PHY TOLOGIA Designed to expedite botanical publication Vol. 9 February, 1963 No. 1 Miss ai MAR 4= 1463 CONTENTS CUATRECASAS, J., Notes on neotropical Compositae, 1 ....... 1 MOLDENKE, H. N., Notes on new and noteworthy plants. XXXIV 7 MOLDENKE, H. N., Materials toward a monograph of the genus Verbena. IX ee ee ee ee DEGENER, O. & I., Arachniodes, a fern genus unrecognized from MEE sn 5 see eoeeee ere eee es ee see ees ewe ee eee eseeee eee ere eee ee eee eh hh Published by Harold N. Moldenke and Alma L. Mokieske 15 Glenbrook Avenue ae = 2 Yonkers 55 New oo Us s. AL — : Price of this number, $5 per vol NOTES ON NEOTROPICAL COMPOSITAE, I José Cuatrecasas The following descriptions, nomenclatural transfers, — mic concepts and other comments refer to the Astereae, especial Flora Colombia in preparation, The genus — za is consider- ed here as defined by Cronquist in Brittonia 632 1943. The work upon which this opty is based has ak sponsored by the National Science Founda Key to the Colombian species of Conyza: 1. Flores feminei capillari-tubulosi. 2. Pappus duplex, exterior minutis pilis coronam formantibus. Plantae lanatae vel lanuginosae. Sec. Laennecia. 3. Capitula angusta subturbinata )-5 mm alta 8-9 mm dian~ etentia (expansa), involucro ) mm alto bracteis lineari-lanceolatis pallide viridibus vel brevissime violaceis, floribus femineis 2-6. Achaenia dense Villoso - hispidula, Pappus pilis exterioribus con- formibus 0.8-1 mm longis. C. filaginoides (DC.) Hier. 3. Capitula — 5-6.5 mm alta 11-15 mm diameten- tia (e » involucro h-5 mm altum bracteis late f Tasco siabta. sursum um purpuraceis, floribus femineis cir- hisp xte minutissimis saepe dilatato-squamulosis 0,2-0.3 longis. Ce gnaphalioices 1K. 2. Pappus simplex, pilis uniseriatis. Plantae haud lanatae. Sec. Conyza 5of mn a aribus. Capit parva 5 mance Yay Fn mm alta s 75-110 floribus genie corolla ¥ lin- bo imfundibuliformi. C. sophiaefolia HBK. (3<12 capitulis). Folia repando dentato-crenata vel oi basilaria 5-20 cm longa. Cap 6<8 mm alta 15-18 mm lata (expansa) semigloboso- 1 2 PHYTOLOGIA Vol. 9, Bie campanulata circa 380 floribus femineis, corolla g Limbo prerpie tubuloso, C. chilensis Spreng. 5. Herba annua stricta ta parce hispidula. Folia serrata vel integra eee 1-6 cm longa. Panicula corymbosa mediocris po brevis. Capitula minuta semigloboso-campanul 225-3 mm alta circa 100 floribus femineis. Scots % limbo infundibuli- formi, C. microcephala Hemsl. 1. Flores feminei minute ligulati. Sec. Coenotus. 6. Herbe suffruticosa ee feliis pinnatipartitis segmen- tis linearibus, itulum semiglobosun a 5 mm altum 110-160 floribus heise, Corolla § tubo stricte tub- uloso. Ovarium dilute adpresse i eo « cardaminifolia HBK. 6. marke annua vel perennis foliis integris dentatis vel lobatis, 7» Herba plus minusve lanata pilis longis tenuibus flexu- lanatae. osis intricatis. Proles a C. prolialba Cuatr. 7+ Herba hispida hirsuta vel villosa. 8. Herba annua erecta stipitato-glandulosa et hirsuta. Capitula 6 mm alta 170-300 floribus femineis. C. trihecatactis (Blake) Cuatr. 8. Herba haud glandul ° 9. Herba basi suffrutescens g O- hirsuta). Capitula 5-7 mm alta 10- mate (expansa). Ce uliginosa (Benth.) Sante, 9. Herba annua simplex. 10. Herba plerumque flexuosa vel erecta pil fo. ula circa mm alta 7-9 mn oa (expansa) « Ovarium dilute by ei llos Ce spent Ne HBK. 10. Herba erecta plerumque robusta foliis lance- olatis vel lineari-lanceolatis vel linearibus, sursum ramis fastigiatis thyrsoideo-flori- 1963 Cuatrecasas, Neotropical Compositae 3 Ms pcos ar mm alta 7-3 mm diametro bracteis viridi-pal prominula glandulosa. Corolla ¥% limbo tubuloso. Ovarium glabrum vel subgla- brum. C. canadensis L. li. Capitula io mm alta, 10-15 mm diametro (expansa) 60-120 floribus femineis, In- volucrum seed te hirsutum vel villoso- hirsutum. Corolla ¥ for cies tubuloso- uliformi., Ovarium e villo-=- sum. C. bonariensis (Le) Cr aren CONYZA PROLIALBA Cuatr., sp. nove Herba annua vel perennans radice fusiformi ramulosa; planta vernalis vel juvenilis parva pulvinata 2=5 cm alta basi ramosa, ramis numerosis congestis densissime foliosis, foliis dense crasseque indumento albo lanato vestitis apicibus ramulorum sterilium floccoso-lanatis; adulta ad 60 cm alta caule simplex vel basi ramoso, caulibus ramisque erectis teretibus vel levi- ter striatis sparse glanduloso-pilosis et plus minusve lanugi- nosis pilis longissimis tenuibus flexuosis intricatis. basi in longum petiolum planum nervatum subscariosum producta lobis h-5 obtusiusculis circa 2 mm longis sienrtined » untringue albo- lanata pilis stipitato-glandulosis brevibus ot pilis longis inosis dense tecta,5-12 mm longa -10 mm lata,petiolo 8-20 mm longo circa 1.5 mm lato basi ampliato subamplectenti. Folia caulina i onga ‘on & erispoque, nervo medio subtus prominenti, nervulis later- = parcis ascendentibus venulisque laxis obsoletis, 1.5-5 em longa 0.5=1 cm lata. Capitula 1-5 breviter pedicellata vel subsessilia in axillis foliorum glomerata, glomerulis secundum formantes; folia (bracteae) subtendentia viridia ovato- neti auriculato-amplectent tia integra margine recurvata costa subtus ed garsat ig glomerulos excedentia vel aequilonga Pedicelli teneri 0.5-l; mm longi tenuiter lanuginosi, bracteoits 4 PHYTOLOGIA Vol. 9, no. 1 subtendentibus linearibus 5-8 mm longis 1-1.5 mm latis. Capitula circa 5 mm alta semigloboso-campanulata circa 6=7 mm lata (expansa 11-12 mn). he fade pallide viride vel pur- purascens ).5-5 mm altum, bracteis 3-seriatis lance-linearibus acuminatis dorso dilute stipitati~-glandulosis et longe flexo- is mar late osis eroso-ciliatisque, interioribus h-l.5=5 mm longis 0.6-0.8 mm latis co apes 325 mm ag Flores radii feminei ligulati 200-260; c minute li gram vel minutissime emarginatam 0. ongam formanti. Rami pe tc stricti lineares circa 0.7-0.8 on po cf exce=- Flores centrales hermaphroditi 10-20; corolla — circa a mm longa tubulo angusto sursum minute pilosulo 2 l= 2.8 mm longo limbo campanulato dentibus oblongis iccoouits 0.7-0.8 mm longis sparsissimis pilis aac igi Antherae bere O.9 mm longae basi rotundatae a gm ys culatae. Rami styli robusti 0.7 mm longi aioe apice pateinaians ex= tus dimidia superiors go arpoogplarmrs ie Ovaria e lana anagem albi- ilibus Pappus 3 mm dus pilis te us strigulosis wmiseriatis, deciduis 8. Receptaculum 2=2.5 mm diamitens alveolatun marginibus alveo- lorum dentatis, Type: Colombia, Magdalena: Sierra Nevada de Santa Marta, ha hig rdsscending 20 a6 uthwestern from Picos Reina and Ojeda: Owy paramos around Laguna Naboba, Laguna Mamo, 4200-1300 malt. Adult leaves light green or lanuginous whitish; involucre ae —— or the tips more or less dark; rack plants densely whi duly, October 4, 1959, coll. J. ph casas & R. Romero ou 24595 (Holotype, US; Iso Other specimens examined; Sierra Nevada de Sante erate wee hg a Hoya del Rio Donachui: Meollaca ky pdéramos about 3400 m alt involucre pod green. October 8, 1959, coll. J. Cuatrecasas & Re Romero Castafieda 2); Santa 3; involucre very li e coming purplish, ein yellow. Cua casas ° ean wets 195 1t.e Pale green herb; Corollas te. November 9, coll. J. Cuatrecasas & R. Romero Castafieda 25260 rik : Cc ers lialba is ere related to C, mima Blake from Venezue by ha always ligulate ray flowers and glabrous achenes, and by being a larger plant. From the 1963 Cuatrecasas, Neotropical Compositae Central American Erigeron subspicatus Benth., the new species differs by its glabrous achene se S$ and more abundant yee both representing probably only vica od nth varieties of the same species; inlet the Bentham epithet is already used for an- other species of Conyza. CONYZA PROLIAIBA var. SUBSPICATA (Benth.) Quatr., comb. nov. Erigeron subspicatus Benth. in Oerst. Kjoeb. Vidensk. Medde 1852 CONYZA i ay cry (Blake) Cuatr., comb. e a Sas m trihecatactis Blake, Journ. fabs ‘ail Sci. 273 Os e CONYZA paar (Benth.) Cuatr., comb. Erigeron uliginosus Benth., Pl. ate 20h. 185. Erigeron popayanensis Hieron., Bot. Jahrb. Engl. 28:586. Eri eron sulcatus var. columbianus Hieron., Bot. Jahrb. Engl. Eri on "wi osus var. hirsutus Hieron., Bote Jahrbe Engl. 203 Erigeron "ponariensis mat meridensis Cuatr., Trab. Mus. Nac. Cienc, ate fard. Bot. Madrid, Ser. ae 333132. 1936. CONYZA preosagpeens var. COLUMBIANA (Hieron.) Cuatr., comb. nov. BBB i var. columbiamus Hieron., Bot. Jahrb. 1901 Le Folia axthine anguste linearia, 1-2 mm lata. Inflorescent- iae plerumque laxiores pedicellis gracilioribus. CONYZA ULIGINOSA var. HIRSUTA (Hieron.) Cuatre comb. nove . Erigeron uiiginosus var. hirsutus Hieron., Bot. Jahrb. Engl. 28: Caules folia inflorescentiaeque densiuscule longeque albido- CONYZA BONARIENSIS var. LEIOTHECA (Blake) Cuatr.,comb. nove Erigeron bonariensis var. leiothecus Blake Contr. Gray Herb. 52: zs Conyza Sic ibaile HBK, Nov. Gen. Sp. Pl. h:73- 1820. ERIGERON CHIONOPHILUS vare LEPTOPHYLLUS Cuatre, var. noVe caespitosa rhizomate verticale vel obliquo ramoso ramis erectis cum vaginis foliorum vetustorum tectis denique nudis apice folia rosulato-fasciculata congeste ferentibus. Folia anguste lanceolato-linearia vel linearia xg acuta basim versus sine sensu longeque attenuata supra basim in vagi- © paulo conspicuo undique pilis rigidis patulis vel subpatulis longis acutis basi incrassatis copiosis tecta, 6 PHYTOLOGIA Vol. 9, now 1 Capitulum 1.5 cm diamitens campanulatum. Involucrum circa 1.3 cm altum bracteis herbaceis viridibus parorree re subaequi- longis lineari-subulatis acuminatis interi us 1] mm longis ad basim 1-1.2 mm latis margine scariosis pet year Aaa dorso villosis pilis longis subpatulis flexuosis, exterioribus 8 mm longa tubulo capillari 3 mm longi glabro vel subglabro neari apice minute 2-3 dentata vel integra O.l mm lata, Pp 5 pilis inaequilongis basi coalitis strigosis n'a ys ex- terioribus vel subexterioribus minutis (0.1-0.3 mm longis) squamulosis, ovario oblongo sublaeve glabro vel praecipue sursum sparsis pilis 2 mm longo. Flores disci hermaphroditi circa 50, corolla .5-5 mm longa parce pilosa parte media tubulo 2-3 mm longo limbo tubuloso-campanulato dentibus 0.8-0.9 mm longis tri- longis acutiusculis, ant 8 longis basi minute auricul kgs 0.8 mm longis oblongis subcom- pressis obtusiusculis extus sursum papilloso-pilosis deorsum gradatim minus papillosis a papilloso-stigmatico, ovario 1 m longo glabro vel parcissimis pilis, pappo cum in femineis. Varietas a specie folia anguste linearia bracteae angusti- ores differt. Type: poles e Sertanars popes montanhosas a cue mn alt., a 2 km del paraje Berlin; flores blancas; suelo sec tthe ee aoe Araque & Speier 18S),09 faiotees: ERIGERON WEDDELLIANUS Hieron. ex Sodiro, Bot. Jahrb. Engl. 19: 89. 189k; 29:21. 1900. oz Erigeron spathulatum Wedd., Chl. And. 1:197. 1857. Not E. spathulatus V. : p ron ecuadoriensis Hieron., Bot. Jahrb. Engl. 21:336. 2 Er eron wee var. rosuloides Cuatr., An. Univ. rep Eri ae. Caldasii Caceres ? “trab. Mus. Cienc. Nat. Jard. Bot. Ser. Bot. 29:23, dhact 6. 1935. In *189h Hieronymus eribed very briefly and imperfectly sti on a tions of esgeaees llitus G well known Erigeron spathulatum Weddell (not Vahl). ORITROPHIUM HIRTOPILOSUS (Hieron) Cuatr., comb. nov. Erigeron hirtopilosus Hieron., Bot. Jahrb. Engl. 213336. 1095.6 Types Stuebel 48 i. Photo F.M. 18h). es Same Mathews, 1862, "Peruvia" (US. 245951). This specimen 1963 Cuatrecasas, Neotropical Compositae 7 perfectly with the description and photograph of type. BACCHARIS INCARUM (Wedd.) Cuatr., stat Baccharis microphylla & incarum incarum Wedd. ie And. 1:170. e type (P) was collected by Weddell in the verve’ de Tacora, Peru. Para s came from Potosi, La Paz, Titicaca near Tioguanaco (collect. Weddell, Orbigny , Pentland). The Peruvian specimens: Macbride & Featherstone 47 from Matucana and Ma aso & Featherstone 2502 from Pomopampa, dis- tributed as B. intermedia DC. atthe sty to B. incarum, which is very different i B. inte NOTES ON NEW AND NOTEWORTHY PLANTS. XXXIV Harold N. Moldenke a MACROPODA f. ALBIFLORA Moldenke, f. nov. ec forma a forma typica speciei corollis albis recedit This form differs from the typical form of the pio in h e * The of the form was collected by Cyrus Longworth Lun- dell fade 14517) in the Yeager Pasture of the South Texas Syn- dicate Ra Ranch, in the vicinity of Experimental Garden, La Salle County, Texas, on aeteber 5, 1945, and is deposited in the Lundell Herbarium at the Texas Research Foundation, Renner, Texas LIPPIA INOPINATA Moldenke, sp. tex; ramulis gracilibus eateagonts subadpresso-villosis, pili seis; foliis s subcoriaceis ovato- ellipticis saepe falcato-conduplicatis ad apic inque acutis, ma sie entis “wisinped —_— e villosis Wo shrub; stems tall, woody, gray, glabrous, subterete; branchlets slender, brownish, obtusely tetrag onal, ons a es subappressed antrorse grayish hairs; principal internodes abbreviated, 8--21 mm. long; leaves " decussate-opposite; petioles us with long 8 PHYTOLOGIA Vol. 9, no. 1 tion more or less subimpressed above and slightly prominulous be- neath; inflorescence axillary, crowded near the apex of the twigs, qual ° : ense spreading-villous; heads subglobose, yellowish, about 1.5 cm. long and wide, densely many-flowered; bracts papery, subobtuse at "the apex, corey pubescent on both surfaces. this 1 gaan was collected by Howard Scott Gentry (no. 18267) i an Opuntia nicket on a rocky slope in cactus- acacia association, altit riettade 6200 feet, Villa Nueva, — Mexico, on November 29 1959, and is deposited in the C. L. Lun- dell Herbarium a + the texas Research Foundation, Renner, Texas. MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. IX Harold N. Moldenke VERBENA CATHARINAE Moldenke TariAcanebane rather few-flowered; peduncles very slender, a- bout 2 cm. long, sparsely hirsute; bractlets lanceolate, a corolla blue, hypocrateriform, its tube about 11 mm. long, puber- ulous on the outside above the calyx, its lobes also pilosulous- puberulous beneath. The type of this species was collected by my good friend, ei Raulino ages (no. C.1561) on a campo at —— * alti- e 900 meters, Ararangud, Santa Catarina, Brazil, on February is, 19K6, aes is deposited in the Barbosa Rodrigues bets wa at Itajai, Brazil. The species is said to be a ruderal one and has been found on campos and rock outcrops, growing at from 760 to bruary ber. Vernacular names ee for it are adinha", "formosa "Ju h "camar. a whic _ apparently given to many other species of ret genus in Braz In all, 10 ‘haglanaes specimens, a otidine the type, have been by m Citations: * BRAZIL: Paran4: Hatschbach 3363 (Z). Santa Catarina Reitz C.1561 (N--isotype, Rd—2121—type); Smith & Klein 8055 (Ok, W—2251336), 11403 (Ok, W—2251777); Smith & Reitz 8917 (Ok, W-- 2251467); Smith, Reitz, & Sufridini 9561 (W—22h9363) ._ VERBENA pecan si ceee, Phytologia 5: 228-229. 1955. ography: , Phytologia 5S: 228--229. 1955; G. Taylor, Ind. Kew. Suppl. Ie: ET "19593 Sires Résumé 9k & hos: 19593 Moldenke, Résumé Suppl. 1: 7 & 25. 1963 Moldenke, Monograph of Verbena 9 Herb, O—5O em. tall; stems simple or branching in the base, tetragonal, deeply sulcate between the angles, aad sh les. leaves decussate-opposite, sessile, elliptic or narrow-elliptic, 6--7 cm. long when mature and 1.5--2.5 cm. wide, the upper ones much narrower and oblong, often recurved, acute at the apex, cuneate-acuminate to the amplexicaul base, sharply and irregular ly serrate from above the cuneation to the apex, scabrous and ob- scurely cia taps above, more densely short~pubescent with glanduliferous hairs eath; midrib slender, impressed above, very prominent ants : deciatoel slender, about ) per side, ascending, reuate, ng in the teeth, impressed above, undant ou re ry prominent beneath; escence terminal. spicate, usually with one shorter side-branch, densely many-flowered; peduncles similar to the stems in texture mm. 1 calyx, pilose on the back ek ers es calyx gone about mm coring The type of this species was collected by Teodoro Rojas {Hassler 259] at Lomal Clavel, latitude S.23°20', in the Gran Chaco, Paraguay, in November, 1903, and is deposited in the herb- arium of the University of California at Berkeley. The species is known from only the type reap aa Only 2 specimens, including the type, have been exam tations: PARAGUAY: T. Rojas 8 s.n. {Hassler 2459] (Ca—929435- type, Z—isotype). VERBENA CHEITMANIANA Moldenke, Phytologia 2: 420--21. 1918. paiognnia made Phytologia 2: 420--,21. A Moldenke, astanea 1 1949; idee, Known “Geogr. Distrib. Verbenac., [ed. 2 ms oe & 197. et Moldenke, Phytologia 3: 135. 1949; E. J. Salisb Kew, 1. li: 262. 1953; Moldenke, Phytologia 5: 2h. 195); aa denice, Shened 127 & 471. 1959; Moldenke, Résumé Suppl. 3: 1 962 Sprawling herb; stems often procumbent, very slender, elon- gate, acutely tetragonal, strigillose or glabrese cent; branches rather few, very slender, elongate, sprawling, cutely tetragonal, often slightly sulcate between the angles, cn densely strig- illose with very short and whitish reflexed ogy nodes es tee be apparently often numerous only poe dare the tips og ai ranches; petioles slender, 1-—-1.5 cm. long, margined, densely 10 SHEET OLO6GIs Vol. 9, no. 1 pilose-pubescent with subappressed, whitish, antrorse hairs; leaf-blades membranous, somewhat lighter green beneath, not brun- nescent in triangular-ovate in outline, 1.5--) cm. op 1--3 te o an 9 mm, long, densely strigose with closely appressed, antrorse, white hairs, its rim unequally gan ye the teeth long-aristate, their tips connivent before and after anthesis, purplish; corol- ery villous at throat within, the limb 7--8 mm, wide, patent, the lobes deeply cordate at the apex, glabrous on both’ surfac The type of this most interesting sas toon ae collected by Robert E. Fries (no. 477) in shady places under Salix humboldti- ana at Piquete, on nm the sandy sabes of the Rio San Francisco, Jujuy, Argentina, on August 21, 1901, and is deposited in — herbazium | of the wie ppt ska Rikemuseum at Stockholm. species one in the general appearance of its enoes of the North American V. canadensis (L.) Britton. It is in honor of Philip Cheitman, American ca nd ardent oad of Nat o has rendered considerable help to me in searches on tropical and extra-t American plants. Pp. The species has been found on hillsides as well as on sandy riverbanks, at altitudes of 50 to 3300 meter rs, es in February, dpril, and November, The Formosa, Argen ecord given by me in my Known Geogr, Distrib. Verbenac., py 2}, 105 (1949) and Résumé 127 (1959) was based on Morel 2003 and 1214, but these two collections are now r regarded by me as representing V. calliantha Briq. The vorgensen 86 cited res po} be from the provinces of Buenos A Cérdoba, Entre Rio os, San Luis, or Santa Fé, since there is : "esaladi lion locality in each of these provinces. In all, 11 herbarium saya including the type, and 5 mounted photo graphs have been me. Citations: ARGENTINA: Cabineres: Brucher & Brucher s.n. [16/ 2/1949] (Lu). Jujuy: R. E. Fries 77 (F--ph Cooshctc of tee of type, N—--iso- 1963 Moldenke, Monograph of Verbena 1l type, N—photo of type, Pc--photo of type, S--type, S— S--isotype, Si--photo of type, Z——photo of type); wiierone “Ek [Herb. Osten 20012] (N, Ug); Venturi 5132 (W--1591)62). Salta: Venturi 5329 (W—1591L69). Tucuman: Venturi 7785 (W-~1591389). Province undetermined: Jorgensen 86 [El [El Saladillo] (W--921685). VERBENA CHILENSIS Moidenke, Known Geogr. Distrib. Verbenac., [ed. 1), bb, 78, & 101, hyponym (March 12, 1942); Lilloa 8: 29. ate ynonymy: Verbena glabrata R. A. P Anal. Univ. Chil. 35: 190-291 {Sert. Mendoc. Alt. 32]. hee hk v. eiabrate H.B.K., 818]. Univ. Chil. Bs “pen Ag (sert. Mendoc, Ait. 1080, ‘i272, & 1198. 1949; Moldenke, 4 ra oe Distrib. Verben- ac., Ted. 21, 105 & 197. a Noidenke, Phytologia : 71. 1952; E. J. Salisb., Ind. Kew. Suppl. 11: 262. 1953; Moldenke, Inform sen Set 51 Spec . h. 1956; Diaeakh. tee 127, 36h, hg, & 71. Semiprostrate shrub or suffruticose, completely glabrous throughout; branches 23--30 cm. long; upper internodes about twice as long as the leaves; leaves up to 26 mm, long and 15 m. into 3 small lobes; spikes exserted, capitate, many-flowered; bractlets ger eee subulate, shorter than the calyx; calyx 9-- 10 mm, long, rous; corolla yellow or yellow and purple or pale-purple, 51-23 ma long, glabrous; anther—appendages clav- ate, exse The type of this species was collected Poi Rudolf Amandus Philippi in Mendoza, Argentina, sometime in 1868 and is deposit- ed in the herbarium. of the Museo Nacional de Historia Natural de Chile in Santiago. Unfortunately, the label of Macbride's widely distributed photograph 1717 of an isotype does not show Philippi's original label and is erroneously inscribed "Chile" -- whieh is the basis for the misleading specific epithet which I chose for the species when it was discovered that P ree pak pa original epi- hom The Gillies s.n. [Mendoza] specimen cited the limb medium-sized, deeply 5-fid; fruiting-calyx oa, twice as long as the fruit; cocci Nethedlavestihenodts te Ve. This species has been collected in thickets, along roadsides, in sandy pl » in deep soil of canyons, and among shrubby lack soil, at altitudes of 3200 to 3600 1 rseaaie growth in fertile flowering in Jan » March to June, September, and etn and fruit: in - Soukup says that it is locally co J » Peru. Gay (189) says menet it occurs "en a Lea iecil de Chile entre Santiago y Mendoza". Common names recorded for it are "cara huanzuy", nies serbanae: "verveine Sanchoouae: and "wedge-leaved vervain" Walpers (185) classifies it in his Section Leeuenes rd section Inermes, Group Foliosae, pages Macranthae, ondary Subgroup fpr with ten other heuer ened places it in et gh Verbenaca, a ei te Pachystachyae, with 21 other speci Schauer iene Dombey 261 7H = & Pavon SM. from Hu4nuco lin and De ers sew ie amba, Per Peru, int er the latter being the type collection of the species. H (1941) cites his no, 1,09 from Cuzco, Peru. Raimondi (193) cites his no, 10255 from se veliea, 1535 & 12032 from Ancash, 2558 from Jw Junin, ai and 8929 from Department ent undetermined, Peru It is aoukle noting that the V. cuneifolia of bakicceine is V. stricta Vent., that of Wallberg i is V. simplex Lehn., and that of Hort. is V. hispida Rufz & Pav The H. H H. "Rusby 911, dapbeibated as V. cuneifolia, is the type collection of Ve V. bangiana Moldenke; 3 Cook & Gilbert 99 is V. hayekii Moldenke; F. i We Pennell 14582 and “Wilkes SMe “Gane [Obrajil- lo] are V. hispida Rufz & Pav.; and Cook & Gilbert + 373 is V. parvula Hayek. In all, 12 herbarium specimens and 2 mounted been examined by me. Citations: PERU: Ayacucho: Killip & Smith 2329) (N, W—-1358- 017). Junfn: Killip & Smith 222k (N, “W—-1357172); Kunkel 383 (Gg); Macbride & Featherstone e 1210 (W—1h71583) ; Soukup mkup 3537 photographs have 52 PHYTOLOGIA Vol. 9, no, 1 Se jared N, W--1930168); Stork 10920 (Ca-—655732). La Liber- : A. Lopez Miranda 310 (N, “Z). Department undetermined: Dombey Son. [Macbride photos 17411] (Kr--photo, N--photo). VERBENA CURTISII Moldenke, Phytologia 2: 147--148. 196. Synonymy: Verbena caroliana L, ex Moldenke, Phytologia 2: 148, in syn. 196. Bibliography: Moldenke, Phytologia 2: 17--18. 196; Moldenke, Alph, List Invalid Names Suppl. 1: 23. 1947; Ho N. & A. L. Mol- 1 ee stems tetragonal, st ramineous, lightly strigillose; nodes with a transverse band of short white hairs; leaves de- pepemrga so Beg only the upper ones in and just below the inflor escence seen; petioles slender, 1—-1l ong, sparsely scattered pilose with s 33 Leahch tation thin-ch light-green, lanceolate, 1.5--l om. long, 7— de, acute or 20 mm. wi subacute at the apex, attenuate into the petiole at the bas x wi te broadly ar forward-pointing teeth, scatter- ed-strigillose on both surfaces with short appressed whitish hairs, more densely so along the venation be eneath; inflorescence abun- t; spikes ylides. numerous, two from each upper node, three at fruited, often branched toward the base » the branches subtend- ed by reduced leaves; peduncles slender, usually abbreviated to a- bout 1 cm., lightly strigillose; bractlets lanceolate, about 2 mm. long, long-attenuate at the apex, ciliate-margined, otherwise glab the frui pex of Phos fruit and forming a distinctly sharp point until the c S ruptured; fruit about 1 mm. iong, smooth, long-coherent. a type of this very puzzling species was collected by Moses Ashley Curtis -- in whose honor it is named — somewhere in "Carolina® [probably North Carolina] and is labeled "Verbena car- herbarium of the Botaniska Tridgard at Gvteborg, Sweden. The spe- S reminds on in its general aspect, but dif- te pubescence, It also resembles Vv. ur- Vahl, from which its non-s abrous leaves at once distingu- ish it. It differs from V- riparia Raf, and V. urticifolia in its dense fruiting-spikes and - from the former in its non-lobed leaves. Its full fruiting-calyxes seem to preclude xV. engelmannii Molden- ke or any other hybrid origin. A common name recorded for it is 1963 Moldenke, Monograph of Verbena 53 "hierba de la mula", In » 3 herbarium specimens, Apelnsi NS the type, and 2 moun- ted photographs have been examined by m tations: NORTH CAROLINA: County mndotennined Me. Ae Curtis sen. [e Carolina] (Go--type, N—photo of t » Z—photo 0 of type). CULTIVATED: Belgium: KM. Martens sen. [h. a i 1845] (Br, N). xVERBENA DEAMII Moldenke, Revist. Sudam. Bot. kh: 18. 1937. Synonymy: Verbena bracteata x ‘stricta Palmer, Ann. Mo. Bot. Gard. 22: 629. 1935 [not V. bracteata x eata x stricta pt eres 1941). Verbena stricta x bracteosa Bebb ex Moldenke, Revist. Sudam, Bot. h: 18, in syn. 1937 [not V. stricta x bracteosa Engberg, 1958, Sudam. Bot. ta x bracteosa steckonada ex Maadenike dk Sudam, Bot, h: 18, i syn. 1937. Verbena stricta x bractatea Rydb, ex Moldenke, Rie vist. Sudam. Bot. h: jot. h: 18, in syn. 1937. Verbena bracteata x stric- ta Gates, Fl. Kans. 190, 190. Verbena stricta x bracteosa Sand- berg ex Molderike , Alph, List Invalid Names Suppl. 1: 27, in syn. 1947. Verbena stricto-bracteosa Patterson ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 1947 [not = stricto- bracteosa Engelm., 190]. Torben stricta x bracteosa Waite ex Moldenke, Résumé Suppl. 3: 4i, in syn. 1962. a a stricta- bracteosa Michx. ex Molaenke, phiiat Suppl. 3: 41, in syn. 1962. oe bractiosa x stricta Holzinger ex Moldenke, Résumé Suppl. as in ae 19624: 8 = = ibliography: E. J. Palmer, Ann. Mo. Bot. Gard. 22: 629. 1935; Woldenke, Revist. Sudam. Bot. 4: 18. 1937; Gates, Fl. Kans. 190, map 1072a. 1940; Moldenke, Prelim. Alph. List Invalid Names Known Geogr. nac el 7," a). en 191,25 Moldenke, Alph. List Invalid Names 50. 19h2; Moldenke, Castanea 10: 38. 1915; Moldenke, ae List Cit. 1: 0 & 181. 1946; Moldenke, Phytologia 2: 327. 1947; Hill & Salisb., nd. Kew. Suppl. 10: 2h2. 1947; Moldenke, seen —, — Names Suppl. 1: 22 & 27. 19h7; eke Alph : 518 & 522 (1948), ¥ 699, 708, 773, 800, 887, 892, & Sore (2985), and ls 1098, 1229, 1255,"% 1261. 1919; Moldenke, Known Geogr. Dis- de: r str e : 131, & 13h (19525 and pr. 2, 3: 127, 131, & 13h. 1958; Moldenke, Am. Midl, Nat. 59: 315, 1958; fell eae Résumé 16; 17, 19; 22; . 25, 359, 375, & UR 88 Moldenke, Résumé S suppl - 3: ‘Se 6, %, & 1962) a Illustrations: peor) in Gleason, ee wee & Br. Illustr. Fl., pr. 1, 3: 134 (1952) and a, Be 2, 33, 13h. 29 This nan e has been proposed for the natural hybrid between V. weiyediien: Lag. & Rodr. and V. stricta Vent., commonly found where the ranges of these two species overlap in the central United 54 PHYTOLOGIA Vol. 9, no. 1 States, The hybrid reneubles V. bracteata in habit, but is stout- flowers, small ally not foliaceous bractlets which are most- « long and are long-attenuate or long-ac a the apex, the spikes usually o -10 m. wide and poorly fruit- ed, and pubescence in the inflorescence only soft lious. ollectors have found this plant in copses on ains, describe it as "decumbent", "procumbent", "ascending", © "erect", and have collected it in flower and fruit from June to Au ae type of ite preesy was collected by Per Axel Rydberg at pomruad’ in the Britton Herbar e New York Botanical Garden. The name, V. stricta x sores Waite is based on a SNe from Mount Carmel, Wabash County, Illinois, collec on August 6, 1887, and deposited in the herbarium of the cara of Ill- inois; V. bracteata x stricta Gates is based on J. B. S. Norton san. from Manhatt , Riley Co County, Kansas, collected in August, 1892, and dapuaites’ in the herbarium of carte State College; V. stricta x bracteosa Sandberg is based on Sandberg s.n. from Hen- nepin County, Minnesota, collected in August, 1881, and deposit- ed in the herbarium of the New York State Museum at Albany; V. stricta x bracteosa Eggert is based on Eggert s.n. from Saint Louis, Missouri, collected in August, 1886, and — in the herbarium of the New York State Museum at Albany; and V. stricto- bracteosa Patterson is based on H. N,. Patterson Pein from Hender- son County, Illinois, collected in July, 1872, and deposited in the herbarium of the ’ New York State Museum a e hybrid is recorded from Jasper County, Missouri Palmer (1935) and from McPherson County, Kansas, by Gates (19h0). Her- barium specimens have been misidentified and distributed in her- baria under the names ve bracteata Lag. & Rodr > Ve stricta Vente, V. bracteosa x stricta Eggert, and ve Seal thond eeetiie Engeln.; apes calls it "V. bracteata x hastata" » Which it certainly is no The so-called V, bracteata x stricta of Schneck is xV, perri- ana Moldenke; the V V. bracteosa x : stricta of Britton, of “Palmer, : _ rydbergii Bborglh Wordsnbss stricta x bracteosa of Hitchcock is xV. pe while that of Engberg is typical V. bracteata Lag. & ig ao The V. stricto-bracteosa of Engelmann is xV. perriana, Wallis describes the plant as "somewhat ascending. The H, Re Bennett s.n. (Brewster Arnold Ranch, August 2. 2, 19571 in the Uni- ted States National Herbarium bears dently a hybrid with Vv. stricta Vent. or some other vBct plttagts but I regard this specimen is representing typical V. bracteata. ARACHNIODES, A FERN GENUS UNRECOGNIZED FROM HAWAII Otto & Isa Degener Thanks to the lucid article by M. Tindale in peta South Wales Nat. Herb. 3 (2): 89--90. i961 we find the Arachniodes Blume as 1. Jav. 21. 1828) to be Neteeeented in the Hawaiian Islands by a form of what has been hitherto known locally as Dryopteris (Polystichopsis) carvifolia (Kuntze) C. ist ° B. P. Bishop Mus. 25: 10. 1925. is perky place this species in its rat genus as Arachniodes carvifolia (Kuntze) Deg. & Deg., comb. n BOOK REVIEWS Alma L. Moldenke "Plant tate of Palestine Israel and Jordan", by Michael Zohary; 33 nd 262 pp., illustr. Ronald Press, New York City, 1962. =| This new member of the worthy Chronica Botanica New Series of Plant Science Books heya readably the dedicated study over few score years of the taxonomy, geogr saber, ecology, physiology and economic im h crew of fellow scientists and a public — locally and abroad — very much interested in these studies. The text is organized ecologically. The sage aie ga esrb is not one of the many still-forming American systems, but th Simplified, the plant habitats are semi- —— — = arid. The rainfall varies annually from rth dowmm to 25 mm. in the south, with aide 75 uaa ae hg VLE fe itation felt in eee. January an ebruary Data is presented so effectively that one can easily envision the plant growth as the following items indicate: There are 718 0 or subalpine flora. The greatest numbers of species are fram the 55 56 PHYTOLOGIA Vol. 9, no. 1 following families in descending order: Compositae, Gramineae, Leguminosae, Umbelliferae and Cruciferae. The genera represented by the greatest mmber of species are Astragalus with hl, Trifol- ium with 0, Silene with 36, Euphorbia with 31 and Allium with 25. Weeds comprise 50 O species; endemics total 160 species; annuals make up 50 sala of the fl The very clearly reproduced ko ge and line drawings of many of the important plants and typical scenes as well as the clarity of the text show some of the c effects of time, earth forces and man from the Tertiary to the Natufian culture of 8000 and on to the present day on area, ally (as co-author with my husband on "P] of the Bible", Ronald ess, New York es my I wish bape there was m en of the Bible times bu h was not the intention e introduced since old and New Testament days? What ones from be hapter on "Man and Ae at ee potent reasons for a ae here The descriptions of desert and steppe plant life are some of the best ever printed. After reading this aren cot book I am all the more eager to visit and study this are "Atlas of Plant Morphology Portfolio IT: Photomicrographs © of flower, f d seed", by Emma L. Fisk and W. F, Millington; 48 pp. Burgess Publishing Co., Minneapolis 15, Minnesota. 1962. $2.75 Fine ra reproductions of well prepared slides of flower, fruit pa sone sections — often with a couple of illustrations — -- make this looseleaf portfolio very attractive and exercises in sever. urses on college campuses and even f high school biology. Students could use _ advantageously in conjunction with their own slid r with their tissue 3 drawings. Students c ertainly should be Pete "to tell the truth, the whole truth, and nothing but the truth" with their 4H pencils and/or stippling pens. an accurate oes of a small Now I am eager to peruse Portfolio I on roots, stems and leaves- PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. mber consists of not less than 32 pages. All manuscript accepted will be sages in the next issue, so that the size of numbers ee vary greatly. A volume will contain about 32 signatures, 512 pages, or a number of pages with an equivalent number of plates. This plan ‘nian immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra charge is made for line d rawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain limitations of size and proportion. An extra charge will be made for half- tones, depending on their size, as fixed by the engraver, with a minimum of about $2.25. a a dealing with research in =e ~— of botany, in any reasonable length, biographical sketches, and critical reviews and summaries of liter- of type, and polemics will not be published. Advice popular : on the stabi of manuscripts will be solicited, if necessary, from quali- - fied botanists. Under the parent Cost of pontine. the basic rate for a page 0} ecdlitic r fraction $3.00 for n of 250 copies. This price is subject — to ‘change without notice, since it a entirely on ae prices prevailing in the Sie industry. will be furnished at cost. A proportionate fraction of the edition ; of 250 copies is also furnished = to contributors. “Upon request, the editors will sn —— instructions concerning the: of manuscript or further information sist it _— magazine. In- oO the < neeganioe @ Or to él ither ec ae ah re Sane EE wipe Silane ens 6 a es Pome See eee ee PHY TOLOGIA Designed to expedite botanical publication ee ereereeteeehienenenessesnespneeneenen — Vol. 9 March, 1963 No. 2 uawcuw RoTANIcAt ROEN LIBRARY CONTENTS BENNETT, R. W., Notes on eastern American species iy PERSIOMON So OD e ood s cee oe es 57 MOLDENKE, H. N., Materials toward a monograph of the Rerar Veriemee Nd oir es Chass pe ene eek ew es OR 59 MOLDENKE, H. N., Notes on new and noteworthy plants. XXXV . 98 LAUGHLIN, K., Quercus x riparia Laughlin .........200+20+40+ 101 LAUGHLIN, K., Malus lancifolia Rebder.......... eee es 108 "Published | by Harold N. Moldenke and Alma L. Moldenke 15 Gleptont A venue Yonkers 5, New hes U. SA. NOTES ON EASTERN AMERICAN SPECIES OF PENSTEMON Ralph W. Bennett * The forthcoming publication of a taxonomic booklet on the east- ern American species of Penstemon suggests the desirability of a few taxonomic and nomenclatural changes being pu now. Most of these changes are in accord with ideas bare ye to me by Dr. David D. Keck in a recent informal conversation. Foll the recommendation in Article 22 of the 1961 Inter- posi Code of Botanical Nomenclature, the following peer s in subdivisions of the genus Penstemon are publis if Poh Eupenstemon is changed to Subgenus Penstemon, Section Spermmculus to Section Penstemon, and Subsection Graciles to Subsection Penstemon, The three names Eupenstemon, Spermmculus, and Graciles were used by Dr. Keck in his Studies in Penstemon VIII, American Widlend Naturalist Vol. 22, No. 1: 128-206. 1945. P, laevigatus Solander, P. digitalis Nutt., P. calycosus Small, P, alluviorum Pennell, and P. deamii Pennell are combined into one species, as follows: P. laevigatus Solander. ~ 88P._ laevigatus P. laevigatus Solander, Ait., Hort. Kew. 2: 361. 1789. ¥v Bas —Sigital is (Nutt.) es comb. gitalis Nutt., Sims in Bot. Maes “Bee pl. 2587. 1825. v S3P+ “eagcons (Seal) Bennett. comb. alycosus Small, Bull. Torre + he Club 25: 470. 1898 / om “sUurioru pment Bennett. comb. nov. agi: Small,.Man. S.E. Fl. 1203. 1953. Vv pes Pristaeag ae amii (Pennell) Bennett. comb. nov. P. deamii Pennell, ges No. 1, Acad. Nat. Sci. Phila. 212. 1935. P, pallidus Small, P. arkansanus Pennell, and P. multicaulis Pennell are combined into one species, as follows; P. pallidus Small Ssp. pallidus P, pallidus Small, Fl. S.E. Un. St. 1060, 1537. 1903. -~ 88p. arkansanus (Pennell) Bennett. comb. P, arkansanus Pennell, Proc. Acad. Nat. eat Phila. 733 494. 1922. Pp multicaulis Pennell, in Proc. Acad. Nat. Sci. Phila. 733: 494, 1922. * Hlykictereg of the American Penstemon Hani 5607 No. 22nd St., Arlington 5, Vi ia. 57 58 PHYTOLOGIA Vol. 9, no. 2 P, australis Small, P. laxiflorus Pennell, and Pie whe rryi Pennell are combined into one species, as follows P. australis Small ssp. australis Loe australis Small, Fl. S.E. Un. Ste. 1060 ,1337. 1903. /s 828s laxiflorus (Pennell) Bennett. comb. » laxiflorus Pennell, Monograph No. ly, grea Nat. Sci. Phila. 229. 1935. P. wherryi Pennell, Monograph No. 1, Acad. Nat. Sci. Phila. 228. 35. P. wisconsinensis Pennell is reduced to a variety of P. grac ilis Nutt., follows: P. gracilis Nutt. v Var. gracilis P. gracilis Nutt., Gen. N. Amer. Pl. 2: 52. 1618. / War. wisconsinensis (Pennell) Bennett. comb. wisconsinensis Pennell, Monograph No. ly, hdd Nate Sci ie Phila., 234. 1935. The following new varieties are published: P. laevigatus Solander ssp. digitalis (Nutt.) Bennett vare angustus Bennett, var. nove A subsp. digitali caulibus dimidio minoribus (ca. 40 cm. longis) et folius multo minoribus et angust- loribus (ca. 2 cm. latis) differt. P, hirsutus Willdenow var. hirs Fe pieces (L.) "eoiteg cure Ley Spec. Plant. ede IV. ae 2e7. w Var. Pygmaeus peceatk 9 Vare nove A var. hirsuti caulibus decumbentibus differt. MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. X Harold N. Moldenke VERBENA DEAMII Moldenke In all, 33 herbarium specimens, LNG the types of all the names invo lived, have been examine r| by m Citations: ILLINOIS: Coles Co.: shles “& Gilpin 7455 (Hi— 103233). Henderson Co.: H. N. Patterson s.n, [July 1872] (Al), S. n.[near Oquawka, July 17, " 1873] (Dt), sen. [Oquawka, July] (W— 149161), s.n. [vicinity of Oquawka] (N, Ur, W—-1323063, 1323123). Wabash Co.: Waite s.n. (Mt. Carmel, Aug. 6, 1887] (Ur). IOWA: Story Co.: C. E. Bessey son. [Ames, Aug. 23, 1878 ] (Va— pes WISCONSIN: Pierce Co.: Schuette s.n. [Bay Settlement, July 9, '83] (Du—122875). MINNESOTA: Henn epin Co.: J. H. Sandberg s. n. [Aug. 1889] (Al, Ca--2517h, Up--17089, W—71937) - theca ab Coe: Holzinger sn. [Winona, July 10, 1888] (Ca--67)16). KANSAS: Riley Co.: J. B. S. Norton s.n. [Manhattan, Aug. 1892] (Ka). Waneste alo ee Coes Rydberg s S.n. (Kearney, June 20, 195] (u—type), s.n. [Kear- ney, June 1895] (N). Lancaster Co.: J. L. Sheldon s.n. [June 2h, 1899] (We). Nemaha Co.: J, L. Sheldon s.n. (Peru, June 20, 1900] (We). MISSOURI: Greene Co.: "Eggert s.n. [Brooklin, 13 aly 1879] (Cm, N, ee St. Louis: Eggert 5321 (N), s.n. fAug. 1886] (Al); Engeluann s - (St. Louis, Aug. 1643] (Pr), sen. (St. Louis, July 1861] (Dt), s.n. [St. Louis, Aug. 1861] (Dt), s.n. [St. Louis, 1861] (Br); M. Martens s.n. (Br). OKLAHOMA: Adair Co.: C. S. Wallis 2385 (st ; ea VERBENA DELICATULA Mart. ex Zucc. in Otto & Dietr., Allg. Garten- zeit. 2: 243 & 245. 183k. ynonymy: Verbena delicatula Mart. & Zucc. apud Schau. in A. DC., Prodr, 11: 555. 18h7. Bibliography: Zucc. in Otto & Dietr., Allg. Gartenzeit. 2: 2h3 ' thing 1834; Schau. in A. DC., Prodr. 11: 555. 187; Jacks., Ind. ew. 2: 1178. 1895; Perry, Ann. Mo. Bot. Gard. 20: 32 & 355. 19335 Moldenke, Résumé Suppl. 2: h, ll, & Softly pubescent throughout; stems short ‘ “erect; leaves petio-— late, broadly cordate-ovate, coarsely serrate, subine ml lacerey at the base; racemes rather slender; flowers erect; calyx angled, long, the rim 5-dentate; corolla-tube narrow, curvate, the ed ly Freiherr Wilhelm barnes von Karwinski + Tehuantepec, Oaxaca, Mexico, in moderately warm places. gestae (1933) says "Un- 59 60 PHYTOLOGIA Vol. 9, no. 2 fortunately the essential characters, which would separate this species from its allies, are not clear rly defined. The description is so much like that of V. barbata that the writer suspects tha t the two species are identical." DELTICOLA Small ex Perry, Ann. Mo. Bot. Gard. 20: 314— 315. 1933. My My Verbena denticola Small ex Cory, Texas Agr. Exp. Sta oie 550: 88, | 88, sphalm. 1937. Bibli graphy: Per erry, Ann, 3 pets Mas 20: 248, 311, 31L-- 315, & 3. 219333 Gory, Texas Agr. Exp. Sta. Bull. 550: 88. 1937; nd, - Suppl. 9: 29). "T5538; Moldenke, Suppl. List Inval- re Names 8. “toll; olderke. Known own Geogr. D Distrib. Verbenac. » (ed. 1], 13, 18, & 101. 192; Moldenke, Alph. List Invalid Names 16. ahs H. S. Gentry, Carnegie Inst. Wash. Publ. 527: 221 & 306. 1942; A.M. T. Davis, Study Boscaje Palma 61. 1912; Moldenke in unde ,eed L}s 29, & 35-—-36. 1942; Moldenke, Phyto losis 2s 3s atl, e 128. 1945; Moldenke, Alph. ates Me 176, 215, 216, 232, é a (1946) and 3t0, 43 3, & bop. 3u8, Moldenke, 1: 230. 198; Yol- “ : 162, 1948; Moldenke Geogr. Distrib. ek : rp hey 32, 1949; Moldenke, Alph 6 781, 785, 795--797, 807, 829, 831, 878 861-883, &'933 (1995 and’: 980, 1071, 1072, ree as ’ gua. 1126, 1170, 1171, a8, 3230, 12h0, 12h6, & 1290. 19h9; &-A. L. Moldenke Anal. . Biol. Mex. 20: $i oe Moldenke, Brac fg 29, 39, 33, & aps nde ; Lewis & + Journ - 48: 639—61. 1961; Moldenke, Résume paring ne 7 . 10. stoma! Annual or eae perennial herb, to 30 cm cumbent or ascending to erect, b ranched, more or — Facer goth i dentate, often 3-lobed, usually thin-textured, sparsely appress- minute bulb aling odorless or fragrant with a faint odor: calyx Teen Mages t sparsely glandular, purple, purple, deep pinkish- pis lavender-blue, Be nip or violet, sometimes pink- berg gins a whitish " or "fuchsia-color", its tube 1— 1.5 cm. long, protruding oe beyond the calyx, pubescent or glabrate on the outside, the limb 7--10 mn. wide, the lobes e- marinate; . cocci about 3 mm, long, subeylindric, reticulate from 1963 Moldenke, Monograph of Verbena 61 the apex to the definitely broadened base, the commissural faces practically reaching the tip of the cocci, muricately scabrous; chromosome number: n = 15 ris and Carl Dudley Duncan (no. 3161) at Las Palmas Ranch, in the vicinity of Br e, eron Coun’ ; en August 1 and 1921, and is deposited in the Britton Herbarium at the New York Botanical - In southern Texas the species is said to Sandy soil, clay, loam, or sandy clay loam en grounds, fields, or orchards, or at the edge of thickets, often covering acres of ground, ing ruary t on the as yet seen by me, and that the species is frequent to abundant in early spring "throughout this region" and "throughout Cameron colonies, covering small or large areas, t t hou acres of ground", Gentry asserts that it is only "casu scattered on milpas in the Lower Sonoran zone" in Sonora, Leaven- tufa worth found it on limestone bedrock covered in spots with in Nuevo Leén, where Mueller reports it as "rare on sand a 8 " and Thompson found it "in red sandy clay soil on rocky mountainside in dense oak-pine woods" eaver found i and W in "loose sandy soil near rivers". Buchholz , however, says "can- mon in region" [Nuevo Leén]. Collectors have found it in sand, sandy loam, sticky black canyons, pine woodlands, clearings, open grounds and fields, or ange orchards, open valleys, roadside ditches, open low pastures, arroyos, and thorn forests on volcanic cerro slopes, at the edge of thickets, among grass, and along roadsides, at altitudes of 10 to 2000 meters. It has been collected in anthesis and fruit in every month of the year. Gentry asserts that the species in Sono- ra inhabits sandy adobe soils in valley margins of the Short-tree It is mentioned by Anna May Tarrence Davis in her thesis "A study of Boscaje de la Palma in Cameron County, Texas, and of Sabal texana" (August 192). Common names recorded for it are "alfombrilla", "bartunicha", "kamiyo", and "moradia", Specimens have been misidentified and distributed in herbaria under the names V. ambrosifolia Rydb., V. aubletia Jacq., V. au- bletia L., V. bipinnatifida Nutt., V. canadensis Britton, V. ciliata Benth., "V. sp. aff. V. ciliata", V. ciliata var. longi- Gentata Perry, V. elegans H.B.K., V. elegans var. asperata Perry, Ws gooddingii Briq., V. gooddingii nepetifolia Tidestr., V. 62 PHYTOLOGIA Vol. 9, no. 2 gooddingii var. nepetifolia Tidestr., Vv. lambertii Sims, Vv. pum ila Rydb., and V. quadrangulata Heller. Nealley 117, Edw. R&S 90, Re Runyon 1943, and J. N. Weaver 667 annotated as V. delticola b; by Perry and cited by her or so identi- fied and distributed, are all V. cameronensis L. I. Davis; G. L. Fisher 45198 is Vv. elegans var. asperata Perry; and J. N. Weaver 1033 is ve. pumila | Rydb. The Edw. Palmer 1045 annotated by Perry as "Aff, V V. delti delticola Small" is V. quadrangulata Heller and the Edw. Palmer 3 34, similarly annotated, is V, andrieuxii Schau. the E. W. “Nelson 6028 and aoe Palmer 3 39 in the Britton Herb- arium are anomalous in appearance and may actually not belong here. Frye & Frye 2310 has a the aspect of V. gooddingii var. nepetifolia, L. Irby Davis, in a letter to me dated July 28, 191, says "I am at present interested in studying the variations in v. deltic 1tice ola Small. There are certain features in pt ae from the type Tocality ( some of which can be observed usually o in fresh material) which I find are not present a specimens called V. delticola ate E. San Luis Potosi and S. Tamaulipas. Also I “feel sure that there are at least two species in San Luis Potosf that are usually identified as V.delticola. There are seasonal, cal- endar and habitat variations in ve wee | in my home county The Ecology Class Univ, Texas xas S.n. sabe in Lundell, Fl. Texas 3 (1): 3% (i9h2) as ve r, delticola is actually Ve c ee Le I. Davis, Similarly, the Nealley 117 and 118 and Edw. Palmer 90 {insofar as seen by me], cited by Perry (193k) as Ve oy aternie® are actually V. cameronensis,. Miss Perry cites the following 15 aces Sunes a not as yet examined by me: TEXAS: Cameron Co.: Ferris & Duncan 3161 (E—- isotype). MEXICO: Nuevo Leén: Arséne 6129 (E, G); Gregg Gregg 202 (E), 752 (E); Edw. Palmer 1051 (G); Pring Pringle 2 2228 (G), 11843 (G). Tam- aulipas: E,W W. Nelson U2) (c), 6628 (E, G); Edw, Palmer 39 (E, F, G). Vera Cruz: Ervendberg eon (G). She says "Superficially, this species bears § strong resenblance to V. canadensis, but usually the leaves are not so deeply incised. Its distinctive character is found in the nutlet. Commonly, in the section Gland- ularia, the schizocarp is shallowly lobed at the apex; hence, e, the ps appears be attached in a very definite depression and the ssural face does not reach the tip of the mut- ee In V. aeitiocia. however, the depression is indefinite, the present." The Liebmann 11313 & 1131}, Nealley 117, and Edw. Palm- 1963 Moldenke, Monograph of Verbena 63 er 90 which she cites are V. cameronensis. She has annotated E. Stearns 10h as V. ambrosifolia Rydb., but I do not think that it belongs there. Instead, I am ci iting it as V. delticola -—- its leaves do not look like those of V. delticola, but its flower Spike does, Of Mearns 591 she says 3 "Of, Verbena ambrosifolia Rydb.", but, again, I doubt that it is that species. E. W. Nelson 6628 has two labels on the sheet which originally indicated its number as "6028", In all, 129 herbarium specimens, including type material of all the names involved, and 1 mounted description have been ex- amined by me, Citations: TEXAS: Cameron Co.: Cory 28301 (N), 28302 oe L. i. Davis s.n, [Olmito, Dec, '1] (Au--122061), son. . (h mi southeast of Rang erville, December 2, 195] (Au—171995); ate & Johnston oo 21 (Au—-122065) ; Ferris & Duncan 3161 (Du--125),2)- isotype, G -31)67—~isotype, N—type) 3 H.C . Hanson 66 (W— 98282h) 5 Hotchkiss 618) (Ar--219537); Me Ce ~ Johnston 54150 (Au-- 122066, St, St); F. L. Lewton 11 (Ar--271586, W—56209); C. Le Lundell 10680 (N, Rf, Rf, W--1926930), 10758 [South. Appal. Bot. Chub 17th Dis Distrib. 1647] (3i--1N8517, i -=1p0510 N, N, Rg, RE, Rf, We); Lundell & Iundell 10018 (Id 18886775; # Parks 2936 (Wi, Wi, Wi), 2937 ‘jawed sie (Wi, Wi); a & Cory 17935 (tr); R. Runyon 327 (Au--122062, Au, "W—1222821),- 1761 (Rr, Rr), 19h3 (N, Rr), 2182 (Rr), 2363 (N, N, N,N, N, N, N), 2587 (N, N, N, N, N), 2693 (N, N), 392 (N); I. Shiller Shiller 33h (W--1812017) Small & Wherry Try 11901 (N). Hidalgo Co.: L. H. “L. H. Bai- 7431 (Ba); Clover 106 (Fs, Gg—-233345), 566 (Fs, Me); L. He fone a 5999 (W—1h68812); Lundell & Lundell 9802 (Ld, N), 950k” (Id, N, " W—1888832), 9828 (Ld, N), 9979 (N); Molby 722h (w— 3h); H. Be Parks 293 2937 [Mission] (Wi); Tharp arp 5999 (Au— 122063) ; peel no (Au--12206,, N). Maverick Co.: Parks & Cory 12422 (Tr). Willacy Co.: M. C. Johnston 5325314 (Au--12202h) MEXICO: eee E. Stearns 3 10h (W--5028),8). Coahuila: E. G. Marsh 118) (st), 1592 (St), 2h7 (St). Nuevo Leén: Abbon 19 [ar- séne | 6129] (N, Ur, , W—1003536); | Arséne s.n. (Monterrey, 5-1911] (B, N)j F. A. Barkley 14361 (Au); Barkley, Webster, & Rowell 7131 (Au-12327h, N); J. T. Buchholz s.n. (Feb. 23, 1936] (Ur); Lye I. Davis s.n, ([Teran, March }, 1959] 9] (Au—-171918) ; Herb. Inst, Biol. Univ. Nac. Mex. 7138, in part (Me); Joes m & Bark- ley 1 16129M (Au); W. C. Leavenworth 119 (Ur); Lewis & Oliver ver Skis” (Nb); (Nb); C. H. Mueller 2058 (Mi); Mu: Mueller & tre 158 ( (Me); Oldendort & Oldendorf s.n. (April 23, 193] (Ok—16316); C. R. Oreutt 1191 (W--1207770); Edw. Palmer 105 setgeagllle 1051 (Pa, W—56180); F. W. Pennell 166L4 (D--733978, Me, Me, W— 64 PHYTOLOGIA Vol. 9, nO. 2 164027h) ; Pringle 2228 (Me), 11843 (Ca—1683)1, ——- Mi, Mi, St, Vt, W-l)62591, W—1586)30); Rowell, Lind, 16M58h (Au); Safford 1221 (W—573262); Tharp 1826 samoce te W. M. W. M. Thompson m 258 (1 (Au——167)18) ; J. Ne Weaver 562 (W—21341)3), 1033 (N); S. S. White 2h (Mi), 1539 (Mi), 1576 (Mi). Puebla: Goldman 42 (W—321,807). = Luis Potos{: Graber 183 (W—2085)22); Edw. Palmer 34 (N). Sonora: R, J. Ferris 8770 (W--1686987); Frye & Frye 226 (W—179098h), “2318 (Du—2854,0, Rs--25552, W— 1790989); H H. S. Gentry 356m (Ak—20069, Mi), 1312 (Ak—-19968, Ca- 646402, Fs, Ge, I, Me, Me, Mi, N, N, S, 5) 92 a in part (Ge), 7942 (Ak—-22061) ; Siveve 6081 (Fs); S. S. White 708 (Mi), 77h (i), 3787 (Mi); Wiggins 6102 (Du—215932), 6288 TGa--S90L50, Mi, W—1739922), 7225 (Mi). Tamaulipas: H. H. Bartlett 10127 (Mi), 10170 (Mi), 10985 (Mi), 11133 (Mi); LeSueur 00 (Au, Fs); Lewis & Oliver 5420 (Nb), 5421 (Nb); EB. W. Nelson 42h (W—-330977), , 6028 (N), 6628 (W-3h7257), s.n, [May 1898] (W—-203186); fate a 39 (Ca—1)5872, Cp, N, W--572267), 90, in part (Cb Lauber, & Taylor 2100 (Du—366279, N, N); Viereck 987 “Cre Vera Cruz: MacDaniels 454 (Ba). State un determined: M {San Bernardino ranch, Mex. bound. line] (W—591720); = 79 ia: MOUNTED DESCRIPTION (N) . VERBENA DELTICOLA f. LILACINA L. I. own Bibliography: A.M. T. Davis, Study Boscaje Palma 62. 192; Moldenke, Known Geogr, Distrib, Verbenac. Suppl. 1: 2 & 4. 1913; Moldenke, Si ge 2: 128. 19453 Moldenke, Alph. List Invalid Names Suppl. 1 : 23. 1947; Moldenke, Known Geogr, Maeett Verben- ee. [ed.2], 23 & 197. 1919; Moldenke, Résumé 29, 363, This form is said to a ae the typical form of the spe- cies in having la The type of the form aaa Peobhestea by L. Irby Davis on Sage says "there must be a gia g ar owering plants of 1. “geltie- ola to one of the lavender form. It seems to be a mutation that is likely to happen anywhere though and once it occurs is likely to spread locally, Although this color form is sometimes found up with masses of vec ar. longidentata plants, it 1963 Moldenke, Monograph of Verbena 65 s just as likely to be found where there are no V. ciliata vari- Suite nes miles around. should be borne in mind that in many cases where a series of pac ts, is given in the Sooasiiian of the flowers of a species, these are taken from notes by collectors on their labels at least, merely re time of their collection not nearly so many colors would corded. Therefore, the corolla-color may in many cases be far more uniform than these notes would indicate. 183. 19530 Phytologia 3: 286, nom. nud, (1950) and 1953 pein 183 Bibliography: Moldenke, Phytologia 3: 286 (1950) and i eign 19533 Moldenke, Biol. Abstr. 27: 1887. 19535 Berea Bot. Gard. 9: 77. 1955; G. Taylor, Ind. Suppl. 12: i. “1989; Moldenke, Résumé 80 & 71. 1959. ange rosttate or sprawling per rennial herb with heavy woody roots stems issuing fram the crow; stems oe branches numerous, slaidaes rather acutely tetragonal, decumbent or ascending, often closely appressed to the ground, minutely stbagtiions with tiny widely scattered hairs or glabrate; nodes annulate, often marked 3 nodes much abbreviated, to 2 cm. long, usually much less; leaves decussate-opposite, subsessile or short—petiolate; petioles fili- form, about 1 mm. long, ilose on the margins with s ae ascending hairs; leaf-blades chartaceous, rather uniform- peleicas npadgya Yea iam a prominulous be- neath; vetnlet reticulation indiscernible on both surfaces; in- many-f1l rate or very minutely wigpe cage acess: rachis Hpelpa il he t; f this species was collected by Wendell Holmes Camp (no. E. B.2010), in the Parroquia Luis Cordero near the village of San | an Marcos, 5--8 km. northeast of Azogues, Caflar, ae on A- a 1, “r9hs: and is deposited in the Britton Herbarium at the ew York Botanical al Garden, The collector notes that the plant is 66 PHYTOLOGIA Vol. 9, no. 2 sometimes used as a fever cure and is called "verbena echada" (i. e., -down verbena]. It has been found among weedy vegetation in hard gravelly soil, at an altitude of 2550 meters, blooming in February and September In all, 6 herbarium specimens, including the type, have been examined by mn me, Citations: ECUADOR: Azuay: Asplund 17801 (S, S); W. H. Camp E.1851 (N, mt Bi trsesee Giler 31 (N). Caflar : W. He Camp E. E.2510 (N—type). Province undetermined: L. Fraser s.n, (Bm). XxVERBENA —— Moldenke, Phytologia 2: 148. 19h6. Synonymy: Verbena hispida x bonariensis DBermen, Cytologia 7: 163, 16h, 16s 170, 171, & 175. pes, Verbena dermenii Moldenke in Chittenden, Roy. rhe Soc. Dict. Gard. 4: 2210, sphalm. 1951. Bibliography: Dermen, Cytologia 7: aes fig. 18, 164, 165, fig. 28 & 29, 170, 171, & 17. 1936; Moldenke, Phytologia 2: "118. 1916; A. L. & He N. Moldenke, Pl. Life 2: 56. 198; Png Known rib. Verbenac., [ed 2, 16h & 197. 1949; Moldenke, Phytologia 3: 289. 1950; Yoldenke in yi ay tel aor a Soc. Qa @ * Dict. Gard. h: 220 345—3k6. 1958; Moldenke, Résumé 115, 118, 223, 309, "366, & in. 1959; Moldenke, Résum eeuppls 3837s 2. fe Ijlustrations: Dermen, Cytologia 7: 163, fig. 18, & 165, fig. & 29. ° his name has been proposed for the hybrid produced artificial- ly in Massachusetts in 1936 by Dermen between V. hispida Ruiz & Pav. and V. bonariensis L. It is named in honor of Haig Dermen, contemporary Armenian cytologist and geneticist, born in Turkey and later research associate at the Arnold Arboretum in Jamaica Plain, Massachusetts. The hybrid has been found nine times already in the wild and I am certain that it occurs c where the ranges of the two . agu. a, ince and 9 provinces of Argentina. It seems very probable to me that some, at least, of the more or less "anomalous forms" of V. bras- iliensis Vell. and of V. litoralis H.B.K. pete in herbaria actu- ally represent this hybr- id, because its general appearance is much like that of a slender-spiked V. brasiliensis or a dense- spiked V, litoralis. ani. atlnge aol It is described by collectors as a suffrutescent herb, 0.5— m. tall, with blue or lilac-blue corollas. It has been cenlacted at altitudes of 100 to 3900 meters, in rinJ » Feb- rui two stinguished from V. hispida by the fact that its Gactiote equal or but slightly surpass + the calyx, and from V. bonariensis by its non-clasping leaves and more elongate pedunculate spikes 1963 Moldenke, Monograph of Verbena 67 which are solitary or in open panicles, Herbarium material has been misidentified rs distributed under the names V. bonariensis L., V. bonariensis var. longibracteata Kuntze, and V. littoralis var. caracasana Kunth. The Morong 1510 eétladtiod appears to be a mixture, because the Columbia Uni University Herbarium specimen of this number is definitely V. bonariensis Im al}. 12 capaiggaa specimens and me \ dbhited photographs have been examined by m Citations: BOLTVEA: La Paz: Asplund 3565 (S, Us); M. Bang 20h (C); Buchtien 459 (W—1177980), 5530 5530 (W—1159359); Mandon ion 523 (Bm); He H. Rusby _ 707 (C). PARAGUAY: | Hassler 3853 (Ca—9l363) ; Morong 15],0 150, in part (W—141672h); T. Rojas Rojas s.n. (Herb. Osten 18192] (F—photo, N, N--photo, Sg—-photo, Ug, Ug, Z—photo). VERBENA DISSECTA Willd. ex Spreng. in L., Syst. Veg., ed. 16 750. 1825 [not V. dissecta pasts | is0h, nor Poepp., 1807, nor Schau., 1959, nor Walp., 1 Synonymy: Shuttleworthia dis mecha” (aia pi, tian Syst. Bot. si 1B. 1845. Shuttleworthia dissecta Walp. Schau. in Mart., Fl. Bras. 9: 194, in syn. 1851. Shut tleworthte dissecta (wanaa’) Walp. apud Moldenke, Lilloa 6: 327, an ape Ste oars laciniata Kuntze ex Moldenke, Suppl. List I , in syn, 194] [not V. laciniata Briq., 1960, nor (Le) Bria ae nor (Lam.) Briq., 1939, 8 — & Moc., 190]. andularia dissecta (Willd.) Schnack & Covas, Darwiniana 6: ue oy ee dissecta Augusto, Fl. Rio gos: do Sul 217, C. 5: srinoides Osten ex Moldenke, Résumé 361, in syn. == Tate srinoides Auth, ex Moldenke, Résumé 363, in syn. 1959 [not V. ve. erinoides Chod., 190k, nor Hook., 1959, nor Hook, & Arn., 1959, nor L., 1959, nor Lam., 1791, nor Poepp., 187, nor Spreng., 1830, nor Willd., 197]. Verbena dissecta Spreng. ex Moldenke, sumé 36 i ee ees, Bibliography: Spreng. in L., Syst. Vegs, ed. 16, 2: 750. Walp., Nova Act. Acad. Nat. Caes. Leopold Carel. oes 5 .. es ? dad: Pings Walp. Repert. Syst. Bot 33. 185; in A. DC., Prodr. 11: 551 & 552. 18h7; C. Gay, Pe vb. & sageot Bot. eg 16. 189 ; AR in Mart., Fl. Bras. 9: 3 ae 1851; Weddell, Chloris Andina [Castelnau Exped. Bot.] - 186 3 Morong, 197. era Jacks., me Kew, 2: 895 & 1178. 1895; Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8: gh 1904 Reiche, Chile 5: 292, 19105 aan: Florula ~ 105. 1930; Le He & Ee 68 PHYTOLOGIA Vol. 9, no. 2 Z. Bailey, Hortus, new rev. ed., 632. 19353; Troncoso, Darwiniana : 483-86, fig. 2 & 3 g——n. 1939; Moldenke, Carnegie Inst. Wash. 9415 Schnac ° Cat hz: 19 & 20, 1942; Moldenke, Known Geogr. Distrib. Verber ac., [ed. 1], 39-42, bh, 7h, & 101. 19423; Moldenke, Alph. List nvalid Names 8. 1942; Moldenke, Lilloa 8: 29. 192; ll: 9h & 97. 19hh; Schnack & h: ISA 945; Ind. Taxonom. i 1y——16): 6. ie Cabrera, Bol. logia 2: 338 & 386. ; » Alph. List Se 2: 355, 364, 367, 375, 378, 426, Lhe 479, 533, 53h, 599, 629, & yl (19 918), 3: 660, 670, 687, oe 697, 703, 706, 732, 735, 7h5, i 748, 765, 766, 7 803, 838, 2. 874, 875, 909 911, 913, pe 963 (1918), and hs 2013, mh one og 0871092, 100 er a i Bi Castellanos & Ragonese, Lilloa 20: 259. igh9; Moldenke, epg ie Reapaccem Bot. 8: 173. 1950; Moldenke, Phytologia aL 167. a Invest. Agric. 5: LO, 76, 83, & 208 (1964) a8 and ake “th, & 206. 1951; Moldenke, Sp. Subsp. Cont. Mold. Set 45 [3]. ene Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard, 6: 2209 & 2210. 1951; Moldenke, Inform, Mold. set h6 Midl, 121 , 127, 223, 295, 3h, 359, 361—363, 38, 379, & kn. 198; Moldenke, Résumé Suppl. 1: 7 (1959) and 2: lle iz 1 j Aneely, Fl. Paran, 16: 78 (1960) and 17: h6. 1961; R , Sellowia 1 3 (13): 110. 1961; Moldenke, Résumé Suppl. 3: 32 5 35 (1962), vas pe 19 2), and 5: 6& 7. 1 Darwiniana much ssa mostly decumben trailing or creeping at the base, rooting adventit- ending rect, slender, subterete, usually 12—60 cm - tall, occasionally tolm, long;” principal internodes short, i—3 on. long, but on 1963 Moldenke, Monograph of Verbena 69 floriferous stems elongated and 4--6 cm. long; aeavest decussate- opposite, ovate, 2.3—l cm. long, cuneate at the bas attenuated and decurrent into the petiole, broadly an eet ie or hirsute above, spreading-hirsutulous or hirsute with hairs (which are * especially dense on the venation and margins) beneath, subrevolute or revolute on the margins, the leaf-segments r sublanceolate to linear-oblong, subobtuse at the a entire or sometimes to 3 inflorescence terminal, the spikes short, fastigiate, 2 or 3 times t. chotomous, very mary: wered, corymbiform, often cymose- culate, not at all or hardly e aft thesis; peduncles short, to ch cute at the apex, densely hirsute or white-pubescent; flowers spreading, not odorous or fragrant with the odor of vanilla; calyx tubular, elongate, 7-8 mm. long, about three times as pink, add bright-rose, clear-rose, or even Lignt-red, $ its tube 10—- « long, slightly longer than the calyx, glab us below and pelaeient above on the outer surface, densely ts within, t t% con ven , the commissural face narrowly acute~ angled, sub-bisulcat smooth; chromosome number: 2n = 10. e of common southern South American speci Osten 3002 from Costa ‘dal Rosario, Sak mas. Uruguay, col- lected on ae Be 189), growing between "V. erinoides" and "VY, chamaedryfolia", I do not believe it to be of hybrid origin; it seems to be Bes V. dissecta. The only true hybrid of this species know to me is with V. temisecta Briq. and is herein discussed under x¥, nequam Moldenke. It is worth noting here that the V. dissecta of Morong is V- 70 PHYTOLOGIA Vol. 9, no. 2 tenuisecta Briq., that of Schauer is V. berterii (Meisn.) Schau., and that of Poeppig and of Walpers is V. sulphurea D. Don; the V. erinoides of Chodat is V. calliantha Briq. and V. storeoclada Briq., that of Lamarck is V. laciniata (L.) Briq., that of Willde now iu V. ciliata Benth., and that of Hooker, of Hooker & Arnott, of Linnaeus, of Poeppig, and of Sprengel is V. berterii (Meisn.) nie and the V. laciniata of Briquet and of "(Lamarck) Brique! ve laciniata (L.) Briq., while that of Sessé & Mocino is adel prismatica var. laciniata Grenz. The V. dissecta f. a ae fera (Sanzin) Moldenke is V. perakii (Covas & Schnack) eres dissecta inhabits dry, dry-arid, cultivated, sunny, and high dry soil, | soil, sandy banks, matorales, dry barren scrublands, quebradas, fields, pampas, and. dunes. It has been collected along railroad tracks and road embankments, in grassy meadows and grassy fields, campos, parks and waste ground, wet ea and rocky fields, * pastures and barnyards, on stony hills and sandy- rocky hills, "an br ambly places and sandy places, = salty ground, among grasses, at the edges of small arroyos, in "campo abierto" and Juncus acutus swamps, often forming handsome pa a in ters. It has been collected in anthesis and fruit in every month of th ra says of it "campestre poco frecuente"; Mal- donado reports it as everywhere" in Santiago del Este- ro3; Schulz that it is abundant around Rfo Sali, but the » Pac - Rosengurtt sistas that in Uruguay it is found "en acon bajos y rastrojos Common names recorded for this sue are "cutleaf verbena", "margarita morada", and "margarita morado", Vervoorst asserts that the color of the corolla is that of "Maerz r mek Pl. i H-3", but that when dried the flowers lose their or Her’ al has been misidentified and distributed un der the names V. cabrerae Moldenk alliantha Briq., V. crinoides Lam., ¥. erinoides Lam., v. Sinaloa Hook., V. laciniata (L-) pei V. microphylla H.B.K., V. pulchella Sweet, V. radi- cans Gill. & Hoo ok., V. sulphurea Sweet, V. tenera Spreng -, Ag temisecta Briq., and ev. aff. erinoides Lam." Osten determined On the other hand, the Claude-Joseph 128, 3548, 456, & 5053, distributed as V. dissecta, are actually V. berterii (weisn.) Schau.; Claude~Jos 988 is V. lipozygicides Walp.; Cook & Gilbert 549 is ve. necroptylis H.B.K.; Sehnen 5052 is V. r, tenera Spreng.; and W. E. Broadway 1997, Faris 478, Lazar 32, Morong 219, Pedersen 1325 & =the Rambo 51652, and F. A. Wetmore B30 1963 Moldenke, Monograph of Verbena 71 are ve tenuisecta Briq Barker asserts that V. dissecta is"a verbena commonly cultiva- ted in Puerto Rico gardens. It occurs in two varieties, pink and pier, " Troncoso says that it is cultivated in Argentina. Appar- it was first introduced into cultivation commercial 1920. The pee pled (1935) assert that the material in the horti- cultural trade passing under this name is actually V. pulchella. The four specimens in the Bailey Hortorium herbarium examined by me, however, under this name, are tru ev. dissecta, so I think that Dr. Bailey was in error is his Fale fo Chicchi 26) is a mixture with V. hookeriana (Covas & Schnack) Moldenke, ° ee (1939) gives the date of Weddell!s reference as "1857", but the part referred to seems definitely to have been published’ eet in 1860. Schauer (187) says of Ve dissecta: "Habitus fere ee (V. erinoides Lam.], a qua vero sicut a praecedente [V. tenera Spreng . } ian ee alia jam ek abe ey pr culata spic ine vix s " In hi plifies this. to "Habitus fere ver benae wis idis, a qua ve Sicut a V. tenera differt; caule suffruticoso, hirsutie wasted; Spicis brevibus fastigiatis neque elongatis saepe in paniculam sam erdum amplam collectis; ab illa insuper recedit caly- oribus late tripartitis waa tadaciies inciso-dentatis, modo pinnatifido- laciniatis,." Troncoso (1939) says of V. dissecta: "nueva para la flora ar- ge . — Esta interesante especie conocida de Chile, Peru, Bolivia, Brasil y Uruguay, no habia sido citada hasta ahora para la Argentina. Probablemente, esto tenga por causa su afinidad con V. tenera Spr. y V. laciniata (Lam.) Briq., con las que puede facilmente co: . No seria de extranar que s herbarios bajo alguna de estas denominaciones. La e por primera vez en las ancas de la localidad bonaerense de Camp- ana y Posteriormente, en Julio del ano pasado, fue he peo el ingeniero Arturo Burkart en las barrancas rib 3082, Troncoso 172, and Burkart 10191 from Buenos Aires, and Fel- ippone 6g, 6975, & 28),7, Hick Hicken 74, and Rosengurtt 2116 1/2 uguay m “all in the herbarium of the Instituto Darwinion at ont. pocos a Ocupado de esta sepebin coinciden en considerala como planta amal, solo Reiche....,duda de esta caracter. Indudablemente es bar planta et se halla en cultivo “ el Darwinion desde mediados del ano pasado.....Es vecina a V. tenera Spr. y V. laciniata ( (Lam.) Briq., de las que se ears por la treba 72 PHYTOLOGIA Vol. 9, no. 2 cia en espigas mas densas y munca alargadas en la fructificacion, y por sus glandulas anterales de mayor tamano y netamente visibles en la garganta del tubo eae ea aoe ejemplar leg. Burkart, SI. 10191, cultivado en el Darwinion, se ha Seaateorass con todo vi- gor, Meese en plena floracion como una gran mancha de color de 1 m de extension, Su rapido Sethaaa 3b su _— pepe apes pad fenktaizes por gajos y sus vistosas flores, react esta planta muy recomendable para ne abcd de Bbecuigurtt = 1946) says "Mala hierba perenne, estolonffera ach- atada contra el suelo, de ciclo indefinido. A Abunda en campos vir- genes y de rastrojo. Es constituyente accesorio de las estructur- as degeneradas, y disminuye en los campos regenerados." In fiis 1943 work he says: "Hierba hemicriptéfita y estolonffera cundid~ e+eseEsta planta fué publicada anteriormente por los aut eee os; de amplia ite ed Sudamérica. En Argentina habita desde hasta Buenos Aires y Rfo Ne sre Walpers a & 185) a peat 157 for this species, but Schauer (187) cites it for us sulphurea, The latter author also (1847, 1851) cites Sellow s.n, from Uruguay, Née s.n. from Chile, Meyen s.n, from near esaoae alt. 15,000 feet, Peru, and Lund SR, Raben s Raben s.n., Riedel s.n., and Sellow s,n. from S&o Paulo, Augusto & Edésio s.n. from Odério, Rio Grande do Sul, Brazil. Schnack (192) refers to a "VY. aff. dissecta x V. peruviana", but just what this hybrid actually is has not yet been determined by me. Briquet (190) suggests that V. selloi Spreng. may be con- Brig. in my Alph. List Cit. h: 1090 (1949), while the . Jorge 1613 cited as V. dissecta in the same work, 2: 599 (198) is the type collection of V. andalgalensis Moldenke. For a key to distinguish V. dissecta from related species, see pages V. laciniata, ,» 268 herbarium specimens and 7 mounted photographs, in- oleate’ t type or phototype material of all the names involved, have 1963 Moldenke, Monograph of Verbena 73 been examined by me, tations: BRAZIL: Rio Grande do Sul: Jttrgens 74 (B); Rambo 10103 (N), 4891) (N). So Paulo: Brade 7000 (N, N, Sp—6728); Col- lector undesignated 217 (Sp—20061) ; Freire & Azevedo 110 (Ja—— 2187); LOfgren s.n. [Itapetininga, Sept. 22, 1887; Herb. Com. Geogr. & Geol. 3685] (N, Sp--15725). State undetermined: Raben 828 (Br); Sellow s.n. [Brasilia] (Br). BOLIVIA: Tarija: Troll 261 (8). Province undetermined: Kuntze s.n. [Macbride photos 1742h] _ (Kr--photo, N--photo). URUGUAY: Collector undesignated s.n. [San Carlos, 8 Dec. 188] (Ug); Commerson s.n. [Macbride photos — in part] (Kr--photo); Felippone 6118 (W--1858371); Gallinal, gone, Bergalli, Campal, & Rosengurtt 1433 (N), 1625 mets x 1821 (N), PE.5262 (Ml, (Ml, N, N, Si); Herter ‘ter 50882 (N), s.n. {Herb. Osten 17657] (Ug); Osten 3002 (Ug), 3305 (Ug), 539 (Ug), S6k2 (Ug), 6386 (Ug), 7793 (S, Ue, Ug), 10356 (S, Ug), 1h676 (S, Ue), 16039 (Ug), 22096 (Ug); Rosengurtt B.2116 (N), B.3756 (M0, N, N)j Seijo 650 (Ug), sen. [Colon, 30 Nov. de 188k] (Ug). CHILE: Province undetermined: d: Née 8.0. pes soos ae Macbride photos 17460] (Kr--photo of type » Z--photo of type). ARGENTINA: Buenos Aires: hag Soy 330. oF ates 10712 (hiv) ; Carette s.n. [Monte Hermoso } (N)3 Cc Collector undesignated s.n. (S); Camerson s.n. [Macbride photos 39502, in part] (Kr— BERR eSieabagh Hauman s.n. [Sierra du Tandil, XII/1905] (Br); - HE, Hunziker 38) (S); Lefebre s.n. [Monte de Talas, La Plata, Tit} (Br); Ee Meyer 6990 (S); Née 30 (Q), 96 (Q)3 Ruiz Huidobro 1176 (S). Catamarca: F. Ae Barkley I Toarsé1 (N); Brizuela 375 (N, Ok), hog (n), LLS (N), 502 (N), Su6 (N), 754 (N), 826 (N), > 872 (N, RE), 883 (N), 993 (N), 1037 (N), 1197 (N); Jorgensen 1613 {Ret Inst. re guel L owe . 31745; Herd. bast a0eF0 676] (H, He N, Ue); una Risso 39) (N 23 (N Malvarez 3; B. L. (Wy), i09 Te hoe Zi11529 THerb. T. [Herb, Inst. Miguel Lillo 26 | (Bu, Ut—11si11b), "h" [La Merced] (Bm), s.n- [18/V/bh] (Ca), s- n. De (Ca); Schreiter 190) [Herb. Inst. Miguel Lillo 32759] (Ug—L912); Sleumer & Vervoorst 2369 (W-—2173085); Ven- Saat — seFsranca a Chaco: Cabrera 3169 (Bt—3535, Ja— » N, S); » Schulz 1470 (N), “172 (N); Ce L. Schulz a @s Venturi 3780 0 (Ge—173b50). Cérdoba: Castellanos s.n. THerb. Mus. Arce: Argent. Cienc. Nat. 31195] (N); Cuezzo 901 (N); Z- Meyer 13726 (N, Sm); Ragonese 6256 [Herb. Inst. st. Bot. Minist. Agr. Nac. sera (N); Ragonese & Piccinini 6394:{Herb. Inst. Bot. Minist, Agr. Nac. 68411] | (N); Ruiz Le Ruiz Leal 1 12098 (Ss); Varela WOk (Ca—165151); Villafafle 29 (Gg——-352669, Ge—353263, N, N), 120 (cg—35326h, N), 33 (N), > 52h (N). Formosa: I. Morel (N). Jujuy: Cabrera L056 (N, N)j Byerdam & Beetle 22317 (Ca— 7h PHYTOLOGIA Vol. 9, no. 2 652218); Schreiter 5013 (Herb. Osten 20754] (Ug); Venturi 5027 (W--159145), 5264 (Herb. Osten 20796] (Ca--3h2620, Ca , Ca—37628, Gg--158394, Ug, , W—1)0785, W—1591)66). La Pampa: Chicchi 26h, in part (N). La Rioja: Afary 15 (Ut--11513b); Castellanos s.n. (Herb. Mus. Argent. Cienc. Nat. 33882] (N); Cuezzo 973 (N)5; Fe Ae Roig s.n. [Ruiz Leal 15453] (Ss). Mendoza: Ruiz ruiz Leal 897 (N), 8825 (N), 8883 (N); Semper s.n. [Ruiz Leal 9833] (N). Rfo Negro: U.S Ss. Expl. Exped. [Wilkes] s.n. (T). Salta: R. M. Aguilar . 286 (Bn, Ca); Barsini 731 (N); Cabrera 300 (N, S)3 Dinelli s [1906] (Bm); Lili Lillo 10871 See Osten 87h] (Ug); Malva: 3 ilalvares or 1 (N), 206 (N), 245 (N), 609 (N, Rf); T. Meyer 12290 .(N); Mo’ Moldenke & Moldenke 1970 (Es, Lg, N, Ot, Sm); | D. ). Rodriguez 1g [Herb. Inst. Miguel Lillo 31698] (N); F. A. Roig s.n. [Ruiz Leal 17880] (R1); Ruiz Leal 12632 (Ss); Torrent s.n. [Laguna El Duraznito, 5-IX- 1945] (N); Venturi 10301 (N); Zabala 116 [Herb. Inst. Miguel Lil- lo 38045] (N). San Luis: Ruiz Leal 11667 (Z). Santa Fé: Kuntze 8. n. [Ceres, Oct. 1892] (W—701722). Santiago del Estero: H. Tes Bartlett 20)16 (Mi, W--1907572), 20428 (Mi); Cuezzo 2302 POR 2325 (Gg—353273, N), 230k (N), 240, (N, ok), 2h92 (N, St), 3302 (8)5 Di Lullo 22 (Ug—6h79); Luna Risso ‘a (N), 83 (N); Valdonado B.185 (N); Pierotti 53 (ca), "h” [El Cimbolar] (Bu, N), "n" (Bm), son, [31-III-19hh] (S), s.n. (1.1V.u4] (W—-193031), sen. [6-L- 4k] (W--193h029); Ruiz Huidobro 3052 (N, Ok), 3108 (N), 3137 (N)5 Venturi 5754 (W--159179); E. W E. Wall s.n. [Inca, 9/11/46) (Br), Se n. [Escolta, 9/11/46] (Ew); Wall & Sparre sen. (Guardia, 9/11/16] (Ew). Tucum4n: Baer 6 (S); Bailetti 231 (Herb. Inst. Miguel Lillo 31690] (N); Bruch Son. [Valle de Tafé, , 1908] (N); JOrgensen 86 [Herb, Osten 1190] (U (Ug); Lillo 1873 [Herb . Osten 8,76] (Ue), 2022 (Herb. Inst. Miguel Lillo 31712; Herb. Osten 8475] (N, Ug), 18286 (Herb. Osten 8454] (Ug), 16128 [Herb. Osten 850] (Ug)s Te Meyer 309) (N), 3368 (N, N), 4268 (Mi, a Moldenke & Moldenke 19717 (N), 19720 (B, Es, F, Fy, Lg, Lm, “Mr, N, No, Ot, Rs, S, Sm, Ss), 19753 (Es, Es, le, N); Monet ti ne [Herb. Inst. " yiguel Lillo 31692] (N), 301, (Herb. Inst. Miguel Lillo 31721] (N), s-M- Herb. Inst. Miguel Lillo 31696] (Du—317602); Moretti 2113 [He ferb. Inst. Miguel Lillo 3169] (Mv); O'Donell 1,600 (Ny); OL Olea ah (Ca-- 165410), 190 (Ca—16L67h); Peirano 9161 [Herb. Inst. Miguel Lillo 32894] (N); Schreiter 190 Herb. Inst. Miguel Lillo 32759] (N, N), 2358 [Herb. Inst. Miguel Lillo 32760] (N), 6522 [Herb. Osten 22968] (Ug), 811) (Herb. Inst. Miguel Lillo 32755] (Ug--L938), 8116 [Herb. Inst. Miguel Lillo 3275h] (N), sen. [Tucumén, Aug. 12, 1917; Herb. Osten 12183] (Ug), s.n. [Taft Viejo, July 8, 1923; Herb. Inst. Miguel Lillo 32758] (N), s.n. [Tapia —cadillal, Herb. Osten 12181] (Ug), s.n. [Cadillal; Herb. Osten 12182] (Ug), s-n- em 1963 Moldenke, Monograph of Verbena 75 ( rechica del Chorro; Herb. Osten 12198] (Ug); A. a. Schulz 2885 a: 2889 (N); Scolnik, Araque Molina, & Barkley 21 21Tu005 (N); c. C. Skottsberg s.n. [Est. Agric. Exp. Puna aati! (Go); “Terribile 162 (N), 269 (N), 325 (N), 338 (N); L. A. Varela s.n. [La Ramada, 8.111.19uL] (N); Venturi 4, THerb. Inst. Miguel guel Lillo 31729] (au, N, W—1591207), lb [Herb. Osten 17255] (Ug), 102 [Herb. Inst. el Lillo 31730; Herb. Osten 17253] (Du—372h07, N, Ug, W— 1591236), 1930 (Herb. Inst. Miguel Lillo 31732; Herb. "Osten ae (N, Ug, W—15912h6), 2303 (W—1591251), 2838 (H-1591269), EO 3180 (Herb. Osten 17252] (Ug, W—1591275), 3579 (Ca—3h2571, 1343300), 4762 (S, W—1591bhh); E. Villa Ta 523 (N), 527 tcp 55382, N), Su (N); Wall & Sparre s.n. [Taff Vieja, 19/11/L6] (Bw). Prov ince undetermined: Friedmann s.n. [1923] (Ba); Hauman s.n, [Serra de Balearca, Nov. 1922] (Br); Moreno s.n. [Patagon. 50/3°, a 4] (MN). CULTIV. VATED: Argentina: Vervoorst 93 (N)- Dominican public: B. Augusto 290 (N). England: Nelmes es 1113 (Ba); jellies Se ne [Cambridge Botanic ic Garden, July 1931] (Ba, Ba). Puerto Rico: E. E. Barker s.n, [Rio Piedras, March 15, 1920] (Ba). LOCALITY OF ee UNDESIGNATED: Herb. A. Jussieu s.n. (N); Tweedie s.n VERBENA DISSECTA f. yest Moldenke, Phytolo gia 2: 22, 1948. Bibliography: Moldenk , Phytolog ia 2: 422. 198; Moldenke, Castanea 13: 119. 198; Meldenke, Known Geogr. D Distrib. At ' hte aad sastica oe typical form of the species in having white corollas. The type of the form was collected by Santiago Venturi (no. 7068) in a prado at Balcozna, dept. Del Alto, Catamarca, argen- tina, at an altitude of 1250 mater, on January 18, 1928, and is deposited in the herbarium of the California Academy of Sciences at San Francisco. The form has heat collected at altitudes of 2 to 1250 meters, gr in dry soil and on sandy shores, flowering in uary and. September. It is said by E. E. Barker to be cultivated along with the typical form in Puerto Rico gardens, It has been misidentified and distributed in herbaria under the name V. lac iata var, albida Herter. In all, 5 herbarium sper IOS a! the type, and ) mount- ed photographs have been exam Citations: URUGUAY: Herter “1805 {Herb . Herter 96556] (Du-- 373695, N, Z). ARGENTINA: Catenaroas hoa 7068 (F--photo i type, Gg--161),52—-type, N--photo —photo of type, Z— photo og 16s Hn asearnteg 5018 SPnz-iso1hs3). XVERBENA DISSOLUTA Moldenke, Phytologia 5: 133. 1955. 76 PHYTOLOGIA Vol. 9, no. 2 ynonymy: Verbena erinoides x phlogiphlora Schnack, Anal. In- st. Fitotéc. Sta. Catalina 4: 20. 192. Glandularia laciniata x phlogiphlora Schnack & Covas, Darwiniana 7: 73 & 75. 195. Verbe- na laciniata x phlogiflora Derinat ex Moldenke, Phytologia 3: 67, nom, md. 1951. Bibliography: Schnack, Anal. Inst. Fitotéc. Sta. Catalina : 20. 1942; Schnack & Covas, Darwini ana 6: 472 (1944) and 7: 73 & 75. 1945; Moldenke, Phytologia 3: 67 (1951) and 5: 133. «19955 Moldenke, scene Abstr. 30: 1093. 1956; Moldenke, Am. 59: 36. 19985 aang Résumé 223, 296, 372, 376, & "LT. 19533 &sumé Suppl. 1960. s hybrid was originally aerate rm Schnack and Covas, who produced it artificially in Argentina in 1945, as being a cross between V. laciniata (L.) Briq. and Vv. phlogiflora Cham., and was so regarded by me in my discussion of f hybridity in the Verbenace- ae (1958). However, it now appears that what Schnack & Covas re~- garded as ve laciniata is actually V. tenuisecta Briq., so the hybrid is actually one between V, tenuisecta and V. phlogiflora. The two parental species occur rather commonly together in Para- guay, Uruguay, three states of Brazil, and at least four prov- inces of Argentina, so natural examples of this hybrid may be ex- pected there. It should certai have considerable horticultural merit, and so deserves further study and experimentation. VERBENA DOMINGENSIS Urb., Symb. Ant. 5: 18h. Synonymy: Verbena officinalis gene: ex an in Leén & Alain, Fl. Cuba h: 281, in syn. 1957 [not V. officinalis Cuevas, 1930, nor L., 1753, nor Lour., 1845, nor Wats., 1942]. Verbena domin- gense aed in herb. Bibliography: Urb., Symb, ew oa aed 1908; Prain, Ind. Kew. Suppl. 4 SOhs. 1913; Perry, 0. Bot, Gard. 20: 262, 263, & 355. 19333 Moldenke, mdlay at mag List Invalid Names 16. 190; Known ie 189, ye z 2 525, “SShy 57, ‘as, “ke, aus, "651, & 652. 1948; Moldenke, Castan be e, Known cong. pistrib. Verbenac., [ed ars mee ie 19h; Moldenke, Alph. List Cit. 3: 880 & 927 (3199 sat in 78 1188, 1256—1359, & 1289. 199; Alain in ta & Alain, Fl. Cu ns ae 1957; Moldenke, Résumé 53, 57, 371, & Erect or sprawling perennial shrubby herb, 30--60 cm Saree arc ing, scrambling, or even scandent and to 1m. long; roots branch- oa producing very many rootlets at the apex; stems numerous from te=stri . oming purp only a few leaves; se glabrous, mostly producing 3 spikes at their apex; principal internodes many times longer than the 1963 Moldenke, Monograph of Verbena 77 adjacent leaves; petioles 1—2 mm. long; leaf-blade s varying from oval-elliptic cr otieipee lane ont ones ) re Tangenlaie or linear (upper ones), 0.7—-l) cm. long, 3--5 mm. wide, obtus acute at the apex, narrowe ed and decurrent Tots the petiole at gre base, sparsely c crenate or incised-crenate along eee with 2—l teeth on each ma margin, sometimes entire, scabro eater with basally inflated hairs, short-pilose beneath bepects)9 on the veins, the upper leaves decreasing in size, sparsely c crenulate or a membrane, the rim short—dentate, the teeth acute; cor- ° lue or dark-blue to purple-violet, purple, purple-red, lavender, or pink, scarcely ) mm. long, the tube cy- lo the other tooth-like and wee Bo lower fruiting-calyxes times shorter, upper ones one-half or one-third as pe as ng tween them; cocci brown, scarce ober 1 mm. long, pe ae on the Nee densely white-striolate with aised striae on mner fac cr @o a The spe ed on a collection by Baron Henrik Franz Alexander Eggers (no, 1828) from among limes tone rocks on the cality in the Dominican Republic, Soh deposited in the herbarium of the Botanisches Museum at Berlin. Urban says "Alia species hu- jus generis in Sto. Domingo sponte crescens, V. officinalis L., caulibus subsimplicibus, foliis pluries majoribus, pinnatifidis v. inferne pinnatipartitis, floribus majoribus: in Valle nuevo ultra sylvam umbrosam in paludosis 1970 ms ge aeeets n. 2175% herb to L Holdridge describes the species as 1m. long, Scandent", whil e Howards say that it he an hing plant in marsh", a "sprawling plant mai s clumps", a "sp pine forests, waste ground, eth abtot fields, sugare ds, on Eocene limestone shin t the edge of brooks, and in fis- sures of coral rock, none ter of 500 to 2500 meters, flower- ing and fruiting from “fink Jan Ekman describes the species as a mweed in geet in Pin- Leo it is "verbena", a applied to —* mae movetenc of the 78 P2704 66-24% Vol. 9, no, 2 identified and distributed under the names V. officinalis L., V. scabra Vahl, and as "Labiata sp, subaphylla", Britton, Britton, & Shafer 292 is a mixture with Stachytarpheta jamaicensis (L.) Vahl. Rugel states that his no. 121, cited below, is the same as his no. 856, not as yet seen by me. _ ay should be noted here that the V. officinalis of Cuevas is Stachytarpheta jamaicensis (L.) Vahl, - that of Watson is Verbena menthaefolia Benth,, while that of Loureiro is V. officinalis L- Perry (1933) Si Sag V. domingensis to syn onymy under V. offic inalis L., saying: On the whole, the specimens from Santo Domin- go and Cuba differ from the typical V. officinalis in their slen- der and more elongate habit; the inflorescence is s cely as glandular, the flowers are smaller and the nutlets ie do not xc V. officinalis. Since many of the specimens are rather poor, it appears probable that oot may very well represent an impoverish- ed condition. Urban himself was somewhat uncertain of ae status of his species as a “appended the following note in a pub- lication: 'An re a formis v. officinalis L. ey igeet To me the species 1 3oke 3 more like Herter's V. gractlescens 7 from as pointed out by Perry, is not vil all typical -- its many stems from a basal rosette of leaves cause it to resemble erehthy Ve Vv. halei Smali; si aaa sheets of ile collection have the typical more spraw rawling erry ise he railowing 5 additional eS not as yet seen by me: CUBA: Havana: Curti Curtiss 677 (D, E, G). Province undeter- — C. Wright 3658 (G). HISPANIOLA: athe E. C. Leonard 3939 In all, 77 herbarium specimens, including type material of all the names ’ involved, and 3 mounted photographs have been examined y me, Citations: CUBA: Havana: Alain A.785 (N); C. F. Baker 2591 (N, Po—63884); Baker, Tracy, & Hasselbring s.n. (Es—-3090); Boldo 94 (Q)5 Britton & Cowell 10326 (N); Curtiss 677 (Cm, Es, Es, N, Vt, W--522300); Ekman 165 (S), 61h (S); Leén 131 (Ha), 6862 (Ha, N), sen. [21-l-1921] (Se—30147, V Vi); Leén & & Edmund s.n, [Playa de M » Abril 21, 1921] (N); Shafer 61 (Cm, Es). Las Villas: Fernando 297 (Ha, N). Matanzas: Britton, Britton, & Shafer 292, in part (a, Es); Rugel 121 (Bm, Cc). ranaog- a Acufia, Alonso, & Pina 18801 (Es); Ekman 7960 (N, S). Pinar d 1 Rio: Ekman 10381 (S), 11196 (s). Province undetermined: c. begsiee 98 [18655 Herd. Sauvalle 1744] (Hv), 3658 (Pa, W--71967). HISPANIOLA: Dominican 1963 Moldenke, Monograph of Verbena 79 Republic: H. A. Allard 12468 (W—195595h), 11838 (N, S, W— 1958659); Chardon 34 (Ba, N, N); Eegers 1828 (C—cotype, Le-- Pp Cc 1711890); Fuertes 1771 (N), 1856 (N); R. A. Howard 12180 (N, N); Hovard & Howard 9056 (N, W--2110920), 9083 (N, S, W--2110930), (W—-1957796) . Haiti: he ee Jr. sn. [July 27, 19h] (W— 1881388); Ekman H.1560 (S, W—1)118hL), H.1569 (s, petri xg H.9314 (S); Holdridge ridge 83h (Gg--316098, Mi, N, W--188021h); E. C. Leonard 3939 (N, W--1076321). CULTIVATED: suena Questel 1686 (W—1781452). VERBENA DUSENII Moldenke, Phytologia 2: 422. 1948, Bibliography: Moldenke » Phytologia 2: 22. "19d; Moldenke, nst. Par 15. 1955; Angely, Fl. Par 12 (1957) and 12: 17. 1958; Moldenke, Résumé 110 & & Wh. "toe8y ane gely, Fl. Paran. 16: 78 (1960) and 17: 6. 1961 Apparently a rather tall erec 3 stems and bran sharply tetragonal, preg puberulent or eventually glabrescent on the larger parts e upper portions often decidedly purplish twiggy; principal Ekecwdes 2--9.5 cm. long; nodes ate; ou te een beneat .5--3.5 cm. long, 1-—- » + fid-dissected, the lobes irregularly incised, acute at the apex, he lower ones widely divergent, v ly oresce: spicate, terminal any ah the upperm subeapitate-flatte ned age anthesis, the aeroe port tion later elongating to 7 am., dense, showy; peduncles rather stout- ish, tetr ragonal, mostly purpli lish, 6-10 cm ag tae rather sparse- i 3 trigillose with reflexed whitish haixs; b actlets conspicu- US, green, rather broadly gs » o--8 mm. ga apiculate, 1.5--2 mm. long, purple; corolla showy, red * rogeeg its tube about 15 mm. long, puberulent above the calyx, 80 Pulb?: f0:5,0 0: 3-A Vol. 9, no. 2 O--13 mm. wide, very sparsely puberulent on the outside, the lobes deeply cordate-notched at the apex; anthers glandular- appendaged, The type of this very showy species was oor by Per Karl Hjalmar Dusén (no. 7108) -—- in whose honor it is _- rather swampy ground at Pinhaes » Parana, Brazil, on EEA tober 29, 1908, and is deposited in the herbarium of the Naturhistoriska Riksmuseum at Stockholm. The species is known thus far only from and has been collected in anthesis only in October. It has been misidentified in herbaria as V. pinnatisecta Schau. In all, 9 herbarium specimens, including the type, and mounted photographs have been examined by me. Citations: BRAZIL: Parané: ape 7108 (Ca SoN688-—tsotypey es photo of type, N--isot; N--photo of t; ype, Si--p of type, W—11,81768—isotype, 2 oaks of gees Ett (N, ete Hatschbach 1519 (N), 2836 ae 3135 (2). VERBENA EHRENBERGIANA Schau, in A. DC., Prodr. ll: 548. 18h7. Synonymy: Verbena ehrenbergiana DC. ex Moldenke, Résumé 363, in syn. 1959. we eee. ste Bibliography: Schau. in A. ont Prodr. <4 Su8 & 555. 18175 Hak in — i; wa ere negra (3a): hate 1894, vac 1178. 1895; Mo. Bot. Gard. ee 2h6, Sata, 2 259, "267-268. & 36. 1933; 1 teadece Known at Distrib.” erbenac, 3 [ed. Lyi de 2B, & lol. rli2g so vane Bot. 2: 57. 1948; Mold er. era Ver = 26, 32, & 198. 199; . caeke, alate ist Cit. 3: “687, 7 “753, » S42, 891, 958, & 363 (1919) and tea soa = “ichéen- ke, Résuné a & 471. 1959; Lewis & Oliver, Am. Bot. 8: 639. 1961. erect, tetrag onal, hirsute, branched in paniculate fashion; leaves decussate-opposite, somewhat ovate, --10 cm. eng somewhat paler and prominently veined beneath, attenuate at @ apex, cuneate at the base and narrowed into the margined pet- ieee trifid, strigillose above, hirtellous (especially on the venation) beneath, the segments. or lobes small, lanceolate-oblong, is ta long as the oe ne or the hse at the apex, ciliate; flowers Ae calyx short, about 1.5 mm. mm » convex on sharply at right angles, almost smooth; chromosome number: n = lh. The type of this species was collected by Carl August ” threnberg 1963 Moldenke, Monograph of Verbena 81 (no. 713) near Los Reyes, in the state of México. Mexico, and was deposited in the herbarium of the Botanisches Museum at Berlin, now destroyed. Schauer (187) tentatively suggests that V. yatha Lehm. may belong in the synonymy of this species, but actu tually os two species are very distinct. He says, further: "Species per e quidem satis insignis, sed quoad habitum fere V, Carolinianae hireutioris forman Schizophyllam exhibens. Spicae in apice caulis au s struentes, pluripollicares, gracil- Takase. floribus suboppositis perexiguis. Calyx 3/l lin. longus, US » bo calyce vix longiore." proaches V, carolina, It ee however, in its tripartite leaves, smaller flowers, d fruits. The collection from Saltillo (Palmer 2037] is much more sae hirsute on all parts, but ap- parently is conspecific. Schauer places the species in his Section Verbenaca, Subsec- tion aeprestachyae. It has been found in moist places, damp riv- eds » and densely wooded waterways, from 1600 to 2025 meters altitude, flower in February, July Au n ti in J and August. The Muellers report it is "rather com- ns t distributed in aeasiaes under the names V. caro olina L. polystachya H.B On the other sale the Kellerman 5825, Ttrckheim 8)2 & IT .651, pase L. V. Velasco 8999, distributed as as V. ehrenbergiana, are ac- carolina L. In regard to the occurrence of V. ehrenbergiana in Arizona, C. T. Mason, Jr., in a letter to me dated January 26, 1955, says: "I have checked our herbarium for Verbena ehrenbergiana, and we do not have any san guaitatives from Arizona. In fact, we have only two specimens, both of them from Mexico." Miss davey cites the following 13 specimens and photographs not as yet seen by me: MEXICO: Coahuila: Edw. Palmer 2037 (G). México: C. A. Ehrenberg 713 (E—photo of type). Nuevo Leén: Pringle 1948 (D, E, F, G). San Luis Potosf: Orcutt 54123 (BE); C- A. Purp Purpus 5451 (E, F, G); Seler 722 (G). Vera a Cruz: Ervendb 153 (D, G). What she cites as "Seler 722" is unquestionably Seler & Seler 722. eon all, 37 herbarium specimens and 2 mounted photographs, in- cluding type or pootetrpe material of all the names involved, examin: have been od Citations: ARIZONA: “county. wee Stalmach 1 198 (Au). rca Federal District: arséne 8829, in part (B rt (B). Hidalgo: M. - Edwards 757 (Au); G. L. Fisher 3762 (W—1725L65), b6169 (W— Tease sen. [Jacala, Aug. 12, 1937] (N); Lietmann 11335 (¥#— 1315096). México: C. A. Ehrenberg 713 [Macbride photos 17u1h] (Kr--photo of type, N—photo of type). Michoac4n: Arséne 8829, in part (N). Nuevo Leén: Mueller & Mueller 156 (Au, Me, . Me, Me), 82 PHYTOLOGIA Vol. 9, no. 2 1132 (Au, Me, Me, Mi); Pringle 198 (Br, C, Ca--10822, Me, a Pa, | 5, Vt, Ww-56 190); M. Taylor 238 (Au). ris bla: F. Salazar s.n. [Patmatlan, 12 July 1913] (Me, W—1169860). san Luis Sater? es A. Purpus 5451 (Ca—156750, N, W--l63961); Seler & Seler 722 (W— 1323087). Tamaulipas: Stanford, Retherford, & Northeraft 891 (Du- 288695, N, Se--69700). Vera Cr aloe aseedbers 153 (Gb). LOCALITY a COLLECTION UNDETERMINED: Ravn s.n. {N. Amer. re) (Cp); Herb. sen. (N. Amer.) (Cp). ~ VERBENA ELEGANS H.B.K., Nov. Gen. & Sp; Pl. 2: 273. 1618. Synonymy: Verbena moranensis Willd. ex Spreng. in L., Syst. Vege, ed. 16, 2: 750. 1825. Verbena elegans Humb. ex Spreng. in L., Syst. Veg., ed. 16, 2: 750. 162 1825. Verbena elegans Hort. ex Walp., Repert. Syst. Bot. 4: 33, nom. nud. 1645. Verb Verbena elegans Kunth ex Schau. in A. DC., Prodr. 11: 55). 1847. Verbena lamberti Ker ex Schau. in A. DC., Frodr. 11: 554. 1847. Verbena moranensis H. & B. ex Schau. in A. DC., Prodr. 11: 55h, in syn. syn. 187. Ver- bena canadensis subsp. elegans Thell., Fl. Advent. Montpel. 428. Montpel. 428, in syn. 1912. Verbena lambertii Ker ex Moidenke, Résumé 368, in syn. 1959 [not V. lambertii Sims, 1903). Bibliograpiy: age te Nov. Gen. & Sp. Pl. 2: 273. 1618; » Syst dl., Reg. 13: pl. 1102. 1827; Hook. & Arn., Bot. Beech. Voy. 305 (18385 and 48). om megs Repert. Syst. a aM a 18h5; chau. in A, DC. - 11: 55h. 1847; Brig. Pe othe & Prantl, Nat. Pflanzenfam. fer (3a): 148. 1894; Jacks., raf Kew, 2: 1178 & 1179. 1895; Perry, Ann. Mo. Bot. Gard. 20: 248, 311, 319--320, & relim, ! 5& at Moldenke, Alph. List Invalid Names Ee se 48. 192; Moldenke, own Geogr. Distrib. Verbenac., [ed. 1], 7h, & 101. 19h2; Moldenke, Alph. List Cit. 1: 53” & 178. 18455 *Moldenke, Alph. List Invalid Names sete sa 23 & 25. 1947; Mol denke, Phytologia 2: 331 & 347. 19h7; . & A. L. Moldenke, Pl. Life 2: hh. 198; Moldenke, Alph. List cit. 72! 393s 467, 573, 60h, 617, & 639 (298) 3: 679, 685, 696, 709, 717, Thh, 752, 767, 830, & £33 Sid ), and kz: 1074, 1076, 1119, l2 oes. 1295; & 1303. 199; Moldenke, P gia 3: 132 & ° tect Mold denke, Known Geogr. Distrib. vanveiaess fed. 2], 33, 16h, & 198. 19h9; Moldenke, Sp. Subsp. Cont. Mold. - 4: 220 - 1951; Moldenke, mm. Mold. 4, (1951) and h9 Spec. 3. 1954; Moldenke, Phytologia 5: 22. 195h3 rte! ae a ae 20; 5," -f raat S08 19 2 1- 3¢.8, 10, .15,.31, We (1962, ve me (i968), and 5 Ne ‘ or prost a neg: Beek stems prostrate-decumbent, procumbent, or ont tending to root at the older nodes, more 1963 Moldenke, Monograph of Verbena 83 or less ep eget sparsely hispidulous-hirsute; bran- ches ascending, purplish, the older ones hirtous or hirsut- ulous, the yo ahi ones hispid. or hispidulous; peo short; leaf- blades ovate or lanceolate-ovate, 2--5.7 cm. long, 2-3. wide, incised-pinnatifid or trifid with the divisions less (deeply branches or fascicles leaves in t S, somewhat appressed-hirsute to strigose-hi ous or —hispidulous on both surfaces, the di lanceolate to broadly obl or line ong ar= peor. rather obtuse at the apex, entire to sparsely toothed or unequally incised-pimatifid, subrevolute-margined, the ultimate lobes spec ot entire, acute; venation prominulous beneath; pina Sepeaase: and axillary, solitary, pedunculate, about 2. g cm, during anthesis, fastigiate, peed Bs or fasc i¢le-like, ir ie tec tlaw red but dense and compact, usually not much elon— gating after per ies pay hirsutulous and with a few glandular hairs intermixed, the pedun uncles about 2.5 cm. long; bractlets ecrong late or subulate-lanceolate, varying — half as long as to two- thirds as long as or about equaling ¢ calyx, attenuate ea the apex, glandular-hirsute or eens , ciliate; calyx 6—10 m. long, more or less densely glandular-hispidulous or -hirsute, the rim very unequally 5-fid, the teeth short, unequal, subulate, the posterior ones shorter, the mouth therefore oblique; corolla hy- pocrateriform, showy, from » Violet, violet-rose, deep-pink, o r vivid-pink to magenta-purple, pavenser ae lavender, or blue, its tube about ce as long as puberulent on the nage fei barbate in the throat, in e iit large, wi appe dric, 3 late’ chiefly on She Spper half, the lower part striate; comnis-— sural face muriculate. The type of this very handsome species was collected by Pinter ats oven ota Alexander von Humboldt and Aimé Jacques wero andre Bonpland between Moran and Regla, Hidalgo, Mexico, at altitude er about 2600 meters. The type of V. moranensis _ sae Bs. se, from Moran, deposited in the Willdenow Herbarium Berli. Ver Ruasa canadensis var. ehrenbergii Thell. was probably based on C. Ehrenberg 130, from Mexico, which is cited by Schauer in A. DC., Prodr. 11: 554 (1847) as V. lamberti Ker, a binomial which he credits erroneously to Ker, Bot. Reg. pl. 1102 and of which he Says "Flores et fructus omnino V. Aubletiae, sed paulo majores" ~~ but the Lindl., vole Reg. 13: pl. 1102 illustration is of "V. Verbena camieais subsp. elegans Thell. is based on V. ele- Penny and of Sims are V. canadensis (L.) Britton, . while that of Sprengel is V. lasiostachys Link, and V. lambertii Sims is also 8h PHYTOLOGIA Vol. 9, no. 2 V. canadensis, Verbena lambertii Ker was erroneously reduced to V. canadensis by me in md Résumé 368 (1959), but actually belongs e@ synonymy of Vv. 8 — elegans ee reg pte fields and mountains, gravelly neds dry slopes, « cliffs, steep mountainsides with oaks and pines, and open forested slopes *with Ppine-oak-madrofio associations, from 1000 to 3165 meters altitude. It has been collected in anthesis as "abun mn sun" in Tamaulipas, while ema reports it "uncommon in s oam open sun on wooded slopes rancas" in Nayarit. It is called "showy verbena", and was intro- duced into cultivation in 1827, The L. P. McCann. specimens cited below were grown from seeds secured from Copenhagen as Vv. cana- densis. Bouchon 1,782 is said to have originated in the southern ae States 2 to be subspontaneous in the Botanical Garden at ° The ePrihets 6308 cited below has its pubescence very short. Née 105 is a mixture with V. teucriifolia Mart. & ~~ Herbarium Aertel of 7. elegans has been misidentified and distributed in herbarium under the names V. aubletia L., "V. au- bletia forma", V. bipinnatifida Nutt., V. canadensis (L.) Britt- on, V. ciliata Ben th, V. delticola Small, V. elegans var. asper- ata Perry, V. tenera Spreng., and V. wrightii A. Gra A. Gray. ~~ On the other hand, the Chute pues, G. L. Fisher 3759, H. S- Gentry 2730, Edw. Painer igi, a and Stanford, Retherford, & North- craft craft 566, distributed as ae elegans, are | are all actually var. as~ perata Pe Perry; R. M. King 42h9 is V. cameronensis L. I. Davis; Arsene 1671 is V vs “eiiata Benth.; and C. R. Orcutt 3551 is V. teucriifolia Mart. & Gal. Walpers (185) coe V. elegans in his Section Verbenaca, Subsection Inermes, Group Fi Foliosae, Subgroup Macranthae, econdary Siberaas: Aubletia with ten other species. Schauer (1847) cites Sweet, Brit. Flow. Gard., ser. 2, h: pl. 353 as V. lamberti Ker, but this plate is cited by me as V. canadensis (.-) Britton. He also cites Mendez s.n. from Vallalpando, but com- praecedentam [V. elegans] e et sequentem [V. canadensis], ab i114 foliis minus divisis, f1 » floribus majoribus, ab hfc vero foliis = 4 ry quodammodo accedens, sed éoret 18 magn& jam satis aia Wal- pers says rons V. erinoidi et te corny a prioris varie- tatibus hance adnumerarem si vidiss em specimen ag 1963 Moldenke, Monograph of Verbena 85 The colored illustration by Lindley in Bot. Ronde’ _ pl. 1102 (1827) often cited as V. elegans is actually V. hastata L. Schau- er (187) mentions a "v. lamberti Ker" on the basis of an Ehren- berg specimen from Me xico Perry (1933) comments that V. elegans is "related to V. cana- densis but is easily separable b by its slender more or less pros- eee habit, few-flowered compact spikes, and shorter r calyx- (e) am ’ th, The mu reticulations in Ve elegans are a little coarser than those in the more northern 5 species." She cites the following 12 addition— specimens not as yet seen by me: MEXICO: Hidalgo: Clokey s.n. {Real del Monte, 9 May 1910] (E); Humboldt & Bonpland 4063 (E— photo); Pringle "6908 (D, E, F, G), 11092 (E, F, G); C. A. Purpus Kk. Purpus 1433 (E). Oaxaca: Gandoger s.n, [Tehuantepec, 1906] (E). Vera Crug: Seler 723 (G). Of Eggleston 10813 she says on the United States National Herbarium sheet "aff. V V. Gooddingii Briq." In all, 96 herbarium specimens and 2 2 mounted ya Scr in- cluding type material of most of the names involved, e been S Citetiaaee h ARIZONA: eer Co.: Eggleston io (W--66),875) 5 A. R. Moldenke 146 (Am, Fg). Pima Co.: Barneby 2506 (N). MEXICO: Chihualma: Knobloch 950 (sm), 590, (W—17910)7) - cahaiias Pur- s 4598 (Ca—1hh758). Federal District: E. lyonnet 173 (N, W— SeaSay )- Hidalgo: Clokey 1857 (Ca—882859); M. Te “M. T. Edwards 862 (Au); Humboldt & Bonpland s.n. [inter Moran et Regla; ‘a; Macbride photos 3979a] (Kr--photo of type, N--photo of type); E. 2231 (W--17),8265); Martinez 11 (W—1791051); Matuda 21650 (N), 29558 (Z); H. N. Moldenke 19842 (Mg, No); Moldenke & Moldenke 198L2 (B, Es, F, Fy, Le, Im, Mr, N, Ot, Rs, S, Sm, Ss), 19843 (Es, Le, N, Sn); Pringle 6908 (Br, Ca--10h710, Cm, Io—38754, Me, Me, Me, Mi, N, Po--O300h, S, Vt, W—33K766), 11092 (Cm, N, W— 46047h) 5 nti 1433 rere gan rt Po—63880), Sn. s.n. (Pachuca, May 1905] (Ca--139757); Rose & Hay 5556 (W—395320); R ; Rose, Painter, & Rose 8668 (N, W—52162); Sh Sharp, Hernandez, & Goodman ELS (N); Yates & Wileox 155 (Ak—31225). México: i i. Querétaro: wee & Wal W159 (Sty, Hepa, pie San Luis Co. Mickel 517 Leth es oa: H. S. Gentry 7181 (Ca—9l6136, Mi, N). , Tamaulipas: King 1513 (N); H. W. Viereck 1OL7 (W--1687519) Vera ear eo Je Sharp 1723 (0).~, tate reiiernie mined: C. A. Ehrenberg 130 (vt); Keerl s.n. (Tlapujah 29] (Br); Née 105, in part (Q)3 H. W. Viereck 253 [c. me eB eee # 128). CULTIVATED: France: Bow chon L782, - in part (La). Germany: Golenz s.n. [Schwiebus, 1872] 86 PHYTOLOGIA Vol. 9, no. 2 (B). Maryland: be P. McCann s.n. [9-15-36] (Md, Md). Massachu- setts: Leavitt s 1. (Cambri dge, 13 Oct. 1898] (Rf). Michigan: C. F. Wheeler s.n. sat (Po--63888). New York: H. N. Moldenke 18462 (Ec, Jr, N, N, Si). Sweden: A. Andersson 5 (Ew); E. Wall 5 (1902] (Ew). Switzerland: Bernet s.n. (S). IA ELEGANS var. ASPERATA Perry, Ann. Mo. Bot. Gard. 20: 319- 320. 1933. Synonymy: Verbena elegans asperata Perry ex ore hae in Chit- tenden, Roy. Hort. Soc. Dict. Gard. 6: 2209. Bibliography: Perry, Ann. Mo. Bot. Gard. es m; ee & 355. 19333 H. S. Gentry, Rio Mayo Pl. [Carnegie Inst. Wash. Publ. 527:] 43, 222, & he 1942; Moldenke, Known Geogr. seaweah: Ver- benac., fed. 1], 18, 7, 80, & 101. 19h2; Moldenke, Known Geogr. Distrib. Verbenac ag 4 uppl. 1: 2. 19h;3; Moldenke, Bot. Gaz. 106: a ta E olog : yl. 32-683, 73; 768, “Ta 09, 807, 847, 879, 933, : 963 (19 and hj: a2" 1071, 1099, 116h, 1170, 1175, 1233, 125, hae 1255, & 1291. i9h9; tents in Chittenden, ee Hort. Soc. Dict. Gard. hh: 2209 & 2211. 1951; Suppl. 2: 3 & 6. 1960; Lewis & Oliver, Am. Journ. Bo t. UB: 639-- 641. 1961; Moldenke, Résumé Suppl. 3: "10, we , & & 37. Tlustrations: Lewis & Oliver, Am. Journ. Bot. is: "60. 1961. 355) at San sntedta: Coahuila, Mexico, on August 31, 18h8, and is deposited in the herbarium of the lieaourt Botanical Garden s tain slopes with cut-over oak woods, rocky slopes ats serts, and pine~oak: fia openly forested slopes, at altitudes of 330 to 3930 meters, flowering from February to October and fruiting from April to October names reported for this plant are "alfombrillo", 1963 Moldenke, Monograph of Verbena 87 "oradilla", and "verbena". Gentry states that it is a characte istic plant of arroyo margins, including meadows in pine forests, blooming in summer. The Mue liers found it aaa Bow i woods" in Nuevo Leén, Stanford, Retherford, & spark ote R a mere it on mountains and in canyons with uxuriant vegetation, and on der found it "abundant on are rocky slopes" in Nuevo Leén. icti is "uncommon" on the Sonora-Chihuahua boundary. Pennington af- firms that "a tea is Breperae from this plant for stomach disor- ders and for catarr r the probable et location of collection of Stanford, Rethersords & Northcraft 566, which came ete 18 km. west of Con- ion del Oro on the Coahuila-Zacatecas border, see my note atae V. canescens H.B.K. The specimens of Weaver 585, cited be- inp iS ist "same as 562", but I have not as yet seen the’ latter sollect Material of this “variety has been much confused, misidentified and distributed in herbaria under the names V. ambrosifolia Rydb., V. ambrosiifolia ae V. aubletia L., V. "Besa teas Nutt., V. canadensis Britton, VY. ciliata Benth., V. delticola Small, V- elegans H.B.K., Ve erinoides Lam., V. erinoides Willd., V. hy- brida Voss, V. multifida Ruiz & Pav., V. pulchella Hort., Fa. 7 Pulchella Sweet, V. remota Benth., V. tenera Spreng., V. venosa GEIL. & Hook., V. wrightii A. Gray, Smibtelmorthés pulctells Meissn., and Shuttleworthia pulchella Meisn. On the other hand, the H. S. Gentry 7181, distributed as V. elegans var. alparatas is typical ve elegans HeBeKe5 Maysilles ree a is V. ciliata Benth.; and Graber 103 is V. delticola Small. variety was apparently intro duced - 7s y cultivation in 1883." The Bouchon 782 cited below is said to represent plants that were both cultivated and subspontaneous oth the Botanical seed which he Garden « oe Bordeaux, France. Dr. Bailey reports that Planted at Ithaca, New York, on April 6 produced plants that owered on August 10. His August 28, 1925 specimen be tation: "gland on back of anther — LHB." he Herb. Prager 18632 does not have any notation on its label sntcating that it came fran Yorks visa material, but I am assuming that it did. Basie 23 was grown from seeds obtained from taster, while no. grown from bettie Sposa from Cluj, Romania. Se ccenntiaione 88 PeEETOLO GTA Vol. 9, no. 2 tedo s.n. came from Kew, England, those for G. V. Nash s.n. [7 5S. 1898] from Geneva, Switzerland, and those for N. Y. Bot. Gard. Cult. Pl. 628 tron mablaess 1s from Cambridge, 23187 fr from Ham- burg, 23352 from Madrid, and 33298 from St. Louis. The seeds are handled by Haage & Schmidt in Erfurt. The Hort. Huber 753 speci- men cited below bears a notation "Verbena pulchella H Hort, est Vv. erinoides Lam. DC. p. 552, no. 64 Ct)" Perry (1933) cites the following 18 additional specimens not s yet seen by me: MEXICO: Chihuahua: Edw, Palmer 295 (G). Coahu- way 2 J. Gregg 355 (E--type); Edw. Palmer er 1052 (G)3 “Purpus 497) (E, F, G). Durango: Edw. Palmer 191 (E, G). San Luis Potosf: Edw. Mite 51 (E, F, G)3 Parry & Palmer 720 (D, E, F); Schaffner 716 (G). Sonora: M. A. Crawford S.n. (Hermosillo, 1888} (G); Lobel Uu5 (G), bh6 (G). She says "This variety, as compared wit. th the species, is stouter and more erect with coarser pubescenc me larger spikes. Palmer 1gl1 differs from the other collections in having very shallowly Tobed ovate leaves. =e & Palmer 720 shows much variation in pubescence, som s being scabrous, others scarcely so at all. Purpus s 97h is oe the coarse pemeorenn® but appears more closely allied here than elsewhere." n all, 153 herbarium specimens and 2 mounted photographs have been examined by me. 7 ns: TEXAS: tora | Co.: Lewis & Oliver x ane (Nb). Hid- aardeetiaane Futian dente 8 ate teens. Knobloch 496 (Z), 1174 (Mi), 1288 (Mi), 1305 (Mi); Edw, Palmer z Be (W—56166); Pennington 68 (Bm), 369 (Bm). Coahuila: Kenoyer & Crum 2692 (Mi); E. W. Nelson 3896 (W—266885) 5 Edw, Palmer 1052 a, ate Purpus 97h (Ca—-153292, W— 842191). Durango: Edw. Palmer 191 (Ca—-13 39760, N, W--571212). Hidalgo: F. A. Barkley 17M009 (Au--123219, Au—~123229, N), 17006 (Au—1700)8, N); V. H. Chase Chase 7116 (Fs, Ld, N, N, Ur, Ur, Ur)s Chute M.31 (Mi); ae Z. Davis 205 (N), sen. [Jacala, June 191] (Au 172667); G. L. Fisher 3759 (W—1725462), 45198 (Ew, W—189h77h), 46151 (En, Gg--353139), 16171 (W—1889831), s.n. [Jacala, Aug. 10, 1937] (Fs, N); Lundell & Lundell 12392 (N, Rf, Rf, Rf, W--1926600); Moore & Wood 3922 22 (Ba); | Rowell 2 2020 20 (N). Nayarit: W. H. Lewis 5378 (Nb). Nuevo Leén: M. T, Edwards 129 (Du--278381); Meyer & Rogers 2579 (N), 3022 (N); Mw Mueller & & ee 328 (Mi); Re he he Schneider 9 927 (Mi); M Me Taylor 129 129 (Ca—665000); J.N. Weaver 585 (N). Oaxaca az M,C C. Johnston 53-691 (Au--122102, St). San Luis ; Po- tosf: MeVaugh | 12274 (Mi); Edw. Palmer 51 (Bm, Me, N, W—397598)3 Parry & Palmer 720 (Io, Pa, W--56186); 3 F. W. Pennell 17879 (Mi, N, x, oe Es Rodehgnes-21 17 Fe Ne, Ras SrreRnrOnee) Rut- 1963 Moldenke, Monograph of Verbena 89 6169 (Ge, N), 7181 (W--1945195). Sonora: Branson s-n. [August 15, 1959] (St); H. S. Gentry 670m (Fs, Mi); Shreve 59L6 (Fs); S. Se White 3085 (Mi), 3964 (Mi), 4178 (W--2132338); Wiggins 11673 (Du- 3600). Tamaulipas: L. I. Davis s.n. [Feb. 16, 1960] (Au— 181592); P. S. Martin 551 (Ii); Rozynski 276 (Fs, Mi), 353 (wi), 691 (Mi, N); Stanford, Lauber, & Taylor 2486 (Ca—-987776, Gg— 381242, N, N, W—-2216750); Stanford, Retherford, & Northcraft 678 (Du—286 303, N, Se--70560, Tu--10059), 1023 (Du--287085, N, Se— 70645, Tu--16242); H. W. Viereck 194 (W--1637114). Zacatecas: Stanford, Retherford, & Northcraft 566 (Ca—713837, Du—288651, N, Se--69687, Tu—10912). State undetermined: Berlandier 1703 (Du— 166490); Herb. Prager 18632, in part (Gg--31L47). SWEDEN: A. An- dersson s.n. (Sept. 1931] (Gg—-222542). FRANCE: Bouchon 4782, in part (Mg—-125). CULTIVATED: Connecticut: Bevoise s.n. [aug. 193k] (Ba). Cuba: J. G. Jack 8617 (Gg—2378l8, N)- Denmark: Lange 5 -Me (Sept. 15, 1919] (Ba). England: L. H. Bailey s.n. (Aug. 8, 1919] Santiago 6831) (Sg); Nelmes 1151 (Ba). France: Hort. Huber 753 (Bm). Germany: Heiland s-n. rt —314)8); Rehder s.n. [Hort. Bot. Berol., Aug. 30, 1885] (Ba). Massachusetts: L. H. Bailey s.n. {August 6--12, 1929] Qu), s.n, [N. ¥. Bot. Gard. Cult. Pl. 114] (N), sen. [N. Y- Bot. Gard. Cult. Pl. 1115] (N), sen. [N. Y. Bot. Gard. Cult. Pl. 1146] (N), s.n. [7 S. 1898] (N); BR. C. Schneider s.n. [N. Y. Bot. Gard. Cult. Pl, 23187] (N), sen. [N. ¥. Bot. Gard. Cult. Pl. 23352] (N). Sweden: Blom s.n. (Go); B. Wall 5 [26/82] (S)- XVERBENA ENGELMANNII Moldenke, Revist. Sudam. Bot. 4: 18. 1937. Synonymy: Verbena alta foliis urticae, floribus dilute coerul- eis spicatim in Summis caulibus congestis Gronov., Fl. Virg-, ed. 2, h. 1762. Verbena urticaefolio-paniculata Engelm., Am. Journ. Sci. 46: 101, 18h). Verbena paniculato-urticaefolia Engelm., Am. Journ. Sci. h6: 101. 18k. Verbena hastata x urticaefolia Brain- erd, Bull. Vt. Agr. Sta. 187: 236. 1915. Verbena hastata x urti- cifolia Dermen, Cytologia 7: 170. 1936 [not V. hastata x urtici- folia Gates, 1959]. Verbena hastata x urticifolia Blanchard ex Moldenke, Revist. Sudam. Bot. 4: 18, in syn. 1937. Verbena hasta- ta x urticifolia Eggert ex Moldenke, Revist. Sudam. Bot. 4: 18, in syn. 1937. Verbena hastata x urticaefolia Norton ex Moldenke, 90 Et T OL OATS Vol. 9, no. 2 Revist. Sudam. Bot. h: 18, in ee 1937. Verbena urticaefolia x hastata Farwell ex Moldenke evist. Sudam. Bot. : 18, in syn. 1937. Verbena hastata x ee ep Perry ex Moldenke, ” Sipe: denke, Suppl. List Invalid Names 10, in syn. 19h1. Verbena hasta- ta x nntdoabtelie Blanchard ex udldenke, Alph. List ; Invalid Name: Names Suppl. 1: 2h, in syn. 1947. Verbena re x urticaefolia Chase ex Moldenke, Alph. List Invalid Names Suppl. 1: 2, in syn. 19h7. Verbena hastata x urticaefolia Dutton ex ‘Moldenke, *,lph. List In- valid Names Sup Suppl. 1: 2), in syn. 197. Verbena hastata x urticae- folia Peck ex Moldenke, Alph. List Invalid Names Suppl. 1: “2h, in syn. 1947. Verbena hastata x urticifolia Farwell ex Moldenke, Alph. List Invalid Names Suppl. 1: 2h, in syn. 197. Verbena has- tata x urticifolia Pringle ex Moldenke, Alph. List Invalid N. Names Suppl. 1: 2h, in — —— Verbena trti.ci folie x hastata Gates ex ee Alph. List Invalid Names Suppl. 1: 27, in syn. 19h7- ena hastata rg we pesto McCoy ex Moldenke, Am. Midl. Nat. 5 Lz, “in syn. a Verbena hastata x ubiieahtaulié Pammel ex Moldenke, Am, Midl. Nat. 59: 3h6, in syn. 1958. Verbena engel- Résumé Suppl. 3: 1, in syn. 1962. Bibliography: Gronov. wr Virg., od, 2; i. aged Engelm., Am. Journ, Sci. 6: 101. 18; E, Brainerd, Bull. Vt. A . Exp. Sta. 187: 238. 1915; Dermen, Cytologia 7: 170. 1936; Moldenke, Revist. S$ own Geogr. Distrib. Verbenac., 6, a idan, spb, List. rreiia Names {,7,” eS : ae "19424 & Friesner, Proc. Ind. Acad, Sci. 51: 98) a a ee 10h. 1983; Moldenke, Known Geogr. Distrib. rte Suppl. 1: 1, 1943; Deam, Kriebel, Yuncker, & Friesner, 5 36, 438, & sau, Ae: ih 6, x eke Gale} 32 699, 719, 70, Tlsy 773, 790, 792s 793, 803, L, $16, 930, & 9h6 (1919), and bs 1119, Ta38, ee in73, "1175; 1176, ey 1201, 1238, 1251, 1255, 1963 Moldenke, Monograph of Verbena 91 1260, & 1261. 1949; Moldenke, Kn istrib. Verbenac., [ed. 2], 3-5, 12--17, 21, 16h, & age tee “B95 “aes Abdi Phytologia 3: 130 & 131 (1949) and 33 t 31h. 195 1950; » Yuncker ,& Friesner, 93 Fl., pr. 1, 3: 130 (1952) ig 2, 3: 130. 19 This is the common natural hybrid between Verbena hastata L. and V. urticifolia L., occurring wherever the ranges of these two very common species overlap, and with intermediate charac- ters. It resembles V. urtic ifolia in habit, but the spikes are aly mostly contiguous, the corollas small, varying from blue, pale- blue, violet-blue, or tint biae to lilac, EERE light the corollas, and from V. urticifolia by its very sparse fruited spikes, colored | corollas, and narrow-elliptic leaves. It is one of the commonest of all the natural hybrids known cata un 5 although they grow most commonly together or in close proximity, they are so very different in general appearance -— Vv. hastata having very dense-flowered spikes of usually deep-blue flowers and V. urticifolia having very loose and lax distant—flowered spikes of white flowers It is worth noting here that the "Verbena hastata x urtici- folia" of Gates, at least insofar as the herbarium specimens Which he so annotated and which have been seen by me are con- cerned, is xV. illicita Moldenke. The type of xV. engelmannii was collected by Carl Andreas Geyer at Beardstown, Cas was designated V. urticaefolio-paniculata It is deposited in the Torrey Herbarium at the New York Botani- 92 PHYTOLOGIA Vol. 9, no. 2 cal Garden. The Verbena alta foliis urticae, floribus dilue coer uleis spicatim in summis cau. Caulibus congestis. of J. F. Gronovius (1762) seems to be this hybrid, but the pre-Linnean and Linnean synonymy which he gives apply to V. urticifolia L. The type of V. panicula to-urticaefolia was collected by George Enge at Saint Loui issouri, in August, 183. The type of V. hybrida Bicknell was collect ted by Eugene Pintard Bicknell (no. 7351) in Bronx County, New York, and is deposited in the Britton V. hastata x urticaefolia Blanchard was collected by William Hen- ry Blanchard in Ben nington County, Vermont, on August 11, 1902, deposited in the herbarium of the University of Vermont; * that of V. hastata x urticaefolia Chase was gathered by Virginius Heber Chase (no. (no. 3230) in Tazewell County, Illinois, and is deposited in the Britton Herbarium; that of V. hastata x urticaefolia Dut- ton was collected by Dura Lewis pastel at Ee at Leicester, Addison County, Vermont, on August ae 1915, deposited at the University of Vermont; that of V. hast x urticaefolia McCoy was gathered by Scott McCoy (no. doy ie pranvtin County, Indiana, deposited Li was collected by John aig Smith Norton (no. 792, in part) in Riley County, Kansas, and is deposited in the herbarium of Kansas State College at Manhattan, ee with xV. illicita; that of V. hastata x urticaefolia Peck was gathered b: by Morton Eaton Peck at Fal Hardin ium of Iowa State College at Ames; that of V. hastata x urticae- folia Pammel was collected by Louis Hermann Pammel at St. Olaf, Cl County, Iowa, on August 10, 1925, deposited at Iowa State College; that of V. hasta ta x urticifolia a Eggert was gathered by Oliv Atkins Farwell (no. (no, 1426) in Wayne County, Michigan, depos- ited in the herbarium of the Colorado Agricultural and Mechanical College at Fort Collins; that of V. hastata x urticifolia Perry was collected by Marcus Eugene Jones at Grinnell, Poweshiek Coun- s ey Pringle at Charlotte, Chittenden County, Vermont, on July 15, 1887, depbatten at the e University of Vermont; and that of V. ux arl » Wabas 1897, deposited at the University of Illinois. t is possible that the perplexing "Verbena diffusa Willd." ex J. Sm. in Rees, Cycl. a ae 20 He See may also represent this plant, oar he says eiisdios to V. urtici- 1963 Moldenke, Monograph of Verbena 93 folia, with very small, somewhat purple, flowers". Rafinesque's Vv. in incarnata may also pelotie here, but until I can see authentic material = is perhaps best a so to dispose of them Collectors have found xV, ngelmannii in low ground and swampy or peaty erie: » on creek eal and bottoms, in marshes, in door- yards and pastures, cow pastures and open ravine pastures, on and san ear out canal s black oak dunes, in clearings ab landings, in marshy fields and ditches, on prairie strips and moist or dry banks, on the banks of field-draining ditc ong roadsides, hes in the vicinity of stockyards, and in low fields associated with Teucrium canadense L., Peat lapathifolia (L.) S. F. Gray, and Carex vulpinoidea Michx., in flower and fruit from July to under the names V. carolina L sea Cheney, ve. hastata var. oblongifolia Nutt., ae officinalis Bag V. riparia Raf., V. scabra Vahl, V. stricta Yent., V. urtic- aefolia L., V. r , V. urticifolia L., V. urticifolia var. riparia Brit- ton, and "Verbena hybrid” said On the other hand, the ee 19083 distributed as xV. engel- s V. montevidensis Spreng.; C. C. Crampton n 369, distrib- uted & as TW urticifolia x hastata, is XV. -—sTlicita Tlicita Moldenke, and the Bogusch 969, similarly distributed, | is V. scabra oats As is to be | expected, collectors' labels record the c names of the parents for this plant, "blue vervain"™ and onatties leaved vervain". Grimes considered his ot cited below, as a hybrid be- tween "V. urticaefolia" and V. stricta, while Eggert identified some of his Specimens as "V. stricta x urticaefolia" and his August 12, 1875, Schneck found the hybrid growing among both parental species in @ dooryard and makes the comment that it "looks like V. urticifolia, but the flowers are more like those of ve. hastata." Shafer also Says that he found his specimens actually growing between both parents. Dean ae his plants as being wide-spreading, "al- most as wide as tall", and states that his no. 1978 is actually only the lower branch of a very large plant. Bicknell describes his plant as as “long and spreading 2 v2 ig surface of bon oer and beng a) within oe puberulent, racem 3 flexuous, spre and drooping, mostly 10 to 12 in. long, greatest spread of pan- icle 22 inches" and "rather more harshiy hairy than V. hastata, leaves with more irregular and —_—!, meeere pm a little Smaller and paler." Ahles found his no. 6581 i a pasture noth parents and V. stricta Vent. Panmel ters to the plant as a weed". G. N. Jones describes his no. 22339 as 2 meters tall; pa i says of his No. 579 "growing hg with ve hastata, but flow- 9h PHYTOLOGIA Vol. 9, no. 2 ers smaller"; Storm reports "plants not numerous, in about half acre of moist — ai ae stream bed." Bicknell 7363a consists of leaves only; th very hairy and peculiar in shape. The Grace 353. an below has very hirsute stems and does not look much like most specimens of this hybrid, Norton 792 is a mix- ture with xV, illicita, while F. A. Smith s.n. [July 1 1896] is a xture with V. arkeoifalie. The Fp en 6450 collection is Bers close to typical V. hastata. His nos. 18613, 18618, 18619, and 18620 were all growing together, he says. Of these, 18618 is typ- ical V. artic’ folie, 18613 is typical V. hastata, 18620 1 looks mach like hastata and had "blue" Comets. whi , While 18619 has more distant sae or "pinkish" flowers. The last-mentioned was re- garded by him as F, material. gee records the hybrid from Craw- ford and Meade Counties, Kansas, e Deam and his peg Sito report it from Fulton and Greene chntida, Indiana. It was ap ently introduced into cultivation in abou Ahles, Bell, & Radford (1958) cite a Collector undesignated s.n. [bank of New River, Solitude P. 0.] from Ashe County, North Carolina, The Walpole Island mentioned below is actually in Lmb- oO. The sp ) Wels cited below, bears a notation "W. F. K. Drawn, Mrs. Dick- 1931", and is’ apparently the basis of an illustration not es is ae seen by me. Iltis records the hybrid from Lafayette Coun- ‘click! until this summer, when I found the single plant of no. 239m The other one you saw, no. 1692W, was also a single plant and all I could find that day. But I had been looking for it all that day because I had found only the single plant, no. 1522W.-- on an earlier trip. You see, the very pale blue — not ot Dap ee at all -- more sky blue color of each of these caught my eye When I got this very rough 1522W I surely thought I had a species that was absolutely new to me (I mean, neither hastata nor es ifolia, and I never thought of the fgbrt rid between the two). after I got 1692W and then this year 23957, with the same ite blue flower and the typical urticifolia inflorescence in the lat- ter number, I thought 1522W was just another form of the same cross. Now. your special comment about it seems very pertinent. Ve hastata tata grows in low places, while this big tall rank apogee was was picked off the top of the bank, so that as I drove peng the car, it towered its whole 5 feet (or about that) above me. The flower color was striking. I wish I could have Pcs ab of it. I looked for it again and found this 2395W this summer not too far from its location. They surely are only single sp ecimens -" In all, 197 herbarium specimens, including the types of almost 1963 Moldenke, Monograph of Verbena 95 all the names involved, have been examined by PR secre ONTARIO: * Walpole Island: Gaiser 1692 (Gp, Gp, Gp, » 2395W (Gp, Mm). VERMONT: Addison Co.: Dutton sen. [Leicester, ane 7, 1915] (Vt). Bennington Co.: W. H. Blanchard 65 (N), S- ne [North Pownal, Aug. 11, 1902] (Dt, Vt, Vt). Chittenden Co.: _ Pringle s.n. [Charlotte, 15 July 1887] (Vt). gross Co.: L. A. Wheeler s.n. [Towmshend, 5/23/23] (Go, S). NEW YORK: Bronx Co.? E. P. Bicknell 7336 (N), 7348 (N), 7350 (N), 7361 (N), 7352 (Al, It); H. N. Moldenke 18463 (Aa, Ec, dr, Jr, M1, " N)- Nassau Co.: E. P. Bicknell 7353a (N), 7353e (N). Richmond Co.: Monachino 579 (N, Z). Tompkins Co.: Cs. Be , Peck s.n. eeeeieae e] ait Ulster Co.: Elting 207) (Al). ~ NEW W JERSEY: Morr [Lake Hopatcong, 4 Aug. 1902] (B); yackensie h7k6 (N, iF wo SYLVANIA: Allegheny Co.: Holz s.n. [Pitt urg, 1031] (S). Craw- ford Co.: Shafer s.n. [July 23, 1901] ae Delaware ere a Ww. Pennell 11695 (Cm). Luzerne Co.: J. K. Small s.n- {L August 15-16, 1889] (W--298653) . Montgomery Co.: o.: Wismer ee (Up). Philadelphia Co.: Bassett s.n. [Pennypack Park, Oct. ct. 5, 1927 (N). Union Co.: Westerfeld 2377 (Ur). County omiat amined Schweinitz s.n. [1929] (Br). MARYLAND: Washington Co.: He Ne ee 21762 (Z). VIRGINIA: Princess Anne Co.: Fernald & . Long 12h53 (W—1971125). WEST VIRGINIA: Pocahontas Co.: C. Be Clarkson 1585 5 (We). Preston Co.: Bartholomew & Vail s.n. [Big g Sandy Creek, August 15, 1959] (We). OHIO: Champaign Co.: E. C. Leonard 20462 (W—20069)9). Lorain Cos: A. A. Wright s.n. Tsuly 30, 1889] (Ob— 75328). Lucas Co.: Moseley sen. (Aug. 2, 1925] (Ob—97293) « NORTH CAROLINA: Ashe Co.: Collector aoe negianated sen. [Solitude P. 0] (Hi—59)76). ILLINOIS: Adams Co.: R. A. Evers 802, in part (Ur, Ur), 807 (I1—15901). ea ied Coe: Geyer 8 $.n. (Beardstown, July 18h (1—-type) « Champaign Co s 2911 (Ur, Ur, Ur); R. A. Evers 160) (Ur); G. N. Jones ae wits Crawford Co.: Ahles 4900 (Ur, Ur, Ur). Hardin Co»: Ahles 2754 (Ur). Henry Co.: V. H. Chase 6910 (Ur, Ur), 6911 (Ur). Iroquois Co.: Ahles 2538 (Ur); Winterringer 6452 (11-3370) . Jo Daviess Co.: Pepoon & & Moffatt 254 (Ur). Piatt Cow: oS 6580 (Ur), 6581 (Ur). Sangamon ;CO.t Go De ). Fuller Ate Se SN. ‘isonet be tle. (I). Tazewell Co.: : Ve H. Chase 3230 (al, ¢ Ca--B8265h,, I1--15816, N, Ur, W--1897066). Vermilion Co.: Storm s.n. [July 28, 1949] "(ur). Wabash Co.: Schneck s.n. [Sept. 1, 1897] (ur); Schneck & Waite s.n. [It. Carmel, Aug. 5, 1887] (Ur—3001) Winnebago Co.: M. S. Bebb s.n. [Fountain indale] (Al). INDIANA: Boone Co.: S. McCoy 529, (Bt—51727). Carroll Co.: Ek se n. [west of Burlington, 8-7-1941] (Bt--576l1). Cass Cos: Friesner liso (Bt—1065). Clinton Co.: C. C. Deam 1978 (Dm, Mg--202) - 96 Paper 710. OO Tow Vol. 9, no. 2 Franklin Co.: S. McCoy 4170 (Dp—l662). Fulton Co.: Friesn a (St). Hancock Co.: Friesner 18619 (Ca--882638, ey N, Pl—153592), 18620 (N, N, We); S. McCoy 828 (Dp—l671). mili Co.: Grimes 2083 (Dp). Warrick Co.: C. C. Deam 29006 (In). IOWA: Clayton Co.: Is’ ff H. Pammel s.n. [St. Olaf, Aug. 10, 192)] (Io—115923). Hardin Co.: Peck seh. {Iowa Falls] (Io). Johnson Cos: Le H. Pammel 708 (Io--119267, Mg—195). Lee Co.: I. Mitchell 172 (Io--115868) . Muscatine Coe: Pammel, Kelso, & Harlan sn. (Jul. 20, 1919] (Io—950h)). Poweshiek Co. Me M. E. Jones s.n, (Grinnell, August 1875] (Po—-71239). Story Co.: J Jo Re -R. Campbell 67 (Io—8561) . TENNESSEE: Carroll Co.: Eggert sen. [Hollow Rock, L & 5 Aug. 1897] (Cm, N, W—-754370). MICHIGAN: Cass Co.: Pepoon 1347 (Mi). Oakland Co.: Farwell 5159 (Mi). Saint Clair Co.: Farwell Farwel 6741 (Mi). Wayne Co.: Farwell 1426 (Fe), 861 (mi), sen. (Detroit, Sept. 25, 1893] (c); H. H, Rusby sen. (Detroit, Aug. 160)] (Mi). WISCONSIN: Dane Co.: Cs ee Albers 33015 (Au); A. B. Seymour 1062 (Ws). Grant Co.: H. H. Smith 7821 (B). Lafayette Co.: Shinners 2358 (Ws). Milwaukee Co,: Lapham s.n. [Milwaukee] (Ws). Vernon Co.: Fassett 20233 (Ws); H. H. Smith 727) (B), 7366 (B). KANSAS: Bourbon Co.: Clothier & Whitford sin. [Aug. 11, 1897] (Ka). Brown Co.: F, Garner s.n, [Summer, 1925] (Ka--72011). Cloud Co.: Carle- ton s.n. ~ [July 18, | 1888] (Ur). Linn Co.: Brooks & Brooks Bint 11928] (Cm). Neches Co.: A. S. Hitchcock s.n. (Dec. 11, 1896] (Ka): Pottawatomie Co.: Kimball s Son. [St. George, Aug. 12, 1888] (Ka). Republic Co.: F. A, Smith s.n. [July 1896] (Ka). Riley Cos: Je Be S. Norton 792, “in part (Ka, N, Sober fe MISSOURI: Greene Co.: 181535, Ca—l25200, Ca—882858, Cn, om, Du 2021 133, G cg—183218, Go, In, lo—79865, " un—6885, N, roigeson, Tl, Up--60775, 754957), san. [St. Louis, Aug.’8, 1875] (Au, B, Ca-—183100, "tes 425202, cm, D Du——202131, Gg— crbioins Io——-79868, Mn—6896, N, Po 192593, Up—60751, Up, W—754954), sen. [St. Louis, As 1875] (Up-60750), sen. [12 Aug. 1877] (Al, I); Engelmann s.n. (St. Louis, Sept. 1841] (W—71971), bay [St. Louis, July “F5i3} (Dt), 8.n. ist. Louis, Aug. 1843] (Br, Pr, Pr, T), sen. [St. Louis, Aug 1865] (Dt), s son. (St. Louis] (Br, S, Sg—-16097); Glatfelter s.n- (near St. Louis, 8-10-9h] (Mi); s.n. [St. Louis, 8-11-91] (W— 3096); Lindheimer s.n, [St. Louis, July 1839] (Dt); M. Martens Sen. (Br). OKLAHOMA: Kingfisher Co.: Grace 353 (Ok—19032). CULTI- VATED: Spain: Herb, Hort. Reg. Matrit. 26 (Q). LOCALITY OF COLIEC- nerTbs Hort. Reg. Matrit. 26 TION UNDESIGNATED: A. Murray 32 (S); Swederus s.n. (S). 1963 Moldenke, Monograph of Verbena 97 XVERBENA ENGELMANNII f. ALBIFLORA Moldenke, Résumé Suppl. 3: 2, nom, mud, trate), "f. nov. Haec forma a forma’ typica hybridae corollis albis recedit. This form differs from the typical form of the hybrid in hav- e type of the form was collected by Sarat Ee Mathewson Lawrence and W. J, Dress (no. 2) ss marshy gr 20h ar west of Chip Bottom, Ocean County, New Jersey, on astra 2 dole, and is deposited in "the herbarium of the a araias of Galiforns at Berkeley. It is described as a perennial 1 meter tall. ie form is known only from the type spec cimen tations: NEW JERSEY: Ocean Co.: Lawrence & Dress 570 (Ca-— 805326 tore). VERBENA EP EPHEDROIDES Cham., Linnaea 7: 260—261. 1832. Syno: : Verbena virgata Sellow ex Moldenke, Suppl. List In- valid Names 10, in syn. 1941 [not V. virgata Rutz & Pav., 1798]. Verbena valerianoides St.-Hil. ex Moldenke, Alph. List Invali Names Suppl. 1: 27, in syn. 197 [not V. valerianoides H.B.K., 1818). Verbena isabelleana Briq. ex Moldenke, Résumé Suppl. 1: 23, in syn. syn. 1959. Bibliography: Cham., Linnaea 7: eg 1832; Walp., Repert Syst. ots : 17. 18153 Schau. in A , Prodr, 11: 543. 18:7; Schau, Mart., Fl. Bras. 9: 191. ets "Lorentz, Veg. Nordeste Prov. Entre Rfos, ed. 1 86 150, & ie 1878; Griseb., Abh. K. in Engl. & Prantl, Nat. Pflansenten h (Ja 3a): “U7. 189); Jacks, Ind, Kew. 2: 1178. 1895; Briq., im. Conserv. & Jard. Bot. Genev. 10: 101. 1907; Glaz., Bull. Bot. Soc. France Mém. 3: Shy. 1911; Seckt, Rev. Univ. Nac. Cordoba 17: 90. 1930; Herter, Florul. Urug. 105. eee Herter, Revist. Sudam. Bot. : 186. aed ee aie i Shera 2: “ts, a & 5. 1815; ‘Woldenke, Alph. List Cit. 1: 195. 338. a Entre Rfos ed. 2 150, & 168. 19473 Mold Castan- ea 13: 117. 1918; coinaes: * ,lph. rig Cit. 3: 687 & £923 ae) enac., a ain & Phytologia 3: 53. 1951; Stellfeld, Trib. Farmac. 19 ‘Qo 3 Srous and spreading; nodes annulate somewhat contracted; princi- Pal internodes 2.5—6.3 cm. long; leaves mostly rudimentary. NOTES ON NEW AND NOTEWORTHY PLANTS. XXXV Harold N, Moldenke CITHAREXYLUM MATUDAE Moldenke, sp, Arbor 7 m, alta; ramis ant iocas: canis’ perobtuse tetragonis albo-griseis, non marginatis vel alatis, glabris; cicatricibus foliorum valde divaricatis percrassis suberosis; 3 sarmentis in siccitate nigrescentibus; foliis ellipticis integris ad basin a- picemque acuminatis, supra glabris, subtus parcissime pilosulis; spicis agp ay terminalibusve solitariis elongatis multiflo- r drying, elliptic, 6—-1, cm, long, 3--7 cm. wide, acuminate at the base and apex, entire, glabrous above, very sparsely and obscure- ly pilosulous beneath, more densely so along the midrib; midr. slender, flat above, prominent beneath; secondaries slender, as- cending, slightly arcuate at the ti ps, *6—10 pairs, flat above prominulous beneath; veinlet re ticulation indisce rnible above, an bundant and plain but not elevated beneath; inflorescence axil- lary or terminal, spicate, solitary; spikes 725-25 cm. long, ed, er i many-flowered and many-—fruit ect anthesis, apparently drooping in fruit; peduncles short or subobsole glabrous, nigrescent i anthesis; rachis rather slender and nig- rescent in ing anthesis, big eavy and similar in tex- ure and color to the branches 3; flowers ae » shortly pedicellate; pedicels slender, pe Be ges anthesis, much thickened and somewhat longer in fruit; baotinks single under each pedicel, 1—2 mn, long, linear-lanceolate or subulate, glabrous; calyx campanulate, 4——-5 mm, long and wae der gk ely puberulous toward and ae ~ rim, otherwi rwise glabra a > —c eatly intieeask and indura ated, 6—8 « Wide, subpat- elliforn, eabrice » Slightly verruculose on the ou pe surface » vhe rim shallowly erose, not distinctly toothed nor pace drup- aceous, gry ht ee “about 1 em. long and wide, nigrescent and much wrinkled The ne of this species was collected by Eizi Matuda (no. 5467) — in whose honor it is named — ina virgin forest at Pina- beto, near the Guatemalan bor near Motozintla, at 2585 meters altitude, Chiapas, Mexico, on eo 9, 1945, and is deposited in 98 1963 Moldenke, New and noteworthy plants 99 the C. L. Lundell Herbarium at Renner, Texas. LANTANA HISPIDA f. ALBA Moldenke, f. Haec forma a forma typica specied pe albis This form differs from the typical form of the pF in having white corollas. The type of the form was collected by Elias Contreras (no. 1056) on the top of Temple Ri Tikal, Tikal National Park, El Petén, Guatemala, on June 9, 1960, and is deposited in the C. L. Lundell Herbariun at slannes? Texas. LANTANA VELUTINA f. ALBIFRUCTA Moldenke, f. n Haec forma a forma typica speciei fructis albis recedit. This form ee from the typical form of the species in having white frui The type of the form was collected by Cyrus Longworth Lundell and Amelia L, Lundell (no, 12320) on bres Sprain ee near Taxco, at km. 157 on the Acapulco Highway, G , Mexico, on August 1, 1943, and is deposited in the cactus ad: the 8 is ‘ies earch Fo oundation at Renner, Texas LIPPIA PRINGLEI f. INTECTA Moldenke, f. Haec forma a forma typica v4 Te ei hentai foliorum subtus pub- erulis pilis brevissimis recedit. This form differs from the typical form of the species in having the ni leaf~surfaces merely very short-puberulent with rather distant hairs. The type of this form was collected by Efraim Hernandez Xolo~ cotzi (no. 125) at El Llano, south of San Blas, Nayarit, Mexico, on Jamary 7, 19h), and is deposited in the herbarium of the Texas Research Foundation at Renner, Texas. It is sai tree 3--6 meters tall, with white flowers, and called "tacote". STACHYTARPHETA GUATEMALENSIS f. ALBIFLORA Moldenke, f. Haec forma a forma typica speciei corollis albis sbatien 8 This form differs from the typical form of the species in having white corollas, The type of the form was pelehl ie by Elias bem (no. 1579) in an airfield clearing at Dos Lagunas, Petén, Guatema- la, on November 3, eine and is deposited in ey * i FP *Tundell He erbari rium at Renner, STACHYTARPHETA PETENENSIS Moldenke, sp. nov erba; caulibus subteretibus parcissime pilosulis vel glabra~ tis; foliis decussatis tenuiter membranaceis Sppomacnes pe subglabri Sada em argute acutis, ad basin lo alatis, wargiiie sse sbi rkeret spicis acacia elong oR dense multifloris, ee ide valde excavato; peated pena tis, ad apicem longe a ttenuatis; corollis Herb, apparently simple or subsimple, with a epee abbreviated branches in the upper axils; stems subterete, in drying, 100 PHYTOLOGIA Vol. 9, no. 2 very obscurely and minutely scat tered-pilosulous or subglabrate throughout; nodes not annulate; principal internodes 5--6 cm. long; leaves decussate-opposite, the lower ones usually with a few small ones in their 8, thin-membr ranous, brunnescent in pape slightly lighter beneath, lanceolate, 5--13 cm. long, = 36 « Wide, acute at the apex, long-acuminate into the ed etioks at the base, coarsely serrate along the margins eth: on e basal acuminati ion, glabrous on both surfaces except f i few scattered minute hairs on the midrib beneath; iafloress cence terminal, spicate, Resrete ad simple, about 1 cm, long; spikes: etna! many=-flowered, erect, rather stout, the rachis nae ¢ mitely pilomious, oe eply excavated; bractlets lanceolate, ut 9 mm. long, lon ~attemate at the a apex, sparsely ciliolate one the margins, i primer patent; corolla red, the tube 10—13 mm, long, the limb about 1 cm, wi The type of this iota was aii cted by Elias Contreras (no. 1563) in zapotal on the Uxactun trail about 2 km. from the village of Dos Lagunas, El Petén, Guatemala, on October 27, 1960, and is deposited inv the-@4-Ls Lundell. Herbarium at Renner, Texas. VERBENA RINCONENSIS Moldenke. sp. n Herba sarmentosis; caulibus Pratttias procumbentibus; ramis gracilibus adscendentibus parce cdiaaie, pilis albidis; internod- dis elongatis; laminis ovatis irregulariter dissectis vel sub-3- lobatis undique parce pilosis; spicis terminalibus abbreviatis longiuscule pedunculatis; bracteolis anguste lanceolatis saepe divaricatis vel recurvatis pilosulis longiter ciliatis Creeping herb; stems rooting, slender, mostly procunbent; branches ascending, very slender, r. rather pas se e with ra~ ther stiff & divergent white hairs; Prapcaped internodes elongate, long, 1.5—2.5 am. wide, e incised, the larger one lobed, rather sparsely pilose on both faces, the lobes & divis- ions oblong or ovate, mostly acute or subacute at apex; oO ence Ss c thesis, elongate-cylindric in fruit; peduncles slender, 1--3.5 cm- long, densely white-pilose with wide-spreading rather stiff hairs; bractlets narrow-lanceolate , u-—6 mm, long, mostly opiate or .e ven r 2 anes corolla salverfom, tube ca. 8 mm NE, aie ide toward apex, the limb to 7 mm. oie Bg ag George B. Hinton (no. 8011) from hill at Rincén, dist. Te mascaltepec, México, exico , August 2, 1935, Sepnes tee. in herbar- ium of the Texas Research Foundati ion, Renner, QUERCUS XRIPARIA LAUGHLIN KANSAS CITY OAK Hibrida neta Quercus rubra L. X shumardii schneckii Sarg. Erythrobalanus Section Coccineae Series Kendall Laughlin (1890- ) 165 Pine Ave., Chicago, I1l. A tree 40 cm. in diameter and 20 m. tall with Slender horizontal branches and ascending limbs in the crown. Bark dark grayish brown, thin, obscurely ridged, a little warty in the lower portion and quite Smooth above. Season's branchlets glabrous, angled, tawny-brown with light lenticels. Terminal winter. buds 7 mm. long, pyramidal, compressed, acute, red- dish brown, puberulous, with about 6 scales. _ Leaf blades 11.5-20 cm. long, 8-15.5 cm. wide, thin, dull moderate yellow green above and slightly lobes, the apex 3-5-toothed. Base usually obtuse. Petioles 2.8-5.6 cm. long, terete, glabrous, greenish yellow. Acorns solitary or paired, 21-26 mm. long; cup bowl-shaped with an incurved rim, 11-14 mm. high, 18- 102 PHYTOLOGIA Vol. 9, no. 2 enclosed in the cup; peduncle up to 12 mm. long, pu- berulous. QUERCUS XRIPARIA LAUGHLIN Arbor 40 cm. diametro et 20 m. alta cum gracili- bus aequis ramis et ascendentibus membris in summo. Cortex fuscus cinereo-brunneus, tenuis, obscure fasti- gatus, in inferiore paulo verrucosus et supra plane laevis, Novi temporis ramuli glabri, angulati, fulvo- brunnei lucidis lenticellis. Extremae hiemales gemmae 7 mm. longae pyramidatae compressae acutae rufo-brun- minus ad costam mediam; lobi infimi acuti et integri; bus; lobi proximi 2.5-3 cm. lati in bene passis foliis et finientes 4 dentibus; lobi supremi latissimi levi- ter dilatantes ad apicem et finientes 4-5-dentato api- nus 10.5 cm. latus trilobatus divisus in 2 interdum latos 2-4-dentatos lateralis lobos, apice 3-5-dentato. Basis plerumque obtusa. eh Petioli 2.8-5.6 cm. longi, rotundi, glabri, gal- ni. Glandes solae aut compositae, 21-26 mm. longae; cupula poculoformis incurvato margine, 11-14 mm. al- ta, 18-22 mm. lata, tecta hebetibus cinereo-brunneis intus glabrata; nux brunnea, insigniter varia pallido cinereo tomentulo, subquadrata, 17-20 mm. longa lata- que, novem-vicesimis conclusa in cupula; pedunculus tenus 12 mm. longus, puberulens. DISCUSSION Sarg.,which has a diameter of 16 inches, a circumfer- ence of 4 feet 2 inches and a remarkable height of 67 feet, is located in Swope Park, Kansas City, Missouri, U.S.A., at the top of the right, east bank of the Blue 1963 Laughlin, Kansas City Oak 103 River west of Lakeside Drive at about 70th Street. A Bur Oak 13 feet 2 inches in circumference, the big- wi to be cursorily regarded as rubra in the field. ut the obovate shape of the leaves of riparia and winter photograph of a typical schneckii about seven hundred feet northwest of riparia appears on LOGIA. sessed by the same species. The acorn-cups of the t brid has lustrous olive green leaves much longer than 10h PHYTOLOGIA Vol. 9, no. 2 broad open sinuses extending more than halfway to the midrib, falcate lower primary veins, yellow petioles and midribs, a broadly cuneate or obtuse base and in- significant tufts of hairs on the ee ase and deep saucer-shaped acorn-cups with gray scale These characters are very different from rigarie as described above. e holotype will be deposited in the Royal Bo- tanic Gardens, Kew,England, and an isotype will be deposited in the U.S. National Het Data, Washington, D.C., by way of the U.S. Forest Service e Gray Herbarium Card Index and Wm. Tre- lease's Mp American Oaks" show "Quercus rubra X shumardi = Q. joorii Trelease.” This rubra” was the ee hern d Oak, now called falcata. Laughlin, Kansas City Oak 1963 2 E+ fx Ou. pt EH fx] joo Ee OF QUERCUS XRIPARIA BA dey! * a » are 7 106 PHT F010 G1 A Vol. 9, no. 2 THE. TYPE TREE OF QUERCUS XRIPARIA RN > 53% of natural size 1963 Laughlin, Kansas City Oak FOLIAGE OF QUERCUS XRIPARIA 21% of natural size 107 MALUS LANCIFOLIA REHDER Kendall Laughlin 165 Pine Ave., Chicago, Ill. lvalus lancifolia Rehder, the Lanceleaf Crab Ap- ple, was originally described in Volume II of C. S. Sargent's "Trees and Shrubs" (1907-13) and was regard- ed as a distinct species until M. L. Fernald got the idea that it was a variety of coronaria. ernald's description of coronaria var. lancifo- iia in Rhodora 49:232 was sketchy. He did not name a single intergrading character. ald's "description" indicated that he got his information from C. C. Deam. Deam's description o coronaria in the 5th edition of his "Trees of Indiana" is also sketchy. Instead of describing the species in the descriptive text, he writes about Crab Apple thickets, wildlife, poisonous spray and general taxo- nomic problems. H es not mention lancifolia. The photoengraving on e 210, represented to be corona- ria, is actually lancifolia. It is evident that Deam n The type material of lancifolia, sent by B. F. Bush to the Arnold Arboretum, was col- lected in rich woods near Courtney, Bush's home, in 1906. Mention is also made in the original descrip- tion of a collection taken near Independence. 108 1963 Laughlin, Malus lancifolia 109 Coronaria is not found in Jackson County, so Elsewhere in Missouri lancifolia has been found only at Williamsville in Wayne County. Arie F. den Boer has written me that he has never found a single better shaped than coronaria. year. The characters of lancifolia as it grows on the eastern outskirts of Kansas City are so different from the characters of co ia in the Indiana Dunes ot e Shows those characters of the two species that at ty of the specimen. have given the greatest weight, 7, to the col- or of the anthers. The color of the anthers is an 110 P EZ? 00 6 FA Vol. 9, no. 2 and found that their color was deep pink. In May of 1960 and found that their color was orange yellow. These ave given the second greatest weight, 6, to the venation of the leaves. This very distinctive character has been overlooked in the past and has ap- parently been the cause of much of the confusion be- tween the two species. he lobation and the shape of the base of the ves, Key can also be used by persons that prefer T to recognize lancifolia as a variety rather than a Species. 1 Some botanists use the generic name Malus and others use Pyrus for the Apples and Crab Apples. In most other professions there is a central committee that defines terms, outlines proper procedures and settles disputes and I think it is deplorable that the professional botanical taxonomists do not see the need for uniform understanding. lil 1963 Laughlin, Malus lancifolia COMPARATIVE KEY TO MALUS LANCIFOLIA AND MALUS CORONARIA SHOWING THEIR DIFFERENCES WEIGHT| CHARACTER}{MALUS LANCIFOLIA MALUS CORONARIA Leaf Blades: 4 Rounded, often Often broadly rec- semicircular. linear-cuneate or nearly so above the narrowly rounded or subcordate base. z Loba- sypicaiee unlobed|4-7 pairs of rec- tion except for 1 pair|tangular, pointed, of lobes 1-2(-4) |toothed lobes or mm. wide (mea- lobules, the — sured horizontal-| pair usually the y but not always| largest and appear- present) at ing at three- distal end of the]|eighths to one-half rounded base; or |the distance from sometimes scarce-| the base to ly lobulate. apex on lobed ence eaves. 5 Dimen- | Averaging 52% to | Two-thirds to four- sions % as wide as fifths as wide as ‘ile i long. long. 1 Color |Moderate yellow |Dark greenish yel- ects SUOVE AZree low or dark yellow. 6 Primary| Sometimes arcu- Straight, leaving veins te, leaving the |the midrib at regu- midrib at irregu-|lar intervals and lar intervals and|extending to the nearly buried in |points of the the parenchyma, lobes. usually forking and not reaching the margin of the ——__ 1 . — 2 |Flowers | 31-45 mm. wide. |30-32 mm. wide. 7 | Anthers |Deep pink. Orange yellow. 3 | Pubes- Glabrous. Pubescent or hoary- cence tomentose. lower Side of sepals 1 | Pome 21-28 mm. high, |20-23 mm. high, 25-36 mm. wide. | 26-31 mm. wide. 2 PHYTOLOGIA Vol. 9, no. 2 weg ee These leaves of MALUS CORONARIA, which are approximately 17% of natural size, were taken from different trees in the Indiana Dunes State Park. Distortion due to the low angle at which the camera was pointed at them causes these leaves to appear to be narrower than they real- ly are. The leaf on the right was taken from the largest tree in the park, 1'1ll in circumference, and shows the lobulate type of leaf found on old trees. These leaves and flowers of MALUS LANCI- FOLIA, which are approximately three- eighths of natural size, were taken from the A.F.A. champion in Blue Valley Park, Kansas City, Mo. PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and epee. his con- tribution. 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An extra charge will be made for half- tones, depending on their size, as fixed by the engraver, with a minimum of about $2.25. Articles dealing with research in all lines of botany, in any reasonable length, biographical sketches, and critical reviews and summaries of liter ; lori : or so-called pe, and | on the cag of noe ES Pe will be solicited, if necessary, from quali- fied botan Under. the peeneot _—* of ; punting, the basic rate for a page or fraction n of 250 copies. This price is subject vas change without notice, since it ‘depends entirely on the prices prevailing in the printing industry. coe EASEMENT REIN «5 Ne Oe ed ee ; fraction of the edition ne of 250 copes as fried pet aa . Upon request the editors will send detailed instructions “concerning he manuscript or further. information about the magazine. in : ae tay be ae to the passages or to either editor PHY TOLOGIA Designed to expedite botanical publication Vol. 9 | June, 1963 No. 3 CONTENTS MOLDENKE, H. N., Materials toward a monograph of the “genus Verbena. XI Oe ee ek enn eee ene an eee 113 MOLDENKE, AOL., Book tevied:. icc. Gees <5 vo se eset ee _ MOLDENKE, H. N., Notes on new and noteworthy pla g MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XI Harold N. Moldenke EPHEDROIDES Cham. Leaves very characteristic, very minute and scale-like, only 1—2 mm. long, acute at the apex, recurved or reflexed, and glab- Panicle usually very anple, fastigiate, loose and open; spikes Single at the apex of the branchlets or distinctly ternate, pedun- culate, 1--2 cm long, densely flowered, not noticeably elongating from king ‘anes white or else ies from about the middle, no~ ‘sen Seca at the base, very slightly puberulous in the ro e limb sm i i + throa all, violet o 1 e, id, the lobes oblong, jpetvers or eg ad - ste ae ell Truit about — mm» the c e and fuscous~ herbarium of the Botanisches IMuseum at Berlin (and there, feta! ately, photographed by Macbride), but now eR nen nc species is closely related to V. alata Sweet and V. anii Briq. Com- mon names recorded for it are "jaqueca", Gee and "yerba de Santa Ana", Tt inhabits fields, wet places, and swamps, and has been “ay ged tp ape of 350 to 1600 *meters, in anthesis es a n Jan and er. teed nategas of this a has been misidentified and Nokes ae under the names V. litoralis Kunth by Widgren, V. —oralis L. by Dusén, and Neosparton ephedroides Griseb. by | ¢ On the other hand, the Claude-Joseph 814, Hutchison 42, and Jersensen 1403, diutributed as Verbena ephedroides, are actually Diostea scoparia (Gill. & Hook.) Miers. Verbena virgata Rufz & Pav. is a syno nym of Aloysia virgata (Ruiz & Pav.) A. L. Juss. The Britton sheet of Glaziou 17716 bears a supplementary sees : , but this is apparently an error Glaziou, in his formal enumeration of his collections, peep that his nimber 17716 was collected by him in S ° Paulo and does not list a number 17713 for this — at ai, "He states that it blooms in Warch at and Avril, and that the flowers are blue. eon riously, he identifies his no. 17716a as V. perennis Woot., a cap Sk Baa | “dia i 11) PHY TOLOG IA Vol. 9, no» 3 species not known to occur in South America. The type of V. vir- gata Sellow is the same as for V. ephedroides; Sellow's herbarium name (cheironym) was discarded by Chamisso because of the earlier homonym of Rufz & Pavon. The type of V. valerianoides St.-Hil. was collected by Johan Fredrik Widgren (no, 1207) in ‘in Minas Gerais, Brazil, on November , 1845, and - aeponited in the herbarium of the Jardin Botanique de at Etat at el Briquet (189)) lapel v. oe in his eile Verbenaca, 2 sect. Junciformes, along with V. alata Sweet, V. sagittalis Junelli are vont & Hook.) foldenke, J. ae jancea toe Gay) Moldenke, and J. spathulata (Gill. & Hook. ) Moldenke. Wal- sia (185) blacks it in his Sect. Verbenaca, Subsect. Inermes, it cea (Gill. & Hook.) Miers and D. scoparia (Gill. & Hook.) Miers, Junellia glauca and J. spa thulata, and Neosparton ephedroides Griseb. Schauer (18,7, 1851) reas a Bacle s.n. from Montevideo, Uruguay, in either the Berlin or DeCandolle herbarium, not as yet is". all, 17 herbarium specimens and 3 mounted photographs, eS type or phototype material of all the names erly oo been e me Spee BRAZIL: Minas Gerais: Widgren 1207 hapten - sen. [18145] (Br). Paran&: Dusén 11083 (S). Rio Grande do S Camargo 62960 (S); Jttrrgens s.n. [Linha Burity, sevege (Tos 2199) Rambo 3093) (N). Santa Catarina: F. Mttller 90 (Ja-- 46529); Schwacke s.n. [SHo0 Bento, 30/VI/1885] (als, Ja, N)- S#o Paulo: Glaziou 17716 (B, N); Schwacke s.n. [Mogy das Cruzes, 19/IV/1889} (Ja—-l658h). State undetermined: Herb, a ee Rio ref. XI (Ja, N); Sellow s.n. {Brasilia meridionali; Macbride asin 1715) (Kr--photo of type, N--photo of type, N--photo of ype). ARGENTINA: Misiones: Ekman 2023 (Mi, S). XVERBENA FABRICATA a Phytologia 5: 133. 1955. Synonymy : Verbena hispida x urticifolia Dermen, Cytologia 7: 170. — ona Raa ae Bib aphy: Dermen, Cytologia 7: 170. 1936; ee Bia bes logia 7 "67 see a 63133. 1955; Moldenke, Biol. A « 30: 1093. 1956; Moldenke, Am, Midl. Nat. 59: 348--3h9. 1958; gobi Résumé 223, 366, 37, & 471. 1959. This is an artifies ially produced hybrid between V. hispida Ruiz & Pav. and V. urticifolia Ley slappers sad Dermen in a Massa- chusetts garden. The two parent species do no nomally grow hace gether anywhere, so it is not icheiia: that this hybrid will ever be found in the wild. It is not likely to possess any horticultur- al merit either. VERBENA FASCICULATA Benth., Bot. Voy. Sulph. 153. 186. Synonymy : Verbena matthewsii Briq., Ann. Conserv. & Jard. Bote 1963 Moldenke, Monograph of Verbena 115 Genév. 10: 10). 1907. Verbena occidentalis Moldenke, Phytologia Jacks., Ind, - 2: 1178. 1899) ree — Cc Ve . Bot Genév. 10: 10), 1907; Prain, Ind - kh: 245. 1913; Mol- denke, Phytologia 1: ur9—ibo (1910) raoets 1s cents I9lil; Noldenke, uppl. List Invalid Names 9. 191; Moldenk Si List Inva oat: ¥olden e, Known mqgezt. Distrib. Verbenac., * an Eyre herb, iat cate 2 That mi often rambling over the und, the stems and leaves with the aspect of Castilleja spe- cies; stems robust, often erect, apparently woody at the base, sometimes decunbent ramose, obtusely angled, Rae pe braited or paeBir i! Ace ota erect or ascending, 15-~31 cm. i dl j di m. Long leaves decussate-opposite, very short-petiolate or subsessile to plainly sessile, the young ones fasciculate in the axils; peti- oles, when present, very short and winged; leaf-blades chartace-— ous, uniformly gray- or ashy-green on both surfaces, about t cm. deeply incised-crenate or pinnatifid, gee 0.5 mm. wide, the primary incisions to 3.5—-5 cm. long, the lobules about ) m. pe obtuse or subobtuse, somewhat revolute he margins, he rest and much n er, lanceolate or line cute, entire or 3--5-fid b and secon es very slender, impress- ed abov » Prominulous beneath; veinlet reticulation in ble on both surfaces or prominulous beneath; spixes terminal, single, ple, sessile or -pedunculate, unbranched, 7- . Siongate d thesis, very densely wie dy agate the: eae S part 205 and 1.5 cm. r about wide ws canadensis but otherwise similar; peduncles tear, wien, 5 5 ~ — densely hirsute-pubescent like the stems; flowers sweet- Snelling, very densely imbricate (except the lowest 2 or 3, which nay be arranged i ‘whorl somewhat separated from the rest of arated so as t ic — ewhat divergent in poset bractlets large, ear to narrow- anceolate or ih ap ence , 5—~8 mm, long, less than 1 om. Wide, acuminate at the apex, equaling or geese dame Hiren the calyx du during anthesis, eae 7 creukalels with gland-tippe hairs like the peduncles or antrorsely pilose, similar to a of 116 Bete? O1L6G:G:14 Vol. 9, no. 3 V. canadensis; calyx oblong-elongate or tubular, 7--8 mm. long wide during anthesis, similar to that of V. canadensis, its tube about 5 mm. long, 5-costate, 5~angled, hirtous or dense- hirsut ing from lavender or purple to deep- or dark-purple, smaller than that of Vv. canadensis, its Pe about 10 mm. long, surpassing the subemarginate; stamens and pistil barely equaling the mouth of the corolla-tube; anther-glands small; anther-cells distinct, the pulvinus curved-acute, ed; cocci oblong, splitting spontaneously at maturity, rugose above with an — dibs tion of veins; testa very slender, membranous; endos) very sparse, but appar- ently present. The type of this species was collected at Huamantango, ype locality of both was in the same region of Huaroch- (L.) Britton, but is more hirsute and has different leaves. He $4 that it is perhaps suffruticose and that Cuming 954, from the coast of Peru, seems to be the same species, with h rather blunter lobes to the leaves The type of V. occidentalis was collected by Andrew Mathews (no. 498) somewhere in Peru in 1834, and is deposited in the Meis~ ner Herbarium at the New York Botanical Garden, while that of V. matthewsii is given by Briquet as "no. kgs" from the environs the Lima, Peru, deposited in the Delessert Herbarium at Geneva. It i also. given as "95" on the labels of Macbride's photograph * of ee type specimen, but this appears to be an error in reading in both a part collection. Also, the ger ininythy surname should be spelled with 4 nly the Linnean Society and one Journal of Botany obituary aa the name with a double sits and there his first name is given as "Alexander The species has ho found in dry clay or clayey soil, sandy — 1963 Moldenke, Monograph of Verbena 117 loam, stony habitats, hot and dry rocky hillsides, and puna, from 550 to 3900 meters altitu de, flowering from F ebruary to April and in October and December erbari rium eaticiar has been misidenti- fied and distributed as V. laciniata (L.) Briq. Tovar 1 351 is said by the collector to have had whitish flowers and may there- fore represent a color form worthy of special designation. Briquet says "Espéce voisine des V. clavata Ruiz et Pav., V. sulfurea Sweet, V. Berterii Schau., v. lipozygioides Walp. etc. -_ anenaneeeneieines — Toutes ces espéces ont des tiges procumbantes, sortant d'un ase ligneuse et un port complétement dif- férent. Toutes aussi ont des bractées bien plus courtes que le alice, L Me avata Ru aVe, a longue er bractées se ants Ss lobes corollins lineares-oblongs, caractéres qui per- nett, rated de l'en distinguer." 3 herbarium specimens, including the type collections of ota Of the names involved, and 2 mounted photographs have been examined by me. Citations: PERU: Ancash: Ferreyra & Cerrate 12191 (Ok). Aya~- cucho: R. Ferreyra 5170 (N, Ug). Cajamarca: Stork & Horton 10032 (Ca-~-639346). Huancavelica: Tovar 1 1351 (2). Lima: R. Ferreyra 1980 (N); Mathews 498 ["h95"; Macbride photos 24691] (Kr—photo, Ky N--photo); S.G io Be Saunders 797 (S), 792 (S); Stork & Vargas 9312 (Ca~-6),7,36) . i erekes to: Née | 99 9 (Q). Province undetermined: Math Mathews S.n. [in Peruvia interiore, 1862] (M); Née 98 (Q)- XVERBENA FECUNDA Moldenke, Am. Midl. Nat. 59: 349. 1958. Syn * Verbena ena gurmdeon nsis x ambrosifolia f. eglandulosa Ke, Résumé 360, in syn. 1959. Verbena canadensis var. cam wor osifolia f. eglandulosa Dermen, Cytologia 7: 168, in textu. 1936; Moldenke, Résumé 360, in syn. 1959. Verbena ¢ canadensis var. compacta Hort. x V.ambrosifolia f. eglandulosa Dermen ex Moldenke, 19 ~--—_—— Résumé 421, in ious Bibliography: Dermen, Cytol ogia 7: 168. ee Moldenke, ee Midi. Nat. 59: 349. July 1958; Moldenke, Phytologia 6: 330. August ae miei Résumé 359, 360, M21, & W71. pees G weLsaniea, go ~ Under : mttdd and 3¢ 3 : ao fertile hybrids produced arti- ficially re Massachusetts by Dermen — V. canadensis (L.) Perry. These t together in at wo parental pr apparen tly grow least two counties of Texas, so it is possible a the hybrid 113 PHYTOLOGIA Vol. 9, no. 3 will be found in the wild state It should have horticultural merit and deserves further study. xVERBENA FERAX Moldenke, Am. pit aks a 349. 1958. Synonymy: Verbena canadens racemosa Dermen, Cytologia 7: 168. 1936. Verbena canadensis var. eae X racemosa Dermen, Bibliography: Dermen, Cytologia 1 168. 1936; prow ee Am. Midl. Nat. 59: 349. July 1958; abide, Phytologia 6: 330. August 1958; Moldenke, Résumé 360, 373, & k7l. "1959; Moldenke, Résumé Suppl 1: 15. 1959. his name are included the fertile hybrids ag arti- riéial ally in race fcsatee by Dermen between V. canad (L Britton [including V. canadensis var. atroviolacea a and V. canadensis var. compacta Hort.] and V. racemosa ert. These two eae apparently grow together in at least two counties of Texas, so re is a distinct possibility that the hybrid may be found in a ite state. It may have horticultural merit and deserves further study. VERBENA FERREYRAE Moldenke, mehegt ig 3: 279. 1950. Bibliography: Moldenke, * Phytol a 3: 279 & 286. "1950; E. de Salat as Ind. Kew. Suppl. 11: 263. o383; Moldenke, Résumé 85 & 71. 1959. Herb; stens and branches erect or ascending, obtusely tetrag- onal, brunnescent in dryi ing, rather sparsely puberulent with The "type of this species was collected by my good friend and 1963 Moldenke, Monograph of Verbena 119 colleague, Dr. Ramén Ferreyra (no. 591) -- in whose honor it is named type collection, and = io rbarium specimens, ime taine the type, have beon examined Citations: PERU: iyathbhot R. Ferreyra 5491 (N--type, Ug-- isotype) . “Secure VERBENA FILICAULIS Schau. in A. DC., Prodr. 11: at 18,7. sh Syni irri fe Verbena pinnatisecta Schau. in A. DC. Prodr. 11: > a3 CAT ee TLR Ie Bibliography: Schau. . DC., Prodr. 11: 549 & 556. oe Schau. in Mart., Fl. a é 192. 1851; Briq. in Engl. & Prantl, Nat. Pflanzenfam. ) (3a): 18. ath Jacks., Ind. Kew. 2: 1178. 1395; Hayek, SP Sa Kais Akad. Wiss. Maths -nat. 79 (1): ust 0, Fl. Rio randie do oy at? < 19,6; ae Alph. Li at. is "56, 166, : 177. 1946; Moldenke, Gastanea 13: 117. 19183 Moldenke, Alph. List Cit. 2: 363, 36h, 366, 367, 375, 53k, 550, 551, & 598 (19k3), 3: 670, 675, 086, 688, 751, 81h, 8b6, 863, 920, & 923 (1949), and 4: 1081, 1097, set 112h, 1180, 1222, 1249, & 1250. 1949; Moldenke, known Geogr. ’ Distrib. Verbenac., » 9a, 106, & 198. 1995 vt oe Phytologia 3: 288 13805 Stell feld, Trib. Farmac. 19 (10): 166 & 267. 1951; Sellowia 7: 260. 1956; angely, Fl. Paran. 7: 12. 19573 Ursa Résumé 110, 127 3725 & ng 1959; Angely, Fl. Paran. 16: 78 1961 be = |e o. 3 @ Qa Cae} oO o 4 Pp. ad 2 late, acute or rather obtuse at the apex, entire or bifid, sub- inflorescenc e ateral, the spikes pedunculate ©: long-pedunculate. the terminal ones single and simple or ternate the Jater nes again ternate or ternate-corymbose, cylindric~ oblong, Oy ‘Lowe: or imbricate, not very ™m ny-flowered, 120 PET O.4:0:G- 25a Vol. 9, no. 3 the a oor acute at the apex, obsoletely ciliate; “ee short, tubular, about 3 mm. long, subcostate, the rim short—den ; varying from blue or red—bl fc) violet, lilac, or red, glabrous on the outside, its tube somewhat surpassing the calyx, the throat closed by dense barba irs, the limb velutinous, lanuginous in the center; stamens inserted rt 7x, about ng; cocci terete, fuscous, obsoletely striate on roi dor- sal surface and areolate at the apex, the commissural fac whitish-leathe Vv. pinnatisecta is based ona aoe made by Lucwis Riedel in wet swampy places 4 or Mugy das Cruces, Sao Paulo, Bra- i Friedrich enaliae (no. 20) cheney in Vinas 1, while Vv. filicaulis was based on a collection made by Peter Wilhelm [und (no. 823) in S%o Paulo, one by Sellow (no. 28) in Minas Gerais, and one by Johan Lhotsky ["Khotsky" according to Schauer] in Minas Gerais. ae distinguishes the Gerais, Zi two by saying for V. filiceulis: "Proxima \V. pinnatisectae, pot- issimum teneritate p. ium, scilicet erp debilibus foliisque minus dissecta laciniis angustioribus, diversa." Personally, think that they are conspecific. Schauer also suggests that his V. pinnatisecta might be the same as V. crithmifolia Gill. & Hook. {"quae vero ex descr. differt foiiorun - ater angustioribus earibus, bracteis linearibu us, calycis dentibus obtusiusculis et corolla. calyce aupib longiore"] and that his V. filicaulis might be the same as V. crithmifolia ¢ minor Gill. & Hook. [=V. hookeriana (Covas & Schnack) Mo Moldenke], whic. h he apparently knew only from the descriptions. These s species, however, as all very distinct from that of Schauer. Verbena filicaulis has been collected in swarps and swampy places, wet crassy campos , bogs, and meadows, on often inundated riverbanks, in fields and bare soil along railroad tracks, and in pinewoods with Butia, at pada of 90 to 11,00 meters, flower- ing in May and iacaat to January. Briquet (189) places _ in his Sect. Verbenaca, eda Leptostachyae. Hoehne says of it "planta meio rasteira" and Pickel says "em solo humido da art What Schauer means by his "caules....lateralibus foliis alternis sub- excavatis, ey subjectis planiusculis submarginatis et subliter Striatis" is not certain. Herbarium material has been misidenti- fied and distributed as V. tenuisecta Briq. On the other hand, the Dusén 7108, distributed “as V. pinnatisecta, is the type col- lection of V. dusenii Moldenke,. Hayek (1903) cites Wettstein & Schiffner s en. [Eutantan, V. 1901] and S.n. [Tieté » VIII .1901) fron So | Paulo, while stelifel 1963 Moldenke, Monograph of Verbena 121 oo) cites two additional Tessman and one Stellfeld 5 on tan aran4 -- Herb. Mus. Paran,. 2232, 2447, and 2527. ride pe diact 17439, ci cited eget is of a specimen ioe in the oer arium, now destroyed. In all, 67 herbarium specimens and 9 photographs, including = or Piotr material of both names involved, have been ex- me. Citations: BRAZIL: Minas Gerais: Henschen s.n. (Caldas, 23/10/ 1868] ee Mosén 969 (N, S); Regnell 11.211 [17/11/1866] (N), 11.211 (Caldas, Oct. 23, 1868] (N, S); Sellow 20 [Macbride photos 17439] eae Kr--photo, N--photo, N--photo, N——photo), , N—photo of cotype, Ut--cotype, Z--photo of co- type); Widirren S70 [185] (W-—-1323092), 1210 (Br), sen. [185] (Br, Ja--6590, N, S), sen. [Brasilia] (S, 8). Paran&: Dusén 7106 (S, S), 15679 (Ca--501671, N, S, S, W--148176h); Hatschbach bach 2561 (N), 2672 (N); Jonsson 1392a (N, 5, W+1L01765); Smith, Reitz, & cassie 5 9568 (Ok, W--2251567); Tessman s.n. (Herb. Mus. Paran. 3395] (N). Rio Grande do Sul: Rambo ambo 35413 (N), 3643 (Ss), 54571 (By, 57237 (S). Santa Catarina: F. 2 F. Miller 9h (Ja--6532) 5 Reitz & Klein 7711 (Nm), 7738 [Herb. Barb. Rodr. th & Ke: Klein 11041 (N, Ok, W--22517h6), 11404 (Ok, W--2251778) ; Smith & Reitz 9122 (Ok, W——2251h93); smith, Reitz, & Sufridini 9544 (W- 2251563, Z); Ule 1536 eeoien aes ten 1301h) (Ug). S&0 Paulo: Brade 2738 (38 2288L, 5, Sp--6726); C. Duarte 196 (N, Sp--15723); Ed- wall s STicun Branca, Ave. ie “1895; Herb. Com. Geogr. & Geol. 3127) (Sp aera), ¥. C. Hoehne sen. [Butantan, Sept. 15, 1917] (XN, N, Sp—~5),3) ; We . Hoehne e 527. (N); Lofgren s.n- {Pinheiros, Nov. 2, 1893; Herb. Com. Geogr. & Geol. 1560] (N GF cae Lund 823 (Macbride photos 7854] (Kr--photo of cotyp of cotype) > Pickel 1759 (N, Sf); Usteri s.n. [Tieté, Lite Pn 5 STI. 1 PoE oastese: De Rodriguez 570 (Herb. Inst. Miguel Lillo VERBENA FLAVA Gill. & Hook. in Hook., Bot. Misc. 1: 170. 1630. onymy: Verbena flava Gill. ex Reiche, Fl. © hile 5: 63. Pig slangulria fl flava (Gill. & Hook.) Schnack & Covas, Darwin- ana 6: 75. " Bibliggs att Hook., Bot. Mise. 1: 170. 18% Walp., Repert Syst. iy he 30 . 1845; Schau. in A adr. ll: 555. 18475 Jacks., Ind. Si 2: 1178. ei Greg — wags on, La Plata 3: wuman a dp tite . Hist. Nat. Buenos Aires 2): 13. 1913; Heken, “Pages is 2: oi a eas - 2. OT; 12195 Stapf, Ind. Lond. 6: 429. 1931; Noldenke, Known ang Dive aps Winiana 9s; 's Seteaek & Covas, Revist. Argent. Agron. 12: 222. 15, Pa a 122 PEP T OLTCrTs Vol. 9, no. 3 Taxon. 1 (14-~16): 6. 1945; Cabrera, Bol. Arg. - Bot. 1: 67. 1945; Moldenke, Alph. List Cit. 1: 93 & 202, ishé; “Moldenke, Phy- tologia 2: 338. 1917; Moldenke, Alph. List Invalid Names Suppl. 1 10 & 2h. 1947; Moldenke, Alph. List Cit. 3: 881. 199; Moldenke, Known Geogr. Distr pd Verbensc._ [ed. 2], 106 & 198. 199; E. J. alisb., Ind. Kew ppl. 11: 101. 1953; Moldenke, ages 127; 296, 36h, & Li7l. isco, eoidenn, genie Suppl. 3: Illustrations: Sangin Anal. G02. Cient. Argent. 583 ae 19195 Schnack & Covas, Darwin jana 7: - 1945. Stems pro cumbent at the “Heh eaee linear-lanceolate, entire, pinnatifid, glabrous, the segments few, linear, trifid, Hm a flowers loosely spicate; calyx elongate, pubescent, the teeth very unequal; bractlets minute; corolla ais llow or Repay: to rose, glutinous; chromosome number: 2n This Andean species was based on rei collections made by John Gillies in the aga valleys near Mendoza and on the hills at Rio del Diamente, Mendoza, Argentina. Walpers (185) places it in his aati ver benaca, * subsection Inermes, Group Foliosae, Subgroup Macranthae, and Secondary Subgroup Hee tis with ten other species. It tas been collecte ted in sandy places at altitudes re) m. and in Sep- Ne ium material has been misidentified and distributed as V. glutin- osa Kuntze. On the other hand, the H. H. Bartlett 1910, .— distrib- uted as V. flava, is actu ally ve. ae Moldenke. Ruiz Leal 13088 bears a notation to the effect that the flowers vary from "yellow to rose", perhaps due to age. 8 all, 22 herbarium spec- Citations: ARGENTINA: Chubut: Graetel s.n. (Br). Mendoza: Carette 339, in part (N), s.n. (Ruiz Leal 2558] (N); Ruiz Leal 6913 (N), 7093 (N), 7117 (N), 73h9 (N), 8503 (N), 8960 (N),9292 (N, N), 9790 90 (N), 13088 (: (Z); Sanzin 310, [Herb. Osten 19637) (Ug). Rfo Negro: W. Fischer 59 (Gm, (Cm, N, Ur, W—-70229, W--70,2 30) « Province undetermined: Kuntze _ son. Fmapas Reise 35° 1.92] (N, W- 701714, W—701715) . A FLAVA var. ANGUSTILOBA Speg., Anal. Soc. a Argent. "534 249 [Nov. Add. Fl. Patag. 2: repr. 72]. Synonymy: Verbena flava angustiloba Speg. ex sound Résumé 364, in syn. 1959. Bibliography: Speg., Anal. =<: Cient. Argent. 53: 249 [Nov- Add. Fl. Patag. 2: repr. 72]. 1902; Hauman-Merck, Anal. Mus. 413. 1913; Sanzin, Anal. Soc. ient. Argent. 88: 127. 1919; Moldenke, Known Geogr. Distrib Verbenac., [ed. 1], hl & 101 (2942) and [ed. 2], 106 & 198. 1995 Moldenke, *pésumé 127, 364, & 471. This variety d iffer ers from the t; mie ie form of the -acoaneaypt in having its branches elongate and slender, the lower leaves tri- 1963 Moldenke, Monograph of Verbena 123 partite with linear segments, the upper ones entire, with elon- gate narrowly linear segments, the corolla yellow, its throat nes, e ina, du 1898. Hauman-Merck (1913) cites a Garbers specimen from Choele- Choel, Rfo Negro [Hauman-Merck 343]. The variety is known to me only from literature. FLAVA var. LETT Speg., Anal. Soc. Cient. Argent. 53: 249 [Nov. Add. Fl. Patag. 2: repr. 72]. 1902. cg ae a flava latiloba Speg. ex Moldenke, Résumé 959. peaaeccanty Speg., Anal. Soc. Cient. Argent. 53: 249 [Nov. dd. Fl. Patag. 2: repr. 72]. 1902; Hicken, Physis 1: [Canist. Neuquen.] 127. 1912; Sanzin, Anal. Soc. Ci ent. nets 88: 127. 1919; Moldenke, Known Geogr. Distrib. Yerbenac., Lh & 101 (1942) and [ed. 2], 106 & 198. 199; Moldenke, geet 127, 36k, &h 1h. 1962. ime I Q magi var, LATILOBA f. ABORTIVA Hicken, Sert. And. 65. Bibliography: Hicken, Sert. And. 65. 19233 Moldenke, Geogr. Distrib. Verbenac., [ed. 1], 4 & 101 (192) and ey a. 2], its anther-glands very mi minute, not at all conspicuous. The leaves are almost iy ar blr P rarely with a few entire of the Pcs was collected by Cristébal Marfa Hicken (no, 138) at Arroyo Morro, on the Rio Diamente, at an altitude of meters, Mendoza, argentina, in December, 1912. The lerealgiy! andul4s abortan er St incontrar una u ot het pecuefia serie on las partes Superiores de los tallos." "The form is know to me only from literature, 12h PRYTTOLOGTA Vol. 9, now 3 VERBENA GALAPAGOSENSIS Moldenke, Phytologia 2: 55. 1 Bibliography: Moldenke, Phytologi a 2: 55. 1941; M Known Geogr. Distrib. Verbenac., {ed. 1), 34 & 101. ee woselel ke, Castanea 13: 116. 1948; Moldenke, Known Geogr. Distrib. Ver- benac., [ed. 2], 71 & 198. 199; Moldenke, Alph, List Cit. 3: 972. 1993 BE. J. salisb.., Ind. Kew. Suppl. 11: "263. 1953; Moldenke, —— 81 & 471. 1959. , nigrescent throughout in drying; stems and branche Prreey acutely tetragonal, often sulcate and striate between the angles, very tga! pilose with short whitish widely scat- tered antrorse nodes annulate; principal internodes 1--5.5 em. long; ae: “decissate-opposite, oe sub-clasping at the base, linear o ry narrowly lan ol e, - long, rather minal, spicate, --15 cm. long, loosely many-flowered; peduncles slender, similar to the branches in “all respects, nigrescent in ing, aso Ae tetragonal, ere ge eter striate, very cs nia scattered-pilose, 1--3 cm. long; bractlets ovate-Lanceolate bout 2.5 cm. ia, ciliolate-margined, acuminate at the ape ex, “ast ally brate or very obscurely ‘pilostous except for vate margins; mature flowers and fruit not The type of this remarkable species was collected by Alban Stewart (no. 3313) at Cowley Bay, Albemarle Island, in the Galap- agos Is lands, on August 10, 1905, am is deposited in the Britton Herbariun at the New York Botanical Garden. The collector states in the past with V. litoralis H.B.K. In all, 6 herbarium speci- 7 Re ra the type, and mounted photographs have been ex- e me. Citations: GALAPAGOS ISLANDS: Albemarle: A. Stewart aan (Bi-- isotype, es ces sf Deer han. Gg--3137h--isotype, N--t --type, photo of isot; --photo of isotype, Z--photo of mae S 3u19 (Bi, 62-1373, W—-921598). ENA GENTRYI Moldenke, Phytologia 2: 27. 1941. Bibliography: Moldenke , Phytologia no! Ce T9is1; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 1 101. 1942; Moldenke, Alph. List Cit. 1: 233. 19h6; H. Ne % Ae L. Mold denke, Pl. Life 2: 60. 1948; Moldenke, Known Geogr. Distrib. beta "fed. 2], 33 . 198, 19195 i Moldenke, Alph. List Cit. : 1246. 1949; E. J. Sal- ew. Suppl . 11: 263. 1953; Moldenke, Résumé 39 & & h7l. Low spreading bush, branched from the base; branches rather slender, tetragonal, often purplish, very sparsely scattered- pilose with rather long weak hairs or glabrous; nodes annulate; principal internodes 1--3.5 cm. long; leaves decussate-opposite; petioles indistinct, to 5 mm. long, winged, ampliate and clasping 1963 Moldenke, Monograph of Verbena 125 the stem at their one sparsely acatveree ee or glabrescent, h ene elli serrate from apex to below the middle, rather abundantly ap- Pee iea apie! gees on both surfaces, more densely so beneath, a cabrous above, not glandular; venation slightly ectdeoesened i ce he » PB pound, akas very slender, to 19 or more cm. long, many-flowered, the flowers closely imbricate before and dur. aie anthesis, rather uni- formly separated in fruit; peduncles 1--3 cm. long, slender, gla~ brate; rachis slender, glabrate; 3; bractlets pe e, very snall, about 1--2 mm. long, sharply acuminate at the apex, sub- glabrate or = minutely ciliolate at the base; calyx narrow- ubular, about 1.5 mm. long , glabrous or gubglabrate; corolla very tiny tii eonee © mm, lon The type of t alo was pe ed 8 by Howard Scott Gentry (no. 920) a ra cra honor it is named — in a moist canyon bottom in the Short-tree Forest zone, a an uasaiee of 1500 feet, in the Sierra Monterey, Sinaloa, Mexico, on March 13, 19h0, lia L., to which it is related. Thus far it is known only from the type collection. ee all, 6 herbarium specimens, including the type, have bee canined by me. Citations: weerco mpereniepee H. S. Gentry 5923 Soars see isotype, tin oBkOdT_Anotype. Fs—isotype, type, Me—isotype, isotype, N--type). VERBENA GLABRATA H.B.K., Nov. Gen. & Sp. Pl. 2: 276—277. 1818 (not V. glabrata R. A. Phil., 1870]. Synonymy: Verbena glabrata Humb . ex Spreng. in L., Syst. Veg., ed. 16, 2: 740. 1025. Verbena glabrata Kunth ex ee in A. DC., Prodr. 11: Sh2, in sp: “mn. 1847. verbena glome rineq & Gerard in Réveil, Régne Vég. h: pl. 18. 1870. jeans S hambeta Auct. ex Stapf, Ind. Lond. 6: 429. ae Bibliography: H.B.K., Nov. Gen. & Sp. P . 2: 276--277~ 1818; Spreng. in L., Syst. Veg., ed. 16, 2: 7h8. 18255 Walp., Repert. Syst. Bot. : 20. 1845; Schau. in A. DC., Prodr. 11: 542. 18475 Hérincq & Gerard in Réveil, Régne Vég. 4: pl. 18. 1870; Hook. f. & Jacks., Ind. Kew. 2: 1178. 1895; Stapf, Ind. Lond. 6: 429. 1931; Moldenke, gr. Distrib. Verbenac. 5 Tey ees Moldenke, Holmbergia ): 151. s WOK, ener ae iced + 162% Moldenke, Alph. Lis pe i 268 (1946), 2: ares 641 (1948), 5: 687, ‘23, hile 805, hs, ist, 871, 872, S01, & 972 (1949), and h: 107% & 1078. 1995 "yoldenke, Known Geogr. Distrib. rbetas ed. 2], 0, Lie a ; 1949; R. Espinosa, Est r 3: 68. 1949; Moldenke, os hytologia 3: 236’ & & 37 E. eo Moldenke, Sp. Subsp. Cont. Mold. sai bs. (3. 1951; Moldenke, Inform. Mold. Set 6 Spec. h. 1951; 126 Part OL OG 1 A Vol. 9, no. 3 Moldenke, Phytologia ): 267. 1953; Moldenke, Inform. Mold. Set Py Spec. [h] (1954) and h9 Spec. 3. 1954; Moldenke, Mem. N. Y. Bot Gard. 9: 177. 1955; Moldenke, Inform. "Mold. Set Si Spec. h. 1956; Moldenke, Résumé 69, 80, 81, 85, 364, & 71. mand | Moldenke, Ré- sum Suppl. 3: 13 & 38 (1962), ft: 16° (1962), 6: 5 & 6. 1963. Tllustrati aire ee & Gerard in Réveil, ae tee ties h: pl. 18 = color). Oo. ent or ei ae perennial herb, 0.5—-1 m. tall, much ser practically glabrous, usually of strapeling aroma habit, with h many stems from a thick root; stems mostly procumbent, branched; branches Letgigconl, usually with ascending or erect tips, to 60 cm. or even 1 m, tall; —— decussate-opposite, mostly brunnescent or nigrescent i n dryi lanceolate-oblong, 5- 6.5 cm. long, 1.8--2 cm. wide, cuneate at the base, sharply in- cised-serrate, reticulate-venos » glabrous on both surfaces or with a few, fine, white, appresse ed hairs beneath and a few coarse hairs above; spikes term ninal, et cylindric-filiform, the lateral ones long-pedunculate and about cm. long, the central stripes, hispidulous; corolla varying from blue pale bluish-violet to purplish, purple, pale-violet, pale-lilat, lilac, pale-rose, or very pale-mauve, sometimes white or — white, the tube often red, slightly surpassing the calyx,. limb often bue-violet, mcportens fruit red-brown, pk et eet included by the igang Mountains of Colombia. Teles (1845) places the species in his Section Verbenaca, Subsection Inermes, Group Foliosae, Biber P Micranthae, and uscridaie eee Gk ey along ag: 2 other species. Hooker & Jackson 895), however, reduce synonymy under V. litoralis H. ae while cahaney (1847) nese it to Vv. littoralis & Pycnostachya Schau., with the comment "Hu- jus modif, monticola, magis compacta et glabrior est V. glabrata Kth." and cites Herb. Willdenow 11133. Truly, the spec ecies is closely related to V. V. litoralis, but I have personally collected both in the field and find that the habit and aspect are quite distinct He rbarium material has been ss dg are and distributed un- der the names V. gracilescens (Cham.) Herter, V. litoralis H.B Ks V. littoralis H.B.K., V. littoralis L., and V. _ polystachya H.BeK. Collectors have found V. glabrata growing as a weed in gard- ens and upon walls, in the crevices of mud-walls, along roadsides, in humid forests, in hedgerows and roadside thickets, the dy soil s side of quebra on paramos and the floor of craters, and in raod-cuts, at ea ape ee 200 to 3835 meters, flowering from November to May and July to September, and fruiting in February, 1963 Moldenke, Monograph of Verbena 127 April, May, and July to September. spain reports it to be medic- inal; Killip & vehi say that it is used in the Supa of mails te Color cular name i ‘ben hould ¥ Soted that the V. plsbrata Pe “Pekin belongs in the ee f V. chilensis Moldenke, whic e. The illustration cited above, unfort a o single Fein and therefore is of little re in identifying ac- curately the plant called V. glomerata. Ball makes the curious statement that the flowers of V. glabrata open "at the tips of t rachis which lengthens and the flowers always open from the bot- oe upwards. In all, 97 herbarium specimens have been examined y m Citations: COLOMBIA: Cauca: E. L. Core 207 (W-~2059688); H. Pittier 128 (W--53162l). Narifio: R. Espinosa 2745 (N); Schultes & Mardoqueo 7422 (W--199614h), 7ub2 (W--1996163) 5 > Sneidern A.31l (S, S). Norte de Santander: Killip & p & Smith 21197 (W--1356107). Putu- mayo: Schultes & Cabrera 18901 (ss, W—2172312). ECUADOR: Azuay: Rose, Pachano, & Rose 22851 (N, W--1022h48), 22939 (N, W--102251h). Bolfvar: Pertand d & Summers 602 (N). Caflar: Rose & se & Rose 22730 (N, W--1022359), 22740 (W--1022367) « Chimborazo: André K. K.721 (N); Wille 39 (N, W--1121)23); Rimbach 671 (N). Cotopaxi: +: Weydahl 7h (S). Imbabura: Asplund 20189 89 (S). Pichincha: Anthony & Tate 333 (W--119210); Asplund 6110 O (S, W-—-2223752), 16130 (Ss), 16139 (8), 17227 (S), 20151 (S); Balls 5801 (N, W—17792%); We He . 17e- (N); Couthouy SiN. . fQuitensian Andes] (T); R. Espinosa osa 2435 (OF Firmin 77 (N, W--13)0L69); F. R. Fosberg 22543 (N, W— 2109606); F.C. Lae 13la 2 (oa)s Ee a Idenke & Woldenke 19783 (B, Bis cb, ay » Mg, “ig N, N, N,N, , No, Ok, “Ok, Ob, Rs, S, ‘ou Ss, ak. “78k oe Bs, y vty, , Hw, Le, Ln, Mg, Mr, N,N,N, Wo, Ok. Ot, Rs, ~ S, Sm, Ss, SS, 2); G. W. Prescott 133 (X). Tunguragua: Asplund 19923 (s); pees: 3 65 (N); Ge H. He Tate 562 (W--119755h). Province undetermined: Iille 16203 (Kr). GALAPAGOS ISLANDS: Albemarle: A. Stewart 3317 (Bi, Ge—31375, N, W—921597). Chatham: Schimpff 132 (N)~ PERU: Arequipa: Alva A. 0189 (W--2270251). Ayacucho: R. D. Metcalf 3029h (W—18 3,981) « Cuzco: Herrera 3362 (N). Lima: 2 Re '. Ferreyra 4087a (N)- San Martin: Ll. Williams 5911 11 (W-~14966L3). VERBENA GLANDULIFERA Moldenke, Phytologia 3: hh. Synonymy: Glandularia elandulifera (Moldenke) pteabon te Revist. Invest, Agric. 5: 83 & 206. 1951. Ss geo Rago af Pape Invest. Agric. 6: 82, sphalm. 1 liography: Moldenke, Phyto legia a via 318) and 3: 76 & Lx, alg; Moldenke, Known Geogr. Di Verbenac., [ed. 2], - 1949; Moldenke, ee ceha. oe "505. 1950; Ragonese, 128 PHY T0166 Is Vol. 9, no. 3 Revist. Invest. Agric. 5: 0, 85, 207, & ee (1951) and 6: 82, 83, & 206. 1951; E. J. Salisb., Ind. Kew. Suppl. 11: 263. 1953; Mol- 195 hairs glanduliferous; a annulate; principal internodes 1-~1.5 1 or vi ilac or violet, its tube ——9 mm - long, very minutely puberulous rate, its limb about 5 m:. wide. The type of this species was Tei aited by Nélida S. Troncoso (no. 292) at Ascochinga, at an altitude of 700 meters, Cérdoba, Argentina, on October 26, 1936, and is cers Gag in ‘he Britton Herbarium at the New York Botanical G hill near a limestone quarry, at altitudes of 700 to 2100 “aur flowering from October to April, fruiting in March and April. Purkart describes it as "common" and Troncoso as "ve ery abundant” in Cérdoba. It has been misidentified and distributed in herbaria under the names V. erinoides Lam., V. marrubioides Cham., Ve mic~ rophylia oe. and "cf. V. eed Cham." _SAGORARS der: ia de las aes de oérdoba y Catamarca, que tinicamente hemos observado en una sola oportunidad en el terraplén de las vfas del ocarril." He pe ees tes ce it from the related ve dissecta Willd. as follow. 1. Calyx with ere hairs; corolla 10--14 WUs FORE Reta Fee la. rea SanriE, but without glandular hairs; gore 7--10 ee re ent ren f - dissecta. In oe herbarium specim Sasiruacee no types of all | the names tisha. and mounted ni rir have been examined by ‘Citations ARGENTINA: Catamarca: Brizuela 69 (N); Cabrera 1963 Moldenke, Monograph of Verbena 129 1099 (Ja~-30392, N); Castellanos s.n. [Herb. lus. Argent. Cienc. Nat. 33877] (N); Jtrgensen 1026 (W-—704752); Ragonese & Piccinini 6032 [Herb. Inst. Bot. Minist. Agr. Nac. 69123] (N). Cérdoba: H. H. Bartlett 20079 (Mi), 2017) (Mi); Burkart 794 [Herb. Osten 23367] (Ug), “10413 (N); Castellanos s.n. sen. [Herb. Osten 15289] (Ug), s.n. [Herb. Osten 15291] (Ug); T. H. Goodspeed 1726 (Ca--656782) Hieronymus sen. [.XI.1876] (W--281545); A. T. Hunziker 7175 (N); Lossen 72 (Ba, Ba, Mg—31); 0'Donell 4h27 (N), bh67 (N); O'Donell & Rodriguez 340 (Ut—115h16b), 493 (Ut--115),07b), 610 (Ut—-115417b); Osten 10613 (Ug), 17091 (Ug); Pierotti 5108 (N); Fe F. A. Roig 10h (Herd. Ruiz Leal —— (Z); Sota 1309 Q); ——— oa 2(F—- photo of type, N-- , N--photo of type, Si--photo type, Z— photo of type). _ reien Wie & Grassi 2028 (N); gee ok A 739 (N). Santiago del Estero: Pierotti "h" [Tazana, oii V/ub; Herb, Inst. Miguel Lillo 9960] (Bm a) pa sen. (Villa Gussay4n, April 9, 19h; Herb. Inst. Miguel Lillo 99587] (S). VERBENA GLUTINOSA Kuntze, Rev. Gen. Pl. 3 (2): 256. 1898. Synonymy: Glandularia glutinosa Schnack & Covas, Darwiniana 7: 72, 7h, & 75, pl. 2B, nom. nud. 195. ae fischeri Mol- denke, Alph. List Cit. 2: 625, hyponym Biblio graphy: Kuntze, Rev. Gen. Pi; = ays 256. 1898; Briq., - Conserv. & Jard. Bot. Genav. 4: 21. 1900; Thiselt .-Dyer Ind. Kew. Suppl. 2: 191. i90h; Sanzin, Anal. Soc. apr Argent. 88: 97, 98, 12-126, & 13k. 1919; Stapf, Ind. Lond 1931; Moldenke, Lilloa 6: 328 (19l1) and *8: 430. 1942; ‘yoldenke, know Geogr. Distrib. Verbenac., [ed. 1], 4h & 101. 192; M na” Lilloa 10: 38. 19h; Schnack & Covas, Darwiniana 7: ne Tu, & 75, pl. 2E. 1945; Schnack & Covas, Revist. Argent Me 12: 223, 1945; Schnack & Co Me pea ine: 7: 75. 19453 Covas & Schnack, Darwiniana 7: 86. 955 loldenke, Alph. List Cit. 1: 96 (1946), 2: L3 & 629 (a8), 31 687 (2949), and li 1255. 1949; Mold ke Known Geogr. Di eS Verbena 106, & 198. 19119; Moldenke, Phytologia 3: 3h & hia (19 919) oe 3: 290. 1950; Moldenke, Résumé 115, "127, & 71. 1959; Moldenke, WW Pp Tllustrations: Sanzin, sage Soc. A Argent. 88: 125. 1919; Schnack & Covas, D ana 7: p rennial very glabrous or Sg N nerd, to xs cm. often retaining bl ains of sand; s many, t to 50 pests ched, glabrous or sc pilose, glutinous or subglutinous throughout; branches often decumbent, . te - long and basal or sub-basal, the margins saga’ involute; cence spicate, Pasion spikes at first densely, finally loosely few-flowered, terminal; p peduncles 1—3 cm - long; bract- 130 Rat oe O LO G.T sk Vol. 9, NOeg lets small, herbaceous, about 2 mm. long, ovate, acute at the pex, ciliate along the margins; flowers about &’p er ie 3 Light ly fracrant; calyx long-tubular, about mm. long and 1 mm. wide, its rim with 2 or:3 linear-subulate teeth, shorter Gin the tube; The e of this species was collected by Carl Ernst Otto Kuntze ce (?) "2"] in the Cordillera at Paso Cruz, 34° S. lat., Mendoza, “Argentina, at an altitude of 1500 meters, in January, 1892, and is deposited in = ogc Herbarium at the New York Botanical Garden. This spec s also the type of V. Ss Leal describes the species as being very comm through the Mendoza region in Cretaceous sand and Sasal it detri- tus and abundant in sandy places on undulating eens Wilczek describes it as "frequent". It has been veto # n green mead- ows and sandy deposits, in arroyos, and on campos, at altitudes of 200 to 1950 meters, flowering from Sot okee to February, in fruit in J ry Leal reports that it forms mats on undu- lating pampas in Mendoza, preferring s. Herba material has been misidentified and distributed under the name V. sulphurea D. Don and Glandularia laciniata Schnack. The rela- tion of the length of style to the size of t ollen-grains is € po discussed by Covas & Schnack (1945). Schnack an Covas (1945) dis- cuss the Se ee of a hybrid between V. glutinosa and Vv. peruvi =e Britton. Gh eae, Briquet (1900) says: "Cette belle espéce pape Racy icine par ses feuilles linéaires allongées, pourvues ga et 14 d'un ou deux segments trés étroits, sa glabréité, ses piendes visqueuses re- tenant les grains de sable de la pampa, ses longs calices tubu- leux 4 dents intedveateaihen et ses élégantes corolles d'un jaune soufré 4 lobes profondément échancrés au sommet....I1 est singulier qu'un type aussi saillant, comme les précédents d'ailleurs ait pu échapper jusqu'a main ntenant aux recherches des botanistes argentins." He cites Wilczek 5h from Pampa de San Rafael in the Delessert Herbarium. Peredo describes the corolla aa T hot i e that the yellow corolias may fade out to a rose hue; in fact, a Leal 9655 bears a notation to the effect that the corolla is "yellow verging to red" or "cream to rose". Kuntze says: "Ge- rt in die Verwandschaft von V. sulfurea, ist aber durch die — linearen Blatter, von denen shine 100 Soin 2--3 nur einen Blattzehn zeigen, die unbehaarte glu bean se Beschaffenheit etc. recht verschieden....V. lavandulifolia Phil. hat Ce ae Blatter, aber kopfige, behaarte ieiien escenz, kurz jan ge Bracteen, V. decurrens hat viereckigen Stengel, staat er mg Le mit aiecraren & Rand und sonstige Untersc - all, 31 herbarium spec » incl the types of all 1963 Moldenke, Monograph of Verbena 131 by me. Citations: BOLIVIA: Santa Cruz: Peredo s.n. [El Concho, h-I- 1947] (N). ARGENTINA: Mendoza: H. H. Bartlett 19521 (Au--195097, Ca--772)83, Mi, Mi, #-150LSL2: “W-=190h5h3) ; gastel ames s.n. [Herb. Mus. Argent. Cienc. Nat. 36864] (N), sn. wn. (Herb. Mus. Ar- gent. Cienc. Nat. 36865] (N); unease 2 (Macbride ae 17418] (Kr--photo of isotype, N--type); F. A. Roig s.n. [Ruiz Leal 11935] (Z); Ruiz Leal 1952 (N), 6936 (N), 7093a (N), 7468 (N), 7500 (N), 7665 (N), 7665/1k [Goodspeed 3101] Ce Nhat Ww 2322928) , 767L (N), 7671/20 [Goodspeed 31020] (Bm), 769k (N), 9455 (N), 9655 (N), 1038) (N); Sanzin sen. [Ruiz Leal 1h1h] (N)5 Wilezek 54 (W--1h73108), 51h (W==1173108) « Neuquen: F. A. Roig Sen. Sen. [Ruiz Leal 18050] (Sm). Rfo Negro: Calcagnini 39 9 (Herb. Os- ten 19358; Herb. Lab. Bot. Buenos gives ara) OR XVERBENA GONZALEZI Moldenke, Phytologia 8: 27h. Synonymy: Glandularia santiaguensis x pecakii eck & Gon- zalez, Rev. Argent. Agron. 12: 290, in ter. 1945. liography:Schnack & Gon nzalez, Rev. ent. Agron. 12: 290. oe! Moldenke, Phytologia 8: 27h. 1962; eotaaeee: et: Suppl. re e has been proposed for the hybrid between v. santia- guensis fears £ , Schnack) Moldenke and V. perakii (Covas Sets Schnack) Moldenke, thus far known only from artificial produc- tion. The hybrid may, however, be expected in the wild, since the two parental opanlee both grow in the provinces of La Rioja and Mendoza, Argentina VERBENA GOODDINGII Briq., Ann. Conserv. & Jard. Bot. Genév. 10: 103~-104. 1907. ynonymy: Verbena verna var. fissa A. Nels., Am. Journ. Bot. 18: 437. 1931. Verbena goodingii Briq. ex woldenke, » Supe List Invalid Nanes 8, in syn. 1941. hen bipinnati §0oddingii (Brig. ) Jepson, Fl. Calif. 3 (2): as : » 19h3- Verbena gooddringii Briq. ex nraie on. List Invalid Names Suppl. 1: 2h, in syn. 1947. Glandularia gooddingii (Briq.) Sol- rig, Madrofia 15: 50. 1959. ees gooddingii x wrightii Kear- ney ex Moldenke, Résumé 364, in syn. 1959. Verbena bipinnatifida Suppl. 3: 38, in syn. 1962. Verbena gooddingii var. gooddingii Briq. ex Moldenke, Résumé Suppl. 3: 30, in syn. 1962. Verbena Soodengii aeorei Briq. ex Moldenke, Résumé S — - = in syn. ae io : Briq. - serv. & Genév. 10: 103-104... 1907 Prain, "tes ee Suppl. hs ais. . 15335 Ay ste a Am. Journ. Bot. 19: 137. 1931; Perry, Ann. Mo. Bot . 20: 132 PHYTOLOGIA Vol. 9, no. 3 243, 248, epee capes & 355. 19335 Laudermilk & Munz, Carnegie Inst. Wash. - 487: 279. 1938; de, Bot. Jahresber . 59 (2): 1939; Sea Prelim. Alph. ist Invalid Names 9. 190; Moldenke, Suppl. List Invalid Names 8. 1941; Moldenke in Lundell, Fl. Texas 3 (1): 19 & 45. 19h2; ee, Alph. List Invali Names 7 & 51. 1942; MWoldenke, Known Ge eogr. Distrib. Verbenac., fed. 1], 10, 11 13-15; 18, & 101. 1942; Jepson, Fl. Calif. 3 5 J 1916; Moldenke, Alph. List Invalid Names Suppl. 1: 22 & 2h. 19h73 ath N.& A. L. Holdenke, Se Life 2: 43 & 61. 1948; Moldenke, Cas- ea 13: 121. 1948; Moldenke, Alph. List Cit. 2: 38, 439, 52, Mes, 467, 472, h74-—1,78, 480, 482--48h, 493--)95, 497, 498, 503-- 505, 521-528, 579, 58h, 58 Wn @ “~) J VL Se) 7 ‘oO ~] 7 o =" oe ge Ov nN Se) — ee ‘oO = CO — ~~ wo ES > 2 1 3 1210, 121h, 1217, 1222, 1225, 1227-1231, 1237, 1246, 1252, 1253, 1288, 1290, 1292, & 1298. 1919; He N. & A. Le Moldenke » Anat. inst. Biol. Mex. 20: 13. 1919; Moldenke, nea Geo, erbenac., [ed. 2], 11, 19, 20, 23, 25--27, 33, 16h, & 198. "1919; “Yoldenk enke, Am. Wild Fis. 292 & 450. 19493 Mol- denke, Phytologia 3: 1m & & 133. Reni. Moldenke in Chittenden, Roy. Hort. Soc. Dict ard. h: 9 & 2211. 1951; Clokey, Univ. Calif. Publ. Bot. 2h: nies : Om. 1951; Moldenke, Phyto 08 a h: 119. 1952; Moldenke, Inform. Mold. Set thy. Moldenke, Journ. calif. Hort. Soc. 15: 82. 19h; i Mattoon, Pl. Buyers Guide, ed. 287. 1958; Moldenke, Résumé 14, 2h, 26, 29, M33, 32, 223, . 36h, 378, 379, & W7. 1959; solbrig, Mad- rofia 15: 50. 1959; Moldenke, Résumé Syppl. 2: 10 (1960), 3: 6, 1963. 8, 10, 32, & 38 (1962), 5: 3 (1962), and 6: 10. sometimes erb, decidedly gray-green throughout, forming rounded clumps or mats 15 cm. in diamet- er, often prostrate; stems usually several a common woody CMs ly ading-pilose or somewhat —villous, often glan ry branches ascending; principal internodes 3--6 cm. long; leaves de cussate-opposite, 3—5 cm. long, about d ay-green on faces, deeply 3-lobed or 3~cleft and softly canescent- pilose or less villous—hirsute on both surfaces, tapering at the base into a short margined petiole, the divisions broadly oblong, about m. long, or sed to deeply promin ‘ florescence terminal, pedunculate, fascicle-like during anthesis5 peduncles 4--8 cm. long, stout, softly cinereous-pilose or vill- ous-hirsute with white — bractlets lanceolate, about 7 mm. rani a little shorter than the calyx, acuminate ab the a pec coarsely spreading-pilose or tee us-hirsute throughout, ciliate on the margins; flowers corymbose during anthesis, "the 1963 Moldenke, Monograph of Verbena 133 corymbs to 2.5 cm. wide, the spikes somewhat or greatly elongat- ing after anthesis and then about 5 cm. long and 1.8 cm. wide; lar, its tube about 7 mm. long and rather prominently costate, the teeth unequal or subequal, slender, lanceolate-subulate, a- bout 2 mm. long; corolla hypocrateriform, showy, varying from red, fe) 5 4 oa ¢ « g B 0 b B at @ ra + ct Ss ® fw > | ® a K 0, oe a ee A corolla-mouth; anther-glands small; pistil barely equaling the corolla-mouth; fruiting-calyx about 10 mm. long; cocci oblong- t mm ly striat e base, retrorsely gui on the commissure in a white Pest med yd conspicuous ban The type of this handsome species “was polerses by Leslie Newton Goodding (no. 645) -- in whose honor named -- at in the Meadow Valley set nae in mene a is de- ire darker or bluish in drying. ogee’ notes for ton 6266 below, that "gland small on stamen, rather large on the Other, but not as high as the Ei a-oolin, The type of V. verna var. fissa, which is said to differ from BE verna in having pinnately cleft leaves, is Kunz, Johnston, & Harwood 4.252, posited in the Rocky Mountain Herbari The so-called hybrid, V. gooddingii x wrightii palisiiiy, was based on Ethel Palmer s.ne , from San Carlos, Gila County, Arizona, Concerning which Kearney says "Corolla tube too long fe) S0oddingii . " He also says of Eastwood 16979 "leaves exceptionally 134 BHYTOLOGIA Vol. 9, no.3 —— but corolla tube not longer than calyx; possibly a hy- brid h V. wrightii" and of Eastwood 17772 "corolla tube 1 ayy times get: possibly a hybrid with V. wrightii". Of Clover 6383 he says "leaves more dissected and pubescence stiffer than usually”. ee eae Nodwell 7086 he notes "gland rather small, much lower than the anther-cells". Loomis 835 was first deter- mined as & anbrosiaefolia pels then as sv. wrightii A. Gray, but Peebles and Kearn ey comm atent it noalyx minutely glandu- lar, pubescent with the oma Pe aa ped hairs; gland very large, considerably surpassing the anther cells. " Kearney & Pee- bles 10993a and Peebles 11332 were first identified as V. cilia- ta Benth., then as ¥. wrightii; so was Peebles 9186, whose bra alee: are actually lon, ee narrow and which may possibly seeesing, a hybrid with Vv. Prager tpi Verbena rooming has been found by collectors on Rest flats, open brushy flats, and canyon Hooves a0} ane lands and pinyon forests, riverbeds, in clay or sandy eg! ranitic soil, along roadsides, among rocks, on ro slopes and gravelly hillsides, on roadside embankments and grassy slopes, ind sandy washes, dry or sandy bottoms, and creek bottoms, on the sides of ravines, in deserts and desert grasslands, on eather ha. gravelly slopes and rocky limestone slopes, san , mountainsides, desert washes, sandy flats, on canyon floors and stony adobe slopes, in Artemisia tri- ky s monophylla and Juniperus association, in juniper and ponderosa pine forests, in the juniper and yellow pine belts of vegeta- tion, on hillsides under oaks, in sandy arroyos, and among the browse-type vegetation, at altitudes of 665 to 5670 meters. It has been collected in anthesis from February to November, and in fruit — March to October s it is found in sandy soil and washes and o tains. Heller states pra the species is "occasional in orev places in juniper and oak belt of the Upper Sonoran Life Zone", Ferris found it "associated with two species of scrub oak, Cer "frequent in washes". Benson says it occurs in the "lime stone soil of an desert vegetation type", the "Sonor~ an desert vegetation type", "in s and gravel in the Lower YT ons n bottoms"; Brass reports it "frequent in damp soil of floodplains"; and Tidestrom says that it is "occasional on slopes". 1963 Moldenke, Monograph of Verbena 135 In Arizona it was found by Demaree in "small fair habitats on dry red ridges". Cutler found it "frequent in pastures" in New Mexico. Wold discovered it in the Lower Sonoran zone "with Joshua trees, Salvia carnea pilosa, in sun, in dry sandy gravel in wash", "in bilge Upper Sonoran life-zone with pinyons and junipers, common cky wash", "scattered in Upper Sonoran life~zo ne with h sunny hillside in deciduous red-browm volcanic rock." revit found it forming groundcover with Helenium hoopesii, Delphinium, Artemisia, and Pteris in pine forests. J. Goodwin Goodwin 63 appears to have had white flowers and may represent a white-flowered form worthy of bisiantidadead de signation. Edw. Palmer 339 is also said to have had white flowers ommon names for the species are "southwestern verbena" "southwestern vervain". and "vervain". Material has been takaee. tified and distributed in herbaria under the names V. am rosiae- folia Rydb., V. ambrosiifolia Rydb., V. aubletia Sher V. aublet- ia Willd., v. aubletiana Willd., V. bipinnatifida Nutt., V. bi- Pinnatifida v var. Pee Perry, alk canadensis (Le) Britton, ve. ciliata Benth, , V- ciliata var. longidentata Perry, V. ciliata var. pubera Pith: } Perry, V. gooddingii var. nepetifolia Tide- str., V. longiflora Lam., V. remota ne V. stricta Benth., V- wrightii A. Gray, and Verbenia wrightii Gray. On the other hand, H. S. Gentry 1312, disteibiked as V. good- dingii, is actually v. “delticola Small; Herb. Chiricahua , Veg .- Study Team 1958-118 is V. ambrosifolia f. eglandulosa Perry; ry; Mc pasts 25 175 & ae are iv ciliata var. longidentete F Perry sGooe G. W. Gultion 4i/, it pee i poae am posbimaneaal Tidestr., os Solbrig ” 2803, Wigeins ee: as DPE oo goodding= ii, is v. patent Cee Nutt. As furth Vidence of how this species has been misunders eetooe by herbarium workers in the past -- Eastwood 17119 was first i dentified as "V, ambrosiaefolia Rydb.", then as V. wrightii ney Kearney; H. H. Rusby 337 was determined by the collector as V. bipinnatifida and by eee as V. wrightii; Kearney & Peebles 10993 & 11118 were first determined as V. ciliata, Zata, then as V. mrightii; while S. W. Hutchinson s.n- [Kay 1929] was annotated by iongiflora, . bipinnatifida, ‘and Glandularia s . Palmer s- Rn. [southern Arizona 186 was first identified as V. aubletia, Grit wrightiin, Of H. He Rusby 337 Kearney says "I make this wrightii", 136 PHYTOLOGIA Vol. 9, no. 3 of T. E. Wilcox 26 and Zuck s.n. (Tucson, May 16, 1896] he says definitely ey, wrightii", while of Eastwood 18032 and T. E. Wil- cox s.n. [near Williams, April 11, 1905] he says nv. wrightii?”. Of Tidestrom 772 Wiss Perry says Mot typical", while Kearney says "V. wrightii?"; of Griffiths 35 Kearney says "V. wrightii?} while Perry says "cfr. V. gooddingii". Of Lemmon & Lemmon $.n. (Peach Springs, June 1884] she says “toward var. nepetifolia". Of Kearney & Peebles a Kearney says "Intermediate between Wright- ii and Gooddingii On the label of c T. Mohr s.n. (Gila River, Feb. Mar. 1873] be saison says: “This species has been recently eye to gardens under the name of Vv. montana. It is very variable in ri oliage an mene " He is here referring, however, not to the oopres comprising his nei but to the name whic h he assigned o it, V. suhathie: Both the name Vv. Piven teines and V. montana Hort. are synonyms or Vi. Ce CLé )B Britton, a species which is, indeed, "very variable in foliage and h item Kearney & Peebles th Tragia ramosa Torr., while Nelson 2 & oe 12g in the herbarium of Catholic veri was mounted wit h Fox oS which is V. ciliata. The Collector un- designated s.n. [Partridge Creek, April 26] is a mixture with V. ciliata var. pubera (Greene) It should be pointed out that tise Newlon 653 from the Santa cited below, could be from either Pima or Santa Cruz Counties, Arizona; Dudley 10, from the Hualapi Indian Re- servation, cited below from Coconino “gaa could as well be from Mohave County; and Eastwood 17119, from the Mazatzal }oun- tains, cited below from Maricopa tee could be from Gila or Ceverat County. Miranda & Deaver 5785 has a label reading "Mo- have or Yavapai County". The Blumer 1345 collection, cited below, was cited by Pe as V. bipinnatifida._ Nattoon (1958) notes that the seeds of V. gooddingii are offered to the horticultural trade by Clyde Robin. Perry (1933) cites the following 20 additional specimens not as yet seen by me: NEVADA: Clark Co.: Goodding 645 (E--isotype, F--isotype, G--isotype). ARIZONA: Cochise Co.: Carlson sn fSulphur Stee Valley, June 3, 1915] (W); W. F. Parish h 196 (4), 197 (G). Gila Co.: Goodman % Hitchcock 221 Tey, 3 Maricopa Coe Bartram 255 (D). Mohave Go.: Kk. 5. Jones s.n. (Hackberry, 52 153] (F). Pima Co.: Hanson & Hanson A. rere 21 (E, F). Yavapai Co.: Coues & Palmer 498 (E); W. W. Jones s.n. ares Divide, 25 July 1921] (E). CALIFORNIA: San Bernardino Co.: Edw. Palmer 337 (E, F), sen. [Rock Spring, May 1376] (G). San Niezo Co.: Edw. . palmer s.n. [southern part of San Diego Co., 1875} (G). MEXIZO: 30: Baja 1963 Moldenke, Monograph of Verbena 137 California: Orcutt s.n. [7-25-1883] (F). She says "In habit V. Gooddingii is much like V. ciliata, but differs in the villous- hirsute pubescence and the larger flowers. It is readily recog- nized in its typical form by the long-ciliate character of the bracts, the short dense villous spikes, and the shaggy pubescen- ce." The Mearns collection from Fort Verde, 1881, and the Mac Dougall 603, which she cites from the Britton Herbarium, are re~ garded as var. nepetifolia by me, and the Edw, Palmer 339 which she cites from the same herbarium, is also regarded by me as var. other herbaria may not represent the Palmer's numbers, assigned largely by Asa Gray, are notoriously full of mixtures. all, 446 herbarium specimens, inc type material of most of the names involved, have been examined by me. Citations: ALABAMA: Bibb Co.: C. T. Mohr sen. (Mi). UTAH: Washington Co.: M. S. Baker 837 (N); Eastwood & Howell 12h2 (Gg- 206895, N); F. W. Gould 1386 (Ca—8826Ll, N), 1730 (Ca--882643, Gg, N, Pl--156720); Maguire & Richards 15921 (Ua—21325); B. Mc Allister s.n. [May 15, 1938] (Ua--),717). County undetermined: J. =. Johnson s.n. [s. Utah, 1876] (W--56196); Purpus s.n. [Cave Spring, May--Oct. 1898] (Ca--10)881). NEVADA: Clark Co.: Clokey 7282 (Ca--900551), 7283 (Bl—L2304, Ca--900553, Ca--909945), Du-- 269985, N, Se--l7h6, St--1911h, Ua--31276), 728h (Ba, Ca—9099L,3, Du~26995, N, Pl--12h972, Se--7605, St--19129, Ua--31283), 7686 (Ba, Ca--598,72, Ca--900554, Du--258943, Gg--257095, Hp, I, La, N, Ok, Rs—-20239, Se--7852, Se--52083, Ua-——253L9, W--1729918) ; Goodding 645, (Ca--508 7--isotype, Du--91127--isotype, N—isotype, Po--6 3875——isotype . W--l85 355--isotype); G. We Gullion 26f (Ca— 195471); Jaeger s.n. (William Ranch] (Po--159913)3 Purpus 6061 (Ca—10816, Po--63967, W--348177); Train 1458 (Ca--900555, Du-— 287457, I, N, Tu--39539), 2009 (N). Lincoln Co.: M. E. Jones S- n. (Calientes, April 29, 190] (Po--71255). Nye Co.: La Rivers & Hancock 01 (Ca—-900552), COLORADO: County undetermined: C- C- Parry 551, in part (W-——-1323045) . OKLAHOMA: County undetermined: G. W. Stevens 36 (Ok). TEXAS: Brewster Co.? J. Fe Brenckle 51083 (Le); Nelson & Nelson 5025 (Ca--71030h, W—1974036). Uvalde Co.: M. E. Jones 28301 (Po--130h27). NEW MEXICO: Catron Co.? De — R. Goddard 32© (Ca--505h36). De Baca Co.: A. Nelson 11307 (S)- = G Mar. 1873) (W—-77187h) « Otero Co.: Cutler 2002 (Ca--8628)7) . County undetermined: H. H. Rusby 337 (IMi)- ARIZONA: Apache Co. F. M, Chamberlain 56 (W--562805)- Cochise Co.: L. D. Benson 10301 (Po--2676L8, Tu—35541), 11110 (Po--267657); Blumer 1345, Part (Du--9535, Se--92553); Borell s.n- [Bonita Canyon, Sept. 8, 1935] (Ca—-77136h); J. I. Carlson S.ne [May 20, 1915] (Ge-- 138 PHYTOLOGIA Vol. 9, no. 3 31279), sen. (Sulphur Springs Valley, June 3, 1915] Sc eis S.n. {south | th of Warren, June 3, 1915] (W--873380), s.n. [south Warren, June 7, 1915] (W--873381); O. M. Clark 8120 (St); terre 4579 (Gg); R. S. Ferris 9933 (Ca—882569, Du--27806h); Foster & Arnold 206 (W--18266)6) ; Goodding 38-50 (Gg—365609, St), 1972 (St), 2231 (Ca--130090); Goodman . & Hitchcock 1211 (Du--192000) 3 Gould & Pultz 3154 (Tu--21550), _ 3169b (Ca--705200, N, Tu--21586); W. We We Jones es S.n, n. [Dragoon, Aug. 18,0 1920] (ca--l079 3h) j Kearney & Peebles 's 15117 (G, (Gg—l18631); Lemmon s.n. [Huachuca Mts., Aug. 1 Aug. 1382] (Bm); W W. F. Parish 190 (Du--90889); W. W. Price s.n. [Apr. 21, 189] (Du--97120) ; in: A. Thornber 2981 (Tu--355hh); Wiegand, Ma-~ guire, Richards, & Moeller 11150 (N), 11162 (N, Ua--47412); C C. Be Wolf 2490 (Ba, Rs--9998) . Coconino Co.: Benitez s.n. [Sedona, 5- 7-50] (Fg--3i60); Clover 6383 (Gg—325858, Mi, Tu--18626); Darrow 3112 (Tu--22019); Deaver 7 (Fg--735-374-111), Mr (Fg--73h-37u- 10), 91 (Fg--797-374-111), 290 (Fg--871-l60-111), 1527 (Fe), 2483 (Fg--2837), 3135 (Fg--3903), 3232 (Fe--3979); Demaree 10269 (Ca), 40973 (Gg), Lo991 (cg), 42557 (Ge), 4310 (s)} Dudley 10 — (Gg--191651); D. R. Goddard 525 (Ca--505526); A. A. Heller 15794 (Ca—726072, N, Se--8 7:92) 5 Jaeger Son. (Flagstaff, Sept. 1723] (Po--6 3852) ; sisi s.n. [Midgley Bridge, /12/50] (Fg—-3459)5 Nelson & Nelson 2082 (Ca--58030h, N, N, S, Ua--23757); H. H. Rusby 337 (W--771879) ; Toumey s.n. aad pevine Cafion, July 30, 1891] (W=-56195); Whiting ng 1047 [cat. 926] (To); T. E. Wilcox S- n. [near Williams, “April - 11, 1905] (W--562357). Gila Co. Collom 21 (3). oh. (Collom Camp] (Mi, N); Crosswhite & Sands 788 (Hi-- (Hi-- 194697); Demaree 13717 (S); Eastwood 16635 (Gg--16199k), 16979 (Gg--167367); Foster & Arnold 26) d 26h (W--1826662); Goodman & " Hiteh- cock roam {Herb. Drushel 8501] (C: (Ca--26359, Du--191998, aa 183906, it » Mn--17980, N, Ur); J. Goodwin 63 (Fg--2253, patie Pics Fr W. Gould 1,707 (Ca--773726); MacFadden 105 "(gn —3169) Nelson & Nelson 1832 (Ca--580305, I, N, S, Ua--23758)3 Ethel Palmer s.n. (San Carlos, April 1932] ee noses), Shreve we 7u51 (Rs—-1479)) . Graham Co.s Toumey s.n. [(Galiuro Mts., J “july 29, 1894] (W--619182), s.n. (Galiuro Mts., July 29, 1895] (can=51305 C. R. Van Cleve 13 (ray. Greenlee Co.: Gould & Haskell 1,023 $y _— re ee Eastwood 17119 ing (ce~-167368); F. R. Fosberg S.1)39 (Up); te ee W. Gillespie 5365 (Ba, Ca--l9972, Du--21 797, N, ary elk 1620207); Kearney & Peebles 10993 (To), 10993a (To); Lawr Surber s.n. [3-27-60] (Hi--19h9l); Peebles 6732 (N); raskins & Loomis 6732 (Gg-—-192659, Tu--35526). Mohave Co.: S. B. Benson 02 (Ca——526616) ; Braem 793 (Fg—9267), s.n. (Apr. 10, 1927] (Du-- 190298), sen. (Apr. 1927] (Du--190297) ; Demaree Lls80 (S); East- Mapping by Cc a . > . y counties done by Andrew R. Moldenke . —— . : Q9 Metri : f ‘3 cooddingii in the United 4 | é aE ] e rT Figure 0. Distribution 0 ena goo & in the United States 1 2 3 4 5 6 7 8 9 10 it 11 ] 12 T T , — : | 2 l 13 14 | 15 if 16 T 1 T T PY T 9 +) T > T T S 19 =a 20 21 22 23 24 | 2 6 2 iN y = SEE EEE —— __ 4A 2 L ) 31 I 2 T = T as T = . % ff ; | \ | } | R , T “* . ° A Is . A | } va a % L BE H i | EN a” Coy ey — oO L U R T A | Ss } 1 8 ~h3= “1S... 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ASHU 6 : . me pany Dees — cust, ca a ese co as ae 2 om 2 . os a 2 ‘es REID FLORENCE Cc | ‘ N ~ — | MARSHALL BURNETT wan vase ot < os 7 7 FJ ne Sumy =e “a og D 4 seprton Ae Como Tncour _ 4 é Vj “y ee com _— Kon a Mexaay A ‘cee TS ie " EDMUNDS Dy 7% aca # an os G : oo. 4 5; afe Mercy SHERIDAN a e . ~ g bute 7 TAYLOR m oa SA et ¥ = somes man Soran | eas oom 3 Dooce wOnROE x see | use —ee\ 0 SANBORN | MINER | LAKE | MOODY PIPE-| Munnar i : wast =a a —— oT ls emit Cs e uncon) SusueTre CUSTER JACKSON TOME cortowsoo0 [waTowman a crt << oO wound 4 Teeny oan TW Fuis - Power AriB0 NATRONA r BRULE [AURORA La eae oe fence | Fond DU at =r Cone Pn) N se ‘ Y CONVERSE ‘SHAR wna ap aon vanson] uccoon | MNNERANA “© BOCK | ROBLES [JACKSON | WARTIN | FARIBAIKT | FREEBORN wok —] =a [oooet wonTicn ee ar (un Frome a rl aa io) Ney — | O M bisa FALL RIVER TRIPP Ln pg! CMAN i 2 — wn ‘ a sf s I N . BENNETT Cranies] DOUGLAS ios . . 4 & s aay mn ene Z 0 nn om jon ~ orn ° f CHELL 7 : a ble Tt) "a eee La G = wx ictaaceadl| Wai a. sczon [ocomson [emct[aassuman| | WT ae on wun _/ pant ware keel 2 o asus ou ‘ & age? ess "ona . i at ma Hy =] reecstom = / 2. 7 - : L saa Tan Uncounty 200K Temen | cur pPAOAT HANCOCK oe flow yous _ er H 7 a on Tarom ¢ & . orn cc =x, e "ame 2 ” e HOMME cuy jury GaaaT TancwoaT] RACINE at weit on SWEETWATER '* C= * cnten | ROCK < aaroue | ‘ ip PLYMOUTH TOREROMEE] BUENA PrO=mOrT=s womancor] WRIGHT | FRAMMLIN BUTLER BREMER (AFAYETTE oy ooo \™ _& A mas ® wor : wn oN . s = ee ag : en a ceoae C x Nene : fone ; P onon?,* WEBSTER a rps Cle aie ‘ woesat | Use ae] Oy ral" = Ll Leribe tes sae — h K 2 : \e be I] ¥ — hag Ra) i ! Suse . = ™ "acer ms Sur Le sth 2 _ ; w E ie UT aw wmasay — A ro © Comes cH nance 2 uaa V s = BUTLER WERCER _ m7 Pon TvaeSTOM wrth ~ Ok . SHALL 3 fel an me = P ‘dis say S wis a maan Tecan uucaS [monn] wareiio | 77e==8 vont =e eres wmoquas Ca oe Dyn ee (1 want Mawar | Mone 2 SEwaa | wancasres cass vs? : iy bal y. * A wang amo ar Tt arioa Temoamio| ocarun [ware prmmoms|oans |e Fe fi _ cm , hey "Vee YY Ste . ner, cay mm | 7 me Raay - pO Tcrcent mesrtt SOTA cy ] ” GAGE NODAWAY worth a i \ : U T Eatery ee cENTRY SULLIVAN Town Tinox =o Gus ~~ _ ar, A a 4 tag ay, H on ‘Maoe ny hy TMA ag Sven = jomne M \ . jeans > — ™% Purr e 0 Bee ae ware = “ * tem Sa juan rT) > Tue N bes? — tall! ames 8S Cass mm 0 7 aan = “Sameer MFALFA | GRANT may OSAGE IMOWATA] CRAIG Ora conten __] BUTLER g ud s Baaey — wry TODDARO voaer \e “4 on) ff 3 L aim iia < 2 "i Taney eam ne . me eae? ap a saggy © 3 a “sn ca aT * eamuey DALLA - Ld . . ELLIS | wOOOWARD 2 seas STONE . eee. Ocont Ad WENRT WSO, saath eo ‘3 bd MV >> mn be ‘one <3 en a) 2 seman Trawsrono Joon ter Turscome warLD | OLE 5 Trocars a ie -_ 2s ST eer , wusaer Ao Coo SOR \ sf our 2 =o som oS BS ee st oe P (4 maeos | SANTA FI et Pt ae ed Naaetacn canon, [acone aon waren} . a ene ona sont cg ‘g fn on ai ane) J a Gree vee ARROLL aes PF N ve - one i ‘SHARP a Cad Tel sf stNAT om = o wamer ‘moon | MTCImmSoN Dewey Bane | nncrisnen | L — ‘ LAWRENCE onmm ecm wu H 5, e AS ws ob hot ak : vat ‘om RoseRTS | MEPL GaN CREEK oe Wtsuncron manison newTON san S 7. fea me Jos sw) we is aot ma ace ts 7 = VEtsicg “oem _ nOGER MLS. UaNCOUN nn PDE PENDENCE ; om ; Bae aa i) S D ean Corry ] — porrer = Carson os cusTER NUL GERY MUSKOGEE Anan Seam JORNSON] POPE | vaN BUREN [CLEBURNE —_— d cet ¥ N GONE (8) ie A H A ‘SEQUOYAK ra cous TOA wom Om ANC ‘CONWAY E W OEAF Sani} Guar a no MeiNTOSH 1 cu ys * eH y \ a smusTrom | Comfy faceasmorn i — HASKELL] : wits cay 3 0) vo od Se ens = as Aga Ar on Pammen | casTeo oe i — a me h4 Bmscoe | HALL —— i COMANCHE PowToToc a meine iy Raimi Q (UNCOUN ~ ° ee STEPHENS Forks ‘PUSHMATAMA POLK Lgncue - maMey | ane McCURTAIN - TEFFERSON | ARKANSAS oe 2 5 — = wines nos worty corm: | “stem ‘a _— rT _ —— ToT sPminc | [GRANT JEFFERSON JOWNSTON COANOMA eee = Nontoe WAKER 1 a) fi *\ Py R eee Foun wenn *Pom, CARTER ficansna? —— even aa nh = cannounomemsan a8. a “9 hae aa one hee, nn KEYED TO COLUMN A ; " | > e Cocumam | HOCKLEY | LumsocK | Crosser — WLBARcER f LOVE ‘ CLEVELAND ¢ U sien 3 mend u 4 “— Ca) ee gaan - Cauton of Cleartype Index Book as) a ° ed] 7 " — mame , ere J rm) nox Baron ARCHER MONTAGUE come pen . ak 4 2 haa vay ou z cay vr Sy Lee * CAMALESTOR e RED RIVER Pa wes BOLIVAR crore PrCRERS | * me Vos eA P fn & R ramon “ |ruscaconse asagoe |" sot sou) ons i e/ 7. youu | Terry. | Lee = 1 : seen Lomnnt’, A : on : - KENT STONEWALL | HASKELL | ™=0cemoro=] YOUNG WACK wst cana ca DELTA ak ORB CUM prov) EAPO v VU Se ta / ~ Dams Ama —~ ‘ ee Co = ]Mns eapene-) rr Col ou a cee ln = aust ot ne“ g q y ge WORDT STEPHENS [PALO puto | PARKER | TARRANT | 0 a Cam TRUGA wort = ; e ALLS ARS W000 TTPSHuRY MARION a NESHOBA PERRY ® ~ OT aaa ae : oe ~ i God saat wsencd | [DALLAS ous we q ss = actmact f Ysemr Phage a a 1SOR ge ha Lt eee ee Reon | Ne i é pare TRADE MARK REG. U.S. PAT. OFF. 8 ee CALLANAN | EASTLAND Tear M000 | dommson J eutis none smimH ‘ esac) MINDS = Lownniss bind — es on a : a. a al eal cal eel et {tet i * COUNTY OUTL CULBERSON fromey . — EcToe Midna ocx [semne | Sar Ou Taylor os Taveenc = s Tot ‘guTLee pwe) BARBOUR, nine a = ecl shi Som j Tames] collin Tenpun\ COMANCHE WiLL HENDERSON SIMPSON — onto on = BOSQUE CHEROKEE wor Aq Feast = : 1 ii — ‘sean! = « = tomato |" = cou oe mer ." onoem | i) eee ia — eaten) wits HAMILTON (fe LIMESTONE ae ease JEFFERSON TMA] ee me rot ane 4 | = t ~ vers — A a \ r “a a ao Acs ea “j gee [ ERY | GREENE 1 — aneve a ed ee seem | oense ol . Pecos CORYELL NOM s cOWNGTON . —< oO passa : E 4 MeCULLOCH iN $ # HOUSTON aMiTE PORE or e e cov pos YacKSON a a Bole l f Mi T a % TAMPASAS FALLS st SABINE WS VERNON ; eUIRSON : na sania nas Loop . s 4 wanton \ Sea) le 0. iles . Scan 7 See be AVOTELLES? 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A ° OWELL FAYETTE CHAMBERS oF * > ee (Se = Ss K * * amar BOua St MARY ‘A care ORIGINATOR ai SOLE PUBLISHERS ¥ ) ° Dima Guaoaure conzmes Pownce sone ‘ial Re > TERREBONNE \ samt Wiss ‘ if tee Ag Sey ! = = lp een eal ae CLEARTYPE MAPS “Nae #7) COLORPRINT MAPS \ 1 ae DE wer WHARTON BRAZORIA PASCO cal — NEW YO! v s : Fm face: ghey = ATASCOSA MOTICE—This is a copyrighted map. The lw a - KARNES VICTORIA wmaTAGORDA = T a ssn peskbins the lopeeboome ar copyag of Sl, . : or any 7 persion thereol. by say process for Per THIRD \ sai / HA w A I I oummat ASME MeMLLEN Bee QOD =, — . 1 : é * / wane cncxcneest Copyright < | w ‘. JUDICIAL DIVISION * a HONOLULU (Ait om oe pyright, American Map Co., Inc., New York, No. 14324 KAUAI 4 il a OUVAL mms mae —— | <4 im WEL gua —— ort U ¢ -. - om ; ; oaHU, euats i , / "a = yenort — a Z / f KALAWAO WLEBERG i FIRST P Ae ZAPATA] id HOGG ome ! SPY ~ ~ git. 7 MOLOKAI cast cue ee a a Jui Scale of Miles 7 . 0 caie of Mies 80 LANAI STARR — 2 rar) WIOALGE ont F rl WiLAcY ‘ ONO KAHOOLAWE - Ps Ls LULU no a PART P a HAWAII > * — ° LISIANSK:, “Ne, ms s ~ “ | - ' = ALEUTIAN ISLANDS . , W~—182003), 91h6 (Du--258595, W—-177508). Nuevo Leén: Arséhe S- n. (Monterrey, 5-1911] (N); Edwards & Eaton s.n. [Monterey, 136] (T); Safford 127) (w--5 bi 1084147); Rose, Standley, & Russell 13449 (N, W——5 35272). Sonora: R. S. Ferris 8770 (Du--229060, N); F. R. Fosberg 7911 (Up); Frye & Evye 2310 (N, Se--65803); H. S. Gentry 356 (Du--263066); Gold- man 292 (N, W—335789); Goodding 952 (W—661707); P. B. Kennedy 1033 (Ca——37359k, Du—217703, Ge—157785, w--128720h); MacDougal 73 (W—57427h); Mearns 190) (Du--2)196, W—-232443), 1914 (Du-- eh197, W--232672); R. Moran 028 (Du--361696); Edw. Palmer 307 (W—56203); Rose, Standley, & Russell 12745, in part (W--635561), L27lba (N), 12840 (N, W--635660); Shreve son. [Apr. 8, 1932] (Cm); Studhalter 13h9 (W--1685673); Wiggins 6382 (Du—253uL2, W- 1739923), T17L (Du—251391), 7225 (Ca~-666016, Du-—261197, Rf, wm 177238), 8201 (Du—264L52, Wi, W—1821,373), 11775 (ca--9315 UNTED INGS: “Wash W--2261,369) . Fo : F. 38: 15. 1925 (m, W). roc. Biol. Soc. Wash. 38 1963 Moldenke, Monograph of Verbena 1h7 XVERBENA GOODMANI Moldenke, Phytologia 6: 330. 1958 Bibliography: Moldenke, ‘Phytologia 6: 330. 1958; ‘Voldenke, Ré&- x 26, 365, 375, & hn. 2 s "4s a ees ag between Vv. stricta Vent. and V. halei $9 with inte pe cnc characters. The leaves are various usly in- cised. and lobed as in V. halei, but are densely canescent—pubes- cent beneath and to 7 cm. ca and 4.5 cm. wide. The stems and ranc he type of this hybrid was collected by George Jones Goodman ia. 64,7) -- in whose honor it is named -- along a roadside 3 miles north of Wi illis, Marshall County, Sor aaa on June 28, 1957, and is deposited in the H. N. Moidenke Herbarium at Yonkers, New York. The collector notes that the hybrid is infrequent or Moldenke and V. simplex x stricta Gates. In all. ~20 herbarium speci imens, including the the type, te, have been examined by me. Citations: OKLAH HOMA : Marshall Co.: Alder s.n. [July 10, 1950] (0k); G. J. Goodman =<. ped os (Ca—81915, Ok, Ur, ur) 647k (Bi--isotype, Ca~-iso -~isot: ype, S--isotype, Z——typ perm isotype, Z-~Asotype), Gu6i ina eg 6517 (Bs, lm, Ok, a. deep Senlanis pre eae: the angles, rather Laas scabrous— pubescent with short whitish hairs; nodes annulate; princi in- ‘ernodes 1. ——5 cm. ot leaves decussate-opposite, sessile; leaf-blades firmly c ceous but fragile, grayish-green on both Surfaces, elliptic em. wid and b vO. teeth, eek Scabrous-pubescent above (the hairs soon deciduous, leaving th e lamina white-pustulate) and subbullate, appressed- Sabakeesn on the venation beneath; midrib slender, * impressed a- very prominent beneath; secondaries slender, 5--7 per side, impressed above, very prominent beneath; b ng, oats at the apes ages long-strigose and also more 148 PHYTOLOGIA Vol. 9, no. 3 The type of this species was collected by Goro Hashimoto (no. 663) at Rio de Peixa, Municipio de bere Gofas, Brazil, on Aug- ust 7, 1949, and is deposited in the herbarium of the Instituto Botanico at So Paulo, Brazil. “pes all, 3 herbarium specimens, in- cluding the type, and ‘3 mounted ste AN have been examined by Seca —— Gofas: Hashimoto 663 (F--photo of type, N—~ iso to of type, Sp--5532——type, Z--photo of type); i 858. aera 5 Mer. ate (Cham.) Herter, Revist. Sudam. Bot. 4: 186—- 193 ynonymy: Verbena officinalis? var. gracilescens Chan., Linn- aea ts 254-255. 1532. Verbena officinalis g Hook. apud Cham., Linnaea 7: 255, in syn. 1532. Verbena officinalis p gracilescens Cham. ex Schau. in Mart., Fl. Bras. 9: 191. 1851. Verbena offic- inalis var. gracilescens Cham. ex Kuntze, Rev. Gen. Pl. 3-42 (28 257. 1898. Verbena officinalis var. Spiné4 pebecs (Cham.) Kuntze ex Seckt, Rev. Univ. Nac. Cordoba 17: 90. 1930- eae graciles- Gens Chan. ek Moldenke, Prelim. Alph. List Invalid Names 6, in syn. 1940; Maria, Pl. Vall. Cochamb. 38. 1962. Verbena "offieina- iis var. gracilescens Hauman-Merck apud Gray Herb. C Index ne a [191 or before]; Moldenke, Alph. List Invalid Names Suppl. 1: 26, in syn. 197. Verbena officinalis var. "b" gracilescens verbena gracilescens vypica ke, Alph. List Invalid Names Suppl. 1: 2h, in syn. 1947. Verbené officinalis var. gracilescens Griseb. ex Moldenke, Alph. List In- valid Names Suppl. 1: 26, in syn. 1947. Verbena officinalis var+ major Osten ex Moldenke, Alph. List Invalid Names Suppl. 1: 26, in syn. 1947. Verbena offi cinalis var. tenuis Cham. ex Moldenke, Alph. List Invalid eran Suppl. 1: 26, in syn. 197. lores eee escens (Cham. & Schlecht. y Hackes ex “appar Résuné 365, - 1959. ae gracilescens f, glabra Osten ex eights ay 365, in syn. 1959, Verbena gracilescens f. subglabra Osten ex Moldenke, Résumé 365, in syn. n syn. 1959. freer gracilescens (L-) Briq. ex voldenke, Résumé Suppl. 3: 38, in ¢ 1962, Verbena officinalis var. minor Osten ex Noldenke, Résuné Suppl. 3? 10, in syn. 1962. eeiesioes gracilescens Cham. & siiiaatt. ex Lia: «1963. Résumé Suppl. 6: 10 Bibliography: Chan. » Linnaea 7: 25)——255. 1832; Griseb., Abh. K. Gesell. Wiss. Gotting. 2) [Symb. Fl. Argent.] 276. 18793; Lor= entz & Niederlein Exped. Rio Ne 1; Fl. Tucuman 9), “ibeay Kuntze, Rev. Gen. Pl. 3 (2): 257. * 1898; 1963 Moldenke, Monograph of Verbena 19 . Soc. Cient. Argent. 53: 2h9 Dove Add. Fl. Gs rhe "19025 | Chod., Bull. Herb. Boiss., sér. 2, 2: 817. is02 ae Plant. Hassler. 9: 196. 1902; Hicken, Chloris Plat. Argent. 197. “a Hauman-Merck, Anal. Mus. Argent. Hist. Nat. Buenos Aires is lak 1913; Hicken, Physis 2: 11h. 1916; Seckt, Rev. Univ Ha. Cérdoba 17: 90, pl. 18, fig. 86. 1930; Herter, Revist. Sud- am. Bot. h: 186. 1937; Gallinal, Bergalli, Canpal, Aragone, & re igs 2 Stud. Nat. Mead. Urug. 81. 1938; Herter, Revist. Suc- s - 6: 939; Moldenke, Prelim. Alph. List Invalid Name 46. 19h0; J. A. Clark, y Herb. Card Index [1941 or before]; onese, 5: 413. 191; Moldenke, Lilloa 6: 328—330 g arwini (1941) and 8: 430. 192; Noldenke, Alph. List Invalid Names 7 & <, Anal. . Fitotéc. Sta. Catalina : 19 & 20. 192; Moldenke, Known fe Seah, Verbenac., [ed. 1], 39--h1, 4h, & 101, 1942; Rosengurtt, Estud. Prad. Nat. Urug. 3: 236 & 237. 193; Herter, Revist. Sudam. Bot. 7: 225. 193; Schnack & ey Darwinian 6: 470 & 473. lols Moldenke, Lilloa 10: 3h6— 347 & 384. 194; Schnack & Covas, Bol. Soc. Argent. Bot. 1: 26h. 1946; Moldenke, Alph. List Cit. 1: 12, 29, 80, 86, 92, 202, 217, 219," & 296. 1946; Augusto, Fl. Rio Grande do Sul 212 & 233. 19h6; Rosengurtt, Estud. Prad. Nat. Urug. 5: [Fl. J. Jackson] 39h. "196; Moldenke, Alph. List Invalid Names Suppl. 1: 22, 2h, & 26 191173 Moldenke, Phytologia 2: 338. 1947; Ragonese & Govas, ae pall LS, LS1, LSS, & 487. 19475 Hill & Salisb., Ind. Kew. Suppl. 10: 242. 1917; Boag tear Castanea 13: 119. 19L8 Moldenke, Alph, List Cit. 2: 358, 36, 371, 37h, 375, 378, 380, 426, 458, 532, 533, 537, “ste, 586, 599, 627, & 628 (198), 3: 662, 672, 673, 732, 733, U5 79, 765, 767, 780, 781, bol 859, 862, 869, 876, 9077 911; 913, 923, 931, & 966 (1949), and bs 8, 1090 1100, 12 5 = i R 115, 118, 119, 127, 357, 364, 365, 371, & W7l. 19595 Maria, Pl. all. Coch abamb. a's 1962; Voldenke, Résumé Suppl. 3: 1h, 38, & LO (1962), bh: 17 (1962), and 6: 10. 1963. Tlustrations: Seckt, ares Univ. Nac. “eérdoba 17: pl. 18, fig. 486 fas v V. offi cinalis] . Annual, biennial, or soe aah and suffrutescent, branching herb, 20 cm. to 1 Me tall: leaves decussate-opposite, the blades rarely linear; flowers small or very small in loose spikes, the cl rom li os dark-blue, rose-blue, lilac-blue, or violet-blue to violet, Clear-violet, pale-violet, bright-violet, dark-violet, violet- core lilac,’ clear-lilac, blue-lilac, or * purple, occasionally white, chromosome number: 2n = The species has been found by collectors inhabiting wet mead- 150 PRY? 6.0 G7% Vol. 9, no. 3 ows and fields, the shores and banks of rivers, wet places along canyon places and ti fields, wet ground among spiny shrubs, low and 8 ng every month of the year, Sad in fruit in August, sopesben ecembe ec Meyer says. that it is a "weed in cultivated places", and Fries states that it grows in eg grassy places in the region of much rainfall. Scheettae calls it a "planta pester while Ruiz Leal erage it "en alfalfares Siac: Eyerdam & ete gay that it is common weed" in Jujuy, while Cabrera G91) says of it: "ulig- inosa ere en los baflados pacid s", nade : Govas (19)7) "Planta bienal o perenne, erguida, ae baldfos, ee etc." and cite Rosetti, Covas, & Rago oe and Covas & Ragonese 377h. Ragonese (191) cites his no. “3088 from Santa Fé; Augusto . (196) cites a Sellow n, from Uruguay and a Kadlets s.n. from Rio Grande do Sul. feat (1943) says "Hierba hemic Sita, florece de fines aa principi- as aja e bosque; es escasa y desaparece en los campos muy trab- ad isso's original Seen (2832) of this plant is worth repeating Sea "E Montevideo mi t Sellowi Wius . Ab europaea "stirpe iv ormiun al sa: no irm ens hujus vilioris eregis proponere speciem. Confer. Hook. bot. misc. 2 Oa. Vv. 0 » quae cum gen in campestribus Americae aust walioris peopenisns. foliis ut etsy nec vero gracilitate et tenuitate partium um differre dici- ur Common names recorded for the species are "camar adinha", "formosa sem dote", "jurube epa", "yerba de la golondrina", "yerba e Santa Ana", and "yerba de Santa Marfa". On Fries 27 a note shaken that the flowers were "rhite or pale-blue"; ae te-flowered 1963 Moldenke, Monograph of Verbena 151 specimens are also represented by Byerdam & Beetle 22553 & 22577, Sota 85, Venturi 5504, and Wall & Sparre s.n. (12/11/46]. Lillo describes his no no. “11996 as "forma foliis strigosis". Archer 627 has linear leaves, perhaps because the plants were chewed ewed off by grazing animals and the specimens represent — growth; it was collected on the banks of the Rfo Paran4. J. West $231 is a mix- ture with V. parvula Hayek. The species is 2 to be mentioned by Martinez Crovetto & Piccinini in Revist. Invest. Agric. 4: 30 & 78 (1950), but this is not true, nor does this name or nyerbena officinalis g" appear in Hook., Bot. Misc. 2: 16 (1831) as it Claimed in the literature Herbarium material of v. gracilescens has been misidentified and distributed by various collectors and museum workers under the names V. caracasana H.B.K., V. aghadsdtden Cham., V. fili- caulis Schau., V. intermedia Gill., V. litoralis H.3.K., V. lit- oralis var. caracasana Briq., ve. prea a th, V. littoralis var. caracassana Briq., V. officinalis L., . polystachya H. 5.K. On the other hand, the Jtrgensen 3767, taiaeeimieet as V. officinalis var. gracilescens, is gray: in part V. brasilien- sis Vell. and in part Ve litoralis H.8B Gd 5 should be noted that the Ve rnasias of Linnaeus and of Michaux are V. simplex Lehm., ~ while that of Miller is Stachy- tarpheta angustifolia (Mill.) Vahl. Bang 1058 was identified as V. polystachya by Rusby in his published list of collections. The eee Herter 95525, previously reported as V. gracilescens, is et : actually V. litoralis H.B.K., while A. Stewart 3317 is V. glabra- Saeko ee ee Verbena officinalis var. minor is based on Osten 3333 from ynan, dept. Salto, Uruguay, collected on October aber 1B, 1896, and gracilescens Cham. forma foliis strigosis subforma minor, flori- e violaceis." His V. gracilescens f. typica is based on Osten 3936 from the shores of the Rfo Y near Durazno, dept. Dur- azno, Uruguay, collected on cig neem 7, 1899, and deposited in the same herbar arium, described b as "foliis subtus in nervis Strigis obsitis, fl. pallide ceteias His V. officinalis var. major is based on Osten 338 from Dayman, dept. Salto, Uruguay, collected on Rega) 18, , deposited at Lontevideo, de- Scribed by him at first as yr erak is glaberrimis", but later as "V. gracilescens © Cham. forma foliis subtus parce strigosis, pes ob Scure violaceis." The type of V. —— escens f. subgla collected by Miguel Lillo (no. 1 225 in cultivated campos Sees t ept. Capital, , Argentina, at an al- titude of 50 meters, in October, io10, and is deposited in the Same Montevideo herbarium, while that of V. gracilescens f. glabra 152 Pur? 61:00 2% Vol. 9, no. 3 was collected by Teodoro Rojas as Villa si abioa sat on the shore of Rfo Alto Paran4, Paraguay, and is also deposit dat xont sei eee Hauman-Merck (1913) cites ae no. 347 from nto. Negro, Argen- ina, while Martinez Crovetto % Piccinini (1950) cite ihete no. ees from Entre Rfos. Maria (1962) cites his no. 173/ha fron en los palmares del oeste y centro de Entre Rfos; se trata de una especie muy comin en casi todos los bosques en gal lerfa de la re- gién m pot ae eae La floracién se produce en verano.! In all, 256 herbarium specinens, including ae ‘ase of most of the va involved, have been examined by m Citations: BRAZIL: "Rio Grande do Sul: Rambo “ase (Rb), 208 (N). Santa Catarina: Reitz 3287 (Herb. Reitz 5006] (Le, S, 20 2027330). S#o Paulo: Tamandaré & Brade 6692 (Herb. Brade 4693] (N, Sp-—6723). State prenmnes: Sellow s.n. [Brasilia] (Br). BOLIVIA: Cochabamba: Asplund 067 (S, Us); Buchtien 21h (W-- 1159361); C4rdenas 2390 (N, | rey gli Tio 11.205 (w— 1542913); urs. R. S. Shepard zu (Gg--31,02, N, W--1197308). Tarija: Pflanz he tonk N--123), 360); as pans te in part (Ca- 56512h). Province = Sg ae Me ng 1055 [Bolivian } Plateau ] (Bm, C, Pa, W--71999, W-~1h167h1); Stone 161) [Herb. Inst Viguel Lillo 31419] (N). PARAGUAY: Fiebrig 4432 (mm, si--1159389); Hassler 2585 (Ca—929)15), 3391 (N), 7593 (Ca--94436L, N); Herb: Herter 5),5)1 (N, N); Kuntze ssn. [Concepcion, IX.92] (N); Iorong 101) (Cc, C); Pedersen 1200 (%--2122710), 398) (W--2283351), 4072 (W==2.23366):) ; T. Rojas 357 (Bm, S), 1668 [Herb. Osten 8568; Ras- sler 1668] (N, S, Ug), 1883 [Herb. Hort. Parag. 10058; Herb. Os- ten 13550) (ue), 2526 [Herb. Osten 18183] (N, Ug), 10353 (N), S: n. [Herb. Osten 7903] (Ug), sen. [Herb. Osten 13550] (HN, (N, S), SeMe [Herb. Osten 13556] (S). URUGUAY: Arechavaleta 39, in part (Ue), LO (Ug); Perro 39 (Ug), 4115 (N), 32h (x ; Castellanos s.N. THerb. Inst. 1 liguel Lillo cage (i, MN), Sen. (Herb. Inst. nue Lillo 15731] (N, N); Gallinal, Aracon alli, Campal, sngurtt 10 (i; worker ION (ler. foter T2230) (, 2 a9, N, N,N, S, W—122056), 186la [Herb. Herter 9931h] (ome, 1934877), 50890 (N), 83241 (N), Sen. [Ierb. Osten 181,90] (S, UE), SN. fees” bet Osten 13491] (Ug), sen. [Herb. Osten 13)99b] (Us)3 Legrand 570 (Ug), 2008 (Ug), 2370 [Isla de Pichén] (Ug), 2572 (Ug), 3h23- (Ug); Osorio 1003 (Ug--13217); Osten 3333 (Ug), > 333k (Ug), 3936 (Ug), 21700 (S, Ug), 21713 (Ug); Rosengurtt @ Gallinal 5803 (N, Ug—1212, W—-1908¢),3 3 W. EB. Safford 90 (—-922260) 5 Schrtder 17 fics. Osten 19921} (gy; Hugh H, Smith 82 (i-- 1193735); Wetmore 877 (#1066026). ARGENTINA; Egenos Aires: Bur- kart 1.330 (Herb. Osten 22469] (Ug), 10062 (Ca--337); Cabrera 1605 (Herb. Osten 22286] (Ug), 5704 ()3 Lanfranchi 450 (N)5 1963 Moldenke, Monograph of Verbena 153 Miers 734 (W—-13h3919); Seta 85 (N); J. Tweedie s.n. [Buenos Ay- res] (Bm); Venturi 59 [He . inst. iiguel Lillo 31418] (N, S). sherry Venturi 7160 ee Chaco: R. M. Aguilar oa s, N); J8rgensen 267 [Herb. Inst. Miguel Lillo © 31430; Herb. ot 1187)] (N, Ug, ;, W--1055177); T. Meyer 514 [Herb. Osten 22698] Ug); Rodrigo 2537 (N); T. Rojas 12038 [Herb. Inst. Miguel Lillo 107795] (Ca—164,357); A. G. Schulz 1437 (N), 1886 (N). Cérdoba: Castellanos s.n. [Valle de los Reartes; Herb. Osten 15285] (Ug); Cozzo s.n. [Barrio Jardin, Oct. 1946] (N); Galander s.n. [Cérdoba, 15.X1.1879] (N); Hieronymus s.n. [Cérdoba, X.1819] (Br); Hunziker 6318 (N, Si); Lorentz 113 (Br), s.n. (Sg--68312); Troncoso 316 (i, 1 N); J. de Valencia s.! son. [Herb. I: Inst. Bot. Darvwinion 16064] (N, N); Villafafie 172 (N). Corrientes: Ibarrola 1901 (S, W-- 20962), 2169 (S); Rodrigo 665 (N). Formosa: I. Morel lol (N), ie9 (Nn); Pierotti 4076 (Bm), > 4256 (N). Jujuy: Byerdam & & Beetle 22553 (Ca--65561,7) , “22577 (Ca--655659) ; R. E. Fries 27 (N, ate. Garolera & Romero s.n. [Laguna Tala, 10-I-19,7] (N); “Venturi 4893 (Ca--376027, Gg—1582h6, W--15911,50), = po Osten 20799} (ca~376213, E--960),21, erage Ug, W--10827). La Rioja: Cuezzo 1006 (N). Mendoza: Née 4 (a); of te Leal 1018 (N), 3057 (N), 8099 (N) = 8h3 (N), 898 (: (tl). Misiones: “Archer 1,627 (67910, “W--1691996); | Bertoni i 2461 (N); D. Rodriguez 69 [Herb. Inst. Miguel Lillo 31425] (N, W--1802571); G. J. Schwarz 129h (N, St). Rfo Negro: W. Fischer 203 (Cm, N, Ur, W-—70l396). . Salta: Burkart 13192 (N); R. E. Fries 577 (S); Lillo 10996 [Herb. Osten SLELT ( (Ug); D. Rodriguez bh (Herb. Inst. Miguel Lillo 31420] (Ca—3320, N, Ug—l9hl, W--1858261); Schreiter 11,66 (Ca, N); pare 1269 (s); Venturi 54,71 (Gg--159311), 550 (Bm, Gg--159153, N,N, N, S, V2 767, Vi, vi, W— W—1h2820, W--1591h7h ; Wall & Sparre Sen. [nfo Calderas, s/afus) (Ew) ; Willink 72 (S). San Juan: Cu- sass 1285 (N), 1321 (N), Uhl (N). San Luis: Cuezzo 2238 (N); de D. Varela 452 (N, Ok, Rf). Santa Fé: Balegno h95 (N); Burkart ser (N, N), 9083 (w--1858301) ; Job 1048 (N) (N); Ce C. Meyer s-n. Sen. (Te Meyer 3001] (N); Ruiz Huidobro 3080 (Ge—35325L, N), 3182 (1, Ok, Rf), 3216 (N), 3381 (N, N), 3418 (N); Terribile 103 (uN), 455 (1). Santiago del Estero: Cuezzso 2382 (N); Lillo 6185 [Herb. Inst. Miguel a 31423] (N); Luma Risso S* (8), 23 (N). Tueum4n: Eaer 39 (S); R. Ee Fries s.n. (near Tuc Tucur vaman, 16.10 3) (S)5 Lillo 10692 [Herb. Inst. Miguel Lillo 31413; Herb. — 8463] (Du— 317607, N, Ug, Ug—li9h0); Lorentz & Hieronymus . [5--22.xX1I.- 72] (Nr); T. Meyer 3421 [Herb. Inst. Miguel tae = eL0h] (En, N)5 Moldenke & Moldenke 19722 (Es, Lg, N, Sm)5 senate gl [Herb. Os- ten 10878] (Ug), 96 [Herb. Inst. Miguel Lillo 3ih21) (KN); Peirano Sen. [Marifio, Octs 1S, 19323 Herb. Inst. Miguel Lillo 32697 (N)5 154 PAY 0.14.06: 1.4 Vol. 9, no. 3 Schreiter 73 (Herb. Inst. Miguel Lillo 32767] (N), s.n. [San Ped- o de Colalao, Dec. 1917; Herb. Osten 12187] (Ug), son. [Tucumn, acts iT, 19175 Herb. Osten 12168] (Ug); C. Skottsberg s.n. [Es- tacion Agricola Experimental, 16/10/18] (Go); Terribile e lbh (N), 287 (N); wee 78 (W—1591211), 462 [Herb. Inst. Miguel 1 Lillo 3128] (N, W—1591220), 2300 (Ca--32547, S, W--1343273)5 Wall & Sparre s.n. [12/11/16] (Ew), sn. [24/11/46] (Ew). Province un- determined: Jameson 125/26 (W—-15))516) ; Tamson s.n. (v—8997) « LOCALITY OF COLLECTION UNDETERMINED: Née 11h lay (Q yore VERBENA GRACILIS Desf., Cat. Pl. Hort. Reg. Par., ed. 3, 393. 1829 [not V. gracilis Cham., 1832]. Synonymy: Verbena rbena remota Benth. , Pl. Hartw. 21. 1839. Verbena parviflora Rufz ex Walp. -, Repert. Bot. Syst. lh: 25, in syn. 1845 [not V. parviflora Larraf., 1923]. Verbena arizonica A. Gray, Proc, Am, Acad. Sci. 19: 95. 1883 [not V. arizonica Briq.e, 1907]. Verbena parviflora Pavon ex Moldenke, Résumé Suppl. 6: U, in syn. 1963. Bibliography: Desf., Cat. Pl. Hort. Reg. Par., ed. 3, 393 1 aca 1845; Schau. in A. DC., Prodr. 11: 59 Proc. Am. Acad. Sci, 18: "hs. “1083, Ae sig! 8 Proc. Am. Acad. Sci. 19: 95. 1883; Hook. f. & Jacks., Ind, Kew. 2: 1178 & 1179. 18955 i. Armstrong, Field Book West. iad Fls. 36. 1915; Stapf, Ind. Lond. 6: 429. 1931; Perry, . Bot. Gard. 20: 245, aky 261, 297, 300-— 301 £3555 19335 1 pilderice: Prelim. Alph. List Invalid Names Lh, {8, & 56. 1940; Moldenke, Knowm Geogr. Distrib. er 08, 73 933, & 963 (199), and h: 996, 1076, 1128, 1171, 117h, 1175) 1180, 122k, 1229, 1233, 1245, 12h7, & 1291. 1949; Moldenke, Kn Geogr. Distrib. Verbenac., (ed. 2], 19, 26, 33, 16h, & 198- 19193 SC. ol. 8: 105. : h9 Spec. 3. 195k; Moldenke, Résumé 2), 32, ? 223 ha? 365, aire & nt 1959; Moldenke, Résumé Suppl.” 2: in 12 226 9 rs a sely branched ammual or perennial herb, varying from upright to pores or pane waren oy stems 26--L0 cm. long oF 1963 foldenke, Monograph of Verbena 155 tall, widely branching, round-angled, whitish-hirsute, more or less. glandular; branches decumbent to ascending, more or less glandular, canescent, hirsute; leaves mccain Arte Ae abun- dant along the stems, » ovate or obovate, 1--8 cm. long, cuneate at the base and narrowed int a marginec a a era fe) to lavender-blue, lilac, pale-mauve, Rs = , its tube Somewhat longer ues the calyx, the limb 2--3 mm. wide; cocci 1.5-—-2 mm. long, finely scrobiculate from the apex practically ’ = the base, the commissural face muricately scabrous. This specie a Cultivated in 1829 in the Jardin des Plantes at Paris and prob- ably deposited in the herbarium of the Muséum National d'Histoire Naturelle in that se The type of V. remota is Hartweg 17) from Aguas Calientes [or Zacateca cording to the Kew speci- men], Mexico, aches (187) toi youl V. remota and V. grac- ilis to synonymy under V. canescens H.B.K. “Hooker & Jackson (1895) reduce V. arizonica A. Gray to V. remota, and then reduce the latter and V V. gracilis to V. canescens. In my Prelim. Alph. List Invalid Names 56 (190), Alph. List Invalid Names \7 Eve and Résumé 365 (1959) I also erroneously reduced V. gracilis to Synonymy under V. canescens. It should be noted here that V. gracilis of Chamisso is a syn- Onym of V. intermedia Gill. & Hook., the V. arizonica of Briquet is a synonym of V. gooddingii var. ne epetifolia Tidestr., and the - os of Larraflaga belongs in the synonymy of V. litoral- Secondary tesa Sree zicoe hg other species. The species inhabi S$ sandy or dry red gravelly soil, sandy lo id il of oven woodla a has been Collected in waste places and grasslands, s anc dry steep- + hills, roadside bankings and moist depressions, roadsides in s grassy 2 fron varch to September. Iueller says that it is rare in 156 PAY TOL06 128 Vol. 9, no. 3 meadows in Nuevo Leén, while Benson reports it from swales in desert grasslands in the Lower Sonoran life-zone. It was cues ly Aer into cultivation in or about 1829. erbarium material of this species has been misidentified and Frere under the names V. bracteata Lag. & Rodr., V. bracte- osa Michx., V. bracteosa var. . bre See Gray, V. canescens Kunth, vy. hastata L., V. menthifolia Benth, » V. multifida Pav., ana Small, V. plicata ae ae even Myosotis sp. in am boraginaceae. On the other hand, the Waterfall & Wallis 13806 distributed as V. gracilis is partly V. canescens H.B.K. and partly V. ciliata hire Conzatti 288 is is V. came camescens; J. R. West 13216 is V. ciliata Parry & Palmer 720 | is V. elegans var. asperata Perry; while Scoditine ding 33h, Herb. Le Roy sen. fAriz. '8h), M. E. Jones s.n, [Colonia Jusreal, Cc. B. Maguire 1038h, and Pringle | S.n. [Santa Rita Mts., May 11, 188] are all V. neomexi- cana var. ". xylopoda Perry. The ¥acDougal 63h ars aa y regarded as as V. gracilis proves actually to be V. neomexicana . hirtella Perry. The J. J. Thornber s.n. ocean Sect . 2; 1905], cited Santa Cruz County, | may actually have been collected in Pima County, a ee pines. Gregory & Eiten encountered it "in we san boulders to river in barranca", NeVaugh reports i ‘ "periodically flooded flats and sandy es entering the flats", while Maysilles found it on "gentle west-facing slopes of broa arroyo, oes Peete tte * Reesor woodland" and describes it as a Piva n open meadows in pine forests." rry (1933) cites the folle owing 36 additional specimens not Parts seen by me: ARIZONA: Cochise Co.: Blumer 1612 (E, F, &)3 Lalla Sen. [Tanner's Cafion, near Fort Huachuca , 1082] (G). Pima Co.: Pi Pringle Son. [Gerdinerts Spring, June 25, 1882] (D, F, G)- MEXICO: Agua Aguas Cal: Calientes: Hartweg 17h (G, K). Chihuahua: Pringle 5h (G), Sen. [near Chihuahua, 7 May ‘ay 1887] (E, F). Durango: Nelson 1593 (E); Edw. Feiner aa '(E, ¥, G), L56 (G). Federal District: Pringle 6539 (D, F, G), 9135 (EB, G). Hidalgo: Pringle 7590 (E, F), 13159 (F, a). Wes xico: Eerlandier 578 (E); Bourgeau 361 1 (G)5 Rose, Painter, & Rose 895 (G). Oaxaca: Seler 22 (G). San | Luis Potosf; Parry & - Palmer 722 (E, G); Schaffner 720 (D, E, G)- She says "Ve grac gracilis is probably a relative of V. canescens and V.- neomexicana. It is easily recognized by its slender habit, usu- ally long-attenuate bracts, tiny flowers, and essentially scrob- iculate nutlets. The specimen from Utah is a ety nemt; but for ems better referred here." n all, 153 her a Specimens have been examined by me. u dberg Citations: UTAH: s an Co.: Ry re & Garrett 9201 (N)- 1963 foldenke, Monograph of Verbena 157 ARIZONA: Cochise Co.: Blumer 2) (Tu), 1612 [N.2h] (Du--95)6, Ka-- 60316, N, Tu, W--561809, W--56338); Goodding 292 (Tu); Mearns 1116 (W--22906); Pataky s.n. [Fort Huachuca, 1890] (W--771871); We W W. Price sen. (Aug. 2h, 1893] (Du--9631); Pringle s.n. [Camp Lowell, June 8, 1882] (Du--91156) 3; Shreve 773) (Fs, Mi); Water- fall 1291) (St); T. E. Wilcox 29 (W——213239), sen. [Ft. Iuachuca, July Te! (Cc), Stn . (Near Fort Huachuca, 189] (W--213031). Pima Co.: Pringle s.n. . [Gardiner's Spring, June 25, 1882] (Cc, Ll, Pa, 565513, 0 Up--17096, Vt, W); J. J. Thornber 5637 (Tu). Santa Cruz Co.: L. Benson 1026 (po--26 7651, Tu-—1082); J. J. Thornber nte sn. [Rosemont, Sept. 2, 1905] (Tu). MEXICO: Aguas Caliel Hartweg 174 (i). Chihuahua: Correll & Johnston 21530 (1d); Le Sueur 8 872 2 (Au); Mexia 2568 Poca tes Gg——286757, Mi); Edw. Palmer 20 200 (W--573677); Pringle 5k (Vt), sen. [2h Julio 1385] (ue), san. [7 May 1887] (Ca--10h858), s.n. [22 Mayo 1887] (Me)5 Shreve 8008 (Fs), 80h) (Fs, W--179160); S S.S. White 2468 (Mi). Coahuila: Stanford, Retherford, & Northcraft 126 (Ca—71389h, Du-~291287, N, Se--70L52) « Durango: J. He Maysilles 7497 (Mi, N), 8097 (1k); E. W. Nelson 593 (W--332609); Edw. sor 135 (Ca—10h859, Me, Me, N, W--30L237), 456 (W~571L01), 911 (W— 304612) ; Waterfall 1sh02 (st), 16163 (Ca), 16178 (Ca); 5 Waterfall & Wallis 1302 (St), 13417 (Sst), 13483 (st), > 13528 (st), 13622 (Ok). Federal District: Balls 952 (W—1793739), B B.952 (Ca 684616) ; Bourgeau 361 (Br), 547 (Br, S); G. L. Fisher sher 35208 (Ew, Mi, N, W--1686256), s.n. [Villa de Chabad 28 jaly 18, 1935] (Fs); Be Lyonnet 1233 (W—1746020), 1562 (W--17L6717); \iranda, Barkley, &) Rowell 7 unde (Au); ceinete | Zt (Br, Ca--104857, Cm, Gg--152313, To--38759, He ,» Me m--15393, N, S, vee » w=287765), 9135 (N);° con 2 “e 21,88" (12.39¢21893 Rose, Painter, & Rose 8195 (W—1,51987) ; A. J» Sharp 228 (N). Se ei "née 28 , 28 (Q)- Guerrero: Hinton 5847 (N, W—-134h912) . Hidalgo: Moore & -& Wood W131 (Ba); Pringle 7590 (B, Po--63968), 13159 (Ge-—U21272, Mi, Ti 35528, Vt, W--l61756), s-n- [El Salto, 15 Sept. 1902] (Vt); Salazar s.n, [Huauchinango] (Me), s.n. [Nopala, Aug. l, 1913] aims Jalisco: Gregory & Fiten n 158 (Mi); McVaugh 16880 ve i). uSxico: Berlandier 675 [27h] (Du--166499, W--1169371)5 ~ourgeau 361 (Cb); Matuda 26: 26165 (Cb), 26302 (2); Je Ve. Santos 2h (i); s Schiefer 39 (Vi). Nuevo Leén: M. M. Te Edwards Lg (De (Du-- 277046); Frye & Frye 2392 (W—1790998); C. He '. H. Mueller 2392 (Mi). Querétaro: Kuntze 26lLh ( (il, N); Waterfall & Wallis 13996 (St). San Luis Potosi: Barkley, Rowell, & Paxson | 826 26 (Au--123256, N)5 stiley, Webster, & Paxson 768 (Au—-123279, N), te oe M5. L. Lundell 5047 (La); Parry & Palmer 722 = ig a Selsey. By mdels BOUT (ia); Fares aes: 11/G7] (T 5: 158 BEY? O1:04-L4 Vol. 9, no. 3 White 947 (Mi). Vera Crug: A. J. Sharp 45487 (N). Zacatecas: Shreve 7199 (Ca—-731765, Fs); Weintraub & Roller 20 (Mi). State undetermined: Halsted s.n. [La Olla] (T); Née 10h (Q); Schmitz 4,78 (Bm); W. Schumann 1070 [Faral] (W--132305)). CULTIVATED: France: Me Martens s. Sn. 1. [he bs. Par. 1835] (Br, Br), sen. [hort. insul. 1836] (Br). LOCALITY oF COLLECTION UNDETERMINED: Herb. Jan. Gen, 22 Spec. 3 (S) VERBENA GRISEA Robinson & Greenm., Am. Journ. Sci. 150 [ser. 3, 1895. 50): Bibliography: Robinson & Greenm., Am. Journ. Sci. 150 [ser. 3, 50]: 142—1)3. 1895; nee Jacks., Ind, Kew. Suppl. 1: 451. 1906; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 34 & 101. 1942; Moldenke, Castanea 13: ae 193; “\Woldenke, Alph. List Cit. 3: 972. 199; toldenke Known Geogr. Distrib. Verbenac., [ed. 2], 71 & 198. 19h9; foldenke, savas 81 & 471. 1959. ensely and somewhat sericeo ously erayish-tomentose, in parts brownish or ferruginous in the dried state 3 stems te tragonal, somewhat furrowed; internodes mostly considerably exceeding the e subtending leaves; leaves decussate-opposite, petiolate; leaf- blades ovate or deltoid in outline, 2.5 cm. or mo fe) densely pubescent as to obscure completely the venation on both surfaces, bininnatifid, the rachis and segments very n ot mm. wide, frequently curled, obtusish at the apex; intioye. cence branched, the flowers small, borne in slender loose elon gating spikes, we acti das spike floriferous agai t from the pase, the others more S pedunculate; bractlets small, subulate, 1/3 to 2/3 the tenet. of the calyx, pubesce oe galye cylindric, about 2mm. long, 5-ribb bbed, hirsute-pubescent on the outer sur- mpi the lee! very s hal lowly and obtusely 5-toothed; corolla arly twi s long as the calyx bescent in the throat, es Linb unequally SLobed, the lobes rounded at the apex; anther: sor ie ndaged. he type of this very disti cette species was collected 0 poseke irae pean os Islands, in August, 1891. The authors say that it i member of the peaks here with much the habit of V. remota Se but differing in pubescence and foliage." The s Species is said around 1250 feet elevation 0 - It has been collected in anthesis in Augus - in August and December. In all, a herbariun gies ie and mounted photocraphs have been examine Citations: GALAPAGOS ISLANDS: Duncan + Stewart 3315 (Ge—- aeodb XN); 3316 (F~—photo, Gg--31277, Se Si--photo, Z-- VERBENA GYNOBASIS Wedd., ae And. 2: 156, a ee bella R. A. Phil., Anal. et Nac. Chil. 1891: 58. eaten ae Wedd., Chlor. And. 2: 156. 1860; R. A. Phil., nal. tus. Nac. Chil. 1891: 58. 1891; Hook. f. & ‘Jacks, Ind. Kew 1963 Moldenke, Monograph of Verbena 159 2: 1179. 1895; Durand & Jacks., Ind. Kew. Suppl. 1: 451. 1906; Reiche, Fl. Chile 5: 235 & 286, 1910; Moldenke, Suppl. List In- valid fae es 7. 191; Herrera, Sinop. Fl. Cuzco 1s" 382, 1941; Mol- enke, Known Geogr. Distrib. ‘Verbenae +, [ed. 1], h2 & 101. 19h2; ictasscn, Alph. List Invalid Names 5. 1942; Moldenke, Lilloa 8: 430 (19,2) and 10: 38). 191; Moldenke, Knowm Geogr. Distrib Vargas, Bol. lius. Nac. Hist. Nat. Chile 25: Sh. 1951; Moldenke, sumé 115, 121, 358, & 71. 19595 Ricardi, Gayana 3: 13. 196; Noldenke , Résumé § Suppl. 3: ly. 1962. Herbaceous perennial or slighty suffruticose, canescent- hirtous throughout, 10--20 cm. tall; stems several from a single ’ margins 5 spikes terminal, short-pedunculate, many-flowered, the florifer- ous portion short, ovate after anthesis, in fruit not over 1.5 Bebe . bractlets linear, nals as sy regis as or slightly shorter alyx than t 2--) mm. long; calyx linear or prismatic, 4—- os tae e rim 5-fid, the tek titeiiee e- or ie haped; cor= olla dark-red, its tube subi dibular, about half again o twice the le of the calyx, the limb rose-purple or deep-red, ginate at the apex; stamens included in the corolla- See mee ke ves unappendaged; one of the sey onbeanabel horn 5 deeply Was collected by Rudo = Apans Philippi at Sibaya in the Cor- dilleras de Tacna, Tarapa hile Herrera (1941)’ perce a NGay Bice wor "Cuzco: Andes del enton i which is probably the type collection of var. strigosa Wedd. (see below). The a re has been gathered at 2400 meters altitude in Bolivia, flowering in April. The o- i s ss LA © 3 fg "1857--61", but seems actually to pote been gare ue eg ee Pts Calyx is described as l-S mm. long n V. gynobasis and long in vy. bella. ete . wibina (1981) cites the 3 following SPecimens and photographs: CHILE: Tarapac4: C. Rahmer S-ne [15. ure) (Sg--5h671), sn. are .1885] (Sg—l2u58); R- Ae Phil- ippi Son. ees Macbride photos 17419] (Sg--phote) - 160 PHYTOLOGIA Vol. 9, no. 3 In all, ie eran specimens anc 2 mounted photographs have been e Citations: BOLIVIA: Santa Cruz: Troll 3312 (B). Province unde- termined: Lobb s.n. ie Bay Herb. Reichenbach f. 158619] (N, V)- CHILE: Tarapac apacd: R. - Philippi s.n. [Sibaya; Macbride photos 17419] (Kr--photo, | fat oto). VERBENA GYNOBASIS var. ‘Sept OSA Wedd., Chlor. And. 2: 156 [as "6 strigosa"]. 1860 in ing the hair on the stems, branches, and teayes nine strigose, rather than hirtous, and the corollas sma The type of the variety was Aptis ted collected by Claude Gay in the Andes of Cuzco » Peru, in 1339, ifw we can assume that the that the first represents the typical form of the species. It is known to me only from the ori iginal description. ee HALEI Small, Bull. Torrey Bot. Club 25: 617. 1898. ynonymy : Verbena leucanthemifolia Greene, Pittonia 5: 135. rc Verbena hallei Small ex Moldenke, eo List Invalid Names 8, in syn. 191. Verbena hallii Small ex Moldenke, Suppl. List rete iebe 8, in syn. 19h1. Verbena halei L. ex Moldenke, Alph, List Invalid Names Suppl. 1: 2h, in syn. 197. Verbena coerulea Vatke ex Moldenke, Résumé 362, in syn. 1959 [not V. coe- —_ rulea Hort., 1959], Verbena haleii Small ex Moldenke résumé 365, 19 Bibliography: : : Foubee Bot. Club 25: 617. 1898; Britton & Br., Illustr, Fl, ed. 1, 3: 70, fig. 3057. 18985 Greene, Pittonia 5: 135. 1903; Small, Fl. South. U. S., ed- 1, _ 1008 £1009. 1903; Thiselt Dyer, Ind. Kew. Suppl. 2: 191. 1903 Prain, Ind. Kew. Suppl. 3: 187. 1908; small, Fl. Southeast. U. Ss ed. 2, 100 1009, 1913; Britton & Bie, Ids Fl ° ‘ 95, fig. 3552. 1913; E. D. Schulz, $00 Wild Fle. 17822179. 19223 ei, , Texas Wild Fls. 337—336. 1928; small, 2 see Bull. 550: 88. 1937; ee Texas Wild Fl. Legends, ed. 2, 113. 19h0; Moldenke, Prelim. Alph. List Invalid Names 47. 1903 ¥ol- denke, Suppl. List I tivelca Names 8. 19)1- Sperry, Sul Ross Bull. hl; 22: - 191; Moldenke, Known Geogr, Distrib. Verbenac., [ed- 1), 1963 Moldenke, Monograph of Verbena 161 hm-6, 10, 11, 13, 1h, 18, & 101. 1942; Moldenke, Alph. List In- valid Names hi7 & one "19425 Moldenke in Lundell, 5 8 Texas 3 (1): 16 & we hes Schnack, Anal. Inst. Fitotéc. Sta. Catalina h: 18, ue ag Davis, Study Boscaje Palma 72. et Saree & Covas, ar a 6: 4,70. 19h; Darlington & Janak o- mosome Atlas "270. 1945; Murrill, Guide Fla. a 18. ToL; jee ke, Bot. Gaz. 106: 160 & 161. 19453 Moldenke, Journ. Bot 609. 1945; Moldenke, Phytologia 2: 68, 8h, 87, 115, & 128 (rhs). : 19h6; and 62-- ; Ge L. Fisher Am. Bot. Exc 1946; Tolstead & Cory, Field & Lab. 1h: 60. 19l6; Moldenke, Alph. List Glue: Le \ a ree. h ’ Sen? Vie ’ > 115, 119, 125--127, 130, 135, 154, 159, 195--197, 199, 202, 203, 209, 216, 226, 267, 270, 276, 280, 281, 291, denke, Phytologia 2: 327—329, 1947; He N. & A. L. Moldenke, Pl. Life 2: 62. 1948; Moldenke, Castanea 13: 112 & 113. 1918; Molden- ke, Wrightia 1: 222--22). 198; Moldenke a. List Cit. 2: 391, a P91» 298, UOT, 126, 138, leims3, eh Woy UML, Uth-U80, sta & 1299. 19h9; ze 1230, 12 Knovm Geogr. Distrib. Verbenac., [ed. 2], >, 10, 11, 18, 19, 21, 23, 25, 26, 33, & 198. 19h9; Moldenke, - Ne e, . Inst. Biol. Mex. 20: 13. 1919; Moldenke, Phytologia 3: 72 & 132 (19h9) | awa 3: 67. 1951; McDougall & Sperry, Plants Big Bend 14,6. hytologia 4: 166. 1953; Moldenke, Inform. Mold. Set is Sc (hy (195) and 9 Spec. 3. 1954; Moldenke, Phytolo- Rec. : 132 & 133. 1955; Moldenke, Biol. Abstr. 30: 1093. te gia 6: 330. 1953; oldenke, Am. Widl. Nat. 59: 342, 3h, & 361- ee Eos 3 ee fs ’yalias Rig 1958; Moldenke, Résumé * Oliver, eget Bot. 8: ae: 1961. Slender, nearly smooth, annual or cena herb, 20--120 cm. fibrous; stems usually several from a woody base, ere Pes 55 Cm. tall, ascendingly branched, glabrous or strigillose above, 162 FT T 01 86 Fs Vol. 9, no. 3 leafy below; bark green; branches erect, about 15 cm. long; leaves decussate-opposite, 3--10 cm. long, strigillose on both surfaces, ive in outlin vi to inating in aes as long as broad; Site trigonous, approximately 2 5 Mim. —— usually prominently s triate, raised-reticulate at the ee he commissural faces muricate; chromosome number: n = 7, 2n “The type of this species was a aa tgs by oar pei ee eo To ugh th cites, in addition, Croft 119, Heller 119, and Palmer 242, wi no specific one “cesignated. The species is named in honor of tke als Demaree— 32070 (Le), 33307 (Cb, cb, 2), 33708 (Ss); S. M. Tracy s.n. [Bil- oxi, oxi, 6/9/00}(N) . Jackson Co.: Demaree 33625 (Cb, Ss), Ss), 34996 (Ss); He I. Jacob 1296 (Hi--19629)); J. Skehan 109a (Au-—-1803h2), 1599 QW), s.n. [Seymour & Earle 109] (Gg--26N185, Hi--59L63, Ur), s- n. [Seymour & Earle 109a] (Id), sn. (Ur—22596). Jefferson Co.: UeDougali 1291 (W--1925706). Stone Co.: Demaree 32532 (Le). Ship Island: Demaree 30585 (Hi--51390, Mi, N, Ok, St, We), 30710 (N). ARKANSAS: Calhoun Co.: Demaree 22711 (N, Sm). Drew Co.: : Demaree 24933 (Sm). Hempstead Co.: Demaree 1270 (Gg). Miller Co.: Demar- £e ee 39156 (Sm, Sm), 0819 (Bm). Stone Co.: Demaree 3187) (Bm). U- nion Cov: D. M. Moore 510154 (St). Washington Co.: Brackett 253 [April 21, 1938] (Au). LOUISIANA: Acadia Par.: Penfound s.ne (A Pril 21, 190} (Tl); F. W. Pennell 10201 (N). Calcasieu Par.: A. R. Woldenke 228 (Fg); Small & Wherry 11774 (N), Some [near Lake Charles, Ap April 11, 1925] (W--173911)). Cameron Pare? alice 2 1953 (W—61,91,3) ; S.M. Tracy 5709 (T1), 8709 (Es, N, Tr, Up-—503L5, eee Iberia Par.: Ashbey 129 le Iberville Par.: Je He aoe sete Par.: prenning sen. [17 190k] Se se a 168 Pat tT OU SOTA Vol. 9, no. 3 Sen. [New Orleans] (T). Plaquemines Par.: Langlois s.n. [28 Mai 1883] (Du-~90896). Rapides Par.: Ewan 19377 (Tl). Saint James Par.: Small & Alexander Sen. [near Convent, Apr. 1931] (Au, H-- 82843, N, Ny N, We). Saint Martin Par.: Langlois s.n. [April 1880] (I). Saint Tammany Par.: Ars&ne 11831 (W--10330%), 122h2 (W--1033052); Cocks s.n, (Slidell, Apr.] (T1)..Tangipahoa Par.: Correll & Correll 925), (H—78937, N). Union Par.: R. Kral 8673 (Hi--151002). Vernon Par.: Cooley & Brass 3762 (Tl). Parish un- determined: Hale s.n, (T—type); Riddell 1268 (N). OKLAHOMA: Ato- ka Co.: Hopkins, Nelson, & Nelson 1092 (0k). Bryan Co.: Ce Se Sheldon 23 (Ob—97289, W). Gaddo Co.: Edw. Palmer 2h2 (Ww). Carter Co.: A. G. Payne 38 (Ok). Choctaw Co.: Waterfall 2102 (N, Ok). Jefferson Co.: Waterfall 7829 (Au--122275, Ok, St)+ Johnston Co! Edw. Palmer 22 [339] (T); G. T. Robbins 2410 (Ca-—966184, Ok) - Latimer Co.: Hopkins & Cross 1732 (Ba, Ok). Leflore Co.: R. Bebb 5398 (Ok, St—-19588). Love Go.: E. E. Dale 82h (Ok); Nelson, Nel- Son, Goodman, & Waterfall 5706 (Ok); Waterfall 725) (Ok). Marsh- all Co.: Alder s.n. [July 10, 1950] (Ok); Cory 58996 (St); Ve» Ee Dowell sen. [April 22, 1955] (Ok); G. J. Goodman 6458 (Bm, Bs, Mm, Ok, Sm); Kelting 193 (Ok); P. B. Riggs s.n. [Lake Texoma, June 20, 1952] (S); Starks s.n. [19 July 1950] (Ok). McCurtain Co.: Waterfall 8057 (0k), 8825 (Ok, St), 10470 (St), 11829 (Au— 122277, Ok, St, W--2231103). Murray Co.: Stratton 62 (St--26122). Muskogee Co.: C. S. Wallis 2219 (St). Pontotoc Cos: G bins 2486 (Ca--966 SN. [Kiamichi, June 3, 1937] (Ms). County undetermined: Cc. S. 531 (Hi—111003, xi, W—2327763); E. G. Marsh 52 (Au--122130), 27-61 (Au--122235); E. D. Schulz S.n. [Palestine, 1930] (Wi); K: E. Smith s.n. (Palestine, 4/21/35] (Au, Bt—25),28, N, st——-925h)5 Tharp sen. [Palestine, 4/21/35] (Du-~362690). Angelina Co.: Parks & Sory 10651 (Tr). Aransas Co.: Cory 1903 (Sm); H. C. Johnston 53253,8 (Au—12216)), 53253.12 (Au=1DO053). arches Cac? Henon 8 n. [Archer City, april 10, 1934] (Au, Au); Whitehouse 9721 (Mi, N). Atascosa Co.: Parks & Cory 1917h (Tr). Rustin Coc: He Wurzlow S.n. [Industry, 1891] (ur). Bastrop Co.: Gc. C. Plant Project e- es ve Ve s * ceil 103uk (Id, N); Be H. Warnock 46283 (au), Gob (Aas By Oks St); Westlund, Rowell, & Barkley 177216 (Au, ii). Dee Co.t Fe Ae Barkley 16T)21 (Au, N, Or—-55L08, S, Vi); Cory 54133 (Rf); Lun- dell & Lundell 10051 (N). Bell co, : Cohn T.11 (Au, N); Cohn & Barkley Tels (Au); S. E. Wolff 2918 (Tr, Tr—1616h, W—162L016), 3491 (Tr, Tr—23397), 3718 (Tr-=19170); York # York 53053 (0k)> Bexar Co.: C. R. Ball 927a (W—50151,7); R. Bebb 2202 (Ok); Burr 1963 Moldenke, Monograph of Verbena 169 U9 (N); L. D. Christensen 38 (Ar--250852); Clemens & Clemens 502 (Po—69601), > 969 (Po--69887), 970 (Po--69572); Groth 75 (Ge— 31399, N, W--717689); Havard s.n. [San Antonio] (W— 15563, W— 218876) ; Headley s.n. [March 2h, ag (I); G. Jermy 87, in part (W--315686), 83 88 (W—315687); L. G. McLean 8 (St); Meadows s.n. (April 20, 19l1] (au); Metz 62 at 75 (I), 3240 (Av, N); Me 0. Rice s.n. [San Antonio, April “as. 1938] (St); E. D. Schulz 103 (W--1102680), 518 (N, W—1O7L9L5, Wi, Wi), 766 ( (N, W--12h3197, i), sen. {San Antonio, h-1)-1922] (his 5 Son. n. (Spr. 1926] (Mi), san, {San Antonio, 1-32] (Mi), son . (9 mi mi. so. San Antonio] (Wi); Cc. Skottsberg s.n. [San Antonio, 2/6/1935] (Go); Texas Agr. Exp. Sta. Sta. Herb. Exchange 3495 (Hp); Whelan 171 (N). Bowie Co.: Correll & Correll 1221 (Ca—90725h, Sm Sm) 3 Eegert Sone {13 June 1898] (N); Milligan s.n. [Dalby, May 1891] (—50335) « panes Co.: B. F. Bush 77 (Hi--59L61, N, S, W--336410); =. J. P 50k (Po--217659); Tharp 50-106 (Au--122217); He S. Young s.ne is 4/22/18] (Po--161329) . Brazos Co.: Buechner s.n. [3-19- 46) (au) ; Chenault s.n. (April 12, 1937] (N)3 Cory sory 50627 (LA); FP. W. Gould 253 (Au-—-12223h), 51,36 (Ca—-917763, S, Ur); Ur); 66bh (Ca-—373888, st St); C. E. Gray 23 3 (Ei); H. S. Jennings s.n. [Col- lege, Mar. 31, '70] (Mi); Lake sen. [May 1890] (Pr); Ness s.n- (Apr. 15, 1926] (Tr); E. J. Palmer 11715 (Ca--l2559l); Le He 1 s.n. [College Station, 7-1388] (Io--1539); H. B. Parks Sn. [)-13-),7] ees Sone [h-30-117] (Au); E. L. Reed sen. , [Col- lege Station, Oct. 1915] (N); Reeves 189 (N), 193 (N), 196 (Cs), 208 (Cs); J. N. Weaver 490 (Ld, Mi). Brewster Coe: Hinckley & Warnock 1096 4096 (N); 0. B. Sperry "983 (W--1699323); B. H. ¥ Warnock ase Gn, s W.538 (Au, N)5 Warnock & Hinckley BG.108 (Rf). ). Brooks - L. Lundell 10811 (Rf, Sm); S S. Williams 96 (Ok). Brown COes Boke L. Sweet s.n. [Brovmwood, 5-28-1931] (Au, Au). Burleson Co.: Herb. Univ. Texas s.n. [Lyons, Spring 1928] (Au), s.n- Hoecreatid ae Spr. 1930] (Au). Burnet Co.: Rose-Innes & Warnock 198 (Au). cathoun Co.: H. S. Gentry 29 (Au, N, S)3 Tharp s.n. SMe [Port O'Connor, 5/19/30] (Ca-—3326))7) « Caldwell Co.: McBryde | Se ne (Spr.-Sum. '31] (Au). Calhoun Co.: Trew 211 (Au--122176) « Callahan Co.: a son. [Putnam, Apr. 1926] ~ (Wi). Cameron Co,.: 4. B. Cannon s.n, = Rosa, Nov. 21, 1926] (Mi); HM. Cannon Sls . [Santa = Feb. 1927] (ii, ui); H. P. Chandler r 7082 (Ca--17,503, 1 eG. Clover 736 (Iii); L. I. Davis Sn. [Palm Grove, Sanne 11] (Au, Sm, St); Ewan 15551 (Tl); i. Ge Johnston 253-8 253-8 (Au--1.22139); C. L. Lundell 10639" (N, Rf); Lun- dell & Tundell 8623 (Ld, Mi, ™); “Yealley 115/370 (W); R- Runyon 1777, in part (Rr, Rr), 2181 (Rr, Rr), 2368 (N, N), Sn. [Browns 170 PUY.T,0L 0G.1.A Vol. 9, no. 3 ville, 1930] (Hp); Tharp 1872 (Au); Tharp, Gimbrede, & Johnston 52-523 (Au--122190). Cass Co.: McClung s.n. [6-15-26] (Au). Cham- bers Co.: L. Barrow 31 (Au, Au); Tharp sen. [h/7--10/36] (Au, Mi). Cherokee Co.: F. A. Barkley 13585 (Au, N); High School Sone (Au) Coleman Co.: B. H. Warnock 16341 (Au, N). Collin Co.: Lundell & Lundell 9318 (Du--321633, N). Colorado Co.: Biology Class 19 (Au); B. HE. Warnock 6430 (Au, N), 46432 (Au); Wild Flower Contest sn. [Eagle Lake, Oct. 1930] (Au). Comal Co.: Landa 11081 (Tr); Lind- heimer 537 (Ka), 1076 (Au, Br, Ca--1)7515, Ca--176385, Me, Me, lie, N, Ok--10360, Up--8537, W--502)9). Cooke Co.: E. E. Russell seme [Gainesville, 5-23-1923] (Au). Culberson Co.: Cory 53037 (N)3 Waterfall 496 (N, Ok, Ok). Dallas Co.: Boll s.n. [1871] (Cb); Cory 53699 (N); Hamric 21 (Au—-168941); Hynes sn. [Dallas, May 3 '25] (Au); M. E. Jones s.n. [Dallas] (Po--70677); Lundell & Lun- dell 83hh (Mi), 8316 (Ld, Mi), 8578 (Ld, Mi, N), 10393 (Id, N, W 1887663), 12107 (N, Rf); J. Reverchon 732 (W); Shinners 8505 (Sm); M. B. Stephenson 200 (W--158053)) ; R. Van Vleet 86 (Au); Walden 13 (Nt); Wilfong 82 (sm). Denton Co.: Mecart 1977 (Au--122231), 2006 (Au, Mi, St--1l716); Whitehouse 15735 (Sm). Del¥itt Co.: Blanch 17 (Ok); A. H. Howell 313 (W--501h62); M. Riedel s.n. [9- 26-H1] (Au); E. D. Schulz 2502 (W—1521151, wi). Dimmit Co.: Al- varez, Guajardo, Salazar, & McCart 7619 (Ok); D. S. Correll 20735 (Rf); Correll & Johnston 19473 (Rf). Duval Co.: Alvarez, Guajardo, Salazar, & MeCart 7671 (Ok); Croft 119 (C, Ga), SeMe {San Diego, 1885] (Mi); R. Garza 7 (0k); H. J. Hamby 77 (Ar— 206292); M. C. Johnston 54107 (Au--122237, St). Eastland Co.! Herb. Univ. Texas s.n. [Sept. 11, 1931] (Au); Mrs. Robinson §-Me (Ranger, 9-11-31] (Au, Au). Ellis Co.: 0. sanders 19 (Ii). Brath Co.z Hoisington 27 (Au—122891), 8h (Ok), 2h1 (Ok). Falls Co? A. H. Howell 362 (W—501510). Fayette Co.: Parkley, Warnock, & Tharp 46350 (Au, N); Parks & Cory 10072 (Tr--16171); Ripple 51+ (Au--1222)8); Lundell & Lundell 10139 (Ld, N); Parks & Cory 12748 (Tr); Wolcott & Barkley 16T265 (Au, N). Galveston Co: Re Bebb 1131 (Ok), 1481 (Ok); Cory 51017 (N, sm); R. L. Crockett S+ n. (Apr. 22, 194] (Au); G. L. Fisher 189 (W--107623), s.n- [Galveston, June 6, 1920] (H—SIi69, Ur); C. S. Wallis 8266 (Sts St). Gillespie Co.: Bryant & Bryant 3 (Ca--990129, Ok); G. Jexmy 203 (ii). Goliad Co.: Lundell % Lundell 10060 (Ld, Ld, N). Gon- zales Co.: M. L. Fernandez 58 (Ok); Parks & Cory 598 (Tr), 13259 (Tr); C. G. Ward 288 (St-—-1687h). Grayson Co.: Center 3 (Nt); He S- Gentry 50-198 (Au—122713), 18¢ (au); G. Graham 39 (Au, Au), S.n. [Denison, 5/2/32] (Au); LiacGarland 18 (Au); O¥- 1963 Moldenke, Monograph of Verbena wal ster s.n. [Denison, May 21, 1883] (li). Gregg Co.: F. W. Johnson 1766 (N); C. L. York sn. (vall 1937] (Au), sen. [h-2=38] (Au). Grimes Co.: : Harding 579 (St--20287); J. N. Weaver 1038 (1); T. V. are 1038 (11). iarris Co.: L. Anderson s.n. [Sept.—-Oct. 19 36) (Au); E. Boon 62 (Au, Ca--8826h6, N, W--231132h), 290 (Au, N); B. F. Bush 28 (N, W-386382); G. L. Fisher 20 (i W-~50309)), sone {Houston, _ Apr. 14, 1912] (Vi), sen. (Houston, Apr. 23, 191h] (Hp), S.n. (Houston, Apr. 9, 1931] (Bt—33327, Du--233966, Gg-—-2225h0, St--17432), s.n. [Houston, May 2, 1934] (Ew, lis); aes 3933 (Bl--l,2306); E. Hall 432 (N, Po--70678, Pr, W); Hayden s.n. [Hous- ton, 1917] (Io—-100110); Joor 121 (li); Killip 45697 Coe Seon Kuntze 233 (N); Penfound 1-19 (Au, Au), s.n. [April 10, 1936] (Tl); Rose-Innes & Warnock 673 (Au, N); Snyder s.n. cypress summer 1905] (Sd—I259), sen. [Cypress, 1906] (Ca--185369); Thu- row s.n. [Houston Heights, April 15, 1923] (Hu). Hays Co.: S. We Stanfield a [San Mar —— July 1897] (N). tanas cae dB 872 (a); ts 3061 (au—180171) ; Lundell & & Lundell 10035 (N); Painter & Barkley 1428 (Au, N); R. Rumyon 2553 (N, N). Hill pales H. Rogers 22 (N, Nt). Hood Co.: Blackwell dy (Nt); Schoot Son. ee = etecine (40) Houston Co.: Lewis & Strip Son Co.: A. R. cians 205 (Fg); Gee 2, Barkley 13A11) (au, 32 A N,N). Jasper Co.: Lundell & Lundell 10513 (Id, N); Mrs. A. C. Rowe s.n. [Jas sper, 1905] (Ms). Jefferson Co.: Je Fe Erenckle W070 (x) (K); Kolthoff s.n. (4 Maj 1927] (S, S)3 C. L. Lundell 11250 (Sm). Jim Hogg Co.: Salazar & Santos 32 (Ok, . Ok). Jim Wells Co.: H. J. Hamby 664 (Ar—206303); kK. Ce Johnston 54089 (Au--1222h7); Lewis & | & Oliver 5431 (Nb); Lundell & Lundell 10076 M5 Muenscher & Kuenscher 14391 (N); Wagner & Barkley 167320 — ) (Au, N), 167321 (Au, gs 167322, in part rt (N). “Karnes Co.: at Be Cc. Johnson 822 (Hk, Id). Kaufman Co.: Lundell & Lundell 8506 (ia, * Ki); Fe PF. Fa Tyler SeN's panes of Terrell, “april 25, 190k) (W-- S00803) . Kenedy Co.: Cory 2805 (N); S. F. Glassman 2201 (0k); KM. C. Johnston $3253.5 (Au——1222bh), 54276 (Au--122h5), 5), 541058 (Au=-122155) ; Lundell & Lundell 10728 (N, Rf), 103b4 (Rf), 10847 (RE), 1984 (Ld), 19847 (N); A Re Moldenke 195 (Fe); R- Runyon 2358 (N, N, N); Tharp 5902 (Au); TW Wagner & Barkley 167392 (Au, (a, §), 19393 (au, N). Kerr Co.? f. E. Jones 2829 (Ge—-237867) Be Je Palmer 10037 (Bi, Du—20559h, S , W--15 32020) . caccsaeet wis 0. Degener S0e6 (N). Kleberg Co.: Benke 517 (N); Cory 5 Sn); J ) Johnson 10 (Au, Au); ke Ge Johnston 5325310 10 ee i188), 172 PEL? 0 150-6: 4 Vol. 9, no. 3 5325311 (Au—-1222h6), 53253.16 (Au-—-1222)1), 53253.16a (Au-- 122239), 53253.17 (Au--1222)9), 53253.17a (Au--122233), 53253.18 2 (Au--122289); R. Runyon 4282 (N, Sm); J. F. Sinclair s.n. 1. (Kings ville, sumer 1910] (Au); Tharp & York 51-230 (Au--1221))5). 1a Salle Co.: F. A. Barkley 17T077 (Au, Au--178081, HN); Lundell & Lundell 10113 (Ld, N). Lavaca Co.: Tharp , Rogers, & York 19162 (Au--122177). Leon Co.: Re We Long 231 (ti —205308, N)3 N)} Lundell & Lundell 10391 (Ld, N); E. C. Smith s.n. [Oakwood, 5-4-19l2] (Fe)5 Tharp, ' Turner Turner, & Johnston 5973 (Au=-12217),). Liberty Co.: A. Re Moldenke 220 220 (Fg). Llano Co.: G. L. Fisher s.n. [Bluffton, Nay 31, 1936] (Ms); | M. E. Jones 29187 (Bm, Po--199866) ; Kelting 38 (0k)5 Lee, Barkley, &T Tharp 47232 (Au, N); Lundell & Lundell undell 90L0 (Ld, WA); Parks & Cory 6280 (Tr); M. E. Watts 28 (Au); S. E. } Wolff 3067 (Tr--16165, Tr—22251, W--1621,021), 3825 (Tr--16166) . ). Mata- gorda Co.: Cory 51091 (sm); Tharp 1403 (Au); 1 Walter 51 (Au); Be H. Warnock 16308 (Au, N), 46312 (Au, N). Maverick Co.: M. E. Jones 28295 (Po- (Po—-187970). McLennan Co.: L. D. Smith 388 (St), 847 (Au--122170); C. L. York 46087, in part (Au), 16100, in part (Au). Milam Co.: Lundell & Lundell 1037) (Ld, N); Wolff 3620 (Tr). Montague Co.: McCart 163 (Au, N, Nt). Montgomery Coe Warner 66 (Hu). Morris Co.: Correll, Johnston, & Edwin 22158 (Ia). Nacogdoches Co.: W. H, Lewis “Lewis 5276 (Nb); Tharp & rp & Brown 53 51 (Au--122078). Navarro Co.: — 51533 (Sm); Mitcham ]2 (Au) 5 Purdue 7 (Au). Nueces Go.: A. A. Heller 1h19 (Cc, Ca--104376, kn- 15397, W--213687); Tharp 5603 ao ae me ciit 16); S. H. Tracy 8-0 /14/05] (Tr--16172), s.n. [Corous Christi, 3-31-1905] (r— 20); B. H. Warnock 20999 (Au). Orange ect Ja Tee Aer some Sic i, 1925] (F1--29058); small & Wherry 11801 (N); Wild YWood Club s.n. [Orange, Apr. 2s, 1931] (Au); li. S. Young 167 a "253 ee a Pinto Co.: Culwell & Timmons is 3068 068 (tit); J =s We Gillespie 14 (Du--210195) ; Lindheimer 77 [I TT [10155] (E- Tie 96), 155 ee, Parker Co.: oa . York 7 (au--122129) os Co.: B. H. Warnock 16166 (Au). Polk Co. Girvin sen. [AP- il 12, 190] (Au, 2 an); Tharp sen. [7-23-39] (Au), 5-n- [b-12- 42] (Au); Tharp, Turner, & Johnston 51704 (Au--122126). Reeves Co.: Waterfall - 1388 (N, Ok, Ok, Tu--31202). Refugio a Lundell : oe 10856 (N, Rf); A. R. Moldenke 196 (Fg); E. J- palmer a Gi, Du—205599, S, W--1531877). Robertson Co.: Rowell eli 302 ‘i, N); W. H. Seaman ae: Scot 11-9-18 W—-78 7347) at : Walker 75 Ci 2TS5IZ8); - Ward s.n. ee Sept. ll, am (W--1, 7568); s. E. wolff i a (W--1623883). Runnels Co.: J. Ce : sosigy oe 650 ince Rusk aa C. E. Porter 51-1830 lie - San Augustine Co.: E. J. Pa —20559 S, W—-1531929). pete saris Cow: ae era sm); 1963 Moldenke, Konograph of Verbena 173 Lundell & Lundell 10082 (N); McKelvey 1725 (W--1581388); E. De Schulz 855 (Wi), 2812 ( (Wi, Wi), sen. {Aransas Pass, May 1926] (Wi); Tharp sen. [Sinton prairie, 3/23/31] (Au); thax & Brown 48-203 (Au~-122179, N, St), 48-203a (St); Williges -W (Au-- 175509), 15 (Au--175628) » ae ek Wolcott & ~& Barkley wre (an, N). Shelby Co.: + Correll, Johnston, & Bonin 22223 (Ld). Fe asian ‘992 (1d, N). Tarrant Cost Fo0s Gates 19133 (Ka— 89647); Killian 6953 (Au, W—1653h0); Lundell & Lundell 3522 ote .. N); Ruth 8), (au), be (L1), 108 (Cm, Io--113613, Ka, N, p, Ur, W--501768), s.n. [Fort Worth, June 5, 1909] (Po- ts) Wilezek 36 (Cb). ape Co.: S. M. Tracy 7996 (Au, Cm, Es, N, N-—photo, Vt, W--500 366, Z-~photo) Terrell Co,: Farr Se ne sane May 8, 1911] (Up—52680). Throckmorton Co.: Wild Flower Contest S.n. (rivockmorten, April 1935] (Au). Tom Green Cow: C. Smith 151 (0k). Travis Co.: C. C. Albers 33016 (Au); Ar- mer 8386 (Au, Au, W—1)38998), son. [Austin, }.23.29] (Go); Barkley | & Rowell 7057 (Au, ll); Biltmore Herb. 14787a (S); Bogusch son. son. [larch 26, 1926] (Ob--50791); L. C. Chambers 3033 (W— Visas Cohn & ee 13253 (Au, Ca--882529); Collector undes- nated s.n. (Au); A. M. Ferguson s.n. [l/27/01] (Au); Lundell & ae 10308 (La, MN, i a: Mahler 1267 (St); McWilliams 4051 (Au); E. E. D. Schulz s.n. [Austin, 6-30-1920] (Wi); Tharp 1529 (au), ae (W--1221373), sen. (Austin, 4/23/29] (au, N, N, N, Pl— 151577, Vi » We), s.n. (Austin, 5/2/35] (au, Bt-—25h2h, st—-9253), S.n. ied 5/9/35] (Au, Du—362688, St--9252), sen. [Deep Eddy, 4-26-36] (Au), s.n. (Austin, 7/15/38] (Au, Au); B. H. Warnock L5- 5 (Au), 4610) (Au, | Ok, S, Vi); Werquithin s.n. [Austin, 5/1/15) (10102277); Westler J. Y. T- Ss sine Ligeia, Ay 1930] (Mi)5 Wheeler, Buck, & Barkley 167248 (Au, N); i. S. Young 77 (Au), S- ne [h/5/18) ( (Au) . Tyler Co.: ‘ow: Tharp, Turner, & & Johnston 548)0 (Au~122108) , Uvalde Co.: M. E. Jones sen. Sool of Uvalde, April 26, 1931] (Ca--l79073, Du—230896, Po—-187973); C. L. Lundell 10965 (N, Rf); E SW St = 1356 (B, W--160387), 33604 a Small & a a (N) see Verde Co. Cory S-n- ate on rr 9893 ( (RE). van Zandt 3 Ee son. (Wills Point, 1930] a ; Heel 33 eae 5699 (WJ—1h65515) 5 B. Williams 10 (Nt ). Victoria Gor F L. Lewton 37 (Ar--271653), 763 (Ar—2717k7) 3 Kaxon 3822 (iso ee MeVaugh 1222 (Mi); A. Re Moldenke 203 oy gee: Co.: Parks & Go 8 (Tr); Tharp sen. [ ry Cory 7883 (Tr), 20233 (Tr); Tharp (Sm) "Waly, Waller Co.: Dooley 2 (Au, Ni); Thurow s.n- [say 10th, 1890] 17h PHEPTOLOGLA Vol. 9, no. 3 (Hu). Washington Co.: C. C. Albers 33017, in part (Au); Brackett 253 [March 29, 1938] (Au), s.n. [April 21, 1939] (Au); U. Lehman (Brenham, 4-21-3)] (Au). Webb Co.: Cantu, Villa, & MeCart 7h (0k); K. K. Mackenzie 96 (N); McCart 7242 (Ok, Ok); Perkins & Hall 2627, in part (Po--256397). Wharton Co.: A. R. Moldenke 209 (Fe); Be He je He Warnock 621 (Au, N). Wichita Co.: Ferris & Duncan n 3337 (Du- 12522, Gg--3140h, N); Whitehouse 10474 (Mi, N). Willacy Co.: Je F. Brenckle 7-361 (N)3; Cory 51489 (sm) (Sm); Davis & Johnston 53253. 22 (Au--122250), 53253.23 (Au--122251); M. C. Johnston 53253.9 nston 5325369 (Au--122161), me (Au--1222)3); Lewis & Oliver 52h (Nb); Lun- dell & Lundell 8786 (Ld, Mi, N); Tharp s.n. (Redfish Bay, War. 5, 1934] (Au), sen. [Nov. 30, 1940] (Vi)~ Wilson Cos: Thompson & Graham 52 (Au--123189). Wise Co.: McCart 167 (N, Nt). Young Co. Correll & Johnston 22127 (Ld); Ss. Hayes 606 (is, N); Parks & Cory =e (Tr); Whitehouse 1532) (N). County undetermined: 7; Belfrage . [Texas, 1860] (S); Berlandier 3011 (Cb), 3016 (W); Black 38 Bang (Be~l6632); Collector undesignated s.n. (Nt); Herb. Univ. Texas s.n. (April 23, 1929] (Au); A. H. Hildebrandt SMe i 329997) ; ene I.155 (W—-187092h), s.n. [Dry prairies on the Brazos, Aug. 133] (E--118198), s.n. [Southern Texas, 18)9--51] (N), son. [Texas] (W--771866); E. Meyer s.n. [Texas, 18h] (W-- 261612); Edw. Palmer 3 [Yaqui River] (i), 1043 [sw. Texas] (WJ, W—-1323112); Reeves 20h (Cs), 207 (Cs); G. Sims h2 (0k--10368) 5 u)3 Le L. Be) Ward s.n. [Texas, Sept. 11, 1877] (W—- 1 Island: Tharp 1622 (Au, Au). El Toro Island: Tharp 49098 (St, W—1972910), 49236 (st). Galveston Island: E. D. Se Schulz s.n. [Galveston ip 7/30/1938] (Wi); S. ke Tracy 7533 (Au, Cm, Es, Io--1,886 , Wbh21 190). Green Island: “Tharp 1200 (Au). icastang Island: = So ha 86 (Au—122173, Sty. NEW MEXICO: Otero Co.: Mh. E. Jones 26229 (Du--235292). ARIZONA: Pinal Co.: Goodding & Lusher 186-15 (J, Or--55219); Peebles & Harrison 422 (La). MEXICO: Chihuahua: Pringle s.n. [Valle magne wile 11, 1887] (Gg--l21277, N, st). eg Ge Le Fish » Ew, N), Lhléb (Ew, Gg—-35310h); E.G. Marsh 1625 one 2128 (st). icvaacae Waterfall & Wallis 13371 (St). Nuevo Le6n: Arséne s {Sta. Catarina, 7~1911] (B, N); Frye & Frye 247 (Se—65787, | ann). Siavbtien: Agniel s.n. [arsene 102h2] (Ur). Sinaloa: H. S. Gentry 7011 (W—-19h5099). Sonora: K- 7 . Se NA Tamaulipas: perlandier on. (Tampico, 21.5.190h] (B)s ing Gr & Johnston wade f= 73 37); LeSueur 399 (Au, Fs); Lew a 3 . is & Oliver 5422 (Mb); _ 3 Edw. Palmer 78 (Bm, Gg——21397, Ny w— 3015). Vera Cruz: eS Davis s.n. [near Tecolutia, 6-2L-L7] (Au--1232),0); Rutten & z Rutten-Pekel nar aring 178 (t—8100b) « LOCALr ee Se ts HID 1963 Moldenke, Monograph of Verbena 175 ITY OF COLLECTION UNDETERMINED: Collector undesignated s.n. [Col. Ter., 1875] (Up--17113). MOUNTED CLIPPINGS: Bull. Torrey Bot. Club 25: 617. 1898 (Ww). VERBENA HALEI f. ALBIFLORA L. I. aefee B ek Leaflet [Lower Rio ysis ita eae Club] 2: [4]. Bibliography: Davis, Natur a aaa ot {Lower Rio Grand Valley iat. “ehb) 2: iu. 19ks Moliden iit: an Jou arn; BORE 984 610. 19453 Kol eogr. Distrib. Verben » (eds 2], ah & 198. *oh95 lice: ieaune 59 py h71. 1959. This form differs from the typical form of the species in hav- ing white corollas The type of the “form was collected by L. Irby Davis near Fal- furrias, Brooks County, Texas, in June, 192, and is deposited in the herbarium of the University of Texas at Austin. The collector States that other specimens of this form have been gathered in the Rio Grande Delta region of Hidalzo County, Texas. As yet have not seen any material of it. ge HALEI f. ROSEIFLORA (Benke) Moldenke, Phytologia 1: 459. 940 Syn mnonymy Verbena officinalis f. roseiflora Benke, Rhodor 35: 4S. 1933. Verben arbors officinalis f. rosea Blackwell ex eae Résuné Suppl. 3: 40, in syn. 196 _ Bibliography: Benke, Rhodora Eo 5. 1933; Moldenke, Phyto gia l; 9 eo) or i: 51l. 19lil; Moldenke, Sup pl. List invalid anes 9 ell 192; Noldenke, ‘kioh. List © Tava tan 49. 1942; Moldenke, Known Je0er. Distrib. Verbenac., [ed. 1], 13 & 101. amr Lio ldenke, Alph. List Cit. 1: 43 (1946) ay iu: 1129. 1949; Moldenke, Known 320gr. Distrib. Verbenac., [ed. 2], 2h * 19 5. i595 ce REé- f this form was collected by Hermann Conrad 5 Benke (no. “aay at Kingsberg, Kleberg County, Texas, om March 25,1920, and is S deposited in the herbarium of the Chicago Natural History “useum. Although known thus far only from three collections, it is : : . tod. I; has been found in degen meadows, blooming fron lk “8Y. Herbarium material has been misidentified and distriou as ¥. halei small and My Peiainatia Le _ all, fel ions specimens, en examined by me. Citations: TEXAS: Goliad Co.: Ce. Be wilitiee 32 (an) Hood Co.: Slackwell 81 (Au, N, Nt). Kleberg Cont Benke 518 8 (w--15711238— gt ee tiie isot VERBENA HARBISONII Moldenke, Phytologia 1: 438--h39. 1 Eibliography: Moldenke, P Phytologia 1: 136-439 tisho) ae i: 176 PHYETOLOGIA Vol. 9, nO« 3 othe Boe tt i Known Geogr. Distrib. Verbenac., [ed. 1], 13 3 Moldenke, Alph. List Cit. 1: aos 19163 "Will & Sal- i pct aoe Supp 1. 10: 2h2. 1947; H. N. & A. L. Moldenke, Pl. Life 2: 63. 193; Koldeake, Alph. List Cit. 2: 183 (1943) eo , 122. me Moldenke, Kno ead Distrib. errae s [ed 33 & 198. 19119; Noldenke, ies é 39 & 72. Herb; stems erect, slende er, rather acu re tetras sonal, ridgee, acta densely hirsutulous-pubescent wi short, divergen Cc escent hairs; branches similar but amen nore * densely canescent hirsutulous, sg) start hi short; internodes on stems usually elon- gated, 4--3 cn. long, more abbreviated on the branches; nodes: plainly annulate; Leaves decussate-opposite; vetioles 3--10 m long, wingec-margined, densely hirsut wlous s—pubescent with ae cent hairs; leai-bl ades chartace eous, uniformly green on both sur- faces, 2--5.5 - long, 2--5 cm. wide, the larger ones deeply Ba parted ES Oe to aie base, the divisions lanceolate in out- 2 ating the upper branchlets; ; thse heart pts slender. acutely tetragon- al, ene, canescent—hirsut vemion 2--5.5 cm. long; spikes oblong, out 1. jr wide, very densely many-flowere@; bractlets lanceolate, equa ving the calyx Bee mm. easels about 1 Sse, A e tube about Pie mm. lon ag 0.9 m mn. wide at the base and 2.2 mm, wide ape el hi me: its rim 5-lobed, the Rage about 2.5 mm. long, setulose, densely pubescent ; corolla hypocrateriforn, ¢labrous on the outside, + tube about 6.8 mm. long, the limb about 5 mm. in diameter, hea rounded and bifid at anex; stamens , didynamous, inserted about 2. and 3.) mn. low the mouth of the corolla-tube, included; filaments fo, very short, 0.2—-0.), mn. long; anthers abo ut 0.9 mm. long a oT mn. wide, oat han fet style capillary, about .9 mm. die, “gle brovs; ovary oblon: b 3 ) og d la- 3 &, about 1.4 mm. long and 0.7 mm. wide, & _ The type of this species was collected ~ Charles Francis Har bison -- in whose honor it is naned -- at Agua del Refusio, Baja in the rm. herbarium of the San Diego Society of ‘Natural Histroy at San Di- ego, California. The species is ouite distinct from all the other own species of this region, alshouch its affinity is apparently with V. setacea Perry, which it re sembles. Its most obvious char are its lo onz-pedunculate, densely-flowered, oblong spikes, ne pee be setaceous bractilets, nik ab tid 3-parted leaves whose lobes are all v plainly acute at the apex. The type collection “en sometimes regarded as Harbison 11303, on this number seems t° 1963 Moldenke, Monograph of Verbena 177 a herbarium number rather than the collector's number. A note on the label of the type says "material from same lot determined as Verbena sp. by Ira L. Wiggins." Ferris identified the Dudley Her- barium specimen as V. setacea Perry. In all, ) herbarium speci- mens, goer the type, and 2 mounted photographs have been ex- amined by m Gitations: MEXICO: Baja California: C. F. Harbison s.n. {Agua del Refugio, beri 1, 1935] (Du--2h976—-isotype, Gg--328000— iso N--isotype, ’ "photo of type, Sd--1180 ila Z--photo of teat: fear grr a in Chod. & Hassler, Bull. Herb. Boiss., - 2, kh: 1056. aks. Verbena reece Chod. aos Briq. in Chod. & mace Bull. Herb. reat sér. 2, : 1056, in syn. 190) [no phlogif lora Cham., 1832, nor Cham. & Schlecht., 1959]. Verbena See riq. acd pale Ind. Kew. Suppl. 3: 187, sphalm. 1908. Verbena hassleri Briq. ex Mioldenke, Suppl. List Invalid Names 8, ins in syn. 191. Verbena paraguariensis var. latiuscula Briq. ex Moldenke, Phytologia 2: 475, hyponym. 1948. Verbena hasleriana Briq. ex Moldenke, Résumé Suppl. 3: 38, in syn. 1962. i C Known Geogr. Distrib. Verbenac. [ed. 1], & 101. 192; Moldenke, Lilloa 10: 385. 19h; Moldenke, Phytolo- gia 2: 75.1948; soldenke, oo List Cit. 1: 263. see ie Ne & ALL. Moldenke, Pl. Life 2: 63. 1948; Moldenke, Castan 117--119. 198; ie: Aliph. Tist Cit. 2: 599 ( 1918), 3: 666, 693, & 916 (1949), and h: 1237 & 1249. 19495 M » Kn H. N.& A. L. Moldenke, Anal. hg et Biol. Mex. 20: 13. 19h9 lh: 188. 1953; angely, Pl. P 1987, oldenke, R llo, 118, 127, %5, 3 van 372, aa, «le. 1959; Angely, Fl. Paran. as 78 (1960) and i "E. 1961; ieee: Résumé Suppl. 3: 38. nnial herb. to 1m. tall, robust; stems tall, green, Plaj pas vane branched; branches ascending, ie se Scabridulous; middle internodes 2--6 cm. long; leaves Re opposite; petioles winged above; leaf-blades oblong or ovate- ob] ng, the 1 ger ones to CM. wide, ase 178 PHYTOLOGIA Vol. 9, no. 3 —° orci slender terminal, spicate, abbreviated, about ong, m or less see Ee ee congested; bract- mm. in asin corolla (eine a varying from violet to rose, bluish-rose, or bluish-red, its tube cylindric, surpassing passing the c pniaetanias by about 5 m The species is based on three cit coeraas by mil Hassler — in whose honor it is named. These collections are no. 3550, from 4 swamp at Tucangua, Second collected in Noveater, 1900, and nose 6401 and 6464, from a swamp near Tobaty, Paraguay, collected in September, 1900, all “deposited in the nie er Herbarium at t Conservatoire et Jardin Botaniques at Geneva. V. paraguariensis var. latiuscula seems to have been based by Briquet on Lindman A+ 3905 from Paraguarf, segaatea in October, 1893, and later re~ determined by him as V. platensis Sprenz. “Briquet says: "Le Ve hasslerana aids ivadets au sroupe de V. megapotamica, chanaedeyier lia, platensis otc. comme affinités générales, mais il s'écarte Pkg. Ba NS inhabits bogs and swamps, campos, anc land adja- cent to rivers, from 750 to 1200 meters ’ altitude. J&rgensen says that it is common in thickets in shrubby marshlands. It has een collected in anthesis from September to Decenber and in Febru A common name for it is "flor de vovia". In all, 20 herbarium Specimens, including type material of all ary names involved and 2 mounted epee a gs have Leen exami ae Citations: BRAZIL: Rio Grande do Sul: ‘eae 52102 (Ss) Catarina: Smith % Reitz "8876 (Ok, 2261128), 8933 fosstii*) Z), 11392 Gia 2oLo SY. PARAGUAY: Hassler 61,6) [hiacbride photos 2685] re Kr--photo of type , N--cotype, a photo of cotype, S—cotype, Tl—~—cotype, “W-2086198--cotyPes S). ARGENTINA: Misiones: Grttmer 104) [Herb. Osten 23151] (N, ne 3 Ruiz Huidobro 753 (Gg—353220, | N); ¢ G. J. Schwarz 340 (N, ——s a reas cca var. OVATIFOLIA Moldenke, Phytologia 6: 330. Bibliography: Moldenke, Phytologia 6: 330. 1958; Koldenke, Ré- opal ct aie eae ae 1963 Moldenke, Monograph of Verbena 179 sumé 110 & 472. 1959. This bet agiffers from the typical form of the species in having its ee es much smaller, ovate, 1.8--3.3 cm. long, 0.6—=2 cm. Wide e filiforn alceds only 4--5 mm. long. The type of eas are was collected by Lyman Bradford Smith and R. Klein (no. 378) in ruderal situations at Rio Negrinho, municipality of Rio o Negrinho, Santa Catarina, Brazil, on Decen- ber 8, 1956, and is deposited in my personal herbariun at Yonkers, , , bo New York. The plant has also been found on riverba nks, and ap— pears to crow at altitudes of ae to 1200 meters. It % has been col- lected in anthesis in December and February. Smith & Klein 1 11638 is a tee with something spas a aceous. get = arium specimens, including the type, have been exmined b Citations: “BRAZIL: Santa Catarina: Smith & Klein 875 (W— 2251403—isotype, Z—-type), 11638, in part (Ok, W—2251793); Smith & Reitz 10203 (Ck, W-<2251662); Smith, Reitz, & Sufridini 95 (W—22h9361). WaRBHAHASTATA L., Hort. Upsal. 8 (178); Sp. Pl., ed. 1, 20. 3 Synonymy: Verbena americana, urticae foliis ete Introd. Gen. Remherb. (ahi, pl. [57]. 1690. Verbena ameri urticae foliis ancustioribus, spica multipli rg a caeruleo Herm., Fl, Tugd.-Bat. S88, 1690. Verbena americana urticae folate angustioribus, spica multiplici floribus purpureis [as "Eadem floribus shatriariten Herm., Fl. Lugd.-Dat. 55. 1690. Ver- in Sitissina americana spica multiplici, urticae foliis an— ager iy espe PU ——eeee oO Verbena altis altissima americana ‘Nemec. Parad. Bat., ed. 3; pl. 2u2 1698. ¥ Verbena americana urticae foliis ue ribus, flore caeruleo | lioris., Pl. Hi Pl. Hist. Univ. Oxon. 3: "408" [=)13]. 1699. Vervena ai tiated americana, spica multiplici, urticae foliis “tgustis, — soueekeis Herm. apud Ray, Hist. Plant. 3: Suppl. 286. 170). verbena altissima americana spica multiphics, urticae foliis angustis, floribus coeruleis Herm., Parad. ed. 2, 242. 1705. verbena altissima, americana Herm. apud ite Ind, Plant. Hort. Acad. Lugd Cana; altiss sima; urticae foliis angustioribus; spicis brevior- ibus; floribus caeruleis Herm. apud Boerh., Ind. eee Plant. Hort aren cep Lege + Acad. Lugd., ed. 2, 1: 186. 1720. Verbena, ameri altissima Herm. apud Boerh., Ind. Alt. Plant. Hort. pee aa ec. 2, 1: 186, in syn. 1720. Verbena; americana; altis sima; Urticae foliis angustioribus; spicis brevioribus; flo1 ‘ibus pur- 180 PHYTOLOGIA Vol. 9, no. 3 pureis Herm. apud Boerh., Ind. Alt. Plant. Hort. Acad. Lugd., ed. 2, 1: 186. 1720. Verbena americana altissima, urticae foliis an- gustioribus, Spicis brevibus. floribus caeruleis Boerh. apud L., ort. Cliff. 11, in ayne 1737. Verbena foliis lanceolatis serra- spicis brevioribus, ees he apud A. van pice. Fl. Leyd. Prod. 327, syn. 170. Verbena tetrandra, spici longis is acuminatis, Peres ear, oo tek, ee a 17:8. FB. verbena foliis rege Pee Sees sies filiformibus paniculatis Ray apud L., Hort. Ups. 8, in syn. 1748. Verbena americana altis- sima, spica multiplici, urticae foliis angustis, Floribus caerul- eis Ray apud L., Hort. Ups. 6, in syn. 178. Verbena americana altissima, apticas foliis angustis, spicis brevioribus, floribus purpureis Boerh. apud L., Hort. Ups. 8. 1748. Verbena americana altissima, Spici multiplici, urticaefoliis angustis, floribus caeruleis Herm. ex L., Sp. Pl., ed. 1, 20, in syn. 1753. Verbena americana altissima, spica multiplici, urticae foliis =a floribus coeruleis Ray apud J. F. Gronov., Fl. Virg., ed. 2, in syn. 1762. Verbena caule triangulo striato, spica muda a Lene tele Plauen ete ee “Totes, ete. eter J. F. Gronov., Fl. Virg., ed. 2 . 1762. Verbena ae Lan., Tabl. Encycl, Méth. [Tllustr. Gen.] 1: 87. 1791 [not Ve pinnatifida M. E. Jones, 1941, nor Nutt., 191, nor Schau., 1895]. Verbena paniculata Lam. Encyel. Méth. Bots 6: 548. 1808. Verbena americana, altissima, spic& multiplici, urticaefoliis angustis, woe, a erm. ap apud Poir. in Lam., Encycl. Méth. Bote : 5L6, syn. 1808. Verbena hastata, erecta, foliis lanceolatss sensin sp is, acutissimis, inciso-serratis; spicis panicula~ cycl. i tis, iinearibus Sees Michx. apud Poir. in Lam., t. - 1308. Verbena hastata, tetrandra, spicis longis, scininat ig foliis hastatis L. apud Poir. in Lam, Encycl. Méth. Bot. : 56, in syn. 1308. Verbena (pinnatifida), Spicis 11s formtbues foliis esses erosse serratis Lam. apud Poir. in Lam., Encycl. Wéth. Bot. 8: » in syne 1808. Verbena spicis filifo rmibus, adauresie; foliis lanceolatis, grosse sats, indivisis pm i apud Poir. in Lam., Encycl. | wéth. Bot. 8: Shié, syn. 1808. Verbena spicis paniculatis, foliis recciats, acuninatis, acute serratis, basi inciso-hastatis am. apud Poir. in Lam.. Encycl. Be thé. in eyns . Verbena hastata 6 Peal ag iar Siam vat ae wee ae clé. 1808- Verbena tetrandra, spicis longis apasaales: fol. hastatis L. = nasya\~ pud H. E, Richter, Cod. Bot. Linn. 35. 1835. Verbena fol. dance serratis, spicis filif. paniculatis Royen apud H. B. Richter, 60d a 1963 Moldenke, Monograph of Verbena 181 Bot. Linn. 35, in syn. 1835. Verbena americana altiss., spica multipl., urticae — a fl. caeruleis Herm. apud H. E. Richter, Cod. Bot. Linn. 35, syn. 1835. Verbena amer urticae w ro) . Ww Nat “ H He B . he oc Ww wal . < Oo | Co fo) S rt) 0) rH . rt) hk co p 0) n . “ ang., spicis breviorib., fl. purp. Boerh. apud H. E. sergio Cod. Pot. Linn. 35, in syn. 1836. Verbena americana altiss., spica mult. etc. Herm. apud Peterm., Cod. Bot. Linn. Ind. Alph. 196, syn. 1340. Verbena americana urticae fol. etc. Noris. apud 8 Fai Cod. Bot. Linn. Ind. Alph. 196, in syn. 18])0. Verbena americana tiss., urtic. fol. etc. Boerh. apud Peterm., Cod. Fo inn. Ind. Alph, 196, in syn. 1840. Verbena hastata Burm. pei Peterm., Cod, Bot. Linn. Ind. Alph. 196, in syn. 160. Verbena paniculata ? pinnatitida (1 (Lam.) Schau. in A. DC., Prodr. 11: 546. 18h7. Ver- 3 Palco Vars.pi ifida (Lam.) A. Gray, Syn. Fl. Ne 2 (1): 336. 1878. varvbens hastata x officinalis Briq. in Sew Prantl, Nat. Pflanzenfam. (3a): 148. 189h. BOOK REVIEW Alma L. Moldenke "Plant Taxonomy", by Lyman Benson, 94 pp. Chronica Botanica Ser- ies, Ronald Pres ss, New York 10; N.Y. i962. $11.50. Through this work Dr. Benson has made available to colleges, universities , botanical institutions, and the private — or and yist an excellently organized, advanced text and referenc Faret. it describes intimately what is involved in botanical taxonomic re esearch; it is effectively and richly pinisteated with his om fine work on the genus Ranunc ulus us. Because this group has Such a common and wide range, some es are known or easily knowable to almost any English Tanciage reader. The same is true ~~. the genus Quercus, which is used very effectively for further illustrative purposes. explains sraphically how data 4s collected from her- = * *ariun preparation and consultation, from field observation, cae ~*SFoscopic morphology, from fossil Se ae oe ie Physiological experiments, from ecology, from gross and oo 12s at Stics, and from a synthesis of ic peatin from all of t disciplines available. ; 1 ta ist and a Only a ver; cood and thoroughly experiencec taxonomis 182 PHYTOLOGIA Vol. 9, no. 3 superb teacher could have written so lucidly the Pag oc on the principles of plant class sification, the techniques of naning plants, the survey of the Code of Botanical omer tater the construction and the use of keys, and the documentation of col- lected plant specimens The final section of the book deals with how materials and methods should be handled in research in this and how they da, should be presented in a treatise or monograph form, even to the etails of correcti alley proof and arranging bibliographies. momy, can make chiwiowed use of their preparation time b staring this book. writer of this review would like at this time to append a note £5: her earlier. basi ? ically very favorable review of Dr. Ben- son's previous book, "Plant Classification". The 2 Jacket s commended this te: r beginners! colle e biology or bot: courses. I thought that it was (and is) the cinaen in the field at a slightly higher level or for course enri stants I now must "dorc-earedn Sodaune of very heavy use by two different nigh school students who have gained very much from itt NOTES ON NEW AND NOTEWORTHY PLANTS. XXXVI Harold N. Moldenke ak CRENATA Moldenke, sp. no tex; ramis crassis Sabie dense pubescentibus; ramun uricostatis dense pubescentibus; foliis Saree: revis ime multiflor on= Shrub; anches rather stout, not plainly tetragonal but PF sed with parallel es ribs, densely pub- escent with i j otherwise $ principal internodes 3--); em. long - 3 petioles very short, 1—3 mm. long, aocealy ’ pray’ Se ee eee 1963 Moldenke, New and noteworthy plants 183 ish-pubescent; leaf-blades osteoid Kenia cepalh ae even in ellipti ic to elliptic-lanceolat r else slightly oblance- olate, 3.5--9 om. long, et cm. wide: aetaee at the apex, reg- i ase astomosing ne e margins; vei t reticulation very abundant, deeply impressed above, prominulous beneath; ane escence 1 ternate, spicate, the spikes elongate, ong, Vv ary, o 22 cm densely many-flowered, the peduncle and r sit aac ate patted aa pubescent; flowers sessile; bractlets subulate, 1--2 mm. long, densely grayish=pubescent, shorter than the calyx; calyx cylin- Cric-cupuliform, 3--); mm. lo ong, about 1.2 mn. wide, densely gray- ish-pubescent, the rim shortly 5-toothed; corolla hypocrateriforn, its tube )--5 mn. long, pubesemt on the outer surface, the limb about ) mm. wide he type of this handsome species was collected by Karl Fie- brig (no. 6137) in the region of the Alto Paran& River, Paraguay, in BPO 9 or 1910, and is deposited in the United States National barium at Yashington. oe Seer cme aca ioldenl e, Sp. Frut an amulisque grac ilibus ieee “tetragonis densius— sos cineteo-mibes samecerst am foliis decussatis vel ternatis; peti- olis gracilibus densiuscule puberulis; laainis chartaceis oblon- sowovatis supra leviter subtus dense puberulis; inflorescentiis wdllaribus terminalibus a cymosis multifloris laxe multibrach- iatis densiuscule re ee calyce obconico densiuscule __ Shrub; bra Pilose with more or less =a rsely subappressed hairs, 128s | so or clabrescent in age, obtuseiy tetragonal, pig youngest o — principal internodes 1--6.5 cm. long a = lance, t osi nate; petioles slender, 1.l--2.l, cm. long, rather pana pilose like the branchlets; leaf-blades thin-chartaceous, oblong-ovate, h.5=-20.5 om. lo 5—5.5 cm. wide, acute at the apex, entire, rather lightly sth ee to ent above, pore densely so beneath; midrib slender, flat above, prominulous e > n b GF ary hte dichotomous, pilose like the peduncles; calyx obconi ano -ts tube about 2 mm, long, the rin 5-aristate, the tube and teeth + 72 Sta * Father densely pilose; corolla herpoeraterifom, its tube vary 18) PAT TOL060 Te Vol. 9, no. 3 slender, about 9 a * the limb about 6 mm. wide. The type of thi ecies was collected by Mrs. Helen G. Faulk ner (no. 2785) in cuertut aes shade or seni-shade, at Chulwani, Zanzibar, at sea-level, arch 21, 1961, and is deposited in the herbarium of the tee cetostee, Riksmuseum at Stockholn. minute strigillosis; laminis folio embranaceis olivaceis ovaq t subacutis vel obtusis, ad ba brupte acutis, margine regu ter serratis, utrinque parce strigillosis vel ragonal, minutely puberulent or glabrescent in age, Sehckege stramineous; nodes annulate; principal internodes 2--3 cm. 10N8}5 leaves decussate-opposite; petioles slender, abbreviated, apes. - long, minutely strigillose; leaf-blades ‘membranous, olive- green even i rying, somewhat lighter beneath, ovate or to t rse gob es strigillose or pilosulous on both surfaces; midrib very pee secondaries 5 or 6 per side, very slender, ascending, not pe arcuate, termi inating in the simises between the : margin nal tee long and nee denote “Satgadiowered; lower btractlot 5 enlarsey ee to 7 mn. long anc 2 ma. wide, attemiate-acute at ra apex, sparsely pilosulous, subequaling the corollas; corolla Ny pocra cageesuiapes rose-color. he ° £ this species was collected by Santiago venturi Bo) -- nan am é baer at Tilcara, dept. Tilcara, at an altitude of 3000 vistors he pod pi obinemanit on Febru ruary et nh and is deposited in VERBENA RADICATA Moldenke, nom. nov. 20 Verbena radicans Gill.’ & Hook. in Hook., Bot. Misc. 1: hoes 1829 [not V. radicans “'oench, Meth. 369. 179]. VERBENA meieasi tos Pree gg (Hicken) Moldenke, comb 903. Yerbera r glabra Hicken, Herwiniane 1s 56: ‘1 Mapping by counties done by Andrew R. Moldenke Te . . . . . y 2 tT —e Figure 9. Distribut f Verbe hal the Unit s+ 5 . Distribution of Verbena halei in the United States x is 7 ar ’ = 9 T 10 ll m _ j [ aS 33 u 35 36 7 % 5 = _ \ I A . A L | 8 ‘83. ( A | oe . 1 2 2 ‘ | “Ar (ee ma NR ro ¥ T - : W AN BoA fe) N — ns e “pe, * nine Suma se | TG / ince . Py * ill ni . tees "a c Ps se | | Sone cg Fe s Hy E 2 f 1 s ) a Uv ™, Uagary ‘a Se Q = Se Sug se Sea POmog es PLLips s Cc my “pe Moen; od Tom oats rT D CnUTEAG } Cd oe | REWRLE | soTTincAU neuen ’ TOWNER | CAVALIER PEMBINA gy KITTSON ROSEAL orn Les i Mc HENRY AScaog MOUNTRAL s PIERCE e WARD RAMSEY WALSH ‘MARSHALL > Fe RGus o ec ome BENSON a D ou MeKENZIE ROYALE as ss ' NELSON GRAND FORKS yg POLK | PEMWINGTOR as 2 re oT Siemon) TH asim Wo COOK QoQ use E | MCLEAN \ J Mee N one cee Hy N = sae 7 oF c ee st fe) = TASCA £ R H D ¥ 5 A MERCER TRAIL » My Bd ¢ pees! wert as N Ame . PLlmcs FOSTER coeccs Noman Punnoute v As » . bat Seti Peséaue | NER sumeice | KIDOER | STUTSMaN STERE coe] HUBOMRO Hs ¢ ° i ee Pp valey custen ' = eC — (se ° *% rm. E mn] Pans - aa ABH caaba ouy (ECKER — .Y WS ” / REASURE WALLEY uw wet . F SWEET Grass Cr) CONTOWAGOR MARQUETTE 4 “™, YELOwSTong SLOPE arias é — Ty ® BAYFIELD \ cnrrewa OS aa Py a (i ) Eamons | LOGAN UA MOURE RANSON OTTER TAR ° qocenie a Mu ase! someon oar = << ton" STUwaree 816 Hoey 5 Soman 1 \ ADAMS svoux ‘oman M I aa. 1RON — » DICKINSON! 7 DELTA = MciNTOSH DICKEY wus fy) cm cs vo ToDo Crom WING iL LE @ OWDER arveR heme) cos e a PINE eaSwBURN| SAWYER SHUAND e <2 PERLINS =- 2 cat | oueus . Pact ( ai 8 seco x oa F — Loh | onsen CAMPBELL ONEIDA FLORENCE ers eet mar ent a TeNTON BURNETT WARINETTE e st e ARK N "ARK seweas és : . STEVENS | POPE STEARMS SANTI — FOREST s 4 —, Loy v wo a MG WORN = 2 TIEBACH ‘apaaesiie g F LINCOLN san KOS STSECO| Somer wer 7 So. Dewey 7% “ _ a aaa i tf ned ‘ e ort RANOFFOR! ANOKA 5 TAYLOR * ri a " 7 oR a ‘ POTTER werner f weiGeT j = ca —] CHIPPEWA ats aan YOSCIOA rc iS ® Kom Siac. MEADE . w e eas JOHNSON ARMSTRONG fa =15 ee ‘SHAWANO ie a a SULLY Wel E00 — . Seovcest_ WL Dabwaw 10500 c WASHAKIE STANLEY aot RERVILLE CARVER PIERCE W I Cc oO N seme < pos meron [mss on e {7 aR . 08 io ‘ Fre . S YELLOW MEDICINE Eau CLAIRE WOOD [PORTAGE | WAUPACA Fait = < ° LAKE oN i : se nor ammenct T . sey) Tae: suauey 7 ay wan | 3 BROWN © sre ° - Loin ce CHEYENNE WALLACE | LoGaN cove Samal cae a TINCOLN Fea poncaanrn mare — pexson seu =, Fg ennct é yn ‘ell [aoa pry - Ny A M Chareey_) FREMONT . pa we... =r RSE r iss 1S, “ 2 Z = : ~ omen wOurs, om : bw Pn N = SAGUAG . WABAUNSEE fee UBAYETIE CALLAWAY Ae > mac ) i ~ . ) wae cneeey | wicwta | scort SAGE | DOUGLAS | JomsON IF cass SON Vialead ees Wansen |crames / cauensnt®, sacaamost Tacs fame pl a, H 4 iS pom wa be =e hv aye 5 povertel 2 orl a . PS — J . P i - Neo Sot “ee rs ares paar ret ] rm — _ Pr CA we mee fees < | ® | | | poe cc) Osase ? hae a Leso = Seay oe am — I | | [cay es MS | SEWARD LARK ADE — ‘ sae u ue " ~ - % be como) aid cowLer pweestea] WRIGHT wnt en, Wwe — SA oo ae bal i) ye Noe f HARPER GREENE ee 80h To ARUN rae easHintO™ \ Sears eockine? ‘] = A wane » Py s ye ‘sure? STORES. ai \ 4 = | | Tamacnct ey ~ pants em oot A ee Laan wa TEAS to % ce ret t WAR, 5 2 BEAVER 3 3 ca aT e a SU GUILFORD nce Mery, 2 woos CHRSTIAN soul “Ss vo om . “s TH “orm a : wae ee = & * _ CARTER BUTLER srr Sam na a — raat scot are ~ em 1 canter es bs yan ie wn on \ 0 a ‘Snooval - 3 Bane = on ‘OREGON ey = Pe Fywmen] wacor | CLAY - car me Scent 43 van ane e a 2 wot % s Oa; —_ 4 a — : = FI BANOOLPH Toya ~ . wORA LAM . . Tyton ofant - - ~~ i x sneman Tsansroro Tocuu reer Turse BUS] wooowaRD camrifid ‘| WORE 3 rer roe = ¥ So von cape) L a = ed (0s wake — ci Sens — in a a . . « : ant} OBION - ay of WILSON, smi q JomnsTO) ed Cale Gort . vig ia PAWNEE as coy cannou | BOONE WON) BAXTER | FULTON RANDOLPH mscor wan PY Ge PUTNAM wart | . | SAN micuey ohana . : = = bi moa c jones wooRe AYE RO SHARP CARROLL ayers, ure cS ue Ss D ie \ YALBICA * ; ral Varia aa sn , wanison)) NEWTON — “a “Ht ‘ape * Quay WAGONER re Searcy ee Vv —s — B racer nis TINCOUN CHEROREE Crennnce oo Se, one sas — , Be Lo ] POTTER CARSON = po oe Pre < aan ecwoeesn) (PERRY VGN gy sited prs v | co - JOHNSON | POPE VAN BUREN [CLEBURNE — “-— mu \ - GUADALUPE 4 H SEQUOYAN pone mars), ra} tes, Carne rfeecnct] GES Sy HOM cESTER eat Tips paann® UNiON Y one 9 ORRANCE iy Bane ae Becenam — | WASHITA CANADY ORLAMOMA ae — THOS aranetn SameY Y FAYETTE arom iwcoun c ancasTen Ee om 4 wren E x ARMSTRONG | Dowity Jooumereonr GRADY LEMOS mn OW saa me TOCaN WHITE i} ” e * me Aarsnew \ AZamncon / pint = I Oo = cater Kiowa _— fwooomurt| ST FRANCIS ° nasot man) YACRSOM LAUREN, ‘ rf wan panic, — may t . Rabe Le Pane MER west ‘ =. nT omenct P3 PARME DE Soro x cA ® [caste | swine —[aecar MecLAIN SOIT TH oe wows es e MALL mn DRESS COMANCHE 5 was” a ACHLANDA rT TiNcouN TACKSON a a) a PRAIRIE = fiat TWPPAK i oon at cnet Rae +. PULASK’ Foro Pen we “A STEPHENS POU rooms UNO ous/%o f in . coseve:t Weaney [uae i tquone FARA | yyavertt \awnenct s MALE Flow ‘ LEE |rewnemtng WINSTON | CULLMAN ct CO Ly ental) Canes Worley [comme 7] “AROeMAN| = © Corton ONTGOMER Cant | RPFERSON | amtansas aT wan in esaras ( HoT spring =| ° or org men 9 5 COAHOMAL sana pau Cc ORANGEBUT J COCHRAN cuy evien ae TRUS - racomusn a nounf ones] MONROE — BOUT) yyut YANO ae re —€ penntitt ag , wooneyiueeocx | caosay | ooans Tame wig CLEVELAND f LINCOLN pa 2 ier saree aan) On Laon LY K arerc| woven KEYED TO COLUMN A raLLanaTe Z = > 7} + knar BAYLOR = 3 — os LEBURNES cag Oh al oo saa LouLETON of Cleartype Index Book i aD erven P wevada TQuAcHIT ° soar] WEBSTER Gar cS cat cana\ Om 4 ay OS ys are ce nin a PADLEY| BOLIVAR FLORE se TEBE PICKENS = — ae nal es CHARLESTON Y rc) GARIA [oT Sean ‘a LEFL ba TYSCALOOSA one ee Fite [re ve CA 2] sence & R § DONA Ata } HASKELL JACK CATE Q ° LOWE! - woctamh own 7 i Ar xi z [ts Pty at § AITALA wonveee a on a — ne) a eel ‘ i S = T - a Z cant, * GAINES ms z bard apes rents CHL TOM wr na ag (on) euncor tae 7 s Bonen] scummy] Fisnen [Jone 2 COLUMBIA s . as sae winSTOM es B AA a A > ¢ ‘ —— aS = 2 WE] CLAIBORNE wennaus J af SpE) Teme Gi a os es ~ = a W008 | STR ~ 4 fancon A De UPSHUR) Sir ¢| AUTAUGA wort (x “as Md MARTIN : : eR M6 ages veswona | _ SEMPER Pena “a of 00 — r\ a poe F 7 — - I sowaro | wiTCHELL SOM UNCOUN 5) sau! TAADISOM ScorT [newTom [LAUDERDALE cen DALLAS eerie WOM css = =e) arta f Yacare 10mm z Fricnano ¢ MAD! J exen) WHOS my, _ ? BuLLOCX scat | oar ae vrompy TRADE MARK REG. U.S. PAT. OFF. . . cAD00 = Lome: sre ae wart ECTOR r MYILLE RANKIN = a - ant T s ie wiotawo | cuasscocx [sremine | Con La os PANDA rs a Tarn Pasren | CLARKE FI wicox ay eee) Se = DE SOTO piven 7 CALOWELL TENSAS 6 punt BUTLER PRE} BARBOUR, pa on oe, WARD a Bosque 87 anit WONROE aoe : F a aa ie REAGAN NE 7. Feasanty ~ JUDI ap ml lea FREESTONE ETT melo Corian cocepones | | caunsmety COFFEE | DALE a Tee amon aan = : CIAL DIVISION : — us \ ANTON LIMESTONE 1 TNC | sex = = ot - é eee Be 4 sem = uu werowt \ E x MeCULLOCH CORYELL “j EORREST | PERRY | GREENE 1 : — Tey La Tocaren [Oe Teas [0% \_ ones bel A 5) PIKE . P-! —_ COWINGT s LOMOLS ass LAMP, s « La = " cRocxeTT aime . HOLMES Dw '. oft Fe ss wore Sm = Xp tt ae Scale of Miles —es SAN SABA ois TRINITY . . a ceonct Hose Son . MASON a" east st WASHIF 2 wer 1’ FOURTH o — ee Feamaf riicame pret ” eel eon WADISON / cyan Te 0 100 200 dh JUDICIAL DIVISION PESO —— we) > \e 45 aver Toe ‘ooms 4 a Pde om st \ ure Cy WAKULLA ny BLANC Ue ag DIN aa remastom ee er arAvETTE [osy MAP NO. 125 owes FRAgKL) Fae om NDALL an = c U RMILION ; Enamit st aes E / ‘ty ~ AMERICAN NY, INC 5 ’ . LEY st wary EDINA bas Txt “ oat eno Ly] foe rms aie ORIGINATORS BLISHERS = x TYPE MAPS 3X OLORPRINT MAPS sce SD w Pan YORK v NOTICE—This is a copyrighted map, The law . 4 a aL SOR poe prohibits the reproduction or copying of sme, H A w A I I MULLEN i or any portion thereof, by any proces for per n Cm sonal use or resale, without permision 4 HONOLULU — 7 7 T r | P PART a > a pot er Copyright, American Map Co., Inc., New York, No. 14324 ' warn ’ % ¥, x M oaHU yy assaih fae we W ‘ Cs - - ‘ 4 = ee, I f * ee TY DP ° FIRST y, KALAWAO a - iviSt FRO = a . JUDICIAL DIVIS 7 MOLOKAI > rary 0 Scale of Miles 99 - ) 7 ees ey ce Y nog a x / Moy, a won, e ovay HONOLULU , ge PART 4 LISIANSKI, . = — . ~w WAYSAN 8 icind | ‘ S., ALEUTIAN ISLANDS ] Pad . THIRD JUDICIAL DIVISION s * ¥ z S aif a NECKER ° ‘ pon - g *e @ Poh . ™= Q Miles 309 — Z 1 mt : i 9 a 10 I ii T 17 ] 18 I 13 T 2 T = T = T = . a u & | 26 it 27 | 2 ] 23 T ) T TT 4 PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among ae contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. number consists of not less than 32 pages. All manuscript accepted will — es in the next issue, so that the size of numbers may beds y- A volume will contain about 32 signatures, 512 pages, or a SI number of with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. NS extra charge is made for line dra wings, such as are ordinarily reproduced in — for diagrams, tables, or eects provided they conten to certain tones, dependi. about $2.25. dione with research in all lines of botany, in any reasonable : sketches, and critical] reviews and summaries of liter- ature will be considered for publicati amateur. r the present ent cost of printing, the basic rate fo a page or fraction — oximat Y $8.00 for an edition of 250 copies. "This price is subject : ce, since it depends entirely on the prices prevailing it PHYTOLOGIA Designed to expedite botanical publication ee October, 1963 No. 4 Wy 2 ‘alee : CONTENTS — LAUGHLIN, K., Taxonomic problems of Crataegus, with special reference to species of hybrid origin ......+++++00++++> eas 185 MOLDENKE, H. N., Notes on new and noteworthy plants. XXXVI... 186 MOLDENKE, H. N., Materials toward a monograph of the genus Dereee A 2 ee ea ee ee 189 MOLDENKE, A. Book reviews ©... ccbc wees es pee ee ‘SMITH, L. B., Notes on Bromeliaceae, XX ..--+ee-++e+eererrrrrt* 242 - EH. LB. 5 ynopsis of American Velloziaceae Supplement Ill...- 262 _ Published by Harold N. Moldenke and Alma L. Moldenke — 15 Glenbrook Avenue, Yonkers 5, New rome; be ‘Price of this number, $1; per volume, $. 75, in advance TAXONOMIC PROBLEMS OF CRATAEGUS, WITH SPECIAL REFER- ENCE TO SPECIES OF HYBRID ORIGIN Kendall Laughlin 165 Pine Ave., Chicago, Ill. nce there seems to be much misunderstanding about Crataegus among uninformed persons, particularly in the East, I submit the following statements and Conclusions regarding the Hawthorns of Cook and Du Page Counties of Illinois: , - There are at least 26 readily identified spe- cles and 5 varieties in addition to the types. 2 i within a Origin are more restricted. : 3. There is no gradation between species. 4. Knowledge of 79 characters is desirable to identify a plant. Some characters are more important than - Mo tinctive that they can be identified with reasonable certainty by their leaves, bark, thorns and manner of growth without recourse to floral characters. : - Nearly one-third of all the described species are of hybrid origin. 8. Individual undescribed F, hybrids are quite rare, but a number have been found. sa s - The taxonomy of species of hybrid origin is no different from pure species except for the problem of deciding what series to place them into where the par- is vitiate ch as n pasaus and no evidence that this genus conforms to endel's law. 2. An inbred hybrid population of great age may have developed characters different from and not 186 PHYTOLOGIA Vol. 9, no. e Ame showed that C. hillii is of hybrid origin, a hypothe- sis morphologically verified by the odd number of its stamens. Its parents were the Downy Hawthorn mollis and the Scarlet Hawthorn pedicellata. But the nutlets of hillii are 9 mm. long, thin, and acute at both ends, a combination of characters not found in either parent or in any other species of its region. - On the assumption that "classical" taxonomy is the proper base, adequate treatment of Crataegus requires detailed descriptions of all species and va- rieties, regardless of their phylogeny. ov? (¢°" NOTES ON NEW AND NOTEWORTHY PLANTS. XXXVII Harold N. Moldenke DOMBEYANA var. ESPINOSAE Moldenke, var. nov. Haec varietas a forma typica speciei calicibus sub anthesin variety having its calyxes at time of anthesis 10--12 mm, long. The type of the variety was collected by my friend and col- os Dr. — Espinosa (1899-1950), no. E.1805 5 7 re © honor it is named — at Zaruma (alturas de Viscaya), # altitude of 1530 meters, El Oro eee. on August 18, 19h7, ane sion was a great loss to South amerix nee an botanical science, "Arbusto de y largas. Hojas color verde claro. corolé color lila muy claro. Frecuente ." TANTANA CUJABENSIS var. PARVIFOLIA Moldenke, var. nov. This ety differs from the typi a gee cies in cal f of the spe paride Fog leaf-blades Veubeiilate. 2: fee ili long, and 1—2e1 2 Miranda was collected by N. Angulo and H. meters, Jalca de (no. 1346) at Jalca, at an altitude of 3360 2 on July » Province of Utuzco La Libertad, Pert, at lakes: srr? sposited in the H. N. Moldenke Herbarium 1963 Moldenke, New and noteworthy plants 187 TRIFOLIA f. RUBIERENSIS Moldenke, f. nov Haec forma a forma typica speciei recedit laminis foliorum ar- gute acuminatis et marginibus grosse crenatis. This form differs from the typical form of ‘the species in having its leaves sharply acuminate at the apex and the margins coarsely crenate, The type of ne form was collected by Mayeul Grisoll (no. 15) at La Rubiera ar Calabozo, Guérico, Venezuela, and is depo jeposit- ed in the United . athe National Herbarium a + Washington. scula ramosa; ramis cand Tbh obtuse tetragonis sulcatis- que ilvaite puberulis; ram ramulis pergracilibus puberulis; foliis op- positis parvis, laminis 6—10 m. longis 3—l, mm. latis obovatis obtusis vel subac ped ad apicem parcidenticulatis » utrinque ute albido-strigillosis; inflorescentiis axillaribus oppositis subsessilibus gereie: multifloris cinereis. Small shrub, app arently considerably branched; branches slen- der, bromish, ’ obtusely tetragonal and margined, striate-sulcate between th the angles, minutely puberulent; branchlets very slender, ternodes rather uniformly 1—1 5 cm. long; leaves decussate-opposite, sometimes somewhat fasciculate, sessile or subsessile, thick-textured, ob- ee 6—10 mm }+l, mm, wide, obtuse or subacute and many-flowered, gra Poorater‘ form, about 5S mm. long, white, densely white- The type of this species was collected by H. Arnaldo Lépez Miranda and A. Sagdstegui A. (no. 3426) 8 Sao, slope be- tween Huaylillas and Tayabamba, province of Pataz, la Libertad, Peru, at an altitude of 2350 meters, on May 21, i961, and is de- posited in the H. N. Moldenke Herbarium at + Yonkers, New York. PARP eo CHOTOMUS var. GLABRESCENS Moldenke, var. noVs b Haec v. vertices a pean typica speciei foliis extra basin gla- ris Soe rec ety aorexs "from the typical form of the species a oe having its leaves, except for the very base, completely on tho type of the variety was collected by John J. Wurdack (no- 1388), locally frequent, in clumps, in the Jalca zone gone 1—5 km. west, of Molinopampa, at a al altitude of 200—2450 meters, prov~ eo , Peru, on July 18, 1962, and is : deposited in the United repens National e o noVe Herba perennis; nny annie page a 8 fe dense folioso; foliis 188 PHYTOLOGIA Vol. 9, no. h oblongo-linearibus usque ad 1 cm. longis ad apicem subobtusis vel is supra dense adpresso-pilosis, pilis albis antrorsis, sub- tus glabris; pedunculis terminalibus solitariis, 3.5--5.5 cm. neis 1.52 mm. longis 0.5 mm, latis ciliolatis. : dais herb, forming dense mats; s numerous, branched, 5 apex; leaves numerous, oblong-linear, very pronouncedly gray- green in appearance, to 1 cm. long, about 1 m, wide, esate or acute at the apex, opaque, not fenestrate, densely appress pilose with white antrorse hairs above, glabrous and shiny be- ppr antrorse hairs, very conspicuously and densely long-barbate at The type of this distinctive species was collected by John J- pitieag pees 1081) -- in whose honor it is named —- locally abur meee clumps, in the Jalca zone along the Yambrashbamba— cha trail between Yanayacu and Pomacocha, at an altitude of 2300—200 meters, province of Bongar& on June 2: onas ’ 28, 1962 i 2d States National Herbar ius at Res sopoeited in the United States Na VERBENA PUMILA f. ALBIDA Moldenke, f, nov. forma a forma typica speciei corollis albis recedit. a This form differs from the typical form of the species in ving white corollas, The type of the form was collected by Robert Runyon (no. 2499 in sandy loam, open sandy ground, at the edge of Kenedy County ene in Lantana. Unquestionably more will be discov” *conatheg ag ec ia Of these and other verbenaceous genera im- eas * field notes become more complete. MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XII Harold N. Moldenke 21. 1552; eee . Pemptad., 149—151. 15835 Collect. 113. 1587; L'Obel, rahe Stirp. 1591; Gerarde, Herbal , fige-1& & 2. oe Rar e ° R ee os 2 re & 360. 1808; Raf., Journ. Bot. (Paris) 2: 170. 1808; urn. Bot. (Paris) 1: 228. 1809; Sims in Curtis, Bot. Mag. er i. S76. 1818; Raf., First Cat. Bot. Gard. Transylv. Univ. 16. 182); Torr., Ann. Lyc. Nat. Hist. N.Y. 2: 23h. 1827; ten, Nov. & eer Cog. cy oe 1: sed 1828; foams i Scart : 82h. a iB, 1656; anohettouns in G, polwetats. Beitr, Fl. esien. 150, 165, 167, & app. iv. 1878; Batt. & Trab., Fl. Alg. 1: ny. 1888 Hook. f.& Jackse, ind. me 2: 895, 1161, a er ial I 12h8, 953 N. L. Britton, Man. 7 - 1900; Fiori & « Paclet ren. Fl. Ital. 2: 380, fig. 3219. 19025 Dusén, Arch. Bo Jan. 13: 36 & 9h. i5035's Schaffner, Ohi o Nat. 73 Bie =3i,. 190 . "556 1s yee ag) « 155. 80 ih, ays: "a ge. 1916; J. H and 2: oe & ° 3 aioe "7b, 655, a. 14, Gard. shes 76: 25. 192h3 Se Arnot ser. 3, 16: 267. 19245 '"C.", Gard. Chron., ser. , 76: 279, fi8~ 99+ etta? Trivetts, Cat. 1932; Wadmond, ra 3h: 19 Triv- ts, Cat. Suppl. 1933 Novelty List 6. 193 40s 2hh— 33 , Rhodora Mo —2h5 « 1938; N. ras Petersen, Amer. B ay : 166-167. 19393 a in Contrib. t : er, 4 Friesner ‘ Acad, Sci. 55: 56. ° “19 Acevedo de varene: Bol. Mus . Hist. ° 5 ey 1951; E H. C. Greene, Am. Midl. Nat. iB: 52. 1952; oH Greene, ‘Biol. Abstr. 27: 1865. 1953; Biol. Abstr. 27: 3735 3768. 1953 [not Rye auper Press, Little 56. 1955; at mee ee 21,0 & 28h. 1955; Mar- Dict. Terms 7 tines ¢ Crovetto & Rojo, Revist. Invest. Agric. 11: 129. 1957; Mat- 189 190 PHYTOLOGIA Vol. 9, no. k toon - Buyers Guide, ed. 6, ge Page 1958; Lundell, Wrighti 2: Wee, 19603 Pe Jaeger, Wonderf. Life of Fls. 12h. 1961; Zohary, Pl. Life Palest. ct 1962; Soret Biota ls 191. tid Bare Angely, Fl. Ao Bs Paran. 22: 39. 1962; N , Bull. t Gard. Lucknow 76: ok. 1962; Serie Résumé rep = 1—ll. 1963; Vaughan's Gard. Illustr. 87:5, 57, & 67. 1963; Gleason & Cronquist, Man. Vasc. Pl. li, 579—581, 807, & ree 1963; Park's 9 Seed iso 53 3 a 1963; Haggerty, Our Parks 29 (1): 1. 1963; pine Ta 43 19635" son Toes ubrosis Semesinnenamenttiiiiee iene Ferber alts — fuchsii Gesn, — in the Lamiaceae Verbena, “floret August August. & Sept. semen fert. ert. Octob. Gesn. — in the Lani Lamiaceae Verbena foemina seu supina fuchsii Gesn. — in the Lamiaceae Verbena lutea f fuchsii Gesn. — in the Lamiaceae Verbena quaedam rario rarior (8 provincia) pulchrior & maioribus Soe “gi bend me mini, floret Iul. Gesn. —- in th imiaces, Oe? et Soukup (1 eee ote As Ginn ck, as "China verbena", but in all probability it is merely xV. hybrida Voss to which = is here The Collector undesignated s.n. {ex Chili] distributed as o, Verbena sp. in the Lund herbarium is actually Phyla nodiflora val rosea (D. Don) Moldenke, Oemler s.n. [1837] is . Persicaria caria hirsuta (Walt.) Small in the Polygonace eas, and D. S. Correll 26588 is not verbenaceous, Nair & Reiman (1962) give an interesting "key" for the identi~ fication of five species of this genus by their pollen-grains: 1. Ectine Rese gramlate, 2. Bndocolpium faint Ve phlogi tiers Cham. 26 oo. clear (lalongate) V. bonariensis L t diam sooresceee® » Longes eter of endocolpiun 12—-16 8 ce. officinalis re L. 3a. Longest diameter of endocolpium below 10 pa eaee aa es rigida ee la. Ectine surface psilate my, pinnatifida". © they later state that the endocolpium of V. bonariensis +f aes * Prong Ag it a not clear how they expect ; the above Ganon 0 be t they are referring by "V. pinnatifida" is Briq. explanation of the abbreviations aig ae ols used for the names of the 269 herbaria in which Verbena material herein in cited 1963 Moldenke, Monograph of Verbena 191 TA 53 Re (1955), 6: 2h2 Bais 73 Zi~98 2 (3959), 7 GB, rer (1960), 7: pel G se a : 95 SS 61), ng additions: Ag = terto Arizona; Hi = = University of North Carolina, Chapel Hill, N Carolina; = He limann Herbarium, CIBA, Bas Basel, Switzerland; Ja= J, de J. Jiménez Herbarium, Santiago de los Caballeros, Dom- inican Republic; Og = P. Olrog Herbarium; Pm = Pharma Herbarium, , Basel, Switzerland; Rf = Texas Research Foundation, Renner, Texas; Ul = Centro de Botanica, Junta de Investigag$es de Ultra- gl Lisbon, Portugal; and Ws = University of Wisconsin, Madison, nsin, I enke Additional bibliography: Moldenke, Phytologia 8: 378 & 396 (1962) and 8: 60. 1963; Moldenke, Rés umé Suppl. 6: 3. hed Howell reports the stems prostrate and rooting, the ores- cences ascending, 40039 (Z) citations: CALIFORNIA: Monterey Coe: J. T. Howell et cee bliography: Moldenke, Phytologia 8: 146--148 (961), anor Hee : 397 (1962), —_ 8s - 1963. imen ed by Macbride's photograph 17399" A$ mints the Berlin one before World War II and is now royed, mae Seadeooner ons: BRAZIL: State undetermined: Sellow 5.n- [Macbride photos 17399] (iphoto) « edt ccoet Collector undesig- hated s.n, monte Video] (Lu siaietoant waiasean Logia 8: 379 & 397 onal bibliogra hy: Moldenke, ris : sey 399 (1962) and 8: 460—L61. 19635 Moidenke, Résumé Suppl. 6: 3, h, 8, & 10, @ditional citations: TEXAS: Jeff Davis Cos: Ae Re Moldenke 68 (8). Pecos Coe: A. Ee soars 69 a. 650 (s). NEW MEXICO: Lin- = ar A. Re Mo decks 637 (S), 638 (S)- Otero Gow: Ae Re Molden ene AMBROSIFOLIA f NDULOSA Perry 1 Additional bibliography: Signy Phytologia 8: 379 & 399——4,00. 962; Moldenke, Résumé Suppl. 6: 3. 1963. ee tional citations: NEW MEXICO: Hidalgo Co.: As Re — 780 (S). Luna Co: A. Re Moldenke 630 (S). ARIZONA: > Cochise a) Ae R, Moldenke 782 (S), 70u (S)$ Moldenke & Woods 611 (S), Be (8). — _{avcama B- eles “acevedo de Vargas, Bol. gee cd Nat ch chile 2 251 50 50. 1961; Moldenke, Phytologia 8: 379+ 1962. 192 PHYTOLOGIA Vol. 9, no. k S. Moo tional bibli tingiaptirs Mold » Phytologia 8: 189--191 & 10 (4962) and 8: 61. 1963; ada: Résumé Suppl. 6: 7 & 8. 1963. The Moore specimen represented by Macbride's photograph ane pes in the Berlin herbarium before World War II, but is now de- Te peat ideas citations: BRAZIL: Matto Grosso: S, Moore 1083 {Wacbride photos 171,03] (N--photo of type). ATACAMENSIS Ri Additional pty Perea Acevedo de Vargas, Bol. . Nac ‘hy ~ peg Nat. Chile 25: 51. 1951; Moldenke, Phytologia iy 379 & YOU. 1 BALANSAE Briq. x) Additional citations: BRAZIL: Santa Catarina: Reits C.583 (N)- BARBATA Grah, Additional bibliography: Moldenke, Phytologia 8: 201. 1962; Moldenke, Résumé Suppl. 6: 10. 1963. VERBENA BERTERII ome Schau 3 Mewortiia berterii Meisn., Pl. Vasc. Gen. 2: 198, 98.1839. Stuttiemortne Additi Les igre Saw f. & Jacks., Ind. Kew, 2: 398 & & 1178. 185 1895; pera Aage argas, Bol, Mus. Nac. Hist. Nat. Chile 2¢: Si--s2. 1951; aaa” Phytologia 8: 380— 381 & Ol (1962) and 8: 461-462. 1963. aterial of of this species been misidentified and distribu- ted as V. radicans Gill, & Hook. On the other — a the Skottsberg & Skotteberg 613 cited by me in Phytologia 5: is actually V. sulphurea D, Don, as is also the Bridges 5-%+ (Chile) distributed as "ve orinoides Hook," Additional citations: CULTIVATED: Sweden: Collector undesig~ nated s.n. [H. H. Loc. 1837] Say BIPINNATIFIDA Nutt. prem rar es synonymy: Verbenia bipinitifida Walker, ee Additional bibli aphy: Mattoon, Pl. Buyers Guide, ed. 0, 287. Pai else ree ogia 8: 230-212, 381, & 01—L0h (1962) and 8: Be 1963; Gleason & Cronquist, Man. V: & i 3 Moldenke, Résumé Suppl, 6: 3 & 10. 1963; Park's pom seb 39. "1963 ae species is described by Park (1963) as "a showy 15 inch lavenderb h lovely fern-Like trade from York & Chicago), w= Thompece patho grr an rea’ sist-h Pearce (Moorestom, N. J J), and Saier Reeginemvelt sei Mich.)« 1963 Moldenke, Monograph of Verbena 193 additional common name is "fern verbena". Gould describes the corollas as light-violet and found the plant in disturbed black t Val Verde County, aeg | tea nag Edmondson refer to it as "in- quent" on prairies The F. W. Pond 27, distributed as V. bipinnatifida, is actual- ly V. canadensis (L.) Britton; Fleetwood BANA is V. ciliata var longidentata Perry; Hayes 610 is V. pumila a Rydb.; and C. dC, Jackson 18h is V. temuisecta : Brig. ~~ Additional citations: KANSAS: Smith Co.: We H. Horr E.108 (Au- 121735). ARKANSAS: Hempstead Co.: Damaree 963 : (S). OKLAHOMA: Choctaw Co,: Nelson, Nelson, & Goodman 5534 (Au—121733). ai Co: Waterfall & Coryell 9921 (Au-121736) Murray Co.: Muncri 72 (du--121734). TEXAS: Bell Co.: Moldenke & Woods 583 (8)- — We Coy: A. Re Moldenke 663 (S). Comanche Co.: Ae TA 's Moldenke 659 (S); Moldenke i. Woods 59 (8). Coryell Co.: Moldenke & Woods (8). Dallas Cos: Zundell & Lundell 11315 (au—196625, 14). Ellis Co.: leas & Woods 57 (S). Erath Co.: A. R. Moldenke 660 (S)- Falls Co.: Moldenke & Woods 590 (S). Garza “Gos! Moldenke & Woods 60k (8). Hartley Co.: York & Rodgers 336 (au—201 201856). Hays Co-t ae) W. Walker 2 (Au—197613) Hill Co.: Moldenke & Woods 3 568 (S)- Jones Coy: N Ne C. Henderson 63-135 (Au). Kent Co: “YWoldenke & Woods 605 (S). Lampasas Co.: Moldenke & Woods 592 (S)- Lynn 1 Coes Wolderke & Woods 577 (S), 397 (8)» 601 (S), 602 (8). McLennan Coy: Moldenke & Woods 579 (S). Menard Co.: A. Re Moldenke 656 (S). Oldham Go.: York & Rodgers 01 lame yaa “Reagan Co. A. Re Moldenke 653 (S). San Patricio Co.: Gould & Hycka 80L2 (ao 19,231). Schleicher Co.: A. R. Moldenke ae (S)- Shackleford Co.: N.C. Henderson 63-95 (Au—-217645)- Terry ¥ Co.: Moldenke & & Woods $03 (8). Throckmorton Co.: F. W. Gould 9080 (Au—1910L0) « Upton Cos: A. Re Moldenke 652 (S). Val =e Co: B. H. Warnock 11349 (4u—198],57). ARIZONA: Cochise Co.: A. Be os , 783 (S)5 Mol- Genke & Woods 612 (S). MEXICO: C , Gs Marsh 1121 ( (au— 22919). Durango: Powell & Edmondsen 961 Tisai5 32 9 ae BIPINNATIFIDA var. LA’ TA Perry- ee Additional bibliography: anyones Phytologia 8: 361 & 405 (1962) and 8: 62. 1963. _ Additional citations: ARTZONA: Cochise Co.: Moldenke & Wooes $10 (s), 614 (S$), 615 (S). Santa Cruz Co.3 A. BR. Moldenke 622 (8). umztco: Michoaedn: Hinton 13125 (Rf). A BIANCHARDI Samer Bs 3 — 302 & Las 0382) and "8: ee. £2. 19635 Gleeson’ & spaces oe . Vasc, Pl. 19h PHYTOLOGIA Vol. 9, no. § BONARIENSIS L. Additional & emended bibliography: Morong, Britton, & Lew Ann, Ne Y. Acad, Sei. 4 197. 1893; Mattoon, Pl. Buyers G ed. 6, 287. 1958; Nair & Rehman, Bull. Nat. Bot. Gard. bucknor %: 3—~ , fig ; - 1962; Moldenke, Vaughan's Gard, Illustr. 87: 67. 1963; Gebers & Cron Vasc. Pl. 580, 1963; Noldenke, Résumé Suppl dditi papncgerers Nair & a bebaas: Bull. Nat. Bot. Gard. Lucknow 7: 5, fig. 2 ° Nair & Rehman (1962) etl ote are grains of thi cies as follows: "size 3 (range 39—}6 )- Endocolpium lalon- wee (5.6 x 16, sd hs and Cite Nat. Bot. Gard. 24619 from Almora slide 2727), not as yet seen by me. Vaughan han (196 vag? offers this ee to the horticultural trade as seed packet 8566 and describe it as "Tall, hardy verbena. Pe: eseselt Pt. Sweet scented rosy lavender flowers . Ager . year from a sowing « seed indoors and set small plants out when ground is ju Peparph n Curtiss 1963 is a mixture ates ec part [19638] from "sandy ridges bordering the ocean, eastern Florida" is V. maritima Small; the part [1963%*] from "roadsides near Charleston” is Ve bonariensis L.; and the part [1963] from "low open woods, Indian River, June" is Ve S Nash. BA sels (1962) cites his no. 182/he from Cochabamba, Bolivia, ae yet seen by me Dusén 14320 and Gould & Reeves 8227, distributed as Ve pouastensiae are actually V. V. brasiliensis Vell., while 0- sols 70 is vy. kotor rocarpa Perry, Seeds of V. bonariensis are offered to the horticultural trade by Vaughan's (New York & Chicago), Burnett Brothers (New York), Roberts Pibvinetan Kent), Sutton & Sons Thomps rgan » Watkins & Simpson (London), Hone Hone stead Nurseries (Boskoop), Hillier & Sons (Rinchester) > oo Moorestom, N. J,), Saier (Dimondale, Mich.), and John Forbes Additional citations: ALARAMA: Winston Co.: A. R. Moldenke 772 (S). MISSISSIPPI: Holmes Co,: A. Re Moldenke 754 (S)- ai ewe Cos: A. Re Moldenke TuO (S). LOUISIANA: Beaure egard Par.: Ae B Moldenke 709 (S). Bienville Par.: A. R. Moldenke 685 (S)« Hate a Pare: Ae Re Moldenke 698 (S), ARGENTINA: Buenos Aires: O° 2231 (S)- hoa * Files, Norlindh, & Weimarck 2655 (Lu), 2647 (14), 4003 ie. CULTIVATED: England: Stearn 5.1 Sen. oe ees “ze Moldenke 1963 Moldenke, Monograph of Verbena 195 VERBENA BRACTEATA Lag. & Rodr. badttiens) nas ag Verbena brevibracteata Eggert ex Prain, Ind. Kew. Suppl. 3: « 19083 j REE — Mo. Bot. aot: 20: 338, in syn. 1933. Eig ies eatun aged ghingene in her Additional bibliography: Perry, Ann. Mo. : 32. 1933; Moldenke, Phytologia 8: 27 —309, ea es, & zWor—in2 fig. 5 (1962) and 8: h6h. 1963; Moldenke, Résumé Suppl. 6: 1—3 10, 1963; Gleason & Cronquist, Man, Vasc. Pl. 579--581. 1963. Perry (1933) regarded Eggert's V. brevibracteata as a of V. pumila Rydb. because the specimens he so identified are that species. He based his binomial, however, on Gray's V. bract~ eata var, brevibracteata and therefore his name has te go 0 into the same synonymy as Gray's : Cronquist found V. bracteata "on lake shore above present water level but well below highwater mark" in Oregon. Demaree re~ ports it as "common" on low sandy ridges in Arizona. Rowell de- Scribes the corollas as "pale—purple" and tank the plant in -—_ f dunes SLopalay Abuphant 4m cis wipes ON = dog tom". It has also been found in sandy soil, sandy loam, and on rolling prairies. Warnock calls it a "frequent annual prostrate - tinestone depressions"; Wiggins & Wiggins found it and Distichlis "on soil flooded when reservoir is Rowell states that it is "occasional" in sandy loam or tight sandy loam near cattle tanks. The Correll & Johnston 21530 distributed as V. bracteata is paged ep V. gracilis Desf., while Waterfall 12703 is V. subulig- es citations: ALABAMA: Lawrence Co.: A» R. Moldenke 768 (S), ARKANSAS: Desha Co.: : Demaree 45863 (S)- COLORADO: Arch- usta Cos: C. F. Baker 56h (Iu). Boulder Co.: pen, Pl. Exsicc. Gray. 1090 (Au—1966h, Rf). TEXAS: Carson Co.: He N- Moldenke 21883 (S). Culberson Co.: B. H. Warnock 14313 (xu——198201) « 8201). Dal- lan ¢ ma C. M. Rowell 538ha a (du-—195968), SuOla (Au—195986) 5 Turner & Melchert 1,800 (Au--209801); York & Rodgers 165 (Au— 201685), 205 (Au—201 7h), 242 (Au--201831). Hemphill Co.: Ce Me oe ee LosT (Au--195200). Lipscomb Co.: C. S. Wallis 7T2h0 (Au— Ys Iubbock Co.: C. M. Rowell 60-02) I (au--195352) - ). Oldham Co.: H. N, Moldenke 2166) “(s) NEW MEXICO: Catron Cow: Ae Re Moldenke 628 (S). Otero Co.: A. Re Moldenke li (S)- Sierra Coe: ae R. Moldenke 633 (S). Socorro Cos: As Re Woldenke 634 (S). ARIZONA : he Co,: Demaree 11,626 (S)- Coconino Co.: + Denaree 42,7 (5). OREGON: Harney Co.: = Gronguist, 8570 8570 (a (Av—207619, Mi N). CALIFORNIA: i oeenae Goce Maggio “& Wiggins 16338 (Av » so0sh8). VERBENA BRAS Additi. ell « Nace bibliography: Hist. Nat. chile j feidente, ee eagta $1 8; 309-323, 38h, & Misia, fig = bi z. dostg uel a gt in oo 1963; Moldenke, 196 PHYTOLOGIA Vol. 9, no. oes raat 6: 2 & 6. 1963; Gleason & Cronquist, Man. Vasc. Pl. 79 1963. Balls found this species in a cultivated field in California in association with Typha, Juncus, etc. Gould & Reeves found it on "open coastal prairie” in Chambers County, Texas, while ee reports it as "very cammon all along the coast" in Harrison County, Mississippi, The specimen of Lorentz 131 which is represented by Macbride's photograph 20316 is deposited in the Munich herbarium. Additional citations: ALABAMA: Co.: Moldenke & Woods 503 (S). Lawrence Co.: A. R. Moldenke 769 (S). Winston Coe: A. Re i Adams a Moldenke 770 (S); Moldenke & Woods 505 (S). MISSISSIPPI: Cos: A. Rp Moldenke 735 (A). Attala Co.: A. R. Moldenke 755 (S)- Choctaw Co.: A. R, Moldenke 760 (S), Claiborne Co.: Ae R. Molden ke 72 (S); Moldenke & Woods 518 (S). Clay Co.: A. R. Moldenke 76 (S)» Copiah Coy: Moldenke & Woods 519 (S). Harrison Co.: Demaree 29009 (Au—121835). Hinds Co,: Moldenke & Woods 520 (S)- Holmes Co,: A. R. Moldenke 753 (S). Humphreys Co.: A. R. Moldenke 749 (S); Moldenke & Woods 51) (S). Issaquena Co.: A. R. Moldenke 748 (S). Itawamba Co.: Moldenke & Woods 509 (S). Jefferson Coe! Ae R. Moldenke 737 (S); Moldenke & Woods 517 (S). Lee Co. Mol~ Genke & Woods 512 (S). Lowndes Go.: A. Re Moldenke 762 (S)+ Nox bee Co.: A. R. Moldenke 758 (S), Panola Co.: Moldenke & Woods 5a (S). Sharkey Co.: A. R. Moldenke 751 (S); Moldenke & Woods 515 (S). Warren Co.: A. R. Moldenke 747 (S); Moldenke & Woods 522 (S)- Winston Co.: A. R. Moldenke 759 (S). Yazoo Co. Moldenke & Woods ARKANSAS (S)- Beauregard Par.: A, R, Moldenke 706 (S). Bienville Pars: A» R. Moldenke 686 (S), Bossier Par.: A, R. Moldenke 677 (S)5 Molden ke & Woods 550 (S). Caddo Par.: Moldenke & Woods 552 (S)- Cald- eo Pare? Ae R. Moldenke 688 (S), Sabine Pare: A» Re Moldenke na (S). Saint Landry Pars: A. R. Moldenke 717 (S). Tensas Pare? Fo Moods 528 (S). Vernon Par.: A. R. Moldenke 703 (S)- ster Par: A. R. Moldenke 682 (S); Moldenke & Woods 548 (S)- TEXAS: Camp Co.: Moldenke & Woods 562 (S)~ bers Coo: Gould £ Peps cael (au—19],206). CALIFORNIA: Sutter Co.: Balls 11992 - BRAZIL: Parand: Dusén 14320 (N), S80 Paulo: Brade 5737 1963 Moldenke, Monograph of Verbena 197 (Herd, Inst. Bot. SXo Paulo 6729] (N); Campos Novaes s.n. [Herb. Inst. Bot. Sfo Paulo 1977] (N); Campos Porto 36 [Herb. Inst. Bot. Geol. 65h; Herb. Inst. Bot. So Paulo 15728] (N). ARGENTINA: Cér- : Lorentz 131 aaaeean photos 20316] (N—photo). VERBENA Semenig Scope dditional b aie ues » Phytologia 8: 417-20 (1962) and 8: iss 19633 sal pet nam age Ba Suppl. 6: 7. 1963. Moat (190) comments as follows about Chodat's original misidentification of this plant as V. erinoides Lam.: "Le V. laciniata Briq. (=Erinus laciniatus L. Sp. Pl. ed. aie Pe 879, am, 1753 =V. erinoides Lamck. Ill. I, 57, ann. 1791) est une plante originaire du Pérou et au Chili a caractéres trés différ- ents. Voy, A ce thee Briquet Lab. Balans. Parag. p- 9. * . — ons: ARGENTINA: Tucum4n: Venturi 3568 (Du-— cat 1 Moldenke, Plyto : 21 —h25 (1962) at 8: Wes. ee Résumé Suppl. é: 3&kh. sag ound s es south-facing gravel slopes in Coahuila, fruiting in Pict Dat misidentified 1b aes Rydb, King found it growing along gravel r roads in San Luis Poto- =, fruiting in February. se - I. Davis s.n, [Southmost, Spring 1)1] isotype cited by on Caor gia 6: Woe (1962) as deposited at Southern Metho- =a on gree tens 1 Herbarium instead, and one the Iundel & Lundell 8698 specimens cited as in the Univer- sy of Michig: = elle Herbarium is now also in the Lundell Herbarium ue Mo ie li E instead. Additional citations: MEXICO: : Rinehart 298 (S). San inis Potosf: R. M. King 3983 (an--189609), LAD Tj2k9 (Au—16979h) » ted Galeotti 777, in part [Macbride photos 24699] (N-—- CANADENSIS (L.) Britton Additional & syn synonymy: Verbena m —— Hort. ex Sieb. & Voss in Vilmorin, Blumeng., ed. 3, 1: §25, in syn. 1895 [not VY. montana Lillo, coon Verbena canadensis ¢ ta grandiflora Hurst ex Mi Mattoon, Pl. Buyers Guide, ed. 6, 287. ane "Verbena ope compacta Hort. ex Mattoon, Pl. Buyers Guide, ed. 6, 287. 1958. Verbena canadensis rosea Hort. &x Mattoon, Pl. wen Guide, Aaa: : Gen. 2: 198. 18393 tional bibliography: Meisn., Pl. Vaacs Phytologia - Buyers Guide 8: L263 (1962) and 8: h65—hO8, fig. 7- 1963; Moldenke, eg oe tn 3, “os a 11. 1963; Gleason & Cronquist, Man. 198 PerTTOLoG Ii Vol. 9, no. Hardin describes the flowers of this species as blue-lavender. ag found it in "annual-perennial asstciation" along roadsides in sandy loam area recently burned in Texas Seeds of this species are available to the horticultural trade from Hav'a'look Gardens, Ridenour, Hillcrest Nursery, Thompson & Morgan (Eng rican Perennial G en City, Mich), air Nurseries (Nichols, N. Y.), J» Herbert Alexander ( ae boro, Mass.), an J (Syosset, Ne ese the "com- ta" form from Pearce (Moorestown, N. J,), Saier (Dimondale, Mich.), John Swain (Bristol), and Norsk Fro (Oslo); the "compacta grandiflora" form from oe ay Son Rice and the "rosea" form from Ridenous, Joseph F and Tucker & Son (Belleville) . The Leavitt s.n. [Caubridse, "13 Oct. 1898] di distributed as V. canadensis is ac actually V. elegans H.B.K., while Ammons, Straus straus- baugh, & Bartholomew s.n. [Presque Isle, July 29, , 1940) is is Ve stricta | Vent. "Additional citations: FLORIDA: Leon Co.: Re Ke r 5uls32 (S)- MISSOURI: Jackson Co.: K. K. Mackenzie 62 (Mi). TEXAS: Mad- ison Co.: F. W. Pond 27 ae Robertson Co.: Hardin 547 (an-—191113) « Tod LOCALITY OF COLLECTION UNDETERMINED: Collector un designated s.n. (Lu). VERBENA CANESCENS H.B.K. 19630 bibliography: Moldenke, Phytologia 8: 1,69--li75- 8 species has been found in sandy clay soil on flats and on south-facing limestone-gravel slopes in Mexico. The C. Te Mohr s.n. [Hunter's Wharf, Sept. 5, 1892] eiateh bated as is Ve r, can escens is is actually Vv. 8s supina L tional citations: TEX TEXAS: Jeff Davis Cos: A. Re Moldenke G45 (S)- MEXICO: Aguas Calientes: Hartweg 177 (Lu). coakuilat Rinehart 221 (S), 29 (S). CANESCENS var. ROEMERIANA (Scheele) Perry Additi tional bibliogrep Moldenke, Phytologia 8: 475--l82- 19634 Moldenke, Résuné ce ies © dis 8 ~~ ‘Additional citations: TEXAS: op oe Co.: A. R. Moldenke 66) (S). Coryell Co.: Mo Moldenke & Woods 587 (S). Menard Co.: As Re Moldenke 657 (S). Schleicher Coe: | A. Re Moldenke 655 (S)- wr pin blicgraphy: Perry . Bot, Gard. 20: 356- 1933) Seiten Phytolo : Suppl. 6: 11, 1963, = 8: LBS 183-195. . 19635 Moldenke, Résumé The 0. M. Clarke 7348 (N) cited in Phytologia 8: 49h (1963) is actually ¥ vy. fe Fecta H,B H.B.Ke The He Je Hamby 187 distributed 3s 1963 Moldenke, Monograph of Verbena 199 V. carolina is actually V. halei Small, 343 is V. neomexicana var. hirtella Pe: Perry, and 5h5 is ve V. plicata Greene, while A. A. Eaton 196 is V. scabra Vahl. Additional citations: ARIZONA: Cochise Co.: A. R. Moldenke 781 (S). MEXICO: Chiapas: Matuda 5295 (1d). Guerrero: Hinton 9287 (Rf). Hidalgo: Lundell & Lundell 12391 (Rf). México: C. L. Lundell 12209 (Rf); Lundell & | & Lundell dell 12355 (Rf). Michoacén: Hinton 13022 (Rf). State undetermined: Collector undesignated s.n. [Chico ma ee eee VERBENA CAROLINA f, ALBIFLORA Moldenke Additional bibliography: MASE py elees 8: h95—h96. 1963; Moldenke, Résumé Suppl. 6: h. 1 HEITMANIANA Moldenke Additional Mb Liogrenurt Moldenke, Phytologia 9: 9—-ll. te Material of this species has been misidentified and distri ted under the name V. santtapanetl Brttcher. It has been aan ed i ee in January. mal citations: ARGENTINA: Catamarca: Brttcher & Brticher an, “GOA/iS) (Iu). VERBENA CHILENSIS Moldenk Additional eibaacececie: Moldenke, Phytologia 9: 11--12. 1963. The correct bibliographic citation for this name is "Moldenke, Known Geogr. Distrib. Verbenac., [ed. EF 78. 192". plo (Ma) citations: ARGENTINA: Mendoza: H. He Bartlett VERBENA CILIATA Benth. ep tional synonymy: Verbena — Smith, 1 dditional bibliography: Hy: Moldenke 963, Moldenke, Résuné Suppl. 6: 3, by ee - King f Reeded to determine if more than one taxa are here repres Specimens of V. uiiate have been misidentified and distrib- as -V. pumila Rydb. The H.C. Hanson 304, distributed as V. ciliata, is actually var. longidentata Perry, while Hayes 610 is V- ;V. pumila Rydb. ier araraeeeeneenneeeaeed Additional citations: TEXAS: Cameron Coe: “Fleetwood 3711 (an- 200 PHYTOLOGIA Vol. 9, no. 192387)» Reeves Co.: A. R. Moldenke 61), (S), NEW MEKICO: Sierra Co.t A. R. Moldenke 632 (S). ARIZONA: Coconino Co.: Demaree 4615h_ (S), b6155 (S), W6191 (S). MEXICO: Chiapas: Matuda 5863 (Id). Co ahuila; Rinehart 217_ 217 (S). Guanajuato: Hartweg 176 z 176 (Lu—isotype). er & Hernandez Xolocotzi 276 1, oe J. Alexander 2287] (N), 277 [B. J. Alexander 2288) (N). México: Hinton 869 (N, NW), 2893 (K, N, Rf, W—1822199), 7466 (K, N, N, N)3 Leeds Bone near Amecameca, March 1h, 1929] (Id); C. L. Lundell 12210 . (Rf). Michoac&én: Iltis, Koeppen, & Iltis 50a . (Ws); Re F. Smith mith M.97 (Au--218258), Nuevo Leén: 3 F, Smith th Moh6k (Au--218066) « Oaxaca: W. He 2581 (Mi, N)3 Conzatti & Camp 5237 (N). Puebla: R. Me King 2627 (Mi). Vera Cruz: C, L. Lundell 12535 (Rf). TA var. LONG GIDENTATA Pe Additional bane A Molde ged , Pavtoloeta 9: 22—26. 19635 oeee cita : : ron Co.: Fleetwood 3) (Au 192006); H. C. Hanson 304 (N), 322, in part (N). VERBENA C - PUBERA (Greene) P CILIATA var Additional bibliography: Moldenke, Phytologia 9: 26-27 19633 Moldenke, Résumé Suppl. 6: ae. & 10. 1963. Additional onal citations; : Lubbock Co.: Demaree 7460 (Au— VERBENA CLAVATA Rufz & Pav, Additional bibliography: Acevedo de Vargas, Bol. Mus. Nac. Hist. Nat, Chile 25; es 19514 Pan Phytologia 9: 29--32- 1963. Additional citations: PERU: Hudmco: Ruiz son, [Macbride photos 17408] (N—-photo of cotype). VERBENA VERBENA CLOVERAE maidens Additional Verbena clovei Smith, 963 Additional bibliography? Moldenke, | piytcloeia vi : 331 19075 Moldenke, Résumé Suppl. 6: 3 & 10, 196 Smith refers to this as a = a herb Additional seinidaae, MEXICO: Nuevo Leén: ‘Re F. Smith M02 Me62 (Au—218096) . i 1963 Moldenke, Monograph of Verbena 201 VERBENA _SLOVERAR f. ALBA Lundell, Wrightia 2: 7 [as "cloveri"]. 1960. Bibliography: C. L. Lundell, Wrightia 2: 47. 1960; Moldenke, Résumé Suppl. 6: 3. 1963. This form differs fran the typical form of the species in having white corollas. The type of the form was collected by Cyrus Longworth Lundell and Amelia A. Lundell (no. 15055) in open brush off U. S. Highway e type specimen. Citations: TEXAS: Zapata Co.: Lundell & Lundell 15055 (Rf— VERBENA COCCINEA Raf., Herb, Raf, 50, nom, nud, 1833. Bibliography: Raf., Herb. Raf. 50. 1833; E. D. Merr., Ind. Raf, 205 & 295. 199, I know nothing about this plant except that Merrill (1919) lists it as having been named and published by Rafinesqe, with the type specimen from "Oregon or the Rocky Mountains". VERBENA COCHABAMBENSIS Moldenke Additional synonymy: Verbena myriofila Maria, Pl. Vall. Cochab. 38, hyponym. 1962. Verbena myriophylla Maria, in herb. Additional bibliography: Maria, Pl. Vall. Cochab. 38. 1962. Maria (1962) says for his V, myriofila "Hierba icada desdo la base, florece en cimas de color lila brillante: es un verdadero adorno que alegra el campo" and cites Hermano Adolfo Maria 180/he from Tarani, Cochabamba, Bolivia, collected on De- cember 16, 195). I have examined a specimen of his no. 180 and found it to be typical V. cochabambensis. Additional citations: BOLIVIA: Cochabamba: A. Maria 180 (2). Additional : : a evedo de V: Bol. Mus, Nac bibliography: Ac e Vargas, ° + ° me Nat. Chile 25: 70. 19513 Moldenke, Phytalogia 9: 39—h0. VERBENA CORYMBOSA Rufz & Pav. ered attoon thi to the horticultural trade by Thompson & Morgan (England), Hill~ ier & Sons (Winchester), and John Forbes (Hawick). VERBENA COVASII Moldenke Additional bibliography: Schnack & Covas, Revist. argent. Agronon, 12: 22h. 19h5. VERBENA CRITHMIFOLIA Gill. & Hook. 202 PHYTOLOGIA Vol. 9, no. k Additional open ab Moldenke, Phytologia 9: 6—h9. 1963; Moldenke, Résumé Suppl. 6: 9 & 10. § The Brttcher & Brttcher s.n. (28/12/48) distributed as V. crith- mifolia is actually V. hookeriana ironed & Schnack) Mol ldenke. Additional citations: ARGENTINA: La Pampa: Burkart s.n. [Herb. ae ghets Darwinion 20575] (W—230690). Rio Negro: We ij. Fischer 90 (N CUMINGII Mo. Additional bibliography: Acevedo de Vargas, Bel. Nace Hist. Nat. Chile 25: 70. 1951; Moldenke, ee 9: 9: 19506 1963; Moldenke, Résumé Suppl. 6: 10. 1963 VERBENA CUNEIFOLIA Rufz & Pav. Additional bibli phy: Acevedo de Vargas, Bol. Mus. Nac ogra: =e Nat. Chile 25: 70. 1951; Moldenke, Phytologia 9: 50-52. 9636 Additional sips: PERU: La Libertad: Lépez Miranda & Sag&stegui 3543 DEAMTI Molden 1 Additional aati ceve thy Gleason & Cronquist, Man, Vasc. Ple 581. 19633 Moldenke, Phytologia 9: 53--Sh & 59. 1963. VERBENA DELTICOLA Small , Additional bibliography: Moldenke, Phytologia 9: 60—6h. 19633 oes Résumé Suppl. 6: . 1963. ss exican Bound. Surv. 8.n. [Plains near Leon Springs, or perhaps was a mixture with that species. Fleetwood 6079 bears a notation by the collector "other plants had more dissected oe leaves = sesters pre rites of V. delticola has also been distri gert. Additional ottetisees TEXAS: Bexar Co.: Herb. Mexican ae: a oa ee near Leon Springs, September 7, 1052) (N as a tata onal bibliography: 1 ocking, Dict. Terms Pharmacog. 26h. 19365 moyas » Phytologia 9: 65-66. 1963. & emended bibliography: Worong, Britton, & Vail, Amn. Ne T. Acad, Sci. Ts on tis actaee & Covas , Revist poste Sse Str enas otis 5 cen oe ten Bl as » Nat. : 9513 Moldenke, Phytologia 9: 67-—/- 19635 Me aldecke, Résumé Suppl. 6: 7 & 8, 1963. en Weiete cee Se ol 1963 Moldenke, Monograph of Verbena 203 1946], distributed as V. dissecta, are actually V. tenera Spr Additional citations: TROGENTINA: Tucumén: Re Rocha ha 3018 8 Poe VERBENA DOMINGENSIS Additional bibliography: Moldenke, Phytologia 9: 76—79. 19635 Moldenke, Résumé Suppl. 6: 8 & 10. 1963. USENII Moldenke Additional bibliography: Moldenke, gape 9: 79—80. 1963. Additional citations: BRAZIL: Paran4: én 7108 (Lu--iso otype)- VERBENA ELEGANS H.B.K. Additional bibliography: Moldenke, Phytologia 9: 82—86. 19633 Moldenke, Résumé sing Bad 6: 8. 19 The R. M. King 513 cited in Phytologia 9: 85 ee as a elegans appears to me better regarded as var. asperata ~~ Additional citations: MEXICO: Coahuila: E, G. Marsh ed (au— 213647), 1258 (Au—2130h)). VERBENA ELEGANS var. ASPERATA Per Additional bibliography: acini Phytologia 9: 86—89. 19633 Moldenke, Résumé Suppl. 6: 8. 1963. Soderstrom states that the corollas of this plant are pink, "turning blue with age", and that the plant is "frequent on rocky Clay sides of arroyo". King 513, previously regarded by me as typical V. elegans H.B.Ke, sage said by the collector to have had oe flowers and to have been abundant in in gravelly soil, Open gs Additional citations: MEXICO: Chihualma: Soderstrom Oh/ 847 (Mi). Tamaulipas: R. M. King 4513 (an-—1899h0, N). LOCALITY OF COLLEC- TION UNDETERMINED: “Gizantoro sen. [9.66] (Lu). XVERBENA ENGEIMANNII Moldenke Addi tional peng AE ena hastata var. p oblong toss Nutt., Gen. 2: ho. 1818. SS a hastata y oblongifo Class-Book, ed. 1, 412. ~ hyl2. 1856. Verbena hastata var. oblongifolia Nutt. apud Small & "Heller, Mem. Torrey Bot. Club 3: 12, in syn. Additional biblio Nutt., Gen. 2: 0. 1818; Wood, Class- Book, ed. 1, 12, ieee, Beall & Heller, Mem. Torr rey Pet = oe 3: a 1892; Perry Ann. Mo. Bote Gard. 203 2676 1933; Mo Phytologia 9s 89—96. 1963; Moldenke, Ri Résumé Suppl. 6 ee w 13835 st, Man. Vasc. Pl. 580. 19633 Ba .8.1.C. ( B.N2. 1963. Small & Heller (1892) an and ey (1933) B ees cere, s name in synonymy V. riparia Raf ttall and himself represents a hybrid between V. hastata L. and Ve urticifolia oa His description of the plant is: "h. hastata. Flowers — 20h, PUTTOLGG Ti: Vol. 9, no. i rarely if ~_ hastate. Of this species there appears to be a very distinct variety near Philadelphia which I shall distinguish ag the name of oe *oblongifolia, aint otlLome-lenonalate deeply serrated mph erely acute, not acuminated; spikes fili- form, panicula 5 Flowers vamaiter, pale blue. May ‘ends be V. pam iculata of rig he flowers are not imbricated, nor r in {usa of the same, but assuredly intermediate, if not a hybrid be- twixt V. hastata and V. urticifolia. It has only occurred to me twice on the banks of the Delaware." It seams obvious to me ated this cannot apply to the V. oe Raf. as published by and tyP: haat tonal citations: MISSOURI: Saint Louis: Engelmann s.n- (St. Louis, Aug. 183}. (Au—12280)) . VERBENA FILICAULIS Sota Additional & em ibliography: Hook. f. & Jacks., Ind. rang 2: 11:78 & 7. 1895; Moldenke, Phytologia 9: 119121. Additional citations: BRAZIL: Minas Gerais: Widgren 5.De (1845] (Lu). Parané: Jonsson 1392a (Lu). GOODDINGII Ge Additional bibliography: Moldenke, Phytologia 9: 131—1h1, fig. 8. 1963, a eim reports finding this plant in rocky soil on top of Additional citations: gre es : Langenheim 3728 (Av— 209322). NEW MEXICO: Luna Co.: Soh, oldenks 631 (S)- ARIZONA: Cow: A. R, denice 62h (S), 626 (S). Pima Co.: A. R. Mol- denke 625 (8). Coe: A. Ry Moldenke 621 (S)~ CULTIVATED Calif : Balls 39 19756 (au—122111) VERBENA GOODDINGII var, NEPETIFOLIA Tidestr. < bibliography: Moldenke, Phytologia 9: 11—16- dditional citati ; 11775 (A ie ¢ ons: MEXICO: Sonora: I. L. Wiggins li/i> VERBENA ended arose SChem.) Herter Verbena gracilescens Cham. ex Hicken, Chlor. ju a PB ine 197. a 1963 toma bibliography: Moldenke, Phytologia 9: 148—15h. A@ASlonal citationas BOLIVIA: Cochabanba: A. Maris 178 (Z)- 1963 Moldenke, Monograph of Verbena 205 VERBENA G D oe SetastnenintG s Moldenke, Phytologia 9: 154-158. 3 Additional citations: ARIZONA: Cochise Co.: A. R. Moldenke 617 (S). MEXICO: Aguas Calientes: Hartweg 17h (Lu). Federal District: Berlandier 578 (Lu). VERBENA HALEI Small : eae bibliography: Moldenke, Phytologia 9: 160—175, fig & Hycka encountered this plant in open oak-mesquite in Texas Additional pttatiines ALABAMA: Cullman Co.: Moldenke & Woods 50h, (S). MISSISSIPPI: Jefferson Co.: A. R. Moldenke enke 730 (s). Warre Memes Co.: Moldenke & Woods 525 (S, Z, Z). Yazoo Co.: Moldenke & Woods 52), (S). LOUISIANA: Avoyelles Par.: A. R. Moldenke 722 (S)« Bisnville Par.: A. R. Moldenke 687 (S); Moldenke & , & Woods (S). Bossier Par.: A. R. Moldenke 679 (S); Moldenke & Woods 549 (S). Caddo Par.: Moldenke & Woods 551 (S). Caldwell Par.: Moldenke & Para: rag R. eo ldeniin 693 (S). Franklin Pars: Moldenke & “Foods 530 (S), 56 (S). Jackson Par.: Moldenke & Woods 532 (8). Lincoln Par: “Moldenke & Woods 538 (5). Natchitoches Par.: A. R. Moldenke $97 (S). Pointe Coupee Par.: A. R. Moldenke 720 (S). Red River Pars Ae R. Moldenke 690 (S). Tensas Par.: Moldenke & Woods 527 (S). Webster Par.: A. R. Moldenke 681 (S); Moldenke & Woods 547 8). TEXAS: Anderson Co.: Hardin 531 (Au--19111h). Bell Co.: ; Mol- denke & Woods 586 (S). Camp Co.: Moldenke & Woods 560 (S)- Camer— on Co Co.: Fleetwood 3508a (Au—-192305). Cass Co.: Moldenke & Woods 558 (S). Comanche Co.: A. R. Moldenke 658 (S); Moldenke & Woods — 595 (S). Harrison Co.: A. Re Moldenke 675 (S). re Cos: Moldenke & Woods 567 (S). Hopkins Co.: Moldenke & Woods 565 (S)» Lym Cost Moldenke & | & Woods 598 (S). Marion Co. on Co.: Moldenke & & Woods 553 (S)- Morris ©. Cos: Moldenke & Woods 557 (S). San San Patricio Co.: Gould & — 790 (Au~—19],66). Upshur Co.: A. R. Moldenke 673 i ee Pag 2 A. Re Moldenke 672 (S). Wood Co.: A. R. Moldenke 670 (8)5 Wol denis & Woods 563 (S) — ta Farwell ex Moldenke, Prelim. Alph. List I ae 46, < syn, 190. Verbena hastata var. pinna Holdenke, Prelim. Alph. List Invalid Names ng in Ce 1940. oe Dena hastata Michx. ex Moldenke, Suppl. 206 PHYTOLOGIA Vol, 9, no. & syn. 1941. Verbena hastata var. pinnatifida (Lam.) Britton ex Moldenke, Suppl. List Invalid Names 8, in syn. 191. Verbena lac- unosa Schrad, ex Moldenke, Alph. List Invalid Names Suppl. 1: 25, in syn. 1947 [not V. lacunosa Mart., 1941]. Verbena lanceolata Boldo ex Moldenke, Alph. List Invalid Names Suppl. 1: 25, in syn. 1947 [not V. lanceolata Willd., 1825]. Verbena paniculata Schau. House ex Moldenke, Résumé 366, in syn. 1959, Verbena hastata var- pinnatifida Britton ex Moldenke, Résumé 366, in syn. 1959. Verbena astrata L., in herb. Verbena hastata var. pinnatifidum (Lam.) Sch. ex Moldenke, Résumé Suppl. 3: 38, in syn. 1962. Liatris eas Nash ex Moldenke, Résumé Suppl. 3: 33, in syn. 1902 Var- esina biserrata Nash ex Moldenke, Résumé Suppl. 3: 42, in syn- 1962. Verbena haetata L. ex Moldenke, Résumé Suppl. 3: 38, in syn. 1962. a ography: P, Herm., Hort. Acad. Lugd.-Bat. Cat. 699. 16873 P. Herm., Fl. Lugd.-Bat. 5-55, 1690; P. Herm., Parad. Bote, ede 1, 242. 1698; Ray, Hist. Plant. *. e . - en”: 412. 1856; a. W. i bok os d. 1, pre i por (2860), "pr. 2, 307 (1865), and pr. 3, 307. 1372; an. Ar. 167lis ¢ 197k; A. Gray, Syn. Fl. N. Am. 2 (1): 336. 1878; Pickering, 1963 Moldenke, Monograph of Verbena 207 Chronolog. Hist. Pl. 949, 968, & 1220. 1879; W. Mill., Dict. Eng. Names Pl. 1] & 261. 1884; A. W. Chapm., Fl. South. U. S., ed. 2, pr. i, i) (1884) and pr. 2, 307. 1887; Rattan, Pop calif. ve ed. 8, li. 1888; 3. W. Chapm., Fl. South. U. Se, wi 2, Pre 3, “307. 1889; A. S. Hitche., Trans. St. Louis Acad. Sci. 512. 1891; Coult., Contrib. U. S. Nat. Herb. 2: 327. 1892; A. . «, Fl. South. U. S., ed. 2, pr. kh, 307. 18923 Know Wild Fls., ed. 1, 252, 291, & 297 (1893) and ed. 7, 252, “201, & 297, pl. 92. 1893 Briq. in Engl. & Prantl, Nat. Pflanzenfam. Clu F Gard. Bot. 31 Rydb., Contrib. U. S. Nat. Herb. My e (1895) and 3: 518. = W. Chapm., Fl. Sou: ue U. fan 8. eA. ge n& i 11, Ann. Rep. Comm. wor, Contrib. U. S. Nate Herb. 6: 3. 1901; H. H. W. Pearson in , co Herb. 7: 383. 1902; , Fl. 3 z : 903 L. He » owa Agric. Sta. 70: 513. 1903; Selby & Hicks, Ohio Agric. ? pl. 4. 1903; Michx., Trav. West = te, 67. 190k; Selby & Hicks, Bull. Ohio Agric. Sta. 175: pl. k. 1906; U. S. Dept. Agr tot Bull. 260: 22. l' epi ag s ae ° "Ue S. Nat. Herb. li: ro "49 MacDoug: Ove * st. 1907; Chadwic - N.Y. Sta na: 12h: 150. 1908; Robin- Mi Wild Fils. owa Bull, k: 263 & 68. 1913; Britton & Br., I Hiinstr Fe, ed. 2, 33 9h, 95, 599, 618, & t 635, es 366. 1913} Small, Fl. Southeast : S., ed. 2 Bendrat, Torreya 13: 36 vell ower Kanda, Bot. Gaz. 69: Sh--T1, fig Su—59, 63, 75, & Ple 5 fe 1920; ; Pellett, De ae at x gE 266. 19205 Fernald, Rhodora 208 PHYTOLOGIA Vol. 9, no. k be e e ne a By th Moldenke, Obs. Fl. peony a. ee z (1933) and ed. 2, we 1933; Junell, a nee Upsal. i: 11, 12, 1h, 19, 16h, 171, & al s ico-c Sa Bull. Hortus, new rev. ed., 632, 1 E. J. Palmer, Amn. poe Bot. Wade 02 6a5. 1935; H. T. st ; = en ihe ae Sci. 22: 62. 1936; acne, Cytologia 7: i153, fig. Os: gtr re ch. Acad, Sci. 22: 88. 1937; *Bausor, T 51. 1937; Cory, Texas ; ta. B ; *550: 88. 19373 Mol- e Sudam. Bot. h: 18—20. 1937; Svensk. Bot 2 Camp mss, 190; Worsdell, Suppl, 2: 4,86. = las Moldenke, Suppl. List 4 st Invalid Names 8. 191; ag Becees . be, Pcie - 219. 19h; C. N. Jones, Ohio Journe tea, 1, 340. 19h 191; , Known . Distrib. Verbenac rates devatia'tmos 7 15 a “Thy & ie shes Moldenke, Alph. os 1 Fi, temas 3 ()i 16 & 2h— . 19h2; Pim, Leonard, cH ): - Wise. ae a. is. “ls ed. 5 nas netanes ab: 47. 1943; Suears College Dict-, Jepson Fl. 19h35 C. N. Jones, Ohio Journ. Sci. 43: 191. a eps: > -C * 3 ( 2): 380—381, figs his. 1943; Ce W. Lowe 1963 Moldenke, Monograph of Verbena 209 List Flow. Pl. Man. 81. ae E. L. Braun, Amnot. Sperm. 116. 193; danemei a ea: - Appal. Exp. Watershed rot & 56. 19h3) Moldenke, Known G eran Verbenac. Suppl. 1: 1 & 2. 1943; E. L. D. seucrrathypse evs - Encycl. eit _9ts Moldenke, Obs. Fl, Warren 27. 15s etienis! Obs. Fl. W Suppl. 1: 11. 19hby Rouleau, Inst. Bot. Univ. Montréal “contrsb. "Eht i, 225, & 36h. 194k; Schnack & at 4 Darwiniana A 9bh; Moldenke, Cas tan- ea 9: 69 (194) and 10: 35--39. ee Kriebel, Yuncker, & Friesner, Proc. Ind. Acad. Sci. 5h: 95. rraghtys Seed Trade Buyers Guide 86. 1945; Darlington & Janaki Ammal, Chromosome Atl. 270. 195; H. E. Stone, Fl. Chester Co. 112h—1126. 19h53 —* 1945; Moldenke Bot. 610. 1945; Ce Ne Jones, ce) he rg Sci. ange 165. 1945; G. N- Jones, Fl. Ill. [An. e Nat. Monog. 2:] 216. 19453 Moldenke, Phytologia 2: 68, sag irs she, & 15, 19Ls5 Moldenke, Am. Midl. Nat. 35: 377. 19h6; G. L. Fish Am, Bot, Exchange List. 196; Moldenke, Alph. mst tk 1: & es td 109-112, 115, lig, 1 | 129-2131, 139-—1h5, 148 i 160, es, teary 131-188" 191, 19h, 195, 197--199, 201, 202, 20-2077 209, 2112-215, 217, 22h--226, 23h, 235, 239, 2h2—2h8, 250, 254—256, 258, 260, 265, 267, 268, 2m: 271, 7k 278-285, 287, & 291-297. 19465 Pellett, in. 8 Pl., ed. h, 420. 19h75 Hotchkiss & Stewart, Am. Midl. * 38: 35. ih 1947; Tatnall, Fl. Del, 218. 1975 Deam, crs peat asi Proc. Ind. Acad. Sci. 56: 110, 1947; Moldenke , PiytoLogta 7 £ 38—321, 329, ges 382, & 383, 197; woldenke, Alph. Names og nt —26. 1947; Moldenke, Wrightia 1: Meh 1948; 7 molaknkel astanea 13: o—~113, 1 9183 Ogden, dame & Hyland, Bull. gears Bot. Soc. 8: 55. 19)8; & Friesner, Proc. Ind. Sek. 57: a (oid) and and ee 1 95. = ola John W. Harshb Bowman's st man, Begin. Guid Pella Sint 75. 1948 prep dis , - Guide Wild Fls. 03 “k bn Mo Cit. 2:°366, 357, 361, 362, 390—L02, bOh—hO7, 10, fal 439, 4h3, bio, be1—b26, 467, h69—b73, 475--478, 480, ub, 4B8, 535, 7s 9° S71, 572, 536, 538, Shb—Sh8, 55s $56, 958-560, 561365 g0, 312s 57h, 576, 2 4-598 OSs) g 519-581, 583-585, 288, tae 6 a i ap ate (13h8), a6 811. 8, 31, 832, 839, 8i1-—-8h3, 851, 852, sl, ae8 » 869, 870, 881, 88h, 887, 892, 896, 90h, 906, 9 15-917 : 92, 974 22t 9325 939 a, 92, 96, 97, Orme 755, 963, 97 970, 9TLy » & 977 (1949), and 97983 985, 998, 999, Soe 1010, 1013-1015, 108k, 1085, up Pale et si! 1138, 1242? 11) nt its, 16, we b 1155, 1262, ’ 16h, ams "e 11h47-- “e ’ ’ 9. 1165, 1169, 1172-1175, 1177, 1178, , 118h—1189 210 PHYTOLOGIA Vol. 9, no. 1191--1193, 1197, 1199-1201, 1203, 1205, 1207, 1209-=124hs 1217, 1218, 1220--1222" 122-1232, 1235--1239, 12h1; 12h6—12h8, 1251- 1253, 1256, 1260° 1261, 1288-~129 2, 129, & 1297--1299 . wish. Be L. Palmer, Handb. Se "Hist. rid & 663. 199; Heusser, Bull. rey Bot. ob». Ginb 76: 393, 1949; Loew, Ann. Rep. Huntington oh Gard. 13: 37. 19h9; H. N. & A. L. Moldenke, Anal. Inst. Biol. » Oregon Bartholomew's Cobble. [pe 3). 1949; Moldenke, Kn Geogr. Distrib. Verbenac., [ed. 2], 2—-8, 10, 12-17, 19-21, 2h,--27, 107, 108, Me & 198. ae pad zs Clausen, Cornell Univ. Agr i Phytol by 29s, . 37h. 1950; Fern. in A. Gray, Man. Bot., O, & *1612 sme o FH 6h. 1950; Zim & Martin, Flowers 60. oe Tatoots: Pl. Disease Handb. 657. 1950; M. S. keith, Kodachr. Ser. Wild’ Fis 8. Ward's Nat. Sci. Estab. LX68-83 & Lx65-8h. 1805 M. T. Grant, + Iowa bnads 66 Sci. 57: a 1950; mere gale” nat Be “ip. 1950; Yuncker, & Friesn Chittenden » Roy. Hort. Bio Dict. Gard. : 2209 & 2211. 1951; Gottscho, Wildfls. 55 & 192. 1951; Moldenke, Phytologia 3: 450 & 467 (1951) and : 67. 1952; H. Ne & A. Le Moldenke, Pl. Bible 5 & 326. 1952; Molderke in Gleason, New Britton & Br. Illustr. Fl., pri 2 fe Proc. Iowa Acad. Sci. 62: 208. 1955; » Dict. 7 og. 7h, 110, , 209, & 2h0. joie eaaatian Nature “ matin? esculentus, and Lythrum and in * , Iowa drift sheet, assoc~ iated with Carex vulpinoidea, Verbena urticifolia, Epilobium col- oratum, Posyecn acre, Aln and Aster novae-angliae” "a in low ground, etc., associated with er 3 Silphium rarer! , Rudbeckia laciniata, Salix longifolia, and Solidago serotina", and Pammel & Spiker found it in "black soil, low ground in pas pastures, associated with Vernonia fascia Poa son roadsides of common in fields", Tosh reports it as "common in all low wet land" in Ra~ leigh Count, West Virginia. In Iowa it was found by Pammel & Zimmerman "in low ground with Convolvulus sepium and Helianthus presierratea; in Oklahoma it was found by Hopkins "in moist damp copses among Salix"; in California it grows with S$ tachys and me according to Nobs & — and in Kansas it is said to be Common in wet places in prairies . Small describes it 8s found "in sand and clay soil atéae roadsides through salt marshes" in New Jersey, while Maguire reports that in Utah it is ste canal t Places", In a prtpebs it was red by Hamerstrom in sandy loam be Soil with Asclepias syriaca incarnata, Lycopus americanus, Phleum oe page sevaieh hh describes it as "abundant at edge of swam G. J eae os fl from June to September e Johnson 1926 says that it flowers and is to be _(2928) aye that 38 £2 + the United States and Cana~ to its FLOWERS. For color and appearance in natural surroundings, 218 PHYTOLOGIA Vol. 9, no. usually seen clinging to the blooming spikes, and ap nieve on then.# - is also stated there that’ "John eames $T- to s . averse arte word for any weed, speaks of its SESE pa knotted threads, that 'make a pretty etching up- on the winter snow.'" The cabs Es here is to the fruiting spikes, which, however, are not "drooping", but firmly erect. was * probably referring to the spikes of V. urticifolia rather than ‘anne of V. hastata. There is no mention in Stapf's "Index Londinensis" of any color plate ee a Rinc 1766 Palmer (1935) states that in Missouri V. hastata inhabits “low open woods, wet prairies, fields, and eae Soaks circum- neutral. General, but commonest in northern and central" portions of the state, - a his 1916 publication he cites his nos. 189, 1085, 1086, and 2483 from Jasper County, not yet seen by me ae 1930 publication he cites his no. 23505 and C. Ke ines —— the species from Hancock County, “Iowa, at cite theiz a 1568; Loew (1949) records it from Huntington C and says it is "medicinal"; and C, N. Jones (a9li1) cites Re wal- ker s.n. from Hocking County, W. Goslin s.n, from Fairfield County, and L. E. Yarman s.n. from Knox County, Ohio. In his 19h3 work he cites | C. 1 H. Jones 8.n. from Marion County and in in his 19l5 erbari 16. Jepson (1913) records V. hastata fron pean beac County, Califor nia; E. L. Braun (1943) records it fron Boone and Morgan Counties, Kentue Dean, Kriebel, Yuncker, & Friesner (1915) age ° 3 st ranklin, ec, Knox 2 Somerset Counties, Maine. Jepson states that t species was goblestes a in the Grand Island, Lower Sacramento, Te~ . Leonaz 1 181 the Leempr: an W: (1943) ree Focdrd it pe Ae Bull Run ui 5 — a illiam County?]. Iowa Acad, ‘Sei. 6g: sa: it 1 (955) it is maid to: be cman Gn Geaccan County, It - 1963 Moldenke, Monograph of Verbena 219 Christchurch, Sou by Druce in R . Exch. Club Brit. Isles wes, 1208 Gems. ‘oansata (1922) Aas Dale seen west of Hants County" in Nova Sc = S. Steele 98, eT in "Camden County, New Jersey, misleading | printed label reading "Plamts of Pennsylvania"; Je Torrey 125, sotinctel « at Hoboken, New Jersey, bears a label reading "Hoboken, N. Y." and "Lect. circa Hoboken} in Canada." The "East Caledonia" Tiger Vermont, recorded for this species by me The Block Island, "“assachusetts", recorded on page 6 of the same work is an error for Block Island, Rhode omraere e a the "Girard" inne to be an er ror of transcription. There is no such county in t the pte nor onl nents a town by that name there, nor a county by that name ere, The species is used locally in medicine in Florida, according to Curtiss, while in Michigan the leaves are "used to make ver- vain tea, an old-fashioned spring tonic". H. N. Webster (192) says that it is "an emetic herb used from earliest times in Col- on 0 late perspiration, while "large doses are emetic". The Black 36-5881 36-5881 from Ohio looks somewhat like xV. engel- mannii aeageriic — as to the difference between V. hastata ata and x¥. e my discussion of the latter in these notes. The etiam 3356, distributed as Y, hastata, is actually Veroniea” longifolia L., while C. M. Long s.n. [Owe Sound, Ont., 1873], also so distributed, is Veronicastrum virginicum (L.) Far- well, both in the Scroplmlariaceas. Verbena hastata apparently hybridizes freely with related species — — its hybrid with V, carolina L. is xV. matritensis Mol- denke, with V, hispida Rufz & Pav. is xV. noacki Moldenke, with vy. oftictoslis Tae e L. is x¥. baileyana Moldenke, with V. simplex Lehm. is xV. blanchardi Moldenke, with V. stricta Vent. is xv. Tydbergii Moldenke, and with V. urticifolia L. is xv. engelmann- ii Moldenke, MacDougal (1907) “suggests that it also . hybridizes with V. bracteata Lag. & Rodr. in the wild. However, he cites no heron, gives no description of the ——. hybrid, and does y that he actually ever saw any such hybrid plants. —, (lish) ree arded V. pinnatifida Lam. as a hybrid between V. has- se cat V. officinalis (that is, as what is now know as 3 xv. baileyana), but this does not seam likely nor, if true , would it pate aes, the latter name, since Lamarck did not propose it asa Nair & Rehman (1962) describe the pollen of 4 "V- TT) aS "Size 43y (range 2-6 ps)» . Endocolpium lalongate (5.2 x a 220 PHYTOLOGIA Vol. 9, no. k Ectine surface psilate (very faint 2 " Paes base this on what appears to have been a cultiva ucknow, India -- "N[ational] B[otanic] G[arden] 166733 “S1[ide} "2729". Not having seen this specimen, I am, of course, not at all certain that the information actually applies to V. hastata, a species not as yet known to me as cultivated in India — while V. temisecta Briq., is there cultivated and me: Bach roggs have been misidentified as "V. pinnatifida" if only the leaf character were considered. It is worth pointing out ent that the homonyms referred to in e py ata td of V. hastata, and elsewhere, are disposed as fol- lows: V. lacunosa Mart. is Stachytarpheta lacunosa Schau.; V+ eet Willd. is is V. litoralis H.B.K.; the V. pinnatifida of M. Ee Jones is V. wrightii tii A. Gray, while that of "Nuttall and of Schau- er are V. bipinnatifida Nutt.; V. hastata x angustifolia Hill - bracteata Lag, & Rodr., while that of Pammel is xV. moechina Mo wol- denke, and that of Rydberg and of Schnack are xV. perriana Molden ke; another "Ve hastata x simplex Gates" is V. simplex Lehm.; Vs ta x spuria Haartman is xV.1 baileyana; the V. hastata x stricta Allen is V. stricta f. albiflora Wadmond, while that of on, of Gates, and of Rydberg are xV. rydbergii, and Lng tina Ravenel is Stytoen carneus (Medic.) Moldenke; the V. hastata x urticaefolia of Blanchard, of Chase, of Dutton, of McCoy, y, of Nor Pammel, sits ge aaa engelmannii; V. hastata x xVe urticifolia of Blanchard, of Dermen, of eat: of Farwell, of Perry, and of Pringle are all xV. engelmannii, while that at Sa is xV, illicita Moldenke; V. hastata var. ta var. oblongifolia andrews is Vv. urticifolia, while that of Nuttall is xV. engelmannii; aon Ss = we ctfolis x hastata Gates is xv. engelmannii. In my Prelim. ph. List Invalid Names 1 erry (1933) in Placing V. hastata f, sabi OS) Stewt Ps gi » paniculé- ell in the | regarded as being in the synonymy of V. hastata !- rosea Cheney. She followed Small & Heller in placing | V, hastate do not it is of hyb rid o: tion 0 type to cue ought tifsae this point. His original description is 1963 Moldenke, Monograph of Verbena 2a Briquet also keeps V. hastata and V. paniculata as separate spe- Roberts (1929) quotes Haartman (1751) to the effect that "Ano- ther , between Verbena hastata female and Verbena paris male is stated to have originated naturally in the Bo Gar- den [at Uppsala, Sweden] in 178, perishing two years tocar, It is recorded as arising in the same bed with the two species named no s as "1764", This hybrid is now knom as xV. baileyana Moldenke. The Henderson 795 collection from "Columbia River Cascades", He anuary 16, 188, says that the species grows in both counties. In Castanea 9: 69 (19k) my no. 120% 4 s cited from Loudon e at Rogersville, Holmes County. The reference is a typographic error for no. 12171. The c.&W. 232 cited below from Roberts and ma: ve b om Big Stone County, Minnesota, tead, The Wright 553 cited Salad from Rolette County, North Dakota, was collected in the Turtle Mountains, which are 60 percent in that county and 0 percent in Bottineau County. he dase 181 and A. S. Hitchcock s.n. [Douglas Co., 8.7.99], cited below as 3 Ve hasta- ta, were jienainvone as V. urticifolia x hastata and as V. hastata x 8 respectively by Gates in his Kansas Flora Kausch s.n, (Rostock, 1869] is a mixture with v. officinalis; Glevenger s.n, (Fletcher, Aug. 13, 1897], A» Fitch sons wn, [East Greenwich, 1665], and Brinton s.n. [8.9.1876] are mixtures with ert s.n. (St. Louis, Aug. 8, 1875] is a mixture with xV. engel- mannii; di; and E. W, Fell 57-736 is a mixture with xV. rydbergit. McVaugh 147672 is not at all typical — its leaves resemble those o of V, urticifolia, but the corolla is said to have been blue~violet. The physico-chemical properties of V. hastata in relation to phy togeog raphy are discussed by J. A. erris ris (193k), 0.27628 f from Bins drift zone of a sandy beach on the Lake Ut 222 P-E-Y;T-0 &-0-G.2-8 Vol. 9, no. k involving the whole plant and apparently very destructive, caused by an acarid, Eriophyes sp., originally published by Chadwick (1908) . Hocking (1955) says that the dried aerial portion of ve hastata is called "verbena" in the pharmaceutical trade and is. used as a diaphoretic, expectorant, tonic, and vulnerary in dia- betes. A Ning! glucoside de called verbenalin" is found richest in low The following fungi have been reported as infesting V. hastata in the wild: Aecidium verbenicola Ellis & Kellern., Dicaeoma ver- benicola Arth., Puccinia sydowiana 2 » P. verbenicola Arth., otanical yordeaed, Thompson & Morgan (England) ,’ and Sater (Pimcmiale. Michigan). The collector, Josiah Thomas Scovell, has his surname mis- covill® in ; spelled "gs he University of Michigan herbarium. Miss Elizabeth Ann earth omew, in a letter to me dated March l, 1961, states that Roy Clarkson's records indicate that his core 272 and 1585, te Pocahontas County, and 3109 from eo 2 County, West ¥: irginia resent this species, but I hav as Mead bot, 9san ) —— co. etic : yee erry 33) cites the following 175 additional spec as yet seen by me: NOVA SCOTIA: Hants Co.: Pease & Long 22360. (G). NEW BRUNSWICK: Gloucester Co,: Fowler s.n. [Napisiguit, uit, 30 July 1873] (E). York co,: Fernald & Pease 25247 fpashesat) £6)» on Co.: Louis-Mari wie 1d (G). Saint Maurice lain & Knowlton s.n. et of Three Rivers, 1 Aug. 1923] (G). Sherbrooke Cow Knowlton s.n. [Ascot, 20 July 1923] ae ONTARIO: Elgin Cos: G. L. Fisher s.n. [Aylmer, Aug- s Sar Aug. era G). Piscataquis Co Fernald 296 (E, G). york Co.: Parlin ioe: rth Berwick, 28 ‘ke 1851) (6); Willian Son, (Eliot, 27 , a NEW HAMP- SHIRE: Cheshire Co,; "Robinson 196 (G); Be 185) (0) [Walpole, 1963 Moldenke, Monograph of Verbena 223 28 July 1901] (G). Coos Cos: A. He Moore 4316 (G). County unde- termined: Jesup s.n. [bank of Connt. River] (G). VERMONT: Addison Co.: Williams s.n. (Weybri ridge, 8 July 1908] (G). Bennington Co.: Day 149 (G). Rutland Co.: I, W. Anderson s.n. (Mechanicsville, 6 i. 1906] (G); D. L. Dutton s.n, [Brandon, July 7, 1921] (E). Windham Co.: Robinson 153 (G). MASSACHUSETTS: Barnstable Co.: oe Ie sen. (Centreville, 20 Aug. 1902] (G); pti 398 (E, G)5 ee 8: s.n. [Centreville, 6 Sept. 1896] (G). Bristol Co.: Stur- tevant s.n. [Nonquit, 29 Aug. 1888] (E). Hampden Co.: Morris $.n. oe July 3897] (E); F. C. Seymour 511 (E). Hampshire Cost Robinson 572 572 (G). Middlesex CGo.: Faxon sn, [Jamaica Plain] (G); sae S.No vert 2 10 Aug. 1878] (G); Morong s.n. [Melrose, 17 July 1876) (E); Sturtevant s.n. (South Framingham, 11 Aug. 1888] (E). Norfolk Co.: ae @ gen. (Dedham, 6 Oct. 1901] (G). Martha's ian tes Te A A C. Seymour 132), (G). RHODE ISLAND: Provi- NECTICUT: paecdind Sox Andrews sen. (Southington, 31 July 1898] (G); Hassel s.n. [Southington, 10 Aug. 1897] (E). New Haven Co.: Eaton s.n. [New Haven, 1858] (E); a sen. (Oxford, 11 Aug. 1888] ay: Tolland Co.: Weatherby 5076 (E). NEW YORK: Co.: Von Schrenk s.n. (Fleischmann's, 15 July 1892] (E). Essex Co.: Yon Schrenk s.n. [Keene Valley, 1 Aug. 1891] (E). Queens Co.: Hexamer & Maier s.n, yerapee Neck, L. & 8 Aug. 1853] (G)- ai padmisP atta onic ier LB7L(G) 5 cus, 154 4873 (G). Washington Co.: J. A. Dru- Shel 6351 (E). ogee Co.: Stabler s.n. [Port Chester, 3 Sept. 1886] (G). NEW JERSEY: Middlesex Co.: J. A. Kelsey 173 (G)- mion Co,: Trelease s.n. [Plainfield, 9 Aug. 1877] (E)- PENNSYL- hin Co.: J. K. Small s.n. [near Harrisburg, 13 July : » MARYLAND: Harford Co.: Shull 156 (E). Howard Co.: Arsene 672 (E). Diente OF COLUMBIA: I, Blanchard s.n. [11 July 1 1891] (E). VIRGINIA Elizabeth City | Co.: W. Palmer S.e » 10 Aug. 1901] an. IA: Barbour Co,: 3 Ae He] H. Moore 2569 (G). Pocahontas Co.: Berkley 130k (E), Woods Co,: Pollock sen. [7 Aug. 1897] (E)- FLORIDA: achicola] (E). OHIO: —" : A. W. Chapman s.n. [Apal hoga Co.: Greenman | SR ORRE E 1379 (E, G), 1382 (E). Portage Co Webb 5441 (G). county unde malaee: Riehl sen. [dry places, July @®). ILLINOIS: Cass Co.: Geyer Ss . [Beardstom, July nag Sy . Champaign Co.: H. A. Gleason son Is (0). Cook Co»: Greenman greemnan 1, (E)5 H, H. Smith 565 (G); Steele 106 (B)- Kankakee Coes 22h PHYTOLOGIA Vol. 9, no» 1649 (E). Pike Co.: Davis 3565 (E). Saint Clair Co.: Engelmam ne n, [banks of the Mississippi, | opposite St. Louis, July eas OF INDIANA: Lake Co.: Lansing 2805 (G). IOWA: Boone Co.: L. H el 1806 (E). Decatur Cos: | J. P. anderson s.n, [July 13, oT Poweshiek Co.: M. E, Jones s.n. [Grinnell, Aug. 1877] (E, G)- Story Co,: Bessey s.n. 8.N. » {Ames, S Sept. 1873] (G); L. H. Pammel (E). TENNESSEE: Carroll Co.: Eggert s.n. {Hollow Rock, 5 Aug. 1897] (E). McNairy Co.: S, M. Bain Bain hl, (G). MICHIGAN: Berrien Co.! Lansing 32h1 (G), s.n. [New Buffalo, 20 July i] (G). Eumet Co.? F. C. Gates 10708 (E). Marquette Co.: Barlow s - (Turin, 9 J 1901] (G). Van Buren Co.: Lansing 3 2340 (6) WISCENSIN: Buffalo Cos: Fassett & Hotchkiss 3322 (G). Sauk Co.: Eggert s.n. [near Mirror ag July 1903] {53 Blaran Co.: E. J» Palmer a 58738 (E). MINNESOTA: Becker Co.: M. L, Grant 3073 (E, G). Hennepin Co.: Ballard s.n, ee Lake, July 1892] (G)» Stearns Co.: Chandonnet S.n. Sen. (Collegeville, 29 July 1912] (E). NORTH DAKOTA: Benson Co.: eet Sate [peninsula of Lake Ibsen, 1 Aug. 1899] (G). Grand DAKOTA: Brooking 1903] (E); Thornber s.n. [Broo okings, 1 Aug. 1893]. Kingsbury C0-! Thornber s.n. [Iroquois, 9 Aug. 189] (G). Sanborn Co.: Visher 4439 (EB). KANSAS: Johnson Co,: A. S. Hitchcock s.n, [Olathe, Avg. 1892] (E). Osborne Co.: Shear 202 (G). Reno Go.: White 174 (E)- Riley Co.: J. Be S. Norton 390 (E, G). MISSOURT: Clark Cow: Be Fe Bush 9165 (E). Jackson Go.: Be F, Bush 221) (E). Jasper Co-? mee 3 honed nis (E), 8 (E), ee 10568 (B); Trelease 716 (2)- 337 (E), s.n. [near st. fei Sept. 181] (E), son. (St. Louis, Aug. 1861] (Gq (G); Kellogg 1131 rn ere n, 16 June 1896] (E); Lindheimer 5.n, Ts. sen. [St. Louis, July uly 1839] (E). IDAHO: Ada Coe? Je Ae Clark 253 (E, a). Canyon Baik Macbride 30 (E). WYOMING: Sheridan Co. A. Nelson 2258 (B, ¢), 8439 (E). UTAH: Salt Lake Co.: Pammel & Blackwood 338 (G). COLORADO: Sore Co.: Be Le Johnson 507 (E). Denver Cos: Eastwood 41 (E, G). Fremont Co.: W. B. Anderson son. [Aug. 8, 1917] hs ob undetermined: J. Fletcher 1303 [Kelowna] (Mg). ISLAND: J. R. Anderson | Son. [21 Aug. | 1905] (P1—-l)5521, wp s822) 4 1916] (Po-179931) ; Macoun s.n, [aug. 12th’2 C). MAINE: An- Con: E. oh Me 757 ( 87 A toock CO «3 G. ai eee oe oe ashburn, Sept. 2, 190] (Ca-——8826)9); Fern Pd ald s.n. (aug. 1893) GY; True ee Up); E. P. Walker 1 (Up). Cumberland ¢ =, ih o: E. B. s.n. [Cumberland, I Aug. 1901] (W—491.252), H. N. Wctenke 15858 (Es, ¥, or, Se, Si, Up)- Hancock Co,: Strobel s.n. [Blue Hill Ti, 1892] (um—23). Oxford Cow: Fe W. Johnson 215 (N); rs 2h7 47h (Up). Penobscot Co.: Harvey : SBT eater 6082517) 5 Hiland s.n, [above Old Town, VII-2>- Js Sen. (n. of Orono ono, Vis-33-1929) (La); L. P. Hynes 1963 Moldenke, Monograph of Verbena 227 1371 (Up). Piscataquis Co.: Fernald 296 (C, W—278h75). Waldo Co.: Friesner 9092 (Bt—27853, Bt--27854). York Co.: Cléonique- Joseph 295), (Vi); F. T. Hubbard s.n, (Seabury, Aug. 15, 1901] (Rf); Parlin 1372 (Hi--22995)3 True 275 (We), 2756 (Up). NEW HAMPSHIRE: Belknap Co.: True 290 (Up). Carroll Co.: Pierce s.n. {August 19, 1889] (Du--2)182). Cheshire Co.: Ottley 5750 (Go). Coos Cos: A. H. Moore 062 (Up); Pease 35460 (Hi—199501). Graf- ton Co.: Bowen 35b (Up); W. V. Brown 1928 (H—63336); C. He Hitchcock s.n. [Hanover] (Dt); Langdon s.n. [Campton, July 25, 1893] (Mi). Hillsboro Co.: Batchelder 2113 (Dt); H. N. Moldenke 1902) (Es, N, Or, Ot, Se, Si, Sm, Up). VERMONT: Bennington Co.: Me A. Day 149 (Vt, W—38488). Caledonia Co.: F, Blanchard s.n. (Barnet, July 188] (Ca~192068), son. [Barnet, Aug. 1886] (Dt); Grover s.n. [July 2h, 1890] (Ob—50799). Chittenden Co.: Core, Frye, & Core 4811 (We); Dole 111 (Vt), 501 (Vt); Fe As Re Sete (Burlington, July 23, '03) (Vt); Grout s.n. (Burlington, July 2h, '91]) (Vt); Herb. Univ. Vermont s.n. [Burlington, July 7, 1907] (Vt); Pringle s.n. (Essex, 18 August 1876] (Vt), sn. (Charlotte, 29 August 1878] (Vt). Franklin Co.: Pease 36488 (Hi— 19922). Orange Co.: J. F. Reed sen. [Fairlee, Oct. 22, 1931] (H- 27587). Rutland Co.: Banker 250 (N); Carpenter s.n. [8/13/05] (Vt), sen. (La); D. L. Dutton sen. (Brandon, Aug. 10, 1910] (Vt), Sen, [Brandon, July 7, 1921] (Cm), s.n. (Brandon, July 9, 1921] (Um—221, va); Eggleston s.n. (Rutland, 8-90] (Dt); Osborn s.n. (Pittsford, July 20, 1980] (Pu). Windham Co.: H. N. Moldenke 9886 (Ur); Moldenke & Moldenke 9886 (Go, N). Windsor Co.: F. Blanchard 8.n. [W. Windsor, 1873] (Dt); Chivers & Poole son. [Norwich, 7/ 19/39] (Go); Loveland s.n, (Norwich, 7-28-1875] (Vt); Roller s.n- (Plymouth, 7/15/36] (Nj). County undetermined: E. Brainerd s.n. (10 Aug. 1879] (Vt); Collector undesignated s.n. [Lk. Champlain, Aug. 17, 1881] (Vt); Dike s.n. [August] (Vt); Herb. Univ. Vermont 9589-17 (Vt). MASSACHUSETTS: Barnstable Co.: Beetle 534 (Pl— 3 Fernald 669 (W--1885809); Fernald & Long 19032 (Al); Br 563 (Up—82h39), 3593 (Up, Up); J. P. Poole sen. [Mashpee, 8 shire Co.: Churchill S.n. [North Easton, July 28, 1893] (Rf); Blomberg sn. [August 1903] (S). Dukes os ee 2h6S (Up); F. M. Jones 5-N- {August 17, 1940] (Up); F. C. Seymour 132 (H—107712). Essex Co.? Je Blake Bete [8.1882] (S); Chivers s.n. [Amesbury, July 7, 1 (Dt, Dt, Dt); Ss S- Clark 263 (La), 263a (La); A. H. Moore 12 (Io—131755); Morong s.n- [Jul. 28, 1668] (N)3 K. L. Powell sone 228 PHYTOLOGIA Vol. 9, no. (Ipswich, Aug. 1, 1921] (Sd—-5965). Franklin Co.: H. N. Moldenke 8846 (N); R. E. Torrey s.n. [July 20, 1910] (Ms). Hampden Co.: Re H. Piper 13]1 [Dorwart or Herb. Amherst Coll. 68337] (Ok); Fe Co Seymour 511 (H—107713, Ms, N). Hampshire Co.: Barlow s.n. [Am- herst, July 11, 1905] (Ms); A. Clark ssn. [Amherst, August 1875] (Ms); Ewer s.n. (amherst, 7/11/28] (Ms, Ur); Goodale s.n. [North Hadley, 31 July 1926] (H--107715); Jesup 1956 (Dt); Peabody s.n. {Amherst, July 187)] (Sd—h253); C. S. Plumb 395 (Ms); F. C. Sey- mour 2669 (H—~10771)); R. E. Torrey sen. [Amherst, 8-22-11] (Ms). Middlesex Co.: H. H. Bartlett 2h2 (Mi); F. S. Beattie 14)09 (Es), Gene (Aug. 8, 1927] (Or—20970), s.n. (Po—1h6577); A« He Clark 160 (Gg—31327); Cushman s.n, [Middlesex Fells, July '89] (Up— 69671); W. Deane s.n- (Aug. 31, 1912] (Dt); G. B. Grant 6111 (Po- 267625); Grover Sen. [July 26, 1895] (Ob—50793)3 F. C. Seymour Sn. Waverley, July 2h, 1908] (H—~-107716). Suffolk Co.: Blazic Sn [July 1922] (Gg—-31329); Boot s.n. [Boston, 1816] (8); —————=— SeRe (Dorchester, Oct. 1, 1898] (Ba); Girard s.n. (Can- peated 35) 'l8] (Bm); E. L. Greene s.n. [Boston, 25 aout 1833] ene V. Brown 3083 (Au); W. F. Carr $- Re [West Sterling] (se—1)937); Hathaway 681 (S); R. H. Piper 52h (Ok), 1177 (Ok). Martha's Vineyard: MacKeever MV Oo (Gg), MV. hO2a (Gg); F.C. Seymour 132) (N). Nantucket Island: E. P. Bick- Rell 7339 (N), son. (Sep. 2, 190k] (N); MacKeever N.286 (N, Um, : D - Uneatena Island: D. S. Francis 37 (Ok). County undetermined: J. Lund s.n, (Edgeworth, Aug. 11, 1882] (0k); J. Torrey soMe 13833) (Br). RHODE ISLAND: Kent Co.: H. N. Moldenke 19038 (B, Es, » Fy, » Mr, N No, Or, Ot, S. Se Si, Sm 8, Up). Block Is- land: P. Spaulding s-n, [Block Island, bees tole) (Gg—31323, W—1118326),. County undetermined: A, Gray sen, [188] (S). CON- NECTICUIT: Fairfield Co.: J. H. Bardhan 5], (N); Bisaillon 70 (Um—51566); S.C. Brooks s.n. (Greenwich, August 25, 1908] (Ca— 377695); Jesup (Westport, Aug. 1850] (Dt); c. L. Pollard 1 (W—3096h2); P. Wilson seed 347 (N). Hartford Co.: M. Brandegee S.n. [Berlin] (Ca—6812]); H. c, Cowles s.n. [Sept. 13, 1892] (Ob—S079k) . Litchfield Co.: J. Dryer 2368 (N); Meredith SNe (ine 23 1927] (8). Middlesex Gon: We Ee Hunt ban [Essex July 15, 1922] (au). New Haven Co.: W. W. Denslow s.n. [New Haven] (Ms); Se rae, [New Haven, July 187] (Ge—31331); J. We yout 2* ne [July 1864] (Ms). New London Co.: aa speek et 1927] (S), See (Groton, July 16, 1929] (Go), sen. (Groton, July 18, to ott)s Sete (Groton, July 28, 1929] (Bw), sen. [Groton, Aug on te cee 22), Sasss [Tymo, hugs 18, 2957) (60); P. J $08 S-ne (Groton, July 17, 1928} (Io—13536) ; Lumsden s.n. [Aug- 7, 1884) (ob—so802) ; M+ A+ Setchell s.n, (Norwich, July 10, 1883] (Ca—25183). Windham Co: J.T. Sheldon s.n. (Central village, —— te S.n,. ee 1963 Moldenke, Monograph of Verbena 229 June 1903] (We). County undetermined: Herb. Gray son. (S); C. Wright s.n, (Cm). NEW YORK: Albany Co.: H. D. House 19599 (a1); Whitney 654 (Al), 1097 (Up). Bronx Co.: Ahles 505 (Aa), 777 (Aa) ; E. P. Bicknell 7341 (N), 73ula (N), 7342 (Al), 734h (N), 7346 (Nj); Clute s.n. (Bronx Park, July 21, 1898] (Bt—22923), sen. (Pelham Park, July 12, 1899] (N); Gilly 105 (N); Holtzoff s.n. (McLean's Wood] (N); L. Johnson s.n. (Wmsbridge, Aug. 1, 1876] (Lh). Cattaraugus Co.: Alexander & House 12481 (Al), 13005 (Al). Chautauqua Co.: G. B. Grant 530 [3059] (Po—267672); Southworth Sn. (Forestville, Aug. 17, 1885] (Mi). Chemung Co.: Munz 173033 (S). Chenango Co.: E. E. Davis 6 (Al), 82 (Al), 129 (A1), 1028 (Al), 1088 (Al), 1119 (Al). Glinton Co.: Whitney 199) (Al). Col- Co.: McVaugh 107 (Al, Up), 14767a (Mi); Moldenke & Molden- ke 11745 (N), 19527 (Sm); C. He Peck s.n. [Boston Corners] (Al); W. A. Weber 11), (B1—-)8178, Io—152915). Cortland Co.: A. Re Moldenke 1 [H. N. Moldenke 2016] (N); C. He Peck s.n. [Marl Pond] (Al). Delaware Co.: Ferguson, Ennis, & Ottley 2685 (Me), Sn, [Ottley 2685] (Io--127769)3; Ottley 2685 (Me); N. Taylor 792 (NX); P. Wilson sen, [Arkville, July 18, 1915] (Al, N). Dutchess Co.: B. Sherwood s.n. [road to Wingdale, Aug. 7] (Sh, Sh, Sh); Standley & Bollman 1193 (W—895512). Erie Co.: G. W. Clinton 5. Rhoades s.n, [Little Falls] (Hs). Jefferson Co.: Keyes SMe [Wat ertomn, July 1889] (Ca—199029); Muenscher & Maguire 2500 (Ca— 513006); True 926 (Up). Kings Co.: Ruger son. [East Williams- burgh] (Te). Lewis Co.: Hotchkiss 871 (Al). Madison Co.: Haberer 2748 (W—1200262); Maxon s.n. [near Breese's Woods, July 26, 1895] (W—671,569). Monroe Co.: Barss s.n. [Rochester, 7 September 1909] (or—8862); Holger s.n. [July 1863] (Al); W. A. Matthews 3154 (Hi—29791, La); As R. Moldenke 8 [H. N. Moldenke 201%] a Co.: G. T. Hastings s.n. [Tully, July 2h, 1895] (N)5 Straub s.n, [p. Syracuse] (W-—317094). Ontario Co.t A. Re MOUCS™ =e [H. N. Moldenke 2043] (ar, = oreeee et 5: Me ee + [Goshen] (N); Raup 7656 (Mi, N). Orleans Uo« leonen 2s be" y 1893] us 03 = Govt Hotchkiss 3059 (Al); 0. E. Pearce psd NT 230 PHYTOLOGIA Vol. 9, no. [N» Hannibal, July 5, 188)) (W—147577); C. S. Sheldon s.n. [July 20, 1878] (Al). Otsego Co.: F. W. Johnson s.n. [Cherry Fam, Aug. 27, 1921] (N). Putnam Co.: H. N. Moldenke 19530 (Ot); Moldenke & Moldenke 11853 (N). Queens Co.: W. C. Ferguson 7732 (N); Hexamer & Maier s.n. {Willet's Neck, L. I., July 9, '53] (Cm). Rensselaer Co.: Banker 1395 (N); Puissant s.n. [West Troy, July 28, 1865] (Br), s.n. [Troy] (Br, I). Richmond Co.: Hollick s.n. {July 15th SS Coe: H. Ne Moldenke 19522 (Es). Schoharie Co.: H. N. Moldenke 19518 (Es). Schuyler Co.: McCarthy s.n. (Watkins, July 1687] (W— 7286 (Po—-267627) ; S. L. Clarke s.n. [Quegue] (N); Disbrow s.ne [East Hampton] (Nm); H. W, Graham sen, (Deer Pond, July—Avg- 1926] (Cm); L. K. Henry s.n. [Deer Pond, July 1929] (Gm), Sele [Long Pond, July 1929] (Gm); Latham 38 (Al); Muenscher & Curtis $430 (Al); C. Skottsberg s.n. (Montauk, 10/8/1926] (Go). Sullivan Coe: Eiten & Jensen 1463 (N); W. A. Weber 208 (B1—-18179, Io— 150795). Tioga Co.: A. R. Moldenke 2 [He N. Moldenke 20420] (N)- T Cos: L. H. Bailey s.n. [Ithaca, Apr. 10, 1919] (Ba), 8: ne (Ithaca, July 27, 1925] (Ba, Ba, Ba); Hoisington 47 (Ok— 1560), 160 (Ok—15659); H. S. Jackson s.n. (McLean, July 2, 190k] (H—107717); J. M. R. san. [July 15, 1926] (Gu—E.1051); X. Z- Lee sn. (Enfield, July 1898] (N); Lennon s.n. {July 23, 1896] (Al); Locke s.n, [Fall Creek, Sept. 19th, '79] (Fe), SMe (Ithaca, Sept. 25th, 179] (Fc); B. Maguire 6662 (Ua—26812)} As Re Moidenke 3 (H. N. Moldenke 2026] (Mg, N), 5 [H. N- Moldenke Saal Macfarlane s.n, [Phoenicia Valley] (Up—l687). Warren co.: Mar- go —~ = (We). Washington co.: S. H. Burnham son. [August 1912] [ He De House 29386 (Al, Ok); Jellitte S.n, [Huletts, July '87] (C). Wayne Co.; H, idenke 2019 (N). Westchester Co.t E. P- N. Mo. Bicknell 7340 (Al, N); Collector undies ted s.n. [Tarrytown, Aug. 3, 196] (N); H. A. Gleason ee ae N, S); Mackaness 1963 Moldenke, Monograph of Verbena 21 Son. [July 20, 1937] (Tl); H. N- Moldenke 19091 (N, Or); F. We Pennell 6681 (N) « Wy Wyoming Co.: Herb. Univ. Te sen. [Wy Aug. , 1859] (Ur). Yates Co.: A. Re Moldenke 10 i. fae sie Moldenke 20448] (N, S); W. S. Phillips 221 (Al); S. He Wright s 15,03). County undetermined: Brourme s.n. [1045] (S); Col. ner undesignated s.n. [July 15, 1896] (N), sen. [August 25, 1896] (N); F. C. Ferguson s.n, [Plattsdale, L. I., ~7=18-19] (N); J. M. fs 8. n, (Renwick, July 26, 1926] (Gu-—E.1050); Ottley 1659) (er) ; Sn, (Tc); J. Torrey "16 (Al, Al), sen. (Pa). NEW JERS: arte -Z Cow: H. Xe Moldenke 21576 (Fg); We Stone 12057 (Up). ae Coe: Dautun s.n, [Fairview, Aug. 2, 1903] (La); G. T. Hastings s.n. [Forest Vie fice Landing, Sept. 5, 1917] (N); K. D. Kimball s.n. [Moon achie, Aug. 12, 1916] (N); ; Niederer s.n. [Carlstadt, July 26, 1883] (Ws Hrdberg sane sen. [Ft. Lee, July 30, 1921] (Po—174819, Um—203) ; r sen. (Ridgefield, J 27 1879] N). Burlington Co.: Fogg 9212 (Up); H. A. Green tg 2 a ae (Cm); W. Stone 12935 (Up). Camden Co.: ‘Fogg 9200 (Up); L. P. Hynes 139 ’ (Up), sen. [Ju- 98 1e 521027). Cape May Co.: Fender Bog (UP)5 Fou 9406 (Up); G. ardsons, Sept. 1889] ia) 6 S.n. (Fickenasces » 8-30-91] (op) - Hud Son Co,: eae s.n. (New Durham, Aug. 2h, 1894] (C)5 J. Torrey 125 (W—26],611); Yon Rabenau s.n. [Herb. Osten 13023] (Uj ie. Hun- terdon Co,: H. ibe “Fisher s.n. (Stockton, 7-18-97] (Ur). Mercer Co.: Collector undesignated s.n. (Princeton, 2 Aug. 1876] (Pr); Macloskie s.n. [Princeton] (Pr). Middlesex Co.: J. A. Kelsey 173 (ui, Ua--11366, Ua--27138, We), s.n. [Halsted 173] (C, Fe, To. 15318, Vt); H. N. Moldenke 1911 (Or); J. A- Small s.n. [Cheese- make, 7/21/39] (Nj ); Thais s.n. (Mt); Vail s.n. son. [New Brunswick, July 18th, 1890] (N). Monmouth Co.: Brinton s.n. {Sea Girt, Aug. 27, 1891] (Ca—25181); Dautun ae cee Cherry Hill, 2 Aout 1908] La); H. N. Moldenke 21731 731 (Sm). Morris Co.: G. V- Nash s-ne . 5, 189 1106 (Up), 1211) (tp), ), 11166 (Up); arpiterse bt: s.n. [Seaside | Park, July 21, 1900] (Up—L361h); L. Pe Hynes S-0e n. [September ll, 1938} (Up); J. A. Small s.n, [Island of Long 1936] (Nj). Passaic Co.: Disbrow s.n- [Greenwood Lake, July ' J (Nm); H. N. Moldenke 21878 (Fg); G- Ve Nash sen- [Clifton, Sept. 4, 1892] (Ka, Lh). Salem Go.t Adams & Adams 1708 (Up); gar 7hO3 ; (E, Go sme = t) Somerset Co.: He i, sous oar on 232 PHYTOLOGIA Vol. 9, no. (Go, N, Um—232, Ur), 8090 (Go, N, S), 20308 (Wh); Shetler 292/1 (N); Sumner s.n, (Bernardsville, Aug. 17, 1897] (Bl—l232h). Sus- sex Co.: Adams, Adams, & Wherry 1521 (Ca—-5885)6); Fogg 12535 oe Lisi 185 (Up); K. K. Mackenzie 833 (N); H. N. Moldenke 21551 (Ss); G. V. Nash s.n. [July 16, 1909] (N); H. H. Rusby s.n. (Franklin, July 26, 1873) (Mi), s.n. [Franklin, Aug. 1979] (Mn— 16086); Witte s.n. [Aug. 9, 1932] (Up). Union Co.: P. Clarke s.n. (Great Island, Aug. 2, 1902] (N, N); Kezer s.n. [8 . County undetermined: Harshberger s.n. [New Jersey, July 1889] (Up- 2153); Paddock s.n. [1838] (Ur); 0. E. Pearce s.n. [N. J+, 1884] (W—218873); W. Stone 3467 [Cateksin Camp] (Up); J- W. Wood sme (Aug. 1842] (Ms). PENNSYLVANIA: Allegheny Co.: Bright 55 (Cm); 0- E. Jennings s.n. [Aug. 22, 1907] (Cm), sen. [Aug. 27, 1921] (Cm), sen. [8-21-13] (Cm); Shafer s.n. [July 22, 1900] (Cm), son. [Aug. » 1900] (Cm). Armstrong Co.: J. M. Fogg 15349 (Up); 0. B. Jem nings s.n. (Sept. 27, 190] (Cm). Beaver Co.: Andriessen s.n. Q. King Auge 1919] (Cm), Berks Co.: Collector undesignated s.n. [prope dor sen. [July 12th 89] (Cm); Shafer s.n. [July 13, 1889] (Cm)- Bradford Co.: 0. E. Jennings s.n. [Aug. 21, 1929] (Cm); H. Ne Moldenke 17080 (Br, il, Nd, Se, St); W. Rhoades s.n, [near Gro- ver, July 1933; He N. Moldenke 17080] (Bt, Bt--299l1, N)3 He Ae Wahl 17597 (Hi—16873h); Westerfeld 817 (Cm). Bucks Co.: Dreis- bach 368 (ui) ; MacElwee s.n. [July 22, 1893] (N); McDowell 453 (Up); O'Neill s.n. [August 16, 1930] (I). Butler Cos: Bright s- n, [Aug, 2, 192k] (cm); J. Me Fogg 15305 (Up); L. K. Henry S-M- lang. 1927] (Cm), sen. (9/16/40) (cm); 0. E. Jennings son. (8 19/22] (Ga); Krouse son. [7/26/i7] (Hi—133086); N. Russell He eel (NR.2355] (We). Cambria Co.: 0. E. Jennings 17 (cm), 5-De July 2h, 1908] (Gm); W. Stone 8870 (Up). Cameron Co.: J» Me ( 11538 (Up), 18096 (Up): Jennings & Jennings s.n. (aug. 20, 1925] (Cm); H. A. Wahl fan (i Ur). eine Cos: M. Te Travis 714 (Up), 1787 (Up); Westerfeld 648 (Cm). Chester Go.: T. W. Ed- Rondson 6337 (N); F. W. Pemell 12262 (N); M. T. Travis 591 (Up). Clarion co,: OQ. E. Jennings s.n, (July 21, 1904] (Gm), 8: QO. Ee J Sen. [July 12, 1908] (cm). Clinton Co.: Wahl & westerteld 12057 (Ur). Crawford Co.: nee J : ] (Cm), s.n. [aug. h, 1909] (Gm); James A. Murray S-le "é- 1, 1928] (Cm); Shafer s.n. [July 23, 1901] (Cm). Cumber- eee ee 1963 Moldenke, Monograph of Verbena 233 land Cos: A. R. Moldenke [H. N. Moldenke 21767] (Bm), 99 (Fg) Dauphin Co.: He Ne Moldenke 19369 (N). Delaware Co.: Eckfeldt 8. s. [July 18, 1938] (Up); F. W. Pennell 14689 (N), 1,754 (Cn, Fl— 31477, B~56567, Io—153886, It, Um--235, Ur, We); W. Stone 552 sn. [July 19, 190k] (Cm) fr Jennings Jennings 8. n, ae {1/267i925} (Cm, Gn); N. H. Phillips 84 (Up), Tee (Up). Forest Coe: Kosinski s.n. [8/25/1940] (Cm). Franklin Co.: 3 Collector un- designated s. sen. (Concord, July 13, 188] (Mi); H. Ne Moldenke 19167 (N). Fulton Co.: A. R. Moldenke 101 (Fg). . Huntingdon Coe: E. He H. Graham sen. [7/12/32] (Up); M. T. Travis 790 (Up); is Ae Wahl 1368 (P1—-129973, Ur). Indiana Co.: 0. E. Jennings [uly 1, 11, 1908] (Cm); Shafer s.n. [Aug. 8, ; 1901] (Cm). ae A Co.: E. H. McClelland gn. [8/7.10/22] (cm). Juniata Co.: J» Me Fogg 15530 (Up). Lackawanna Co.: Charnell 569 (eis Glowenke 7772 We); Twining s.n. [July 15, 1907] (Cm). Lancaster Co.: o.: Brubaker né (Up), 1315 (Up); F. W. Pennell 13093 (Cm, a Sanaa irs (Up), s.n. [July 28, 1937] (Up). Lawrence Co.: L. E. Fox Al Herlen 60 (Cm); 0. E. Jennings s.n. (Sept. 18, 1919] ore Whit, tenberger s.n. sn, [August 30, 1939] (Up). Lebanon Co.: C. H. Kauff. . man s.n. [Lebanon, August 22, 1928] (Mi); H. N. Moldenke 19370 (N). Lehigh Co.: O'Neill s.n. (Allentown, August 19, 1927] 7] (Fl— 21112, I). Luzerne Co.: Glowenke 2875 (Up) » Lycoming Co.: J. Me Fogg 11588 (Up), 17719 (N)j We Re Tayler gon, [Essick Heights, 1912] (Up—-75580) . McKean Co.: 0. Jennings s.n. [July 21, 1904] (Cm), sen. (Sept. 6, 1937] eat Mercer Co.: L. K. Hi (Ca); 0. E, Jennings s.n. "Tang. 30, 1923] (Cm); Shafer s.n. [July 17, 1902] (cm). Monroe Co.: Bernhardt oe [Paradise dise Falls, Aug. 18, 1928] (Mi, Up); Gress, Jennings, | & Jennings S.n- [Tobyhanna ] (ca) ; Henry & Graham s.n. [Aug. ug. 1930) ee Jennings, Jennings, & Gress s.n. (Aug. 1920] (Cm); Langman 112) (Up). Wont gonery Coes I. Me Fogg 911 (Up); MacElwee s.n. — 26, 1892] (N); Wismer 547 (Up). Northampton Co.: Tyler 216 (Br), s.n. (Easton, July July 1h, 1896] (N). Northumberland Co.: J« M. Fogg 15013 (Up). Perry Co-: A. A. Heller 14222 (Du—157211, Se——81873)5 A» Re sacle 17 (H. N. Moldenke 20459] (F); H. He Eee s.n. (Sept. | (Up). Philadelphia Co.: Brinton s.n. [6.9-1070) Neate I, Burke 3.n, (Commons, Phil.] (Up—17108); Breisbach 1702 (Up); Mack™nee MacElwee 1074 ee G. Watson s.n. [Philadelphia] Gotclg seg Pike CO.t Kill Co,: Arndt s.n. [Auge 17, 1 1915] Tip) ; P. Re Wagner 668 2258 (Up), L878 a $636 (Up), 7265 (Up)- Suyder Co.t He Ne H. Ne 234 PHYTOLOGIA Vol. 9, no. & Moldenke 20479 (N); Wade & Wade 1385 (Up). Somerset Co.: 0. E. S Sen. sen. [Sept. 15, 1919] (Cm); B. H. Patterson s.n. [July 19, 1872] (Cm); Van Dersal sn. [7/31/34] (Gm). Sullivan Co.: W. Stone 5877 (Up), aene (Up). Tioga Co.: Gress Gress, Jennings, & Jen- nings s.n. (Sept. 2, 1920] (cm); H. N. Moldenke 2066 (1 (N). Union Co.: ee este 2 (8/25/36 (Cm). Venango Go.: Wurdack s.n. [9/6— — arren Co.: 0. E. Jennings s.n. [8/22/29] ] (Cm); Jen- cmetgass Sen. [Iron : Springs, ngs, 1928] (G (Cm); H. N. Moldenke — TeGee se . 17696 ( (Br). Washington Co.: Jennings & : McKee $.n. (Aug. 10, 1919] (cm, Up). Wayne Co.: Dix 125 (Up); Jennings, Jen- nings, & Gress s.n. [Aug. 21, 1920] (Cm). We Westmoreland Co.: Block SNe [July 188 1881] (Cp, I); Demsire S Sate [8/20/35] (Cm); L. Ke. Hen- Fy 551 (ca—882656, Bi—192860, v 0935), ane [7/29/37] T (Ga), she A. D. | ay ae S.n. iene BASAL} (nye Tox “Cont re = Lloyd s.n. [Aug, 6, 1889] (Cm); W. Stone 2164 (Up). County unde= termined: Eifrick ssn, (Mt. Lake Park, Aue. ~~ 25, 1901] (Cm); ‘pe Schweinitz s.n. Pennsylvania, 1829] (Br); C. W. Short s.ne sylvania] (Iu). DELAWARE: kK Kent Co.: E. L. Larsen 310 310 (Up), 12h0 (Gu--138L0, Up). New Castle Co.: & . L. Larsen 662 (Up). Sussex Cos: N. L. Britton 27 (N); R. Re Tatnail 1 4302 (Up). MARYLAND: Al- legany Co.: Shriver s.n. [May 189] (N). Baltimore Cos: Boldo 13 (Q). Caroline Co.: E. C. Earle 1566 (Up). Cecil Co.: W. L. abbott 2448 Eich, keotara 1383] (W—2160h60); H. N. Moldenke 13602 (2). Garrett Co.: F.C. Lane Lane 1721 (Ur, Ur); J. D. Smith s.n. [July 15, 98] (130355 3)- Hanover Co.: A. R. Moldenke 491 (Fg). Harford Co. Shull 156 (N); Tanger s.n. [Aug. h, 1937] (Up). Howard ably A» Re Moldenke 489 (Fg). Montgomery Co.: Tidestrom s.n. (Chain bridge, Oct. 1 1897] (I). Prince Georges Co.: T. Holm s.n gn (Juni 1896] (I); Hyacinth 1457 (Gg—31321); A. R. Moldenke 492 (Fe)3 Petrak s.n, - (20. ¥Ir 1950) (S), sn. [27.vI1I.1950] (S)3 ° True = (Up). Talbot Co.z E. C. Earle 1596 (Up), 1603 (H—-S2hik) . Wa ected Beceend 198 198 (I). DISTRICT OF COLUMBIA: F. Blanc THB) (renee T1)) ts ele ga ite HDs Gamby y sn. [D- Crs ig ¢ Holm s.n. [18/7/1911 I, s Kearney 5.ne poo ais Opa iecnard Las bs (Cn)3 7 F. sags Sargent 7608 j Be Se Steele san. ray 36, 2 556. (Ob—50806); Wismer 368 ip). VIRGINIA: aed City Co.: A. R. Moldenke denice 190 "fie 36681 (N). Fauquier Co.: Allard 3251 (¥~1734757) « Floyd Co. Bar2 (Bm). an “ss rea Re Moldenke 48 [H. N. Moldenke 21772] a P. Schallert Ur. s.n. (Winchester, 8/27/39] 1963 Moldenke, Monograph of Verbena 235 (Ky). Giles Co.: J. M. Fogg 13011 (Up). Grayson Co.: Radford Radford 2613 (Hi--35271). James City Co.: Artz 1131 (Up). Lo adoun Cov: A. A. R. R. Moldenke 68 [H. N. Moldenke 21793] (Bm). Montgomery Co.: Re Kral 959 (Hi—201637); Murrill s.n. (Blacksburg, August 1894] (I10—30879). Patrick Co.: A. R. Moldenke 49 [H. N. Moldenke 21773] (Bm). Roanoke Co.: Fauntleroy 659 (W--694518); H. N- Mol- denke 19248 (N); C. BE. Wood 6239 (Hi—51277). Smyth Co.t A. R. Mol- denke 60 (H. N. Moldenke 21785] (Bm); ze K. Small oa (st. Clair's Bottom, July 30, 1892] (C, Ca—25182, Up—-17106, W). Washington Coes A. Re Moldenke 58 tH. N. Moldenke pat ] (Bm). Bee Cos: As R. Moldenke 62 [H. Ne Moldenke 21787] (Bm). WEST VIRGINIA: Barbour Coe: A. H. Moore 2569 (Po—-152h5). Berkeley Co.: A. R. Moldenke 4? (H. Ne — 21770] (Bm). Cabell Co.: F. A. Gilbert 1000 (We). Fayette : L. We Nuttall 345 (We); smithson s.n. (1) July "36] (We). sapanse Co.: Frye 1,83 (We). Hancock Co. Sumpetine Sen, [July 1938] (We); R. West 551 (We, We). Hardy Co Core 3758 (N); L. W. Wilson 130 (We), s.n. [July 20, 9h ti). Kanarha C Co.: Mrs. Es Re Yates sen. [summer, 190] (We) « Mercer em 8 s.n, [Hartmans a heraiad 1952] (We). “Monongalia Co: er 345 (We); pina aie 345 (We). Nicholas Co.: E. A- Har- Hac san. [July 19, 1955) (We); Lyman s.n. eee re 8=1h-79) cee (We); Guttenberg sn. [July 9, 1878) att Pocahontas Co.? Clarkson & Clarkson 612 (We (We) 3 E- Le Core 3540 (Ob—50795)5 Herbe West Virg. Bot. Exped. s.n. [July 29, 1930] (S)~ Preston Go.: Dar vis & Davis 1ys8 (We); J. 2 Sheldon 1882 (We, We); Spangler 271 (Wie, We). Putnam Co.: E. = ,. core 6463 | (We). Raleigh Co.: Tosh 535 (ca661131, Ky, ee eajea ne Cos: Co Be eee 1775 (We). Summers Co.: Boone 250 (We); Tosh 77 ’ (Ky)- F Goes a 9070 (W), 9119 (W), 92 a (W—-1872154), 12500 (W._1895058), 2168 21 134497); | Ce Be Glarkson 1478 (We); Davis & Davis 765 ( Webster Co.: C. F. Millspaugh 563 (We). Wood Co-: W. Me Pollock S.n. (Aug. 7, “1997] (W—-315367)~ NORTH CAROLINA: Ashe Radford ord 38460 (Hi—104888), 38677 (Hi--104889). Orange Co: "Bad- ford & Stewart 200a bone (ieeeaiaaeas 97). Transylvania Co-? Blonguist U2 st L712 Cow: E. J. Alexander B.D. 1i/eB7e3) ] (Hi—59L83). Watauga Co.: Ah- Jee & Duke nie ts 08 (Hi--104890); ashe s.n. [July 19] (Hi—59Li85]- County unde termined: Ashe s.N. pia ee AP f N. e (H—327850) « SOUTH CAROLINA: Charleston Co.: aibbes af sn. Para — (N)- GEORGIA: Chatham Co.: g.n. (BY); | : Duval Co 8.n. (Jacksonville, ieee ( Oa). OHIO: reer : H. N. yoidenke 236 PIE T.0 L061 Vol. 9, no. 12171 (Co~y5, Da). Ashtabula Co.: I. E. Diehl s.n. [Rock Creek, July 3, 1901] (Ca--373442). Butler Co.: Bridge sn. [7/30/11] (Cn); J. Ferguson s.n. [July 19, 1932] (Ob--80596). Champaign Co.: Demaree 11689 (Ca——591988, W—163219); E. C. Leonard 1609 (Herb. Leonard 350] (W—2162185). Clark Co.: F. P. Davidson s.0. (Springfield, July 1885] (Or—-8861). Columbiana Co.: Nicely Bs arden, July 20, 1958] (We); E. Wilkinson s.n. [31 July 1887] (Ob—8059). Coshocton Co.: H. N. Moldenke 13149 (Co—1290), 13380 (Br, Co--153}, Da, Ec, M1, Nd, Se, St). Crawford Co.: Benson & Yaussy 309 (Po—267071). Erie Co.: C. L. Booth sen. {August 2, 1901] (Ob—75335); Friedel s.n. [near Huron, August 25, 192] (We) Monosmith s.n. [July 13, 1900] (Ob~-75337); York sen. [Cedar Point, 8-1-1902] (Up), s.n. (Cedar Point, 8-3-1902] (Dp). Fair- field Co.: Horr s.n. (Pickerington, Sept. 7, 1833] (Io—153h7) - Franklin Co.: Condit s.n. (Westerville, July 25, 1909] (Ca— 456152); E. B. Williamson s.n, [July 1897] (In). Hamilton Co.: L. Braun s.n. [July 10, 17] (cn), sen. [8/10/17] (Cn), sen- [ 10] (Cn)s R. Buchanan s.n, (L1); M. Mohr 65 (11); C. W. Short +h Bend) an C E. ig — (Co—-l135, Da). Knox Co.: H. N. Moldenke 12118 (Co—lilh), 12172 (Co—L6, Da). Lake Co.: H. G. Beardslee S.n. wis 2, 1870] (Ob—- 60598). Licking Co.: H. L. Jones san. [hug 10, 1889] (Ob—80595). Lorain Co.: H. C. Cowles s.n. (Oberlin, July 21, 1893] (Mi, Ob— 15333) Me Ee Day son. (Ang. 29, 191] (ob—99796); W» M- Dick Be {August 6, 1894] (Du—296882, Ob—75330); Grover s.n. [July 23 1900] (0b—75338) ; F. D. Kelsey s.n. (Ob—50802); Monosmith 5-De es Hollow, July 13, 1900] (iim—18)91, Um—215); 5. W. Phil- ZPE sane [10/8/1892] (cm); A. E. Ricksecker 286 (Ob—75339, Ob— 89h] (Se—28246, St—23390), S+ Fullerton s.n. (aug. 7, 193 ie er 5. letches a. e 7, 1939] (En). Miami Co.: Clevenger 5: 3 i" » Ang. 13, 1897] (W—-152),139). Ottawa Co.: E. Se Ford T9N0] (We) wpe ee Sats [July 29, 1940] (We), sen. [July 31, . (We). Richland co.; Je C. Myers s.n. [August 7, 190] (We)s se The (Cm, La, Ob—80597). Wayne Co.: Le Se Hopi Ok). Jobngon Talamis; sn. (W. Salen, July 31, 1887] (H--2h535, x » July 18, 190k] (Up). ILLINOIS: Adams Co! 2063 (15879) ; 199 (MN, Ur), 802, in part (Ur); Erers; 1963 Moldenke, Monograph of Verbena 237 Jones, & Jones 572 (Ur); A. B. Seymour s.n. [Northeast Turnpike, 20 Jane . 1876] (H—-106462) 5 1 Wehmeyer 19 (Mi, Mi). Boone Co.: Fell & Fell F469), Siatacigib Brown Co.: Winterringer 5253 3(n-- 35417). Cass Co.: Geyer s.n. [Beardstom, July 1842] (S, Vt); Wir terringer 515) (135658) Champs ep Cos: Dembski 67 (Ur); H. 7 Jewett 435 (Mi); G. Ne Jones iene (ur), 1271. (ur), wn8 (Ur), 12805 (Ur), 16557 ( (Ur), 16568 (Ur); Knibloe s.i sen. (Herb. Ill. Ind. Univ. 1760] (Ur) (Ur); Perino s.n. [August 3, 1950] (Ur); saad sen. (Urbana, July 1881] (Sd—l25)); Taft s.n. (Champaign, 18 1876—-79) (Ka) ; Waites sen. (Ur—29661); Winterringer 72 (Ur). Clay Co.: Winterringer 2152 (I1~—2703). Clinton Co.: Spiess 298 — 22631). Coles Co.: Umbach 11)5 ed Cook Co vee +e ° cock 9281 (Ba—23753), Sn. [Chi Jul. is, 187] ( em) Bakker s.n. [Riverdale, July 1912] | (S123 Re Bebb 1686 (Ok), 2109 (Ok); H. R. Bennett s.n. (Barrington, : ugust 2), 1957] (Go); Blatchford 2515, in part (Il--15820); Ss. A. Cain say sn. [southern Chicago, 1926] (Bt—2k72, Bt--275), s.n- [August 1926] (Bt— 2465); Dixon & Gage 767 (W—6O9L7); H. S. Fawcett s.n. (Berwyn, 3/1/05) (Io—28h08); G. D. Fuller 2292 (T1—15880), 8090 (1— 1588h); F. C. Gates 1871.2 (Ur), 1873 (Ur); Lansing 1399 (Ur); Munroe s.n. [Chicago] (Po—-70996); Sherff 1850 (Ur); H. H. Smith 5993 (Up); E. S. Steele 106 (W—608865); L. F. Ward s.n. [Chica- go, August 16, 1897] (W—298088) ; Waugh aay (Ka); Wessén s.n. (Chicago, VII 1899] (Go). Crawford Co.: Ahles 4850 (Ur); eS ringer 3753 (P1—29199). Cumberland Co.: Spooner s.n. [Haz Dell) (15872); Winterringer 3403 (I1—28729), acid rare nhs) De Kalb Co.: P. B. Whitford s.n. wn. [July 19, 1946] (Ur). Douglas Co.: G. N. Jones 8 17523 (Ur). DuPage Co.: G. D. Fuller 1706 (I1—15883) 5 W. S. Moffatt t 573 (Ur); Umbach s.n. [Naperville, July 23, 2897] (W—339h79). Edwards Co.: Winterringer 4177 (11--30075). EBffing- ham Co.: Ahles 2606 (Ur, Ur); Winterringer 5877 (Il—22151). Fay- ette Co.: McDougall 91 (Ur); O'Dell 335 (Ur), -), 336 (Ur). Co.: Ahles 5525 (Ur); } Winterringer Singer 3522 fae tghls Henry at Winterringer 52h1 T3558). Iro Iroquois Co.: Win er (I1—3)123). oe Coe: es & Welch 268 as", Ey yee ee 1021 (II—15875); J. W. Voigt 985 (Il—36535)5 Ge on 1021 (Ur). Jefferson pte. aoe 5496 (Ur); Winterringer 770% - 178k (— 37432) . Jersey Co.: P. Jacobs 31 (Wr). Jo Daviess Co.: Ahles (Ur) ; Pepoon & Moffatt ee 255 (Ur); Winterringer 8156 (1—37H58)- ease er & ee — ea (1135095); &. Je Hill 222.1878 (Ur); G. Ne ip Be deem 15936 (Ur), 16046 (aS “Ur). Lake Co.? R. K. Brown 90 (Ur), 560 (UF), 238 PHYTOLOGIA Vol. 9, no. 4 son. (Sept. 5, 19h9] (Ur); F.C. Gates 3211 (Mi, Ur), 1349h aa 73469); Gleason & Shobe 161 ( (Ur); L. P. “Richardson s.n. (Augus 15, a (St). LaSalle Co.: G. D. Fuller 1797 (11—15082); Pe "Be rd 311 (T1—32222). Lawrence Co.: : Sivert s.n. (Sept. 1h, eazy wis Winterringer 6191 (1133522). Lee Lee Co.: V. H. Chase 530 (Ur); Keithley 29 (I1—-20017), 272 (T1~—31488), sn. (duly 12, 1945] (Ur), s.n son. [July 28, 1946] (Ur); Lyman s.n. [Amboy, 21-9),] (Dt). Livingston Co.: G. D. Fuller 9149 U9 (1115856, Ur). Logan Co.: Ahles 3050 (Ur). Macon Co.: Re Ge Mills s.n. (Decatur, Aug. 10, 1939] (N). Marion Co,: Winterringer 2563 5963 (I1-—-33306) . Co.: F. C. Gates 3887 (Ur). McHenry Co.: ? Nason s.n. on- quin, Aug. 6, 1878] (Ur), sen. (Algonquin, Aug. 26, 1879] (Ur). McLean Co.: Hitierinees 5071 (I1—35890). Montgomery Co.: Win- terringer 3880 (I1—29176). Moultrie Co.: Winterringer 9107 (n— 39392). Ogle Co.: Blount 120 (T1—3083); A. L. Hills 3b3u-0 (I1- 15876), 3452-0 (Ur), 36) 3640-0 (I1—-15858). Peoria Co.: Heading 8M. $Me [1875] (Ur); F. E. McDonald s.n. (Peoria, July 1886] (ur), 8 SoBe {Peoria, July] (Ur). Perry Co.: I. L. Forbes s.n. [Duquoin, vIII- 16-'27] (Io—129208); steiner s.n. [July 1, 19h8] (Ur); Winter- ringer 3945 (I1—29075). Piatt Co.: G. N. Jones 2163 (Ur)5 r)y Win- re Se (Ur). Te Co.: Rapp & Rapp 56 p 56 (N, Ur), 57 (N, es + Pulaski Co.: Winterringer 5992 (I1--33180). Richlan i "Bgeey 3173 1173 (Ur, Ur), 117) (Ur); Scherer 06 (2659, Ur). Saint Clair Co.: J. Neill 1299 geen ar 2 Saline Co.: Winter == 3511 (I1—28927). Sangamon Co.: G. D. Fuller 5039 sa » 5330 (T1—1586), 5572 ieee 9593 (1115870), B49 (1115861), 6099 (I1--15862), 6156 1 e568) He Ne J denke 21882 (Ss). Stark Co.: V. H. Chase 981 (Ur), s.n- [July 10 as ie thy Ur). Stephenson Co,: Ahles 4500 (Ur); Serf 98h (a Winterringer 8100 200 (I1—37535). Tazewell Co.: V. r, H. Chase 3229 (Al, T1—15878, N, Ur, Ur). Union Co.: G. D. Fuller 67h (Ur) -_— —_— eee Alma L. Moldenke "The Growth of Plants", by G. E. Fo ” ° ° eg 268 pp. Penguin Inc., Baltimore 11, Maryland. 1963, $1.68 paper back. ent elementary, modern, thorough, introduc text to fie field ot plant piysiclogy this book ist The oresniz®- tion unlike most standard texts in this field and is planned 1963 Moldenke, Book reviews 239 to emphasize the interrelations among processes, functions and structures from the atom to the organism in its complex environ- ons factors controlling a “! materials and with correla- ale of gener subject in an ascen ml difficulty and then lists seout a thi tl j f publication, out the author lists at the end of t bibliog easly the main sources for locating such papers, pe: oEb- Ological Abstracts, etc. he gost me effectively illustrated with plant photographs and diagrams wi 2 eee legends. The print is easily readable. In the introduction ¢ e survey work on cytology and histology could also be found i By Tar old —- A tissue is defined as " Joined togethase and algae as "a group which includes the sea~- weeds and their microscopic petal ay These are poor "slips" — a scientist of high caliber work and of ordinarily careful anguage use. It was wise to mention the scientific names of rhaste along wh their common ones; the latter often vary from Place to place or represent a whole genus instead of a single Species. Why the author used the spelling Lyriodendron on page 251 is not clear. These points of criticism are for minutiae; portant content is laid before the readers in excellent manner. u easy up~ Scientifically accurate "paper backs", could give a good founda- tion in biology. Tomorrow's studen have even greater ne for better trained classroom leadership. Ses fede opeape cet aa Isoetes", by Dominick J. ee Ft , Univers “Tilinois Press, Urbana, Illinois 1963. $2000" paper he one cloth bound. done with the precision This is New World, present day study at ster, Scans f man-hours of related past work done by the Old World and others, This newer work wisely omits 20 PUTT OLOGY: Vol. 9, no. executed on finely stippled drawings, but it is even more effect- ively illustrated by photography through an ortholux fluorescent microscope, by modern differential slide staining, by clearcut cell outline drawings, and by clear orthostichy graphs in the mathematical patterns of our day. Comparison studies of Isoetes howellii of different ages were made to reveal the nature and significance of sporophyte development especially. The literature in this field that covers a of a century we an s < nf sa, e ted and organized concisely. The author's main new contributions (1) a substitution of a group of apical initials in the shoot fram the time of first re- Cognition for Hofmeister's single enduring apical cell concept, (2) the ground meristem and the procambium developing from his logically undifferentiated tissues underlying the shoot apex, (3) regarding lateral and basal meristem as derived froma si cambium, ()) i produced to Side or the other of the basal meristem, vision for & termino’ t may avoid semantic difficulties. The publishe of this No, "Manual of Vascular Plants of Northeastern United States and Ad- ge Canada", by Henry Allen Gleason and arthur Cronquist, 10 pp., Van Nostrand Co., Princeton, N. J., 1963. $11.75 S is a portable-to-field. -to-c sroom = se Sation of the 1952 three volume reference work of the senior autir wo ew ton ated Flora". Any ions 48 + author. ~ book encompasses much of the same area covered by Fernald's "Gray's Manual of Botany", edition 8 (1950), but with more com ous keys, with the introduction of tyP®, ving abbreviations, and with the regrettable if erns only. "Re" could have been used as a space Saving substitute for the frequently used "descriptive of". It 1963 Moldenke, Book reviews 21 is also most regrettable that recommended common names are omit- hi of how taxonomists may shift a given taxon from one genu ther. The flexible cover is handy for field work. It is eae that the back binding will hold up over the years of handling like those of the last few editions of the Gary's "Manual" did. The but clear print is a necessary concomitant to keeping the book portable. Bt? a technical work that will become a "bible" for many the xt has a surprising mumber of printing errors not checked on galley proof. The misspellings of ercer aa and Cleroden- drum on pages xxvii and 582 are examples. For mi tings af- fecting scientific terms there should be a i a list sup- Plied with all copies of the book sold in the future. Cc + s done uniformly by zoologists and cpelecehelow wise and all sles ag names (like Lablatae and Gramineae) are not given the uniform eae ending The now=in-vogue St tet of a specific epithet as a S$ a varietal name prodices In ludicrous tautonyms and Gertrude Steinian poly- , for instance, Solidago rugosa rugosa rugosa OM P. 715. Would. not the designation of a logotype for quences rubra = ee the rejection of this well know name as a "nomen am long know escapes from cultivation as Acer palmatum Parthe- nocissus tricuspidata, and Platams acerifolia. A few species, old Polypodiaceae into smaller f es? The evidence presen by Canadian sts that our common northern pipewort is genet~ oe and phenotypically distinct from the eptangulare has been ored. Such a tremendous peo of careful and valuable scientific work has gone into the production of this book and its volumes, which will be used by so mamy pers to come, that it seems sad to that the authors still follow the outm stem of classification, rather than the the authors’ Englerian far more realistic Hutchinsonian. In any event, dS efforts will be appreciated by many users of these books years to come. NOTES ON BROMELIACEAE, XX Lyman B. Smith COSTA RICA - Smith, sp. baie Say Mez, cui habitu valde imulans, bracteis flo- eostae eubsuplo majoribus angustioribus yekls subaequantibus, pps paar onge Sige agg entre shes 2 ly but inconspicuously appressed-lepidote, reddish cope benessty Sen above; scape straight, stout, glabrous; scape-bracts dens completely covering the scape, broadly ellip- tic, apiculate, purpl, on covering apex spr med Fi inflorescence densely bipinnate, slenderly fusiform, 32 cm. long, 8 cm. in di- ameter, glabrous; primary bracts s broadly ovate, apiculate, slightly but consistently shorter than the branches, subcoria- ceous when dry, probably subfleshy in life; bran suberect ed iaxly few-flowered with ¢ terminal aT quite equaling the sepals toward apex; pedicels rather sle ender, to 8 mm, long; flowers not secund; sepals narrowly e 8 oa pitas, 24 mm. long, 10 mm. wide, coriaceous = base; stamens in ed. Pl, z ranch x 1/2; og % 3 Sepal x COST. CA: Rare in cloud f —— of El Alto de la Palma, alt. 1500-1550 m., C. K, Horich fhe type). Cultivated and flowe in the University of Californi Beooie Garden, Septenbe ap te accession — r 58, jee. The ectlecter report, ~ Species of frogs live and apparently breed in these "pls 4 AMAZONIAN SOUTH AMERICA NEOREGELIA to thane a eee orations in the upper Amazon Basin have brought r of new species of Neoregelia that fortify Ule's Soncept of a subgeneric group. They all in having & slender hat merges wi he the 1963 Smith, Notes on Bromeliaceae 2h3 1. Leaf-blades ay pt pedicels 15 mm, long; inflorescence com- pound, many-flowered; sepals free, 21-26 mm. long. . N. eleutheropetala 1. gS gaa narrowly triangular, long-attermate; pedicels 2. Sepals free; inflorescence compound. 3. Sepals glabrous; leaves uniform...........2. N. aS 3. Sepals lepidote; inner leaves basally pink...3. N. wurdackii 2. Sepals connate for 2-7 mm.; inflorescence compound or simple. + Leaf-blades 5 mm, wide; sepals connate for 5-6 m. 4. » pendula 4. hal pec 10-30 mm. wide. 5. Sepals 18-32 mm, long; inflorescence compound; leaf-blades 5-L0 mn. wide 6. Sepals 32 mn. long, lepidote; leaf-blades much pene than the sheaths AF rosea 6. Sepals 18 mm, Lene 2 Pha eeinacet leaf~blades moetiy’ short- er than the s -6. N. peruviana 5. Sepals 10-13 m. peor inflorescence simple. 7. Stolons 8 mm, thick; leaves numerous in each rosette, the sheaths ca. 15 cm. lo OnE blades ae inflo- rescence few-flowe Sey mooreana 7. Stolons 4 mm. thick; pene about 5 in each SE IR the sheaths 3-5 cme long; blades reeroneesutoe: inflores- cence 2-flowe: . stolonifera 1, NEOREGELIA ELEUTHEROPETALA (Ule) L. B. Smith, Contr. Gray Herb. 104:78. 1934. la. NEO NEOREGELIA THEROPETALA . ELEVTHEROPETALA ‘isvlarius ee le, Verh. Bot. Ver. Brand. 48: bregolia eleu ala Mez ex L. B. Smith, Contr. Gray Herb. 98:5, pl. = fin. 4-6. 1932. Leaf-blades essentially concolorous. , COLOMBIA, PERU, lb. NEOREGELIA ELEUTHEROPETALA var. BICOLOR L. B. Smith, Phytolo- gia 8:227. 1962. Leaf-blades strongly BLealeres } because of the dense minute Sa: scales on the under surface 2. NEOREGELIA MYRMECOPHILA (Ule) L. B. Smith, Smithsonian Misc. Coll. 126:30. 1955. Miditaviae ayremnoridins Ule ex Karst. & Schenk, euing ai bild 3: under pl. 3, 4. 1906; Verh. Bot. Ver. 48: 132. i 1 plone hila Mez, Engler Pflanzenreich IV, 32:52. 1934. Additional material may indicate stronger division between 2h PHYTOLOGIA Vol. 9, no. Ule's two forms, latifolia and angustifolia. 3. NEOREGELIA WURDACKII L. B, Smith, sp. nov AN. myrmecophila (Ule) L. B. Smith, cui affinis, sepalis le- pidotis, foliis interioribus basi roseis differt Plant epiphytic, stoloniferous; stolons flattened, 11 m. wide; leaves about 15 in a funnelform rosette, to 8 dm. long, subdensely appressed-lepidote with brown-centered ng a the in- ner ones pink toward base; sheaths suborbicular, ca. 8 om. long, apex; blades linear, sivediale: is mi. wide, sp 1.5 mm midrib, 25 mn. “long, sparsely Léphhate sepals Pro stro asymmetric, 14, mm. long including the 2 mm, long blunt mucro, © green, sparsely lepidote, the cape geal ones carinate with the keels decurrent t 3 pet white, the als » naked, blades acuminate; ovary subcylindric, mesharine *with the short in- distinct pedicel. Pl. I, fig. 3: Inflorescence x 1/2; fig. 4 Floral bract xl; fie. ch Sepal x 1 : AMAZONAS: rin B. grormocse het aya na Se . ss Sea rt affinis, sepalis basi siti foliorum laminis angus tioribus differt. Srp epiphytic, stolo pra as stolons elongate, pendent, 2- ‘4 - in er; leaves about 15 in an erect ellipsoid rosette 4 olon, sparsely lepidote soon becoming glabrous; ecieraes broadly ovate, -5 om of cen ent linear, attenuate, 5 mm, wide, f broad flat spreading eae » Simple, 1 om. in diameter; floral bracts elliptic, the ovary, pale, membranaceous, ote; sepals strongly asymmetric broadly acute, 21 mm. Jeng, comate Te for 5-6 m., glabrous; ovary cylindric, merging in rd very short pedicel; seeds few, ellipsoid, 5 mm. long, epalis purpureis. wines + long; eras bred green; sepals dark pur~ a es » Bagua: Low epiphyte, rainforest along see 5 ; in. “above mouth, alt. raven 8-13, 1962, J ee aoe k 2166 (US, tyee). 250-360 m. 1963 Smith, Notes on Bromeliaceae 245 4b. NEOREGELIA PENDULA var. BREVIFOLIA L. B. Smith, var. nov. Foliorum laminis eetinies sepalis viridibus. Leaves not over 18 cm. tong; H OC ote vo, ed tas except near apex; sepals green; petals white (! PERU: LORETO: Prov, Alto Amazonas: ee iphyte, rainforest m, 00-550 m., October 19-21, 1962, J. J. Wurdack 2345 (US, type; “vsM). 5. NEOREGELIA ROSEA L. B. Smith, sp. AN. mooreana L. B. Smith, cul arttnbe, foliis utrinque den- sissime cinereo-lepidotis, inflorescentia composita, sepalis sub- duplo majoribus differt. Plant epiphytic; leaves rosulate, many, 5-6 dm. long, covered n pressed browm- ° h sides with ap b ~centered cinereous .scales; sheaths elliptic, ample, 18 cm. long, dark castaneou cept near apex; blades very narrowly t ar, attenuate, 25-40 mm, wide, laxly serrate with antrorse dark 3 florescence the center of the rosette, c owered, 5 cm. diameter; pr broadly elliptic, m- Tonate, 5-6 cm. long, exceeded by the sepals, subd ly a s str for 6-7 mm., lepidote; ovary slenderly cylindric, 32 ma. long, merging with the short obscure pedicel. Pl. I, fig. 8: Inflores- cence x 1/2; fig. 9: xl. PERU: LORETO: Pro rate Amazonas: High epiphyte, rainforest = — north Anes of Cerros Campanquiz at Pongo de Manseri- °. o Maranon, aa 550-750 m., October 22, 1963, "3 - J, Wurdack 2389 (US type). 6. NBOREGELIA IA PERUVIANA L. B. Smith, sp. AN. mooreana L. B. th, cui affinis, foTidees laminis vagi- nas subaequant ibus vel brevioribus = inflorescentia prefameann Dy aa Palis majoribus glabrisque differt. Plant epiphytic, stoloniferous; stolons § mm, in diameter; leaves many in a broadly funnelform rosette, ca. 15 cm. aes ei covered with appress nereous scales 8 sheaths elliptic, ample, mostly longer than the blades, vontire except at apex; blades suberect, tri , attemuate, pungent, ma, wide, flat, repand-serrate with broad flat spines 4 m. a inflorescence sunk in the center of the rosette, depauper— ately d, rather few-flowered; primary bracts seit ? Bm, long x 1/2; fig. 11: Inflorescence x 1/2; fig. 12: Floral bract 2h6 PHYTOLOGIA Vol. 9, no. flower x 1 PERU: LORETO: Prov. Alto Amazonas: High e peste locally fre- et rainforest on i ieer northwest slopes of Cerros Campanquiz, Maranon just so $e Pongo de Manseriche, alt. 350-350 m., 0C- tober 17, 1962, J. J. Wurdack 2320 (US, type; USM). 7. NEOREGELIA MOOREANA L. B. Smith, Phytologia 8:227, pl. 2, fig. 6. 1962; Graf, Exotica ed. 3. 1659. 1963. Neore elis ossifragi Hort. ex L. B. Smith, Phytologia 8:227. 3; Graf, Exotica ed. 3, 441. 1963. Nomen. 8. a wagers ay cetera L. B. Smith, sp. nov. ss » mooreana L. B, Smith, cui affinis, stolonibus gracilibus, rolls oe =F * “scints retrorsis, inflorescentia biflora ylindric, merging Sepal obscure pedicel, Pi. I, fig. 13: Habi: it x 1/2; fig. 14: ie =, : LORETO: Vicinity of Iquitos (7), 1963, Lee Moore 50 \'™s VENEZUELA TTCAIRNIA BREWERI L. B. Smith, sp. no AP. heliophila L, B. Smith, re yea affinis, foliis utrin- que glabris, sepalis acutis diffe. Z nt wholly glabrous at anthesis, Figmoing 2. 1m. high; iors slightly narrowed lot flat, very Ty serrate with slender antrorse spines 2 ™ foli scape erect, 6 mi. in eter; scape-bracts strict, sub- rrulate, all but the highest exceeding the inter parte “ + long, ecarinate; i 2 fig. 1: Branch of 23 vary 2/3 super x Te ie ovules alate. Pl. 2% ig 1963 Smith, Notes on Bromeliaceae 2h7 VENEZUELA: BOLIVAR: Cardona camp, near Chivapure, on the Rio Blanco near the headwaters of the Suapure and Parguazo, 5 km from Cerro Guanay, latitude 6 30' N., longitude 66 30! ae “alt. 400 m., August 4, 1962, C. Brewer 31 (VEN, type). COLOMBIA TA DELEONII L h, sp. anac omnibus speciebus gp toranlies sapaits longe mucronatis ert Stem] emles flowering shoot 25 em. high; leaves about 10 in a funnel-form rosette, over 5 dm. long, mich exceeding the inflo- not petiolate, 6 am. wide, sparse ge green except for a 3 scape ere 8 slender, covered with white sub scales; scape-bracts erect, densely in- green, white-lepidote, laxly serrulate; inflorescence simple, re Soe pet » 6 em. long, densely white-lepidote except the pe- oral bracts like the nae etn bay: about ad Mat ck Onger, thin, densely lepidote; inflorescence over . ? decurved, laxly bipinnate, except for the petals covered with - ferruginous stellate subappressed scales; axis ane < Pom Spikes; spikes , to 55 m. long including the short na- 6, su TiPicws tet vs mm, ena ng rae > nse AOA » flexuous, ral bracts broadly ovate ee to 15 mm. long, exceeding flowers, , ecarinate, membranaceous, prominently nerved 2h8 PHYTOLOGIA _ flowers not secund, subsessile; sepals feet meme b qQuadrate, cuspidate, 9 mm. long; petals barely ex eeding i se-~ pal » fig. 5: Primary psa and spike x 1/2; in 6: Sepal x Ls ECUADOR: COTOPAXI: Scrub (fire climax?), Pilalé, alt. 2200 m, August eg 1962, Mrs, Amy Jean Gilmartin 799 (US, type). PERU IA XIPHOLEPIS 1S L. B, Smith, sp. nov. AG. nubicola L. B. Smith, cujus bracteas primarias spicasque mi imitans, an florigeris acutis sepala superantibus dif owerin, tic, 25 cm. long, cm, ame densely and aay cppressed— (-} te into a tube, the A poi spreading, lan a a 3; stamens exnerted from the ae of aceite ne on, than the petals. Pl. II, fig. 7: Prima- ract and spike x 1/2: fig. 8: Sepal x 1 PERU: AMAZONAS: Prov . Bongarés Epiphytic 5-10 m, above ground, ocally frequent in moist forest, hills west-northwest vee ) of Pomacocha, alt. 2300-2700 m., June 19, 1962, J. J. Wur- Gack 917 (US, type; Usx), a CLAVATA L. B, Smith, sp. nov. nigra André, cujus feted imitans, foliorum petiolis integris, 1 ~lanceolatis utrinque Longe at~ ges serrulatis differt, ~caulescent, flowering 95 cm es leaves all alike, 1 dm. long; sheath nar Py Moy ut 15 _ long, entire, merging with the petiole, covered Sabon with : sed Scales; petiole 5-10 mn. wide, channeled, entire; blade e t: 6 cm. en laxly and finely 2 tenuate at both ends, flat, eae 1963 Smith, Notes on Bromeliaceae 29 3; inflorescence simple, nahh Gi ated » broadly ellipsoid, Re cms long, glabrous; floral bracts broadly ovate, triangular- da forming stame’ 2 inferior; ovules obtuse. Pl. II, ‘ie. 9: Tarleneseeass : "V2; fig. 10: Sepal x PERU: LO : Prov. Alto Ama zonas: Terrestrial, rainforest at upper end of Pongo de Manseriche, Rio Maranén, alt. 250 m., Octo- ber 26, 1962, J. J. Wurdack 2419 (US, type). PITCAIRNIA MELANOPODA L. B. Smith, sp. nov minimi em. hi em. long, entire; sheaths ovate, small, nearly black, densely in- bricate and forming a narrowly ovoid or subcylindric pseudobulb 4m6 cm. long; blades sessile, deciduous, linear, atteniets. 2 mm. wide, white-floccose at base, soon glabrous; scape erect, 1.5 mm. in ’ te-floceulose especially at the nodes; scape- bracts much shorter than the internodes, ovate, acuminate; ; inflo- rescence ng eg ee ublax, 7 cm. long, whit e-floceulose; | epete bracts like the nr scape-bracts, to 13 mm. long, — ea ne hich sepals; flowers divergent; pedicels prises 2 mm. long; sepals rohing: rounded and ee? 7 mm. long, eca- ; petals 12 mm. long, yellow, naked; ovary 4/5 superior; ovules not seen. Pl. II, fig. 11: ‘eat whale x = 25 fig. 12: PERU: AMAZONAS: Prov. Bongara: Serrett stale wath Jalca zone along nggrieccprerst bien gt 10 trail be Yanayacu and Pomaco- cha, alt. 2300-2400 m., June 26, 96a J. J. Wurdack ae Wurdack 1070 (US, FULGENS AB cergitiaee (as 0) boas: f., cui affinis, foliorum laminis longioribus angustio ribus supra dense lepidotis, floribus fe Flowering 6-11 dm. high; leaves many in a dense rosette, to 5 dm. long; sheaths seersie te? 25-30 mm. long, pale e toward base, dark us toward mead ntire, glabrous; blades very nar- rowly triangular, attenua > pungent, 10-15 mm. wide at base, glabrous beneath, covered te, Ponetth whitish subappressed scales, serrate with. subs straight slender brow 10 mn. blade of the scape; inflore pectinate-serrate, exposing much ape; cence cylindric, 3-5 dm. long, simple or pseudo-simple with short inconspicuous branches at base, lax except at apex, white-floccu- 250 BEY O0LOQGIA Vol. 9, no. k lose soon becoming glabrous; primary bracts like the upper scape- bracts, covering the few-flowered axillary branches; floral bracts of the main axis like the primary bracts but merely acumi- nate and wholly brown, about gupeting the sepals, floral bracts of the lateral branches b adly acute, entire, distinctly exceed- ed by the sepals; pedicels sleniee” 10 mm. long; flowers spread- 3 Sepals elliptic, oe 15 mm. long, me eee at apex; petals naked, 25 mm. , pale green. Pl. II, fig. 14: Leaf- base x 2s fig. 15: ay bract and flower x ipo fig. 16: PERU : taiveds : Prov. Chachapoyas: Terrestrial, abundant, sum mit of Cerro Malcabal (Cerro Tumbe) 3-6 km. sou outhwest of Molino- Pampa, sa 2850-2900 m., July 20, 1962, J. J. Wurdack 1458 (US, PUYA MARIAE L. B. Smith, sp. A P. isabellina Mez, cui affinis , foliorum laminis haud repan- 2 oe vs an © lepidotis, pedicellis elongatis, pets maj bd Florering 7-20 dm. high (! Brdeek); leaves 1 m. long; blades narro with wiy triangular, attermate, 5 om. wide, covered beneath fine appressed ciner es, laxly dactite’ with flat pale a trorse 5 mm. long spines; 3 Scape erect, 25 mm. diameter near middle, densely and fine ely white-lanate; scape—bracts erect, gream (1 Wardaak)e Pas eign ty eee nn ee ae sky tupe of Cae Sse Tal are fetta ect See Cultivated b; aa? Wonka ae araack 602 rent: type; USM). wa Eek sina aL. 3 “Ba th, eal inflorescentiam imitans, pedicellis similar Gensély whitish-lanate except the petals and the old primary bracts ovate, aeeitngte: ca. 5 cm. long, much 1963 Smith, Notes on Bromeliaceae 251 shorter than the usually naked sterile bases of the branches, dark brown, becoming nearly glabrous; branches spreading, to 3 dm. long, lax except at apex, 10 mm, in diameter; floral bracts broadly ovate, acute, 20-30 mm. long, about equa stou ore edicel; sepals broadly elliptic-oblong, broadly ro mm. ed, mm. » green. Pl. III, t apex, 40 mm. long; petals nak lon, fig. 1: Branch of intisttetaine x 1/4; fig. 2: Sepal x PERU: AS: Prov. Chachapoyas: Locally frequent, approach- ®s to Cerro Campanario north-northeast of Diosan, alt. ” 3200-3500 m., August 3, 1962, J. J. Wurdack 1600 (US, type; USM). = WURDACKII L. B. Smith nov. AP. pyramidata (R. & P. ry “Schult, f., cui affinis, foliorum laminis longioribus angustioribus subtus ate lepidotis, sepalis lineari-lanceolatis differt. Flowering 1m. high; leaves many in a dense spreading rosette, 3 dm. long; sheaths reniform, 5 cm. wide, glabrous, entire, pale slender brown 6 mm. long pie scape erect, slender, very sparsely uhite-stellates scape-bracts erect, exceeding the inter- nodes, the erg subfoliaceous, densely imbricate, the upper sub- orbicular w. ith a narrowly triangular Eee pepsin eee ee and sing peg Tg of the scape, thin-coriaceous, brom, sublustrous to » very sparsely Ty ante teak: gee rescence —? Ret oe: "20-35 cm - long, lax except at apex, sparsely and obscurely white- stellate; floral bracts like the upper scape~bracts, about equaling the sepals; pedicels slender, 10 mu, long; flowers completely vere ck after anthesis; sepals Lnear-lanesolate, Bet pant, 25 mm. long, white-lanate at apex; petals nak » long, green. Pe ree fig. 3: Floral bract and flower x a); “tig. ks: Sauk s PERU: AMAZONAS: Prov. Bongara: ene locally fre- quent , ee pny _along Yambrasbamba-Pomacocha trail between t ocha, alt. 2300-2400 m., June 26, 1962, J. J. Wurdack 1066 (us. "type USM). pwd CARNOSA L. B. Smith, ‘ rauhii L. B. Smith, cud ‘ili vaeesndl foliorum laminis pina » spicis longe stipitatis differt Flowering 1-2 m. high with the nutant inflorescence extended; leaves many in a dense rosette (! Wurdack), over 9 dm. long, co- vered throughout with appressed cinereous seales; sheaths ellip- tic, 25 cm. long, 14 cm. wide dark cast eene. blades ligulate, covering the scape; inflorescence pendent, a4 laxly bipinnate, (3 Wurdack) ramones glabrous on the outer sur- se aye 252 PHYTOLOGIA Vol. 9, no. ches spreading, to 55 cm. long, the sterile base about twice as long as the spike, laxly bracteate; spikes lance-oblong, 18 cm. long, 6 cm. wide, very densely 24-flowered, strongly complanate; rhachis nearly straight, stout, alate; floral bracts densely im- pe Leveptely Ret cial ches ptic, obtuse, to 5 cm. long, equaling or ex- sepals, ecarinate, fleshy (! ? Wurdaek) , stron, Rabeenatie we drying; flowers short-pedicellate, not at all al urp. tic, 22 mm, long; stamens and sty e slightly exserted at anthe- sis. Pl. TIT, Fig 5: Branch of inflorescence x 1/4; fig. 6: Sepal x 1. ERU: AMAZONAS: Prov + Chachapoyas: Abundant on cliff faces, rocky slopes of Cano Santa Lucia just east of Chachapoyas, alt. Al Se the subgenus Tillandsia, the petals are not tightly rolled into a tube as in that subgenus but are much more like those in the sub- genus Allard Allardtia. TILLANDSIA CUSPIDATA L. B. Smith, sp. nov. AT. tetrantha R. & P., cui affinis, ‘tne ovaaoustiad ramis 4 strictis bracteas primrias multo superantibus, bracteis sepalis -8 mm sl Sepals, thin, ecarinate but with a strong dark median pine eg all secund, spreading t ' 3 P inel luding : orescence x 1; fig. 8: peng ? Prov. Chachapoyas: Epiphyti c, summit o Maleabal (Ce rro Tumbe) 3-6 km. southwest of Molinopampa, ). 0-2900 m., July 20, 1962, J. J. Wurdack 1457 (US, type; vs biases DIFFUSA L. B, Smit th, sp. n I. pallidoflavens ens Mez, cud ks nt ene similans, scapi 1963 Smith, Notes on Bromeliaceae 253 bracteis remotis, inflorescentia densissime pallido-lepidota diffe Stemless, flowering over 1 m. high; leaves about 20 in a sub- bulbous rosette, to 32 cm. long, covered with fine appressed ci- nereous scales; sheaths broadly elliptic, 8 cm. long, dark casta- neous turning to purple at apex; blades subligulate, we Sar ame- in ter, cinereous-lepidote becoming glabrous with age; scape-bracts very tripinnate, over 5 dm. lo ong, > S eenbany cinarceeesl enacks ex- amply cept the petals; axis slender, slightly flexuous; primary bracts ut equaling the sterile bases of the branches, like the scape- diy Ovate, acute, 5 mm ts ong, about equaling the sepals at anthesis, even, ecarinate; flowers s spre » su paecaiis. sepals free, on ° y to section Faoudo-Catopets is (Contr. Gray Herb. 89:16-19), this species f to T. micrantha, a + much more closely poéublet t. vens because of its acuminate rn We oe , very lax non-flexuous spikes and stiffly spreading flowe a LAMINATA L. B. Smith AT. pectinata André, cui aliquid affinis, foliorum laminis angustissime triangula gularibus, scapi bracteis omnibus longissime Flowering 1.3 m. high; leaves many, over 1 m. long, co tliicaghtut sith fia6 ehnesly appieaees whitish’ goales; anakhe Ovate-elliptic, to 13 cm. long, dark castaneous; blades very nar~ rowly triangul imary bracts; axis slender, near red-tinged (1 Wurdack) ; lower primary bracts like dea oranda cts with long blades os ceeding the axillary braces - "the upper bladeless, much Seronen “7 and scarcely different from floral bracts; branches su all but the uppermost asad near the base, to 9 cm, long; se- 254 PHYTOLOGTIA Vol. 9, no. a bracts like the floral bracts: 3; spikes lax at base, sub- dense toward apex, about 10-flowered; rhachis flexuous » strongly m. long, convex, often bicarinate at base, strongly nerved; pedicels stout, angled, ca. 7 mm. long; flowers suberect ir imes remai 80; sepals free, asymmetric, lo: e when dry; capsule slenderly fusiform, 2 cm. long, see ae fig. 11: Branch of inflorescence x 1/2; fig. 12: Sepal : AMAZONAS: Prov, vane pores Epiphytic, scrub forest a- long igre Ventilla 1-2 km, west of Molinopampa, alt. 2350-2100 m., July 23-25, 1962, J. J. Wurdack 1502 (US, type; USM). TILLANDSIA WAGNERIANA L. B. Smith, sp. nov famaleana E, Morr., cui valde affinis » bracteis florige- ris rosels apice valde incurvatis, petalis minoribus diffe ert. owe bf a pies ovate, acuminate 5 dulopabcekon laxly bipinnate, inear-lanceolate, acute, to 10 pei tee 2 cm. wide, strongly complanate, dense ely many-flowered or ope at base; floral bracts ov ~connate, lan aieecckt acute, 21 mm. * long: aid seg be G Naciay the blades ae broadly elliptic, 9 ong; stamens s deeply included, exceeding the pistil. Pl. hy fie. 1: Spike x x 1/2; fig. 2: Sepal x 1; fig. 3: Petal and st PERU: Vicinity a Tquitos Lee Moore 310 (US, type, cultivated by David Barry, J wle. ih the rac request of Lee Moore, this species is dedicated to the memo memory of his friend, Ronald Thonae Wagner. TA WURDACKII L. B. Smith, sp. nov. I. orbicularis L, B, Smith, cui verisimiliter affinis, in- thorescentia laxa, spicis patentibus eutmilt iflorie bracteas pri- marias plerumque superantil a dir » obs ly the erect densely imbricate subfoliaceous racts; inflorescence laxly bipinnate, about as long as the 1963 Smith, Notes on Bromeliaceae 255 scape, glabrous except for the blades of the lower primary bracts, deep red to scarlet except the petals (! Wurdack); axis pe- stout, nearly straight; lower primary bracts like the sc bracts » longer or shorter than the spikes, pair primary bracts suborbicular and apiculate, less than s long as the spikes; ’ paratype, : ene 3 stamens included, filaments straight. Pl. IV, fig. NAS: Prov. Chachapoyas: On trees, freqient, upper=- most attics and summit of Puma-urcu southeast of rng rd alt. 2900-3150 m., June 7, 1962, J. J. ae 801 (US, type; USM). On tree limbs, freqient, moist scrub forest on south side of Sasi Stiopanre-Divens pass, alt. 2700-3100 m., August 8, 1962, J. comes increasingly difficult and dangerous to eens novelties from the complicated maze of Tillandsia subgenus Andes such t ° riation in characters e and in on Poh length of primary bracts and spikes that Mez and I ked; spikes spreading. Costa oa Panama. var. costaricensis Sve =3 3. Sterile base of spike wholly wiaere spikes suberect. cuador eeeese . eeeene = 2. Spikes floriferous to base (or sometimes “the lowest sith a single bract) and without sterile bracts. 4. Spore spreading, rather many-flowered, mostly longer than rasa) inflorescence lax. Peru..T. 4 Spikes erect, -5-f1 , mostly shorter than the primary cts pad oid. Colombia sieht Saeesices densely ellips a ae orbicularis P oat 1. Sepals at least hal f-comate posteriorly. 5. Inflorese bracts shorter than the spikes. 256 PHYTTOLOGIA Vol. 9, no. k Colombia, sa ms Peru, Bolivia T. confinis 5. Inflorescence dens 6. Primary bracts “shorter than the spikes; inflorescence panes tate. Colom eecseeeete SucSCana 6. Primary Cech pe most of the spike 7. Lower primary bracts triangular-ovate, long-attenuat e. Ecuador. . confertiflora 7. Lower primary bracts broadly elliptic apiculate ° Caleta’, . strc enesiviri . dichrophylla VRIESEA TILLANDSIOIDES L. B. Smith A VY. harmsiana (L. B. Smith) L. e Pomith, cui valde affinis, spicis paucifloris fere omnino eccultis, bracteis Daria? den- pf onan tenuibus ad fa pong nervatis, sepalis minoribus f Flowering about 3 dm. high; leaves rosulate, many, 35 cm long, cinereous-lepidote throughout; sheaths broadly ovate, ee 7 em. long, kc 2) erence. blades very narrowly trian a Papi ey pats long, 15 mm, mide, densely 5-7-flowered; rhachis alate; fl bracts lanceolate, acute, to 23 mm. long, exceeding the sepals, sharply car inate, dens sely cinereous-lepidote, thin, nerved toward long, ecarinate, glabrous; petals tubular-erect ct, green, bearing 2 acute scales near base; stamens exserted, Pl. IV, fig. 6: Spike : PIURA: On trees, near Huancabamba, September 1956, N Rauh P-304 (US, type). Cultivated and flowered in Heidelberg, West Germany, in 1962, BOLIVIA T = L. B. Smith, Phytologia 4:218, pl. 2, figs- Tillandsia subtropicalis L. B logia 8:505, Pl. 3; dg aubtropicalte Be ae of the species of Tillandsia — SSardtia with bicolorous triangular leaf-blades (see je re dsia wurdackii above), I find that T. subtropical is mo more than a reduced form of T. confinis. BRAZIL Nee a ag TRIANGULARE Smith & Reitz, sp. no Surantiace E. Morr., cujus habitum Sen ctneee simlans, 1963 Smith, Notes on Bromeliaceae 257 foliorum laminis anguste triangularibus, scapi bracteis serrula- tis, co rbonmepeene wages “Tepidote differt Known from separate leaves and flowering shoot, plant over 3 dm, high; leaves 3 dm. long; sheaths elliptic, 14 cm. long, entire except at apex, dark purple except near base, sub- wears vestite with fine brown appressed scales; blades narrow. ar, attenuate, pungent, 3 cm. wide, cover th appress- ed brown-centered cinereous scales, becoming glabrous especially above; scape straight, 4 er, ferruginous-lepidote to scape—bracts ere olding the scape, elliptic, apiculate, se ferruginous-lepidote, the upper imbricate; inflorescence very densely se, 5 cm. in diameter inclu- subco ‘ ding the bracts, many-flowered; outer bracts suberect, broa ovate, exceeding the flowers, entire, red, subdens ew appressed- lepidote; floral bracts oblong-elliptic, acute, muc slightly exceeding the sepals, thin, obscurely Laihietes sepals free, asymmetric, oblong, truncate and mucronate a 14 mm. long, strongly nerved, obscurely lepidote; petals and stamens unknown; ovary ggg 3 placentae elongate; epigynous tube crateriform, 4mm. high, Pl. IV, fig. 8: A Apex 0 of leaf x 1/2; fig. 9: Inflo- rescence x 1/2; fig. 10: Se : ESPIRITO surto: ere , Castelo, Forno Grande, December 6, 1956, E. Pereira 2246 (HB, type). NIDULARIUM SEIDELII Smith & Reitz, sp. nov omnibus speciebus Nidularii scapo subnudo cum inflorescen- tia elongata differt. / Stemless, flowering 35-40 cm. Fae} leaves rather many form rosette, 5-6 dm. long, much exceeding the in- florescence; sheaths elliptic, 12 cm. long, entire, densely ves~- wee pn nearly naked, invade Porcigisode-liaeahe We becoming more or 8 glabrous; scape-bra cts 2, remote, the upper broadly ovate, os. 35 mm. long, serrulate, lepidote; inflorescence su beylindric, U-15 cn. long, 5 cm. wide; primary bracts suberect and densely c lowers, yell Vinate, few-flowered; floral bracts elliptic, acute, about ae ing the sepals, thin, entire, arog d acpagees flowers sessile, pale-ferruginous- -lanate toward base; sepals dly spatulate, ly as 5 1.5 mm. high; a al. oe fig xt ous and °) Em. centae apica - . : isfloreecmce = 1/2 fig. 12: Sepal x 1; fig. 13: Open corolla @ stamens x BRAZIL: SiO PAULO: Near Ubatuba, A. Seidel 6-20 (HB, type; US) 258 PHYTOLOGIA Vol. 9, no. Plate I 1963 Smith, Notes on Bromeliaceae 259 Plate II Fig. 2: 2 Pitcairnia breweri; fig- 3, 4: Ronnbergia deleonii; , 6: Tillandsia gilmartiniae; fig. 7, 8: 7 eeant: fig. = 10: eine * clavata; fig. nhl melanopoda; fig. 1h-16: Puya fulgens; fig. 17, 16: 260 PHYTOLOGIA Vol. 9, no. Plate III Fig. 1, 2: Puya ramosa; fi 6: 8- 3, 4: P. wurdackii; ie D3 Tillandsia carnosa; fig. ¥i-8 a: T. cuspidata; fig. 9 » 10: T. diffusa; fig. 11, 12: T, laminata. 1963 Smith, Notes on Bromeliaceae Plate IV 261 Fig, 1-3: Tillandsia wagneriana; fig. 4, 5: T- wurdackii 6, 7: Vriesea ioide mai fig. "9, 10: oa 7 tillands SYNOPSIS OF AMERICAN VELLOZIACEAE SUPPLEMENT III Lyman B, Smith Collections received from Dr. G. Pabst of the Herbarium Brade- anum and Dr. J. Murga Pires of the Universidade de Bra silia have yielded further noteworthy Velloziaceae as follows 6. VELLOZIA a epee = Mart. Fl. Bras. 3, pt. 1:75. 1847, end. L. B. tietseaktaacent,. pag or Seas leaves 20-30 on each branch, densely fasciculate; sheaths ovate, 10 mm. long, closely d ramin and coarsely nerved, st eous, lust rous; blades long-persiste linear, attenuate ely obtuse apex, 4 cm. long, 2 m. wide, conduplicate or splitting and ap ’ serrulate on th and beneath on the keel; glabrous; scape solitary, 25-40 mm. long, very slender, glabrous; ovary or campanulate, 6 mm. ’ re pals elliptic, obtuse, 18 mm. long, blue; stamens 6, 10 mm. long, O mm 1: wey at base, unappendaged. Fig. Plant; Cs ok Tepal and s amen, as Gerais: pean: da a Piedade, alt. 1800 m., March us a ae proceeding to emend a eoias without nits its in this description of aa sellovii Seubert. Furthermore, Sellow collect- ed in the area from which this later material came, so that it is not alee: that it is topotypic. + BARBACENIA Serer ey L. B. ert. B. saint oy Taub. C systema mea = sed sca~ heey brevioribus, antheris sinum ppm ee superantibus Caudex Lege ares in t leaves nu- ype but at least 15 mm. long; aa ous, densely massed, their order not evident; sheaths ovate, “ 5 aed ie very closely nerved, glabrous; blades linear, scum nate, 3 shen se @, closely nerved on both 8 : sparsely denticulate on kee] ‘ fe otherwise smooth and evolute; scape solitary, 7-9 cm. long, black stipitats glands especially near the age rs purple ({ Fed af L perianth-tube 10 mm, long; ovary ds; a = ostae about 20, covered with black stipitate a uae tube buliform, 4 mm. high; tepals su Pi pres 2 Gimme Long acute, 11 mm. long, the outer narrower; apices; 1963 Smith, American Velloziaceae 263 base of the filament-sinus; style exceeded by the anthers; stig- mas linear, terminal. Fig. 3: Leaf-blade; fig. 4: Flower; fig. 5: Stamen (abaxial) ; fig. 6: Stamen (adaxial). BRAZIL: Minas Gerais: Serra do Caparao, near the two lakes, above the stone house, March 2, 1960, Flavia Térgo rt "CHB, type). 2la. BARBACENIA INVOLUCRATA L. B. Smith, Ste NOV. fulva Goeth. & Henr. in systema mea proxima sed ab omnibus speciebus involucro magno haud foliaceo aiftert. curved with age; scapes very short; flowers several fascicle, each almost completely covered by an involucre of large pe=b » brown ubcharta » glabrous; periant Slenderly subclavate, to 75 mm. long, slightly const tween the ovary an 15 mm, 1 epigynous tube, covered with ong all; ant hens linear, 16 mm. long, a equal the ay apex of the filament but Ag og ie well below its ber it Fig. a IL: Gerais: Serra “‘o Cipé, March 3, 1958, Heringer & Castellanos 594, (US, type: Univ. Brasilia). 50a. BARBACENIA GRAMINIFOLIA L. - Suit, P+ nov. tubo. > Lomentosa Mart. in systema proxima se ag aaviey? ss, aaallige guar ase angustioribus differt. A B. Goeth, & Henr., cui parum affinis, scapo altiore “alee: at bus minorius differt. us, densely rosulate; sheaths broadly re, Hse ag 2 cm. : . long Pals erect, oblong, broadly acute and apiculate, 17 mm Ds filaments cad Taare with short acute lobes, 2.5 mm. long in all; anthers linear, 11 mm, long. Fig. 11: Section of leaf-blade; fig. 12: Flower ; fig. 13: Tepal; fig. 14: Stamen anon, (adaxial - : BRAZIL: Minas Gerais: Serra do cips March 3, 1958, Heringer & Castellanos 5950 (Univ. Brasilia, type). 26h, PHYTOLOGIA Vol. 9, no. 4 American Velloziaceae Pig. 1, 2: Vellozia sellovii: f Barbacenia ionantha} tig. e-10: B. involuerata, let ii-14: B. Serene al coles Figure 10. Distribution of Verbena hastata in the United States Herbarium curators who have material of this species from additional counties are asked to seng it to the author for verification and record, so that future editions of this map may be more complete, Mapping by counties done by Andrew R. Moldenke . = me, FRC Ras Ss nr re . MAT. Pape i? a a SE a 1 uM I Te Gates aE’, — a. YANKTOR ek, Min ae ERO a c Teron) SO serra] 1 Swat Preece] CRESS CGN SNE Ce SO a eee recs | fumoneesteRSsnet mARDIN wonrce i CLEARIXPE COUNTY OUTLINE UNITED STATES Scale of Miles 100 a ar: ae {tenes 0 200 TP aeet ee MAP NO. 125 on Oe ; anERiCan wag dowrany, 1Nc . SECS SOLE PUBLISHERS \ CLEARTYPE MAPS Sei COLORPRINT MAPS SEE: map. The law prokibics the reproduce or copying of mma, or any ere! process for per ew, \ sonal use or resale, without —™. Copyright, American Map Co., Inc., New York, No. 14324 PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the éntire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary greatly. A volume will contain about 32 signatures, 512 pages, or a smaller number of with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be — = to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in. ; in zinc, or for diagrams, tables, or charts, provided they conform to certa limitations of size and proportion. An extra char ade for half- tones, oo on their size, as fixed by the engraver, with a minimum of ‘Mbout $2.25. dealing with research in all lines of botany, in any reasonable isi sketches, and critical reviews and summaries of litet- 2 re will be be considered for publication. Floristic lists, casual notes of am , ie ani to pe eae of popular type, and polemics will not be published. Advice ; a8 the sviaiiy of Manuscripts will be solicited, if necessary, from quali- fied | ‘botansts- PHY TOLOGIA Designed to expedite botanical publication — Vol. 9 November, 1963 No. 5 CONTENTS BANERJI, M. L., Studies on Indian Acers - Il. A new variety of Acer oblongum Wall. from India ...-+-++++++20007 ttt MOLDENKE, H. N., Notes on new and noteworthy plants. XXX VII.. 266 MOLDENKE, H. N., Materials toward a monograph of the genus Verhend NIN) ee eee 267 Published by Harold N. Moldenke aa pine + Ee eae : 15 Giogaek pie Ae oe Yookers 5, New Yok, U. =e oe —S | Price of this , umber, $1; per volume, $5 75, in adv “Volume 9, Ne in STUDIES ON INDIAN ACERS - II A new variety of Acer oblongum Wall. from India. « Le Banerji University of thekvadic Kalyani, India. Pax (1902) has mentioned five varieties of Acer oblongum Wall. of which var. glaucum Graf c. Schwerin is the most common and has a wide range of Seer bates being from Kashmir to Central China. Another Indian variety is var. microcarpum Hiern, which is based on Griffith 935, " eartacted from Mishmi Hills. Two other varieties, latialatum and concolor, are eeaiizéted to Hupeh in Central China, we the tifth, var. horizontale, is based on a cultivated plant in the botaedaalt garden at Florenz. The var rietal differences, designated by Pax are the leaf and fruit characters. The widely distributed var. glaucum, which has leaves that are glabrous on the upper surface and slaucous beneath, has been i nt The and at its Paap point it measures 9 to 12 m. The var. micro- carpun has fruit wings just 1 cm. long, with a width of h—5 mm. ~~ None of ae five varieties mentioned by Pax has thin membran- ous leaves, A specimen collected from Jharipani (Ifusso orie) has characteristically membranous leaves, and also the size of the alae of the fruit is shorter than that of var. glaucum. Material in the Central National Herbarium, Calcutta (CAL.) and in the Forest Herbarium, Dehra Dun (DD.) have been examined. In the Cen- These specimens are floriferous and the floral structures are actly like those of var. glaucum. It has, thus, been thought vest to describe this pices at with thin membranous leaves as a new variety under A. obl Acer oblongum Wall. in DC. pbk, LEU: _ Hiern in Hook. . Fl. Brit. Ind. ie 693, 1875; Pax in Engler Bot. Jahrb. VII: 1886; et pera. @ 1902. (=A. lauri Pie Eig . Don, Prodr. Fl. Nepe 2h9, 1825, A. buzimbala Buch.—Ham « ex D. Don, loc. cit i Yor var. membranaceum var. no Accedit ad var. glancum aaa in notis, ab eo tamen dif- fert foliis membranaceis oe ae peal alis fructuum minoribus Rempe. 2 se cm. longis, 9--12 mm. ad medium latis. ec ad Jharipani prope oo in montibus Hin- alaicis aditentatetien ab oped ee Gollegii Meerut die 29 2 266 PHYTOLOGIA Vol. 9, no. 5 martii anni 1957, et positus in CAL sub accessionis nume 331441; paratypi, Mckinnon s.n. lectus die 15 aprilis pas 1896 ad Mussoorie, et Mckinnon s.n. lectus die 3 aprilis anni 1898 eodemque in loco, servati in CAL, primus sine accessionis numero, alter vero sub accessionis numero 62A. Acer oblongum is an evergreen tree about 12—15 m. tall with esting Rigen long-pointed leaves. Flowers in hairy seed ak appear f. February to April and fruits from “Apes to July in the North W Western Himalayas. Flowering and fruiting is late in tern Himalayas. Distributed in the temperate Himalaya, ae m. (2,000-—-6,000 feet) above sea leve his new variety approaches the typical variety glaucum in nost of the characters but clearly differs in having thin the fruit wings, which are 2.3~--2.5 cm. long and in the mi 12 mm . wide. The au uthor is deeply indebted to Rev. Fr. H. tapau, Direc- tor of the Botanical Survey of India, for the Latan translation of the diagnosis of the new variety and for going through the paper and improving its quality. NOTES ON NEW AND NOTEWORTHY PLANTS. XXXVIII Harold N. Moldenke age segpiaonr LONGICAULIS var. GLABER Moldenke, var. nov. a forma typica speciei foliis vaginisqe oncogenes laberrimis recedit. mr variety differs from the typical form of the species — its peqiees sheaths, and aneiee completely glabrous. of aitues Herbarium at Yonkers, New Y ALYX MICRANTHUS f. ALBUS Moldenke, f. is Bie forma a forma typica speciei floribus Heiss alb This form differs from the typical form of the species in a its flowers and fruits, including the membranous wings, of the forn fs consented by Andrew Ralph sag re Sie: a5 on the aa + Bingham, Socorro County, N August 1, — the H. Ne ey denke ete at oe oa asi = 2 cans in the MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XIII Harold N. Moldenke VERBENA HASTA’ pret oliegg ee ey ILLINOIS: Union Co.: Fuller & Fisher 299 (I1--15881), 674 (I1—15900). Vermilion Co.: G. N. Jones 13010 (Ur); Storm sen. [July 28, 199] (Ur, Ur, Ur). Wabash Co.: Ahles 4637 (Ur); Schneck s.n. ore 6, 1888] (Ur), s.n. [Aug. 12, 1889) (Ur), sn. [July 5, 1900] (Ur), s.n. [July 1900] (Ur); Shearer s.n. [Mt. Carmel, July 20, 1900] 10] (Vi). Wayne Co.: L. Car ter 111) (11--2)9)3); M. Walker s.n. [Sept. 10, 1949] (Ur); Win- terringer 3248 3248 (I1—-28508). White Co.: Winterringer 3628 (I1— 28831), 3645 (I11--288)2), 3755 (I1—291h2), 7775 (I1—37h18) Whiteside Co.: Ahles 1203 (Ur); V. H. Chase 12952 (Ur); Winter- ringer 292) (11—2779). Winnebago Co.: E. W. Fell 51,02 (I1— 38331). Stony Island: F. W. Johnson s.n- ~ [Stony Island, 7-19-13) (N). County undetermined: as - McCree 903 (Ur); Umbach 4059 [Wheat- land} (Ca—l1526). INDIANA? Allen Co.: C. C. Deam 10962 (Dm). Bartholomew Co.: Friesner 12973 (Bt--143). Blackford Co.: C. Ce Deam 181 (Dm). Boone Co. McCoy 5292 (Bt—52109). Brown Co: ae cD Deam am W145 (Dm); Friesner 1167 7 (Bw), sen. [Trevlac, 7/19/30) Bt--5099, Bt Bt--5100). Carroll Co.: Daily s.n. [Lake Freeman, aa (Bt—3226); C. C. Deam 49299 (Dm). ed Co.t riesner Au, Mi, Ob—201962, Ok, S). Clin : Ek s.ne TAT “te ceI0 1038) . rage ae Co. Ao escoag 12628 " (Be--1913) « Dubois Co,: C. C. Deam 52652 (Dm). “gikhart Gos: te C. Deam 20993 (Dm); Friesner 15608 ( (Bt—-5622). Fayette Co.: Knipe | Son. (In 2506). Floyd Co.: C. C. Deam 41551 (Dm); Friesner SNe gg Gheen Sn. (1n--231); ee 50s (In—10 018). Hamilton Co.: S.C. C. Deam 53038 (Dm); Friesner 116% (Bt—3799l). Hancock Co-: Friesner 18613 (N). Henry Co.: Black 36-7099 (Be— 14,7804). Howard Cov: Ek sen. [8-1-1939] (Bt—56802). Jasper Co: C. C- Deam (Dm); “Friesner 14565 (Bt—50399); Welch 12h (Dp)- Jefferson 00.: ce C. Deam 50850 (Dm). Jennings Co.: C. C- C. Deam = pa eoy Li? (Dp--l672). Knox Co.t C« Ce Deam 29287 (Dm); S- Modoy Be ee tan, Kosciusko Co.: C. C. Deam ; 384 (Dm eds ese Sty (Pu), 684 (Pu); Friesner 12659 (Bt--l1421); M. Morris (Ba); Templeton 115 (Pu); Umbach ach 1145 (Ka—621)20) cyst Jo COet 7 Cas 268 PHYTOLOGIA Vol. 9, no. 5 C. C. Deam 32480 (Dm); Friesner 12119 (Bt—l1866). Lake Co.: C. C. Deam 168 (Dm); F. W. Johnson 391 (N, W—1653653) 5 Lansing — 2805 (Up, Ur, W--753089). Laporte Go.: C. C. Deam 7063 ; (Dm), 21126 (Dm); Friesner 1380 (Bt—L7129, B Bt--7140), 14200 wae 7725). Lawrence Co.: Kriebel 953 (Bt--19476). Madison Co Friesner 1708 (Bt--62701); S. Nc McCoy 2237 (Dp--13913) - Marion Coes H. H. Bartlett s.n. [July 20, , 1903] (Mi). Marshall Co: C. Ce Deam 34733 (Dm); Evermann Ths (W--35759); Scovell & Clark k 7h (56) (Du--24181). Miami Co.: es Ek sen. [7-23-1936] (Bt—h2532), Se n. [8-16-1936] (Bt~-2531). Monroe Go.: C. C. Deam 1459 (Dm); Friesner 3200 (Bt--8706); Gullian s.n, ([Ellettsville] e] (In—2%); Lipps s. son. [July 17, 1941] (Ok). Montgomery Co.: Grimes 2h2 (Dp), 627 (Dp). Newton Co.: C. C. Deam 21478 (Dm), 58511 (Dm, N, N, N, N)5 Friesner 11669 (Bt--50638); Loughridge 1365 (Bt--17892); M. McKee 1329 (Dp). Noble Co.: 638) Lovehridee 1355 ee C. Deam s.n- ney oi, 21, 190] (Dm); eS jer 8963 (Bt—-27356) site. Go G. Ga 52662 (Dm). Porter Co.: C. CG. Deam 39518 (In); Friesner e- Bi00 (Bt- 16551). Pulaski Co.: ve C. Deam 46373 (in); Friesner 352 (Bt— 7742), 9768 (B1l--4.2323, Ca--597262, Cm, Gu—1h737, H--u1100, lo—- 146962, La, N, Pl--83176, St--92h9, Ur, We). Putnam Co.: ae a (Bt—67692) ; ; Geihee 65, (Dp). Randolph Cost Ce & (N). Scott Co.: C. C. Deam 34496 (In). Shelby | a Deam 13711 (In); Peieehi g SoBe [Morristown, 8/12/30] 00 Seat, Bt—5052). Spencer Cos: C. Deam 37454 (In). Starke Co.: Ce Ce Deam 21100 (In); Pelaenay “10012 (Bt—3302h, Bt--33$25) . Steuben Cow: G.-6y C. Deam s.n. [August 11, 1903] (In); ig 15680 (Bt— 54759). Sullivan Co.: Friesner 661 (Bt--154)1); S. eee Th Coy 1265 (Dp——673), B B.26 pet aeeey Switzerland Co.: C. Cc. Deam 5814) 5eihs (N). et gece Co.: Dorner 306 (Pu). Tipton Cot Se i Dean 13909 (In), san. [C. C. Deam 13623] (Pu). Union Co.: McDougal 8+ Ri ae ete (Dp). Vanderburgh Co.: C. CG. Deam 1677 (Dm). ViE° Co.: Friesner 6667 (Bt—1),60). Wabash Go.: Fri Friesner 16018 (Bt— 53180). Wayne Co.: Aakick 167 (Ea); M. E. Chambers ambers 153 3 (Ea)5 Greek 157 (Ea), 159 (Ba); H. Hudson 16, (Ea); L. J. King 1690 (Ea), 203 (8), 25 2569 (Ea), 2570 (Ea), 2651 (Ea); Knipe sme jane (Ine 2505). Wells c er a C. Deam sen. [Sept. 13 13, 1896] ~(in)- White Cow: C. C. Dean 3936 39362 (In); | Friesner 3h)3 (Bt—736). Whitley C+! CG. C. Deam 4909 (In). County undetermined: J. He an eng [King's Creek] ( (Ka). IOWA: Boone Co.: e uchanan, 3872 (Io——31763) ; aes & Macklin s.n. Fier: 1912] (ose 08, : Bode son. [Lanont, Aug. 13, 1919] Saget Fy = ; ss Pomel 155 5 (io—is9622). Butler Co.: Whaley . [July “4, IsiBY (To (Go~138153) Cedar Go.: Fay 1126 (a-565063), 1320 (Vi) Crat: eh. [Clear . 1918] J (Io 95673), L. H. Pammel a (Io); — reba 1922] 1963 Moldenke, Monograph of Verbena 269 (N). Chickasaw Co.: Pammel & Spiker 605 (Io--118829); Rolfs s.n. (Io—15321). Clarke Co.: Pammel & Pammel s.n. [Osceola, Sept. 26, 192] (Io--11)368). Clay Go.: A. Hayden 3002 (Io—149003), 300k (Io—-149005), 3008 (Io—18886, N), 3011 (Io—1)8889), sen. [Aug- ust 7, 1934] (N). Clayton Co.: L. H. Pammel 624 (Io). Clinton Cow L. H. Pammel s.n. (Clinton, Sept, 10, 192] (Io—116730). Decatur Co.: J. P. Anderson s.n. [July 13, 1903] (Io—52106). Dickinson Co.: Shimek s.n, [Aug. 19, 1909] (N), sen. [Aug. 23, 1919] (Ur). Dubuque Co.: L. H. Pammel s.n. [Dubuque, 6/20/91] (Io—-15302)- Emet Co.: Cratty s.n. (Armstrong, August] (Io—-92223); Pammel & Fisk s.n. [Dolliver, July 6, 1926] (Io—12622h). Fayette Co,: Fink 185 (July 19, 189] (W--21)903), 185 [Aug. 16, 189h] (W-— 219903), son. [Fayette, Aug. 1893] (Io—28698), s.n. [Fayette, July 19, 1899] (Io—26815). Floyd Co.: Arthur s.n. [Charles City, July 6, 1881] (Ah), s.n. [Charles City, 5/15/81] (Ah) H. N. Mol- denke 7663 (N). Guthrie Co.: Fay 3629 (Ca—5356); L- H. Panel 569 (Io—118719). Hamilton Go.: A. Hayden 10338 (Io--149577);5 Pammel & Zimmerman 30 (Io—118093). Hardin Co.: C. M. King sme [Steamboat Rock, 9-12-03] (Io—6231). Iowa Co.: Easterley 791 (We). Jasper Co.: Van Bruggen 1365 (N). Johnson Co.: P. Barker sn, [Iowa City, 7-25-29] (Au); Loufek s.n. [Lake Macbride, June 1938] (Io—176h7). Jones Co.: C. E. Bessey S-Ne [Monticello, July 10, 1876] (Ua—11385). Louisa Co.: R. A. Davidson 2725 (We 2235842). Mitchell Co.: Cratty sen. [Osage, July 17, 1918] (lo— 77816); F. M. Tuttle 229 (1o—77516)- Muscatine Co.: Estle & Brown s.n. [south of Muscatine, summer 1935] (Io—-146],29) ; Pam- mel, Kelsom, & Harlan s.n. [Jul. 20, 1919] (Io--950h2)3 Shimek Sen. (July 21, 1923] (Ur), sen. [Aug. 25, 1925] (N). Palo Alto Co.: Cratty san. [West Bend, Jul. 10, 1919] (Io—-96034); L. He Pammel s.n. [Lost Island, July 18, 1920] (Io—97806), S-n- (Jul. 18, 1920] (Io—97735). Poweshiek Co.: M. E. Jones SNe (Grinnell, August 1875] (Po—71001); Macy-Noyes 211 (Ok--10356). Scott Co.: Hull s.n. [New Liberty, Sept. 28, 1913] (Io--7h99h) 5 C- C- Parry 8.n. [Davenport, July 1886] (Io); E. A. Ross S-Me {Davenport, July 1890] (Ca~199030); Suksdorf s.n. [July 20, 138423), san. [Aug. 2, 187h] (Pl—130u23)- Sioux Ot be n. (Hull, 6-1895] (Io--20785). Story Co.: Beyer S-ne LANES, BO” ust 1 ; To—50263, 1o—84555), 137 888] (Io—-45075); P. Clayton 37 ( hee. 182} (Jo--18320), [ames, July 23, 1907] L. H. Pammel 78 (Io— (Ok—~10351 ne 1 Io—75051); L. H. Pamme” ge a ares ease ae $1,789); Pammel & Max- endry fe & 11 s.n. [Kelley, Sept. 1911] (Io—-84557) - T Cow: Pen ee 270 PHYTOLOGIA Vol. 9, no. 5 icarenaits July 1933] (Io—-140891). Wapello Co.: L. H. Pammel s. Eddyville, 9-16-12] (Io—76523). inch aioe aaa Goddard Sen. 1-20-95) (Io—-72079) ; Holway s.n. (Decorah, Aug 1879] (Io 38755); Tolstead s.n. [July 11, 1933] ig Si "Worth Co.: Cs S. Wallis s.n. [Aug. 3, 1950] (st), sen. [Aug. 21, 1950] (St). County undetermined: Cratty s.n. (13 July 1886] (Ob——50800) ; So- mes 3490 [Butler's Landing] (W—672273). KENTUCKY: Bath Co.: Ply- male s.n sen. [July 16, 1939] (Ky). Fayette Co.: C. W. Short s.n. (Pr). Henderson Co,: A. R. Moldenke 10 (Fg). Hopkins Co.: A. Re Moldenke 107 (Fg); Shacklette 620 (Bt——59 308, Ky). Laurel Co.: Peter s. $.n. 1. (Pittsburg, July] (Ky). Menifee Co.: F. T. McFarland 2314 (Ky); lcFarland & Lyle 09) (Ky). Powell Co.: F. T. licFar- land 410 (Ky, Bet <0). Oe Union Co.: Shacklette 531 (Ky, N). Coun- ty undetermined: - Short s.n. [180] (W--26618), sen. (18h2] (W—7718L8) , ooh eatery (Lu). TENNESSEE: McNairy Co.: S- M. Bain Ll (C). MICHIGAN: Alger Co.: E.G. Voss 2486 (Mi). Arenac Co: McVaugh oh (Mi). Berrien Co.: F. W. Johnson 1316 (N); Lansing 32)1 (W—753381); Merkle s.n. [August 1939] (Ok). Calhoun Cos: Potzger s.n. [Duck Lake] (In--11170, In—11171). Charlevoix Coe: J. He Ehlers 643 (Mi); Yuncker 1530 (Dp). Cheboygan Co.? Je He Ehlers 1626 1626 (Mi); Gates & Gates 9779 779 (Ur); Kofoid s.n. [Aug 25, 1890] (Ob=-50808) ; Swallen s.n Sen. (Douglas Lake, July ly 192] (W- ae a Co.: eas: J. Hermann ermann 7092 (Mi). Clinton Co. ras O (Mi). Eaton Co.: McClatchie s.n. (Olivet, July ll, EL} rey. Toumey s.n. [Grand Ledge, July 19, 1891] (Ca- 25146). Emmet Co,: J. He Ehlers 1195 (Mi); F. gs getss lal (Ka- 65927); Gleason & ae Ciaeee | 162 ( (Bt—3083, Dm, Go, N, We). J. Gogebic Co.: M. K. Cleme Son, [Ironwood, July 28, 09 Genasst) He Bailey sun. s.n. [Lans sig: Aug. 11, "yest (Ba); Lae Pe "eke sm [heric. College, 1890] (Po—63902); Cordley s.n. [East ‘st Lansing, 6-27-95] (Or—8860) ; E. B. Lake san. 17227 88] (Or--8866), Sts Baile S-n- [Hubbardston] (Mi). Jackson Co.: Collector undesignated s-M+ (Jackson, July 10, 1838] (ii). Kent Coe: He ie Bailey Sut (Grand ms a Aug. 15, 1891] (Mi); Bazuin 3626 (ii); Sones s.n- sn. [July » 1892] (PL—22627) Livingston Co.: Hamerstrom 192 (a) Bay Ray~ ack Co.: Allabach 9 (Cm); P Coe? Setabach 49 3 Potzger L553x (Bt--21680). Mason eeney, 3k ie 81 (NN). Menominee Co.: Grassl 2610 (Mi, * Oakland Co.: Ameel s.n. [Union Lake, Aug. 25, 1926] 2 Ug» ys neni Son. (Franklin, Sept. 22, 1918] (MMi) « Roscommon CO.: 1963 Moldenke, Monograph of Verbena 271 Dreisbach 5299 (Gg--319619). Saginaw Co.: Dreisbach s.n. (2h/7/ 1927] (Ew). Saint Clair Co.: C. K. Dodge s.n. [near Port Huron, 7-27-92] (Mi), son. [7/28/95] (Ob--87589), son. [Aug. 29, 1912] (Mi). Saint Joseph Co.: Hanes S.J.13 (Mi). Van Buren Co.: Lansing 3340 (W--753461). Washtenaw Co.: E. C. Almendinger s.n- {Ann Ar- bor, July 6, 1860] (Mi); D. Carter s.n. [July 1932] (Ok——10358) 5 Collector undesignated s.n. [Ann Arbor, Aug. 17, 1916] (Mi); Dod- der s.n. [July 1931] (Ok--10359); Frickelton s.n. [July 27, 1931] (Ok—-10357); Friesner s.n. [Ann Arbor, 7-20-18] (Bt--270), sen. [Ann Arbor, 8-13-16] (Bt--2l69); Gustafson s.n. [Ann Arbor, July 22, 1923] (Mi); Kauffman & Kanouse s.n. [July 29, 1923] (Ka-- 68880, Mi); C. D. LaRue sen. [Ann Arbor, July 17, 1915] (Mi), S- n. (Ann Arbor, Sept. , 1915] (Mi); Podolski s.n. [July 25, 1917] (Mi); Walfrole s.n. [Aug. 7, 1918] (Mi); Whited 105 (P1--22570) , 2105, (Or--12380). Wayne Co.: B. F. Chandler s.n. {near Trenton, Aug. 13, 191)] (W--983198); Foote s.n. [Detroit, July 10, 1060] (Mi); Gilaman s.n. (Br); Grassl 5905 (Mi); H. H. Rusby s.n- [De- troit, Aug. 198h) (Bz—23752, Se—14939); J» M. Sutton 689 (Mi); E, Wilkinson s.n. [W. Salem, July 31, 1887] (li). Sugar Island: F, J. Hermann 7251 (1). County undetermined: M. Craig 1207 (Mich. Agric. Coll.] (Ga--367052); Miller s.n. [Bass Lake, Auge 12, 1928] (Gu—E.1048); Umbach 7111 [Magician Lake] (Au). WISCO SIN: Ashland Co.: E. Beals S.N. [June 21, 1959] (Ws); BE. Me Christensen 3537 (Ws); Fassett 9534 (Ws); Ugent, Weber, & Zimmer- man s.n. [September 2, 1959] (Ws). Bayfield Co.: Cheney 14328 (Ws); Fassett 8161 (Ws), 816) (Ws); Horner S.ne [Sept. 5, 1955] (Ws); Shaughnessy & Shaughnessy s.n. [Oct. 3, 159] (Ws); Weber, Ugent, & Zimmerman s.n. [September 3, 1959] (Ws). Brown Co. De Byle s- n. [3 Aug. 1952] (Ws, Ws); Halloin 181 (Ws); T- S- Kellogg S-0e [De Pere, August 1, 1888] (W—-259280); P. O- Schallert 32 (H— 5470); Shuette s.n. [Green Bay, Aug. 4, 1887] (N)- Buffalo Co. Fassett & Hotchkiss 3320 (Ws); H. H. Smith 7069 (B). Burnett Co.: Fassett 6163 (Ws); M. Johnson sen. [Sept. 17, 1960] (Ws); W. T. McLaughlin s.n. [Sept. 1, 1929] (Ws); E. Newcomb sen. [Aug- ust 7, 1959] (Ws). Calumet Co.: Benke 386 (W—1521469); We Be Rogers s.n. [July 6, 1932] (Ws); A. Smith 86 (Ws). Chippewa COs Bjerke 8 (Ws); Te A. Ebert son. [10 July 1960] (Ws)- Lelchert ae ne eg Schmidt sm. [August 1, 1958] (Ws); J. Wickham 33 (Ws). Columbia Co.: Re Le Burgess 19 ~~ p. Chandler 38 (Ca—5180); Titis & Koeppen 8160 (Ws); H. L. Russell 100 (Ws). Crawford Co.: He He N- aa jan w: Ce Ge Albers 33018 (Au); Backus Se Smith 7478 (B, Ws). Dane Co.: Ce Ceo - Ds 8 a n. (Aug. 5] (Ws); N. L. Britton s.n- [Lake i (C)5 We. H. Chane son. [iiadison, 7/27/1877) (Ws), Sens [7/ 29/1877] 272 PHYTOLOGIA Vol. 9, no. 5 (Ws); H. K. Henry 101 (Up); Herb. Univ. Wisc. s.n. {Sun Prairi Aug. 29, 1893] (Ws); Hunter s.n. [Oct. 6] (Ws); G. N. Jones 17655 (Ur), 17712 (Ur); Lapham s.n. (Ws); L. He Pammel s.n. [Madison, Sept. 188)] (Io-=153):6) 5 R. F. Peters 3 lykb (Ws); A A. B. Seymour 1062 (H--107718, Ur); H. L. Smith s.n. [July 12, mee (Ws) 5 (Ws); Ugent sen. [July 7, 1960] (Ws); S 3. _W. Williamson s.n. [July SOT xk (Ws). Dodge Co.: Bubbert s.n. - [Auge 1928] (Ws); Iltis ; Koeppen 8557 (Ws, Ws); Martz s.n. [August 25, 1958] (Ws); J. W. W. Rhodes Rhodes 371 (is); Shinners 728 (Ws). Door Co.: E. Anderson s.n. [July y ib, 1956] (Ws); Lellinger 123 (Ur); McCabe s.n. [27 August 1945] (Ws). Douglas Co.: Thomson ¢ & . Jacobson 5345 (Ws). Dunn Co.: F. A. Bach ach- sen. (Aug. 9, 1928] (Ws); Bachmann & Patrick s.n. . [duly y 13, 1928] (Ws); Buss s.n. [July 29, 1940) (Ws); D. 5. Meyer 23 (Ws); Rossler s.n. [Aug. 7, 1928] (Ws); Weinzirl s.n. “Taug. 7, 1892] (Ws). Eau Claire Co.: Kunz 198 (Ws). Florence Co.: Riffle se {Aug. 1, 1960] (Ws). Fond du Lac Co.: P. Mueller s.n. [Sept. 25, 1960] (Ws); Parenteau s.n. [8/23/39] (Ws). Forest Co.t Re Ac 3° El- der s.n. [Argonne, 1936] 6] (Ws). Grant Co.: Fassett 14282 (Ws); He W. Graham s.n. [July 2h, 1928] (cm); Tntiey Long, Long, & Rudolph 13794 1379 (Ws); H. H. Smith 7680 (B, Ws); Wunderlin s.n, [July 11 “Ti, 1960) (Ws). Green Co.: E. W. Fell 58-86 (Ws), 38-59 (Ws); stunts st: 8 Me [Aug. 20, 1891] (Ws). Green Lake Co.: Kunz s Me [aug. 25, 92) (Ws); R. F. Peters liha (Ws, Ws); Ugent s.n. . [duly 1h, tens (Ws) + Iowa Co.: Berlin sen. {[Sept. 19, 1957] (Ws), sn. [July 30 , 1957] (Ws), sen. (Aug. 28, 1957] (Ws); Iltis & Noamesi 7068 (Ws); A. Me Peterson 228 (Ws); H. H. Smith 6885 (B (B, Ws). Jefferson Co.: Al~ berts s.n. [Sept. 10/5] (Ws); G. V. Burger 83 (Ws); Kuhlman $2: Taug. 26, 195h] (Ws); Loewenberg son. [17-1X.60] (Ws); Metzner 5« n. (Ft. Atkinson, July] (Ws); Schempf s.n. [July 25th '95] (Ws)- Juneau Co,: B. 0. Dodge s.n. (Sept. 1935] (N); Iitis & Noamesi 6931 (Ws, Ws); Lemke 129 (Ws); Wills s.n. [June 25, 1957] (Wis) « Kenosha Co.: Grace 34 (Ok). Kewaunee Go.: Iwen 156 (Ws). La Crosse Cos: T. Hartley su Sen. [Blue Lake marsh, 9/24/55) ~ (W=2261658) 3 a H. Pamel s.n. (Holman, Jul. 27, 1923] (Io—111)07); A+ M. Peter son = ih 325 (Ws). Lafayette Co.: Cheney s.n. [Auge 20» 1869) (lis); We 2 Ee sen, [Shullsburg, July 2, 1003] (N)s P+ Be Nelson s.n. [22 July 196] (Ws, Ws); Shinners 2356 (N, WS)3 5 Be Wagner sen. [8/8/59] (Ws). Lincoln Co.: Iwen 350 (Ws)5 peroutky & (Ws); F.C. Seymour 15))2 (Ws); “Re. Weber 38 (Ws) + = Co.: Iltis & Univ. Wise wise: Botany 130 Class 14210 etd athon Co.: Cheney 3082 (Ws); Schlising 706 (Ws); Schlist ie to (Ws). Marinette Co.: Tltis & % Rudolph 177% 77. 5 56 (Ws, Ws), 173 (Ws), 223 (Ws, Ws); Ware son. [August 17, 1956] (ie). Marquette Co.: Fassett 8795 (Ws); Hartwell s-ne [Ger 1963 Moldenke, Monograph of Verbena 273 mania] (Ws); Iltis, Bell, Melchert, Patman, & Witt 12361 (Ws); Iitis & Buckmann apa (Ws); Noamesi 6299 (Ws); Parenteau 32 (Ws). Milwaukee Co.: J J. S. Douglas s.n. [Wis. 3.) (Je—3158); thy, Bemers s.n. [17.VII.86] (Ws, Ws); Lapham s.n. [Milwaukee] (Ws); Strong s.n. (Wauwatosa, July 26, 1891] (Mi). Monroe Co.: A. M. Peterson 9 (Ws). Oconto Co.: Gillman s.n. [July 26, 1868] (Pr); E. E. Honey s.n. [Keshena, oy ch 19S] (Ws) 5 Schlising & Schlising 2 1017 Outaganie Co.: foetoel Sone [9 July '55] (Ws). Polk Co.: Mrs. C. F. Baker s.n, [June '99] (Po—6465k), son. [7-11-99] (Po—6L655); Benner & Benner 328 (Ws); M. Johnson s.n. en. [Aug. 15, 1960] (Ws), sen. [Aug. 26, 1960] (Ws); W Ws (Se McLaughlin 11h (Ws), son. (Aug. 26, 1929] (Ws); Re We Pohl 166 . (Ws). Portage Co.: Goder s.n sen. (Ju ly 1955] (Ws). Price Co.: Escoll 502 (B). Richland Co.: M. Me A. rosvere 336 (Ws); H. P. Hansen 502 2 (Ws); Iltis & Salaman . 9939 3 Tutton s.n. [Aug. 13, 1959] (Ws). Rock Coe: E. W. Fell ‘ell 57- 613 (Wis), 57-734 (Ws), 57-736, in part (Ws), 57-739 Ly — (Ws). Rusk Co.: Co.: Iltis, Iltis, & Noamesi 7880 (Ws). Saint Co.: W. T. McLaughlin 1)12 (Ws); N. Rus: N. Russell s.n. [July 19b8)} sw Sauk Co.: Iitis & Koeppen 11818 (Ws); Koeppen s.n. [August 1956] (Ws); Schaars s.n. [August 17, 1958] (Ws); He He Suith 7906 (B, Ws); Umbach 12273 (Ws); J. H. Zimmerman 11 L (Ws). Saw yer Co.: Ugent, Weber, & Zimmerman s.n. [September a 1959] (Ws). Shawano Co,: Hein 66 (Ws); E. EB. Honey sen. [July 11, 193k] (It); Melchert, sen. [9/3/58] (Ws), 3. [early July] (Ws). Sheboygan Co.: Goessl son. Sete 1919] (Ws); Kuhlman s.n. [Sept. 16, 195k] (Ws, Ws). Taylor Co . Anderson 139 (Ws) . Trempealeau Co. i- tis & Koeppen SS See Vernon Co.: Fassett 20234 (Ws); K. Ge Foote aon; (Ws); He 3; He P. Hansen 129 (Ws); > Melchert & rt & Witt sone Tsep. 20, 1958] (Ws); He He Smith 727h (Ws), 7366 (Ws); Stegner Sens [Sept. 25, 1960] (Ws). Vilas Co.: Potzger 8750 (Bt—60877, we); Stearns 85) (Ws). Walworth Co.: Almon zal ae ap July » 1926] (Ws); 1 $n. st 2 a ‘s); ] (Ws); Fernholz s.n. [Augu a eT ae ter 53 (Gg—31325); Kleeberger s.n. [Whitewa » Hlg-1 2) : 2 ssi 8162 (Ws); 332); Wadmond 17734 (Ky, S). Washburn Co.: W. T. McLaughlin 1413 (Ws), 1418 (Ws); H. Stephens S-n- n. [July 12, 197] (is). gre i Co.: Grotjan 182 (Ws), 186 (Ws Ws). Waukesha Cos: Gull 238 (Ws); Ehrler son. [Aug- 22, 1928] (Ws); Scimite S- ne (Sept. 1959] (Ws); Sheaffer s.n. [July 31, 1957] (Ws), a PRE CAngust 26, 1957] (Ws). Waupaca Co.: Melchert pe — (Hs) ; Rill "ye (Ws), SoM. {July 16, 1959] 59] (Ws); Ray pas BeBe (dine 23, 23, 1957] (Ws) Waushara Co.: Doll sen. (July *, DST Ws); Pochmann s.n. (Ws); Snell s.n. [August 17, 1957] (Ws). Win- nebago Co,: M. M. James s.n. n. [Aug. 21, 189)4] (Ws); Kaspar $-ne 274 PHYTOLOGIA Vol. 9, no. 5 [July 24, 1953] (Ws). Wood Co.: M. Johnson s.n. [July 23, 1960) (Ws); M. B. McMillan s.n. [Wisconsin Rapids, 1900] (Ws); Stolle sen. [Nekoosa, Aug. 3, 1957] (Ws). County undetermined: Barnston s.n. [Lake Superior to L. Winnipeg] (Ws); R. Sprague s-n. [1926] (Cn). Farmer's Island: Botany 162 Class s.n. [Sept. 27, 19h1] (Ws). Goose Island: Hartley 1375 (Ws) . Manitou Island: F. CG. Lane 2591 (Ws). MINNESOTA: Anoka Co.: Stoesz s.n. [1926] (Ob——70106) « Becker Co.: Buell 2019 (H—-71943, No—-8612); M. L. Grant 3073 (Ca—l38910, In, Io—139963, N, Ob--50796, Vt, W--187037); Moyle 732 (Ca--882651). Cass Co.: Pammel & Pammel 669 (Io--58202). learwater Co.: Deaver 5355 (Fg—8h55). crow Wing Co: E. B. Wate son s.n, [Brainerd, 1902] (Io—-l,5603). Douglas Co.: E. D. McDon~ ald Jr. sen. [July 11, 1940] (W--1888)85). Goodhue Co.: Sandberg Sen. [Red Wing, Aug. 1885] (B). Hennepin Co.: F. H. Burglehaus 861 (Se--14938); Daubenmire s.n, [8-1-33] (Bt—18l91); Mearns 526 (C); J. H. Sandberg sn. [Aug. 1889] (Ca—251h5, Ca--25150)- Sueur Co.: Shunk & Manning 132 (W—16232))3). Meeker Co.: He Le Dale s.n, [July 13, 1925] (Du--1509)9). Mower Co.: J. F. Brenckle 7-547 (N, Si). Nicollet Co.: C. A. Ballard s.n. [Swan Lake, July 1892] (S, W). Ottertail Co.: Chandonnet s.n. [July 11, 1911] (UP 60383); P. Johnson 545 (Bt--63915, Ky, N). Pine Co.: Lakela 1170 (P1--12989)). Polk Co.: Childs s.n. [Aug. 7, 1898] (Ob—-5083) - Ramsey Co.: Buell 723 (No—-0611); Mearns 526 (W--64,9507, W— 649508), son. [Fort Snelling, August 31, 1691] (W—670316)5 Rama = 351 (Bl--,2322). Rice Co.: G. Ke Knight sn. [Northfield, 8/ pe ots toate + Saint Louis Co.: C To—l9039); Lakela 2108 (ca—7027, H-—-560h6, N, Pl— 100355). Stearns Co.: J. 5 ee fig MGS July 1896] nona Co.: Holzinger s.n. [July 1901] (N). Yellow Medicine Co-! Ballou.s.n, [July 27, 192k] (Du—1)0432); J. F. Brenckle 1,608 (N). County undetermined: J. E. Campbell s.n. (Girard, July 1897] (Ob—50792); Roman s.n. [29/7/1893] (S). NORTH DAKOTA: Benson Cos: Iunell 516 (Ur), sen. [June 2hth, 1899] (Dt), sen. [July 1% tt Cos: W. B. Bell 1348 (Ca--58776 : Pp. Met- a2 B: Bel -- 3). McLean Co.: F. Fe Se calf 127 (R—1070875). Pembina Co.: Bergman 2256 (Cm). Richland 1963 Moldenke, Monograph of Verbena 275 (Gu-21830), sen. [Kindred, August 20, 19] agg i Rolette Co.: Wright 553 (N). Wild Rice Co.: Lee sen. [Wild 1891] (La). SOUTH DAKOTA: Brown Co.: Over 17223 (Ue230)3 Petrak sen. [16.VIII.1950] (S). Custer Co.: Barr sen. [July 25, 190] (Ba); Yuncker & Welch 4100 (Dp). Deuel Co.: Sanborn 5.n. {August 9, 1918] (Du--92151). Fall River Co.: Rydberg 933 (W—210965) . Grant Co.: Kienholz s.n. [Big Stone, July 1914] (Ur). Kingsbury Cos: J. J. Thornber s.n. batik a ig is (Ca—10)810). Law- rence Co.: F. Le Bennett 1093, in - Pennington Co.: F. L. Bennett 1093, in part (Bt--58323, hk en ee Roberts Co.: Corbett & Williams 232 [Herb. Corbett 345] (We); Over 1438) (W— 12h,2262)5 T. A. Williams 232 (Du—9551). Spink Co.: ot de Fe ~F. Brenckle — (Ba, pie N, Up), &7-532 (N), ace [Mellette, Aug. 1939) 39019 Petrak SNe ae () oe S); A. E. Ricksecker icaheseee Ob--50809, Ae 215260, tmz), 0 tl > Todd Co.: Tol- Stead s.n. [Aug. 1 ont h *(Zo~- 16166) « Tri : T. A. Williams 78 (D Du--9552) « Washabaugh Co.: Over 7079 Ghee - Se—1h935)- : Atchison Co.: Horr & McGregor s.n. [6/26/46] (Lw). Brown Co.: F. Garner s.n. {summer 1925] (Ka—72012). Chautauqua Co.: A. S. Hitchcock 5 sen. [Chautauqua Co., Aug. 8, bak he (Ka). a Co.: Re L. aay 3797 ol: Cloud Co.: S. V- Fraser 439 (Ka— 78939) . Comanche Co.: A. A. S. Hitchcock s.n. ae +. ee 17, 1896] (Ka). Crawford Co.: seas 6064 (Je—686); A. S. Hitchcock Sen. [Cherokee, Aug. 1892] (Ka); N. B. Jacobs 181 1 (Ka—897\1). Decatur Co.: A. S. Hitchcock s.n. Dense tur Co., June 2, 1897] (Ka). Doniphan Co.: Agrelius, Hall, Lovejoy, & Maroney sen. [8-7- 13] (Iw); Clothier & Whitford s.n. (oy, 31, 1897] (Ka). Douglas Co.: A. S. Hitchcock s.n. [Douglas Co., 8.7.99] (Ka); R- L. Mc Gregor 1570 (Hi--198622, W--2327196); Snow 1956 (Lm). Ellis Co.: E. Run Runyon 280 (Cm, Ok, Se--69270, St--9250, Vt). Ellsworth Co.: A. S. Hit Hitchcock s.n. (Ell swort rth Co., July 1895] (Ka). Finney Coes Sahat Sen. [Garden City, Oct. 10, '92] (Ka). ae Coes Re McGregor 12699 (W--2230)03). Harper Co.? A Ss. arper Co., Aug. ts 1896] oe Harvey Co.: Re L. MoGregor 371 (Iw). Jackson Co.: Norton & Clothier s.n. [July y kth, 1895] 1895] (Ka). Jefferson Co.: W. H. Horr 228 (N). Johnson Co.: Pellet s-n- {sum- mer 1890] (Ka). Kingman Co.: A. S. Hitchcock s.n- [Kingman Co., Aug. 23, 1896] (Ka). Lincoln Co.: A. S. Hitchcock s.n. [Lincoln ro July 1895] (Ka). Linn Co.: ¢ Clothier & meierord 8 sn. [Aug- 9, 97] (Ka). Logan Co.: Rydberg & Imler 1 1080, part (Lw). Meade *? A. S. Hitchcock s.n. [Meade eg sept. 1, B97] (Ka). Miami ches Clothier & Whitford s.n. [Aug- 8, 1897] (Ka) 5 oyster 6516 6516 (Herb. Prager 18625] (Ge—31318, Ge—31319) 5 Rohrer 9 (Ka—9566 8). Worris Co.: Westgate son. [Aug. h, 1898] (Ka). Neosho Co.: As S- Hitchcock s.n. [Neosho oe Aug. 1896] (Ka). Osage Co.: Clothier 276 PHYTOLOGIA Vol. 9, no. 5 & Whitford s.n. (Sept. 2, 1897] (Ka). Osborne Co.: Shear 202 (W— 227262). Ottawa Co.: A. S. Hitchcock s.n. [Ottawa Co., July 1695] (Ka). Pottawatomie Co.: Herb. Bot. Club s.n. [St. George, Aug. 1, 1893] (W—-2)h181). Pratt Co.: A. S. Hitchcock sen. [Pratt Co., Aug. 1896] (Ka). Rawlins Co.: As S. Hitchcock s.n. [Rawlins Co., June 1897] (Ka). Reno Co.: A. S. Hitchcock s.n. [Reno Co., Aug. 25, 1897] (Ka); R. L. McGregor 3931 (Iw). Republic Co.: J. B. S- Norton s.n. [Republic City, Sept. 23, '95] (Ka). Rice Co.: A. S- Hitchcock s.n. [Rice Co., August 1895] (Ka). Riley Co.: Kimball 5687 (Ka); re B. S. Norton 390 (Ka, N, W—353117)5 C. He Themen sen. (Manhattan, Aug. 9, 1892] (Ka). Rooks Co.: E. Bar n. [Aug. 3, 1891) (Ka). Saline Co.: Hancin 456 Gn7e792). Scott Coe: V. L. Harms 1107 (N); R. L. McGregor 2169 (Lw); Rydberg & Imler 1080, in part (Ka—-75039, N). Sedgwick Co.: T. L. ews sen. [Whichita] (o—28227) 5 aseiiee & Whitford s.n. [Aug. 2h, 1897] (Ka); Herb. Wichita Univ. 29 (Lw). Shawnee Co.: Maus S-N» {[Auburn, July 20, 1927] Ka 75015) 3 E. A. Popenoe s.ne Topeka, July 20, 1879] (Ka) » Sheridan Co.: C. Weber 35 35 (Ka—701L8) « Stafford Co.: A. S. Hitchcock s.n, [Stafford Co., Sept. 1, 1697] (Ka). Trego Coe: m S. Hitchcock s.n. [Trego Co., J (Ka). Woodson Co.: Clothier & Whitford son. [Aug. 29, 1897] (Ka)- Wyandotte Co.: K. K. Mackenzie s.n. [Rosedale, July 12, 1896 (Ka, We). MISSOURI: Callaway Co.: Schwab s.n. [July 192k] (Up) + Clinton Co.: C. R. Barnes s.n. [Converse, 8/,/77] (Pu). Clark Co.: Be Ee Bush 9165 (Po- (Po—127528); Steyermark 68897 (S). Greene Co.: P. C. Standley 9732 (W--6886)9). Jackson Co.: Co.: Be F. Bush 32321 roe Jasper Co.: E. J. Palmer 10568 (Bi, Du—205610, , 8) Johnson Co,: Steyermark 21,528 (Ky, N). Lincoln Co.: J. D Davis 9h (Ur). Marion Co.: J. Davis 3578 (Ur), FDL (Ur), XV (Ur), 5-ne (Hannibal, 8-7-13] (Au); R. Gmelin s.n. (Hannibal, Aug. 1879] (1991592). Piltte ch: RadloiF 3k [July 26, 1956] (Ur). Ralls Co.: J. Davis 4799 (Mi). Saint Louis: Eggert s-n- [4/8/75] (Go), san. [St. Louis, Aug. 1875] (ca—181533, Ca—l25203), S-Be (St. Louis, Peat 8, » 2875) (Cm, ee ie sen. [St. Louis, Avg. 1875] (Au, B, Gm, Du—202130, Gg-—183215, To—-79816, wn—5893 o--97l10, Po—15259, Up--60752, W-—-751953); Engelmann 337 (Pr), Sn. [St. Louis, Sept. 181] (W); M. Martens sen. (Br, Br)» CONT ty undetermined: J. Davis s.n. [Oak Boulder, 51T.1915) (Au); Le- jeune s.n. (Br). LOUISIANA: Orleans Par.: Cocks sen. [New OF rican, Apr] (Tl). MONTANA: Cascade Co.: F, W. Anderson 332 (in—11928) 5 eis i 692 (N). Gallatin Co.: Blankinship s.n. [Manhatt@% ° 98] (Fc); D. B. Swingle s.n. rks, July 21, 1933] (Mn—23399, Pl—71262, Sees sachet tt jefferson COs: 1963 Moldenke, Monograph of Verbena 277 R, Adams s.n, [Fish Creek, Sept. 8, 1871] yeas 3 B. T. But- ler 292 (Du—-123),86, N). Lewis & Clark Co : F. De Kelsey SNe [duly 26, 1892] (ob—50872), sen. [Helena, ‘ Gotober 17, 1892] (Du- 226538, Po—19762),) « Ravalli Co.: Hitchcock & Muhlick 21937 (N)- Sanders Co.: M. E. Jones 8963 (Mn—7890, Po-—-72923) « County unde- termined: F. ¥.. Anderson 108 [Great Springs] (Ca—25147) « IDAHO: Ada Co.: crise & Ward 8386 (N); J. A. Clark 253 (Ba, Ca--168207, Du—2186, S, W—5)43138) 5 Mulford s son. (Boise, 5, July '92] (C). Canyon Co.: hae 304 (Ca— —160621, | Cm, Du-—2)185, N, Pl-— 22566, Ur, W-—542255). Kootenai Co.: H. J. Rust 209 (W--870175). Latah Co.: J. H. Christ 6881 (N). Lemhi Co.: Je He H. Christ 5962 (N, N). Nez Perce Co.: J. He Christ 12958 (N)5 Heald 3h) (Pl--10h727). WYOMING: Sheridan Co.: cle Nelson . 2208 (N), 2258 (W—28),732) 5 Tweedy 266 (N). County undetermined: McCosh s.n- (Pr). UTAH: Box Elder Co.: C. B. Maguire 21402 (Ua--47397)3 Zundel 252 (Du— 75588). Cache Co.: Cronquist 1095 9 patible Gessel SMe [July 29, 1939] (N, Ua--b739); Ce Be Maguire 3731 (Ua—11388), 20098 (N, N, Ua--h7395), 21101 icra tre fee C. . Pith age (Du- alish, va--1236h). Davis Co«: Garrett 7868 (Gg--321166). S Lake Co.: M. K. Clemens s-n. (Murray, Sept. 25, 1909] a apt Me. E. Jones 3 487 (Po—70992); F. E- Leonard s.ne [Salt Lake City, July | 17, 188] (Io—131122, Ka--81)00, 00, Ob--5087h) 5 a & Black- wood 3638 (Io—28507, Io--83751) ; Rydbere sen. [July 1 188) (Di-337390) 4 S. Watson 822 (W). Utah Coe: Abrams 7213 "“pa--89892)5 J. A. Harris 0.27628 (H-—40515)5 Be Fe Garrioon 9217 (ca—8826h2, at 11865 (W—2098708); M. 5. Jones Jones 1487 (Br, Go, Po— 70993, Sg--16089, Ua—11383); Tidestrom 1718 Ts wells Woodcox 55 SS (Au—122283). County undetermined: F. E. Leonare [Jordan Valley] (Ob—30873); E. L. Little son. [July 3, art Tox —10352). NEVADA: Lincoln Co.: M. Es 2 Jones Son ‘Selle Sag, sn par ( 5, 1912] (Po+-69600). COLORADO: mot Coe: Ea Racked in Ca- 450176). pein Cost Le Kelso Wit (Hp) - er Cow: He W Campbell 121 (B1—1,2308); McFarlane s.n- 1881] y (eae 3 Ram- aley 740 (Bl—2320); W. We actiee 2554 (B1—-h2318), 5732 (B1— 42309); Tweedy 512h (N). Clear Creek “Gos: Bethel s.n. [Clear Creek, Aug. 9, 1916] (Gg--31326); Heustis s.n- [Clear eee . 19-16} (Bl--2317), sen. [Clear Creek ‘eek Junction, Aug. 1 (Bl—12321) ; Me ae ve [Georgetowmn, 7-2-1878] (Po—83731). Denver Co,: Bethel sen. [Denver, 7.1906] (Fc)j Eastwood 1, in part (B1—I2319, Gg--31325, St--229k6, W-582287), Sone [Denver, Aug. 1890] (Bl1—2336); McCosh & Greene SoBe [Denver, Auge 22) (Ec, Jr, M1, Pr, Va); He Ge “G. smith dre 8 s.Me n. (Denver, (W), sn, (Berkeley, Aug. 2, 1901] (Bl-—-42312)5 H. S- Nee = Be n. [Denver, July 29, 1933] *tee-17875) Douglas Co.: R. B. Living 278 PHYTOLOGIA Vol. 9, no. 5 ston 457 (H—77h53). El Paso Co.: I. M. Johnston 2633 (Ca—206896); R. Sutton 143 (H—77463). Fremont Co.: | T. Se . Se Brandegee 46h (Ca~ 169172). Gilpin Co.: Scovell Son. [Central ¢: City, 1869] (Mi). Jef- ferson Co.: L. S. Ehlers 688 (i); 3 Ewan 14552 (Bl—-52992, Gg-- 345128); M. E. Jones 523 (Br, Du—151783, N, Po--7099h, Ua-— 11382). Larimer Co.: C. F. Baker s.n, [Fort Collins, 8-25] (Po 63901); Collector undesignated 2015 (N, Vt); Je He Cowen 05 (W-- 254915), sen. [July 11, 1893] (N); Crandall 201 (Fe), § Sone n. [Fort Collins, Aug. 3, 1898] (Io—22911); Herb. State Agr. Coll. 11. Colo. 2015 (Fe, W—l89)91); W. F. Marshall 20 2016 (Fe, Fe, c, Mn—6086, Pl— 22569, W-l89192); Osterhout sen. [July 22, 1901] "(N, Po--6 3893) 5 Preston s.n. [E. of Fort Collins, 7-31-1921] (Fc). Las Animas Co.i C. M. Rogers 981 (W—-2053370). Logan Co.: Osterhout 7740 (Po-- 21665). Weld Cos: Ewan 12267 (En, Pl—122581); M. T. James s-n. (Roggen, Oct. 3, 1931] (Bl--1231h); E. L. Johnston 507a (N), 507% (W—768891); Osterhout s.n. [Aug. 5, 1905] (Du—77582); Ramaley 12393 (Bl—l.2316), 12890 (B1—12890). Yuma Co.: Rggleston 15551 (Po—63862) ; Ramaley 127)1 y 127h1 (B1l—-2366). County undetermined: Crandall 175 (Fc); W. F. Marshall s.n. [Mason's River Front Farm, July 22, 1899] (N). NEBRASKA: Buffalo Co.: W. E. Be son. [Avg- 1922] (Ok—-10353) , Cass Co.: J. L. Morrison 1200 (Go); Te A+ Williams s.n. [Weeping Water] ~(W—75008) - Dawson Co.: Wahl | Sete [Willow faiand) Jul. 1922] (Io—~102)2)). arcane: Co : Enander S- Se ne [31/8/1921] (S); Kleeberger s.n. [Omaha] (Gg—31333) « )» Hooker Coe: Rydberg 1515 (N, W--210370). Kearney Co.: H. Hapeman S-Me (Minden, ise. 23, 23, 1939] (Hp, Ky). Knox Co.: F. E. Clements 279 (Ka, W). Lancaster Co.: J. L. Sheldon s.n. {Lincoln, 7-20-9 6) (We, We); me J. Webber s.n,. ~ [July h, *85] (Cc). ape Coe: a Le Sheldon s.n. [Peru, July 7, 1900] (We). Otoe Co.: Sen. Sen. [Talmage, 1890] (Je—58)0), s.n. [Talmage, or aT Ge 3156, Je—3157). Sheridan Co.: R. E. Buchanan s.n. [Aug 2--20, 1905] (Io—-51233), sen. [Spade Ranch, 1905] (1o--l9012); J. Be Hatcher s.n, [on the Running Water, VIII.1--10.1886] ere ’ Sen. [July 1886] (Pr). Thomas Co.: Blumer sn, [Halsey Oct. 1902] (To—82631); Krautter s.n, [duly 23, 1907] top lst, 5745). Washington ¢ 0: Lallman s Sen. [August 19, 1936] (Bt— 45hh7). county SP teeetccn Pre . E. Abbott s.n. [1907] (Ge)s Re oe 158 [Rat Lake] (788961), 302 [crescent Lake] (N, W-- 789095) . OK * Cherokee Co.: C. S, Wallis 669 (St). Craig pte Brensing 21 (Ca—882658, st—15268); & E. B. Miller 63 (St--25006) - Clark 3354 (B). Delaware Co.: C. S. Wallis 2534 (St). Ellis Co.: G. W. Stevens 2901 (Du--68056, Ok, Ur), A-20L (St--92h6). Grant co.: G. W. Stevens 1791 (Du—77110, Ok, “Ok 1963 Moldenke, Monograph of Verbena 279 21305, Po—63860, St--92h7, St—92h8). Harper Co.+ Stratton 100 (St--26127, St—26128). Muskogee Co.: E. L. Little 2435 (Ok— 10311); C. S. Wallis 7728 (Au--170667, St); Waterfall 1028) (st), 10298 (St). Oklahoma Co.: M. Hopkins 121 (Ok); Waterfall 2462 (N), 3074 (Ok). Okmulgee Co.: Stemen 547 (Ok). Ottawa Co.: G. W. Stevens 2308 (W—589895). Payne Co.: Van Vleet 399 (Ok—-20508) . Sequoyah Co.: C. S. Wallis 5557 (St), 1736 (St). County undeter- mined: G. W. Stevens 2145 (Ok). TEXAS: Hartley Co.: Cory 16431 (ud); Parks & Cory 16428 (tr), 16429 (Tr), 16430 (Tr). Hemphill Benton Co.: Gotfredson 90 (Po--16360). Clark Co.: C. English Jr+ U77 (W--1620266); Gorman 5069 (Du--189721, Or—22h21). Douglas Cow: Cotton 1396 (Pl--22583). Kittitas Co.: C. V. Piper sone [E- nsburg, July 9, 1897] (P1--22630). Klickitat Co.: Suksdorf 5862 (PL~138i18), 7972 (P1—-138419), 12072 (P1—138i20), Sane [Nov. 189] (P1—-138)2)), sen. [July 8, 1095] (P1—138h25), son. [Vil- 190] (N). Lincoln Co.: Suksdorf 150 (P1—-13822) . Okanogan Cos: Dillon 842 (Ca—739564, P1--151100, Se--94450); Fiker 332 (Pi— 7619). Skamania Co.: Suksdorf 6658 (P1—138L17). Stevens Co.: Beattie & Chapman 2188 (Ca——002061, Pl—2257h, Um—22h, Um-—-233, W—187Lj10); Kreager 46k (Ca—882660), h69 (N, Pl—22577, Se— 68697, Un—225, W— , Whitman Co.t Co Ve Piper son. [Almota, ug. 26, 1894] (P1—2262h). Yakima Co.: T. Se gee SMe annam Creek, 1880] cas ess 3 Cotton 767, in part (P1—22582) 5 L. F. Henderson s.n. [June 19, '92] (Se--14943); Ge Ve Piper s.n. [Aug. 6, 159k] (Pl—22625); Watt sen. [North Yakima, Aug- 2 a (Pl--22626), County undetermined: Lake & Hull 605 [Rock Lake] (P1l- 22628). OREGON: Clackamas Co.: Suksdorf sn. [Aug. 2, 1896] (Pi—- 138421). Columbia Co.: L. F. Henderson SMe [Columbia R., August 12, 1885] (Du--90899). Linn Co.? F. B. Lioyd sen. (11 Juy 1894] (N). Marion Co.: J. C. Nelson 005 (Al). Wultnamah Co.: Gilkie s- Re (Portland, July 31, 1935] (Or—3349h, We); Be Be Sheldon 11167 Du—178070). Wasco Co.: L. F. Henderson 798 Island: Je Te. Howell s.n. [Sauvin's Island, 280 PHYTOLOGIA Vol. 9, no. 5 CALIFORNIA: Contra Costa Co.: Condit 253 (Ca—l5)110); V. Jones Sen. [Orwood, Aug. 29, 1920] (Gg—31320); Nobs & Smith 942 (Ca— 19206) . Mendocino Co.: Chesmt 58), (W—-l,30636). Sacramento Co.: Jepson Sone [Lower Sacramento, Sept. 1891] (C); Wilkes SMe {marshes on the Lower Sacramento] (W). San Joaquin Co.: Crampton 31L9 (Gg); Jepson 5693 (Ca), 10215 (ca); E. B. Stanford 721 (Du— 170466, Po--147778). Shasta Co.: M. S. Baker s.n. [Fall River Lake, July 15, 1899] (Ca—-li50175). Solano Co.: Jepson sen. [16 Sept. 1891] (Up--17105); H. L. Mason 12666 (Ca--192063, Du—15507, Ge 417883). Yuba Co.: Eastwood s.n. [Marysville, Sept. 1919] (Gg-- 31322). Grand Island: Jepson 21229 (Ca, Ca), sen. [17 Sept. 1891) (Ca~-251)9). JAMAICA: Pessin s.n. (Tl). GREAT BRITAIN: England: Re Campbell s.n. [Bristpool] (Mm—20l,78). SWEDEN: Blom s.n. {2/9 /1939] (Go); Schontz Sen, (Go). GERMANY: Kausch s.n. [Rostock, 1869] (La). CULTIVATED: Austria: Herb. Mus. Bot. Stockholm s.n. [Hort. Schtnbr.] (S). Belgium: Herb. Jard. Bot. Brux. se. (BY, Br); M. Martens s.n, [h. b. L.] (Br), sn. [seed from St. Louis, Mo.] (Br). England: J. D. Hooker s.n. (Br), eus s.n. [Hort. Son. [h. f. 18] (Du--166)36); Herb. Hort. Paris. s.n. [Aug- 1820] (W—617239), son. [Jul. 1822] (W—617239). Germany: Herb. Hort. Gast. sen. [1806--12] (W—617453); Herb. Martius son. (BY)j - Mus, Bot. Berol. s.n. [Sept. 182] (Br); Herb. Prager 18639 (Gg—3157); Schulz & Sohuts saps 11.8.1897] (B)+ Maryland: McCann s.n. [9-15-36; seed from Copenhagen] (Md), S-Be (9-15-36; seed fran Gradiva Botanica, Bucharest] (Md), SMe (9~ 15-36; seed from Helsingfors] (Md), Netherlands: Burmeister 5-De (S). New York: G. V. Nash san, [N. Y. Bot. Gard. Cult. Pl. 20853] (N), Sen. (N.Y. Bot. Gard. Cult. Pl. 20972] (N); Re C. Schneider Se—119)1) ; Je Fowler son. (Ms); Herb. Coll. Phar Herb. W. H. Harvey son. (Du—166L39)5 Herb + t YPe, Z——photo of isot : Herb. Meisner (M); Herb. Mus. Bot, vPe)s ee Selle — Stockholm s.n, (S, S); Herb, Saldanha 2765 1963 Moldenke, Monograph of Verbena 281 (Herb. Mus. Nac. Rio Jan. ref. VI] (Ja); Herb. Swartz sen. (S, 8); Herb. Torrey s.n. [N. W. coast] (T); F. W. Johnson 770c [Fitches Bridge] (N); L. A. Koehler s.n. [U. S.] (Pa); M. Martens sno (Br); Née 61 (Q); New ‘ew York Bot. Gard. Econom. Mus. 5749 (N); Par- ———— SS Pollart de | de Caniéri ate (Br); Schweinitz s.n. (T); Short s.n. a: $8); Je: J. Torrey s.n. (Br); Whipple s.n. [13-1] Semgale Bape ILLUSTRATIONS: E. J. Alexander, Addisonia 13: pl. 422. 1928 (N)3 Am, Mus, Nat. Hist. color slide 27621 pats 2388),0] 5 “toto kiss distribution map (Ws); Rydb., Fl. Prairies & Plai - 479. 1932 (N); Natl. Geogr. Mag. pl. 3, 7/22 (N)5 anpubl sp (N); unpubl. pencil drawing (N); Wild Fls. Canada pl. 9 (N)- Casing re f£, ALBIFLORA Moldenke, Am. Midl. Nat. 2h: 752. 940 Synonymy: Verbena; americana; altissima; urticae foliis angus- tioribus; fiictns albis Herm. apud Boerh., Ind. entities , Plant. Hort. Acad, Lugd., ed. 2, 1: 187. 1720. Verbena alba Moldenke in Chittenden, Roy. Hort. Soc. Dict. egret Pe “2211. Bibliography: Boerh., Ind. Alt. Plant. pg Acad. Lugd., 2, 1: 187. 1720; Cheney, Rhodora : 26. 1902; Robinson & = aa ant Known Geogr. Distrib. cag {ed. 1], 2& Phytologia 23 478. 1bL8; pndetke. Alph. ist 2 Gat, 2: 507 & 536 (1948) rage Geogr ° sy J The type of the form was collected by Ellsworth Jerome rome Hill (no. 9.1888) in wet ground at Attica, oe o County, New York, he er of the Uni- i us far only s fron Massachuset k. New Jersey, Iowa, and singh aa along ro roadsides, and at the edges his rowing fend I see no evidence of hybridit, fos is = ection seems typical of the albino form of Te i eet 2 t Dees rollection cited below was distributed original as V. urticifolia L. In all, 9 herbarium specimens, including the type, and 3 moun- 282 PHYTOLOGIA Vol. 9, no. 5 ted photographs and clippings have been examined b Citatio: HUSETTS: Berkshire Coe: upidenke | & "Yoldenke 1171 (N); W. A. Kober 1590 (Bl—18180). Worcester Co.: G. EB. Stone s.n. [Proctor's jor's Pond, about 1890] (Ms). NEW YORK: Wyoming Co.: E. J» Hill 9.1888 (N—-photo of type, Ur—type, Z—photo of type). ] NEW JERSEY: Ocean Co,: Lawrence & Dress 570 (Bm). IOWA: Des Moines Co.: Davidson & Werlein 979 (W—2261038) . WISCONSIN: Shinners son. [Aug. 31, 1938] (Ws). Outagamie Co.: J. H. Schuette s.n. (Kaukauna, Aug. 7, 1889] (W—752023). MOUNTED CLIPPINGS: Am. Midl, Nat. 2h: 752. 190 (Ww). VERBENA HASTATA f. ROSEA Cheney, Rhodora ): 2h5-—26. 1902. Synonymy: Verbena hastata var. pani ta f, rosea Farwell, Papers Mich. Acad. Sci. 2: 37+ 1923. Verbena hastata f. rosea Farwell ex Moldenke, Prelim. Alph. List Invalid Names 46, in in syn. 19)0. bliography: Cheney, Rhodora ): sonata ey Robinson & Fern. in A. Gray, New Man, Bot., ed. 7, 689. 1908; ell, Pa- Invalid i 46. 190; Moldenke, Known Geogr. Distrib. Verbenats, fed. 1], 2, 3, 6, & 101. 1942; Moldenke, Alph. List Invalid re 7. 19h2; theta Am. Journ. Bot. 32: 609. 1953 Moldenk hes List Cit. 1: 11, 208, & 2h.(i9h6), 2: 506, "568, & 581 (1948), and 3: 740, 803, & 948. 199; Moldenke own Geogr. Distrib. Verbenac., [ . 2 ’ S, ee gk & 196 19195 He a : € ba ‘yol- denke, Anal. Inst. Biol. Mex. ee 3. 19493 Man. Bot., ed. 8, 1210. 1950; Woldenke, ARE 5! "er. “T9825 enke in Gleason, New Britton & Br. *Tliustr. Fl., print. 1, 1952) and 223s "129. 1958; Moldenke, Am. wal, Nat. 59: 37. 1958; Moldenke, Résumé 6, 7, 16, 19, 366, & 19595 Uoldenke, Résumé Suppl. 3: 1, 2,’h,’e 5° (1962) and $3 Te s form differs from the typical form of the s sere in hav- ing the corollas pink Ae e-pink, mud on the margin of a lily pond in closest proximity to typical reville, Barnstable County, Massachusetts. a a most careful examination and a ee no difference was detected excepting in . » Manner of growth, form and texture ° alike. I counted sixteen plants of the rose~ a small space 2. form has been collected in low scat along with the tyP- form — in wet ditches, at altitudes up to 00 meters, eg Connie September, in fruit in July and August. specimens ha ve been examined by me. 1963 Moldenke, Monograph of Verbena 283 tions: CONNECTICUT: New London Co.: Jansson s.n. [Aug. 9, an posanpnnere NEW YORK: Madison Co.: H. D. House 23127 (N, Up, W--1653793). Oneida Co.: H. D. House 26197 (Al); C. H. Peck san. [Trenton Falls, Aug. 18, 102] (Al). Queens Co.: E. P. Bick- hell 7353b (Al, N), 7353¢ (N). Tompkins Co.: S. J. Smith 1L6h (Al, It). NEW JERSEY: Monmouth Co.: 0. E. Pearce san. [Swartzwood, Aug. 1, '8h] (W—117578). PENNSYLVANIA: Huntingdon Co.: B. H. Graham Ae [7/12/32] (Cm). Sullivan Co.: F. R. Fosberg 15770 (Up). WEST VIRGINIA: Preston Co.: J. L. Sheldon 1881 (We). ILLIN- OIS: Champaign Co.: Ahles 2918 (Ur). DuPage Co.: W. S. Moffatt 573 (Ur), 574 (Ur). MICHIGAN: Monroe Co.: Me McDonald a 5650 (Mi). SCONSIN: Buffalo Co.: Fassett & Hotchkiss wae (N, Ws). Polk Co.: Re W. Pohl 515 (Ws). KANSAS: Franklin C oe: Re Le McGregor 12700 (W--2230L0h) . sho age deca » Résumé Suppl. 3: 6 & 8, ye Collectors describe the sigh as "very tall", the unbranched - from a crown, and the 3 in pjeds ditches and wet places, Swampy fields, on pela oA along highways, blooming from June to September, "fruiting from July "to October, from abies altitude. It is S lied "blue vervain". Elm as "very rank along irrigation ditches", while Cory says of it Scattered in canyon bottom near stream bank, Carex, Juncus, Bidens, and Hordeum in moist coarse loam soil." J. A. Harris C.28865 was used in 4 study of osmotic pressure. Kreager 75 is a mixture with V. stricta Vent. It is very probable that some of the northwestern specimens cited by me on previous pages under typical Ve Ve hastata belong by me and and returned to owners in case convinced of the validity of this taxon. They need r bari’ aa ee the type, have 28h, PHYTOLOGIA Vol. 9, no. 5 (W——-2310063). County undetermined: J. Macoun s.n. [Sicamous, Aug. Ujth 1889] (W—219611). NORTH DAKOTA: Richland Co.: 0. A. Stevens sen. (Aug. , 1936] (Hi--1920)6). MONTANA: Gallatin Co.: : Blankin- ship s.n, [Logan, July 23, 1906] (W—5l1111). IDAHO: La Latah Co. Le F. Hen Henderson s.n. [Juliaetta, Aug. 31, 9h] (W—22669) . Owyhee Coe: J. M. Johnson 254 (Hi--192662, N). UTAH: Utah Co.: Gal 2192 (Ca—911080); A. 0. Garrett 3149 (ca~hS369), 7868 (Ca-— 882659); J. A. Harris C.26865 (Ca-—882655). TEXAS: Hemphill Co.: Cory 50298 (Ca--754878, Du—345342, N, Sm, W—18859))). NEW MEXI- CO: Socorro Co.: Wooton s.n. [Crain's Ranch, July 1, 1900] (W#— 56228, W--736876). WASHINGTON: Okanogan Co.: Elmer 537 (N, Pl 22629, Po—-63903, Vt, W—352309). Skamania Co.: 3 Suksdorf 10572 (Ca—755098, N, N, P P1--139857, S). Stevens Co.: Boner & Weldert 202 (Ca-—7079156, Du—301168, Gg—-310858, N, P1~93879, Se——69128, Ua--3810) ; Rronger 475, in part (W--)1291). bey! Walla Co.? Ry M. Horner 17 (W—26 ). Yakima Co.: Cotton 787, in part (W— 525258), 1396 spent oot Kruckeberg 255 Cree wi ORE- GON: Multnomah Co.: E. P. Sheldon $.11167 (W—34857). Umatilla Co.: Leiberg 2630 (N, | W=2 75970). CALIFORNIA: Modoc Co.: He Le Mason 13502 (Ca—192061--type). San Joaquin Co.: H. L. Mason 5h25 (Ca--635119). Shasta Co.: Nobs & Smith 100 (ca—192062, Du— 415615, N). County undetermined: J. S. Newberry s.n. [Upper — of Reb River, Aug. 8] (W). VERBENA HATSCHRACHI Moldenke, Lloydia 13: 226. 1950. Synonymy: Verbena hastchbachii Moldenke ex angely, Fl. Paran 12: 17, sphalm, 1958. Verbena hatschbachii Moldenke, Résumé 128, in syn. 1959. Sete aor Bibliography: Moldenke, Lloydia 13: 226. aes Moldenke, tologia 3: 377. 1950; setiree Trib, Farmac. [Vellozoa] 19 * flo): 167. Bae Ee Je Salisb. , Ind. Kew. Suppl. 11; 263. 19533 AngelY, 12°(1957) and 12: 17. 1958; Woldenke, R&sU- 2é 110, 0, 18, ‘etre? 1959; woldenke, Résuné Suppl. 1 23. 19593 Angely, - Paran. 16: 2B d17: 6. 1 gee t herb; stems very slender, tatragonal, — lent, rooting at the nodes; ‘ivi ascending, slender, 78" ther’ acu tely tetragonal like the stems, pube prod Fe with more or upwardly curved hairs; nodes annulate; principal internodes 1--k.5 cm. long; leaves decussate-opposite; petioles slender, 3— 5 mm - long, sparsely strigillose, margined; leaf—blades membran- es "bright~green on both erent ovate in outline, 6—16 m- long 6— Cy Ate slightly prominulous veinlet reticulation indiscernible on Spoth surfaces; in- 1963 Moldenke, Monograph of Verbena 285 florescence terminal, spicate, pds on As & the floriferous por- tion subcapitate when young, elongating to 2 cm. in fruit; pedun- cles very slender or subfiliform, egret 5 cm. long, acutely tet- ragonal, whitish-puberulent with somewhat antrorsely curved occas- ionally gland-tipped hairs; bractlets lanceolate, about t 5 mm and 1 mm, wide, attenuate-acute at the apex, often pu lish, white-ciliate along the margins, otherwise sent 3 calyx 4-5 mm. long, purplish, 5-ribbed, white-ciliate on the ribs, otherwise glabrous, its rim une 5-apiculate; coral. cae ocrateriform in 1621) — in whose honor it is named -- along the road from Rio Taquari to Rio D n the municipalit quara, Par Brazil, on tenekinns “23, “1919, and is deposited in the Britton He rbariun at the New York Botanical Garden. The species has. been collected at altitudes of 1000 to 1300 meters, blooming in July and November, In all, a herbarium specimens, including the type, be Citations: BRAZIL: es Hatschbach 1621 (N—type), 4895 (Sm). Santa Catarina: Smith & Klein 7376 (N, W--2251273, Z)- VERBENA HAYEKII Moldenke, Phytologia 2: 146. 196. Syno; : Verbena procumbens Hayek in Engl., Bot. Jahrb. 2: 163. 1908 [not Ve eaten Forsk., 1775]. Bibliography: Forsk., Fl. Aegypt.-arab. 10. 1775; Hook. f. & Jacks., Ind. Kew. 2: 1179. 1895; Hayek in Engl., Bot. Jahrb. 12 Geogr, Distrib. Verbenac., [ed. 1], 35 & 102. 19h2 enke, 23 + 1946; Moldenke, Alph. List pl. 1: cas 1947; Moldenke, comiagee : 116. ae ; - Ne & Ao Le Moldenk 2: 43, , 191,83 Moldenke, Alph. List| Pee 4 3 hh, Cit. 2: E51 & 602 (1948), 3: be. : "987 (dsl), aa "hk: 105k, & 1229. 1919; Moldenke, Known Geogr. Distrib. Verbenac., ed. ae bs = ak 19h95 Be Je Salisb., Ind. Kew. oe 1l: wet ae tua ja 5: 2h. 195): vee Inform. Mold. te be “Ure. 1959. s trate, rooting, glabrous; branchlets erect, shar’, twigey, : sparsely setose; leaves cuneate-oblong or obova vate, narrowed a the base, sessile, coarsely incised-dentate, strigose-hirtous on coming ‘Lindri both ce es the base of the branches; bractlets ovat te-lanceolate, | ge ‘ew the apex, setose, subequaling the calyx; sane angulat i eee : » the rim from 600 meters, Junin, Peru, and was deposited in the herbarium of the 286 PHYTOLOGIA Vol. 9, no. 5 Botanisches Museum at ea gi where it was photographed by Mac- ride, but is now dest This high alpine species ha s been found on turf hillsides, in meadows, along roadsides associated with Ranunculus eee grasses, on grassy-shrubby steppes, in cultivated soil, and in thin soil over limestone, at altitudes = ga to 5660 Larne peru es in January. Stork & Horton call it "a lowly pasture herb, possibly car gy pate grass" and describe the flowers as "white and hairy in the center, the ss obes lilac or pink". Ochoa 268 is said to have se white Pow The species has peek reidiented in anthesis from November to Jarmary and in March and April, in fruit in January, and has been misidentified and distributed in herbaria under the names V.- cun- eifolia Rufz & Pav. and V. hispida Rufz & Pav. A common name re- ported for a is "canserccora®. Forskal's V. procumbens is a syn- onym of v.58 Le ites te comments follows about our plant: nsteht ent- schieden der vorigen ape Ive weberbaueri] nahe, ist aber durch die verltmgerten, wurzelnden Aste, die grtsseren Blttten der anders gestaltete Inflorescenz ured andere Bl&tter verschieden." all, 2 herbarium specimens and 3 a photographs, in- cluding phototypes, have been hesornen Citations: PERU: Cuzco: F. L. Herrera 7409 (W--1283221) « Hu ancavelica: Stork & Horton 10820 (a bB6765); Tovar 115 (W-- 2100h09, 2), 855 (Ss). Junin: W. H. Hodge 6237 (Ms, N)j A. Se Kalenborn 36 (Du—103350, N, W--l0W28h), 176 (Du--103355)5 Kal enborn & Kalenborn 36 (N); Kunkel 375 (Gg), > 305 (ag), 81h (14), 817 (Mi); Ochoa 268 (1 (W--1950563) ; Meberbauer 2573 — photos nie} (ie photo of type —-photo of type). - Aguilar F. fe Aaa ee °1918} 1a; "Fe W. Pex nell hoe Ate Soukup 259 Soukup 259 Reaidione: W—1775352) 3 De D. Stafford 27 (K). Department undetermined: Cook & Gilbert 99 [Sicuant Vilcarnota Valley] (W--603386); Cc. Watkins SoMe 'S Sen. [Tuapata, 1916] (W—1059586) . Moldenke ex Herter, Revist. Sudam. Bot. hs 186, ios. me 19375 Moldenke, Revist. Sudam. Bot. 5: 2——l)3 « nymy: Verbena dumetorum Osten ex Moldenke, Alph. List In . ie Bibliography: Herter, Revist. Sudam. Bot. h: 186 . aft tee o . nace, [ed. 1], i? Ri Sul Suppl. 1: 23. 1947 HLL & Salist : 10: 2h2- 19h75 He Ne &A- Le “tate 3: 6h, 1968; Moldenke, Alph, List cit. 2: 367, 366 368,’ 632 (1948) and 3: 700 & 875- 1949 1963 Moldenke, Monograph of Verbena 287 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 100 & 198. 1949; Moldenke, Phytologia 3: 377. 1950; Moldenke, "Résumé 110, 119, 363, & 473. 1959. Low herb, 15-60 cm. tall; stems and branches prostrate, creep- ing, tetragonal, very slender, va from lightly puberulent to glabrate or glabrous; internodes abbreviated; leaves small; peti- oles elongate, 4-10 mm. long, glabrate, slightly margined above; leaf-blades thin-m embranous, oblong or elliptic to ovate, 8—1) mn. long, --10 mm. wide, o obtuse or very obtuse at the apex _ acuminate or cuneate at the base and protracted into the petiole, ly te al margins teeth per side, occasionally oblong with more or less paralle entire margins and 3-toothed only at the apex, entire from the Widest part to the base, glabrous or glabrate on both surfaces; peduncles —— slender, ).7--7 cm. long, erect, tetragonal, ut 2— mi. long, 5-costate, membranous between the ribs, sparsely pilose with ascending white hairs, its rim slightly were enh = : i from long h idynamous, the upper pair each with a long clava sppendage} * pistil included, about 9 mm. long, ciocele bifid at “th e oes of this distinctive ee was collected by my good friend ah colleague, Wilhelm Gustav r (no. 9793 Herb. Her- ter mall in sandy banks, at an baer a 50 meters, Artigas Truguay, in November, 1927, and is deposited in the Britton Her- barium iv the New York Botanical Garden. The aeacies differs fron all other known Uruguayan species of the section G 288 PHYTOLOGIA Vol. 9, no. 5 violaceis; appendicibus antherarum clavatis; corolla longe exser~ ta." In all, 13 herbarium ae cimens, Sti the types of both names involved, have been examin y m Citations: BRAZIL: ran Grande do win “Rambo 26125 (Rb). URUGU- AY: Berro a (N); Herter 979 [Herb. dice yi ements B-—isotype, -323326—-isotype, It-- —isotype, Te—9208 1 ttectype), 979a (Herb. Spe eee) ray, rae 3128 (Ug), 3326 (Ug); Rosengurtt B.3770 (N). VERBENA HIRTA Spreng. in L., Syst. Veg., ed. 16, 2: 7h9. 1825 [not V. hirta Grah., 1959]. Synonymy : Verbena pohliana Mart. ex Moldenke, Alph. List Inval- id Names Suppl. 1: 1. 1: 26, in syn in syn. 1947. Bibliography: Spreng. in L., Syst. Vege, ed. 16, 2: 7h9- 18255 Cham., Linnaea 7: 255—256. 1832; Walpe, Repert. Bot. Syst. h: 22- 23. 1645; Schau. in A. DC., Prodr. ll: 50. 18h7; seas in Marte, Fl, Bias, 93 185. “1851; Hook. age & ra acks., Ind. Kew. 2s 1179+ 18955 Arch. ac. Rio . 13: 36. 1903; Hayek, Denkschr. Kaiser. Akad. Wissen. S neeh oo 79 (1): 291 6 1908; “yee ie i - « ® = ° Grande ke, Alph. List Invalid Names oe tard - = a spiders tanea 13: 117. 198; Moldenke, Alph t Cite 2: 362— 370, 402, Lbh, bus, hho, 486, "932, a "s51—353, . - (1948), 3 670, 675, 687, 750, 772, 81h, 815, 8h6, 855, 900, & $63 (1949), and bi 1015, 1077, 1081, 1113, 12h 1180, eo 03, 120h, 128-1251, 1287, 1300, & 1301. 199; Moldenk , Known Geogr. Dis~ trib. Verbenac. {ed. 21,9 94, 106, 163, & 198. "199; Moldenke, Phytologia 3: 7. 1919; Moldenke in Chittenden, Roy. Hor ots Soc. Dict. Gard. 4: 2208 & 2211. 1951; Ste aT Trib. Farmac. 1 (10): 166. 1951; Moldenke, Phyto oo gia h: « 19533 ideale In- form. Mold. Set 8 Spec. [h] (1951) aan oe Spex. 3. 195k; ° Sellowia 7: 260. 1956; Angely, Fl. Paran. 7: 12. 19573 \oldenke, Résumé 110, 127, 223, 366, 372, & “ine 1959; ok Levant. Herb. Inst. Agron. \inas 150. 1960; Angely, Fl. Paran. 16: 78 (1960) Low perennial herb, sometimes shrubby, usually to 20 cm. e. sometimes to 1 m, tall, densely hirsute, villous or villous~ ous-Lana t% globose, soon pean cylindric, always dense~ » cymose, axillary and terminal, capeeite-oonnested wd 1963 Moldenke, Monograph of Verbena 289 long-pedunculate and racemose, ternate; bractlets lanceolate, cute at the DORs _ciliate-margined, equaling or slightly shorte er than th ranous, about mm. long, plicate- costate, subglabrate me the ’ angles , hirtous on the herbace- ous angles which terminate in subulate subequal very corolla all g from blue, blue-violet, or violet to lilac, rose, clear-rose, reddish, red, clear-red, or purple, ali htly surpass— the calyx, pubescent or Lanugino us above on the outer surface, villo us at the insertion of the stamens within, the mouth bearded with short and uniform submoniliform hairs, the limb about h e, the lobes retuse at the apex; filaments distinct, the upper ones longer and cernuous; anthers muticous; cocci linear, fuscous and Srstriate on the back, areolate, the commissural face broadly white-gran hand, the Dusén 6798 distributed as V. hirta is actually vare gracilis Dusén, The so-called V. hirta of Graham is actually Lan- tana hirta Grah. " ‘The type of V. pohliana was collected by Johann Emanuel Pohl — in whose honor it was named — in 1839 somewhere in Brazil, & Bailey 82h, according to L. H. ey, daged. The Rombouts s.n., cited below, exhibits one dark-green very hairy stem and one light-green lightly hairy stem while Hauff 16-35 shows one extremely hairy sterile stem and one less floriferous stem. The Otto s-M- cited from eae in Germany was ee from plants gr grown from sgt sen Botanisches. il in Berlin, where it was pnotographed. by Mac- bride, but is now Sean "It may have been the ual type of the species. Smith & Reitz 8916 is a mixture with ¥- “Tobata var. Sessilis Mo ldenke. Schauer (187, 1851) oe Sellen sone {Mogy], Pohl sen. [Reg- isto Velho et Queluz], Ne gen. [castro o], and Lund s.n- [Cas- tro] from S#o Palo. Hayek ry cites Wacket 8-Me ne [inter et Alto da Serra], sen. [prope S- J, and s.n. [prope Campo Grande], and Campos Novaes S-ne [Campinas], also all from 290 PHY:TOLOGIA Vol. 9, no. 5 S&o Paulo, Brazil. In all, 210 herbarium specimens and 3 mounted photographs have been examined by me. Citations: BRAZIL: Minas Gerais: Aragto 23 [A. Lutz 2287] (Lz); Brade 1566, [Herb. Rio Jan. 32898] (B); Burret & Brade 1602 (Herb. Rio Jan. 35233] (N); P. Clausen 2)6 [18)2—L3] (Br), See [aug.—April 180] (Br); Collector undesignated s.n. (Ja—32328, Ja); Dusén 2036 (S); Henschen I1.209 (W--2096)2); F. C. Hoehne 5 n. [Pocos de Caldas, Jan. 9, 1919] (Sp——2685); Mendes Magalhaes 531 (Herb. Jard, Bot. Belo Horiz. 34357] (N); Mosén 966 (N, S), 967 (S); Occhioni s.n. [Herb. Rio Jan. 1658] (N); Regnell 1335 [Caldas, Jan. 1066] (Ja--148)9), II.209 [Capivary, 1862—63] (S), 11.209 [Caldas, 7/1/1868] (S), II.209 [Caldas, 187] (N, S); Wid gren 1212 [185] (Br), 1213 [1815] (Br), sn. [18h5; Herb. Rio Jan. 148]3] (lu, Lu, N, S, S, W-—-1199650, W--1323095), sen. [Bra- silia] (S). Paran&: Braga & Lange 221 (Bm, W--23693l7), 329 (W— 2369351), sen. (30/8/59; Herb. Inst. Hist. Nat. 5267] (sim); Bra- ga, Moreira, & Lange 329 (Gg); Collector undesignated s-M- [Ponta Grossa, 1874] (Ja--l6601); Dusén 8 (N, S), 2577 (Ja-—l6566, N), 2733 (Ja—6568), 3865 (Ja--6800), 7533 (Mi, N, S), 13239 (4, N), 13478, in part (S), 151K9 (S), sen. [Pirahf, 28.12.1903] (5), son. [Curityba, 25 Feb. 190i] (S), sn. [Villa Velha, 13.3.190h (S), sen. (Curityba, 7.X.1908] (S), sen. [Jaguariahyva, 2l1/10/ 1910] (S), sen. (Ja—31105, N); H. M. Filho 87 [Herb. Mus. Parane 5314; Herb. Curso Farmac. 720] (S); Hatschbach 599 (N), 100) (N), 1453 (N), 2913 (N), 3369 (Sm), 3760 (Sm), 6903 (Ii), 8201 (Ca)s Hatschbach & O'Curial 599 (N), 1599 (S); Hertel 2 [Herb. Mus. Paran. 1932] (N); F. C. Hoehne s.n. (Curitiba, Oct. 18, 1928] (N, N, Sp--23052), son. [Ponta Grossa, Nov. 2, 1928] (Sp—-23330)5 M: sen. [So Luiz, Aug. 27, 1939] (N, Sp--41532)5; Lange 1048 (Herb. Inst. Hist. Nat. 5260] (Mm); Lourdes Camargo 42 (Herb. Mus. Paran. 476] (N); Mattos 720 (N, 8), Sen. [R-20, no. 11; 9-11-51] (Sm); E. Meyer s.n. (20/12/58; Herb. Inst. Hist. Nat, 5259] (Mm); E. A. Moreira 27 (W--2369333), 5ly (W--2369337), 113 (W-2369342), sen. [2/1960] (Ih—-5912); Moure sen. [R-20, 7° 10; 2h-9-52] (Sm); Princesa Imperial s.n. [Campos Gerais, Dec- 84; Herb. Saldanha 8788] (Ja—j6542); Reiss 16a (I, N, S)5 Reitz 5722 (Cb); J. E. Rombouts s.n. [Ponta Grossa, July 17, 19385 Herb. Inst. Agron. Est. S. Paulo 2603] (Sp--l)1084, Sp)s Ne Santos s.n, [Araucaria, Feb. 191] (Ja—l6556); Stellfeld 1180 _ (Herb. Mus. Paran. 2232] (N), 2232 (S); Tessmann s.n. [Herb- eae eiro: 0 Pusén 2035 (Ja—L6563); Glazi ; Segadas-Vianna Ga), SIN3 (Ga); Sogadas-tianna, Bas, Orson i Waokrine 502 1963 Moldenke, Monograph of Verbena 291 = Ule 640 (March 1894] (Ja--6525), 640 [Dec. 27, 1895] (Ja— (Herb. Barb. Rodr. 739] (Le, W—214173h), 4516 (Herb. Barb. Rodr. 4742] (Le, N, 8, W—-21)1737), 4750 (W--2268919) , C.310 [Herb. Rio Jan. 51359] (N); Reitz & Klein L750 (Ok), 4865 (W--2252076), 52hbs (W--2252079) , 5270 (W--2252080), 6458 (Rd); } Smith & Klein 6428 (W--229358), LiWkS (Ok, W—-2251783), 12118 (Ok, W--2251953);_ Smith & Reitz 8916, in part (W—2251h66), 9162 (N, ri hare ce i 9226 (Ok, W—2251513), 9839 (Ok, W—-2251615); Smith, Sufridini 9315 (Ok, W—2251531), ait (Ok, W--22 Sassy =A Feiss Bailey & ] & Bailey 82h (Ba, Ba, Ba), 921 (Ba); Brade 5739 (N, S, Sp- 6713), 570 (S), 12 12393 (Ja--46495); Campos Novaes 726 (#389893); (N); Ha * April 1, 197; Herb. Inst. Bot. S. Paulo 54036] (N); Hashimoto 30h (Sp-42939); Hauft 16-35 (Sp--3059, Sp); Herter 4360 (N)j Fe G1 Hoehne s.n. [Sio B Bernardo, Oct. 23, 1918] (N, N, Sp——2535), SM. [Santo Am Amaro, Oct. 21, 1917] " (sp--758), sen. [Campos do Jord#o, Sept. 19, 1923] (N, Sp--28656); W. Hoehne 1865 (N), 2270 (Mg, Wo), 2275 (Mg), 2276 (N); Hoehne & Gehrt s.n. [Santo Angelo, Oct. ve "1938 ] (Be--39685, N, N, Sp 36577) Me M. Kuhlmann s.n. [Umua- rama, Feb. 7, 1935] (K, Sp--32hh5), su. [Santo Amaro, Aug. 5, 19133 Herb. Inst. Bot. S. icine 16360] 0] (N); Leite — (El), xxvii (N); LOfgren & Edwall s.n. [Santo Amar 0, July 29, 189k; Herb. Com, Geogr. & Geol. eat “CM, ET P. We tis SMe (Mugi, Novembr. '33] (Cp, Cp); A- Lutz 517 (Hk); Be B. Lutz O.Bs [Serra da Bocaina, Jan. 1925] pete’ Woldenke & Moldenke 19650 (Es, Mg, N, No, Ot, Sm), 19652 (Es, N, Sm); Le Netto son. (Ja=-6796) ; Pickel 1287 (N), et (Ny 32), "197 (N, Sf), 5370 Leiepentstt 5509 (N); J. E. Pohl son. [1839] 9] (Br); L- Riedel 7 (N)3 L. Roth 361 [Herb. Inst. Bot. S. Paulo 4808] ( 5 Schwacke Sen. [Mogi das Cruces, April 19, 1889] (Ja--6587)5; Us- teri s.n sen. [Herb. Inst. Bot. S. Paulo 15713] (N), son. [Modca, Oct. 2, 2, 1905] (N, Sp--15720), sn. [Vila Mariana, Aug. 26, 1906] (Sp-—- 15715). State undetermined: 1d: Bowie ningham S.Ne Lesecsieead (Bm); Ule s.n. [Herb. Rio Jan. mn. 640] (N). ARGENTINA: Niederlein 1742 (Ra—23608). CULTIVATED: Germany? eta son. [Mac- bride pate ete! oigag ata Ne-photo, N—phot oto). LOCALITY oo UNDETERMINED: Herb. Jard. Bot. Brux. Se Sr s.n. [18h] (S). VERBENA HIRTA var. GRACILIS Dusén, Arch. e 1903 Biblio graphy: Dusén, Arch. Mus. Nac. Rio Jan. 13: = ah Moldenke, Known Geogr. Distrib. Verbenac., [ed- ii. Oye Schau- Mus. Nac. Rio Jan. 13: 292 PHYTOLOGIA Vol. 9, no. 5 1942; Moldenke, Alph. List Cit. 1: 170 & 172 (196) and a: ip & e branched; the branches 20--30 cm. long, ascending, 1-~2 mm. in diameter ; agen sometimes very shortly petioletas otherwise typical in form; inflorescence terminal, less robust than in the typical fo bes The type of the variety was collected by Per Karl Hjalmar Du- sén in a campo at the hill called Pedra Assentada, at an siti ade of about 2300 meters, already past anthesis in May. Dusén 232, . below from Minas Gerais, may actually have been n collected in he state of Rio de Janeiro -- its label indicates that it was sutinties at Serra de Pedra, Serra do Itatiaia. ety has been collected on campos, shrubby area: and lilac or rose in color. Common tah recorded by Reitz are "callan radinha", "formosa sem dote", and "jurupeba", names which, however, he lists for all the species in the genus from southern rn Brazil. Herbarium material has been misidentified and distributed as Ve Spreng. and V. strigosa Chan. Weir 319 was originally iden- tified as "Verbena 5} sp. near V. lobata Vell.", but is actually very typical vy. hirta v. var. gracilis. In all, 36 herbarium specimens have been examined by me. Citations: BRAZIL: Minas Gerais: Dusén 232 (Ja—31106, S)- Parand: Ayrton de Mattos 281 (N), 59h (N)» 4595 (N)3 Dusém 297 (Ja—1u86h, Ja, S, W—1199]60), 6798 (S, W—1L81766), 8567 (N, ), 73239 (S}, 13478, in part (S); Hatschbach 512) (Sm), "6162 (Mi), 6796 (ui); Mattos s.n. [R-20, no. 9; 5-11-50] (Sm), sn. [R-20, no. 12; 18-h-51] (Sm); i x Moreira Filho s.n. [2/1960] (Ih--5910) 5 Tessmamn sen, [Herb. Mus. Paran. 2516] (N); Weir 319 (Bm). Rio Grande do Sul: Rambo 35421 (N), 5190 (N, W-—2102250). Santa Cat~ arina: Reitz C.310 [S. F. 51359] (N), C.1542 (Rd); Reitz & ne 733 (Mm), TUES Qin), 7645 SUS (um), 7934 (in)s Sait & Rett go 8 (W225 ermined: Dusén 2687 [Se errinha, Jan. 14, 190)] (Ja—L6799) » Sen. (Herb. Rio Jan. 31106] (N)- VERSENA HISPIDA Rufz & Pav., Fl. Peruv. & Chil. 1: 22, ple 3la- Synonyuy: Verbena tetrandra, ee is ternis, cylindricis; fol- oS cratic is oblongisqe, integri 1963 Moldenke, Monograph of Verbena 293 Bot. 8: 550, in syn. 1808. Verbena glandulosa Moris, Ann, Stor. Nat. 4: 39. 1830. Verbena cuneifolia Hort. ex Walp., Repert. Bot. Syst. : 22, in syn. 1845 [not V. cuneifolia Raf., 1608, nor Rufz & Pav., 1798, nor Wallberg, 1959]. Verbena glandulosa Morren ex Walp., Repert. Bot. Syst. : 33, nom. m. nud. 185. Verbena clandes- tina Mart. ex Moldenke, Alph. List Invalid Names Suppl. Suppl. 1: 23, i 23, 4B syn. 1947. reset hispida Pav. ex Moldenke, Alph. List Invalid Names Suppl. 1: sl: 2h, in syn. 19,7. Verbena bonariensis x officinal- is Regnell ex Uoldenke, Résumé 359, in sy in syn. 1959. Bibliography: Rufz & Pav., Fl. Peruv pow is - 3ha. 1798; Poir. in Lam., Encycl. “apt Bot. ‘gt 550—— oT fohs J. Sm. cl. 2: 7h9. 18253 wae Ann. Stor. Nat. h: 39. 18303 2 Don in Loud., Hort. Brit. Suppl. 2: 630. 1839 Walp., Nova Act. Acad. Nat. Caes. Leopold.—Carol. Cur. 19: suppl. 1: 38: wa Walp., Repert. Bot. 5 th < 22 & 33. 1845; Schau. in Prodr. 11: 5),2—543. oo & “28h. t. 19205 Herzog, Meded. Rijksherb. Leiden 29: 3. . Soc. Cient. Arg. Buenos Aires 88: 97, 98, ie 146, & “13k. "39193 "Jaffuel & Pirion, Revist. Chil. Hist. Nat. 25: 387. 1921; Hicken, Darwiniana 1: 33. 1922; Seckt, Rev. “Univ. Nac. Cérdoba 17: $0, 1930; Stapf, Ind. Lond. 6: h29. 1931; R. Bspino- sa, Okol. Stud. Kordillerenpfl. 3% & 39. 1932; Junell, Symb. Bot. Upsal « hz: 11—1), 16, 19, 164, 166—168, 170, Bip igh Me. > ie e He Ze Bai ° ue hae 1s, “18. een eae vg andl. Verbenac. . J ON Ww e B toe ~e WwW WL we tc _ fee) .* . 231. 1938; P. M. Hall, Rep. Bot. Exch. Club Brit. Isles 1938: 51. 1939; Robledo, Lecc. Bot. er 97. 1940; F. L. Herrera, se Cuzeo 1: 352. 19); NoldseRe., Lilloa 6: 330—331 (19/1) and a 430. 192: Schnack’ Anal. Inst. Fitotéc. Sta. Catalina h: 17—1). Darwiniana 6: 70 73. 19h; Moldenke vb bagg : 87 & 115. 19h5; Darli Pu Amal, Atlas 270. 1945; P. I. Acufla, Cat. Fl ae 1955 Sehnask Covas, Bol. Argent. Bot. 1: 283 & 28. Reet wolde e Alph. List if a) 19. 20, 28, 72 76, 79 251 & 27h. . Cc a 1s 3; ; ’ ’ ry , Teg 19k6; Moldenke. Phyto 2: 338, & 348."1987} canta, ks lordeste prov. ‘wate ee oie "122 & 150. 9hi73 buns, *glph. 29h, PHYTOLOGIA Vol. 9, no. 5 List Invalid Names 23 & 2h. 1917; H. Ne & A- L. Moldenke, Pl. Life 2: hh. 1948; Moldenke, Castanea 13: 117, 119, & 121. 1948; Mol- dale, Alph. List Cit. 2: 356, 357, 372—3th, 377, 378, 02, 4, » OT2, 69 eat — ‘Xo oO ° ge as =~ = & ww ~ Ww RIG o On oO ‘e Ww ie Ov ne) ft. ‘e ~ Oo wy oy & L6 7 869, 872, 880, 881, 883, 888, 893, 908, 913, 952, 957, & 968 and : 1010, 1079, 1088, 1090, 1091, 1093, 1133, 1158, 68, 1175, 1192, 1203, 1215, 1233, 1257, 1293, & 1302. 199; o cS »?) ~~ “ = (1949) and 3: 291 & 377. 1950; Acevedo de Vargas, Bol. Mus. Nac» Hist. Nat. Chile 25: 5). 1951; Moldenke, Phytologia 3: h67. 1951; foldenk Chi ort. Soc. Dict. Gard. : 2200 & 2211. 1951; Moldenke, Phytologia : 267 (1953), 5: 75 & 96 (195k), and 5: 133. 1955; Moldenke, Biol. Abstr. 30: 1093. 1956: i 359, 362, 36h, 366, 425, & 472. 1959; Moldenke, Résumé Suppl. 1: § (1959), 2: 11 (1960), 3: 13 & 41 (1962), and \: 5. 19625 reer, Pl. Vall. Cochab. 38. 1962; Moldenke, Résumé Suppl. 6: 6 (1963 and 1% Te 1963. Illustrations: Rufz & Pav., Fl. Peruv. & Chil. 1: pl. 3la- 1798; Sanzin, Anal. Soc. Cient. Arg. Buenos Aires 88: 116. 19195 y tall, grayish or yellowish in drying; stems 1 to ou hirtellous-pubescent on the lamina beneath, the teeth a: impressed above; iri tent cula very dense and crowded, long- or short-P te cylindric; 3 ’ a: ‘ ’ sing the c dular-hispidulous; ©#” ig 2—2.5 mm. long, glandular-hispidulous, its teeth short, acute hairy; corolla varying aad clear-blue, bluish, very pale-blue, or purplish-blue to bluish- ©, purple, lilac e-lilac, lavender, violet, pale-violet, light-violet, clear-violet, dark-violet, pinkish- rose, or "steel-color", its tube about twice as 10né 1963 Moldenke, Monograph of Verbena 295 as the calyx, the limb 5-lobed, very small and inconspicuous, the interior of the throat sometimes white, the outside of the flower or base of the tube often white and the remainder of the tube ae ET edema comma én = his comm wid ead species was collected by sea bi me (no. 1/3 31) pects torrents, in sandy soil, at Hu- anuco, Peru, and is o teres a in the herbarium of the Jardin Bot- anico at Madrid; the unnumbered Rufz & Pavon collections cited below are probably. parts of ari same collection. An isotype in the h herbariun of the Botanisches Museum at Berlin was ee in ~ nd cultivation in or about 1835 and is now know from gardens in ney s of the world. Bailey (1935) lists Kew as handling the Seeds. In its native haunts it inhabits both cultivated and unculti- cities, at the edge of montes, fertile campos, — places, san- dy playas, among rocks, in ary stony places, mountains, dry river valleys, campos , among weedy vegetation i hard gravel- ly ground, along ee sa of old canals, at the foot of nas rancas, near or betw cultivated areas and in old fields, growing at Bee ee 10: token wate Tt Uae ee collected in anthesis in every month of the year, and in fruit in er d from July to December. liandon asserts that it grows tivated ground. Ruiz tani states that it is ab wet mat- torales in the mountains, Eyerdam t associated with Stipa, Juncus, Salvia, etc., among stones in red cl Rauh & Hirsch ating it in the Adesmia zone and i Rojas 13561, from Paraguay, is said to have "petals white Side of tube pale lilac". Stafford 1206, at Kew, from an aititac “ 10,500 feet at Arequipa, ai Pikes that are not so densely Material of this species has been misidentified and ee in herbaria under the names V. alata Cham., V. bonariensis Le, 296 PHYTOLOGIA Vol. 9, no. 5 bonariensis var. SSeS Kuntze, Ve cuneifolia Rufz & Pave, V. gracilis Cham., V. hastata L., Ve lasiostachys Link, V. litor- alis Kunth, V. Hiteets EE eh VY. littoralis Kunth, V. rigida var. hanes (Gill. & Hook.) Farwell, and V. venosa Gill. & Hook, On the other hand, the Buchtien s.n. Waa iia, and Venturi 525, distributed as V. hispida , are V. bonariensis L.; C4rdenas 3019 is V. brasiliensis a Vell.; oan ieean “803 35, and He Krause s Sone n. [Coral] are V. corymbosa Rufz & Pav.; Soukup up 259 is Vv. hayekii | Moldenke; Killip & Smith 21925 and Pérez Arbeldez 1207 are V. litoralis H.B.K.; ten 135 & 5887, Herrera S-N- [cuzco, July 1923], Killip & Smith 215), 21855, & 2212, and Ledig 33 are V. parvula Hayek; and Rose & Rose 19072 is V. villifolia Hayek. Junell (193k) discusses and illustrates the gynoecium morpho- logy of V. hispida. Common names recorded for the species are "bristly vervain", "hairy verbena", "verbena", and "vyerveine hispide", The type of V. clandestina is Herb. Martius s.n., collected fron material grown in the botanical garden at Munich, Germany, in 1839, and de Sicetien in the herbarium of the Jardin Botanique de 1'Etat at Brussels. The type of V. bonariensis x officinalis is Regnell s.n., collected in August, 1897, from material grown in a garden in Sweden and deposited in the herbarium of the Nat- urhistoriska Riksmuseum at Stockholm. It is aged of note here that the V. cuneifolia of Rafinesque, referred to in the synonyuly, is V. stricta Vent., that of Wallberg is V. simplex Lelm., while that of Rufz & & Pavon is a valid st fai Gay (1849) reports that "Esta es muy comun en las provincias centrales, Val , ete. Florece en set~ iembre, etc." Reiche (1910) says "Desde el Pert al traves del ee ta pro ntrales; tambien en las te cordilleras bajas." Its hybrid with V. bonariensis L. is xV. der- meni Moldenke, with VY. hastata L. is x xV. noacki Moldenke, with V. etrix Moldenke, with v. simplex Lehm. is xV. prostibula Moldenke, and with V. urticifolia L. is xV. fabricata Moldenke, which se®- Walpers (18L5) cites Poeppig -I.158 "in cultis" near Concan, Chile, and Schauer (187) says of this collection "forma fo liis eximie laciniat + i §-n. from Bolivia; Mertens s.n., Philippi s.n., and J. Styles Se n. from Valparats so, Chile; Bertero s.n. from Q ota and from Santiago, Chile; and Meyen s.n. from Mellipilla. Herrera (19h) eee Hicken wit ee and Herrera s.n. (Oct. 1929] from Cuzco, - Raimondi (1943) cites his nos. 7372, and 7727 from Cajamarca, 9587 and 10513 from Puno, nes eee neo, an and et 10693 from an undetermined department, = 2 ent, Peru. Cabrera (19 1963 Moldenke, Monograph of Verbena 297 cites Keidel s.n. from Jujuy, Argentina. The species is recorded by P. Me M. Hall ~ (1939) from Jackdaw Lane, Oxford, where it was col- oe by J. P. M. Brenan and J. Chapple, and is o recorded as uralized in England by Douglas H. Kent ina letter to me dated Sy at 28, 1949. Maria (1962) cites is no, 181/d from Cocha- bamba, Bo livia. The Lo ud., Hort. Brit. Suppl. reference given in pri bibliograpty o of this Hear ie is sometimes erroneously accred- =~ . Acevedo de Vargas (i991) ete the following 13 specimens not as yet seen by me: CHILE: meagua: Landbeck s.n. [XII.1860] (Sg—5h692, Sg--5695). Hcl ha, Reiche sen. [1.1899] (Sg—68332) Linares: Collector undesignated sen. [1.1877] (Sg--42309, Se— 54693). Maule: Collector undesignated sn. [Constitucién] (Se— 68329), sen. (Sg—5l696). O'Higgins: Bertero sen. [IV-V.1828] (Sg—5691). Santiago: Collector undesignated sen. [X1.1895] (Se- oso. Sg—Sl69h). Tarapacd: Kuschel sen. [5-X11.19k6] (Se— 201,) ; Rahmer s.n. [III.1885] (Sg——lj2308). Valparaiso: Buchtien iio (Sg—68331). In all, 216 herbarium specimens, including type material of all the nanes involved, and 1) mounted photographs have been ex- amined by m pistince ECUADOR: Azuay: Asplund 17605 17805 (S); ——- 23 [S-- 2059686). PERU: Arequipa: Munz 15/87 (N, N, Po—255349); Sandeman 3793 (K), 3850 (Kk); D. Stafford 622 (K), 1206 (K). adhe Rose & Rose 19072 (N) (N). Hu4nuco: Pavon | sen, [Macbride photos 21,686] (K (Kr—- photo, N—photo); H. Rufz 1/3k [Macbride photos 29686] (Kr—photo of type, N--photo of type, Q—-type), sn. [Peruvia et Chilis; Mac- ore phate 17555] (Br--isotype, F--photo of isotype, gow of +; si—phi otype, N—-photo of isotype, N--photo of iso ole , of yield 2 ants of isotype) . Junin: So 18 (W- 1561077). oe 1310869); ; We Lima: Kunkel 388 (Gg); F. W. Pennell 14562 (N, S, E. Safford s.n. [Rimac, Aug. 29, 1087] (C)s Wilkes sme gee oa To] (W—65025). Puno: Sh am 11 (N); Soukup sn. sen. [July 8, 1938] (Ew), Tacna: H. H. 09 (C, Mi, Pa). a). Department undete ed: Haenke 1762 (N); fe 497 [Peruvia interiore, Meyen s.n. (Br); Rauh & Hirsch h P.592 [Chiquate] (Z)- BRAZIL: So Paulo: Pickel 9h6 (Sf, W—1564303), son. [Herb. Serv. Flores’ S. Paulo 995] (N). BOLIVIA: Cochabamba: piers 18 (Ko); Hyerdam aus (Ca—6),9066); A. Jimenez 96 (W—2369935)5 JuLlo itec— 1512919); Senn pe (ii, W—-2031,330); Steinbach 8729 (W—-15574h0) . a Buchtien 5888 (W—1159357)- La ein Ua Fes: Asplund 22 i re (Us), 938 (Us), 3750 (S, Us); M. Bang 965) ; aoe Rs ee (V), 196 Hs 128 3 (V, W—1159356) , 4683. (H— Linas en. f Sen. [La Paz, Mirz 191 » 5, Ug)3 Hamnarlund 167 (8), 196 (S), 516 (S)5 Raine (S)s "Re So Millions 5 (WH) 298 PHYTOLOGIA Vol. 9, noe 5 Oruro: Asplund 3352 (Us). Potos{: C4rdenas 327 (W--157331)). Tar- ija: Hammarlund | 288 (S). Titicaca Island: Bandelier 13 (N). PARA GUAY: Hassler 2459 (N, V); Pedersen 4,018 (W—22836L)) Te Satan 313 (S), 355 (B, Bm), 1890 [Herb. Hort. Parag. 10063; Herb. Osten 13561] (N, Ug, W—1858377), 251) [Herb. Osten 18176] (N, dag? 13561 (S), sen. [Hassler 2459] (S)» CHILE: Aconcagua: Reich (N). Arica: Jaffuel 1625 (W—16178l)6). Colchagua: Barros Valenzu- ela 8048 (N), 8049 (N); Bertero s.n. [Racangua] (Br). rr). Concepcion: Barros Valenzuela 822 (N). San Santiago: Re A. Philippi 1 1300 (W— 1323096); Sparre 2346 (S); Wilkes s.n. “Tsantiago] (W—-71952) » Val- afso: Buchtien s.n. [18.1X.1895] (W--1159366); Claude-Joseph 3549 (W--1283380); Garaventa 2106 [Herb. Looser 027] (N)s Looser 4505 (N). Province undetermined: | Cuming 417 (Br, F--photo, Wee photo, Si—photo, Z—photo), sen. [1035] (Br). ARGENTINA: Cata~ marca: JOrgensen 1021 (Herb. Osten 1127] (Ca—195443, Ug, W—- 704748); Venturi 7229 (Gg--16138)). Chaco: Buratovich 17 (N)5 Pa rodi 12811 (N); A. G. Schulz 1482 (N), 6387 (N). Cérdoba: Burkart ae (N)5 Gesvellance SDs, sen. [Herb. Osten 15285] (Ug); E. Fielding sen. [Cordova] (Bm); Osten 10611 (Ug); zrongees Ed (N), 6025 (W- 1558367), Villafane 171 (N). Formosa: J&rgens 32 (W--1065618), 2638 (Herb. Inst. Miguel Lillo 31322] (N, N); ce os 77 (N), 25 (N), 311 (N), 1177 (N), 1854 (N); Pierotta L080 Bat PSS Claren 11558 (s); R. E. Fries 509 (N, $); Garolera & R (N); Kuntze s.n. [Jujuy, Oct. 1692] (N, W-—701710); Moldenke sjalions mke & Moldenke 19750 (Lg, N); Venturi 5370 (Ca--376)16, Du——372500, Ge- 158645, Vi, oho ate Sha “Wendoza: Dawson & Pujals 19 (N)j Jen a oe a . [Santa Rosa, 190] (W-—-108108)); Ruiz iz Leal 1172 (N), 2026 (HN), | “305, (MN), $593 (M), 6761 (w), 8422 (MN), 8800 (Ns 8987 (N), 90h9 (N), cs Ws 9337 (N); Semper son. [7-V-19h] N). Misiones: Ekman 2027 (S). Salta: Garolera & & Romero see [17- I-1947] (N), sen. “ag eae (N); Lillo 11013 [Herb. 0 Osten 8167] (Ug); T. Meyer 3uk6 (Herb. Inst. Miguel Lillo 1733k) (Cca-88287% N), 4870 (Ca—90660, N, Si); Moldenke & Moldenke 19735 (Es, 1g» Ws OrDoned Lc 4801 03 D. Rodriguez 161 [Herb. Inst. %. Miguel nee (Ca—3315, N, W--1858262 S, S); Venturi 20° (Da STAI, Wo ASLO) Ea eS AT Se ON ar Salt, 4/12/h6] (Ew). San Juan: Castellanos 15191 2 a eS 1596 (N), Santa Fé: Balegno 496 (N)3; C. Meyer Ze (Tr. Meyer 3273] We Tucumfn: Baer 7 (S); Bruch 272), (N); Lillo 5 a “Seten : aie, ] Ge): 220 5201 [i ipolens Miguel Lillo raise (8), om ’ Herb. Osten 8 15290 [Herb. 08- ten 8453] ire Monetti 359 [Herb. ee abeee} (Ug), 1807 beg ao ee Lillo 31319] (N), 3015 [Herb. Inst. Miguel Li } (N)5 Schreiter 2359 [Herb. Inst. Miguel Lillo 32785] (x, N), 1963 Moldenke, Monograph of Verbena 299 sen. [Herb. Osten 12179] (Ug); Venturi 53 [Herb. Osten 1721] (Ug, W—1591210), 520 (W—1591222), 825 (Herb. Inst. Miguel Lillo 31331] (Ca—-3311, N, N), 2767 (W--1591267). GERMANY: E. H. L. Krause 29135 (B), 31938 (B). SWITZERLAND: Aellen s.n. [Basel, Schutt bein Klein-Riehen] (Go); Collector undesignated sen. (W— 261440). CULTIVATED: Belgium: M. Martens s.n. [hort. insul. 1837] (Br), s.n, [h. b. lov. 180] (Br). England: Nelmes 1202 (Ba). France: He: Herb. Harvey son. [h. Re P. 1835] (Du——166);72) » sen. [he Re P. 183] (Du—16671), Sone a P. Septb. 189] (Du—166)69); Herb. Jard. Bot. Brux. s.n. . (Br). Germany: Herb, Gtting. 2109 (S); Herb. Martius son. [hort. Monac. 1836] (Br, Br), 5.n. ~ [he Monac. 1839] (Br), sen. [hort. Monac. 18)3] (Br). Italy: Herb. Harvey s.n. [Bellovaco ex seminib. h. R. P. 181] (Du—166;70) « Massachusetts: L. H. Bailey s.n. (Cambridge, June 21—23, 1919] (Ba), san. (Cambridge, Aug. 31, 1921] (Ba), sn. (Cambridge, August 6--12, 1929] (Ba). New Jersey: Guyot s.n. (Pr). New York: Bastedo s.n, [22 Au. 1897] (N); Burdic k sn. (Sept. 26, 1922] (Ba). S ). Sweden: Blom s.n. [1929] (Go); Collector unde signated sen. Sen [e semina Helsingfors in H. L.] (Lu); Regnell s.n. [8/1897] On Vestergren s.n. [25/10/1923] (S). switzerland: aes Hort. Bot. Basil. sen. [5.7br.180] (M). TY OF CO ON DNDET: ED: Beyrich s.n. [1833] (S); Herb. Meisner s.n. (15 .7he 2) (ui); Née 110 (Q); E. W. White 2), [Yacusula] la] (Bn). = eee (Covas & Schnack) Moldenke, Phytologia 2: 118. egayi Verbena crithmifolia var. minor rele & Hook. in Hook., Bot. Misc. 1: 169. 1829. Verbena crithmifolia g minor Walp., Repert. Bot. Syst. : 30. 1845. Verbena cri olia ® minor Schau. in A. DC., Prodr. ll: 556. 18h7. “i8h7. Glandularia hook- eriana Covas & Schnack, Revist. argent. Agron. 12: 57—59, fig. 1. 1945. Verbena crithmifolia f. minor Gill. & Hook. ex Covas & Schnack, Revist. Argent. Agron. 12: 59, in syn 1945. Verbena crithmifolia g minor Gill. & Hook. ex Moldenke, Résumé 363, in syn. 1959. Verbena erythmifolia Gill. & Hook. 6x Moldenke, Résu- mé Suppl. 6: 10, i 10, in syn. 1963. Verbena eurythmifolia Gill. & Hook, ex Moldenke, Résume Suppl. 6: 10, in syn = 1963. — Biblio erepty: Hook., mee) Misc. l: 169, 93 Ma eee TBhT; Bot, be 18. chau. in A. DC. : at nt ’ neg ecg Ua Ar nt. Agron. to: e7—59, fig. i 19h5; Cabrera, Bol. Argent. Soc. Bot. 1: Sy Pe s k & age: 7 ’ 9. hy 75 Suppl. - 10. 1 197; He Ne . Moldenke, Moldenke, Alph. risk OH: 3 oes 735, 896, 303, 910, & 951 300 PHYTOLOGIA Vol. 9, noe 5 (1948) and li: 1172, 1196, 1197, & 1200—1202. 198; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 106 & 198. 19493; Mol- m Mold. Set 48 Spec. [h]. 1954; Moldenke, Résumé 127, 296, 363, h72. 1959; Moldenke, Résumé Suppl. 3: 1h & 37 (1962) and 6: 7. Illustrations: Covas & Schnack, Revist. Argent. Agron. 12: 58, fie. 4s ‘ Prostrate spreading herb or else subshrub or shrub to 80 cm. tall; stems sometimes very woody, cylindric-tetragonal, pubescent : ng intermix- ed with glanduliferous hairs about 0.05 mm. long; leaves decus~ sate-opposite; petioles 0.8--1.2 cm. long; leaf-blades ovate in outline, 2.5-- cm. long, 0.8--2.5 cm. wide, laciniate, pubescent, at mm. wide, obt + revolute along the margins, irs simple, subappressed, abo - long, intermixed with 4 very few s gl rescence spicate, the spikes breviated, soli elongate after anthesis; ciliate on the margins with simple hairs about 0.2 mm. long, pi- lose on the back and margins with glanduliferous hairs about 0.05 flowers very fragrant; calyx tubular, 6—7 mm. long, s er: 2n = 20. The of this species was collected by Guillermo Covas (n0e aguadero, in the department of La Paz, : gentina, on December 15, 1943, and is deposited in = erbarium of the Instituto de Bot4nica Darwinion at San Isidroe | Walpers (185) and Schauer (1847) describe their V- crithmifo™ ia p minor as differing from typical V. crithmifolia in having 1963 Moldenke, Monograph of Verbena 301 tionship to the species which he suggests except for a very sup- erficial leaf-resemblance. Covas & Schnack (195) make Wg following comments: "Esta es- pecie es affn a G. crithmifolia (Gill, et Hook.) Schnack et Covas, de la cual fué considerada como una forma por Gillies y epigabae diferencias entre ambas entidades son lo suficientemente cadas como para considerarlas especies distintas. Ambas pue orig diferenciarse asf: A. Hojas con la s generalmente menores de 2 mm. de ancho; pe- los foliares dibioest e« An superiores sin apéndices ola en su mayor parte glandulosos. Mericarpios de largo con el 4pice recurvado hacia la cara ventrales.seesssee Ge co AA. Hojas con lacinias generalmente mayores de 2,5 mm. de anc Pelos foliares erectos, rigidos, muy cortos, dando a la te mm, de bane con el Apice raped seit gata thmifolia." They comment further: "G. aicbaeiacs y Ge oy cas tienen 2n chrom siendo io ellos - menor - tamafio que en las ni at diploides a Blacdeiarte (2n = 10).....Al parecer las 4reas dispersién de G. Hookeriana y de G. crithmifolia no se superponen. G. Hookeriana crece en las sierras is de Cérdoba y San Luis y en la llanura la regién limftrofe de Mendoza y San Luis. G. crithmifol- ia crece, dentro de la Prov. de Mendoza, en la regién precordill- erana y en la llanura adyacente." Verbena hookeriana inhabits campos, barrancas, flat uplands, and dry land, often in the mountains, and has been found at alti~ tudes of 50 to “3500 meters, blooming ” from September to May and in Jwy, fruiting from January to May and in November. Bartlett found it "in the uplant poe 7 called 'monte de cal- den'", in loess soil of flat cuneifolia association", and growing with Gomphrena and ices megapotamicus. Eyerdam, Beetle, & Grondona state that it forms tee paces sometimes — abundant over wide areas in semi-open fiel a ee a ering al4, Catamarca; Kurtz says that it is very common in dry ground at Los Romeros, Cérdoba; Senn found it ii comm La Pampa; and Hunziker reports it as de Agua, Santiago del Estero. A common "cola leche", Herbarium material has been misidentified and distributed un- der such names as V. crinoides Lam., Ve crithmifolia Gill., V. crithmifolia Gill. & Hook., V. critmifolia Gill, V« erinoides 302 PHYTOLOGIA Vol. 9, no. 5 Lam., and Glandularia laciniata (L.) Schnack & Covas. Covas & Schnack (1945) cite also a Ruiz Leal s.n. [Herb. Esc. Agron. Men- doza 3098] from Mendoza, not as yet seen een by me. Chicchi 28 exhib- its very woody stems, while Vervoorst 1,252 and Sanzin 1563 show very plainly the upper stamens without the glandular appendages so rent as in V. crithmifolia » 137 he barium specimens, including type ee of most se the names involved, have been examined by m Citations: ARGENTINA: Buenos Aires: H. He Batiste 19918 (Ca-— 772196, Mi, W—1904717); Castellanos s.n. [Herb. Osten 15287] (Ug); wee 28 vd oid Eyerdan, Beetle, & ao one Ee N), 516 tN), 870 (N), 9 978 (N), 9 996 (N), 1257 (MN); * Brttcher & Brt- cher s.n. [25/12/48] (S), s S.n. [28/12/48] ] Gas Castillén 1210 [Herb. Osten 849] (Ug), 1212 [Herb. Osten 88] (Ug), SMe (Que- brada del Tala; Herb. Inst. Miguel Lillo 31738] (N), S-M- n. [Herbe nst. Miguel Lillo 31740] (Du—317596, N, N); Hessling & Barkley 19hr619 (N); Job 1337 (N); JUrgensen 1019 [Herb. Inst. Miguel Miguel Lillo 32795 Herb. Osten 92969) (N, N, soe Se [Herb. Inst. eae Lillo 32861] (N, N, W—71974), sen. (Herb. Inst. Mig uel Lille 86242] (ca, Ca—16l80l)); Soriano 8h7 (N); Venturi 6729 esac: Vervoorst 252 (Le); Wall & Sparre s.Me Andalgala, 27/11/46] (Ew, Ew, Ew). Chubut: Scolnik 358 (Er, N)- N). Cérdoba: He H. Bartlett 20149 (Ca—772h77, Mi, N, W—1904869); Burkart 2091 (Av—175]68); Cabrera 6391 (N); pA We sen. [Herb. Mus. Are Cienc. Nat. 31202] (N); A. Wee Hunziker 77h a5 Isler 117a e-. 1473338); F. Kurtz 840 (N), sen. [Los Romeros, Jan. fan. 18, 1 j, 1895) (Ja--1886); Lorentz s.n. [Herb. Krause 16831] *(ca—liS3755) Sede [Stdlich von El Chaflar, 28 Nov. 1871] (W--1134903); Lorentz & Hieronymus s.n. [1873] (C); Lossen 9 (Um—-52); O'Donell 474? 1525 (Ns O'Donell & Rodriguez 358 TERY. 597 (Ut——115L20b) 5 ) 5 Sota —— a ane Herb. ius. ‘Argent. Ciene. Nat. 33882] (N, oy MU. Pe Gomez 5.n. [Iliar; Herb. Inst. Miguel Lillo 10684] (Ga--16L602, 8); ; Become & Niederlein s.n. [Herb. Osten een said N, N, W—19091h1), 1286 (ca—79153, Ca—90 933, Ns Venturt 766 (W—1591392). Mendozar Ruiz Leal 8990 (N), 2400 (™)s 1963 Moldenke, Monograph of Verbena 303 9510 (N). Rfo Negro: H. H. Bartlett 19955 (Ca--772025, Mi, W— at O'Donell 1553 (W—1931)65), 1693 (S); Scala 11 (N, Ug, Ug). Salta: Venturi 6939 (W--1591505) . San Juan: H. “He He . Bartlett 19550 (Mi, W—190L559). San Luis: Gillies s.n. [Sn. Luis] (Lu); Kuntze s.n. the José de Morro, 3/1/92] (N, N, i: W—701713), Miers sen. [Rio Quinto] (Bm); Ruiz Leal 10h) (N); Sanzin — Look Osten 1,639] ] (Ug); Terribile 765 (N); Troncoso 1858386); J. De « Varsta TST Ci ), 48h (Ge—35: 5268, 3), ‘Sa , 5 aut (Ss), ao (S), 653 (S), eo (S). Santiago del Estero: A. T. 1 (N). Province undetermined: Jameson SMe {in R Bey Argentina, 18),6--69 ] (RRL VERBENA HUMIFUSA Cham., Linnaea 7: 271-272. 1832. Synonymy: Verbena chamaedrifolia Briq., Arkiv Bote 2 (10): 10, in syn. 190) [not V. V. chamaedrifolia Juss., 1821, nor L., 1878]. Tecan reineckii Moldenke, Phytologia 1: 480. 190. Verbena hunitusa Augusto, Fl. Rio Grande do Sul 211, sphalm. 196. Verbe- ha reineckii (Briq.) Moldenke, Résume Suppl. 3: 40, in syn. 1962. Bibliography: Cham., Linnaea 7: irae og 1832; arte ae pee Pro 187; aos in Mart., Fl. Bras. 3 18h. 18 Baa: in rane 105. stant, Ind. Lond. y# 129» 1331; uel sacks, Phytologia 1: 180 "Ex9u0) and 1: 511. 191; Moldenke, Lilloa 6: 331. 191; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 39, Wi, & 101. 192; Augusto, Fl. Grande do Sul 209, 211, & 232, fig. 96. 1963 Moldenke, Alph. List ‘age 1: 26h. 1946; Moldenke, — L Ans Tee ames Suppl. 1: 24. 19)7; Hill & Salisb., Ind : 1907 Moldenke, Castanea 33: DLT. 1gh8; ane Alph. pare on : 467, 485, "gon (1948), 3: £08, Oe ou & £983, C9kS) 5 see h: Tish, 1286, & 1303. 199; Moldenke, i Sable 100, & 98. 1995 ee orm. Rambo, Sellowia 6: 60, 8h, & 153. 195k; Angely, s 126 1957; Moldenke, Résumé 110, 118, 119, 366, 373, & @ ire. = 19595 hs gely, Fl. Paran. 16: 78 (1960) and 17: 46. 1961; Moldenke, R mé Suppl. 3: 37 & 0. 19 2. Illustrations: Lindman, Veget. Rio Grande do Sul 110. 170035 Augusto, Fl. Rio Grande do Sul fig. 96. 196. fruticose-decumbent herb, creeping; stems slender we nig form, often whip-like, very long, prostrate, ascending at th tips, more or less hirsutulous = puberul tellous, less so in age; nodes y nodes more or less elongate, Oita greatly so, us 1 fx Ong; branches ascending; leaves decussate-opposite; petiole ae very slender, to 5 mm. Sig and sparsely hirst sutulous, or obsolete; 304 PHYTOLOGIA Vol. 9, no. 5 leaf~blades chartaceous, rather uniformly bright-green on both surfaces, ovate or ovate- elliptic or even subrhomboid, 0.5--2.5 cm. lo ~—2 cm cute or subacute at pex or some~ times rounded-obtuse, cuneate~atternmate into the petiole at the b entire-margined, widened toward the apex, sp scabrous=pilose with bulbous—based s ove or strigose-hispid gins, hirs ous or strigose-hispid along the vena~ tion and g beneath, the ins coarsely dentate or crenate- serrate towar e apex, nee deeply laciniate or trilobate with broad obtuse lobes, the be 3-5 , the later ones usually 2— antate H aaah 3-plinerved, the midrib and the cate, all impressed above, sacs pa prominulous beneath; veinlet reticulation very sparse, usually obscure; inflorescence axillary and terminal, subcapitate-spicate, 8—15-flowered; peduncles slender, hirsutulous, solitary or ternate, 1—2.5 cm. long, usu- lets lanceolate or ovate-lanceolate, about 1/3 as long as the calyx, about ) mm and 1 mm. wide, hirsutulous, pie uate at the apex; flowers subopposite, patulous , eventual rs separated by distinct sympodia; calyx tu ae ’ hi colored, herbaceous-costate, the ribs ending in very ere - 7 often unequal mucros; corolla varying from blue to violet 1 cm, in ameter, lobed, the lobes emarginate; style sometimes 5 scarcely. exserted, sometimes long~exserted from the calyx after the get " yx t d bout i mm. long, included by the fruiting-calyx; i pale, biculate~areolate on the dorsal Set Py the com= ate. _ This species was based on several collections made by Fried- lected in grassy fields, dry grassy fields, on sunny hills, in dry sandy-gravelly soil’on hills, and on campos, flowering ‘from September to No iced (1845) erin the species in his ‘open verte sa n Inermes, Group Foliosae, Subgroup Macran Sec ondary seating Aubletia, with ten other species. aa (1895) places it in his Section on Verbenaca, Subsection Nobiles. The ve Chamaedrifolia of Jussieu and of Linnaeus, cited in qin the synonymy above, are both synonyms 8a var. reticulata Moldenke is a synonym of V . marrubioides eee: Herbarium material of V. humifusa has been a misidentified and distributed under V. chamaedrifolia Juss., "V. chamae- — the names Grifolia Juss. forma, V. erinoides Lam., V. marrubioides ; Chamtey 1963 Moldenke, Monograph of Verbena 305 and V. supina L. On the other hand, the Dusén 1571) distributed yumifusa i actually V. marrubioides Chan. __ bret Ta50l,) comments as follows: "C'est & tort que nous a- ns jadis réuni Soha espéce (Reineck et Czermak, Pl. exsicc. soar merid. n. bet Fe ec le = chamaedrifolia. Elle s'en distin- gue en effet suffisamment s tiges couchées ou rampantes, illes ProgensGaent eri lobeee et incisées -crénulées, son calice & c8tes longuement poilues-coliées, 4 poils m@lés de quel- i ac stipitées, 4 champs int erneuraux glabrescente, etc. Augusto (196) cites Augusto & Edésio s.n. Nes de Gléria, Porto Alegre] from Rio Desine do Sul, and spells Carl Axel Magnus Lindman's surname Ser ndman 1283 is cited by Briquet Fig eg as "Exp. I. Regn. sige? In all, 29 herbarium specimens and 5 “nounted photographs, in- cluding Sins material or phototypes of all the names involved have been examined b i Citations: BRAZIL: o Grande do Sul: Collector undesignated 554 (Ja--h659h:) ; Henz Ee (N, Rb); Jurgens 20 (Ja—17763) 5 Leite 662 (N); Lindman 283 (Ja--28163), A283 -(S, S); Moldenk ter 19855 (i); Rano 29105 (N), 43261 (), 13845 (N), TB a N), 48911 (N O13 (Im, N N), 37298 S)3 Reineck & — (N), 48911 Os Aaa Ait Blas 58 (5) 5 Bee BG, erb. Osten i160] (N, N-—photo, h, = Ug). a gel orca ey Sellow s.n. setae de Brasil merid- ional; pers nee 17421] (Kr—photo of cotype, N—photo of cotype, N— of cotype). PARAGUAY: Hassler 9479 (Bm, N, N, 8, Vv). a ge < = . praenin’ sen. [Monte Video, 3852] (Ss). HYBRIDA Voss ex Rttmpler in Vilm ed. 1, 1263--126l, [as "V. hybrida Hort.."]. 71873 rae Ye V. V. bybrida Bicknell 191]. nymy: Verbena buistii Harrison, Floricult. Cab. 8: 160. 180. Verbena hendersonii Hort. ex Harrison, Floricult. Cab. 8: 160 & 18. 180. Verbena teucrioides var. anais Univ. 3: 9. 1842. Verbena teucrioides var . anats Lem., Univ. 3: 9. 182. Verbena melindres Tati folia ja Bohn, Florists Journ. 5: hl. 184. Verbena anaeraeeee Bohn, Florists! Journ. 5: 41, fig. 1. 18h. Verbena hybrida var. vriculiflora Hort. ex i Verbena hybrida var. Hort. ex Vilm., Illustr. Blumeng., ed. l, 1265. 5. 1873- Vi i 28: 372. 1879. Verbena Lo — pride Hort. ex ngl. Svensk. Vetens .~Akad. ° (4): 9 & 16—17, pl. 1. 188). Verbena hybrida grandiflora elegans Regel, Gartenfl. 36: 1887. Verbena ida var. auriculae- brida erecta compacta Blanc, emeagere Verbenas. 1888. Verbena grandiflora auriculaeflora Blanc, Mammoth Verbenas. 1858. Verbena 306 PHYTOLOGIA Vol. 9, noe 5 hybrida var. atrocaerulea oculata Hort. ex Vilm., Fleurs Pleine Terre, ed. 3, Suppl. 195. 189). Verbena hybrida var. a fleur d'auricule Hort. ex Vilm., Fleurs Pleine Terre, ed. 3, Suppl. 195. 189. Verbena hybrida alba Haage & Schmidt, Cat. Général 1898: 93. 1898. Verbena hybrida atroviolacea Haage & Schmidt, Cat. Gén- éral 1898: 9h. 1998. Verbena hybrida candidissima Haage & Schmidt, Cat. Général 1898: 9). 1898. Verbena hybrida candidissima fol. aureis Haage & Schmidt, Cat. Général 1898: 9h. 1898. Verbena hy- brida carminea foliis aureis Haage & Schmidt, Cat. Général 1898: 9h. 1898. Verbena hybrida cinnabarina oculata Haage & Schmidt, Cat. Général 1898: 9. 1898. verbena hybrida coccinea Haage & Schmidt, Cat. Général 1898: 94. 1898. Verbena hybrida coccinea fol, aureis Haage & Schmidt, Cat. Général 1898: 9h. 1898. Verbena hybrida coccinea oculata Haage & Schmidt, Cat. Général 1893: 94. 1898. Verbena hybrida coerulea Haage & Schmidt, Cat. Général 1898: 9h. 1898. Verbena hybrida coerulea oculata Haage & Schmidt, Cat. Général 1898: 94. 1898. Verbena hybrida compacta Haage & Schmidt, Cat. Général 1898: 9. 1898. Verbena hybrida compacta candidissimé Haage & Schmidt, Cat. Général 1898: 9. 1898. Verbena hybrida ca pacta cupreata Haage & Schmidt, Cat. Général 1898: 9h. 1898. Ver~ bena hybrida compacta Défiance Haage & Schmidt, Cat. Général 1898: 94. 1898. Verbena hybrida compacta erecta atrosanguinea Haag° & Schmidt, Cat. Général 1898: 94. 1898. Verbena hybrida com s recta carminea Haage & Schmidt, Cat. Général 1898: 9h. 1898. OF bena hybrida compacta erecta coccinea Haage & Schmidt, Cat- Gén- éral 1898: 9. 1898. Verbena hybrida compacta erecta coccinea ocm ulata Haage & Schmidt, Cat. Général 1898: 9). 1898. Verbena hybri- da compacta erecta violacea Haage & Schmidt, Cat. Général 1898: 9u. 1898. Verbena hybrida compacta rosea Haage & Schmidt, Cat. Général 1898: 9. 1698. Verbena hybrida compacta violacea oculata Haage syne ocusate & Schmidt, Cat. Général 1898: 9. 1898. Verbena hybrida Dé fiance purpurea Haage & Schmidt, Cat. Général 1898: 9. 1898. Ver bena hybrida foliis aureis Haage & Schmidt, Cat. Général 1898: 9h. 1898. Verbena hybrida grandiflora Haage & Schmidt, Cat. Général 1898: oh, in syn. (1898) ; Michell, Grower's Cat. 19h: 23. 19h. Verbena fybrida lutea Haage & Schmidt, Cat. Général 1898: 9) 1898. Verbena hybrida nigro-violacea albo-oculata Haage & schmidt, Cat. Général 1898: 9]. 1898. Verbena hybrida striata Haage & cei i Cat. Général 1898: 9l,. 1890. Verbena hybrida compacts pacta , Gartenfl. 51: 556, fig. 120. 1902. Verbena hybrid® com= Scharlachrosa Wittmack, Gartenfl. 51: 557. 1902- Verbena hy- ee ee eee Ber Hot Belz. 29s 5-72 1903. verbe- Hc miteae Vilm. in Vilm. Andr. & Cie, Fleure Pleine Terre, ede 1963 Moldenke, Monograph of Verbena 307 5, 1126. 1909. Verbena hybrida compacta lutescens Haage & Schmidt ex Heinemann, Rev. Hort. pen 35: 45. 1909. Verbena hybrida com- pacta vesta Hort. ex Heinemann, Rev. Hort. Belg. 37: 72. i9ll. Verbena hybrida gigantea aaa Rev. Hort. Belg. 37: 36h. 1911. Verbena hybrida alba Thompson & Morgan, Descript. Seed List 1930: 94. 1930. Verbena hybrida caerulea Thompson & Morgan, Des- cript. Seed List 1930: 9. 1930. Verbena hybrida purpurea Thomp— son & Morgan, Descript. Seed List 1930: 9h. 1930. Br SS rosea Thompson & Morgan, Descript. Seed List 1930: “9h. 1930+ Ver- bena hybrida colossea Thompeon & Morgan, Descript. Seed “ae 1930: 9h. 1930. Verbena hybrida compacta Hort. ex Stapf, Ind. Lond. 6: 429. 1931. Verbena hybrida compacta lutescens Hort. ex Stapf, Ind. Lond. 6: 430. 1931. Verbena hybrida elegans Hort. ex Stapf, Ind. Lond, 6: 430. 1931. Verbena hybrida grandiflora elegans Hort. ex : 430 Stapf, Ind. Lond. 6: 31. 1931. Verbena hyb: Haage & Schmidt ex L. H. Bailey, Cat. “Cat. Florists Handl. Verbenac. mss. 1935; Moldenke, Alph. List Invalid Names Suppl. 1: 2h, in = 1. he a purpurmantei Haage & Schmidt ex L. H. Ba ley, Cat. Florists Handl. Ver rae mss. 1935; Moldenke, Alph. List Invalid Names saat 1: oh aa . 1947. Verbena grandiflora Bodger ex Harrow, Journ. Roy. Hort. Lt 61: 399, 1936 [not V. Grandi flora Michx., 1821, nor Ort., 1797, nor Sessé & Moc., 1089, nor Steud., 1895]. Verbena lutea Waller-Franklin ex Harrow, Journ. Roy. Hort. Soc. 61: 399. 1936. Verbena luminosa Waller-Franklin ex Harrow, Journ, Roy. Hort. Soc. 61: 399. 1936. Verbena rosea stel- sate Bodger ex Harrow, Journ. Roy. Hort. Soc. 61: 399. 1936. Ver- — coccinea Bodger ex Harrow, Journ. Roy. Hort. ey 61: 00. 1936 {not V. coccinea Waterer, "19471 « Verbena violacea stellate Bodger ex Harrow, Journ. Roy. Hort. Soc. 61: 00. oe. . Verbena auricula Park, Illustr. Flower Book 48. 1937. Verbena striata Park, Tilustr.. Flower Book l8. 1937 [not V. striata Vent., 192]. Verbena Goiounas Park, Illustr. Flower Book 6. 1937. Verbena anafs Hi Hort. ex Moldenke, Suppl. List Invalid Names 7; Isl. Verbena hortensis L. H. Bailey ex Moldenke, Suppl. List In- valid Names 8, ~ 1941. Verbena hortensis Hort. ex Moldenke, Suppl. List eae ie 8, in syn. 191. Verbena hybrida var. alba Hort. ex Moldenke, Sana List Invalid Names 9, in y, in syn. ighi. Verbena hybrida var. caerulea Hort. ex Moldenke, Suppl. List In- valid Wy Names 9, in syn. 191. Verbena a var. colossea Hort. €x Moldenke, Suppl, List Invalid Names 9, - hybrida var. purpurea Hort. ex Moldenke, eer. List Invali Nanies ames 9, in syn. yn. 191. Verbena hybrida var. rosea Hort. @ ke, Suppl. List Invalid Names 9, in syn. 19)1- I. Verbena hhybrida var. syn. 308 PHYTOLOGIA Vol. 9, no. 5 striata Vilm. ex Moldenke, Suppl. List Invalid Names 9, in syn. i9hl. V Verbena teucrioides x venosa Hort. ex Moldenke, Suppl. List Invalid Names Names 10, in syn. i9hl. ve Verbena voglioni Hort. ex Molden- Suppl. List Invalid Names 10, in syn. 191. Verbena hybrida « grandiflora Michell, Grower's Cat. 19h: 23. 19kb. Verbena seas var. hybrida Mill. ex Moldenke, Alph. List Invalid Names Suppl. 1: 23, in syn. 1947. Verbena drummondii Sweet ex Naldenke, gar List Invalid Names Suppl. 1:23, in syn. 197 sat - drummon , 1941, nor Hort., 1839, nor Lindl., 1947, no: oe ) ft eRe nor (Lind1.) ot 1903]. Verbena ee f, chlorina Winge ex Boitanke, Alph. List Invalid Names Suppl. appl. 1 25, in syn. 1947. Verbena hybrida f. nana Beale ex Moldenke, Alph. List Invalid Names Suppl. 1: on in syn. 1947. Verbena 1uminosa Ce Invalid Sa Suppl. 1: 28, in syn. 1947. Verbena hibrida Voss ex Stellfeld, Trib. Farmac. (Velozoa] a (10): 167, sphalm. 1951. Verbena ena hybrida var. nana Hort. ex Moldenke in Chittenden, Roy Hort. Soc. Dict. Gard. : 2211 sentin Verbena hybrida var. coc= cinea Hort. ex Moldenke in Gilt ciekion, “Roy. Hort. Soc. Dict Gard. 4s 2211. 1951. Verbena hybrida var. lutea Hort. ex yoldenke in Chittenden, Roy. Hort. Soc. eg Gard. h: 2211. 1951. Verbena lacea Hort. Soc. Dict. Gard. 4: 2211 year: Verbena hybrida a grandiflora alba Kennedy, Kwality Seeds et a 1951: 45. 1951. Verbena hybri- da grandiflora luminosa Kennedy, Kwality Seeds Spring 1951: U5. 1951. Verbena hybrida ¢ grandiflora violacea stellata Kennedy, Kwality Seeds Spring 1951: 45. 19S1. verbena hybrida gigantea al al- ba Vaughan, Gard. Illustr. 1957: 0. 1957. Verbena teucrioides var. auriculiflora Regel ex Moldenke, Am. Midl. Nat. 59: 349, 48 syn. 1958. Verbena buistii Hort. ex Moldenke, Am, Midl. Nat. 59: 349, in syn. 1958. Verbena hybrida var. gigantea alge ex cunege ke, Am, Midl, Nat. 59: 319 349, in syn. 1958. Verbena hybrid grandiflora elegans Regel ex Moldenke, Am. Midl. Nat. aH 319, 8 syn. 1958, Verbena hybrida var. compacta vesta Hort. ex Se Hort. ex Moldenke, Am. Midl. Nat. 59: 349, in syn. 1958. Verbené = _—* nigro-violacea Haage & Schmidt ex Moldenke, Am. Midl- Nate 59: 349, in syn. 1958. Verbena hybrida var. lutea Haage & Schmidt ex Moldenke, Am. Midl. Nat. 59: 349, in syn. yn. 1958. Verbens Veryens hybrida var, candidissima Haage & Schmidt ex Moldenke, Am. 1963 Moldenke, Monograph of Verbena 309 Nat. 59: 349, in syn. 1958. Verbena hybrida var. coccinea Haage & Schmidt ex Moldenke, Am. Midl. Nat. 59: 349, in syn. 1958. Verbe- na hybrida var. compacta Hort. ex Moldenke, Am. Midl. Nat. 59: 3L9, in syn. 1958. Verbena hendersonii Harrison ex Moldenke, Am. Midl, Nat. 59: 349, in syn. 1958. Verbena melindres var. latifol- ia Bohn ex Moldenke, Am. Midl. Nat. 59: 349, in syn. 1958. Verbe Verbe- ha gigantea alba Hort. ex Mattoon, Pl. Buyers Guide, ed. 6, “287. 1958. Verbena gigantea monstrosa Hort. ex Mattoon, Pl. Buyers Guide, ed. 6, 287. 1958. Verbena gigantea rosea stellata Gulde- 1959. Verbena hybrida var. gigantea Burpee ex Moldenke, Résumé 7, in syn. n syn. 1959. Ve: Verbena hybrida var. variegata Hort. ex Mol- denke, Résumé 367, in syn. 1959. Verbena lindleyi Helo ex Mol- denke, Résumé 368, in syn. 1959. Verbena hybrida compacta Pyn- aert ex Moldenke, Résumé 366, in syn. 1959. Verbena as oak. gantea Hort, ex loldenke, nérans 366, in syn. 1959. Verbena hy- brida var. nigro-violacea Hort. ex Moldenke, Résumé Suppl. 1: 23, in syn, 1959. Verbena hortensis nana compacta Hort. ex Moldenke, Résumé Suppl. 2: 11, in syn. 1960. Verbena cyanea Mazziari ex Moldenke, Résumé Suppl. 3: 37, in syn. 1962. Verbena —— f. Tosa cardinal Wall ex Moldenke, Résumé Suppl. 3: 38 395 yn. 1962. Verbena hybrida var. auriculata Vilm. ex Moldenke, ce Suppl. 3: 39, in syn. 1962. Verbena teucrioides hybrida Vilm. ex Moldenke, Résumé Suppl. 3: ll, in syn. 1962. Verbena hybrida mltiflora Putz, N. Y. Herald Trib., sect. h, p- “p. 16, in textu, February , 1962. Bibliography: oneness Floricult. Cab. 8: 160 & 18h. 1840; lem., Hortic. Univ. 3: 9—11. 1842; Bohn, Florists! rel we ll, fig. 1—3 Syl thcanabs Gard. Chron. 18)6: 18 nay on Rev. Hort. : 5-12. 1852; Florist. yous anaes a . Tel a 5 Viln., Fleurs Plein enema I » & 288. 187; Regel, Garte: ustr, , ustr. Blumeng., ed. 2, 10h5- rel: 285 & 573. 1879; Viln., si Fl. 5s 81. 187918873 os Hulme & Hibbord, Familiar Gard as sos, oS. sane Tllustr. 1: 730. _— pnt OR "-Akad. Handl. 21 a (i): . 3 Lindman, Kongl. S of Fls. 356. 1887; Re- 9& Es oe: Piste or Ingram, repose gel, "te ah es oe ‘Baby i. ners Vilm., Album de Clichés on ee 310 PHYTOLOGIA Vol. 9, no. 5 2565, einen mee ‘ 25661. ee P. Henderson, Types of New Mamno er 1—15. 1 ; Blanc, Mammoth bedi 1888; Re S. Mill., “Tick's “Tlustr. tenth Mage 1) (4): 117. 18913 Vilm, Fleurs Pleine Terre, Suppl. 195. 1894; Hook. f. & Jacks., e 3 Ind, Sak: ora T?. $895; Haage es Schmidt, Cat. Général al 1898: 93 9h. 1898; a Mem. Torrey Bot. Club 9: 169. 1900; Wittmack, Gartenfl. 51: 556--557, fig. 120. 1902; Heinemann, Rev. Hort. Belg. 29 Sh—55 (1903) and 35: ki--hs. DI Vilm. in Vilm. Andr. & Cie, Fleurs Pleine Terre, ed. 5 1909; Russell, Bean, ‘& erz. Samen & Pfl. 95—96. 191 in Barron, Pocket Gard. Lib. 1: (Gard. Fls. Spring] 205 & 230. 1917; Olmsted, Covill Kel- sey, 1. Names, ed. 1, 521. 192); Britton & Wils., Scient. co 6: 138. 1925; Haage & Schmidt, Gen. Cat. 192/ 70. i927; Hegi, Illustr. Fl. Mittel-fur. 5 (3): 220. 19275 Free n eful Pl. Trin. 165. 1928; Guilla d J 2 [Encyel. Prat. Nat. 16]: pl. 43. 1 Thompson & Mor gan, Descript. Seed List 1930: 9—95. 1930; Stapf, Ind. Lond 29-431. 1931; Gard. Chrons, ser. 3, 9: 226 & 231. 1933; Scheff 3 Soc, 59: 301. 193k; L. H. Bailey, Cat. Florists Han mss. 1935; Thompson & Morgan, Descript. — rat ev “DI. = a9 L. He & E. Z. Bailey, Hortus, new rev. ed. & 752. 19395 men, Cytologia 7: 160—163, fig. 1& 2, 16h, ee 167, 173, ’s "YB. 1936; Bedevian,. apie Polyglott. Dict. Pl. 9364 Harrow, Journ Morte Sou” 61: 398--40L. 19363 Noack, Biol. Zentralbl. 57: 305-388. 1937; Emsweller & Blodgett, Proc. Amer Soc. Hort. Sci. 35: 822—82h.. a Goss, Calif. Dept. Agric. 26: 326—333. 1955 rack, Fog ee tr. Flower Book 1937: 48. 19373 Herter, Revist. Sudam. Bot. : 186. 1937; Troncoso, D ana 3 53. 1937; J. v. Watkins, at Pg Fla. Agric. Exp. Stae Bull. 89: 36. RS B fe) Inst. 10: 01, 11, 2h, & 425. 1939; Moldenke, Alph. List 3 O°. Common Names 3—5 t3, 18-20, 23, 25—28, 32, & 33. 1939 a Moldenke, Lilloa {: 333. 1939; Moldenke, Suppl. List Common Nam ; ice, & Scott-Moncrieff, Journ. Genet. bi: 65-— f . : 1941; Moldenke > pistes Verbenac., (ed. 1], 3, 7, 13, 7h, & 101. 1942; Moldenke, AlPDe ,), 19h2; Fl. Te 3.0% 18, 37—38, & ba 19h2; se s cata- bi 3 Schnack, Anal. Inst. Fitotéc. Sta. : 18, 19, & 21. 19h2; Moldenke, Known Geogr. Distrib. Ver 1963 Moldenke, Monograph of Verbena 311 oward, arman, Am. Journ. Bot, Soe toe at 228. 1946; Anon., ard. Gate 1 (1): n.p. 196; Moldenke, Am. Midl. Nat. 35: “ST. 7. 19h; Alph. List I alid Names 28 "T9UT5 Dower, . Sun, May 17. 1947; G. G Ne Y eral Trib., sect. 5, p. lh, December 21 (1947) and sect. . 17, June 27. 1948; Stumpp & 33 op Shr 67h, 678, 715, 716 7h3—7. 736, 837, BL, 89, 852, ats Bop 1950; 3a, sensi 976 ( “Gsis5, and en “abot; 1075, 1076, 108h, 3, 1168, 1179, 1199, 1207, 1208, 1210, 11, 1233, 1253, 125k, 1259, 1296, & 1303. 1919; H. Ne & Ae Le Moldenke, Anal. Inst. Biol. Sect., 30 & 5, March 19. 1950; Moldenke vartateele 3: 67. 1951; stellfeld, Trib. ones taaseoial 19 (10): 167 & 172. 1951; W. A. ee, Burpee Seeds 1952: 6. 1951; Kennedy, kwali s i in coop Roy ruary 3. 1952 r Gard. 1952: 22. “hs eetdenie » 2, 38 227, ia Fe g. 19538 15. 1953; Henderson, Stumpp, & Walter, Garden 1953: 16c & 17- 1953; W. A. Burpee, Burpee Seeds 1953: 18 (1953) and 195i: U8. 19535 Moldenke, “Pagtologss hs 190 190 (2953) and 5: heer . Montevid. 232. 195k; W. A- How to Have Better 312 PHYTOLOGIA Vol. 9, no. 5 —— — ats 1954 D. Jenkins, d Trib., sect. h, P- _ ebruary 21. 195; Biol. foote, a 3501. 195); Moldenke, « Calif, Hort. Soc. 15: 1954; F. C. Chandler, U. 5. Pat- ek + Office Pl. Pat. aoe oa “f "1955 Moldenke in Cheeseman, Fl. Trin. & Tob. 2 (6): 38 32 1955; Moldenke, Fam. 2 Verbenac. 2. p. 12, Mésch = 1956; F. Cx ace er, ee Boyce Thompson a 18: ae ng 1956; J. R. Dalzell, N. Y. Herald Trib., sect. - 18, Jun » 1956; Westcott, Garden. Bug Book 7, pe 72, 7B “ia, 27, “sin, & 507. 1956; measeaasiy in ners Mal T&S 19563 W , Burpee sas 19568 50 (1956)? 19ers 2 & 128 (1957) go apete "ho & 39. 1987) Vaugh- an's Gard. Illustr. 1957: 0 & 98, figs 5738 & 57h9 2357) ye 1958: ho & 98, a Ai & 579. 19573 Camp, Boswell, & Magness, i ur Gard. 82 & [8 1957; er World 3] 5. W. Johnston, N. Y. H raid Trib., sect. h, p. Feb Shinners, Spring Fl Dallas 325. 1958 ae Pl. Buyers Guide, ed. 6 288. Se i. hinhitie Boo . _ 327. 19585 mnagee 3 + Midl. Nat. 5, 3h7, 349—352, 362, ; Vaughan's - Tllustr. 19593 0, fig. 5738, 57h, & og. isis) pie “Bh: 38, fig. $738, 5740, & 57h9. 1999 6 Fl abe A 26; 3 "110, 129, 1h2, 156, 223, %60--%63, 366-369, 373, 376, 378, bl, 436, & W72- 19593 pan ams Résumé SRT 9, 23 (1959) and 2 - 1960; T. H. Everett, New Illustr. Encycl. Gard. 13: 2h0h, pl. 13-10a. 1960; Severy, Nat. Geogr. ait: bios 1960; Anone, Ne Y. Herald Trib., sect. 2, ps 6, ; Vaughan's Garde Titustr, 85: 61, 68, & 70, pl. "STO : stig ‘a962) aot on er’ Tl, pl. 5749. 1962; Moldenke, pésuné Su . 3: 3,8 Pare ciel 36-1 (1962), h: 2& 19 (1962), and Gs 4% "19625 L _ Parks 28 (1): 1. 1962; Park's Flower B Book 1962: 36 & "ST. + 19625 A. Putz, N. Y. Herald Trib., sect. . 18, February . 19625 ‘Leak Bae at Foyt wad 2058, & 2060. 1963 57, & 67, figs 5740 & S7h9- ce - ci 6 vietaaevcnget Ks 29 (1): ¥p Been: packs oon ~s 3: Haggerty, Our Par 1. 1963; Mold denke, Résumé Sup abs ta ht (1963) and 7: 10- 1963; De Me Ferry’ & reitaps Detroit, — _ neo seed pack- ustr. Blumeng., ed. 1, 126). 18 ea — oe seh 1879 pe o ay : so ot pore dt Gart eo ous Hulme & Hibbord, Familiar Gard, Fl. ots tin in Ah tore Me 79-1087; , Kongl. Svensk, Sees camca ana, 21 (ks ple 2» ab 1963 Moldenke, Monograph of Verbena 313 rea Gartenfl., 36: 668. 1887; P. Henderson, Types of bg Man- Verbenas fig. 1—15 [in color]. 1888; Blanc, Mammoth Verbenas. 1868, mana Album de Clichés, fig. 26)5, 25655 ITN & 25661. 1888; seo —- 1 gp Month. Mag. if, (4): 117. 18915, Witthack, + 556, 120. 1902; Heinemann, Rev. Ho Belg. 29: BS ( (1903), = is (1505), and 37: 72 & 36k. 1911; + ON, Pocket Gard. Lib. 1: (Gard. Fls. Spring] 205 [in color]. 3 Guillaumin pare des Jardins 2 [Encycl. Prat. Nat. 16]: pl. iB [in hare 1930 or] & 293 [in colori. “Toa; Bedevian, Illustr. Polyglot. Dict. Pl. 13 [in color], 1h, & 190; Ge . Y. Herald Trib., Jam- ary 31. 1943; Michell's Growers Cat. 19hh: 23. 19h; T. He Fls P) Y. Herald nips & Walter, Seed Annual 1948: 12. 19 ee, e 1950: 47. 19h9; capes & Walter, Seed Annual 1950: 9. 1950; W. A. Burpee, Burpee Seeds 1952: 6. 1951; Moldenke in Casson, New Britton & Br. Illustr. Fl., print. 1, 3: 13h. 1952; J. We J ston, N. Y. Herald Trib., sect. h, D. “nt — ae 19535 Hen- derson, Stumpp, & Walter, Garden Annual or]. 1953; W. A. Burpee, burpée Seeds 1953: 1B "G353),, "gst rf (1953), and io58. 9. 195h5 F. C. Chandler, U. S. Patent 1. 1330—133) [all in color]. 195 Stanek, Beauty mia 25h. 1955; Moldenke in Humbert, Fl. Madag. 174, fig. I, 1& 2+ 1956; = P., Ne Y. Herald Trib., sect. h, p. 12, March 57; Cam amp, ease agni rid in Your Gard. [83] [in color]. 19573 Vaughan's Gard. Illustr. 1957: fig. 5738 [in color] & 5749 [in color] (1987), ache fig. 5738 {in color] & 5749 [in color] (1957), a - 5738, 570, & S7L9 ae in armas 1958; Moldenke in G eason, — Britton & 3: 13h. 1958; vaughan' ant oe Bh: us fig. £738, eto, 6 “Smid {all in color} «2: 9593 Tllustr. Encycl « Gard. rd. 13: 20h, pl. 13 — "fan parson aah or non., N. Y. He Phy sec jae 6, . ou « Geogr {in color]. 85: pl. 570 [in color] & 5749 [in oer es go Tees oe ae in color]. 19623 s Flower Boo 7 [in ane eg — and 1963: 39, fig. 2043 [in color], 2 coe “s 7060. i s ard. Illustr. a7: 57, fig. 5740 [in color] & 57h9 ere 1963; Pelletized Seed Co., s — packet P.4370 [in ee 19635 Steckler — Hee 1 packet 602 ee a: ned.j De Me Ferry i pte in ie) . t, entities moun aan ricer tems from gardenin ee Britton Herbarium;1 mounted illustration fram greeting card in © Moldenke Herbarium, Perennial herb, usually treated as an annual in gardens in -_ Perate climates, vaughe oUe in protected places, of 314 PHYTOLOGIA Vol. 9, no. 5 pact snbe 15—-l;5 cm. tall when in bloom; stems mostly pro f spreading hairs; nodes slightly annulate; principal inter- nodes 1--5.5 cm, long; leaves decussate-opposite, numerous; pet- ioles slender, abbreviated, 2--7 mm. long, densely hirsute with spreading white hairs, margined; leaf-blades chartaceous, u thick-textured, rather uniformly dark-green on both soe or rarely entirely chlorotic, those ay the ra of the branches smaller, base with numerous acute and spreading, often lobe~like and doubly dentate teeth, more or less densely soft-pubescent or hir- sutulous on both surfaces with rather coarse whitish hairs which from ly flat co the floriferous portion .5—6.5 cm. long, 5.5--9 cm. wide, a 55-flowered; peduncles tetragonal, 3-10 or more cm. long, caer ly white-hirsute or -villous like the branches; spikes at first late depressed and corymb- ns er. » r elongating, very ely many-flowered, s closely imbricate and © more or less fr 3 bractlets lanceolate, much shorter than the calyx, 5 ma Y uate at the apex ely 80 riform from white or cream-color to practically all shades of pink, » red, crimson, scarlet, yellow, maroon, light-vermillion, deep-blue, or purpie, some forms described as "between prune aeyt ple and fluorite ag po, "light carmine varying to spectrum red toward the mar , “Bugenia red — lighter areas toward the whi tish l a ned 2 om lo enish or ? rous or ot ng t=] j1-—~2 ly pilos wlous on the outside , the « cm se 9 of oped ppendaged; pollen often 38--78 percent aborted; do be ee p in diameter; chromos some numbers * 1963 Moldenke, Monograph of Verbena 315 exact ancestry of the plant is not certain, but it is supposed to be a multiple hybrid between V. platensis Spreng., V. phlogiflora Cham., V. incisa Hook., and ve. , peruviana Kee) ) ibaa all native to South America. The first-named "parent apparently the source of the white corollas seen in such gi elites) as alba and candidissima mentioned in the extensive synonymy above and the fragrance characteristic of most varieties. The second- named "parent" apparently brings in the rosy and purple shades seen in the varieties caerulea, rosea, nigro-violacea, and purpur ea. The last two, names V. incisa and V. peruviana, are undoubt— edly the source of the red colors in var. coccinea, as well as the general leaf~shape and procumbent habit of the plant. Collectors, Vi he lists the "aurikel-blttigen Verbenen" and "getugte Verbenen" as having sab pre from V. phlogiflora crossed with what he calls V. teucrioides It should be noted here that V. hybrida Bicknell is xV. engel- mannii Moldenke, V. coccinea Waterer is V. peruviana (L.) Britton, V. striata Vent. te V. stricta Vent., V. grandiflora Ort. is Priva grandiflora (Ort.) Moldenke, V. grandiflora Sessé & Moc. is amoena Paxt., and the V. grandiflora of Michaux and of Steudel are V. V. canadensis (L.) . . The recently produced hybrid be- tween - xV. hybrida were Voss and V. tenuisecta Briq. is xV. teasii Mol- denke, which see. There are are at least five names proposed for this cultivated plant which are earlier than the one of Yoss here a- dopted, but xV. hybrida seems to be the first actually dah once tore, of the present tion of the Internat- onal Rules of Botanic Nomenclature is correct, 4 waa rte pted. The dar iier See to have heen pro Species, subspecies, or varieties, not as xVerbena kybriaa lise been collec anthesis from January to May and from July to Dec g at altitudes from sea~ level the latter in Colombia um material has been misidentified and under the following names: VY. aubletia Jacq., Ve. ¢ olia Juss., Ve © “o ws Juss., V. incisa ey Ve marrubioides yon V. peruviana L., V. peruviana (L.) Britton, V. phlogifiors © ts V. phlogiflora Cham. & Schlecht., V- platensis Spreng-, ¥- supina 316 PHYTOLOGIA Vol. 9, noe 5 Le, V. teneroides Gill. & Arn., V. teucrioides Gill. & Hooke, Ve teucroides Gill., and V. venosa Gill. & Hook. The type of the V. hybrida var. variegata mentioned in the synonymy above is Herb. Hort. Bot. Bogor. XV.K.A.XLV.12 in the Buitenzorg Herbarium, while that of V. lindleyi is Herb. Hort. Bot. Calcutt. s.n. in the same herbarium. The type of V. voglioni is Herb. Hort. Bot. Basil. sn. [Aug. 181] in the Meisner Her- barium at the New York Botanical Garden, inscribed "videtur hybd- rida e V. teucrioides et V. venosa?" The type of V. anais is an- other sheet of Herb. Bot. Basil. s.n. [Aug. 181] in the Meisner erbarium, but inscribed "Hybrida inter V. teucricides et Ve (e-B-, labruscana, x0 morifolium, xGladiolus hortulanus, xDelphinium cultorum, xVerbena hybrida, etc.) for use h preserved mate needs to be cl varietal and form designations will serve to cover all the 125 or more horticulturally named trade varieties (cultivars): var+ CO pacta Wittmack, of compact growth; f. nana Beale, of dwarf stat~ ure; var. auriculiflora Vilm., with a conspicuous central "eye" ed; var. alba Thompson & Morgan, with white corollas; var. caerir lea Thompson & Morgan, with blue corollas; var. coccinea Haag? & Schmidt, with scarlet or red corollas; var. lutea Haage & Schmidt, Schmidt, with violet or dark-violet corollas; var. purpurea ThomP” son & Morgan, with purple corollas; and var. rosea Thompson & Morgan, with pink or rose-colored corollas. Horticulturists wishing to use trade names will find the fol- lowing list of value. Descriptions of the plant, as given by the ermany; Harrison, New York city, New & son, L ’ » Davington, Kent, England; Saier, Dimondale, Michigan; oe 1963 Moldenke, Monograph of Verbena 317 Steckler Seed Company, New Orleans, Louisiana; Stumpp & Walter Seeds, New York City, New Tork Sutton & Sons, Reading, England; John Swain, Bristol tea Terrace Gardens, Youngstown, Ohio; Otis S. Twilley, Salis , Maryland; Vaughan's Seed Company, Chicago, Illinois, eg aie York City, New York; Vilmorin-Andrieux & Cie, Paris, France; W Waller—Franklin Seed Company, Guadalupe, California; and Watkins & Simpson, Drury Lane, London, England. Alba (Haage & goimigt) — rein weiss; blanc pur. wide, alnost white, shading to delicate cameo-pink the Atrorielacen ( a (igs & Schmidt) —- rein dunkelviolett; violet eg (art — in many colors, the corollas with a large distinct Neye!, Basilisk —- clear pink, with a white "eye". Beauty of Oxford (Buchman's; Burnett; Hasting's Seeds; Paul J. Howard's California Flowerland; Vaughan's; Waller-Franklin) = , 12 inches tall, flowers 3/4 inch wide, rich -salmon, ’ variable in shade. Sookie of Oxford Hybrids the Be Henderson; ome: 4 ener Stumpp & s tall, flowers over Walter) — 18 inc wide, rose-pink to r allioar "khe brightest rose vaiiiin ev- er produced from seed, a cross between Beauty of Oxford and ae Beethoven -- rich scarlet. el Blue Defiance (W. H. Simpson) — spreading, 1) inches tall, flowers 3/ inch wide, rich violet-blue, variable in shade, with a large creamy-white teye", This is identical with the Franklin, Blue Peter (Demarest) -- rich blue, with a white pte Blue Sentinel (Burpee; Michell) — 10 inches tall, broad ae tke sig OR p vhalet-blve or royal Epepratt ale ripe mea St "Medal Wimner See . Blue Shades Franklin) — ee 1 s tall, ers 1/2 Spares wide, violet-blue, variable in corte Apparently the same as th of er. ge Blue py (Bodger; Macdonald) — spreading, ers 1/2 inch wide, og very variable in shade, with a large creamy—white (Guldemand's Seeds; Bests Kennedy; Toogood & Sons) 318 PHYTOLOGIA Vol. 9, no. 5 — 6 inches tall, flowers deep rich flame-rose, with a med- ium-sized white "eye" Brilliant Giant (iichell) - -- flowers rich deep flame-rose, wth medium-sized white "eye". Apparently the same as "Brilliant". Bush or Multiflora (Putz) —- Ase but with a spread of only 15 c 0 d heightening the e ant ieee ently the same as “Bush Type". Bush Type (Park) — broad flattopped bushy plants, 8--10 inches , 12-15 inches across, the flower-heads almost touching and covering the Bush Type Mixed (Park; Vaughan's) —- with uniform broad~topped bushy habit, 8—-10 inches tall, 12—15 inches across, the flower-heads practically touching each other, the flowers vivid-red, salmon-pink, mid-blue with a white Neye", and clearest white. on purple, light-pink on deep-rose, and pink on white. vanes icc ase odger) — compact, prostrate, to 6 inches tall, the s 7/10 inch wide, soft blush-pink. a ar age Saat & Sohmiat) -- sehr grossblumig, reinstes weiss; b ? s fl ° Cardinal eee Ai oe inches tall, flowers 5/8 inch wide, rich scarlet, with a small cre amy—whi te te "eye"; a var~ lable stock. Carmine Ball (Bodger AR ogi sen Be ae ge Franklin; W & Simpson) — act, 9 inc Sahay toe tower 2/2, inch wide, brat oo a aig pa "variable in shade, ™ a white "ey Carmine Rose (Bodger) — n-compeentiag, the flowers rosy~carmine and cerise, Carter's = owe i oo eapities or dark ——— flowers+ Carter's Dwarf Compact — purple pees Carter's Holborn Mammoth -- pink fl Century (Henderson) -- flowers aaa aie with a clear white canter, Cerise Queen (Waller-Franklin; Burnett; Guidemand's, ner Wat- kins & Simpson) — spreading, 12 inches tall, the flowers 3/k inch Wide, rich rosy-cerise, with a small pet rT "eye"; a stock with color rogues. sisscos © Queen (Kennedy) — long stems, with imense individual over 1 inch wide on giant trusses, soft salmon~ Chamois (Bodger) — compact, 16 inches tall, flowers 2/5 inch wide, not opening ¢ completely, soft be fnareer a true even stock, Coccinea (Waller-Franklin) — spreading, 16 inches tall, the 1963 Moldenke, Monograph of Verbena 319 Hewes =r inch wide, dull scarlet, with a small creamy- white Coccinea (Beager) — reading, e dwarf, 12 inches tall, the flowers 3/l, inch en bright SOE: Coccinea Cage & Schmidt) — rein scharlach; écarlate pur. Coccinea oculata (Haage & Schmidt) -- rein scharlach mit weissem Auge; écarlate pur a oeil blanc. Coerulea (Haage & Schmidt) — rein dunkelblau; bleu foncé pur. Coerulea oculata (Haage & hee cremation — dunkelblau mit weissem uge; bleu foncé & oeil blanc. Colossea (Park) — efaigs over 1 inch wide, in large umbels, in lovely pink shades, also rose with a white "eye". Creamy White (Burpee) — dwarf, compact, 6 inches tall, 10-12 inches wide, forming neat little mounds covered with medium sized flower-heads, the flowers creamy white. Crimson Glow ee Michell; Stumpp & Walter; Vaughan's) -- ro- bus : flowers over an inch wide, forming a 9 Crimson Seedling (Bodger) — spreading, 12 inches tall, the flow- ers 3/4 inch wide, deep ruby-crimson. Crom Prince (Bodger) — compact, 10 inches tall, the flowers 3/8 dnch wide, te Fy Sven ees eee with a small creamy-white eye", a good e ock. crystal (Breck; ‘asain; Herbst; Park; Saier; Terrace Gardens; Vaughan's) — compact habit, 9 inches tall, spreading only 10--12 inches, with 10—15 flower-stems per plant, th flowers pale glistening snowy-white in immens maculate Cupreata (Haage & Schmidt) — kupferscharlach; écarlate cuivré. Danson peered — flowers rich purple-mauve, with a clear white Dannebrog Sapo Daehnfeldt; Wa & Simpson) — less com- pact, 12 inches tall, the ror 1/2 inch wide, rosy: crimson, with a creamy-white "eye", the stock ale in Dark Blue (Bodger; Hurst) — spreading, 16 inches tall, the flow ers 1/2 con side, poms variable in oer Apparent- ly the same as the Blue Shades of Waller- aa solid lasses )— yer compact habit, t-red, with no "eye", "so ees you "need sung to 1 look at itu yee Defianc iance] (Hurs han's; Waller-Franklin spread— py pmpnarae e shies’ da: 7 the flowers 4/2 = wide, rich scarlet, te teye". This is the Défiance of Haage & ome. Re-selected (Watkins & Simpson) — apparently the same as the Defiance of Waller-Franklin. 320 PHYTOLOGIA Vol. 9, no. 5 ——_ i ea & Schmidt) ~- feurig scharlach. This is the same he Defiance of Waller-Franklin. peiNaeas purpurea (Haage & Schmidt) — purpurroth. Delight ineoe Vaughan's) — see Dwarf Delight. oe er Kronprinz (Haage & Schmidt) — listed as a form of Ver- ma hybrida compacta erecta. Dwarf re 3; Vaughan's) —- see Sparkle. Dwarf Senne (Vaughan's) -~ 6-7 inches tall, the flowers clear oral-pink, suffused with salmon. a, Master Blend (Vaughan's) -- very compact and dwarf, the flowers appearing when the plants are very small, retail taining their color and habit throughout the season, with a wide range of color, many with "eyes", Dwarf Mixed (McAllister) -- spreading to form circular bushes 4 bout 2) inches in diameter, compact, the stems stiff and 6 rect, the flowers large and varicolored. Dwarf White Compact (McAllister) — short sturdy habit, very » the stems about 6 inches long, flowers white. Edith _(Hentereon) — flowers salmon, shaded carmine, with 4 white mfin Scarlet oreary aa ee pa Ester Waller—Franklin) — com pact, the red, in shade, with color rogues « Elite (Ohlsens cay © ints “is inches "tall, the flowers 5/8 inch wide, mixed colors, Ellen ee (Bodger; Hurst & Son; John Swain; Thompson ” Mor~- an; Watkins & Simpson) — spr 12 inches tall, the ers HGS inch wide, rich rosy salmon-pink, with a small "eye", a true stock, but Een nf El Mahdi all le ea royal~purple Emily (Henderson) — flowers royal-purple, with a clear larg® center, white Buily Dale (Demarest) — flowers beautiful pink. Etna (Michell; Waller—Franklin) — spreading, 12 inches tall, the flowers 3/4 inch wide, rich Soneiet or intense geranium~ red, with a small creamy-white " ¢ Etna (Kelway; Macdonald; Watkins & Pau inferior stosks © the above. Btna (Stumpp & Walter) — flowers vivid scarlet, with a small yel low "eye", Fairy Queen (Bodger) -- compact, a mixed stock. “ Famy (Henderson) — flowers violet-rose, with a large white "67° Fine Mixed (Ferry) — various in color, shades, and striping, 16" ally running pure white to deep-purple. Finest Mixed (Burpee) — a mixture of Apple Blossom, co nie cane ’ trum Red, and i ips Glory, Royale, 1963 Moldenke, Monograph of Verbena 321 Fireball ee Watkins & Simpson) — compact, 10 inches t tall, the flowers 3/l inch wide, bright rosy-red, a good e stock, Fireball (Burpee; Stumpp & Walter) -- 6 inches tall, covered with brilliant scarlet or flame-scarlet flowers. Firefly -- bright scarlet. Firelight (Henderson; Park; Stumpp & Walter; Vaughan's) — 10 in- ches tall, broadly flat-topp hy, the flowers a most vivid rich scarlet, intensified by the absence of an "eye", Flamingo (Stumpp & Walter) — erect, 11/2 feet tall, the flowers vermillion— a Floradale Beauty (Burnett; Burpee; zones) S gem Fro; Saier) — robust spreading g rowth, the flowers an inch wide, oa shades of Beauty of Oxford os, in extra large tates rose from bright rose~pamke to deepest rose-red, ; All- America saikeer Medal tandet Galatea (Henderson) -- yiakers bright scarlet, evenly striped with white, th salmon- Giant (Burpee) — 12 inches tall, with a wide range of colors. Giant Erect Mixed (W. H. Simpson) -- compact, 18 inches tall, the flowers 1/2 inch wien; in mixed colors. Giant Salmon Pink (Vaughan's; Waller-Franklin) — spreading, 1h inches tall, with flowers 5/8 inch wide, ing rosy-salm with a mali c reamy-white "eye", a good eve stock. Gigantea Alba (Guldemand's 8 Seedes Garden Trails pete; Husbands Rockcliffe Gardens; La Germinadora; Vaughan's) -- robust ha~ bit, 12 inches tall, with flowers 1 inch wide, pure white, very fragrant. Gigantea Annapolis Blue -- see "Annapolis®. Gigantea Beauty of Oxford — see "Beauty of Oxford". Gigantea Brilliant -- see "Brilliant". Gigantea Ellen Willmott —- see "Ellen Willmott". Gigantea Giant Salmon Pink — see "Giant Salmon Pink". Gigantea Lavender Glory -- see "Lavender Glory". Gigantea Lucifer —- see "Lucifer". Gigantea Rose Queen -- see "Rose Queen". Gigantea Royale -- see "Royale" Gigantea Scarlet Queen -- see Te Pe Queen". Gigantea Snow Queen — see "Snow Queen". Gigantea Vivid — see "Vivid". Glow—worm (Henderson) — flowers brilliant scarlet, of perfect form Golden n Queen Bodger; Kelway; Park; er) — spreading, 15 hes Siar) ~- spreading, 18 sachet i the flowers 5/8 inch wide, scarlet, with a large c white "eye", a true s ° Scarlet King (Michell) — the flowers bright-—red. Scarlet Queen (Hurst; Park; Thompson & Morgan; Unwin; Watkins & Simpson) —- the flowers huge, vivid-scarlet, with a white "eye *, Scharlachrosa (Wittmack) -—- auffallend schtne, niedrige Verben® Verbena von aufrechten Wachstum; die zahlreichen, an straff aufee- richteten Stielen siteenden Blumen sind von leuchtend scharlachrose Farbe mit weissen Auge™. Schlosser's Pride (Michell) — the flowers dark-red. Sedan (Demerest) -- the flowers blood-red. Silver Plume (Danerest) -- the flowers finest white. Simoon aoe) -- the flowers brilliant crimson, with 4 dark "eyet Snowball (B Bodger) -~ compact, 15 inches tall, the flowers 5/8 ull white. inch wide, dull Snowdrift (Breck) -- flowers purest white. Snow Queen (Harrison) -- spreading, 18 inches tall, the flowers 3/ inch wide, — with color rogues. The same as ora White o £’ Bodger Snow Queen (Hurst; eat John Saudis Thompson & Morgan; Watkins & pA on son) — giant—flowered, the flowers a 2 sch Snow riod — see "Snow-white". Snow-white (Burnett; Henderson; Park; Stumpp & Walter; ¥ Yaughan's = 10 inches tall, Lasadly flat~topped, bushy, 12--15 inches Sparkle (Bracks renee Herbst; sags —* Saier; Terrace inches mghan's) -- dwarf, of uniform tall, the flowers brilliant cachet or eight soeny red, with a ‘Large gleaming white "eye", 1963 Moldenke, Monograph of Verbena 327 Sparkle (Putz) — only 10 inches tall, so age covered with flowers that only an occasional leaf shows t ea to softer salmon, lavender and white, mid-blue and ae many with bright-hmed combinations. This is probably t Sparke of Putz. Special (Burpee) — a mixture of Apple Blossom, Royale, Spectrum Red, and White. Spectrum Red (Breck; Burnett; George Delbard; Guldemand's Seeds; Herbst; Norsk F105 Park; John Swain; Vaughan's; Waller Franklin reading, we brilliant scarlet or intense rich scarlet, with no Spectrum Bs Red (Watkins & Simpson) -—- a less regular stock of the Spectrum Red (Henderson; Stumpp & Walter) — 6 inches tall, the owers a dazzli pectrum Bet 3 Anatkg Vaughan's) — dwarf compact habit, the flowers royal-purple, accented by a clear snowy—white "eye", starlight & (endereeny Mercia Park; Stumpp & Walter; Vaughan's) O inche » broadly flat-topped, bushy, 12—15 Fan Sn g across, pha oe at intense blue or bright mid-blue, with a creamy-white "eye". Steckler D — flowers white. Steckler EB — flowers pink, with a white " wrist (Park) — flowers of red and rose eae charmingly riped. Striata (Haage & Schmidt) — italienische gestreifte in allen Farben gemischt; striés en mélange. ~— Suttonts — Burpee; Herbst; Norsk Fro; Park; Sutton the se royal —blue ee solid mid-blue, in rounded trusses 2 1/2 to 3 inches long and wide. Union ee ) — flowers with an enormous white eye encirc- ed by a rim of violet~blue. Yes. (Burpee) — dwarf, erect, 10 inches tall, the flowers soft chamois=-pink, the jnnermos ones suffused with copper. Violacea Stellata (Bodger; K ay spreading, the oe vi- olet or deep—violet, with a white Sot, a mixed st aS Violet Bouquet (Waller-Franklin) — 12 inches 4d flowers sf inch wide, bright fem at with a creany- white "eye" Vivid (Garden pxtiounes 3 Vaughan's) —— brilliant sunproof ned, po a bright Pree color, in large uniform trusses s sore Angatenes car= 328 PHYTOLOGIA Vol. 9, no. 5 Weisze Zwerg-verbene — see "Dwarf White Compact". White (Burpee) ~- flowers white, White Ball (Michell) -- petite, 7 inches tall, with 20 or more f erect branches, each with 3—~5 clusters of 2-inch white flowers, White Beauty (Henderson) — 6 inches tall, the largest and finest te variety ever introduced, the flowers pure-white, blooming continuously. White King (Stumpp & Walter) — flowers pure sparkling white. Zwerg Verbenen — see "Dwarf Mixed" In addition to the above, the following horticultural names have also been found in the trade: All-color Giant, American Gardens), Spies Cerise Beauty of Oxford, Choise Mixed, Coccinea Dwarfer, Comp ure, Danneborg [error for Dannebrog], Dwarf Coerulea, Der t ‘ tea Monstrosa (Toogood & Sons; Watkins & Simpson), Gigantea an Glory (John Swain), Gigantea Rosea Stellata (Guldemand's Seeds), Gigantea Sutton's Pink (Guidemand's Seeds: Sutton), Lag mere ie olacea Stellata (Guldemand's Seeds), Holborn ab ear bore onsets (Norsk Fro), Hortensis Hybri ra (Saier), Hybride ot carlet Shades train, ru moth Rose Queen, Mammoth Scarlet Queen, Mammoth Snow Queen, “wi ilyn (Burnett; Terrace Gardens; Vaughan's), Master Blend mee man's; Terrace Gardens), Miss Millmott [error for Miss Willnovt), Delight, . Mantle, Red, Red Emperor, Rose, Rose~red, Royal Blues, scarlet, Scarlet Beauty of Oxford, Scharlach, Schlosser's Cerise-rose, Schlosser's Orchid, Shylock, Snowstorm, Spectrum, Sultans, SY ton's Giant Pink, Sutton's Giant Royal Blue, Sutton's Giant White, Tom Thumb, Triumphator, White-eye Blue, White Giant (Has ting's Seeds; Herbst), and Zeuxes. Unfortunately, I as yet 18° any descriptions of these. names for XV. da are also numerous, including "alfombrilla", "aurikelblumige Verbenen" » “barbfng", "camaradin- ha", "common garden verbena" » "common verbena", "corbeille preci "es "Eisenkraut", "garden verbena", ngerden- artenverbenen* verbena", ” ltssiga verbene", ngermanderblttttiga verben4; “garten Eisenkraut", "gttvercin otu", "gttverein otu®, bicodil 1963 Moldenke, Monograph of Verbena 329 bena", "lipia", "malvira", "margarita", "shah's favourite", "sha- passan", verbeina", me bs rbéna", "yerbena comune", "ver= bena de cae, tyverbena ne tyerveinel, tani ape anais", "verveine commune", *verveines des jardins", "verveines hybrides? and "verveine hydri de a fleur d'auricule". Bedevian (1936) gives two Armenian common names for plant in Arm Sscripte Bailey (1935) lists the following ete ed varieties, with the Bote or seedsmen the seeds of each at that time: V. grandiflora (ichell; Rose Valley, Lyon, New York; Schling); ve hybrida (Hunt; Muller-Seely; Stecklers; bomen enemas North Abingtm, meer Oscar H. Wills Company, Bismark, North Dakota); V. hybrida alba (Thompson & Morgan); V. hybrida caerulea (Haage & z Schmidt; siineon & Morgan); Ve hybrida candi- Gissima (Haage & Schmidt); V. hybrida coccinea (Haage & Schmidt); V. hybr: hybrida compacta (Haage & & Schmidt) ; ve terse grandiflora (Haage & Schmidt; Huntington; Francis C. Stokes, Philadelphia, Pennsylvania); V. hybrida lutea (Haage & Schmidt); V. hybrida nigro-violacea (Haage & Schmidt); V. hybrida purpurea (Thompson & Morgan); V, hybrida purpurmantei (Haage & Schmidt); V. hybrida rosea (Thompson & Morgan); V. hybrida striata (Haage & - Schmidt ); V. luminosa (Chester J. Hunt; Waterer); V. rosea stellata (Hun- tington; Waterer; Kunderd Nursery, Goshen, Indiana); V. violacea stellata (Kunderd; Waterer). In regard to Lemaire's V. teucrioides var. anais, the comments of the oh tepid are of interest: "Parmi les nombreuses variétés ou s jardins. Mais, en cette occasion, nous nous som mes poets au sentiment de M. Paxton, ae décrivant de son c6Oté une belle espéce de Verbena nous et tout a l'avantage de la derniére.".sees l'historique et la description du type de la variété nate es S'agit, Celle-ci am diffare par ses feuilles manifestenen 330 PHYTOLOGIA Vol. 9, no. 5 s (et qui nous feraient croire que nota a presidé 4 sa ss 9 nm c'est-a~dire qu'elle serait née . teucrioides et dtune espece congénére fort voisine), matin Ciel et méme on plante, dont la floraison dure plus de deux mois, et qui, plantée en terreau de bruyéres pur, prospere dans un endroit chaud et péré Rtmpler (1873) discussed this plant as follows: "Einjuhrig, im Gewtichshause ausdauernd. -- Es ist heutigen Tages sehr schwer, mit Sicherheit zu ermitteln, welchen botanischen Arten die un men. Man wird jedoch mit der Vermuthung nicht fehlgehen, da als Stammeltern die Verbena chamaedrifolia (V. melissoides und ihre rothe Variet#t Verbena melindres Gillies, ue tani und incisa, die unter Nr. . 5 und 6 beschriebenen Arten, und @ lich Verber Verbena phlogifera Chamisso, welche am Ende eat "plosse Formen einer und derselben Stammart sein kOmten, anzusehen haben. "Die botanischen Merkmale der Gartenverbenen sind dieselben, wie wir sie fur Verbena teucrioides angegeben haben. "Die Gartenverbene ist so sehr vertnderlicher Natur, dasz man bei jeder Aussaat von Samen, die man von Varieté#ten sortenweise Bestand der Collectionen sehr wanderbar, z f- Variet#ten, wenn ihre blumistischen Werthe in neueren ther eo fen sind, die meisten aufgegeben und nur einige bres 8 aa rragen elb und des Schwarz sind bei der Gartenverhene alle Farben ates und durchlaufen vom reinsten Weisz und Indogoblau bis zum leuchtendsten Roth die ganze Far~ benscala; man hat eee: blitwliche, lilafarbige, amaranth- rothe, purpurrothe, carmcisinrothe, barune, schieferfarbige ¥+5- w. in den verschiedens tad eee schtnen oder auffallen- zigem as die Farbe betrifft, $0 ist rade Iechatey neue oder a uffallende Colorit werth erhalten zu werden iat man denjenigen Blumen den Vorzug, bei denen a . des Kronensaumes von der des Schlundes > verschieden is um welchen herum die walbena des Saumes enen und Klar abgesetzten Zirkel bilden musz.” 1963 Moldenke, Monograph of Verbena 331 On page 126 he lists var. auriculiflora Hortorum ("Aurikel- scharfer Lini farbe ws airmail so — — verhYltnismaszig groszen Blumen denen ainer Aurikel in Etwas “hnlich sind; diese Race ist sehr beldebt. mad pihieuhcaad On page 1265 he lists var. eae a Dy ee a ers ausgezeichnete Varietttten. Bei einigen anderen sind die Blumen halb von diefer, halb von einer anderen Farbe und es ist gar Richt selten, auf einge elnen StUcken und selbst in den Blttthen- bouquets eines und desselben Stockes in vollkommender Weise ge- streifte, zweifarbige und einfarbige Blumen zu finden, letztere ad in der b arbe. t zuverlussig ist, vorzugsweise durch Stecklinge vernehrt werden, Seit einigen Jahren hat man jedoch auf dem Wege einer recht umsichtigen Auslese eine fact erzielt, sage aus Samen ganz befriedigende Resultate giebt. Die italienis en Verbenen wi Shivers diese nat der Blume h di ichtigkeit ihre P auszerordeniliche Dau- erhaftigkeit, durch den Reichtum ihres Flors, mit dessen Schon- , ch dadurch, dasz sie wegen der Weise ihrer Anordnung zur Be- rae, von Bouquets, zur Ausstattung von Vasen u.s.w. geeignet "Diese Pflanzen geseihen in jeder sonnigen und freien Lage und in allen lockeren, leichten und gesunden Bod prametine? Einige Frische und Dtmger sind ihnen vortheilhaft. Bodendeckung und Be- gieszen im Sommer werden immer von den besten relia ad eee tothe’ e Tep- ereitet aus den Verbenen Einfassungen, Gruppen und plctbeete von hoher Schtnheit, deren Eifekt sich durch —. schung oder durch contrastirende bandftrmige Pflanzung vi ai fach vertidern luszt. Wegen der groszen Menge von heckenaegnt Garten egg eye, ame oa Wenden, ohne Einfv: 4t befttrchten gu mtissen. Diese Pflanz breiten sixh rasch ther den Boden aus, in dem sie sich einwurzeln und den sie mit ihren ausgebreiteten Zweigen (einige jedoch haben frech ende Z und b auf en, welche von Juli bis in den Sp&therbst umun 332 PHYTOLOGIA Vol. 9, no. 5 einander folgen. Auch in TUpfen kultivirt sind sie reizende ten, = u.seW. leisten sie gute. Dienste mmehrt die Yerbenen durch Aussaat mad durch Stecklinge. Die Aus sagt welche den Vorzug verdient, weil man dadurch kr#f- de oder Lauberde; die noch ganz jungen Pfltmzchen pikirt man in die kleinsten Tupfchen und thherwintert sie dicht unter dem Glas eines Lauwarmhauses; man entspitzt sie, wenn es ntthig sein sollte, e s Keimen bisweilen seht eigensinnig; die am sorgf%ltigsten gepflegten Samen versagen mitunter ganz und gar, wtthrend die in das freie Land ge s ch und Farben » Wie sehr auch die Samentr&ger durch Farbenpracht i _auszeichnetenj immer aber wird die Aussicht, schtne Sam zu erzielen, grtszer sein, wenn man die Samen von Pflanzen erste® Ranges und von Varietitten mit lebhaften und dunklen Blumen sai~ melt. "Die schtnen Varietuten, die man aus Samen ee hah, > OOF servirt man, idem man die durch Stecklinge verm und das see fast in jeder Zeit des Jahres peeenenens de pS aber bewirkt man diefe Vermehrung im Frtthjahr 1 in Mistbeete o- im Gew8c nach jeder Seite hin, je nach dem Zwecke der 1963 Moldenke, Monograph of Verbena 333 ZUNE « Nese G&rtner machen ihre Stecklinge vorzugsweise im No- vember in December in Schalen im Warmbeete aus noch krautig woichen ag tees von Pflanzen, die man in kalten Kasten eae r angetrieben hat. Jede Schale besetzt man mit eine groszen Menge solcher Stecklinge und pflanzt diese zu Ende jee Andere ziehen es vor, Ste cklinge in Februar-Murz im Warmbeete zu erziehen aus ee Trieben, die von im Gewtchshause tierwinter- ten Sttcken in "Seit ‘algo s sake cai hat man sich mit Erfolg bemttht, gewisse Variettten soweit besttndig zu machen, dasz sie sich, wenn auch noch nicht durchweg, doch zum grtszten Theile aus Samen farben- treu wiedererzeugen." It is worth noting here that the Verbena hybrida names pub- lished in Rev. Hort. Belg. 35: 1—-l5 5 (1909) me vane Hh--365 (1911) are in a series of advertisements pu by Haage & Schmid are to be eas to raphe 5 pebehaier of the firm, not to Pynaert. Lemaire's V. colsieee var. Divy its variant designations) have berry, been regarded aye os pli gered with V. platensis Spreng., but upon "pke say of the original description and plate, as well as what may be au thentic material of the plant — wath Saris Basil. s.n. [Aug. 1841] in the Meisner Herbarium, said on the label to be a — between V. teucrioides and V. tweediana — I feel now that Britton & Wilson (1925) refer to our plant as V. ch iia Juss., while Calderén & Standley (191) refer to it as ve teucrioides Gill. & Hook. L. H. Bailey describes all his col- lections fran enemy California, Alabama, and North cosy eee as "perennial" or "wintered over". He notes on P. L. Hall 6 that —_ pret eateng are definitely not appendaged. a ti9h3) ™ dr ns ornhi (1880) Soh eqyie ova verbena wal mae Formerly O22 seaseied st coon to Serious drawback to its culture, many Continue to be raised, and those who S aaia know the ieee As surprised and ¢ armed ee as the show varieties are to be A The culture ge & Verbena is so simple that it an easily undertaken by anyone possessed of an ordinary frame R. S. tee (2891) i nyerbenas may be slipped seid eng for winter blooming. e really elegant window plants." 334 PHYTOLOOG IA Vol. 9, no. 5 in g in poor soil, and withstand drought remarkably well." Peter Hen- derson says "Sow in shallow boxes of light soil in the greenhouse, hot-bed or light window in a temperature of from 60 to 70 deg +, usually flower well in August from seed sown in open ground May. Germination will be hastened if the seed is soaked in luke- warm water before planting. It is desirable to have the soil well fertilized and carefully pulverized. Cover the seed about one~ the yo to cover the ground completely. Earlier blooming may be had if e ts are started indoors and transplanted in the open ground after danger of killing frost is over." Burpee advises "Sow the the sp 1 s s na win r transplanting into the garden when the maples are in full leaf; or, the seeds may sown outdoors dur- ing April and May, either in a special seed-bed or right where the ts are to bloom. Keep the seed-bed moist, because the hi s 12 inches apart in a row. Soaking the seed in water 12 hours before sowing usually helps to quicken germination. Verbenas Vaughan (1959) says "These colorful annuals originated as des rt plants, thrive e sun, flowering continuously. They & n beds, bouque sow the seed outdoors until the soil gets warm (May 15th to June 1st). May also be i rs d e. Gr inches tall but free~flowering, and with a considerable range of color making it excellent for sumer b j some forms are pleasantly poented. The vari strains offered in the trade are doubtless of hydra origin, the result of crossing V. peruviana, V. incisa, V+ phlog- iflora, and V. platensis in various combinations. Trea 1963 Moldenke, Monograph of Verbena 335 other half-hardy annuals, the seedlings should be planted out a- ne the end of May about 1 foot apart. By this time they will be eginni grown as pot plants with the necessary stopping and supports to counteract their spreading habit. Verbenas appear to be worthy of wi ’ ment. They are very desirable for massing in beds on the lam. e large clusters of showy flowers are borne in succession through a long season until frost. The seeds sell in the United States at 15--50 cents per packet, 70 cents to $1 per one-quarter ounce, -50 to $3.50 per ounce, and rooted cuttings in 2 1/l inch pots for $2.50 to $6 per 100, or $20 to $55 per 1000. ink family union. Under the name of V. teucrioides Gill. & Hook. the longevity of this plant's seeds is reported on by Barton (1939). She used the variety "Royal Bouquet Mixed" from W. Atlee Burpee Seed Com- er pigments of Verbena [hybrida] have been established: (a) ee anthocyanidins may be pelargonidin, delphinidin, or mixtures these with each other or with cyanidin (i.e., all combinations ex~ cept cyanidin alone). (b) The anthocyanins may be 3:5-dimonosides. (c) The anth may or may not be ee beets (d) The non—acylated types have a more alkaline cell-sap, then the pigment is more densely aggregated. (e) The ne 1‘ may be partially or completely inhibited; if the forme, ~— an increase in anthoxanthin copignentation. (f) The anthoxan nine hocyanin) may be (in flowers not containing ant ) luteolin (in te? low or an ermined flavone (in white flowers) « : inheritance of these variations is contrary Plants in the fo : (a) Pelargonidin oot ay are Sometimes dominant, sometimes recessive to delphinidin. page Sides are sometimes , sometimes ive anne 336 PHYTOLOGIA Vol. 9, nO» 5 c) Mixtures of anthocyanins occur, due to incomplete dominance or to modifying factors. These irregularities are considered to be due to the interspecific origin of the garden Verbena. The other variations are not abnormal as regards dominance, direction of mutation, or gene-interaction, in so far as they can be com pared with homologous differences in other lants ." Beale (190) says "There is little doubt that crossing ha been attempted by horticulturists. Lecocq (1852) reports having crossed V. incisa, V. tteucrioides' [=V. platensis] and V. 'Mel~ indres' [=V. peruviana], givan an account of a technique for e- masculation and pollination much the same as that used in this work. Some of the variations of the modern garden strains appear to have arisen as mutations after the original crosses were made e.ge: (1) Pure white flowers. (2) Yellow flowers (said to have this hybrid as follows: V. peruviana -- V® pros= trate habit, flat cotymbose inflorescence, bright scarlet flow ers, with a very yellow "eye" with a dark inner rin, troduced in about 1826 from Argentina; V. phlogiflora ~~ upright habit, the spikes longer than those of V. peruviana, the flowers scarlet-magenta, with a medium-sized yellow "eye", introduced 2 isa xV. hybrida: "Compact" is dist he norm ing shorter internodes and no runners; "Long spike" has the — florescences as long as 20 cm the corolla-tubes longer than oro in the normal; "Stellate" has the petal-lobes bent downward om © radial axis, while the whole plant is reduced in size, bears few » and sets little seed; "Wrinkled" has both the gre - wrinkled, ; e soeerns "Chlorina" or "Bleached dwarf" is a very small partially orotic plant, the cotyledons and first leaves greene PHYTOLOGIA is financed entirely by its contributors, each onc paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. Each number consists of not less than 32 pages. All manuscript accepted will be published in the next at so that the size of numbers may Vary greatly. A volume will contain ut 32 = 512 pages, or a smaller number o' with an pagel ae number of plates. This plan insures immediate publication of all accepted mice Ilustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced ” tad propo of on their size, as fixed by the engraver, with a minimum oF | z meee so-called popular type. and polemics will not be sages Advice ie “Under send present coat 0 of ‘Printing, the tone Fae rate ee : ae — fracts ct PHYTOLOGIA Designed to expedite botanical publication Vol. 9 January, 1964 No. 6 CONTENTS STEYERMARK, J. A., and collaborators, Notes on Ecuador plants... 337 MOLDENKE, H. N., Notes on new and noteworthy plants. XXXIX.... 350 MOLDENKE, H. N. , Materials toward a monograph of the genus Verbena NIV. 5525 occa Ge ee en nee 351 MOLDENKE, A. L., Book reviews... c-+--0c0-scucue-trteto® ve++- 407 —______.__ Published by Harold N. Moldenke and Alma L. Moldenke tee 15 Gioia pee ae ‘Yonkers ae teoeg bi s a ‘Price + of this umber, $i; ae volume $ NOTES ON ECUADOR PLANTS Julian A. Steyerma Instituto Botanico, Ministerio de iiecsan y Cria, acas, Venezuela and collaborators Between March and December of 193 the author was in Ecuador, engaged in the collection of Cinchona na and other members of the Rubiaceae. In addition, other “plants » were collected as time af- forded. Some of these have been found to constitute new taxa, and have already been published by various botanists. Although a large umber uadorian pose Linen of the ohieage Gattiral History Museum. CYPERACEAE CAREX AZUAYAE Steyermark, sp. nov. Planta stolonifera, culmis trigonis ere vel basi 1-2- foliatis 20-30 cm. altis 2 mm. diam., surculis sterilibus phyllo- podiis, laminis erectis 15-5 cm. longis 3-5 m . lathe non ’ septato-nodulosis marginibus serrulatis; BE perce OE aang ores es parte clausa ca, 2 mm floris tenuipedunculatis; spicis terminalibus asculis ca. longis, lateralibus plerumque andr anes ari-cylindricis h-5.2 cm. longis 3-3-5 mm. diam. cum terminali longa; glumis foemineis atropurpureo-castanei obtusis perigyniis brevioribus 2 prea carinntis,. © carina lido-viridescente Biv brumea 8 rigynii cis _ e rg eee ET {naequaliter bi- wc ae bicostatis dor=- 0 5 -0.2 ] “a e SCUADOR: es Azuay: p4ramos in vicinity of T Molieturo and Quinoas, alt. 3785-3900 m., dune 15, Steyermark § 53105 (holotype, F). ~~ ‘This we. well-marked species is characterized by the pedunculate, long slender spikes, obtuse pistillate scales, ve as Toughened, Parsonees Frese eo erveless or obscure Toreador, be 1943, Julian eas 338 PHYTOLOGIA Vol. 9, no. 6 rved perigynia, reticulate surface of achenes, elongate leaves, stolontferous, stout, elongated culms, and the lowest bract con~ spicuously exceeding the inflorescence. This is a member of the Subgenus Eucarex, Sect. Acutae, Sub- section Vulgares, apparently related to C. Andersonii ii Boott and C. decidua Boott of Chile and Argentina on the one hand, and to Cc. strictior Dewey and related species on the other ean It re- sembles the latter in the granular-roughened peri t dif- fers from it and related members of the group to to which "at paees in the phyllopodic culms and long-stipitate perigynia. From bs Andersonii Boott, C. decidua Boott, and C. Hindsii Clarke, and others, it differs in its very granular-roughened ghened perigymte eg spikes, and stouter culms and leaves X TOREADORA Steyermark, Planta stolonifera, culmis 4g ety cm. longis 1.5-2 mm. pats ee angulis striatis, infra spicas minute sspaceiepsty foliis oC ms lo 898 2-3 mm. latis culmis plerumque breviori scabridulis, supra canaliculatis, cits are es na spice foliosis; inflorescentiis {anceclato-ovatis 1.5-2 cme longis glumis infinis brevissime bra is; spi -5 ice ovatis densifloris 6-8 mm. 1 s 2. mm. sated spicae aAaee bractea a -5 mm. a aristata; glumis fo pall long emine ferrugineis vel rufis late suborbiculari-ovatis apice rotundatis vel obtusis usque breviter mucronato-acutis perigynii bre ° This taxon is characterized by the conspicuously rugose upper leaf Surface, densely brown-tomentose lower leaf surface, the short racemes, and the small flowers. APOCYNACEAE Robert E. Woodson, Jr. MANDEVILLA VERSICOLOR Woodson, sp. nov Frutex volubilis, ramulis élongatis “crassiusculis pilosulis 348 PHYTOLOGIA Vol. 9, no. 6 cortice rubro-brunneis. Folia opposita petiolata, lamina ellip- tiso-ovata acuminata basi cordata 10-15 cm. longa iy em. lata membranacea utrinque pilosula supra nervo medio basi pauci- glanduligera, petiolo 1-1.5 cm. longo, appendicibus interpetio- om. longo basi ca, 0.2 cm. diam. dilute viridi et rubromaculato, faucibus conicis ca. 1.5 cm. longis ostio ca. 0.6 cm. diam. basi Staminigeris, lobis oblique obovatis ca. 3 cm. longis sordide croceis margine interiori salmo-rubro patulis. Anthera anguste oblonga basi truncata 0.8-0.9 om. longa g . Ovarii carpella chlongatdes oa Pas cm. longa glabra, nesterias 5 plus min' cm. longis, stigmate wmbraculiformi ca. 0.l “m- eked longe Popes a |ADOR: Prov. El Oro: between Paccha and aac ‘cag ie _ rb a de Pueblo Viejo, Cordillera ; slash e 30 me, A e 26, a ulian Ae ‘aioe (holotype, F). pa. 2 Obviously allied to the western South American species of Subgen. Beeverite, Sect. Laxas, particularly Me Me ery but differing in the unusually large flowers and lax, pend inflorescences each at first sight pain’ the genus feanwe— Pemnaden SOLANACEAE yebrer sce il gsc Steyermark, nov. s dense fulvo-villosulis; foliis petiolatis, iar i rare om. longis dense velutinis, laminis simplici ovatis apice subito subacuminatis basi subcordatis vel late ro- tundatis 6-7.5 cm. longis 3) cm. latis supra plus minusve dense dense velu tu 15-17 mm. longis dense pilosulis; floribus secundis fere Scorpioideis; calycis tubo breviter cupulato-campanulato 1 m. alto 4 mm. lato dense eae = 5 anguste triangulari- canpanslat tis 11-12 mm. iongis, tubo rishi mm. longo basi 3 mm. lato fauce 8 mm, lato, lobis eS ovatis subacutis 7.5 mm. lon~ gis 5-6.5 mm, latis extus villosulis intus glabris laxe reticu- ‘venosis; bus exsertis, peas oer oblongis apice ro~ 196) Steyermark, Ecuador plants 349 tundatis 5-6 mm. longis 2.5 mm. latis glabris; filamentis 3 m. longis glabris basi corollae insertis; stylo 10-10.5 mm. longo Pilosulo; ovario glabro ovoideo-conico; bacca globosa 1.5 cm. longa 1.5 cm. lata glabra. ECUADOR: Prov. Loja: region — Las Chinchas, alt. 2250 ms, Abril 12, 1944; arbustiva; flores moradas; b ’ redo ndas, Pheed M. Acosta-Solis 773 Asin: F). Local name: "sabal- ueur This species differs from C. hypomalaca Bitter of Ecuador in the shorter petioles, smaller lnef-tledes, £ fewer-flowered inflor- escence: «3 shorter pedicels, longer, nar calyx-lobes with attenuate tips, and differently shaped aaeotes fruit. NDIFLORA Steyermark, sp. no mh ligneosa, ramulis apic BR peels stulis ceterum glabris; ro floribus pat pedunculatis; calyce tenes caipanatake tau re- ticulato-venoso basi l-carinato 5.5-6.5 cm. longo basi 1.7-2 lato a cm. lato, lobis ovatis subacutis vel obtusis 2.2- cm. longis 1.5 cm. latis; corolla * o longio: a; antheris lineari-oblongis 16 m. 1 mm. la- tis; filamentis glabris; ovario conico 9 mm : Prov. Pic ha: Bone de Loche a Condor Mackay, Cordillera Occidental, alt. 2700-3100 m., Enero 15, 1944; arborea ° arbustiva; flores grandes sag peciolos largos colgantes; cor= olla verdosa; Me Acosta-Solis 7009 (holotype, F). The large calyx and corolla are characters of this species. NIEREMBERGIA ESPINOSAE Steyermark, gs pt calyce tubuloso ee 6 mm. Tree ee ee at ; . is subacutis vel Shetiagcbita. teri-infun- extus minute ia tam 5-7 mm, po 6 mm atin subito dilatato extus pilis glan- bus remote obtecto apice in lobos breves late triangulares 2.5 mm. latos 0.5 mm. longos leviter desinente; ovulis ca. 20; Seminibus ferrugineis subrhomboideis obtuse sub extreni- oe subtruncatis vel obliquis 1 mm. longis; testa alveolato- osa, 350 PHYTOLOGIA Vol. 9, no» 6 UADOR: Prov. Loja: en prados, Namanda, al 130 a Nov. as , 19465 yerba menuda, rostrera; * poco ee adas, color blanco con ligero tinte rosado; Reinaldo a HE A o Espinosa 107% Rn, F). ~ Phis species is characterized by the diminutive size of the plant and proportionately large size of the corolla. NOTES ON NEW AND NOTEWORTHY PLANTS. 0Odx Harold N. Moidenke FRIOCAULON BREVIPEDUNCULATUM var. LONGIPES Moldenke, var. n0V- Haec varietas a forma typica speciei pedunculis 3.55 cm. longis recedit. This Bhs farts differs from the typical form of the species in having its peduncles 3.5--5 cm. long, usually subequaling the leaves. The type o - the variety was collected i R. D. Hoogland and Re Schodde (no. 7647) in a swampy patch in treefern grassland, t an altitude ~ of about 8500 feet, in the sie: saneseng are’ (source of Lagaip He. Subdistrict, Western Highlands New Guinea, on August 31, 1960, and is deposited ei gin United States Herbarium at Washington. LIPPIA LASIOCALYCINA var. SAINTHILAIREI Moldenke, var. noVe Haec varietas a forma typica speciei bracteis "late ovatis ad apicem acutis vel subacutis recedit. variety differs from the typical form of sy species in having its bracts broadly ovate and merely acute or subacute * The type of the variety was sei mabe tap by August Frangois esar Prouvengal de Saint~Hilaire - whose honor it is nam named em at Olho d'agoa, Minas Gerais, med between 1816 and 182h, s ted MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XIV Harold N. Moldenke leaves practically white, and it does not flower; "Strap" is a narrow rib Plant are tonpenheas; 2n = 20, while her Rose Red, purple, Magenta Verbena Plants are triploids, an = 15. Salmon (1900) states that the fungus Microsphaera ferrugines Erikss, infests this plant, while in Biol. Abstr. 28s 3507 (195k) it is given as the host for a species of Erysiphe. ¥ Westcott (1950) reports that it is attacked by Flower Blight (Botrytis cinerea) odera marioni) in Maryland, by Stem Charcoal Rot (Macro ee ma, by three kinds of Root Rot -- Phymatotrichum canivorum in is basicola er Yellow Wooliybear phair a ae oa Morningglory Leafcutter (Loxostege obliteralis), Verbena Leafminer (agromyza artemis ; Oblique-banded Leafroller (archips rosaceana), Cyclamen Mite (Steneotarsonemus pallidus), Broad Mite Mite (Hemitarsonemus latus latus), Two-spotted Mite (Tetranychus telarius), Ferm Nematode (Aphelen- Choides olesistus), Greenhouse “Orthezia (Orthezia insignis), Flower Thrips (Frankliniella tritici), Greenhouse Thrips (Helio~ thrips haemorrhoidalis), eer — 50 Whitefly (Trialeurodes vapor Srey, ret id ere i), oes us persicae), Melon isha (A » moth (Endothenia Saresieney. Tees hed Plant-bug (Lygus lineolar is), Foxglove Aphid (Myzus solanii), Cottony cushion Seale (108 Ya purchasi), Gray Blister-bee beetle, and Snapdragon Lace-bug. Lundell rts that it is nplanted for or ornament throughout the seed of Texas. The Keeling 64 64, cited below, is labeled as e from cultivation. The Collector undes ignetes 2 22 from Mexico, the Carr s.n. en eas 352 PHYTOLOGIA Vol. 9, no. 6 s.n. [Peking Union Medical College] from China, and the L. H. Bailey 7155 and 7156 from Mississippi do not have any indication on their labels that they were taken from cultivated material, but I am assuming that they were. Neither the Herb. Est. Cent. Agron.s.n. (Nov. 1909] nor the Fortune & Arias . 7066, “ited below, ear any indication on m their labels that they came from cultivated material, tn Dr. Acufla assures me that they did. below, is a mixture with V. canadensis fo ) Britton. Perry says "Probably cultivated — cf. - V. peruviana (Le) Britt." for the Kerber 1,70 collection cited hereinafter, but there is no indica~ tion on the label that it came from cultivated material. Similar ly, there is no evidence that the G. W. Barclay 528 cited below was cultivated — its Lana reads "herbaceous creeper, flowers rose coloured, soil - sandy loam - found in open habitations in the woods of Machala, Colombia, Oct. 1836." Norton found xv. hybrida growing in low sandy bottomlands and open woods, while Radford found it in waste places in South Car=- olina. Stellfeld ( 1951) cites Stellfeld s.n, [Ilha do Mel, 5-ll- 94; Herb, Mus. Paran. 2075]. Schaffner (1933 & 1934) cites L. Be Hicks s.n. as escaped in Astabula County, Ohio. Herbarium material of xV Ve rida is often found in herbaria under the name Y. incisa Hook. On the other hand, the Dole 33, distributed as xV. hybrida, is ceoresed Gaillardia pulchella a var. picta A. Gray in the Carduacea Standley (1938) applied ay name V, teucricides Gill. & Hook. to the commonly cultivated garden verbenas of Central America which are actually xv. hybrida. i says "Verbena. Cultivated lant. Native of southern wit from of Central America, eaenetaliy in the temperate regions. The pal aed specific name of the garden plants, which may be in par ae Ages weer is somewhat uncertain." xamined by m wees —. PENNSYLVANIA: Beaver Coe: Shafer s.n. [Beaver Falls, Sept. 6, 1900] bh SOUTH CAROLINA: Darlington Co.: J- Be Se Sonton sen. [Apr. 8, 1921] (Hi~59)62). Kershaw Co.: A. E- Radford 20772 (Hi—92953). FLORIDA: Brevard Co.: J. K. Small S7U8 La atta OWA: Boone Co.: Pammel & Pammel — ey 192k] (Io—116119). OKLAHOMA: Payne Co.: K (St—17567). TEXAS: Val Verde Co.: Parks ay 9003 Clute 900k (Tr). CALIFORNIA: Ventura Co.: He M. Pollard Son. {San Antoni 8 Creek, June 25, 1948] (Gg—343936). MEXICO: Nuevo Leén: noyeal z 1964, Moldenke, Monograph of Verbena 353 (W—192)914). Vera Cruz: Kerber },70 we ec te PANAMA: Panam4: Paul 39 (W—-1586935). CUBA: Havana: Van Hermann 867, in part (W— 115108). COLOMBIA: Antioquia: Toro Toro 935 (Fo —1663, N). BCUA~ DOR: El Oro: G. W. Barclay 528 (Bm). BRAZIL: Federal District: Fraz&o s.n. (Herb. Jard. Bot. Rio Jan. 702] (N). Minas Gerais: Sampaio 21 (Ja—l6750). Rio de Janeiro: Rocha e Silva 54 (Ja). Sio Paulo: Santoro s.n. [Campo Experimental; “Herb. Inst. Agron. S. Paulo 7613] (Be—35758). SWEDEN: Nilsson sen. (Sept. 1907] (Go)5 Wide s.n, [Malmb, Okt. 1897] (Go). CONGO LEOPOLDVILLE: Bredo 1153 (Br). TANGANYIKA: A. Peter 37891 [V.168] (B)~ MADAGASCAR: R: Baron X19 (K, P)s Herb, Jard. Bot. Tananarive 1322 (P). CULTIVATED: Alabama: L L. H. Bai. ley 7 ey 7048 (B (Ba). Amoy ). Amoy Island: H. H. Chung 1609 (Ca—225555), Argentina: Burkart 18105 (N); Herb. Univ. Mass. 5. n. (Aug. 186] (Ms); T. Meyer 12620 (N); Moldenke & Moldenke 19723 (N). Arizona: A. R. Moldenke 620 (S). y. Austria: “Khek 5.1 SNe [aug. '72] (vV—11289). Barbados: Herb. Bot. Stat. Barbados 322 (N). Bermuda: Brown & Britton 1728 (N);Brom, , Britton, & Bisset 2209 (N). Brazil: Etzel s.n. [Prefeitura Municipal, July — 1937] (N, Sp—38719); W. Hoehne 2557 (Bh, N); Moldenke & Moldenke = Ds Stellfeld 1095 [Herb. Mus. Paran. 1095] (N). “califor- ailey & Bailey 7960 (Ba), 9743 (Ba); o: Fe P. Bradbury 5-n- Pye Hollywood, 1916] (Ba); W. R. Dudley sen. [July 1893] (Du— 362568) ; Eastwood s.n. [Inverness, May 13, ia] (Gg—2142h0); H. Me Hall 3819 (Ca—59951); P. L. Hall 63 (Ba); Lenington s-n- [Po- mona, Dec. 15, 1927] (HP), Sen [Pomona, Feb. 16, 1938] (Hp); Woodcock 979 (Ba). Canada: Desrochers 16-52 (ug), ba 417-52 (Mg), Li8-52 (ug), 19-52 (Mg), 991-52 (Mg), 592-52 (Me), 25-52 (ME), 736-52 (Mg); Herb. Marie-Victorin s.n. (Vi). China: Chiao Chiao 259 [Herb. Univ. Nanking 12028] (B2—23750, Ca—294567)5 "Herd. Inst. Bot. Acad. se 1459 [275] (S); Lauphit 113 (Ur); Lin sone san. [Pe edical College] (Ba). J). Colombia : Daniel 2: 0S (F— 78707) ; ‘4 Garcfa y Barriga 11619 (N, y—185Li15)5 To = 1220 (Fn—1657). Congo Leopoldvillet RR. PP. Salésiens 291 (Br). Suioitinc saa Enequist 152 (N Herb. Est. Cent. ee 1909] (Bs); Ponce & Ramos L15 Delaware: Herb. Sulasgant sen. (Pa). EL Salvador rte (Vi). Florida: ( ), 7022 (Ba Ba), 8-Re see, ey 3] (a, Ba); G. B. Grant 2503a [10678] (Po); i. S. Watkins s.n. [ e, 3-20-31) (Fl—21113)- snag Bouchon s.n. Um—29); Herb. W. He F s.n, [b. n (Daffour 5029] ( ) bs TE ie — On Ss R. P. 1837) (Du—166485); Herb. Saldanha 2769 ( 354 PHYTOLOGIA Vol. 9, now 6 L. Krause 29280 (B). Georgia: Cuthbert s.n, [Augusta, June 1, 1900] (Fl—2111)). Germany: Bothe s.n. [Juni 1903] (B, B)3 Golens s.n. (Schwiebus, August 1872; Herb. Prager 18627] (Gg--31450), 8 n. [Schwiebus, 1872] (B), son. [Schwiebus, Sept. '73] (B), SMe {August] (Gg-—31L9), s.n. [Schwiebus] (La); Lohmeyer 8.n- {cham aedrifolia var. hybrida] (B, B, Du), son. [teneroides] (Du); C- J. Mayer s.n. [Mtmchen, Septbr. 1895) (Mi); Schulz & Schulz 5-M- [Berlin, 13.9.1896] (B), son. (Berlin, 1.6.1897] (B). Hawaiian Islands: Neal s.n. [June 25, 19h2) (Bi). Illinois: Herb. Div. Floric. Univ. Ill. 176 (Ur), 273 (Ur), 886 (Ur); Herb. Univ. iu. s.n. [Oct. 1, 1870) (Ur); Stenstram (Ur, Ur). India: Herb. Horte Bot. Calcutt. s.n. (Bz—23717, Bz-~23787). Iowa: Harmon SMe [Marshalltom, Oct. 7, 1929] (Io—-133933). Italy: Gresino 5.Ne [29.V.1938] (N); Herb. Harvey son. [hort, Bellorae, jun. 1042 (Du-—-166458). Japan: Collector undesignated s.n. [17 Juni 1908] (W—113306k). Java: C. A. Backer 12523 (Bz—23706); Herb. Horte Bot. Bogor. XVK.A.XLIV.5 (Bu—-26425), XVK«A-XLIV.9 (Bu-~26L26), XVKA.XLV3 (Bz--23755), XVK.A.XLV.12 (Bz—2376u), XVKA-XLV-L3 (Bz—264h0, Bz), XVKA.XLV.20 (Bz--264)5), XVK.A-XLVI.19 (B2—- 26470). Kansas: E. A. Popenoe s.n. [College Greenhouse, ApTe -» 1901] (W—-111,010). Louisiana: Langlois s.n. [August 1878] (I)- Madeira: E. Wall 5 [27/329] (Bw). Massachusetts: L. H. Bailey 5: n, (Cambridge, June 21-23, 1919] (Ba), s.n. [Cambridge, August 6-12, 1929] (Ba, Ba); Carr s.n, [Clinton] (Se—1h9li2); A- Clare son, (Amherst, July 1876] (Ms)} He L. Jones s.n. [Oct. 1693] (Ob—50810); Kidder san, [7 Aug. 1929] (Oa--10740)3 Torrey & Cross s.n. (Oct. 3, 1936] (Ms, Ms). Mexico: Collector undesi ted 1 (I), 59 (I); M. B. Foster s.n. [Veracruz] (Ba); Halbinger & Reko 312 (N); Moldenke & Moldenke 1982h, (N). Mississippi: L- E> (Nm); H. N. Moldenke 3030 (N), 7858 (N). New Mexico: Arséne & Benedict 21573 (W--1696551). New York: L. H. Batley s.n- [Sept- 1, 1915] (Ba), s.n. (July k, 19193 Vilmorin-Andrieux 80070] (Ba, BA), 13, 1924; seed sown May 6; Ferry] (Ba), s.n. [Sept. 1, 192k] (Be photo, Z—-photo), s.n. (Sept. 12, 192k] (Ba), Si eT pe ee : Sept. 20, 1925: : 6 =~ s.n. [July - 53 Haage & Schmidt 5203; seeds — April 6] (Ba/s Steckler E] D) (Ba), son. (Aug. 31, 1927; Carter 5805; seed som April 1] (Ba), sen. [Nov. 1, 1927; Carter 5807; seed sown May 17] (BA)» 5+ 196k Moldenke, Monograph of Verbena 355 n. [July 29, 1928] (Ba, Ba--photo, Ba--photo, N--photo, N--photo, Z=-photo, 2—photo), s.n. [Aug. 5, 1928] (Ba), s.n. (Sept. 2 1928] yeh 3; Burdick s.n. ene (July 2, 1923] (Ba); Collector undesig- hated s.n. g eae, S Rug. 1, 196] (N)3 Herb. N. Y. Bot, card. Eethe sane [hugiat , 1896] (N); Jacobi s.n. [N. Y. Bot. Gard. Cult. Plants 26575; es/i0] (N), san. [N. Y~ Bot. Gard. Cult, Plants 26575; 7/8/10] (N), sn. [N. Y. Bot. Gard. Cult. Plants 26575; 8/ 1/10; seed from Leiden] (N); H. N. Moldenke 8242 (N), 8243 (N), 82h (N), 1189) (st), 11895 (St), 11897 (St); Moldenke & Moldenke 11887 (N), 12868 (N), 11893 (N), 11694 (N), 11895 (N), 11896 (N), 11897 (N), 12898 (1), 11699 (N), 11900 (N), 11901 (N); N. XY. Bot. Cult. Plants 111 11100 (N—photo) ; W. He "He Rhoades s.n. sen. [near R - Rochester, July 1938] (Hs); Re C. Schneider s.n. (N.Y. Bot. Gard. Cult. Plants 26575] (N)3 A. Wood s.n. (Pa). North Carolina: L. H. Bai~ ley s.n. (Washington, May 17, 1930] (Ba)3 P. 0. Schallert “T7805 (0k, Ok), sen. [8/10/32] (Or——-28)33)5 Me We . Ward Son. rsen. [oct. 21, 1938] (Hi—5083); Ziegler s.n. [Oct. 16, om) (Hi--069), 3.0. [Oct. 18, 1938] (Hi—-06h)~ North Dakota: St. Lawrence L9 (I). Ohio: E. ’ Wilicinson sen. [25 Sept. 1887] (E8055) Oklahoma: 0. He —— + 220 (St—2l366); Murphy 99 (Ob—35109). Oregon: Ge ie a estas - 8, 1933] (Or—-h:2525) . Panama: Maurice 896 ee Pennsylvania: Burpee & Co. S-NM- [Doylestown, stom, Fordhook Famous] (Ba), s.n. [Doylestowm, “Scarlet Defiance] (Ba). Philip- pine Islands: M, S- Clemens 18h31 (Ca—368058). Pitcairn Island: pe & Christian 11199 (Bi). Puerto on Britton & Boynton 825k (N); Moldenke & Moldenke 19537 (N)- : Br. —- = aie ton 237 (N). St. Vincent: Morton 5457 + (y--186,366). Sweden Bagge 8 son. = (aie (Go); Blom s.n. (Aug. 1931] (Go); Herd. n. [Holmiae, 180] (S); Lindstrom 5 Sah (En); Eaters Sen. [31/7/1886] (Go), sen, [13/9 ] (Go); Tro- lander 5 (Bw); E Z. Wall 5 [18/84] (Bw, agi 2 [1/941] (Be, Bw), aes 941] (Ew). Switzerland: Herb. Hort. t. Basil. son. [Aug. 18L1} (u), son. (Aug. 1839] (M), Sete eras I) GO, s-n. QM). Texas: L. H [. Bailey 7398 (Ba); Ce L. Lundell 10936 (N, Rf), & Lundell 10939 (N, RE), > 10940 (Sm); Lundell & 9763 ie N)5 Reeves 190 (N). Uruguay: Herb. Herter 6171y (N); Rosengurtt Be 3k28 (N). Venezuela: Bailey & “galley 88 (Ba), ise eyes (oe); Elias 158 (0a—13986). Vermont: Dole 3u6 (Vt); Loveland Loveland $.ne 3-22 2-92] (Vt cality of Llection undetermine LS egenen a dine Fe: Harb, waseiart 1685 (S zoe (s), 356 PHYTOLOGIA Vol. 9, no. 6 » 3. Exquisite (N); P , Mammoth erbenas, copy coe 1888 (N); embossed sae plate (N); “Yerbena Extra Choice Mixed 2 pts: (N); 25645. Verbena hybrida (N); Verbena. Packet, 20 Cents (N); No. 88. Verbena. a 0O (N); 25660. Verbena hybrida, var. erie (N); 25661. Verbena hybrida, var. striata (N); 25662. Verbena hybrida, var. sivtake (N); Verbena. Garden Hybrids (N); Hybrid Garden Verbenas (N); Tom Thumb Verbena (N); 25665. Verbena hybrida, var. auriculaefiora (N); Verbena (N); Verbenas. be est of our General seopientl is (N); P. Henderson, New Verbena "Union" (N)5 No. 3097. N erbenas,. $1.2 25 (N); McAllister, Dwarf neat cereee Verbena (N); leciatagees: New Mammoth Verbena (N)3 A- of Mammoth Verbenas (N); L. H. Bailey, Ithaca, Sept. ? “92h “Ws Le He » July 29, 1928 (N, N); New York Bot. Gard. (N » 956, Verbena randifiora Brightness (N); 4521, Hende American Beauty Collection of N pores (N, Ds 2 are Verbena All-color Giants (N, N, N)3 > did N)5 Verbena (1); 12, Verve a i a (N); 79906-79907 Verveine hybride variée extra (N)3 Store, Vaughan's Best Verbenas erbena trum Red (N); 2761. Blue eae Verbena (N); Verbena ae "Beauty of Oxford" (N); New Giant-Flowered Verbenas Gs "Flamingo" (N); Attraction - A New Hardy Verbena (N) 5 New York Bot. Gard. color slide 6073 (Z). XVERBENA ILLICITA Moldenke, Revist. Sudam. Bot. l: 18--19. 1937- Synonymy: Verbena urticaefolio-stricta Engelm., Am. Journ. Sei. 6: 101. 1844, Verbena stricto-urticaefolia Engelm., Am. Journ. Sci. 46: 101. 18h). Verbena stricta x % urtiesetolss Mac- kenzie ex Moldenke, Revist. Sudam. Bot. 4: 19, in 1937 [not ¥. stricta x x urticaefolia Eggert, 1958, nor Ap ee 1958, nor Pammel, 1958]. Verbena stricta x urticaefolia Pond ex Mol- denke, Revist. Sudam. Bot. 4: 19, in syn. 1937. Verbena strich™ stricta x urticifolia Britton ex Moldenke, Revist. Sudam. Bot. pot. bh: 19, 12 syn. 1937 [not V. stricta x urticifolia Glatfelter, 1958, nor is 1, in syn. 1937. sav urticifolia x stricta Eggert ex Mol- » Revist. Sudam, Bot. 4: 19, in syn. 1937. Verbena strict Ryans Septende Gates, a Kens. 191. 1940. Verbena hastata gn ticifolia Gates, Fl. Kans. 190, 190[not V. hastata x a x urisolfole Blanchard, 1937, nor Dermen, 1936, nor Eggert +, 1937, nor 19k7, nor Perry, 19), nor Pringle, sghTl Verbena stricta x ue ticifolia Sehneck ex Moldenke , Suppl. Invalid Names ames 10, in 196, Moldenke, Monograph of Verbena 357 syn. 1941. Verbena stricta x urticaefolia Gates ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 1947. Verbena stricta x urticifolia Bush ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 19k7. ——- urticifolia x stricta Gates (in part) ex Moldenke, Alph. List Invalid Names Suppl. 1: 27, in syn. 197. Verbena urtic Seatin ; éiststa Norton ex Moldenke, Am. Midl. Nat. 59: 352, in syn. 1958. Verbena stricto-urticifolia Patterson ex Moldenke, Résum6& 375, in syn. 1959. Verbena stricta urticifolia Gates ex Moldenke, Résum6 375, in syn. 1959 gn ae a Engelm., Am. Journe Sci. 46: 101. 18443 Molden- 03 Mol us K denke, Prelim. Alph. List Invalid Names 48 & 9. 19L0; Moldenke, Suppl. List Invalid Names 10. 191; cones Alph. List Invalid Names 50 t 51. 1942; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 6—11 & 101. 1942; Deam, Kriebel, Yuncker, & Salisb. Moldenk + Cit. a a a 399, 470 516, & (198), 3: 699, 721, 773, 7 , 792, 800, 822, 826 sey 90 (919), and ibs 1217 » 1255, 12h, f "1298. aes eee 7, cee Scr 19135 ikeas: piptaiadhe 4s ni l49193 and : 67. 1952; Y tniile in Gleason, New Britton & Br. Illustr. Fl., print. 1, 3: 126 aie) 3 on, New tton & Br. Bod: ° 3 130. 1958; Moldenke, am. Midl. Nat. 59: 347 & 362—353. 1958; Moldenke, Résumé 16, 17, 19, 21, aay 26, 365, 375, 377, & 72. nquis ons: Moldenke in » New Brit tton & Br. Illustr. Fl., print. 1, » 3: 130 (2952). sa print. 21 3 130. 1958. This is a een Ve Vent. and V. ur- ticifolia L., "eal é eas ge eo rica of the two species overlap in the central United States. It is based on the 7 V. urticaefolio-stricta of Engelmann, of which the type was col- ected by George Engelmann at Saint Louis, Missouri, in September of 1841. It resembles V. urticifolia in habit, but has the spikes denser, not greatly elongated after anthesis, very a and ly fruited, the mature calyxes Bm ostly contiguous, the flowers lareer, the leaves broader, and the stems, branches, in- florescences, and leaves densely and coarsely long-pubescent . The de: as blue Poindexter 1962) compares this hybrid with its two parents as follows: Leaf ra oe Ve urti teifolia = 1.9--2.6 (average 2.2), xV¥- 358 PHYTOLOGIA Vol. 9, no. 6 illicita = 2.1--2.7 (average 2.2), V. stricta = 1.3--2.1 (average 1.8); le leaf-shape: V. urticifolia = broadly adly lanceolate to ovate, xV. illicita = ovate to broadly lanceolate, V. stricta = elliptic to ovate; leaf-base: V. urticifolia = rounded, decurrent into the ear xVe illicita = = sharply attenuate to sessile, V. stricta attenuate to sessile; leaf-apex: ve urticifolia = grad- any acuminate, xV. illicita ita = gradually to 0 abruptly acuminate, Vv. stricta = suave to . rounded; stem pubescence: V. urticifolia = hirtellous, xV. illicita = hirtellous to hirsute, » V. stricta = hirsute; nutlet length: V. urticifolia = 1.6—1.9 (average 1.7), xV. illicita = 1.6—2.) (average 2.2), V. stricta = 2.2--2.9 (av erage 2.6); markings on back of nutlets: V. urticifolia = moder ately ribbed, xV. illicita = moderately to 0 densely ribbed, V. stricta = artecle: ribbed; pollen fertility: V. urticifolia = 83— 99 percent (average hte? percent), xV. illicita = 19-6], percent (average 34.7 percent), V. stricta = 50-100 percent (average 90 percent); corolla-tube length: V. urticifolia = 1.5--2.5 (average 2e1), XV. illicita = 3.l-—-l.5 (average i), V. stricta = hk. 55 65 (average .9); calyx length: V. urticifolia = 1 1.72.3 (average 2), xV. illicita = 2--.5 (average 3.2), V. stricta = 3.8--5.1 (average 4.3); petiole-lateral vein measurement: V. V. urticifolia = Uy--25 (average 19.6), xV. illicita = 315 (average 9-7), Vs stricta = 3—6 (average 3. .. aes ~~ Storm reports the hybrid as "not numerous", growing among Ys stricta Vent., V. hastata L., and V. urticifolia L. Pammel refers to it as "a common weed" (but probably is referring here to the true V. urticifolia with which he has mixed it on herbarium sheets) in sandy clay soil, associated with V. stricta, V. hast® ta, Anthemis cotula L. = Persicaria mitis Gilib., Ambrosia elatior® @ elatior L., and Coreopsis palmata Nutt. Other ‘er collectors have found it a it in wet or sandy soil, pastures, meadows, waste grater and river~ ron September. Ahles found it in a pasture with V. stricta, Ve hastata, and V. urticifolia, ae anes: : Lge panne it in Stricta Vent., Vv. stricto-paniculata ce = ark mm. wide; stamens istil included. ype of this little-lmown species was Bisse cted by Hassler (0. 3164) on the campo near Tobaty, Paraguay, and is de- posited in the Delessert Herbarium at the Conservatoire et Jardin otaniques at Sage Briquet notes that "Cette curieuse espéce le Snse, et ses bractées ellip cleus, subobtuses au sommet (lancé~ olées-acuminées dans le V. Scuatheneda). Thus far the sutiee Ts known only from the type specimen, not as yet seen by m VERBENA INCISA Hook, in Curtis, Bot. Mag. 65: pl. 3628. 1639 [not V. incisa Chod., 190k]. Synonymy: ? *Yerbena arranana J. Harrison, Floricult. Cab. 6: 7. 1838. ?Verbena tweedieana var. arraniana Niven ex Maund & Henslow, Botanist 2: pl. é0, in textu. 1830, Verbena triumphans Marnock, 2 3 Verbena arrani ana Hort, 4: "Bote, i iy 18h1. verbena arran= Seana vr gg ex ab si, in Mieks. rs Bras. 9: pega Se Verbena triumphans triumphans Hort. ex Stapf, Ind. Lond. 6: 431. 1931. Yer bena tweediana arr arraniana Niven ex Stapf, Ind. Lond. 6: 431. 1931. Verbena fulgens Bonpl. ex Moldenke, Lilloa 6: 331, in syn. 19h. Verbena incisa incisa Hort. ex Moldenke, Alph. List Invalid Names Suppl. Z iven ex + 255 in syn. syn. 1947. Verbena tweediana var. arraniana N Ucidenke, Alph, List Invalid Names Suppl. 1: 27, in syn. 1947. Verbena arraiana Hort. ex Moldenke in Chittenden, Roy. Hort. Soc. 362 PHYTOLOGIA Vol. 9, no. 6 Dict. Gard. : 2210, sphalm. 1951. Verbena briquetiana ana Osten ex Moldenke, Résumé 360, in syn. 1959. Verbena na briquetiana f. cam- briquetiana f, silvatica Osten ex Moldenke, Résumé 360, in syn. 1959. Verbena chamaedryfolia a melindres f. siccanea Osten 6x Moldenke, Résumé 362, in syn. 1959. Verbena chamaedryfolia hybrida Osten ex *Moldenke, Rhouné 362, in syn ~ 19596 Nezpaos nopapotanice var. truncatula Brig. ex Moldenke, ghee 369, Ver- bena melindroides f. briquetiana Osten ex ada Résumé 369, in syn. 1959. Verbena melindroides f. silvatica Osten ex Moidenke, Résumé 370, in syn. 1959. Yerbens megapotamica f. Lakh Briq. ex Moldenke, Résumé Supp Suppl. 3: 40, in syn. 1962. Verb Verbena ediana var. grandiflora ae ex Moldenke, Résumé Suppl. 3: syn. 1962 Bibliography: J. Harrison, Floricult. Cab. 6: 47. 1838; gi arnoc « DC., Prodr ten ¥ 1851; Vilm., Fleurs Pleine Terre, ed. 1, 939. 1865; Rttmpler a Viln., eras? Peeeue's ed. oF 1263. 1873; Regel, Gartenfl. <0! - eee 19023 Ed. isis ea sihectaait: gh 1902: 11h. 1902; Chod» Plant. Hassler. 9: 197. 1902; Briq., Ann. Conserv. & Jard. Bote Genév. 7-8: 288. 190k; Briq. in Chod. & Hassler, Bull. Herb» Boiss., sér. 2, hk: 1061—1062. 190k; Brigq., — Bot. Stockh. (10): 1012. 190k; L. H . Bailey, Cycl. Amer. Hort., ed» 4, 1... 1912, pl. 2649. 1906; Gerth van Wage, Dict. Plantnames 1397--1399- 1911; L. H. Bailey, Stand. Cycl. Hort. 6: 34h5, pl. 3910. 19175 C. L. Pollard in Webster's New Internat. Dict. Eng. Lang. 2275+ | 1917; Herter, Florul. Urug. 105. 1930; pit, “tnd. Lond. 6: 1430 431. 1931; Parodi, Rev. Argent. Agron. 1: 202. 193k; Le H. & B- % cea ove sihget | new rev. ahi; 6326 19355 ensures as i Inst. ty veld’ Nene 19405 ae O, 348, ¥ & 365. 19) 1940; woldenke, Liliza Seay eee! seh} ond “a, 10, 38», 2 ie eogr. Distrib. Verbenac., [ed. 1], ear Th, & for. "T9h25 Moldenke, Alph. List invalid Names 196), Moldenke, Monograph of Verbena 363 1942; Schnack, Anal. Inst. Fitotéc. Sta. Catalina h: 19423 Moldenke, Lilloa 10: 385. 19hh; penne eg ogia Ot "19 & 115. 1945; August 0, Fl. Rio Grande do Sul 209 & 232. 19116 Moldenke, Alph, List Cit. 1: 12, 22, 47, 212, & 262. 19183 Moldenke, Alph. List Invalid Names Suppl. "1: 22 & 25. 1947; corte Aiea on 2: 348 (1947) and 2: h25. 19h8; mele —_ a 13: ao Moldenke, Alph. List Cit. 2 : 356, 360, 37h, 318, ‘a » Wa, 481, "533, 535 SP, ae 598, & 599 “(ua 3 703, 731, 132 748 749, 780, 1832+ 860, 875, 893, 909, 223 66 (1949), 1080, '1088, , hae 7292, 1a > 6h 1172, 1207, 77, 1233, 1237, 1283, & 1293 191594 Maldenke oes *Geogr, Distrib. rbenac. , 16k, & 198. 1949; Are [ed. 2], 9h, 98—~ Piytologit 3: 135° (sis), 3: zal (1980), and 3: usr. 1951; Stell- 9 (10): enke in Chitt: a feld, - 1951; Mold Roy. a yom ae a: Me? 208, 2209, & 2211. 1951; on in roe my ae 19565 Moldenke , Inform. Mold. S 51 S 3 Moldenke, rs Midl. Nat. ‘so sto & & %2- 0) olen, Résumé He 135, +105 ae eas 23, 367, 3695 421, & h72. 19595 ‘angels, Fl. aran. 16: 78 (1960) an 447 73 vue ase ‘sinus Résumé Suppl. 2 & & 36—la (1962), liz 19 (1962), 5: 2 (1962), and 6: 10& tions: J. Harrison, Floricult. Cab. 6: me {in wes ae hee 1838; Marnock, Floricult.Mag. 2: pl. 2h, fig. 1 [in color]. 1838 Maand & & Henson, Botanist 2: pl. 60 [in color]. 1838; “Hook. in it icle at the tips of the branches ope cle~like, later prongs tirectiets orere, 1/k to 1/3 as ong as the calyx, densely whitish-pubescen owers . about O mm. ae dense t .e-pubescent or rim with short acuminate teeth; 36h PHYTOLOGIA Vol. 9, no. 6 red, clear-red, dark-red, blood-red, bright-red, fire-red, or vivid-red to cinnabar-red, scarlet, *prilliant scarle t, ri or bright crimson [other colors are reported, see below], tube about 15 mm. long, glandular-pubescent, about 3 times fe long as the calyx, the limb large, ss mm, wide, the lobes obo- vate, deeply emarginate at the apex This rather perplexing but comm on species was based on two ee of James Tweedie (nos. 50) and 505) from "Alacripori- » Rio Grande do Sul, Brazil. The collector was cited as Wtweed® by Schauer in 18h7, but this error was corrected by him in his 1851 work. The same author cites also Treedie | 460 "in cam- pis vig dad Sin A Platensium, ad S. Fé arana. Its + the tes aeweg name proposed for this patra may vasilitte be be vecheta arranana. The article in which this name is proposed is unsigned. 1 assume that Joseph Harrison, editor of aA the journal, was the either, te this is just an assumption bas on a memorandum from Dr. H. W. Rickett wherein he cree Sa e of the specific epithet to "arraniana"” and credited it merely to horticultural origin. Other authors credit it to Niven. — & Henslow (1838) say "This variety, which Mr. Niven has mmed Arraniana......seeds sent by Mr. Tweedie, in 1834, to Mr. baie of the Glasnevin Botanic Garden, Dublin, by whom a single plant was raised." They call it "ur. Tweedie’s Vervain, Purplisir flowered variety", but V. incisa does NOT have purple flowers. whenc e the lovely V. tweediana, and where grown in contrast with the other ri Will have a pretty effect. It has been named in compliment to the Earl of Arran." Walpers (a8he) classifies V, incisa in his Section Verbenaca, Subsection Inermes, Group Foliosae, Subgroup Macranthae, and Sec- ondary Subgroup Aubletia, along with ten other species. Schauer (1847) placed it in Santiok Verbenaca, Subsection Nobiles, with 6 other species. He points out that it is closely relat ated to Ve Osten Bait’s = Bailey 1234 "pink", on Herb. Osten 13572, Schreiter 5151, and An- isits 1976 "rose", on T. Meyer 277 "pale "rose", on Lillo o 6181 "purple", and on Herter r 8Lh32, Rojas 11785, and Ruiz % Huidobro ———— 196), Moldenke, Monograph of Verbena 365 3565 "violet", I am not convinced that these collections are true Verbena incisa —- they may represent hybrids with other species - although Eyerdam & Beetle on the label of their no. 22954 say that the flowers aft V. incisa show a "large range of color varia- tion, pink, scarlet, dark red, lavender, etc." Briquet nel gives the following characters for V. incisa: "les inflorescences.....non pas en capitules corymbiformes...les e cali t cumin olle deux & trois fois plus grande [than in V. tomophy limbe étalé mesurant 1,5 cm. de diamétre, et , glanduleuse extér- sie tonal en, in a memorandum written in Montevideo and dated January, a, cave "Zu Herb. ea caer No. 8046. Verbena 'Briquetiana'. Cf. Briquet Verbenac anae in Anmulaire du Conserv. de Genéve gt hen (War-W211) - 288 « a Cf. Briquet in Arkiv f. Botanik II no. « 190k. Arbeiten von Chodat (Plantae Hassler- eine grossere von Verbenen gesandt mit meinen Bemerkung- en, habe aber nicht einmal Bmpfangsanzeige erhalten. An Hassler senate on ihm schriftlich die Nac , dass e rnardino, ich habe ihn leider nicht sehen k@nnen). "Briquet behauptet, dass V. chamaedryfolia in Paraguay nicht vorkomme., Das ist richtig soweit es sich um V. Melindres Gill. handelt, V. melindroides Cham. habe ich von Misiones und dttrfte edenf: (turma, grex) V. chamaedr. zusammenfasst, so gehtrt diese Form 8046 jedenfalis « dazu. Ich habe hier in Uruguay, in Paraguay, in Argentinien gefunden dass - Bltttenfarbe der einzelnen Arten sehr constant ist, dass dieselbe Art rot, violett, weiss bitht, ist absolut pesceniiaane (et. Chodat in Plant. Hassl.t). So-. weit ich aus dem Beschreibungen Briquet's ersehen kann, hat er diese Form zu phlogiflora rp eS ica var. Tweediana) als forma truncatula gezogen. Sie hat nichts damit zu tun. Ve phlog- i de: B80L6 in meinem Herbar als 'Briquetiana' bezeichnet und halte sie ftir den Uebergang von V. incisa Hook. zu der Ve scrobiculata Griseb. (symbolae no. 1736). V. incisa Hook. aus der zone, s/Br. Uruguay {nur fluss s littoralt]; Ve scrobiculata = It erenah 3. are Verbena incisa Chod. is actually @ synonym of V. tomophylla Briq. The The type of V. fulgens is a specimen collected by Aimé Jacques Alexandre Bonpland in Cor 366 PHYTOLOGIA Vol. 9, no. 6 rientes, Argentina, and deposited in the Bonpland Herbarium the Botanisches Museum in Berlin. The type of V. incisa ek ye Herb. Martius s.n., collected, presumably by Carl Friedrich pp von Martius, from a cultivated plant in the Munich Botan- ical Garden and deposited in the herbarium of the Jardin Botan- ique de 1'Etat at Brussels, with a note on the label that it is perhaps a hybrid with V. teucrioides. The type of Vv. chamaedry- folia < melindres f. siccanea was collected by Cornelius Osten (no. 10612) at La Falda, at an altitude of 1000 meters, in the Sierra Chica, on April 20, 1917, and is deposited at the Museo de Historia Natural at Montevideo. The type of V. megapotamica f. truncatula was collected by Emil Hassler (no. 567) on a campo at roneously listed by me (1959) as a synonym of V. scrobiculata sco ee but this is not true unless Osten is correct in n his interpreta- tion of that species as be e common Tucum4n plant -—- an th c P opinion which I cinhees when I first examined ae’ identified ve Soe material, but to which I do not now hold e type of V. briquetiana ana and of its f.c ampestris is ap- parently Osten 8016 from along a roadside at San Bernardino, Par aguay, collected on August 18, 1915, and deposited at the Natur~ kho historiska Riksmuseum e that of V. briquet iané f. silvatica and of V. melindroides f. briquetiana is Osten 9017 places at S. San Be 0, collected on August 11 ’ 1916, and deposi in the same herbarium. ° ese names re erroneously ref to V. scrobiculata in my 1959 Bailey & Bailey 925 and 925b are typical V. incisa, but no» 925a — "runners and flowers from one plant, fem colonized along railroad" -- is d doubt fully oe Presumably, however, all were collected at the same spot. R. E. Fries “ser may be @ hy- and the col- rar a 6: 331 (1941), as well as in my Prelim. Al Invalid Names 6 (1940) and Alph. List Invalid Names re (isk), by me as : incisa f. albiflora Osten & Moldenke, which see- Morong, g, Britton, & Vail (1892) cite Morong 51 as V. peruviana oath be noted here that the name, Verbena incisa, appa site should be accredited to Marnock who first first published it with a color plate, not there credited to Hooker or anyone else, 196) Moldenke, Monograph of Verbena 367 in 1838, It is also very possible that the proper name for this taxon is V. peruviana (L.) Britton, a name based on Lychnidaea, veronicae f folio, Floss coccineo Feuill. (1725), to which Linnaeus in 1753 merely added a Latin descriptive sentence and the binom- ial, Erinus peruvianus, Feuillée's type was collected on the north bank of the La Plata River in Paraguay (not in Peru) and his description points to V. incisa rather than es the taxon now passing as V. peruviana. I hesitate, however, to make this change ere made, then what is now passing come V. peruviana (L.) ne and what is now passing under the latter name would become vec hamaedryfolia A. L. Juss. Numerous vernacular and common names have been recorded for V. al “rae. SSedavidlienans "cut-leaved verbena", Ncut-Leav— - veingeschni ttenbiattrige Verbene", "jurujuba", , margarita macpeitan! margarita del cam punzo", "margarita punzé", Pectin roja", "Mr v i eer "scarlet sip ero, "verbena roja", "verbenen", and "verveine a feuilles incisées Herbarium material of this rn PEO has ar "widely misidenti- fied and distributed under such names as V. chamaedrifolia Juss., V. chamaedrifolia Le, "Ve face mers lia var. nov.", V. chamae- Gryfolia Juss., V. chamaedrifolia var. melindroides Schau., V. melinires(Gsils olia ioides Cham., V. megapotamica Spreng., V. meg tweediana (Niven) Kuntze, V. melindroides Cham., cz savtheua Spreng., V. peruviana (L.) “Britton, V. peruviana R. & Pe, V- platensis Spreng., V. scrobiculata Griseb., V. tenuisecta Briq., V, tweediana Hook., and even Lippia nodiflora (L.) Rich. Osten annotated Rojas 3395 as "V. chamaedryfolia Juss. hybrida?", while on Jacinta s.n. [Herb. Osten 7079) he says "V. chamaedryfolia Juss. ad melindroidem vergens On the other hand, the vt Hort. Bot. Basil. s 1839] and the J. K. Small 8745, distributed in Pin herbarls es V. in cisa, are eager Ac! TV. hybrida Voss, Ekman H.12615 - ue seme tenn secta Briq. Face ensen 2636 cited in my Alp s : 2: 599 (ge). ete Osten =e 11582, and Herb. Inst. Miguel Lillo are V. tomophy ll. e (19 190) Yegards V. incisa as barren’ Eghied f tecunes with flat rents ope ° = a ee eee mining c+ patty 8. rados, in dry er eds and fl , Sdge of r, along dry roadsides, on uncultivated campos, on dry 368 PHYTOLOGIA Vol. 9, no. 6 sandy banks and dry _ banks, in waste ground, and along rail- road tracks, mostly in full exposure to the sun, growing at alti- tudes of 20 to 3100 meters, flowering and fruiting in every month of the habe’ Troncoso (1937) reports it as cultivated in Buenos Aires, Argent erdam & Beetle found it in sandy alluvial soil near stream- beds in the shade of shrubby yellow-flowered mimosas, while Ritter found it growing in a region of 1.5 m. rainfall and 10--35° temper ature variation. Schwarz reports it as "rare" at Gob. Roca, Mis~ iones, but Quiroga says that it is "abundant in high sandy waste soil" in Misiones. Rosengurtt found it "common along roadsides aioe shrubs", Jtr pein reports it "common on the campos oe at oh larica®, common at Formosa, and common as El Candado in Catamare Morong says "this pied -tleeiced trailing Verbena seems to mn all over Paraguay and [blooms] nearly all the year round. I found it not only in copses about Asuncion, but also in the streets of = city, and far up on the Pilcomayo River. It is equally com- a hundred miles east of Asuncion. The stems sometimes climb ay among bushes for 6 dm. or more." Mexia describes it as "all along the roadside from Tucum4n to Racas....one of the commonest flowering plants." Schulz reports it as "abundant" at Colonia Benitez in Chaco, and "in fertile soil of fields, abundant" at des 3 Hunz Herzog says that it is very common in the potrero at Charagua, and eyer a that in gard disturbed ground” at Resistencia. He says for his no. 277 that it is "very rare", but since he describes the flowers rs of this collection as being pale-rose it may well be that it represents 4 hybrid. G. J. Schwars o8 is anomalous -- its leaves are not 49 ovate as is species Hooker & Jackson (1895) state that V. incisa is native to Panama, but this is certainly an error. The es is know to in Italy in 1842, and in Spain in 1849. It is recorded as cultiva~ ted in Argentina by Parodi (193) and Troncoso (1937)- Rtmpler (1873) says: "Brasilien. — Einjxhrig, im Gewttchshause ausdauernd. —- Diese Art, obschon aie ziemlich allgemein nur als eine Variettt der vorigen [V. platensis] betrachtet wird, unter- rosenrothen und geruchlosen Blumen. Aus Samen Smee diese Art mit allen ihren Merkmalen unverndert wieder und niemals bemerkt unter den Stmlingen Pflanzen, welche mit Verbena teucrioides OPP Ee h#tten. Auch von dieser art hat man durch Kreuzung ent der vorigen Art eine Anzahl von Varietaten erzielt, pai A txt der rosenrothen Ferbung a0 cue, mana und Verwendung 196), Moldenke, Monograph of Verbena 369 wie bei Nr. 7 [x¥. hybrida]." Augusto (1916) cites an unmmbered Tweedie collection from "Brasil meridional" ne as well as unnumbered collections by Augusto, Augusto & Edésio, and by Bormmeller from Rio Grande do Sul. The Seiskeehess £. 180879, cited below, does not bear any indication ya ep label t seat Ih it was from a cultivated plant, but I am assum- The pertan Bot. . 65: pl. 3628 reference as sometimes cited as "1837" or ni858", ee this plate appears not to have en published actually until 1839. It should be toate: also that Marnock, in his Fags ron 5: iy ol (1840) cites his Verbena trium umphans to "fig, 3" of plate 5. The numbers, however, is sean to have become tr ransposed on the plate — "Ve tri illustrated as fig. 1, fig. 3 being Ve. rigida Spreng. and fig. 2 being Lythrum roseum Marnoc The Florida record for V. “incisa given by me in my Résumé ao 5: 2 (1962) was based on J. K. Small 8745, which I now regard as xV. hybrida Voss. In all, 310 herbarium specimens, cad ol type material of most of the names i have been ed by me. Citations: BRAZIL: Paran4: Dusén Leh (8 (S, W--1481775); F. Co penne 8.ne s.n. (Curitiba, O6ks 18, 1928] (N, oS 95)) Rio Grande © Sul: Collector undesignated gee [Herb ac. Rio ref. V] ms Henz 33436 (N); Leite 250 (N); Yalme 6u0 by 127h (S)5 Rambo 8990 (Sp--50983); Ritter 35460 (S); Je Vidal s.n. (Ja— T6518). sa Santa Catarina: R. Fischer 3 sen. {Lencol, ol, Feb. 8, 1920] (Sp—3679); F. Miler 96 . (Ja——6534); Reitz 2233 (N), C ¢.882 (Ja- 55301, N). SHo Paulo: Bailey & Bailey 925 (Ba, a, Ba), 9258 (Ba), 925 (Ba); A. Betzler s.n. (Herb. Jard, Bot. Rio Jan. 61346] (Br, Vt), BOLIVIA: Santa Cruz: W. M. A. Brooke 5550 Herzog 1217 (S). Tarija: R. E. Fries 110k (N, S)+ PARAGUAY: Ce Alvarez s.n. [Asuncion, 195k] (Je—0503, Je—8657), sen. [Piri- bebuy, 195)] Soest Anisits 1978 & 2238 (S)3 Archer L666 (N, N, W—1705489); Fiebrig & ig & Osten s.n. [Herb. Osten 8062] (Ug); var BE 567 (N), 258h (N), 9252 (Bm), + 133s Lice poli’ = f 0 sten 2 > 4, at pega 3772 7 lies rae S, 8)j Ae Late 1505 2)" he Meyer ten (K), > 16162 ne Worong 51 (C, Mi, » ¥—-71998, ¥-117059)5 Osten 806 (N, 5, Ug), 9017 (S, Us); Bax E Edw. Palmer 8- = (Capt. Page Exped. 185k] (W—227576), cg amen nakien . age La Plata Exped. 227578 T. Herb. 7899] (Ug), 1894, ifn dete “ Be a isneen 13572] vg), 3395 {Herb. Osten p05) (Oe ate SMe cnt Apa 35 nah Hicken 12: 92] Ug), SMe Herb tier’. Osten iet2) (0)) Sandeman 4861 1s be A. G. Schuls 7815 ~ 370 PHYTOLOGIA Vol. 9, no. 6 (es) Woolston 731 (W--233828)). URUGUAY: C. Alvarez s.n. [Trini-~ Junio 1954] (Je—8703); H. H. Bartlett 21096 1096 (W--1907618) ; ie ie (Herb. Herter 829)1] (B, W--1422055), sn. [Herb. Os- ten 19218] (Ug); Osorio sen. (Ug—139h3, Ug—13989); Osten 3196 Oe UB, Ug); Rosengurtt B.22hh (N, Ug--5606). ARGENTINA: NA: Buenos ham s.n. [Buenos Ayres] (Bm); Sota 5 (N), 51 (N)- ptt Brttcher & Brttcher s.n. [21/1/19] (Er); Jorgensen 1028 1028 [Herb. Inst. Miguel Lillo 31350; Herb. Osten 11421) (Av, Ue)s Rodri Vaquero 907 (Gg—~353257, N); Schreiter 61,37 (Herb. Inst. Miguel Lillo 32771] (N); Sleumer 269 (ca--l)9220, W— 2168457); Venturi 7164 5: casey Chaco: Re Me A Buy (S); Boffa 1090 (N, N); JUrgensen 266 [Herb. Inst. Miguel Lille 313753 Herb. Osten 11866] (N, N, N, Ug, W--1055176); Malvarez 1365 (Gg--352673, N); T. Meyer 143 143 [Herb. Osten 22661] (Ug), 277 [Herb. Osten 22945] (Ug), 290 (Ug—10932), 2673 (N); T. Rojas 11785 (MN); A. G. Schulz 277 (Herb. Osten 22915] (N), 1475 (N), 3988 (N); C. C. Schulz 783 (N, St); Venturi 33 (W—-1013609) « Cérdoba: Caputanelli s s.n. (Ruiz Leal Leal 15990] (Ss); Lorents 13 ee 436 (Sg—68311); O'Donell & Rodriguez 1,81 (N, t—-72505d) j 10612 (Ug), 17086 (Ug); Rodrigo 246 (S); Ruiz Tesh 12099 ar Corrientes: Bonpland s.n. (N); Ibarrola 770 (W—-19 31120) 3 Rodrigo 705 (N, N), 855 (N). Entre Rfos: T. Meyer 10154 (N); Ruiz Huidobro 3493 (N), 3565 (N), 3603 (N). Formosa: ca a (Herb. Inst. Miguel Lillo 31495] (N), 5151 [Herb. Osten 2001 13) (Ug); Venturi 5270 (Ca~-3h2587, Gg--158412, N, W-—-1591468)3 Je =e (Ca—561720). Misiones: Bertoni 19h2 (NW), 2252 (N)>_ Ana, 15-III-194h; Herb. Inst. Miguel Lillo 98)12] Seapine Ekman 1978 (N, S); Grtmer 6 [Herb. Osten 23182] (Ug); J. H oer 773 (N); Jacinta sn, (Herb. Osten 7079] (Ug); iris = aes of Colonia Bonpland] (S); Montes i (Nw), 2435 ae N)3 Quiroga 16) a, Osten 7663) (U (Hs 4 Ge suas 6869 (Sz); G. J. Schwars n 7653] (Ue); Rodesgues = (Ss), 1243 (S), 2052 pS “WY, 2780 (5), 3000 (N), 312h (N, 2), 2162 3162 (N)- Salta: R. E. Fries 557 (S); Hunziker 1033 33 (A 3 ios!) uae & Moldenke 19739 (N); Ruiz Leal 12627 HT ()) Herb. Inst, Miguel Lillo 31357] (N)~ San Luis: Pastore 207 i) Sen Santa Fé: R. Alvarez 858 (S), 909 (8, N)i blag WF ts), Job 1071 (N, S); Ruiz Leal 14321 (Ss); Terrib- seceteeen camer nemneR 196k Moldenke, Monograph of Verbena 371 ile 366 ft Ok), 448 (N, Ok), 476 (N), 492 (N); Venturi s.n. fHerb « Argent, Cienc. Nat. 23672] (N)- rpg del Estero: H. H. Dartlatt 2021 (Mi); Maldonado R RAAT? (N); E. Wall s.n. [Es- colta, 9/11/h6) (Bw) (Ew); Wall & Sparre s.n. wn, (Guardia, 9 ) (Ew). tusumdn: Bailetti 58 (N); Bellowe Sone a Th2/ra/igle) (N)5; Borsini 3 (Herb, Inst. Miguel Lillo 2003] (Ba, aggtadg Ut--115423b) 5 Descole 1548 (N); Descole & ins s.n. (Herb. Inst. Miguel Lillo 6 86115] (Bm, Ca--164751); Ferrey: Ferreyra a 059 (ok); R. E. Fries 8. n. (near Tucumfn, 16-10-8] (S); Lillo | Lillo 3220 220 {Herb. Inst. Miguel Tillo 31360] (N, N), 6181 [Herb. Osten 8158) (Ug); Lourteig 187 (N); Mexta 7832 (Ca--64y015, N); T. Meyer 9799 (N); Moldenke & Moldenke 19721 (N), 1972h (N), 19754 (Es, lg, N, Sm); Monetti 162 (Herb. Osten 1087] (Ug), 1623 THerb. Inst. Miguel Lillo as (N), 8.n. (Herb. Inst. Miguel Lillo 3135h] (N); a8 (Herb. Inst. Miguel Lillo 32193] (N); Rocha 3693 ( 200i), 8. n, [san Augustin, s-X-1951] (Ba); Ruiz Leal 12401 (Ss), 12621 Sade Schreiter 810) [Herb. Inst. Miguel Lillo 32610) (N), oe Villa 453 (N), 680 (N); Wall & Sparre s.n. [Sabadilla, 19/11/16) 46] (Ew). Province undetermined: Friedmann s.n. [1923] (Ba). CULTIVATED: Austria: Reichenbach f. 190879 Se Belgium: Herb. Martens s.n. [h. b. lov. 1630] (ar, Br); Me Martens s.n. [h. b. lov. 1839] (Br). Brazil: Bailey & Bailey 123k (Ba)s ° 38 (Ja—-6197), 7647 (Ja—-h651h), 7675 (Ja--h6515, N), 7705 (Ja— 46517, N). France: Herb. Harvey son. [h. R. P. 1836] (Du-166L59), S-n. [h. Re P., julio 10k)] (Du--106460). Germany: Herb. Martius San. (h. Monac.] (Br, Br); Herb. Mus. Bot. Lund. s.n- n. (Schwiebus, Aug. '72] (Lu); Herb. Prager 10613, in part ; (Gg). Italy: Herb. s.n. [hort. Bellovae 1042] (Du—166460). Spain: Herb. rb. Hort. Matrit. 23 (Q). LOCALITY OF COLLECTION UNDET : Co ee undesignated 20 ae 21 (Q), 22 (Q), 2k (Q)5 Herb. Mus. Bote Stockholm 2109x emt ( D.Sc 2 VERBENA INCISA f. RA Osten & Moldenke ex Moldenke, Phyto- logia 6. a 1949. = Y ciee ian isa lus. albiflora Osten ex Moldenke, res - Aiph. List I List Invalid tases 6, in age: 190. Be . Los. ‘sip a 48 ig. 19h Moldenke, = oe 4 idenke , Alph. List rear Names 48. ; - Sande o: 1%. 199; Moldenke, Known Geogr . Distrib. Verbenac., 372 PHYTOLOGIA Vol. 9, no. 6 {ed. 21, os & 198. 1949; Moldenke, Résumé 127, 367, & oe E eo This form dif from the typical form of the species ing a sing The type of the “form was collected by Teodoro Meyer (no. 518, in part) at Fontana, Chaco, Argentina, in October, 1931, an dis deposited in the Osten Herbarium at the Museo de Historia waisted at Montevideo. The collector calls it "verbena de flores blancas" but now feel that it is worthy of nomenclatural recognition. In all, 5 herbarium specimens, including the types of both names involved, and mounted photographs have been examined by me, Ci : ARGENTINA: Chaco: T. Meyer 518 [Fontana, Oct. 19315 Herb. Osten a 22609] (F—photo of type, N-photo of type, Sg—photo of type, Ug—type, Ug—isotype, Z—photo of type), 518 [C Colonia rary Dec. 1932; Herb. Osten 2296] (Ug); A. G. Schulz 1478 > XVERBENA INHONESTA Moldenke, Phytologia 5: 133. 1955. Syn : Verbena urtiet Polis x prostrata Dermen, Cytologia 7: 170. 1936. Verbena 1 lasiostachys Link x T. urticifolia L. ex Mol- denke, Résumé 368, 368, in syn. 1959. oe a urticifolia L. x V+ iostachys Link ex sorte: Résumé 377, in syn. 1959+ Verbena ur ticifolia x lasiostachys Senet ex “hdres Résumé Suppl. ppl. 2# 13, in ac liogra: » Cytologia 7: 170. 19363; Moldenke, Phyto- logia 3: 68 (9s) and and 5: 133. 1955; Moldenke, Biol. Abstr. 3: 1093. 1956; Moldenke, Am. Midl. Nat. 59: 353. Arg rrr R _— 223, 368, 377, & 172. 1959; Moldenke, Résumé 13+ This is the hybrid between Verbena urticifolia L. and V. lasi- men Vassac cies happen b intro- duced at some future date in each other's territory. I doubt if ++ would have any horticultural merit. INTEGRIFOLIA Sess6é & Moc., La Naturaleza, ser. (Pl. Nouv. Pree ed. 1] 6 es sroerbenat 1889 {not Verbens verbens ridged : Fbene integrifol olia pg ber & Moq. ex Moldenke, Pgh?” eo =p a AS é& Cgraphy: Walp., Sans Bot. Syst. ): 18. phos 6 Sess Woes, La Naturaleza, ser. 2, 1: app. [Pl. Nouv. Hisp., aR} Os 196 Moldenke, Monograph of Verbena 373 1889; Sessé & Moc., Pl. Nouv. Hisp., ed. 2, 6. 1893; Hook. f. & Jacks., Ind. Kew, 2: 1179. 1895; Hill, Ind. Kew. Suppl. 7: 2h9. - 20 pi mt Known Geogr. Distrib. Verbenac. Suppl. 1: 2. — Mol- enke, logia 2: 87 & 115. 1945; Moldenke, Alph. List Cit. 1: 233. ish mei vets Alph. Niet Invalid Names Suppl. 1: 21. 1947; Moldenke, Known - Distrib, Verbenac., Ted 2), 33 & 198. 199; Voldenke, Ste 39, 35 t 472. 19595 Lewis & Oliver , Am. ie, Bot. 8: 639. 1961; Moldenke, Résumé Suppl. 3: 10 & 39. Perennial bush; stems mmerous, to 1.8 m. tall, sharply ous, shiny, deeply sulcate; nodes contracted, annu- ea sa and t » Spicate, numerous omewhat paniculately disposed at the apex weit ha ee. slightly sca a pair of foliaceous bracts S—16 mm. long at t filiforn es slender, original very fragmentary description of this little-known np is "Verbena tetrandra spicis filiformibus foliis oblongo- ellipticis integerrimis. Fl. Mex. Habitat in Queretari circuitibus. Floret Maio. y.m The species is knom two collections, one of Which und open rocky soil in the pine forest area on open ky knolls in humid mountain eys, to 2330 meters altitude, flowering in and the other at $300 feet alti- tude, in June. Perry (1933) places it in her list of "Doubtful or Little- es" says of it "No known Mexi- Can species of Verbena has entire 1 ° » une try specimen cited below shows them very clearly and is said to have been taken from an "old" bush. The V, integrifolia of Michaux is a synonym of V. lex Lehm. and was published by Walpers (Repert. Bot. Syst. hit | ) in 374 PHYTOLOGIA Vol. 9, no. 6 nymy only, so therefore does not invalidate the use of the same reeithet by Sess6 & Mocino, Hooker & dake ag errone~ mely give the page reference of the Michaux nam » im stead of "18", According to Gentry V. integrifolia is called "verbena" local~ ly, as, of course, are many members of this — asian genera. Only 2° herbarium specimens have been examin an Michoacfn: R. L. Oliver 19. (Z)« Sinaloa: 174 (N). H. S. Gentry 617) xVERBENA INTER : in Chod. & Hassler, Bull. Herb. Boiss», sér. 2; ery tts Synonymy! erbena bonariensis x ovata Briq. in Chod. & Hassler, Bull, Herb. Boiss., sér. 2, 4: 1057, in syn. 190k. Verbena bonar- iensis L. x V. ovata Cham. ex Moldenke, Résumé 359, in syn. 1959. Verbena ovata Cham. x V. bonariensis L. ex Moldenke, Résumé 371, in syn. 1959. Verbena tobaty Davis ex Moldenke, Résumé Suppl. 3: hl, in aSGES os Bibliography: - in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, 2 1057. 190k) ip i in Chek & Hassler, Plant. Hassler. 10: k79. 190k Prain, Ind. . Suppl. 3: 187. 1908; ae ’ Lilloa bat 332 (191) and 8: “ID. 1942; Moldenke, Known Geogr Distrib, Verbenac., [ed. 1], 41 & 101. 192; Moldenke, Aiph. List Cit. 1: nor & "26h. "1916; Moldenke, Alph. List Inva nvalid Names Suppl. 1: 22 & 26, 197; Moldenke, Aaph. eee Cit. 2: 599 & 600 = and 3: “687. 1919; Moldenke, Distrib. verter ed. 2), 99, 106, & 198. 19495 Sioaaasae » Prytelogia Ae 75 % (1949) cm 32/289. 1950; Acevedo de Vargas, Hist. Nat. Chile 25: 57. 1951; Moldenke ~ a nara hs he 188. «19535 Moldenke, Am - Midl. Nat. 59: sal 35h. 1958; LD gpeey at résumé 11 a th sere 359, 371, & 72. 1959; Moldenke, Résumé Suppl. 3 Coarse erect suffrutescent herb, 0.5--1.5 m. wales stem pale ellow-green; leaves decussate-opposite, very stiff, pale yellow” green, paler beneath; bractlets pale yellow-green o purple-bros on the margins; calyx pale yellow-green; corolla blue or viole t if : i (no. 3324) on a wet campo at Caraguatay and (no. 619) in swamps near Tobaty, Paraguay. Briquet says: "Cette plante ext > intéressante posséde 4 peu prés l'inflorescence du V- 0 ovata, & bractées d'un violet foncé, mais les épis ne sorte que partiellement subsessiles et s en anbelle, ninsieurs dtentre aux s en » @n corymbe Lache. En les feuilles n'ont plus la forme largement eee si caractérist rapprochent distingue bien encore la VY. intweceians du V. bonariensis, sins toutes ses formes, par la texture plus Spaisse ot plus raide des 1964 Moldenke, Monograph of Verbena 375 feuilles, la face supérieure un + luisante, la rugosité plus 2s 2 cm de surface); la partie cupérieure de la tige est presque re les épis eal trés hérissés-glanduleux. Ces caractéres V. intercedens une segs intermédiaire entre les V. vale tabat et V. ovata, entre lesquels il osville par tous ses caractéres. Comme les V V. ovata et bonariensis croissent e 1 es cul ompliqu piesa l'interprétation exacte des membres de ce g entre les V. bonariensis et ovata." Jorgensen reports that ve intercedens is "common" on the campo at Villarica. It inhabits low campos, wet places, and swamps in various stages of periodic drying-up, and has been eal iasted in anthesis and fruit in oie Novemb J the names V, bonariensis L., V. bonariensis var. longibracteata Kuntze, Ver rigida Spreng., and ° Vv. venosa Gill, & Hook. On the other hand, the JOrgensen 3768, distributed as xV. intercedens, is sctualay ¥ ve a Le “TL. The Berro specimen | cited below is very abno e and may be virus-infec . Acevedo de bine ll “(951) 8 stakes aha mothers Pd photograph 17422, cited below usly distributed as V. landbecki R. A. Phil. in her h eathape It is of interest to note that V. ovata — one of the reputed ts" of this has c , although it has been found in the other localities where the hy- has been e ne would suppose that bo 18) Would be more plentiful than the hybrid everywhere where it oc- curs and would, pape aig She ected sooner and m ften. Peds there seens to po ubt whatever that V. ovata is one of he "parents" of thi ve fart Jorgensen 377k sellestic, cited below, is 4 mixture with V. bonariensis var. conglomerata Briq., prey as a result = other cases a In all, ig herbarium specimens, including type ma material 0 all the names involved, and 2 mounted photographs have been exam- ined x pee Cita : BRAZIL: Rio Grande do Sul: Rambo 43510 Ser Go), 49696 re PARAGUAY: archer 1,821 (W—1709560); Hassler 6149 [Mac- 376 PHYTOLOGIA Vol. 9, no. 6 bride photo 17422] (Ca-—9)361—cotype, Kr--photo of cotype, N— cotype, N--cotype, N—photo of cotype, S~-cotype), 893k (V); Jorgensen 3770 [Herb. Osten 222)5] (N, Ug), 3774, in part (Du-~ 29026, N, S)3 Woolston 785 (B, W—~2338285). URUGUAY: Berro 1626 (Ug); Rosa-Mato 728 (Ug--100k7). ARGENTINA: Misiones: Ge ei somam 1 1896 (N). CULTIVATED: West Virginia: Davis & Davis 3 8061 INTERMEDIA Gill, & Hook. in Hook., Bot. Misc. 1: 166. 1829 bast V. intermedia Penny, 1832]. Syn Verbena gracilis Cham., Linnaea 7: 261--263. 1832 (not Vv. pantie ae Desf., 1829]. Verbena tenuis Steud., Nom. Bot., ed, cae 2: 751. 181. Verbena chamissonis Walp., Repert. Bot. Syst. h: 18, 1845. Verbena intermedia Gill. ex Lorentz, Veg. Nordeste Prov. Entre Rios 150. 1878. Verbena intermedia Gill. & Arn. ex R. A. Phil., Anal. Univ. Chil. 90: ~~ 903 616. 1896. Verbena in- termedia Schau, ex R. A. Phil., Anal, Univ. Chil. 90: 616. 169 Verbena bonariensis a brevibracteata Kuntze, Rev. Gen. Pl. 3 (2): 254-255. 1898. Verbena bonariensis p longibracteata Kuntze, Rev- eata Kuntze ex Briq., Arkiv Bot. Stockh. 2 (10): 10. 190k. Ver- bena carollata Briq., Ann. Conserv. & Jard. Bot. Genév. 10: “jo1-- 102. 1907. Verbena corollata Briq., Ann. Conserv. & Jard. i Genév. 10: 102, sphalm. 1907. Verbena bonariensis var. longi: bracteata Kuntze ex Moldenke, Prelim, Alph. List Invalid Nam Names L5, in syn. 19h0. Verbena carrollata Briq. ex Moldenke, Lilloa 1h: 53, sphalm. 1948. Ver! Verbena bonariensis f. brevibracteata : Kuntze ex Moldenke, Résumé 35 359, in » 1959. Verbena bonariensis f. aes oe Kuntze ex di Rag e's 6 399, in syn. 1959+ Verbena is Schau. ex Moldenke, Résumé Suppl. 2: 10, in a7. 1960 Toot V. bonariensis Dill., 1938, nor L., Sart tt 4, nor Vell. 1959]. Verbena rigida var isa, & Hook,) Farwell ex sP). tetoons cigs Suppl. cr aoe os , ography’ Gill. & Hook. in Hook., Bot. Misc. 1: 166- soa A. DC., - 11: 5h1. 1847; Schau. in Mart., Fl. B : 188. 851; Lorentz, Veg. Nordeste A os, ed. 1, 19, 4, 150, & 172. 1878; poe Abh, K. Gesell. Wiss : ise e Fi. Argent.] 276. 1879 ape & en ° Hegre 2 (bot): 26h. 1861, Lik hho, Fl. Tuc Britton, & = £ Vail, Ann. N. Soe Acad, Sci. 7: "198. 18933 Hook. acks., Ind. Kew. 2: 1178 & 1179. 1895; R. A. Phil, Anal. uav 196) Moldenke, Monograph of Verbena 377 Chile 90: 616. a Ba og Rev. Gen. Pl. 3 (2): ser iey riba Speg., Contrib. . Fl. Tandil. 41. 1901; Brig. in Ch Hassler, Bull. ae Boiss. +, s6r. 2, kh: 1057. 190k; vag in toa. & Hassler, Plant. Hassler. 10: 479. 1903 Briq., Ann. Conserv. & 1. 190k; t . Bot, Genév. 7-8: , Arkiv Bot. Stockh. 2 (10): 10. 190); Macloskie in W. B , Rep. eton Univ. ott Patag. 8 (2): 686. 19055 Briq., Ann. Conserv. & Jard. Bot. °.05 101--102. 1907; Prin, Ind. Kew. Suppl. 3: 187. 1908; Hi » Fl. Pile. 102. 1909; Hicken, alee Plat. Argent. 196. 19103. TG Mém. Soc. Bot. France 1: 5h. 1 Hauman-Merck, . Mus. ent. Hist. Nat. Buenos Aires te 3. 1913; Prain, Ind, Kew. Suppl. h: 245. ; Anal, Soc. Cient. Argent. 1913; Sanzin, 4 uenos Aires 88: 98, at & 13h. 1919; ao. Physis 5: 22. Monti oa 3: 359. 1938; Herter, Re Bot. 6: 97. 1939; Troncoso, Darwiniana 3: \ o—fs2 & 485, fige- 1&3 e& f. 1939; Moldenke, Prelim. Alph. List Invalid ; denke, Known Geogr. Distrib. Verbenac. ed. 2 4, ete isha; ite ey He Bip sat Invalid Names isis < bi. i9li25 enke Lilloa 10: 346 & itn 1glh; Cabrera, DAGI Pub. Tec. 3: “tS & 75. 19453 Moldenke, Alph. “Aguas Gab 1: er DASE 197, 219, & 266. 19h6; Ro t, {63 Schnack agus). 31 : eon 693, 703, 70h, 781, 782 a 923 (1 ali), ye 1128, 1196, & 1300. 1949; H ldenke, eek oe 20: 175. a Ros oa et ae Mas ’ trib. years {ed » 105, oi & 198. 19493 Moldenke, da 3: "6 & 13 (asia), are oe 89, & 305, 1950; Martinez aki . » k: 178 & 226. 1950; Martinez ese Bavist ric. kh: 362, 386, & 2 1950; Rambo, Sellowia 7 ng ne : » 110, 119, 126, 127, 359, 362, 7 67, 3 37 ae 19595 wo Moldenke, a 2: 5, 10, & 12 (1960), & 1962), and k: 10. 1962. (962), an bs 30. 3: 481, fig. ee. 3 e & f. 19393 Schnack & Covas emania 32 8, fig 21. 197. Tall {biennial or perennial herb a woody creeping 378 PHYTOLOGIA Vol. 9, no. 6 rhizome, grayish-green throughout; stems agent simple or some- times branching, stout, strict, 0.5--1.5 m , green, sharply tetragonal with rather prominent angles, very T rough on the acute sulcate and longitudinally striate between the angles, subco very short; leaf- blades creck. stiffly coriaceous and rather rig- id, uniformly dull- or light-green on both surfaces, not brunnes~ cent in drying, 1--8 cm. long, 2--13 mm. wide, decreasing in size upwards, the lower ones cuneate-oblong or elliptic-oblong, rarely those immediately above, usual ly 3--8 cm. long and 5--10 a wide, those on the middle portions of the stem linear, oblong, lanceo~ cuminate-mucrona ° even eamcgt sparse in the inflorescence, linear, 1--5 mm. pn set mm, wide, teictuat-aake; stiff, entire, ciliolate; midr? rather crowded, few (3—5), di ays many-flowered, the one 28 heigl sessile and the lateral ones shor’¢~pedun- culate, rather thick, 1.5-—-8 cm y ietee normally 8—10 or more Fol lindric, glandular-hirtellous throughout, flowers and fruits closely imbricate; peduncles slender oF rath- sharply t olet, a tubes, sometimes with the pares ccoteyme purple 196 Moldenke, Monograph of Verbena 379 of this often pr Semone species was collected by nae Gillies on the pampas of Buenos Aires, Argentina ("in Provin- cia Bonariae, et in ee non raro occurrit"), and is deposited in the herbarium of the Royal Botanic Gardens at Kew. Cotypes of V. gracilis were collected by Friedrich Pye at Barado and ne rupa "in cam ampo de Tacagua grande", Brazil, are deposited the herbarium of the Botanisches Museum at er Cotypes a ve tandilensis were collected by Carlo Luigi Spegazzini (nos. 2408 and 2109) in January, sn at Tandil, Argentina, The type of V. is given erroneously as "100—101" in eee 8: 29 (1942). 4—6)", revibracteata is based on undesigmated collec- * unén, Argentina, from Sica stg pretest and fram Montevideo, U and is described by - ruguay, flo s breviores vel subaequilongi." Seckt (1930) gives V. caneocene H.B.K. as a synonym of V. intermedia, but this is er US » : en misinterp reted the statement of Schauer (1852) egarding V. gracilis Desf. (not Cham.) being in his opin- ion conspecific with V. canescens, a distinct species of N Texas, and Mexico, Verbena bonariensis Y ong iractenss Walp. 4s conspecific with typical V V. bonariensis L Verbena intermedia ran anges from sookheen Brazil through Urugu- et celaiiathieihiatediiniacreteeaiiias ay far into central Argentina. In Uruguay it has been collected at ne of 30 to 50 meters, and in Argentina at 50 to 2000 meters, the a pels from the hig numerous other collections -acehpcweas of the spikes on these specimens are to very co The apeotes fa has s been found ty collectors in meadows and dry meadows, sy places and savannas, rocky places, rocky or stony hills, toc pat or dry campos, stony or dry rocky campos eta sunny campos, pempas, 5: swamps, grassy slopes, hills and ar- royos, dunes, riverbanks, borders of "acequias", in the wet 380 PHITOLO@T 4 Vol. 9, no. 6 soil of meadows near rivers, in grassy rather wet campos, on mountains, in "lomas" [dry places], and on the shores of rivers. It is oh to be het as in fields, and has been found also on ly near rivers, and has been collected in anthesis from october to July, and in fruit in January, February, October, and November. pos= ites, and refer to it as an "annual", Hunziker found it “rather frequent" on the shores of Rfo Amzacate and also "rather frequent" on railroad embankments at Frenque Lanquen. Rosengurtt reports it as "common along railroad tracks" and, - a 1946 work, de~ scribes the plant as "Hierba frecuente en campos poco pacidos, enes o de rastrojo". Ragonese & Covas (19h7) refer to it as "Especie bienal o perenne, de tallos erectos, frecuent et collection from Ne anges do eee ridional", e of Rosebgurtt Cerro Uruguay, ro) Sorter from iyare , Uruguay. Ragonese (19h) Cites Govan & & Ragonese 3505 from Santa Fé, Arg » While Martinez Crovetto & Piccinini (1950) cite ele uns from Entre Rios. They cite the sas publication of t cies as "1830" and describe the plant as "Planta ae eg per enne, con flores aguladas, comin en oan, toda la Mesopotamia; Briqet's comments fan his V. Sats are perhaps worth repeating here: "Nous avions rapporte cette espece en 1899 (in Amn. Cons. et Jard, bot. Geneve, IV, a ey V. littoralis Kunth, concu & cette eopque dans un ses trés vaste. Une oe plus at~- tentive de ce » fait depuis lors, nous a a detacher du V, littoralis plusieurs types parfaitenent Poprereti (v. isa- bellei, ¥. cordobensis). Le V. coroliata s'ecarte a son tour du guement exsert (depassant a peine les dents calicinales dans le ve. ite oo son gos ies oblong, plus grand, ses epis presque _, leet (1896). was nig ai opinion that the plant Eaaertinter + intermedia by Schauer (187) is not ar MEF with tha - oa & Hooker (1829), but the bt to me that they are t - Rosengurtt (1943) describes the species "Hierba hemicriptéfita, florece en y os de otofic. Habita ras 7 e, erecta, de alrededor de 1,5 metros de ura, con tallo cuadrangular y hojas opuestas, sesiles, lanceo 196) Moldenke, Monograph of Verbena 381 gs rer Pte oe violdceas, et ly ge en —_ largas que m panojas definidas. Frecuen tepa patie pie it growing with vy. ie vets i and pianrd gor that it might be only a form of the latter or a hybrid with it, cam- (1943) report that its chromosome picture is on » 28, but in a 4 ort for "V. . = of 2n = 56, based on a plant cultivated at Pie Mendoza, originally from Sierras de Balcarce, Buenos Aire ard. Bo essentially characterized | by its short bractlets a hardly e- qual the flowers in length or very slightly surpass them, the corollas being small, exceeding pve Ringe betege by only 1—1.5 m, the limb being 1~1.5 mm, in diam r, and that therefore it is exactly synonymous with typical ve app Beer a type, however, on which we must base the name regardless of the descrip- tion given of it, and which I have examined, tedies to be V. temedia. Monticelli (1938) says of V. intermedia: "Es afin a la V. bonar bonariensis, de la que se aparta por la corola larga dos veces la longitud del edliz -" fTroncoso (1939) differentiates the two lows "A. Tallos estriados longitudinalmente, glabros, con las aris- tas de los angulos © menos asperas. Limbo corolar bien desar- rollado. Caliz y ener con pelos glandulares. pina grees Parciales distanciadas, c/u de 3--5 espigas, generalnente minal spin at las laterales " pusiaaiae. Hojas inferiores a S—-8 cm. de largo x 0,8--1 cm. de lat., borde dentado en su mitad superior, prea subiguales. Plantas ferde-centctentag = edia. "B. Tallos liso enero tos principalmente en las -aristas de loe angulos. Limbo corolar reducido. Caliz y bracteas sin pelos glandulares, Pdi ccdiaasaiah i parciales aglomeradas, c/u de ysl espigas subsesiles. Hojas inferiores de 9-13 cm. de long., 1,5 cm. de lat. , borde casi totalmente dentato, dientes dosigu ales, Plantas de color verde Wikebetis cab aaesiets nariens icons & Burkart (1946) say "V. intermedia....se ee aparte de los rasgos ya comentados, por flores més grandes y vistosas [then V. tristachya], de color violfceo intenso, limbo orolar bien desarrollado, c4liz y br&écteas con dig eacxageoae ms ones breve y estambres fijos en en la mitad inferior del tubo lar.......V. in y V. montevidensis pe pa "los tallo oe Macizos,..,..E1 estudio anatémico del tallo de V. tristachya pre- senta como caracterfstica interesante, que la capa de enorofiliano e es continua, es decir, que no sé nm los éngulo. ae s? ber of spikes as nintinisaat, Walpers (1845) places his V. chamissonis in Section Verbenaca, 382 PHYTOLOGIA Vol. 9, no. 6 Subsection Inermes, Group Foliosae, Subgroup Micranthae, and Sec- ondary Subgroup Holophyllae, while V. intermedia is plac ced in Subgroup Macranthae, Secondary Subgroup Melindres. Schauer (1847) correctly combines them in his Section Verbenaca, Subsection Pachystachyae. The V. intermedia of Penny re eferred to above is 8 synonym of V. canadensis {L.) Britton. A reference to V. intermedia in "Briq., Verb. Balans. Parag 4" has not been verified by me as yet. In Lilloa 10: 3h7 (1944) it is stated that Gallinal, Aragone, Bergalli, Campal, & Rosen~ gurtt PE.4605 is a mixture with V. 4th V. carollata Briq., Zq., but I do not now believe this and re regard it now as all V. intermedia. The com- mon names "berbena" and "verbena" are recorded for V. intermedia, tut are used also for many other species in this and d related gen- era. The Steudel reference given above is sometimes cited as "18,0", but appears to be more correctly dated 181. Jorgensen 2h68 is definitely V. bonariensis in the Montevideo herbarium and is definitely ve. intermedia in the United States National — ium — another example of this collector's habit of mixing § al collections under the same number! Hunziker describes the corolla on his no. 4,003 as being clear "morada" on its lower half and the rest ng ogee Tes ag & Arnott record the species from Chile, apparent erbarium material has been misidentified pai “distributed un- der the names V. bonariensis L., V. brasiliensis Vell., carollata Briq. On the other hand, Jtrgensen 3774, ‘Activate as V. intermedia, is actually ve. bonariensis var. " conglomerata Briq.; Kuntze s.n. (Montevideo, , 7/12/1891) is ve brasiliensis Vell.; Herter 1055 and Herb. Herter 82656 are Vv. gracilescens (Cham.) Herter; and Legrand . 1992 & 992 & 3809 a: are V. , montevidensis Spreng. Glaziou (1911) cites his no. 1162a from Rio de Janeiro, Brazil [his no. 14162 he identifies as V. litoralis H.B.K., but I regard it as as V. teats Vell. — 1 have not seen his no» 14162a], while Briquet (190k) cites Hassler 1026a and Regnell A.3645. Troncoso (1939) cites the following: URUGUAY: Rosen- Bart 3.765 (Si), B.2513 (Si). ARGENTINA: Buenos Aires: 3: carette 497 ee Hicken 475 (Si), s.n. Saat mg (gi-~3825) ee 3.M- -xI-1937] (S (Si—110); Pr Pelosi 18 (Si); spegazzini 2409 (Si), 2h09 (Si), s.n. [I.1901] (Si); Troncoso sone (2u-X1-1937] (Si--12h6); Valentini 502 (Si). Chaco: 3 Lynch some sane (19-1-190k] ee: Cérdoba: Burkart 7493 (Si); Nicora 1574 (Si). Co + Burkart 8053 3 (Si); Hicken s.n. [Bonpland, 1-1929] (S30) La Pampa: Monticelli C.17 (Si). Mendoza: Ruiz Lest , all, 122 herbarium specimens and 10 m ounted photographs, "type or or phototype material of most gage the names invo 196k, Moldenke, Monograph of Verbena 383 ved, have been examined by me, Citations: BRAZIL: Rio Grande do Sul: Malme 136 (S), 136b (N, S), 808 (N, S); Rambo 9718 (Sp--50987), 34723 (N); A. R. Schultz TU6 (Nl); Schwacke 11.336 (Ja--\6576); Sellow s.n. [Bras. —e Macbride photos 17L23] (Br, ig F--photo, Kr—-photo, N--photo, Photo, Bain oto, Z——photo); Vidal s 3.n. (Ja=~36857) « Saunt: : - [Sta. "Elena, Nov. aan 2] (Bm); Arechavaleta 3h (Ug), 32 (Ug), ai (ug), kl (Ug), 102 (Ug, Ug), 3135, in part (Ug); Berro 37 (Ug), 4423 (N), 8472 (N); Cabrera 3890 (N)5 CastelLance 8.Ne (Herb. Inst. Miguel Lillo 11756] (N, ); N); Gallinal, Aragone, Ber- galli, Campal, & Rosengurtt PE.2621 (N), PE. PE.2002 (N), PE 5, PE aos (i (nN, ed Herb. Her ee pe Bb (N); Herter 1155 (Herb. Herter 83295] (Ca- Herb. Osten 17032] (Ug), ae {[Herb. Osten (Ug), 4127 (eve 3 Montoro Guarch 574 (N); "oslo 736 (Ug--13265) ; Osten 3661 (Ug), 3863 (Ug), 3947 (Ug), l3h9 (Ue, Ue), 6567 (Ug), 7197 (Ug), 22669 (Ug, Ug), 21868 (Ug), 22650 (Ug), 22650a (Ug); Rosengurtt 6018 6018 Leaggeennns ), B.765 ene N, N), B.1915 ra 23642 (Ca--623654); Gillies s.n. [Pampas of Buenos Ayres) (N-- photo of type, Z—-photo of type); Hauman s.n, (Sierra Baya, V. ae. (Br); Hauthal Barthel Glo (N); Hunziker 4003 3 (Ns A. Ge G+ Some 5697 (Sz); Troncoso (W—-1858 34.7) 5 “Wall & Sparre s e erra Bac hiche, 2/11/16) (3), 8 son, (Sierra EL Volcan, 3/11/16] (Ew). Chaco: Jorgensen 21,68, part (W--1055178); A. G. Schulz 1481 (N). Cérdoba: Burkart ae fis re Osten 23368] (N, Ug); A. T. Hunziker 701 (N)j O'Donell & R 454 (Ut—115h05b) ; Os- ten 1311] (Ug); Pierottd — 3 Rodrige 20 (N); Sparre 122 2 2 (eo 1300 m.] (Bw)« Corrientes: Burkart 6053 (Ca—337h, W—1858299); Parodi 12h (N). Entre Rios: Ruiz Huidobro (N), 3550 (N), 3649 (N). Formosa: Jorgensen 3108 ) (W--1065729) « La Pampa: Fortuna 22 (Ca) A. G. Schulz 5842 (Ca—166282). Men- doza: D. 0. King 137¢ (Bm); “Lourteig 852 Y (Ca——166002); Ruiz Le- al 110] (N), 1171 (N), 1498 (N), 2980 (N), 5598 ( (N), 708% (N), 5500), 11388 (Rl); Ruiz Leal & Roig s.n. (Ruiz Leal 18809] (2); Sangin 3120 [Herb. Osten Sia) (Ug); Semper 343 (S); Wil- hoto) . Misiones: czek 56 [Macbride photos 21,683] (Kr--photo, N——photo Burkart 154k9 (N, W--2056387); Ruiz Huidobro 4995 1998 (Ws 2 a Schulz 6 Schwacke 22h9 Herb. Mus. Nac. Rio ref. (Ja). — stg c Roig 1327 1327 se Ruiz Leal 19077] (0k); Varela 566 (S); Viguts DP 7029 (W—-1858375). Santa Fé: Feddersen 36h, PHYTOLOGIA Vol. 9, no. 6 sen. (Aurelia, near S. Fé] (Cp); Ruiz Huidobro 3138 (N). Province undetermined: Wall & Sparre s.n. (Ew). I » ALBIFLORA Moldenke, Phytologia 3: 176. 19h9. Bibliography: Voldenke, Phytologia 3: *176. 1993 Moldenke, Kn Geogr. Distrib. Verbenac., [ed. 2], 106 & 198. 1949; Moldenke, Ré&- sumé 127 & 472. 1959. differs from the typical form of the species in hav- oro. ° ) The type of the form ap cutee ad Arturo Burkart (no. 4179 ve been examin Citations: ARGENTINA: Entre Rios: Burkart 1179 [Herb. Osten 22466] (F--photo of type, N--photo of type, ‘Sg-—photo of type, Us type, Z--photo of type). ENA INTERMEDIA f. GLABRESCENS Hauman-Merck, Anal. Mus. Argent. Hist. Nat. Buenos Aires oh) “103.29 1914. Bibliography: Hauman-Herck, Meabonds pat Nat. Bue- nos Aires 2h: 413. 1911; Molde oo - Diatrib. Verbenace, [ed. 21, 106 & 198. 19h:93 Moldenke, ar nae 137 "ge W72. 719596 This differs from the typical form of the species in being ving n aly subamplexicaul at the base, the flower spikes 15--18 mm. long and 4 mm, wide, the lanceolate bractlets, and the corolla often more than Aaa (but never 3 times) as long as the calyx. atic of the form was collected by Lucien Hauman—Merck (nO. = in part) near the shores of the Rfo Negro and its islands, in protected spots, Rfo Negro, Argentina, where the author says a = the species itself is very common [but from where I have 8° ens the corolla. No | ateiak = as yet been seen by me. VERBENA INTERMEDLA var. LANUGINOSA Moldenke, Phytologia 3: 118. 949 Bibliography: Moldenke, Paytologia 3: 118 & 135. 19L95_ Molden” , Known Geogr. Distrib. +» [eds 2], 106 & 198. 19495 Moldenke, Résumé 127 & hie. 19. This variety differs from the typical form of the species > having its s 8, peduncles, and leaves Laugier gs be aN sactasay very densely lanuginous. The corolla i s said to US » The type of the variety was collected by Adrian Ruiz oe es 2981) at the edge of a dry river at La Pampa, San Pablo, tied . Argentina, on February 6, 1935, and is depos in the Britton Herbarium at the New York Botanical variety is known thus far only from the original collection, of 1964, Moldenke, Monograph of Verbena 385 which 1 ee specimen and ); mounted photographs have been exami. ned by m Citations: " khoueriaa Mendoza: Ruiz Leal 2981 (F--photo -~ wee, N--type, N--photo of type, Sg—photo of type, Z—-photo YPe) VERBENA TR DTRGTS Moldenke, Phytologia 2: 237. 1947. Synonymy: Verbena pusilla Dusén ex Moldenke, Résumé 373, in syn. 1959. Bibliography: Moldenke, Phytologia 2: 237. 9475 Moldenke astanea 13: 117. 198; Moldenke, op nny ist Cit. 2: 598 (asta) Se he 12h9 & 1287. 19195 3; Moldenke, Known Geogr. eens ace, [ed. 2], 9h & 198. 19h9; pho Phytologia 3: 134 (199) and” Fd ole i955 spietite ld, Trib. Sc egady 19 (10): 166. aga E. . Kew. Suppl. li: ore 953; angely, 1 13. 195 1957; “cheney & Résumé 110, 373, & 1,72. 1959; Angely, ‘Le Paran. 16: 78 (1960) and 17: 6. io8l sci Fe herb, about 16 cm. tall, * sbundantly b fr the base, bushy; branches decussate-opposite, a Pest tetragon= al, rather densely arte ee brown when dry; principal ternodes 1--2 cm. long; leaves decus osite, subsessile; petioles 1 mm, long or less, ee piety Leaf-blades | ovate, tite, 7—25 mm. long, 5--15 mm. wide, each of the 3 divisions again lobed or incised, the lobes narrow and obtuse, more or less Hing llary, racemose, the racemes few-flowered, 1—2 em. long is less asus 1 cm. long during sigh Ppa cles filiform oe pies anthesis, to 1 cm. long ru: pube og as; bractlets ovate, about ’ ing-pubescent, otherwise glabrate, the apiculation ter anthesis; corolla hypocraterifom, th with parallel longitudinal ridges on tee lower half, glabrous, pg a type of this most interest and distinctive species was collected by José Eugenio Leite ee 37h) in wet places fields, Campos do Jordfo, at an elevation of 1600 meters, Sfo 386 PHYTOLOGIA Vol. 9, no. 6 Paulo, Brazil, in April, 1945, and is deposited in his herbarium at Novo Friburgo, Rio de Janeiro, Brazil, The conspicuous disks beneath the ovary, which remain in the axils of the reflexed bractlets after the fruiting-calyx and fruit have been shed, ren- der this species most remarkable and show its relationship to V. thymoides Chan. The species has ae collected in wet and grassy places fields, and low woods, on campos, and on rocky slopes, at alti- tudes of 750 to 1821 Ph eoitey blooming in January, April, and Sep- tember to November. Rambo found it "in campo alte graminoso sub- humido", while Leite says "very rare, near wet places". The type of V. pusi ijlla was collected by Gusta. Jtnsson (no. 1010a) in gras- sy places at Itaperussu, Paran4, Brazil, on September 27, 191b, and is deposited in the herbarium of the Naturhistoriske Riks- museum at Stockholm, In all, 28 herbarium specimens, including the types of both names involved, and mounted photographs have been examined by tations: BRAZIL: Parané: Dusén 7138 (N, S); Jénsson “= 10104 (S). Rio Heatag ts eck Rambo 36410 (F-~photo, N, N, “W, photo, S, Sg--photo, Z--photo). Santa nta Catarina: F. Midler 93 Cmca N)3 Reitz, & Klein 868 (N, Ok, B-2250703). ), 5193 (Ok, S, H— 2268975), 5422 (ok, We-2269003) ; Smi th & Reitz 10360 (W—-22L9372)» 10382 (Ok, W—225167h), 10385 (W——22h9 se}, AOL (H201097) 10489 (W—22h937k) 5 Smith, Reitz, & Klein 7 B3 (W——-2251305 . a S80 Paulo: Leite 347) (El—type, (nee Ate {Campos Jord#o, I-1949] (N, W—1953453). xVERBENA sige Moldenke, Phytologia 148. : Verbena racemosa x erinoi Be Mak “oytologia 73 163, fig. aa & 170. 1936. Verbena racemosa x erinoides alba Der men, cytologia 7: 170. 1936. Verbena racemosa x laciniata ta Dermen ex Woldenke, Am. Midl. Nat. 59: 354, in syn. 1958. Verbena FEES race- mosa Eggert x V. tenuisecta ee. ex Moldenke, Résumé mé 373, 10 syn. 1959. Verbena temuisec sa x Ve. racemosa Eggert ex Mol- Soc. Dict. Gard. hk: 2211. 1951; Mi . Nat. 59% ae A9535 Malderke Résumé 223, 373, ss & “he Reps Voldenke, R 12) * a Illustrations: Dermen, Cytologia 7: f 203s ig Be 19 36- . This plant is a hybrid between V. rt and Ve tenud secta Briq., produced artifi en a pointe in 1936, and said to have intermediate acters. The two paren” es both grow wild in Texas, the former native and the lat roduced. As yet I have no record of them from the same county» 196), Moldenke, Monograph of Verbena 387 but V. tenuisecta is spreading very rapidly and it is probably only a matter of time before both will be growing in close prox- imity to each other. The rid may then be expected in the wild. It ought to have considerable horticultural merit. VERBENA KUNTZEANA Moldenke, Pyles 2: Bliss diz" 198. rigged Verbena megapotamica hybrida Osten ex Moldenke, Ré- sumé 369, in syn. 1959. pig eicaetans "Yolde nke, Phytologia 2: WTh—b 75 (1948) and 3: 75, 13k, & 136. 1949; Moldenke, Known Geogr. pir Verbenac., [ed. 2]° 99, 106, & 198. 1949; He N. & A. Le ia Inst. Biol, Mex. 20: 13. 1993 Es Je et Ind. = 1l: 263. 1953; Moldenke, Résumé 110, 118, $e & ire. 79595 Moldenke, Résumé an: (ede 3: 14 (1962) and ‘1: eg Suffrutescent herb or subshrub, 3—6 dn. Spars poeeeitelieks ed from the base; stems fae branches rather ts Boinae tetragonal, the sides often sulcate, densely hirsutulous popes oe with widely divergent, wii > and peels esa ipped hairs, the lower part of the stems often decumbent; ago’ sz hirsutulous; floriferous portion of the inflorescence about 3.5 cm. long, to 2.5 om, wide during anthesis; bractlets elon- gated, linear-lanceolate, 11—13 mm. long, densely glan somewhat inc rs have fall tubular, the tube about 15 mm. long and 2 mm. wide, dense. glandular-hirsutulous with whitish hairs, the rim 5-apiculate, , densely glandular- hirsutulous with whitish hairs, the longest ones about ), mm. long; Corolla blue oe violet-blue to rose-lilac, long, dense hort-pubescent on the outside, the limb mm. wide, sinataly puberulent on the outer surface, within, by Carl Axel » in October, um of the ie dabariaiariale age ope poe tes that the native ° —eoaag notes Linge It was first identified The type of this distinct species was ee Magnus Lindman (no. A.3649) at Mintacctig he 1893, and 4 deposited in Riksmuseun +ockholm name for the afer is "flor de vovia". 388 PHYTOLOGIA Vol. 9, no. 6 by Briquet as V. paraguariensis var. latiuscula Briq. and then 4s V. platensis Spreng. It is named in honor of Carl Ernst Otto Kuntze, who did such splendid collecting of Verbenaceae and rela- ted groups in Asia, Africa, and America, whose keen insight has detected so many varieties and forms wo worthy of nomenclatural re- cognition, and who fought so heroically and eloquently for fair Lard and honesty in botanical nomenclature, albeit a losing The species has been collected in fields, campos, wet campos, and on high rocky ground, at 1200 meters altitude, blooming in "Verbena megapotamica Spr. hybrida". The Pedersen 4069, distrib- uted as "Verbena aff. kuntzeana Mold.", is actually V. nana Mol- eC. In all, 27 herbarium specimens, including the types of both paeet involved, and 6 mounted photographs have been examined by *“catat ions: BRAZIL: Minas Gerais: Widgren s.n. [1845] (Lu). Parané: Braga 1030 1030 (Z); Braga & Lange 255 fe 255 (Gg). Santa Cat Moreira & M 23h (Gg); Smith & Reitz 10161 mee por PARAGUAY: Sesh 5699 (W--113806); Hass: Dialer er 393% (N), 14u28 (N), 5758 (Ca—9 Fisbris $52 "), ‘ee (N), 662k (N), 11052 (Bm, Cb, oc a N, N--photo, Z--pho 3 Linden K.3649 — 0 ves type, isotype, N—-photo o we a S—type, S—isot —photo of type, Z—photo of type); T. Rojas 31406 mee arta 17906] (N, N, Ug), sn. gic Osten 15638] (N, N, Ug). ARGENTINA: Salta: Schreiter TF 1167 7 (N VERBENA LACINIATA (L.) Briq., Am. Conserv. & Jard. Bot. sae 7-8: 296--297. 190k nok A laciniata Kuntze, 19h1, n Raf., 1833, nor Sessé & Synonymy: Lychnidaea gplirtn ae tenuifoliae folio, vulgé Sandia-Laguen Feuill., Saae um. Obs. Phys. Cotes Orient. = Lah er 36. 1725. lychnidaea Verbenae termifoliae, vulgo Sandia sepe Feuill., Journ. Obs. Phys. Cétes Orient. [3]: pl 91.6: 255 25, (fig 2}- 1725. Erims laciniatus L., Sp. Pl., ed. 1, 2: 630. 1753- 1, 2: 630, in syn. nical Verbena erinoides Lan., Tabl. Ency¢l. 35 oepp o: 1947}. Verbena multi tiga Rufz & Pav., Fl. Peruv. & Chil. 1: 21, Pl. 33c. 1798. Erinus (laciniatus), foliis laciniatis L. apué 196), Moldenke, Monograph of Verbena 389 Poir. in Lam., Encycl. Méth. Bot. 8: 547, in syn. 1808. Lychnidea verbenae tenuifoliae folio, vulgé sandia-laguen Feuill. apud Poir. in » Encycl. Méth. Bot. 8: 547, in syn. 1808. Verbena (multi- fida), g Ai aN floribus capitato-umbellatis; foliis semitrif- idis, iaciniis bi trifidis Rufz & Pav. apud Poir . in Lan. Z m., Encycl. uéth. Bot. 8: | 547, in syn. 1808. Verbena spicis eoibiaaide, corollarum laciniis emarginatis; foliis foliis laciniatis, gubsessilibus Lam. apud Poir., Encycl. Méth. Bot. 8: 547, in syn. 1808. Verbena selloi Spreng. in L., Syst. Veg., ed. 16, 2: 750. 1825. Verbena odorata Meyen, Reise 1: 480—-),81. 183 [not V. odorata Desf., 1821, nor L'uér., 191]. Lychnidea verbenae tenuifoliae folio, vulgs Sandia Laguen Feuill. apud Lindl. in Edwards, Bot. Reg., 21: pl. 1 1766, in syn. 1835. Verbena erinoides L. ex Walp., Nova Act. Acad. Nat. Caes. Leopold.—Carol. Cur. 19: Suppl. 1: 379. 1843 ne Zz worthia selloi (Spreng.) Walp., Repert. Bot. Syst. k: 13. “13. 1645. lychnidea Verb: Verbenae tenuifoliae folia Feuill. apud Walp., Repert. Bot. Syst. : 31, in syn. 1045. Shuttlworthia selloi Walp. apud erinoides C. 189. Verbena rbena erinofdes PE prostrata C. Gay, Hist. Fis. Chile Bot. 5: 10. 18h9. Lychnidea verbenae tenuifoliae folio, vulgo Sanda- 937. 1865, Verbena multifida Hort. ex Regel, Gartenfl. 28: 372-- 373, in syn. 1579. Verbena erinoides var. multifida Rufz & Pav. ex C. Ludwig, Wien. Ill. Gart.-Zeit. 20: lh3. 1595. Shuttleworth- ia selloi Walp. ex Jacks. in Hook. f. & Jacks., Ind, Kew. 2: 3} S, in ‘syn. 1895. Verbena pulcherrima Haage & Schmidt, Cat. Général 1898: 93, 1898, Verbena geraniifolia Haage & Schmidt, Cat. Génér- al 1898: 93, in syn. gyn. 1898. Verbena crinoides Lam. ex Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, 4: 1059, sphalm. 190. Schulltievorthia selloi Walp. e x Briq-, Ann. Conserv. & yulgo denke, Résumé 315, in syn. 1959. Verbena erinoides var. laciniata (L.) Brig. ex Moldenke, Résumé 36h, in syn. 1959. iychnidea, dea, Ver- 390 PHYTOLOGIA Vol. 9, no. 6 benae tenuifoliae folio, vulgd Sandia-Laguen Feuill., ex Moldenke, Résumé Suppl. 2: 9, in syn. m. 1960. Bibliography: Fevill., Journ ag Phys. CStes Orient. [3]: 35- 36, pl. 25, (fig. 2). 1725. Le, Spe Pl., ed. 1, 2: 630. 17535 Lames Tabl. Encycl. Méth. Bot. [Tilustr. sie 57. 1791; "Rutz & Pave, Poir : Enc usin Bet. 8: ited 1808; Mirbel, Ann. Mus. Hist. Nat Paris 15: 3 Lindl. in 1766, in textu. 1835; G. Don in Loud., Hort. Brit. Suppl. | gekig and V. fasciculata Benth., species that have nothing whatever t do with this complex. Hayek (1908) says "Verbena ¢ ii Lame, Tll., I. p. 17 (1791), ist wahrscheinlich identisch mit Verben® Berterii (Meisner) Schauer, da ja der Autor die Pflanze aus agg anfthrt. Es scheint daher qveckaites’ got die sttdbrasilianisc Pflanze as Verbena Selloi Spreng. zu beseiciaen,” coment (ae) — il sivent yee duplo con tele pa: corollis minoribus." Sanzin (1919) claims that V. mendocina Phil. is conspecific with what he calls V. erinoides. He states that its characters, as seen by him on Ma’ oides and his V. erinoides var. now Ts dissecta f, glandulifera (Sanzin) Moldenke]. It seems probable that Sanzin's V. erinoides is actually y. dissecta. Neal (1948) reduces poo Regel (1879) places V. multifida Hort. (as well a oe bind Hort 1964 Moldenke, Monograph of Verbena 395 and V, tenera Spreng.) in the synonymy of V. pulchella. Rufz & Pavon's V. multifida seems to be ,nothing more than a new name for Erinus laciniatus L., which they cite as a synonym. They also cite the common name "sandia-laguen", but the specimen which they ree oe came from Concepcion, Chile. The "Index flowers, growing in to the sun on a mountainside. Pips Brighet (190) has the following ge say: "On a souvent confon~ vec les V. tenuisecta, dissecta et tenera une espece voisine, Sacrer les notes ss Bees Gi St..s--Cette espéce a 6té basée par Lin- né sur une plante décrite sous le nom de Lychnidea, etc. dans le sree de Observations cba ete. de Feuillée (III, a8 or: a et figurée sur la planc pat: fig. 1. Au sujet de la pa & cette plante, e iinekour oe 'Je trouvai cette plante on ee > nes du royaume de Chily, Ki 38 degres 28 minutes de hauteur ~ Péle austral.' La figure représente trés exactement le es ort une verveine 4 corolle de coleur incarnat assez omni F ae et qui differe totalement des V. tenuisecta et V- es feuilles cuneiformes 4 la base, pinnatilobées, "ay soonents un pet tad plus larges, Feuillée dit la tige 'parsemée d'un cha d'un vert meee pep ae Cette tige + rend sa cou. be en effet complétement hérissée de poils srt en plus enses encore que dans le V. dissecta, tandis ie “Te tenui- hum wuleux, Secta et tenera ont un indument apprime. Le santos est a angles 5 aillants entiérement hérissé de poils étalés mélangés nayh des glandes stipitées plus ou moins abondan ndantes. Les dents lane sont courtes et tri eolées. Nous ne connaissons cette espéce — impossible a a con- transandines on= fondre avec gelles des plaine lorsqu'on lta c Prise - que di (nombreuses prouvenances h. Delessert) et (Bonpland in h. DC.). Elle est , il est ; pe nel e Bresil meridional en général, et en anes s ben aa ms ne sont pas dues a variant num s t Bri t tems rom 4 quet's seer . a ve laciniata is fig- of the sketches Feuillée's work — ure 2 on plate ne - peravians (which Briquet's description ieties of V. erinoides, but oak tact (a8ks) xt, rte “five var to give them nomenclatural designation: lanceolatis inciso-serratis vix trifidis; p Foliis Losier andl idis subtripartitisve, laciniis incisis, | ultimis ovatis “avy ex pl. Chil. no. 159%]5 LEED dao - ” ultimis lin Regist. t- 396 PHYTOLOGIA Vol. 9, no. 6 1766. — V. Sabini Hort. — Sweet, Brit. flow. gard. (2 Ser.) IV. t. 3h7;3 SFO: Foliis iis bipinnatifidis, segmentis paucis linearibus elon- gatis vel etiam foliorum segmentis anguste linearibus; ¢ Foliis bipinnatifidis, segmentis linearibus brevibus, bracteis plerumme calce longioribus. -- An specie diversa? -- Crescit in Chili, Per- a idionali." His third variety is discussed by me under V. laciniata var. contracta (Lindl.) Moldenke. The Mexican plants to which he cai are certainly not conspecific we the rest of the material are probably V. teucriifolia Mart. Briquet (1904) says "Schauer a od dubitativesent au Ve laciniata (V. erinoides) deux plantes qui en sont probablement différentes: les V. Selloi Spreng. (Schuttlevorthia Sellot Selloi Walp.) et V. Sabini Sweet. -- Le | V. Selloi est décrit par Sprengel (Syst- veg. II, 750) comme ayant des tiges hispides, des fascicules flor=- pédonc 6 hirsutes et des segments foliaires lin 3 es trés entiers. Ces indications, en particulier les dernicres, ne cadrent le Vv. iniata: elles s'appliqueraien au V. dissecta. Mais Walpers attribue a la plante de Sellow des corolles deux fois plus courtes que le calice (op. cit. a cage 6 : dans de 1 caractére, qui ne se retrouve aucune autre espéce tion, acheve de rendre le V. Selloi entiérement obscur: cine = p Stre 6lucidé que par 1'examen des Gay (1849) lists two eties of his V. erinoides, — he distinguishes as follows: "x erecta —- s plus minus ‘a atis; foliis confertis.....f pro rostrata —- ramis eae us, debilibus; foliis remotis, superioribus tripartitiss spicis — ternatis.” The first of these is probably what is now called Ve laciniata var. contracta (Lindl.) Moldenke. Vi laciniata has been found by collectors on the interany dine highlands and sandy fields, on open hillsides and —— in full exposure to the sun on mountainsides, on sandy plains ty * grassy northwestern slopes, in sandy es and stony © places, on dry pam aaa open rocky pl - on are and gravelly ground, in s sandy 2 dk volcanic soil, on dry or dry desert slope, and in the Opuntia-Baccharis zone, at nititades of 1165 to to 14350 rui ve e and ting in January, April to June, August, September, and November. Common and vernacular names reported for it are Routleaf verbena", "Et inus-like vervain", "mamitac ttoccaynin", "margarita norada", "ye verbena" is the name recommended by "Stan~ ents ae Pie pon att Herbarium chem ‘has been abundantly misidentified and dis- 1964 Moldenke, Monograph of Verbena 397 tributed under such names as V. aubletia L., V. berterii (Meisn.) Schau., V. chamaedryfolia x erinoides Osten, v. crinoides Lam., V. diffusa Willd., V. dissecta Willd., "V. mr. . elegans", ve ans cides Hook., V, erinoides L., V. inflata H.B.K. [by I. M. John On the ot other hand, the Pat NG hep Bot. Berol. sen. [180] and Lorentz 1053, distributed as V. laciniata or one of its synonyms, are ac are actually ve. ee S. Moore; memes Son. (8.1x.1895], s.n., and R. A. Philippi 1298 are re V. berterii (Meisn.) Schau.; Ga Galeotti 736 “$56 is V. ciliata bentley Yona Venturi 4h, 1930, 2303, 2838, 3579, & 526) are V. dissecta Will ; Hieronymus 8.n. (hex. 1876} is V. glandulifera Moldenke; a 2698, 4063, L607, & 8356 are a. lilloana Moldenke; Cook & & Gilbert Bhg and and F. L. Her- rera 8.Nn. , (Cuzco, July 1923] are V. microphylla a H.BKe} Safford S.n, (Independencia, Oct. 2h, 1886] is V. pulchella Sweet; — ee ceeeenenee! | caenmee amine ~abnnenianen 19868, ee » Britton, & Boynton ton 8296, ck 2h, Castellanos s.n. ere + ee Reartes, Dec. ec. 15, i919], Garnier 615, J. G. Gilmore 1792, Goode 145, Hartling s.n. [N. Y- < eee Gard. Cult. Pl. 42722], Herb. Meisner s.n, [20 Jul. 1637], Herb- oasae 13675, 15290, & 20hi5, Herter son. n. [Paysandé, Sept.--Oct 1907], JOrgen 2470, Jtrgensen 2h65, Marcus cus 1962, McCann s.n. (9- 15-36], H. Ne Moldenke 106ha, N.Y. Bot. Gard. Cult. Pl. . L722, Cory - Store & Tr 96h, P. 0. Schal- atthaus Parks & Cory 2273h & 22735, Pastore & = & rremoee lert ; 251 & s.n. [April 2, 1911), » Mosier, & —& Mat L778, E E. C. Stewart 683, ‘tone m9 a uston, ton, June 1926], Teiss- 2 ee ee mmm ? Tiel as and Valeur LLB are V. , tenuisecta Briq.; Venturi 2651 is a Briq.; and Venturi 9h9 & 3677 e. Balls rts that the species makes “loose mats to several Seats across where plants intertwine, growing chi among thin turf and against rocks o cag a slopes Soil." Steyermark desc aes having "stems han banks". Ragonese & pat “Gan eon ta a as a gt Perenne, de flores vielhootay 9 camin ae eee _ i ne os, culti bundant at La Ollada. pe Si pig a "Hierba perenne, con 398 PHYTOLOGIA Vol. 9, no. 6 para purgaciones, Sus hojas menos profundamente partidas y con ve anchos y muy cortos 7 mismo las bracteas la dis- mitad tan largas como el c4liz" and tos apéndices de las anteres mui cortos e inclusos en el tubo." He asks "En V. multifida.-«-« as anteres se han jado sin apéndices, por omisio: W As its distribution, he says "Planta del Peri il, de la Arjen tina i de Chile; aquf desde el Norte hasta la Araucania; tambien en las cordilleras bajas. V. multifida fué descrito de Concep- cion." Pte Ty Haage & Schmidt 2870) describe their V. erinoides — based om their no. 5919 —- as "purpurviolett, reichblthend,” while Park (1963) describes it as "12 inches (talie]", with purple flowers, and offers 100 seeds for 25 cents, 200 for 5 cents. Mattoon (1958) tells us that the species sé offered to the hortic icultural trade by Vaughan's (New York & Chicago), Sutton & Sons (Reading), Thompson & a Pearce (Moorestown, New Jersey), and Saier (Dimondale, Mic vas & Schnack wel 95) discuss the length of the pistil in re- lation to the size of the pollen-grains. Peck (1937) offers etn and calls Stile -iersaeks eae in A.DC., prodr. ae ee (18),7) says fe V. teucriifolia Mart. & Gal. is ae to modo accedens, sed corolla magna jam satis distincta.” The pear epithets of V. erinoides a V. selloi are often upper sed, Walpers (185) places this species in his Section Verbenacé, Panties | Subsection Inermes, Group Foliosae, Subgroup Macranthae, and °° te) Subgroup Rubletia with 10 other species. Perry (1933) places it in her list of "Doubtful or Little-Known Species" and says "This species established itself in several places [im the United States]. It belongs to a South American species complex needing critical study to determine accurately its real identity". The plant to which she refers is V. tenuisecta Brige, not V. laciniata. Tae ee The Hicken, ees ome - 65 reference given pin is often dated "1922", but as far as I am able to determine is more correctly dated 1923. The Loud., Hort. Brit. Suppl. 2: “680 (187) * poe ence cited above is often credited to W. Baxter, 196 Moldenke, Monograph of Verbena 399 erence mention several times on prec Schauer (1861) 4 "nFeuill. H Hist. Peruv. Chil." A 4 Linnaeus (1753) as "Few. peruv." The original publication for Erinus lac- iniatus is ccaatsaees given as page "879" in error, while Te Poiret gives it as "Sp. Pl. as 177". The Bot. Reg. 21 reference is some~ times misdated "18 The Weydahl 160 pee cited below, may have come from either Cotopaxi or Pichincha, Ecuador, as the locality of collec- tion lies directly on the border between these two provinces. Osten identified his no. 3523 as Ve chamaedryfolia x erinoides mms that he found it growing among these two species. Hi error. The F = Journ. Obs. Phys. Cotes wesc (1725) ref- s is cited by - Org patie an Mera Hers icult. Assoc. Kua yumpur (9k0)" v. ghee hor is said to be pag ete in Malaya. Troncoso de V. tenera SPR., especie muy afin a la ied: y qe dado lo sintético de las diagnosis originales me ha — per sible dif- erenciarlas, Problema que me queda por resolve Hegi (1927) says of the plant to sri he ee be the name V. erinoides: "soll die Menstrationen ftrdern." — Frag fairy niien collections of Martius, Riedel, and Sel- low from , Brazil, and Bonpland and Meyen collections Grande do Sul, Riedel from S&o pope and Martius from par gh vice (18),3) cites Herb. denow (Lindl.) Moldenke. Hayek (1908) cites Wacket and Schiffner collections from S#o Paulo; Herrera (igla) cites his nos. 18 and 342 and Hicken s.n. [1903 3] fr Raimondi (I9L3) cites his 10703 from Arequipa, 10712 and 10751 from An- Pi 7373 from Cajamarca, an and 1069 Province termined, Peru; Cabrera (1915) cites his no. hss Augusto (196) cites a Sellow collection fram Perto de S. |, Indcio, Rio Grande do aL, Herter s.n. from pera and Augusto & & Edésio s.n. from Bananeiras. Ragonese & Covas (1947) cite y okt Besete™ atti, Covas, & Rago- — 3795. Lorentz & padaninin (1881) cite a sical lesteaste te o Negro entina, whic spec Vv. berterit (uetane) Schau. rey: apply ‘0, (1951) cites the fol specimens: Santiago: Collector undesignated s. R. [x-1882} prenaaa 2 2 Sg—68h0L) 5 Looser 4222 (Sg—67501)5 Re A. P. gan. [X1.1855] (Sg—5l760)- Llay-llay: Collector = an if 3) (s Sg—68L05) . chy ES Collector un ota] (Se—5u75u) « z Enis Sceignated $.n. designated s.n. (Sg—6806); Germain Sone 4,00 PHYTOLOGIA Vol. 9, no. 6 Antofagasta: Collector undesignated s.n. [Paposo] (Sg—l2h76) Santiago: Collector undesignated s.n. , [Salto de Conchali, 1976] (Sg—5h737) Covas & Schnack (19));) ares V. laciniata from some of its closest relatives as follows e Stems erect or suberect, not creepi 2 eb ph plant covered by a dense pubescence 0 2 mixed simple glanduliferous hairs; leaves tripartit coreeeisger 2a. Entire plant covered by a sparse yee a: ° of simple hairs; leaves pinnatisect.......--++-V« mendocins. + Stems creeping, rooting at the base and iscesae gz at the tips. ny Mericarps 2 mm. long h. rete one-third as long as the calyx; spikes med —_ ating after anthesis ivbeske = ee Bractlets more than one-half as long as the oa e elongating after anthesis.....ssssoceesesVe anaes Meri Ng « ub 5. Corolla glabrous outside; glandular anther-appendages $ ylindric, scarcely visible from the outside or included; pubescence composed of hairs appressed to the epidermis «++ Ve laciniata. Sa. Corolla pubescent outside; glandular anther-a ad ¢ much compressed, plainly exserted; pubescence composed 0 erect or oblique hairs coeeeVe parodii- Rttmpler (1873) says of V. laciniata: "Brasilien. -- - Einjenrig : — Steifhaarige, stark verzweigte Pflanze mit liegenden, sp#ter am fiedert, mit linienftrmigen, spitzen, ganzrandigen oder kaum o zzhnten. Lappen. Blumen violettroth, in wenig nga 9 we e execheinen von Juni bis Oktober in reicher Folge. Sie ist hawt s¥chlich zur Bildung kleiner Gruppen und zur Ausstattung der Rabatten geeignet,” In all, 97 herbarium specimens, aniiatinn nue types of same of the names involved, have been examine - Citations: ECUADOR: Azuay: Steyermark 53129 729 9 (i) « Chimboraso? Asplund 6862 (Lu, S, W—-1930933), 2063 (S); S); Fagerlind & Wibom 799 (Iu, 8), 837 (S); Hartweg 1351 (Lu, N); Mille LO (Ny — Tia, . ); Penland & Summers 1,66 (N); Rimbach 175 ). erarracn ge , 423 (Mi, N, S)3 Rose & Rose 22312 = w—1021979) « Weydahl 160 (s). Pichinchs: a ce (Ge— SeM86T, s, Fe hh Tunguragua: Asplund at eT (3)3 ot & Rahn 97 (S); A. S. esteneock 21 Ze (N, W—11 E.G. Lehmann 1i2 (Bm, W—13231)6); Pachano 155 no 155 (W—-10LL636) 5 196 Moldenke, Monograph of Verbena 401 Steyermark 51853 (N, S, W--1901701). Province undetermined: André 3609 [Alto de Guaillabamba] (N); Spruce 5065 [In Andibus Ecuador- ensibus] (Lu, N). PERU: Ancash: Ferreyra . 7333 (Ss). Arequipa: El- lenberg 182 eateay rib Ayacucho: Ferreyra 7216 (W—2028320) . Huancavelica: Rauh & Hirsch P.375 (Z). Junin: Macbride & Feather- stone 1260 (W— sinker R. W. Pearce sn. (Palca] (pm). La Liber- tad: Lépez Miranda 15hh (S); Je West 8115 (Ca--565221). Lima: As- plund 11322 (Gg—03668, S, W—222l08); J. Ball s.n. (Chicla, Aprili 21, 1882] (Cc, #1323080) ; Diers 962 (Ko); Ferreyra 6502 (N); Killip & Smith 21619 (N, W—1356700), 2175b (N, S, N, S, W--1356893); Macbride & Featherstone 678 (N, N, W-—1830560). Puno: P. Aguilar F.2 (0k). Department undetermined: Mathews 1018 {Peruvia interi- ore] (Bm, M). BRAZIL: Rio Grande do Sul: A. R. Schultz 701 (N)- State undetermined: Sellow s.n. [Brasilia] (Br). , BOLIVIA: La Paz: suenicn 758 (V). Oruro: Hammarlund 123 (N, S); Kuntze Sone {Oru- 3/92] (W--702211). Province undetermined: Julio 133 {La obs (W--1192719). URUGUAY: Osten 3235 (Ug), 3 283 523 (Ue), 8723 els 8726 (Ug). CHILE: Bfo-Bio: Pfister 2177 (N). Ci ion: Barros Valenzuela 876 (N)3 Junge 971 [Looser ser 1,030] , 1805 ae O'Higgins: F, W. Pennell 12237 (N). Santiago: Alcibiad Santa Cruz s.n. {circa mons Caracol, X.193h] (Ba); Looser 7038 (N). Talca: Alcibiades Santa Cruz s.n. [1938] (Ba)- Valparaiso: Garaventa 2672 [Looser ioe] (N); Gillies sen. (1); Looser 4032 (N). Province undetermined: Dusén s.n 3.n. [ad Lotam urbem, em, 10.X. 1896] (Lu); C. Gay 792 ae Herb. cae sn. [Talcaraque] (C); Herb. Mus. Nac. Santiago 1 [Condes] (N)- ; ARGENTINA: Jujuy: Balls 598k (W—177777h) . Mendoza: Rufz Leal 13612 bis (Rl); Sanzin 8. n. ae (Ruts Leal 1413] (N). Salta: T. Meyer 12088 (Gg——-353269, 3069, NJ : Schreiter s.n. [Cumbre Alta; Herb. b. Osten 72196] (Ug)- CULTIVATED: New York: R. H. Ward s.n. (Troy, July 1868] (Ur). ae OF COLLECTION UNDETERMINED: Collector undesignated 10 ‘A var. CONTRACTA (Lindl.) Moldenke, Phytologia Synonymy: Verbena multifida var. contracta Lindl — — Bot. Reg. 21: pl. 1766. 1835. Verbena erinoides Willd. apu s : : ~ IS. 2: 750. Loe [not Ve erinoides Auth., 1959, nor Hook., 193, nor Hook. = Arn., 1959, nor oe 1959, que Lam ee wea no r Poepp., 13k7, r Spreng, 1830]. bena Wieiskstins Willd, ex Lindl. gee te Bot. Reg. 21: i 1766, in textu. 1835. Verbena mult aatida PB contrac rants Lindl. ex Sweet, Brit. Fl. Gard. 7 Ler: 2, bj: pl. 347. 18 . Verbena er- incides B® contracta Schau. in A. DG, prodr. 11: 3. 1647. Ver- bee. 9 erinoides & erecta C. Gay, “wist. Fis. Chile Bot. 5 ue] d 5 ; i BR “8 402 PHYTOLOGIA Vol. 9, no. 6 Bibliography: Lindl. Bot. Reg. 21: pl. 1766. 1835; Sweet, Brit. Fl. Gard. 7 lear: . “his pl. ate 18 36; Walpe, Rep- Fl. Gard. 1: pl. 48. 1854; Briq., Ann. Conserv ard. Bot. év. 7-8: 297. 190); Stapf, Ind. Lond. 6: 29 & oe: 1931; pies ke, Phytologia 3: 26 & 167 « 1951; age bgt Rak Abstr 3081. 1951; 3 Moldenke in Chittenden, Roy. Hort. Soc. Dict. G kh: 2211. 1951; Scars, Résumé 223, 36h, 30, Py 472. 19595 Mol- denke, Résumé Suppl. 3: 1h, 29, & 37 (1962) and k: 5, 6, & 15. 1962. Illustrations: Lindl. in Edwards, Bot. Reg. 21: pl. 1766 [in color]. 1835; Sweet, Ornament. Fl. Gard. 1: pl. 48 tin color). 1 This variety differs from the typical form of the species in it more compact growth, the stems more or less abbreviated, and the leaves crowded. Schauer (18h7) describes the variety as follows: "foliis tri- itis, laciniis inciso-pinnatifidis, ultimis Linsari—oblong'# acutis" but includes V. sabini arrte on its synonymy. It i doubtful whether the leaf charac which he gives really 4 mes to this variety. Briquet oaks 6 she "Quant au V. multifida var- contracta Lindl, ...... dont Schauer a fait son v. erinoides Pp contracta, c'est évidemment, d'aprés la figure, une forme forme alpine réduite du V. laciniata.”" The Lindley, Bot. Reg. reference giv- above is often dated "1836", but apparently should be 1835+ Gay (1849) describes this plan ant "ramis plus minus abb bbreviatis; foliis confertis", Lindley affirms that it is native to the "Alps of Chile and Mendoza", and calls it "Dwarf purple vervain! esto gy sei var. SABINI (Sweet) Moldenke, Phytologia 3: 9 Synonymy: Verbena erinoides var. sabini Sweet, Brit. Fl. Gard. 7 (ser. 2, h]: pl. 347. 1836. Verbena sabini Hort. ex Sweet, Brit. Fl. Gard. 7 eg 2, 4]: pl. 347, in syn. 18%. Verbena multifida sabini D. Don ex G. Don in Loud., Hort. Brit. Suppl. 2: 680. 1839. ae sabini Sweet ex Schau. in A. Des Prodr. 11: 553, in syn. 1847. Verbena sabiniana Hort. ex Brides Ann. Conserv. & Jard. Bot. Genav. 7-8: 297. 190). Verbena erin Qides sabinii D. Don ex Stapf, Ind. Lond. 6: 29. 1931- Verbena laciniata var. sebintt Moldenke in Chittenden, Roy. Hort. 50Ce paniples rnetind Brit. Fl. ¢ 7 (eer~ 2 2, bjs pl. 347+ 1836; Walp., Repert. Bot. Syst. hj: -. "1845; S in Ae Prodr. li: 553. 1847; Regel, Gartenfl. 28: ots “38793 Hook. re * Lond. 6 19315 we A ste mee fete 3: 426 & Lb7~ 195k woldenke, Biol. str. = « 1951: Moldenke,. Ré & 22 373 19595 Moldenke Risk 8 Suppl. 3: 29. 37, 33° 0, *e Ll. 1962- 196) Moldenke, Monograph of Verbena 403 oh gerare Sweet, Brit. Fl. Gard. 7 [ser. 2, k]: pl. 37 Hes fe. 1836 his variety differs from the typical form of the species in b e dwarf in habit, denser in growth, more glabrous, and with dee terole corollas Briquet (190) sata WLe V. Sabini Sweet.....est représenté dans nos collections par des 3 échantillons cultivés sous ce nom au ap ge de Paris (ann. 1836, Verbena Sabiniana): ils appartien~ nent 4 une forme presque identique au V. = pulchella Sweet et peu- vent rte Os ertande an V. tenera a Spreng." The original publica~ tion by Sweet is often dated 1838", but appears to be more cor rectly dated 1836. The "Hort. Brit. Suppl." reference listed a- bove is sometimes credited to W. Baxter, apparently in error. var. contracta Lindl." Regel (1879) places V. sabini Hort. as Synonym under his V. pulchella "Spreng.", where he also Tay ¥. mitifida Hort. and Vv. tenera eee VERBENA LANDBECKI R. A. Phil., Linnaea 33: 194. 186k. : Verbena lamdbecki R. A. Phil. ex Acevedo de Vargas, Synonymy: Verbena lamdbecki Bol. Mus. Nac. Hist. Nat. Chile 25: 57, sp Bibliography: R L * 19h. 1861; Hook. f. & acks., Ind, Kew. 2: 1179. 1895; Reiche, Fl. Chile 5 287. 1910; Moldenke, eogr. Dis + -, Led. 1. 19h2; H. Ne & 1d e 2: 68. 1948; Molden- €, Alph. List Cit. 3: 687 & 813. 1949; Moldenke, . Distrib. Ver" , [ede 2], 101 & 198. 19h9; Ace meg ag Bol, Mus. Nac. Hist. Nat. Chile Bet 56--57. et Moldenke, 2 gente & 172. 1959; Moldenke, Résumé Suppl. 3: Us (1962) and P wists plant, about 25 cm. tall, grayish-hispid, mec Stemmed; leaves d ite, sessile, more = less semi- 5 s decussate-oppos °, teal twigs in their axils, the segnents linear, “obtuse s the apex, revolute along margins; spikes terminal, $ rere aes red; bractlets awl-shaped, 5 tly shorter than flowe gh calyx; calyx about 8 mm. long, its rim with 5 short triangular teeth; corolla lilac or blue, its tube about 1 cm- limb 8 mm, wide, the 5 lobes cuneate te and emargina The type of this very rare species was collected by gra Ludwig Landbeck near erage Acancagua [Coquimbo, ac smmkay fs - Acevedo de Vargas], Chile, in January, 1863, md is Gspos Too” the herbarium of the Museo Nacional de Historia Natural ago, Chile. The species has been found at t alti = 1350 meters, in flower in January and October. 2 herbarium specimens and 3 phototypes Have been Reiche (1910) says of it "mui Berterii." Acevedo de Vargas (1951) cites the following roeslean - Specimens: Coquimbo: Landbeck $-Me (1.1863) (sg—5h672—tyPe)» Ss Lok PHYTOLOGIA Vol. 9, no. 6 n. (Concumen, 1863] (Sg--54736, Sg—68372). Field Mus. 17425 (Sg). She notes that "La foto del Herb. Berlin, distribuida | par ar Field Mus. Nat. Hist. 17422, no es V. lam dbecki. En nuestro herbario museal ha sido confundida con V, laciniata." The Macbride photo- graph to which she refers actually is of a cotype of V. interced- ens Brig. Citations: CHILE: Aconcagua: Landbeck s.n. [Choapa; Macbride photos 1725; Herb, Mus. Nac. Hist. Nat. Chile 5672] aaa of isotype, N—-photo of type, N--photo of isotype). Biese 1875 (N). Province undetermined: Claude-Joseph 18s. (Bureo) (W—-13]2397) . Mo tess cera: her' Se Link, Enum, Hort. Berol. 2: 122. 1822. Synonymy: na prostrata R. Br. in Ait., Hort. Kew., ed. 2, hk: 41. 1812 er V. prostrata Benth., 1959, nor G. Savi, 1802]. Verbena rostrata Felt, Bull. N. Y. State Mus. 200: 181, sphalm. 1917. Verbena Tasiostachya Link ex Moldenke, Alph. List + Invalid Names Suppl. 1: 25, in syn. 197. Verbena prostrata Ait. ex Mol- denke, Alph. List Invalid Names Suppl. 1: 26, in syn. 197. Ver- bena Lambert’ Spreng. ex Moldenke, Résumé 368, in syn. 1959 Tnot Lem iamberti Ker, 1827, nor Penny, 1832, nor Sims, 1820]. Verbena lasiostachys var. prostrata (R. Br.) Wall ex Moldenke, Résumé 368, in syn. 1959. Verbena leptostachya Winblad ex Moldenke, Ré- suné Suppl. 3: 39, in syn. 1962. Verbena prostrata Greene ex Moldenke, Résumé Suppl. 3: 0, in syn. 1962. Verbena prostrata A. Br. ex Moldenke, Résumé Suppl. 6: 11, in syn. 1963. Verbena iasiostachys var. lasiostachys [Raven], Aliso 5: 336. 1963. 827; Sweet, Hort. Brit. 418. 1830 & Arn., Bote Beech. Voy. 256 un 303 (1838), and ish. ial; “Walp, Repert> Bot. Syst. hs 53 Schau. in A. DC., Prodr. ll: 547. “Am 933 B am. l, (3a): 148. 18943 Hook. f. & Jacks., ew. 23 1179. 1895; Richter, Calif. Agr. Exp. Sta. Bull. bi7e 583 & 1017. 1911; Felt, Journ. N. Y. Ent. Soc. 20: 150-151. 1912; M. Amm- strong, Field Book Test. Wild Fls. 435. 1915; Felt, Bull. N. Y« State Mus. 200: 181. 1917; Smiley, Monthly Bull. Dep. Agric ailey, List Flor : erbenac. mss. 1935; Munz, Man. So. Calif. Bot. gh 19353 196), Moldenke, Monograph of Verbena 405 Dermen, Cytologia 7: er 1936; Perry, Rhodora 38: 272. 19363 oldenke, Prelim. Alph. List Invalid Names 48. 1940; Worsdell, 3 sc . iis Yotderke, Alph, List Invalid Names 9. 1942; Moldenke, mn Geogr. Distrib. Verbenac., [ed. 1], 2, 15, 18, 7h, & 101. i Jepson, Fl. Calif. 3 (2): 381--362, fig. et 194.3; Bower- » Flow, Pl. Ferns Mt. Diablo 72. 1943 San Dimas Exper. Forest anak om: some Si 270. 19hS Moldenke, Alph. List “cht rap Es 6 Be, i 9 ee eo 135, sate Sh, A 18. 191, i, 191, 20 a’ » 11, 120, 1H 1305 150s 756, 761, 7th, 779, 785, phe 800, 803, 816, 825, 831, 8 3. 850, His 871, 918; hat 953, Re & 976—978 (89), ant bs Son Bh 989, lok, 114: 1138, ye Ay. 3 oy ’ ry , 117, 176. 1197, iat 1216, 1217, tr ae 1230, 1232, 12h3--12h5, 12h7, 12)8, 1252, 1253, 1288, 1290, & 1292 sig; H. Ne & A. Le Moldenke, Anal. Inst. Biol. Mex. 20: 1h. 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2], 3, h, 27, 33, Pi & "198. sg 4 Moldenke in Chittenden, Roy. Hort. Soc. Diet, & . hz 2209 & 2211. 1951; Moldenke, Phytologia 3: a New Bri Gleason, New Britton & Br. Illustr. Fl. Pa in, & 132, 1958; Mattoon, Pl. Buyers Guide as gs “287. 19585 * yolden- e, Am. Midl. Nat. 59: 353, 35h, & 361. 1988; *yoldenke, Résumé 5, 37 résumé g Je, 33 Suppl. 2: 13 (1960), 3: 8, 9, 29, 3%, & hO (1962), 6: 11 (1963), wt 7: 9. 1963; Gleason & Cronquist, Man. Vasc. Pl. 561. 1963; Ven, Aliso 5: 299, 302, & 336. 1903. Illustrations: prichenb., Hort. Bot. pl. 6) [in color]. os Reichenb., Iconogr. Bot. Exot. 1: pl. 64 [in color]. 1627; M C Dep. Agric. Calif ci Me ete fig. 807. "1925; Junell, Symb. Bot. 19 . So. Calit. Bot. 437. 19353 Jepson, Fl calif. 3 are ‘fig. 20. 20. 19h3i e in Gleason, New Britton & Br. Ill- Pi Bt Le 32 (1952) and nd print. 2, 2, 3 132. 1958. Baer alg sparsely Vi r vill eon ennial, strongly pungent, eae es to 1.3! Re 1,06 PHYTOLOGIA Vol. 9, no. 6 wide-spreading, more usually semi-decumbent or prostrate, much branched from the bares stems at first erect or ascending, usual- Spars S}3 ae es decussate-opposite, mos brunnescent drying; leaf-blades oval broa o oblong-ovate 0 oblong, 4--10 cm. long, obtuse or acute at the apex, cuneately abteanats at the base into the margined petiole, coarsely and im te-dentate with apiculate teeth, incised, or lac- iniately lobed, the lower one ae -divided, the venation appressed above, r less prominent beneath, both surfaces mostly soft~puboscent. 0 or Pilose to pilose-strigose or ous, somewhat ru e above, scabrous; spikes t minal, solitary or more Re loosely paniculate, dense before anthesis, becoming much elongat ad open in 5-3 cm. flowers and fruits mostly imbricate; bractlets lanceolate- late, 1/3 to 1/2 as long as the c calyx, villous and pa 08 ely c the limb 3--5 mm. wide; cocci oblong-triquetrous with a convex back, 2—-2.5 mm. long, slightly thicker at the distal en raised-reticulate above, striate below, often oe the striations fading out toward the base, the commissural face more or less woesagiied, ges amagige A reaching the tip of the mitiets chromo- e mumber: = 7, Qn = ae “grade - and rather pe lexing western species collected in dry fo Pans A we hills, brushy hills pnt anil sides, and steep or dry hillsides, in ret places as well as ound, on flats along riverbanks, in 2--3 inches of soar near tide line, and washes, at the e of foot- hills and at the edge of woods, in te places and moist ravines, on flats and flats, in seeps at the base o ean cliffs, near creek beds, in woods and pine woods, on damp sunny south and stream aks, in low areas near rivers, on creek bottoms and gravel bars, along ‘rivers especially near the ocean, in stony river flood beds, in marshes and an inhabitant of the Upper Sonoran Zone, usually in dry ope? hill country, from sea-level to altitudes of 2995 meters. It has been collected in anthesis from March to October and in pe sgntecl and in fruit from May to October and in December. Johnston repe it as "common" in wet places in the San antonio Mountains; pepe i Says "plants occurring spontaneously in artificial streams in mois heavy soil with Mimulus cardinalis Dougl." 196), Moldenke, Monograph of Verbena 07 Ewan states that this species is frequent in the moist soil of backwater channels in Los Angeles County and in dry grassy clear- ings in Baccharetum with Navarretia and Gnaphalium in Contra Cos- ta County, California. Harrington calls it "a bush about 3 feet e Du barium has fasciated spikes. The Ogden s.n. [Ventura, ried & Aug. 2 and L. S. Rose 39140 collections have ave the calyx slightly r is usual for the typical form of this sane s and nay possibly represent var. septentrionalis Moldenke. Draper 25, distributed as this se So ieast in the Britton Herbarium), is actually Marrubium vulgare L. in the Lamiaceae. BOOK REVIEWS Alma L. Moldenke "Wild Flowers of America", edited by H. W. Rickett, 471 pp., 400. color plates. Crown Publishers, New ror city; 3rd printing. 1963. $15. This book is a wonderful means of making the famous M. V- onian In- family of the SCient. the names Pig ovr ramiggdacts sc i. t wal have been added. 408 PHYTOLOGIA Vol. 9, no. 6 "Tree-Sort Guide", by J. E. Forester, 155 pp. or cards. Crown Publishers, New York City; illustr. 1963. $4.95 n today's world of I B M machines and push buttons these e lished excellent work of the U. S. Forest Service. The most- ly dichotomous key leads to easy and Bbiece® es ge The cards are protected with a good plastic case "Look at a Flower", by A. 0. T. Dowden, 120 pp., illustr. Thomas A, Crowell Company, ea York City. 1963. $1.50 This charmingly and ti oosguen prapege tis book is the work of 4 fted botanical illustrator and has older juvenile or ia amateur level. The mv pe sh acs as is vouchsafed care checking before it seg to press by outstanding authorities. ee information on flower structure, function, and classification is very clearly given ga word and pict ure. "The fapste-s Seite to she American Trees", by C. C. Curtis and e504 3h2 illustr. Collier Boake teat New York me "1363. ;: 98 paper back. The immediate ancestor of this pocket-sized paper back book is the now depleted hard cover larger sized version of 1943; and t one, in turn, is derived from the also depleted but “former y _— used and admired "A Guide to the Trees" of 1925 and from t of the senior author en The or Faces work = Dr. Curtis was limited edaraal to t southern and ek parts of our country, so that now the descriptions of genera and species are clear-cut resting} both scientific common names are given along with the thorities for the former; the on oll OO ne —— are diagnostically accurate and natural in appearan ree Introduced and cultivated trees and borderline arborescen nt $ are not included. It is good to have this fine work, initiated by Dr- curtis again available for carrying in one's pocket on local fie 1d es trips in different areas of our continent, and for the home hool library, PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue Each number consists of not i than 32 pages. All manuscript accepted will be ctidied in the next issue, so that the size of numbers ae vary greatly. A cg will contain sae 32 signatures, 512 pages, or a smaller number of pa with an equivalent number of plates. 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Uaet the pemeat ck of pri ting, the basic rate for a page or fraction ae is approximately $3.00 for an edition of 250 copies. ‘This price is subject =e change without notice, since it — entirely on the prices prevailing ae See oes er sprints will be furnished at cost. A proportionate fraction of the edition — et ee Soe tres aa PHYTOLOGIA Designed to expedite botanical publication Vol. 9 _ February, 1964 No. 7 CONTENTS WURDACK, J. J., Certamen Melastomataceis VIII ....--+--++++++++7* 409 HARKNESS, B., A checklist of the cultivated woody plants of the Rochester parks. Part | .....+2+e-+e0202°° ee MOLDENKE, H. N., Materials toward a ee of the panes ; Verbena. AV Sa eae Ua eat ad oes Re a we SR a Oe OPE ee aie ae oeeee Published by Harold N. Moldenke md Alon L. Moldenke t Bora of. eee 15 Glenbrook Avenue, s ae EB 1Y 1964 Yonkers 5, New York, u. Ss. 5 Ae S CERTAMEN MELASTOMATACEIS VIII. | John J, Wurdack Dept. of Botany, U.S. National Museum the assistance of a grant from the National Science Museo de Historia Natural — [=] — o J ba wt = © 4 a] nn e @ mm o eon ias see 1962 at the Peruvian road construction scadas de Mayasi; the flora of this moist region aioe the a ad ° = ct a 7) ca | | ° “ D * aa Le | is") a] rh Wer n. the zone, the eats Soeatne from the xerophytic associ iation along the Maranon © Pongo de Rentema is startling and topographically seeming ly Asie chat : Tex . Turner and Mr. Mike Powell at the pease d 44 i S were “able to obtain chromosome ante a wo of the Lao whose anthers were sent to Austin face sage 7 basing were n = ca. 12-16 for Miconia ¢ centcaghora aud. Wurdack 517) and n = 19 for Graffenrieda calyptr a iat oides oP £ tly are the first familial Me ies of South American lastomataceae. For 1506, Dr. Taree reported very small poorly stained chromosomes and for 1 meiotic cells with e ay ees that the tha family cytologically, pre ; an relatively easy like the pens? iat (where he obta chromosome counts on 58 of 117 vials shipped from Ps oe iter MERTENSIOIDES Wurdack, Sp- no A, dependenti R, & P. affinis se “glabra et foliis minoribus sicut folia inflorescentiarum ramique primum i teretes in seats stipuli- cm tenuiter coriacea apice basique acu jugo interiore 0.5-0.7 cm supra basim inserto jugo amen = debili 09 410 PHYTOLOGIA Vol. 9, no. 7 inframarginali nervulis | ag ksh. a Jos aie sinuato- On serrulata de ntibus Shieh ca. mm altis et 2-3 mm inter se distantibus. Pani fis 10 cm longa; Fiave es eats deniantaatel glabri Soe: a 0.8-1.3 cm longis. Hypanthium (ad torum) 2.6-2.8 X 2.5-2.7 mm; calycis tubus 0.5-1 mm apice trun- data vel vix emarginata. Stamina isomorphica; filamenta 4. longa; antherarum thecae 4.5 X 0.8 mm apice dorsaliter sag ig uniporosae, appendice dorsali inflata 5 X (lateraliter) 2 basi 2 ats Stylus 15 X 1-0.4 mm; stigma 0.4 mm latum; ovarium 4- ra oak glabrum apice truncatum; fructus 4-valvatus; semina non Le Type Collection: J. J. Wurdack 1467 (holotype US 2404308; isotype USM; 14 isotypes to be distributed), collected in scrub forest along Rfo Ventilla 1-2 km west of Molinopampa, Prov. 2 400 nt blue at anthesis ; anthers purp le-brown with burnt-orange vaca ehhi bulb” The cific epithet refers to the developmental change in petal ex, rather like that in the floodplain bluebells of eastern U.S.A, A. glandulosa has a rather sparse tomentum of leaf surface veins and veinlets, and inflorescences; the lea blades are 7- = Xx (2.5-) ES fiat While the correct ape ee tion of the names A. dependens d A. glandulosa R. & P still debatable (with Raytarat's s sill Eves’ applications tie same as the Flora Peruviana Vol. 4 plate s), for convenience Eves' Slopes of Puma-urcu above Gnickibeies (Wurdack 657 & 789), where the large pendant panicles (up to 5 dm lo ny sited the flowering plants somewhat the aspect of tree wisteria AXINAEA N Wu ik hs. F from the jalca zone south of Laguna Pacha ue: Prov. Bongard, Amazonas, is more glabrous than either t the © Species, rath than aving a slight infiltrat of A. gor ong baer eet genes. No pla of A. tomentosa were seen in the immediate vicinity of No. 954. A. nitida was frequent in all undisturbed jalca zones; a close relative is 196k, Wurdack, Certamen Melastomataceis 411 A. lanceolata R. & P. (sensu Eves), with quadrisulcate branch- lets and deeper hypanthia. AXINAEA TOMENTOSA Cogn. Wurdack 1623, from the south side of the Molinopampa-Diosan (isotype US), but has terete to obscurely rounded-quadrate (rather than sulcate-quadrate) branchlets, somewhat longer pubescence, and leaf blades above with veinlets densely raised- e. e na MERTANIA TETRAGONA (Cogn.) Wurdack, comb. nov. Axinaea tetragona Cogn., Bot. Jahrb. 42: 137. 1908. Meriania raimondii Mgf., Notizbl. Bot. Gart. Berlin 13: 1937 Wurdack 1250, from the middle eastern Calla-Calla slopes, has been compared with the type collection of M. raimondii (USM). no doub to the synonymy. Certainly, the species 1s Placed generically beside its near relative, M. radula (Benth.) Bia acbride (also fairly common in upland Amazonas), with which it shares the f re of ornamental red petals. The eat smaller Ovate cordate strongly bullate leaves with glabrous Petioles separate M. tetragona from M. tetraquetra. MERTANIA RUGOSA Mgf. i é Two collections (Wurdack 1483, from the Rio Ventilla scrub forest west of Molinopampa; Wurdack 1617, from scrub forest north Raimondi specimens, are comparable with those of my own flowering Collection, 2.5-3.5 X 1.5-2.5 cm. M. rugosa must be a close 12 PHYTOLOGIA Vol. 9, no. 7 relative of M. tetraquetra Tr., known to me only by description and photograph; the basally nerved leaves and tetraquetrous branchlets, as well as the smaller flowers, are ample distinc- tions from M. rugosa. MER TANIA HEXAMERA Sprague Macbride had tentatively recorded this Colombian tpesiee Pomacocha and Yambrasbamba) seems to confirm this distribution, although the leaves on No. 1050 are only 7-8 cm wide. Cuatrecasas 9139 (US), from Caquetd between Sucre and La Portada, Colombia, in young bud, has also been referred to M. hexamera. GRAFFENRIEDA CALYPTRELLOIDES Wurdack, sp. nov. De affinitate mihi incognita sed certe in syst Cogniauxii a G. patenti et G. weddellii Band: valde welsiiasinl Rami graciles teretes glabri. Petioli 0.7-1 cm longi glabri; lamina membranacea 4-6 X 1-2.5 cm eH de basi apiceque i a vel non articulatis. Hypanthium (ad torum) 1.5 X 2 mm inconspicue 10-costatum; calycis tubus 0.1 mm altus, lobis interioribus 0.3 mm altis late triangularibus apice apiculatis, dentibus exterioribus 0.3-0.5 mm eminentibus acutissimis. Petala X 4.2-4.6 mm oblongo-lanceolata apice anguste acuta. Stamina isomorphica; filamenta 1.7-1.8 mm; antherarum thec 4) Ghachapeyas, Depto. Amazonas, Peru, elev. 2350-2400 es 1962. “Shrub 1-2 m, most ly in bud, frequent. Petals w ite”. Within Graffenrieda, although the aspect of G. 7 Oides is most like that of G, moritziana Tr. 6, a (Griseb.)'Tr., the Peruvian material totally lacks o Galyptrella. € general habit of the Molinopampa endemic, especially the te well-pedicelled flowers, is rather like that 1964 Wurdack, Certamen Melastomataceis 413 of Calyptrella tristis Tr. and C. gracilis Tr. The inflorescence of G. calyptrelloides is also reminiscent of that of G. candelabrum Macbr. That species, however, surely belongs in the Miconieae, probably in Leandra (but not near the vegetatively n = iJ) io) i") n” co ~ < is") = ~_ iO o ~ (9°) | . — ™ on | wa oO i=] =" t —_ i] rh 3) ba | ~te ° | ° . e Rathke! Arnold Arboretum 1915 purpurea Speeth Nur. 1905 p- 'Bley' ex Crittenden Blvd. 1950 "Red Jade! Brooklyn Bot. Gard. 1953 ‘Red Tip! Arnold Arboretum 1919 robusta r. 'Dulcis' Spaeth Nur 1902 re 0 Rochester barks seedling re pereietfolta ° ore 1920 Boerner Bot. Gard. 1960 “gna PO 1d Arboretum 1909 » 'Rosea! B. HE. Slavin sel. 1919 Sie Arnold Arboretum 1902 - arborescens Arnold Arboretum 1918 sikiimensis Arnold Arboretum 1911 spectabilis Arnold Arboretum 1909 s. 'Plena! 196), MALUS, MARSDENIA — Asclepiadaceae Harkness, Cultivated woody plants con sublobate: "Cashmere! Z. 'Bobwhite! Z. 'Calocarpa Z. 'C. Golden Hornet! erecta METASEQUOIA — Pinaceae glyptostroboides MORUS ~ Moraceae albus a. tatarica australis MYRICA — Myricaceae cerifera pensylvanica NEILLIA ~ Rosaceae longiracemosa sinensis thibetica (N.) NEVIUSIA — Rosaceae alabamensis NYSsa — Nyssaceae Ogeche sylvetica ORIXA - Rutaceae japonica OSTRYA — res ee Japoni nein ai XYDENDRUM — Ericaceae PACHISTIMA — Celastraceae Canbyi arboreun — Buxaceae Arboretum Goluchow Arnold Arboretum Votcutt Nur. Arnold Arboretum C. Y. Chiao, Nanking Arnold Arboretum Arnold Arboretum Meehan Nur Arnold ehh Arnold Arbore U. of Wash. yi a Meehan Nur. C. C. Laney coll., Fla. Arnold Arboretum Arnold Arboretum Arnold Arboretum Spaeth Nur. Parsons Nur. Arnold Arboretum Holden scare J. Daws 456 PAEONIA — Ranunculaceae Delavayi suffruticosa 'Banks! PARROTIA — Hemamelidaceae persica PARROCTIOPSIS — Hamamelidaceae Jacquenontiana tricu: t. 'Lows PARTHENOCISSUS -— Vitaceae uspidata PAULOWNIA — Scrophulariaceae tomentosa FETTERIA — oa rament PEELLODENDRON - Rutaceae amurense poem piriforme ie inensis FHILADELPHUS — Saxifragaceae "Belle Etoile! "Bicolore! brachybotrys caucasicus (N.) mee us c. 'Deutziaeflorus! c. 'Duplex! ce. 'Primulaeflorus! PHY ?T0:406:2 a Vol. 9, no. 7 1905 Veitch Mur. 1892 Freiburg 1952 Horton Nur. 1958 Arnold Arboretum 1913 Arnold Arboretum 1916 Arnold arboretum 1912 Arnold ce 1915 Arnold Arbo 1929 Bs meee tee: Topsfield 1904 Wayside Gard. 1949 Lemoine Nur. 1919 Lemoine Nur. 1914 Bautumi 1961 Brown Bros. Nur. 1943 Leroy Nur. 1892 Ellwanger & Barry 1892 Spaeth Nur 1892 Teehiion edits 1892 Lemoine Nur. 1914 Lemoine Nur. 1905 Lemoine Nur. 1908 Arnold Arboretum 1920 Oslo 1957 Arnold Arboretum 1909 Spaeth Nur. 1892 Spaeth Mur. 1892 Hllwanger & Barry 1892 Arnold Arboretum 1915 Arnold Arboretum 1929 Transon Nur. 892 1964 PHILADELPHUS, age L. 'Ere Harkness, Cultivated woody plants 's Avalanche! us! L. ‘options Lewisii L. Gordoni lanus = dient Ti Mt. a nivalis pekinensi tat (N.) Ss pendulifoliu polyanthus . oh pubdescen ih acianae bees macdialee ‘Sylviane! Salicifol satsumanu satsumi Schrenkii Favorit erbe de aiee Blan Ss ‘Atlos® e! lon Blanc! ius s A ong ith ‘Stlver & aia! (N.) - 'Minne ve. 'Pyram: v. 'Slavi PHOTINIA — Rosa Beauvardi vil 8 v ie) ve ese Snowflake! —_ ana = aptawde (N.) wicziana (N.) sinica PHYSOCARPUS — Rosaceae amurensis (N.) bracteatus Lemoine Nur. Lemoine Nur. Wayside Gard. oine Nur. croton ee Lemoin BHS ey pp gr NH. Y. Bot. Gard. Vv. Gibbs Morton Arboretum Arnold Arboretum O. E. White Arboretum Andrews WV of Glacier 1954 1914 1959 1915 1961 1914 458 PEYTOLOGIA oe ~s glab intomedin parvifolius opulifoliu Oy ethene! PICEA — Finaceae Abies A. 'Auresc he Spates Sa (N.) i Ae icrosperna' A A. 'Tiditorets} (Ne) A. 'Frocunbens! (N.) Pia lat A. irene A. 'Viminalis! sperata a. heterolepis a. ponderosa glauca &- albertiana g- 'Coerulea! &- conica Glehnii jezoensis j- hondoensis likiangensis (N.) mariana 'Doumet! Maximowiczii ova Vol. 9, no. 7 Andrews Nur. 1914 Arnold Arboretum 1921 J. Dawson 1892 Brown Bros. Nur. 1943 Ellwanger & Barry 1902 Westbrook Gard. 1923 Mayfair Nur. 1958 Arnold Arboretum 1922 Arnold Arboretum 1905 Parsons Nur. 1905 Parsons Nur. 1905 Barbier Nur. 1911 Barbier Nur. 1911 Mayfair Nur. 1958 Parsons 1905 bier Nur. 1911 Andorra N 1907 Mayfair Nur. 1958 Arnold Arboretum 1906 Veitch Nur. 1905 Arnold Arboretum 1908 Arnold Arboretum 1915 Arboretum 1917 Arnold Arboretum Arnold Arboretum 1906 Biltmore Nur. 1895 Ellwenger & Barry 1896 Jack coll., Kalispell 1921 Arnold Arboretum 1900 Veitch Nur. 1999 Manda Nur. 1905 Barbier -Nur. 1902 Veitch Nur. 1899 1d Arboretum 1896 Ellwanger & 1899 little : 1897 Arnold Arboretum 1896 MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XV Harold N. Moldenke LASIOSTACHYS Link The type of V. leptostachya was collected by Ynes Wilton Win- n San Luis Obispo tinge hE California, on June 17, pera and is Reptas ted in the herbarium of the Califo rnia Academy of Sci- snces at: San Francisco, while that of V. lamberti Spreng. "s Le- jeune s.n., collected fron cultivated material in Belgium and de- 3 species here under discus discussion, is actually V. elegans H.B.K., While the V. lamberti of Penny and of Sims is v. V. canadensis (L.) Britton; the V. prostrata of Bentham is V. lasiostachys var. sep- tentrionalis Mo. Moldenke, while that of G. Savi is V. bracteata Lag. & Rodr. Verbena lasiostachys var. abramsi onlaene Jepson is herein regarded as V. abramsi Moldenke, wh: which s Verbena lasiostachys has been employed by eicinatiee in suc- represent natural hybrids with related taxa. Hybrids definitely kKnom thus far are with V. halei Saas (=xV. scorta Moldenke), with V. hispida Rufz & Pav. (=x¥. lecocgi Moldenke), and with v. urticifolia L. (=xV. ee poy a It almost certainly hybridizes with V. robusta Greene and with V. abramsi in the wild where its range overlaps that of those species. Mattoon (1958) | states that V. lasiostachys is offered to the horticultural t hy. Cijde Robin, Wlpse (1845) places V. prostrata R. Br. in the synonymy of V. aubletia L. with a question. Jepson (193) reduces var. ably in the process of becoming stabilized. Herbarium material of V. leptostachya has been misidentified and distributed rather widely under such te sg as V, ciliata Benth., V. hastata L., V. menthaefolia Benth., V. officinalis L., V. officunalis L., Ve polys stachya H.BeK., "Ve eet var", ve robusta Greene, and V. urticifolia L. On the other hand, the Bradshaw 68, Hoover 3612, J. McDonald Sn. [Big Sandy Creek, June 1915], C. V. Meyer - 2h0, S. B. Parish 11590, L. W. Reinecke s.n. [September 5, 19371, Rutter 17, L. Ce L59 60 PHITTOLOGIA Vol. 9, no. 7 Sratiay bans etSa ion: 1893], & s.n. (Wiech co ae June 1921], irs. R. M. Austin sits M. S. Baker 1) & s.n. [Morley's Station, May 22], W. H. Baker 4207, Braunton 100 & 378, W. H. Brewer 31, ae Butler 1621, H. P. He. P. Chandler 1353, Chesnut & Drew s.n. [Hy-Am-Pun, am, July 21, 3508), 4 Le Constance 8, W. B. Cooke 15149, Mrs. E. Cooper s.n. sta. bara, June 1879], Coville & Funston 1110, Davy & Blasdale 1 S87, Ge W. Dunn Eastwood 166, Eggleston 972), Elmer 3846, 566, aon. (Santa. Paula, 1881], Ewan 8401, Gifford 197, H. M. ae , Hall 175, Hansen 1823, A. A. Heller 5785 & 5919, T. M. M. Hendrix 269 & Y& 1043, oy Calif. estypenhe Se School s.n. [Poly Canon, May 1h, 1905], sen. (Santa Barbara Forest, May 2h, 1933], J.T. Howell TZ & SoM. adits Pass, July 3d, 1887], Johannsen 6, A. A. Kellogg sen. (near San oe ie 1866], H. C. Lee ies Ss C. C. Marshall s. ee ens ee tener =, £8 = & Bee pits of LAs bei & Soe eam s.n. Palmer [Way 1902], Rothrock sn. (Santa Barbara, June 1575], Schulthess son. (Kelseyville, July 3, aeaty H. K. Sharsmith 1139 & 3187, iin s.n. (U. S. Dept. Agr. Forest Serv. 19562], Mrs. R. W. Summers son ~ [June 13, 1887], J. P. Tracy 2775, 5059, 12632, & & 12965, Ge G. asey S.n. Lear SO 1875], H. A. Walker 211, and H. S. Yates Sei are V. lasiostachys var. iy septentrl duals Moldenke; A- (Menifee, 1893], HL . Mason 8149, Edw. Palmer 309, , S» B. Parish s.n. (ouly 1887] & ‘a.n. [hug. 1 1809], Parish & Parish 969, sie tacchi 628, and G. R. Vasey s.n. [San Antonio River, 1500] are V. abramsi Moldenke; Suksdorf s.n. [July 22, 1881] is V. nae ata Lag. & Lag. & Rodr.; D. R. Harve Harvey 539 is V. neomexicana var. " xyopoda Perry; Hansen 477 is V. officinalis L.j and T - S. Brandegee s.n- [1881], Chesnut s.n. Forge a Clokey "sola, Gi. ey Davy 278 76, 25 259, & 865, Elmer 9 L950, Epling, Darsie, e, Knox, & R & Robison son. [Flinn pant Ractnaststeal- Midlit_ta ssh 8698, C. Re Orcutt 1301, be We W. Reinecke s.n. fiedudod City, June ak pre Schreiber 30 307, He A A. Walker 696, Wiggins & . L741, Wiggins & Gillespie 39 3977 are V. robusta Gr = Teaaae (1923) states that Vv. V. lasiostachys yields veecesaéaabie in some parts of California. Reichenbach (1827) describes the plant as follows: "Eine neue Verbena aus Californien, e 196), Moldenke, Monograph of Verbena hé1 Sweden (Uppsala). Bailey (1935) says that Kew handles the seeds. Common names recorded for it are "California het vervain", "prostrate vervain", "tibinagua", and "woo olly= spiked vervain"., I+ is mentioned by’ Sharsmith (1345) as occurring in the Mount Hamilton Range, California Eastwood (191) included ue lasiostachys as reported fram the Channel Islands under the name of V. prostrata in her concept of V. robusta Greene, She isceraha excludes V. lasiostachys from V. lasiostachys and not Vv. ro Verbena lasiostachys is often i Tivaated by the fungus, Pleo- spora herb: herbarum (Fries) Rabh. Felt (1912, 1917) describes a 3a gall in the form of a cylindric swelling of the flower-stalk caused by the Itonid gall-midge, Lasioptera ra verbenae Felt, based on a Specimen collected by Pi Hs Timberlake in the in the Puente Hills at Whittier, California, on November 13, 1910. The larvae hibernate in silken tubes attached to the walls of t gall. Schauer Seal. cites a Douglas collection in the DeCandolle Herbarium at Geneva from California. He affirms that he has seen cultivated aitert of the species and that Aig is related to V. stricta Vent. "sed jam crescendi modo et pasta i diversa.™ pei (1933) cites the following 22 additi ecimens not yet by me: OREGON: Curry Co.: J.C. Welson ms29 (G). Jackson te M. E. Peck 8702 (E, G). sade Co.: J. T. Howell 12h9 (BE). CALIFORNIA: “Lake Co.: Blankinship Sone [Clear Lake, Lake, 12 July 1929] (E); A. A. Heller 5919 (E, G); ite rrey 417 (G). Los An- eles Co.: Fritchey 29 (E). Mendocino ¢ aes Chestnut s.n. [20 May- 20 June 1898] (W). Monterey Co.: Elmer 05 (G); Hartweg 192 (); A. A. Heller 6778 (E, F, G). San Diego Co.: M. B. Spencer 994 (G), 1164 (G). San Luis Obispo Co.: irs. R. W. Summers 5.n- (39 gune 1867] (E). Santa Clara Co.: Panmel 187 (E). Santa C Co.: M. E. Jones sen. ae Cruz, 21 June 1801] (G, Po). Ventu- ra Co.: Abrams & MeGregor 5 (G). She says: "Although, from the original publication, one would naturally infer that V. prostra- Robert Brown, some without reference to og Desc rose t 462 FELT OL Care Vol. 9, no. 7 closely resembles v. robusta, which is much harsher and of limited distribution." The | Gniwertods 42lo, Edw. Palmer 309, S. B. Parish 2809, Parish & Parish 969, and Parry & Le Lemmon 3)j2 which she cites eing different from V Vv. lasiostachys, but - but to which she does not i a name, are V. abramsi Mo idenke, as is also the Vasey s.n. {San Antonio River, “1880). The Parish 11590 which she cites is re- arded by me as being V. lasiostachys — chys var. ar. scabrida, while Palmer 342 and Torrey 16 are V. robusta, and Clemens s.n. [Wood's Creek, Fresno Co., 19 June 1910], Sraanton 378, Brewe Brewer 31, Elmer 386, Hansen 96 & 1823, Heller 5785, J.C. Nelson 2669, Piper 6160, J. W. Thompson hjl2, C. Ep . Epling 545, J. T. Howell T7h & s.n. (Grant's Pass, July 30th, Te and Hammond _ 322 are V. lasiostachys var. septentrionalis. The Butler 1621 is in part var. scabrida and in part var. septentrionalis, while Wilkes 1661 is in part V. abramsi and in part Vv. Tasiostachys var, septentrionalis. Prof. P. H. Re ven, in a letter to me dated November . serge states that there is Pd sheet of Munz 673 in the Pomona herbarium, The following collections need to be re-examined. Although cited by me herein as Y. lasios stachys, at least in it is most (probably var. scabrida or var. septentrionalis], Heller 5919 {perhaps var. s Septentrionalis], J. T. Howell 11518 [probably var. Scabrida], L. C. Wheeler 666 [probably var. septentrionalis] and 2a8 [probably var. scabrida], and Wiggins 8957 [probably var scabrida or var. septentrionalis]. The Nobs & & & Smith 868 collection, tes regarded by me as var. septentrionalis in the California Academy of Sciences herbarium, is definitely typical V. lasiostachys the University of California herbarium| In all, 283 herbarium specimens of what I have re as the Citations: NOOTKA ISLAND: Née 112 (Q). YORK: faites Coe: Ruger s.n. [Brooklyn, Sept. 16, 1873] (Cc). pote Douglas Co.: Me E. Jones s.n. (Glendale, June 19, 1902] (Po—70666). Jackson Cost Applegate. 2228 (W—1h3273k); Cusick 4812 (P1—12111); E. C. Jomston s.n. [Trail Creek, July 31, 1931] (Gg--192085); Me E- Peck 8702 (Ba, N). Josephine Co.: L. F. Henderson s.n. [Dickson & Drake s.n.) (N). County undetermined: Abrams © lic McGregor 689 (N)§ M. E. Jones 2906) (North fork of Noyo River] (Po—-168l58)- CALI FORNIA: Alameda Co.: MacNeil s.n. [King's Run, May 9, '96] (Ge—- 31351). Calaveras Co.: Bigelow 5.n. et i Hill) (T); Blais- dell s.n. [Mokelume Hill] (Gg--31335). Lake Co.: A. A. Heller 5919 (N, Po--Su561, Ur); Mrs. L. R. Reynolds s.n. [foot of Mt. Sam parsing Teng gle (Gg--31354). Los Angeles Co: pane 1058 105 2h W—-13273k) ; M. L. Bryant 88 (Po—1557 3)3 Cohen (Po—2676lh); Cranford & Hiatt s.n. [Senta Monica Mts-, 2/26/16] 196) Moldenke, Monograph of Verbena 463 (Po—l008); Elmer 386 (N); Ewan 750 (En); F.°R. Fos S$ -30h2 (N); G. B. Grant 1365 (Du—756531), 2575 (Du-—75585); H. E. Hasse s.n. [Los Angeles, September 1887] (N, Pa, W—-71922); D. Haynes 3.0. [Pasadena, June 1893] (Po—-015); Hilend s.n. [Topanga a yon, May 19, 1933] pe ge ek I. M. Johnston 1320 (Du—10 Mi, Po—-l011, W—-1103173); MacFadden 926 (Ca--l,50803, dai McClatense s.n. [Pasadena, ae 2] (N); 3 Steele & Pratt s.n. [May Ly, *33] (Ob—69L:78);. 2L. C. Wheeler 666 (Po—21)392, Rs— 9719). Mariposa Co.: L. J. Fox s.n. [Yosemite Valley] (I). Mendo- cino Co.: Chesnut 1 (W--430451); Rattan sen. [June 1882] (Du— 19182); L. S. a 2 elke (Ca—882662, Ew, Ky, N, W—1897881, We). Marberey Coe: = (Du~6560) 3; Chisaki, Sharsmith, & Lag Hartweg 192) (N); A. A. Heller 6778 (Ca——58422, Du--2h203, 75576, Du--75587, N, Po--64519, W--467197)3 J» T. Howell ns62 (Ge—21257h, Po-—20175), 11575 (Gg—212573, Mn—20852), 11578 (Gg—-212582, Ss, Um-—-18), 11580 (Gg—212358), 39561 (2), 395958 (B), Loogs (B); Knoche 234 (Du); Mallory s.n. [May h, 1920] (Du— . 99)} Michener & Bioletti s.n. [Moss Beach, July ~~ (se— 95720); Ottley 1276 (Ws); Parry sen. (Monterey, 1850] (N); Re Ae Plaskett U2 (i, eae eae S. Rose 33338 (Vt), 53042 (B, Go, Go); Let L. C. Wheeler 29h (Ca--604725); Willey s.n. [Monterey , 1819-—i855) (Dt); Youngberg 228 (Ca--59770L, Po—2311h2), sn. [July 1938] (Po--258037, Ua--39522). Orange Co.: Booth 1171 (Ca, Po--201796); W. H. Brewer 717 (W--322h01); E. H. Cam Cupiett Ii 91 (Ge--31338)5 D. De L . Crawford s.n, [near Laguna, “July 2 (Po 4010); F. R. Fosberg 5.935 $.935 (Up); Munz 6734 pacers vung, (Ca—25160); 5. T. Howell =e (La); Pieters s.n. [June 1, 1910] (Mi). San Bernardino Co.: I. M. Johnston 1,07 (Ca--205350), 1608 (ca--205359, ca—212188, ‘po83815, Polo?) 5, Lim 30 (Du--143859); Munz 12313 (N); S. B. Parish (Du--91159), 9479 (Vi), sen. [Sept. pt. 1900) (Da—13858)3 EB. Le Peterson 831 (En); F. ll. Reed 7990 (Bt—15802). San Diego Co.: L6h, PEeE-? O'R, 0.0.1-& Vol. 9, no. 7 Mills sn, (Ur—-22835); Orcutt s.n. [Cuyamaca Mts., Sept. 1882] (Sd--6779, Sd—6787), s.n. [1890] (Mi); Edw. Palmer 18), (Sd—-6780) ; Snyder son. [La Jolla, May 1897] (Sd——252) ; M. E. Spencer 99h (N, Ob—508)2, Po—l7698), sen. [10/10/1918] (Ob—-50825); Stokes son. [Smith Mt., Aug. 1898] (Sd--13)70); Stover s.n. [Point Loma, May 4, 1938] (Sd--21028); L. F. Street sen. [May 12, '17] (Du—107329); Wiggins 261) (Au—-19)526, Ca--1612, Ca—165, Du—366023), 2967 (Ca——1695, Ca—l29803, Ca—73771, Du—182564, Du—3660)5); Wood & Negley son. (Palomar Mt., Sept. 20, 1936] (Sd--1605). San Luis Obispo Co.: L. D. Benson 7998 (Po—267639); 0. Degener 4685 (Ms, N)3 Fe H. Elmore 349 (Ak—10958); R. S. Ferris 9770a (Du--269819); Ferris & Rossbach 943 (Ca—60),9),0, Du—256))8)) ; F. R. Fosberg S. L981 (N); Gander 753) (Sd--25195); J. McDonald s.n. [July 19, 1925] (Gg--129)83); Nobs & smith 826 (Ca--191997, N), 868, in part (Ca—191998); Nordstrom 1132 [U. S. Dept. Agr. Forest Serv. 19818] (Ca—12hh07); Steele & Pratt s.n. [June 2, 1933] (Ob—-50823); Mrs. Re W. Summers s.n. [June 19, 1887] (Po—6l653); Winblad son. [June 17, 1937] (Gg—253028). San Mateo Co.: Blasdale s.e [Pilarcitos, June 23, 1893] (Ca—h50179); W. R. Dudley s.n. [June 18, 1905] (Du-—27863)); L. S. Rose 32321 (Al, Ms, Se—l159),); Walther s.n. (Woodside, June 9, 1919] (Gg——313h6), s.n. (Woodside, May 5, 1920 (Gg—31362); Wiggins 3782 (I, La); C. B. Wolf 607 (I, Rs--18513). ta Barbara Co.: R, S. Beal Jr. s.n. [Santa Barbara, 13 May 1918] (Ca—942561); Bingham 155 (Ka); Demaree 18103 (Du--2970h9, Io— 152354, Ok); Eastwood LOl (Gg—31358), 818 (Gg—3136h), s.n- Sant ta Barbara, July 2, 1917] (Gg—313h0); J. L. Fox sen. [June July 1930] (I); Kincaid s.n, [Santa Barbara, July 5 1935) (Se 34654, Se-—-3h655); Munz 9295 (Po—98527); G. Newell s.n. [San Is- idro, Aug. 1913] (Gg--31356); S. B. Parish 11061 (Du--91183); Suksdorf 179 (P1—13806), 179b (P1--138)05). Santa Clara Co.: A- brams sen. [Stanford University, April 1898] (Po--87969); We A» Atkinson s.n. [April 1900] (Du—2)216) 3 He A. Barker 608 10374); Collector undesignated 196 (Po--126)76) ; H. A. Davis S-Re (near Lagunita] (Se—l6752); Demaree 9205 (Io—-147325)3 We Re Dudley s.n. [June 1899] (Du--278632), san. [June 3, 1900] (Du-—- 278631); HT. E sen. [June '77] (N); Hewatt 162 (Au, Nt); Ingels s.n. [Lathrop Hills, Apr. 17, 1908] (Dp); J. D. Randall L22 (Du—63390, Du--63391); W. H. Rich san. [June 8, 1912] (DU 98022); Re J. Smith s.n. [July 1906] (Du—7758); H. S. Yates 5528 [U. S. Dept. Agr. Forest Serv. 14953] (ca--12h)16). Santa Cruz Co.: Akey 15h [U. S. Dept. Agr. Forest Serv. 11685] (Ca-- 12h40h); N. K. Carlson kh [U. s. Dept. Agr. Forest Serv. 11692] (Ca~—12h03); H. Davis s.n. (Glenwood, Sept. 1907] (Du—278635) O- Degener 4)82 (Ms, N); M. Epling 8310 (La, or—20188, Or-- 1964, Moldenke, Monograph of Verbena 465 20235); M. E. Jones 2215 (Bm, Br, Go, Po—71013); H. L. Mason 3365a (Ca--635116) 5 Pringle sen. [Aptos, 11 July 1882] (Vt); M. Rendall son. [5.VII.30] (Ca—95557); E. E. Stanford 88 (or— 20802). Shasta Co.: Nutting s.n, [May 22] (Po—126l75). Siskiyou Co.: Edw. Palmer 25298 (W— (W--277855); ?7L. C. Wheeler 3315 (St-— 9255). Sonoma Co.: H. Edwards Sone (way 1877] (Im). Tulare Co.: Culbertson 4210, in part (Po--63007); Steele & Pratt s.n. [July 20, aan (Ob--70165) . Ventura Co.: Abrams & McGregor 5 (Du--2420h, N, 612916); W. He Brewer 229 (W—-322h00); Dudley & Lamb 1,808 (Ca— 1687, Du--9558); Farr sen. [Ventura, April 9, 1911] (Up—52603); J.P. Harrington s.n, [April 1955] (W--2176802); M. E. Jones s.n. [April 27, 1926] (Po); Kendall s.n. (Ventura, },-13-23] (Po—17L16) ; ae 101 (La); Munz 13151 (Po--213656); Ogden s.n. ag July E Rug. 1892] (Cm); S. F. Peckham s.n, [Ojai, May 15, 1866] (W— ae H. M. Pollard s.n. [Oak View, Oct. 8, 19h] taccamioe Purer 6703 (Du-—271012), 67 6755 (Bt—-29702) ; Thacher 40 (Ca); Wood & Jon Jones §.1 S.n. (La). County undetermined: Ae Fitch s.n. (T); Herb. State Si Survey 717 (Ca--25161); Jepson 21226 {between Sims & Sisson] (Ca). CHANNEL ISLANDS: San Clemente: Munz 673) (Ca—2875) 5 Mur- barger 218 (Ca—557827); Trask 7 (w—-l682). MEXICO: Baja Cali- fornia: Fish Sen. [Oreutt s.n., 4-16-1885] pe aegis CULTIVATED: (Md). Spain: Herb. Hort. Matrit. hi 5 (Q)- “LOCALITY OF COLLECTION UN- DETERMINED: Collector undesignated 1558 (Sg--1611)); Herb. Vahl s. ne (S). TRISH se ed Ree Ee VERBENA eee f. ALBIFLORA Moldenke, Phytologia 2: 329, nan. anea "3: 113. 19L8;, widen, iph. List Cit. : ASE ogg dl & 1252. 1919; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2 27 & 198; Moldenke, Résumé 33 & 72. 1959. his form differs from the typical form of the species in hav- ing white coroll The type of the “form was collected by Le Roy Abrams (no. 5109) in Portola Valley, San Mateo County, California, on June 2, 19 and is deposited in the dig pas Herbarium at Stanford University. It has been collected in flower and fruit in June. In all, ) her- partie specimens » including the type, and 3 mounted phot ographs ve been Citations: CALIFORNIA: San pogo pS ee cee type, F—photo of t N—photo “J—photo o Jini Barbara ¢ a apart 579 fer). yoda Cow: A. Le Grant 66 PHYTOLOGIA Vol. 9, no. 7 s.n. (Grant Ranch, near Columbia, June 11, 1915] (Po). County un- determined: M. L. Kendall s.n, [l-13+23] (Po--17h16) « a Cts been var. SCABRIDA Moldenke, Am. Midl. Nat. 2h: 753- 19 Synonymy: Verbena SESS scabrida M. S. Baker, Part. List Seed Pl. North Bay 33. 195. Bibliography: Perry, Ann. Mo. Bot. Gard, 20: 291. 1933; Molden- ke, Am. Midl. Nat. 2h: 753. 1940; Moldenke, Known Geogr. Distrib. Verbenac., [ed. ae ns “aed & 101. 1942; Moldenke, Known Geogr. Distrib. Verbenac. : 2. 1943; Jepson, Fl. Calif. 3 (2): mae = P having the mature leaves dec ecidedly scabridous on the upper surface. The type of the variety was collected by Samuel Bonsall Parish (no. 11590) in a dry meadow at Monterey, Monterey County, Califor- nia, in August, 1917, and is deposited in the herbarium of Pomona College at Claremont. The plants so designated may actually prove to be a hybrid with V. robusta Greene, which they closely resemble. The plants often have their bractlets as long as the calyx (cfr. C. V. Meyer 240 at the University of California), as in that spe- cies, not shorter as in typical v. p saptowtactyss yet Bradshaw 68, Hoover 3812, C. V. Meyer 240, and L. CG. Wheeler 3315, with their open open elonga ated spikes, are @ surely mS conspecific with V. robusta! The variety has been collected on dry sunny flats and dry mesas, in sloughs and dry mead ows, on damp harks eae stream banks, along rivers and wet roads, and on damp sunny south slopes, at altitudes of 8 to 3500 feet, in flower and fruit from May to pobre: Herba- rium material has been misidentified and distributed und names V. casseoenciys Link » Ve prostrata R. Br., and ve ean Greene. Material of L. Cc. "wheeler 3315, Wiggins 8957, » and C. Be Wolf 2252 has been cited by me herein also under vy lasiostachys and its var. Septentrionalis and should be re-examined; it will probably all prove to be var. scabrida. Perry (1933) cites spec- imens of G. T. Butler 1621 in the Missouri Botanical Garden United npreragren ational herbaria as typical V. lasiostachys. In 3 herbarium specimens, including t of both nanes involved, and 2 anced photographs have gree pen tnienr >a Aclilisees | OREGON: Curry Co.: Kildale 6055 (Du). Jackson Co.! C. B. Wolf 963 (Du~-159333). CALIFORNIA Mlameda Co.: Wiggins 1964 Moldenke, Monograph of Verbena 467 8957 (Du--266760, Gg--310868, Rs——-23159, Se--5214h). Colusa Co.: R. Stinchfield 375 Boeri tenct N, Po—63857). Fresno Co.: J. T. Howell 34053 (2), 3h 090 (Gg); J. McDonald s.n. [Big Sandy Creek, June June 1915] 15) (Gg-—313h2, W—-1089985) . Lake Co.: M. S. Baker 8677 (N); Bradshaw 68 [U. S. Dept. Agr. Forest Serv. 20535] (Ca 12401); R. F. Hoover 3812 (Ca--766869, N, N); Schulthess s.n. (July 3, I931] (Du——2740)0). Los Angeles Co.: Abrams 2y81 (Du-- 24214, N, Po—156337). Monterey Co.: J.T. Howell 6539 (Ge-- 2071 9h) ; S. B. Parish 11590 (Ca—-205362——isotype, eritets of type, Po--63863-—-type, Z—photo of type); L. W. Reinecke s.n. (September 5, 1937] (Ca—770047); L. C. Wheeler 294 yee Cost C. Ve M Meyer 240 (Ca, Ca—-488893); C. B. Wolf 2252 (Ba, Du— 207790, Gg—173929, Rs--9996). San Mateo Co.: Barry 215 (Du— 279302) . Santa Clara Co.: Newell s.n. [Los Gatos, os, June 18, 1914] (Gg-—31353); Rotter 17 (W—308h27). Santa Cruz Co.: J. T. 11615 (Gg--212563, Rs—8h77); He A» Walker 730 (Po—126)7h). Sis- kiyou Co.: G. D. Butler 1621 (Du--2207, Po--69590); Eastwood 2102 (Gg——31357); Kildale 919h (Du); L. S. Rose 45098 . (Gg——-329h75, Goys 1. Ge Wheeler 3515 (Ca—882795, Gg—2330L1, , 13898). umne Co.: Mrs. W. J. Williamson 189 (Ca--882797). Ver tura at 3 T. Howell - 1028 . (R8--653) » VERBENA LASIOSTACHYS var. SEPTENTRIONALIS Moldenke, Am. Midi. Nat. Me 753. 190. Synonymy: Verbena lasi ostachys septentrionalis M. S. Baker, Part. List Seed Pl. North Bay 33. 1945. Verbena nevadensis Kearney ex Moldenke, Résumé 370, in syn. 1959. Verbena prostrata Benth, ex Moldenke, "Résumé 372, in syn. 1959 [not V. prostrata Alph, List ¢ 3, 2555 et & 285. 196; Moldenke, Phytologia 2: 336 (dot) and 2: - 1002, 111 vat iL. ? 17h, 1177, 1198, 1199, 1211, 121, 1216, 1222-1230, 1232, 1236 12381212" 121),°-12h8, 1252, 1253, 1259, 1261, 1288, & . 68 Poe? Os OLA Vol. 9, no. 7 1949; He Ne & Ae Le Moldenke, Anal. Inst. Biol. Mex. 20: 1). 199; Moldenke, Known Loner» pistriv. Verbenac., [ed. 2], 27 & 198. 191095 Yoldenke, Am, Mi 2 59: 342 & Aer. 1958; Moldenke, R& sumé 32, 33, 368, 370, “3i2, oe & 72; vee Résumé Su Suppl. 2: 10 & 12. 1960; Moldenke, Phytologia 8: 120, 121, a and 8: 206, 267, 279, 280, tle & 407. 19625 er ‘ae Sf THE TYPE TREE OF QUERCU BARK S XCOLUMNARIS ACORNS X7/10 493 Lok PHYTOLOGIA Vol. 9, no. 8 beaas Upper Surface 22% of natural size Lower Surface 27% of natural sine LOWER LEAVES OF QUERCUS XCOLUMNARIS 196, Laughlin, Caldwell Oak Upper Surface 23% of natural size Lower Surface 23% of natural size UPPER LEAVES OF QUERCUS XCOLUMNARIS THE GENERA SESBANIA AND BAUBENTONIA ON COASTAL CENTRAL UNITED STATES Clyde F, Reed The = Sesbania is represented in central eastern United States by S. exaltata (Raf.) Cory. The distribution given in Fernald (8th. *s Poa Ray p. 902, 1950) and Gleason (Britton & Brown, Thins. ahoma, being adventive from southeastern Pennsyl uae: to southeastern New York; also in the southwestern States and Tropical America The following are records of adventive specimens of Sesbania Po oP (Raf.) Cory from Maryland and Virginia. The folwers are yellow, mottled with purple; the pods are 10- 0-20 em, long, about 3 m. wide, tapped with the sloneatel slender style, subtended by the per- sistent calyx. (S. macrocarpa Muhl.). cote tip 19 n, Newkirk Street, Baltimore, on ehrome ore piles. Oct , 1958, 2 Reed h11,6. Virginia: Newport News, Warwick Coe, on 3, quite common. Aug. 7, 1959. Reed L0l5; Mov. 15, 1969. Reed a hs872, The gems Daubentonia is represented along the Atlantic Coast of United States by De punicea (Cav.) DC. (Mem. Legs, p» 286. 1826) The plants —> pd S Ttlantic C east and the Gulf Coast have gone under se as Daubentonia longifolia DC. (Jour. Ags Res. 20: $07-513. Bis. 190) and 5 seetent aa ca esii Watson. Recently, Wilbur (Legum, PL North Carolina, N.C. Agr. Exp. Station, Tech. Bull, No. 151, 1, 1963) mapped the distribution of D. punicea in that state. Let ertenta vee eftarnatr and Finn Sse me is a shrub or small tree, with th 20 to 60 leaflets, which are ob- ttyl gia and pointed; pet nes in color from scarlet to yellow mostly scarlet), in bin shorter than the leaves; the are oblong, 5-7 cm. long, compressed with four wings from the e valves and produced beyond the sutures; the seeds are ated Peet one another by sé ions, Very Poisonous ly sheep. (Marsh & Clawson, 1920, Jour. Ags Research, 20(6): 507-513, plate, as D. ja DC.3 Small, Flore ta, p. 615. 1913). FI to central Texa Mexico; adventive northward to North Carolina. North Carolina: Dare Co., Wanchese, Roanoke Island, common in swamps on south of island, everyone's yard, a8 Sate trees up to 10 ft. Aug. 28, 1958, Reed 11982, Beaufort Co: shruby 5-7 » cultivated near Bath. Aug. 7, "1957. Reed 39626. wastes, Rt. 172, 6 mi. W of Swansboro. Sept. 25, 1960, Reed "8309, (Used for Cancer Pro ject). 496 SELAGINELLA DENSA RYDB., AN ILLEGITIMATE NAME Clyde F, Reed Sela nsa Rydb., a species widely distributed in Western Nor ca ne aemtberas Alaska, through the western third of Canada down through the Rocky Mountains nn of western retns States to New Mexico, hasan and Texas, is ille antedated by S. densa Hort. ex Sim et) 3 mae re arate from Brazil, The next available name for our western plant is 5. engelmannii Hieron., S. longipila and S. haydents being reduced te synonyms for the reasons stated below seein densa Hort. ex R. Sim, Cat. No. 6: 60, no. 797 (descr. ). 3 abbey in Journ. Hort. & Cult. Gard. 6: eiid (Descr.). 1864; ex Cz. & Warcz., Cat. Hort. Cracov., p. 13 (nomen). 1864; Nicholson, Diets aaa ae 09 ("a garden name fer S. apus"). 1887. (Brazil). oe used by Lauche, Verzei 88 same s also Hort. Augustin., p. & poem 1856; Hort. ex A.Br., Ind. Sem, Hort. Berol., App. p. 23 (sub S- apus). 1857; Backh. Cat., p. 22. 1857. Hier (Hedwigia 39: Lageigh Dees 28, 1900) described three New species, all of which ref same species, S. densa Rydb. _ SELAGINELLA aaa cg Hieron. (l.ce, pe 291) is discarded because i@ lecali onymms "Bhotan", even vaaeals the specimens ag ala Ann. aaa Bot. ‘card 42: 70. 1955) are actually S. densa var. on selected a type, Saskatchewan, Fort Carlten, Bourgeau no ", » 1858, selagioniia longipila of authors, as Alston (Proc. Nat. Inst. Sci. . is S. indica (Milde) Tryon from om Nepal, Bhutan and Khasia. SELAGINELLA HAYDENII Miaron:: (lee, pe 296) is discarded because type 1 ties, — those given are very gener ageoricy ag? eee age Nebraska", Black Hills, n 1853-5), es at ex HY, 1859, ae US; paratype: B . Oregon (then Washington 3; fragn. in SELAGINELLA ENGELMANNII Hieron, (l.ce, Pp. 291), based on a speci- men from Colorado is the most acceptable name to replace 5: 3. densa Rydb. Several of our floras from western United States have used S, densa Rydb, for this species. Rydberg in his Flora ef the Rocky Mount: reduced S,. e Imannii « bow’ uii and e Hieron, to S. i Ere col T,a,faton under S. = p= J Hieron. (6 BY. pe 293) agg: Se eoLumbiana nar er) ongs under Maxon Se te S. aes Pe 297) prem, [iaten te North Am erican Ferns, p. 159. 1933) maintains Wallacei over 2- mace a no Misrom under S» rupestris (L.) Spring. 497 9 498 PHYTOLOGIA Vol. 9, no. 8 During my studies of Selaginella several collections and locali- ties in addition to those given y Tryon (1955) help te fill in the distribution ef S. e imannii and its varieties, I wish te thank the euraters in the gry herbaria for permitting me to anno- tate the specimens in theit herbaria: Univ, ef South Dakota (SDU); om. of North Dakota (NDA); Univ. of Southwest Louisiana (USWLA); Delaware (U.Dela.); Univ. of Connectieut (CONN); ssoghe of Pish "& Wildlife Raekdine at “wer nt, Maryland. Specimens Reed Herbarium are indicate d by REED and those in the United States National Herbarium by US. SELAGINELLA ENGELMANNII Hieron., Hedwigia 39: 29h, no. 5. (Dec. 28), Colerdde near Empire City. G. Engelmann. Aug. 27, ton (3B, holotype; US). SELAGINELLA ENGELMANNII sor) “poy yee Syn.: S. densa Rydb., N.Y. Bot. Gard. 1: 7. 1900, S. densa Hort, ex Sin, 1853. etna; Little Rocky Mountains, Drs ie Y. Havard, 1889 (NY, holotype; fragm, at US; paratypes: Nevtrerrys W. 53h3 172; Montana, Mrs. Jennie Moore). (Watson in 1850, GH is S. ei Hieron.). S. densa Var, densa; Se Amn, Missouri Bot. Gard. 2: 68, fig. LL, 05, 07, map L5. 19 aD dom ila Hieron., Redwigta 39: 291. 1900 (Pass 28). Sas- tien, Bour Mer. 29, 1858; + Kew Exs, No. 1239 , iw, °?, US, YU), wro = beled " 3 paratype: iffith, Bho otan, B; fragm, NY, Tryon ses p 70) states cimens. of Hieronyms species, S. longipila, are sige S. densa var, densa esidiicts tik ipa for forma Wicieipdla A.Br, ex Hieron, ex He 39: oer. enii Storen. in Hedwigia 39: 296 (as haydeni). Sl (Dec, 25). Tactotrpe type: S. Dakota, Black Hills, 1863- oh. F.V. @ Mo, ~ tag » US); paratype: "Oregon", 19 N. Lat., "16859. A a B $° is var. densa (Rydb,) Clute, Fern Allies, kon (Hieren,) Clute, in Fer Bull. ak es ; 3’ : rv stris forma Hieron.) Clute, in Fern 16: 52, 2 SRSEH oe ? Canada (Manitoba, Se Alberta, British Columbia); United States (Montana, North Dakota, South Dakota, Oregon, Wyo- ming, Colorado, Nebraska), Bennie oe i soil. ag Ar ; mage 5 23 3 Swift bare — 8. June 2), 1917. pitied 1 oF feat (nko. REED Man oo oem sand (Wa, REED 196), Reed, Selaginella 499 on short grass prairies, 2 mi. S of Da » Medicine Lake, Nat. Wildlife Refuge, Sheridan Co. Aug. 10, hd. Neil Hotchkiss 6931 (Fish, & Wildlife, Patuxent); Middle Creek, Gallatin Valley. July enshi hillsides at Atkinson e Rec. Atkinson, Holt Co. Sept. 21, 1960. John W. Thieret “6325. (Ustia, REED), North Dakota: Dry » Towner, McHemry Co. Aug. cag L.R. Eales 2337. (NDA); peneaias "VeHenry Co, Aug. 12, 19086 De ° ONN, REED); 15 mi. S of Sea McHenry Co. re Bie 135. om Stevens 1535. (NDA) s . hf ° phe 1952. ary prairie, E of Marmar Robt. T. Clausen 8200. (NDA, REED); ary cate » Epping, W Co. Aug. 1h, 1941. 0.A.Stevens. (NDA, REED); on * rocky mein of )3 top ef rocky hill, Powers Lake, Bai Co. a Sas 30, “1916. ”0,A.Stevens. (NDA); prairie gprsndse pie in oakwoods, ? Co. 12 i960. 0.A.Stevens & D.R.Moir. (NDA); gandy pr , Hamm, Eddy Co. July 11, 1959. O.A.Stevens 2110 )3 on gumbe in , Wade, Grant Co, July 12, W.B.Bell 632. (NDA); prairie, Ross, Mountrail Co. July 12,1960. oA. & D.R.Moir. (NDA); dry sandy 4e ridge, Akr . 0.A.Stevens = ); ary prairie at foot of butte, Black Butte, Slope Co ,» 1960. 0,A.Stevens ee sete dry rocky "hills oidas raring Stark June 21, » 190% Je : (NDA), South Dakota: Slim Buttes, Harding Co. » Age 20, 1910. S.S. Visher oH (SDU, REED); common on high Tablel Harding Co - dy 15, 1920. W.H.Over & Solem 358, (SDU, REDS valley near Game Lodge. June 2b, 192h. Ar MeInto; cm (SDU) hills jut W of Deerfield. July et Ce Intosh A-1;6, (SDU). ¥ SELAGINELLA ENGELMANNII VAR. lia ae Norse Reed, comb. oo Syn,: S. densa var. scopulorum (Maxon) Tryon, Anne Missour Bot. Gard. 2: 67, Mg ee soe The 19556 orum om in Amr, Fern Journ. 11: 36. 1921. Standley per (Holotype: c, Cracker Lake 15732 (US); ‘peratypen: i listed by Tryon, 1955). s Nutt, ex Baker, Handb. Fern Allies, De um ? 352 1 ByNe)e (pcinen at Kew; fragm. = heal $3, columbiana A.A,Eaton ex Maxon, Amer, Fern Journ. 11: 37. 1921 syn.).» (hyidently based on Shaw 398 and fd Heacock 235, both (in from British Columbia). Canada (Alberta and egg oi United States (Montana, Washington, tg ce gh wromnte + Utah, Colorado, California, Texas, Arizo ae ye Trail, Belt Mts. ae + 1925. es) Montana: Swif (U.Dela., REED); Roek Palas, bane = Seu Deion Perk, Jul: 26, « Th M.J.Elrod. (U.Dela. Oregon: ground, “id if Mack Rock Lookout, eley . 500 ft. Aug. é erie ae ae geal St. Coll. eat REED). 500 PHYTOLOGIA Vol. 9, no. 8 ee sags LAURE var. STANDLEYI — oe comb, Syne: tandleyi Maxon in Smiths. Misc. C 72(5): 9. 1920, (Holstyper Yee eaten! vic. Sexton Glacier, aera it. — Aug. 7, 1919, standley uae (US); pasties listed be: Tryon, 1955). Southern Alaska; Alberta and British Columbia; Montana and Colorado, Colorado: Pikes Peak, at timberline, S of Windy Point Seetion House, 11600 ft. elev. Sept. 35°3922, zeMe Johnston 3908. Os BED, Inter- mediate ——s en var. stand vare sco pulorum: tral sandy soil on sand hills just N of Gothic, 9500 ft. elev., peeeetn Co. July 16, 1937, E.T.Wherry. (REED). NOTES ON NEW AND NOTEWORTHY PLANTS. XL Harold N. Moldenke aa oe var. PARAGUARIENSIS (Briq.) Moldenke, comb. trina var. paraguariensis Briq., Ann. Conserv. & ney Jard. Bot. aS 7-8: 305. 190k. ALOYSIA TISSIMA var, REVOLUTA (Moldenke) Moldenke, comb, nov. rs iycioides var. revoluta Moldenke, Phytologia 3: 108. 19h9. ALOYSIA GRATISSIMA var. SCHULZAE (Standl.) Moldenke, comb, nov. pis = trina var. schulzii Standl., Field Mus. Publ. Bot. he . 1929. LIPPIA HIRSUTA var. GLABRESCENS Mo idenke, var, Haec varietas a forma typica speciei foliis eiptis vel glab- rescentibus recedit. his variety differs from the typical form of the species in having the leaves glabrous, subglabrous, or glabrescent beneath. The type of the variety was collected by Ellsworth Paine Medellin to Rionegro, at an altitude of 2000 meters, Antioquia, Colombia, on November 17, 198, and is deposited in the Britton Herbarium at the New York Botanical cal Garden, CANESCENS , f. nov. Haec forma a forma typica Gientae corollis albis recedit. This form differs from the typical form of the species in 196), Moldenke, New and noteworthy plants 501 having white corollas. The type of the form was ected by C. E. Smith, » Fe A. Peterson, and Narcisso sedge ei S940} in gray to setts soils Ss and MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. XVI Harold N. Moldenke VERBENA LIPOZYGIOIDES Walp. weer literature: F. Phil., Cat. Pl. Vasc. Chil. 220. Fruiting-calyx connivent at the —_— schizocarp half as long as the mature — cocci black o: er-red, oblong, areolate, Sulcate on the dorsal surfac 27, 33, 3, ogr. Distrib. ., [ede 2], 18, 19 ho 61, 65, 70, TL, 73, Is 3e——io2, 106, 122, 150, 154—156, 16h, & 198. 19L95 2, Phytologia 3: 75, 76, 133, 136, 140. 1949; Cabrera, oa 20: 175 & 337, ‘ae 7 ’ a aes Neal, In Gard. Hawaii 636. 1949; R. Espinosa, Sur Ecuad. 2: 68. 19h9 [to be continued] 506 PHYTOLOGIA Vol. 9, no. 8 Index to Authors in Volume Nine Banerji, M. L., 265 07 Bennett, Re We, 57 Moldenke, H. Rts, 7 » 98, Cuatrecasas, J., , 182, 186, He? ee 267, Degener, 0.& Ley 55 350, 351, 159° 500, 501 Eiten, L. T., 481 Reed, C. F., 496 497 Harkness, B.,: Schery, R. Laughlin, K., 101, 108, 185, Smith, L. B., 2h 2 Maguire, B., 35 Moldenke, A. Le, 55, 181, 238, Wurdack, J Index to Supra-specific Scientific Names in Volume Nine Abelia, 28 Amorpha, 31 Abeliophyllum, 28 Ampelopsis, 31, 432 Abies, 28 Anacardiaceae, 39 Seaeias 15, 199 Andrachne, 432 antho' » 428 Aneilema, 339 isan, és » 266, 428-30, Annonaceae, ),32 themis, 358 Aceraceas, 1,28 Antirrhimm, 335 Achyranthes, 165 Aphanoides, 345 Actinidia, 430 Aphelenchoides, 351 Actinidiaceae, 430 Aphis, 351 Acutae, 338 Apios, 2h0 Adesmia, 295 Apium, 165 Aecidium, 222 oliaceae, 48 Aesculus, 431 Arachnoiies, 55 Agromyza, 351 Aralia, Ailantius, 31 irzlintess, 428, 432, 447, 49 Archips, Aregelia, 2)3 Albizzia, 31 Aristolochia, 432 chemilla, 36 ene 432 Allardtia, 252, 256 Aronia, » 240 Artemisia, “2h, 135, 199 Aims, 217, 431 Asclepiadac 453 Mloysia, 182, 500 Asclepias, a7 Sasaies 366 Asinina, 13 417 Aster, 21 » 16h As pine, 7 ey’ sh. \aee A letia > 7; a ts , Imelanchier, 431 "30h, 36h, 398 1964 Index 507 Axinaea, 09-11 Cedrus, 435 Baccharis, 7, 396 Celastraceae, 35, hhh, L55 Barbacenia, 262-26 Celastrus, 35 Berberidaceae, 32, 53 Celtis, 435, 436 Berberis, 432, 33 Centronia, 413, Berchemia, 33 Cephalanthus, 36 Betula, 433 Cephalo Ss, Betulaceae, 55 Cercidiphyllaceae, 36 Bidens, 283 Cercidiphyllum, 36 Bignoniaceae, 43h, 135 Cercis, 436 Blakea, 25, 26 Cercocarpus, 134 Botrytis, 351 Chaenomeles, Bouchea, 70, 165, 393 Chamaecyparis, 36, 437 Bouteloua, 15 na Se Bromeliaceae, 25, Chimonanthus, 37 es ae oes ten aocentina” 1437 Chrysanthemm, 316 eae: 768 Cinchons, 337, 3hh Bro ussonetia, 433 Cirsium, 164 Brunellia, 3h, 3h5 Citharexylum, 98 Buchnera, 215 Glasrants 437 Buckleya, 33 Clematis, 37 Buddleia, 433, 434 aisredeniron, a, 43h Clerodenae > 183, 168, 2h1 Buxaceae, 3h, 55 Cl 1, > 43h Sean 1,38 Calceolaria, 333 C ’ Callicarpa, 188, 3h Cc Calochortus, 199 Colutea, 1,38 canthaceae, 43h, 137 Compositae, 1, 56 Calycanthus, 43k Comptonia, 438 Calyptrella, 12, 413 Conostegia, Campsis, volvulus, 217 Canistrum, 256, Conyza, 1- Cc yatorsacese, 128, 1h3, bbb, Coreopsis, og Co ay, Goriariaceae, 138 caragin, 9 x Gornacene, 438, bh? Carex, 93, 93, 217, 283, 337, 338 Cornus, 438, 439 Carpinus, 13h ta ee C D , Se as, 188 Corylaceae, hw, 433, 43h, 1,39 Castanea, 43 Coryiopsis, 439 Castilleja, 115 gory 439 Sees Costus, 339, 340 Ceanotims, 435 Cotimus, Gedrela, 135 eee, 508 PHYTOLOGIA Vol. 9, no. 8 ee 109, 185, 186, Ericaceae, 437, hhh, hho, 450, Cremanium, )21 ee at See Erinus, 197, 388, 393, 396, 399 3 preccau’en, 21, 350 ——s ophyes, 222 ee Erysiphe, 222, 351 roe ig tno obalanus, 90 raceae, 1151 =H Cyperus, me Coph a, 348, 39 Eucommiaceae, ))j) eee | Eumandevilla, 348 Cyriliacess, h)3 —— Cytisus, Lh3 Euphorbia, 56 ee Euphorbiaceae, 432, Lh6 Datura, 16 eee Daubentonia, 96 Brochorda, peated Daucus, 2L0 Fagaceae, 435, hh5 Davidia, 3 See? Decaisnea, )3 Floscopa, 339 Delphiniua, 135, 316 Foliosae, 14, 4h, 47, 51, 8h, Denteies TG x ae 155, 213, 289, 30h, Diacrisia, 351 waibese’ ifs Dicaeoma, 16, 222 Foresticra, hs Dichorisandra, 339 Forsythia, bhs Didymella, 222 Fothergilla, bl Diervilla, 3 Frankliniella, 352 Diolena, )1))-l16 Fraxinus : Diospyros, hhh Gaillardia 382 Diostea, 13, nh, 479 Genista, f Dipelta, hhh Geranium, 15 Diplarpea, 15 Ginkgo, Nh Distichlis, 195 Ginkgoaceae, 6 bryopteris s 55 Gladiolus, = ng a, 186, 188 = a ee ee ahs ei 13, y*, 299, 301, 02, $9, Elaeagnus, nar Gleditsia, iS Eleocharis, 81, 483, 48h Glochidion, Elsholtzia, po Gnaphalium, 407, 69 Endothenia, 1 ‘é ee hhh Graffenrieda, 09, 412, 413 77 eae, 0, 196) Guamania, 248, 259 Gymnocladus, 41,6 dron, 6 Hamamelidaceae, 439, lS, bh7, Hierobotana, 31 Hilaria, 28, 199 Hippocastanaceae, 130 Holodiscus, 47 Holophyllae, 126, 213, 269, Indigofera, )8 Inermes, 1), 31, hh, oh 51, » Lh, 122, 126, 155, 213, 287, 30h, 36h, 382, 398, 501 Inula, 241 2 Jamesia, ))8 Juglandaceae, 35, Lh Jugians, Junceae, 11) Junciformes, 11, 478 Juncus, 70, 196, 283, 295 bh, Juniperus, 13h, 135, bb8, bho Kalmia, bh9 Kalopanax, Lh9 Index Keithia, 76 ia, “ho Laennecia, 1 Lamiaceae, 190, 07 Landularia, Lantana, 7, 99, 18h, 166-188, 209 Lardizabalaceae, 31, 43 axa’ eandra, 13 Teguminosae, 56, 431, 43h, 436, potto2e hh3, Lh6, Lh8, 450, 2 > Leitneria, )50 Leitneriaceae, 50 , 903 393, he Li » 451 Loganiaceae, 33 Lonen’ , Lychnidaea, 388, 389 510 PHITOLOG@G7 4 Lychnidea, 388, 389, 39h 4s2 2 Macranthae, 14, 44, 47, 51, 64, 122, "30h 36h, 362, 398 Macrocentrum, {17 Macrophomina, 351 Macrosiphon, 351 Magnolia, 52, 453 Magnoliaceae, 51, 452 Mahoberberis, 53 Mahonia, 3 Malus, Halus, 108-112, 453-455 hh? onolena, Moraceae, 33, hh2, 452, ki Morus, 455 5 » Joe ay Myricaceae, 438, 55 ityrmidone, 2h Vol. 9, no. 8 igus » 351 amy pee 4,07, 69 Neillia, Neoregelia, 22-26, 258 Neosparton, 113, 11h, 479 Neviusia, 55 Nidularium, oy) =p 261 Hioreaberete, Nobiles 30L, i, 365, 375 Oleacea Rigasaas: 428, 437, bb5, 451 1e ry 2h, 48 ’ 199, 5 Fasiystasiye, 2, 51, 382 Paeonia, Soa 187, 266 Parrotia, Parrotiopsis, 56 Parthenocissus, 21, 456 Paulownia, Penstemon, 57, 58 Persicaria, 93, 190, 358 158 Phellodendron, 456 Fhiladelpins, 456, 457 Phieum, Phlox, a Photinia, 157 Phyla, Phyla, 165, 188, 190 Phyllosticta, 222 196), Phymatotrichum, 351 Physocarpus, 57, 458 Picea, 58 Pinaceae, 4,28, me Dat 42, LL8, 450, 455, Pinus, 1 135° Piptadenis, 3 a Pitcairnia, 246, 248, 249, 259 Platanus, | 2 Pleospora, 461 Poa, 21 Polemoniaceae, 16, 17 Polygonacess, o Polygonum 217 Polypodiacese, 21 Polystichopsis, 55 Priva, 44, 31 Prosopis, 28, 143, 163, 3h7 Pseudo-Catopsis, 25 Pteris, 135 Puccinia, 16, Puya, Puya, 219-251, eS 260 Pyrus, Quercus, aes 181, 21, Rhizoctonia, 3 247, 259 Rosaceae ha, b32, ys 4,39, » Who, 453, rae: Salpingia, 416, 417 Salvia, Salvia, 135, 199, 295 Scirpus, 217, 203 Scrophulariaceae, 219, 456 Solidago, 217, 21 Stachys, Stashyterpheta: 78, 99, 188, 220 Stenotarsonemus, 351 Stevia, l Stipa, 29 Stylodon, 35, 215, 220, 4,78 512 Tillandsia, 27, 251-256, 259~261 Tococa, 22-25 Tradescantia, 339 PHYTOLOGIA Vol. 9, no. 8 355=369, 371-377, 379-391, 469, 471, 473-480, 500-503, 0 vemenaca, ri 31 h2, hh, 47 Tragia, 1 ? ’ ’ ’ one 36h, 381, 382, 398, u7hs, i787 Triolena, 15 501 Tripogandra, 339 Verbenaceae, 215, 388, 3h, 437 Tripterocalyx, 266 Verbenia, 192 Typha, 196 Verbent, 206 Uerbena, 372 Verbina, 165 Ulmaceae, 35, Vernonia, 213, 217 Verbena, 8-19, 21-5), 59-67, Viola, 69-73, 75-95, 97, 100, 113- Vitaceae, 431, 456 131, 133-137, , 143- Vitex, se NM ay He, 195, 197-207, 2 Bh Saee, 22, 256, 258, 261 > 271, Vulgare ee, Nebsteria, 481, 483, 8h Xanthiun, 299-309 Yucca is 28, 134, 135 15- nt, sxe, Zapania, 0 ; Zizyphus, 2) Publication Dates of Volume Nine No. 1 = February 1963 PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. 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