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Hoover Director Paleontological Research Institution 1259 Trumansburg Road Ithaca, New York 14850 U.S.A. 607-273-6623 VOLUME 90, NUMBER 324 MAY 9, 1986 Neogene Paleontology in the northern Dominican Republic 2. The Genus Strombina (Gastropoda: Columbellidae) by Peter Jung Paleontological Research Institution 1259 Trumansburg Road Ithaca, New York, 14850 U.S.A. Library of Congress Card Number: 86-61110 Printed in the United States of America Allen Press, Inc. Lawrence, KS 66044 U.S.A. CONTENTS Page а. 5 Subgenus Spiralta, new subgenus ..................... 13 Бестен е ЭМ сү ed Ro хул итын 5 Sozuppyi Woodring, T928 и 13 A E ааа Mud T A E 5 Subgenus Sincola Olsson and Harbison, 1953 .......... 14 A ЦЭЛ girl o cea еі 5 Se eurabensis Maury, ТО ү M ect errem TET 15 Боа Етар Руни a ae ЕН 6 S. pseudohaitensis (Maury, 1917) ................... 16 о (ru с НЕГ a a 7 БИСИ а а DD TS ха ан цайн a 18 Abbreviations of Repository Institutions .................. 8 S. bassi (Matty 19у 6 cen Hog ee 19 Systematic Paleontology Senanniebellae (Maury, 1917) 72" ОС 20 Я Au re Bay Ree 8 Subgenus Bifurcium Fischer, 1884 .................... 21 Genus Strombina MOTHS OVER ео 10 S. nuestrasenorae (Maury, 1917) .................... 22 Е ve. а Lo cbe oe 10 SECHNAEFNIEWESDELIEST. м ен ен. 23 5. cyphonotus Pilsbry and Johnson, 1911 ............ 11 Referencest@itede ss аре е eae 24 S. prisma Pilsbry and Johnson, 1911 ................ 12 А Ва ТЕГЕ А Жена ы ЫКЫ sque 26 LIST OF ILLUSTRATIONS Text-figure Page ] mer SOU Destion Oi шхезловесате оо NIC 002.55.006 60066 Tun Eth n кубы ае 6 2 Part ol RO udo. Section shoning ranges of species обоа sesi srar nee ee S QUA ee 7 3. Раш ol Rio Cana selon SITO WIS Tall ges: Ob Species OL ЭО озу... жуз у. ne een ea 7 4. Diagram showing relative frequency. of subgenera апа species Of SIFOMDINA osi reeet errie renent t even qnt here anne rernm 8 5. Intt ЖОНЕ of а Сабора showing mode of counting volutions ~. erri cece etre meena rey reir Mut Meee EE E EO UI 9 6. Graph showing (restored) heist ла тејацопенр 06.5. сурлопогих апа 5. Pris... ce ice rer tec teense ene sete nes 13 А ОТОО Э ИРОН ДОО о 15 В Heiphe Widtn.dlagr. nn В ҒИМА 12.20... 16 9. The two fields of size variability о} 9, gurabensis.and Su DSCUAONGILCNISIS SO 16 10. Apical vrewscot the protoconchs:0% S. orzabDensis.and S pseudohaitensis“ ао SHEER LEES И] ЇЇ Side views or the proloconchs 0652207627195 Andes. Pseudohanensiser..... nennen г Wels 127) 12: Height аралады Э САТРО оа 700000222222 72 18 Tx (PHO О Са ео ао 22 19 so Herm Int ата ES DOS STE ое а о АЕ 19 15 РОО Е 522542 он A AS e a pn ne ae CO er DL SET бойын olas 19 10: (Restored) иен Ауа арта 06.5 ranmiebellaen. Aone зуу узу nennen en LIP ae ta creed 20 Do ЁИНОСОБЕН ОЕ NO Teee ar ne na ka re samen o 21 18. (Restored) Вери тартан Of S- nuesiraserorae. | camus u. quaes echec a e a a наре Denn ost astuce Qe « 22 DE ОВ dria lua оғын ez m EL 23 20. оао size Уа а or S. NUestrasenorae aid S. сапа se eei e 0.10.6 609650666660 600006666 ети 23 27. Apical views OF thie Proloconc OD 5. MUCSITOSCHONAC andas CUNAC 2922220: Deere usine t me eh tet en 23 NEOGENE PALEONTOLOGY IN THE NORTHERN DOMINICAN REPUBLIC 2. The Genus Strombina (Gastropoda: Columbellidae) By PETER JUNG Naturhistorisches Museum Augustinergasse 2 CH-4051 Basel Switzerland ABSTRACT Ten species of Strombina are discussed and their type specimens refigured. S. cyphonotus and S. prisma, two species belonging to the subgenus Strombina, although always listed in Dominican faunas, in all probability do not occur in the Dominican Republic at all. The stratigraphic range and the mode of occurrence of the remaining eight species is indicated. One subgenus (Spiralta) and one species (S. canae) are described as new. The subgenus Sincola dominates the fauna both in number of species and in numbers of individuals. The subgenus Bifurcium is represented by two species, the subgenus Spiralta by one. RESUMEN Diez especies de Strombina son discutidas y sus ejemplares tipos refigurados. S. cyphonotus y S. prisma, dos especies perte- necientes al subgénero Strombina, aunque siempre han sido listadas en la fauna dominicana, mucho паз probablemente no aparecen en la República Dominicana. La distribución estratigráfica y el modo de aparición de las restantes ocho especies son indicados. Un subgénero (Spiralta) y una especie (S. canae) son descritos como nuevos. El subgénero Sincola predomina en la fauna tanto en número de especies como en número de individuos. El subgénero Bifurcium está representado por dos especies y el subgénero Spiralta por una. INTRODUCTION This is one of a series of taxonomic papers dealing with Neogene fossils from sections situated in the Ci- bao Valley, northern Dominican Republic (Text-fig. 1; Saunders, Jung, and Biju-Duval, 1986). The well-pre- served faunas from that area have been famous for a long time. The earliest collections of molluscs have been described Бу С. В. Sowerby II* (1850), and Pflug (1961) revised some of Sowerby’s species. The species described (but not figured) by Gabb (1873) have been redescribed and figured by Pilsbry (1922). However, all these papers are characterized by the lack of in- formation on geographic locations and stratigraphic Position. In this respect we find an improvement in Maury’s papers (1917a, 1917b) with more, although Not necessarily detailed information. The material studied in this paper has been collected from measured sections. The geographic location of the investigated areas is shown in Text-figure 1. For detailed information as to geographic location and Stratigraphic position of all the collecting stations, as well as to the general biostratigraphic framework and —_ * There were three persons by the name of G. B. Sowerby (father, Son, and grandson). They all published in the field of malacology, and it is therefore necessary to distinguish them. This has been done by Roman numbers. For additional information see Palmer (1966). the ages, see the paper by Saunders, Jung, and Biju- Duval (1986). Formational names have been used with care, because correlations from section to section are not certain. ACKNOWLEDGMENTS The material on which this paper is based was col- lected during field work carried out in the years 1978, 1979, and 1980. The field work was made possible by a grant from the Swiss National Science Foundation (Grant 2.646-0.76). The financial help and the assis- tance in the field provided by Institut Francais du Pé- trole are gratefully acknowledged. I am indebted to the following persons for the loan of specimens under their care: Messrs. John Peake and C. P. Nuttall, British Museum (Natural History), Lon- don, England; Dr. Robert Robertson, Academy of Nat- ural Sciences of Philadelphia, PA, U.S.A.; Dr. Tom Waller and Mr. Warren Blow, National Museum of Natural History, Washington, DC, U.S.A.; Dr. Peter Hoover, Paleontological Research Institution, Ithaca, NY, U.S.A.; and Dr. Emily Vokes, Tulane University, New Orleans, LA, U.S.A. The manuscript was criti- cally read by Richard J. Erickson, Tulsa, OK, U.S.A., and Christopher L. Garvie, Long Branch, NJ, U.S.A., and I am grateful for their suggestions. 6 BULLETIN 324 GUAYUBIN uber VS 2) Pro, Gua | 26/77 - xe MONCION Q 10 20кт 4 Rio Cana 2 Rio Gurabo En Р З Rio Mao ррег Cenozoic 4 Rio Amina 5 Cañada Zalaya 6 Rio Yaque del Norte 7 City of Santiago 8 Arroyo Puñal 9 Rio Verde Ї Oligocene - Early Miocene ? Mesozoic Text-figure 1.—Index map showing location of investigated areas in Cibao Valley, Dominican Republic. In addition I am grateful to Mr. Wolfgang Suter, photographer at the Naturhistorisches Museum Basel as well as Dr. Richard Guggenheim and Mr. Marcel Diggelin, both of the Scanning Electron Microscope Laboratory, University of Basel, Switzerland. Dr. Fred Rogl, Naturhistorisches Museum Vienna, Austria, has dated a sample from the type section of the Rio Banano Formation in Costa Rica. BIOSTRATIGRAPHY Out of the eight species of Strombina occurring in the Neogene of the Cibao Valley five are restricted to single sections. The other three species occur in more than one section. S. guppyi is represented by a single specimen which was found at the stratigraphically highest locality of the Rio Gurabo section, i.e., in the middle to late Plio- cene Mao Formation. S. pseudohaitensis is known from the Rio Mao sec- tion only and has been found over a horizontal distance of only about one kilometer, i.e., between Maury’s Bluffs 2 and 3. The dip of the beds in which it occurs is less than 5°, and its vertical range does not exceed 15 m. These beds are part of the Cercado Formation. 5. caribaea is restricted to the López section along Río Yaque del Norte north of the Village of Baitoa. This section exposes part of the Baitoa Formation (see Saunders, Jung, and Biju-Duval, 1986, text-fig. 25). The thickness of the sediments with S. caribaea is a little more than 40 m. The remaining five species occur in the sections of Río Gurabo and Río Cana. Their stratigraphic ranges are shown in Text-figures 2 and 3. S. gurabensis is restricted to the top part of the Cercado Formation of the Río Gurabo section (thickness of sediments about 50 m), whereas 5. nanniebellae only occurs in the up- per part of the Cercado Formation of the Río Cana section (thickness of sediments about 130 m). S. bassi, S. canae, and S. nuestrasenorae occur in both the Río Gurabo and Río Cana sections. S. bassi occurs in the Cercado Formation of the Río Cana sec- tion and in the top part of the Cercado Formation and the lowest part of the Gurabo Formation of the Río Gurabo section. S. canae has mainly been found in the Cercado Formation of the Río Cana section, but there is a single specimen from the Gurabo Formation of Río Gurabo. 5. nuestrasenorae mainly occurs in the Gurabo Formation of the Río Gurabo section; a few specimens are known from beds in Arroyo Zalaya (Saunders, Jung, and Biju-Duval, 1986, text-fig. 36), which are tentatively assigned to the Gurabo Forma- tion, and there is a single specimen from the Cercado Formation of the Río Cana section. With the exception of S. guppyi which also occurs in sediments of Pliocene age of Jamaica and Costa Rica, all the species of Strombina here described are so far known only from the Neogene ofthe Dominican Text-figure 2.—Part of Río Gurabo section showing ranges of — species of Strombina. Note the short range of S. canae at the top of this section. The range of S. bassi is discontinuous. DOMINICAN REPUBLIC NEOGENE. 2: JUNG 7 | Republic and hence are of only local biostratigraphic importance. | PALEOECOLOGY 8 The eight species of Strombina occurring in the areas S studied are grouped in three subgenera: Spiralta (one 300 m - ? | G species), Sincola (five species), and Bifurcium (two t species). Each of these subgenera has at least one living кррсад representative neither of which is found in today’s Ca- BARA ribbean faunal province, but only in Eastern Pacific Y PX waters. ty = Y = = ә Сэт = m = ас == 2 8 2 ота < = 5 == & 5 de 20 oS i ui S o = =P с O ER © п. —=> 5 © о aoe D 5 са = 2 $ < Сері ; v ic ек т (02) E EAT c 212: 3 © a Tos © 200 m 4 5 ЖУ Е = ТЕ © О % - < | > із a = © ја ц. = О ee à c = < 4-4 9 o © > a Фф HOC 5 5 > 5 о с RON Y = 5 © = o z | S © с E Y < = 150 m — i O Lu. ©) < © cc Lu O Text-figure 3.— Part of Rio Cana section showing ranges of species of Strombina. Note the short range of S. nuestrasenorae in the lower part of the Cercado Formation. The following all live in relatively shallow water: 5. (Spiralta) тасшоза (С. В. Sowerby I, 1832) lives “оп mud flats and offshore to depths of 37 m” (Keen, 1971, p. 601); S. (Sincola) gibberula (G. B. Sowerby I, 1832) lives “intertidally and in depths to 100 т” (Keen, 1971, p. 600); and S. (Bifurcium) bicanalifera (G. B. Sowerby I, 1832) has been “dredged in sandy mud at a depth of ten fathoms” [= 18 m] (original description, p. 113). The bathymetric range of these species is there- fore from 0 to 100 m. The representation of the three subgenera among the fossils from the northern Dominican Republic is very unequal. Text-figure 4 shows that the majority of the species are assigned to Sincola, and that in terms of numbers of specimens they represent the bulk of the available material. The mode of occurrence of each species (scattered in the sediment, in lenses, etc.) is mentioned in the taxonomic part of this paper. With the exception of the single specimen of S. guppyi, found in sands with pebbles, all the other species were associated with silty sediments. The majority of the specimens, however, has been found concentrated in shelly lenses, thin shell beds, or in conglomeratic layers. Thus there is little doubt that most of the Strombina material has been transported over some distance before final deposition. The species here described seem to have lived in shallow water of normal salinity. As mentioned under 5. bassi, some specimens of that species from Rio Cana have been found associated with large numbers of Arca 0 500 1000 1 дирру! Spiralta gurabensis pseudohaitensis e 2 caribaea -8 y ü bassi nanniebellae | nuestrasenorae Bifurcium | canae Text-figure 4. — Diagram showing relative frequency of subgenera and species of Strombina. BULLETIN 324 patricia G. B. Sowerby II, 1850, and many, but not all of the shells from Río Cana are smaller than those from Río Gurabo. It is possible that the smaller size of these specimens is due to the influence of brackish water. Considering the fact that practically nothing is known about the ontogeny and the life habits of living species of Strombina, it is difficult to make paleoecological statements of any precision. In addition it seems wiser to wait with paleoecological conclusions until the ma- jority of the groups of fossils distributed to specialists for study is reassembled again, and the respective pub- lications available. One conclusion concerning the early ontogenetic de- velopment of the species considered herein can, how- ever, be drawn with some confidence. A number of protoconchs have been studied with the aid of scanning electron microscope photography, and some of the photographs are reproduced on the accompanying plates. Two types of protoconchs can be observed: the first type shows a sinusigerous outer lip and is found in all the species of the subgenus Sincola; the second type has an opisthocyrt outer lip and has been observed in the two species assigned to the subgenus Bifurcium. According to Lebour (1945), Thorson (1950), and Shu- to (1974), both types of protoconchs would point to pelagic larvae, or— more precisely —to planktotrophic larvae, i.e., larvae that feed on phytoplankton (Thor- son, 1950, p. 10). The protoconchs with a sinusigerous outer lip are ““considered to be limited to the plank- totrophic larval type of long pelagic period" (Shuto, 1974, p. 246). ABBREVIATIONS OF REPOSITORY INSTITUTIONS The following abbreviations for repository institu- tions are used in this paper: ANSP: Academy of Natural Sciences, Philadelphia, PA, U.S.A. BMNH: British Museum (Natural History), London, England, U.K. NMB: Naturhistorisches Museum Basel, Switzerland (the letter H after NMB stands for gastropods) PRI: Paleontological Research Institution, Ithaca, NY, U.S.A. TU: Tulane University, New Orleans, LA, U.S.A. USNM: United States National Museum of Natural History, Washington, DC, U.S.A. SYSTEMATIC PALEONTOLOGY INTRODUCTION In this paper a total of 10 species ofthe genus Strom- bina is described. Two species of the subgenus Strom- DOMINICAN REPUBLIC NEOGENE. 2: JUNG 9 bina are discussed, and it is shown that in all proba- bility these do not occur in the Dominican Republic. The remaining eight species belong to three subgenera: Spiralta (one species), Sincola (five species), and Bi- furcium (two species). The field work on which this Paper is based has provided ample opportunity for extensive collecting with the result that many of the species found are represented by large “populations”. An attempt has been made to consult and refigure type specimens of all the taxa involved and to compare them with the “populations” of the new collections. Being aware of the problems of biological species, the concept of population, when used in connection with fossils, has always been put in quotation marks and the approach in the following systematic part is largely morphological. This 18 obviously unsatisfactory to the neontologist, who would like to assess the amount of ecophenotypic variation within a given species. Sim- ilarly the application of “biological species” in pa- leontology meets with great difficulties. In the frame- work of the project on Neogene paleontology of the northern Dominican Republic it is certainly premature at this moment to interpret faunas in detail. Certain groups of fossils have been sent to various specialists for study. Once a good proportion of the taxonomic work has been published it may become possible to interpret more reliably the reassembled assemblages as to species concept, paleoenvironment, апа bio- Stratigraphic usefulness of individual species. A good example illustrating the difficulty of applying a clear species concept is provided by two species of Strombina: 5. gurabensis and 5. pseudohaitensis. Both species are represented by more than 1000 specimens. Both lived during the deposition of part of the Cercado Formation, i.e., penecontemporaneously, but not in the same area. 5. gurabensis occurs in the Rio Gurabo section and S. pseudohaitensis in the Rio Mao section, i.e., only about 10 km from each other. In addition their mode of occurrence in the silty sediments is sim- ilar. On the other hand there are consistent morpho- logical differences described in the systematic part of this paper which persuaded me to keep the two species Separate despite the initial temptation to place them in Synonymy. Similar circumstances could be listed for 5. nuestrasenorae and 5. canae. 1 am waiting for good arguments against this practice. Wherever possible the protoconchs of the species described below have been studied. The number of volutions of protoconchs is an integral part of their description. In order to avoid misunderstandings the method of counting volutions is shown in Text-figure 5. All the material from NMB localities is deposited at the NMB and only type, figured, or specially-men- tioned specimens carry individual catalog numbers. The heading “Diagnosis” is reserved for the descrip- tion of supraspecific categories, whereas the heading “Description” is used only in species-level taxonomy. Under the heading “Description” follows a description of the species, not of specimens. If there is a (partial) description ofa specimen, this is found under the head- ings “Remarks” or “Comparisons”. The equivalent descriptive terms “volution” and “whorl” are both used, especially if they occur in the same sentence. All generic, subgeneric, and species descriptions start with a general statement referring to size (e.g., “of small size”, “of medium to large size”, etc.). In the case of generic and subgeneric descriptions these statements are not quantified, because only a few species ofa given genus or subgenus are dealt with in this paper. A re- vision of the genus Strombina and related groups in- cluding both fossil and living species is now being pre- pared by the present writer. Although that study is still incomplete, the following preliminary figures for ranges of heights can be given: subgenus Strombina: 11-39 mm (small to large); subgenus Spiralta: 18-37 mm (medium to large); subgenus Sincola: 7-12 mm (small); subgenus Bifurcium: 7-15 mm (small). The general statement referring to size in the de- scriptions of species is quantified under the heading “Measurements”. The measurements of most species are plotted in a graph. The term “restored height” occurs frequently in this paper. It refers to measured specimens, the tip of which is incomplete (part of the protoconch, the protoconch, or even early spire whorls missing). The total height has been measured by extrapolating with the naked Text-figure 5.—Initial whorls of a gastropod showing mode of counting volutions. In this schematic drawing there are two-and- one-quarter whorls. eye. The accuracy thus reached is sufficient for material of the size category reported on in this paper. The heading “Оссштепсе” is followed by detailed geographic and stratigraphic information on a given species within the studied areas of the Dominican Re- public. Under the heading “Distribution” there is gen- eral geographic and stratigraphic information on a giv- en species within the Dominican Republic plus such information outside the Dominican Republic. Genus STROMBINA Morch, 1852 Strombina Mörch, 1852, р. 85. Strombocolumbus Cossmann, 1901, p. 241. Type species (by subsequent designation, Bucquoy, Dautzenberg, and Dollfus, 1882, p. 78).—Columbella lanceolata G. B. Sowerby I, 1832, p. 116. Recent, Ec- uador and Galapagos Islands. Diagnosis. —Shell of small to large size. General shape ranging from fairly stout to slender. Spire usually high. Sculpture may be lacking except for a varying number of spiral cords near the base, but axial ribs are the predominant sculptural elements. Outer lip generally thickened, with or without denticles on inner surface. Parietal callus present. Anterior canal more or less re- curved. Remarks.—The genus Strombina lives in tropical waters of the Western Hemisphere. Before 1950 the living representatives of the genus were thought to be restricted to the Eastern Pacific, an area from which more than 20 species have been described. For this reason Rutsch (1934, p. 68) considered the occurrence of Strombina in Pleistocene sediments of the Cabo Blanco area of Venezuela as remarkable. This Pleis- tocene species was later described аз S. caboblanquen- sis Weisbord, 1962 (р. 323, pl. 28, figs. 25-30, pl. 29, figs. 1-4). Woodring (1964, p. 252) reported on the discovery of S. pumilio (Reeve, 1859): “In 1950 R. W. Foster ... dredged specimens off Scarborough, To- bago, at a depth of 36 fathoms.” Gibson-Smith and Gibson-Smith (1974, p. 52) listed additional localities on the mainland of Venezuela and the Island of Mar- garita, where Recent specimens of S. pumilio had been found. According to these authors S. pumilio also oc- curs in Holocene beach deposits of mainland Vene- zuela, in raised beaches of the Araya Peninsula and the Island of Margarita, and in Pleistocene beds of the Araya Peninsula. Vink (1979, р. 7) reported on live 5. pumilio from Playa Santa Cruz near Cumaná, Уепе- zuela, found at a depth of 3 m “where it crawled in a habitat of coarse sand and fine algae”. A second Recent Caribbean species has been described from Cayo Francés, Islas Los Roques, Venezuela, under the name of S. francesae J. Gibson-Smith in Gibson-Smith and Gibson-Smith, 1974 (p. 54, pl. 2, figs. 1-4). BULLETIN 324 Petuch (1981, p. 323, figs. 35, 36) reported on the occurrence of living S. caboblanquensis Weisbord in depths of 15 to 40 m along the coast of Venezuela and on an unidentified species of Strombina (Petuch, 1981, p. 323, figs. 39, 40) from a depth of 35 m in the Golfo de Triste, Venezuela. These records increase the num- ber of species of Strombina living in the Caribbean Sea to four. The geographic distribution of these four Re- cent species as now known is restricted to a relatively small area within the southeastern Caribbean Sea. In contrast to the weak representation of the genus in the Recent Caribbean fauna, Strombina is wide- spread in Neogene sediments of the area, thus reflecting the closing of the isthmus between the two Americas during the Neogene. Almost 50 names of species of Strombina have been proposed for the eastern part of the Tertiary Caribbean faunal province alone, and the total age range of the genus is early Miocene to Recent. Subgenus STROMBINA sensu stricto Diagnosis. —Shell of small to large size, slender. Spire high. Sculpture on spire whorls predominantly axial, often becoming less conspicuous on late spire whorls. Aperture long and narrow. Parietal callus prominent. Outer lip thickened, with denticles on inner surface. Body whorl higher than half the height of the complete shell. The thickening of the outer lip and two additional thickenings at intervals of 120° in most species result in triangular cross-sections of the body whorl. Anterior canal recurved. Suture of body whorl somewhat as- cending. Remarks. — S. pumilio, S. francesae, S. caboblan- quensis, and Strombina sp. of Petuch (1981, p. 323), the four living Caribbean species, all belong to the subgenus Strombina s. s. According to Radwin (1977, p. 407) S. pumilio prefers a sandy substrate and pref- erably lives on the middle shelf, i.e., in depths of 22 to 80 m. The following ecologic information on 5. francesae is based on notes by Willard Brownell dated February 25, 1976. These notes were kindly sent to me by J. Gibson-Smith. Brownell states that S. francesae is relatively common in the western part of the Los Roques Archipelago. He had found about 20 live spec- imens buried in loose sand 1 to 4 cm below the surface at water depths ranging from 1 to 18 m. In addition he found numerous shells of S. francesae inhabited by hermit crabs. The fossil record of the Tertiary Caribbean faunal province contains about 20 species of this subgenus. According to the literature two species of the subgenus Strombina are said to occur in the Neogene of the Dominican Republic: S. cyphonotus and S. prisma. As discussed below there is a high probability that neither of these species occurs in the Dominican Republic. DOMINICAN REPUBLIC NEOGENE. 2: JUNG 11 Strombina (Strombina) cyphonotus Pilsbry and Johnson Plate 1, figures 1-9; Text-figure 6 Strombina gradate [sic] Gabb, 1873, p. 221 (in part). Not Columbella gradata Guppy, 1866, р. 288, pl. 16, fig. 10. Strombina cyphonotus Pilsbry and Johnson in Brown and Pilsbry, 1911, p. 353 (footnote), pl. 25, figs. 6, 7; Maury, 1917a, p. 97, pl. 15, figs. 7, 8; Pilsbry, 1922, p. 351; Weisbord, 1929, p. 42, pl. 7, figs. 9, 10. Description. —Shell of medium size. Spire moder- ately high. Protoconch not known. Up to eight post- nuclear whorls, Early whorls almost flat in profile, later Ones slightly convex. Spire whorls smooth except for faint, somewhat prosocline growth lines. On later spire whorls there may be a rudimentary, inconspicuous ax- ial sculpture consisting of narrow, distant axial ribs on the upper half of the whorl. Dorsal hump with a few Narrow axial ribs. Base of body whorl sculptured by Spiral cords. Anterior canal moderately long, slightly recurved. Outer lip thickened, with five to nine den- ticles on inner surface. Callus on inner lip prominent, lacking denticles. Lectotype (herein selected). — ANSP 3293 (РІ. 1, figs. 1-3). This is the specimen illustrated by Brown and Pilsbry (1911, pl. 25, fig. 6). Pilsbry (1922, p. 351) mentioned “the type and eight other specimens are no. 3893 A.N.S.P.” [instead of 3293], but did not designate a particular specimen as the “type” and therefore did not select а lectotype. Pilsbry and Johnson in Brown and Pilsbry (1911, p. 353) gave the measurements of the specimen figured by Brown and Pilsbry (1911, pl. 25, fig. 6) and probably considered this as the “type” without explicitly stating it. After these measurements (which are not quite correct) they continue: “Other specimens of the type lot measure: . . .”. For this reason the specimen mentioned above is selected as the lec- totype. Dimensions of lectotype. — Height (incomplete): 23.5 mm; width: 11.2 mm. Type locality. —Not known. Maury (1917a, p. 97, pl. 15, figs. 7, 8) mentioned and figured S. cyphonotus, but said little about the species and gave no locality information as she usually did for other species. In 1974 I received three specimens of S. cyphonotus from J. W. Wells, which are said to come from C. J. Maury’s collection, but were actually part of Gabb's collection as stated on an old label accompanying the specimens. This seems to be proof that Maury never found S. cyphonotus in the Dominican Republic, but received some specimens from Philadelphia labeled as coming from “Santo Domingo” (without any further locality information). State of preservation and coloration of these three specimens are exactly the same as those of the specimens of the type lot. S. cyphonotus is thus not represented in the material Maury collected in the Dominican Republic; nor has it been found by E. H. and H. E. Vokes, nor by our own field party. It is therefore highly probable that S. cyphonotus (as well as S. prisma: see below) does not occur in the Dominican Neogene. The specimens must have been mislabeled at an early stage: at least this seems to be the most probable source of confusion. Material. —A total of 13 specimens have been stud- ied: the lectotype, eight paralectotypes, the specimen figured by Weisbord (1929, pl. 7, figs. 9, 10), and the three specimens from Gabb's collection mentioned above. Measurements (in mm). — ratio of restored height restored height width height to width ANSP 3293 (lectotype) 23.5 24.5 11.2 22 [see РІ. 1, figs. 1-3] ANSP 60173 (paralectotype) 19.8 21.5 10.2 24 ANSP 60173 (paralectotype) 217 23.4 11.2 2.1 ANSP 60173 (paralectotype) 21.8 23.4 10.1 2.3 ANSP 60173 (paralectotype) 202 22.6 10.3 22 ANSP 60173 (paralectotype) 22.4 23.8 10.6 2.2 ANSP 60173 (paralectotype) 2257 эл 11.1 2.1 [see Р]. 1, figs. 4—6] ANSP 60173 (paralectotype) 23.3 24.7 11.2 22 ANSP 60173 (paralectotype) 23.4 24.6 10.4 2.4 PRI 22962 (figured by Weisbord, 1929) 20.5 22.7 10.4 2.2 [see Pl. 1, figs. 7-9] NMB H 17147 (from Gabb collection) 18.0 19.5 8.6 213 NMB H 17148 (from Gabb collection) 21.6 23.0 10.0 2:3 NMB H 17149 (from Gabb collection) 2912 28:2 10.6 2.2 Remarks.—The axial sculpture on the late spire whorls is variable. When present it is weak, but on Some specimens it is absent altogether. Although the cross-section of the body whorl is basically triangular, the thickening to the left of the aperture is more or less well developed. 12 BULLETIN 324 Comparisons. — S. cyphonotus has a stouter арреаг- ance than S. prisma. Its apical angle is larger than that of S. prisma, and its anterior canal is shorter. On an average S. cyphonotus is smaller than S. prisma (see Text-fig. 5). Occurrence. — As stated above it is most likely that S. cyphonotus does not occur in the Neogene of the Dominican Republic. Distribution. — The only more or less reliable record of this species is the single specimen described and figured by Weisbord (1929, p. 42, pl. 7, figs. 9, 10, refigured here on РІ. 1, figs. 7-9). It was collected ““be- tween Las Perdices and Pto. Colombia, Dept. of At- lantico", Colombia, and the age is given as “Miocene”. Strombina (Strombina) prisma Pilsbry and Johnson Plate 2, figures 1—9; Text-figure 6 Strombina gradate [sic] Gabb, 1873, p. 221 (in part). Not Columbella gradata Guppy, 1866, p. 288, pl. 16, fig. 10. Strombina prisma Pilsbry and Johnson in Brown and Pilsbry, 1911, p. 352 (footnote), pl. 25, figs. 9, 10; Maury, 1917a, p. 97, pl. 15, figs. 97 10; Pilsbry, 1922; p. 551. Description. —Shell of medium size. Spire high. Pro- toconch not known. Up to eight-and-one-half post- nuclear whorls. Early spire whorls almost flat in profile, late spire whorls slightly convex. Postnuclear whorls practically smooth except for slightly prosocline growth lines; a faint spiral striation may be present. Dorsal hump with a few short and narrow axial ribs or ac- centuated growth lines. Suture between early spire whorls in some cases channeled. Base of body whorl with strong spiral cords. Anterior canal long. Outer lip with six to 10 denticles. Callus on inner lip strong, without denticles. Lectotype (herein selected). — ANSP 3292 (PI. 2, figs. 1-3). This is the specimen illustrated by Brown and Pilsbry (1911, pl. 25, fig. 10). Pilsbry (1922, p. 351) mentioned “the type and six other specimens аге no. 3292 A.N.S.P.", but he did not designate a particular specimen as the “type” and therefore did not select a lectotype. Dimensions of lectotype. — Height (incomplete): 26.0 mm; width: 10.6 mm. Type locality. —Not known. Maury (1917a, p. 97, pl. 15, figs. 9, 10) mentioned and figured S. prisma, but said little about the species and gave no locality information as she usually did for other species. Al- though not absolutely certain, it is highly probable that Maury did not collect the species herself, but somehow had received specimens from the Gabb collection in Philadelphia. As no later collectors have ever found S. prisma again it is concluded that S. prisma most prob- ably does not occur in the Neogene of the Dominican Republic at all (see also under S. cyphonotus). Material. — Only the lectotype and six paralectotypes have been studied. Measurements (in mm).— ratio of restored height restored height width height to width ANSP 3292 (lectotype) [see Pl. 2, figs. 1-3] 26.0 26.6 10.6 215 ANSP 60174 (paralectotype) 22.0 24.1 912; 2.6 ANSP 60174 (paralectotype) [see Pl. 2, figs. 7-9] 22.8 24.1 10:35 235 ANSP 60174 (paralectotype) 24.0 29:6 10.8 2.4 ANSP 60174 (paralectotype) 29519 27.0 9.8 2.8 ANSP 60174 (paralectotype) [see Pl. 2, figs. 4—6] 2027 2742 10.0 257 ANSP 60174 (paralectotype) 29.0 30.0 120 2:4. Remarks. — The seven specimens of the type lot show some variability. The prominence of the axial sculp- ture on the dorsal hump is variable, but it may be absent altogether. Furthermore the suture between ear- ly spire whorls is channeled in some cases. The lec- totype does not have a channeled suture, but several of the paralectotypes do (see РІ. 2, figs. 4-6). Comparisons. — 8. prisma is most closely related to S. lessepsiana Brown and Pilsbry, 1911 (p. 352, pl. 25, figs. 11, 12) from the Gatun Formation of the Panama Canal Zone. According to Woodring (1964, p. 253, pl. 40, figs. 22, 23, 30, 31) S. lessepsiana is quite abundant in the Gatun Formation; specimens from the lower part of the formation are larger than those from the middle and upper parts. The axial sculpture is strongly variable: the whorls of specimens with axial ribs are somewhat shouldered, but if the axial sculpture is ab- sent, the profile of the whorls is almost straight. None of the specimens of the type lot of S. prisma has axial sculpture. The type lot of 5. lessepsiana (ANSP 1718) consists of seven specimens: four of them are frag- mentary; the remaining three shells lack axial sculpture and are thus so similar to S. prisma that one is tempted to consider them as one species. 5. waltonia Gardner, 1947 (р. 513, pl. 52, figs. 26- 28) from the middle Miocene Shoal River Formation of Florida essentially is a smaller species. Most of the many specimens of S. waltonia from the Shoal River Formation of Florida at hand, which were kindly sent for comparison by E. H. Vokes, have axial sculpture on later spire whorls and are shouldered to some de- gree. On some specimens the axial sculpture is absent. S. striatocostata Marks, 1951 (p. 110, pl. 7, fig. 7) from the middle (?) Miocene Daule Formation of Ec- DOMINICAN REPUBLIC NEOGENE. 2: JUNG 13 uador is another similar species. It is smaller than S. prisma and has a more pronounced axial sculpture. Occurrence. — Not known (see above). There is a high Probability that this species does not occur in the Neo- gene of the Dominican Republic. Distribution.—As the seven specimens of the type lot of this species are the only specimens known, and as their type locality is not known, the distribution of the species is not known either. Subgenus SPIRALTA, new subgenus Etymology of name.—L. spira = spire; L. altus = high; gender feminine. Е 2 30; Е ‚2 o “ә 25- 20- О =cyphonotus A = prisma ST Ї Ї Ї 8 10 12 width in mm Text-figure 6.—Graph showing (restored) height/width relation- Ship of S. cyphonotus and 5. prisma. Type species (herein designated).—Strombina ma- culosa (G. B. Sowerby I, 1832, p. 116). Living, Gulf of California to Ecuador. A syntype of S. maculosa is figured here (РІ. 3, figs. 1-3). Diagnosis. —Shell of medium to large size, slender. Spire high, considerably higher than height of aperture. Sculpture predominantly axial, consisting of straight or somewhat sigmoid axial ribs or knobs just below the suture, which may be axially-elongated. Axial sculpture weak or absent on early spire whorls. Spiral sculpture restricted to base of body whorl. Outer lip somewhat thickened; denticulation on its inner surface varying from almost absent to moderately well devel- oped. Parietal callus present but not prominent. Cross- section of body whorl circular except for the moderate thickening of the outer lip. Remarks.—The subgenus Spiralta is easily distin- guished from Strombina s. s. The proportion of the height of the aperture to the height of the spire is very different: in Spiralta the spire is much higher than in Strombina s. s. In addition Spiralta has a prominent axial sculpture on later spire whorls which is usually lacking in Strombina s. s. Furthermore Strombina s.s. has a dorsal hump on the body whorl resulting in a triangular cross-section of the body whorl, whereas the cross-section of the body whorl of species of Spiralta is circular due to the lack of a dorsal hump. As now known the subgenus Spiralta comprises two fossil species: S. falconensis H. K. Hodson in Hodson and Hodson, 1931, from the late Miocene Caujarao Formation of Falcón, Venezuela, and S. guppyi Woodring, 1928, from the early Pliocene Bowden For- mation of Jamaica. In addition three Recent species from the Eastern Pacific are assigned to Spiralta: S. elegans (G. B. Sowerby I, 1832), the type species, 5. maculosa (G. B. Sowerby I, 1832), and S. расеапа Dall, 1916. According to Keen (1971, р. 601) S. ma- culosa lives on mud flats and to depths of 37 m. S. maculosa has also been cited from the Pliocene of Car- men Island, Gulf of California, Mexico, by Hanna and Hertlein (1927, p. 147), but not from Pleistocene de- posits of that area (Grant and Gale, 1931, p. 699). The stratigraphic range of Spiralta is therefore late Miocene to Recent. Strombina (Spiralta) guppyi Woodring Plate 3, figures 4-6 Columbella ambigua Guppy, 1866, p. 288, pl. 16, fig. 8; Guppy, 1874, р. 439. Columbella [Strombina] ambigua Guppy. Pace, 1902, p. 52. Strombina ambigua Guppy. Gabb, 1873, p. 221; Olsson, 1922, p. 126, pl. 10, fig. 9. Not Columbella ambigua Kiener, 1841, p. 11. Strombina guppyi Woodring, 1928, p. 282, pl. 17, fig. 2. 14 BULLETIN 324 Description. —Shell of medium size, slender. Spire high. Protoconch consisting of one-and-one-quarter whorls. About eight postnuclear whorls. Spire whorls straight to slightly convex in profile. Axial sculpture consisting of 20 to 23 straight to slightly sigmoid axial ribs per whorl extending from suture to suture. On the body whorl the axial ribs may be restricted to the upper part. On early spire whorls the axial sculpture may be absent. Base of body whorl with some spiral cords. Outer lip only slightly thickened, having a few elongate denticles on its inner surface. Parietal callus not prom- inent. Body whorl circular in cross-section. Holotype. — USNM 115514. Dimensions of holotype. — Height (incomplete): 27.8 mm; width: 10.2 mm. Type locality. —Bowden, Jamaica. Bowden Forma- tion, early Pliocene. Material. —A single, incomplete specimen (NMB H 17059; see Pl. 3, figs. 4-6) from the Dominican Re- public is available. Measurements (of NMB H 17059).—Height: 22.6 mm; width: 8.6 mm. Remarks. —The only available specimen consists of four whorls showing remnants of a reddish-brown col- oration. It is somewhat more slender than the three topotypes at hand. The early spire whorls of one of the topotypes are smooth. Woodring (1928, p. 283) de- scribed the protoconch of S. guppyi as consisting of one-and-one-quarter whorls. This cannot be confirmed here as none of the specimens at hand have their pro- toconch preserved. The specimen from the “Gatun Stage” of the “Ba- nana River” section of Costa Rica described and fig- ured by Olsson (1922, p. 126, pl. 10, fig. 9) is also incomplete. According to Е. Rögl (written commun., 1973) at least part of the Banana River section falls within the Globorotalia margaritae Zone (early Plio- cene). Gomez and Valerio (1971, p. 43) include this section in the Rio Banano Formation. Comparisons.—The species most closely related to S. guppyi obviously is S. falconensis H. K. Hodson in Hodson and Hodson, 1931, from the late Miocene Caujarao Formation of Falcón, Venezuela. The late spire whorls of S. guppyi are slightly convex in profile, whereas those of S. falconensis are practically straight. The 15 topotypes of S. falconensis at hand as well as the topotype figured by Gibson-Smith and Gibson- Smith (1974, pl. 3, figs. 1, 2) show that the axial ribs do not extend from suture to suture as in S. guppyi but are restricted to the upper part of the spire whorls. The tendency towards slightly sigmoid axial ribs in S. gup- pyi is not seen in S. falconensis. No specimens of the Recent Eastern Pacific species S. elegans (G. B. Sowerby I, 1832) are at hand for comparison. According to the figure given by Keen (1971, fig. 1271), S. elegans has the same type of axial sculpture as S. guppyi but is larger. Occurrence. —Rio Gurabo: NMB locality 15833 (see Saunders, Jung, and Biju-Duval, 1986, text-fig. 4). This locality is at the top of the measured section and falls within the Mao Formation (middle or late Plio- cene). Distribution. —Jamaica: Bowden Formation (early Pliocene). Costa Rica: Rio Banano Formation (early Pliocene?). Dominican Republic: Mao Formation (middle or late Pliocene). Subgenus SINCOLA Olsson and Harbison Sincola Olsson and Harbison, 1953, p. 230. Type species (by original designation and tautony- my).— Strombina sincola Olsson, 1922. “Miocene” of Costa Rica. Diagnosis. —Shell of small size. General shape rang- ing from moderately stout to slender. Height of aper- ture about half the total height of shell or somewhat more. Sculpture predominantly axial. Axial sculpture usually restricted to early spire whorls. Spiral sculpture usually restricted to base of body whorl. Outer lip thickened, mostly with prominent denticles on its in- ner surface. Anterior canal short and recurved. Remarks. —Among the Strombina material from the northern Dominican Republic the subgenus Sincola is by far the most prominent group. There are five species represented by numerous specimens. Sincola is widely distributed throughout the Neo- gene Caribbean faunal province. There are almost 20 species names available for this subgenus alone. A sin- gle species survives in eastern Pacific waters: S. gib- berula (G. B. Sowerby I, 1832) which according to Keen (1971, p. 600, fig. 1273) ranges from Baja Cali- fornia to Peru and lives intertidally to depths of 100 m. S. gibberula is also recorded from Pleistocene beds of Lower California, Mexico, by Grant and Gale (1931, p. 699) and from the Pliocene of western Ecuador by Pilsbry and Olsson (1941, pp. 7, 35). The protoconchs of the species of Sincola to be de- scribed all have a sinusigerous outer lip. They are as- sumed to be pelagic larvae (Thorson, 1950), and this particular type of protoconch is interpreted as a plank- totrophic larva with a long pelagic life (Shuto, 1974, p. 246). Before considering this type of protoconch as char- acteristic of the subgenus Sincola, additional species of Sincola and in particular the living S. gibberula would have to be studied. DOMINICAN REPUBLIC NEOGENE. 2: JUNG 15 Strombina (Sincola) gurabensis (Maury) Plate 3, figures 7-9; Plate 4, figures 1-9; Plate 6, figures 1-7; Text-figures 7, 9, 10, 11 Columbella (Strombina) pseudohaitensis gurabensis Maury, 1917a, Б a ngula. Description. —Shell of small size, moderately slen- der. Protoconch smooth, consisting of about three vo- lutions not counting the sinusigerous outer lip. Post- nuclear whorls five to six. The first one to three Postnuclear whorls sculptured by axial riblets. There are 12 to 14 axial ribs per whorl. The earliest axial riblets may be slightly opisthocyrt, otherwise they are orthocline to somewhat prosocline. The whorls having axial ribs are shouldered, and the ribs themselves usu- ally are more prominent toward the lower suture. Later Spire whorls somewhat convex in profile, their surface smooth except for growth lines. Base of body whorl Sculptured by a varying number of spiral cords. Outer lip thickened, with or without denticles on its inner surface, Holotype. —PRI 28734 (see РІ. 4, figs. 1-3). The ho- lotype is not quite an adult and its outer lip not yet thickened. Dimensions of holotype.—Height (not quite com- plete): 7.3 mm; width: 3.4 mm. Type locality. — Río Gurabo at Los Quemados, Do- Minican Republic. This is not precise. For this reason the type locality is here restricted to NMB locality : s (Saunders, Jung, and Biju-Duval, 1986, text-fig. 124 114 height in mm 101 T Т Т Т “БМЛЕЕКТЕТЕ ЕН ОЕ width іп mm Text-figure 7.—Height/width diagram of S. gurabensis with line indicating heights corresponding to two widths (h = 2w). Material.—Maury (1917a, p. 96) stated that this species is rare. The present study of this species on the other hand is based on well over a thousand well- preserved specimens. At the type locality alone it is represented by at least 300 specimens. As shown by the scanning electron micrographs (РІ. 6, figs. 1-7) the protoconchs are somewhat corroded, and their surfaces are thus not very well preserved. Measurements. —The large amount of material available makes it possible to draw meaningful height/ width diagrams. In Text-figure 7 the measurements of “populations” of 50 adult individuals from ММВ lo- calities 15903, 15904, and 15907 (Saunders, Jung, and Biju-Duval, 1986, text-fig. 4) are plotted. The three “populations” have not been marked individually in Text-figure 7, because they do not show different pat- terns; on the contrary, quite a number of specimens have identical measurements. The line indicating heights corresponding to two widths (h = 2w) cuts al- most across the middle of the field of size variability, which means that the shells are about twice as high as wide. Remarks. — S. gurabensis is a fairly variable species. The denticles on the inner surface of the outer lip may be well developed, there may be only a few, or they may be absent altogether. Another variable feature is the axial sculpture on the early spire whorls. The de- velopment of the callus on the parietal wall is also variable. Figures 3 and 7 on Plate 6 suggest that the length of the sinusigerous outer lip of the protoconch is not constant. S. gurabensis has only been found in shell beds in- terbedded with silts and in conglomeratic shell beds. Thus it seems that the shells have been transported over a short distance before final deposition. Comparisons. — S. gurabensis is closely related to S. pseudohaitensis. The relationship of the two species will be discussed under the latter. S. lampra Gardner, 1947, from the middle Miocene Shoal River Forma- tion of Florida has somewhat similar proportions but lacks the axial sculpture on the early spire whorls. In addition S. /ampra is smaller than S. gurabensis and the profile of its whorls is almost straight, whereas in S. gurabensis it is somewhat convex. S. anomala (Gardner and Aldrich, 1919) from the late Miocene Duplin Marl of North Carolina is a small- er and stouter species. It has fewer whorls, but like 5. gurabensis its early spire whorls carry axial sculpture. Gardner (1948, p. 232, pl. 30, fig. 32) recorded S. anomala also from the late Miocene Duplin Marl of South Carolina and compared it with S. gurabensis, which she erroneously cited as occurring in the Gurabo Formation of the Dominican Republic. According to Maury (1917, p. 96), S. gurabensis only occurs in her 16 BULLETIN 324 “Zone С”, which is part of the Cercado Formation (Maury, 1917b, table). 5. sincola Olsson, 1922 from the “Miocene” of Costa Rica is a smaller species and its axial sculpture is not restricted to the early spire whorls but extends down to the penultimate whorl. Occurrence. — S. gurabensis occurs only in the sec- tion of the Rio Gurabo. It has been found at the fol- lowing localities: NMB 15896, 15897, 15900, 15903, 15904, 15906, 15907; 15910; 15911, 15912, 15913, 15914, 15915, 15916. All these localities are situated in the upper part of the Cercado Formation just below the unconformity marking the base of the Gurabo For- mation (Saunders, Jung, and Biju-Duval, 1986, text- fig. 4). The thickness of the sediments containing S. gurabensis is a little more than 50 т. Distribution. —So far S. gurabensis is not known out- side the Dominican Republic. Strombina (Sincola) pseudohaitensis (Maury) Plate 5, figures 1-9; Plate 6, figures 8-16; Text-figures 8-11 Columbella (Strombina) pseudohaitensis Maury, 1917a, p. 95, pl. 15, figs. 12, 13. Description. —Shell of small size, rather stout. Pro- toconch smooth, consisting of a little more than three volutions, not counting the sinusigerous outer lip. Five- and-one-half to six postnuclear whorls. The first three postnuclear whorls are always sculptured by 14 to 16 orthocline axial riblets per whorl. The axial sculpture may continue after the first three postnuclear whorls; if so, the axial ribs are less prominent and gradually become prosocline. On the body whorl the axial ribs height in mm Т Т Т T T 7 3 4 9 6 width іп mm Text-figure 8.—Height/width diagram of S. pseudohaitensis with line indicating heights corresponding to two widths (h = 2w). Pe 0 аге always absent. Spire whorls shouldered, their pro- file straight to slightly convex. Body whorl convex, smooth except for growth lines and spiral cords near the base. Outer lip thickened, with a varying number of weak, moderately prominent or heavy denticles on its inner surface. Parietal callus moderately prominent. Lectotype (herein selected). — PRI 28733. This spec- imen has been figured by Maury (1917a, pl. 15, fig. 12) and is refigured here (see РІ. 5, figs. 1-3). The only paralectotype (PRI 30582) has also been figured by Maury (1917a, pl. 15, fig. 13) and is refigured here (РІ. 5, figs. 4-6). Dimensions of lectotype.—Height: 8.8 mm; width: 5.1 mm. Type locality.—Cercado de Mao: Bluff 3 of Maury on Rio Mao, Dominican Republic. In order to be more precise the type locality is here restricted to NMB lo- cality 16913 (Saunders, Jung, and Biju-Duval, 1986, text-fig. 29). Material. —This species is represented by about 1000 well-preserved specimens. The type locality has yield- ed about 120 specimens. Measurements.—In the height/width diagram of Text-figure 8 the measurements of “populations” of 50 adult individuals from NMB localities 16913, 16923, and 16927 (Saunders, Jung, and Biju-Duval, 1986, text- fig. 29) have been plotted. The three “populations” have not been marked individually in Text-figure 8 as m Е 12- 11- height T T Т 6 width іп mm Text-figure 9.—The two fields of size variability of S. gurabensis (larger field) and S. pseudohaitensis with line indicating heights cor- responding to two widths (h = 2w). N- о- > о 0 1 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 14 they do not show different patterns. The line indicating height corresponding to two widths (h = 2w) lies on one side of the field of size variability, i.e., the width of the specimens is more than half the total height of the shell. Remarks. — S. pseudohaitensis is somewhat variable in its development ofthe axial sculpture and the strength of the denticles on the inner surface of the outer lip. S. pseudohaitensis mainly occurs in thin bands large- ly consisting of molluscs, and in shell beds interbedded With silts, but it has also been found scattered in silts. Maury included Planaxis crassilabrum Guppy, 1874, in the synonymy of her S. pseudohaitensis, stating that Guppy had described a young specimen. Guppy's fig- ure is indeed so bad that it does not allow for the recognition of details. However, S. crassilabrum is a distinct species of the subgenus Sincola in Trinidad (Jung, 1969, p. 508, pl. 53, figs. 14-20). Comparisons. — S. pseudohaitensis is closely related to S. gurabensis. It differs from S. gurabensis in being Stouter; its axial sculpture differs in details of its de- velopment and covers more spire whorls. On an av- erage there are more axial ribs per whorl in S. pseu- dohaitensis, and S. pseudohaitensis is smaller than 5. Surabensis, although Text-figure 9 shows some overlap of the two fields of size variability. In addition there Seem to be some differences in the development of the Protoconch of the two species. Text-figure 10 shows apical views of the protoconch of both species. The Protoconch of S. pseudohaitensis has slightly more Whorls, but at the same time has a somewhat smaller initial whorl than that of S. gurabensis. Furthermore the largest diameter of the protoconch of S. pseudo- haitensis is larger than that of 5. gurabensis. The po- Sition of the two specimens of Text-figure 10 is ad- mittedly not analogous, but this does not change the fact that the protoconch of S. pseudohaitensis is stouter than that of S. gurabensis. a b Text-figure 10. — Apical views of the protoconchs of S. gurabensis (a) and S. pseudohaitensis (b). a. from NMB locality 15914 (Río Gurabo); x 60 (see PI. 6, fig. 4; NMB H 17022). b. from NMB locality 16913 (Río Mao); x60 (see PI. 6, fig. 8; NMB H 17027). The well-rounded body whorl of S. pseudohaitensis is somewhat similar to that of S. harbisonae Olsson, 1964, from the Angostura Formation (middle Miocene according to Olsson) of northwestern Ecuador. S. har- bisonae carries axial sculpture on most of the spire whorls like S. pseudohaitensis, but it is larger and tends to have more postnuclear whorls. Olsson described the protoconch as consisting of about two smooth whorls. I have examined the holotype of S. harbisonae (USNM 644034) and noted that its protoconch consists of at least three whorls, although counting is difficult, be- cause the initial whorl is not well preserved. Occurrence. —5. pseudohaitensis occurs only in the lower part of the section of Rio Mao. It has been found at the following localities: NMB 16912, 16913, 16914, 16915, 16916, 16917, 16918, 16922, 16923, 16924, 16926, 16927, 16928, 16929, 16930, 16931, 16932, 17269, and TU 1294. All these localities are situated between Maury’s Bluffs 2 and 3 (Saunders, Jung, and Biju-Duval, 1986, text-fig. 29). The beds with S. pseu- dohaitensis probably are of about the same age as those carrying S. gurabensis. Distribution. —So far S. pseudohaitensis is not known outside the Dominican Republic. с Text-figure 11.—Side views of the protoconchs of 5. gurabensis (a) and S. pseudohaitensis (b). a. from NMB locality 15912 (Rio Gurabo); x 60 (see РІ. 6, fig. 2; NMBH 1 7019). b. from NMB locality 16913 (Río Mao); x 60 (see РІ. 6, fig. 13; NMB H 17029). с. overlay of a and b to show difference in size and proportions. Strombina (Sincola) caribaea Gabb Plate 7, figures 1-12; Plate 8, figures 1-7; Text-figures 12, 13 Strombina caribaea Gabb, 1873, p. 221; Maury, 1917a, p. 98, pl. 15, fig. 6; Pilsbry, 1922, p. 350, pl. 18, fig. 9. Strombina caribaea micra Pilsbry*, 1922, p. 351, pl. 18, figs. 10, 11; Not Strombina caribaea Gabb. Woodring, 1928, p. 284, pl. 17, fig. > Description. —Shell small and stout. Protoconch smooth, consists of two-and-one-half whorls not counting the sinusigerous outer lip. Postnuclear whorls six, smooth except for faint, almost orthocline growth lines. Spire whorls somewhat convex in profile and slightly shouldered. Body whorl moderately inflated, with amore or less well-developed, broad dorsalhump, and a narrower hump on its left side. Base of body whorl sculptured by about six strong spiral cords. Outer lip greatly thickened, with a few denticles on its inner surface. Parietal callus well developed. Posterior end of aperture usually somewhat channeled. Columella with a few elongated denticles. Lectotype of $. caribaea.—ANSP 3996. Pilsbry (1922, p. 429, pl. 18, fig. 9) selected and figured the lectotype (refigured here: Pl. 7, figs. 4—6). Dimensions of lectotype of S. caribaea.— Height: 10.2 mm; width: 5.2 mm. Type locality of S. caribaea.—A type locality has never been selected; it is here restricted to NMB lo- cality 17288 which is situated within the section south of the village of López on Rio Yaque del Norte, north of the village of Baitoa (Saunders, Jung, and Biju-Du- val, 1986, text-figs. 21, 25). Holotype of S. caribaea micra.—ANSP 2800. The only specimen of the type lot is refigured here (РІ. 7, figs. 1-3). Dimensions of holotype of S. caribaea micra.— Height: 5.8 mm; width: 2.7 mm. Type locality of S. caribaea micra.—A type locality has never been selected; it is here restricted to NMB locality 17288 (see Saunders, Jung, and Biju-Duval, 1986, text-figs. 21, 25). Material. —S. caribaea is represented by about 300 specimens. Most of these specimens have been col- lected at NMB locality 17286 but many of these are incomplete and not very well preserved. The type lo- cality has yielded only about 20 well-preserved spec- imens. Measurements.—The dimensions of a number of complete specimens are plotted in Text-figure 12. Remarks. – Тһе type lot of S. caribaea consists of six specimens, three of which are incomplete. The ho- * 5. caribaea micra was ascribed to Pilsbry and Johnson [italics mine] in the explanation of plate 18. P.J. BULLETIN 324 lotype of S. caribaea micra is an immature S. caribaea; it has only five postnuclear whorls instead of six and therefore lacks the dorsal hump on its last whorl (see РБ 3L 119) 2). The record of 5. caribaea from the early Pliocene Bowden Formation of Bowden, Jamaica, is based on a single specimen (USNM 369458) (Woodring, 1928, p. 284, pl. 17, fig. 5). Its protoconch is said to consist of one-and-one-half whorls; Dominican specimens on the other hand have a protoconch with two-and-one- half whorls (see also Text-fig. 13). Woodring men- tioned additional differences. I have not seen the spec- imen from Bowden but it obviously represents some other species. S. caribaea has mostly been found scattered in silts and more rarely in sands, but it has also been collected from conglomeratic layers and from shell beds domi- nated by Turritella. Comparisons.—The Recent Eastern Pacific species S. gibberula (G. B. Sowerby I, 1832) is similar to S. caribaea but considerably larger. Both species lack sculpture on the spire whorls. In contrast to S. cari- baea, S. gibberula has no or only weak indications of denticles on the inner surface of its outer lip. This latter feature, however, does not seem to be significant tax- onomically as shown by the variability in other species of the subgenus Sincola. 5. galvestonensis (Harris, 1895), from the “Deep Well at Galveston”, Texas, has axial sculpture on the early spire whorls. Harris stated that the axial ribs are “уегу rarely as strong as indicated in the figure”. The age of height in mm = 9 4. 6 width in mm Text-figure 12. — Height/width diagram of S. caribaea with line indicating heights corresponding to two widths (h = 2w). шш = 0 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 19 a b Text-figure 13.—Protoconch of 5. caribaea from ММВ locality 17288 (Río Yaque del Norte); х 70. a. apical view (see РІ. 8, fig. 7; NMB H 17034). b. oblique apical view of same specimen (see РІ. 8, fig. 6). the beds carrying 5. galvestonensis is given as late Mio- сепе. Occurrence. —S. caribaea only occurs in the section south of the village of López on Río Yaque del Norte, i.e., north of the village of Baitoa (Saunders, Jung, and Biju-Duval, 1986, text-figs. 21, 25). It has been col- lected at the following localities: NMB 16936, 16938, 16942, 17283, 17284, 17285, 17286, 17287, 17288, 17289. All these localities fall in the Baitoa Formation. Distribution. —So far S. caribaea is not known from Outside the Dominican Republic. Е Е 13- £ 5 124 Ф С 11- 10; 9. су U < О 1 23 Y 4 400 width in mm | Text-figure 14.— (Restored) height/width diagram of S. bassi with line indicating heights corresponding to two widths (h = 2w). Strombina (Sincola) bassi (Maury) Plate 8, figures 8-14; Plate 9, figures 1—6; Text-figures 14, 15 Columbella (Strombina) bassi Maury, 1917a, p. 96, pl. 15, fig. 17. Description. —Shell of medium size, moderately slender. Protoconch consists of slightly less than two- and-one-half smooth whorls not counting the sinusig- erous outer lip. Up to six-and-one-half postnuclear whorls. Profile of spire whorls straight to slightly con- vex. Axial sculpture prominent. Axial ribs orthocline to slightly prosocline. There are 13 to 17 axial ribs on early spire whorls and 15 to 21 axial ribs on the pen- ultimate whorl. Spire whorls somewhat shouldered; on the body whorl the shoulder is accentuated by a spiral cord. Axial ribs absent on last half of body whorl. Base of body whorl sculptured by a varying number of spiral cords. Outer lip thickened, with denticles on its inner surface. Parietal callus moderately prominent. Colu- mella with a varying number of denticles. Holotype. — PRI 28736 (refigured here: РІ. 9, figs. 1- 3). Dimensions of holotype. — Height: 11.0 mm; width: 5.3 mm. Type locality. — Río Gurabo at Los Quemados, Do- minican Republic; Maury's Zones D and E. It does not seem advisable to select a more precise type locality, because our collections do not contain 5. bassi from Maury's Zones D and E, which extend from the bridge of the Los Quemados-Santiago Rodriguez road up- stream (southward) over an airline distance of a little more than 2 km. Zones D and E include parts of the Gurabo Formation (Maury, 1917b). Material. —'This species is represented by about 70 specimens, many of which are immature or have a broken apex. Measurements. —'The dimensions of the specimens that can be measured with reasonable accuracy are plotted in Text-figure 14. a b Text-figure 15. — Protoconch of S. bassi from ММВ locality 16856 (Río Cana); x 70. a. apical view (see РІ. 8, fig. 14; ММВ Н 17092). b. oblique apical view of same specimen (see РІ. 8, fig. 12). 20 BULLETIN 324 Remarks. —This species is variable as to prominence and number of the axial ribs on the spire whorls. Ac- cording to Maury this species occurs only in the section of Rio Gurabo. The material at hand is partly from Rio Gurabo but also from the section along Rio Cana. In fact the best preserved protoconchs (Text-fig. 15; Pl. 8, figs. 8-14) are from Rio Cana. S. bassi usually occurs scattered in silts. Some spec- imens from Rio Cana have been found associated with large numbers of Arca patricia Sowerby, 1850. Many but not all of the shells from Rio Cana are smaller than those from Rio Gurabo. Comparisons. — S. gunteri Mansfield, 1930, from the late Miocene Choctawhatchee Formation and the Plio- cene of North St. Petersburg, Florida (Olsson and Har- bison, 1953) is more slender and somewhat smaller than S. bassi. Both species lack axial sculpture on the last part of their body whorls. The protoconch of S. gunteri is said to consist of three whorls. When I ex- amined the holotype of S. gunteri its protoconch was no longer preserved. S. gunteri leonensis Mansfield, 1930, had been collected from about the same level as S. gunteri; it most probably is an immature S. gunteri. The proportions of S. lloydsmithi Pilsbry and Brown, 1917, from beds of uncertain age (Miocene?) of north- ern Colombia are similar to those of S. bassi. Like S. bassi, S. lloydsmithi lacks axial ribs on the last part of the body whorl except for one axial rib behind the thickened outer lip; but its body whorl is sculptured by prominent spiral grooves from base to suture except on and just before the dorsal hump. When examining the holotype and only specimen of 5. lloydsmithi I noticed that its protoconch is broken off. According to the original description it consisted of three whorls. Another species lacking axial ribs on the last part of its body whorl is S. cunninghamcraigi (Rutsch, 1942) from the late Miocene Savaneta glauconitic sandstone Member of the Springvale Formation of Trinidad. The holotype (NMB H 6177) and many topotypes are avail- able for comparison. It is smaller than S. bassi and has fewer postnuclear whorls. Its protoconch consists of more than three whorls and its siphonal fasciole is somewhat swollen. | Occurrence. — S. bassi occurs in the sections of Rio Gurabo and Rio Cana and has been collected from the following localities. Rio Gurabo: NMB 15882, 15899, 15902; Rio Cana: NMB 16841, 16848, 16854, 16855, 16856, 16857, 17001, and TU 1230. Most of the spec- imens from Rio Gurabo have been found in the top part of the Cercado Formation (NMB locs. 15899, 15902) and only the two specimens (NMB H 17145) found at NMB locality 15882 come from the lower part of the Gurabo Formation. All the material from Rio Cana is from the upper part of the Cercado For- mation with the exception of a single specimen (NMB H 17146) from the Gurabo Formation (NMB loc. 16869) which has been identified as S. aff. bassi. This specimen is larger and more slender than the others and carries axial ribs also on the last part of its body whorl. Distribution. —So far 5. Баяя 18 not known from out- side the Dominican Republic. Strombina (Sincola) nanniebellae (Maury) Plate 10, figures 1-13; Plate 11, figures 1-9; Text-figures 16, 17 Columbella (Strombina) nanniebellae Maury, 1917a, p. 96, PETS, figs. 15, 16. Description. —Shell of medium size, stout. Proto- conch consists of three smooth volutions not counting the sinusigerous outer lip. Up to six-and-one-half post- nuclear whorls. The first one or two postnuclear whorls sculptured by orthocline axial ribs numbering 13 to 15 per whorl. Remaining spire whorls smooth except for prosocline growth lines. Profile of spire whorls some- what convex. Body whorl inflated, almost circular in cross-section, sculptured by spiral cords near the base and a more or less prominent spiral cord adjoining the suture. Outer lip thickened, with no or only a few den- ticles on its inner surface. Parietal wall with a callous thickening near the adapical end of the aperture. Lectotype (herein selected). — PRI 28735. This is the specimen figured by Maury (1917a, pl. 15, fig. 15). There is only one paralectotype (PRI 30583). The lec- totype is refigured here (Pl. 10, figs. 1—3). height in mm о eG. 7778 width іп mm Text-figure 16.—(Restored) height/width diagram of 5. nannie- bellae with line indicating heights corresponding to two widths (h = 2w). DOMINICAN REPUBLIC NEOGENE. 2: JUNG 21 Dimensions of lectotype. — Height: 12.4 mm; width: 6.8 mm. Type locality.—Rio Cana at Caimito, Dominican Republic, Maury’s Zones H and I. The type locality is here restricted to NMB locality 16856, which is situ- ated 2.5 km upstream from the old bridge of the Los Quemados-Santiago Rodriguez road at Caimito. This locality is in the Cercado Formation (Saunders, Jung, and Biju-Dival, 1986, text-fig. 15). Material. — As stated by Maury this species is abun- dant. There are a little more than 1000 specimens in our collections, most of them well preserved, but many incomplete. The type locality has yielded 120 speci- mens. Measurements. — The dimensions of a number of specimens have been plotted in Text-figure 16. No significant differences in size were found in different “populations”. The line indicating heights correspond- ing to two widths (h = 2w) lies on one side of the field of size variability, i.e., the width of the specimens is always more than half the total height of the shells. Remarks. —The development of the axial sculpture of S. nanniebellae is quite variable. Usually the first two postnuclear whorls carry axial ribs, but in some Specimens they are restricted to the first postnuclear whorl; in other cases they extend over the first three postnuclear whorls. On certain specimens the axial sculpture may be almost missing. In fact the type lo- cality has yielded many specimens with practically no axial ribs. On the other hand there are shells with axial wrinkles on parts of their body whorls. In those cases a low and broad dorsal hump is usually developed. The outer lip of S. nanniebellae is usually smooth On its inner surface, but a few inconspicuous denticles may be developed. The parietal callus is only promi- nent near the adapical end of the aperture. The protoconch is unsculptured and consists of three whorls (Text-fig. 17). The length of its sinusigerous Outer lip seems to be constant. There is one specimen a b . Text-figure 17.—Protoconch of S. nanniebellae from ММВ local- Пу 16856 (Rio Cana). a. apical view (see РІ. 11, fig. 5; ММВ H 17045); x70. b. oblique apical view of same specimen (see Pl. 11, fig. 4); x65. from NMB locality 16839 with a protoconch consisting of only two-and-three-quarters whorls (see РІ. 11, fig. 9). Does this mean that the number of protoconch whorls is not quite constant in a given species? S. nanniebellae mainly occurs in silts, be it scattered or concentrated in lenses or individual layers. It has also been found in conglomeratic lenses. Comparisons. —5. pigea Olsson, 1964, from the An- gostura Formation (middle Miocene according to Ols- son) of northwestern Ecuador is larger and more slen- der than S. nanniebellae. S. pigea has no axial sculpture at all, but the cross-section of the body whorl of both species is similar. 5. pseudohaitensis (Maury, 1917a) (discussed earlier in this paper) is consistently smaller, has a more prom- inent axial sculpture on its spire whorls, and less con- vex whorls. Occurrence. — This species occurs only in the section along Rio Cana. It has been found at the following localities: NMB 16834, 16837, 16838, 16839, 16842, 16844, 16853, 16854, 16855, 16856, 16857, 16984, 16986, 16989, 16995, 17003, 17004, and TU 1230, 1282, and 1301. All these localities are situated in the upper part of the Cercado Formation (Saunders, Jung, and Biju-Duval, 1986, text-fig. 15). Distribution. —So far S. nanniebellae is not known from outside the Dominican Republic. Subgenus BIFURCIUM Fischer, 1884 Bifurcium Fischer, 1884, p. 638 Bifurcina Cossmann, 1901, p. 230. Type species (by original designation and mono- Туру). — Columbella Бісапа ға С. В. Sowerby I, 1832. Recent, Eastern Pacific. Diagnosis. — Shell of small size. General shape rang- ing from stout to moderately slender. Sculpture pre- dominantly axial. Axial sculpture usually restricted to early spire whorls. Spiral sculpture restricted to base of body whorl. Outer lip thickened, with or without denticles on its inner surface. Outer lip extended adapi- cally, bordering a prominent posterior canal. Anterior canal short and recurved. Remarks. —This subgenus is represented in the ma- terial from the northern Dominican Republic by rel- atively few specimens. They are assigned to two species. So far the only species that has been included in Bifurcium is its type species, the Recent species Col- umbella bicanalifera G. B. Sowerby I, 1832. It had been dredged in the Galapagos Islands in sandy mud at a depth of 10 fathoms (= 18 m) according to the original description. According to Keen (1971, p. 587) it is known from the southern part of the Gulf of Cal- ifornia to Ecuador and the Galapagos Islands. Pilsbry 22 BULLETIN 324 and Olsson (1941, pp. 8, 35) cited this species (?) under the name Bifurcium bifurcium Fischer (a name that does not exist formally, ¡.e., a nomen nudum) from the Pliocene Jama Formation of western Ecuador. If the species described below are included, the stratigraphic range of Bifurcium is late Miocene to Re- cent. As mentioned below the protoconchs of 5. nuestra- senorae are not well preserved, and better specimens are needed to determine the shape of the outer lip. The protoconch of S. canae has an opisthocyrt outer lip, which would point to a pelagic-planktotrophic larval life according to Shuto (1974). Whether S. bicanalifera has the same type of protoconch remains to be seen. Strombina (Bifurcium) nuestrasenorae (Maury) Plate 11, figures 10-12; Plate 12, figures 1-9; Text-figures 18, 20, 21 Columbella (Strombina) neustrasenorae [sic] Maury, 1917a, p. 98, pl. 15, fig. 11 (nuestrasenorae in explanation of plate). Not Strombina neustrasenorae Maury. Woodring, 1928, p. 284. Description. —Shell small, rather stout. Protoconch smooth, consisting ofa little more than two volutions. Five postnuclear whorls. The first two or three post- nuclear whorls sculptured by orthocline to slightly pro- socline axial ribs. There are 14 to 20 axial ribs per whorl. Later whorls smooth except for faint growth lines. Profile of spire whorls straight to slightly convex. Whorls somewhat shouldered. Body whorl moderately inflated, its shoulder accentuated by a spiral depres- sion. Body whorl with spiral cords near its base and one or two more or less well-developed axial swellings on its left side. Outer lip strongly thickened, extended adapically, with prominent denticles on its inner sur- face. Parietal callus well developed, with a ridge form- ing the left border of the posterior canal. Columella with some denticles. Anterior canal recurved. Holotype. — PRI 28732. This was apparently the only specimen Maury had when she described this species. It is refigured here (see Pl. 12, figs. 4—6). Dimensions of holotype. — Height: 7.8 mm; width: 4.2 mm. Type locality. — Río Gurabo at Los Quemados, Do- minican Republic; Maury's Zone G. The type locality is here restricted to NMB locality 15882 which falls in the lower part of the Gurabo Formation (Saunders, Jung, and Biju-Duval, 1986, text-fig. 4). Material. — Maury stated that this is a rare species. Our collections have yielded only 25 specimens, many of which are incomplete and (or) not well preserved. In particular the protoconch is never well preserved, and the shape of its outer lip cannot be determined with certainty (see Pl. 11, fig. 10). Measurements. —The dimensions of a number of more or less complete specimens are plotted in Text- figure 18. Remarks.—' The axial sculpture of this species is variable: it is usually restricted to the first two or three postnuclear whorls but may be faintly developed on the fourth postnuclear whorl as well. The number of axial ribs per whorl is quite variable. The prominence of the axial swellings on the left side of the body whorl influences the general shape of the body whorl. A single specimen (NMB H 17086) from locality 16856 (Río Cana, Cercado Formation) has a consid- erably stouter appearance than the other specimens. This is largely due to the two strong axial swellings on the left side of the body whorl. It has been identified as 5. aff. nuestrasenorae (see Pl. 13, figs. 1-3). S. nuestrasenorae has been found scattered in silts, concentrated in thin shell beds, and in silts with bur- rows filled with small molluscs. Comparisons. —'The type species of Bifurcium, the Recent Eastern Pacific species S. bicanalifera (Sower- by, 1832), is more slender and larger than 5. nuestra- senorae. One of the three syntypes (BMNH 1966312) of S. bicanalifera from the Galapagos Islands is figured here for comparison (РІ. 13, figs. 4-9). The protoconch of S. bicanalifera consists of three whorls, and there are six postnuclear whorls and not five as in 5. nues- trasenorae. The outer lip 1s not as greatly thickened, there are no prominent axial swellings on the left side of the body whorl, and there is no strongly-developed parietal callus. The figured syntype has only weak den- ticles on the inner surface of the outer lip; in the other two syntypes they are more prominent. S. canae, n. sp., from the Cercado Formation of Río Cana is described below. It is essentially smaller than E E 114 i: £ 5, 10- Ф e 9- 8- о Ah BE width in mm Text-figure 18.—(Restored) height/width diagram of S. nuestra- senorae with line indicating heights corresponding to two widths (h = 2w). DOMINICAN REPUBLIC NEOGENE. 2: JUNG 23 5. nuestrasenorae and its axial sculpture is more strongly developed. Occurrence. —S. nuestrasenorae occurs in the Río Gurabo, Río Cana, and in the Arroyo Zalaya and has been found at the following localities. Río Gurabo: NMB 15869, 15873, 15878, 15882; Río Cana: NMB 16857 (one specimen only); Arroyo Zalaya: TU 1227A (see Saunders, Jung, and Biju-Duval, 1986, text-figs. 4, 15, 36). The specimens from the Río Gurabo section have been found in the lower part of the Gurabo For- mation and the single specimen (NMB H 17085; see Pl. 12, figs. 7-9) from the Río Cana section in the Cercado Formation. Distribution. —So far S. nuestrasenoraeis not known from outside the Dominican Republic. Strombina (Bifurcium) canae, new species Plate 13, figures 10, 11; Plate 14, figures 1-9; Text-figures 19-21 Etymology of name. — Named after Río Cana (gen- itive singular). Description. —Shell small and stout. Protoconch Smooth, consisting of one-and-three-quarters volu- tions. Five to five-and-one-half postnuclear whorls. Spire whorls straight in profile, sculptured by ortho- Cline axial riblets. Axial sculpture on last spire whorl may be faint. There are 13 to 16 axial ribs per whorl. Outer lip strongly thickened, extended adapically, usu- ally with prominent denticles on its inner surface. Body Whorl moderately inflated, with a varying number of Spiral cords near its base, and with three conspicuous axial swellings on its left side. Parietal callus well de- veloped, with an axial ridge forming the left margin of height in mm = о ee ar on width in mm 1 Text-figure 19.— (Restored) height/width diagram of S. canae with line indicating heights corresponding to two widths (h = 2w). 11 height in mm Т Т Т T 0 35.4 - Б 4 width in mm Text-figure 20.—The two fields of size variability of S. nuestra- senorae (upper field) and 5. canae with line indicating heights cor- responding to two widths (h = 2w). the posterior canal. Columella with a few denticles. Anterior canal recurved. Holotype. —NMB H 17041. Dimensions of holotype.—Height: 8.1 mm; width: 4.5 mm. Type locality. —NMB locality 16837: Rio Cana, Do- minican Republic; upper part of Cercado Formation (Saunders, Jung, and Biju-Duval, 1986, text-fig. 15). Material. —This species is based on about 160 spec- imens, many of which are complete and well preserved. The type locality alone has yielded about 140 speci- mens. Measurements.—The dimensions of a number of specimens are plotted in Text-figure 19. a b Text-figure 21.—Apical views of the protoconchs of S. nuestra- senorae (a) and 5. canae (Ъ). a. from ММВ locality 15873 (Rio Gurabo); x60 (see РІ. 11, fig. 12; ММВ H 17035). b. from ММВ locality 16837 (Río Cana); x 65 (see Pl. 13, fig. 11; ММВ Н 17038). Note that the views on both specimens are not exactly apical, but slightly oblique. 24 BULLETIN 324 Remarks. —The axial sculpture of 5. canae is some- what variable. Although it usually covers all the spire whorls, it may be reduced or even missing on the last and rarely on the last two spire whorls. Whenever the axial sculpture is missing on spire whorls their profile is somewhat convex. The number and strength of the denticles on the inner surface of the outer lip is also variable. On the left side of the body whorl there are usually three axial swellings, but in some specimens there are only two. In some specimens there are ad- ditional axial ribs on the front (ventral) side of the body whorl. S. canae has been found scattered in silts and con- centrated in conglomeratic layers, in shelly lenses, and in thin shell beds. Comparisons. — 5. canae is obviously closely related to S. nuestrasenorae; in fact it looks like a miniature S. nuestrasenorae. However S. canae is not only small- er than 5. nuestrasenorae but its proportions are some- what different: in S. canae the width of the shells is always more than half the total height, which is not the case throughout in S. nuestrasenorae (see Text-fig. 20). In addition the axial sculpture of S. canae covers more spire whorls on an average. In S. canae there are usually three axial swellings on the left side of the body whorl, in S. nuestrasenorae only two. Unfortunately the protoconchs of both species are not very well preserved, but it seems that both do not have a sinusigerous outer lip. The protoconch of S. nuestrasenorae has a little more than two whorls, that of S. canae only one-and-three-quarters. The speci- mens in Text-figure 21 are shown at slightly different enlargements, but it is still obvious that the initial whorl of S. canae is somewhat larger than that of S. nuestrasenorae. S. nuestrasenorae mainly occurs in the lower part of the Gurabo Formation of Río Gurabo, whereas most specimens of 5. canae have been collected from the upper part of the Cercado Formation of Río Cana. Occurrence. — S. canae has been found at the follow- ing localities. Río Cana: NMB 16835, 16836, 16837, TU 1230, 1301; Río Gurabo: NMB 15842 (one spec- imen only: see Pl. 14, figs. 7-9). The specimens from the Río Cana section have all been found in the upper part of the Cercado Formation and the only specimen from the Río Gurabo section in the Gurabo Formation (Saunders, Jung, and Biju-Duval, 1986, text-figs. 4, 6, ISO) Distribution. —So far S. canae is not known from outside the Dominican Republic. REFERENCES CITED Brown, A. P., and Pilsbry, H. A. 1911. Fauna of the Gatun Formation, Isthmus of Panama. Acad. Nat. Sci. Philadelphia, Proc., vol. 63, pp. 336-373, pls. 22-29, text-figs. 1-3. Bucquoy, E., Dautzenberg, Ph., and Dollfus, G. 1882-86. Les mollusques marins du Roussillon. Tome premier: Gastropodes avec atlas de 66 planches. Paris, 570 pp. The individual fascicules appeared as follows: 1 (pp. 1-40, pls. 1-5): Feb. 1882; II (pp. 41-84, pls. 6-10): Aug. 1882; MI (pp. 85-135, pls. 11-15): Feb. 1883; IV (pp. 136-196, pls. 16-20): Aug. 1883; V (pp. 197-222, pls. 21-25): Jan. 1884; VI (pp. 223-258, pls. 26-30): Feb. 1884; VII (pp. 259- 298, pls. 31-35): Aug. 1884; VIII (pp. 299-342, pls. 36- 40): Sep. 1884; IX (pp. 343-386, pls. 41-45): Feb. 1885; X (pp. 387-418, pls. 46-50): Aug. 1885; XI (pp. 419-454, pls. 51-55): Jan. 1886; XII (pp. 455-486, pls. 56-60): Apr. 1886; XIII (pp. 487-570, pls. 61-66): Oct. 1886. Cossmann, M. 1901. Essais de paléoconchologie comparée. Quatriéme livrai- son. Paris, 293 pp., 10 pls. Dall, W. H. 1916. Notes on the West American Columbellidae. Nautilus, vol. 30, No. 3, pp. 25-29. Fischer, P. 1880-87. Manuel de conchyliologie et de paléontologie conchy- liologique ou histoire naturelle des mollusques vivants et fossiles. Paris, 1369 pp., 23 pls., map. The individual fas- cicules appeared as follows: I (pp. 1-112): 21 Sep. 1880; II (pp. 113-192): 16 Mar. 1881; III (pp. 193-304): 28 Jul. 1881; IV (pp. 305-416): 5 May 1882; V (pp. 417-512): 21 Feb. 1883; VI (pp. 513-608): 20 Dec. 1883; VII (pp. 609—688): 30 Jun. 1884; VIII (pp. 689—784): 29 Jan. 1885; IX (pp. 785-896): 31 Aug. 1885; X (pp. 897-1008): 30 Apr. 1886; XI (pp. 1009-1369): 15 Jun. 1887. Gabb, W. M. 1873. On the topography and geology of Santo Domingo. Am. Philos. Soc., Trans., new ser., vol. 15, pp. 49-259, 2 maps. Gardner, J. A. 1947. The Molluscan fauna of the Alum Bluff Group of Florida. Part 8. U. S. Geol. Survey, Prof. Paper 142-H, pp. 493- 656, pls. 52-62. 1948. Mollusca from the Miocene and lower Pliocene of Virginia and North Carolina. Part 2. Scaphopoda and Gastropoda. U. S. Geol. Survey, Prof. Paper 199-B, pp. 179-310, pls. 24-38. Gardner, J. A., and Aldrich, T. H. 1919. Mollusca from the upper Miocene of South Carolina with descriptions of new species. Acad. Nat. Sci. Philadelphia, Proc., vol. 71, рр. 17-53, pls. 1-4. Gibson-Smith, J., and Gibson-Smith, W. 1974. The genus Strombina (Mollusca: Gastropoda) in Venezue- la, with descriptions of a new Recent and some fossil species. Asoc. Venez. Geol. Min. Petr., Bol. Informativo, vol. 17, Nos. 4-6, рр. 49-70, pls. 1-4, text-fig. 1. Gomez P., L. D., and Valerio G., C. E. 1971. Lista preliminar ilustrada de los moluscos fosiles de la formación Rio Banano (Mioceno), Limon, Costa Rica. Inst. Geogr. Nacional (Costa Rica), Informe Semestral, pp. 43— 62, 42 figs. 2 Grant, U. S., and Gale, Н. R. 1931. Catalogue of the marine Pliocene and Pleistocene Mollus- ca of California and adjacent regions. San Diego Soc. Nat. Hist., Memoirs, vol. 1, 1036 pp., 32 pls., 5 text-figs., 3 tables, diagrams A-D. DOMINICAN REPUBLIC NEOGENE. 2: JUNG 25 Guppy, В. J. L. 1866. On the Tertiary Mollusca of Jamaica. Geol. Soc. London, Q. J., vol. 22, pp. 281-295, pls. 16-18. 1874. On the West Indian Tertiary fossils. Geol. Mag., decade 2, vol. 1, pp. 404-411, 434—446, pls. 16-18. Hanna, G. D, and Hertlein, L. G. 1927. Expedition of the California Academy of Sciences to the Gulf of California in 1921. Calif. Acad. Sci., Proc., ser. 4, vol. 16, No. 6, pp. 137-157, pl. 5. Harris, G. D. 1895. Neocene Mollusca of Texas, or Fossils from the Deep Well at Galveston. Bull. Am. Paleontol., vol. 1, No. 3, pp. 1- 32, pls. 1-4. Hodson, F., and Hodson, H. K. 1931. Some Venezuelan mollusks. Bull. Am. Paleontol., vol. 16, No. 59, pp. 1-94, pls. 1-24. Jung, P. 1969. Miocene and Pliocene mollusks from Trinidad. Bull. Am. Paleontol., vol. 55, No. 247, pp. 289—657, pls. 13-60, text- figs. 1-4. Keen, A. M. 1971. Sea shells of tropical West America. Stanford Univ. Press, Stanford, Calif. (second ed.), 1064 pp., pls., figs. Kiener, L. C. 1841. Species général et iconographie des coquilles vivantes ..., Columbella. 63 pp., 16 pls. [not seen] (fide Sherborn, C. D., and Woodward, B. B., 1901, Proc. Malac. Soc. London, vol. 4, pp. 217, 219). Lebour, M. V. 1945. The eggs and larvae of some prosobranchs from Bermuda. Zool. Soc. London, Proc., vol. 114, No. 4, pp. 462-489, text-figs. 1-43. Mansfield, W. C. 1930. Miocene gastropods and scaphopods of the Choctawhatch- ee Formation of Florida. Florida State Geol. Survey, Bull. No. 3, pp. 1-185, pls. 1-21. Marks, J. G. 1951. Miocene stratigraphy and paleontology of southwestern Ecuador. Bull. Am. Paleontol., vol. 33, No. 139, рр. 271- 432, pls. 1-9, 12 figs. Maury, C. J. 1917а. Santo Domingo type sections and fossils. Pt. 1. Bull. Am. Paleontol., vol. 5, No. 29, pp. 1-251, pls. 1-39. 19176. Santo Domingo type sections and fossils. Pt. 2. Bull. Am. Paleontol., vol. 5, No. 30, pp. 1-43, pls. 1-3, 1 text-fig., 1 table. Morch, O. A. L. 1852. Catalogus conchyliorum quae reliquit D. Alphonso d'Aguirra et Gadea Comes de Yoldi. Hafniae [= Copenhagen], fasc. 1, 170 pp. Olsson, A. A. 1922. The Miocene of northern Costa Rica. Bull. Am. Paleontol., vol. 9, No. 39, pp. 1-168, pls. 1-32. 1964. Neogene mollusks from northwestern Ecuador. Paleontol. Research Inst., Ithaca, NY, pp. 1-256, pls. 1-38. Olsson, A. A., and Harbison, A. 1953. Pliocene Mollusca of southern Florida, with special refer- ence to those from North Saint Petersburg. With special chapters on Turridae by William G. Fargo, and Vitrinel- lidae and freshwater mollusks by Henry A. Pilsbry. Acad. Nat. Sci. Philadelphia, Monographs, No. 8, pp. 1-458, pls. 1-65, text-figs. 1-2, 2 maps. Pace, S. 1902. Contributions to the study of the Columbellidae. Malac. Soc. London, Proc., vol. 5, pp. 36-154. Palmer, K. V. W. 1966. Who were the Sowerbys? Sterkiana, No. 23, pp. 1-6. Petuch, E. J. 1981. А relict Neogene caenogastropod fauna from Northern South America. Malacologia, vol. 20, No. 2, pp. 307-347, figs. 1-130. Pflug, H. D. 1961. Mollusken aus dem Tertiár von Santo Domingo. Acta Humboldtiana, ser. geol. palaeont., No. 1, pp. 1-107, pls. 1-26, 1 text-fig. Pilsbry, H. A. 1922. Revision of W. M. Gabb's Tertiary Mollusca of Santo Do- mingo. Acad. Nat. Sci. Philadelphia, Proc., vol. 73, pp. 305—435, pls. 16-47, text-figs. 1-48. Pilsbry, H. A., and Brown, A. P. 1917. Oligocene fossils from the neighborhood of Cartagena, Co- lombia, with notes on some Haitian species. Acad. Nat. Sci. Philadelphia, Proc., vol. 69, pp. 32-41, pls. 5-6. Pilsbry, H. A., and Olsson, A. A. 1941. A Pliocene fauna from Western Ecuador. Acad. Nat. Sci. Philadelphia, Proc., vol. 93, pp. 1-79, pls. 1-19, 2 text- figs. Radwin, G. E. 1977. The family Columbellidae in the Western Atlantic. Veli- ger, vol. 19, No. 4, pp. 403-417, 2 pls., 4 text-figs. Reeve, L. A. 1858-59. Monograph of the genus Columbella. Conchologia Iconica (London), vol. 11, 37 pls. Rutsch, R. F. 1934. Die Gastropoden aus dem Neogen der Punta Gavilán in Nord- Venezuela. Abh. Schweiz. Palaeont. Ges., Bde. 54— 55, pp. 1-169, pls. 1—9, 20 text-figs. 1942. Die Mollusken der Springvale-Schichten (Obermiocaen) von Trinidad (Britisch- West-Indien). Verhandl. Naturf. Ges. Basel, Bd. 54, pp. 96-182, pls. 3-9, text-figs. 1-2. Saunders, J. B., Jung, P., and Biju-Duval, B. 1986. Neogene paleontology in the northern Dominican Repub- lic. 1. Field surveys, lithology, environment, and age. Bull. Am. Paleontol., vol. 89, No. 323, pp. 1-79, pls. 1-9, text- figs. 1-39, tables 1-4, appendices 1-5. Shuto, T. 1974. Larval ecology of prosobranch gastropods and its bearing on biogeography and paleontology. Lethaia, vol. 7, pp. 239—256, figs. 1—8, 3 tables. Sowerby, G. B. I* 1832. New species of shells collected by Mr. Cuming on the west- ern coast of South America and among the islands of the South Pacific Ocean. Zool. Soc. London, Proc. Comm. Sci. Corr., Part II, pp. 113-120. Sowerby, G. B. II* 1850. Descriptions of new species of fossil shells found by J. S. Heniker, Esq., in Moore, J. C., On some Tertiary Beds in the Island of San Domingo: from notes by J. S. Heniker, Esq., with remarks on the fossils. Geol. Soc. London, Q. J., vol. 6, pp. 44—53, pls. 9-10. Thorson, G. 1950. Reproductive and larval ecology of marine bottom inver- tebrates. Biol. Review, vol. 25, pp. 1-45, text-figs. 1-6. Vink, D. 1979. Reeve was right. It’s Strombina pumilio. Hawaiian Shell News, vol. 27, No. 2, p. 7, figs. * See footnote, page 5. BULLETIN 324 Weisbord, N. E. Woodring, W. P. 1929. Miocene Mollusca of northern Colombia. Bull. Am. Pa- 1928. Miocene mollusks from Bowden, Jamaica. Part 2, gastro- leontol., vol. 14, No. 54, pp. 1-57, pls. 1-9. pods and discussion of results. Carnegie Inst. Washington, 1962. Late Cenozoic gastropods from northern Venezuela. Bull. Publ. No. 385, 564 pp., 40 pls., 3 text-figs. Am. Paleontol., vol. 42, No. 193, pp. 1-672, pls. 1-48. 1964. Geology and paleontology of Canal Zone and adjoining parts of Panama. Description of Tertiary mollusks (gas- tropods: Columbellidae to Volutidae). U. $. Geol. Survey, Prof. Paper 306-С, рр. 241-297, pls. 39-47, 1 table. EXPLANATION OF PLATE 1 Figure Page 1-9: Strombına (Strombing) cyphonotus Pilsbry AAA ни ous says cute 11 1-3. Lectotype, ANSP 3293; locality not known; specimen figured by Brown and Pilsbry (1911, pl. 25, fig. 6); 1. front view; 2. rear view; 3. from right side. height: 23.5 mm; width: 11.2 mm. 4-6. Paralectotype, ANSP 60173; locality not known; specimen figured by Brown and Pilsbry (1911, pl. 25, fig. 7); 4. front view; 5. rear view; 6. from right side. height: 22.7 mm; width: 11.1 mm. 7-9. PRI 22962; specimen figured by Weisbord (1929, pl. 7, figs. 9, 10) from “between Las Perdices and Рю. Colombia, Dept. of Atlantico”, Colombia (“Міосепе”); 7. front view; 8. rear view; 9. from right side. height: 20.5 mm; width: 10.4 mm. PLATE 1 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 2 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 22 EXPLANATION ОЕ PLATE 2 Figure 159 Strombina (Strombina) prisma Pilsbry and Johnson ce... ee nen 12 1-3. Lectotype, ANSP 3292; locality not known; specimen figured by Brown апа Pilsbry (1911, рі. 25, fig. 10); 1. front view; 2. . rear view; 3. from right side. height: 26.0 mm; width: 10.6 mm. 4-6. Paralectotype, ANSP 60174; locality not known; 4. front view; 5. rear view; 6. from right side. Note channeled suture of early spire whorls. height: 25.7 mm; width: 10.0 mm. 7-9. Paralectotype, ANSP 60174; locality not known; specimen figured by Brown and Pilsbry (1911, pl. 25, fig. 9); 7. front view; 8. rear view; 9. from right side. height: 22.8 mm; width: 10.5 mm. BULLETIN 324 EXPLANATION OF PLATE 3 Figure Page 1-3. 4-6. 7-9. мэртипа US pidlia) таслах (Ge B. SOwerDy I). Ko pees os еее, 15 BMNH 1982136; syntype. “Guacamayo, Central America, sandy mud, 10 fathoms”; type species of the subgenus Spiralta; 1. front view; 2. rear view; 3. from right side. height: 23.9 mm; width: 8.2 mm. Оа (90 раа) SUPP VEIN OGLE ... 22222222... ИН 0... 15 ММВ H 17059; from NMB locality 15833: Rio Gurabo; Mao Formation (middle ог late Pliocene); 4. front view; 5. rear view; 6. from right side. height: 22.6 mm; width: 8.6 mm. а (ПШАШГУ) ... 00 оюм уйкун сүх нх ухаа неон. 15) NMB Н 17063; from ММВ locality 15907: Rio Gurabo; upper part of Cercado Formation; 7. front view; 8. rear view; 9. from right side. Specimen without denticles on inner surface of outer lip. height: 10.1 mm; width: 4.8 mm. PLATE 3 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 x10 x10 x10 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 4 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 29 EXPLANATION OF PLATE 4 Figure 129, Strombina (Sincola)sZurabensis (ао) о оета ИОГ er. 15 1-3. Holotype, PRI 28734; Rio Gurabo at Los Quemados; upper part of Cercado Formation; 1. front view; 2. rear view; 3. from right side. Specimen is not quite adult. height: 7.3 mm; width: 3.4 mm. 4-9. Front views of six specimens showing variability of general outline, sculpture on early spire whorls, development of outer lip, and development of denticles on its inner surface. 4-6. All from NMB locality 15912: Rio Gurabo; upper part of Cercado Formation; 4. NMB H 17060. height: 9.7 m; width: 5.2 mm; 5. NMB H 17061. height: 8.9 mm; width: 4.6 mm; 6. NMB H 17062. height: 9.2 mm; width: 5.3 mm. 7-9. All from ММВ locality 15907: Rio Gurabo; upper part of Cercado Formation; 7. ММВ H 17065. height: 9.5 mm; width: 4.9 mm; 8. NMB H 17066. height: 9.8 mm; width: 4.4 mm; 9. NMB H 17068. height: 10.1 mm; width: 4.7 mm. 30 BULLETIN 324 EXPLANATION OF PLATE 5 Figure Page 1-5: тот (9 тон psendohawWensts (Maut)... une na MS, не... 16 1-3. Lectotype, PRI 28733; Rio Mao, Bluff 3 of Maury; Cercado Formation; 1. front view; 2. rear view; 3. from right side. height: 8.8 mm; width: 5.1 mm. 4-6. Paralectotype, PRI 30582; Rio Mao, Bluff 3 of Maury; Cercado Formation; 4. front view; 5. rear view; 6. from right side. height: 8.7 mm; width: 4.9 mm. 7-9. Front views of three specimens showing variability of sculpture on spire whorls, thickness of outer lip, and development of denticles on its inner surface. All from NMB locality 16924: Rio Mao; Cercado Formation; 7. NMB H 17075. height: 8.4 mm; width: 4.6 mm; 8. ММВ H 17076. height: 7.7 mm; width: 4.4 mm; 9. ММВ H 17077. height: 7.6 mm; width: 4.5 mm. PLATE 5 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 31 EXPLANATION OF PLATE 6 Figure Page 1575 STrOmbina SACO EAS ау. 13 1-3. NMB H 17019; from NMB locality 15912: Rio Gurabo; upper part of Cercado Formation. Note long sinusigerous outer lip of protoconch; 1. x30; 2. x60; 3. x120. 4. NMB H 17022; from NMB locality 15914: Rio Gurabo; upper part of Cercado Formation.Very slightly oblique apical view; x60. 5-7. NMB H 17021; from NMB locality 15912: Río Gurabo; upper part of Cercado Formation. Note sinusigerous outer lip of protoconch which is somewhat shorter than that of the specimen shown in figures 1-3; 5. x 30; 6. x60; 7. x120. 8-16... Strombinal(Sincola)) pseudonaitensisi(Mauty) e cree ec ee 24502222222 аа. 16 8. NMBH 17027; from NMB locality 16913: Río Mao; Cercado Formation. Very slightly oblique apical view; x60. 9-11. ММВ Н 17028; from ММВ locality 16913: Río Mao; Cercado Formation; 9. x12; 10. x30; 11. x60. 12-13. NMB H 17029; from NMB locality 16913: Río Mao; Cercado Formation; 12. x30; 13. x60. 14-16. NMB H 17030; from NMB locality 16927: Río Mao; Cercado Formation. Note sinusigerous outer lip of protoconch; oblique apical views; 14. x35; 15. x70; 16. x140. 32 BULLETIN 324 EXPLANATION OF PLATE 7 Figure Page 1-12: 3StrombinaSmeola):caribaen аа 18 1-3. Holotype of S. caribaea micra Pilsbry, ANSP 2800; precise locality in Dominican Republic not known; 1. front view; 2. rear view; 3. from right side. height: 5.8 mm; width: 2.7 mm. 4—6. Lectotype of 5. caribaea, ANSP 3996; precise locality in Dominican Republic not known. Specimen figured by Pilsbry (1922, pl. 18, fig. 9); 4. front view; 5. rear view; 6. from right side. height: 10.2 mm; width: 5.2 mm. 7-9. NMB H 17081; from NMB locality 17288: Rio Yaque del Norte (López section); Baitoa Formation. Immature specimen corresponding to S. caribaea micra; 7. front view; 8. rear view; 9. from right side. height: 6.1 mm; width: 2.9 mm. 10-12. ММВ H 17078; from ММВ locality 16942: Río Yaque del Norte (López section); Baitoa Formation; 10. front view; 11. rear view; 12. from right side. height: 9.1 mm; width: 4.8 mm. BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 7 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 8 1 х 30 2 х 60 3 х 125 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 33 EXPLANATION OF PLATE 8 Figure Page 1-7. Strombina (Sincola) caribaea Gabb ....................................................2.............42.1.2г.124лй......-. 18 1-3. NMB H 17032; from ММВ locality 16936: Rio Yaque del Norte (López section); Baitoa Formation. Note sinusigerous outer lip of protoconch; 1. x30; 2. x60; 3. х125. 4-5. NMB H 17033; from ММВ locality 17288: Río Yaque del Norte (López section); Вайоа Formation; 4. x30; 5. x 60. 6-7. NMB H 17034; from ММВ locality 17288: Rio Yaque del Norte (López section); Baitoa Formation. Note sinusigerous outer lip of protoconch; 6. oblique apical view, x 70; 7. apical view, x 70. 8-14. Strombina(Sincbla)sbassulMaury)i а 19 8-9. NMB Н 17090; from ММВ locality 16856: Río Cana; Cercado Formation; 8. х 35; 9. x70. 10-11. NMB H 17091; from NMB locality 16856: Rio Cana; Cercado Formation; 10. x30; 11. note sinusigerous outer lip of protoconch, x65. 12-14. NMB H 17092; from NMB locality 16856: Rio Cana; Cercado Formation. Note sinusigerous outer lip of protoconch; 12. oblique apical view, x 70; 13. x140; 14. apical view, x70. 34 BULLETIN 324 EXPLANATION OF PLATE 9 Figure Page 1-6, DI Оба 19 1-3. Holotype, PRI 28736; Río Gurabo at Los Quemados; Maury's Zones D and E; 1. front view; 2. rear view; 3. from right side. height: 11.0 mm; width: 5.3 mm. 4-6. ММВ Н 17087; from ММВ locality 16857: Rio Cana; Cercado Formation; 4. front view; 5. rear view; 6. from right side. height: 12.2 mm; width: 6.2 mm. BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 9 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 10 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 35 EXPLANATION OF PLATE 10 Figure Page 1-13 x StrombinaSiicola)manniebellae (Maury), ee 20 1-3. Lectotype, PRI 28735; Rio Cana at Caimito, Maury's Zones H and І; specimen figured by Maury (19172, pl. 15, fig. 15); 1. front view; 2. rear view; 3. from right side. height: 12.4 mm; width: 6.8 mm. 4-10. Front views of seven specimens showing variability of sculpture on early spire whorls, development of outer lip, and development of denticles on its inner surface; all from NMB locality 16856: Río Cana; Cercado Formation; 4. NMB H 17047. height: 10.9 mm; width: 6.6 mm; 5. NMB H 17048. height: 11.2 mm; width: 6.9 mm; 6. NMB H 17049. height: 11.2 mm; width: 6.6 mm; 7. ММВ Н 17050. height: 11.9 mm; width: 7.2 mm; 8. ММВ Н 17051. height: 12.8 mm; width: 7.2 mm; 9. NMB H 17052. height: 10.6 mm; width: 6.2 mm; 10. NMB H 17053. height: 12.2 mm; width: 6.9 mm. 11-13. NMB H 17054; from NMB locality 16844: Río Cana; Cercado Formation; 11. front view; 12. from right side; 13. rear view. height: 10.8 mm; width: 6.5 mm. 36 BULLETIN 324 EXPLANATION OF PLATE 11 Figure Page 1-9. Strombina (Sincola) nunniebellae Машат)... ке 20 1-3. ММВН 17044; from ММВ locality 16856: Rio Cana; Cercado Formation. Note sinusigerous outer Ир of protoconch; 1. 23072 60: 3. x 120: 4-6. NMB H 17045; from NMB locality 16856: Rio Cana; Cercado Formation; 4. oblique apical view; note sinusigerous outer lip of protoconch, x65; 5. apical view, X 70; 6. apical view, х 140. 7. NMB H 17058; from NMB locality 16854: Rio Cana; Cercado Formation. Note sinusigerous outer lip of protoconch, x 140. 8-9. NMB H 17056; from NMB locality 16839: Río Cana; Cercado Formation; 8. oblique apical view; note sinusigerous outer lip of protoconch, x 70; 9. apical view, x65. 10-12. Strombina (Bifurcium) nuestrasenorae (Maury) .................................................... 66.61 152222222... 22 10-11. NMB H 17036; from NMB locality 15873: Río Gurabo; Gurabo Formation; 10. x60; 11. posterior canal, x30. 12. NMB H 17035; from ММВ locality 15873: Río Gurabo; Gurabo Formation; apical view, х 60. BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 11 2 x60 3 x120 6 x 140 7 х 140 11 x30 BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 12 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 37 EXPLANATION OF PLATE 12 Figure Page 1-92 Strombina CBifürciuummiestrasenorae (Many a о ыу ы ы MEE Ea A 22 1-3. NMB H 17083; from NMB locality 15873: Río Gurabo; Gurabo Formation; 1. front view; 2. rear view; 3. from right side. height: 9.6 mm; width: 5.4 mm. 4-6. Holotype, PRI 28732; Río Gurabo at Los Quemados; Maury's Zone G; 4. front view; 5. rear view; 6. from right side. height: 7.8 mm; width: 4.2 mm. 7-9. NMB H 17085; from NMB locality 16857: Río Cana; Cercado Formation; 7. front view; 8. rear view; 9. from right side. height: 10.0 mm; width: 5.6 mm. 38 Figure 1-3. 4-9. 10-11. BULLETIN 324 EXPLANATION OF PLATE 13 Page Strombina (By urcmm) ай: nuestrasenorae (Maury). era A ee een 22 NMB Н 17086; from NMB locality 16856: Rio Cana; Cercado Formation; 1. front view; 2. rear view; 3. from right side. height: 9.8 mm; width: 6.6 mm. Strombma (Винт) bicanalifera (G, B, Sowerby D ТМ. u ae 77772727 22 Syntype, ВММН 1966312; Recent, Galapagos Islands; 4–6. surface not whitened; 7-9. surface whitened; 4, 7. front views; 5, 8. rear views; 6, 9. from right side. height: 12.3 mm; width: 6.1 mm. 23 РГО ПО (BUNCH) СИЛИ ПЕ ISDE 6525 200 2, reso Saks en А einer e, Paratype, ММВ Н 17038; from ММВ locality 16837: Rio Cana; upper part of Cercado Formation; 10. oblique apical view, х 70; 11. apical view, x65. BULLETINS OF AMERICAN PALEONTOLOGY, VOLUME 90 PLATE 13 PLATE 14 о с ш = = E O > © © © = O = Z б [53] - < 29 2 < Е z Щщ = < ш O N 2 E ш 4 d 2 m DOMINICAN REPUBLIC NEOGENE. 2: JUNG 39 EXPLANATION OF PLATE 14 Figure Page 1-9. Strombina:(Bifurcium) canae; now: species et on ee онен nenn EIE TT АЫ ОЕ 23 1-3. Holotype, NMB H 17041; from NMB locality 16837: Rio Cana; upper part of Cercado Formation; 1. front view; 2. rear view; 3. from right side. height: 8.1 mm; width: 4.5 mm. 4—6. Paratype, ММВ H 17040; from ММВ locality 16837: Rio Cana; upper part of Cercado Formation; 4. front view; 5. rear view; 6. from right side. height: 8.2 mm; width: 4.4 mm. 7-9. Paratype, ММВ H 17043; from ММВ locality 15842: Rio Gurabo; Gurabo Formation; 7. front view; 8. rear view; 9. from right side. height: 9.4 mm; width: 5.3 mm. 40 BULLETIN 324 INDEX Note: Page numbers are in light face, plate numbers are in bold face type; principal discussion pages are in italics. ambigua, СОЛИЛВЕЛЕ углаас СМ га a ain si vow ceeds bse 13 Columbella (SWOMIDING) nee ee seen 13 SU OWIDDIBE san en ee erstes 15 Angostura: Formation: cc, ыы сна ДЕТ АІ anomala, Strombina: (тео. 15 ANSP (Academy of Natural Sciences, Philadelphia, PA) ..... 8,11, 12,18 Ataya РЕША анын, 10 Arca patricia G. By Sowerby TE 18802. 2. nn men. 8,20 ATIOVO А. 6:29 Da dao Ы ы ООА 1122 Ballon na... о өй 6,18,19 o e cse oce een 6,19 Bua Cai ENA A o doces c 2227 ee 14 bassi, о (TOT IDIO).... cou эши Uadec Уг 22077102 19 Strombina (51100103 tiritas гь. 80-222 6,18,19,20 bassi (alle); Ола а Ион OSIACOLO) „а. „оаа 20 bicanalifera, ОСИ E 21 Sirombina UB ата ТТТ ee coats Denen ІЗ 8,22 БЛП ЖОБА 1901 т 222202 2! ПРИ ОЛИ ОО НТ 222: 22 ВЕСЕ ЕССЕ ПӘ Цагтаа Эл лла 241222 Аа 2 МОК ЧЭ -21-82222-12222 о 9 о. 5 ВИ 2 АМЕН een. nn es ee 17 I: ved Eli Nucl а up MCN M то 16,17 BMNH (British Museum (Natural History), London, БЕВАИ у ааваа 8,22 И o a ТС a Cs Le Коса 14,18 Bowden a o о ОО е 13,14,18 Brows: and PUSbry СТ 11,12 а Үй зы ине нна N у шәл о В 10 Bucquoy, Dautzenberg, and Dollfus (1882) .......................... 10 Cabo: Blanco ares cn АА ren 10 caboblanquensis, Strombina (Strombina) 22 10 A т ен. 21 canae, Strombina (Bifurcium) ............ 13,14 .... 6,9,22,23,24 caribaea, ГОЮН: a он. 18 Strombina (Sincolay a... aan ТЕ лу 6,18,19 (42141220 404411: (44218840 421111217:10 аана, 18 Caribbean л ца provinte о 7 Carmen от 0 еттен оа ае Т? 13 Сашатар oa cio A ЕИ теье 13,14 Сау N О АНА АНИ л REL a OA 10 СОГАП ШЕ ан 16 ESTOS ВОИ un. ann 6,9,16,20-24 Choctawhatchee Formation... еее 20 Сараа Valley. nee A O 5.6 Colombia ion E AA Ucn БЕЛЕ омар СЕ 12 ЁО ORE CS а ы йена ce een 12 Trento СО deer NN RU ec ее ене EZ Columbella amorrua СТАН УД бб en. ee. ambigua Kiener, 1841 bicanalifera С. B. Sowerby I, 1832 ......... i gadu Guis OO 2025 2 2 lanceolata Gebesowenbyel, РОЗ, Columbella (Strombina) ambigua Guppy, 1866 bassi Maury, 1917а ............. s nt ae 19 nanniebellae Maury, 1917a 20 neustrasenorae Maury, 1917a 2/2) pseudohaitensis Maury, 1917a 16 pseudohaitensis gurabensis Maury, 1917а ......................... 15 (оопа таа ОО nee Aes 10,21 (05481012 ceo о UE eret 6,14,16 crassilabrum, оо Т-ға” 17 SIFOMBIR (SIN COLD) ono ыл орну Ыбы EE TS 17 (:1111141183 dana АЛ анна nt dese oil «+ 10 cunninghamcraigi, Strombina (Sincola) ШЫ 20 cyphonotus, Strombina (Strombina) ............. y A ЗАО 12 DA (ЧОТ ун АС а Е... 13 Daule Formation ne нь 12 Dominican Republic APOYO АЈА Дре case cement ee 4... 6,23 Бапоаыассем елен... 6,18,19 ВЇП САО МАСУ Әл... 17 BO ОЕ ен 16,17 СаО ана 21 Cercado IM merrer Se ен 16 Cibao Valle Е ннат 5,6 Topez e E eL eR eset Rods: 18,19 ПОБ (Олий а ааа O a 15:19:22 ТО CALI ota УМ са OT TOT 6,8,20-24 Вах. 6,8,14-16,19,20,22-24 рн. 6,16 Т0) Ука е 61У То о 6,18,19 О ДЫ ЕГУ белу е сү а еи 15 ТИСЕН VINE АСТ LL RN d [у T T 6 Easter PACIO e co ERE ean nc DROP T da 7,14 EGUAG Ole eret Е 10,12,14,17,21,22 Ealapaco а 10221222 dedans IBAI SRA a А 13,14 E KEO ENa A od 5 ооо Мах 13,14 Taleonensisy Strombma(Spiralta) zu... men 13,14 Fischer (1884) То OA North St. Petersburg OBIS АМ ооо ds 10 Гайса ОНОО (Отта en. Е. 10 СЛОВО АНТЫ 27 б ЭП? стајараров ІЗ (6 а. о ооа аи 10:21:22 PIPE TONES ISS IO MDI CNCA) S DESEE 18,19 DOMINICAN REPUBLIC NEOGENE. 2: JUNG 41 Gardner (O47) rere ea МО ae Gardner (194971 те ne Gardner and Aldrich (1919) .. za 28 15, САВУ о, EA tuae Эн о GatundFormauon «ooo ERR Un M аел ае Sa ба Stage оО ee Sete es gibberula, Strombina (Sincola) Gibson-Smith, Jeu did ня stem 10 Gibson-Smith and Gibson-Smith (1974) ......................... 10,14 Globoroialia margaritae Zone ORO DT OR 14 Golfo: dev иер на анау и ымын кы hom ode t eme 10 Gomez and Valenin DID e a r 14 gadaa бог г1а2 алты тет 11:12 adale SIYombinde x оме 11:12 Gram and Galo (ТЗ 22:12 12:20 Оа е 13,14 CUBE Rere oreari иене 6 ОШО от М ree e uu Ue 13,21 ПЕШ Soy oy ym spleen) As cct M Tes RU ee 2) gunteri leonensis, Strombina (5іпсоа).. Ашны лан 20 Guppy: Обо OG CES 11:12 CN лы еу E Ey guppyi, SANA ее 13 SIDRA SPD EM 2. З 6,13,14 gurabensis, Strombina (Sincola) .......... ЗА бен: 6,9,15,16,17 Сиаро Оо ОИ а cs 6,15,16,19,20,22-24 Наша айа Неше Ө? 22244202 A 13 HAV DISONGESS ОТІП ena ПИЕНО 17 АЛЕ ОО Mies е eerte rette bleed o э. 18 Eiodscnrandehlodson. OB ке ылас Ө E 13,14 ПРЕ ТИ DO а e SE EN 5 Pas Бов Код нов «irn eicere t NER, 10 dama Formation «ceste ette e ettet eee ES 22 Јаптатсани ент e cL eR 6,13,14 Bodensee EE NINH RTT. 14,18 JODE (1969) cit d акысын EUM ҚЗ т 17 Keen: О ӨЛ л ну ынды RR ERU 8,13,14,21 КЧепег (184 De ner s SIUE S 13 lampra; SINOMbDINAaUSINCOlA) e e ЛО Л К ОО ОЛОК ДЕ И 145) LANCEO GIA оштро ал AAEREN he 20022207: 10 MEAS етсен 555 осол ое. I2 ЁЄро (1045) oe с det nv NES SNR 8 lessepsiana, Strombina (Strombina) 222222209 12 ПОРАЗИ Отоа (исо) 20 Тори ENTRIES 18,19 Bopez sectione ice AER ner 6,18,19 Јов пета обилна eer 15:19:22 BostRoquessArchipblagor nn en an И 10 maculosa, ООО ee Strombina (Spiralta) .... Mansfield (1930) ............. Мој ЙАШ io OS АРАТА eae ee ee eh ies TO HE окы Marks (1951) ...... Маш етт Maury (19172) ..... 5,11,12,15,16,19-22 ЮКО ПОТЛЫ) у и ол оу 5,16,19 Mexico Baja California... laser ал 14 A RT а 13 Gulf of California 1324 IN кни тайн A oe EN 10 nanniebellae, GOLUMI DELIA SU OPI ee ee 20 ОН (СЯ 10242552 6,20,21 neustrasenorae, СӘК ИС (СОО 2222222 22 Strombina North: Carola. dte I li. ee 15 North SU Petersburg оо ие 20 nuestrasenorae, Strombina (Bifurcium) ......... JB PS AA 6,9,22, 23,24 nuestrasenorae (aff.), Strombina (Bifurcium) .......... а nds 22 не EDEN 5) (9155011:019:22) A ЛІ ee de ТЗ эла 14,16 (15501111 90425542 dad хаанай насан 14521 Olsson and Harbison: (1953) Бент с бе Аан 14,20 paceana, Strombina (Spiralta) Ае ОСО aerial at у кезе апаа апа ове en anne IMA VEA ете ae A es us сла Uc Lge oti Beake: № ut a a 13618210:141: Че а baron о, E қақы томар EE AT BEIUCHN еште ыл een EA НИС ПИОНЕР Салты ТА олы қыша іы ee ПИВО УРОТА ОО а ақы, IAL SUV a (UO ee Т е ое аа та отео ЈЕО БӘ ОП ИЯ ТО сн Pilsbry E ОЈ One A сата аа ПНА CVSS UCONN SUD Y Пон Blanktoltophielarvae sen nn RE ENS ee PRI (Paleontological Research Institution, Taca NIS USA name 8,15,16,19,20,22 prisma, Strombina (Strombina) pseudohaitensis, Columiela (ONOMA) nen a ig ole a 16 УТО ТРИ (SINCOLA) are SO и 6,9,15,16,17,21 pseudohaitensis gurabensis, Columbella (Strombina) ............. 15 BUSTO Colombia ее a гарснаа 12 DADES SAD ANS A ше. 10 Radwin (1977) RES О 10 Rio Banano Formation s оне 205605 6,8,20-24 БӨ Cana SOHO 2222-2222. 022. 22 6,20,21,23,24 КООШ АЙК Renee 6,8,14—16,19,20,22—24 ВО CURED ЗО ен 6,9,16,20,23,24 ПРОТА оу а а а 6,16,17 КТО МА ОДО О ее 6,9,17 Ка Е О. ен КӨСЕ M ee ЁС ОВА ТЕУ танана ЁС О еН 42 BULLETIN 324 Saunders, Jung, and Biju-Duval (1986) ............ 5,6, 14-19,21-24 Savaneta glauconitic sandstone Member of Springvale КОШПОЙ... ненае а сылды E ET cena SCAPDOLOURN сз ТТ МКТ А А Shoal River Formation cin oM AN ааа а sincola, Сиам АЛ НТ ауа ыы 14 ОИ О О о vector rane 16 MOOD AO lacie ы Не қалын TD LN сү. T5 БОХ р 0522) ыы ел RN 8,13,14,18,21,22 MOEN DY Als GES IO) A A аны М 5,8,20 SPEINEVAlE-FORHANONER aati Ино Л, 20 Savaneta glauconitic sandstone Member .......................... 20 З 01005210, потоа (SITOMDING) onen. ae 12 ЗОО 16902 „ен еке. к, И ен НЕ 10 амилсца Чарру, 860. .... COREL NARRA, ren COMICO ADO ек deme NUMEN E ME D TO CONDAL СНО РИ ру A922. 109.3 202: een {со 2112. GAMO 878 NAM Сны A Барга ОЧЕР, 1908. nennen maculosa (G. B. Sowerby I, 1832) ИНО аку rennen Strombina (Bifturcium) Fischer, 1884 5 4-21 5.7.9721 aff. nuestrasenorae 13 5 22 bicanalifera (С. B. Sowerby I, 1832) ................ 13. e 8,22 COAG AU ое с оо i314 5,6,9,22,23,24 nuestrasenorae (Maury, 1917) ......... 31,127 е 6,9,22,23,24 Strombina (Sincola) Olsson and Harbison, 1953 ......... 5,7-9,14, 17,18 A ЕЗ RE И 20 anomala (Gardner and Aldrich, 1919) ............................. 15 ШИ (МАШ LOUD) ее ВИ 6,8,19,20 COTVORCH GADD, 051002 Ж КЕКСЕ КӨТЕН NEE 6,18,19 TAN (04054 OY, 1974). A E nennen er ilg E a meraror (Rutsch, ЭД). 20 POWVESTONCTIONS, (TES 1899) nennen на er aterra езине. 18,19 20906): (G. B-Sowerby L 1832) ое 8,14,18 Barton: пав теј 1030 ее. 20 umeri leonensis Mansiteld, 193022222222... enone 20 gurabensis (Maury, 1917) ..................... 3,4,6 ...... 19 75.17 EY YO RADA A 1908 ann inne секер RINA А 1 ЦООР CAIN hes LOA o ena at ce tee ee 19 JOVIS in Pılsbiy and Brown, 1917 wann ee een 20 nanniebellae (Maury, 1917) ................... 10:11 2... 6,20,21 O A NEN 21 pseudohaitensis (Maury, 1917) ....... 0 з» ООВ ИСО SSO Me 1022 a РАИ И 14,16 SITOMUINA рата Нар n. SUDBen. е LS eeoa G. O da E EIL) i nee 13,14 falconensis Н. К. Hodson in Hodson and Hodson, 1931 ucl I MEE CU ОЕ ое 13,14 Strombina (Spiralta) maculosa (С. B. Sowerby I, 1832) ..................... ӘУЕ 8,13 равев та а 1946 атаманы А ОН N 15 Srombmma sp- ОЕР ЧОНУ LISI ee 10 Strombina (Strombina) Mörch, 1852 ........................... 9,10,13 caboblanguensis.Weisbord, 1962 2222.01. 00000 10 cyphonotus Pilsbry and Johnson, 1911 ...... У а SUIT, francesae J. Gibson-Smith in Gibson-Smith and Gibson- Sith Al As eee па tie ARR EOS CT 10 lessepsiana Brown апа Pilsbry, 1911 .................. Y RAUM 12 prisma Pilsbry and Johnson in Brown and Pilsbry, 1911 ТТІ 77-2 DES LOBEN 723 DUO (К еее зо). ао ehe 10 SIrtatocostatasN aus 1951. пера теку тата ие eem 12 Walton Gardner ПОД nenne ee ee 19 StrombocolumbusCossmann, 190222222222... 10 SUVs dl татран ао RD. NA 6 ЕВ acc trab RL Шораро лалы CE Trinidad TU (Tulane University, New Orleans, LA, U.S.A.) 1M ect UE Тел бир 8,17,20,21,23,24 ТИКО ы ко c A АН Сы 18 USNM (United States National Museum of Natural History, Washington DO ОВ 200501002004 8,14,17,18 Venezuela Ay ESAS 222220200000 10 САБО БАШ АНАУ да Ык оо 10 (ЕЕ ОЕ esee еб ee лк еөз тты 10 САША усе лау ук ст 10 AIG OT ы у AU хо у сло Та 13,14 (ОНО 11516 ео cc E Tu 10 ПЕЕ ОО ЕДИ ПЫ ааа о е 10 Los Roques Archipelago Ио) ее AES LEN 10 Blaya Santa оа ee dM INI ЭЛ 10 ОО 10 VOKOS 8 е тает ИТ УОЗ ое 11 Wallet Me 0000 от 5 wakoma, Stromibina (тоттан 12 Weisbords 929) an id ок 11,412 МЕТРОА ШОС) ceo ee ee o BR 10 О. 11 Уссат ОМА 13,14 Moodxring (19064 аар 10,12 PREPARATION OF MANUSCRIPTS Bulletins of American Paleontology usually comprises two or more sep- arate monographs in two volumes each year. 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