PHETOLGIA Soiaial 8 1961 - 63 Missouri BOTANICAL GARDEN LIBRARY PHY TOLOGiS Designed to expedite botanical publication wf Vol. 8 September, 1961 No. 1 CONTENTS SMITH, L. B., Notes on Bromeliaceae, XVII .....+.++++:- eee ne 1 MOLDENKE, H. N., Notes on new and noteworthy plants. XXVIIL.... 14 MOLDENKE, H. N., Additional notes on the genus Aegiphila. XVI... 15 MOLDENKE, H. N., Additional notes on the genus Vitex. I ...+++++> 21 oct 13 1961 ARDEN BRAT Published by Harold No id Alma L. Moldenke — 15 Glenbrook Avenue Yonkers 5, New York, U.S. A. Price of this number, 75 cents; per volume, $5.75 in advance _ Vol. 7, No. 9, was issued July 5, 1961 fo NOTES ON BROMELIACEAE, XVII Lyman B. Smith MEXICO and CENTRAL AMERICA HECHTIA In making keys there is usually the problem of whether to at- tempt a natural arrangement with the simple and presumably primi- er in my key in the North American Flora (19:84. 1938), meayen for the nl i emphasis on an inferior ovary. This last the advantage o s Seinaivg together the southernmost species, rs = chroantha and H, guate’ emalensis, as they obviously » should The natural relationship of the groups is probably t ae EE =. Ew a8 —— V is —— speed of the inferior ovary vi its devel- t has outhward. Group III has developed P -gaeras er protection te its Posner enabling it to spread north cooler climates pe iid or its remnant almost or wholly inferior (the remnant aring to be an apical swelling of the pedicel in the Geinthaanir lowers)......00. eoeeess ccubkey V 1. Ovary - its t lar, or wholly superior, oe remnan gely S when present not abruptly swollen at 2. Secaus” sea floral bracts drying anon roseate with no hy- margin, ‘ ovat eseeeeee scubkey IV 2. Sepals and floral ‘bracts brow, s stramineou ous or hyaline, or if roseate (H. elliptica, H. scariosa) then with h hyaline mar- 3. Floral Breve strongly convex, ample, always exceeding the pedice d mich of the re est IIt 3. Floral oe flat, usually mich shorter than the sepals >» or if convex then shorter than the pedicels; flowers dis- ct pecies 4. Leaves densely serrulate with teeth not more than 0.5 mm long; plant delicate, less than 1 m. high; ‘it linsianane Ba FRYETOLOG IA Vol. 8, now 1 very lax L. Leawes laxly eeseee .Subke y II and coarsely serrulate or “where not known the plant coarse and over 1 m. eeccccccccccccccsendurkey I Subke 1. Inflorescence slenderly cylindric; branches simple or 3-parted at base, not over 12 cm. long and 2. ee, internodes. i - pedicellata 2. Branches densely flowered, or flowers oubverti cliitte e. 3. Rhachis mch flattened; branches often 3- —— from the base 3. Rhachis subterete; ; branches s simple 4. —— pals ually mch less. owered; pedicels slender, 38 mm. long; cape-bracts mich exceeding the mg 2. H. podantha -5-4, mm. long, Sbttabes 3; rhachis stout, 5. Speen strongly repand; rhachis subalate. 5. 3. H. subalata psy with straight margins between the spines; S nearly even.... 4. Sonate —— 1-2 mn. long. 6. Rhachis -seeesehs H, melanocarpa to 10 cm. long, ascending; lower primary bracts oe from an ovate base; floral bracts shorter than Srasutens the pedice 6. Rhachis 25 cm. CU nbee ts cade - caudata 7. Floral bracts anarg shorter than the pistillate pedicels; rhachis to 5 em. lo * ee 6 H 7, — bracts equaling or exceeding the pistillate pedi- 3 rhachis lo cosceseels H. 1. Inflorescence pyramidal or if subeylindric then broad with elongate branches, Key re price plants. — nies se equaling or shorter phi: 25 pedicels; 8 evenly flowered in most s 10. Pedicels continuous with the Matin. gone articulated. li. The pedicels stoutly veiess 3.5 mm. long; seeds with a thick blunt wing.. ll. The a slenderly 12. Leaf-sp 8. H. roseana y cylindric » 4-10 mm. long. 5 mm. long; at Scape-bracts remote; pedice cela ine — long, slender...........9. a2. ype nope - long; s bricate; montana scape—bracts all large and in- ue 10 mm. long, almost filiform. 10. gimp 10. H. matudae articulated with the rhachis. 233 6 even exce ernpes ddan Nee pt for a median — pedicels wpa 13. Carpels reticulate; pedicels. sulcate, ‘gubterete. 12. H 9. Floral bracts mch exceeding the thesis; racemes flowers strict. ‘ reticulata short pedicels at an- interrupted pote more or less moniliform; 1961 Smith, Notes on Bromeliaceae 3 14. Rhachis slender, even or faintly angled. 13. H. stenopetala 14. Rhachis stout, ee sulcate.............14. H. galeottii 8. Key to staminate 15. Sepals acute. 16. oe In@ludedscs sie sss «209. H. montana 16. Stamens exserted. i aly deen mich thickened toward base; orgs = stout; cemes suaveolens 17. rede ea greg pedicels Shenae racemes 18. Rhachis strongly compressed througho 19. Stamens included; floral bracts Sauriig or shorter than pedicels eell. H. glabra 19, Stamens exserted; floral bracts exceeding the pedicels. 17. H. conzattiana 18. Rhachis compressed only a 20. The rhachis seers Ha oe “faintly so 2 a ee ce S..H. " sphaeroblasta 21 - Stamen 13. H. stenopetala 20. The thactia het deeply ilestd....- : Ak H. galeottii Subkey II 1. Sepals ovate, broadest at about one-third their length; s 3m, 1 lin ong... H. dmanioides 1. Sepals cubtriangular, broadest at base; gies 5 mm. long. 2. Pedicels 3 mm, tong» more than twice as long as the floral acts. Siu Ae one 4 4s aos H. tillan oS 2. Pedicels less “than oy mm, “long » equaling ae shorter than t floral bracts........ 21. H. Tindal tisiun Subkey II 1. Flowers 8-10 mm, long; floral bracts stramineous or roseate with broad scarious margins. 2. Sepals at least as broad as long. 3. The sepals acute, 6 mm. long.......ccsecceeee22e i. texensis 3. The sepals obtuse, 325 mm. Long...++++.6...46023. H . mexicana + Sepals much longer than ad. 4 whe Loopels elliptic, oehyees leaves splashed rh bright eeeereeesesn one peeveuame H. elliptica hk. the. pei acute; leaves "gree 5» Floral bracts and nepea age with broad ark Marg. ene eoeeee ee eeaeeene - He carios sa 5. Floral bracts Bate, atramiiecus............s%26: H. seeeeaice 1. Flowers 4-7 mn. 6. Pere sisnguedy Tak or sublax except sometimes at extreme 7. Inflorescence amply tripinnate with the branches eee ely divided cocceee?. H. schottii 7. Inflorescence bipinnate except ‘at ‘base “where the branches are palmately 3-parted.......sccsccceccce2he H. fosteriana h PHYTOLOGIA Vol. 8, no. 1 = Spikes ee aes dense. 8. Sepals acu 9. Leaves Bets lepidete on both sides.. esi H. argentea 9. Leaves = abo . - H. capituligera 8. Sepals obtus 10. Primary “al ghatof ovate or suborbicular, narrowed abruptly to a ex. ll. Leaf-blades ees oes 3 branches short, mostly ex- ceeded by the 3 cm, long primary bract cts..31. H. confusa ll. Leaf-blades densely pale-lepidote on both sides; stami- nate branches to 55 mm. Sone and pistillate to 40 mn. both mostly exceeding t sae? sad bracts 32. H. namier-lapostollei 10. Primary braoke lanceolate or narrowly triangular, incon- spic 12. Leaves all coarsely serrate...........+..33. H. glomerata 12. Leaves (only the inner known) finely serrate. 34. H. gamopetala Subkey 1. Scape-bracts mich exceeding the internodes, foliaceous - H. desmetiana 1. Scape-bracts mich shorter than the internodes, vaginiform. Ze FAOWEER BUNOGRNLIO.. 8. ios civtunsac....... 2... --36. H. rosea 2. Flowers distinctly pedicellate 3 3. Floral bracts equaling or exceeding the pedicels. 7. H. meziana 3. Floral bracts about half as long as the pedicels. 38. H. laxissima the ovary remnant pgarelry to ees of the upper part of the pedicel; petal See bee bbe 008s 639% ec aatiionn e. Toning the ovary at ee pistil- nate flowers and nearly equaling the sepals of the stami- 3 flowers not more afhtia spreading....40. . “dichroantha 2 yee bracts inconspicuous; flowers mostly reflex 41. H. pan ae 1. HECHTIA PEDICELLATA S uae fete ae 1 Wats. Proc. Am. Acad. 26:155. 1891; e ray er Lie ds fi . . ° . Fl. 19: :89. 1998. p. figs. 30, 31. 1937; No. Am co. 2. HECHTIA PODANTHA Mez in DC. Monogr. Phan. 9:549. 1896; Contr. Pg Sop aA7tD> Bt. 1, figs. 27, 28. 1937; No. Am. Fl. 19: Hechtia liebmannii Mez Bot. Jahrb. 30:Beibl. 67:6. 1901. Hechtia tehuacana B, is Rob 5 3 Soon inson, Proc. Boston Soc. Nat. Hist. 1961 Smith, Notes on Bromeliaceae 5 MEXICO: Hidalgo, Morelos, Puebla. 3. HECHTIA SUBALATA L. B. Smith, Contr. Gray Herb. 117:15, pl. il, a = 22. 1937; No. Am. Fl. 19: 88. 1938. : Duran™go, Zacatecas, 4. HECHTIA MELANO OCARPA L. B. Smith, Contr. Gray Herb. 161:32, pl. 4, figs. 8, 2 1mé. MEXICO: Guerr 5. EGCEETEA CAUDATA L. B. Smith, sp moo. e« nov. reana L. B. Smith, cui affinis, foliis mlto ma vidal pi longioribus, floribus subverticillatis nullo e- ee tres oe 12 dm. ey (! Foster) ; ae over 5 dm. long; abeeine subo EN pole to 9 cm. wide, except at apex entire, brown and lustrous; blad 8 narrowly ppc col 55 mm. wide, red, suc- culent (1 tens covered with coarse appressed cinereous scales on both sides, partially glabrous april with age, laxly serrate mm pex brous; scape-bra linear subentire Lepidote blades; inflorescence bipinnate, cylin- dric, glabrous; lower primary bracts like the scape-bracts, the upper much reduced et bladeless; branches to 10 cm. long, sl der, slightly compressed at base, rather dense with the flowers eubverticillate; floral bracts ovate, apiculate, flat, af . long, the pedicels; pean s slenderly obconic; ovoid, acute, 11 mm. long, yellow-brown. Pl. I, fig. i: Section of staminate branch x 1; fig. 2: Staminate flower > airy Be Staminate sepal x 2; fig. 4: Pistillate flower (fruit) x i; teh 53 Old pistillate sepal x 2. Type in the U. S, National Herbarium, collected on rocks, be- tween Tehuantepec and Tuxtla, State of Oaxaca, Mexico, are 300 meters, pty 8, 1957, by Mulford B. Foster and 0. Hyning (No. 2999). 6. HECHTIA MOOREANA L. B. Smith, Contr. U. S. Nat. Herb. 29:522, aie. eaccnadl 7. HECHTIA oso L. B. Smith, sp. nov. - Smith, cui affinis, foliis mlto majori- bus, Scand Gract i bracteisqus lanatis, inflorescentiae ramis breviori- diffe -60 cm lanate at first (1 Foster) ; sheaths renttiels » 6cm. high, densely ro c. tan neous, strous; blades narrowly > 7 cm. wide, covered beneath with appressed cinereous scales, with age glabrous and lustrous 6 PHYTOLOGIA Vol. 8, no. 1 above, densely serrate at base, laxly so elsewhere with ae ve curved spines 5 mm. long; scape central "1 Foster), 5-6 diameter, lanate at first (! Reston); scape-bracts mare equaling the internodes, ovate with a linear lanate blade; od rescence slenderly cylindric; primary bracts like the upper scape-bracts, about equaling the branches; branches 2 cm. long, densely fl owered; floral bracts exceeding the pedicels but narrow and nearly flat, serrulate; pedicels narrowly obconic, the stami- nate 2 mm. long, the pistillate 2.5 mm.; sepals broadly ovate, obtuse, 1.5 mm. long, serrulate; staminate petals obtuse, 4 mm. long; stamens included; capsule ovoid, acute, 9 mm. long, casta- neous, even, lustrous. Pl. I, fig. 6: Staminate flower x 2; fig. 7: Staminate sepal x 2; fig. 8: Pistillate branch x 1; fig. 9: Qld stern sepal x 2. ype in the U. S. National Herbarium, Speneaees | on rocks in a very area, near Tequisistlan, State of Oaxaca, Mexico, al- titude 2400 eh April 1, 1957, by Mulford B, Siete er amd 0. C. Van Hyning (No. 2934). 8. on IA ROSEANA L. ms Smith, ai et are I172i75: pl. 2, cv aner Pueb va. 9. HTIA Ee ae Brand. Erythea 7:9. 1899; Contr. Gray Herb. i; 16, 1, figs. 38-40. 1937; No. Am. Fl. 19:90. 1938. echtia hts cellata sensu I. M. Johnston, Proc. Calif. Acad. " « 12:995. 1924. MEXICO: Baja California, Sonora, Sinaloa. 10. fags ia aia L. B. Smith, Phytologia 5:395, pl. 1, figs. i, 2. 1956. MEXICO: : Morelos. 11. HECHTIA GLABRA Brand Prigt Calif. Publ. Bot. 7:325. 1920; Contr. Gray Herb. 117: 117 sl, figs. 43-45. 1937; No. Am. Fl, 19:91. 1938. : ; MEXICO: Vera Crus. 12, HECHTIA RETICULATA L. B. Smith, Contr. Gray Herb. 117:17, pl. 1, fig. 46. 1937; No. Am. Fl. 19:91. 1938. MEXICO: Colima. 13. HECHTIA STENOP OPETALA Kl. Allg. Gart. 3:402. 1835; Contr. Gray ae 117:18, pl, 1, figs. 47-50. 1937; No. Am. Fl, 19:91. Hechtia cordylinoides Baker, Bot. Mag. 107: pl. 6554. 1881. pees pringlei Robinson & Greenman, Am. Journ. Sci. III. 50: MEXICO: Puebla, Oaxaca. 14. HECHTIA GALEOTTII Mez in Fedde Rep. Spec. Nov. oe das 1919; Contr. Gray Herb. 117:18, pl. 1, figs. 51, 52. 1961 Smith, Notes on Bromeliaceae 7 Fl. 19:92. 1938. MEXT 15. HECHTIA SUAVEOLENS E. Morr. ex Mez in ery Monogr. Phan. 9: 550. 1896; Contr. Gray Herb. 117:18, pl. 1, fig. 53. 1937; No. Am. Fl. 19:92. 1938. MEXICO: Indefinite. 16. HECHTIA REFLEXA L. B. Smith, Contr. Gray Herb. 117:18, pl. 1, figs. 54, 55. 1937; No. Am. Fl. 19:92. 1938. MEXICO: Michoacan, 17. eae CONZATTIANA L. B. Smith, Contr. Gray Herb. 117:19, 1938. rent era eee 1937; No. Am. Fl. 19:93. 18. HECHTIA SPHAEROBLASTA B. L. ee ae Proc. Am, Acad. 35:323. 1900; Reta Gray Herb. 117:19, 1, fig. 57. 1937; No. Am. Fl. 19: 93. 1938. MEXICO: rrero, 19. HECHTIA LINDMANIOIDES L - Smith, Contr. Gray Herb. 117:14, pl. 1, figs. 24-26. 1937; as Am. Fl. 19:88. 1938. MEXICO: Vera Cruz, : 20. HECHTIA TILLANDSIOIDES (André) L. B. Smith, Contr. U. S. Nat. Herb. 29:431. 1951. Bakeria tillandsioides André, Rev. Hort. 61:84, pl. 1889. Hechtia purpusii Brand. Univ. California Publ. — 73325. 290s 4 ses MEXICO: Vera Cruz. saschantns tillandsioides L. B. Smith, Contr. Gray Herb. 104: 193k. 21. HECHTIA LUNDELLIORUM L. B. Smith, No. Am. Fl. 19:97. 1938. MEXICO: San Luis Potosi. 22. HECHTIA TEXENSIS S. Wats. Proc. Am. Acad. 20:374. 18 85; Contr. Gray Herb. 117:19, pl. 1, figs. 58, 59. 1937; No. Am. Fl. 19:93. 1938. UNITED STATES: Texas. 23. HECHTIA MEXICANA L. B. Smith, Contr. Gray Herb. 117:19, pl. 1, fig. 60. 1937; No. Am. Fl. 19:94. 1938. MEXICO: San Luis Potos{. 24. HECHTIA CA L. B. Smith, Contr. Gray Herb. 117:20, pl. L, figs. 61 a. » 62 19375 No. Am. Fl. 19:94. 1938. CO: 25. fecur A SCARIOSA L. B. Smith, Contr. Gray Herb. 117:20, pl. 1, figs. 63, 64. 1937; No. Am. Fl. 19:94. 1938. 8 PHYTOLOGIA Vol. 8, no. 1 UNITED STATES: Texas. MEXICO: Coahuila, Tamaulipas. 26. HECHTIA ZACATECAE L. B. Smith, Contr. Gray Herb. 117:21, pl. 1, figs. 65, 6. 1937; No. Am. Fl. 19:95. 1938. MEXICO: Zacatecas. 27. HECHTIA SCHOTTII Baker in Hemsl. mel, Centr.—Am. Bot. a as 1884; is tad Poni Herb, 117:22, pl. figs. 69, 70; No Fl. 19:96. Hechtia Magda Mez in DC. Monogr. Phan. 9:550. 1896. Hechtia myriantha nents Bot. Jahrb. 30, Beibl. 67:6. 1901. Hechtia macrophylla Greenman, Field Mus. Publ. Bot. 2:247. Se) MEXICO: Vera Cruz, Yucatan, Chiapas. 28. HECHTIA FOSTERIANA L. B. Smith, sp AH. schottii Baker, af affinis, inftteedvibh ies ramis sim- plicibus vel infimis basi 3—partitis diffe Flowering 15 dm. high; leaves over 1 m. Longs sheaths reni- form, 10 cm. wide, brous except at apex, very dark castaneous and lustrous on both sides; blades narrowly triangular, 7 cm wide, glabrous above, covered with appressed cinereous scales be— neath, densely serrate at base, laxly so above with curved an- trorse spines 6 mm. long; scape ores and flattened at base and presumably pital li mm. wide at apex; scape-bracts broadly tr. xpos ing almost the Bas ood scape but their linear mostly adine blades equaling or exceeding the internodes; Sai rescence (pistillate only) Mptivaté except at base, 9 dm. 1 14 cm. long, rather evenly and sublaxly flowered, sulcate, flat- tened at base; floral bracts broadly ovate, acute, i, mm. long, much exceeding the stout obconic pedicel; sepals ovate-oblong, rown - long en, sublustrous. Pl. I , fig. 10: Section of pis- tillate branch x <3 fig. 114-018 Aaadeie eo sepal x 2, ype in the U. 8. National Herbarium, collected on rocks » near Tequisistlan, State of Oaxaca, Mexico, altitude 2100 mt ers, pa 1, 1957, by Mulford B. Foster and O. C. Van Hyning (No. 29. HECHTIA sage ae r ex Hemsl. Biol. Centr.-Am. Bot. 3:317. 1884; Bot. Mag. 122: pl. 7460. 1896; Contr. Gray Herb. “17:21. 1937; No. Am. Fl, 19: 95. 1938. Aes argentea K. Koch, Wochénechr, Gartn. 7:176. 1864. omen, MEXICO: Indefinite, 30. HECHTIA CAPITULIGERA Mez in - Monogr. Phan. 9:546. 1896; Contr. Gray Herb. 117: 21, pl. 1, figs. 67, 68. 1937; No. Am. 1961 Smith, Notes on Bromeliaceae 9 Fl. 193 95. 1938. MEXICO: San Lugs Potosi. aie gree CONFUSA L. B, Smith, Contr. Gray Herb. 117:22, pl. 1, ao a 2 12 Pal No. Am. Fl. 19:96. 1938. 32. HECHTIA MARNIER-LAPOSTOLLEI L. B. Smith, Bromel. Soc. Bull. 11358, figs. 1961. MEXICO: Indefinite, 33. HECHTIA GLOMERATA Zucc. Abh. Akad. Milnch, 3:240, pl. 6. 1840; Contr. “Gray aes 117:23, pl. 1, figs. 73, 7k. 1937; No. Am. Das lirion eer ater oe ium Karw. & Zucc. ex Zucc. Allg. Gart. 6:258. 1838, not Hechtia eae Verlot. 1868. Hechtia ghiesbreghtii Lemaire, Dl. Hort. 10: pl. 378. 1863. Hechtia morreniana Mez in DC. Monogr. “cl 9:547. 1896. UNITED STATES: Texas. pe Tamaulipas, San Luis Potosi. GUATEMALA: Quiché, Huehuetenango. 34. HECHTIA GAMOPETALA Mez in DC. Monogr. Phan. 9:549. 1896; Contr. con Herb. 117. 23, 1. 2, figs. 1-3. 1937; No. Am. Fl. 19:97. 1 MEXICO: raiti aia 35. HECHTIA DESMETIANA (Baker) Mez in DC. Monogr. Phan. 9:551. 1896, Contr. Gray Herb. 117:14. 1937; No. Am. Fl. = 86. 1938. Dyckia Gesmetiana Baker, Bot. Mag. 120: pl. Le Bromelia desmetia metiana Baker, Bot. Mag. 120: under or ain. 1894, in he 6 ynonyny. ? 36. HECHTIA ROSEA E. Morr. ex Baker » Handb. Bromel. 140. 1889; Contr. Gray Herb. 117:14. 1937; No. An. Pi, 19: 87. Ae Hechtia roezlii Baker, Handb. Brom. 140. 1889, in synonymy. MEXICO: Indefinite. 37. HECHTIA MEZIANA L. B. Smith, Proc. Am. Acad. 70:149, a Hass 2, 2- 1935; Contr. Gray Herb. 117:14. 1937; No. Am. Fi. 7 MEXICO: Oaxaca, Chiapas. 38. HECHTIA LAXISSIMA L. B. Smith, Contr. U. S. Nat. Herb. 29: ie toate o. aca 39. HECHTIA EPIGYNA aa Notizblatt 12:531. 1935; Contr. Gray Herb. 117:16, pl. 1, figs. 34-37. 1937; No. Ams Fl. 19:89 MEXICO: Tamaulipas. 10 PHYTOLOGIA Vol. 8, no. 1 40. HECHTIA DICHROANTHA Donn. Smith, Bot. Gaz. 42:299. 1906; Contr. Gray Herb. 117:14, pl. 1, figs. 19, 20. 1937; No. Am. Fl. 19:86. 1938. GUATEMALA: Baja Verapaz, Chiquimmla (7). HONDURAS: Morazan. 41. HECHTIA GUATEMALENSIS Mez in Fedde Rep. Nov. Spec. 3:14. 906; Contr. Gray Herb. 117:16, pl. 1, figs. 32, 33. 1937; No. Am. Fl. 19:89. 1938. ATEMALA: El Progreso, Zacapa, Chiquimula, Guatemala. SALVADOR: Achuapan, Salvador. name tification although there are no notes to indicate distinguishing haracters. There is great variation in the size of the bran- ches, density of the flowers, and size and shape of the capsules. However, intergradation is so complete that I hesitate to make even varieties. 2 4 TA REMOTA Wittm. in Engler, Bot. Jahrb. 14, Beibl. 32:6. 1891; Mez in DC. Monogr. Phan. 9:637, 673. 1896. Tillandsia tenuifolia sensu L. B. Smith, No. Am. Fl. 19:133. 1938, in part, as to Tillandsia remota Wittm. In describing Tillandsia remota, Wittmack did not see complete stamens and did not commit himself as to their length, although he surmised that they about equaled the petals. Mez made the ra- ther natural assumption that, like the majority of species in co ral America, it had exserted stamens. Furthermore the habit of T. remota is so close to that of T. tenuifolia that it is difficult to distinguish them without good flowers, Mez's key character of spike position being worthless, Actually the those of T. ten the axes still more slender. Also T. remota has distinctly nerved floral bracts and posterior sepals mo connate, but the spreading pet and inclu , tig. 3) 1/2; fig. 13: Inflorescence (Foster & Van Hyning 2986) x 1; fig. 14: Petal and stamen x 2, hae Lae ILLANDS AT. brachycaulos Schlecht, » cui affinis, foliorum vaginis haud distinctis, laminis latioribus » bracteis florigeris sepalis- ess; leaves numerous, suberect, 16 cm. long, covered with 1961 Smith, Notes on Bromeliaceae pia appressed cinereous scales, reddish; See merging with the blades; blades narrowly triangular cuminate, ca. 2 cm. wide, slightly recurving; scape very radar wholly concealed by the leaves; inflorescence depauperate-compound, few-flowered, capi- tate, much exceeded by the leaves; primary bracts foliaceous but the sepals, carinate, membranaceous, bdensely lepidote with pale brown appressed scales; flowers baseuaiel et sepals free, olle Cerro de Escasu, Province of San José, Costa Rica, August 1935, by Fernando Solis (No. 314). ECUADOR hay ag HARLINGII L. B. Smith, sp. nov. P. cuatrecasana L. B. Smith, cui affinis, inflorescentiae intnazie ferrugineo, sepalis acominatis differt. Known only from one leaf and an inflorescence but flowe wering shoot probably over 1 m. long; leaf linear-lanceolate, acuminate at both ends, oe 15 dm. long, 5 cm. wide, laxly serru- late throughout, glabrous above, covered with pale ferruginous blades; inflorescence simple, 4 ry long, lax, ferruginous-lepi- dote except the petals; floral bracts triangular-ovate, acumi- nate, to 55 mm. long, exceeding the lower pedicels; flowers sade pnts at anthesis, 7 cm. long, seemanewts orange (1 Raxid ling); o y 2/3 inferior; ovules alate. Pl. I, fig. 18: Flower * “1 23 te 19: Sepal x 1; fig. 20: Base of petal x l. Type the Naturhistoriska Riksmuseum, Stockholm, collected in thicket at river, El Topo, Provi nce of Napo-Pastaza, Ecuador, a 1250 meters, November,28, 1958, by Gunnar Harling (No. PITCAIRNIA UNILA IS L. Ab omnibus speciebus cies peeeabia’ a aie laminis lineari- bus integerrimis, inflorescentia simplici dense resin ica ’ width of the blade at base; scape unknown; inflorescence evident- ly simple, 38 cm. long, densely secund-flowered, except for the 12 PHYTOLOGIA Vol. 8, no. 1 petals densely and finely white-arachnoid; axis 7 mm. in diame- ter; floral bracts triangular, to 15 mm. long; pedicels slender, long, pale yellow, bearing a suborbicular crenate scale at base; ovary more than 1/2 inferior; owles caudate. Pl. I, fig. 21: Flower x 1/2; fig. 22: Sepal x 1; fig. 23: Base of petal 3s Type in the — eb io oie Rikemuseum, Stockholm, ary from steep rocks by river, Rio Daule below Pichincha Santa Barbarita, Province of Guayas, Ecuador, April aes, "i96oe by Gunnar Harling (No. 4785). BRAZIL ae om ICATA aye Cam.) L. B. Smith, comb. nov cata Arr Aechmea stephanophora E. Morr. ex Baker, Handb. Bromel. 67. chevelieria ee - Morr, ex Baker) Mez in DC. re rae cated quite a I believe, that Anana ricatus and Chevalieria stephanophora are identical. Frias “situa tion necessitates a new combination in Aechmea to which g eria has been reduced. ANANAS MONSTROSUS (Carr.) L. B. Smith, comb. nov Ananassa monstrosg Carr. Rev. Hortic. 42: 288, fig. 43 B aca Ananas thi go, Arquiv. Jard. Bot « Rio ~~ Janeiro Ih: 281. 1956, without Latin diagnosis. As demonstrated by Camargo, Ananas monstrosa is a perfectly typical pineapple except that it completely lacks the foliaceous apical coma on the inflorescence. This lack of a coma would in- dicate that A. monstrosa is probably the most primitive species in the genus. 1961 Smith, Notes on Bromeliaceae 13 Plate I Pig. 1-5: Hechtia caudata; fig, 6-9: Hechtia lanata; fig, 10, i: Hechtia fosteriana; fig. 12-14: Tillandsia remota; fig. 15-17: Sia abdita; fig. 18-20: Pitcairnia harlingii; fig. 21-23: Pitcairnia unilateralis. NOTES ON NEW AND NOTEWORTHY PLANTS. XXVIII Harold N. Moldenke CITHAREXYLUM ROSEI var. PILOSUM Moldenke, var. ni Haec varietas a forma typica speciei laminis ney po subtus parce ne a pilosis recedit. This variety differs from the typical form of the species in puis of Hacienda Las Rosas, which is about 20 niles south of ientes, at an altitude of 1950 meters, Jalisco, Mexico, on dines 18, 1958, _ is deposited in the herbarium of the Uni- versity of Michigan a ARBOREA f. DENTATA Moldenke, f. n Haec forma a forma typica speciei Tanto foliorum parce Oe. iii cboddac tentatts recedit. rm differs from the typical form of the species siti ' its " Tentobheeien marked with 2 or 3 large lobe-like teeth on the ma The type of “the form was collected by Azizullah Shah a Jhajra, Siwalik & Jaunsar, India, on March 1), 1922, and oe de- posited in the herbarium of the hisvanattey of California at Ber- ey. CONGEA ste Moldenke, sp. n Fru scandens; ramis eracthchink dense fulvo-tomentosis; foliis Sppouitees laminis anguste ellipticis vel elliptico- ——— vel suboblanceolatis acuminatis integris, ad —_— sub- tis, supra nitidis, subtus dense fulvo-pubescentibus; in- peptroiee! Hip axillaribus simplicibus valde elongatis 6--10-cap- itulatis densissime fulvo-tomentosis et parcissime villosis, sympodiis valde elongatis, bracteis involucrantibus 3 3 parvis, s cent in ing, very shiny above, lighter benea th, narrow elliptic or ellip- tic-oblong, sometimes” sone oblanceolate, 5510.5 cm. — wide, acuminat the gins and puberulent midrib densely fulvous-pubescent beneath the hairs longer on the midrib; inflorescence in the upper leaf- axils, Po ye serie 7 ree sr elongate » Widely divergent, with 3-5 pairs of small opposi pEueus emma, about 3.5 cm. 1961 Moldenke, New and Noteworthy Plants 15 long, densely fulvous-tomentose and scattered long-villous; sym podia similar to the peduncles, greatly elongated, .5--7.5 cm. long, with a pair of foliaceous bracts at each node; bracts green, vous- pubescent on both surfaces, short-stipitate, acute at the apex; ar sessile, 3-flowered; *involucral bracts 3, narrow-elliptic, O--1) mm, long, mm, wide, obtuse or © bate at the apex, very neath; calyx campanulate, Seok mm, ay very densely fulvous- villous with long forward-pointing and very conspicuous hairs; corolla undeveloped. The type of this very distinct species was collected by Joseph Francis Charles Rock (no. 1677) — in vee: honor it is named — between Ta Kaw and Meh Soi, Thailand, on January ), 1922, and is deposited in the herbarium of the Chivereity of California at Berkeley. ADDITIONAL NOTES ON THE GENUS AEGIPHILA. XVI Harold N. Moldenke IPHILA Jacq. Additional literature: Aristeguie eta, Clave & Descrip. Fam. Arb. Venez. we apiece 230, & 277. 195k; A. ‘ Schultz, Inst. Tec. Rio Grande di - 30: 15. 1960; Rambo, Pesquisas }: 18. 1960; Angsty, end Bot. he Peg & 37. 1960; Moldenke, Phy- tologia 7: anal & 467—506. Vernacular names as bay ge by pene for sms of this genus in Venezuela are "chicharro" and ob An additional excluded species is: Aegi nila sartian el fame — Alph. List Cit. 2: 445, sphalm. 19,5 = mar- Schau a8 AEGIPHILA ACULEIFERA Moldenke Austin Smith adds the following notes about this species: tree with base 1 m. wide; limbs long, lateral; bark dark-brown; leaves membranous, rather fleshy, dark-green, ning, very glabrous, the venation reticulated, the midrib Tight preah, buds round or flat-t topped, pure white; calyx and ovary pale- green; pedicels darker green; petals opening flat or slightly reflexed, 8 m - long, pure white; stamens and pistil long- exserted. He says that it is a showy tree when in bloamm, inhab- iting wet loam soil in semi-shade of the subtropical zone. Additional citations: COSTA RICA: Alajuela: A. Smith P.2035 (Ca~-12690) , PHILA AUSTRALIS Moldenke Additional literature: A. R. Schultz, Inst. Tec. Rio Grande 16 PHYTOLOGIA Vol. 8, no. 1 do Sul Pub. 30: 15. 1960. Schultz, in the reference cited above, reduces this species with a ney ae to A. integrifolia (Jacq.) Jacks. AEGIPHILA BOGOTENSIS (Spreng.) Moldenke Additional citations: ECUADOR: Tunguragua: Asplund 7980 (S). IPHILA BRACHIATA Vell. eet synonymy: Aegiphila aes at apud A. Re Schultz, Inst. Tec. Rio Grande do Sul Pub, 15, sphalm. 1960. Additional scearevaeee Rambo, Pesquisas bs ‘By 960; A. Re Schultz, Inst. T Grande do Sul Pub 15. pre rat Molden- ke, Phytologia 7: a. 1961; Angely, Fl. — 17: 46. 1961. AEGIPHILA CANDELABRUM Brigq. Additional citations: PARAGUAY: Woolston 844 (Ca—86),05). AEGIPHILA CORDATA Poepp. Woytkowski describes this species as a tree 12 meters tall, with yellow-green flowers in September, and says that it is rare in forests at 1000 meters altitude. Additional citations: PERU: Junin: Woytkowski 5977 (Z). AEGIPHILA COSTARICENSIS Moldenke ‘i Se ae citations: MEXICO: Chiapas: Matuda 2101 (Ca— AEGIPHILA CRENATA Moldenke Additional literature: Angely, Fl. Paran. 17: 6. 1961; Mol- denke, Phytologia 7: 464. 1961, aaa t eg erotik hissy Fl. Paran. 17: 46. 19613; Mol- denke, Phytolagte 7: 72, loele” , oo HASSLERI Briq. tional literature: - Paran., 17: 46. 1961; Mol- denke, Phytologia 7: helt. tae : AEGIPHILA LHOTZKIANA Cham. Additional literature: Angely, Fl. Paran. 17: 6. 1961; Mol- denke, Phytologia 7: 48h. 1961. . AEGIPHILA MEMBRANACEA Turoz, Steyermark & Nilsson describe this plant as a vine, with chartaceous leaves, the peduncl es, pedicels, and calyx pale- green, the corolla pale~yellow OW, the style whitish, and the two tional citations: VENEZUELA a son 193 (2). ae aapemmsid'< & Nils 1961 Moldenke, Notes on Aegiphila 17 eee OBDUCTA Vell. tional literature: Angely, Fl. Paran. 17: 46. 1961; Molden ke, "waysaacse 7: 492. 1961. AEGIPHILA PARAGUARIENSIS Brigq. Additional literature: Angely, ke, Phytologia 7: 9l--95. 1961. AEGIPHILA S area Re Additional litera Angely, Fl. Paran. 17: 6. 1961; Molden- ke, Phytologia 7: eo E02, 1961. Fl. Paran. 17: 6. 1961; Molden- AEGIPHILA TRUNCATA Moldenke Additional literature: Moldenke, Known Geogr. Distrib. Verben~ » [ed. 2], 59 & 175. 199; Voldenke, Phytologia h: 41S. rigs Yoldenke, Résumé 65 & 42. 1959; Moldenke, Phytologia 7: 506. Me tealf & Cuatrecasas report that this ’ species is a tre ey . tall, the fruits green with brow spots in February, Piuhd at al- titudes of 20 to 260 meters. Additional citations: COLOMBIA: Antioquia: Metcalf & Cuatre- casas 30072 (Ca--68722)). AEGIPHILA sebetiarnaeete Mo. rye terature: Molde Phytologia 5: 225. 1955; Moldenke, Biol. Abstr. 30: 39). 1958; “foldenke, Résumé 82 & 42. 1959. Ferreyra describes the flowers of this slant as purple outside, sy reir pinki Cita : PERU: Hudmco: F 2190 (W—2026718); Stork & Horton 9198 (Ca—6h.7102—-type = Card. Ind. Gray Herb. 1903; Prain, Ind. Kew. Suppl. 3: h. 1908} Moldenke, Brittonia 1: 25h, 264, 360-—-362, & et 193k; Molden- ke, Ghicaies 1: 271-272. 1935 Moldenke, et ot Distrib. Avi- cenn. 6. 1939; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 25 & 85. 19125 ernest Alph. List Cit. 1: 28. & 303. 1946; Moldenke, Known Distrib. Verbenac., [ed. 2], 46 & 175. 19,9; Moldenke, is 5h & Lhe. 1959. AEGIPHILA VALERII Standl Additional and enended literature: Moldenke, Brittonia 1: 252, 258, 263, 289--292, & 476. 19343 Moldenke, Yeogr. seat Avi- cenn. 13 & 17. a Moldenke, Known Geogr. Distrib, Verbenac., e bs, 1942; Moldenke, Alph. List Cit. 1: 325& 326 (1946) and dad 2s ‘uz 29104 Be He & he by ldenke, Pl. Life 2: 87. 1948; Moldenke, Alph. Cit. 3: 7h, pls & 962 (1949) a: 4: 1051-1053 & 1082. 19193 Moldenke, Known Geogr. Distrib. » (ed. 2], 28, = & 175. 1907) Mg mo wh Phytologia ]: Is. + 19535 Moldenke, Résum & 3h, 46, ustrations: Moldenke, Phytol ate ‘Dg, f e 5. 198. peered citations: MEXICO: Chiapas: Matuda 11876 76 (W— 18 PHYTOLOGIA Vol. 8, no. 1 2087069) . PHILA VALLENSIS Moldenke Additional iach ee Alph. List ae ks: 1214. 199; Moldenke, Known Geogr. enac. eos elena, Paytclogis Sa "ase 1953; Soldecks ; "Résumé 65 "& 1959 ro es Litera Je lark, ray Herb. 1932; ari ae Brittonia 17 19h (1332) and 1: 2 io3, 126, 437-436, & 1 Moldenke, Phytologia 1: 272. 1937; Moldenke, ae er = ed. lj, 5. (ash) and i 1032. 1948; Moldenke, Known Geogr. Distrib. Verb- » [ed. 2], 71 & 175. 199; Moldenke, Résumé 82 & 2. 1959. 975 & 76. ° “Tol ; Moldenks. Kn er. strib. erbenac., (ed. 2], 62 & 175. 1949; Moldenke in $ k, Fieldiana Sly 515. 195 enke, Phytologia : 415, 1953; Steyermark, Field 3 Mo 28: 1178. 19573 Moldenke, Résumé 70 & 2. 1959 Collectors describe this plant as a shrub or slender shrub, 2.55 m. tall, or a small tree to 12m. tall, the slender stem about 1 cm, in’ pes eter, the leaves firmly menbranaceous, dark- or deep-green and shiny above ve, paler green beneath, the stems, petioles, bg peduncles appressed buff-tomentose, the rachis 8S gray-green, the calyx pale-green and appressed lobes mm. long and 2.5 mm, wide, the filaments white, the an- thers buff-brown, and the fruit turning yellow, It is said to be occasional in scrub forests and quebradas, in upper mixed Clu- sia-Magnolia forests, and along streambeds, at altitudes way 1600 to 2300 meters, blooming in Jamary, Febru ruary, and Jun and fruiting in December and January. It has been Aisidenti fied in herbaria as a species of pepboopeds » Additional ig grocer oar ee Cowan & 31298 (N); Maguire, st Wardack 30042 (N). Bolfvar: 325k 39, sak 0), 75664, (N), 75835 (Z); mgzize Jobo ; “Siegert see (N) 75838 ¢ j UELENSIS Literature: a P a e wits & he noe . weiytatnete, a 323 353, 1958) 3 Moldenke, R& llectors report that this plant is a shrub with white flow- = occasional in talus forests, at altitudes of 1300 to 1500 rs, blooming in January. The type was collected in an upper- Citations: onas: Maguire, Wurdack 36952 (N-type); agatine: Wurdack, & Bunting 12582 ‘ny 1961 Moldenke, Notes on Aegiphila 19 AEGIPHILA VERRUCOSA Schau. Literature: Roem. & Schult., Syst. Veg., ed. 1, h: 698 (1819) and ed. 2, 2: 82, 1820; Schau. im A. DC., Prodr. i: 650. 1847; Karst., Ausw. New. Schtn. Blttt. Gew. Venez. 31—33, pl. 10. 1848; Bocq., Adansonia 3: 188, 1862; penth. in Benth. & Hook. f., Gen. Fis 23 1151. 1876; Jacks., Ind. eat ls 7 & 3u5. 1893 93; Briq. in Engl. = Prentl Nat. Pflanzenfam. ) (3a): 166. 18935 Pittier, PL 6. 1926; Moldenke, Brittonia 1: 258--260, 263, 285-289, Le, 47h, & 475. 1934; Moldenke, Phytologia ct 272. 1937; Moldenke , Alph. List Common Names 8. 1939; Moldenke, Geogr. i Invalid A. L. Moldenke, Pl. Life 2: 2 ‘ » 29d ldenke Bye Distrib. Verbenac., [ed. 2], & 175. “T9h9 93 Nol Alph. List Cit. 3: 69h, 19 819, “fe “oth (1949) a ks 1038 & 1049. 1949; Moldenke, Résumé 65,70, 239, 2h0, & Whe. fod Tllustr. trations: Karst., Ausw. Beni 86 — "tome. Gew, Venez. pl. 10. 1848, The wood of this species is reported to be white, and the ies is said to grow at 200 meters altitude. It has been misidentified in herbaria as Metteniusa sp Additional citations: VENEZUELA: aerid, Bernardi 6087 (N). AEGIPHILA VERTICILLATA Vell. Additional synonymy: Aegiphila veryicillata Vell. ex Molden- oo Alph, List Cit. 1: 171, sphalm (19. and 3: 670, sphalm. 9 Addi tional and emended ens Jacks., Ind. Kew. 1: 7. e Moldenke aie eae ad a: aan vetle ie 1 age 06° ; eee a 2: 343, "40° 3 7 243 535, 561, an, "ie, 2'933 nats 3: 663, 670, 67h, 675, 690, 692; 710, 711, 751, 81h, 816, 838, SLh—bh6 858, 890, 915, 921, & 922 (1949), and ti 1046, 1058, 1081, uf 1106, 1123, Lik, & 1301. 19h9; prea Known Geogr. 9 peat er [ed Py 75 ry 9 r & 175. 19. ne Tease beh logia h: 434. 19533; Moldenke » 229, 2 28h, & l2. 1959; angely, Ind. Ang. 10. 19595 Renn, Levant. rhe Inst. Agron. Minas 149. 1960; Angely, Fl. Paran, 16: 3. onal citations: BRAZIL: Minas Gerais: Macedo 43h (S). Paran&: Hatschbach 3757 (Z).» S#o Paulo: Brade 13095 [Herb. 20 PHYTOLOGIA Vol. 8, no. 1 Jard, Bot. Rio Jan. 29513] (B). ARGIPHILA VILIOSs Lan ) Gmel. onal and emended liter, prec svecitrs Ind. Kew. 1: 7 (1893) and 2: 160. “189k; Moldenke, Brittonia 1: 259, Oey 268, om, 339, gieeraies & h7h--76. 13343 Moldenke, Alph. List Common ames 6, 22, & 33. 1939) Moldenke, Geogr. yo Pris ryt 21& 2 1 1939; Moldenke, Prelim. Alph. Hep Invalid N 26 1910; Moldenke in Pulle, Fl. Surin. (2): 30h. "19b0} Moldenke, Alph, List Invalid Names at & a ae aE Re coee Geogr. : 15, 37 (19 62h, 631, & 60 eae, : 655 709, 72s, 6, 8s, 853, '& 608 (19h9), and 4: 106 & 4097, 19195 Mo , Known Geogr. Distrib. ; wd oagerh ue sls 65, or %, & 5. BrER He Ne & Ae Le Mol- , Anal, » Bio » 20: 2. 1 9493 Moldenke, Phytologia be 13 . 13-37, 1953; Mol danke, Résumé 7h, 78, 86, 231, 319, & Ba 9 reports that this is a tree 6 meters tall, fruiting in Novenber, found in the primary forest at altitudes of 70 to 300 meters dditional citations: BRAZIL: Amap4: Cowan 38260 (N); Frées & Black 27481 (2). eats, 037% AEGIPHILA A VILLOSISSIMA Moldenke Literatu 1 Mold e, Bull, Torrey Bot. Club 601 392. 19335 ttonia 1: 296. 1938; ol asnike. Geogr, Distrib, 25. 1939; Mol- « Distrib. Verbenac. & 85. 192; H. N. & A. L. Moldenke, Pl. Life 2: 6. 198; "Eats List Cit, 2: 621, 198; Moldenke, Geogr. Distrib. Verben- Known aCe, [ed, 2 2], 5 & 175. 19,9; Moldenke, Résumé 86 & 2. 1959; ely, Ind. "ang. 10, 1959. mae Fs te engin Seat by s doubtfully valid in 2 eel ener 1s ey & “YT (93h), Geogr. “Distrib. Avicenn, 25 (1939), and ph. List Cit. 3: 737 (1949), has proved to belong in the syn- onymy of Arrabidaea corchorioides (Cham.) P. DC. in the Bignon- IPHILA VITELLINIFLORA Klotzsch Additional and emended literature: pa » Ind. Kew. Ps Py: & 7. wey, Moldenke, Brittonia 1: 259, 279, 21 426, L776. 193k; Moldenke, Alph. List Common Names 7 & 2 19395 Moldenke, Geogr. Distrib, et i 28, & 35. 1939; Moldenke, Pr ° Alph, List Invalid Name & 2h. 190; Moldenke, : ke, Alph, List » ok, 78, 107, 114, 135, 226, 236, 237, 263, 26h 28, "3257 & 326 (i9h6)? 2: 529, 331, 1961 Moldenke, Notes on Aegiphila 21 » 361, 41h, 432, 485, a ‘ geet Pee 3: 68h, 726, 736° 801, 814, 838, 856, ; 920, & 922 2 sks), eer h: 982, 1086, 1106, 1113, & it 719h9y as ldenke, . Dis- trib. Verbenac., [ed. 245 755905 96, Las & ets, chi Voldenke, Phytologia l: 37—38. 1953; Moldenke, 86, 113, 115, 212, ay Porn alles pees & Lh. 1959; Angely, _— ao 10. A704 « tations: BRAZIL: Bahia: Blanchet 904/129 (N), eeoats oe vt eerie Pires 3662 (Z). AEGIPHILA WIGANDIOIDES Lundell Additional and emended literature: Moldenke, Alph. List Gide 3: 715. 1919 Moldenke, Known Geogr. Distrib. Verbenac., [ed. 28 ck ree 1995 MeL, Phytologia : 438. 1953; Moldenke, abe & Whe. 1959. dditional citations: MEXICO: Chiapas: Matuda 4253 (Mi—type, W—1B913I1-—iectype), ADDITIONAL NOTES ON THE GENUS VITEX. I Harold N. Moldenke These nadia are supplementary a ate information Se in my monograph the genus as publish reliminary f in Phy tologia 5: eer --176, a oe7-= 280" 293-336, 313-393, Oh—h64, & 465-507 and 6: 13—6h, 70—128, 129—-192, & 197—— 231 in Sy period of ory to 1958. VITEX Tourn Additi tdonal synonymy: Vitee L. ex Moldenke, Alph. list Cit. 3: 89, ry 199. Addi and emended literature: A. Cunn., Precur,. no. 390, OVe App. 160. 1880; Forbes, Nat. Wand, East. oat a ona Campbell & Watt, Descript. Cat. — Prod. & 55. 1886; Moloney, Sketch Forest. W. Afr 623 & & Sohe “1887; core & pected Fl. Kais. Wilhelmsland 121. 1889; Durand & , Bull. - Roy. Bot. Belg. Compt. Rend. 38: 133-134. 1899; Warb, in Seeii Bot. Jahrb, 13: 428. 1891; K. Scum. & Lauterb., Fl. Deutsch. p Ste: “— 523-5 oh, 1900; Cheesen, » Man. New ¢ Fis, » 565. 1906; Laing & & Black- 22 PS PTeOL0 OLA Vol, 8, no. 1 well, Pl. New Zeal., ed. 1, 350--351, fig. 11). 1906; Valet., Bull. Dept. a Ind. Néerl. 10: S153, 1907; Sim, For. Fl. Cape Colony 2 287, pl. 120. 1907; Pulle in Loren rentz, Nova Guin- ea 8 (4): i 1912; 0 ee Distrib. V Veg. Fl. New Zeal. 1). 1919; Bri del, Bull. Soc. C e Biol. 33 1226 19.23; E. De Merr., Philipp. Flow, Pl. 3: sab a 1923; Setchell, Carnegie Inst. Wash. Publ. 31 (Dept. Marine Biol. 20]: 55. 192h5 Cheesem., Man. New Z e Fl., ed 9 6 65 925; ry ° New Zeal., ed. 3, 354-356, fig. 127. 1927; Crevost & P&telot, Bull. Econom. In 39 7. 19343 Moldenke, Brief Course Syst. Bot. Lect. 1, rev., 5. 1938; Moldenke, Brief Course Syst. Bot., ed. 2, 7. 1939; Oppenheimer & Evenari, Bull. Soc. Bot. 31: 0; Laing & 2; Pl. New Zeal., ed. h, Genev. 363. hed Blackwe 372—373, fig. 139. 190; Doney, Brooklyn Bot. Gard. Rec. 30: 23. 1941; A. R. Schultz, Introd. Estud. Bot. Sist., ed. 2, 8h, 519, & 562. 1943; E. L. D. Seymour, New Gard. Encycl. 1292. 19h; Karrer & Schmid, Helv. Chim. Act. 29: £25, 1965 Holdridge & al., Forests West & Cent. Ecuador 23, 25, 6, & 67. "19473 Mol- ae Alph. List eps 3: 849. 1949; Sevim, Orman Fakult. berets si & oth, Deutsch, Med. Wschr,. 79: 1271. 19 : & Descript. Fan. Arb. Venez. 192, 193, 209, 52, 36, & ° 19543 Hansford, R 3: 75—-112. 1954 eda, Caldas- ia 7: 49--50. 1955; Cavaco, Bull. Mus. Nat. Hist. Nat. Pari sér. 2, 27: 91. ; » Journ. Jap. Forest. Soc. 37: 334-—- 335. a 53 eames Phytologia 5: 343-3 LOh—6,. 1956; : 260. 1956; e ° Cavaco, 293 2& 3630. 1957; Van Steenis, Fl. Males. Bull. 2: 302, h7h, 529, & 19 tapau, Fl th Pac Ste Phytologia 5: 465—507 (1957), 6: 13—6h & 70—~128 (1957), and ¥- 129—192, 197—231, & 332. 1958; as — Tas. Fil. Afr. Trop. Index te 61. 1958; Cavaco « Broter, 13: 1h. 1958; Biol. tr. 30: fn ha & 1398. 1908; Alain, Revist. Soc. Cub. Bot. 15: ite 1958; Moldenke, Am. Midl. Nat. 59: Ange 19585 Lombardo, Arb, Cult. Pas, Pub, 255—256,. 1958; Brizicky, S tern, & Cham- bers, Trop. Woods 109: 79. 1958; Assoc. Etud. Tax. Fl. Afr. " India Res.] 9: 233, 248, 250, 251, 276, 285, 302, 309, 319. 321 2 355, 356, 379, > 395—399, 109; ih? & hth. 15595 loidetce, £ Résumé Suppl. 1: 5, 713, 15, 23, & 25, 1959; Reitz, Sellowia 1961 Moldenke, Notes on Vitex 23 ae List a Trees [1 0}. 1960; Assoc. Bea Tax Ange Fl 2h. 1960; Angely, Liv. Gen. Bot. Bras. 35 & 58. Hae ely, Fl. Paran. 17: ils 1961; Menninger, Trop. Tree Seeds [2]. 1961; Tel- Aviv Univ. Ind . Sem 1960: 3. 1961; Kelsey Nursery Service, Short Guide Cat. 167: 33. 1961. Additional excluded species: Vitex americana L, = Buddleja am- ericana L., Loganiaceae; —-- lanceolata Turez. = Psychotria sarmentosa Blume, Rubiacea This is a diverse genus cap about 380 known specific and sub- specific entities, mostly native to the tropics and subtropics of both the Eastern and Western Hemispheres; a few also f in the temperate portions of Europe, Asia, and South Africa. At least 56 are cultivated, of which the Mediterranean V. agms-castus and the yelcrrp Vv. negundo are the most commonly seen, each exist- ing in several popular horticultural varieties. sees occasionally escape i cultivation and become naturalized in areas far re- moved from their original home. Five species are know in the fet o vest Pinta from Cretaceous to Recent formations in Europe ame: The generic name is derived from the Latin, vieo, to bind, as plants. Members of the gemus are now commonly yeferred to in Eng- lish ste-trees", in French as "gattilier", in German "Keuschbaum", and throughout most of South America as — Aristeguicta, in the reference cited above, lists the commo names “aceituno", "escobillo", "guarataro", and "totumillo" an members of the genus in Venezu Menbers of the group are most ly trees or shrubs, very rarely woody vines, with opposite or pe gel aoret palmately com- pound leaves of 1—7 leaflets, and c e, axillary or terminal, usually racemiform or thyrsoid to late aittuse panicles of blue, Violet, yellowish, or white mint-like flowers, rarely solitary or contracted to form heads, An Amazonian species is said to grow to a height of 200 feet, Certaun species have been and are still & more or less dome-shaped upper lip, is sometimes segregated as the separate genus Chrysomallum, On the other hand, the Asiatic genus Pseud idium were formerly included in Vitex. The Hinton EE and Loveless 277, both distributed as species 2h PHYTOLOGIA Vol. 8, no. 1 of Vitex, are not verbenaceous, while Peter 13705, distributed as Vitex gomphophylla Baker, is actually Combretum brosigianum Engl. & Diels. in the Combretaceae. VITEX ACUMINATA R. Br. pa is described by Mrs. Clemens as a medium-sized tree, grow- ing am mangroves on the bank of a canal. It has been collected in antheale in December and in fruit in December and February rate Fe ia te have sometimes been misidemtified as V. eas ra ° ~~"Kddi tional “ oitits bed AUSTRALIA: Queensland: M.S. Clemens s. n. [1 December 196] (Mi), son. [1 December 197] (Ca—81173), s s. ne [9 November 1918] (Mi); Mrs. Re o L. Higgins s.n, [near Rock- hampton, 2/1922] (Ca--226265); Kajewski s.n. ({Gympie, Decr. 1922] (Ca—31)11); C. T. White 1423 (Ca—226203). VITEX AGNUS-CASTUS L. Additional synonymy: Vitee agmus-castus L, ex Moldenke, Alph. List Cit. 3: 849, sphalm. 19]9. Addit eer onal literature: Griseb., Abhand. Ktnig. Gesell. Wissen. GUtting. el: 1857; Loret & Barrandon, Fl. Montpel. 532—533 & 918. 18%; Bridel, Bull. Soc. Chim, Biol. 3: 722. 19233 Molden- ke, Brief Course Syst. Bot. Lect. 1, rev., 5. 1938; Moldenke, Brief Course Syst. Bot., ed. % Te 19395 ens Brookiyn & Bot « 3 igshis iol oe 150, & 186-196 (955) 363° & al 1986 stdene Shean tr. 30: 170 19565 Moldenke, Phytologia 5: 165, Usa : & 50h (1957) af 6: 17. 1957; Moldenke e, i: Midl, Nat. 59: 365. 19583 Price, New Arz 8, 10—12, 1h, en. ie oP 77, lu, 128-132, 131, Be om 165, 172, is, 225, 232,28), 335, 379— 9, 420, 426, & denk otal Shih.’ 1s 6, Sos8, BPEDe et eeen The species is said to grow tat roadsides at Macon, Geor and in talus in Spain, Waby found to be a tree in sa Turner's Hall woods Barbados, =e nisidentitiea it as a species of Inga (Mimosaceae}, Dempster says that it grow hilly country in Epirus, while Balls found it on dark oie anic sandy oil by seashores in Turkey, Allard 11391, cited below, bears no indsdetion on its label that it is from a cultivated plant, but I am assuming that it is. McAfee found the species growing in 8 in Oklahoma spreading by underground runners, the inflorescences 6—~12 in- a ae ee a ee 1961 Moldenke, Notes on Vitex 25 ches long, flowering in August, the flowers "in shades of pale- lilae or Javender-blue, each bush a separate color," Dempster says that the flowers vary from "dark-purple to nearly white", Allard calls them "purplish" or "blue-purple", while Chiao de- scribes them as re te with offensive odor", Allard found the plant blossoming i The Stefani ae hg Mai 1903] from Corsica, distributed as this species in the herbarium of the pccmemr ged of ees is not verbenaceous —- it probably represents ase posed labels. The Mori atte [Agosto 1933] (Ew) cited = me on page 175 of Phytologia, volume 5 (1955) as from Italy, is actu- ally from a cultivated plant and should be shifted to the "CULTI- VATED" section of the citations. Hinmsel & Winde, in the refer- ence cited above, describe a new glycoside, which they call “agnosid" or "agnoside", from Vitex agnus-castus. It is said to be a pseudoindicane and possesses the formula Co5H9¢0)], with a molecular weight of 166.3. In the crystalline form it melts at 146° C. It is the ester of p-oxy-benzoic acid with aucubine. Additional citations: GEORGIA: Bibb Co.: Biltmore Herb. 1786a (S). Decatur Co.: Curtiss 6813, in part (Ur). FLORIDA: Santa Rosa Co.: Curtiss 6813, in part (Ur). ALABAMA: Lee Co.: Earle & Baker s.n. , [Auburn, , 10/9/1898] (Dt). OKLAHOMA: Payne Co.:McAfee 85 (Ca—862863); w. L. Spears s.n. [July 10, 1935] (Um—179). TEXAS: Travis Co.: ? Harpin, Waldorf, & Barkley 13081 (Ca~~-882862, Ok). WINDWARD ISLANDS: Barbados: Waby 105 (Bm). FRANCE: Albert sen. [Magnier 1261] (Um—177); Autheman 51,67 (Um—178); Despaty son. (Duffour 1871] (Vi); Hérail s.n. 1 s.n. (Um—180); Monnier s.n. 11832] (vi); F. Schultz a (B); Théoeneau s.n, [Juli 1 (B); Tholin s.n, [22 Julii '82] (B). SP, csr Sennen 429 (S), s.n. (Duffour 1871 bis] (Um—176). GREECE: Cyrén s.n. [Agrinion, 15/ oom. Go); L. Dempster 2467 (Ca—179508); 8.n. 8.190] ‘s) sn. [23.8.190)] (B); Heldreich 599 (B); Mattfeld 2035 (Ca—3l7616), ABGEAN ISLANDS: Sanos: B. Brown 9. [June & July, 192h] (Ca—986781). ITALY: Bigo sen. [16/8/1871] (B). CYPRUS: H. Lindberg s.n. [22.7.1939] (Go). SICILY: H. Ross 865 (B). CORSICA: 0. Debeaux 62 (B). CRETE: E. H. L. Krause 16803 (B); E. Reverchon 185 (B). JUGOSLAVIA: Dalmatia: M. singe (B), 373 (B)3 0. Krebs gen. [Junio 1901] (B). Fiume: Noé 138 (B). Herzegovinia: Engelhardt s.n. [31/VIII/1900] wag Istria: elhardt s.n. ra/SPGOS ) (B). fr: Trieste: Hrouven s.n. [1h/7/ 1889] (B); Vatoda s.n. [9.VIII.1921] (Um—171, =171, Vi). B arGULGARTA: (Ca—79895) ; Bornmtfller 1236 (B, B); Callier 171 (B, » B)3 Hal- &csy 1,37 (B)} B. Post son. [Aug. 11, 1935] (Dt). ISRAEL: Field & Lazar 233 (Ca—583310, -“Ga--620197). CULTIVATED: Brasil: G. Ae 26 P HIT0LO0G612 4 Vol. 8, now 1 Black 18-3635 (Ca—72808). California: W. R. Dudley s.n. [July 1895] (Du--362565). China: Chiao 52 [Herb. Univ. Nanking 1076] (Ca—-281054). Florida: McFarlin 5571 (Mi). Germany: Baenitz s.n. (15.9-1910] (B); Rettig s.n. [Koehne 321] (B). Kansas: Gates 15025 (Um—167). North Carolina: Biltmore Herb. 1786 [July 27th] (S), 1786 [September 27th] (S). Oklahoma: Felkner 10 10 Sas al ". Vv. He H.* 93 (Ok); Hopkins, Nelson, & Nelson 654 (0 (Ok). Texas: Knobloch 187 (Ok); Tharp s.n. , (Austin, 8/20/41] ( (Ca-~882866, Vi). Virginia: H. A. Allard 11391 (Ca~793723), 20670 (We). VITEX AGNUS-CASTUS f. “en P oadloh ) Rehd, hae cheetinaee synonymy: Vitex agnus-castus f, albiflorus Carre, in non tonal “i = Sees ye Bot. aoaee - 30: 23. “toll; » New Gard. cycl» 292. 19h Molsees = pon Ri * sont: 1s a tional citations: eaius it “Peter 45229 [V.362] (B). ITALY: Frank s.n. (Messina, Julio 1882] (B). VITEX AGNUS-CASTUS f. LATIFOLIA (Mill.) Rehd. Additional FA gl bnigtad nds Brooklyn “6 Ee ard, Rec. 30: 2 Seymour, Encycl. 1292. 19h; Moldenke, Phytologia 5: 190—193, 195 é & 289 (ise pearance 5: 553 (1956), and 5: 465. 1957; Lombardo, Arb. - 7? & 255. 1958; Molden- ‘ 130—132, 158, 25, “307, "352, "379, 380, 385—387, — 427, & 475. 1959; Menninger, 1960 Price List t Flow, Trees [10 1960; Kelsey Nurs Service, Short Guide Cat. 167: 336 1961, ca tional illustrations: Lombardo, Arb. Cult. Pas. Pub. 255. The Sciulz & Schulz s.n. specimen cited below has most of its » With showy blue flowers from July to frost » growing well in sun and in acid soil, called "chaste tree™s 18—2))-inch piants 6s (B). CULTIVATED: Germany: Bornmffller s er 8.n. (18782 1882] (B re Schuls & Schulz s.n. [2).9.1897] (B). Jamaica: G. R. Proctor 21205 (Bm). New York: He N. Moldenke 21251 (Bs, “Mm, Ok, Sm), 21572 (Sm). Virginia: He rv Allard 1. 11391 ( (Gg—381232) . A 4 ehd, Additional synonymy: Vitex astus var, rosea Doney, Brooklyn Bot. Gard, Rec. 30: 23, 19d, Additional literature: Doney rooklyn Bot. Gard. Rec. 30: 23. 191; Moldenke, Phytologia 5: oe 196. 19855 “Woldenke, Résumé hy ieee 386, "26, & nis. 1959; Moldenke, Résume Suppl. 1: 8 & 1961 Moldenke, Notes on Vitex 27 dditional citations: JUGOSLAVIA: Dalmatia: Brunnelle s.n. [5/ 7/1886) (B). VITEX AGNUS-CASTUS var. CAERULEA sept Pe gles pene literature: Moldenke, Phytologia 5: 190. 1955; Mol- Résuné 2h, 30, 120, 131, 228, 379, 380, 383, 126, & hs. 1999; *Moldenke, R é sum Suppl. « 1959. The Heldreich 599 sexinasions., cited below is inscribed snp ah bus coeruleis et carneis" so may represent a mixture of this iety and of f. rosea Rehd, Additional citations: GREECE: Heldreich 599 a ede Dalmatia: Brunnelle 2014 (B). CULTIVATED: North Carolina: P. 0. Schallert 351 (Ok), Son. sen. [June 20, 1941] (G2--882865) VITEX AGNUS-CASTUS var. Additional literature: Moldenke, prying "Ss 1955; Molde Nat. 59: 36 aides: Ré- sumé 132, 158, 159, 379, 38h, 388, 427, & 475. 19595 Moldenke, Résumé Suppl. 1: 11. 1959 » Rec. 1959; Tel-A Ind. Sem. 1960: 3. 1961. that V. hausknechtii Bormm. is identical with this variety. Additional citations: UNION OF SOCIALIST SOVIET REPUBLICS Turkmanskaya: Bornmttller 917 (B). JORDAN: Kneucker 4,30 (B). TRAQ: Field & Lasar 735 (Ca--620082), 818 (Ca~-620076). IRAN: Bornmtfl- ler 5128 (B); Gauba 902 (B); Rechinger & Rechinger 5778 (Ca— 27h0); T. Strauss s.n. [Juli 1897) “3897) (B). AFGHANISTAN: N: Gabriel & Gabriel 3 (B). BALUCHISTAN: Rechinger, Rechinger, Aellen, & Es- a 3996 (Ca——l2739). PAKISTAN: Sind: Stapf s.n. [13/VII/ ) (B) VITEX AGNUS-CASTUS var. SERRATA Moldenke Additional literature: Moldenk Sachem. 5: 196. 1955; Moldenke, Résuné 225, 232, 365, & “h7s. Petry specimen cited below shows ha “serrate leaflets. Adaiti onal citations: CULTIVATED: France: Petry s.n. [Septbr. 1900] (B). AJUGAEFLORA Dop Additional literature: Moldenke poveee 5: 353. 1956; Moldenke, Résumé 1Tt, epee a & & 5. nen oe eyes es this as a meters tall, flower- ing in eae apked Pie prt rt seams to be very oleuene pointed to V. pedun duncularis Wall. and may Sa be con- specific with it. It has been collected in fruit Additional citations: INDOCHINA: Annam: Poilane 2 68, (B~ cotype). Cochinchina: Pierre 24 [Bois 2h] (B), B), 1509 Y (Ca—514869— cotype), s.n. [Gia Pau me, 9/1865] (Ca——54870), s.n. [10/1866] be oR: Bot lane 4076 (B). CULTIVATED: Indochina: Pierre 1509 (B— 28 PHYTOLOGIA Vol. 8, no. 1 VITEX ALTISSIMA L. f. Ee gee cepa cau oe score & & Watt, rnp AF Cat. Econom. Nagpur 55, ae coe Phytol gia 5: 197—202 3s), 5: 2 Bees 5 b, 16s (1957), and "es 13. 1957; Mol- denke , 188, 191, 202, 225, 319, 380, 361, 36, "sat, & his. Campbell & Watt, rain eitaioaie cited above, record the ver- nacular names "bhad They state that the wood is The Menninger material cited below illustrates the dimorphic character of the leaves of this species. The flowering branches exhibit hardly any wings on the petioles, but the non-flowering branches have conspicuous wings, Menninger penornt that the two kinds of branches grow side by side on the The Herb, Lemmon s.n. [Mangalor, Canara] tb the herbarium of the wesvensty (oe 2 of California, determined as this species, is not verbenaceou Additional citations: PAKISTAN: East Bengal: Griffith 6066 ("Hk. and Bth. no. 11") (Ut--11518). INDIA: Bombay: Hohenacker 115 (B), s s.n. [Prope sien Mangalor] (B). Madras: Yeshoda 18h (B). State undetermined: Re Wight 2325 (B). CULTIVATED: Florida: M Men- ninger s.n. [Moldenke | aaa (Bs, Ok). ALTMANNI Moldenk Additional literature: Moldenke » Phytologia 5: 203. 1955; Mol- denke, —— 191, 196, & 475. 1959. This p ant has’ been’ misidentified in herbaria as V. parviflora a " (pdittonel citations: PHILIPPINE ISLANDS: Luzon: Loher S.n. [Rizal Province] (Ca—229199) . err Pos go fe 5s Additional literatur eae | P lo 20 19 —— —— 5 & & Ls. 1959. n bers ene Serer rs describe this species as a tree, 8 feet tall, with a single » rn and mauve aromatic flowers, bloo Oct ober, roups in rich brown soil in thickets at altitudes of 33 to $00 1 i Tanner reports the common name name "mnegeg says that the roots are boiled and the resulting liquid drunk in the treatment of sharp pains in the stomach, It has been misiden- tified in herbaria as V. amboniensis Gtirke, me eee TANGANYIKA: A. Peter 3659 [0.1.105] (B, Z); Tanner VITEX AMBONIENSIS Gturke Additional literature: Moldenk 1 6) and 3 : igs 1957; Moldenke, Résuaé 3. Me iis ack: Oafe ie Peter describes this plant as a shrub 3 meters tall or a tree 1961 Moldenke, Notes on Vitex 29 6 omatere tall, with on nahh? paonnintne in pgs He found i t+ in February, and June, growing at altitudes Pg pe to 1180 meters. The Tanner 3315 distributed as this species is V. amaniensis Pi Additional citations : TANGANYIKA: Engler sen. [7.II.1905] (B); A. Peter 21267 [0.117.202] (B), 22531 [0.IV.255] (B), 23015 [0.IV. 279) (B), 23001 [0.IV.313] (B, B), 3 35068 [V.127] (B), 35486 [V. 135] (B), 35642 [V.136] (B), 35700 [V.137] (B), 36020 [V.1h2] (B). VITEX AMBONIENSIS var. AMANIENSIS Pi Additional literature: swoneeeete Phytologia 5: 206. 19553 Mol- denke, Résumé 145 & 1:75. Citations: TANGANYIKA: ‘iat. Inst. Amani 221 (B). VITEX BENTHAMIANA Damin Additional literature: Moldenke, Phytologia 5: 355. 1956; Mol- denke, Résumé 211, 390, & 475. 1959. The original description 0 of Bentham's V. trifolia var. parvi- flora is "erect; leaflets 5 or sometimes Je ovate or ovate- lanceolate, acute e; flowers much smaller; reeenhl ing V. Negundo from which it is scarcely distinguishable." VITEX BREVILABIATA Ducke Additional literature: Mol hytologia 5: 218—220. 1955; Moldenices Am, Midl. Nat. 59: %5. "$968; Moldenke, Résumé 111 & 75. 19 Coelho gecarshee the flowers of this plant as rose-colored, blooming in Augu Additional citations: BRAZIL: Amazonas: Coelho s.n. (Herb. Inst. Nac, Pesq. Amaz. )02lj] (Bs), son. (Herb. Inst. Nac. Pesq. Amaz. 5729] (2 ). VITEX BUCHANANII J. conga Additional reetemtent ldenke, Phytologia 5: 220--221. wee Wega Résumé ils, 150, 11, 381, & 475. 1959. dditional citations: TANGANYIKA : Schlieben 1614 (B). VITEX BUCHANANII var. QUADRANGULA (Gttrke) Piep Additional literature: Moldenk Piytologia “bs 221. 19553 Moldenke, Résumé 145, 150, 381, 368, & 1959. Additional citations: BRIT TISH a na Stolz 512 (B). Eln. Additional and emended literature: - Bot. 10: 3798-3799. 1939; enya. payeciseta’ st "223. 1955) Wolden- ke, gene 185 & 475. 1959. 1 suberect tree; trunk 3 dm. in diameter 6 n. oe sub- fovea branched chiefly hee Ns top; wood very hard an brittle, dull-brown except the thin dingy- or yellowish-whi ite sapwood; bark smoothish, gray, witzecas except the epidermis; 30 PETTOLOGTIA Vol. 8, no. 1 branchlets ascending, rigid, breaking with a snap, gray, glabrous, the young twig portion covered with a ferrufinous pubescence which soon wears off with age; leaves not numerous, opposite, ter c=-obl , Subchartaceous, entire, shallowly folded upon the upper sublucid surface, obtuse at the base, mostly acute and recurved at the apex, only slightly paler green beneath, cur- tan=b: ° » the p the ultimate ones very fine; petiolules 1.5—l cm. long, radial sp » Strict, yellowish-brown-pubescent especially at their thickened bases, relatively stout, canaliculate on the upper sur face; flowers not known; infructescence erect or suberect, term inal or axillary, about as long as the subtending leaf, sparingly paniculately branched, 1-3 dm, long, branched from below the f t Elmer (no. 17004) in secondary woods, at an altitude of 1750 ; @ province of Sorsogon, Lu- zon, Philippines, in August, 1916. It is very probable that this is actually a species of Teijsmanniodendron. In fact, Elmer states that it is closely related to Vitex curr. He. J. Lam the veinlet reticulation quite evident beneath, Additional citations: PHILIPPINE ISLANDS: Luzon: Elmer 17004 (Ca~-272090—isotype, Um-~17l,—isotype). VITEX BURMENSIS Moldenke, sp. nov. Arbor excelsa; foliis 5-foliatis; foliolis ellipticis acumina- tis integris, ad basin acuminatis » Supra breviter pubescentibus, subtus dense fulvo~pubescentibus; inflorescentiis axillaribus abbreviatis ubique dense pubescentibus T. onal, ght less so in age; twigs very acutely tetragonal and ed, ve. ns rown-puberulent, sulcate in drying; leaves decussate- opposite, 5-foliolate; petioles slender, 6.5—8.3 cm. long, 1961 Moldenke, Notes on Vitex 3 densely brown~pubescent; leaflets unequal; petiolules i—17 m, long, densely brown-pubescent, posgehy eee ve and Pit ghtly canalicu- late above; leaflet-blades membranous cm. wide, the rest proportionately smaller; inflorescence axil- lary, shorter than the petiole, about 7 cm. long, rather loosely ~flowered, cymose, rg brown-pubescent throughout; ped- uncles slender, flattened, 2.3--3.3 cm. long, usually one long shallowly dentate; corolla about cot or 8 mm. long, densely fulv- ous=pubescent on the outer sur. The type of this species was paeconne by R. S. Cooper (ee 5969a) in the mixed forest around Kanpelet Station, Mount V: toria, Burma, on May 27, 192), and is deposited in the “Naat of the University of California at Berkeley. eaayet re BURMA: Upper Burma: R. S. Cooper 5969a (Ca—17027) —type). o Pros pint pe Sandw. tional ems Prsmeann Phytologia 5: 356 & Ok. 19565 Moldenke, Biol. Abstr. 30: 170). 1956; Moldenke, Résumé i iui, 387, £15, & h7s. 19835 Moldenke, Résumé Suppl. 1: 5. 9596 Maguire, Wurdack, & Bunting describe this species as a sprawling tree 3-10 m,. tall, the corolla blue-purple, its throat yellow, or purple-blue, the te lip with a yellon spot at its base. Green fruit was present February. It has been found at altitudes of 100 to 10 arose and is said to be "oc- casional" in distribution. Additional citations: VENEZUELA: Amazonas: Maguire & Bun 37402 (N, S), 37413 (N, 8)5 Wurdack & Adder rley 13038 ee (WW, S). SURINAM: ouw & Lindeman 2662 (N, Ut—16655b). BRAZIL: Amazonas: William s.n. [Herb. Inst. Nac. Pesq. Amaz. 2977] (Z). VITEX CANESCENS Kurz Peerage literature: Moldenke, Phytologia 5: 259—261 ae 3 and 5: eatny 1956; Moldenke, Résumé 16h, 166, 171, 17h, 177, 178 =] & ecent coltectors describe this as a shrub 6 feet tall or a ‘ne 4 m. tall and 90 cm. in trunk circumference, with white, light-yellow, or yellowish-white flowers in August, Green fruit has been collected in February and is described as "bitter", The species is said to be common in mixed deciduous forests and scrub jungles in Thailand. It has been misidentified in herbaria as V. pubescens Vahl. Additional citations: BURMA: Upper Burma: R. E. Co 6107 (Ca—1702)7). CHINA: Kwangtung: Peng, Tak, & Kin 883 Herb. Can- 32 PHYT0L0G TT & Vol. 8, now 1 ton Chr, Coll. 12882] (Ca--27)712). Szechuan: T. C. Lee 3321 (W— 1990034); W. Wang 611) (W—199070)). HAINAN ISTAND: How 70722 (B). INDOCHINA: Annam: Poilane 102 (Ca--38883), 6275 (Ca--5398h) . | aren bodia: Bejaud 522 (B); Thorel sn. [Me-Kong] (B). Cochinc Harmand . 1889 (Pierre 64,8] (B, B, B, Ca—53979); Thorel 1591 (B). THAILAND: heng 1167 (Herb. Roy. Forest Dept. 22862] ve Khantchai Herb. Roy. Forest Dept. 15970] (Sm); Winnit 3) (Ca—31)8889) . VITEX CAPITATA Vahl Additional literature: Moldenke, Phytologia 5: 261-26) O92) 5: 356 (1956), and 5: 66. 1957; Moldenke in Stey ana 28: 1082. 1957; Moldenke, owe 62, 63, 69, 7h, 1, 208, 309, 330, 381, 382, 391, & 475. 1959. Additional citations VENEZUELA: Bolivar: Ll. Williams 1162 (Ew), 12046 (En), 12696 (Ew). Gudrico: Aristeguieta ta 1183 (Bm), 4187 (Bm). SURINAM: Rom Rombouts 151 (Ut—-1)172b). BRAZIL: Rio Branco: J. G. Kuhlmann 2915 (S, S—photo). VITEX CHRYSOMALLUM Steud, a literature: Moldenke, Phytologia 5: 269—271. 19 Additional citations: MADAGASCAR: Decary sen. [1 Mai 1928] (S). VITEX CILIATA Pierr Additional hitevahianes Moldenke, Phytologia 5: 356. 1956; Moldenke, Résumé 140, 382, & 75. 1959. Additional citations: EQUATORIAL AFRICA: Gabun Heintz 10 (B); Klaine 2809 (B), 3227 (CaShl10)3 Le Testu 1701 oc Pe rpeomige Britton & P, Wils, tional baer tine 3 Meldonkes 2 hytologia 5: 2 27h. 19555 4 Alain in Leén & Alain, Fl. Cuba lj: 317 & 318, Wos7; Mol- denke, Résumé 53 & WTe. 1959. VITEX COFASSUS Reinw, Additional literature: Moldenke, Phytologia 5: 3&6 (1956) : 13-1). 1957; Moldisike, “een e% =? 195, 198, 199, Additional syno: itt a eg x Moldenke, to 20 m. in diameter, often s Sparse, the canopy irregular; bole 3—26 m. tall, usually crooked and’ twisted, fluted or 1961 Moldenke, Notes on Vitex 33 deeply fluted and channeled, 30—85 cm. in diameter at breast height, to 225 cm. in circumference at 2m. height, with high uted buttresses 2-6 m. tall, the buttresses very much grooved and flanged, fluted to the first branch, to 1.2 m. ge trunk freely branched, 30-—-50 cm. in diamet er; bark close, varying fron light gray-browm, pale brownish-gray, or brown to fad ovate or white, 5--13 mm, thick, scaly or papery but of very fibrous tex- ter bark heart- wood ee fron dark- or chocolate “brown to o light-brown, often sharply defined, separated tangentially with some yellow; branches twiste d, heavy; leaves rather thin, dull, the blades 1 ple, or p ish~blue to violet, pale-violet, or mauve, sometimes described as "lavender and white"; fruit flestw, green or light- green ay-green when immature, dark-blue black when ripe, Sad guiets inside, ovoid, surrounded on the gyitws half by the ee fruit species has been found by collectors in clay or clayey soil, sf tert river flats, dry gravelly flatland in rainforests, flatl and planting areas, swamp forests, primary forests, seco and near river- i gf él 7 tion compared to the number of flowers produced. In addition to the months mentioned in “ped sagemees St 2% (1955), the species has been cevnd in anthesis in June and October, and in fruit Jamary, March to May, September, er October. _ very durable timber is used to make house posts, drums, plates, ax handles, and p A Additional vernacular names are "bitim", "garamut", "gowaha", os arte speaker, Mjangor", "la vase", "mokuli", "nar-a-vain' tgarikon", and "sido: in, The Kanetira 3 distributed as this species is var. puberula H. J. pt perp citations: CELEBES: Bttinemeijer 10773 Conshzoest) ; Politan 6 [Boschbouwproefst. BB.31891] (Vi). MOLUCCA I << Ceram: Kornassi 451 (Ca--234911), 1039 (Ca--265952). sn 3h PHYTOLOGIA Vol. 8, no. 1 Main & Aden 926 (Ng--16903); Tangkilisan 19 [Boschbouwproefst. BB. 33731) (Ng--16870), 239 [Boschbouwproefst. BB.33909] (Ne—16959) . NEW GUINEA: Batanta Island: Van Royen 3247 (N). Dutch New Guinea: Brouwer BW. 809 (Ng—-2022)), B.W.543 3 (Ng—-20225) ; Pleyte 1033 (Ng—168h6, | Ng) 5 i poppers Bt Gee B.W.690 (Ng—-169h,). Northeastern New Guinea: Barrett 18 (Ng—16977); Cavanaugh 2022 (Ng—-6567, Neg), 4052 (Ng—6573); C saomens 1g (8); Zeya 3388 388 (Ne—16911); Jagat land 4869 (Ng—832 2s Pike 250 ag Ng); MeViegh & Ridg- well 7366 (Ng—16900); ™N. G "N. Ge Fo" 866 (Ng—6564); A. C. Richard- son 1086 | sapiorsitg 3 J. C. Saunders rs 527 sepriycees Be F. T. Vick- ery LAL (Ne—6565, Ne); White, Dadswell, & Smith 1616 (Ng—6566); Womersley 2913 (nee 2989 ( (Ng—655 2) 3897 (Ng—6571). Pap- ua: Brass 21909 (Ng—17165), 21: 2950 (Ng—17163). 1 BISMARK ARCHIPEL- AGO: New Britain: Floyd 6471 (Ng--16888), 6633 geitE)s Wom- ersley & Kazakof 7082 (Ng—16899). AROE ISLANDS: Kobroor: Buwalda Ng—16956) . SOLOMON ISLANDS: Bougainville: "N. G. 73 577 (Ng—6563). inte pen Java: Herb. Hort. Bot. Bogor. san. (B)s fll ath Ninth lla tint Dokl VITEX COFASSUS var. PUBERULA H. J. Additi literature: ah a ae Phytologia 5: 280. 19553 Mol- pee ef Rome! 3 20h, & 475. 1959. erbariun material of this variety is often misidentified as witees ve cofassus Reinw, 2 oe oe citations: CAROLINE ISLANDS: Pelew Islands: Kanehira Ca VITEX COLUMBIENSIS Bhan da Additi litera idenke, Phytologia 5: 280 & 293-295. 1955; Moldenke, Biol. A bar 30: 170h & 3551, eed ere Abstr. 30: 4370, 1958; Moldenke, Résumé 69, 419, & 475. VITEX Re pba A Turez, onal literature: Moldenke, Phytologia 5: 295—300. mr Moldenke, Résumé 62—6),, 69, 7h, 76—78, 85, 111, 225, 276, Me 22; & 475. 1959, tional ¢ 1687 z), citations: BRITISH GUIANA: Jenman 7637 (S); Little VITEX pepe oe She ea & Th. prod 5s) pe 4 7. 2 1956; Noldenke, Meme’ 137, 135. agit on, 383, We Additional citations: BELGIAN CONGO: Callens 1,97 (Z). Mage oo see Standl, onal literature: Moldenke, Phytol « 1955 Moldenke, Résumé 43, hh, 47, 49, & a uTS, Ise. a 1961 Moldenke, Notes on Vitex 35 The species is said by Allen to have pale-blue flowers and to in pastures. Additional citations: NICARAGUA: Zelaya: Shank 1172 (Ca— 29618). COSTA RICA: Puntarenas: P. H. Allen 6250 (Ca—10782). : Canal Zone: Stern & Chambers 159 (N). Bert. Additional literature: Moldenke, Phytologia 5: 357. 1956; Moldenke, Résumé 69, 7h, 111, 115, "18, 128, a 301, 382, ’ 383, 387 & 475. 1959; Angely, Fl. Paran, 16: 80° (19 60) and 17: 7. eee Vitex cimosa Bert. ex Angely, Fl. Paran. 16: 120, wie dditdonal iCal BRAZIL: Par&: Krukoff 5903 (Mi). PARA- aie Hassler 12307 (Go). CULTIVATED: Brazil: Bot. Gard. Goeldi tree VITEX DIVARICATA Sw. Additional literature: Moldenke, Phytologia 5: 357. 1956; Maldacke, Réaeb 56, 57, 59-63, Th, 225, 23), 353» 363, 391, & 76. 1959. - The Williams specimen cited below bears striking resemblance to V. orinocensis var. multiflora (Miq.) Huber. Additional citations: PUERTO RICO: Otero MoL7 (Mi. VENEZUELA: Aragua: Ll. Williams 11119 (Mi). VITEX DIVARICATA var. CUBENSIS Urb. Additional literature: Moldenke, Phytologia 5: 319-321. 1955; Moldenke, Résumé 53, 57, jd 383, & ne 1959. Howard describes this as shrub 7 feet tall, growing on lime- stone outcrops. Additional citations: CUBA: Havana: P. Wilson 1066 (Ca--68327h —cotype). Las Villas: C. F. Baker 3409 (Ca--663293); Howard, Briggs, Kamb, Lane, & Ritland 59 59 (Ca——998135) « VITEX DONIANA Swe Additional i ieaeaks Moloney, Sketch Anker tba W. Afr. 523. 1887; paisenks, Phytologia 5: 357— =: pre ores Alain, Fl. Cuba h: at & 318. a715 6 53, 1,0, 12, 13, 145--147, 150, 1 Oss, SST, ‘oes 227, 582, “a, 387, 390, 26; & 476. wi igegs See line Résumé Suppl. 1: 9 & 10 3. Vu q Alain recards the vernacular name "of6én" from Cuba, Peter British Nyasaland. Dr. Buchheim reports that there is a specimen of this species collected by Engler on October 7, 1905 [A. Peter 51820] preserved in the Berlin herbarium. ~~ Recent con ecer describe this species as a medium-sized much-branched tree, completely deciduous, the trunk 30—80 cm in diameter, the bole thick, giving thick cover, bark gray- “lin with brown slip; wood wood white, soft to cut; s ap colorless; leaves 36 PHYTOL OG A Vol. 8, nod small, ic; fruit edible, blackish-purple when ripe. Peter 8279 and “1821 exhibit leaves that are abundantly galled, the gal: galls pr ropinent and rounded on the upper leaf-surface. In addition to the months listed in ny monograph, it has been found in anthesis in January, April, June, and November, and in fruit in June and July. It has been collected at altitudes of 300 to 2165 meters, on hard stony soil in open parkland and in gullies in Brachystegia woodland. It has been misidentified in ooh a as V. madiensis Oliv. Additional citations: LIB e 1905 Ae De ae COAST: Aubréville 806 (B); Serv. saver + Pemraee 3908 : Zenker 742 (B); Zenker & Staudt 302 (Ca—617110). Sisaayten: ngler | n. [A. Peter 51818) (B), sn. [A. Peter 51819] (B); Faulkner Ws (S), 1202 (Ss); Goetze 1397 (B); Grote son. (A. Peter - 51809] | (B)3 A. Peter ter 9b {0.1 .137] ( (B), 7250 [0 (0.11 .115] (B), 7299 [O.II. 117] (B), 8279 8279 [0. III.30] (B), 8334 [0.11.33] (B), 12817 [0. III.162] (B), | 14950 [0.111 .23)] (B), 16207 [0.Iv.35] (B), 16351 (0. IV.43] (B), 17281 [0.I1V.78] (B), 31639 [V.55] (B), 32109 [V. 66] (B), 51821 [0.11.30] (B); Schlieben 1: 1282 (B); Stuhlmann Son. [A. Peter 51817] (B); Tanner 1758 (Ca--183319). ZANZIBAR: E. Hee L. Krause 1680) (B). ANGOLA: Lunda: Gossweiler 11109 (B). BRITISH NYASALAND: Brass 1707) (Ca—1707k); Stolz 402 (B (B). VITEX DONIANA var. PARVIFOLIA (Engl, ) Moldenke Additional literature: Moldenke, Phytologia 5: 326—-327. 1955; Moldenke, Résumé 139, ae oe ee & 76. “1959. Additional citations ONGO: Callens L778 (2). VITEX DUCKEI Huber 3s) Additional literature: Mold enke, Phytologia 5: se (19 and htt 358-—359. 1956; Moldenke, Résumé lll & 476. 1959. tional citations: BRAZIL: Paré: Murga Pires, Black Black, Wur- dack, & Silva 6523 (Rn, Ny). VITEX EPIDICTYOIDES Mildbr, Additional aD NolAmke Phytologia 5: 332. 1955; Moldenke, Résumé 112, 3 250, & h76. ‘353. #. Peter. describes a as a tree, 2m. tall, fruiting in February, growing at an ‘ltitace of Bho meters, Additional citations : TANGANYIKA: A. Peter 3700 (V.158] (B), BRITISH NYASALAND: Stolz 556 (Be-isotype) . VITEX ERIOCLONA H onal literature: Moldenke, Phytologia 5: 332—333. 1955; Tree; ; branches subtetragonal “the younger parts striate, ful- vous— or cinereous-farinose; leaves 1-foliolate, petiolate; pet- ioles very slender, 1.5—3,5 om. long, with an indistinct articu- 1961 Moldenke, Notes on Vitex 37 lation at the apex, sulcate above, fulvous- or cinereous-farinose; leaflet-blade sessile, ovate-elliptic, 9e5—15 om. long, 4.55.7 cm. Wide, acute at the apex, entire, acute or broadly cuneate to rounded or truncate at the base, glabrous (except for the puberu- lous midrib) and brunnescent abo ove in drying, very densely yellow tomentose beneath; secondaries 7-12 per side, promimulent; in- florescence terminal, paniculate, foliose at the base, rather conspicuously bracteolate, fulvous- or cinereous-farinose through- ong * ong 3 bractlets foliaceous, ovate-lanceolate, ad mm. gh 1.5-—2 mm. ar Fg A. R. Schultz, Inst. Tec. Rio Grande do Sul Pub. gin tog Pesquists obs 18. 1960; Angely, fe Paran. 16: 30 ‘aseoh and 17: 7. tional ae neem recorded for this plant are Ns preto", "tapinho#", "taruma duro", and "tarum& It has been collected at sea-level, in pinewoods, residual — ests, riverbank forests, and "in mato branco without Araucaria". The flowers are described by Brade as yellowish and blue, while Reitz & Klein describe them as white and red or white and light- Violet, and the fruit as dark-red. They found the species on varzea "land, fruiting in February. poner citations: ARE ire Parand: H. M. Filho 7) [Herb. Mus, 316; Herb. Curso F 719] (S)» Rio de Janeiro: Brade a2 aera Rio de Jan. 29519) () 3 Santos Lima & Brade 13295 [Herb. Rio de Jan. 29518] (B). Rio Grande do Sul: Rambo ener (Gg—-354595); Sehnem 3516 (Gg—356389). Santa Catarina: ein 232 (Herb. Barb. Rodr. 1610] (N), 303 (Herb. Barb. Rodr sat sa» 1852 [Herb. Barb. Rodr. 10069] (N, peigeinly 1858 20609); Rambo 31521 (Ur); Reitz & Klein 3009 [Herb . Barb. a ens (N, Sm), > 5915 (sm), 5968 [Herb. Barb. Rodr. 16273] (N, Sm), 6409 (Sm), 758 ( (im), 7613 (Mm), 8506 (3m), 9325 (Gg); Smith & Klein 8027 (N, Ok), 826), ( (Ok), 10685 ( coe 11928 (Ok) . PARAGUAY : Hassler 11417 (T1). URUGUAY: Herter 1 2 [Herb. Hert- er 88856] (N), 1392 (Herb. Herter 902] (N)- mani Uru- guay: Herter 1392d [Herb. Herter 98517] (N). 16 PHYTOLOGIA Vol. 8, no. 1 VITEX MEXIAE Moldenke Additional literature: Moldenke, Phytologia 5: 71--73. 1957; Moldenke, Résumé 111, 225, 381, 389, & 77. 1959. Irwin describes the corolla a as tena 8 ef ooten that the plant is common in red clay soil of secondary f ts. Additional citations: BRAZIL: Minas Gerais: teen “2062 (N, W- 2281262); Mexia 5251 (a0), Su7k (Go). RANTHA Gtirke Additional literature: Moldenke yb 5: 473--h 7h. 19573 Woldenke, Résuné 137, 7, 139, 385, & 77. 1959. An ad ar name recorded for this plant is omeeeerees Additional citations: IVORY COAST: Aubréville 105 (N); Banco 386 (B, Ca—53500). VITEX MOLLIS H.B.K. Additional literature » Phytologia 5: 76—181. 1957; Laven — 40, 205. 216, 38 381-387, 38s 390, & Lit 1959. Hinto ound this s growing in coconut orchard spec : 20 cette ates other nat or ieahectrs have found it scatter ed small canyons, in rocky deciduous woodlands, abundant near the summits of ravines, in open brushy palm savannas, an uncommon tree in loam soil on old lava flows in o sun, in gravelly- granite soil on hillslopes, om arid slopes with low scattered trees, and in grazed areas with the vegetation m or | penn leg- t as "a f. tree.. fruit sold in markets" in Jalisco. Turner reports that it is used by the natives in Michoacdn in treatment of stomach ache, n 106 is a mixture with something non-verbenaceous. He a that the fruit is “very refreshing, eaten raw or added to goat milk; leaves are boiled and put on rheumatic patients". Recent collectors describe the crom as 25 m, wide; corolla » the lower lip darker, the tube purple-lined within; fruit globose, brown or black when ripe, reddish on the upper a to 2.5 em. wide, re- sembling a small apple (Malus pumila), It has been found in anthesis also 808 July and November. _ itional vernacular names are "atoyac", "higualmo", and "j The M. E. Jones 23032 distributed as this species is REA ts Tabebuia chrysantha tha (Jacq.) Nichols. in the Bignoniaceae. Additional citations: MEXICO: Chilmatma: Hartman 1 1034 (Ca-- 702713, Du--351432); ze ton 106 (Au--122928). Guerrero: Hint 10989 (Ca—97688, uh, - Jalisco: McVaugh 11973 (Mi), se (a); Wilbur & Wilbur 2018" oes. Michoac4n: McVaugh 15051 (Mi), (a) B. L. Turner 2183 (Mi); Leavenworth & Hoogstraal 1726 17% (is). elos: Pringle ngle 6993 93 (B). Nayarit: Feddema 16 (Mi); M. fa Be Jones 22698 (Gg—1 7286) ; MeVaugh 12078 (Mi), 15222 (Mi). Oaxaca: R. Me King 1259 (W—-2301502); Wigeins 13286 (Du--l0053). Sonora: Rs We 1961 Moldenke, Notes on Vitex 7 Moran 02) (Ca--1758, Du--353268); Wiggins & Rollins 452 (Ca— 721797). VITEX MOMBASSAE Vatke Additional synonymy: Vitex mombassae Rich. ex Moldenke, Résu- mé 386, in syn. 1959. Vitex tex mombassae Vahl ex Moldenke, R 386, in syn. 1959. near Add itional literature: Moldenke, Phytologia 5: eel pols tiga Résumé 143, 145, 146, 18-151, 383, 386, 389, & 77 Recent collectors describe this species as a small spindly tree, to 5m. tall, with gray smooth bark and pale-biue or white and blue flowers, blossoming in March and April. It has been collected in fruit in January and August at sitienise to 1640 m. Bullock reports it as common in the scrub on hillsides. Additional citations: TANGANYIKA: A. A. Bullock 2372 (S); Busse 278) [A, Peter 51808] (B); Holtz s.n. [A. Peter Fash (B); A. Peter 22959 [0.IV.278] (B), 34223 [V.115] (B), 35007 [V.126] (B), 35332 (V.132] (B), 39641 [V.208] (B), 44138 [V.32s] (B), 44232 [V.3 [V.325] (B); Schlieben 1477 (B), 5353 (B). BRITISH NYASA- LAND: Stolz 1737 (B). SOUTHERN RHODESIA: Wormald 7 79/51 (Ca—— 10495). VITEX MONROVIANA Pieper Additional literature: Moldenke, Phytologia 5: 84-85. 1957; Moldenke, Résumé 137 & 77. 1959. e describes this 8 a neditm-sised tree, ferruginous tomentos ehirdite throughout, growing at an altitude of 20 m. It has been misidentified in herbaria as V. rufa A. Chev. A NEW COLOR FORM OF THE GROUNDNUT Fred W. Oswald APIOS AMERICANA f. KEIHNERI Oswald, f. Haec forma a forma typica specied Soeollip nisi quod in gula macula parva alba ubique indico— usque ad atro-corinthiano- purpureis recedit. This form differs from the typical form of the species in that the flowers are completely Indian Purple to Dark Corinthian ood nd inside and out, except for a small white area in the completely destroyed in creating a parking lot before big cia material could be collected and described. On August 48 PHIYTOLOGIA Vol. 8, now 1 20, 1960, the plant was rediscovered and collected by myself a- long a nameless rill east of the Bickford Pond -- Boren Road and west of Mill Brook, in Porter, Oxford County, Maine, and t Gees is deposited in the H. N. Moldenke Becseae age at Yonkers, New York. The plant is named in honor of Robert Volk Keihner (1891-- ) as a token of appreciation for his many decades of teaching the sciences to the youth of the Hackensack (New Jersey) High School, the author included, and for giving so many of these young people countless hours ° s own free time to further student interest and knowledge in the fields of nature and conservation. rs of flo have been variously de- The col scribed by authors, but few of these descriptions are, to an ar- tist's eye, complete ly accurate. Utilizing COLOR STANDARDS AND COLOR NOMENCLATURE, pA Robert Ridgway, to match the exact colors against the flowers of growing plants at Commack, Long Island, New York, at Ten eg River Scout Camps, Narrowsburg, New York, at Salis sbury, Massachusetts, at North Windham, and at East on, Maine, the month of August, "561, tis cde of the typical form. of the species were obtained as follows: However, upon closer scrutiny, and by u using the color charts re- ferred to above, it will be discovered that the taplon externas color of the standard blends from Pale Vinaceous to Vinaceou the tip White to Dull Green-yellow. The internal colaxe of i standard are Garnet-browm to Acajou Red, with the throat White and/or Pale Green-yellow. Both surfaces of the wings are Oxblood Red to Dark Perilla Purple, while — keel blends from Dark Vin- aceous to Pale Dull Gre en-yellow r White, To obtain the exact colors, the standard, vine and keel had to be matched separately, that is 8, each part r ved and color- tested without the other parts present to oe a false tone. Each side of every fl part was thus ed. In checking the colors of flowers, each c many may possess a dark or light blend of one color, though orn slight, and these actually become two or more different colors with each bearing a different name, and this fact must be taken into con- sideration. Freshly opened and normally developed flowers only nati ow, Medium gray-toned paper (i.e., Pale Smoke Gray) is an excel- im one to use as a near background for the flower-parts to be PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain limitations of size and proportion. An extra charge will be made for half- tones. ing on their size, as fixed by the engraver, with a minimum of about $2.25. Articles dealing with research in all lines of hotany, in any reasonable , biographical sketches, and critical reviews and summaries of liter- ature will be consid amateur or so-called popular type, and polemics will not be published. Advice on the suitability of manuscripts will be solicited, if necessary, from quali- fied botanists. der the present cost of printing, the hasic rate for a page or fraction thereof is approximately $3.00 for an edition of 250 copies. This price is subject to change without notice, since it depends entirely on the prices prevailing ind Reprints will be furnished at cost. 4 proportionate fraction of the edition of 250 copies is also furnished gratis to contributors, Upon request, the editors will send detailed instructions concerning the preparation of manuscript or further information about the magazine. In- _ quiries may be addressed to the magazine or to either editor. Bs i a FHY TOLOGLA Designed to expedite botanical publication a Vol. 8 November, 1961 No. 2 Nov 6 - 1961 G CONTENTS SROEN LIBRA LITTLE, E. L., Avicennia nittte-tNom actrees. oe 49 MOLDENKE, H. N., Notes on new and noteworthy plants. XXIX ...... 37 OSWALD, F. W., A new color form of groundnut from New York ....... 61 MOLDENKE, H. N., Additional notes on the genus Vitex. Il ......... 61 MOLDENKE, H. N., Materials toward a —— of the genus POOR Fe Fb) ok EER wh 0 oes ae ek 95 eee by Harold N. Moldenke and Alma L. Moldenke 15 Glenbrook Aveane Yonkers 5, New York, U.S. A. Price of this number, $1; per volume, $5.75 in advance noo Nel. @ 5, pectoral October 6, 1961 _ AVICENNIA NITIDA (NOMENCLATURE) Elbert L. Little, Jr. The common black-mangrove of ah ing s rican s has long been known as Avicennia qe Recently W. T. Ste has proposed for this ier the ombination A. germinans (L.) Stearn. However, the b A. is L. was validly published by Linnaeus nearly ra 3 earlier for a larger group with broader "clremscription Thus, A. nitida Jacq., - almost universal use for two d years, name. The clature is of special interest, because this species is en _~ —— — and publications about vegetation and merally as A. nitida, Moldenke (Phytologia 7: 7: 165-168, 179-181, 198} Cited many references. This common species is the most widely distributed and hardi- est of the New World mangroves. It extends the pce ade north in = United States to Texas, Louisiana, and Mississippi, as well e native Atlan this species ranges from Be ‘ northern Mexico south to Br (Espirito Santo) and on the Pacific Coast from northern Mexico south Gi Islands, The r is usually to include west- ange is stated erm Africa also, However, Moldenke in ping 7: 149, 1960) refers the west African plants to A. africana P. Beauv., which is “admittedly very closely related." The gems Avicennia L. was revised by gt C. Bakhuizen van den Brink (Revisio Generis Avicenniae (cum tationibus diversis). Buitenzorg “goo Bot. Bul. ser. aM 33 199-223, illus. 1921), A recen —— a is by — N. Moldenke (Materials Fed f the gems Avicennia, I-III. es gr 7: 123-168, 179-232, 259-293. 53.1960). This gems is place family Verbenaceae, though Moldenke Pye cone others pth ab gn the nanreetae family Avicenniaceae with this single genus, Avicennia L, (Sp. Pl. 110. 1753; = Pl. ed, » : Sige honors Abu Sina, or a (9 80-1036), Bokhara, guished era ian and aut loasehet: ie pane Fcacs ge tirst propo d in 1735 by Linnaeus (Syst. “Wat. ed, a (folio). 7353 ¥g 2, 1B. 1740) without description as a — nou Fs ower dete noepata 8 Ds was a in Tetrandria M oe wae) ption, Limnacus (Gen. Pl. ed. 1 ° cited Rh Rheede's Hortus Indicus Malabaricus (lj: t. hS. 4g 50 FPRITTOLOGIA Vol. 8, no. 2 included this description of the seed, which was repeated in later editions (Gen, Pl. Ass 5, es ish): "SEM: — rae presmm, ne germinans intr ricarpium." Avicennia mentioned algo in other pdsiteations oe Linnaeus (rissa Zeglanioa 235 1773 Materia Medica 2. 17h9). In 313 the starting point of botanical nomencla ote Dc species, A. officinalis L. fc ooh PL. 10 1753), This name, according to W. T am (Kew Bul, oe "35. 5. 1958) is to Bh typified by nOepata” sterions (1683) ted type-locality the coast of Cochin, southern ene : Lananen Seiearien in London has specimens of this species and A. nitida, as cited by Spencer Savage (A catalogue of the Linnasan Herbarium. 225 p., illus. 195). I have seen the new microfiches of these specimens and in 1956 photographed the two flowering specimens of A. nitida (813.1, 813.2). The essential is given below. two monographs. synonymy of the common New World black-mangrove Additional later synonyms were cited in the AVICENNIA NITIDA Jacq lack-mangrove Bontia b germinane 1 Loy Syst. Nat. ed. 10: 1122. 1759. Tvi Jacq., Emm. Pl. Carib. 25. 17603 also “Sie Br, Ty. Nat. Hist. Jamaica (ed. 2), 7 Avicennia gérminanis xs Steara, Kew Bul. 1958: 3h. 1958. All the works by Linnaeus and Jacquin cited above should be orated Gn indi in chronological order, Limaeus inter- ted this species differently in each of four important books published within the five-year period 1759 to 176k. Patrick Browne Wand Nat. Hist. gees 263. 1756) called ve mangrove tree, described it clearly in and Tend observed that the seed s and germi- nates before it falls," and described the seed in athe’ a follows: *Unicun germinans, lobis foliaceis." This work, which did not adopt binomial nomenclature, put this 8 doub alone in the genus “BONTIA?". B. daphnoides d. lig ae Popeye applied to the eres Nyate ode 10, 21 Wor. meee Syst. Nat. ed. 10, 2: lize. 21759) as “A.B. 20 - Index Kewengsis ga «Oppo » pedunculis spica a later hetten® (le, Sp. ed. 2; 891. "1763) 1961 Little, Avicennia 51 and listed this name a t 68 . fA. officinalis L ci a5 genus Bontia Plum. L. Sp. PL 3 17533 Gen. P1. ed. 285. 175k) jot Thmaecs a ynamia Angiospermia and the family Myopor now has a sblagle species, Bontia da ides L. (Sp. Pl. 1, 638. “1953) In a family otherwis soTEstor te Ss World, this 8 was described from Barbados and is — rg shores in West Indies and meirhag South Ameri men of Bon daphnoides the Linnaean Herbarium = 12.1) a" W. T. Stearn (A Kew Bul. 1958: 33~37. 1958) noted that under Bontia gersinane L. (1759) were orme. references, "Brown. a= « 193" r from Jamaica an tter probabl n Cumané Venezuela, He stated that the name Bontia germinans L. is typified by 's specimen in an am 3 Avicennia)., The Browne's plan bulk of were purchased by Tinnasus in 1758, according to Spencer Savage (A catalogue of the rbarium, p. vi, 1945). Donatia Loefl. (Iter. published tly was a new name for Bontia, which was cited, Jacquin in 1760 first placed this New World ere aed in the ms Avicennia, He distinguished two species, tida Jacq. ’ be. 25. 1760; ree 1762) end i." toneons Jacq. (Enum. Pl, Carib. 25. 1760). The la included an in- to Bon and another to 7: 185. 1960)Jaequin's types of wnt species, now in the British =i pal of Natural History, represent variations of a single spec Limasus in the second edition of his Species Plan (Editio Secunda. 168) Holmiae, 1762-63) in the second hve ssed his genus Avi Pe lume (p. 890-891) suppre and A, officinalis, Bontia had two species, in ides (p. ° Antiliés now added Jacquin's first species nitida) to the » B. germinans (p. one "of the Tnites now Included a reference to eicommts tn the first edition as "Sp. pl. 1. p. Lion the citations of A. oftisianlas from the tion, Patrick Browne's Latin description, and Jacquin's second species (A. tomentosa). Linnaeus added that he reduced this species from Jacquin's personal observation. Jacquin in his Selectarum Stirpium Americanarua Historia (1763) retained the gems Avicennia with enlarged Latin Hart scriptions of his same two species. A. nitida Jacq. of was and flower, A. tomentosa Jacq. of the Cari eign continent was illustrated by a single obtuse leaf, 52 PHYTOLOGIA Vol. 8, no. 2 References under gecond species included Browne, Loefling, Linnaeus! 8 dnecription of Bontia sara Wa s ("Linn,. syst. A. 1122"), Jacquin obse a ar Lieven ae c sentanee iach & To point ay the difference illustrated also the flower and fruit of B. daphnoides (t. 113, fig. 46). edition Fialebed. 176k) followed soon after the second edition (ir60-65) and. was almost identical. Sa dean OM I Boe eres third edition el a reprint of the seco: Cabaanene of the Works of Linnaeus (ed, - p. 15. iS. 3). " have hen enna yey two copies of this edition, which Bakhnut: B cite Bontia (p. 890) in the third edition was restricted to its seapetee B. da noldes of the Antilles excluding Deinadads oie species of Limmaeus restored his yrs pes Piaiy real 891) with tao apecies, A. nitida Jacq. and rminans of the Indies, and now under his Didynamia ospermia, A. germinans had almost the same synonymy as B. rminans (1763, no except for the omission of Linnacusts “BONTTA tion and was not corrected to Index In the sixth edition of his Genera Plantarum (176)), rs we Tn this work on genera, the species were not Then, in the Appendix of the same volume, the sixth edition of his Genera Plantarum (p. 5 on. 176), Linnaeus restored and de d the geme Avice mais again en his da « He th "Bontia germinans, &- =? ‘ag ate @ synonym and A, nitida Jacq. of M 7g Be Bontia Linmnae Bonts ops artinique a us rejected py own rig germinan 1961 Little, Avicennia 53 » in a later edition of Systema Naturae (ed. 12, 2:126- wre Th67) and a reprint (ed.13, 2: h26-h27. 1770), Linnaeus accepted Avicennia with 2 species A. tomentosa Jacq. and A. nitida cove Mahe eget with only B. sL. The spec pted also urray (Linns, Systema pani = cy ing ed. 1h, p. 579. 178) and a later reprint. (ed. 15, p. 579. 1797). mera Plantarum (ed. 8) and 8 (ed. 2), Linnaeus retained Bon ides L. in its original scp a sg cs and B. IT wish to report that the binomial Avicennia germinans, which was onitted from Index Kewensis, wil svtnste pena Jamaica (1189) publi shed the year before his death. edition (1756) did not employ binomials, the pad cattion almost identical, according to the title page "added complete an indexes." The binomial Avicennia germinans as the Linnaean name without author appeare spermia, on the twelfth unnumbered s of Index I with Bontia and on the seventh unmmbered page of Index II with re ence to Bontia on page 263, Avicennia P. Browne (1789) ree and his type specimen even daniion die 4 thee Bai C (813.2), also the type of Bontia germinans L. The confusion of the two unrelated genera Avicennia and Bontia by Patrick Browne and Linnaeus was odd. — (1756) correctly in genera the other sa ne ted common the gems A was known Species, In 1753- oh ae and earlier, Linnaeus put the two gene different classes. In 176 he restored Avicennia from syno- it next to Bontia under his Didynamia s with 2-14 sh corolla tinged with purple. Bontia, which has been called olive, grows aca high shrub along shores near 5h PHYTOLOGIA Vol. 8, no. 2 mangroves but has been planted elsewhere as an ornamental. One wonders if specimens might have become mixed. omitted from Index Kewensis, a =e L. van di Bakhuizen van den Brink meh 200) traced the great mr in the specific epithets to Linnaeus pia Citing for rminans L. the “1763 reference as did Index Kewensis, fo ently overlooked the first publication of that name in 1759. He mentioned the "new name Avicennia gemminans xplained: "However, in the 3d edition of the Spec. Plant. (176)) p. 281, LINNAEUS ... gave again the generic name Avicennia, plants, as synonym to be attributed to A. tomentosa JACQ. from America. This fact was understood by LINNAEUS himself, who in the 6th edition of a Gen, Plant. (76) pe 579 are back to the American species its original name Avicennia tomentosa JACQ." Harold N. Moldenke (The Verbenaceae Avicenniaceae of Tobago, Lilloa hh: 283-336. 29) cited as syno- nyms of Avicennia nitida Jacq. (p. 33h) bo rminans L. and Bontia germinans a In the latest nonopraghes gems Moldenke(T: rminans (L,) Stearn and distin- guished 11 species been a a easetase BP corse. Se rena) " Francis Macbride adopted "7 cennia germinans (L.) Stearn in his Flora of Peru Field Mus. Wat. Wat. Sete Mer, 13 (pt. £, no. 2)t 721, Stearn in publishing the new combination Avicemmia germinans observed that this name had been attributed to Linnaeus Bakhuizen van den Brink and by Moldenke, However, he apparently overlooked the citation and comment by Bakhuizen van den Brink quoted above as well as Linnaeus's publication of this binomial. com bination Avicennia germinans (L.) Stearn (1958) must be tiove en lished twi before by ieagane (176) and by Patrick Browne (1789). 1961 Little, Avicennia 55 ation t0 be settled now is the disposi tio m of hvioemie germinans L, (Sp. Pl. on Ses 891. 76h). Is*this a iow DEAS cfs lew c. ombination? binomial to be rejected The binomial Avice rminans L. following Bontia rminans L » eugeeste So a new com tion because the author pea epithet. In new combinations published before 1953, and pil agp gr piri car og 5 ay D ge a spite of a synonym of intervening date in the second gemus. I conclude that Avicennia Sonatina mus regarded as a mast ot eae as illegi- 3 d 4 oe b L. (Syst. Nat. ed. 10, 2: 1122. » ate) nor = Scien » koe Pl. eds: 25 oe 1763) and neither OF these eariier works containing this same epithet. Likewise, Linnaeus in publi ae 2703) did not cite B. germinans L, (755) ae only the later = peblication, the ‘iy hapa preg apelbparont eptoees for r example by woe Mae ph van den Brink (1921), until brought to light Stearn (1958). Thus, B. germinans L, (1763) could be Path api ry 1958). altversat nase Troe B. ieee L. (1759) py rejected as a later homonym. Then if rminans L. were regarded as a ge there would be two pos nek. has a conscrip- tion more more nearly like that of B. Sintcans (1763) eee with the latter as its basionyn would as a new name in 176. Perhaps sae most coenesint omen is Linnaeus's citation of his ow earlier wo the name A. germinans L. (sp. P rage ed, 3, $91. "T76h) Fe FLimeeus —_ neither reference with rminan, partly upon A. officinalis ts own citation, has the Scere 56 PHYTOLOGIA Vol. 8, no. 2 game type, the Indian plant "Oepata" Rheede aioe with re- stricted type-locality the coast of Cochin, southern India (Code, Appendix IV, 2, 3d). L. (176k) as not be ees as a new com- ping tsoe ate Actas Article 55 (last paragraph) because Linnaeus deliberately cited a page sneon, rence to his Attymie earlier we, Teo same time made no reference to B. germinans L. 1759, 1763). As a new name, A. germinans L. (1761) is ns nomenclaturally exper Flaoa Wen pub lished (Art. 6) (1)), be= cause the Naxerslacts group as bed Linnaeus included the type of the hope name and epithet A. officinalis L. (1753). In other words, Linnaeus when wrap two taxono-_ c¢ groups from Old and New World should have retained the oldest legitimate name and epithet A. officinalis L. (Art. 57). So far as I know, Linnaeus accepted A. germinans in only one obscure, overlooked edition of one ae He rejected this binomial could not be rejected for that reason. authors, such as Willdenow (Sp. Pl. i h, 3: 39L- 3950 "1000) followed Linnaeus in accepting A. nitida and A. ga instead of A 5, Sermanane So far a2 TS, Pa Patrick thor re Stearn to pets up A. nitida Jacq. (1760) has been in almost universal use for about two hundred years. A. tomentosa Jacq. (1760), was published on the » is now generally treated as a synonyn, a pubescent variation. I do no turally superfluo cause its synonym included an Br. (1789) be sa Senoa + ge L. (176) (arte EF 55 (ty, “6h ( (3). conpens rminans (L.) Ste 1958) could not re- aged L3G) be ce re oe ioe ( hie d be f F. ani cause 0 ° gg ply Rietgg mai A; @eemizane (1.) Steam. 1961 Little, Avicennia 57 Thus, Avicennia nitida Jaoq., the name in almost universal use for about two hundred years, remains the correct name for black- most umeed A. tminans would serve no useful purpose. If application of the Gols -shoaTa be doubtful, then the interpre- tation should faver retention of the name established in usage. However, as complete agreement may be lacking, the nomencla- ture merits further attention under Resolution 2, accepted by the on of ternational Botanical Congress, Montreal 1959 (Taxon 8: 2h5. 1959). This resolution invites sons * ral of plant names for which stabilisation seens important, whether or not these names are now threatened ..." A nitida Jacq. is the name of an tant species for Fy and which is now threatened. Therefore, I am submitting this name for considera In the meantime, while action at the tion. Tenth International Botanical Congress is pending, the name long established in usage should be re = Forest Service, United States Department of Agriculture, m, D.C. “—e- eweee we @&S @& es = = NOTES ON NEW AND NOTEWORTHY PLANTS. XXIX Harold N. Moldenke CALLICARPA ERIOCLONA var. PAUCINERVIA (Merr.) Moldenke, comb. nov. Callicarpa paucinervia Merr., Philip. Journ. Sci. 9: 13h— 13 e 19 e : vari bus anguste ellipticis parvis longe acuminatis, ad basin cuneato- acuminatis, supra basin mimte adpresso-serrulatis utrinque recedit This variety ‘differs from the typical form of the species in ves thin-memb 2——ly 5 1 lig Ch. Ong , The type of the variety was collected by Hyon Pia Chong at Wan-Do, Korea, on October 29, 1950, and is deposited in the herbarium of the University of California at Berkeley. The Kor- te be "chhom-chaksal-nanu", 58 PHYTOLOGIA Vol. 8, no. 2 HOLMSKIOLDIA SANGUINEA f£. CITRINA Moldenke, f, nov. Haec forma a forma typica speciei corollis calycibusque yee rec T form differs from the typical a of the species in mbes its fresh corollas and calyxes ame lemon-yellow col=- or, the older ealyxes becoming Tacaatcyelton and drying light- brown. The type of this handsome form was collected by Colin Potter (no. FL.1264) in outdoor caiecauet in the Foster Botanical corner of Nuuam cay eben keg , - aaeeye Oamm, Hawaiian Islands, on September 20, 1961, deposited in the H. N. Moldenke herbarium at Yonkers, New Y at LIPPIA ADOENSIS var. PUBESCENS Moldenke, var varistas a forma typica speciei renis pet ee et lam- This variety differs from the typical form of the species in havings its stems and branches, as well as the under surface of and shrub t! thickets, subject to burning, at Katera, tude 1200 meters, Masaka District, Uganda, on Betober i; "1953, and is deposited in the herbarium of the Botanisches Museun a Berlin. STACHYTARPHETA KINGI Moldenke, sp. nov. Herba erecta; caulibus crassis obtuse tetragonis sulcatis pilosis; ramis paucis prebiagion gracilibus pubescentibus; fol- iis late ovatis acutis grosse crenatis, ad basin subtruncatis vel subcordatis, in petiolum PP pita breviter protractis, ut~ rinque densiuscule puberulentis; spicis solitariis te tea escentibus, rhachide non incrassato; calicibus adscendente di- vergentibus Erect herb, branched toward the apex only; stems stout, stramineous, obtusely tetragonal, Pe Sela ulcate toward the 1961 Moldenke, New and Noteworthy Plants 59 fine, not very conspicuous on either surface; inflorescence term- inal, spicate, solitary, densely many-flowered, to about 11 cm. long (or longer when in full anthesis or fruit); peduncles very short, about 1 cm. long, along with the rachis rather slender and at type or “this handsome species was collected by reperds Merrill King (no. 877) —- in whose honor it is named — clay loam soil, in open sun, on a flat grazed area 6 kn. WEEE of Juchitén, * along the Pan-American Highway (Route 196), = Fe ele- ~o of less than 50 phi among vegetation mai f legum- inous s and cacti, » Mexico, on July 29, 1958, pe is depootted ¥ the erbatiuc Bf of the University of Michigan a + Ann VERBENA ARENARIA Moldenke, sp. nov. Herba; ae at ramosis 5} ramis gracilibus glanduloso- b paulo Potentis multifloris dense hirsutis brevi dunculati Herb, eS low aes wide-spreading; stem much branched; 7 tr. giandular- mmly er surfaces, ovate in outline, irregularly laciniate-dentate or the larger ones sub-trifid or 3-lobed at the base, the lobes ed ci along the » cuneately narrowed into a petiole which is 5-10 mm. long and scarcely distinguishable from the lamina~base; in- florescence axillary and terminal, capitate, slightly elongating rgen often Corolla hypocrateriform, blue, the tube about 10 mn, long, glabrous outside, the limb about 7 mm, wide. he type of this species was collected by Santiago Venturi 60 PHYTOLOGIA Vol. 8, no. 2 (no. 2hil) at Rio Lali, dept. Capital, Tucumd4n, Argentina, along the So bake of the river, altitude 150 meters, on Dec pr and is deposited in the United States National Herbarium at fashington. x VERBENA ARGENTINA Moldenke, hybr. nov. Herba hybrida V. cabrerae Moldenke et V. calliantha Briq. in- termedia; foliis parvis ovatis incisis, inflorescentiis albo- strigosis, pilis arcte adpressis. Herb, probably a natural hybrid between V. cabrerae Moldenke and V. calliantha Briq., with the tomentum of the former and the leaves of the latter; stems apparently sprawling or decumbent, e nodes, branched from the io or near the base; branches slender, obtusely t cattered base, the lobes 2-toothed, rather sparsely short-strigose on both surfaces — inconspicuous appressed antrorse ately narrowed at the base into the petiole; venation somewhat prominulous beneath; inflorescence terminal, solitary, spicate, sparsely ing-pubese: long, much et aye than the calyx, acuminate, densely long=- ciliate on the margins from the base to the mid-point; calyx » 9 mm. a densely white-strigose with closely ap- pressed. antrorse » 5-apiculate at the apex, two of the apic- ulations longer Poser the other three; corolla hypocrateriform, its a about 14 mm, long, glabrous outside, the limb about 12 ~= ° The type of this hybrid was collected by pi agg Venturi (no. 14) at Mocovi, Santa Fecino, Chaco, Argentina, tember Uy 1903, and is deposited in the United States diaapat Herbar- ashington,. VERBENA NIVEA f. ROSEA Moldenke, f, nov. Haec forma a forma typica speciei corollis roseis recedit. This form differs from the typical form of the species in rose-c rollas, The type of the form was collected by Santiago Venturi (no. a in apie land betwe im pastures, at 3600 meters altitude, del Cajon, dept. Taf{, Tucum4n, Argentina, on April 20, 1926, and is — sited -— the United States National Herbarium A NEW COLOR FORM OF GROUNDNUT FROM NEW YORK Fred W. Oswald APIOS AMERICANA f. M°CULLOUGHI Oswald, Ve Haec forma a forma typtca speciei tal aly coloribus exterior- ibus vexilli pallido-purpureo-griseis vel pallido-vinabso-griseie marginibus Acajou-rubris flocculosis, coloribus interioribus vex- ili maroccano-rubris vel atro-rubris. This form differs from the typical form of the species (cfr. cfr., Sate oes Vol. 8, no. 1, p. 48) in that the external colors of standard are Pallid Purple-drab to Pale Vinaceous naceous-drab, pyres about the edges with Acajou Red, while the internal colors of the standard are Morocco Red to Maroon. Both surfaces of the wings are Victoria Lake to Maroon, while the keel is Pale Vinac oe to Deep Purplish Vinaceous. The calyx is Acajou Red instead of green. This plant was collected by myself on the west side of Davis Lake, also known as Davis Pond, a bs Delaware ade at a of nat the principles of conservation to the Boy Scouts of the Greater New York Councils, and elsewhere. In 1926, as a Boy Scout, he received the Eagle A e founder of Scouting. In 1925 he received the Hornaday Award for 4 ion, and, in 1957, the Ho: ° In 1960, for oute he the Silver B e servic It seems fitting, then, that this new form which was discover a Greater New York Council Scout Camp, should be named for & man who has lived Scou outing much of his life. ADDITIONAL NOTES ON THE GENUS VITEX. IT Harold N. Moldenke VITEX Tourn Additional and emended literature: Mann, Enum, Haw. Pl. 19h. 1867; Drake del Castillo, Fl. Polynés, Franc. 151. 1893; Guill- aumin, Ann. Mus. Colon. Marseille 9: 66 & 205. 1911; me: a eee Enum, Born, Pl, 514. 1921; S. Moore in Rendle, Journ. 63: Suppl. 81. 1925; Neal, In Gard. Hawaii 63% & i163, fee 61 62 PHYTOLOGIA Vol. 8, no. 2 2756 a ba &Ie cre seg Pac. Disc. 1) 8 ge 15. 1961; F. anda, - Bot. Mex. 26: 142, 1s, 148, 152, 167, 172, & t 17S. 19615 saline, Phytologia 8: 21-47. isha. VITEX ACUMINATA Re Br. Additional literature: F. M. Compreh. Cat. Queensl. Pl. 386, fe 362. 1913; mune eid PB 8: 2h. 1961. % Tilust rations: F. M. Bailey, Compreh. Cat. Queensl, Pl. fig. 2. 1913. AGNUS-CASTUS L. Additional literature: Moldenke, Phytologia 8: 2-26. 1961. The Raizada 85 cited in Phytologia 5: 187 1988) as the typi- se form of this species appears, on further examination, to be Haus: da Oradea citations: ITALY: Savi 1995 (B). UNION OF SOCIALIST ee REPUBLICS: Transcaspia: Michelson son 87 (N, N). Turkmanskaya: twinow 21,07 (N); Zapriagaev 1 (N). CULTIVATED: Texas: P. 0. saationt 3 (8). TEX AGNUS-CASTUS v. . PSEUDO-NEGUNDO H Additional Tiveraterss Moldenke, th weeny "8s 27. wees This “a str been collected at altitudes of 1200 t meters. The Raizada 85, cited below, was incorrectly cited by me in Phytologia as: 187 (1955) as typical V. sgms-castns Le Additional citations: IRAN: Rechinger & Rechinger 5778 (B) BALUCHISTAN: Rechinger & Rechinger 3996 (B). CULTIVATED: India: VITEX ALTISSIMA L. f. Additional literature: Moldenke, Phytologia 8: 28. 1961. The corollas of this species are described as violet, and it has been found in anthesis in August. pete gon Fe to ——— hergrins Hinds sor peemogh oe ag xy Ping Additional citations: INDIA: Andhra pduayneciel octh an 37h al; (B). VITEX BARBATA Planch, periopne pre giartoeri proc Moldenke, P Phytologia map tag ge 1956. 2), onal c ons: MALI: Sou 5032 (2), S179 (2). sl ernie VITEX BUCHANANII J. G. Baker Additional literature: Moldenke, Phytologia 8: ae ee Schlieben describes this as a tree 6--8 m. tallm with white » growing in parklands. Additional citations: TANGANYIKA: Schlieben 6008 (B). VITEX Turcs. Additional literature: shame Ae Phytologia 8: 34. 1961. Additional citations: SURINAM: Wood Herb. Surin. 3l9 (N)- 1961 Moldenke, Notes on Vitex 63 DONIANA Sweet tase literature: Moldenke, Phytologia 8: 35-36. 1961. Additional citations: MALI: Soudan: Jaeger 3377 (Z). VITEX FERRUGINEA Schum, & Thonn. id tacces -oFloce sentag Moldenke, Phytologia 5: esa 1955. Additional citations: CONGO: Corbisier~Balanot 1293 (B). VITEX GAMOSEPALA Griff. Additional literature: Moldenke, Phytologia 8: 37-—-38. 1961. Collectors describe this as a shrub 5 m. tall or a tree 10 m. tall, scattered in evergreen fconsten with yellow flowers in July and October . Additional citations: THAILAND: Thaworn 1,83 [Herb. Roy. Forest Dept. 23012] (Bk, Z), 156 [Herb. Roy. Forest Dept. 15001] (Bk, Z). GAMOSEPALA var. SCORTECHINII King & Gamble Additional synonymy: Vitex gamosepala var. scortechini King & nro ex S. Moore in Rendle, Journ. Bot. Lond. 63: Suppl. 81. Additional literature: Moldenke, Phytologia 5: 366. 1956. VITEX GAUMERI G Additional Teharetaee: ae Phytologia 8: 38. 1961; F. Miranda, Bol. Soc. Bot. Mex. 26: 152, 167, 172, & 175. 1961. Miranda records the edditic nal vernacular names "carrete", “crucillo", and "yashnike" for this species. ron (3). citations: HONDURAS: Morazén: Williams & Molina VITEX GLABRATA R Addit liters ture: F. Me Bailey, Dompesiie Cat. Queensl. Pl. tp oe F saguit sak Phytologia 8: 39. 1 reference cited above, ate the additional elie ike eeerer VITEX KENIENSIS Turrill Additional literature: Moldenke, Phytologia 8: 1. 1961. Recent collectors describe this plant as a small » about 6 m. tall; trunk about 15 om. in diameter at breast height; bark owe with scattered trees or tree-clumps of Ss and oth= ers, ot an adtd tarde of 1600 meters, flowering and fruiting in Saliva: citations: KENYA: Maas-Geesterams 6282 (S). VITEX Lo PALA King & Additions Eye ad « Maldenke, Phytologia 8: 43. 1961. 6h PHYTOLOGIA Vol. 8, no. 2 Recent collectors describe this plant as a medium-sized tree, ng in evergreen jungles, with yellow flowers in February. Addi tional ather teas THAILAND: Premrasmi s.n. (Herb. Roy. Forest Dept. 3266) (2). VITEX ree pace Te — Additi litera In Gard. Hawaii 643. 19483 Mol- denke, Pigtologia 8: ey 1961. ackwell report that the wood of this tree is valuable b However, logs of this wood are often perforated by the larvae of the Puriri Moth, which destroy the wood value, The tree is a surface-rooter, and is often blow over by gales. It is endemic and restricted to the northern part Med North Island, New Zealand, but is easily cultivated elsewhere. VITEX MONROVIANA Pi es literature: a mone ra ge eg 7. 196 Additional citations: LIBERIA: Dinklage 219) fll ge se B= isotype), 3058 (B). VITEX ees gre oo Gttrke Die. Sade Addi ure: Moldenke, Phytologia 5: 485. 1957; Mol- denke, be ser is. 1 el, & h77. 1959. Additional citations: TANGANYIKA: Schlieben 5792 (B). VITEX MOSSAMBICENSIS var. OLIGANTHA (J. G. Baker) Pieper Additional ene: Vitex mossambicensis var. oligantha (Gttrke) Pieper, in herb. Additional literature: Moldenke, Phytologia 5: 85-186. 1957; Moldenke, Résumé 145, 387, & 477 sand The Schlieben 5792 ios as this variety appears to be the typical form of the species. VITEX NEGUNDO L. ee synonymy: Vitex negundo Royle ex H. J. Lam, Verben- - Malay. Arch. 369, in syn. 1919. Vitex negunda L., in herb. peagerones Saeed literature: Campbell & Watt, Descrip. Cat. Econam. - Chutia Nagpur 39. 1 ° — 9: 66 & 205. 1911; E. D. Merr., A - Pl. 514. 19215 oL. Gard. Encycl. 1292. 19. Neal Hawaii 641 & 643. 1948; Hansford, R sa 3 75=-112. 195h5 — Abstr. 30: 1,18. 19563 2 19573 Santapau » Phytologia 5: 186-198 and na ti nied eT Biol. Abstr. 30: 4370. ie Moldenke, Am. Midl. Nat. 59: 365. 1958; Moldenke, nes thy 62) 123, ~ ee Aram 167, 320, 321), 177, te Potlane 147k specimen in the Berlin herbarium has the low- er leaves conspicuously toothed, but the upper ones entire. San- 1961 Moldenke, Notes on Vitex 65 “pes Begorss that this species is now growing "all over the hill t Pur » but was possibly earpepet he planted ree alae check as a tlie eres). & & Wall, in the reference cited above, report the ashes of the being used dyeing and the roots Polunin, Sykes, & Williens describe the calyx as white and the corolla mauve, They found it growing at 2500 meters altitude in Nepal. Hansford, in the reference cited above, describes a new species of fungus, Irenina depokensis Hansford, from this plant in Indonesia. Guillaumin cites a Pennel 300 not yet seeh by me. The Conklin s.n. [Mt. Yagaw, May h, 1953], distributed as this species, is actually V. trifolia L., while Brass 28095 is V. tri- folia var. bicolor (Willd.) Moldenke, and Chiao 1879 is Vv. 2 negun- do va: var. intermedia (P'ei) Moldenke. Clemens 5924 is host to a species of Cronartium fungus. Tsui 617 seems to be in part the typical form of this species and in part var. intermedia. Lau describes this plant as "semi-woody, 1m. diam.", and Mer- rill says that it is "widely distributed in Old World tropics". Rock 1065 presents much the appearance of f. alba Ptei. Sven white (Tak & Ch ow 2856). The plant is said to be a tree with sieges or "4])~smelling” flowers. I, has sometimes been distributed in herbaria as var, incisa (Lam.) C. B. Clarke. The Allard 11390 cited below does not , bear on its label any in- dication that it came from cultivated material, but I am assum- Py that it did. The additional common name "samaloo” is record- haasttonal citations: FLORIDA: Polk Co.: McFarlin 6112 (Mi). BALUCHISTAN Harsukh s.n. [Duthie 20615] (Ca—294806). PAKISTAN: Swat: S. Aun S.n. [10.68.52] (Vi); Rodin 5427 (Ca--36603). NEPAL: Polunin, Sykes, & & Williams 5833 (Bm). INDIA: East Punjab: Koelsz 1628 (us Mi), 4137 (Mi); Nasi: Nasir s.n s.n. [R. R. Stewart 147h3b) (Ca— 972843). Madras: Janaki 776 (Mi). United Provinces: Kaul 97 [fls. wn 3, 1922] (ca~—228229), 97 [fr. Nov. 1, 1922] (Ca--228229); U. Singh 216 (ca—361112), West Bengal: Hosein son. [Chanderna- gore, 1902] (B). State undetermined: K, Singh 5-76 [Bindalnala] (Ca 304520); Wight s.n. [Peninsula Ind. Orientalis] (Ca— 2991). CHINA: Anhwei: Herb. Univ. Nanking 1726 (Vi); Ling 1307 ake Univ. Nanking 785] (Ca--250772). Chekiang: Cheo & Wilson 66 PHYTOLOGIA Vol. 8, no. 2 227 (Herb. Univ. Nanking 12873] (Ca--316391, Ca—-35u6l1); Ching 2h29 (Ca—28181)); A. N. Steward s.n, [Herb. Univ. Nanking 2387) (Ca--230043). Fukien: H. H. Chung 1073 (Ca--225837), 3452 (Ca— 42034). Hupeh: Sun 10b > (N). Kiangsi: . Ip 85 [Herb. Univ. Nanking 7698) (Ca~-258860) . Kwangsi: Ching 51,50 (Ca--09697) « Kwangtung: S. Y. Lau 699 (N), 20133 (Ca--611)60); Tak & Chow 2856 [Herb. Canton Chr. Coll. 1717] (Ca--319390); Tso 20940 (N); Tsui 617, 2 part (Ca--612i16, Ca--882867); Ying 975 (Ca-—38892h). Ssechu- an: Fang 9122 (Du—333631, N), 9277 (Du--333626, N). Ytmnan: A. 13210 (N), s.n. [Yunnan] (N, N--photo); J. F. C. Rock 10L65 (Ca--327h51). AMOY ISLAND: A. N. Steward 3137 (Ca—hh 741), FORMOSA: H. H. Bartlett 6267 (Ca--325501); Hayashi 21212 (Ca— 344512); A. Henry s.n. 3.n. (Takow] (N); Tanaka & & Shimada 17878 (N). pr es “C. Ford s.n. (N). LANTAU ISLANDS: Taam 1728 (B, Ca-- 278). HAINAN ISLAND: W. Y. Chun 189 (Herb. Univ. Nanking ne} (Ca—-242587); Chun & Tso 43701 (Ca—59486h); S. K. Lau 293 (B, Ca--525009, N), 1209 (N); C. 1. Lei 202 (B, N), 700 (B, Ca—— 612480, N); Liang 64661 (N); May 123 (Ca—373948); Tsang 622 (Herb. Lingnan Univ. aes (B); 2 33162 (N). INDOCHINA: TA: An~ nam: Soll ails 147k (Herb. Chevalier 14,39] (B, Ca--53752). Cochin~ china: Godefroy s.n. AA es 4 de Saigon, 1875] (Ca--5L799)5 Thorel 120 (B). State undetermined: Petelot 1170 [Cho Gouh) (Ca- 2277h6). MALAYA: Penang: Haniff 239 (Ca--355089) « PHILIPPINE IS- LANDS: Luzon: Clemens 2 [Herb. Philip. Bur. Sci. tae sore 2h, Ca—268)12) ; fois: Clemens 16803 (Ca--2853hh) 5 5611 , 8125 (N), BAS" (Ca--271,0h, N); Haenke 80 Fin ant Langlassé 72 72 (B); Lete 173 (Ca--301962); Mangubat g.n. [Herb. aie 1 ge Sei. 133) (N); R. c. wmoregae era (Ere Philip. Bur. 43532) (Ca~-237688) 3 E. D. Merrill 1503 (N), 1636 (N), 2320 om. 2876 (N), 3429 (NW), 3627 (N), Sp. Blanc. lO (N)3 Van- overbergh 1397 (Vi). SUMATRA: D, Fairchild 1076 (Ca—301280). SARAWAK: M 5° (B, Ga-83750). Virginia: % Ae [. A. Allard | 11390 (Ca-—-793722) + Hrs ie Pei Additional literature: Moldenke, Phytologia 5: 98--L99. 19573 Moldenke, Résumé 171 & 77. 1959. Collectors describe this plant as a shrub, 4-6 feet tall, with persistent aromatic foliage, inhabiting open scrub at alti- tudes of 9000 to 10,000 feet. Maire B.3663 has the leaflets toothed like thosein var. intermedia (P'ei) Moldenke. It has been distributed in herbaria as V. n L.3 on the other hand, Rock 10K65, cited by me as typical V. negundo, may be f, alba instead. 1961 Moldenke, Notes on Vitex 67 The form has been found in anthesis also in May. Additional citations: CHINA: Ytmnan: Forrest 10123 agit ~~cotype); Maire B.3663 (Ca--386972); J. F.C. Rock 3} 3880 (Ca~- VITEX Serney f£. LAXIPANICULATA Pte Additi literature: tee nitemaee 6: 20. 1957; Mol- denke, Abound 171 & 477. 1959. EY owers are said to be “blue or white, blossoming in June at altitudes of 3 to 900 meters dditional citations: CHINA: Ytmnan: « Henry 13210 ( cotype); Maire 625), pentape Te 3 Yen 31 (Ca——h87585) VITEX NEGUNDO var. CANNABIFOLIA {side & Zucc.) EEC eens _literatures Neal, In Gard. Hawaii 6 19185. “yol- aS: 1499-502 and 6% 14. 19573 uoliene 7B, 152, "Yon, fn, 1273, 17h, 178, 292, 225, 361, » 382, “ss, 387, & 477. is 9593; Moldenke, Résumé Suppl. 1: 11. fe ent ve describe the flowers as lavender “or mauve Sides. An additional common name is "hemp-leaved vitex". Additional citations: NEPAL: Stainton, Sykes, & Williams 2737 (Bm). CHINA: Chekiang: Cheo & Wilson 76 [Herb. Univ. Nankin, 1272h] (Ca--354601, Ca-—316390). Fukien: L. Y. Tai 11712 (Um— 173). Kiangsu: Herb. Univ. N 659 (Ca==2300K2); Ling 2382 (Herb. Univ. Nanking 12334] (Ca--316390). Shantung: Chiao 270k (B). Szechuan: F Fang 5279 (Du--333777). CULTIVATED: New jew York: Moldenke & Moldenke 11865 (N). NEGUNDO var. HETEROPHYLLA (Franch.) Rehd. Additional and emended synonymy: Vitex negundo Curtis apud Rehd. in Sarg., Pl. Wils. 3: 33, in syn. 1916 [not V. negundo L., Pee Vitex incisa Bunge apud Rehd., Man. Cult. Trees, ed. 77, in syn. 1927 {not V. incisa Thunb., 1947, nor Wall., akos]. Vitex negundo var, incisa Clarke ex Moldenke, Résumé 387, in syn. 1959, Additional Doney, Brooklyn Bot. Gard. Rec. 30: 23+ 1941; Moldenke, fe ee aS: 502—507 and 6: 1-15. 19573 Moldenke, Résumé 15, » 30, Th; 267; 17, 185, 225, 232, We; 382, 384--387, 389, & yay 1959. Recent collectors describe the flowers as pinkish, grayish- blue, or purple, and encountered the plant in open dry loess, under pines, and on terrace walls in mountains, ascendi za std altitude. Herbarium material has been distributed as obs Addit io SY itei cae: CHINA: Hopeh: Mrs. W. Granger 8.n. 68 PHIYTOLOGIA Vol. 8, no. 2 299 [277] (S), 341 [278] (852 T. F. Li 63 (Herb. Univ. Nanking 10897] (ca--299830); F. N. Meyer 1008 (Ca--180786). Shansi: Ling 1,67 [Herb. Univ. Nanking g 9113) (Ca—-263675), 1721 [Herb. Univ. Nanking 9366] (Ca-—-266638), 1741 [Herb. Univ. Nanking 9386] (Ca-— pena Shantung: Chiao 3052 (B, Mi). CULTIVATED: Germany: Hei~ . (B). Massachusetts: Herb. Jesup s.n. [Camb. Bot. Gard., TY to, Dt); Rehder s.n. [Aug. 5, 1098) (B). Oklahoma: Duff 101 (Ok); 0. Me King 5.n. mn. (June 21, 1955] (Ok); D. McLean 86 (Ok). Virginia: H. A. Allard 11390 (Gg--381231). ope ee » INTERMEDIA (P'ei) Moldenke ional literature: ae in Leén & Alain, Fl. Cuba 4: 318. 1954 pear Bs Phytol : 16--20. ot Lombardo, Ay _ ogia « Pub. 7 & 255--256. i958 Moldenke, Résumé 1, 30, 53, on 131, 158, 165, 171, 172, imu, 178, 188, 191 j 198, 908 ety 2 7 & L77. Illustrations: Lombardo, Arb. Cult. Pas. Pub. 255. 1958. Recent collectors describe the foliage as aromatic, the corollas as light-purple or "white and purple", and the fruit sh. ; mountain forests, and on open ridges, ascending to 2800 meters altitude, Herbarium material has been misidenti- fied and distributed as V, cannabifolia Sieb. & Zucc. The © 1653 cited below was collected on the campus of Fukien Christian College and so may have come from cultivated material. Ying 1263 looks very much like typical V. negundo L., while Young 3236 3236 ap- proaches var. heterophylla (Franch, ) Rehd. Additional citations: INDIA: East Punjab: Koelz 4592 (Mi). U- nited Provinces: Gill 35 (B). CHINA: Anhwei: Herb. Univ. Nanking 1726 (Ca~-23004); Ling 1309 (Herb. Univ. Nanking 76,7] (Ca-- 25877h). Fukien: Chang & Metcalf 275 (Ca—2h9191); C. Bs sheng 1653 (Ca--563707), 1711 ~ (Ca==28 7025) 3 H. H. Chung 2371 2371 (Ca-- 233089, Ca—l20367); Metcalf & Chang 31 (Vi). Honan: A. fe: Stew- ard 1603 (Herb. Univ. Nanking 9729] (Ca--2706l9). Kiangsus S. S$. Ne Lei s.n. (Herb. Univ, Nanking 2990] (Ca--225100); Ren & Tao ate (Herb. Univ. Nanking 12581] (Ca--299863)3; G. D. W Wilder s [Herb. Univ. Nanking et (Ca—280986); Young 3236 3036 (ca--382758)- Kwangtung: Peng, Tak, & Kin 671 (Herb. Canton ton Chr. Coll. 12670] (Ca-~27h996)3 Tsui Tes (Ca--612522) ; Ying 1059 reggie: 1263 (Ca--360 . Sikang: Chiao 1 1879 (Bw). Yttnnan: Maire 1776 ° ined: David 833 (Hiokiang] (Ca~-192799). HONAM ISLAND: E. D. Mer- rill 10073 (Ca--992h3h) . Mee pyrene Zimmermann 2 (B, B). JAVA: Bakhuizen - van den Brink Gg . CULTIVATED: California: McCaskill 316 Ce Ses te 3 Seo) 1961 Moldenke, Notes on Vitex 69 VITEX ORINOCENSIS var. MULTIFLORA (Miq.) Huber Additional literature: Moldenke, Puytologia 6: 27—31. Aah Moldenke, Résumé 69, 7h, 76--78, 85, 111, 112, 226, 386, 387, & 477. 1959. records the vernacular name "aceituno" and calls the plant a riparian tree. The corolla is described by recent col- iene. as blue-lilac or purple. The tree has beenfound fruiting in M citibinea citations: VENEZUELA: Amazonas: Maguire & & Wurdack 34759 (N, S). Barinas: Bernardi 6849 (N). Bolivar: Bernardi 2915 (ad) 5 Ll, Williams 13295 (Ew). Federal District: Aristeguieta as (N). SURINAM: Florschttts & Florschttts 1228 (N); Lanjouw & eman 1862 (N), 1983 (N). BRAZIL: Amazonas: Murca Pires & ae Ke 968 (ut—~98028a, 2). Maranhfo: Frées 1952 (Mi) (Mi). CULTIVATED: Brazil: B Bot. Gard. Goeldi tree 719 (Bm). VITEX OXYCUSPIS J. G. Baker Additional literature: Moldenke, Phytologia 6: 31--32. 1957; Moldenke, Résumé 137, 5 his 113, 148, 426, & 77. 1959; Moldenke, misidentified in herbaria as V. madiensis Oliv., V. madiensis var. angustifolia Pieper, and as V. madiel ¥. madiensis var. schwe [I Pieper. dditional citations: SIERRA LEONE: D. Small 613 (B). TANGAN- vies A. Peter 17900 [0.1V.108] (B), iste y [O.1V.115] (B), 18833 [0.1V.136] (B), 202h9 (0.1V.183] (B), 276 (O.IV.231] (B, ai 23046 [0.1¥.279] (B), 39654 [V.209] yey VITEX PACHYPHYLLA J. G. Baker Additional literature: ears Phytologia 6: 33--3h. 19573 Moldenke, “Résumé 140 & 477. 1959. E um material of ae pi has been misidentified as V. obovata Ee i Mey. and as V, cienkowskii Kotsch. & Peyr. ~~ Additional citations: GABON: Debeaux 416 (B); Klaine 2656 (N)- apn eee Lecomte S.n. [Fernand-Vas, Avril 1692] (Ca— ba pegs H. Hallier erature: Moldenke, Phytologia 6: 3h. 19573 Mol- Pr Résund 188. & 77. 1959. This pret g eugene ook with a trunk 1 foo bane growing on red soil in virgin forests, pea dtiing in tees ember and Dee Additional citations: SUMATRA: Krukoff 4339 (Ca—534213)- VITEX PANSHINIANA Moldenke 70 PHYTOLOGIA Vol. 8, no. 2 Additional literature: Nik oe Phytologia 6: 34-36. 1957; 5 marl Résumé - & 477. 1959. acedo f ae ge ee blooming in July. It has been mis- stamarend in herbaria as V. polygama Cham. by G. M. Barroso. Additional citations: BRAZIL: Goyaz: Macedo 3897 (W--221,8967, W—228968) . VITEX PARVIFLORA A 200, 203, ri & [Sold Si 959; Menninger, Trop. Tree Seeds (2). 1961. Elmer de Ss ow - » heavy, te, the heartwood yellowish, odorless, tasteless, the bark thick, yellowish (even the thinly scaling surface), the branches forming a flat c crown; leaves smooth, horizontal or HE he, subcoriaceous, usually conduplicate; inflorescence » the green stalks becoming purplish-tinged; pretends erect, deciduous, pee oe slightly odorous. Recent collectors describe this a large spreading tree to 20 m tall, violet corollas, blooming also in December, inhabiting forests, streamsid , reminds of commerce, used for posts and construction. The Loher s.n. [Rizal Prov.] distributed as this species is V. stinennt WoTdenke- Additional citations: PHILIPPINE ISLANDS: Luson: Bawa (Herb. Philip. Forest Bur. 2602] (Cam—238972) 5 conite sae (Herb. Philip. Forest. Bur. awn (Mi); M. S. Clemens 19006 (c (Ca~ 2957h1), S:- [Bulacan Province, Sept.-Nov. 192) (ca~-268139); Elmer 17286 ( (Ca—272333, te—168), 18089 (Ca-~270776); G. Edafio 8.N. gen. (Herb. Philip. Bur. Sci. 1,876] (Ca~—321808) Haenke Ue (Ca—~280927)3 & Hagger 255 (Ca—7031h); sone aa (Ca—2),3058), gn. ae, May 1914] ( (Ca—229200); Quisumbing 2210 (Ok—172h6), son, (Mt. Maguiling, sora 1925] (ca SSL r Sulit & Canicosa s. n. [Los Belton, & aguna Province, March 1926] (Ca——308189 ). Mindan aot Ablasa 73 (Herb. Philip. Forest. Bur. 30281] (Ca——32108); F. Crus s.n. | s.n. [Herb. — Forest. Bur. 23893] (Ca--205896); Elmer 13440 (Ca—272778); D - Miranda s.n. (Herb. Philip. Forest. Bur. 20573] fussed: i: hase Phasis s.n. (Herb. Philip. Forest. Bur. 26939] (Ca-205)27); Ramos & Edafio s.n s.n. [Herb. Philip, Bur. Sci. 49518] (Ca—323889); Wenze: onze ~ 2759 ( (Ca——31 703). Negros: Perlas s.n. (Herb. Philip. Forest. Bur. ae (Ca--358560); A. Reyes s.n. (Herb. Philip. Forest, Bur. 29932] (Ca--2680)1). MARIANNA ISLANDS: 1961 Moldenke, Notes on Vitex 71 : Re V. Moran 4597 (Ca—52220). MOLUCCA ISLANDS: Buru: us Seprin [Toxopeus s]) 52 (Ca—23921) . HAWAIIAN ISLANDS: Oahu: R. C oes 7, 194k) (Du--377961); Degener & Park 9509 (Mi). CULT: San eaT Florida: R. F. Thorne 10062 (Ca-= (Ca--965097). Hawaiian Islands: “% Fe Judd s.n. [Bishop Estate | Meals Dec. 1, 1930] (Ca-- 50239h) . VITEX PARVIFLORA f. STERILIS H. J. Lem Additional literature: Mol enke Phytologia 6: 42), 1957; Moldenke, Résumé 185, 226, 23, & "L77. 1959. Recent Ler” "describe this plant as a tree 10 m. tall, the trunk 0 cm, in diameter at breast height, the wood strong, good thamane; considered the best wood for posts in Luzon, and ibaa in house construction. The flowers are white to bluish slightly adorous, and the fruit is fleshy, on nae Both rea", flowers and fruit, acco to ero, smell Clemens reports it as the host of a parasitic fungus on the leaves. A vernac "molave", It has been collected at name is an altitude of 110 meters, in fruit in October. Herbarium mater- ial has been distributed under the name V. parviflora Decne. Additional citations: PHILIPPINE ISLANDS: Leyte: F. Franco s. n. (Herb. Philip. Forest. Bur. 26408] (Ca-239317). Luzon: H. H. Bartlett 14940 (Ca—~16h199); M. S. Clemens 1870 (Ca—268139); Miras 3.1 Sen, (Herb. Philip. Forest. Bur. 25697] (Ca-—-239368); Parros & N & Nera l, (Herb. Philip. Forest Bur. 30143] (Ca—291921); I. Pas Pascual 8. sen. [Herb. Philip. Forest. Bur. 23063] (Ca—205891); Quisumbing 8.n. (Philip. Nat. Herb. 6579] (Ca—985LU2). Mindanao: Cordero 1 [Herb. Philip. Forest. Bur, 29816] (Ca-~256837). CULTI- VATED: Philippine Islands: Sulit 13 (Ok--17189, Ok--17233). PATULA E. tien Taiteeeunes Moldenke, Phytologia 6: hh--l5. 1957; are! Résumé ryt 15h, & 477. i959. e Rodin 4232 ributed as this species is V. harveyana H. ri W. Pearson : ee VITEX PAYOS (Lour.) Merr Additional ia veratine: Moldenke, Phytologia 6: 5—h7. ase Moldenke, Ré 5, 148 “ul, 226, 233, 301, 385, 389, 416, & 77. 1959; Moldenke, Résumé Suppl. 1: lo. 1959. Peter desc s of this plant as bluish-whi -purple, or pale-pink, and found them in an- thesis in Jam wary, March, April, November, and December. It has been encountered at altitudes of 150 to 1300 meters. Herbarium material has been misidentified me distributed as V. hilde- brandtii var cens ©? Additional citations: TANGANYIKA! H. G. Faulkner 1331 (B, 8) A. Peter er [0.III.219] (B), 16210 y [0.1V.35] (B)y “24179 (0.IV. 319] (B), 330h7 [v.90] (B), 33lkl (V.93) (B), 33896 [V.111] (B), 40654 (V.2hk] (B), 46477 [V.208] (B). SOUTHERN RHODESIA: A: Byes 1387 (Mg—-172); A. Peter 51119 (S.6] (B). PORTUGUESE Mozambique: A. Peter 51168 [S$.66] (B). MOUNTED Tate Photograph of type and original description (N). VITEX PAYOS var. GLABRESCENS (Pieper) Moldenke Additional literature: Moldenke, Phytologia 6: or 1957; eet Résumé 1)3, at 149152, 38h, & 477. eter describes thi 3 plant as a tree » growing a 2 ing altitude of 280 meters. Herbarium material has been misidentified and dis- tributed as V. yolkensii Gtirke. Additional citations: MALI: Soudan: Jaeger 3769 (Z). TANGANYI- KA: A. Peter 20936 [0.1V.196] (B). VITEX PAYOS var, pig entag Moldenke, var Haec varietas a forma typica speciei foliolis perspicue stipi- This variety differs from the typical form of the species in berans long leaflets plainly petiolulate, the petiolules being 5—- 0 _ of the variety was collected by Ernest Henry Wilson (ei "Told in thickets around Kampala, Uganda, on December 22, 1921, three ps is deposited in the herbarium of the University of at Berkeley, citati pase UGANDA: E. H. Wilson 19) (Ca-~381,08--type) . VITEX PEDUNCULARIS Wall. Additional literature: Moldenke, Phytologia 6: )9--51. aw} eget Résumé 159, 165, 166, 177, 178, 226, 380, 386—388, Recent a pies describe the bark of this tree as blackish, the trunk to 1.1 meters in circumference, and the fruit globose, with on, bitter. They have found it in mixed yg paged forests Quercus, in dry deciduous forests, in dry dipteroe o aati x “Woceasional" in open Quercus-dipterocarp fats and "common" in mixed deciduous forests, ascending to 850 meters al- titude, flowering also in February, and fruit ting also in April and July. Khant reports that its wood is used for buil and other purposes, It is possible, in my estimation, that V. ajugae- flora Dop is conspecific with V. peduncularis. 2 U 53762), 2856 (B). Cambodia: Poilane 14937 (B), 17320 (B); Thorel 1006 (B). Cochinchina: Poilane 17320 (Ca— (Ca—3887h) ; Thorel 1006 (Ca—38106). dial Bunchuai 161 [Herb. Roy. Forest Dept. 22656] (Gg); i Serene , Larsen, & Hansen 956 (Bm), 2598 (Bm), 2642 (Bm), 3900 a VITEX PETERSIANA Klotzsch 1961 Moldenke, Notes on Vitex 73 eee’ literature: Moldenke, Phytologia 6: 56--57. ree 3 em. Soc. Broter. 13: 7h. 1958 j Moldenke, Résumé 11,8, 151, 36h, a "UT. 1959. VITEX PIERREANA neni Additional literature: Moldenke, Phytologia 6: Barkin 19573 Moldenke, Résumé v7, 177, 388, 19, NO, & 477. 1959. Pierre reports that the trunk o tree is to 18 m, tall and «in ° tional citations: INDOCHINA: Cochinchina: Pierre s.n. [ad flumen Saigon, 3/5/1866] (B, B, Ga--£L810) . VITEX PIERREI Crai Additional Le Moldenke, Phytologia 6: 60-61. 1957; Woldenke, Résumé 177, 178, L19, & 477. 1959 Recent collectors’ desc ribe this as a tree 6 m. tall, with a crooked and twisted stem, gray bark, and yellow or dirty-yellow flowers, bloaming in March and ADE ig scattered in mixed deciduous forests at an altitude of 300 m. vernacular name g ton" is recorded. Pierre 5218 is marked "sp. nov." on the labels, but apparently is not one of the cotype collections Additional citations: INDOCHINA: Annam: Evrard 1512 (B). Coch- inchina: Evrard 2766 Salen Pierre 5218 18 (B, C3887, Ca—— 54956). THAILAND: Smitinand 2551 (Z); Suvarnakoses 1430 [Roy Forest Dept. 16559] (Z). PINNATA L. onal literature: S, Moore in Rendle, Journ. Bot, Lond. 63: Suppl. 81. 1925; Neal, In Gard. Hawaii 643» 198; Moldenke, Phytologia 6: 61—6l, & 70--79. 1957; Moldenke, Résumé 76, 157, 159, 16e—-167, bg or Wg eee 8 185, 187-189, 191-196, 198, 226, 23h, 302, 335, 3617 382° 38h--386, 388, 393, 19, 427, & hi77- 1959; Moidenke, Résumé Su Suppl. 1: 12. 1959. onal synonymy : Vitex pubescens var. genuina Hochre, Plant. Hochr. 33 nhs ae Oe Sieg Recent collectors describe this as a —. tree, with a trunk 100--120 cm. in circumference, the lower 5 m. without any *purp. or even yellow; ow; fruit dark-purple, Sentialeck: or black when mature, The flowers are sometimes described as pale-violet or borne and the young fruit as globose, green, topped with €s of sea~water, on plains, and near beaches. Elmer says that it is "a small-sized tree everywhere in eg woods" in — North Borneo, while Henderson describes it as a "small tree common on the seashore" in Pahang. It has been vest in sabhaete also in 7 PHYTOLOGIA Vol. 8, no. 2 November, and in fruit in July and November. In Thailand it is "amuraon", "binh linh", "culimpapa", "kara", "ketileng", “kulipapa", and "Malayan teak". Hochreutiner describes his V. pubescens var. genuina as fol- lows: "Flores striato-coeruleo-violacei; folia trifoliata inflor- escentia plusminus corymbosa, i.e. minus thyrsoidea quam in var. alt. ca. 60 m., 20 octobre 190). Additional citations: INDIA: United Provinces: Khant 132 (N). West Bengal: Helfer 132 (B). INDOCHINA: Annam: Pierre 2224 [Lign. coll. 187] (B, B); Poilane 6259 (Ca—-38173). Cambodia: Chatillon s.n. [Prey Kedey, 19-l~1907] (Ca-~53757); Poilane 13892 (B) .co- chinchina: Germain s.n. (B); Pierre 222) (Ca—53981, Ca--5h797); Poilane 10852 (Ca—54816); Bordeneux s.ne {Bencat, aout 1905] (Ca—38886); Thorel 100 (B, Ca—5375i). Laos: Poilane 11399 (B), 16103 (B). Pulau Condor: Perry sen. [7/1870] (Ca—5i,813). Tonkin: Pierre s.n. (B, B). State undetermined: Collector undesignated Uy (8). THartanp: Bunpheng 982 [Roy. Forest Dept. 1606] (Sm); Kerr 10885 (B), 11132 (B); Put 894 (B); Rabil 2h3 (B); Smitinand 2968 [Roy. Forest Dept. 12670] (Su); Winit Wanandorn 2,9 [Herb. Roy. Forest Dept. 1994] (Vi). MALAYA: Pahang: M. R. Henderson 10750 (Ca-~360665), 1866 (Ca--318553). Penang: Hanif? 12h (Ca— 355297). Selangor: Burkill & Shah 947 (Ng--20206). Singapore: H. Ne Ridley s.n. [Bukit Tunah, 1900] (Ca—267600). PHILIPPINE IS- LANDS: Palawan: Cenabre s.n, (Herb. Philip. Forest. Bur. 29995] (Ca--29179h); Cenabre, Gellidon, & Paras s.n. (Herb. Philip. Forest. Bur. 27872] (Ca—213949); Edafio s.n. [Philip. Nat. Herb. 13999] (Mg); Re C. McGregor s.n, (Herb. Philip. Bur. Sci. )596)] (Ca--309255). Sulu: Ramos & Edafio son. [Herb. Philip. Bur. Sci. 44138] (Ca—257651). Palawan: R. B. Fox s.n. (Philip. Nat. Herb. 13428] (Mi). SUMATRA: Bartlett & La Rue 351 (Ca—2h3872); Krukoff 314 (N); Toroes 3% (Ca--91889); Yates 611 (Ca--232531), 663 (Ca- 232451), 1267 (Ca—251149). KARIMANDJAWA ISLANDS: Karimandjawa: Karta 311 (B). JAVA: Boer’ L6ha (Ca—918503); Koorders 97)1b (Ca--2659h5), 13595b (Ca--265960), 15661b (Ca—-231899); Saimoendt 38 (Ca-—307850). KAMBANGAN ISLAND: Hochreutdiner 2301 (Ca—L1278). BANGUEY ISLAND: Castro & Melegrito 1511 (Ca—21389). JAMBONGAN ISLAND: C 3801 [P.T.0. 515] (Ca--3i7069). BRITISH NORTH BORNEO: Elmer 20101 (Ca--228922, Um-—169), 21095 (Ca—312133); Ginang 163 [71; D. D. Wood 2527] (ca—32025)); Goklin l (D. D. Wood 1868] (ca—2hh262); orolfo sn. (D. De Wood 1096) (Ca— 1961 Moldenke, Notes on Vitex 7 249064); Puosa s.n. [D. D. Wood 2289] (Ca--28672h); Sales 3906 {field no. 468] (Ca--347177). LESSER SUNDA I : Timor: Bloem- bergen 19 [Boschproefst. BB.27071] (Vi). CULTIVATED: Hawaiian Is- _— H. Ne Moldenke 21851 (Z). Java: Herb. Hort. Bot. Bogor. s. Ne es VITEX PINNATA var. ALATA Moldenke Additional literature: Moldenke, Phytologta 6: 79. 1957; Mol- denke, Résumé 177, 191, 423, & 478. 1959. This plant has be ollec fruit in June and September, and has been identified in herbaria as Y. pubescens var, ptilota Dop (which actually may prove to be the same taxon). Additional citations: INDOCHINA: Annam: Poilane 21 (B), 1685 (B). State undetermined: Evrard 2525 [C&y Binh Linh, descente de Cana] (Ca—53767). oe. VITEX POLYGAMA Cham. Additi 1959; Renné, Levant. Herb. Inst. A Fl. Paran. 16: 80 (1960) and 17: 47. 1961. Reitz & Klein collected the species at 10 meters altitude. Additional citations: BRAZIL: Rio de Janeiro: Brade 14158 (Herb. Rio de Jan. 29520] (B). Santa Catarina: Reits & Klein 4210 [Herb. Barb, Rodr. 16275] (N, S, Sm). State undetermined: —— 8.n- [Macbride photos 17565, in part] (N—photo of co- VITEX POLYGAMA var. BAKERI Moldenke Additional literature: Moldenke, Phytologia 6: 86-87. 1957; Moldenke, Résumé 111, 226, & 78. 1959. Hatschbach & Lange state that the flowers of this plant are VITEX POLYGAMA var, DUSENII Moldenke Additional synonymy: Vitex laciniosa Turcz., Bull. Soc. Imp. Nat. Mosc, 36 (2): 225, 1863. Vitex laciniata Turcz. ex Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 39 & 103, sphalm. 19l2 (not V. laciniata Hort., 190, nor Host., 197]. Additional literature: Turez., « Soc. Imp. Nat. Mosc. 36 (2): 225, 1863; Moldenke, Alph. List Invalid Nanes 53. 19125 Moldenke, Known Geogr. Distrib, Verbenac., [ed. 1], 39, 103, & 10h. 1942; Moldenke, Alph. List Invalid Names Suppl. 1: 29. 19h7; Molde 9 Geogr, Distrib. Verbenac., [ed. 2), 95, 201, 202. 19493 Moldenke, Phytologia 5: 29—l30 (1956) and 6: 87-88. 19573 Moldenke, Résumé 111, 385, 387, & 178. 1959; Moldenke, R&- sumé Suppl. 1: 7 & 23. 1959; Renné, Levant. Herb. Inst. Agron. 76 P Eee OL OG-IA Vol. 8, no, 2 Minas 151. 1960; Angely, Fl. Paran. 16: 80 (1960) and 17: 7. 1961. Additional citations: BRAZIL: Bahia: Kegel 12319 (Z--photo, Z- photo). VITEX POLYGAMA var. HIRSUTA Sc Additional literature: Moldenke, Phytologia 6: 88—89. 19573 Moldenke, Résumé 111 & 78. 1959. Additional citations: BRAZIL: Espirito Santo: Sellow s.n. (Macbride photos 17565, in part] (N—photo of cotype). VITEX POOARA Corbishley Additional literature: Moldenke eae ad 63 89=-90. 19573 Molienke, Résumé 149, 15h, 388, & 478. 1959 Theron describes this as a a large tree with pitch-black stone- fruits ep agin of bedbugs. He found it fruiting in December, and Le name ‘nm ao ittonal Skee UNION OF SOUTH AFRICA: Transvaal: Theron 2049 (S VITEX PYRAMIDATA B. L. Robinson Additional literature: Mold Phytologia 6: Rew Af ee Moldenke, Résumé 1,0, 387, 388, 0 & 78. 1959; oc 2 i, bh, & & 148. 1961. trunk diameter of 30 om. at breast height. It has been found in in fruit in November, at altitudes of 66 meters, Miranda explains that the name “yashcabté" means literally "green-branched-tree". Additional citations: MEXICO: Guerrero: Hinton 9990 (Ca-— 939136), 10005 (Ur). Jalisco: Herb. Univ. Kans. Mexico Exped. E. 57 (Mi). Michoac4n: Hinton 16235 (N (N); Leavenworth & Hoogstraal 1707 on); Metaugh 15020 18020 (Mi). Sinaloa: H. S. Gentry 5073 (Ca-- 651836, VITEX QUINATA (Lour.) F. N. ¥ eee a literature: Neal Gard. Hawaii ant 19483; Mol- denke, Phytologia 6: 98-103, issn Moldenke, Résum 171, 172, 17h, 177, 1, 181, 185, 188, 191 "192, oh, 195, 198, 200, 226, 276, 361, 38 —386, 388390, 78. 1929. Recen t Steck cnamberd the bark’ “a dark-gray, the leaves light shiny-green, the flowers pink, lavender-tinged, drab", or "green and drab", the immature fruit pale-green, and the common names ar. Additional citations: CHINA: Chekiang: Ching 1987, (ca-B1 76); Ling 107) (Herb. Univ. Nanking 7311] (Ca--252262). Fukien: H. He Chung 3003 3003 (Ca--2),36)8), Kiangsi: Hu s.n. [1920] (Ca--232221) . 1961 Moldenke, Notes on Vitex 77 Kwangsi: 7309 (Ca--410038). Kwangtung: W. T. T 2119) (Gansta) eal (Ca--11589); Ying 770 (a3), os (Ca %-—-359931). AMOY ISLAND: H. H. Chung 793 (Ca—~h20312). ISLAND: E. D, Merrill 9996 (Ca--992150) Ping 18h9 [ter | cata chr. Coll. 1367h) (ca (Ca-—287160) . HONGKONG: Taam . 1532 (Ca—-82712). HAINAN ISLAND: W. Y. Chun 2118 (Herb. Univ. Nanking 7096] (Ca-- 24.3589); Chun &T Tso 0 4673 (8); Fung 20120 (B, Ca—11390); F. ce How 70858 (Ca—59482h), 7 71073 (B)3 How & Chun 70248 (B); Lau (B, Ca——52536), 153 (B, “Ca=—5252hh) 5. Lei 66 (B, ee Fh (B)s Tak 178 (Herb. Lingnan Univ. 15677] (Ca-—32608h) , 223 (Herb. Lingnan Univ. 15722] (Ca--326086), 719 (Herb. Lingnan Univ. 171,68] (Ca--356748), 868 [Herb. Lingnan Univ. 16367] (Ca—326085); Tsang 786 (Herb. Lingnan Univ. 18320] (B). INDOCHINA: Annam: Clemens & Clemens 3480 (Ca--3)0209). Laos: Poilane 15435 (Ca--51,808). Ton- kin: Ba Balansa 4885 (Ca--53755, Ca—-5907, Ca-—54812). State unde- termined: Collector undetermined 1081 [Phu Ho] (Ca—227709); Petelot 963 (Ca--223917). PHILIPPINE ISLANDS: Mindanao: Wenzel 2523 (B (B, Ca—31688), 2760 (Ca--316628), 2912 (Ca-~316955), 3430 3430 (Ca—351967) . Samar: Jager 935 (B). CULTIVATED: China: H. He Chung 2517 (Ca—233121). Hawaiian Islands: Judd s.n, [Bishop Ee- tate Nursery, Dec. 1, 1930) (B). QUINATA PUBERULA (H» J» im) Moldenke i a ages al Literatures CNC, 4 1b, 185, 6 6: ets ae pS a Be by «Oe Wy 177 ct 192, 19h 196, 200, 202, 203, 207, 226, 38 Pgh “tar, ‘ti Recent collectors describe this plant as a tree with a bole 12 m. tall and 50 om. in diameter, with buttresses to 1 m.; bark yellowish-brown, deeply fissured, in strips 2 and mm. thick, the living bark 5 mm, thick, yellowish; wood or te; brown-reds leaves light~green; inflorescence gray; flowers s odorless; calyx dull violet-tinged or yore cor plla white, "creamy— “white, deep-blue inside", "white and yellow", ed at the inside of the i equal petals, the lip violet-blue"; stamens long, white, or the filaments dull viol let-tinged and the anthers violets pollen whitish; style sordid-whitish; unripe fruit gray~green or greenish, dark-brown when ripe. It has been collected dense jungles and in open forests. On Morotai it is found in stony clay soil of primary forests in flat July, and in —— in November. Additional common names are "drai tolu® 7. Herbarium material has been misidentified and distributed as Le Sumatrana var. urceolata (C. B. Clarke) King & Gamble. Merrill says of Castro & Melegrito 1592 "a form with trifoliolate 78 PHYTOLOGIA Vol. 8, no. 2 leaves, India to Formosa through oe to Celebes" — see his Emm, Philip. Flow. Pl. 3: 3% (1923 Additional citations: CHINA: Kwangei: Ching 5552 (Ca-~10181). Kwangtung: Peng, Tak, & Kin 61) (Herb. Canton Chr. Coll. 12613] (Ca—27hh34). FORMOSA: Matuda 9 901 (Ca—345187). HAINAN ISLAND: Lei 71k (Ca--612)95). INDOCHINA? Annam: Poilane 1640 (B), 1107h (B). Tonkin: Balansa 1,885 (B); Pételot 977 (Ca——223923)3; Poilane 13060 (B). Fiona Bunclmai 76 (2). (Z)~ MALAYA: Perak: King's col- lector 75 529750). JAVA: Backer 17102 (B); Koorders 9620b Teac) re (Ca--2659)7), S8Lib (ce~ (Ca—2659h9); Saimoendt 63 (Ca~~307849). BANGUEY ISLAND: Castro & Melegrito 1592 (Ca-— (Ca-— 241557). BRITISH NORTH BORNEO: M. S. Clemens 1027) (Ca—-214817); Sales 3912 (Ca-—3)7168). BORNEO: | M. [. Jacobs Estan (Ba); Kostermans 6895 (Ng--1687h, Ng, Ng). CELEBES: Koorders 195S1b (Ca—265951). MOIUCCA ISLANDS: Morotai: Tangkilisan 200 (Ng—16935). Obi: Beth. Forest. Serv. BB.23819 (Mg——-56523) .NEW GUINEA: Dutch New Guinea: C. Koster BW.1121 (Ng—16910); Mangold B.W.2082 (Ng—20199), B.W.2106 (Ng—20193); Schram B.W.1848 (Ng—-20200), B.W.1869 (Ne— es B.W.1907 (Ng—-20198), B.W.1965 (Ng—-20196, Ng). Flgl IS~ ti Leva: O. Degener 14h81 (Ca- ome J. We Rat 2117); A. C. Smith 1307 (S), 6295 (S). CULTIVATED: Java: rat: Hort. Bot. B Ore | S.n. ee (B). TEX RADULA Mildbr, Additional literature idenke, Phytologia 6: 108. 1957; Moldenke, Résumé 15, "350, ". iy i959. er citations: H NYASALAND: Stolz 1398 (B= VITEX RAPINI Beauvi Pre yee pasion Vitex rapinii Beauvis. ex Moldenke, Ré- sumé 388, in syn. 1959, Additional literature: Guillaumin, Marseilles 4 205. 1911; Moldenke, Phytologia 6: rahres 1957; “VWoldenke, » 342, 388, & Me 1959. Additional citations: NEW CALEDONIA: Pancher s.n. (B). VITEX RAPINIOIDES Guillaum. Additional literature: Moldenke, Phytologia 6: 109--110. 19573 sa pm Résumé 205, stg & 78. 1959. tional citations: NEW HEBRIDES: Efate: Kajewski 211 (N). Eromanga: Kajewski 299 (MN). REHMANNI Gttrke Additional literature: Moldenke, 112—113. 19573 Phytolo ae Abstr. 29: 3602, 19575 Moldenke, be eine gee 388, & 478. 1961 Moldenke, Notes on Vitex 79 The species has also been collected in anthesis in January and in fruit in June. Additional citations: UNION OF SOUTH AFRICA: Transvaal: Sidey 2477 (S)3; Smts & Pole-Evans 860 (S); L. E. Taylor 482 (Ca— 9560L5) EX REHMANNI f. SUBTOMENTOSA Moldenke Additional literature: Moldenke, Biol. Abstr. 29: 3602. yal? Sinaia 6: 113--11h. 1957; Moldenke, Résumé 15), 388, & 478. 1959. RIVULARIS Gttrke Additional literature: Moldenke, Phytologia 6: M-«116, 1957; Moldenke, Résumé 137-139, 143, 382, & 478. 1959. ditional citations: CAMEROONS: Zenker 2691a (B), sen. [Bi- pindi ] (Ca—620105). VITEX RUBRO-AURANTIACA DeWild. Additional literature: Moldenke, Phytologia 6: 117-118. 1957; Moldenke, Résumé 143 & 478. 1959. The species has been found at 70 meters altitude, blooming in Sep ° Additional citations: CONGO: Louis 29h, (Ca-—962327), 5786 (Vi, Vi), 5900 (Ca--962275)., arc Additional literature: Biol. Abstr. 30: 1703. 1956; Assoc. Etud. Fl. Afr. Trop. Ind. 1955: 63. 1956; Moldenke, Reytaices 6; 118— 119. 19575 A. Fernandes, Mem. Soc, « 13: 3h. 1958; Biol. Abstr. 30: 4370. 1958; Moldenke, Résumé 137 & 178. 1959 A » the Cavaco, Bull. Mus. Nat. Hist. Nat. Paris, sér. 2, 27: 91 reference cited by me in Phytologia 6: 118 (1957) as "1956" should be dated "1955." denke, Résumé 171 & 1,78, 19 Additional items of description: bracts and bractlets linear, 3- ong ° , ehtly cucullate, glabrous inside, the small lip reflexed, cleft, rous 80 PHYTOLOGIA Vol. 8, no. 2 Additional citations: CHINA: Kwangtung: Ying 782 (Ca-~358297). VITEX SCHLIEBENI Moldenke Literature: Moldenke, Phytologia 7: 85-86. 1959; Assoc. Etud. Tax. Fl. Afr: Trop. 1959: 53. Synonymy: Vitex schliebenii Moldenke ex Assoc. Etud. Tax. Fl. Afr. Trop. Index 1959: 53. 1960. See the first reference cited above for complete description of this species. Citations: TANGANYIKA: Schlieben 6008 (B—type, Z—isotype). VITEX SCHOMBURGKIANA Schau. Additional literature: Moldenke, Phytologia 6: 125--126. 1957; Moldenke, Résumé 76, 112, 323, 382, & 478. 1959. Irwin describes this as a low tree es found it on a savanna at the edge of a rainforest , growing in ine brown sand. Herbar- ium material has been misidentified and rpttt as Ve capita- ta Vahl. ~~ Additional citations: BRITISH GUIANA: Irwin 889 (au—165609). VITEX SECUNDIFLORA H. Hallier tional literature: Moldenke, whys 6: 128 (1957) and 63 129. 19583 Moldenke, Résumé 191 '& 478. 1959. ies has been encountered at alti tades of 100 to 300 me ° Additional citations: BORNEO: Dachlan 37 [Z. 0. B. 35133 Boschproefst. BB.10715] (B, Ca——358),0). SELLOWIANA Cham, Additional literature: Moldenke, Phytologia 6: 130=-132. 19985 Moldenke, Résumé 112, 226, mr & “ly78. 19595 Renné, Levant. Inst. Agron. Minas 161; VITEX SIAMICA F, ee — Additional li , Phytologia 6: 132-133. 19583 Moldenke, Résuné 178 178, 1 181, " LTB, “Soe9, This ies has been looming also in N ovember Additional citations: sors Langkawi Islands: Holttum 15095 (Ca--283987). Selangor: Nur 34381 (Ca—3105). VITEX SIMPLICIFOLIA Oliv. Additional literature: Mol P dag 6: 133-135. 19583 ed Résumé 133, 13h, Die, 383, 389, & 478. The Latilo & Daramola 31,90 distributed as this is cited by me as var. vogelii OF Ge Baker) Pieper. VITEX SIMPLICIFOLIA var. VOGELII I (J. G. Baker) Pi Additional literature: Moldenke Phytaloess 6: “T3516. 19583 Moldenke, Résumé 138—1)0, 391, & “478. 195) Recent collectors have found this aks oe open savannas, with 1961 Moldenke, Notes on Vitex 81 very small green fruits in Fe ebruary. Herbarium material has been misidentified and distributed as typical ¥V. simplicifolia Oliv. Additional citations: MALI: Soudan: P. J Jaeger 5778 (Z). CAMER- OONS: Latilo & Daramola 34490 (B). RUCEI Additional literature: Phytologia 6: 137139. 19583 Moldenke, Résumé 112, 387, ‘2, & y, & hb. 1959 Collectors describe owers of this plant as white and ar- omatic. It blossoms —— 7 pf Additional citations: BRAZIL: Amazonas: Coelho s.n. (Herb. Inst. Nac. Pesq. Amaz. 1984] (Z); Due Ducke s.n. [Herb. Rio de Jan. 22577] (B) $ Francisco s.n. [Herb. Inst. Nace Pesq. Amas. 2063] ar 3 Frées 2226) (Ut—951,52a); Herb. Inst. Nac. Pesq. Amas. 1836 Mati SPRUCEI var. VAUPESENSIS Moldenke my ied Moldenke, Phytologia 6: 332. 1958; Moldenke, Résu- ns Py & 478. 1959. S variety differs from the a form of the species in ae ng its leaflets uniformly 3 in number. The type of the variety was collected by Bassett pe oat John J. Wurdack, and W. M. Keith (no. 41839) on a small savanna 0.5-- 1.5 km, north of Puerto Colombia, on the nfo oak a opposite to » V Colombia, at an elevation of 130 meters, on Octo- ber 12, 19k7, and is aeons in the Britton Herbarium at the New York Botanical Garden, The collectors describe the plant as tree pee meas tall, the corollas white, the lip with a cieeh ow spot, d state that it is frequent in the type locality. Citations: ci COLOMBIA: Vaupés: Maguire, Wurdack, & Keith 18 (¥—type, N--isotype, S—isotype, S--isotype). VITEX STAHELIT Moldenke Additional Aiterature: Mol denke, Phytologia 6: 140—1)2. 1958; Moldenke, Résumé 524 Thy Ty 7, 389, & & L78- 1959 Additional ci ta Amacuro: Squires 257, in citatlons! Wenz: be VITEX STRICKERI Vatke & Hildebr. Additional synonymy: Vitex parvifolia me ex aren Résu- mé 387, in syn, 1959. vitex strieker! Vatke in he Additional literature: 1 odie 6: "Gr 1 1958 Mol- denke, Résumé 145, 146, 387, 389, & 478. 1959. Recent collectors describe this plant as a liana to 15 meters shru + Rusby & tree to 8 m, tall, profusely leafy, the whole plant aromatic, the young stems with dense brown indumentum; stems erect, sharply ar- fmatic; sap crite tips of twigs with orange pubescence; cal- browmish-gr. orolla » whi een cream-colored to Aceon 2 or pale~pink, slightly aranatic; fil- 82 PHYTOLOGIA Vol. 8, no. 2 aments cream-colored; anthers brown or buff; fruit green with a 8. species has been collected in scattered-tree grasslands, at the Linea of pi pen forests, at the margins of thickets in woo el and black cotton soil in thick forests, st on loose brow soil at the edge of thick cover, from sealevel to 1275 gs altitude, blooming in March, April, and July to September, and December, “fruiting in April, June, and July. The leaves are cooked and the resulting water a used to rinse the mouth in the treatment of geet lon gums. name is of Peter 3322) is the type collection of V. parvifolia Peter. Herbarium material has been misidentified and distributed as V. lamiana Pieper, Premna sp., Clerodendron sp., and even Allophylus sp. in the Sapindaceae Additional pitaticne’ TANGANYIKA: Busse 2163 [A. Peter 51807] (B); Drummond & Hemsley 1810 (S), 3579 (S) (S)3 Het H. G. Faulkner 563 [12.444.1950] (8), 563 [5.iv.1950] (S); Merker Tih (B)5 Ae (B); A. Peter 1417 [0.1.35] (B), 16335 [0.1V.43] (B), 16684 [0.IV.52] ~(B), 20672 [0.IV.191] (B), 2088 [0.IV.195] (B), 248h5 [0.IV.340] (B), 3322) [V.9h] (B), 39460 [V.203] (B), 42239 [V.282] (B, 2); Schlieben 5623 (B); Tanner 630 (Ca--178110), 2065 (Ca—170h65). KENYA: Bogdan B.57 (Ca~-9h35h6). VITEX SUMATRANA Miq. Additional literature: S. Moore in Rendle, on Suppl. 81. 1925; woldence, Ph Prytologia 6: "sits. ° a & Bakh., Bull. Jard, sér. 3, 3: Sh--55. 19213 Moldenke, Résumé 166, 7288, "pl, 1,365, & 478. 1959. VITEX SWYNNERTONII S. moni Additional literature « Broter. 13: 7h. 1908 Moldenke, Phytologia 6: Us 1s6. 1958; ee, Résumé 151 & 478. 1959. VITEX TANGENSIS Additional literature: Moldenke Ber talens 6: 16-147. 19585 Moldenke, Résumé 5, U6 4 tSby, 388, & . 1959. Peter describes t neat 3 m. tall, with pink flowers, blooming yo ror ee Additional citations: TANGANYIKA: A, Peter 39696 [V.211] (B, Were Tare BSTYLOgA Pe aie s terature: Mold Bae sg ia 6: eR a j Moldenke, Résumé 136-139, 3. 2 15k, 387, 389 Additional citations: IVORY COAS : pbepeie neon a peek OONS: Zenker & Staudt 355 (B). 1961 Moldenke, Notes on Vitex 83 TOMENTULOSA Moldenke dditional literature: Moldenke, Phytologia 6: 15—155. 1958; ; Moldenke, Résumé 53 & 78. 1959. Alain describes this as a shrub, 2 m. tall, inhabiting the coastal thickets, Additional citations: CUBA: Pinar del Rfo: Alain 6915 (Bm). VITEX TRIFLORA Vahl onal literature: Moldenke, Phytologia 6: ag 1958; ae Résumé 7, 76-78, 85, 1i2, 115, 226, = 27h, 319, Asplund describes this plant as a tree 20 a , With the corollas pale-lilac, their lobes light~blue. Additional citations: BRITISH GUIANA: Little 16618 (Bm). SURI- MAM: Be Maguire 21,837 (Ur). PERU: Hudrmco: Asplund 12729 (S). BRAZIL: Amazonas: Cha; Chagas s.n. (Herb. Inst. Nac. Pesq. Amas. 3970] (Bs); Luis s.n. (Herb. Inst. Nac. Pesq. Amaz. 1165] (Bs). VITEX beearte var. KRAATZII Additional literature: eaaence Phytologia 6: 163-16). 1958; Woldenke, Résumé 112, 383, & 1,78. 3959 96 12972), tional citations: BRAZIL: ard: N. T. da Silva 2 (Ca~ TRIFLORA Additional literature: M , Piytologia 6: 164—-165. 1958; Moldenke, Résumé 85, 112, 390, & “e y8, 1959. tional Aisations : BRAZIL: Acre Territory: Krukoff 5765 TRIFOLIA L easy synonymy: Vitex trifolia var. trifoliolata H. J. in herb, Additional and emended literature: Drake del ene Fl. Polynés. Frang. 151. 1893; Valet., Bull. Dept. Agric. Néerl. 10: 51, 1907; Guillaumin, Ann. Mus. Colon. uatseitic 93 6 & 206, 1911; F. M. Bailey, nee he Cat. Queensl. Pl. 386. 1913; Neal, In Gard, Hawaii ond veo E. H. Walker, Proc. Pacif. Sci. Cong. : 102. 1957; Menninger, 1958 Price List [7]. 19583 Menninger, 1959 Price A “téle 19583 Moldenke, Piggelen gla 6: 165~17),, 1958; Moldenke, Résumé ish, 1 157—159, 165—16 171, 173, 175, 178, 181, 182, 185, 186, 188” 189, 191. 932196, ar? 200, 202—207; 211, 212, 226, 249° 2507 302, 335, 379, 380, » 385, 389—391, 4,23, 27, & Ai. 19593 Moldenke, Résumé Suppl. 1: 10 & 12, 1959 Hara geog. Japan 8. 1959 Price tie Flow, Trees [10]. 1960. Recent collectors call this a s e shrub, or a tree li ie chee ae in January laces open p in Photograph accompanies the Yates I9lit specimen in the Uni- versity of California herbarium . Hara, “in the reference cited 8 PHYTOLOGIA Vol. 8, no. 2 above, records u — from the island of Amani-oshima in the Liukiu Islands. have not seen the material Pi which he bases this record, oy Sonos that it will prove to be var. bicolor (Willd.) Moldenke. The Boeea 551, Brass Bsus, Blackburn s.N., Bangham & Bangham 615 & 508, Cha Chapin n 853, 0. D 0. Degener 9505, Franc 102h, K Kuntze ze bbkS & & 5817, H Herb. Mus. Yale , School Forest. 2266) A. C. Smith Smith 4559, 6078, ° & 6610, F. K. Ward 8929, and Waterhouse 60, “all I previously di “distributed and even in some cases cited =: as V. trifolia, are var. subtrisecta (Kuntze) Moldenke, as is also a a Se Sone [Timor], distributed as V. trifoliata Lam. . (Stuart, April 17, 1945] cited by me in agit a 6: ae Gsz8) as V, trifolia is actually var. subtri- secta, as are probably the plants referred to by Menninger rin the several works by him listed in the above ty ep I The V. trifolia of Neal, In Gard. Hawaii 641, called by he hlue vitex", is var. subtrisecta, as examination of ‘eae speci- mens in the herbarium of the Bishop Museum, Honolulu, has shown. The Kanehira 2273 distributed as V. trifolia is actually vare bicolor, while e Kajewski 8 is var. , variegata Moldenke. Drake del Castillo records v. trifolia from Noukahiva, Mar quesas Islands, on the basis of a “Le Bastard" collection not seen by me. He says for its geographic distribution "Regions chaudes del'Anci - Monde", Guillaumin cites a Pennel 300 also not yet seen by m Additional Peeeitiais MADAGASCAR: Viguier & Humbert 1,39 bis (B). SEYCHELLES ISLANDS: Mahé: Desc BeMe . (22 fév. 1892) (B). INDOCHINA: Cochinchina: Thorel 2092 (B)- PHILIPPINE ISLANDS: Mindoro: Conklin s.n, (Mt. Yagaw, May h, 1953) (Mi). SUMATRA: Dorsett & . Fairchild 1a 199 pe Te Yates 524 (Ca--234112), 1,80 (Ca—263981), 263981), 1941 (Ca--287207). JAVA: Koorders 28038b (Ca— 265959) « nay Btnemeijer 10633 10633 (Ca--234985). NEW | HEBRIDES: Tanna: Kajewski 66 (N). NEW CALEDONIA: Franc 56, in part (Ca 390572, N), 1201 (Ca—390603). TONGAN ISLANDS: 3: Bua: H, EB. Parks 16178 (N). FIJI ISLANDS: Ovalau: J. W. Gillespie 4503 3 (N). SAMO- AN ISLANDS: Tutuila: Garber 611 (N) . AUSTRALIA: New South Wales: Boorman s.n. [Tweed Heads, 1-191)] (Ca-—18268)). CULTIVATED: Indochina: Pierre 389 (B). TRIFOLIA var. BICOLOR (Willd.) ggg 2 ——— literature: Guillaumin, 3 19 3 genet pares 6: "e380. “195 Pact: + Midl. Nat. 59: 366. 1958; Yuncker, Bisho 220: 232. 19595 uoidenis, Résumé 155, 157, 165, 187, 25, 181, 282, 185, 186,189, 191, 19)--196, 200, 202-207, 211, 212, 226, 380, 381, se 3072-389, 427, ‘: he. 1959. Recent collectors descri Te IO Fe a paets 0 1961 Moldenke, Notes on Vitex 85 —_ slemder, much-branched tree, the bole to 1.5 m, tall ° 20. cm. in eter at breast height; bark smooth, cinera- pa ag sapwood hin, brown; heartwood yellow-brown; leaves green or pale gray-green above, varying from me ae grayish-green, glaucous-gray, or white beneath, those close to the inflorescence 0. pale lilac-blue, ns ty ane ee "blue hite", "deep-blue paler outside", or tish when Pate [Evangelista 362 32); young fruit Seigete:, Soph It has been found by c seri actet on eet seashores, dry shore lines, seacoasts, along the strand, in forests and secondary for- ng s in seas hi of mangroves"; on Hayman Island it is "common on beach at edge of light Taliihobeks #2 and ly the Pelew Islands "very common on the seashore”, It has been collected in flower and fruit in July. th this concoction and tied tight. ahs Franc 56 cited below from the University of California her- os Ae is definitely this variety even though other specimens of the same number are cited by me as the typical form of the species. Guillaumin cites a Pennel 300 not yet seen by me. Additional common names are “drala", "dralakaka", "gregit", tahi®, "mulokaka", and "namrlega". iadititna citations: INDOCHINA: Cochinchina: Godefroy s.n. (Ba yoo] (Ca—53719). PHILIPPINE ISLANDS: Mindanao: Ahern 671 (Ca—260119); Guerrero 25 {Herb. Philip. Forest. Bur. 30388) (Ca- 321155); Florita 20 [H 20 [Herb, Philip. Forest. Bur. 29757] (Ca— 257032). Panay: C. Edafio s.n. [Herb. Philip. Bur. Sci. 46232) (Ca—309671). MARIANNA ISLANDS: Tinian: R. S. Cowan s.n, [April 3, 1945} (Ca—-995191); Kanehira 2273 (N). CAROLINE ISLANDS: Uman: BANGUEY ISLAND: Castro & 0 & Melegrito to 1636 [Melegrito 130] (Ca—- 21652), BRITISH NORTH BORNEO: Apostol 29 [D. D. Wood 1820] (Ca-— 244323); Evangelista 362 [D. D. Wood 2589] (Ca-320376); Ginarg 193 [D. D. Wood 2502] (Ca—320280). CAROLINE ISLANDS: Arekalong: Taman 1697 (Ca-~805550). MOLUCCA cama Ceram: fornasst = autten 111 (Ca—-2659h6). NEW GUINEA: Dutch New Guinea (Ne—16912), Papua: Brass 21928 vate erg » 255U8 a tag (S)5 Womersley & & Gray 8617 y 8617 Y (Ne—168hh, Ne--168h))~ SOLOMON ISL ISLANDS: 86 PHYTOLOGIA Vol. 8, no. 2 Guadalcanal: J. C. Riley 52 (Ca——7h903). NEW CALEDONIA: Frane 56, in part (Ca--390609); C. Skotteberg 9 (Go). TONGAN ISLANDS? Ton- gatapu: Setchell & Parks 15298 (Ca—l03218), 15551 (Ca--297563), 15578 (Ca--297638). FIJI ISLANDS: Ovalau: Je W. Gillespie 503 (Ca—l,8981, B). Ththia: E. H. Bryan 556 (Ca—805552). Kandavu: A. C. Smith 31) (Ca—601858). Varma Levu: Degener & Ordon 1,058 (N)5 Ae Co Smith 6622 (S). Viti Levu: 0. Degener 13691 (N), 150hh (N)3; Ordonez Sn. (0. Degener 13620] (§); H. E. Parks 20857 (Ca-- 47555). SAMOAN ISLANDS: Safotu: Vaupel 389 (Ca—-882869). Savaiis Christophersen 2849 [J. Bower s.n.] (Ca--592222). Tau: D. W. Gar- ber 611 (B). Tutuila: Stttupe 531 (Ca--216023). Upolu: A. J. Eames 36 (B, Ca—$92h,36). NIUE ISLAND: Yuncker 100k] (ca—-882668). AUSTRALIA: Queensland: F. Mueller s.n. [Endeavour River] (Mn-—- 1536). HAYMAN ISLAND: C. T. White 10167 (NW). COOK ISIANDS: Raro- tonga: Parks & Parks 22112 (Ca—393363), 22573 (Ca-—393331). VITEX TRIFOLIA var. BICOLOR f£. ALBIFLORA Moldenke, f. nov. Haec forma a forma typica varietatis corollis albis recedit,. This form differs from the typical form of the variety in having white corollas, The type of the form was collected by Harold Ernest Parks (no. 16178) at the strand margin, Ohoma, Eua island, Tongan Islands, on June 27, 1926, and is deposited in the herbarium of the Uni- —s of California at Berkeley. It is described as a shru ) di form, since the collector notes that the flowers were "whitish when fresh", ‘. 5 hic se TONGAN ISLANDS: Bua: H. E. Parks 16178 (Ca—297255- ype ‘ ee Restennmnteneeeneell Benth. ex F. M. Bailey, Compreh, Cat. Queensl, Pl. 386. 1913. Vitex rotundifolius Hara, Outline eogr. Japan 56, sphalm. 1959. Vitex rotundifolia var. rotundifolia Mizushima, in herb. Vitex trifolia var. ovata Schan., in herb, Additional literature: Doney, Brooklyn Bot. Gard. Rec. 30: 23. 19h1; Neal, In Gard, Hawaii 61,2, fig. 275. 1948; Ikeda, Journ. Jap. Forest. Soc. 37: 334-335. 19553 Satoo, Biol. Abstr. 30: 2695. 1956; Biol. Abstr. 30: 1370. 1958; Moldenke, Phytologia 6: 183-192 & 197. 1958; Hara, Outline Phytogeogr, Japan 55, 56, 58, & 86. 1959; Moldenke, Résumé 112, 157--159. 166 167, 171--175, 177, 179, 181, 182, 185, 189, 191, 193, i9h, 196, 198, 200, 202, 203, 205, 211, 212, 226, 379° f 390, & 479. 1959; O.&I. Degener, Pac. Disc. ly (55: 15. 1961. Additional illustrations: Neal, In Gard. Hawaii fig, 275. 198. 1961 Moldenke, Notes on Vitex 87 Doney, in the reference cited above, says that the hardiness of this plant cannot be depended on in Broo ooklyn, New York. Ikeda reports an unidentified species of Bacterium and of Alternaria as parasites on this plant in Japan. The M. R. Henderson 18415 has galled leaves. Recent collectors describe the Aas as a prostrate "herb" or strongly aromatic decumbent shrub » sometimes woody oe erect, with stems several meters te peottie at the nodes, th leaves aromatic, green or gray-green above, glaucous-tamentose a neath, and the corollas lavender, mauve, deep~purple, or light~ throat violet, lemon-yellow in the e It is described by collectors as common on coastal dune » sandy beaches, sand near shore, sandy places near the seashore, on “top of the beach", and prostrate on sand just above highwater mark, It has been found at altitudes of sealevel to 185 meters (the latter in Chekiang, C Additional vernacular names are "homa-go" and "pohinahina", McClure describes it as a "vine cre reeping on dry beach". Clemens & Clan ens 3028 specimen in the University of California herbarium wg a notation by E. D. Merrill "Vitex trifolia L.; Lour. p. 390 pro parte, quoad habito", The Alexander & Kell & Kellogg 2300 label says "not in bloom", but flowers in full anthesis are quite in evidence on the University of California specimen of this numbert Chun 212, Pételot 1409, and M. R. Henderson 1815 have petioles longer and more slender than is 3 typical for the variety. The Hom A.387 [Herb. Lingnan Univ. 1879h] cited in ee 6: 19 197 (1958) as this variety is actually var. subtri- (Kuntze) Moldenke. Secta ~~ Additional aitatioie: CHINA: Chekiang: Ching 1967 tg 281898). Fukien: H. He Chung 2031 (Ca--232779). McClure 117) [Herb. Canton Chr. Coll. 13095) (Ca—287863) . Fi tung: F. i. Meyer 380 Serge sdb AMOY ISLAND: A. N. Steward 30h (Ca-lis773). QUELPART ISLAND: In-cho 112) (Mi). FORMOSA: 8. Sasaki Sen. [Kelung, July 31, 1927) (Ca——344332). JAPAN: Honshiu: Dahistrand s.n, sn. (Kanagawa, 23/9/1950] (Go); Hayakama 1.2502 (Ca~ 320631); Kobayashi 1033 (s), 14133 (S); Lindquist s.n. [Akita, 8/9/1952) (Go), s.n, [Nakagoya, 9/9, hele 1: Go); Maruyama & Oka- eee Ca—163615, Go, N, S); Sasaki & Togasi 606 (B, Ca 355 Go, Mg, S, Vi); S. Suzuki 136 [Sep. + 1952; Herb. Sumuki is50ih) (¢ (Ca—-6032), Kiushiu: Ichikawa 200132 [Tanaka 120] (Ca— 2182). Island perp pie “Hiroe 5887 [Karo, prov. Inaba] Rectan ns : Mrs. K, R, Smith s.n. [August 1929] (Mi). HONGKONG: Chin 2 212 12 (caBIG tae LANTAU ISLANDS: Taam 1702 (B, Ce-~62298). Ta INDOCHINA: Annam: A. Chevalier 1418 (B); | Clemens & ens as (Ca——305)); Pételot 109 (Ca——223788). Cochin china: | sen, [1 oct 1875] (B). MALAYA: Pulau Tioman: M. R. ees 18L15 (Ca~-318680). PHILIPPINE ISLANDS: Luzon: De 88 PHYTOLOGIA Vol. 8, no. 2 la Rosa s.n, [Gates 849] (We); Haenke 576 (N). NEW GUINEA: Dutch New Guinea: | H. S. McKee 1769 (Ng—-16879). HAWAIIAN ISLANDS: Kauai: A. Ae Heller 2 2731 (Ca—139766) ; C. He Hitchcock SMe (Dt); Ae Re Moldenke ke 86 [H. N. Moldenke rr (2); M. Reed M. Reed sen. [Aug. ql, 139 139 (Go); M. Reed son. (Apr. 13, 1905] (Ca, Ca—237091); Setche: Setchell & Setchell s.n. [Kailua Beach, July 12, 192k] (Ca—-2h6961); R. Sin sn. [Kruckeberg 129] (Du—377938) 5 Topping 3012 (Ca-—305369). Island undetermined: C. N. Forbes s.n. (Ca--502395). AUSTRALIA: ro haere Territory: Specht 677 677 (Ng--6500) . Queensland: Brass 1919 VITEX TRIFOLIA var. SUBTRISECTA (Kuntze) Moldenke, comb. nov. Synonymy: Vitex agmus-castus ¢ subtrisecta Kuntze, Rev.-Gen. Pl, 2: 510 & 511. 1891. Vitex rotundifolia var. heterophylla Mak., Ill. Fl. Nipp. 186. 1940. Vitex trifolia var. heterophylla (Mak.) Moldenke, Phytologia 3: 178. 1949. Vitex Vitex trifolia var. subincisa Kuntze, in herb, ? Vitex trifolia var. ar, acutifolia Benth. in A e 53 ‘le Bailey 1913; Moldenke, Phytologia 6: 180-183. 19585 Moldenke, Résumé 157, 165, 166, 171, 173175, 177, 17, , 182, 185, 189, 191, en 198, 200, 202, 206, 207, ou, oe 26, sc 388, 390, & 478. Recent collectors describe this plant a spreading bush or shrub, 1 to k m, tall , or less, of rather straggling growth when not kept trimned, or a small tree to 6.5 m. tall sp leaves dull-green and aromatic or very pale-green to even whitish beneath; flowers with a spicy fragrance; corollas bilabiate, pale- purple, purple, reddish-purple, ligh t—purplish, or purplish-blue to pale-blue, lilac, orchid-color, heliotrope, or pale-blue with a white throat, the labellum dark- -purple above; filamen pci? pale-purple; anthers deep-purple; style pale-blue; unripe green. er, and in fruit in May and from October to Decemb s, while in N ern detten it is said to be very common on the sandy banks of rivers; in Fiji it grows on grass-covered slopes. It is very often culti- vated as a hedge plant in the Hawaiian Islands » Where it was ob- served by my wife, my son, and myself many times in August and 1961 Moldenke, Notes on Vitex 89 September, sir apes In Queensland it occurs on golf courses and is widely plante a hedge; on Rurutu it occurs along roadsides Vanua Levu. The Banghams encountered it in rocky pasture land on Sumatra at 3800 feet altitude, where, they say, iti reg: often used for hedges by the natives. rack rege’ names recorded for it are ee "lagondee", ", "salah puteh", "salt-bus y and Meietcanate®. Moninger reports it as used Ha medicine on H I and refers to the fruit as "a hard *® (actually, sland it is a drupe). The "flowers" are said to be "green" by Native collector [Muka Hill ground, 20.1.28], but surely this is an er- ror for "fruit", Herbarium material has been misidentified and distributed as V. negundo L., ¥. rotundifolia L. f., V. trifoliata L., V. tri- folia L. a L. and its var. ovata Thunb., var. simplicifolia Cham., var . unifoliolata Schau. » and var, variegata Moldenke. Buchholz 1536 was originally identified as "a plant of the mint family". The | A. C. Smith 1559, cited below, was erroneously cited by me in Phyto- logia 6: 173 (1958) as typical V. trifolia, as was also Sen. (Stuart, April 17, 1945] on page 17h, while Hom A.387 was Seeprutr cited on page 197 as var. sinpltseteiae: appears to me now that I was in error opting Makino! wither "heterophylla" for this taxon in a rtloei 6: 180-~183 (1958). Makino's V. rotundifolia var. heterophylla was published pub. ust used. Kuntze's type has been examined by me and is not "typical V. trifolia" as stated by me in Phytologia 6: 172. Linnaeus' type of V. trifolia, also examined by me, definitely shows all the leaflets distinctly long-petiolulate. It seams to me that the trifoliolate specimens with sessile leaflets, er regarded as typical V. trifolia, mst be excluded from that taxon and placed in var, subtrisecta, which has its leaves eA 1- or foliolate. When only 1-foliclate, the leaflet is distinctly long- petiolate, while a var. simplicifolia it is sessile, subsessile, or very short-petiolate. Je ¥. Olllesie 4380, M. S. Clemens 11067bis, Collector undes- ah a 902, Hiroe 13289, a and d Moninger 219 have the leaves all 1- ° Olate an and give the appearance of a lon dolate form of var. simplicifolia. et pp ong-pet: Bentham, in the reference cited above, describes his var. acu~ tifolia as decumbent or erect; leaflets 3 or sometimes 5 or only 1, ovat Noe or ovate-lanceolate, acute or acuminate, the middle one ° we are now calling var. subtrisecta, but not hav- 90 PHYTOLOGIA Vol. 8, no. 2 ing seen the type as yet, I hesitate to say so definitely. If it proves to be the same, then Bentham's varietal name will have to be the one adopted as the oldest one in that category. Additional citations: FLORIDA: Monroe Co.: Stern & B. 301 (Du--)18359). BURMA: Shan States: Rock 2325 (Ca~-26,322) . CHINA: Kwangtung: F. A. McClure 1208 (Herb. ~Ganton Chr. Coll. 13131] (Ca—-2876)1). FORMOSA: Collector undesignated 902 (Ca— 345188). JAPAN: Honshiu: Hiroe 13289 (Ca—-180307). HAINAN ISLAND: Fung 20300 20300 (B, Caw-11545)3 Lau 270 (B, Ca--525029). INDOCHINA: Annam: Eberhardt 3291 (B). Laos: F F. K. Ward 8929 (N). State unde- termined: Thorel 2092 (Ca--53760). PHILIPPINE ISLANDS: Luzon: El- mer 1523 (Ca--272331, Um--170). SUMATRA: Bangham & Bangham 65 (NW), 806 (N); Boeea 551 (N); Toroes 910 (Ca——l,2233). JAVA: Kuntze 5817 (N-type). JAMBONGAN ISLAND: Cabiling 3691 [555] (Ca—- 347149). SARAWAK: Native collector sn. [Muka Hill ground, 20.1. 28) (Ca--357311). LESSER SUNDA ISLANDS: Timor: Herb. Mus. Paris s.n. [Timor] (T). NEW GUINEA: Northeastern New Guinea: M. Me ag? Clemens 11067 bis (Ca--180821); Hoogland 5172 (Ng—-8325); Womers Womers- ley 47 711 (Ng--16953). Papua: Je Ce Saunders 1: 127 (Ng--16823) . HAW- ATIAN ISLANDS: Kauai: H. N. Moldenke 21847 (Z). Molokai: H. N. Moldenke 21832 (Z). SOLOMON ISLANDS: Bougainville: Waterhouse 60 [Herb. Mus. Yale School Forest. 2266)] (N). NEW CALEDONIA: J. Ts Buchholz 1536 (Ca--79926); C. Skottsberg 5 (Go). LOYALTY ISLANDS: Franc Frane 102) (N). FIJI ISLANDS: Vanua Levu: A. C. Smith 6610 (N). Viti Levu: J. W. Gillespie 1380 (Ca—l)8812); H. E. Parks 20800 (Ca-hh7572)5 A. C. Smith L559 (N, S), 6078 (N). SAMOAN ISLANDS: Tutuila: Kuntze 23006 (N). AUSTRALIA: Northern Territory: Specht 42 (Ng—6589). Queensland: M. S. Clemens s.n. [12 November 1918] (ih); Podenzana s.n. [Cooktow, | “|. 1892 +1892] (Go). AUSTRAL ISLANDS: Rurutu: Chapin 853 3 Chapin 853 (N). CULTIVATED: Australia: C. T. White 12402 (Ca--937616) . — Hom A.387 (Herb. Lingnan Univ. 18794) (MN) Florida: Buswell s.n. (Miami, June 1h, 1941] (Gg--339298); Men- Sen. lieu “pri 17, 1945] (H)» Hainan Island: Moninger 219 (Ca——223866). Hawaiian Islands: 0. Degener 9505 (N). Java: Kuntze Wbh8 (N). Malaya: Haniff 29 (Ca--355172). New Guinea: M. S$. Clemens 8209 (B). LOCALITY OF COLLECTION UNDETERMINED: Black- burn s.n, 8.n. (tT). VITEX TRIFOLIA var. SUBTRISECTA f. ALBIFLORA Moldenke, f. nov. ae — 2, toma typica varietatis corollis albis recedit. orm ers from the hbatng weate eects typical form of the variety The type of the form was collected by Harold Saint John (no. 16705) on top of a coral sand beach, altitude 2 meters, Peva Rurutu island, Austral Islands, on kiguat 28, 1934, and is depos- 1961 Moldenke, Notes on Vitex 91 ited in the herbarium of the University of California at Berkeley. The collector notes that the plant was forming a thicket and was in the form of a shrub 3 m, tall. It has been distributed as ve. trifolia var, trifoliolata “Schau. Citations: ASUTRAL ISLANDS: Rurutu: H. St. John 16705 (Ca— 805710-— LPR ARR es VITEX eg var, VARIEGATA we Additional literature: Neal, In Gard. Hawaii 61. 1948; Molden- ke, Re iocts 6: 198. 1958; Moldenke, : peas 226 & 479. 1959. "Collectors describe this as a shrub, to km. tall, the leaves ] 2 vari variegated or only slightly edged with white corollas blue, It has heen found blooming in September, Kaj reports it as common in the rainforest on the New Hebrides, where it occurs also in native Nal pete It is very peeked Ce raeateabe as a hedge on the Hawaiian Islands, where 1. ct s taxon is a form of V, trifolia var. subtrisecta, but in keeping with widespread horticultural practice to simplify nomenclature of strictly horticultural subjects, it is here regarded as a "variety", Additional citations: HAWAIIAN ISLANDS: Kauai: H. Ne Moldenke 21848 (Z). Maui: He N. Moldenke 21825 (Z). Oalm: H. N. Moldenke 21862 (2). CULTIVATED: Florida: CS 58162 (N); P. 0. Schallert 23077 (S). New Hebrides: K Kajewski 8) ( ie. = TEX TRIPINNATA (Lour.) M Additional literature: Mo ol denk Phytologia 6: 198--200. 1958; rerarre Résumé 175, 177, 179, >, on, 353, 355, 381, 385, his, & Collectors describe this as a medium-sized tree, with "sweep- ing" branches, yellowish blaze, and scented, white and yellow flowers, apr ine Eeetioge arch and April, and fruiting in August, scattered in open deciduous forests. Contrary to what is said in pam ig # 199 Ebi ine euler fos this supical form.af the labro Clemens Sy ~seyde 17 zal 17901, gow meseat “23 cited on page 200 are all actually var. clemensorum, the first-mentioned being the en pe ccllestion of that variety. tional vernacular name is "neklek —, woiens 323 in the Berlin herbert arium is a mixture with some material; Poilane 8565, in the same herbarium, re labeled mnyitex V. tripinnata has been collected at 600 meters altitude of 600 meters. Ga material has been misidentified and distributed as V. peduncularis Wall Addi ditional citatio : Annam: Colani 4040 Sy Poilané 2717 (B), 11261. (B), 3), 22283, (B, Ca--53908). Cambodia: 92 PHYTOLOGIA Vol. 8, no. 2 Jjaud 223, in part (B). Cochinchina: Poilane s.n, [Arboretum de Trang-bom) (B). Tonkin: Pételot 040 (Ca—5798). State undeter— mined: Eberhardt 105 (B); Pierre s.n. (B). THAILAND: Charoenmayu 386 (Royal Forest Dept. 5166] (Sm). PINNATA var. CLEMENSORUM Moldenke, var, nov. varietas : forma typica speciei calyce extus dense aureo- rece This variety ref eee from the typical form of the species in having the calyx densely golden-glandulose on the outer surface. The type of the variety was collected by Joseph and Mary Kna Clemens (no. 3394) in open ground along a river by a forestry lodge and native house at Tourane, Annam, Indochina, between June : and 13, 1927, and is deposited in the herbarium of the Univer- sity of California at Berkeley. Previous to this writing I had not distinguished between the very glandulose and the glabrous forms of this species. C of low flowers in June. Citations: INDOCHINA: Annam: Clemens & Clemens 339) (Ca—339h89 —type, N--isotype, Uh emcees ype); Poilane 17 (N), 17901 (N). Cambodia: Bejaud 223, in part (N) VITEX UMBROSA Sw. Additional literature: Moldenke, Phytologia 6: 201-203. 1958; Moldenke, Résumé 56, 321, & 479. 1959. Additional citations: JAMAICA: We Harris 11975 (Vi). VITEX URCEOLATA C. B. Clar Additional oh gh sotdinwe » Phytologia 6: 205-206. 1958; Moldenke, Résumé 165, 166, sh 172, 179, 181, 189, 191, 19h, 959.6 320, 38h, 385, 583, & 479% Rec ent c ollector describe this tree as having a trunk diameter of 12 inches, gray bark, cut whitish, white flowers, and globose fruit which is green when immature and black when mature, They report the tree to be common in lowland evergreen forests and at the edge of forests, at altitudes of 100 meters, flowering in June, and fruiting in September, Specimens have been misidentifi- ed in herbaria and distributed as NV. glabrata R. Br. Additional citations: ANDAMAN ISLANDS: South Andaman: Prain's collector 29 (N—-photo). INDOCHINA: Tonkin: Bon s.n. (Ca—53985) THAILAND: Thaworn 152 (Herb. Roy. Forest Dept. 11552] (Z). SUMAT- RA: Yates 1609 (Ci (Ca--26 39,3). VITEX VANSTEENISI Woldenke literature: Moldenke, Phytologia 6: 206-207. 1958; Noidanks, Résumé 189 & 79. 1959. tional citations: SUMATRA: Van Steenis 900 (N--isotype)- 1961 Moldenke, Notes on Vitex 93 VELUTINA (Koord. & Val.) Koord. dditional literature: Moldenke Phytologia 6: 209-210. 19583 Moldenke, Résumé 185, 189, 191, 38h, & 79. 1959. eats nal 7036 (Ca—265985—— otype 2703 VERMOESENI DeWild. Additional literature: Moldenke, Phytologia 6: 211--213. 1958; Moldenke, Résumé 143 & 79, 1959, Additional citations: CONGO: Vermoesen 17h2 (Ca—6192—cotype, Z——cotype), 1933 (Ca—6190—cotype) . VITEX VESTITA Wall, Additional literature: Moldenke, Phytologia 6: 213216, eth § ropeenkes Résumé 165, 166, 171, 177, 179, 181, 187, 189, 191-19 198, 226, 383, 385, 386, 390, 391, & h:79% 1959. Recent collectors describe this plant as a bush or shrub 2,5— t 4m. tall, wi k-green leaves that are dull to slightly glos- Sy above, paler and dull beneath with b sh-green % green when immature (June). Sinclair f it a- long wooded paths on steep hillsides in Perak. It is found forests on and is said common in deciduous forests in Thailand, growing at altitudes of 25 to 3000 meters, ing also in February, March, June, July, and No ‘ Additi names are "khot ngu” and noi". Its roots are employed in Thailand against snakebites, It has gant misidentified by Dop as V. sumatrana var, urceolata King & amble, ——— age Srna Additional citations: INDOCHINA: Tonkin: Bon 6202 (B). MALAYA: Wnre: Mur & Kiah bin Hadji 772 (Ca--231469). Negri Sembilan: Nur 11508 (ca~-3)6227) . Pahang: Best 13892 (Ca—-237335). Perak: Pranit’ 39 [Royal Forest Dept. 10775] (z). SUMATRA: Bosea 7049 (Ca-—1662); Toroes 28h (Cal226h), L497 (Ca—530917, S); Yates TB (Ce--261070), 21,0 (B, Ca—300515)+ BRITISH NORTH BORNEO: M.S. Clemens 11272 (Ca—-214991), 51590 (Ca--541713); Clemens & Steens 26679 (Ca—Sh1269), 29768 (B Ca—5h1962, Ur). | VITEX VESTITA £, GLABRESC Moldenk “aga Soma literature: Moldeaice, Phytologia 6: 216. 1958; Mol- : 189 & 479, 1959, ne ee) onal citations: SUMATRA: Toroes 1698 (Ca--530521— 94 PHYTOLOGIA Vol. 8, no. 2 VITEX seconde is WINKLERI enke Additi terature: Moldenke, Phytologia 6: 217. 1958; Mol- denke, aaat ibs, 192, eae & sagt 1959. This goonies has been in fruit in October. Herbarium acne has be SF Moteibated ca V. vestita Wall. dditional ssiaieia BRITISH NORTH BORNEO: M. S. Clemens 9813 (ca-=21,990). VITEX VOIKENSII Gtirke Additional literature: Moldenk ogia 6: 218--219. 1958; Moldenke, Résumé 13, ede Ws 383, 3 neh, 2 “79. 1959. Schlieben describes tree oar © - tall, with hard wood, and the Py cote g = Basi ty growing a + 1200 m. ’ alti- tude. Additional citations: TANGANYIKA: Scheffler 199 [A. Peter 51814] (B); Schlieben 3201 (B), 5910 (B). VITEX WELWITSCHII Gttrke Additional literature: Moldenke, Phytologia 6: 223. 19583 Mol- denke, Résumé 139, 143, 145, 148, 9s 151, 385, 389, & 79. 1959- pris ee this plant in + in October. onal citations: CONGO: Ss ps 2931 (Ca—55590)3 Louis 2163 (ni), 3 3513 (B). VITEX WILMSII var. REFLEXA (H. H. W. Pearson) Pieper Additional L synonyay Vitex wilmsii var vars reflexa He He W. Pear- son, ddi terature: Moldenke, Phytologia 6: 225--227. 19585 Moldenke, “ateun nb 152, 15h, 388, & ‘bp 96 Recent c to's buh 8 feet tall, grow ing in pede se Sees ai evihreeade ys locally frequent on rocky slopes in Transvaal, flowering in re aig Additional citations: SWAZILAND: Rodin UShh (Ca-—802799) » UNION rl ce AFRICA: Transvaal: E. E, Galpin 262m 282m (Ew); Meeuse 10259 VITEX YUNNANENSIS W tional es ew Woldenke, Phytologia 6: 228--229. 19583 Moldenke, Résum& 171 & 79. 1959. Recent collectors describe Sy as a small tree, gar violet 326752), 23725 (Ca--516551), 21,298 (Ca--S1BEE0), "Ytmnan: Maire 3647 (Ca—386798); Forrest 11349 (Ca—231330); J. F. CG. Rock 5063 (Ca—327150), 10564 (Ca--327553) . bag ans oe Gttrke “8 wo} literature: Moldenke, Phytologia 6: 230. 1958; Mo+ Additional citations: CAMEROONS: Zenker 139 (B), 5315 (Ca 1961 Moldenke, Notes on Vitex 95 62031). VITEX ZEYHERI Sond, Additional literature: Moldenke, Phytologia 6: 230—231. 1958 ; Moldenke, Résumé 152, 15h, & 479. 1959. Recent collectors describe this as a tree, 6m, tall, with cream-colored flowers, cammon in Transvaal at altitudes up to dditional citations: UNION OF SOUTH AFRICA: Transvaal: Sn. [6/4/1935] (S)s Schlieben 7733 (B); Wasserfall & von Niekerk h (S). MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA, I Harold N. Moldenke This is the twenty-eighth in my series of works of monographic nature on the genera of Verbenaceae, Avicenniaceae, Stilbaceae, and Symphoremaceae. Previous genera so treated were Acantholippia Griseb., Aegiphila Jacq., Amasonia L. f., Avicennia L., Baillonia Bocq., Bouchea Cham., Casselia Nees & Mart., Castelia Cav., Chas- canum E. Mey., Citharexylum B. Juss., Cornutia Plum., Dios Miers, Dipyrena Hook., Hierobotana Briq., Parodianthus Troncoso, Petitia Jacq., Petrea Houst., Priva Adans., Pseudocarpidium Millsp., Recordia Moldenke, Rehdera Moldenke, Rhaphithamnus Miers, Stylodon Raf., Svensonia Moldenke, Tectona L. f., Vitex Tourn., and the New World and cultivated members of Callicarpa L. Full explanation of the abbreviations employed herein for the names of the 260 herbaria whose material was examined, in whole or in part, in the preparation of these works will be fo Phytologia 5: 15159 (1955), 6: 2h2 (1958), 7: 91--92 (1959), 7: 123—12h d 7: 343 (1961) with the addition of Ko = Botani ln, Germany. th 5k (1960), an Sches Institut der Universitet, Ktln, VERBENA L., Sp. Pl., ed. 1, 18. 17533; Gen. Pl., ed. 5, 12. 1754 (not Verbena Dill., 1763, nor Pluk., 1763]. Synonymy : Obletia Rozier, Introd. Obs. Phys. Hist. Nat. 1: 367. 1773, Verbena Tourn. ex Adans., Fam, Pl, 2: 12 & 198. 1763. ratya Neck., Elem, Bot. 1: 296. 1790. Aubletia Neck., Elem. Bot. 1: 356—357. 1790. Glandularia J. F. Gmels in L., Syst. Nat., ed. 13, 2 (2): 920, 1791. Billardiera Moench, Meth. Pl. %9. 179k. Suttleworthia Weisn., Pl, Vasc. Gen. 1: 290 and 2: 198. 1839. = Bunge, Bull. Sci. Acad, St. Pétersb. 7: 278. 180. Schattleworthia Meisn. apud Schau. in A. DC., Prodr. ll: 535, in 96 PHYTOLOGIA Vol. 8, no, 2 syn. 1847. Shuttlworthia Walp. apud Schau. in A. DC., Prodr. ll: 553, in syn. 187. Aubletia Jacq. apud Wittstein, Etymolog.-bot. Handwtrterb. 85. Sipe Uerbena Sessé & Moc., Pl. Nou. Hisp. 1: 6. 1887. Helleranthus Small, Fl. Southeast. U. S., ed. 1, 1011 & 1337. 1903. Veebena Vent. ex Moldenke, Alph. List Cit. 1: 96, sphalm. 1946. Verebna Augusto, Fl. Rio Grande do Sul 217, sphalm. 19h6. Verbenia L. ex | L. ex Moldenke, Alph. List Invalid Names Suppl. 1: 28, in syn. 19h7. Vernena L. ex Moldenke, Alph. List Invalid Nemes sient. 1: 28, in syn. 19h7. Verbent L. apud Deam, Yuncker, & Fries- ner, Proc. Ind, Acad. Sci. 58: 95, | 95, sphalm. 19,8. Verben Perry & Fernald ex Moldenke, Alph. List Cit. 2: 58), sphalm. 198. Grandularia o5 Bekions Crovetto & Piccinini, Revist. Invest. Agric. h: 77, sphalm, 1951. Verbina L. ex Woldenke, Résumé 379, in syn. _ _Siandlons P Penfound, in herb. Verbinia Wahlgemt, in herb. erature: erature: Clus., Rar. Stirp. Hisp. Obser. Hist, 2: 372--373. 1576; C. Bauhin, Pinax Theatr. Bot., ed. 1, 269. 1623; en Parad. m tb. —389. 1732; L., Syst. Nat., ed. 1, ti as L., Hort. Cliff. ll. 17373 Clayt., Fl. Virg. 8. 17433 Ley Hort. Upsal. 8. 178; Haarte 1 *9 Pl. Hybr. 7-8. 1 13 Le, e e ® 2 & 630. 1753; bes Gen. e, ed. 5, i2?y rise in es Am o A ° 33 35 & {3, pl. 2. 1756; tz in e 6: 81. 17603 *9 Sys e Nat., ed. 10, 23 eh Py "1759; Lig oF PL., ed. 2, 29. 17623 o, Fam. Pl. "2s 12 & 198. 1763; o Vive 28 t. Veg., ed. — 62. 177k; Fl. Dan. h: pl. 628. 1775; “Medi, Act. A -Palat. 3 (Phys.): 19h, ay a 1775; Forsk., Fl. t ewarab, 17753 Aubl., Hist. P Ls oe i: 75; Hasselt ‘Biyacl, Teut. Ling. anes —— Fl. Brit. Indig. (a. 92). 17783 chri . 1779; Houttuyn ya = “2 "1779; Zorn, c oo Med, 1: pl. ee rt195 decay Select. ° Hist. Picta pl. 9. 1780; . 11 & 12. 1786; Walt., Fl. L Medic., Bot. Beobacht. 1783: 13. 270 Gilib., Fl. Lituan. 1: 92+ Nov Carol. 164. 1788; Jacq., "coll. 23 See 1788; Plenck, Ic. Pl- 1961 Moldenke, Monograph of Verbena 97 Med. 1: pl. 17. 1788; Gaertn., Fruct. & Sem, 1: 315, pl. 66. 178 Jacq., Icon. Pl. Rar. 2: pl. 207 & 208. 1789; A. L. Juss., Gen, Pl. 109. 1789; Hoppe, Ect. Pl. Ratisb. pl. 332. 1789; P. Brome, Civil & Nat. Hist. Jamaic., ed. 2, pl. 3. 1789; Neck., Elem. Bot. 1: 295-296 & 356-357. 1790; J. F. Gmel. in L., Syst. Nat., ed. Hoffm., Deutsch, Fl. pl. 2. 17913 Le. Ce Rich., Act. Soc. Hist, Nat. Paris 1: 105. 17923; c. K. Spreng., Entd. Geheimn. Natur pl, Po. 1 R > 5 6 Spe T3 pl. 435. 1806 Moriart; Col, Plates e Pl. pl. ho. 1806 Poir,. Mth. Bot. 8: 516—550 & 813—8h). 08; Roques, Pl. Usuel. In- + & Exot. 2: pl. 59, 1808; § » Bot o, Od. 2, pl. 3. 1808; Raf pos. N. Yo, sere 2, 58 352-8 360 26085. e, ed. 2; 2 pl. e 3 -s “2 port. Berol. 632633, 1809; Jaume St.-Hil., Pl. France {ple 9]. » Ann. Mus, Hist, Nat. Paris 15: 260, pl. (2), fig. 1-5 & 7, 1810; R. Br., Prods. Fle Nowy. Holl., ed. 1, alk. 18105 A. & F. Plée, Herbor.’ Parte (pl. 27]. 1811; Ait., Bin; KOMes ed 2, hz 362, 16125 Hill, Family Herbal pl.’ 9. cn Vigneux, Fl. Pitt, Envir. a wie 9 + Inst. France 12: 157 & 162, pl. 1. 18123 Stokes, Bot. Mat. . 1812 Brit. Fl. Tai7?', 1816-18295 curtis, Fl. Lond., od. 2) 1 (1): pl. li. mw Hopkirk, Fen pl. 7, fig. 5 & 6, 1827; J. Sm. in Rees, > %, no. 128, 1817; H.B-K., Nov. Gen. & Sp. Pl. 2: pl. 98 PHYTOLOGIA Vol. 8, no. 2 39-37. 1817; Hayne, Getreue e 5: pl. meron, - 63 pl. rg Fa 1818; Curtis, “Bot. re “is pl. 1576. 1 8185" Bet sg Fay Gen. & Sp. Pl ®» quart., 2: 272—< 8; edwards, Bot. Reg. hg pl. 29h. 18185 277, pl. ee 181 Sims in reng. o, Syst. Veg., ede iE, 20 Th? 752 & 763, coats ant Fl. Flum, 16-18, 1825; Reich- enb » Hort. Bot. pl. 6h. 1825; Lehm., Ind. Sem. Hort. Hamb. 17. . ° 1 43 1827; Reichenb., agcnna Bot. Exot. 1: xiv & 8, pl. 64. 1827; Sweet, Brit. ¢ 3: pl. 202. 1827; Eadie & Sm., Fl. Graec. 6: sp. 553 & ssi. 182 1827; Spreng, Syst. Vi Bey Cur. Post h (2): 30-231. 1827; R. Grah., Edinb. Now Philos, Journ. (Oct.-Dec.]: » ed. 27 Reg. 13: ms te 1102 ° (1827) and. 14: = aah. 38 gl anew Consp. Reg. Veg. 1: 117. 1828; Linnaea 3: —— ho. 1828; Curtis, . Mag. 56: pl. 2910. 1829; Desf., Cat. Pl. Hort. Reg. Paris, a 3, 393. 1829; Loud., Encycl. Pl. 520--522. 18293 Sweet, Brit. - Gard., ser. 1, 3: pl. 295 (1829) and [ser. 2, 1] ° 29 “ . : pl. 151) & 1550. Froriep, Notiz. Gebiet Wi Heilk., ser. 1, 23: 292. 1829; C. McIntosh, Flora & Pomona pl. 11. 18293 Hook., Bot. Misc. 1: 159-172, 18 1829; Maund, Bot. Gard. 3: Pl. | 52 & 70. 1829--1830; M. Roscoe, Fl. Illustr. Seasons pl. xu. 1831; Sweet, Hort. brit., ° 418, 18 Epi Carol . 4 830; Moris, Gewtichsk 2 e Engl. Fl. Gard. pl. 262. 1833; Hook. & Ara., Bot. Beech. Voy- 156. 1833; Baxter, Brit. Bot., ed. 2, 1: pl. 26. 1834; Dietr., Fl. e Bot. 2 23 ies 38. 18 3h; este . Gard. 6: (ter. 2, 12! pl. 22 221. 183; Maund » Bot. i ee che 106, & 112, 183h--1835; Lindl., Bot. Reg. 21: pl. rey “a 1766. 18353 1961 Moldenke, Monograph of Verbena 99 18 ; Linnaea 10: pretend 115. 1836; Sweet, Brit. ard. 7 i, 2, kj: pl. 318 & 347. 1836; Hook, & Arn sch. Voy. 205 & 268. 1836; orists Mag. 1: 5. 1836; Hook., Comp. Bot. Mag. 1: 176. 1836; Hook. in Curtis, Bot. Mag. = pl. ; h, Flora 19: 390. 1836; Zenk Tausch, Fl er, t Pl. 321. 1836-1855; Edwards, Bot. Reg. 23: pl. i928. 1837; Floricult. Cab. 5: 2h. 1837; Bojer, Hort. Maurit. 25). 18375 E. Mey., Comm. Pl. Afr. Austr. 1: 27). 1837; Nutt., Trans. Am. Phil. Soc., new ser., 5: 18. 1837; Sweet, Brit. Fl. Gard. 7 [ser. 2, lJ]: pl. 3 eae and pl. 363. 1838; Paxt., Mag. Bot. : 5 (1838) and 5: 23. Maund Hoo. e 36 Marnoc Flo i ms ° 1838; Jun pons } Bop 216 & 33. 518368 Endl., Pl. 1: 633- 63h.) 1885 1 Floricult. Cab. 6: 47 (1838) and 7: bo, "18393 Meisn., Fi. Vasc, Gen. 1: 290 & 299 and : er re 96 1839; G. Don in Loud., Hort. Brit. Suppl. 2: 680. 1839; Royle, Illustr. Bot. Him- wh 18393 Maund, Botanist 3: pl. 129. 1839; Benth., Pl. H ook. in Curtis Bot. Mag. 65: pl. 328 & 369h. 1839; Maund, Bot. Gard. 7: pl. 11: pl. 197. 1839—-1851 ; e Sci. Acad. St. Pétersb. 7: 278. 180; H. E. Richter, Cod, ° Linn, 868. 1840; Eaton &W Wright, N. Am. Bot., ed. 8, 6 ews 180; Fisch. & Mey., Mém. ate St. Pétersb., et yh (2): 153 ~-156. 18h0; Marnock, Floricult. Mag. 5: 87--88, pl. aan 1840; Paxt., Mag. Bot. a 3. 180; Harrison, Floricult. ae ‘Bs i & _ 180; R. Grah., Edinb. New Phil. Journ. 29: "1th--175. ook, a Bot. E Univ. 2: 135. 18h1; Nees, Gen. Pl. Gamop. 2: pl. 50. 18l1; Paxt., Mag. Bot. 1: 173—17h & 278. 18h1; Steud., Nom, Bot., ed. 2, 2: 2: pl. 91. 1844; Bohn, Florist's Journ. es “W—h2, pl. "eh, fig. 1—3. 18h4; Benth., Bot. Voy. guigh. 1 153. 18h; Hassk., 2e Cat. tens. 131. 18h; Im., Am. Journ. Sci. 6: oer 4, 5, 7-9, 11-36, 13, & 7—=50- , Hist. Nat. Vég. Phan, Atlas pl, 8), ; Duchesne, Pl. Util. pl. 56. 1816; Sieb. & Zucc., Abh. Muench. (3): 152. 1846; Benth., “st oy ny as 152—~153. 18h63; Gard. Chron. 18h6: 11h. Je We —_ ha s2 Pp. ho. 18,63 Schau. in A. as $3566. 18475 Walp., Repert. 6: 686--687. 18473 W act ry wes Pn — 1873 Scheele, Linna t 7556 Annal , » P 189; a ‘shes she. inusidlsy ¢ Ce “nee, Hist, Fis. Chil. Bot. S: 7—2h. Gard) ay. 169. 69. 1850--18e4, Lem., Jard. Fl. 1: Misc. 85. 1851; 100 PETT?0L06:2a Vol. 8, no. 2 Sweet, Ornament, Fl Pp. 1855; "Fl. des Serres, 8 str. 2 1: pl. 1129, 1856; ae Schweitz, Pfl e~ldiotikon,. 88, any & se 18563; A Gray, Man., 9 ae 1856; Wood, Class-Book, "ed. hi, 412, 18%6, Schnitzlein, "Teonogr. 2: pl. 137. 1856; rote ss Fl. Ind. Bat. 2: 908. 1856; Reichenb., Icon. Fl. Germ. 18: pl. 1292, = ar Organog. Comp. Fl. pl. 115. 1857; R. A. eet » Linnaea 21. 1857; Wedd., Chlor. Castelnau u Exped. Bot. J 2: ar peor r. Muell. 1: 60. 1858; Ettingsh., K.-k. Geol. Reichsanstalt Jahrb, 8: 79 1&15. amatty, Brit. Wild Fls. fig. 981. 1858--1860; Torr, in eines Rep. U. S. & Mex. Bound. Surv. 2: 128 18595 A. W. Chapm., heast. U. S., ed. 1, pr. 1, 306—307. 3 Wedd. erg, fig. 282, 1861; Walp., Annal. 5: 706. 1860; R. A. Phil., Fl. Atate LO, pl. 5. 1860; R. A. Phil., Viaje Des. Atac. 20, & 21). 1860; Bocq., Adansonia 2 : 105, pl. ne 1862; T. Moore, Field Bot. Hogg & Johnson, Wild Fl. Gr. Brit. 1: pl. 59. 1863; Gaud., Soc. Helvet. Sci. Nat. Nouv. Mém. 20 (3) [Contrib. Fl. Foss. Hal. 6]: Us Bij Brit. Fl. fig. 81). 1865; Schubert, Naturg. Pflanzenreich, ed. 2 Pl. 2, fig. 7. 1865; — Fleurs Pl. Terre, ed. 1, 936--9l2. 18653 Miq., Ann. Mus, Bot. Lugd.-Bat. 2: 97, 1865; Syme, Engl. Bot. 6: pl. 1018. 18665, se heer & Naud., Man. Amat, Jard. 2:3 492. 18663 H. — — Am. Acad. 7: 19). a : — & Gibelli, Comp, tal. pl. 49. 1867--1886; + Sy Acad. Sci. Philed. iser 21. 1868; Le Maout &. seis aité Ste: - 1868; R. A. Pui Anal. Univ. Chil. 35 191-192. j Dupuis & Réveil in Réveil, , Gerard, & Hérincq, Régne bi Flor, Med. 3: pe = ae i 1870; Hérineq & Gerard in — Régne Vég. h: pl. 16 & 18, 1870; Belg. Hort ae : Alt. 32—33]. 1870; Benth. & Muell, » Fl. Austral. 5: 35. 1870; Van Hall, Kruidtuin Rijks Hoog. Burgersch. — g- 1871; Ae W. Chapm., Fl. Southeast, U. a _ on gwen 18725 Ulrich, Internat. Worterb. Pfl.’ * is—-350. 872; Rtmpler in Viln., Tllustr, Blumeng., ed. L Ch Sem 9 1873, Pratt, ow. Pl. Grasses Sedges & Ferns, ed, 3, ks vidi & 208-211, P i. t. 88, 1874; Cusin, Herb, Fl. Frang. 18 [Verb.]: pl. 1. 1875; Ulrich, ternat. Worterb. Pfl., ed. 2, 249-260. 1875; Lesacher & Mares 1961 Moldenke, Monograph of Verbena 101 chal, Nouv. Bot. Méd. ls {pl. 8). 1876; Benth, in Benth, & Hook. f., Gen. Pl. 2 (2): 1146, 1876; Vatke, Ind, Sem. Hort. Berol. 1876: App. 1. 1876; J. G. Baker, Fl. Maurit. & Seych. 252. 1877; ed. 1, 1: 26. 1877; A. Gray, Syn. Fl. Ne Am. te Prits. & Jessen, Deutsch. Volksnamen Pfl. 1 ier, Verh. Natur, Ver + Rheinl, & Westfalens 39 (1): pl. 1. 1882; Wi » Ill. Deutsch, Fl., ed. 2, 617. 18823 Spe . Clent. Argent. 15: 112--116. 1883; W. Robinson, Engl. Fl. Gard. Pl. 262, 1883; S. Wats., Proc. Am. Acad. Sci, 18: 13%. 1883; A. Gray, Proc. An, Acad. Sci. 19: 95, 1863; W. Mill., Dict. Eng. Names Pl, & 261, 188k; Lindman, Svensk, Vetens. Akad. Handl. 21 (lu): 9 & 16—~17, pl. 1. 1884; Ball, Journ, Linn. Soc. Lond. Bot. 21: 230-—231.°188h; Macoun, Cat. Cahad. Pl, 2: 379. 188h; A, Le Chapm., Fis Southeast, Ue S., ed. 2, pre l, 306—~307. 2 in Hook, f "¥1 3 g tag 1885 Denkschr, Akad 7 = ay - Brit. Ind. k: e 3 Stapf S ° gf Ween 50: 3h. 1885; Baill., Ic. Fl. Frang, h: pl, 376. 1865--189h3 Sterne, Herbst- & Winterbl. pl. 16, fig. b. 1886; Macoun, Cat. As e Chapm., Pa Southeast. U. S., ed. 7? he af 307. 1887; Sessé & Moc., Pl. Nou. Hisp., ed. 1, * 6. 1887; Moloney, Sketch Forest. W, Afr. 523-52). 18873; Reg- Sl, Gartenfl, 36: 668, 1887; Nicholson, Illustr. Dict, Gard. }: 145. 1887; Samijn, Decken de Bots Kruidwordenb. 147. 1888; ages Pittonia 1: 156, 1888; Lillo, Fl. Tucumén 9. 18 ; inane’ Album de Clichés 707, fig. 25645, 25655, 25660, & 25661. + eg Rattan, Popular Calif ‘ ; +» Fl. Haw. Is. 30, 1888; Dujard-Beaumetz & Egasse, Pl. Méd. c Sessé & Moc., Pl. Nou. His 2, 1s ° ° ° pan., ed. 1 [La Naturaleza, ser. 18bg. O¢ 18895 Lorents & Niederlein, Exped. Rio Neer> 26-266 Southern, Fl» Deutschl. kt pl. 531. 18895 A. W. Chapm., Fl. - U.S., ed. 2, pr. 3, 306—307, 636, 692, & 698. 1889; ray Pa Sin. 2: 252. 1890; Wats. & age hs A ss od, 02. 1890 Paolucci, Fl. Marchig. ° ° 1890; Re A, Phil, anal. Mus’ Nac. Chil. Bot. 1891: 58—59. 1891s 102 PHYTOLOGIA Vol. 8, no. 2 Bulg. kl. 1891; Ball, Journ. - Soc. Lond. 273 IT obo; Baill., Hist. Pl. 11: 78—79. 1891; A. S. Hitche Trans. Louis Acad. Sci. Ss “12, 18913 Re Se Mill., Vicks Tl- lustr. Month. Mag. 1, (4): 117. ee, Webber, Trans. Acad. St. Louis 6: 0. 18923; Bergen, Journ. Am. Folk-Lore Apr.-June 1892, p. 102. 18923 Baill., Dict. Bot. hz 256. 1892; A. We Chapm., Fi. s east. U. Se, ed. 2, pre h, 306—307. 1892; Coult., Contrib. U. S. Nat. Herb. 2: 327--328. 1892; Morong, Britton, & Vail, pe: ms 3. 1892; Lub 2: 370. 1892; Small & Heller, aig Bot. soo zt ies 1892; Sess6 & Mi rr Pl. Nou Hispan., e 6. 1893; J , Erythrea 1: 12, 1893; G. Beck, Fl. Nied.- Oesterr, 2 (12): “OTS « Fed Masclef, Atlas Pl. France 70. 1893; Stewart, Bull. Iowa Agric. Sta. 21: 814. 1893; Hook. & Jacks., Ind, Kew. 1 (2): mgt & 1032. 1893; Dana, How to Know Wild _Fis., ed. 1, 90, 252, & 297. 1893; Britton, Ann. * Y. Acad. Sei. 73 192. 1893; Heckel: Prov. List Indig. Fl. Pl. amaic. 30. 1893; Schimpfky, Uns. Heilpfl. 2: pl. 42. 1893-189); oad, Dict. Hort. 1189, 1893-1899; Dana, How to Know Wild Fls., ed. ¥ pl. acks., France 527. 189); Britton, Mem. Bot. Club 5: 276. 189; Coult., Contrib. U. o Nat. fac 2: 2: 687. 189); a et Surv. Nebr. 3: 18. 189); Vilm., Fleur. Pleine Terre, ed. 3, 195 & 1198, 1894; Bretm, Mery. Nat. Monde des Pl. 2: 36. 25sL--1856 arcke, Ill. Fl. Deutschl. 489. ve barat Blumeng - 3 [Sieb. & Voss], 1: 825. 18953 Karst 1.Deutschl. 24 eh. 18953 Rydb., Contrib. U. S. Nat. Herb. 3: hearse 1895; Dana, How to Know Wild Fls., rev. ed., 122, 31h, 365, te: : 372. 18953; Robin- s 1y2 spats & ageers Bull. Kans. Agric. Sta. 50: pl. 4 fig. 91 & 92 8953 Re Ae , Anal, Univ. Chil. 90: [Pl. «] 607— as, pl. 1. 1895)" He He Rusby, Mem Torrey Bot. Club hs 2hi--2h5 8953 Heller, Contrib. Herb. Franklin & Marshall Coll. 1: ot pl. 1895; S. Moore S. Bo ad, ser. he : ° 18955 L. H. Bailey in A. Gray, Field Forest ot 3 & 518. 1895; nes ites Pl. Jard. pl. 227. reel Bull. Torrey Bot. Club 23: 10k. tree Rév, Hort. 1896: 188. "48965 Re Ae Phil., + Univ. “onal : 609 & 616, 1896; ny Bull : . Sta, st "28, pl. 12 & 13, fig. 137-10. 18965 Si tlas Poche Pl. - & Bois, sér. 2, 1896; Heller, Minn, Bot. St. 1: 882. 1897; A. pm : o U. S., od. 3 369. 1897; Step, Fav. Fl. Gard. & 563. aig ateaae Mittheil. Bot. 973 Phage Le 376; oad pull. Soc. ae oan "223 77, 189 97; Hitehe . & Norton, Agric. S ta. 66: pl. 12, fig. 137-10 (1897) and Paige pl. r pert i: 47 & 48. enh, 3: pl. 22. 1897; Speg., Rev. = tee oa Plata 3: 559— Thttrimg. 1961 Moldenke, Monograph of Verbena 103 1898; Wooton, Bull. Torrey Bot. Club 25: 262, 1898; Speg., Comm. - Buenos Aires 1: 137. 1898; Haage & Schmidt, Cat. Général 1898: 93-9. 1898; Schimp., Pflanzen-geogr. 7))2. 1898; Kuntze, Rev. Gen, Pl. 3 (25: 255——258. 1898; Greene, Pittonia 3: 308 309. 1898; Lyst Volksnamen Genees. Utig. Ned. Pharmac. 1898; Small, by, Bull. Tor ub 27: 80. 19003 Farwell, Ann. Rep. Comm. Parks & Boul. Detroit li: 82. 1900; Lindman, Veget. Rio Grande do Sul 110. 19003; Salmon, Mem, Torrey Bot. Club 9: 169. 1900; Journ. Agric. Soc. Eng ser. 3, 11 [61]: 738. 1900; Gartenfl. 49: 585. 9003; Blanchan, Nat. Gard. 40. 19003 Greene, Pittonia h: 152. 1900; Baker & Stapf in Thiselt.-Dyer, Fl. Trop. Afr. 5: 286—-287. 1900; Rev. Hort. Belg. 36: 76—-77, fig. 15. 1900; Bull. Soc. Tosc. Ortic. 26: %. Speg +, Contrib. Fl. Tandil 41. 1901; Eggert, Torreya 2: 123 Earle, Bull. Ala, Agr. Exp. Sta n ks 2y5—216. 1902; Cockerell, Am. Nat. 36: 809. 1902; Chod., Bull. Herb. Boiss., sér. 2, 2: 817--818. 1902; Chod., Plant. Hassler. 9: — 1902; Wittmack, Gartenfl. 51: 556—~557. 1902; Bull. Speg., Nov. Add. Fl. Patag. 2: 66-72. 1902; C. T. Green, Fl. Liv erpool Distr. fig. 487. 1902; Josef von Oesterreich & Marg. von Und Taxis, Atlas Heilpfl. pl. 181. 1903; Voss, Salomon's Wrterb. Deutsch. Pflanzennamen 37 & 251. 1903; Losch, Krtuterb. Pl. 61. 1903; Sturm, Fl. Deutschl., ed. 2, 11: pl. 22. 1903; Pan- mel, Bull. Iowa Agric. Sta. 70: 513. 1903; Dusén, Arch. Mus. Nac. Rio Jan. 13: 36. 1903; Selby & Hicks, Bull. Ohio Agric. Sta. 12: Pl. k, 1903; Heinemann, Rev, Hort. Belg. 29: 5li-—-55. 1903; Small, pie Southeast, U, S., ed. 1, 10091011 & 1337. 1903; Greene, i 2? 135-137, 1903; Briq., Ann. Conserv. & Jard. Bot. i — a) t= ’ Ind. Kew, Suppl. : 191. 190; Michx., cg o, ed. T 67. 1 “pata 3: 436. 1905; Rendle, Journ. Bot. 3: 33. 1905; Mac- in W. B. Scott, Rep. Patag. 8 (2): 1906; Kirby, Brit. Fl. pl fer 1906 Merino, Fl, Galicia 2: 223° 1906; Durand & Jacks., I > > 1: 451. 1906; sleby & Hicks, Bull. Ohio Agric. Sta 10h, PHYTOLOGIA Vol. 8, no. 2 175; pl. he 1906; Ue Se a ae Agr. aeep res 260: — Clark & Fletcher, Farm Weeds Canada, ed pl. 5h 31. 1906 F. M. Bailey, Weeds & Poison. Pl. ped; "139 & “o F ares De Te » Bot. Gaz. 43: 57. gee Lazaro, Comp - Espafi. 2 alte 1907} Heukels, Woordenb. erl. Volksnamen i 1907; Beuten- Conserv. & Jard. Bot. Genév. 10: 100—10);. 19073 F. Ee Lloyd, Car- e Kew. Su « Wash. Publ. 82: pl. 3. in ° . 3: 187 Urb., « Antil. 5: 48h. 1908; M. E. Jones, Con- trib. West. Bot. 12: 78. 1908; \ - N.Y. : 1 inson & Fern, i G ew Man. Bot., ed. 7, 688, A. 689, & 92h, fig. 878-880. 1908; Hayek, Denkschr. Kaiser. Akad. Wissen. Math.-nat. 79 (1): a a M. A. Day, Check List 128. ek in - U 35: ree 1909; Clark & Fletcher, A: Weeds Canada, ed. 2, pl. 75, fig. 61. 1909; Beille, Bot. Pharm. 2: 1075 & 107%. 1909; Black, Nat. Fl. Se Austral, 127. 1909; Hassler, Flor. Pilc. 10l— 102. 1909; sg 8 al Fl. Nederl. 3: 270. 1909; Vilm. Andr. & Cie, Fleurs Pleine Terre, ed. 5, 1126. 1909; Hicken, Chlor. Plat. Ar- gent. 195197. 1910; P. atieeeiete Handb. Pl., new ed., 75—l76. 1910; Backer, Ann. Jard, Bot. Buitenz - Suppl. 3 (1): 419. 1910; Oesterr. Gart .~Zeit. es ne 1910; Reiche, Fl. Chile 5: 296, 300, 302, 30h, & 463. ; Dinand, Taschenb. Heilpfl. pl. fig. i, i910; abelian (ator Fl. Bolivia 1: 165. 1910; Fed- tse er., Fl. Eur, Russia 7 3 H. H. W. Pearson Thiselt.-Dyer, Fl. Cap. 5: 181 & 207—-210. 1910; Russell, » & — in Webster's Imperial . 10-1911; Beal, Mich. eeds [Bull. Mich. Agric. Exp. Sta. 2673] goa eg aiegge Pamel, Seeds bien & Gard. 218. 1911; Daniels, Fl. ~-205. Soc. F Sp. Nov. 9: 362. 19113; Correvon, Fl. Champs & Bois pl. 83. 19115 Lindman, Vi och Vara Blom pl. {2. i911 19133 Creevey, Harper's Guide Wild Fls. 331-332. ee Somoku Dausetsu, ed. Makino [Ic- on. Pl. Nippon.] oe pl. 0. 1912; Thell., Fl. Advent. Montpel. 428. 1912; Beal, Ann, Rep. Mich. Agric. Sta. 25: 385—386. 1 19135 Woot. & Standl., Contrib. U. S. Nat. Herb. = 166. seks 4 Pammel, Iowa Geol. Surv Surv. Bull. ls 263, 265-267, 468, & 46% 1913; Britton & Br., Illustr, Fl., ed. a 32 94-97, 599, 638, Nat. Buenos Aires 1913; Teangee. ra ei Wissen, Wien 89: 638. 1913; Klein, Unsere Unkrauter pl. h6. 1 1913+ PHYTOLOGIA is financed entirely by its contributors, each one paying — 23 in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the apheie = upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, aie his-233 part of the expenses and sharing in the profits, if any accrue. ! will be published in the next issue, so that the size of numbers may vai tly. A volume will contain about 32 signatures, 512 pages, or a smaller number of pages with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. extra charge is made for line drawings, such as are ordinarily reproduced it zinc, or for diagrams, tables, or charts, provided they conform to limitations of size and proportion. An extra charge will be made for h tones, depending on their size, as fixed by the engraver, with a minimum about $2.25. bopaphion sketches, and critical reviews ature will be considered for publication. Floristic lists, pare notes pe ar amateur or so-called popular type, and polemics will not be published. Ad on the suitability of manuscripts will be solicited, if necessary, from fied botanists. the present cost of printing, the basic rate for a page or to change without notice, since it depends entirely on the prices pre in the Peocing industry. Reprints will be furnished at — fraction of che: 3 of 250 copies is also furnished colle, to s conibusars _ Upon request, the editors will send detailed instructions concerning. preparation of manuscript or further information about the quiries may he addressed to the magazine or to either editor. Piy TOLOGIA Designed to expedite botanical publication & CS \S RPE CEN am ie Vol. 8 December, 1961 No. 3 CONTENTS _ BROWN, H. D. & DUPUY, H. J., An observation concerning the development of excised snapdragon buds in the presence Ze iis RRSP NMIOPYUIADIDH OFS ooo So i . 105° MOLDENKE, H. N., Materials toward a moroetieh of the genes oe Verbena 108 — fished & Ragold N. Moldenke and Alma Ey Moldenke 1S Clesbicsh Avenue <2 : Yonkers 5, New yok: U.S. AL = Vol. 8, No. 2, was issued October. "Price of this number, 75 cents; per Selon $5.7 75 in advance AN OBSERVATION CONCERNING THE DEVELOPMENT OF EXCISED SNAPDRAGON BUDS IN THE PRESENCE OF RESPIRA TORY INHIBITORS Brown, H. D.' and H. J. Dupuy During a study involving the isolation of young snapdragon buds from the stem and their maintenance, floating, upon solutions containing respiratory inhibitors (Brown, 1959) it ted that freq development would occur. More interesting, the activity of several inhibitors resulted in flowers which were characteristically misshapen 41 1 | J avuael itial to give what might be described as a gaping appearance. The fact of interference with the metabolism of the developing floral parts by known respiratory inhibitors is not in itself, of course, particularly surprising. Had the outcome of contact with the inhibitors been solely a slowing down of the pattern of development or its cessation, this would have accorded with expectation. But the regular peculiarity of the resulting malformation, indicating a specialized or localized activity by the inhibitors, led us to seek information about the developmental pattern of the flowers in this abnormal environment. MATERIALS AND METHODS Snapdr agons, Antirrhinum majus L,. cultivar. Candlelight, a tetra- Ploid, were grown to flowering under greenhouse conditions. The young buds, averaging about 15 mm. were carefully cut from the plant and, after the Sepals were coated with low melting-point paraffin, were mounted on lucite plates with the floral stalks protruding through holes drilled in the plates, The mounting was sealed with additional paraffin 80 that when the plates were floated, only the floral stalks would be in 1 PRES node ADDRESS: goTANY DEPARTMENT, SOUTHERN ILLINOIS UNIVERSITY, WARDSVitie, is 1LLINO 105 106 PHYTOLOGIA Vol. 8, no. 3 contact with test solutions for approximately 72 hours. Record of development was made by time-lapse photography. A Grass kymograph motion picture camera was actuated by a rotary switch adjusted to expose one frame every twenty minutes. The light of three one-hundred-watt lamps, diffused, provided illumination, Exposure was f22 for 1 second with Eastman pantomic-x film. Buds were floated on test solutions in small glass vessels. The vessels and buds, enclosed within the humid atmosphere, provided by a glass aquarium tank, were photographed through the wall of the tank, Solutions were 10 percent sucrose for controls, 10 percent sucrose and either .1 or .05 percent sodium azide for the experimental plants. In the earlier study, which had not been directed toward floral morphology, twelve inhibitors had been applied under conditions similar to those described here. RESULTS AND DISCUSSION It had been noted that at .1 percent concentration, sodium arsenate, sodium azide, and sodium fluoride caused, with varying intensity, @ characteristic deformity of developing excised snapdragon flowers. In the photographic observations .05 as well as .1 percent concentrations of sodium azide were found effective. Study of the photographs indicates that it is abnormality of the uppe? lip of the bilabiate corolla which is largely responsible for the character of the "gaping" flower. Figure 1 illustrates this type of flower with its upper lip folding backward so that essential parts are largely exposed where they would be concealed by erect stature of the upper lip in normal flowers. Azide inhibition, frequently linked to phosphate transfers (James, 1953), might be expected to affect petal development. The present observations are, of course, amenable to more than 4 single hypothesis. The characteristic malformation may result from ee 1961 Brown & Dupuy, Excised snapdragon buds 107 the indirect effect upon an auxin transport system or, more simply, relate to the distribution of inhibitor consequent upon the vascular anatomy of the flower, Figure 1, Normal and ‘gaping’ flowers (1, lucite plate; u, upper lip; p, palate). The authors find attractive also the possibility that a relationship of endogenous respiration to the efficacy of the inhibitor is involved. Siegelman, Chow, and Biale (1958) found, in rose, that the effectiveness of a metabolic agent, was inversely related to the respiratory rate of the tissues, Hence it seems not unlikely that, within the petal, localized differences in endogenous rates could, with the inhibitor present, effect the developmental pattern reported here. Loyola University, New Orleans LITERATURE CITED Brown, H. D. (1959) Effects of respiratory inhibitors upon nectar Secretion in Antirrhinum. Torrey Bull. 86: 290-295, James, W, O. (1953) The use of respiratory inhibitors, Ann. Rev. Pl. Physiol, 4: 59-90, Siegelman, H. W., Chow, C. T., and Biale, J, B. (1958) Respiration of developing rose petals. Pl. Physiol. 33: 403-409, MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. II Harold N. Moldenke L. Additional and emended a epee Dod., Stirp. Hist. Pemptad. re cae 1583; Clus., Rar. Plant. Hist. h: xlvi. 1601; R. Moris- on, Pl. Histor. Univ. Oxon, 3: goo Ba & sae sec. ll, pl. 7, & 8. 16993 1748; bes Spe Pl., ed. 1, 16--21, 630, sr i "879. i783 gre Beary Linn. Pfl zensyst. 5: pace 1779; Christm. & Panzer Pb Prin syst. eae 5: 119--130. 1779; Lour., Fle. ochinch. 1: 33. 1793; Lindl. in Edwards, Bot. Reg. 21: pl. 178 & 1766. 1835; Lorentz & Niederlein, Exped. sie Negro 2 (bot.): 263--266, a0 12. 1881; Ind. Kew. Suppl. 4: 245. 1913; Druce, Rep. Bot. Exch. Prain, Club Brit. Isles 3: 425. 1914; gph andy Fl. Moiss. 50, fig. 32, pl. 46. 1914; Perrin & Boulger, Brit. Flow. Pl. i: pl. "338 & Nac. 2 3: 158. 1915; Fyson, Fl. Nilgiri & Pulney coasikore 2: 213-- 1915; nerd, Bull, Vt. Agr. Exp. Sta. 187: 238. 1915; Woot. W: Fl e > 3 ’ Tr ’ Field Book West, Wild Fls. 435. 19153 Garden 79: 510. 1915; Bl ’ Wild Fls. [New Nature 63] 1916; H. A» Gleason, s Be Ann, Mo. Bot. Gard. 3: 292. 1916; Garden 80: 355. 1916; Herzog, ~— Rijksherb. Leiden 29: 1,3. 1916; Free in Barron, Pocket Gard. - 6: [Fls. Winter] 112 & 206. 1917; Shaw in Barron, Peake, @ Gard. - 1: (Gard, Fls. Spring] 205 & 230 and 2: (Gard. Fls. Summer 13 "& 251. 1917; L. H. Bailey, Stand. Cycl. Hort. 3: 3hh5--3uL7- 1917; C, L. Pollard in Webster's New “Internat. Dict. Eng. Lang. 2275 & 2278. 1917; Felt, Bull. N. Y. State Mus. 200: 162. 19175 Rydb., Fl. Rocky Mts., ed. 1, 740. 1917; Bonnier, Name this Flower pl. 45. 1917; Basu, Ind. Med. Pl. pl. 732. 1918; Lindman, Svensk rogamf. {6l;. 1918; N. L. Britton, Fl. Bermuda 310-311. 1918; F gamf, Tm House, Wild Fls. N. Y. 2 (Mem. N. Y.’ state Mus. 15]: 23h, 3%, & 6, 361, pl. 18 3b. 1918; H. J. Lam, V enac. 7, 9 1 .8 en : 8: 96--99, & 134, fig. 11--35. 19195 Lovell, Flower & Bee 255. 19193 Fiteh . Sm., Ill. Brit. Fl., rev. ed. h, iss. 2, fig. 622. 19193 Hauman, ak, Soc. Cientif. Argent. Buenos Aires bo: 22. 1920; Kanda, Bote Gaz. 69: Su—-71. 19203 Pellett, am. Honey Pl., ed. l, 19205 Molfino, Physis 5: 22 & 285. 1921, Murbeck, Acta Univ. Tund. ay eee - iT eae. _ieecieeeeees 1961 Moldenke, Monograph of Verbena 109 Univ, Arsskr.], new ser., 17 (1): 19. 1921; Fernald, Rhodora 23: 289. 1921; C. A. Weatherby, Rhodora 23: 172. 1921; Jaffuel & Pir- li: 203, 1922; Blanchan, Wild Fls. Worth Knowing 185, 186, & 270. 1922; Dixon & Fitch, Personal. Pl. 97, 150—-151, & index. 1923; Hicken, Sert., And, 65. 19233 K. V. O. Dahl - 1923: 10 & 12, 1923; Crawford, Addisonia 8: pl. 258. 1923; Larrafi., Escritos D. A. Larrafi. e Inst. Hist. Geog. Urug.] 2: 9+ 1923; Rep. Bot. Exch. Club Brit. Isles 1922: 617. 1923; Pell- et ney P 2, 353--354 & 391. 19233 Saunders, West. Fl. Guide 46, 169, & 285. 1923; Olmsted, Coville, & Kelsey, ; Pl. Names, ed, 1, 520--521. 192) Mak., Ill. Fl. Jap. [215]. 192k; r, Ohio F : Argent. Bot. A he 1926; A. W. Hill, Ind. Kew. Suppl. 6: 210. 1926; ow, 6s, Our Roman Legacy 83. 1926; Hegi, Illustr. Fl. Mittel-Eur. 5 (3): 2238 —22h3, pl. 222, fig. 3172. 1927; Bonnier, Fl. Compl. France ams, Guide Roy. Eot. ; Trin. 5. 1927; Sandw., Kew Bull, 1927: 184—187. 1927; Haage & Schmidt, Gen. Cat. 1927: 70. 1927; Gonzalez, Copp., & Lom- bardo, Fl. Urug. 1: 9 & 29. 1928; E. D. Schulz, Texas Wild Fls. 335338. 1928: E, J, Alexander, Addisonia 13: pl. 22. 1928; Roads, Ohio Journ. Sci. 28: 211° 1928; Freeman & Williams, Useful Pl. Trin, 165, 1928; Molfino, Flora Advent. 2: 20. 1 sonst, Contrib. Gray, Herb. 81: 96 (1928) and 85: 100-101, 929; H. F. Roberts, Pl. Hybrid. before Mendel 2. 1929; Frear, Fan. Isl, Trees 146, 1929; Standl., Field Mus. Publ. Bot. h: - 19295 Seymour, Host Ind. Fungi N. Am. 587—588. 1929; Bloss- “ dt, Art Forms in Nature pl. 39. 1929; A. W. Hill, Ind. Kew. — 7: 2h9, 1929; Stebbins, Rhodora 31: 85. 19293; Roberts & Rey ’ An. Pl. for Am, Gard. 23, 87, 96, & 130. 19293 Seckt, peg Univ. Nac, Cordoba 17: 87—90. 1930; Besant, Gard. Chron., ht bg-'700? 133+ 19303 Strausbaugh & Core, Proc. W. Va. Acad. sci. » 1930; Thompson & Morgan, Descript. Seed List 1930: 94-95. sv8) Nomb. Vulg. Pl. Silv. Chile, ed. 2. 1930; G. T. Ste- , — Flow. Pl, Middle Atl. & N. Eng. States pl. 129. Publ pece Mold. Southeast. Set 10. 1930; P. C. Standl., Field Prat, Bot. 3: 103, 1930; Guillaumin, Fleurs de Jardins 2: [Encyc * Mat. 16] pi. 43. 1930; Herter, Florula 105. 1930; Stapf, 110 PHYTOLOGIA Vol. 8, no. 3 Ind, Lond. 6: 03 & ae Oy es aon Schwencke, Zytol. Untersuch. Verbenac. 1--36. 1931 Elliott, Gard. Chron., ser. 3, 90: 378. Moldenke, Obs. Fl. Watchung, [ed. 1), 7 “ipa and ed eo an. Southeast. Fl. 1136-1139 & 1508. 1933; rack Nippon Shokubutsu Zufu (Jap. Bot. Illustr. Album] 1020. 933 Steyermark & Moore, Ann. Mo. Bot. Gard. 20: 805. 1933; Perry, Ann. Mo, Bot. Gard. 20: 239-—-362, pl. 13 & 15. 1933; ouard, Zoocéd. Pl. Amer, Sud 3). 1933; Schaffner, Ohio Journ. Sci. 33: 293 (1933) and 34: 172. 1934; Moldenke, Obs. Fl. Wat- c » ed. 3, 39. 1934; Dormon, Wild Fls. Louisiana pl. 19. 3h; spa ere in C gute Bot. Mag. ‘157: pl. tg 1935 Journ, Arnold Arb, 16: 133. 193k; House, Wild Fls. , 3h 336, & 361, pl. 183b. 193k; Hochr., Candollea $: 188. 193h3 P- Ce Stand. 293, & 320. 193); Patermann, Beitr. erbe 1--56. 19355 Molfino, Trab, Inst. Bot. & nah Sh: 112. 1935; L. He a Cat. Florists Handl. Verbenac., mss. 1935; Moldenke, [Syst. Bot.] 1 (1935) and Quiz 10 [Syst. Bote] 3. 19353 Yuenscher, Weeds N. Y. [Cornell Univ. Agr. Exp. Sta. Bull. 635:] 10, 16, & 387. 1935; s he _— er, Ann, Vo. Bot. Gard. 22: 629. 1935; Journ. oy. Hort. S nd. 61: L01. 1935; Thompson & Morgan, Des script. Seed List 1538; Bt. eg Moldenke, List Pl. Obs. Pines 7. 19355 orsmo, Ugressfrt pl. 17. 1935; er PE Phytologia 1: 171. 1935; Dop in Lecomte, Fl. Gén. Indo-chine ): 773. 19353; Munz, Man o. Calif. Bot. 37. 1935; Svenson, Am. Journ. Bot, 22: 253 & 271. 19353 Le H. & E. Z. Bailey, Hortus, new rev. ed., 631--632 & 752+ 19353 Moldenke in Fedde, Repert. Sp. Nov. lil: 5, 9, 12, 1s, 17, ng- ton, Papers Mich, Acad. Sci. 22: 62. 1936; Perry, Rhodora 38: 271 1936; Bedevian, Illustr. Polyglott. Dict. Pl. Names 609. 19365 ies Harrow, Journ, "Roy. Hort. Soc. Lond. 61: 398--l01. "19365 K. G 1961 Moldenke, Monograph of Verbena 111 368. agate Ensweller & Blodgett, Proe. Ame er. Soc. H Sci. 4 822--82). 1937; we rst Darwiniana 3: 9, 50, & 5335, 19373 Sudam. p. 16, April 18. 1937; Je ‘ Watkins, Univ. ria: Agric. “Exp. sta. - 89: 36. 1937; Pulver, Horticulture 15: 333. 1937; Pérez Ar- beldez, Plant. Medic. Colomb. 40 & 20. 1937; Von Hagen, Off with their Heads 86. 1937; Latzina, Lilloa 1: 189. 1937; Hermann, Pa- pers Mich, Acad. Sci. 22: 88. 1937; Cory, Texas xas Agr. Exp. Sta. Bull. 550: 88--89. 1937; Bausor, Torre eya 37: “a Bre Moldenke, Syst. Bot. Lect. 22: a 1938; Little, Am. Midl « 38: 385. Linn, 1938; Laudermilk & Munz, Carnegie Inst. Wash. Publ. 87: 279. 1938; Crocker, Bot. Rev. 4: 256. 1938; Monticelli, Lilloa 3: 358. 1938; A. W. Hill, Ind. Kew. Suppl. 9: 12h & 29)--295. 19383; E. D. & Bergalli, Campal, Aragon & Rosengurtt . Nat. Mead. 2h, 81, Su, 102,’109, 112° 116, 118, 128, TMT, 150, 153, 15h, & 156. 1 1938; Re * oe jjertoorn, Union of S. Afr. Dept. Agr. & Forest. Bull. 1853 fig. 90. 19383; P. C. Standl., Field Mus. Publ. Bot. 18: oue“a0ns: 1938; Farwell, Papers Mich. Acad. Sci. 23: ty 1938; Fernald, Rhodora 0: 371 & eS. - 1938; G. Grim, N. Y. 93 9393 Diet. Bot. Names, th pr., 118. 1939; Moldenke in Fedde, Repert. Sp. Nov. 6: 201. 1939; Rep. Bot. Exch. Club Brit Isles 1938: 51. 19395 Fedde, Bot. Jahresber. 59 (2): 417. 19395 Moldenke, Brief Ann & - 190; ¢ 32, maps 1718-172. 190; Potter, Wo io, d, Rhodora |2: 46. 190; Moldenke, Prytologia 1: "132, 138— 1940; is, & 27 MTT—iBo. 1940; Gates, Fl. Kans. 190, maps 1 1066--1068. Darwiniana 4: hl & 55. i9hios Ranson, Texas Wild Fl. 112 PHYTOLOGIA Vol. 8, no. 3 Legends, ed. 2, 113. 190; Madeoke, List Obs. Fl. Watchung }1. 190; Moldenke, Am. Midl. Nat. 2h: 750--753. als Fassett, Man, Aquatic Pl. 286, 380, & a ree Oppenheimer & Evenari, Bull. Soc. Bot. Genév. (Fl. Cisiord.] 31: 363. a) Marchionatto, Las Malegas 33. 1905 Monachino, Torreya 0: 83. i9hO; Moldenke in Fedde, Repert. Sp. Nov. h8:" 25; aT & 29 (1940) and 49: 97--101, 109, 117, 119--121, & 125. 19h0; Beale, Price, & Scott-Moncrieff, Journ, Genet. hl: 65--7h. 190; hl nig pees & Sturgess, Bio- chem. Journ, 3: 268-271. 190; Lawrence & Price, Biol. Rev. 15, no, l. ay deep Journ, Genet. 0: 337358, ple 13-15. 1910; Moldenke, negie Inst. Wash. Publ. 522: 18—150, uae uty 173, 17k, 176, 178-180, 185, & 223. 1940; Moldenke in Pull . Su (23: 268, 272-276, & 279 1903 Moldenke, Lilloa omy 392— ko3 oN) and 6: 321—334. 1941; Gard. Chron., ser. 3, 110: 110 Leaflet Lower Rio Grande "wat. Club 1: 13. 1913 ry ney List Invalid Names 3, 7--10, & 12. 1913 Gongalves de Cunha & Goncalves Sobrinho, Contrib. Conhecim. Fl. Agores 52. 191; Fe Le Herrera, Sinop. Fl. Cuzco 1: 352-~353. 191; : Sperey ul Ross Bull. 22: oe 19hi1; Moldenke, Darwiniana 5: . 19h; a ge Oy 19h1; Calderse & ‘stand, Fl. Salvad., ed. 2, 239. rte a ed. 1], 313, lenhe Lp 50, 50, ¢3, 53, 5é'61, 63, 6h, 66, 70 lol, & 102. i9h2; a & Dawson, Punta Lara 357. 1925 Ripper- ton & Hos — cua Haw. gate Exp. Sta. 89: 55. 1942; Levan, Her~ oditas 28: + dln 1942; Moldenke, Addisonia 21: 59—60 3 67: 303. 19h2; & Peebles, Misc. Publ. U. S. Dept. Agr- 423s 764 (19g) and and 857. sks fe ey ivr £, 3(2)t Poe 382. 193; ndell, Fl. Texas siah3 132. 19133 Little & Campbell, Am. Midl. Nat. 30: 66h. 19135 weno ap - Dicty ed. 5, 1112 & 1115. 1943; G. Grima, 31. 1943; Ramfrez Canté, Anal. Inst. Biol. Mex. 1h: 06. 19435 Sampaio & Peckolt, Arquiv. Mus, Nac. Rio Jan. 37: sen isae 1943 j —— ee a 1961 Moldenke, Monograph of Verbena 13 E. L. Braun, Annot. Cat. Sperm. Ky. 116. 1943; Beetle, Bot. Rev- iew 9: 67h. 19h3; Moldenke, Fl. N. Appal. Exp. Watersh. 28 & 56. 1943; Moldenke, Known Geogr. Distrib. Verbenac. Suppl Journ. Sci. 3: 191. 1943; Niemeyer & Stellfeld Arquiv. Mus. Parans 3: 18. 19433 Allard & Leonard, Castanea 8: 17. 1913; Pots- ger, Trans. Wisc, Acad, Sci. 35: 1)5. 1943; Raimondi, Bol. Mus. Rosengurtt, Estud, Prad. Nat. Urug. 4: 8. 19h3san Dimas Exper. Forest Herb. & Symb. 21, 1944; Fagerlund & Mitchell, Checklist - Nat. Park 53. 194k: EB. L. 1 1279 & 1281. 19h; Moldenke, Iilloa 10: 345—-3)8 & 381--385. 19h); Moldenke, Castanea 9: 69 s Rouleau, Inst. Bot. Univ. Montréal a9 Contrib, 5): 161, 225, & 36. 194k; Bowerman, Flow. Pl. Ferns Mt. Diablo 49, 51, Tl, 72, & 221. 19h; Moldenke, Obs. Fl. Warren 27. 19h; Moldenke, Obs. Fl. Warr Suppl. 1: 11. 194k; Stelifeld, on, Rev. Mus ta, ser. 2, Bot. 5: 356=-357 & 381. 19h; Moldenke, Phytolo- 5, 87~-89 lington & Chranosome Atlas 270. 1945; Parodi, Bol. Soc. — Schnack & Covas Darwiniana 7: 71—79 & 85--86, 5, fig 1& 5. pl. 1-- 2. 1945; Moldenke, am. Journ. Bot, 32: 609 & 610. 1945; Seed Trade ers Guide 86. 1945; c. M. Wils., New Crops 2:2 & 295. ent In- A Ce, Philad., seed packet 2780. 1945: T. H. Everett, Guide Gard. Fls. 39, 19h 65 Cabrera, DAGI Pub. Tec. 3: 27, 5, & 75. 19LR o ty Acufia Cat. Pl. Catamarq. 30 & i bd . 95; Dean, Kriebel, Homans’, & Friesner, Proc. Ind, Acad. Sci. 5h: 95. 19h5; W. Le Renae Luth. Burbank 87 & 505. 19h 3 G. N. Jones, Fl. Ill. [Am. areltts Monog. 2:] 215-216 & 316. 1915; L. I. pavis, Nature Jones, nteower Rio Grande Valley Nat. Club} 2: [h]. 1945; C. Ne 168-3 Ohio Journ, Sei. 45: 165, 1945; Moldenke, Bot. Gaz. 106: denke 7, 19453 Moldenke, Castanea 10: 353 & lS—-l)6. 1945; Mol- enke, Holmbergia 4: 151--152. 1945; Moldenke, Torreya h5: 50 & nke, Am 09--6 Marae Bot. 32: ‘ » Nat. 32: 576 (1945) and 35: 377. 196; Anon., Through 0. t (1): n.p. 1916; Merr. & Reeder, Bartonia 2h: 76. 19h65 eta aSher, Am, Bot, Exchange List.196; Tolstead & Cory, Field - Lk: 39 & 60, 1946; B. P. Reko, Bol. Soc. Bot. Mex. : 35. ny PHYTOLOGIA Vol. 8, no. 3 1M A. C. Martin, Am. Midl. Nat. 36: 608. 1916; Augusto, Fl. Grande do Sul 208--21h & 233, ‘fig. 99. 196; *stellfeld, Vell- site h (5): 99. 1946; W. C. Leavenworth, Am. Midd. Nat. 36: 187. 1946; Rosengurtt, Estud. Prad. Nat. Urug. 5: [Fl. J. Jack- ss ween: fig. 3%. 1946; Schnack & carers Bol. ie Argent. 282-28). 196; Troncoso & Burka: 218. “3906; ne =n up . List Ai li 313 \s--h7, \9--57, al, ish--1h5, 226, 229--236, 239--251, 253--271, 27h, & 276— am aly Pree. Ind. Acad. Sci. 55: 56. 196; Barton & Garman 33: 227-228. lohé3 Moldenke’ Phytologia 2: eecish, & Past shy (1946) and 2: 235=-2h1, fig. 2, 321--332, 335--33, 347, 348, $io- 38h, 386, . 387. 19473 Deam, Yuncker, & Friesner, Proc. "Ind. Acad. Sci. 56: 110. 1973 Falco, Guia —— Jard. ete Roi i 42. 19473 Hill & Salisb., Ind. Kew. 10: 242. 19h7; nese & hee Darwiniana 7: 4h3, WS ws, "ke Wee ights Lor= sacl Veg. Nordeste Prov. Rntre Rfos, ed. 2, 19, 86 » 107, 150, 165, 167, & app. iv. 19473 Hotchkiss & Stewart, Am, Midl. Nat. di h, i, h20--421 & 466. 1 9473 Schnack & Covas, Haumania 1: 5 & 8, fig. 2h&i. 19h7; Daniel, verb, Cent. Antiog. 1--2. 1947; Moldenke, Brief Course syst. Bot. 21 & 92. 1947; B. P. Reko, Bol. Soc. Bote Mex. 5: 34. 1947; Kuhlmann & Ktthn, Flor, Dist. [biti 117 & 182. ke, List Pl. Obs. Branchville 12. 198; cabienes: Cone 13: 116-121, 1948; J. Hutchinson, Brit. Flow. Pl. 129 & 37h. 19485 & e e ° Acad. Se. 198. 1948; Moldenke, Alph. isis Cit. 22 35-375 3712-382; 38h, 389-407, 409-111, 413, hak, 116, 1426, Lar, tk 438, 139, hh 9 bh6, L858, 462, 463, h65—-hBL, 486--198, 508, 510—-5hi2, Si~-562, 565, 567-577, 579—~589, 592-602, 60l 624-635; » Shb--6 65 2. 1948; Deam, Yuncker, & Friesner, OC » Ind. Acad, ia e's 8h. 1948; Neal, In a ‘koma’ re rie 636 & favs Fagor of hn W. Harshberg Lite 23 a6, ie 25, 30, 33, 36, 39, W--hh 73--80, 82--90, oh, & 2 $5. 19483; Mold enke, Phytologia 2: 419--h28, 466, bilie=L79, MB2, & a 511-512 (2948) 32 Lb—-6 (1948), and 3% 63-6, 71-76, 120, 30-13, 138--1s1, 167, & 176—175+ 1949; Re T. aaa pn Sle 11 bare * Exp. Sta. Mem. 291: h2- 1949; Delect. Sen. Hort. Bot. Pisa ilo: 13. 1919; R. Espinosa, 1961 Moldenke, Monograph of Verbena 115 Est Ecuad. 2: 68. 19493 Moldenke, Am. Wild Fls. 291— 293, 36k, & 450. 19493 E. L. Palmer, Handb. Nat. Hist. 297 & 663. 1949; Troncoso, 8: 481--85. 1949; Wan o Buy ted to Buy, Ore- gon Biol. Supply Co. 12. 19493; Rehd., Bibl. Cult. Trees 583. 19,9; Moldenke, Orig. Cult. Pl. and & 19493; Boelcke & Echeverria, Lilloa 8: 2h1 & 250, cuadro 8, 9, 17, & 26. 199; Waterfall, Rhodora 5l: 20 & 27. 19h95 H. N. & A. L. Moldenke » Anal. Inst. Biol. Mex. 20: 12--1). 1949; Cabrera, Lilloa 20: 175 & 337, cuadro I, III, IV, VII & XVII. 1949; Ramfrez Cantié, Anal. Inst. Biol. Mex. 20: 223, 19h9; Castellanos & Ragonese, Lilloa 20: 259. ges W. P. Eaton & al., Bartholomew's Cobble [p. 8]. 199; Moldenke, Known 999, 1001-1005, 1010, 1012--1017, a 1024, 1054, 1055, 1060, 1062, , 1069, 10711095, 1097—1100, 110 31130, 1132-1155, 1158, 1160—-1189; 1191--119h, 1196-1208" 1207=-1233, 1235-1262, 1283, & 1286130. 1949; Heusser, Bull. Torr. Bot. Club 76: 393. pre Loew, Amn, Rep. Huntingto n Goll. Bot. Gard. 13: 37. sh + A. Burpee, Burpee Seeds 1950: 7. 1949; H. S. Gentry, A Hancock Pacif. Exped. 13 (2): 33 & 2h. 19493 F. A. Barkley terminac, Fac. Nac. Agron. Medellin 1 (1): 2& Pa (9t9) and or be nig. 1949; Fernald in A. Gray, Man. Bot., ed. 8, 1208--1211 & Torre Bot. Club 77: Woleeipe 19503 N. Y. Bot. G Exch, List 1951: 4. 1950; Matuda, . Nat. bb: 576. 19503 Zim & Mart- owers 60 & 157, 19505 Noldenke, Lloydia 13: 226. 1950; Stefferud, How to Know s. 100 & 143. 19503 Moldenke, Rev. Sudam. Bot. 8: 165—-169" & 173. pt Turri Curtis, Bot. Mag. 167: vas P “ee 3 M. L. Grant, Proc. Iowa A . Be Sei. 59: 51, — rac & Walter, Seed Annual 1950: is. ; s t, : 145. 19 ep Chile 25: 60. 1951; Moldenke, List Obs. Fl. Watchung Suppl. 206, @-issue: 46. 1951 1; Ragonese, Revist. Invest. Agric. 5: 40, 83, & 208 (1951) and 6: 7h, 76, 82, & 87. 1951; Deam, Yuncker, & 116 PHYTOLOGTIA Vol. 8, no. 3 esner, Proc. Ind. Acad. Sci. 60: 87. 1951; Gottscho, Wildfis. 55 & 192. 1951; Moldenke, Inform. Mold. Set 6 Spec. 4. 1951; N. Y, Bot, Gard. Seed Exch. List 1952: h. 1951; Stellfeld, Trib. Farmac. [Vellozoa] 19 a 166--167 & 172. 1951; Martinez Cro- vetto & Piccinini, Revist. Invest. Agric. 4: 77. 19245 W. A. Bur- ds 1952: 46. 1951; Kennedy, Kwality Seeds Spring : 1. Bot. 2h: 198--199 & 27h. 1951; Moldenke, Sp. Subsp. Cont. Mold. set 5 [3]. 1951; Moldenke, Biol, Abstr. cin ee 1981; McDougall & Sperry, Pl. Big Bend Natl. = 46. 1 om 3 Anal, Inst. Biol. Mex. 22: hl. 1951; Rioja & Herrera, Anal. » Biol. Mex. 22: 58. 1951; He N. & A. L. Moldenke, “pl. Bible si & 326. 1952; Lid, Norsk Fl. 530 & 771, fig. 303a. 1952; Deam, Yuncker, & Friesner, Proc. A- cad, Sci. 61: 75. 1952; G. Grimm, N. Y. Herald Trib., sect. h, p- 9 ’ 3 & B77. 1952; Moldenke, Phytologia : 58, 66--68, 70, Ti, 7h, 119, 121, & 128-129 (1952) and h: “nike, 188, 190, 199, *266, 267, 293, & = ko-ace, 19533 Moldenke, Biol. "abstr. 27: 1807 & & 3121. 19533 Cabrera, Man. Fl. Alreded. ra ee Aires 298. 19533 J» We Johnston, N. Y. Herald Trib., sect. h, p. 638 Feb. 15. 19535 Henderson, Stumpp, & Walter, Garden Anmal 1953: 16c & 17. 19535 arden, a a Statesman, Aug. 25. 1953; Schnack & Sol- brig aon aon Plata 29: 255—266, fig. 1--h. 19535 C, Skot TYecets 3 uan Fern. 922 & a a 1953; Moldenke, Hor- eiihive 3: 178. 19835 E E. J. Salisb., Ind. Kew. Suppl. 11: 101 & 262-263. 1953; ere aes mat . 409 & & 909 (2953) and 2: en tossed Wherry, » Club 80: 228. 1953; We Ae Burpee Seeds ers iB Cl (1953), 95h: 48 (1953), and 1955: et 19¢h; Lombardo, Invent. Pl. Cult. Montevid. 232, 248, & 262+ J 195h3 Moldenke, Journ, Calif, Hort. Soc. at 80 & 33-83" 195h3 Je Rzedows ki, inal. ani Nac. Cienc. Biol. 8: 105 & 112. 195h; A. R. Magalhies, Flores Bras. 1: 20. 195k; Moldenke, Inform Mold. Set 1,8 mt tid (h). 95h5 Biol. Abstr. 25: 062 (195k) and = Stell & 3534. 195h5 Rast , Sellowia 6: 60, 8h, & 153. 195h5 rne, Am, Midl, 6a, t. Fle 5h; Hylander 0, pl. 166. 195h5 Fe Ce Chandler, U. S. Patent off. Pl. 1330-~133. 1954; Breck, ra to _ oo Gard. 1954: 27- 195h5 Do. i Peay rd _ it eg i 15, Feb. 21. 195h; « Sci. ath 70. “ashy "Woldenke, Phytologia be 198, TL, 95, n96, "498, 501, 502, & 512 (195k), 5 i 2h, » 70 iS iB, 92, 94-97, & 1 ‘ (195k), and St 132——. 13h, & 227-229. 1 9553 Stanek, pad re Nat. 25 & 370. 1955; Biol. Abstr. 27: 3735 & 3768. 19553 He He Sme, Bull. Torrey Bot. Club 82: 307. 1955; Moldenke, Mem. N. Y- Bot. 1961 Moldenke, Monograph of Verbena 117 19553 Gatsadiuntt ature 17: iok, pla 12% 1955; Rzedowski, Ciencia 15: 146 & 148. 1955; Moldenke in Chees poo pe neo - & Tob. 2 (6): 39be-395 & 397. 1955; Moldenke, Fam. 1unthyd 36, eit Moldenke, Biol. Abstr. 29: 293h (958) nin 30s "1092—~1093. 1956; angely, Gat. Estat. Gen. Bot. 17: 6+ 1956; McVaugh, Edw. Seabee . pee, Moldenke, Inform. Mold. Set 51 Spec. h & 5. 19563 & 271. 1956 Amaral *Lisbéa, Anais V Reun. Anual Soc oc. Bot. Bras in Camp, Boswell, & Magness, World in You Bhp 82 & (831. “ws Biol, abstr. 2 29 "3591 & 3630. 1957; reece - Purandhar 1 10h & 158. 1957; Lehr, Bull. Torr are Bot. Club a 312. 1957; ch State Univ. P. Notes Pl. 2: 1 & 6, 1957; pecs & G. C. Rzed ; jf Potos. 1: 39 & 55—57. 19 7; Cabrera, Reviet “intost 332 & 398. 1957; Alain in Leé ain t 279 —— 281, fig. - 19573 Vaughan, Gard. _lilustr. 1958: ile “Sh, ‘ 38 fig. 5738 & Ae 1387; Rzedows W. A. ee Seeds eT: 33 ; "38. (1957) and Cake 12 & 39. 1967; . 5. Ww. Johnston, N. Y. Her rib., sect. k Pe 12, Feb. 2h. 957; Benson, Pl. Classif. 62, 112, 227, hs, & 667. 1957; Cour- en, 36 (1): . 1958; R. S. Lemm e 16—17, pl. 11. 1988, .: Zs n, Rhodora 60: 69. 1958} Moldenke, Am. Mid]. Nat. 59: 33h & 3h1--370. 1958; Angely, . 17. 1958; R er, Exkursionsfl, Deutschl. 373 & 501, fig. 373/ » 1958; De Roon, I ect. Spec. 229, 19 ne a ot + Re Va. 3: 772 & 774—777. 1958; Ahles, Bell, a on, Wasmann Journ. 16: 5. 1958; Dandy, List Brit. Vasc. Pl. 101 & 176. 1958; Shin- ners, Spring fi, Dallas 323-326. 1958; Moldenke in Gleason, New ardening I 5749. 1958; Biol, Abstr. 30: 38 3856, Woldenke, Phytologia 61°235, 33° & 331 oe 1959 >, 93 “yee Bull. pre club "86: UNS 19593 G. Taylor, Ind. Kew. Suppl. 12: 63. & 19. 19595, Vaugh- roo othaty ening Illustr. 8): "3 ‘& 51, tie. 5738, 570, & 519. Limis, ‘spat ae aes me 2 i 80, 81, 8h, 8%, 109421 a8 160" uir—a22, "126-138, ike, 13, ahs, Ue, hy, 45h, 156, 158—~ 16h, 166, BT, eam ri 1 181, 182, 185, 188, 191, 118 PHYTOLOGIA Vol. 8, no. 3 aioe 222—-226, 23h, 235, 237, 2h0, 275, ont = se em 318, » 343, 3hh, 355, 357--379, 393, 397, 399, 405, 15-28, 4 moiith & 193-95, *Verbena teasii Moldenke = V. hybrida Voss x V. temisecta Briq. xVerbena torpa Moldenke = V. neomexicana (A. Gray) Small x V. ur- ~ticifolia T L oie ee gr Verbena transitoria Moldenke = V. megapotamica Spreng. x V. tem- ~ isecta Briq. xVerbena - trinitensis Moldenke = V. incisa Hook, x V. tenuisecta erbena re See Moldenke = V. peruviana (L.) Britton x V. — sae Moldenke = V. megapotamica Spreng. x V. santiaguensis (Covas & Schnack) Moldenke *Terbena wingei cineie V. canadensis (L.) Britton x V. temi- 122 PHITOLOGIA Vol. 8, no. 3 secta Briq. An intergeneric interfamily hybrid has also been described and illustrated, xVeronicena haartmani Moldenke, said by Plantae hybridae 7--8 (1751) and Amoen. Acad. 3: 35, ple 2 (1756), to be a hybrid between V. officinalis L. and the scro aceous Veronica maritima L. The status of this veto: however, is highly questionable, as has been pointed out by me in Am, Midl. Nat. 59s the 171,938 herbarium specimens of my groups that have passed through my hands in the past 32 years there were two specimens which may possibly have represented this same or a similar taxon. ! 2 ented a plant much like the one figured ° The sheets on ch they were mounted -- as ecall them now were both quite old and had appa been bandied about from one group. I recall "Scroph.?", "Labiat.?", and even "Lobeliac.?" in- scribed on at least one of them, with each suggestion later strick- en out by pencil. Perhaps the curators of the herbaria in ques pres on reading this, may recall the specimens and may re-examine in the light of Haartman's claims. Unfortunately, having aaateee material of my groups from 260 po ee I cannot now recall to which herbaria they were returned As far as I am aware, this claim of Haartman is the first sty hybrid between eye belonging to entirely different families either plants or animals, Generally such an inter-family yn is regarded as biologically re Bag FT and this alone tends to ¢ much doubt on Haartman's claim. It is very possible that his con- cept of a "hybrid" plant mae quite different from our modern con Verbena and Veronica as genera, and the Verbenaceae and Sc ular Gaceae which they a tlie The chromosome complements of both supposed parents are also Cabrera, Covas, Dawson, “Gaelin, “Vattfeld, Nuttall, Ragones®, Schnack, Rozier, Small, and S Solbrig have all suggested the segregs tion of the genus Glandularia J. F. Gmel. from Verbena. Baia I think that this is quite justified, but because of the great Tit ber of nomenclatorial changes which this would nece ceaiuate tn er ly known species, including cultivated ones, I have no not followed this course in the present work. Schnack & Covas, in Bol. Soc. ots gent. Bote 1: 282--28) (1946), summarize the situation as follows "El estudio de mumerosas especie s inclufdas en el género verde (sensu pencdines en D. C., Prodr., ll: 550, 1847) nos ha pe - t 1961 Moldenke, Monograph of Verbena 123 confirmar el cardcter artificial de esta entidad, y correspondi- entemente la validez del género Glandularia, recientemente rehab- ilitado.....y que comprende una serie de antiguas Verbenas. En efecto, hemos podido establecer la concomitancia de una serie de caracteres de la m4s distinta naturaleza dentro del género Glandu- laria, por una parte, y del género Verbena (en su sentido actual dularia, y s e a los caracteres de las especies de Verbena, que no vacilamos en distintas y naturales, en las que fenéminos de convergencia de formas conducen a una similitud pio de especies que pertene- cen a dos lfneas evolutivas distin They summarize the differences as Tens in Verbena (1) the saps are plainly quadrangular, (2) the chlorophyllous ‘lous cortical mchyma is discontinuous (interrupted at the angles), (3) the att ccustanas is paniculate, many-spicate, (l) the oss funnel~form, (5) the thecae of the anthers do not surpass thi connective, (6) the style becomes approximately 3 times as sie ricarps are the th the pericarp not prolonged either laterally or at the base of the spor sector, and (8) the chromosomes are x = 7 and - InG a, on the other hand, (1) the stems are v: (2) the chlorophyllous cortica longer than th » (7) the mericarps are brown-blackish, with the pericarp dilated laterally or rather prolonged ben embryonal sector, fo an excavated b (8) the chromo- 6, and Somes are x = 5 and comparatively larger. Mattfeld (1951) says: ry bisher mit Verbena (x = 7) vereinigte Gattung Glan - (x = 5)......musz als eigene Gattung abgetrennt werden." Meioner and Walpers used the generic name Shuttleworthia and Bunge proposed Uwarowia for what is essentially this group. In this connection it is interesting to note that beg de have reported the fo Ollowing chromosome numbers for species Verbena ( a, ding Glandularia): n = 5, _ A Gissecta, V. erinoides eS, tennisecta], V. erinoides var. alba (probably tenuisecta var, al alba], V. flava, V. gran Probably cenadensis), ve. Hitt, V. laciniata, V. mendocina, ¥. microphylla, V V. parodii, V. pe ¢ per rakii, V. peeee, Ys V. phlogi- flora, ¥. platens ais, ve. radicans, V. tia onal, 2 , tenera, ee us crithmifolia, v. hookeriana, V Facemosa, and V, wrightii; n = 1 ~s oe oe kA ree ty ve 12h PHYTOLOGIA Vol. 8, no. 3 ambrosifolia f. eglandulosa, V. bipinnatifida, V. cameronensis, V. 8 var, atroviolacea, Ve canadensis var. -, compacta, V. can- adensis, V. ciliata, V. delticola, V. elegans var. aspera rata, Ve gooddingii - var. nepetifolia, Vv. racemosa, sa, and Vv. tencriitelias | n= 7, 2n = 1) — V. canescens var. roemeriana, Ve , carolina, V. clow erae, Ve halei, | V. hastata, V. hispida, V. macdougalii, ve. neamex- icana, ‘Ve neomexicana var. var. hirtella, V Ve officinalis, ve perennis, v. plicata, V. prostr V. prostrata foe ah V. simplex, V. stricta, and V. oe n= 1), 2n= 28 —V. bonariensis, V. brasili- ensis, V, ehrenbergiana, and V. litoralis; n= 21, n= 42— Ve pa ATER V. rigida, V. mutha. It is to be noted that workers have obtained different chromosane mumbers for V. canadensis an * V. racemosa, perhaps due to misidentification of the material Walpers (185) classified the genus Verbena as follows: Sect. i. pineate ides Walp. Sect. 2. Verbenaca Walp. B § 2. Inermes penteps * Folia fasci Ae Relagindidies Walp. B,. Thymoideae Walp. #* Folia opposita non fasciculata A. Junceae Dp. B. ee eae ls a elindres Walp. p « Aubletia (Neck.) Walp. In this classification his Section 1 is synonymous with the genus tains 2 species of Diostea, 2 of Junellia, and 1 Neosparton, arton, as well as 3 true species of Verbe: Verbena. a (1847) divided the genus as follows: - 1. Verbenaca W; Subsect. 3. Pachystachyae Schau. Subsect. h. rere ioree Schau, Subsect . 5. Schau. Sect, 26 Gieeeieia 73 (Je F. Gmel .) Schau, 1961 Moldenke, Monograph of Verbena 125 In this classification his Section Pungentes is now a part of the genus Junellia, while his Section Verticilliflorae is divided a- mong Diostea Miers, Junellia Moldenke, and Neosparton Griseb. In the sy: system adopted by myself in 1959 and later weg the genera aan subgeneric groups of the Tribe Euverbeneae are guished as follows: 1. Calyx more or less tubular; fruit dry, included 2 the fruiting- calyx, usually separating into cocci when ripe ns h. + Woody shrubs or subshrubs of desert and wpm regions; leaves usually greatly reduced; inflorescence reduced. 4. Calyx deeply 5-parted almost to the base, with long fili- form lobes, densely long- villous; stigma obliquely capi- tate, apiculate; leaves densely imbricate; Chile. Genus Urbania Re As Phil, ha. Calyx m mostly only 5-toothed, with short, often obscure eeth, rarely long-toothed, usually not "long-villous 3 ration not usually closely ame ae Argentina, Chile, Bolivia, and Peru. Genus Junellia Moldenke. 5. Leaves. alternate, simple, fascicles. Group Selaginoid- eae Walp. Sa. Leaves opposite. 6. Flowers in small dense heads; leaves parted or entire, more or less stiff and thorny. Series Acerosae Briq. Te go distinct from the foliage. Group Seriphioid- alp. 7a. Seties and bracts spinescent. Group Erinaceae Walp. 6a. Flowers in 1- or 2, rarely few-flowered cluster: oe at the tips of the branches; 3 low, often tufted or sod- eee eae plants with tangled etchaas S Pungentes Schau, 3a. letdieae plants or subshrubs, not limited to desert or high-alpine regions; leaves and inflorescences usually well developed, 8. Fruit not beaked, composed of cocci, easily separable when ripe. Genus Verbena L. 9. Anthers unappendaged. Subgenus Verbenaca (Walp.) Lewis. 10. Plants often subshrubby, usually with greatly reduc- ed or even scale-like leaves. Series Junceae Walp. 10a. Plants mo stly herbaceous; foliage mostly not reduc- ed. Series Foliosae Walp. ll. Young spikes fastigiate-capitate, Subseries Mac- ran Pe 12. Leaves crenate, dentate, or subincised. Group Melindres Walp. 12a. Leaves incised-lobed, aegig or bipinnati- fid. Group Aubletia (Neck. ) Wi lla. Bie: spikes not ANA se EA corollas Ostly small and indi vidually not showy; calyx rare- 126 PHYTOLOGIA Vol. 8, no. 3 ly twice as long as the fruit, ia contorted be- yond it. Subseries Micranthae 13. Spikes abbreviated, the Frees er: Group Pachystachyae Schau. 13a. Spikes elongated, filiform, the flowers scat- tered. Group Leptostachyae Schau. 9a. Some anther-connectives usually appendaged; corolla mo large and s 3 calyx usu than twice as long as the fruit and constricted or contorted be- yond it. Subgenus Gl ia (J. F. Gmel.) Lewis. 8a. Fruit beaked, nutlet-like, not readily separating. Gems Stylodon Raf, 2a. Fruiting-calyx accrescent, broadened and inflated at the base; stamens 2, Genus Hierobo tana Briq. la. Fruiting-calyx broadly campanulate; fruit a Ae oe Arne, with ste or siege fleshy exocarp and hard endocarp. Gen Ghinia § It is worth es here that Verbena Dill. is a synonym of Priva Adans. and that Verbena Pluk. is synonymous with Stachytar- heta Vahl. The gems ; Verbena L. was placed in Family 75, Labia~ tae, Section Verbeneae, “by Hei Heinrich Gottlieb Ludwig Reichenbach in Messler, Handb. Gewichsk., ed. 1, 1: xcocvi (1827) and ed. at li Lexy (1833) » and also by him in his se Regni Vegetabilis » Flor. Dist. Ibitd 182 (1947) say "Verbena spp. -- As herbas agrupadas neste genero aparecem geralmente em e em terren- os baldios, cultivados e nos quintais. A 'camaradinha' ' juruj- ba' € cultivada nos jardins. Produzem flores de colorido * varidvel conforme a espécie ou variedade, também visitadas por Lepidépteros e 'Abelhas'. Os frutos sao secos, nada demneciando de aprecidvel suas relacoes com a Fauna," Rehder (1949) includes in the synonymy of Verbena the following genera: Burseria oe [actually a synonym of Priva "Priva Adans.], Sty- a Raf., Styleurodon Raf. [a synonym of Stylodon], and Junellié@ unellia °. csr is "n egligib le". Junell (193)) Giscusees the gynoecium morphology: Schauer (1847) says that there ar 2 stamens in one species — f the genus (1%) that the posi oe very r ed. Schau now abe botana, Junellia, and Stylodon. The Gmelin reference cited by the bibliography (on previous pages) is often cited by other auth- ors as "Gmelin, Syst. Veg. 920. 1791", The genus Billardiera sometimes cited to "Moench, Meth. 366. 1794", but "369" is the CoP” rect page citation, Verbena is sometimes cited to "L., Syst. Veee ed. 1. 1737", but I have thus far been unable to trace this refer metre & ges! 1961 ve a very interest: a of the genus under V, mite ot "This eka rout in ast-central 1961 Moldenke, Monograph of Verbena 127 South 7 ae Aa might have arisen as an amphidiploid in a a between an n = 5 and n = 7 species followed by genomic to reach this polyploid — Dermen's (1936) inability ¢ to pro~ duce an intersectional hybri d, the rare occurrence of 1 parent in South America, and the unquestioned classification of V. ovata in the section Vickenace (the taxon re ae morphologically . intermed- iate) are all evidences against h an origin, Alternately, V. to Glandularia., Additional cytological studies might reveal other 'relic' taxa in South America, but the very existence of V. ovata — the hypothesis of an ancestral x = 6 stock. Since many the wides in the tropical enviro: and lowlands characteristic of this period, With the aaa of the Cordilleras during the Lamaride Abaca the phylad was reduced considerably in distribution. € of the new colder and drier environments adjacent to the Guy in the late Cretaceous and Plaeocene gave the neces- Sary stage for the evolution of new taxa. On the sprang b aries of the phylad's now limited r e, the ascendi A sate ee originated, perhaps during the Sorte dl iy) Tertiary. This volution gave rise to the now dominant Sort i ratty Similarly, the x = 5 descending wnewietts a tee in South America was evolved (section Glandularia) and became a- rene to the more temperate environment in that —— By hy: could not but Levan a and if the high level of eulgplalay vensind | ta V. ovata is an emample, this factor also may have contributed to the near ex- tinction of the x = 6 phylad. The final result is an amphitropi- cal distribution for Verbena. "The question of both Verbena sections occurring in North and South America @ remains to be answered. Migration of diploid or tetraploid Glandularia, from South America via trans-tropical ridges, similar to the distribution of V. litoralis, or by ng-range dispersal, are plausible explanations. Probably this sigrtion took Place during the later Miocene or Pliocene in a of extension of the upland temperate regions into . Geoptet forests. Once established and after becoming dis- Sar taxa in North America. Very likely a migration southward °f one or more North American Verbenaca also occurred during 128 PEYTOLOGIA Vol. 8, no. 3 these periods and formed the basis for the high level of poly- ploidy in South America currently unknown to the north. These species, new to both continents, successfully pth anirl with the taxa present, wi po lyploidy as an evolutionary mechanism had a — advantage over the widespread (diploid) popula- tions rnacular names for the genus as a whole are "aublétia", vaubléticr, "camaradinha", "camaradinhas", "cambarés", " cedrén" P "i jzerhard", “iron-weed", "jujubas", "jujubes", "jurujba" garita colorada", "margari z6", "moradilla", "salvia montés", "Shah vourite", "Sha-passan", "tep ", "tlal- a ahuehuetl", "verbena", "verbenas", "vervain", "vervains", "verveine", "vervéne" » "verveyne", "wijkruid", and "yapau". Actu- e give these names as generic designations. It is worth noting here that "false verbena" is Cochranea anchusaefolia (Poir.) Gttrke in the Heliotropiaceae and "sand-verbena" is the Parse abronia Juss. in the Nyctaginaceae, while the name "verbena" is often applied also to Bouchea prismatica (L.) Kuntze, B. prismatica var. longi-~ rostra Grenz., Clerodendrum fragrans var. pleniflorum Schau., Priva la ea (L.) Pers., “ Stachytarphe eta cayennensis (L. C. Rich.) Vahl, s. fg na Moldenke, S. frantzii Polak., S. jam aicensis (L. ) Vahl, and S. guatemalens sis Moldenke, as well as to Salvia polystachya. Ort. in the Lamiaceae; "verbena celeste" is Stachytarpheta frantzii, "verbena de playa" is S. janaicens”*, "verbena manza" is Ss. Jenni ensis and Bouchea prismatica var longirostra, "verbena cimarrona" is Bouchea prismatica, nyerbena de flos grande" is Bouchea rusbyi Moldenke, and " verbena morada" is Stachytarpheta frantzii. "Verveine" is oe applied to Phyla nodiflora (L.) Greene and to Stachytarpheta jamaicensis, Wyerveine queue de rat" is S. jamaicensis, "verveine 4 fleurs rouges" is S. mitabilis (Jacq.) Vahl, "verveine & trois feuilles" is Aloysia triphylla (L'Hér.) Britton, "verveine Levi and "verveine queue de rat" are also S. indica (L.) Vahl, citronelle", "verveine Reeve “Wverveine du Pérou" are Aloysia triphylla, *verveine du Mexique" is Priva mexicana (Le) Perse, "verveine du pays" and "verveine sauvage" are Phyla nodiflora “verveine faux-gerva6" is Bouchea fluminensis (Vell.) Yoldenke, "verveine caraibe" is Wedelia car: carnosa Rich, in the Carduaces®, "verveine puante" is Petiveria alliacea L. in the Petiveriacel’: According to Augusto d'Halmar's "San Miguel y las Verbenas madri~ i lefias", there are some fiesta dances in Spanish-speaking countr known as "verbena". In Chilton County, Alabama, there is a tow? and post office named "Verbena", ilieieiaiiaeicieeieiabteamiien Mapping by county done by Andrew Re Moldenke Figure 1. Distribution 9f Verbena in the United States So oem se boots TOWNER | Cavauirr ear ER) 7 S O's ty q PINE f RAINS KEOKUK | "Se THAYER a coma s Pog vi | a ! F a Lar Ra a es 3 27 ek emer “a. , Oe was ae Pe aha & CLEARTYPE COUNTY OUTLINE ne ‘ a abe , be UNITED STATES | Scale of Miles 2 0 100 200 MAP NO, 125 COPYRIG: u AMERICAN MAP COMPANY, INC. ORIGINATO Spee Sou = ee ars Sx CLEARTYPE MAPS S270 COLORPRINT MAPS TRADE MAAN REGISTERED tr evtiratenteach erga “a SPO Y = GO Roe ee ea ei ase: sik eae Oe aie Leb osc USUNEES, ? . ' he Lay ny’ e ix e pS IN Ek ort Sol ALEUTIAN ISLANDS +o | ae. - O, P-., THIRD JUDICIAL DIVISION Ste = >» o 2 ° s j ; ; “ Sd f* ®; am Copyright, American Map Co., Inc., New York, No. 14324 Scale of Miles a) aoe ) z 100 200 et z. y RL = & EES L 2 l 3 | 4 [ 5 i! G I ST I 2 I 3 I u T 7 T % 1961 Moldenke, Monograph of Verbena 129 The maps appearing in this monograph are designed to show at a glance the known distribution of the genus and its constituent species in the United States of America, based on the specimens thus far examined by me. I am deeply grateful to my son, Ralph Moldenke, for the hundreds of hours of his time which he sacrificed from his entomologic studies in order to prepare these maps for this work. No attempt has been made to indicate the rel- ative abundance of records from a given county -- there may be records within the county or parish. A county or parish (or is- land) is blackened on the map to indicate that we have a record this country. Counties where more intensive collecting is requir- ed are pl sh this map, since it is hardly likely that the genus does not occur in counties completely surrounded by counties from which we have records of its presence. Students living in or near these counties are urged to search in wu for specimens of this genus and communicate them to me so that may eventually fill in the empty spaces in a future edition "of this map and the maps that follow. In the Linnean Herbarium at London the specimen number under genus 35, Verbena, is Stachytarpheta indica; number s is S. jamaicensis -- and they are very different from each othert Number "3" is labeled "prismatica" in the handwriting of Solan- der and bears the annotation "Br." [=Browne; Patrick Browne's herbarium per purchased for Linnaeus in 1758) tion in the handwriting of Smith "Stachytarpheta prismatica Sm. in R. Cycl. No. 11." It is actually Bouchea prismatica (L. Kuntze, with the beaks of the cocci 1.52.5 mm, long. Number "lh" 4s labeled "mexicana" in Linnaeus! handwriting and is Priva mexicana (L,) Pers. Number "5" is ee "lappulacea" in the | handwriting of mes i and bears the notation "Br." It is Priva handwri tion "Br." [=Browne] and "Rolander" [Rolander collected in Suri- nam after 17h}; 4 it is Salvia occidentalis Pits Number "7" is abeled "stoechadifolia" in the handwriting of Solander, and is Phyla stoechadifolia (L.) Small. Number "6" is Taeean todifiora” in the handwriting of Solander, and is Phyla nodiflora (L.) Greene. There are more specimens following of this and other spe- Cies. On specimen number "1" vanes genus 755, Phryma, Linnaeus awri ng Verified by Mr. Savage, of the Linnean Society, in each case]. On — reverse side of sheet number "3" there is a description in an "Verbena. Cal, l-phyllus cylindricus, etc." The specimen ed "hians" on the front of the shea by Linnaeus (never aan 130 PHYTOLOGIA Vol. 8, no. 3 under Phryma until by B. D. Jackson in his Index of the Linnean Herbarium) and is oe Chascanun arith ta (L. f.) Moldenke. The appa fet specimens in his herba rue verbens na species will be discussed hereinafter und taxa invol Following is an aiuhabetais "Ee of names 5 eee to species and varieties excluded from the gems: Verbena alatocarpa Troncoso = Junellia alatocarpa (Troncoso) Mol- denke Verbena alpigena Walp. = Aloysia scorodonioides (H.B.K.) Cham. Verbena alternifolia Herb. = Junellia selaginoides (Kunth) Molden- ke Verbena alternifolia Hort. = Junellia selaginoides (Kunth) Mol~ denke Verbena alternifolia Sessé & Moc, = Stachytarpheta incana var. angustibracteata Moldenke Verbena ameghinoi Speg. = Junellia ameghinoi (Speg.) Moldenke Verbena americana Mill, = Stachytarpheta jamaicensis (L-) Vahl Verbena americana flore coccinea spicato Breyn. = Stachytarpheta ann Seeneenemmenepeneeatlimemesmnemmemnemmeesnmmnenee essere atti ee tee a eee — angustifolia Mill. = Stachytarpheta angustifolia (Mill.) Hews aphylla Gill. = Neosparton aphyllum (Gill. & Hook.) Kuntze Verbena aphylla Gill. & Hook, = Neosparton aphyllum (Gill. & Verbena arborea H.B.K. = Petrea arborea H.BeK. Verbena arborescens Bojer = Lippia javanica (Burm. f.) Spre Verbena aretioides R. E, Fries = Junellia aretioides (R. E. grt Mo eV Verbena acaoncsip es Hayek = dJunellia hayekii Moldenke Verbena aretoides costed = Junellia | hayekii Moldenke Verbena aristata | Vahl = tachytarpheta orubica orubica (L.) V. Verbena asparagoides mp = Junellia asparagoides Phyo & Hooke) ~~ Moldenke Verbena asparagoides Gill, & Hook. = Junellia asparagoides (Gill. ~~ & Hook.) Moldenke Verbena asparagoides var. frmetinensiis Hieron. = Junellia aspara~ goides (Gill. & Hook.) Moldenk Verbena aspera Gill. & Hook, = Junellia aspera (Gill. & Hook ») Moldenke Verbena aspera Htgel = Priva cordifolia (L. f.) Dru Kt azorelloides Speg. = Junellia azorelloides (ania ) Molden- Wetues bifurca Benth. = Bouchea prismatica (L.) Kuntz Verbena bisulcata Hayek = Junellia bisulcata (Hayek) yoldenike Verbena bonariensis Rendle = Junellia ia patagonica (speg.) Moldenke Verbena briquetii Macbr. = Junellia s aeite ies (Gill. & Hook.) 1961 Moldenke, Monograph of Verbena 131 Moldenke Verbena bryoides Echeg. = Junellia asparagoides (Gill. & Hook.) Mol denke Verbena bryoides R. A. Phil. = Junellia bryoides (R. A. Phil.) Moldenke Verbena caespitosa Gill. & Hook. = Jumellia caespitosa (Gill. & Hook. } Moldenke Verbena cajanensis L. C. Rich. = Stachytarpheta cayennensis (L. C. Rich.) V Verbena cajenensis Rich. = Stachytarpheta cayennensis (L. C. Rich.) Vahl Verbena callicarpiaefolia Kunth = Lippia callicarpaefolia H.B.K. Verbena cannabifolia Sieb. & Zucc. = Vitex negundo var. cannabi- ~ folia (Sieb. & Zucc.) Hand.-Mazz Verbena ca capensis Thunb. = Lippia javanica (Burm. f.) Spreng. Verbena capensis f. capillaris Ray = Hebenstretia erinoides L. f., aginaceae Verbena capitata Blanco = Phyla nodiflora (L.) Greene capitata Forsk. = Phyla nodiflora (L.) Greene Verbena capitata Hort. = = Lippia javanica (Burm. f.) Spreng. Verbena carnea 1 Medic. = Stylodon carneus carneus (Medic.) Moldenke Verbena carolinense (Walt.) Gmel. = Styl Stylodon carneus (Medic.) Mold, Verbena carolinensis Gmel. = Stylodon carneus (Medic.) Moldenke ena carolinensis Small = Stylodon carneus carneus (Medic.) Moldenke Verbena carolinensis Walt. = Stylodon carneus (Medic,) Moldenke Yerbena arotinensis (Walt.) J. F. Gmel. = Stylodon carneus (Medic.) Mol denke Yerbena eeeenensis (Walt.) Small = Stylodon carneus (Medic .) Verbena pertiiia Hort. = Stylodon carneus (Medic.) Moldenke Verbena caroliniana L. [ex Raf.) = Stylodon | carneus (Medic.) Mol- soe enke rete caroliniamm L. = Stylodon carneus aaa Moldenke Verbena caroliniensis (Walt.) J. F. Gmel. = Stylodon carneus nk Verbena carroo Speg. = Junellia tridens (Lag.) Moldenke Verbena carréo carréo Speg. = Junellia tridens (Lag.) Moldenke Verbena castra castrata Sessé & Moc. = Stachytarpheta cayennensis (L. C. Rich.) Vahl Sieh: Verbena Caudata Salisb. = Stachytarpheta indica (L,) Vahl Yerbena cayame ets Rich, = Stachytarpheta @ cayennensis (L. Ce h.) Vahl Verbena ¢ cayennensis L. C. Rich. = Stachytarpheta cayemensis (Le 132 PETIT OLO Gas Vol. 8, no. 3 C. Rich.) Vahl Verbena cedroides Sandw. = Junellia cedroides (Sandw.) Moldenke Verbena ceratophylla Bonpl. = Hierobotana inflata (H.B.K.) Briq. Verbena chubutensis Speg. = Junellia wilczekii (Briq.) Moldenke Verbena cinerascens Schau. = Diostea cinerascens (Schau.) Molden- Verbena citriodora Cav. = Aloysia triphylla (L'Hér.) Britton Verbena citriodora Ort. = Aloysia triphylla (L'Hér.) Britton Phil colchaguensis (R. A. ee) Moldenke Verbena comberi Sandw. = Junellia comberi (Sandw.) Moldenke Verbena eee recess oats = Junellia connatibracteata ~~ (Kuntze) Moldenk Verbena Rainatehirectaate f. glomerata Monticelli = Junellia con- natibracteata f, glomerata (Monticelli) Moldenke — Verbena connatibracteata f. normalis Monticelli = Junellia connat- Verbena connatibracteata f. rosulata Monticelli = Junellia connat- denke : : 4 2 |i » p E e a # Bo i 58 Verbena cordata Hort. = Alo; Aloysia chamaedryfolia Cham. Verbena cordata Mackay = Aloysia chamaedryfolia Cham. Verbena cordifolia Bourgeau = Priva mexicana (L.) Pers. Verbena cuneata Willd. = Phyla nodiflora (L.) erent? Verbena curassavica L. = Ghinia curassavica (L.) Mi Verbena decurrens Moench = Stachytarpheta ey oes (Le ) Vahl Verbena dentada Vivian, = Aloysia chamaedryfolia Cham. Verbena dentata Hort. = Aloysia c chamaedryfolia Cham. Verbena dentata Juss. = Aloysia chamaedryfolia sal Verbena dentata Visian. = Aloysia chamaedryfolia Verbena di diandra Sessé & Moc. = Bouchea a active var vare ; previrostra ee Grenz Verbena diandria Larrafi, = Bouchea prismatica var. brevirostra Grenz. Verbena dichotama Larrafl, = Stachytarpheta cayennensis (L. C- ~ Rich.) Vahl Verbena dichotana Rufz & Pav, = Stachytarpheta cayennensis (L. Ge ~ Rich ¢) Vahl Verbena Sigitata R. A. Phil. = Junellia digitata (R. A- Phil.) ~~ Moldenke Verbena pepe Duchassaing & Sagot = eta jamaicensis = ) Vahl Vahl Verbena diversifolia Kuntze = Junellia diversifolia (Kuntze) Mol- enke Verbena dolichot Sandw. = Junellia dolichothyrsa (Sandw.) Mol denke ymeliis dole 1961 Moldenke, Monograph of Verbena 133 Verbena echegarayi Hieron. = Junellia echegarayi (Hieron.) Mol- Verbena echegayari Hieron. = Junellia echegarayi (Hieron.) Mol- Serten echinata R. A. Phil. = Junellia seriphioides (Gill. & Hook.) Moldenke Verbena elliptica Willd. = Phyla nodiflora var. canescens (H.B.K.) Moldenke Verbena ellipticifolia Stokes = Stachytarpheta indica (L.) Vahl Verbena erinacea Gill. & Hook. = Junellia erinacea (Gill. & Hook.) oldenke Verbena etchegarayi Hieron, = Junellia echegarayi (Hieron.) Mol- denke e Verbena fluminensis Vell. = Bouchea fluminensis (Vell.) Moldenke Verbena foliis cordato-ovatis, floribus spicatis, calicibus in flatis, seminatis echinatis P. Browne = Priva lappulacea L.) Pers. jeu Pave verticaliter ovatis, spicis globosis L. = Phyla nodiflora (L.) Greene Verbena foliis verticaliter ovatis » spicis solitariis ovatis Royen = Phyla nodiflora (L.) Greene Verbena forskaelaei Vahl = Priva cordifolia (L. f.) Druce Verbena forskaelei Vahl = Priva adhaerens (Forsk.) Chiov. Verbena fruticosa Sessé & Moc. = Stachytarpheta incana Moldenke Nerbena gayana Kuntze = Junellia glauca (Gill. & Hook.) Moldenke Verbena glaberrima Gill. & Hook. = Dipyrena glaberrima (Gill. & Hook, ) Hook, Verbena glauca Gay = Junellia glauca (Gill. & Hook.) Moldenke Terbena glauca Gill, & Hook. = Junellia glauca (Gill. & Hook.) oldenk Verbena glauca var. cisandina Niederlein = Junellia glauca var. cisandina (Niederlein) Moldenke Verbena glauca g cisandina Niederlein = Junellia glauca var. cis- andina (Niederlein) Moldenke Verbena globifera L'Hér. = Lippia alba (Mill.) N. E. Br. Verbena globiflora L'Hér. = Lippia alba (Mill.) N. E. Br. “erbena globiflora Nocca = Phyla nodiflora var. reptans (H.B.K-) Oldenke 134 PHYTOLOGIA Vol. 8, no. 3 Verbena globulifera Auct. = Lippia alba (Mill.) N. E. Br. Verbena globulifera Spreng. = Lippia alba (Mill.) N. E. Br. Verbena globuliflora L'Hér. = Lippia alba (Mill.) N. E. Br. Verbena grandiflora Ort. = Priva grandiflora (Ort.) Moldenke Verbena gratissima Gill. & Hook. = Aloysia lycioides Cham. Verbena hastata x stricta Ravenel = Stylodon carneus (Medic .) oldenke Verbena heteraphilla Willd. = Hierobotana inflata (H.B.K.) Brig. Verbena heterophylla Willd. = Hierobotana inflata (H.B.K.) Briq. Verbena hirta Grah. = Lantana hirta Grah. Verbena hispida Sessé & Moc. = Priva lappulacea (L.) Pers. Verbena hyderabadensis Rottl. = Svensonia hyderobadensis (Walp.) Moldenke Verbena hyderobadensis Rottl. = Svensonia hyderobadensis (Walp .) M Verbena hyderobadensis var. maysorensis H. Wight = Svensonia hyderobadensis (Walp.) Moldenke Verbena hydrabadensis Rottl. = Svensonia hyderobadensis (Walp-) Moldenke Verbena hystrix Briq. = Junellia seriphioides (Gill. & Hook.) Mo 3 Verbena hystrix R. A. Phil. = Junellia hystrix (R- A. Phil.) Mol- denke C) Verbena illapelina R. A. Phil. = Junellia illapelina (R. A. Phil.) Moldenke Verbena imbricata Sessé & Moc. = Stachytarpheta mutabilis (Jacqe) Vahl Verbena inconcinna Brig. = Junellia ligustrina (Lag-) Moldenke Verbena indica Jacq. = Stachytarpheta angustifolia (Mil1.) Vahl Verbena indica L. = Stachytarpheta indica (L.) Vahl Verbena indica lanuginosa fl. rubente Barthol. = Manulea rubra Verbena inflata Humb. & Kunth = Hierobotana inflata (H.B.K-) Bria Verbena inflata H.B.K. = Hierobotana inflata (H.B.K.) Bria Verbena inflata Kunth = Hierobotana inflata (H.B.K.) Bria Verbena integerrima Larrafi. = Aloysia lycioides Cham. rbena intricata Briq. = Junellia intricata (Briq.) Moldenke Verbena jamaicensis L. = tachytarpheta jamaicensis (L.) V jamaicensis Vahl = Stachytarpheta jamaicensis (L.) Vahl Verbena jamaicensis Vell, = Stachytarpheta cayennensis (Le Ce Rich.) Vahl Verbena jamaicensis var. campestris Griseb. = Stachytarphet@ Fann tralis Moldenke Verbena javanica Burm. f. = Lippia javanica (Burm. f.) Spreng Verbena jancea Gill. & Hook. = Dicstea juncea (Gill. & Hooke) Miers er 1961 Moldenke, Monograph of Verbena 135 Verbena _ imei tormis Phil. = Junellia pseudo-juncea (C. Gay) Mol- oS Sage Junipertna Gill, & Hook. = Junellia juniperina (Lag.) Verbena py See Lage = Junellia juniperina (Lag.) Moldenke Verbena juniperina var. campestris Griseb. = Junellia juniperina (Lag.) oanerge Verbena juniperina var. clarenii Kurtz = Junellia juniperina (Lag. te: eel rae Verbena juniperina var. grisea I. M. Johnst. = Junellia juniper- dna - grisea (I. M. Johnst.) Moldenke Verbena lesiniate Sessé & Moc. = Bouchea prismatica var. lacinia- ae Grenz Verbena lacunosa Mart. = Stachytarpheta lacunosa Mart. Verbena laevigata Sessé & Moc. = Bouchea smatica var. lacinia- Verbena heed Willd, = Phyla nodiflora var. canescens (H.B.K.) enke 1d Verbena lancifolia Moench = Stachytarpheta indica (L.) Vahl Verbena lancifolia Steud. = Stachytarpheta indica (L.) Vahl Verbena lantanoides Willd, = _Lippia sibs alba (Mill. ) N. E. Br. ve Ve iapulacea Sessé & Moc. = Priva lappulacea (L.) Pers. Verbena lapullacea L. = Priva lappulacea (L.) Pers. “ene savanculactolte R. A. Phil. = Junellia lavandulaefolia - Phil.) Moldenk Verbena’ Mr folia se colchaguensis Reiche = Junellia lav ot adalan folia var. colchaguensis (R. A. Peep ) Moldenke Verbena lavandulifolia R. A. Phil. = Junellia lavandulaefolia (R. Verbena ligustrina Lag. = Junellia ligustrina (Lag.) Moldenke —_— ligustrina Lay. = Junellia a ligustrina (Lag.) Moldenke Verbena lindleyana Walp. = Buddleia lindleyana Fortune, Logania- ceae s Verbena lobelioides Grah. = Castelia cuneato-ovata — Verbena lobelioides Hort. = Castelia cuneato-ovata C erbena lorentzii Niederlein = Junellia ligustrina bad: ) Molden- Vertiens lorentzii Niederlein & Hieron, = Junellia ligustrina ( a) Moldenke 136 Purr OLogrs Vol. 8, no. 3 Verbena luppulacea L. = Priva lappulacea (L.) Pers. Verbena lupulina Cham. = Lippia lupulina Cham. Verbena lychnitis Mart. = Stachytarpheta lychnitis Mart. Verbena Maximiliani Mart. = Stachytarpheta lactea Schau. Verbena Reysorensts Wight = Svensonia hyderobadensis (Walp.) Mol- Verbena =e Sessé & Moc. = Priva mexicana ie ) Pers. Verbena mexicana L. = Priva mexicana L.) Pers Verbena mexicana trachelii - Gils tes fructu adie Dill. = Priva mexicana (L.) Pers. Verbena micrantha R. A. Phil. = Junellia micrantha (R. A. Phil.) Moldenke CU microphylla R. A. Phil. = Junellia minutifolia (R. A. Phil.) Moldenke Verbena minima Meyen = Junellia minima (Meyen) Moldenke Verbena minutifolia Lag. = Junellia minutifolia (R. A. Phil.) Men CO: oldenke Verbena minutifolia R. A. Phil. = Junellia minutifolia (Re Ae wee idenke Verbena morenonis Kuntze = Junellia patagonica (Speg.) Moldenke Verbena mulinoides Speg. = Junellia mulinoides (Speg.) Moldenke Verbena muscoides Phil, = Junellia mimtifolia (R. A. Phil. ) Mol- ~~ denke Verbena mutabilis Jacq. = Stachytarpheta mutabilis (Jacq.) Vahl Verbena mutabilis Willd. = Stachytarpheta mutabilis (Jacq.) Vahl rie oe Wight = Svensonia hyderobadensis (Walp.) Mol- Verbena nyssorensis Wight = Svensonia hyderobadensis (Walp.) Mol- ~~ d@enke re Verbena negundo Royle = Vitex negundo L. Verbena nervosa Scheele = Buchnera elongata Sw., Scrophulariaces’ Verbena nigrescens Mart. = Stachytarpheta glauca Cham. Verbena nispida Sess6 & Moc. = Priva lappulacea (L.) Pers. Verbena nodiflora C. Bauh, = Phyla nodiflora es ai Greene Verbena nodiflora L. = Phyla nodiflora (L. ora (L.) Gre Verbena nodiflora Petagn. = Phyla nodiflora (L. ) “tee eene Verbena nodiflora arborescens Mart. = Lippia javanica (Burm. f -) reng . verbena nodiflora var, sublanceolata Née = Phyla nodiflora tice ea (Ds Do Don) Moldenke werkiae te Gann Rich, = Phyla nodiflora (L.) Green mn Verbena aaa ena Poepp. = Junellia caespitosa (oil. & Hook.) Mo} eee We sess Verbena see} pt Speg. = Junellia spegazzinii Moldenke Verbena nudiflora L. = Phyla nodiflora (L.) Greene 1- Verbena ochreata Briq. = = Junellia s spathulata (Gill. & Hook.) Mo or gigantea 1961 Moldenke, Monograph of Verbena 137 Verbena odorata Desf. = Lippia javanica (Burm. f. m Spreng. Verbena odorata L'Hér, = Lippia alba (Mill.) N. re Verbena officinalis Cuevas = St: tachytarpheta jamicons (L.) Vahl Verbena officinalis L. x Veronica maritima L. = xVeronicena ~~ haartmani Moldenke Verbena orchioides Hort. = Castelia cuneato-ovata Cav. Verbena orchioides Walp. = Castelia cuneato-ovata Cav. Verbena orubica L. = Stachytarpheta orubica (L.) Vahl Verbena orubica, teucriifolio, primulae veris flore Hem, = Buiatog 3 IY: achytarpheta mutabilis (Jacq.) Vahl Verbena ourostachya Briq. = Junellia echegarayi (Hieron.) Molden- Verbena oxytetragona Mart. = Stachytarpheta gesnerioides var. cuneata Schau. Verbena patagonica Moldenke = Junellia patagonica (Speg.) Molden- ke Verbena patagonica Speg. = Junellia patagonica (Speg.) Moldenke Verbena pil Lippiane Kuntze = Junellia mimutifolia (R. A. Phil.) ee, fain serene Turcz. = Junellia selaginoides (Kunth) Mol- a ae ee enn Verbena polycnemoides Lag. = Junellia minima (Meyen) Moldenke ? Verbena prichardi Rendle = guneliia patagonica ( ee! ft ee Verbena prismatica Jacq. = Bouchea prismatica (L.) K Verbena prismatica a = “sowsien pr primatic (L.) Kuntze Verbena prismatica Willd. = Bouchea prismatica (L.) Kuntze Verbena See =a = ae a ea prismatica (L.) Kuntze Verbena ritchardi Rendle = Junellia patagonica (Speg.) Moldenke Verbena psamophila Mart. = Stachytarpheta canescens H.B.K Verbena pseudogervao A. St. Hil. = Bouchea fluminensis (Vell.) 2 oe 1 denke Verbena Pseudogervaé St.-Hil. = Bouchea fluminensis (Vell. ) Mol- e “ertena pectco~gervac A. St.-Hil. = Bouchea fluminensis (Vell.) Terbena poeuo=gervio St. Hil. = Bouchea fluminensis (Vell.) Mol- “ebena ecto tsnne Clos = Junellia pseudo-juncea (C. Gay) Mol- Verbena pseudo~juncea C, Gay = Junéllia pseudo-juncea (C. Gay) pene ee Verbena Punctulata Hieron, = Junellia punctulata Hieron, & Mol- Wevers hs nke Verbena - Fries = Junellia pygmaea (R. ree) Moldenke Verbena pag salen. = Phyla nodi. nodiflora (L.) Gree Verbena repens repens Larrafi, = Phyla nodiflora (L.) ROE 138 PHYTOLOGIA Vol. 8, no. 3 Verbena repens Savi = Phyla nodiflora (L.) Greene Verbena repens Ten. = Phyla nodiflora var. rosea (D. Don) Moldenke Verbena reptans Loisel,. = = Phyla nodiflora var. reptans (H.B.K.) Mo "Yoldenke Verbena rhinanthifolia Mart. & Gal. = Priva grandiflora (Ort.) enke Verbena rubiginosa Gill, = Acantholippia seriphioides (A. Gray) Moldenke Verbena rubiginosa Hook, = Acantholippia seriphioides (A. Gray) Mo. 2 Verbena salviasios Hook, & Arn. = Aloysia salviaefolia (Hook. & Arn.) Mold Verbena sanguinea ARES = Stachytarpheta sanguinea Mart. Verbena Sarmentoda Willd. = Phyla nodiflora (L.) Greene Verbena Scabra Muhl. = Phyla lanceolata (Michx.) Greene Verbena scirpea R. A. Phil. = Diostea juncea (Gill. & Hook.) Miers Verbena scoparia Gill. & Hook. = -= Diostea scoparia (Gill. & Hook.) Snes Verbena scoparia Hook, = Diostea scoparia (Gill. & Hook.) Mi yates ES a Hook. & Gill. = Diostea Diostea scoparia (Gill. & eck.) at ve Gill. & Hook. = Diostea scoparia (Gill. & Hook.) Miers Verbena selaginoides Kunth = Junellia selaginoides (Kunth) Mol- denke Verbena selaginoides var. illapelina Phil. = Junellia illapelin® (R.A. Phil.) enke Verbena selaginoides var. illapelina Reiche = Junellia illapelin® te As A. Phil.) ) Moldenke Verbena seriphioides Gill, & Hook, = Junellia seriphioides (Gill. ~ & Hook, ) Moldenke Verbena seriphiodes Gill. & Hook, = Junellia seriphioides Gill. & re eka) Mold enke a == (Speg.) Speg. = Junellia serpyllifolia (speg-) a & ° ee Verbena "serpyliifolia Speg. = Junellia serpyllifolia (Speg-) Mol serpyliitoits Verbena serpyllifolia (Speg.) Speg. = Junellia serpyllifolia (spegs) ~~ Woldenke ae Verbena silvestrii Speg. = Junellia silvestrii (Speg.) Moldenke Verbena spartioides Turcz. = Diostea cinerascens (Schau) "yoldenk Verbena spartoides Turcz. = Diostea | cinerascens (Schau.) Moldenke A Verbena spathulata Gill. = Junellia spathulata (Gill. & Hooks ) wo ~~ denke Verbena spathulata Gill. & Hook. = Junellia spathulata (Gill. & ~~ Hook.) Moldenke Verbena spatmlata Hook. = Junellia spatimlata (Gill. & Hooke) Mole 1961 Moldenke, Monograph of Verbena 139 denke Verbena spathulata var. ones es Schau. = Junellia spathulata ~~ (@ill. & Hook. ) Mold Verbena specs ta var. i EE Reiche = Junellia pseudo- cha sau ea (C. Gay) Moldenke Verbena oes, a grandiflora Schau. = Junellia spathulata +; hola & gs Moldenke Verbena spathulata parviflora Schau. = Junellia spathulata (Gill. & Hook.) Moldenke Verbena spicata jamaicana Pluk, = Bouchea prismatica (L.) Kuntze Verbena spissa Sandw. = Junellia spiss pissa (Sandw. ) Moldenke Verbena a Jacq. = - Elytraria a saute (Jacq.) Lindau, Acan- Verbena sten ahachiite Scheele = Becnnera lavandulacea Cham. & ~~ Schiec echt., Scrophulariaceas Verbena stoechadifolia Br. = =e stoechadifolia (L.) Small Verbena stoechadifolia L. = Phyla stoechadifolia (L.) Small Verbena struthiomm Speg. = Junellia struthionun (Speg.) Moldenke Verbena subfruticosa Aubl. = Phyla stoechadifolia (L.) Small Verbena succulentifolia Kuntze = Junellia succulentifolia (Kuntze) Moldenke Verbena suffruticosa Aubl. = Phyla stoechadifolia (L.) Small Verbena suffruticosa Steud, = Phyla stoechadifolia (L.) Small Verbena Supina Trag. = Bidens tripartita L., Carduaceae Verbena Verbena tetragonocalyx Troncoso = Junellia tetragonocalyx __ (Troncoso) Moldenke vonvene thymifolia Lag, = Junellia thymifolia (Lag.) Moldenke Verbena olia var, minutifolia Sao Speg. = Junellia oe tifolia (R. A. Phil.) Moldenk Verbena Liynoides Phil. = Junellia eimmtitotia (Re. A. Phil.) <= Meid denke Verbena Sstiaceuise Cham. = Lantana tiliaefolia een Li Verbena toninii Kuntze = Junellia ameghinoi (Speg. "Woldenke Verbena tridact; tridactyla Phil. = = Junallia tridact; tridactyla (R. A. Phil.) Mol- pce: nk — tridactylides Lag. = Junellia tridactylites (Lag.) Mol- e Tarbene tridactylites Lag. = Junellia tridactylites (Lg.) Mol- ” denke eer gesie unrest camer ees metic Jerbena tridactylitis Lag. = Junellia tridactylites (Lag.) Mol- ike ——SwweSS eee Verbena tridens Lag. = Junellia tridens (Lag.) Moldenke Verbena cee peme: = amelie std Ss (Lage) Moldenke 140 PET TOLD OGIA Vol. 8, no, 3 aceae Verbena sri furcate Re Ae Phil. = Junellia trifurcata (R. A. Phil.) ~~ Moldenk verbena tripartite Phil. = Junellia trifurcata (R. A. Phil. ) Mol- erbeon triphila Willd, = Aloysia triphylla (L' Hér.) Britton Verbena triphyla Willd. = Aloysia triphylla (L'Hér.) Britton Verbena triphylla L'Hér. = Aloysia triphylla (L' Hér. 2 bein Verbena tuberosa R. Graham = = Castelia cuneato-ovata Verbena tuberosa Sessé & Moc. = Priva er grandiflora foes ) Moldenke Verbena ulicina R. A. Phil. = Junellia aaiicand ulicina (Re A. Phil.) Mol- ae. pa Verbena uncinata Waterfall = Priva mexicana (Le) Per Verbena uniflora R. A. Phil. = Junellia uniflora past: ‘i Phil.) Molaenke Verbena uniflora var. glabriuscula Kuntze = Junellia uniflora var. ", glabriuscula (Kuntze) Moldenke Verbena valparaisos Lejeune = Phyla nodiflora var. rosea (D. Dor) Moldenke Verbena varsabilis Hort. = Stachytarpheta cayennensis (L. C. i ee VV Verbena rerbonaefoLio Michx. = Stylodon carneus (Medic.) Mol- Verbena aa eae L. = Stylodon carneus (Medic ») Moldenke Verbena violacea Walker = Stachytarpheta urticaefolia (Salisb.) Verbena a virgata Ruiz & Pav. = Aloysia virgata (Ruiz & Pav.) As le Varta witezsitt te = Junellia wilezekii (Briq.) Moldenke Verbena yamaicens s L. = Stachytarpheta jamaicensis (L.) Vahl "Vervain — is Salvia verbenacea L. in the Lamiaceae, "vervain mallow" is Malva alcea L. in the Malvaceae, and the ph iia manele ae is sais minima of Jamaica. The “lav- r verbena" of Andreas nger Taiotoera pher) in Life 28 te): 65 & re Mier: [color cnmteation’. April 10, 1950, from the Borrego Desert, San Diego County, Calif lifornia, is Abronia villosa S. Wats. in the taginaceae. The Hicken, Sert. And. references in the bibliography of this Seg often dated "1922", but should be 192 and m., Fl. South. U.S. (1860, 1865, 1872, 388i, 288%, "1889, 1898) references in the generic bibliography on p ve be are there erroneously written "Fl. Southeast. U. 3. and should a he Sharp, Clebsch, & Clebsch \9i\8, distributed as nverben& aw appears to to be a a species of of ‘composite; the Mrs. Neff ff Bakker s.n. [Herb. San Diego Soc. Nat. Hist. 12751], also so “distributed, is Teucrium occidentale A. Gray in the Lamiaceae, while Goodspeed <- 1961 Moldenke, Monograph of Verbena 1y1 s.n. [Ft. Lauderdale, Mch. 1930; Herb. Iowa State Coll. 136138] is Pycnothymus rigidus (Bart.) Small in the Lamiaceae, Shreve & Tinkham 9896 is Marilaunidium hispidum (A. Gray) Kuntze in the Hydrophyllaceae, F. C. Gates 2112) is Veronica officinalis L. in the Scropmlariaceae, Hitchcock & Stanford 6952 (Po--266725, Se-- 58765) is Schobera angiosperma (J. Murray) Britton in the Helio- tropiaceae, Rowell 2308 is Elytraria bromoides grst. in the Acan- thaceae, and Edw. Palmer 13 is something not verbenaceous. In various herbaria sheets of Mimulus moschatus Dougl. (Scrophulari- aceae), Hemizonia fasciculata T. & Ge (Carduaceae), and Conoclin- ium coelestinum (L.) DC. (Carduaceae) have been distributed as eee herbaria, from Tacora Peru, is an as yet unidentified species of Verbena, Raimondi (19f,3) cites his number Cajamarca, Peru, as "Verbena sp.", but these have not as yet been seen by me for further identification, The "Verbena sp." re- corded by H. S. Gentry in his "Rio Mayo Plants" (192) is actual- ly two species -- his no. 557 from Mesa Colorado is V. canescens H.B.K., while nos. 1522 and 1758 from Sierra Chacuro and no. 1923 from Cerro Guicorichi are V. pinetorum Moldenke. Pulver (1937) says that wire hairpins are useful in pegging down verbena stems, but he is here referring to the prostrate- branching cultivated species such as V. hybrida, V. tenera, V. tenuisecta, etc. Cheesman says that "Verbena spp." are common in gardens on Trinidad. Hoehne (1939) reports that plants of the Verbena and Mint Families are used in Brazil to make perfumes called "cambar4s" and herva cidreira", Von Hagen (1937) affirms that a Verbena species which the Jivaro Indians [Amerinds] call "yapau", when mixed with one of the Amaranthaceae » is efficaceous evers uador. Noack (1937) discusses the genetics and chromosome numbers in he genus, as do also Schnack & Covas in their various publica- tions. Junell (1934) discusses the gynoecium morphology. Rosengurtt Gurvich ( n near the point of Tybee Island, Savannah River, Georgia, no. V.19-755 (1). The genus is mentioned by L. R. Parodi (193) as n the climax formation in the Elionurus viridulus ee an t rri wong with Cyperus, Fimbristylis, Juncus, Nothoscordim, Spergularia ge Melochia, Eryngium, Buchnera, Wahlenbergia, Ver- Ronia, a, and Baccharis. 142 PET TO L:6:6:-74 Vol. 8, no. 3 Although we now recognize 363 taxonomic entities in the genus, Bentham in 1876 and Pearson as late as 1901 regarded the genus as comprising 0 80 species. Westcott (1950) lists as diseases attacking the native species in the United States: Plasmopara halstedii (downy mildew, in New Mexico), Ascochyta verbenae (leaf=spot, in Wisconsin), Cercospora verbenicola (leaf-spot, in Alabama, Louisiana, and Texas) and other species of Cercospora (in Illinois, Kansas, and Mississip- pi), Phyllosticta texensis (leaf-spot, in Texas), Septoria verb= enae (leaf-spot, from Vermont to Mississippi, Texas, and South Dakota), Erysiphe cichoracearum (powdery mildew, general distri- bution), Puccinia aristidae (rust, in Ohio [January] and Arizona [February & March] on grasses), and Puccinia vilfae (rust, in 0- Bank, winters on 4 Verbena species in the Paraguayan Chaco. Verbena leaves are said Ho 933) says that insect galls occur on the leaves of erbena species. Seymour ) reports that on species of thi genus may be found the fungi Rhizoctonia solani Kuhn and Sphaero" theca Iumuli. 182 (1917), describes a gall on Verbena, reared from the rolled leaves, caused by a probably predaceous itonid gall-fly, Leste him in Bull ne an irregular oval stem-gall 3--5 mm. long, caused by an itonid, Cecidomyia sp. (described by Felt in Journ. Econ. Ent. 4: 47h. 1911), the other, reared from marginal leaf rolls, being an itonid known as "Verbena leaf midge", Itonida verbenae Beutm. (described by Beutenmueller in Canad, Ent. 39: 306. 1907)- ABRAMSI Moldenke, Am, Midl. Nat. 2h: 750--751- 19,0. Synonymy: Verbena lasiostachys var. abramsii (Moldenke) Jepst™ par 3 ee 381. 1943. Verbena abramsii Moldenke ex Jepson, - Calif. 3 (2): 381, in syn., sphalm. 19]3. Literature: Perry, Ann. Mo. Bot. Gard. 20: 292 19335 woldeni™s Am. Mid], Nat. 2h: 750--751. 1940; Moldenke, Known Geogr roma. Cit. 1: 2, 3, 56, 72, 121, & 135. 19h6; Moldenke, Phytologia 2; 383. 1947; #11 & Salisb.’ Ind. Kew. Suppl. 10: 242. 1975 Molen ke, Alph. List Invalid Names Suppl. 1: 22 & 25. 19L73 Moldenke, 1961 Moldenke, Monograph of Verbena 143 Castanea 13: 113. 1948; H. N. & A. L. Moldenke, Pl. Life 2: 8. 1948; Moldenke, Alph. List Cit. 2: aed 439, 65, 467, 472, 482, 488, 495, 500, 560, 570, 576, & 587 (1948), 3: 718, 76S, 706 79, 799, 608, 832, & 8hl (19 949), and k: 980, 1119, TT, Jer 1182, 1215, 1225— 1050, & 1252. 199; Moldenke, Known Distrib. Terbenso., [ed 2), 27 & 197. 1949; Moldenke, gee a3; 959 6 Bushy perennial herb, often somewhat woody at the base, 5—7 dm. tall; stems erect or bran anching from the base and laxly pros- trate-ascending or prostrate, slender, tetragonal, usually thick- margined on the angles, den ensely s short-hirsute with browmish or canescent beire issuing at right angles to the stem, less so in age; branches similar to the stems, but even more dens and olny ts short—hirsute; leaves sessile, oblanceolate or obo- vate, 1.5--9.5 cm. long, 0.3 -5 cm, wide, usually small, dark- green on both surfaces, often brunnescent in drying, acute at the gular the cuneate base, sometimes the 2 lowermost divisions ypev rom irs and sc any-fl1 8 not se era bractlets ovate, about 2 mm. long, acute at the apex, densely hirsu: i pag ee erm flowers 3 calyx very small, 2—3 mm. long in all, densely hirsutulous, its rim very minutely toothed with very pager d obscure Saathe teeth; corolla purple or lavender to blue, a tube about mm nec Be its limb about 2.5 <_< in diameter. of this distinc a was — ted by Edward Palner (no. 309) at Hot Springs, in the southern part of San Di- ego County, California, in 1875, and is aapotitted in the Morong Herbarium of Barnard College at the New York Botanical Garden. The species has in the past been confused and united with V. iasiostachys Link, from which its small and mimutely toothed Collectors have found this plant on dry hillsides ny rc cover and desert slopes, on gentle slopes near ponds, earth, damp meadows, or damp land in general, meadows - oo nenthaefolia Benth., V. officinalis AA V. polystachya H.B.K., V. Prostrata R. Br., V. scabra Vahl, V. urticifolia L. found it "spontaneous in Botanical “Gexian trickle", Los Angeles County, se says that it is "The prevalent form “here (San Ber~ ne ty], meadows, etc." Twisselmann found it "growing in mud of whe seep in open pasture, scarce in dense growth of Mimulus Wh rer tOLioegra Vol. 8, no. 3 guttatus, Scirpus sp....in grassland of Upper Sonoran" zone in San Luis Obispo bispo County. The label on Blankinship s.n. [Sept. 23, age ~ohe tage ently an introduced weed" n 4210 is a mixture with V. lasiostachys, and H.210 ie Brandegee s.n. [Jolon, June 19, 1908] are mixtures with V. adiectacies var. -, septentrionalis Moldenke, while Abbott son. [Monterey, 1889] is a mixture with V. robusta Greene. “The K ee son. [Aug. 1905], at first determined as v. ortioinalal 7 wy G L. We. R [, Reinecke, was later re-determined by hin mas Ve menth- Cats Benth. He says that Miss Perry cites this collection as V. menthaefolia in her eMcpee of the genus in Ann. Mo. Bot. Gard. 20 (1933) » but she does 7 aoe Perry (1933) noted the differences: between this species and V. ens but did not give a e to the plant. She says ; that "The following specimens differ ean the a in oe scarce inflorescences ruiti ng ca 2.5-—3 mm. long shortly conniving beyond the fruit.." and rg cites five of the collections which I enumerate herein under V. abramsi, with the following 6 additional spec cr which I have not as yet seen: CALIFORNIA: San Bernardino Co.: S. B. Parish 2819 (E); Parish & Parish 969 (F, W). San Diego a Edw. Palmer 309 (E, G). Tulare Co.: Culbertson 210, in part (E). “She also cites Abrams 2481 from Los Angeles County, but I regard this as ve lasiostachys v var, scabrida Moldenke. In all, 62 herbarium specimens, guelnt ine the types of all the names inv nvolved, have been examined by m Citations: CALIFORNIA: Fresno Co.: x. "prandegee s.N. See [Fresno, July 21, 1905] (Ca—185315). Inyo Co.: Horn s.n. ane ns Valley and at Fort Tejon, 1863] (N). Lake Co.: Blankinship 5-n- oe 23, 1928] (Re—2766) . Los Angeles Co.: Abrams & yenregor oat a 2205, N); E. Be Babeoet s.n. [near Pomona, Aug. 2h, 1902 56930) 5 E. K. Balls 19393 (Au--122335); Braunton 520 oe T598 Du—78780), s.n. [Nov. 1902] (Du—-75595); J» B. Davy 2859 (Ca 25165); Dressler 69 (Ak-—-29781, Ak—2978h) ; ~tiningworth 74 (Po 4009); Munz 6919 (N, Po—-18355); Pruett 61 (Po--l1126); Re ', B. Rate dall 282 (Sd—h255); L. C. Wheeler 13 r 13 (La). Mariposa Co-: Congdon C.95 (N). Monterey Co.: E. = K. Abbott 1689 (Um—19), S-M+ semagee ey, 1889] (Mn--23135, N); K. Brandegee s.n. (Jolon, June 19, 1908] pegstraer sy H. L. Mason nis (Ca—912303); Simontacchi 020 628 [U- - Dept. Agr. Forest Serv. 23020] (Ca—12h18). Orange Co-? ay Te Brel LL8 (Rs--65l), 2806 ee teeny Riverside Co.: M. he King s S.n. n, (Menifee, 1893) (Ca—-75796); McClatchie s.n- {near El- sinore, 5/16/1892] (N). Sacramento Co.: Bruce 1977 | (Pon TAs San 0 Cost Se Bs Parish 2819 (Gb—30826, Po—H002, Sy Url 6295 (au), 1058 (Du--91160), s.n. [July 1887] (Car-2516), este TSept. 1887] (I), sn. [Aug. 1869] (Dt), son. [July 189b] (ca) 1961 Moldenke, Monograph of Verbena 145 sen. (6 June 1898] (N), son. [May 29, 1901] (Du—-1)3856); Parish & Parish 969 (Al, Br, Ca--192067, Du—91137, Io--92219); Pequeg- hat s.n. S.n. [Coldwater Canyon, 25, VIII.37] (La). San Diego Co.: G. LA s.n- [San Diego] (Ca~~104851); Edw. Palmer 309 309 (Be--type, Ca—187503--isotype, Pa—-isotype). San Joaquin Co.: Suksdorf 3d (P1--138),10). San Luis Obispo Co.: Eastwood 11984 (Gg—-1h7762); Twisselmann 1485 (Gg-~391013). Trinity Co.: £25 P. Johnson 1163 (Rs--1109). Tulare Co.: K. Brandegee s.n. (Aug. 1 1905] (Ca: (Can 185373); Culbertson 4210, in part (Po--63887), H.210, in part (Gg—31352). County undetermined: Parry & Lemmon 32 (C, Io). XVERBENA ADULTERINA Hausskn,. in Nyman, Consp. Fl. Suppl. 2 (1): he 1889 [not xv. eal penta Woldenke, 1955). Synonymy: Verbena officinalis x supina Haisskn. , Mittheil. Thttring. Bot. Ver., new ser., 10: 65. 1897. Vitex adcl.tarine Hausskn. ex A. W. Hill, Ind. Kew. Suppl. ay 210, = 210. 19 - Dist Verbenac., [ed. 1], hh & 102 fy9h2) "antl fed. 2 2], “108 “& 200. 15195 Moldenke, * Phytol og ia 5: 132. 1955; Moldenke, Biol. Abstr. 30: 1093, 1956; Moldenk , Am, Mid]. Nat. 59: 3hl--3h2, 3h, & 365. ot prune néenas 357. 1959; Moldenke, Résumé Suppl. 2: 6, ” a Se 15. This Big pa "proposed by Haussknecht in Nyman's 1889 work, ited 2 save: for the natural hybrid between V. officinalis L. and Vv. su upina L. which he found growing wild among the two parents in Greece. It is again discussed by him in the 1897 ref- erence listed above and is correctly listed by Durand in 1906. Known Geogr, Distrib. Verbenac., [ed. 1], hh & 102 (1942) a led. 2], 108 & 200 (1949). Hill credits the name to t Haussknecht reference, but in this reference = sancti ad Plainly Proposing a Verbena hybrid. There is no such thing as a “Vitex adulterina". Not kn knowing of Haussknecht's Verbena adul- terina at the time, in 1955 I proposed another "xVerbena erbena adulter- ina" for the hybrid between V. officinalis 1. and V. halei Small. The ska is now scat as xVe conata ri be discussed by me in my treatment of oe ess species. Because of this fact. ] pellined Haussknecht's © synonymy under officinalis in my Résumé in 1959. pee en cai v. officinalis and V. supina grow together in many parts of Burope and the Ori- 146 PHYTOLOGIA Vol. 8, no. 3 ent, where they are in most places the only representatives of the. genus. A hybrid between them is certainly to be expected. It dy named Myarieties" and "forms" of these two species may in actuality re- present the hybrid instead. Experience with natural hybrids tween other members of this genus tens xV. engelmannii, xV. ecentio xV. rydbergii, etc.) has shown that the hybrid popula- tion, when extensive , usually shows every intergradation between the two parent species. It is possible, therefore, that the par- ticular specimen of the type collection which I have seen repre- sents a plant that was on the V. officinalis extreme in its sup- i insane om n Haussknecht's original description is as follows: "inter par entes pr. Karditza Th[essalonica], — Im Allgemeinen besitzen diese hybriden Formen mehr das Aussehen der V. officinalis, un- terschéiden sich aber durch ktirzere gedrungenere Stengel, durch kleinere, feiner zertheilte Blutter, auch auffallend kttrzer ge~ en e stellten Blttthen, die nur in der unteren Htifte lockerer sind als bei V. supina; ferner durch breitere, etwas tiefer gespaltene Kelehzipfel und vor allem durch das die ganze Pflanze tfiberzieh- ende graue hispide Indument, welches jedoch schwicher ist ane bei zs Spi Die gu Th, anscheinend entwickelten Samen sind im eren Se Sweet, Brit. Flow. Gard. l [ser. 2, 1]: pl. 4. Synonymy : Verbena alata Otto ex Sweet, Hort. et ed. 2, 418. alata Link & Otto ex G. Don in Loud., Hort. Brit. — I: 602. 1832. Verbena alata Hook. ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syn. 1947. Verbena alata Cham. & Schlecht. 6 9. rature: Sweet, Brit, Flow. Gard. [ser. 2 1]: pl. le 1830; Sweet, Hort. Brit., ed. 2, 418. 1830; G. Don in Hort. , Bot. Gard. 5: pl. 98, 1834-1835; Walp., Repe 1845; Schau. in A. DC., Prodr. 11: sh3.° 18173 Schau. in Marte, » 9: 190. 1851; Briq. in Engl. & Prantl, N re 3a): 891; Jacks., Ind. Kew. 2: 1178. 1895; Stapf, Ind. : 428. 1931; Mol 2 own Geogr. Distrib. Ve oy Lome I, & 101. 192; Augusto, Fl. Rio Grande do Sul 211, 215, & 233 Rio fig. 100. 1946; Moldenke, Alph. List Invalid Nanes Suppl. 1: 22 nt Mold enke, Phytologia 2: 336. 197; Moldenke, Castanes 7 - 1948; Moldenke, Alph, List Cit. 2: 357 (29l8), 32 687, 14s aie, 922, "& 923 (19l9), and hs 12h, 1180, 1233, grt ee ig 5; oldenke, Known Geogr. Distrib. Ve rbenac., [ 93; “ & 197. "19193 Moldenke, Phytologia 33 "134 (1949) and 3: “bs. os Stellfeld, Trib. Farm. 19 (10): 166. 1951; pape ie : Roy. Hort. Soc. Dict. Gard. 2208-2210. 1951; Ramb , Sellen a nee 1961 Moldenke, Monograph of Verbena 147 153 & 165 (195k) and 7: 288. Rai, gamers prey: 109, 222, 357, & 470. 1959; Moldenke, Résum é suppl. es. o, & 15. 1960; ely, Fl. Paran, 16: 78 (1960) and is fe.’ sae Tllustrations: Sweet, Brit. fe ge rep ‘i [ser. 2, 1]: pl. Erect, suffrutescentt, rather robust perennial bush or subshrub, to 2m. tall, or suffrutescent only at the base, with a peculiar broom-like appearance; stems subfastigiate, very sharply ag= onal and broadly winged, brachiate; branches sharply tetragonal or the larger ones subalate, appearing as though leafless, the sides smooth and canalic culate, slightly striate, the angles some- times scabrous; nodes contracted; principal internodes 7.510 cm. long; leaves very small, sessile, mostly ovate or triangular- o§ inflorescence a terminal cymose panicle, subfastigiate, the spikes short and dense, at first ovoid and about 6 m. long, later sub- cylindric and elongating to 2.5 cm.; peduncles slender, sharply l- angled and canaliculate, Teel ign seers, bractlets lanceolate, calyx, its apex drooping; stigma declined; cocci a- bout 2mm. long, dark on the back, striate, the commissure angu- = arc. - Humnemann, who rece it from the Royal Botanical Garden at Berlin in 1628". He ade that "it is a native of Monte Video". Actually, I ha material of the species from Uruguay as yet, and the pines & mblish it validly. The species has been widely cultivated, being own from gardens in England and Germany as early as 1828, in 148 PRETOLOGIA Vol. 8, no, 3 Italy in 1849, and also in New Jersey. In its native haunts it is said to inhabit marshes, swamps, and pine woods, the edges s of swamps, rather marshy places, woods on varzea aan wet il 1 and rather open wet places in general, as pre and along streams in small woods, at asta of rey to ers. It has been collected in anthesis from September to in fruit in “winged-stem vervain® in England. According to Maund the plant will grow best ina rich light soil, attaining a height of 6 feet, becoming bold and handsome, although its flowers are not very conspicuous. It is not bushy and needs to be staked up if grown in the open. It may be raised from seeds or occas jonally increas- division of the roots. It needs a little spe kesiksha in win- ter if grown in areas where there are severe frosts The species is mentioned by Irm&o Augusto in his “wFlora do Rio Grande do Sul", pages 211 and 233 (1946), ~~ his illustration is mis-labelled "Verbena Lindmanii". He says that Emrich collected the species at Arroyo Tiririca, Caracol, Grande do Sul, but this specimen has not as yet been seen by me. Material of this species has been misidentified in herbaria as V. ephedroides Chan., V. litoralis L., V. litoralis Kunth, V. brasiliensis Vell, and even (by Dusén) as Neosparton ephedroides Ss Griseb. n all, 5 herbarium specimens, including type material of all the names ” invol » and 3 mounted photographs or illustrations Citations: ‘BRAZIL: Minas — F. C. Hoehne s.n. [Pouso Al- gre, May 2, 1927] (La, N, Sp—19358); Hosta én 968 (N, S); Netto 33h (Ja wT ses0, N); Regnell I 7326 [8/2/1865] (S), 1.326b (S)5 ss aes hoo (Lu, (lu, Ut—1158h), sen, [Brasilia] (N, S)- én 26 (S), 2788 (S), 3061 (S), 5061 [Herb. Mus. Nace Rio Jan, ars (N), 7541 (S), 10753 (S), 18047 (S)5 a 3X6 (Z), 35956 (Rb); Tessmann 3399 (Herb. Mus. Paran. ] (N)s 3400 (Herb. Mus. Paran. 3400] (N). Rio Grande do Sul: daw ses 411 [Herb. Hort. Bot. 1315] (N); Kleericoper 12 [Herb- Mus « Rio Jan. 18452] (N); Rambo aes part (N), 44083 (N), 497i? 9 (is 54735 (B). Santa Catarina: . Mtler s.n. [Herb. rb. Mus. Nac. ec. Rio Jan. 31564] (N); Reitz 1966 ¢ (x), ), 6831 (N); Reitz & Klein 5u8h (Ns Ok); Smith & Klein 8065 (Ok), 9059 (Ok); Ule e 1068 (W-- W—132 31h) « Sto Paulo: Brade 5737 (S); Leite 37h (El, N). State unde versie’ Sellow s.n, (Brasilia; Macbride photos 17399) ae 5p N, Vt). CULTIVATED: France: Herb. Martens s.n. [h. b. Pare 10 (Br). Italy: Herb, Harvey s.n. [h. P. “Ps Sept. 5 i8h9) (Du-—166469» part). New Jersey: Guyot s.n. (Pr). MOUNTED ILLUSTRATIONS: Maund, Bot. Gard. 5: pl. 98. 183)--1835 (N). VERBENA ALATA f. ALBA Moldenke, Phytol 26. 1951. Literature: Moldenke, Phytolo a at Br ghee 1951; Moldenke, 1961 Moldenke, Monograph of Verbena 149 een “Hors te 25: 3051. 1951; meen Résumé 109 & 470. 1959. s form differs from the typical form of the species in hav- a eee or whitish corolla The type of the form was co ollec ted by Anders Frederik Regnell (no. 1.326, in part) at Serra do Caldas, Minas Gerais , Brazil, on October 16, 1861, and is deposited in the Britton pesbabbin at the se York Botanical Garden. The same number, collected on February » 1865, appears to be typical V. alata Sweet. The white-flowered ria is known thus far only from Minas Gerais. In all, herbar- ium specimens, including the type, have been examined Citations: BRAZIL: Minas Gerais: Regnell 1.326 [16/10/1862] (N--type, S--isotype, S—isotype), 1.326 [2 [20/11/1867] (s). VERBENA AMB LIA Rydb. in Small, Fl. Southeast. U. S., ed. 1, 1011 & “1331. 1903. Synonymy: Verbena ambrosiaefolia Rydb. apud Wooton & Standl., Contrib, U. S. Nat. | ae Herb. 19: 549. 1915. Verbena ambrosiafolia Rydb. ex Moldenke, Suppl. List Invalid Nanes 7, in syn. 19 Verbena ambro siifo lia Rydb. ex Moldenke, Suppl. List Invalid. Names 7 7, in syn. 191; Rzedowski, Ciencia 15: 147. 1955. Verbena androsaemifolia Ricker, Kodachrane Slides of Wild Fis. (J, hypo nym. 1953. Verbena eabronio! ded Rydb. ex Moldenke, Résumé 357, in syn. 1959. Verbena ambrosifolia Perry ex Moldenke, Résumé 357, in syn. 1959. Verbena bipinnatifida x wrightii Wooton ex Moldenke, Résumé 358, in syn. 1959. Verbena bipinnatifolia var. latifolia Scholl ex Moldenke, Résumé 358, in syn. 1959. Verbena ambrosae- folia Rydb. ex Moldenke, Résumé Suppl. 2: 10, in syn. 1960. Ver- bena | bipinnatifida var. latifolia Scholl ex Moldenke, Résumé _ ata Southeast. U Se ed. i # 1011 & 1337- 1903; Prain, Ind. Kew. Suppl. 3: 187. 1908; small, Fl. Southeast. U. S., ed. 2, 1011. 1913; Wooton & Standl., Contrib. U. S. Nat. terb. 19: 549. 1915; Rydb., Fl. Prairies & Plains 678. 19325 Perry, Ann a Sh 6a 10: 39. 1945; Darlington & Janaki Ammal, Chromosome Atla D “ F ‘. eL 19465; Moldenke, Alph. List Cit. 1: h, 1h, 23, 3h, 99, 12%—129, » 15h, 158, 175, 182, 2h2, & 256. 196; Moldenke, mytaloeis 2: oe (2948) and 2: 327 & 328. "19h:73 gebaga Ye Healing Herbs 150 PHYTOLOGIA Vol. 8, no. 3 539, 549, 572, 59h, 618, & 8 (1948), 3: 656, 677, 679, 708, 713, ; 5 i, 787, , bo1, 805, . 199; Moldenke, Wrtsioats 3: 132. 1995 eee aa Ge- ogr. Distrib. Verbenac., [ed. 2], 16, 18, 20, 21, 23, 25, 26, 32, & 197. 1949; H. N. & A. L. Moldenke, Anal. Inst. Biol. Mex. 20: 12. 19L9; aks er, Kodacrome Slides of Wild Fis. [h]. 1953; C. M. Rogers, Rhodora 56: 207. 195k; Rzedowski, Ciencia 15: 17. 19553 Moldenke, Résumé 20, 22, 2h, 25, 28, 30, 31, 36, 357, 358, & 470. 1959; Moldenke, Résiné Suppl. “4.3 & 1 Annual with’ short taproot or perennial bushy herb, usually low growing and spreading in dense circles to 2m. in diameter; stems more or less diffusely branched from the ban mostly spreading to about 25 om., oacoang erect and 30 cm. tall or loosely de~ cumbent-ascending, somewhat hirsute; leaves - dewadeats epee short-petiolate, tne “blades 2-6 cm. long, bipinnatifid with the subulate and often ands nied cocci subcylindric, slightly broadened base, 2.5--3 mm. long, meee rg above, striate below, the commissural faces muricately scabrous; chromosome number: 2n = 30, The type of this species was collected by George Everett Os- terhout at Rocky Ford, Otery County, Colorado, © codons 8, 1900, and is deposited in the Britton Herbarium at the New Y Bo 2 cal Garden. The type of V. bipinnatifida var. latifolia is scholl University of Texas at Austin, while that of V. b bipinnatifida x wrightii is Wooton 642 from Dona Ana County, New Mexico, in the Britton Herbarium at the New York tee Garden. rs homonymous designation, V. ambrosifolia Perry, was used by Demaree on his no» 13335 from Cimarron County, Oklahoma ~~ Miss Perry says "An examination vi the type specimens of V enbrostfolis and V. wrightii reveals differences probably suffic- ent to suggest distinct but closely related fonaes Typically the sommes is characterized by long calyx-teeth and coarser pubes~ cence; whereas the latter has Pre ea eet, finer and some~ 1961 Moldenke, Monograph of Verbena 151 what viscid pubescence. On the other hand, a study of the aggre- gate specimens of the two shows a high variability in the combin- ation of these characters; moreover, their geographic ranges practically coincide. For these reasons, V. am — sifolia is main- ed as a species with very doubtful status. V. ambrosifolia is a fairly widespread ence, resembling V. canadensis (L.) Britton in its calyx-lobes and V. wrightii A. Gray in its leaves and glandular pubescence. It is sometimes difficult to distinguish from the latter species, as indicated by the psa from Miss Perry. Typical specimens of the two spe- cies are, however, very distinct. Intermediate material m may well be the ni Siri of natural hybridization, since hybridization seems to be an established characteristic of the genus a since the ranges of the two species seem to be practically identical. V. ambrosifolia has been found by collectors on Paes soil and deep nar canyons on igneous canyon slopes and dry spt land, along s r sides and moist streamsides, on prairies and high or open sandy Plains, banks, limestone hillsides, alluvial flats near arroyos, red and blue shale banks, grassy or alkali flats, bluffs and rocky canyon walls, dry ridges and mesas, dry hillsides and foot- hills, railroad embankments and shady slopes, canyon pine slopes “ digi breaks and plains, in arroyos, on steep shale 8, along roadside ditches and dry creek beds, and at the ine ere hills." Cory says it is "frequent on roadsides and shoulders of highways", Goodman found it i "loose soil near highways", Warnock Says it is Sooper pee along roadsides, infrequent", and porter re- i it "common along roadsides on the plains." rweeeralld found * growing with grass and Flourensia on the Verhalen clay, on cal- ha soil frog ditches, on gravelly calcareous soil along ek bed S, at the base of cliffs in yellow pine forests, in gyp- seous soil a level plains in Prosopis-Gutierrezia tpt ridges in fine possib siferous soil. H and sg Yelkerburgh pipet 4a eee thee selamets sone aha we ta t on dry xeric plateaus, while the Moores say it is Mul i Res get Sg gravel soil on highway embankments”, or found it "sparse on limestone", while Parks & Turner de- 152 PHYTOLOGIA Vol. 8, no. 3 scribe it as "infrequent on limestone soil along highways" and Webster found it "locally common around edge of intermittent pond? In Texas it grows in fields and prairies, on gras flats, deserts, and mesas, and along roadsides a Hudspeth and Presidio to Val Verde Counties , blooming and fruiting from March to December. I+ has been found at altitudes of 50 to 8500 feet. Rogers lists it with See ieee torreyi (Kunth) A. S. Hitche., Oryzopsis hymenoides (Roem. & Schult.) Ricker, and Chry- r s tha of this fact it enters Oklahoma on the prairie grassland of the plains the Mesa and on the level top where mature soils have developed and where climax vegetation is found under present climatic conditions The common name "moradilla" is recorded for this species, and Curtin reports that "If a Spanish New Mexican has pains in the back, he makes a plaster of pitch combined with powdered moradil moradil- la and places it on the affected part of his body. But if he “he has Therves' or rheumatism, he prepares a bath with moradilla leaves, and drinks a tea siadlarly made. Although a number of other dis- eases have been confused with leprosy, the ground dry leaves {of this plant] are rubbed on spots that are supposedly symptomatic of this diabolical malady." The Arséne 16555 and Tis28 collections cited below are labeled "PL. du | MEXIQUE", but were actually made in New Mexico in the U- nited States. Perry states that Mearns 2677 is "atypical". The M- E. Jones s.n. [Colorado Springs, May 1878] is also not typical —~ it was actually previously cited as V. bipinnatifida Nutt., but to this identification Weber objects! Ramaley 15322 was also pre~ viously cited as V. bipinnatifida; its glands are very sparse ~~ possibly it may represent ve Sabcromifo lia f. eglandulosa Perry: The Utah State Agricultural College 11391 specimen is not num bered on the collector's label, “put seems definitely to be 4 part of ue M. E. Jones 122 [sollection, and is so cited by me herein- afte . Holmgren hia vert vied this. Nelson tis glee on the sheet’ of Archibald A.255 seen by him that it probably represents a "new species". The Lundell & lundell 7056, previously cited as V. ambrosifolia, actually is V V. bipinnatifida. T e Hitchcock & Stanford 6782 > specimen in the herbarium of Or econ eee College seems definitely to be V. ambrosifolia -- not V. wrightii A- Gray as are the rest of the specimens of this collection in oth- densely canescent form of V. ambrosifolia. The W. A. Weber 4588 pried RT distributed as V. ambrosifolia, are actu~ ally V. bipinnatifida; Deaver 3133 and Ensor 7 [111] are V. cil iata Benth.; J. W. Gillespie espie 8503 and Pas hg & Hitchcock 1297 ay wri are 1- gosdaiagis wuiz oss while St Sturgis s.n. (May h, 1902] is V wrigntit Mapping by county done by Andrew R. Moldenke ie) i stribution of Verbena ambrosifolia in the United States Dee ett nmr! Figure 2. Di — eee J + ° ry ry 7 i : D 10 > | 7" I a) T z rT] 2 es 2 | 2 = A % 3 2 It 3 I 34 if a I x 7 H \ o * | } ’ | ! "\ b j 3 AAs | I oO a Co { ER? ; j I A { ° K | f A M A N y T 5 ] THE Wan i! N TOBA ) N : Hy ed } ry os 2 i | H owe j ae Che : } 7 . te f 9 © mam | ' a sug "9 Gammy 9 y ; } 1 My oa = bd uaeary T ees ee . . — 1 ang . — 5 2 : | as ee \ Pomoens mat mane = 5, 12 ° pre = Sanden tor = 5 Dancer | omneg on oer cro maa mEMrALE | sormmcay mount = | rower | cavaee Cn Pitey | Peng & bet y™ stsrmt males } MOUNTRAR soma e D LO eae Chicane Py war peace wusty | wae i AL o > “a > ‘BENSON aT Slama, wscon Fexaus cong TOU lees “ oee=—1 eu 2 Deeacn |e oy 7 ‘SHERBURNE é exTRet A oO any 4 ¢ co Ss mee 2 ee 5 2 me Se mak yp rT ’ a m 7 SHERIDAN — ae a SUTTE : saps sat P| . CHPPERA cron) Coal Fhe =, s — POTTER FAUU GRANT s 0 Eunoeg San MONT Ig ee CAMP EEL pad “7 = K NK oe ,; ad ie" crock | DUNN WARATION 90 | coer ot 2 Ps burg] PFs wD C7 = s Leg] h. © aan pe ee © Th cuh on 4 bl s ry WaNCOCK re Gia . ows TION % aoe * \ m Box fone ® x , he Fson [oawes = eum. 5 crecory san Trweccm [uncanny St forpeen | Cust fu oat os i ne rr Cool % conta ror eal fend Fy, me TY yi . =e RAN DAN —s CERRO 1 = —_ Cal “ Py e & Ny os CHERRY hie s . ona HOMME ms Ej 5 oe san |x Tan WORT] RACINE “4 ain oT 7 = ea _ on a7rk Rt) ETWATE, e 7 Som ‘ert a re I Se (tor 2 : ord Pure | cosHen BROWN war S; rent pesareed Ree =a foe beresel bcanal Fe : carton Sa 4 ae = a eT ina & Me cane oc \ es ¥ ars 4 fe ALBANY nox CEDAR A a = DELAWARE] ° oo te Seren | UKE ot =aet{ won yj vi ones tor - yj aS . {| a | mx OUT ed al wane BLACK nue 30 paness } 5 = oo some oan Fepancd “= A ar post! a mace ae UiNTa bad JANTELOPE) PIERCE cont] _ ae Tones * CoOK est" a Tg ence fn are cr i-¥ . Ja weeee WAYNE cunoun TOM Tha | BExTON | LINN ToRROAL | OLE ae BRNO oom oo iy ind ! at aa4e- SCOTTS BLUFF] won cant | nooner THAWFORG [CARROL [GREENE [BOONE | STORY JMARSHAL fer ihe ee mn toe che lay i i Le- |e ees AS nn J \ q vis{ Mote 2d . ol Fee] oMAS | Busine [tour [canrieno] wena waoisox [arom] Come FA WwW ): a ee - Ce De en coaaal (sna OAs on gas = | \F Sm ® ey namie BANNER ‘ARTHUR | McPHERSON bem oe Wuonison | SHELBY —piimmae TOMAS [POUR [ASPER [romtswcn| 1OWA 7 y | wa vr aE am sence Ex pe. aT bony fen 4 eo . e us JMB nan® " . SAT Lug ee * ew ey H IE B ARs A aay ec eee hace | seed TRISCATINE aA HENRY | BUREAU See oat forTuey} ALLEN gy nineons yt reas ont] Sp lone ws 4 ae : wt Py, eee we K s KIMBALL CHEYENNE STARK os r woul) aad tn war ocr Pedi ==" 9 me freed LINCOLN WK sspees WS FoTTAWATTAMIE | CASS | ADAIR | MADISON] WARREN } MARION WARASHA KEOKUK | *ASINS mck oI PULASA [PATON pa was | ast onan & aS, 4 ‘ef ray tour . SsweRman| wowano | ANCE poren | woe? LOUK we ERCER saa seal - ms Tesow ? ae ah 9 pee ee = a] wwermicn POX WHOL warsrann] [EMMSSTON Lr recoois want [ASS Lie a if ry opt . rr a ae —— Teas PwOnROE | WAPELLO ervensOn | HENRY TT eT iad ye bet 5 prensa od qn) on ot SS =] ba 2 pecegenest opto BaEIOR, BurFALG MALL Yor, | Sewanp |uncana cass wer yd meer sere anno. ay ee = a toe - mrt * *. " ral ti i" Tr) 3 BY LOGAN x mee q A =) PHILLIPS porn yon Faamont| Pace | tancon[rimacoub| DecaTUR | ware [arrenoost| DAVIS foun [UE fa ] bez TORO = ool me F pion Dee int cc carer AS wm a, cr) a 0 Buc CSE rm ae coma prs) at ee “hed cuasarnsok ros =. 7 AY fast GRAN CLAY | FILLMORE | SALINE * nigh ig vat ie ei, pecs M wire CaRon ] WASHINGTON J YOMA sont nODAMAY [WORTH [RARRISOML MERCER menace Oe ig we rAZEWELL Cuan . I FROIN) ro ea Coe OT | gon Ms . z oo L UND) ADAMS OE DONOUGH, LOGAN Tog wiT < rT’ ama an] eure =] = - \ oe ARE U = fn 1 Fowor = saan bon tee 008 | omen wasn on pont Ka ae a er > a > Of Sa i. : pEL Ww T ; cure rn ae | eon Pamnar oo GRUNDY vows Thus ENA at Ss Kramer anon am |e ; cncemt_|'* a Y “ee amoral cost mere sal om Uncooy A H fextan ce Losartan reetrt CHARDSOM' WROREW DAMIESS — . 3 oF a em or WARIO 4 aT onan CUMTON mS Fearon Sarvs er a * A ced eal * 4 er Tr} SUMMIT . “ it fe DE MALS —= Fr SHELBY [MARION acre Pe 2 = TROON ByTLER WIE gr fa am an m ~ ‘ws ; TOES WIG. sen 8y foun game te 7 soe ry é-\ omg we iw ctaney Prraay ELBERT urna MORGAN om mf Cc um LINCOLN RIT Cason SHRM CLINTON CHARITON mas sor z SHELBY we on H “ at Ears vont ng’ ane 2 . . THOMAS CLOUD ATCHISON pwcmanan| CARROLL GREENE (CHRISTIAN. ae OWEN ri 3 scx0TO pa ber fom ? aa wane I O DOUGLAS I] ron Gat RE crawatouie| OAS fr! MONROE “4 sonreontay cet (oc a ee mun yer i >» sworn . Le or or ey Or) fesent SS M ond . TA uNsom D Eo i [aed RUDRA _ FAYETTE. [errno ASPER eo ee hare ne ya os RX s OTTAWA BOONE - ul pore A . aon je ew O csaeur ecru oe ne S (7 unre at Rr CHEYENNE ences | Lo nee Sa H ons “an geonano knoe | =" pies papisonT 8° cor tect mapas | coh oe fot Pn eZ Saw TF. nose ER = m ~~ mane | to CUR, wasecson nl cunt aattt voy eneet? Cod faae coat Sa, * : 2 . CHAFFEE 4 GEARY Peasaunsct © wacxson Parris mann : } ae » — kee — Oe Co an roe = ~ = uno o aire = sf i Ouray SAGuACHE TSWORTH yonnson fi cass PSO — Fee ¥ me ee at xen es : et To - ° ! gt mc CROWLEY | Kiowa ed fede SALINE LYON owreau | re st, Lous N@ en re e Pp oeot pes et La N X\ a ASHINGTOn | MME —o HINSDALE CUSTER Ss S VENRY ] a ad came feat ih Cea \e by tus y/ . a a ae en Wa = ae WASHINGT ane Wate ‘\ Pe TEZUMA = ry ee | mean x fest ted ms aes Lon a ln ti A 2 Se SS, acts = r 2 aay . eco0wt rt nee ~ lessor — N 4 ‘a won Juan MePHERSON Cuicay) J ¢ - xp e : ». An Nou SS ad | Pare 80 GNOE F a HANEY ST cuaik | pexTon wer oar) ee fh = * at cad, ° MS Sgn ss] ee BS, ts : = . ! . GREENWOOD Froooson] ALLEN | BOURBON Mf VERNON HICKORY | CAMDEN ne antnest A Cervo & ety 7 OMD Cort ts a, sis, me me oe oy — * s 9 START — ceOAR DENT apa wi POPE or \ ron f eat . <¢ heat) fe hn iJ : jon | GRANT POLK = manson es — rr) . : Soon me. s SO NC ert v. | bed aan “ft Coneios f Cosma ENCAN WLson co - c een) Nee en wa we bat) ee oe ! say =e» oe ARF OROS BARTON} —— acue [rows g x a wnt wont wt, 4 ws Ln bs ort soto as i—-| a \) . XN STEVENS ae WRIGHT UMNO FI wr? ak) . % . . suet sso [Ww rl ] worron | 5 mn | comer GREENE aN Scere e lp? fomstan oo bog AE, Corie aaa ise: . AN = 08 Mery ‘. ) me Sarma} | YMEROREE AS Tarn : Scot ee eet og Ses) Lm en 8 OS FT is $A Pe ons ome \ ° nt . facsa- cy oy re , scott cusoret vo mo) nee ome oe at woe ht ‘ > a CHRISTIAN = [oouaias HOWELL rm aca = . 7 keg srs . vs \ TEXAS a NEWTON carver [BuTLER mae s 8 epeEuncr] woe cw e=) Fay a ‘Gace | 4g pave ROO Fearn i Neg i ° { bs ALFALFA | GRANT kar (OSAGE Bpnowaral cease | oTrawal oar OREGON = ened fees) ae, Ses ot : am 9 “ Ey ne Z zn, fe we 0 \ t ; TwNEY [OZARK a ie —= cS ee oar Aaah * v St, S) Jae o> ae a * . 0808 — WILSON sant pTmaad COCA acs, = ~~ sown | moot wae | . McKomury MOA LAM wrung s . © mmm ELS | wooowand GARFIELD NOBLE * Troctes anny s ny , iz vet nf © Deeg E DEA SMITH ee CTT) mi arr 5 notin) mS eon} msTONE J) gt Aone . , Clow Ww ‘ARMSTRONG | DONLEY — HUGHES jwoopruFt| ST, FRANCIS. . . TORN sat, ‘ ac wane wort y s Greer S yoowa rorta= PITTSBURG pl YE O77 dente DE SOTO. I =< Ce Coty an iN ho suwTt ben y | curey ue arcane LATIMER . Te MARSHAL a i couent nd ose pele ol ee naervns eneags J ANP T meron Q cf j i PARMER — | CASTRO SWISHER | BRISCOE HALL oa H SALINE PRAIRIE] —= H TATE oan FRANKLIN oa Part) << ee cS “pam SF piecing A 2 TACKSOM one Pe Feast er’ a > > STEPHENS PUSHMATARA POLK i RANA} : if or] COLMAN coun de eal ent ae oes ee are FERSON | ARKAN: ae moosevent W paxey [uae ‘ MURRAY McCURTAIN COME age 2 S Coes sxnatt m * toe ° Stim e ~! ma flow [worry [come] Macau] pein [: 1 PIKE woe 7] oUNT ks ora hee KEYED TO COLUMN A ae — i onusm WORROE wane i. ‘Saunt YY CALHOUR we re | b, imac ned 7 aM x ieee entre one on EN ast gpm of Cleartype Index Book R a wewta ~ F cuy a MARSHAL —"BRYAN | CHOCTAW CLEVELAND p! LINCOLN eines master ux = ret) yar, ont mos SC COLLETON comes \ . , oN town |wooaty fussoex —Tevossr [nme Pane —Tarop So f.. e a SA tie H aos RaW 3 ee a a a a w ie R ARCHER MONTAGUE UMAR WEVADA ] QUACHITA (HCHENS, Mite [rat cooKe GRAYSON Qe ume ence ou BOLIVAR uervonel TUSCALOOSA nitro won fy, J 2 bom A | ane -< CALHOUN] ’ ‘ow aes sae Hest cm \ er earn = 4 ee — band BT? Ue Ra [KENT Delt . ITM ‘pes Consh ) ° x fe pe ee te 9 Scale of Miles 4 7 Leh rd Fi —S—S——— LANAI ? STARR HIDALGO ’ ®, re > ) 4 WaLAcY rex 2 * rons aot ul ~_, = 2 io LISIANSHI, — “~s : ° ’ bea) : ] . 8 é¢ . ry eo oe) a oC Os, ALEUTIAN ISLANDS “ . > oF. oe 2 THIRD JUDICIAL DIVISION NECKER wt Pa . oe MIHOA “KAUAI ° ad a —s wine Fy Cored Scale of Miles Pte 0 = geen” hae * Joan? ° Copyright, American Map Co., Inc., New York, No. 14324 Se, ) ee é~ _ ee, KAUN, Sf” a) HONOLULU PY . Pa 7 f SEE SST Se ry Te Soe u 3 26 27 23 Fr) er ee eS | I 3 T 7 T 5 T ; I 7 a ee 7 T 7) T 7] T 7] T is T 17 1 18 19 T 20 I 21 I 2 ] 2 | L it | i T EJ 31 f 3 I x | 3 | s PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- ections have been deducted, will be distributed among the contributors upon : the completion of a volume, in proportion to the space which they have used. Fach contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. Each number consists of not less than 32 pages. (ll manuscript accepted will be published in the next issue, so that the size of numbers may vary greatly. A volume will contain about 32 onder as 512 pages, or a smaller num of pages with an equivalent number of plates. This plan insures se immediate publication of all accepted sianiectiok ad Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provided they conform to certain lim itations of size and proportion. An extra charge will be made for half- tones, dep ing on their size, as fixed by the engraver, with a minimum of about $2.25. Articles dealing with research in all lines of botany, in any reasonable length, Biographical sketches, and critical reviews and summaries of liter- ature will be saaibtoniect os publication. Floristic lists, casual notes of an — amateur or so-called popular type, and polemics will not be published. Advice _ on the suitability poss manuscripts will be solicited, if necessary, from quali-— fied botanists. = Under the present cost of printing, the basic rate for a page or fraction thereof is approximately $3.00 for an edition of 250 copies. This price is subject — to change without > since it depends entirely on the prices prevailing Z in the printing in ustry. os Reprints will be furnished at cost. A proportionate fraction of the edition of 250 copies is also furnished gratis to contributors. wes Upon request, the editors will send detailed instructions concerning the Pecperatien of manuscript or further information about the ee In- quiries ma aditremed io ‘the magazine or to either editor. they TOLOGT. Designed to expedite botanical publication Vol. 8 January, 1962 No. 4 CONTENTS : LAUGHLIN, K., Quercus xmegaleia Laughlin, Hibrida nova.... eeeee 153 ; MOLDENKE, H. N., Notes on new and wolewecthy slatta: XXX. ae . 159 WURDACK, J. J., Certamen Melastomataceis VI] .....-...0.05- vee 165 MOLDENKE, H. N., Materials toward a monograph of the genus Se Verbena, Il 7 CESONRES CHR64 «PEe Aha ewe abet ey smnens ee eee 6 175 Published by! Harold N. Moldenke and Alma 1. Moenke QUERCUS XMEGALEIA LAUGHLIN Hibrida nova Quercus lyrata Walt. X macrocarpa Michx. Kendall Laughlin (1890- ) 165 Pine Ave., Chicago, Ill. upper portion nearly smooth and interrupted by narrow vertical cracks. Branchlets glabrate at the end of the season, led, white lenticels. Terminal winter buds 4 mm. long, mm. wide, ovoid, broadly rounded at the apex, light brown, puberulous. Leaf blades 10-20 cm. long, 4.5-13 cm. wide, firm, dull moderate yellow green on both sides, gla- brous above, soft-pubescent beneath with a tomentose Petioles .4-1.5 cm. long, slightly angled, pubes- cent or nearly glabrous late in the season. d castaneous, puberulous, short-oblong, rather peice impressed at the apex, which is eccentric in a ation to the rim of the cup, about two-thirds en- osed in the cup; peduncle 11-22 mm. long. 153 15h PHYTOLOGIA Vol. 8, no. QUERCUS XMEGALEIA Laughlin Arbor 1.09 m. diametro et 29 m. alta cum gracili- bus extendentibus ramis 2 m. altioribusque super basim et pinguibus ascendentibus membris. Cortex pallidior cinereus et tenuior; inferior leviter fastigatus, su- perior fere laevis et angustis ad perpendiculum direc- tis rimis interruptus. Ramuli temporis fine glabrati, angulati, cinerei lutei brunnei numerosis albis lenti- cellis. Extremae hiemales gemmae 4 mm. longae, 3 mm. latae, ovoidae, in apice late rotundata, pallidae brunneae puberulentes. ia 10-20 cm. longa, 45-13 cm. lata, firma, in DISCUSSION newly discovered hybrid of the Overcup Oak This lyrata Walt. and the Bur Oak macrocarpa Michx., which h feet 7 inches 1962 Laughlin, Quercus xmegaleia 155 huses are of about the same width. In one rare form the leaves resemble the Post Oak. The leaves of mega- n its general appearance megaleia shows a strong su- perficial resemblance to the White Oak. The White Oak t found in this park and occurs but sparingly Megaleia's acorns are odd. The awnless cups are wider than the average of lyrata. Both ends are im- ing smaller species in the north central states., Both ted in large numbers and reach prodigious size- in iquoria, macrocarpa having acquired exceptional tol- france in competition with large Southern species in these bottoms. The holotype will be deposited in the Royal Bo- deposited in the U.S. National Herbarium, Washington, —:\:, by way of the U.S. Forest Service. One and one-half miles from the Mississippi Riv- er; dgscribed in PHYTOLOGIA 4:154-155. he maps in Figures 18 and 19 on pages 14 and 15 ce, U.S. De- partment of Agriculture, Revised September 1959, are very instructive in showing the complementary (vicari- ous) ranges of these two species. Vol. 8, no. } PBI 20:LO06 TA 156 THE TYPE TREE OF QUERCUS XMEGALETIA ings the foreground . Seedl in 1962 Laughlin, Quercus xmegaleia 157 FOLIAGE OF QUERCUS XMEGALEIA 21% of natural size 158 PHYTOLOGIA Vol. 8, no. Longitudinal Section of an Acorn ACORNS OF QUERCUS XMEGALEIA One-third of natural size J 1962 Laughlin, Quercus xmegaleia 159 35 See the table of dimensions on page 376 of Vol- ume 6 of PHYTOLOGIA. 4MIQUORIA" is a name coined by the author to take the place of "Big Oak Tree State Park," being brief, euphonious and descriptive. It is made u up as follows: MI from Mississippi Co. and River; QU from Quercus; ORIA from Hicoria NOTES ON NEW AND NOTEWORTHY PLANTS. XXX Harold N. Moldenke igs eager Fagg ep ad var. BARBATUM Moldenke, ri a forma typica speciei calycibus Aateé patente- que “hispido-barbatis recedit. This variety differs from the typical form of the species in having the calyx densely hispid-barbate with white hairs spread- ing at right angles to the calyx. The type of tis “vas variety was collected by Karl Rudolf Afzelius at Ay isc in the province of Tulear, Madagascar, on January 10, 1913, and is deposited in the herbarium of the Naturhistoriska Riksmuseum at Stockholm, CLERODENDRUM SUMATRANUM var. FAIRCHILDI Moldenke, var. nov. Haec varietas a forma typica speciei petiolis laminisque glab- ris fe es iety differs from the typical form of the species in havitie its petioles and capiavners peared e type of the variety was collected by David Fairchild (no. 488) — in whose honor mee is named -- “rbnugh first discovered oF James a Dasestts near the trail from eae to the Turpentine Still n Take engon » Sumatra, on March 5, » and is deposited in the harkérisa of the University of atiPene at Berkeley. yo? ERIOCAULON MINUSCULUM Moldenke, sp. nov Herba annua parvissima 1--2.3 cm. alta ubique glaberrima; fol- iis linearibus filiformibus 7--13 mm. longis; pedunculis solitar- Ld t+ 1 om apex, the lobes erect; heads solitary, black, 1—-3 mm, wide, Owered, glabrous; involucral bractlets erekaay obovate, shiie” 160 PHYTOLOGIA Vol. 8, no. stramineous or brunnescent, about 1 mm. long and wide, rounded at the apex, completely glabrous, shiny. ype of this very distinctive species for the region was collected by Harry Smith (no. 12123) in a swamp at Sunglingku, at about Vad meters aes ha between Taining (Ngata) and Taofu eptember 13, 1934, and is deposited in ty r-heads on the type collection. The species cannot be mistaken for any other in the region, LACHNOCAULON ANCEPS f. GLABRESCENS Moldenke, f. n forma a forma typica speciei pedunculis sidlieie vel glab- rescentibus recedit. s form rs from the typical form of the oR - hav- ing its peduncles either glabrous from the beginn paride e be- coming glabrous, subglabrous, or glabrescent in The type of the form was collected by Samuel Malis Tracy (no. 7586) at Braidentown (now called Bradenton), Manatee County, Florida, on July 23, 1901, and is deposited in the herbarium of the New York Botanical Garden - It bs originally distributed as L. digynum Holn, later re~determin d by Dr. J. K. Small as Le anceps eps (Walt. ) Morong with a aaeatine There is a possibility that it may represent a natural hybrid between L. anceps and L. glabrum Ktrn. pEerser LANTANA CAMARA var. TERNATA Molde nke, var. nov Haec varietas a forma typica speciei foliis ternatis laminis cmneieeryis hirsutulis supra scabris subtus resinoso-glandulosis recedit This variety differs from the typical form of the giaaty: in having its leaves ternate and the blades more densely hirsutulous on both surfaces » scabrous above, and olor pa pe Sane The type of the variety was collected by Otto Bnery Jennings (no. s8) 0 on the beach at Siguanea City, Isle of Pines, Cuba, on May | ee 1910, and is deposited in the Britton Herbarium at the peat York Botanical Garden. The collector states that it is a low LANTANA HIRTA f. TERNATA Mo idenke, f. aec forma a forma typiea speciei foliis ternatis plus elonga- tis recedit This form differs fron hg typical fom of the species in hav- ing ternate leaves, at least on the main stems, and in wing the leaf-blades thinner in arta, more elongate, "lanceolate, 3--5 45 cm. long, and 1.5--2.5 cm. wide. The type of the form was collected by Cornelius Herman Mull (no. 2686) on openly wooded or unwooded banks near Monterrey, in the municipality of Cafion Diente, Nuevo Leon, Mexico, on December 8, 1939, and is deposited in the herbarium of the University of California at Berkeley. The collector notes that it is an abund- 1962 Moldenke, New and Noteworthy Plants 161 ant shrub 2 to jj feet tall, with orange-yellow corollas. ee sernggeo “tase sp. n ramulisque ears obtuse tetragonis sulcatis; folie. pieietaas petiolis ca, lcm. longis glabris; laminis foli- orum membranaceis ovatis utrinque glabris, ad sepa. acutis, ad basin longe acuminatis; inflorescentiis quam foliis brevioribus; meee perconspicuis. foliaceis hrub; stems and piteeore apparently glabrous throughout, ob- Sites tetragonal, sulcate in stramineous or brownish h; nodes somewhat annulate; ata ipal internodes --7.5 cm. long; leaves mostly ternate; 2 ag slender, about 1 cm. long, gla- ha rous; leaf-blades membran somewhat lighter green beneath, ovate, m. lo Sack cm, wide, rather coars e fron slightly below the widest part apex, acute at the a- pex, long-acuminate at the base, rous on both surfaces, eu flowered, hemispheric in anthesi s, later somewhat elongate an and loose; bractlets very numerous and conspicuous, foliaceous, --7 mm. long, 1--2 mm. wide » very minutely strigillose, irregularly spreading in fruit; corolla aa ones 4--5 mm. long, very minutely strigillose on the outsi The type of this species was aniseiesd by ees Merrill King (no. 1775) -- in whose honor it is named — in clay loam in o- pen sunshine alo ong the Pan-American Highway vate 190) 2 km. South of the village of Niltepec, altitude 50 meters or less, on tep is deposited in the herbarium of the University of Texas Austin. The collector states that the plant was common oe ee var. SUBGLABRESCENS Moldenke, var’. ari a forma typica speciei foliis najoribus ‘subtus pilbplatessoe tins recedi This variety differs from the typical form of the species in having its leaves larger, to ) cm. long and 2 cm. wide when ma- ture and then subglabrescent beneath. The type of the variety was collected by Peggy E. Ellis (no. 23h) southwest of El Rago, at 6°35! N., Wh 3' E., in the Ogaden, bu describes it as a woody herb, with weak chia 3 feet tall, Gadd often growing under shrubs, the leaves very rough, and the °wers cream and pale-mauve in color. LANTANA YELUTINA t. VIOLACEA Moldenke, f. nov os ikea a forma typica speciei corollis “violaceis recedit. rm differs from the typical form of the species in hav- ing tt its aity light-violet or light-purple in color, with a 162 PHYTOLOGIA Vol. 8, no. yellow center. The Ne of the form was collected by Thelton W. Rk: and Chester M. Rowell, Jr. (no. 3481) on rocky soil in a ravine, at an crested of 5800 feet, L miles =. of Chilpanc ee reeset sited of Mic Pp age lorescences is exactly of L. montevidens (Spreng.) Briq., but the plant nd doseribed ty the collec coi as a tree 6 feet tall! ss vag ARGYRODERMA var. BREVIPES Moldenke, var. nov ¢ varietas a forma typica Sotbiet peduneulis reat em. long- This variety differs from the typical form of the species in having its peduncles only --5 cm. long during anthesis. The type of the variety was collected by Auguste Frangois Marie Glaziou (no. 9000) on the campos i Itatiaia near a little house called "Bueno el", Minas Gerais, Brazil, on November 20, 1876, and is deposited in the herbarium a the Muséum National d'His toire Naturelle at Paris. LEIOTHRIX FLUMINENSIS var. PUBERULA Moldenke, var. nov Haec varietas a forma typica speciei foliis ubique minute puberulis rec This variety differs from the typical form of the species in having the leaves minutely puberulent throughout. The type of the variety was collected by Jean ane Antoine among Gaylussacia plants at Restinga de Tocaia, Brazil, si is deposit- - in the fleet mid-g of the Muséum National d'Histoire Naturelle t Paris PAEPALANTHUS BALANSAE var. DENSIFLORUS Moldenke, var. n aec varietas a einen typica speciei capitulis panei ehoree ora eae 1 cm. diam. t. his variety differs from the Weottet fom of the species in having its heads very densely many-flowered, completely globose and 1 cm. in diameter during anthesis, not compressed in drying. dos, P Brazil, in October, 1958, and is deposited in the He N. Moldenke Herbarium at Yonkers, New York. CUMINGIANA f. CORDATA SCaarest f£, nov. Haec forma a forma typical speciei laminis foliorum ad basin valde cordatis rec This variety differs from the typical form of the species in oe its leaf-blades conspicuously and deeply cordate at the ase. The type of the variety was collected by Mary Knapp Clenens on Mount Apo, province of Davao, Mindanao, Philippine Islands, in 1962 Moldenke, New and Noteworthy Plants 163 June, 192), and is no. 15630 in the herbarium of the Philippine Bureau of Science, rae een in the Britton Herbarium at the New York Botanical G. PREMNA INTERRUPTA var. SMITINANDI Moldenke, var. nov. Haec varietas a forma typical speciei laminis foliorum supra velutinis subtus dense velutino-tomentosis recedit. eneath, ype of the variety was collected by Tem Smitinand -—- in whose honor it is named -—— and I. Alsterlund (no. 6783) on a rocky ridge at Doi Chiengdao, at an altutude of 1600 to 1800 m., Chiengmai, northern Thailand, on April 26, 1960, and is deposited in the H. N. Moldenke Herbarium at Yonkers, New York. The plant is described as a common tree, 8--10 m . tall, wi whiti flowers, PYRAMIDATA var. VITIENSIS Moldenke, var. no Haec varietas a forma typica speciei laminis raf han supra glabris subtus leviter puberulis rit ie is variety differs from the typical form of the species in having its leaf-blades glabrous on and merely lightly puberu- lous beneath, The type of the variety was collected by John — Gillespie (no. 3400) in woods in the vicinity of Nasim, 9 miles from Suva, at an altitude of 150 meters, in the province of Naitasiri, Viti Leva vu, Fiji Islands, on Oct tober 20 » 1927, and is deposited in the herbarium of the University of gal rg at Berkeley. PREMNA SUBSCANDENS var. GLABRESCENS Moldenke, var. nov. Haec varietas a forma typica speciei laminis foliorum glab- is recedtt, vel in venis principalibus parce breviterque pilosu- re y differs from the typical form of the species in having oc a. blades glabrescent on both surfaces, mostly only , Iuzon, Philippine Islands, on February 25, 1926, and is deposited res the herbarium of the University of rke PREMNA WOODI Moldenke, sp. n Frutex scandens; ramulis praciiibes ip ag foliis * Sea brevipetiolatis; laminis folio irme chartaceis bu Woody vine. to 35 ew i6 v9 eter; ° 2 ngs yl to FS CM. - am ri branchlets very slender, the young er parts densely yellow- or 164, PHYTOLOGIA Vol. 8, no. fulvous-hispid or -hirsute; principal oe ~_— 5 cm. long; leaves decussate-opposite; petioles short, slen ——6 long, densely yellow- or fulvous-hirsute; ictae fi rly chartaceous, brunnescent or nigrescent in drying, lighter beneath, shiny above, oblong-oval, widest = or ier encore paecd the middle, fe) acuminate pilose — most of the venation beneath with fulvous hairs; midrib slightly impressed above in drying, prominulous beneath; SeceAdiMees: “slender, about 5 per side, ascending, not emg arcu- ate, slightly impressed above in drying ; prominulous ben veinlet reticulation rear ba above, rather conspicwous beneath; inflorescenc terminal, flowered, small, and wide; tataiie slender, in, Tm» long , dens neely tulvous hirsute; pedicels slender, 1—2 m., pee rather densely s pubescent or puberulent; Pata linear, minute; calyx c pi late, nigrescent, about 1.5 mm. long and anaes irregularly labi- ate, densely puberulous; corolla~tube about mm, long, the limb about 3 mm, wide, The type of this species was collected by A. Mail (no. 12) at Teng meh | British North Borneo, on August 27, 1926, distributed as D. D. W 31 -- in whose honor it is named —- and is de- poakies in the herbarium of the University of California at er’ ‘ PAEPALANTHUS CHASEAE Moldenke, sp. nov Herba perennis parva; caule ut videtur radicante subterraneo lignoso crasso toto Tonet tuaivie radicifero; radicibus p coma ke brevibus; foliis graminoideis 1—-2.5 cm. longis , 1--1.5 mm ilibus 2~-5 cm. longis ‘ ecta tie tortis wee bris stramineis tured | capitulis hemisphaericis dense multifloris albis 5--7 Im, s,. hortatelat is just beneat the surface o e bab sol bearing a large number short roots on its soos bag ae roots straight, unbranched, usually extending directly 3 leaves numerous, ical, grass-like, uniformly bright-green on both surfaces, t ewhat spreading, 1--2.5 cm. long, de, acute and subulate at the apex, entire, apparently clebrees and shiny on both surfaces, but ly ing a ve ny ac ely scattered, long, whitish hairs; sheath 1 cm. long, clekreas or sparcely pil ose; oo bractlets in many series, overlapping, nigrescent, puberu e, Agnes Chase 10358 in pote t mixomn let near open summit of Serra de San Antionio, and Wy tn “eg , Minas Gerais, Brazil, December 27-30, 1929, deposited. in herb. University pes wichig an. CERTAMEN MELASTOMATACEIS VII. John J, Wurdack Dept. of Botany, U. S. National Museum CHAETOLEPIS SANTAMARTENSIS Wurdac k, SD. In indumento se heed C, micro yilee pool! dies distanter affinis, cum foliis amp us 5-ne 8 vel breviter “S-plinervatis, floribus anplioribus, petalis "feerlb. mata leviter barbellata, Rami cum petiolis lami- — — subtus dense strigosi vel longo-strigu- na 8-18 X 5. rir modice aac tanar? subtus in superficie modice setulos Flo ramalor apices 1-3-aggregati, cum bracteis foltacsis es involnerat bracteae ad 8 mn longae ovato-oblongae, supra apic tul Petala 9 X 8 mm obovata asymmetrice truncata ciliola Filamenta Palas mn paws antherae 3-3.5 mm longae oblongae apice mimate uniporosae, connectivo non prolongato sed dor- Saliter leviter stevetst Stylus 8-9 mm longus; stigma puncti- forme; ovarium -loculare apice dense setulosu ection: J. Cuatrecasas & R. Romero Castaneda 24669 (holotype US 2325392), collected in subparamos at "hoya del Rig Do so pte da Santa Marta, Dept. Magdalena, Colombia, elev. 3070- 3100 m, Oct, 9, 1959. "Tree 2m, Leaves yellowish dark-green sore, yellowish beneath, Petals lilac-violet, Anthers yel- basal connective @ppendage; both of these more-distant rela- ves are otherwise quite different. 165 166 PHYTOLOGIA Vol. 8, no. } CHAE TOLEPIS PERIJENSIS Wurdack, sp. omnibus ae non Suis deaseenbachendtiins differt foliis uninerva 1g Scene 0.3-0.6 m apres ramli acute quadrangu- 8 hypan lata setulis appressis 0,1-0.2 mm longis caducis alioqui ge Flores in aepennis brevibus solitarii terminales -meri ad torum) 2.3 X 1.8 mm; calycis tubus 0.2 mm altus “pte ag unisetulosus setula 0.2 mm longa, lobis 1 m lo bus J gna is appresso-ciliolatis apiculata eciliata a? na isomorphica glabra; fila- menta 3.5-4.2 mm; cntherarun thecae 2,1-2.3 mm anguste oblon- gae mimte sae ivo basi inconspicue (0.1-0.2 mn) 3 stigma Benen fem 3 ovarium h-loculare apice breviter perp wa et modice strigulosum Sabslae 0-2-0. mm longis; semina ag x ‘ee mm cochleata lae lection: J, at ne &R. yonere Castaneda gait (hot (fetotype S 2339 > collecte etwe ro Avion, pti de Peri jé, : Dept. Seale lena, ep Tg elev. 3270-3350 m, Nov. 8, 1959. "Leaves light green, Peta 8 and stamens yellow." Paratypes Chet from Sierra de Perijé, Dept. Magdalena): Gas temineneee & Romero aa tanede. 2 082, from Saba bana Rubia east o « 3000- mR; M. L. Grant 10870, from péramo 12 kn cash mactieace of Manaure 1 km from Venezuelan border, elev. 3000 m C, loricarella Tre, the only other species peg — not s h squamose pubescence, does m to be a very Pe relative. C, He nsis should be pin ee sac with C, alpina Naud,, C, » C. cufodontisii e sessil of these * four Species have setulose branch nodes, 3-nerved usually broader > lead blades, much more prominent intercalycine setae 0.6-1.1 mm long. MICONLA ARCHERI Wurdack, sp. nov. Sect, Tamonea, M, kraenzlinii Cogn. affinis sed cum folii Frutex 1-2 m; rami teretes cum foliis subtus inflorescen- tiis bracteolis hypanthiisque densissime cum pilis patulis a 1962 Wurdack, Certamen Melastomataceis 167 stellato-plumosis plerumque 0,3-0.4 mm longis vestiti. Petioli 0.5-1.2 cm; lamina 5.5-11 X 1.5=-) cm elliptica apice ss (O.4-1 en): ‘graditerque acuminata basi obtusi vel vix ndata coriacea, supra primum sparse ssdhgrrenie marginem conc gracili-setulosa (pilis i. s. flavidis ad 1 mm lon- gis) et modice stellato-puberula demum sere et vix rugu- losa, S-nervata nervis secundariis supra inconspicue impressis subtus leviter elevatis et ca. 2 mm inter se distantibus ner- reg subtus inconspicue reticulatis, ad margines denticu- ta distantibus, Panicula 1.5-3 cm ee 3-6 cm longa pyramidato-oblonga; ares plerumque 5-meri in ramis lateral- ibus 5-10 mm longis glomerati (plerumque in dichasiis) ses- Siles; bracteolae 2,5-); X 0.5-1 mm lanceolatae tarde deciduae, Hypanthium (ad torum) 3.5 mm longum campanulatum; calycis tu 1.5 mm altus extus et intus lobi interioresque densissime stellato-puberulus, lobis interioribus 0.5 m altis late tri- angularibus vel oblatis apice obtusis vel rotundatis, denti- bus exterioribus callosis non eminentibus per pilos occultis. * anay 6.5-7 X 3=3.5 mm obovate-oblonga apice pico extus modice stellulato-furfuracea ad cu ye a stellulato-ciliolata intus glabra. Stamina in dimen- Sionibus vix dimorphica in forma isomorphica; filamenta 6 vel 0.3 mm lato et cum setulis ca. 8-15 glandulosis 0.1 mm lo ornato, Stylus 7.5 X 0.4-0.6 mm basim versus sparse vel modice glanduloso-puberulus; stigma truncatum; ovarium 5- loculare 1/3 inferum movin: soatoen set “a eae 0.6 mm altum re Ca tylum pro Type Co Sectieass W. prone re 2 Shoasine US ee eee me Bello, — ° ombia, elev. 1800 m, June 19, 193 Paratypes dail D Dept. poteelgh Archer 768, from Medel- lfn, elev. 1500 m; Molina & Barkle 1640368, Tron ce rro El Volador east of Mede TCH elev. m3 Bro. Daniel 1223, withens parinite locality. mzlinii has as only stellulate hairs on the upper “pm Nthicr, pinoid with very short peor on the 168 PHYTOLOGIA Vol. 8, no. pleiostemonous species with somewhat more compact cence, shorter nthia, and oblong non-attenuate rate Asplund (in litt.) several years ago noted that M. s is synonymous with M. versicolor, The Orinoco res Bonpland Melastomataceae ae: Graffenrieda ro (Bonpl. ) tee Brachyotum folium (Desr.) Tr. as trtted et by me, See Frmongepoi Wurdack, sp. nove Amblyarrhena,. a annulatae Tr, distanter affinis sed a tole cen Gaialsribas peticeliis longioribus antheris brevioribu ex 1-3 m inter; rami teretes nodis incrassatis. pedicelli 5-10 mm longi ad medium vel paulo supra medium arti- culati, bracteolis )-5 X 0,-0.7 mm lineari-oblongis apiculatis h X mm suborbicularia cucullata apice vix emarginata incon- spicue Sooo ad margines cum glandulis sessilibus minntis dense obsita, Stamina isomorphica; filamenta ca, 2 mm longa 0.5-0.6 mm aban: stigma ataatias ovarium ——— 1/2 in Pog. sik har soe conum glabrum 0.9 mm al pro liection: Bro. Daniel 162) eee 1S *LTasBL2) , collected at Abejorral s Dept. Entioquia, Colombia, Aug. 1938. "Corolla red; fruit viole Paratypes (all Dept. apolar’ Bro. Tomas 1517, from Yarumal; Metcalf & Cuatrecasas 30111. from between Valdivia and Yarumal, € Ve 2000 m3 Hod 7, Bartley t Johnson 252, both from be nts of Ia Ceja, elev. 1800-2500 m. While M. rhodantha shows the same branch node swellings 45 M, annola — 4 Gleas,. as well as the same general inflorescence type as ¥, atypifol ja Naud, and M, livida Tre, 1962 Wurdack, Certamen Melastomataceis 169 the placement remains dubious, The anthers are rather like those of Sect. Chaenopleura, but not at all rimose, The leaf texture and venulation ” (bat not the floral features) are reminiscent of that of M. cundinamarcensis Wurdack, while the nthium and of M, compressicaulis Wurdack, From Gleason's photograph, M nodosa Cogn. is somewhat similar vegetatively, but with pli- nerved leaves furfuraceous on the veins beneath and smaller pes flowers, MI CONTA is sian Wurdack, sp. Sect. Cremanium, M. co heal era Wurdack et M, gonio- cladae Tr, affinis sed cum TL Tribus maioribus, Fere 1 vix cordata rigida i. s, lutea, 5-nervata nervulis subtus inconspicue retic ulatis, ad aig inconspicue dense- que serrulata dentibus a ppressis ca. 1 mm inter se distantibus. — 2 Sate om longa Ss uciflora; peaicelis 7-9 mm longi apice vix retusa glabra. e ” 4 somorphica aatives pela: menta 3 m longa; antherarum ated 1.9-2 X 1 mm obovato-oblon- gae late biporosae, connectivo non prolongato dorsaliter basi aoe 3 oe: truncatum; ovarium )(?)-loculare 1/2 inferum lto gla TO s ectiont J. Cuatrecasas 21667 (holotype US Seah SE eoLisctea at "La Palma, lado derecho del rfo Pich- indé , hoya del rfo Cali, Cordillera Occidental, vertiente oriental", Dept, El Valle, sore elev. 2500 m, Jul. 2h, 19h6. "Frutex gant 2’m alt. con ramificacion terminal. ee envés, Céliz andri lo claro. Pétalos id. Frutos rojos, maduros negruscos nicest relatives have hypanthia only atch 2 Bal Similar apart. Suatrecasas 21667 had been distributed as M. coctageiae ¥., 170 PHL TOLOAGLA Vol. 8, no. species of Sect, Amblyarrhena og sessile large thin leaves and calyx lobes at anthesis only 0.5-0.7 mm long. MICONIA PURACENSIS Wurdack, s sparse arachnoideo-furfuraceis non costatis staminum maiorum culato. tex -8 m; rami obscure rotundato-quadrangulares cum foliis subtus inflorescentiisque densissime cum indumento amorphico arachnoideo demum granuloso-floccoso fulvo appresso obsiti. Petioli 0.3-0.5 cm; lamina 2-3.5 X 1-1.8 cm elliptica obt creberrime impressis subtus non vel vix elevatis, Panicula 2-l, cm longa oblongo-pyramidata; flores 5-meri; pedicelli 1-2 mm 1 crassi paulo supra culati, bracteolis non visis, Hypan torum) 1.9-2.2 X 205-26 urceo 2.3-2.l) mm suborbicularia apice vix emr losa, Stamina vix dimorphica glabra; filamenta 1.9-2 a longa; antherarum thecae oblongae 1,1-1.3 X 0.7-0.8 mm apice late (0.4-0.5 mm) dorsaliterque biporosae, Stamina antesepala: connectivum ventraliter 0.5 mm prolongatum et vix Sa roteitaban dorsaliter cum appendice descendente truncata 0.8 X 0.6 mm ro el-0.5 mm et 0.3 mm elevato hebeti armatum, Stylus 2-3 X O.h-0.5 mm glaber; stigma truncatum; ovarium iS 8 atealereae 2/3 inferum eee & Lg in conum crassum 0.) mm altum circum stylum protract Type Collect. tion: J. Cuatrecasas 23419 (holotype US pei isotype F), collected at "filo de la Cordillera Cen- N. del vole4n Puracé, alrededores La de San Rate fael, péramo 3350 m E hgit "Dept. Cauca, Colombia, Jan. 29, 197. "Arbolito m alt. Ho oja “poipene. verde clara, bril- te at colin en Pétalo cos. : Cauca: Giatrecasas hehe (#), from Soph ge del Puracé ar sur grt Volcén e Francisco, 3450-300 m mr i “Corstens caseral: 33 "Jule 19133 . Antonio C, 75 (F), from "Péramo de las Papas, "elev. 3100- 1962 Wurdack, Certamen Melastomataceis 171 3500 m", Jan. 197. El Valle: as 2030 (Ff the at from "cabeceras del rfo Tulud, Queb: s Vegas 300-3500 m alt., phi tara ben tees 2 vertiente i grote " 22 Mar. 196. While M. s losa has similar (albeit ae larger) foliage, its eaets are sessile and erect or patent, the hypanthium is densely granular-squamulose and usually basally 10-costate, and the anthers have connectives with latero- ventral lobes but no large dorsal appendage. The more dis- tantly related M, ochracea Tr. has shortly acuminate dentice ulate leaf blades, glabrous hypanthia, and a capitellate stigm, HETEROTREICHUM RACEMOSUM Wurdack, sp. De congeneribus differt saiecsiecnall racemiformibus,. Frutex 2- m; rami primum rotundato-quadrangulares de teretes betioktans dense persistenterque epee ene pel pilis gracilibus fuscis vel rufo-fuscis ca. 0.5-1 mm longis. Petdioli oe cm; lamina 7-12 X 3.5-6 cm elliptico-ovata apice acuta basi 0.5-1 mn lo ongis, 5- vel sub-7-nervata nervis secundariis supra impressis subtus vix elevatis ca. 2 mm inter se distantibus. te oblongo-lanceolatae extus senanstonie intus glabrae per- sistentes, Hypanthium (ad torum) 3.5 X 2.5 mm calycis dentes exterioresque modice brevi-strigosums calycis tubus 0.6 m tot lobis interioribus 1,2-1.5 mm longis anguste triangu- 1 apice truncata vel vix e marginata "glabra ciliata, Stamina Gimorphica glabra, filamentis 5,9-6.1 vel "bide 5.5 mn, connectivo ad filamenti insertionem dorsaliter inconspicue tuberculato hecae ‘i longis acutis, Stamina minora: antherarum thecae 3.6-3.8 mm longae vix curvatae basi ventraliter cum popcorn coalitae et Prolongatae, connective basi ventraliter ad apicem bilobu- lato lobis acutis 0.1 mm longis, Stylus 6 X Gui 5 = mm glaber; Stigma vix clavato-expansum 0.9 mm diam; 3 ovarium 6-loculare 172 PHYTOLOGIA Vol. 8, no. 1/4-1/3 inferum vix 12-sulcatum sparse glanduloso-strigulo- sum apice in collum 0.6 mm altum glanduloso-ciliolatum pro- tractum, Type Collection: H. Pittier & Me Nakichenovich 151,69 laine Us 19337335 3 isotype ected at "alturas de Las Vueltas, — e Rancho Grande*, Jaa Nacional Henri Pittier, Edo, Aragua, Venezuela elev. 1500 m, July, 1947. Paratypes (ait Edo. Aragua}: T. Garcia 72 (VEN), from El Paraiso, Ranch a mentor Foldats “San (EN) from Cumbre Choronf; F, Fernandez 101 (JS), from "cumbre, pea Ae. Maracay-Choroni", elev. 1775 m All other Secret species of Heterotrichum have cymose or cymse-paniculate inflorescences, Vegetatively, the Aragua endemic resembles » Be zoe treton (Naud.) Gleas., which has cor- ssc leaves and 8-: s flowers with very short external ca eth, mie a ghee scail exappendiculate anthers. Among the eglandular species, H. racemosum resembles in exter- teeth H, strige s > h erved leaves), H. macrodon e Tger flowers and exappendiculate anthers), and H. i Bi Tr. The latter ssness is probably the closest re relative, having ve ventrally a ed stamens but capitate-cymose inflorescences and 7-9(-11)-nerved leaves with only inconspicuously ekg pe upper surfaces. CLIDEMIA AGUACLARENSIS Wurda urdack, sp. C. coronatae Gleas, affinis — iss foliis vix cordatis pedicellis panle infra hypanthium insertis, setulosi pilis muriculatis ad 1 mm longis; 6-12 X 265-5 anguste ovata apice graditer maeaxsenne 2 hebeti-acuta basi 5 m) cordata membranacea 7- v = ne 8 demum glabrata, gubtas da narvis arimeriie ¢ sparse setulosa a pean fe laevibus ad 1 mm longis in superficie glabra, nconspicue SoG Blas ine Bin, A dentibus setulosis pst Sparesiie ca. 0.3 om nese Inflorescentia diffusa in ramorum axillis e basi 3-5-ramosa, ramis divaricatis ad 7 cm rioribus callosis non emine entibus; torus sparse pi 1962 Wurdack, Certamen Melastomataceis Petala 1.7-1.8 X 1 oe mm oblongo-obovata apice rotundata glabra, Stamina fere isomorphica glabra; filamenta 1.1-1.2 mm; antherarum thecae x. vel 0.8-0.9 mm longae crasse oblon- gae apice ca. 0.1; mm latae et minute (0.1 mm) uniporosae, Type Collection: J. Cuatrecasas 22067 (holotype US 2338667; isotype As ocliasted Weerca del puente de Agua- clara, hoya del rfo Anc hicay&, Cordillera Occidental, ver- tiente occidental”, Dept. El Valle, Colombia, elev, 120 m, Sept. 29, 196. nSutratex robusto, 1 m alt. Ramas divari- cadas, densas. Hoja cartd&cea, verde oscura has, verde pélida envés, Pétalos blancos, Antera anerie las sta Rican relati ngs iiy broadly pital le af ong ted at the h thium base, C, £ sa Donn, more distantly related, hawing purple- e branchlets amd lacking stellulate hair rp pe Lemaherie: of the three species “s also in the next-described species) are actually terminal, Coming pseudolateral by epi eterpir branch overgrowth. general aspect of C, aguaclarensis is rather like that of the two h-merous species urpureo-violacea Cogn. and C, rami- flora Gleas,, as well as Saat Species of Ossaea. Sponrer a at Wurdack, sp. mm longis) et modice rence stellulato-puberuli. Pett 0.8-0.9 mm longae crasse "oblongse ro 0; *> ae ti ngato dorsaliter 0,05 vel 0.15 mm calcarato. tylus 3.4 X 0.25 mm glaber; sti catum; ovarium 3- ri loculare 2/3-3/h inferum apice conico sparse cum glandulis minutis clavatis et circum styli basim sparse cum setulis 173 17h PHYTOLOGIA Vol. 8, no. ornato ype Collection: J. Cuatrecasas 237h2 ear US 2338702; isotype F), collected in "quebradita del km. 51, rdo del rio San Juan en la regién ie Queremal , hoya del rfo Digua, Cordillera Occidental, en occidental", Dept. "EL Valle, Colombia, elev. y15h0-1650 nm, Feb. 25, 1947. "Frutex de ramas bejucosas, Hoja rugosa, de la inflorescencia violéceas. Pétalos blancos. Anteras amarillas," Despite the marked pubescence disparity, C. ee seems to be merely a higher-elevation version o clarensis. In Costa pra C. reitziana Cogn. & Gl, ex Gl. ronata Gl, are a similar pair of eerie with however the reverse Sibi satthihessence relation er ate Pages Wordack, Sagraea, C, domnél nant th i Cogn, distan aevtias oe Cum foliis minoribus sessilibus glabris postion thii pilis petalisque Sheep teretes foliaque glabri, Folia sessilia vel vix (ad 1 mm) petiolata; lamina 3-7 X 1,5-3 cm subcoriacea apice longe (1-2 cm) graditer angusteque acuminata basi vix (ad 2 mm) cordata S-nervata nervis secundariis non vel indistincte bus persistentibus; flores -meri, Hypanthium (ad torum) 1.9 mm longum dense setulosum setulis simplicibus 0,8-1.3 mm eminentibus linearibus anguste acutis sparse setulosis; torus glaber, Petala 2,3-2.5 X 0.9 mm oblonga a —_ rotundata gla- bra, Stamina isomorphica glabra; filamenta 1.3 mm longa; poet re bia oe 7 ma longae oblongae minute (0.1 mm) uni- » connectivo non prolongato non appendiculato, Stylus an x 022 mm nt stigma truncatum; ovarium h-loculare omnino inferum glebrum, Collection: J, gosteepasts 19987 (holotype US 233866) , collected at "que = te : he Bu » Costa del Pacffico," yrs El “Valle, por elev, 0-10 m, Feb, 24, 196. "Fruticuletum epf{fito d 1962 Wurdack, Certamen Melastomataceis 175 colgantes, Hoja verde clara. Céliz verde claro, Pétalos blancos. Filamentos blancos, Anteras rojizas," Vegetatively, the suggested Guatemalan relative, with its setose branchlets and larger petiolate strigose leaves, does not resemble C, fausta; however, the floral structure r la rom calyx teeth and larger (3.7 X 1.1 m) petals, eral vegetative aspect of the Buenaventura "lucky") species is e the rather like that of Mouriri parvifolia Benth., whil inflorescence and long sepals reca issaea. MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. III Harold N. Moldenke L. Additional & enended literature: Brunfels, Herb. Viv. Icon. 1: 119--120. 1532; Fuchs, Hist. Plant. Basil. 592--593. 15:2; Ruel, Ped. Dioscor. Anazarb., ed. 1, 317.(15i7) and ed. 2, 519— 521. 1551; Tragus, Stirp. 210. 1552; Gesner, Collect. Stirp. 113. , Hist. Plant. Univ., ed. 1, 3:hh3—hbb. 3 Bont., Hist. Nat. Ind. Orient. 150. 1658; K. Bauhin, Pinax Theatr, Bot., ed. 2, 269. 1671; K. Bauhin, Prodr. Theatr. Bot. 125. 1671; Imperato, Hist. Nat. 673--67). 1672; Matth., Apolog. 1. 7, fig. 7. 1692; Petiv., Mus. Petiv. 60. 16953 Pluk., Almag. Bot. 283 & 362; pl. 321, fig. 1. 1696; Sloane, Cat. Pl. Jamaic. 64. 1696; Magn., Hort. Reg. Monsp. 203. 16975 P. Plant. ‘ 285-~288 & App. 249. 170; P. Hermann, - Bot., ed. 2, 2h2, Pl. 242, 1705; Sloane, Hist. Nat. Jamaic. 1: 171, pl. 107, fig. 1. 1707; Boerh., Ind. Pl, Hort. Lugd. 1: 187. 1710; Munting, Phytograph. Cur. 784. 1711; Barr., Pl. Gall. Hisp. 30, pl. 116. 17k; Vaill., Serm. Struct. Flor. 49. 1718; . Plant. Hort. Act. Lugd. 1: 186--187. 1727; P. Hermann, Thes. 176 PHYTOLOGIA Vol. 8, no. h Zeyl. 228. 1737; L., Hort. elitt; 10-11. 1737; Breyn, Prodr. $y e v9, & “e Toh. 1725 ae Plant. Gall. 30, Pl. 055 & 116.’ ATs 145, 28, & we. "1751; his Sp. Pl., ed. 1, side 18--21 & (1783) aka 27° 6c, 630, 660, 398 & 879. 1753; Fl. Suec., ed. 2, 10. 1755; Kniphof, Bot. Orig. Herb, Viv. = “ital, 1757; Do- dart, Mem, Serv. 1'Hist. Pi. _ 3, 627, pl. 35. 17583 J. Fe Gronov ae Bak oa. 2, hk, & 146. 1762; A. Haller, Stirp. t. 1: 96--97. "sake, f. ou Murr. = —_ Syst Veg., ed. 13, ee 1774; Sabbat in Martelli, Hort. - 3: 10--11, pl. S4--56. 1775; Oeder, Fl. Dan. h: pl. 628. 17753 0 O. F. Mtll., Icon. Plant. Fl. Dan. hl: 5, fon 628, 7715 gee —— Fl. 9, ple zs ee Desf., a ant. 1: 15-17. 1800; G. 3: 502--503. 1805; Poir. in pees tigi “yéth. “Bot. 8: eh 581 1808; H.B.K., sa Gen. & Sp. » 22 272-- 277. 1818; Hook. , Bots - OF 966. Bias, “Gabrera, Bol. Argent. Bot. 1: 67. 1 An eae abbreviation employed herein is Ws = University of as teae tae Madison, Wisconsin. Pefialosa 871 dist ributed as "Verbena sp." is actually pouches Steet prismatica var, ees Grenz._ Additional excluded species Verbena amer, frutescens teacei a foliis & facie, floribus caeruleo-purpurascentibus amplissimis Breyn = Stachytarpheta mutabilis (Jacq.) Vahl Verbena asparagoides Gill. © Hook, = Junellia asparagoides “(Gilly & a Moldenk Verbena caroliniana, icenbe; foliis oblongo-obovalibus, obtusis obtusis; Spicis pihdtaa icuiciseinis: Stine Wichx. = Stylodon carneus (Medic) Moldenk ee connatibracteata var, piahetoe Skottsberg = Junellia erina ~~ junipe Lag.) Moldenke ecere erioclona Schau. = Callicarpa erioclona ose ) Verbena fruticosa Houst. = Phyla nodiflora var. s (H.B-K- ~~ Moldenke, in part, & P. strigulosa (Mart. & oa Ty yisldenke, in Verbena fruticosa Mill. = Phyla nodiflora var. reptans (H.BéKs ) " *Moldenke, in part, & P, strigulosa (Mart. & Gal.) Moldenke, Verbena mas Fuchs = Erysimm officinale L., Brassicaceae Verbena mexicana trachelii folio, fractu aparines Dill. = Priva ~~ mexie mexicana (L. ana (L.) Pers. vaelion eek nodiflora Matth. = Phyla nodiflora (L.) Greene Verbena nodiflora curassavica foliis menth menthae Norison = Ghinia 1962 Moldenke, Monograph of Verbena 177 curassavica (L.) Millsp. Verbena nodiflora foliis minoribus viridibus Moris. = Phyla nodi- flora (L.) Greene __ Verbena orubica teucrii folio » primulae veris flore, siliqis & serizioas 1 longissimis; caudo muris vulgo Parkins. = Stachytar- phe pheta mutabilis (Jacqe) Vahl Verbena racemo simplicissimo, floribus sessilibus, calycibus » Exootus ret pores recenoqe appressis Gronov. = Phryma lep lepto- Stachya LD vihacs ie eslaghioTdes a th = Junellia selaginoides (Kunth) Mol- Verbena spathulata var. pseudojuncea Remy = Junellia pseudo juncea Beay (Px Gay) Moldenke Verbena thymifolia Cav. = Junellia thymifolia (Lag.) Moldenke ABRAMSI Moldenke Additional citations: CALIFORNIA: County undetermined: Newlon 271 [Alexander Valley] (Wi VERBENA AMBROSIFOLIA Rydb, Additional literature: Daniels, Fl. Boulder Colo. 20). 1911. Waterfall found Vv. ambrosifolia in Mexico on flat mountaintops with acacias and on stony flats with acacias, mimosas, and cacti, while Wynd & Mueller found it in moist places on deserts. The Cory 53507 and Wynd & Mueller 150, distributed as V. ambrosifolia, are actually Vv. ~bipinnatifida Nutt. On the other hand, herbarium material of V. ambrosifolia has been Breidentl find and distributed as V. ambrosifolia f. a. losa Perry, V. aubletia L., V. bipinnatifida Michx., V. bipinnat ifida Nutt., V. bipinnatifida var. latifolia Perry, V. bipinmtt fida var var. latiloba Perry, V. bipinnatifida var. latilobata Perry, Ve bi eereee ichx. » Ve canadensis (L.) Britton, V. ciliata Benth., V. gooddingii nents V. pinnatifida Nutt., V. racemosa Eggert, and V. wert PE al Re ca he only recorded common names are "moradilla" gonnegvecteion tee vervain", "verbena", and "western pink verbena", but, according t his plant is called "é co.yin’az The Whitehouse 8741 ee btw atl below may eet have been collected in Santa Fé County -- its label pets fe "Las Vegas -~ Santa Fé"; while Whitehouse s.n. [9.3.1929] may > bave Come from =~ Miguel County -- its label merely con "Santa Rosa een Co.: E. C. Smith s.n. [South Den- co ver, 8 June 1891} (E). Fremont Co: T. S. Brandegee s.n. [Caffon 178 PHYTOLOGIA Vol. 8, no. City, May 1871] (E). Otero Co.: Osterhout s.n. [Rocky Ford, June 8, ne Saat ate. Rose & Fitch 1750 (E, W)~ Pueblo Co.: Greene’ t 5 (G)3 E. L. Johnston 9 986 (E). County undetermined: Os- terhout 3165 [New Windsor] (G (G). OKLAHOMA: Cimarron Co.: G. We. Stevens 48h, (E, G).» TEXAS: Hudspeth Co.: Ferris & Duncan , 247 (E); Orcutt 6096 (E), 6197 (E). Presidio Co.: Eggert Sn. . [near Marfa, 14 May y 1901] (E). Val Verde Co.: Moore & Steyermar (E). ERE undetermined: Moore & Steyermark 3622 [McKittrick Canyon] (E). NEW MEXICO: Bernalillo Co.: E. J. Palmer 31192 (E)5 Rose & Fitch 17801 (E, W). Colfax Co.: Mathias 5u5 ( ). Don Dona Ana Co.: Wooton 642 (E, W), sen. (Organ Mountains, 16 Aug. 1893] (£). Eddy Co.: Wooton s.n. [Queen, 2 Aug. 1909] (E, W). Grant Co.: O- B. Metcalfe 1231 (E). Lincoln Co.: Earle & Earle 236 (E, Po); Wooton 363 (E, G).» Quay Co.: G. L. Fisher s.n. [Nara Visa, 21 A- pril 1911] (£). Rio Arriba Co.: Eggleston 664 (W). ee g 4% Ars8ne & Benedict 16306 (E). San Miguel Co.: Anect 100 (G), G)3 P. C. Standley 951 Fr. @ anta Fe Co.: mnge s.n. [about Santa ae Sept. e83) (E); Wislizenus 525 ( = Socorro Co.: Vasey sen. [Socorro, May 1881] (BE, F, W)- nee) ae Wooton s.n. [near Fairview, 18 July 190] (W). County un mined: Fendler 586 (D, E, Fy G, W); Griffiths Lakh fBouth spring] (W). ARIZONA: Coconino Co.: Edw, Palmer s.n. mer SM. [Little Colorado, 1869] (W). Navajo Co.: L. F. Ward s.n. [June 17, tet (W). Pima Co.: Mearns 2677 (E, W). MEXICO: Chihuahua: Stearns 10) (W). Coan huila: Ee W E. W. . Nelson 6789 (W); Edw. Palmer 1050 Sr aio) (G)3 Pringle 3467 . 3467 (F), 13157 (F, G, W). Nuevo Leon: Edw. Palmer 1050 Lote d a! (E, W). It should be noted that she cites lore r 586 " this species, but does not cite this r for any otliae species of the gems in her alphabetic list of stationed She reg Visher 2132 as ve. bipinnatifida. In all, 351 i specimens ; gene the types of all the names * invo ve been examined by m Citations: SOUTH Sines Washabaugh Co.: ie 2132 (N)- KANSAS: Douglas Co.: Yuncker & Welch 1,019 (Dp). Neosho Sho Co«t Je E. Miller SNe (May 1888] (Cm). . ARKANSAS: I Izard Co.: Demaree 1700 (N). COLORADO: Adams Co.: N. K. Berg 4185 ee mi JE Grant s.n. [Quinby] (Se--28358). Archuleta Co.: Ferril [May 31, 1902] (Bl—l22)3); Weber & Livingston 6258 Te 3812h8). Baca Co.: W. A. Weber 3286 (Au--121563, Gg—370l 73, . Boulder Co.: Ewan 11188 (En); H. C. Hanson s.n. (Boulder, June 26, 1920] (N); J. R. MacFarlane s.n. [1861] (Bl--l2276); Penard aus "(c); Ramaley 15322 (Au, Bl—h2365, Gg—-267607, Mn—32166); Tweedy 5125 (N); A. G. Vestal s.n. [June 11, 1913] (Du—1)3286) Ems. po = Bee (Du—1)3577). Denver Co.: Eastwood 140 en B1l— ——|\2 Ca—~1),8188, Gg--31 sn, [Denv 1882} * pl —loale}, ro ee : 31209) gone Thee ay, N), “sens 1962 Moldenke, Monograph of Verbena 179 {in the mts., July] (Bl--l22)2); Ferril s.n. [May 13, 1902] (Bl-- 4220). El Paso Co.: H. Hapeman s.n, [Colorado Springs, July h, 1920] (Hp); M. E. Jones 122 (Br, Du—151798, Go, Po—711sh, Ua— cing Va-—-11392), sn. [Colorado Springs, May 1878] (B1—12277) 3 F, D. Kelsey s.n. [May 1878] (Ob—50860); Ownbey 718 (P1--98766) ; sitet 15h8 (Bl—-l2275). Fremont Co.: T. S. Brandegee 5 (Ca— 177631) » Sn. [Canon City, Apl.-May 1871] (Pr), s.n. [Canon City, 1877] (Ca--169645); J. H. Ehlers 7516 = Mi); A. Nelson 10548 (Ca—500825, Du--252578, Me—37, Mn——20027, N, Po—190666, S);_ Osterhout s.n. [June 22, 1926] {po--11373053 Penland 1209’ (Gg— ale Redfield 521 (Pr). Gilpin Co.: Scovell s.n. [Central City, 1869] (Mi). Huerfano Co.: Shear nied ALD A sen. [6/ 16/3] (Bl—22h)), Jefferson Co.: Je 7 (Bl—2300); Huestis s.n, [Morrison, June 10 190) CAL ae Pp ated Sons Deer Creek, May 25, 193k] (Gg—-220518 ). Larimer Co.: J. He Cowen s.n, (Herb, State Agr. Coll. Colo. 545] (Fe); Crandall s.n. (June 23, 1896] (Fc), s.n. [June 27, 1896] (N). Las Animas Co Archibald A.255 (Bl—l2356); Beckwith 139 (N), 162 (Di--123523); Eastwood 5568 (Gg—31288); Ewan 1295) (En); C. Ms Rogers 6015 (Au--12156l,). Otero Co.: Huestis s.n. [June 17, 190k) (Bl-— 42283); Osterhout s.n. [Rocky Ford, June 8, 1900] (N--type); Le Paull 3 (B1--l22)5), 58 (ie koe70) Rose & Fitch 1750) (MN). Pueblo Co.: J. H. Ehlers 6805 (Mi, Mi), C6lL (M)5 Ban Ewan 11995 (En), 14157 (B1—53188, | En); Hicks & Hicks sn. [Pueblo 90] (Dt, Ob--97282) ; W. W. Robbins 5011 (Bl--)2357, at eset Re C. Rollins 1209 (N); Rydberg & Vreeland 5676 (N); Skjgt- Pedersen 1220 (Cp); ; Waterfall 10858 (St). Weld Co.: Osterhout 3.n. [May 27, 1908] (Du—91153); Ramaley 10439 (B1—-]236h). County undetermined: Adney s.n, [Colorado] (Ob--50859); E. = peniera sen. [July h, 1897] (Vt); J. H. Cowen 1186 [Soldi Canon] (Fe, N), son. [Soldier Canon Road, June 20, 2, 1699) (To 22986); Vasey 827 TRocky Mts,, lat. jo--1°} (Pa), OKLAHOMA: Cimarron Co,: Demaree 13335 (N, Ok); G. J. Goodman alae (0k) 5 Goodman & Waterfall 1811 (Ok); Hopkins & Van Valkenburgh 5809 (Ok); G. W. Stevens 8k (ok, Ok, St—9217, Ur); Waterfall 7437 (Ok), 9070 (Ok, st), 9737 (St). Comanche Co.: M. K. Clemens =& = oe, ides Jest oe an He T. Robbins 2335 (Ok). 7 (N); Cory 28677 (N), 28678 (H), 25ee1. (nt), "39868 5 ti) “TO3 367 7 N), 4L80h (au); G. F. Fisher s. ne he (Alpine, Aug. Aug. 23, 1932] (Ge—23786)); E. J. Palmer 342h0 (WN); Rose~Innes & & Moon 1172 (au); 0. E. Sperry 7.1072 (Ca—802805) 5 Steiger 1097 (N); B. H. Warnock 20121 (Au), 20921 (Au), 7.66, in Part (Ai (Au), W.283 (Au, Ca--882560, Ca—-8826)9, N, N), W«625 (N); GL. Webster LL66 (3 (St). Culberson Co.: M. A. Chase 6034 (Mi); Gory 53007 (N); Hitchcock & Stanford 6782, in part (Ca--0826L8, 180 PHYTOLOGIA Vol. 8, no. Or--512h); Moore & Steyermark 3622 (Ca--71229, Du--2277, Gg-- 194779, Mi, N); Scholl 8 (Au); Sperry T.129 (Om); Tharp & Janszen :9=1169 (Au--121572); Waterfall 3775 (St), bu58 (N, |, Pl--151556, St); Whitehouse 1689) (Sm), 16928 8 (Sm); M.S. = Sone ii Kent, 8/6/16] (Au, Au, Au). oe Co.: Ferris & Duncan 27h Du--126178, Gg—-31431) ; C. H. Muller Bois (St) ; far TS TE (an, Ca-~882559, N), 46212 (Au), 46217 (Au), 46235 (Au, N); Waterfall 3851 (St), 4895 (N, St). Jeff Davis Co.: “Gory 40365 (Au); Moore & Moore 2 (Mv); Scholl 22 (Au); Warnock 21676 (Au); u); Waterfall 4722 (N, Tu-30988) ; Me $33 Young s.n. n, [Davis Mts., 9/18/18] (Au, Ca-- 8828hh). Pecos Cos: Ge Le Fisher s.n. (Sheffield, July 20, 19 (B); Parks & Cory 18352 (Tr); Tharp 43-796 (Al, Au, N, Ok), 43- 197 (Au, N, St). Reeves Co.: Cory ory 52132 132 (N). Terrell Co.: Parks & Turner 79 {Warnock 2050] (Au--1228)8). NEW MEXICO: Bernalillo Cost C. C. eg Fay BS 17 (N); Koelz s.n. [Sandia Mts., June 28, 1926] (Ii), Soe . [Sandia Mts., July 5, 1926] (Mi); Nelson & Nelson 169 (S)3 E. J J. Palmer 21192 (Gg--151769); Rose & Fitch 1 T7801 (N). Chaves Cos: F. S, Earle 322 (Vt); Earle & Earle 322 (Bm, N, Po—6 3867) 5 N. A. - Palmer 60, in part (Go, N, S). Colfax Co.: Ewan 13137 (En); Mathias 515 (Po--180697). De Baca Co.: A. Nelson 11307 (Ca--500721, Mg--l,0). Dona Ana Co.: Archer 7349 (Ca—382650); Wooton 62 (N, Ur), s.n. [Organ Mts., Sept. 17, 1893] (Ur), sen. [May 26, 190k] (Mi). Eddy Co.: A. Nelson 11401 (Mg--38); Waterfall 3720 (st); Wilkens 1793 (W--1650339). Grant Co.: 0. B. Metcalfe 126 (Ca 130179, Ca--882853, Mv, a; Mulford 681 (Io—33158, Ur); We Stewart s.n. [Mimbre Mts., June 26, 1936] (Ca—8825)6). ocadat- upe Co.: 0. Degener })515 ( pe A. Nelson 11338 (Mg—l1); White~ house s.n. [9-3-1929] (Au). Hidalgo Co.: F. A. Barkley 11786 (au). Lea Co.: Eastwood isla? (Gg—-161887); G. L. Fisher r O11 (Ew), 40119 (Ur). Lincoln Co.: F. S. a 638 (N)5 Ee Beelester 18917 (W—153%27) ; ii tcwsbek. Rethke, & Van Raadshooven } (Du--256178, La, pL—O9793, Se-=18010, GaebeuT)y Toten (Po—61523), 33 (Ca--10831, N, Po--71165, Ur); Wooton & Stand ley 3635 (Or-8875), 3691 (or—8858, Or--8859); Worth sen. [Bon nell's Ranch] (Ru). Otero Co.: Cutler 2002 (Gg--2712h2) 5 )) Bie ley s.n. [White Mts., June 193%] (Au, Au); Waterfall 12971 (St Quay Co.: Bailey & Bailey 101251 (Ba); G. L. Fisher s-M- jen. [Nara Visa, Aug. 11, 1910] (Bw); G. J. Ikenberry s.n. (May 1, 1937] (St). Sandoval Cos: Be P. Keesecker 53 (Ok); A. A. De Read s-fe (June 27, 1915] (Io-=109)57). San Miguel Co.: Anect 13 (MV), 179 (N); Ars8ne 18257 (To—128668), 18528 (B, N), 18615 (N, Po— 18765), 18793 (Po--1h8318), 21058 (Mi); Bacigalupi 61 (Ca— 882562, Du--286399); Cockerell s.n, [Las Vegas] (Gg-— 31290); Ge J. Goodman 2308 (Io—~1h2528, Ok—19552); A. Nelson 10548 (0 Up), 1962 Moldenke, Monograph of Verbena 181 11566 (Mg—36); C. L. Porter 3016 (Pl—1177l8, Se--7896, St-- 25230) ; P. C. Standley 4951 (N); Sturgis s.n. [May 4, 1902] (Gg); Whitehouse 871 (Au). Santa Fe Co.: Arséne 16555 (B) 3 Bacigalupi 265 (Du—286)01); Bertaud 36 (N); W. R. Dudley sen. [1908] (Du-- 9547); Koelz sen. [Santa Fé, August 1927] (Mi); A. Nelson 11757 (Mg--39); Snow s.n. [June] (F1--21083) 3 Vasey son. (Glorietta, June 1881] (Du--90892). Sierra Co.: Beals s.n. [Lake Valley, June 190h] (Mi); 0. B. Metcalfe 1090 (Ca--852566, Mv). Socorro Co.: I. Ee Diehl 386 (Po—-71132), 491 (Po—71131); C. Be Wolf 2890 (Ba, Du-~190958, Gg—~17h785, Po--174759, Rs--9993). Torrance Co.: F. Clark 60 (St--9202); G. L. Fisher s.n, [Torrance, July 30, 1907] (B)» Union Co,: Brenckle 8081 (N); Demaree 1361 (Du--2970h6, N); Eggleston 20107 (W--1523089). County undetermined: Arséne s.n. {Tijeras Canon) (Ca-~594,662); Benedict 2329 [San Jeronimo] (8); Fendler 586 (Pr, S). ARIZONA: Coconino Co.: Clute 132 (N); H. E. Lee s.n. [9/17/36] (Tu); Shapley s.n. [Sunset Crater Natl. Mon., July 14, 1938] (N); Shreve 4772 (Fs). Graham Co.: R. J. Davis 213-35 (Ld), 225-35 (N). Navajo Co.: L. F. Ward s.n. [June 17, 1901] (N). Pima Co.;: H. W. Graham s.n. [near Stratton, 8-30-46] (Ca--882561); Mearns 2677 (Du--958, N); Shreve 517 (Fs). MEXICO: Chihuahua: H. S. Gentry 7998 (N); LeSueur 1528 (Au); E. Wilkinson S.n. (Chihuahua, 26 Sept. 1885] (Io--92227). Coahuila: Herb. Inst. Biol, Univ. Nac. Mex. 7138, in part (Me); M. E. Jones 634 (Po— T1154); Marsh 707 (Au); E. W. Nelson 6789 (N); Pringle 3467 (Io— 38747, Me, Me), 13157 (Gg--4219h6, Mi, Vt); S. S. White 1743 (Mi); Wynd & Mueller 176 (Mi, S), 572 (N, S, St, Ur, Ur). Durango: H. W. Viereck 395 (W—-1687222) 3; Waterfall 16167 (Z). Nuevo Leon: Frye & Frye 246 (Du--289708, N); Heard, Webster, & Barkley 14512 (Au, N); R. Schneider 927 (N). Querétaro: Agniel s.n. [Arséne 10252] (Ur); Waterfall 1655 (ca). San Iuis Potosi: C. L. Lundell 2590 (I). Sinaloa: J. Gonzalez Ortega 710 (Me). Sonora: H. 5. Gen try 2122 (Ge), 7971 (N) . State undetermined: Fendler 586 [Mexique, 1850] (au—12282h,). LOCALITY OF COLLECTION UNDESIGNATED: H. R. Reed 31,063 (Ur). VERBENA AMBROSIFOLIA f. EGLANDULOSA Perry, Ann. Mo. Bot. Gard. 20: 328, 1933, Onymy: Verbena ambrosifolia var, eglandulosa Perry ex Mol- denke, Suppl. List Invalid Names 7, in syn. 191. re: Perry, Ann. Mo. Bot. Gard. 20: 312, 328, & ae 18 & 42. 1942; Schnack, Anal. Inst. é 92; Moldenke, Alph. List Invalid Names }5, 1942; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 11, 12, 14, 18, & 101. 182 PHYTOLOGIA Vol. 8, no. 4 1942; Moldenke, Alph. List Cit. 1: 4, 157, & 161 (19h6), 2: 72, Mids & ve (1948), Db: 12 15 ctedsc lone 1g. foes 1335 & a82 (1949), and h: 990, 991, & 1150. 199; Moldenke, Known Ge istrib. Verdenac., [ed. 2], 19, 21, 23, 25, 26, 32, & 197. 19493; Moldenke, Am, Midl. Nat. 59: 349. 1958; Sere ak Phytologia 5S: 330. 1958; >, 9 = Moldenke, R6 25, 28, 1 357, 421, & 470. 195 This differs from the typical form species in its more hirsute- escence on the flowers and in the absence glands. eohees ers describe it as a low, ascending or erect, branched, and spreading ann’ erb, with fibrous roots and odor less flowers, the topetias Vairine from deep vivid-purple or aoe es reddish-purple or lavender. The chromosome number s 2 = The type of this form was collected by John Michael Holzinger at Santa Rita, Grant County, New Mexico, on August 1, 1911, and is deposited in the herbarium of the Missouri Botanical Garden at St. Louis. The plant has been found growing along roadsides, in open age. and waste ground, juniper Rete , open ground and fields, Sandy Kelis nak" Die banks of arroyos, in sandy, clay, or black clay soil or sandy loam, at altitudes of from 10 to "3933 meters, blooming from February to October. It has been collected in fruit in April, July, and August. Runyon says that in Cameron County, Texas, it is "very abundant.....It covers acres 0 ground and is widespread"; in Bes County he says it is rere in open fields in the Beeville region". Mueller reports tha Nuevo Leon it is "common in upper pine woods", while peer says it is "abundant on open rocky slopes" there. It is separable from V. bipinnatifida only by its shorter bractlets. Herbarium material has been misidentified and distrib- uted as V, ambrosifolia Rydb., V. ambrosiifolia Rydb., V. bipin~ natifida Nutt., V. canadensis (L.) Britton, V. ciliata Benth., the, Ve ciliata var. longidentata Perry, and V. wrightii A. Gray. The pene between this form and V. canadensis is know as xV. fecunda ch see, ined mare Miss Perry cites the following 1) specimens not yet seen by me: TEXAS: El Paso Co.: Wagner 97 (W). NEW MEXICO: Bernalillo Cos: E. J. Palmer 31153 (B). Dona Ana Co.: P.c.S Standley 5-Me [mesa just west of Organ Mts., 13 June 1906] (W); Wooton age? (Organ Mts., 1) July 1897] (W). Grant Co.: Holzinger s.n- [san Rita, 1 Aug, 1911] (E—type, W--isotype); Mearns 2278 (Wr). mer algo Co.: Mearns 2486 (W). Socorro Co.: Wooton 2635 (W). ARIZONA: Cochise Co,: Carlson | sen, (Douglas, 13 May 1915] (G, W); W. We Jones 185 (G). Pima Co.: Toumey s.n, [Santa Catalina Mts., » Sept. 15, 1896] (W). MEXICO: Sonora: Mearns 1138 (W). The Wagner 974 cited above may actually not have come from El Paso County s since its label says "El Paso to Monument no. 53". In all, 29 bitin sole specimens have been "examined by me. Citations: WYOMING: Fremont Co.: T. A. Williams s.n, [Wind 1962 Moldenke, Monograph of Verbena 183 River] (N). OKLAHOMA: Tillman Co.: Demaree 12188 (N). TEXAS: Bee Co.: Re Runyon 2187 (Rr). Cameron Co.: R. | Runyon 1782 (Rr, Rr), 218) (Rr). Childress Co.: Tharp s.n. (Childress, Apr. 1930--31] (8). Culberson Co.: M.S. Young s.n son. [near Kent, 8-6-16] (Mi). Dickens Co.: J. Engleman s sen. [March 25, 1940] (Ok). Hardeman Co.: E. O. Hughes s.n. 3 sen. [April 30, 1950] (Ok). Jones Co.: B. Adams 9 (Nt). Nueces Co.: Tharp 5602 (Au). Rockwall Co,: E. Me Moore 8. 8. n. [5 mi, east of Rockwall, 8-8-38] (Nt). Webb Co.: Tharp s.n. fuarch 15, 1931] (Au). Young Co.: McCart 836 (Nt). County unde- mined: Herb. Univ. Texas s.n. (Au, Au). O: Grant Co.: I. E. Diehl 9) (Po—7116h); 0. B. Metcalfe 1231 (Gg—28271h). Hi- dalgo Co.: Antisell 186 (N). Luna Co.: Munz 1231 (Po—123619). oe Cochise Co.: Griffiths 1919 (N). Pima Cos: ae SM. (Santa Catalina Mts., Sept. 15, 1896] (N). MEXICO: vo Leon Herb, Mexican Biol, Exped. Univ. Ill. 927 (N); Ce fa "Mueller 2275 pueatieestinn -deeiilceges., conememnene: ceoumees. Sac os (Fs, 1 Mi); Re. R. A. Schneider 927 (Ur, Ur); A. J. Sharp p 45665 (N 65 (N)e VERBENA AMOENA Paxt., Mag. Bot. 7: 3. 180. Synonymy: Verbena grandiflora Sessé & Moc., Pl. Nou. Hisp., ed. 1 [La Naturaleza, ser. 2, 1: app.) 6. 1889 [not V. grandi- SS Bodger, 1936, nor Michx., 1821, nor Ort., 1797, nor Steud., 1895} Literature: Paxt - Bot. 7: 3. 180; Marnock, Floricult. — 5: pl. 52. 1803 Gerkiecttce Univ. 2: 135. 1811; Sessé » Pl. Nou. Hisp., ed. 1 [La Natu veers ser. 2, a appe] 6 1889; Sessé & Moc., Pl, Nou. Hisp., ed. 2, 6. 1893; Jacks., Ind. Kew. 2: 1178, 1895; Hill, Ind. Kew. ri 7: 29» 192 9; Stapf, + Lond. 6: 428, 1931 , Ann. Mo. Bot. Gard. 2k 250, 312, » & 354. 1933; Moldenke, Pre - List Invalid Names 46, 19) rst Alph. List Taveled Names 47. 1912; Moldenke, Known r. Distrib. Verbenac., & 101 (1942) and [ed, 2}, 32 & 197. 5125 uolderke, P aa't List Cit. 3: 831. 1949; Moldenke, Résumé 38, 365, & 70. 1959; Moldenke, R&-~ sumé Suppl. 2: 8, 1960, Illustrations: Paxt., : 3 (in color). 180; Mar- hock, Floricult Mag. 5: ae sé ait sciet: 18,0; Horticult. Univ. 2: 135 (in color). 18h, Herbaceous perennial; stems about 5 dm. tall, partially re- oO 5 & af Af sf fe 5 © 4 : p —6 cm. lo Pinnatifid or "bipinnatifid, with the lower part entire, forming & broadly margined subauriculate and seni-anplexicaul base, the lobes or divisions oblong or linear-oblong, remote, sparsely in- Cised, re apa spototeadiadaacact above, hi spidulous beneath, the midrib secondaries prominent; spikes terminal, dense fascicle-like, during anthesis, long and erect, bracts numerous, sues anoneh ihe or awl-shaped, subul ®scents calyx about + 10 mm. long, densely pubescent fagpestally 18) PHEYTOLOG ZA Vol. 8, no. 4 along the veins), glandular, the teeth slender, subulate, similar in form and pubescence to the bracts; coroll a hypocra ateriform light- or pinkish-purple, its tube slightly longer than the calyx, pubescent outside, especially around the throat, the limb about 9 mm. wide, the lobes 5, eee sty aneeosSia enlarged distally, bifid; anthers unappendaged; mature fruit not kn The type of this Slaeia species was cates ted from cultiva- t Chiswi England, by Mr. Edmonds, gardener to the Duke, in the autumn of 1839. The type of V. grandiflora was collected by Martin Sessé y Lacasta and José Mariano Mociflo (no. 99) at Puruandiro, Michoac&n, Mexico, and is deposited in the herbarium of the Jardin Botanico at Madrid —a PD tbe of the type is in the herbarium of the Missouri Botanical Garden at St. Louis. Paxton describes the leaves of this plant as stipulate, with the "stipules much more hairy than the blade". This is an * error in observation. There are no true stipules in the family Verben- aceae, What Paxton saw was the subauriculate and semiamplexicaul leaf-base. Perry says very truly that "This unique species is readily recognized by its coarse habit, together iii its bipin- id, subauriculate and semiamplexicaul leaves." The Pringle collections cited below agree well with Paxton's illustration and with the photograph of the Sessé & Mocifio type. Pringle's material was collected on hi on at a Mexico, ‘flowering in July and jasaat, fruiting i Paxton's discussion is es coe ae here: "Beautiful as are many of the innumerable hybrid Verbenas now in the market, eans c positively cumbered with names, commemorative only of the indi- viduals who raised then, rs the plants, in not a few instances, can scarcely be distinguishe "We venture these ae "not because we are averse to the practice of hybridization, for we greatly desire its more gener- al ado option; but solely to deprecate the best f a name and wo know several places in which this plan is judiciously followed, and trust, for the prevention of further confusion, it will be still more regarded in future. "Of the very ereite species herewith published, its appear~ ance is so far removed from that of all others yet introduced, that it is unnecessary to point out its distinguishing traits. respect, some resemblance will be a parent between it and V. tenet pesadees ao but this is of the slightest order, 1962 Moldenke, Monograph of Verbena 185 as the blossoms of V. teucrioides are particularly distant conm- pared with the present plant, and it wants those nunberless bracts which are here so conspicuous. Indeed, V. amoena is alto- gether more aig and interesting than the — above aaa! to, but is destitute of its sweet fragrance "Of its introduction to England, we have’ no very authentic information, We saw it first in the garden of the Horticultural Society, and fran some plants which were obtained thence by Mr. Edmonds a s wick, our figure was taken in the decline of last autumn. We have "It is fully as hardy as V. teucrioides, or other species, simply requiring a place ina a protected frame during winter, flourishing in the open border through the summer months. On ac- colours e Ra pee which have their flowers arran, heads, engthy spicate disposition of the blossoms by Which it is pecaltaciy characterized, This must be affected by id ion, and it is an object worthy of eels assiduous attention. Common abi sse a is performed in t u- si "The specific Shafinatton te “that by gate bo plant is own the Horticultural Society's Garden, and expresses its Particular "pleasing ve The Shove 3 hone 1 ae and ogres ea are repeated here as an the good writing which formerly “Teokine, ot scien- ° ly Pie pl ay aspect." He proposes "oh common name all, 6 ce rbar arium specimens and as mounted photographs and Siieeitat tans have been examined 186 PHYTOLOGIA Vol. 8, no. Cita : MEXICO: México: Pringle 13h3h (Au--181669, Gg— waa, ioeea506ia3); 13k34a (N, N—photo, Vt, Z—photo). MOUNTED ILLUSTRATIONS: Paxt., Mag. Bot. 7: 3. 18))0 (N). VERBENA ANDALGALENSIS Moldenke, Phytologia 5: ae “aren Literature: Moldenke, Phytologia 5: 2 eae 8. yeni aylor, Ind, Kew. Suppl. 12: 149. 1959; Moldenke, Lesions 26 & ii70. 1959. Perennial herb growing from a woody tapro sending up many stems from its apex; stems slender, often pte te: © th mostly erect, to 20 cm. tall, apparently much grazed and more “stubby; a internodes 5-~10 mm. long; leaves decus- sate-opposi ous, sessile, broadly linear-oblong, thin- uniformly gray-green 8 on 1l=-2 cm. long j peduncles very Slender nde cm. lone, densely pa ai nee a with white hairs; practlets very short, lanceolate, about 2 m. long, white-strigose on the back and long-ciliate along the margins, attenuate to a very sharp a- 1 ab ong . » wide, densely white-pilose, its rim 5-to othed; corolla hypocrateriform, bluish-rose, its tube glabrous, projec ting 1--2 mm. from the calyx-mouth, its limb 5--6 mm. wide. The type of this species was gollected by Pedro JUrgensen (no. 1613) at Pampa del Arenal, at an altitude of 2700 meters, Andalgal4, Catamarca, Arg dntten, in March, 1916, and is deposited ii the herbarium of the University of California at Berkely It s known thus far only from the type naam ol pf ae arium pa inclu the type, have been examin Citations: ARGENTINA: Catamarca: JOrgensen 1613 (ca198365— type, Z-~isotype). ve oe VERBENA ANDRIEUXII Schau. in A. DC., Prodr. 11: 553. Synonymy : Verbena andriaei ei Schau. ex Moldenke, Tis = "Met Invalid Names 7, in: T, in va a. 191. nt a Schau. - DC., Prodr, 11: 553. 181575 Jacks «, 1178, 1898; da Pa Mo. Bot. Gard. 20: 312, 333, ‘ "3c. 1933; Moldenke, Suppl. List Invalid Names 7+ 19111; Moldenke, Known Geogr. Distrib. Verbenac. [ed. 1], 18 & 101. 192; Moldenke, Alph. List Invalid Names 4S. 19423 he pre Alph. List Cit. 1: 11. 196; Moldenke ler 2: 363. 19473 He Me & A. L. Moldenke, Pl. Life 3s 48. 1948; Yoldenke, Know - Distrib. Verbenac., [ed 32 & i. 199; Moldenke ish, “List ae 3: 686 (2949) a se 1171. 1 5 Mondanke, 8. ian. branches opposite, elongate, cumbent, oi tetragonal, hispid; leaves sessile, about |, cm. long, cuneate sh 7 1962 Moldenke, Monograph of Verbena 187 2.5 cm. long, short-pedunculate, oblong, crowded, hispid, subglan- dulose; bracts subulate-lanceolate, one-third as "long as the cal- yx; calyx about 8 mm, long, the rim subulate-5-fid; corolla-tube limb appendages; cocci half as long as the c . The type of this species was collected by G. Andrieux (no. 138) -- in whose honor it is named —- between Puebla and Cnmnoa, Mexic 0, and is deposited in the DeCandolle Herbarium at the con- servatoire et Jardin Botaniques at Geneva. Material has been mis- identified and ig aaa in herbaria as V. aubletia L. and V. lamberti Ker. The Kraus © 31939 » cited below, has a label stating that the sient was "verwildert in Bot. Gart. Stras eae: The Barkley, Webster, & Paxson 857, also cited below a long 8 — it was collected in odk: dosdanidl Ser Wa ge aie. are in August In ala, 5 herbarium specimens and 3 mounted phototypes have been examined by me. Citations: MEXICO: Oaxaca or Puebla: Andrieux 138 [Macbride photos 7852] (Kr—photo of type, N—photo of type, N--photo type). on ints © otos{: Barkley, Webster, & Paxson 857 (Au). wk x f these @ 31939 (B). CULTIVATED: D: Germany: Golens yin) 57 (B); Gran Grantzow s sen. (Sept. '66] (B), sen. [Aug. ae R. A. Phil., Anal. Univ. Chile 43: 520--521. Synonymy: Verbena aurantiaca Ph., in herb. [not V. auranti- it... acl. terature: R. A. Phil., » Chile 43: 520-521. 1873; Jacks., Ind. Kew. 2: irre “1895; Reiche, Fl. Chile 5: 289 & 290. 1910; Moldenke, Known Geogr. Dis Verdenac., [ed. 1], 42 & lol. + 942s Moldenke, "Tilloa 8: 28 “G9i2) and 10: 361. 19h; Moldenke, Known Geogr. Distrib, Verbenac., [ed. 2], 101 & 197, 295 Woldenke, Alph. List Cit. 3: 687, 696, & 613. 1949; » Phytologia 3 493 Moldenke, Résumé 121 & i70. 29595 Noldenke, Résumé Suppl. 2: 5 & 10. 1960. hairy ( variab Sbovate, cuneate at the base, entire or trifid with the middle Segnent larger than the others, ovate, and sometimes also some- what parted, the lateral segments oblong-linear, the whole leaf Me: 4 cm, long, with the petiole comprising about this yb terminal, capitate, slightly separated from the eafy region; bractlets ovate, about 2 mm. long, sagt pa calyx 188 PHYTOLOGIA Vol. 8, no. } prismatic, 3--10 mm. long; corolla-tube slender, about 15 m. long; corolla-limb yellow, 5-6 mm. in diameter, composed of 5 cuneate divisions; anther-appendages thick, projecting et the throat of the corolla. hilippi remarks that he found only a single specimen of this plant, and that this was 15 cm. tall, but that it is cer- tainly a very characteristic species ["especie mui caracteristi- ca"], The type of this little-known species was collected by Ru- dolf amandus Philippi in the Araucaria region at Cupulhue, Chile, and is deposited in the herbarium of the Museo Nacional de Histor ia Natural de Chile at eee ean st species has been collected at altitudes of 1800 to 2500 me », blooming in Jamary. Herbar- ium material has been misident seed” and distributed as V. alter- nifolia Herb. and V. erinoides Lan. igs Macbride 343) photograph cited below shows 00 collector's or number, and the label is inscribed "Argentina indi- pod that the s specimen was in the Vienna herbarium aan it was photographed. In all, 2 herbarium specimens and 6 raetease photo= graphs, including a Phototype, | have been examin Citations: CHILE: Bio=Bio: R. A. Philippi s.n. rs, Herb, Mus, Nac. Hist. Nat. Chile 4273 & 54770] (N--photo of type, N-~photo of iso say Linares: Re Ae Philippi s.n. [Lin- ares; Macbride photos 1700] (Kr—-photo, N--photo . Santiago: Grandjot & Grandjot ret (N)3 Macbride Shotos 34344 (Kr—photo, W—photo) ; C. Reiche hk ( (N). ENA ARENARIA Moldenke, Phytologia 8: 59-60. 1961. Herb, apparently low and wide-spreading; stem much-branched; tivensbes slender, subtetragonal, more or less densely g ubescent and also whitish-hirsute with perpendicular ar hairs, less so in age; nodes somewhat annulate; principal internodes el 4 where, about equaling th calyx tubular, 5-6 mm. long, densely shent-qeibees ent ‘int reas sparsely hirsute with much longer divergent hairs, often somewhat glandulose; corolla hypo crateriform, blue, the tube about 10 mm, long, glabrous outside, the limb about » Wide. The type of this species was collected by Santiago Venturi (no, 2hhi) at Rio Lali, dept, Capital, Tucum4n, Argentina, along 1962 Moldenke, Monograph of Verbena 189 the banks of the river, at an altitude of 450 meters, on December 18, 1923, and is depo sited in the mn cdi Bee en a jum at Washington. son 2 specimens Citations: ARGENTINA: Tucum4n: Ventura Zbl (ieeliglason fetes Z—-isotype) . XVERBENA ARGENTINA Moldenke, Phytologia 8: 60. 1961. Herb, probably 4 natural hybrid between Vv. cabrerae Moldenke and V. calliantha Briq., with the tomentum of the former and the leaves of the Latter species; stems apparently sprawling or de- cumbent, root at the nodes, branched from near the base; te at petioles plaster: 3--9 mm, isha, rather sparsely strigose with rse hairs; leaf-blades chartaceous, somewhat lighter green beneath, ovate in outline, small, 1958 cm. long, 1? cm. wide, irregularly incised-laciniate or the larger ones trifid at the base, the lobes 2-toothed, rather sparsely short-strigose on rs ately narrowed at the base into the petiole; venation somewhat Prominulous beneath; inflorescence terminal, solitary, sareiet a ate, ely long ciliate on the margins from bi base to the mid-point; calyx tu- tube about 1) mm. long, glabrous reper the limb about 12 mm. The type of this hybrid was ote by Santiago Venturi (no. 1h) at Mocovi, Santa Fecino, Chaco, Argentina, on S , 1903, and is deposited in the eeiise States National Herbar- ium at Washington, ane 2 specimens, including the type, have been examined by me Citations: ARGENTINA: Chacos Venturi 1, (W--10),3597--type, Z~—isotype) . VERBENA ARISTIGERA S. Moore, Trans. Linn. Soc. Lond. Bot., ser. 2, be —— 1895. : S. Moore, Trans. Linn. + Bot., ser. 2, ks 439, 10953, Durand a Jacks., Ind. Kew. a wanes “Ls USL. 1906; ldenke, Known Geogr. pte Verbenac., [ed. 1] & 101. 19h2; 3 Moldenke Ai Lundell, Fl. Texas oa (ij: hi. 19/25 Moldenke, ea 3: 117 & 118. + Fe Moldenke, Alph. List Cit. 2: 552 (9k8), 3: 687, 705, 748, 761, & 916. (1949), and ks 1092, 1165, : "awe 1949; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 2 935 97, 99, 105, & 197. 19h93 Mesietiee, Pan Jo ogia 3: 135 (18.3) and’ 3: 289, 19503 Moldenke, Résumé 109, 115, 117, 119, 190 PHYTOLOGIA Vol. 8, no. — & —. 1959. 15--50 cm. tall; stems procumbent at the base, woody; ois Saeeian swollen; branches wide~spreading, assurgent or e- od tetragonal, about 2 mm, in diameter, obsoletely striate, final 3 simaruie s3 princi internodes });--l,.5 cm. long; leaves decus- sate-opposite, petiolate; petioles to 1 cm. long, subalate; leaf- ure, pinnatifid-trisect, to 3.5 cm. long and cM. wage strigose-pubescent with appressed hairs, the segments te, ne, aoctemate at the apex, often Fictice armen ae varying in length to to 1 cm., not over 1 cm. wide; inflorescence spicate, the spikes at first abbreviated, a- bout 1.5 cm. long and wide, finally elongating to 7 are densely flowered, ate gone-cavescetts peduncles about 1.5 m. young, later alonenting to 6 cm.3 bractlets rather Ie subu- late, ascending, + mm. long, prapeaig te eg ag oe tubular, tas ake about 6 mm. long and 0.7 mm. wide, narrow, 2=—3 mm. ing; corolla varying from blue or lilac to violet, pink-purple or purple, ane tube greatly surpassing the B.crlyts slightly over 1 cm. long, about 1 m, wide at the base and almost 2 m. et at wide, retuse at the apex; stamens short, inserted about § mm. a- . the base of the corolla-tube, the filaments slightly short- he acute anthers; anther—connective not appendaged; ea elongate, about 9 mm. long, much longer than the ovary, minutely pube than wide, longitu The type of this species was collected by wig 4 le Marc Moore (no. 1083) near Mt. P&o d'Assucar, between Coimbr4 and Rio Apa, in Matto G Grosso, sage and was deposited rks - herbarium of the Botanisches Museum at Berlin, now destroyed. The species d wet sandy campos, and sandy soil along rail Saiebes Srackivie curt ane See titudes of 250 to 700 meters, blooming from July to May. It is very closely telated to V. tenuisecta Briaq., it has been suggested that the ye are conspecific (in which case this name would replace that of Briquet). However, it seems to me that they are sufficiently distinct to maintain as ig trig taxa. Moore compares the species with V. laciniata (L.) Briq. 4 follows: "A V. crinoidi, Lam., cui sat similis ob calycis dentes longe ac patule aristatus sae antherarum connectivum eglandu- eg V. aristigera is called Nearaditai. and herbarium material wor 8 been misidentified and distributed as Be dissecta Willd. and - tenuisecta Briq. Anisits reports it as "frequent" at Estan- Sis Sate Maza, Paraguay, while Meyer as that it is "common" 1962 Moldenke, Monograph of Verbena 191 at Resistencia, Chaco, and vicinity, growing in campos and mat- to: 686 In all, O herbarium specimens and 3 mounted phototypes have been examined me. Citations: BRAZIL: Matto ph ogg 4 eS C. Hoehne, Com. Rondon 67 anos) (Es (Kr—photo of ant of type, W f£ type). : Oriente: Cardenas Es ( (N) « PARAGUAY : eter “3591 (KN, ae Haste « & Osten san. (Herb. Osten 806] (N, Ug); Hassler 1012 (N); A. Luts 150h (z); Malme s.n. [27/3/1903] (N, S); Morong 219° [Herb. Osten 13013] (Ug); 0 Osten en 9013 (Ug), 9016 (Ss, Ug); Te Rojas 153 (S), s.n. [Hassler 1602; Herb. Osten 8383) (N, Ug), SeNe . [Has- Sler 2650) (S). URUGUAY: Berro 4746 (N); Castellanos s.n. . [Herb. Inst. Miguel Lillo 15051] (N), sn. (Herb. Inst. . Inst. Miguel Lillo 15126] (N). ARGENTINA: Chaco: T. Meyer 292 (Ug—10954,); Venturi 9780 (N). Formosa: Ryerdam & Beetle 2293 (Ca-~-652375); I. Morel 70 (Gg—-353256, N), 1127 (N), 1305 (Gg—352672, N), 1760 (N), 1S he 2261 (N), 2605 (N); Pierotti 1176 (N), 4220 0 (NM); Re- ales 35 (N). > oem Reiche, Anal. Univ. Chile 123: 369 & 371. Synonymy: Verbena stacamensis Reiche ex Moldenke, Résumé 37h, in syn. 1959. eae Tee Oooo ab ae A. Phil Fl. Atac. hO. 1860; Re Ae Biel Vi- , 25, & 21h. 1860; Reiche, anal. Chile 133: 37 37. 1908; Specks. Fl. Chile 5: Prec cas ed prein, Ind. Kew. Suppl. : 245. 1913; I. M. Johnst. Gray Herb. 85: akpreaul « de Moldenke ®, Lilloa * p sei—322. e iplay Mo Known Geogr. Distrib. 1), 42 & 101. i Molden- od! tisha List ce taek 1: "261 (19 *Caoks) pe z'1116. 1949; Moldenke, Known Ge . Ver Tera og lea. 2], 101 & 197. ery Wol- denke, “Béowsé 121 121, , 3th, & Wats Perennial plant ola" ean: green, hispid; stems vari- seg , obtuse, entire or with a al lobule, roughened at the mar , With leafy — issuing their axils; inflorescence long- itate, but with » cap the lowest flowers somewhat distant; Seastiots lanceolate, reach- ing to about the middle of the calyx; calyx prismatic, about 6 short th; beet white to oe » mark projecting; The type of this species was “eakanved by Carl Friedrich Reiche in the southern part of the province of Atacama, Chile. Species has been found on rocky slopes and open west-facing 192 PHYTOLOGIA Vol. 8, no. Slopes of quebradas, at altitudes of 50 to 150 meters, blooming in tember and October. Worth & Morrison report it as "not com- mon", soe has been misidentified and distributed in herbaria as V. porrigens Phil. and V. sulfurea Don. Johnston (1929) says that the V. sulphurea and Ve . erinoides of R. A. Philippi (1860) are actually V. atac: ve. Sees: In all, 8 herbarium specimens have been Citations: * CHILE: Antofagasta: Wall & Sparre 789 (Ew); Nerder- mann 789 (Gg--1)30)1, N, S). Atacama: Werdermann 1632 (N, Worth & 1 & Morrison 161. (ca—630350), 16169 (Ca--6 30339) « VERBENA AURANTIACA Speg., Rev. Agron. La Plata 3: 563. 1897. La ta Literature: e, Rev. Agron, 3: 563. 18973; Speg-, Soc. Cient. Argent. 53: 249. 1902; Thiselt.-Dye: Suppl. 2: 191. 190k; Macloskie in Scott, » Princeton - Ex nk 191; Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1}, bh & 101, 192; Moldenke, Alph. List Cit. 1s Yl 2 19h6; Moldenke, Geogr. Distrib. Bora » fed. 2], 105 & 197. 19493 Mol— ree ——— 33 76 (19h9) and 3: 205 aed: rapyen ns mé ; & 170. 19593 tig Résumé Suppl. 21 5 & 10. 1960. er m pani at — stems moa branches ot = surfaces; inflorescence terminal few lateral twigs near the apex of the branches, fasc ate-subcapitate, rauner many=4 ; les slender, subterete, . long, very minutely and ig eg pub rulent; bractlets lanceolate-ovate, 2--3 mm. lon ong, e e base, SS the apex, strigillose on aie back, “ciliolate 21006 the ; lindric, about 1 cm. lo lainly 5-rib and deeply 5-sulcate, white-pilosul us ons passly on the ou » the teeth small, i subulate; corolla hypo crateriform, creamecolored or yellow, ite tube cylindric, about mm. long, glabrous on the outside, the limb about 6 mm. wide, bearded at the mouth inside anthers. AG ht exserted or sub- equaling the corolla-tube. : oe The species is found in dry sandy scrub and on ridges above the barrel-cactus zone, at altitudes of 2300. me ters blooming from November to January. It has been misidentified and distributed in herbaria as V. flava Gill. & Hook. In all, 1962 Moldenke, Monograph of Verbena 193 herbarium specimens have been examined by me. Citations: CHILE: Malleco: Behn 8031 (S). ARGENTINA: Q'Donell 31,80 (N, Tl). Mendoza: H. H. Bartlett 19410 icmapeneed, Neuquen: Senn 1326 (N). Santa Cruz: “Donat 219 nat 219 (Ca—L70k6h, Gg—- 194031, Go, N, N, S). sa” Mee es var, GLABERRIMA Moldenke, Phytologia 5: 3h1— 20 Liter. een enke, Phytologia 5: 341—3h2. 1956; Moldenke, Inform. me “set 51 Spec. . 1956; Moldenke, Résumé 126 & 4,70. Fe his variety differs from the typical form of the species in ae completely glabrous throughout. The corolla is described as yellow or lemon-yellow The type of the v: ariety was collected by Adrien Ruiz Leal and F. A. Roig (no. 15723) at ae ag Alto Valle del Atuel, at an es as of 2100 meters, Mendoz , Argentina, between January 9 and 17, 195, and is deposited ia th e H. N. Moldenke Herbarium at cig New Y ork, The variety hes been found by Ruiz Leal © at the edge of the crater of the Biamente volcano, at 1800 waters altitude. It has been found blooming in January and Feb- po ee hexbarium um specimens, including the type, have been takes ARGENTINA: Mendoza: Ruiz Leal 17000 (Ss); Ruiz Leal & Roig 15723 (Z—-type). ACA £. ROSEA Moldenke, Phytologia 5: 3h2. 1956. Liter : Moldenke, Phytologia 5: 32. 1956; Moldenke, In- oe, Wold. § Set Sl Spec. . 1956; Moldenke, Résumé 126 & 4,70. This form differs from the typical form of the species in having rose-colored corollas. The type of the form was collected by Adrian ped — (no. 14703) between Estancia Pfo-Pfo and Colonia Sarmiento, ora vadavia, Chubut, Argentina, on pepe at 1953, ary is de- Posited in the H. N. Moldenke Herbarium at Yonkers, New Yo Ruiz Leal also collected it along city sirests. sald by herbarium VERBENA AUSTRALIS Moldenke, Phytologia 2: h19--l20. Literature: Moldenke , Phytologia 2: 419-20. 19485 "Yolen, Castanea 13: 117. 1918; "Mo idenke, Known Geogr. Distrib. Verben- ; Kew. Suppl. 11: 262. 1953; Angely, Fl. Paran. 12: 17. maa uoldenke Résumé 109 & 70. 1959; angely, Fl. Paran. 16: él. b Herb with decumbent s tems, ending at their tips, not much Tanched; stems and heaiabea. Pepieing acutely tetragonal, sulca~ 19h PHYTOLOGIA Vol. 8, no. te, lightly rea se en eer with spreading or subappressed irs; principal internodes 28.5 cm. long; nodes annulate; Leaves oo sparse, maimezooncetiey sometimes with a few s in their axils; petioles slender, 3—5 mm. long, mar~ gined, ‘aodgilacens leaf-blades thin-chartaceous, rather uniform- ly bright-green on both surfaces, ovate in outline, 1--3 cm. and obscurely strigillose above, somewhat more densely so and punctate beneath, the very «gaa midrib and secondaries often slightly subimpressed above and prominulous beneath in drying; veinlet reticulation ergata on both surfaces; inflores- cence spicate, termi and e uppermost axils, densely con gested during anthesis, later e ating to about 3 cm.3 pedun- les slender, tetragonal, rather densely strigillose-pilosulous th v hort white cae hairs; rachis densely puberulent; bractlets lanceolate, 3-- about 1 mm, wide, acute a the apex, g exe fr “the white-ciliolate margins; tubular, about 5 mm. long, ely strigillose on _ Ba ng ribs or ’ glabrate, the teeth short, acute, not appendag branous and purplish between the ribs toward the apex; ete" tube about 6 mm, long, very sparsely and mimutely pilosulous on the outside above the’ calyx; corolla-limb about 5 mm, wide, very Dusén (no. 13190) in wet almost swampy soil at Ja Paran&é, Brazil, on October 10, 1911, and is deposited in the herbarium of the Naturhistoriska Riksmuseum at Stockholm. A a known thus far only from two collections all, 3h barium e pineam 2 oe the type, and ), mounted photographs have been by m Gitations: BRAZIL: Parand: Braga, Moreira, & Lange 328 (2); Dusén 13190 (F—photo of type, ie-isctyps, N—photo of "type, S— type, Si-photo of type, Z—photo of type). XVERBENA BAILEYANA Moldenke, Phytologia 2: 235--236. aais Synonymy: Verbena tetrandra, spicis longis acuminatis, foliis multifido-laciniatis Haartman, Amoen, Acad. 3: 43. 7B. Verbena cad. 33 ° H berts, » Hybrid. Before Mendel 2h, 1929; Moldenke, Phytologia 2: 235- 238. 19175 ‘pide Alphe L List A ope — $ Suppl. 1: 2h & 26. 19bTs HN. & A. ent 19185 Moldenke, he List cit. 21 £356 56 (1948 and ht 1232. 11949} oldenke, Geogr, Distrib. ed. 2], 163 & 4ou95 Woldenke, Phytologia 3: 66 & Lev, “4961; Moldenke in "nittenden, Roy. Soc, Dict. Gard. 6: 2209 & 2210. 1951; E. J. Salisb. inte Ker. Na Moldenke, Résum6 222, 365, 366, 371, 374, 376, => 470. 1959+ Herb; stems Pci branched rather s harply t rag onat, 1962 Moldenke, Monograph of Verbena 195 cate between the angles, strigillose with short, stiff, white, antrorsely subappressed hairs 8; nodes annulate; principal inter- nodes 2.5--5 cm. eens leaves decussate-opposite, usually with several small ones on very abbreviated twigs in their axils; pet- ioles obsolete or fee 2 cm. long and broadly winged, merging in- st 1 lades c us above when the finger is drawn from the a apex toward the peoey midrib slender, impressed above, prominulous beneath; secondaries nent, L or 5 gularly bran reticulation subimpressed above, plane but visible beneath th; in- florescence a terminal panic le, the lowest branches of which are axillary to the uppermost much reduced leaves, the panicle about 15 cm. long and 5 cm, wide, its branches erect or ascending, strigillose-puberulent; bracts lanceolat te, 5--8 mm. long, dense- ly atrigtl inet, bractlets similar but smaller, slightly shorter than the » densely al in the Jardin rr Plantes at Paris in 1819 and ve deposited the Dudley Herbarium at Stanford University. It is spetently a hybrid raeoeae wee piri esealie L. and V. hastata L. acl with, general, haracters. It is named in honor of Day Liberty’ oven paikey ( 1858--1951), a as amar pas crepe — — expert on Carex, Rubus and palms, kindly gentleman end The hybrid is said to have been growing in Swedish gardens as mond as 1748. Normally the two parent species would seldom be growing in close proximity to each other -- one being Euro- eae and the other North American. The former, however, has been introduced and grows as a weed in many parts of eastern ern North An- erica where ¥. hastata grows padecetig, and the latter is some- times cultivated (and even naturalised) in Europe where ¥. offic- inalis is very common. The supposed hybrid shows a marked domin- ance o of V. nastata characters. The Austrian specimen cited below Haartman states in — Acad. 3: 3 (1756) that this hybrid originated naturally in the Botanical Garden at Uppsala, pres "bu semi no one had the seeds here hitherto, nor through a planting of it, since it had not previous sly been seen within the country." In’ all, 3 herbarium specimens, —— the type, and 3 moun- ted photograph¢ hé have been examined by me. 196 PHYTOLOGIA Vol. 8, no. k Citations: CULTIVATED: Austria: Herb. Reichenbach f. 160270 (N, V). France: Herb. W. H. Harvey 5.n. gen. {he Re Fe 1819) (Du-~ 1664,38--type, F--photo of type, N—-photo of type, Z—photo of type) « VERBENA BAJACALIFORNICA Moldenke, Phytologia 2: 22-~23. 191. owes VE ae seas Phytologia 2: 22--23. mp eee Known G . Verbenac., [ed. 1], 18 & 101 (1942) and fed. 2, 50 ro 97. 15195 Moldenke, Alph. List Cit. 3: 933 oe 12kh 3 Shreve & Wiggins, Carnegie Inst. ht & iz? 1951; E. J. ena Sg Ind. Kew. Suppl. ll: Kee ago: scattered, stiff, whitish, non-glandular hairs about 1 mm. long, glabrescent in age, sesger ag decumbent at the very base and out roots the lower nodes; leaves secusente opposite, petiolate; Petbeben distinct, very slender, 1--10 m. a densely or sparsely hirsutulous with stiff, white » non- & surfaces, more densely so along the midrib and larger veins be- neath, abundantly pinnatifid-incised, sometimes obscurely with the divisions again abundantly pinn nnatifid-incised, and even after anthesis or the 2 or 3 lowermost to 5 m. 4 in fruit; bractlets very small, lanceolate, 2--3 mm. re = half the length of the calyx, attenuate at reer except for the long-ciliate margin; calyx t , b- 5 mm. Lon, projecting from the calyx, its tube slightly puberulent at the ue 1059, in syn. 190) [not V. thymoi-~ des Cham., 1832 “Literature: Chod., Bull. Herb. Boiss., sér. 2, 2: 818. Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8: 293-29), ipdks Me in Chod. & Hassler » Bull. Herb. Boiss., sér. 2, hz 1059. 190k; . & Hassler, Plant. Hassler. 10: 481. 190k3 Briq., nke, Known Geogr enac., on 1), » 41, & 101. 1942; Moldenke, Alph. List invalic ane Sl. 19423” Be Alph. List Cit. 1: 26 & 26h. 1963 # enace, ed. al» 93 Phytologia 3: 288. 1950; stellfeld, Trib. Farmac. 19 (10 ie peta, ge aaa ll: 57 & 13h. 1959; Moldenke, Résumé 109, 117, 119, 126, 376, & 470. 19593 Moldenke, Résumé Suppl. 1: 7 (3959) an and 5, 5.1 360; Angely, Fl. Paran. 16: 78 (1960) and 17 Pe eotis herb or subshrub, to 50 cm. tall; caudex rather thick; stems erect, virgate, thick, simple or with elongate- ascending branch hes, tetragonal, shortly substrigose-canescent pal internod won Easeicpe unculate, few-flowered, subcapitate during anthesis, out, with no or very sparse glan e about 5 mm, long, the teeth 5, une lanceolate-subulate, about 0. - lo qd anthesis, but to 1 ate after anthesis and then connivent; corolla varying from lilac or blue to white, Blabrous outside, the tube oe by the calyx, the limb about 4mm. wide, the lobes sub subequal, obovate, emarginate at the apex, barbellate—villous with blue = Me a the throats; stamens 5 or 4 198 PHYTOLOGIA Vol. 8, no. h didynamous, ee style included; ovary l-celled; cocci ea peiegin| yellowish, to 3 mm. long, with elevated reticulations on the outside. types of this species were collected by Benedict Balansa (no. 1028) —- in whose honor it is named -- on the campos at Caa- din collection has blue flowers and the latter white, so if th white-flowered form should be separated, 1028 would become the type of the species and 1163 would become the type of the white- flowered form. This handsome species is very closely related to V. : on Cham., which is also characterized by leaves tripartite to the very base, ee ae ecies are easily dist ata whe length of the calyx. In V. thymoides it is shortly campanulate-tub and not over m, long, including the teeth, while in V. balan- sac it is 6—7 mm. long. Briquet verified this by examination of an original ~ of V. thymoides in the DeCandolle Herbarium. He remarks that . thymoides seems to have a low prostrate habit, while V. Peis is erect, erect, but admits that he is not sure 0 of what value this character is in view of the wide vartabey an characters — in no, 1028 the leaves are "courtes appliquées et trés grises", perhaps due to its being a sunshine form, while in = So they are "deviennent plus étalées, plus longues et plus vertes", perhaps ae to it being a denizen or inhabitant of less arid situations. He cites Lindman A.1189 as "Exp. 1. Regn. A ay par et ne nous paraft pa pouvoir @tre réuni avec le V. thymoides Cham. & cause des dimensions du calice, yt — "état actuel de nos connaissances." Apparently due to typographic errors, Briquet, in his original description, says that the leaf-segments of V. balansae are "0,2—1 cm." wide and the cocci "ad - cm. altae." Doubtless "mm," was sctenios in both instances. Ve balansae is said by collectors to inhabit campos and high ary campos, fields, dry grasslands, and rocky places, blooming campos", Vernacular name " n, "formosa sem dove”, and " a", It has been widely misidentified and “aistributed in herbaria as V. thymoides, while the Hassler 4951, determined as V. balansae in the Britton Herbarium, is is something non-verbena~ ceous, The species has been recorded from Uruguay on the basis “ Osten 22375, but this collection appears to be ve thymoides te albiflora Moldenke instead. In all, oP ae apnea and mounted photographs, including 2 of c manineds Citations: BRAZIL: Parand: Dusén Ss is. °3 190k] 8) pie 1962 Moldenke, Monograph of Verbena 199 [Herb. Mus. Nac. Rio Jan. 16565] (N, S), 7692 (S), 7971 (S), 10467 (N, 8), 10847 (N, N, S), 16907 (S), 17923 (8), sen. [17/2/ 11] (S); Hatschbach 253) (N), 2765 (N), 5467 (Z). Rio Grande do Sul: Lindman A.1189 (N, 8); Malme 916 (S), 96a (S), 9h6b (N, 8), gbc (S)3 As Re Schultze 91 (N). Santa Catarina: Reitz 2% 0.883 (N)3 Tle 1175 [Herb. Osten Osten 13010] (Ug). gpa: Be Balansa 1028 TWacbride photos 21,680] (Kr——photo of cotyp --photo of if eateeae) ; Hassler 343 (Mi), 4640 (N), 7104 (N), ie {cb, N--photo photo, V, Z= Z——photo, ~Z——photo) 3 3 Jorgensen 1582 {Herb. Osten eee (N, S, Ue) « ee Corrientes: Pedersen en 2906 (N, S). Misiones: Eman 2021 (Mi, Dense ne mci amas vaca at the base, forming d tufts to 1m. in diameter; stems simple or very sparsely cot § 15--22 cm. long, densely retest throughout nodes annulate; ous, cuneate-obovate, 1--3 cm. long, 1--2.3 cm. wide, rounded in ou at the distinctly 3-lobed, long-cuneate to the base, dense rt-pubescent with brownish or grayish often glad-tip hairs on both surfaces, so in age and then often the escence beneath limited to the larger venation, each lobe often again 3-lobulate, the lobes oblong and blunt; ies impressed above, promin- ped Corolla rose or creamy-pink to blue, its tube about 1 cm. long, the Pape about 6 mm, in diameter. 7 apt of ho showy and most distinctive species was col- se JW Panand « Balls (no, 6036) -- in whose honor it at oes on dry s exposures in shaley formations welt Site ene nad moisture es tn the Chabet oe near Tilcara at an altitude of 12,800 feet, Jujuy, Argentina, on February 13, 939, and is deposited in the United States ws oan Herbarium at Washington, It has also been found in mead quebradas, blooming in F February and March, Apparently it grows at altitudes 200 PHYTOLOGIA Vol. 8, no. of 3000 to 4265 meters. In all, 6 herbarium specimens, ae the type, and 2 mounted photographs have been e Ce Citations: ARGENTINA: cA Balls 6036 (Gan-€83618-—teotzpe, N--photo of type, eggs Z—photo ype). Sleumer & Vervoorst 3006 (N, x, 7422173298) ; antes én am 1591190) . VERBENA BANGIANA Moldenke, Sponge 3: 63--6h. 199. Literature: Moldenke, Phytologia mi 63—6h. ini MeLdenke, Known Geogr. Distrib. is. [ed. 2], 97 & 197. 19,9; E~ Je Salisb., Ind. Kew. Suppl. ll: 262. 1953; Moldenke, Résumé 115 470. 1959. Herb; stems and branches rather oo tetragonal, scattered- pilose, more densely so on the youngest parts, often more or less hollow and sam deeply sulcate in drying; nodes annulate; princi- pal internodes 3—10 cm ones 2 — decussate-opposite; pewted les 5-10 mn. ragesise broadly wi ed and merging into the blade; leaf-blades chartaceous, fathead uniformly bright-green on both surfaces, — or slightly obovate, acute at the apex, petal spar’ slender midrib and 6—-8 secondaries subimpressed a promi= inent beneath; larger parts of the veinlet reerrarsrt ie often m™m ; parts of 1—7 cm. long, pilose; floriferous portion of the spikes short dense-flowered, 2--6 cm. long, pen mmpoionay pteenyl bractlets oblong-lanceolate, about 5 mn. long, acumi a apex, ciliolate-pilosulous, or the Séniet pair to 2 cm. pA and contato, SPophcciates cofelis-tats barely surpassing the CHl7% pos - sing the ¢ its limb about 1.5 a a 18 Pere ee The type of this s a em a collected by Henry Hurd Rusby (no. 911) at Sorata, at an altitude of 8000 feet, La Paz, Boliv- honor of Miguel Bang, ~F collected so extensively in Bolivia for Dr. Rusby, The species resembles V. litoralis H.B.K- in gen eral habit, while its sot maaiC re Yesemble those of 7. ie pida Ruiz & Pav, It may possibly prove to be a natural hybrid between the two, the same as the hybrid produced artificialy by Dermen and named xV. bealei Moldenke, the type of whic not as yet seen. V. '. bangiana is known thus far only from the type reerrpeitigier. of which 31 3 mounted specimens have been examined. Citations: BOLIVIA: La Paz: H. H. Rusby 911 (C-type, Paes: isotype, Preciadtiie’ e 1962 Moldenke, Monograph of Verbena 201 VERBENA TA Grahe, Edinb, New Philos. Journ. h [Oct .-Dec.]: tei. 1827. ure: ahe, Edinb. New Philos. Journ, [0ct.-Dec.]: 6-177. 18273 Sweet, Hort. Brit., ed. 2, 18, 1830; Walp., Repert. ): 33. 1845; Schau. in A. De., Prodr. ll: 555, 187; acks., Ind, Kew. 2: 1178. 1895; Perry, Ann. Mo. sd bs ghomny Cie st spreading hairs; leaves petsatsts, Sopte site, decussate, spreading, cordate-ovate, reticulate-v eined, pubescent on Mhgpes surfaces, the margins rather unequally crenate-serrate; sp oli- bay Sagano bractlets subulate Figg than oe short pedicel; han solit alyx g the 1 Denirtts of the bractlets, pebestteh “With erect ialce: corolla pale-pink, ular, pubescent with reflexed hairs, its tube twice as long as the calyx, the limb erect; anthers included; filaments inserted on the Shexiinciebar style filiform; stigna hooked, exserted just before the bud fully expands, but after- and rig many stems, without being farther divided, it is possible that both the woody structure, and the branching, may have ates says from the leading shoot having been cut down." He r. hat "The species has no beauty, nor does it possess any interest Sxcept that it is new. We received the plant from Mr Hogg at New York last s ; er no name, but with the information that it had been procured from Mexico." Sweet calls it the "Bearded Ve rry says "This tedly a member of the Section Verbenaca, but the debsriphine is too meagre to identify it." She suggests bagesieg that V. delicatula Mart. & Zucc. may be the same taxon Weldon, Chittenden, Roy. . Dict. gard. 6: 2210. 1951; ges ; YP Midl.’Nat. 59 "32. 1988; Map ebay Résumé 223, 366, sp ibisstraltone Dermen, Gytologia 7: 163, fig. 21, & 169, fig. This is a hybrid between V. hispida Rufz & Pav. and V, litor- alis H.B.K., with intermediate characters. It originated in a garden in Massachusetts in 1936, having been produced artificial- 202 PHYTOLOGIA Vol. 8, no. ly by Dermen in his experimental work on the yes espe ee? of Ver- bena. There is plenty of opportunity for this hybrid to occur in Nature because the two parental species occupy the same vereialbee: ic areas in Paraguay, in S8o Paulo, Brazil, and in at least three orms ¥. litoralis seen in rtabberte secirivent this hybrid. It is also very possible that ve. bangiana also belongs here. V. litoralis is apparently very widespread and variable; it should | receive much more intensive study, and when this is done the pone of hybridity with related species should be kept in VERBENA BERTERII (Meisn.) Schan. in A. DC., Prodr. ll: 551. 18h7. Synonymy: Verbena erinoides Spreng. ex Hook, & Arn., Bot. Beech Voy. 41-2 (1830) and 48h. 1841 [not V. erinoides Au th., 1959, nor Chod., 190), nor Lam., 1791, nor Willd., 1947]. Shuttleworthia berterii Meisn. ex Walp., Roper’ h: 13 (1845) and Meisn., Gen. Pl. ere 198. 1846. Verbena erinoides Poepp. ex Schau. in A. DC., Prodr. 11: 551, in syn. 187. Schuttleworthia berterii Meisn. apud C. Gay, Hist. Fis. Chile Bot. 5: 9, in syn. 1819. Verbena erinoides L. ex Lorentz & Niederlein, Exped. Rio Negro 2 (Bot.): 266. 1881, Verbena berterii Schau, apud Jacks., Ind. Kew. 2: 1178, 1895, Verbena berteri Schau. ex Hauman-Merck, Anal. Mus. Argent. Hist, Nat, Buenos Aires 2h: 415. 1913. Verbena erinoides Hook, ex Moldenke, Lilloa 6: 332, in syn. 193). Verbven ‘Verbena dissecta Schau, eterna Boe sagen 363, in syn. 1959 [not V. % Ve dissecta Morong, 190k, » 1847, nor Spreng., 1959, nor bind? 1849, nor witid., : 182), * verbena erinoides Hook, & Arn. Mol- denke, Résumé 36), in a Literature: Feuill., + Obs. Phys. Cotes Orient» [3]: 35» pl. 25, fig. 1. 1725; ook & 2 "arn., Bot. Beech. Voy. ae 1830; Maund, Bot. Gard. 5: pl. 106. 1831835; Hook. & ies Meisn., Gen. Pl. Comment, 198. GREE esheld.- sh . m ‘ br. Pro i - Us 551. 1847; Cc. Gay, Hist. Fis. cate’ Bot. 5: past eas, 18195 ke, Bot, , 106s 162, 1945; Moldenke, Alph. List Cit. 1: 39 & 19,8; denke, Ple denke, Alph 113 28 i bes (1949) and h: 1017, 1078, 1116, 1175, 1187, 1215, 1 1297, 1302, & 130k. 19h9y befdenke’” sreesaeene’ 3+ 7H’ (A999 and 1962 Moldenke, Monograph of Verbena 203 3: 290 & 377. 1950; Moldenke, Sp. Subsp. Cont. Mold. Set }5 (3]. 1951; Moldenke in Chitte enden, Roy. Hort. Soc. Dict. Gard. 6: 2 a orm. Set 51 Spec. 4. 1956; Moldenke, Résumé 84, 121, 223, 358, 363, Hh, & W70~ 1959; Woldenke, Résumé Suppl. 2: 5 & 9—11. Dlustrations: ere Bot. Gard. 5: pl. 106 (in color) [as vy. radioans]. af 3 835. ~~ Anmal o oe procumbent or prostrate herb, often slightly woody or it tite at the base; stems red, creeping, spreading, ascending at the tips, covered with short white hairs; branches as ascending, tsttactint: whitish-hirtous throughout; leaves decussate-opposite, petiolate, pinnately or bipinnately parted tri innatif and entire a r fid-p » attemate the base, strigose-hispidulous on both surfaces » the anceolate to a or oblong-lanceo » short, rather the branches, eet not enlarging’ after anthesis; bractlets lanceolate or linear-lanceolate, 1/2 to 2/3 as long as the calyx, te-hirsutulous or hispid-canescent, the teeth unequal, linear, rather obtuse; corolla hypocrateriform, 2=-3 times as long as the calyx, varying fron yellowish, pinkish, or rose to light lilac, The type of this a was collected ep entie moa Ber= tero (no. 7) -- in whose honor it is named -- in Col A Chile, and is deposited in the Meisner Herbarium at the New York Botanical Garden. Schauer cites also Bertero 1390 from (Colchagua, Chile], Poeppig 159 from Concan (Valparaiso, Chile], Seudichad s.n. Sen. and Style s.n. n. from at and Filter s.n, from Herbarium at the Conser- taniques at Geneva. ag Lilloa 6: is (1941) I cited eatong 1322 and United States Exp Exploring Exped. [Wilkes] s.n. from Valparaiso ~ Bridges ¢ idges 602 from Valdivia, but these collec-_ tions, on furthe study, ha’ ve proved to be other species. a °ere Distrib, Verbenac. and in my Résumé » Peru, rror. It is said to be cultivated in England, but I have as yet 20h, PHYTOLOGIA Vol. 8, no. seen no cultivated material of it. The species is said by Schauer to be closely related to V. lipozygioides Walp., but is very distinct in the character of its corolla (which he says is apparently whitish) and anthers. He al- so compares it with V. ines se tM pointing out that it dif- fers from the latter in 1 flower, and indumentum characters. sed pubescence, rose ead of yellow flowers, and above all in its anther-appendages being not visibly exserted. He notes that "El sefior Schauer le da ademas como carActer propio el de tener el caliz contornado sobre el fruto, pero esta particul aridad sé observa Saabien: on las V. sulfurea, erinoides, etc." Pa as material has been abundantly so aoe and dis- tributed as ve erinoides Lam., V. laciniate (L.) Briq-, Ve and therefore cannot say if identified. Lorentz & Niederlein (1881) phy saeeee 4 aieeas V. erinoides Le] ies om Rio Negro, but they may be referring to V. laciniata (L. ) In all, 98 herbarium specimens, including the types of ees of the nanes involved, and 5 mounted photographs and illus tions have been exam by me, tations: PERU: Ayacucho: R. D. Metcalf 30322 (W--1834982) « Huancavelica: R. D. Metcalf 30267 ’(W—1034979)» CHILE: Aconcag- ua: E. M. Kausel 2558 (N), sen. (Zapallar, 10.x.1948] (N); Looser 550 (N), 5505 (N)5 Schwabe 120 (N). Antofagasta: ra Valenzuela le £005 (N)3 Tenvermann meres 708 . (Ca--289 369) « Bio-Bio: DUSt Dusén 7853, in ah (Sartre of fit Sa W--type, N—photo of ise ype, Pp @), 8.n. [Racangua] (Br) ‘ Gal gpchae undesignated 2 2 (N)e aes cepcion: Barros Valenzuela 8029 (N N)3 J Junge 971 [Herb. Jard. Bov- Rio Jan. 26790] (N); Skott matte & & Skottsberg 1491 (Go, S)- 1962 Moldenke, Monograph of Verbena 205 quimbo: Biese 2238 (N); Skottsberg & Skottsberg 843 (N); Worth & Morrison 1660 (Ca—631791). Curicé: Barros Valenzuela 8036 (N). Maule: G. Kausel s.n. [Constitucién, 2--9.2,19]9] (Ku); C. Reiche 9 (N). Nuble: Barros Valenzuela 805) (N); Puga 8 (N). Santiago: Biese 21 (N, W—1931303); C. Gay 108) [18; Herb. Mus. Nac. Hist. Nat. Chile 5473)] (N); G. T. Hastings 190 (Ca—66365); E. M. Kaus- el 306), (N), 3147 (N); Looser 222 (N), 5507 (N), 5509 (N), 5511 (N); He N. Moldenke Sen. [Kausel 2679] (N); Moldenke & Moldenke 19758 (Es, Es, Lg, Mg, Mr, N, No, Ot, Sm); J. L. Morrison 16771 : (Ca--632907); Re A. Philippi 73 (S); Skottsberg & Skottsberg 975 (Go, S). Talea: Collector undesignated 7 (N). Valdivia: Bridges sn, (Valdivia, 1862] (i). Valparaiso: Barros Valenzuela 8036 (N); K. Behn s.n. (1) Sept. 1930] (ca~-L],6699), sen. [3-X1-190] (N); Bertero 1389 [Macbride photos 7853, in part] (Kr—photo, M, N--photo); Bridges s.n. (Br); Gtmther & Buchtien s.n. [IX.1923] (Ew, S); Killip & Pisano 39710 (N)} Moldenke & Moldenke 1976h (N), 19936 (B, Bs, Es, F, Fy, Hk, Hw, Le, Lg, Lm, Mg, Mm, Mr, N, n, [Valparaiso] (T). Province undetermined: Bridges sen. [Chile] VERBENA BERTERII f. ALBIFLORA Moldenke, Phytologia 3: 278. 1950. Literature: Moldenke, Phytologia 3: 278 & 290 (1950) and 5: 964 195k; Moldenke, Résumé 121 & 70. 1959. his form dif. from the typical form of the species in on the western slope of the Cuesta de Zapata, Valparaiso, Chile, on November 3, 1948, and is deposited in the Britton Herbarium at Gard VERBENA BINGENENSIS Moldenke, Phytologia 2: 145--1h6. 19h6. 206 PHYTOLOGIA Vol. 8, no. Literature: Mol » Phytologi 145--146. 19163; Moldenke, Alph. List Cit. 2: ae (298) | and + “pe. 19493; Moldenke, Known strib. Verbenac., [ed. 2], 26 & 197. 1993 us as Salisb., Kew. Suppl. 11: 262. "1953; Moldenke, Am. Midl. Nat. 59: 3h2- 313. 1958; Relieee, Résumé 32 & 70. 19593 Moldenke, gh ci Suppl. 2: 10 a thanatied herb; stems medium, slightly woody at the base, obtusely tetragonal, brown, often blotched, rather abundan more dense te-pubescent with soft hairs; padi: ascus ee ate- oppent +83 ssisatée short, 2—5 mm. long, winged, in et he blade; Leaf-blades firmly aceous, brittle ove (when the finger is dram d and slightly so wat ovate in outline, cm, long, mostly about 1—1l, he largest to 3 cm. wide at the Paros lobe~like and spreading on the largest leaves, abundant ly strigose-pubescent on both surfaces “ the hairs very variable in length beneath and densest on the venation; venation impressed branched, in the axils of all bes upper leaves; peduncles slen- der, 1—5 cm, long, sharply tetragonal, densely short-pubescent; rachis slender, densely pibescent with rather stiff forward- pointing white hairs of vari ous lengths; spikes to about 10 cm. long, seh many~flowered, apparently setting seed very poor . ma spicuous, very variable in size, --8 mm. long, lanceolate, the lowermost slightly foliaceous, attermate-acuminate at the apex, out 1 mm. wide at the base, more or less es not keeled calyx; calyx 3—l m. long, densely strigose; corolla-tube slightly surpass~ ing the calyx, densely puberulent on the outer surface, its limb about 2 mm, wide, The type of this natural hybrid was collected by Wilhelm N. Suksdorf in bottomlands at Bingen, Klickitat County, Washington, on yon 9, 1898, a is deposited in the Britton Herbarium - the New York Botanical arden, It was identified originally 4s officinalis L. The a appears to be a natural hybrid petteok v. Lage & Rodr. and V. lasiostachys var. $ tentrional- is Spey The two parental taxa Be gated in at least 9 counties of California, but thus far only one supposed parent, ve caaeata, has been seen by me from Rickitat County, Wash- , where Suksdorf collected the type. This fact militates against the hypothetical second parent, but possibly pre Sayer sive collecting may reveal its presence there. Suks was 4 very intensive collector and one would rete that is youd have collected material of a parents if they occurred t in the bottomlands at Bingen in 1898 when he iva the "ype of the supposed hybrid, The hybrid nature of this plant, there 1962 Moldenke, Monograph of Verbena 207 fore, and, if it is really a hybrid, ae owe by identity of the second parent, are still in doubt. An experimental geneticist might try to make the suggested cross ae see if the result re- sembles this plant. It is know thus far only from the type col- lection, of which E, herbarium specimens, including the type, have been examined by Citations: WASHINGTO TON: Klickitat Co.: Suksdorf s.n. [ July 9, 1898] (Ca—755099—-isotype, N--type, ae ER Sree . ee wigs pps Nutt., Journ. Acad, Nat. Sci. Philad. 2: 23. 1821 Synonymy: Glandularia eee Rei & ) Nutt., Phil. Soc., new ser., 5: 104. 1037- sipimmatitis ylang Schau, in A. DC., Prodr, 11: 553. Fee Sirs bipinnati- fida Nutt. apud Schau. in A. DC., Prodr. 11: 553, in syn. 1647. Verbena bipinnatifida Schau, apud Jacks., Ind. Kew. 2: 1178. 1895. Verbena pinnatifida Schau. ex Briq. in Engl. & Prantl, Nat. Pflanzenfam. }; (3a): in nota. 1895 [not V. pout Lat., 1791, nor M. E. Jones, 19h]. Verbena demareei | , Am. Midl. Nat. 2h: 752, 1940. Verbena bipennatifida matt. ex Moldenke, Suppl, List Invalid Names 7, in syn, 191. Verbena bipinnalifida Nutt. ex Moldenke, Suppl, List Invalid Names 7, in syn. 191. Verbena bipinnatifida Michx. ex Moldenke, Suppl. List Invalid Names Names 8, in syn. 1941. Verbena bipinnatifidus Nutt. ex Moldenke, Suppl, List Invalid Names 8, in syn. 191. Verbena pinnatifida Nutt. ex Moldenke, Suppl. List Invalid Names 10, in syn. 1941. Verbena bipiinnatifida Nutt. ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syn. 1947. Verbena artemisiaefolia Clemens ex Moldenke, Résumé 358, in syn. 1959. Verbena bi-pinnatifida 1. & Wright, N. am. peek ot a) 4,70. 180; Walp., Repert. k: 1815; Schau, in A od. 11: $53. 1875 A. Grey, Sym, Fi. Ne Am. 2 (1): 337, 1878; Wats. & coult. An A - 6, Neg Penge Coult., Contrib. U. S Na t. ; “i Bibs Hook. Hook. e, Ind, Kew. 1 (2): 1032 (1893 paces 953 Hi : papas” Bull Ss. Me bile 32 ( 57: pl. 13, Se 140 (dase) an and 66: pl. 12, fig. 1h0. 1897; Britton & Br., rags r. Fl., ed. 1, 3: 72, fig - 1898; A. S. Hitchc., Fl. Kans. pl. 15. 1899; , «, ed. 1, 1010. 1903; Briq., Ann. Con- serv. & Jard, Bot. Genav. 10: eee Robinson & Fern. in A. Check List 128. 1908; Britto 7 Bre r. Fl. , 32 97, fg. 3559. 19 in, Fl. Bre ; san 101d. 1913; - De 00 Wild Fls. 177--178. 1922; Olmstead, Co Kelsey, Stand. Pl, Names, ed. 1, 521. 2924 Ee Ds Schulz, T cy, 3 :. exas Wild Fis, 335-3. 1928; Seymour, Host Ind. Fungi N. Am. 587. 1929; Rydb., Fl. Prairies & Plains 678--679. 1932; Perry, Ann. Mo. 208 PHYTOLOGIA Vol. 8, no. Bot. Gard. 20: 2h3, 2hh, es a, oe 3255 328, 331, Se : 305. 1993 E, J. Palmer > Journ. : 133. ig93h; iley, Hortus, "new reve ee OG5ins63, 19353 L. i, gE Cate vinweite mss. 19353 Eo Je Palmer, Ann. Mo. Bot. Gard. 22: 629. 1935; red Texas Agr. Exp. Sta. Bull. 550: 88. 1937; Mol- enke, t 108 & 113. 1939; Eee ida 4 List Common coi! a «1905 wotdente, Prelim, Alph. List Invalid Names 26. 1940; G - 190, map 1066. 19)0; Fl. Indiana 795, 1083, "2 1232. “1910; Moldenke, spe rs t Tavalid Names 7, 8, & 10. ila; vege Weeds Kans. li2 & 2 + 1941; Moldenke in Lundell, Fl. Texas 3° (2): 18, 38--h0, & 2. 19423; Moldenke res Geogr. Ee woud eee [ed, 1) L—6 —6, 8-12, 1h, 18, 15. 2: 70, 71, 71, $ Lh, & niraey 19S; Moldenke, Bot. Gaz. 106: 161. 19h; Motibeice , Castanea 10: fo. 1955 5 Ados ato, 189; 0, 2 3, uth, 184, 157-169, 161, 162, 166, 17h, 178, 163, 193, 194, 97? 199—202, 20h, 209, 211, 213, 217, 225, 226, 231, 20, 2ha ahh, 250, 255--257, 267, 270, Jt 280—282, 288, & 297. 19465 Merr. & Reeder, Bartonia 2h: . i9h6; Tolstead & Hgts shige 1 » 1hs 39 60. 196; ey Alph, List Inva 1: 22. 9h7y Moldenke, Phytologia 2: 328, 329, a7, 7 28) ‘Gein : 159--160, 19483 H. Ne & A. L. Moldenke, Pl. Li 55 , 19185 arg Wrightia 1: 231—232. 198; “toudone, ‘caste :112& 1948; Moldenke, Alph. List cit 356, 591, 393, 397, 38, hob—hO7, 139, 450--L55, L57, 170, Wt2--h76, 522, 525--527, 529, 536, 538, Shs any zi8 260) sur-238, 574 > page 0 9; 3 , ots, 281, 585, 588, 593, iy 6--598, 60h--607, 617, 621, 630-- 69, & 6hO (1948), 3: 66 166 prey 67h, 678—663, 698, 5 apa o2, 723,77 7 3 1, 1183, 1187, 1191, 11 1202, 1205, 1207, 1210, 1% as? 1217, 1220, 1221, 1225, 226, 1229, 1230, 1233, 1237-1237 ys p27, 1260, 1283, she" 1290, 1292, & 1296. 19h9 % st. Molde gre « Ver > [ede 5, 2 163, & 197. 19195 olden, prytatoest, ps 72, 79) 234, aise" 19h; Moldenke, Am, Wild Fls. 291 & 450, pl. = 199; Deam, Yuncker » & Friesner, Proc. Ind. hak: Scie 59% Sle 1962 Moldenke, Monograph of Verbena 209 1950; Fern. inA » Man. Bot., 1209, 1211--1212, & 1612, fig. 1170. "1960," Noldenke, ade rs 3: 37h (1950) and 3: 450. 1951; McDougall & Sperry, Pl. Big Bend Natl. Park = A9Sks Moldenke an Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 2 2210. 1951; Moldenke, Inform. Mold. set 6 Spec. h. 1951; Molden i enke in t- ton & Br. Iylustr. Fl., pr. 2, 3: 127, 133, & 13h. “19585 neh na Am, Midl. Nat. 59: 357. 1958; Moldenke, Résumé 28, 31, me 223, 295, 358, 363, 372, & sae 19595 “ual deuke, Résu- mé Suppl. te 10, & 12. *19603 Lewis » Am. Journ. Bot. 48s Ps (aa asset te ense Hitche. & Norton » Bull. - Agric. Br., Tllu ustr. Fl., ed. Lj 33 a 306). 1898; A. S. Hitehe » Fl. Kans. pl. 15. 1899; Robinson & Fern. inA. aray, New ana Bote, » fig. 880. 1908; Britton & Br., Illustr. Fl. geet fig. 3559. 1913; E. D. Schulz, Texas Wild Fis. 336. $9385 *Volden- » Am. Wild Fls. pl. xx. 191195 Fern. in A. Gray, Man. Bot., ed. Coarse annual or simieaie, “herb , sonst erect or ascending to 6 dm, tall, more often procumbent or nearly prostrate with from the base; roots fibrous; stems at first sprawling or procum- bent, later loosely a ascending, occasionally rooting at the "lower harply tetragonal, spre ding, about 2 dm. long, with erect Ps og rather densely hirsute with canescent hairs 1—1 e-5 mm, long and standing at right angles long, usually with several much reduced ones fasciculate in their axils, peti- olate; petioles 3—10 m, long, alate-margined, hirsutulous; leaf-blades thin-chartaceous » unif: green on both surfaces, ovate in cutline, 2--6 cm. long and wide, bipinnatifid or tri- partite with the divisions again more or less bipinnatifid, Sometimes 2- or 3-cleft, often very deeply 3-parted to the base glabrous above and hirsutulous along the margins and venation th long and subacute, often sharply cut and toothed; inflorescence Spicate, terminal, pedunculate; peduncles 2—-7 cm. long, densely canescent-hirsute like the stems; spikes at eos he “fascicle-like or subcapitate, later elongating to 8 cm., very densely many- flowered, about 3. 5 em. wide during anthesis (1. cm, wide ex- ets narrow-lanc eolate, 9 mm. long, mostly longer than the calyx, Sikdinteatave” and long-attemiate to the setaceous apex, 210 PHYTOLOGIA Vol. 8, no. densely canescent—-hirsutulous or ciliate along the margins; flow- ers with an uncommonly large amount of nectar, cae with a slight sweet odor or odorless; ype 6—-7.5 mm. long, sho pubescent or sparsely sho. ort-hirsutulous, its teeth stuaneous 3 eo wtcoe surface, the limb 8--13 mm, wide, the lobes emarginate; lands much smaller and considerably longer than the anihaereitia 3 fruiting-calyx 8.5--10 mm. long, pubescent, hispid- hirsute along the veins, the teeth very slender, —— subu- late-setaceous from a broader base; cocci cylindric, 2-- long, slightly broadened at the base, reticulate-ecrobiculate a- bove, striate toward the base, the commissural face muricately . ; ous; chromosome number: n = 10 (Solbrig) or 15 (Lewis & Ol- ver * n sunny stone nilis along the Red River, Arkansas, by saan Nuttall and is probably deposited in the herbarium of the Academy of Natural Sciences at Philadelphia, The type of V. demareei was collected ri Delzie Demaree (no, 17198) -- in whose honor it was named — in chalk and marl ditches at Okolona, altitude 350 feet, Clarke County, Arkansas, on June 30, 1938, and is deposited in "the Britton Herbarium at the New York Botanical Garden. This Demaree collection combines the ordinary leaf-characters of ve bipinnati- fida with the large flowers of V. canadensis (L.) Britton and may possibly prove to be a natural hybrid between them if the corolla limb, states, of V. bipinnatifida varies only from 10 mm, in midi Those specimens with the limb up to 13 mm. in d then fall into the V. demareei category. It is possible, however, that the apparent corolla-limb size may be due in some cases to an accident in pressing. The actual knowm hybrid a If V. demareei should prove to be this same ple a nee wet affect the nomenclature of the hybrid, since V. d posed as a true species. Demaree 16114, aes hot on T and abeled a by the collector as "Verbena demareei cacy ae coll.", and for se a calcareous s oy dark or black loam, —_— soll, = lly sandy or dry sandy soil, limesto soil, red clay or red sandy soil, shallow xeric rocky limestone 5 1962 Moldenke, Monograph of Verbena 211 meadows, s pl deposits. It is to be expected along gravel or calcareous road- edges of dried-up streams, alo rights-of-way, on gravel sandbars and in gravel pits, on rolling or broken plains, on rocky slopes and prairies and along sandy roadsides in prairieland, on dry banks, blackland and disturbed prairies, laws and gardens, valleys , thin black soil over lime- stone, open woods, yellow pine forests, and open meadows without crops, on open xeric calcareous hilltops , and in xeric overgrazed pastures, in stony pinelands, and on canyon slopes in Pine forests, in chalk and marl es, near running streams and in red river bottoms, in muddy depressions on plains and de- Pressions in prairies, on high limestone prairies and limestone hilltops, in open dry pastures and rocky sunny fields, in can- yons anc on black rocky clay prairies, in rocky xeric limestone gullies and open rocky meadows, in rolling antidesitic pine- ds and sedimentary badlands » On open mesas and flood plains, grazed parks, characteristic of open ground, fields and uplands, cry me ows, rocky prairies, and high or broken plains, often in limestone soil mixed with sand. It is found on hillsides under oaks (Quercus) and in rich cork elm (Ulms thomasi Sarg.) — hickory (Carya) — oak woods. Ruth port, very throughout" Tarrant County, Texas, "very common throughout the state", Smith reports it as "infre- quent" in Sonora, Mexico. Molohon notes that in Gila County, Ari- it is slopes in Leptochloa-Verbena- ember, and ro on top of dry sterile sandstone dunes in association with mes- quite and cactus in Jackson County, Oklahoma. It has been found 212 PHYTOLOGIA Vol. 8, no. commonly on the Viola and Arbuckle limestone formations in Okla- homa, Warnock & McBryde describe it as "frequent" in Terrell and Val Verde Counties, York as "common" in Travis County, lli- ges as "fre undisturbed caliche" in San Patricio County, S. Turner & Johnson encountered it in native 1 County, Texas; Horr paacionated it in dry loam soil in Kansas. In Jalisco it grows, according to Barkley, Webster, & Rowell, in volcanic soil on mountainsides. Kruckeberg says that it occurs in the yellow pine - douglas- fir - Gambel oak association in Arizona, while Cutter found it soil along edge of pages pg fields" in Runnels Coun Setaria vtvaremapen and Cenchrus in Karnes County, Texas. Tharp, | is "abundant on prairies" in — County, PB reports it as "common" in Bexar and Grayson Counties of same state. It has been collected at eitttades : of 30 to 10 500 ae blooming from February to December, and fruiting from h to December. Deam (190) states that it has been reported rein the Calunst District be papirage growing on railroad embankments, oe gards it as a "migrant" there. Deam, Yuncker, & Friesner (1950) record it dvds Orange County, Indiana. I have not as yet $ “purple verbena", "sm everes "verbena", " na", "sweet William", tye rbena vervain", and "wild verbena". vaece recommended by Standardised Plant Names" is "Dakota Herbarium specimens hav: entified an distributed +e Herburti ae x i dientastaien eas 05 Vs ambr ae folia Rydb., V. ambrosiifolia Rydb., V. angustifolia Michxe, Ve aubletia L., V. bipinnatifida var, latilobata Perry, V. bracte- ata Lag. & Rodr., V. bracteosa Michx., V. canadensis (L.) Brit- ton, V. ciliata Benth., V. ciliata var. es Perry, Ve drummondi, Baxt., V. drummondii (Lindl.) Baxt., V. halei i Small, V. multifida Pavon, V. multifida Rufz & Pav., Ve r, pubera Greene, Ve pumila Rydb., V. racemosa Eggert, V. teucriifolia Mart. & 1962 Moldenke, Monograph of Verbena 213 Gal., V. wrightii A. Gray, and even Abronia umbellata Lam., Ger- aniun carolinianum L., and Teucrium laciniatum Torr os ~~ On the other hand, the A. Nelson 11307, Eastwood “uo, Wooton 363, and Wynd & | Mueller 176 distributed as V. bipinnatifida are actually Vv. ambrosifolia Ry Rydb.; Hartman 906 i is the type collec- tion of V. r, bipinnatifida var, latilobata Pe: Perry; Friesner 22735 is V. canadensis (L.) Britton; Deaver 313k, W. We Jones s.n son. ” ([Pres- cott, May 30, 1922], Mexia 2537, A A. Nelson s.n. ~ [Schnebly Hill, 5/23/40], Edw. Palmer 35, Purpus 1095 and 52h, and E. Wilkinson sn. (santa Eulalia Plains, Sept. 26, 1885] are all v. , ciliata Benth.; J. B. Davy al, Forrer sen. [Sierra Madre, Sept. & Oct. 1881], oe A. . Heller “1388, and and M.S S. Young 113 are all V. ciliata var. longidentata Perry; Purpus se eee + Ve | elegans H.B.K.; ae Brandegee s.n, [Barnwall, May 15, ], Epling & | Stewart sn. (Patagonia, Sept. h, 1936], verses oe Goodding 65, Ee Hl Johnson 962, Nelson & Nelson 136 and 1543, and Toumey s.n. [Gal- liuro Mts.) are all V. gooddingii Briq,; M. Hopkins 609) is xV. oklahomensis Moldenke; G. T. Robbins 2h9 is V. pumila lage and A. Nelson 11388, 11)01, 11566, and 11757, H. H. Rusby 337, and Snow s.n. Son. [Santa Fe, Aug.] are all ve V. wrightii A. Gray. ~~ As to the oc occurrence of V. ‘biped ifida in Utah, a letter re- ceived by me from Seville Flowers, ——e of Botany at the University of Utah, dated January 25, 1955, says "Dr. John D. Spikes handed me your letter inquiring about Verbena bipinnatifida @." My own recording of pasteiietace or the Vorbenknans (Gai & 1919), an anbome (19 (1959) was based on M. E. Jones s8.n. int peng Sta oa herbarium, but on oon nina n I now r ropa this spec- en as representing Vv. wot ry an as actually being a Part of his no. 122 from El Paso County, Colora a Dr. LeRoy H. Harvey, Curator of the Herbarium State University, in a letter to me dated eee “a6, “1955 re- Ports that he has checked through the herbarium there and finds no Montana specimens of this species and "from knowledge of other i e Rodr., Benson 9573 is a mixture with Vv. macdougalii Heller, Haw- —w8 37 is a mixture with V. pumila Rydb., and Pitts s.n. [l/19 By isa mixture with xVe oklahomensis Moldenke, , while @ Fleming 132 is mixed with material of Melosmon laciniatum (Torr.) Small, Herb. Harvey s.n, [h. R. P., Julio 18h3] is Brees sith De Dracoceph- yal PHYTOLOGIA Vol. 8, no. alum sp., and Rydberg & Imler 1166 is a mixture with Hedeoma camporum Rydb, The Osterhout | SMe sn. (Ok--20)06) from Norman, Okla- homa, distributed as Verbena bipinnat tifida, is actually Phlox ilosa var, fulgida Wherry. Iherry. The specimen Trea the Delhi Universi- ty Herbarium in the Gtteborg herbarium is something brassicaceous. M.S. Yo 113 has the leaf-characters of V. ciliata var - longi- dentata and may actually prove to belong to . that taxon. The Ge Je Goodman 5286 collection cited below bears striking resemblance to Ve V. bipinnatifida var. latilobata,. The Lundell & Lundell 10363 cited by me in the "Flora of Texas" is actually V. ve. ciliata v var. longidentata. The Small & Wherry 11968, cited below from Bexar County, may actually have pies bees Tected in some other county of Texas, since its label says m "San Antonio to Austin" and Dr. Small's original field book pe no further information, Lindheimer 1072 and 1073, cited from C "Comanche Spring: Bra is of the opinion that they came from Comal Sovstioy 2 Palmer 47, cited by me from Caddo County, Oklahoma, has a label inscribed "between rt Cobb and Ft. Arbuckle", The Tracy & Earle 106 cited here is not the true no. 106, which is the type collection of v. inconspicua Greene and very ry different in many characters. The He inscribed "Santa Rita Mts.", so could actually have come from ore Rooters 2 Ruth 107 le labels in some herbaria that originally ed "Plants of Tennessee", but the collections seem catiettens: ros have been made in t wo Texan counties Gentry, in his "Rio Mayo Plants", lists V. bipinnatifida tras con tiieerees Sonora and cites his no. 2122 as a Pe are bushy oO op: found the species both in the Transition and the Upper * sono oran lize zones in Arizona, Warnock describes it as "infrequent in eS soil" in Burnet County, Texas. Gates 16666 and 16684 represent a very canescent form, as does also one stem on a sheet of ney 16492 in the Kansas State College herbari @ A. S. Hitchcock s.n. (Winona, May 1895] and ‘Be Bartholone® 8.n, (Rockport, June 12, 1089] were.erroneously cited in the "Kan- sas Flora" as V. canadensis (L.) Britton. Stratton 1,302 looks 4 bit like xV. oklahanensis. Forrer s.n, is also atypical —- Perry identified it as var. latilobata, but the leaves are not broadly lobed. Pringle s.n. [April 13, 1881] is also anomalous —- it has much of the aspect of V. gooddingii Briq. Hinton 13125 greatly resembles V. ciliata var. longidentata Perry, The "March 25, 1617" date on the E- D. Schulz specimen in the University of Michigan herbarium is surely erroneous, yet someone has deliberately emphasized it by going over it with a pen. The collector's name of Nickels s.n, in the University of Pennsylvan- 1962 Moldenke, Monograph of Verbena 215 ia herbarium is there misspelled "Nickles". C. B. Wolf 290 looks much like V. ciliata var. pubera (Greene) Perry, while Peebles & Fulton 11489 in the herbarium of the United States Hels Sie Station at Sacaton was first identified as V. ciliata var. pubera, then as V. wrightii A. Gray, and finally as Vv. -V. bipinnatifida. 7 R&mer sen, in the h herbarium of the California Academy of Sciences, cited below as from Brazos Count; » Texas, may well be from anoth- er county since its label merely states "Colorado & Brazos rivers! Berlandier 14,9 is labeled "Bejar a Austin Mexico", but pro was collected in Travis County, Texas. Carlson en (Bisbee, June 2, 1915] is sterile, but 1 think that it is this species, in spite of the fact that Kearney feels that it is ve cilista Benth. forms a groundcover with Helenium hoopesii A. Gray, Del Artemisia, and Pteris in pine forests. sts. Stanford, Retherford, = Northeraft found it in Coahuila in playa valleys with consider- able drainage from the surrounding hills, with Larrea and Acacia, and with Bisttotous weeds common. Morgan found it growing with Bermuda-gras s [Cynodon dactylon (L.) Pers.] in Oklahoma. re & to introdu ction from farther west, as, indeed, was pointed out by E. J. Palmer in 1935, That the’ jobitte 2 considerably cultivated, and could escape from gardens in suitable localities, is iisiosehd By by the fact that Bailey's notion "Catalogue of Florists Handling Verbenaceae" lists W. A. Bridwell, Forestburg, : A. Barr, Prairie Gem Ranch, snithwick, S. D., an Cronamere as offering its seeds. It is grown as an annual in = trade, and has been in cultivation en: 1819, The specific nam is often capitalized, for no valid reason. The poerg en is orten pnfested by the fungi Phyllosticta texensis Seaver and P. ver- enicola Tharp, CT erusal of the accomp tributional map will indicate & most remarkable disjunct or ei rte i ii This, I feel Sure, is due to inadequate collecting. Also, in spite of the fact that I have examined over eleven hundred herbarium specimens of the species, there are still mumerous institutions in the peihan’ which have not submitted their material to me for —o in spite of eaten ted formal invitations to do so. It Seams most probable to me that opentathey we will find the spe- cles in most of the counties of Texas, New Mexico, Arizona, Ok- g]9 216 PHYTOLOGIA. Vol. 8, no. lahoma, Nebraska, and southern South Dakota. It must be remember- ed that the present map is based only in the specimens examined by me to date. In regard to the accredition of the binomial, Verbena bipin- natifida, Nuttall's (1821) generic entry is "Verbena, ena, (subgenus Glandularia.) Genus Glandularia, Gmelin", and under this entry he has "—- Gl. * bipinnatifida." As Merrill & Reeder (19)6) have pointed out, - say dash clearly indicates Nuttall's intention to publish this as a Verbena, not as a Glandularia. He did not pub- lish the Glandularia combination until 16 years later (1837). agers "anual", editions 5 (1829) to 8 (18)0), eropely credits binomial to Nuttall, although the "Index Kewensi oe it “ pehalat (1817). In addition to the ate a the Beart gern! pany atl 161 following ones fa : Marengo Co.: E. J. Palmer 209. (E); th 536 (5), "19hé (E). WISSISSIPPI! Jackson Co.: sie 47 (G). LOUISIANA: Rapides Par.: Ball Ol (Z, G, W). Saint Mary Par.: Langlois s.n. [17.VII.1893] (Ey W)- MISSOURI: Jackson Co.: B. F. Bush 351 (B, G), 4029 (E, G, W)- County undetermined: Geyer sen. {Upper Missouri, 21 June 1839] (W). ARKANSAS: Hempstead Co.: Eggert s.n. [near Homan, 10 June 1898] (E); E. Je Palmer 8045 (E). Miller Co.: Letterman S.De [Texarkana, Aug. 180] (E). County undetermined: Nuttall s.n SMe {Red River] (G--isotype). SOUTH DAKOTA: Hughes Co.: ico Northwest Exped. s.n. [Fort Pierre] (W); T. A. Williams $-n- [2 Sept. 1891] (E). Lyman Co.: Over 3177 (W). Fall River Bap oe Ryd- berg 935 (G, W). Mellette Co.: Over 2103 (W); T. A- Williams Ss. ny n, (White River, 16 July 1896] (N). Washabaugh Co.: Visher 2132 (F). County undetermined: 0. E. White s.n. [near McClure, Ture, June S901): NEBRASKA: Boyd Co.: F. Ee a Es Coenmtte 2776 (G, W)- Custer Os: Bates s.n, (Callaway, July 1901 KANSAS: Barber (0-3 Smyth 306 (W). Cloud Co.: Carleton sen. [19 May 1888] (E)- Ellis Cos: As Se ‘Biteheock 393 (E, G, W); yd Rydberg & Imler 1253 (E)- Cos: Shear ‘z (G, W). Riley Co.: A. Nelson 5245 (E, G, W). county : A. 82h5 E, G C. S. Sheldon ]8 (W), sen. [Caddo, 22 June 1891] (BE). cimarron Co.: Stratton LL9 (£). Comanche Co.: M. K. Clemens 11751 (E)« Craig Co,: Gurney & Monell s.n. (Vinita, ~3 Aug. 1 1877) (B)- _ Co.: Clifton 3085 (G). Grant Co.: White 197 (BE). Harmon C0-: ge We teas EO 1108 ( (G); Stratton 317 (E). Johnston Co.: Griffith G). Murray Co.: Buig 399 (E 88 (E); G. W. Stevens (E, G, W); Stratton 10 Gl. te 2 Se bo ‘ S Stevens 427 (c)- Woods Co.: G. W. Stevens os (E, G, W), 2050 (G); White loo te 165 (B)- TEXAS: Austin Co.: Lindheimer 16 (B)s Bell Co.: Ward SMe [Killeen, 1 Oct. 1891] (W). Bexar Co.: Ball 909 (W)- Figure 3. Distribution of verbena bipinnatifida in the United States Mapping by county done by Andrew R. Moldenke LR I 13 it 4 T 15 i * 2 \, SK A 'e Ulkeag 2 a Fin Ty : 23 eee Mor ® Laeery z Lt s a Lung a an ee iy aay VaLLey So oes» 2m Ung Dames ae she ° &> Croatoa 5) OTT INEAY POLETTE = | TOWNER | CAVALIER PEMBINA s LITTSON ROSEAU OOSEVELT WiLL aes } ° Los any Cascang b MOUNTRAA, ManENRY - = oe, rs wey Slang vita mausey | WASH Wa eh. ae %, A KOOCHICHING 7 Eres Meccone RCRLAND SON eon ST. Lou ° > ~ Menez e ISLE ROYALE M aries ' MELSON [GRAND FoRxs @ POLK | PENNINGTON an 6 ee ly ro. ceeevieiar? Ra UOT asin ao WALLY Cay McLEAN mn N PETROL DAWSON . N SHERIDAN aie on . oUF ER MEAGney T ITASCA £ ie) th i— H D O a Wer N iste 7 bruncs =z rosTea on NORMAN Pumncuen wv E og a 0 T cnicas “teen oe ie) a, USSEL EL Rosepue | Over pumcce | KiDoER | stuTswan STERE copa) HUBBARD as ie &» “ var CUSTER . cra CASS — WATER is *, oe Ce YER bo CALATINT Pha wreaux [)o.0en ; MORToR ben i TREASURE VALLEY TKN : i FALLOW ONAGON wo . rT Sweet SOP é = WARQUETTE MADtsom Suss YELOwSTong HeTTiNGER — s _— com oI ® BAYFIELD oor Eumoees | LOGAN Us MOURE RANSOM ~ WGLAS CG ae ACER Po cenrT . oman iG oe TR SCHOEN" Pace STnaree BIG Hon ADAMS SUX ottn M I N-NE-] Ss ——# 1RON = DICKINSON] baa MCINTOSH DICKEY WAS A PZ * CARTER ome a SARGENT To00 ‘CROW WING rie POWDER Riven a *AROING c= 2 MORRISON | LACS Pamtec PINE WaSnsuen] SAWYER | ASHLAND . e © PERKINS sos c GRANT | DOUGLAS 5 ~~ SOM =e 2 c— speed Price soron } = CAMPBELL - s caus . s - ONEIDA Fu a es ry Ppl tr MCPHERSON BROWN MARSHALL ‘i MAD NSTONE 2am » ri '. 7 BURNETT MARINETTE pe Pi vi EARN > 4 Lad Aa i he may fi eT raaecast | STEVENS | POPE ers sat = a FORE 2 $f sam TR > 8 WoRn — 2E8ACK | Dewey WORTH EDMUNDS Day i BURN i ¥ UNCOLN FyLAne 4s / ~ pani oT SEDO] wary = GurtE roserts ® 7 afe em 4 RANDITOM ARORA TAYLOR a POTTER a MEEKER WRIGHT g CHIPPEWA me COMA ae 7 ee] ocy ck cur Tc CHIPPEWA ” E fi je™ croix | OUNN MARATHON FeAWANO — ord ert E OOINGTON sawed B OCONTO JOHNSON . ARMSTRONG sr |g CLARK lac ont iY DEUEL ry MeLEOO —r— I N fe ng. |fanerest_]KALKASAN = plbed Nascad ans at FOWILLE conven PieRCE wil c|O N or = LE oa ASHAKIE 3 (@) al YELLOW MEDICINE Th] EAU CLAIRE WOOO | PORTAGE WAUPACA er rita = fa) Simian Frei Ty en scott’ oT > roan me Lawes HAAKON A| K/O od SuBLEY mo} 3 a wanistE - TT) HOT spp WESTON NCE REDWOOD [t7 GOODHUE 2 ThKSON Oo Tart | OScEGA TUARE NGS NICOLLET / SUEUR RICE 2 aT mam TOWOC, 2 PENNINGTON BEADLE KINGSBURY bes = Waseda Porn BAO =>) ASO ay mes BUFFALO a ADAMS rai - us Sut JERAULD SANBORN | MINER | LAKE MOOOY gy PIPE-| MURRAY WASECA DODGE MONROE Th | (SABELLA POwER CARiaoy NCO] SUBLETTE = TACKSON td corroneaos [watonman Ci eoaen oT = a eran WNT] EEO 7 n CONE: AURORA BLUE EARTH STEELE! OLMSTED WINONA ie oO wis Y: ted Ni nmsO™| HANSON] McCOOK | MINNEHAHA gy ROCK | KOBLES — | JACKSON | MARTIN FARIBAULT | FREEBORN | MOWER — | FILLMORE JUNEAU Stee | FOND OU LAC — aa a Coosa pam lOBRARA a - worTe ya ee I Th sad VERIO COLUMBIA | 00066 itm] = = cnesee (PER Scan a vom | ~ e ONE Ing . . s RICHLAND 2 auseroom] KEN . s s s s s é 7 SEAR ig G ean s HUTCHINSON TURNER A LYON OSCEOLA ns mson | EMMET | KOSSUTH] = E842" | WORTH | MITCHELL HOWARD — oA TORI CLINTON ad DOUGLAS x - ") s n\amant SAUK_ dant siraason] wee asst] ORL t @ : 2 ©, c ri w sy pla wo SOX | Dawes TenOAN = oes erecony soma: Tynan Luncoun fy S!0UX O'BRIEN | CLAY PALO ALT HANCOCK Floyd [omcmasaw oe 1OWA ie - cc \ =e Swe CHERRY 2 ‘ON oT a Mw my YEATON e Chee bo ET WATER. Te Poe Ly HOMME cay [=— CERRO FAYETTE cant LEON BAR < cual -/ CARBON PLATT GORDO -maLWOaTH] RACINE . ALBANY 3 eet @ HOUT ° PLYMOUTH [CHEROKEE] BUENA [roc=ontas] womsocor] WRIGHT | FRANKLIN] BUTLER | BREMER e Taraverte. | OREEN be < TASHTENAW | WAYNE 7) ARSON e nnox td VisTA CLAYTON a mn pawn ¥ - 80x BUTTE oxo. *, WEeSTER aocpanan [DELAWARE] “5 . . s er ha carro @ UNTs 2 Da oN - MO DAVIESS | STEPHENSON wt BAG [reine ALOU Lcatt] wONNat tne 7” weeea aN Lorn DUBUQUE parson] fi --<-- Het Plaid poe *%} pene Ud i? Ss [sours com e rane Aegean \ vis] MORGAN SCOmTs ature] wonms cunT pension Ge CALHOUN — TaUA[BENTON | LINN] JONES xeon caRROLL [OGLE TE Tone J 000K pewnck —_* anole wen ‘ bed a | CARDEN MoonR | THomas” T Buane | tour MONONA [CRAWFORD [CARROLL] GREENE | G0ONE | STORY [MARSHAL IO KALB oo sc RrieN . a Rives — cs fume» GARFIELD} WHEELER MADISON [5™™0708] CUMING A CLINTON srt aes] FULTON i. omnes | Susy * my Unnag BANNER surT + CEDAR WHITESIOE if ani Toga on i “| JACKSON a 2 i! x HOWARD € MURDERS ssn WS POTTAWATTAMIE ] CASS] ADAIR] MADISON | WARREN | MARION [MAHASKAT REOMUK } wAo¥™ mex ISLAND 7 3 a moan! ‘ a ma : ‘ool, BUTLER Soucias LOUISA SO cRCER on a ss mer casas a WELD bad omens @ MERRICK“ POLK LEN again HOLMES [9 age LOGAN . 1) [iwineston SPER 4 cod SEDGWICK PERKINS DAWSON BUFFALO HALL Mais] geofly ADAMS [UNION [cLARKE [LUCAS | MONROE WAPELLO | ¢FF8RS0") HENRY} — aus aiae LL ma TROQUO'S. E were [CASS wl , i rr = J SEWARD | Uncastin cia so TOOOFORD tit SoM @ FO Banco PHILLIPS. BERTON CARROLL ft GUNDET NAMA MUERFANO, ] RENO HARVEY ST CLAIR | BENTON ah Fat OR me ANDOU l % an POO™ = Waa Ke es a = BE “ on : Oa oo EDWARDS BUTLER | GREENWOCD FroooSonE ALLEN | BOURBON ff VERNON HICKORY Tcawpen — * Tourasei] PHELPS one corer ad ae ~ fang ey fom 5 Paty OT ay oe US Animas STANTON | Grant | Hasxi PRATT DALLAS] Jwasmincromy %. ‘yWion ag on N AT om rant 2 CONEIOs ) COSTILLA euL DENT shincons PERRY st) we q rt oe é ent Cnn ON a 2 As a KiOWA KINGMAN WILSON | NEOSHO fay ROM os ao a oa eursen font 1art08 feasce ¢ eo Lowe) Fh NE ; 20 agg.) © ce STEVENS | Se CLARK Bs rte Lente fo ee aiene eae wen AT gure \ ey, ron) {ross ae s sand SUMNER Z e ~ on gnctOn =r © oom fwensten] WRIGHT sea . asin em un Anse ee Loe COAL a to 2 cae pom eras rte YNE ; a cA ® i oth aesTIAN me PALE out paid & wautavoual | &® HEROKEEASPER wa Lot OA ites em Se) a wee a a , an Lege S\ tracen cum —, a ptt 7 re xen ot tom . iJ HKINS wil == 5 . a ic CHRISTIAN = oquGLAs HOWELL end locum sare |S io ons Gam} ET CLAVBORNE he a FA (OSAGE ‘Pp NOWAT TAWA ~ ecaon TER _} 8 aan “5 mp LS “ho eis cute = e ea Oo WiPLEY STODDARD mame iemant womens 9 foe oe - ry) “o) may DAL 5 McDONALD ca ryaton e . node] sacason oe A owen. Uo 5 MORA PPotraes ‘SHERMAN n z ELAR ONE . “ ome 7 HENRY cH emnenses J NORA oc « 2, Ne ® HARING IPSCOMa ROGERS 2 . a s a Pn LcuAY pnt) OBI nary js) aarion [ocson Fj WILSON \ seat Fer nAM cocKt Sb, Z deport % 103 auabos| Pane MAYES cee ‘CARROLL | BOONE MARION] BAXTER | FULTON RANDOL romscon A: i} sno commem.ant i) sm . + me ees wait SAN HAR GREEN DYER - u cunt a HARTLEY — PAYNE - SHARP oe “3 cu8son CARROLL euMePRE YS ar Eta mutt RRO H ~n 7, ‘mane BOUNT] g' % N eT ~ MADISON NEWTON ; aay a TULSA] WAGONER, srnsancton seancy I CaNGREAD | MISSISSIPPI & ay Wannen has monroe ce SHA tn, if aero pn eo ROGER MiLL CINCOM CHEROKEE FNOEPENDENCE sehr orcs) “ . dont] YORK f BO Wisvoewo tno Cow i corret Owe (a) oN é ware \o ee Le 2 Sacetson P RMALILLO ITTER MUSKOGEE NSEIT sarwose then maul i cnn) 7S s wae et ity ry *s WHEELER L (MUL GE! Aan © craweons (Rar JOHNSON] POPE [VAN BUREN [CLEBURNE THT MADISON, nt TOL A Cant YP cnc] GES C4 4 rak CHRON AI as c mESTERFILD ond PENOER GUADALUPE A H M. A ‘SEQUOYAN otis A ve Ri ota rr) Fy . PICK UNION cya sTER On A TORRANCE ; DIAN ANOMA CONWAY CROSS reenoing gueiey | FAYETTE tft LuNcou as = ‘! aMcast ave cous McINTOSH . Seog 2 (wun ane 17 cero \ Agauncem\, / oat ARMST HASKE! LoGaN WHITE s 8 KSON id FNP ELD E xX I sod ms a eooonut| ST FRANCIS _~ om 8 mm NEROALE ron | MADISON I came by ANDERSON Lore it we agg! prunn, ee votta= PITTSBURG UE FLORE YELL FAULKNER s bh ‘CORN > i ret fasaenck Feld arose ir PERRY DE Soro wera Se bal pry = mT ciety ea) wore OE Baca Pana pom 0) 1 73 “ z = = > A SHE 0 7 . : nd { suMTE R | BRISCOE Tart inom Dou Enid i ~ een PRET H cont ‘sascvtl ak Cs cl GARLAN! 7 {1 2B FRANKL! ste PUSHMATAHA POX ONROL FLIPS. an) — TNION ammenct|_ MOREA ‘= Q ROOSEVELT LONOKE Un yawn Ton [CULAR ano NYfe as w00n faa aastns AF BAILEY | Lame HALE TAARION | WONST fi s FLOYD MOTLEY COME ‘OTTON RAY = McCURTAIN ] ONTGOMER maint JEFFERSON | ARKAN yr ATOKA IT SPRING td | TEFFERSON HOWAROY PIKE coanonal QRANGERURG : Facopusna = TaNER went Seay FoarD ae SeviER us a caLnouaT oncuses | MON oo — COCHRAN acs CLEVELAND f LINCOLN po ecm FaverTe figsrtRSOm KEYED TO COLUMN A , TN Taway ~ 3 cemna | - Sa LETON of Cleartype Index Book = * ea NEVADA T OUACHITA Dew rq =r Ss Mh : ¥ BRADLEY} BOLIVAR erLore] PICKENS een + CHARLESTON rOAKUM | TERRY 1 wv R CALHOUN STONEWALL | HASKELL Lf CARROLL, HOCTA ey JASPER 2 . 0 AY ro NOKUBEE Hau : BTTALA fre! z TTe here oy peo s (a P; 8 pod tein DAWSOH pestis! Fy gs coLUMBIA . sete WeNSTON wae Scurry | Fi € * . . s —=s a a mE.FOn0 T STEPHENS as Came 8-] CLAIBORNE | UNION worcnouse J 2f DE Uae “i wooo UPSHURY WaRION bs § ; yaz00 FH | ccsrosr| _xeween RY ng MARTIN | HOW FS Sibu NEWTON | LAUDERDALE! ro MITCHELL HARRI LINCOLN a » mms 0 ‘swith RICHLAND f MADE J saan) WHOS ™y’ TRADE MARK REG. U.S. PAT. OFF. "2 J VAN ZANDT GREGG MITA, 3 2 s Lansing yasPER | CLARKE ES wiLCO COL J} ITY OT JTI } JE ® ovine Loe MIDLAND cs vl fms RED = 4 es 5 ‘CURE I ‘ WOMRLER STERLING | CORE vevocnson | wt 910 2 river wnt By eed SIMPSON wow: Bosque WORROE . WAYNE “ + re Chant | urron eal! TEMITOCH he jes 2 COPIAR Tan] JONES. a SECOND | > di (er ‘ atm FREESTONE SABINE ioe serrenson Seca] te if onc FRANKLIN ane Cun a JUDICIAL DIVISION . nie “a4 Fa *y Tomes ae oR ore xX MeCULLOCH HOUSTON ry = MARION . % A FALLS pen ESCMB rors : 19 A or see eG Be ee wry Swenson x es Scale of Miles ai ROBERTSON mn NEWTON® ed a GEORGE t ms SAN SABA MADISON INITY PK ES SPER mt FEAL ecerpmiienl STONE ° 0 100 200 mh I S LLANO MILAM ‘caro | ALLEN EVANGE~ PEARL ‘ . FOURTH PRESIONG Y me) A lent, ote STNG RIVER [7 T HARRISON own ® ers J ’ JUDICIAL DIVISION te oo tos rancid MANY Qc es i wngston \PAHOA fs - Seem] | fn Fos _A MAP NO, 125 = a d ° avs, gat, BERVILE cannon oh a reacts] MONTCOMER} atte bo) H cm wan —N COPYRIGHT U CAMERON * = st i SERNARO wt 7 : g¢ kK “A ! *, erm : N MPANY, INC. OURCHE ‘ORADO errs ust ay } com au Lode > sax «\ ORIGINATORS XS SOLE PUBLISHERS a : a ( 3) c bp CLEARTYPE MAPS 4NS27) COLORPRINT MAPS = ‘WHARTON ed That ame OTST COLD ~ et TRAD mana FRO » "7 t arascosa vse hee RK v WATAGORDA oy NOTICE—This is a copyrighted The law THIRD KAUAI / | ; lc? Cs I the meee, or copying of same, Wi ’ OM any portion thereof, any process for per % JUDICIAL DIVISION / HAWATI f youn tonal use er rene, without = aun: HONOLULU ise: = eer 4 C / PART , 7 exes NUIHAU oT) ] M3 ’ i 4 Te 0 ! e iy Li wr” prc) et pa A Ww ! @n° / OAHU. 4 ano . ons t A / Zz / x t . * i ¢ FIRST u of KALAWAQ vm 1 = ---. ' mn phy eas ii s TAPATA Ji HOGS ous fo ° Sos ee - JUDICIAL DIVISION, “5 : 7 MOLOKAT t. es HENEDY Lit) XN 9 Scale of Miles 80 ie cou | rar) *, ———————————— ANS a’ d ee KURE Ae ? STARR pant ¢ \ ae | 7 * x . Po * ) o wiLACY 1 °”, lng; HONOLULU < 7 Y s * lo aes . NN PART , > USIANSKI, >) ~ = . — LAYSAN : if - . bd * ° ALEUTIAN ISLANDS \ ] : THIRD JUDICIAL DIVISION Sas \ S ue : NECKER . : ea a o? : ~ ¢ > (Een x ? g Copyright, Ameri M c ’ \ oe pyright, can Map Co., Inc., New York, No. 14324 ° Ua “= fee os . . A f : 9 10 1 7 | 1 | 12 | 13 | 14 | 15 | 18 if 17 | 18 | 19 | 20 i 21 T 2 T rr T rT) T 7. T - 1 ° “— T os as a eee i { 2 il Fy T u T uy T M PHYTOLOGIA is financed entirely by its contributors, each onc hs in advance for the entire cost of printing, binding, and distributing his c tribution. All money received from subscribers, after the expenses of = ions have been deducted, will be distributed among the contributors upon the etion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the m eS, assuming his part of the expenses and sharing in the profits, if any accrv Each number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary greatly. A volume will contain about 32 signatures, 512 pages, or a smaller number of with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be angi according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or ton provided they conform to certain limitations of size and proportion. An extra charge will be made for half- tones, depending on their size, as fixed by the engraver, with a minimum of about $2.25. dealing with research in all lines of botany, in any reasonable biographical sketches, and critical reviews and summaries of liter- ature will be considered for publication. Floristic lists, casual notes of an amateur or so-called popular type, and polemics will not be published. Advice on the suitability of manuscripts will be solicited, i. necessary, from quali- fied botanists. ‘ Under the present cost of printing, the basic rate for a page or fraction — Pee = Spetorinnies $3.00 for an edition of 250 copies. This price is subject to change without notice, since it depends entirely on the prices prevailing in the ‘printing industry. Reprints will be fucaiahed at cost. A proportionate fraction of the edition : (250 copies is also furnished gratis to contributors. Upon. request, the edit ~ ill send detailed instructions concerning the tion of manuscript or further information about the magazine. In- quiries seas be aera to > the magatine or to either editor. &: PHY TOLOGIA Designed to expedite botanical publication ae Vol. 8 February, 1962 No. 5 CONTENTS SMITH, L. B., Notes on Bromeliaceae, XVHI]..... 0-0 -ceeecceeters EET MOLDENKE, H. N., Materials toward a — of the genus Gobend: Le ee ee re Pere Pe ee Soe se ZQUR! 80T As eA eCe! Ve, MAR 12 1962 Ea ee) ee ee a oe of sal ill re EDN ee | ie Seay ing ae SEE oe =: « SPOEN sent Published by Harold N. Moldenke and Alma L. Moldenke 1S Glenbrook Avenue — Yonkers 5, New York, U. s ia ___ Price of this number, $1; per volume, $5.75 in advance — NOTES ON BROMELIACEAE, XVIII Lyman B. Smith TROPICAL AMERICA GRAVISIA heady Brazil consists of more taxa than were porns i in seed, and that detailed field observations ons may reveal still others, Our present admittedly incomplete knowledge may be sum- marized as follows: As Flowers on elongate axes, forming na spikes. - Floral bracts minute, not covering the genioulate rhachis, . G a 2. Floral bracts 15 m. sation imbricate and concealing t A Cc _ eS ee ee ane ouwit 1. Flowers on short axes , forming stout spikes or serine 3. a of the floral bracts much longer spon. § he body of the (eee 3 Spies of the floral bracts. much shorter a8 the body of t act. 4. Lower branches of the par arercenas equaling to mch exceed~ the Pp 5- Floral and se condary rae s firm, even or nearly so, yel- low, 2 cm. long; sepal-wing distended basally. ts ral ring 4. G. ee 5+ Floral and secondar ary bracts subchartaceous, nerved, ate, 3 cm. long; sepal-wing cuneate basally. 5. G. rubens 4. eee branches of the inflorescence only about half as long 6. Flo al tel ribo: b or. se@con cts ae ae a cas ega var. aquilega 6. Floral and secondary bracts epg seh 6b, G. ag ee var. chrysocoma 1. waters CONSTANTINII Mez, Repert. Sp. Nov. Fedde 14:2h5. BRAZIL: Indefinite, from cultivation. 2. GRAVISIA LANJOUWIT L. B. Smith, Act. Bot. Neerlandica 5:93, eae pe 1956. 3. a BRASSICOIDES (Baker) Mez in DC. Monogr. Phan. 9:173. 96. 217 218 PUYTOLOGIA Vol. 8, no. 5 Aechmea brassicoides Baker, Journ. Bot. 20:329. 1882. BRITISH GUIANA 4. oe FOSTERIANA L. B. Smith G. aquilega (Salisb.) Mer, cui sftinte, » bracteis primariis S quam ramis brevior vel eos tibus, bracteis se- cundariis florigerisque Dts aureis, floribus haud fascicula- tis differt. Flowering plant over 1 meter high; leaves over 8 dm. long, bearing appressed white scales with brown centers; sheaths broad- pe. brous; primary bracts like the scape-bracts but much smaller, ty- t small specimens; branches Seager the lowest blcecine enlarg- ed to 15 cm. long and with the spikes somewhat separated; floral axes short but distinct; Picaaee in broad digitately clustered mm y te toward apex, even or nearly so, glabro » yellow; sepals asymmetric with a broad wing that is er tgrcsitoa nt uriculate at base, 15 pies including the short muc short—connate; eer appendageds Basal branch of inflorescence x ome fig. 2: Sepal x oe YT the Gray Herbarium, cage on fst ound in low Srey June 10, 1939, by M. B. and R. Souter (No. 8 BRAZIL: Bahia: Dunes, Boca do Rio, near ae, May 4, 1952, L._ B. Smith, Seabra & Leal Costa 7114 (US). In sand in thickets of Cocoloba, on coast 10 miles north of Salvador, May 29, 1939; M, B, & R. Fos ster 45 (a, small specimen with short ene res- cence). Agua Preta, 1939, M. B. & R. Foster 60 (US, flowered in cultivation, inflorescence small, axes flocculose at ae 5+ GRAVISIA RUBENS L. B. Smith, sp. nov. AG. Suess ( — Y Mez, cui affinis, bracteis primariis inf imis amis brevioribus, bracteis secundariis florigeris~ que wubslartacets rubentibu , floribus haud fasciculatis differt. Flowering plant over 6 dm. high; leaves (only one known wn) Cas 55 cm. long, bear ring saath ssed white scales with brown centers; sheaths broadly subedliptic, ipa aieene from the blades, ca. 2 dm, long, dark browm ligulate, broadly rounded and cuspidate, to 12 cm, wide, aremalis serrate with dark spines 3 1962 Smith, Notes on Bromeliaceae 219 bracts, shorter than the branches, deep rose; branches spreading ascending, to 14 cm. long, the lowest with the spikes somewhat mucronulate mm. long, subchartaceous » stro rved, broad- ly convex, slightly carinate toward a ex, glabrous, the ou fs) Sp rose with narrow yellow margin r progressive- hi + ly paler; sepals asymmetric with a large semi-elliptic si that is cuneate at bas @, 22 mm. long including the short mc ro, short- connate; petals appendaged; pollen grains ie multiporate; Ovary glabrous p the epigynous tube 2.5 mm. high. Pl. I, fig. 3: Basal branch of inflorescence x 1/2; fig. 4: Sepal x 1. ype in the U. S. National Herheniun. from salbsyett on in the Jardin Botanique de Montréal (No. 2284-57). Supposedly t came from Mulford Foster's Brazilian collections, but so far I can find nothing in his herbarium vouchers that resembles it. 6. git AQUILEGA (Salisb.) Mez in DC. Monogr. Phan. 9:173. 1896, 6a. So ahecteg AQUILEGA var. AQUILEGA rane romelia es pees Parad. Lond. pl. 40. COSTA RIC VENEZUELA , TRINIDAD, TOBAGO, GUIANA, BRAZIL. 6b. sop ba AQUILEGA var. CHRYSOCOMA (Baker) L. B. Smith, comb. pl. 65. 1892, In liv ge material, the bright yellow secondary and floral bracts of var. chrvsccoma are strikingly different from the dull green ones of ‘typical Gravisia aquilega, but the distinction is Ost in most herbarium specimens. RAZIL: ivated, Devansaye (LG, type or type strain). PER- NAMBUCO: Afogados to Boa a Viagem, Baker & Collins (GH). Caxagua, Ridley, Lea & Ramage (BM). BAHIA: Jacobina, Foster 91 (GH). Teena obra CEA Sw. Fl. Ind. Occ. 1:593. aa aonealmia recurvata 4. L, Sp. Pl. 287. Tillandsia oe sensu auctt. plur., non L. 1753. Linnaeus" original publication of Tillandsia tenuifolia in the first edition of the "Species Plantarum" (p. 286) was based on rences, Van Royen and Plumier, In the second edition in 41,0 ct sao @ e re acta Bulchella Hooker and curiously ane: already has been 220 PerTtorocirs Vol. 8, no. 5 TILLANDSIA TENUIFOLIA L. Sp. Pl. 286. 1753. TILLANDSIA TENUIFOLIA var. LIA Tillandsia foliis lineari-subulatis integerrimi s imbricatis, seu oe > Spica s simplici laxa Van Econ Prodr. Lugdb. 2.1 pissaet pulchella Hook. Exot. Fl. 2:pl. 154. 1825. ier sib pase alae a var. DISTICHA (L. B. Smith) L. B. Smith, Tillendeta ae ae a var. disticha L. B. Smith, Arquiv. Bot. Estado Sao pate nov. ser. 1:114, pl. 117. 1943. TILLANDSIA TENUIFOLIA var. SAXICOLA (L. B. Smith) L. B. Smith, comb, nov. Tillandsia pulchell a var. saxicola L. B. Smit h, Arquiv. Bot. Estado Sao Paulo nov. ser. 1:115, pl. 118. 1943. TILLANDSIA TENUIFOLIA var. SURINAMENSIS (Mez) L. B. Smith, comb. Tillandsia pulchella var. surinamensis Mez in Mart. Fl. Bras. 3, pt. 3:603. 1894. TILLANDSIA TENUIFOLIA var. VAGINATA (Wawra) L. B. Smith, comb. nov. Tillandsia pulchra as ae a Wawra, Oesterr. Bot. Zeitschr. 30:224. MEXTCO PITCAIRNIA MATUDAE L. B. Smith, sp. nov. ee densiflora sensu Matuda, Bromel. Chiapas in Anal. "Inst. Biol. México 23, No. 1, 2: :96, 1952, as to specimen. . densiflora Brongn., cui affinis, foliorum petiolis serru- isthe, " scapi bracteis haud Poryertic eed aber bracteis florige- Pina patentibus vel reflexis differt g shoot erect, 35 cm. high; leaves over 1 m, long, esate acbhedins the inflorescence; sheaths broadly ovate, 3 om glabrous, narrowly channeled toward base; scape ca. 8 mm. in a ameter, brown-flocculose at first; scape-bracts a erect, densely all d ary 2/3 superior; ovules caudate. Pl. I, fig. 5: Inflorescencé x "V3 fig. a Sepal x 1. 1962 Smith, Notes on Bromeliaceae 221 e in the Gray Herbarium of Harvard University, collected on eke vetting State of Chiapas, Mexico, April 9-12, 1937, by E Matuda (No. 1854). VS opregae DRESSLERI L. B. Smit h, = sticha H.B.K. in systema "Meelis | siicdes sed bulbo elon- fie laminis majoribus reflexis, sepalis subaequaliter Tet differt. AT. balbisiana Schult. cujus habitu valde imi- yd » Staminibus quam petalis brevioribus (Subgen. Allardtia) rt. emless, flowering 35 high; leaves bulbous-rosulate, co- vered with minute sabioeresess cinereous scales; sheaths ovate, d forming a narrow- spikes, very densely lepidote; i rs divergent, oblong- lanceolate, 3-8 cm. long, 15 mm. wide, to 16-flowered, strongly complana anate; rhachis flexuous, winged, phe 9 and obscurely Punctulate-lepidote; floral bracts densely imbricate but narrow and more or less exposing the rhachis, ovate, acute, to 18 mm long, mich exceeding the sepals, sharply carinate, subcoriaceous, Tous, lustrous; flowers ee sepals equally subfree, So erg broadly a , 12 mm. long, glabrous, nerved; petals blue-purple o “Dreseler), "the blades divergent, nearly 2 3 long, cnet 4 tamens, Pl. I, fig. 7: Inflorescence x “Type in ~~ her barton of the Missouri Botanical Garden, col- lected on tre 8 miles south of Agua Nueva, State of Sinaloa Mexico, fieteitber 31, 1949, by Robert L. Dressler COSTA RICA ora prone its finis, foliis ro- + Subsecunda Wittm. cud Mt aledianie af. s, foliis tundatis longe eploulatinque, floribus per anthesin vix secundis, Sepalis extus dense punctulato-lepidotis differt. lowering shoot 2 23 em. high, leaves about 15 in a subspreading aa to 25 cm. long, the lower half covered on both sides Mgt essed white scales; sheaths ovate, ca. 5 cm. long; arrowly and long, exceeding the sepals, ecarinate, subcoriaceous, very pale green, » Punetulate-lepidote, flowers subsessile, not secund at an- 222 PHYTOLOGIA Vol. 8, no. 5 thesis; sepals broadly ovate, obtuse, 23 mm, long, 15 mm. wide, ecarinate, punctulate-lepidote on the outside; petals white, bearing 2 lanceolate scales at base, the blades 5-6 mm. long; stamens included. Pl. II, fig. 1: Apex of leaf x 1/2; fig. 2: ay wash ay x 1/2; fig. 3: Sepal x 1. ype in the U. S, National Herbarium, collected in Costa Rica, by Robert G. and Catherine Wilson (No. 31), and flowered in cul- tivation. VRIESEA HELICONIOIDES (H.B.K.) Hook. ex Walp. var. POLYSTICHA L. B. Smith, var. nov. A var. heliconioides floribus plus quam distiche ordinatis differt. Differing — the typical variety in having the flowers in more than 2 r ype the v. ‘8. National Herbarium, collected in Costa Rica, in 1960, by Robert G. and Catherine Wilson (No. 29), and flowered in cultivation. VRIESEA HYGROMETRICA uees! L. B. Smith & Pittendrigh var. ANGUSTIFOLIA L. B. Smith, var. nov - hygrometrica foliorum laminis angustioribus differt. _Jeat-biaites only 25 mm. ° Type in the U. S. National Herbarium, collected in Costa pre by Robert G. and Catherine Wilson (No. 32) and flowered inc tivation. WEST INDIES PITCAIRNIA The West Indian species of Pitcairnia are all endemic t that area with the single exception of P. integrifolia cant which scattered through the Lesser Antilles, except the already noted E. as rifolia which is just in Trinidad and the adjacent main- and, In spite of their geographic separation, the West Indian spe cies of Pitcairnia show few consistent characters to enable one to distinguish them taxonomically from the remainder of th =| e o > a a @ a @ ny £8 Sek Qa ga @ fer g ct A hdl x i be p, @ ral ° 3 ct S t= 5 0 pet Antegrifolia, The seeds are narrow with a very narrow lat section of the wing and usually a broad flat — section. This last siuaee is teisiy distinctive but is n typical of P. albucifolia. The group is revised as follows 1. Racemes dense serenely toward apex; inflorescence simple. Lesser Antilles 1962 Smith, Notes on Bromeliaceae 223 2. Floral bracts broadly ovate.....c.cececesee. eoool.s P. spicata 2. Floral bracts lanceolate........e.cccceeees 2. P. albucifolia 1. Racemes lax throughout. 3. Pedicels 15-30 mm. long, stiffly spreading. Jamaica. 4. Leaf-blades 15 mm. Wid@......cecececeucus 3. P. bromeliifolia - Leaf-blades to 32 m, wide. teecccececesessehs P, platyphylla ds "Pedicels not over 12 mm. long, o or when rarely longer, then curved and secund. Non-Jamaican West Indies. 5. Leaves evenly spinose-serrate throughout, not more than 20 mm. wide, 6. Leaf-blades 10-20 mn. wide; flowering plant 1-2 m. high. rto Rico, Lesser Antilles socvesvecceds P. angustifolia 6. Leaf-blades i mm. wide; oman plant 95, em. high. nican "La pao P. elizabethae 7. The leaves of the fertile rosette dimorphic and the inflo- rescence simple or few-branched; leaf-blades to 18 m. de, wi 8. Sepals 14-16 mm. long. Cuba 12. P. cubensis 8. Sepals 17-25 mm, long. Hispaniola. ge: red; sepals acute..... 7. P. fuertesii . s yellow or cream; sepals obtuse..8. P. samuelssonii 7. Leaves of the fertile rosette all alike or the inflores- cence much branched, 10. iagere: to 25 mm. long; leaf-blades mostly more than 20 r Antilles + Wide, Lesse il. Viewers erect or suberect Leaves all alike; inflores- nce simple or few-br 9. P. latifolia ll. Flowers spreading or n : —— ecund; leaves dimor- phic & Sepals 14-20 mm. long; le af-blades "16-19 mm. wide, + Inflorescence much branched. Trinidad. ll. P. integrifolia 12, rng piar simple or few-branched. 13. Sepals 14-16 mn, Jong; inflorescence moni. simple ee Recess ai Pe cubensis 13. Sepals 20 mn, long; " inflorescence weakly, branched. minican Republic....... P, jimenezii i: PITCA TENA SPICATA (Lam.) Mez in DC. Monogr. Phan. 9:396. 896, la, PITCAIRNIA SPICATA var. SPICAT Prpmelia a spiesta Lam. Encyel. 1: RIS 1783. Lesser Antilles: St. Kitts, Guadeloupe, Dominica, Martinique. 1b, PIFCAIENIA roar var. SULPHUREA (Andr.) Mez in DC. Monogr. 896, Phan. 9:393, Piteairnia & mies Andr. Bot. Repos. 4: pl. 249. 1802. 22h PHYTOLOGIA Vol. 8, no. 5 als yellow. Lesser Antilles: Dominica, St. Vincent. 2. PITCAIRNIA ALBUCIFOLIA Schrad. Blumenb. > 1827. Lesser Antilles: Martinique, St. Vince 3. PITCAIRNIA BROMELIIFOLIA L'Hérit. Sert. Angl. « 1769 (fide International Code Bot. Nomencl. al 1 956)5 “ite Hort. Kew. 1:401. 1789; Oi: Fl. Ind. Occ. 1:580. 1797. Hepetis angustifolia Sw. Prodr. 56. 1788, non Pitcairnia an- gustifolia Ait. 1789. In both of his monographs, Mez identifies Hepetis $ angust foils Sw. with the species of Porto Rico, in spite of the fact z cited "Jamaica" in his original description and Pot duced his own species to Pitcairnia bromeliifolia. Records pr than Jamaica would appear to be misidentifications. 4. ri “pang PLATYPHYLLA Schrad. Blumenb. 26. 1827. amaic re no oie distinction than a broader leaf it is doubtful if this merits separation from Pitcairnia bromeliifolia, but it would be well to examine recent material before reducing it. 5. PITCAIRNIA ANGUSTIFOLIA Ait. Hort. Kew. 1:401. 1789. Pitcairnia ramosa Jacq. f. Eclog. Pl. a 154, pl. in: 1816. Pitcairnia tomentosa Dietr. Lex. Nacht iy Fad 20, in sy - 9:405. 1896. Porto Rico, beh Islands, and doubtfully elsewhere in the Lesser Antilles In using g Pitcairnis angustifolia as a new geesaer yeti" Mez nytt tay both in the application of the basionym, Hepetis an stifo- lia Sw., and in the use of a later homonym of Pitcairnia angusti- felis of Aiton, 6. Aides y a L. B. Smith, Bromel. Soc. Bull. 8: 20, figs. 1 Dominican en Barahona. 7. PITCAIRNIA FUERTESII M be we Rep. Pune Nov. 12:415. 1913+ Pitcairnia tomentosa send % z in DC. Monogr. Phan. 9:389- etr. ex Beer 1857. Pitcairnia fulgens sensu Mez in Urban, Symb. Ant. 8:87. 19%; non Dene. ex Dietr, 1851. Dominican Republic, Haiti (?). Urban cited Taylor 109 from the vicinity of San Lorenzo on ~ mana Bay as the at for reporting Pitealmnia fulgens Dene. fr the Dominican Republic. Pitcairnia fulgens Dene. is pro bably 3 the synonym of the Brazilian P. flammea Lindl., and Taylor 109 a New York Botanical Garden is P. fue: P, fuertesii Mez. IRS. ~rntenseniaeetitnahiakescliiitamiicitititalIN cnn aii 1962 Smith, Notes on Bromeliaceae 225 Plate I VE . | | BEE OA 5 i ag Fig. 1, 2: Gravisia fosteriana; fig. 3, 4: Gravisia rubens; fig. 5, 6: Pitcairnia matudae; fig. 7: Tillandsia dressleri. 226 PeriToLocra Vol. 8, no. 5 8. PITCAIRNIA SAMUELSSONII L. B. Smith, Contrib. Gray Herb. 117: 28, pl. 2, ra 24-26. 1937. Bicester ca hoca os sensu Mez in Urban, Symb. Ant. 8:87. 1920, non ig Dominican Republic, ittt: 9. PITCAIRNIA LATIFOLIA Ait. Hort. Kew, 1:401. 1789. Lesser Antilles: Montserrat, Guadelou —. Dominica, Marti- “a St. — a, St. Vincent . Grenadines » Grenada. t has not been possible confirm records of this species FO Porto Rico and Dominican is epublic and it is suspe they are based on misidentifications, probably of P. fuertesii in the case of the Dominican Republic. 10. PITCAIRNIA GRACILIS Mez in DC. Monogr. Phan. 9:407. 1896. Lesser Antilles: Montserrat, Guadeloupe, Dominica, St. Lucia, St. Vincent, Grenada. 11. PITCAIRNIA INTEGRIFOLIA Ker-Gawl. Bot. Mag. 36: pl. 1462. Trinidad; Paria Peninsula, Venezuela. 12. PITCAIRNIA CUBENSIS (Mez) L. B. Smith, Contrib. Gray Herb. wee” :2h, pl. 2, figs. 12, 13. 1937. sey: ca. rnia tatits var. cubensis Mez in DC. Monogr. ; he Cuba: Oriente. 13. PITCAIRNIA JIMENEZII L. B. Smith, Phytologia 7:1, pl. 1; figs. 3-5. 1959. Dominican Republic: Puerto Plata. PERU TA AMAZONICA L. B. Smith, sp. nov G. danielii L. B, Smith in systema mea (Contrib. U. S. Nat por 29:286-292) proxima sed bracteis exterioribus dense lepido- tis, sepalis majoribus basi solum castaneis differt. lepidote above, sparsely so beneath, at base subdensely serra ate with dark straight or curved spines 5 mm, long, involute when : the outer bracts peer e subtriangula ar, acumina , mcronate, 25 mm. long, alate~ carinate, green except the eter Se base, glabrous or near 4 80; petals 35 mm, long, naked; pollen grains spherical, smooth. 1962 Smith, Notes on Bromeliaceae 227 Pitt, fig. Je Inflorescence x 1/2; fig. 5: sce Type in the U. S, National Herbarium, No. 40,435, collected in Amazonian lowland, Peru, 1960, by Lee apa Fee A-120) NEOREGELIA 1 ee ga (Ule) L. B. Smith var. BICOLOR L. B. isa ae sesataha foliorum laminis subtus dense adpresse- que Se ee pa diffe Leaf-blades s — bicolorous because of the dense minute whitish scales on t shy pe surface. ype in the U. S. National Herbarium, Nos. 2,343,779 and 2534357 780, collected re the vicinity of Loreto, Peru, by Lee arr MOOREANA L. B. Smith, sp. myrmecophila (Ule) L. B. Smith, "as affinis, laminis foliowum valde recurvatis, inflorescentia simplicissima, sepalis connatis lepidotis differt ant propagating vegetatively by stolons 8 mm. in diameter; leaves about 20 in a dense rosette, 3 dm. long; sheaths elliptic, about as long as the blades » subdensely lepidote with brown-cen- tered “seri are recurving, linear, ee » 15 mm de, rate with dark antrorse spines 2 mm, long, sparse- ly pale-lepidote above, glabrous beneath; inflorescence sunk in the center of the rosette, simple, few-flowered; floral bracts elliptic, 3 om, long, about rT the sepals, cucullate, connate tear 4 es sparsely lepidote; als 30 long, the blades narrowly e ve ereoly 2 ridot claaeas cylindric, stout 1 “do mm. long; Rpated aylae ovules apres 2 gro conical, short. Pl. II, fig. 6: Leaf apex x 1/2; fig. 7 Pare bract and flower x 1; fig. 8: Sepal x es Type in the U. S, National reprint No. 2,338,767, collected in small trees » upper reaches cal Napo River n near Iquitos Department of Loreto, Peru, in 1960, by Lee Paes is the same species that oe Moore as s distributed | moll » Neoregelia ossifra Since the earl validly published, 1 Eire his request not to use it. PUYA VARGASTANA L. B. Smith, sp. nov. P. bicolor Mez atque P. araneosa L. B. Smith, quibus affi- nis, bracteis Ged rigeris maximis sepala po leinanyg|g sepalis lineari~lance atis acuminatis differt wn only. ys es but certainly over 1 m. high; leaves ca. 5 dm. long; blades owly triangular, brown-lanate os at Narrowly ovate with. long linear subentire blades, sub- vnsely pectinate-serrate, red, finely browm-lanate, becoming glabrous with age; inflorescence laxly bipinnate, finely brown- 228 PHYTOLOGIA Vol. 8, no. 5 ae cated the petals; 9 | bracts lanceolate, acuminate ong, at least half as long as the axillary branches, - nite po red, the lower ones sparsely serrate; racemes with a long sterile naked eset arched~spreading, laxly few-flowered; floral bracts more or less secund with the flowers, Tanesolats ; a 3 ng, pedicels inconspicuous, obconic, 5 mm. long; sepals agheg lance-triangular, filiform-acuminate, 43 mm. long; p (dmma~ ture) naked, Pl. II, fiss 9: Branch of inelotentatest : xP fig. a. 22s Type in the Gray Herbarium of Harvard University, collected at Amaibamba, Santa Barbara, Province of Convencién, Department of Cuzco, Peru, December 30, 1948, by C. Vargas (No. 7558). BRAZIL thc vias EGLERI L, B. Smith, sp. AP. torresiana . Smith, ig affinis » foliorum laminis persistent ibus basi seratis diffe Stemless, flowering 35 cm. high but the leaves exceeding the inflorescence; leaves dimorphic, all persistent, to cm. long; e, laxly r me, 7 - long, rather few-flowered; floral bracts spread- ing, Tanneel shes to 8 mm. long; pedicels ascending, slender, 15 mm. long, much exceeding the bracts; flowers red; sepals pores broadly rounded at apex, 13 mm, long, ecarinate; petals long, bearing a scale at base; — slightly more than 1 1/2 supe- rior; ovules caudate. Pl, II, fig, 11: Base of plant x 1/2; fig. 12: Inflorescence x 1/2; fig. i a pie x 1; fig. 14: Sepal x 1. Tapajos, State of Para, Brazil February 3, 1960, by W. A. Egler a iat 4 Raimundo, Isotype in the ituseu Paraense "Emilio ” species is dedicated as a sorrowful tribute to my friend and colle eague, the late Dr. Walter Alberto Egler, who lost his life on August 28, 1961, while exploring the "Rio Jari. ARGENT INA PITCAIRNIA ee th sp. nov. » Bs Sat ana pet cud affinis, scapi bracteis remo~ — rt. g 1m. high; leaves dimor hie, some reduced to » other or rp functional with slender serrulate petioles; blades ny pt 1962 Smith, Notes on Bromeliaceae 229 Plate II Fig, 1-3: Vriesea apiculata; fig. 4, 5: Greigia amazonica; fig, * Neoregelia mooreana; fig. 9, 10: Puya vargasiana; fig. 11-14: Pitcairnia egleri; fig. 15-17: Pitcairnia oranensis. PHYTOLOGIA Vol. &, no. 5 mm, wide, entire, channeled toward base, covered beneath brous above; scape erect, § mm. 230 long, 24 with white appressed scales, gla sterile laxly ; pedicels ascending at anthesis, 15 mm. long, s s oblong- lanceolate, obtuse, ong, ecarinate; petals obtuse, naked; L rior, Pl. II, fig. 15: Lateral branc of 800 meters altitude, October 31, 1938, by W. J. Eyerdam and A. A. Beetle (No. 22795 in Goodspeed Expedition). MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. IV Harold N. Moldenke ENA BIPINNATIFIDA Nutt. A continuation follows of the listing of specimens cited by Miss Perry, but not as yet examined by me: TEXAS: Bexar U0-: Berlandier )29=71749 (G), Us9=189 (G); Eggert sen. [San Antonio, April 1901] (G); Groth 30 (F, G, W); Moore & Steyermark 3001 ct 0, April 1919] (@)i Lind- W). De Witt Co.: Howell 308 (W). Edwards Co.: M. E. Jones bee” “saa o.: Berlandier 2,20« 17 June 1897) (E)- Howell iu ee 00 E); Ee Jd h May 19 1s Fy (E). Gillespie Co.: G. Jermy 183 (E). Goliad © 998 (E, G). Hays Coe: Trelease s.n. [San Marcos, Hemphill Co.: Eggert s.n. (Canadian, 10 Aug. 1900] (E); (W). Hood Co,.: Eggert s.n. {near Granbury, Palmer 6510 (E). Kaufman Co.: Tyler s.n. [vicin May 1904] (W). Kinney Co.: Mearns 1252 (W), 127) (W), 139% ag Navarro Co.: E. J. Palmer 7821 (E). Parker Co.: Se Me ). Taylor Co.: EE- E gert s.n. [near Abilene, 7 June 1900) (E); irs = sees organ Eggert F). Travis Co.: E. J. Palmer 33389 (E, G). Victoria oe bee J s.n. (near Victoria, 10 April 1900) (E); Maxon 3808 (W)s B+ os 1962 Moldenke, Monograph of Verbena 231 Palmer 9098 (E). Walker Co.: Dixon 561 (F). County undetermined: Lindheimer 232 (E), 289 (E, F, G, W), 307 (G). NEW MEXICO: Chaves Co.: Earle & & Earle 526 (E). Santa Fe Co.: Earle 8h (£). County undetermined: W. Wright 1502 (G, W), 1504, in part (D). ARIZONA: Cochise Co.: Blumer 1345 (E, F, G); Goo Goodding 251 oS a a Goodman & Hitchcock 1211 (E). Coconino Co.: } PEs T. Mac al 317 (D, F, | F, G). Pima Coe: “Pringle Sen. [April 13, 1881) GRE Santa Cruz Co.: Peebles, Harrison, & Ke Kearney 4549 (W). MEXICO: Chihua- hua: Nelson 6117 (G, W); To W); Townsend & Barber 139 (E, F, G, W). It should be noted that the New York specimen of Visher 2132, which she cites here, is regarded by me as V. ambrosifolia Rydb., and those of Bush 4029, Blumer 1345, Earle & Earle 526, Goodding 251, and G. W. Ste Stevens ens 1h are regarded by me as Vv. bipinnatifida var. latilobata Perry. She cites Hayden s.n. (E) “from Montana with a question — Haas label oq fang pa says "sandy soil along Yellowstone and Upper Missouri", She cites M. E. Jones 26228 ani 26229 as "in part", = tomas as a opr re with V. ve pumila Rydb. and the latter with V. halei Small, She cites Berlandier ndier 1)9=189 in the Gray Herbarium as from "San Antonio de Bexar” [Bexar Co. mar Co}, b but I cocliggaaas no. plans) in the Lund herbarium as from Travis Miss Perry annotated a specimen of Mearns 159 from Flagstaff, Arizona, collected on June 17, 1887, in the Columbia University roe em at the New York Botanical Garden as "V, bipinnatifida Nutt." in 1932, but does not cite it under that species in her monograph it is p robably the Mearns s.n. ch she cites there under var. ietitopate, and I am also so regarding the collection. She cites an an Eggert s.n. from "along seen eat, Miller Co. [Ar- kansas], 19 June 1908" as in the New York herbarium -- actually the date of this collection is "1898", not "190 08", excellent monograph of the North An- Miss Perry, in her erican. species of Verbena, notes that "In the specimens Arizona and Me exice. =the weenenes is hispiculous and = most co) nized by the hispid-hirsute pubescence, partic florescence, the lack of glands, the long floral bracts, the e- t the deeply cut leaves, and a rather coarse ha-~ ciliata in northern Mex- canadensis i in the southwestern part of its range. Neve rtheless, oe spite of the many intermediate phases, it seems preferable maintain = group as specific entities." 232 PATFTOLOOIA Vol. 8, no. 5 Berpes. 25 5 (N). Greene Co.: R. M. Harper 3183 (Ba, N); We He = 6s s.n. (Clinton, July 1927] (Hs, N). Montgomery Go.s Le He alley U5 (Ba); L. H. Pammel s.n, (May 30, 1921] (Io—-102h36); W. = Rhoades s.n. TH. N. Moldenke 17089;Montgomery, July 1931] (Bt, ¥, St). Sumter Co.: We He Rhoades s.n. [near Gainesville, July 1927] (N).» MISSISSIPPI: Lee Co.: R. D. Ray <058 (N); Be Shelton 19 ook Lowndes Co.: Diener 930 (T1). cerbicie Co.: Weatherby & Weatherby 6313 (N, Up). Scott Co.: We H. Rhoades s.n, [Morton, July 1927] (N).» Sharkey Co: J. De Ray 4923 (N). SOUTH DAKOTA: Brookings Co,: Gunn s.n. [Brookings, 7-10-1895] (Io--1533%). Brule Cos: E. Ne Wilson sen. [July 1893] (Ob--50770). Custer Co.: Hay- ward 2562 (Se--77056). Fall River Co.: Rydberg 935 (C, N)- Hughes Cos? T. A» Williams s.n, (Pierre, Sept. 2, 1691] (N), Sen. [Blunt, Aug. 16, 1892] (P1--22605); Williams & Wilcox s.n. [Pierre, 1891] (Ob—50769) « Lyman Co.: Over sen. [July 1h, ISI] (Se—1L956) » Mellette Co,: Brenckle 51323 (Le); E. J. Wallace s.n. [White River, 7/16/1896] (N). Shannon Co.: Visher 2177 (N). Todd Coo: C. J. El- more sen. [Rosebud, Aug. 24, 191)]) (Je—315h). aad undetermin- ber Co.: 3 We | “te Horr 3569 (Iw). Cloud Co.: Carleton sM-. ig July 20, 1883) (Ur), sen Sone (Aug. 20, 188} (Ka), SeNe {June 1886} (Ka, Ur, Ur, Ur); S. Ve Fraser Uh2 (Ka--78951)- Decatur "Cost A. S. Hitchcock s.n, [J Jennings, June =. 1892] (Ka). Ellis Co.? Bondy s.n, sen. [June 7, 1935] (Se-—5l978); G. C. Deane S.n- {June 2 1882] (Dt); F. C. Gates 16931 (Ka--76208); A. Se “S. Hitchcock 393 (N), Sen. [Plains, July 16, 1895] (Ka); Imler BeNe - [May 18, 19 1928] (Lw); Kellerman 9 "(N )3 Rydberg & Imler 1166 , in part (N), 1183 (Ka—-75067), 1212 0K), 1253 (Lw, N)3 Sci. i, Exped. Mee sen. [June 2%, oe) (Ka); H. Shepherd 63 (Cm, Dm, En, Ms, Ok, St--9211, Vt). rth Co.: M. Becker s.n. [Kanopolis, Summer 1896] (Ka); Ae 5: ane sen, (Ellsworth Co., July 1895] (Ka). Geary Co.: Fe Ce Gates 17500 (Ka—79l08). Gove Co.: A. S- Hitchcock 5.Meo [Gove Coe, July 1895] (Ka). Graham Co.: Imler 21 (Ka—7707), 2021 422h7), sen. [Bogue, Sept. 12] (Lw). Hodgeman Co.: As Se Hitchcock s.n, [Hodgeman Co., August 1895] (Ka). Jewell Co.: W. H. Horr 1,368 (N); B. Osborn 863r (Ka—-88363). Kearny Co.: A. S . Hitchcock S-Me [Kearney Co., Aug. 29, 1897] (Ka). Lane Co.: F. C. Gates Gates 1668 (Ke 75855). Lincoln Co.: F. C. Gates 16492 (Ka-—75906); A» Se $. Hitch- cock sen, [Lincoln Co., July 1895] (Ka). Logan Cos: Ae er “qiteh- cock s.n, (Winona, May 1895) (Ka). Meade Co.: A. S . Hitchcock soMe [Meade Co., Sept. 1, 1897] (Ka); W. H. Horr Sen. Te /T/z] (im). Mitchell Co.: Carleton sen. [May 19, '88] (Ka). Morton Co.? Ae Se Hitchcock s.n. [Morton Co., August 1895] (Ka). Ness Co+s Av Ss Hitchcock s.n, [Ness Co., August 1895] (Ka). Norton Coe? F. Ce 1962 Moldenke, Monograph of Verbena 233 Gates 17456 (Ka--7933)), 18081 (Ka--88035). Osborne. Co.: Ae Se Hitchcock s sen. [Osborne Co., dt June 20, 1897] (Ka); Shear 38 (C). Ottawa Co.: Godlove ve 319 (Ka--93071). Pawnee Co.: A. S. Hitchcock 8.n. [Pawnee Co,, August 1895] (Ka). Philipps Co.: . Ae | S. Hitch- cook Son. ipa sa ne June 29] (Ka). Republic Co,: A. S. a cock sen. [Court » June 2), 1892] (Ka). Riley Co.: Birch s [5-22-1899] ae oe bine s.n. (Manhattan, May 31, 1886) (1 me an. Sen. [May 31, 1886] (Ur); J. We Finley sen. [se. of M May 2 tan, 7) (X8)5 A. Nelson 82h5 (N);3 S. Stewart s.n. [May a1) (Ka). Rooks Co.: E. Bartholomew sn, [Rockport, June 12 1, 1889] (Ka); Clothier s.n, [Aug.-Sept. 1898] (Ka); Imler s.n. (sep. 12, 1929] (im). Rush Coe: A. S. Hitchcock sen, [Rush Co., August 1895] (Ka). Russell Co.: Carleton sen. [June 25, 1885] (Ur); Kellerman s.n, a 25, 1885] (Ka). Saline Co.: Carleton s.n. [7/28/1892] (Ka); Hanein 251 (Ka-~76788). Sheridan Co.: C. Weber 36 (Ka—781,7). Smith Cov: We H. Horr £.108 (B, Mv, Ok, S, St, Tl, Ur, We), W. ~ 8 (Ca—bs951)3 3 Kellerman s.n. [Smith Center, July 10, 1888] (Ka). amas Cos: A. S. Hitchcock s.n, [Thomas Co., June 27, 1897) (Ka). a © Co,: F. Cad Gates ates 16666 (Ka——75838); M. Reed s.n, [July 8, 1892] (Ka), son. [July 1392] (Ka). Wabaunsee Co.: Bartley 120° N)j; We H. Horr s.n. [April 28, 1939] (Lw). Washington oh Dodd 35 (Ka—-B30595), 58 (ka—80595). County undetermined: W. S.n. [western Kansas, July 1871] (Pa); 3 Oyster 4672 (N)5 = s. Sen, [n, west Kansas, 1878] (Cm). MISSOURI: Jackson Co.: B. Pe Bush )0: 4029 (S). County indabiniaeds L. He Pammel s.n. [5/20/ 00} (To (Io--6199). ARKANSAS: Clark “tg Demaree 1611) (N), 17198 (Du~-25656, Gg—267601, It, Ky, N, N, N, Ok, Ur). Hempstead Co.: E. J. Palmer 8045 (Gg—182183, fen’s9}. Izard Cos: F. We 20696 (H), 2 (N), 17004 (Gg—267608), Miller Co.: Eggert sen. [19 June 18598] (N). County undetermined: Nuttall s.n, [Red River] (C—iso- ms Wibbe sen. [9-10-79] (Al). LOUISIANA: Caddo Par.: Cocks s. : [June 1908] (T1). Kast Feliciana Par.: Drummond 194 (Lu), 212. i Ta). R Rapides Par,: Ball 101 (N). Saint Mary Pars lois s.n. [17.11.1893] (I, N). Vermilion Par.: Langlois son. [28.V-1003) 1). COLORADO: Baca Co.: W. A. Weber 1589 (Ca--945347). NEBRASKA: Boyd Cow: F. E. Clements BF (Gg T92) 2776 (Gg--31292, Ka). Furnas Co,: J, We Barnha: Barnhart son. [Hendley, june 4, 1885; Herb. J. H. Barnhart 13) (N). Sherman Cost Jo. Ba son. [June lL, 193} (0b—50773). Valley Co.: C. nig sen. [Ord] Saige Webster : ®mmined G. W. Stevens Mm mele) Beckham Co.: Demaree 12h51 (ou--20575 Ok-=22233) 5 : 1737 (Ok); J. R. McFarland 64 (Ok); Neill 27 (0k); ee (M1, N, St--26135, St—26136); Waterfall 1998 (Ok, 23h PHYTOLOGIA Vol. 8, no. 5 re Blaine Co.: J. Engleman 1272 (Ok). Bryan Co.: W. S. Blain s. . (Durant, april “10, 1934) (Bt--29693); Brage 2 oe arg: ag na Sheldon 18 (Se—11957) « Caddo Co.: Graumann 17 (St--17578)3 oa Palmer 47 (N). Carter Co.: Demaree eo 12296 (Ba, Ok—-22206, St- 12969); Demmer 31 (Ok), 46 (Ok); M. Hopkins ins 4691 (Ok), 4714 (Ok, Ok), 4837 7 (OK), ~ 6094 (Ok) 5 Hopkins, Yelson, & Nelson 1131 31 (0k); Marian an 30 (Ok); Ae Ge Payne 18 18 (0k); Ea 193 (Ok). Choctaw Co.: Estes 93 (St—-17572) 3 Hopkins, Nelson, & s& Heise a aa? 742335, Ok); | McAfee 6 (St--21503); Nelso eis Nelson, 23: (Ok); Smith & Randall 73 (Ok); Spears 77 Sot eT Bt = . Cimarron Coe: Rigney y 129 (St—-20028) 5 Smith & Rinehart a2, in part (0k); Stratton ‘ten bio 9 (St—26130), 5195 (N, “st--26153, St—2618h)5 Materfal. a oh (Ok), 9049 (0k). Cleveland Co.: Babb 3 (Ok—-10277); Re Ee @ 50 (Ok--10281), 97 (Ok--10280); Fielder Fielder sen. [5/16/27] | icon Hibbard 17 (Ok--10282); Louise e Martin se en-s) (Ok--10271); Pitts sen. [/19/15] (Ok--20h77, | ae ). Comanche Co.: Buttram & Buttram O (0k); M. K. Clemens er yond Gg--37142), s.n. [Fort Sill, Apr. 27, 1916] (Ba); Ce T. Eskew 1663 (0k), 1675 (0 (0k), 1750 (Ok); Hopkins, Nelson, & Nel Nel- gon 801 (Au, Ca--742279, N, Ok, St—-26115); M. L. Nelson felson 9 (Ok) 5 C. S. Sheldon 48 (Al); Waterfall 13097 (St). Custer Tr COs: Mericle en (0 or) ee 1265 (Ok). Davis Co.: M. Hawkins 37, “in part ‘ton 3085 (Po--127527), SeMe aleve 3085) ee nian Cos: Gephardt 1016 (Ok). Garvin Co.: Kennedy 31 (Ok); E. J. Palmer 12063 (Ga-—534758). Grady ag Ee Morgan a (Ok). Greer Co.: Bailey ¢ & baler 10157 (Ba); Herb. , Univ. Oise 14899 (Ok). Harmon Go.: Sherrill 205 (0k); G. W. Stevens (0k, St--9222); puateeet aT (St--26126); Waterfall 8330 oy St). Harper Co.: G. W. Stevens 305 (Du-—652h6, Ok, Ok, St--9218, Ur). Jackson Co,: Demaree 12134 (Ok--2220h), 12138 (ok——-20380) , 12216 (Ok—-20360); M. Hopkins 1024 (Ok), 3067 (Ok). Johnston Co.: Hop” kins, Nelson, & Nelson 99 (Ok). Kay Cos: Kelting 231 (0k ). Kingfisher Co.: J. Engleman 1271 (Ok); Grace SoMe oe 27, 1935] (0k—-19063). Kiowa Gos: G. J. Goodman 6272 (Ok). Lamar Co+? Der aree 12003 (N), Logan Co.: J. Engleman 1273 (Ok); C. C» Smith 43 (0k), 673 (Ok). Love Co.: McLennan 3 mnan 3 (Ok); Nelson, “Nelson, Good- = bee ee 5695 (Ok). Major Co.: Je Re Mc , McFarland 30 aes aterfall & Coryell 9921 (Ca--993896, . M Oot Ge Je Goodman 5817 (Ok), 5865 (Ca—-81889, jy atari 698 - -Oi)s Ae Wilson s.n. Same 12, 1950] (Ok). McClain Co.: Demaree @ 12859 (Ok— 22017, St--12968). McCurtain Co,: Keeling 8h ” (gt—=17570) « )). Murray Co.: Beard 65 (St—-9233); R. Bebb 10 Wako (0k); Becker, Waterfall, & se ee Kennedy 5299 (Ok), 5342 (Ok); Brenckle 8065 (N); Broadbent, Lé e)3 +118 (Ok), 295 (Ok); Coleman 19 (We); E. E. Dale 50 (Ok), 111 (Ok), 118 (0k), 22 1962 Moldenke, Monograph of Verbena 235 (Ok); Demaree 1222 (Ok—-22221), 12277 (Ok—22162); Enig 43 (Cm), 788 (Cm); Fielder 4 (Ok--10261); Flowers sen. [May 3, 1939] (Ok, Ok); Goltry 10 (St—1230); G. J. Goodman 2078 (Mn—-26180, Ok—- 17969); G. BE. Hall 59 (Ok); M. J. Hall 13 (Ok); Harden 108 (st-- i ccauealilicemnndtiiceanaeeaeeaee — 25040); M. Hopkins 2765 (Ok, St--15021), 2730 (Ok), 3959 (Ok), 4736 (S), 4771 (Ok), L823 (Ok), 5178 (Ok), 6075 (Ok); oe 280 Cross 6421 (Ok); Hopkins, Nelson, & Nelson 667 (Ca—742200, Ok), 709 (Ca-—72338, Ok, St--26116); Jeffs s.n. [May 15, 1918] (Ok— 10270); Kinnan 8 (Ok); Marian 25 (0k) 3G. M. Merrill 283 (N); W. S. Myers 31) (Ok); P. M. Nelson 102 (Ok); Randal hh (Ok); Rice 22 (Ok); K. Starks sn, [16 July 1950] (Ok); G. W, Stevens (Du—66651, Ok, Ok, St—9216, Ur); Stratton 10 (N, St--2118, st— 26119); Van Valkenburgh 81 (0k). Oklahoma Go.: Stedmen 79 (Ok); Waterfall 1982 (N, Ok, St--21491). Payne Co.: Ray 110 (0k); Stiles Sen. (May 20, '97] (St--21930). Pontotoc Co.: Bs E. Cole 94 (St-- 21678) D. McCoy 1149 (St); Prater 10 (0k); Preston 90 (Gg— 316093). Pushmataha Co.: Parker 21 (Ok). Smith Co.: G. W. Stevens 699, in part (Ok). Texas Cos: G. W. Stevens 27 (Du~-65),05, Ok, St--9212, Ur), Tillman Co.: Fleming 132, in part (Ok); C. C. Smith 1005 (0k). Woods Co.: G, W. Stevens 699, in part (Du--66663, N, Ok—-21373, Ok, St—-9213, St—-921h, Ur), 2850 (Ok), A.2850 (St— 9215). Woodward Co.: G. J. Goodman — (Ok). County oo aon Carleton 177 (Ka, Ur); C. Te Eskew (Ok); Ge W. Stevens (Ok), 4.1063 (ok), 43088 (ok); Van Vieet s.n, [iit. Sheridan, 7/ 4/03} (Io—11 7045). TEXAS: Anderson Co.: Plank s.n. (Palestine, April 19, 1895] (N). Andrews Co.: Lundell & Lundell 11105 (Sm). Austin Co.: F, W. Pennell 1028) (N)3 Turner & Johnston 54345 (Au— 121642); H. Wurzlow s.n. [Industry, 1691] (Ur). Bandera Co.: L. H. Seana Bailey 7482 (Ba), 7501 (Ba), 7503 (Ba). Bastrop Co.: ° SelM, (Bastrop, 1928] (W==1),38667); Marrs son, (Smithville, -10-1930] (Ca——882571); Tharp 2817 (Du—206897). Baylor Co.: Bridge sme a Seymour, 6/16] (Cn). Bee Co.: F. A. Barkley 167423 (Au, N, Or——55386)5 F. We Gould 6060 (Au—-121653, Ca——985733); Me Ce John Ston 5141 (Au--1216h3, St); Lundell & Lundell 10055 (14, N); York & York 54035 (au--121637). Bell Co.? Lundell & Lundell 10135 (N); McDaniel 1, (Nt), son. [Salado, Aug. 6, 1930] (Nt)j C. L. York 46389 (Au); York & York 5022 (Au--12166), Bexar Co. Balcom SoM. (San Antonio, April 1917] (Up—-75578)3 Ae Ball h (Av--121651); Barkley & Parsons 1 (Au, N); R- Bebb 2272 (Ok), Son. [San Antonio, May 1, 1905) (Ur 3 Bogusch 199 (Ur); Clemens & Clemens 976, in Part (Ba, Gg—3713%6, Po—69803); Collector undesignated s.n. [San Antonio, 1926] (Wi), sen. (Sr); Groth 30 (N); Ge Jermy sen. [190k] (N), Sen. [San Anton.] (N); I. S. Jones 2209 (Po~-158869), Sn. (Laurel Eeights, Mar. 12, 1926] (Wi); May some [3-18-26] (Wi); 236 PHYTOLOGIA Vol. 8, no. 5 McCullough 6 (0k); Metz 79 (Gg—310895, I, N, Ok, Se—57949); Pa~ gel s.n. [San Antonio, Mar. 1, 1926] (Wi); L. Richter s.n. [San Antonio] (Au); E. D. Schulz s.n. {San Antonio, March 25, 1817] (Mi), sen. [San Antonio, March 18, 1918] (Mi), sen. (San Antonio, 3-15-1921] (Wi), sn. [San Antonio, 4/13/1922] (Wi); C. Skotts- berg s.n. [Sibolo Creek, Selma, 1/6/1935] (Go); Small & Wherry 11968 (N). Blanco Co.: Bogusch s.n. [April 8, 192h] (ob—5077h) 5 Iundell & Lundell 1027) (Id, N). Borden Co.: Whitehouse 16716 (Sm). Bowie Co.: Letterman s.n. (Texarkana, Aug. 1880] (N). Bra- zoria Co.: Bush hig (N). Brazos Co.: 0. M. Ball 150) [Reeves 199] (Cs); L. He Pammel son. [71888] (Io—15341); Romer sen. (Herb. Prager 1863h] (Gg—~31291). Brewster Co.: Warnock W. Bh, (Au) » Brown Co.: Brenckle 18078 (N); F. W. Pennell 10181 (N); K. Stan- ford s.n. [Brownmood, 5-5-1931] (Au, Mi). Burleson Co.: Reeves 185 (N). Burnet Co.: C. C. Albers 38003 (Au, Au); Barkley & Cope land 36a (Au, N); F. W. Pennell 10h69 (N); Warnock W.1096 (Au). Caldwell Co.: McBryde s.n. [Spr.-Sum. 1931] (Au, Au); Mercer s.n. (Fentress, 3-29-1931] (Au); Straudtmann s.n. [Dec. 00, 1936] (au). Cameron Co.: R. Runyon 2495 (N, N), B3u0 (Br, M1, N, Ug); G» He Thurber s.n. [Fort Brom, Feb, '57] (N). Carson Co.: A. Me Ferguy gon s.n. [4/3/01] (Au). Castro Co.: Rachaner 102 (Au). Childress Co.: Beazley s.n. (Carey, May 16, 1929] (Au); Wild Flower Contest sen. (Childress, May 1930] (Au). Coleman Co.: Pruitt 228 (Au— 121661); Shinners 26392 (Ur). Collier Co.: Lundell & Lundell 9295 (La, Mi, N), 9296" (Mi); McCart 1541 (Nt); F. J. H. Merrill 5.ne [May 1, 1886] (C). collingsworth Co.: Bailey & Bailey 10143 (Ba)- Comal Co,: W. H. Kellogg 8 (Au); Lindheimer 10 (Ka), 1072 (Au, (N, Up 48540); F. W. Pennell Shih (N, Up—72053); Rauterberg 1593 (Sm). Comanche Co,: Wild Flower Contest s.n. [Apr. 30, 1930] (Au). Cooke Co.: Correll & Correll 12969 (Sm); Newton 10 (Nt). Coryell Cos: Warnock 46413-(Au, N); Wolff 3458 (Tr). Crockett Co»: Cory 35469 (N, N); Warnock & McBryde 15313 (Au--123235). Crosby Go: C. 0. Erlanson 1135 (Mi), 1160 (Mi). Culberson Co.: Hitchcock & Stanford 6762 (Po—265508). Dallas Cos: R. Bebb 2456 (Ok), 1310 (Ok), 1374 (Ok); Boll s.n, [1871] (Cb); E. Brainerd s.n- [ , March 28, 1908] (Vt); Bunker s.n. [May 'lj2] (Ob--50762); Collec” tor undes s.n. [Highland Park, 329-19] (Wi); A- Be Couch 3 (ur); I. B. Diehl 13 (Po—71128); L. Goodwin 12 (au--1669L5); Hamric & Thompson 6 (Au—-172518); Hendrix 16 (Nt); Hynes s.ne [5 18-26] (Au, Au), s.n. [Spr.-Summer 1926] (N); B, Jones 6 (Nt) 5 Iundell & Lundell 7056 (Mi, N, N), 8333 (Id, Mi), 853h (1d, MA, N), 9161 (Id, Mi, N, N), 9137 (1d, MA, N), 9771 (wi), LOWI6 (LA, 1962 Moldenke, Monograph of Verbena 237 N), 10568 (Ld, N), 11315 [Plant. Exsic. Gray. 1275] (Al, Au, Ca— » Dp—31802, Gg—-308962, H—~106661, I, Ka—95212, Ld, Mi, Tl, Ur, Vi, Vi, We), 1170) (N), 12132 (N); Ness s.n. [Auge 7, 1925] (Tr); Re L. Oliver 76 (Nb); F. W. Pennell 5399 (Up~72783); J. Reverchon 739 (Up—1708h), 1962 (Up—17087), 1962* (Be, C, Ca~ 445518), s.n. [Curtiss 1962] (Cm, I, Pa, Vt), sen. [Dallas, May 1876] (Be); 0. Sanders 113 (Ld, Mi); Shinners 8576 (Sm); R. Spen- cer 5 (Nt); Storm 1019 (Ur); R. Van Vleet 78 (Au—-121656), 120 (St). Deaf Smith Co.: C. L. Lundell 11457 (N). Denton Co.: Dob- bins 16 (Nt); B. B. Harris s.n. (Spring '26] (Nt); Lundell & Lun- dell 9535 (Ld, Mi, N); McBryde 811 (Nt); McCart 1593 (Nt); White- house 15590 (N). DeWitt Co.: M. Riedel s.n. [7-18-l1] (Au). Dick- ens Co,: Parks & Cory 13620 (Tr). Duval Co.: M. E, Croft 20 (C), Sen, [San Diego, 152) (Mi). Eastland Co.: Hodge Oak School 18 (Au, Au); Mrs. G. Robinson 18 (N); Warnock 16376 (Au). Edwards Coe: Cory sen, [11-3-1936] (Md); Cutler 778 (Ba); M. E. Jones 26228 (Du—235289, Gg—237865, La, Mn—-22402); Parks & Cory 6633 (Tr), 9035 (Tr), 20843 (Tr); Shreve 9925 (Fs). Ellis Co.: Cory 39243 (Au); Lundell & Lundell 936 (Id, Mi, N); 0. Sanders 21 (Mi); Shackelford 3 (Nt); Turrell s.n, (Waxahachie, 3/18/1898) (Tu), Erath Co.: Gough sn. [3-12-21] (Au); Hoisington 9 (0k), 208 (0k), 140 (Ok), 242 (Ok); Lundell & Lundell 10167 (Ld, N); Whitehouse 20675 (St); G. Williamson 13 (Nt). Fannin Co.: McCart 2032 (Au—-121638, au). Fayette Co.: Parks & Cory 1007 (Tr); Ripple 51-580 (Au--1216)7). Garza Co,: Ruth 107, in part (N). Gillespie Co.: F. A. Barkley 14522 (Vi); G. Jermy 182 (Ka); Mainland & Barkley 1522 (Au). Glasscock Co.: Cory 42069 (We). Goliad Co.: Lundell & Lundell 10059 (Ld, N); C. B. Williams 13 (Au). Gonzales Coe: Ge Ce Albers 35006 (Au); Friesner 10376 (Bt- 40558); Pladeck s.n, [near Gonzales, 3-30-1910] (Bt—51l6h). Grayson Co.: Anderson & Hood 312 (Ok); R. Bebb 110 (0k, Ur); aoe 17 (Nt); Harold Gentry 51-183 (Au—121611, Au—~121683), a1 71 (Au—121615); C. S. Sheldon sen, (Denison, June 13, 1891) (Al); Sowder 2), (Au); M. M. Taylor 11 (Nt); Woodall sen. [Deni- son, Apr. h, 1930) (N), Son. [Denison, Apr. , 1931] (Ge— 5)» Grimes Co,: Sixth Grade Navasota son. [April 1934] (Au, Au) 5 Wild Flower Contest s.n. [Navasota, Apr. 1934] (Au, Au). Guadalupe Co: Johnston & McCart 5023 (Au--121613). Hansford Co.: cutter 33, in part (Ok), 38 (S), 220 (Ok); Ge Je 5286 (Au—-121652, N, Ok). Hardeman Co.: Whitehouse 1076) (Sm). Harris pest Rifrig Sen. [Houston, June 1926] (Ca—882581, Mi); E. Hall ie in part (Pr); Parks & Cory 8221 (Tr). Hays Cos: G- L. Fisher <20U7 (Go); R. M. King 2615 (Au--101196); Krodel & Warnock 16529 238 PHYTTOLOGIA Vol. 8, no. 5 (Au, N); Stanfield s.n. [July 1896] (N); Warnock & Krodel 19 (Au, Ml, N, N). Hemphill Co.: Tharp s.n- [Red Deer Cr., 6.16.29] (Au). dalgo Cos: Clover 1450 (N). Hill Co.: K. E. Smith s.n. [Itasca, 12~1)-33] (Au). Hockley Co.: Rachaner 131, in part (Au, N). Hood Cos: Blackwell 64 (Nt); E. J. Palmer 6510 (Po--217509). Irion Co.: Wild Flower Contest s.n. [Mertzon, April 1932] (Au). Jack Co.: Me Cart 892 (Nt); Whitehouse 15291 (sm). Jefferson Co.: Brenckle 18069 (N); Lundell & Lundell 11232 (N). Jim Wells Co.: Freeborn & Freeborn 347 (N). Jones Co.: Mahler 1367 (St). Karnes Coe: Je Ce Johnson 805 (Au--121660, Hk). Kaufman Co.: Ashworth 4 (Nt). Kerr Coe: Ve Le Cory 51759 (N); M. E. Jones Son {between Kerrville and San Antonio, April 23, 1931] (Po—188428); Kelting 17 (0k)5 White- house s.n. [June 8, 1929] (Au). Kimble Co.: Straudtmann 5-Ne [Aug 19, 191] (Au). Kinney Co.: Cory 28993 (N); Mearns 127, (Du-- 2h19h), 139 (Du--2h192). Kleberg Co.: G. J. Harrison s.n- [Rivi- La Salle Cos: Mmermann 12 (Au); Penfound s.n. (Cotulla, April 13, 1936] (T1). La Vaca Co.: Johnston & MeCart 5205 (Au--121607) « Leon Co.: Tharp, Turner, & Johnston 51,986 (Au--1216h4). Liberty Cos: Small & Wherry 11781 (N). Live Oak Co.: Tharp sen. [March 22, 1931] (Au, Au, Au). Lubbock Co.: E. L. Reed 3943 (Au, B1-—1,221:8) « Lynn Co.: Rose-Innes & Moon 1052 (Au, N). Mason Co.: Dapprich 7770 (Sm). Maverick Co.: M, E. Jones 26300 (Ca—b79297, Ge—237869, Po--188h26), sen. [Eagle Pass, April 27, 1931] (Du--239762);5 Schott son. [Eagle Pass] (N). McLennan Co.: Pudig & York 6382 (Au); Le De Smith 21 (Au—121610, St), 197 (Au--121609), Li0 (St), 730 (Au—-121619), 905 (Au--121614); C. L. York 46087, in part (N), 46100, in part (Au). McMullen Co.: Thompson & Graham 65 (Au-- 123199). Menard Co.: Bottimer 7.20 (Au--172720); Y- Jones SMe [Fort McKavett, l-1-1931] (Au); Mahoney sen. (Menard, May 1, 1933] (Au, Au), Mills Co.: Egg sen. [Jan. 7, 1931] (Au, N)5 Knowles Se n, [Goldtimaite, h-10-1931] (Au). Montague Co.: McCart 1632 (Nt) 3 Satterfield 21 (N, Nt). Navarro Co.: I. B. Bryant 12 (Nt); Gahr wiler 1) (Au); Matlock sen. [Frost, 5-10-1932] (Au) 5 L. M. Mc Glure 5 (Nt); E. J. Palmer 7821 (Gg--182152). Nolan Co+: Mrse Ue E. Stanfield s.n. (Sweetwater, 4-5-1931] (Au). Nueces Co.: Ae Ae Heller 1365 (Es, Mn--15396, Tu); F. W. Pennell 10337 (N); Tharp aes [11-9-39] (Au). Ochiltree Co.: C. S. Wallis 885 (St). Old- Cos: Lambert & Lambert son. (August 19, 1939] (Up); Lundell & Lundell 11450 (N), 1lu62 (N). Orange Co.: Herb. Univ. Texas Se n. (Orange, Spring] (Au), Palo Pinto Co.: Anderson & Harty 10 (Nt); Harold Gentry 1282 (Au--121662); J. W. Gillespie 5223 (Du-- 210177). Parker Co.: S. M. Tracy 7999 (Au, Cm, Es, N, Vt); Ce be 1962 Moldenke, Monograph of Verbena 239 York 22 (Au--1216)5). Randall Co.: Lundell & Lundell 114)) (sm), Reagan Co.: Cory 53507 (N); Herb. Univ. Texas s.n. Sen. [Best, April 1931] (Au). Red River Co.: E. Adams 10h (Nt). Reeves Co.: reser & Earle 106, in part (Au). Roberts Co.: C. S. Wallis olh-1, part (St). Runnels Co: F, W. Gould 7065 (Au--121626); J. rae Johnson 611 (Au--121608); Lundell & Lundell 10160 (Ld, N). Patricio Cos: F. A. Barkley 16709 (Au, N); Williges 3% isis 179760). Sutton Cos: Cory 39625 (Au, N, » N)5 Geib 4251 (Tr); Gould 18806); Rohrbaugh 390 (Ok) » Tarrant Co.: F. C. Gates 1913) (Ka— $9646); H. Hapeman s.n. {Fort Worth, May onme )3 He es VYolden- ke 695), (N); E. Nash 13 (Nt); Ruth 83 (40), 92 cia), 307 , in part (Cm, Io-~1136], Ka—67107, Mi, Mi, N, N, Sm, Up ge oe (Up), 110, in part is sen. [Fort Worth, ie 19, 1909] (PO=BETET Steinert sn. [May 5, 1946] (0k); L. Fe Ward son. [Fort W iia 9, 1877] (Bc), s.n. [Fort Worth, Sept. 1077] (Pa). Lp Co.: Lundell & Lund Lundell 1 11364 (N); Mahler 1312 (St); Tolstead 5783 (Ca—l8977), 6927 (Au, Ca-—882576, N); S. M. Tracy 8000 (Gm, N);_ Warnock 6352 (Au, N). Terrell Co.: Warnock | & McBryde 14866 (Au— 123218); G. L. Webster 150 (Au—12168) . Throckmorton Co.: Wild Flower Rites! Son. [April 1934] (Au). Tom Green Co.: Cory 50566 50566 (N, Su); F. We F. W. Pennell 10493 (N); R. W. Pohl 4741 (Sm); C. Smith 7 (ok), Tie (0H) wal Wolff 1159 (Tr). Travis Co.: C. C. Albers 33021 (Au), 34011 (Au, Au, Au); Barkley & Copeland 5 (au, N) N), 76 (an, §); Barkley & Rowell hl (Au, N, N), 103 (Au, N)j Berlandier 1Usb9 (Lu); Biltmore Herb. 365kb (S, S), Sen. [6.VI-1903] (S)5 Birge 2957 (au); Bogusch 423 (Me—27), sen. [w. of Austin, 2-1 1926] (Mi); Bogusch & Molby 2927 (Ur); Bot. 23 Univ, Texas son. ee Eddy, 1-30-36] (Au); Brues 1928 (Sm), 19797 (Sm); 5 Sint, Buechner, & Barkley 16T2L9 (Au, u, Ok, S)$ Cohn & B 13252 ao Ca—882568, i); A, i. Ferguson 189 Lotiagywete sen. [Oct. 12, 1900] rete Herb. Hort. Bot. ee s.n, [Austin, June ®, exas F. W. Johnson sn. (Austin, Oct. 6, 1913] (N); J. C. Johnson ree ell & & Lunde. 6826 (1, Mi, N,N), (i a j Mahler 2258 (st)5 St); McWilli 2 (Au); R. H. Painter - 71738)5 R F Faseon 2 Gs} Ripper & parkley Ish2a (Au, N); aoae & psoas 24129 (W—-1369:62) ; Rowell 7058 (Au); Rowell & aus . (Au, N); Rugel s.n, [Austin] (Ii); rs. saith sane [i [austin, (Au, Bt--2,727, St—9207); Straudimann $.n. 1940} (a Au); L. C. Taylor 3034 (ca—697809); T harp 1,090 fn », Ste (Austin, Spring 1920] (Ca—882579, dT son, [Austin, 10.20, ] (Go), s.n. (austin, 4/12/35] (Bt—-25427, N, Sm, St—9208), 8 i aye 2/9/35) (Au, St. 9209), son. (Austin, 31-37] (Ca— 2574), sen. (Austin, 2-23-39] (Mi)» S- en. [Austin, 4/10/41] (Ca- 2h0 PHYTOLOGIA Vol. 8, no. 5 882573, Mv); Tharp & Warnock 1,608) (Au); Tharp & York 50-168 (Au- 121639); Thonpson & Hamilton 3451 (Se--28357); Warnock 107 (Au), 16033 (Au, N); Webster 5 (Au, N); Werkenthin s.n. {Austin, 2.26. 15] (Io—102276); CG. L. York 4900} (Au--1216L0); H. H. York SMe [3-18-08] (Au); York & Wolf s.n. [Sept. 29, '08] (Au); M- Se Young 113, in part (Ur)~ Uvalde Co.: Cory 6133 (Tr); Lundell & Lundell 10263 (14, N), 10955 (N), 10958 (N), 10975 (N)5 Parks & Cory 18103 (Tr). Val Verde Co.: Farr sen. [Del Rio, May 9, 1911) (Up--52679); M. E. Jones 26229 (Po--178358) Parks & Cory 2082 Tr); Perry, Bigelow, Wright, & Schott 830 (C); Warnock & McBryde 5073 (Au—-123265). Victoria Co.: B. J. Palmer 9098 (Du-—205507, S); Shiller s.n. [Dacosta, May 2, 1928] (Au). Walker Co.: Dixon 561 (La, Po--15905, Po--1593k, W--1521886); Warner SNe {Hunts- ville, March] (Hu). Washington Co.: Brackett 253 (Au), SMe ([Apre 1928] (Au, Au), son. [4-10-33] (Ca——852572); Reeves 186 (N)- Webb Co.: Nickels sen. (Laredo, April 1880] (Up—-17085). Wichita Co.: Parks & Cory 13233 (Tr), 16234 (Tr); Whitehouse 9589 (N), 9609 (N). Williamson Co.: Bodin s.n. [Georgetom, Dec. 1889] (Ka); Lundell & Lundell 10369 (Id, N)3 C. L. York 46207 (Au). Wilson Co. Bogusch 764 (Mg--35); Edw. Palmer 1049 (Pa); Thompson & GrS- ham 50 (Au—123190). Wise Co.: Newton 20 (Nt). Young Coot Se Hayes 610 (Ms, N). County undetermined: Capt. Bolton s.n. [April 1895] (Ka); T. Drummond 19) (S), s.n. [Texas] (Sg—68315); Eifrig s.n, (Texas, 192k) (Ca--982582); L. Kern sen. [May 17th] (Av); Lindheimer 232 (Ka), 289 (N, Pa, S), SNe {Soutehrn Texas, go= 51] (N), 11-209 (Ca—24023h); E. Meyer s.n. [184] (M); Ruffner s.n. [N.W. Texas] (Pa); Schlottmann s.n, [1836] (Ah); Teas S-De (So. Texas, June 1926] (Ba, Ba). NEW MEXICO: Dona Ana Co.: Re Se Ferris 169 (Du-—-91550). Grant Co,: E. L. Greene SMe {Near Sil- ee ver City, Aug. 11, 1880] (Pa); 0. B. Metcalfe 1231 (Bl—503l1)5 Mulford 896 (Ur); W. Stewart s.n. [June 26, 1936] (Gg--267605, La); C. B. Wolf 2598 (Du--190979) « Hidalgo Co.: L. Benson 7376 (Po—267660); Tucker 2h (S, Z). Lincoln Co.: Wooton 363 (Ka, Fo —26767). Luna Co.: Treakle s.n. [Deming, April 9, 1911] (Po- 73867). Santa Fe Co.: F. 5. Earle 8, (N); Heller & Heller 35% (Herb. Barnhart 290,] (NN); A» Nelson 11757 (8). Socorro Co.: Re S Ferris 1187 (Du--91552); Ferris & Duncan 2292 (Du—126170); He He Rusby 337, in part (C, Pl—72002, Pr); Vasey sen. [Socorro, May 1881] (S); Wooton s.n. [Devil's Park, Aug. 9, 1900] (We)- Valencia Cos: Condit s.n. (Laguna, July 8, 1912] (Ca—195627); O- Degens® 4853 (N); Nelson & Nelson 2187 (N). County undetermined: Lambert & Lambert s.n. [August 30, 1934] (Up); Mearns 108 (Du--2h167)5 Parry, Bigelow, Wright, & Schott s.n. [below Donana] (C)j He He Rusby 213 (Fe) C. Wright 1502 (T). ARIZONA: Apache Co.: Le Bensc® 1962 Moldenke, Monograph of Verbena 2h1 9573, in part (Po—267652); R. S. Ferris 1218 aie si Co- chise Co.: L. Benson 9467 (Po=-267662) ; Blumer 135, in part (Ka- 60177, Tu); Carlson s.n. [Bisbee, June 2, 1915 1 (Gg-—31h26) ; 0. Me Clark 4880 (B); R. S. ferris 9933, in part (Mi); Fowler aia [Barfoot P Park, 25.V1.1933] (La, La, La); L. N. a Lis (nN); Goodman & Witchocck 1211 (Ca~-.26361, Mn, N); Hilend Th7 (La); Kearney rney & & Pesbles 1423), | (To); Kruckeberg 386) (N); Kusche son. (Cave Creek Canyon, July 1927] (Gg--1h7070); ‘ 3 Thornber s.n. [June 1, 1903] ae Wiegand, Maguire, Richards, & Moeller 11150 (N); C. B. Wolf 2490 (Du—19087h) . Geonetne Oe Soles 902 (N te o tr MacDougal 317 (Ca—104868, Dp, Io-~3876 , Tu); He 337, in part rt (Ca=--104830) ; Sobre 203 Peake ars “a— matshon 52 (Fe); Peebles & Smith 1) 093 (N); Whitehead 1645 (Mi). Graham Co. : Darrow, Phillips, & | & salt 1122 (Tu—10327); ; Kearney & Pesbles 14153 (To); ome 12989 (To); Shreve 4367 (Ca—195155, Fs). Greenlee Co.: L. D. Benson 9467 (Du--326794, N); D. R. God- dard 650 (cess: Navajo Coe: H. S. Gentry 3822 ors hag Livingston, Livingston, & Thornber s.n. [Apache Camp, 8/10/06] (Tu); Luck s.n. [Taylor, . ange 2 1 Bd (Cm). Pima Co,: L. Benson 3322 Fon aeieish, H.W. G 8-30-16 (Cm); Nelson & Nelson 1513 (Ur); Peebles & | Harrison ee (La, S); Pringle s.n. » [April Ws 1881] (Il). Santa Cruz Co.: He H. W. Graham 7-1)-2 (Gm); Loonis, Peebles, Harrison, & Kearney 1348 (To); Peebles & Fulton 1,89 — (To). ¥ Yave vapai Co.: L. Benson Benson 7460 (Po—267661) . MEXICO: Chihuahua: pote 35 (Au); M. E. . Jones s.n. (Sept. 17, 1903] ON he Ae Lee L2 (au, N); F. W. Pennell 19252 (Me, N)5 Tomsend & 139, in part (Ca--138810, Me, N, Po—71126, Ur, Vt); S. 5S. White 2393 (Fs), 2543 (Fs); E. "Wilkinson sen. (Sept. 26, 1885] (Al). Coahuila: F. A. Barkley 16057 (Au); B. G. Marsh 1121 (St), 2146 (St); Edw. Palmer 1050 (Pa); Stanford, Retherford, & Northcraft 241 (N, Ur); Wynd & Mueller 150 (Mi, S, Ur, Ur). Durango: Forrer S:n. [September and October, 1881] (I, N)3 H. S. Gentry 6870 (ak—22789), Je He Maysilles 7039 (N), 7057 (im, N), N), 7472 2 (ui). uanajuato: G. L. Fisher sn. [July 28, 1926] (Mg--3h). Jalisco: Bartley, Webster, & Rowell 7659 (au—167038, N); L. I. Davis s.n. (Jaday, July 6, 1989] (Au—171900, Au—171909). México: Gs L. Fisher’ 8.n, [July 2h, 192k] (Mv); Matuda 2638h (Cb). Michoac4n: Hinton 13125, in part (Mi). Puebla: Barkley, ey, Rowell, & Webster 2u58 (Au—123215, N); L. 1. Davis 226 (8), 233 3 (N)« Snteor pe ®). cur nag CULTIVATED TED: Belgium: Herb. aed Bot. Brux. $.n. — @: Boucher s.n. [1819] (Du-—166h90). Italy: Herb. Harvey 8. n, a i AL 1819] (Du—166491), s.n. [he R. P., julio 1843] (Du—r6¢ 92). Massachusetts: Re Ee '. Torrey sn. {Amherst, Sept. 6, 2h2 PHYTOLOGIA Vol. 8, no. 5 190] (Ms). New Jersey: Go H. M. Lawrence P.237 (Ba), P.353 (Ba). New York: H. N. Moldenke 1126) (N, N, N). South Dakota: Barr 101 (Ba). West Virginia: Davis & Davis 6259 (We). LOCALITY OF COLLEC- fie UNDETERMINED: F. V. Hayden s.n. [Yellowstone, etc., 1853--L) Dt). VERBENA BIPINNATIFIDA var. LATILOBATA Perry, Ann. Mo. Bot. Gard. 20: 325--326. 1933. Synonymy: Verbena bipinnatifida var. latiloba Perry ex Molden- ke, Alph. List Invalid Names Suppl. 1: 22, in syn. 1947. Verbena bipinnatifida latilobata Perry ex Moldenke in Chittenden, Roy. Hort. Soc, Dict. Gard, 6: 2209. 1951. Literature: Perry, Ann. Mo. Bot. Gard. 20: 311, 325—326, & 355. 1933; Sperry, Sul Ross State Teach. Coll. Bull. 22: 1941; Moldenke, Known Geogr, Distrib. Verbenac., [ed. 1], 12 18. & 101. 192; Moldenke in Lundell, Fl. Texas 3 (1): 18 & ho. 19h,2; Moldenke, Phytologia 2: 127. 19h; Moldenke, Castanea 10: . 1945; Moldenke, Alph. List Cit. 1: 143, 16h, 169, 193, , 2h2, 2h, 270, 27h, & 290. 1946; Moldenke, Alph. List Inval- id Names Suppl. 1: 22. 197; Moldenke, Phytologia 2+ 37. 197; 1593 Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6+ 220? & 2210, 1951; Moldenke, Phytologia 3: 167. 19513 Moldenke, Résumé rages hah Rl, 38, 223, 358, & 470. 1959; Moldenke, Résumé Suppl. his variety differs from the typical form of the species in its }cleft (trifid) leaves, with the segments remotely incised or lobed, the ultimate segments much broader than in the typical equali: the calyx, the floral and fruit characters in general similar to those of the typical form, the corolla described as light-rose, sags lavender, or purple, and the connective-glands (acco T Car (no. 906) between San P and Fronteras, Sonora, Mexico, be~ tween elly hillsides, burned-over pinelands, subalpine meadows, rocky slopes, eni-timbere woodlands and waste places, along roadsides, at the edge of ounsi sane and in rocky terrain on canyon slopes, at altitudes of 1350 to 8 a 1962 Moldenke, Monograph of Verbena 2h3 feet. Brown found it to be "sparse in pasture" in Brewster County, Texas. Ferris reports that it forms a groundcover with Helenium hoopesii A. omer perphinn, Artemisia, and Pteris in Cochise County, Arizona; in the same county Bluner found it in "rolling andesitic pineland reais lumbered", Tucker encountered it on steep southeast-facing slopes in shrubby growth of Quercus ariz- onica Sarg., Q. emoryi rag and Pinus cembroides Zucc. Mueller begged it as “spars se slopes" in Chihuahua. e variety differs chiefly re the typical form of the spe- cies a its foliar characters. It has been collected in anthesis n fruit in February, April, and June to October. Material in febarks has been misidentified and rare as V. aubletia Jacq., V. bipinnatifida Nutt., V. canescens H.B.K., Me ciliata Benth., V. elegans H.B.K., and V. wrightdi A. Gray. Davis enpha- sizes that it has "perfectly upright growth", but Hinton says "general umbent", The Bush 4029 ma piesa” from Missouri is from "introduced" plants, according to the collector, but I as- Sume that he does not mean cultivated; ape Perry cites this collection, as well as Earle & Earle £26 526, as typical V. bipinn- atifida, The Whiting & Sanders rest from Oak Creek Canyon, cited below as from Yavapai County, Ari actually have Come from Coconino County, and the M. Ee “jonae 6 ois {Santa Rita $+, Aug. 2h, 1903], cited from Pima County, may possibly have rio collected in Santa Cruz County. The C. Wright 1503 cited by Miss Perry "in part" apparently is a mixture with | ve , tumidula Perry and V. wright4i A. Gray. She cites the following 1) speci- mens not as yet seen by me: ARIZONA: Cochise Co.: L. Ne Ne coodsing 883 OF Griffiths 1,833 (W); T. E. Wilcox s.n, [Fort Huachuca, Sept. 189h] (Ww). Pima Co.: Griffiths & Thornber 159 (W); M. E. Jones s.n, (Santa Rita Mts., 2) Aug. 1903] (W). MEXICO: Chilma- ma | Nelson 856 (G, W); C. Wright 1503, in part (G, N). Durango: Forrer s.n. sen. [Sierra Madre west of Durango, Sept.—Oct. 1881] (F); Gareia 331 (W). Sonora: Hartman 880 (G), 906 (G—type); Murdoch S.n. S.n. [Cananea, 20 Aug.—1 Sept. 1914] (F). Miss Perry cites a Mearns S.n, (Flagstaff, 17 June 1887] on the basis of a specimen in the New York Botanical Garden herbarium — the specimen there, however » is Mearns 1 159 and is certainly this variety, but was annotated by her in 1932 as "Te bipinnatifida Nutt." In all, 9 herbarium specimens have been & Citations: MISSOURI: Jackson Co.! Bush 1,029 (N)- ORLABOLA: Greer Cows Logan 12h 12h (Ok). Murray Cot 3 Ge We W. Stevens 14 (N) TEX- AS: Brewster Co.: Ge G. W. Brown B.107 (Au); S perry 1492 (Om). Dallas Gos: Bs Jones 6 (Ok). Hidalgo Co.: Co.: L. I. Davis 199 9 (N). NEW MEX~ ICO: Chaves Coe: Earle & Earle 526 (N). Grant rant Co.: Blumer 60 (N). Otero Co.: Spalding 28 (Tu)~ County undetermined: C. Wright 1503, Part (Ca--2h:7679), 1602 (T). ARIZONA: Cochise Cos: F. A. Bark- Aey 1NA611 (au, ca-—71,3022); Blumer 1345 (N); Re S. Ferris 9933, 2hh PHYTOLOGITIA Vol. 8, no. 5 in part (Rs—26916); Fowler s.n. [Barfoot Park, 26.VI.1939] (La); L. N. Goodding 37 (N, Tu), 251 (w), 883 (N, Tu); Goodman & Hitch- cock 1211 (Gg--183908); L. Henderson 26 (Au, au); Hilend 678 (La); M. E. Jones 8.n. [Sept. 3, 1903] (Po--71156); Kearney & Peebles 5117 (To); Kusche sen. [Cave Creek Canyon, June 26-29, 1927] (Gg—-17069); W. W. Price son. (aug. 25, 1893] (Du--97116); Je Je Thorrber 702) (Tu), sen. [Fort Lowell, April 25, 1903] (Ca-— 882580), s.n. [Dragoon Summit, Aug. 19, 1905] (Tu); T. E. Wilcox Bone (Ft. Huachuca, Aug. 1893] (C). Coconino Co.: Fulton 9672 (To); Mearns 159 (C, N). Gila Co.: Dunkle 5226 (H—L91); He We Graham 7-26-3 (Cm); Peebles & Smith 1093 (To); E. Sanders 803/ 2339 (To); J. Whitehead 1645 (N). Graham Co.: Humphreys Anderson, Rinehart, & O'Donnal 712 (Tu); Kearney & Peebles 9751 (To), 9802 (To); Pultz & Phillips 18h9 (Tu—16175). Pima Co.: M. BE. Jones 9» n, (Santa Rita Mts., Aug. 2h, 1903] (Po--71159, Po—71160); C- Be Maguire s.n. (March 2h, 1935] (N); J. J. Thornber s.n- [Fort Lo- well, June 1, 1903] (Tu), s.n. ([Tanque Verde, June 8, 1913] (Tu). Pinal Co.: Porter, Peebles, & Harrison 171 (To). Santa Cruz COe? Stewart son. (Patagonia, Sept. h, 1936] (La); H. S. Gen — — & try 3295 (Ak--2313h); Kearney & Peebles 10130 (Gg—267609, Po 208646), 10548 (Mi, To); Peebles, Harrison, & Kearney 4Su9 (To); ee M. E, Jones 5-Ne [Sept. 11, 1903] (Po—69327), son. [Sept. 16, 1903] (Po——69 320); Le Sueur 305 (Au), h61 (Au); C H. Mueller 3347 (Ca--720107)5 Fe Me part (ile); S- S- White 2393 (Iti), 2456 (Mi), 2543 (iH)- Durango: F, W. Pennell 10264 (D-—-733977, W-—16)1017) « Waichoac4n: Hinton 13125, in part (Au, N, N, N). Sonora: Donnelly 5 (Ca--16398) , 6a (Po—-)\003); Hartman 906 (Ca—92h053——isotype) ; Tucker 3h78 (Du—399532, Gg—13693)5 S. S- White 3131 (Mi, Mi), 3261 (I), 3731 (Mi). State undetermined: Née 109 (Q)- CULTIVATED: Belgi- um: M, Martens s.n. (Br). Mexico: Halbinger & Reko 313 (N)- 10- R CALITY OF COLLECTION UNDESIGNATED: Fremont s.n. [Expedition to California, 1845~—7] (T); Herb. Bonpland s.ne (S)- XVERBENA BLANCHARDI Moldenke, Revist. Sudam. Bot. h: 18. 1937 Synonymy: Verbena angustifolia x hastata Blanchard ex Molden- ke, Revist. Sudam. Bot. h: 18, in syn. 1937. Verbena angus ti fol ia x hastata Hill ex Moldenke, Suppl. List Invalid Names 7, a2 syn, 191. Verbena hastata x angustifolia Hill ex Moldenke, Suppl.List Invalid Names 8, in syn. 191. Verbena blanchardit Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 2209 & 1962 Moldenke, Monograph of Verbena 2hs5 2210, sphalm. 1951. Verbena hastato-angustifolia ~~ in herb. ure? Kanda, Bot. | Bot. Gaz. 69: Sh--71, pl. 6, fig. 8 & 9 S. 190. 1940; Moldenke, Suppl. List Invalid Names 7 & s. a Mol- me Geogr. Distrib. Verbenac j & 101. 192; it a a Alph. List “Invalid Names ig ’ ire ihes Moldenke, Phytologia 2: 68, 72, & 11h. 195; Moldenke, ot List Cit. 1: 30, a 3. 43. 19h6; "anaes: Phytologia, 2: 326. my Moldenke, Aaphe List In Invalid Names Suppl. 1: 2h & 1947; ae aE bs ra « Suppl. 10: 2h2. magi ie A & r x Mol- up - life “1988; Moldenke, Alph. . 23 359, 5, ". 638 (sl), 38 828 & 879 (19495, and i 08. ‘isles Nol den e Zi : pated b. Verbenac., ed. 09 & ans was Moldenke in Gleason, New Bri sag & eg Tainst gf oe » 32 126, 129, & 130 (1952) and pr. 2, 3: 126, 129, z "130. "1568; ‘Sienna Am. Midl. Nat. 59: 3h3. 1585 elienke, Résumé 5, 9, 15, 18, 20, 26, 223, 357, 365, 366, & 470. 1959. Dilsetratt one: Kanda, Bot. Gaz. 69: pl. 6, fig. 8 & 9. Bes a Moldenke in a pom New Britton & Br. Tllustr. Fi; pre 4, 5% 130 (1952) and pr. 2, 3: 130. 1958. e of this hybr brid was ee lieutenant Ruffner at Prieta, Rockbridge County, Virginia, and is deposited in the Torrey Herbarium at the New tide Botanical Garden. It is ed in honor of Sec Henry Blanchard (1850--1922) — botanist and educator, well-known for his work on and o New England plants. “tha name XV. blanchardi has been a roposed for the natural hybrid which occurs between V. simplex Lehm. and VY. hastata L. where the — of these two species cies overlap in the eastern United state , and for artificial hybrids produced ne ia between rend two species. The cejaepelac grey resembles "parent", senotinet 5 the somes in gen The Rcks ox = gene nerally b ader, more acute, and with coarser Pubescence and larger Soith han in V. simplex, more narrowly lanceolate or oblong-lanceolate than in V. hastata and never in- cised nor hastate. The fruit is produced sparingly and irregu- ly, whereas in both "parents" fruit production is usually heavy. Kanda (1920) gives a nyt 9 discussion of the hybrid. Was apparently intvcdnied to (or originated in) cultivation tn 1854, but the Herb. Hort. at. Berol. S-M- (185)] specimen Was annotated as Vs hastata L. by Miss Perry. The type of ve - hastato—angustifolia was collected by Michael Schuck Bebb at Fountaindale, Winnebago County, Illinois, and is deposited in the Jesup Herbarium at Dartmouth College. The "Ver- bena simplex x hastata" of Gates, Fl. Kans. 190 (1940), if one” may Judge Adige ee the herbarium specimens which he has s0 annotated, § a mixture ve simplex ex Lehm. and V. simplex vare eggerti Moldenke, I neve not as yet seen any ‘Specimens | so annotated lobated by 2h6 PHYTOLOGIA Vol. 8, no. 5 Gates that are the hybrid which he suggests. It may, therefore, be that the Kansas records which he gives -~- an which I have Do actually do sot 4 apply t o this ert Similarly, the "Verbena angustifolia x hastata" of Jacobs is typical V. simplex x Lehi. Collectors report finding xV. blanchardi in n dry iry fields, grassy ground, and on roadside banks and railway generals $ flower- ing from June to August and fruiting in June and A e They re=- port the common names "blue vervain" and "false Recent -- the former also applies to V, hastata L., and the latter to V. hasta~ ta and to V. litoralis H.B.Kk. In herbaria it has been variously misidentified and distributed as V. angustifolia Michx., Ve engel- blue or violet to lilac-blue. In all, 15 herbarium specimens, in- para the types of all the names involved, have been e ed Citations: VERMONT: Bennington Co.: W. H. Blanchard 162 (N). VIRGINIA: Rockbridge Co.: Ruffner s.n. (T--type)- ILLINOIS: Cook bee E. J. — 161.1898 (Ur), 162.1898 (Ur, Ur). Winnebago Co. . S. Bebb s.n. [Fountaindale] (Dt). Stony Island: H. He Smith ae (Ga=88583), 5982 (Ca-—882823). KENTUCKY: Warren Cos: Se Ne Price Se [Bowling Gre. Green, July 29, 1899] (N). OKLAHOMA: Latimer Cos: F. A. Barkley 1219 (Ok—-10312), sen. [6/20/28] (Ok—-10313) Marshall Co.: G, J. Goodman 5418 (St). Wagoner Co.: Re Bebb 4 (Ok). CULTIVATED: Germany: Herb. Hort. Bot. Berol. sen. {185] ( reams OF COLLECTION UNDESTGNATED: Collector undesignated VERBENA BONARIENSIS L., Sp. Pl., ed. 1, 20. 1753 [not V. bonari~ ensis Rendle, 190h,, — Schau., 19 ° = Syno: : Verbena bonariensis altissima, lavandulae canarien- sis spica multiplici Dill., Hort. Eltham, 06. 1732. Verbena bonariensis. Lavandulae canar, spica multiplici Dill., | Hort. El- tham. pl. 300, fig. 307. L732. Verbena foliis lanceolatis, flor- ibus congestis fastigiatis L., Hort. Hort. cliff. ll. 1737. Verbena bonariensis altissima, lavandulae canariensis spic& multiplict Dill, ex L., Hort. Cliff. 11, in syn. 1737. Verbena tetrandra, spicis fasciculatia: foliis lanceolatis amplexica exicaulibus L., Hort. Upsal. 8. 1748. Tastand bonariensis altissima, aves ee canar- iensis folio, spica lavandulae Dill. apud L., Sp. ed. 1, i, 20, in syn. “1753. Verbena bonariensis altissima spice sale Dill, ex Sabbat in Martelli, Hort. Roman, 3: 10. 1775. Verbena elongata Salisb., Prodr, Tle 1796. Verbena cog hotoma a Moench, Suppl. Meth, Plant. 131. 1802. Veronica bonariensis, ‘altissima, 1962 Moldenke, Monograph of Verbena 247 lavandulae canariensis folio, spic& lavandulae Dill. apud Poir. in Lam., Encycl. Meth. Bot. 8: 549, in syn. 1808. Verbena spicis brevibus, subfasciculatis; foliis oblongo-lanceolatis, amplexi-_ caulibus Lam, apud Poir. in Lame, Lam., Encycl. Meth, Bot. 8: 549, in syn. 1808. Verbena (bonariensis), tetrandra, spicis fasciculatis; foliis lanceolatis, amplexicaulibus L. apud Poir. in Lam., Ency- cl, Meth, Bot. 8: 549, in syn. 1803. Verbena quadrangularis Vell., Fl, Flum. 16 (1825) and Icon. 1: pl. 39. 1827. Verbena bonarien- OS Walp., Repert. 4: 19, in syn. 1845. Verbena corymbosa Hort. ex Walp., Repert. 4: 19, in syn. 1845 [not V. not V. corymbosa Cham., 1832, nor Relh., 197, nor Ruiz & Pav., 1798]. “Verbena bonariensis y longibracteata Walp., Repert. ): 20. 1845 {not V. bonariensis var, longibracteata Kuntze, 1893]. Verbena bonariensis Dill. ex Goncalves de Cunha & Goncalves Sobrinho, Rev, Fac. Ciencias 3: 10 & 43. 1938, oe fonariensis L. ex Moldenke, Suppl. List esate Trib. Para, [Veli oz0a] 12: 45, sphalm. 1944. Verbena bon- atiensis L. ex Moldenke, Alph. List Cit. 1: 28h, sphalm. I9h6. Berbena bonariensis L. ex Moldenke, Alph. List Invalid Nanes Suppl. 13 2, in syn. 1947. Verbena bonariense L. ex Moldenke, Alph, List Invalid Names Suppl. 1: 22, in syn. 197. Verbena hasta L. ex Moldenke, Alph. List Tnvalid Names Suppl. 1: 2h, in syn. 197. Cymaria idjenensis Koord. ex Moldenke, Résumé 276, in syn, 1959, Verbena bonaerensis Larrafl, ex Moldenke, Résumé 358, in syn. 1959. Verbena bonariensis Vell. ex Moldenke, Résu- = 359, in syn. 1959. Verb Verbena bonariensis f. umbrosa Osten Woldenke, Résumé 359, in syn. 1959. Verbena littoralis H. Betch. ex Moldenke, Résumé %68, in syn. 1999 [not V. littoralis Humb. ad ol6] H. & B., Hs nor E.B.Ke, as, nor Kunth, 10h7, nor Litera : Dill. . Eltham, 06, pl. ae ee 387. 1732; “ag ies cliff. eth "1737; A. van Royen ree gal ee "17h0; L L., Hort. Upsal. 8. 1785 Le, SPe are 1753; Kniphof, Bot. Orig. Herb. Viv. pl. (2 28h). inst 2, 8 eal Denom. Linn. pl, 300. 17h; Sabbat in Martelli, Hort. Ronan. 3: ©, pl. Sh. 1775; Christm, & Panzer, bei ecg oe S: 125-~126. 1779; Gaertn., Fru 15, fig. 1. 1788; Lam., Tabl. mncycl. yéth. Rote ~ fritases%) a 57, pl. 17, fig. a 1791; Re As Sila Prodr. ven -, Spe Pl. Lele “118. 1 1797; Thunb., Prodr. Pl. Cap. 96. 1800; Moench, ae ant. 131. 1802; Poir. in Lam., cl. Méth. Bot. 8: reiet ee 1808; Mirbel, Ann. Mus, Hist. Nat. Paris 15: 260, pl. 1h (2), fig. "4. 1810s, J. Sm. in Rees, 2u8 PHYTOLOGIA Vol. 8, no. 5 x «et no. 6. 1817; Steud., Nom. Bot. 1, 87h. 18213 Thunb., a. , ed. somites, uu? & 799. 1823; se "Pl. Flum. 16 "Br ’ Prodr. 11: 541-52. eu; C. Gay, Hist. Fis. Chile Bot. 5: 21-- os Higa d Schau, in lf Fl. Bras. 9: 187 & 189. 1851; Re Ae , Linnaea 33: 195. e. i86h; H. . Haws Pl. 19h. 18673 i = *P ° Univ FS ee zh 70; Ulrich, Internat. 2 gent.]: ger ae Vell., Fl. Flum. repr. Linn. . Bot. 21: 230. 188; C. Be Clarke in Hook. f., Fl. Brit. Tod . "56s. 1885; Lillo, Fl. Tucuman 9. 1888; A. We 18893 Ball, Journ. Linn. & Vail, Ann. (Plant, Hassler. 9: 197]. "1902; Speg., - C 53 (Nov. Add, Fl. Patag, 2: repr. 72]: 2h9. “1902; Small, Fl. : Conserv Boiss., sér. 2, 4:1057. 190); Briq. in Chod. & Hassler Hassler. 10: 479. 190); Briq., Arkiv Bot. 2 (0): 10. $905 wae lo W. B. Scott, Rep. Princeton Univ. - Pat 8 (2): 68h. 1 05; Agric. Gaz. N. Ss Wales Lit 800. 1906; Fe M. Ba ? _ & Pois eensl. 143. : schr. ees d. Wissen, Math.-nat. 79 (1): 29h. 1908; Hayek oy BO Jahrb. 2: 1 908 ; Hassler, Fo : 1909; Reiche, Fl- Chile 5: 283 & 28h. 1910; H. H. W. Pearson in Thiselt.-Dyer, Cap. 5: 207--209. 1910; Hicken, Chloris Plat. oi pig 195+ 1910; Gerth van Wijk, Dict. Plantnames 1397—1398. 1911; G , Ann, Mus. Colon. Marseille 9: 206. 1911; goFke southeast. U- S., ed. 2, 1010, 19133 Hauman-Merck, Anal. Mus. Argent. Hist. Nat. Buenos Aires 2h: 415. 1913; F. M. Bailey, Compreh. Cat. Queens]. Pl. 362. 19135 Fyson, Fl. Nileiri & Pulney Hill-tops 2: 21. 1915; Hicken, Physis 2: 114. 1916; Britton, Fl. Bermuda ee ; Sanzin, Anal. Cient. Argent. Buenos Aires 88: 96, 98, llh, at : 13. 4 Molfino, Physis 5: 285, 1921; Pellett, am. ed. aes 19233; Rep. Bot. Exch, Club Brit, Isles 1922: "ihe 19235 tearn, Flora Batava 27: pl. 2093. 1925-1930; J. M. Black, Fil. 1962 Moldenke, Monograph of Verbena 2h9 South Austr. 3: 79. 1926; pret Illustr. Fl. Mittel-Eur. 5 (3): 220. 1927; Seckt, Rev. Univ. Nac. Cordoba 17: 90. 1930; Herter, Florula 105. 1930; Stapf, Ind. Lond. 6: 429 & 431. 1931; St. John H Eastwoo 1138. 1933; J. A. Harris, Physico-chem. Prop. Plant Saps 218. 1934; L. H. Baile ey, Cat. Florists Handl. Verbenac. mss. 1935; we 5 632. Dermen, Cytologia 7: 161-163, fig. 19, 16h, & 172. 1936; Herter, Revist. Sudam. Bot. h: 186-187. 1937; Cory, Texas Agr» Exp. Bull. 550: 88. 1937; Gallinal, Bergalli, Campal, Aragone, & Ro- sengurtt, Stud. Nat. Mead. Urug. 112. 1538; Moldenke, cuit. Pl. 35. 1938; Gongalves de Cunha & a So obrinho, Rev. Fac. Ci- encias 3: 10 & 3. 1938; Moldenke, Annot. & Classif. List 1 1939; Herter, Beih. Bot. Centralbi. 39: is. 1939; Moldenke, Darwiniana 3: st Comm Moldenke, Suppl. List Invalid Names 8. 191; Gongalves de Cunha Anal, Inst. Fitotéc. aaa eet a 18-20. 19h —— Estud. Prad. Nat. Urug. 3: 235. 193; Molde: ae oe Sa e0gr. Distrib, Verbenac, Suppl. 1: 1 & 2.1 1 9433 A Re Schultz, Introd. 3 943; Raimo 2,22; th, & 27. 1973 Moldenke, Phytologia 2: 326, 328, 335, 3 115, 118, 120, & 121. 1948; ge Alph. 356, 357, 360, 363, 367, 369--311, 3 375, 399, 02, ik? 426; 438; 439, 4s2, 455-158, N67, 76, 78, 79. ur 7. st : 403, os » 483 487, Loh, 95, 5 ’ 2 5 95 520, 7, 52, 2 5 3 551, 554, 559, 570, 571, 57h, 579, 584-586, , 626, 628, & 638-611 (1948), 3: 655, 662, 663, 666, 669, 673, 676, 18, 72 9 1335 7. Js 75, 7h7, 79, 750, 765, 766, 7 8, 7155 779, 781, of > 9 812, 813, 832, 8h3, 86, 847, 89 852, 857, 860 ’ 871, 87-877, 896, 897, 899, 9 932, 93h, 935, 91 +coe>e rks , & 968 (1949), and h: 979, 1, 992, 1069, soo, “beh, 508 094, 1107, 1116, 1117, 1119, ; nil, 1163, 116k, 1166, 1168, 1172, 1177- 1180" 1183, 1192, aoe 1199, ee 1207, 1212, 1215, 1233, 1237, 1239, 12h 1298, 1300, & 1302. 19 9 1213, 12h, 1256, 1257, 1283, 1290, 1296, if aumin, Bull. if 95 Paris, sér. 2, 21: 261. oe Boelcke & Echeverria, Lilloa 18: 2h1, cuadro 8'& & 9. 1949; Waterfall, Rhodora 51: 20 & 27. 1993 Voldenke e, — Geogr. Distrib. Verbenac., [ed. 2, Shy 39, 24,23, 26, 27, ay 47, 61,°79 99-101, 338, 107,108, 119, 122——125, 128, 130, 1b, 15)--156, 163, & 197. i9h9; He Ne & Ae = toldenke, Anal. In +t Biol. Mex. 20: 12. 1949; Cabrera, Lilloa 20: cuadro III I & XVII. 19h9; Delect. Sem. Hort. Bot. Pisa 19k93 13. 1949; Moldenke, Am. 7 Wild Fls. 293 & SO. 1919; Moldenke, Phytologia 3: 72, 7h, 135, 136, & 11 (1949) and 3: 289, 291, & 378. 1920; N. Y. Bot. Gard Seed Exch. List 1951: (1950) and 1952: 4. 1951; et Trib, Farmac. [Vellozoa] 19 (10): 166. 1951; Molde oc. Dict. Gard. 6: 2208--2210. on Molden son, New Britton & Br. Illustr. o 1, 3: 126--128. 19525, bo, Sellowia 6: 60, 8h, & 123. en ad "Mo idenke, Phytologia 9h. 195k; Moldenke, Inform. Mold. Set 48 Spec. [h]. 195h5 yoI= denke, Inform. Mold. Set 9 Spec. 3. 19543; Moldenke, Journ Calif. Hort. Soc. 15: 80. 1954; Thorne, Am. Midl. Nat. 52: "53s his 1954; Amaral Lisb8a, Aspect. Fl. Ruder. Ouro Preto 3. 195); Am aoa Liebe, Anais V Reun. Anual Soc. Bot. Bras. 1h0. 19563 Mol- , Inform. Mold. Set 51 Spec. rs "1956; Moldenke in 8. 1956; Vaughan, Ga rd. Illustr. 1958: 5h 9: 33h, 3L6 Gard. Illustr. 8h: 51. 1959; Reitz, Sellowia ll: 57 & 13h. 1959; Moldenke, R6- sumé 6, 10, 11, 13-15, 18, 22, 23, 26, 28, 32, 33, 9, 55» 6% ? 1 1, 191, 202, 203, 206, 0-212, 3; 276, 358, 359, 363, 36h, 366, 373, 376, 377, 470, & Lid. 1959% W enowiL 223, 237, 2: 221. 19593 Moldenke, _— ne re i * 2 & Kunkel, 12 (1959) and 2: 1, 5, 10, & ll. "19603 Ange ly, (1960) and 17: 6. "1961; Vaughan! s Gard. pnt ey 86: ners 19615 ; 78 1962 Moldenke, Monograph of Verbena 251 oS ea Suppl. 3:°3, 7,8, 1%, 1k;AS, 25, , Oy my, & Fama & ‘alinstretions: Dul., Hort. Elth 300, fig. 387. 1732; Kniphof, Herb, pl. [28h] (in colér) ta787 and 2: pl. 1169 (in color). 1763 “pill., Hort. . Add. Denom. Linn. pl. 300. ; it : . obo LTT “rege Fruct. & Sem. 1: oy 66, fre se 17885 Oo Soc. Cient. Argent. Buenos Aires 88: 115. 1919; Stearn, Flora Batava 27: pl. 2093. 1925--1930; St. ust HOT Basin I \ j.P 5 MeN 5 U R 1 SHERIDAN or N PETROL Danson . N oe ber oe 1 TASCA ie SiR conn T T H Di A K— M s MERCER O s¥ TRAIL » ise a ‘ie N Panag s BILLINGS FOSTER crices RORMAN prerry v 4 & J Tittsom | "ACMA Mussa Sel Poseaye | OuvER Bumeic | moor | stur: sTeeue cxsan) MUBBARD fee ‘4S +, a ta o.De z NES cass one (a Lovee VALLEY CustEn ‘STARK cur BECKER Q SiLvER Aan |p, wigaur prop MORTGA — — ; '. r SEA 80m! fol PReASURE ae ALLEY : ont oxrtt o ‘parhigll ‘A Mn 2 SWEET Gauss YElLowsr S.0PE HETTINGER TRANT . abc a CARLTON . x wan —— pu CHIPPER \ Mots one Bawons | Loan Ta mOURE TaNSOM fda om DOUGLAS iY coors =a XY Seno cred ww BOWMAN wks ‘bacel! LA ARON DECKINSOM| DELTA . Pu Pm a on “=| M I N-N~ES => 7 . McINTOSH DICKEY SARGENT ooo roROW WING T ie sac =, Pema DAWSON BUFO | wat York | Seward was [ase [ous [non |e snes @ 5 POR woDDFORD aT erm % wor we Aen on . ye er provost cass s CARROL a copiot ton cx a ‘ROCKLAAD s C s s PHILLIPS Geamont| PAGE T Tavton Recor] O€CATUR | warwe —[47PAn0056) DAVIS pate: = iu foe — cra (aad TICKING bogrmave | GUERNSEY 7 somensel o* CASE aie REMON TAYLOR |RINGGOL Buren [LEE Py a i at son] ums i pert 2S " PASSAIC ee mare) GRAND WoRan My nAYES | FRONTIER ——}__| OTOE Feu Sot) piri) NTON ST TIFTON v un A Ce ane ‘BREN PHELPS Txeanney GAY [ruumone] SALINE ° ¢ a CT eos WOR =. ee wom — BOULDER WASHINGTON J YUMA murat SCOTLAN! e CHAMPAIGN s on recrtth * uJ AoA Gace Trounson TERSONG nooAWAY.[ woara Tuannisonh MERCER) PUTA ~ ~ a ie FRANRLIY arnit ORNL NORE am on 8 { get L Dag SuNoY wiTcHcocK | RED] FURKAS “ CUARK DONOUGH, wason | LOGAN Tg wiTT y 4 WANE sorrento ar woul vars foots . willow HARLAN | FRANKLIN wucnous | THAYER ‘GENT! SULLIVAN Foun KNOX ANCOCK Tec HUYLER. wont- [BOONE jw” - Coal FUASHINGTON Berit onl om oso 4 orm air ADAMS: LE JEFFERSON pom sn GRUNOY LEWIS TONKS ad ATT sham coment been macOen? HENRY . ayer & op — oa ae) prc : é : = re | grembes . WEBSTER Y ENS. I must a wt i ‘ge 7 CHEYENNE | Ra RICHARDSON aNOREW DAVIESS on ben EDGAR wanionl 3 i saaata] CLINTON OSS ‘cm SUMMIT, : . WLiNS | DECATUR WACOM = j— A te TON} ATHENS ae, PIT PARK rT = wonton] pups | sir JEWEL REPUBLIC ped aa weney: ues we MACON Pow" rate; G rota oe 7 FRAALIN: gy guTLER WIGS gry Cy i pe on 4 med — na PIKE aos C 160 er oan a ss we UNCOLN [kr oie Deel ORGAN — Tao, KE ° (=. ® cs! = SHERMAN [THOMAS aon caro CHARITON | me 0) var CLA cote H 5 |e u cst mr — (@) SHERIDAN | Granam | Rooks T osboene Twit 7 ITCHISON evcmaman CARROLL ANDOU A GREENE CHRISTIAN mean owen H | Eo wont ® eee unison D I Cun LE orrawatomie| AS ere! MONROE rere ee Pets boa 5 3) acount “3 ‘oe wsot fas > TEER "RUDRA = FAYENTE. [ersoncuaie] JASPER jmonec DT wAcKSOn ae 3 aoa ean - oy M son ¥ he S&S @ ° EL Pas wi OTTAWA INE BOONE UNCON LARENCE Lisson |e Fra Kom att oc va, om aur — CHEYENNE MLUACE | LOGE Ove Taco ae ee boos A NCIMARD a JERSEY Tro nant 08M RGK S. faxvon fated Meuse wen ry cHarees ) FREMONT : ci eae _———— fo - a end muse” | scott kK * << sans -. > , TOE J . — WABAUNSEE Bac —— = ot. s MARION bs pi fe Eohaatay saitestcgr ghee PUEBLO - creeuey | wicmta [scart ELLSWORTH. MORRIS OSAGE |] DOUGLAS | JOHNSON HT cass cooPER bicep + ame I - " a) snet ot ROWLEY | KIOWA NESS SALINE LYON FRANKLIN | MIAMI st Louis frees Bo ¢ rg va ance RasOaLE CUSTER s K £ N S , A S E ce im OC a Ad ea ue ee te “ a BATES aor wHTE TA | ato! y * uf MINERAL oreo PROWERS HAMILTON [KEARNY | FINNEY mocha) att BARTON a COFFEY. Ginoer: URN rss bee re) ~ hor rer Te facet \d Kouk J STAFFORD McPHERSON occ ot R10 GRANDE Huei WARYE ton M4 ” Mawoan RAND RENO spa ant o Arne os hn TT es EDWARDS we GREEMHOO0 Faroopson| ALLEN | BOURBON Il VERNON teres om a eine ARCHULETA satin SEDGMICK a C3 BCA GRANT | HASKE: PRATT wrt fexsey Comtuos } COSTRLA mirc : WOWA KINGMAN Twason | neosno coed Forte tome § . ICRAWFOROI BARTON a 08 ary PA ‘ommaeg aK ee ce RO Arey mS © emcee ‘ wonton ] sTEvens | sewaro | MADE 4 Cuan BARBER Sumner | come = | a » ° Cy ee Tras a 2 COMANCHE HARPER wip? one cast amt ns “6 Courax | toms cHauTAvUA| &*T [CHEROKEE ff JASPE rom oor cut aro won 2 se 5 lh 2 el 5 Ponce pa oe Jae “ runes ros a Tous os | ol arson ons om! m1 vi BEAVER HARPER REWTON \ s out " woo0s + a x] wacon ) “Towne . ALFALFA | GRANT MAY OSAGE B{NOWTAY CRAG | OTTAWA! DaRRY OREGON RIPLEY /ODDARD) < . STONEWALL | HASKELL | ?80c=mo=To=] YOUNG JACK wist DENTON DELTA LAFAY ATINA Be COOSA haciss aman, rv » Tor COLLIN HUNT ‘= | Titus = UNION CHICOT Jp wasnne- ae ame on CHILTON my aes ra BOT cues, (nid sad = ‘ z = 5 rom HOLMES Z CANES MOPRINS 1 a3 COLUMBIA Alcs — B_A- 7 fant o oe ~ 2 DAWSON | BORDEN] ScuRRY | Fi £ bo . s LJ ™ ry he! TEAKE fancon TS ous nor SHER | JONES SHACKEVORO] STEPHENS [PALO Pinto | PAPKER [TARRANT [OALLAS [rors an CAMP WEB-] CLAIBORNE | UNION MOREHOU if TUTAUGA awott he 78 Pact. S ‘ =F Fan amon STER Aa) D | eswoon| newer vost S 7] - FC os: RUDEROALES waco cs ee ee a MOREWS Pa rr A Bossies ay FS Staal BADISON Scart [NEWTON re wanenco | [DALLAS eoercouet » = russE gt mt thes apy TRADE MARK REG. U.S. PAT. OFF. =) oe | Hem CALLAHAN | EASTIAND—[eraTH M002 | JOHNSON [ELLIS nauFMan! ioe . “i ste MOST Hewsamcn} WHOS : LOWNDES tu a Sava sare 2 ~ a No Nee NO Se axe [ome Ef fm es Wl Ged ca ono = COUNTY OUTLINE =i, cuu or = PAROUA NexSOn pone Ly BERSON Loving ‘ WINKLER eed MIDLAND | GLASSCOCK | STERLING CORE —s Ss Pat NAVARRO HENDERSON De soro ) RED CALDWELL! TEXAS 4 cunt Lay et Loa (al one Ls | COLEMAN COMANCHE s ee wae cus biel 0) he BROWN CHEROKEE en WOMROE ak at . ems Fans wird CANE pen TERITOCHE 9 re) eS : yeney ae Caan = o, UPTON REAGAN Tom GREEN ra FREESTONE “s © SABINE Jw TINCOUN | Bsc 7] “4 Cohws | ow MILLS LTON ‘Met LIMESTONE. ” ere i ‘COMECUH ci \ E = a | ee ae eee Mss rece mecuLLocH HOUSTON * 5 SCAMBIA s . r « re . % X A S\ Ao wom \ See» me 8 re Scale of Miles ‘ < eet XMS agin WF vexnon roa sae aah a, ca ‘ a ROCKET | SCMEICHER ae pei ne . = . ORGE gy 'e son wate 0 100 200 \ “EFF bavis SAN SABA ed ATP TanTy Pou pAPiOEs: ae ATO Ot WASHINGTON gy STONE SON WADisOn / gang antl The a bese TANG cara | mien | EVANSE PEARL TACKSON TOR es : % ‘aeewsre TERRELL SUTTON nt Ai Gaimes | WALKER . we) O \remt, RvR . panwn % t WORALA ‘court! ‘cx! FOUR og \ : ( s El wos ie ofc MAP NO, 125 i sunt! aye, LI mance Aaa, Tee \ BURLESON tga dicta LCad faa Gor” rome (RU % . aon aa aa WALES sane OQ ber COPYRIGHT = 7 . WASHINGTON ("waiie| MONTGOMERY se Na waROn VaUSA AL Remaats, hays ‘BASTROP F\ia) x E IC \ hase camera verison /IBERIA ae ‘ lent . LA) BANDERA COMAL ow A M R A N : S K A > \ = CAL OweLL SC FaNETTE san Sire oat ORIGINATORS INNEY rt < er 2 SEN \ ry wee wroina Guaonure /eonzaes fiavaca one FORT BEND CLEARTYPE MA '° LEAR PS . waSoN Res TRADE ame PE GTTERED Of WITT WHARTON. BRATORIA PASCO NE\ YORK: nd pivencn | ravan oy an i SON NOTICE—This is a copyrighted map. The law . \e KARNES VICTORIA MATAGOROA 1 PASO BREVARD, prohibits the reproduction or copying of same, . FI ot any portion thereof, by any process for per KAUAI 1 Le! sonal use of resale, without permision. " Wi \ / HAW le oe Pee ee a . *% JUDICIAL DIVISION " / . Aa, . ot ve : / HONOLULU ee a KAUAI m7 UVE 0K mm ware 4 - / PART we cro . NUHAU a : u : ae / / 5 =a aa Cicer ' ) —_— - fi eo Stent -—ys, / a NUECES wos ! an? 0. *. t 2 Coe Cc / * A vt own! “ > . = : — x f vi % B- / e' \ LEBERG woo Sane H ° FIRST ON L.. s mata Pam woce TGaoaas 41) bil&tah do.}? izfgi", and is said to be used by the Navajos to stop postpartem hemorrhage. Seymour (1929) records the following fungi as infesting it: aecidium verbenae Auct, Amer., A. verbenicola Ell. & Kellerm., Dicaeoma verbenicola Arth., Puccinia sydowiana Diet., P. verbenicols Arth., Erysiphe cichoracearum DC., and Septoria verbenae Robe It was introduced into cultivation in 1801 or earlier, Bailey (1935) lists Cronamere as offering it to the horticultur- . Publ. Bot. 2: 198 & 27, (1951) is probably a misprint for Clokey 8,73 oe eta MacDougal (1907) suggests that what he calls V- bracteosa Michx. (now more correctly known as V. bracteata Lag. & Rodre) may hybridize in the wild with V. canadensis (L.) Britton, with V. hastata L., and with what he calls V. angustifolia Michx. (now more correctly known as V. simplex - Lehm.). However, he cites no specimens, gives no descriptions of the supposed hybrids, and does not say that he actually ever saw any such hybrid plants. I he only hyb which I recognize are xV. bingenensis Moldenke [V. bracteata Lage & Rodr. x V. lasiostachys var. septentrionalis Moldenke], V+ deamii Moldenke [V. bracteata Lag. & Rodr. x Ve stricta vent.], and xV. perriana Moldenke [V. bracteata Lag. & Rodr. x Ve urtici- folia L.J. e 1962 Moldenke, Monograph of Verbena 281 A. H. Smith (1868) records V. bracteata as "rare" at vet Point, Philadelphia, Pennsylvania. Jones (191) records it from Ross Coun ounty, Ohio, on the basis of a Bartley & Po aitsiie collection. Lowe (193) says it a occasional on eB eas and waste ground at Elgin, Lena, Lyleton, and Ninga, Manitoba -- I have not yet seen material from the first two of these localities. Jepson (193) rae it a Kern County, California; Braun (193) from Mc Cre serge d Harper (192) from Scioto County, Ohio; peel Sr ti en County, New Jersey; Monachino (1960) from Bronx Coun i car x Taek; Porter, Wo » Pride, & Howard Ty, (190) from Worcester Co ounty, Massachusetts; Deam, Yu it is "Native from Ohio westward and southward; locally introduc- ed in the northeastern states and on the Pacific coast." Groh (19hh) says that in Canada it is an annual, found in sod and waste places in dry land. He cites the following material not as yet seen by me: ONTARIO: Lambton Co.: Groh s.n. [Pt rd, 192k]. Middlesex Co.: Groh s.n. [Strat hroy, L9oh]}. Westmeath"; Grylls none [1935]. MANITOBA: Brandon Co.: Groh s.n. [Wawanesa, 1932]. "Long River Valley": Burman s Sn. SASKATCHEWAN: Maple Creek Co.: Campbell 95. Regina Co.: “Carmichael 89. Swift Current Co.: Groh sen. (Cabri, 1933]. "Forget": Russell & & Ledingham 135. ALBERTA: Lethbridge Co.: Fyles s.n. (Lethbridge, Biesen'E roh Ss. Se n. (Lethbridge, 1929]. North Battleford Co.: Carmichael 7. "Cavendish": Campbell 4. BRITISH COLUMBIA: Cariboo oat “Fletcher Sn. [Lake Kamloops, May 29, 1885]. Yale Co.: Eastham s-ne 1s [0S0— yoos, 1938]; Groh 243. He notes that the Point “Edward "infesta- tionn dates rem to 0 1889, at least, since the species was there by Dearness, according to Macoun's Catalogue. David Macoun also collected rte "near the forks of the Red Deer and Saskatche- - 8, 1879." Douglas H. Kent, in a letter to me dated mevetet 2 28, 199, Says that the species is adventive in England, Thur n'& igurs (1922) cite a Harvey collection from Bodmin Road rake i Corn- wall, collected in 1918, Gay (38h9) says "En la quinta noticia de las plantas raras del jard ot4nico de Ginebra, A. de Can- dolle indica esta especie como y ire igualmente de Chile, lo qe es muy dudoso." I have not as yet seen any material of it from rom Chile, wild or cultivated. Perry (1933) cites the following 320 additional specimens not #8 yet seen by me: ONTARIO: ae on Co.: Macoun s.n. [Point Ed- » Lake Huron, 19 July 1901] (N). BRITISH COLUMBIA: Cariboo Cou: Macoun s.n. [Kamloops, apie 13th 1889] (W). MAINE: Cumber- 282 P8I:7:0 £.0-.6-1-4 Vol. 8, no. 6 land Co.: Chamberlain 118 (G). MASSACHUSETTS: peohig Co.: Rich sen. [Dedham, Aug. 1900] (G). Worcester Co.: Dewhurst s.n. (Worcester, 27 June 190] (G). CONNECTICUT: oueteid Co.: Eanes 8137 (G). Litchfield Co.: Fernald s.n. [Winsted, Winchester, YES Sept. 1909] (G). NEW YORK: “Suffolk Co.: Von Schrenk s.ne [Mon- tauk Point, 23 July 1895] (E). NEW JERSEY: Y: Canden Co.: Parker s. ne [Kaign's Point, Camden, 20 July 1866] (BE, G). Hudson Co.: . Cos: Van Sickle sen. (Weehawken, Aug. 189),] (W). PENNSYLVANIA: Lancaster Coot Ae Aw A. Heller s.n. [Lancaster, 29 Aug. 1900] (W). Lehigh Co. Churchill s.n. [Cementon, 20 Aug. 1923] (G). NORTH CAROLINA: Ire- dell Co.: Ruger s.n. (Statesville, June 1872] (W). GHORGIA: Bart- ow Co.: Ravenel s.n. [Cartersville] (N). DeKalb ERs Eggert S.ne [Stone Mountain, 2)) May 1897] (£). Gwinnet Co.: H. Allard | 207 (E, W). Lee Co.: J. D. Smith s.n, (Smithville, bin rend 1885] (E)- : Franklin C Co. Biltmore Herb. 1082b (W). County undeterm- ined: Buckley SN. (E); Chapman s.n. Sen. (EB). ALABAMA: DeKalb Co.: Ruth 518 (g). Etowah Co.: Eggert s.n. (near Atalla, 9 July 1 898] (E, N). Jackson Co.: Ward sen. [Stevenson, 5 Sept. 1877] (W). Lee Co.: Earle & Baker s.n. [Auburn, 13 May 1898] (W). Mobile Co.: Mohr s.n. [Mobile, 22 May 188] (W). County undetermined: Rugel sen. [Sept. 1843] (E). OHIO: Erie Co.: Moseley s.n. [Sandusky, sky, Us Aug. 1920] (G). ILLINOIS: Kane Co.: Sherff 1797 (E). Mason Co.t June 1886] (G). Winnebago Co.: M. S. Bebb s.n. [Fountaindale, 1887] (G). INDIANA: La Porte Co.: Mell & Knopf s.n. [Michigan a 6 aes 1903] (E). Steuben Co.: C. C. Deam 1272 (E, W)- OWA: Black Hawk Co.: Burk Sl2a (E). Chickasaw Co.: Rolfs SMe tlarier, 1890] (G). Fayette Co.: Fink s.n. [30 July 1894) (W)- ottawat Co.: Vasey sen. {Council Bluffs] (G). Story Co-? ball Some Laat. 5 Aug. 1896] (E.G. k, ws H. Se Fawcett ie (E, G, W : Jefferson Co.: Be sn. 1, [southern ‘thern Lowis- vinle, ia Aug. 1892] (E). County si rmntipoet Rafinesque SNe» (G). VESSEE: Cheatham Co.: Eggert s.n. [Kingston oe 3, 19 Aug. oe (E). Franklin Co.: Eggert s.n. [Sherwood, 1897] (E). Shelby Co.: Fendler s.n. (preter 22 June ? B61} (G)- MICHIGAN: Cheboygan Co.: J. He J. He Ehlers 627 (G). Manistee Co.? Morong s.n. [Manistee, 3 Aug. 1882] Se WISCONSIN: Dane C0.: Chure Sen. [Madison, 27 Aug. 1893] (G qe Sauk Co.? Eggert o-e Eh (near Mirror Lake, Be ott 903] (E). County undeterm Schuette s.n. [Fort d, 22 July 1887] (, W). SErMITRSOTA! Hen- nepin Co.: Mearns 5 al (W)3' Sctmette Schuette s.n. [hills between li and Fort Snelling, 5 July 1588] (G). Mille Lacs oa BE. P. Shel- don s.n. (Princeton, July 1892] (G, W). Stearns Co-: Chandonnet 1962 Moldenke, Monograph of Verbena 283 sen. [Collegeville, 29 July 1912] (E). Wabasha Co.: Manning s.n. [Lake City, 25 July 1883] (G). NORTH DAKOTA: Benson Co.: Lunell S8.n. [Leeds, 10 Sept. 1896] (G, W). Morton Co.: Sarvis 122 2 (W). Richland Co.: Bergman 2370 (E). Stark Co.: Holgate s.n. ~[Dickin- son, 1), Aug. 1908] (G). SOUTH DAKOTA: Lawrence Co.: Carr 179 (E, G, W); Rydberg 93) (W). Meade Co.: Forwood 298 (W). Walworth Co.: wishut 3359 (E (B). KANSAS: Grant Co.: Ce He “C. H. Thompson 42 (E, W). Hamilton Co.: Rydberg & Imler 1008 (E). Kearney Co.: A. S. Hitch- cock 1129 (E). Morton Co.: Rose & e@ & Fitch 17106 (W). Riley Coe: J. B.S S. Norton 392 (E, G, W). MISSOURI: Butler ,Co.: Kellogg 15276 (EB). Greene Co.: P. C. Standley 9362 (W). Jackson Coe: B. F. Bush 2109 (BE), 4010 (E, G), 8537 (E). Jasper Co.: E. J. Palmer ish (); W. Trelease 9), (E). Jefferson Co.: F. Wislizems 26! 284 (E (E). Marion Co.: Je Davis 2691 (E), 3587 (E), 9039 (E). Pike Co.: de Davis 2h1) (E). Polk Co.: W. Trelease 715 (E) (E). Taney Co.: B. F. Bush Bush 601 (E). Wayne Co.: E. J. Palmer 6106 (E). Saint Louis: Eg- gert S.n. [Jefferson Ave., St. Louis, 2 July 1875] (E); Engel-_ mann sen. [St. L Louis, Sept. 18)1] (c, W); Riehl 9 (E). ARKANSAS: Craighead Coe: Tenixee 3353 (E), 3371 (E, G G). Hempstead Co.: Be F. Bush 103) (E). Independence Co.: W. Trelease s.n. [Batesville, I sept. 1897] (E). Sebastian Co.: Bigelow s.n. [Fort Smith, 1853- 54] (W). Washington Co.: F. L. Harvey vey 62 (E). LOUISIANA: Rapides Pare: C. Re Ball 553 (E, G, W). MONTANA: Dawson Co.: J. Ball s.n. (near Glendive, 19--21 Aug. 188k] (G); Ward sen. (Glendive, 17 July 1883] (W). Sanders Co.: Sandberg, MacDougal, & Heller 972 (G, W). IDAW: Ada Co.: Jo A hs Clark & ( (B, G, W), 279 (E, G, W)s Mulford s.n. [Boise, July 1892] (E, G). Canyon Co.: J. F. Mac- bride 200 (E (E, W). Fremont Co.: Merrill & Wilcox 776 (G, W). Nez Perce Coe: Sandberg, MacDougal, & Heller 264 (E, G, W). County undetermined: L. F. Henderson 070 (W). WYOMING: Albany Co.: A. Nelson 7671 (E, G, W). Natrona Go.: Goodding 207 (E, G, W). Yel- lowstone National Park: Mearns 2698 (W), 5009 (W). County unde- termined: A. Nelson 320 (E, G, W). UTAH: Beaver Co.: Goodding 1022 (E). Emery Co.: “Me E. Jones 5478 (E, W). Piute Co.: Rydberg & Carlton 7043 (G, W). Salt Lake Co.: Pammel & Blackwood 3589 (E, G); Ss. Watoon 823 (G, W). Sanpete Co.: Ward 256 (G, W). NEV- ADA: Clark Co.: Goodding” 2312 (E, G); S. B. parish 10288 (E, G). COLORADO: Archuiiste Bon Co.: C. F. Baker 56h (E, F, G). Denver Co.: Eastwood 25 (E, G, W). El Paso Co.: Co.: Clements 9 (E, G, W); Drush- el 4887 (E). Gunnison Co.: C. F. Baker 920 (E, G, W). Larimer — Co.: G. C. S. Sheldon 38 (G). Montrose Co.: + F. Baker 288 (E, G). Otero Go.: Rose & Fitch 17503 (E, W). NEBRASKA: Cedar Co.: Clem- ents 2612 (G, W). Kimball Co.: A. A. Heller 14303 (E). Lancaster Cow: Webber sen. [Lincoln, 20 June 1988) (W). Cow a undetermined: 28h PHYTOLOGIA Vol. 8, no. 6 Hayden 10 [banks of Yellowstone River] (E). OKLAHOMA: Comanche Cos: M. K. Clemens 11750a (E). Creek Co.: Bush 36 (E). Kiowa Co.? G. W. Stevens 1192 (G,N). Leflore Co.: Blakeley 162 (G). Texas Coes G. W. Stevens 391 (E). Woods Co.: G. W. Stevens 680 1/2 (G), 2889 (G). TEXAS: Dallas Co.: Reverchon 736 (E), 2116 (E). Bl Paso Cos: C. Wright ok (G, W), 567 (G), 1499 (E, G, N). Howard Co.: Havard s.n. [west of Big Spring, Sept. 1881] (W). Lubbock Co.: E. J. Palmer 12)96a (BE). Randall Co.: Eggert s.n. {banks of Red Riv- er, 13 Aug. 1900] (E); E. J. Palmer 13861 (E). Tarrant Co.: Ruth 109 (E, G, W). Taylor Co.: S. M. Co.: Parry, Bigelow, Wright, & Schott s.n. [below Dofiana] (W). NEW MEXICO: Bernalillo Co.: Arséne & Benedict 16600 (E); C. Ce Ellis 221 (E, W); E. J. Palmer 31117 (E). Chaves Co.: Earle 352 (E, W). Colfax Co.: Griffiths 5560 (E); P. C. Standley 6268 (W)-. Dona Ana Co.: Wooton 09 (E, G, W); Wooton & Standley 3330 (E, F). Eddy Co.: Wooton s.n, [Queen, 2 Aug. 1909] (E, W). Grant Co.? 0. B. Metcalfe 137 (E, G, W). Sandoval Co.: Arséne & Benedict 16790 (D). San Juan Co.: Wooton 2831 (W). San Miguel Co.: Arséne 18399 (D), 18407 (E); P. C. Standley 5136 (E, W). Santa Fe.: AP sine & Benedict 1573) (E)3 P. C. Standley 58 (E, W), Lobe (B, % W)» Sierra Co.: 0. Be Metcalfe 897 (E), 1008 (E, G, W). Socorro Co.t H. H. Rusby 3% (E). County undetermined: Fendler 587 (D, By G, W), 592 (E). ARIZONA: Apache Co.: Goodman & Hitchcock i317 (E). Coconino Co.: Clute 93 (E, G, W)5 H. C. Hanson A147 (E)5 MacDougal 286 (D, G, W). Yavapai Co.: Peebles, Harrison, & Kear ney 4243 (W). WASHINGTON: Benton Co.: Cotton 621 (BE, G, W Douglas Co.: Sandberg & Leiberg 340 (E, G, W). Kittitas Co? Cot- ton 1363 (G, W). Stevens Co.: Kreager 47h (G, W); Lyall SoDe (Cascades Mts. to Ft. Colville, 1860] (G). Whitman Co.: Elmer 32h (B). Yakima Co.: Vasey 68 (G, W). OREGON: Clackamas Co.: Ee Fe Tro" Thompson 880 (E, G; W)- Umatilla Co.: H. E. Brom 58 (E, W); Griffiths & Hunter 18 (W)5 Leiberg 902 (E, G, N, W)- Wallowa Co.: 5. P. Sheldon 8202 (5). Wasco Co.: Abrams 91,87 (E); Engelmann s.n. [The Dalles, 2 Aug: 1880] (E). County undetermined: W. C. Cusick 1967 (E, G)5 Be Hall 393 (E, G). CALIFORNIA: Humboldt Co.: Abrams S.Me Longe 2 Lake, 1 April 1903] (E, N). Inyo Co.: Coville & Funston 953 Los Angeles Co.: Abrams 257) (E, G). Monterey Go.: Bolander 4<0 (G); Vasey 512 (W). Orange Co.: D. L. Crawford s.ne [Laguna Can- yon, 26 July 1916] (£). San Bernardino Co.: Parish & Parish 2171 (W). Tulare Co.: Palmer 2699 (W). MEXICO: Baja california: C. Re Orcutt 1228 (E), 1546 (E), 1549 (E), sen. [Tia Juana] (F, W). Chihuahua: Gregg 433 (E); Stearns 7 (F, W). Coahuila: Greee 636 (E); Edw. Palmer 1040 (D, G, W)5 Purpus 52h, in , (w) . Sonora: Kennedy 7032 (W). CULTIVATED: Spain: Herb. Bote 1962 Moldenke, Monograph of Verbena 285 Gard. Madrid s.n. (E--photo of type). In aged I have examined 2293 herbarium A ge i of this spec~ ies, including the type on pe of most of the names rt, jou ions: CANADA: ONTARIO: Lambto: roan ta C. K. Dodge s.n. (Point Edward, 7-15-96] (Fe, ast sen. [Pt. Edward, 7/8/ 97) ( (Au, a Mi, Ob--874)5), sen. [Point Edward, Aug. 12, 191h] (Mi, Mi); -L. Fisher s.n. (Sarnia, July 10th, 1902] (S); Gaiser 230) PE ose Gp, Mm), 2517 PE (Gp, Gp, Mm); RS. & Montgomery 2237 PE (Gp, Gp, Mm); J. Macoun 5635 (N). Middlesex Co.: Dearness & Col- grove s.n. [northeast of Strathroy, 3lst August 193] (Mg). County undetermined: We Dearness 84 (Mg). MANITOBA: Crystal City: G. Thomas s.n. [Crystal aed Le ies Secures oe are 12135 (N)5 ioace is Campbell 95 (1in—38075) . ALBERTA: Lec Cos: Macoun . 21268 8 (N). Macleod Co.: E. H. Moss 20) (N). Medicine Hat Dist.: Boivin & Alex 9177 (N). North Vattleford Co.: Breit- ung 15643 (N, N). County undetermined: Degener & Degener 25650 [Frank] (N, W). BRITISH COLUMBIA: Cariboo Co.: Macoun son. [Kam loops, dune nie 1889] (C). Yale Co.: Macoun 76775 (N)j Je Le Thompson 11909 (cm), 1437) (Ba, Ca—882591, 591, Gg——31086h, It, Se--201)2, Se—577h9)» UNITED STATES OF AMERICA: VERMONT — tenden Co.: Muenscher, Manning, & Maguire 9 (Vt). Windsor Co.: de P. Poole Sen. [West Hartford, ~8/12/19h6) ( (Dt). County undeter- mined; Herb. Carpenter s.n. (vt). CONNECTICUT: Fairfield Co.: Eames 8137 (( (N)5 Jansson s.n. 2s pat June 23, 1928] sey Peck s.n. [West Troy] (Al); 3 Puissant s.n. [West Troy, Aug. 30, 1870} (Al (Al, Br). Saratoga Co.: H. D. : H. D. House 2911 (Al, Au, Au, B Ca—882600, N, Ok, Ur); 0 . Ps Phelps son. [July 2h, 1925] (Al). Suffolk Co.: E. Je Alexander son. {Nontauk Point, Aug. 20, 122 (Bt—35372, Dp, H--h0668, I, Io—1h5160, Ka—90369, N, Ok, Um—251, Up, ur), s 8.n,. sy om *1936] (N). NEW JERSEY : aids Co.: Van Sickle s.n. [Weehawken, July 20, 189] (Tc). Sussex es +t Abies SUB ( (tr); Clausen & Edwards 3538 aes PENNSYLVAN- Phere, oma Fogg 9870 (Up); C. F. Parker son. [July 28, 77) (Pr), Sen. [July 25/76) ] (Pr). Wyoming Co.: Glowenke 11298 286 PHYTOLOGIA Vol. 8, no. 6 (Ur). DISTRICT OF COLUMBIA: Mrs. L. Me Jenkins s.n. (Washington, — ee July 1911] (Up--58797). VIRGINIA: Accomac Co.: He A. Gleason 857 1012 (C, Mm—1539, Up—17088). WEST VIRGINIA: Hampshire Co.: BE. L. Core 4391 (We); Frye 1198 (We). Monongalia Co.: Davis & Davis NORTH CAROLINA: Clay Co.: Huger 18 (N). Forsyth Co.: P. 0. Schal~ lert 11832 (No—23996). Iredell Coe: M. E. Hyams sn. (Du-—2))161). McDowell Co.: LeRoy & Ruger sn. [Foot Black, June 1872] (N). GE- ORGIA: Bibb Co.: J. K. Small s.n. [July 8-9, 1895] (C). Catoosa Co.: J. K. Small s.n. [August 6-12, 1895] (C). Charlton Co.: E. M. Hall 511 (N). Clarke Co.: K. M. Drewry E.1988 (Up); Mrse Ke Me Drewry s.n. [Aug. 25, 1929] (Gu-—E£.1988); W.H. Duncan Tis (Gu) DeKalb Co.: J. K. Small s.n. [Stone Mountain, July 1--6, 1895] (C, Io—-20750, S). Fulton Co.: T. P. Cleveland 5.ne [Atlanta, 1860] (F1--21086). Gwinnett Co.: H. A. Allard 207 (N). Hancock C o.: Cuthbert s.n. (Mayfield, May 1903] (Fl--21087). Jefferson Co.: He M. Hopkins sen. (N). Muscogee Co.? Boykin s.ne {Columbus (C); Saurman s.n. [Columbus] (Pr, Up——17098). Screven Co.s Jo Me Reade s.n. [May 18, 1923] (Gu--E.8353). FLORIDA: Franklin Co.? Chapman s.n. [Biltmore Herb. 1082b] (N), S.M. [Apalachicola] (C)- County undetermined: Buckley s.n. (Io--153h2); Chapman 5 De (Fl rida] (C). ALABAMA: DeKalb Co.: Ruth 95 (N). Etowah Co.? Eggert s.n. (8 July 1898] (N); Vasey son. (Gadsden, 1878] (Pr, Vt). Lee Cost C. F. Baker 914 (C); Earle & Baker Sn {Auburn, 10 May 1896] (Cm), sen. (Auburn, 5/13/1898] (Cm, Mi, Po--63897, UF, Vt)5 Earle & Underwood s.n. [Auburn, 20 May 1896] (C). Tuscaloosa Co? llard & Maxon 333 (Gu--E.7022, N). Wilcox Co.: Buckley S$.m- [May 180] (T). County undetermined: Rugel s.n. (Alabamam mediam, Sept. 1843] (Br, Ca—-80)78, Go, M). OHIO: Butler Co.: Demaret 27351 (N, Qu); J. Ferguson s.n. [July 6, 1932] (Ob--80 91). Ham Zlton Co.: Aiken 10676 (Ll); E. L. Braun s.n. [6/12/06] (Cn, cn); Ce G. Lloyd 1962 (Mi), sen. [July 9, 1861) (Al), SMe [July 12, 1883] (Go). Lake Co.: H. CG. Beardslee s.n. [June 7th, 1871] (0b— 80592). Richland Co.: Moseley sen. (0/2h/2h] (Ob--97292) « ILLIN- OIS: Adams Co.:Evers, Jones, & Jones 1207 (T1—2561L, Ur); A- Be Seymour s.n. [Camp Point, Aug. h, 1877] (H--106598), S-Ms (Ur— 1175). Alexander Co.: H. M. Franklin 62 (Ur); Winterringe® 3137 (I1--28358). Bureau Co.: Winterringer 7996 (1L—372bh) « carroll perme Co.: Ahles 322 (Ur); Winterringer 2770 (1128112). Champaign Co.: Ahles 6299 (Ur, Ur), 6300 (Ur, Ur); M. H. Bell 131 (ur); Ge N. Jones 11815 (Ur), 12457 (IT1—158h5, N, Ur), 1665 (Ur)s stubee 87 (We); Winterringer 022 (I1--29322). Cook Co.: He ie Babcock 925 (Ba—-237k0), Sen. [Hyde Park, June 19, 1871] (N), S-0e [gute 1962 Moldenke, Monograph of Verbena 287 18, 1874] (Sg--16115); L. H. Bailey s.n. (Chicago, July 27, 1880] (B a); H. R. Bennett s.n. [July 6, 1957] ne M. A. Chase 1570 (Ur, Ur); 6 Collector undesignated s.n. (Cn); .D. Ful Fuller 165 (T1--15852), 11205 (I1—19h17); Lansing 2616 (( ie “2616 bis ( rr), H. N. Patterson s.n. (Bz—237h9). DeKalb Co.: Ge. N. Jon Jones s 17260 (Ur). DuPage Co. W. S. Moffatt 383 (Ur), s.n. (Warrenville, July 11/93] (Ur); Moffatt & Moffatt 318 (Ur). Fayette Co.: A. Hoffman 191 (Dt). Hancock Co.: Mead 18h2 (' (Tl), son. [Augusta, 1842] (Vt). Henderson Co.: H. N. Patterson s.n. [July 1877] (Cm). Iroquois Co.: Winterringer 6462 (I1—3375y). Jackson Co.: G. D. Fuller 319 (Ur); Fuller & Welch 319 (T1--158)9). Jo Daviess nite ‘i hes 4366 (Ur). Kane — Sherff 1 1797 (Ur). Kankakee Co.: E. Hill 71. 1871 (Ur). Lake Co.: R Ra Kas Brom s.n. [July 9, iat (Ur); ae 5812 sokasikiaiy 6562 (N). LaSalle Co.: Kofoid s.n. [Aug. 1889] (ob—50780 0). Lawrence Co.: Sivert s.n. [July 25, 1952] a. Lee Co.: Keithley 113 (I1--158)), son. [June 23, 1943] (Ur). Liv- ingston Co.: G. N. Jones 18955 (Ur). Macon Co.: R. G S.n. (July 3, lok] (Ur); Weigel s.n. [July 2h, 1950] (Ur). Mason Cos: Ables 2010 (Ur), 4068 (Ur); Burrill & Clinton s.n, (Havana, June 22, 189k] (Ur); He. A. Gleason 9183 a, son. (Havana, Aug. 18, 190h] (Io—176180) . McDonough Co.: G. N. Jon Jones 22408 (Ur). olenty Cow: Ve H. Chase 952) (Ur); Nason s.n. , [Algonquin, August 2, 1878] (ur), sen, [Aug. 23, 1912] ore Vasey 30013 (Ur), Sen, [Ringwood] (Pa, Up--17090). McLean Co.: - De . D. Fuller 173 (1581.7), Menard Co.: E, Hall s.n. et "jaly 1863] (Pa). Monroe Co,: Winterringer 3921 (11--29051), 3955 (I1—29021). Mor- gan Co.: Winterringer 52 5252 (T1--35h16). Ogle Co.: A. L. Hills 23-0 (T1—-158)3); Waite s.n. (Ur--1176). Peoria Co.: V. He Chase 9707 (Ur, Ur), 1heL8 (Ur (Ur); M. Fisher s.n. [6-13- ae eae [oF Heading s.n. [187 5) (Ur); F. E. McDonald s.n. [Peori June 1887) (Ur), son. [Peoria, July 1890] (Ca-—67601), Son. " [pe- oria, June 1903] (N), sen. [Peoria, Jul. 1903] (og 258), Behe [Peo eoria, Aug. 1913] (La, Um—2h0, S). Randolph Co.: V. A. son 15196 (I1—31496). Richland Co.: Scherer 316 (I1--26357, Ur). Rock Island Co.: G. D. Fuller 13805 (T1--3134h); H. N. Moldenke 1656 (N). Saint Clair Go.: Eggert sen. [East St. Louis, June 11, 1879] (Au, Ca—181583, Cm, Io—79813, Mn—688h, Po-~161455, Up— j Je Neill 31 (I1—-23391). Sangamon Co.: G. D. Fuller 5086 (15816), S10¢° (I1--15848), 8795 (Il--16015, Ur). Shelby Co.: Batson s.n. (N); | Ge. D. Fuller 13123 (I1~23793). Stark Co.: V. He Chase _ SN. en. [July II, 1896] (Ur), ae - arwess oe re (Ur). Stephenson Co.: c. Wi Solute sen. [F » 1898] (N). Tazewell Co.: V ,H. Chase 1112 1/2 ck ie py! 0, 13539 (Au~123201); Cin Churehiili Sen. [E. Peoria, June 9, 1912] (Ba). Ver- 288 PHYTOLOGTIA Vol. 8, no. 6 million Co.: W. S. Moffatt s.n. (Danville, Aug. 3, '39] (Ur). Wa- bash Co.: Schneck s.n. [June 15, 1879] (Ur), sen. [July 8, 1880] (Ur), sen. (Aug. 6, 1888] (Ur); Shearer 191 (Dt), sone [June 10, 1900] (La), sen. [Mount Carmel, July 4, 1900] (Ur, Vi). Whiteside Co.: Ahles 426) (Ur). Winnebago Co.: M. S. Bebb son. [Fountain- dale, 1571] (Dt, Pa); R. Bebb s.n. [Spring 1579] (Ok); Fell & Fell F.46358 (Il—2123h); Fuller & Haime 28f (I1--15851), 2528h (Il— 15850). Woodford Co.: V. H. Chase 9697 (Ur). County undetermined Collector undesignated s.n. (Gg--31297); Marie-Victorin 6914 (Vi); Seymour s.n. [1881] (Sd—251); Underwood s.n. [Auna, June 1882] (Cp); Vasey 6 (I1—16005), s.n. (Pr). INDIANA: Bartholomew Co.s C. Ce Deam 36915 (Dm); Friesner 19217 (Au). Cass Co.: Ek 5.Me [near Logansport, 8-25-19)2] (Bt—64072). Clark Co.: J. F. Baird sen, [Charlestown, 5-30-77] (Mi). Crawford Co.: Co C. Deam See [July 12, 1899] (Dm). Daviess Co.: H. J. Clements S.ne (7/31/95) (Pu); C. C. Deam 25582 (Dm). Elkhart Co. C. C. Deam 34430 (Dm), 66163 (No—-19967); Ek sen. (July 27, 1942] (Dp--33216), See [Go shen, 9-1-19h2] (Bt—63357); H. N. Moldenke 7559 (N)~ Fountain 3923 (Dm); Friesner 13937 (Bt—h7968, Fn--1656). Gibson Co.: Ce C. Deam 4.3828 (Dm). Grant Co.: Ek sn, [8-15-1938] (Bt—56919)- Hendricks Co.: G. W. Wilson s.n. [Danville, ] ard Co.: Ek san, [6-16-1939] (Bt—l9325). Jasper Co.: C. C. Deam 12087 (Dm); Friesner 11593 (Bt--$0734); W. H. Welch 12h (Ur), 20 (Dp), sn. (In--6176). Jefferson Co.: C. CG. Deam 13409 (Pu); Ae H. Young s.n. (Hanover, 1875] (In—10225, Pr). Knox Coe: Co Co Deam 54330 (Dm); Lipps s.n. [August 28, 191] (Ok); S- weCoy 1906 ( 669), sen. [near Bruceville, 6-3-3] (Bt--18),28) « Kosciusko Cos: C. C. Deam 5132 (Dm); Friesner 1929 (N, Ob—201452, Ok, St). Lagrange Co.: C. C. Deam 14934 (Dm); Friesner 15977 (Bt—5LOLL, N); Weatherwax s.n. Tontario] (In--3666). Lake Co.: M. Ae Chase 100 (Ur); Umbach s.n. [Lake Station, June 6, 1896] (N, N)» Laporte ines OSs Deam 53971 (Dm); W. S. Moffatt 1685 (se~95718, usrg c. P. th s.n. [July 2h, 190] (Mi). Lawrence Co. Kriebe 3098 (H--56280); Wible 72 (In—-H58h). Madison Co.: GC. Re Barnes Sens (Madison, 6/18/77] (Pu). Miami Co.: Ek sen. ([6-Lb-19h2] (Bt—-63700, Pl--132302, We). Monroe Co.: F. C. Anderson S-Re [Bloomington] (In—-1729); Lewis & Brown s.n. [East Pike, dee 106] (Au). Momtgomery Co.: We Rhoades 971 (Bt--14479, Dp-— )e Newton Co.: C. C. Deam 50589 (Dm). Noble Co.: C. C. Dea 1690 (Dm); Van Gorder 139 (Pu). Perry Co.: C. C. Deam 25057 (Dm). Po~ sey Co? Ce C- Deam 43500 (Dm); W. H. Welch 6899 (B1—li2290, 00" 882597, Dp--1h065, Ky, We). Pulaski Co.: C. C. Deam 29861 (Do/} Friesner 1,73 (Bt--50777). Putnam Co.: Friesner 17832 (Bt--6 750, 1962 Moldenke, Monograph of Verbena 289 Mi); Grimes 671 iy Yuncker 10180 (Ca—7097, Dp—1h53). Starke Co.: S. M oy 4199 (Dp--13912); W. H. Rhoades s.n. [Kno July 1932; vaidsnike sake 1707] se Hs, N, Ray s.n, [near Knox] (Hs). Steuben Co.: C. C. Deam 1272 (Dm, Mi, N)3 Friesner 6071 (Bt-- 13377, Bt—13380, | ae ce -SLOKShS, 15717 (Bt--55228, Ca-- 882598). Tippecanoe Co.: H. J. Clements s.n. [6/17/96] (Pu); C. C. Deam 17735 (Dm), sen. [June 23, 1901] (Dm); Dorner 205 (Pu); Ek s.n. [June 10, 192] (Bt—63512, Dp—32006); Friesner er 22856. (S8)} He Herb. . Purdue Univ. s.n. [7/29/91] (Pu). Vermillion Co.: C. Cc. Deam 11 Ti910 (Da (Dm). Warren Co.: Umbach sen, [Pine, June 29, 1898] (Ka—62598). White Co.: M. McKee ihm (Dp). County undeter- mined: Grimes 2160 (Dp). IOWA: Allamakee Co.: A. Hayden 5019 (Io- 14880). Benton Co.: Folck s.n. [Sept. 15, 1932] (10139569). Black Hawk Co.: Burk 5]2a (Ur). Boone Co.: Pammel, Buchanan, & King 3882 (Io—31762). Buchanan Co.: Bode s.n. [Lamont, Aug. 13, 1919] (Io—-97035). Carroll Co.: L. H. Pammel s.n. (Carrollton, Jul. 21, 1919] Saag: Cerro Gordo Co.: L. ES Pammel s.n, [Wheelerwood, Sept. 20, 23, 1902] (Io--82h2h). Chickasaw Co.:— Rolfs s.n. [Lawless, 18637" (Io--15318); Spiker s.n. [1925] (Io-- 118281). Dickinson Co.: Conard s.n. [July h, 1 aes (Se--19992) ; te 2 Be Fox 2 (We). Dubuque Co.: Mertz s.n. thug. , 2871] .. [Dubuque, Aug. 15, 1877] (Vt). Emmet Co.: rein au coral 165173), s.n. [July 16, 1930] (Go). Fayette Co.: stte, Aug. 1893] (Io—28697), sn. [Fayette, Ee 30, 1 sar ras 2682). Greene Co.: J. B, Hatcher s.n. [Cooper] (Ca—25179). Guthrie Co.: L. H. Pammel 562 (Io—118733). Hamilton Co.: Pammel & Zimmerman 300 (To--118096). Hardin Co.: L. H. Pammel s.n. [Steamboat Rocks, 9-12-12] (Io—-52668). Henry Cons ot Je He Mills Sen. [Mt. ent rat 1897] SNotige 5+ Iowa Co.: Shimek s.n. [July ri ] (Ur). Johnson : Anderson & Harrington s.n [May 30, 1932] (Bt—58018) ; "pape % Fitapetrick s.n. [6- 17-1901} (Dm); A. S. ee ee City, 6/8/89] (Io— ° Co.: Fults 116 (Io—-138969). Linn Co.: Treat s.n. Brae: June iss Lah Mahaska Co.: Augustine 397 (Ok, Ok). arshall Co. ammel s.n. (Marshalltowm, litown, Jul. 13, 1929] ‘or (To--13),205, ‘etre 38), Muscatine Co.: Shimek s.n. [June 28, 1913] (Ur), s.n. (sept. 2, 1922] (N). Pottawattamie Co.: Demaree 23679 (Sm). Story Co.: arthur s.n, [Ames, June 12, 1871] (Ah, Ah), s.n. (Ames, July 17, 1877] (Ah, Ah, Gm); C. R. Ball 86 (Fe, Fl—-21088, F1--21090, Io—15316, N); C. E. Bessey eae. Cratty son. [Ames, Jul. 29, 1925] (Io—117331); J. B. Elder 365 (St); H. S. Fawcett 12 (F1—-21090, To—76853, N); A- Hayden 30 (Au, To—-1h2h37, N), s sen. [October 15, 193] (N, N); Herb. ‘Towa 290 PHYTOLOGIA Vol. 8, no. 6 state Coll. 7654 (Io); A. S. Hitchcock s.n. Ure, June 18382] (Io—-15312), s.n. [Ames, 1889] (Io--15317); L. H. Pammel s.n. fames, 1909] (Io--7663), sen. [Ames] (10--20752, 1057705); We ae Neber 43 (Bl—l8182, Io--152969), 723 (B1--18181) . Ue apd Coe: sen. [Creston] (Io—-22908) . Webster Co.: Blumer is (Io--10769h); Paige s.n. [Fort Dodge] (Io—-13101); Somes Pisin 8. ne n. [7/27/05] (B1—2295). Winneshiek Co.: Goddard s.n. (Decorah, 6-1-95] (Io--72021); Holway s.n. (Decorah, Aug. 79) 92225), sen. (Decorah, June 1}, 1881] (I0--38751); 3 Tolstead s.n. {Decorah, July 10, 1933] (Ca--88259, Ew), s.n. [Aug. h, 1933) (Io--14)285) . Bint undetermined: Suksdorf s.n. [July 26, 1873] (Pl1—138h29). KENTUCKY: Anderson Co.: Wildburger 1962 (Ka) « Hick- man Co.: Anderson & McFarland 230 (Ky, Vi); McFarland & Anderson 230 (Ss). Jefferson Co.: Gun J 189 (N); C. W. Short S.n, [Louis- ville] (L1, Pr). Union Go.: Shacklette 500 (Ky). County undeter- mined: C. W. Short s.n. [west of Kentucky, 1831] (T), s.n. [sandy the Ohio & Salt Rivers, rial 1835] (iy—-1778), 8 » SNe [185)] * (ca-=379995) 5 Snow sn. (S). TENNESSEE: Blount Co.: 2 Ruth 3260 (N). Cheatham Co.: tO, S. Rect iecach s.n. [Kingston Springs] (N). Decatur Co.: G. L. Ames s.n. [May 9, 1855] (Mi). Hamilton Co.: Collector undesignated s.n. (Chattanooga, Aug. 1868] (Pr). Humphreys CO.: Galloway s.n. [June 8, 1889] (0b—-50783). Montgom- ery Co.: T. L. Anderson s.n. (Clarksville, 186] (Io--32956) Obion Co.: : Jennings & Ji & Jennings s.n. [Reelfoot Lake, 192] (Cm, Cm). County undetermined: G. B. Grant 2810 (Po--26760); Nason Se ne [May 9, 1855] (Ur). MICHIGAN: Allegan Co.: F. T. McFarland 8» a... Douglas Lake, Aug. 15, 1915] (Ky). Cass Cow: Fe. C. Gates 1321 aii). Cheboygan Co.: J. H. Ehlers 472 (Mi), 627 (Mi), 1771 771 (Mi)3 F. C. Gates 13222 (Ka--66182); M. McKee 613 (Dp). Dickinson Co.? C. A. Davis s.n. (Norway, July 2h, '05] (Mi); Fernald & Pease 3L95 (i). Emmet Co.: E.G. Voss 2420 (Mi). Houghton Co.t Ce De Richards 2710 (Mi). Tosco Co-: C. A. Davis sen. [Oscoda, Sept 25 = Hanes s.n. "tras, Aug. a 1953) (Mi). “Tivingston Cow: De Lynch son. [September 9, "19h7] (An--121789) . Menominee Co.: Grassl 2620 [ (is); Mulliken & Cole s.n. [Grand Haven, Aug. ll, 1896] (Mi) « Saint Joseph Co.: Hanes sn. [Three Rivers Park, Aug. 22, 195k] (Mi). Van Buren Co.: W. H. H. Rhoades s.n. [between Papaw and Kala~- mazoo] (Hs). Wayne bon Parker sen. [Fort Gratiot, 1829] (C)- County undetermined: F. D. Kelsey s.n. [August 6, 1899] (Ob-- epg WISCONSIN: Adams Co.: R. T. Brown 116 cis EK. He 0 nis 356 (Ws). Brown Co, : Gillman s SNe TAug. & ] (Pr); Shinn stor (Ca--882593, Ws). Buffalo Co.: Iltis at, 9352 ( (Ws iF as 1962 Moldenke, Monograph of Verbena 291 tis & Noamesi 8136 (Ws). Columbia Co.: C. C. Albers 31,010 (Au); H. P. Chandler 37h (Ca--25178); Marx s.n. [6/3/58] (Ws); F.C. Seymour 10279 (Ws). Crawford Co.: Fassett 9789 (Ws); D. Houghton S.n. [Prairie du Chien, Aug. 1831] (Mi); H. He Smith 755 (B, Ws). Dane Co.: C. C. Albers 36007 ruld pees Sen. (Madison, July 7, 196] (Ws); 1 Bokde, LTP rt s.n. [June 9, 1957] (Ws); R. Burton 123 (Ws); Cull 692 aye . J. Hale s.n. [Madi- son] (Ws); Heddle 706 (ee) ae Te wis) oo7h (Ws); S. H. Watson 8.n, Amiel (Pa). | Dunn Co.: E. A. Baird s.n. ([Caryvi ville, Au Aug. 15, 1920] (Ws); Buss s.n. [Falk City, July 8,,190] (Ws). Grant Co.: Busch 25 (Ws). Green Co.: G. Cottam 5h (Ws); E. W. Fell 58- 729 (Ws); ; PP W. Thomson s.n. [south of Albany, September OG. 19L9] este Green Lake Co.: Hein 65 (Ws); Shinners 1 (Ws). Iowa Co.: Clik » Drescher, Gisvold, “& Leiderman s.n. [Blue Mounds, June 10, et (Ws). Jackaan Co.: Grether 6585 (Ws), 7194 (Ws); H. He smith 6835 (B). Jefferson Co.: P. 0. ~ Schallert B51 H-~ 50k Oh). Juneau Co.: J. W. Thomson s.n. ~ [Armenia, Aug. 18, 18, 1938] (Ws); Wills s.n. {Camp Douglas, June 25, 1957] (Ws). Lasietee Co.: Hartley 2929 (Ws); A. M. Peterson 223 (Ws). Lafayette Co.: L. S. Cheney s.n. (Fayetteville, Aug. 27, 1888] (Ws); P. K. Nelson Son. (Darlington, July 22, 196] (Ws). Lincoln Coes Peroutly s« sen. (Ws); F. C. Seymour Ujhh2 (Ws). Marathon Co.: L. S. Chene Cheney 3286 (Ws). Marinette Co.: J. We Thomson s.n. [Camp Bird, July 13, 13, 1946] (Ws). Marquette C Co.: Grotjan 156 (Ws); I1- tis, Bell, Melchert, Patman, & Witt 12369 (Ws); Titis, Trenk, . & Voanesi 6163 (Ws); (Ws); Koeppen 382 (Ws). Milwaukee Co.: J. S. Doug= s.n. [Wis.] (Je--3152); Shinners s.n. (Milwaukee, Oct. 2, 1937] (Ws). Pepin Co.: Kinney s.n. [Sept. '96] (Ob--99769) . Pierce Co.: Weinzirl s.n. [River Falls, Sept. 3, '92] (Ws). Por- tage Co.: F. W. Stearns 797 (Ws); W. R. Taylor 2608 (Up—75579)5 = u Thomson s.n- [a rnott, Aug. 2h, 1938] (Ws). Ra tA Co. Je vis s.n. (Racine, Jany 7, 1879] (Ur, Ws); Wadmond s.n. ti 778) (al). Richland Co.: M. A. Fosberg 335 (Ws). Rock Co “Cons Re B. Anthony s.n, [Bebit, June 19, 193k) (Ws); E. W. Fell 57-652 (is), 57- 57-741 (Ws); Sleavienn tks (Janesville, June 27, '89] 89] (Ws). ak Co.: C. C. Albers 33019 (Au, Au, Au), 33020 (an); Fassett 2687 (Ws); G Greene & Curtis 5093 (Ws); True S.n. Sen (Greenfield, _ July 3] (Ws), Sheboygan Co.: Goessl s.n. [Plymouth, Aug. 191] (Ws). Trempealeau Co.: Hartley 372 (Ws); F. Je Hermann 895k (N, Ws). Vernon Co.: J. B. Marks s Sone n. [Coon Valley, 1939] (Ws). Wal- worth Co.: Almon s.n. (East Troy, July ), 1926] (Ws); H. C Benke 6077 (N); Blatchford s.n. [July 19, 1875] (ca--882555, N); Kleeberger s.n. [Whitewater, June 7, 1878} (Gg—31300) ; ve Ne Thomson s.n. [Lake Geneva, Aug. 5, 19,0] (Ws); Wadmond 3765 (Ky), 292 PHYTOLOGIA Vol. 8, no. 6 1303 (S, Ws). Waukesha Co.: Lapham s.n. [Ottowa & Genesee, Aug. lith, 1847] (Ws), s.n. [Eagle] (Ws); R. W. Pohl P.1203 (Bt-~-5092, 158] (Ws). County undetermined: Univ. Wisc. Econ. Coll. sn. [Warwatosa, July 18, 1890] (Ws). MINNESOTA: Anoka Go.: Cratty s. n. (Anoka, Aug. 20, 1918] (Io—96199). Crow Wing Co.: A. B. Sey- mour s.n. (Brainerd, Aug. 22, 188] (H--106L67); E. B. Watson Ss. ne [Brainerd, 8-l-1902] (Io—h5775). Goodhue Co.: J. H. Sandberg s.n, [Red Wing, July 1885] (S). Hennepin Co.: G. B. Aiton 982 (Ur, Ur), sen. [Aug. '90] (Dt); Buell 551 (H—58006, No—860h); Fe He Burglehaus 855 (Se—1,961); E. J. Hill 269.1889 (Ur); Ramaley }20 (B1—-h228l); J. H. Sandberg 267 (Fe--2279, To—1135, Ka), S+ n. [Aug. 1889] (Ca—251))1). Jackson Co.: Cratty 5-n- [Jul. 2h, 1923] (Io—109581). Mille Lacs Co.: E. P. Sheldon s.n. (Princeton, July 1892] (N, S). Ottertail Co.: Chandonnet s.n. [July lh, 1910] (Dm). Ramsey Co.: F. Blanchard s.n. (St. Paul, July 31, 1890] (Dt); Saint Lawrence 2 (N). Rice Co.: G. K. Knight s.n. [North~ field] (Bt—31376). Rock Co.: H. A. Gleason 939k (N}. Stearns Cos: Re Gmelin s.n. [St. Cloud, Aug. 1875] (Io--94752)5 Eo Be Watson s.n. (St. Cloud, 7-16-02] (Io—82h0h). Winona Co.? Holzin- ger s.n. [July 1901] (N, Vt). Wright Co.: Jesup 1962 (Dt). NORTH DAKOTA: Benson Co,.: Bolley s.n. [Leeds, Aug. 1h, 1891] (La); Lun- ell s.n. (Leeds, July 2nd, 1898] (Dt), sen. [Leeds, July 2hth, 1898] (Ur), s.n. [July 12, 190k] (Ok), son. (Leeds, July 2h, 1906] (Du—77501, S), sn. [Butte, July 28, 1907] (Dm, N), SMe (Leeds, July 22, 1909] (Ur), sen. [October 16, 1916] (Dm). Bil- lings Co.: Facey 98 (Au--121808). Bottineau Co.: Boivin & Dore 8198 (N). Burleigh Co.: F. P. Metcalf 366 (Gg—31311). Cass Co.: Bergman & Stevens s.n. {[Fargo, July 19, 1910] (Ca—587762, Cm) « Stark Co.: Holgate son. [Aug. 1h, 1908] (Gg—31306, N, Pl— 103657). County undetermined: Brenckle 2282 (Dm). SOUTH DAKOTA: Brookings Co.: F. C. Gates 166h7 (Ka--81020); E. C. Panmel Soe (Brookings, July & Aug. 1902] (Io—22907). Brule Co.: EB. Ne Wile cox s.n. (Chamberlain, July 1893) (Ob—50772). Butte Co.: Fe be Bennett 618 (Gg—310863). Custer Co.: Degener & Peiler 16061 (Ny, Si); Jones & Jones 10657 (Ur); Over s.n. {June 19, 1914] (Se— 14959). Deuel Cow: C. & We 348 [Herb. Corbett 1039] (We); Te As Williams s.n. (Clear Lake, 20 August 189] (N). Fall River Co. Whitham s.n. (Hot Springs, Sept. 15th, 1909] (Io—50180). Hanson Cos: G. F. Weber s.n, (Farmer, 8-18-27] (F1—21085). Hughes Co.s L. Adams s.n. (6/27/9h] (Ur). Lawrence Co.: W. P. Carr 179 (Vi), sen. (Vi--6012); C. M. King s.n. [Spearfish Canon, Aug. 1916] (Io—-7h097); Rydberg 93h (N). Lyman Co.: Over s.n. (July 1, 1914] (Se~-14956). Meade Co.: A. D. Pratt s.n. [Piedmont, June 1962 Moldenke, Monograph of Verbena 293 1895] (Ob—50782, S). Pennington Co.: Enander s.n. [1-h/9/1921] (S); J. W. Leedy 431 (Bt-—-55627). idberte tee Co.: Over 14387 (Du-- 14612). Shannon Co.: Visher 2028 (N). Spink Co.: Brenckle 39-6 (Sd—26485), 39-7 (N); Petrak s.n. [20.VII.1950] (S); A. eee Ricksecker 66 (Ca—-75503, Ob--50781, Po--2152h9, be ae, Um- 2h, Ur). todd Co.: Tolstead -527 (Io—15162). reseed ee Fes- senddé sen. [Hurley] (Io—70756). County undetermined: Clark 6059 (B); Petrak 50072 [Badlands National Park] as 3.n, (Bad Lands, 3.VIII.1950] (S). KANSAS: Allen Co.: A. S. Hitchcock s.n. [Allen Co., July 1896] (Ka). Atchison Co.: ‘Glothier & Whit- ford s.n. [Aug. 2, 1897] (Ka); A. S. Hitchcock s.n. [Atchison G6: Oct. 3, 1896] (Ka); Jeanette 162 (N). Barton Co.: A. S. Hitchcock s.n. [Barton Co., ., August 1895] (Ka). Bourbon Coe: it Bs De. Thompson 57 (Lw). Butler Co.: A. S. Hitchcock s.n. [White Water, July 1892] (Ka); D. J. Johnson s.n. [July A) (Ka—71718) . Chautauqua Co.: A. S. Hitchcock sen. [Aug. 8, '96] (Ka); We He Horr s.n. [July 5, 1930) (Lw). Cherokee Co.: A. A. Jacobs 102 (Ka—71,038); Kellerman s.n. [Galena, July 8, 1887] (Ka). Cheyenne Co.: F. C. Gates 16835 (Ka—76132); Steller & Jacobs 337 (N)j Ne White s.n. [June 16, 1931] (Ka——77629). Clark Co.: A. S: Hitch- cock sen. (uimecla, July 1892] (Ka). Clay Co. Imler Ce 1 (ka 93368) Decatur ey gsi eme man s.n. (Oberlin, July 1l, 1 se (Ka); R. Le ucGregor 2116 (Iw). Dickinson Co.: A. S. Hitchcoc SMe (Dickins n'G0.; arm 1896) (Ka); Imler s.n. [6/8/29] (Li). Doni- rH oo: —— son a sn, [July 31, 1897] (Ka). Douglas [july 6, 1896] (Ka); W. H. Horr E.570 (Ca-ie08s, “Ge 3S 19 o--16637, Ok, S, St, Ur, Vi, We); R. - Re be MeGregor 16 (Ly) ; Snow s.n. [August] (F1--21092). Edwards A. S. Hitchcock s.n. [Edwards Co., Sept. 1, 1897] (Ka). nik doe Clothier & Whitford s sen. [Auge 20 & 21, 1897] (Ka). Ellis Co. Bondy 269 (S), sen. [June 7, 1935] (we), son, [August 19, 1935] (Go); A. S. Hitchcock s.n. [Ellis Co., July 1895] (Ka); Imler s. n, [May 12, 1928) (Lw). Ellsworth Co.: A. S. Hitchcock s.n. [Elle worth Co., July 1895] (Ka). Finney Co.: Menke s.n. [Garden City, May 31, 1891] (Ka); Wilson & Miller s.n. [7-10-12] (Lw). Ford Co.: F. C. Gates 15973 (Ka——7161); A. S. Hitchcock s.n. [Bucklin, July 1892] (Ka). Geary Co.: F. C. Gates 17838 (Ka), 10696 (Ka); Gayle 511 (C); Imler s.n. (May 1929] 9] (Lw). Gove Co.: rhs 8 S. Hitch- cock s.n. [Gove Co., July 1895) (Ka); Inkenberry 6 (St--15962). Graham Co.: A. S. Hitchcock sen. (Graham Co., June 20, 1897] (Ka). Grant Co.: C. H. Thompson 2 (N), 212 (Ka). Gray Co.: A. S. Hitch 294 PHYTOLOGIA Vol. 8, no. 6 cock sen. [Gray Co., Sept. 1, 1897] (Ka). Greeley Coo: J. P. An- — derson sen. [Aug. 26, 1908] (Io—53321); M. Reed s.n. [Tribune, July 29, 1892] (Ca-—10826), sen. [Tribune, Aug. 3, 1892] (Ka). Greenwood Co.: W. He Horr sen. [June 28, 1930] (Lw). Hamilton Co. Clothier sen. [May 17, 1898] (Ka); A- S. Hitchcock s.n- [Cool- idge, July 4, 1892] (Ka); Rydberg & Imler 1008 (Iw). Harvey Co.: Clothier & Whitford s.n. [Aug. 2, 1897] (Ka); Doell 40 (Ly); Mrs. F. Muck 53 (S)- Hodgeman Co.: As Se Hitchcock sen. [Hodgeman Co., August 1895] (Ka). Jackson Co.: Clothier & Whitford s.n. [July 28, 1897] (Ka). Jewell Co.: W. He Horr 439) (N)3 J- Be Se Norton s.n. (Webber, Sept. 23, 1895] (Ka). Johnson Co.: Clothier & Whitford s.n. [Aug. 7, 1897] (Ka). Kearney Co. A. S. Hitchcock i129 (Ka——7963, Ka, Mi, N), sen. [Kearney Co., Auge 29, 1897] (Ka). Kiowa Co.: Kellerman "a" (Ka). Labette Co.: Gwar 5 (Ka- 94531). Lane Co.: A. S. Hitchcock s.n. [Lane Co., August 1895] (Ka). Leavenworth Go.: A. S. Hitchcock s.n. [Leavenworth Co., May 1896] (Ka); W. H. Horr s.n. [6/23/30] (Iw); R. L. McGregor 828 (Iw). Lincoln Co.: A. S. Hitchcock s.n. [Lincoln Co., July 1895] (Ka). Linn Co.: A. S. Hitchcock s.n. [Pleasanton, Aug. 1892] (Ka). Logan Co.: A. S. Hitchcock s.n, [Logan Co., July 1898] (Ka). Marshall Co.: Clothier & Whitford son. [July 25, 1897] (Ka); n, [July 1885; Herb. Prager 18631] (Gg—31298). Mitchell Co. He V. Dwyer sen, [July 1896] (Ka). Montgomery Co.: As se Hitchcock Sen. [Montgomery Co., August 1896] (Ka); W. He Horr s.ne {July >» 1930] (lw). Morris Cos: A. S» Hitchcock s.n. [Morris Co., August 1895] (Ka); W. H. Horr son. [July 17, 1929] (Lw). Morton Cos: Ae S. Hitchcock s.n, [Morton Co., August 1895] (Ka); J. N. Rose 17106 (La); Rose & Fitch 17106 (N). Nemaha Co.: Clothier & Whit- ford s.n. [July 27, 1897] (Ka); A. S. Hitchcock s.ne {Nemaha Co«, Dec. 11, 1896] (Ka). Neosho Co.: Hummer son, [1898] (Ka). 0sage Cost E. Ge Spalding 258 (Ka—92565). Osborne Co.: Neher 94 (Ka- 79838). Ottawa Co.: A. S. Hitchcock s.n. [Ottawa Co., July 1895] (Ka). Pawnee Co.: A. S. Hitchcock Sen. [Pawnee Co., August 1895] (Ka). Pottawatomie Co.: G. Een Son [56521950] (S); Imler SMe [June 7, 1929] (Lw); Norton, Clothier, & Pond s.n. {July 189 ] (Ka). Pratt Co.t A. S. Hitchcock san. [Pratt co., aug. 1696] (Ke)- Rawlins Co.: L. B. Fry 8.n. [Atwood, May 1891] (Ka). Reno Co.? A. S. Hitchcock Sen. (Turon, July 1892] (Ka). Republic Co.: De K. Thomas s.n, [Scandia, 1890] (Ka). Rice Co.: Ae S- Hitchcock Se n. [Rice Co., August 1895] (Ka). Riley Co.: F. C. Gates 12655 Tit), 12748 (Ka—7283, Un—57); Imler sen, 15/27/21) (Lm)s Naso® 1962 Moldenke, Monograph of Verbena 295 June 25, 1889] (Ka); Mayhew (Ky). Rush Coe: A. S. Hitchcock s. n. [Rush Co., August 1895] (Ka). Russell Co.: ic Ly Hitchcock as ne [Russell Co,, July 1895] (Ka). Saline Co.: Eccles 10676 (11); Hanein 18 (Ka—76789); A. S. Hitchcock sen. [Saline Co., June 1896] (Ka). Scott Co.: F. C0. Gates 10608 (Ka~-75859); A. S. Hitchcock s.n, [Scott COs, “Rugust 1895] (Ka). Sedgwick Cos: T. Le Andrews s.n, [Wichita] (Io--l8)85); Aquinas 122 (Ka—87579); A. _ S. Miller s.n. (Wichita, Mar. 1892] (Ka). Seward Co.: Kellerman sen. [Arkalon, June 27, 1888) (Ka). Shawnee Co.: Norton & Cloth- ier | sen. [July 4, 1895] (Ka). Sheridan Co.: A, S. Hitchcock SoM. [Sheridan Co., June 21, 1897] (Ka). Sherman Co,: | A. Se Hitchcock Sone [Goodland, June 26, 1892] (Ka). Smith Co.: Kellerman SNe [Smith Centre, ory, 10, 1888] (Ka). Stafford Co,: H. T. ? H. T. Porter Se n, (Stafford C , duly 13, 1896] (Ka). Stanton Co.: A. S. Hitch-— cock s.n, Facies Co,, August 1895] (Ka). Stevens Coe: are. Henry 103 (Vi); A. S. Hitchcock s.n. [Stevens Co., August - 1895] (Ka). Sumner Co.: A. S. Hitchcock s.n. (Wellington, July 1892] (Ka). Thomas Co.: A, S. Hitchcock sen. [Thomas Co., June 27, 1897] (Ka); W. C. Wallace 10 (Ky). Trego Cos: As Se Hitchcock s. n. [Trego Coe; July 1895] (Ka). Wabaunsee Co.: Tas 8 Ss. Hitchcock — S.n, [Wabaunsee Co., Aug. 23, 1896] (Ka); Norton & Clothier s.n. [7-L-1895] (Ka). Wallace Co.: Demaree 2911 (Le); : Sci. Exped. . 85 Sen. [July 5, 1885] (Ka, Ka). Washington Co.: Imhoff 6 (kKa— 93160), Wichita Co,: A. S. Hitchcock s.n. [Wichita Co., July 31, 1895] (Ka). Woodson Co.: G, M. Clark sen. [Batesville, Aug. 20, 189] (Ka); W. He Horr sen. 17/11/30] (Lw). Wyandotte Co.: K. kK. Mackenzie s.n. [Argentine, May h, 1895] (C), sen. [Kansas city, Aug. 15, 1897] (Ka, We). MISSOURI: Adair Co.: C. S. Sheldon 1962 (N), 6512 (N). Atchison Co.: H. A. Gleason 9285 (N). Clay Co.: Broussard s.n. {VII .2.19)7] (Je--5959) ; C. Je Elmore s.n, [Liber- ty, Aug. 17, 1923] (Je—3151); Gier 8077 (S, S), sen. [VII-10-50] (Je-—-6780); K. K. Mackenzie 109 (S). Greene Co.: Blankinship s.n. (Springfield, 6-11-1807] (Ka). Jackson Co.: B. F. Bush 207 (C), LOlO (N), 12675¢ (Bt--1552h); K. K. Mackenzie s.n. [May iB 1.5 1896) (N); G. Smith s.n. [Kansas City, July 26, 1871) (Up--17093). Lin- ocin Coe: J. Davis s.n. (Whiteside, 1-6-15] (Vt). Marion Co.: J. Davis s.n, spree) 46 (Au), Sete (13-6-18] (Gg--31312). Mercer Co.: Palmer & Steyermark },1310 (N). Pike Co.: J. Davis (Du-- 87277). Platte Co.: Radloff sn. [June 29, 1956] (Ur). Saint Louis Co.: Flatfelter s.n. [Near St, Louis, 7-16-92] (Mi); Letterman s. & re) almer Steyermark },1085 5 (N). Wayne Co.: E. J. Palmer 6106 (Po—21766k). Saint Louis: Drummond SMe {[St. Louis] (Au—121010) ; Eggert SNe 296 PHYTOLOGIA Vol. 8, no. 6 [Herb. Geete 5699] (Go); Engelmann 335 (Pr), sen. [St. Louis, — 188] (Au--121797) ; Letterman 78 (Po—-267658); M. Martens s. . (Br); Riehl 9 (Br, M). “RRKANSAS: Bradley Co.: Demaree 25503 Faas 25505 (Sm). Clark Co.: Demaree 17826 (N). Conway Co.: ye: Dem- aree 25085 (Bm). Craighead Co.: Demaree ee 7027 (N), 27014 (N, Ok, St). Desha Co.: Demaree 26522 (N). eng = Demaree 18053 (n), 24901 (Sm). Garland Co.: Demaree 2012 » Sm). Hot Spring Co.: Demaree 17958 (N, Pl--131512), 512), 1869 an "19335 (N, Sm). Lawrence Co.: Demaree 30767 (N). Logan Co.: Os: Mrs. H. S. Young s.n. [1925] (Cm). Newton Co.: D. M. Moore 32347 (Au). Prairie Co,: Demaree 37763 (Sm, Z). Pulaski Co.: Denaree 8623 (Du—231hh, Ge--237051, N); Hasse Sih [May 9, 1885] (N). Saint Francis Co,: Demaree 11415 (Bt— —31371, V); S. McDaniel 112) (N). Scott Co.: ; Demaree 18179 (N). Stone Co: Demaree 42729 (Gg). Washington Co.: F. Le Harvey 62 (Cm, Ca, Mi, Ur), Sen. (Curtiss 1962] (I), s.n. TFay- etteville, May] (Ur). County undetermined: Welch 1108 (Mt. Maga- zine, Ozark Mts.] (Dp). LOUISIANA: Orleans Par.: Cocks s.n. [N. se July 1901] (T1). Rapier Par.: C. R. Ball 553 (N)- MONTANA: oadwater Co.: Hitchcock & Muhlick 11525 (N). Carter Co.: Lovi- sik 911 (Mn—35077), 1098 (Mn—35078). Cascade Co.: Fe We aAnder- son 102 (Ca--251)0), 333 333 (Mn--11789), sen. [Sand Coulee, , sept. 1888] (Mn—-15380); V. L. Marsh 668 (un--26731). Custer Coe: me gemt 17 (Mn—35076). Dawson Co.: wie H. Pammel s.n. (Glendi Aug. 1h, 190k] (Io--l629). Flathead Co.: Rogers & Rogers 1068 (Pl—12762). Gallatin Co.: J. W. Blankinship O (Me, Mn——0082, S, Vi), sen. [Bozeman, July 22, 1898] (Mn—30578, N); Hitchcock & tuhlick 12)78 (N); W. W. Jones sen. [Bozeman, July 1 15, 1901) (Ca- 164937). Jefferson Co.: H, Re Flint 20-31 (4n—316)7). Lake Co.! B. T. Butler 637 (N), 639 (N); M. E. Jones son, [Ravelli, ih, 1909] (Po-- Pie “Lewis & Clark Co.: B, T. Butler 110 (N); Cramer sen. [Herb. U. S, Dept. Agr. 016] (Mn— Mn—-1016h); Jo | Je Te Howell 7899 5 (a haces; G. N. Jones 5351 (Ur); F. D. Kelsey S+ ne ENA (Ob—50867). Liberty Cos: Rishoff 627 (ln—~26733) + Missoula Co,: Bache 270 (Mn—-27227); Fe Ae Barkley 1685 (Mn-- 28558); M. E. Jones sn. [Fort Missoula, August 2, 1909] ee 74943); Kirkwood 2095 (Ca—35233h, ca~-882602, uin—16133, N); Ve L. Marsh 591 (ln—337h2); F. H. Rose 213 (uin-—3607h) , 533 (Min-- 33088, P P1--9876}) ; Je H. Sandberg s.n. (Missoula, Aug. 1 1892] (Ca—-25139, Pl—1,503). Park Co.: Hitchcock & MuhLick 13567 (N)5 Ce W. Simpson 7a (Ca--882607, N); Suksdorf 917 (P1—139839) « Powder River Cos: F. He Rose 335 (Mn—3292, a BteaOeGen Powell Co.: Hitchcock & Muhlick 11520 ( (N). Rosebud Co.: J. W. Blankin~ ship sen, [Forsyth, June 29, 1898] (Se—L35l5), s.n. “Trosebud, June 29, 1898] (Se—l3359); B. D. Lund 49310 (un--8987). Sanders 1962 Moldenke, Monograph of Verbena 297 Co,: Sandberg, MacDougal, & Heller 972 (N, Po——-8)18)). Silver Bow Coe: Babich 381 (Mn--26732). Stillwater Co.: E. W. Elliott 29 (Mn— 35083, N)3 G. Peterson 1006 (Mn--35071) . ~ Sweetgrass Co.: Eggleston 7970 Os Hitchcock & k & Muhlick 13305 (Ca--757473, Ge, N, S). Teton Teton Co.: » Franklin, & Gillett ; 566 (Vi). Toole Co.: Rishoff 575 Rc ee Wheatland Co.: C. L. Hitchcock 2335 (Mn- 23760, Po—22151h, Rs——1)0h6), 2h27 (un-—239],7, Po—222072, Rs-— 13978) . County undetermined: Engelmann s.n. [July 1839] (Br); Pe D. Kelsey s.n, [Montana] (0b--50865); Lamson-Scribner 209 [Indi Creek] (Pa); Lejeune s.n. (Br); 3 Severy 17 [Kootenai National vis est] (Mn—1309, Ob--5086)) ; Wied—-Neuwied sen. [1833] (Br). IDAHO: Ada Co.: J. A. Clark 55 (Ba, Ca—l50173, st ge N, Po—63861, 8), 279 ee “Ca--h50172, Ca-~882615, Du--24159, Fg—l31l, N, N, P1968, Po—-63855, S); Mulford s.n. [Boise, yelp 192] (C). Adams Cost . H. Christ 12950 (N); Re J» Davis 2412 (P1--10993). Bannock Co.: J. H. Christ 2321 (N), 9003 (N)3 Cronquist 213-36 (Va--l697, Ua--l6498); R. J. Davis 69-3 (N), 327-37 (Id); In- termountain Herb. 3730 (Ca--52859,, Ua-—11381). Benewah Co.: ws Epling s.n. (Santa, July 1929] (La). Boise Co.: R. J. Davis 162- 37 (Id (Id); Hitchcock & Muhlick 9960 (N, P1--155093, | Se—9320h). Bonner Co.: Js He Christ 1155) (N); C. Epling 10380 (La), s.n- [Granite Station, July 1929] (La); Pammel & Dudgeon s.n. (Sand Point, 8-05] (1093832), Canyon Co.: J. F. Macbride 200 (Cm, N, P1—22665, Ur). Clark Co.: J. H. Christ 5807 (N); Cronquist 751-37 (Ua--305h6, Ua-—6499); H. J. Rust 57 (Gg--31305). Frank- lin Co.: C. B. Maguire 17171 (N, N, Pl—12h7i7, Ua—h7h22). Fre- mont Co.: “Cronquist 1473 (Ua--33710); Merrit & Wilcox 776 (N)- em Cos: A, Beard 11862 (N). Gooding Co.: E. Oe: BE. Knox s.n. [Gooding, June 2, 1931] (Sd--23977). Idaho Co.: 7 H, Christ 12668 (N); Colson’ s.n. [Horseshoe Bend, May 17, 196] (S); R. Je J. Davis 3207 (F1127067); Q. Jones 366 (N). Kootenai Co.: J. He Christ ist 10256 (N), 12973 (N); H. J. Rust 457 (Gg——-31301). Latah Co.: Midrich 13677 (N)j J. H. Christ 16k6 (N); Stillinger 309 (Mn—-35080);— Bak. L. Turner 2279 (Tl). Lewis Co.: Hunter 102 . (P1—6881). Mini- doka Cos: Christ & Christ 18535 (N). Nez Perce Co.: W. H. Baker 5895 (N), 5918 (N); J. He Christ 2529 (N), 3955 (N), 9066 (N)5_ Gail 13679 (N) Ownbey & & Weber 27h6a (Gg--381233, N), 27u6b (Ge- 381231); Sandberg s.n. [June 1892] (La), 26h (Gg-—31309); Sand- berg, MacDougal, & Heller 26h (Du—2h160, Fe, Me, N, Po--8L181). Payette Co.: J. H. Christ 9578 (N); R. A. Harper sn. [Payette, Aug. 4, 1890] (S, Um--2l1, Um—2)7). County undetermined: L. F. Henderson 4070 [Link of Big Lost River] (Mi); Sandberg 972 . (Ge— 1308). WYOMING: Albany Co.: Benson 5059 (Po--267 3 A. A. Nelson 1652 (Po—706L6, Ur), 7671 (Cm, 1o—22910, N, Po—-63871); L. O- 298 PHYTOLOGIA Vol. 8, no. 6 Williams 1060 (Ca--882611, Gg—210261, Go, I, N, S, Ua-—11380). Carbon Co.: Cantelow s.n. [Rawling to Laramie, July 15, 192] (Gg—316096). Crook Co.: L. W. Carter s.n. [Devil's Tower, 8/1h/ 1897] (N). Goshen Co.: Petrak k 50081 (N), sen. [hVIII.1950] (S)- Hot Springs Co.: 0. Degener 1300 300 (N). Johnson Co,: Tweedy 3365 (N, Se—20576). Laramie Co.: R. J. Davis 319-W (Id); Frem Fremont s.n. [Evening Camp from N. to South | Fork, July 12] (C); F. We Johnson 13 (N); Ownbey 29), (P1--98767). Natrona Co.: Goodding 207 (Du— 75655, N, Po—63896). Niobrara Co.: F. H. Knowlton milton 148 (N). Park Co.: Marston 359x (S). Platte Co.: C. ra Porter 3987 (Ca--71)781, N, S). Sheridan Co.: Reppert, Paddock, & Ellis 83 (Io--111,62); Tweedy 2663 (N). Yellowstone "flational Park: Bright s.n. [7-30- 19] (Cm); Mearns 3432 (Du—2)163), 5009 (N). County undetermined: Black 36-5182 [Sherman Pass] (Be—}5869); Hayden Survey son. [Aug. I = Sept. 13, 1870] (N); A. Nelson 320 [Blue Grass Hills] (C, Ca, as B. Taylor 26 (Cherokee, Park Colorado] (Ok). UTAH: Beaver Co.: Parker, McClintock, & Robbins 6347 (Ca-—~736876, N). Box El- der Co.: Brendel s.n, [Brigham City] (Ur); B. Maguire 228 (Ua 11372). Cache Co.: A. D. Smith s.n. [August 3, 1936) (Ua—-l7458) . Emery Co.: Clover & . Jotter 193k (Mi); M. E. Jonee Sa Srisiild 159677, N, Po--70155). Garfield Co.: ee A. B (Mn 35070) . Grand Co.: Rydberg & Garrett 8365 (N = sea tee Coe: eaeats & Starr 39 (Vae-11370); Cutler 2432 (Ca--882609, Gg—2712b1); Maguire & Becraft 2786 (Ua--11379). Kane Co.: B. Maguire 18937 (Ua--l,7)21); F. M M. Stone 316 (N); Tidestrom 2390 (Mi). Millard Cos: D. D. Keck 632 (Ca--,23603, Du--199162, Po--19725); Maguire & Becraft 2785a (Ua—11371). Morgan Co.: Pammel & Blackwood 3960 (To—32026). Piute Co.: Rydberg & Carlton 7043 (N). Rich Co.: 3 Me E. Jones 1488 (Ua--11373), s.n, [Alta Wahsatch Mts., Aug. 1879] (Br, Go, N). Salt fase. Ce Co.: M. S. Baker 1,835a (Ro), 14835 (Ca—-L21125) 5; M. K. Clemens s.n. [July 10, 1908] 8] (Du--2162); Ae O. Garrett 1726 (N), “Sent (ca--88260); Jepson 21228 (Ca); M- Ee Ee Jones s.n. Son. [City creek Canon, Sept. 1, 1900] (Po-—7069), SMe {city Creek canon, Sept. h, 1905] (Gg—153863) ; F. E. Leo 85/83 (Po—21h222), sn. [July 11, 1885] (Ob—-$0868); Pammel & Blackwood 3529 (Io—83752), 3589 (Io—28506, Io--7028); Rydberg & Carlton 689h (N); S. Watson 623 (C). San Juan Co.: Rydberg & erg & Garrett 9200 (MN). Sanpete Go.: Olsen s.n. [July 12, 193) (N, Ua- 78 (Cm, N), (Cm, Gg—-236313). Utah Co.: H. Re ca roams coe) ween ces faiecaen : Larsen 775) (Ua-- 35864); Waterfall 15097 (st). Washington Co.: S. B. Benson 32 (ca--5S638 26); Degener & Peiler 1676 (N); Gierisch 569 (N, Ua-~ 4719); F. W. Gould 2027 (Ca--852592, N); Edw. Palmer 1,01 (C, N, Pa), s.n. (St. George, 1875] (Bc). County undetermined: i: Capt. Ni % jee : ! 4 2 p21 7 fe i} Pe : 7>0O #9 ok e z if to : cap (A ae iD SS z wos : o i 4 a ei 3 G ij £ i | B ~ u 8 ic @ © at. S bn | $ re ints bp aaa we par} é Br : Cy = out cn MOU TS aye ORNS : = Weg @ : ofp ie Te ee ae Tec : 1 ae U Lapa, 2s : Sen | ALY aor 7 ie ca : = Vy, 3 Bs bal Bc bracteata in the United St ates Verbena oS ee je Figure 5 Distribution of 1962 Moldenke, Monograph of Verbena 299 Bishop s.n. [1872] (Ka); G. A. Black 36-7647 (N); E. W. Davis Se ne n. (Cove Fo re Fort, 11 July 1927] ' (Um—2h9); Elrod s.n. [Utah, '86] (iin—6881) ; edget sod 83 [Preuss ee near Clay's Ranch] (I); Kellogg 1739 (Ca—€h1322); ae 557. (Ba, B1—h2253, aaa Ca—-9009 39, 39, Du--26996h, N, cree Ua—-31277), 8096 [Herb. Forestry Serv. dries (ax"=25513, AL , Au, Au, Ba, Bt--58933, Ca-- Ob—2011. , or== 23943, S, Se——20)25, Se--54.785, St, St--19052, Ua--39356, Up, Ur, , f09t (cae -poocte N), 8473 5 (os 500515, Dp—1043, Gg-- 310860, N N, Pl—-110326, S, Se--84785, Ua--l3899), 873a. (Ca— 700513)5 T. Craig 1168 (Po—-18576); Goodding 2312 (Ca--1662h, N); Louise Kellogg 1739 (Se--62361); LaRivers & Hancock 57h (can-862 9, Ca--900550); S, B. Parish 10258 (Ca-~183650, Du-- 91086); Tidestrom 9582 (S); | sis 1913 (Ca--9005L8, 359), 198), (Ca--900540, Du--286897, N, N). Elko Co.: A. H. Holmgren 159 (Ca--676171, N); P. Be ial te 3069 (Du--1.630) « annoy Co.: M. E. Jones s.n, (Palisade, June 1h, ~1882}] (Po—70652). Humboldt Coss Archer 1 196 (Mi) ; Hillman s.n, (sept. 1, 1897] sitlagaige Lander Co,: Co,: Goodner & penning 1097 (H=-69222) « Lincoln Co.: G. W. Gullion 582 (Ca=52111); M. E. Jones sen. [Panaca, Sept. 6, 1912] (Po--69911); Minthorn 253 . (Ca--501 74). Lyon Coot Re Ae Allen 259 (Ca~-882590). Nye Gos: Bentley s.n, [Currant, June 1916] (Du- 11,0008, N, Po—-129057, Po—-2676)2); Goodner & Henning 1329 (Ca— 900517, Fs); LaRivers & Hancock 716 (Ca--9005l)9). Storey Co.: C. A. Purpus 5946 (Ca--10813). White Pine Co.: M. A. King s.n- {Warm Springs, Oct. 1917] (Gg—~—31310). COLORADO: Adams Co.: Dee aree 29526 (Le, Sm, Sm). Arapahoe Co.: Garfield Univ. Sci. Exped. 332 (Ur). ‘Archuleta ae C. F. Baker 56h (B, Ca--10860, N, Po-- 64520, S); F. S. Earle 25 (N); Schmoll 1074 (Bl--42302). Boulder Cos: Re E. Buchanan s.n. (Ward, 5/10/1901] (Io—107610); H. W. Campbell “22 (B22; A. T. Evans 189 (B1—2266), 239 9 (Bl-- 42268), 273 (Bl--2267); Ewan 11418 (Gg——345033), Pl. Exsicc. 300 PHYTOLOGIA Vol. 8, no. 6 S. Young s.n. [Boulder, Sept. 1913] (Au). Chaffee Co.: Waterfall 10906 (Ss (St). ¢ Cheyenne Co.: Cory sen. [9-3-19)0] (N)- Conejos Co.: Ramaley 1298) (Bl--h2286), 14147 (Bl—2288). Crowley Co.: M. Te Jones s.n. (May 28, 1932] (Bl—-2361). Delta Co.: Burritt 97 7 (B1- 2252). Denver Co.: Babcock s.n. [Cory 27781] (N); Bethel s.n. (Denver, June 1893] (Fe), sen. [1910] (Gg--31302); Demaree 15271 (NW); Eastwood 25 (B1—2260, Ca—-148189, Gg--31317, Ok), See [Denver, July 1886] (B1—1,2256) ; Ferril s.n. [July 29, 1901] (Bl- 42253); Heustis s.n. [Platte near Riverside, June 26, 1916] (Bl-- 2261), sen. (Riverside, 6-26-16] (B1—-l2251); p24 Holm 61 (I), 8s n, [6/8/1896] (S), son. [11/8/1896] (I); Redfield 521 (Pr); Schmoll 735 (Bl—-h225h); H. G. Smith s.n, “[Denver, Aug. 17, 1888] (Ga, Pl—22622). El Paso Co,: Atkinson s.n. [July 15, 1895] (Pl— 86890); Bacigalupi 919 (Du--195802, Po—172348); Biltmore Herb.’ 1082e (S); S. L. Clarke s.n. [September 19-21, 1895] (C, C); Clements & Clements 9 (Du—2h158, Io—-23730, N, Ob--86567) ; Fry He Ehlers Ehlers 7630 (Mi), 7677 (Mi, Mi), 7761 (Mh, Mi); Harper & Harper Sn. S.n. (Manitou, July 12, 1900] (Io—133189); M. E. Jones 143 (Br, Go, N, Po—183131, Ua-—11377); Loftfield 119 (Bl—-2205); R Rusk + 8s ne [c. C. campus, 7/10/1942] (Je); A. H. Smith s.n, [Manito Springs, August 1878] (Up--1709)); M. S. Young sn. (Green eae tain Falls, 9/25/17] (Au). Fremont Co.: T. S. Brandegee S.Ne {Canon city, 1877] (Ca--169222); Waterfall 10877 (St). Gunnison Co.: C. F. Baker 920 (N, Po--63872); H. N. Wheeler 626 (Bl— 42263). Huerfano Co.: McAllister s.n. [Cuchara Camps, 8-16-23) (Au); Rydberg & Vreeland 5675 (N); Stigall s.n. [6/16/3h] (Bl-- 42269). Jefferson Co.: L. S» Ehlers 218 (Mi); L. H. Pammel s-Me (Golden, Aug. 1919] (Io—100972); Ramaley 9906 “(al=-1,2273) , 9913 9913 (Bl--2272); W. W. Robbins 5780 (Bl--42271); Zobel sn. [Deer reek Canyon, May 25, 1934] (Bt—2h675, Gg—220519). La Plata Co. Moinat 22 (S)5 Way sen. [Oxford, 1916] (pl -12559) "Tariner Con: C.F, Baker s.n. (Fort Collins, 6-18-93] (Po--65183), SMe [Fort Collins, 7-21-93] (Po--6389)); M. S. Baker 763b (Ca-- 421120, Ro); Collector undesignated s.n. [Ft. Collins, July 1926] (3) J. H. Cowen 2018 (Fc), 2019 (Fe, Mn--6883, P1l— 22568), s.n. [Fort Collins, June 12, 1895] (N), sen. [Fort Co, July 29, 1896] Somme wi orandall 172 [Halsted's Am. Weeds 172] (Ah, C, Fe, I “Va--11375, Ua--27139 We), 31 (F ¢); buteo: me ag Soruag eat. July 1926] 722 Vio 2285 Lappin 185 (Ok), 188 (au, Ok); W. F. Marshall 2017 (Fe); Panmel & Johnson 1749 (Io—2h47h); Preston s.n, [0-1-1921] (Fe); Ramaley 2758 (Bl—-L2259); C. S. Sheldon 38 (Al), s.n. [12 July 188k] Lb) Om )3 G. F. Weber s.n, [Estes Park, 8/26/39] (Fl—21091). Las Animas Co,: Beckwith 161 (Ge—31307), 163 (Ca—-21675h). Logan 00+? E. J. Kraus s.n. [Sterling, June 19, 1926] (H--107720, “um--2h)5) « 1962 Moldenke, Monograph of Verbena 301 Mesa Co.: S. G. Stokes s.n. [June 15, 1900] (Du—-2h17h, N). Mon- tezuma Co.: | Bader son. (Bl—)227)). Montrose a9 C. F. Baker 28 (Ca~-350)2, Ga—10l885, gia oo Ny Ar 2256 Payson 93) (S); W. Stewart sen» [Ute Pas aes 1937] (La); B. P. P. Walker er 02 BS sie): rosase 06 53 ‘pons & Fitch 17503 (N). Rio Grande Co.: Ramaley & Johnson 14568 (Bl—l2289). Saguache Co.: Waterfall 1217 (Au--121809, Ok, St). Washington Co.: Ram 1262 (B1—I2287). Weld Co.: Ewan 12129 (En, Pl--122568); Oster- hout s.n. [New Windsor, June 17, 17, 1901] (I0-~20753, Po--63899), Ss. n. [June 18, 1905] (Du—-91087) . County undetermined: J. E. Chase s.n. [Mamnoth, 1904] (Ms); J. M. Coulter s.n. [Platte River, June 26, 1873] (N); Degener & Degener 270h2 (1 (N)$ McCosh & Greene s.n. [July-Aug. 1877] (M1, Pr, Va); L. H. Pammel s.n. [Lookout Mt., Sept. 25, 1927] (To0-—103087) ; 3 Scovell s.n._ mans [Colorado] (Mi); Vasey 527 (Ms). NEBRASKA: Adams Co.: R. de Lima 150 (I), 233 (I), hab (Ny). Antelope Co.: Wernecke 82 (Gu--E.77h5). Buffalo Co.: 5.3. R. Holmes s.n. [Kearney, Aug. 1889] (Fe--2506); Pammel & Brownlie Se n. n, [Kearney, 6-27-99] (Io—-15343). Cass Co.: M. E. Day s.n. s.n. [June 28, 1916] (Ob--99770). baie a Engberg s.n. rreaiot 189] (Je). Douglas Co,: Eastman s.n. (Omaha, June 26, 1915] (Ur); Kleeberger s.n. [Omaha] (G Ti aiose) Fillmore Cos: Muenscher & Muenscher 11607 7 (Ca-~727596). Kearney Co.: C, Hapeman s.n, [Min- den, May 31, 19 '91] (Hp); H. Hapeman 8451 (H—-559), Sa Site Triage Sent. h, 192h] (Bt--5069, Up), sen. [Minden, July 10, En, Pl--61802, St--9230), s.n. [June 27, 1931] (Au), s.n. [Min- den, June 1S, 1933] (Ms, N)} Herb. Univ. Nebr. 258 (Au, Au). Keith Co.: C. 0. Erlanson 2253 (Ob—50776), 2255 (Mi). Kimball Coot Ae rt Heller 11303 (Au, Du--156892, 3 BEG) Lancaster Co,: E. Armstrong SoM, sen. [June 26, 1923] (Go); Herb. Okla. Coll. Women 312 (Ok); J. L« Sheldon sen. [Lincoln, September 1897] 97] (We), S.n. Sen. (Lincoln, 7210.98] (We). Lincoln Co.: Plank s.n. [July 1896] (N). Madison Co.: F. J. Hermann 50 (Mi). Merrick eat ure E. 5 sees sen. [E. J Hill 2001902] (Ur). Nemaha Co.: . Elmore Sete Johnson, Oct. 1, 1890] (Je—3153); us L. Sh sep sn. (Peru, June 16, 1900] (We); Shubert s.n. [July 10, 1933] (0b—590777). oe Co.: A. E. Allen s.n. [Talmage, June ert 1891] (Je—3150); E. H. Jones 8 (Se—5653). Phelps Co.: McMillan sn. [Holdrege (To—53119 - Platte Co.: Clevenger s.n. [Columb bus, June 10, 190k] (La). Thomas Co.: Krautter s.n, [Halsey, June rien 190k] (Up- 17132), sen, [Halsey, July 26, 1907] (Up—l6608); H. J. Webber s. n. [Aug. 7, 1889] (c). ee Co.: Lallman s.n. [June 15, 9%) (Bt--5451). York Co.: M. K. Clemens s.n, [York, June lh, 909 } (Io--107). Platte Islands [Buffalo Cos): E. R. Holmes s. Kearney, June 23, 1887] (Ca—216740). OKLAHOMA: Beaver Co.: Hinman 202 (St); Penfound P.387 (Ok). Beckham Co.: P. Hart 37 302 rorrtOLUuGcia Vol. 8, no. 6 (Bt--)8615); Fe - Pennell 10557 (N). Bryan Co.: W. S. Blain s.n. (Durant, July 6, 1935] (Bt--29682). Caddo Co.: Demaree 12533 (N, Ok); Edw. holga 2l6a (N); Van Vleet 32 (0k--20),80). Cherokee Co.: C.S. pees 66 (Sst). Cimarron Co.: Demaree 13290 (Ok, Ok), 13307 (N, Ok, Ok, Po--251719); F. C. Gates 20968 (ka—92216) 3 3 Ce M. Rogers 668 (Ok) 5 Smith & Rinehart 62, _ 82, in n part (0k); Stratton 5197 (i, N, W, St--261%); Waterfall xe 9 (Ok, St), 3147 (Ok, St), 10756 (St). Cleveland Co.: Bamford s.n. [April 30, “911] (Ok-- 20481); F. A. Barkley s.n. [5/19/28] (Ok--10289); Bayliff s.n. [July )] (0k--1029h); Bruner sen. (Norman, 8/2/2h] (Io—-1170Lb, Ok--10287, Or--18975); Demaree 1271 (Ok—-22198); C. T. Eskew 1938 (Ok), 2016 (Ok); Harness s.n. [May 5, 1911] (Ok—20)82) ; gains & Van Valkenburgh 5526 CGE) Keesecker & Osborne 56 (Cee Osborne & Keesecker 56 (St-~12769); Steen 207 (0k--20507), Se ot ne [6/28/19] foes 10296) Pusonet 30 (Ok—10293)5 Stemen 251 (0k); C. Whaley l,2 (Ok—-10291). Comanche Co.: M. K. Clemens 11750a (Ba, Gg—37ihil); W. Se Myers 31, (0k); . “Reber tson 75 (Ok). Creek Co. A. J. Sampson 73 (Or--0069). Delaware Co.: C. S. Wallis 2381 (St). Ellis Co.: Bruner s.n, (8/2/2h)] (pn—178618)- Garfield Co.? sag ige 379 (Ok). Garvin Co.: R. W. Pohl 5203 (Sm). Greer Co. (Ok--10286). Harmon Co.: G. W. Stevens 1051 (Ok, St-~ 9225). H Harper Co.: McMurry 1180 (Ok). Kay Coe: Byler 277 (St)5 M. G. Wolfe s.n, [Sept. 15, 1931] (Ok--10273). Kingfisher Co.: Grace s.n. [June 20, 1935] (Ok—19055). Kiowa Co.: G. W. Stevens 1192 (Ok). Latimer Co.: Hopkins, Nelson, & Nelson 517 (Ok). Le- flore Co.: Blakley 1462 (Du—66336), San. (Ge W. | Stevens 1162] (St—9223, Ur). Lincoln Co.: 0. M. Clark 2223 (B); Payton 85 (St—-922h). Logan Co.: E. Beck 47 (We) ;Keyser s.n. Tc. W. Stevens 6044] (N, Ok). Mees Co.: C. S. Wallis 3439 (St). McClain Co.: Me Hopkins 12 (Ba, Ok, Ok, St—-12975), 13 (iin--33215, Ok, Ok, St 1297), 13360 Se Rs Ok). McCurtain Co.: H. W. Houghton $.Ne [G. W. Stevens 3726] (N). Murray Co.: Broadbent t 83° (Ok); M. Hop- kins 6108 (Ca—882596, Ok, St—-2118, We); Hopkins & Cross 6l51 ios; Stratton 732 (St--261h1, St—261)2). Muskogee Co.: Ee . Little 22h (0k--10285), 1585 (Ok—10290). Noble Co.: Harding a (St—-17568), Oklahoma Co.: W. S. Myers 90 [10-27-2h] (Ok), Se n. [2-22-25] (Ok); F. W. Pennell 10579 (N); Waterfall Mh2 (N, Ok, Si). Payne Co,: Ambrose 18 (Ob—81690); C. 0. Chambers SNe [Summer, 1919] (St--9231); Davy 102 (Au); Fry 110 (Au), Sen. 19 20-35] (Au); Gardenhire 77 (Ok); Graumann 39 (St--17553); Harding 257 (Ca--882606); Mader Mader 22 (Ok); D. McLean 15 (Um—2h8); Powers 31 (N); L. E. Rose sen, [August 2, 1939] (Bl—)2296); Waterfall 13 (St). Pittsburg Co.: Be. P. sesbiokes 39 (Ok); E. Rogers SMe 1962 Moldenke, Monograph of Verbena 303 [June 12, 1957] (St). Pontotoc Co.: D. McCoy 722 (St), 1703 (St). Pottawatomie Co.: E. D. Barkley 296 (Ok—lhh7h); F. A. Barkley 1452 (Mn—-28598). Roger Mills Co.: J. Engelman 0) (Ok). Rogers Co.: Buffington 99 (Ok). Swanson Co.: G. W. Stevens 1192 (N, St—- 9229). Texas Co.: G. W. Stevens 391 (Du--65310, Ok, Ok, St--92)0, St—-92h1); Q. Williams 36 (Tr—19291). Tulsa Co.: M. B. Clark 363 (St). Woods Co,: G. We Stevens 680 1/2 (0k), 2889 (Ok). Woodward Cos: P. J. White s.n. [July 9, 1900] (Dm). County undetermined: Gowan 26 (Ok--10292); "R. E. J." sen. [7/1/19] (Ok--10295); Jeffs Son, [Winding Stair Mtns., July 1, 1919] (Mn—25705); B. A. Kee- Secker s.n. [June 11, 1936] (Ok--20601); G. W. Stevens A.1)62 (Ok), A.156) (Ok). TEXAS: Bailey Co.: Harns 82 (Um—-2h3). Bexar Cow: J. B. Davy 11 (Ca--25138). Bowie Co.: Correll & Correll 12398 (Ca—-907252, Sm). Brazos Co.: Reeves 192 (N). Brewster Co.: Collector undesignated L.20-l159 (Sr); Cory 9147 (Fs); Parks & Cory 9147 (Tr). Cochran Co.: B. B. Harris 526 (Nt). Collin Co.: Timmons 743 (Au). Comal Co.: Depprich 7767 (Sm). Culberson Co.: Tharp 43253 (Au, N). Dallas Co.: F. C. Gates 20972 (Ka--92225); Cs L. Lundell 11015 (Sm); J. Reverchon 120 (Pa); Shinners 26626 (Go). Denton Co.: B. B. Harris sen. [7-20-27] (Nt). El Paso Co.: B. Barlow s.n. (7 July 1911] (Ca--882613, La); Me K. ER ee [El Paso, June 7, 1917] (Po--175086); Lee, Berkman, & Tharp 16199 (Au, N); Pringle s.n. & Paso, 11 et (Vt); Shinners 5911 (Sm); Thurber 90 (T); Vasey s.n. [El Paso, May 1881] (S), Son. [El Paso, 1881] (Fl1—z1089); B. H. Warnock 1000/5818 (Au--122572); Waterfall 6622 (N); C. Wright 45h (Ca--215384). Erath Co.: Hoising- ton 83 (Au, Au, Ca--733348, Ok). Garza Co.: Tharp s.n. [7/9/L1] (Au, N). Grayson Co,: Anderson & Hood 313 (Ok); R. Bebb 1412 (Ok). Hale Co.: J. F. Brenckle 48080 (Gg--38123, N); Waterfall 3703 (St); Whitehouse 9926 (N). Hansford Co.: Cutter 33, in part (Ok); G. J. Goodman 5289 (0k). Hockley Co.: B. B. Harris s.n. [10-10- 27] (Nt). Huspeth Co.: K. B. Reed Sen. (Grayson, Nov. 3] (Au); Tharp 46152 (Au, N). Hunt Co.: Legget son. [Loneoak, 7/15/1927] Au); Tharp 2929 (Au, Au). Jeff Davis Co.: T. Head 10 (St). John- Son Co.: Whitehouse s.n. (Cleburne, 6-15-1930] (AU, Ca--882603, Mi, N). Lamb Co.: Demaree 0876 (Gg); Tharp 627) (Au). Lipscomb Co.: C. S. Wallis 7118 (St). Lubbock Co.: Demaree 7562 (Au, Du-- 205782, H—52537), 770 (Du—205758); Fe W. Gowld 7122 (Au— 123217, Ca—38005); E. J. Palmer 12h96a (Au--121765, Ca—h25608) ; ZL. Reed 3940 (B1—]:2292); Tharp 620) (Au). Menard Cos: Co 29734 (N); Parks & Cory 9765 (Tr). Mitchell Co.: R. W. Pohl 4998 (Sm). Moore Gos: Jespersen & Jespersen 2708 (Ca--77L452, N). New- von Co.s Tharp sen. [7-23-39] (Au). Ochiltree Co.: C. S. Wallis 30k PHYTOLOGIA Vol. 8, no, 6 4755 (St), 4762 (Au--123217, St). Potter Co.: York & Rogers 1 (au--123198). Presidio Co.: Hinckley 69) (N, N, Sm), Son. (Marfa, July 1936] (Au, Au, Fc); Steiger ger 1308 "(N). Randall Co.: Co Cory 13530 (Md); E. J. Palmer 13861 (B); Parks & cs 13529 sa Rea- gan gan Co.: Parks & & Cory ory 1890 (Tr). Roberts Co. : Ce S. W 494-1, in part , (St). Sherman Co.: Weaver 17793 os oe OB. Swisher Co,.: Whitehouse 9961 (N). Tarrant Co.: Ruth 109 (Cm, Io-- 113679, Ka—67108, N, N, Nt, Ob--50779, Up, th. Be Whitehouse 16263 (N). Taylor Co.: S. M. Tracy 8001 (Au, Cm, Es, , Vt). Terry Co. E. L. Reed 3774 (Au); Tharp 14506 | (Au, Um-~2)6). yh Green Co.: Cory | eor® (Po—19226) . Travis Co.: Herb. Univ. Texas s.n. [Lake Austin, 5/10/33] (Au), sen. [5/10/33] (Au). Walker Co.: yo: Warner 5. sville, May 20th] (Hu). Wheeler Co.: Parks & Cory 1615 (tr) » 16146 (tr). Wichita Co.: Mahler 1258 (St). Wood Co.: C. Le Lundell 12081 (N). County undetermined: i: Thurber 607 [Valley of the Rio Grande] (N); Warner s.n. as NEW MEXICO: Baca Co.: Her- shey 2352 (Sm). Bernalillo Co.: | C. C. Ellis 221 (N); M. Ee sae s.n. [Albuquerque, 9-l)-188)] (Po--26766) ; Koelz Koelz sen. [Sandi Mts., June 28, 1926] (Mi), s.n. [Albuquerque, e, September 8, 1926 (Mi). Carson tess Hitchcock, Rethke, & Raadshooven 1180 (Ca 603818, Du—257%7, Gg——267612, ads i Se—l15728) . Chaves Cos: Earle & Earle 351 (N, Po-—63868) $ N. A. Palmer 60, in part (S). Colfax Co.: oy ¥. F. Baker s.n. ([Cimarron, Jul. Ol] (B); Lucas 16, (Au); P. C. pale 14249 (Du--12287h). Curry Co.: White- house 8738 (Au). Dona Ana Co.: Archer 335 (Mi); Campbell & Keller 735 (Ca--882608) ; grec ea 09 (Ca--10811, Du--75569, Ka, N, “Po ay sn. (Mesilla Valley, May 17, 1893] (Ur), s.n. [Organ €ts., Aug. 28, 1897] (Po——70878, Tu--98879), sen. [June 7, pet (N); Wooton & Standley 3330 (au, Du--2)177, Fs, coeer Mea G809 5B Richards, & Moeller 11963 (N, ‘YN, Ua—-;718) ; 0. B. PE 137 Caer scotia , Ca-—882810, Dm, Du--9115 ’ N, Po—70887, W. S. Stewart s.n. [Pinos Altos, June 26, 1364 (Bl—l2297, Fn, La, La, Mi). ago Go.8 Geb. Fisher sen. [Nara Visa, July 8, 1911] (Ew) « Rio Arriba Co.: Waterfall 1111) (St); C. B. Wolf 2925 (Ba, Du~191126, Gg—-170h6, Rs—9995). Sandoval Co.: Koelz SMe (James Springs, July 4, 1926] (Mi); Rousseau 35089 (vi). Sa San Juan Cos: Wooton 2831 (Ob--50876). San Miguel Co.: Arsene 169h (B), 18343 (Gg—187237), 18543 (Po—-148758), 18593 (Po—1h8392), 18634 (N, Po--1,8686), 18885, in part (Po—-1)8523), 18961 (Po--1,8657)» 18964 (N, Po--1486)3), s.n. es Vegas] (N); P. G. Standley 51 (N); Whitehouse 8739 (Au). Santa Fe Co.: Avatce: 15 1 1573 eer 158h0 (N), sen. [vicinity of Santa Fe] (B); Clear s.n. [July 6 197] (Gg); Marcelline 1819 (Mi); Renner 29) [Herb. Forest Serv. 1962 Moldenke, Monograph of Verbena 305 43460] (Ca--137930); P. C. Standley Mss a, USh9 (N). Sierra Co.: Beals s.n, [Lake V Valley, August 190 ] (ua); ci Cutler 2057 (Ca- 882610); 0. B. Metcalfe 897 iauceaie” “revit 718), 1008 (ce— 31303, N, “Po—63858, Po Po—8}185, Vi). Socorro C Eggleston fae (W—1524)13); H. H. Rusby 81 (St—2h0s), 336 i, Mi, Ob—50869, — Pr, Se—1960), san. [Ft. Craig, Aug. 1888] (Dt). Taos Co.: Thi tehouse 8740 (Au). Union Co,: Demaree 12278 (N, 0k), 13278 (0k, P1--8),105, Sd——21049). Valencia Cos: O. Degener 873 (Ms); Wooton S.n. (Aug. 1, 1892] (C). County undetermined: Englemans.n. [Therma, August 17, 1934] (Ok--18362); Fendler 585 (Pr), 507 (Ca-- 711045) ; C. Wright 199 (T). ARIZONA: Apache Co.: Collector 1 undes- ignated s.n. - [Navaho Reservation, July 1918] (Ca—202336); eae Goodman, & Payson 2951 (Du—-272803) ; Deaver 3902 (Fg—-5966), (B, Fe- Fe—-£967); Goodman & Hitchcock 1317 (Bt--12931, ee Du--191999, Gg--183907, Mi, Mi, N); Herb. Univ. Arizona s.n. [Naw aho Reservation, July 1916] (Fs, N); Peebles & Smith 13568 (1 (N, To); L.c. Whitehead s.n. [Navajo Reservation, June-Sept. 1916] (Ca—882612). Cochise Co.: Darrow, Phillips, Gould, & Pultz 1486 Pomel Tu-—-10880). Coconino Co.: Aubineau 5 (Fg—-8037); Re R. S. Beal s.n, [Flagstaff, 30 June 197] (Ca—9h256) ; Benson 9618 (Po==2676L1), 9649 (Po—267655); Clover 5070 (Mi, Tu--19627); Clute 93 (N, Ur); | Collom s.n. [Grand Canyon Village, Sept. 1, 19h3)] (Fe—91h5); Deaver 2509 (Fg-~3096), 378h (Fg--5253), 136) (Fe—8038); Demaree 2782 (Gg); Ellison 1107 (La); Fulton 6283 (To), 9654 (Ca--50265); Goodding 45-48 (Gg--41599), 26319 ( (8), 1,92 (Fe~8035), 1595 (Fg--5030); H. C. Hanson A.147 (Au, BL T229k, or—20),8 9, Ur); A. A. Heller 15783 3 (Ca--7 201 2h, N, al S791); Herb, Univ, Ariz. s.ne [Flagstaff, June-July 1915) (Tu); J. Te Howell 2hho5 (Gg—34261h), 24600 (Gg—3k2813); H. E. Lee Sony (Flagstaff, 9-23-%] (Tu); MacDougal 286 (Ca—-187k71, Dp, To--387h9, N)5 Peebles 14346 (To); He H. Rusby sen. [Oak Creek, May 30, 1883} (C, Mi); P. 0. Schallert s.n. [5/30/43] (N, Ur), Sn. [mest of Winslow, 5/15/43] (N), sen. [6/16/43] (P1--133007, Ur), sen. (9/1/43) (au, Ok); J. J. Thornber s.n. [Flagstaff, Aug- ust 5, 1907] (N, Tu--35537). Gila Coe: F. W. Gould 3918 (Ca— 73717h, N); Parker, McClintock, & Robbins 6111 (Ca—736880). Gra- ham Co.: Bohrer 393 (Ge—377372); 3 J. J» Thornber 5554 (Ok), 5674 (Tu). tee Co.: Ahles 9223 (Ur, We), 9408 (Ur). Navajo Coos Eastwood & Howell 8872 (Dp--h677, Ge-—261698) ; L. Hanilton s.n. (K. F, Perker 533 8338] (S); G. J. Harrison Sug (To); Herb. U Herb. Univ. Ariz. s.n. [Navaho Indian Reserv., July 1916] (Tu); Mrs. Re We Hoyt 8.n. en. (Ft. Apache, 1893] (C); Klinger s.n. “[Newcomb's, June ah, 193k] (Tu); Peebles & Fulton 9636 (To); Pultz 1714 (Tu--16148) ; Whitehead s.n, s.n. [Navajo Indian Reserv., June-Sept. 1916] (Tu); 306 PHYTOLOGIA Vol. 8, no. 6 Whiting 85/2798 (To). Pima Co.: Darrow & Haskell 2072 (N), 2172 (Ca——70522), Tu-—-161)3). Pinal Co.: Peebles & Harrison 1754 (To), 2061 (Io—128)75). Yavapai Co.: W. We Jones s.n. [Verde Valley, July 25, 1920] (Ca~-07936), sen. (Camp Verde, July 7, 1922] (Ca- 407935, Ca--07937); E. B. Kurtz s.n. [July 18, 1937] (Up); Loomis = pay 3 Peebles, Harrison, & Kearney 2692 (To), 42h3 (La); He He by 336 (Pa), 780 80 1/2 (1 (Mi), sen. (June, 1883] (Cc, (c, Pr), a Tsay (Up—-17091). Yuma Co.: S. B. Parish 890 (Tu--98876) . WASH : Adams Co.: Jeffrey s.n. [June 22, 1946] (Ca--755256); St. ie Beedon, Davison, Ransom, om, & Ridout Be Cores eigen Cos: nticheock & Muhlick 831 (Ca—-9684Lh, N » P1—-15509 ag gn Co.: Cotton 621 (P1l--22585) ; Gort 136 (a Sane Ro); . Henderson s.n. [Prosser, May 26, '92] (P1--22621); G. Ne tee a7 (P1—-h1136) ; Cc. S. Parker 653 (P1—-l, 7538); L. S. Rose 8153 ( (Go, N, S, S); R. Sprague s.n. [Aug. 16, 1938] (Or 37663). Chelan Cos: iuenscher & Muenscher 11215 (N); re 7192 (Du--268787, N, Sd—-2h252); Raven 10112 (Gg--l09688); J. W. Thompson 6901 (Du—208092, Or—-28268, Se--21)78), 6930 there 21477), 10808 (St—15931); Whited 1373 (P1--22581), sen. [June 21, 1904] (Or—-12376). Clark Co.: Suksdorf s.n. [Vancouver, Aug. 16, 189] (P1—138591). Douglas Co.: Sandberg & Leiberg 30 ae ca—170667, Gg—31316, u, P1—-22616, S). Franklin Co.: | 694 (Pl—100118) ; Jennings & Jennings 8888 (Cm), 8911 (Cm); Rav Raven 10363 (Gg—-1,0851,5) . ak Cov: H. W. Smith 1178 (Pl—8)538)5 Je W. Thompson 6732 (Se--21h79). King Co.: Otis 603 (Gg—31313, Pl— 128712). Kittitas Co.: Cotton 1363 (Pl--2258h); Otis 2030 (Pl— 93351); H. W. Smith 1410 (Se--30501), 1776 (Pl--0h537); Js We Thompson 9067 (Ga——518319, Du—-22),006, N, ey vores Se—-21))80) « Klickitat Cos: F, E. Lloyd s.n, [Dalles, July 189] (N), SMe [23 Aug. 1894] (N); Suksdorf 207 7 (Pl--138)28), =e a: P1--138363)5 Sen, [July 22, 1881] (Ca--192061, Cm, Mi, Pl1—7200 be J. a Thompson 11109 (Se--15933). Okanogan Co.: Batie pril 2h, 1922] (Bt--3589h); Dillon 832 a Be etiol)5 Fiker uy (Pl--76371); R. A. Harper sen. [Aug. 9, 1890] (S); Me Be a a June 26, 1911] T (Po--69916) . Spokane Co.? urge 1420 (Se--108)7), s.n. [May 1930] (Du—-209173, Se— 21876), Sen. [June 1931] (Se--21922); Suksdorf 1976 (N, Pl— 13803). Stevens Co.: Kreager 7 (Ca--582614, N, Pl--22580, Se— 68369, Um—-22), 87 (P1—-22576), s.n. ae "aug. oe, 1902] Pl—22575), Wallawalla Co.: Wilkes S Expl. Exped. s.n [Wallawalla & Kooskookoo] (T). Whitman Co.: Eastwood & Howell 07h ae 1962 Moldenke, Monograph of Verbena 307 Yakima Cos: L. F. Henderson s.n. [June 12, '92] (Se--1)9h9); Suksdorf s.n. [June 7, 188) ~(P1--138)30) ; 3 Vasey 468 (N, Pl— 22579); Watt s.n. [Aug. 1895] (Pl--22617). County undetermined: T. S. Brandegee 1032 (Ca--190923); J. H. Howell s.n. [Lower Cas— cades, June 1506) (P (Pa); Jennings & Jennings s.n SNe (Cm). OREGON: Baker Co.: W. A. Weber 348 (N). Clackamas Co.: | E. P. Sheldon 8856 (N). Deschutes Co.: tas F. Brenckle 51440 (Le). “Gillian ©. Co.: Applegate 6519 (Du~239351)5 & C. D. Mell s.n. (Arlington, Aug. 26, 1903] ie “Hood River : Gorman s sen. [Lower Cascades, July ' 90] (Se--1))9)0); L. F. ahi oes m 797 (Or--8853); J. T. sowail s s.n. [Cascades, June 1586] (Se—21h76); Lyman s s.n. [Cascade Locks, 7- 30-95] ( Dt); E. P. Sheldon 12179 (Du—92289). Malheur Co,: Ferris & Lorraine 12993 (Du--37802h); Fleischman s.n. (Ontario, 5-20-30] (or--35766); R. A. Pendleton s.n. [September 22, 195k] (N). Mor~ row Co.: Hills 6 (Or—25509), 14 (Or--25512), 26 26 (Or—-25510); L. S. Rose Lihl3 (A 3 (al, Se ae “Gg-—-310865); J Je a Thompson 1,880 (Du--179362, Po--215928). Multnomah Co.: Suksdorf 3014 (Pl— 138413), s.n. [Aug. 15, 89h] (P1—-138)27) « Sener oe Co.: We Ee Lawrence 13 (Du 100815) . Umatilla Co.: H. E. Brown 58 (Dm, N); Dana sen. Stteaasaten, Oct. h, 1957) (Or—-34h36); Evinger s.n. Pendleton, 9/3/27] (Or—-31167); Gorman s.n. [near igh hai » '05] (Mn--16973) ; Griffiths & Hunter 16 (N); W 806 (Du--10h71); Leiberg 912 (N); W. A. Weber 3106 jose hs N)5 Zivney 5 (Or--38566). Wallowa Co.: E. P. Sheldon 8202 (N). Wasco Coe: Abrams 9487 (Du——133965, Po--127890) ; T. S. Brandegee Sn, [The Dalles, 1832] (Ca—169223); Gilkey s.n. [the Dalles, — June 120, 192) (or—18252, Or—l5975); Me EB 3 M. E. Jones s.n. [The Dalles, jane 27, 1897] (Po--7065h) ; Lyman | Sen. [The Dalles, 7- 30-95] (Dt); E. P. Sheldon 12179 (Du--75590); Suksdorf s.n. [June 8, 1897] (Pl——130h26); J. W. Thompson 2821 (Du—171167, Se--21) 17h). Washington Co.: Solbrig 2909 (Ca--177947). Wheeler Cos: Ebert s.n. [July 20, 19hh] (Or--9935). Hayden Island [Coos Co.J: J. W W. Thompson 3789 (Du--171163, Se--21473), 3909 (Du-- 171801), Sauvin's Island [Multnomah Co.]: J. T. Howell s.n. (June 1886] (Or—8852). County undetermined: W. C. Cusick 1967 pconie Oregon] th Bh Ewoetines ; E. — == (N, Pa). IFORNIA: Kern randegee s.n. (Came: on Lake, March 2, Le (ca--10h81.7); epee wT 2667 (c-89516, co 16650, ms Re Short 295 [U. S. Forest Serv. 21066] (Ca--661057); Twissel- 126, (Gg--391012). Los Angeles Co.: ee 343 (Du--1],3860) , Tit oe 143861, Po—l005), 257) (Du—2hl6k, N, Po--156282), s. ne og Aaya pine’) (Ca-—205361); M. K. seaports son. [Bixby, August 1, 1910] (Ca—185367); Braunton s.n, [May 10, 1902] (Du— 9559), Sen. [5/15/1902] (Du--75592); L. T~ Chamberlain sn. (N); 308 PeEETOLUGIA Vol. 8, no. 6 Fosberg S.l105 (N, S, Up); Condit s.n. [Gardena, July 27, 1908] (Ca——156150) ; Bs M. Hall aa sericea I. M. Johnston s.n, [Nigger Slough, s. Calif., 111, 1917] (Da—10859h, Bos McClatchie s.n. Fane teas. 5/3/18 96] (N, N); Parish & Parish 1596 (Br, Du--91089). Madera Co.: H. Barnes s.n. [Madera, Sept. 25, 1930] (Ca—l20788). Merced Co.: Eastwood 17956 (Gg——-176070) ; Jepson 1748) (Ca); Mason & Smith 8252 (Ca—192059); Nobs & Smith (Ca—192060, Gg—l17903). Modoc Co,: Woods & Tidestron 2719 Be Orange Co.: D. L. Crawford s.n. [Laguna gg 7/2/16) (Ca--205360, Po--l00); Fosberg S.1020 (Up); E. R. Johnson 11285 (Rs—30h)5); Munz 2207 (Po--LOly), 63h9 th 6598 (Po 17hk1l); E. L. Paddock 12733 (N); peirson 4651 gerne En). Riverside Co.: Ewan 5220 (Ca—l72156, Du—2l27i1, —19)565, Po=~182),70). San Bernardino Co,: S. B. Parish 81,90 "(pu--91088), son. [3 June 1891] (Ca); Parish & Parish on (Ca--25135, Du-- 90891, N, Or—885), Pl1—22620, S, Ua-—11376, Ur—17h58). San Di- ego Cos: ’ abrams 3198 198 (Du-- —-2)166 , N); Cleveland ia "33 (Sd--6788) ; Fenoge 1235 (Po--1)8016) ; peut s.n. [San Diego, 7-8-1884] (Cm), Sen. SoM. [Se De, 9-8-188)] (Mi) » 3, [San Diego, Sept. 188) (Pa). San | Joaquin Co.: M. K. Brandegee s.n. [Lathrop, Sept. 1907] (Ca-- 139765); Jepson 10285 (Ca); Suksdorf 7a (Pl—138h08), 7b (Pl— 138407). San Luis Obispo Co.: C. Dudley S.n, [Atascadero Lake, July 4, 1936] (Gg—235651); R. J. , Ferris 9770 (Ca—652680, Du-- 269818, Gg—310871, Mi, Se--52775); Jeps Jepson 15888 (Ca); Edw. Palm- er 342 (Pa). Santa Barbara Co.: J. L. Fox s.n, [Santa parbara] _ (I); # H, M. Pollard s.n. [Aug. 6, 1952] (Gg--385809). Siskiyou Cot W. B. Cooke 15356 (ca—hs319, Du--277307, N, Or—-l2075) « Stanislaus Co.: Hoover 9 (Ca, Ca--767060, N). Tulare Co.: Cong- don s.n. (Oct. 1881] (Du—2h165). Ventura Co.: Fendge 1838 (Po 15297); H. M. Pollard s.n,. ee Lake, May 3, 1946] (Ge-—- 333017). County undetermined: M. K. Brandegee s.n. [Bissell Sta~ tion, Mojave Desert, July 16, sat (Ca--185368); Edw. Palmer S- ne [1876] (Da--200619) ; Vises sen, (Bl--lj2291). CHANNEL ISLANDS: Santa Catalina: Fosberg $.503 (N). MEXICO: Baja California: Mearns & Schoenfeld 2925 (Du—9556); C. R. Orcutt 1228 (10), 15h9 (Dt, Ur), s.n. (Tijuana, l-7-1885] (Ca—-2136), sen. [AUg. Ae 1889] (iti). Coahuila: E. G. Marsh 121, (St); Edw. Palmer 1018 (Io, Pa, ee Federal District: Rutten & Rutten-Pekelharing ng 378 (Ut). San Luis Potosf: Parry & Palmer m5 (Pa). Sonora: Gregg Se n. [near St, —— Soe 2, 1819] (T); P. B. Kennedy 7032 032 (Ca 373595) . SWEDEN: 8.n. [VIII.1923] (Bw, Go, S); C. Sandberg sen. [6/9/1913] Gre aes, Kausch s.n. [Aug. 1915] (La). CUL- TIVATED: Belgium: Lejeune s.n 3.n. (Br); P. Martens s.n. [h-b-L.] (Br). Denmark: Herb. Schumacher s.n. (Cp). France: W. H. Harvey 1962 Moldenke, Monograph of Verbena 309 s.n. [Julio 1818] (Du—166l95). Germany: Herb. Molliano 528 (Br); nated: Herb. W. H. Harvey s.n. (Du—166191) . LOCALITY OF COLLEC- TION UNDETERMINED: Burke 1827 [Rocky Mountains near California] (Ut—-97722); Collector undesignated s.n. (Je); Cranwill s.n. [July 23, 1917] (Ur); F. V. Hayden s.n,. [Sargent's Bluff on banks of Missouri, July 1856] (Dt); Herb, Coll. Pharmacy s.n. (Pa); Herb, Decaisne son. (Br); Herb. Mar Martius s.n. (Br, n. (Br, Br); Herb. Mol- liano 288 (Br), s.n. (Br); Herb. Mus. B Bot. Stockholm 2109 09 (S), 8. 8. n. (S, S)} Herb. Parry s.n. . (Io); Herb. Torrey sen. [Rocky Moun- tains] (T), sen. [N. W. Coast] (T); M. Martens s.n. (Br); C. R. Oreutt sen. (Ca—10),882, Ca--10)883); Edw. Palmer 341 (California, Arizona, etc.] (C); Prier s.n, [Tablequah] (St—9227); 3 Schleuse 35007 (An); 3 Show s.n. .. [Kansas & Nebraska boundary] (C); Wallber Wallberg S.n. [1808] (Lu); A. Wood s.n. (Pa). VERBENA eeecry f£. ALBIFLORA (Cockerell) Moldenke, Phytologia 2: ak Synonymy: Verbena bracteosa albiflora Cockerell in Daniels, Fl. Boulder Colo. 20). 1911. Verbena bracteosa var. SaaS Cockerell ex Moldenke, Prelim. Alph. List Invalid Names 5, i syn. 190. Verbena bracteosa f. albiflora Cockerell ex Coton Alph. List Invalid Names Suppl. 1: 22, in syn. 197. Literature: Cockerell in Daniels, Fl. Boulder Colo. 20h. 1911; Perry, Ann. Mo. Bot. Gard. 20: 30h. 1933; Moldenke, Prelim. ph. List Invalid Names 1,5, 190; Moldenke, Alph. List Invalid Nanes ls. mies § Moldenke, Byte ogia 2: 146. 196; Moldenke, Alph. List Invalid Names Suppl. 1: 22. 19473 “Moldenke, hiph. List Cit. 2: lois & hos Gis) and : 981. 1949; Moldenke, Known ——— Dis- trib. Verbenac., [ed. 2], 26 & 197. 199; Moldenke, Résumé 3 cock & L7l. 1 1959; Moldenke, Résumé Suppl. 2: 3 (1960) and . “*, aint form differs from the typical form of the species in hav- ing white corollas, The type or the form was collected by Theodore Dru Alison Cockerell on the campus of the University of Colorado at Boulder, Boulder es Colorado, on July 15, 1908. It is said inhabit ot the followed it with the expression "n, var.", indicating that a var- ietal status was intended. The only recorded common name is "white-flowered. large-bracted vervain". Three herbarium specimens ae examin tations: WASHING NGTON: Klickitat Co.: Suksdorf s.n. [Binge bie 11, 1895] (N, P1—1380h), son. (Aug. 2h, 1098] (Pl1— 138366) . VERBENA BRASILIENSIS Vell., Fl. Flum. 17 (1825), Icon. 1: pl. 0. 310 PHYTOLOGIA Vol. 8, no. 6 1827. Synonymy: Verbena littoralis @ pycnostachya Schau. in A. DC., Prodr, 11: 542. 187. Verbena littoralis f. angustifolia Chod., Bull, Herb. Boiss., sér. 2, 2 2, 2: 818, hyponym . 1902. Verbena approx- imata Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8: 292--293. 190k. Verbena litoralis var. pycnostachya Schau. ex Briq., Ann. Conserv. & Jard. Bot. Genév. 7-8: 292, in syn. 190. Verbena litoralis var. brasiliensis (Vell.) Briq., Ann. Conserv. & Jard. Bot. Genev. 7-8: 292, 190). Verbena cordobensis Briq., Ann. Con serv, & Jard. Bot. Genév. 10: 100--102. 1907. Verbena boliviana Briq. ex Moldenke, Suppl. List Invalid Names 8, in syn. 19}. Verbena brasiliensis Mart. ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syn. 197. Verbena on Vell. ex Molden- ke, Alph, List Invalid Names Suppl. 1: 22, in syn. 197. Verbena litoralis f. pycnostachya Schau. ex Moldenke, Alph. List Invalid Names Suppl. 1: 25, in syn. 1947. Verbena littoralis f. pycno- stachya Osten ex Moldenke, Alph. List Invalid Names Suppl. 1. 1: 25, in syn. 1947. Verbena hasta L. ex Moldenke, Alph. List Invalid Names Suppl. 1: 2h, in syn. 197. Verbena litoralis f. angusti- folia Chod. ex Moldenke, Known Geogr. Distrib. Verbenac., [ed. 21, on & 198. 1949. Verbena littoralis var. pycnostachya Schau. enke in Humbert, Fl. Madag. 174: 9, in syn. 1956. Verbena rae sis var. veris Nideetar ex Moldenke, Résumé 359, in syn. 1959. Verbena officinalis var. gracibras Lelm. ex Moldenke, Ré- sumé 371, in: in syn. 1959. Verbena Litters nok pycnostachya DC. ex seer Sop mort Suppl. 3: erat t Velo, | Fl. Flum. hy 1825), oon 1: a hO. 18275 Schau, ee rg r. li: 542. 1847; Schau. Bras. 9: 189; 1851; stot diag = Kew. 2: 1178. 1895; j chide ‘Bulle Herb. Boiss., sér, 2, 2: 818. 1902; Briq., Ann. Conserv. & Jar Bot. Gen&v. 7-8: 292--293. 190k; Brig., Arkiv Bot. Stockh. 2 (10): 11. 190h; Briq. in Chod. & Hassler, Plant. Hassler. 10: 180--L81. 190; Briq., Ann. Conserv. & Jard. Bot. Genév. 10: 100--102. 1907; Prain, Ind, Kew. Suppl. 3: 187. 1908; Cee Chloris oebee els Shivs I Southeast. ~ 10. 1930; Stapf, Ind. Lond. 6: 29. 19315; Perrys Ann. Mo. Bot. Gard. 20: 252, 265--256, & 355. 1933; Fern., Rho- dora 0: 371 & 455. 1938; Moldenke, Annot. & Classif. List 108. 1939; Moldenke, Prelim. on List Invalid Names bh, 56. — Moldenke, Suppl. Invalid Names 8. 191; Moldenke, loa 6: 32h—325 rr9ka) pe 8: 428. 1923 Moldenke, __ Ge- ca Distrib. Verbenac., [ed. 1], 3, 5, 6, 9, 10 HET 5 BOs vt lk, 7h, 80, & 101. 1925 woldenke Alph. List an alid 5, ae & 48. 1942; Jepson, Fl. € a (2); 380. ee poe Geogr. Dis rt Meee a 28 2s. 2983 Sampaio & Peckolt, Arquiv. Mus. Nac. Rio Jan. oi 3926 19135 1962 Moldenke, Monograph of Verbena 311 Moldenke, Lilloa 194 gone & 382. 19hhs Cabrera & Dawson, Rev. Mus. La Plata, ser, 2 » 5: 357 & 361. 19h; Moldenke, Castanea 3 ee 1,7, 158, aha 172, 177, ig92, 196, . 200... hap thd pte ka ce ee it 263, 282— -26l,, 288, 289; 291; 296, & 298. 196; K & Flor. a hs bits 4117. 1947; Moldenke, Alph. List Invalid cong » 2h, & 25. 197; Moldenke, Phytologia 2: 327-- tanea 13: “112, hie. 116, 118 , & 121. 19183 Molders, Alph. List 815, 837, fia’ ie? ane’ 852, ope 1, 863, 869, a7 em om 885, 89, 899, 913, 935, 9195 8 * 966 (1949), : 980 80, 9 3. "3853 Look, ’10107 107k, 1 7, 1081, 1087, 1059, 1106, 1 108, 1 116, 1123, 11 aah. "18, 1142, 1147, es ust, 1166, Tt ry rip” 120); 1213, 1222, 122h, N. Molde e, Anal. Ins t. Biol. Mex. 20: 12. 19195 eee: Pca es coer Distrib. Verbenac., [ed. 2], get et” » 7, 61, 73, 9h, 97, 99--101, ”105 to. 153? 197, & 196. “Biss *Moldenke, Sein go 3: 135 (199) and'3: 288, 290 29h, t 305. Me, Moldanks, Revist. Sudam. Bot. 8: 165. 1950; Fern, in A. aseeeaP ed. 8, 1209--. ; Noldenke, Prytoldg! anes 450, & h67. 1951; Stellfeld, Trib. Farmac, fYelionoal 19 a isy 1s "1951; Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 6: 2209 & 2210, 1951; Moldenke in Gleason, New Britton & Br. Il Rp Beet Pe Bee , 19525 Moldenke, Phytologia h: 266 (19 $ and 5: 26. 195h; » Am, Midl. Nat. 52: 313. 1954; Moldenke, Inform. Mold. Set 18” A 99 [4] (195), 49 Spec. 3 (195k), and 51 Spec. h. 1956; Moldenke in Humbert, Fl. Madag. 17h: 5, 7, & 9--10, fig. Ty 3h from s ea~levél to 900 m. altitude, as a feet, in California from 80 to 700 feet, South Africa from 200 to 5500 feet, in Bolivia from 6500 t 12,000 feet Chile from 200 to 1200 m ters, in Arg “1d 60 to 1620 meters, in Mexico from 1 e from 1000 to 3700 meters, an Colombia from 1700 to 3250 met- om April to Decem United States it blooms throughout the year, but mostly from A pril to September. In Louisiana Ewan describes it as "common in grassy places" that it occurs mixed with Ambrina ina ambrosioides in sugarc fields and ped ditch banks. Demaree avers that it is Filla 1 Texas it was found by Gould in cartiis shade along roads through pine forests, while fers As it as a "weed in vacant lots" in Sacramento County. How- ell reports it as a "roadside weed" in Stanislaus County. This potisercess no. 30055, by the way, is said to be from the type region of V. hansenii Greene. Feddema says that the species is common in wet meadows in loam soil exposed to the sun in Nayarit and in grasslands with occasional shrubs on likes clay loam in open sun in Michoac4én, In the Transvaal it occurs in mountainous areas with many a ptayee and streams, some wooded and some grassy areas, on basic- dolomites and Limestone covered with deep humus in many Places, acco: Collectors in South carolina report it there from waste ground, roadsid des, railroad yards an rights-of-way, warehouse areas creek bottom woods, fields, oi forests and forest margins, and in sandy marl along waterways. In Ala been found along roadsides and on old Amerind shell mounds. Kral & Redfearn say that it is "rare on wet womies Hiteiees of nant oheees in Florida. In 316 PHYTOLOGIA Vol, 8, no. 6 North Carolina collectors have found it along railroads and at railway Nera in waste places, about homesites, along road- sides, and o savannas . The GL. "Fisher 48132 at GUteborg, cited below, is definitely ve brasiliensist Perry states that this is "A species very similar in habit to V. litoralis, but readily dis tinguis hed by the differences in in- orescence. * the spikes of V. brasiliensis are short, compac i: sessile, and regularly arranged in open cymes, whereas thos v. litoralis are longer, compact or somewhat pg ick pecincend ‘and arranged in more or less paniculate cymes Common names recorded for V. brasiliensis fe cas "Brazilian vervain", "camaradinha", "erva de pai caetano", "formosa sem dote}! "herva do pae caetano", "jurupeba", "oi", Mquininha", and "ver- as a whole, as well as to V. bonariensis L., V. bracteata Lag. & Rodr., V. corymbosa Ruiz & Pav., V. hastata L., xV. hybrida Voss, V. litoralis H.B.K., V. officinalis L., V. peruviana (L.) Britton, V. platensis Spreng., V. stricta Vent., V. urticifolia L., Bouchea prismatica (L.) Kuntze and its var. longirostra Grenz., Clerodendrum fragrans var. pleniflorum Schau., Priva lappulacea (L.) Pers., Stachytarpheta calderonii Moldenke, S. cayenne ensis (L. C. Rich.) Vahl, S. frantzii Polak., S. guatemalensis Moldenke, and S. jamaicensis. (L.) Vahl. Sampaio & Peckolt (1943) reduce V. brasiliensis Pp synonymy under V, litoralis, as do also Hooker & Jackson in the "Index Kew- ensis" (18 95), while sonmoee (18,7) reduces it to a mere variety of Vv. tiara. Ry species are, however, reasonably distinct. The f following b: ief ia ~ ee to distinguish V. brasiliensis from its imm imeetiate relati 1. Leaves sessile, clasrine 8 + the bas 2. a small, ——l,.5 cm. long, pt pe . wide, subcoriaceous, rrate at Pa apex only or entire; pubescen ence on rachis, “oon +. and calyx g rlandulose Ve termedia. 2a. Leaves opie 5--10 cm. long, 5—22 mm, wide, coarsely ser rate bas ence rachi acon and calyx not glandulose........V. bonariensis. la. —— distinctly petiolate or cuneately teste to a sub- lar bas petio 3. op penta “lax, elongate; pity distant; pareonnst on inten bractlets » and calyx v minute and closely 4 pre x literalis. 3a. Inf erleomanae dense, contracted; flowers ae ss Lv conepeberts pubescence on rachis, bractlets, and calyx brasiliensis: Glazion (1912) calls his no. 14162 "Verbena littoralis H.B.Ke" and 1)162a "V, intermedia Gill. & Hook.", but the former of these numbers is herein cited as ve. brasiliensis. Villafafie describes 5 1962 Moldenke, Monograph of Verbena 317 the flowers on his no. 970 as "amarilla", but this must surely b an error, The printed label for the ious sen. (Sept. 2nd 1903) collection cited below is inscribed "Plants of Mariposa Co.", but B sland is located at the mouth pe the San Joaquin River ange -~ it is very leafy, like some forms of v. oar =e Key yet its inflorescences are very pubescent. Hammarlund 631 much of the aspect of V. parvula Hayek and may prove to be gece species. Tate 206, from an altitude of 12,000 feet at Pongo, Bol- ivia, may prove to be V. hispida Rufz & Pav. The Sprague s.n. 1185 is a mixture with V. litoralis, while Worth & Morrison on 16498 is a mixture wi . donarlensis Lc Glaziou (1911) Tdenti tien nie no. 356 as V. bonariensis, but it is actually Vv. brasiliensis. Specimens of this species have al- so been misidentified and distributed in herbaria as V. angusti- folia Michx., V. bonariensis L., V. bonariensis var. brevibracte- ata K Kuntze, V. bonariensis var. longibracteata Kuntze, V. ephed- roides Cham. V. hastata L., V. hispida Rufz & Pav., V. . interme- dia Gill. & Hook., V. ae aes V. litoralis H.B.K., V. ae Kunth, ve “itera. aracasana Briq., = ‘atecmiie caracasana (H.B.K.) aay v. oe var. carac runt V. littoralis H.B.K., V. officinalis L., V. aerate var. gracilescens Kuntze, Vv. polystachya H.B.K., Ws stricta Vent., ¥. venosa Gill. & Hook., and even Verbesina occidentalis Walt. On the other hand, the Hassler 706, distributed as V. approxima- ta, and the Bassler 6685, dist distributed as V. litoralis var. bras- Titers Briq., are both ve montevidensis S Spreng. it is the ttc tall verbena of ey, common al - road tracks and between cane fields." In Minas Gerais, Brazil, it is said z to be found "in red clay soil, ee especi t tina. Miss Mexia reports it as "common and widespread in hedge- “ows and cultivated fields" in Minas Gerais. In Chile, Elliot 318 © FeerTroOLuoGai: Vol. 8, no. 6 says it is "eaten by cattle", Bryant tells us that in the Cape of Good Hope it is a "large regular but open growing bush 3 fe vi gh and several feet in circumference." Fasciation is seen Barros Sulinanele 8002 in the Britton Herbarium at New York. “The Sprague SMe, cited hereinafter as Fact Meta, Colombia, may a have been collected in Cundinamarca -- str label merely stat "between Villavicencio and Bogota, surname of the sol 1sotes F iss, is misspelled "Freyreis" ot gene Stockholm sheet oc stastiative. of the difficulty of identification attending some specimens of this species is Rojas 105 -- this specimen was originally named v. bonariensis by the collector; then as V. bonariensis var. brevibracteata by Hassler; then as V. litoralis by Osten, and finally as V. bonariensis var. veris, also by Hass- ler. Sanzin 27) was identified as V. littoralis by Hauman, Rojas 1887 as "V, litoralis Kunth vel affinis" by E. Almquist, and Herb. Osten Rie littoralis Kth. ad V. bonariensem L. spectans" by Oste Troe et It is of sakaeaat to note that Small (1933) retiorstee the spe- cies from only Louisiana; ve gee (1950) mpeg mange in the United States from "Fla, to La. and n. to se. Va. ge res in 1952 I further extended it to puree Arkansas ie gt diet California. This is apparently an index to the rate with which it is spreading in the United States Briquet (190) cites Hassler 1027b and 1027d from Paraguay an Regnell A.475 1/2, A.3647, and A.3651 from Brazil, Fernald 1938) sp rginia, while Kuhlmann & Ktthn (197) cite M. Kuhlmann 66 from SHo Paulo, Brazil. Ahles, Bell, & Radford (1958) cite Be Bell 1 2520 from Beaufort County, South Carolina. Perry (1933) cites the following 28 additional specimens not as yet = by me: NORTH CAROLINA: New Hanover Co.: Randolph 100) (G); J. K. Small 5707 (E, G). GEORGIA: Chatham Co.: Kolthoff SNe Sone [27 liay i927) eee FLORIDA: Escambia Co.: Curtiss 61,90 (E, G, Ws S. M. Tracy 8706 [E, F, G, W). Washington Co.: F. S. Blanton 6598 (E, W). ALABAMA: Mobile Co.: C. T. Mohr s.n. (l July 1893] (i). LOUISIANA: Plaquemines Par.: Lloyd & Tracy 20 (E, F, G, W)- BY r she cites the Louisiana specimens under Alabama. What she akie a as "Small 5707" seems to be Herb. Biltmore 5707, at least in the Britton Herbarium at aod York, as well as in the Univers- ity of North Carolina Herbari In all, I have examined S13 Nevbardtie specimens, erie the collections of many of the names involved, and } m photographs. Citations: VIRGINIA: Warwick Co.: Fernald & Long 7585 (N)- NORTH CAROLINA: Bertie Co.: Ahles & Duke 46161 (Hi--121529). Bla~ en Co,: Ahles & Haesloop 29296 (it TOG POY 3 Brunswick Coe: Ce Re Bell 13163 (Hi-121530); Fox & Godfrey 2823 (N, No--19606); Ae vo Ae Heller 1065 (Du--157392, ae oor 0). Buncombe Co.: Biltmore Herby 1962 Moldenke, Monograph of Verbena 319 1089b (Up—-11832). Columbus Co.: C. R. Bell 12800 (Hi—-110659). Jones Co.: Radford 3722 (Hi--10)728). New Hanover Co,: C. R. Bell 12874 (Hi—110660); Biltmore Herb. 5707 (Hi—59L68, N, N); Blon- quist 711 (H--19220), s.n. [Aug. 15, 1930] (Ob—-70515) ; Randolph & Randolph 100) (Ba); P. 0. Schallert s.n. [Wilmington, S=31<37) (tin—29556, “Sd=-25691, Tr--19773, Vt); A. W. Totten s.n. [Fort Fisher, Sept. 29, 1935] (Hi--59l,87). Northampton Go.: Fox, Boyce, & Moreland 2097 (N); Fox & Godfrey 2279 (N). Onslow Co.: Ahles & es & Haesloop 28262 (Hi--10)862). Pender Co.: Ahles & Haesloop » 28062 (Hi~10)861); Coker & Totten s.n. [July 17, 1932] (Hi—59L82) . Robeson Co.: Britt 211), (Hi--177879). Scotland Co.: Ahles & Haes- loop 28803 (Hi--10876). Union Co.: Ahles & Haesloop . 27637 (Hi-- 104877). W. Wilson 1 pees 35706 (Hi--104978). SOUTH CAROLINA: liendale Co, 3975 (Hi--92962). Banberg Co.: Ahles & Haesloop 25938 fester etree Hi--10733). Beaufort Co.: C. R. Bell a3 (Hi--927h9). Berkeley Co.: Ahles & Haesloop 2667 (Hi 04731). Calhoun Co.: Ahles & Haesloop 25540 (St), 30276 (Hi— soasea Cherokee Co,: Ahles & Haesloop 26978 ae Ae O. Me Freeman 56425 (Hi--9272h). Chester Co.: 0. M. Freeman (Hi-= 92933). Clarendon Co.: Radford 2392 (Hi--92911). Dillon Co.: Ahles & —— 27677 (Hi--104865). Florence Co.: C. R. Bell TE (Hi—-92852). Greenwood Co.: Radford 2296 (Hi--92912). Hamp- Co.: A Re Bell 2595 (Hi~—92750). Horry Co.: C. R. Bell 7710 (goss Lancaster Go: Ahles & Haesloop 27233 3 (Hi——10486h) « Cos: Radford 2h18) (Hi—92915). Lexington Co.: Radford 23392 (Hi—9291)). Marion Co.: C. Rs Bell 6409 (Hi--92916). Marlboro Cos: Radford 15482 (Hi--9293)). Newberry Co.: C. R. Bell 9820 (Hi--92853) . Orangeburg Co.: Ahles & Haesloop 25203 (Hi--104363). chland Co.: C. R. Bell 16326 (Hi——180053). Sumter Co.: 0. M. Freeman 56809 (Hi--92935). Union Co.: 0. M. Freeman 56402 (Hi— 92856) . er ari Co.: Beacons 2119 (Hi—-92917). GEORGIA: : Chatham Co.: D. E. Eyles 7339 (Sz)3 Kolthoff s.n. [27 May 1927] (S); Miller &M Maguire 1260 0 (Ua--16575). Glynn Co. : H. N. Molden- ke 118) (£, Go, H—-5h61, N N, S, Up, Ur); Pyron & ucVaugh h 142k (Gu==13995 a FLORIDA: Escambia Co.: Curtiss 690 (Ca—10L820, To—38750, N, Ur, Vt); Gander 7673 (Sd--25560); S. M. Tracy 8706 sal Tr, ae Bi ag Holmes Co.: Kral & Redfearn 2075 (Hi— 135). Sas Co.: Re K. Godfrey 53362 loaiasonees, 0, N). Taylor oe R. K. Godfrey 55761 (N). Washington Co. : F. S. Blanton 6598 (Ba, En, 4 Ny Po--215350, Um—22); O'Neill 6598 (N). ALABAMA: Escambia abe Ahles 7030 (Ur), 7158 (Ur); 0. - Blanton 114 (I). Mobile Co.: Demaree 33639 (Cb, Ss); Diener 637 (Tl); Shinners 158) (No—21011). Sumter Co.: S. B. Jones s.n. (3 July ly '60) GS 210902). Dauphin Island: S. B. Jones 602 (Hi~-210617). MS- 320 PHYTOLOGTIA Vol. 8, no. 6 SISSIPPI: Forrest Co.: F. H. Sargent s.n. [Hattiesburg, Sept. a 1932] (N). Hancock Co.: “Demaree 29590 (N), 32987 (Le). Harrison Coe: Demaree 28385 (N), 29009 (N, Ok, St), 31897 (Le). Jones Co.: W. H. Rhoades s.n. [near Laurel, Sept. 190] (N). Lee Coe: J. De Ray 5871 (Hi—199698, N); B. Shelton 85 (St). Pine Co.: Je D. Ray 543 (N). County undetermined: W. H. We He Rhoades sen. [9-190] (Bt-- 61223). LOUISIANA: Ascension Par.: Ashbey 42 (Tl), 92 (T1). As- sumption Par.: Perkins & Hall 2621 (Po—256899). East Baton Rouge Pars: C. A. Brown 1008 (N). Jackson Par.: Shinners 2016 (Ur). Jefferson Par.: H. P. Riley s.n. [Barataria, Apr. 27, 1937] (Or-- 4,063). Lafayette Par.: Perkins | & Hall 2622 (Um--25). Lafourche Par.: Ashbey 16 (Tl). Orleans Par.: L. H. Bailey 7324 (Ba); Cocks sen. (ifay 1901] (Tl), s.n. [New Orleans] (Tl); Bwan 18736 (Tl); __ Langman 19)0 (Up); H. P. P. Riley sen. [Apr. 20, 1937] (En, Se— 47522), sen. [Apr. 25, 1937] (Se--4752h); Ee ‘Wall 5 (Go), SMe [New Orleans, 10/5/31] (Ew). Plaquemines Pare: Ewan 1 1747) (Au-- 1217 = Lloya & Tracy 20 (N); Tracy & Lloyd 20 (Gm). Sai Saha Oe Ber- : Demaree 13493 (W--1699223) . Saint Charles Par.: 17707 (11). Saint John the Baptist Par.: Ashbey la (Tl), 7h “THi). Saint Martin Par.: S. W. Hutchinson 7878, in part ‘t (En). Saint Tammany Par.: T. F. Hall s.n. [April 30, 1939] (Tl). Tensas Pare: Je F. Brenckle . 40180 (N) (N). Terrebonne Par.: Arceneaux 394 (It); Ashbey 65 (Tl); Bynum, Ingram, & Jayres sen. [Houma, April 23, 1923] (N); W. H. Rhoades s.n. {Houma, August 1936] (Bt--71463, Hs, N); Wurzlow s.n. [Houma, May 29, 1914] (N, N). Vermillion Pars: Ashbey 11), in part (Tl); Constantin 46 (Hi--188555) - AR- KANSAS: Chicot Co.: Demaree 25689 (Sm). Cleveland Co.: Demaree 23329 (Sm). Saint Francis eg Demaree 15103 (Gg--267610, Os TEXAS: Hardin Co.: C. L. Lundell 1083 (N). Harris Co.: Fitz gerald 8¢ (Bl—-23L7), ac L235); W. H. Lewis 5301 Gib) Jefferson Co.: J.F. oe 48073 (N)3 | Ge be: Beg Fisher 51042 (Go) 5 Harold any 53-230 (Au—1217h3); c. L. Lundell 11136 (ND. Liberty Co.: F. W, Gould 6925 (in TA MAY; Grates Cor @ Cos: Cory 508h0 a. Sa); 6 G. L. Fisher 8132 (Go). Wharton Co.: Warnock 16420 (au, N, Or—-55399, S, Vi). OREGON: Multnomah Co.: Suks- dorf 1980 (Pl—136)01). CALIFORNIA: Amador Cos: J. T. Howell 29996 (Ge—338069), 30055 (Gg). Butte Co.: W. Anderson S.-M [Gridley, Oct. 1959] (Gg) (Gg); J. T. Howell 30519 (Gg), 34611 11 (Ge). Eldorado Co.: Crampton 2020 (Gg--368055). Nevada Co.: M. S- Ba ker 9971 (N). Sacramento Co.: Crampton 2228 (Gg--389520, Ge 369573). San Joaquin Co.: Crampton 2260 (Gg--389519, GE-— 389572), 2260a (esi). — ano CO.: mae SoM. pene. Rio Vista] (Du—-10001), s Co.: Hoover 93 (Ca 767079); J.T. 7. Howell doo12 ( (Ge--388070), sc107 732). 50697 (Gg- 1962 Moldenke, Monograph of Verbena 321 409322); Nobs & Smith 206 (Gg--1790h); Wiggins 14369 (Du 409390, N). Sutter Co.: Everett & & Balls 2390 (Hi--198835) ; Mason & Grant 13039 (Ca--192057). Yuba Co.: Crampton 133 (Gg), 3h (ce); ; < 8080 (Gg); L. S. Rose 39200 (Al, 0 (Al, Bt—51937, Ca-- 882583, I, Mi); C. B. Wolf 1437 (Ba, Du—231h26). Bouldin Island [San Joaquin Co.}: Congdon sen. (Sept. 2nd, 1903] (Ca, Ca~~138621). MEXICO: Michoac4n: Feddema 23 23 (Mi). Nayarit: Feddema 583 3 (Mi JAMAICA: Orcutt 2736 (Du-—206013). COLOMBIA: Caldas: Dryander 2751 (N). Cundinamarca: V. Grant 7352 (W—1802322). Huila: mila: Pérez Arbel4ez & Cuatrecasas 8361 (Jc). Meta: T. A. Sprague s.n. [be- tween Villavicencio and Bogot&] (K). Santander: Killip & » & Smith 17340 (N). Valle del Cauca: Killip & Garcia 3389 (W--1770719) . PERU: Cajamarca: Woytkowski 1751 (H) « ye Hammarlund 631 (S). Hudnuco: Sandeman 3486 (K). Junin: A. S. Kalenborn mborn 160 (Du-- 103207); Kalenborn & K. & ISLE 160 (N)j Ochoa 121. (N). La Liber- tad: Ochoa 1) Uhh (N); Smyt 6 (N). Lima: Ellenberg a3 (0t— 115378b). BRAZIL: a District: Harrison s.n. (Br); B. Lutz 578 (Ja--2h126), sen. [Rio de Janeiro, May 1926] pete Se n. (Ja--l6519); Moldenke & Moldenke 19591 (N). Minas Gerais: Ackermann 2831 (Br); Bai. Bailey & -& Bailey "1212 (Ba); J. F. de Castro 43 (Sp-~31291) ; P. Clausen s.n. [Aug .-April 180] (Br); Heringer Sen. [Herb. Esc. Agr. Lavras 263; Lavras, Nov. 8, 1939] (N, Sp— 26h); Irwin 2116 (N, W-—2281271); Macedo 2889 (N); Mello Bar- reto 8140 [Herb. Jard. Bot. Bello Horiz. 23376] (N); Mello Matt- 0s S.n, [Itabira ee ote mn 1902] (Ja--l6555); Mexia 5275 (Gg—286630, Go Go, I, Mi, » Vi); Regnell 1.326 [Nov. 19, a (Ja-~148)5), I1.210 rater (S), Sen. (Caldas, 186); He Rio de Jan » UBL] (N ); Sampaio 138 (Ja—6h99), 2h ( ‘a--li6500), at (Ja--6509), 006 (Ja——h6510), 6023 (Ja-—h6511); Widgren 1211 (Br), s.n. [18]5] (Lu, N, S); L. 0. Williams 5002 (G, (G, N). Frees Dusén 1 1320 (S), sen. [Herb. | Rio de Jan. 31553] (N); Hatschbach 379 (Sm), 3753 (Sm); R. B. Lange 1110 (Hatschbach al (Sm); Moure 989 [Herb. tus. Paran. 1879] 71 Os Steliteds 288 [Herb. Mus. Paran, 2538] (N). Ric de Janeiro: 1033 (Br, Br); N. J. Spe ete: (1851) (Lu, ee , Lt Bailey & & Bailey 569 (Ba); Dusén s.n. [Rio de Jan ulio] (la); Glaziou 356 (Br, Br), 162 (Br, N)3 P. P. toa a. S.n. [Ipiabas, Feb. 19),3] ered B. Lutz 1165, in part (ny, Sen. [Herb. Rio Jan. 24126] (N); Raben 521 (Br), = (Br); Sampaio 72 (Ja——l6)98), oy (Ja--l6502), 3099 (Ja—l6506), 192k (Ja—l6518); J. Vidal sen. [MaticS, March 2h--26, 193) (Ja—39278) ; Warming s.n. [Rio de Janeiro] (S)- Rio Grande do Sul: Reineck & Gzermak 63 (S); Sehnem 70 (B). Santa Catarina: 322 PHYTOLOGIA Vol. 8, no. 6 Reitz C.821 (Le); Reitz & Klein 292 (N), 1206 (Gg, Sm). SMo Paulo: Brade 5737 (Sp--6729) 5 Campos Novaes s.n. [Campinas] (Sp--1977); Campos Porto 36 (Sp--15730); A. Carvalho 190 (Ba); Costa & Viégas sen. (Herb. Inst. Agron. Est. 234] (N, Sp--31016); Edwall s.n. [Piritu uba, Oct. 30, 1893; Herb. Comm, Geogr. & Geol. 65] (Sp— 15728); Emelen 121° (Sp--28929); Freire & Azevedo s.n. [Herb. Rio Jan. 31561] (N); Hashimoto 2 (Herb. Jard. Bot. Rio Jan. 2711] ("), 285 (Sp—l2920); F. C. Hoehne s.n. [Alto de Serra, Jan. 10, 1919] (Sp—3035); W. Hoehne 22 (N); Leite 3725 (El); Lofgren s. n. [Sorocaba, May 25, 1887; Herb. Comm. Geogr. & Geol. i (N, Sp—1571h); Moldenke & Moldenke 19667 (Es, Lg, Mg, N, Ot, Sm); Pickel 14320 (Sp—I|0189), 5459 (N); Russel 27 Ba fates: Usteri S.n, (Barra Funda, Oct. 2h, 1905] (N, Sp) Venfinciio ae (Hk). State Disteinined: Freyreiss s.n. [e Brasilia ia] (S)3 Gardner 233 (N); Martius 1033 (M); Sampaio Sey (Ja—~l65 nae * 3096 (Ja—],6505) ; Sieveking eking 6 (S). BOLIVIA: Cochabamba: Kuntze s.n. (Cochabamba, III.92] (N). La Paz: Asplund li (S, Us), 812 (Us), 1259 (S, Us), 1564 (S); M. Bang 8, in part , (Pa), 136 a Ka, Pa); uessler 3 (Ca—-91),362), | 3853 (N, N), 4887 [uacbride photos 21679] (Kr—-photo, N, N-~photo), 11539 (S, W--1056970); Jorgen- sen 3767, in part [Herb. Osten 22210] (N, N, S, Ug); Morong 128 (C); Ponder s.n. [Asuncion] (Je--761h); T. Rojas 10h5 [Hassler 209; Herb. Hicken 5; Herb. Osten 7538] ( (N, Ug), “1877 (Herb. Hort. Parag. 10069; Herb. Osten 13562] (S, Ug), 1887 [Herb. Hort. Parag. 10072; Herb. Osten 13557] (Ug), 2522 [Herb. Osten 18180] (N, Ug). URUGUAY: Berro 871 (N); Cabrera 3890 (N); Gallinal, Ar agone, Bergalli, Campal, & Rosengurtt B.765 5 1/3 (m1, N); Herter s.n. [Herb, Osten es? 7] (Ug); Kuntze s.n. [Montevideo, 7/12/ 1891] (N); Osten 293 (Ug), 22921 21 (S, Ug); Rosengurtt B.865 (N), B.336h (N). CHILE: Co Colchagua: Barros Valenzuela 8047 (N). C Con- cepeion: Barros Valenzuela 805 (N); Junge 2323 (Ba); Scott Ell- iot 77 (N, Pr). Coquimbo: Barros Valenzuela 5002 (N); inet hai ¢ (N); Worth & Morrison 16360 (Ca--629836), 16498, in part (Ca-~631730). 6 Curicé: Barros Valenzuela 8051 (N), +8227 ae 8228 (1), 8229 (N). Malleco: Kunkel 27 270 (Sm). Santiago: Looser }361 (N); Moldenke & Moldenke 19760 (Es, Lg, N); Pahlman s.n. “sen. [Los Nogales, 26/1/1913] (Iu). Valdivia: Hollermayer son. [Werdermann 1362] (Gge—1h990h, Ut—-91273). Valparaiso: Behn sen, [1h Dec. 1930] (Ca—)1)6698) . Province undet Sareceee Guzman n 22 ees Mus « Nac. Hist. Nat. Chile 59934] (N). ARGENTINA: Buenos Aires: Re Alvarez 190 (S), 228 (Ga-n166257), 31 311 (S), 503 (S)5 jurelsus $+ é: a! ———— 1962 Moldenke, Monograph of Verbena 323 n. (S); Cabrera 2159 (N, S); Cannelle s.n. [Punta Lara, IV.19)3] (iv), son. [San Isidro, 10.X.19]3] (Mv); G. Dawson 865 (N), 918 (N); Floyer 18 (os ; Hauthal 649 (N); Née 115, in part rt (Q); Rodrf- guez Vaquero 17 (S); Rodrigo © 222) (N)3 Ruiz Huidobro 1473 (S); Troncoso 195 (N). Catamarca: Cabrera 1186 (N); Luna Risso 380 935 (N); Pierotti s.n, [Merced, 19/2/hh] (Ca); Villafaffe 1222 @), 1263 (N). Chaco: Ae Ge Schulz 1485 (N). Cérdoba: Belem eno 1096 (N, N); José Gutierrez 232 (N); Lo Lorentz 131 [Macbride photos 20316] (Br, Kr-—-photo, N--photo), Bel ( Bel (vt), SoM mates Ger- mania, June--Dec. 187] (Sg--68310); Osten 13110 (Ug), 13112 (Ug); Villafafie 335 (Gg—-352670), 517 (N). Corrientes: T. Meyer 6616 (Ut--72507, Ut—-115)09b); Pedersen 1911 (N, S). Entre Rfos: T. Meyer 10098 (m), 10155 (N). Jujuy: O'Donell 291) (Gg--06037, S). La R Rioja: T T. Meyer 245 (Ca—-77209). Mendoza: a: carette 3050 (N, N, Ug); G. Dawson 966 (N); E. M. Garcia 5 (N, Ok); Jensen- Haarup Son. [Santa Rosa, 1905] (S)} O'Donell “1067 (Ut--115h19b) . BOOK REVIEWS Alma L. Moldenke "The Story of Pollination", by Dr. J. D. Meeuse, illustr. by Hilda Kern, 23 pp. Ronald Press, New York. $7.50. There is mch here to delight the hearts and sige of a. of all ages and of little or much biological trai Ther careful organization of the subject matter, baited with sapkyate ing chapter titles like "Flashing eae ‘and Dashing Flyers", "Unbidden Guests Beware", and "Of Saus and Bats". feeling of having a ringside seat at a circus of fascinating nature adventures. For all the story-telling the scientific con- tent is accurate, definitely valuable, and is good leavening for one's om orientation or for one's teaching of young people from gece hades through college levels. must have direc ad his way, along i wl a ete on the city or ng snilet The ri het cent eeelle consist o al clear-cut, helpful, attractive drawings and a few fine Siekograbhs that do much to. 32h PHYTO LOE TA Vol. 8, no. 6 clarify the text. Unfortunately two labels were interchanged in figure 28, but the clear text and legends prevent any serious confusion for the readers. A fine bibliography of wide scope is iven The pollinating work of the skillful hymenopterans and lepid- r rns, is all effectively described. The "blundering beetles" codons have been belittled somewhat unfairly. These ancient pol- linat oo ere. is an interesting discussion of wind pollination as a derived rather than a primitive condition. Experiments on colors, olving in- erns, rs, sect pollinators and non-pollinators are convincingly phan Fe The price seems quite high for a book of this small size. "Tree Growth", edited by Theodore af Kozlowski, 2 pp., illustr. Ronal Press, New York. $12, This book is an outgrowth of what must have been an unusually worthwhile roup of sessions at the International Conference on i i — been well recognized for his valuable text Piste of Tre This work is definitely not to be confused with published re- cords of some other conferences. where much coverage is given to Typically there are good charts, diagrams, and pho otographs nriching the text. I was particularly impressed with the photo- graph of the "pancake" root system o: . ° V many aspects of tree growth ple discussed in concise e c axonomy, chem al influences, root grafti anatomical cassia and Specialization, and physiological problems of auxins, photoperi- odicity, photosynthesis » and mineral nutrition, The book is dedicated to the great man of American forests, PY be and it is a very worthy tribute to a very worthy st NOVELTISS IN THE FLORA OF SANTO DOMINGO (HISPANIOLA) II José de Js. Jiménez 1 Regarding the large collections of plants of the Island of Santo Domingo (Hispaniola) deposited in inchanne such as the Gray Herbarium of Harvard Daivevild $f: in Mass., the Smithsonian Insti- tution of Washington, the New York Botanical Garden, to cite but a few among the most important, three main facts — elf- ivitint to any student devoted to our Flora, namel l. New records of already known plants not mb published, as far as I am informed; 2. Many ithe ties specimens; and 3. A lot of non-identified specimens, many in tron fe waiting for a careful study or requiring further collectio both in flower and fruit to resent accurately whether hey are mre to A new records or new spec These facts represent huge ia pee ise apg Soe more adequate SeRwottene garnering more capable students, and b ing in mind sane criteria for the modern met of our ea pen A short y tion sometime pen Brg last June and July has en me the oppor- Th for a better secuhanat ete flora, the list of phanerogams existing now-a-days in the whole Island At the same time I have selected for my ge tepid a small new lot of comb. nov. because of transferring the Anastr: aphia spp. to the genus ti While bent = the ow Herbarium of Harvard University ago out that Roy Jervis had annotated many of I wrote to two years Ir the Anastraphia sina “under the genus Gochnatia. lr, Jervis asking him to inform me about his decision to publish them s I am greatly indebted to Dr. Howard and Mr. Garay of the ae Arboretum and Oakes Ames Herbarium respectively and to me to study the specimens cited in this paper and for their ef- — to make my cr sattabis stay with them a very delightful one tana ; Taxonomist of the Instituto Boténico "RAFAEL i. MOSCOSO", UNIVERSITY O07 SANTO DOMINGO, Dominican Republic. 325 326 PHITOLOGIA Vol. 8, no. 6 Orchidaceae e up-to-date and relies Schultes) "Native Orchids of Trinidad vom Tobago", pase 93, the geographic a of Hexisea reflexa (Lindl. ) Red dhs f., he cites Hispaniola among the countries where it has been record A careful search in the botanical Wéeratuss a with Hispaniolan floras was entirely negative. So ural ques- tion is where did Schultes find this "unrecorded" rec priser ing my last visit to the Oakes Ames Herbarium of Harvard oe June 5-8, Dr. Garay and I found the correct answer. c re to two different tenorde in the Dominican Republic and Haiti, viz: DOMINICAN REPUBLIC: La Cumbre, in forest, Cordillera ones ral, Santo Domingo Proviine: c. 300 m, 5. Ve. 1929, Ekman H 12374. Herbarium Oakes Ames No. 45126. Det. Mansf., 1950+ s On sures are odes , Riviére Glace, 2200 feet, May 7, 1949; J. = Curtis E-20. Herb. Oakes Ames Ho. 64210. Det. Schweinf ur ae ect A. Garay in his monumental work "Venezuelan Or- ", page 372, 1959, prefers the binomial Reichenbachanthus vaca (Lindl. + ie Brade.e SPIFANTHES CALCARATA (Sw.) Jiménez, comb. Neottia calcarata Sw. Flor. Ind. Occ. 3: "7413 to. 28 fige Ce 1806. SPIRANTHES LAXIFLORA (El, &: a ) Jiménez, comb. epgponen 3 laxiflorum Ekm. & Mansf. in Ark. foer Bots 22A6 Compos itae During one of my ale oi trips to the capital I noticed among ithe trimmed grasses of the lavms of different Vv rostrate residences te ne ry common p Compositae with small yellow heads enti liy oat osity became aroused. I picked up several spec at home examined them very thorou chly. There was no ; ona but that rnd Lae cen ep to the tribe of the Helianthea prise, none of the genera of our epaihs could ade- eveay natch my little plants. They wore then sent er the Division of Phanerogams of the Snitheonian Insti SpE for study by Emery Leonard and Dr. le Be who found them to be the Mexican Blainvillea tempicana ( Less. ) —~ & Jackson ‘ - dn. Flore, Sertes II, part 2, page 4 — ere Johnston ae exander includes this name in the synonymy Ree Sal wd RIVER win ‘ANDERSON SHELBY oMTOCME! TACORDOCRES © SABINE : HOUSTON + sapine BS VERNON ANGELINA NEWTONS uy POLK, JASPER RAPIDE! “er THUR fe Graves | WALKER eee ear CALCASIEN moc) a COME} CAMERON HAMBERS FORT BEND eu ta BRAZORIA TAGOROA TEFTERSON | WAUKESHA 1 H 5 . s 3\"" ‘COOK BLACK HAWK ‘BENTON | LINN ror TUADISOR| WARREN | MARION | MAHASKA] KEORUK } "ATI? WOMROE | WAPELLO [ e7FERSON | HENRY FAYETTE | crvimcnans] ASPER hw se wee x Le) Sua Tone, Z x C o ~ 5 = A w Hy 3 i) = < OSHA | a wwe a) coo wi. a re Ff s FORD aa i) s PUTT) |S DOUGLAS) . cues SHELBY og orca fousren |_| to" Carne RARDIN veasTeR cry TEBE i mots ATTA a WINSTON. LEAKE i.) ESHOsA KEMPER ‘corr NEWTON | LAUDE! Tr a 8 a NE GREENE saan CLEARTYPE PHYTOLOGIA is financed entirely by its contributors, each one Hee: in advance for the entire cost of printing, binding, and distributing his tribution. All money received from subscribers, after the expenses of ot lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. ach number consists of not less than 32 pages. All ae accepted be will be published in the next issue, so that the size of numbers may vary Teatly. me will contain about 32 signatures, 512 pages, or a smaller number of with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. nergy sine be ape = to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in zinc, or for pie tables, or Enh provided they conform to certain limitations of size and proportion. An extra charge will be made for half- tones, depending on their size, as fixed by the engraver, with a minimum of about $2.25. Articles dealing with research in all lines of botany, in any reasonable length, biographical sketches, and critical reviews and summaries 2 liter- ature will be considered for publication. Floristic lists, casual notes of an amateur or so-called popular type, and polemics will not be eI", Advice Hae oi of manuscripts will be solicited, if necessary, from qua uali- as d the ‘Present cost 0 t of printing, the basic rate for a page or for an edition of 250 copies. This price is subject to change without r ieeice, since it depends entirely on the prices prevailing — ee in the printing eee: "Reprints will be furnished at cost. A proportionate fraction of the ca i of 250 ole eS eee gratis to contributors. se Upon request, the editors will send detailed instructions concerning the iis may be addressed. to ne € magazine ¢ or = either editor. PHY FOLOGIiA Designed to expedite botanical publication — Vol. 8 June, 1962 No. 7 : CONTENTS | BARKLEY, F. A., Anacardiaceae: Rhoideae: Lithraea...... PONE ico 329 LAUGHLIN, K., Common names for Midwestern forest trees ..++++++ 366 ALAIN, Bro., Novelties in the Cuban flora, XIV..-.+++++++++ Means 30 MOLDENKE, H. N., Materials toward a monograph of the genus Verena VT US nd ee eee ce ee et Oe ee 371 URI BOTA Ne RDen speeniabied by Harold N. Moldenke and ‘Alma L. —— ae Glenbrook Avenue Yonkers eee New York, U. S. ee | Price of pope $1; per volume, $5.75 in advance ANACARDIACEAE: Rhoideae: Lithraea* Fred A. Barkley Of the ee 38 ia of the tribe Rhoideae et are four which have stamens twice many as the petals, the tricar- ce ae tanons and ovary: Lithraea, Mauria, Schinus and Pachycormus. ea differs from the other three in that it has basal seed and ee leaves have ne mar The first entity of this genus to be known was apparent] described by Molina as Laurus caustica in his Saggio Storia Nat- urale del Chile 176, in 1782 as "Taurus fol. ovat tbus rugosis peesuce antibus, flor. quadrifidis". Vellozo described an entity from Brazil in 1825 as Schinus molleoides and this was followed by the de escription 4 n 1833 of Schinus ternifolius Gillies ex Hooker & Arnott, two entities variously treated, but historical- ly mostly Hct as Synonymous. 1826 Miers published Lithraea venenosa in Trav. in Chile 529, thereby establishing the genus Lithraea, by which name ase em have been consistently known Since that time. genus is represented by southern Brazil, Paraguay, Bol- ivia, ieceny, Argentina ar Chile by a erked of polygamo-dioce- cious poisonous-sapped tree LUIERABA Miers Miers, Trav. in Chile, 2: 529, 1826. Endl., Gen. 1130. 1840, aed Rep. 1: 551. 1842. Lindl., Veg. King. 467. 1847. Blume, Mus. Bot. Lugd. 1: 15, 1850. : Bentham 2 | Hooker, Gen.Plant. 1: 418. 1862. larchand, Rev. Anacard. 9, 182. 1869+ Engler in Martius & Eichler, Fl. Brasil. 127: 394. 1876; in de Wat. Ee Monogr. Phaner. 4: 345. 1883; in Engler und Prantl, cttanzenfam. 35;°164. 1892 a cumén. The writer is agen to the rector, Dr. H. R. Descole, © the 1 ° 1 s in the 3 330 Parto.ogi.a Vol. 8, no. 7 Cabrera, Rev. Museo La Plata n.s. 2: 35. 19385 Rev. Argent. Ag- ron. és Pome tb Sate pee Os Flora Chilena 2: 43. 186. » Bot . Mise «at 275« 183). with ER sap and slender branches. Leaves alt- ernate, spidnnatea, simpl rnate or pinnately 5-foliolate, the c r alate rachis and often alate petiole. Panicles usually lateral ee clustered in the axils o p most leaves, rarely e bracts deltoid and caducous.e termi Pedicels usually nha Kove the abscission plate. Calyx usu- ally with 5 semicircular or broadly deltoid lobes, glabrous ex- cept the margin of some ciliate. abs _— 5, valvate, lanceolate, acutish, dt The s ns twice as many as the calyx lobes, in stamina e flowers siieeely: shorter than the petals, many lobes as the stamens. ‘The pistil in pistillate flowers with three hemispherical stigmas, one style, and a subspherical ovary with one cavity and one basal ovule, the staminate flowers have much smaller and abortive pistil with two points to each stigma. The fruit has whitish epicarp readily separating from the dark resinous mesocarp, the mesocerp adhering to the bony, smooth to rough endocarp. @ species: Lithraea venenosa Miers, Trav. in Chile as 529. 1826 (- Lithraea causti Ole) Hook. & Arn two red problems which confronted the present author were the specimens of the mountains of northwestern Argentina mens it has seemed better to segregate the Bolivian-Argentinian Series of tall trees with large fruits as specifically distinct ith small rom the similar but smaller pied he er fruits from the lower tropical regio Tiis m nous species 6 first de- scribed fe Gillies so publdahed by ser pad a as Schinus - three iisvinet Bey one leaved species L. brasiliensis March. only mor or less S aaencls from the one or two pinnately compound leafed species. ‘The pinnately compound-leafed species, Lithraea molleoides (Vell. ) tu This is common in southern Brazil. In northeastern Argentina is & plant with linear-lanceolate acuminate leaflets, ee ae mus named Lithraea Lorentziana. I+ has been included i raea molleoides by most authors and is mited to that et “tet by ® large series of intermediate specimens. To the author ee ee ee OD 2 ck TOT eee ce oa Ue ge ew Gepem eae ce are 1962 Barkley, Lithraea 331 or cytogenetic studies may show that they are aa variations . a single species. At present it seems to the writer best to treat them as two species and one variety, ahenctiol Lithraea brasiliensis is scarcely better differentiated from ace molleoides than Lithraea molleoides var. Lorentziana. KEY TO THE TAXA 1 Leaves simple or mostly soe Leaves ovate, heavily coriaceous, the margins entire 0: slightly irregular, inclined to be scalloped; fruit ‘avis Chile. 3 Leaves and branches glabrous, leaves often larger and more coriaceous, the margin more Wee ae LITHRAR CAUSTICA 3 Leaves and branches pilose, leaves usually smaller and oped slightly thinner and less scollo 4a. LITHRABA CAUSTICA var.PILOSA 2 Leaves oblong or ovate-lanceolate, the margin entire or ir- r regular, rarely scolloped; fruit small: Brazil and Argentina. LITHRAEA BRASILIENSIS ATHRASA Siiol 1 Leaves compound or mostly s0o« 2 Fruit small, 4.5-5.5 mm. in diameter; lowlands of north- astern Argentina to Brazil. 3 Leaflets usually ovate-lanceolate, usually obtuse but often acute al LITHRAEA MOLLEOIDES 3 Leaflets usually linear-lanceolate, acuminate. 2a. LITHRAEA y MOLLEOIDES var. LORENTZTANA 2 Fruit Borat 7-9 mm. in diameter; highlands of western Argen- tina and Boli 3, LITHRASA TERNIFOLIA 1. Lithraea brasiliensis March., Rev. Anacard. 185, 1869.6 = yenulosa Spreng. ex Engl. in Mart. & Eichl., Flora Brasil. in synon. pe australiensis Engl., Engl. Bot. Jahrb. $ “ah 1881; Engl., in de Candolle Deas teeea ~ ks ShT. 3, in syn. Becta verrucosa Miers ex Engl. a de te Monogr. Phaner. rgreen tree f 7 or 8 m.s branches slender, glabrous or sparsely puberulent, iid brown; the leaves simple, or leaves 332 PayT 02 60:2 Yol. 8, no. 7 in part with leaflets 2, 3 or 5, leaves thinly coriaceous, the margin white, entire or somewhat irregular, glabrous or ve sparsely puberulent on the veins, obtuse at apex or rarely sub- acute, cuneate at base and decurrent on the 4 10 mm petiole, long, to 3 -« broad, the compound leaves 4 to 9 cm with leaflets sessile, similar to the leaves, the petiole and pat ous, deltoid, often one larger (5X7 mm.) and two smaller bracts (2 X 4 m.) clustered together, glabrous except for a few cilia on the margins; pedicels about 1.5 mm. long, sparsely pilose- puberulent below, glabrous and shining above, in fruit to 3 mm. long; sepals 5, rounded-deltoid, glabrous, margin more or less irregular and with very few cilia, 1 X 1 mm; petals glabrous, th stigmas 3, flaccid, .2 mm. in diameter; the pistillate flowers with petals .7 X 1.8 m., stamens with anthers sterile, oD Ime. filaments slender .5 mm. long, the disk 1.2 broad, less lobed than in the staminate flowers, the three stigmas large, thestyle .4 m. long and the ovary sph dia meter; fruit 4 to 5 mm. in diameter, slightly flattened, sty persistent and brownish, epicarp creamy yellow, r separat- ing from the thin resinous mesocarp; seed joneeenet slightly larger at one end, 4 mm. wide, 5 mm. long, 2.2 mm. thick, essen- tially smooth, brownish black Saud’ cheud 925 from Sao Paulo, Brazil, in the Paris Herbarium © range ms i. species is from IMinas Geraes, Brazil to wssiones, in ataoag "Aruera" "S,litre", “Iaurel del Monte", role, og “beber", "Arbol malo", "Litre", "Aroeira Braba", "Pan e Bugre", "B ratoa", eeetunks examined BRAZIL: ae me 18199 (K); be se rerio ark SNe (u) and 1820, M); Riedel sen. ane S oNe S ones j Setters Bays (si); DINAS GeRARS jee rrisire 1762 (RT BM J. EO: Cape Fri p ~ 3; Cabo Frio, ; Cape =v BR : Sept eh, Bi; Glastou 15947 (F, Us) and 12547 (R), and 1882, Glaziou Se remerraes 3 Ca o, 18 zt 0, Septe 1881, Netto, Glaziou y Schwacke S-¢ . trouch in der Restinga bei Cabo Frio, Octe 1899, Ule 4714 (corp, R); SAO os Sorocaba a za deiro Folios, es: Curityba, 25 Nov. Mee *pusen 2229 (2), 1908, 6849 (S perussu, 18 Nov. 1908, 7144 r ur (S, US), Pinchaea, 13 Noy -TSTL, 13353 (F, S), and Rio Capinsal, 29 Febr. 1916, Dusen 17883 (S); Sao bea 26 Dec. 1929, Gunzel Sen. (RB) ; Nove 1936, 16102 (2B); Curityba, Bstacao me guys ct. 1946, Hatschbach 421 PB) , 100 oe Jonsson 1ll2e (F, , Wich, NY, S); SANA CATARINA: Herve Jinto 1011 Dusen sen, (S); 1837, Caudichaud 269 (F, GC, v8) tyba, 19 Oct. 1909, 8740 1962 Barkley, Lithraea 333 Esprsanopolio, 16 Sept. 1145, Joao y Rohr 504 (LIL); Somprio, 23 Nove aan Reitz c708 (RB) and Sombrio, 10 m alt., 19 Sept. ae C1227 (LIL); RIO GRANDE DO SUL: Miamao, Campo do Varejao, Oct. 1945, Emrich 1167 (LI L); Sao Joao do Montenegro, Pareci- al Oct. 1944, Eu ae ee (LIL); = Joao do Montenegro, Pareci=Novo, 16 Oct. Henz y Rambo 29655 (LIL); Sao Leopoldo, 20 me alt., 10 Oct. aa Eugenio 35403 (LIL, S) ics Oct. 3003 (GH); Porto Alegre, 3 Nove 1892, Lindman a581 (S); Rio pt 29 Octe 1901, Malme 63 (S), 30 Gct. 1901, 86 (S), Porto Alegre, 16 Dec. 1901, 860 (S) and Porto Alegre, 5 Nov. 1892, Malme 258 (S, US); Norse Sapueais, 29 deni 1948, Bo acios & Cueto Man (LIL Porto Alegre, Morro da Policia, 2 Febr. 6 steza, 3 Febr. 1948, 698 (LIL) and Porto huis: tristeza, 3 Febr. 1948, Palacios & russes 740 on » U)s Cai, Morro Cauastra, 16 Mar. 1933, o 1720 o da Gloria, 16. Oct. 1945, 29302 (LIL, S), nresa, ot Oct. 1861, 51333 (US) and 10 Sept. 27, Bese. (Us), Porto Alegre, Morro da Gloria, Octe 1944, 27097 (LIL), Sa Leopoldo, Sapucaia, 5 det -1945, Rambo 29486 (LIL), Sao Franses de Paula, Vila Oliva, 8 a 1946, Slloe (LIL), Aprados da Serra, Bom Jesus, 1000 m. alt. nero 1947, 35008 (S), Bom Jesus, Faz. Belem Velho, Jan.1947, Seanes (LIL), Porto Alegre, 2 Oct. 1948, 37793 (LIL), Garroupilha, 29 Jan. 199, 0338 (LIL) and Porto Ele- gre, 30 Sept. 1949, 3667 (LIL), Sao Leopoldo, 17 Oct. nica 43915 ), Lagoa dos Barros, Osorio, 2h Nov. 194, 4509 (LIL) and Fur- na do Tigre, Itapoan, 19 Nov. 199, Rambo L500 TL LIL); Bélem Velho, Oct. s02s peneck & Czermak Si) a Se al da Floria y Beis em Velh a ; Ee Thering 1 (R); _Arochavelota sen. (CORD); Berro 883 (G, fad a Feb. j SEROLERED: “torre ™ 1926, 1892, Trscheva Teta Sen (CORD); CO de las Cue tas, 25 Febr. 1988, Rose gorct 32583 ) and eee (LIL), a at 38 Dec. 1938, B-2796 “ Palma, Nov. 1934, “sae 1537c (US); Arroyo vansavd Nagra maps Castro: Rincon de Santa slena, “ov. 1946, Rosenzurtt & Gallinal 5744 (LIL, S$, US); ALLEJA: Sierra de Wins, Wovs, Arochevalsta s. 1 eli Minas: Jena 200 m. alt., 1-2 Nov. I See (F, Ge U); MALDONADO: Piria- polis: Cerro del Toro, 10 Febr. 1933, Clos 4971 (BAB ; Cerro de S, 7 May 1936, Legrand 809 (F); *Sferra de Animas: Nov 1922, Hauman sen lt Sept eNe 3; Cerro de las Ss, 500 me alte, vept. 1954, Rosa 526 (BA, LIL); Punta Ballena: Bosque Lus Febr. 1943, Posengurtt B-4520 (LIL): ROCHA: an Vicente: 2 8 Jan, 1921 1, “store BESTS EL HS: Tacuari, Nove 1933, Herter on Ale Ge. U splnpak UC and 1537 San } = ‘+ ees Niederlein 713 (1591) on x Fl Lithraca molleoides (Vell.) Engl. in pantie’ = aes. 122; aga. t.83. 1887; Cabr., Rev. 'useo de la Plata “Se 2: 38, 938, DDE. Schinus mollecides Vellozo, Florae Fluminensis 10: pl. 154. 334 PHYTOLOGIA Vol. 8, no. 7 1825. Rhus clausseniana Turcz., Bull. Soc. Nat. Mosc. 1: 469+ 1858, fide Engl. in DC. Monogr. Phanor. 4: 346. 1885. Lithraea Aroeirinha March. ex Warm.in Kjoeb. Vidensk. Meddel. 6l. 873. q Schinus liolle Hort. ex Engler in DC., Monogr. Phan. 4:346. 883, Ss aos ee Mart. ex Engl. in Mart. & Eichl. FleBras. 122, 394. te cole Pepe Mercheex Cabr., Rev.Museo de la Plata nS. 2: 41, ae wus 5-5 meters tall with slender branches at first tan, finely and (usually) densely puberulent, later glabrate and creamy- tan; leaves ternately compound, or rarely 5-foliolate or very rare~ ly some Nasheg leaflets sessile, thinly coriaceous, terminal usu- ally slightly ery es cme long, 1+8-2.2 om. broad, lateral 5-525 cm. long, 1.5 - broad, ovate to ovate-lanceolate, mucro- nate, typically catty obiuee, but often subacute to acuminate, base broadly to narrowly cuneate,entire to sli the branch Shortly pilose, 5-6 om. long; bracts caducous, broadly wade Sparsely pubescent on outer surface, spars n6 eset with simp hairs; pedicels 1 mm. long, increasing to 2 fruit, en te cent below, glabrous above; ca calyx lobes very paca deltoid, 1 mm. broad, 5 mm. high, glabrous except margin sparsely ciliate with simple hairs; petals tet vate, 1.8 mm. long, »8 mm. broad, acutish, glabrous; stamens in the staminate flowers with filaments .8 m. long, anthers ehIbNE, -8 mm. long, in the pistil- late flowers much smaller 3; disk about .8 mm. broad, lobes as many 08 t smaller; fruit 5 mm. long, 5.5 mn. broad, 4 mn, thick, epicarP creamish, readily separating from the dark resinous mesocarp, the mesocarp adhering to the bony endocarp; pec eed 3.5 mm. high, 2mm. thick, and 4.5 mn, long, br s JXPE: Warmin cpt from Minas tenes; ies Common names: "Aroeira branca", Aroeira brava", ie branca", "Aroeira miuda", epshnd ry "Aroerinha", “Anieira" , Aroeira" . "Chichita", "Yolle Guazu" ange: from l'inas Geraes, Brazil, to kisiones, Argentina. 10486" cimens nal LL: Cuming sen. (UPS) G a an n Jose dos Campos, 8 Sept apa 339 plete: “Telncto, 1837, a3 4 Gn 25 Jan. 1859, fania, alt. 700me» eat 1962 Barkley, Lithraea 335 1839, sen. {s), 2 3 (G) and 1841, Claussen 8 (NY); Jul. 1842, Gardner 49 7 (G)3; Grei de Barbacena, 25 Jun. ni i Glaziou 2 fe US); Prata, Triangulo Mineiro Tabouri au sen. (RB); Caldas, Sept.1854, Li indbors “zi (s)5 Bello coke aad Villa Novo Horizonte, 25 Jul. 750 (F. R.)3 Bello Horizonte, Engenho Nogueira, my Fe 1942, Vagalhaes 3155 (BEL); Caldas, 20 Sept.1873, Mosen 338 (UPS, S) Sept. 1862, Netto sen. (R); Caldas, 29 Feb. 1843, Regnell 63 (S, UPS, US), Caldas, Rens 1846, II 43 (vrs), ees 1847, Il 63 (HY, UPS), Caldas, 5 Oct.1858, s.n. (US) and II 63 (S), July 1848, II 63 (NY, UPS), nthe Il 66 (R), 128 oe spt. 1874, II 63 (UPS), and 1875, Regnell 63 (UPS); 1822-1828, Riese) Ser (S, US); Rio das Vehas, 1 Sect. Schwacke sen. (R); oa Santa, 1 Sept. aan Warming s.n. (S--IOSTYPE); 1645, Wideren ae 5 (BR, U, UPS) and Caldas, “Tas, 1082 (S); ; Sena tle te le de Belo Hori- zonte, Pe das Pedras, 1000 malt., 4-5 Sept. 1945 eo ohne & Azzis 7481 (LIL, US); SAO PAULO: Botucatu, Nov. SNe and ane Feio, 15 lov. O¥s tees 159 (SP); 18 gi “eT Fandro 73 (PF, and Sone tt pbs - iene ra, Joao Jorge, 25 Aug. 1904, iotge r 161 (S); Buta Aug. 1917, se nn (SP), 23 N ~“TS17, 942 (SP, ie fe is Sept.1931, (F); ake 19 Oct. et 9, ene Franco sen. (SP); "errs ~ Cantereira, Koscins tapetininga, 28 Nov. 1943, Lima s.n. (RB); Sedo 22 Sept. 1887, Lofgren s.n. (SP) and San Jo se dos Campos, 8 Sept. 1909, 359 (EB); Taubate a Sao 2 do Parahytinga, Sept. 1892, Lofgren y "dwall s.n. Ypiranga, ae 1913, Luederwaldt S.n. (SP); 20 Jun Mosen 5855 (S); Helvetia, 6 Dic. 1941, Pickel 5547 “cs)s" Sellow 3 Campinas, 8 Sept. 1873, Severin and 12 Sept. 1873, 60 (UPS); Campo Lim po, 13 Sept. 1934, Saith 2 (SP); Oct. 1912, Tamandare 125 (RB); Capital, Avenida Paulista, 18 Nov. 1906, Usteri s.n. (SP); Jan.-Febr., Vecchi s.ns (R); Loreto, 1909, Vooeh See (SP); Campinas, Joaquim Bgidio, 27 Aug. 1939, Viegas s.n. (SP); PARANA: Ponta Grossa, 12 Dec. 1903, Dusen = x 2 3 Ponta Grossa, 18 Marzo 1904, s.ne (S) and cal rossa y Acuariahyua, 26 Sept. 1911, Dusen 13059 (S); Palmyra, & » Nello 5262 (F, R) an Inzias Capeiras Baccaria, 19 Sept. 9206 (F)s oe gion 5 Jan. 1902, Malme 928 (s) and iiss (8); Pinhal, 6 ‘ar. 1948, (LI Palacios uezzo 2090 (LIL); pu as Parana, 1909-1910, Fiebrig 5666 (L, LP GUIARA: Villarica, Jan.1905, Hassler 8729 (G); Canon Villa eT 0 Oots 1928, Jorcensen 3428 (BAB, SABosq, LIL, LP, ny, US) and Sept.1S31, Bree, S)s EAACURL: oe Sept. 1941, Rojas 915T (LIL); ‘O: Sierra de Ma yu, San = séanss te o, Hassler 41592 0% OC); YHU: Nov. 1874, ai go28 (4,8). aclrt oes b 336 PAZTtALO GIA Vol. 8, no. 7 eGUAd: ie ieecheraiel o9 (G) and 191 (Cc); CaRRO Lagcos oO Necr Palle — 1936, RoSengurt tt A705 (us); DURAZI °O0: rincon 5 pens ners a rio Negro, Dec. aru: Lerrand 350 @)s FLORIDA: Rio Yi: ca del Arroyo Timote, 2 Nov. 1046, C stella = paces eee curl) "Rincon ee. Sante Blena, 21 one Sent — cetiine nf love (vt Eason me A178 (LiL) and 93-4178 “CLIL); RIVERA RIVERA: Galgo, 240-300 mealt., 2-4 neces 1933, Herter O77a (F, G, U, U, RB, Cc US); TACUARELEO: Oct. 1895, Arechavaleta 279 (G)3 Valle iden, aoe Seas yas tel tance iets MISTOUSS ¢ THGUAS : Puts Rico, 12 Septe 194% (LI LTL) and hes Encantado, 230 m alte, i Jul. ort a (LIL); CANDeLARIA: Santa Ana, 8 Sept. 1912, Rodri- guez 556 = (Fac CBA). Smee 2ae lLithraea mollooides var. Lorentz ziana Hieron. exe Lillo, Contre Conoce Arbe Arm. 2 1910 . Arbor 5 m- alta; foliis- : foliola ee a ahopolatis: acuminatis; drupis 4.5= -5 mn, in diametro Lithraea Lorentziana Hieron. ex Niederl., Bot. Mens. ot. seg ee: 3 292 . 18925 ex yiederl., Result. Lithraea chichita Speg. in Speg. & Girola, Cat. Descr. Maderas . . Re thraea molleoides Engl. ex Cabrera, Rev. Museo de La Plata neS. ° 1933, pr. vi oar: rah o 5 meters with slender glabrous branches gray to a ae the leaves alternate, glabrous, 5-foliolate or ere 3-foliolate (very rarely with a simple leaf); leaflets about equal, 4-7 cm. lon, 6-10 mm. broad, linear-lanceolate, acute to acuminate at the apex, narr a ip am at the base, ee very sparsely and shortly pilose; bracts caducous, eltoid, ntti except ciliate on the margins, .5 mm. broad, ye = long, usually associated with two caducous beasnectes Ey Brag pedicels about 1 mm.long in flower, increasing to ficee, pat ye sparsely pilose below, enlarged and os wide 5 yx lobes deltoid-semicircular, glabrous, « ice, 5 mnelong; stamens usually 10, in the staminate apy oee t 1 mme broad, lobes as many as the it age stigmes 3, style 1, ovary 1 +t : ty eee and one seeded n the pistillate Powers about a ie: much smaller in sm paph flowers; fruit 5 mm. long, whites 4mm. thick, with smooth amish or preapenrp te ad, mm. thi carp eptenrp which readily separates from the resinous meson ; F) © mesocarp adhering to the endocarp;. seed tan, smooth Mme long, 4 mm, broad, 2 m. thick ea Neeeme icicles iil tect 1962 Barkley, Lithraea 337 TYPE: Niederlein 1590, Esteros y Paso de Ibera, ¢ October sui in the Herbarium of cme rene ore Museum in Buenos Aires. "Chich "Chicha", "Aruera", i. melo" 2 ‘Molle: eaat Vole ais", ; “hyotte®, “Corazon de bugre The areuiee ranges from xouleains Brazil, through Uragusy and Paraguay, to northeastern Argentina. Specimens examined:= BRAZIL: RIO GRANDE DO SUL: Dowie estancia do Jarau, 26 Dec. “1944, Rambo 26310 (LIL), Jan. 1945, fe a )s n valle fluminis Y-acas Valenzuela, Jan. 1900, Ss 6945 (Oh 3; Alto Parana, pe 3 ig 6666 (LIL); ENCARNACION: rana, Jan. 1943, Jimenez 10 3 Sept.1915, Hassler 1368 (G); Jan. 1944, Ro ioe 108 a bir ); 28 Jan. 1907, Spegaz— Zini sen. (BAB); 10 so arse nastier 3; Villarica, il 1941, ‘ojas 8 sae 7 (LP) and Sept. i DIST (LP); YHUs Cosme, entre V rcuae: Telanse 2325 (BAF); + ARTIGAS: Santa Rosa Guareim, Nov. 1927, Herter 977, (, G, LIL, NY, S, U); CERRO LARGO: Rio Negro, Palleros, Tan. 1936, Gallinal, Ara — Bergalli, Campal & Rose sat a an erter 16444 )s to eG y A° Palleros, if wke. 1936, Eeeens tt 1126 $i) ; : Minas, Argua, Mar. 1920, Sch au? Chapicuy, A° Guariyu, Kilda, a tt B-4919 (LIL); ARGENTINA: MISTONES: Denis s.n- Ry Gi Big Sen. Azara, 14 = oe 291 (LIL) and 31 Jan. 1945, CAINGUASs Puesto Mineral, 250 m. alt., 23 Aug. 1950, Schwarz 10654 (LIL); Puerto Rico, 5 Mar. 1948, Schwindt - (TIL); CANDELARIA Loreto, 8 Dec, 1947, Cosso entre Garupa y Candelaria, 9 Oct. 1943, Gavio s.n- (BA); septs 1939, Goytia Sen. (BAB); Estacion Experimental de Loreto, 9 Mar. 1950, Gruner 85 (LIL); Bonpland, wa 1910, Jorgensen = (as) Santa 2 Dece 1904, 723 (BAB) a nta (BAB, G)s Loreto, 220 m, Eas 53999 (BA), rebebyri, "5 Tori 1945, 796, ia, LIL), Loreto, 218 m-, 14 Jul. 1945, 1000 (LID, Santa oy 2, Roart z 556 (LIL, NY; SI), and Seats om 3 bea ois” 12574 7 Santa Ana, 4 Feb. 1907, Spegazzini s.n. (BAB, LP), Sante Ana, 21 Jun.1909, Venturi 96 (LI $ Posadas, Zarman, 27 Aug.1945, Bertoni 1626 (LIL); Fosadas, La Granja, 5 Dec. 1907 7-(S); Posadas, Oct.l Gallardo 3739 (EA); eee Febr.1944, Meyer 5427 (LIL) ‘nd Arroyo Vera, 24 Jan. 1947, Meyer 11400 3 Posadas, Jan. 19 907, Ne EOS 1349 (LP); got par S Febr. 1948, Schulz 6916 (LIL); IGUAZU: Salto nua April 338 PHYTOLOGIA Vol. 8, no. 7 Rodriguez anata {L0L); Bl Dorado, 160 m, 26 Sept.1949, Schwindt ACIO: Ihaca-guazu, 11 Sept. mae? Bertoni 1876 (Cc, LIL); 1 Febr. 1945, Descole 3275 (LIL); San Ignacio, ove 1922, cals 156 (BAB) and I56B (BAB); San Ignacio, 27 Jan. 1947, Meyer 11604 Segre sibs Teyuanare, 222 m., 5 April 1946, Montes 2107 (LIL), San Ignacio, 4 Febr.1947, sn. ( and 8 Febr. 1947, s.n. (BA); i. Sept. 1913, ie @ Sene (BA); camino a la Plantadora, Jul. 1927, Scala s. ne ore Gober= nador Roca, 8 Sept. 1945, Schwarz 12 Balsa Nancan- guazu, 31 Aug. 1945, 1188 (LIL), Gisela, 14 Sept. 1946, 3374 (LIL), Santo Pipo, 3 Oct. 1947, 4873 (LIL), Nancanguazu Oct. 1947, 4994 (LIL, U), Gobernador Roca, 13 Nov. 1947, 5158 (LIL), Santo Pipo, 27 Sept.1948, 6175 (LIL) and Santo Pipo, 4 Oct. 1948, Schwarz 6281 (LIL); SAN JAVIER: Concepcion de la Sierra, 1 Febr.1945, Sesmero 360 (LIL); SAN PEDRO: San Pedro, 4 Nov. 1886, Niederlein 718 (81); El Alcazar, 1 April 1949, Schwindt 1449 (LIL); Campo Cumprido, 22 Sept. 1945, Bertoni 2102 3 s Esteros y Paso de Ibera, 7 Oct. Tes6, Niederlein 1590 (BA, SI) and Esteros y teed de ta era, 7 Octe 1886, 719 (BA); BERN ASTRADA: Dern de Astrada, 20 Oct. 1949, Schwarz 6358 58 (LIL) and 8361 (LIL) CONCEPCION: Loma Alta, 17 Mey 1944, Tbarrola 540 (LIL) and 2 9 Aug. 1945, Ibarrola 3191 S)s ESQUINES | (F, LIL rranca rio Parana: Puerto Mercedes, 24 Jul. 1922 (BAB); rana: Puerto Mercedes, 24 Aug. 1922, Ro as 4337 (BAF); GENERAL PAZ: Lomas Vallejos, N r 14 Nov. 1949, Schwarz 8707 (LIL); 8 AN LUIS: San Luis Palmer, Cerrudo Gue, 5 Oct.1945, Ibarrola 3434 Aa sit S), or MARTIN: tres Cerros, 8 Nov. 1036, Burkart (LP ar Sc 8094 (st); Yapeyu, 2 Jen. TSES— Toarrola sie el IGU reto, 8 Sept. 1944, Ibarro ll Santo Tome, 26 Nov. 1947, Cozzo BA); 9 Fobr. “1910, Venturi 444 (LIL); Santo tome: 25 Nov. 1944, Barre 142 (LIL); TTATI: Itat ti, 20 Jul. 1930, Fonseca Sen. pe URUGUAY: Salto Grande, 5 Pe Eee Taste Lancs Sle (8A); Concepcion del Uruguay, Dec. 1917, Hauman ssn. (BA, LP); Concepcion = Uruguay, Oct.1877, Lorenta (UPS) and ont 1878, Bee ae D), 1665 (BAF, CORD, K), * and IG65E (L); iClA: Fontana, Nov.1935, rer 2014 (LIL); — sare 343 mae aa Schulz 34 (BAB ) ontana, May 1928, Schulz olig (Gillies) Barkley, Rom. n. combe Schi pepe Toye mbfolia Gillies ex Hook.s Arn., Bot. Miscs 3: 177. Schinus tenuifolius Steud., Nom. ed, 2. 2: 530.1841 Lithraea Gilliesii Griseb., Goett. Abh. 19: 114, 1874. ene ee ; 1962 Barkley, Lithraea 339 Schinus Terebinthifolius var. ternifolius Ma roche, Reve Anac. Tree to seven meter rs, branches tannish gray, at wit ly compound (or rarely the first leaf on a branch simp ee elabroue ‘; alternate; leaflets 3-5, thinly coriaceous, ovate-lanceolate, +4 mm. broad (usually associated with 2 similar but smaller bracteoles), glabrous except with a few sessile glandular cilia on the margins, deltoid; pedicels about 1 mm. long or in fruit to 4 mm. long, at first sparsely pilose but becoming glabrate; the calyx with 4 to 5 glabrous lobes 1 mm. broad and .5 mm. long, with margins bearing a few sessile and glandular trich- mm. mm. long, 6 mm. broad, 2.8 mm ck. 8 wie e Probably exists in the Kew Herbarium a spec met, t been seen by the present writer, nor any direct “citations. a it n names: "olle de beber", "Molle blanco", "Molle duloe", Aaya de Cordoba", "Molle", "Palo Negro", "Molle de bebe," This species Sanens from Central Bolivia to Central Arg=- entina, mostly in the mountains Neb atualne ox nace ~ BOLLVA Bridges S.ne oO: oe BAUBA: Vile-Vila a tacos. 400 n. == STIS LP) larecaja: San Pedro cerca Sorata, 2650 m., 9 Nov. 1897 & es ig 898, setae 767 (F, G, NY, S, US); : Valle- 00 m, 13 Mar. np teinbach 3725 (IIL, WY, “US) a and Bergwald, 1600m., 13 0 5 A, LIL, ‘Ny, 8); Pt ae Tambo, feet si tambo y Narvaes , 2 one 873, Lorentz ; Hieronymus s.ne (CORD); 73, Lorentz & Hleronyms a pitas _ Ledesma: eGerr. de Calilecua T, uri S871 (F, LIL, S, st, US); SALTAS CAbmEARTAN mastrade c Un- 340 PHYTOLOGIA Vol. 8, no. 7 quillo, 1200 n., a Nov.1931, Schreiter 6644 (LIL, NY) and 6647 (LIL, NY); CHICOANA: San Pedro de Hscoipe, 29 Aug. 1949, Hueck 231 (LIL); COROWEL OLDES: La Hoyada, 14 Nove. 1942, Burkart 13148 (SI); GUACHIPAS: Quebrada del rio Alemania, “bosques xerofilos", 12 Auge 1936, Cabrera ors (LP, NY, SP); Pampa Grande, Nov. a Lee en Pampa Grande, Jan. 1897, Spegazzini ina: Jane *1897, Spega- zzini 2M 3 Ile Vina: Jan. 1897, Spegeezint 1342 (LP); o del foes 9) 1100 m., 26 Nov. 1888, Lillo 1015 iy ee Cuesta de las Juntas, 1500 2 23 Dece lode, T383 (LIL); BURRU He Cerro del Campo, 1200 m., Mar. 1918, te ti 915 (LIL); Rio del Nio, 10 Febr. 1914, Monetti 3; Cerro del Campo, 1000 m., 14 Sept.1928, Venturi 7Slo (BAB, F, US); CAPITAL: Alto de la Polvora, 500 0 iy. Oct. 1924, Venturi 3322A (BAB, LIL, LP, S)3 CHICLIGASTA: Alpachirin, 600 m, 7 Mar. 1924, Venturi 3322 (LIL, US); Rio Cochuna, 10 Nov. ey Villa 645 (LIL); TAFI: Cabro- harco, 12 Nov. 1944, 4 TERE; eas CAS: Thi myics tna 1300 m, 13 Jan. 1 1888, EETTo 1876 (LIL); Lomas de 24 Sept. jue Schreiter 9 and 1431 (LI); CATAMARGAL Jun. 1942, Schaefer sen. 3 Febr. 1897, Sperazz 40 LP); Anbetoetemteo eT eg hi Pobre TSE Parodl 14009 (LP); ANCASTI: "dol fruto se prepara una bebida per- manente llamada "aloja Jan. 1915, cones Soe Bate Huillapinna, desde eee. hasta Pampichu 1909 § zzini sene (BAB); ANDALCALA: 21 Sept. 1945, pa Ae 5.Ne ‘acS); Andalgala, 10 Oct. 1915, Jorgensen 969 (BA, LIL, a ae and Dec. 1914, 969 (HA); Fuerte de imialgaia, Oct. p | ° $b . o ° , ee 3 oe w pe fe} dq se ala, 0 on 165 (1339) (LIL) and 21 Jun. 1908, 291 de la Guchi lla del Cura, Jane er Oo, Ss egazzini Ss Ne (BAB); EL ALTO: Bl Alto, 7 Nov. 1948, Arganaraz 720 Ts Sierra de Gracian, 15 sity Castellanos s.ne (BA RIOJA: Quebrada de Talampaya, 28 Jane 1906, Kurtz ito, CAPITAL: Pere de la in ce al K 9-10, 450 m., 10 oct 941, Alaniz 30 (LIL, LP, U); GENERAL BELGRANO: Sierra de olte, ete del Medio, 3 Febr. 1940, Castellanos s.n. ( and 8 Febr 1940, sen. (BA, LIL); GENS 2 Ghepes, 10 Nov. 1945, Cuez2Z0 1004 (ITEJ; GOBERNADOR GORDILLO: Retamo, 220 m, 17 iM Ms jo ssyenehs 20 (BAB); SANAGASTA: Sierra Velasco; Yacuchi, ) de Te Casa de Piedra, 2100 m., 28 Febr. 1908, Kurtz aber "(cm ; Comechingones, Castellanos sen. (BA); r ot 7 —- 1882, Ga Galander s * (CORD, G, OC), Bajo Velis, - 19 Febr. 1895, thax (iF) and 8532 (Gono) anc 8578. (cond); quebrada de los Cordones, 4 Sept. 1 “ L); AYACUCHO: 24 Nov. 1926, Castellanos Sn» Gait 1962 Barkley, Lithraea 341 BELGRANO: Caleras del Gigante, 14 Nov. 1926, Castellanos 8.De (BA); hego2A, 1922, Gez 70, (SI); CAPITAL: Sierra Varela, C. SeNe ap JUNIN: Piedra Blanca, 2 i 2 1910, Gerth see Febr. 1942, Digilio s.n- ); Merlo, Nov. (L)s Rincon del =ste, 950 Br 10.Febr. 1948, Meyer 13795 (LIL) 12 Febr.1948, 13857 (LIL); P,edr anca ra Sete 1942 re 21 (LiL); PaDERNERA: El pie ner’ Jan. 1933, Vigna nati 36a (LP); PINGLES: Cerro en la Aguada, 5 Nov. 1940, Burkar 10950 (LIL); Quebrada de los mae ress 12 Febr. 1925, Cast- @llanos Sen, (BA); Cerro de la Aguada, 5 Nov. 1940, Covas 1095 LP, NY); CORDOBA: ira Bruch SeNe (Ba) and 30 (LP)7 Castel=— lanos 711 (F, aa F 36 (LP); Lorentz sens (SI); 25 Mare, Lossen 453 (F, SI); aetternrca, Cordon Criental, 17 Aug. 1926, Millan sen. (BA); CALAMUCHITA: Sierra Chica, Arroyo Cachicural, 1000-1800 m., Mar. 1894, Bobenbender s.n. (NY, R) and 8206 (CORD); Sierra Chica, Los Cocos, Mare 1940, Cabrera 6388 (F, NY); Sierra Chica, Estancia Ila Reduccion, 600 m, 28 Dece 1935, Burk- art 7303 (F, SI); Capilla a Los Cocos, Nov. 1915, eee Suis (BA); Sierra Chica, entre Dique y Casabamba, Sept. 1904, Holm- berg sen. (SI); Stocba Chica, La Falda, 1000 malt. 8 lay Tol’, Osten 10445 (LP); Sierra Chica, Nov. 1888, Spegazzini 1497 (a, Sie ca de Cordoba, 3 Oct. 1904, Stuckert CAPITAL: Altos Sud de Cordoba, 17 Febr. 1897, Stuckert 1809; alrededores de Cordoba, Nove 1902-Febr. 1903 ZzZin LP); COLON: Sierra de Cordoba, Carlos Paz, 14 Febr. 1939, Brid= Sorts 595 (LP); Ser pein A i ae Sen. (LP, S); Unquillo, ol Gonzalez Bono (FACCBA) ; Raailic: ¢ Villa Diez, vee, Gristiont Bit 74 (BA); camino Bl Cuadrado, voy 1958, Danes 5 (LF; Wo Ceball allos, Oct. 1875, Hiere us 2 F > CORD); cer a (CORD); Rio Ceballos, Jun. 1939, Lahitte sen. (babyy Siorre de eee? Oct. 1872, Lorentz 381 Tt Sates puedes, 26 Mar. O'Donell & Rodi uez 691 Sreek. rinoe Cabana, 5 Febr. Pool » Scolnik, Araque ¢ Darkle oe LIL); Sierra de Saldan, 97, Stuckert 3123 Casa Bamba, 23 April ieee a basis rt 6805 con ISCHILIN: i? Sept. 1945, Hunziker 6100" ( 3 Jas Penas, Febe 1871, lorentz 00 . 381 (CORD, GC); Gui Ting @ San Jose, 390 m., 17 Dec. 1947, Meyer 15471 (LIL); 3e Stoker 0 (CORD, LIL) ana 13 Nov. 1896, Sen. (CORD); P Capi shay reise Nove il. Bodenbender s.Ne (BAF) ; Ca pila del oe Toma, 15 Jun. 1945, Castiglioni 1908 (BaBosa) 5 Cap= 2a, del Monte, er 1909, Guyssegir e. 2 : eee la Canada, n las pendientes inferiores de e Azucar cerca de Rio srhion, 20-23 Deo, 1878, Hieronymus 76 (CORD); Los Cocos, Tro Caravuelta, Valle de Punilie, ept. 1917, Hosseus oat (CORD): Cuesta de Co baaiia 23 Jan. 1885. Kurtz 2 » Pan © Azucar, 1 Febr. 1885, 319 (corD) and eng’ ada i T Pantano oo San Roque, Sierra Chica, 14 hag. 1505, urte & Glaser we38 (coRD); Cosquin, Pan de Azucar, 1000 2. akaeaFob, 138 , 3h2 PHYTOLOGIA Vol. 8, no. 7 Rodrigo 550 (LP, N Y, 8); vicinity of Cosquin, 10 ~— 1915, Rose 21081 (NY, US); Ie Cumbre, 25 Jan. 1948, de la Sota 466 ‘ ahiay 27 Jane 1948, 485 (LIL) and Parque Los Cocos, 14 Febr. 1948, de la Sota 720 (LIL, U); Huerta Grande, 20 Febr. 1897, shirt 2108 (CORD), Ochoa, 17 Nove 1902, * 12055 (CORD, G, LIT), choa, Sierras re Dec. 1900, 998 ees LIL), Ochoa, 3 ee 1904, 14399 (G, LIL), los Conc pril 1909, 20244 (CORD), and Huerta Ubande, 10 May 1910, Stuckert 21548 ); Sam Sebastian, 22 Jan. 1947, Vi liafane at0° (C, LiL); SAN ALBERTO: Cordon Central, Pampa de Achala, 9 Febr. oe oceans sen. (BA) ; Mina Clavero, 25 saat vee egre, 3 Febr.1948, Meyer 13296 (a1); is “Hevero, 1000 me, 16 Dece 1901, Stuckert 10603 (CORD, cerca me Clavero, 1000 m., 18 Jan. 1950, Vervoorst bet M34 SAN : San Javier, 700-1200 m, 30 Dec. 1926, (LP); 20 ae core Castellano sen. (BA); Dique de la Vina, 30 Jane 332 (LIL); SANTA 1948, Meyer MARTA: entre Potrero Fa og ay y los Wettace 2 Nove 1949, precer.ckas 6509 (LIL); © o la Potrerillos, Jun. 1938, Devo to oF (BABosq); TOTO TOBA fede? chinga, 20 Oct. 1h Giardelli 482 (SI); TULUMBA o Col- orado, 16 Oct. 19 > wane ter 776858 (CORD); Cerro colerades 25 Aug. 1939, age hse: 120 (BOC, LP, SEN sceptowmremncenerrauter ann eT re 4. Lith raea caus (Molina) Hooker & Arnott Bot. fT OAL 175. 1833; Sngler in Martius & Eichler, Flora, 2°; 396. 1876; Engler in De Candolle, Mo ono Phaner. 4: ri 1883; oe Anal. Univ. Chile 47: 62. Reiche, Flora de Chile 2: 26, T8S0: Cabrera, Reve Museo de la P Plata Des. 23 Wonk To38 Sap es & Cass le del Chile ———, ca bapa Seggio Sulla Storia Naturate Co” Persea cnoethon ‘kaa Syst. Vegeot. 2: 269. ot Lithonek Nenenosa liers, Trav. Chil. 2: 529+ 1826 Rhus caustica Hooke & 4r Bot. Bote pt. Beech 1s, tr ) nott, ot. of Ca ech Voye Duvaua ua pleuropogon Turoz, Bull. Soc. Net. Moses I, 51s 467. 1858, prep. : j Schinus Lithi Domb. ex March., Rev. Anace 183. 1869, in Syne Mpuria sim spoitolis Humbe & Kunth. ex Hook. & Arne, Bote Misc. 3: 175. in 3 ongl. in eg & Bichl., Fl. 3rasil 1e*s 396. 1876, ins ib or vin tt tree to four meters tall, evergre en; b ranch= es (mostly growing in length in February) ereenish-tan and mod~ apes thick, later becoming tan and then dark gra o etana or emarginate at apex, cuneate +a truncate at base, usually 21 3 x 45 mm. but rerely from 16 x 25 m, to 7 1962 Barkley, Lithraea 343 Ime, veins white and ee margin thick and white, entire to irregular and subrenate, petiole 3 to 15 m. long, thick, often winged almost to base by the decurrent lamina, glabrous; panicles axillary, long, very sparsely pilose at the base of t lorescence; flowers yellow; cts lanceolate- deltoid ownish, ab ng, persistent; pedicels in tw m olate, glabrous, obtuse and slightly inrolled at apex, 3 mM. long, 1 mm. wide, stamens 10, anthers 1.6 mm. long, filaments 1.5 mm. long, enlarged twoard the base; disk cream-colored, 10 lobed, 1 mm. in d diameter; pistil 1 mm. long, three stigmas and ep pcr yellow, readily separating from the black thin resinous meso- carp, Seed tan-brown, very slightly rugose, 4.5 mm. high, 5-5 Mm. wide and 2.5 mm. thick. Typical specimen: from las planicies del Rio res opel. Chile, 18 January 1941. nnames: "Litre", "Lithi", "Llith the sDecies is limited in ter tenier pre to central Chile. pecimens examined:- CHILE: _—— sen. (G); la = 1828, ne 413 ey p. (G); Dombe ); "Pero iP Dombey-884 (C, I) and 885 (L, NY); "sTUst. 1896, Dusen bape S); Gay sen. (G); Compania, Nov.1853, Germain seme (SGO); Pavon 994 (€); Chile austral, Poeppig Sen. (L); Pawel tucioa, Sept. 1907, Reiche sen. (SGO ; Soar O: Illapel, Pichidangul, 5m. alt., 15 Oct. 1948, Looser 5632 SGOLooser); OVALI Quebrada del engeoeel: 12 Nov.1937, Ibarren sen. (SCO); ICON SAGA: Punta es, 15 Oct.1948, “forma rostrera", Kausel 2665 (ScOKause1); los Molles, Nov.1862, Landbeck Sene (SGO) 5 Punta Molles, 50 m. alt., 15 Oct. 1948, Toossr S881 (S20Lo0ser); Tih : : 8 Oct. 1895, bb S. 3 rane 3 cau — 3 325, Macrae s.ne (G Sebi. ertvero Sele 3 ' Nov. 1918, Claude Jose n 843 US), Jan. 1919, 762 (US) Snd 859 (US); Quebrada de Pinalolen, 850 m. alt., 1l Tar. cp te ‘ “arkle 210H040 (SGOKause1) and 21ch041 (sco Ms. ce (SGOKausel) and alt., Kausel & Barkle 21ch367 (S0CRause1) 870 m alte, 20 “Nov. 927, Montes 456 (F); Cerro San Cri Astobal, Mie servi yaar Sen. (BA); Cerro san hnavctal. 1858, Ph ‘tipi SoNe Checabuco, April 1879, s.n. (Sco); Apoquindo, 929, peotfond 3h PHYTOLOGIA Vol. 8, nos? ski Sone (SGO@F); Quebrada Dormida, 8 Mar. 1951, Sudzuki, Sotome & Barkley 21chO09 (SGo); TALCAs Talca, 16 Dece 1925, cuncke- aig eemnennrs O'HIGGINS: May 1829, rertere 413 Pepre 3; Termas de Cauquenes,750 m alt., 21s 15 1946, Looser siés (SGOLooser); Oct. 1890, Philippi 378¢ Gr 3 ak 3000-5000 m. alte, 14-17 May 1882, Ball oif acienda Los Tralhuenes (Hiuque), 15 Febr. 1946, oa sek 1792 eeopi cael) Cutemu a Quirahue, 150 malte, 17. Febr. 1946, 1852 (SGOKausel), and Hacienda Lolol de Cerro Robles, 200-900 m. alts, 19 Febr.1946, Kausel 1895 (SGO Kausel); Banos de Cauquenes, Se cURRens del rio Cachapoa A 219 Llico, Dec. 1861, Miers Sene (LP); LINARES: Cordillera de la Philippi sen. (US); aah Nov. 1926, CLON: Talcahua - 1896, Dusen ol, 15 Nove 1923, soit 29 ene Tsla, Quriguina, 23 Nov. oie, 398 (SCOGunckel) and P Pedro del Rio, Gualpen, 150 m.alt., 1 Jan.1941, ci 95918 (LP, SGOGunckel); Chiguayante, 21 Nov. 1934, zune Hualqu mvyida, 12-25 F Febr « 1945, 1578 (LIL ) and 1580 (LIL), and lera- queti, 14-27 Febr.1949, KauseT 2800A (LIL, —SGokausel); Cerro Caracol, 7 April 1924, Looser 5649 (SCOLooser); Concepcion, Mertens sen. (NY); Fondo renal, 21 Nove 1932, on 2530 (BOG, RB); San Vicente, Oct. 1890, Philippi 378a (Sc — back of Jalcahuano, 22 Nov. 1916, Skottsberg « Skotts= sen. (S) and 1366 (F, NY); CAUTINs Almagro, 50 m. alte, I “ br. 1943, Gunckel 14181 Se eee): Se MALLECO: Cordil- lera, Nahuelbuta,8 Dec. 1943, Lavin s (scOGunckel); Angel, 10 Febr.1892, Kuntze s.n. (NY, US)e Lithrae veR pilosa. Engle in Mertius & ichler, icreccts 6. 1876. Lithraea caustica £ ilosa (ingl.) Cabre, Reve Museo de bmn th ies. ae rigid branches y branches ten and densely pilose at first, later becoming tannish-gray eventually glabrate; leaves simple, oblong-obovate, 4-7 cm. long, 1.5-3.5 om. broad, coriaceous, margin somewhat waxy but rarely plicatulate, te white corneous, the veins whitish, the veins above, and the under surface, soft pee at least at first, eventually becoming glabrate, apex truncate or emargin= ate, base broadly cuneate and slightly decurrent on th 7 rminal contracted panicles; axis not the sna long pilose; lenceolate-, Pe a July 25. & 37h PHYTOLOGIA Vol. 8, nosc? 1951; Connor & Adams, Poison. Pl. N. Zeal. 106 & 135. 1951; Ahles, Bull. Torrey Bot. Club 78: 269. 1951; Howell, Wasmann Journ, Biol. = a a R. Mohr, Deutsch, Med. Wochenschr. 79: ola 2955 Karrer, Helv. Chim. Act. 37: 790. 195; Univ. ladt of Sootia..7, » 1985; Kasapligil, Pl. Jordan 102, hice, Journ, Fac. Sci. ott PRPs 3 ser. 3, eect 36. 1956; Brain-Slabut & Bolés, Anal, E - Exper. Aula Dei 5: 1, 62, 68, 1 202, pl. 8 & 25. 19575 Univ. Hull List of we “Ts i908, od Sellowia 11: 5, 20 79, & 134. 1959; Winde & H&nsel, rob. Pharm, 293: 556—567. 1960; H. V. Sm., Mich, Wildfls. 313—315, 317, & Leyd, Prod. 326--327 . 530, (arto) is “sonctines or ted as "Royen vats 161"; L., Mat. « 6, 208, 8 (17h9) is pena tiaee cited s page "38": L., Sp. oy ed. 1, ov yer 1 & 32 and ea hiss. & 660 (1753) is sometimes cited as "878 & 879%; Kniphe, Orig. Herb. Viv. pl. [284] (1757) is soetines cited as Poth : 12, no. 99"3 J. A. Murr, in L., Syst. V ed. 13, 61--62 (177) is sometimes cited as page "66"; Lour at; & Cochineh. eo, Ode 26-~27 & 721 (1790) is sometimes cited as page "33", which is the page in edition 2; Burbidge, Book Scented Gard. 7h & dere (1909) is sometimes cited with the letters "n. e"; while the Tre Jard, Bot. Buitenz. 3: 77--87 (1883) soney eee: pean He the ee of this gems contains no reference whatever to it. dditions and emendations to the List of Exp luded Species: Verbena unerins frutesc, etc. Breyn. = Stachytarpheta mutabilis __ (daeqe) Vahl Verbena americana ee etc. Breyn. = Stachytarpheta mute mutab- a tt ilis (Jacq.) Vahl Verbena a americana Sexcican , teucrii foliis & facie, floribus ee, ~~ gaeruleo— scentibus amplissimis Breyn. m Stachytarph- eta mutabilis eESe a. Verbena americana frutescens, teucrii foliis foliis, floribus 5 coer ascentibus amplissimis Breyn. = nm, = Stachytarph ta mutab- ilis (Jacq.) V. Verbena americana latifolia etc. Magn. = Stac eta mutabilis (Jac acq.) Vahl Verbena americana latifolia ica simplici longissima nigro- satifolia, spica simplici longis Vahl ~~ purpu purea Magn. = Stachytarpheta mutabilis (Jacq 2) V Verbena americana media annua ocimi folio Siar Tomgsueo, flore 1962 Moldenke, Monograph of Verbena 375 purpureo amplo Breyn. = Bouchea prismatica (L.) Kuntze Verbena americana media annua, ocimifolio lanuginoso, flore reo amplo Breyn. = Bouchea primat ca (L.) Kuntze eine, cecieeiemniicenmiaens -mmmnimamacnamemmnrs mmamacapea ztrLore coccineoe spicata = Stachytarpheta mutabilis (Jacq.) Vahl Verbena americana, veronicae foliis, flore coccineo spicata breynii Conall. = Stachytarpheta = matebilis (Jacq.e) Vahl Verbena oguvoides major canadensis Munt. = Laportea canadensis aceae L.) Gaud. Verbena bryoides eee, Munt. = Laportea canadensis (L.) aud., Urticaceae Verbena capensis folio capillacea Ray = Hebenstretia erinoides f., Selaginaceae Verbena caudata americana flore pallidé coeruleo Munt. = Stachy- tarpheta jamaicensis (L.) Vahl Verbena caule repente, foliis eee superne crenati pedunculis solitariis Gronov. = la nodiflora c :) Greene Verbena caule repente, foliis pet superne ne crenatis edunculis solitariis capitatis Gronov. = Phyla nodi flora (L. Greene Verbena curassavica scutellariae foliis, flore purpurascente H tze Verbena Vahl Le tidiipatininmia mena a P ° Verbena diandra, spicis laxis, calycib. alternis prismaticis ~~ truncatis aristatis, fol, ovatis obtusis L. = Bouchea » spicis laxis, calycib. fructus reflexis rotundato-didymis hispidis L. = Priva mexicana (L.) Pers. Verbena dianc diandra, spicis longis., calycib. aristatis, fol. ~ ovatis argute serratis L. = Ghinia curassavica (W.) Millsp. Verbena diandra, spicis longiss. carnosis mudis, fol, lanceolate ovat., oblique dentatis, caule laevi L. = Stachytarpheta indica (L.) Vahl ee Gre ee aie Verbena diandra, spicis longiss., foliosis L. = Stachytarpheta orubica (L.) Vahl Verbena diandra, ep ovate, fol. lanceolat. serrato-plicatis, eave frut fruticoso Royen = Phyla stoechadifolia (L. all Verbena di. dispermos americana lignescens veronicae foliis Moris. = ~ Bouchea prismatica var. longirostra Grenz Verbena femina Brunf. = Erysimum officinale ‘ , Brassicaceae Verbena fol. ovat. acuminatis, spica foliolosa L. = Stachytar- -_pheta o orubica (L.) Vahl erbena Zp fol. subrot. subrot. serrato etc, Sloane = Stachytarpheta jamai- , geese, censis (L.) Vah Verbena fol foliis lanceolatis plicatis, spicis ovatis, caule fruti- 376 russ. VnPe UL Ss Vol. 8, no. 7 coso Royen = Phyla stoechadifolia (L.) Sm Verbena foliis cmiaetiatis. spica carnosa Sanat ae = Stachytar- ceeceienincapenane tangas” cigs aim ena ee ce mr ee es e V Verbena folio itrctundo serrato, flore caeruleo Sloane = Stachy- tarpheta ahi Verbena folio folio subrotundo serrato flore coeruleo Sloane = Stachy- " tarpheta jamaicensis (L.) Vahl —~S Verbena fruticosa Millsp. = Phyla nodiflora var. reptans (H.B.K.) ~~ Moldenke & P. strigulosa a (Mart. & Gal) denke Verbena indica Bont. = Stachytarpheta jamaicensis (L.) V: Verbena indica lanuginosa Bod. = Manulea rubra L. f., sso " daceas ae a BR Verbena mar mariana, rosae chin, fol, etc. Petiv. = Phryma leptostach- ~ Phrymac Bleasdale dead oe L., Phr Verbena wekiban: L. = Priva mexicana (L.) P Verbena mexicana, trachelii fol. ete. ones = Pelee mexicana (L. ) ers. Verbena mexicana, trachelii fol., fructu aparines Dill. = Priva mexicana (L.) Pers. Verbena minima, chamaedr, fol. Sloane = Bouchea prismatica (L-) ——$——— ee Kuntze Verbena minima, chamaedros fol. Sloane = Bouchea prismatica (Le) Kuntze Verbena minima chamaedryos folio Sloane = Bouchea prismatica (L.) 7a REE ee ee ere eee Verbena; nodiflora C, Bauh. = Phyla nodiflora (L.) G Verbena nodiflora capite oblongo, seu V. nodiflora tapas ae = Phyla nodiflora (L.) Greene Verbena nodi flora curassavica, foliis menthae Herm, = Ghinia COP” assavica (L.) Mill Verbena : nodiflora, ‘foltis ‘nigeceinn viridibus Moris. = Phyla noe nod= iflora L. en alee Verbena, no retiflora,, foliis minoribus, viridibus Moris. = Phyla “nod #1 ra (L. J Greene Verbena aie ii incana curassavica latifolia Herm. = Phyla nod- ~ dflora ra (L.) Greene Verbena nodiflora incana, Ce latifolia Herm. = Phyla ~~ nodiflora i. oa CC Verbena orubica, teucrii $68 etc. Pluk. = Stachytarpheta MIE mutab- ee “Vahl Perel TOE lis ee ct nm tenement 1962 Moldenke, Monograph of Verbena 377 seminib. longissimis Pluk. = Stachytarpheta mutabilis (Jacq.) Verbena orubica, teucrii folio, primulae veris aoe a s& ~~ geminibus . Aine inept Pluk. = Stachytarpheta mu (Jacq.) Verbena racemo simpliciss. etc. Gronov. = Phryma leptostachya L., FP eae Verbena racemos simplicissimo, floribus sessilibus, calycibus "fructus reflexis, racemoque appressis Gronov. = Phryma ‘lep- acea Verbena recta & mas Gesn. = Erysimum officinale L., Sa Verbena repens nodifl. Burm. = Phyla nodiflora (L.) Green Verbena repens nodiflora Park. = Phyla nodiflora (L.) avels Verbena rubra | Rumph. = Aerva sanguinolenta (L.) Blune, Amaranth- Bouchea Verbena scutellariae foliis dispermos americana Pluk. = prismatica (L.) Kuntze Verbena scutellariae s. 28 Ee fol, etc. Pluk. matica (L.) Kuntze si hae. =the, scutellar nl 8. ot fol. fol., dispermos americ. Pluk. = Bouchea coer pant (L.) Kun Verbena ae sive — wens dispermos Pluk. = Bouchea "prism ca (L.) Kuntz Verbena spats jamaicana ate Pluk, = Stachytarpheta = Bouchea pris- jamaicensis ees Verbena betaates jamaicana teucrii pratens’s = folio, dispermos Pluk. = "= Stachytarphe ta jamaicensis (L.) V Verbena spicata jamaicana, teucrii serene folio, dispermos Pluk. = Stachytarpheta jamaicensis (L.) Vahl Jerbena tetrandra, calyc. subro mane erectiusc., sem. echinatis L. = Priva lap , 8 Verbena tetrandra, spicis ss Mighecsteaia, 20045 foliis serratis, caule repente L. = Phyla nodiflora (L.) Greene Verbena tetrandra; spicis capitato-conicis; foliis fuse serratis; ~~ Gaulle repente L. = Phyla nodiflora (L.) Green The following thirteen pre- pie 8 Epes have pie as yet been satisfactorily identified by me. 7 readers of this Paper, specialists in the regions tot these names actually apply, it would "i yar gc appreciated if this information were communicated to me: Yerbena americana radice grumosa Munting apud Ray, Hist. Plant. 3: Suppl. 285. 170h. Verbena foliis planta sesso quadricapsularis, flore parvo Calathoide coeruleo Pluk. apud Ray, Hist. Plant. 3: Suppl. 286. 170). Verbena fructu in singulis capitulis echinatis, unico flore albo 378 Pe y f:O LOG Fa Vol. 8, no. 7 Sabbat in Martelli, Hort. Roman. 3: ll. (eon tn Oe TI. 3: Suppl. Verbena bond foliis ses 8 ae Boe. Spei Petiv. apud Ray, ~~ Hist. Plant. 3: Suppl. 286. Verbena indica, rotundifolia, Jasle comosis Mus, Zeyl. pag. 58 Se ee Se Pa snrnin ee eeper es eeeornn Speenens. fo oe 3, pa Tag Herm., Thes. Zeyl. 228. 1737. Verbena eae incana minor Indiae Orientalis, Poordela Mala- ~——parorum Pluk. apud Ray, Hist. Plant. 3: Suppl. 266. 1704. Verbena nodiflora major indica flore ad hermanni Morison, Pl. st ee eee Ray, Hist. Plant. 3: Aap. 287. te Verbena pervana perpetyo virens J. Bauh., Hist. Plant. Univ. 3s 2% 5 pai —ihs. Verbena scorodoniae folio spicata marilandica D. Sherard apud ~~ Ray, Hist. Plant. 3: Suppl. 287. Verbena taxi foliis, marginibus lanuginosis Petiv. apud Ray, Kae ee eee ct ce Pl Py t Verbena trachely, ruvidis foliis, fructu aparines H. Eyst. apud Sabbat in Martelli, Hort. Bua ot Ll. I7%s. Additions to the list of known hybrids: xVerbena gonzalezi Moldenke = Verbena santiaguensis (Covas & Schnack) Moldenke x V. perakii (Covas & Schnack) oldenke xVerbena perplexa Moldenke = V. gooddingii Briq. x V. bipinnati- fida Nutt. Nutt, xVerbena tentamenta Moldenke = V. perakii (Covas ¢ Schnack) Mol- hse x - = (L.) Britton note that Lamarck & DeCandolle, Fl. Frang. 3: SOs 30% (A808) ae a Verbena: "Ce genre, placé par Lamarc arck dans les Labiées, et par Jussieu dans les Fyrénacbes a eaidat la liaison naturelle de ces deus familles.' VERBENA ABRAMSI Moldenke Additional literature: Moldenk Fa arg cua 8: 142--1h5 (1961) and 8: 177. 1962; iloldene , "Résumé Suppl. 3: 8. 1962. s formosum H eB.K., and Gnaphalium chilense Spr enge ae citations: CALIFORNIA: Madera Co.: Brock 297 (Hi-- Ol XVERBENA ADULTERINA Hausskn. od In my Alph. List Invalid Names Suppl. a 22 Sigel I reduc this binomial to synonymy under V. officinalis L. and fo lLlowed this disposition of it as late as my ar in isso. It is only 1962 Moldenke, Monograph of Verbena 379 since the latter date that I have decided to keep it apart. VERBENA ALATA Sweet Additional & minitied literature: Briq. in Engl. & Prantl, Nat. orn 4 (3a): 147. 189); Reitz, Sellowia 13 (13): 110. 1961 VERBENA AMBROSIFOLIA Rydb. Additional literature: Moldenke, Phytologia 2: 126 (195), 8: 149--152 ‘pete oe and 8: 177--181, fig. 2. 1962; Moldenke, Résumé Suppl. 3: 7 VERBENA “sesso f. myer ire A ional literature: Moldenk , Peytolocia 8: 181—183. 1962; aAderike, Résumé Suppl. 3: 37. 1 VERBENA ARAUCANA R. A. Phil. Additional literature: Acevedo de Vargas, Bol s. Nac. Hi Nat. Chile 25: 50. 1951; Moldenke, Phytologia 8: “187-188. 1962." cevedo de Vargas states that al tipo de esta especie, con- has Py NA ARENARIA Mol eee rature: Moldenke, Phytologia 8: "gave: oe and 8: 188-- 189, 1962 Moldenke, gy RA Suppl. 3: 1h. ENA ARGENTINA Moldenke Literature: Moldenke, Phytologia 8: 60 (1961) and 8: 189. 1962; Moldenke, Résumé Suppl. 3: 1k. 1962. VERBENA ATA’ TACAMINSIS Reic Additional literature: as do de Vargas, Bol. Mus. Nac. Hist. Nat. Chile 25: 51. 1951; Moldenke, Phytologia - “191—192. 1962+ of this species was coll lected in September, 1900, e al in Santiago, Chile. Ace argas Re A. Philippi s.n. [Paposo, A56 _Pablished ~ Harold N. “Moldenke and Alma 1, Moldenke - 15 Ghesbrock Avenue ee! Ooms 5] New York, U. S.A ey Price Sf sii oumber $1; per humts75. a advance Vol. 8, no. 6, was issued May 1, eer oe ie 8 no. 7, was ‘issued Brel 26, em NOTES ON NEW AND NOTEWORTHY PLANTS. XXXII Harold N. Moldenke AEGIPHTLA SESSILIFLORA var. CUATRECASASI Moldenke, var percrassis usque ad 17 m, longis, calicibus fructiferis campanu- latis percrassis usque ad 10 m, longis et 17 mm. latis verrucosis secdeerg lobatis, et drupis carnosis 2 cm. longis latis sque This variety differs from the typical form of the species in having solitary fruits, the fruiting-pedicels very thick, to 17 mm. long, the fruiting-calyx coriac aceous, campanulate, to "10 mm. oe 17 mm. wide, verrucose, and the drupes fleshy, 2 cm. long e The type of the variety was collected by José Cuatrecasas Arumi and Rafael Romero Castafieda (no. 25404) in the — at Seckacae Nuevo Pe face El Podrido, Sierra de Perija, of Manaure, 3 Colombia, on November 16, 1959, and ag poirot in. the H. N. MolAenike Herbarium at Yonkers, New Yo rk. ~via VITELLINIFLORA var. EGLERI Moldenke, var. Haec varietas a forma typica sooner calicibus hadi pil- Osis ie subglabratis recedit. S variety differs from the typical form of the species in th anthesis very sparsely pilose with long Pi &§ Th y was collected by H. S. Irwin, W. A. Fgler, and J. Murga Pires (no. 47162) on riverbanks between Olapoque and Santo Antonio, on the Rio Oiapoque, Amapd, Brazil, on July 26, 1960, and is deposited in the Britton Herbarium at the New York Botanical Garden, It is named in ren and commemoration of Walter Alberto Egler, whose tragic while ona botanical expedition was a great loss to A a, b CALLICARPA ERIOCLONA f. GLABRESCENS Moldenke, Résumé Suppl. h: 9, nom. nud. 1962; f. nov. * forma a forma typica speciei foliis maturitate subtus 8 parcissimeve stellato-pubescentibus vel glabrescentibus re- av This form differs fran the ec form of the species in hav- ing the mature leaf-blades varying from rather sparsely to very ely s tt i tangs or even eh cbedendal and resinous- genie e lower surface. type oar the form was collected by Takahide Hosokawa A 8398). at Tadiu, on Truk island, Caroline Islands, on August 11, 1936, a * deposited in the herbarium of the Bernice P. Bishop Museun onolulu, 385 386 PHYTOLOGIA Vol. 8, no, 8 ERIOCAULON CONGOLENSE Moldenke, Résumé Suppl. ): 7, nom. nud, 19623 sp. nov. Herba parva annua; foliis numerosissimis lineari~filiformibus erecto=-patentibus pellucidis glabris fenestratis apiculatis; pe- dunculis numerosis erectis filiformibus tricostatis stramineis glabris parce contortis; vaginis laxiusculis; capitulis oblongo~ S. ufted herb, apparently annual, to 5.5 _ tall; leaves very numerous, Lineat-fh45 fo0e; hpi i FE e3--2.4 Cm. ’ ’ glabrous or practically so, slightly twisted; sheaths rather loose, about equaling the leaves and similar to them in texture and color; heads oblong-obovate, 1.5--2.5 mm. wide, 2--3 mm. long, few-flowered; involucral bractlets obovate-elliptic, golden- stramineous, about 2.3 mm. long and 1.) mm. wide, acute at the a- pex, glabrous, shiny; Sao glabrous; “receptacular bractlets ~~ about 2.1 mm. long and 0.7 mm. wide, cream-colored, glab- staminate florets not seen; pistillate florets: sepais 3; sonmate, linear, hyaline, about 1.), mm. long and 0.1 mm, wide; peal separate, linear; ovary nor eile about 0.5 mm. long rou 11642) at a yas Selina Handa and Niangara, province of stanley- ville, Congo, on J 30, 1956, and is a in the herbarium of the Tnststut 1 Francaise d'Afrique Noire at Dakar. ERIOCAULON GUINEENSE Moldenke, Résumé Suppl. 4: 6, nom. nud. 19625 Sp. nov. pedunculis erectis filiformibus stram oe ps tricostatis paulo contortis valde patenti-pilosis ‘ apo pe glabrescentibus3 cap~ tifl itulis hemisphaericis albis dense fo) Annual herb, 1)--17 cm. tall; leaves rosulate, erect or g, rather few, sei 2--3.5 cm. long, 2-3 Wide, attenuate at the everal- to many-striate, glabrous} peduncles 5=-10 per plant, erect, filiform, stramin - 15 cm. long, 3-costate, slightly twisted, ab reading-pilose, the hai ng off in age; heads hemispher- staminate florets long-pedicellate, the sepals 3, separate, ob- long, whitish, about 1.) mm, 1 -35 mm. wide, d te; petal , oblong, whitish, about 1.1 mm S 3, separate, oblo whi long and 0.35 a wide, densely white-barbate; pistillate eget ts: sepals 3, separate, oblong, hyaline, about 2.8 mm. long 0.35 mm, wide, acute at the apex, long-ciliate at the apex only, — lS 1962 Moldenke, New and Noteworthy Plants 387 otherwise glabrous; petals 3, oblong-linear, about 0.7 mn. long small and - Wide, densely white~barbate; o glab = about 0.35 mm, 1 de, 3-sulcate, 3-celled, 3 te. of this species was coll ismare (no. 386) on wet sandy soil at Toumou, Friguiagbé, Republic of Guinea, on September 13, 1943, and is no. 390) in the James Chillou h 8 que Noire at Dakar, ERIOCAULON HIRSUTULUM Moldenke, Résumé Suppl. hj: 6, nom. nud. 1962; sp. nov. Herba parva annua; foliis caespitosis graminoideis Patentibus viridibus attenuatis glabris nec fenestratis non Striatis; pedunculis erectis gracilibus stramineis dense paten- teque Sericeo-hirsutulis; vaginis laxiusculis multostriatis gla- bris; capitulis hemisphaericis stramineis. Small annual herb, 7--9.5 cm. tall; leaves tufted, grass- pc erect—spreading, 1.3--2.2 cm. long, about 1 mm, wide, green, a 8 se many= striate, glabrous, the blade short, erect, plainly apiculate at the apex; heads hemispheric, about 3 mm, wide, stramineous; in- volucral bractlets oblong, stramineous, about 2.3 mm, long and 1 wi s al at the apex; petals 3, separate, linear, stramineous, about 1 m. long and 0.1, mm, wide, laciniate at the apex, glabrous; ovary Subglobose, about 0.5 mm. long and wide, glabrous, 3-sulcate, }- celled, 3-ovulate The type of this species was collected by J. G. Adam (no. 18370) at Asamane, Senegal, on October 8, 1961, and is deposited the H. N. Moldenke Herbarium at Yonkers, New York, SRIOCAULON LATIFOLIUM f. PROLIFERUM Moldenke, Résumé Suppl. h: 6, nom. nud. 1962; f. nov. Haec forma a forma typica speciei capitulis plerumque foliaceo- edit. form differs fran the typical form of the species in hav- ing its heads often proliferating into a cluster of leaves 5--20 ng gal, on Decenber 9, 1951, and is no. 10358 in the herbarium of the Insti- tute Francaise d'Afrique Noire at Dakar. This same tendency to vavipary or proliferation is seer in E. brevifolium Klotzsch, E. Jongifolium Nees, E. modestum Kunth, E. sexangulare L., Paepalan- 388 PHYTOLOGIA Vol. 8, no. 8 thus capillaceus Klotasch, P. fasciculatus (Rottb.) Ktrn., and Syngonanthus caulescens (Poir. ) Ruhl. ERIOCAULON SEXANGULARE f. VIVIPARUM Moldenke, Résumé Suppl. 5: 6, nom. nud. Net 2. nov Haec forma a forma typica speciei capitulis plusminusve vivip- aris recedit. This form differs from the typical form of the species in hav- ae its nes more or less conspicuously viviparous. collected of thoi +1663) in mud by a stream at Rengam, Johore, Malaya, on Oct~- ober 22, 1941, and is deposited in the herbariun of the Naturhis- to riska’ Riksmuseum at Stockholm. LIPPIA | MONAT Moldenke, Résumé Suppl. 4: 4, nom. mud. 19625 sp. g tiflerts bracteis ‘aig papyraceis ovatis; “corollis albis. 12 m, tall; trunk 25 cm. in diameter at breast em. long, glabrous; leaf-blades thin-chartaceous, h surfaces, ic or ovate, 2—-6 cm. long, 13.5 cm. wide, acute or short-acuminate at the apex, s e with re somewhat antrorsely appressed teeth fran below the widest part to the apex, cun ac te at the base, glabrous on both surfaces; midrib very slender, flat above, prominent beneath; secondaries very sle ; de, arcua ing, n stonosing, inflorescence axi » near and at the apex of the branchlets, one per axil, opposite and decussate; peduncles ere ee cm, long, glabrous; heads densely many-flowered, 2.5 pie bracts large, ovate, papery, greenish-white, at? Tile Long, mn. wide at the base, acute at the apex, sessile, densely short- The type of this species was collected by Rogers McVaugh Walter Norman Koelz (no. 1660) on low mountain summits e 7 miles north of Santiago, on the road to Durazno, in deciduous woodlands growing with Cordia, Brosimum, and Platyniseim, at an elevation of 200 meters, Colima | Mexico, on Decembef 10, 1959, and is pos ited in the herbarium of the University of echt gae at Ann Arbor aT: Michigan. The collectors note that it was abundant at the type cality, It is a pleasure to name this fine soeetit in honor of oY good friend, Rogers McVaugh, who has done such n aon the taxonomy of vascular plants, especiall of ths Lobeliaces®: at 1962 Moldenke, New and Noteworthy Plants 389 CHILLOUI Moldenke, Résumé Suppl. h: 6, nom. mud, 1962; sp. nov. Herba parva anma 3 foliis rosulatis patentibus capper or vel pellucidis apiculatis glabris nitidis; pedunculis nomullis stramineis filiformibus }-costatis glabris benbomtte- erectis; i albis multifloris; bracteis involucri e ellipticis stramineis, Diminutive herb, epee Tt 3—~8 cm, tall; leaves basal, tufted, spreading, anous or _pelluetd, 5-9 mm. long, less than 1 m. wide, . ng a ’ short~apiculate at the apex, more or less spre ‘ e oth surfaces, not te nor fenestrate; peduncles very slender, stria erect, 2-6 per plant, 13—30 cm. long, golden-stramineous, more or less densely pubescent or hirsutulous, very obscurely 2-sulcate or subterete; sheaths rather closely appressed or somewhat loose, surpassing the leaves, 5-6 cm. long, obscurely striate, slightly twisted, densely hirsutulous with fulvous hairs, the apical limb ovate, erect, acute at the apex; heads obconic when young, la durin; 5 mM. wide, acu 1. at the apex, with broad lighter margins, densely long-pilose with rse hairs on the backs receptacle long-pilose; reget red, about 0.72 mm, long and 0.35 mm, wide, dens stamens 6; anthers yellow; pistillate florets: sepals apparently 1962 Moldenke, New and Noteworthy Plants 391 surmount late red lobes about 0.7 mm. long and 0.35 mm. wide, densely barbate~fringed; ovary globose, about 0.5 mm, long and wide, glabrous, 3-sulcate, 3-celled, j-ovulate; style filiform, about 0.7 mm, long, glabrous; stigmas 3, minute, erect, papillose or finger-like, The type of this ing and beautiful species was collected by James ohilioe (no. 1838) on very wet sandstone at the planta- tion of | Friguiagbé, in the valley of Quatamba, Re- Public of Guinea, o pbtich ee 10, 1940, and is deposited in the H. N. Moldenke piesa t Yonkers, New York. The floral morph- ology of this plant is not clear. Some flower—heads seem to have ing these are appendages to the receptacular bractlets or to the Sepals is not clear. Almost certainly a new gems is here indi- cated, PAEPALANTHUS BELLUS Moldenke, Résumé Suppl. 4: 5, nom. nud. mo Sp. nov. @ parva caulescens; caulibus brevibus ater eees — 8 onge pilosis, caeterem glabris; pedunculis filiformibus Grisostatis glabris stramineis; vaginis arcte adpressis glabris; capitulis hemisphaericis albidis; bracteis involucri obovatis brunneis. Small caulescent herb, to about 10 cm. tall; stems short, very slender, oye 12.5 cm. long, ascending or — leaves quite uniform -5 cm, long, less than 1 m A opaque, severcaabantell patnay blunt at the apex, ieaechaines on the e margins toward the base, otherwise glabrous; peduncles fili- form, 4-9 cm, long, 3-costate, glabrous, stramineous; sheaths closely appressed, shorter than the leaves, glabrous, the apical e i o long and 0.8 mm + wide, glabrous; receptacle Long-Pilose; recep— paced tract lets oblong, dark-brown, about 1.5 mm. long and glabrous; staminate : brom, acute at the apex, white-barbate at the apex on the back; petals 3, connate into a hyaline tube about 1.) mm. long, glab- rous; stamens 3; pistillate florets: sepals 3, separate, obovate, dark-brown, acute at the apex, white-barbate on the back at the ) long=pilose; ovary subglobose, stramineous, about 0.7 mm. long and oom glabrous, 3-sulcate, 3-celled, 3-ovulate; style about mms long. The type of this species was collected by Gert .Hatschbach (no. 7391) at Rio Coutinho, municipality of Guarapuava, . » On October 21, 1960, and is deposi ved in the H. N. Mol- denke herbarium at Yonkers, New Yor: 392 PHYTOLOGIA Vol. 8, no. 8 PAEPALANTHUS DIFFISSUS Moldenke, Résumé Suppl. : 5, nom. mud. striatis minute puberulis versus apicem fissis; capitulis 1--3 per diviso pedunculorum. Herb; stems apparently thick but abbreviated, very densely leafy; leaves densely cespitose, erect or ascending, 6—10 cm. long, lanceolate, to 1 cm. e at the base, uniformly taper. flat, many: a ery minu apex into 2—-l| divisions of equal or very unequal length, each termina du less obconic or hemispheric, h--9 mm. wide, bractlets numerous, closely imbricate, ovate-lanceolate or tri- angular-ovate, 2,.5--3 mm. long, 1-2 mm, wide, abruptly acuminate from the broad base, ciliolate along the margins, otherwise glab- rous and shiny; receptacle long-pilose; receptacular bractlets broadly elliptic, dark-brown, about 2.{ mm. long acute at the apex, white-barbate on the back at the apex; apex; petals 3, hyaline, forming a tube about 1.7 mm. long; sta~ mens 3, included; pistillate florets not seen. The type of this species was collected by Vareschi (no. 5357) in Espeletieta graminosa, at 3200—3600 meters altitude, Po. El Batallon, T4chira, Venezuela, on October 3, 1956, and is deposit- ed in the Herbario Nacional de Venezuela at Caracas. PAEPALANTHUS MUSCOSUS var. TACHIRENSIS Moldenke, Résumé Suppl. i: » nom. mad. 19623 var. nov. Haec varietas a forma typica speciei pedunculis usque ad 12 em. longis, caulibus usque ad 13 cm. longis, et foliis usque ad 1.7 cm. longis recedit. This variety differs from the typical form of the species in ts cles to 12 om. long, the stems to 13 cm. long, and the leaves to 1.7 cm, long The type of the variety was collected by Vareschi in Espelet- ieta graminosa, at an altitude of 3100 meters, at Po. El Batalion, T&chira, Venezuela, on March 10, 1956, and is deposited in the Herbario Nacional de Venezuela at Caracas. PAEPAIANTHUS POLYTRICHOIDES var. DENSUS Moldenke, Résumé Suppl. 5: 5. 1962; var. nov. Haec varietas a forma typica speciei foliis densissimis arcte adpressis imbricatisque recedit. This vari fers from the typical form of the species ms having its leaves extranely dense, very closely appressed, eg: tightly overlapping during anthesis. 1962 Moldenke, New and Noteworthy Plants 393 The type of this variety was collected by José Cuatrecasas Arumi and Roberto Jaramillo Mejfa (no. 25737) scciaa the Laguna de Chisacé, at an altitude of 3650 to 3700 meters, on the Péramo de Chisacé, Macizo de Bogot&, Cundinamarca, Colombia, on December 29, 1959, and is deposited in the H. N. Moldenke herbarium at Yonkers, New York. PREMNA TAITENSIS var. VITIENSIS Moldenke, Résumé Suppl. i: 9 and 5: 6, nom. nud. 1962; var. nov. Haec varietas a forma t ica speciei venis amine foliorun hi te the apex in attenuate-acute or subacuminate fashion, In var. rimatarensis the leaf-blades are usually more subrotund or oblong in shape, usually abruptly acute or obtuse in submucronate fash- ion at the apex in the style of P, obtusifolia R. Br. The t of the variety was collected by Albert Charles Smith (no. 7029) at the edge of a forest, at an altitude of 100 to 200 meters, in — Rigmigy of Ndakaivuna, in the hills east of the Fainimb , Tailevu, Viti Levu, Fiji Islands, between April 1 and 26, 1963, “and is deposited in the H. N. Moldenke herbariun at Yonkers, New It seems ea “that Brown's var. marchionica, var. rapensis, ant variety or form of P. obtusifolia. However, not ciggane as yet seen the actual types of | P, taitensis, P. ma rP. gaudichaudii, I hesitate to make the toenbfore’ at this ioe It is also not at all clear to me why P. integrifolia L. should not be retained as the valid name for what is now passing as P. ob- tusifolia, The P. Nelson collections distributed and annotated by E. D. Merrill as "TOPOTYPE, typical of Prema gaudichaudii Schau- er" seem to be typical P. mar ee SUAKIEAA Moldenke, Résumé Suppl. 5: 6, nom, nud. 962 gies 8 Frute rg ieasians; ramis subtetragonis glabris; ramulis puberu- lentis; et ceo iee! foliis decussatis; petiolis gracilibus 6—9 mm trigilloso- o-puberulent bus} laminis chartaceis cates: ad basin rotundatis, supra sparsis- Sime strigillosis is, ite had roa nitidis, subtus dense pubes- centibus; inflorescen speech terminalibus paniculatis; bracteis in- 394 PHYTOLOGIA Vol. 8, no. 8 slender, subtetragonal, densely brown-puberulent; nodes annulate; principal internodes 2 37 cm. long; leaves decussate-opposite; petioles slender, 6--9 mm. long, strigillose—puberulent with an- trorse hairs; leaf-blades chartaceous, dark-green above, lighter sely st nealing’ or glabrescent above, densely brown-pubescent beneath; midrib slender, flat above, prominulous beneath; second- aries slender, about 3 per side, ascending, flat above, perorn § prominulous beneath; sattiet reticulation abundant, fo rather conspicuous network; inflorescence terminal on the nancies lets, paniculate, and also a few small cymes in the axils of the uppermost leaves; prey about 23 cm. long and 12 em. wide; cymes pedunculate, about 5 cm. long and 3 cm. wide; sympodia tet~ ragonal, slender, densely brownish-strigillose; peduncles very slender, about 1.5 cm. long, brown-strigillose; sympodia 1--3 cm. long; involucral bracts 6, oblanceolate, to 18 mm, long and 6 mm. wide during anthesis, densely strigillose on ge surfaces, rounded at the apex; calyx cupuliform, about l mm. long and wide, ae canescent-puberulent, the rim conspicuously lobed; corol- The type of this species was collected by my good friend and eee Tem Smitinand (no. 852) — in whose honor it is named attered in an old Sheeeing at Rachasima, Pak Chawng, at an altitude of about 200 meters, in eastern Thailand, on September , 1958, and is acosktek in the H. N. Moldenke herbarium at Yon- ae New York. STACHYTARPHETA INCANA var. JALISCANA Moldenke, Résumé Suppl. hi: h, nom, nud. 1962; var. nov. Haec varietas a forma typica speciei bracteolis angustissime Lnsarttienceutuese et corollis albis recedit. This variety differs from the typical form of the species in having its bractlets very narrowly linear-lanceolate, except for the Pris base almost filiform, and the corollas ° e type of the variety was collected by Rogers McVaugh (n0- 19919), who says that it was locally abundant in wooded ravines In the mountains north of Autl tl4n, 3--5 miles above Mina San Fran: amy (Cia. Minera de Autlén), in the —— paged forest riumfetta, at 1500—16 SYNGONANTHUS PULVINELLUS Moldenke, sp. Herba parvissima acaulescens; foliis vrosulatis linearibus orth ris nitidis apiculatis, ad basin densissime albo-lan ee vaginis arcte adpressis griseis tenuiter membranaceis multistriat paulo peter ou glabris; pedunculis solitariis gracillimis erectié € pilosis tricostatis paulo Sbehae tint capitulis subg*o” roe Rg sin phe bracteis involucri membranaceis lanceolatis aristatis 1962 Moldenke, New and Noteworthy Plants 395 Dwarf acaulescent herb, fo tufts; leaves rosulate us <- erect loosely long-pilose, 3-angled, slightly twisted; heads solitary, subglobose, 3—-l mm. long and wide, erays involucral bractlets membranous or subhyaline, lanceolate, about 1 mm. long, aristate at the apex, broadly white-margined, is a dark “central midrib leading into the apical point, glabrous on both surfaces; recep- tacle pilose The type of this very distinct and unmistakable species was collected by Bassett M Maguire, Geraldo Mendes Magalhfes, oe Celia K. Maguire (no. 49183), occasional on sandstone on eee oad be- tween Diamantina and Gouveia, Minas Gerais, Brazil, on August 12, 960, and is deposited in the Britton Herbarium at he New York Botanical Gar SYNGONANTHUS XERANTHEMOIDES var. GRAHAMAE Molde enke, var. nov Haec varietas a a ee * specied foliis temiter chartaceis submembranaceis flaccidis 5—9 cm. longis 0.5--2 mm. latis ad api- anthesis thin-chartaceous or submembran- ba flaccid, 5—9 cm. long, 0.5—-2 mm. wide, and ahaa narrow a The type of the Rac.oye at was jooteuas by V. Graham (no. 15) — whose honor it is named -- on brown sand at the Mahdia gold- field, Potaro River, British Sapa on July 31, 1958, ami is de- posited in the herbarium of the R Royal Botanic Gardens at Kew MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. VII Harold N. Moldenke VERBENA L. Additional and emended bibliography: Leche, Kongl. Swensk, Wet e Aca d. Handl, S: 262. 17hh; J. A. Murr. Prod, Desig. Stirp. 7 23 if me b. 6: 3M cote and 3: pl. 129. 1839; Gard, Chron. *, Fl. Frang, 2: 718. 1852; Strobl, Flora van Admon 8 Melker Gymnasium. 1882; F. Muell., Syst. Cens. Aust. Pl. 102. 396 PHYTOLOGIA : Vol. 8, no. 8 1882; J. Ball, Notes Naturalist 293. ei F. M. Bailey, Cat. In- dig. Nat. Pl. Queensl. 35. roel teres sch. Pik Monatsc 20: 52. 1902; L. H. Bailey, Cycl. Amer. rie he he 1912— 1916, pl. 26l (1902) and ed. Rie 6: 1912--1916, wie 2649. 1906; Fedde in ba ee: preteeta 33 (1): 632-633. 1906; E. H. L. Krause in S$ me tes - 15: 305. 1907; Murr, Alig. B Bot. Zeitschr, ie 9 (19 08) ae are 187. het Druce, Rep. Bot. Exch. Club Brit. att 1913 ae 425. 1914; L. He Bailey, — Cycl. Hort. 6: 3445-3447, pl. 3910. 1917; "Yolfino, wert : 103. 1923; Britton & P. iils., Scient. Surv. Porto Rico 6: 137, 138, 10, & 142-1. 1925; iarra, Atlas Bot. i 42. 19273 Gonzales, Copp., & Lo TUg « 1: 9& mm 306 1928; He F. Roberts, Pl. Hybrid. before Mendel by FET Ls a & 18. 19293 Herte: , Es a t. Reg. Urug » An. kel, Ber. Schweiz. Bot. Gesell. a eri. & Arbustill. Pas. Publ. 255. 1961; Assoc. a, Trop. Index 1961: 60. 1962; McCabe, Our Par ks 28. (jet ee H. N. Moldenke, Résumé Suppl. h: 1--21 i962) and 5: 18. 1962; Pearce Seeds ants, Our Am, Wild Fls. 10. 1962. Verbena mendocina R. A. Phil. x V. parodii (Covas & Schnack) Mol- d Verbena peruana, quae hispanicae sinilis, ex Monardi, Lugdenensis Teferet C. nae: Pinax igh tr. Bot., ed. 1, 269. 1623. It is of interest to note that "Verbena" is now being used a8 the name of a Ad (of blankets sbeioas by John Wanamaker, Phil- adelphia, 62). The Waterfall & Wallis 13176, distributed as &@ species ~~ Tetons is actually Priva iva grandiflora (Ort.) woldenke. ABRAMSI Moldenke Additional bbiioere phy: Moldenke, Résumé Suppl. h: bed eee The Wilkes 1661 collection cited below fag a mixture lasiostachys var, far, septentrionalis Moldenk Additional citations: CALIFORNIA: Sings ‘Co. : Kearney 2> pale og NI) —xenmeasimnanietistinnemimneniainininiiasinietssiints 1962 Moldenke, Monograph of Verbena 397 410850). Los Angeles Co.: Abrams & McGregor 689 (W--61339h); Braunton 520 (W—h65502). Monterey Co: G. Re V. Vasey s.n. (San An- tonio River, 1880] (W—5622l). San Bernardino Co.: mish ae Par- ish 969 (W—7718L9). County undetermined: Wilkes 1661 1661, Lower Sacramento] (W—71925). VERBENA ALATA Sweet alae and pga ses Phi vg otic’ Moldenke in Chittenden, ort. oy Dict. Gard. h: —-2210. 19513; Rambo, Sellowia 7: 60 & 288. 1956. Additional citations: BRAZIL: Minas Gerais: Henschen 1.326 poet 8 1855] (W—2096L3); Regnell 1.326 (Caldas, 5/2/1305) (w— ny ay ¢ Additional biblioceriet erie) apap 3 si wots Specimens of this species have also been misidentified and dis- tributed in herbaria as V. ciliata var. pubera (Greene) Perry, v. hastata L., and V. pinnatifida Lam. The Wooton 642 cited below was was identified originally as "V. bipinnatifida - x wrightii", The A. H. Howell 159.1 cited below is a mixture with V, bracteata Lag. & Rodr, The Mearns 2677, also cited below, is annotated by Per erry as "atypical", while Kearney says of it Owhy not V. gooddingiion Moore & Steyermark 3622 is annotated by Perry as V. bipinnatifida var, latilobata Perry. In the United States National Herbarium she has annotated 0. B. Metcalfe 1231 and Wooton s.n. [Ojo Calien- te, July 18, 190k) as Ve ambrosifolia, but 1 see no no glands on then, so I am regarding than as f. Trisndnek Perry. The Clawson 13923 and 0. B, Metcalfe 1090 annotated by her as V. wrightii A. Gray and so © cited by her (1933) are V. ambrosifolia, at least a sofar as the United States National Herbarium specimens are cerned, as is also Oo. Be Metcalfe 126. The Wooton 364 cited nti fae and by me as V, wri wrightii may be a mixture, because the United Pape: National a and Britton Herbarium specimens are def- the pele V. ambrosifolia, T 12i51 & 12h5la, Geyer 107, A. Nelson 82h5, and Rose & Painter Loy 6427, distributed as V, ambrosifolia in the United States National Herbarium, are all V. bipinnatifida Nutt.; S| L. Fisher 319, Shreve 8797, and Wynd & Mueller 150 are V.c pentay Es We Nels Nelson bhoh & s.n. (May 1698) are ots f , aie 8 _— M. Chamber erlain 5 is V. ooddingii Briq.; Ge a a sv. gooddingii var. newiteteliz sidestr ,» While the | Eggleston 10737, annotated by Perry as V. ambrosifolia, is actually V. bipin- hatifida var, latilobata Perry. The Kellogg specimen cited below was originally distributed as 398 PHYTOLOGIA Vol. 8, no. 8 V. tenera Spreng. The C. Thomas s.n. [Colorado Terr., 1869] col- lection is a mixture with V. bracteata Lag. & Rodr. The sheet of E. W. Nelson 6189 in the United States National Herbarium bears this number on a tag attached to the plant and also on the label, but someone — changed it on the label to read "6789". V. ambrosifolia has been collected on sandy loam bottoms and in fertile alluvial soil of the coastal plain in cg Additional citations: pee Denver Co.: J. Coulter s.n. Pe - Cowen 03 [Foothills] ST (e201) Ce Thomas s.n. Fectseats tot 1869] (W—71930 30). TEXAS: Bexar Co.: Havard | se [San Antonio] (W—155939). Brewster Co.: ‘he R. Moldenke 171 (Fe)3 7.126 (W—-16792h9). Culberson Co.: “Moore & S & Steyermark 3622 (W—1566713). Dallam Co.: A. H. Howell 140 (W—-500265) . Hudspeth | Coe: A. R. Moldenke 162 (Fg), 163 (Fg), 16h (Fg, Z). Jeff Davis Co.: A. Re Moldenke 165 (Fg, Z, Z)» Presidio Co.: A. R. Moldenke 167 (Fe, 2). Ward Co.: Clawson 13923 (W-1085078). NEW m MEEIOO® rnalillo Co.: Harward 9 (W—370690), 14 (W--370695); Ros Fiteh 17801 (W— 760880). hi eo ae ' & Earle 322 (oso Seteiioe 32th (W—L96289). Colfax Co.: Gr ai Feiths 5515 (W--196705). O.? A. R. Moldenke 132 (Fg); Wooton 6h2 Be an). Eddy ee P. oe Standley 40334 (1 (W--1221731) ; Wooton s.n. [Queen, Aug. 2, 1909] (W—736222). Grant Co.: 0. B. Metcalfe 126 (W— 495213). Lincoln Co.: A. R. Moldenke 130 (Fg, Z)3 Wooton 363 (W—- 330588), 36 (N, W—330589); Wooton & Standley 3635 (Ww—-562150), 3691 (W—561431). Otero Co.: Wooton Sen. n. [Hess Ranch, Aug. h, poe (W--56)652). Quay Co.: G. L. Fisher 89 (W—-660028) ; ae Moldenke 123 (Fg). Rio Arriba Co.: Eggleston 5957 oe ’ 91 (W—660685), 661) (W--660857) oa he Cost ailey 529 (W360), $36 (W—Wl36L7); P. c. standl ost (r=198973)- Santa Fe Co.: Wooton 2832 (W—736802), 2033 (W--736801), S.De (Glorieta, aug. 2), 1910] (W—736885). Sierra Co.: 0. B. Metcalfe ae (w—L98278). Socorro Co.: C. L. Herrick 513 (W—737261)3 Ge R. Vasey s.n. (Socorro, May 1801] (W--155910), s.n. (Socorro, 1881] (W). Torrance Co.: G. L. Fisher 18 (W—562733); A» R. Mol- denke 128 (Fg). Union Co.: A. He Howell T 159.1, in part (W— pene C. Standley 6087 (#--605123)- county reir 86 (W). ARIZON. ARIZONA: Apache Co.: Peebles & Smith 13972 (W— EE Coconino Co,: age Si s.n. [Little Colorado, ado, 1869 (W—-56179). Navajo Co.: L. F. Ward 8.n, [Two miles south of Hol- 1962 Moldenke, Monograph of Verbena 399 brook, June 17, 1901] (W—10320) » Sen. (Woodruff, June 20, 1901] (W—1,10319) ; Zuck sen. [Holbrook, Aug. 10, 1896] (W—66),1,66) , 8. n. (Holbrook, Aug. 16, 1896] (W—66,65), son. (Taylor, Aug. 1, 1897] (W—3)9023), Sen. (Holbrook, Oct. 9, 1897] (W—-66hL6)) . Pima Co.: Mearns 2677 (W—2344h6). MEXICO: Chihuahua: H. S. Gent- 2y 7998 (W--1978779). Coahuila: E. W. Nelson 6139 (W—3h7hi8); Pringle 13157 (W—~6175); Wynd & Mueller 572 (W—1639970). Nuevo Leén: Heard, Webster, & Barkley 14512 (W—-213)281); Edw. Palmer 1050 (W—-1323139). San Luis Potosi: G. L. Lundell 6290 (i 1638806). Sonora: H. S. Gentry 7971 (W-—-1978765). CULTIVATED: pi J. He Kellogg s.n. [Mo. Bot, Gard., Aug. 25, 1911) (W— 1325356). VERBENA AMBROSIFOLIA f. EGLANDULOSA Perry Additional bibliography: Moldenke, Résumé Suppl. 5: . 1962, hrey found this plant in yellow pine forests. Additional citations: TEXAS: Deaf Smith Co.: A. R. Moldenke 120 (Fg). El Paso Co.: F, Wagner 97k (W—228855). NEW MEXICO: Dona Ana Co,: A. R. Moldenke 133 (Fg); P. C. Standley s.n. [mesa just west of Organ Mts., June 13, 1906] (W-560976); Wooton Sn. {Organ, July 1h, 1897] (W—73688h). Grant Co.: Holzinger s.n. (Santa Rita, Aug. 1, 1911] (W—-660557); Mearns 2278 (W-~233966). Hidalgo Co.s Mearns 21,86 (W—234697); A. R. Moldenke 143 (Fg), ill (am, Fg). Sierra Co.: 0. B. Metcalfe 1231 (W—I96301); A. R. Moldenke 159 (Z). Socorro Co.: A. R. Moldenke 158 (Fg, 2, Z); Hooton 2835 (W—736933). ARIZONA: Cochise 0o.: J. I. Carlson s. nh. (Douglas, May 13, 1915] (W—~-873379, W—882856); Cazier s.n. mi. E, Portal, Aug. 27, 1955] (Am); Herb. Arizona State Coll. S.n, [near Southwestern Research Station, May '57] (am); Herb. Chiricahua Veg.-Study Team 1958-118 (Am), 1959-30 (Am); A. R. Moldenke 13 (Fg, Z, Z), 136 (Am, Fg), 139 (Am), 2L0 (Am). Greenlee Co.: R. R. Humphrey s.n. [9/2/3L] (W—1697613). Pima Co.: Toumey s.n. [Santa Catalina Mts., Sept. 15, 1896] (Ww— 61918L), s.n. [Santa Catalina Mts., Sept. 20, 1896] (W—Ll1757). Yavapai Co.: Fulton 9672 (W—162378). County undetermined: Edw. Palmer s.n, (Arizona, 1869] (W—56218). MEXICO: Sonora: Mearns 1138 (W#—2291,08) . |_,Waterial of this form has also been misidentified and dis- tributed in herbaria as V. bracteata Lag. & Rodr., V. ciliata var. pubera (Greene) Perry, and V. gooddingii Briq. The 0. B. Metcalfe 1231 and Wooton s.n. [Ojo Caliente, July 18, 190h] were regarded by Perry and annotated by her as V. ambrosifolia Rydb., but I see no glands and so I regard them as f. e dulosa. Of at S.n. [Santa Catalina Mts., Sept. 20, 1896), Edw. Palmer S:n. (Arizona, 1869], and one sheet of J. I. Carlson s.n, [Doug- 00 PHYTOLOGIA Vol. 8, no. 8 las, May 13, 1915] in the United States National Herbarium Kear- ney’ says "Why no da? Bracts equalling or nearly e- qualling the a t e af Toumey s.n. [Santa Catalina Mts., Sept. 15, 1896] he says et bipinnatifida. Bracts nearly equalling calyx. Anorfoschsa] ve ciliata in other characters." Of another sheet of J. I. Carlson s.n. (Douglas, May 13, 1915] he says "V. ciliata? glandular lar pubescencet", but I fail to see the glands; 3 of R. Re R. R. Humphrey SNe ae he says "V, bipinnatifida?" I regard all these ese specimens - eglandulosa The F, Wagner oh ¢ cited above from ‘nl Paso County, Texas, may pee actually have come from that county, as its label is inscrib- ed "El Paso to Monument 53". AMOENA Paxt. Additional citations: MEXICO: México: Pringle 13434 (W--l62365). VERBENA ANDALGALENSIS Moldenke Pago Sin Tiecvecty: Voldenke, Alph. List Cit. 2: 599. 19h8. e JUrgensen 1613 collection is incorrectly cited 1988 publication as V. dissecta Willd. 1613 ( Additional citations: ARGENTINA: Catamarca: JUrgensen 1613 922058—-isotype) . VERBENA ANDRIEUXII Schau. This species has been collected at altitudes of 5200 to 6800 feet, flowering and fruiting in October Additional citations: MEXICO: Oaxaca: E. W. Nelson 1513 (w— VERBENA ARISTIGERA S. Moore and Herbarium material of this species has been misidentif ied distributed as V, erinoides mies and as V. mit ifida Ruiz & Pav. Additional citations: : Chaco: Venturi 9780 (W— 1591421). Entre Rfos: ribo s 1053 * (W-1233656) « CULTIVATED: Ge any: Herb. Mus, Bot. Berol. s.n. [180] (W—5)6086) . VERBENA ATACAMENSIS Reiche A 4 i dditional citations: CHILE: Antofagasta: Werdermann 789 (W#— 474,301). ge EB AURANTIACA Speg. Additional citations: ARGENTINA: Neuquen: Senn 326 (W--203433)- VERBENA BAILEYANA Moldenk Additional mah liseventei H. E. Richter, Cod. » 33° 1835; H. F. Roberts, Plant Hybrid. before ‘Mendel 28. 19295 uo}s 19513 ke in Chittenden, Roy. Hort. Soc. Dict. Gard. h: 2209 & 2210. “77% a e Richter reference given above it is stated "In Diss. dé 1962 Moldenke, Monograph of Verbena 401 Pl. hybrid. (Am. acad. tom. 3. p. 43) commemor. planta inter hanc et V. spuriam enata, solis fol. multifido-laciniatis ab hastata diversa, nom.: 'V. tetrandra, spicis longis acum., fol. multif.- lac.' mox periit, hinc ignota." VERBENA BAJACALIFORNICA Moldenke Additional citations: MEXICO: Baja California: Shreve 7169 (W- 1749073——isotype) . < MRT eee BALANSAE Briq. ; Additional bibliography: Fedde in Just, Bot. Jahresber. 33 (1): 32. 1906. Additional citations: PARAGUAY: J&rgensen )582 (W—-1511008) . ARGENTINA: Corrientes: Pedersen 2906 (W—-2283152). VERBENA BALLSII Moldenke onal citations: ARGENTINA: Jujuy: Balls B.603% [U. S. Nat. Additi ENT Arb. 163273] (W—22502h3—isotype) . XVERBENA BEALEI Moldenke Emended bibliography: Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. 4: 2210. 1951. VERBENA BERTERII (Meisn.) Schau. Additional and emended bibliography: Moldenke in Chittenden, Shae? Soc. Dict. Gard. : 2209 & 2210. 1951; Moldenke, Résumé fA alr ge this species has been misidentified and distributed as V. dissecta Willd. and as V. lipozygioides Walp. Additional citations: CHILE: Santiago: G. T. Hastings 190 (W— 530087); R. A. Philippi 1298 (W--1323088); Wilkes s.n. [Between Valparaiso and San Diego] (W—-71950). Valparaiso: Buchtien s.n. (8.1X.1895] (W--1159365); Claude-Joseph 3548 (W--1283379); Killip & Pisano 39710 (W—1953999). Province undetermined: Claude-Joseph Lol3 TBureo) (W—-1281313), 50 [Vicufla] (W—1283608). VERBENA BIPINNATIFIDA Nutt. Additional synonymy: Verbena bipinnata Nutt., in herb. Additional and emended bibliography: Moldenke in Chittenden, Roy. Hort. Soc, Dict. Gard. : 2209 & 2210. 1951; Shinners, Spring Fl. Dallas 325. 1958; Moldenke, Résumé Suppl. : 21 (19625) and 5: 3 & h. 1962; Pearce Seeds & Plants, Our Am. Wild Fls. 10. 1962. This species has been collected on ballast about eastern ports in the United States. The Pearce horticultural supply company de- Scribes the species as an "ever-blooming annual", The corolla is de- Scribed as "deep-pink" by Kearney & Peebles. The Geyer 107 cited be- low has been previously identified as V. bracteoga Michx., V. auble- ta L., and V. ambrosifolia Rydb.t The E. Hall U3 collection is a mixture with V. pumila Rydb. and was identified by the collector as ¥. bracteosa-aubletia. 02 PHYTOLOGIA Vol. 8, no. 8 The Earle & Earle 322, Eastwood 140, Fendler 586, G. Le Fisher 18, Griffiths “yh, Harward 9 & ih, Cc. L. ~ Herrick $13, P P. Standley 4951, “Tweedy 102, Wooton nm 363, 2832, 2833, and s.n. wren ieta, Aug. . 2b, 1910], and Zuck s.n. . [Holbrook, Aug. 10, “1896], distributed as V. bipinnatifida, ar are all V. aubropifolsa Rydb., as is also Wooton 642, distributed as "verbena bipinnatifida x wrightii". The Cazier s.n. (6 mi, E. Portal, Aug. 27, 1955], Mearns 2186, 0. B. Metcalfe 1231, P. C. stand s.n. [mesa just west of Organ Mts., June ~June 13, 1906), and Wooton s.n. (Organ, July 1, 1897] are all V. ambrosifolia f. eglandulosa Perry, while Rose & Fitch 17106 is V. bracteata Lag. & Rodr.; Waugh 160 168 is V. canadensis (L.) Britton; Maysilles 7039 7039 is V. ciliata Benth.; and Zuck s.n. (Tucson, May 16, 1896] are all V. gooddingii Briq.5 Lindheimer s.n. [Texas] and ce T. Mohr s.n, (N. Orleans] are V. halei Small; Kearney & Peebles 9602 and E. W. Nelson 4856 are Ve bipinnatifida var. latilobata Perry; G. pee Fisher 207 & “& 223, May- silles 768), Edw. Palmer 7, and Pringle 1117 are V. V. ciliata Benth.; J. K. Small s.n. [near Wharton, April 12, 1925] is V. ciliata var. longidentata Perry; and Mearns 109, F. G. Plumes s.n. [Lincoln National Forest, 1903], and “and Wooton £285 1. [Divide a- bove Mescalero Agency, June 23, 1895] & s.n. n. [White te Sands, Aug. 25, sett are V. wrightii A. Gray. earney says 3 of Toum oumey S.n. [Santa Catalina Mts., Sept. 20 1896] and Edw, Palmer s.n. (Arizona, 1669] in the United states National Herbarium "Why not V. bipinnatifida? Bracts nearly equalling calyx", but I regard both collections as V. ambrosifol- TS aes ee ia f. eglandulosa Perry. He also says of Toumey Sele , [Santa Cata- Tina Mts., Sept. 15, 1896] in the same herbarium "V. bipinnati- fida. planta nearly equalling calyx. Approaching] Ve V. ciliata in glas 1915] "V. bipinnatifide?" ey eon both fenesatea as Ve ambrosifolia f. seen The te a & & Mueller 450 cited by me in Phytologia 8: 251 0 ‘ation will probably prove to be V. ciliata, as are the REE pa es 7039 | & 7057 there cited. Additional citations: ALABAMA: Dallas Co.: R. M. Harper 19 (W —1365890). MISSISSIPPI: Oktibbeha Co.: Weatherby & , & Weather’ 6313 (W—-160087). INDIANA: Clark Co.: Umbach s.n. ~ (Clarke, June 29, 1898] (W—~339478). SOUTH DAKOTA: Fall River co.: Rydberg 938 (W—210967). Iyman Co.: Over 3177 (W-582996). Stanley Cos G&S 107 (W). Washington Co,: W. H. Over 2103 (W—582936) « KANSAS: Barber Co.: B. B. Smyth 306 (W). Douglas Co.: W. H. Horr 1,368 (FF 1962 Moldenke, Monograph of Verbena 4,03 2097550); Re L. McGregor 12211 (W—2230321). Ellis Co.: A. S. Hitchcock 393 (W—-353120). Meade Co.: W. H. Horr 348 (W"—-2030691). Morton Co.: Carleton 177 (W, W); Rose & ey ine (W-~-6392h)9). Osborne Co.: Shear 38 (Ww (W—227316) , Riley Co.t Ae Nelson 8245 (Ww— 48579). Scott Co.: F earing & Latham s.n. [August 1h, 1950) (W— 2030815). Smith Co.: W. H. Horr E.108 (W--2031069). Wabaunsee Co.: Bartley 12,0 (W—1973587). MISSOURI: Jackson Co.: B. F. Bush 1,029 (W--606119). ARKANSAS: Hempstead Co.: E. J. Palmer 1531506). LOUISIANA: Rapides Par.: C. R. Ball 401 (1 38275h) . Saint Mary Par.: Langlois s.n. [Cotes Blanches, 1 17.VII. 1893] (W- 1465962). NEBRASKA: Boyd Co.: | F. E. Clements 2776 (W). Sherman Co: J. M. Winter 75 (W—1679019). OKLAHOMA: eee Co.: Demaree 1251 (W—168),883), 12h5la (W—168)88)). Bryan Co.: C. S. Sheldon (W). Comanche Co.: H : Hopkins, Nelson, & Nelson 801 Taniseress: Waterfall 13097 (W--2260931). Custer Co.: A. Re Moldenke 115 (Fe). Harper Co.: G. 1 G. W. Stevens 305 (W-~589065) . Kingfisher Co.: ? Blan- kinship s.n. (Huntsville, Apr. 16, 1896] (W—313601). Lamar Co.: Demaree 12003 (W—1683782) . nerat Co.: J. F. Brenckle 48065 (W— 2003822) ; Merrill & Hagan 303 (W—1653007); “G, W. Stevens 1h (Ww 588951). Woods Co.: G. W. Stevens 699 (W—589223). TEXAS: Bell Co.: L. F. Ward s.n, ~ (Killeen, October 1, 1891] (W—~1L7559); s. E. Wolff B76 (Woe (W--162),007). Bexar Co.: C. Re Ball 909 (W--501971) ; Groth 30 (W—71761,9) ; G. Jermy 86 (W—315685), sen. [S. Antonio] (W—392397); Larrabee s.n. [Alamo Heights, May 1, I, 1900] (Ww anges Mrs, H. D. Slater s.n. [San Antonio, 1918] (W~891772). Blanco Co.; Lundell & Lundell 1027) (W--1888919). Brazoria Cos: B. F, Bush 206 (W—386LL.7), LL (W--3865LL). Comal Co.: G. Le Fisher 2 g = 11ch890) 5 Lindheimer 1072 (W—502LL5), 1073 (W--SO2L]6). : Iundell & Lundell 10416 (W—1887677), 10568 (W— 1836956). 11315 [Plant. Exsice. Gray. 1275] (W—1925785); J. Rev- crchen 739 (W—117561, W), sen. [Curtiss 1962*] (W—1323050, W); M. B. Stephenson s.n. [vicinity of Dallas, Apr. 2, 1929] (W— Tass DeWitt Co.: A. H. Howell 308 (W-S01L57)« Fisher Co.: I. M. Shepherd s.n. (Grady, April 1h, 1901] (W—L13915). Frio Co.: Rose & Painter 6427 (W—L51706). Guadelupe Co.: F, S. Earle 430 (W—Ts0901). Hanphill Co.: A. H. Howell 111 tSEE Ds tho Hood Coe: E. J. Palmer 6510 (W~128101h). Kaufman Co.: F. J. Tyler s. aR [vicinity of terres May h, 190] (W-—-500802) . “Kerr Co.: Mun- aon & Hopkins s.n, [Ke errville, Jul. 1889] -(W--219990). Kinney Co.: Wearns 1252 ( 1252 (25189), 1274 (W—229520), 139 (W--2300K2). Kle- berg Co.: G. J. Harrison san. (Riviera, 2/24/30] (W—15323L0). Liberty cont iT ane & Wherry 11781 (W—~1738883) « Navarro Co,: Joor 107 (W); E. J. Palmer (W—I531827). Nueces Co.: Marlatt S.n. TMtecestown, May 27, 1856] (W—-266973) + Oldham Co.: C, L, Lundell To) PHYTOLOGIA Vol. 8, no. 8 11,62 (W—19269,2). Parker Co.: S. M. Tracy 7999 (W--500368) . Tarrant Co.: Killian 6779 (W—1L65227); McKelvey 1679 (W—1581392); Ruth 107 (W--501767, W—-1488932); L. F. Ward s.n. [Fort Worth, Sept. 9, 1877] (W—1h7557). Taylor Co.: S. M. Tracy 8000 (W-— bh2h52). Tom Green Co.: Cory 50566 (W-—-1586002); Tweedy s.n. [Kni- ckerbocker Ranch, May 1880] ( —1)7562). Travis Co.: E. Hall 431, part (W). Uvalde Co.: J. M. Milligan s.n- (Uvalde, April) (W— 50,333). Val Verde Co.: G. L. Fisher 85 (W—1366096); Wooton s.n- [Del Rio, Nov. 5, 1913] (W--662058). Van Zandt Co.: J. B. Hender- son 5698 (W—168822). Victoria Co.: Fe Le Lewton 60 (W); Maxon 3808 (W--50033); E. J. Palmer 9098 (W—153107%h). Wichita Co.: Whitehouse 9589 (Mi), 9609 (Mi). County undetermined: Lindheimer Tir .289 (W—197100), W). NEW MEXICO: Chaves Co.: Earle & Earle 526, in part (W—382588). County undetermined: C. Wright 1502 (W— 56225). ARIZONA: Apache Co.: Edw. Palmer s.n. [Willow Spring, May 1890] (W--56212). Cochise Co.: Blumer 135 [108] (W—561617, W— 563255); Kearney & Pecbles 1423 (W—1760656); H. N. Moldenke 21880 (Am, Fg); Ordway s.n. [Rustler Park, July 30, 1955] (Am). Coconino Co.: Kearney & Peebles 13983 (W—1732901); MacDougal 317 (W—33k412); Mrs. C. D. Walcott s.n. [Grand Canyon, Aug. 1938] rc em eter nnmnarticne (W—173090L). Gila Co.: Collam 248 (W—1683284). Graham Co.: DAE” rison, & K Leh9 (W—136800l). MEXICO: Chihuahua: E. W. Neo son 6117 (W—359829); Townsend & Barber 139 (W—383221). MOUNTED DESCRIPTIONS: Am, Midl, Nat. 2h: 752. 1940 (W). row, Phillips, & Pultz Peebles (W--1732962); Peebles 12989 (W-—-1679125). Pima Co.: Peebles, Har- W. VERBENA BIPINNATIFIDA var. LATILOBATA Perry Emended bibliography: Moldenke in Chittenden, Roy. Hort. Soc Dict. Gard. 4: 2209 & 2210. 1951. The Moore & Steyermark 3622 annotated as this variety in the United States National Herbarium by Perry is actually V. ambrosi- folia Rydb. On the other hand, of the Eggleston 10737 identified by her in the same herbarium as V. ambrosifolia Kearney says "I make this [to be] V. bipinnatifida var. latilobata" and I agree with him! C, Wright 1503 in the same herbarium was misidentified as V. ambrosifolia. eee Additional citations: ARIZONA: Cochise Co.: Eggleston 10737 (W-664802); Griffiths 4833 (W—L96451); Herb. Chiricahua Veg-— Study Team 1999-lh2 (Am); A» R. Moldenke 138 (Am), 145 (Am)j OF way s.n. [Rustler Park, July 30, 1955) (Am); Te E- Wilcox SDs {Ja Co.: Peebles & Smith 14093 (W-17329l3). Graham Co.: Ke & Peebles 9802 (W- 162yh16). Pima Co.: Griffiths & Thornber 159 (W-—W97223) ME: Jones s.n, [Santa Rita Mte., Aug. 2h, 1903] (W--856989) « Sante 1962 Moldenke, Monograph of Verbena 0S Cruz Co.: Kearney & Peebles 10130 (W—162),502). MEXICO: Chihuahua: E. W. Nelson on 856 (W--332903); C. C. Wright 1503, in part (W—71927). Michoac4n: Hinton 13125 (W—1891037). Sonora: Hartman 906 (W— 18925),2—-isotype). XVERBENA BLANCHARDI Moldenke nded synonymy: Verbena blanchardii Moldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. Gard. hz 2209 & 2210, sphalm. 1951. Emended bibliography: rte in Chittenden, Roy. Hort. Soc. Dict. ae ard. h: 2209 & 51. E. W. Lathrop 3 & L88, distributed as xV. blanchardi, are ieee xVerbena ras Moldenke, VERBENA BONARIENSIS L. Additional synonymy: Verbena bonaerensis L., . a Additional and emended bibliography: C. Murr ur. 9975 Spreng. in L., Syst. Veg., ed. 16, 2: a8. “ia2ss 1 ihe ®,s Fic Austral. G3 y— 37. 1870; Ee H. L. Krause turm. nh. pene. 15: 305. 1907; Moldenke in lygeseeyes ". Hort. Soc, Dict, Bard. : 2208—2210. 1)Ls Rambo owia 7 les Moldenke, Résumé Suppl. : 1—5, 1h, is. & 19 (1962) and 5: ahaa “4l.ustratdons: Moldenke in Chittenden, Roy. Hort. Soc. Dict, Gard, h: 0. 1 An additional — name is "cluster-flower'd vervain". Th species was apparently already in cultivation Aly Eng in 1799 according to the Murray reference cited above. It has been collected on dry hillsides in Jamaica For those students interested in noting the rapidity rp ob spread spec of introduced plants, the following date en this es was first collected in a given county(as far as I an n be o North Caro. Bladen, C k, and New Hanover c houn County - 1947; Alabama: Tuscaloosa County ~ 1938; Mississippi: Ra County - 192 - Arkansas: Saint Francis C , Ashley and Drew Counties - 1938, B ey County - Bermuda - 1921. The E, Carter 91, E. S. Ford 3785, Godfrey & Shunk 4072, and L. S. Rose 39200, distributed as Ve oe are all V. brasilien- sis Vel Vell., as are also Os Novaes 916, Ae arvalho s.n. {Herb. Inst, Bot. S. Paulo 190], Claude—Joseph oh 1286, eke “Dusén 14320, Julio 2, Mexia 5275, T. Meyer "12hs, Miers s.n. \ Vaenional,, and Semper 293. The Buchtien 11683 | and Cardenas 327, distributed as V. bonariensis, are V. hispi hispida rp & Pave; G. L. L. Fisher 168, A. A. Heller 20h6, NT ead Hosaka 1354, and Mendes Magalhfes 3151 are ev. litoralis H.B.k.; and 406 PHYTOLOGIA Vol. 8, no. 8 Barkelew [not "Barkeley" as stated in Phytologia 8: 257. 1962] 232 an aes 0. Solis 20 are V. sphaerocarpa Perry. ham (1870 0) cites the following additional specimens not as Ei a wg me: AUSTRALIA: New South Wales: Beckler s.n. (Hastings river]; F. Mueller s.n. [near Sydney]; C. Stuart s.n- . [New Eng- land]. Queensland: F. if ueller s.n. {Brusb sbane river, Moreton Bay]. Additional citations: NORTH CAROLINA: Currituck Co.: We Be Fox s.n. (1/2 mi. n. of Point Harbor] (W--22651,67). Wayne Co.t Ae Re Moldenke 163 (Fg ). SOUTH CAROLINA: Charleston Co.: Curtiss 1963+ (W), s.n. (Charleston, 1875] (W); L. R. Gibbs sn. (W—-203816, W- Ww 203817); W. Palmer s.n. token teeta; June 2--10, 1902] (W—513666) « Columbus Co.: Ae + A. R. Moldenke Moldenke 452 (Fg). Florence Co.: A. R. Molden- ke 453 (Fg). Georgetown Co.: Godfrey & Tryon 268 (W--1337175) - Horry Co.: A. R. Moldenke 45k (Fg) - Kershaw Co.? A. Re Moldenke bh3 (Fg). Lee Co.: A. R. Moldenke 6 (Fg). Lexington Co! rey ey Voldenke 3 (Fg). GEORGIA: Bibb Co.: A. Re Moldenke 371 (Fg) - Bleckley Co.: A. R. Moldenke 379 (Fg). “Brooks Co.: A. Re Moldenke 308 (Fg), 312 (Fg). Burke Co.: A. R. Moldenke 406 (Fe). shoo Co.: R. F. Thorne 3659 (W--1929865). Grady Co.: 3 A. R. Molden 298 (Fe g). Jefferson Co.: A. R. Moldenke 405 (Fg)- . Lanier 5 = = R. Moldenke 327 (Fg). Monroe Co.: A. R. M Moldenke 387 (Fg). Scre- m Co.: A. R. Moldenke Lats) (Fg). FLORIDA: Bradford Co.: Murr 53h (W--192835h). Liberty Co.: A. R. Moldenke 278 (Fg). Marion Co: Gist 105 (W--1653820) . ALABAMA: Tuscaloosa Co.: R. M. Harp- er 3672 (W--1731623). MISSISSIPPI: Madison Co.: McDougall 1 1383 TW-—192487k). LOUISIANA: Ouachita Par.: J. A. Drushel 9478 (W-- 1654135). Saint Tammany Par.: Arséne 11859 (W--103187 3), yy L253 (W--122296). TEXAS: Angelina Co.: Tharp 2558 (W--1221348). Har ris Co.: A. R. Moldenke 216 (Fg). Jefferson Co-: Je F. Brenckle 18071 (W—2003825); A. R. Moldenke 222 (Fg). CALIFORNIA: Marin Coes Cot Je T. Howell 19323 . (W—187259) . Yuba Co.: A. A. Heller 19162 (W--197)207) BERMUDA ISLANDS: Main: Brown & Britton 98 (W—-13h1080); F. S. Collins 266 (W--717560). JAMAICA: Je He , Hart 3.n. (W--1323053)3 Killip 307 (W--105096); Maxon 2615 (W-- 52058); Maxon & millip 30 69 (W—106299); C. Re Oreutt 2736 (W- 141166) . : Minas Gerais: F.C. Hoehne Sn. . (Herb. . inst. Bot. S. Paulo 2912] (N); Kuntze s.n. , {Contendos, Dec. '92] (N)5 Regnell II.210 [Caldas] (N), 11.210 210 [1843] (W--209641). Rio Janeiro: Holway & Holway 1160 (W--118822h), 15) (W--119929h) 5 Lund s.n, tel {ys Rio Grande do Sul: Rambo Silshh (W--2102030) So Paulo: F, C. Hoehne s.n. (Herb, Inst. Bot. S- Paulo 1361) (N); Usteri s.n. dicta Esc. Polytec. 25ib; Herb. Inst. Bot. S- Paulo 1572] (N). BOLIVIA: Santa Cruz: C&rdenas 5130 (W-~ 225068) . PARADA. 2 Orgensen 3768 (W-1183025). URUGUAY: Herter 1962 Moldenke, Monograph of Verbena 407 268 (Herb. Herter 81709] (N, en etart torent z om (W—-1233689). CHILE: Curicé: Herb Herb. Mus. Nac. Santiago ) ( (N). Lu quihue: Gunck- el 16865 (W--1997763). Valdivia: Sasnetan s.n. REET 1898] (W--1159 368); 3 H. Krause s.n. [Coral] (W--1690233). ARGENTINA: Buenos Aires: Brizuela 159 (W--2198122). Chaco: Venturi 8), 1043709). Corrientes: Pedersen 1333 (W—-2283062). Misiones: Ped- ersen 43hl (W--2283778). Tucum4n: Venturi 123 (W--1591212), 525 (W--1591223) , 2936 (W--1591273). AZORES ISLANDS: Faial: Ce Ba Brown 215 (W--262278). REPUBLIC OF SOUTH AFRICA: Natal: Sidey 2058 8 (S). Transvaal: Schlieben 7176 (W--2272136). INDIA: Uttar Pradesh: R. R. Stewart 16007 (W--1992151). State undetermined: C. Be Clarke 24)76G [D nites (W—802905). MELANESIA: New Cal- edonian Islands: Ile des Pins: M. R. Germain s.n. [U. S. Nat. Arb 218555] (W--2185103). we Caledonia: Franc A.1693 [U. S. Nat. Arb. 218556] (W—-2185088). POLYNESIA: Austral Islands: Saint Mary's: Eights s.n. (W--920098). VERBENA BONARIENSIS f. ALBIFLORA Moldenk oe bibliography: Moldenke, Résuné Suppl. 4: 5 & 6. 962 VERBENA BONARIENSIS var. CONGLOMERATA Bri Additional bibliography: Moldenke, Résum eo Additional citations: PARAGUAY: cuogecete 37h fe asbhorlo. ° VERBENA BRACTEATA Lag. & Rodr. Additional synonymy: Verbena bracteosa brevibracteata Gray, fa ———_—— in Additional and emended beta Spreng. in L., oom As Pa ed. a te re 749. fine Moldenke in eaperig om Hort. Dict. 209 & 2210. 1951; pore ers, 325. 1955) ‘woldeake, Résumé Suppl. hs aM wee S% (2962) and 7 1 5&7 . Torr. Bot. Club 89: 56. S.n. [near Southwestern Research Station, 4 lay ' a distributed as as v. bracteata, is actually V, ambrosifolia f. eglandulosa Perry, While Pataky SNe Sescby Huachuca, 1090] is V. gracilis Desf.; Rugel s.n, [ad vias et in incultis arenosis, per Alabamam median, Sept. 18]3] is v. caliccetn (L.) Britton; E. W. Hammond 322 is V. lasiostachys var. septentrionalis Moldenke; “and J. Le! Je Let 3. Sen. [San Diego, Nov. 9] 4s V. menthaefolia Benth. The A. H. Howell 159.1 & 159.2 and C. Thomas s.n. [Colorado Terr., 1869] collections are mixtures with ve ambrosifolia Rydb. The H. R. Bennett s.n. [Brewster Arnold Ranch, August 2, 1957] specimen cited below bears a notation, probably by the collector, "evidently a hybrid with V. stricta Vent. or some other species” but I do not agree -- to me it seems to be typical V. bracteata. 1,08 PHYTOLOGIA Vol. 8, no. 8 A common name for the species in Mexico is "chichiro", Lehr (1962) records V. bracteata from Rockland County, New York. additional citations: ALBERTA: Macleod Co.: BE. He Moss 20) (W- 1896368). BRITISH COLUMBIA: Cariboo Co.: J. Macoun SNe {Kamloops, Jone 13th 1889] (W--219610). Yale Co.: Calder & Savile 9668 (W— 2310062). NEW JERSEY: Hudson Co.: Van Sickle s.n, [Weehawken, July 20, '9h] (W-30969), sn. (Weehawken, Aug. 1891] (W--2))3276)- (W--,07053). Philadelphia Co.: Martindale s-n» [Phila., 8/78] (We 1756). VIRGINIA: Southampton Co.: A. A. Heller 1012 (W). NORTH CAROLINA: Iredell Co.: Ruger s.n. [Statesville] (W-—310734). Coun- ty undetermined: Ashe s.n. [Southeastern N. Cel (W——3278L9) , GEORGIA: DeKalb Co.: J. K. Small s.n. [Stone Mtn., July 1—6, 1895] (W--298966). Gwinnett Co.: H. A. Allard 207 (W--513129) « Lee Coo: J. De Smith 151 (W--1323056), 1961 (W—-1323057) » FLORIDA: Franklin Co.: Biltmore Herb. 1082b (W—332102). ALABAMA: DeKalb Co.: Ruth 501 (W—345682). Jackson Co.: Le Fe Ward s.n. [Steven~ son, Sept. 5, 1877] (W—1h756h). Lee Co.: Earle & Baker sen. [Au- burn, 5/13/1898] (W—3l4792). Mobile Co.: C. T. Mohr son. [Mobile, May 22, 1884] (W--77188h). Morgan Co.: J. D. Smith 152 (W--1323055). Tuscaloosa Co.: Pollard & Maxon 333 (w——L2883) . ILLINOIS: Cook Co.: H. H. Babcock s.n. [Chicago] (W--787353)- Menard Co-: Ee Hall s.n, [Athens, Sept. 1668] (W). St. Clair Co.: Eggert son. [E- St. Louis, June 11, 1879] (W--751560). County undetermined: G. R. Vas~ ey s.n. [Illinois] (W-—-31)208). INDIANA: Howard Co.: Ek 16 (W— 1819280). Lake Co.: Umbach s.n. [Lake Station, June 6, 1896] (W— 339477). Steuben Co.: C. C. Deam 1272 (W—-607256) « Vigo C0.: Ever- mann s.n, [Terre Haute, June 17, 7889] (W-—-6l15927). County unde- termined: H. W. Clark 175 [Keen’s, Lake Maxinkuckee] (W—L07572) « IOWA: Fay’ ) Coe: Re Ae 2456 (W--2261079). Johnson Co.: Somes 3696 (W—67232h). Story 0°: G. Re Ball 86 (W--2802h3); H. S. Fawcett 12 (W—-L93389) « KENTUCKY: Jefferson Go.: C. W. Short sen. [borders of the Ohio below Louis- ville] (W—771885). WISCONSIN: Trempealeau Co.: Hartley & Hartley 372 (W--2261772); F. J. Hermann 8954 (W—1732399) « County undeter- mined: "J. H. S." son. [July 22, 1007] (W-—752025). M As Mille Lacs Co.: E. P. Sheldon s.n. (Princeton, July 1892] (W)- Ramsey Co.: Mearns 52 (W—6L950h). NORTH DAKOTA: Benson Co.: Lun- ell s.n. [Leeds, October 7, 1915] (W—893170). Cass Co.: O. Ae Stevens s.n. {Fargo, hug. 17, 1949) (W—1973377)- Morten ©Ee! — vis 122 (W—S8L95S) - SOUTH DAKOTA: Custer Co.: Over 1779 ag B82008). Jackson Co.: J. Fe Brenckle 39-7h (W—1026779) « Lemans Co.: W. Pe Carr 179 (W—751175); Rydberg 93k (w—210966) Heade Co. Forwood 298 (W-~317830). Washington Co.: Over 2369 (W#— 58296). KANSAS: Douglas Co.: W. H. Horr E570 (W-—-2031273)5 We Se 1962 Moldenke, Monograph of Verbena 09 Stevens s.n. [Lawrence] (W—216102). Finney Co.: Coville 2h (W—- aml Meuble s.n. [Garden City, June 7, 1891] (W-—2)1965). ant Co.: C. H. Thompson 42 (W--270168). Jewell Co.: W. H. Horr Ll Bir P07 T905) Reape Kearney Go.: A. S. Hitchcock 1129 (W—353932). n Co.: We. H. Horr 3472 (W—-2030700). Morton Co.: Rose & iten ape (W--639250). Osborne Co.: Shear 62 (W--227339), 101 (W—22731h). Riley Co.: J. B. S. Norton 392 (W-—353119). Woodson Cos: E, W. Lathrop 1276 (W—2235252). Wyandotte Co.: G. Smith s. Be (Kansas City, July 26, 1871] (W-—2311016). MISSOURI: Greene : P. C. Standley 9362. (W—688277). Saint Louis: Eggert s.n. [rate places, St. Louis, 2 July 1875] Sere ae (St. Louis, Sept. 18,1] (W). Wayne Co.: E. J. P (71581018). ARKANSAS: Craighead Co.: Demare 2 10er USN) ski Co.: Demaree 8623 (W-—1582392). is Co.: F. C. Hi i81 (W—~1651,399) « Sebastian Co.: Bigelow s.n. [Fort Smith] ww, LOUISIANA: Rapides Par.: C. R. Ball 553 (W-—382825). MONTANA: Dawson Co.: L. F. Ward s.n. (Glendive, re, July 17, 1883] (W— 147563). eis Co.: Blankinship I.410 (W—5L0L6h). ees patie Co.: P. C. Standley 1876 6 (W—1026L6 65). Jefferson Co.: R. Sn, Teetever Madison & Jefferson rivers, Sept. 7, '71] “Hi 235943). Powell Co.: E. Le Morris son. [Gold Creek, August 189)] (W—2328727). Rosebud C: Co. H. Re Bennett sen. [10 mi, east of Lame Deer, 7-25-57] (W--2300907) , son. [Brewster Arnold Ranch, August 2, 1987] (W—2300906). Sanders Co.: Sandberg, MacDougal, & Heller 972 (W--238177). Sweet Grass Co.: Eggleston 7970 (W— 5ul156). IDAHO: Ada Co.: J. A. Clark 55 (W--543356), 279 279 (W#— wciog ge 1872] (W— Sete SU3K55) » cog Co.t J. Me - Coulter s.n. - Fe iiidaretin ‘OTD =a det. Fremont Co,? Es De Merrill “Idaho Co.? eidaracin sn. 9. Jones Lewiston, June nt 19h) (w226168); Sanbeee MacDougal, & Heller 26) (W—21.2923). WYOMING: Albany Co (W—1,33812). Natrona Co.: Geodsing Et 207 (i--185108). , Niobrara Cos F. H. Knowlton 148 (W—-272918) . sper ones — Mearns 302 E.0.S. (W--488963), 2658 (W—l87268), 5009 (W— 188519). County coe enined A. Nelson 320 (W--281,505) Ape Co.: M. E. Jones 5178 (Ww—2),9911) . “Grand Co.: Rydberg Garrett 8365 (W—765012). Kane Co.: pee 2390 a ae Piute Co.: Rydberg & Carlton 70l3 “Salt ae ot S. Watson 823 (W). San Juan Co.: Rydbe rg & Gareett 9201 (W—~ 7651 on 823 (W)- Se Tidestrom 161 (W—506968); L. F. Ward “F. Ward 256 (W—11,7566, W—221200). Utah Co: Tidestrom 1735 (W—5072h2). NEVADA: Clark Co.: Alexander & Kellogg 17397 (W—1826991); Clokey 410 PHYTOLOGIA Vol. 8, no. 8 8096 (W--1785517). Lander Co.: A. E. Hitchcock 583 (W--765930) « Storey Co.: Purpus 5946 (W--38137) « County undetermined: S. Ge stokes sen. [Grand Junction, June 15, 1900] (W——lO499) « COLORADO: Archuleta Co.: C. F. Baker 56, (W— 368089). Boulder Co.: Ewan Sl. (Pl. Exsic. Gray. 1090] (W—-1828391) . Conejos Co.: P. c. Standley 6496 (W--685518). Denver Co.: Eastwood 25 (W--582271); He Ge Smith Jr. sen. [Plains near Denver, Sep. 9, 1888] (W). Eagle Co.: Killip 3u5u (W--1829030). El Paso Co.: Biltmore Herb. 1082c [1- 3k) (W—980795); Clements & Clenents 9 (W--580010); Curtiss SNe [Colorado Springs] (W-—20220). Gunnison Co.: C. Fe Baker 920 (W- 12533). La Plata Co.: Tweedy 433 (W--370LU5); Wooton 2836 (wW— 562272, W—737206). Larimer Co.: J. H. Cowen hOk (W—25)916), 2019 (W—l89193). Montrose Co.: C. F. Baker 200 (W——l)11953); E. P. Walker h02 (W—5),3789) . Otero Go»: Crandall sn. [Rocky Ford, June 13, 1891] (W—-216753); Rose & Fitch 17503 (W—760579). Wash- ington Co.: H. L. Shantz 108) (W—583884), 1200 (W—58)00)), 1229 (W—58)011). County undetermined: C. Thomas s.n. {Colorado Terre, 1869] (W—71930). NEBRASKA: Buffalo Co.: J. H. Holmes 12 (W), 20 (W). Cass Co.: T. A. Williams s.n. [Weeping Water, June 1885] (W—750381). Cedar Co.: F. E. Clements 2612 (W). Lancaster Coes Seal s.n. (Lincoln, 6-10-9h] (W--231105); H. J. Webber s-n+ {Lincoln, June 20, 1888] (W--216319); T. A- Williams sen. [(Lin- coln, June 5, 1889] (W—750382). Thomas Co.: Rydberg 1306 (W— 210371). OKLAHOMA: Beckham Co.: A. R. Moldenke 16 (Fg). Caddo Cos: Demaree 12533 (W—~16838l5). Cimarron Co.: C. M. Rogers 4668 ——— (W-—2053022). TEXAS: Deaf Smith Co.: A. R. Moldenk 21 (Fg) Donely Co.: A. R. Moldenke 119 (Fg). El Paso Co.: B. Barlow SMe [Canitallo, 7 July 1911) (W—15217k1); Dieffenderffer 90 (W— 2311831); G. R. Vasey son. [El Paso, May 1881) (W—155632)3 C- Wright 45 (W). Grady Co.: A. R. Moldenke 118 (Fg). Hale Cost Whitehouse 9926 (Mi). Howard Co.: Havard san. [w. of Big Spring, Sept. 1801] (W--155631, W—-221203). Lamb Co.: Tharp 6274 (We 164159). Lipscomb Co.: A» H. Howell 63 (W--500186). Lubbock 00: Denaree 770 (W—169002)j Es Ls Reed 3058 (W--1520199), 3291 (W- 1520198); Tharp 6284 (W--1L65165)3 Wooton s.n. (Lubbock, May 3, 1925] (W—12),608). Moore Co.: Jespersen @ Jespersen 2708 (a 2003035). Potter Co.: C. R. Ball 1675 (W—503005). Randall Cos: J. F. Baker 2 (W--SOO}1);). Tarrant Co.: Killian 6784 (W--1465230 )s Ruth 109 (W—501769); Whitehouse 16263 (wi). Taylor Co.: S. Me Tracy 8001 (W253). Val Verde Co.: Parry, Bigelow, Wright, = Schott 829 (W—56193). Wheeler Co.: A. Re Moldenke 117 (Fg). coun- ty undetermined: Havard s.n. [Texas, Spt. 1861] (W-—221199)- NEW MEXICO: Bernalillo Co.: C. C. Ellis 221 (W-662639). Catron 00+? ete A. R. Moldenke 157 (Fg, Z). Chaves Co.: Earle & Barle 351 (W— 1962 Moldenke, Monograph of Verbena 411 382500). Colfax County: Cockerell s.n. [Raton, Aug. 29, 1900] (W- 56227); Griffiths 5560 (W—l96719); P. C. Standley 6268 (W— 685299), 1h2h9 (W--897032). Dona Ana Co.: P. C. Standley 47h (W— 564402), sen. Sen. (Mesilla Valley, June 15, 1906] (W—560977); W Wooton Sen. (Mesilla Valley, May 17, 1893] (W—2h1169), sen. [San Augus- tine Ranch, Aug. 28, 1897] (W-—-736942); Wooton & Standley s.n. (Mesilla Valley, July 28, 1907] (W--56129). Eddy Co.: Wooton s. n. [vicinity of Queen, Aug. 2, 1909] (W--56655, W—736936). Grant Co.: 0. B. Metcalfe 137 (W--L9525h). Guadalupe Co.: A. Re Moldenke 125° (Fg). Lincoln Co.: A. R. Moldenke 131 (Fg). Quay Co.: G. L. Fisher 152 (W—660110); A. Rs ~ Moldenke idenke 122 2 (Fg). Rio Arriba Co.: Cockerell s sn. [No Agua, Aug. 1912] (W--691096) ; P. C. Stand- ley 6602 (W—68562)) ; Standley & Bollman 10852 (W—6895 38) 5 Woot on s.n. wn. fAug. 16, 190) J} (W--736937) « San Juan Co.: P. C. Standley 6927 (W#—685936), 7252 (W--686223), 7896 (W—68683); Wooton 2831 (W—736935), 283 (W—=736910) , Sen. TFarmington, Aug. 8, 190h] (W--56225). San Miguel Co.: P. C. ~C. Standley SUS (W—-1.98776), 3136 (W--99113); Studhalter 3.3028 (W--1735151). Santa Fe Co.: P. ec, Standley 458 (W--98751); Wooton s.n. (near Tesuque, Aug. = ~190h] (W—736941). Sierra Co.: > 0. B B. “Hetcalfe 1008 (wW— 497890) . Socorro Co.: H. H. Rusby 6512a (W—1h7507). Torrance Co.: A. R. Moldenke 129 (Fg). Union Co.: A. He Howell 159.2 (W—L950l47). County undetermined: Fendler 587 (Ww); Wooton SNe ~ [Inscription Rock, Aug. 1, 1892] (W—730939), Sen. [Gallo Spring, July 13, 1906] (W—736938) . "ARIZONA: Coconino Cos: Clute 93 (W--1070295); Collom 723 (W-~17294,8); Fernow s.n. [Little Colorado River, August 196] (W=-1,79895) ; MacDougal 286 (W-—334382); A. R. Moldenke 149 (Fg); G. A. Pearson ), (W—-56305h), 355 (W—5632h6). Navajo Co.: Mrs. Hough Sen. [Holbrook, June 1905] (W~890166); Peebles 9636 (Ww 162366); L . F. Ward s.n. (Holbrook, June 15, 1901 1901] (W--10318) ; Wooton s.n. . [7/19/1892] (W--2)1170) . vevanes. Co.: Peebles, Harri- son, & & Kearney 23 (W--1367578). WASHINGTON: Douglas Co.: Sand- berg & Leiberg 3h0 (W-—288317). Franklin Co.: Leekenby s-n. . [Pas- co, Aug. 8, 1898] (W—6221)8). Kittitas Co.: Cotton 1363 (W— 525159); J. W. Thompson 9067 (W—-1h1979). Klickitat Co.: Suks- sore s.n. [July 22, 1881] (W--147585). Stevens eh Kreager 7h 290). Walla Walla Co.: A: ie Me Horner 18 ( —28149;); Wilkes S.n. [Walla Walla, Oregon] (W ie man CO.? po 32h (W-- 277025). Yakima Co.: Cotton 621 TH 526183). Gounty undetermined: E. P. Sheldon 3202 (W—329122); G. R. Vasey 468 (W——-29674,7) . ORE- “mgr rrow Co.: J J. W. Thompson 4,880 1,880 (W—1)36931). Sherman Co: W. E. Lawrence 413 3 (W—98372h) « Umatilla Co.: H. BE. Brown 58 (W— aes Griffiths & Hunter 18 (W--527103); Leiberg 902 (w— 288318). Union Co.: ae By P. ~ Sheldon 8856 (W—528)97) . Wasco Coe: 12 PHYTOLOGIA Vol. 8, no. 8 Isat s.n. [Williamson's Exped.] (W). County undetermined: aes 1967 [eastern Oregon] (W—35),693). mers era Inyo Co.: & Funston 953 (W--56220). Kings Co.: Kearney Snes “Ton ‘Angeles Co.: .: Braunton 379 (W--1,65367). Monte ane Co.: G. R. Vas 612 (W—56219). San B Bernardino Co.: Parish & Parish 2171 ( 56201, W—-56222), Tulare Co.: Edw, Palmer 2699 (W—277861). Coun- ty undetermined: Griffiths & Morris 19 [Humboldt River bottoms near Minnemucca] (W—l025L6); L. . Schoenfeldt 2925 (W--2)0859) MEXICO: Baja California: C. R. Orcutt 223 (W—1323062). Chihuahua: E. Stearns 7 (W—502757). ~ Coahuila: ila: Edw. w. Palmer 1048 (W--5617h) . Sonora: P. B. B. Kennedy 7032 (W—-1287203) « CULTIVATED: Switzerland: Herb. Hort. Basil. s.n. [Aug. 1839] (W—-264622). LOCALITY OF COL- LECTION UNDETERMINED: Havard s.n. [Penitence] (W--218868); P. C. Standley s.n. [along the Rio Grande river, May h, 1906] (w= 564,309). MOUNTED CLIPPINGS: Lag. & Rodr., Anal. Cienc. Nat. hs 260. 1301. (W); Woot. & Standl., Contrib. "U.S. Nat. Herb. 16: 166. 1913 (W). A BRASILIENSIS Vell. Additional and emen hisses bibliography: Fedde in Just, Bot. Jahr- esber. 33 (1): 632. 1906; Moldenke in Chittenden, Roy » Hort. Soc. Dict. Gard. h: 2209 & 2210. 1951; Moldenke, Résumé Suppl. h: 1—3 (1962) and 5: 1--5. 1962. The Hammarlund 631 in the Stockholm een cited in Phyto- logia 8: 321 (1962) as this species, is actually V. arvula var. the Montevideo herbarium, cited on page 322 of the same work, seems, on re-examination, actually to represent V. litoralis "HB. = The Ae S. Kaltenborn 160 [Kaltenborn & Kaltenborn 160] ci y me in pa as List Cit. 2: 602 (1948) and in Phytologia 8: 3a (96) 3 is V. parvula H ae) He ayek. laterial of V. brasiliensis has been misidentified and distributed as V. bonariensis& brevibracteata Kuntze, "Ver benia litoralis tag and *Stachytarpheta jamaicense (L. Vahl". A common e for the plant in Peru is said to be Mpepermo" For students aitabeb te in the speed of spread of this specy ies in the southern United States, the following dates are Piven s Geo ’ Glynn County - 1923, Chatham County - 1927, Bacon, Bibb, Bleckley, Clinch, Dodge, Jeff Davis, Jefferson, Lowndes, Putnam, Screven, Telfair, Thomas, Twiggs, and War @ Comties - 1962; Florida: Escam- bia County - 1899, Washington noite 1930, Leon County 19555 Taylor County - 1957; Alabama: Mobile County - 1893, Sumter Co” nty - 1960, Baldwin County - 1962; Mississippi: Forrest County ~ 1932, 1962 Moldenke, Monograph of Verbena 413 Jones County ~ 190, rors aga - 1946, Harrison County - 199, Hancock County - 1950, Lee and Pike Counties - 1955, Jack- son and Pearl River Counties - el, Louisiana: Plaquemines Par- ish - 1900, Orleans Parish - 1901, Ter rebonne Parish - 191), East Baton Rouge Parish - 1927, Jefferson Parish - 5g Saint Tammany Parish - 1939, Tensas Pari sh - 1940, Calcasieu, Cameron, Iberia, Jefferson Davis, Saint Mary, and Washington Parishes - 1962; Tex- as: Hardin, Jef ferso on, and Orange Counties - 19)5, on County - 1916, Victoria ral - 1962; California: San Joaquin County - 1903, Hovada County - 191, Stanislaus County - 1958, Butte Coun- - 19 be ddit ie 1 citations: VIRGINIA: Warwick Co.: Fernald & Long 7585 (W—-1699825). NORTH CAROLINA: Bladen Co.: A. R. 1. R. Moldenke mke 1461 (Fg). New Hanover Co.: Biltmore Herb. 5707 (W--33210h); Godfr acess & Shunk 072 (W—1767208); McCart: McCarthy 8 (W). SOUTH CAROLINA: Ban Coe: AeR. R. Moldenke 27 (Fg). Barnwell Co.: A. Re. Moldenke 22 (Fg). Columbus Cow: FE R. Moldenke 1,57 (Fg). Dillon Co.: A A. Re Re Moldenke 450 (Fg). Horry Co.: A. R. Moldenke 55 (Fg). Kershaw Cost A. | R. Moldenke 42 (Fg), lS (Fg). Lee Cos: A. R. Moldenke 436 (Fg). Lexington Co.: A. Re Moldenke 133 (Fg). Marion Co.: A. R. | Moldenke 9 (Fg). Orangeburg Co.: A. Re R. Moldenke 435 (Fg). Richland Co.: A A. R. Moldenke 439 (Fg), bil (Fg). GEORGIA: Bacon Coo: A. Re Moldenke 346 (Fg). “Bibb Co.: eS R.- Moldenke 369 (Fg). Bleckley Co.: A. R. Moldenke 376 (Fg). Chatham Co.: Kolthoff s. n. (Savanna, 27 Maj 1927] (W—1367190). Clinch Co.: A. R. Mol= denke 329 (Fg). Dodge Co.: A. R. Moldenke 363 (Fg). Glynn Co. UcAtee 3295 (W—111)063); H. N. Moldenke 116) (W—1581762). aie Davis Co.: | A. R. Moldenke 3 354 (Fg). Jeff Jefferson Co.: A. R. Molden- ke 40h (Fg). Lowndes Co.: A. R. Moldenke 320 (Fg, Z). Putnam Co.: A. me Moldenke 389 (Fg). Screven C0.: Cont Ae Re R. Moldenke 413: (Fg). retoet ir Co.: AoRs R. Moldenke 362 (Fg). Thomas Co.: ; Cost Ae R. R. Moldenke 06 (Fg). Twiggs Co.: A. R. Moldenke 390 (Fg). Ware Co.: Stic, 336 (Fg). FLORIDA: Escambia Co.: Curtiss 6490 (W—357553) ES em Tracy 8706 (W--510598). Washington Co.: Oe? F | Ss. » Blanton anton 6598 shiaes3s); E. S. Ford 3785 (W--2230914). — hao Cost ir R. Moldenke 262 (Fg, Z)~» Mobile Co.: C. T. Mohr s.n. [July 1893] (W—771881) ; A. R. Moldenke 261 (Fg). esisstirt: Harrison Co.: A. R. Moldenke 253 (Fg). Jackson Co.: A. R. Moldenke 258 (Fg). Jefferson Coe: McDougall 1036 (W--1925222). Pearl River Co.: A. R. Moldenke 249 (Fg). LOUISIANA: Assumption Par.: A. R. Mol- denke 236 (Fg). Calcasieu Par.: A. Re Moldenke 226 (Fg). Cameron Pars: A. | Re Moldenke 233 (Fg). Iberia Par.: A. R. R. Moldenke 229 (Fg). Jefferson Davis Par.: A. R. Moldenke 235 (Fg). Lafourche +? A. R. Moldenke 20 (Fg) Plaquemines Par.: Tracy & Lloyd 20 (W—383532). Saint Charles Par.: A. R. Moldenke 2h1 (Fg). t Mary Par.: A. R. Moldenke 231 (Fg). Saint Tammany Par.: A. 41) Fret OL Costs Vol. 8, no. 8 = Moldenke 243 (Fg). Terrebonne Pare: A+ R. Moldenke 238 (Fg). ermilion Par. A. R. Moldenke 234 (Fg). , Washington Pare: ALR. Mr 2h7 (Fg)» 28 (Fg, 2). TEXAS: Hardin Co.: C. L. Lundell 7033 (W--223597h). Harris Co.: A. R. Moldenke 213 (Fe, Z, Z)> Jefferson Co.: J. F. Brenckle 48073 . (W——2003826)5 Co C. L. Lundell 14136 (W--1927003); A. R. Moldenke lenke 221 (Fg), 22h (Fe, “Z). Liberty Co.: A. R. Moldenke 217 (Fg), 218 Z), 218 (Fg). Orange e Co.: Cory 5080 (W--1886061). Victoria Co.: A. R. Moldenke 202 (Fe, 2)- CALIFORN- IA: Fresno Co.: E. Carter 91 Kayenta Yuba Co.: L. S. Rose 39200 (W). ECUADOR: Azuay: Rose, Pachan » & Rose 228L5 (W— (W--170501h). Paran&: Dusén 14320 (W--181776). Rio de Janeiro: Dusén 8 (W--1159395); Holway & & Holway 1272 (W--1188243). S&o Paulo: Campos Novaes 916 (W--389887) 5 A. Carvalho s.n. [Herb Inst. Bot. S- Paulo 190] (W—1592285). BOLIVIA: Cochabamba: car denas 3019 (W—159)061). La Paz: Julio 49 (W--1192637). PARAGUAY: Morong 126 (W—-1)17048). URUGUAY: Kuntze s.n. (Montevideo, 7 7/12/ 1891] (W—700691). CHILE: Valdivia: Claude-Joseph 261 16 (Ww 1198543). Province undetermined: Claude-—Joseph 1286 [Ramon] (W-- 1059300). ARGENTINA: Buenos Aires: Pedersen 349 (W--2122390) - Corrientes: Pedersen 1911 (W—2283117). La Rioja: T. Meyer hak (W~-1909186) . Mendoza: Miers s.n. [Mendoza] (W—-121161) 5 ; oa 293 (W—-20)9690). Saritiago del Estero: Luna Risso (Gg--352007, ¥, N), ko (N); Pierotti s.n, (Quebrada Tazana, a, April h, sth, Herb. Inst. Miguel Lillo 100888] (S). Tucum4n: Bailetti 233 [Herb. Inst. Miguel Lillo 31392] (N); Gillegan & Ros SNe re [9 octe ubre 1950] (N); Venturi 50 (Herb. Inst. Miguel [illo 31391; Herb. Osten 17263] (N, Ug). REPUBLIC OF SOUTH AFRICA: Cape of Good Hope: a3 SS 1098 (Ka~-92330); C. N, Lake san. [3-h-1922] (S)- Natal: J. M. Wood 12122 (Vt). Transvaal: Repton 1300 (Z); Rodin 3917 (Ca--802719); Schl Schlieben 7691 (N, W-2272h7 8). MADAGASCAR: Cours 863 (P); Decary 17970 (P, S), 18178 (P)- POLYNESIA: Hawai- dan Islands: Oahu: A. A. Heller 20b6, in : in part [Oahu] (Ca—~101819) 5 M. A. King s.n. (Oahu, ~1902] 02] (Ca--73591) « CULTIVATED: Brazil: Du- 3én 8 (S). Mary Maryland: McCann s.n. [9-15-36; seed from Brussels] (ud), sn. (9-15-36; Seed from Montevideo] (Md). ete 950. var. SUBGLABRATA Moldenke, Phytologia 3: 0, raphy: elaeoks Phytologia 3: 278 & 290. 19503 Molden ke, “about 121 & 71. 2 959. ‘This v ety differs from the typical form of the species in having i iieeaae and stems oo or subglabrate and its epeticien uniformly twice furcate, the primary peduncles 1962 Moldenke, Monograph of Verbena 41s glabrous or subglabrate, the secondary peduncles and rachis puber- ulent, the bractlets non-ciliate, very minutely and obscurely puberulous or ii the calyx very minutely puberulent and ake “i all strigos type of the mney: was collected by Ernesto Barros Valen- ee (no. 8050) at San Fernando, Colchagua, Chile, on February V5 1930, a and is deposited in the Britton ———s at the New York Botanic al tcc den, The variety is known thus far only from the type spec Citations: ee Colchagua: Barros Valenzuela 8050 (N—type). VERBENA CABRERAE on ee Phytologia 1: 478—l79. Bibliography: Moldenke, Phytologia 1: Ty Bi9 a le) and 1: 511. 191; Moldenke, Lilloa 6: 325. 191; Mold a oer. Distrib. Verbenac., [ed. 1], hh & 101. 1925 woudetie r 10: 346 & 382. 19lih; Moldenke, Alph. List Cit. 1: 8h. nee ret] Salisb., Ind. Kew. apes 10: sae rs f He ae & me a Leni Py. Life 2: 52. 198; pk fy ; Moldenke Alph, List Cit. 2: 37 on OE z "599° (98), 43 808 & oa (1949), and : 1077 & 1090. 199; Moldenke, Known Geogr. Distrib. Verben- Spec. 3. 1954; Moldenke, Résumé 115, 126, & 71. 1959; Moldenke, Résumé Suppl. 2: 5. 0. Perennial, prostrate, creeping or semi~creeping herd; stems and branches slender, obtusely tetragonal, rather sparsely appressed-pilose with whitish hairs, more densely so toward the apex on younger parts; nodes annulate; principal internodes 2--l, cm. long; leaves decussate-opposite, usually with several smaller ones on very much abbreviated twigs in their md. ieee ioles slender, 5--10 mm. long, winged, why vemmlrcaeee Som blades very thin-chartaceous, uniformly green on both ace, aa 6 cm. long and wide, Seenty 3-parted to the base, the divisions laciniate or pinnatifid, the lobes oblong and hairs on both pele na prefs and secondaries plane or subim- pressed esos: prominulous beneath; veinlet reticulation arr cernible on both surfaces; ; inflorescence terminal; pedunc Slender, appressed-strigose, 2.5—l. 5 em. long; spikes imesiae. ed, congested, many-flowered; flowers perfumed o ect calyx narrow=tu bular, about 9 mm. long and 1 m, wide, white- strigose; corolla varying from violet, heer a eeigiebaliey ° blu lilac- elilac white, purple, or even vivid rose-red, its tube about 12 m. long, the Limb mostly blue, 7-9 mm. wide. The type of this species was collected we — Iulio Cabrera (oe 4199) -- in whose honor it is naned — at Quebrada del Rfo 416 PHYTOLOGIA Vol. 8, no. 8 and November, at altitudes of 10 to 1200 meters. It has been found in quebradas, in fertile soil at the edge of cultivated ground, in savanna fomations, in sandy loam od See and "very rare, perfumed". Both collections were identified as "Verbena tenuisecta Briq. forma" by Osten. The Venturi 4607, cited in Lil- ik 6: 325 (191) as a satrerse is actually the type collection of lilloana Moldenke ; was originally identified as "Verbena er- Siciiee Tan Lam, aff." Ute ia the basis for the Tucum4n records rds Listed by me for V. cabrerae in previous publication Herbarium material of V. cabrerae has been "misidentified and distributed in herbaria as 's V. calliantha Briq. In all, 2) herbarium specimens, including the type, have been examined by me tations: BOLIVIA: Santa Cruz: CArdenas 4617 (W-2027163) » 5235 (W—2250761); Peredo s.n- [Capital, Gtal, 27111-1946) (S, S)s Se n, (Las Luntas, 20.1 “I9h7] (Gg--353270, N); Steinbach 2533 (Herb. Osten 14799] (Ug), 2673 [Herb. Osten 1801] (ug), Subk [Herb. Os- ten 16227] (Ug). ARGENTINA: Buenos Aires: A. T. Hunziker 3861 (S); Pastore 1163 (Go). Catamarca: Pierotti : son, [La Merced, ed, 19- V-I9lis] (N)s Chaco: T. Meyer 678 [Herb. Osten 22702] (Ue), S788 (Herb. Osten 22701] (Ug). Formosa: J&rgensen 21,70 [Herb. Inst. Miguel Lillo 31725; Herb. Osten 14554] (N, Ug)- ). Salta: Cabrera 4199 (N--type); A. T. Hunziker 1212 (W--180)100); Pierotti oo (N); Schreiter Ba (Herb. Inst. -. Miguel Lillo 31492] (N)5 As Ge Schulz 5299 (cb), 5337 (Z). Santiago del Estero: Ruiz Huidobro 3079 (Gg--353272, N). VERBENA CALIFORNICA Moldenke, Known Geogr. Distrib. Verbenac., [ed. zi; 719.6 ve Bibliography: Moldenke, Known Geogr. Distrib. Verbenac., [ed- 1], 79, 80, & 101, 19425 Moldenke, Know Geogr « Distrib. Verbenac. Suppl.'1: 3. 19h3; Moldenke, Alph. List Cit. 2: 559. 1985 Mol- denke, Known Geogr. Distrib. Verbenac., [ed. 2], 27 & 197; 1995 E. J.’Salisb., Ind. Kew, Suppl. 11: 262. 19535 wordenke, résumé 33 & L71. 1959; saat ne rental = - i en in af-blades firmly chartaceous, rather —_ oblancee- mpi darine Srerye.) 1 oe} cm. long, 5--10 mm. wide, te r subacute 1962 Moldenke, Monograph of Verbena 17 at the apex, irregularly serrate with obtuse or subacute, more or less appressed, antrorse teeth except toward the base, attenua with rather straight and subparallel margins to the rather broad clasping base, decidedly revolute along the margins from base to apex, scabrous and rather densely hispid with long white bulbous-based hairs on both surfaces, especially above, the hairs 4 ten slightly promimlous beneath; inflorescence terminal, spicate; spikes usually in 3's, the outer ones often again singly branched, similar to the stems in all respects, but usually muc shortly “pengere: bendy pe at the base; bracts aby foliaceous, 1b long o mm, wide, firm s and - long, t eursuie “densely beset with appressed white strigullae exce ee oy ok oe thie species was collected by Robert Francis Hoov- = ton! 3870) three miles north of Keystone, Tuolumne County, California, on September 30, 1938, and is deposited in the herb- arium of the University of California at Berkeley. The species has aes collected at an altitude of 1300 feet, flowering and in June and July, but I have not as yet had time to aeenaat the flowers. Herbarium material has been misidentified and distributed as V. carolina L. and V. polystachya H.B.K. In all, 5 herbarium specimens and 2 mounted photographs, including the type, have been examined by me Citations: CALIFORNIA: Tuolumne “Cos: Belshaw 2357 [Herb. U. S. Dept. Agr. Forest Serv. 16881] (Ca--12h399); Hoover 2382 (Ca— 76699), 3870 (Ca--type, N—isotype, N—photo o of type, Z--photo of bth CULTIVATED: California: Belshaw s.n. [May 25, 1937] 360). (Ca—56 Briq. in Chod. & Hassler, Bull. Herb. Boiss., é k: 108 1060, 190 ser, a5 1059——1 Syno onymy : Verbena erinoides Chod. (in part) ex ota in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, h: 1059, in syn. 190} [not v. ra Auth., 1959, nor "Hook., “4959, nor Sook k & “arn., 1959, nor L., 1959, nor vets 1791, nor Poepp., 1847, nor Spreng., 1830, 418 PHYTOLOGIA Vol. 8, no. 8 nor Willd., 1947]. Verbena chamaedryfolia x tenuisecta Briq. in Chod. & Hassler, Bull. Herb. Boiss., sér. 2, 4: 1059, in syn. 190). Verbena calliantha var. genuina Bri Bull. Herb. Boiss., sér. 2, : 1060. 1904. Verbena melindroides x temuisecta Osten ex Moldenke, Résumé 369, in syn. 1959. Verbena chamaedryfolia var. bipinnatisecta Kuntze ex Moldenke, Résum6 362, Bibliography: Bri hod. & Hassler, Bull. Herb. Boiss., sér, 2, : 1059--1060. 190); Briq. in Chod. & Hassler, Pl. Hassler. 10: 481--I82. 1904; Prain, Ind. Kew. Suppl. 3: 187. 1908; Moldenke, Pre Alph. List Invalid Names 5. 190; Moldenke, Su pl. List Bt Moldenke, Lilloa 10: 346 & 382. 19h; Moldenke, Alph. List Cit. 1: 16 & 263 (1946) and 2: 374, 375, 378, & 384. 1948; Moldenke, Cas~ tanea 13: 118. 198; Moldenke, Known Geogr. Distrib. Verbenace, 197. 1949; Moldenke, Alph. List Cite 3: : 2 Set 9 Spec. 3 (195i) and 51 Spec. . 1956; Moldenke, Résumé 109, 115, 118, 126, 350, 362, 363, 369, & 71. 1959; Moldenke, Résumé Suppl. 1: 7. 1959; J. A. Clark, Card Ind. issue 26. n.d. Suffrute olm ascen -stout, pubescent with antrorse hairs; branches ascending; middle inter- nodes cm. long; leaves decussate-opposite; petioles to 2 cm. long, shorter than or subequaling the lamina, antrorsely pilose- pronim ; florescence terminal, spicate; peduncles elongate, 3--15 cm. long} spikes congested, eventually to 5 cm. long; bractlets lanceolate, h—-S mm. long, half as long as or subequaling the calyx, densel arely mixe subsessile black glands, at maturity enlarged-urceolate, the mouth oblique, the teeth lanceolate-subulate, the anterior ones longer and to 2 mm, long, the posterior ones shorter and 1--1.5 F 3 é F E tq : : e a a P | : a z A limb , g mn, wide, spreading, the lobes broad, obcordate, the larger ones 3——l, mm. long and mm, wide, the terminal emargination to 1 m. 1962 Moldenke, Monograph of Verbena h19 deep; style exserted a mm. beyond the calyx-teeth after mn sis, incrassate toward t Pay x and there subbilobed; seeds su linear-oblong, yellowish, 3--3.5 mm. long, 0.8 mn, wide, oe fully elevated-reticulate a the top. The species is based on several collection made by Emil Fane er in Paraguay: no. 1782, collected on the campo at Sapucay in Jam ary, no. 3196, ¢ collected on the shores of Rfo Limpio in August, and no. 0. 7615, collected in thickets near Concepcion in October, all ‘deposited pee — herbarium of the Conservatoire et Jardin sek wg Sam at G species had en found a collectors in fields and campos, rac ream riverbanks, and growing as a ses in cultivated fields. Pedersen t ow pasture on sis from August to March, altitudes of fram 700 to 2700 meters. A vernacular name ponders is "margarita". Meyer states that in eee it is frequent on railroad embankments, in abandoned soil, dite oe notes that "Le v. calliantha est voisin de V. tenuisec- ta, dont il se sépare trés facilement par l'indument, le mode de division des feuilles et l'ampleur relative de leurs ne CSe » la forme des bractées et des dents calicinales etc." What appears to be a natural hybrid between this species and V. cabrerae is dis- cussed herein under xV, argentina Moldenke. Herbarium material of V. calliantha has been misidentified and distributed as Vv. chamaedryfolia Juss., V. cheitmaniana Moldenke, "V. erinoides Lam. aff." , and V, santiaguensis (Covas & Schnack) Moldenke. The Fiebrig & Osten s.n. (Herb. Osten 8063] was claim- growing, he says, with no. 9013 ee con end —, was identi- fied by him first as V. or secta Briq. and then as V. chamaedryfolia va T einteak ease as (probab- x an error for "subpinnatisecta Ra His no, 9015, type of V. melindroides x tenuisecta, was growing among 4a peruviana and :. temisecta. The Santa Cruz, Bolivia, record for V. calliantha given by me in my Résumé and elsewhere is based on Peredo sen. [Las Juntas, 20-I-191,7] which now proves to be V. cabrerae instead. The Monet- ta S.n. (Herb. Inst. Miguel Lillo 31696] and Lillo 3989 [Herb. Inst. Miguel Lillo 31691] cited in Lilloa 10: 346 (19k) are actu- ally V. lilloana Moldenke, and the Bailetti 231 [Herb. Inst. Migu- el Lille 31690) and R. Seeeeiter 2358 (Herb. Inst. Miguel Lillo 32760] are V. dissecta Willd., as are also the Zabala 116 [Herb. Inst. Miguel Lillo 30045) eae rs Lilloa 10: 362 (194) and the 4,20 PHYTOLOGIA Vol. 8, no. 8 T. Meyer 309) cited in Lilloa 6: 326 (191). On the other hand, Cabrera 3169, cited by me in — Alph, List Cit. 1: 8) (196) as v. dissecta, is diene’ vec antha, while Venturi 526, cited as V. calliantha in Alph. List == Lh: 1090 (1 99), is V, diss- ecta. ~~ Tt is perhaps worth noting here that the other part of V Vv. erin- oides Chod. is V. storeoclada Briq., while V. erinoides Auth. is V. dissecta Willd., V. erinoides Hook., Hook. & Arn., L., Poeppe, and Spreng. are all Ys berterii (Meisn.) Schau., V. erinoides Lam. is V. laciniata (L.) Brig., and V. erinoides Willd. is V. ciliata enth. In all, 32 herbarium specimens, including the type collections of all ito a0 names involved, and 2 mounted photographs have been 8 Bikextons: BRAZIL: Rio Grande do Sul: Schreiner 15 gr Schwacke sen. (Herb, Saldanha 6326] (Ja—-)6578). BOLIVIA: Tarija Pflanz 4056 (B, W—13)1238). PARAGUAY: Fiebrig & Osten s.n. [Herb. Osten 8063] (N, Ug); Hassler 3196 [Macbride slaw es ol: a photo of cotype, N—-photo of cotype), 7615 (N—cotype); T. Meyer 16035 (N, W—-20)9913); Osten 9014 (Ug), "9015 (Ug); Pedersen 3210 (W—2283617) ; T. Rojas 2539 (Herb. Osten 18187] (N, Ug); A+ i Schulz 7789 (Cb). ARGENTINA: Catamarca: Brticher & Brttcher 8 aon. [30/1749] (Ss). chaco: Cabrera 3169 (Ug); T. Meyer Ul [ke (Herd. Os- = 22664] (Ug); A. G. Schulz 1477 (N), TUS (235 Corrientes: SS 1844 (Hi Hi—193251, N <8 $, W—-2283108); Ybarrola 542 (W— 29). Formosa: Morel 1003 (N), 121) (N). Misiones: Yontes oe (W—1988288); D. Rodriguez 5u8" [He: {Herbd. Inst. Miguel “Tillo 31367) (N). Salta: T. Meyer 1,608 . (Ca-=906601). Tucumn: Olea 66 (Ca); Ruiz Leal 12618 (Ss); Venturi 3568 (Ss). VERBENA CALLIANTHA var. MICROSOMA po gg in Chod. & Hassler, Bull. erb. Boiss., sér. 2, h: 1060. 1 Bibliography: Briq. in Chod. & Hassler, Bull. Herb, BoisSe, sér. 2, 4: 1060. 190; Brig. in Chod. & Hassler, Pl. Hassler. "10: 482 * Bo; Moldenke, Known Geogr. Distrib. Verbenac., [ed. a la ton (a9 (a9l2) and [ed. 2], 99 & 197. 19195 Moldenke, Résumé This ae “differs from the typical form of the species in its smaller stature, being only 8--20 cm. tall, its smaller leaves, the blades of which are only 2 cm. long and 1 cm. wide, and their narrower segments, the shorter ototas, which are on~- pd 5 Ba long, the more slender peduncles, and the more reduced spikes. the type of this variety was collected by Emil Hassler (no. 1272) in thickets near Cerro Pyta, Paraguay, in October, and d is deposited in the ene of the Conservatoire et Jardin Botani~ ques at Geneva, I have as yet seen no material of this plant and 1962 Moldenke, Monograph of Verbena yal suspect that it may be merely a depauperate ecotype of the spe~- CAMERO Is L. I. merne Nature Leafl. ones Rio Grande Valley Nature Club 1, pr. 1, [2-3]. Augus ae ae Verbena lundelliorum Moldenke, Sach ne 2: “oh. A t 26, 19h1. Verl Verbena lundellorum Moldenke ex A. M. Davis, Davis, Nature Leafl. Lower Rio Grande Valley Nature Club 1, pr. 1, [2-3] pe and we 2, — a3 Fangs Moldenke, Phytologia 2: 2h. 19h1; W . Ind. 305. 19h1; Moldenke in Lund- ell, oy tag 2 (ie 18 & 3h. 3912; A. M. L. Davis » Study Bosca- je Palma Cameron Co. 31, 39, \i1--43, & 61, pl. 10, mss. 1942; Moldenke, Alph, List Invalid Names 18. 19 trib, Geogr. Distrib. Verbenac. Suppl. 1: 2. 1943; ea 10: 4O, 1945; Moldenke, 2 Ame a pod (195) 2: 160. 19h6; G. L. er, - 1946; Moldenke, = age List Cit. 1: 14) (igus) and 2: 518. 19h; “ichaenke, , Wrightle 229-—~—230. 1948; H. Ne & Ae L. Moldenke, Pl. Life 1918; Woldenke, Alph’ List Cit. 3: 678-681, 723, 76h, & 708 “ata and 4: au, ‘123, 1218, 125, & 1291. 2 19h9; Moldenke, Known Geogr. d. Py ae 32, & ior. 1949; Moldenke, Phy- tologia 3: “Be 1949; Ee Ai Salisb bes Ind. Kew, Suppl. 11: 262 & 263. 1953; Moldenke, Résumé 28 369, & h71. 1959; Lewis & Oliver, Am. Journ. Bot. 8: re ta 1961. Illustrations: A. M. L. i a Boscaje Palma Cameron are 10, Mss. 192; Lewis & Oliver, Am, Journ, Bot. 48: 60. Sprawling or prostrate perennial herb (when large), sometimes covering an area over 3 m. wide, or bushy upright and to 5 cm. ed, obtusely tetragonal, hirsute or ogg pe al ; branches numerous, very slender, erect or ascending, obtu vasely "tetragonal, more dense- ly hirsutulous; principal internodes )—10 cm. long; leaves de- cussate-opposite, numerous; petioles siender or very slender, 0.5—2.5 cm, long, hirsute or a hirsutulous like the . baer urants thin-chartaceous or mem 1i, r green beneath or else mitein, 2 above h blunt or subacute teeth, appressed-hirsute or sparsely scattered- Pilose on both surfaces with aye wea Lg hairs; ~ ores— ant: esis, cence spicate, abbreviated and 2 densely many-flowered, the flowers ciowely tieloate before and during anthesis, somewhat more separated in fruit, elongated to 22 PHYTOLOGIA Vol. 8, no. 8 15 em. in fruit, with few Spessinly yagi to 7) open flowers at any one time; peduncles al te ) and rachis very slender or filiform, ports acutely renga ge Bore or a- bundantly hirsutulous like the branches, often with shorter glandular hairs Eitaneresds “heactieth linear-lanceolate or linear-attemate, 6--8 mn. long, slightly shorter or longer than the calyx, setaceous at. the apex, hirsute or densely Ne aa long-ciliate along the margins with stiff white hairs; calyx t ular, swollen, mm. long, about 1.5 mm. wide, hirsute or wide’ at the base, the commissure] face asic ation reaching the apex, muricately scabrous; chromosome number The type of this interesting species was haath by L. Irby —. eas a palm grove on the a aqui: Plantation, Cameron Coun- ’ , in April, 191, and is deposited in the herbarium of hoe univeaty of Texas at Austin, The type of V. lundelliorum ected by Cyrus (os 3658) -- in whose honor “3 was named — in a clearing at the palm g grove, south of Brownsville, Cameron County, Texas {in other words, at the — tooetityl; a on May 4, 1940, and is deposited in the Britton Herbarium at the New York ce of Peo Garden. Lundell 10011 is said a have come from the type colony. It is rather singular that both binomials proposed for Pan nd = botanical libraries twelve days before mine was, so his name stand te Date ites Noe name for the plant in question. This my booklets "The Known Geographic ae Avicenniaceae", p. 8 (March 14, 1942), and in Phytologia 2: 127 (Jamary ], 1 Ths) and so (aay 8, 1946). Anna May Tarrance Davis, " her thesis entitled " study of Boscaje de la Palma in Came eron County, Texas, and of sabal texana", pp. 31, 39, l1l—li3, & 61 sooo 192), gives valuable additional information about this s and on pl. 10 a splendid illustration of it. Te species has been found by collectors on praseise, Oo e roadsides, in woodlands, fields, and an ground of tennis courts, at altitudes of 20 to 3100 feet, blo in . every month of the n Texas it is said that March and Apr are usually the months of best bloom, and good fruit has been col- lected in July. In Mexico it has been found in fruit in June, 1962 Moldenke, Monograph of Verbena 423 growing in sticky black non-wooded valley soil. Davis reports that the January and February spikes renee look quite unlike the later ones "for tee have as many as seven flowers open at one time, a circle around the dgihical mass of buds. Later e- longated spikes have only three or four flowers open at one time. It i t grows more like V. ciliata and the leaves are smalle 4 Po thick- er. In Cameron County, Texas one plant, growing in the open, covered an area ten feet across and sh howed extensive ce a at the nodes." King calls it an "abundant weed in open sun" and " bundant in partial shade" in San Luis Potosi. evens Y eauitie the plant as oe "along roadsides at palmgroves thrown in road making, in light brown silt in Sabal texana — Celtis mississippiensis complex", while Cory reports it "fre- quent o n grounds of tourist courts." King found it in fields with abundant weeds in open sun The Ecology Class Univ. Texas s.n. cited by me in Lundell, Fl. Texas 3 (1): 36 (192) as as Ve delticola Small, as well as the Liebmann 11313 & 11314, Nealley 117 & 118, and Palmer 90 cited by Perry (1933) under that species, are gE v. cameronensis " erbarium material of V. cameronensis has also been misidentified and distributed as V. aubletia L., V. canadensis (L. ) Britton, V. delticola Small, and " V. elegans H.B.K. Galeotti 717 is the type collection of V. teucriifolia Mart. & Gal., but the he sheet of it photographed by Macbride as his photograph no. 2h699 is V. camer- onensis; the real type of V. teucriifolia is the sheet with this @ number in the herbarium of the Jardin Botanique de 1'Etat at foskecla, seen and photographed by me. series of letters to me, Mr. Davis gives quite sam e in- contained several errors (one of them rather bad) seaeciene 50 or more......were mailed out to a first preferred mailing 1 which was.....made up mostly of friends and a few specialists. a piesa" printing was then made and it is being sent to all those aig and institutions interested in botany and of course to thos who irst . makes the entirely uncalled for caustic comment that "Because of certain Post Office laws our opinion of such media for the publi- 2h PHYTOLOGIA Vol. 8, no, 8 cation at sear species cannot be expressed here." Davis continues: "Verbena cameronensis is the first new spec- ies I ever ast and I have delayed describing it for years while I was studying its variations and distribution. I; is rather com mon in parts of Mexico....I have numerous photographs of it grow- ing here and in different places in Mexico (some of these pictures show V. cameronensis growing in mixed beds with other species which thus show the characters most decidedly by contrast with i known species). I have considerable data on gio and asonal variation, and also a number of notes on differences with Ve delticola to present later. I could not present this v oluminous material in the original publication for reason of cost; conse- quently the rather unique 7 ae of publication was used... The 'yine-like' verbenas that occur naturally in Cameron fore Texas may be separated by the “teenporn scheme: V. s the corolla limb not more than 1/2 as wide as the calyx = long; Vv. Sasnonahets has the corolla limb usually exactly equal to the length of the calyx but may in extreme cases be 3/4 as wide as the calyx is long; and the others (V. ciliata, V. delticola, wee have the corolla limb considerably Wider than the calyx is long He goes on to say that the pubescence of V. cameronensis and T. delticola "is almost exactly the same even m though the plants are so decidedly different in the field here that it seems impossible that the two could be confused. Except in very young plants the leaves of delticola tend to be ag more irregular in outline than those of cameronensis here Texas —- most plants have 4 number of leaves that are ecaaae, three~divided in delticola.«- This difference will sae nable you to separate the plants without even looking at the perciae I find that in certain Mex- ican states the leaves of delticola become more regular in out- line but there are ey er other characters to readily separate the species. In the field cameronensis and delticola are easily separated ni the pris: es in color or by the difference in appearance of the centers of the heads of the spikes when pose), I think the Southmost area much more interesting. pes tropical plants that do not grow in the Rabb area are found in the Southmost bend. There V. cameronensis is quite common. PS dam noticed that the description of your V. lundellorum sounded ex- actly like V. cameronensis except for - the bt eh were ab~ surdly small for cameronensis. After studying i efully wife and I discussed it and decided it could va consibly be the 1962 Moldenke, Monograph of Verbena 2s as the corolla limb was only 2 mm. wide in your species Last April [191] we had made a study of the variation of camer~ onensis while on a trip into central Mexico...,and found that plants growing very poorly on dry hillsides had corolla limbs of 6--7 mm.; so we felt that it Seuid be impossible to have a cor- olla less than 6 mm, wide. That will be unfortunate if they have such bag ee corollas. If they a e same it certai for all the taxonomists that I carefully sent specimens to always called it 'delticola®! As far as can Hind it is the only ver- bena in certain places in Mexico all, 58 herbarium Beta ne including the types of all the names involved, and 9 mounted photographs and descriptions have been examined by Mee Citations: TEXAS: Cameron Co.: Cory 51353 (Ca——75),8)2, Du-- 345338, N, my W—-1885955); L. I. Davis s.n. s.n, [Southmost, Spring 1) (Au—-1218 38--type, F--photo of Serres N--isotype, N--photo of isotype, Si--photo of isotype, otype, Z--photo of iso- type), son. [Southmost, Feb. l, Soho} mt sen. (March 7, 19h2] (N), s.n. [Southmost, March 22, 192] (ag—-295228, M1, N, W— 1828599), s.n. [Bayview, Yarch 3, 1946] (Au--171996); Ecology Class Univ, Texas s.n. [Palm Grove, 3.1.30] (Au--121839) 5 e Le Fisher 1031 (Au--121537, Ge, S, Liars C. Le Lundell 10771 (N, W--192693)); Lundell & Lundell 8698 (Ld, es Atl N, i- photo, Z-—-photo), 10011 “(id, Mi, Mi, N, W—198887h), 1001, (Ld , Mi, N, W--1888876); Nealley 117 [1368] tke We-£6207), 118 (Up—-17081); R. Runyon 193 (W--173)131); Traverse 1107 (Au-- 179110, W—2339918). MEXICO: Hidalgo: H. E. Moore 2917 (Ba); A. J. Sharp 441820 (0). Oaxaca: Liebmann 113 a asie8lo) . San luis Potos{: L. I. Davis 213 (N); Re M. King 3983 (N, 2), L2h9 (N); C. = Lundell 12256 (N, W--1927026); Re Le L. Oliver 125 (Z)5 J. N. Weaver 667 (W--2134162), 677 (N)- Tamaulipas: L. 1. I. Davis 2k (N); ee 2° in part % (Gg——31h66, N, W—-l63027). Vera N—photo); Liebmann 11313 (W—-1315091); Medellin 0 (Me). MOUNTED DESCRIPTIONS: Le 1. Davis, Nature Leafl. Lower Rio 0 Grande Valley Nature Club 1: [2—3]. 191 (Ur); Moldenke, Phytologia 2: 2h. 191 (W). VERBENA CAMPESTRIS Moldenke, Phytologia 3: 117--118. 1919. Synonymy: Timotocia c nena ides (Cham.) Mold. ex Yoldenke, Résumé Suppl. 1: 23, in syn. Bibliography: Moldenke, Paptahoete 3 oo a (1949) and 3 ont 19505 E. J. Salisb., Ind. K e ll: 262. 1953; mite, Sellowia 11: 57 & 13h. 1959; on call gen 110 & 471. 1959; 4,26 PHYTOLOGIA Vol. 8, no. 8 Moldenke, Résumé Suppl. 1: 23. 1959. Low her rb; stems creeping, very slender, shortly appressed- pubescent; branches procumbent or ascending, very peal Pe rather finely strigillose above and on the venation beneath; midrib, secondaries, and veinlets very slender, often subimpressed above prominulous beneath; inflorescence terminal, spicate, the florif— bout mm. long, S-angled, 5-costate, 5-apiculate, spreading ciliolate on the angles; corolla igpconeian’ oes blue, its tube about ) a Sates 7 strigillose above the calyx, its lobes about 2 prore long, emarginate at the apex, finely strigillose on the out- s The type of this very distinct species was collected by Raulino Reitz (no. 2392) on a campo, at an altitude of 2000 meters, at Campos dos Padres, Santa Catarina, Brazil, on December 16, 1918, and is deposited in the Britton Herbarium at the New York Botani- cal Garden. The species has been collected in anthesis also in January. Common names recorded for it are "camaradinha", formosa sem dote", and ajuepaba® -- ao whic owever, = ’ applied exanined b BRAZIL: Santa Catarina: Rambo 60168 (S); Reitz 2392 (ivig pels "2505 (N), 2536 (N). ome eo ENSIS (L.) Britton, Mem. Torrey Bot. Club 5: 276. Synonymy: Buchnera canadensis L., Mant. 88. 1767. Verbena aubletia Jacq., Hort. Vindob. 1: pl. 176, and 2: 82. 1772. Obletia Rozier, Introd, Obs. Phys. Hist. Nat. 1: 367, ple 2. 1773. Verbems oblaetia Retz., Svenska Vet. Akad. Stockh. Nya H Handl. 34: 143, ple 5. 1773. Verbena obletia Medic., Act. Acad. Theod.-Palat. 3 (Phys)! 19h, pl. 7. 1775. Anonym Anonymos caroliniensis Walt., Fl. Carol. 16h- 1788. Verbena longiflora Len., Tabl. Eneyel. Méth. Bot. (ILlustr.] 1: $7. 1791. Glandularia caroliniensis J. F. Gmel., Syst. Veg. 2 (2): 920. 1791. Billardiera explanata Moench, dani. 9. 179. Verbena lisb., Prodr. 71. 1796. Verbena setreniet spicis solitariis; c corollis fascioulaties Folits foliis cordatis, inciso~ serratis L. f. apud Poir, in Lam., Encycl. Méth. Bot. “i ~ 8: 5u7, in syn. 1808. Verbena (aubletia), tetrandra, spicis laxis, § solitariiss 1962 Moldenke, Monograph of Verbena 427 foliis trifidis, incisis Ait. apud Poir. in Lam., Encycl. Méth,. Bot. 8: 547, in syn. 1808. Verbena (aubletia), assurgens, foliis varié dissectis; spicis solitariis, peduncularis, imbricatis, grandifloris Mich. apud Poir. in Lam., Encycl. Méth. Bot. 8: 5h7, in syn. 1808. Verbena (longiflora), spicis solitariis, densiuscw ‘ lis, corollarum laciniis emarginat, s; foliis ovalibus inciso- serratis, petiolatis Lam. apud Poir, in Lam., Encycl. Méth. Bot. 8: 547, in syn. 1808. Buchnera (canadensis), foliis laciniatus, caule dichotano L. apud Poir. in Lam., Encycl. Méth. Bot. 6: 57, in syn. 1605. Verbena anonyma Bart. ex Raf., Suppl. Fl. Ludovic. 139, in syn, 1817. Verbena aubletia Wild ex Raf., Suppl. Fl. Ludovic. 139. 1817. Glandularia carolinensis J. F. Gmel. apud Raf., Suppl. Fl. Ludovic. 139, in syn. (1817); Steud., Nom. Bot., ed. 1, 873. 1821. Verbena lamberti Sims in Curtis, Bot. Mag. 8: pl. 2200. 1820 [not V. lamberti Ker, 1827, nor Spreng., 1959]. Glandularia obletia Rozier ex Steud., Nom. Bot., ed. 1, 873. 1821, verbena grandiflora Michx. ex Steud., Nom. Bot., ed. 1, 873, in syn. 1821 [not V. grandiflora Bodger, 1936, nor Ort., 1797, nor Sessé & Moc., 1889]. Verbena intermedia Penny ex G. Don in Loud., Hort. Brit. Suppl. 1: 602. 1832 [not V. intermedia Gill. & Hook., 1829]. Verbena lamberti Penny ex G. Don in Loud., Hort. Brit. Suppl. 1: 602. 1832. Verbena lamberti var. rosea Sweet, rit. Fl. Gard., ser. 2, 4: 363. 1836. Verbena aubletia var. drummondi ” in Edwards, Bot. Reg. 23: pl. 1925. 1037. Gland- ularia aubletia (Jacq.) Nutt., Trans. Am. Phil. Soc. 5: 18h. 1837. Verbena drummondi Hort. ex Lindl. in Edwards, Bot. Reg. 23: pl. 1925, in syn. 1837. Verbena aubletia drummondi Lindl. ex G. Don in Loud., Hort. Brit. Suppl. 2: 680. 1839. Verbena lamberti rosea D, Don in Loud., Hort. Brit. Suppl. 2: 680. 1839. Verbena Grummondii Hort. ex G. Don in Loud., Hort. Brit. Suppl. 2: 680, in syn, 1839 [not V. drummondii Sweet, 197]. Anonymos carolin- ensis Walt. apud Walp., Repert. k: 31, in syn. 1845. Verbena aubletia L. apud Schau, in A. DC., Prodr. 11: 554. 1847. Verbena obletia Retz. apud Schau. in A. We, Prodr. 11: 554, in syn. I8L7. Verbena longifolia Lam. ex Rtmpler in Viln., Ijlustr. Blumeng., ed. 1, 1259, in syn. 1873 [not V.longifolia Mart. & Gal., 18h]. Verbena montana Hort. ex Sieb. & Voss in Vilmorin, Blumeng,, ed. 3, 1: 625, in syn. 1895. Verbena grandiflora Steud. ex Hook. & Jacks., Ind. Kew. 2: 1179, in syn. 1895. Verbena aubletia grandi- flora Haage & Schmidt, Cat. Général 93. 1898. Verbena drummondii (Lindl.) Baxt. ex Small, Fl. Southeast. U. S., ed. 1, 1011. 1903. Verbena lambertii Sims ex Small, Fl. Southeast. U. S., ed. 1, 1011. 1903. Verbena canadensis Britton apud Durand & Jacks., Ind. Kew. Suppl. 1: 451. 1906. Verbena aubletia var. lamberti (Sims) M. E. Jones, Contr. West. Bot. 12: 75. 1908. Verbena canadensis var. 428 PHYTOLOGIA Vol. 8, no. 8 lambertii Thell., Fl. Advent. Montpel. 428. 1912. Verbena aubletia drummondii Lindl. ex Stapf, Ind. Lond. 6: 29. 1931. Verbena lambertii rosea D. Don ex Stapf, Ind. Lond. 6: 430. 1931. Verbena aubletia var. lamberti M. E. Jones apud Perry, Ann. Mo. Bot, Gard. 20: 343. 1933. Glandularia lambertii (Sims) Small, Man. Southeast. Fl. 1138 & 1508. 1933. Glandularia canadensis (L.) Small, Man. Southeast. Fl. 1139 & 1508. 1933. Glandularia drummondii (Lindl.) Small, Man. Southeast. Fl. 1139 & 1508. 1933. Verbena canadensis var, drummondii (Lindl.) Baxt. ex Junell, Symb. Bot. Upsal. y: ll. 193). Verbena canadensis var. atroviolacea Hort. ex Dermen, Cyto logia 7: 16l1—163, fig. 10, 16h, 165, fig. 26, 168, & 17h. 1936. Verbena canadensis var. compacta Hort. ex Dermen, Cytolog : 161, 164, & 168. 1936. Glandularia drummondii (Hort. ex Baxt .) small apud A. W. Hill, Ind. Kew. Suppl. 9: 12). 1938. Glandularia lamberti (Sims) Small apud A. W. Hill, Ind. Kew. Suppl. 9: 124. 1938. Verbena bracteosa Pursh ex Moldenke, Prelim. Alph. List In~ valid Names 45, in syn. 190 [not V. bracteosa Michx., 1803). Anonymos caroliniensis J. F. Gmel. ex Moldenke, Prelim. Alph. List Invalid Names 5, in syn. 190. Glandularia aubletia Nutt. ex Moldenke, Prelim. Alph. List Invalid Names 26, in syn Suppl. List Invalid Names 7, in syn. 19)1. Verbena aubletia vare drumnondii Lindl. ex Moldenke, Suppl. List Invalid Names 7, in syn. 1941. Verbena aubletia var. grandiflora Hort. ex Moldenke, Suppl. List Invalid Names 7, in syn. 191. Verbena drummondii Baxt. ex Moldenke, Suppl. List Invalid Names 5, in syn. 1941. Verbena drummondii (Lindl.) Bart. ex Moldenke, Suppl. List In- valid Names 8, in syn. 191. Verbena drummondii var. montana Hort. ex Moldenke, Suppl. List Invalid Names 8, in syn. 19). Verbena canadensis L. ex Moldenke, Alph. List Invalid Names 46, in syne 192. Verbena canadensis var. lamberti Thell. ex Moldenke, Alph. List Invalid Names 6, in syn. 19h2. Verbena aublettia Roig, Plant. Medic. Cuba 671 & 811, sphalm. 1945. Bttchnera canadensis L. ox Moldenke, Alph. List Invalid Names Suppl. 1: 3, in syn. 1947. Glandularia aubletia Gmelin ex Moldenke, Alph. List Invalid Names Suppl. 1: 9, in syn. 1947. Glandularia canadensis XYZ ex Moldenke, Alph. List Invalid Names Suppl. 1: 9, in syn. 197. Glandularia lamberti Small ex Moldenke, Alph. List Invalid Names Suppl. 1: 10, in syn. 19),;7. Glandularia obletia Rozier ex Moldenke, Alph. List Invalid Names Suppl. 1: 10, in syn. 197. Verbena aublattia Role ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syne 19hi7 « Verbena aubletia Ait. ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syn. 1947. Verbena aubletia (L.) Britton ex Moldenke, Alph, List Invalid Names Suppl. 1: 22, in syn. 1947. Verbens 1962 Moldenke, Monograph of Verbena 29 aubletia Michx., ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, in syn. 1947. Verbena aubletia atroviolacea Haage & Schmidt ex Moldenke, Alph. List pidge Names Suppl. 1: 22, in syn. 1947. Verbena aubletia grandiflora Huntington ex Moldenke, Alph. List Invalid Names Suppl. 1: 22, ie syn. 1947. Verbena carthaginensis Berg ex Moldenke, an List naghee A Names ; Suppl. 1: 23, in syn. 1947. Verbena compacta Schling ex Moldenke, Alph. List Invalid Names Suppl. 1: 23, in = syn. 1947. Verbena drummondi Baxt. ex Mol- denke, oe List Invalid Names Suppl. 7 23, in syn. 1947. Ver- bena aii Lindl. ex Moldenke, Alph. List Invalid Names Suppl. 1: a3 in syn. 197. Verbena lambertier Berkimer ex Mol- denke, Alph. List Invalid Names Suppl. 1: 2 25, in syn. 1947. Ver- bena Jongiflora L. ex Moldenke, Alph. List Raa Names Suppl. 1: 25, in syn. 197. Verbena cadihininls s var. grandiflora lMoldenke in Chittenden, Roy. Hort. Soc. Dict. Gard. h: 2210. es « Verbena aublettia Jacq. ex Roig, Dicc. Bot. 1: 909 & 2: lil, 1953. Verbena canadnesis L. ex Moldenke, Résumé 360, a a 1959. Verbena obliqua Prager ex Moldenke, Résumé 370, in syn. 1959. Verbena aubletia compacta Hort. ex Moldenke, Résumé Suppl. 2: 10, in syn. 1960. Verbena aublettia L. ex Moldenke, Résumé Suppl. 2: 10, in syn. 1960. Verbena bracteosa x urticifolia Hall ex Moldenke, Résumé Suppl. 2: pl. 2: 10, in syn. 1960 [not V. bracteosa x urticifolia Eggert, 1958, nor Rydb., 1932, nor r Ste- vens, 1958]. Verbena canandensis (L.) Britton ex Moldenke, Résu- mé Suppl. 23 10, in in syn. 1960. Verbena drummondi (Lindl.) Baxt ex Moldenke, résumé Suppl. 2: 11, in syn. n. 1960. Uwarowia anita Meisn. ex Moldenke, Résum$ Suppl. 3: 35, in syn. 1962. Glandulosa canadensis Penfound ex Moldenke, Résumé Suppl. 3: 32, in syn. 1962. Verbena aubletia Gray ex Moldenke, Résumé Suppl. 3: 3, in _ 1962. Verbena lambertiana Sims, in herb. Verbena canadensis L.) Small, in _in herb. Verbena aubletiana Willd., in in herb. Verbena aubletia Willd., in herb. Bibliography: L., Mant. 88. as Jacq., Hort. Vindob. 1: pl. 176 & 2: 82. 1772; Retz., Svens . Akad. Stockh. Nya Handl. : 143—14h, pl. 5. 1773; Aner eng beara Obs. Phys. Hist. Nat. 1: 367, pl. 2. 1773; Medic., Act. Acad. Theod.-Palat. 3 (Phys.): K. uoench, mak. PL ek 5, Usteri, Ann. Bot. 9: pl. 2. 1. 3. ua 1795; Re A. Salisb., Prodr. 71. 1806; Poir. in Lan., Encycl. Méth. Bot. 8: 547—5h8. 1808; J. Sm. in Rees, Cycl. 36: no. 27. 1817; Edwards, Bot. Reg. he pls 1,30 PHYTOLOGIA Vol. 8, no. 8 29. 1818; Sims in Curtis, Bot. Mag. 48: Bike 2200. 1820; : Stats Nom. Bot., ed. 1, 873. 1821; eres in L., Syst. Veg., ed. 16, 2: Tho & 750. 1825; *vaund, Bot. Gard. 3: pl. "Be. 1829-18303 Sweet, i Hort. Brit. Suppl. Am . aoe 180; Hook. & Arn., Bot. Beech. Voy. h8h. 181; Walp., Rep- tis Se 33- 185; Spach, aoe Nat. Vég. Phan. Atlas pl. 8h. 1Bhe5 Schau. in A» DCe, Prodr. 11: 554. 1847; Schnitzlein, Iconogr. 2: pl. 137. 1856; A. W. gag Fl. South. U. S., ed. oy yy 20t (1860) and pr. 2, 307. 1865; Viln., oti Terre, Cea . 2 ; at W. Chapm., Fl. South. U. S., ed. 1, pr. 3, 3 307. 1872; Decaisne, Naud., & Hemsl., Trees Shrubs & Herb. Pl. 357. 18733 poe r in Vilm., Illustr. Blumeng., ed. 1, 12 259--1260. 1873; C. H. Harris, Floral Kingd. Wile ak 1876; Meehan, Fls Se & Ferns U. S. 1220. 1879; We Mill, the te i Names Pl. 141 & 261. 188h; A. W. Chapm., Fl. South. U. S., ed. 2, pr. 1, 307 (188k), pr. 2, 307 (1887), and ee "33 307. 1889; baill., Hist. Pl. ll: 79. 1891; A. S. Hitchc., Trans. S ad Louis Acad. Sci. 5: 512. 18915 A. We Chapm., Fl. South. U. ed. 2, pr. h, 307. 1892; Coult., Contrib. U. S. Nat. Herb. 2: ite. 1892; Hook. & Jacks., Ind. Kew. 1 (2): 1032. 18933; Briq. in Engl. & Prantl, Nat. Pflanzenfam. (3a): 148. 189); _ Mem. Torrey Bot. Club 5: 276. 18945 Hook. & Jacks., Ind. Kew. 2: 1178 & 1179. 1895; L. H. Bailey in he oats ete aoe & Gard. Bot. 31. 18955 Sieb. & Voss in Sou th. ve Se, a. a 368. 1897; step, ae _ Gar d. be Greenh. 33 ig 20. 18975 Britton & Br., Illustr. Fl., ed. 1, 33 69, 71, ° 1: h . 3 lee 1 Bailey, ° er. Horte, ed. L, 6: rise. 1906; D. T. MacDougal, Bot. Gaz. 33 Ae 1907; Brie, . & . Bot. so 10: 103.6 7 J s Robinson & Fern. in A. Gray, New Man. Bot., ed. 7, 689 & 92h. 1908; P. Henderson, Handb. Pl., new ed., 75. 1910; wettstein, ulde er [New Nature Lib. 6:) 2. 1916; L. H. Bailey, Stand. crcl Hort. 6: 3hh7. 1917; Olmsted, Covilie, & Kelsey, Stand. Pl. Names, ed 1, 520 & 521. 192h; Hegi, Ill. Fl. Mittel-zur. 5 6): 2237 & 22h0, fig. 3171. 1927; Roads, Ohio Journ. Sci. 28: 211. 1928 5 1962 Moldenke » Monograph of Verbena 431 our, Host Ind. Fungi N. Am. 587. 1929; Blossfeldt, Art Forms vane 30. -- Nature pl. 39. 1929; Stapf, Ind. Lon 1931; Gier, Ss. Kans. Acad. Sci ay 261. 1931; B ot. Gard. a1: ri6; 1932; a aya ries & Plains 678. aaas Perry, 343503 35h, 355, pl. +3: fig. 9-17. 1933; Je . * Smal is Southeast He eee 1139 & 1508. 19335 Junell, "one De ae. pial: li--16, 19, 16h, 166--168, 170, 171, & 210, fig. 8, 11, & 22. 193 Hebert, An. uid). Nat. 15: 339. 1934; Dornon, Wild 1934; 73 38: 385. 1938; Woldenke, Alph. List Common Names 10, 25, 26, 29, & 32. 1939; Moldenke, Brief Course Syst. Bot., ed. 2, 20. 1939; C. C Suppl. List Invalid Names 7--9. 191; Gates, Weeds Kans. 219. 191; Worsdell, Ind, Lond, Suppl. 2: 186. 19); G..N. Jo ones, Onio Journ. Sci. 42: 209. 1942; Moldenke in Lunde ell, Fl. Texas 3 (1): 18 & 36-—-37. 192; Moldenke, Known Geogr. Distrib. Verben- ac., [ed. 1], 3-8, 10, 11, 13, 1h, bh, 7h, & 101. 192; Molden~ ke, Alph. List Invalid Names , 6, 7, 25, a 5--50. 1942; Molden- Seymour, New d. Encycl. 1279. 19h; Schnack & Co + ana 6: 169, | un, tly & “Wis, fig. 3-4 & B (29bb) and 77 TL. 19455 Moldenke, Bot, Gaz. 159~-161. 1945; Moldenke, Castanea 10: 36—39. 1953 wsidenke, “am. Journ. Bot. 32: 609 & 610. 195; G. N. Jones, Fl. Ill. [Am. Midl. Nat. Monog. 23] 215. 1945; Darling- ton & Janaki pik ier ge Atl. 270. 1945; Roig, Plant. a, 107, 1io, 411,°119 ‘st, 1 126, 129, 140 We 9, 152, 153, 1572-160 166, 175, 181, 183, 192, 193, ob 197, 201--20h, aor ’ , ot s Suppl , 9, 10, & 22-25. 1947; Moldenke, Phyto- logia 2: 326—- —329, 3h7, "382, 7 383. 1917; Daniel, Verb. Cent. Antiog. 2. 1947; Moldenke, Castanea 13: 112. 19hb; H. N. & A. Le 432 PHYTOLOGIA Vol. 8, no. 8 Moldenke, Pl. “eae 49, 56, & 68. 1948; Moldenke, etl ee? arene 31. i , Trans. Kans. Acad. Sci. 50: 198. eg: Lite a al dy 50 Lh, » 547, 549, 559, 560, 568, 569, 571, 572, 57h, 577 579-581, 5%, 592, 59h, 598, 605 9, & aly bo, a 9 WL ‘° Mh Ler) ay Ww VL Lee) o- ~ Wr pa fa 657, 660, 665-668, 680 682, 683, 698, 699, he ao lteg tees. Toes 738, 747, 7 , aes ’ 74, re 776, 783, 7 S, 788--79 ’ 792, 7 ’ ’ 798; 800, 801, 803, 805, 806, 809, 810, 822, 826, 628, 831, 90h, 952, 953, 96h, 970, & 978 (199), kt 979, 985, 986, 989-993, 997, 998, 1002, 1003, 1010, 1013, 1085, 1086, 1105, 1107, 1109, ii} 113 2 h7, 1ii9, ; 1 11 1187, 1188, 1191, li92, 1197, 1200, 120, 1205, 1207, 1210, RE 121), 1217, 1218, 1220, 1221; 1226, 1228-1230, » 1233, 1235, 1 12h3, 1248, 1252, 1255° 1289, 1291, 129k, 2997, 19493 He Ne & A. L. Moldenke, Anal. Inst, iol. Mex Xe ea. “sles ee 1d d. 2}, 12:31. | 28, 19,235 9h, 97, 99-101, 105, 122, 163, & 197. 1949; Moldenke, Phytolog- ia’ 3: 72, 79, & 132 (1949§ and 3: 28h, 374, & 382. 1950; Fern. 209, 1211, & 1612, fig. 169. in A. Gray, ‘ ag: O008, 1209, & 16 1950; Stefferud, How to Wild Fls. 100. 1950; N. Y. Bot. Gard. Seed Exch. t 1951: 4. 1950; Kennedy, K Jy Let (asst) er h: 67 & 68. 1952; Moldenke in Gleason, New Britton Fl., pr. 1, 3: 127, 131, & 134. 1952; Deam, Yunck- end ner, ° e Bot. 1: 799 & $09 (1953) and 2: 1114. 1953; Moldenke, Phytologia h: 185 & 186. 1953; Moldenke, Inform. Mold. Set 8 Spec. [4]. 1954; Moldenke, Journ. Calif. Hort. Soc. 15: 80. 195h5 gr eR Macmill orne, » Nat. 52: 313. 195k; Buser, Proc, Ind. Acad. Sci. 63: 70. 195k Yoldenke, Midl. Nat. 59: 3h3—3hh, 349, 350, 357, 358, & 363- 19585 itton & Br. Illustr . J sh-Brown, Am. G iad rev. ed., 383. 1958; Moldenke, Phytologia 6: 330. 1958; Straus~ baugh & Core, Fl. W. Va. 3: 77h, 776, & 777. 19585 Shinners, 8 Pp. Andrieux & Cie, seed a 82, 30 50 & bol. a 1962 Moldenke, Monograph of Verbena 433 Spre . Ge - Nat pl. 1, fig. 23. 1793; Usteri, Ann. t. 9: pl. 2. 17943 Curtis, Bot. Mag. 9: pl. 308 (in color). 1795; Moriarty, V. Col. Plates Greenh. (in Sims coo cree Bot. Mag. 48: pl. 2200 (in color). 1820; Morris, Fi. spic eae ‘es color). 1825--1830; Maund, _ Gard. 3: pl. oe a ar as —-1830; Mag. Bot. & Gard. 2: pl. 38 (in color). 183); a Mag. is Aw (in color). 18365 aaak, Brit. Fl. Gard. 7 [ser. 2, h]: pl. 363 (in color). 1836; Edwards, Bot. Reg. 23: pl. 1925. 1837; a Hist. Nat. Vég. Phan. Atlas pl. 8 (in color). 186; Schnitzlein, Iconogr. 2: pl. 137 ian color). $ Dle val . 2 79. 191; Bois, “atlas Pl. Jard. pl. 227 (in “tolor). 1896; i aby. Hort. 1896: 188. 1896; Step, Fav. Fls. Gard. & Greenh. 3: pl. 2 (in es 18973 Britton & erat Ti gate. Fivy. Oded 3h te ot 3063. 1898; L. Cycl. Amer. Hort., ed. 1, 6: 1916 (1902) and ed. i 1916 i ete Wettstein, Handb. syst. Bote, ° ed. 2, 739. 1911; Britton & Br., Illustr. Fis, 2 ee BY f 3858. 1913; L. H. Bailey, Stand. Cycl. Hort. 6: 3ui7. 1917; Hegi, Ill. Fl. uittel-sur. 5. (3): 2239, fig. 3171. 1927} rae 26. 1936; Sichnbolk 'ee-€ 6: th, fig. 3A & Be 19h; Life 2), fai): [62] Gin sen 9 oles Moldenke, Am. Wild Fls. pl. (in color). 1949; Jaques, Pl. Fam., ed. 2, fig. Ui *; & Br. Illustr. Fl., pr. 1, 3: 134 (19 1952) and pr. 2, 3: 13k. “1958; Strausbaugh & Core, Fl. W. Va. 3: 777. 1958. Erect or ascending herb of a pag a habit, to 60 cm. tall, annual in the northern part of i rennial in the south- ern part; roots fibrous; rhizomes sometimes present; bark green; rooting at 43h PHYTOLOGIA Vol. 8, no. 8 ares about 1.3 cm. wide; calyx “aon rie 10--13 mn. , its lobes very slender, subulate-setaceous, u r pu: & hatjadicet bright-lavender, lilac, pinkish-lavender, ec gy ore er, orchid- colored, soft pale blue-violet, reddish-violet, purplish, purple, or blue, sometimes varisgated, opel lavender or purple with a reddish eye, its tube about twice as long as the calyx, varying from glabrous to finely pubescent or lananias on the outer surface, its limb sometimes orchid-purple (HCC.31), 11- : rpm cbt iesapiacoe base, reticulate-scrobiculate from the apex o the base, the commis sural faces muricate-scabrous; chromosome adie an = 30. This common species grows from Illinois to Tennessee, southern Pennsylvania, Virginia, and Florida west to Colorado and Texas, is sa have been given to various forms, but they do not seem to be suf- ficiently schatant to warrant nom ne i ae designation. In the horticultural trade, for instance, plants with dark-violet pete las are know as var. atro visiases those with compact habit ar var. compacta; those with sopacisiiy large corollas are var. grandiflora; those with bipinnatifid leaves and the calyx only 10 mm. or less in length are var. drummondii; and those with nar- es pinnatifid lanceolate or ovate-Lanceolate leaves, cuneate at e base, and the calyx 12 mm. or more in length are var. Lamberti (anbert rosea is said to have paler, fragrant, larger flowers}. Typical V. canadensis has pinnatifid broadly rei: to deltoid pape Feats or broadly truncate at the base, the calyx 12-- . long. Small differentiates these forms (as scestas from sae near relatives as follow 1. Corolla rose-purple, pink, 2. Leaf-blades teukneds Lieed, Poked! or pinnatifid, the see~ ments bro 3. Tip of the longer calyx-lobes subulate, 1 mm. wash ay te Vy. mari V. ae 3a. Tip of the longer calyx-lobes bristle-like, 2--3 mm. long. 4. Leaf-blades pinnatifid or inci sed-pinnatifids cocci 265" rat - long, stout, pitted from near the base to the a- 5. teai-tiades once pinnatifid; calyx 12 mn. long or lon- 6. Leaf-blades eagbomeaes to arabenlanenne + cuneate the Ve lambertii- ae por pager ad 6a. teat-olades 6 broadly ovate to deltoid, senda or a broa neate at the Samesk ays Aald anadensiS« Sa. of eiad ou bipdimasisias calyx 10 mm. ea or lesS-s eeonee 1962 Moldenke, Monograph of Verbena 435 drummondii. vy. ha. rite Bey serrate or incised-serrate; cocei about slender, pitted from about the middle to the tam apex o-Ve pensis. 2a. Leag-blades ipiteiatédect; the aie terns linear to Ve tenuisecta. la. cebote red siVu peruviana. Among the numerous common names recorded for V. canadensis are the following: "clump berbena", "cl bena", "clump vervain", "common verbena", "common wild bena", "cut-leaves rose ver=- vain", "cut-leaved vervain", "flowering verbena", "Lambert's ver- vain", "large flowered bd as "large-flowered verbena", "large flowering verbena", "large flower verbena", "perennial verbena", "pobladora", "pobbeeten", "purple-flo wered verbena", "purple verbena", "Rocky Mountain vervain", "rose-coloured ver= vain", "rosenrotes Eisenkraut", "rose verbena", "rose vervain", "small-flowered verbena" [surely in errort], Ngweet William", tyerbenen", "verbenia', "vervain", "verveine & bouquet", "verveine 4 bouquets", tverveine citronnelle", "ver— veine de Drummond", "verveine de Miquelon", "verveine de Miquelon r Indes", "verveine du Canada", "verveine du Pérou", and "wild ver- bena", It must be pointed out, however, that the pte etre and "vervain" are applied to many other species of the g and to the genus as a whole — as well as to species. of sained genera [see under V. bonariensis L.]. Specimens have been a aee angustifolia Michx., V. bipiinnatifida - Nutt., Ve ip Sas Nutt., V. bracteosa Michx., V. chamaedrifolia st ES ciliata Benth., V V. denticola Small, ¥s erinoides Lam., V. halei i Small, ' Vv. rida var. compacta Hort., “y macdougalii Heller, Ve —T. officinal- is L., V. phlogiflora Chan V. pumila Rydb., V. stricta stricta Vent., i is Nash, V. venosa Gill. & Hook., and V. r, xutha Lehm. The V. bracteosa x Se ecifalsa of Eggert, of Rydberg, and of Stevens are all xV. perriana Moldenke; V. lamberti Ker is V. ele Bans H.B.K.; Ve Lamberti Spreng. is V. lasiostachys 3; Vv. aubletia var. maritima Curtiss is V. maritima Small; V. eeiakia Jacq. x V. tenera Spreng. is xV. wingei Moldenke; and V. canaden- sis var. ~ehrenbergii Thell. is Vs elegans H.B.K. ~~ Rttmpler (1873) calls this plant "Aubletiaartiges Eisenkraut", "verveine de Miquelon", "verveine & bouquets", and "Aubletia~like vervain", His description is "Einjtthrig.-- Mehr oder weniger e ige, von G auf verstelte Pflanze mit vierkantigen, ausgebreiteten, dann aufrechten, 30--50 Centimeter hohen Aesten. on So npr ed stark genervt, lt&nglich h-eirund, halbgefied- ert, mit 3 ungleich gez%hnten Lappen. Blttthen zahlreich, sitzend, in aheas mehr oder weniger dichten und doldenftrmigen, dann 4% PHYTOLOGIA Vol. 8, no. 8 er ee Aehren; Kelch behaart, rthrig, gegen 1 Centimeter lang, mit 5 kleinen Z%hnen und mit einem linienftrmigen Beckblatt- getheilt, von denen 3 grtszer; im Schlunde ein kleiner Kranz weiszer Haare; Staubgefusze , odr 5, eingeschlossen; Griffel dttnn, ebenfalls von der Rthre eingeschlossen." He lekctes on page 1260 var. drummondii Hortorum (which he calls "lilafarbige Abart" as PM Aes "Unterscheidet sich von der vorigen nur durch ihre wolebieklateuiges Blumen, welche zu ge- Pen Stunden des Tages schwach dufte f the species he says "Diese i chene und ihre Abart sind durch den Reichthum und die Dauer ihres Flors ausgezeichnet, welcher in Juni beginnt und je nach der Zeit der aaa sich bis zum Oktober verlungert. Sie sind zur Bildung kleiner und groszen Gruppen, zur Ausschmttckung der Rabatten, wie auch zur Anlage htfbscher Einfass- ungen sehr beliebt. Man sie sie: -- 1. Im Murz-April in das Mist- beet, pikire sie in das Mistbeet oN in TUpfe, die man in das Im September auf ein Gartenbeet; man pikire die Pflunzchen in Schalen, die man unter Glasfenstern tierwintert und pflanze sie im Mai mit 0--50 Centimeter allseitigem Abstande an den Platz." The J. B. H. sen. [Bad Lands, June 1886], Carleton 177, Demaree 12003, Geyer 107, GC. S. Sheldon 323, B. B. Smyth 306, L. L. F. Ward s.n. (Fort Worth, Sept. 9, 1077], and CG. W Wright ight 1502, dis tributed as V. aubletia or V. ee are actually ve peipiaeeti Nutt.; T. S. Brandegee 5, Je M. Coulter s.n. [Denver, May 17, 1873), Je aly H. Cowen 403 & s.n. wi i June 1h, 1892], and 0- Be Metcalfe 1090 are V. ambrosifolia Rydb.; S. B. Parish s.ne [July 1882] is V. ciliata Benth.; Goldman 2, C. R. Orcutt routt 1191, Pringle 11843, and Tharp 1826 are ve. delticola Small; J. E. J. E. Johnson sone [s. Utah, 1876], Lemmon & Lemmon sn. [Peach " springs, June 188h], G. T. Mohr sen, (Gila River, Feb. War. 1873] & s.n. [Plains of Sonora, 20-29-80], Edw. painer s.n. [Southern Arizona, 1869], Purpus 6061, and Toumey 305a & s.n. [Grapevine Cafion, July 30, 1891] are all V. gooddingii Briq.; Griffiths 3489, Hough S -Me [Sabina Cafion, April , 1696], Baw. Palmer 307, C. C. Parry 55 in part), M. E. Spencer 92, Toumey s.n. son. [Camp Verde, Aug - ug. 7 1891] & s.n. [Sabina Cafion, April , 1896], and G. R- Vasey Sele (Tucson, ~1881] are all v. gooddingii var, nepet a Tidestr -5 and Parry, Wright, & Schott s.n. [Frontera, Mar. 2 , 1852] is 4 cotype collection of ve wrightii A. Gray, while A . ‘teach 20, He Manning 33, and S. Taylor 27 are V. tenuisecta Briq.; J- Le "carl- son s.n. (Douglas, May 13, “1915) is V. ciliata Benth. or v. = brosifolia f. eglandulosa Perry; Liebmann 11313 & 1314 and Neal- ley 117 are V. cameronensis L. I. Davis; Tharp harp 5602 is 2 is Ve ciliate 1962 Moldenke, Monograph of Verbena 4,37 var. longidentata Perry; E. J. Palmer 11843 is V. pumila Rydb.; C. Thomas s.n. [Colorado Terr., 1869] is in part | V. bracteata Lag. & Rodr. and in part V. ambrosifolia Rydb.; Collector undesignated s.n. [Partridge Creek, April 26] is in part V. gooddingii Brig. and in part V. ciliata var. pubera (Greene) Perry; and C. He Barrett 13, Beckwith 772, and Mattern & Mattern s.n. [Titusville, ua 3y * 191] are are V. ‘, tampensis is Nash. ur hybrids involving this species are known thus far: V. iebtaats crossed with V. ambrosifolia f. eglandulosa Perry y is xV. fecunda Moldenke, crossed with V. bipinna tifida Nutt. is xV. oklahomensis Moldenke, crossed with V Vv. racemosa Eggert is xV. ferax Moldenke, and crossed with V. tenuisecta Briq. is Rei wingei Moldenke. I regard the V. canadensis var. compacta x x am— brosifolia f, eglandulosa Derman and the v. canadensis var. atro- violacea x ambrosifolia f. eglandulosa Dermen as xV. fecunda, too, and the V. canadensis var. compa acta x racemosa Dermen and and the ve. canadensis var. atroviolacea x racemosa Dermen as xV. ferax. The V. canadensis var. candidissima Haage & Schmidt is treated ated by me under ve canadensis f. candidissima (Haage & Schmidt) Palmer & Steyerm., whic Bartram GIs) per bei ve canadensis at Pointe package Louisia- and describes it o n page 436 (not "136" as ted by Raf ote: Suppl. Fl. Lu rit gt 139. 1817) of his *travels" as fol- lows: "here is likewise a new and beautiful species of Verbena, with decumbent branches eae lacerated deep green leaves; ; the | branches terminate with corymbi of Violet blue flowers, this pretty plant grows in old fields where there is good soil" and blooms in Autumn. The species was introduced into cultivation in or about 1770. It has been found growing at altitudes of 30 to 2800 feet, blooming from January to deeiber: and fruiting from Febru- gravelly limestone, red clay, red sand, s loam soil in general, in dr. land and rocky ground, on n prairies, sand granite outcrops, dry shale bluffs, and es, on dry banks and knolls, limestone barrens 1 and hillsides and open slopes, in wet meadows gravel clay along railroad tracks and soil, on r dry rid railroad embankments, in rich or moist sandy wo and oak-eln soil, along log train tracks, on rocky-s tch banks, in orange groves and sandy cre Nore on low hills, 438 PHYTOLOGIA Vol. 8, no. 8 dunes, and limestone ridges or hills and bluffs, limestone thick- ets and es, along roadsides and roadside ditches, at the edges of fields and prairie pastures, in grasslands and hammocks, along sloughs, in stubble fields and roc » on open bank ridges or dry rocky fertile hillsides, in meadows and dry waste land, on low wooded eous intrusive areas, in rocky woods and cleared nd ou e ridges and dry rocky hills, and in the foothills. Chase calls it a "weed" in cornfields in Illinois, while Jones (1945) says that it grows in open woods from Menard County southward, blooming from May to August there. Doellinger, Moore, & Barkley found it "on Boone limestone, Eureka formation”, In Kansas Hale says it grows along roadsides and in rocky places, while Bailey & Bailey refer to it as "common and showy" there. Houghton found it to be "common dant in Cameron County [Texas]; it covers large areas of open ground and fields; flowers in terminal panicles; leaves dissected into many lobes; fruit a cylindric capsule." His no has in Texas, while Furlong describes it as "occasio sar roadsides" and Marsh reports it "common in woods under tree there. In Oklahoma it was found by Hopkins in a "dry steril in clearing of oak-hickory forest", while Waterfall found clay and shale in crevasses of mountainside" and "in sand at edge of post oak woods". Godfrey says that it inhabits grassy strips 1962 Moldenke, Monograph of Verbena 4,39 along open fencerows, pits open bottomlands, and pine woodlands in Florida. In Iowa it grows among viney pea, * wood-sorre 1, and violets according to West, while in Alabama it is said by * cooley and the Harpers to be Mot common on steep cera Huss found it in sand in the "little bluestem association" The type of V. aubletia grandiflora is Haase & & “Schmidt 5917, collected in cultivation at Erfurt, Germany, in 1898 or earlier. The Deam 21736 and Ek s.n, [October 3, 1936], from In are both said to be shea from cemeteries", and Gates 13773, from oecrers County, Michigan, has a note on the label stating that it cul ti that "this verbena is common along the track from $ - Louis to Galveston", but his April 13, 1880, collection was adi ed from front yard" in Wabash County, Illinois. Davis & Davis snr = it is "a hardy creeping rockgarden verbena bought as Vv. venosa and cultivated at Morgantown, West Virginia. It appare ntly es- es- capes from pes aise in other yg: og too, because Gibbes found it on "hills n.e. of Hamburg", and B Beauregard re- ports it wtih cult. fields" in France. “Hoes eel. (1927) says “verwil- dert in der Mark Brandenburg unweit Nauen." Jansson's collection rom plo gy gti apparently from escaped “material, bears a note "native in the southwestern states." The L. H. Bai ley sen. [Sept. 30, 1926] bears a notation on its label stating "gland on back, anthers .§ appendaged", but Churchill s.n., according to Bailey, ha s them "not appendaged”. Lawrence reports that his P.58) has ike connectives without gland." MacDougal (1907) suggests that V. canadensis may hybridize in the wild with V. bracteata Lag. & Rodr., but he cites no examples, ives no description of the eg hybrid, and does not say that he actually saw any such hybrid plants. Britton & Brown (1898 and 1913) make the same statement as a fact, but there are no such hybrid specimens in the Britton Herbarium *at the New York er hybridize with each other, and certainly have ry er seen any specimens which could be regarded as such a hybri T sen. [June 6, 1932] has massive ee and the flowers exceptionally large. McBryde s.n. er - also anom- — in having large, thick, s Pern fly incised lea reminding ne of V. tampensis Nash. The L. H. Bailey s.n. anion ail Ai 30, 1919, July h, 1919, July 33, 1985» Ang Fort 7, 192, 22, 1926, ise 5, 1926, September yee and October 8, nee also have the leaves especi yas owly lobed. Tracy 8707 looks anidaxt like V. rersvecke: but apparently is not that species. Burk s.n. (Girard Point is a mixture with V. Everett & Quinn s.n. [N. Y. Bot. Gard. Cult. Plants a) is called : the "red form" on its label, while Quinn s.n. (N. Y. Bot. 40 PHYTOLOGIA Vol. 8, no. 8 Gard. Cult. Plants R.38/36] is the "purple form", The Herb. Calif. Acad. Sci. 31446 does not actually say "cultivated" on its 1 its label, but I assume that it represents cieteeiad material. Croom sene, from Florida, distributed as V. canadensis, is actually Phlox pilosa var. virens (Michx.) Wherry in the Polemoniaceae, “while A. S. Hitchcock s.n. [Fowler, July 1, 1892], also distributed as this species, is Chamaesaracha panties (Moric.) Britton in the Solanaceae. The verbenaceous portion of Demaree 17004 is anomal- ous -- it looks like a cross with V. bipinnatifida da Nutt., but its corollas are smooth (not hairy, as >in xV. oklahomensis Moldenke) . Van Vleet s.n. [Mt. Sheridan, 7/4/03), however, is apparently a mixture with xV. oklahomensis. The parents of this hybrid were probably growing in close sed to it, accounting for the mixture. Cain (19) notes that from xV. wingei Moldenke (which is the cross between V. canadensis and V. , Sormloacte Brig.) "it is possible to peeceents both parents in: pure fo R. W. P a letter to me dated pesesel 27, "1951 says of v. canadensis: it Mis found in this vicinity [Ames, Iowa] either Vi sonitttork? The label on Winterringer 5331 31 bears this notation: "evidently trash had been dumped here". "Hibbard 87 was first i- dentified as V, bipinnatifida, then as V. bracteata, and finally as V. canadensis. Hitchcock & Stanford 6782 is very * goubtfully this | species, V. canadensis was recorded by me in my "Known Geographic tribution", edition 1, page 1) (1942) from Pima Coun unty, Arizona but this record was based on Stiffler sen. [1925] in the cornell University herbarium, which has since proved to be Ve. gooddingii Dis- ya actual: - have | rom to McVaugh (1956) ae indicates that Torrey included Palmer's Or nal nos. 260 and 390 in his "247" — 260 was collected in Gar vin County on June m4 while 390 was collected on July 1-either in Atoka or in PRET Ee Coun ¢ is perhaps worth noting here that the specific epithet of this species is very often written with a capital initial letter, but this practice has no basis in Latin grammar nor sreniy a ee sana: County, Arkansas "Ho Springs County". “Banee 98h) records V. canadensis ‘trom Fountere County, Indiana; Hebert (193) from Berrien County, Michigan Roads (1928) from Pike County, Ohio; a (1 19365 stot he Anne County, Virginia; C. N. Jones (1942) from Darke County Ohio; and Deam, Yuncker, & Friesner (1952) from Blackford and Delaware Counties, Indiana. As yet I have seen no mat terial from 1962 Moldenke, Monograph of Verbena hi these counties, but have no reason to doubt their reco The reference to Loud., Hort. Brit. Suppl. of on a), yg i ofte Baxter, but apparently in error. The "Verbena Shas iiy ra beef of Dermen, Cytologia 7: 162--163, at 7 (1936) is apparently an error for. V. grandiflora Michx., since Ortega's name applies to a species of P. Priva that would igedity ne been mistaken for a Ver- bena by Dermen. ~~ Verbena canmiikte is often attacked by the fungi Dicaeoma verbenicola Arth., Erysiphe cichoracearum DC., and Septoria ver~ benae Rob. In the Linnean Herbarium, sheet number 7, under Buchnera, is cag "Buchnera canadensis" in eee ae own handwriting, while note in pencil in Smith's handwriting V. Aubletia J. E. $. " Sheet number 8 under Buchnera is lubeled "obletia" in Linnae- us' handwriting and bears a supplementary not e "Touretta" and an unidentified plate, with figures 1 to 7, caated rd it. Jackson does not know the source of this plate. the specimens in both cases are definitely Verbena canadensis, Schauer ia A. W., Prodr. 11: 55, (1847) credits the name Verbena aubletia to Linn- aeus, but Linnaeus in the reference to which Schauer refers, Suppl. 86--87 (1781), plainly credits it to "Jacqu. hort. v. 2. oo ua. t, Las” a a: Phe to note that at the nd of his treatment of this inus peruvianus an hujus varietas?" Pdiset: 73808) aches a6 latte Rozier, which he in- cludes here, to "Journ. d'Hist. Nat. Nat. (1771)". Verbena intermedia Penny is listed in the "Index Kewensis" with an "x", indicating hybrid origin. It is said by Don to be Ac aoynmeatan between V. aubletia and Vv. lamberti Penny", but he rtainly does not claim hybrid origin | for it! The name Verbena sree rected is cited by me in Lundell, Fl. Texas 3 ti) (1): 36, syn. (1942), to "G. Don in Loud., Hort. Brit. Suppl. 2: 680. 1835, but the pied Kewensis" affirms that W. Baxter was re~ sponsible for it there. The original description of V. aubletia var. drummondi pare is simply "floribus lilacinis suaveolent- ibus," Seed of V. canadensis is handled by many dealers, including Borsch, Cronamere, Poughkeepsie, Stumpp & Walter, Breck, and Weston, as well as Wayside Gardens of Mentor, Ohio, Gardenside Nurseries of Shelburne, Vermont, and W. H. Simpson of Birminghan, England. Vilmorin-Andrieux & Cie., in Paris, offer the drummondii form; Huntington offers grandiflora; Schling offers compacta; Sutton offers drummondii; while Haage & Schmidt in Erfurt, Germ any offer atroviolacea, compacta, drummon ndii, and montana. Vilmorin-Andrieux give the following cultural directions on their seed packets: "Semer: 1° en auot-septembre en pépiniere, en pots ou en terrines; repiquer en pots; hiverner sous ch&ssis ou en sierre tempérée au pius pres de la * umiere; mettre en place en 4h2 PHYTOLOGIA Vol. 8, no. 8 fin avril-mai. -- 2° en mars-avril sur couche; repiquer en pépin- iére ou en pots, sur couche; mettre en place en mai (ou mai-juin). — 3° en avril-mai en place ou en pépiniére. —- Fl. de juin- juillet en octobre." The species has been in cultivation in Eng- and since at least 1799 according to Murray (1799). Roig (1953) says: "La Verbena canadensis Britton, es muy emple- ada en jardines para cubrir los canteros a mane - bros, y por la rapidez con se extiende y cubre el suela muchos la di peng th. ago de las Vegas, y en ot pueb- las de la provincia de La Habana.....cultivada como ornamen los jardines. Tiene las flores en umbelas de diversos colores." I have never seen any C material of this species and am fair its "Prairies, fields, glades, and rocky slopes. Calciphile to circumneutral. General, i Mo." Schauer (187) cites a Lindheimer s.n. from Texas, Douglas s.n. from Louisiana, Frank s.n. from Ohio, and Beyrich s.n., Fra~ ser s.n., and Walter s.n. from "Carolina", all deposited in the Berlin and DeCandolle herbaria. He says "Forma vel varietas — major, speciosior est V. Aubl. var. Drummondi Lindl.!....Planta taria hortis europaeis familiaris, ad affinibus foliis minus dissectis discernenda." Walpers (1845) classifies it in hts Sect. Verbenaca, Subsect. Inermes, Group Foliosae, Subgroup Mac- ranthae, and Secondary Subgroup Aubletia with ten other species. Schauer places it in his Sect. Glandularia. Perry (1933) cites the following 10 additional specimens not as yet seen by me: NORTH CAROLINA: Vance Co.: Sturtevant S.M- (Kittrell, 11 April 1889] (E). SOUTH CAROLINA: Abbeville Coe? Hexamer & Maier s.n. [Abbeville District] (G). GEORGIA: County undetermined: Rugel s.n. [lower Georgia and middle Florida] (E)- IOWA: Pottawattamie Co.: G. R. Vasey son. [Council Bluffs] (G)- KENTUCKY: Shelby Co.: Flint s.n. [Shelbyville] (G). TENNESSEE: Rutherford Co.: Eggert s.n. [ May 1898] (E). KANSAS: Cherokee Co.: A. S. Hitchcock 790 (E, G). Miami Co.: Rydberg & Imler 22 (E). MISSOURI: Barry Co.: E. J. Palmer 2980 (£). Carter Cot Be F. Bush 318 (E). Henry Co.: A. S. Hitchcock s.n. [Windsor Springs, 30 May 1890] (E). Iron Co.: Drushel 2762 (E)3 Engelmann s.n. [Pi~ lot Knob Creek, 9 Sept. 1859] (E), sen. [Pilob Knob Creek, July 1867] (£); C. C. Epling 613 (E); Greerman 3870 (E), 4076 (8)+ Jackson Co.: Broadbent sen. {28 March 1864] (E)3; Be Fe Bush 283 (E, G), 1933 (E), 2901 (B, G), k922 (EB), 6439 (E)s Tindald SDs (Independence, 1 June 1895] (E); Trelease 1161 (E). Jasper Co? B. F. Bush s.n, [16 Aug. 1893] (E); E. J. Palmer 30h (B), 357 1891] (B), 2996 (B). Jefferson Co.: Drushel 3659 (£); Eggert S-De [Crystal City, 19 Aug. 1886] (E), s.n. [Crystal City, 6 May 1962 Moldenke, Monograph of Verbena 43 (E, E); A. S. Hitchcock s.n. [Victoria, 10 May 1890] (E); Johnson s.n. [Kimmswick, 30 April 1905] (E); F. Wislizenus 285 (E). Madi- son Cos: Greenman s.n. [Mine La Motta, 19 May 1927] (E), sn. [near Silvermine, 20 May 1927] (E). Phelps Co.: J. H. Kellogg s. n. (Jerome, 5 April 1914] (E). Saint Francois Co.: Trelease 71 (E). Shannon Co.: B. F. Bush 1168 (E). Stone Co.: E. J. Palmer 5682 (E), 24616 (E). Taney Co.: B. F. Bush 569 (E). Washington Co.: J. H. Kellogg 1958 (E). Saint Louis: Eggert s.n. (Allenton, 27 April 1887] (E), sn- [16 May 1879] (E); Engelmann 33h (E); Sherff 333 (G). County undetermined: Eggert s.n. [Silica, 18 A- pril 1896] (E); Greenman 231 [Shot Hill, near Selma, 30 May 1923] (E). ARKANSAS: Carroll Co.: E. J. Palmer 29333 (E, G); Trelease s.n. [Eureka Springs, 19 April 1899] (E). Garland Co.: Trelease s.n. [Hot Springs, 31 Oct. 1899] (E). Hempstead Co.: E. J. Palmer 507 (E). Hot Spring Co.: Letterman s.n. [Malvern] (E). Nevada Co.: B. Fe Bush 545 (E). Sebastian Co.: Benke 566 (G). County undetermined: Pitchard s.n. [Red River] (G). LOUISIANA: Avoyelles Par.: Hale s.n. (G). Cameron Par.: S- M. Tracy 8707 (E, G, N). East Feliciana Par.: Drummond s.n. {Jacksonville] (G). Natchitoches Par.: E. J. Palmer 7001 (BE), 7071 (E). Webster Par.: Trelease s.n. (Minden, 15 April 1901] (E). COLORADO: County unde- termined: G. R. Vasey s.n. [Rocky Mountains, lat. 0-51', 1868] (E, G). OKLAHOMA: Cleveland Co.: Bmig 415 (E). Creek Co.: Be Fe Bush 37 (E, G). Garvin Co.: G. W. Stevens 109 (E, G)- LeFlore Co.: Blakeley 1h71 (G). McCurtain Co.: Houghton 3643 (E, G). Muskogee Cos: E« J« Palmer 14283 (E). Ottawa Co.: G. W. Stevens 2340 (E, G). Payne Co.: Waugh 168 (E). TEXAS: Bowie Co. Eggert sen, [T. & C. Junction, 5 Sept. 1900] (E, G). Dallas Co.: REVer™ chon 740 (E), 2533 (E), sen. [Dallas] (G). Galveston “ Van Huff s.n. [Galveston] (E). Grayson Co.: Reverchon s.n. [Denison, 15 Warch 190] (£), sen. (Denison, 22 April 190L] (E). Gregg Co.: Eggert s.n. [Longview, 21 April 1915] (E); E. J. Palmer 712) (E). Harris Co.: G. Le Fisher 707 (W), sm. [Houston] (E); E. Hall 435 (E); Lindheimer s.n- {Houston, Feb. 182] (G). Houston Coz: E. d. Palmer 12056 (E); Tharp 836 (G). Liberty Co.: E. J. Palmer 11979 (E). Montgomery Co.: E. Je Palmer 33319 (E, G). Polk Coe: E. J« Palmer 5185 (E). She comments that nVerbena Canadensis is one of the most easily recognized of the North An- erican species belonging to the section Glandularia. It has a at maturity, an elongated spike with somewhat remote fruits an i Conspicuous calyx-teeth. The leaves are variously lobed or cleft. Although these variations have been used at sundry times to seg- by PHYTOLOGIA vol. 8, no. 8 recognition." The G. L. Fisher 707 which she cites is doubtless a typographic error for his no. 1707, cited below. In all, 1183 herbarium specimens, including type material of most of the names involved, and 22 mounted photographs and illus- trations have been examined by me. Citations: CONNECTICUT: New London Co.: Jansson $.n. [East Lyme, July 8, 1931] (Go). NEW YORK: Suffolk Co.: John Jones s.ne [Bay Shore, May h, 1951] (N). PENNSYLVANIA: Lancaster Co.: Tanger s.n. (June , 1938] (Up). Philadelphia Co.: Ie Burk s.n. [Girard Point] (Up--17095). Washington Co.: Shafer s.n. [July 8, 1907] (Cm). VIRGINIA: James City Co.: Eggleston s.n. [ Jamestown- Williamsburg, May 16-18, 1907] (Dt). County undetermined: Herb. Wallberg s.n. [1808] (Lu); Stebbins s.n. (Virgin.] (T). WEST VIR- GINIA: Lewis Co.: West Va. Univ. Bot. Exped. 1651 (We, We). NORTH CAROLINA: Alamance Co.: Ahles & Bell 16928 (Hi—92858); Radford 10M) (Hi--92858). Brunswick Co.: C. R. Bell 11601 (Hi--121527) « Cumberland Co.: M, G. Brooks s.n. Trayetteville, April 7, 190] (We). Orange Co.: F. Faust s.n. [10/25/37] (Hi—59l.70). Randolph Go. Collector undesignated s.n. [June 30, 1891] (Hi—-59460). Rutherford Co.: C. R. Bell 2109 (Hi—92956). Washington Co.? Rad- ford 3519 (Hi—104886). SOUTH CAROLINA: Abbeville Co.: O- Me Freanan 5771 (Hi--92955); F. M. Hexamer 289 (Br); Radford 20026 (Hi—9295h). Charleston Co.: K. Hunt 356a (It). Jasper Co.: Ahles & Haesloop 22295 (Hi—92851). McCormick Co.: Radford 20175 (Hi-- 9285). Orangeburg Co.: Ahles & Haesloop 21682 (Hi--16196). i Cos: C. Rs Bell 5696 (Hi—-92957) « GEORGIA: Clarke Co! Barfield s.n. [May 8, 1917] (Gu—E.102); Cleckler s.n- {April 10, 1917] (Gu--E.10)1); K. M. Drewry s.n. TApr. 16, 1929] (Gu-- £.8541)5 Elrod sen. [May 3, 1917] (Gu--B.10h6); Fe Ce Garrett s. n. (May 3, 1917] (Gu--E.10)3); C. B. Ingram sen. [April 30, 1917] (Gu--E.1039); Miller & Maguire 1256 (Ba, Ua—16572), 1257 (Ua— 16573), 1258 (Ba, Ua-—1657); Palfrey sn. [April 15, 1917] (Gu~ E.1045); Pyron s.n. [Simpton's Bridge] (H—-1652); J. M. Reade Se n, [April 22, 1911) (Gu—1176), s.n. [May 8, 1911 (Ga--B.1616) 3 Thornton s.n. [May 3, 1917] (Gu--E.10)0); A. C. Welch s-ne [May 13, 1917] (Gu—E.10h7); Woodard s.n. (May 7, 1917] (Gu--117h7). ugherty Co.: Thorne & Muenscher 2475 (Ca—906198, N, Vi, Lacs 1929830), 9198 (Vi). Hancock Co.: F. J» Hermann 101h9 (N). Jack- son Co.: Clausen & Trapido 3802 (Ba, N). Putnam Co.: Cronquis 4238 (Ca-=789071, Gu-—20028, N, Tl, W--1927967); We H. Duncan 4900 (NN); Pyron & McVaugh 2396 (Gu—16648, H—-52338). Randolph Co.: Thorne & Muenscher 7790 (Gu—-30795). Richmond Co. cuthbert sen. (Augusta, May 1897) (Fl--2110k); L. DeWitte s.n. [August 1638) (31). Sumter cos: Huger son. (Americus, Apr. 1697] (C)i Lawrence & Moore 672 (Go). FLORIDA: Alachua Co.: Murrill 11 (# 1962 Moldenke, Monograph of Verbena Lhs 1846518), 421 (W--19282)3), 422 (W—-19282h)); Van Hyning s.n. [A- pril 3, 192k] (Ob—50811), s.n. [May 2, 1924] (Ob—50763); G. F. Weber s.n. (Gainesville, -1-28] (Fl—20973, F1--2097h). Brevard Co.: A. A. Baldwin 70 (Up--17080). Broward Co.: G. L. Bates sen. [Heath, Feb. 26, 1889] (Ur). Columbia Co.: Rolfs s.n. [Lake City, May 1893] (F1--21133). Dixie Co.: Hume & sen. [Oldtom, 29 March 1936] (Fl~—11)66, F1--20970). Duval Co.: T (Jacksonville, 1833] (S); G. H. M. Lawrence 193 (N)3 Lighthipe 599 ee ee Brass 15267 (W—2065)5). Jefferson Co.: R. K. Godfrey 54436 (Hi— 16081, N); Lighthipe s.n. [Monticello, March 13, 1891] (Ur). Lee Co.: E. C. Marquand s.n. [March h, 1930] (N). Leon Co.: R. Ke Godfrey 5432 (Hi—155443, N). Marion Co.: J. D. Smith s.n. [0c- ala, April h, '79] (W—13230kh). Saint Johns Co.: J. D. Smith 475 (W--1323047), sen. [Mous Creek, April 15, 179] (W—13230h6) . Suwannee Co.: Exploration Party 1939 s.n. [17 Mar. 1939] (Fl— 30166). Volusia Co.: Berkimer s.n. [Daytona, 3--22-'25] (Vt); S- A. Deam 1831 (Ur, W—753721)3 G. B. Grant 3063 (Po, Po—267663) ; A. S. Marsh s.n. [Deland, 1923] (W--1220068) ; B. H. Patterson s- nh. (Daytona, Feb. 23, 1918] (Cm). County undetermined: Herb. Chapman s.n. [Florida] (C). ALABAMA: Jefferson Co.: Cooley, Har- per, & Harper 171 (T1); Harper & Morgan s.n. (March 22, 1941]~ (I). Madison Co.: R. M. Harper 3958 (N, W—-189917). Tuscaloosa Co.: Clausen & Clausen 5753 (N); Re M. Harper 3018 (Ba, Mi, N); E. A. Smith s.n. (Tuscaloosa, May 8, 1875] (W—87684L)5 L. Fe Ward s.n. [5 mi. above Tuscaloosa, April hi, 1892) (W—-147558), sn. [Tuscaloosa, April 1892] (Au—-18)936, W--2558)0). Wilcox Co.: Buckley s.n. [May 180] (T), sone [May] (W--771877). County undetermined: R. C. Alexander s.n. [May 16h1] (Ca—37999k), S.n- Oktibbeha Co.: S. M. Tracy 1659 (W). Pearl River Co.: Ashe s.n. [Apl. 18, 1931] (I). Pike Co.: F. S. Earle Son. [McComb City, A- pril 3, 1878] (cm). County undetermined: Collector undesignated 8.n. (N). OHIO: Clermont Co.: E. L- Braun s.n. (Milford, 5/4/12) (Cn), sen. [Miamiville, 5/23/17] (Cn, Cn). Hanilton Co.: Aiken s.n. [May 13, 1905] (Po—267628); C. G- Lloyd sen. [IV-23-05] (Ll). ILLINOIS: Adams Co.: Brinker 3379 (I1—1585L, Ur). Cham- paign Co.: Honeus s.n, [May 20, 1892] (Ur); L. A. MacGregor S-ne 46 PHYTOLOGIA Vol. 8, no. 8 (May 25, 1892] (Ur); Schwickrath s.n. [July 11, 1955] (Ur); Slim- mer 33 (Ur). Cumberland Co.: Winterringer 5331 (I1--31672). Kane Coe: Springer s.n. [September 6, 1955] (Ur). Lawrence Co.: Sivert s.n. (May 8, 1949] (Ur). Menard Co.: E. Hall s.n. [1861] (Ur, W— 1070016, W--120143), sn. (Athens, June 1866] (Ms). Peoria Co.: V. H. Chase 8710 (Ur), 14197 (Ur). Shelby Co.: G. D. Fuller 12713 (I11~—19209, I1-—-19210). Union Co.: Ahles 3630 (Ur); F. S. Earle s.n. [Pine Hills] (Ur); G. D. Fuller 11630 (I1—19098, I1--19099, T1-—-19100); Winterringer 7059 (I1--3516,). Vermilion Co.: R. Bell son. [June 9, 195] (Ur). Wabash Co.: Schneck s.n. {April 13, 1880] (Ur), sen. (May 1900] (Ur), sen. [May 1h, 1904] (Ur); Shear er s.n. [May 15, 1920] (Vi). INDIANA: Brown Co.: Lipps S-Ne [June 1h, 1942] (Ok). Clark Co.: J. F. Baird s.n. [8-l-78] (Mi). Howard Co.: Ek s.n, [near Greentown, 6-23-1936] (Bt--l.2530), son. [Octo- ber 3, 1936] (Dm). Jennings Co.: C. C. Deam 2):736 (Dm). Monroe Co.: Blatchley s.n. [May 7, 1887] (Dp). Orange Co.: Friesner 22735 (B). Putnam Co.: Yuncker 1380 (Dp). Wayne Co.: Greek 5.ne [Nutter's cem., 8/4] (Ea). IOWA: Story Co.: W. W. Thompson SMe [8 June 1951] (N); J. W. West s.n. [June h, 1951] (Gg--,06217) . KENTUCKY: Edmondson Co.: Tix 363 (W—-19)8018). Fayette Co.? Did- lake s.n. [Lexington, July 15, 1909] (W--592232). TENNESSEE: Da~ vidson Go.: Dermon 12 (Hi--192179); Ge We Hubbard 1963 (Cm, In— 9679, Ka, Vt); C. S. Williamson s.n . [Nashville] (N). Montgomery Cos: T. L. Andrews 151 (Io--32955). Rutherford Co.: Eggert SMe [4 May 1898] (N); Gattinger s.n. [Cedar Glades] (Du--90902); Mor- timer 38 (W~—1814)L9); Quarterman 1638 (Va); Sharp & Shanks 1,39 {Herb. South. Appal. Bot. Club 110 & 11.149] (Au, Au Bt--65318, Dt, Gg—310876, I, N, N, N, Ok, Or—l6592, P1—133778, St--23610, We). Wilson Co.: C. C. Deam 61270 (Dm, N), 61351 (Dm); Re McVaugh 5560 (Ca—72h961); F. H. Sargent 6119 (St). County undetermined Gattinger s.n. [Cedarbottom, June '79] (Vt), SMe [Middle Tenn, Aug. 1879] (Cm), sen. [Cedar barrens] (Al). MICHIGAN: Cheboygan Co.: F. C. Gates 13773 (Ka--679L1). Genesee Co.? Eggleston S-Me {Flint, 5-9-79] (Dt). MINNESOTA: Ramsey Co.: Eggert SMe [Sst- sen. [Anderson Co., July 1896] (Ka); W. H. Horr son. [duly 22, 1929] (Lw). Bourbon Co.: Kellerman s.n. [Fort Scott, July 15, 1887] (Ka); Z. D. Thompson 163 (Lw), 545 (Lw). Brown Coe: Mrs 0. Olson 2 (Ka-—88181). Butler Co.: R. L. McGregor 1048 (Ly). Chautauqua Co.: A. S. Hitchcock sen. [Chautauqua Coo, AUS> Ys 1896] (Ka). Cherokee Co.: A. S. Hitchcock 790 (Ka, Ny, W——353581)5 A. A. Jacobs 50 (Ka—77569), 98 (Ka-—T02)3 Re Le McGregor 1332 (W—D3hS). Coffey Co.: Eilford sen. [Sept. 198] (Ka). Cowley Co.: Clothier & Whitford s.n. Aug. 22, 1897] (Ka); Ryde 1962 Moldenke, Monograph of Verbena Lh7 berg & Imler 508 (lw); M. White s.n. [Apr. 198] (Ka). Crawford Co.: A. S. Hitchcock s.n. [Crawford Co., Aug. 1896] (Ka). Douglas Co.: Derr O (Ka); A. O. Garrett 8452 (Se--1963), s.n. [June] (Ob--50767); D. L. Marsh 1522 (Hi—198639, N); Re L. McGregor 31 (Iw, S), 14289 (Hi—198613, W—2327112); F. R. Schallert Lol (H—-5L57); Snow 1963 (Lw). Edwards Co.: A. Finch 45 (ka—80195). Elk Co.: Clothier & Whitford s.n. [Aug. 20 & 21, 1897] (Ka). El- Co.: C. S. Sheldon 323 (Al). Franklin Co.: W. E. Castle s.n. [April 10, 1890] (Ob—50768); A. S. Hitchcock s.n. [Franklin Co., July 1896] (Ka); Street s.n. [Princeton, May 1), 1919] (Po-- 63842). Greenwood Co.: Q. Gillett 36 (K); A. S. Hitchcock s.n- [Eureka, July 1892] (Ka); W. H. Horr sen. [June 28, 1930] (Lw, Iw). Johnson Co.: W. H. Horr s.n. [Sept. 20, 1928] (Lw); Pellett s.n. (Summer, 1890] (Ka). Labette Co.: A. S. Hitchcock s.n. {La- bette Co., Aug. 1896] (Ka); D. Nelson s.n. [May 3, 192] (Ka-- 71452); Paull s.n. (Oswego, 5/29/1886] (Cm); Rydberg & Imler 323 (Lm), 379 (Lw, N)3 Schneck 97 (Ur). Linn Co.: B. F. Bush 15259 (Ka--89270), 15503 (Ka--09203); Clothier & Whitford s.n. [Aug. 9, 1897] (Ka); Gier 7956 (S). Miami Co.: Clothier & Whitford s.n. [Aug. 8, 1897] (Ka); Oyster s.n. [Spring, 1885; Herb. Prager 18629] (Gg—3145), sen. [May 1686] (Ob—50771); Rydberg & Imler 22 (Ka--7h6l5, Lw, N). Montgomery Co.: Bailey & Bailey 1023 (Ba); Clothier & Whitford s.n. [Aug. 19, 1897] (Ka); N. Be Jacobs 318 (Ka——92925) 5 F. W. Johnson sen. (Cherryvale, Sept. 16, 1900) (N). Morris Co.: Good 4 (Ka—91088), 25 (Ka--91087). Neosho Co.: Broadbent 22 (Ka); W. H. Horr s.n. [7/25/29] (Iw); Maus 1102 (Ka~ 72792, Ka--72793) . Osage COs? Z» De E. Brown s.n- [July 28, 1696] (Ka); F. C. Gates 17317 (Ka~79799)} Re L. MoGregor 1561 (LW). Riley Co.: Burtner sn. [May 10] (Ka); F. C. Gates 12681 (Ka— 72873, Ob—50775) ; Kellerman sen. (Manhattan, May 18, 1884] (Ka); L. H. Pammel s.n. [Manhattan, May 15, 1927] (Ca--351348, Io-- 129735); J. F. Ross sen. [May 15, 1899] (Ka); Walters sn. [Man- hattan, May 2, 1691] (W--22232h). Shawnee Co.: Chamberlin 27 (Pl—35099); G. Hale 35 (Ka--786L5); V. L. Harms 1379 (Hi— 198197, N); Maus son. (Auburn, July 20, 1927] (Ka—72218); Pel- lett s.n. [Topeka, 1928] (Io—13283h); F. R. Popenoe SoDe [Tope- ka, June 2, '80] (Pu). Wabaunsee Co.: A. Se Hitchcock s.n. [Wa- baunsee Co., Aug. 23, 1896] (Ka); Maus 36 (Ka—72175), 239 (Ka-- 72679). Wilson Co.: Haller sen. [Apr. 23, 1896] (Ka); W. H. Horr S.n. [July 12, 1930] (Lw). Woodson Co.: G. M. Clark s.n, [Bates- ville, Aug. 20, 189k] (Ka); M. Hale son. [1 May 1955] (We); We He Horr s.n. [7/11/30] (Lw); E. W. Lathrop 3h (W—223)918), 169 (W— ——<—— —— 2231972), 556 (N), 1028 (N), County undetermined: Herb. ee S.n. (Hp); Herb. Kans. State Agr. Coll. s.n- [June 15, 1591] ( Lh8 PHYTOLOGIA Vol. 8, no. 8 MISSOURI: Bates Co.: Broadhead s.n. [May 1871] (Cm). Boone Co.: Rickett 533 (0k), sen. [Bear Spring, April 23, 1927] (H--106),68). Clay Co.: Broussard 22 (Je—5131); Gier 8106 (S); G. Moon sme [Liberty, 5/14/190] (Je—-3155). Greene Co.: Blankinship S.n» [26 -)-79] (Io—3h748), s.n. (Willard, July 20, 1919] (Po-—636h3); B. F. Bush 286a (Ca—25130); P. C. Standley 8367 (W--687327), 9171 (W--688089), son. [vicinity of Springfield, May 1905] (W—735592) - Iron Co.: Churchill s.n. [Ironton, May 2h, 1918] (Ba); Je A. Dru- shel s.n. [5/28/1916] (Ur); C. C. Epling 6134 (La, Or--20105, Or- 20141). Jackson Co.: B. F. Bush 7 (Up--17079), 286 (C), 2901 (N, Ur), 4922 (W—606h03), son. [/9/1882] (Cn, Cn), Sone [Courtney, 5-19-91] (C); K. K. Mackenzie 62 (Dm, I, N, S), 811 (N); Co Ae Ripley s.n. [May 1890] (Ob--5076). Jasper Co.: Storm 2 (Ur). Jefferson Co.: Eggert s.n. [Crystal City, 19 Aug. 1886] (Cm, Du-- 123487); G. J. Goodman 962 (Ok); Goodman & Hitchcock 962 (Po— 195k); Lloyd Williams 1468 (I). Johnson Co.: G. W. Stevens 4,308 (N). Lafayette Co.: Bailey & Bailey 1026) (Ba). Lawrence Co.? We Flint s.n. (Dt). Madison Co.: J. M. Greenman 8.n- [Mine La Motta, 19 May 1927] (Mn--18638, N). Ozark Co.: B. F. Bush 13329 (Up). Phelps Co.: J. H. Kellogg sen. [Jerome, Apr. 5, 1914) (N); Ee Ce Leonard 6592 (W-=216L960). Saint Clair Co.: B. F. Bush 13298 (8), 13298a (Ka—8)565). Saint Francois Co.: Ahles 2012 (Ur). Sainte Genevieve Co.: Brinker 1533 (I), 1638 (I); Koelling 31 (Ur). ra Louis: H. N. Andrews Jr. ape [/29/36} (Ms); Begert Sone Stony hills, 16 May 1879] (Se—95715, W-—-518722); C. C. Epling 6412 (La); Herb. its en (Ka); Letterman gen, [11-90] (N). Saint Louis Cos: Eegert s.n. (Allenton, 27 Apr. 1887] (Au, Ca--183199, Cm, Dm, Du--122672, Gg—31281, Io—79867, un-—6688 Up—60737); Glatfelter s.n. [Allenton, 5-23-92] (Mi, W308 363) Letterman s.n. [Allenton, May 1888] (Ur); Ohlweiler s.n. [Allen ton, -18-09] (Au); L. H. Pammel s.n. [Allenton, 5-1887] (Io— 15338), son. [Valley Park, 5-1007] (I0—20748), sen. [Glencoe, Apr. 22, '88] (Io—207h7). Saline Co.: B. F. Bush 13439 (Au, S)- Stone Co.: E. J. Palmer 5682 (Po--217515, W--1581050). Taney Co-* W. W. Eggleston 12240 (N, W-—6250). Vernon Co.: Palmer & Stey- ermark 12162 (N, W-—160)823). County undetermined: Collector un- designated s.n. [s.w. Mo., 7-89] (Pl—22610) ; Dapprich 1772 (Sm)5 James s.n. [near the Lead Mines of Missouri] (T); Je Re Lawrence sen. (Apr. 7, 1929] (Ms); Letterman 1 (Po); Fe peck 1963 (Dt), 5 Sen. [Missouri] (Pa). ARKANSAS: Baxter Co.: Demaree 2693 (N)» 289Ll, (Hi--S1L55, Mi, N, Ok, We), 28971 (N, St). Benton Co.t EY ans & Morgan 26 (Ok); Heacox s.n. [July h, '89] (Ob—50766) 5 De F. Morgan 15 (Ok); Plank s.n. [1899] (N). Bradley Co.: Dena’ 24217 (Sm). Calhoun Co.: Demaree 1399 (I, N)- Carroll Co. Del- 1962 Moldenke, Monograph of Verbena yg linger, Moore, & Barkley 15008 (Au, N, N); A. Nelson 10859 (Du 230822, Um——5, Up); E. J. Palmer 29333 (N) Chicot Co.: Demaree 20933 (N, Sm). Cleburne Co.: Demaree 45072 (Ca). Conway Co.: Dem- aree 20916 (M1, N, Sm), 37506 (Sm). Crawford Co.: Demaree 6411 (W—-1,37257); Shearer sen. [Dyer, Aug. 28, 1919] (Vi). Drew Co.: Demaree 16566 (Du—256559, En, H--97668, N, Ok). Faulkner Co.: Demaree 5962 (St); F. C. Haas 1674 (W--1651393); Haas & Haas sn. (April 7, 1934] (Dp). Franklin Co.: Demaree 20890 (N, $m). Fulton Co.: Demaree 25864 (N). Garland Co.: Demaree 16683 (Du--256562, N, Ok, Se——69268). Grant Co.: D. Taylor s.n. (Grape Vine, 5/2/38) (N). Hempstead Co.: E. J. Palmer 5407 (Po—217508, W—1580971). Hot Spring Co.: Demaree 1301 (Gg—-267603, N, Ok), 14878 (Ky, Ok, P1—131513, Ur), 16734 (N), 16920 (Du—256566, Gg~-267602, N, Ok), 16943 (Bt--52485, 1o—-152222, N, Ok), 17270 (N), 18830 (N, Sm), 18880 (N, Sm), 19020 (N, Sm); Letterman s.n. [Malvern] (W--985805). Izard Co.: Demaree 1700, in part (Du--256565, Ok). Lawrence Co. Demaree 25957 (N), 304u9 (Au--121882, Hi--51)37, Mi, N, Ok, We); (i a, «sl Co.: Demaree 168h9 (Du--256563, N, Ok). Perry Co.: Welch 1205 (Dp). Polk Co.: Demaree 36603 (Sm, Z). Pope Co.: G. M. Merrill 202 (Il--19853, St--16757, Ur). Pulaski Co.: R. J. Bailey 5.ne TApril 17, 1937] (Cm); Demaree 18794 (N, Sm); He B. Hasse s.n- [Little Rock, June 1886] (N); G. M. Merrill 1566 (Ur). Scott Co.: Demaree 18161 (N). Sebastian Co.: Bigelow s.n. [Fort Smith] (W), s.n. (9 miles w. of Fort Smith] (W); Demaree 18143 (N), 20881 (N, Sn). Sevier Co.: Brinkley 62 (au, Au); Demaree 36578 (Sm, Sm). Sharp Co.: Demaree 26056 (Le). Washington Co.: L. H. Bailey 7565 (Ba); Fassett 17530 (Pl—76609); F. L. Harvey 18 (Cm); Be Je Turner s.n. [Greenland Township] (Au); E. M. Wells 3 (W—981703) . Yell Cos: Demaree 20907 (N, Sm), 22777 (Sm)3 S. Watson s.n. [Mar. 28, 1931] (St—92h2). County undetermined: F. L. Harvey 30 (C), S.n. (Ka); L. McLean s.n. {Blue Springs, 8-20-31] (St—-14177) ; L. F, Ward s.n. [Arkansas, 1877] (W-155629), sen. [Arkansas] (W— 1599h6). LOUISIANA: Acadia Par.: EB. Brainerd s.n. (Crowley, March 19, 1908] (Vt); Chambliss s.n. (Crowley, Apr. 5, 1916) (TL); Fe W. Pennell 10189 (N). Avoyelles Par.: Hale s.n. [Red River] (W— 314317). Cameron Par.: Cocks sen. [July 1903] (Tl); S. M. Tracy 8707 (Bm, Es, N, Tl, Tr, Up-—503u4, W--510599). Bast Baton Rouge Par,: Correll, Correll, & Brown 10393 (H—7393h) . Lincoln Par. Bardin s.n, (Ruston, April 2, 1951) (Tl). Natchitoches Par.: Cocks son. [Mar. 15, 1911] (Tl), sn. (Apr. 15, 1911) (Tl), sen. emma 4,50 PHYTOLOGIA Vol. 8, no. 8 [Apr. 1914] (Tl); L. E. Fox 401 (Ne), 403 (Ne), LO4 (Fx), 09 (Fx); E. J. Palmer 7001 (Gg--1822h0, W--1531581), 7071 (Ge—— 18219). Orleans Par.: ot Be a 436 (Au--121796)3 J J. Torrey 8. n. (Br). Ouachita Par.: al 840) (Hi—178)0) Plaquemines Pare? Langlois sen. [Febr. $8807 ST (up -7077) Rapides Par.: Charette 57 (Vi) (Vi); “Small & Alexander s.n. [Sandhills near Lena, May 1931] (N). Saint Landry Par : Biltmore Herb. 3655¢ (S). Saint Martin Par.: Langlois s.n,. (8 March 1998] (1). Saint Mary Par.: Penfound s.n. [Cote Blanch Bay, )/3/32] (Tl); Perkins & Hall 2623 (Po— 256898) ; Rose-Innes & Warnock 701 (Au, N). Sain int Tammany Par.: A- nect 20 (W--103665), W—226141h); Arséne 11093 (W--1031525), 11982 (W--1033039). Tensas Par.: Ewan 1 wan 19048 (T (T1); Penfound s.n. [3-25-37] (Gu--19186, Tl). Terrebonne Pare: Amognesne 290 (It). Union Par.: Cocks s.n. [Apr. 1913] (Tl). Winn Par. Penfound s.n. [March 25, 190] (Tl); N. F. Petersen s.n. neanetela: 19-19) (N). Parish undetermined: Hale s.n. [w. La.] (Pr); Herb. Hapeman sen. (Hp); C. Peck s.n. (Ky); Penfound s.n. [C6te Blanch Bay, L/ 3/32] (71); Short son. [Louisiana](Cm, Ll, M, Pr); A. Wood s.n. [March] (Pa). COLORADO: Boulder Co.: Ramaley 15322 (Ok). County undetermined: G. R. Vasey 526 (Gn, Ms, Pa). NEBRASKA: Franklin Co.: Harper & Harper sen. [July 2h, 1897] (Io--133188). Furnas Co.: Lallman sen. [May 20, 1936] (Bt—-1)6800) « OKLAHOMA: Adair Co.: M. Hopkins 3111 (Ok) 5 C. S. Wallis 2859 (St), 6314 (Au 171251, St). Atoka Co.: Folsom s.n. [h-6-23] (Ok--10279)5 Me Hopkins aie (ok), 285 (N, Ok); Hoskins Nelson, ede Nelson 1080 Gibh (ok & Van Vaikenouregn + 85 stat son. (Durant, Avril 20, 193k] (Bt—2969h) 5 Ce Se arate os (Al, Ob--97281, Se—1h962, W--56206). Caddo Co.: Edw. Palmer 2h6 (T). Cherokee Co.: M. Hopkins 390) (0k); Little es Little 19 (Ok), 37 (Ok); W. C. Rhoades 49 (Ok); C. S- Wallis 6L57 (St), 666) (St). Choctaw Co.: H. R. Griffith 21 1 (St--20095)5 W. Be Hughes 25 (Au, Ok); J. R. McFarland 53 (0k); “Van Valken- gh 56 (0k). Cleveland Cos: Fea. Barkley s.n. [5/1/28] (Ok 10272); Emig 15 (W—6),3862); F. As Herzog son. (Helsel, April 28, 190k] (Wi); Hibbard 87 (0k--10283); Jeffs 9 (Ok--10283), Ss n. [5/6/1921] (Ok—10310); Jeffs & Johnston s sen. [5-1-28] (Ok-— 10275); Kitchens 27 (0k—-20505); Mauk sn. (Norman, 5/1L/1] (Ok--20506); Que Quesenbery 18 (We). Coal Co.: Demaree 11976 (Ok— 22229); S. Fry 133 (St—1009); Van Valkenburgh 16 (Ok, St— 2128). Comanche Go.: M. kK. Clemens 11750 (Gg—37139); Daw Davy 186 (St--20238); Demaree 11957 (Au, Ca--8)1)0, ca—71779h, Ny XN, Ok— 22155, St--12971, W--1683779), 13168 (ok--221L8, St—--12973) 5 Ferguson, Buttram, & Buttram 67 (Ok); M. Gates s.n. [Apri il, 1950] (Ok); Hopki Fy Hopkins, Nelson, & Nelson & Nelson 965 (Ok); D. McLean “ott 1962 Moldenke, Monograph of Verbena 451 (We); F. B. McMurry 657 (0k); W. S. Myers 337 (Ok); E. Le Rice s. n, [April 23, 1938) (Ok); Me 0. Rice 52 (Ok) Van Vleet s.n. [ht. Sheridan, 7/4/03] (Du--17264L, Ok--10269, W—12l,2502); Waterfall 291 (Ca-—-882622, Ok). Craig Co.: Carleton 7 (Ka, Ur, Ur). Creek Co.: Bailey & Bailey 10220 (Ba); B. F. Bush 437 (C, Ok, W— 217401); Schendel 33 (St--17917); Shinners, Cottam, & Stephens 3577 (Ca—882617). Delaware Co.: Blevins 6), (St—18580); R. Good- ner lj), (Ok); Reno 26 (Ob--68605); O. S. Wellis 2912 (st), 3769 (St). Garvin Coe! dw. Pelmer 247 (1); Ge We Stevens 109 (Du-- 65695, N, Ok, Ok, St--9245, Ur), Ae115 (St=--9245). Greer Oo.: Hetchett 20 (St). Haskell Cos: Clark & DeVitt 26 (Ok); Zachritz 15 (St). Hughes Co«! He Goodman 81 (Ok). Johnston Oo«? Me Hopkins 4864 (Ok); Hopkins, Nelson, & Nelson 1024 (Ok). Kay Coe! Davy 79 (St--17575)3 Me Be George 4 (Ok); Randel 15 (Ok). Kingfisher Go«: Le Ae Blenkinship sen. [Apr. 18, 1896] (Ob=-50788 ). Letimer Coe? R. Bebb 8 (Ok); O. Me Clark sen. [6/ 12/ 30] (Ok-=10301 ); De He Dunn 19 (St--20073); M. Hopkins 1864 (Ok). LeFlore Coe? Blekley sen. [G. We Stevens 1471] (Ur)s Demaree 12104 (N, Ok--22146, St-- 12972); Ge Je Goodman 2525 (Ok); Hopkins & Grogs 1528 (Ok); Re Fe Meson 119 (Ok), 128 (Ok, Ok, St--14979), 158 (Ok, Ok), Sens [A- pril 12, 1937] (Ok), sen. [April 17, 1957] (St--12970 )3 Pe Me Nelson 46 (Ok). Lincoln Ooe? Le Se Rose 1/0 (St--19869 ). Mayes Coe: Re Bebb 2726 (Ok); Oe Me Clark 3452 (B); Spesirs 78 (Ok, St- 17564); Tripp 166 (St--9257), 167 (St--9258); Ge Se Wallis 6678 (Au--171160); Waterfall 15292 (Go, Hi--193032, W--2331771); Wherry sen. (Pryon, llay 2, 1925] (Up). McClain Oo»: Ven Velkenburgh 1 (Ok). iicQurtein Cos: Demaree 12055 (Ok--21772); Goodmen & Berkley 2061 (Ok--17714); Me Hopkins 2865 (Ok); Hopkine & Cross 2503 (Ok); Hopkins & Pollock 6291 (Ok), 6302 (Ok); Houghton sene [G. W. Ste~ vens 56435] (N, Ur); Little & Olmsted 271 (Ok--10305), sen. (June 16, 1950] (Ok--10504); Cs Ge Ward 296 (St--16870). McIntosh (0+? 8. Breinerd gen. [aufele, April 4, 1908] (Vt); Stretton 5060 (M1, N, St--26149, St--26150). Murrey Cost Flowers Sele (May 3, 1939] (Ok). ihuskogee Cost Re Bebb 3690 (Ok), 3816 (Ok), 3868 (Ok, Ok); Be Le Little 56. (Ok--10303 ), 1577 (Ok--10508 ), 1578 (Ok--10299), 1579 (Ok--10298 ), 1869 (Ok--10300), 2614 (Ok--10509); Se Je Palm er 14285 (B); We Park 103 (Um--26); Le 3+ Rose Sens (May 6, 1941] (St--19871); Turley 85 (Ok); Ge Se Wallis 3791 (st), 6221 (Au-~ 171257, St), 6462 (St). Okfuskee Coo? O. Me Clark 2387 (Ok-- 10302); Rey 14 (Ok, St--18078); Me Whiteheed 26 (Or--41265). Ok- lehoma Oo.: Harding 81 (St--17577); MeLennen 88 (St--24887 ); Ste~ men 89 (Ok); Weterfell 2676 (Ok). Okmulgee Oo.+ De Williens 16 (Ok). Osage Cow? UO. He Calvert 64 (St--25435)3 Te Johnson 115 (Ok), 118 (Ok); Liebenheim 49 (St); Perrin 100 (St—-9234); Rice 452 Pon OL O'G-t a Vole 8, noe 8 137 (Ok); Sizemore 195 (St); Ge We. Stevens 2048 (N, Ok), A.2048 (St=-9259); Stout 18 (St——-24202); Stratton 5095 (N, St--26151, St--26152 ). Ottawa Coe! Brenging 28 (Ok); Inkenberry 27 (St-- 18145); Ge We Stevens 2540 (Du~-66385, N, Ok, Ok--21022, W-- 589900), A.2340 (St--9235); Ce Se Wallis 3731 (St); M. C. Whaley 85 (Ok-~10307). Pawnee Coe: Prewett 136 (Au). Payne Coe: Bogue 1364 (No—8609); Ce O. Chambers sen. [June 19, 1915] (St--9228); Fe Clark sense (Id); Cowley 55 (Ok); Deonier 107 (Bt--48281); Div- ine 104 (Gg--267604); Felts 60 (St--9210); We Be Hughes 30 (Ca-- 882621 ); Ce De Leern sen. [Mey 1, 1915] (St--9226); Preston 93 (B1--42299); Rees 141 (Ok); O. C. Schultz sen. [June 13, 1927] (St--92352); Stretton 236 (St--25124); Waterfall 104 (St); Weugh 168 (Ka, W--266913); A. D. West sen. [April 18, 1959] (St). Pitts- burg Ooe! Ae He Brown 25 (Ok), 60 (Ok); Me J. Hall 26 (Ok); Hop- kins & Cross 1638 (Ok); Fe W. Pennell 10599 (N); BS. Rogers Sens [4~21-57] (St). Pontotoc Ooo: De McCoy 527 (St), 1288 (St), 2265 (St), 2282 (st), 2449 (st), 2481 (St); G T. Robbins 2335 (Ca— 903931); Ven Valkenburgh 2 (Ok). Pottawetomie Cow: 5. De Barkley 27 (Ok--16762); Demaree 11969 (Ok--22231); Feulkner 28 (N), 30 (Ok); Me Te Jacobs 46 (Ok); Pe Je White gene [Auge 8, 1900] (Dm). Pushmateha Cos? Mehler 378 (W-~2331708); Parkey 22 (Ok); Van Val~ kenburgh 51 (0k); Waterfall 633 (Ok), 10550 (St), sen. [Albion, June 5, 1937] (N). Rogers Co.: iiaterfell 1456 (Ok)-» Seminole oe! Demeree 11960 (Ok-~20365). Sequoyah Qos: C. Se Wallig 3680 (St), 6225 (Au--171258, St). Tulsa Oost Pe V. Beck 156 (Ok), 187 (Ok)s Burwell 42 (St--9236); C. Cameron 65 (Ok—10276); Constance aeNe (March 25, 1958] (lim--32156). Wagoner Gost Re Bebb 3935 (Ok), 3955 (Ok). Washington Cos! Rees 246 (Or~-40101). County undeterm= ined: Beiley & Bailey 10215 [Tala] (Ba); Re Bebb 2160 [Florel] (Ok); Butler 10629 [Indian Terr.] (In); Le Gould 59 (Ok—10274); Je Ae Howard sen. [April 21, 1935] (Ok—~18242); We Se Myers 230 (Wichite Mte.J (Ok); Fs He Rose 7 [Wichita Forest; Herb. Us Se Foreat Serv. 42887] (Ca~-445582); Mrs.e 0. St. John sen [Indian Be Ma spss Be SeNe Bryde sene [5/15/31] (Nt). Cameron Cost Ms Cannon sen. [February 1962 Moldenke, Monograph of Verbena 453 9, 1927] (Mi, Mi); Gory 17932 (Tr), 17935 (Tr), 17934 (Tr), 17936 (Tr), 28307 (N); Re Runyon 2497 (N, N, N). Chambers Co.: Tharp 56006 (Au), sens (4/7--10/56] (Au, Au, Mi)e Comal Cos: Thurow se ne (Spring Brench, April 29th, 1923] (Hu). Culberson oe: Hitch= cock & Stanford 6782 (i!). Dallas Coe! Reverchon 2533 (N). Denton Coe: Ae Nelson 10819 (Du--230823, 5, Um—~1l, Up). Falls Coe: Lun- dell & Lundell 10101 (Id, N, W--1888891); whitehouse 19453 (Mi, N). Freestone Co.: Lundell & Lundell 10392 (Ld, N, W—- 1087659), Grayson bo H. Gentry ios Tt Do oie g Co.: E. J. Palmer nek (Gg—182118, W--1531810); C. L. York es [3-20-38] (Au, Au, Gg-- 316097, N, Sm). Hardin Co.: Whitehouse 8748 (Au). Harris Co.: Biltmore Herb. 3655e bid G. L. Fisher 33 (W--503107), LO (W-~ 1203310), bl (W—120 “U7 (W--1287656), 52 (Vi), 1707 (W—~ 719766) , puch (ribess2hy, 1905 (Go), sn. [Houston, March 31, 1912] (Ew), sn. [Houston, Mar. 2, 1913] (Vi), s.n. {Houston Apr. 3, ee Tish68, Hp), sen. [Houston, Apr. 3, 1915] (bo— 172646, Mi), sen. [Houston, Mar. ar. 31, 1918] (Ca--882623), s.n. (Houston, Apr. 3, 1920] (Ur), s.n. [Houston, Apr. 22, — 3) (Hp), son. (Spring, May 4, 192)] (Du--172647, Ew, H—-5L73, Ur), S.n- TMar. 17, 1930} (1), 82 s.n, [Houston, March 2s, 1938) ” (Gg-—310875) 3 Gaston 1l (Au); E. H all 35, in part (N, Pa, Po—71257, Pr, W); Joor ou (W—771876), 8.) Sen. (Harrisburg, August 10, 1875] (Mi); Lindheimer 137 (W--1870841), s.n. [near Houston, * March 1842 (Sm); E. Meyer s.n. n. [Buffalo Bayou, Feb. 18h) (C). BOOK REVIEWS Alma L. Moldenke oe oes of the World for Tropics and Warm Climates", by enninger; 335 pp., 425 color plates. Hearthside Press, - ey 1962. $18.95 The author would have been most pleased with this review if it could have been rendered as a "play~back" of a tape recording of t e "ahs", "ohs", and more achacndeke and laudatory comments of the botanically minded people, lovers of nature and appreciators of the beauty in color photograph and art who have given an al~ cently presented. Mr. Menninger states that "the purpose of this book is to put the finger on tne flowering trees that somebody thinks are the 454 PHT70L0G0 12-4 Vol. 8, no. 8 most beautiful in the flowering world" of tropical and warm cli- mates. That "finger" is guided by a highly skilled brain, a great heart, and a most perspective eye for beauty, and is accompanie selected one thousand species in 500 genera to describe, ofte the words of the plant explorers themselves, and 25 to *iinateeee by means of beautiful color photographs. So many popular books use only confusing, vig empte and over- lapping common names, known often only locally. So many scientif- but not aga a Na attractive, illustrations. This book serves both gr unusually well because the flow cba ais trees are grouped a their families alphabeti cally, e given common well as scientific names, with iracttes and synonyms. The 5 also includes a thorough seeen? a helpful bibliography and a highly and variedly useful series se classification tables based on flower and leaf c coloring, minimum survival temperatures, growth rates a sizes, anh a to wind, cunie spray and fire, odors in flowers and eav es, nomic us 3, shade and moisture relationships, a a rie et and geographic ranges, etc. rb even a chapter on "left out" flowering trees to help pacify readers who find a few of their favorites omitted. It is gratifying to recall that from the author's experiment- n means of beautifying_so much of that Sunshine State. The dedica- tion of this book, like the dedication of the author to his ow the number of its color plates are conside ned eas of the plates are exquisite; just a few have lost a little clarity in reproduction. "Michigan Wildflowers", by Helen V. Smith, ee by Ruth P Brede; 65 pp., 231 line-drawing plates, 16 color plates. Uni- versity Publishers, New York atv $5 .00 This is a pleasing and iS ae book because e its natural appearing drawings and color s, clear text and easy-to~ follow keys and descriptions. 7% lists each plant ty re or more common names, by scientific name without synonymy, 4 fay: statement about the number of genera and species in it graphic distribution, and a special comment about a few members well known for their attractiveness or economic importance. Many fascinating bits of plant lore are given under some of the spe~ 1962 Moldenke, Book reviews 455 cific descriptions. e is a fine rine for the conservation of certain vanish- ing wildflowers, and some good suggestions for wildflower gard- e diagrams of flower and leaf parts and types all aid in making the book useful to the erpieer’ as well as to the local botanist. The bibliography * Rating ies, 1 natural fran and 3 named forms and varieties in the state, one occurring in at least le counties. Phyla lanceolata, wn. "tr om This book's extremely modest price in terms of today's book rears Fund to the sal onyesi 3 Institute of Science. This is a rthy service because it a good book within easy reach of ae more amateurs than ae otherwise be able to afford it. "Photochemistry ee es sgn vente quarterly j al, Vol. O. -M sr . Pergamon Press, 3 A. edito. ew York and Heri 1962. UL bec seers ioah $10, in- stititional $20. This new journal was spawned by the recent Photobiology Con- gress held in Copenhagen to provide a spec ecific organ for papers in Rico —S expanding field and "to bridge the gap between osing views of nature, name ely, the physical an and room biological systems", stressing the quantum aspects of the sub- ject. Rapid beget ts is an expressed goal; this first issue has achieved it The first eight articles emanate from four universities S the United States, from Australia, from Warsaw, from Ein ndhoven from Frankfurt. Two of them are in German; and tucleic acids" cent lit terature in photobiology organized alphabetically by ea sent sage is listed on a few pages. There is a photobiology news page, The printing is clean. The instructions to contribu- tors are nee organize This journal will probably prove to be much in demand in all related fields. BOOK REVIEW Gerald S. Hanusik* "A Dictionary of Terms in Pharmacognosy", by George M. king ; 28 pp. Charles C. Thomas, Publisher, Springfield, nth 1955. The author has here compiled a dictionary which h will undoubted- be of value to students in any one of the health professions. It seems to be designed to augment the information given in text- books, and to provide the student with a wi Sie and quick reference W r ’ than it is to subjects more easily accessible in the many refer- ences available. This compilation of words an i a uinely revealing work which gives a tite coverage to a fie heretofore deprived of any single reference which would be ade- quate. A work of this magnitude is without question a monumental task. An author who undertakes to write a scientific dietiotery mist first be a scientist who is qualified in his specialty, and, secondly, he must be a man of diligence and fortitude. The writing of a dictionary is not an easy task. It therefore goes without saying that omissions and mistakes will o bake his di tionary is quite thorough, and this reviewer could find sad a few omissions of information in some of the unease the author covers. One of these omissions is under the waa of A Here he has failed to distinguish between Sweet Almo ond (Prunus amygdalus dulcis), which is a food, and Bitter Almond (Prunus amygdalus amara), which is poisonous due to the presence of gdalin, a a substance which also gives to Bitter Almonds their abe taste. his omission is small when thine ay se o the magnitude of the dictionary ant: umber of cakemaiteh covered. - He must ha g through the pon of the students whom he will indirectly este with this book, “B.S. in Pharmacy, Fordham University College of Pharmacy; at present a pharmacist with Reife's Pharmacy, Yonkers, New York. 456 enke Mapping by counties done bY Andrew R. Mold Figure 7. Distribution of Verbena canadensis in the United States i 10 I oo UPRGEe Ray Cag eee Se rring ue Devel Bows pant Soe WOO0T ap PPE: De0GE ial a lil Crt Sx | ig ROK | HORLES FLLMORE iJ s s MIRMEKAHA ‘FREEBORM 1 E on oy ra nine ee Lia iN | i H : i a | i [DEN a | Fi Aeon Wivat RES ait ef rl | [Sal Gale bite a sg COUMERT FRANKLIN fe {i A . Te ‘ . Lp “En ar rot «] JEFFERSON C LE A RTYP E COUNTY OUTLINE UNITED STATES ; Scale of Miles ry Bere Els cau =o ae oly. < ag | . s . ‘wae TON 100 aoe | & pee 4 MAP NO. 125 ol ee OTA Feo aes £ > : we AMERICAN MAR OMPANY INC Ce | @~ Sy oe ll U8 Lee aa ee ol IGINATOR ORAS SOLE PUBLISHERS J 9 on CLEARTYPE MAPS RSGz/ COLORPRINT MAPS YORK BONE Tes 5 sopieidhted sag. The law 4 ose of resale, sathout De: ase PHYTOLOGIA is financed entirely by its contributors, each onc paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if amy accrue. Each number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary greatly. A volume will contain about 32 signatures, 512 pages, or a smaller number of pages with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra pu is made for line drawings, such as are ordinarily reproduced in zinc, or for diagrams, tables, or charts, provi ded they conform to certain limitations of size and proportion. An extra charge will be made for half- tones, dep on their size, as fixed by the engraver, with a minimum of about $2.25. tales icles dealing with research in all lines of paleonnes in any reasonable eh, bigaphic sketches, and critical rev and eT S liter- att ered for publication. paki Poe casual no! an amateur or so-called popular ‘type, and polemics will not be a oink Advice on the suitability of manuscripts will be solicited, if necessary, from quali- fied botanists, ‘Under the present cost of co the 0 rate for a page or y $3. 00 fo of 250 copies. This price is spree? to change without notice, since it eas gee on the Aogh — in n the printing ' i ; / Reprines will be furnished. at cost. A proportionate, of 250 btu is also geass gratis to CREED OLOERE ne bpm request the editors will send detailed instructions conenmnind the ipt or further information about the maga PHYTOLOGIA Designed to expedite botanical publication Vol. 8 January, 1963 No. 9 CONTENTS KOYAMA, T., Nomenclatural remarks on the genus Scirpus. 1.... 457 MOLDENKE, H. N., Notes on new and noteworthy plants, XXXII. 459 MOLDENKE, H. N., Materials toward a monograph of the genus Verbena, Vill o.oo ree oo ve a ee eS he es SMITH, L. B., Notes on Bromeliaceae, XIX ....++++-+++05 ee ey 497 SMITH, L. B., Synopsis of American Velloziaceae Supplement Il . 507 3 Author index to Volume Eight ss cccevccceess: Rr eee 2 os ee 3 by Index to supra-specific scientific names in Volume Fight ......- 515 Publication dates of Volume Eight .....++++++++0+000 oe 520 Published by Harold N. Moldenke and Alma ie Moldenke 15 Glenbrook hea =e = gee : Yonkers 5, New York, U. S. A - a Price of. this number, $1; per clone $5. 21k pe Volume 8, No. ae was s issued pos ove 10, wns fee ala, eee ed ae oe ee NOMENCLATURAL REMARKS ON THE GENUS SCIRPUS I Tetsuo Koyama The following notes are based on some syno- nyms that were overlooked, and the examination of a number of critical specimens and old liter- ature referencesthat were unavailable to me when I was preparing a treatment of some of the aphyl- loid species of Scirpus that occur in North America, lf -Scitpous juncoides Roexb.;, Hort. Beng: St, ing, ed. 1814, nomen nudum, & Fl £4 2, Lee. var. DIGYNUS (Béckeler) T. Koyama, comb. nov. nus Bockeler Scirpus supinus Linn. f digy. in Linnaea 36: 700. .1869-70 Bockeler's var. digynus was a new name based on S. debilis Pursh, non Lamarck, and has priority over var. williamsii. The synon- ymy and status of var. williamsii have been fully discussed in a recent paper (T, Koyama in Bot, 49:.9145.9190 270e1 Can, Journ, var, juncoides. 2. Scirpus juncoides Roxb, Scirpus supinus Linn.velatior Béckeler, in Linnaea 36: 700, 1869-70. In major part, excl. syn. 2» wallichii. Bockeler's var, elatior was based on $. juncoides, and could be included in the synonymy of S. juncoides var. juncoides,. 3, Scirpus lacustris Linn., io Pig ed, 1. 48.. 1753. eubsp, CREBER (Fernald) T. Koyama, comb, nov. Scirpus validus Vahl, 1806, Scirpus validus Vahl, var. creber Fernald in Rhodora 45: 282. eee | = . Schoenoplectus validus (Vahl) A. Love & ; Lowe in: Bali. ents. Bot, Giub, St; 35. 1954, Enum. Pl. 2: 268. — 457 458 PHYTOLOGIA Vol. 8, no. 9 Schoenoplectus validus A. Love & DD. heave subsp. creber (Fernald) A. Léve & D. Thee 4. co 8h oo, 1994 This new combination replaces Be lacustris ssp. validus and has priority as creber was first treated as a subspeci es by Léve and Love, a Scirpus lacustris Linn, subsp, Koyam var, LUA creber T, a URIANS (Miquel) T. Koyama in Can, thutn> Bot. 40: Jey. 1962, valido, em, hoc. loco. Scirpus tabernaemontani C.C. forma luxurians sub ssp- Gmelin Miquel, Ann, Mus. Bat. Ligd.-Batav. 2: 143, 1865, Scirpus hippolyti V.- Krecz. in Not. ex Herb, Inst. Bot. Acad. , Ti 28 OSS. (Bynes NOV. Schoenoplectus hippolyti Vj Ketece. eadem loco. onymis. The holotype of S. hippolyti, Central Asia: Kagaktania, FN. Rusanov 2491 (LE), has been It was, as suggested in my previous paper (T. Koyama, 1962.), identical with var- luxurians of southeastern Asia. Syst. Sci. URSS. Nomen invalidum ex syn- examined, NOTES ON NEW AND NOTEWORTHY PLANTS, XXXIII Harold N, Moldenke CITHAREXYLUM POEPPIGII f. ANOMALUM Moldenke, f, no Haec forma a forma typica speciei alabastris Ahi heisetcrea tis, ad apicem conicis vel attematis, et calicibus elongatis cylindricis 6--9 mm. longis rece This form differs from the typical form of the species in having its unopened flower—buds elliptic-elongate, gray-puberulent, conspicuously Mg or ee ee at the apex, and the calyx elongate-cylindric, 6--9 mm long. The type of this puz zling form was collected by A. L. Bernardi (no. 2142) in wet tropical woods at Camp No. 3, 33 km. from El the process of being built, Bolfvar, Venezuela, April 9, 1955, and is deposited in the Herbario Nacional de Venezuela at Carac or pi CITHAREXYLUM POEPPIGIT var. CALVESCENS Moldenke, var. nov Haec varietas a forma typica speciei foliis subtus minute puberulis calvescentibus re cedit. This variety differs from the typical form of the species in having the lower surface of its leaf-blades only very minutely ae ‘haga or glabrescent. type of the variety was collected my good friend and redid José Cuatrecasas pee (no. 15949) in woods at Veneral, at an altitude of 5 to 50 meters, Rfo Yurumanguf, on the Pacific coast, Valle del Cauca, Colombia, between January 28 and February 10, 1944, and is deposited in the Britton Herbarium at the New York Botanical Garden CLERODENDRUM SERRATUM var. VELUTINUM Moldenke, var. Haec varietas a forma typica speciei laminis era pra —— — inflorescentiisque dense velutinis et foliis parvis phe variety ag pb from the typical form of the species in having only small leaves at time of anthesis, 4--7 cm. long, 2--3 cm cence, bracts, and calyxes also densely veluti The type of the variety was collected by F. Saadentare (no’, <733), growing in an old recently abandoned "taungya" on open hillsides covered with grass and tangled thickets, at Mindat, at about 5000 feet altitude, in west central Burma, on April 26, 1956, and is deposited in the herbarium of the British Museum ature) History) in London. The collector describes the plant 88 a soft-wooded shrub with sre stems. 9 MATERIALS TOWARD A MONOGRAPH OF THE GENUS VERBENA. VIII Harold N. Moldenke ENA “ts Additional and emended bibliography: Hook., Bot. Misc. 173, pl. Ven k8. 1829; Maund, Bot. Gard. 9: ple 197. ee Lem., Horticult. Univ. 3: 9. "1842; Se Wats., Epos « Am. sows Sci. 18: 136--136. ie Briq., Arkiv Bot. Stockh. 2 (10): 7--lh, pl. 3. 190k; R » Exch. Club Brit. Isles ats tee. oa re Florula 1059108. "1930; parodi, Rev. Argent. Agron. 1: 202. 193k; Beale, Chron, Bot . ot £17. 1939; Herter, Revist. Sudam. Bot. 7: 225, 1943; Barton “¢ Garmon, Contrib. Boyce Themen PSs Wy: 243, 26, & 252. 1946; Angely, Fl. Paran. ce h & 12--13. 19573 To He Everett, New Illustr. Encycl. Gard. Zuo 2106, 1960; vennin- ger, Flow. Trees World 18, 281, 286, 308, & 335. 1962. Additional abbreviations used for the names of herbaria in this series besides those listed or referred to in Phytologia 8: 95 (1961) are Am = Southwest Research oe American Museum of Natural His- tory, Portal, Arizo = University of North carolina, Chapel Hill, North Carolina = P, Olrog Herbarium Pm = Pharma Herbarium, CIBA, ggetacted ay gr ER Rf = Texas Research Foundatio Texas Ws = University of Wisconsin, gooey "Wisconsin Oo ol b VERBENA ABRAMSI Moldenke Balls found this plant growing in moist heavy soil along with Mimulus cardinalis Dougl. Additional citations: CALIFORNIA: Kern Co-: Groesbeck s-M- [Glenville, 6.6.1954] (Bm). Los Angeles Co.: Balls 10065 (Bm) « VERBENA ALATA Additional ilsigventirs Angely, Fl. Paran. 7: 12. 1957- Material of this species ae been misidentified iol distribu- ted in herbaria as V. littoralis H.B.K. Additional citations: BRAZIL: Minas Gerais: Regnell 1.3¢0 326, in part [19-11-186] (W--119969). Santa Catarina: Reitz & “Klein 548) (W--2269015); Smith & Klein 8065 (W—--2251339) 5 9); smith & h & Reitz 9059 (W--2251h87) . VERBENA AMBROSIFOLIA Rydb The E. steams 10h, annotated as V. ambrosifolia by P does not seem to belong here. I am cit iting it under V. siti seie Small, as also ye learns 591 which she annotated as 3s "Cf. Verbena bena ambrosifolia We Su a2 Additional citations: OKLAHOMA: Cimarron Co.: Demaree 13335 1,60 1963 Moldenke, Monograph of Verbena 461 (Rf). TEXAS: Brewster Co,: C. H. Muller s.n, [Cat-tail Canyon, 7- 6-32] (Ar—199985); B. H. Warnock 10562 (RF), 10882 (Rf), W.283 Rf). Culberson Co,: “Correll & Jo & Johnston 19136 ( Rf), 20370 ( Rf); Correll & Rollins 23785 (Ld). Dallam Co.: D. . D. “ Correll Fire (Rf). Hudspeth Co.: C. H. Muller 8215 (Rf). Pecos Co.: 1 B. H. War- nock 5169 (Rf), 11851 (Rf). Presidio Co.: Correll & Johnston 21852 (Ld (Ld). Reeves Co.: Correll & Johnston 18592 (RE); B. H. War- nock 10185 (rf). Terrell cos: Warnock & McBryde 14750 (Rx) MEXICO: Nuevo Leén: Edw. Palmer 1050, in in part {Monterey ] (Ww (W--56188). Tam- aulipas: Rozynski 312 (i 312 (W=-1588),39) . ee ANDRIEUXII Schau, he type collection of this mange was gathered Foci Be Chila (dit. de Puebla) et Huanapan (dit. de Oaxaca)" according Schauer. The species has beer souk on poms ridges at alti- tudes of 6500 to 8000 feet, flowering in June and November, Perry annotated Edw. ee imer 3) as wAff. V. delticola Small" Additional citations: MEXICO: Oaxaca: E. W. Nelson "1895 (W-- 250226). Oaxaca or Puebla: Andrieux 138 (W—-photo of - of isotype). San Luis Potosf{: Edw. Palmer 3h (W--1,70888) . VERBENA ARISTIGERA S. Moore Additional citations: PARAGUAY: T. Meyer 153 (Bm). ARGENTINA: Chaco: Venturi 9780 (Bm). VERBENA AUSTRALIS Moldenke Pa ci bibliography: Angely, Fl. Paran. 7: 6, 7, & 12. 1957. VERBENA BALANSAE Briq. Additional bibliography: Angely, Fl. Paran. 7: 12. 1957. Additional citations: BRAZIL: Se asd Dusén 16907 (W-- 1481761). is an "herb with pinkish lilac flowers growing in full e Very widely distributed in this locality Galoarateis Rasarvedsl® or with "pale lilac or cream-coloured fine epee ” picetng big sb Miers adtnedtion from Conean is marked "sp. nov ture with V. sulphurea D. Don. Herbarium Site ee bees aoe identified and distributed as V. dissecta Wil Additional citations: CHILE: ": Coquimbo: dieihécdoenct 4hs1 (W-- 1283809), 5053 (W--1)22160). Santiago: Claude-Joseph 2898 (W— 1233408) . Valparafso: J. Anderson s.n. [Valparaiso] (Bm); Bridges tan (Prov. Valparaiso, , 18hh] (Bm); A. Calvert s.n. [Valparaiso] Bm) ; Claude-Joseph 3769 (W—-12816L6) ; Cuming Sn. [Valparaiso] 62 PHYTOLOGIA Vol. 8, no. 9 (Bm); Miers s.n. [Concan] (Bm, Bm); Sandeman 175 (Bm), 197 (Bm). Province undetermined: Bridges 455 (Bm); Claude-Joseph 128 [Cos- ta] (W--105926)), 4456 (W--128379) 3 Collector undesignated 623 (Bm); Miers 602 (Bm); E. W. White 22 (Bm). VERBENA BIPINNATIFIDA Nutt. Additional bibliography: S. Wa US ey, Proc, Am. Acad. Sci. 18: 136. 1883; T. H. Everett, New Illustr. E neycl. Gard. 13: 206. Collectors have found this plant also growing in black clay, sandy black soil, and grayish calcareous soil. Specimens have been misidentified and distributed in herbaria as V. officinalis L. On the other hand, the Hamby 514 and Ibana Garefa 331, dis- tributed as V. bipinnatifida, are Ve. bipinnatifida var. latilo- bata Perry; Purpus 452) (in part) is V. bracteata Lag. & Rodr.; and Hinton 1191) is V. teucriifolia Mart. & Gal. Additional citations: TEXAS: Andrews Co.: Lundell & Lundell 11405 (Rf). Atascosa Co.: C. D. Marsh 11087 (Ar--11813) . Bexar Coe: Reynaldo Ramirez 36 (Ok). Concho Co.: Hamby 94h te Croc Ar— 206756 kett Co.: Warnock & McBryde 15219 (Rf), 15313 (Rf)- Dallas Co.: C. L. Lundell 1170h (Rf); Lundell & Lundell 11315 (Bm), 12132 (Rf, Rf); 0. Sanders sen. [Ledbetter Drive, April 10, 1957] (Rf). Deaf Smith Co.: CG. L. Lundell 11157 (Rf). DeWitt Co: Blanch 50 (Ok); Correll & Schweinfurth 15556 (Rf). Gillespie Co.: Correll & Johnston 19558 (Rf); Hamby 899 (Ar--2067L6) + Hays Co+: Re M. King 2615 (Id). Jefferson Co.: Lundell & Lundell 11232 Rf)« ielullen Co.: Alvarez, Guajardo, Salazar, & MeCart 7710 (Ok); Paez 112 (Ok). Medina Co.: Hamby 595 (Ar-—~ 0670h); Co De Marsh son. (Near Hondo, Apr. , 1908] (Ar--7l91). Oldham Co.: Ce L. Lundell 11462 (Rf); Lundell & Lundell 11450 (Rf). Pecos Co. B. H. Warnock 13356 (Rf). Randall Co.: Lundell & Lundell 11 (Id). Tarrant Co.: Ruth 107 (Rf). Taylor Co.: Lundell & Lundell 1136 (Rf, Rf). Terrell Co.: Warnock & MeBryde 1666 Rf). Travis Co.: Barclay & Perdue 778 (Id). Uvalde Co.: C. L. Lundell 10955 (Rf), 10958 (Rf), 10975 (Rf). Val Verde Co.: Warnock & McBryde 15073 (RE); Warnock & Turner 703 (Rf, Rf). Victoria Cos: F. L. Lewton 60 (Ar—271672). Ward Coe: He E. Wheeler s.n- [near Mona- — ee hans, April 21, 1938] (Ar—22l580). Zavala Co.: B. Stewart 52 (0k). a VERBENA BIPINNATIFIDA var, LATILOBATA Perry Additional citations: MEXICO: Chihuahua: C, H. Muller 3347 (Rf). Durango: Ibana Garcfa 331 (W--1013337) - XVERBENA BLANCHARDI Moldenke The E. W. Lathrop 369, distributed as this hybrid, is actual- ly V. canadensis (L.) Britton. 1963 Moldenke, Monograph of Verbena 463 VERB. BONARIENSIS L. Additional rey emended bibliography: Hook., Bot. Misc. 1: 166. an Angely, Fl. Paran. ae 125195 7g.%. He Everett, New Illustr. cl. Gard. 13: 206 eee (1829) says of ee species "Hab. Vulgaris, praecipue cis cultis. In Bonnaria et Mendoza. I possess this plant or more places, most readily on each side of the shorter tooth, which often thus constitutes a separate leaflet, The Verbena littoralis of Reece seems to be a variety of this with shorter spikes than u The Osten “E301, satitemmidl as V. bonariensis, is actually var. conglomerat Briq.; Hollermayer s.n. [Wwerdermann 1362], me G. Houk - (Herd. Inst. Bot. S. Paulo 20), and Jensen-Haarup [San aa Rosa, 1904] are V. brasiliensis Vell.; Claude-Joseph 1] is V. corymbosa Rufz & P, Pav.; Buchtien 59 and Morong 15,0 (in part) are x xV. dermeni Moldenke; Jorgensen . 268 is V. intermedia Gill. & Hook.; 1 Widgren idgren 898 is Vv. " Lindbergi Moldenke; and 0. Solfs 70 is ve sphaerocarpa Pe Preys... leo Ss Additional citations: BERMUDA ISLANDS: Main: Brown & Britton 98 (W—-52181)); Harshberger s.n. [Pembroke's Swamp, June 21, 1905] (W—8)7616) . JAMAICA: W. Harris 9132 (W—52l,608); Harris & Law- rence C.15120 (W-—79l,026); Nichols 163 (W—-L29097) ; Alexandrina Taylor Lah], (w—8)8565). BRAZIL: Santa Catarina: Reitz 3321 (Herb. Barb. Rodr. 993] (W-~2141662); Smith & Klein 1139) (Ww— 2251776); Smith, Reitz, & Sufridini 9426 (W—2251518), 95L5 (W¥— 225156). CHILE: Valdivia: R. A. Phidippi 1303 (W—1323051). ARGENTINA: Buenos Aires: R. Alvarez 759 (Bm) ; | Boffa s.n. {Beris- so, XII.1944] (Bm); Rufz Huidobro dobro 1666 (Bm), 1695 (Bm), 1771 (Bm 2). Rfo Negro: W. Fischer cher 236 (W—-704)27). Tucum4n: Descole & Schreiter 10752 (Bm); Ventu Venturi i 2612 (W--1591268). SOUTHERN RHOD ES IA: Ne Cc. Chase 1 (Bm, “Bm). Pt PORTUGUESE EAST AFRICA: Lourengo Mar- ques: F. A. Mendonga 2973 (Bm). REPUBLIC OF SOUTH AFRICA: State undetermined: Ecklon Ecklon & & Zeyher 8.n. (W—-1578500). MASCARENE IS- LANDS: Mauritius: Ro: Roxburgh s.n. (Bm ). INDIA: Uttar Pradesh: B. L. ee s.n. (Mussoorie, Sept. 1927] (W--127233)). NEW CALEDONIA! “tye Caledonia: Le Rat 2345 [U. S. Nat. Arb. 218543] (W— 2185073 73). AUSTRALIA: New South Wales: Constable 19183 (Ww— 2126578) ; Goode 82 (Bm VERBENA BO SIS var. ,omuagno, ae ee icy bibliogra Ang 3 > . citat pectin Spy eyed aasias r 893k (Bm). URUGUAY: Osten § 5391 (W--11779 73) » 6), PHYTOLOGIA Vol. 8, no. 9 VERBENA BRACTEATA Lag. & Rodr. Additional and ehh bibliography: » Proc. Am. Acad. Sci. 18: 136. Hate H. Ve Sm., Mich. a 313, 315, 317, & 65, is ape « Additional hee oe H. V. Sm., Mich. Wildfls. pl. 155. 1961. The Canby, Sargent, & Trevease 1gh, distributed as this spe- cies, is V. canescens var, roemeriana (Scheele) Perry; MacDougal 73 is V. gooddingii var La areTOLin Tidestr.; Edw. Palmer and W. Schumann 1070 are V. gracilis Desf.; and A. W. Chapman 69 is xV. perriana Moldenke. Additional citations: ALABAMA: County undetermined: Buckley S- n. [Alabama] (Bm); Rugel sen. [ad vias et in incultis arenosis per Alabamam mediam] (W—21881)1). ILLINOIS: Tazewell Co.: V. He Chase 13539 (Rf). County undetermined: Meade s.n. (Bm). KAN KANSAS: _ Kiowa Co.: Horr & McGregor 3790 (Rf). UTAH: Rich Co.: M. Ee Jones sn, [Alta, Aug. 2, 1879] (Bm). COLORADO: Boulder Co.: "Ewan Sn. {Plant. Exsicc. Gray. 1090] (Bm). OKLAHOMA: Murray Co.: Ge | elidel Merrill 28 (Rf), 747 (Rf). TEXAS: Andrews a Seudday s [September 1957] (Rf, Rf). Culberson Co.: B. H. Warnock ae (Rf), 14313 (Rf). Dawson Co.: Lundell & Lundell ee Rf), 16973 (Rf)- Hudspeth Co,: B. H. Warnock 93h6 (Rf). Presidio Co.: Warnock & McBryde 1)5)2 (Rf). Tarrant Co.: Cory 54522 (Rf). Wheeler Cost Correll & Johnston 17044 (Rf). Wood Co.: ati: Cs L. Lundell 12081 (Rf). NEW MEXICO: MEXICO: Dona Ana Co.: Wooton SMe [Mesa west of of the aes an Mountains, Aug, 26, 1899] (Bm). WASHINGTON: ede Coe: - K. Beattie 11728 (Rf ). MEXICO: Coahuila: Edw, Palm r 1048 (W— TSe5e8y; Par Purpus 452, in part (W--81695). Sneea Germany Bilimek san. [6/8760] (Bm VERBENA BRASILIENSIS Vell. rarer bibliography: Angely, Fl. Paran. 7: 12. 1957. « Ge Houk specimen cited tae was ig iaies identified as ne eintpaoahata jamaicense Vahl" and later as Verbena bonara ensis L. Additional citations: TEXAS: Hardin Co.: C. L. Lundell 14083 (Rf). Jefferson Co.: C. L. Lundell 11136 (Rf). COLOMBIA: Santan- der: Killip & Smith 1730 (W—-1352959) « | BRAZIL: SHo Paulo: We Ge Houk s.n. (Herb. Inst. Bot. S. Paulo 2),0] (W--1592309). Santa Catarina Island: Reitz & Klein 292 [Herb, Barb. Rodr. 6318) (W-— 2321087). BOLIVIA: £1 Beni: Buchtien 1688 (W--1159360). La Pazt M. Bang 136 (W—~72000). PARAGUAY: Morong 128 (W--2l3599) - CHILE: Aconcagua: R. C. McLean Q,6 (Bm). Valdivia: Hollermayer s.n. (Werdermann 1362] (Bm, W--1516867). Valparafso: Cuming 352 (Bm). Province undetermined: E. C. Reed s.n. [Chil i] (Bm). Bm). ARGENTINA: Buenos Aires: H. N. Curran s Sone [oct. 19, 1913] (W—920853) - 1963 Moldenke, Monograph of Verbena 65 Mendoza: Jensen-Haarup s.n. [Santa Rosa, 1904] (W--1081083); Sem- per 342 (Bm). VERBENA CALLEANTEA, 9 riq. Additiona tat tons: ARGENTINA: Tucum4n: Venturi 2922 (Ww 1591272), 3568. amt ees ee ee VERBENA CAMERONENSIS L. I. Davis The specimen of Edw, Palmer 90 in the British Museum herbarium bears a notation "conspecific with this is a specimen at Kew col- lected by Palmer in 190. I, is unnamed. L. M. 4/1920." Santos (Rf); Traverse 1107 (Rf). Hidalgo Co.: M. Santos 2 (0k). MEXICO: San Luis Potosf{: C. C. L. Lundell 12256 (Rf). Tamaulipas: Edw, Palm- sr 20 (Bm). BENA CAMPESTRIS Moldenke earns citations: BRAZIL: Santa Catarina: Reitz 339 (W-- 2104858 nt CANADENSIS (L.) Britton @ E. W. Nelson 6628 [6028] and Pringle 11843, distributed as xed species, are actually V. delticola Sma. Small, while Lemmon Lemmon s.n. Paul Springs, June 108) is Ve gooddingii Briq. The Lathrop col: collection cited below was originally distributed as xVerbena blanchardi Moldenke ditional citations: KANSAS : Woodson Co.: * W. Lathrop 369 IT nd eSUESTAN Tensas Par.: Ewan 19048 (Bm). TEXA TEA ee dell & Lundell 11 On ey aa Harris Co.: Penfound $.n. rest ene ton, April 10, 1936] (Tl, Tl); Ravenel s.n. (W); Snyder s.n. (cypress, 1906] (Ca—185370); Tharp & Barkley 171030 030 (Ca—-768197, Ur). Henderson Co.: Cory 53953 (Rf); “Lundell & Lundell 81,89 (Ld, ii, N); O. Sanders 151 (Mi, Sm); Tharp s.n, ) Sen, [June 6, 1932] (Au, ax) Tindel 5 (Nt) (Nt). Hidalgo Co.: Clover 566 (Mi); Re Runyon 2625 » N). Houston Co.: A. Freeman s.n. [2.28 28.1930] (Au); E. Je - Pal- mer 12056 peg Ca——25591); Tharp 838 (N, W—-1104761). Hunt Co? Stran tman s.n. [Reeves 201] A} (Ge)? vasper Goer So 52728 (N, ieee 52864 (N, No—15l13); Lundell & Lundell 1118) (Sm), 11203 (N, Rf); Tharp sen. [4/13/41] (Ca—882619). Jef- ferson Co.: Bray Bray 65 (Au). Kendall Co.: H. B. Parks sen. [Cory 29520] (N). Kleberg Co.: G. L. Fisher 1065 (W--18255hl). Lamar Co.: Brozleton s.n. (Paris, 1933] (Au); G. Jones 2 (Ok); McMullen S.n. [Paris, 3/13/27] (au), sen. [Paris, 3/15/27] (Au). Lee Co.: Lundell & Lundell 1009) (Ld, N)- Leon Co.: Correll, Johnston, Edwin 22)03 (Ld); Lundell & Lundell 10382 (Ld, Ld, N). Liberty Co.: 1,66 PBT O:1i0sGt& Vol. 8, no. 9 Ee Je Palmer 11979, in part (Au--1218)6, Ca--h25593, Po--22020h). Wadison Co.: Huss 91 (Ca--37893). Marion Co.: R. Bebb 2645 (Ok, Ur); Correll & Iundell 18791 (Rf); B. B. Harris Lh6 (W—1L66779), 456 (W=-1)6676), s.n. [1-20-30] (Nt). Milam Co,: Lundell & Lun- dell 10096 (Ld, N); Wolff 026 (Tr). Montgomery Co.: D. S- Cor- — a rell 14168 (Sm); E. J. Palmer 33319 (N); Tharp & Barkley 17T030 (Au); Warner son. [Willis] (Hu). Nacogdoches Co.: Lundell & Lun- ’ dell 9721 (Ld, NN), 9751 (N). Newton Co.: C. C. Albers 39009 (Au); Cory 52606 (N, W--1925316). Orange Co.: Bray 65 (W—358206). Polk Co.: Girvin 2000 (Au), sen. [March 15, 1910] (Au); Lundell & Lum dell 9723 (14, N), 9724 (Id, N), 11291 (N, Rf); Ee Je Palmer 5105 (Po--217712, W--1581012); Rose-Innes & Warnock 765 (Au, N)3 T sen. [h-25-36] (Ca--882620), s.n. (3/15/1] (Au, Au, Ca--882618, Gg—31506h). Red River Co.: E. Adams 69 (Nt); Sartwell s.n- [Red River, red] (Ru--2159)). Robertson Co.: Hardin Su7 (Hi—111002, Mi, W—2327778); Lundell & Lundell 10098 (N), 10099 (Ld, N), 10380 (N); Reeves 184 (N); Rowell 8001 (N). Rusk Co.: Henderson School 7102 (W—1h65393); Herb. Univ. Texas 7102 (Au). Shelby Cos: De S. Correll 16167 (Rf). Smith Co.: Cory 54999 (W--2007302). Trinity Co.: Parks & Cory 7963 (Tr). Tyler Co.: Lundell & Lundell 9731 (Ld, N), 10945 (N, Rf); Whitehouse 87147 (Au, N). Upshur Co. D. S. Correll 13161 (Sm). Van Zandt Cost Je Be Henderson 5697 (W--1168813); Juanita Johnson sn. [Wills Point, 4.17.1930] (Au); Lundell & Lundell 9698 (Ld, N, W--1887576), 10437 (Id, N); Peck- ham s.n, [May 1930] (Au); Raborn & Reynolds h2 (Au), 8746 (Au). Wood Go.: McMullen s.n. (Golden, Aug. 3, 1926] (Au). County un- determined: Belfrage sen. [20--24.).1860] (S), sn. (S); Brace sen. [1872] (W—310733); Collector undetermined 263 (Au—1218)8) 5 Drummond II.s.n. (Lu), IIT.263 (Lu); B. B. Harris Lié (Nt), hoo (Nt); Ness s.n. (Tr); Normand s.n, [Muskogee Railroad, 3.13.27] (au). FRANCE: Beauregard B.132 (M). GERMANY: Gibbes 13-1 (N). CULTIVATED: Belgium: Herb. Jard. Bot. Brux. s.n. [1h Sept. 1889] i RT tens s.n. [h.b-Lov. 1839] (Br), son. [h.b.L.] (Br). Connecticut: Herb. Denslow s.n. [June 1865] (Ms). District of Columbia: Vasey sen. (Washington, from seed] (W). France: Collector undesignated Sen. [jardin de Mr. Clément 4 Montpellier] (Au--1218)8), s.n- {h. P.] (Sg--1609k); Herb. Berg sen. [1839] (Du--166L83) 5 Herb. De- caisne s.n. [h. P. 1829] (Br); Herb. Dumortier s.n- [1826] (Br); Herb. W. H. Harvey s.n. (hort. Burdig. 1815] (Du—16682), 5-De (Du--166)81); Herb. Saldanha 2770 (Ja); B. Jussieu s.n- {1770] (S). Georgia: E. C. Marquand s.n. [April 18, 1931] (N). Germany? Herb. Calif. Acad. Sci. 3146 (Gg); Herb. Martius s.n. (Br, Br); ee etna ttm atta ee Sree an 1963 Moldenke, Monograph of Verbena 67 Herb. Molliano s.n. [h. E. 1807] (Br); Herb. Mus. Bot. Stockholm 8.Nn. gen. [e horto Hamb, 1829] (8S), sen. [h. Berol.] (S); Herb. Prager 18643, in part (Gg--31)58). Illinois: Mrs. Parker Earle s.n. (Aug. 1h, 1880] (Ur); E. Hall 1963 (Dt); Vasey s.n. (Ur). Iowa: Elrod s.n. tone “May 21 21, ae ee 3 Mrs. Fischer s.n. (Bz—-26),27) , XV.K A -XLV. ), (B2--2629). Kan: Sicidad E. Hall s.n. [cult. sp. 1868] (W). Kentucky: C, Peck s.n. [seeds from Louisiana, Aug. 1835] (Ky); C. W. Short sen. [1835] (W--771875). Maryland: McCann sen. [seed from Copenhagen, 9-15-36] (Md). Massachusetts: L. He Bailey s.n. [June 1883] (Ba), s.n, [June 21-23, 1919] (Ba). New Jersey: Ee T. Clausen H,532 (Ba); | Guyot s.n. (Pr); G. H. Lawrenc P.392 (Ba), P55) (Ba). New York: be Bs i. Bailey s.n. [Haage & Schnidt seed 5196; July 25, 1926] (Ba), sen. [Haage & Schmidt seed 5196; Aug. 15, 1926] (Ba), s.n. [Haage & Schmidt seed 5197; July 22, 1926] (Ba), sen. [Haage & Schmidt seed 5197; Sept. 8, _ ] (Ba), son, [Haage & Schmidt seed 5198; July 23, 1925] (Ba), . [Haage & Schmidt seed 5198; Aug. 7, 1925] = “ acts Peta oa ert seed 5198; Aug. 8, 1925] (Ba, N--photo, N--p photo, Z—photo), s.n. fHaage & Schmidt seed 5198; i gts 1926] (Ba), sen. [Haage & Schmidt seed 5198; Oct. 8, 1926] (Ba), S.n. [Vilmorin-Andrieux seed 80050; June 30, 1919] ” (Ba), Sone gets in-Andrieux seed 80050; July 4, 1919] (Ba), sn. [Vilmor Andrieux seed 8006); June 28, 1919] (Ba), s.n. ng {vinortinrs seed 8006; Aug. 31, 192)] (aa), Sen. {root from Shreve, Ark., July 25, 1926] (Ba), s.n. [root from Yom Shreve, Ark., Aug. 15, 1926] (Ba), san. [root from Shreve, Ark., Sept. 30, 1926] (Ba), sn. [root from Shreve, Ark., July 22, 1927] (Ba); R. T. Clausen m H.516 (Ba); Everett & Quinn s.n. [N. Y. Bot. Gard. Cult. Pl. R-ll-36) (Ba, N) Gilly 213 (N)} Moldenke & Moldenke 1188) (N); Quinn s.n. [N. Y. Bot. Gard. Cult. Pl, R.30/36) (N); Servinghaus s.n. [N. YX. Bot. Gard, Cult. Pl. R.51/36] (N); Swift s.n. [N. Y. Bot. jot. Gard. Cult. Pl. R.38/36] (N); Van Melle s.n. wn, (Sept. 10, 1931] erg Me Zimmerman 38 (Ba). North Carolina:L. H. Bailey s.n. [May 1 1930] (Ba); Whiffen s.n. [October 19, 1938] (Hi—3917). ak: Pia R. E. Griffith sen. (Ky). Spain: Herb. Hort. Reg. Matrit. C. C. Albers 3 wate ee famacat Pringle Sept. 1879] Ped West petty Davis & Davis 1165 a: "2083 (We). LOCALITY OF COLIECTION UND hb peered te Il. 31 (C); Bosc s.n. (Caroline, 18)9] (Da—166187 7)3 (tr); ie J. Brewster s.n. (Minn. to Kans.] te Soo e281); 68 PHYTOLOGIA Vol. 8, no. 9 Collector undesignated 5 (Q), 8 (Q), 9 (Q), 10 (Q), sen. (Cn); Fremont s.n. [Exped. to . Calif. 7 (C)3 Herb. W. | He Harvey s.n. (Du- 166,86); Herb. Linnaeus G.790, S.7 (Ls--type, N--photo of type, Z—~photo of type), Ge790, $8 (Is, N--photo, Z--photo); Herb. Mus. Bot. Stockholm s.n. (S, S, S); Herb. Swartz son. (S); Herb. Univ. Ill. 17456 [Plains] (Ur); Herb. Univ. Kentucky s.n. (Ky)3 McCord s.n. (Cn); Rugel s.n. [Ad vias et margines agrorum, per Georgiam infer. et Floridam med., Mart. 183] (Go); Shearer s.n. [Lower Fordyce Bridge, Lower Wabash weet 5/15/1920] 0] (Ur); Thunberg s.n. (S); A. Wood s.n. [Ga. an a.) (Pa). MOUNTED ILL- USTRATIONS: Curtis, Bot. Mag. 9: pl ae aS (N, W), Bs pl. 2200 (N); pps by Prof. Soomtapae “donated by B. G. Amend (N); ds, Bot. Reg. 23: pl. 1925 (Ba, N--photo, N--photo, Z-~ photo Z—phote) Life 2h cli) (62) (N ds Mag. Bot. & Gard. 2: pl. 38 (N); » Fls.&F 2 Se Ae 21. 1879 (N, W)5 2 4 Source dines onl Beas 1h (N), fig. 25635 (He a ens Noldenke, kod- achrome slide 57 (Z). VERBENA CANADENSIS f. CANDIDISSIMA (Haage & Schmidt) Palmer & Steyerm., Rhodora 57: 316. 1955, Synonymy: Verbena aubletia candidissima Haage & Schmidt ex "Y. Ne", Rev. Hort. Belg. 26: 76—-77, fig. 15. 1900. Verbena aubletia candidissima Hort. ex Stapf, Ind. Lond. 6: 29. 1932. Verbena canadensis var. candidissima Moldenke in aitittensén, Roy. Hort. Soc. Dict. Gard. lh: 2210. 1951. Verbena canadensis var, candidissima Haage & Schmidt ex Moldenke, Résumé 360, in syn. 1959. Bibliography: "V. Ne", Rev. Hort. Belg. 26: 76—77, fig. 15. ists Hand Stapf, Ind. Lond. 6: 429. 1931; L. H. Bailey, Cat. Flor- a ldenke, Résrmé Suppl. 2: 3 & 8 ( 1960) and 3:37. 1962. au ations: "V. N.", Rev. Hort. Belg. 26: 76, fig. 15. This form differs from the typical form of the species i having its corollas white, the center sometimes pinkish, Ao Odell sometimes whitish and the tube whitish suffused with pale- Although originaly published as a trinomial in the ge ne om manner of subspecies, the name was apparently intended to h varietal rank, as the use of the exe nyariété" in the aamet sion by the author plainly indicates. The note is signed "V. Ne" so it may well be that the name should be accredited to him since it is said to have been first proposed in the a current 1963 Moldenke, Monograph of Verbena 69 Haage & Schmidt catalogue ra which I have not seen). The original statement reads "La variét nouvelle qui est annoncée par MM. HAAGE et SC re dans leur dernier te tre mace d'un coloris blanc éclatant. 11 rit nds that when growing this is white, or, eee the center has the least pinkish tinge imagineable; itt erm. ish and the corolla tube is whitish with pale lilac suffused throughout." The form is apparently cultivated in En gland and Belgium, as is evident from the 1900 and 1951 items in its liter ature; it is recorded from cultivation in England by me in my "Ré offered by Stecklers and by Haage & Schmidt, but those of the former have proved to be V. platensis Spreng., of which V. candi- dissima Stecklers is a synonym. Cora Steyermark found the form growing wild on an expose limestone bluff alongside of a road ad =e to Long ¢ Creek, F just north of Oasis, in Taney County, Missouri, blooming in April. Palmer & Steyermark (1955) cite ised’ collections not red yet seen by me: C, Steyermark s.n. [T22N, R22N, sect. 3, 10, and 11, April 28, 1919] from Taney County, isso uri. In all, 3 herbarium specimens have been examined by me, Citations: TEXAS: Red River Co.: Sartwell s.n. [Red River, white] Fe CULTIVATED: New York: Sartwell s.n. [seeds from Red River] iy etree ENA CANESCENS H.B.K., Nov. Gen. & Sp. Pl., ed. fol. 2: pl. 136 (1817) & 2: 27h. 1818 [not V. canescens Chapm., 1860]. Synonymy: Verbena canescens Humb. ex Spreng. in L., Syst. Veg., ed. 16, 2: 750. 1825. Verbena canescens Kunth ex Schau. in DC., Prodr. 11: 548. 187. Verbena sanescens H.B.K. ex Mol- escens var. prostrata Don ex Moldenke, Résumé 361, in syn. 1959. een cauesceus oe * + ex Moldenke, Résumé 361, in ~ 1959. ibliography: Nov. Gen. & Sp. oo ed. fol. 2: pl. 136. (1817). & 2: oth 3348) and ed. quart., 2: 27h. 13185 ‘Spreng. in L., Syst. Veg., ed. 16, 2: 740. 1825; rab Hort. Brit. 00s ey B Pl. Hartw. 21. 1839; Walp., Repert. h: 2h. 1845; Schau. in A DC., Prodr. 11: 548--549. 1847; A. W. Chapm., Fl. South. U. S., ed. 1, pr. 1, 307 (1860), pr. 2, 307 (1865), and pr. 3, 307. ra + Wats., Proc. Am. _— gs 18: 135--136. 1883; A. W. Jy Fie Sous V. a r. 1, 307 (1884), pr. 2, 307 (188777 pr. 3 "307 (18895, yer tes l, 307. 1892; Coult., Contrib. U. S. Nat. Herb. 24 327i 1892; Briq. in Engl. & Prantl, Nat. 1 Ind. Lond. 6: 429° 1931; Perry 0. Bot. Gard. 20: 248, 261, 297, 301—302, & 355. 1933; iis: Prelim. Alph. List Invalid Names 56, 190; Moldenke, Suppl. List Invalid Names 10. 191; 4,70 PHYTOLOGIA Vol. 8, no. 9 H S. Gentry, Rio Mayo Pl. [Carnegie ened Wash. — 527:] 222. 192; uoldenke, Alph, List Invalid Names 7 & 50. 19:23 Moldenke, Known Geogr. Distrib. Verbenac., Tea 2) 33535528; 101. 19425 Moldenke in Lundell, Fl. Texas 3 (1): 17, os & eae 32. 1942; Northeraft, Contrib. Dudley Herb. 3: 310. rob * 9 a7. 196; Moldenke, Phytologia 2: 331. 1917; Moldenke, Alph. sige — ere Ley omer a Saag Peso ¥ me & A. L. Moldenke Pl. elles t Git. 2: 393, 79, 497, ser, 8, “393 Gsuby, °3r 63, wa, "772, Toh, 118%» 199 802, 807, iB 2ale 269 94), and hs 1071, 107, 1076, "216, 1289, 1291, & 1295. fous; Moldenke, 7 Rzedowski, Act. Cientif. Potos. 1: 39. 1957; Moldenke, Ame “ms Nat. 59: 356. 1958; Moldenke, Résumé 2, 28, 39, 360 373, & Tle 1959; Moldenke, Résumé Suppl. 2: 3 & 11 119605, 2 & 37 (1962), and h: 1962. Illustrations: ie. Nov. Gen. & Sp. Pl., ed. fol., 2: pl 1%. 2 AG Low, coarse, perennial herb of spreading habit, often with the pel Ps of ltyosotis, sometimes prostrate, usually 15—l6 cm. tall, canescent-hirtous throughout, much branched; stems ee? tetragonal, branched, procumbent or decumbent to ascending 0 rect, canescent-hirsute; branches slender, sulcate, about 1s Pe long, rarely almost simple; leaves rigid, * mostly sessile, gray- hairy, oblong-lanceolate, 1--5 cm. long, acute at the apex, re~ motely incised-dentate or subpinnatifid, contracted or attenuate into a margined and entire semiamplexicaul or petiolar base, lineate-rugose and hirtellous above, the hairs often with minute bulbous bases, canescent-hirsute and somewhat conspicuous veined beneath, revolute along the margins, the laciniations spreading and acute; spikes sessile or short-pedunculate, mostly solitary, filiform-elongate, weak, 5--7.5 cm. loose- owered or compact, glandular-hirsute; lower bractlets often filiform, the upper ones lanceolate, acuminate, variable th it, canescent-hirsute, with glanduliferous hairs intermixed, cil- iate; —— ee anes calyx 2--3 mm. long, canescent—hi with glandu intermixed, subtruncate, the herbaceous veins eiantine into very short inconspicuous mucros or teeth; corolla varying from blue, deep-blue, ote ete ‘ pygore or ie aaah gre to blue-purple, Pp rple, purple, ‘ve-purple, or purple-violet, its tube pelightly ee pei "the calys, its limb l--6 mm. wide, villous in the throat, the lobes the not reaching the tip of the cocci. The species is based se a collection made by Friedrich Hein- rich Alexander von Humboldt and Aimé Jacques Alexandre Bonpland 1963 Moldenke, Monograph of Verbena 471 in Guanajuato, Mexico, and is no. 1111 in the Willdenow Herbar~ ium, Schauer (18,7) pte en the bractlets of this species lanceolate or ovate says r "spicae....in alii is mead remotiflorae bracteisque foliaceis interstinctae, in = pedun: culatae bracteis et infimis integerrimis et nunc * omnibu ns, Coal excedentibus nunc imis tantum calyce paulo serge ease cit ibus vero eumdem modo aequantibus," This of the characters of typical V. canescens pla of its variety roemeriana (Scheele) Perry. In its typical form, this species has been found by collectors r e sandy or sandy clay loam, on roadside banks and in moist depres- ions, dar savannas and in cactus-acacia grass with coarse rocky soil, in sandy loam bottoms, in scrubland on lime- stone mountainsides, on cliffsides along arroyos and on Oa erate banks in deserts, and in clayish soil in desert scrub, er at altitudes of 170 to 2665 meters, It has hehe collected = anthesis and fruit from February to November, In Lundell's ora of Texas" it is said to be found "in sand and oe seg careous hills, valleys, and canyons, blooming fr pril to October. McVaugh found it on brush-covered hills ~ shrubs and small trees of Acacia, Mimosa, and Opuntia. Runyon re- rts it as = eing "common on acuntainsiien®, Stanford, Retherford, & Sorthozer’ say that it is found "on valley floor sparsely cov- ered by Yucca and Larrea"; Muller found it "abundant in neh colonies" in Nuevo Le én, while Mueller & Mueller report it "sparse" in the same state. Wiggins encountered it on Riteeetene imen. ve been misidentified and distributed in herbaria as V. swam yer Michx., V. canescens var. roemeriana (Scheele) Perry » V. gracilis Desf., V. neomexicana A. Gray, V. neo-mexicana A. Gray, V. neo-mexicana (A. Gray) Small, V. plicata Greene, V. remota Benth,, and V. xutha Lehm, The Moore @ & Wood 113) is a mix- ture w. with V. menthaefolia Benth. Heard Webster, & Barkl 1,509, “A. A. Heller 1732, Li C. Hinckley 125), Lindheimer 107 & 111-500, Edw. Palmer 10LL, Rozynski 13, Stanford, Lauber, & “& Taylor 2252, Tweedy 113 & 246, and M.S. Young 112, all distributed as as Ve canescens, are e actually v. reais, (Scheele) Perry; Pataky sen. [Fort Huachuca, 1890] and Pringle s.n, (Gardiner's Spring, Jyne 25, 1882] are V. gracilis 472 PHYTOLOGIA Vol. 8, no. 9 Desf.; L. F. Ward s.n. [Hearne, Sept. 11, 1877] is partly V. halei small and partly V. xutha Lehm.; hi M. Ferguson s.n. ~ [April 1901], Parry, Bigelow, Wright, . & Schott s Son. [Valley of the Rio Grande below Doflana], a and S. S. White 2202 are V. neomexicana (A. Gray) Small; McKelvey 2046, Ce ae Mueller 2391, P Purpus 308 & 3927, Tharp sen. [March 21, 1932], S. S. White 2168, and E. Wil- = Sone sta. fireanes hills, July 30, ~1885] are V. neomexica- irtella Perry; M. E. Jones ot. (koe 5; 1903] & sone net ‘yale, April 28, 1903], oe 270, Shreve 9097, Stan- ford, Retherford, & Northeraft 426, J. J. Thornber 8lll, E. Wil- kinson 37, and T. E. Wilcox s.n. [Fort Huachuca, Aug. 1392) are V. neomexicana var. 's Xylopoda Perry; Havard s.n. (Guadalupe Mts., Oct. 1881] is V. perennis ‘ennis Wooton; M. E. Jones 28298 is V. quad- rangulata Heller; S. Ge “S$. G. Stokes Sone (Grand Junction, June 6, 1900] is V. bracteata L Lag. & 2. & Rodr.; and F. Salazar s.n. [Nopala, Aug. l, 1913] and Shreve 80 80h, are Ve gracilis ia The species is placed by by Walpers “3 (1845) in Section .Verbenaca, Subsection Inermes, Group Foliosae, sae oe Second- ary Subgroup Schizophyllae, along with V. xutha Lehm., Ve radi- cans Gill. & Hook., V. microphylla Wiax.: i soles Desf., V- officinalis L., V. supina L., V. bracteata Tag. & Rodr, and what is now called Hierobotana inflata (H.B.K.) Briq. Briquet (1894) speaks of a "V. neo-mexicana Briq.” which he says is "V. canescens x officinalis", by which I assume that he means V. heomexicana (A. Gray) Small, but I do not regard this taxon as The nam a hybrid. name V. gracilis Desf. is given as a synonym of V. canes~ cens by Schauer (157), by Hooker & Jackson (1895), and by me me & Jackson also cite V, remota Benth. as a synonym of VY. canescens, and the two latter authors dispose similarly of V. parviflora Rufz, but these two names belong in the synonymy of ve. gra sa Goulter (1892) eager cited in the bibliogra aphy ab ometimes erroneously given as page "587" and dated mag", or) the barton (1839) reteretee as "18,6", It appears also that, in spite of the title page date of "1817" on the H.B.K. reference, the text page appeared Psi in 1818, while only the illustrative plate was issued in It should be noted here that the V, canescens Chapa. is Ve bracteata Lag. & Rodr., V. canescens var. arizonica Gray is pws neomexicana var. xylopoda Perry, and V. canescens var. neomexica- na eon y Vv. neomexicana (A. Gray) Small. Ac n name recorded for the plant is "canescent verrehie Williges ante on his no. 427 "probably the same as no. 333, but 1963 Moldenke, Monograph of Verbena 473 the fruits seem to be spaced farther apart and the calyx looks smaller, and the red lines on the x seem to be ipa beh " Stanford, cae & Northeraft 486 appears to be a mixt with V. | perennis var. johnstoni Moldenke, while their no. 198 as well as Waterfall & Wallis 13806 are mixtures with ve. V. ciliata eee The L98 collection, ci cited hereinafter from Zacatecas, may tually have « come from Coahuila -- its label reads "15 kilo wet of Concepcion del Oro on the Coahuila-Zacatecas border, lat. 2°56)" N. long. 101°)5' W." The town referred to is in the state of Zacatecas and this state's territo ory — for a considerable distance west and a fair distance north of t e town, so the chances are that the collection was made in rel state. Howard s. Gentry 557m, cited below fron eae is from Mesa Colorado on the Sonora-Chihuahua boundary an may vi tualy hav come from Sonora, In his "Rio ee plana" *Ca9h2) esis lists this collection as "Verbena sp." and cites it along with his nos. 1522 and 1923 (which are V. V. pinetorum Mébaetiee} and no. 1758 ~ (which I have not seen as yet). He sa He says that they inhabit "Open slopes and meadow margins in Pine Forest. 1,000--6000 feet. Peren- me, as well as Hartweg 17h, which, however, is the type collection of V. remota and is therefore V. gracilis. He saw these collec- tions in the Berlin herbarium, now destroyed, Perry (1933) cites the following 27 additional specimens not as yet seen by me: MEXICO: Aguas Calientes: Hartweg 177 (G); Rose & Painter 771 (G). Coahuila: Adole 22 [Arsene 10626] (E, F, G); Gregg 99 (E), 2h6 (E), SUS (E), 740 (E); Edw. Palmer 25 (E, F, G), 10h7- (D, G). Oaxaca: Pringle 478k ( (D, E, G)3 Le Ce Smith 412 (c) . San tas Potosf{: Purpus us 5515 Y, (F, 6); J. Ge Schaffner 719 (D, G). Vera Cruz: Seaton 322 es a trae esete Edw. Palmer bac = F, Pablo i on eae her c cteristic compact habit. The specimens cited from Coahuila are meine nt examples of phases intermediate between the species and the variety. The collection from Nevada seems somewhat out of ranges. Fopesly it was carried in as a weed or escaped from cultivation . ase cuunetipine ens Saigon: ‘oak Co.: M. E. Jones 55h (Po 2N6e36). TEXAS: Bander «: Johnson & Webster 566 (Au--121910) . Bexar Co.: Metz 3019 sere hin SEE Coe: Reeves 206 (Cs). Guadalupe Co.: John- 47h PHYTOLOGIA Vol. 8, no. 9 ston & MeCart 5096 (Au--121953). Presidio Co.: M. C. Johnston 352 (Au--168707). San Patricio Co.: Williges 286 (Au--179721), 333 (Au--179805), 427 (Au--179846). Willacy Co.: M. C. Johnston 53253. (Au--122)28). County undetermined: Mrs. G. A. Black s.ne —<—$_—— I ————————— weg 177 (N); R. McVaugh 16639 (Mi); Rose & Hay 6212 (N, W-— 396025); Rose & Painter 7741 (N, W--l51358). Chitmahua: Howard S» Gentry 557m (Mi); Gregg s.n- [San Pablo, 29/47] (1); E. Wilkinson sen. [30 July 1885] (I), s.n. [Santa Eulalia Hills, Sept. 19, 1885] (Ca--192070). Coahuila: Adole 22 [Arséne 10626] (Bm, Cm, N, Ur, W--1030229); Barkley, Webster, & Rowell 7189 (Au--170081) ; Hinton 16636 (W—1947239); Johnson & Barkley 16259M (Au); Kenoyer crum 2579 (MH), 2951 (Wi); E- Ge Marsh 278 (Ok), 575 (Ok), 1687 (St), 210 (St); R. McVaugh 12310 (Mi); Edw. Palmer 25 (Me, N, S, W—3%010), 1047 (Pa, W--5619h, W--13231h1); F. W. Pennell 17305 (Me, Me, N, N, W--1640L95); R. Runyon 1321 (W--147)119); Stanford, Retherford, & Northcraft 426 (Du--286)19, Tu--10892), 486, in part (Du--288703, N), 498 (Ca--713893); Warnock & Barkley 14852 (Ca—7h78)2), 14852 (Au). Guanajuato: Spivey 151 (Ca--91678) « —__ eee H. Mueller 2305 (N); Mueller & Mueller 465 (Mi), Sn. [Mayo 19, 193) (Me); F. W. Pennell 16792 (D—733972, W-—-160232), 16881 (D-733973, W—160299); Waterfall 15325 (Ok); Waterfall & Wallis 13217 (Ok). Oaxaca: C. Conzatti 19h (Me, W--1082260), 4288 (Me, W—1082312), 4992 (W—1696339); Galeotti son. [Tlajiaco, Juin 18h4] (Br); Pringle 78 (Br, C, Ca--10481h, Cm, Io-—38732, Me, Me, Mi, Mm—15391, S, Vt, W-—251676); C. L. Smith 221, in part (Ca—-975386, N). Puebla: Arséne sen. [15/6/1907] (Br); Balls 5266 (W—1793806), B.5266 (Ca—68L513); Chute M.163 (Ii); Nicolas 80 (Gg—-31273), sen. [Cerro Guadalupe, 20/6/1909] (W--li6L379), Se n. [Rancho Posadas, 20/6/1909] (Br, Br), sen. [Atoyac, 20/6/1910] (W~L64380); H. Pittier ih (W--570809); Wiggins 13261 (Du— 404631). San Luis Potosf: G. L. Fisher 3729 (W--1725138), 37029 (Ew, Ew), son. (Cardenas, Aug. 21, 1937] (Fs, N); Parry & Palmer TS (Io); Purpus 5515 (Ca—-157217, N, W—l6001); J. G. Schaffner 719 (Me); Shreve 934) (Ca--731688, Fs, Mi). Sonora: Gregg S-M.- TBuenavista, 22/7] (T). Tamaulipas: Graham & Johnston L707 (Au-- A TD 1963 Moldenke, Monograph of Verbena 475 174972); Painter & Barkley 1373 (Au); Rose & Russell 2317 (w-- 1369606); Rozynski s.n. (B); Stanford, Lauber, & Taylor 217ha (N), 2252 (N); H. W. Viereck 229 (W--1687119). Vera Cruz: L. I. Da Davis 203 (N); F. Miller 1215 (i); H. E. Seaton 322 (W--56175). Zaca— tecas: i S. Gentry 8555 (Ny, " We-2022211); 5 Edw. Palmer 303 (N, W--471173); Purpus urpus 167 (1 (W—,70278), 469 (Ca--138833); Stan- ford, Retherford, & Northeraft 98, in part (N, N, Se--70h51); Waterfall 15570 (Ok); Waterfall & Wallis 13806 tn part (Ok). nee aes undetermined: Galeotti sn. (Br, Br); Halsted sen. [Mexico] VERBENA CANESCENS var. ROEMERIANA (Scheele) Perry, Ann. Mo. Bot. Gard. 20: 302. 1933. Synonymy: Verbena roemeriana Scheele, Linnaea 21: 755. 188. Verbena cauesceus var. roemeriaua (Scheele) Perry ex Moldenke, Alph. List Invalid Names Suppl. 1: 23, in syn. 19)7. Verbena canescens var. roemeriana (Gray) Small ex Moldenke, Résumé 31, in syn. 1959. Verbena cauesceus var. roemeriana agai, Perry ex per nag, Résumé 361, in syn. 1959. Verbena cane riana (Scheele) Perry ex Rzedowski, Act. ant Potos. lj: Ui, 6 65, & 91. 1961. Verbena texana small ex Moldenke, Résumé ppl. 3: 1, in vi 1. 1962. —— 30 Re w re . we ww a 2 5p - 4 Pe 3 © 3 a a dg a enke aes er. Dis Verbenac., [ed. 1], 13, 18, "a hol A9e Makers , Ciatates, 10: 2 3 Moldenke bw : 328. 19h7; uoldenke, Castanea 13: 112 & 113. 1948; He Ne & A» Le Moldenke, Pl. Life 2: 79. 1948; Molden- ke, Wrightia 1: 228--229. 1918 exe a jAiphe List Cit. 2: 370, 396, 470, b7k--47, 79, 83, Lohe 5 306» Sly 516, $20, 522, 527, 221 a1, 238 ; 272, 55, 592, : 33 (1918), 3: 656, 66h, : » 678, 683, 708, ne, 19, spe Bog » 12, Be 752, 773, 78h 788 789 70, 796, ’ 843, 850, 878, 879, 881-883, 891, vy 8, & 934 (as 9), h: hy: 982, 989-993, 100k, 1014, 10 78, 1098, 1099, 11 1103, h 2 38 1s, | 1150, ’ si 1155, 1158, 1162-116, 1170, 1171, 1182, 1189, 1207, —— 1217, 1231, 1238, 1239, 1246, 1253, 1254, & 1289. 1949; e@, known Geogr. Distrib. etigewet feds 2], - 23,‘ 27,: 32, & 197. 1949; H. Ne & Ae Le Moldenke, Anal. I 12. 1949; Moldenke, Phytologia 7 2. 1949; Shinners, Spring 4,76 PHYTOLOGIA Vol. 8, no. 9 Fl. Dallas 326. 1958; Moldenke, Résumé 28, 33, 39, ky :3I3 5m 71. 1959; Moldenke, Résumé Suppl. 2: ll. 1960; Rzedowski, Act. Gientif. Potos. h: hh, 65, & 91. 1960; Lewis & Oliver, Am. Journ. Bot. 18: 639-~641. 1961; Moldenke, Résumé Suppl. 3: 7, 37, & 1 962 Illustrations: Lewis & Oliver, Am. Journ. Bot. 8: 60. 1961. This variety differs from the typical form of the species in ere 30 cm. tall; bark pale or gray; roots fibrous; leaves oblong in outline, pinnatifid, canescent; flowers odorless, in terminal spikes, open n the morning; corolla varying from blue, lavender-blue, blue- Le, purplish-blue, pinkish-purple, or lavender with a whitish "eye"; an = 14 9 * The variety is found in dry rocky or sandy soil on clay slopes and hillsides, in ravines and open woodland and in cedar cal prairies, showing a preference for dry rocky hillsides, in Texas, blooming there from February to October (mostly from March to June), and is widespread in central, southern, and western parts Counties. Collectors have found s dry limestone or clay soil, black or dry calcareous soil, in deserts and arroy' b s, in sandy loam bottoms or in clay-loam soil, on limestone am granite outcrops, along roadsides and railroad rights-of-way, on creek banks and limey-clay hillsides, near ponds, among rock and gravel in xerophytic canyons, in open vated or waste fields, limestone rock and gravel soil, or chapar- ral, on forested hills and rocky hilltops, in rocky limestone valleys, among coarse gravel and alluvium, in wet ravines and desert scrub, in scrub on ridges, and in xeric serubland and limestone-clay loam. Clemens & Clemens say that in Texas it inhabits the Tropical Life Zone; Warnock found it "infrequent on cemetery knoll" in 1963 Moldenke, Monograph of Verbena 477 in "extremely dry areas, old fields" in Tamaulipas, while Water- fall found it in a "flat valley with acacias and other shrubs" in Coahuila and "on Baie desert with Larrea and Fouquieria" in Chihuahua, Tra e found it in deep low scrub on buff-gray silt in Celtis Ra Ssehep ss the Websters encountered it on xeric limestone hillsides; Graham & Johnston in brush with isaciks rig- idula Benth. on a hills and on caliche spots; and M. C. John- ston in shallow sand over caliche in short brush-Calliandra erio- oLyiten oat gat torh Johnston & McCart report it as growing in loam with much 1 stone gravel on gentle slopes. McVaugh says that. it ia? tainindant and weedy" on rocky limestone hills in Val Verde County, Texas. It has been collected in anthesis and fruit from sca to December. It is most probable, therefore, that it flowe fruits throughout the year. rt has been collected at aithtualbe of 1.2 to 150 meters. Material of this variety has been misidentified and distribut- herbaria under the names of V. bracteata Cav., V. bracteosa Michx., V. bracteosa var. brevibracteata Gray, V. canadens is B<) Britton, “2 canescens H.B.K., Vv. caroliniensis (Walt.) Gmel., ve cloveri Moldenke, V. hastata L. - V. imbricaria Woot. & Standl., V. neomexicana (A . Gray) Small, V. , neomexicana Small, V. neo-mexicana (A. Gray) Small, Ve. n neo-mexicana Small, Ve neomexi- cana var. hirtella Perry, V. neomexicana var. TS ry Perry, V. officinalis L., V. plicata Greene, V. urticaefolia L., V. xutha Lehm., and even Phyla Sp. On the other hand, the Warnock & Barkley 1852 and Waterfall & Wallis 13217, distributed as as V. c. canescens var. roemeriana, are actually typ typical V. canescens y HeBeKe3 Hamby 1637 is V. cloverae Moldenke; Bryant & & Bryant 3 is V. halei i Small; 0 Owens 3125 is V. neomexicana var, xylopoda Perry; and Tharp 43-801 is is V. V. plicata re The “varkety is named in honor of Karl Ferdinand Roemer (1818-- 1891), German professor of geology A paleontology at Breslau, genus as a whole, as well as to in it and even in related genera (see under V. bonariensis The Hinckley, Warnock, & Tharp §.n. (8/22/L7] collection ‘te very glandular and looks like a form of V. neomexicana, but its bractlets are very large and broad’ Perhaps it represents a hy- brid, Bray, Ramsey, Beck, Long, | Wild, & Ferguson suggest that their col lection, cited be low, "may yh stout form of V. canes- cens." Warnock s.n. [May 3, a7 J. Reverchon 737, and Tharp 13-800 are mixtures with ve. plicata G Greene, Parry, y, Bigelow, 478 PHYTOLOGIA Vol. 8, no. 9 Wright, & Schott 827 is a mixture with ¥, xutha Lehm., while G. Jermy Jermy 87 | and M. | M.S. Young s.n. [4/5/18] are mixtures with V. halei The Small & Wherry 11969, cited by Perry (1933) and var by Texas, may actually have come from some other county, ‘anes its label *merely states "San Antonio to Austin" and Dr, Small's orig- inal field book gives no further information about it. Whitehouse SMe, soni from Menard County, may have come a Concho, MeCul- loc loch, n Counties since its label merely says "Maso n--Eden®, but the Srageaher tells me that it was "probably" can laned in Menard County. Similarly, Tharp 8588, cited from Terrell County, may be from Val Verde County -- the the label is ae scribed rg afon ae A mn", but Dr. Tharp, in a letter to me, iad *probab=- ly in Terrell County". Lindheimer 107k was apparently not collect~ ed in Comanche County, as some workers have thought, but in Comal County, because "New Braunfels" is definitely unde rlined on the label of the University of Texas sheet. The "J. F. Norman" appear~ ing on labels in the same herbarium is apparently an error in transcription for J. F. Normand. Tharp s.n. [June 19], Ecology Class Univ. Texas s.n., and Painter r 29 were cited in Lundell's "Flora of Texas" as = neomexicana ree hirtella, but prove to be V. canesc meriana inst Perry 71533) tee the Fotlowine re — specimens not as yet seen by me: TEXAS: Bexar Co.: Clemens & Clemens 97) (E); Groth 16 (F, G); E. J. Palmer 9183 (E). ae Co. Co.: EB. J. J. Palmer 13283 (E). Brown Co.: i de J. Palmer 29535 (E)j Je Reverchon 1961 (G). Comal Co.: Lindheimer r 29h (E), “1074 (E, F, G). Duval Cos: Clark 401 (£). Gillespie Co.: Jermy 182 (EB). Goliad poe Eggert sen, [near Goliad, 9 April 1900] (E). Hays Co,: Trelease 5.n- ‘San Marcos, 6 Nov. 1897] (E). Kerr Co.: A. A. Heller ry > (E, G) E. J. Palmer 10002 (E). Real Co.: Berlandier 6h, in part (G), 205h (G). San Patricio Co.: McKelvey 1710 (G). Terrell Co.: Me Ee Jones 26226 (E). Tom Green Co.: ‘O.: Reverchon 3903 (E). Travis Co.t E. E. Hall 1 430 (E, F, G); M. S. Young 112 (E). ", Uvalde Co.: Moore & Steyermark rk 3005 (); E. J J.P “Palmer 13563 (E), 33605 (E). Val Verde Co.: Eggert son. {Devil's River, 10 Sept. 1900] (E); Orcutt 6111 (E). County undetermined: Lindheimer 500 (E, G); Reverchon Bi ’ in part [1885] (E). MEXICO: Nuevo Leén: Gregg 791 (E E); Edw. , Palm er 104) (D, G); Wislizenus 325 (E). Tamaulipas: 3; Berlandier ay (E, G)5 Edw. Palmer 82 (E, F, G); Rozynski 13 (F), 17 (F), 135 (F). She comments that the variety is "Differing from the species chiefly in its coarser less gl ar pubescence and ovate a te bracts. A few of the Spiegel speci- mens show transitional phases, but, as a whole, are closer to the variety". The "Berlandier 827m which nf cites is - Asiaactbed "8 3h- 1963 Moldenke, Monograph of Verbena 79 827" in the Torrey Herbarium, Her "Canby 19k" is cited hereinaf- ter as Canby, Sargent, & Trelease 194. In all, 458 herbarium specimens of the variety have been exam- ined by m Citations: TEXAS: Bandera Co,: Bogusch s.n. [April 10, 1926] (Ob--50813); V. Sutton sen. (Medina, 10.12.1929] (Au). Bee Co.: J. S. Williams 39378 (Au). Bell Co.: McDaniel 12 (Nt); Wolff 539 (W—1623900), 3L8h (' (Tr); York & York 54448 (Au—121903). Bexar Co.: R. Bebb 2251 (Ok, Ur); Bogusch s.n. [April 16, 1925] (Ob-- 50813); Clemens & Clemens 971 (Ge——37132, Po—-6959h), 972 (Po— 69598), 973 (Ba, Mi, Po—-69592), 97h (Ba, Po--69602), s.n. [San Antonio, “April 22, 1911] (N); Groth th 16 (N, W--717636); G. G. Jermy St, in part (W. 315686), s sen. [San Ant.] (N, W—392396, a7 2 J. Jermy s.n. [Bexar Co., 1903] (N); Mauldin s.n. [May J (B1—-L2332); Metz 76 [Aug. 21] (I), 76 [Aug. 2 ay (1), 3a oy 2478 (N), son. [July 17, 1931] (N); H. N. Moldenke 6981 (N); E. J. Palmer 9183 (Du--205598, re )j F. W. Pennell 10358 (N, RE); 3 E. D. Schulz 526 wrens Wi), s.n. ~ [lassen Ranch, May Whelan 196 Gn: Brewster Co.: Parken 3683 (Au); G. W. Brown B, 102 (Au); Goodman & Waterfall 581 (0! (Ok, St); Hinckley y 61h (w— 1972322); Hinckley, Warnock, & . Tharp SN. (8/22/47) ( (Au); Lewis & Oliver 5156 (Nb); Tharp 253 (W—1873782), s.n. [6/19/31] (N), s- n. n. [10-9-36] (Au, Au, Ca—082669, Mi, Pl--151576, Sm); Warnock s Se re [May 3, 1937] (Au). Brooks Co.: Perkins & Hall 2637 (Mg--51, Po--256895) . Brown Coe: J. Reverchon m 737, | in part (W), sen. [Cur- tiss 1961] (Bc, C, Cm, Ka, Pa). Cameron Co.: C. L. Lundell ell 10702, in part (Sm), 1077 (N, Rf, W--1926935); Lundell & — 10023 (Ld, N), 10760 (Sm); Re Runyon 337 (Au, Au, Au), 629 (Rr, W- 1224222), 1532 (N), 1778 (Rr), 1779 (Rr, Rr), 2186 (Rr), 2u92 (N, N, N), 2518 (N, N); Tharp 1873 (Au); Tracy s.n. [Olmito, [7is/05} (Tr); Traverse 1093 (Au--179278). Coleman Co.: Warnock 16334 (Au, N). Comal Co.: C. C. Albers 38004 (Au); Lindheimer 294 (Ka), 107 (Au, Br, Ca--1)7486, Me, Me, N, Ok--10349, Up— 39, W—502L)7) ; B. H. Warnock 462k) (Au, Au, N, N, Ok, S, St). Coryell Co.: John- ston ¢ & McCart 5409 (Au—-121955) « Crockett Cost M. E, Jones 26222 (Po=~1783)1) ; Warnock & McBryde 15175 (Rf). Dallas Co.: Collector undesignated s.n. (Hight and Park, 3-29-19] (Wi). Dimmit Co.: Cory 29418 (N), 29419 (N). Duval Co.: Ripley & Barneby 9027 (N); Tharp 5903 (Au). Edwards Co.: Cory 903k (Fs); M. E. Jones 26223 (Po 1783)6) ; 3 Parks & Cory 20841 (Tr--16189); S Shreve 9928 (Fs). Gilles- bade Co.: Whitehouse s.n, [near ore ee June 6, 1929] os uadalupe Co.: Wagner & Barkley 16738 N). Hays Co.: G. Le Fisher 67 (W._1101872 72); Lundell & & Lundell 10278 (Ld, N); S. We 4,80 PHYTOLOGIA Vol. 8, no. 9 N, N). Hidalgo Co.: Cameron 239 (Au), obs (Au); Clover 1839 (Mi); Pande & Lundell 9819 (Ld, N); R. Runyon 2733 (N, N, N, N) Mrs. E. de Walker 30 (Au, u, N), 3h (Au, “gu), kg (Au, Au, N), s.n. {Feb. 9, 1942] (Au, Av, N); MeL. Walker 66 (Au). Howard Co.: .: Havard san. {w. of Big Spring, Sept. 1831] (W—-221212) . Jeff Davis Co.: M. Me E. Jones 2622), in part (Po--178345); M. S. Young s.n. Beis Mts., 9/5/18] (Au). Jones Co.: B. Adams 22 2 (Nt). Karnes es Co.t Je Johnson 1525 (Z). Kendall Co.: .: Parks & Cory 12929 TS Kerr Co.: A. A. Heller 1732 (C, Ca--104815, Mm--15395, W--213L2); Milligan s.n. (Kerrville, May 1898] ae eee E. J. Palmer 10002 (Du--205613, S, W—1532015) . Kinne : D. S. Correll ee (Rf); Mearns 1237 (W--229621), 1338 rg 1456 (W— 30088); A. R. Moldenke 175 (Fg). Kleberg Co.: Bogusch 5.61 (W-— panes a Re R Runyon 4283 (N, Sm); Tracy & Ness s.n, [Kingsville, 4/10/1905] (Tr). Live Oak Co.: M. C. C. Johnston 52054 (Au--121920, St). Llano Co.: Correll & Correll 12701 (Sm); Wolff 3064 (Tr-- 16140). Mason Co.: Dapprich 7766 (Sm); Westlund 85 (Au). Medina Cos: C. L. Lundell 10986 (N, Rf); Small & Wherry “11980 (N). Me- nard Co.: Whitehouse sen. [Mason-Eden, 8. 31,1929] (Au). Mills Co.: Egg s.n. [Mullin, Apr.-May 1931] (Au). Nolan Co.: Herb. Un- iv. Texas s.n. (Sweetwater, 5-20-1931] (Au). Pecos Co.: Tharp 43-800, in part (Au, St); Warnock & McBryde 14908 (Rf). Presid- io Co.: Hinckley 125) (N, N), 2709 ~(N), 3218 (N). Red River Co.: Gregg 791 (T). Reeves Co.: Tharp sen. [Pecos, 6.1h.31] (Au). Runnels Cos: Lundell & Lundell ell 10162 (Mi, N); Quillin s.n. [Bal- Linger, 5/24/31] (Wi, Wi). San Saba Co.: Wolff 238 (Tr). San Patricio Co.: McKelvey 1710 (W—-1581393). Scurry Co.: B. Be ris s.n. [7-)-27] are Starr Co.: D. S. Correll 14887 ( (RE, RE)5 Ecology Class Univ. Texas s.n. [2.28.30] (Au); — & Lundell 9796 (Ld, N, W--1857595), 9930 (Ld, N, W—187)87); R. Runyon 2559 (N, N). Sutton Co.: Babcock sen. [Cory 27780] ae 0. Deg- ener 5078 (N); G. L. Fisher 50101 (Ew); R. W. Pohl 760 (Sn); Rohrbaugh 385. (Ok). , Terrell Cos: M. E. Jones 26226 (Po—1783L8) 5 Tharp 8588 (A (Au). Tom Green Co.: F. re We |, Pennell 1050 (N); Ce Smith 19 (Ok), 143 (Ok); Tweedy 113 : (Ww), ), 26 (W--1)7571), SMe [1879] (T). Travis Co.: Adameik & Barkley 16T2h0 (Au, N)j Ce t, 5. Albers 32016 (Au), 34009 (Au); Albers, Barkley, & Warnock 6h55 55 (Au, N); Armer 5379 (Au, W--1)38993), s.n. [Austin, l-2-29] T ON); Barkley & Rowell 11 (Au), 42 (Au, N), 7061 (Au); Bogusch 76 (W--1285721); Bray 100 (N); Bray, Ramsey, Beck, Long, ong, Wild, & Ferguson s.n. [mouth of Bell Creek, Oct. 12, 1900] (Au); Cohn & & Barkley 13148 (N), 1518 (au); E. Hall 430 (N, Pa, Po--69310, W); Joh Johnson & Barkley 167198 (Au); Johnson & MeCart 057 (Hi-- Stanfield s.n, [San Marcos, 1898] (N); B. H. Warnock 16538 (Au, 1963 Moldenke, Monograph of Verbena 481 117747); Lundell & Lundell 8860% (Mi), 8876 (Ld, Mi, N, N); Me bre 1639 (Au); R. H. Painter 249 (Au), 392 ta; Ky--2716) ; C. M, Ro Rogers 6726 (Au—-121937); Rowell & Barkley 16T214 (Au, N) ; Trak (Au, N); Rowell & Mann 13 (Au); R. Runyon 1532 (S); Tharp 1099 (Au, N), Sen. Austin, 5727/20] (Ca--882668, Po: o—-1613 i i gia Ur), | » SoM [Austin, | Spr. 193] (Au), sen. [11-17-39] ae, Sm), son. [7-18-l,0] (Au, Au, N), sen. 1s [7-19-10] (Au); Tharp, Barkley, | & Warnock 46271 (Au); B. He H. Warnock 86 (Au, N), 46272 (Au), héus (Au); Webster & Webster 115 (Au--1219),7) ; Me S. Yo Young ll2 (Ca--882626, Ur), s.n. - [10/10/13] (Au), sn. (Mt. Bonne nell, 11/22/13] (Au), sen. (Austin, )/25/1)] (Au), sn. [L/29/1h] (au), sen. [Austin, )-1-1918] (Au, Mi, Mi, Mv, N), son. [4/5/18] (Au). Uvalde Co.: M. E. Jones 28292 (Po--18797h), 28296 (Po—-188286, Po—-188287); C. L. Lundell 10956 (N, Rf), are 972 (N, Rf, W— 1926938); Lundell & Lundell - 1026h (Ld, Mi, N, Rf, W--1888913) ; E. J. Palmer 33605 (N), 3366 (N); H. Re ee 33618 (N, NW). Val Verde Co.: D D. S. Correll 14970 (Rf); Hamby 977 (Ar--206763); M. E, Jones 26225 ; (Po—1783h7), 28296 (Ca—l79298, Gg--237868) ; Kc Vaugh 10599 (Mi); A. Re Moldenke nke 17h (Fg); Parry, Bigelow, Wright, & Schott 827, in part (W--56198); Tharp s.n, [Langtry-- Dryden, 6-1)-1931] (Ca-—-882670); W. M. Thompson 222 (Au—16h)96); « H. Warnock 11238 (Rf); Warnock & & McBryde ryde 15046 6 (Au--1232%, RE) ‘ » 15101 (Rf). Waller Co.: Normand s.n. [Waller Creek, h//28] Au). Wharton Co.: Small & Wherry 11826 (N), Sen. Ger Wharton, April 12, 1925] (W—1739113)- Wichita Co,: Tharp 1361 (w— 111412). Williamson Co.: Bodin s.n. [Georgetowm, = 22, 1889] (Ka). County undetermined: G. A. Black 36-692) (Be—1:7626) ; Gol- lector undesignated s.n. (Wi); Herb. Univ. Texas s.n. (Au); irs. W. Re Hughes 26 (N); Lindheimer ITI .500 (Ca—2h62hh, | W), Sn. {Southern Texas, 18)9--51] (N); Mebane s.n. (Spring, 1926] (Wi); Normand s.n. [Central Texas, June 28, 1927] (Au); Small & Wherry 11969 [San Antonio to Austin] (N); C. Wright s.n. [Texas] (T). CALIFORNIA: San Diego Co.: Edw. Palmer 312 - 312 (Pa). | County undeter- mined: Eastwood s.n. [Southern Pacific Railroad, April 1908] (Gg—3131h). KEXICO: Chihuahua: Waterfall 1287 (St, Z); E. Wil- kinson s.n. [19 Sept. 1885] (Io—9222h). Coahuila: Herb. Inst. Biol. Univ. Nac. Mex. 7138, in part (Me, Me); Hernandez Corzo, Rowel Rowell, & Barkley 16M500 (Au, N); Johnson & Barkley rkley 1622hM (Au), 16260 (Au); Kenoyer & Crum 2587 (Mi); E. Ge Marsh 278 (Au), oh (Au), 1687 (Au); Warnock & Barkley 14852 (N); Waterfall 15302 (St); s.s S. White 1759 (1h); Wynd & Mueller 93 (Fs, I, Me, Me, Mi, N, Ur, Ur, W--1639573), 193 (Fs, 1, N, S, St, Ur, Ur, W— 1639685), 253 (Fs, I, N, Ok, S, Ur, Ur, Wi—-1639 706) « Hidalgo: 1,82 Fea t-% OL:0-62% Vol. 8, no. 9 e sen, [El Salto, 15 Sept. 1902] (Me, Me). Jalisco: Mearns 1) Ti89 (W--228 336). Nuevo Ledn: Alonis, Johnson, & Barkley 16185M (Ax); Barkley & Barkley 17317M (N); Barkley, rkley, Webster, & Rowell ter, & Rowell 7181 (Au--123252, N)3 J N); Je Te “T, Buchholz Son. Sen. [Feb. 23, 1936] } (ur), son. (Feb. 2h, 1936] (Ur); Canby, Sargent, & Trelease 19h (t= 369718); L. = Davis s.n. {Santa Rosa Canyon, = 22, 196) 235 hlcsyangol.lyjeca Son. sentheney, July 26, 1926] (pu—15),568, Fs, Hp, Mg-—26); Frye rye & Frye 2362 (Ca—-88262h,, Du—285),81, Hp, N P1—111152, Rs--25233, ~ ge=-65689, W—1790993); Howard S. centers) 6627 (N), 6727 (Ak—22936, W--1978725); Graham & Johnston )586b (Au--171597); Heard, Webster, & Barkley 14509 (N, W—213L279) 5 Hernandez Corzo, Rowell, & Il, & Barkley 16M5 16M53 (Au); Johnson & aa 16041M (Mi, Ok, Ur), ry, 160aLI" (Au) j (Au); Lewis & Oliver 541) (Nb Mueller & Mueller 155 (Au, M (Au, Me, Me); Painter, Luca Lucas, & Bat 11,276 (. (Au); Edw, Palmer 104 (Pa, W—56192, W—1323067); Salil Salinas, Rowell, & Barkley 16M371 (Au); Tharp 1807 (Au); Webster & . Barkley 193 (Au); S. Se S. 5. White 1407 (Mi, Oa), 1ky2 (Mi), 1572. (Mi, Oa, Tu—35535). Tamaulipas: H, H H. Bartlett 10186 (Mi, W——11913h2), 10610 (Mi); Berlandier 83h (T); Graham & Johnston 384F (Au— 174501) ; LeSueur 1,02 (Au); Edw, Palmer 8 (N), 82 (Ca--145769, W— 572312), 103 (Gm); Rozynski 13 (Ca—l69826, Mi); Stanford, Lauber, & Taylor 217ha (N), 2252 (N (N, W—221673l); He W. Viereck 867 (W— (w— 1687429). VERBENA CANIUENSIS Holdenke, Phytologia : 128--129. 1952, cine ie be Moldenke, Phytologia 4: 121 & 128-129. 1952; he 7, & & 12 (3997) and 12: 17. 1958; he Pee Ind, Kew, Suppl. 12: 149. 1959; Angely, Fl. Paran. 1 #3 (19608 and 17: 6. : creeping herb; ee Care tetragonal, rooting at cm cul. - bove, sparsely strigillose; leaf-blades rather firmly chartaceous, rather uniformly green on both surfaces, ovate-subrotund, 1—2. cn, and wide, acute at the base or * slightly prolonged into the petiole, rounded at the apex, rather coarsely dentate along 3 illose with whitish ascending re like ches on the stems, pe~ 1963 Moldenke, Monograph of Verbena 4,83 duncles, and leaf-blades; rachis very Slender, strigillose like the peduncles; bractiets lanceolate, about 2.5 mm. long, equaling or slightly shorter than the calyx, or only 1.5 mm. long and much shorter than the calyx, acuminate at the apex, sparsely strigil- lose; calyx cylindric, 2--3 mm. long, 5-ribbed, scarious between the ribs, ng ects with ascending whitish hairs on the ribs, 5-apiculate, he lowermost ones often ipl ecading’ corolla the limb about 3 MM» wide. The type of this interesting species was collected by Gert eat ol (no, 2560) in "terreno umido da mata ciliar", Rio anit, Palmeira, Parané, Brazil, on November 11, 1951, and is de- posttea in the Britton Herbarium at the New York Botanical Garden. th is in October and November, Thus far only he herbarium specimens, including the type, have been examined by Citations: BRAZIL: Parand&: Hatschbach h 2566 (N--type), 3374 (Z). VERBENA CAROLINA L., Syst. Nat., ed. 10, 2: 852. Syno! f in Ray, Hist. Plant. 3: Suppl. App. 249. 170). Verbena carolinensis, melissae folio aspero Dill., Hort. Eltham. 07, pl. 301, fig. 368. 1732. Verbena caroliniana Murr. in L., Syst. Veg., ed. 13, 62. 1774 [not V. caroliniana Hort., 1847, nor L. ex Raf., 1825, nor Medic., me nor Michx., 1803, nor (Walt.) Michx., 1960]. Verbe- ene ee ent ooo Syst. aoe ed. 13, 62, in syn. 177]. corer carolini ane Willd., Sp. 119. a Eaton & pgs "N. Am. Bot. «8, &10. 180. pert carolin phn serrato, rei apud Poir. in Lam., = oh Bot. 5: 548, in syn. os Verbena tetrandra, spicis filiformibus; foliis indivisis, lanceo- latis, serratis, obtusiusculis, subsessilibus L. apud Poir. in Ene Méth. Bot. 6: 548, in syn. 1608, Verbena polystachya H.3.K., Nov. Gen. & Sp. Pl., ed. quart., 2: 27). 1618 [not V. polystachya Jep- son, 197]. Verbena biserrata H.3.K., Nov. Gen. & Sp. Pl., ed. quart., 2: 275. 1818. Verbena veronicaefolia H.3.K., Nov. Gen. & Sp. Pl., ed. quart., 2: 275. 1818 [not V. veronicaefolia J. Sn., 185]. Verbena vancnssiitol te Humb. ex Spreng. in L., Syst. Veg., ed. 16, 2: 748. 1825. Verbena caroliniana L. apud Hook, & Arn., Bot. Beech, Voy. 153. 1833. 1833. Verbena tetrandra, spicis filif., fol. indivisis lanceol, serratis obtusiusc. subsessilib. L. apud eel ieieemsimnantnemmenmmennamnanndlitlieneaanenaaeenaenena melissae fol. aspero Dill. apt it. E. Richter, hter, Cod. B 35, in syn, 1835. Verbena caroliniana, fol. integro serr. icatiic 48), PHYTOLOGIA Vol. 8, no. 9 Ray apud H. E. Richter, Cod. Bot. Lim. 35, in syn. 1835. Verbena carolinensis, meliss. fol. asp. Dill. apud Peterm., Cod. Bove Oo Linn. Ind. Alph. 196, in syn. 180. Verbena caroliniana, fol. integro etc. Ray apud Petern., Cod. Bot. Linn. Ind. Alph. "196, in syn. 180. Verbena mollis Mart. & Gal., Bull. Acad. Brux >. ae: (2): 323. 18) [not V. mollis ollis Raf., 1832]. Verbena paucifolia tarts & Gal., Bull. Acad. Brux. 11 (2): 32h. 18h. Ve Verbena veronicaefolia Humb, & Kunth apud Benth., Bot. Voy. Sulphur yr 152. 186. Verbena caroliniana Spreng. apud Schau. in A. Dc., Prodr. 11: 56, in § in syn. 1847. Verbena aes etc. Dill. apud Schau. in A. DC., Prodr. ll: 546, in . 1847. Verbena biserrata Kunth apud sohan. inA. DC., Prodr, 11: et in syn. 187. Verbena polystachya Kunth apud Schau, in A, DC., Prodr, 11: 546, in syn. 187. Verbe- na veronicaefolia Kunth apud Schau. in A. DC., Prodr. 11: 56, in syn. 15)7, Verbena pauciflora Walp., Repert. 6s 687. 1847. Verbe- na paucifolia Turcz., Bull. Soc. Nat. Mosc. 36 (2): 196. 1863. Verbena caroliniana f, polystachya Loes. in Fedde, Repert 9: 362. Ion. V Verbena caroliniana var. far. polystachya Loes. in Fedde, Repert. 9: 362. 1911. Verbena pauciflora Mart. & Gal. ex Moldenke, Prelim Alph, List Invalid Names 47, in syn. 1940. Verbena carolinensi Dill. ex Moldenke, Suppl. List Invalid Names 6, in syn. 19)1 [not V. carolinensis Gnel., 1959, nor Small, 1933, es a ie 1959, nor (Walt.) J. F. Gmel., 1903, nor (Walt.) Small, 192]. Verbena caroliniana f, polystachya (Kunth) Loes. ex Moldenke, Spates List Invalid Names 6, in syn. 1942. Verbena caroliniana stachya (Kunth) Loes, ex Moldenke, Alph. List Invalid Names —~ in syn. syn. 192, Verbena carolinea L. ex Moldenke, Résumé Suppl. 2: il, in syn. 1960. Ver’ Verbena caroliniana, folio integro serrato seabro Ray apud Moldenke, Résumé Suppl. 2: 2: 11, in syn. 1960. __ Bibliography: oad Hist. Plant. 3: Suppl. App. 29. 1705 Dill., Hort. Eltham. 07, pl. 301, figs a 1732; Le, Syst. Nate, ed. 10, 2: 6 117595 Ue, Spe Pl., eds 2,29. 1762; Mill., Gard. “s 873 & a7. 1 21; Bpbeiees mL , syst. Tees, ed, 16, 2: 7,8. 1825; Cham. & Schlecht, Linnaea 5: $8, 1830; Sweet, Hort. Brit., ed. 2, 18. ete Hocity el Bot. Beech, Voy « 156. 1833; H. E. Richter, Cod . Linn, 35. 1835; Hook. & Arn., Bot. Beech. Voy. 305. 18385. nt & Wright, Ne Am. Bot., ed. 8, 70. 180; Peterm., Cod 1963 Moldenke, Monograph of Verbena 485 Bots Linny ind; en te 196. es re: & Arn., Bot. Beech. Voy. 484, 1841; Mart. & Gal., Bull. d. Brux, 11 (2): 32h. 18k); Prodr. 11: 56--57. 1847; Turcz., Bull. Soc. Nat. Mosc wae (2): 196. 1863; Pickering, Chronol. Hist. Pl. 962 & 1220. 18595 Ss. Wats., Proc. Am. Acad, Sci. 18: 135. 1883; Briq. in Prantl, Nat. Pflanzenfam. (3a): 148. 189); Hook. f. & axes Ind. Kew. 2: 1173--1180. wena Loes, in Fedde Fone ~— — 9: 362. 1911; Stapf, Ind. Lond. 6: 429. 19 31; P Gard. 20: 243, 24527, 259 9, rk 213, 2, 6 te 5. "1335 a D. Merr. Journ. rno x 9: a Bree bs ; Moldenk Alph, List Invalid Tene & 49. "190; pasaindes & Standl, Fl. Salvad., ed. 2, hens folder, Suppl. List Invalid Names 8. 19)1; iors Alph. List Invalid Names hS—l9 & 51. 19h2; Moldenke Ab ban dell, Fl. ee 3 (1): 2h. 1942; Howard s. Gentry, Rio Mayo Pl. [Carnegie Inst. Wash. Publ. 527:] 222. aghes Moldenke, Known Geogr. Distrib. Verbenac., [ed. 1], 11, Pons 18, 20=-22, & 101. 192; Ramfrez Cantii, Anal. Inst. pets o Lys 406. 1943; Moldenke, i - rh & a 19453 We Co sep flr ce ort idl exe » 3665 370-2372, 37 78, & L7l. 19595 aa. Résuné Suppl. 2: z 10--13, 1960; Lewis & Oliver, Am. Journ, Bot. a 1961; Ps Résumé Suppl. Jt 37, ty.& ue (1962), 7 tay 39; & 20 (1962), and 5: 6. Aled a! bie, ant Eltham. pl. 301, fig. 388. eed, Dill., Hort. Eltham, Add. Denom. Linn. pl. 301, fig. 388. Mirbel, Ann, Mus. Hist. Nat. Paris 15: pl. ( (Pigs 5. B10; Lewis & Oliver, Am. Journ. Bot. 48: 60. 1961. On % 1,86 PHETOLOGIA Vol. 8, no. 9 Coarse, erect, aromatic herb, mostly annual or biennial, some- times perennial, slender, 30 cm. to 1.55 m. tall, usually 0 i usually tary, sometimes s rom the crown of a thick rootstock, tetragonal, sulcate, branched, rom escent to more or less villous-hirsute or hirsute-hispid; princi- es 1 long, sometimes 3 times as long as leaves; leaves numerous but often remote, arying from short-petiolate to subsessile or sessile; leaf-blades vary f eolate or lanceolate-oblong vate-lanceolate or a3 wide, acute or obtusish at the apex, cuneate narrowed to the base, coarsely and irregularly serrate or biserrate to crenate- serrate along the margins, lineate-subrugose, varying from ap- pressed-pilose or appressed—pubescent to strigose and somewhat pustulate mostly hirsute-strigose (especially along the , lavender, light-lavender, pale- or very pale lavender to blue, pale-blue, light~ or very light-blue, or even ocarp; cocci trigonous, dark-brown, barely 1.5 mm. long, the dorsal surface convex, faintly striate and obsoletely reticulate, the commissural faces meeting at right angles, almost smooth; chromosome number: n = 7, 2n = lh. The species is a very variable one and has been much confused n given forms which are not sufficiently constant to merit nomenclatural recognition. The type of V. biserrata was collected by Friedrich Heinrich Alexander von Humboldt and Aimé Jacques Alexandre Bon- pland near the city of Mexico, Federal District, Mexico, and is de- posited in the herbarium of the Muséum National d'Histoure Natur- elle at Paris. The type of V. veronicaefolia was collected by the same distinguished explorers at Moran, Hidalgo, and that of V. polystachya on the slope of the volcano Xorullo, Michoac4n, and are also deposited in the Paris herbarium. The type of V. mollis was gathered by Henri Guillaume Galeotti (no. 737) in three places: (1) in wet places in the Cerro de San Felipe, Guanajuato, (2) near Oaxaca, nd (3) in the mountains of Yavezia and Castrasana, at 7500 1963 Moldenke, Monograph of Verbena 87 to 8000 feet altitude, described as having red flowers from May to September. Martens & Galeotti say of it "Affinis Verbenae nas at the German colony of Mirador, Vera Cruz, at 3000 feet al- titude, and is said to have bluish flowers in August. The authors of the name say "Affinis Verbenae polystachiae ind oe foliis cesarean caule minus folioso praesertim diver s of V. carolina have been abundantly sa taibaseeee and aueernten under the names of V. barbata Steud., V. ciliata Benth., V. ehrenbergiana Schau., V. elegans Humb. is Voldenke, Ma os halei Small, V. hastata L., Vv. littoralis H.B.K., Vv. menthaefolia Benth., "y, paniculatae affine", V. recta H.B.K., ¥. scab: , V. urticaefolia L., V. urticifolia L., Stylodon car- neus (Medic.) “Moldenke, and even ‘Ipomoea SD. ~ Qn the other hand, the P. C. Standley 21331, Steyermark whe Tucker 1307, and S. Se White 3715, distributed as typical ae Olina, are V. carolina f, albiflora Moldenke; G. L. Fisher Hee and Tietmann 11336 are V. ehrenbergiana Schau.; | Rose, Pachano, | & Rose 22851 & 22939 and Rose & Rose 22730 are V. glabrata H.B.K.; M. Bang 1058 is V. gracilescens (Cham. am.) | Herter; T. S. Erenceeee S.n. (Julian, June 18, 189] and Doten 21 are V. rT, lasiostachys var. septentrionalis lioldenke Arsene s.n. yao etn 925, and F. W. Pennell 1770 are V. litoralis H.B.K.; H. H. Rusby Rusby 6121 & sen. [Clarke's ” Valley, Aug. ve 1883] are V. ae Heller; Conzatti 207, Pringle 9529, H. E. Seaton m 7, and S. S. White 410) are Vv. ~menthaefolia Benth.; E. W. Nelson 1105, “Pefialo- ¥; recta HeBeKe; a fda le Blankinship s.n. fperkeley, tals 22, 1891) is vy. V. robusta Greene; 3 Ne L. Britton 1527, Britton, Britton, & Shafer 292, Bi 292, Gritton & Hollick 2704, Britton & Wilson 5771, Brown & Britton 373, W. Harris 9937, Leén 685 & 685 & 1343, and Edw. Palmer 10h0 are V, scabra Vahl; and W. C. C. Coker s.n, [July 8, 8, 1909] is Vv. u urticifolia var. letocarpa Perry & & Fernald, while ths Herb. Hort. Bot. Genev. s.n. [18 Aug. 1826] is in part V. recta and in part | Vv. ur urticifolia var. leiocarpa. The Seler & Seler r 43k7 dis- tributed as "Verbena spec. , aff. V. caro aroliniana L." and l194 as "Vv. caroliniana L. form. vel var. polystachya (Kunth) Loes." are Vv. longifolia Mart. & Gal. The GL. Fisher 320 and Pringle 769 4 dis- tributed as "Verbena aff. V. ~ polystachya H.B.K." are are V. recta HBK. Concerning the closely related V. ehrenbergiana Schauer says: "Species per se quidem satis insignis, sed quoad habitum fere V. carolinianae hirsutioris formam Schizophyllam exhibens." Perry disti shes the regecols reished ceeedgs of this affinity by means of the following 4,88 PHYTOLOGIA Vol. 8, no. 9 1. — very slender, elongated anc graceful, usually with re- te fruits 2. te aves l- or 2=pinnatifid, }-5-cleft, or deeply incised. 3. Corolla-limb not less than 3 mn . broad; cocci 1.5 mm. or more in leng . Schizocarp a little anne than broad. 5 Bnb.og escence usually densely glandular and somewhat offi scid-pubes oot ip Sand eoeeVe O LC inalis. Sa. inflorescence sparsely, if at all, sepennel es and rigillos Ve enthaefolia. ha. Slicsere about twice as long as br — 6,-Leaf-blades diverse in outline, the basal ones incis- ed-dentate, the middle ones 4 or 2=pinnatifid, the upper ones * sparingly toothed or entire; bractlets half as long as the calyx during anthesis.V. halei. 6a. eee ee all similar in outline; bractlets about s long as the calyx during anthe Sis...Ve riparia. 3a. corshla-Linb carcely more than 1 mm. se cocci r less in leng Aes ciabtine siete lle ehrenbergiana. 2a. “ar serrate sed. .- Leaves subsessile or snort patlolate with an attenuate a 8. Stems J gobo x ee: leaves mostly crenate-serrate; calyx hi spidulou eeeee ee «Ve caro olina. 8a. tty glabrous ; cueeh acutely serrate; calyx practic-— ally g labrous TO) BREN! 9, mgifolia. Ta. eee obviously petiolate, with rounded-cuneate base. 9. Fruiting-calyx spreading; calyx-lobes conn yeohs stig- aes surface subtended by 2 sterile style ere scabra. 9a. pita -calyx ascending; calyx-lobes not connivent; stigmatic surface subtended by a eerete sterile style-lobe ‘ urticifolia. urticitols la. oeree thicker or densely flowered, usually eet contiguous 10. so Ne oe pert or very short—pedunculate, fo orming a2 id ngested inflorescence, not strict; corolla-limb 2 mm, road oeVe recta. 10a. Spikes or ae geod sooner not congested, strict; cor= Limb broad , hastata. Schauer (187) Sch ¥. recta to synonymy under V. carolina, he is followed in this by Hooker & Jackson (1895), but uct ria species are amply deatease: as has been pointed out by P in her excellent monograph. s Sathens worthy of note here that V. caroliana L. is Ve pores Moldenke; V. carolineana Hort. is V v. officinalis 1; Ve carolina var. glabra a Hultén is V. longifolia a Mart. & Gal; Ve caroliniana f, recta and var. recta (Kunth) Loes. are V. recta 1963 Moldenke, Monograph of Verbena 489 «Ke; V. veronicaefolia J. Sm. is V. peruviana (L.) Britton; V. or er Ste Jepson is V. scabra Vahl; V. mollis Raf. is V. stricta Vent.; and V. carolinense (1 jalte) Gmel., Ve carolinianun L Le, ne TE caroliniensis (Walt.) Je Fe Gmel., Ve carolinensis Gmel., Small, Walt., (Walt.) J. F. Gmel., and (Walt. ) Small, and V. caroliniana Hort., L. ex Raf., Medic., Michx., and (walt.) Michx. are all Stylodon carneus (ledic.) Moldenke. The Verbena caroliniana, erecta, foliis ‘oliis oblongo-obovalibus, obtusis; spicis filiformibus, iopinsinis distinctifloris Michx. cited by Poiret in Lam., En- cycl. Méth. Bot. 8: 5,8, in syn. (1808) as a synonym of V. caro- lina, actually belongs in the synonymy of Stylodon carneus. ~The bibliographic citation for the rg synonym onym given in i. eaet synonymy is sometimes given as page "),06", apparently in Te ° Iybrid of V. carolina L. with V. hastata L. is known as xV. matritensis, which see nets carolina fihabites rishi or volcanic soil, brown volcanic ac- idic. soil, moist sand, and moist brownish sandy-loam, rich clay- ish loam along lake shores, among granitic rocks, in clay or red meters. It has been found in anthesis in every month of the year, and in fruit from April to February. It pro obably also fruits throughout the Sosy Peflalosa describes it as a "weed between and he and Hoggett report it as "very common in yards". Liac n Standley to be ta weed in vacant lots" and "common on open weedy mt i cker, D n e plant in waste places" in Honduras. Mueller & Mueller asse that it is "very abundant on waste ground" in Nuevo Len, It is to be a "street weed" | in Honduras and "common in waste slaces" in 4,90 PHYTOLOGIA Vol. 8, no. 9 sun of wooded slopes and barrancas and abundant in sandy loam a- long roadsides in open sun on precipitous mountain slopes; Muller say. 4 it is "abundant throughout open pine forests flowers white to light-blue"; McVaugh & Koelz found it "weedy a- forest road on steep slopes ophytic est - r near sum mits of barrancas in the pine zone"; Wilbur & Wilbur aver that it is "very common in bunchgrass meadow in mountains"; Leavenworth & Hoogstraal discovered it to be "very common in yard behind house"; and Fearing & Thompson found it in black soil of volcanic origin 8. The specific name is sometimes written with a capital initial letter. T. H. Kearney, on the label of Harrison, Kearney, & Hast- 6y), says "the distinction between this meet oe and v. = ticaefolia seems slight." The tallest specimen yet recorded is Hartman 94, said by the collector to have been l--5 feet tall. The species has been in cultivation appar eine 4 = sphere or Common and vernacul es recorded t+ are vervain", "kasar&é §ijékame" (which m mane H main bracts very broadly nd “pleulate, sacar half as long as the branches and equ the lower spikes » punctate toward apex; branches sg ted pony mostly of 3 subfasciculate spikes with the 2 the terminal; te sepals, sharply carinate, coriaceous, nerved, green; opens na Racto 5¢ sepals free, elliptic, acute, ar mm, long, coriaceous, nerved, the pos- terior Repeeses petals eh = longer than the sepals, pale yel~ iow, bearing nh es stamens included. Pl. II, fig. 6: Apex of leaf x *€ 17% re Ne 4 of inflorescence x 1/2; fig. 8: Sepal x 1 e in t pre es National Bestexse, collected on tree, vici- nity of Cebolleta, Department o Magdalena , Colombia, altitude 2400 m., Faienien 1, 1959, by :. Romero~Castaneda (No. 7199). PERU POSTERELLA L. B. Smith Schidospermm Griseb. in Lechler, Berberid. Amer. Austr. 56. romans” L«Be Smith, Phytologia 7:171. 1960. iling a series of type photographs of Liliaceae, I was aoe by the familiar appearance of ye igen poe Schidospermum Chlorophytum. The hlo name must be aetaaeet as follows: FOSTERELLA SCHIDOSPERMA (Baker) L. B. Smith, comb, nov Schidospermim sanseviera Griseb, in Lechler, Berberid. " Amer. 1963 Smith, Notes on Bromeliaceae 501 Austr. 56. 1857. Nomen. athe schidospermum Baker, Journ. Linn. Soc. Bot. 15:326. 18 Cottendorfia rusbyi Baker, Bull. Torrey Bot. Club 29:697. 1902. Not Lin rusbyi Mez, 1901. Lindmania weberbaueri Mez, Fedde Rep. Spec. Nov. 12:417. 1913. Fosterella weberbaveri (Mez) L. B. Smith, Phytologia 7:172. 1960. Junin: Chanchamayo Valley, 1500 m., Schunke 273 ("). a Merced, alt. 700 my, ak tng 5352 (F, GH). Rio Paucar Valley, near Perené Bridge, alt. 700 m., Killip & A. C. eens 25326 (NY, US). Vitoc, 13 km. from San Ramon, Soukup 2446 (US). AYACUCHO: Prov. Huanta: By the Rfo Pieni, Weberbauer 5635, (B, type of Lindmania weberbaueri Mez; F photo no. 11460). San Gaban, July 1854, Lechler 2382 (G, type number of ccthieeadiind mum sanseviera Griseb. “and of Chlorophytum schidospermum Baker; F photo no. 25148). BOLIVIA: La Paz: Yungas, alt. 2000 m., Rusby 2541 (NY, type of Cottendorfia rusbyi Baker; US). ae COMMIXTA L. B. Smith, sp. n A P. lasiopoda ahs B, Smith, a “valde affinis, spicis subses- silibus differt. Flowering 6 dm. high; leaves 2 dm. long; sheaths ample, dark castaneous, forming a bulb 5 cm. in diameter; blades very narro i > 15 mm, wide at base, densely cinereous-lepidote beneath, glabrous above, laxly serrate with slender sprea brown spines 4, mm, long; scape slender, soon glabrous; str flowers erect; pedicels obconic, 3 mm. long, inconspicuous; se- pals linear, sag ee nearly 4 cm. pens 5 4 mm, wide, ecarinate; petals strongly contorted and purple when dry. “79 Hi, figs: 9: Branch of ae eto: x ane fig. 10: Sepal x des in the U. S. National Herbarium, oe on dry open de among rocks, fog belt, on trail, 2-6 km. from Oconeque, of Pun m., May 22-25, 1942, by R. D. Metcalf (No. 30591 of the Universi- ty of California, Third Botanical Garden Expedition to the Andes pace et by T. H. Goodspeed). Isotype in the University of Cali- fornia. PUYA LOPEZII L. B. Smith, sp. nov. A P. lanata (H.B.K.) Schult. f. , cui valde — bracteis florigeris api apice contractis minoribusque, sepalis oblongis dif dl Known only from fragments but probably flowering over 1 m. 502 PST OLOG Ix Vol. 8, no. 9 high: leaves rosulate, ca. 8 dm. long; sheaths suborbicular, 10 em. long, the basal part yellow and glabrous, the apical dark castaneous and covered with rie al white appressed scales; blades narrowly triangular, 5 cm. wide, covered on both sides with ap- pressed white scales, laxly poet Se ‘vith flat brown ns inate spines 8 mm. long; upper part of scape 8 mm. in diameter, soon glabrous; upper scape-bracts ovate with a long narrowly triangu- lar apex, exceeding the internodes but expos most of the scape, entire, rugose when dry as if formerly fleshy; inflores- rru 5 cence simple, subdensely cylindric, int over dm. long, densely and finely white-stellate; floral bracts broad- ly ovate, contracted to a narrowly triangular ’ 8 ecarinate; petals 40 mm. long, yellowish; stamens included, Pl. II, fig. 11: Floral bract and flower x 1/2; fig. 12: Sepal x 1. Type in the U. S. National Herbarium, collected on dry slopes, Huaylillas, Province of Pataz, Department of La Libertad, Peru, sneer a m,, May 24, 1961, by A. Lopez and A. Sagdstegui oo. PUYA SAGASTEGUII L. B , Sp. AP. lanata (H.B.K. co ociedlt im ey ue affinis, bracteis flori- geris | apice co contractis, sepalis angustis minoribusque, petalis purpureis diffe: — Known om fragments but er over 1 m. high; leave over 8 dm. ~ dap blades narrowly triangular, 25 mm, wide, odvanet on both sides with white appressed scales, laxly serrate with slender brown uncinate spinea 7 mm. long; scape unknown; inflo- rescence simple, subdensely many-flowered, densely and finely pale~lanate, becoming subglabrous, the base unknown; flo mm. long; sepals linear-1 lanceolate, 23 mm. long, 3.5 mm. wide ecarinate; petals with lateral folds, contorted after, anthesis. Ly Ii, fig. 13: floral bract and flower x 1/2; fig. 14: Sepal Type in the U. S, National Herbarium, collected on dry slope Huay. Province of Pataz, Department of La Libertad, Peru, ape m., May 24, 1961, by A. Lopez and A. Sagaste TILLANDSIA LOPEZII L. B, Smith, Sp. nov. AI. kuntzeana Mez, cui affinis, laminis foliorum subglabris, ~ subdigitata differt. rent ote, dark purple; blades ligulate-triangular, acuminate, 45 mm. wide, flat, green with purple splotches, very obscurely lepidote beneath; scape erect, 7 mm, thick » labrous; scape-bracts erect, imbricate, the lower foliaceous but reduced, the upper ovate, apiculate, lustrous; inflorescence subdigitate from 6 spikes; 1963 Smith, Notes on Bromeliaceae 503 primary bracts a » broadly ovate, apiculate, to 4 cm. — spikes subsessile, narrowly lanceolate, acute, 7-9 cm. long, 2 cm. wide, slightly odeuinaahs, densely &~-9-flowered; floral bracts broadly elliptic, acute, equaling or exceeding the sepals, sharply carinate, except for the apex and narrow margins dr nearly black and lustrous, coriaceous, glabrous; sepals elliptic, apiculate, 30 mm. long, glabrous outside, the posterior carinate, connate for 3 mm. Pl. II, fig. 15: Spike x 1/2; fig. 16: Poste- rior sepals x l, in the U. S. National Herbarium, collected on rocks, road to Cujibamba, Province of Bolivar, Department of La Libertad, Pe- ru, altitude 3300 m., June 1, 1960, by A. Lopez and A. Sagastegui (No. 3307). ILLANDSIA SAGASTEGUII L. B. Smith, sp. nov AT. plat lla Mez, cui affinis, foliorum laminis acumina- tis, sepalis bracteas florigeras superantits differt. Flowering to 2 m. high; leaves over 1 m. long; sheaths oblong, scarcely distinguishable from the blades except by the brownish color, over 12 cm. wide, densely vestite with minute brown ap- pressed scales; blades narrowly triangular, acuminate, flat, green, obscurely lepidote eet glabrous above; scape not seen; inflorescence — ipinnate, essentially glabrous, glau- cous; lower primary bracts naiud tanner nearly eq the axillary branches, the upper primary bracts caudate from a broad- ly ovate base, shorter than the one-bracteate sterile base of the branches; branches of 4 subdigitate spikes, ascending, to 25 cm. long; spikes linear, complanate, to 15 cm. long including the the sepals at anthesis, ecarinate except near the ap y in- curved apex, coarsely and faintly sulcate; a inconspicu- ous, 2 mm, long; sepals free, oblong, su - long, sub- coriaceous, glabrous outside, minutely hepiaees pened » the pos~ terior carinate; petal-blades elliptic, 18 mm. long, violet; sta- mens included. Pl. II, fig. 17: Apex of leaf x 1/2; fig. 18: grag te nti fig. 19: Sepal x 1. Type in the U. S. National Herbarium, collected on —— » near Santiago, Gusmango, Province of Contumaza, Department Cajamar~ ca, ae altitude 2600 m., August 3, 1960, by A. aa tainaas (No. 3390 — oe pri L. B. Smith, sp. AT. era Mez, cui affinis, ayia. laxiore, rhachi haud obtecta, yt liberis, petalis albis differt. C escent; stem over 15 cm. long, 8 ug in diameter; leaves densely imbricate, recurved-spreading, 20 cm. long, densely ap- pressed-lepidote; sheaths elliptic, ca. 5 cm. long, merging with the blades, ferruginous; blades very narro riangular, long- acuminate, 20 mm, wide at base; scape distinct, 15 am. long; 50h PHYTOLOGIA Vol. 8, no. 9 scape-bracts erect, densely imbricate, densely lepidote, the low er subfoliaceous but much reduced, the upper broadly elliptic and api te; inflorescence simple; broadly maneenaite acute, 11 cm. long, 4 cm, wide, strongly complanate; rhachis flexuous, winged, soon glabrous; floral bracts erect, many times longer e me _ narro than the interno at base and exposing most of the rhachis, obovate, > 4 em. long, minutely incurved at aatihe ecarinate, densely. Iepidote ; evidently fleshy because rugose drying; pedicels 2 long; sepals free, elliptic, broadly ss e and spioulate ee 28 mm. se subcoriaceous, subdensely lepidote; als 5 mino de -_ Quishuas, Province of Bolivar, Department of La Li- Peru, altitude 3000 n., ie 31, 1960, by A. Lopez and A. Sagistegui (No. 3296). TESEA CEREICOLA (Mez) L. B. Smith, Phytologia 6:194. 1958. Tillandsia cereicola Mez in Fedde, Rep. Nov. Spec. 3:34. 1906. Vriesea lopezii L. B. Smith, Phytologia 4:218, pl. 2, figs. 8-10. eee ty" Vriesea lopezii was described before I realized that Tilland- sia cereicola was a Urieses. Now I can find no good distinction between the two species BOLIVIA epee. MICROLEPIS L. B. Smith, sp A B. formosa Ule, cui affinis, Ferien i laxa elongata, sepalis minoribus differt. own only from fragnents but apparently the habit typical of Billbergia gubgemms Helicodea; leaves 50-75 cm, long, densely eee on both sides, white-banded ‘beneath; 3; sheaths ellipt r 26 cm. long; blades ligulate, subacute and apicu- late, 5-7 = ate laxly serrate with flat black cc ra teeth 1.5-5 mm. long; sc ape slender, decurved, finely pale-lepidote; sdape-bracts elliptic, acute, to 13 cm. long, the lower erect, eae broadly ovate, apiculate, the lowest almost e ovary, the others much shorter; flowers sessile; i subequal, lance-cblong, attenuate, 20-24 mm. long, soon glabrous except at base; petals linear, over 14 cm, long, greenish when dry, recoil- ing in a tight spiral at anthesis, bearing 2 truncate dentate scales at base; ovary ellipsoid, sulcate with dark lines against the white indument., 9 mm. high exclusive of the 3 mm, short- cylindric epigynous tube. Pl. III, fig. 3: Floral bract and a fig. 4: Sepal x 1. Type in the Gray Herbarium of Harvard vn 3 th » collected on tree, it ter (about 10 km. south of Mapiri), Province of La~ recaja, Department of La Paz, Bolivia, saad Sade 850-950 Mey October nr 15, 1939, *by B. A. Krukoff (No. 11158). 1963 Smith, Notes on Bromeliaceae 505 aoa CARDENASII L. B. Smith, sp. nov. P. palmeri S. Watson, cui valde affinis, rhizomatibus aucta, Pn integerrimis differt. Reproducing vegetatively by slender scaly rhizomes, flowering cm. high; leaves forming bulbous rosettes that soon appear rhi long, strongly nerved, castaneous toward base; blades dimorphic, some persistent, reduced, broadly triangular, thin, stramineous, others deciduous along a straight transverse line, linear, atten- uate, 10 cm. long (probably not fully developed), 2 mm, wide, fasciculate; scape very slender, white-flocculose especially to- rd base; scape-bracts erect, the lower imbricate, subfoliaceous and with deciduous blades, the upper narrowly triangular, shorter than the internodes; inflorescence secundly racemose, r sparsely white-flocculose; floral bracts narrowly tr. jangular, ex= long ceeding the slender 6-8 m. pedicels; flowers suberect; se- pals oblong, acute, 18 mm. long, carinate at base; petals linear- - rhpionsb » rounded-api mm, long, naked; clu- te, 4 3 st ded; ovary 4/5 superior; seeds caudate-appendaged. Pl. III, fig. 5: Leaf-blade x “1/23 fig. 6: Inflorescence x 1/2; fig. 7: Sepal x Type in the U. S. National Herbarium, collected in thicket on wet rocky slopes, near Cuesta de Monos, Department of Santa Cruz, Bolivia, altitude 1000 m., November 1950, by M. Cardenas (No. rn ° Originally I confused this specimen with Pitcairnia rigida Mez, giving rise to the dubious citation of that species in R. C. pty Catalogue of the Ferns and Flowering Plants of Bolivia pe ILLANDSIA STREPTOCARPA Baker, Journ. Bot. 25:2h1. Tillandsia soratensis Baker, Journ. a 253235. i687. Tillandsia condensata Baker, Handb. =e rat 1889. Tillandsia apoloensis Rusby, Bull. N. Y. Bot. Gard. 6:489. ares 18 BOLIVIA: La Paz: Apolo, alt. 1440 m., September 31, 1902,:R. S. Williams 1483 (NY, type a t. apolo&nsis). Cochabamba: Tuna- ri, alt. 1300 m., Kuntze s, n. “(NY, type of T. soratensis). ta "cruz: West Velasco, alt. 200 m., Kuntze s. n. (NY, type of T. condensata). On attempting to differentiate the above species in a key, the characters previously used proved to be quite trivial ge unre] i- able when compared with those of the ag of Tillamdsia strepto- carpa (Balansa 615, from Paraguay, & ox TILLANDSIA SUBTROPICALIS L. B. Smith, sp. nov. AT. kuntzeana Mez, cui aff finis, foliorum laminis supra glabris, scapi bracteis glabris, bracteis florigeris haud incur- vatis, flo: ribus mlto minoribus emless, flower 3 dm. high; pit 15 or more in a funnel- form rosette, 20-23 cm. long; sheaths broadly elliptic, 6-7 — 506 PHYTOLOGIA Vol. 8, no. 9 long, dark castaneous, densely and minutely appressed-lepidote; blades ligulate with a long attenuate apex, 15-20 mm. wide, covered beneath with fine appressed cinereous scales, glabrous ca concealing the scape, the lowest subfoliaceous, the others ellip- tic, glabrous and lustrous except for the small narrow blade; in- florescence laxly bipinnate from 5-7 spikes, glabrous; primary bracts ovate, about half as long as the spikes; spikes subspread= ing, subsessile, lanceolate, acute, 45 mm. long, 12 m. wide, strongly complanate, dense; floral bracts imbricate and wholly conce. eabie the rhachis, ovate, acute, 18 mm, long, sharply cari- nate with a straight apex, co oriaceous >» even, lustrous; sepals lanceolate, acute, 15 mm, long, the posterior carinate, 2/3 con- nate. Pl. III, fig. 8: Apex of leat x 1/2; fig. 9: Inflorescence x a) fig. 10: Posterior sepals eU. S. National Bicbarai, S teas in the sub os ae ciation, at Hacienda Casana on the road to Tipuani, De- partment of La Paz, Bolivia, 1400 meters altitude, August 5, 1922, by Os Buchtien (No. 7185). : Same as type, October 15, 1922, Buchtien 7187 (US). BRAZIL AECHMEA aes (L.) Griseb. var. AEQUALIS Smith & Reitz, var. A i " mdicaulis scapi bracteis fere aequalibus, supremis non aggregatis differt. Scape-bracts small, nearly equal, not massed below the inflo- rescence; lowest floral bracts elliptic, attenuate, seg RE the ovaries. Pl. III, fig. 11: Scape and inflorescence x Type the U. S, National Herbarium, collected in STE State of Espirito Santo, Brazil, by Alvim Seidel (No. 6-14). QUESNELIA SEIDELIANA Smith & Reitz, sp. nov. A Q. lateralis Wawra, cui valde ar tsnte; scapi bracteis valde remotis, inflorescentia floribusque parvis differt. Stenless, flowering 5 dm. high; | eaves few in a fasciculate rosette, 35-40 cm, long, covered wi theook closely aera sed dark-centered scales; sheaths wtispete » nearly 20 cm. long, tinged with dark purple especially otopr blades rigiite, sessile; sepals oblanceolate, acute, 15 mm. long, connate for 3 mn., white-lepidote; petals "sky blue", 25 mm. long, bearing 2 fimbriate scales at base, the blades elliptic, obtuse; stamens included; ovary ellipsoid. Pl. III, fig. 12: Apex of leaf x 1/25 1963 Smith, Notes on Bromeliaceae 507 fig. 13: Inflorescence and upper scape x 1/2; fig. rg Sepal x l. in the U. S. National Herbarium, collected the State of Rio de Janeiro, Brazil, by Alvim Seidel (No. saan. VRIESEA BARILLETII E. Morr. Belg. Hortic. 33: si pl. 3. 1883. BRAZIL: Cultivated in the botanical gard f Liége, 1883, and supposedly c g from Ecuador, E. Morren (LG, Pog oh GH). Central Brazil, lyin Seidel (US). Alvim Seidel, in “dn his search for new and rare species of orna- mental b romeliads, has discovered aie barilletii growing na- turally in ak spoken Brazil. This is not altogether surprising since all its near relatives are Brazilian and the record from Ecuador has long been regarded with suspicion. SYNOPSIS OF AMERICAN VELLOZIACEAE SUPPLEMENT II Lyman B. Smith As frequently happens, publication in a new field is followed ¥ offers of more material to study. In this instance the study of the unidentified American Velloziaceae from the herbarium of the Royal Botanic Gardens, Kew, has yielded nine tga epi and pecies,. The Pes wpe here are numbered in conforaity with the an in my original paper in Contributions Hat Sonal. Hertart, volume 35, part 4. The figures are ngs 5 natu- Fal size la. ap oy 0 JA LAEVIS L. B. Smith, sp. nov ay, edoni Woodson, cui affinis, staminibus 6 basi exappen- Genistse a pray browm, lustrous, glabrous, dividing and recurving at apex with age; blades Sed covatetenk linear, attenuate to a rounded re- tuse apex, 6-8 cm. long, 1.5-3. 3 mm. wide, laxly serrulate on the thickened margins and beneath on the keel, glabrous; scape soli- tary, 8 cm. long, very slender, glabrous; ovary subglobose, 4 mm. r. ila aged. Fig. 1: oe fig. 2: ose ke tepal, and stamen BRAZIL: Minas Gerais: Among rocks, Caraga, 1824, Riedel 8. n. Vol. 8, no. 9 P SEot OO. GT & 508 Plate I fig. 4-6: Vriesea ampla; hainesiorum Pig. 1-3: Tillandsia leucolepis; fig. 7-10: Wa 1963 Smith, Notes on Bromeliaceae 509 Plate II Sa , = 4 ee oe, edt, arn OE Oe eee ok SS eee 2S wl Fig. 1, 2: Vriesea leucophylla; fig. 3-5: Tillandsia romeroi; fig. 6-8: Vriesea enae; fig. 9, 10: Puya commixta; fig. 11, 12: P. lopezii; fig. 13, 14: P. sagasteguii; fig. 15, 16: Tillandsia lopezii; fig. 17~19: T. sagasteguii. 510 PHYTOLOGIA Vol. 8, no. 9 Plate III Fig. 1, 2: Tillandsia truxillana; fig. 3, 4: Billbergia microlepis; fig. 5~7: Pitcsirnia cardenasii; fig. 8-10: Tillandsia subtropicalis; fig. 11: Aechmea mdicaulis var. aequalis; fig. 12-1): Quesnelia seideliana. 1963 Smith, American Velloziaceae 51 lla. VELLOZIA CRISPATA L. o Smith, sp. A V. compacta Mart. ex Schult. r. et affinis, floribus folia I or Caudex simple cm. long, terete and 13 mm. in diameter in- cluding the “neg ret leaves very densely imbricate, the com— oe ones 15-25 at the top of the caudex; sheaths ample, ca. 3 em. long, very closely and coarsely nerved, yellow, lustrous, glabrous, the apices split into fibers and recurved in tight spi- rals with age; blades linear, attenuate to a blunt retuse apex, » subden trichomes; ovary obovoid, trigonous, 16-18 mm. long, scabrous on the base and angles; tepals elliptic, 5 cm. long; stamens numer ous, probably included, the phalanges bearing numerous short teeth; style 25 mm. long. Fig. 4 Apex of leaf; fig. 4: Ovary; stam an ens : Rio de Janeiro or Hine Gerais: "Environs de Rio de Janeiro et d'Ouro Preto, 1883-84," Glaziou 15501 (K, type). 20a. VELLOZIA DROSEROIDES L. B. Smith, sp. AY. echinata Goeth. & Henr., cui Pe ats ovario multicostato satin fe) rt. Caudex simple, 3 cm. long, terete and 15 mm. in diameter in- t cluding leaf-sheaths; leaves very densely imbricate, the com- plete ones 10-15 at the top of the caudex; sheaths 2 cm. long, closely nerved except near t » yellow-browm, a gla » With deeply divided into fine st es b linear, attenuate to a t apex, 7- mn, wide, on margins and beneath on one keel, abrous; scape 6-10 cm he; ee Shdek,: oedl co ly veatite with short~stipitate glands; ovary ellipsoid, 6~8 mn. 1 12-costate with the stipitate glands gui ong, 10- irregular 8s but well ptyreees tepals elliptic, 2 om. long. Fig. 6: Apex of leaf; fig. 7: Ov ip ecitas Serra da Lapa, November 1824, Riedel 1053 (K, For We 27>. VELLOZIA VESTITA L. B, Smith, sp. nov. AY. AEE colore L. B. Smith, cui affinis, foliorum vaginis anatis per aetatem grosse fibroso—divisis, laminis angus- Pryde differt. 8 @, 10 cm. long, t e and 3-4 cm. in diameter in- cluding the Leaf—sheaths; palit "a densely imbricate, the com- plete ones about 12 at the top of the caudex; sheaths ample, 25 throu, to a minutely blunt apex, 24 om. long, 14 mm, wide, entire, green lar; ovary subglobose, 10 mm. long, completely covered with very fine stipitate glands; tepals elliptic, 5 cm. long, the outer 512 Pry T0450 6 TA Vol. 8, no. 9 densely glandular; stamens included, 12 or more. Fig. 8: Apex of leaf; fig. 9: Flower. BRAZIL: Minas Gerais: 1892, Glaziou 19935 (K, type). 3la. VELLOZIA METZGERAE L. B. Smith, sp. n AV. pilosa Goeth. & Henr., cui artinis, - “folie dense cinereo- ie » floribus folia subduplo superantibus, ovario densissime glanduloso-piloso differt. Subacaulescent, simple; leaves very densely imbricate, the complete ones 4-5 at the top of the caudex; sheaths ample, 10-15 ed, b mm, long, very closely and coarsely nerv dark brown, sublus- trous, glabrous except the pilose apex, with age splitting into straight coarse fibers; blades linear, 5-7 fe) mn, wide, - thick, vestite especially toward apex with short- sine elacsia} rey globose, 5 mm. pes very densely and bs pilose-glandular 3; tepals en af elliptic, 15-20 mm, long red, glandular at base and the o at apex; wegen Serge style 14 mm. long. Fig. 10: nthe of leaf; fig. BRAZIL: Bg as Gerais: Serra da Lapa, November 1824, "Riedel Se n. (K, typ This s Sables is named in honor of Frau Crista Metzger in r aia of her help in making type-photographs of Pelipathtens available 38a. VELLOZIA BREVIFOLIA Seubert var. ANGUSTIOR L. B. Smith, var. pe. ar. brevifolia laminis foliorum longioribus angustioribus- que ‘a fert Leaf=b blades lin linear, 5-7 cm. long, 3 mm. wide. Fig. 12: Apex of leaf; fig. 13: Flower. 8 Gerais: Serra da Lapa, November 1824, Riedel s. n. (K, pra 38b. VELLOZIA ARENICOLA L. B. Smith, sp. nov AV. brevifolia Seubert, cui affinis, caudici simplici, scapo brevissimo, ovario ellipsoideo differt. Subacaulescent, simple; leaves very densely imbricate, the complete ones 10-13 at the top of the caudex; sheaths ovate, 2 em. long, yellow, lustrous, very closely nerved, glabrous, curved at apex ne ng and more or less divided; blades linear, obtuse, 6-10 cm. long, 3.5 mm. wide, serrulate on the poiamg margins and been, ~ the keel, glabrous; scape solitary, very short; flower hidden among the terminal leaves; ovary ellipsoid, nar- numerous, about half as long as the tepals, denticulate-appen~ daged at base. hee 14: Apex of leaf; fig. 15: Ovary Gerais: In sand, Ti juco tales December 1824, Riedel shi ns (K, +; ° 1963 Smith, American Velloziaceae 513 49. VELLOZIA GRANULATA Goeth, & Henr. gore 23373. = Leaf-blades as short as 2 em. and as narrow as 4 mm. Fig. 16: Apex of leaf; fig. 17: Flower BRAZIL: Minas Gerais: Serra da Lapa, November 1824, Riedel s. 60, VELLOZIA SWALLENII L. B, Smith, Contrib. U. S. Nat. Herb. 35: 265, pl. 9, figs. 9, 10. 1 1962. BRAZIL: Golds: Goias City to Cavalcante, 1828, Burchell 7586 (K). Maranhao: Carolina to Para, 1829, Burchell 8972 (K). ‘ar gee BLANCHETIANA L. B. Smith, sp. nov. « sWallenii L. B, Smith, cui affinis, foliorum vaginis vix civisis » laminis margine serrulatis, ovarii tuberculis truncatis if rt. Caudex at least 4 cm. long Sepeeageate , Ogre let . leaves very densely imbricate, the complete o at t of the caudex; sheaths e, long, d - brous, closely nerved, splitting but slightly with age but then pectinate-fibrous by t rsistence > -strands ; blades » attenuate to a filiform apex, ong, . rulate on the thicken [4 an curely so on the keel beneath, otherwise smooth and glabrous; scapes 5 cm long, 3 ; and finely truncate-tuberculate especiall ovary subcylindric, ° » de y bases known); style at least 35 mm. long. Fig. 18: Section of leaf-blade; fig. 19: Ovary. ad BRAZIL: Bahia: Serra Assurua, near Rio Sao Francisco, 1828, Blanchet 2814 (K, type). 46a. BARBACENIA PLANTAGINEA L. B. Smith, sp. nov. Ex affinitate B. hilairei Goeth. & Henr. atque B. oxytepala Goeth. & Henr. sed ab omnibus speciebus foliorum laminis brevibus amplis apice late rotundatis et acuminulatis differt. Subacaulescent, simple, glabrous; leaves about 7; sheaths broadly ovate, 2 cm. long, closely nerved except the broad thin margin, yellow-brown, lustrous; blades oblong, broadly rounded then long-acuminate, 6-9 cm. long, to 2 cm. wide, serrulate on t ed 10-13 em. long, 1.3 mm. thick, curved; flowers red; perianth-tube very slenderly and regularly cylindric without enlargement for the ovary, curved, 5 cm. long, 4 mm, in diameter; tepals sub~ erect, oblong, obtuse, 25 mm. long; " stamens 12 mm, long; fila- ments oblong with flaring base and large suborbicular lobes; anthers linear, exceeding the filaments by 5 mm.; ovary 2 cm long. Fig. 20: Apex of leaf; fig. 21: Flower; fig. 22: Stamen (abaxial side). BRAZIL: Minas Gerais: Serra da Lapa, December 1824, Riedel 1061 (K, type). 51h PHYTOLOGIA Vol. 8, no. 9 American Velloziaceae a) ng ae aoe age —_ PS tie Bide , Vou, sr net Kia Se Mi foe _— © — —_ —y BR irs << Fig. 1, 2: Vellozia laevia; fig. 3-5: V. crispata; fig. 6, 7: V. droseroides; fig. 8, 9: V. vestita; fig. 10, 11: ao Sis. i, 13: V. brevifolia var. angustior; fig. 14, 15: V. arenicola; fig. 16, 17: V. granulata; fig. 18, io: ¥ - blanchetiana; fig. 20-22: Barbacenia plantaginea. 1963 Index Author Index to Volume Eight Alain, Bro. -- 369 Barkley, Fe. Ae -~ 329 Brown, H. De, & it H. J. —- 105 Oswald, F. W. - Smith, L. Paget, 0 an 497, 507 Wurdac. ok, Jede nt reeds 273, “bh, in, ~ 388° 395, ids, 515 57 zo 95, 108, 159, 175, 230, Index to Supra-specific Scientific Names in Volume Eight Abronia -~ 128, 140, 213 Acacia — 215, h71, 77 Acanthaceae -- 139, Acantholippia -- 95, 138 cerosae —- 125 Acopa -- 1)2 Aechmea -—~ 12, 218, 219, 506, 10 Aecidium ~- 280 Aegiphila -- 15-~21, 95, 385 Aerva -- 377 Agrimonia -- 1,39 Allardtia -- 221 Allophyllus -- 39 Allophylus -- 82 Moysie | -~ 12h, 128, 130, 132-- a. --10 Boma -- 12) Alternaria -—- 37 Amaranthaceae —- 141, 377 Amaryllidaceae -- 369 Amasonia ~- 9 Amblyarrhena -- 168, 170 Ambrina -- 31) Amomis -- 370 Ananas -- 12 Ananassa Anonymos — -- 12 fects -- 325, 327, 26-428 Antirrhinum -- 105, 107 Apios -~ 7, 61 Aquifoliaceae ~- 369 20 Artemisia —- 215, 277 Ascochyta —- Ascochyta -- 14 Aubletia -- 95, 96, leh, gas ae Avicennia -- 9--57, 95 Avicenniaceae — 9, 422 Baccharis — 1,1 Bacterium -- 87 Baillonia -- 95 Bakerantha -- 7 Bakeria -—- 7 Baptisia -- 277 Barbacena -- 513, 514 Berbena =-- 268 Bidens -- 139 54, 328 516 Bignoniaceae -- 20, 2, 46 Billardiera -- 95, 126, 26 Billbergia -- 50), 510 Blainvillea -~ 326 70 Bouchea -- 95, 128--130, 132, $995.1.95,' 437, 239, 196, 257; 273, 260, 316, 375—377 370 225, 2b. "239, “is, “199-301, 503, 50 05, oe » 509 Bros == ~ a 139, 141, 426, Bursera -- 0 Burseria -- 126 Byrsonima -- 369 ’ Callicarpa -- 57, < te 273, 8 Carya ~- 211 Caryophyllaceae -- 369 Casselia -- 95 Castelia — 95, 135, 137, 140 Catabe: 2 Ceci 142 69 Chaetolepis -~ 165, 166 PE 70 LOCA Vol. 8, no. 9 Chascanum -- 95, 130 Chamaesaracha —~ },),0 Chrysothamms -~ 152 Citharexylum ~- an 38, Pby:hb? Clerodendron ~- Clerodendrum ~- of 132, 159; aoe 316, 1 — 2h Ce ae — 326, 370 Congea -- Conoclinium — 1);1 Cordia — 388 Cornutia -- 95 Cottendorfia ~- Ro remanium -- 169, 170 Cressa -- 276, 277 Cuscuta -- 12 Cyclopogon -= 326 ymari Daphnopsis — 18 Dasylirion -- 9 Delphinium — 215 Dicaeoma — 280, Didynamia 4h a Angiospermia -- 51— 53 Diodia —- 276 Diostea — 95, 12h, 125, 132, 138, 143 Dipyrena -- 95, 133 Donatia -- 51 Dracocephalum — 213 1963 Drymaria — 369 Duvaua ~- ae 354, Dyckia Brstis - -- Be 35h Elionurus — Wy Elytraria — nk 141 Epicampes -- Epidendrum — at Eragrostis -- 277 Erinaceae — 12), 125 Eriocaulaceae -- 55 Eriocaulon -- 159, 386--388, Eryngium ~- 11 Erysimum ~ 176, 375, 377 Erysiphe ~- 142, 280, 41 hrina -- 1, 6 Eupatorium — 327 Euverbeneae -— 125 Ficus — Fimbristylis —- 141 Flourensia — 151 Foliosae — 12), 125, h2, 472 368 Forestiera —— 366, Fosterella -- 500, 501 Fouquieria — 77 Frankenia -- 276, 277 Galium -- 39 Gaylussacia —- 162 13 Ghinia ~- 126, 132, 176, 375, 3 Glandularia — 95, 119, 122-- imu, 12, 127, ts 216, 27h, 26 28; 4h2, 43° es 1, Gnaphalium —- 277, 378 Gochnatia — 325, 327, 328 Graffenrieda — 168 Grandularia -- Gravisia ~- 217--219, 225 Greigia — 226, 229 Index Guarea -- 326 Gutierrezia ~- Hedeoma — 21) Helenium —- 215 Helianthus —- 25) Helicodea -- SO Heliotropiaceae -- 128, 1) Helleranthus -- 9 Hemizonia — 11 Hepetis —- 22) Heterotrichum — 171, 172 Hexisea — 326 Hexiseae -—- 326 Hierobotana -- 95, 126, 132, Holmskioldia -~ 58 Holophyllae -- 12) Hydrophyllaceas — ll Hymenocallis -~ 369 Hypericum -- 378 flex —— Tnermes — 12h, 42, 472 inga — Ipomoea —— 87 Trenina -- 65 Juncus ~— 11, » 140, 142 Lan 13h, gi 160, 162 Laportea -- - Larrea ~- 215, 277, 471, 477 Laurus — 329, 342, 354 Lavandula — 257 TeLothrix — 162 518 PHYTO LOCA Leptochloa -~ 211 Leptostachyae —- 12h, 126 Lestodiplosis — 12 Liliaceae -- 500 Lindmania -- 501 Lippia —~ 58, 130, 131, 133— 3h1—3h5, » 39, 351, 353-355, BP 3595 61, Lobeliaceae -- 388 Lotus -- Lycopus — 279 Machoania -— 370 Macranthae — 12), 125, lhe wa peshhaceas — %9 M -- 16 osa — iecanats —~ 170 Myrtaceae -— 370 Vol.. 8, no. 9 tekmania -- 370 Myurteimanda Neoregelia -- 227, 2 Neosparton -- 12h, 125, 130, 8 Meoktin => 326 Nobiles -- 12], Nothoscordum -- 141 apceaia tases — 128, 140 Obletia -- 95, 2 Opuntia — 71 Orchidaceae -- 326, 369 Oryzopsis -—— 152 Ossaea — 173, 175 is -- 11 Pachycormus — 32 Pachystachyae — 12), 126 Paepalanthus -- 162, 164, 387, Pancratium -— 369 Parodianthus -- 95 Petiveriaceae — 128 Petrea —- 15, 95, 130 Phlox — )0 Phyla -- 128, 129, 131~-133, terme -- 129, 15 279, 376, aes —- 177, 279 Pinosia — %9 Pitcairnia —- 11, 13, 220, 222——226, 228, 229, “30s. 510 Plasmopara -- 142 Platymiscium ~- 388 7 ee Polemoniaceae -- Premna -- 62, 162, wae 273, a | Priva -- 95, 126 , 126-130, T32--1 136, 138, 1,0, 176, 1963 257, 280, 316, 375-377, Prosopis — 151 Prunus -—- ),56 Pseudocarpidium -—- 23, 95 Psychotria — 23, he Pycnothymus -- Quercus -- 72, 153, 211, 366, 367 Quesnelia -- 506, 510 Recordia -- 95 Rehdera -—- Reichenbachanthus -~- 326 Rubus —- 19 Rumex -~ 25) Sabal -~ ):22--h2h Sagraea -- 17h Salvia ~- 128, 129, 10 Sapindaceae -- 82 Sarcobatus — 275 Schizophyllae ~- 12h, 472 Schobera -- Schoenoplectus — 157, 458 Schuttleworthia -- 95, 202 Seriphioideae — 12h, 125 Index 519 Setaria —- 212, 25h Shuttleworthia — a 123, 202 Shuttlworthia -- Sida -- 276, 71 Spergularia -- 11 Spermacoce -- 370 Spl haerotheca — 12 Sphenodesme ——- 393 Spinosae —~ a Spiranthes -- 326 Stachytarpheta — 58, 126, 128--138, 1 10,176, 177, 257, Pe 316, 374377, 394, 412, Stevia — 141 Stignaphyllon —369 Stilbaceae -- 95 Styleurodon -- 126 Stylodon -- 95, 126, 131, 13h, Svensonia 5 i 136 Symphoremaceae 95 onanthus -- 388, 395, 396 epee —- 2, h6 Tamonea — 166 Taxodium -—~ 252 Tectona —- 95 Teijsmanniodendron ~- 23, 30, 273 Tetranychus — 142 Seles | honeys — 9 Teucrium -~ 140, 213 eel ae “an Tillandsia — 10, 13, 219-221, 22) 197, 499, 502-505, 508—510 Uerbena -- 96 — 211 Urbania — 125 Vellozia — 507, 511—51y Velloziaceae -- 507, 511, 513, 520 PHYTOLOGIA Vol. 8, no. 9 514 Verbent -- 96 Verben —~ 96 Verbesina -- 317, 370 Verbena -- 59, 60, 95, 97, 99, Verbina 101, 103, 108, 109, 111, 113, Verbinia — 96 117, 119--127, 129-152, Yerebna —- 96 175-179, 181-207, 209-216, Yornona — 96 Diy Gay Cady C39, C31, OD; 21-2117, 29, 251, 253-259, Vernonia — 141 261, 263, 265, 267--269, 271, Veronica —- 122, 137, 139, lui, 272, 27h--281, 283, 285, 287, 471 7 L779; ieee! 495, 496 Verbenaca -- 12),, 125 2227, 201; r% Verbenaceae eho, Sh, -95, 119, 422, 2 ish, ues "MSS Verben toe _ oy 279, Veronicena mem 122,157 Verticilliflorae -- eae 125 Vitee -—- Vitex -- 21, 23--l7, 61-~95 ) Vriesea -—- 221, 222, 229, 497, 198, a, Soh, 507-~509 Wahlenbergi ~- ii Wedelia -—- 28; 370 Yucca — Zapania —— 268 Zygopetalum -—- 369 Publication dates of Volume Eight No. 1 -= October 6, 1961 No. 2 — October 27, 1961 No. 3 -— December 19, 1961 No. -- February 2, 1962 No. 5 -— February 17, 1962 No. 6 — May 1, 1962 No. 7 -— May 26, 1962 No. 8 -— December 10, 1962 No. 9 -- January 22, 1963 PHYTOLOGIA is financed entirely by its contributors, each one paying in advance for the entire cost of printing, binding, and distributing his con- tribution. All money received from subscribers, after the expenses of col- lections have been deducted, will be distributed among the contributors upon the completion of a volume, in proportion to the space which they have used. Each contributor is therefore a shareholder in the magazine, assuming his part of the expenses and sharing in the profits, if any accrue. Each number consists of not less than 32 pages. All manuscript accepted will be published in the next issue, so that the size of numbers may vary greatly. A volume will contain about 32 signatures, 512 pages, or a smaller number of pages with an equivalent number of plates. This plan insures immediate publication of all accepted manuscript. Illustrations will be published according to the desires of the authors. No extra charge is made for line drawings, such as are ordinarily reproduced in inc, or for diagrams, tables, or charts, provided they conform to certain limitations of size and proportion. An extra charge will be made for half- depending on their size, as fixed by the engraver, with a minimum of about $2.25. Articles dealing with research in all lines of botany, in any reasonable gth, biographical sketches, and critical reviews and summaries of liter- ature will be considered for publication. Floristic lists, casual notes of an amateur or so-called popular type, and polemics will not be published. Advice on the suitability of manuscripts will be solicited, if necessary, from quali- fied botanists. Under the present cost of printing, the basic rate for a page or fraction thereof is approximately $3.00 for an edition of 250 copies. This price is subject wy. change without notice, since it depends entirely on the prices prevailing in the printing industry. _ Reprints will be furnished at cost. A proportionate fraction of the edition | of 250 copies is also furnished gratis to contributors. Upon request, the editors will send detailed instructions concerning the Pe aGOn: of manuscript or further information about the magazine. In- quiries may be addressed to the magazine or to either editor.