste argentino hasta la regién orno, p gandose f4cilmente por estacas y divisién de matas." Rosengurtt (1919) describes it as "sufritice recostada entre arbustos de bosques riberefios; poco frecuente" in Uruguay. In Cuba it is described by Roig y Mesa e en algunas de las Antillas Menores, en las Ba- hamas y en la América tropical continental ." In Texas it is said to grow in woods and semi-moist situations arc ties. It has been widely confused there with the Florida and West Indian Lantana involucrata L., which is not known from Texas. Small (1933) and Cory (1937) record Lantana involucrata from the state, probably on the basis of these misidentifications. Pedersen says of it "fairly common in sunny pastures » seems to vador; King avers it to be locally abundant in Tamaulipas. Lombardo (1954) states "Se le encuentra en algunos alambrados y tejidos viviendo en estado espontd4neo y subespontdneo" in Montevideo. The Brittons found it "established after cultiva- tion" on the island of Trinidad. Duss says "planted about all the dens Le it is "usually in gardens [in Cuba], en be introduced." Le Cointe (1934, 1947) speaks of a Lippia species commonly cultivated in the city of S&o Paulo -- this probably is L. alba. Pittier states that the plant is "pigly aromatic", but exactly what he means by this is 60 PHYTOLOGIA Vol. 12, no. 1 not clear since all other authors describe the odor of the plant as quite pleasant. Nair & Rehman (1962) give palynograms of the Dusén 11170, Kuntze s.n. [VII.92], and Miers 1206; and the small- leaved northern form by Herb. Osten 16169 & 17309 and Schroder Se n. (Colonia, Jan. 30, 1922] & sen. (Paysandfi, Dec. 192]. Herter 747, Herb. Herter 82833, and Pflanz 2093 are anomalous. In the event that horticulturists should eventually wish to eparate color forms of this species, the following specimens are described by their respective collectors as having had corollas of the following colors: "blue" — Soeprato 138, Tresling 156; “white, blue, and yellow" -- podson & Thien 1281; "mauve" -- Britton, 768 & 2787, Pittier 13118 3 "light-violet" -- Lanjouw 6 ,» Versteeg 6h; "light-violet with yellow throat" —- Asplund 17605; "violet- purple" -- Grisebach; "pale-purplet — Herb. Herter 62833, Herter 747; "light-purple" -- Pedersen 1906, G. R. Proctor 9421; "purple" —— Dodson & Thien 1260, Hinton 102), T. Meyer 135, Raimondi 750, R. Runyon 893 & 2177, Standley & Padilla V.3230, Wonderly 52; __ "pink-purple" — Leonard & Leonard 15,70, R. Runyon 2603; Wlight- pink* -~ Herzog 1429; "pinkish" -- S. S. White 5679; "pink, yellow or yellowish within” — Rojas 106 & 8 af er 2428; "pink" — Correll & Johnston 17926, Correll & Robbins 20956, R. S. Ferris 5927, Hinton 5720, 126Lh,, 13149, 13585, & 13962, A. S. Hitchcock 20206, Howard S71, Kappler 1960, Mexia 819, F. W. Pennell Tad3, Ruiz Leal 1,266, ass Santos 2533; "rose" -— W. E. Broadway 99, Hassler 7367 & 7397, Herb. Osten 13562 & 20678, Jorgensen 2471, Koelz 22192, Macedo 3280, T. Rojas 1874, 1875, 12212, & s.n. (Concepcion, Dec. 1916] , Rosengurtt B.l232, Ruiz Hui- dobro 2037, 2180, 3078, 3116, 3309, 3343, 3679, & 3088, Sagot L69, Schreiter s.n. (Embarcacion, Dec. 20, 1925], Wroth 59; "pale-rose” ~~ JOrgensen 2171; "lilac-rogen —~ Rodrigo 952; "roge-lilac" -- T. Rojas 12082; "rose, yellow inside" == Herb. Hicken 22, Herb. Os- ten 7534, Te Rojas 406, 495, 187ha, & s.n. (San Bernardino, June 1925); on reddish-violet" — Ekman H.351h; "lilac" —- Bailey & ey 1236, Britton, Britton, & Wilson 5)92, W. Harris 11911, Re A. Howard 1855, Klug 1637, Maced "2612, T. Meyer 10058, 10099, & ee 1965 Moldenke, Monograph of Lippia 61 100), Mexia 1929, Steyermark 35447; "light-lilact — Asplund 14375; "pale-lilac" —- Steyermark 50859; "purple-lilac, white in- side, later yellow" -- Herb. Osten 19033, Herter s.n. (Nueva Pal- mira, April 1926]; "lavender" -- W. E. Broadway 36, | Correll & Sclweinfurth 15661, W. H. Cowgill 2073, Hyerdam & Beetle 22993, R. A. Howard 11718, R. M. King 035, Koelz 2168, E. C. Leonard 8515, Leavenworth 200, Lundell & Lundell 8685, Plant Quarant. 050726, Trin. Bot. Gard. Herb. 8199; "lavender with a yellow eye" on Ga. dee Lundell 1064); "lavender and white" —- Gilly & Hernandez Xolocotzi 2533 "tube lavender, large lip purple, throat orange" ~~ Feddema 578; and "white" [perhaps worthy of a form name] — Angulo 1325, Barbosa da Silva 2, W. Harris 11737, Irwin R.120, Maxon & Killip 16%, Raimondi 10576. It must always be remembered however, in most cases where color variation is given on the basis of various collectors! reports, many of these apparent variations may be due o to differences in interpretation of the same col- or by different collectors or even to defects or imperfections in their color-vision. Common and vernacular names for this plant are very numerous, including "ajeujo cimarron" » “alfombrilla", "alfrombrilla", "alfronbrilla", tanfs de Espafia", "anfs del Espafia", "beukes bosjie", "blakka tiki ment", "bushy lippia", "cariaquito” te 1, : d a : : z ix. ampo" d "hierba del negro", "hierba negra", "hinojo de anfs", "juanilama", "leppie rude", "lippie rude", "malmequer do mato", "malva", ‘mal thee", "mas "sonora", "sonora lila", "St. Mario", "tarete", "te", "te castillans", »t6 de Castilla", "té de Castillo", "té del pais", "te del pafs", "te de playat "toronjil", "toronjil americano", ", "yerba buena americana", ty ématica mato flor morada", "yerba buena cimarona", "yerbabuena cimarrona", "yerba cidreira", "yerba cidrera", and "yuanilama", The name "poleo" is also applied to L. affinis Schau., L. grisebachiana Moldenke, L. integrifolia (Griseb.) Hieron., L. turbinata Griseb. and its varieties, and Mentha pulegium L., while "romero" is ap- 62 tea TOLOOGTIA Vol. 12, no. 1 plied also to Baccharis rosmarinifolia Hook. & Arn. and to Senecio rosmarinus Phil. Johnson reports that L. alba is an aromatic sudorific used by the natives of Campeche in an infusion called a tea, "palisado", by them; Hinton reports that a brew is made from the very fragrant leaves in Mexico and used to treat stomache-ache. Asenjo 30 was actually purchased on the drug market in Puerto Rico. Hocking (1955) refers to the species as a diaphoretic, antispasmodic, em- menagogue, and stomachic. Biolley refers to it as a "medicinal cording to Duss. Its leaves are used for tea in Colombia, accor- ding to Proctor. Lados says of it "planta aromatica, estomacal, ports that a tea of the leaves is taken in the same country to counteract the effects of purgatives. It is "medicinal" in Argen- ina, according to Wroth & Morel. In Surinam, according to Lan- ouw, the herbage is used for bathing in the treatment of fevers. Roig (1945) says "ln Camagttey la llaman Quita dolor y la toman de re Menta americana. En San Antonio de los Baflos lo llaman Hinojo de anfs y lo usan en infusién contra los célicos hep4ticos. En otros lugares de la provincia de la Habana la llaman poleo y la usan camo pectoral. En Puerto Rico también la llaman Poleo. Segfn Standley la planta tiene propiedades sudorfficas, anti-espasmédi- cas estimulantes de la Salvia de Europa y se usa en los mismos casos a la dosis de un manojo de la planta en una botella de agua hirviendos tomando por tazas la infusién endulzada en el término del dfa." In his 199 work he says that it is used as an anti- dysenteric in Cuba and South America; in his 1953 work he comments "us. como medic - En la parte Sur de se le encuen- tra en antiguos bateyes. Su cocimiento se usa contra los dolores de est a." It should be mentioned here that the illustration labeled "Lippia asverifolia" by Briquet in Engler & Prantl, Nat. Pflanzen- fam. (3a): 150, fig. 58c@D (1895) appears to represent L. javanica, The H.B.K., Nov. Gen, & Sp. Pl. 2: 266 (1818) reference ab erence "Humb, & Kunth, Nov. Gen. 2: 21)" occurring in literature has not been located by me, nor have I been able to trace "Lam. Til, 2: 10"; LtHér., Stirp. Nov. 1: 23, pl. 12 (1786) is often misdated "176" or "i785"; Marths, Cat.’Pl. Jacd’ ued. Paris 67 1965 Moldenke, Monograph of Lippia 63 (1801) is often referred to as "Rich. Cat. Hort. Med. Paris 67" Coult., Contrib. U. S. Nat. Herb. 2: 328-329 (1892) is sometimes erroneously cited as "1891" or "1894"; Ragonese, Darwiniana 5; 413 e J. K. Small, Fl. Sout east. U. S., ed. 1, 1012 (1903) and ed, 2, 1012 (1913) references are often incorrectly cited as page "101)", Material of Lippia alba has been misidentified and distributed in herbaria by various workers under the names Aloisia citriodora Ort., Aloisia sp., Goniostachyum graveolens (H.B.K.) Small, anescens Kunth, L. canescens H.B.K., L. chamissonis D. tana c Dietr., L. citrosa (Small) Moldenke, L. indica Roxb., L. involu- crata L., L. lilacina Desf., L. odorata L., L. odorata Sessé & Moc., L. origanoides H.B.K., L. originoides H.B.K., L. recta Ait., L. reticulata Pers., L. sellowiana Link & Otto, L. trifolia L., pa EN th ha L. velutina Mart. & Gal., Lantana sp-, Lippia balansae Brigq., L. even Bacopa aquatica Aubl., Hyptis stricta Hoffmgg., Mikania hasslerana f. cuneifolia Hassler, and Waltheria americana L. Fries 1415 is the type collection of L. chacensis Moldenke; Aguilar Hidalgo 0, M. E. Jones 29186, Millspaugh 41, Parks & Cory 19425, Small & Wherry 11921, Tharp 590k, and Vogd s.n. (Pharr, June 18, 197] are L. graveolens H.B.K.; Tamayo 326 is L. origanoides H.B.K.; M. E. Jones 23231 is L. palmeri S. Wats.; Archer 4667, Herb. Inst. Miguel Lillo 32056, Pedersen 1163, and Rodriguez 23h are L. recolletae Morong; Venturi 354 & 1789 are L. suffruticosa (Griseb.) Kuntze; Lorentz & Hieronymus 236 & 373, Jorgensen 102), and Werdermann 47h L. turbinata Griseb.; Bai- letti 37 and Herb. Inst. Miguel Lillo 32219 are Lantana aristata var. angustifolia (Kuntze) Moldenke; Archer 227 and Pittier 1289 are L. boyacana Moldenke; Killip & Smith 16399, Miller & Johnston eats Cette Cr tee pet cae eet 2172 are L. canescens H.B.K -3 Pittier 6172 is L. fucata Lindl.; Acle & Guerrero 15, tu, Covell, & McCart 45, Cory 51332, H. C. Hanson 346, M. E. Jones 29186, Martinez & Trevino 29 & 13 » McCart 6h reas OL0 OTA Vol. 12, no. 1 7395 & 7460, and Tharp 590 are all L. macropoda Torr.; Gaumer & sons 478 is. L. microcephala A. Rich.; | Miers 85 is Phyla nodiflora var. rosea (D. Don) Moldenke; and Killip 11616 is Hyptis lacus- tris St.-Hil. ~The W. H. Cowgill 2073, cited below, cultivated in Maryland, was raised from seeds of Koelz 2168 Fart Dacca, Bengal, India. Range (1935) cites his no. 3325 1 Box cites his no. 1570 from An- dies Ragonese (191) cites his - no. 2137 from Santa er Argenti- na; Calderén & Standley cite this species from Santa Tecla, El vador. Matuda (1950) cites his nos. 1478, 5213, bar 16645 Mexico, and Rosengurtt (199) his B.4232 from Uruguay. Hodge (1947) cites Archer 22 227, Giraldo s.n., and To: Toro 315 from Colombia. Raimondi (1943) cites the following: PERU: Cajamarca: Raimondi Jou. Lambayeque: Raimondi 750. Lima: Raimondi pen & 11879. Loreto: Raimondi 217h. Province undetermined: Raimondi 1287, 2175, 9890, & 11 & 11222. Augusto (1946) ¢: cites: BRAZIL: Parand: Pio Corr€a s.n. Rio mee do Sul: arguese s.n. [Pelotas, cércas]; Augu: Augusto & Edésio ——— ee ee de Porto hay URUGUAY: Herter s.n. fier Glaziou (1911) sitar his nos. 11330, 11331, 1sh, 14159, and 21899a from Minas Gerais, Brazil » all as L. geminata except no. wash which he identifies as L. EE lag aaa ~~ Schauer oe. cites cultivated specimens from Paris as auth- entic material of Lantana mollissima Desf. and of Lippia asperi- folia A, Rich., deposited in the DeCandolle Herbarium at Geneva; Herb. Willdenow 11512 & 11611 as authentic mterial of Lantana lavandulacea W Willd. and Lippia citrata Cham., respectively, in the Willdenow Herbarium at Berlin; Wei, Weigelt s.n. from Surinam; Ber- landier 2306 from Mexico; and Poeppig s.n. s.n. from Cuba. Also, under L. geminata, he cites Bacle S.n. from Buenos Aires; Joannes de Santa ta Barba: Barbara s.n., Sellow s.n., & Gardner s.n. from Rio Grande do Sul; 1 Riedel s.n. from S&o Paulo; Martius s.n. from Bahia; Martius SNe, “Pohl : S.n., Riedel s.n., and Sieber 38.n. from "secundum flumen Amazonum"; Kegel s.n., Perrottet s.n., Schomburgk s.n., and Weigelt SM. from "in omni omni Guiana'; Humboldt & Bonpland s.n. n. from Venezue- 1a; | Dombey s.n. from Peru; Berlandier s.n. from "in oris maritimis", Mexico; Poeppig S-n. from Cuba; and Bertero s.n. from Puerto Rico. Under L. asperifolia he cites Gillies s s8.n. from Cérdoba, Martius 8. n. from | Minas Gerais, Leblond Son. from British Guiana, Humboldt & Bonplend 5.n. from Venezuela, and several African collections. The E. Pe Johnson 50 & 8.n., cited below from Tabasco, cova as 1965 Moldenke, Monograph of Lippia 65 well have been from Yucat4n -- their labels read "Yucatan & Tab- asco"; S. Watson 387b, cited below fram Chiquimula, is labeled "Eastern portion of Verapaz and Chiquimula"; Commerson S.m., cited from Buenos Aires, Argentina, is actually labeled "Buenos Ayres et Montevideo"; and Langlassé 168, cited by me from Michoac&n, c Mexico, may actually have been collected in Guerrero, since its rilla". In all, 788 herbarium specimens, including types or phototypes of many of the names involved, and 22 mounted photographs and il- by me. Citations: TEXAS: Cameron Co.: Bogusch & Molby 1193 (Ur); Clo= wer 297 (Mi), 962 (Mi); Correll & Johnston 17926 (Rf); Correll & Robbins 20956 (Ld); Cory 28309 (N), Siuh5 (Sm); A. M. Davis Son. {Palm Grove, Sept. 'l1] (Au); G. L. Fisher 41011 (Ew, Gg--316087), Son. (Brownsville, aug. 16, 192] (Hp), sn. (Apr. 20, 1941) (au, Au); H. C. Hanson 434 (G); Herb. Univ. Texas s.n. [Bromsville, 1907] (Au, Au); C. L. Lundell 106k (N, Rf); Lundell & Lundell 8685 (G, Id, Mi, Mi, N); Nealley 113 (C, Up--17030); Perkins & Hall 2618 (It); R. Runyon 226 (Au, Au, Au, N, S), 898 [2177] (Rr), 2603 (N), 4361 (Br, N, Ug), sen. (Brownsville, 1930] (Hp); Schott L6 (T); Tharp 1847 (Au, Au); York s.n, (Brownsville, 3-19-1907] (Au, Au). Hidalgo Co.: Clover 907 (N), 113 (Mi, ) ; Correll & Schweinfurth 15661 (Rf); Cory 17 (G), 36095 (G), 36288 (N, N); G. Le Fisher 1096 (W--2261197); A. R. Moldenke 186 (Fg); M. L. Wal- ker 3 (Au). Wharton Co.: J. Ks Small san. (near Wharton, April 1925] (G); small & Wherry 11825 (N). Willacy Co.: M. Cc. Johnston 5325330 (St). Zapata Co.: Clover 962 (N). County undetermined: Havard s.n, [Sept. 188] (Mi). MEXICO: Campeche: Houstoun s.n. [Bailey Herb. neg. no. 5057] (N--photo of type). Chiapas: Breed- dove 6192 (Z); Matuda 5213 (Ld); Nakamura 178 (Mh, N). Colima: Edw. Palmer 1303 (G, N). Guanajuato: Collector undesignated 18 (Q)- Guerrero: Hinton 5720 (N, Rf), 9242 (K, N, N, Rf), 1h02h (a, Mi, N, N, Rf); N. L. H. Krauss 361 (Mm). Jalisco: B4rcena 225 (Me), 54h (Me); Diquet s.n. [Chapala] (N); Galeotti 788 (Br); E. W. Nelson 11h2 (G), 6520 (G, N); Edw. Palmer 33 (C, Ca--192610, G, Pa, Sg--16083), 686 (C, G, Me, Me, Pa); per 11085 (Cm, G, It, Me, Me, Me, Me, Me, Me, N, Vt), s.n. [26 July 1693] (Me). México: Hinton 1162 (a, N, N), 1812 (G, N, N), 4101 (N, N); Mrs. D. H. Sheldon s.n. [winter of 1672-3] (G). Michoacdn: Hinton 1264) (Du-—34585), La, Mi, N, Oa--6687), 13149 (G, Mi, N, x, at S), 13585 (G, N, N, Rf), 13962 (au, G, N, N, Rf); Langlassé 168 (Cb, G). Nayarit: Feddema 576 (Mi); R. S. Ferris 5927 (Du— 185341); M. E. Jones 23230 (Ca--00878, Gg—172662), 23563 (Po— 153921); Mexia B19 (G, G, Gg--1558h9, N), 1929 (Du--195L9L, G, G, Gg-~157098, La, Wi, N). Oaxaca: Conzatti 579 (G); Ghiesbreght s. 66 PHYTOLOGIA Vol. 12, no. 1 n. [Oaxaca] (Ca--322965). San Luis Potos{: V. H. Chase 762 (G, Ur); Crutchfield & Johnston 5135 (Au--177650) 5 N. L L. H. Krauss Krauss 273 (2); W. C. Leavenworth 200 (Ur); Purpus 5289 (Ca--157335, G, N). Sinaloa: | J. Gonzalez Ortega 5650 (Me); Rose, Standley, & Rus- sell 112) (N). Tabasco: Gilly & & & Hernéndez Yolocotad tzi 253 Git 123195, Mi » N); E. P. Johnson 50 (C), sen. (C). Tamaulipas: Ber- landier 2 (Vv), 874 (G, N), 230i (Du--166557, G, M, T); Re M. King 1035 (Nv (N); Edw. Palmer 102 (G, Ge Gg--31156, N); Pringle 1960 © (Br, te Ca——10L919, G, G, Io—38721, Me, Me, Me, Mi, Pa, S, “S, Vt), sen sen, [8 Aug. 1888] (0b—-50727) . Vera Grass Ervendberd 112 (G), 162, in in part (Cb); Medellin 5 (Me); J. V. Santos 2533 (Mi). State undetermin- ed: M. Be Jone Jones 663 [Armeria] (Po—69159) ; Sessé, Mocifio, Castillo, & Maldonado 2195 (Q), 2196 (Q), 2210 (Q, Q, Q), 2215 (N--photo, Q, S--photo, Z--photo), 2216 (Q), 2221 (Q), 2572 (Q). GUATEMALA: Chi- quimula: S$. Watson 367 (G). El Petén: C. L. Lundell 1,232 wre: Escuintla: | 3, 0. a D. Smith ith 2062 (C,:-G). Huehuetenango: Steyerma 50859 (N). “Suchitepéquez: | Steyermark 3547 (F--1058716) . ecapa: AL = Bartlett 360 (N); C. C. Deam C. Dean 360 (G, Mi), 6261 (Ca—-201)28, Mi, N). BRITISH HONDURAS: C. L. Lundell 1819 (Au, Au, F--685351, ny 1138 (F--689),39, Me, Mi, 3), “L232 (F--689)53, Mi); Peck 798 (0, N); Schipp 755 (Ca--65288, F—712290, G, G, Mi, N, S)- ALVADOR: La Libertad: Calderén 1507 (G). Santa Ana: Standley & sali V.3230 (N). NICARAGUA: sees: E. Wall s.n. [Corinto, 15/4/28) (Ew). Managua: A. Garnier 308 (N). Department undeter- mined: Lévy 261 (Cb, Cb); C. Wright $n. [1853-6] (G). COSTA RICA: Alajuela: Brenes 14320 (6), 2169) (N), sen. (N). Guanacaste: H. Pittier 2869 (Br, Br); Tonduz s.n sen. [H. Pittier 13627] (G). San bee pei eS 987 (Br); H. Pittier 344 (Br); M. Valerio 37 (Br). moe H. Pittier 3329 He Colén: Fendler ~ 220 (7 ara CUBA: Havana: Leén 7530 (H » N). Las Villas: Britton, Britton, & Wilson 5192 (N); Ekmai on 16588 (S), 18893 (N, S)3 Re Ae soar’ Ls 4855 (N, N), SU? (N); Jet G. Jac’ aux 8107 (N). Matanzas: A- : ais et 28 (Ha ms) gan soar a Diaz Barreto 17367 (Es); arabia Ha); Ekman 1578 (S); Herb. Martius sone [Macuriges, Apr. 162] (Br); Hioram oran 1,882 (N), 67 2h (Ha); Leon ae (Ha, N, N), 11917 (N), 16740 (Ha, N); vistous 2141; (Ha). Pinar del Rio: Leén 19725 (N); Leén & Alain 22855 (N); Leén, a are Acufla, & Alain 2340) (N); Leén, Victorin, & Alain "19705 (fia). Province un- determined: C. Wright AR) LO [1860-~6l; Her Herb. Sauvalle 1756], in part (Hv), 3157 (S), 3159 [Herb. Sauvalle 1755] (Hv, N, Pa). JAL- AICA: Ne L. Bri Britton 3917 (N); W. Harris 11737 (N), 11911 (Gg 3116, N, 8); Harris & Britton 1 10593 (N); Maxon & Killip 1636 (Ur)j G. R. Proctor 9421 (N); Stearn 675 (Bm). HISPANIOLA: Dominican 1965 Moldenke, Monograph of Lippia 67 Republic: Fuertes 586 (N, S). Haiti: Ekman H.351; (S), H.8282 (S); Leonard & Leonard 4431 (Ca—l39879). PUERTO RICO: Asenjo 30 (As); Barrus xi) (It); Bla Blauner 175 (Cb), s.n. [1852-53] (M); Britton, Britton, . & Brown 6070 ( (N); 1 Eggers 86, | (Br); Sintenis Bind (S)3 J. “ke Stevenson 32h2 (N); Vélez 1) (N). VIEQUES: Shar 228 (Ge—-31163, N). LEEWARD ISLANDS: An: Antigua: Box 1570 = veg Duss 23 (N). Guadeloupe: Duss 3281 (N, N) » 4552, in rt (N). WINDWARD ISLANDS: Grenada: W W. E “E. Broadway 1825 (N). echsatinas Duss 416 (N, N), 452 (N); R R.A 5; Howard "Howard 11718 (N, N). Herb. 8533] (R), Sen. [September 25, 1932] (I, I); Cruger Ben. (Trin. Bot. Gard. Herb. 213, in par +] (R). MARGARITA ISLAND: Je R. Johnston 81 (Ca—16677, N, W—531920); 3 Miller & Johnston 125 (N, W--h17629) . PROVIDENCIA ISLAND: Proctor 3169 (W--1979286) .— WEST INDIES: Island undetermined: Swartz rtz son. (S). COLOMBIA: Ama- zonas: R. E ; Sohutes 8198 (W--1996275); Schultes & Black l6-Lhha (W--1998675) ates tioquia: Gallego sn. [Julio 1946) (Fn, N); Gir- Gir- aldo s.n. rostaat 1945] (Fn—-3023) ; Tomas 955 (N); Toro Toro 315 __ (Fn—-16)6, N). Atl4ntico: Araque Molina & Barkley 19At082 (N, N); ). We Pennell 4143 (W--10)3881). Caqueté: Klug 1637 (N, S). Magda- lena: Pérez rez Arbel4ez 516), (W--16938)8). Putumayo: Klug 1637 (W— 1517962). VENEZUELA : Angostura: Bailey & Bailey 12% (Ba), 123%a (Ba). Apure: Chardon 22 (Ve), 23 (Ve); Humboldt & Bonpland sn. [Macbride photos 39485] (Kr ~-photo, N--photo, N--photo). Aragua Vogl 1538 (N). Marat Holt & Gehriger 1) (N, Ve, W—-1471858) ; F. W. Pennell tas (N); Tamayo 3430 (Ve, Ve). Delta Amacuro: Bor nd, Gillin, & Brown 166 (N, W—1189880). Falcén: Tamayo 729 (Ve—-12768). Fedaral tistrict: Boldingh 1129 (Ut—12855); Burkart 16013 (Ve). Lara: H. Pittier 13118 (N, Fons, W—1h0h18); Saer | 196 (Ve, W—1188507). Yaracuy: Burkart 16500 (Ve). Zulia: Nooquerye 851 (S, W--2282)8), W--23830]9). State undetermined: ocquerys s.n 8.n. [Duaca] (W-- 2282483, W238 3051). BRITISH GUIANA: 5 R.120 (A (Au--165428); Jenman 5450 (C, W--1322921), 5617 (Cc). SURINAM: Co: Collector undesignated T2hd (Le); Collector undesignated Suriname 80 (Ut); Domburg s.n. [Tulleken 180] (Le); | Re ae (Le) ; 321 (Ut) (Ut); H Herb. Acad. Rheno.-Trai. 123 (N); Herb. Coll. PI Sen. (Surinan] (Pa); Hering s.n. (Le); H Hostmann Bs en = mann & K er 565 (S); Lanjouw 8 (Ut); Menge sen. (Br); Pulle (UE)s soeprate Se (ve), IB TUOG stare oe Ge eee (Ut); Versteeg 6h (Ut, Ut), 258 (Ut), 290 (Ut); Weigelt 8.n. [1827] (Br, Br, 8), S.n. (Gt, Gt, Le); Wullschlagel 407 (Br). FRENCH GU- IANA: W. E. Broadway 99 (N); Lemoine 76hh (N); Perrottet s.n. (Mi); 68 PRY-TOLOGIA Vol. 12, now 1 Sagot 69 (S, T). ECUADOR: Bolfvar: F. L. Stevens 22 (Ur). Guayas: Asplund 17605 (S); Dodson & Thien m 1260 (Ca (Ca), L 1281 (E); Fagerlind & Wibom 138a (S), 138b (S); A. S. Hitchcock 20206 - 20206 (N, W--1195201); Mille = (W-1573127). Los Rios: Asplund id 5520 (S); Harling 152 (S), LL9 (S); S. S. White 5679 (Tl). PERU: Hudnuco: Soukup 2231 (W—1876975). La Libertad: Née 107 (Q), 132 (Q). Lambayeque: Angulo 1325 (S). Lima: N. J . Andersson sen. [Lima, 17 Mars 1852] (N, S, S)3 Barranca 30 mor Ridoutt s.n. [Oct. 1939] (Ok); Wilkes s.n. [Lima] (W--72761). Loreto: Asplund 14375 (S, W—222],902); Tessmann 3082 (S); Ll. Williams 1559 (F--612695),. 2340 (W—195220), 1975 (F—62h953, S). San Martin: Allard 2056) (W=1999787), 22176 (W—-2025)01); Ferreyra 10113 (Ss); Ll. Wil- _ liams 5581 (F—-623160), 7325 (F--62h511), 739 ( (F--623609, W—~ 1498343). Department undetermined: J.T. Baldwin Baldwin 4730 [Ramon Cas- tilla) (Mi); Herb. U.S. Expl. Exped, (Wilkes] sn. [Peru] (T). BRAZIL: Amazonas: Poeppig 26h (¥-=212658, V, v). B nee A. Lutz 211 (Hk), 26 (Lz, A ey Ceara: Drouet 2561 (I, 126 ¢ Migaity S, Sp—383]9, Sp). Gofas: Macedo 2612 (N, S), 3280 res 39h3 (s). Maranh4o: Ozimo de garvaihe 6 (N, (N, Sp—3462). Matto Grosso: 0: Fe Ce Hoehne Uom., Rondon 7 (Sp); Kuntze 8.n. [Mattogrosso, VII.92] ay! N, W--701326); Malme 1880 (S), 16 1880a (N, S, W—-1483440), 2787 (S), S.n. [on banks of Rio Cuyab4 near Santo Antonio, July 12, 1903] (S, S). Minas Gerais: Heringer s.n. [Herb. Est. een: café 378) (N, Sp—h56h); Ule s.n. [Herb. us. Rio de Jan. 1,900] (W—- 1199834). Par&: Archer 8266 (Be--12100, N); Black & Ledoux 50- 10292 (Be--61583); Monteiro da Costa }3 (N); Sampaio sen. [Herb. Rio Jan. ores i” +3 Schwacke 2 25t ()s — ase! — comm. Beak 3752 (sm); Ha Tataclace & Pereira 7863 [Herb. ate RS ~~ (Lw); No Nogiri 19 (Gg); Stelifeld 1253 [Herb. Mus. Paran. ] (N). Pernambuco : G. Gardner 1104 (Cb). Rio de Janeiro: patal Marques s.n. [march 1915] 1915) (Je--735) ; M. Kuhlmann s.n, [Par- ahyba do Sul, Dec. 2, 193k] (x, Sp--32170, Sp); Mello Barreto 1365 (Herb. Jard. Bot, Belo Horiz, 23266] (Ja--L5896)3 Seg Segadas-Vienna, Dau, Ormond, Machline, & Lor€do 59 (Ja). Rio Grande do Czemak & BR, & Reinec k 340 0 (Cb); Henz 35351 (N); Joannesa de Santa Bar bara s.n. = Alegre, ae) S.n. (1836) (Br, F--photo, N— photo, Si--photo, Z-~photo); Leite 660 (N); Malme 768 (S, S)} Ram bo 38012 (Nn), h969h (N); Relte Tae (N); Sehnem 3072 (B). Rio Negro: G G. H. H. Tate 130 GD. Santa Catarina: L Lorentz sen. [Con- cepeion del 1 Uruguay) (B); Reitz & Klein 2387 (Gg). Sfo . Paulo: A: S- Costa Serra s.n. (Campinas, June 21, 21, 1935] (K, Sp-—33h0, Sp)5 1965 Moldenke, Monograph of Lippia 69 Je ZT. Lima sen. (Herb. Jard. Bot. Rio Jan. 8998] (N); Luederwald Sen. (N, Sp--20080); B. de Oliveira s.n. [Capital] (Sp--30276, Sp); Pereira 926 (N, Wh, Wh), s.n. (Capital, Jan. 13, 19);2] (Sp~- 46292). State undetermined: Collector undesignated 183 (C); J. photo, N—photo, Si--photo, Z—photo), sen. (Br); Sellow s.n. [Bras- ilia] (Br, Vt); Ule s.n, (Ja——32325). BOLIVIA: El Beni: M. C4r- denas 3761 (S); Fleischmann 107 (S); H. He Rusby 916 (C, Pa, Pr); Werdermann 2410 (E—-999966, S). La Paz: Buchtien 1458 (W-1177996); H. H. Rusby 70h (Mi, W--1232237); 0. E. White 90 (ii, N, Pa, W— 1232269). Santa cruz: T, Herzog 1h29 (Ss). Tarija: R. E. Fries 1425 (S); Pflanz 2093 (W-—-123341). PARAGUAY: Hassler 960 (N, N), 7367 (Ca-—9LL3h2, N, S), 7397 (Ca—9hb3h1, N, S), 11112 (Ga— 930219, Go, N, S, W--1056911); Kuntze s.n, (Concepcion, Nord Para- guay] (N), s.n. [Concepcion] (N); Malme 902 (N, S, S); Edw. Palmer sen. [Paraguay River] (W--l;1139); T, Rojas 06 [Herb. Osten 7896] (ug), 495 (Herb. Hassler 495; Herb. Osten 8326] (Ug), 1406 [Herb. Osten 7898] (S), 187) [Herb. Hort. Parag. 10060; Herb. Osten 13566] (N, Ug), 187ha (Herb. Hort. Parag. 10060a; Herb. Osten 13567] (Ug), 1875 (Herb. Hort. Parag. 10061; Herb. Osten 13563] (Ug), 1875 (Herb. Hort. Parag. 10065a; Herb. Osten 1356)] (Ug), 8326 (Go), S.n. [San Bernardino, June 1915; Herb, Hicken 242; Herb. Osten Sen. (Concepcion, Dec. 1916; Herb. Osten 13563] (S); Sandeman L870 (K). ISLA DEL TORO: Grtmer 1027 [Herb. Osten 23169] (Ug). URUGUAY: Arechavaleta 8 (Mercedes, Feb.] (Ug), 8 [Paso de los Toros, April] (Ug), 1132 (N), s.n. [Pocitos, Buceo, & vic. of Montevideo; Herb. Osten 3753] (Ug); Berro 86 (N); Cabrera 2581 (N, N, N); Castellanos s.n. [Punta Piedras, Nov. 10, 19]6; Herb. Inst. Miguel Lillo 15761] (N) » Sen. [Bella Unién, Jm. 28, 198; Herb. Inst. Miguel Lillo 150h4] (N); Chebataroff s.n. [dept. Rio Negro, Jan. 1939] (Ug—h736); Collector undesignated s.n. (Maldonado, 25 Dec. 188] (Ug, Ug, Ug), sn. (Ug—-5705, Ug, Ug)s Gibert 7 (Ug); Herb. Herter 10053 (N), 95310 (N); Herter 7h7 [Herb. Herter 82833] (B, Ca—32333k, N, N, S, W--134h15S), 7h7a [Herb. Herter 82658] (N), s.n. [Nueva Palmira, April 1926; Herb. Osten Unién, Jan. 28, 1918] (N, Ug—13930); Rosengurtt 5.950 (N), B.1959 (N), B.h232 (Ug--8278); Schroder s.n. [Colonia, Jan. 30, 1922; S); Commerson s.n. [Buenos Ayres et Montevideo] (N); Lefebre s.n. (Ensenada, 1891] (Br); Venturi 117 (S). Chaco: Jorgensen 271 (Herb. Osten 11876] (Ug, W—1055179), sen. [Herb. Mus. Argent. Ci- 70 PERTOLOGQIA Vol. 12, no. 1 enc. Nat. 23944] (N); T. Meyer 135 [Herb. Osten 22658] (Ug), 8790 (N, S); T. Rojas 12082 (S); A. G. Schulz 134 (Herb. Mus. Argent. Cienc. Nat. 6471] (N), 1500 (N), 1501 (N); C. L. Schulz 863 (N); Venturi 39 (W--1043612). Corrientes: Cabrera 11623 (W—-2197996); Ibarrola 59) (W—1934140); Malme sen. [Empredados, 5/1902] (S); Pedersen 1153 (N, W-—2283036), 1906 (N, S, W--2283115); Rufz Hui- dobro 2037 (Gg—l06032, S), 2180 (Ss), 3879 (N), 3888 (N); Rufz Leal 11266 (Ss); G. J. Schwarz 126 (Ca), 250 (Ca), 559 (Ca); Wroth 59 (S). Entre Rfos: T, Meyer 10058 (N), 1040 (N), 10580 (N); x Morel 19), (N); Rodrigo 952 (N); F. A. Roig 1263 [Herb. Rutz Le 18362] (Sm); Scala 2003 (N, N, N, S, Ug), sen- [li-XI-29] (La). Formosa: Ryerdam & Beetle 22993 (Ca—-6522)); I. Morel 19h (Ss), 690 (N, Ur), 1120 (N), 131h (N), 1u91 (w), 1636 (N); Pierotti 1087 (N); Reales 80 (N). Misiones: Ekman 2015 (N, S); Montes 990 (Ca); D. Rodriguez 616 (Ca--3490), 6161 [Herb. Inst. Miguel Lillo 3221] (N), sen. (Herb. Mus, Argent. Cienc. Nat. 23771] (N). Sal- ta: Schreiter s.n, [Bubarcacion, Dec. 20, 1925; Herb. Osten 20678] (Ug). San Juan: Cuezzo 1127 (N). Santa Fé: R. Alvarez 909 (N)s Job 650 (N, N), 18 (N, N); Ragonese s.n. [Herb. Mus. Argent. Ci- enc. Nat. 19776] (N); Rufz Huidobro 3076 (N), 3116 (N), 3230 (N, S), 3292 (N), 3309 (NW), 3343 (N), 3367 (N); Venturi s.n. (Herb. Mus. Argent. Cienc. Nat. 23767] (N). Tucum4n: Lorentz & Hierony- mis 1174 (N); E. Romero s.n, [28/1/9h5] (ca); Venturi 29h (N)- Chica Island [Misiones]: Montes 990 (S). Laguna Oca Island [For- mosa]: T. Rojas 12212 (S). Puentes Island [Entre Rfos]: T. Meyer 10099 (N). PAKISTAN: Fast Bengal: Koelz 10667 (Mi). ‘INDIA: As- sam: Chand 7692 (Mi); Hooker & Thomson 5 (S); Koelz 2292 (Mi). Khasi States: G. Mann 5.n. [Khasia] (Bz--2177;). AUSTRALIA: Queensland: Baron s.n. [Rockhampton] (Sg--16072). CULTIVATED: Argentina: Miers 1206" (W—1343921) . Belgium: Crépin s.n. (Br). ampai tal, Dec. 7, 1942] (Sp--7560); Tiberio de Moraes 1980 (Sf). Co- lombia: Cuatrecasas 2283) (F—-128003]). Costa Rica: Tonduz 6723 (Br, Br, N). Cuba: Acufla 16063 (Es); Elman 1031) (N, S)5 Leén 11917 (Ha); Marie-victorin 21351 (vi) » ell (Vi); Moldenke & Moldenke 1988) (Es, Lg, N, Sm). Dominican Republic: Ekman H.15802 (N, S)- England: Hort. Barclay s.n. [15-6-30] (Cc). France: Herb. Jewett son. [h. P.] (Mi). Germany: Lucae s.n, {[Hort. bot. berol. ee ss ss gent Duss 4552, in part (N). Haiti: Ekman z . S), H.9808 (S); E. C. Leonard 8515 (N). India: Herb. Horte Bot. Calcuttensis s.n. (B2—-21608). oon. Duss 1,765 (N)« 1965 Moldenke, Monograph of Lippia 71 Maryland: W. H. Cowgill 2073 [Koelz 2168; Plant Quarant. 050726] (Ba). Mexico: | Wonderly 52 ( (Mi). Surinam: Kappler 1860 (Cp, Gt, S, Ut); Kegel 12h0 (Gt). . Texas: F. B. Jones 4128 (Fj). Tortue: Leonard & Leonard 15470 (N). Venezuela: Tamayo 1891 (Ve--12756). LOCALITY OF COLLECTION UNDETERMINED: Collector undesignated 85 (Q), 150 [Mastrante] (Q), sen. (Q); wee 99 (Q), 118 (Q), 129 (a); Tweedie | S5.n. (1837] (Cc). MOUNTED ILLUSTRATIONS: “L'Héritier, Stirp. Nov. 1: pl. 12 (S); Pohl, Icon. Plant. Brasil. 302 ( V-~drawing). TPPIA ALBA var. CARTERAE Moldenke, Phytologia 7: 30. 1961. sinlineranny; Moldenke, Phytologia 7: 1:30. 1961; Moldenke, Ré- sumé Sw page 3: 9. 1962; Hocking, Excerpt. Bot. AS: hh. 1962. variety differs from the typical form of the species in haytiie clear-yellow corollas. The type of the variety was collected by Annette rte -- in whose honor it is named -- and Roxana Ferris (no. 3864) in the rs in the ©, bordered with Prosopis, Acacia brandegeana, Jatropha cinerea, Lophocereus schottii, Opuntia, and Bumelia, at _— El pa northeast of San Yavier, at an altitude of 35 me S, C Sier Giganta, south tude "25083.2 oft Fody longitude 111°33' W., on March 17, 1960, and is me. Citations: MEXICO: Baja California: Carter & Ferris 386 (Ca-- 199588--type, Z--isotype) . LIPPIA ALNIFOLIA Schau. in A. DC., Prodr. ll: 588. 187. Synonymy: Lippia ston Mart. & Schau. ex Schau. in Mart., Fl. Bras. 9: 242. 1851. Lippia brasiliensis A. S. Mtiller ex x wol- denke, Known Geogr. Distrib ve Verbenac., [ed. 1], 37 & 95, mud. 1942. Lantana odora Mart. ex Moldenke, Résumé Suppl. "10; 5, 196)7 Bibliography: Schau, » Prodr. 11: 588. 1847; Schau. in +, Fl. Bras. 9: ‘ale. *ia61)" Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 95. 189i; Briq. in Engl. & Prantl, Nat. oe f (3a): 151. ees Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 37 & 95. 192; Moldenke, Alph. List eves Names 13. buts op vero Bol. Soc. Venez. re a 11: 46. 1947; Moldenke, Distrib. Verbena 80 8 189. 19695 Moldenke, Alpe Lint Cit. 3: 691, 712, & rh isles oldenke, REsuné 93, 311, & 160. 1959; 3 Moldenke, Résumé Suppl. 10: 5. 98h. BOOK REVIEW Alma L. Moldenke "Plant Biochemist: ate D. D. Davies, J. Giovanelli, Ap Rees, xit he illust. fF. A. Davis Co. age eee Pennsylvania. Pe $10.50 These authors from "down under" have compiled an excellen constructed text to "supplement existing ones on plant slwadoleey® 6-65 presupposing a background of at least two of modern chemistry. Plant biochemistry can exist as distinct from animal (and protist) biochemistry only in those few and minor cases where the end products and pathways involve botan- ically uniqie ones. Surely students advanced enough to use this text would understand that most reactions and products are common to all living things. As some of the following chapter headings indicate, the book stresses those biochemical pathways and phosphorylation with its coupling to hydrogen transfer, (6) photosynthesis whose basic reaction is the phosphorylation of ADP to ATP cyclically, (7) lipid senennt ies (8) isopentane compounds, (9) alkaloids, (10) amino acid metabolism and (11) nucleic acid and protein synthesis. The chapter on carbohy- drates is probably the best one. markings should have been clearer on p. 169), a limited index, one a well chosen "reading list", - the ryt: "literature cited" which would have been more useful if there were connected text reference numbers used. All Shp abbreviations are identified at the beginning of the book. PHY TOLOGIA Designed to expedite botanical publication Vol. 12 May, 1965 No. 2 CONTENTS DEGENER, O. & I., The Hawaiian genus Neowimmeria (Lobeliaceae). MOLDENKE, H. N., Materials toward a monograph of the genus PURE Ts oe og oo oo a hn COL es wane nes eat aes MAGNE A Legh reviews. 6s ols ee eet 73 Published by Harold N. Moldenke and Alma L. Moldenke ee lo 15 Glenbrook Avenue = Yonkers 5, New York, U.S.A. Bice of this number, $1; per volume, $5.75, in advance, close of volume Winton Porawe, or $6 at close o} JUN 7. 1065 : THE HAWAIIAN GENUS NEOWIMMERIA (LOBELIACEAE) Otto & Isa Degener Thanks to Dr. R. Melville's note (Kew Bull. 19: if te we here belatedly typify the Hawaiian genus Neowimneri species Lobelia grayana E. Wimm. (in Annal, =a Sy he pat 56: 369. 1948) = N. grayana (E. Wimm.) Deg. & Deg. in Fl. Haw. Ky¢, March 29, 1963. The type for the species was collected by the United States Exploring Expedition of 1838-182, and exists in fragnentary condition x the Gray Herbarium of Harvard Univer sity. The above typification of the gemus should legitimize the species names N. costata, N, dunbariae, N. ebrandii, N. ieuca, Ne niihauensis, N. remyi, N. tortuosa and N. Fasoskase, all o of which are new combinations p: proposed oposed by Degener & Degener in the same publication on March 29, 1963. MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. IT Harold N. Moldenke LIPPIA ALNIFOLIA Schau, Tall shrub, glandular—dotted throughout; branches strict, as thick as a goose-quill, subterete, fuscous, strigose-pubescent and scabrous; princi internodes shorter than the leaves; leaves decussate-opposite, ovate or subrotund-elli sea about the size and shape of those of Alnus glutinosa (L.) Gaertn., at bove, pemninerved, coarsely crenate, venose and rugulose, peony Fi rs simple or _subpapillose at the base, Tap ant pom branches; peduncle » 2-- per axil, 2.5 cm. long, thickened upwards, patulous, pubescent; heads 1 ap gs a gh about 8 mn, wide; bractlets membranous, broadly ovate, ) mm. or more in length, short-acuminate at the apex, ¢ Meenas , many-nerved, pubescent, ciliate-margined, loosely imbricate, shorter than the ped long, obcordate-oblo ong, very smooth, fuscous, included by th fruiting-calyx, easily split es 2’ parts. 73 7h PHYTOLOGIA Vol. 12, no. 2 his apparently rare species was based on several collections made by Carl Friedrich Philipp von Martius "In campis graminosis Brasiliae, in prov. Minarum e. gr. ad Munbucas deserti Minarum novarum, Julio, nec non inter frutices montis Serra de Sincora prov. Bahiensis, Octobri", deposited in the Martius Herbarium in the Botanical Museum at Munich. Lippia brasiliensis is based on A. S. Mifller 69 from Vicosa, Escola, Minas Gerais, Brazil, col~ lected on April 18, 1933, and deposited in the Britton Herbariun at the New York Botanic al Garden, gathered for the fungous infec- tion on its leaves. Lantana odora is based on a Martius solseal tion from Minas dered deposited in the Munich herbarium and there photographed by Macbride as a cotype of Lippia alnifolia alnifolia under his type photograph number 20321 The species is placed by Schauer (1852) in Section Zapania Subsection Axilliflorae. He says of it "trita odorem fortem sub- amp tia." In um specimen and 3 pe la photographs of a cotype collection have been examined Citations: BRAZIL: Minas Gerais: Martius ash, {Macbride photo a (It~—photo of cotype, Kr--photo of c: cotype, N- “4. $ enez Verbenac. Cent. oO 1947; Moldenke, Alph, List Invalid Names 1 & 15. 19473 Moldenke, Phytologia 2: 38h z 385. 1947; ieee Bol. Soc. Venez. Cienc. ea 13: 116. 1948; Moldenke, Alph. List Cit. 2: 361, 126, Sil, $58, & 610 (ste), 3s 693, 82 "32, O95, 90%, WS, ‘Sut, (sis, : 1148, 116k, 1208, > i048, 1269, & 1261, ne - & A. i ueldenke, anal; aw "Biol. Mex. “the 9. td hs denke, Known Geogr. Distrib. Verbenac., [ed. 2], 30, 3, 70, 72, & 189. 19119; Moldenke, Phytologia 3: 285. 1950; "yoldenke, Bol Cienc. Nat. 11: 289. 1950; F. rege Deter- : nac. ikea tale Moldenke, Résumé %, 1,8, 67, 71, 80, 84, 312, 113, 316, 1959; Troncoso, Darwiniana 12: 267. 1961; Moldenke, “a Suppl. 82h (196) and 10: 2 & 5. 196h 76 FarreLOGIa Vol. 12, no. 2 Illustrations: Houst., Reliq. pl. 12. 1781; see toahg Fruct. & Sem, Se i: pl. 56, fig. "2. 1788; Gil & Xuarez, Osserv. pl. 5. 1790; Lam., Tabl. Encycl. Méth. Bot. (Illustr.] 3: pl. 539, fig. 1& 2. 1797. herb. woody or somewhat woody below, 7. a mri sprawl- ing, arching, scrambling, or scandent shrub or undershrub, 1--5 m. 9 sometimes 15--20 cm. in diameter; branches elongate, drooping or pendulous to subscandent, light-green, the adult ones subterete, yellowish, and glabrous; branchlets tetragonal, sparsely appressed- strigose and glandular—punctate; leaves decussate-opposite or ter- re- ly angulate; bractlets cuneate~obovate, subequally imbricate in 8-farious fashion, recurved, abruptly cuspidate or broadly very -5 mm. long, equaling bifid, bifariously villous with long hairs on the keel, otherwise shortly white-pubescents corolla small, hypocrateriform, not con- spicuous, varying from pale-green or greenish-white to cream, white, or pale-yellow, its tube very exiguous, the limb hirtellous or pubescent on the outer surface; fruit. paterenie fuscous or black, am oe at the base of the T g ae ot E a 3 3 g ing i gb g o Fe ct cae a a 8 oa e as about 1/2 inch long, the leaves exactly like those of the plants currently passing as L. hemisphaerica Jacq. from Colombia, poe the general habit of the plant seems to be like that of L. alba ory ) M. E. Br. One wonders if Linnaeus was in error in ascribing th type specimen to Houstom es Mexico The type of L. pauciserrata was gathered pi tage Joseph Schlim (no. 27h) at Aguachica, Ocana, Santander do N ot at an altitude of 500 feet, Colombia, in November, between 1816 and 1852 -—- an 1965 Moldenke, Monograph of Lippia 77 isotype in the Delessert Herbarium at the Conservatoire et Jardin Botaniques at Geneva having been photographed by Macbride fm his type photograph number 24657. The type of L. hemisphaerica collected "in sylvaticis regni N ovo-Granatensi pr. a3 of according to Jacquin (1760). sehier (187) npg the branch- lets of L. hemisphaerica as hispid-pilose, which raises the possi- bility of f this name belonging to the synonymy of L. ssestaatie f. hyptoides (Benth.) Moldenke. Perhaps his descriptive term, how- ever, applies to the Ruiz collection from "in Peruvia ad Guaya- qouil" which he also Saeret rather than to Jacquin's type. He says of L. hemisphaerica "Simillima praecedenti [L. diari dant? tamen distincta facilis capitulis obtuse-bracteatis ss haud angulat is, calyce minore brevius villoso, Calyx semilineam longus, acute bi- fidus, villo b na non excedente. Cor. calyce duplo eae pu The nam Lippia fruticosa Kunth, seems to be based on an un-~ numbered soLection made by Gustav Karl Wilhelm Hermann Karsten at Diballa, Rio Hache, and Paes cpg: in the herbarium of the Na~ turhistorisches Museum a The L. floribunda of AIRE nese referred to in the synonymy a- bove, is actually a synonym of Aloysia reichii Moldenke, that of Briquet is Lippia moritzii Turez., Z., while that accredited to "Hort." is Lantana achyranthifolia Desf. Walpers (185) places Dalea lippiastrum Gaertn. in the synonymy of Lippia americana, but ac- cording to — (1823) this name belongs to Microdon ovatus (L.) Choisy in the Selaginaceae. Lippia canned by Briquet (1895) in Subgenus Za~ ia, Section Dipterocalyx. Kunth (1818) states that it is closest to L. hirsuta L. f. The L. hyptoides Benth., often in- cluded in its onymy, is actually the basinym of what is here c spe- cific epithet, as also that of L. hemisphaerica, is somet times ee ris ased, raat Lippia americana seems to inhabit thickets and dry thickets, woods and deep wooded apres saps the edges of woods, goign +t 1665 meters, cieetiig in March and from May to December, fruit from October to January, in June and August. Common names recorded for it are "barelago", "belf{ta", "lippie yee geal oy hémisphérique", "mano de Dios", Noreganito mac ia", "tr Haught describes the nr as "abundant in this rence eee ajira, Colombia] and "abundant on roadsides", while H. a aeen ophytic scrub" in Cajamarc “oDandele (3su7) 8 sa of it: "En los lugares how Sta Antioquia] 78 PHYTOLOGIA Vol. 12, no, 2 y en sitios arenosos se suele encontrar otra forma que llama la atencién por su simetria y por la disposicién regular de sus ca- pitulos al parecer esféricos; es Lippia hemisférica Cham., que tiene sus flores blancas agrupadas en capftulos casi sésiles y La muestra en estudio proviene de Megua y se debe su coleccién al diligente entusiasmo del Rdo. Hno. Elfas mientras es- tuvo en quilla por los affos de 1933." The corollas are described as "cream" on Asplund 7685; as "greenish-white" on Asplund 15733; as "pale-green" on Asplund 15879; as "pale-yellow" on Allen 1094; and as "white on Asplund 15752, Cuatrecasas & Castafieda 24879, Elias 109), Haught 407 & 6726, F. W. Pennell 1206h, and Schlim 27h). It is of interest to note here how Miller (1768) differentiates between L. americana and L. hemisphaerica and what he says of them: "1. Lippia (Americana) arborescens foliis conjugatis oblong Lippia with hemisphaeric heads. The first sort in the country of its native growth, commonly rises to a height of sixteen or a- bout the size of a large gray Pea, in which are many small yellow flowers appearing between the scales, which are succeeded by the seed~vessels, € second sort grows naturally in Carthagena in New Spain, where it rises with shrubby stalks ten or twelve feet high, sending out slender branches toward their top, garnished with oval spear shaped leaves three inches long, ending in acute points, smooth on their upper surface, which are Placed opposite; the foot-stalks of the flowers come out opposite just above the leaves, each sup- a dal head of white flowers which peep out from the scales of the head; these are succeeded by capsules having two cells, including small seeds. "The seeds of the first sort were sent by Dr. Houstoun to sev- eral curious gardens in Europe, where some of the plants have spar 3 they will not live through the winter. e , specially while they are young; but when they have acquired strength, they may be preserved 1965 Moldenke, Monograph of Lippia 79 in a less share of warmth. As the plants advance in their r growth, they should be shifted into larger pots, but this should not be too often repeated; for if they are removed into new pots eve = will be as often as they require; so that when these, and many other exo » are too often removed, they do not ve so well as when they are permitted to fill the pots with their roots. The best time to shift these plants is in April, at which time the tan of the hot-bed should be stirred, and fresh tan mixed i $55 it, to increase the heat. The earth "an which these plants are placed, should be fresh and light, but not too rich." Lamarck (1797) distinguishes the two as follows: L. h hae- rica -- "capitulis hemisphaericis", L. americana — "capitulis ovato-subrotundis", Lam., Tabl. Méth. Bot. 3: 91, pl. 539 (1797) is often cited as "Lam. Ill. 3. pl. 529", but the plate O67, ai referred to is 539. Gaertn., ee & Sem o Pl. ls *966--267 56, fig. 2 (1788) is sometimes erroneously “cited as plate abet erbarium material of this species has been misidentified and agcie tex: in herbaria as "Rubiaceae". On the other hand, the - Andersson 83, A. S. Hitchcock 20111, and Hartweg 686, dis- emi. mies as L. hemisphaerica, are actually L. americana f. hyptoides (Benth.) Bike the last-mentioned collection being the type of the form, and Holway & Holway 807, one & Smith wee, Re & 1518, Pittier 10541, and Killip, Dugand, & Jara- 3833) are L. americana f. pilosa uote, the last- men sontioseee collection being the type of the form, while the Moc~ querys s.n. [Duaca, 1893-9h] rhs sce oy - distributed as this by Troncoso, is a species of Cordia in the Ehretiaceae; the Collector undesignated i and Herb, Mus. Paris 1, distributed as L. flori- bunda by her, are L. moritzii Turez., "while Galeotti 787 is L. pringlei Bri Schauer (1847) cites under L. americana a Linnean specimen from Vera Cruz [doubtless the How toun collection which is the type of the species], Billbery s.n. [doubless an error for "Billberg"] from Cartagena, Colombia, Mutis s.n. from "in regno Novo-Granatensi", and Hartwe 686 from . Guayaquil, sorsegh all except the first deposited in the Berlin herbarium st royed. The last~mentioned collection, however, is the type Seiigatton of f. cides. Under L. hemisphaerica he cites a Jacquin plant from "in sylvis regni Novo-Granatensi pr. Carthagenam" and a Rufz collection from Guayaquil, also in the Berlin herbarium, now de- Stroyed. Pollen of P. H. Allen 109) is preserved in the slide Collection of the marred Unim University herbarium. In all, 63 herbarium specimens and 8 mounted photographs cluding type materiel of most of the names involved, have been examined Cesticca MEXICO: Vera Cruz: Houstoun s.n. [Herb. Linnaeus G. 801, = 1] (Ls--type, N--photo of type, Z——photo of type). PANA- MA: Herrera: P. H. Allen 1094 (G, N, Nj. COLOMBIA: Atldntico: 80 PEYI-T.O0LOGIA Vol. 12, no, 2 Araque Molina, & Barkley 19At017 (N, N); F. W. Pennell 1206) (Ww 11,3339) . Bolivar: Ara Molina & Barkley 19B0016 (N); Billberg (C). Goajira: Cuatrecasas & Castefieda 25518 (Fg); Haught )07 (N, N, W--17093)6). Magdalena: Cuatrecasas & Castafieda 21,879 (2); Espina & Giacometto B.la [Herb. Mus. Yale School Forest. 20986] (¥, N, W--IL6L81L); Haught 3810 (N, S), 3873 (W), 6726 (Hw); H. He Smith 450 [Herb. Hort. Then. 1.625)] (Br, Cm, N, N, S, Vt, W-- 532973). Santander: Araque Molina & Barkley 185318 (N). Santan- der do Norte: Schlim 27) [Macbride photos 24657} (Br, Br, F--photo, Kr—photo, Kr-~photo, N, N—photo, Si—photo, Z--photo). Depart- ment undetermined: Collector undesignated 199 (F~-1337816); F. C. Lelmann 7914 {in Colombia et Ecuador collectae] (W—~1323012). ECUADOR: Guayas: Asplund 768% (S), 15879 (S). El Oro: Asplund 15733 (S), 15752 (S). PERU: Cajamarca: Sandeman dln (kK). partment undetermined: Rufz 783 [Peruvia et Chili] (W--2h9771). LOCALITY OF COLLECTION UNDETERMINED: Collector undesi (Q); Herb. Bot. Mus. Lund. s.n. {ex America meridionali] (Lu); Karsten s.n. [Diballa, Rio Hacha] (V). LIPPIA AMFRICANA f. HYPTOIDES (Benth.) Moldenke, Phytologia h: 292. 1953. Syn Lippia guayaquilensis Turcz., Bull. Soc. Nat. Mosc. 36 (2): 202. 1863. Bibliography: Benth., Pl. Hartw. 122, 1843; Turez., Bull. Soc. Nat. Mose. 36 (2): 202. 1863; Jacks. in Hook. f. & Jacks., Ind. K Known Geogr. Distrib. Verbenac., [ed i}, & 95. 1942; Moldenke, Phytologia 2: 385. 197; Moldenke, Abstr a 272 (1953) and 5: 2h. 195h; Woldenke, Résimé 80, 81, 84, 313, 460, & 461. 1959; Moldenke, Résumé Suppl. 1: 19 & 25, 1959. This form differs from the typical form of the species in having its branches and branchlets rather densely hispid. The plant is a shrub or subshrub, 2-~3 ot 7. ran on both surfaces, short-petiolate, acuminate at the apex, serrate- ong the margins, t t with short white conic tuberc white hairs, strigose on the venation beneath, otherwise glabrous 1965 Moldenke, Monograph of Lippia 81 and smooth, "venulae tamen etiam minores pilis nomullis hinc inde hirsutae", the upper leaves among the inflorescences smaller, ion Hartweg (no. 686) in woods near Guayaquil, Guayas, Ecuador, and is depos- ited in the herbarium of the Royal Botanic Gardens at Kew. The type of L. guayaquilensis was gathered by William Jameson (no. 890) in the ook age of Guayaquil in May of 18)5. Turczaninow describes his plant follows: "Lippia ramis obtuse tetragoni is, su tum pilosulis; capitulis axillaribus pedunculatis }--6 elssets petiolum subaequantibus; bracteis trapezoideis setae ine florum; calycibus dorso bicarinatis; carinis longe ciliatis; Corollae tubo brevi calycem excedente; coccis maturitate seceden- tibus. Lippia asperrifoliae Rich. affinis; at folia vulgo majora integerrima aut vix serrulata The form has been collected ty river valleys, at altitudes from sea-level to 2200 meters, flowering in March, May, and June, fruiting in May. Benthan says *wspecies pluribus notis L. lasio- us al has been misidentified se distributed in herbaria under the names L. americana L., L, alba (Mill.) N. E. Br., L. henisphaerica Jacq., and Lantana sp. ~The subspecific epithet is astern uppercased. herbarium specimens and 1 mounted photograph, in- siestnens type or phototype material of all the names involved, xami have been e ne me. naa Fe ECUADOR: antcdet - Andersson 83 (N, S, S, S); H 6 (N--isotype); A ition 20111 (N, W—1195152); Jameson 390 $6 (W—15LL5h9), 30% (g—photo) “PUNA ISLAND: Eggers 14739 (W1322867). PERU: Piura: Ellenberg 1193 (Ut—-115373b) ; Weberbauer 6013 (W-~1197159), 6346 (F—TN9ELED), LIPPIA AMERICANA f. PILOSA Moldenke, Phytologia h: 292. 1953. Bibliography: Moldenke, Phytologia 4: 266, 267, & 292. 1953; 82 roe F O's O- OD A Vol. 12, no. 2 Moldenke, Biol. Abstr. 27: 3121. 1953; Moldenke, Résumé 67, 80, & This form differs from the typical form of the species in hay- ing its young branches conspicuously long-pilose, the hairs most- ly appressed. The plant is an erect herb, 1—1.6 m. tall, woody below, or a spr shrub to m. tall, "with white, greenish-white, or crean- colored flowers The type of the form was collected by Ellsworth Paine Killip, Armando Dugand ih and Roberto Jaramillo Mejfa (no. 38334) ina ’ deep wooded canyon, at 380--600 meters altitude, Quebrada Cabafla, Haci- enda El Gechase: between Tocaima and Pubenza, Cundinamarca, * Colom bia, on May 8, iol, and is deposited in the Britton Herbarium at the New York Botani cal Garden Common names recorded for the plant are "belfta" and "mano de Dios". Collectors often say “heads white" or "inflorescenc certai in thickets, along trails, at the a of woods, and in deep wood- canyons, ’ at altitudes of 30 to 1075 meters, flowering in May, July, Au ugust, and November to January, fruiting anuary, May, and November, Killip & Smith 14342 is Pang pect piloset Herbarium material has been misidentified and distributed in herbaria under the names L. anorscane L., L. hemisphaerica Jacq., and L, hemispherica Cham, Ina 15 herbarium specimens , inclu- ding the type, SHS stan anletana'uy ans seed eee Atl4ntico: Elias 873 (W--1543601). Boli- Killip & Smith 14238 (nN, pemeceeeh tee > 14342 (N, W--13503%), 1618 Pe SEORSY Cundinamarca: Dugand, & Jaramillo 38334 (tape) Méta: F. Re wosbaca 135 1396 ae ment undetermined: Hi feibecee © lid1 (W—~1067132). VENEZUELA: ahh He pidtsee 105h1 (N, V 1078), 1 W-—-1187218). ECUADOR: Guayas: Asplund is ( 1931119); Holway & Holway 807 (N); Schimpff 12h 33). of is (W--1619 LIPPIA ANGUSTIFOLIA Cham., Linnaea 7: 377. 1832 [not L. angusti- folia stent 190h]). Synonymy: Lippia graminifolia Kuntze, Rev. Gen. Pl. 3 (2): 253, in textu. 7898 Lippia angustifolia Cham. & Schlecht. ex Molden- ke, Alph, List Invalid Names Suppl. 1: 13, in syn. 19k7. Lippia stifolia H.B.K. ex Moldenke, Alph. List Invalid Names Suppl. 1: 13, in syn. 197. Riedelia angustifolia Cham. ex Moldenke, Alph. List Invalid Names Suppl. 1: 19, in syn. 19h7. Lippia = gustifolia Schau. ex Moldenke, Résumé 310, in syn. 1959. Bibliography: Cham., Linnaea 7: 377. 18325 ——. Nom. Bote, ed. 2, 2: 5h. 180; Walp., Repert. Bot. Syst. lh: hh. 18455 shat. in A. DC., Prodr, 11: 588. 1847; Schau. 4 Fl. Bras. 9: 242. 1851; Hiern, Vidensk. Meddel, Kjfbenh. 1877-1878: 99. "18773 1965 Moldenke, Monograph of Lippia 83 Morong, Britton, & Vail, Am. N. Y. Acad. Sci. 7: 196, 1892; acks. in Hook. f, & Jacks., Ind. Kew. 2: 95. 189k; Kuntze, Rev. Gen. Pl, 3 (2): 253. 1898; Brig, in Chod., Bull. Herb. Boiss., 23 - in riq., Conserv. & Jard. Bot. Genév. 7-8: 31). 190k; Herter, Flérula 10. 1930; Mold » Lilloa 5: 19 & 2h. 1940; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], sf; a Bh. & 95. 192; Fl. : 481. 1948; Moldenke, Alph. List Cit. 2: ; 1, 772, 86, 847, 856, 922, & 923 (199) 250, & 1257. 1919; Moldenke, Known Geogr. Distrib. Verbenac., fed. 2}, 80, 99, 10h, & 189. 1949; Moldenke, Phytolo- gia 3: 376. 1950; Stellfeld, Trib. Farmac. 19 (10): 168. 19$1; Angely, Fl. Paran. 7: 12. 1957; Moldenke, Résumé 93, aran 7 & + b - 1961; Troncoso, Bol. 12: 258, 259 Bot. 9: 18. 1961; Troncoso, ana 9, Résuné Suppl. 8: k. 196f, Ti Soc. Argent. 286, 287, & 289, fig. 15. 1961; Moldenke, I, te simple or racemose-branched at the apex; branches few; lower in- shorter than the subtending leaves; leaves decussate-opposite, sessile, the blades lanceolate, 5--6.3 cm. long and 8--10 mm, wide or larger and elongate to 10 cm., a- hes, de bractlets ovate, cuspidate-acuminate at the apex, strigose- Canéscent on the back, closely imbricate, shorter than the corol- la; flowers dioecious; calyx very small, bifid, villous; corolla infundi toward yellow, bular, abo the apex, the lowest lobe of the limb expanded, subcordate, and ~ te; fruit obcordate-subglobose, adhering for half its length to the diphyllous fruiting-calyx, easily splitting; cocci dark and subnitid on the back, the commissural surface oblique and whitish, This species is based on several collections made by Friedrich Sellow "in montibus Serra da Lappa dictis ad praedia Baixis et Vi- but where one was photographed by Macbride as his photograph mm- Per 17h95. Lippia angustifolia Chan. & Schlecht. is based on Du-~ 8h PHYTOLOGIA Vol. 12, no. 2 sén 2334 and L. angustifolia Schau. on Dusén 1 1236a, both from Pa- ran&, Brazil, “and deposited in the United States National Herbar- ium at Washington, but are mere misaccreditions; L. angustifolia H.B.K., another misaccredition, is based o nm Dusén 11 from a rather marshy campo at Desiro Ribas, Sacank “collected on February 17, 1911, and deposited at Stockholm. The name, L. angust ite olia Morong, belongs in the synonymy of L. morongii Kuntze Lippia angustifolia has been found on campos, wet campos » and rather marshy — » Open moist banks and riverb: swamps = ager dcp hes, dry gras a Bey and almost marshy thickets, 00 meters alti tude, flowering from September to April. Schauer (18h 7) places it in his Section Zapania, Subsection Paniculatae. He cites Riedel s.n, and Lund s.n. n. from "in agro Sorocabensi, Dec- embri-Martio", S%o Paulo, and Sellow Sen, and Pohl sen. from "ad fluvios Taboca et Paracatu" in VWinas Gerais. Hiern (1077) also cites lund s.n, from Sorocaba Troncoso (19 (1961) places the species in her Subgenus Lippia, Sec- tion Lippia, Series Paniculatae. She reduces L. bothrioura Briq. to synonymy here and says "Establezco esta sinonomia bas4ndome en el ejemplar Dusén 11365 del Brasil, determinado como L. a ti- folia Cham. y que se se halla en el Herbario de Kew. Un . duplicado del mismo se encuentra en el Darwinion y coincida exactamente con la descripcién de Chamisso. ee dioica. Los ejemplares ex- aminados, Hassler 7471 y Dusén 11365, son 9, en cambio ez los estambres, en cambio no ha visto fruto, lo que me hace pensar que su ejemplar era©. Lippia angustifolia pertenece junto con L. scaposa Briq. a la serie Paniculatae (Sch.) Briq. caracteriza- por por inflorescencias teeter ie Pog _cabesuslas pip totsorege nis o menos racimosas o cimoso raimos She cites: BRAZI Dusén 11365 (K, si). PARAGUAY: Haseler TTL (Cb); T. aus 3627 (Si). ARGENTINA: Misiones: Burkart 119 (Ss. (Si); F. Me Rodriguez 129 (Si). I am, however, ma maintaining Briquet's taxon ——- of which Hassler 771 is the type collection -- as distinct, at least for the time bei. e um material of L. angustifolia has been misidentified and distributed in herbaria under the names L. hieraciifolia SP, and Verbena sp, On the other hand, the Morong Cham, » Lantana s 860, distributed as L. an angustifolia, is actually L. morongii Kuntees ed 43 herbarium specimens and 5 in- all, and 5 mounted photographs, siting type material of all the nanes involved, have been exam- 1965 Moldenke, Monograph of Lippia 85 ined by m Citations: BRAZIL: Minas Gerais: Mosén 970 (S, S); Regnell 111.947 [Uberava, Nov. 29, 1848] (Ss, W--201129), "rr1.9h7 [oal (Caldas, Jan. 7, 1868] (N, S), 97> (S); Sellow s.n, tab inp meres Macbride photos 17495] (Kr—-phote of cotype, ovale’ N—photo of cotyp ype - a (N, S, W—132288),). Paran4: Dusén 123a (N, (N, W—-148 (Herb, Rio de Jan. 1,860) (N, S, W--1199),28) » 11365 ra —— bach 661 [Rambo 36858] (N, Rb); G. J&msson 1236a (S), 1333a 3a (S). Rio Grande do Sul: Rambo 25780 (Rb). S&o Paulo: Riedel & Lund 2074 (N). State ese Har Rio de Jan. 32216 (Ja). PARAGUAY: Fiebrig 468) (cb “N--photo, Z—photo); Hass- ier 9117 (V), 7971 (N, 7 $61)5 Tina 4325 (N, w 200577). ae CENA Corrientes: Ibarrola 3956 (N, (N, S). Misione : Ekman O (Mi, N), 2011 (S), 2012 (N, S); Grttmer 713 [lisa . Osten #55) (Ug); Niederlein son. (Herb. Mus. Nac. Hist. Nat. Buenos Aires 2360] (Ra). LIPPIA Stier Loes. & Moldenke ex Moldenke, Phytologia 2: 19-20. 19, Synonymy : goa antaica Loes. ex Moldenke, Suppl. List Inval- id Names 5, is. Statatets i Mol denke, Phytologia 2: 19—20. 29115 Pegg Suppl. List Invalid Names 5 191; Moldenke inte aoe _* Verbenac., [ed. 1], 35, 80, & 95. 19h2; Moldenk ieee valid Names 29. 192; Moldenke, Alph. List cit. "hs 305 (pkey ae hk: 1023. 1949; Moldenke, Known G eogr. Distrib. Verbenac., [ed. 2], 73 & 189. 1949; E. J. Salisb., Ind. Kew. Suppl. 11: 138. 1953; Moldenke, nies 8h, 310, & 460. 1959. Shrub, 2m. anches slender, apparently Sroeh and virgate, ee Steams, brownish, op Peau weaice outs principal internodes 2-—3 cm. long; et annu- ripe leaves decussate-opposite, eae 3 petioles se Shier 1— long or obsolete, densely s St Spee. eaf-blades fimiy chartaceous or subcoriaceous (when mature), flabelliform or subrotund, obovate when immature, to abort 2 om. long and wide base), regularly and uniformly crenate-serrate from the base to the apex with rounded teeth, the margins revolute, very scabrous and bullate above, puberulen nt on the yagi densely pubescent se many-flowered, about 8 mm. long and 10 mm. wide; brac tlets ons, 2.5—3 mm. long, subacute at the apex, densely short-pubescent; 86 PHYTOLOGIA Vol. 12, no. 2 i h--5 mm. long, its limb about 3 m. wid he type of this rare species was collected by August Weber- pee oet or (ae. 218) below Limatambo, at an altitude of 2300--21,00 » cuze co, Peru , in June, 1911, and is deposited in the herbar- Fen ey the Chisago Natural History Museum. The species has been collected at altitudes of 2100 to 21,00 meters, flowering Ap. and June. In all, 2 herbarium speci- mens and 2 mounted photographs, 7 ag type material of both names involved, have been examined by m Citations: PERU: Cuzco: Vargas 929 (it); Weberbauer 5918 (F—— 62884,3—type, N--photo of type, Z--photo of type). LIPPIA ARECHAVALETAE Moldenke ex Herter, Revist. Sudam. Bot. h: ¥2} Bot. 7: 225. 19433 Moldenke, Lilloa 10: 342 & 377. 19h: ueadenk, ph. List Ci Moldenke, Phyto 3%. 1 Moldenke, Bol. Soc. Venez. Cienc. Nat. ll: 6. 19475 He in 4 H L. Lilloa 14: 0. 1948; oldenke, t Cit. 2: 357, a 457, ~ 532 (1918), 3: 780, o75, B76, & t 528 ste), ‘ ae 1166. 1949; Moldenke, Known Geo Dis Verbenac 0, 100, &l "k 199; Sopanaee Lilloa 20: 137, 136, & 17h. 193; Sstelifeld. Trib’ Farmae. 19 (10): 168. 194; Moldenke, Phytologia ly: or. 1953; E. J. Salisb., . Kew, Suppl. ae 1953; Angely, Fl. Paran. 7: 12. 1957; Moldenke, Résumé 93, » 119, & 460, 1959; Angely, Fl. Paran. 16: 59 (1960) and 17: "ie 1961; Troncoso, Bol. Soc. Argent. Bot. 9: bine 1961; Moldenke, Résumé Su Suppl. 10: 4. 196. erenni herb, woody at the base, or low subshrub; stems signee densely puberulent with very minute appressed hairs and parallel, impressed above; inflorescence axilla » abundant, solitary in each axil, decussate-opposite, erect, 1--3.5 cm. long, surpassing ves le puberulent and strigose; bractlets densely glandular-—pubescent; atten soaps corolla yellow or golden-yellow to orange or The type of this distinctive species was eae ta José Arechavaleta (no. 11) — in whose honor it was te COEtY) Tacuarembo, Uruguay, in November and December, se, eat” is de- posited in the Delessert Herbarium at the Conservatoire et Jardin 1965 Moldenke, Monograph of Lippia 87 Botaniques at Geneva. The species has oe collected in rocky pastures, arroyos, and dry clearings, on rocky hills, stones and shrubbery, and on c campos, at altitudes of *500 to 1026 calls The flowers are described as "yellow and red" or “orange, yel- low, and red" on on Rosengurtt B.3921, Veottusalions on Herter 60165, and "yellow" on Berro 3007, 3007, 3008, 5786, & 7480, Hatschbach = Herb. Osten 3756, and Smith & Klein 11485. erbarium material of this spe species has been misidentified and aicocane: in herbaria under the names L. pseudo-thea Schau. and Ver Verbena ar echavaletae Osten. A por on Osten 3255 reads "cf, L. intermedia Cham. sed flor. axillare In all, 27 herbarium specimens a 2 mounted photographs, in- cluding the t type, have been examined by me. Citations: BRAZIL: Paran4: Batechbach 3370 (Z); Hertel 6 (Herb. Mus. Paran. 1960] (Mp). ta Catarina: Smith & Klein 1485 (W—22h9377) ; Smith, Ear A & Sufridini 9278 (W—22],9358). State undetermined: Sellow Sen. [Brasilia] (V (Vt). PARAGUAY: Fie- brig 6493 (W—1159378). URUGUAY: Se 11 [Briquet 23] (Cb—-type type, Cb—isotype, N--isotype, N--phot aM, tape, Ug—isotype, Ug--isotype, U g--isotype, Z-—photo of a « (Herd. Osten 3756] (Ug); Berro 3007 (N), 3008 (N), 5786 ay “Fido (N), 7481 (N); Chebataroff | s.n. [San Javier, Jan. 1939] (Ug—-4697); Collec- tor undesignated s.n. [Arenales de San Javier, near the Rio Uru- guay, Feb. 190] Taes-£866); Herter 60165 (N); Osten 3255 (Ug); Rosengurtt ae ae » B.3921 (N, N, Si). ARGENTINA: Entre Rfos: Herb. Hert Herter 4491 (N) LIPPIA eRe cu var. MICROPHYLLA Moldenke, Lloydia 13: 223. 1950 Bibliography: Moldenke, Lloydia 13: 223. 29505 Moldenke, Phy- tologia. 3: 376. 1950; Moldenke, Résumé 93 & 1959. variet; having its leaves unifomly malier, ee Ergo Ape SS m. ong mm. wid T © of the variety was collected by Padre Balduino Rambo (no, 2751) in dry thickets a Morro dos one near So Leopoldo, Rio Grande do Sul, Brazil, September 9, 1936, and is deposited in the herbarium of the Colegio inchiota, at Porto Alegre, Brazil. pe far the taxon i S know only from the type collection. _ all, 2 herb. arium Wm specinens, ee rie type, and mounted phot SPaphs have b een examined y m Citations: BRAZIL: Rio Grande do Sul: Rambo 2751 (E—photo of F--photo of type, N—isotype, N——photo of type, Rb—type, Z~-photo of type). 88 PHYTOLOGIA Vol. 12, no. 2 LIPPIA ASPERRIMA Cham., Linnaea 7: 215--217. 1832, Synonymy: Lippia asperrima var. asperrima Moldenke, Résumé 310, in syn. 1959. Bibliography: Linnaea 7: 215--217. 1832; Steud. Bot., ed. 2, 4 ee 1810; D. Dietr., Syn. Pl. 3: £97. 183, Walp, Repert. Bot. S = m LS 1845; Schau. in A. DC., Prodr. ces 1847; Morren, Belg. Hort. 1: 18). seo: Schau. in sler, e s5 86r. eA 190); Briq. in Chod. & Hassler, Pl. Hsseler. 2 "2 Gis" tg 193. 1908; Te Herzog, Meded. Rijksherb. Leiden 29: hh vist. S e 7: 225 33 Rosengurtt, Estud. Prad. Nat. Urug. 3: 2356 19,3; Moldenke, Lilloa - 342 (1944) and 12: es 1946; Moldenke, Alph. List Cit. 1: 12, 27, 39, 51, 95, 201, 26h. 1946; Moldenke, Phytolo- gia 2: 308 & 386 (1947) and Pi tet, 1948; Moldenke, Lilloa 1h: 4O—}1, 43, Lh, & wee Sige Moldenke, Alph. List Cit. 2: 376, 31, 585, 587, 598-600, 627, & 628 (198) 3: 3: 105, 1732 133, 3, ts, viele st, 875, 910, 911, 913, & 967 (1949), and 4: 1088; 1237, "1256, 1257, & 1302. 1949; Noldenke, ee ae ae erbenac., [ed. 2], 80, 97, 99, 100, 10h & 189. 1919; Rosengurtt, Lilloa 20: 132, 137, & 17h. 1949; Mo ldenke, Phytologia 3: 290. 1950; Stellfeld, Trib. Farmac. 19 (10): 168. igs, Moldenke, Inform. Mold. . Set & le Spec. 2. 195h; 2 ely, Moldenke, hate 93, 114, 16, 125, 10, & her 1959; Angely, - Paran ee 59 (1960)’ and 17: rte. 1961; Troncoso, arwiniana erennial glutinous herb, often rough, creeping and suffrutes- cent, or at ite gp or ’ shrub, (@) 4 ks - tall, with a wo xylopodiun ts woody, thick; stems herbaceous above, erect, fastigiate? pubtotnigey, glandular-hirtellous and viscid, the hairs short, patent, ghee rigid, and anduli ferous; _prineipal » axillary heads subglobose, involucrate, 8—10 mm, wide, convex during anthe~ sis, many-flowered, ovoid in in fruit; bractlets ovate, acuminate at the apex, loosely imbricate, half equaling the head in involucrate fashion at the base, subequaling the corolla~tube, strigose; eg very Micah corolla hypocrateriform, from y low, clear-yellow, or yellow. orange, to rosy~yellow, orange, °F "yellow and red", yi tube villosulous at the apex, the limb vil- 1965 Moldenke, Monograph of Lippia 89 losulous on the back, very finely glistening hairy above, unequal- ly l-fid, the lobes sho rt, Sea ate-obtuse, the throat villosu- lous; fruit hidden among the qually long bractlets » pubescent, easily splitting into often Bis per Si nutlets. Th of this common species was collected by Friedrich Sel- low "e Brasilia tropica" and was deposited in the herbarium of the Botanisches Museum at Berlin, now destroyed. Schauer OBB: places the species in his Section Zapania, pmo; on Axillifl t on rocky and grassy slopes at Arroyo Tres Arboles. In his 199 work he says of it "campestre oes? principalmente en pedregalesY A. G. Schulz says about his n . 8689: "flor amarilla, luego anaranjada y finalmente rojiza. H “ig muchas flores sin caliz; otras con un solo sépalo y otras las m4s escasas, con 2 sépalos ph and | "bosquecillos ralos en suelo arcilloso, duro; flor amarillo; luego anaranjado y ap i caliz de sépalos libres, mu- Chos flores sin caliz." For his no. 8692 he says: "Flores amar- illas; pasada la antesis son cnn te luego rojizas" and "bosquecillos ralos; suelo arcilloso BE, pent 3 flor amarillo luego anaranjado y rojizo; caliz grande, entero." Venturi 2h32 has the leaves extra long and narrow. For horticulturists, interested in color variants, the follow- ar lLlow" G range" — Hassler 6340; "yellow-orange" -- Anisits 2252; "yellow and red" — Venturi 2432; and "yellow" — R. Mf erro 1968, Burkart ee Herb. Osten 12177 & pn. W. Hoehne “2662, 2503], rei ah 3152, Moldenke & Moldenke 1977, Osorio ca te- 1947] & sen. [Cerro Tob: Tobatinga, Feb. ree Ns ATED pele 6568, ie 179, & 9180, Rambo 3040, Rosengurtt B.3002, Schreiter 5 5082 & 8.n. [Muflecas, Nov. a C. L. Schulz Amite 1010, spare 1238 1238, Terribile 381, Venturi 7359, and J. West 6285. Ragonese, Darwiniana 5: 13 Za (gt) = refereaoe ce in the biblio~ graphy above is sometimes a cited as Santa Fe (191) as T. Herzog, Meded. Rijksherb. rat iss 25 TER AGas\ ie douse thes 90 PHYTOLOGIA Vol. 12, no. 2 ted as "Herzog, Bolivia III (1916) 44"; and Briq. in Chod. & Hassler, Pl. Hassler. 2 (11): 493 (1904) tae Hasslerianae XI (190)) el 93" 6 Ih originally inscribed "Peru", but this was later crossed out. Briquet (190) compares his Lb. trachyphylla with the present species as follows: "Espéce tres caracterisée » voisine du L. as- Cham. dont elle différe par ses feuilles ovées et fone lées, ses tiges rampantes diffusement rameuses, ses pédon plus c courts, a indument strigoso-applique 4 glandes cexpaieia trés courtes et localisées sous les ep vag corolles d'un rose- violace (jaunes dans le L. asperrima) e Herbarium material of L L. asperrim asperrima fai ee misidentified and distributed in herbaria under the names L. asperifolia Rich., "Le aff. asperrima Cham," » L. imbricata Kuntze, L. recolletae Morong, L. reptans Kunth, L. turnerifolia Cham., L. turnerifolia Cham, & Schlecht., L. turnerifolia var. camporum Griseb., amd even Mimosa bimucronata var. hexandra (the last-ment ioned probably a case ge of label-crossing). On the other hand, Venturi 531, 1375, & 7359, distributed as — asperrima, are L. contermina B: Briq.; Herb. . Osten 18188, Herb. Mus. Argent. C: ey Nat. 16270, D. Rodriguez ]09, Te Rojas jas "28h, and R Rosengurtt B.3303 are L. morongii Kuntze; Kuntze; and ate Inst. Bot. S. S. Paulo 23346 and F. Cs , Hoehne | son. (villa Vel- ha, P Ponta Grossa, Nov. 3, 1925] are tL. ila Cham. Schauer (1851) cites Riedel s.n. and Sellow s.n, from "In prov. S. Pauli et Matto Grosso pascuis hm humilis e. gr gre pre pr. Camapuan, Santa Fé, th while Rosengurtt (199) cites his B33 — the eeuren ioned, however, is actually L. morongii. Troncoso (1961) te Hassler 7456 as this species, even though Briquet (190) had regarded it as a cunie of his L. phaeocephala, which Troncoso maintains that it is In all, 151 herbarium Ghanian me 2 mounted photographs of L. asp errima have been examined Tabi ition: BRAZIL: Parand: a(S). Rio Grande do Sul: Bornmtfller 206 (C, Cb, v--6631}79 3 see Sen. [Cerro Tobatinga, Feb. 2h, 1947] (Tg—1h21) 5 Rambo 300 (Rb); Swallen 7687 (We-1933118) « S#o Paulo: W. Hoehne 2662 (Bh, N). BOLIVIA: Cochabamba: M. C denas 5292 (W--2250812). Santa Cruz: Steinbach 2712 (N, Ra-—33057)- re tee undetermined: D'Orbigny 11h6 [Herb. Reichenb. f. 127452] N). PARAGUAY: Anisits 1925 (S), 2252 (s); Fiebrig 4777 (Cb)5 Hassler 6348 (Cc a~-94,337, N, N, S), 9312 (N, $aphote, V, Z—photo), 11590 (Ca—93025h, Go, N, s, W--1086991); J Jorgensen 3775 [Herb. Osten 22239] (Du--197838, Ms S, Ug, W—1)83630), s.n. [Herb. Mus. Argent. Cienc. Nat. 28/; (N); Kuntze s.n. [Std Paraguay, IX 92] (N); Osten 9178 (ug), 3179 (S), 9180 THerb. Osten 9012] (s), 9182 (Ug); T. 1 Te Rojas 12967 G, | Ur), s.n. [San Bernardino; Hassler 1965 Moldenke, Monograph of Lippia 91 2403; Herb. Hicken 21; Herb. Osten 7539] (Ug). URUGUAY: Arecha- valeta 1 (Cb), 15 (Ug), 16 (Cb, Ug), 17 (Cb, Cb, Ug, Ug); H. H. Bactlets 21030 (W--2320218); Berro 1968 (N); Gallinal, Aragone, , Campal, & Rosengurtt ~ 1923 (We-17i2788); Legrand 2007 RES 2012 ~ (Ug), ” 3869 (Ug); Osorio s.n. [Valle Eden, Feb. 18, 1947) (U (Ug—-13839) ; Osten 3321 ee ~ 6565 (Ug); Ros rtt B.2356 (N), B.3002 (N); Rosengurtt & allinal 5730 (Spa); Schroder ay tates Ysabel; Herb. 0: Herb. ate 16592] (Ug). ARGENTINA: Chaco: Re i 468 (N); JOrgensen 2631, in part [Herb. Inst. Mig- os: Lillo 321 273, Herb. Osten 11916) (1 (N, Ug); Malme s.n. [Las Pal- mas, 20/3/1903] (S); T. Meyer 267) (N); A. G. Schulz az 1497 (N), 2916 (N), 8689 (Cb, Hk), 8692 (Cb, Hk); C. , L. Schmalz 1010 (N); Venturi 36 (W—103610). Corrientes: Ibarrola 1300 (8), 227 (N), = a3 Os M)» 3883 (N), 4348 (N, S). psi Rios: Be 9 [Herb Ug). Formosa: Jtrgens in part = 4 Osten 169) (oe); <. —— 96 (N) 7B Gr 9), 1s (N), 834 (N), 1237 (N), 1288 (N, Vi). Jujuy: E. K. Ball B.5931 (Ca—603618); Kuntze s.n. [Jujuy, Bet: 1892] (N, etta 8 eas & Moldenke 18745 (N); J. West 6285 (Ca-—561723). Misiones: 3: Bertoni 191 Th9l (Ca); Ek- Ek- man 1997 (N, S), 2013 (S); Grtmer 701 [Herb. Osten 23168] (Ug); Montes 225 (Ss), 1008 (N, Ur); G. J. Schwarz 783 (Ca); Vattuone & Bianchi 1.28 (W-=101,3492) « Salta: Job 1543 (N); T. Meyer 3452 [Herb. Inst. Miguel Lillo 35569] (En, N); Moldenke & -Moldenke 19747 ~ Lg, N, Sm); O'Donell 4648 (N); Pierotti 219 (N), 1232 (Ca); ; Ringualet 115 (N)3 J. Te ". Roig 15427 15427 (Bs); Schreiter 5062 pales Osten ten 20647] (Ug), 3.n. [Cerro San Lorenzo, Oct. 18, 1925; Herb, Mus. Argent. Cienc. Nat. 26/1342] (N); A. G. Schulz 6 ase arre 1176 (S), 1238 (S); Wall & Sparre s.n. n. [4/12/k6) (Ew), 5.n. AEABT t (Ew, Ew, Bw). Santa Fé: Castel lance sen. [Herb. Mus. sls ent. Cienc. Nat. 1961] (N); Feddersen s.n. [Aurelia, near S. Fé} (Cp); Job 801 (N); Kuntze s.n. [Ceres, X92] ns Ruiz Leal Us2h5 (Rl); Terribile 381 (N), 122 (N). Tucunén: kley & Garcfa 128 (N); Moldenke & Moldenke 19716 (N), 19755 (Es, Es, Ig, §); Monet ti 1038 [Herb. Inst. Miguel Lillo 32176) (N); Rufz Leal 12157 (Rl); Schreiter see [Mufiecas, Nov. 15, 1917] (Ug), Sens , [Mutiecas, Dec. ae 1923; Herb. Inst. Miguel Lillo 3799] (N, Ug—L9bh); Ven- ane 232 [Herb. ste 17235] (Ug, W--1591257, W--1591258), 2627 Gan ios, W-1591261), 361s (W--1591277); Wall & Sparre son. [207i hs (Ew), son. [2h/11/06) LIPPIA ASPERRIMA var. LONGIPEDUNCULATA Moldenke, Lilloa 5: k2l. 19h0. Bibliography: Kuntz — Gen. gn 3 (2): ath 2 Thiselt .- r, Ind. Kew. Pi 2s << my oa 5: 421 & 423 (1940) and 8: A ss 19h2; enke, Kn nei hone opr Verbenac., fed. 1], ho, 1, 43, & 95. erry Gote. Revist. Sudam. Bot. 7: 225. 92 PATO LOGIS Vol. 12, no. 2 p 3 + 689, “693, “859, snore ‘ “OLS We and a bs ae 19103 Moldenke, Phytologia 3: 7h. ee er own Geogr Verbenac., [ed I "80, 97, 99, agit a 90, 19193 iosenmertt. Lilloa 20: iz, 137, & 17h. 19193 uoldenke, | my ae 93, ei 116, ig, 126, 310, & 461. 1959; Moldenke, Résumé 10: 3—5, This variety differs from the typical form of the species in that its peduncles far exceed the leaves prints shred ng being or 0 The type of this variety was collected by José Arechavaleta (no. 12) at Salto, Uruguay, in September, 1898, and is deposited in the Delessert Herbarium at the Conservatoire et Jardin Botan- iques at Geneva. The bibliography of this taxon is inextricably mixed with that of L. imbricata Kuntze, with which this plant was for a long time confused. Seca examination of Kuntze's type, however, deposited in the Britton Herbarium at the New York Botanical Garden and col- hotographed by Mac bride as his type photograph no. "11516, has convinced me that it is conspecific with the widespre d Lantana achyranthifolia Desf. On the other hand, the cihvent cited as Lippia imbricata in Lilloa 5: 23 ayes 10: 378 (194k), and 20: Eas (1949), and Holmbergia h: 150 (1945), and in my Alph, List Cit. [see biblio- bove) -- with the exe oridana Kuntze. The Herb. Inst. S. Paulo 5421 and Rosen- gurtt & Gallinal 5730, distributed and cited as L. imbricata, are typical L. asper: asperrima Cham. The Arechavaleta 1 is; “distributed and cited by me in my Alph. List Cit. 1: 12 (isk6), is actually L morongii Kuntze, The Castellanos s.n. [Florida, 2.11 .19h6; Herb. Inst. Miguel Lillo 15773], Son. [Artigas, 30.1.1948; Herb. Inst. Miguel Lillo 15764], & s.n. [Santa Ana do Livramento, 2h.II-19h7; Herb. Inst. Miguel Lillo 15763], Gallinal, Aragone , Ber rgalli, Campal, # Rosengurtt PE.5171, Legrand 38 3489, and Rosengurtt B.4 582, Stated bated and cited by me as L. imbricata, are all Le all L. villaflor- The variety has been found on dry rocky sunny campos, in spiny 1965 Moldenke, Monograph of Lippia 93 grassy meadows, in fields, and adventive especially a~- ee roadsides, at altitudes of 60--80 meters, flowering in April, si gales", "frecuente en los campos, sobre las mérgen ine Gel 1 Rio Uru- guay", "en los campos rocosos", “sobre las lomas se ae oot tas de pasto a lo largo del Rfo Yi", "comin en pra crytéfitas", and "frecuente en laderas", Schulz dete ant his no. 1499 "planta abundante; pelos no glandulosos; terrenos arcilloso- haléfilos, duros y secos." The corolla is described as "orange" on A. G. Schulz 199, as "yellow with red spots" on Rosengurtt B. B.225h, and as "yellow" on Anisits 1876, Rodrigo 60), and Rosen- gurtt B.4210. The Rosengurtt, Estud. Prad. Nat. Urug. 3: 235 geo): cited in the bibliography above boxe a ited as "Rosengurtt, Flora de Palleros ( 193) 2 Herbarium material has ode misidentified and distributed in herbaria under the names L. imbricata Kuntze, L. turnerifolia Cham., and "Phyla nodiflora var." On the other 7 hand » Arechava- ieta 15, distributed as L. asperrima var. longipedunculata, lata, is reset L. morongii Kuntze. In all, 17 herbarium aes apolmere the type, and 2 mounted a have xamine Citations: PARAGUAY: anisite 1876 (S). URUGUAY : Arechavaleta 12 (Cb~-type ‘ Cb--isotype, N--i N--isotype, N--photo of isotype, N-- ae of type, Ug--isotype, Ug-isctype) Gallina = srege galli, Campal, & Rosengurtt 1616 (N); Herb. Mus, Hist. Nat. Mon- tev. 8607 (Ug), 5605 (Ug); Herter 2033a (Herb. Herter rter 50765] (8); Rose! Rosengurtt B.2180 (N, N), B.2254 (N), B-4210 (N). ARGENTINA: Chaco: T. 1 T. Meyer leyer 113 (Herb. Osten SS } (Ug); A. G. Schulz 1199 (N). Corrientes: A A. P. Rodrigo 60h (N). LIPPIA ASESAILTA var. ROTUNDATA Moldenke, Phytologia 1: )66--l,67. 1 9 Biblio jowrenton Moldenke, Phytologia 1: Rageaaeel (1940) and 1: 50h. 19h1; denke, Known Geogr. Distrib. Verbenac. 37 & 95. isha, Moldenke, sph. List Cit. 2: eI & 552. 1948; Miol- Geogr. Distrib. Verbenac., » 80& ihe » 19L9; oldenke, Phytologia 3: 287 & a0. 1950; ha Trib. Farmac 19 ci "168. eoegee ely, aran. 7: 12. 1957; Moldenke, RE ene 125, « bei. is 1959; ee Fl. Paran. 16: aes 960) and this + having its leaf-blades obtuse or rounded at the apex. It Srennial herbs petioles 1--); mm. long, margined; leaf-blades oid liptic, 1 8-347 em. long, 1.2--2.) cm. wide, regularly s pais almost to t he base with biunt teeth, acute at the base an longed into the short petiole; venation decidedly Fai Ser: Only the midrib and secondaries slightly prominulous beneath; 9 PHYTOLOGIA Vol. 12, no. 2 peduncles 3--3.5 cm. long. The branchlets are densely g pubescent with short dark hairs and are also copiously hirsute poe white spreading hairs twice as long, less densely so in age. he hairs on the upper surface of the leaves apparently become bulbous-based in age, first along the margins of the leaves, but final ver the whole lamina, but the upper leaf-surface is scabrous ihe ee times. The type of this variety was collected by Frederico Carlos Hoehne at Ponta Grossa, Paran4, Brazil, on November 1, 1928, and is deposited in the herbarium of the Instituto Botanico at So Paulo, Brazil. The plant has been found in rocky places, flower- ing in October and November. In all, 3 herbarium specimens, including the type, and 2 moun- ted photographs have been examined by me. Citations: BRAZIL: Paran4: F. C. Hoehne s.n. [Ponta Grossa, Nov. 1, 1928] (N--isotype, N—photo of type, Sp--23268—type, Z-- photo of type). ARGENTINA: Misiones: Grter 935 (Herb. Osten 2317k] (Ug). LIPPIA BALANSAE ene Ann. Conserv. & Jard. Bot. Genév. 7-8: 305--306. Synonymy : eas “ealliclada Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8: 306--307. 190h. Lantana jorgenseni Moldenke, Lilloa 5: 2-413. 190. ae jorgenseni Moldenke, Suppl. List In- valid Names 5, in Ey ee Bibliography: Briq., Ann. Conserv. & Jard. Bot. Genav. 7-8: 305-—-307. 190k; ments dn Chod. & Hassler, Pl. Hassler. 2 (10): 490. 190k; Prain, Ind. Kew. Suppl. 3: 10h. 1908; Moldenke, Lilloa Moldenke, Alph. List “ages ge Names 31. 192; Moldenke, Lilloa 0: 423. 1942; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 4o, ki, oh, & 7s 1325 Moldenke, Alph. List Cit. 1: 26 & 27. ghé Mo bBo Bb HF He Ne & Ac Le surge ae ri ulent or short-pubescent, especially when young, the pubescence longer and denser at the nodes; middle internodes 2--l cm. long; leaves medium-sized, fragrant or very aromatic, with the odor of witch-hazel (ameelta) when éetamed. decussate-opposite or ter- nate, subsessile or short-petiolate; petiole slender, 2--5.5 mm. long, rather saasely short~pubescent with crisped hairs; leaf- 1965 Moldenke, Monograph of Lippia 95 blades rather nee and heavy, firm-textured or subcoriaceous, sordid-green above, paler beneath, broadly ovate to elliptic or mm. long and 0.2—-1 mm, apart), na coarctate as well as ent at the base, often cuneately n rate the petiole, short-pilose and very scab- e beneath; midrib, secondaries, and veinlet reticulation ae above, prominent beneath; inflorescence axillary, capi- more or less racemosely grouped toward the apex of the branches; 3 peduncles very slender, 0.8-~3 cm. long, subequaling or slightly surpassing the subtending leaves, tetragonal, short- ent with etre t: or somewhat cri hairs or sparsely puberulent, varying to glabrescent, sometimes sparsely glandulose, 6--8 per nodes heads small, ——— » about the size of a large pea (Pisum sativum), 5-6 mm. long, 6-8 mm, wide, many-flowered, conser ait Se asiee among the beactletas bractlets small, eolate or ovate-elongate to ovate-acuminate, 2--l, mm. long, acute or subacute at the re age lightly or densely short—pubescent with bromish hairs on outer surface, glabrescent on the ner surface, the lower ee scarcely s sing the flowers, the minute, Q-1i oro teh ne lobulate, exalate, antrorsely pubescent on the outer surface; corolla hypocrateriform, pale-yellow to rose, pale-rose, or pinkish th surpassi e calyx only its tube exserted, puberulent aap escent he the limb un- equally l-lobed, the posterior lobe longer, 0.7--0.8 mm. long, the rest seers 0.5 mm. long; stamens and pistil included, nor= f e genus; nutlets small, ovate, 1--2 mm. long, hard, ae — @ type of this mieten species was collected fe eepnict Balanse (ne 1033) -- in whose honor it was na named -- a Th Jorgenseni was collected by Pedro Jorgensen “ (no. ce — in whose age it was named -—— at Villa Rica, Paraguay, in October, 1930, and is Lea aaeyaigay in the Britton Herbarium at the New York Bo- tanteal srt ah Aatontven the plant as very common; Archer notes that this. pant could easily serve as basis of extraction of an essen- 96 PHYTOLOGIA Vol. 12, no, 2 tial oil". It has been collected in anthesis in October and from December to February. The flowers are described as "rose" on Archer 826 and ae 6735, as "pinkish-rose" on Balansa 1033, and as "pale-yell » pale-rose, or pink" on Jtrgensen STs Briquet (190)) eesti the venation of the leaves of his L. ° us valde minulo" and the fruit as "mericarpio parva dura a, cum calicis i adpresse prorsus pubescentis laciniis secedentis" and s of it "Cette espéce se plac ae prés du L. rubiginosa Schauer...dont sl. pailis da ee says: "Le L. calliclada est fort voisin du b. paloma Nous aurions été tenté d'y voir une variété re- marquable, si l'absence de formes intermédiaires ne s'opposait, au moins dans 1! état actuel de nos connaissances, 4 cette réduc- lansae par seabed elliptiqes, cunéiformes et plus nettement pétiolées 4 laissent aucun doute sur leurs vraies affinités." He describes its leaf-venation as Pliage medio pennato, omnibus t¢ reticulescen- tibus sed laminam parum rugantibus" and the fruit as tcum calice ete: extus pilis Tenet prorsus versis vel patulis obtecta sece- Troncoso (1961) says "Especie my caracter{stica por sus hojas generalmente ternadas y capftulos pequefios, subesféricos, peduncu- vi i po de Paris pero no me fue posible dibujarlo. Reproduzco el fototipo del Chicago Nat. Mus. No. 2645. L. calliclada Briq., segin su autor es una especie muy afin a 1 . balansae. Considero que se trata de la misma especie, Los caracteres que da Briquet para dif- erenciarla, forma y tamafio de las hojas y capfitulos a veces en ed e se tratan de variaciones dentro de e. The Briq. in Chod. & Hassler, ~ Hassler. 2 (10): 490 (190) | reference cited above is sometimes bid 3 E (190) 90" or as "Briq., Plant. beclawea 2: 490." barium material of this taxon has been misidentified and dis- herb under the name L, germinata H.B.K. On the other hand, the Jtrgensen 2h71 and Harb. Osten 11876, distributed 5 rpet : t f 1965 Moldenke, Monograph of Lippia 97 as L. balansae, are actually L. alba (Mill.) N. E. 6h Troncoso (1961) cites the Following specimens, seen by me, from Paraguay: Balansa 1033 Bernie ; *wasslar b735 (cb, P), 8653 (Bm, Cb, P), 12h8 121,8), inten » Bm, Cb); J0rgensen ensen 3778 ( (P, Si); T. | T. Rojas 4826 (si), 48 4893 (Si), 9211 (si). In a all, 1, 2h herbarium specimens and 3 mounted photographs, cluding type or add material of most of the names pe ere have been examined Citations: BRAZIL: Minas Gerais: Heringer 3835 (Z). PARAGUAY: Archer 1679 (W--1705198), 4826 Se wn 07362)4 Bale Balansa 1033 [Macbride photos 2645] (It—-photo of ty of type, N—photo of type); Hassler 4608 (N), 6738 (C (ca--91L338,, N,N, S, * 7-260), 8653 (Ca--929819, N, N, “N, V—1209), 1248 (Ca~-929699, N, hla Jorgensen 3778 (Herb. Osten 22236] (Du—197832, It, N, N g, W—1453813), s.n. [Villarica; Herb. Mus. Argent. Cienc. et4 287 2070] (N). LIPPIA BAUMII Gttrke in Warb., Kunene-Sambesi Exped. 350. 1903. Synonymy: Lippia asperifolia Hochst. ex Moldenke, Alph. List Invalid Names Suppl. 1: aaj in — 1947 [not L. esperi falta Benth., agit nor H.B.K., 1863, li , 1832, nor Reichenb., 1828, nor A. Rich., 1801, nor L. "Ce R , 1959]. pibliogra phy: Gtirke in Warb., Kunen cote Exped. 350. 1903; Prain, Ind. Kew. Suppl. 3: 10h. "1908; Moldenke, Known Geogr. Dis- b, * Verbe: enac pees ra a & 95. 1525 Moldenke, Phytologia 2: oe Distrib. Verbenac ie 2 9 Ab & 189. foligs i Wolent, Alph. List Cit. ahi 996 & hs. 1919; Moldenke, Résumé i, Us 1h7, 152, 310, & 61. Shrub} to 0.5 m. $l; "h hirt Seermpennests branches elongate, divaricate; 3 leaves Bictasate-eppoaite. .t ssile, lanceo- late, to 6 cm. long and 2 cm. wide (the upper ones Sonal Aekebly smal le er), rather obtuse at the apex, serrate along the margins, ni ott a he on both surfaces with rather abe appressed TS, Canescent beneath; inflorescence axillary in the upper Pb esi cy extrao: ordinarily long-pedunculate; peduncles very slen- to 11 cm. long, solitary, often twice as long as the subten- ves, hirtellous-pubescent; bractlets closely imbricate, ermost ovate-lanceolate, to 12 mm. long - wide, acumin- ate at the apex, decreasing in size and e dth toward the apex of the s, the uppermost or lanceolate- linear, 3--5 mm. long’ 1m. , acute at the apex; calyx 1. mm. long, 2-parted almost to the base, not alate, the lobes broad- ong ly ovate, acute at the apex, strigose-hirsute on the outer surface; — yellow, its tube 5 mm. long, 3--l; times longer than t The type of this species was collected by H. Baum (no, 515) — in whose honor it was named -- in sandy soil at 1200 meters alti- 98 PHYTOLOGIA Vol. 12, no, 2 tude between Ungombekike on the Rubango and Kuito, in Southwest rica, on December 9, 1899, and was deposited in the herbarium the See ches Museum at Berlin, now destroyed. The name, L. asperifolia Hochst., appears to be based on Baum 250, also from Southwest Africa, deposited in the herbarium of the he Jardin Botan- ique de 1'Etat at Brussels. The L. asperifolia of H.B.K., of Poeppig, and of A. Richard are synonyms of L. alba (Mill. ) Ne Ee Br., that of Bentham and of Reichenbach are Phyla scaberrima (A. L. Juss.) Moldenke, and that of L. C. Richard is Lippia javanica Burm. f.) Spreng. Gtirke (1903) says of this plant: "Die Art gehtrt zur Untergat- ung Zapania (Scop.) Benth. § Euzapania Briq., Gruppe der Axilli- rere (Schauer) Briq. Sie ist verwandt mit der weit verbreiteten Lippia asperifolia Rich., obwohl sie habituell gar keine Aehnlich- keit mit ihr hat. Die neue Art hat einen auffallend sparrigen Wuchs mit weit abste henden Zweigen, eed Behaarung und sehr lang gestielte pitig tad. auch betr> die Linge des Kelches nur 1/3 derjenigen der Blumenkro ” The species has beni collected at altitudes of 1200 to 2185 meters, flowering in October and December, fruiting in October. Herbariun material has been tiisidentified and distributed in her- baria under the names L. asperifolia A. Rich. and L. wilmsii H. earson. In all, 4 herbarium specimens and 5 mounted ———e in- — type material of both names involved, have been examined Citations: TANGANYIKA: Haarer 2263 (Br). KENYA: Thorold 2733 (Br). SOUTHWEST AFRICA: Baum 250 (Br (Br), 515 (Br--isotype, type, F-—photo of isotype, N--photo of isotype, N—photo of isotype, Si—photo of isotype, Z—photo of isotype ay; PPIA BELLATULA Moldenke, Résumé Suppl. 10: 3 & 5 (196k), Phyto- logia 10: 170. 196k. Synonymy: Lippia bicolor Mart. & Schau. ex Schau. in A. We, Prodr. 11: 587. 187 [not L L. bicolor Kunth & Bouché, 185, nor Kunth, yes sippis bicolor Mart. ex Moldenke, Résumé Suppl. 10: ry in syn. Bibliograpiy: "punt & Bouché, Ind. Sem. Hort. Berol. 1845: 10- 1815; Schau. in A. DC., Prodr. 1): 587. 1847; Schau. in Mart., Geo 89 Alph. Li Cit. 3: 691 & 711. 19h9; Moldenke, Résumé 93 & 61. 1959; Molden- mpd Peter sumé Suppl. 10: 3 & 5. 196; Moldenke, Phytologia 10: 170. Shrub, to 1 m, tall; stems squarrose, branched; branchlets sub- tet onal, hirtous; Leaves small, varying from decussate-opposite or more or less approximate to ternate, sessile, variable in size and shape, ovate or subrotund to subreniform, h--8 mm. long, + 1965 Moldenke, Monograph of Lippia 99 mm, wide, rigid, patulous, acute at the apex, crenate alo yn the strigose-hispidulous above and along the revolute eee. , hirtous-toment ose and som t+ canescent beneath; inflore cen @ axillary; heads subsessile, solitary, 5- or 6- flowered wae eare Py about if aggregate at the apex vl the babe sr ad conglo-~ bate, subinvolucrate; bractlets herbace rigid, vate, sub- acuminate at the apex, loosely anrtaucl, iasphantous, about half as long as the corolla-tube; calyx hardly more than 1 mm. long, ifid, hirsute, the lobes somewhat kideetates corolla rose, yellow in the throat, pubescent on the outer surface, its tube straight, cylindric, about ); m. long, subventricose at * the middle, the limb somewhat larger, the lobes rotund; fruit medium in size, subglobose, dark, obscured by the mature calyx; cocci subequal, separating with difficulty. The type of this rare species was collected nA Carl Friedrich Philipp von Martius (no. 1983) in fields at Villa do Rio de Contas, Bahia, Brazil, and is deposited in the ety of the Botan- isches Museum ’ at Munich, where it was photographed by Macbride as his type photograph number 20322. The L. bicolor of Kunth & Bou- orae. It has been collected in anthesis in October and is knom thus far only from the type collection, of which 3 mounted photographs have been examined by me. Citations: BRAZIL: Bahia: Martius 1983 [Macbride photos 20322) (It—-photo of type, Kr--photo of type, N--photo of type). LIPPIA BOCAINIENSIS Glaz., Bull. Soc. Bot. France 58, Mém. 3: 51, ll. Bibliography: Glaz., Bull. . Bot. France 58, ° 3: et rt by Ind. Kew. * Suppl. ri 153. 1921; 3, Fg strib. Verbenac., [ed. 1], 37 & 95 “(a9h2) and [ed.'2}, 80 189. 1; ictaenie, Alph. List Cit. h: 1300. 199; Moldenke, Résumé & 959 Nothing is know to me of this supposed species except what little is said about it in the gia sa port where it is recorded as "L. bocainiensis Glaz., n. sp. ?" and the type is sta- ted to be Glaziou 11340 from "Campos ae Bocaina, au Sobrado, ane Aofit- septembre. C", oye specimens deposited in the herbaria at Paris, Berlin, Kew, Copenhagen, and elsewhere, the Paris one prob- vo to be regarded a the actual type; the Berlin one is now de- BOLIVIANA Rusby, Mem. Torrey Bot. Club k: 243—2hbh. 1895. Bibliography He Hs Rusby, Men. Torrey Bot. Club h: 243—2hh. & cks. K Conserv. & Jard. Bot. Genev. 7-8: 305. 190k; H. H. Ru Y. Bot. Gard - Ls 432. 19075 Hayek in Engl, Bot. tat. Gees 1908; Wataenk Lilloa 5: 22. (29405 F. L. Herrera, Sinop. Cuzco 1: 31. igh1; Moldenke, Geogr. Distrib. Terbveriac., {ed. 100 PHYTOLOGIA Vol. 12, no. 2 1], LO & 95. 192; Moldenke Lilloa 10: 378 (1944) and 12: 148. 1946; Moldenke, Alph. List st Cit. 1 28, 92, & 201. 19h65 Moldenke, gaan 2: 386. 19L7; , Lillo oa 14: Wl. 19185 Moizenke, - List Ci 108, & 1293. i993 Moldenke, Bhd Geogr. Distrib. "yore { ; Oh, & 189. 1949; He N. & As ee nk gaan Biol. Mex. 20: 9. 19l9; Moldenke, Résumé 125, & "fi6le 1 5555 A. Maria, Pl. Vall. Coch. 38. 1962; ae Résumé Suppl. 3: 13. 1962. nst. tall, loose and spreading in growth; wide, round apex, obscurely serrate toward the apex with blunt thick and rigid teeth, lelctwatetok, cuneate eis the base and subpetio- late, rugose and papillose above, sas ae ticulate oben peduncles axillary, solitary, very slen Sgn - long, papil- lose; , flattened, about “foag! gal). et gone bractlets closely appressed, oval-obovate, strongly c cute at the thr oy ciliate; calyx ovoid, nearly 2 mm. note “gatid, densely hi the teeth minutely notched; corolla hypocrateri- form or infinaibalar, white to whitish to yellowish-white or pink- white, about mm. long, pubescent, the limb ];-lobed, the upper lobe deeply cleft, the lower obovate, longer than broa vary O- al the’ stigma lateral, not so large as in Lantana achyranthifolia Desf.; mature fruit not seen. The type of this species was collected by Miguel Bang (no. 979) in the vicinity of Cochabamba, Cochabamba, Bolivia, in n 1891, and is deposited in the Columbia University herbarium at ba mo York Botanical Garden. The original publication is cited Instituto Miguel Lillo as "Rusby, Bolivia Memoirs IV (1895) ) a" and Dr. Rusby's 1907 publication as "Rusby, Bolivia (1907) 432." The specific name is often uppercased. The plant has been found in rocky soil, stony soil among cacti, dry bush country, and exposed declivities in the mountains, on — and hillsides, bare stony open hillsides, and rocky slopes, mts, de hojas lan eoladas diminutas, atrae la atencién y se pre senta agradable ie vista por su penacho de flores blancas dimin utas. Es bast pp comin en las laderas del Tunari y Alalay" * and cites his no. 175/3a. oer agp material has been misidentified and distributes the s L. suffruticosa (Griseb.) Hieron. and Aloisia sp. 0 the Aas hand, the Ruiz Leal & Roig 1889, distributed as L- bo- 1965 Moldenke, Monograph of Lippia 101 liviana, is actually the very closely related L. integrifolia (rise. ) sig n. of Argentina. The rs are described as "pink-white" on R. E. Fries 1222, as oof egeieegen to white" on Eyerdam 21,885, and as "white" on Balls 6296, Cérdenas 76, 2378, & . 276, | Kuntze s.n., and T. Meyer Lsh7. aa all, 2% herbarium specimens, including the type, have been e b Gatet fone, P PERU: Cuzco: F. L. Herrera 153) (W--1342313). BOL- IVIA: i Asplund 1166 (Ss, Us); Balls 6296 (W—-177783h) ; M. Bang 979 (C--type, Pa- wii tha M. C4rdenas 7h 7Th6 (W--17h,3896), 2378 (W Cae 2876 (W-~1877089) ; | Eyerdam yerdam 21885 (Ca—65837h, Ew); Kuntze s.n. [Rio Tapacari, 19 Maerz 1692] (N); A. Maria 175 (2); Troll 1391 (B). Potosf: Fiebrig 3111 [Herb. Osten 15223] (s, SP, Sp—3350h, | Ug, W--1178002). Tarija: R R. E. Fries 1222 (S, S). Province undetermined: M. Bang 2531 (C, Pa, W--1322890); aie S.n. [Parotani, 20 Maerz 1992] (N). ARGENTINA: Salta: T. 157 7 (Ca—906130, N). San Juan: H. H. Bartlett 20558 (Ge T7500). LIPPIA hiaigetarca Briq., Bull. Herb. Boiss., sér. 2, h: 1162. 190 Bibliography: Briq., Bull. Herb. Boiss., sér. 2, : 1162. 190k; Inds in Chod. & sb cpea Pl. Hassler. 2 (in): 496 ag Brain, _ Mate Suppl. 3: 1 oh *1908; Moldenke, Known Geo [ed. 1], bi & 95. 4942; Moldenke, Lilloa 8: 723, (1942) Ly 10; 378. 194; Moldenke, Alph. List Cit. 1: 26h. ae Molden- ke, Phytologia 2: 337. 197; jMoldenke, Lilloa 1s: 41. 1948; Mol- denke, Alph. List Cit. 2: 488 (1948) and 3: 692 & 911. 1949; Mol- denke, Known Geogr. Distrib. vere) {ed. 2], 99, 100, & 189. & ase, antrorsely strigose or tr et Re midrib and ies culate, prominent be- heath and often somewhat impressed aber; inflorescence terminal, nore elonga ate to 10 mm. "long an and 2 mm. wide, thickening er an- thesis; bracts or bract-like leaves lanceolate, more or less can- 102 Patt OLoe rs Vol. 12, no. 2 escent, shorter than the peduncles; bractlets canescent on the outer scar ry securely 2-lipped, repandly l-denticulate, pubescent apex on the outer surface; corolla hypoc raterifarm, yellow, pubes- cent above, exserted 2--2.5 mm. from the calyx mouth, its tu ne 0.5 mm. long; stamens and pistil normal for the genus, included; cocci minute, 0.6 mm. long and 0.5 mm. wide — with the canescent mature calyx segments, smooth, excavated on the inner surface, The type of this species was collected by Emil Hassler (no. 7471) in a swamp in the region of the upper Rfo Apa, in northern Paraguay, in November, between 1885 and 1902, and is deposited in the Set gboect Herbarium at the Conservatoire et Jardin Botaniques at Geneva, where it was photographed by Macbride as his type photograph ‘2 2646. Briquet ( medio et nervis lateralibus acroscopis [=macroscopis? } sande re- ticulescentibus subtus prominulis, paginem superiorem saepe alia. ent " rt oides Chan., L. oxycnemis Schauer et L. herbacea Mart. Les deux premiers s'écartent inmédiatenent par leurs aaa eye strobilacés et le dernier par les capitules tétragones-ovoides, outre de breux autres caracteres. Dans le L. bothrioura, les capitules s'allongent un certain temps par le | sommet, ale leur forme gén- pole ne change aa he les fleurs et fruits tombent 4 la “partie Les organ a un litrement creusé de fossettes 1 pagel (d'ou le nom spécifique bothrioura), plus volumineux que le pédoncule, d'un aspect fort élégant." The Chod. & Hassler, Pl. Hassler. 2 (11): 498 (1904) reference Listed above is cited at ie Instituto Miguel Lillo "Plantae Hasslerianae XI Aber 498 The species has been wing oad i in swamps and on the banks of rivers, flowering pw November to January. The Schroder col- Troncoso (1961) rotaie. 2 the species to synonymy under L. angus eee Cham., but it seems sufficiently distinct to me. 1m all, 5 herbarium specimens, including t; material, and mounted photographs have been e Lag tations: PARAGUAY: Hassler 771 [Macbride photos 246h6] (C@~ 9h4340--isotype, It—photo of t of type, | Kr——photo of type, N--isotyPe, 1965 Moldenke, Monograph of Lippia 103 N--photo of type, S—-isotype). URUGUAY: Berro 4757 (N); Schro- der s.n. [San Gregorio, Jan. 1926; Herb. Osten 1 94,36] (Ug). LIPPIA BRACTEATA Carr., Rev. Hortic. 53: 380. 1881 [not L. brac- teata Clausen, 1 i7]. ae Synonym nymy : iipota bracteata Hort. ex Jacks. in Hook. f. & Jacks., Ind. Kew, 2: 95. 189). Bibliography: Carr., Rev. Hortic. 53: 380 & 78. — Jacks, in Hook. f. & Jacks., Ind. Kew. 2: 95. 189k; Moldenk Known Geogr. Distrib. Verbenac., [ed. iy, 73 & 95 (1942) and fed. 4, 162 & 189. 19,9; on Résumé 220 & 461. 1959; Moldenke, Résumé ppl. 10: h& 5. Nothing is 20 to ‘me of this taxon except what is stated in the original description: ee hehe, alge fortement villeuse, laineuse sur toutes ses parties erbacées. Rameaux en ib sieeus: a face pty Space tomenteuse par des 7 pels argentés laineux. "Inflorescence capitiforme rappelant celle des Lantana, subsphérique, au sanmet d'un pédoncule d'environs 5 centim ingtres. de longueur; bractées nombreuses, peoreereees accrescentes, ses- ou viol eurs par quatre divisions gece? trés-petites, ovales arrondies, d'un rouge brique ou fauve. Cultivée avec soin’ et soumise a des pincements appropriés, cette espéce pourrait devenir une belle Plante d'ornament pour les serres d'abord, puis pour les jardines, ou, pat en ee terre pendent 1'é6té, elie fleurirait contim- ellemen chaude, serre tempérée ou a -h sui qu'on ne re avoir des fleurs pendant l'hiver The article in which this description appears is not signed, but the editor-in-chief of the journal pet ate h it appears was at that time Elie Abel Carriare, so it seems likely that the name Should be accredited to him. The L. Scaskeane, of Clausen is a Synonym of L, iupulina Cham, LIPPIA scape (Mart. & Gal.) Moldenke, Phytologia 2: ~ orl Synonymy: Lantana bracteosa Mart. & Gal., - Acad. Brux., sér. 2 il te ll (2): 326. 1844. Lippia nutans ose Seah? bien. Am. Journ. Sci. 150 [ser. 3, 50]: 162. 1695. Lippia bracteosa Mart. & Gal. ex P. C. Standl., Contrib. U. S. Nat. Herb. 23: 12h6, in syn., sphalm. 192). Lippia nutans Bob Greenm. ex Moldenke, Ré- 196k. Bibliography: Mart. & Gal., Bull. Acad. Roy. _— _ ce | (2): 326. 18h; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 28. Gr 953 Durant & Jacks., Ind. Kew. Suppl. 1: 250. 1903; P. C. Standl., Contrib. U. S. Nat. Herb. 23: 12hh & 12h6. 192k; Moldenke, Prelim. Alph, oe Invalid Names 30. 190; Moldenke, Known Geogr. Distrib. bie » (ed. 1), 17 & 96. 19425 Moldenke’ Phytologia 2: 226 & 10 PHYTOLOGIA Vol. 12, no. 2 330. tdi, we pas al Alph. List Invalid Names 11, 13, & 15. be Molde t cit. 2: 393 & 473 (1948), 3: 829, 830, 63h 335, & 872 (1919), and k: 1211 & 1294. is Moldenke, Known Ge- ogr. Distrib. Verbenac., [ed. 2], 31 & 189. 199; E. J. Salisb., petiolate; petioles to mm. long, canaliculate and hispid-pubes- cent above; leaf-blades thickish, ovate-oblong or ovate, about 2.5 cm. long and 1.7 cm. wide, acutish at the apex, cuneate at the base, serrate along the margins, rugose or iene prsatigl , hispidulous and slightly shiny above, paler densely pubescent or canescent-villous beneath; tes slender, ab os 2 iF =) @ nate & 5 co 5 p: @ i A age upper leaf-axils, commonly recurved or smOeING, erage fo - ipeas3 rge, thin, pale, ic ceous or scarious, 8—12 m. long, 4--6 mm. wide, aa at. the seer reticulate, glandular- pubescent on both surfaces, ciliate, e lowest ovate, the upper more elliptic; flowers s Sieh oaaesetinte: about 7 m. "long; ¢ densely pubescent; corolla yellowish, shorter than the bractlets. The type of this distinctive species was collected by Henri Guillaume Galeotti (no. 761) on calcareous mountains at Tehuacan we las Granadas, at 6500 feet altitude, in the Cordillera of ~ ca, Mexico, in August, 180, and is deposited in the herba = the Jardin Botanique de 1'Btat at Brussels. The type of be mitane was gam by Cyrus Guernsey Pringle (no. 5650) in Hoyas Canyon, at 1500 feet altitude, Oaxaca, Mexico, in etait 18h, an _ is Bet Reon deposited in the Gray Herbarium of Harvard versit Martens ‘& Galeotti (18h) make this comment: "Species nostra bracteis magnis imbricatis quae cum bracteis coni Humuli lupuli L. similitudinem quamdam gerunt a caeteris speciebus facile dis- tinguitur." The species has been collected in dry bushwoods and on the dry Soe on canyons, at altitudes - 1000 to 2665 meters, flowering d fruiting in February, July to September, and November. us 10116 and 10360 could have been collected either in Chiapas or Jalisco -- the labels carry both state names. In my 1940 and 19h2 publications I regarded Lantana bracteosa as synonymous with Lip- pia callicarpaefolia, but this was in error. Herbarium material has been misidentified and distributed in herbaria as L- calli- carpaefolia H.B.K. and L. cardiostegia Benth. In all, 27 herbarium 5} Bogle niy and mounted photographs, in- ores type material of all the names involved, have been exal~ e me. Citations: MEXICO: Chiapas: M. C. Carlson 1648 (Mi); Purpus 10146 (Ca--278031, S). Jalisco: Purpus 10360 (N, Po—1)6)21). 1965 Moldenke, Monograph of Lippia 105 Oaxaca: Ss Conzatti Be wh), 42h8 (Me); Galeotti 761 (Br--type, Rabi f type, N--photo of type, Si—photo of type ype); "eisnets 5650 A Me, Vt), eee Me, Me, Me, Mi, Mm—151,08, S, Vt); Rose & Rose 11368 (N). Puebla: Purpus 2570 (Ca—139667, N), 3926 (Ca-~1 39666) « LIPPIA BRADEI Moldenke ae Phytologia 3: 61--62. 19h9. Bibliography: Moldenke, Phytologia 3: 61—62 & 7h. 1949; Mol- denke, Known Geogr. Distrib. Verbenac. [ed. 2], 80 & 189. aes E. J. Salisb., Ind. Kew. Suppl. 11: 138. 1953; Moldenke, Résum 93 & 461. 1959. Shrub; br anchlets slender, obtusely tetragonal, densely short- pubescent with erect glandular-capitate hairs, the brownish bark splitting irregularly into longitudinal fissures; nodes rather obscurely annulate; principal internodes 1--2.5 cm. long; leaves decussate-opposite; petioles very slender, about ho ta ’ 2 sing divergent in fruit; 3 peduncles slender, Sis, elongate to cm. in fruit, densely EyAnen tic pabesceht like the branchlets; heads hemispheric, 1--1.5 cm. wide, densely many-flowered; bractlets large, elliptic, about “C1 m. long and 1.5 mm. wide, “densel glan dular-pubescent, subacute at the apex; th mn lar- Pr tO, ML B deposited in ae Britton Herbarium at the New York Botanical Gar= ed o from mounted hoes otypes have been examined by m Citations: BRAZIL: Minas Gerais: eae, eo Mello Barreto, & Brade 3369 [Herb. Jard. Bot. Rio Jan. 40009] (F-—photo of type, type, W. N-~photo of type, Sg—-photo of type, Z—-photo of type). LIPPIA BURTONII J. G. Baker in Thiselt.-Dyer, Fl. Trop. Afr. 5: 281. 1900. 106 PHYTOLOGIA eal Biblio J. G. Baker in Thiselt.-Dyer, ope 3 281. isoor es hum, in Just, Bot. Jahresber. Qe (a): tse. "19085 Thiselt.-Dyer, Ind. Kew. suppl. 2: 106. 190); Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 49 & 95. 1942; H. i. tk Moldenke, Pl. Life 2: 52. 1918; Moidenke, Known wines oo Verbenac., [ed. 2], 115 & 189. 199; Moldenke, Phytol : 29 & 293. 1950; wordenke, Résumé 12, 148, 426, & 461. 19595 Molden- ke, Résumé Suppl. Undershrub; Srenenlece woody, ee slightly bristly; leaves sessile, ternate, oblong, crenate along like eath le node, very s pedunculate, globose, about - in diameter; Drace yee ovate, acuminate a The type et this species was collected by Sir Richard Francis Burto ose honor it was named -~ at Mondanda in the Re- public of aa Leopoldville in September, 1863, and is deposited in the herbarium of the Royal Botanic Gardens at Kew. The species h. Deschamps says of it: "plante an- nuelle, eur 0 em., foutiage vert clair, fleurs mauves ” in- florescence, Usage: sert dans les maladies des yeux." Quarre 1370 is placed here tentative Herbarium material has been misidentified and distributed in herbaria under the names L. asperifolia Rich., L. ukambensis Vat- ke, and Lantana salvifolia Jacq. On the other hand, the Peter 2229, distributed as L. burtonii, and 2873 as "Lippia n. sp. aff. L. L. burtoniin, are both L. Tavenica (Burm. f.) Spreng. Baker T1900) separates the 8 tropical African species of this ems known to him more - a as follows: 1. Bractlets orbicular, 2e les very eae lL oatesii Rolfe. 2a. Peduncles long L. radula J. G. Baker. la. giclee cuspidate, 3 k. area orbiculax or obovate L. somalensis a ha. Leaves oblong, Tugose....L. javanica (Burm. f.) Spreng lb. Bractlets acuminate. 5. Leaves opposite -L. ukambensis Vatke. Sa... “Le burtonii i J. G. Baker. le. Bractlets acute hadi 2 " plicata J. Ge Baker. all, 6 herbarium specimens, g pte the type, and 9 moun- ted photographs have been examined Citations: NIGERIA: Dalziel 696 fh-ochote, K, N, N—photo, SE~ photo, $1-~photo, Z--photo). CONGO LEOPOLDVILIE: R. F F. Burton 8» fi. (Sept. 1863} (F~-photo of bee K--type, N--photo ‘of type, type, Sé— Photo of type, Z--photo of type); L . Deschamps LS (Br); Quarré 1370 (br). TANGANYIKA: Peter 51781 (0.1.05) (B)- 1965 Moldenke, Monograph of Lippia 107 LIPPIA CAFFRA Sond., Linnaea 23: 88. 1850. Bibliography: Sond., —— 23: 88. 1850; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 9 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2, 52 : i “@le) and fed. 2], 122 & 189. 19495 Moldenke, Résumé 153 oy arama oon a “inches + tall, with the general st and foliage of pt caag rugosa Thunb. , but differing in pubes- » the hairs subvillous and spreading; ri terete or obso- yrs tetragonal ; mirieaes and branchlets arvenes villous- ary, e very villous; flowering heads subglobose; bractlets herbaceous, ower acuminate at the apex, pubescent, the outer ones involu- rate” and 6 mm. long, the inner ones 2 mm. long and somewhat ee than the corollas; calyx about 1 m. i vey hirsute, much shorter than the corolla-tube; corolla m. long, pubescent on the outer surface, the limb short, somewhat 2-lipped, the up- per lip subemar emarginate » the lower lip 3~fid. The type of this little-known species was collected a, ont Ludwig Philipp Zeyher along the Fo River, Transvaal, Republic of South Africa. It is know to me only from tes otte~ inal description. LIPPIA nye Moldenke, porielowe: a 378, nom. mud. (1950); . Bot. 38: 326-327. “Se eae oldenke, Phytol —_ " 378. 1950; Moldenke, Am. Journ. Bot. 38: oer els G. Taylor, ‘ite Kew. Suppl. 12: 82. 1959; Moldenke, Résumé 12 ae 9 ow, very densely Srelnt theres sets nodes annulate; principal internodes elongate, 6--18 om. long; leaves decussate- e-opposite, e 6——13 om. long, 1——-3 cm. wide, obtuse or subacute at the wets crenate srl the margins from the base to the apex with r » divergent teeth, attenuate-acute at the base, ve a ly pubescent ba puberulent above, more densely ort-pubescent on the venation beneath and pilosulous on lamina, deci flat above, pr eath, not ana osing; veinlet retic- lation abundant, obscure above, prominulous beneath; cence axill, and termi -pedunculate, paniculate-spicate; peduncles stout, w 6-~12 cm. long, sulcate, Pw: densely pubescent with brownish malticellular hairs, t one usually terminated by a pair of foliaceous bracts similar to the leaves in all respects but only 1.3—2.-5 cm. long and 1--1.2 108 PHYTOLOGIA Vol. 12, no. 2 cm, wide; — very densely flowered and white-lanate, subcap- itate, 8-12 mm. long, 8--13 mm, wide, 1--3 at the apex of the peduncle, obnerie or short-stipitate on axillary inflorescences, the stalks 1--2.5 cm. long on the terminal antiopesdence and ens ovate, foliaceous, about 1 cm. long and almost as wide, a by about I = pe of this very distinct species was collected by Father H. carta (no. 2275) -- in whose honor it was naned —— at Panzi, Republic of the Congo, on February 8, 1950, and is de- posited in the Britton Herbarium at the New York Botanical Gar- den. In all, 3 herbarium specimens, including the type, have been examined me. Citations: CONGO LEOPOLDVILLE: Callens 1272 (N), 2275 (N—- type), 3966 (N). LIPPIA CAL 1951. Bibliography: 2 Sudania 1: 2 « 1941 Woldenke, ati Abstr. 25: 3051. 1961; “\oldenke, peytitortd 3: 19 & 56 (19 and h: 189. 1953; Woldenke, Résumé 140, 142, & 61. 1959. its heads smaller and denser, less than 1 cn - wide, and the pubescence - the stems fulvous-villous, very dense and velutin- apex. The type of this variety was aya by W. Mullenders (no. 2093) ina pode roe savanna, at altitude of 870 meters, “at Kassendjim, aut Lomami, Repu abide Br the Congo, on February 27, 198, oa Bepontted in the herbarium of the Jardin Botanique de 1'Etat at Brussels. The plant is described by agree as he We flowers ng gr in the savanna formations, flowering in January and agttag? © Cie *(a91) collected it in December and cites his no. 6550 as Icacina sene- galensis Juss. of the Icacinaceae. In all, 5 herbarium specimens, woypea the type, and 2 mounted photographs have been examined Citations: CENTRAL AFRICAN seg Sep Soe A. Cheval ier 6550 (Br, N). CONGO LEOPOLDVILLE: Mullenders 2093 ) (Br--tyPe, N—photo of type, Z—photo of ae 2351 31 (Br Heng | a ee H.B.Ke, Nov. Gen. & Sp. Pl. 2: 268-269 « Synonymy: + pete callicarpaefolia Humb. & Bonpl. ex Steud., Nom. vehi So » 2, 22 54. LOO. Lippia mexicana Cav. ex Steud., Nom. +, ed. 2, 2: 54, in syn. 18)0 [mot L. mexicana Hocking, 1955, 2 nor wine 1947}. Lippia bicolor Kunth & Soaché, Ind. 1965 Moldenke, Monograph of Lippia 109 Sem. Hort. Berol. 1845: 10. 1845. Lippia calocephala Zucc., Del. Sem. Hort. Monac. . 1846. Lippia callicarpaefolia Humb. & Kunth Lippia iodantha Robinson & Greemm., Garden & Forest 9: 103 & 105, fig. 11. 1896. Lippia bicolor Kunth ex P, C. Standl., Contrib. U. S. Nat. Herb. 23: 1246, in syn. 192). Lippia callicarpiaefolia Kunth ex Hegi, Illustr. Fl. Mittel Eur. 5 (3): 220. 1927. Lipp callicarpifolia H.B.K. ex A. B. Seymour, Host Ind. Fungi N. Am. 588. 1929 [not L. callicarpifolia Schm., 1959]. Lippia arborea Sessé & Moc. ex Moldenke , Prelim. Alph. List Invalid Names s 30, in syn. 1940. Lippia involucrata Sessé & Moc. ex Moldenke, Prelim. 19,0, Lippia arborea Pavon & Moldenke ex Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1] & 95, nom. mud. 19l2. List Invalid Names Suppl. 1: 1), in syn. 1947. Lippia callicor- paefolia Kunth ex Moldenke, nbcuns ea a 5, in syn. 196. Bibliography: H.B.K., Nov. Gen. & Sp. 2: 268—269. 1818; ud - Bot. o°2y 2 oh. 180; sets. Repert. Bot. Syst. - 5-55. 18h5; Kunth & Bouché, Ind. Sem. Hort. Berol. 18h5: 10. As DC > Pro - Nat. Mose, 6 (2): 204. 1863; Bocq., Adansonia 3: 1863; Podiiesctskt, Zeitschr. Allg. Oesterr. Apoth. Ver. 21 (30): 7l. 1883; Jacks. in Hook. f. & Jacks 3., Ind. Kew. 2: 95 139k; Briq. in Engl. & Prantl, Nat. Pflanzenfam. W{ 3a): 152. 1895; Robineos: & Greenm., Garden & Forest 9: 103 & 105, fig. ll. 1896; B Barnhart, Bull. Torrey Bot. Club 29: 597. 1902; Loes. Moldenke, Alph. List Common Names 27. 1939; Moldenke, Prelim. Alph. List Invalid Names 29--31 & 36. 190; Leon Alvarez, Co Contrib. own Geogr ere Phytologia 2: 330 & 3h7. 19175 yaa, aphe: oo - Dn 1, Fagot “ yt se 19h,7; » Alph. ae 352, 357, , 526 , Bios, "573 (918), 2 “685, cask or 7h, 301" 820 oy O78, 919, 926, 960--962, & 2), and i: 999, 107h, 1081, 1235, 1256, & 129k. 19495 a Geogr. Dis ret See » fede 2], 31, 35, 73, 182, & Ie “19; H. N. & A. L. Moldenke, Anal. Inst. Biol. Mex. 20: 9. ach = 110 PHYTOLOGIA Vol. 12, no, 2 1949; Moldenke, Inform. Mold. Set 49 Spec. 2 (195k) and 51 Spec. 3. 19563 Moldenke, Résumé 36, 2, 8h, 220, 310--313, 33h, 360, 460, & 461. 1959; Moldenke, Phytologia 8: 131. 1961; Moldenke, Résumé Suppl. 3: 33 (1962), 8: h (196k), 10: 2, 3, 5, & 6 (196k), and 11: 7. 1964; Langman, Select. Guide Lit. Flow. Pl. Mex. 421, ho, 588, 596, & 1010. 196). Illustrations: Robinson & Greermm., Garden & Forest 9: 105, fig. 11. 1896. Shrub or low bush to tree, 1—.5 m. tall; branches tetragon- al, short-hirsute or hirtous-tomentose, canescent; twigs slender, tetragonal, densely tomentose; leaves decussate-opposite, petio- cm analiculate adi teeth, rather densely hirsute-pubescent or softly pilose with bulbous-based hairs above, becoming rough and reticulate-bullate 7.5 cm. long, the branches subverticillate, tetragonal, hirtous- tomentose and incanous; peduncles very slender, numerous, 5--6 em. long, densely hirsute-tomentose, ascending; heads semi- globose, 1--2 cm. long, 1.5--2.5 cm. wide, about the size of a Prunus spinosa fruit, pedunculate, purple; bracts at first pale- green, then orange, finally rose-purple, ovate or broadly ovate, the outside, twice as long as the flowers, imbricate; flowers sessile, 3 mm. long; c campanulate, compressed, sub-bidentate, very densely white-pilose; corolla infundibular, pale-yellow or yellow to orange, or whitish and turning brown [also described 45 ; vate ou mustard seed (Sinapis arvensis), invested by the fruiting-calyx, 2-locular, smooth, glabrous, the cells l-seeded. The type of this very handsome species was collected by Fried at an altitude of 2336 meters near Mexico City, Fede strict, Mexico, floweri May, and is deposited in the 1dt and Bo herbarium at the Muséum National d'Histoire Naturelle 4t Paris, where it was photographed by Macbride as his type pho 1965 Moldenke, Monograph of Lippia li graph no. 39486. The type of L. arborea was collected by José ree Pavon, probably in Mexico, although the label is in- bed ru", and is deposited in the Deles —— rt Herbarium at the iiiaeova tales et Jardin Botaniques at Gene The name gigantea, seems to be based on Herb. W. H. pt sen. [Hort. udin, France, ae and is deposited in the Dudley Herbarium at Stanfo rd Univer ity. The L. Paes of Hocking, referred to in the synonymy above is a synonym of P vis scaberrima (A. L. Juss.) Moldenke, while that of Moldenke is Li Lippia tepicana Moldenke; L. callicarpifolia Schau. is a synonym of L. pringlei Br Briq. The reference, dieoseraces on & Greem., Garden & Forest 9: 103 & 105, fig. 11", rred to a- bove, is sometimes cited oy "Pringle, Garden & Feat 9: 103". Lantana bracteosa Mart. & Gal. and Lippia bracteosa Mart. & Gal. are sometimes included in the synonymy of L. Cae ~ as, for instance, by Standley (192) —- but belong to Lipp bracteosa (Mart. & mt 4} wets which see. Walper a 3) Te= duces L. callicar © synonymy under L. mberiatd Cav. comes ( (1 ani T} Siac eo newt in his Section Rhodolippia, sectio The settles “has been eit in oak woods and forests, damp pine and oak forests, open or low spiny deciduous woods, damp quebra- das, dry bs ” and rocky rubble at the base of waterfalls in canyons es of oak forests, ar” on bushy or brushy hill- sides, beiater or thickety slopes, mountain or bracken cos anna rocky slopes in oak fore rests, exposed open slopes, hills and hills, barrancas and the slopes of barrancas, at altitudes of 500 to 3000 meters + rine « October to March and in May, fruit- ing in November and oe describes the last as "not very frequent" in Guerrero, reports that the plants are cropped by livestock. Hogi (1927) says that it "wird wie Salvia bemtzt". Seymour (1929) affirms that it is ha by the fungi Prospodium lippiae (Speg.) Arthur, Puccinia lippiae Speg., and Uredo lippiae Speg. The flowers are described as "pale-yellow" on z Ce. ee 58896, "yellow" on Steyermark 31462, "orange" on ae 11156 & 11868, "whitish turning brown" on Steyermark 3391), and "purple" on Hinton 11100, 11633, & 129h5. Herbarium material has been none eee and distributed in weloaae under the names "L. involucratae affinis", L. lupulina +, L. umbellata Cav., and tite =e the other r hand, the Pittier 1,29, eietribeib as L. callicarpaefolia, is actually L. es Moldenke; Purpus 21 is L. hypoleia Briq.; ortts 1640 collection of L. moritzii ’ TUrCcz 5 urcz.3 Edw. Palmer Pringle 1733 & 27h3, Purpus 503 s 508 & 8206, and Seler & . Seler ist are 112 PET TO 1:04:54 Vol. 12, no. 2 L. pringlei Briq.; Linden 1j1 is L. substrigosa Turcz.; and M. E. fones Ee is L. tepicana seni Tn al. iy vor herbarium speci specim d 7 mounted photographs and other A rp ; ge ge “type of any oh £0: material of most of the ape: involved, have ed by m : MEXICO: ’ Sexctal: Districts Hunboldt & Bonpland s.n. tuantetae o pote 394,86] SSL ana of type, N—photo of sa Bs i photo of type). Guerrero: Frye & Frye 3133 (G, Se--6338); Hin- ton 9838 (Au, It, K, Rf, Ur), mace (Fs, La, N), ae cee 11302 (G, Ia, N), 11633. (G, La, N, Rf), 148 68 (G, N, N, Rf); Ly- onnet & Elcoro 1163 (W--17L5973); "Vexia 9103a (G, ate He Be Moore’ 5567 (G, N); Nufiez son. [Huitzuco, 1922] (Me, Me); Rowntree s.n- {near Taxco, Dec. 6, 1938] (Fs); Seler & Seler 252 (G, Gg- 25895); Urbina s.n. [Enero de 1886] (Me). México: Hinton 2620 (A, N, N, N, N), 2989 (A, N, N, N); ao a (cb), 27853 Ties 30020 (se), 32091 (Ss). wtohvebdete Hinton 12383 (G, N, WN, R aie Tt, Me, We, Mi, Min—15):10, S, vt), nek (cm, G, en N); Schiefer 18h (c, vi). Vera Cruz: née 96 (Q y. State undeter- mined: Bartling s.n. (M); Ce Ae Ehrenberg s.n. (Vt); Haenke 1574 (N); Herb. Meisner s.n. (M); Lemmon & Lemmon 20) (Ca—100955, > GS Pavon s.n. ["Peru"] (Cb, N, N—photo, N--photo, 0, Z—-photo); Quar- les ¥ van Ufford 135 (Ut); sessé, Mocifio, Castillo, & Maldonado 2217 (Q), 2220 (Q). GUATEMAIA: Baja V Verapaz: P. C. Standl 69695 (N). Chimaltenango: P. C. Standley 7 798k (N). Chiquimula: Steyerark 30975 (F--10%176), 31462 (F--1047003). Quezaltenang®: Hartweg 560 (N); Steyermark 220th (Benn057261) - Sacatepéquez: P- = C. Standley 58896 (N). ee P. C. Standley 62750 (N). CULTI- VATED: France: H Herb. W. aa 8.0. thot Daudin, 1848] (Du-- 166571). MOUNTED IL TiLSta TONS: Robinson & Greenm., Garden & For- est 9: 105, fig. 11. 1896 tpocauetanye LIPPIA CAMPESTRIS Moldenke, Phytologia 2: kik. re per Moldenke, ’ Phytologia 2: lk. 1548 Moldenke, Contanes 13: « 1948; Moldenke, Alph. List Cit. is A251. 19495 at foe - Distrib, Verbenac., [ed. 2], 80 & 189. 29195, seaalteag, rs Farmac. 19 (10): 169. 19515 A Je oes ts 19 Fl. : 6, (ssn) pe 12: 17. iss Votan Résing 5 93& “él. 19595 Angely, Fl. Paran. 16: 60 (1960$ and 17: Dwarf perennial, woody at the base; ais numerous, simple, 15-~17 em, tall, rather densely glandular—pubescent and also hir- sutulous with much longer, divaricate, white, glandless hairs; nodes not noticeably anm a ygemae leaves decussate-opposite, sessile) leaf-blades ovate, 7--16 mm. long, 10 mm. wide, the lower pairs increasingly smaller, “subacute or obtuse at the apex, rounr 1965 Moldenke, Monograph of Lippia 13 ded or cordate at the base, entire, ciliate, hirsutulous Sciescect on both surfaces and some ewhat glandulose tenet lar- ger bhai he! rather obscure above, very faintly subprominulous beneat yay oe axillary, 2 per “a en the median or upper axils; peduncles very slender, 1.5--2. - long, rather densely Bath lnaoccabesoert and nfSiateibas! heads “banepherie, less than 1 cm. long; bractlets ovate, about 5 mm. long, 2 mm. hi 5 long, the tube about equaling the bractlets, unifomly puberulent outside, the limb puberulent on the back, whist gab within. The be of A pee species wan collected by Per Ki almar Dusén on a campo a t~ Jaguari , Parand, Brazil, oe October 9, 1911, ar" is deposited in the herbarium of the Naturhistoriska Riksmuseun at Stockholm. Th nis thus far only from the type collection, In all, 2 asta specimens, including the type, and mounted photographs have been examined by me. 1911] (F—photo of type, N—isotype, N--photo of type, S—type, Si—photo of type, Z=—photo o of type). PPIA CANDICANS Hayek in Fedde, Repert. Nov. Sp. 2: 86. 1906. Bibliography: Hayek in Fedde, Repert. Nov. Sp. 2: 86. 1906; ? - Kew. si me kk: 142. 1913; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 37 & 95. 192; Moldenke, Alph. List Cit. 1: 233 (19k6)° and 3: 695. 1949; Moldenke, Known Geo wi ede Dis- trib. Verbenac., [ed. 2], 80 & 189. 1919; Moldenke in ee Los Angel. Co. Mus. Contrib. Sci. 7:10 & 11, fig. 2. 1957; uel- denke, Résumé 93 & 161. 1959; Moldenke, Tales © Suppl . 8: 2. 1964. a in ce Moldenke in Dawson, Los Angel. Co. Mus. Contrib. fig. 2 Shrub : ; branches tetragonal, appressed puget Signa pened leaves decussate-opposite, petiolate, ovate-lanceo anceo- late, acute at the apex, attenuate into the petiole white tued corolla hypocrateriform » surpassing the subtending bractlet, red- ets thant pilose on the itches e. The this handsome species was collected by G Gustine (no. 3942) in Gofas, Brazil, and is deposited res the her- barium of the Nat’ Naturhi storisches Masse in Vienna, where it was Photographed by Macbride as his type photograph no. 34329. Hayek 3) Says of it "Affinis L. sericeae Scha., a qua habitu grac- iliore, indumento breviore adpresso et eset mate minoribus valde 2 t ering in April. In all, 3 herbarium sp etnies, including the bh Collection, and 3 mounted chbtnerethe have been examined by 114 PHYTOLOGIA Vol. 12, no. 2 Citations: BRAZIL: Gofas: E. Y. Dawson 14695 (Z); G. Gardner 3942 (Macbride photos 3329] (It--photo of type » Kr--photo of aps. M--isotype, N--photo of type). Minas Gerais: Heringer 3789 (B). ate LIPPIA CARDIOSTEGIA Benth., Bot. Voy. Sulphur 153--15). 186. Synonymy: Lippia brenesii Standl., Field Mus. Publ. Bot. 18: 1009. Bibliography: Benth., Bot. Voy. Sulphur a 18,6; Jacks. in Hook. f. & Jacks. so inde Kew, 2: 95. 189); P. C. Standl., Field Mus. Publ. Bot. 18: “Loo9. 1938; Moldenke, Alph. List Common Names res 1939; B. H. Davis, Mycologia 32: 170. 1940; Moldenke, Phyto- logia 2: 21. 1941; Moldenke, Known Geogr. Di istrib. Verbenac., fons soekag 17, 20--23, & 95. 1942; Anon., Homenaje al Prof. A. M. . 19hh Moldenke Phytologia 2: 70 & 107. 195; Moldenke, Cit. 1: 23, 2h, 192, & Pte nats as J. Salisb., Ind. Kew. Suppl. 10: 13). 197; Moldenk Invalid Names 13. 1947; Moldenke, Phytologia 2: an rs 3 Ble “uty rs Bol. Soc. Venez « pee tye li: hh. 7; Ho N. & A. L. Mo jenke, Pl. Life 2: 51. 1948; nen, ele Cit. 2: 351, 390, 408, 69, “ae oh 2}, et 35, 37—39, & 189. 199; H. Ne & A. L. Moldenke, . Inst. Biol. Mex. 20: 9. 1949; Matuda, Am. Midl. Nat. hh: 576. 1950; Moldenke, Phytologia 3: be 1951 arty ce ire Bot. 8: 228. 195); Molderke, Résum tik 461. 1959; Moldenke, Résumé Suppl. 7H % t3360,° - an & 1 1 (2962), and 11: b. 196. frutescent bush, shrub, or tree, 1--5 m. tall, densely and strictly branched , strong-sce ented with a pungent marigold (Tage~ tes) odor; stems slender, arching; bark pale-gray or gray, smooth labrous; eranshleve slender, rather densely strigose with short whitish hairs; principal internodes short or elongate; brown, or glabrate, to ie mn. long in the fruiting stage, Scere a ged; perianth not surpassing the restate 3 calyx compressed, ra 1965 Moldenke, Monograph of Lippia 5 dentate, scarcely more than 1 mm. long, densely white-pilosulous; coro rolla’ hypocrateriform, white or greenish-white to pale greenish-yellow, Pt ob 3 mm, long, the tube slender, glabrous on the outer surfac e limb 1.5 mm. wide, the lobes broadly rounded, Potoplas baxauaxaet on the outer and sparsely puberulent on the inner surface. Bann type of this species was collected at the "Gulf of Fonse- » between Honduras and El Salvador [not "Mexico" as stated in the’ Index penny td and is deposited in the Bentham Herbarium at the Royal Botanic Gardens, Kew. The type of L. brenesii was col- lected by Alberto Mora Brenan (no. 6477) along the Camino de San Gerardo, Santiago de San Ramon, Costa Rica, in December, 1928, and is deposited in the herbarium of the Chicago Natural History Museun Lippia cardiostegia has been tee growing on hills, dry rocky and brushy rocky Slopes, brushy and dry or damp brushy aareee een grassy open slopes, in hedges and thickets, dry o grassy thickets, damp or wet thickets, rocky fields, wy Rebeati- s around base or hills or peaks" in Nicaragua. The specific name is sometimes uppercased for no valid reason. Common names repor- ted for it are "chiligua", "coronchoch", "oreganillo", "oregano montes", and "orégano montés". The name "Yoreganillo" is also ap- plied to L. graveolens H.B.K., L. myriocephala Schlecht. & Cham., and L. umbellata Cav. ee The Kellerman s.n. [Laguna, January 20, 1906] is the type host collection for the fungus Cercospora cardi ceteriae B. H. Davis. G. F. Baker 206 in the Britton Herbarium has the spikes abnormally elongated. Matuda 743 gives no indication on its label as to the exact locality of its collection, but it probably was gathered in Chiapas. Some collectors refer to the leaves of this species as "membranous", but this is a a palpable error, since they are con- Herbarium material has been misidentified and distributed in herbaria under the names L. asperifolia Rich., L. ocephala Cham, & Schlecht., L. umbellata Cav., and Lantana involucrata L. On the other hand, the M. C. Carlson 168, distributed as Lippia Cartlostegia, is actually L. bracteosa (Nart. & Gal.) Moldenke. Standley (1938) says material of this plant has been refer- red previously to L. myriocephala Schlecht. & Cham., from which is amply distinct » and to . Bs oe Rich." He cites the ollowing collections from Costa Rica, not as yet seen by me: Brenes 5855, Pittier 7519, P. “Ce Standley 1133, Tonduz 13631, 116 PAT .T 04:06. 2 Vol. 12, no. 2 and M. Valerio 1521, presumably in the herbarium of the Chicago Natural Histor History Mus Museum. In all, 103 herbarium specimens, including the type collection of L. brenesii, have been examined by me. Citations: MEXICO: = aes Matuda 743 (Mh, Mi, N, N), 938 (Mi, N), 1670 (A, Mh, Mi, N, N, N). GUATEMALA: Chiquimula: P. Co Standley 74739 (N); Stayeiiark 30305 (F—1056265), 30569 (F— 105201). Escuintla: P. Cc. Standley 60217 (A, N). Guatemala: P. C. Standley 61418 (N). Huehuetenango: Steyermark 50770 (N). Ju- tiapa: P. C P. C. Standley 7500) (N), 75922 (N). Santa Rosa: Heyde & Lux 4385 (G); P. C. Standley 60713 (N), Troe Sai 78412 (N), 78809 (N). Suchitepéquez: P. G. Standley 62061 (N). Department undetermined: Friedrichsthal 15 (V), 878 Tier tpelne) (v), 1118 {Lapunte] (V, v), a a (V, V, V); Kellerman s.n. {La- guna, January 20, 1906] (N, N). HONDURAS: Gracias: acias: Hjalmarson son. [1852] (S). a SALVADOR: Ahwachap4n: P. C. Standley 19860 40, 8), 20222 (G, N). La Libertad: Stork & Horton n B6L6 ( (Ca-- 643478). La Unién: P. Cc. Standley 20803 (G, N). San Salvador: Herb. Lab. Quimico 1h (N); P. C. Standley 22677 (G, S), 23262 (G, S). Sonsonate: ay, ve Hartman 13h (S). Department un ndeter~ mined: Kovar 1090 (N, Rf). NICARAGUA: Carazo: N. L. H. Krauss 381 (Z). Chinandega: C. F. Baker 13 [766] (Ca-—199193, Du-- 76202, Gg--31150, Mi, N, Po--64698, W--862825). Grenada: C. Fe Baker 206 [660] (a, i, lie "Lavy 241 (Cb, N). Managua: Garnier 264 (Mi), 641 (I, N, N), 100) (Mi); C. L. Smith 108 (Ca- 975391, G, N, N, N, Tl, Vt). Ometepe Island [Rivas]: Shimek & Smith 8 (Ca--975390). COSTA RICA: Alajuela: Brenes 1) (N), 4756 (N), 6477 [332] (N), 13238 (255; 13358] (N, N), 14339 [20] (N, N), 17509 (N)s A. Smith P. P.2346 (A, Ca—-12709). Cartago: Brenes sen. [San Mateo, 1903) (N)} H. Pittder 2519 [Herb. Instit. . Phys- ico-geogr. Nat. Costaric. 13215] (Cc, G). P G). Puntarenas: Holm & Il- tis 313 (N). San José: Biolley & Tonduz 7218 (Br); Brade 2135 (Br) H. Pittier 602 (Br), 7218 (Br), 6452 (Br); Tonduz 1160 (Br, Br), 3159 (Br); Tonduz & Pittier 8452 (Br). Department undeter- mined: H. H. Pittier s.n. (ra =; LIPPIA , CARTE UDORA Meikle, Kew Bull. 1948: ,67-~1,68. 19L9. Bibliography: Meikle, Kew Bull. 19))8: 167 See vies ‘Molaeckt, ll: = Spun smelling subshrub, to 1.5 m, tall, many-branched; bark prea rsa at first pale, later gray or dark-brown; branches spr eading-erect, very slender, purplish when young, subterete °F 1965 Moldenke, Monograph of Lippia 117 obtusely Marieke, bape sparsely eo avepbenie] Jae glabrescent, densely covered with shiny glands; lea s decussate-opposite, ess ot fs Taher petiol ate petioles alacisar, about 1 cm. long, pubescent, canaliculate above; leaf-blades ovate, to 2.5 cn. long and i -7 cm. wide, gray green, a tira at the apex, subtrun- cate at the base and er radually narrowed into the petiole, rarely shortly decurrent, crenate or amie serrate along the margins, rugose and venose ’ above ve, appressed-pubescent and sparsely glandu- lose above, densely canescent-puberulent beneath and covered with shiny glands; inflorescence axillary, solitary, short—pedunculate, ed; peduncles slender, to 7 mm. long densely white-tonentose; spikes to 5 mm. long and 7 mm, wide; astryee twice anthesis and 2 mm nate hi pex, densely glandulo ose, sparsely feral te sigt after anthesis greatly enlarged, finally membranous, venose, hirtellous, erose margins, acute at the apex, cordate at the base, to 8 mi. long and 15 m. wide, resembling the strobilus o iupulus; calyx minute, to lm. long and wide, membranous, dense- white-hirsute, divided into 2 obtuse or mnarginate lobes, en- larged after anthesis and entirely covering the fruit; corolla e long, rather dark, striate, subglabrous at the base outside, Sparsely pilose Bok " the lobes l, rounded, whitish, incurved, undulate and cremate along the margins, livid-anmiate in the bs within; stamens , didynamous, two inserted in the upper Serted at the middle of the tube; filam ve anthers ovate, 2-celled; style to 2.3 mm, long; stigma obliquely declined, rather ate or subemarginate . the apex; Blobose, 2-celled, about 1 nok wi ae mn. wide, ine: dry, uded s taanaiton fru ~calyx at d by the stoiveted arate splitting into 2 mtlets; pericarp da ~ type of this npoigroe species was collected sg Hs aogier Talker at Wamba, in the Northern Frontier District, anuary |, 1918; and a deposited (dried and alcoholic e material) Speciebus africanis valde dissimilis. This is a very distinct Species, quite unlike any other African eather but bearing a of been d ce tations: KENYA: Jeffrey 5.42 (S); Ge “Walker s.n. [Wamba, l. 1948] (N—isotype, N—-photo of type) « 118 Pansat Os 0-G 1 A Vol. 12, no. 2 LIPPIA CARVIODORA var. MINOR Meikle, Kew Bull. 19,8: 68-169. ynonymy : Lantana microphylla Hutch. & Bruce ex Meikle, Kew ait 1948: WB i) in syn. 1949 [not L. microphylla Cham., 19h7, nor Franche | 1882, nor Mart., 1828, nor Peter, 1959]. Lantana petitiana soi ex Meikle, Kew Bull. 1 948: 468, in syn. 1949 ew Bull Phytologia 3: 136 & 137. 199; Woldenke, Know Geogr. Distrib. Verbenac., [ed. 2], 110, 118, & 189. 1919; Moldenke, — 135, 959; Cuf., Bull. Jard. Bot. Brux. 32: Suppl. 791. and is deposited in the herbarium of the Ro oyal Botanic Gardens "at Kew, where it was received on January 26, 190). “Meike 92) notes that he var. minor, with pale whitish bark, small leaves and fruiting bracts, looks distinct, but some specimens (e.g-, Bally 3029 and Edwards 298h) approach typical L. carviodora, and it ma: may be merely a starved or a ab state of this species’ The whole plant emits a powerful odour reminiscent of caraway ~seed (especially noticeable in the var. = iiy minor), an and is said to be used by the Somalis as a seecdaphishin for ne Vernacular names redial ded are "ged hamar", ed-hamar", "ged hamer", and "ged—hamer." T 3 he Lantana mi matpyod of Ghaniesc, referred to in the syno- nymy above, is a synonym of Lippia peels Cham., that of — is Lantana petitiana A. Rich., that of Martius is Verbe- microphylla H.B.K., and that of Peter is Lantana rugosa var Seaieenee Moldenke, Meikle cites also Gillet 4373, Bally 3029 3029, and Glover & Gilli- land 48 from British Somaliland, and D. De ds Es Edwards 298) from Keny4: Bally describes the plant as "very c -ee+ein seni—desert scrub" at 00 meters altitude and describes the plant as a s ut 12 dm, tall. It has been collected in flower in September, October, January, and March. Thus far I ae seen only a single phototype. Citations: BRITISH SOMALILAND: D. Thomson 72 (N—photo of type): LIPPIA ae Chev., Etud. Fl. Afr. Cent. Frang. 1: ale, onym rig 6 ely A. Chev., Etud. Fl. Afr. Cent. Frans. 1: 242. 193i, Prain, Ind. Kew. Suppl. 5: 153, 1921; Moldenke, Known Geogr. Dis- trib. Verbenac., [ed. 1], 8 & & 95 (2942) and [ed. 2], 11h & 189. 1949; Moldenke, Résum$ 1h0 & 1965 Moldenke, Monograph of Lippia lg All that is known about this ght plant is what is stated by Chevalier, who bases the name on his no. ay which was collected in clearings at Bangui, ia dle Ubangi, Uba: Ubangi- chari, Central African Republic, on December 18, 1903. He de- scribes its inflorescences as white, but gives no further de- scription, LIPPIA CHACENSIS Moldenke, Phytologia 2: ik—}15. ae: Moldenke Looe ia 2: path pe io 5 Moldenke, Lillo ldenke, densely appressed-strigillose above, very densely velutinous with very short flavescent hairs beneath; the slender midrib and or 6 pairs of secon daries, and often the larger veinlets, slightly subimpressed above, prominulous beneath; inflorescence axillary, 1 or 2 per node, shorter than the subtending leaves; peduncles very slender, 1--1.5 om. long, heads pereheric or _cblong, to man 1 cm broadly ovate, lh. - long, about 2 mm, wide at the base, a- cuminate - the fans pte, strigose or strigillose, canescent, glandulose; corolla-tube about 6 m long, very densely strigose’ on the outside, the limb about | mm. wide, glabrous or slightly pubescent at the base. type of this species was collected by Klas Robert Elias Fries (no. 1445) in an open grassy campo at Tatarenda, Gran Chaco, Santa crus, Bolivia, on March 22, 1902, and is deposited in the herbarium of the Naturhistoriska Riksmuseum at Stockho collector describes — ein tee as “rare” and the flowers as "liv- It r only from the type collection and we iain ities a as and distributed under the name B.K canescent-strigillose; - long; tlets L. geminata In all, Xp herbarium specimens, including the type, and ) moun- ted photographs Imve been examined by me Sb cimcraee BOLIVIA: Santa Cruz: R. E. Fries prnny ae. of » N--isotype, N—photo of type, S--type, Si——photo of t ~phots af types. hee 3 ys CHEVALIERII Moldenke, Phytologia 2: 313--21h. (1 blio gna ¢ Moldenke, Phytologia 2: *13--31h, 39, “340 fea hr) od : 136. 1949; Moldenke, Known Geogr. Distrib. Verbenac ., 103, 111, & 189. 199; Moldenke, Alph. List Cit. h: 1197 120 Paws ULUGISA Vol. 12, no. 2 & 1201, Bj Moldenke, Phytologia 3: 291. 1950; E. J. Salis es Be 138. 1953; Moldenke, Heal aK pe OP a & él. "1559. Herbaceous (?); branches very slender, stramineous, subterete, striate, rather obscurely strigillose; nodes ann nulate; principal internodes elongate, 5--13 cm. long; leaves ternate or in 's. BOOK REVIEW Alma L. Moldenke "A Selected Guide to the Literature ey the Flowering Plants of Mexico", by Ida Kaplan Langman, 1015 pp. University of Pennsylvania Press, Philadelphia, Pennsylvania. 196. $25.00 in 1948, when my husband and I were visiting Mexico's ier de Biologia, we first met Mrs. Langman as she was in- triously starting this useful bibliographic contribution to botanical 1i bated The idea for this work started its gesta- gman i to produce this magnificent compilation of the literature on Mex- ican flowering plants, their description, their taxonomy, their phytogeo, ee their economic importance, their mention in books of travel, their indigenous names in various local languages, and | their role in the nistory of botany -- surveying a tremendous tines of printed and manuscript information over a long period of sible to avoid in a work encom y de from so many di + sources. ce the ub r has offered to print 4 list of errata soon for distribution wit! book there is no Both reading and print is clear and of a suitable size and format. The index 4 lone covers 157 pages of four columns each. o many present and future botanists and other researchers Te ferring to this book will be ever so much in the author's debt because of all the sources made so readily available here, 4 all th? q ch author for her persevering dedication, PHY TOLOGIA Designed to expedite botanical publication Vol. 12 May, 1965 Fae as CONTENTS REED, C. F., Distribution of Salvinia and Azolla in South America and Africa, in connection with studies for control by insects... 121 MOLDENKE, H. N., Materials toward a monograph of the genus APU Ts oe eke be OAs eee = wigs ese ee ne 130 | MOLDENKE, A. L., Book review...... a Oe ee Ee 182 elt _ Published by Harold N. Moldenke and Alma L. Moldenke ‘ 2s = 15 Glenbrook Avenue Pee ; Yonkers 5, New York, U.S.A. Price of this number, $1; per volume, $5. 75, in » advance, tee or $6 2 at close of volume URE BoTanicaL DISTRIBUTION OF SALVINIA AND AZOLIA IN SOUTH AMERICA AND AFRICA, CONNECTION WITH STUDIES FOR CONTROL BY INSECTS Clyde F. reead/ In 1961, Dr. Fred D. Bennett, Commonwealth Institute of Bio- logical Control, Trinidad, W.I., made observations on the insects attacking Salvinia spp. in British Guiana and North Brazil. In ? 8S) an de San Migual near Chuy). From October to December, 1964, Dr. Bennett conducted feeding tests with insects attacking Salvinia suriculata at Belem, Brazil. The ultimate aim of these investigations by Dr. Bennett was to obtain some organism At ap of effecting control of Salvinia Suriculata in Kari Rhodesia, Africa. An edi- torial, entitled "Weed rg Bes River Fight Zambezi Power Dam”, in the Baltimore Ey Evening Sun (p. 3, ov. 18, 1961), released by ~ 1d Pellers, kills fish and could eventually turn the lake into a Swamp, since it is so thick in places iy Sine Ghat Se crs St and trees and shrubs can grow on it. Salvinia was first dis- covered in this area of Africa near Victoria Falls in 1949 and was rie Foe at that time as the harmless species, Salvinia hastate. varieties In South America, Dr. Bennett found that huge areas of lakes and reservoirs and other sources of drinking water are being covered by Salvinia auriculata. Also water-ways are being Clogged by Pistia stratiotes be Eichhornia crassipes Solms and Alternanthera ra philoxeroides Gri There, these species, al- though not en entirely held in tmeag by insect enemies, do have a ee 2/ Botanist, crops Research Division, Agriculturel Research Ser- Vice, U. S. Department of Agriculture, Beltsville, Maryland, and Reed Herbarium, rag » Maryland. 21 122 PATTOULOUGIA Vol. 12, no. 3 complex of insects which attack them and which in turn are attacked by parasites and predators. In the various reports from 1961 to 1965, Dr. Dennett found that there are four species of in- sects that feed on Salvinia and ornate: by their feeding habits or by their pela rsae ol = be restricted to an aquatic environment. These seumingte es isi et Pauliniidae by some enthors), © » ar rdbeais ¢ Curculionidae), Samea mulisElicalis ection). sf Lipostemmate purpurata meets Esuris is a synonym of Lipostem- me Paulinia acuminata grist’ attacked Salvinia auriculata and Pistia stratiotes around Belem. In ee lad adults were found on Temma and Azolla, as well as Salvinia an d Pistia. repo: Carbonell (1964) reported this species on Azolla filiculoides Iem., as well as Hydromysteria stolonifera F.G.W.Mey. (Hydro- lata. In the laboratory this insect also attacked Eichhornia qrasetpes s and Nymphaea sp., and to a limited extent a species of Samea multiplicalis was found to attack Pistia stratiotes in the field in Trinidad, British Guiana and Brazil. In the labora- tory it also laid eggs on Salvinia and Pistia, and a larvae fed pean Lyon oe Eichhornia crassipes. 1 . Those feeding on Lemna died in 8 Cyrtobagous s 8 is known from Corumba, gees —— (S.W. Brazil, orig. loc.), “pritieh Guiana and mente (found Salvinia). This species seems to prefer Salvinia; it does not attack Pistia or other aquatic plants; and the vas did not fete oe laced among ic and Eichhornia. Iipostemmata purpura used the tests at Belem. ia cet tests conducted at Belem in 1964 indicate that Paulinia mL crop plants tested at the same time (rice, lucerne, watercress, cotton and sugarcane). Hence, introduction of these species of insects into the Kariba Lake would not endanger any agricul crops either near the lake or farther down the watercourse. Singularis appears to be very restricted on its aicions F habits and is recorded as iter destroyed stands of winia in small ditches near Georgetown, British Guiana. My interest in this project has been the identification of cimens of Salvinia and Azolla found by Dr. Bennett new in his investigations. Meany localities for the in South America now may be recorded, based upon his collections vet » Uruguay and British Guiana. sil studies were deing made in conjunction with control of Salvinis 1965 Reed, Salvinia and Azolla 123 Suriculata in Africa, specimens were also sent to me by the Kariba lake authorities for identification. Additional records for some of the species concerned, represented in the Reed Herbarium, are also included. In South America three species of Salvinia are frequently confused, namely, Salvinia auriculata Aubl., Salvinia late and S. radula d re . and S. rotundifolia rarely form extensive mats, as s. asuriculata usually 4 Also leaves Ss. s. rotundifolia are s . Weatherby (1937) pointed out the differences among these three species, indicating that the taxonomy had become quite entangled and confusing. He discussed the problems and indicated the following characteristics as clarifying the species. Salvinia rotumdifolia Willd. -- Floating leaves comparative- ly thin, average 1 cm. in diameter and drying dull green; some on regular hairs, which are wholly free and somewhat spreading; the tips aapetans and colorless; sporocarps containing the macrospores sessile. Salvinia auriculata Aubl. -- Floating leaves thick, boat- or prevailingly widest at the broadly cor- 3 papillee we center of the leaf; leaf-tissue usually glabrous Pepillae; sporocarps containing the macrospores stalked. radula Baker -- Floating leaves flat, averaging mich r in S. auricula broadly elliptic or oblong- elliptic to obovate, widest at o the 3 pep Or sometimes nearly obso te; leaf-tissue often with single tri be row: pa: ° these charac- teristics, none of on st of caries bodies, S to be a shade-form of S. ta, as suggested by Weatherby. imens appear intermediate be it s. of the collected in shaded areas, or completely intermingled with 12h PHYTOLOGIA Vol. 12, no. 3 S. auriculata, which could provide a shade situation by itself. The geographic ranges for S. auriculata and S. radula — and ° lata. When the complete life cycle of S. auriculata is thoroughly studied, S. radula may prove to be nothing more than a phase in its life life cycle. Salvinia auriculata Aubl. TRINIDAD: #3. Collected in the sea, Balandra Bay, Trinidad, W.1 a. 8, haton reer quantities of Salvinia have been noted on o sea from in Trinidad, or conceivably from the Orinoco River in Venezuela. F.D.Bennett. (Reed Herb. 39379). #+. Collected from the "Old Pond", St. Mary's Estate, Bonase in Southern Trinidad. Ang. 27, 1961. cae “pond of considerable size many years ago, is now overgrown with grass; small quanti- ties of Salvinia being present. F.D.Bennett. Herb. Trin. No. 16405. (Reed Herb. 39380). ee DLS ak #2. Botany Greenhouse tank. July 15, 1961. W.D.Richardson Herb. Trin. No. 16271. (Reed Herb. 38782). BRITISH GUIANA: #10. Botanic Gardens, Georgetown. Oct. ll, 161. Drainage trench. Sporocarps peeeent F.D.Bennett. (Reed Herb. 39931). #14. Canals and ditches, same loc. Oct. 13, tr F.D.Bennett. (Reed Herb. sne8 fis. Same loc. Oct. 15, 1961. F.D.Bennett. (Reed Herb. 39926; #l2 and #15. Spee Ogle Sugar Estates, about 5 from Conga e: Oct. 12, 1961. F.D.Bennett. (Reed sea 3993! and BRAZIL: #19. Belem, Inst. Agron. Small stream. Oct. 20, 1961. along edge. +t 40349). #37. Belem, Inst. Agron. do Norte. May 1963. Spore- carps abundant. F.D.Bennett. (Reed Herd.). ya al, Museu Para. #21. ificial pond. 196s Pee B.D. Bennett oe Herb. 40343); 18. Artificial fi : - 18, 1961. F.D.Bennett. (Reed Herd. 40344); # 23. 1965 Reed, Salvinia and Azolla 125 Artificial pond. Oct. 22, 1961. ocesegRennett (Reed Herb. 40351); #2h. Artificial pond. Oct. 23, 1961. Aa (Reed Herb. 40348); #43. In tanks, sporocarps present. Feb. 1963. F.D.Bennett. fegcd Herd. ); "438. In ae g April 1963. F.D.Bennett. (Reed Herb e Janeiro, Botanic Gardens. #29. Ponds. Nov. 4, 19613 P.D. Bennett. (Reed Herb. 40345) June 12, 19h0. J.G. G. - (Reed Herb. 39062); #85. In swampy area in pas- ture field near Universidad Rural "Km 47", Rio de Janeiro. Mar. 1963. F.D.Bennett. (Reed Herb.). Curitiba, Parana. #33. In small lake. Mar. 26, 1963. F.D. PpAiemate. ” (Reed Herd. i291). #28. Museu Nacional, Quinta Boa Vista, small concrete ponds, Rio de Janeiro. Nov. 3, 1961. F.D.Bennett. (Reed Herb. 40347). Sao Paulo. #34. In small lake about 1 mi. from Institute Butantan. Mar. 20, 1963. F.D.Bennett. (Reed Herb.); in garden pool in garden of director, Instituto de Botanico, Parque Estado. Nov. 1961. F.D.Bennett. (Reed Herb. 35321). Recife. #36. In pond in Recife, Pernambuco. Sporocarps present. fructifications on the leaves. Feb. 1963. F.D.Bennett. (Reed Herb. 45292). © Alegre to Pelotes. #32a. In rice growing in ditches. Apr. 4 "1983. F.D.Bennett. (Reed Herb. 54725). Obidos. #45 and In lake Maraurti. Sporocarps present. a gr Azolla vet pay te also present. F.D.Bennett (Reed Herd. 45289). pea rrerme Manaus, Amazonas. #50 and 52. In small cross channel be- tween Rio Negro and Rio Solimoes. May 1963. F.D.Bennett. (Reed Herb. 45200), ee bg (Northern Rhodesia): Floating, very abundant in Zambezi t Katambora, Dist. Katambora. Nov. 5- ul, "1989. 0.West 3050. (Feed Herb. 42764; GHSR-34026); upstream end of Kandakan Island, Estuary, Kariba » 1960. J.B.Phipps 2810. (GSR). 126 PHYTOLOGIA Vol. 12, no. 3 RHODESIA (Southern Rhodesia): Kariba Lake, Lat. 17°S - Long. Bumi River, which enters Lake Kariba, Dist. Kariba, elev. 1500 ft. July 1960. B. Goldsmith 92/60. (Reed Herd. 42763; a Victoria Falls, rapidly colonizing areas r permanent spray-action water. Dist. Wankie. Oct. 22, 1959. H.Wild 4851. (GHSR); floating water plant, Zambezi near Boat Landing Victoria Falls. Orta Flying Stage, 1954. 0.West 3234. (GSR); aquatic floating weed in masse | River, Dist. Wankie, Victoria Falls. Mar. 3, 1955. N.C.Chase 5773. (GHSR): Park River, Umtali, Di st. Umtali, alt. float Dist. Ut . Umtali, alt. ft. Aug. 1952. N.C.Chase "4615. (Gask-note we Alston, "native of fmerica" ’ abundant on drying on at Schanarea fi Nov. 6, 1953); re dam near Odzi and small dam eee near Odzi, also about f Odzi, Di alt. 3300 ft. Aug. 20, 1959. J.B.Phipps 3171. (GHSR); on bank at mouth of Br ba Dem, lant forming dense mats in protected area , alt. 1500 ft. Sept. 1, 1960. D.S.Mitchell 580. (HER); floating down the Zambezi, at Chirundu since Kariba flow ancreased about 10 days ago, Dist. . July 16, 1959. G.R.Bates 97192. a Peg se (E.C.Harrington), Silverbow Road, Rusape, don dam, Dist. Makoni. March 1957. H.Booy 7h3Te. (GHSR). CAPE PROVINCE: Kuysua, Gouna Forest Preserve, introduced, rae y. pool near the preserve. Apr. 1957. D.S.Mitchell 150. Salvinia radula Baker TRINIDAD: Floating in Botany Pond, Trinidad. July 15, 1961. W.D.Richardson. Herb. Trin. No. 16272. Alt. 100 ft. (Reed Herb. 30701). BRITISH GUIANA: #79. Botanic Gardens, Georgetown. Drainage trench, juvenile plants. Oct. 11, 1961. F.D.Benmett. (Reed River, floating on on swampy lake, Karenambo. Oct. 9-13, 1937. A.C.Smith 2229. (Reed Herb- 1965 Reed, Salvinia and Azolla 127 BRAZIL: #2. Belem, in pond at Museu Goeldi. Apr. 1963. F.D. Bennett. (Reed Herb. 45293). #hh. Ma, » about 1 —. from Obidos. Apr. 1963. F.D.Bennett. (Reed Herb.). #47 Mixtures of S. radula and S. auriculata. (Reet Herb. >. Salvinia rotundifolia Willd. BERMUDA: #5. Collected from an unspecified location in Bermuda re I.W. 8, Dept. Agr., Hamilton. Aug. 1961. (Reed Herb. 361; U3). Rio de Janeiro, Catumby, in stagnant water. Oct. 15; Br "Z. G.Mosén 2738. (Reed Herb. 39060). Pelotas, R.G.S. #31. Collected in small pond near Inst. Agron. do Sul. Apr. 8, 1963. F.D.Bennett. (Reed Herb.); #32. Collected in rice growing area between Pelotas and Porto Alegre, R.G.S. Apr. 9, 1963. In ditches. F.D.Bennett. (Reed Herb.). Belem. Museu Goeldi. #39 and 40. In small tanks. Feb. 1963. Spo: 6 present. F.D.Bennett. (Reed Herbd.); #1. Som loc., collected by Dr. Walcante. Mar. 23 an... ro- t. (Re Mn ’ ed Herb.); #22. Museu rete tank. Oct. 21, 1961. F.D.Bennett. (Reed ieiak. ko3ia); #e0. Same loc. In artificial pond. Oct. 21, 1961. F.D.Benn (Reed Herb. 40346). Sree = Obidos. #49. In Lake Maraurt. \ aa among Salvinia culata. - 1963. ° - 24, 1963. F.D.Bennett. tere ery 963. Dupl. Aug. 24, 1963 i #51. In small cross Manaus, Amazonas. 1 between Rio Negro and Rio Solimoes. Apr. TE Dupl. Ang. oh 1963. F.D. Bennett. (Reed Herb. ). In South America the most frequent species of Azolla S8sociated with Salvinia is Azolla filiculoides Fe In Africa either Azolla nilotica Decaisne or Azolla pinnata var. africana (Desv.) Baker has been found associated with Salvinia auriculate and Pistia stratiotes. Azolla filiculoides Lan. ae Watercress beds, 100 ft. elev. Watercress has been Srown re since 25 yi te probably the the Azolla was brought into the area with it, po Valley Road. May 8, 1961. F.D. Bennett. 128 PHYTOLOGIA Vol. 12, no. 3 Herb. Trin. No. 16380. (Reed Herb. 39549; US); collected at West Indian Station, Imperial College of Tropical Agriculture. Sept. 21, 1961. (Reed Herb. 39550). BRITISH : In canals, Ogle Sugar Estate near George Oct. ie — F.D.Bennett #lla. (Reed Herb. 39933); aor Water pond, s Estate, Dept. of Agriculture. Oct. 16, 1961. F.D. Boact hg (Reed Herb. 39936). BRAZIL: In small channel between Rio Negro and Rio Solimoes Manaus, Amazonas. May 1963. F.D.Bennett #52a. (Reed Herb. ); in Iake Mareurfi near Obidos, Amazonas. Apr. 1963. D. Bennett - (Reed sate): in Leke Maraurt, near Obidos, we Apr. 1963. F.D.Bennett #419a. ome Herb.); pond near Inst. Agr. do a ecw R.G.S. Apr. 8, 1963. F.D.Bennett #3la. (Reed Herd. ). eer ee URUGUAY: stagnant water, on granite, Pocitos, Dept. video. Apr. (San 1924. Guil. Herter 70518. Seve Sea eal stagnant water, Layago, pire Montevideo. Oct. 1924. Guil Herter aA oand * (Read Herb. ). CHILE: Valparaiso, Laguna bei Vina del Mor. Jul. 1900. Dr. Otto Buchtien. (Reed Herb. 1082 - as A. magellanica). ARGENTINA: Cordoba, prope urbem. pe ate 19, 1877. G. Hieronymis (Reed Herb. 33909 - as A. in Wasserlachen am Rio Primero bei Cordoba. septs ib Tet 1877. G. Hieronymms. (Reed Herb 33910 - as A. magellanica; Herb. U. Coimbra). PERU: In Apr. 20, 1915. 0.F.Cook. & G.B. Gilbert 241. ad ea hi “yh Azolla nilotica Decainse NYASAIAND: Chiromo, Shine River. may 1957. W.8.fe cey. (GHSR- 87005); Dist. Port Herald, Chiromo, floating on water at edge of Shine River, alt. ann ft. Mar. 22, ON igen —_ Rivers, at fi ant 150 18, ear f af 3163 erry, . ft. uh s.C. (Git: Heed Mack’ 5), July 958. 5.0. Seegrtef MOZAMBIQUE: Dist. Gorongoza, on open pool among scrubby palm trees, alt. 300 ft. . 26, 1953. N.C.Chase 5079. (GHSR- —_ Herb. 54732); on md banks, Uruna River and pools, 1 on = ag bank on mud or floating on water, Zambezi River, _ ft. Dist. Buroma Prov. Aug. 7-8, 1950. N.C.Chase 1965 Reed, Salvinia and Azolla 129 (GHSR; Reed Herb. 54731); on surface of maddy pool, Gorongosa Game Reserve. July 14, 1957. N.C. Chase cae and Rt een. UGANDA: Lake Kioga, Namasagali, vase water in sheltered bays and gulleys where it forms a cove ring layer with Pistia Stratiotes. Nov. 8, 1951. Mrs. E.M.Norman-S29. "(Reed Herb. Azolla pinnate var. africana (Desv.) Baker CONGO tems 1, alt. 900 ft. Sur les flaques d'eau embie les mollbaires - Fr. Hens No. 5. (Hb. Univ. Coimbra). ZAMBIA: Edge of Bulosi Plain, below Mongu, Dist. Mongu. Nov. 9, 1959. ED. teieak & A.J.Cookson 6272. Cie): ‘patrese end of Kandaker Island, near Victoria Falls, Dist. Livingstone. July 25, a ay G.F.Cunningham yan Someren. (GHSR GHSR-87308; Reed Herb. MOZAMBIQUE: On lake Nhauvine, Gorongoza Game Reserve. Sept. 27, 1953. N.C.Chase 5080. (GHSR; Reed Herb. 54729). SOUTH WEST AFRICA: ‘Inseln des Ouavange. 39 1900+ H.Merxmller 1970. (GHSR; Reed Herb. any BAROTSELAND: Bulozi Plain, Dist. Mongu. Jan. 10, 1960. W.G .OLlass atte (GHSR; Reed Herb. 54727). Seems to be same locality GOLD coast: Ga surface of wai in se nal pond, alt. 10 m., 16 mi. on Ada Road. P.Cudjoe, Oct. 22, “1951. Collected for C.D. D. Adams 983. Pees Herb. 33007). Bibliography Baltimore Evening Sun, Associated Press. se 38, 1961. p. 3. Helps River Fight Zambezi Power Bennett, F. D. Investigations on Salvinia in pg a British Guiana and Brazil during October to December, 1961. (Unpub- lished report). 1961. Se mitten Salvinia in Trinidad, British and Brazil October to December, 1961. (Unpublished report). e192. rt and peg aareyea te 130 P EE Y-20:).0.6.13.4 Vol. 12, no. 3 Pre report on a second survey of the in- cts attacking Salvinia spp. in South America, February to May, 1963. (Unpublished report). 1963. report on the insects collected in associa- tion with Salvinia in Brazil during February to May, 1963. (Unpublished report). 1964. tests with insects attacking Salvinia auriculata undertaken at Belem during October-December, : lished report). 1965. Carbonell, C. S. Habitat, etologia y ontogenia de acuminata (DG.) (Acridoidea, Pauliniidae) en Rev. Soc. Ent. 6: Lo-k8 1964 Paulinia el Uruguay. Weatherby C. A. A Further Note on Salvinia. Amer. Fern Jour. 27(3): 98-102. 1937. co Sua MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. III Harold N. Moldenke LIPPIA CHEVALIERII Moldenk: more densely so benea very slender, us e above, prominulent beneath; s 8 very slender, or 5 per side, ascending, scernible above, prominulous beneath; vein ry permost 2 or a horter than the nding leaves} peduncles slender, 5-15 mm , densely white-pubescent with $3; heads oblong, cylindric, h--11 ng, mm, » acumina y, tomentose on the back; corolla about ) mm. long in all, its lib t 2 mm, wide. The type of this species was collected by August J. B. Cheval- ier (no. 67) — in whose honor it was named — at Toukota, Soudal, Mali, on December 26, 1893, and is deposited in the herbarium of the Jardin Botanique de l'Etat at Brussels, Vernacular names Te corded for the plant tea" and tts ", In Gambia beehives are smoked with this fragrant herb before being placed "P in trees. It is claimed that the fragrance tims body 1965 Moldenke, Monograph of Lippia 131 In all, 11 herbarium specimens, including the type, and mounted photographs ha ve been examined by me. Citations: MALI: Sitiigidbhid Thierry 227 (K). Soudan: A. Che- valier 67 trectype. F=-photo of type, N--isotype, N—photo of _ type, Si—photo of type, Z--photo of type). SENEGAL: Heudelot 103 (K), 107 (N). GAMBIA: Brown-Lester l (K, N), 28 (K); Dawe 30 (K); Ingram s.n. (K). REPUBLIC OF GUINEA: Caille 1h772 (Br, K). LIPPIA CHIAPASENSIS Sage Verh. Bot. Ver. Brand. 53: 78. ng pevesoureplys: 10 S-, Verh. Bot. Ver. Brand. 53: 78. 1912 n, In ° 13 Known dese. trib. Verbenac., [ed. 1], 17, 20, & 95 (192) and [ed. 2], Ss, & 189, 1949; Moidenke, Alph. List Cit. 3: 669, 919, & 962. ish; M. C. Carlson, Bull. Torr. ae Club 81: 391 & 396. 195k; Molden- ke, _pieont 37, h2, & 461. 1 » to 1.5 m, tall; branchlets * first tetragonal, finally aloes rather pated densely hirt the older ones nat mm. in diameter calvescent; peor bes ad internodes 1,5--6 cm. long ; tbibes decussate-opposite, short—petiolate; petioles ),--7 sl vate, rigidly ito sf 3.5--5.5 cm. long, 1.5--3 cm. wide, acute at the apex hirtous leaf-axils, sabe ioneta: to about 1 ca. wide; peduncles 1.32 cm. long, densely hirtous; basal bractlets aaa cert rather thick, ovate~subdeltoid, rather obtusely subacuminate a2 hirt n the 7 ous o r and densely oped on the outer Surface, about 5 mm. long and 3 mm. wide, densely ciliate, sub- el-veined, + ng bractlets thinner, almost hyaline remaini ; much broader, subreniform-cordate, ahout 5 mm. long and to 6.5 mm. wide, more or less palmately venose, Pere reticulate near the and n: — than the others; fruit similar to a pear seed, about 2.5 * song, 2=-seeded, the cocci easily separating. The type of this species was collected by Georg Edouard and Caecilia Seler (no. 227k) on a hill above Huiztan, in the central the » Mexico, in March, 1891, and was creme in herbarium of the Botanisches Museum a + Berlin, where it eo cca Me Macbride as his type acetates no. 17499, bat te 132 PHYTOLOGIA Vol. 12, no. 3 The species has been found along highways, on slopes Quercus, and on brushy hillsides, at an altitude of ge acme Flowering and fruiting in December and January. The labels ac- Macbride's phototype are inscribed "Sellow 227)" and tear cites the type collection as ayo ps 2274", but these are th palpable errors for Seler & Seler 227k. M. C. Carlson (195h) cites her no. 22h6 from Guatemala as this species. Loesener (1912) says brome Art scheint der L. bicolor Kunth et Bouché nahe zu stehen, die durch l&mgere Ktpfchenstiele, die weniger dicht und anderseits tae drtisig behaart sind, und durch grtssere, leb- haft Involucralblutter von L. Chiapasens. ensis abweicht." In all, 5 pally specimens and 5 mounted phototypes have been examined ° Citations: MEXICO: Chiapas: Breedlove 7895 (Ac), 7929 (Ac); Langman 373 (W--197635); F. Ramirez s.n. es {Diec. 3, 1951) (2); Seler & Seler 2274 ({Macbride photos 17499] (Gere s of type, Kr-- photo of type, N—-photo of type, N—photo o Ug--photo of type). GUATEMALA: Huehuetenango: P. Ce pre ied ey 61963 (N). LIPPIA CHRYSANTHA Greemm., Proc. Am. Acad. Sci. 39: 87. 1903. Bibliography: Greenm., Proc i “ncad. Sci. 39: 87. 19033 Prain, Ind. Kew. Suppl. 3: 104. 1908; , ws Standl., Contrib. U. S. Nat. ab 23: 12h & 12h6. 1924; Moldenke, Knowm Geogr. Distrib. var- » (ed. 1], 17 & 95. aaa Moldenke, Alphe List Cit. 2: 66. 191185 ‘Woldenke, Known Geogr. Distrib, Verbenac., [ed. 2], ae 1949; Moldenke, Alphe List Cit. 3: 683 & "Bo. 19493 "Woldenke, 8 37& 61. 1959, » 1.5--2 m. tall; stems much-branched, covered with gray ish bark; we vinate branchlets hirs rsute-pubescent; leaves decussate- opposite, rt-petiolate; petioles 5 mm. or less in length; leaf- blades pence to lanceolate-oblong, 2.5=-8 cm. long, 1--3 a wide, acute or obtuse at the apex, finely and evenly crenate~ dentate along the margins, cuneate or rounded at the base, regen tendency to become reflexed; inflorescence in a t crowded spikes, wr shorter than the subtending leaves, pubescel with sp hairs ~dentate; corolla a ae mm, long, the tube slender, dark-yellom, externally , gradually ampliate above into the limb, which is yellow or peak Thay pom to white, l-lobed; ovary and style glabrous, The of this rare species was collected by Cyrus G ernsey Pringle (no. 8679) on limestone hills, at an altitude of 1500 “e ters, near Yantepec, Morelos, Mexico, on October 26, 1902, f deposited in the Gray Herbarium of Harvard University at seaprids , Massachusetts. The species has been collected alo ong ra e ant 1965 Moldenke, Monograph of Lippia 133 on limestone hills, in flower and fruit in October. Greenman (1903) says "A species well characterized by the sessile short inflorescences disposed in the axils of the upper leaves. L. Chrysantha is apparently nearly related to L. oaxacana Rob. & Green." In all, 17 herbarium specimens, a gs type material, and 2 mounted photographs have been examined me, Citations: MEXICO: Morelos: Lundell & Lundell 12507 (N, Rf, Rf); Pringle = (Ca--139662—isotype, Cm--isotype, G--type, It- isotype, Me- isotype, Me-~isotype, Me-~isotype, Mi--isotype, N— isotype, “ae? 5a of isotype, Po--63773-~isotype, S--isotype, S— isotype, Vt~-isotype, Z=--photo of isotype). LIPPIA CIPOENSIS Moldenke, Phytologia 6: 6. 1959. Bibliography, Moldenk enke, Phytologia 6: 6. 19595 Moldenke, Ré~ sumé 425 & 161. 1959; Moldenke, Résumé Suppl. 1: 6. 1959; Hocking, Excerpt. Bot. A.5: hi. 1962, Shrub, about 1 m. tall; stems and branches slender, gray, densely appressed whitish-strigose with antrorse hairs, twiggy; branchlets and twigs very slender, densely pee nk with ie tly lig 6 mm. long, 23.5 mm. wide, abruptly acute at the apex = base, e The Henrique Lahmeyer de Mello Barreot and Alexandre vk (no. Uh) at km. 142, serra do Cipé, Minas Gerais, ary on April 16, 2935, and is deposited in the herbarium of the Botanisches Wuseum a t% Berlin. The plant has been collected in flower in A~ Pril and June. In all, 5 herbarium specimens, including the type, have been ed e. Citat ete: BRAZIL: Minas Gerais: Brade 13899 [Herbd. Rio de Jan. 29525] (B, Z), 13900 (Herb. Rio de Jan. 29526) (B), 13901 [Herb. Rio de Jan. 29527] (B); Mello Barreto & Brade shh THerb. Rio de Jan. 29521] (B-type). 13h PHYTOLOGIA Vol. 12, no. 3 A CONTERMINA Briq. in Chod. & Hassler, Bull. Herb. Boiss., wey 2, lz 157." 190s e Synonymy: Lippia turnerifolia var. camporum Griseb., Pl. Lor~ entz. pir 187k. Bibliography: Griseb., Pl. Lorentz. 195. 1874; Briq. in Chod. & patel er, Bull. Herb. rg sér, 2 1157. pees Briq. in Chod. & Hassler, Pl. Hassler. 2 (11):" 493. 190); Prain, Ind. Kew. Suppl. 3: 10h. 1908; T. Mey., Rev. Argent. Agron. 2: 356. 19 36; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 20 & 95 (19h2) and [ed. 2], 99, 10h, & 189. 1919; Moldenke, Phytolo cae 3: 75 & 76 (1919) and 3: 290. 1950, Woldenke, Résumé Ly 1959; Troncoso, Bol. Soc . Argen ent. Bot. 9: 18h. “eg Troncoso, pardon 12: 258, 69-2 Tl, & 275, fig. 6. 1961; reste Rée- 12: os: 5. Pte Troncoso, Darwiniana 12: 270, fig. 6. 1961. mal or perennial herb, medium-sized, 0.2--1 m. tall, glut- inous, dioecious; rhizome creeping; gyre stems simple or slight~ ly branched, mostly 30--50 om. tall, tetragonal, viscid- a glandular-pilosulous throughout; middle internodes h—-6 cm leaves decussate-opposite, the lower ones reduced; petioles t 2 em. long, viscidulous, glandular—pilosulous; lower leaf-b. va e on both surfaces, rather rigid, subacute at the apex te con~ vex toward or below the middle, prolonged at the base into the S species was based on two collections made by Bnil H assler in Paraguay —~ no. 912 from thickets at Fort Lépez, in September and no. 6941 from the campo near Valenguela in the valley of she The species has been found on campos, riverb the edges of woods, at altitudes of 300 to 1000 meters gee trea _ 1965 Moldenke, Monograph of Lippia 135 tember to verges and in April. Meyer refers to it as a very com- mon plant and frequent in high campos in the tye A vernacular name recorded for it is "salvia amarilla". The corollas are de- scribed as yellow on Meyer 13h, Osten 91080, and Venturi 7559 7559, but as orange on Schwarz 3575~ as "Pl. nee? . 2: 493. 190k" "Plantae Hasslerianae .m (190k) 93" Briquet says "Le L. ein et est assez voisin c apparence du L. modesta Briq.; il en différe en premiére sper par son indu- ment, glanduleux ux qui recouvre les pédoncules, les on et les petioles. Ce caratrére rapelle le L. asperrima, dont il s'ecarte par la forme des feuilles, les capitules deux fois plus petits, les bractées plus larges i indument non strigneux etc." siliensis Link. However, it is based on a specimen collected | by Paul Gttnther Lorentz (no. 5h) near Pozo del Alto, Tucum4n, Argen- tina, on November 22, 1871. Miss Troncoso has examined an iso- type and has verified its conspecificity with L. contermina. She says: "Especie ates en los ejemplares Hassler | 912 y 69h. Es- tudiados detenidamen laos feet or de Ginebra, | he elegido como lectotipo Hassler 69h1, tratarse de un ejemplar m4s completo, Y coincidir en todo con ia aitaogits de Briquet. apse conter- es una especie dioica, el lectotipo Hassler 6941 es un ejem- plar © y consiste en una rama joven en Baecst, a ejemplar Hassler 912 del Herbario de Ginebra, a primera vista bastante diferente del tipo, parece ser sin embargo, ea una rama ro=- busta, sin es, de un pie 6 de la especie. @jemplar que he podido consultar en los herbarios de Kew y British en omo en todas las especies atedoke citadas en esta nota, el pistilo no es funcional y las flores caen sin fructificar." She States that this species is very closely related to L. modesta Briq., differing in the pubescence: "L. modesta presenta pelos estrigosos en tallos, hojas, pedtinculos y y brdécteas, a veces peli- tos glandulosos muy breves en el &pice de los pedfinculos y de las brécteas. L. joemins Ageing ee tallos, ct braécteas, P), 69h (Bm—~cotype, Cb--cotype, K-—-cotype). ARGENTINA: Chaco: =: Meyer 13h, (Si); Rodrigo 2707 (Si). Corrientes: Sota 1035 (31). 136 PETTOLOGIA Vol. 12, no. 3 Misiones: Schwindt 189 (Ml). Tucumén: Lorentz 54 (Cd); Venturi 531 (Si), 1375 (K, (K, Si), 7359 (K, Si). She states that Hassler 69h1 and Venturi 1375 & 7359 are pistillate; Hassler 912, Meyer 13k, and Rodrigo 2707 are staminate; and Venturi ari 531 is a mixture of staminate and pistillate. Hassler 2359 and Venturi “531, 1375, & 7359 have been formerly regarded b; by me me as representing L. asperrima C Cham. Other specimens cited by me as that species, and not now available to me for com- parison because of ing been returned to herbaria in various parts of the world, rd also actually be L. contermina. More work pee to be done on the L. asperrima, L. contermina, L. modesta, L. morongii, L. turnerifolia, L. villafloridana group of species. The Vienna specimen of Hassler 2359 bears a label originally read- ing "Mimosa bimucronata var. hexandra f. viperes n. f.", but this is doubtless a case of aE cage iga 5 herbarium specimens and , mounted photographs have In been examined by me. Citations: PARAGUAY: Hassler 2398 (Cb, V—1095); Osten 9179 (Ug), 9180 (F--photo, N, N--photo, 8, Si-~photo, Ug, 7Z——phote) « ENTINA: Chaco: T. Meyer 134 (Herb. Osten 22669] (Ug), 2558 (Herb. ols 22948) (Ug). Misiones: G. J Schwarz 3575 ihn of cumén: Venturi 531 (W-~159122h), 1375 [ (Herb. Inst. Miguel 32167; Herb. Osten 17237] (N, Ug, W-—15912h0), 7359 (eg aaeOL. LIPPIA CONTROVERSA Moldenke, Phytologia 1: 23--l2h. 19h0. weoacoaa : Lippia pinetorum Moldenke, Phytologia 2: 20--21. bliography: Moldenke 4 eee 1: 23--h2h (1940), 1: 50b own Geo et ° asia), and 2: 20--21, 19h Moldenke, gre D ed. 1], 17, 20, & 36. ies I mervet Phytologia 2: 3a. 3. 19L7; E. J. Sali ye Ind. - 10: 13h. Bs ff Mol- enke, Alph. List Cit. 2: 5% as ona) ot nh, ; 960, &. 962 (ioh9), and 4: 999 & oe 19h9; Moldenke, Known Geogr. Dis- trib. Verbenac., [ed. 2], 31, 35, 37, 39, 189, & 190. "1919; Mol~ 6, Phytologia 3: 73. sane tu Am, : . 3 Mold . Moldenke, Phytologia 4: 68, 1952; E. J. Salisb., Ind. Kew. Suppl. 11: 138.1953; Moldenke, Phytologia ©: 9E. 195); Moldenke, Résu- es a - bh, 47, 316, & h61. 1959; Moldenke, Phytologia 12: — 1965 Moldenke, Monograph of Lippia 137 thick-chartaceous, uniformly gray~green on both surfaces or dark- sia above and lighter beneath, often brunnescent in drying, the lowermost 8—10 mm. long and 5--6 mm. wide, acute at the a- pex, densely bikie oe Weare or puberulent with brownish gland- tipped hairs and scattered-hirsutulous with longer white hairs; Corolla cream-colored, Decin p ta d pale greenish-yellow, or “right-yellow, hypocrat type of this cactnton’ was collected by Paul Carpenter Standley (no. 58067) on a brushy slope near Las Lajas, at an al- titude of about 1200 meters » Sacatepéquez, Guatemala, on November » 1938, and is deposited in the Britton Herbarium at the New York Botanical Garden. The type of L. pinetorum was collected by Eizi Matuda (no, 3925) ina pineland on Mount Ovando, Chiapas, Mexico, ned November 1) and 18, 1939, and is deposited in the Same herbari The tex are “has been found on brushy slopes, in pinelands and damp thickets, and in stony fields, at altitudes of 500 to 1300 meters, fl and fruiting in November and December. Stan Fa i 8 Be g gah ek. L. cardiostegia cage L. substrigosa Turez., L. umbell , and nutans wh, N, RE). GUATEMALA: Escuintla: P. C. Standley 58223 (N). Jutiapa: Pe C. Standley 77587 (N). Sacatepéquez: P. C. Standley 58067 (N-- ‘ype). Santa Rosa: | Heyde & Lux 1,387 (C, G, N--photo, Z——photo) ; E.G. Standley 77777 (N). HONDURAS: Morazdn: P. C. Standley 11183 138 PHYTOLOGIA Vol. 12, no. 3 6; ~ Ag i KMoldenke & Moldenke 19813 (Es, F, Fy, Lg, Mg, > 8 , Sa “COSTA RICA: Puntarenas: H. Pittier Ulid9 ter}; Taciant uheo Sm eee var. BREVIPEDUNCULATA Moldenke, Phytologia h: Bibliography: Moldenke, Phytologia : 56 & 68. 1952; Moldenke, Biol. cgi 26: 1471. 1952; Moldenke, Seer 37 & 461. 1959. variety differs fron the typical form of the species in having ie peduncles under mature heads only 1 cm. = or less. It is a shrub about 1.5 m, tall, the flowers opening bright- yellow, fading to orenicblc or in age. The type of the variety was collected by Edward Johnston Alex- ander (no. 192) on a riverbank at Nizanda, Oaxaca, Mexico, and is deposited | in the Britton Herbarium at the New York Botanical Gam den. The taxon is know thus far only from the original col lection. In all, herbarium specimens, including the type, have been exam Citations: MEXICO: : E. J. Alexander 192 (N--type, N— isotype, W—isotype, Sep peut LIFPIA CORIACEA Briq. in Chod. & Hassler, Bull. Herb. Boiss., - 2, k: 1160—1161. 190k. : Lippia coriacea f. angustifolia Briq. in Chod. & Hassler, Bull, Herb. Boiss., Boiss., sér. 2, : 1161. 190}. Lippia cori- acea f. latifolia Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér. 2 aoe 1161. 1904. Lippia coreacea Briq. ex Moldenke, Résumé 31, in syn. 1959. Bibliography: Briq. in Chod. & Hassler, Bull. Herb. bt sér, 2, h: 1 160—1161. 1904; Briq. in Chod. & Hassler, P. Hassler. 2 (11): 496. 190k; Pr . Kew. Suppl. 3: toh: i908; Moldenke, Suppl. oe t Invalid Names 5. 191; Moldenke, Know Geogr. Distrib. Verbenac., [ed. 1] v hl & 95. at Moldenke, Alph. List Invalid Names 30. 1942; Moldenke, Lilloa 8: Le peg. Moldenke, Ai List Cit. 1: 39, 263, & 264 (1946) and 692 & 693. 199; Moldenke, Known Geogr. Distrib. Verbenac., fet. 21, 2 & 189, 1349; Moldenke, Résumé 116, 311, & 61. 1959; Trom Bol. Soc. Argent. Bot. 9: 18h. 1961; Troncoso, Darwinians 12; 258, 271, & 272, fig. 7. 1961; Moldenke, Phytologia 12: 2h. Illustrations: Troncoso, Darwiniana 12: 272, ni 7. 1961. Medium-sized shrub, 0.5--1 m. tall; branches erect, obtusely n nce c te ous-pubescent, the pubesce omposed of & few antrorsely strigose hairs and short cri sp: liferous ones 3 e internodes 2--5 cm. long; leaves osite or often approximate, subsessile, pba or elliptic-lanceolate to lanceolate, —6. - long, 1--2. » acute or short-ac te at the apex, entire or Sapértictal ly hicclatecadntate ‘touar toward the apex, revolute and somewhat convex along the margins, rounded-exte enuate 1965 Moldenke, Monograph of Lippia 139 at the base, thick-coriaceous, shiny above, paler green beneath, ovoid-subglobose, about 1 cm. long and wide; outer bractlets ovate, acuminate at the apex, more or less nigrescent in inner surface of the cocci excavated. This species was based by Briquet on two collections made by Bil Hassler in Paraguay -- no. 73 from sandy places near the two collections is also the type collection of Briqet's f. angus- tifolia and has been designated as lectotype of the species by Miss Troncoso. The second collection is the type of f. latifolia. The species is placed by Miss Troncoso (1961) in her Subgenus Lippia, Section Lippia, and Series Axilliflorae. She says: fototipo del Museo de Chicago No. 2650 comprende les dos formas. He seleccionado como lectotipo Hassler 73, cuyos detalles he dibujado. Especie dioica. Los dos ejemplares estudiados del Her- bario de Ginebra son femeninos y se hallan en floraci6én. No he Visto frutos," Briquet's original publication is sometimes cited as "Briq., Plantae Hasslerianae XT (190k) 496" or as "Briq., Plantae Hassler. ad ces, dures, luisantes en dessus, 4 marges enroulées en dessous, to me only from 10 mounted photographs representing type material of all the names involved. a eyed PARAGUAY: Hassler 73 [Macbride photos 2650, in Part] (It—photo of cotype, Kr--photo of cotype, N--photo of co- type, N—photo of cotype, W--photo of cotype), 6798 (Macbride Photos 24650, in part] (It--photo of cotype, Kr--photo of cotype, 140 PHYTOLOGIA Vol, 12, no, 3 N--photo of cotype, N—-photo of cotype, W—photo of cotype). oe CORTMBOSA C » Linnaea 7: we 1832. onymy: Lippia vase perm Mart. ex Hiern, Vidensk. Meddel. pean n. 187TaB 99, in syn. B9T: bliography: Cham., Linnaea 7: 219. 1832; Steud., Nom. Bot., Bs 2, Ey Sh. 18)0; D- "Dietr., Syn. Pl. 3: 597. 1843; Walp., Re- - Syst. hk: Sis. 18455 Schau in A. e, Prodr. li: 590. “187; Schau. in i Fl. Bras. 9: 245. 1851; Hiern, Vidensk, eddel. Kjgbenh. 1877-18787 99. 1877; Jacks. in Hook. f. & Jacks., te. Kew. 2: 95. 189; Glaz., Bull. Soc. Bot. France 58, Mém. 3: 542. 1911; Moldenke, kooan oe pk Patras stat i [ed. bps & 95. 1942; Moldenke, Alph. List s 14. 197; Moldenke, Alph, List Cit. 2: 4b fait 94,8), a: oh, ane é "91s (1995, and ): roe 1203, & 1204. 1949; Mold , Known G eogr. Distrib. Verbe: ac., [ed. 3], 80 & 189. 1949; pa Résumé 93, 1959; Renné, Levant. ect Inst. Agron. Minas 150. 1960; ‘ideas, hirtous ou a s below, simple, corymbosely twiggy above; leaves ternate, rather rigid, very short-petiolate; leaf-blades ovate, acute at the apex, te-rugose, rather shiny and strigose- se-hirtellous a- ones on t about 1.6 cm 1 cm. wide, the upp aller; inflorescence corymbose-paniculate, canes~ cent with whitish hairs; hea ; pedunculate, solitary in the leaf-axils, equaling or surpass the subt aves, congested at the tips of the branchlets, about 1.3 an. long anthesis, lax, finally much elongated and subracemose or spike= like; bractlets membr ranous, somewhat colored, lanceolate, 3j-nerved, » alm qual the corolla reddish or rose, almost 8 mm. long, villosulous on the isa at the tube ventricose on the outside, the limb rath- large, lig velutinous on the outer surface, une h-fid, chat dorsal lobe Linge cee lateral a truncate. spec Ber. ed, although Chamisso says simply "E i orp *misit Sellowius nos- ter." The type of L. monticola was collected by Carl Friedrich Philipp von Martius (no. 1035) at or near one do Campo in the sane state, in March, 1839, deposited in the Martius Herbarium 4 ot Munich, where it was photographed by Macbride as his type graph no. 20323. The species is classified si Schauer in his See- tion Zapania, Subsection Corymbosae. It has been collected in anthesis from November to January and in March. Schauer (1851) cites also P. Clausen s.n. and Lund s.n. from Itabira do Camp? and Riedel 3.n. from sandy wet places in the Serra da Lapa. 1965 Moldenke, Monograph of Lippia 141 Material of L. corymbosa has been misidentified and distribu- ted in herbaria under the name L. pseudo—thea Leut. Glaziou (1911) cites also his nos, 1773, 19713, and 19714 from Habira do Campo and Pinhei near Diamantina, Minas Gerais, and describes the plant as "Frutescent, fls. roses, jan.--mars." He reco the vernacular name "c "de pedestre",. In all, 14 herbarium specimens and mounted photographs, in~ cluding type material of one of the names involved, have been ex- ed by me Citations: BRAZIL: Gofas: Glaziou 21895 (Br, N); uacede 3492 (N, S, S, W--2197143). Minas Gerais: Black, Magalh%es, & Graf- Linger 51-12217 (Be--700hh); P. Clausen 64 (Br); artins eo [Macbride photos 20323] (Br, It--photo, Kr—photo, N—photo, W—~ photo); Mello Barreto 10703 [Herb. Jard. Bot. Bello Horis. ia (N); Murga Pi: Pires & Black 3348 (Be--63792); Regnell s.n. Mus. Nac. Rio Jan. 14857) (W--1199338); Schwacke s.n. tsa-1h888) 5 Ule 2669 ( Ja-~14882), LIPPIA COSTARICENSIS Moldenke, Phytologia 1: 42h—l25. 19)0. Bibliography: Moldenke, Phytologia 1: a—I25 COW and 1: 50k. 1941; Moldenke, Known Geogr. Distrib, Verbenac., alisb., « Kew, Suppl ke, Alph. List Cit. 3: 90 (1949) and \: 999. 199; Known Geogr, iis teib. Verbenac., [ed. 2], 39 & 189. Shi a icky, Stern, & Chambers, Trop. Woods 109: 78. 1958; Moldenke, R sumé Ul, hy & 61. 1959. e, to "18 m, tall; trunk to 33 cm. in diameter; branchlets stoutish or rather slender, aatcly 6 tragonal, often rigs maa or less substrigose-pilose; nodes annulat conspicuously so; principal internodes 2=-;.5 cm. long; Laie de- cussate-opposite, with a mint-like arama; petioles slender, 1—2 cm. long usually short, more or less appressed-strigose; leaf- Wikdea © chartaceous x rather uniformly green on both surfaces, nar- c The type o this spe cies was collected by Alexan Frank a (no, 2292) at an altitude of 915 meters in ee yaninity of eneral, San San José, Costa Rica, in December, 1935, and is depos~ 142 PHYTOLOGIA Vol. 12, no. 3 ited in the Britton Herbarium at the New York Botanical Garden. The species has been found in forests, clearings, and on volea- no slopes, at altitudes of 915 to 2000 meters, flowering in June, August, and December, fruiting in December. A vernac name recorded for the plant is "caragra", Material has been misiden- tified and distributed in herbaria under the names L. myrioceph~ ala Schl. and "Lippia (near umbellata Cav.)". “~Jn all, 11 herbarium specimens, ne the type, and 4 mounted photographs have been examined by m Citations: COSTA RICA: Alajuela: Skutch 3097 (A, N, S). Car- tago: J. A. Echeverria 6 (Ca—776208). San José: E. W. D. Holway 307 iis? are 2292 (F—-photo of isotype, “G——isotype, N= type, N——photo sotype, e, S--isotype » Si—photo of isotype, Z— photo of pat: the a ee 1262 (Br). PANAMA: Chiriqui: Stern & Chambers 88 (A, BY ao. LIPPIA CURTISIANA Moldenke, bie 1: 425-26. Bibliography: Moldenke, Phytologia 1: 1 =-—\26 he hig fe pp 1917; H. N. “Py a Moldenke, Pl. Life 2: 55. 1918; Aiph. List Cit. 2: + WT (a9k8) anc 3: 775, 786, 962, ‘ 73, 3. 13g enke, Known G Dist: erbenac., [ed. 3 Pe # Ye 19195 We Moldenke, Résumé at iz "& “Wel. 1959. hrub, 1--2 m. tall, eae odor of Lantana; branches and branchlets melty a sely tetrago. er onal, onal, densely short- pubescent with sordid appr sna hairs glabrescent in age; nodes pal ™m densely short—pubescent with appressed so: airs; leaf-blades eous or thickish-c eous, often falcate and condupli- cate in drying, rather unifomly sh-green on both surfaces 8 ng teeth from almost the base to the apex and usually pine ait Neeechish ty acute or rounded at the base, strigos cabrous-bullate above, short-pubescent beneath ade along the veeutson midrib very slender, subimpressed above, P lous beneath; secondaries very slender, 8 per side, “ascending, not h rominulo 5 e type o. f this species was collected by Edward Palmer (ne W79)" at the city of Durango and its vicinity, at an altitude 1965 Moldenke, Monograph of Lippia 143 6207 feet, Durango, Mexico, between April and November, 1896, and deposited in the Britton Herbarium at ap New York Botanical Garden. The one is named in honor and grateful appreciation of Dr. Carleton Clarence Curtis (1862-195), for many years pro- fessor of botany at Columbia University, author of valuable texts, inspirational teacher, and valued friend and advisor of scores of the present generation of American botanists, who will a honor and revere his memory. The species is known from the margins of spring-fed swamps, Wet meadows along streams, in pine-oak riccbensa pine forests, sr open limestone hillsides at altitudes of 1500 to 3000 neters, in J be used as a condiment at meals in Durango. Material has been misidentified and distributed in herbaria as L. umbellata Cav. Steyermark 36533 is placed here tentatively. In all, a herbarium specimens, including the type, have been examined Bats ons | MEXICO: Durango: Ochoterena s.n. [Octobre 1910] (Me); Edw. Palmer 79 [April to Nov. 1696) (Ca—1Ol9kl—isotype, ieee “Gis ype G--1s0 ype, Me-—isotype 64105—isotype) . ge E pn beret P. Ce Standley 91197 91197 (N). Huelmetenango: 2044 (N). San Marcos: ea rmark 36533 ( eae ee P. C, Standley L076 (N). LIPPIA DAUENSIS (Chiov.) Chiov., = Somala 2: 359. 1932. : Lantana dauensis , Ann. Bot. Roma 10: 402. igl2, Sonia ath ock ellenbeckii Loes, ex A pe Résumé Suppl. 1: 19, in syn. rind Lippia ellenbeckii var. pinnatifida Loes, ex Mol- denke, Résumé Suppl. 1: 19, in syn. 1959. : Bibtiography: Chiov., Ann. Bot. Roma 10: a t nd. CW he . Somal § Ao He Bilt, Ted. vb 147 er Chiov., zs choos Be » minute, rigid bristles which are and abruptly bent upwards; leaves decussa teenies or ternate; peti- cles short, 2-~l, mm. long; leaf-blades lanceolate, l--5 cm. long and (not including the teeth) 6—10 mm. wide, rather broadly acute pr obtuse at th gradually at mucronulate, the petiole at the base, nigrescent in drying, glabrous and smooth » Slabrous beneath, but with sparse and short, yt antrorse- ly recurved, bulbous-based bristles on the venation and deeply dentate-lobulate along the margins with 6-8 triangular- BENT PHYTOLOGIA Vol. 12, no. 3 linear teeth which are acute and divaricate, the median ones to 6 mm. long; midrib and secondaries impressed abo ove, prominulous be- neath; inflorescence solitary, at first globose and )--5 mm, wide, eventually cylindric-elliptic, 10 mm. long, and 4.5-—5 mm. wide; peduncles slender, 2 or 3 per node, 3—6 cm. long, tetragonal, ‘ ° subapiculate at the apex, densely pubescent; calyx -pAhing bi- lobed, densely short ~pilose on both surfaces with flexuous white hairs interspersed with sessile yellow glands, about Cee mm. long and e during anthesis, not venose, very closely appres ssed to the nak. the lobes very small, rounded, very deeply split above; corolla hypocra teriform, white even when out 1.5 mm. long, th pe Ruspold (a8 1613 (1054]) in "Ogaden: sulla sponde del Daua a Veldi", Ethiopia, on April 21, 1893. Chiovenda placed the species in Section Sarcolippia of Lantana and says "Specie rilevante per yi | —— assai sp , Spugnoso, per il quale si differenzia dalla specie di Lippia che pres sentano invece il pericarpio sot- tile poco sensibile. Le foglie per la loro dentatura ricordano quelle del Lycopus oe Cufodontis a oe oe any "locum typi cl. auctor errore in gaden putavit." ) places the type locality in "Sama~ liland". Lippia rerehlots var. pinnatifida is based on Ruspoli & Riva 1459 from Ethiopia, deposited in the herbarium of the Botan- isches Museum at Berlin. This specimen also has the numbers W054" and "1613" on its labels, apparently indicating its conspecificity with the type of L. dauensis. Cufodontis gives "105" as a supple mentary number for the type. type. Lippia dauensis has been found growing on lava plains, at an al- titude of 3200 feet. A vernacular name recorded for it is "rehan"- = _ said to be of value for an essential oil. Material = tributed in herbaria as "Lippia cf. L. carviodora In all, 5 herbarium specimens, ooh ed As material of one of the ry oe ” involved, mae been examined B tions: ETHIOPIA & Riva Z). KENYA: Jo Be Gillet 13928 (B, Ss, z). ‘pom abs @, 2) nie re Glaz., Bull. Soc. Bot, France 58, Mém. 3 Bibliography: , Bull. e 58, : 542. 1911; Prain, Dd eons ” suppl. e isn. igai; hs Moldenke; ee Geogr 1965 Moldenke, Monograph of Lippia 145 Distrib. Verbenac., [ed. 1], 37 & 95. Ba. Moldenke, Alph. List Cit. 1: 238 (196) and 3: 731. 19493; Moldenke, Known Geo eogr. Dis- « Verbenac., fed. 2], 80 & 189. Pare Moldenke, Résumé 93 & 461. 1959; Renné, Levant. Herb. Inst. Agron, Minas’150. 1960. Conspicuous, impressed above, prominent beneath; veinlet reticu- lation mostly indiscernible above, the tertiaries prominulous be- neath; inflorescence axillary, in the upper leaf-axils only, 2 per node, greatly nig a leaves, showy; peduncles pose Slender, 1--2.5 cm. long, densely s hort—pubescent with spreading hairs; heads subglobose, iene cm. long and wide, ge es many- flowered; bractlets membranous » very large and conspicuous, red- ? rather sparsely short-pubescent on both sur surfaces; corolla hypo- craterifoma, lavender or bright~pink, its tube slender, about 8 mm. Fry spreading-pubescent on the. outside, ampliate at the a- Pex, the limb about 7 mm. wide, venose, very ’ sparsely pilosulous Kea ee beneath, glabrous above; stamens and pistil in- The “type of a rare species was collected by Auguste pid Marie Glaziou (no. 19712) on a campo at Curalinho, near Diamantina, Minas Gerais, Brazil, in 1892. The original publication nigel states n. sp.t in herb. Paris., Berol., Kew., os Ste rux ralinho, prés Diamantina, dans le campo, MINAS, no. ign2. aioe cent, fl. roses, Mars-avril. R." The species is said to occur oc- casion nally and scattered in open gravelly pastures and on sandstone Slopes and and summits, from 900 to 1170 meters altitude, flowering in , June, and August. Material has been misidentified and distrib- uted in herbaria as as L. gardneriana Schau, On the o —Tkgraf, Brade, Mello Barreto & Mendes Magalhties 3273, dintributed as L. ai L. Hanantinensis, is actually L. hederaefolia Mart. & Schau. Th herbarium en, including the type collection, have Goes examined by m 146 PHYTOLOGIA Vol. 12, no. 3 Citations: BRAZIL: Minas Gerais: Brade 138 Laas Rio de Jan. 29529] (B); Glaziou 19712 (N--isotype); A. Lutz 1096 (Z); M Mendes Magalh&es, & & Maguire 49255 (N); Mexia s8 505 8 (Gg--286543, Go, N, S). LIPPIA DOMINGENSIS Moldenke, Phytologia 2: 31)--315. 19h7. Bibliography: Moldenke, Phytologia 2: 31y—315. 1917; Moldenke, Alph. List Cit. 2: 569. 198; Moldenke, Known Geogr. Distrib. Ver- benac., [ed. 2], 48 & 189. 199; E. J. ae Ind. Kew. Suppl. 11: 138. 1953; Moldenke, Résumé 57 & 161 . 1959. 3 ly bromish-puberulent, resinous, glabrescent in age; branchlets and twigs numerous, very slender, tetragonal, densely brownish- puberulent, resinous; nodes annulate; principal internodes care reer 3-20 mm. long; leaves decus sate-opposite or ternate, n very small; petioles very slender, 1--2.5 m. for ote {Mlose“piberulent, resinous; leaf-blades very small, her thick- een above, somewhat lighter omen oma or sthorbicilar, 15 mm. long, 3-11 mm. wide, rounded at the apex, t the base, deeply bullate above, regularly dentate with h te the v ender, , rather straight, deeply impressed above, very P ben alee bat nghtay very verry connecting the secondaries and at right angles to them, rather straight and subparallel, deeply impressed above, prominilent beneath; inflorescence axillary, sparse, less pr 1 cm. long in cae about > the subtending fruit, only flowers 6 lanceolate, 2.5--3 mm, long, 1 mm, wide, obtuse or subacute at the apex, resinous-puberulent; corolla white, its tube 3}-l mm. long, oes surpassing the subtending bractlets, the limb 1~1.5 m.- The type of this species San collected by Richard Alden ier and E. S. Howard (no. 8110) at the aie of a limestone ravine in inewoods along the trail between Pedernales and Aceitial, Bs ang altitude of 4200 feet, in the province of an Re~ is thus far knom only from the type locality. In all, of — specimens, including the type, have been : Dominican Republic: Howard & Howard suo. veteen. B25. i), 8192 (N, S). LIPPIA DRACOCEPHALOIDES Turez., Bull. Soc. Nat. Mosc. 36 (2) 205° 1965 Moldenke, Monograph of Lippia 147 Bibliography: Turez., Bull. Soc. Nat. Mosc. % (2): 205. 1863; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 95. 1894; Moldenke, ri nace, [ed. 2], sumé 93 & 61. 19595 Moldenke, Résumé Suppl. 8: 2. 196, iS} 3 stems rather acutely tetragonal, pubescent—hi leaves decussate-opposite or ternate; petioles very short; leaf- blades linear-lanceolate, acuminate at the apex, attenuate at the base, serrate along the margins except at the base, pubescent on both surfaces; inflorescence axillary, capitate, the heads (espec- ially the lower ones) long-pedunculate, nodding; bractlets ovate, te at the apex, fuscescent. The type of this distinctive species was collected by George Gardner (no. 433k) type in the Delessert Herbarium at the Jardin Botanique in Geneva collection, and ) mounted photographs have been examined by me. Citations: BRAZIL: Brasilia: Murga Pires, Silva, & Souza 9652 (2). Gofas: G. Gardner 433) [Macbride photos 26h8] (It—photo of isotype, Kr—photo of isotype, N—isotype, N=--photo of isotype, W--photo of isotype). LIPPIA DUARTEI Moldenke, Phytologia 10: 170--171. 196). Bibliography: Moldenke, Phytologia 10: 170—~17l. 196k. A a perennial plant with rather strict stems or ted a. Sate-opposite, the lower ones often with abbreviated leafy non- floriferous twigs in their axils; petioles slender, conspicuously Slongated on upper mature leaves and there about 1.5 - long, e hairs very den; c c Straight and antrorse; leaf~blades chartaceous, dark-green above, ; cm when wate-acute at the apex, rounded at the base, uniformly crenate a- ’ ppre a3 heric, many-flowered, 5--10 mm. wide; bractlets ovate, the outer about 2 m, long and 1 m, wide, acute at the apex, appressed- srulent, 148 PHYTOLOGIA Vol. 12, no. 3 The type of this curious species was collected by Apparicio Pereira Duarte (no. 7851) -- inwhose honor it was named -~ at phen da Palma, Faz. M#e d'Agua, Minas Gerais, Brazil, on April » 1963, and is deposited in the Britton Herbarium at the New “hs Botanic al Garden. The species is known to me tms far only aor hee type specimen tations: BRAZIL: Vinas Gerais: Duarte 7851 [Herb. Bradeanun 27896] (N-type). LIPPIA DUMETORUM Herzog, Meded. Rijksherb. Leiden 29: 5. 1916 Bibliography: Herzog, Meded. Rijksherb. Leiden 29: 5. 1916; A. W. Hill, Ind. Kew, Suppl. 6: 117. 1926; Moldenke, Known ee Distrib. Verbenac., [ed. 1], 0 & 95. 192; Moldenke, Lilloa 8: bau (292) and 10: 378. 19h Moldenke, Alph. List cit. 2: 535 (1948) and 3: 692. 1995 Moldenke, Known Geogr. Distrib. Verben- ac., [ed. 2 7 & 189. 199; Troncoso, Darwiniana 10: 72. 19525 ise Résumé 11) & 61. 1959; Moldenke, Résumé Suppl. 11: 5. anched shrub, 3—-l; m. tall; yo oe prinehes obtusely angular Pn sulcate, loosely poe een eaves decussa eate-opposite; petioles medium in t , 8-1 mm. long; gab chartace- ous, ovate-oblong, eras cm. long when ma » 2-5--2.8 cm. wide, subacute at the apex, crenate along subobtuse and bobt contracted at the base, reticulate and glandulose on both surfaces, softly ee aeocest4 beneath; inflorescence spicate, densely fasciculate in the leaf-axils, short—pedunculate, erect, usually at the apex of the branches, mith a paniculate aspect; peduncles 4-5 mm. Longs spikes narrowly oblong, l-farious, about 1.5 cm. long (including the Rrsaot ns teatiiie imbricate, Srekeas ovat o~ triangular, acute at the apex, cancave, 3--l, mm. long and 3 mm. wide, equaling the corolla-tube, short-villous; calyx scneee? compressed, narrow, puberulent; corolla-tube about 3 mm. long, somewhat dilated at the middle, minutely puberulent ee corol- la-limb 2-lipped, crisped. type of this poorly understood species was collected by Theodor K. J. Herzog (no, 1851) at an altitude of 1600 meters at pu Lillo as "Herzog, Bolivia ITI (1916) 5". The species is nom tims far only from the type collection. For a comparison of this species with L. hickenii Troncoso, see the latter species in these 2 herbarium specimens and }; mounted photographs of the Citations: BOLIVIA: Province undetermined: Te K. Je Herzog 1851 [Macbride photos 2649] (It--photo of isotype, Kr—photo of test tll ype, N--photo of isotype, S—isotype, W--photo 1965 Moldenke, Monograph of Lippia 1h9 LIPPIA DURANGENSIS Moldenke, Phytologia 1: 426. 190. Synonymy: Lippia pennellii Moldenke, Alph. List Cit. 3: 786, byponym. ee Lippia durangense Moldenke, Résumé Suppl. 6: 10, in syn. ae Bibli sae, Moldenke, oe ee a | Rssstivall (1940) and 1: 50. 19a Moldenke, Known + Dist erbenac., [ed. 1], 17 & 95. 1942; Eads Salisb., mgd Kew. hae 10: 134. 1947; Mol- denke, Alph. st Cit. 3: 786 & 807 (1949) and h: 1071. 19) 3 somenae ane 3: 132. 1949; Mold eogr. Distrib, Verbena & 189. 1949; Moldenke, Résumé 461. 1959; Soldeaxs? Foie Suppl. 6: ) & 1 3) and 8: 5. 196). » 1h); » Slender; branchlets slender, rounded and often appressed teeth, acute or acuminate at the base, very rough-scabrous above, rather’ densely puberulent and resinous— Punctate beneath; midrib slender, plane or subimpressed above, c 3 peduncles very slender, 1—-3.5 » 2--5 or more in each leaf-axil, more or less densely Seger et ago with strigose whitish hairs, o 80 te; heads 6--9 Tejamén, Dy yeh s Mexico, between August 21 and OT, 1906. Lippia du rangensis has been found in oak woodlands and grama Srasslands, on a volcanic rim in grama grassland, and in volcan- de g from August to Oc Octo- ber, fruiting in March and august to October. _ Waterfall found it *n a stony hillside with grass, cactus, and a Material has been misidentified and atetinted in herbaria as saitana velutina Mart. & Gal. On the other hand, the H. S. Gen try 1222, distributed as Lippia durangensis, is actually L. umbellata 150 PHYTOLOGIA Vol. 12, no. 3 In all, 15 greicdess specimens, aie the types of all the names sraeeet, have been examined Citations: MEXICO: iene Correll & ae 22751 (14); ee & Arguelles 17919 (Ld, Ld), 18064 (Ld (Ld). Durango: H. S eet te 6907 (Ak—22073, G, Mi, N); E. We mide 4725 (W—~332779) ; Edw. Palmer 1496 (ca—139735, N); F. W. Pennell 16109 (W—16)0972— type); Waterfall 15534 (St). Zacatecas: H. Se H. S. Gentry 8516 (G, N)- LIPPIA EKMANI Moldenke, Phytologia 2: 365--366. 19:7. Bibliography: Moldenke, Phytologia 2: 365--366 & 386. 19473 Moldenke, Lilloa 1h: 42. 198; Moldenke, Alph. List Cit. h: 1256. 1949; Moldenke, Known Geogr. Distrib. Verbena, Sts 2], 10h & 189. 199; Moldenke, Phytologia 3: 376 & 3 1950; E. J. Salisb., Ind. Kew. Suppl. 11: 138. 19533 Moldenke, Sa 93, 125, & tél. 1959. Perennial herb or shrub, 0.8—5 m. tall; stems apparently simple and erect, subtetragonal, stramineous, very sparsely and obscurely asperulous above, becoming smooth in age; se ncipal in- ternodes elongated, 7-8 oe em. long; leaves ternate; petioles in- conspicuous or obsolete, to 2 mm. long, subglabrous or pe — scattered hairs; leaf-blades Spat eat uniformly bright- on both surfaces, elliptic, 2.3--l.5 cm. ag 1.3--2. 3, cm. “ite, acute or submucronate at the apex, subentire or with a few very much appressed teeth above the middle, Seanded ; or subacute at the panna r pag racemiform, 15--20 cm. long, the l--6 straight erect sympodia 2—l, cm. long, tetragonal, rather densely resinous- Poy and puberulent; peduncles similar to the stems, cm. ong, tetragonal, asperous-puberulent and resinous-glandular} erous, gags aguanig ies: pun gr iar arene | the lower in wheels of aS the upper sessile or on slender pilosulous and resinous stalks 2--6 mm. long, the lower on stalks to 2 cm. ar the individual heads hemispheric, 1--1.5 cm. wide, eventually * bracts 3 at each node ode of the eats Fs ted by Erik Leonard Ekman (ms 1974) -- in whose honor it was named — at Posadas, Misiones, Argentina, in 1907 or 1908, amd 4# 1965 Moldenke, Monograph of Lippia 151 deposited in the herbarium of the Naturhistoriska Riksmuseum at im. The species has been found on shrubby campos, at an In all, 4 herbarium specimens, including the type, and moun- ted photographs have been examined me. tations: BRAZIL: Rio Grande do Sul: J ttrgens 454 (B); Rambo 3041 (Rb). ARGENTINA: Misiones: Ekman 197), (F=photo of type, N— isotype N--photo of type, S--type, Si Si-—-photo of type, Z—photo of type). LIPPIA ELEGANS Cham mes Linnaea 7: 225—226. 1832. Bibliography: ¢ resco T: 225-226, 1832; Bot., ed. 2, 2: sh. mT D. Dietr., Syn. Pl. 3: do. w 1835 “Talp., Repert. Bot. Syst. h: Sl. es Schau. in A. DC., Prodr. 11: 575. 1847; Schau, in Mart., Fl. Bras. 9: 223. 1851; Jacks. in sited f. & Jacks., Ind. Kew. 2: 95, 1894; Glaz., Bull. Soc. rsd ches ae > 540. 1911; frie ree a Soc. eee 46. 197; Moldenke, Alph. . 2: 36k, sel, "583; 581 (1948), 3: 669, 7 751, “oar “aber 900 (ioi9)", and’he abi, ah 1132, 1134, 1135, 116 ; 1204, 1208, 1293, 1296, & » Kn oe Ce 1949; Moldenke at mk "Distrib. Verbena » [ed. 5. 80 ‘ke 189. 1949; H. x. i af Ws Seeenes, Anal. Inst. Biol. Mex. es 1949; F. C. Hoehne, Ind. Bibl. & Mum. Pl. Col. Com. R 1951; Molde nets Résumé 93 & h6l. 1959; Renné, ° Herb. Inst. Levant + Minas 150, 1960; Moldenke, Phytologia 12: 7. 1965. Shrub or suall shrub, 1—-2 mn . tall; branches obsoletely tet- ragonal, decidedly virgate, strict, erect, subtament ose-pubescent; si bicolored, ovate-oblong, about cm. long and 1.5 cm. wide, about terno ne tra c Squally l-fid, the dorsal segment 2-lobed, the lobes obtuse and entire; stamens didynamous, inserted above the middle of the corol- rotund. ype of this species was collected by Friedrich Sellow some- Where in Brazil 1 (WE B Brasilia misit Sellow"], probably at Villa Rica, Minas Ger. erais, sited in the herbarium of the Botanisches Museum at ee hers now destroyed. 152 PHYTOLOGIA Vol. 12, no. 3 Se ete says "Genus in fructu deficiente recouneneen ro fructu vel germine grossificato habea corollae iste sumect oe transformatum, obovoideum sl SM i tubi immitata parte aca Se specie apiculatum, hirsutum; sectione instituta carnosum, c co vacuo pervium also notes "Habitu Dy iereene sans superioti ([L. sidoides] sin- ilis". Lippia origanoides H.B.K. is given as a synonym of L. el- egans by Schauer (1847 & 1851), but is actually a separate, “albe- ica. Lippia elegans has been found in stg’ ta sandy rocky soil, in woods and ind cerrado, on campos and waste land, on dry open river banks, and among limestone rocks, at watt iden’ of 1000 to st nb meters, flowering from September to June. cula corded for it is "camara braba", so has been meepEP oF and distributed in herbaria under the names L. mollis Mart., L. origanoides H.B.K., and L. aan Cham. On the other hand, the Ule s.n. [Herb. Rio de Jan. 31635], distributed as L. elegans, is actually L. rubiginosa Schau. The Glaziou 15335, cited below, is listed by Glaziou (1911) as L. salviaefolia Cham., as is also his no. 16286 (from Sao Paulo] — the latter ccobelly correctly identified. Schauer (1851) cites also J. E. Pohl s.n, (Correa et Formigas], L. Riedel s.n. [Villa Rica], : and Sellow s.n. [Villa Rica] from Minas ; Gerais, as well as J. E. Pohl s.n. [Serra do Caxorro] from Gofas. herbarium specimens have been examined by m Citations: BRAZIL: Bahia: Blanchet sen. [1827] (Cb); i. “Luts 131 (Lz); Schery 507 (N). Matto Grosso: Collector undesignated 32 (Br); F.C. Hoehne, Com, Rondon 1307 (5 de Ge Kuhlmann s.D- Aquidauana. Sept. 1914; Herb. Comm, Linhas T 1306] (Sp-- 31979). Minas Gerais: Bailey & Bailey 107) teat “Brade 13457 [Herb . Rio de Jan, 29531] (N), /1390h {Herb. Rio de Jan. 29530] Br, Br), sen. [Aug.—-April 160] fe “Br, Br, N, N); tDuarte 7848 (Her . Bradeanum 27899] (N); Gehrt & Hoehne s.n, {Turvo, A ©, April 12h, 1926] (N, Sp—-17h62); Glaziou 15335 (Br); F. C. Hoehne s.n. [Sante Barbara, Jan. 10, 1921] (N, N, Sp--k859); Jarneby s.n. [near Dura dinho, X.1927] (S); Mendes Magalhiies 1360 [Herb. THerb. Jard. Bot, Bello Horiz. 0016] (N, Si); Mosén 971 (N, S, S); Pizarro 118 (Se—- 68299); Regnell III.9h2 (Cp, Cp, Lu, N, S, S), Vy TEE ozo ( (S), ), gh2x (N, S, "8, :6)% Warming sen. [Lagoa Santa] (N); L. 0. Williams 50h7 (G, N), 5311 (G, N); Williams & Assis 581 (G), ~ 6026 (G, N)s 6418 (G, N), 6469 (G, N), 7559 (G, (G, N, 8). S#o Paulo: “Freire & Zeredo B.n. [Herb. Rio de Jan. 31563] (N); Later 24,8 y (Ne Sta undetermined: P. Claes 161 (V¥); Collector undesi gnated S.De at 2s Garsner 27h 27a (W—1LL0307); Raben 529 (Br); Teaberlisk Bole re Br 1965 Moldenke, Monograph of Lippia 153 LIPPIA ELLIPTICA Schau. — A. ig Prodr. 11: 593. 187. ae tae phy: ae in A. DC., Prodr. 11: 593. 1847; Schau. in Mart., Fl. Bras. 9: 250. issa;” Jacks. in Hook. f. & Jacks. Ind. Kor, 2: 95. 189); Hayek in Fedde, Repert. Nov. Sp. 2: 87. 1906; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 37 & 95 (1942) and fed. 2], 80& "189. 19193 Moldenke, Alph. List Cit. 3: 689 & 82h. 1949; Moldenke, Résumé 93 & 61. 19595 Renné, Levant. Herb. Inst. Agron. "Minas 150. 1960; Moldenke, Résum & suppl. atk. 196; Moldenke, Phytologia 12: 2h. 2 965.6 3 stems erect, several feet long, Lange culate and dense- Sf foliose sth de glandular-hirtellous; leaves decussate-opposite, rnate, in )! ss or approximate, spreading, reir ke eg ’ and nitidulous beneath, glandular-pubescent on the veins Bénkash, those on the stems larger, 7.5—9 cm. long and 2.5=—5 cm. pel those on the maa smaller, decreasing in size upwards and there oblong; inflore paniculate, spread- ing, foliose, knotty; peduncles filiform, eae glandulo se~ hirtello ous; heads globose, about the size of a cherry (Prunus a- vium) ; teattlets loosely inbaloate , thin-membranous, 5--7-veined, venose-reticulate, lilac, subrotund-ovate, Taran 1--1.2 cm. ng 8 mm. wide, acuminate at the apex, glandular—pubescent and also marked with scattered non-glandular hairs, ciliate along the margins, f somewhat accrescent; pedicels short; calyx bifid, hirsute, the lobes shortly bid entate; corolla Lilac, vil- =e on the outside above, the tube ampliate above, somewhat in- d, th . wide, venose, the upper lip short, rounded and enarginate at the apex, the lower lip with the middle lobe ded and obovate, the lateral Thee turned forward, ovate, acute sn the apex The of this rare species was collected by Johann Emanuel Pohl ess 160) between Alegeres and Trinidade, Gofas, Brazil, and was deposited in the herbarium of the sg erie Museum at Ber- lin, where it was photographed by Macbride as his type photograph species in his Sect ion Rhodoli » Subsection "BY, Hayek (1906) Says that his L. reticulata ion from this in having much me in previous publications from Amazonas, apparently in mente Only 1 herbarium specimen and ) mounted phototypes have examined by me, . Citations: BRAZIL: Gofas: JE. Pom 160 160 ae sane ae t~-photo of type, Kr—photo of type of type, W— CHa) Minas Gerais: A. P. Duarte’ 7350. Tuer: Bradeanum 37897] LIPPIA EUPATORIUM Schau. in A. DC., Prodr. 11: 592—593. 18h7. Bibliography: Schau. in A. DC., Prodr. 11: 592--593. 18k7; Schau, 15h PEYTOLOG@IA Vol. 12, no. 3 Bras. 9: 250. 1851; St. Lag., Ann. Soc. Des — 8 ek “4380; Jacks, in Ho ok. f. & Jacks Sey Ind, Kew. 2: 9 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 37 & - taake) and [ed. 2] "80 & 190. 1949; Moldenke, Alph. List Cit. 3: 695 (199) and ti: 1120 1949; Moldenke, Résumé 93 & 61. 1959; Molden- ke, Résumé - 8: h. 196k; M Moldenke, Phytologia 12: 2h. 1965. Small tall, hirtous- s along the margins, attemate at the base, penninerved, venose~ rugose, subcane scent beneath; inflorescence paniculate, indefin- itely branched, foliose, centripetal, the branches elongate; pe- duncles elongate, alternately in 2 or 3 pairs, remote, the upper a large cherry (Prunus avium), not much enlarged after anthesis; bractlets imbricate, subherbaceous-membranous, rose-colored, el- liptic, subacuminate or obtuse at the apex, glandular—pubescent, with longer eglandular hairs interspersed, ciliate along the mar- gins, finally somewhat accrescent and almost 1.2 cm. long and 0.8 « Wide; calyx very villous; corolla 6--8 mm. long (when dried), deskupetida: yellow in the throat, glandular-pubescent outside, the tube erect, somewhat ampliate. above, the limb small, the lobes short, rounded. This species is based on a collection made by Johann Enanuel Pohl along the road from Fazendina to Serra de Caretas, Gofas, one by George Gardner (no. 3408) from Piauf or Gofas in 18hh, and an ‘ oto=- winged ge Macbride in the Vienna herbarium as his type photograph no, The species is classified by Schauer (187) in his Section Rhodolippia, Subsection "B", Material has been misidentified and distributed in herbaria as Hyptis longipes St.-Hil. In all, 5 herbarium specimens, including oatee —— as well as li mounted photographs have been examined b Citations: BRAZIL: Gofas: Glaziou 21901 (W—-1123623) ; J. Ee Pohl s.n, [Fazendina ad Serra de Caretas; Macbride photos 34330] (Br—cotype, F--photo of cotype, It—photo of cotype, Kr——photo 2 3408 ( Br--cot; photo of cotype, "guighote. of coer: Minas Gerais: A. Saint- Hilaire c).538 (W—1706203). FELIPPEI Mol » Résumé Suppl. 10: 3, nom. mud. aid 20 1965; Moldenke, _ Piytologia 10: i7ie-172, June 18. li Bibliography: Mold sumé Suppl. 10: 3. 196k; Moldenke, Phytologia 10: iT192. m foray 1965 Moldenke, Monograph of Lippia 155 Small shrublet, with a woody < opodium; branches few, erect, simple or subsimple, about 60 cm. long, slender, stramineous, te) Glandalése-nivtellous ; Neatalates isin schaitaseeta » bright~ et above, lighter beneath, cordate-ovate, 1.7--3.2 cm. long, 1.4--2.7 cm. wide, acute at the apex arly and coarsely ser- ri glandular-pilose with erect gland-tipped hairs on both surfaces, soft to touch, the larger venation slightly Fg a ahove and prominulous beneath; inflorescence axillary in the uppermost leaf~axils, solitary, surpassing the subtending leaves, ascen- 3 peduncles very slender, 1--2 cm. long, densely glandular- hirtellious with soft A whitish hairs; heads globose, a~ bout 2 om. long and wide, apparently rosy-purple, showy; bractlets very mumerous, herbaceous, rdenbely imbricate, ovate, to 12 mm. long and 6 mm, wide, venose, acute or a at the a- ais, presi, on March 20, 1963, and is deposited in the United States agers Herbarium at Washington. The species is known thus far only from the type collection. In all, 2 herbarium specimens, including the type, have been examined ed by Citations: . BRAZIL: Felippe 140 (W--2h26087--type, Z—-isotype) . LIPPIA FERRUGINEA HB, Nov. & Gen. Sp. Pl. 2: 265. 1818. Synonymy: Lippia fe sorugines Humb. ex Spreng. in L., Syst. Veg, ed. 16, 2: 752, apie: Lippia ferruginea Humb. & Bonpl. ex Steud., Nom. Bot., ed. 2, 2: 5. 1640. Lippia ferruginea Hub. & ie ex D. Dietr., syn. Pl. 3: = ee 1843. Lippia ferruginea Kun 6x Schau. in e DC., Prodr. 11: 588. 1 hee aes =n A. D., Adansonia 3: 2), 1863; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 95. ae oe Phytologia 2: 22. 1941; Moldenke, Suppl. List Invalid Names 6, 1941; Moldenke, Known Geogr. Dist rib. ae * ed. u,3 3 8 5 & 95» 1923 Moldenke, Alph. List Invalid Nan 19425 - Mus. Hist. Nat. Jav. Prado 7: 22. 19435 “yol- denke, re List Cit. 1: 318 (1946) and 2: 361 & 573. nt He i. ea. L. Moldenke, Pl. Life 2: 88. 1918; Moldenke, Castanea 116. 156 PHYTOLOGIA Vol. 12, no. 3 1948; H. N. & Ae L. Moldenke, Anal. Inst, Biol. Mex. 20: 9. 1949; oldenke, Alph. List Cit. 3: 689 & 893 (1949) and 4: 1067 & 1113. 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 73 & 190. 1949; Moldenke, Phytologia h: 70 (1952) and 5: 2h. 19545 Yol- rapeht een Résumé 8h, 318, & 461. 1959; Melcenks, Résumé Suppl. (19645 and 11: h. 1 sé Moldenke, Phytologia 12: 2h. 1965. Ss small about 1.5 Me fragr. wi strong spall g which persists even after drying; branches very slender, tetragonal, slightly sulcat rown tely vap- 3 pressed-puberulent on the youngest parts, glabrescent in age i twigs short, appressed-puberulent; principal internodes 2--20 mn. long, very short — lateral twigs; nodes not plainly annulate; leaf-scars comparatively large, elongate-elevated and prominent, divergent; leaves decussate-opposite, small; petioles very slen- der, 1--3 mm. long, appressed=puberulent; leaf-blades thin- chartaceous, grayish-green above, light-green beneath, obovate or obovate-elliptic, Aale csen cm. long, 3—11 mn. wide, rounded at lent above ®. more densely appressed-puberulent ( minu ) andulose beneath; midri a ght as- ending ie mdaries, and the veinlet reticulation rather deeply impressed above and prominulous beneath; ary, about 5 mm. long, its ‘Lim 32 mm. in dianeter. The type of this species was collected by Friedrich Heinrich Alexander von Humboldt and Aimé Jacques Alexandre Bonpland at Huanacab , in the province of the same name, » Peru, at an ne gee of 1900 to 2000 meters, and is deposited in the Bon~ lected by August Weberbauer (no. 6066) at the same locality in April, 1912, and is deposited in the herbarium of the Chicago N4- ural History Museum The species has been — ae full Soc ae tah to the sun at al- Mt of 1900 to 2650 m flowering in April, May, August, ovember, and December. pha (1847) classifies it in his Sec~ sane Zapania, Subsection Axilliflorae. A vernacular name record- ed for it is "poleo del monte". Raimondi (1913) cites his nos. 3875 from Cajamarca, 1057 from Amazonas, and 2325 from Piura, a, Pe s inet previous publications I recorded it from Junin, apparent- 8 herbarium specimens and 8 mounted photographs, in- oitea: type or phototype ae igh of all the names involved, have been examined by m Citations: PERU: joes Ferreyra 7103 (W--2101305). Apurimac: 1965 Moldenke, Monograph of Lippia 157 Vargas 998) (N). Piura: Humboldt & Bonpland s.n, [Huanacabamba; Macbride ph photos 17503] (It-=photo of isotype, ~ Kr—phot o of iso- type, N—photo of isotype, N-~photo of iectype, W-~-photo of iso- rm pure. us Mart. & Schau. ex Schau. in A. DC., Prodr. 11: Synonymy: Lippia filifolia Mart. ex Moldenke, Résumé Suppl. 11: 7, in syn. 19 " Bibliography: Schau. in A. DC., Prodr. 11: 586, 18475 Schau. in Mart., Fl. Bras. 9: 239. 1851; Jacks. in Hook, f, & Jacks., Ind. di List C a en 712, & 7 sinh 9), and i "1204. 1949; Moldenke, Known Ge~- enac., [ed. 2], 80 a 190. 19h9; Moldenke, Résu- me 93 & hél. "1959; Moldenke, Résumé Suppl. 11: 7. 196k; Moidenke, Phytologia 12: 2h, mee Bush, subshrub, o shrub, 0.5—1 m. tall, aromatic; stems woody, very etee grr De sulcate, few-b ranched above, gla- bk or r glandular—roughen ed; leaves approximate, ternate, or rticillate, sessile, subtetragonal-filiform, in clined, bright- green, 1.2—-2.5 om. lo ong, 1 mm. or less wide, obtuse at the apex, attenuate at the base, wider and flat on the. sides, punctate, glandular-roughened or glabrous » deeply sulcate and very narrowly ewed from ab ughened or glabrous; heads hemispheric, about 6 mm. ract= lets imbricate, subherbace S, ovate-subrotund, obtusely subacum- inat pubescent on both surf as the > ac .) corolla-tube, the outer ones larger and involucrate; calyx hirsute, acutely bifid; corolla yellow or orange, puberulent on the out- de, also pulve erulent~glandulose above, the tube about 6 mm, long, subventricoge Po the middle, the limb undulate, velutinous, the throat pubescent, This most adetince species is based on two collections made by Carl Friedrich Philipp von Martius on desert campos in Minas Ger- ais, Brazil — at Serra as S&%o Antonio and between Lavras de Pin- and S%o Domini Ri hi mr ge _Stasteamiung at Munich, where it was photo Macbride as his type photograph no, 2032h. Schauer (1847) classifies the species in his Section Zapania, Subsection Axillifiorae. It has been found also in open gravelly Pastures, at ay meters altitude, flowering from May to July and in Sag and is said by M exia to be "frequent" along the rail- Pe, Ax the. Diamantina district. » 15 herbarium specimens and 6 mounted photographs, in- sain a Phototype material of cocoate names involved, have been exam- 158 PHYTOLOGIA Vol. 12, no. 3 Citations: BRAZIL: Minas Gerais: Brade 13902 [Herb. Rio de Jan. 29528] (B); P. Clausen 635 (S), son. (Cb, Cb, Cb, N); Glaziou 19535 (Cp, N—photo, S--photo, , Z——photo); | Martius s.n. [Macbride phot photos 2032] (Kr--photo of cotype, N—photo of cotype, W--photo of co- type); Mexia 5889 (Au--12116), Ca—509370, Gg—286581, Go, I, Mi, N, S, W—1705025). nate FLAVIDA Urb., Symb. Antill. 7: 353. l Bibliography: Urb., Symb. Antill. 7: 353 11912) and 8: 59h-- 595. ye ioe Ind. vet a 153. 1921; Moldenke, Known Geogr. str » [ed ho; Moldenke, Alph. List Cit. 1: “88. (96) and hie *ibs8 & 1259. 19493 ee Known Gee- ger. Distrib. Verbe wa ae [ed. a], 48 & 190. 19h9; He Ne & Ao Le oldenke, Anal. Inst. Biol. Mex. 20: 9. 1949; Moldenke, Résumé Shrub, 0.6--1.3 m. tall; young branches obtusely angular, densely short~pi losulous with simple white hairs, the older ones more or less plainly tetragonal and glabrous; principal internodes tic, 1.5--2 cm. long, 0.8--1.2 cm, wide, obtuse or subobtuse at the apex, coarsely crenate along the complete margins, obtuse at the base and slightly prolonged into the petiole, scabrous above, very densely short-pilose beneath; midrib and the 6 or 7 secon- daries on each side impressed and. reticulate-joined above, with lobe pe cular, the anterior. short-ovate; lower stamens serted at the middle of the corolla-tube, the sro higher; fila~ ments very short; ac about 1 mm. long; stigma oblique, almost pers 2 a eral; ovary suborbicular, wider and dg tie at the top, The type of this species was collected by Wilhelm Buch (no- 394) in dry places near Ennery, at 1,00 ogpeck altitude, Haiti, Hispaniola, in June, and is deposited in the herbarium cB sches Museum at Berlin, Orban agia). “says of it "Habitu similis A. geminatae H.B.K. var. mic Griseb., quae fol- iis apice acutis, supra ra laevibus aeera majoribus, bracteis triangularibus v. semiorbicular ibus apice zat v. acuminatis, 1965 Moldenke, Monograph of Lippia 159 corollis violaceis, calyce ovarium obtegente inferne tubulos superne bilobato omnino discrepat." The abbreviation, "A.", so the generic name in this quotation is obviously an error for "L." ([=Lippia]. chee description of 5 calyx of L. flavida being kman fe) ve nacular name "San José", Urban cites also Buch 130. The flowers are said to have been white on Buch 1304 and yellowish on Buch . cee In all, 8 herbarium specimens, ice. material of the type collection, have been e Cit + a M srtgreis Dominican Republic: Ekman H.11927 (S). ti: W h 394 (N--isotype); Ekman H.1817 (Mi, S), H.2k71 (S), oe (fan ee S); E. C. Leonard 8901 (N). LIPPIA FLORIDA C ments Linnaea 7: 221--222. 1832. e, Linna Bot., ed. 2, 2: ‘she I 1840; D. aes, ee Pl. 3: br. Tals Bot. Syst. ’ Prodr. ll: 592, 187; Schau. + 93 249. 1851; Hiern, Vidensk. Meddel. ig See 187721878: "BO, 1877; Jacks. in Hook. f, & J Jacks., Ind. Kew. 2: 95. 189); Glaz., Bull. sto al tr Verbenac., [ed a - 1942: - List Cit. 1s 106 (19465, 3: 689, 7a ; 921 (1 1319), and gory ge bs 2075, 1135, 1 1203, & 9; Moldenke, : (ed. 2], 80 & 190. 1993 Mendes win el ” anais V. Reun. Soc. Bot. spain 268— 269. 1956; Moldenke, *nésumé 93 & 461. 1959; . Herd. Inst. Agron. Minas 150. 1960; Moldenke, Ste age 4 2h. 1965. Shrub, 10--80 cm - tall, subtomentose-pubescent with glanduli- ferous hairs or subglandular-tomentose; branches elongate, ae “0 long or longer, erect, simple, strict, ales aa Secondaries 6-10 per side; pedeieae llary or Speeutcan tary, much surpassing the leaves, 1~—8 in the upper - axils, congested at the tips of the branches and surrounding the 160 Peay 7 0 LOO T's Vol. 12, no. 3 terminal bud, l--5 cm. long; tei conspicuous, hemispheric, about 5 cm. wide, nutant, abundant, pilose-pubescent with hairs less abundant than on the leaves; “aad flowerless sits to 1 cm. long; bractlets petaloid, rose~lilac or rose, t embranous, broadly ovate, very large, about 2.5 cm. lo ong, ni Sell . wide 7-ve gins, the outer ones smaller, firmer, more densely pilose, and very obtuse at the apex, the inner ones acute or subacuminate at the apex; calyx small and short, villous or very villous, bifid, obtusely l-dentate; corolla hypocrateriform, "rose or rose-lilac" filaments twice as long as the anthers; anthers ovate; style and fruit typical of the gem The type of this very Hittin ct species was collected by Fried- rich Sellow (no. B.1439 c.495) at Serro do Ouro, Minas Gerais, Brazil — the original publication states nerely "E Brasilia nie sit Sellow" -- and was deposited in the herbarium of the Botan- —_ Museum at Berlin, where it was photographed by Macbride as ype photograph no. "17506, but is now destroyed. Chamisso (8324 cape the bractlets as ‘imprimis autem in exteriori, qua nervos venasque densiores opacant" and "Capitulis floridis cum aitiguanentt specie [L. lupulina Chan.] caeterum diversissima conveniens." rie aa Ein classifies it in his Section Rhodo- lippia, Subsection ~~ The flowers are ple as "yellow" on Williams & teste 7160 and as "yellow and violet" on Mendes Magalhles aoe seems probable to me that the corolla is actually y yell * eho pr conspicuous subtending bractlets are rose, mobic ens eel or vio- The species has been found on ong and yo campos, at an alti- tude of 1300 meters, flowering and fruiting in April and fram June to October. Mater ial has been rere nF and distributed herbaria as L. bracteata Carr. Clausen's surname is misspelled "Glaussen" in the Vienna herbarium. M. B. Foster 713 is 4 mixture with L. lupulina Cham. Schauer [1851] cites Tis a fs Clausen 390 and Schott s 8.n. from Minas Gerais and two collections of L- Riedel from "in c campis arenosis inter Rio S. Francisco er Curvel- los" and "in campis siccis prope Caitete" in Bahia, deposited in the Berlin, Leningrad, Martius (Munich), and Vienna herbaria. Glaziou (1911) cites his no. 15337 fron Minas Gerais and says of it irae fl. vine . juin-juill et. R rbarium In all, = he pecimens and hototypes have ‘caunte a or eke an, uacded peur Citations: BRAZIL: Minas G : P. Clausen 221 (V), 390, nat 609 [10] (8), son. peer “oy Gre P. Duarte 305k 1965 Moldenke, Monograph of Lippia 161 (Herb. Jard. Bot. Rio Jan. 71926] (N); Dupré s.n. [Brésil méridi-~ onal, 182] (N); Heringer 5290 (Z); Mello Barreto 3275 {Herb. Jard. Bot. Bello Horiz. 612] (N); Mendes Magalh%es 301) (Be-— 13913, W--2124262); M. B. Foster 713, in part (W--1779145); Sellow B-1439 ¢.h95 [Macbride photos 17506] (Kr—photo of type, N—photo of type, W--photo of type); Williams & Assis 7160 (G, N). LIPPIA FORMOSA T. S. Brandeg., Proc. Calif. Acad. Sci., ser. a; 3: 163. 1891. o 1), 14 & 95. 19 M. Martinez, Bol. Soc. Bot. Mex. 2: 2& 13. 1945; Moldenke, List Cit. 1: 233 (19h6), 2: L7h, 489, & 595 (19h8), 3: 933 and h: 1121, 1181, 12h], 1242, & lai. 1949; H. Ne & A. Le » Anal. Inst. Biol. Mex. 20: 9. 1949; Moldenke, Known Distrib. Verbenac., [ed. 2], 31 & 190. 1949; Calpouzos, Econan. Bot. 8: 228. 1954; B. H. Johnson, Wasman Journ. Biol. & a GT tara Résumé 37 & 461. 1959; Moldenke, Phytologia : 4&7. 1965. Rather slender erect shrub, 2--l m. tall, scabrous-pubescent; bark smooth, white or mottled-gray; stems many from the ground, slender, erect; leaves decussate-opposite, obovate-cuneate, 2-- cis cm. long, coarsely crenate-dentate, prominently pinnate- Alph. (1949), Molden- Geo Pubescent with white hairs; corolla light rose-colored or pale- ink to » or "from pale to deep pink with yellow ring at mouth of throat", 6 mm. long, the tube twice as long as the calyx, ampliate above; fruit 2--3 mn. long, the cocci easily separable. T ec owns ran- Cap Baj California, Mexico, in or before 1891, and is probably deposited in the herbarium of the California Academy of Sciences at San Francisco, The plant has been found along the edge of sandy arroyos broad open washes or sandy valley lands near washes, at the base tober and from December to March, and fruiting in February and March. It should be noted that the original description says "2--l mm. high", which I assume to be a typographic error for "2—-l; m." The Whitehead 862 collection cited below bears a notation veame as number 663", but I have as yet not seen the latter col- lection. The Gradendtrfer s.n. distributed as L. formosa is actu- 162 Patt oloez a Vol. 12, no. 3 Lantana camara var. mista (L. ) L. H. Bailey. Lippia formosa formosa is one of the many plants called "oregano", "oregeno®, “Worigano*, or ser tle Others are L. affinis Scha,, L. fragrans Turez., L. graveolens H.B.K., L. micromera era Schau., L. micromera var. helleri tevitien) "woldenke, L. origanoides B.beke, L. palmeri S. Wats., L. palmeri var. spicata Rose, Lanta- na involucrata L., L. achyranthifolia Desf., L. L. trifolia L., Coleus amboinicus Lour., Hedeama floribunda Standl. » H. patens Jones, Hyptis albida Hyptis albida H.B.K., He K., He americana (Aubl.) Urb., H. capi: H. capi- tata Jacq., He ” suaveolens (Le ye Poit., Limnophila stolonifera (Blanco) Merr., es austromontana Epl., Ocimum basilicum L., Origanum vulgare L., Le, O. m majorana L., Poliomintha lomgiflora Gray, and Salv Salvia sp. Martinez states that (195) Lippia formosa is the Norégano" | of the Todos Santos region, while that of "Miraflores is L. palmeri and that of Comondii and Mulegé is Aloysia barbata (T. 8. Brandeg.) Moldenke. In all, 34 herbarium specimens, including numerous topotypes, and 2 mounted photographs have been examined me. Citations: MEXICO: Baja California: T. S. Brandegee s.n. [To- dos Santos, 1890] (G—-isotype), s.n. [Fescadero, Nov. 1892] (G), s.n. [Pescadero, Oct. 23, 1893] (Ca--1091)), s.n. [Pescadero, Sept. 20, 1893] (A, G), s.n. [Pescadero, Nov. 1902] (Vt); Carter, exander,, & Kellogg 2291 (Ga-—760098, Du--3)9169, G); H. S Gentry Wi (Du--261276, | re, Ge); Bemseriy 256 256 (Gee-294851)5 Me Egos ones 24112 (Ca—l00367, Fs, La, Mi, N, N, N—photo, 1352, Z--photo), 2357 We Fs, Be iesoks, $d--25):70) ‘peril 6979 (G); Nelson & Goldman 7334 (G); Shreve 7215 (Du--21)2676, Fs, G, Mi); J. Whitehead d 862 (Du--2485)6, La); Wiggins 14536 (Au 190209, GJ. = LIPPIA FRAGRANS Turez., Bull. Soc. Nat. Mosc. te at Le oo a Synonymy : Buddleja ‘peleooans Ro A es fi: 111. 1895. Lippia geisseana (R. A. Phil. 7 se ages sat Herb. Boiss., sér. 1, 6: 623--629. 1898. Buddleia geisseana Re A. Phil. ex Soler ered., Bull. Herb. Boiss., sér. 1, 6: 62h, in sya 1898. Lippia geisseana Solered. ex Thiselt .-Dyer, Ind. Kew. Suppl. 2: 106. 190h. Buddleyia geisseana Phil. ex Moldenke, Ré- sumé Suppl. 11: 6, in ie 196). Bibliography: Turez., Bull. Soc. Nat. Mosc. - (2): 203. 18635 acks. in Hook, 7% & Jacks., Ind. Kew. 2: 95. 1893 R. A- Phil-, - 91: 111. 1895; Solered., e ge Bas Boiss. 2, Names 16. "190; Mold enke, Known Geogr. [ 1], h2 & 95. 1942; Moldenke, Phtolncle. “si icf site j Moldenke, 1965 Moldenke, Monograph of Lippia 163 Castanea 10: he. rong Moldenke, Alph. List asians l: tay 1946; Moldenke, Alph. List Invalid Names 1h. 1947; « Le Molden- k : 60. 9h ac., [ed. 2], 101 & 190, 1919; Calpouzos, Econom, Bot. 8: Berd: 195k; Mufioz Pizarro, Sin. Fl. Chil. 199, 19595 Moldenke, 121 & 461. 1959; Moldenke, Résumé Suppl. : 7, 16, & 19° (ips) and 11: 6. 1964; Moldenke, Phytologia 12: pn 1965. Glabrous shrub, t t la o 60 om. tall, pers s terete, intricately branched, often leafless below; bran e 6—-8-suleate, bout 1.5 mm. in diameter, completely glabrous or glandul, erulent within the sulcations, the lateral fl e late to oblong-obovate, 1.2--1.8 cm. long, about mm, wide, ob- tuse at the apex, entire, lo ip Set ee fo at the base, subcoriace- ous, sticky with microscopic glands, hirtous above with simple a » ternate, ts) congested above, centripetal, subglobose, as long as or longer than the peduncies, 5--7 m. "wide; bractlets concave or more or less ah oblong or broadly ovate, almost or completely hiding the flowers, imbricate, sticky, densely long-pilose on the outer surface Nay on the margins, the lower ones obovate, 3—l; mm. long, 2-—3 mm. wide; flowers small, s essile, solitary in the axils of the bractlets; calyx 2-fid, aes sepals 2, oblong, 3.5--l mm. long, about 1 mm. wide , truncate and more or less bidentate at the apex, densely villous on the outer surface and on the margins, with Pp’ uliferous at the apex and hardly exserted; style Short, included, about 1 mn. long, incrassate at the apex; stigma declined, lateral; ovary small, surpassing the style in length by 1.5 m., entire, 1-carpellary, 2-celled, with one owle in @ach cell; fruit unt know The e type of this curious species was collected by J. 2 oe as 1347) in Coquimbo, Chile. It is most probable that —uming 1347, cited below, is part of the sane oe perp on type of Buddleja geisseana was collected by Wilhelm Geisse (no. 49) — whose honor it was named — near Bandurrias, Atacama, , Chile, be-~ 164 PHYTOLOGIA Vol, 12, no. 3 tween — and 1887. Turczaninow (1863) says of the species: “Pp r plures annos exsiccata odorem suavem spargit, species Pete cela ramis intricatis, i anntly hee saepe aphyllis, Lippia trifidam Gay in mentem re common name "orégano" is ebcadied for this species. In fact, this is only one of many plants called "oregano", "oregeno', to origano", or “oragano", Others are L. affinis Schau., L. formosa T. S. Brandeg., L. graveolens H.B.K., | L. . micranera SoM; ‘Tes Le. micromera var. helleri (Britton) Moldenke, L. origanoides H.BeKe, L. palmeri S. Wats., L L. palmeri var. apdinta: | Rose, Lantana invol- H. Pa toa tis: pater: ttcstiensncedt neste! bh tis albida H.B.K., H. ‘sy He americana (Aubl.) Urb., | H. capitata Jacqe, H. H. suaveolens (L.) Poit. » Limnophila stolonifera (Blanco) Merr., Monarda austromontana Soles , Ocimum basilicum L., Origanum vulgare Le, Ley Ou majorana L., Polianintha longiflora Gray, and Salvia sp. solareday (1898 ") says "Die Bezeichnung 'Or ‘Oregano’ und die vei te Bezeichnung 'Oreganillo' wird in Chile und aha n auch fttr einige andere starkriechende Pflanzen gebraucht, o fur Origanum majorana L........und Oreganillo ftr Lippia peer Steud......und die Labiate G ardoquia Gillesii Gay.....Der Geruch der Lippia Geisseana ist ein ganz charakteristischer und nicht gerade ang ang enehmer; e: er erinnert mich an den Geruch der harzigen Larrea mexicana Moric. (Zygophyll.) der Yuma- und Arizona-Wuste und domo anderer nord-amerikanischer Wisten-pflanzen." He © says: "Vidi specimina a cl. Geisse lecta in Herb. Berolin- ensi, 'Herb. of the California Academy of sc.' ('Chaflarcillo') et tHerb. Columbian College New-York' ('W. Geisse no. 9, 1885-- 87, Desert of Atacama')." In all, 3 herbarium specimens and 3 mounted photographs, in- cluding type or Heap material of all the names involved, a been examined by m ons: CHILE: Atacama: Geisse 9 (C); Re A A. Philippi 1265 A398) Coquimbo: Bridges 1347 (Z—photo of type, z—photo isotype, Z—photo of isotype); Cum: Cuming 1347 (N--isotype) - PIA FRANCENSIS Moldenke, Phytologia 2: 20. loll. Bibliography: erreea malar res 2: 20. 1941; Moldenke, Known Geogr. Distrib. » led. 1], 37 & 95. 19h2; Moldenke, _— List Cit. 1: 231 Gig), and 2: wo 1948; Moldenke, Distrib, Ver 1], slender, 2--5 mm. long, densely white-hirsute; leaf-blades iow’ aceous, " elliptic-subrotund, somewhat lighter beneath, 1.2-h 1965 Moldenke, Monograph of Lippia 165 long, 12.8 cm. wide, rounded at the apex and base, regularly serrate from the apex to the base with rounded revolute-margined llary tate; peduncles very slender, 0.8—3.5 cm, long, very dens hirsutulous with stiff white gland-tipped hairs; heads hemispher~ ic, about 2 cm. in diameter, many-flowered; bracts large, folia- t ceous, vate, to about 10 m, lo: 8 mm, wide, blunt at the apex, densely pubescent with short silky mostly gland-tipped » Ciliate-margined; corolla ye llow and rose", r "ve The type of this handsome species was collected by Guilherme Gehrt [Herb. Inst. Bot. S8o Paulo 4037] in fields at Franca, S&o Paulo, Brazil, on April li, 1920, and is deposited in the Britton Herbarium at the New York Botanical Garden. It was or. distributed as L. lupulina Cham., to which the species is obvi- ously closely related. Thus far it is know only from the orig- inal collection, of which two herbarium specimens have been ex- amined me. Citations: BRAZIL: So Paulo: G. Gehrt s.n. (Franca, April 11, 1920] (N—type, Sp--l037--isotype). LIPPIA GARDNERIANA Schau. in A. DC., Prodr. 11: 592. 18h7. Bibliography: Schau. in A. DC., Prodr. 11: 592, 187; Schan. in » Fl. Bras. 9: 28--29, 1851; Jacks. in Hook. f. & Jacks., Ind. Kew. 2: 95, 189); Moldenke, Known Geogr. Distrib. 1 222, 1946; H. N. & As L. Moldenke, Pl. Life 2: 60. 1948; Moldenke, Alph. List Cit. 3: 693 (19h9) and h: 1296. 1949; Moldenke, Known Geogr. Distrib, Verbenac., [ed. 2], 80 & 190. 1949; Moldenke, Ré- sumé 93 & 61, 1959; Moldenke, Phytologia 12: 2). 1965. Shrubby, pilose-hirsute and more shortly {prrenys aeeendg beneath and especially pilose; peduncles axillary and t 2 Single, subequaling the subtending leaves, subpaniculate at the andu faces very hirsute Slightly less than 2 mm. long, short-dentate; corolla tube about 8 mm. long, curvate, 166 PHYTOLOGIA Vol. 12, no. 3 ventricose below the middle, pubescent above. The type of this rare species was collected by George Gardner (no. 3407) —- in whose honor it was named ~~ in Piauf or Gofas, Brazil, in 18h), and is deposited in the i pani i he Bean heir} et Jardin Botaniques at Geneva, wher ographed by Macbride as his type A Recntis no. aL6s2. "sehen rey Be places the species in his Sebsection "B" of Section Rhodo~ lippia and notes "Proxima habituque simillima L. lupulinae, “ged foliis bracteisque aliena." It us known thus far only only from the original collection. The A. Lutz 1096, distributed as L. gardner- iana, is actually L. diamantinensis Glaz., while J. E. Pohl sn. 8.n. (Herb. Caes. Vindob, 18h7 comm.] is L. hederaefolia Mart. & & ‘Schau . In all, 2 herbarium specimens and Pe: ae photographs of the xamined Citations: BRAZIL: Gofas or atte lies Ge ~ Gardner 3407 {Macbride photos cea Spratt F—photo of isotype, It--photo of type, o of type, N--isotype, N--photo of type, N--photo of rene a pats of isotype, W-~photo of type, Z——photo of isotype LIPPIA GEHRTII Moldenke, Phytologia 1: ps Sh. Bibliography: Moldenke, Phytologia 1 Pe tes and 1: 50. 1941; Moldenke, Known Geogr. Dist: trib. ver erbenac., [ed. 1], 37 . 95. = Moldenke, Alph. List Cit. 1: oe & 290. 19h6; ‘Re de oye cw, suppl. 10: Aare 19473 He N. & Ae Le uoldenke, - 1948; Moldenke, Alph. List Cit. 2: ae 36 (agus), 3: ‘she (2919), and lj: “top 1995 Moldenke, Known Geogr. > » [ed. 2], 80 & 190. 1919; mide "abs 93 s 3 branches and branchlets slender, arg tetragonat, densely hispid with harsh white divergent hairs sae mm, longj nodes hidden, but apparently more or less annulate; principal i ternodes 0,.9—3 cm. long; leaves decussate-opposite; petioles slender, 3-7 mm. long, canaliculate above, more r J ss wing- marge a with rounded or subacute, broad, short teeth, truncate or cate at the base (varying to actminate and prolonged into the pet iole on young twigs), abundantly hirsute on both surfaces with long white hair similar to that on the branchlets; midrib slender impressed above, prominent beneath; secondaries slender, | pe 1965 Moldenke, Monograph of Lippia 167 long and 5—10 mm. wide (or larger), short-stipitate, densely hispid on both surfaces; axillary heads also often present in the uppermost leaf-axils beneath the panicle; inflorescence-branches denseiy white-hispid like the branchlets 3 heads oblong, 1.2--1.7 cm. long, 1—-1.5 cm. wide, densely many-flowered; bractlets large conspicuous throughout the head, ovate, sessile, very thin- e C) th textured, 5--7 mm. long, t + Wide at ase, sharply a cute at the apex, softly pubescent with multicellular sh Somewhat bulbous~based which are dense on the more scattered on the body of the bractlets 3 corolla hypocrateri- form, rose, about 8 mm. long, its limb a mm. of this species was collected by Guilherme Gehrt [Herb. Inst. Bot. S. Pa n whose honor it is named -- at Itarapina, S&o Paulo, Brazil, on April 29, 1923, and is de- posited in the Britton Herbarium at the New York Botanical Garden. aria In all, eae oe including the type, and 2 mounted photographs have been examined b ° ations: BRAZIL: So Paulo: G. A. Black 51-10973 (Ca--91512); G- Gehrt s.n, [Itirapina, April 29, 1923] (N-type, 313-- isotype); Regnell 111.946 [7/1855] (N, S, W~—201215), I11.9h6x [Batalaes, Feb. 1849] (S), ITI.9h6x [inter Canna Verde & S80 Jolo de Jeguaty, May 5--8, 188] (N, S), II1.9l6xa (S); Viegas & Nor manha s.n. (Herb. Inst. Agron, Est. 8. Paulo 5553] (N—photo, Sp— L3h82, Z-~photo) . LIPPIA GENTRYI Standl., Field Mus. Publ. Bot. 17: beige tae Bibliography: P. C. Standl., Field Mus. Publ. Bot. 17: 207-- 208. 1937; H. Ss. Gentry, Carnegie Inst. Wash. Publ. 527: aos (ee3, 281, & 306, 1942; Moldenke, Known Geogr. Distrib. Verbenac., [ed. wetane & 95. 1942; Moldenke, Alph. List Cit. 1: 232 & 233. 19h; fe) e * — branched rather spindly shrub, 1--5 m. tall, sometimes bushy and densely branched; branches terete, ochraceous, very cidedly rugose, ve 11 above and rough to ry densely glandular-villous the touch, gray beneath and very densely pilose with eglandular hairs or sometimes with a few glanduliferous ones interspersed:, 168 PRY TOLO@IS Vol. 12, no. 3 midrib slender, prominent beneath; secondaries about 10 per side, rather straight~ascending or s subarcu ate, impressed above; veinlet reticulation impressed above; infloresc cence e spicate, the spikes if an i phi eiitel bracts e-green, ph accrescent after anthesis, broadly ovate or rounded-ovate e, acute or acuminate at the apex, very numerous, closely imbricate, densely pilose with ous rs or mea white with tinge of green", 6—-7 mm. long, the tube very slender, glabrous outside, the lobes very short, densely white- The type of this species was collected by Howard Scott Gentry (no. 3039) on a tolerant oak slope at Agrimincor, on the he Rio Mayo, Sonora, Mexico, on April 6, 1937, and is ig Sa in the herbar- ium of the Chicago Natural " History Mus Standley (1937) states that the spactee is related to Aloysia barbata (f. S. Brandeg.) Moldenke, but this relationship is cer- tainly not close nor obvious! Gentry (1942) states that it grows a t part in a bates it is sone —. othe as individu owers 3 spr. ummer, This shrub Sacnok salamsle-ailted. t0 cae odie seer as suggested by Standley, nor is it to be confused with the tree Lippia Pringlel Briq. Though the flower structure of the latter and of L. centryi may be very similar, no close observer would take then for the same species in the field. It has recently been collected in northern Sinaloa, Puerto a Tamiapa." He also refers to it as 4 characteristic pagel oe the Oak Forest in canyons and on mountain slopes, but on the label of his no. 1385 he says "I found only @ few plants in one locality." Collectors have pekemagee the species on oak mesas, shady rocky slopes, steep s clay slopes in oak forests, md near the base of cliffs Shy talus slopes among pines a says "common in fairly dry locations" in Durango. collected in anthesis and fruit in March and April, at areata ef {oe to 6210 feet. Material has been misidentified ae pepe: aas 1. pringlei Briq. and L. umbellata Ms the other hand, the H. S. Gentry 2037, distributed as L. gem tryi, is actually L. pringlei. In all herbarium specimens, including material of hate as collection, and 2 mounted photo ig im have been examined b Citations: MEXICO: Durango: J. Gonzalez Ortega 5373 (Me, - 120750); R. P. Hurd 67 (Mi). Biavecr i 5. Gentry 5034 (Ak 1965 Moldenke, Monograph of Lippia 169 20456, Du--274085, Fs, G, Me, Me, Mi, N); I. L. Wiggins 13187 (Du—-10629) . Sonora: H. S. Gentry 1385 (A, Ca--6)63h5, Fs, Ge, I, Me, Me, Mi, Ss), 3039 (, (A—isotype, “Ca—58207h—isotype, Fs~- isotype, Ge—ieotype, Me--isotype, Me--isotype, N--photo of iso- type, S--isotype, Z—photo of isotype). LIPPIA GIANDULOSA Schau. in A. DC., Prodr. 11: 577. 18h7. ynonymy: Lippia microphylla var. alpestris Beyrich ex Molden- ke, Alph. List Invalid Names Suppl. 1: 14, in syn. 1947. Lippia efendiede Schau. ex Renné, Levant. Herb. Inst. Agron. Minas 1 0, sphalm, 1960, Bibliography: Schau. in A. DC., Prodr. 11: 577. ony Schau. in Mart. » Fl. Bras. 9: 227 & 228, "1c; poe in H - f. & —* Ind, Kew. 2: 95. 189i; Mol denke » Known GNal ey [ed. a; 0 e Volde nke, Alph. List Cit. 3: 689, he & 824 (1949) and h: 1132, 1167, & 75 e 199; rai Résumé 93, 31h, & 61. 1959; Renné, Levant, Herb. Inst. . Minas 1 160. i860; Moldenke, Résumé . elt et (1962) ~ “is kh. 1963 Moldenke, Phytologia 12; 23 branchlets slende er, slaaea floriferous; leaves small, ussate- et yg: petiolate, bec ecoming smaller ein leaf-' ‘ines. ihe ) cent~ tomentose beneath, more hirtellous on "the Lime meren Seapeedl th shiny resinous glands, the immature ones throughout, the mature ones pubescent only on the venation, the Z g F ea e @ vac) o rous; secondaries 8 per side; peduncles binary or ternate, sur- pes the subtending leaves, filiform, about 8 mm. long, * pubes- of the cor Olla~tube, bifid, villous-pubescent; corolla hypocrater- iform, white or whitish to white-cream or yellowish, not blacken- ng, its tube about 3 mm. long, straight, *ventricose at the middle, glandular-pubescent on the outside above the e middle, Limb small, conca m, aggrapaaalap pan gl Poe on the back, the rile rounded, retuse at the apex, crisped; stamens didynamous eo style filiform, smooth, almost equaling the corolla stigna unilateral at the e apex. 170 PHYTOLOGIA Vol. 12, no. 3 This species was based by Schauer on J. E. Pohl 132 and s.n, from Rio Paracatu and on L. Riedel s.n. from Morro de S%o Jeroni- mo in the Serra de Chapada, Minas Gerais, Brazil, deposited in the Vienna r a. A duplicate cotype of the first-mentioned collections in the herbarium o. otanisches Museum “a was photographed by Macbride as his type phot: h no. 1 the dela in his Section Goniostachyum and says of it "Habitu L. microphyllae Chan. similis sed foliorum figura et indumento, glandulis copiosis resinosis bracteis et corolla flavescente nec siccamo nigrescente, distincta." The type of L. microphylla var. alpestris was collected by Heinrich Carl Beyrich in the Serra dos Orgaos, Rio de Janeiro, Brazil, in 1833, and is deposited in the herbarium of the Natur historiska Riksmuseum at Stockholm pe eneciee.in has been found in woods, campos, and on mangan~ outcrops, at 1100 to oo eeeess sitiinde, flowering from Conaber to Fe and in April and May. Material has been mis- identified and distributed hs severe as L. microphylla Cham., L. origanoides H.B.K., L. sidoides Cham., and Hyptis sp. It is closely related to the three species just stg gages as well as to L. graveolens H.B.K., L. affinis Schau., L. dumetorum Herzog, L. elegans Cham., L. gracilis Schau Schau., L. grata =e L. hick hick- enii Tronc Troncoso, L. laxibracteata Herzog, | L. martiana Schau., Le mattogrossensis i Moldenke, L. obscura Briq., L. pendula Rusby, Le rubiginosa Schau., L. salviaefolia C Cham., L. schomburgkiana Schauy L. stachyoides Cham., and L. velutina Sc hau. -- a group of very closely related species which is badly in need of more intensive study. The Lofgren Son. [Picos dos Marins, Jan. 10, 1897], Herb. Come Geogr. & Geol. S. Paulo 3195, and Herb. Inst. Bot. S. Paulo aulo 15659, distributed as L. glandulosa, are L. iodophylla Schaue In all, 23 herbarium specimens of L. glandulosa, including type material of ali the names involved, and Ti mounted photographs Citations: peazi Minas Gerais: Assis 8 (G, N)3 A. Pe Duarte 7849 [Herb. Bradeanum 27898] (N); A. Lutz 1690 (Z); Mello Barreto Barreto 3209 (Herb. Jard. Bot. Belo Horiz. 2048] (N)j Mendes “Vagalntes— 1361 (Be—1,569) , 1699 (Be—14559), 172 (Herb. Jard. Bot. “Bot. Belo Horiz. 1453] ("), 1887 (Be~1h706), 171: 17193 (S); Murga Pires & Black 2838 (W—2221531); J. E. Pohl 132 [Macbride @ photos os 17508] Glasteu 14155 (N); Santos iia 32) (Herb. Jard. Bot. Rio Jan 72] (N)j Santos Lima & Brade 14290 [Herb. Jard. Bot. Rio Jan. 1965 Moldenke, Monograph of Lippia 171 29539] (B). State undetermined: Mello Barreto 2049 [Parque Vera Cruz] (N). 3: 5h > hypo Bibliography: Glaz., Bull. Soc. Bot. France 58, Mém. 3: 5h0. 911; Prain, Ind. Kew. Suppl. 5: 153. 1921; Moldenke, Known Geogr. Dwarf pleasantly fragrant shrublet; stems apparently simple, unbranched, strict, virgate, slender, erect, obtusely tetragonal, rather densely and uniformly pubescent with short brownish hairs standing at right angles to the stem; nodes not annulate; princi- pal internodes 0.8--2 cm. long; leaves decussate-opposite or the rather num Ssprea rownish, eglandular; b and secondaries very Slender, plane or slightly impressed above, slightly prominulous 2 t econdaries ascending, et reticulation very fine, s impressed above and rather conspicuously pro. ous be- neath, terminating in the margi 3 oresce or cr in the uppermost leaf- - ; spikes subsessile ylin- to 1.5 cm. long, about 1 cm. wide, many-flowered; bractlets large, imbricate, membranous, surpassing the flowers, conspicuous- ay venone, conspicuously pubescent; corolla hypocrateriform, rose- Colored, This sp Auguste Francois Marie Glasiou — in whose honor it was named — in Brazil -- the first, no. 19170, collected among rocks at Biri- biry near Diamantina, Minas Gerais, and the second, no. 21918, at Serra dos Veadeiros, Gofas, deposited in the herbarium of the Muséum National d'Histoire Naturelle at Paris. A specimen of the Second-mentioned collection was photographed by Macbride in the herbarium of the Botanisches Museum at Berlin as his type photo- &raph no, 17509, but is now destroyed. Loesener's original description, while very fragmentary, seems — to be adequate under the International Rules of Botanic No- 172 P.B.F:T 0.1,0.0:1-4 Vol. 12, no. 3 cotype tcp: have been examined by me. Citations: BRAZIL: Gofas: Glaziou 21918 [Macbride photos 17509] (Br--cotype, F--photo of cotype, pe, It--photo of cotype, Kr-—photo of cotype, N—-photo of cotype, N--photo of cotype, N-—cotype, Si-- photo of cotype, W--photo of cotype, Z—photo of cotype). Bibliography: S$. Moore, Journ. Bot. 57: rte iets a. - Kew. Suppl. 6: 117. 1926; Good, Journ. Bot. 6 , ee 2: 139. 1930; Moldenke, Geogr. Distrib. Verbenac., (ed. 2) mer & 95. ect branched peotlen, branches leafy, pubescent; leaves de- cussate-opposite, sessile, * oblong or oblong-obovate, about 2 an. 1 and 7 mm. wide, gray in drying, obtuse at the apex, denticu- late along the margins or subentire, firmly membranous, scabridous above, pubescent beneath; inflorescence long-pedunculate; peduncles ascending-patulous, about 5 em. long, epamsenpek 3 spikes many- flowered, obovoid or subglobose, 1—1.5 cm. long, about 1 cm. wide; bhractlets lanceolate, surpassing the Hone acute at the apex, pubescent on both surfaces, about 1 cm. long when mature; calyx a~ broadly ovate, emarginate at the apex; corolla hypocraterifom, white, its tube about 3 mm. long, puberulent on the outer surface, about 0.3 mm. wide at the base, ampliate to 1 mm. above, the lobes pacha’ than the tube, the anterior one about 1.25 mm. long and 2 . wide, the e posterior one 1.25 mm. long and wide, and the later on ones 0.5 mm. long and 0.8 mm. wide; stamens Seer ee » bout 1 mm. long, glabrous; stigma oblique, about 0.7 mm ovary ovoid, about mba a long, shorter than the style; “nutlets sheik 1.25 mn. The type of this 1st lacknows species —_ ——S - John Gossweiler (no. 238) — in whose honor it w. amed - open thickets near Munong Rare rie Moore "2939), cea we at is "Easily told fran L. v wilmsii viet. . by the entire leaves rick ge: the bracts; the calyx of the two eile another point of con- ras Nothing is known to me of this species except what is stated i? the literature LIPPIA GRACILIS Schau. in A. DC., Prodr. ans pis— ait 1847. Synonymy: Lippia salviaefolia var. parvifolia Mart. ex Moldem ke, Suppl. List Invalid Names 6, in syn. aU, Lippia parviflora Gardner ex wm gee Alph, List Invalid Names Suppl. i; it, syn. 1947 [not L. parviflora Sessé & Moc., 1940]. Bibliography: Schau. in A. DC., Prodr. 11: 576—S77- lane bea in Marts , Fl. Bras. 9: 226-227. 1851; Jacks. in Hook. f+ Kew. 2: 95. 1894; Luetzelburg, Estud. Bot. Nord fate. 3: “bole 1923; Moldenke, Suppl. List Invalid Names 6. 1914 1965 Moldenke, Monograph of Lippia 173 Moldenke, Known Geogr. Distrib. Verbenac. [ed. 1], 37 & 95. 192; Moldenke, Alph. List Invalid Names 32, 19/2; Molderke, Alph. List Cit. 1: 223. 1946; Moldenke, Alph. List Invalid Names Suppl. 1: 15. 1947; Moldenke, Castanea 13: 116. 1948; Moldenke, Alph. List L85 (1948); 3: 691, 695, 712, & 770 (19b9), and he 1203, 1949; Moldenke, Known Geogr. Distrib. Verbe & hrub, to 2 m, 1, with a pleasant aroma almost like that of thyme (Thymus); branchlets slender, tetragonal, sulcate, pubescent- omentose; leaves small, very variable in size, decussate-opposite, petiolate; petioles more than one-third as long as the blades; leaf-blades thin, ovate or oval to obtuse, 0.6--1.6 cm. long, 3— pex, pennine long the margins, rounded at the base and contracted into the pet- i ovate, imbricate, subacute at the apex, connate at the base, ve g ous; flowers aromatic; corolla small, hypocraterifom, te; stamens yellow. Schauer (187) places this species in his Section Goniostachyum. He bases mn in Munich, where Macbride photographed it as his type photograph tenus ad inferior- * ° ibus tegentur albo-villosae". The chant hes Hook found on campos and "terra firma", flowering in April. A vernacular name recorded marajé", Gardner's L. parviflora was based on his no. 2289, collected Somewhere in Piauf, Brazil, and deposited in the herbarium of the Naturhistorisches Museum at Vienna, where it was photographed by as his type photograph no. 34327. The L. parviflora of Sessé & Mociffio belongs in the synonymy of L. umbellata Cav. Citations: BRAZIL: amazonas: Herb. Inst. Nac. Pesq. Amaz. 1035 (Ok); Herb. Jard, Bot. Rio Jan. proc. no. III [Pirarara] (Ja). Bahia: a. Lutz 113 (2); Martius s.n. [Macbride ene ~ oe to of t Baca “We-photo of type). Minas Gerais: P. Clausen 618 (Or Beit Be dectitee 2289 [wacbride photos 7] (It—photo, Kr—photo, M, N, Ne-photo) ; N. Netto 67 (Ja— 14892) . CULTIVATED: Brazil: Chagas s.n. [Herb. Inst. Nac. Pesq. Amaz. 1978] (Ok), son, (Herb. Inst. Nac. Pesq. Amaz. 3959] (Ok). 17h Pees CLOers Vol. 12, no. 3 LIPPIA GRANDIFLORA Mart. & Schau. ex Schau. in A. DC., Prodr. 11: 591. 187. Synonymy: Lippia grandiflora Mart. ex Schau. in Mart., Fl. Bras. 9: Bee "1851. Bibliography: Schau. in A. Prodr. 11: 591. Perf: Schau, in Bee) Fl. Bras. 93 2h7. isea;” geod in Hook. f. & Jacks., enke, e (19 943) and fed. a 80 & 190. ee Moldenke, Alph. List Cit. 3: 691 & 710. 1949; Moldenke, Revist. Sudam. Bot. 8: 167. 1950; Mol- denke, Résumé 93 & 61. 1959; yi Résumé Suppl. 8: 2 & h. 1964; Moldenke, Phytologia 12: 2h. 1 Dwarf subshrub; roots thick, Wesieiiaw: stems issuing from 4 woody xylopodium, about 30 cm. "lo ong, erect, fruticose, weak; branches tetragonal, hirtous; floriferous branchlets issuing from the lowest nodes of the stem; leaves decussate-opposite, very eure, Gaeta 3 leaf-blades’ lanceolate-oblong, recon at the apex, nate-serrate toward the apex, entire toward the base, 3-pli- merida pinnately veined, the young-ones eitdpaasteeeeas above, pube scent beneath and strigose-hirsute on the venation; peduncles axillary, solitary, elongate, bse peel heads compact at the s of anthesis, later more elongate, finally loosely racemose or subracemose; rachis villous; brastiete alternate, membranous, vate, concave or almost folded, acuminate at the apex, shorter cm. in diamete ed, 1 middle Lobat broadest, transversely elliptic, and the lateral ones broadly ovate and obtus Se. The type of this species was fhe scl, by Carl Friedrich Philipp von Martius (no. 157) in fields near Contendas, Minas Gerais, Brazil, is a very immature specimen, and is deposited = the Martius Herbarium at the Botanisches Museum in Munich, where section Corymbosae, He comments (1851): "Species ad typum L. lasiocalycinae condita, a qua differt villositate totius plantee, bracteis et corolla pro genera permagna." Material has been misidentified and distributed in h under the name Lantana sellowiana Link & Otto. Tastnatoll re (123) records the species fr from the Serra do Benedicto, Bahia, and fras Campinas de Duro, Gof as. I herbarium specimens and ); mounted photographs, in- = phototypes of both nanes involved, have been examined by “Citations: BRAZIL: Gofas: Macedo 369) (S, W--2059795). Minas Gerais: Heringer 3506 (2); Martius 1571 (Macbride photos 20326] 1965 Moldenke, Monograph of Lippia 175 (It--photo of type, Kr--photo of type, N--photo of type, W—photo of type); Saint-Hilaire s.n. (N). State undetermined: Glaziou 21892, in part (Br). LIPPIA Sane ate Hochst. ex A. Rich., Tent. Fl. Abyss. 2: 167-- 168. 1851 Synonymy : Lippia adoensis R. E. Fr. apud Robyns, Fl. Sperm. Pare Nat. Albert 2: 138, in syn. 197 [not L. adoensis Hochst., 1845, nor L. adoénsis Hochst., 181]. Lippia adoensis adoensis - ex uf ., Bull. . Jard. Bot. Brux., 32: § rag oo i in syn. 1 syn. 196 Bibliography: Walp., Roert. Bot. Syst. hi: 55. 18h5; me Rich., Tent. Fl. Abyss. 2: 167--168. 1851; Pha bere in G. Schweinf., Beitr, Fl. Aethiop. 278. 1867 ° Ind. look. f. & Jacks. Kew, 2: 95. 1894; K. Schum, in Just, Bot. Jahresber, 28 (15: 496. 1902; Moldenke, Known Geogr. Distrib. Verben enac., [ed. 1], 49, & 95. 19h2; Moldenke, Alph. List Cit. 1: 98. 196; Robyns, Fl. Sperm, Parc Nat. Albert 2: 138--139 & 616. 19h:7; —— Phytologia 2: 30. 197; Moldenke, Alph. List Cit. 2: 572, 6 hs 1032, 1084, 1174, 1177, 1182, 1202, 1218, 1220, 12h8, 6. l , lary, 1), in each upper leaf-axil, not developed in t cymes but in an elongate naked slender and distant-flowered ra- came; heads globose to globose-ovoid or cylindric, very short— pedunculate; bractlets oval or oval-oblong, long-acuminate at the apex, imbricate, equaling or almost equaling the corolla; flowers small; calyx in-Dillon ‘ a, e made by Geo Heinrich Wilhelm sabaigion (no. 73h) on the mountainsides adjacent 176 PHYTOLOGIA Vol. 12, no. 3 to the deep valley of Feurfeura. Schauer Spiel and Schumann (1902) both reduce this species to synonymy er L. adoénsis Hochst., of which the type is Schimper 1079. pia however, describes L. grandifolia and comments about it as follows "L. ramis strictis, “Yectis obtuse hexagonis bipedalibus tuberc lato-hispidis; foliis ternis basi vix petiolatis ahi iptaceehioa aut lanceolatis acutis, basi sensim attenuatis, margine grosse ser- ratis, utrinqe hispidis, asperis; capitulis globosis brevissine pedunculatis, in ane foliorum superiorum minimorum solitariis aut binis et inde emum elongatum nudum gracilem distantiflorum constituentibus; spaerats ovali-oblongis longe acuminatis undique imbricatis, flores aequantibus; calice compresso lanato, corolla dimidio brevio florens (Schimper). Observation: Je ne partage s l'opinion de M. Schauer....qui réunit cette espéce a la précédente [L. adoen- sis]. Elle en est distincte par des caractéres estrémement tranchés. Ainsi ses feuilles plus longues et plus larges et a dentes écartées, ne sont gaufrées; elle sont couvertes de de si que tige; les capitules sont portés sur des pédoncules trés-courts et ordinairement soli es a l'ais- selle es supérieures réduites 4 de trés-petites dimen- sions; le cal est oh omprimé et tout couvert de poils lanug- eux, trés-longs; 1 re la corolle est er t+ionnellemen b oup plus court que l'espéce pr The species has been collected on previously siaaieed land al- lowed to go back to bush, on tree sa my savannas, savannas with eollit on hills covered with herbs, n , at altitudes of 650 to 1900 gate flowering fran September to July, and fruiting from December strom? May, isk and September. Myers found it sept y among c grasses in neglected clearings; Hauman calls it ; aioe while Frisk preoree it as "arbuste atteignent la hauteur d'un homme". ema says “suffrutex érigé de 50 cm. de haut, petites fleurs vertes en chatons; stigmate trifide." Chapin notes "flowers small and white in heads that look mostly green". Robyns (1947) ongo Belge oriental et du Moyen-Katanga. En dehors du Congo Belge, elle semble confinée a l'Abyssinie et 4 l'afrique tropicale centrale et orientale." The L. adoensis of Hochstetter is a synonym of L. abyssinic® (Otto & Dietr.) Cuf. Cufodontis (1962) includes a "L. adoénsis-- Baker 1900 in part" in the synonymy of L. grandifol lia. The Drummond & Hemsley 4496 collection, cited below, is labeled "Sp nov." Vernacular names recorded for the species are nakuetria", “bulamurima", "dianga-bilondo", "djanga-malondo", t™mogira-ronda", "onga~ai", "umzirarondwe", and "wakayo". Material has been ni identified and distributed in herbaria under the names L. adoe=r 1965 Moldenke, Monograph of Lippia 177 sis Hochst., L. radula J. G. Baker, Lantana salvifolia Jacq., L. trifolia L., and Labiatae. all, 66 herbarium specimens, including type material of all the names involved, and 3 mounted photographs have been examined by me. Citations: SUDAN: Mongalla: J. G. Myers 7731 (K)3 Ne. D. Simp- Son 7333 (K). ETHIOPIA: Schimper I1.73h (F--photo of isotype, M~ isotype, N--photo of isotype, S--isotype, Si--photo of isotype, Z--photo of isotype). CONGO LEOPOLDVILIE: Bequaert 3296 (Br); Besquiere 5930 (S); Blommaert 185 (Br); Brédo 1307 (Br), 1400 (Br), 1170 (Br), 1555 (Br), 19ih (Br); Chapin 237 (N); Claessens 83 (Br), 1737 (Br, N); Debraene 182 (Br); Dewevre 758 (Br); De Witte 1651 (Br); Germain 508 (Br, Br); Ghesquiére 1,268 (Br), 5930 ( brun 350) (Br), 3678 (Br), 4588 (Br) ; Lejeune 3 (Br); Luxen 358 (B, Br), 543 (Br); W. Mullenders 596 (Br, Br, Br, N), 601 (Br); Seemed (Br, N); Robyns 2267 LIPPIA GRANDIFOLIA var. ANGUSTISPICATA Moldenke, Phytologia 3: 419-20. 1951. Bibliography: Moldenke, Phytologia 3: )19—h20 & 56. 1951; Bet tees Biol. Abstr. 25: 3051. 1951; Moldenke, Résumé 12 & ° 9 9. @ apex, canescent—strigillose on the back. It is a shrub, 1—1.5 m. tall, with creamyellow flowers. The type of the variety was collected by Van der Gucht (no. 364) in the Republic of Congo in 1912 and is deposited in the her- rium of the Jardin Botanique de 1'Etat at Brussels. It is knom thus far only from the original collection. Two herbarium speci- mens, including the type, and 2 mounted photographs have been ex- amined by me. Citations: CONGO LEOPOLDVILLE: Van der Gucht 364 (Br--type, N— isotype, N—photo of type, Z—photo of type). —— GRANDIFOLIA var. LONGIPEDUNCULATA Molderke, Phytologia 3: 71. 1950, 178 PHYTOLOGIA Vol. 12, no. 3 liography: Moldenke, Phytologia 3: Bt (1950) and 3: 56. 1951; aoe, Résumé 142, 183, & 61. This v ariety differs from the typic cal ited of the species in having its peduncles regularly up to i cm. long, It attains a on of feet and has white flowers. he type of the variety was collected by Richard Arnold Dttmmer (we 54) in elephant grassland, at 000 feet elevation, at Kepayo, Uganda, between July and September, 1911, a is deposited in the herbarium of the Royal Botanic Gardens at Ki In arium specimens, including tee type, and ), moun- ted photo er aphs have been examin y me. Citations: CONGO LEOPOLDVILLE: baciager 25 (Br). UGANDA: Dt mer 5), (F--photo of type, K--type, “W--isotype, N--photo of type, Sg--photo of type, Z—photo of type). LIPPIA GRATA Schau, in A. DC., Prodr. 11: 576. 187. Bibliography: Schau. in A. DC., Prodr. 11: S16. + 1875 Schau. Mart & Jacks. in 1. Bras. 9: 225-—226. 1851; Jacks. in Ind. Kew. 2: 95. 189; Moldenke g i ee ecaeaed [ed. 1], 37 & 95 (1942) and [ed. 2], 80 & 190. 199; Moldenke, Aiph. it. 3: 691 & 712 (1949) and h: 1176. 1949; Moldenke, 71 Résumé 93 & 61. 1959; Mold ones Phytologia 12: 2 ,» With the young parts sericeous-hirsutulous; branchlets numerous, slender, strict, spreading, pubescent; leaves decussate- posite, petiolate; leaf-blades ovate, about 2-5 cm. long and ide on 1 45, a. ° as @ _ < » ct ct B » on Ra re) e ot © » ct ct bm © oO ® rs) sessile a, shorter than the corolla-tube; calyx oblong, very shortly l-dentate, pubescent; corolla hypocrateriform, its te tube straight, about l, mn. long, the upper part surpassing the he bract~ lets, ventricose and pubescent at the middle, the limb nigrene; in drying, the lower lobe subquadrate, retuse and mucrona tanisches Museum in Munich, where it was photographed by ae as his type photograph no. 20327. Schauer (187) places the ref cies in his Section nm Goniostachyum. He describes the leaf-pube re cence as "pube plane relatis deinceps magis conspicuis odore oe 1965 Molcenke, Monograph of Lippia 179 pollentia" and says "Species indumento dense velutino glandulis ge 7 a ag interstincto et bracteis omnibus per paria connatis bus eximie distincta," It has been collected in anthesis in March. Material has been misidentified and distributed in herbaria under the name L. salvi- aefolia Cham. & Schl. In all, 1 herbarium specimen and 5 m mounted photographs have been exami neti by me. Citations: BRAZIL: Bahia: Blanchet 3886 (Br); Martius s.n. {Villa Nova da Rainho; Macbride photos 203 ye (It—photo of type, Kr--photo of type, N--photo of t; type, N--photo of type, W—photo of type). Bull. Acad. Brux, 11 (2): 327. 18h). Lippia graveolens Kunth ex Schau, in A. DC., Prodr. 11: 575. 1847. Lippia berlandieri Schau. in A. DC., Prodr. 11: 575—576. 18)7. Lippia berlandieri Torr. apud Coult., Contrib. U. S. Nat. a 2: “2: 328, in syn. 18 Goniostachyum ¢ raveolens (H.B.K.) Small, Fl. Southeast. U. S., ed. 1, 1012 & 1337. 1903. Lippia pinata: eri Millsp. apud P. Cc. Stand1., Field Mus. Publ. Bot. 3: 402, in syn. 1930. Lantana odorata Sessé & Moc. ex Moldenke, Prelim. ay List Invalid Names - Alph. List Invalid Names 32, in syn. 190. Goniostachyum berlandieri (Schau.) Tharp ex Moldenke, Suppl. List Invalid Names tee 3, in syn. 19h1. Lippia amentacea M. E. Jones ex Moldenke, Suppl. aa one Names S, in syn. 1941. Lippia sinveolans H. BK. ex M nke, Suppl. List Invalid Names 6, in syn. Hippia Suppl. 1: 10, in syn. 1947. Lantana \ avabeniaie H.B.K. ex Molden- ke, Résumé Suppl. 3: 32, in syn. 1962. Lippia geminata Schl. ex Moldenke, Résumé Suppl 1l: 7, in syn. ag st peg L. geminata H.B.K, 1818, nor Humb. & Bonpl., 1841, nor Humb. & Kunth, 156, nor Kunth, 1825, nor Millsp., 1930]. Bibliography: H.B.K., Nov. Gen. & Sp. Pl., ed. fol. pict., 2: 215-216 (1817) and ed. quart., 2: 266-267. 1818; Spreng. in Ly Syst. Veg., ed. 16, 2; 792. 38255 Steud., Nom. Bot., 5. Revod wart = &Gal., Bull. . Brox, 11 (2): et Bs Wap 3 t. ‘ tah Schau. in A. 576. 1847; ay tig tae oe “BLS; sen uéd, Mex. 13 "132—135. 1856; J. Torr...” Bot. U. . es - Bound. Surv, ~ 127. 1858; Bocq., Adansonia 3; 2),),. 1863; A. ray, Syn. Fl. N. Am. 2 (1): 338. 3°783 Rio de la Loza, Farmacia 2: 3-7. 1892; Coult., Contrib. U. S. Nat. Herb. 2: 328° 189); "hie ait took. & Jocks. , Ind. Kew. 180 PHYTOLOGIA Vol. 12, no. 3 23 hike 1894; Millsp., Field Mus. Publ. Bot. 1: 2 (1895) and 1: bo6; Kuntze, Rev. Gen. Pl. 3 (2): 252. 1898; Rose, Scatiel ‘Ue S. Nat. Herb. 5: 227. 1899; T. S. Brandeg.y 2 Zoe eld e 7; dui Ross State Teach. Coll. Bull. 19; 68. 1938; . C. Standl., Field Mus. Publ. Bot. 18: gern ensich 1938; loldente, "glph. ae, Wash. si 1. List & 6. 1941; H. S. Gentry, Carnegie Inst. ie Publ. 527: 223 & 306. 1942; Moldenke in Lunde il. Fl. Texas 3 (1): 53--55. 19423 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 12, 1h, 17, 20, 22, 23, & 95. 1942; Moldenke, Alph. List Invalid Nanes sd 28--30, & 32. 1942; Moldenke, Phytologia 2: 7h, 85, 88, 107, & 125 (1945) and 2: 156. 19h65 Moldenke, Alph. List Cit. 1: h, 23, 38, 111, 125--127, 148, 152, 176, 203, 216, 218, 227--229, 250, cere 299, 300, & 507. igh6; Reko, Bol. Soc. Bot . Mex : 26& ; M : - 1947; Moldenke, Alph. List Invalid Names Suppl. 0; 3 gh7 Moldenke, Bol. Soc. Vene i 2—-hh. 1917; Moldenke logia 2: 330, 331, 3h7, & 38h. 19L75 Ho NW. & A. L. nke, Pl. Life 2: 50. 1948; Moldenke, Wright 1: 237-238. 19,8; Miranda, Anal. aust Biol. Mex. 19 (2): ane 1948; Moldenke, Alph. List Cit. 2: 351, 370, 371, 3 468, scat "LT, 477, lioh, ee z2 A 538, 540—5h2 se 92, h, 59-596, 598, & 607 (us), : 656, 658, i660, 681, 683 5 708, 709, 738, 752, 765, 769, 773, 785, 787, 798—800, he 10, 817, 828--830, 833, 871, 876, a8 882, eh 914 ‘929, 925, 93e— 93h, 94h, 98, 954, 961, 963, & 973 (1949), and 4: 990, 991, 1099 8, 1159, 86 125, 1289, ayo, 1303. 1919; Moldenke, Known Geogr. Distrib. 6 > ’ 1d an, t. Guide Mex. Flow. Pl. 153, mss. 1950; ‘voldenke, PhY- ecrPicg 32 375+ 1950; Calpouzos, Econom, Bot. 8: 223-226 oe 230, & 232, fig. 1 & 3. 1954; J. Ruedowski, Ciencia 15: 118. 1955; Moldenke, Inform. Mold. Set 51 Spec. 3- 1956; i. « ae 1965 Moldenke, Monograph of Lippia 181 dowski, I ed ski, Act. Cient. Potos. 1: 49. 1957; Moldenke, Résumé 27, 30, 37, h2, bh, 4S, 47, 220, 297, 298, 306, 310, 312, 317, & h6l. 1959; Moldenke, Résumé Suppl. 3: 32 (1962), h: h (1962), 8: h (196k), th, 5, & 7. 196s C. M. Rowell, Sida 1: 268. 196k; Lan Select. Guide Lit. Flow. Pl. Mex. 177, 628, & 1010. 1964; Molden~ ke, Phytologia 12: 23, 47, 48, 56, 58, 63, & 115. 1965; Standl. & Calderén, List Prelim. Pl. El Salvador. n.d. 7 hapa Calpouzos, Econom. Bot. 8: 223 & 225, fig. 1& 3. 195k. and low slender aromatic shrub, usually 0.3--3 m. tall, or rarely & small tree to 9 m. tall (Standley), strong-scented with a sweet spicy fragrance like that of cinnamon (Cinnamomum zeylanicum), rs; leaves most s ant on odor; leaf- blades taceous, oblong or ovate-oblong to elliptic, 1—6.5 cm. long, 0.5—-3.5 om de btuse or rounded (sometimes a- cute} at the apex, rounded or subcordate at base, somet les axillary Per node, spreading, u ly equaling or slightly longer than the petioles, appressed-pubescent or strigose; spikes oblong, 4-~12 m. long; bractlets conspicuously l-ranked, conduplicate-carinate, mnate an ally apex, not venose, gl ose, more or less densely hirsute- to yel -ow in age, mostly turning brown in age, fragrant, hypocrateriforn, its tube 3--4 mm. long, canescent-strigose on the outside. Campeche, is probably deposited in the Bonpland herbarium at the Muséum National d'Histoire Naturelle at Paris. Schauer (1847) places the Species in his Section Goniostachyum. L. amentacea is based on M. = Jones s.n. from Laredo, Texas, deposited at Pomona College. BOOK REVIEW Alma L. Moldenke aoe ae Biochemistry and Serology", edited by Charles E. Leone, a + 728 pp., illustr. Ronald Press, New York 10, New York 196l. $16.50 the International on Taxonomic Biochemistry, Physiology, erology at the slaieatter of Kansas in 1962 as a first effort at gathering toge rmation on molecular omy to " n 8 place for Gtedente and scholars who feel that the next great pi dloyicas synthesis will be at the molecular level". PART I : PRINCIPLES OF SYSTEMATICS The old. was shown by Arthur Cronquist to be an absorption of several ne new sets of principles — Linnean, deCandollean, Darwin- ian, eae cytogenetically enriched, populations centered, alo ong wi th all present new forms which surely will not be the last, nor eytit they be the replacement for the basic classical work of the past and present. So also claims Ernst Mayer who describes the new as different from the old mainly in viewpoint which is biological rather than nominalistic-topological. He stresses the use of ever increasing number of kinds characters, a ready acceptance of new tools ti t larifica taxon VS« category, nested fs of the subspecies as a category ee 5 not an evolutio query age and causes for similarities and differences among t: reminds his readers that "we can never base Fp i pt on seriaieke (since we do not know it), but from our classification we can draw inferences on the relative proba- bility of various possible phylogenies". Contrary to the Adanson- onc eature weigh le gories the dotegrating power of the brain of an intellig experienc axonomist", Rather than inductive ‘no weighing" and deductive "a pute weighing", he c ions inductive-deduc "a posteriori weighing" pane is basically the experienced scien- tific method. He concludes with a brief critique of numerical ney but ea- ewpo ¢ jects any "weighing" and is limited to the pre ae ) phenetic distances between units and groups. re The future, predicted by Robert Sneath, will "include 4 m0 intensive stu aba of evolutionary mechanisms, particularly P tion genetics, both under field o nidis ne ane in the Laboratory under realistic assumptions." "Taxonomy of the higher categor® 182 1965 Moldenke, Book review 183 be revised by some kind of — taxonomy." Development i objective machine methods for erecting taxonomies will remove f the personal and subjective sae with poche ha no- aerenatured, stability. Scientific names of organisms ha ur great museums to the conventionally preserved material (to be actasa rather than discarded, I hope) will have to be added specially preserved newer materials such as voice tapes or r cordings, chromatography, etc., records, along with information storage and retrieval machinery. If this none becomes modi- fied in the Cain & Harrison manner of phyletic weighing or that of Mayer explained above it will be more sccoptable jd the func- tioning biosystematists of yesterday and today, their hopes for se future. : PERSPECTIVES IN MOLECULAR TAXONOMY Marcel Florkin maintains that "the greatest virtue of a so- called natural system of taxonomy resides in the fact that clas- ——— gous in all cells, but not always homologous as = it may be the Product of the action of a chain of biocatalysts an Spyoclyess ° or differently of oxidative phosphorylation -- evidence of conver gence at the molecular level. Alan Boyden explains the value of systematic serology through Precipitin testing with first homologous and later heterologous antigens, yie elding a relative placement series which is — useful in showing relationships in blood-containing animals mrad Scheer describes the role of comparative physiology “in system ro myriad forms of life. Leaders in the bio-courier project explain ompara: es. r discuss the problems of intraspecific hy- 18h PHYTOLOGIA Vol. 12, no. 3 bridization in the comparative chemistry of Baptisia. By chro- genous anthocyanins, shows them to be red-violet chromo~alkaloids and limited to 9 or 10 families in the Centrospermae and closely allied orders. ey and Hopkins' chromatographic studies on the Rigidae and Retusae sections ou on genus Haworthia gave differe nt patterns on hho Big A the lant when treated at the same time. iaston, Mave toetig a biochemical pro ee i = Bip ay est da pool of chemical constituents of B. violet absorbing compounds are by far ve pagers. practical for ready development of biochemical profiles, for eee studies, ec Moritz confirms the idea that serological hams IE tend to show evolutionary convergence. Lester's double diffusion and immunoelectrophoresis of protein extracts of ten South American and twenty North American species of Solanum found the former all to have general uniformity ° antigen igen and the latter to have two main antigens. By similar further testing 130 encion of the genus seventeen series were Fairbrothers and Johnson's precipitin test results on the Cornaceae and Nyssaceae show close agreement with present taxo- nomy. PART V : COMPARATIVE BIOCHEMISTRY OF ANIMALS VI : SEROLOGY OF ANIMALS PART VII : MOLECULAR TAXONOMY OF MICROORGANISMS PART VIII : TAXONOMIC ELECTROPHORESIS OF ANIMALS Common problems and principles of classification, common bio- chemical pathways, common testing techniques, equally or standing authors make the papers in these sections valuable an interesting to botanists. 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Under the present cost of printing, the basic rate for a page or fraction thereof is approximately $3.00 for an edition of 250 copies. ‘This price is subject to without notice, since it depends entirely on the prices prevailing in the printing industry. Reprints will be furnished at cost. A proportionate fraction of the edition - of 250 copies is also furnished gratis to contributors. Upon request, the editors will send detailed instructions concerning the preparation of manuscript or further information about the magazine. In- quiries may be addressed to the magazine or to either editor. PHYTOLOGIA Designed to expedite botanical publication : / Vol. 12 June, 1965 No. 4 CONTENTS FISCHER MATTOS, N. , A new species of Bauhinia for the flora Set Paulo. oo eee 185 MOLDENKE, H. N., Materials toward a monograph of the genus Pe ee 187 BeOHLl. C. 1.” Additions. io the Lamubede of Guatemala ....... 243 PeOkE, Ai 15 Both poles 246 a S Published by Harold N. Moldenke and Alma L. Moldenke 15 Glenbrook Avenue Yonkers 5, New York, U.S.A. Price of this number, $1: per volume, $5.75, in advance, or $6 at close of volume Vol. 12, No. 1, was published May 12, 1965; Vol. 12, No. 2, - < 0m May 25, 1965; and Vol. 12, No. 3, on May 28, oe {op . Wi=0um poramenr JUL 4 12 1965 __SARDEN Liprany A NEW SPECIES OF BAUHINTA FOR THE FLORA OF SHO PAULO Eng? Agr® Nilza Fischer Mattos BAUHINIA HERMESIANA N,Mattos Spe ne — Arbor aculeate circiter 10 m alta (ex collector). Ramuli terminales ter PA pilis brevibus cinereis adpressis dense obtecti deinde Seber soihag et rented et é peect cae ad extremitatem peulo incrassatis et sulca- tis, usque 3 cm seeks s Svea uaine 653 - longa et 9,5 cm datng: triangulari, 9~ palmatin adpresso-cinereo-pilosi » Pluriflori. Alabastra elliptico-lenceola - > j eahbe obtugo-apiculata. Flores pedicello cc. 12 m longo dense e ful ua erne S Tregulariter rumpenti, dente uno bidenticulato, petalis 5 vexillo seyoscents excepto albis, distincte brunneo-vervatis, extus pilis in im af: bl i stipite 4-5 nk longo dense adpresso-piloso suffulto, stilo 9 mm longo inferne Piloso,. Zypua: Hermes M, de Souza see BRASIL — Estado de Sio Paulo: Munic{pio de Pereires, entre a e Pereiras, Fazenda Sio Iuiz, 2/TI1/1964. "Na mata secimdé- * “rvOre de mais ou menos 10 m de altura. Flores cas ®standarte amarelo", Hermes M. de Souza sen. (SP 30242, HOLOTYPUS), In this work we present a new species of the genus Baybinia, 185 186 PHYTOLOGIA Vol. 12, no. of the Femily Leguminosae. A species was sent to us by our tree = pina gre = fe) the genus called our attention, since it s s different from the species already kmown and frequently id atitiad. for peti State of Sao Faulo. After an extensive bibliographic research, we reached the conclusion that we had a new species of the genus. The e was chosen in honor of the collector, Agronomist En- geneer Secade Monsire de Souza. We take the opportunity to give the bangers ae Mr. Oswaldo Handro many thanks for the Latin diagnosi Bauhi -Mattos.- a) stem with leaf; b) 3 ¢) flor Saubinia-hemesiana ¥ gynecium; e) calyx; - eae ae Nj ; MATERIALS TOWARD A MONOGRAPH OF THE GENUS LIPPIA. IV Harold N. Moldenke LIPPIA GRAVEOLENS H.B.K. Lippia berlandieri is based on three collections: (1) collected Vittoria, Botaniques at Geneva, where it was photographed by Macbride as his type photograph no. 33929, (2) collected by G. Andrieux (no. 166) near Oaxaca, Mexico, before 1835, and (3) collected by Carl August Ehrenberg (no. 720) on mountainsides near San Bartholomo, the last two originally deposited in the herbarium of the Botanisches Museum at Berlin, but now destroyed. The type of Lantana origanoides was collected by Henri Guillaume Galeotti (no. 784), but Dr. Robyns has reported to me in a letter dated April 3, 1939: "We have been unable to find in our herbarium (Jardin Botanique de 1! Etat, Brussels] the type specimen Galeotti 784 which is probably lost. But the specimen I am sending [Gale- otti 756] has been authenticated by Martens & Galeotti." Lippia Suaveolens is based on a collection made by Marcus Eugene Jones (no. 29373) at Laredo, Webb County, Texas, and deposited in the herbarium of Pomona College at Claremont, California. It is worth pointing out here that the Lantana odorata of Aiton, of Kunth are also Lippia alba (Mill.) N. E. Bre, while that of Millspaugh is Lantana microcephala A. Rich. Lippia graveolens has been collected on mountains and low moun- tains, hills, limestone or limestone-shale hills almost destitute of soil, low or rocky hills, gravel or gravelly hills, "dry crevic- ious hills", steep or bushy hillsides, gravelly hilltops, mesas, deserts, rocky or dry rocky slopes, and brushy plains or woods. grows » shrub gras Sociations with oak, pine » and juniper on mountain summits, among a heed oaks, at the base of rocky slopes, along rivers and creekbeds, at altitudes of 50 to 3100 meters. In Texas it in- habits dry and ro » valleys, arroyos and inc and open desert scrub from Brewster to Cameron northeast to Austin 187 188 PHYTOLOGIA Vol. 12, no. & and Houston Counties. In Yucat4n it grows in rocky soil and on the slopes of hills and mesas. It grows from southern Texas south to Costa Rica, flowering and fruiting in every month of the year. In Texas it is said to bloan from shone to December, while Palmer & railroad Terrell County, a describes it as "frequent" and abundant in gravelly hills" in Starr County, while Hanson reports it "fre- in canyons" and "common in roc eys" Brewster County Warnock says it is "infrequen mt along roads", "Ninfrequent... ong kley Cae and widespread in "fete soil. on low sedimen Tn Wes it was collected by Waterfall in "clay valleys with acacias and other shrubs" in Coahuila; Hinton found "small colo- nies in rock-jumbled canyons" and Muller says it is an "abundant constituent of the desert shrubs in Larrea-Flourensia = dese in the same state. In Durango it occurs in "shrub-grassland on the south slope of limestone cerro" according to Gentry; in in Zacatecas McVaugh describes it as "occasional in stream valley nearby rocky oak-covered mountainside", while in the state = pra it inhabits "deciduous spiny woods in barranca at edge of river" = "wet roc 8 c cording to Crutchfield & Johnston, it lives on "limestone hills with short brush", Rzedowski found it in "ladera caliza con vee? tacién de Helietta parvifolia" and in similar localities with Agave striata. Standley found it "abundant on danp brushy slopes" in gardens.....The aromatic leaves are used in Costa Rica for fla voring food, also medicinally." Calderta & Standley (1941) state “ 1955 states that in 27 years of botanical stietien 12 4n New Mexico he never seen this species in that state. It is possible that a the two collections cited below from New Mexico are mis-labeled 29371, for instance, is a form with large, thin, hairy leaves} o L. Fisher s.n. (Aug. 20, 1937] has the leaves dark, thin, aa hairy, and is certainly not very similar to e 11083. It been widely misidentified — Millspmgh (1895) cites it as Le = a eee 1965 Moldenke, Monograph of Lippia 189 anoides, while Lundell (1937) cites it as L. alba. Standley (1930) ds in error in stating that the L. geminata of Millspaugh (1900) and of Millspaugh & Loesener (1905) is L. graveolens, Ac- tually, the specimens cited in both these references are Lantana microcephala A. Rich, His comment (192k) » however, to the effect that it is very doubtful whether L. berlandieri differs in any im- portant characters from L. graveolens, is quite true. Following eries of specimens from Texas, Mexico, and Central America has two are c ic. Collectors differ considerably in their description of the flower color, The corolla is described as "yellow" on Muller 3002, Stanford, Retherford, & Northcraft 96, and Warnock & Sur- ratt 9827; "yellowish* on Lundell & Lundell 9792; "cream-color" on Correll & Johnston 18041; "creamy—white with a yellow eye” on Iundell & Lundell 9906 & 12340; “cream changing to yellow in eye" on Sharp 3 “cream-white with yellow throat" on Feddema 12,7; "white with yellow eye" on Lundell & Lundell 9807 and McVaugh 17665; and "white" on Clover 39 & 1676, & 2353, Hinton 1S7L, 13061, 13162, & 13318, Lundell & og & Barneby 13279, Runyon 1721 & 2538, Smith, Peterson, & Tejeda 3977, Standley 71025 & 7094, Steyermark 29371, Tharp, Johnston, & Tur- ner 3509, and White 2072 & 2120. ernacular names recorded for the species are "ahuiyac xihuitl "erégano", “hahauiyac xihuitl", "hierba dulce", "hierba olorosa", " ‘ano ocephala Schlecht. & Cham., and L, umbellata Cav. Standley (1930) states that the application of the names "tabay” and to this species is doubtful. Calpouzos (1954) says "Oregano is becoming a popular herb in America, The botanical identity of the plants sold as oregano in doubt. Most of i} the genus Lippia, of the family Verbenaceae, and come from Mexico. The rest of our supply of this herb comes from Europe. These latter Plants belong to the gems Origanum of the family Labiatae. Species in other genera are also called ‘oregano’, or some variation of this nam, bis Lippia affinis Schau., L. formosa T. S. Brandeg., L. 190 PHYTOLOGIA Vol. 12, no. k Turez., L. micromera Schau., L. micromera var. helleri (Britton) Moldenke, L. origanoides H.B.K., L. palmeri S. Wats., Wats., L. palmeri vars spicata Rose, Lantana achyranthifolia Desf., L. involucrata L., L. trifolia L., “Coleus amboinicus Lour., Hedeana floribunda Standl., H H. patens Jones, Hyptis albida H.B.K., H. 3) He americana (Aubl.) Urb., H. capitata Jacq., H. suaveolens (L. L.) Poit., Limno- phila stolonifera (Blanco) Merr., “Vonarda austromontana Epl., Ocimum basilicum L., Origanum majorana L., 0. vulgare L., Polio- mintha longiflora Gray, y, amd § Salvia sp. sp. ~~ Gaumer (1907) states: "Lippia gi graveolens: stimulant, expector- and tonic. The fluid extract of Tippia is a diffusible stin- ulant which is employed with excellent carminitive effect against flatulency and nervous meteorism, is appreciated greatly in primi- tive fevers. It is also much used in the non-febrile state of cholera morbus and against all kinds of excessive fluxes, also 4- g ertain and stupor of the senses consequent upon serious illnesses and the ss which appears during convalescence; against headaches of debilitate, delicate, and nervous persons, eye-ache, vertigos ob- served in people dedicated to intellectual pursuits, and produ a happy cure for essential hypochondria. It has expectorant properties and effects changes in the bronchial mucous membranes. It is an excellent remedy for chronic bronchitis, and also for some catarrhal affectations of the respiratory system. Dose 1 to sed as ac ent; also use (192k, 1930) confirms that the species is used in aqeeete medicine t4n stinul , and de- » tonic, expectorant, mulcent, especially in the treatment of mi ta morbus, fevers, bronchitis, and catarrh, and that the aromatic leaves are use ed for flavoring food. Material of L. graveolens has been misidentified and distribu tod se oa under the names L. alba (Mill.) N. E. Bre, Le : geminata H.B.K., L. lantanoides Coult., L. origanoides HeB-Key poe wrightii A. Gray, Lantana involucrata fag, L. macropoda Torre, Xs be montevidensis (ee) Briq., L, sp., Larrea tridentata DC, Tournefortia = Sichocalycine DC., Aloysia sp., and Satureja 5P- "On the other hand, the V. H. Chase 7h82, R. S. Ferris 5927, Me E. Jones 23563, and York s.n, (Brownsville, 3-19-1907], 907], distribe ted as L. graveolens, are es alba (Mill.) N. E. Bre; a 1s7L & 2983, Galeotti 752, and Edw, Palmer 1137 are L. hypole’s Briq.; Carran & Haman 606 & 9 966, Dawe 199, ‘and Saer 62 are Le origanoides H.B.K.; Galeotti 795j is L. pringlei Briq.j Shreve 1965 Moldenke, Monograph of Lippia 191 6150 & 6173 are Lantana velutina Mart. & Gal.; and Whitehouse s.n. {El Paso] is not verbenaceous, possibly a mint or a species of torium a * Tharp son. [6-16-1928] is a mixture with Acie ene iat Gray) Heller, Frye & Frye 2353 is a mixture with Torr. and L. microcephala Ae A. Rich., Warnock 831 = a oe nore Lantana macropoda Torr., and Steyermar k 1,6285 is a mixture with Phyla scaberrina (A. L. Juss.) Moldenke. ~~ The label on E. W. Nelson 6802, cited below, originally was in- scribed "6201" in the Britton Gerbart in. Merdedes 8 Aguilar Hidalgo's name is often written "Mercedes Aguilar® or "Mercedes Aguilar H." E. P. Johnson 81 bears a label inscribed "Yucatan & Tabasco", but Millspaugh (1995) cites it as from Yucatén. The Ecology Class (isis Univ. texas §.n., cae below ate = Webb County, may pes: have come from | Zapata e label ds "Laredo—Zapata Glaziou (911) sities his no. 9987 from Espirito make. Brazil, but this is obviously a misidentification. Gentry (19:2) cites Shreve 6150 and 6173 from the Shreve Herbarium » but these collec- tions are both Lantana velutina Mart, & Gal. The records of L. graveolens from La Libertad, Guatemala, in my previous publications were based on Aguilar Hidalgo 40, which proves to have come from a locality "La libertad" in El Petén. In all, 440 herbarium specimens, including type or authentic material of most of baad names involved, and "e mounted photographs auton eearan igel re & Schott ons: TEXAS: ft Co.: Parry, Bigelow, 819 wae Brewster Co.: H. C. Hanson on 61h (N), 709 (Ka—6065L) 5 ey 1555 (Warnock 161052] (N); Hinckley & 5 Serieer a ae peel 13065 (Rf), W.631 (N), s.n. (Sperry sky, (on)s Earnest W603 Teco 3730 (N), 3739 (N), 461031 {Hinckley u, 8), Leto [Hinckley 1550] (Au, Ss), 161052 52 [Hinckley 1555] (Au). Cameron Co.: Nealley s.n. [Pt. Isabel, 1891] (Au); R. Runyon 154 (8); A.C. V. Schott s.n, [Rio Bravo del Norte] (Cc). El Paso Co.: g werent co aie mead, G, T)s Hidalgo Co.: Clover 39 (Mi, N); Johnston 18041 (RE); 3 Cory 51337 (Sm); Je a Drushel 6323 (U0 oy Lundell & Lundell 8803 (G, Id, Mi, N), 9607 (Id, N), 9906 fa, N)3 I. Shiller 736 (Au); Small & Wherry 11921 (N)s Tharp ae 51-252 (St); west 2h sen. (Pharr, June 18, ae (S); Mrs. E. Houston a cs M. Tracy 9158 ae o, It, N, Tr, Tr, Tr, Tr, Up— ns Ws). Jim ii Hogg Cot 3 Tharp 590k 590 (an). Maverick Co.: Pringle 2034 (G, It, Me, Me, Me, Vt). Presidio Co.: Havard 2h (G). Starr Cows Clover 1375 (Fs, N), 1676 (I, Mi, N)3 D. S. S. Correll 1,895 192 PHYTOLOGIA Vol. 12, no. k (Rf); Inmes 397 (G); Lundell & Lundell 9792 (N); A. R. Moldenke 162 (Fg); Ripley & & Barneby 907 9047 (N)3 R (N)3 Rose & Russell 24363 (G, ¥— 1369640); R. Runyon 1721 (Rr), 2538 (N, N); Tharp 5905 Tharp 5905 (Au, Au), Son. [North of Rio Grande City, 6-16-28] (Mi, N); C. C. Wright Sn. [East side of Rio Grande, 1848] (G). Terrell Co.: “Warnock | & . Sur ratt 9827 (Rf). Val Verde Co.: Cory 3416 (G), 19427 (G), 7 (G), 26273 (tr), 38065 (N, N), sen. (Langtry, July ay 192k) (Tr), ; Son. [Langtry, Aug. 192h) (Tr); G. L. Fisher s.n. (Langtry, July 18, |, 1922] (Hp, Vt); Parks & Cory 19425 (Tr), 26719 (Tr); Rose & Fitch 17978 (¥)5 E. D. Schulz 2012 (Wi). Webb Co.: Baird s.n. sn» {Fall, 1936] (N); Canta, Covell, &1 & McCart 33 (Ok); Ecology Class Univ. Texas s.f+ [Laredo-zapata, 2.29.30] (Au); M. BE. Jones 29186 (G, It, P Po 199863), 29373 (Po—-200131, Po—201279), Sone i DEES ash 199873); Martinez & Trevino 30 (0k); Tharp, J Johnston 3509 (St). Zapata Co.: D. S. Correll 20782 athe cory BL oy tH), 35930 (N, N); Tharp 3697 (Au). County undetermined ard S.De [1881) (Mi); Nealley ek) (Du--90916); Neatby s.n. nee Texas, Oct. 1890] (N). NEW MEXICO: County ileterianedy LeRoy 5-0. (Pa); C. W ee 1507 [Paneter Caves, N. Mex.] (G, T). MEXICO: “Campeche: a S. 5S. Waite 206 3 (G, Mi), 2072 (Du--278210, Abi. 21.20 (po--267908, G, Mi). Coahuila: Aguirre 06 (N); Aguirre & Rekp 37 (N) Herbe (G, N)j I. M. Johnston 7038 (G); M. E. Jones 361 (Po—68825)5 Kenoyer & & Crum 2622 (a, MA), 3164 () (Mi); E. J. eae 418 (St), ple (au); M. Martines 715 (Me); 0. H. eps 2 , in part (Ca— 719562, Mi); E. W. Nelson 6802 (N); ae 362 (G), 370 (Ca—882h81), 736 (A, G, G, N), ere (Cc, G, 10, Pa, Vt), 1026 (G, Pa); Bees co) Pringle 25 (G, Vt); 1 Ee ene hee Fs, Mi); Shreve & Tinkham 9579 (Ca--661998, G, Mi); Re H, Schroe sohroe- der 31 (os te sale, Retherford, & Northeraft 96 (Ca— a-—715862, Du—291288, G, N, Se—70); 9h); Warnock & Barkle -1y82lm (Au); Wa terfall ast (St); S. S. White 1938 Git 0658); Wynd & Muel- ler 67 (Fs, G, I, N, S, St). Durango: H. S. Gentry ‘B33 (Ak 22803, G, be 3 E. ¥. Nelson 169 (G); Edw. Palmer 539 me 10L9L8, G, N, S, S). Federal District: Northcraft 7 (Ia Bectatonee. 1616 (Kr). Guerrero: Degener & Degener 20210 “26216 Gs, 0s Hinton 6902 (A, K, N); Lemmon & Lemmon 203 . (Ca--10095h, Sh, G)5 Bs := Moore maa (Ca--918895, G, N); Edw, Palmer 167 (C, C ca—104927, & 1965 Moldenke, Monograph of Lippia 193 dell 12340 (Ld, Ld, Ld, Rf); Matuda 2956) (z); Purpus 141) (Ca~~ 139656, G, N, Po--6)3h). México: Hinton 2655 (A, N, N), 5269 (G, N, N, N), 6370 (G, N); Matuda & al. 31723 (Ss), 31934 (Ss); Northcraft 7 (N). Michoac&n: Hinton 13061 (Au, Du--290363, Mi, N, N, Rf, S, Ur), 13162 (G, Mi, N, N, Rf, S), 13318 (G, Mi, N, N, W Rf, W-~2020809). Morelos: Frtderstrtin & Hultén 189 (S); Moldenke & Moldenke 19831 (Es, Lg, Mg, Mr, N, No, Ot, Sm); Pringle 11083 (Cm, Fs, G, Gg—l21291, It, Mi, N, Vt). Nayarit: Feddema 12h7 (Mi). Nuevo Leén: F. A. Barkley 196A (G); Fernandez C. & Bark- ley 1h96a (Au, N); G. L. Fisher s.n. (Monterrey, July 1h, 192)] (Cm, Du~—175925, Fs, Vi); Frye & Frye 2353, in part (It, N, Pl— 111102, Se--6559); Heard & Barkley TLSh7 (au, N); Johnson & Barkley 16035M (Au, Ca—76328), G, N); Kenoyer sn. [Sabinas Hid- algo, Sep. 15, 1937] (Fs); Mueller & Mueller 159 (Au, G, Me, Me, Ur); Pringle 193, (Br, C, Ca—104917, G, Mi, Pa, S, Vt), s.n. (near Monterey, l, August 1889] (Ca--104918); Quarles yan Ufford 22 (Ut); Seler & Seler 1053 (G), 1092 (G); Shreve & Tinkham 9613 (Fs, G, Mi). Oaxaca: C. C. Deam 100 (G); Galeotti 756 (Br, Br, N- Photo); A. Gonzalez 410 (G); V. Gonzalez 19 (G); H. S. McKee 21001 (Lw); E. W. Nelson 1210 (G), 1977 (G)j Pringle 6258 (Br, C, Ca~-104928, Cm, G, G, Gg——152308, Gg—162998, Io—30719, Me, Me, Me, Mi, Mm-~15),09, S, Vt); Rowell, Webster, & Barkley 17490 (Mi, N); Seler & Seler 1391 (G); L. C. Smith 696 (G, Me). Puebla: Bravo Hollis 7iu5 (Me); Patoni 1007 (Me); Purpus 462 (Ca—13966k), 3401 (Ca~-13965)); Rose & Gay 5930 (G); Smith, Peterson, & Tejeda 2977 (G, N, W—239799h). Quintana Roo: G. F. Gaumer 1527 (F— 58325). San Luis Potosf: G. L. Fisher sen. (Aug. 20, 1937] (Fs, N); Edw. Palmer 711 1/2 (Pa); C. G. Parry 7111/2 (Io); J. Rze- dowsici 61,79 (Au--170136), 6686 (AU--170117). Sinaloa: T. S. Brandegee 8.n, [Cerro Colorado, Nov. 1, 190k] (Ca—10h916); J. Gonzalez Ortega 6602 (Du—17330h, G); M. E. Jones 23233 (Po— 153925); Edw. Palmer 2771 (G). Tamaulipas: 0. M. Clark 6641 (N); Crutchfield & Johnston 1986 (Au--175055); Kenoyer & Crum 3545 (G), 36uL (6); Edw, Palmer 520 (G); Perkins & Hall 332k, in part (It); Stanford, Lauber, & Taylor 2226 (Du-366137, N, N). Vera Cruz: 19h PHYTOLOGIA Vol, 12, no. (Ca--368992); R. McVaugh 17665 (Mi); J. N. Rose 212 (G); Shreve 94,00 (Ca--731697, Fs, G, Mi). State undetermined: Berlandier 832 [de Santander a Vittoria] (G, T), 2252 (Macbride photos 33929] (4, Kr—-photo, N--photo, ger T); Coulter 1162 (G); C. C. Deam gon, {Salina Cruz, Dec, 21, 1898] (Mi); pees, OB g lh3 (G), > 496 (@); Haenke 1470 (N), 1583 (N); piel B.T.815 (V); ¢ Ce Ce Parry SMe [1852] (10); Purpus 1181 (Ca—139657); sary Mocifio, Castillo, | & Maldonado 2126 (Q), 2196bis (Q). GUATEMALA: El P a Pet “Aguilar Hidalgo 0 (F— (F—713089, G, 1, Mi, N, N); Steyermark 46285, in part (Mi). Zacapa: P. C, Standley 7028 (W--18),29),0) , THO, (N) 5 Steyermark 29371 7. (F=~104 3423). BAY ISLANDS: Mugeres: es: Ge F, Gaumer son. (Sg--66302). NICARAGUA: Grenada: Lévy 250 (Cb, Cb, Cp, “Cp, N--photo, Z--photo). COSTA RICA: Alajuela: Brenes 23110 (N). Guanacaste: Brenes s.n, [1910] (N, Si). San José: H. H. Pittier 1681 (Br). CULTIVATED: Costa Rica: Brenes 12238 [117; 717] (N)- Hl Salvador: Calder6n 77 (G). LIPPIA GRISEBACHIANA Moldenke, Phytologia 1: 279. 1938. Synonymy: Lippia lantanifolia Griseb., Abhand. Kaiser. Gesell. Wiss. GOtting. 19: 242, 187) [not L. lantanifolia F. Muell., 1868). Lippia asperifolia argentiniensis Gill. & Schau. ex Gri- seb., Abhand. Kaiser. Gesell. Wiss. abtting. 19s 242, in nits a Lippia lontanifolia Griseb. ex Alvarez, Fl. San . del Estero 1 sphalm, 1919. Lippia lantanaefolia asiek: ex ey “tik List Invalid Names Suppl. 1 ‘plate in syn. night Bibliography: F, Muell. - 6: 151. 1868; Griseb., Abhand. Kaiser, Gesell. Wiss. oettine. “Ss 2h2. 1874; Griseb., Pl. Lorentz. 19h=-195. 1874; Lorentz, Veg. Nordeste Prov. Entre Rios, ed. 1, 150. 1878; Griseb., Abhand. Kaiser. Gesell. Wiss. G&tting. 2h: {Sym ymb. Fl. Argent.) Tr. 1879; Lillo, Fl. Tucuman 9h. 1888; Jacks. dense zin, 102 & 103. 1919; Alvarez, Fl. Sant, del Estero 106. 1919; Stapf, Ind. Lond a. bs 125. 1930; Moldenke, Phytologia Ls 279. 1938; Mol- Alph. List I ames 31. 190; —_ S$: ho bos ‘ peoaden 190; Moldenke, My tiliela 1: ‘Al. “Told; Mol- » Known Geogr, Distrib. Verbenac., Se a P oes ghee ete 0: Yoldenke, Alph, List Invalid Names m Fis 343. 194; Moldenke, Phytologia 2: 83 & tor. ashy Fe BF rf 2, denke, Alph. List Invalid Names Suppl. i: 14. 1947; Moldenke, Phytologia 2: 386. 1947; Moldenke, Lilloa ly: 33, %, & 2. 19b8 . He Ne eh Le Moldenke, Pl. Life 2: 62. 1948; ne Fo Algh. Cit. 2: 368, 377, 378, hhO--hh2, 535, £86, 600, & 620 (1948), 661, 662, 673, 732, 73%, 76, 80h, 863—865, & 907-910 C1918)» 1965 Moldenke, Monograph of Lippia 195 and 4: 1080, 1089, 1091, 1092, 1148, meni nome 1203, & a3 ee Moldenke, Known Geogr. Distrib. Ver » [ed. 2], 10h & sooad by Résumé 125, & Erg 19593 Woldenke, Résumé Su Suppl. 10: 6 (196) and’ air 5: 1964; Moldenke, Phytolo gia 12: 47 & 61. 1965. Tlustrations: Sanzin, Anal, Soc. C Gientif. Argent. Buenos ; manera (im—iv): 103. i919; Troncoso, Darwiniana 10: 74, fig. Aromatic woody or subligneous shrub, 0.3}--3 m. tall, branched; branches cance uae branchlets subtetragonal, stri ate, with reddish-bro bark, peta pa? or with er dispersed long rter the smaller ones, reticulate-veined, with short, rigid, callous- pian hairs along the margins above, conspicuously venose and » 4-6 mm. long, 1.5—-1.8 mm, wide, sparsely pubescent and ciliate, = Be dager ge midrib and 2 or 3 visible lateral ’ glabrous but PT BR ciliate; calyx tubular, 2.5--3 m. apt glandular-punctate, hirsute on the lower half, the upper half, the lips membranous, a densely c ciliate, the teeth very short, irregular; orolla hypo— Srateriform, va from purple, pale-lilac, or vieiet to pink, rose, bluish-white, white, or even yellowish, 2-lipped, the tube ate , pubescent and glandular-punctate on the upper Portion outside, "the upper lip short, entire, the lower lip much 1 bed frui » Oboval » 1.8--2 mm. long, 1--1.2 h, inc by the fruiti ‘ into 2 cocci, convex on the back, flat on the vent ace, the co surf. celled, 1-seeded; usually only 1 seed produced, oboval, with en- This species was based by Grisebach on two collections made by Paul Gtnther amg (1) in thickets at Cueste de = caches cuman, and (2) frequent in the valley of Granadillas, Catamarca, Argentina, Troncoso (1952) says of the endosperm "por f menos Set aaa antes de la madurez una delgada capa de albumen rodea The species has been collected in fields, thickets, and dry o- 9 Places, on banks and verbanks, slopes, and mountainsides, aon Leyes and below m the sa a plateaus, at altitudes of 700 to Tornneters, flowering from October to May, fruiting in March. kart 9432 has cnpantitiy large leaves, The corollas are de- 196 PHYTOLOGIA Vol. 12, no. scribed as "purple" on Bartlett 19619, Meyer 3090, and Rodriguez 1281, "violet" on Venturi Ti 4290 & 7808, "pale-lilac® on Herb. Os~ 12200, "pink" on Job 1375, "rose® on Olea 93 and Venturi 2689 & & 9138, * "yellowish" on n Venturi 10193, "white" on Venturi uri 4296, and "white or inewyreniges te" on Fries 1. Possibly several “color pomities re represented here, but some of the variation is probably due to fading of the che with age or to differences in termin- ology on the part of the collectors. The L. lantanifolia var, crenata Griseb, is a synonym of L. junelliana (Moldenke) Troncoso. "Vernacular names recorded for the species are "cedrén del monte} "palo amarillo", "poleo", "salvia amarilla", and "salvia morada". sima (Gill. & Hook, ) Troncoso, while "poleo" is also applied to L. affinis Schau., L. alba (Mill.) N. B. Br., L. integrifolia (Griseb.) eerie L. turbinata Griseb. and its varieties, and Mentha Lippia we etacht ana is used as a blood-purifier in Argentina. Material has been misidentified and distributed in herbaria under the names Lantana balansae Briq., Cordia sp., and Verbena sp. On the other hand, ¢) the Ventur Venturi 815 distributed as L. grisebachiana is actually L. juneliiana ma (Moldenke) Troncoso. Te “Meyer s.n. [San oo de Colalao > y-1-1940) is a mixture with Lantana aristata var. Moldinke. Job 1375 was erroneously cited in Lilloa 5: 405 ee as Lantana balansae Briq. In all, 76 agp have seek k specimens and 2 mounted photographs Citations: ARGENTINA: Catamarca: H. H. Bartlett 19619 (Mi, W—- 1907566); Castellanos son. (Gracién, “Jan. 2, 19h03 Herb. Mus. Ar- gent. Cienc, — 33876) (N), sn. [Pomancillo, Jan. 11, 19105 Herb. Mus. Argent. Cienc. Nat. 33679] (N)$ Job 1375 (N) 5 Jorgen- sen 1025 [Herb. Osten 11423] (Ca~-192503, ee ig, W—704751), 8.0. 5 ote [Andalgal4, Feb. 1916; Herb. Fac. Cienc. Med. B. Aires 1025] 5) (N-- photo, Sp—25778, Z—photo); Luna Risso "656 (N); Peirano 6.n- (Londres, Jan. 22, 1934; Herb. Inst. Miguel Lillo 320h5] (N), aot n. [Los Boganes, April 2h, 1935; Herb. Inst. Miguel Lillo 3286 (N)5 Rodriguez Vaquero 902 (N, S); Schickendantz a poner Schreiter 10550 [Herb. Inst. Miguel Lillo 32058) (N). Cérdoba: Castellanos s.n. (Herb. Mus. Argent. Cienc. Nat. 31196) (N)- u- Juy: Claren 1179) (S); Venturi 9432 (W—-1591N19), 10493 (S)- ae Rioja: Castellanos s.n. (Herd. Mus. Argent. Cienc. Nat. 27/202h (N), sen. (Herb. Mus. Argent, Cienc. Nat. 28/325) (N); Venvure ee 7808 (Gg—-166712, S); Yepes s.n. (Herb. Mus, Argent. Cienc. Nat. 18855] (N). Salta: R. E. Fries 1 (N, 5); Garolera & mona Ste (Cuesta del Obispo, 21/1/17] (N), s.n. [Cuesta del Obispo, 23/1/ 47] (N)5 Lorentz & Hieronyms s.n. [San José, I1.73] (3)5 D. Rod- 1965 Moldenke, Monograph of Lippia 197 riguez 1281 (N, N), son. [Herb. Mus. Argent. Cienc. Nat. 23770] (N); Venturi 6929 (W--1591500). Tucumén: Baer 98 (S); Bailetti 33 Geeta tet Inst. Miguel Lillo 32399] (vi); Bruch s.n. [Valle del Taff, 08} (N, Ug); Castellanos s.n. (Herb. Mus, Mus, Argent. Cienc. Nat. 34815) (N); Krapovickas povickas 3240 (N (N); Lillo 430h (Herb. Inst. Miguel Lillo 32382] (au, N, W—18025)6), 7lL5 {Herb. Inst. Miguel Lillo 32393] (N); T. Meyer 3090 (N, N), 9766 (N, Vi), 11989 (N), sen. [San Pedro de | Calalao, l-~I-19),0] (N); | Vonetti 1625 [Hi (Herb. Inst, Miguel Lillo 32353] (nN), 1935 [Herb. Inst. Miguel Lillo 32388 (NM); Olea 93 (S); R. Rocha 3825 (Ba); D. Rodriguez ES [Herb. Inst, 1 Lillo 3239 91] (N), s.n. (Herb, Mus. Argent. Cienc. Nat. 23891] (N); Schreiter 372 [Herb. Inst. Miguel Eris 32622] (N), 969 [Herb. Osten 15087] (Ug), 2356 [Herb. Inst. Miguel Lillo 3262) (N, W--21,07899), 4920. [Herb. Inst. Miguel Lillo 3230] (N), 9732 [Herbd. — aes Lillo 32828] (N, eats a Gualinchay to Mal P » Dec. 1 as Herb. Osten 12200] (Ug); Venturi 2689 (Herb. Outen 17261} (N, Ug, W--133291, sat BS 1290 (ca—31.2555, N, N, W-1591437), W2s6 (W—-1343316), 9138 (W—- 1591418), s Son. (Herb. Mus. Argent. Cienc. Nat. 27/11) (N); E. Villa 661 @®. ar HASSLERTAMA Chod., Bull. Herb. Boiss., sér. 2, 2: 821. 902 Synonymy: Lippia hassleriana Briq. ex Moldenke, Suppl. List Invalid Names 5, in syn. 19]. Bibliography: Chod., Bull. Herb. Boiss., sér. 2, 2: 821. il Briq. in Chod., Pl. Hassler. 1 (9): 200. 1902; Briq. Ann. Conserv & - 7-8: 30 meer SL, & 263. 3 19h; Moldenke oa Uy: h2. 29185 H.W. Bee denke, Pl. Life 2: 63, slah op toe Alph. List Cit. 3: 693 (1949) and i: 1269. 19495 Maldenke, Known Geogr. paacah, yecben ces [ed. 2], 80, 9, & Eo, 1949; Moldenke, Résumé 93, 116, rat WN7, & h61. 49593 Trencoso, Darwiniana 12: 257, 262, & 263, pl. 1 fig. 2. 1961; Holienke , Phytologia 12: 24. 1 196, trations : Troncoso » Darwiniana 12: 262, pl. a, fg. 2. = shrub or eter ied, Rear _ Boiny stems woody, 2 7 mm. wi th spars leaves a decussate-opposite. fe runt "genie or subsessile, "hs ades coriaceous, broadly elliptic, 3.3—l.2 cm. long, 2.6--3 cm. ane often apiculate at the apex, sparsely serrate above © along the margins, shiny and rugose above, paler and scarcely 198 PHYTOLOGIA Vol, 12, no. or not at all rugose and setulose beneath, with 5 veins issuing from the base, 2 of which follow appro ximately the basal of the blade, the other 3 suberect but not extending a great dis- tance from the base; inflorescence distinctly paniculate, ample, pedunculate; bractlats coriaceous, very squamose, or ahyens on the back or glabrescent; calyx. 2-alate, glandular, the wi membranous, vesiculate, long-ciliate, but less so than ‘ L, hir- ta; corolla hypocrateriform, white or rosy-white. This species was based by Chodat on two collections made by was named NO. or 1899, and (2) no, 592k, collected in fields near the Rfo Capi- bary in December, 1900, both deposited in tg Delessert soepoe: at the Conservatoire et Jardin Mega sage: t Geneva, where latter was photogra abel by Macbride as his ics photograph beg 24653. Troncoso #11961) cae to the 592 collection as lecto- type and 5685 as syntype. The — 1902 and 190 references given in the bibliography above are often cited as "Plantae Hass~ leriana IX (1902) 200" and "xX (1904) 489", Chodat & Hassler (1902) say "Species affinis L. hirtae, dif- fert foliis subtus et superne minus nervosis, minus “plicatis P bracteis glabrescentibus pilis basi multo minus bulbosis, facie superiore foliorum laevi minus rugosa....A L. glabrescente Cham. et Schld. forma foliorum et eorum facie superiore nitida habitu- que sat distincta." The species has been collected in fields and shrubby campos, flowering from November to March, fruiting in December. The co- rolla is described as "white" on ” palansa 1030 and Hassler 592h, and as "rosy-white" on Hassler 5685. Br Briquet (190) says "Type de premier ordre, rapproché par M. Chodat des L. hirta Cham. et Le glabrescens Cham., Ce rapprochement nous paraft + fondé en ce sens que le L. Hassleriana appartient comme les deux espéces citées 4 la section Dipterocalyx 4 cause de son calice bicaréné. Mais il s'écarte d'ailleurs not tablement par l'ensemble de ses caracteres de toutes les autres espéces de la section Dipterocalyx et rapelle plutét nae igh Corymbosae de la section Busepania- of sp (1961) : "Especie descrita por Chodat basfndose en “Be tipos Gabeior 592 y 5685. Estudiados los agree oes del ‘aoe de Ginebra, aunque ambos representan exactamente la especie, ha seleccionado como lectotipo, Hassler 5924, por hallarse en 1a coleccién de fototipos de la serie del Museo de Chicago, No- sees hassleriana pertenece a la seccién Dipteroc oiliadas, ter considerado por Chamisso de gran valor taxonémico, pues le si rear el género Dipterocalyx por su autor. Segfn la ilustracién dada por Chamisso, las ae aparecen como un simple repliegue del eéliz por el contra trata de dos alas membranosas, vesiculosas.. Este c4liz vesiculos° 1965 Moldenke, Monograph of Lippia 199 es probablemente un medio de aur ya que las semillas su madurez permanecen encerradas por el mismo. Igual tipo de c4liz presenta L, hirta (Cham.) Sch. (Dipterosalyx hirtus hirtus Cham.) yl. sclerophylla Briq." She cites the following 20 spec: oe not as yet seen by me: PARAGUAY: Balansa 1030 (Cb, K, P); Fi rig S022 (Bm, Cb, K, Si); Hassler 5685 (Bm--cotype, Cb--cotype, ~ cotype, P-~cotype) » 392h (Cb--cotype, K--cotype, P—cotype), 8988 (mn), 9168 (Bn, Cb, K)s T. Rojas 4136 (Si); Sparre & Vervoorst 28 (Ma). In all, 17 herbarium specimens, including type material of h names involved, and 6 mounted photographs have been examin mae Citations: BRAZIL: Rio Grande do Sul: Bornmtfller 372 as N, N-~photo, Z--photo); Rambo 10004 (Rb). PARAGUAY: ARAGUAY: Balan: (Br, N, N), sen. (187h--7] (8); Fiebrig 6022 ee ae Hassler 5685 (Ca-—938078—~c ~-cotype, N--cotype), 5924 [Macbride photos 2653 (Ca—-935076—cotype, bE agate of ae wet no of cotype, N— cotype, N--photo of c otype, W--photo of co- type), 8988 (Ca--92900.. Py; “ Rojas "4136 136 (ede). PIA HEDERAEFOLIA Mart. & Schau. ex Schau. in A. IC., Prodr. 11: 593. “i Moon : Lippia hederifolia Mart. & Schau. ex Stapf, Ind. Lond, hi 12 125, 1930. Lippia hederaefolia Schau. ex Moldenke, Khom Geogr, Distrib. Verbenac., [ed. 1], 37 & 95. 192. Lippia hederaefolia Mart. ex MSR a ak Ft 10: 5, in syn. 196k. ae gg 4 Schau. in A. . ll: 593. 1847; Schau. * Mart., Fl, Bras. 93 250--351 & 307, re 41. 1851; Bocq., Adan- nia 3: 2hh. 1863; Jacks. in Hook. f. & Jacks., “ Kew. 2: 95. 18945 Briq. in Engl. & Prantl, Net. Pflansenfan (3a): 152. 1895; Hayek in Fedde, Repert. Nov. Sp. 2: 87. 1906; Glaz., Bull. Soc. Bot. France 58, Mém. 3: 5i2. 1911; cocker ee Estud. = Nordéste 3; 20, 19235 s gi Ind. Lond. kh: 125. 19 te Dep osoaes own Geogr. Distrib, Verben « (e603), 37 .. 4g = ok eed woldente, Alph. List cit, 2s “Ws (1918), 3s 689, Fig & 8 1135 & 1301. 1919; Woldenke, Know Geogr. Dis i. te to ted. 2], 80 & 190. 1919; Moldenke Revist. Sui. Bot. 8: 167. 7. 29905, Holdenke, Ré smé 93, 1. 19593 R vant. H - Minas 1 Mtn halaens, dir gree suppl. ar 2&h t G96) and 10: 5, 96h; “Woldenke, 1 Phytologia 12: 2h, 1h5, & 166. 1965. Illustrations: Schau. Mart., Fl. Bras. 9: pl. i. 1851. » margined, scabrous, foliose above; leaves ’ decussate-opposite, te short-petiolate; leaf-blades ovate or Subrotund, 6-12 mn, 4-10 mm, wide, only 1/) as — as internodes at the bess of the branc hes to equaling arther Up, acute at the apex, entire at the base, Se taier or 200 PHYTOLOGIA Vol. 12, no. subcuneate at the base and narrowed into the F apease ig incised- serrate and subrevolute along the margins with 2- -l, lobe-like the leaves, often e zt 1.6--2 cm, wide; bractlets large, showy, lilac or red, petaloid, Pel crate, ovate, 8--12 mm. long, acute or rounded at the apex, 5--T- veined, reticulate, pubescent, ciliate, finally slightly enlarged nee about 3 m ye hispid, the lobes acute, 2-dentate; co- large, showy, r e to violet or ng or hypocrateri fom, tabaes a on iesie eetaada: its tube about 8 m. long, e rate; stamens didynamous the hs Fc filaments short; anthers subdidynamous; style and normal for the genus; fruit unknown. This very distinct species was based by Schauer on several ections from Minas Gerais, Brazil. He a? (1847) ve campis deserts rok dictis prov. Minarum e (Mart.$ Pohl! th. h. bras. 196)" de aceite in the Senor" aoe DeCandolle her= Mrohat and (3822) bh ganic in campis deserti Serro rov. Min- dium: Martius; inter hehe et Vieira thier ~ i=} ec lippia. Briquet (1895) also classifies it in Subgenus Zapania, Section Rhodolippia. The J. E. Pohl 167, cited below, is probab- ly the Pohl collection referred to by Schauer -- it was photo~ graphed Ww Macbride in the herbarium of the Botanisches Museum Berlin as his type photograph no. 17512, but is now destroyed. The species has been found on apes ar and seeest fields, at 1000 meters altitude, flowering f. July, 1 98 in September and November. Teatesibure (2923) records “it fram the caa The flowers are described as "purple" on Williams & rasta 7050, "violet" on asa Magalh%es 1769 & 4,308, and "rose with a yellow throat" by Schauer. Material 1 has been misidentified set oie Sinigtes ted in herbaria as L. diamantinensis Glaz. and L Schau. Hayek see} ares Sat that his L. reticulata differs from Le raefolia in having the leaves reticulate-venose and pilose be- _ and the boshaabaracag hirsute. , 9 herbarium specimens and } mounted photographs, incla- ding | dine -ponctoee material of all the names involved, have been & y m Citations: BRAZIL: Minas Gerais: Heringer 5293 (Z); Marke were’ Brade, Mello Barreto, & Mendes Magalhfes bes 3273 } [Herbd. Jard. Bot. 1965 Moldenke, Monograph of Lippia 201 photo of cotype, N-~-photo of ¢ type, N—photo of cotype, W—photo of cotype), s.n. [Herb, Caes. Vindob. 187 comm.] (Br--cotype) ; Williams & Assis 7050 (G, N). State undetermined: G. Gardner 0). 512k (W—702lh0) ac ic sagtie Mart. ex Schau. in A, DC., Prodr., 11: 589--590, Bibliography: Schau. in A. DC., Prodr. 11: 589—590, 1847; Schau. in Mart., Fl. Bras. 9: 2hy——2),5. 1851; Jacks. in Hook. f. » 12-2 m, tall; root perennial; stems many, herbaceous, erect, obtusely tetragonal, sulcate along the ove, in Pwards; leaves decussate-opposite or approximate, ses- Sile, mostly deflexed, ovate~lanceolate, 5--6.5 cm. long, about 2.5 cm, Wide, gradually reduced in size upwards, acute at the a- longer appressed hairs, the floral leaves (bracts) lanceolate, the uppermost Ones smaller or even obsolete; panicle ample, cen- tripetal, na » 8p » Pyramidal or subfastigiate, 5--60 cm. long, the branches Slender or filiform, opposite, tetra, ; “preading, rigid, glandular-hirtellous, the sympodia 5~-7.5 cm. > ading, ragonal~filiform, glandu- larhirtellous; heads small, tetraquetrous-ovoid; bractlets herba- is Ovate, imbricate, acute at both ends, strigose-hirtous qu ng the corolla~tube: ¢ short, bifid, hirtous; co- rolla Small, rose-colored, about 1.5 m. long, its Purple, subinfun » ampliate above, curvate, puberu~ le lent above, the limb rose-colored, the lobes subemarg tellous, the lowest wider; fruit normal for the genus, rather large, obcordate, Separating into two halves. This curious Species was based by Schauer on several collections “ip Brazil: (1) by Carl Friedrich Philipp von Martius (no. 1705) w the west bank of the Rio Sao Francisco, Minas Gerais, and (2) by Johann Enamel Pohl also in wet places on the campo in Minas Gerais, deposited in the Munich and Vienna herbaria respectively. In his 1851 Publication Schauer cites also unnumbered collections by Pohl near Pillar in Gofas and from elsewhere in that state. Th Martius cotype in the Munich herbarium was photographed by Macbride 4s his type photograph no, 20328, The species has been collected in fields and wet places on cam- 202 PHYTOLOGIA Vol. 12, no. pos, flowering in April and August. In all, ) herbarium opecinete and mounted “photographs, including phototype material, been e ed by m Citations: BRAZIL! Minas Gerais: Macedo 221) (N, S), 2299 (N), 2h (W--2196854); Martius 1705 [Macbride photos 20328] (It-=photo of ¢ hag: 44 Kr-~photo © of co cotype, N--photo of cotype, W--photo of cot; LIPPIA HICKENII Troncoso, Darwiniana 10: 69--72, fig. 1. 1952. a eens, Darwiniana 10: 69--72, figs ls 1952; G. Tay Ind. Kew, Suppl. 12: 82. 1999; Moldenke Résumé Suppl. 7: 2 ehwis Vol denke, Phytologia 12: 18) &1 70. 1965. mei a Savesshann ¢°70, Lig s 15 195Se 7 shrub; branches mane woody, subcylindric, striate, with ena caper te this year's branchlets straight, subtet- ragonal, 8 sparsely glandular—puberulent, the oerecentd formed of era hairs with resinous glands visible only under a hand-lens; rincipal internodes to 3 cm. long; leaves Serussatexoppolies) shaeuoranintahes petioles 2.5--5 mm. long, ae leaf-blades somewhat bicolored, oval or oblong, 1--2 O.l--1.2 cm. wide, subacute at the apex, truncate an a broadly " gub- and re cuneate at the base, finely and regularly crenate along the mar- gins except at the entire base, trinerved or pennin os especially along the marg inely reticulate, densely glandular-punctate and short-pubescent above, opaque and incanous~ pubescent beneath especially on young leaves, glandular-punctate between the pubescence; venation impressed above, prominent be- neath; inflorescence axillary, on this year's branchlets, fascicu- late; peduncles short, h—-9 mm. long; heads dnb per axil, oval, 5--6 mm. long, hening to 10 mm. during anthesis, 10--l2- flowered; bractlets membranous, stramineous, decussate in 5 or 6 3 y smooth, glabrous, he the style persistent on its apex, about 1.5 mm. ae (without th ‘ style) and 0.8 mm. wide, with a marked long h l-celled, the asset subcoriaceous, dry, thin, smooth and shiny within; immature seeds ellipsoid, white, 1-2 mm. long, basifixt The type of this species was collected by Lorenzo Raimundo P 1965 Moldenke, Monograph of Lippia 203 di (no. 14864) at an altitude of 1100 meters at Diquelos Sauces, Ta Rioja, Argentina, on February 16, 194), and is deposited in the he uto on son, chica ues Guasayan, Villa La Punta, Santiago del Estero, deposi ted as sheet no, 47556 in the herbarium at Buenos Aires. She says: "El ej erencias con as vigorosas con ho Ojas mds grandes, hasta de ,3 cm ms long. x 2,7 cm, de lat. (sin el peciolo), éstas son de superficie menos rugosa; hojas la pubescencia sericea, p L e en ° e ores; las br&cteas son m4s subuladas y plegadas en el &pice, has- ta de 7,8 mm de long. No sponi de ha re- c gién queda para m4s adelante valorar estas pequefias diferencias." She iso 6a comments that "L, Hickenii pertenece a la Secc. Gonio- Stachyum Schauer.....caractenzada por presentar dos o varios ana, Lippia dumetorum Herzog. La comparacién de — at Esta diferencia fundamental la he podido comprobar por én gracias a la gentileza del Dr. F. C. Hoehne, da So fal, = me enviéd material de L. sidoides determinado por el ua Mol- enke y procedente de su 4rea general tf{pica (L. sidoides C il, Sao Paulo, Iti, A. Russell No. 82, X-1597; S80 Pau alo, So- Tocaba, A. Lofgren No. "28, X-1887; du dupl. SI.)" LIPPIA HIERACIFOLIA Chan. +, Linnaea 7: 375. Onymy : ods 'C;: 2¢ 54. 18]0, Lippia hieraciifolia Cham, ex Schau. inA DC., Prodr, 11: 588, 1847. Lippia alegrensis Briq. ex Augusto, Fl. Rio Grande do = 235. 196. Lippia og Rete Briq. Moldenk nke, Alph. List Invalid Names Suppl. 1: syn. ihT. pie Matastsruiss Cham. & Schl. ex Moldenke, ‘ptuiet Suppl. 1: "nib Sogr ted 5. 1832; Steud Nom. Bot., eraphy: » Linnaea 7: 375. ; 3, se i 2, 2: 5h. 1840; 5 Malp., Repert. Bot. Syst. h: 55. j Schau. A. IC., Prodr, 11; 588-589, 1847; Schau. in Mart., ee 20h, PHYTOLOGIA Vol. 12, no. 9: 2h2—-243. 18513; Jacks. in Hook. f , Ind. Kew. 2: ay 189; Bri a Arkiv Bot, Stockh. 2 (19): 20. x156h3 Herter Urug, 105. 4930; Herter, Revist. Sudam. Bot. h: 185. 1937; Mol- ” denke, Lilloa ae “i Paeyse saps, gate List Invalid Nam 5. 1941; Moldenke ex trib, Verbenac., [ed. 1], 31, ee pes & 95. 95. 9hg5 alohaeakes - List Invalid Names i. i923 enke, Lilloa 1 noble Sohleeks Liphe List Cit. 1: 12, es 192, & 195. 3916 co i » Fl. Rio Grande do Sul gio ie Molde enke, sctet ae “Davalia Wan ames Suppl. 1: 13 & 1h. 1947; denke, Lilloa 3. 1948; Moldenke, Alph. List Cit. 3. 366 Li2, & 47 caol8), 31 7 ees, 689, 691, 703, 7h5, Bié, 810, Bh8, 8 Bel, 921 (19h9), eis 1257. 19h9 Voldenke, Known Geogr. Di ous suffruticose perennial; s mga = he base, @& rect or arg mostly simple, slender, rte Site 0.5--1 m. , hirsute, leafy below, almost leafless and scapose above an almost tuberous caudex, savas with some axillary e apex; principal internodes few, greatly elongated; leaves few, decussate-opposite, ype the blades obovate-oblong or lanceo- late, 2.5—-12.5 cm. long, 1.3--3.2 cm. wide, acute at the ag ous oz smaller; peduncles axillary at the upper nodes, soli- tary, 1.3 ca. “, the upper ones shorter; heads globose, about wide, few, racemose or congested at the apex of the stem; rect lets’ ovate, almost 3 mm, long, acuminate at the apex, strigose- canescent on the back, closely fatirteate! equaling the corolla; ost 6 c raterifom, its tube straight, slightly ampliate a ly puberulent in the throat, the limb very short, epee the lobes subequal, rounded. This species was based by Chamisso on several collections made by Friedrich Sellow (nos. 1698 & 3260) in southern Braz and deposited sae the herbarium of the Botanisches Museum in Ber lin, e photographed by Macbride as his type est cae mumber 1751h,, but are now destroyed. The type of Le al was collected by Eduard Martin Reineck and Josef Sree Gon at Porto Alegre, Rio Grande do Sul, Brazil, and is de~ a in the Delessert Herbarium at the Conservatoire et J es eva; an isotype in the Munich herbarium was photographed by nang as his type photograph number 20320. appears as "Lippia alegrensis Briq. sp. nov.” 0M the printed labels. Augusto (1 sto (1916) says "segundo Borsmael} 6 encontra-se a mesma espécie [L. hieracifolia] no herbario 1965 Moldenke, Monograph of Lippia 205 Reineck e Czermatk sob Li Lippia alegrensis Briq.....Rio Grande do Sul, campos arbustivos 'forma proxima a Verbena: littoralis e Dotlkrtensagt confere com Lippia hieracifolia.” Schauer (1847) places the species in Section Zapania, Subsec- a Paniculatas. It am been found growing in bushy or shrubby ds, dry or o Open campos, 8Sy er’ in general, at an altitude of 200 meters, flowering from November to F February and in May. Rambo encountered it in a region of 1.5 326 distributed as L. hieracifolia is actually L. angustifolia Cham. The Leite 245 collection cited below is marked "Bugenio 245" on the original label, but was collected by Padre José Eugen- Leite, — (1946) cites an