JOURNAL OF THE oo. ELPHIA BOTANICAL CLUB) THE SECOND ONE HUNDRED YEARS OM The Second One Hundred Years of Bartonia oe — a os ge oF eliminary Catalog of New Jersey Hornworts and Liverw ee wee reece eee shyt sss co BERRY BARRINGER teh es ee oe ee St ie Re eee sie BARTONIA Journal of the Philadelphia Botanical Club Since its founding in 1891, the Philadelphia Botanical Club has offered outstanding Programs, field trips, and other opportunities for those with an interest in plants to meet and exchange information. Monthl meetings (www.philbotclub.org/meeting.html) feature speakers from various botanical backgrounds. They are held at 7:30 p.m. on the fourth Thursday of the month in September, October, and January through May and the third Thursday in November and December, usually at the Marvin Comisky Conference Center, One Logan Square (one block east of the Academy of Natural Sciences of Philadelphia). From April to October, expert field botanists lead field trips in the mid-Atlantic region and occasionally elsewhere in North America or overseas (www.philbotclub.org/field_trips.html), Officers President: Robert J. Holt (2007-2009), Janet Novak (2009-2012) Other Members of the Executive Committee: Vice President: Janet Novak (2007-2009), Tatyana Livshultz (2009-2012) John C. O’Herron II, Ann F. Rhoads, Treasurer: Alfred E. Schuyler (2007-2010) David Hewi (2010-2012) William H. Roberts, Robert J. Holt 012) Corresponding Secretary: Dennis Gray (2007-2009), Alina Freire-Fierro (2009-2012) Editor, Bartonia: Gerry Moore (2007-2011), Recording Secretary: Pat Gordon (2007-2008), G. Russell Juelg (2008-2012) Sasha Eisenman (2011-2012) Appointees Herbarium Curator: Joseph Arsenault (2007-2012) Endowment Fund Director: John C. O’Herron II (2007-2012) Program Committee Chair: Ruth Dorn (2007-2008), Environmental Committee Chair: Ann F. Rhoads (2007-2012) G. Russell Juelg (2008-2012) Constitution Committee Chair: William H. Roberts (2007-2012 Field Trip Coordinator: Janet Ebert (2007-2011), David Lauer (2011-2012) Herbarium Committee Chair: Ted Gordon (2007-2012) Bartonia, in publication since 1909, was named for William P. C, Barton (1786-1856), Professor of Botany at the University of Pennsylvania and author of the first local flora (1818), Compendium Florae Philadelphicae (and nephew of physician and botanist Benjamin Smith Barton, who was a mentor of the explorer Meriwether Lewis). The journal began as an annual abstract of the Club’s proceedings with short articles on the plants of the Philadel- phia area. Its scope has broadened to encompass original research in plant systematics, plant ecology, and plant conservation biology with articles on floristics, distribution, methods, biography, bibliography, history of botani- cal exploration, and other topics of botanical interest ranging throughout—and well beyond—the mid-Atlantic region. Editor: Gerry Moore Corresponding Editor: James Lendemer, The New York Botanical Garden, Bronx, New York Associate Editors: Timothy A. Block—Director, Pennsylvania Flora Project, Morris Arboretum, University of Pennsylvania, Philadelphia John Dighton—Director, Pinelands Field Station and Professor, Rutgers University, New Lisbon, New Jersey Wayne R. Ferren J i P.A., Red Bank, New Jersey Carl S. Keener—Professor Emeritus of Biology, Pennsylvania State University, University Park Robert F.C. Naczi— The New York Botanical Garden, Bronx, New Yo Emily W. B. (Russell) Southgate—Faculty Member, Notre Dame Academy, Middleburg, Virginia A subscription to Bartonia comes with Club members} ip ( 1 dues are $20.00 for regular members, $12.00 for full-time students) and th $25.00 per issue for non-members and institutions. The members ip year is from 1 January through the following 31 December. The Club is exempt from federal i der section 501(c)(3) sages oe or of the Internal Revenue Cod t are tax deductible Membership/subscription correspondence: Manuscript submission: Subscription Manager Sasha W. Eisenman, Editor, Bartonia The Philadelphia Botanical Club Temple University Dept. of Landscape Architecture and Botany Department, Academy of Natural Sciences Horticulture 1900 Benjamin Franklin Parkway 201 Dixon Hall Philadelphia, PA 19103-1195 580 Meeti Bartonia No. 65: 1, 2011 The Second One Hundred Years of Bartonia The first issue of Bartonia was published in February 1909. In an “Introductory,” we learn that the title was chosen to honor Prof. Wm. P. C. Barton, “Professor of Botany in the University of Pennsylvania, 1815, and author of the first local flora relating to this vicinity, the COMPENDIUM FLORAE PHILADELPHICAE, published in 1818.” Early papers in Bartonia were focused on local botany in southeastern Pennsylvania, southern New Jersey, northern Delaware, and northern Maryland. In 1924, Francis W. Pennell succeeded Stewardson Brown as the second editor of Bartonia, a position he held from 1924 (No. 8) to 1949 (No. 25). In Bartonia No. 9 (1926), Pennell published a paper on Benjamin Smith Barton and his herbarium, which was his first of many on people who made important contributions to botany. These included Lewis David von Schweinitz (No. 16), Elias Durand (No. 17), Thomas Nuttall (No. 18), Witmer Stone (No. 20), and Constantine Samuel Rafinesque (Nos. 23 and 25). Edgar T. Wherry edited Bartonia from 1952 (No. 26) to 1970 (No. 39). He authored numerous papers between 1924 and 1978, mostly on Polemoniaceae and Phlox. In later issues he published check-lists of plants in Delaware (No. 37), Philadelphia (No. 38), Montgomery (No. 41), and Bucks (No. 43) counties, Pennsylvania, and one with Otway Brown for Cape May County, New Jersey (No. 40). Alfred E. Schuyler was editor of Bartonia from 1971 (No. 40) to 1996 (No. 59), Roger Latham from 2000 (No. 60) to 2007 (No. 63), and Gerry Moore from 2009 (No. 64) to the present. The current issue (No. 65) begins the second one hundred years. Sasha Eisenman will be the editor for the next issue (No. 66). Since the early 1970s the journal has included papers on diverse aspects of plant systematics and ecology that go beyond the original focus on local flora, although this subject area continues to be of interest today. ALFRED E. SCHUYLER Curator Emeritus of Botany Academy of Natural Sciences MISSOURI! BOTANICAL MAR 1 2 2012 GARDEN LIBRARY Bartonia No. 65: 2-19, 2011 A Preliminary Catalog of New Jersey Hornworts and iverworts KERRY BARRINGER Brooklyn Botanic Garden, 1000 Washington Ave., Brooklyn, New York 11225, USA kerrybarringer@bbg.org ABSTRACT. Based on a survey of herbarium collections, three species of hornworts (Anthocerotophyta) and ninety-three species of liverworts (Marchantiophyta) are know to have grown in New Jersey. More than half of these records are based on collections more than 50 years old. Eighteen species of liverworts are potentially rare. INTRODUCTION Liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) have been largely ignored in studies of the plant diversity of New Jersey. Though they are widespread and sometimes abundant, they are usually overlooked by collectors. Most specimens in herbaria were collected before 1950 and many genera have not been sampled for more than 100 years. The only lists for the state are also more than 100 years old (Parker 1881, Rau 1889), and Formann (1998) mentions only a few species occurring in the pine barrens. We have almost no understanding of the current status and distribution of these plants in the state. To gain a better understanding of liverworts and hornworts in New Jersey, there is a need for new collections of these plants throughout the state. Both liverworts and hornworts, like mosses, can be conveniently collected into envelopes or packets (Hicks 1992). They should be sought in wet to moist habitats, especially along stream corridors and on fallen, rotting branches and trunks that have lost their bark. They are frequent in all kinds of swamps and wetlands, and many interesting species can be found growing in wet portions of fallow fields (Crum 1991). Identifications are not difficult with fresh material and proper magnification. Most of the common species can be reliably identified even in sterile condition using Hicks (1992), Conard & Redfearn (1979), Crum (1991), and Lincoln (2008). For detailed information and keys to all of our species, Schuster (1966-1992) is unsurpassed. Within the next few years, a definitive treatment of the liverworts and hornworts will appear as part of the Flora North America series (Zander & Eckel 2007). METHODS The following checklist was compiled from specimens at the Academy of Natural Sciences, Philadelphia (PH), Brooklyn Botanic Garden (BKL), and the Chrysler Herbarium at Rutgers University (CHRB). In addition, database records of the specimens at the Lewis Anderson Herbarium of Duke University (DUKE), the University of Michigan Herbarium z A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS 3 (MICH), the Missouri Botanical Garden (MO) and the New York Botanical Garden (NY) were checked, as were the lists of representative specimens in Schuster (1966-1992). RESULTS Based on a survey of herbarium records, three species of hornworts in three genera and 93 species of liverworts in fifty genera are known from New Jersey (Appendix 1). For all of the hornwort species and 55 liverwort species (about 60%), the most recent collections were made before 1950. DISCUSSION With so few recent collections of hornworts and liverworts available, it is difficult to determine which species are truly rare in New Jersey. Vascular plant rarities tend be either plants that occupy very specific or uncommon habitats, or plants whose range has been severely limited by habitat change, or plants near their climatic or distributional limits (Breden, et al. 2006, Snyder 2006). If we apply these criteria, a few liverwort species stand out as potentially rare species for the state (Table 1). Table 1. Species that appear to be rare. Aneura pinguis Cenbaloviclix clachio Cephaloziella rubella iloscyphus minor Chiloscyphus pallescens Fossombronia brasiliensis Fossombronia foveolata Frullania inflata Herbertus aduncus Jubula pennsylvanica Lepidozia reptans Mannia fragrans Marsupella emarginata Mylia anomala Pellia epiphylla Preissia quadrata Scapania glaucocephala Tretnamare, orta Four species appear to be limited to cool sphagnum bogs similar to those found in extreme northern and northwestern New Jersey (Breden, et al. 2001). Cephaloziella elachista (J. B. Jack) Schiffn., C. rubella (Nees) Warnst., Mylia anomala (Hook.) Gray, and Scapania glaucocephala (Taylor) Austin were collected from cool bogs in Sussex, Passaic, and Bergen counties. All but C. rubella are limited to this habitat throughout their range, which tends to be northern, with an extension south along the Appalachians to Virginia. The historical records from Bergen County seem odd today, when no suitable sites are known, but other historical collections in the county, notably the historical occurrence of Cornus canadensis L., provide further evidence for the occurrence of cool bogs in Bergen County in the past. 4 BARTONIA Other species share the Appalachian distribution of these species, and they appear to be rare even though they are not limited to bogs, because they are at the eastern edge of their range. Chiloscyphus pallescens (Ehrh.) Dumort., Frullania riparia Hampe, Herbertus aduncus (Dicks.) Gray, Lepidozia reptans (L.) Dumort., Mannia fragrans (Balbis) Freye & Clark, Preissia quadrata (Scop.) Nees, and Tritomaria exsecta (Schmidel) Schiffner all occur in the Appalachian mountains and are found at the edge of their range in northwest New Jersey. Limestone endemics and calciphiles are usually rare in New Jersey because there is so little undisturbed habitat (Breden, et al. 2006). Aneura pinguis (L.) Dumort. and Preissia quadrata have been collected twice in New Jersey, and are known to be limited in the northeastern United States to wet, basic substrates, like wet limestone, which are uncommon in New Jersey (Schuster 1992). Two of the species of Fossombronia with historical records in the state, F brasiliensis Steph. and EF foveolata Lindb., prefer clay soils. The known collections were made on the inner coastal plain, which is known for its clay deposits (Breden, et al. 2001). This area has been subject to extreme population pressures and it is likely that habitat destruction has severely limited their distribution. CONCLUSION Further study and collecting will greatly expand our understanding of New Jersey’s liverworts and hornworts. It is very likely that there are more species in the state than are included in this list. For example, six species that have been collected on the Pennsylvania side of the Delaware Water Gap have not been found on the New Jersey side: Barbilophozia barbata (Schreb.) Loeske, Frullania brittoniae Evans, Metzgeria crassipilis (Lindb.) A. Evans, M. furcata (L.) Corda, Pellia endiviifolia (Dicks.) Dumort., and Tritomaria exsectiformis (Briedl.) Schiffner. Nevertheless, suitable habitat exists on the New Jersey side, so the proximity of good populations strongly suggests that they will be found there. Some species that are widespread in other states are known from only one or a few counties in New Jersey, so it is likely that further collecting will show that these species are more widespread. For example, Cephalozia catenulata (Huebn.) Lindb. appears to be limited to white cedar swamps on the coastal plain in New Jersey, based on the records we have now. However, Hicks (1992) points out that it also grows in inland cedar swamps and, therefore, should be searched for in northwestern New Jersey. Also, our only record of Cololejeunea biddlecomiae is from shaded ravines in Bergen county. South of New Jersey, this species grows on the coastal plain (Hicks 1992). Finally, we cannot be sure that many of the species listed here still occur in the state, because many have not been seen in more than 50 years. We have no idea how the rapid development in the state has affected the populations of these organisms, but it is likely that many are now rare or endangered (Breden, et al 2006). Only additional collecting and study can begin to answer the many questions still outstanding on the distribution and health of the hornwort and liverwort populations in the state. I hope that this list will make that work a little easier. ACKNOWLEDGMENTS I am grateful to the curators of CHRB, DUKE, MICH, MO, NY, and PH for allowing me access to their collections or to the data in their collections databases. I thank Dr. Walter A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS 5 Bien, of Drexel University, for sharing his knowledge of the liverworts of southern New Jersey and bryophytes in general. I also thank Mr. Basil Shanahan for his careful analysis of Austin’s Hepaticae Boreali-Americanae. I am grateful to Alina Freire-Fierro for her help during my visits to PH and Sasha Eisenman for his help at CHRB. I am also grateful to the editors and reviewers whose comments and suggestions helped me greatly improve the manuscript. LITERATURE CITED BREDEN, T. F, Y. ALGER, K. S$. WALZ, AND A. WINDISCH. 2001. Classification of vegetatio communities of New Jersey: EE Iteration. New Jersey Department of Pavitcninedtal Protection, Trenton, New Jersey. 230 pp. BREDEN, T. F., J. M. HARTMAN, M. ANZELONE, AND J. FE. KELLY. 2006. Endangered plant species populations in New Jersey: Health and Threats. New Jersey Department of Environmental Protection, Trenton, New Jersey. 194 pp. CONARD, H. S. AND P. REDFEARN, JR. 1979. How to Know the Mosses and Liverworts, revised ed. William C. Brown Co., page, lowa. 302 pp. CRUM, H. 1991. Liverworts and »f Southern Michigan. University of Michigan Herbarium Ann Arbor, Michigan. 233 pp. FORMAN, R. T. 'T. 1998. Common bryophytes and lichens of the New Jersey Pine Barrens. Pp. 407- 424 in R. T. T. Forman (ed.), Pine Barrens. Ecosystem and Landscape. Rutgers University Press, New Brunswick, New Jersey. 601 pp. Hicks, M. L. 1992. Guide to the Liverworts of North Carolina. Duke University Press, Durham, North Carolina. 248 LINCOLN, M. S. G. 2009. lees of New England. Memoirs of The New York Botanical Garden. 99: T-161, PARKER, C. F. 1881. Hepaticae. Pp. 154-159 im N. L. Britton (ed.), A Preliminary Catalogue of the Flora of New Jersey. New Jersey Geological Survey, New Brunswick, New Jersey. 233 pp. RAU, E. A. 1889. Class 3. — Hepaticae. Pp. 346-355 in N. L. Britton (ed.), Catalogue of plants found in New Jersey. Pp. 25-642 in G. H. Cook (ed.) Final din af ig ~_ Geologist. Volume II. The John L. Murphy Publishing Company, Trenton, New Jerse SCHUSTER, R. M. 1966, 1969, 1974. The Hepaticae and . of ‘North America. Volumes I, II, III. Columbia University Press, New York. SCHUSTER, R. M. 1980, 1992a, 1992b. The Hepaticae and Anthocerotae of North America. Volumes IV, V, VI. Field Museum of Natural History, Chicago. SNYDER, D. B. 2006. Special plants of New Jersey. New Jersey Department of Environmental Protection, Trenton, New Jersey. STOTLER, R. E., B. J. CANDALL-STOTLER 1977. A checklist of the liverworts and hornworts of North America. The Bryologist 80: 405-428 ZANDER, R. H. AND P. M. ECKEL. 2007. Preface for Volume 27. Pp. xiv-xv in Flora of North America ge! Committee (eds.), Flora of North America. Volume 27. Oxford University Press, New York. 734 pp. 6 BARTONIA APPENDIX CHECKLIST OF THE HORNWORTS AND LIVERWORTS OF NEW JERSEY Nomenclature generally follows Stotler & Crandall-Stotler (1977) unless a more recent treatment is available. If there has been any confusion, or a recent nomenclatural change, synonyms are cited. Accepted names are in boldface. When there is no field number for a collection, the collection date is cited. If the specimens are part of a numbered distribution of exsiccatae, the exsiccata number is cited. The two distributions cited are C. F. Austin’s Hepaticae Boreali-Americanae (Hep. Bor.-Amer.] and Caroline C. Haynes American Hepaticae [Amer. Hep.]. Because of the paucity of specimens for some species, habitat notes are drawn from the literature as well as from specimens labels. Hornworts (Anthocerotophyta) Anthoceros (Anthocerotaceae) hoceros agrestis Paton capsules: fall and early winter on dripping basalt cliffs, also reported from harvested cornfields (Schuster 1992b) Bergen: Wiegmann 1212 (BKL, N Monmouth: N. Britton s.n., 2 Oct 1886 (CHRB) Anthoceros laevis L. (see Phaeoceros laevis) Anthoceros punctatus L. (ours are Anthoceros agrestis) Aspiromitus (Anthocerotaceae Aspiromitus punctatus ssp. agrestis (Paton) R. M. Schust. (see Anthoceros agrestis) Notothylas (Notothyladaceae) Notothylas orbicularis (Schwein.) Sull. capsules: Aug-Nov on wet soil at the edges of corn fields, in ruts and trails Bergen: C. E Austin s.n., Oct 1863 (MO, NY) Hunterdon: E. Britton s.n., 28 Sep 1886 (CHRB) Sussex: Nearing s.n., 27 Sep 1947 (NY) Phaeoceros (Anthocerotaceae) Phaeoceros laevis (L.) Prosk. capsules: fall and early winter on wet soil in harvested cornfields Monmouth: Haynes 1638 [Amer. Hep. 20] (BKL, NY) Morris: N. Britton s.n., 21 Sep 1886 (CHRB Sussex: N. Britton s.n., 20 Sep 1887 (CHRB) Schuster (1992b) reports an Austin specimen from Bergen County. Li (Marchantiophyta) Aneura (Aneuraceae) Aneura pinguis (L.) Dumort. capsules: Mar-Ma wet limestone or traprock ledges and on mossy soil (Schuster 1992) Bergen: Austin s.n. [112 Hep. Bor.-Amer.] (BKL, NY) A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS Morris: R. Torrey s.n., 8 Jan 1922 (NY) Asterella (Aytoniaceae) Asterella tenella (L.) P. Beauv. capsules: Mar-Jun on soil-covered ledges on trap rock or limestone, usually dripping wet Bergen: M. A. Howe 16 (NY) Essex: Underwood s.n., 9 May 1897 (NY) Hudson: J. Torrey s.n. , s.d.(NY Somerset: Wiegmann 5002 (NY) Schuster (1992b) reports a specimen from Passaic County. Bazzania (Lepidoziaceae) Bazzania trilobata (L.) Gray on wet soil, rocks, decaying logs, or tree bases in moist forests, swamps Bergen: C. F Austin s.n. [79 Hep. Bor.-Amer.] (BKL, NY Burlington: H. Webster s.n., 27 Feb 1968 (CHRB) Cumberland: Long 1376 (PH) Gloucester: Long 1395 (PH) Hunterdon: Cantlon & Moul 1001 (CHRB) Monmouth: Cantlon & M. Buell s.n., 13 Apr 1949 (CHRB) Ocean: Long 2232 (PH) Passaic: Barringer 20493 (BKL) Salem: Long 995 (PH) Sussex: Barringer 20623 (BKL) Blasia (Blasiaceae) Blasia pusilla : capsules: sp wet, pea ore ditches, rarely on rocks in streambeds Bergen: Wiegmann s.n., 12 Oct 1916 (NY) Essex: Haynes s.n., 10 May 1902 (NY) Monmouth: M. A. Howe s.n., 15 Jun 1918 (NY) Morris: Britton s.n., 21 Sep 1886 (CHRB) Sussex: Lawton 544 (NY) Warren: Henley s.n., 1940 (NY) Blepharostoma (Pseudolepicoleaceae) Blepharostoma trichophyllum (L.) Dumort. usually — with bryophytes or on moist, shaded rocks in tains (Hicks 1992) Atlantic: M. Robertson 7206 (PH) Badionee: Thiers 1254 (NY) Cumberland: Thiers 1255 (NY) Gloucester: Rau s.n., Jul 1882 (NY) Ocean: Matzke s.n., Oct 1933 (NY) Union: Barringer 20532 (BKL) Calypogeia (Calypogeiaceae) Calypogeia fissa (L.) Raddi on various moist substrates in shaded woods, esp. near streams Bergen: Beals s.n., 28 Nov 1946 Monmouth: Haynes s.n., 28 May 1908 (NY, PH) Passaic: Barringer 20503 (BKL) Somerset: Stevenson s.n., 27 Mar 1962 (CHRB) Sussex: Barringer 20630 (BKL) Union: Barringer 20528 (BKL) 8 BARTONIA Calypogeia muelleriana (Schiffner) Mull. Frib. s on soil of moist streambank Hudson: Austin s.n., 1862 (NY) Monmouth: Barringer & D. Barringer 21594 (BKL) Passaic: Barringer 22356 (BKL) Sussex: Barringer 21003 (BKL) Union: Barringer 21485 (BKL) Warren: Barringer 22091 (BKL) Calypogeia sphagnicola (Arnell & J. Perss.) Warnst. & Loeske peat bogs and Sphagnum stands Burlington: Thiers 1249 (NY) Camden: Barringer 20826 (BKL) Cape May: Wonderly 818 (NY) Gloucester: Rau s.n., Jun 1882 (NY) Middlesex: Moul 6995 (CHRB) Monmouth: Haynes 1123 (BKL, MO, NY) Calypogeia sullivantii Austin on soil or acid rocks in shade Bergen: Austin [74b Hep. Bor.-Amer.] (BKL, NY) Burlington: Evans s.n., 12 Aug 1892 (NY) Somerset: Stevenson s.n., 19 Mar 1962 (CHRB) Calypogeia trichomanis (L.) Corda (ours are C. muelleriana or C. fissa) Cephalozia (Cephaloziaceae) Cephalozia bicuspidata (L.) Dumort. on shaded, wet, usually peaty soils along stream and trails, rarely on acidic rocks (Schuster 1974) Bergen: Wurdack & Cowan 7 (NY) Gloucester: Barringer 21205 (BKL) Hunterdon: Barringer 20752 (BKL Monmouth: Barringer & D. Barringer 21592 (BKL) Ocean: Gilly s.n., 25 Oct 1941 (NY) Sussex: Barringer 20627 (BKL) Cephalozia catenulata (Huebner) Lindb. on decaying logs, Chamaecyparis swamps Bergen: Howe s.n., 22 Sep 1898 (NY) Burlington: Evans s.n., 12 Aug 1892 (NY) Monmouth: Howe s.n., 15 Jun 1918 (NY) Ocean: Torrey Botanical Club s.n., 29 May 1896 (NY) Cephalozia connivens (Dicks.) Lindb. on soil in swamps and bogs with Sphagnum Atlantic: Long 1984 (PH) Burlington: Barringer 20696 (BKL) Camden: Barringer 20846 (BKL) Gloucester: Rau s.n., Jun 1882 (NY) Mercer: Long 1885 (PH) Monmouth: Barringer & D. Barringer 21599 (BKL) Ocean: Barringer 20921 (BKL) Cephalozia fluitans (Nees) Spruce (see Cladopodiella fluitans) Cephalozia lunulifolia (Dumort.) Dumort. on peaty soils, rotting wood and in bogs and swamps with Sphagnum Bergen: Barringer 22301 (BKL) A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS Burlington: Underwood s.n., s.d. (NY) amden: Barringer 20881 (BKL) Gloucester: Barringer 21230 (BKL) Ocean: Matzke s.n, s.d.. (NY) Sussex: Barringer 21906 (BKL) Cephaloziella (Cephaloziellaceae) Cephaloziella divaricata (Sm.) Warnst. on soil in woods Bergen: Howe s.n, s.d.. (NY) Burlington: Evans s.n., s.d. (NY) Ocean: Matzke s.n., 1 Nov 1947 (NY) Cephaloziella elachista (J. B. Jack) Schiffner in Sphagnum bogs often with Mylia Bergen: Austin [54 Hep. Bor.-Amer.] (BKL, NY) Cephaloziella rubella (Nees) Warnst. on peaty soil, wet roadbanks, and sometimes on rotting wood Sussex: — 20595 (BKL) n: Barringer 20985 (BKL) Chiseaaea boc Chiloscyphus cuspidatus (Nees) J. Engel & R. M. Schust. on Be — along streams in uplands n: Barringer 21218 (BKL) PP tse minor (Nees) J. J. Engel & R. M. Schust. on soil and tree bases in coastal plain swamps Burlington: Barringer 20700 (BKL) Monmouth: Barringer & D. Barringer 21590 (BKL) Passaic: C. Gilly s.n., 18 Oct 1941 (NY) Chiloscyphus pallescens (Ehrh.) Dumort. on moist soil and decorticated logs in swamps Passaic: Barkley s.n., 10 Oct 1958 (NY) Chiloscyphus polyanthos (L.) Corda ergen: Matzke s.n., 10 Apr 1947 (NY) Burlington: TP James s.n., Jul 1851 (NY) Chiloscyphus profundus (Nees) J. J. Engel & R. M. Schust. on soil and tree bases, occasionally on rotting wood Atlantic: Long 2036 (PH) Bergen: Wiegmann 5008 (NY) Burlington: Burlington 20682 (BKL) Camden: Burlington 20851 (BKL) Cumberland: M. Robertson 3145 (PH) Essex: Haynes s.n., 10 May 1902 (NY) Gloucester: Long 1926 (PH) Hunterdon: Barringer 20797 (BKL) Mercer: Conard 0-1521 ( Middlesex: E. Britton s.n., 19 Apr 1884 (CHRB, NY) Monmouth: Barringer & D. Barringer 21591 (BKL) Morris: Barringer 20115 (BKL) Ocean: E. Britton s.n., 28 May 1887 (CHRB) Passaic: Barringer 20603 (BKL) Salem: Long 2117 (PH) Sussex: Barringer 20586 (BKL) 10 BARTONIA Union: Barringer 20096 (BKL) rren: Barringer 21231 (BKL) Chiloscyphus rivularis (Schrad.) Hazsl. (ours are C. polyanthos) Cladopodiella (Cephaloziaceae) Cladopodiella fluitans (Nees) Jorg. peat bogs, in sun (Schuster 1974) Bergen: Austin [35 Hep. Bor.-Amer.] (BKL) Burlington: Leonard 6395 (NY) Ocean: Allen 16892 (MO) Cololejeunea (Lejueneaceae) Cololejeunea biddlecomiae (Austin) A. Evans on tree bark, rarely on rocks in shaded ravines Bergen: Howe s.n., 5 Nov 1898 (NY) Sussex: Barringer 22359 (BKL) Conocephalum (Conocephalaceae) Conocephalum conicum (L.) Underw. (not in NJ, see C. salebrosum) Conocephalum salebrosum Szweyk., Buczkowska and Ordzyoski on soil or rocks along streams and in swamps Bergen: Wiegmann 5005 (NY) Essex: Haynes s.n., 10 May 1902 (NY) Hunterdon: Barringer 20743 (BKL) Mercer: Long 1886 (PH) Monmouth: Haynes 12 (NY) Morris: Barringer 18238 (BKL) Ocean: Barringer 22149 (BKL) Passaic: Barkley 36 (NY) Sussex: Barringer 18056 (BKL) en: Barringer 21226 (BKL) Diplophyllum (Scapaniaceae) Diplophyllum apiculatum (A. Evans) Stephani on peaty soil on roadbanks, thin soil over rocks, steep roadcuts Bergen: Howe s.n., 6 Oct 1898 Monmouth: Barringer & D. Barringer 21606 (BKL) sete Barringer 21445 (BKL) arren: Barringer & D. a 21 ste (BKL) fesle e obtusifolium (Hook.) Du Bergen: Austin [20 Hep. Bor. pee (NY) Fossombronia (Fossombroniaceae) Fossombronia brasiliensis Stephani moist, exposed clay of ditches and old fields (Schuster 1992) no county designated: Austin s.n, s.d. (NY) Fossombronia cristula Austin (see FE. foveolata) Fossombronia foveolata Lindb. on exposed clay and moist sand (Schuster 1992) Burlington: Austin [121 Hep. Bor.-Amer.] (BKL, NY) Monmouth: Haynes s.n., s.d. (BKL) Frullania Qubulaceae) ia asagrayana Mont. on bark and rock (Schuster 1992) Atlantic: H. A. Green s.n., 1882 (NY) Bergen: Austin s.n., 1861 (NY) A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS Gloucester: Rau s.n., 1882 (NY) Frullania eboracensis Gottsche on tree bark, especially maples, birches, poplars Bergen:: Austin s.n., 17 Jun 1865 (BKL, NY) Burlington: Barringer 20900a (BKL) Cape May: Long 2012 (PH Cumberland: Long 854 (PH) Essex: Haynes 510 (NY) Gloucester: Long 918 (PH Mercer: M. Barbun s.n., Jul 1900 (PH) Middlesex: Barringer 21764 (BKL) Monmouth: Barringer & D. Barringer 21606 (BKL) Morris: Barringer & Zielinski 21667 (BKL) Ocean: Copeland 660 (NY) Passaic: Barringer 21468 (BKL) Salem: Long 927a (PH) Sussex: Barringer 20327b (BKL) Warren: Barringer 20986 (BKL) Frullania ericoides (Nees) Mont. on rocks and tree bark Bergen: Austin s.n., Sep 1867 (NY) Burlington: M. Haines s.n., s.d. (NY) } he on dry rock walls, often traprock (Schuster 1992) Bergen: Austin [104 Hep Bor.-Amer.] (BKL, NY, PH) Frullania plana Sull. calciphile, on dry, exposed rocks (Schuster 1992) Bergen: Austin [102 Hep. Bor.-Amer.] (BKL, NY, PH) Frullania riparia Hampe On vertical faces of shaded rock (Schuster 1992) Bergen: Matzke s.n., s.d. (NY) Schuster (1992) reports this species from Ogdensburg in Sussex County. Frullania tamarisci (L.) Dumort. (see F asagrayana) Frullania virginica Gottsche (see F. eboracensis) Geocalyx (Geocalycaceae) Geocalyx graveolens (Schrad.) Nees on rotting wood or humus in conifer woods, usually mixed with bryophytes Ocean: Matzke s.n., 1 Nov 1947 (NY) Gymnocolea (Jungermanniaceae) Gymnocolea inflata (Huds.) Dumort. on soil in Sphagnum bogs, swamps, occasionally on wet rock faces Bergen: Austin [34 Hep. Bor.-Amer.] (NY, PH) Burlington: Thiers 1253 (NY) Ocean: E. Britton s.n., 30 Jun 1900 (NY) Passaic: Nearing s.n., 15 Dec 1946 (NY) Herbertus (Herbertaceae) Herb aduncus (Dicks.) Gray on rocks and trees in conifer woods, in gorges on rock underledges Passaic: Underwood 52a (NY) Jamesoniella (Jungermanniaceae) Jamesoniella autumnalis (DC.) Stephani 11 12 BARTONIA coastal plain swamps or black spruce swamps, on soil or rotting logs (Hicks 1992) Bergen: Howe s.n., 22 Sep 1898 (NY) Cumberland: Long 853 (PH) Sussex: Buser 12033 (NY) Jubula (Jubulaceae) Jubula pennsylvanica (Stephani) A. Evans on wet rocks along streams in shaded ravines (Hicks 1992, Schuster 1992) Bergen Matzke s.n., Oct 1931 (NY Schuster (1992) reports a specimen from Gloucester County. Jungermannia (Jungermanniaceae) oe biformis Austin (see J. hyalina) ungermannia crenuliformis Austin on acid rocks along shaded stream Bergen: Austin [31 Hep. ok Asaei) (BKL, NY) Burlington: Moul 7982 (CHRB) Hunterdon: Cantlon 1024 (CHRB) Middlesex: Moul 11052 (CHRB) Monmouth: Haynes 6626 (MO) Ocean: E. Moul 2667 (CHRB, NY) Jungermannia divaricata Sm. (see i divaricata) Jungermannia fossombronioides Aus thin soil over acid rocks, Espbcially tes streams (Hicks 1992) Bergen: Austin [32 Hep. Bor.-Amer.] (BKL, NY) Jungermannia gracillima Sm. on moist soils and acid rocks in woods Bergen: Barkley s.n., 3 Oct 1958 (NY) Burlington: Barringer 20693 (BKL) Monmouth: Haynes 809 (NY) Jungermannia hyalina Lyell on soil over rocks in deciduous woods . Austin [28 Hep. Bor.- Speist (BKL, NY) rren: Barringer 15988 (BK Jomeethlaise lanceolata L. (see J. look Jungermannia leiantha Grolle on wet soil, usually near streams and falls, on wet rocks in ravines Hunterdon: Barringer 20743b (BKL) Sussex: Nearing s.n., 27 Sep 1947 (NY) Union: Stephenson s.n., Mar 1962 (CHRB) Jungermannia laxa Lindb. (see Lophozia laxa) Jungermannia novae-caesareae A. Evans (see Lophozia capitata) Jungermannia pumi. on damp, shaded, cide rbeks along streams (Hicks 1992) Bergen: Austin [33 Hep. Bor.-Amer.] (NY) Kurzia (Lepidoziaceae) urzi (A. Evans) Grolle on peaty soil beside streams, sometimes mixed with bryophytes on logs Bergen: Cowan & Wurdack 22 (NY) Burlington: Barringer 20707 (BKL) Camden: Krout s.n., Aug 1914 (BKL, CHRB, PH) Gloucester: Raw s.n., 13 Jul 1882 (NY) Mercer: Long 1886 (PH) A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS Middlesex: Wiegmann 5001 (NY) Monmouth: Haynes s.n., 15 May 1910 (BKL, NY) Ocean: Cowan & Wurdack 135 (NY) Sussex: Barringer 21917 (BKL) Union: Rudolph & Rissanen s.n., 19 Jun 1949 (NY) Lejeunea (Lejeuneaceae) Lejeunea cavifolia (Ehrh.) Lindb. on bar Passaic: Barkley 215 (NY) Union: Rissanen & Rudolph s.n., 19 Jun 1949 (NY) Lepidozia (Lepidoziaceae) Lepidozia reptans (L.) Dumort on soil, tree bases, rotting wood (Hicks 1992) without county designation: Austin [75 Hep. Bor.-Amer.] (BKL, NY) Leucolejeunea (Lejeuneaceae) Leucolejeunea clypeata (Schwein.) A. Evans on trees, but in uplands can grow on rocks Cumberland: Theirs 1257 (NY) Gloucester: Barringer 21201 (BKL) Warren: Rau s.n., s.d. (NY) Lophocolea (Geocalycaceae) Lophocolea bidentata (L.) Dumort. (not in NJ, ours are Chiloscyphus profundus) Lophocolea heterophylla (Schrad.) Dumort. (see Chiloscyphus profundus) Lophocolea minor Nees (see Chiloscyphus minor) Lophozia (Jungermanniaceae) Lophozia bicrenata (Schmidel) Dumort. moist, acid, leached soils, abandoned farmland Bergen: Austin [40 Hep. Bor.-Amer.] (BKL, NY) Monmouth: Haynes 1490 (BKL, NY) Passaic: Cowan & Wurdack 95 (NY) Lophozia capitata (Hook.) Macoun moist, sandy barrens and dune swales Atlantic: Evans s.n., Apr 1893 (NY) Burlington: Evans s.n., Aug 1892 (NY) Lophozia laxa (Lindb.) Grolle peat bogs Bergen: Austin [46 Hep. Bor.-Amer.] (BKL, NY) Lunularia (Lunulariaceae) Lunularia cruciata (L.) Dumort. weedy in greenhouses Passaic: Nash s.n., 29 Nov 1893 (NY) Mannia (Aytoniaceae) Mannia barbifrons Shimuzu & S. Hatt. (not in NJ, ours are M. fragrans) Mannia fragrans (Balbis) Frye & L. Clark on rock or thin soil over siliceous rocks in sun (Schuster 1992b) Bergen: Austin s.n., May 1858 Warren: T: C. Porter s.n., 4 May 1869 (PH) Marchantia (Marchantiaceae) Marchantia polymorpha L. on soil or rocks in disturbed sites, a weed in greenhouses Atlantic: Moul 6138 (CHRB) 14 BARTONIA Burlington: Krout s.n., Jun 1907 (BKL) Camden: E. Gadsby s.n., 15 Jun 1910 (PH) Essex: Nash 910 (NY) Gloucester: Lippincott s.n., 3 Jun 1894 (PH) Hudson: Torrey s.n., s.d. (NY) Mercer: Stowell 2 (NY) Monmouth: Haynes [11 Amer. Hep.] (NY) Passaic: Nash 625 (NY Salem: Heritage s.n., 14 ee ey (CHRB) Somerset: Moul 10827 (CH Warren: Barringer 21210 ee Marsupella (Gymnomitriaceae) Marsupella emarginata (Ehrh.) Dumort. on wet rocks and thin soil along streams (Schuster 1974) Sussex: Barringer 21875 (BKL) Warren: Buser 12048 (NY) Metzgeria (Metzgeriaceae) Metzgeria conjugata Lin shaded rock outcrops ‘nd on trees in uplands Bergen: Wiegmann s.n., 9 July 1915 (NY) Hunterdon: Barringer 20742b (BKL) Sussex: Barringer 21549 (BKL) Union: Wynne 2788 (NY) Warren: Barringer 21217 (BKL) Mylia (Jungermanniaceae) Mylia anomala (Hook.) Gray in sphagnum bogs in uplands Bergen: Austin [25 Hep. Bor.-Amer.] (BKL, NY, PH) Sussex: Beals s.n., 11 Oct 1947 (NY) Nardia (Jungermanniaceae) Nardia geoscyphus (De Not.) Lindb. thin soil over acid rocks, sandy streambanks Bergen: Cowan & Wurdack 18 (NY) _— Evans s.n., Aug 1892 (NY) nmouth: Haynes 803 (BKL, PH) Nardia sat (Austin) Underw. (not in NJ, ours are N. geoscyphus) Nowellia (Cephaloziac aceae) “ llia n de , rotting logs, rarely on adjacent soil Bova Kuwahara 6954 (NY) Hunterdon: Barringer 22168 (BKL) Middlesex: Barringer 21763 (BKL) Morris: Barringer & Zielinski 21646 (BKL) Ocean: Barringer 22125 (BKL) Passaic: Barringer 20668 (BKL) Sussex: Barringer 20989 (BKL) Warren: Barringer 21230 (BKL) Odontoschisma (Cephaloziaceae) Odontoschisma denudatum (Nees) Dumort. A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS 15 on soil and rotting logs Atlantic: Long 208 (PH) Burlington: Barringer 21149 (BKL) Gloucester: Barringer 21197 (BKL) Monmouth: Haynes s.n., 29 Apr 1908 (BKL, NY) Ocean: Cowan & Wurdack a (NY) Cumberland: Thiers 1256 (NY Odontoschisma prostratum (Sw.) revi Bergen: Barringer 20280 (BKL) Burlington: Barringer 20697 (BKL) Camden : Barringer 20844 (BKL) Cape May: Long 2154 (PH) Cumberland: Long 1175 (PH) Gloucester: Long 2115 (NY, PH) Mercer: Long 1888 (PH) Monmouth: Barringer & D. Barringer 21595 (BKL) Ocean: Barringer 20920 (BKL) Passaic: Barringer 21475 (BKL) Salem: Long 941 (PH) ex: Barringer: 21090 (BKL) Pallavicinia (Pallaviciniaceae) Pallavicinia lyellii (Hook.) Gray on soil and bark in wet woods, swamps, cedar swamps, pond edges Atlantic: Underwood s.n., 6 Sep 1884 (NY) Bergen: Barringer 21730 (BKL) Burlington: Barringer 16714 (BKL) Camden: Barringer 20872 (BKL) Cumberland: Long 1253 (NY, PH) Gloucester: Barringer 21200 (BKL) Hudson: s.l. s.n., s.d. (NY) Middlesex: Barringer 21810 (BKL) Monmouth: Barringer & D. Barringer 21598 (BKL) Morris: Barringer 17793 (BKL) Ocean: Barringer 22110 (BKL) Passaic: Barringer 20612 (BKL) Sussex: Barringer 20616 (BKL) Pst Moul s.n., 16 Apr 1947 (PH) ren: Barringer 22082 (BKL) share (etncn llia epiphylla (L.) Corda on soil along streams and on the edges of lakes and ponds Bergen: Barkley s.n., 3 Oct 1958 (NY) Essex: Haynes s.n., 10 May 1902 (NY) Mercer: s.l. s.n., 27 Apr 1909 (CHRB) Monmouth: Barringer & D. Barringer 21598 (BKL) Morris: Barringer 18238 (BKL, CHRB Passaic: Barringer 21427 (BKL) Sussex: Barringer 21569 (BKL) Union: Rissanen s.n., 19 Jun 1941 (NY) 16 BARTONIA Warren: Barringer & D. Barringer 21844 (BKL) Pellia neesiana (Gottsche) Limpr. on wet soil, especially along streams (Schuster 1992) Bergen: Wurdack & Cowan 9 ( Schuster (1992) reports specimens from Camden and Burlington Counties. Plagiochila (Plagiochilaceae) Plagiochila porelloides (Torr.) Lindenb. on shaded rocks along streams Bergen : Wiegmann s.n., 19 Jul 1915 (NY) Burlington: Montaigne s.n., Apr 1851 (NY) Morris: Austin s.n., Aug 1867 (NY) Porella (Porellaceae) Porella pinnata L. on rocks along streams Hunterdon: Cantlon 1010 (CHRB) Morris: Barringer 18505 (BKL) Ocean: Barringer 22109 (BKL) Passaic: Barringer 18182 (BKL) Union: Barringer 18133 (BKL) Somerset: Barringer 18129 (BKL) Sussex: Barringer 20308 (BKL) uses : Barringer 18133 (BKL) n: Barringer 21211 (BKL) Porella orp (L.) Pfeiff. on trees or occasionally rocks in moist woodlands Atlantic: Long 2033 (NY) Bergen: Haynes s.n., 29 Nov 1902 (NY) Burlington: Stowell 1 (NY) Essex: Haynes s.n., 10 May 1902 (NY) Gloucester: Rau s.n., Jun 1882 (NY) Hudson: Brainerd s.n., s.d. (BKL) Hunterdon: Cantlon 1011 (CHRB) Mercer: Torrey s.n., 1831 (NY) Morris: Barringer 18505 (BKL) Ocean: Cowan & Wurdack 129 (NY) Passaic: Barringer 20607 (BKL) Warren: Barringer 21227 (BKL) Porella platyphylloidea (Schwein.) Lindb.] Hunterdon: Moul 6789 (CHRB) Ocean: Cowan & Wurdack 129 (NY) Sussex: H. Webster 170 (CHRB) Warren: Matzke s.n., 10 Nov 1946 (NY — (Marchantiaceae) eissia quadrata (Scop.) Nees val a rocks near running water, ravines, dripping cliffs (Hicks 1992, Lincoln 2008) lesex: Vail s.n., May 1891 (NY) Ptilidium (Ptilidiaceac) Ptilidium pulcherrimum (Weber) Hampe mi 1 *, ; 1 DY on fallen | z A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS Atlantic: Long 1996 (PH) Bergen: Howe s.n., 6 Oct 1898 (NY) Camden: Long 1527 (PH) Cumberland: Long 774 (PH) Hudson: Austin s.n., s.d. (NY Hunterdon: Cantlon & Moul 1022 (CHRB) Middlesex: Barringer 21784 (BKL) Morris: Barringer & Zielinski 21675 (BKL) Passaic: Barringer 20657 (BKL) Sussex: Barringer 21490 (BKL) Warren: Barringer 21250 (BKL) Radula (Radulaceae) Radula complanata (L.) Dumort. on tree bark and rock outcrops (Hicks 1992, Lincoln 2008) Bergen: Austin [86 Hep. Bor.-~Amer.] ( unterdon: Cantlon 1008 (CHRB) Radula obconica Sull. on rock outcrops or bark in deep shade along streams (Hicks 1992, Lincoln 2008) Bergen: Austin [88 Hep. Bor.-Amer.] (MO, NY) Reboulia (Aytoniaceae) Reboulia hemisphaerica (L.) Raddi on soil over rocks, humid sites; a calciphile, but not exclusively (Schuster 1992b) Bergen: Wiegmann 5018 (NY) Hudson: Torrey 3 ( Hunterdon: Barringer 20742 (BKL) Warren: Cowan & Wurdack 174 (NY) Riccardia (Aneuraceae) Riccardia chamaedryfolia (With.) Grolle on dripping rocks near streams (Schuster 1992) onmouth: Haynes [14 Amer. Hep.] (NY) Riccardia latifrons (Lindb. oe (ours are R. palmata) Riccardia multifida (L.) G in N on soil or os in ae seeps and near falls, in S on trees in cedar swamps (Schuster 1992) Bergen: Austin s.n., 1873 (NY) Burlington: Evans s.n., 10 Aug 1892 (NY) Gloucester: Rau s.n., 1882 (NY) dson: Austin s.n., 1861 (NY) Riccardia palmata (Hedw.) Carruth. on wet, rotting wood, especially in streams, shaded gorges Bergen: Howe s.n., 22 Sep 1898 (NY) Hudson: Austin [15 Hep. Bor.-Amer.] (BKL, NY) ussex: Barringer 21130 (BKL) Riccia (Ricciaceae) Riccia austinii Stephani (see R. lamellosa) Riccia beyrichiana Hampe on soil in wet fields and ditches, usually in partial shade (Hicks 1992, Lincoln 2008) SS Austin ne a Bor.-Amer.] (NY) bifurca pint fall ie 2008) on wet soil at edges of plowed fields or along streams (Hicks 1992, Lincoln 2008) 18 BARTONIA Bergen: Austin [141 Hep. Bor.-Amer.] (MO, NY) Monmouth: Haynes1649 (NY Riccia cavernosa Hoffm. (in NJ?) spores: Sep-Oct on wet or seasonally wet soil in disturbed sites and pond edges (Schuster 1992b) no county designated: Austin s.n., 1863 (MO) Riccia fluitans L. on still water of ditches and ponds or along their margins Bergen: C. C. Curtis s.n., 22 Sep 1898 (NY) Burlington: R. Sim s.n., 1934 (PH) Camden: Underwood s.n., Oct 1855 (NY) Cape May: Heritage s.n., 7 May 1857 (PH) Gloucester: Long 2905 (PH Hudson: Howe s.n., 22 Sep 1898 (NY) Ocean: Cowan & Wurdack 136 (NY) Salem: Long 1246 (PH) Sussex: Barringer 19930 (BKL) Riccia huebeneriana Lindenb. on soil in grassy ditches or at the edges of ponds Atlantic: A. Tuckerman s.n., 12 Oct 1970 (CHRB) Bergen: Cowan & Wurdack 27 (NY) Mercer: James s.n., 1867 (NY Monmouth: Haynes [116 Amer. Hep.] (BKL, NY) Riccia hirta (Austin) Underw. on wet soil in stubble fields or along the banks of streams (Schuster 1992b) Bergen: Austin 669 (NY) Riccia lamellosa Raddi on thin soil of exposed outcrops and overgrazed fields (Schuster 1992b) Bergen: Austin s.n., 1864 Riccia membranacea Gottsche & Lindenb. on soil in periodically flooded sites (Hicks 1992, Schuster 1992b) Bergen: Austin [150 Hep. Bor.-Amer.] (BKL, NY) Riccia sorocarpa Bisch. on soil in ditches and periodically flooded fields (Schuster 1992b) Bergen: Austin [139 Hep. Bor.-Amer.] (BKL, NY) Monmouth: Haynes 1493 Riccia sullivantii Austin (see R. huebeneriana) Riccia tenuis Austin (see R. membranacea) Ricciocarpos (Ricciaceae) Ricciocarpos natans (L.) Corda floating on still water or on soil along ponds and swamps Bergen: Wiegmann 1119 (BKL, NY) Burlington: Stowell 3 Mercer: Underwood s.n., Dec 1888 (CHRB) Morris: Moul 73-2 (CHRB) Sussex: Nearing s.n., 27 Sep 1947 (NY ) Scapania (Scapaniaceae) Scapania glaucocephala (Taylor) Austin on rotting wood in shade, usually evergreen logs (Schuster 1974) Bergen: Austin s.n., s.d. (NY) Scapania nemorea (L.) Grolle A PRELIMINARY CATALOG OF NEW JERSEY HORNWORTS AND LIVERWORTS on soil in seeps and moist roadbanks Bergen: Barringer 22300 (BKL) Cape May: Long 870 (PH) Cumberland: Long 1111 (PH) niet gi Long 2060a (PH) Hudson: Williams s.n., 7 Nov 1899 (NY) eae 1 Barringer 20746 (BKL) Monmouth: Haynes 987 (BKL, NY) Passaic: Barringer 20648 (BKL) Sussex: Barringer 20550 (BKL) Warren: Barringer 20925 (BKL) Scapania nemorosa (L.) Dumort. (see S. nemorea) Scapania undulata (L.) Dumort. on rocks along streams, always near fast running water (Schuster 1974) Bergen: Wurdack & Cowan 19 (NY) Burlington: Moul s.n., 23 Jul 1950 (PH) Sussex: Barringer 21037 (BKL) Union: Stephenson s.n., 27 Mar 1962 (CHRB) rren: Barringer 21248 (BKL) Telaranea (Lepidoziaceae) Telaranea nematodes (Austin) M. Howe in swamps Burlington: Buck 28290 (NY) Gloucester: Barringer 21206 (BKL) Monmouth: Haynes s.n., 30 Sep 1905 (BKL, NY) Trichocolea (Trichocoleaceae) Trichocolea tomentella (Ehrh.) terre along cool, shaded streams, swamp Bergen: Howe s.n., 6 Oct 1898 (NY) Hudson: Austin s.n, s.d. (NY) Middlesex: Marshall [56 Amer. Hep.] (BKL, NY) Monmouth: Haynes [9 Amer. Hep.] (NY) Passaic: Barringer 22543 (BKL) Sussex: Moul 7186 (CHRB, NY) Tritomaria (Jungermanniaceae) Tritomaria exsecta (Schmidel) Schiffner on rocks or tree bark in uplands (Hicks 1992) Bergen: Austin [46 Hep. Bor.-Amer.] (BKL, NY) 19 Bartonia No. 65: 20-28, 2011 Contributions to the Lichen Flora of Pennsylvania - Rare and Important Lichen Habitats and Lichen Communities: Part 1, the Northeastern Counties JAMES C. LENDEMER Cryptogamic Herbarium, Institute of Systematic Botany The New York Botanical Garden Bronx, NY, 10458-5126, USA jlendemer@nybg.org ABSTRACT. A summary of three biologically significant lichen habitats found in the northeastern counties of Pennsylvania is presented. INTRODUCTION The last several years have seen a significant increase in our knowledge of the lichen biota (lichenized, lichenicolous, and allied fungi) of the Commonwealth of Pennsylvania. Much of this information has been published in the form of floristic reports and checklists (Harris and Lendemer 2005, 2006; Lendemer 2005, 2008a, 2008b, 2009, 2010a, 2010b; Lendemer et al. 2009; Lendemer and Macklin 2006a, 2006b) and a number of new taxa have been described based on material collected in the state (e.g., Knudsen et al. 2007; Lendemer 2010a; Lendemer and Elix 2010; Lendemer and Harris 2007, 2009; Lendemer and Hodkinson 2010). Until 2007, the majority of the counties in Pennsylvania were represented by <100 known voucher specimens (historical or modern; see Plate 1). In an effort to remedy this situation the author (chairman of the Bryophyte and Lichen Technical Commission [BLTC] of the Pennsylvania Biological Survey [PABS]) in collaboration with the Western Pennsylvania Conservancy (WPC) began to systematically survey the lichen biota of imp | hroug! | The goal of this survey is to gather data relating to rare and significant lichen habitats in the Commonwealth while at the same time documenting occurrences to determine the conservation status and potential needs of the taxa that comprise the lichen biota in Pennsylvania. The first phase of this study was conducted during the 2008-2009 field season and focused on the northeastern counties (Plate 1). The present communication is intended to be the first in a series of reports documenting rare and significant lichen habitats and communities in Pennsylvania. Further phases of this study were completed during the 2009-2010 and 2010-2011 field seasons. These phases focused on the southeastern counties and north-central counties respectively. Comparable summaries of the rare and significant lichen habitats in those regions are currently in prepatation. MATERIALS AND METHODS a ee : 44. aes E 1 on a survey of natural areas conduced by the author in the northeastern counties of Pennsylvania. Field work was carried out between June 2008 20 CONTRIBUTIONS TO THE LICHEN FLORA OF PENNSYLVANIA 21 2010-2011 H 2008-2009 BM 2009-2010 Wi < 2008 Plate 1. Degree of completion of lichen county inventories to date. Counties represented by >100 modern vouchers are shaded (above), with the hue corresponding to field seasons. and June 2009 and represents two months (60 days) of person time. Survey localities were chosen by consulting the files and staff of the Western Pennsylvania Conservancy (WPC as well as the Natural History Inventories for each country that are available through the Department of Conservation and Natural Resources (DCNR) of Pennsylvania. Each locality was visited and an attempt was made to document total diversity by searching for all taxa of lichens and allied fungi. While it is impossible to document the entire lichen biota of a given site, it is assumed the repeated sampling of the same habitats and communities in a small geographic area gives a reliable picture of the total diversity of lichens and allied fungi. Voucher specimens were identified upon return to The New York Botanical Garden and deposited in the herbarium of that institution (NY). All vouchers were also databased with the data (including georeferenced localities) available online via the C.V. Starr Virtual Herbarium (http://sciweb.nybg.org/science2/VirtualHerbarium.asp). While filing the vouchers from this field study, all additional lichen specimens from Pennsylvania at NY were databased, and collectively these records have been used to create a dynamic checklist of the lichens and lichenicolous fungi of the state which can be viewed online at : | Aiea UES fo 9/1 1/1 / // the following address (http:/, asp) rs o RARE AND SIGNIFICANT LICHEN HABITATS AND COMMUNITIES I) Exposed Rocky Summits The summits of many mountains and ridge-tops in northeastern Pennsylvania, especially on the Pocono Plateau, are exposed and rocky and in these habitats lichens can represent a 22 BARTONIA SULLIVAN LUZERNE MONROE Plate 2. Detail of study area. significant amount of the diversity and biomass of the community. This habitat type roughly corresponds to the “Ridgetop Acidic Barrens Complex” of Fike (1999) and in Pennsylvania is frequently characterized by a scrub oak (Quercus illicifolia) - pitch pine (Pinus rigida) forest with an understory of ericaceous shrubs (Vaccinium, Gaylussacia, Kalmia). This habitat type is found throughout central and northeastern Pennsylvania and extends both northward and southward along the Appalachian Mountains where it is also characterized by a distinctive and significant lichen biota that often includes significant disjunct populations. In Pennsylvania the lichen biota of this habitat type is frequently not as diverse as others discussed in this paper, but rather often hosts disjunct populations of taxa at the northern or southern ends of their ranges (Lendemer in rev.). Presumably the persistence of large open rocky areas and pockets of humus allows species to become established and persist. As most of these habitats were glaciated during the last ice age the present day lichen biota represents relatively recent introductions since the end of that geologic period. Species with notable disjunct occurrences in this habitat type in Pennsylvania include Cladonia crispata, C. stellaris, Lecidea auriculata, Parmeliopsis hyperopta, and Porpidia contraponenda. Interestingly, this habitat type also supports populations of Cladonia atlantica and C. floridana, species of the coastal plain which also have disjunct populations in the Shawangunk Mountains of New York State (Dirig 1994), In the study area, Bartlett Mountain in State Game Lands No. 57 without a doubt represents the most significant exemplification of this habitat type and should be considered of high conservation priority. Another important example of this habitat on public land is the summit of the north end of the Moosic Mountains in State Game Lands No. 300. An excellent example of this habitat type also occurs on Bald Mountain in Lackawanna County. Of immediate concern to the site in SGL 300, and other ridge tops and rocky summits, is the installation of wind turbines for the generation of electricity. Further study of the impact of wind turbines on the lichen biota of this habitat type is clearly needed. In CONTRIBUTIONS TO THE LICHEN FLORA OF PENNSYLVANIA Plate 3. Rare and critical lichen habitats in northeastern Pennsylvania: exposed rocky summits. Fig- ures 1, 2, 4, 5, Moosic Mountain TNC Preserve, Lackawanna County. Figure 3, Moosic Mountain, summit of N end, Lackawanna County. addition to the disturbance caused by the installation and maintenance of these structures there is the possibility that the change in wind currents created by the turbines could result in the establishment of new fungi altering the integrity of the community. 24 BARTONIA Plate 4. Rare and critical lichen habitats in northeastern Pennsylvania: Boulderfields and talus slopes. Figures 1-2, Bald Mountain, W-slope, Lackawanna County. Figures 3 and 4, Boulderfield Natural Area, Hickory Run State Park, Carbon County. (Figure 4 illustrates a cool, humid, protected micro- habitat in between and below two large boulders.) II) Boulder fields and talus slopes The potential biological significance of boulder fields and talus slopes to the lichen biota of Pennsylvania has been outlined by Lendemer et al. (2009). These habitats are not typically included in present community classifications in the state, likely because they are only marginally vegetated. It is also likely that they are included in other community types as they frequently occur as part of larger community complexes such at that outlined above. Regardless of their classification in other schemes, they warrant particular attention here because of the diversity of lichens and lichenicolous fungi they host. CONTRIBUTIONS TO THE LICHEN FLORA OF PENNSYLVANIA 25 In northeastern Pennsylvania boulder fields and talus slopes are usually associated with flowing water or cold air flows that alter the temperature and humidity regime significantly from the surrounding area. It is almost certainly this factor that fosters the persistence of disjunct populations of species, particularly those with arctic-boreal affinities. The lichen biota of boulder fields and talus slopes stand in contrast to those of exposed rocky summits in that they are more diverse and frequently harbor the bulk of this diversity as isolated populations of variable (usually small) size. Notable species found in this habitat in northeastern P centrifuga (common N of PA, arctic-boreal), Chrysothrix chlorina (rare, boreal), Cladonia stygia (common N of PA, arctic-boreal), Fuscidea recensa (common, northern-boreal), Lepraria cryophila (common throughout Appalachians), L. morozit ined. (rare, Appalachian endemic), Melanelia culbersonii (rare, eastern North American endemic), M. panniformis (common N of PA, arctic-boreal), M. sorediata (common N of PA, arctic-boreal), Parmelia neodiscordans (rare outside of PA, eastern North American endemic), PR omphalodes (common N of PA, arctic-boreal), Phlyctis petraea ined. (common, undescribed eastern North American endemic), Porpidia cinereoatra (common N of PA, arctic-boreal), P contrapondenda (same as previous), P. soredizodes (i in PA, distribution elsewhere unknown), P. tuberculosa (common N of PA, arctic boreal), Protothenella corrosa (rare, scattered occurrences in boreal North America), and Stereocaulon glaucescens (rare, eastern North American boreal endemic). As a result of the fact that the rare or disjunct species in this habitat type occur as small scattered populations many occur only at one or a handful of localities, contributing to a high heterogeneity among sites. The best two examples of this habitat type reviewed during the present phase of the study were in Rothrock State Forest (Huntingdon Co., see Lendemer et al. In press) and on the W-facing slope of Bald Mountain (Lackawanna Co.). Boulder Field Natural Area in Hickory Run State Park (Carbon Co.) hosts populations of significant lichen taxa however is heavily visited by hikers and thus large portions of the site have been degraded by foot traffic and the movement of rocks which damages slow growing lichen communities. The first two sites mentioned above should be given high conservation priority and the climate regimes protected from future alteration. III) Rock Overhangs ] ia include Arctop li, Rock overhangs represent a common habitat in North America that is frequently overlooked by lichenologists. Such habitats, especially when occurring on steep slopes near flowing water, are characterized by high humidity and relatively stable temperatures. In these respects they are similar to the crevices between and under rocks in boulder fields or talus slopes and as such there is some overlap in the species that are typically found (e.g. Lepraria normandinoides, Phlyctis petraea, and Psilolechia lucida). In the study area however, acidic rock overhangs are often composed of sandstone rocks with weakly calcareous cement that facilitates the growth of a unique community of lichens. Most of these species are widespread, and occur where the habitat occurs, however populations of rare lichens are also often found intermixed with the more common species. Species that are typical of acidic rock overhangs in northeastern Pennsylvania include the following: Amandinea punctata s.|. (saxicolous populations), Botryolepraria lesdainu, Caloplaca flavocitrina, C. flavovirescens, C. subsoluta, Phlyctis petraea ined., Psilolechia lucida, Ramalina intermedia, and Physcaa subtilis. Rare species that occur in this habi ber of species with proposed relictual distributions reflecting a wider distribution in the past. These species BARTONIA er. SPAS GMM a REP ” Plate 5. Rare and critical lichen habitats in northeastern Pennsylvania: rock overhangs. Figure 1, Wal- lenpaupack Ledges, Lacawac Sanctuary, Wayne County. Figures 2 and 4, middle slopes of Moosic Mountains, State Game Lands No. 300, Lackawanna County. Figures 3 & 5, Bowmans Creek Ledges, Wyoming County. include: Diplotomma alboatrum, Lecanora cinereofusca var. appalachensis, Lepraria adhaerens, Ramalina petrina, and Caloplaca reptans ined. It should be noted that this habitat type supports several occurrences of the only known extant member of the Pannariaceae in the commonwealth, Vablhella leucophaea. Though common elsewhere this species and all other members of the CONTRIBUTIONS TO THE LICHEN FLORA OF PENNSYLVANIA 27 family which were historically at least widespread in Pennsylvania seem to have been extirpated. The reason for the disapearence of these taxa is not clear, however it likely was a result of the extensive deforestation that occurred in the state. A species that should be given protection if extant populations are found is Santessoniella crosssophylla (see Lendemer and Anderson 2008), it is globally rare, and extremely rare in North America. A historical record from Glen Onoko, Carbon County is known, however the species has been extirpated from that site. Acid rock overhang communities are extremely fragile because they rely on a narrow environmental resigeme, changes to which can alter the community and cause rare species to be extirpated. Several species (Botryolepraria lesdainit, Lepraria adhaerens, Caloplaca reptans) often occur on loose pebbles and soil in these overhangs and any disturbance to populations of these species should be avoided. Excellent examples of this habitat type occur in State Game Lands 300 in the north end of the Moosic Mountains, State Game Lands 57 near the Wyoming/Luzerne County line, along Bowmans Creek in Wyoming County, on the middle-low slopes of Bald Mountain in Lackawanna County, and on the property of Lacawac Sanctuary in Wayne County. CONCLUSION Lichens and allied fungi represent a biologically diverse group of taxa that has already suffered, and continues to suffer, significant negative impacts from anthropogenic changes to the natural landscape of Pennsylvania. Widespread deforestation throughout the commonwealth caused significant declines in the populations of many species (Lendemer unpublished data). It is also likely that this factor, combined with air pollution and acid rain following the industrial revolution that caused further declines and extirpations of other species. Unfortunately study of this phenomenon is hampered by a lack of adequate historical data except for a select group of large (and thus frequently collected) ecologically sensitive macrolichens such as those in the genera Ramalina and Usnea. The contrast between the historical abundance of these taxa in our forests, and their near absence in modern times is almost certainly only a hint at the degree of change that has occurred to the lichen biota of Pennsylvania, and eastern North America as a whole, during the last three centuries. Outlined above are three habitat types that are critical to the management and preservation of the present-day lichen biota of the northeastern counties of Pennsylvania. Evaluation and protection of occurrences of these habitat types is necessary to maintain the diversity of lichens, lichenicolous fungi, and allied fungi of the region. A habitat based approach te the conservation of the lichen biota of th rst, the small size and isolated scattered nature of the populations of some of the rarest species Vig a species based conservation approach untenable. Second because of their small size : many species cannot be reliably identified, in the field or the lab, by anon y ahandful of experts in North America. Third, pecies often occur together at a given locality because the factors allowing one taxon to survive facilitate the survival of others. By conserving exemplary examples of significant habitat types, it is possible to conserve an entire — lichen community that is often dependent ona specific long-term climatic regim After three years of intensive field work it is nothing short of acitedile that collecting even in the best known areas continues to result in the discovery of previously unreported taxa, extant populations of taxa known only from historical reports, and even taxa that are new to science (see citations above). The shear number of new reports is a clear indi a) how poorly studied and under collected Pennsylvania remains at the end of the first decade 28 BARTONIA of the 21* century. This paucity of data is particularly troubling in light of the rapid pace of development of natural resources in the Commonwealth as well as the large-scale changes to forest structure that are projected to occur due to the spread invasive species. It is clear that this significant gap in our knowledge of the biota of Pennsylvania will only be filled through continued large scale field inventories. LITERATURE CITED DiriG, R. 1994. Lichens of pine barrens, pine pin and “ice cave” habitats in the Shawangunk Mountains, New York. Sra ai 523-558 FIKE, J. 1999. Terrestrial and E ities of Pennsylvania. Harrisburg, Pennsylvania. 86pp. Harris, R. C. AND J. C. LENDEMER. 2005. Contributions to the lichen flora of Pennsylvania: A checklist of the lichens collected during the first Howard Crum pir ae Workshop, Delaware Water Gap National Recreation Area. Opuscula Philolichenum 2: 1-1 Harris, R. C. AND J. C. LENDEMER. 2006. Contributions to the me flora of Pennsylvania: Additions to the checklist of lichens of the Delaware Water Gap National Recreation Area. Opuscula Philolichenum 3: 69-78. ARRIS, R. C. AND J. C. LENDEMER. 2009. The Fellhanera silicis group in eastern North America. Opuscula Philolichenum 6: 157-174. KNUDSEN, K., J. A. ELIX, AND J. C. LENDEMER. 2007. Lepraria adhaerens, a new species from North American. Opuscula Philolichenum 4: 5-10. KNUDSEN, K. AND K. KOCOURKOVA. 2010. Pyrenidium aggregatum, a new species from North America. Opuscula Philolichenum 8: 71-7 LENDEMER, J. C. 2005. Contributions to the lichen flora of Pennsylvania: The lichen flora of the diabase region of northern Bucks and Montgomery Counties. Opuscula Philolichenum 2: 21-26. a Shae J. C. 2008a. A note on the occurrence of Psora icterica in eastern North America. Evansia 25(1): 1 alan C. 2008b. New and i eer records of lichens and lichenicolous fungi from New Jersey and Pennsylvania. Evansia 25(4): 102-109. LENDEMER, J. C. 2009. Contributions to the bikes flora of ee — Further new and interesting reports of lichens and lichenisolous fungi. Evansia 26(3): 131 LENDEMER J. C. 2010a. N Lepraria s.1. (1 cae A ) in North America: new species, new reports, and bat aes keys. Brittonia 62(3): 267- 292. LENDEMER, J. C. 2010b. Contributions to the lichen flora of Pennsylvania. — Further new and interesting reports of lichens and lichenicolous fungi. Evansia 27(2): 47-64. LENDEMER, J. C., H. M. EDENBORN AND R. C. HARRIS. 2009. Contributions to the lichen flora of Pennsylvania: Notes on the lichens of a remarkable talus slope in Huntingdon County. Opuscula Philolichenum 6: 125-136. LENDEMER, J. C. AND J. A. ELIx. 2010. Two new species of Chrysothrix from eastern North America. Opuscula Philolichenum 8: 51-58. LENDEMER, J. C. AND R. C, HARRIS. 2007. Lepraria normandinoides, a new widespread species from eastern North America. Opuscula Philolichenum 4: 45-50. LENDEMER, J. C. AND B. P. HODKINSON. 2010. A new perspective on Punctelia subrudecta (Parmeliaceae) i in North America: Previously anjected ne characters re t 42(4): 405-421. LENDEMER, J. en "AND J. A. MACKLIN. 2006a. Contributions to the lichen flora af esePartin A preliminary checklist of the lichens of Nescopeck State Park. Opuscula Philolichenum 3: 41-48. LENDEMER, J. C. AND J. A. MACKLIN. 2006b. Contributions to the lichen flora of Pennsylvania: A preliminary checklist of the wPeast of Worlds End State Park. Opuscula Philolichenum 3: 53-64. Bartonia No. 65: 29-96, 2011 An Overview and Checklist of the Vascular Flora of the Delaware Water Gap National Recreation Area in New Jersey and Pennsylvania RICHARD STALTER', YANA MICHAELS', TOMAS Mujo', M. H. CHOO', PATRICK LYNCH? AND STACY JOERGLE-GILDEA? ‘Department of Biological Sciences, St. John’s University, Jamaica, New York 11439 *National Park Service, Delaware Water Gap National Recreation Area, 294 Old M ilford Road, Milford, Pennsylvania 18337 ABSTRACT. A survey of vascular plants of the Delaware Water Gap national Recreation Area in New Jersey and Pennsylvania was conducted. A total of 1,271 species (in 154 families and 546 genera) were identified, with 1,055 species being native to the region. Overviews are also provided on vegetational history, current vegetational communities, and management of invasive species. INTRODUCTION Delaware Water Gap National Recreation Area (DEWA) was established by an act of Congress on September 1, 1965, for public outdoor recreation use and for the preservation of scenic, scientific and historic resources (Public Law 89-158). The federal boundaries extend to over 27,200 ha. in northeastern Pennsylvania and northwestern New Jersey (see www.nps.gov/dewa/planyourvisit/maps.htm) . The actual “water gap,” 40.979 N 75.143 W, is a distinctive geologic feature cut across the Kittatinny Ridge of the Appalachian Mountain chain by the erosive action of the Delaware River. Natural areas within this national park area include uplands associated with the Kittatinny Ridge, wetlands, and riparian areas surrounding the Middle Delaware Scenic and Recreational River and its tributary streams. The areas also include hemlock ravines and rhododendron glades endemic to the New Jersey Highlands and the Pocono Plateau, several waterfalls and upland native grasslands, and river valley bottomlands. These areas have been historically farmed and are now maintained as open space and farmland to maintain the pastoral scene under DEWA’s open space mandate. Vegetational History Pinus strobus and Tsuga canadensis were the former climax species in the region (Kuchler 1964). Sites that were disturbed by windthrows or fire were populated by hardwoods (Trembly et al., 1963). In the 1840s P. strobus was cut for lumber. Debris from pine logging provided fuel for fires that ravaged portions of the cut forest. Succession at burned sites produced a different community from the original pine forest. Hardwoods, especially Castanea dentata, replaced the old pine community. C. dentata was decimated by chestnut blight (1910-1920). With the demise of American chestnut, many landowners converted 30 BARTONIA their hardwood woodlots to pine forests since P. strobus grew faster and was a more valuable timber crop than the slower growing hardwoods. Floodplain forest The floodplain forest borders the Delaware River and its larger tributaries. Common species here are Acer saccharinum, Betula nigra, Populus spp., Salix discolor, S. eriocephala, S. nigra, and Platanus occidentalis. Ulmus americana was more common in the early 1960s but has since been largely reduced due to Dutch elm disease. Successional Fields Fields in various stages of succession are populated by grasses notably Schizachyrium scoparium. The park has over 1,093 ha. of cropland and over 1,619 ha. of transitional fields. Toxicodendron radicans, Rubus allegheniensis, R. flagellaris, R. hispidus, R. occidentalis, and Rhus glak hrubs while Juniperus virginiana and Fraxinus americana are pioneer tree species. On higher-drier sites grasses predominate, including Schizachyrium scoparium. Aster (e.g., Eurybia, Symphyotrichum) and goldenrods (Euthamia, Solidago) pi i late summer and early fall. Common woody invaders are Populus grandidentata, P. tremuluides, Betula populifolia, Juniperus virginiana, Sassafras albidum, Prunus serotina, and Pinus rigida. P. rigida is common and occurs sporadically on the driest sites. Upland Forests Quercus alba, Q. coccinea, Q. montana, Q. rubra, Q. velutina, and Acer saccharinum are the dominant trees at upland forested sites occasionally mixing with Pinus strobus. In moist ravines, Tsuga canadensis dominates often forming pure stands. The park has over 140 distinct hemlock stands totaling over 1,241 ha. The hemlock woolly adelgid (Adelges tsugae) is a recent alien pest on T. canadensis that threatens this species. Scrub Oak Barrens Quercus ilicifolia, scrub oak, is the dominant species in the barrens. Pinus rigida is also present at the driest sites while Populus spp. occupy the wettest sites in the barrens. Ericaceous species, especially Vaccinium pallidum and V. angustifolium, may be locally abundant. Cliffs A prominent feature of DEWA are the nearly sheer cliffs with talus slopes at their base (Trembley et al. 1963). Juniperus virginiana forms nearly pure stands at times at the top border of the cliffs and also grows in crevices in the cliff walls. Some individual trees may be much older than the J. virginiana in the successional fields since the cliff trees are protected from fire and have little competition from hardwoods. Opuntia humifusa often thrives at the xeric cliff edge. Talus slopes The vegetation of the talus slopes is sparse because of the unstable nature of the habitat. Juniperus virginiana and Populus spp. may invade the area but their footholds here are tenuous. OVERVIEW AND CHECKLIST OF THE VASCULAR FLORA OF THE DELAWARE WATER GAP 31 Ravine Banks Ravine banks are characterized by their steepness and lower amounts of sunlight. Pinus strobus may be abundant on the sunny drier north facing slopes, while oaks (Quercus montana and Q. rubra) and hemlocks (Tsuga canadensis) are more common on the moister south facing slopes. Oaks may become more abundant on the south facing slopes in the future because of the demise of T: canadensis due to the woolly adelgid. River Banks and Islands Flooding and ice scouring have a powerful effect on the vegetation of the Delaware River floodplain forest. Successional species such as Populus deltoides, Salix eriocephala, and S. nigra often form dense thickets on the Delaware’s banks. The islands in the Delaware River are forested by Acer saccharinum, Betula nigra, Populus deltoides, and Salix spp. A stand of Liriodendron tulipifera occupies Depue Island, the only site within Delaware Water Gap where Liriodendron occurs in pure stands. The exotic Japanese knotweed (Polygonum cuspidatum) has invaded these areas. Ruderal Sites While not included in Trembley’s list of habitats, ruderal sites support a rich and varied group of vascular plants including many non-native species. Ruderal sites include roadsides, old habitations, disturbed power line right-of-ways and other disturbed habitats. Non- native species of grasses and composites are generally found here as well as members of the Amaranthaceae, Brassicaceae, Chenopodiaceae, and Polygonaceae families. A common grass genus is Setaria. Microstegium vimineum is rapidly invading shaded roadsides and may become more abundant at these sites in the future. Ponds and Lakes There are 256 ponds and lakes within DEWA. These water bodies range in size from 0.01 hectares at Mashipicong to 5.9 hectares at Hidden Lake. Within lakes and ponds Nymphaea odorata, Nuphar microcephala, N. variegata, Potamogeton crispus, P. epihydrus, and P. nodosus flourish while many species of Cyperaceae, Juncaceae, Malvaceae, and Poaceae populate their moist shores. EXOTIC SPECIES MANAGEMENT Park staff |} 1 the foll native va ular pla nts for control in 2005 (Lynch et al. 2005): Elaeagnus umbellata, Ailanthus alesis; Rosa multiflora, Polygonum cuspidatum, Lythrum salicaria, Lonicera spp., and Humulus japonicus. Detailed information in Lynch et al. (2005) on the biology and management of Lythrum salicaria, whorled loosestrife, was prepared by Shreiner (2005). The park hopes to reduce L. salicaria’s spread and restore sites severely degraded by loosestrife. Research on the effect of defoliation of Tsuga canadensis by the woolly adelgid was initiated in 2003 to determine the effects of defoliation on the growth of alien Japanese stiltgrass (Microstegium vimenium). Preliminary results suggest that M. vimenium is more abundant under adelgid denuded T. canadensis, and may become more widespread in the future (Evans, 2005). Park staff predict that all 7; canadensis will succumb to woolly adelgid infestation in 25 to 35 years. 32 BARTONIA CLIMATE Northeastern Pennsylvania and northwestern New Jersey are classified as having a humid continental climate (Garwood 1996). Three weather stations bordering the Park: Stroudsburg, PA, Monroe County (40.59N, 75.24W); Port Jervis, Orange County, NY, (41.23N, 74.60W); and Sussex County, NJ (41.13N, 74.75W) provided long term weather data for the region. Rainfall is evenly distributed throughout the year. The wettest months are May (112 mm. Port Jervis), June (118 mm Sussex), and August and September (115 mm Stroudsburg). February is the month with the lowest average rainfall at all 3 stations ranging from 72 mm. (Port Jervis) and 75 mm. (Sussex and Stroudsburg). Yearly rainfall ranges from 1124 mm. (Port Jervis) to 1227 mm. (Stroudsburg). Winters are cold with January means of -8.5° C at Stroudsburg to —4.8° C at Sussex. Snowfall was highest at the northern station, Port Jervis (1155.7 mm.) and least at the southern Stroudsburg station (962.7 mm.); no snowfall records were available at Sussex. July was the warmest month at all stations with a mean of 23° C at Stroudsburg, 22.4° C at Port Jervis, and 21.6° C at Sussex. METHODS The objective of the study was to prepare a list of the families, genera, and species of vascular plants found within DEWA. Collecting trips were made to DEWA in September 2003, October 2003, May 2004, July 2004, August 2005, October 2005, and August 2006. Objectives for each were the collection of voucher specimens and information on habitat preference for each species. Approximately 400 species were collected by the senior author. These specimens and species lists generated by investigators (National Parks Service 2006), who provided most of the data, form the basis for this flora. Taxonomically problematic specimens were sent to various experts for identification. Experts consulted were [Ihsan Al-Shebaz (Brassicaceae), Jim Montgomery (ferns and fern allies), Eric Lamont (Asteraceae), Charles Sheviak (Orchidaceae), and Gordon Tucker (Cyperaceae). Voucher specimens collected in the present study are deposited at the National Park herbarium in Bushkill, Pennsylvania. Catalogue numbers have been assigned by the National Park Service to the primary sets of specimens at the herbarium in Bushkill, Pennsylvania and are available on request from the National Park Service. Nomenclature follows Rhoads and Block (2007). Non-native plant status was determined by Gleason and Cronquist (1991); these taxa are preceded by an asterisk. RESULTS AND DISCUSSION A total of 154 families, 546 genera, and 1,271 species have been identified. One thousand fifty-five species were native to the region. Largest families in the flora were the Cyperaceae (121 species), Asteraceae (114 species), and Poaceae (96 species). Altogether these families composed 26% of the flora. The largest genera were Carex (90 species), Polygonum (17 species), and Salix (15 species). Non-native taxa composed 17% of the park’s flora. While non-native taxa composed a small percentage of the flora, their potential impact on native species may be considerable. Some additional gh seagate non-native vascular plant species, not discussed earlier were Acer plat. , Celastrus orbiculatus, OVERVIEW AND CHECKLIST OF THE VASCULAR FLORA OF THE DELAWARE WATER GAP 33 Lonicera japonica, and Rosa multiflora. A. petiolata is one of the most common species in the most northeastern woodlands. A. petiolata continues to colonize the floor of the forest and may become the most common herbaceous understory species. Two woody lianas, Lonicera japonica and more recently Celastrus orbiculatus, have invaded woods and fields smothering, shading, and out competing native species. Acer platanoides leafs out earlier than native trees and holds its leaves in the fall longer than most native species. Few native plants can survive in the dense shade of Acer platanoides. Rosa multiflora is common in fields and open woodlands. Microstegium vimineum, a native of tropical Asia, is rapidly invading moist open woodlands, especially hemlock woodlands near the Delaware River. Microstegium out competes native grasses and forbs, and may have a greater deleterious impact on native vascular plant species at DEWA in the future when Tsuga canadensis death reduces shading and enables M. vimineum to become more abundant on newly sunny opened portions of the forest floor. CLASSIFICATION OF VEGETATION TYPES Kuchler (1964) classified this region as “Appalachian oak forest” within the Eastern deciduous forest zone. Drier ridge tops and slopes are populated by a preclimax forest of Quercus montana and Q. alba, while Q. rubra is more common on more mesic soils. Trembly et al. (1963) defined eight major plant communities at the Delaware Water Gap National Recreation Area (DEWA). We expanded these to include ten general habitats: old upland fields/successional fields/meadows, ruderal sites/roadsides/disturbed sites/old home sites/power line rights-of-way, slopes/upland forests/woodlands, lowlands/swamps, lakes/ponds/marshes/bogs, river banks/islands/floodplain forests, ravine banks/ravines, scrub oak barrens, cliffs/ridges, talus slopes. While our general habitat descriptions follow Trembly et al. (1963), our habitat classification also recognizes: ruderal sites and lakes, marshes, and bogs. The abundance classes for vascular plant species at each habitat are based on personal observations. The greatest number of vascular plant species were observed in slopes, upland forests and woodlands, old upland fields, successional fields, meadows, and ruderal (disturbed) roadsides and old home sites. The unstable talus slopes support the fewest vascular plant species (Table 2). The vascular flora at the ten sites may vary in number and kind, as conditions (drought, insect damage) vary at each site from year to year. There are 167 listed species of management concern throughout the national park. Table 1. A summary of the vascular flora of Delaware Water Gap National Recreation Area. Ferns Fern allies Coniters Dicots Monocots Totals Families 17 4 4 107 22 154 Genera 23 6 9 385 123 546 Species a? 20 16 842 354 1271 Native species 39 20 16 670 310 1055 Non-native species 0 0) 0 173 45 218 34 BARTONIA Table 2. Number of vascular plant species at the ten recognized habitats. Habitat codes: OSF, old upland fields, successional fields, meadows; R, ruderal sites, roadsides, disturbed sites, old home sites, power line rights-of-way; SUF, slopes, upland forests, woodlands; LS, lowlands, swamps; LPM, lakes, ponds, marshes, bogs; RIF, river banks, islands, floodplain forests; RB, ravine banks, ravines; SOB, scrub oak barrens; C, cliffs, ridges; TS, talus slopes. OSF R SUF LS LPM RIF RB SOB C TS Spore Plants 12 10 32 7 15 3 5 1 15 2 Conifers 3 4 7 2 2 0 2 2 3 0 Dicots 327 336 421 111 118 94 45 28 22 6 Monocots 85 72 136 50 144 7 7 8 3 4 Totals 427 424 S95 170 278 104 61 39 42 11 ACKNOWLEDGMENTS Appreciation is expressed to Superintendent John Donahue, Delaware Water Gap National Recreation Area, for unrestricted access to the park. We thank Jeff Shreiner and Patty Tipson, DEWA, who assisted us with the project. For assistance in the identification of several taxa we thank Ihsan Al-Shebaz, Eric Lamont, Jim Montgomery, Charles Sheviak, and Gordon Tucker. Finally PE | support of St. John’s University. LITERATURE CITED GARWOOD, A. N. (Ed.) 1996. Weather America. Toucan Valley Publications, Milpitas, California. 1412 p. GLEASON, H. A. AND A. CRONQUIST. 1991. Manual of Vascular Plants of Northeastern United States and Adjacent Canada. The New York Botanical Garden, Bronx, New York. 910 pp. KARTESZ, J. T. 1994. A Synonymized Checklist of the Vascular Flora of the pas States, Canada, and Greenland. Volume 1-Checklist. Timber Press Inc. Portland, Oregon. 622 p. KUCHLER, A. W. 1964. Potential Natural Vegetation of the a. United States. American Geographic Society, New York. 116 LYNCH, P. J., A. AMBLER, J. SHREINER, L. HILAIRE, R. EVANS, D. COOKE, L. MORLOCK, P. TIPSON. 2005. National Park Service, Delaware Water Gap National Recreation Area, Division of Research and Resource Planning Ann ; NATIONAL PARKS SERVICE. NPSpecies Proper: hci on ~ The National Park Service Biodiversity Database. Desktop version 2.21. (accessed Au ). RHoaps, A. F. AND T. A. BLOCK. 2000. The Pets “of Pennsylvania. University of Pennsylvania Press, Philadelphia. 1061 p. RHOADS, A. F. AND T. A. BLOCK. 2007. Pennsylvania Flora Project Database. Morris Arboretum, University of Pennsylvania, Philadelphia. . (accessed February, 2007). TREMBLY, F. J., J. M. FRANCIS JR., B. H. KNApp, J. W. WHITEHOUSE, B. W. PARKER, H. A. WAHL. 1963. A Natural History Survey of the Proposed Tocks Island Reservoir National Recreation Area. National Park Service and the Water Resources Association of the Delaware River Basin, Columbia, New Jersey: UNITED soe DEPARTMENT OF AGRICULTURE, NATURAL RESOURCES CONSERVATION SERVICE, NATIONAL PLANT DaTA CENTER. 2009. PLANTS Database. . tatésened January,2009). 25 Saal Habitat codes: see Table 2. Frequency: V, very abundant; F, frequent; I, infrequent; R, rare. Other codes: *, populations not known native to DEWA; O, observed. State status in New Jersey: NJE, endangered. State status in Pennsylvania: PE, lacued: PT, threatened; PR, rare; PX, extirpated. State (S) and Global (G) ranks given follow NatureServe (www. natureserve. ee aay Citation of a plant rarity status for NJ or PA does not imply that the taxon was found from that state within the DEW. OSF R SUF LS LPMRIF RB SOB C TS -¥ state status Ferns and fern allies Adiantaceae Adiantum pedatum L. a. F Cheilanthes lanosa (Michx.) D.C. Eat R §2 (NJ) Phegopteris connectilis (Michx.) Watt O R §2 (NJ) Phegopteris hemperontere eas )Fee(T Rk F R hexagonoptera (Michx.) T. Pellaea atropurpurea (L.) ae R R Aspleniaceae Asplenium bradleyi D.C. Eat R NJE, PT, S1 (NJ, PA) Asplenium montanum Willd. R $2 (NJ) Asplenium pinnatifidum Nutt. R NJE S1 (NJ), $3 (PA) Asplenium platyneuron (L.) BSP. F F Asplenium rhizophyllum L. R R Asplenium ruta-muraria L. R Asplenium ee R R L. ssp. trichoman Asplenium X pee t R.R. Scott (pro sp.) R [A. ebenoides Blechnaceae Woodwardia areolata (L.) T. Moore I ‘slayorsq ul papraoid useq sary ($661) ZSe1IeY 10/pue (1661) asinbuoiy pur uosea;s wor sawepyy ‘ue synuaios aya sapnpour Anjua yoeq “Ayeonoqeydye posurse aie exe] Jamo] pur ‘sarpruey ‘A108a7e9 ‘A10893e9 yova uTYI/K *s10D0UOUI pur ‘sj0orp ‘surads -ouwAS ‘sare Ud} pur SUID} :sa1IOSaIVd ZULMOT[O} ay 01 poSuLIe UIEeq savy exe) JUETd IJepNOSeA DY], ‘SaIOddS AO LSITIOAHD GALVLONNY XIGNAdd VY $¢ dV YALVA AUVAV TAC AHL AO VYOTA UV TNOSVA AHL AO LSITIOAHD ANV AAIAYAAO OSF R° Sie LS LPM RIF- RB SOB” C’ FS: =stare statis Woodwardia virginica (L.) Sm. I Dennstaedtiaceae Dennstaedtia punctilobula (Michx.) F Vv T. Moore Pteridium aquilinum (L.) Kuhn F V F Dryopteridaceae — Cystopteris tenuis (Michx.) Desv. F Diplazium pycnocarpon (Spreng.) Braun a 4 Dryopteris clintoniana (D.C. Eat) Dowell R $3 (NJ), S2(PA) Dryopteris marginalis (L.) Gray O F F Dryopteris X slossoniae Wherry ex Lellinger R Dryopteris X triploidea Wherry R R R Gymnocarpinm dryopteris (Rupr.) Ching O R $182 (NJ) Matteuccia struthiopteris (L.) Todaro O R O Onoclea sensibilis L. i. On F Polystichum ichoides (Michx.) Schott F O Woodsia ilvensis (L.) R. Br. Vv Woodsia obtusa (Spreng.) Torr. O Equisetaceae Equisetum arvense L. F F Equisetum fluviatile L. R F Equisetum hyemale L. R R R Equisetum laevigatum A. Braun Equisetum * litorale Kihlewein ex Rupr. (pro sp.) I Equisetum sylvaticum L. RoR HO $3 (NJ) VINOLYVa OSF R Sup is LPM RIF RB SOB C TS state status Hymenophyllaceae Isoetaceae Lycopodiaceae Ophioglossaceae Osmundaceae Equisetum variegatum Schleich ex F. Webert & D.M.H Mohr Trichomanes intricatum Farrar. Isoetes engelmannii A. Braun Isoetes riparia Engelm. ex A. Braun Diphasiastrum digitatum (Dill. ex A. Braun) Holub Lycopodium complanatum L., Lycopodium d. Dill. ex A. Braun] Diphasiastrum tristachyum (Pursh) Holub [Lycopodium t. Pursh] Huperzia — (Michx.) Trevisan [Lycopodium lucidulum Michx.} Huperzia porophila (Loyd & Underwood) Trevisan [Lycopodium porophilum L. Loyd & Underwood] Lycopodiella inundata L. Lycopodium clavatum L. Lycopodium dendroideum Michx. Lycopodium obscurum L. Botrychium dissectum Spreng. Botrychium multifidum (Gmel.) Trev. Ophioglossum pusillum Rat. Osmunda cinnamomea L. Osmunda claytoniana L. Osmunda regalis L. oo 2 ew <_ a SS. we A VO VO $2 (NJ) $3 (NJ) PE, S1 (PA) $1 (NJ) NJE, St (NJ) $3 (NJ) Z¢ dVD WALVA FAVAV TAC FHL AO VUOTA UV INOSVA FHL AO LSITIOAHSD GNV AAIAWAAO OSF KR. ap 15 tee RF RW SOK CO TS gacdu |= % Polypodiaceae Polypodium virginianum L. oO oO Selaginellaceae —_ Selaginella apoda (L.) Spreng. oO oO Selaginella rupestris L. R R $2 (NJ) Thelypteridaceae — Thelypteris palustris var. pubescens (Lawson) Fern. Vv [Dryopteris thelypteris auct. non (L.) Sw.] Thelypteris simulata (Davenport) Nieuw. R Gymnosperms Cupressaceae Juniperus communis L, R R Juniperus virginiana L. F Vv *Thuja occidentalis L. O° R e Pinaceae Abies balsamea (L.) P. Mill. 3 Fs *Larix decidua P. Mill. O.=8 9 *Picea abies (L.) Karst R > Picea glauca (Moench) Voss R Picea rubens Sarg. R NJE, $1 (NJ) Pinus resinosa Soland vo NJE, S1.1 (NJ) Pinus rigida P. Mill. F Pinus strobus L. F *Pinus sylvestris L. Oo RO Pinus virginiana P. Mill. O We Tsuga canadensis (L.) Corr. Vv Taxaceae Taxus canadensis Marsh. Vo oO $2 (NJ) OSF R SUF LS LPM RIF RB SOB .C TS state stetus Angiosperms Dicots Acanthaceae Justicia americana (L.) M. Vahl. F Adoxaceae Sambucus canadensis L. O F/O F/O E i Sambucus racemosa L. var. pubens (Michx.) Koehne R R Viburnum acerifolium L. Vv Viburnum dentatum L. HO. 0 Viburnum lantanoides (Michx.) [V. alnifolium Marsh.] R R NJE, $1 (NJ) Viburnum lentago L. YO °Ue oO * Viburnum opulus L. O Viburnum plicatum Thunb. R Viburnum prunifolium L. F Rs) Viburnum rafinesquianum J. A. Schultes R R Viburnum recognitum Fern. [V. dentatum var. O oo... lucidum Ait.] Viburnum trilobum Marsh. [V. opulus L. var. R R R $3 (NJ),S1S2 americanum Ait.] (PA) Altingiaceae Liquidambar styraciflua L. R R Amaranthaceae = *Amaranthus albus L. F *Amaranthus hybridus L. F Anacardiaceae Rhus copallina L. var. copallina Vv R/O Rhus glabra L. Vv O Rhus typhina L. [R. hirta (L.) Sudworth.] Vv O 6¢€ dV9O YALVA AUVAV TAC AHL AO VUOTS UV TINOSVA AHL AO LSITIOAHO UNV AAIAYAAO CSF R Me 18 LOM RIF RE SOB C TS © state stone Toxicodendron radicans (L.) Kuntze [Rhus r. L.] Vv F F Toxicodendron vernix (L.) Kuntze O° oe Annonaceae Asimina triloba (L.) Dunal R NJE, S1 (NJ) Apiaceae Angelica atropurpurea L. 1/O Oo fk Angelica venenosa (G y) Fern vO VO $182 (NJ) Chaerophyllum procumbens (L.) Crantz var fe) fe) $3 (NJ) ocumbens Cicuta bulbifera L. R R 1/O Cicuta maculata var. maculata L. O F Cryptotaenia canadensis L. DC. O O * Daucus carota L. V F Heracleum maximum Bartr. R O Hydrocotyle americana L. R Vv Osmorhiza claytonii Michx. C. B. Clarke F/O F Osmorhiza longistylis (Torr.) DC. F Oxypolis rigidior (L.) Raf. R VO oO *Pastinaca sativa L. FO *Pimpinella saxifraga L. ee Le Sanicula canadensis L. var. canadensis F Sanicula marilandica L. F Sium suave Walter F/O F Taenidia integerrima(L.) Drude R - oO VINOLYVE OSPF R SUP LS LPM RIe RB SOB GC TS state status Zizia aptera (Gray) Fern. R cho Zizia aurea (L.) W. V. J. Koch Vo R Apocynaceae Apocynum androsaemifolium L. F O R Apocynum cannabinum L. r 39 Apocynum cannabinum L. var. hypericifolium [A. I sibiricum Jacq. Asclepias exaltata L. R R Asclepias incarnata L. ssp. incarnata Oooo Asclepias purpurascens L. R R/O Asclepias syriaca L. 47.20 Asclepias tuberosa L. Vv oO Asclepias variegata L. R $2 (NJ) PE, S1 (PA) Asclepias verticillata L. R R §2 (NJ) Asclepias viridiflora Raf. R *Vinca minor L. F/O F/O Aquifoliaceae Ilex laevigata (Pursh) A. Gray VO Ilex montana (T & G.) A. Gray R NJE, $1 (NJ) Ilex opaca Ait. R PT, S2 (PA) Ilex verticillata Ait. FO fs Nemopanthus mucronatus (L.) Trel. o VO. Vo Araliaceae *Aralia elata (Miq) Seem R tO Oo Avalia bispida L. F F/O R Ir dV9 VALVA FUVAV TIC FHL AO VYOTA UV INOSVA FHL AO .LSITHOAHO AUNV AAIAYHAO OSF R SUF LS LPM RIF RB SOB C_ TS statestatus Aralia nudicaulis L. Vv Aralia racemosa L. F Panax quinquefolius L. VO G3 G4, $2 (NJ) Panax trifolius L. F/O Aristolochiaceae Aristolochia serpentaria L. R $3 (NJ) Asarum canadense L. R R Asteraceae * Achillea gatas 1. ¥ 33 Ageratina altissima (L. King & H. E. Robins. Vv [Eupatorium rugosum Houtt.] Ambrosia artemisiifolia L F Vv we Ambrosia trifida L F F F a Anaphalis margaritacea (L.) Benth. & Hook. r © 3 y Antennaria neglecta Greene F oO z Antennaria plantaginifolia (L.) Richards F oO Arctium minus Schk. F Oo Oo *Artemisia vulgaris L. F F Bidens bipinnata L. FIO O O Bidens cernua L. F/O F/O F Bidens connata Muhl. ex Willd. oO oO F Bidens coronata L. Britt. R R Bidens frondosa L. F F *Carduus nutans L. F F OSF R SUF LS LPM Ei RB SOR C JTS state status *Centaurea stobe ssp. micranthus [C. maculosaLam.] _— F F O F *Cichorium intybus L. Vv Cirsium altissimum (L.) Cronq. F Oo O $X.1 (NJ) *Cirsinm arvense (L.) Scop. Vv F Cirsium discolor (Muhl. ex Willd.) Spreng. R Cirsium muticum Michx. F F Cirsium pumilum (Nutt.) Spreng. F Oo VO *Cirsium vulgare (Savi) Ten. Vv. FIO Conyza canadensis (L.) Crongq. var. canadensis F V Doellingeria infirma (Michx.) Greene [Aster infirmus R R Michx.] Doelling rbellata (P. Mill.) Nees [Aster O oO umbellatus P. Mill.] Erechtites hieraciifolius (L.) Raf. F F O Erigeron annuus (L.) Pers. F F Erigeron philadelphicus L. F F Erigeron pulchellus Michx. F F O Erigeron strigosus Muhl. ex Willd. var. strigosus Vv R Eupatorium altissimum L. R/O R/O R Eupatorium perfoliatum L. F/O Eupatorium sessilifolium L. R R Eurybia divaricata (L.) Nesom [Aster divaricatus L.] Vv Cr dVO VALVA TUVAV TAC FHL AO VYOTA UV TINOSVA SHL AO LSITIOAHO GNV ATIANAAO OSF KR Se (5 LP RIF RB SOB CT) meen «6 Eurybia macrophylla [Aster macrophyllus L.] 11f@) Eurybia schreberi (L.) Cass. [Aster s.] R Euthamia graminifolia (L.) Nutt. F F F Eutrochium dubium [Eupatorium d. Willd. ex Poir] aa: Extrochium fistulosum (Espatorium f.] Bartatt /O VO O Eutrochi, l [Euj jadelph | tO. F (L.) King & H. E. Robins. Expatorium m. L.] Eutrochium purpureum L. [Eupatorium p. L.] F F/O O *Galinsoga quadriradiata Ruiz & Pavon oo. oy *Gnaphalium uliginosum L. [Filaginella uliginosa(L.) R R ww Opiz.] a Helenium autumnale L. R R g Helianthus decapetalus L. SR - Helianthus divaricatus L. R R Helianthus strumosus L. R R R Helianthus tuberosus L. R R Heliopsis helianthoides L. [Helianthus h.] R R R *Hieracium caespitosum Dumort [H. pratense Tausch] V EO Hieracium paniculatum L. fe) *Hieracium pilosella L. R R Hieracium scabrum Michx. Hieracium venosum L. F OSF R SUF [IS LPM RIF RB SOB C TS state status Tonactis linariifolius (L.) Greene [Aster |. L.] R Krigia biflora (Walt.) S. F. Blake WO VO" Krigia virginica (L.) Willd. R R Lactuca biennis (Moench) Fern. F/O R Lactuca canadensis L. F F Lactuca floridana (L.) Gaertner var. floridana R R R *Lactuca serriola L. F V F *Lapsana communis L. R R ooh’ vulgare Lam. [Chrysanthemum Vv F oO leucanthemum L.] Liatris spicata (L.) Walt. var. spicata F/O KR $3 (NJ) * Matricaria discoidea DC. {M. matricarioides (Less.) VO Porter} Oclemena inata (Michx.) Greene [Aster VO acuminatus Michx. = Packera anonyma (Wood) W.A. Weber & A. Live R R R SU (NJ), PR, $2 [Senecio anonymus A. Wood (PA) Packera aurea (L.) A.& D. Léve [Senecio aureus L.] I Packera obovata (Muhl. ex Willd.) W.A. Weber & A. R R R Léve [Senecio obovatus Muhl. Packera paupercula (Michx.) A.& D. Léve [Senecio R R $3 (NJ) pau, i *Picris hieracioides L. R VO Prenanthes alba L. R R/O Prenanthes altissima L. R St dV9 YALVA FAVAV TAC FHL AO VYOTA UV INOSVA AHL AO LSITIOAHD UNV AAIAYTAO OSF BR SUR 48 42M RIF RB SOB -C TS wttesuns 8 Prenanthes trifoliolata (Cass) Fern. V F R Pseudognaphalium obtusifolium (L.) Hilliard & Burt. FF O [Gnaphalium o. L.] Rudbeckia hirta L. var. hirta Vv F Rudbeckia laciniata L.var. laciniata R R Rudbeckia triloba L.var. triloba R Senecio vulgaris L. Vv Sericocarpus asteroides (L.) B.S.P. [Aster paternus O O O Crongq.] Smallanthus uvedalius (L.) Mack. ex Small [Polymnia R NJE, $1 (NJ) u. Lam.] $3 (PA) 5 Solidago altissima L. [Solidago canadensis var. scabra V F a Torr. & Gray] re) Solidago bicolor L. F O c Solidago caesia L. Vv Solidago canadensis L. var. canadensis Vv F Solidago flexicaulis L. O Solidago gigantea L. var. gigantea 1/O Solidago juncea Ait. fe. De a Solidago nemoralis Ait. yy Or Solidago odora ssp. odora Ait. R R Solidago patula ssp. patula Muhl. ex Willd. © VO R Solidago puberula Nutt. R R R Solidago rugosa Mill. ssp. rugosa F F O OOSF KR. SUF LS. LPM Ri RB SOB C TS state status Solidago speciosa Nutt. var. speciosa Ro OR R PT, S2 (NJ, PA) Solidago uliginosa Nutt. R *Sonchus arvensis L. ssp. arvensis F Vv Symphyotrichum cordifolium (L.) Nesom [Aster Fk - 0 2. cordifolus L., Aster lowrieanus T. C. Porter] Symphyotrichum ericoides (L.) Nesom ssp. ericoides R $182 (NJ), $3 [Aster e. L.] (PA) Symphyotrichum laeve var. laeve (L.) A.& D. Live I F F Nesom [Aster Jaevis var laevis L.] t lanceolatum F oO oO a ) Nesom Asi lanceolatus Willd., Aster simplex Willd.] Symphyotrichum lateriflorum (L.) A.& D. Léve [Aster F/O F/O F lateriflorus (L.) Britt., Aster vimineus Lam.] Symphyotrichum novae-angliae (L.) N [Astern.L.] F F ae patens var. patens (Ait.) Nesom F O [Aster p. Ait.] aie: pilosum var. pilosum (Willd.) Nesom O Vv O [Aster pilosus Willd.] Symphyotrichum pilosum var. pringlei (Gray) Nesom [Aster pilosus Willd. var. demotus, Aster pilosus var. pringlei (Gray) Blake] Symphyotrichum [st gna (Muhl. ex Willd.) R §2 (NJ) Nesom [Aster p. Muhl.] VO W/O Symphyotrichum puniceum (L.) A.& D. Léve [Aster puniceus § } ich (Ell.) Nesom [Aster ~ tf racemosus Ell.] Ze dV9 WALVA FUVAV TAC FHL AO VYOTA UV TNOSVA FHL AO LSITIOAHD GNV AFTIAYTAO Osr K SF LS LPM RIF RB SOB. C state status Symphyotrichum trad (L.) Nesom [Aster R $2 (NJ) lateriflorus (L.) Britt. var. hirsuticaulis, Aster t. L.] *Tanacetum vulgare L. rR VO Taraxacum officinale G. H. Weber ex Wiggers F Vv *Tragopogon dubius Scop. F * Tragopogon porrifolius L. R R *Tragopogon pratensis L. R R *Tussilago farfara L. fe) Verbesina alternifolia (L.) Britt. ex Keamey R Vernonia noveboracensis (L.) Michx. F. WO *Xanthium strumarium L. oO: of Balsaminaceae Impatiens capensis Meerb. : O Impatiens pallida Nutt. O° 60 Berberidaceae *Berberis thunbergii DC. O F F *Berberis vulgaris L. FO “O° oF Caulophyllum thalictroides (L.) Michx. R Podophyllum peltatum L. F Betulaceae Alnus incana (L.) Moench. ssp. rugosa (Du Roi) VO Clausen [A. rugosa (Du Roi) Spreng. ] Alnus serrulata (Aiton) Willd. 2. “VO Alnus viridis ssp. crispa (Ait.) Turrill R PE, $1 (PA) Betula alleghaniensis Britt. [B. lutea Michx.] 11@) Betula lenta L. FE 8h VINOLYVE OSF KR SUF 15 LPM RF KB SOD € TS Same vtatus Betula nigra L. HO... F Betula papyrifera Marsh. R S2 (NJ) Betula populifolia Marsh. Vv O Carpinus caroliniana Walter F Corylus americana Walter F Corylus cornuta Marsh. fe) Ostrya virginiana (Miller) K. Koch VO Bignoniaceae *Campsis radicans (L.) Seem. R28 *Catalpa speciosa (Warder) Warder ex Engelm. R R Boraginaceae *Echium vulgare L. RB TG Hackelia virginiana (L.) I. M. Johnson F/O Hydrophyllum canadense L. VO NJE, $1 (NJ) Hydrophyllum virginianum L, 1V/O Mertensia virginica (L.) Pers. 1l/O O * Myosotis arvensis (L.) Hill. R R Myosotis laxa Lehm, V/O * Myosotis scorpioides L. R O Myosotis verna Nutt. R R *Symphytum officinale L. Rn. 8 Brassicaceae *Alliaria petiolata (Bieb.) Cavara & Grande V Vv F * Arabidopsis thaliana (L.) Hegnh. V Arabis canadensis L. YO.78 6¢ dVO YALVA FTUVAV TAC FHL AO VYOTA UV TNOSVA FHL AO LSITIOTHO AGNV AAIAYAAO SUF LS LPM RIF RB SOB C TS _ state status Arabis glabra (L.) Bernh. Arabis laevigata (Muhl.) Poir. var. laevigata Arabis lyrata L. Arabis missouriensis Greene *Armoracia rusticana P. G. Gaertn., B. Mey & Scherb. *Barbarea vulgaris R. Br. * Berteroa incana (L.) DC. *Brassica juncea (L.) Czernj * Brassica nigra (L.) Koch *Capsella bursa-pastoris (L.) Medik. Cardamine bulbosa (Schreb. ex Muhl) B. S. P. Cardamine concatenata (Michx.) 0. Schwartz Cardamine diphylla (Michx.) A. Wood Cardamine douglassii Britt. Cardamine pensylvanica Muh. ex Willd. *Cardamine pratensis L. Cardamine rotundifolia Michx. *Cardaria draba (L.) Desv. *Coincya monensis (L.) Greuter & Burdet *Erysimum cheiranthoides L. * Hesperis matronalis L. ee V/O R__ $1.1 (NJ), PE, $1 (PA) $3 (NJ) $2 (NJ) NJE, S1 (NJ) 0s VINOINVd OSF R SUF LS LPM RIF RB SOB C TTS _ Statestatus *Lepidium campestre (L.) R. Br. oO YO * Lepidium densiflorum Schrad. R * Lepidium ruderale L. R Lepidium virginicum L. F Vv Nasturtium officinale Ait. f. [Rorippa nasturtium- 11f@) aquaticum (L.) Hayek] Rorippa palustris (Butters & Abbe) var. glabra (O. E. F Schulz) R. Stuckey [R. islandica (Oeder) Borbas]} *Rorippa sylvestris (L.) Besser R *Sisymbrium officinale (L.) Scop. R *Thlaspi arvense L. Vo Cabombaceae Brasenia schreberi J. F. Gmel. 18) *Cabomba caroliniana A. Gray R Cactaceae Opuntia humifusa (Raf.) Raf. F PR, $3 (PA) Campanulaceae Campanula americana (L.) Small [Campanulastrum R americanum (L.) Small] Campanula aparinoides Pursh R R *Campanula rapunculoides L. R Campanula rotundifolia L. R R Lobelia cardinalis L. F F/O F Lobelia inflata L. F/O F Lobelia kalmii L. R R R PE, Si (PA) Lobelia siphilitica L. R Oo VO 1S dV9 WALVA FYVAV TAC AHL AO VYOTA UV TINOSVA AHL AO LSTTXHOAHSO UNV AAIAYTAO OSF R SUF LS LPM RIF RB SOB C_ TS | state status Lobelia spicata Lam. var. spicata R R Triodanis aan = y aah var. perfoliata O F [Specularia p.(L.) A Cannabaceae Celtis occidentalis L. R F Humulus lupulus L. R R R Caprifoliaceae Diervilla lonicera P. Mill. I/O Lonicera canadensis Barter. ex Marsh. R NJE, $1 (NJ) Lonicera dioica L. var. dioica R R *Lonicera japonica Thunb. F F O *Lonicera maackii (Rupr.) Maxim VO VO 38 *Lonicera morrowti A. Gray V/O I/O R Lonicera sempervirens L. R *Lonicera tatarica L, R R Symphoricarpos albus (L.) Blake. var. albus R R Symphoricarpos orbiculatus Moench R 1/O Triosteum aurantiacum Bickn. R R Triosteum perfoliatum L. R Caryophyllaceae *Arenaria serpyllifolia L. F Cerastium arvense L. ssp. arvense y iy O *Cerastium beeringianum Cham. & Schlecht. *Cerastium vulgatum L.[C. fontanum ssp. triviale] F *Dianthus armeria L. F cs VINOIYVE OSF R SUF LS LPM RIF RB SOB C__ TS | state status Moebhringia lateriflora (L.) Fenzl VO 1/O Myosoton aquaticum(L.) Moench [Stellaria aquatica V/O R (L.) Scop.] Paronychia canadensis (L.) A. Wood F Sagina procumbens L. F *Saponaria officinalis L. F O *Scleranthus annuus L. F R Silene antirrhina L. F Oo 22 *Silene latifolia Poir. ssp. alba (S. pratensis (Rafn.) F O Godr. & Green.] Silene nivea (Nutt.) Otth. R R NJE, S1 (NJ) Silene stellata (L.) Ait. f. R R R *Silene vulgaris (Moench) Garcke R/O R *Stellaria alsine Grimm R *Stellaria graminea L. VO VO Stellaria pubera Michx. R NJE, SH (NJ) Celastraceae *Celastrus orbiculatus Thunb. F F Celastrus scandens L. R Euonymus atropurpureus Jacq. VO fe) S1? (NJ) Ceratophyllaceae Ceratophyllum demersum L. VO Chenopodiaceae — *Chenopodium album L. var. album Vv *Chenopodium ambrosioides L. O F *Chenopodium pumilia R. Br. R €¢ dVS WALVA FUVAV TAC JH.L AO VUOTA UV TINOSVA AHL AO LSITIOTHO UNV MAIAYATAO OS F. Sue. LS. LEM A RS state status * Dysphania ambrosioides L. [Chenopodium a. L.] be * Dysphania pumilio R. Br. [C. pumilio R. Br.] R Cistaceae Helianthemum canadense (L.) Michx. V/O Lechea intermedia Leggett ex Britt. VO $2 (NJ) Lechea tenuifolia Michx. NJE, $1 (NJ) Clethraceae Clethra alnifolia L. Vv F Convolvulaceae —_ Calystegia sepium (L.) R. Br. F F Convolvulus arvensis L. F F Cuscuta gronovii Willd. V/O Ipomoea pandurata (L.) G. F. W. Mey. F/O Cornaceae Cornus alternifolia L. O Cornus amomum Mill. ssp. amomum F P Cornus amomum ssp. obliqua (Raf.) J. S. Wilson F F Cornus canadensis L. R R $1S2 (NJ) Cornus florida L. O F Cornus racemosa Lam. iy F O Cornus rugosa Lam. R Cornus sericea L. [C. stolonifera Michx.] R 1/O Crassulaceae *Sedum acre L. V/O *Sedum sarmentosum Bunge R R *Sedum spurium M. Bieb. +S VINOIUNVa OSF R SUF LS LPM RIF RB SOB C 4&5 _ state status *Sedum telephium L. R R Sedum ternatum Michx. Cucurbitaceae Echinocystis lobata (Michx.) T. & G. R VO l/O Sicyos angulatus L. R R R Dipsacaceae * Dipsacus sylvestris Hudson F R Droseraceae Drosera intermedia Hayne R Drosera rotundifolia L. VO Ebenaceae Diospyros virginiana L. R Elaeagnaceae * Elaeagnus umbellata Thunb. Vv Ericaceae b daphne calyculata (L.) Moench. var. F angustifolia Chimaphila maculata (L.) Pursh Vv Epigaea repens L. RR: YO Gaultheria procumbens L. F Gaylussacia baccata (Wangenh) K. Kotch F Gaylussacia dumosa (Andr.) T. & G. PE, SH (PA) Gaylussacia frondosa (L.) T. & G. 1/O Kalmia latifolia L. F Kalmia polifolia Wangenh. R NJE, $1 (NJ) Leiophyllum buxifolium (Berg.) Ell. PX, SX (PA) Leucothoe racemosa (L.) Gray R Lyonia ligustrina (L.) DC. F Monotropa hypopithys L. R ¢S dV9 UWALVA AUVAV TAC AHL AO VAOTA UV TNOSVA AHL AO LSITIOAHO GNV AAIAYATAO OSF KR Sur tS Lem Rif RB SOB © FS. state status Monotropa uniflora L. vO Orthilia secunda (L.) House vo $2 (NJ) Pyrola americana Sweet [P. rotundifolia L.] F Pyrola chlorantha Sw. R NJE, S1 (NJ, Pyrola elliptica Nutt. F/O is Rhododendron canadense (L.) Torr. Vo R NJE, $1 (NJ) Rhododendron maximum L, e. 6 Rhadodendron pervicl be eitk Chtnn R Rhododendron prinophyllum (Small) Millais R S3 (NJ) Rhododendron viscosum (L.) Torr. F F Vaccinium angustifolinm Ait. O F Vaccinium corymbosum L. ie) F Vaccinium macrocarpon Ait. F/O Vaccinium myrtilloides Michx R Vaccinium oxycoccos L. R $2 (NJ) Vaccinium pallidum L. F F Vaccinium stamineum L. F F Euphorbiaceae —- Acalypha rhomboidea Rat. 2 Fo Acalypha virginica L. Oo FO R R * Euphorbia cyparissias L. F/O F/O R Euphorbia maculata L. [Chamaesyce m. (L.) Small] V * Euphorbia marginata Pursh. R wn fon VINOINVa Ur LS LPM REE RB SOB C TS state status Euphorbia nutans Lag. [Chamaesyce n. (Lag.) Small] Fabaceae *Amorpha fruticosa L. Amphicarpaea bracteata (L.) Fern. Aptos americana Medik. Baptisia tinctoria (L.) R. Br. Cercis canadensis L.' Chamaecrista nictitans (L.) Moench. Crotalaria sagittalis L. Desmodium canadense (L.) DC. Desmodium canescens (L.) DC. Desmodium ciliare (Muhl.) DC. Desmodium cuspidatum (Muhl.) Loudon Desmodium glutinosum (Muhl.) A. Wood Desmodium marilandicum (L.) DC. Desmodium nudiflorum (L.) DC. Desmodium paniculatum (L.) DC. Desmodium rotundifolium DC. Desmodium viridiflorum (L.) DC. Gleditsia triacanthos L. Gymnocladus dioicus (L.) K. Koch *Lathyrus latifolius L. Lathyrus palustris L. " Material my be introduced. I/O F/O R 1/O Oo E F/O ‘y R R R R V/O r 1/O O R R R NJE, S1 $2 (NJ), SU (PA) ZS dVO YALVA FTUVAV TAG AHL AO VUOTA UV TNOSVA AHL AO LSITIOAHO GNV AAIAYAAO OSF R SUF LS LPM RIF RB SOB C [5 __ state status Lespedeza capitata Michx. F/O O * Lespedeza cuneata (Dum.) G. Don. F F Lespedeza hirta (L.) Hornem F F/O Lespedeza procumbens Michx. vO F..-9 Lespedeza repens (L.) Barton R Lespedeza violacea (L.) Pres. R Lespedeza virginica (L.) Britt. R R * Lotus corniculatus L. I F * Medicago lupulina L. F I *Melilotus alba Medik. pee * Melilotus officinalis (L.) Lam. F F Phaseolus polystachios (L.) BSP. R $2 (NJ), $182 (PA) Robinia pseudoacacia L. O F F Senna hebecarpa (Ferrnald) Irwin & Barneby R Tephrosia virginiana (L.) Pers. VO * Trifolium arvense L. F F * Trifolium aureum Pollich R I * Trifolium campestre Schreb. I I * Trifolium dubium Sibth. I F * Trifolium hybridum L. F Vv R I * Trifolium pratense L. VINOLUV4 OSE ..R.. SUR..0S . DPM RI RB. SOs. CS. same stars * Trifolium repens L. 7 Y¥ Vicia americana Muhl. F F $1 (NJ) "Vicia cracca L. F F O *Vicia tetrasperma (L.) Moench. VO VO R * Wisteria sinensis (Sims) Sweet R R Fagaceae Castanea dentata (Marsh.) Borkh. Vo *Castanea mollissima Blume R *Castanea sativa P. Mill. R Fagus grandifolia Ehrh. F *Fagus sylvatica L. R Quercus alba L. F Quercus bicolor Willd. F Quercus coccinea Muenchh. F Quercus ilicifolia Wangenh. Vv oO Quercus montana Willd. (Q. prinus L.] F Quercus mublenbergii Engelm. R $3 (NJ) Quercus palustris Muenchh. oO F Quercus prinoides Willd. I RR R Q bra L. var. ambigua (Gray) Fern. [Q. F borealis Michx. f.] Quercus rubra var. rubra L. (Q. borealis Michx. f.] F Quercus stellata Wangenh. R 6S dV9 WALVA FAVAV TAC AHL AO VUOTA UV INOSVA FHL AO LSITIOAHSD GNV AAIAWIAO OSF R SUF IS LPM RIF RB SOB C TS . state status Quercus velutina Lam. Vv Gentianaceae Bartonia virginica (L.) B.S. P. R Gentiana linearis Froel. R NJE, SH (NJ) Gentianella quinquefolia L. R R $2 (NJ) Gentianopsis crinita (Froelich) Ma. R R Geraniaceae *Erodium cicutarium (L.) LHer. Re Geranium carolinianum L. R 6 Geranium maculatum L. Vv *Geranium robertianum L. F *Pelargonium X hortorum Baily (pro sp.) R Grossulariaceae = Ribes americanum P. Mill. VO Ribes cynosbati L. F/O SU (NJ) Ribes glandulosum Graurer R NJE, $1.1 (NJ) Ribes hirtellum Michx. R Ribes rotundifolium Michx. R *Ribes rubrum L. [R. sativum Syme] R Ribes uva-crispa L.var. sativum [R. grossularia L.] R Haloragaceae Myriophyllum farwellii Morong R PE, $3 (PA) Myrtophyllum heterophyllum Michx. R $2 (NJ), PE, $4 Myriophyllum humile (Raf.) Morong R ro *Myriophyllum spicatum L. VO FP Proserpinaca palustris L. var. palustris on) oO VINOLYVa OSF EK Me ts LPM RIF RB SOR CC TS state status Hamamelidaceae Hydrangeaceae Hypericaceae Juglandaceae Lamiaceae Hamamelis virginiana L. *Deutzia scabra Thunb. Hydrangea arborescens L. Hypericum ascyron L. Hypericum ellipticam Hook. Hypericum gentianoides (L.) B. S. P. Hypericum mutilum L. * Hypericum perforatum L. Hypericum prolificum L. Hypericum punctatum Lam. Hypericum pyramidatum Ait. Triadenum fraseri (Spach) Gleason Triadenum virginicum (L.) Raf. Carya cordiformis (Wangenh.) K. Koch Carya glabra (Miller) Sweet Carya ovalis (Wangenh.) Sarg. Carya ovata (Miller) K. Koch — tomentosa (Lam. ex Poir.) Nutt. [C. alba (L.) El Nutt. 1] he. cinerea L. Juglans nigra L. Agastache nepetoides (L.) Kuntze Pe a Pee ee V 1/O F/O I/O 1/O 1/O F/O I/O O 1l/O $2 (NJ) NJE, S1 (NJ) $3 (NJ) $3 (NJ) $2 (NJ) 19 dVO WALVA AUVAV TAC FHL dO VYOTA UV TINOSVA AHL AO LSITIOAHO GNV AFIAYTAO OSF R SUF LS LPM RIF RB SOB C._ TS § sstate status Agastache scrophulariifolia (Willd.) Kuntze Re $2 (NJ) * Ajuga reptans L. I I Collinsonia canadensis L. F Cunila origanoides (L.) Britt. R *Galeopsis bifida Boenn. [G. tetrahit L.] F *Glechoma hederacea L. F Hedeoma pulegioides (L.) Pers. I I *Leonurus cardiaca L. F Lycopus americanus Muh. ex. W. Bart. F Lycopus uniflorus Michx. Lycopus virginicus L. Mentha arvensis L. F * Mentha spicata L. F *Mentha X rotundifolia (L.) Hudson R Monarda didyma L.' $2 (NJ) Monarda fistulosa L. I I *Nepeta cataria L. I I * Physostegia virginiana (K.) Benth. Pycnanthemum incanum (L.) Michx. F Pycnanthemum muticum (Michx.) Pers. R R I Pycnanthemum tenuifolium Schrad. * Material my be introduced. a No VINOLYVE SUF LS LPM RI RB SOB C TS State status Lardizabalaceae Lauraceae Lentibulariaceae Limnanthaceae Linaceae Lythraceae Magnoliaceae Malvaceae } IY OUND, (L ) Durand & B.D Jackson Scutellaria galericulata L. Scutellaria incana Biehler Scutellaria lateriflora L. *Stachys palustris L. var. palustris Teucrium canadense . virginicum (L.) Eat. Trichostema dichotomum L. *Akebia quinata (Houtt.) Decne Lindera benzoin (L.) Blume Sassafras albidum (Nutt.) Nees. Utricularia macrorbiza Le Conte [U. vulgaris L.] Utricularia purpurea Walt. Floerkea proserpinacoides Willd. Linum striatum Walt. Linum virginianum L. Decodon verticillatus (L.) Ell *Lythrum salicaria L. Liriodendron tulipifera L. “Magnolia macrophylla Michx. *Abutilon theophrastii Med. Hibiscus moscheutos L. (H. palustris L., Malva m. L.] F/O F/O R VO F/O v/O F/O $3 (NJ), PR (S4) €9 dVD WALVA TUVAV TAC FHL AO VAOTA UV TINOSVA FHL AO LSITIOFHS GNV AAIAUTAO Ost KR GUP 25 LOM RIF KB SOB C tS state status Menispermaceae Menyanthaceae Molluginaceae Moraceae Myricaceae Myrsinaceae Nyctaginaceae Nymphaeaceae *Malva neglecta Wallr. Menispermum canadense L. Menyanthes trifoliata L. Nymphoides cordata (Elliott) Fern. *Mollugo verticillata L. *Morus alba L. Morus rubra L. Comptonia peregrina (L.) J. M. Coulter Myrica gale L. Myrica pensylvanica Mirbel. *Anagallis arvensis L. Lysimachia ciliata L. Lysimachia hybrida Michx. Lysimachia lanceolata Walt. * Lysimachia nummularia L. Lysimachia quadrifolia L. Lysimachia terrestris (L.) B. S. P. Trientalis borealis Raf. % Ad sw olecls: ~ re pee | \ Mac Millan < ° x iv Nuphar advena (Ait.) Kartesz & Gandhi (N. lutea ssp. advena (Ait.) Ait. f.] V/O O PR F V F/O R R R R /O © F F R 1/O R/O $3 (NJ), PT, $2 (PA) $3 (NJ), PT, S2 (PA) $3 (NJ), S1 (PA) S1 (NJ) an ac VINOLYVa OSF. R sur DS LPM ‘RIF (RB SOB 43) state status Nuphar po 280 (Pers.) Fern. [N. lutea ssp. pumila (Timm) E. O. Beal] Nuphar variegata Dur. [N. lutea ssp. variegata (Dur.) E. O. Beal] Nymphaea odorata Ait. Nyssaceae Nyssa sylvatica Marsh. Oleaceae Chionanthus virginicus (L.)! *Forsythia viridissima Lindl. Fraxinus americana L. Fraxinus nigra Marsh. Fraxinus pennsylvanica Marsh. *Ligustrum vulgare L. Onagraceae Circaea alpina L. Circaea canadensis (L.) Aschers. & Magnus ssp. canadensis (C. lutetiana ssp. canadensis (L.) Aschers. & Magnus, C. quadrisulcata var. canadensis (L.) Hara] Epilobium ciliatum Rat. [E. ciliatum ssp. glandulosum (Lehm.) Hoch & Raven, E. glandulosum Lehm.] Epilobium coloratum Biehler Epilobium leptophyllum Rat. Epilobium strictum Muhl. ex Spreng. Gaura biennis L. Ludwigia alternifolia L. Ludwigia palustris (L.) Ell. ' Material my be introduced. 1V/O F/O F/O 1V/O 1/O F/O V/O F/O NJE, SH (NJ), $1 (PA) 53 (NJ, PA) $2 (NJ) S2 (NJ), PE, $3 (PA) 53 (NJ) 69 dVD UALVA FUVAV TAG AHL AO VYOTA UV TINOSVA FHL AO LSITIOTHO GNV MATAYHAO OSF R SUP ES 12M RIF RB SOB C «TS state status Orobanchaceae Oxalidaceae Papaveraceae Oenothera biennis L. Oenothera perennis L. aad paupercula (Gray) Britt. [Gerardia p. (Gray) itt.) ae purpurea (L.) Pennell Agalinis tenuifolia (Vahl.) Raf. [Gerardia t. Vahl.] Aureolaria flava (L.) Farw. var. flava Aureolaria pedicularia (L. ) Raf. [Gerardia p. L.] Aureolaria virginica (L.) Pennell Castilleja coccinea (L.) Sprengel Conopholis americana (L.) Wallr. Epifagus virginiana (L.) Barton Melampyrum lineare Dest. var. americanum Orobanche uniflora L. Pedicularis canadensis L. Oxalis stricta L. [O. fontana Bunge] Oxalis violacea L. Adlumia fungosa (Ait.) Greene *Chelidonium mayus L. Corydalis flavula (Raf.) DC. Corydalis sempervirens (L.) Pers. Dicentra canadensis (Goldie) Walp. 1 ve FO - R $2 (NJ), PE, S1 (PA) $2 (NJ, PA) S2 (NJ) NJE, $1 (NJ) lo oO VINOLYVd O6F -R-..SUP...LS..LPM.-RIEF-. RBSOB....C TS state status Parnassiaceae Paulowniaceae Penthoraceae Phrymaceae Phytolaccaceae Plantaginaceae Dicentra cucullaria (L.) Bernh. Dicentra eximia (Ker Gawler) Torr. Sanguinaria canadensis L. Parnassia glauca Rat. *Paulownia tomentosa (Thunb.) Steudel Penthorum sedoides L. Mimulus alatus Ait. Mimulus moschatus Douglas Mimulus ringens L. Phryma leptostachya L. Phytolacca americana L. Callitriche palustris L. Callitriche terrestris Raf. [C. deflexa A. Braun] Chelone glabra L. *Cymbalaria muralis P. Gaertner, Meyer & Scherb. * Digitalis purpurea L. Gratiola neglecta Torr. Linaria canadenis (L.) Chaz. [Nuttallanthus c. (L.) D. A.S utton *Linaria vulgaris P. Mill. Lindernia dubia (L.) Pennell var. dubia Penstemon canescens (L.) D. A. Sutton F/O R F/O FE R F I/O F/O F/O F/O F/O NJE, SH.1 (NJ), PE, S1 (PA) PE, $2 (PA) $3 (NJ) $2 (NJ) $2 (NJ) Z9 IVD VALVA FUVAV TAC AHL AO VUOTA UV TNOSVA FHL AO LSITHOAHSO UNV AATAYAAO OSF R Sum LS LPM RIF RB SOB. C IS _ statestatus Penstemon digitalis Nutt. YO. & Penstemon hirsutus (L.) Willd. O RR 4A *Plantago aristata Michx. F Oo *Plantago lanceolata L. F Vv *Plantago major L. F Vv Plantago rugelii Dene. ee Plantago virginica L. R R *Veronica agrestis L. R Veronica americana (Raf.) Schwein. R I VO *Veronica chamaedrys L. Rok *Veronica longifolia L. R * Veronica officinalis L. R F R Veronica peregrina L. ssp. peregrina R. WO Veronica scutellata L. co 37> Veronicastrum virginicum (L.) Fern. R R Platanaceae Platanus occidentalis L. oO F Podostemaceae Podost tophyllum Michx. $2 (NJ) Polemoniaceae Phlox divaricata L. ssp. divaricata R R NJE, S1 (NJ) Phlox paniculata L. VO V/O Phlox subulata L. ssp. subulata VO VO Polygalaceae Polygala paucifolia Willd. F P F/O Polygala sanguinea L. 89 VINOLYVE OOSF RK SUP 1S LEM Rip RE S05 G TS state status Polygonaceae Polygala verticillata L. var. verticillata Fallopia cilinodis (Michx.) Holub [Polygonum cilinode Michx.] *Fallopi. loulus (L.) A. Love [Polygonum c. L.] *Fallopia j a ‘nae ) Dene. [Polygonum cuspidatum Sie Fallopia —o a) ci [Polygonum s. L.] patie amphibia (L.) S.F. Gray p.p. [Polygonum ibium var. emersum Michx.] Persicaria arifolia [Polygonum arifolium L.] Persicaria hydropiperoides (Michx.) Small [Polygonum h, Michx. var. hydropiperoides| * Persicaria lapathifolia (L.) S.F. Gray [Polygonum lapathifolium L.] *Persicaria longiseta (de Bruyn) Moldenke — caespitosum Blume. %*D ] [Dp]. “ 4 78 P L.] *Persicaria orientalis (L.) Spach [Polygonum orientale {. Persicaria a (L.) G. Maza [Polygonum pensylvanicum L.] Persicaria punctata (Ell.) Small [Polygonum punctatum Ell.) Persicaria sagittata (L.) Gross [Polygonum sagittatum L.] oleae virginiana (L.) Gaertn. [Polygonum virginianum L.] Polygonum aviculare L. z O O O F 12/2 ea Vv F # 1/O $2 (NJ) 69 dVO UALVA TAVAV TAC AHL AO VUOTA UV INOSVA AHL AO LSITIOTHD AGNV AAIAYAAO OSF R SUF LS LPM RIF RB SOB C_ TS. statestatus © R Feheonae tenue Michx. R *Rumex acetosella L. ¥ F *Rumex crispus L. F F/O *Rumex obtusifolius L. VO R Portulacaceae Claytonia virginica var. hammondiae (Kalmbacher) R NJE, G5T1, $1.1 Doyle, Lewis & Snyder (NJ) Claytonia virginica L. var. virginica R F *Portulaca oleracea L. V Ranunculaceae —Actaea pachypoda Ell. [A. alba (L.) Miller] F Actaea racemosa L. [Cimicifuga r. L. Nutt.] R/O R Actaea rubra (Ait.) Willd. R $2 (NJ) e Anemone americana (DC.) Hara [Hepatica a. (DC.) vO o Ker-Gawl., Hepatica nobilis var. obtusa (Pursh) £ Anemone quinquefolia L. F Anemone virginiana var. alba (Oakes) Wood Oo Anemone virginiana var. virginiana L. ie) Aquilegia canadensis L. R VO R Caltha palustris L. O FO F/O Clematis occidentalis (Hornem.) DC. [C. verticillaris R/O R $2 (NJ) es terniflora DC. R R Clematis virginiana L. R R FO Coptis trifolia (L.) Salis. ees OSF R SUF LS LEM RIF KB SOB: © TS _ state status Rhamnaceae Rosaceae Ranunculus abortivus L. *Ranunculus acris L. Ranunculus aquatilis L. var. diffusus [R. longirostris Godr.]} +P an lee bulba us L Ranunculus fascicularis L. corer: hispidus Michx. var. caricetorum [R. ricetorum Greene, R. septentrionalis Poir.] Ranunculus hispidus var. hispidus Michx. Ranunculus recurvatus Por. *Ranunculus repens L. Ranunculus sceleratus L. Ranunculus trichophyllus Chaix. Thalictrum dioicum L. Thalictrum pubescens Pursh Thalictrum revolutum DC. Thalictrum thalictroides (L.) Eames & Boivin [Anemonella t. (L.) Spach] Trollius laxus Salisb. subsp. laxus Ceanothus americanus L. Rhamnus alnifolia VHer. Agrimonia gryposepala Wally. Agrimonia parviflora Ait. r O r F F/O R $3 (PA) F R/O R R NJE, $1 (NJ), PE, $1S2 (PA) ie) F R F/O $2 (NJ) F/O F o <6 R R ie) :. & NJE, PE, G4 73, S1 (NJ, PA) ie) O R O fe) F/O ie) IZ dV9 UALVA FUVAV TAC FHL AO VYOTA UV INOSVA AHL AO LSITIOAHD AUNV AAIAUATAO OSF R SUF LS LPM RIF RB SOB C TS State status Agrimonia rostellata Wallr. R R Agrimonia striata Michx. V/O Amelanchier arborea (Michx. f.) Fern. VO Amelanchier laevis Weig. VO O Crataegus coccinea L. R 1/O R Crataegus intricata Lange R R Crataegus punctata Jacq. R R $2 (NJ) Crataegus succulenta Schrad. ex Link R R R NJE, $1 (NJ) * Duchesnea indica (Andrews) Focke R/O F Filipendula rubra (Hill) B. L. Robinson R R NJE, SX (NJ), $1S2 (PA) *Fragaria vesca ssp. vesca L. R Fragaria virginiana Duchesne F Geum aleppicum Jacq. Fr FQ Geum canadense Jacq. F/O F Geum laciniatum Murray R R Geum rivale L. $3 (NJ) Gillenia trifoliata (L.) Moench R Photinia melanocarpa (Michx.) Robertson & Phipps VO VO [Aronia m. (Michx.) Ell., Sorbus m. (Michx.) Heynh.] Photinia pyrifolia (Burm. f.) Nakai [Aronia arbutifolia R/O F R (L.) Pers., Sorbus arbutifolia (L.) Heynh.] R R Physocarpus opulifolius (L.) Maxim. VINOLUVd OSF R SUF LS LPM RIF RB SOB C TS state status *Potentilla argentea L. F O Potentilla arguta Pursh. R R $2 (NJ) Potentilla canadensis L. r F/O Potentilla fruticosa L. [Dasiphora f. auct. non (L. PE S1 (PA) Rydb., Pentaphylloides floribunda (Pursh) A. Love nom. super. } Potentilla norvegica L. ssp speliensis (L.) Ascher. R R & Graebn. *Potentilla recta L. x Potentilla simplex Michx. Vv F Prunus americana Marsh. R *Prunus avium L. 1V/O Prunus pensylvanica L. f. Oo F * Prunus persica (L.) Batsch. R Prunus pumila L. var. depressa (Pursh) Bean [P. $1 (NJ, PA) depressa Pursh] Prunus pumila L. var. pumila Prunus serotina Ehrh. Vv F Prunus virginiana L. O Oo VO *Pyrus communis L. [P. pyrifolia (Burm. f.) Nakai] R *Rosa canina L. R Rosa carolina L. F *Rosa eglanteria L. ae *Rosa gallica L. R €Z dV9 VALVA FUVAV TAG FHL AO VOTH UV TNOSVA AHL AO LSITIOAHS GNV AAIAYAAO OSF R SUF LS LPM RIF RB SOB C._ TS _ state status *Rosa multiflora Thunb. Vv F F/O Rosa palustris Marsh. F Vv Rosa virginiana Mill. R R S1 (PA) *Rosa wichuraiana Crép. R Rubus allegheniensis Porter F O Rubus canadensis L. Vo NJE, $1 (NJ) Rubus flagellaris Willd. F F F/O Rubus hispidus L. F F Rubus idaeus L. ssp. strigosus (Michx.) Focke [R. VO VO strigosus Michx.] Rubus occidentalis L. F oO Rubus odoratus L. V/O R *Rubus phoenicolasius Maxim. O F/O FO Rubus pubescens Raf. aR Sanguisorba canadensis L. O I/O VO Sorbus americana Marsh. m2 $2 (NJ) *Sorbus aucuparia L. R Spiraea alba Du Roi F R $1 (NJ) Spiraea latifolia (Ait.) [S. alba var. latifolia (Ait.) F R Dippel] *Spiraea prunifolia Siebold & Zuce. R Spiraea tomentosa L. F F/O Spiraea X billardii Herincq. (pro sp.) [S. X vanhouttei R (Briot) Carr.] VINOLYVE OSE RK MP LS LPM Be BB SOR h. (SS satate status Waldsteinia fragarioides (Michx.) Tratt. R R $2 (NJ) Rubiaceae Cephalanthus occidentalis L. F/O Galium aparine L. F F Galium asprellum Michx. F O Galium boreale L. fF 2 VO Uo $3 (NJ) Galium circaezans Michx. var. circaezans 11f@) Galium lanceolatum Torr. O *Galium mollugo L. HO. OF Galium obtusum Bigelow R R Galium palustre L. $3 (NJ) Galium tinctorium L. F Galium triflorum L. R *Galium verum L. F O Houstonia caerulea (L.) Hook. [Hedyotis c. L.] F 1/O Mitchella repens L. Vv Salicaceae *Populus alba L. VO VO Populus balsamifera L. PE, S1 (PA) Populus deltoides Marsh. R VO Populus grandidentata Michx. F/O R *Populus nigra L. R Populus tremuloides Michx. F Populus X jackii Sarg. R CZ dVD VALVA FYVAV TAC AHL AO VYOTA UV TNOSVA AHL AO LSITIOAHO ANV AAIAWAAO iS 47M RIF: RE SOR CC 'S state status Santalaceae Sapindaceae *Salix alba L. Salix bebbiana Sarg. Salix candida Fluegge ex Willd. [S. candida var. enudata Andress.] *Salix caprea L. Salix discolor Muhl. Salix eriocephala Michx. [S. rigida Muhl.] Salix exigua Nutt. [S. interior Rowlee] *Salix fragilis L. Salix humilis Marsh. var. humilis Salix lucida Muhl. ssp. lucida Salix nigra Marshall Salix petiolaris L. *Salix purpurea L. Salix sericea Marshall Salix serissima (Bailey) Fern. Arceuthobium pusillum Peck Comandra umbellata (L.) Nutt. Acer negundo L. Acer nigrum Michx. f. Acer pensylvanicum L. VO F/O i i Hee 14 @ YD 1/O $2 (NJ), PT, S1 (PA) $12 (NJ) S2 (NJ), PT, S2 (PA) NJE, S1 (NJ), PT, S2 (PA) ™N oO VINOLUVE OSF R GUP i5 LPM RE RB SOB © TS state status Sarraceniaceae Saxifragaceae Scrophulariaceae Simaroubaceae Solanaceae *Acer platanoides L. Acer rubrum L. Acer saccharinum L. Acer saccharum Marshall Acer spicatum Lam. * Aesculus parviflora Walt. Sarracenia purpurea L. Chrysosplenium americanum Schwein. Heuchera americana L. Mitella diphylla L. Saxifraga pensylvanica L. Saxifraga virginiensis Michx. Tiarella cordifolia L. Scrophularia lanceolata Pursh Scrophularia marilandica L. *Verbascum blattaria L. *Verbascum lychnitis L. *Verbascum thapsus L. *Ailanthus altissima (Mill.) Swingle *Petunia X hybrida Vilm. [P. axillaris (Lam,) B. S. P.] *Physalis alkekengi L. 1/O F/O I/O iy V/O 1/O V/O V/O NJE, $1 (NJ) ZZ dV9 WALVA FUVAV TAC AHL AO VUOTA UV TINOSVA AHLL AO LLSITIOAUHO GNV AAIAYATAO OSF R “SUF LS LPM RIF RB SOB C TS state status Physalis subglabrata Mackenzie & ‘Bush [P. longifolia R var. subglabrata (Mackenzie & Bush) Cronq.] Solanum americanum Mill. F/O _F R Solanum carolinense L. F/O F/O *Solanum dulcamara L. FO: oO *Solanum melongena L. *Solanum nigrum L. F/O fF O Staphyleaceae Staphylea trifolia L. R/O R R R Thymelaeaceae Dirca palustris L. R $2 (NJ) Tiliaceae Tilia americana L. var. americana V/O Ulmaceae Ulmus americana L. R VO oO *Ulmus pumila L. | eS Ulmus rubra Muhl. R F/O R Urticaceae Boehmeria cylindrica (L.) Sw. var. cylindrica F/O F Laportea canadensis (L.) Wedd. V/O VO Parietaria pensylvanica Muhl ex Willd. F/O. R Pilea fontana (Lunell) Rydb. R Pilea pumila (L.) A. Gray R F/O *Urtica dioica ssp. dioica L. F Urtica dioica L. ssp. gracilis (U. procera Muhl. ex F/O Willd. Verbenaceae Verbena hastata L. F/O F/O Verbena urticifolia L. var. urticifolia VO VINOLYVG OSF R LS: LPM RIP RB SOB: -G TS state status Violaceae Vitaceae *Vitex agnus-castus L. Viola bicolor Pursh Viola blanda var. blanda Willd. Viola blanda Willd. var. palustriformis Gray [V. incognita Brainerd] Viola canadensis L. Viola cucullata Ait. Viola labradorica Schrank [V. conspersa Reichneb.] Viola macloskeyi F. Floyd ssp. pallens (Banks ex Ging.) MLS. Baker [V. pallens (Banks ex DC.) Brainerd] Viola palmata L. Viola pedata L. Viola pubescens Ait. var. pubescens Viola rostrata Pursh Viola rotundifolia Michx. il sagittata Ait. var. ovata (Nutt.) Torr. & Gray [V. ga sororia ae [V. papilionacea Pursh p.p.] Viola striata Ait. Parthenocissus quinquefolia (L.) Planch. Vitis aestivalis Michx. Vitis labrusca L. Vitis riparia Michx. Vitis vulpina L. 1/O V/O I $3 (NJ) V/O NJE, S1 (NJ) $3 (NJ) F/O V/O 6Z dV9D YALVA AUVAV TAC AHL AO VUOTA UV INOSVA FHL AO LSITIOAHOD ANV AAIAYATAO OSF R Slr LS CPM KIF RE SOB C TS: statestatus Monocots Acoraceae Acorus americanus (Raf.) Raf. R $1? (NJ), PE, S1 (PA) Acorus calamus L. R/O Agavaceae *Hosta lancifolia Engl. *Yucca flaccida L. [Y. filamentosa] R Alismataceae Alisma subcordatum Raf. F Alisma triviale Pursh R NJE, PE, S1 (NJ, PA) Sagittaria latifolia Willd. var. latifolia Vv Alliaceae Allium canadense L. V/O ler f Laren | eum L *Allinm schoenoprasum var. sibiricum (L.) Hartman Allium tricoccum Ait. F *Allium vineale L. F/O *Galanthus nivalis L. *Narcissus pseudonarcissus L. Araceae Arisaema dracontium (L.) Schott Vv F Ari phylum ss dsonii (Britt.) R $2 (NJ) Huttleston Arisaema triphyllum (L.) Schott ssp. triphyllum Vv Calla palustris L. R R $3 (NJ) Vv Lemna minor L. VINOIUVE Osr FF SUF Is LPM RIP RB SOB C TS state status Colchicaceae Commelinaceae Cyperaceae Lemna trisulca L. Orontium aquaticum L. Peltandra virginica (L.) Schott & Endl Spirodela polyrhiza (L.) Schleiden Symplocarpus foetidus (L.) Nutt. Wolffia borealis (Engelm. ex Hegelm.) Landolt ex Landolt & Wildi Wolffia brasiliensis Weddell [W. papulifera Griseb, W. ] punctata C. Uvularia perfoliata L. Uvularia sessilifolia L. *Commelina communis L. R Bulbostylis capillaris (L.) C. B. Clarke F/O Carex aestivalis Curtis ex Gray Carex albicans Willd. ex Spreng. Carex annectens (Bickn.) Bickn. Carex arctata Boott ex Hook. Carex argyrantha Tuckerm. Carex atlantica Bailey ssp. atlantica Carex baileyi Britt. V/O Carex bebbii Britt. 1/O Carex bicknellii Britt. 1/O 1/O 1/O VO R R O R V/O Vv R VO R R TOO V/O I/O $2 (NJ) PR, $4 (PA) NJE, $1 (NJ) S2 (NJ), PE, S1 PA $2 (NJ), PE St (PA) 18 dVD YALVA FUVAV TAC AHL AO VUOTA UV TINOSVA AHL AO LSITIOAHO GNV MAIAYAAO OS? R Gur 0S LPM RiP RB. SOB CC ES state patus Carex blanda Dewey VO R Carex bromoides Schkuhr ex Willd. R R Carex bushii Mackenzie R R NJE, $1 (NJ) Carex canescens L. var. canescens VO I/O Carex caroliniana Schwein R R SU (NJ) Carex cephalophora (Dewey) Dewey Vo $3 (NJ) Carex collinsti Nutt. R PE, S2 (PA) Carex communis Bailey Vo Carex comosa Boott 1/O Carex conoidea Schkurh ex Willd. F/O Vv $2 (NJ) Carex crawei Dewey R R NJE, $1 (NJ) Carex crawfordii Fern. VO YO $2 (NJ) Carex crinita Lam. var. crinita PN Carex cristatella Britt. F/O V Carex cryptolepis Mackenzie R $2 (NJ), PT, $1 Carex cumulata (Bailey) Fern. R NJE, - (NJ) Carex davisii Schwein & Torr. R Carex debilis Michx. var. debilis Vo Carex deweyana Schwein. var. deweyana R NJE, S1 (NJ) Carex digitalis Willd. R Carex disperma Dewey R S1S2 (NJ), PR, $3 (PA) oO i) VINOLUV OSF RK. SUF .LS PM RE RB SOB: .C AS state status Carex echinata Mutt. var. echinata R R Carex emoryi Dewey R R Carex flava L. R PI S2(PA) Carex folliculata L. VO Carex glaucodea Tuckerm. ex Olney. [C. flaccosperma V/O Dewe Carex gracillima Schwein. VO Carex granularis Muhl. ex Willd. var. granularis R Carex grayi Carey R Carex grisea Wahlenb. [C. amphibola var. turgida V F/O Fern.] Carex gynandra Schwein. F F Carex haydenii Dewey R R NJE, $1 (NJ), $1$2 (PA) Carex hirsutella Mackenzie [C. complenata Torrit R VO Hook.]} Carex hirtifolia Mackenzie R Carex hitchcockiana Dewey R S2 (NJ) Carex hystericina Muhl. ex Willd. R R Carex interior Baily E/Q.. oF Carex intumescens Rudge. F/O F Carex lacustris Willd. RO a I, g (Kul h.) Mackenzie R Carex lasiocarpa Ehrh. fe) PR, S3 (PA) €8 dVO WALVA AWVAV TAC AHL AO VYOTA UV TNOSVA FHL AO LSITHOAHO GNV AAIAYAAO OSE: Raga. te. cee AIP. BO ot: GC... 0S. ceeete eeatus Carex laxiculmis Schwein. var. laxiculmis R R Carex laxiflora Lam. R R Carex leptalea Wahlerb. 11f@) Carex leptonervia (Fern.) Fern, VO O NJE, S1 (NJ) Carex lupulina Muhl. ex Willd. F Vv Carex lurida Wahlerb. F F Vv Carex muhlenbergii Schkuhr ex Willd. VO *Carex muricata L. ssp. lamprocarpa R Carex nigromarginata Schwein 1/O Carex pellita Muhl. [C. lanuginosa Michx.] F/O ¥ Carex pensylvanica Lam. Vv Carex platyphylla Carey R Carex polymorpha Muhl. R NJE, PE, G3 (NJ, PA), S1 (NJ), S2 (PA) Carex prasina Wahlenb. R Carex radiata (Wahlenb.) Small. R Carex retroflexa Muhl. ex Willd. VO R Carex rosea Schkuhr ex Willd. [C. flaccidula Steud.] 1V/O Carex scabrata Schwein R Carex scoparia Schkuhr 1/O Carex siccata Dewey 7O VO2s NJE, S1 ae PE, S2 (PA V/O Carex sparganioides Muhl. ex Willd. co b VINOLUV a OSF R SUF LS LPM RiP RB SOS ©C F5 state eatus Carex sprengelit Dewey ex Spreng. R R S3 (PA) Carex squarrosa L. mm. VO © Carex sterilis Willd. R $2 (NJ), PT, St (PA) Carex stipata Muhl. ex Willd. var. stipata F V Carex stricta Lam. O F Carex swanii (Fern.) Mackenzie 1/O Carex tenera Dewey var. tenera R R $2 (NJ) Carex tetanica Schkuhr. R R PT, $2 (PA) Carex torta Boott ex Tuckerm. Oo ¥ Carex tribuloides Wahlenb. var. tribuloides VO Carex trisperma Dewey R Carex typhina Michx. R VO $3 (NJ), PE, S2 (PA) Carex utriculata Boott [C. rostrata var. utriculata V/O (Boott) Bailey] Carex vesicaria L. VO $2 (NJ) Carex virescens Muhl. ex Willd. Vv R Carex viridula Michx. var. viridula R S2 (NJ), PE, S1 (PA) Car CX Ip | id Michx R Cyperus bipartitus Torr. (C. rivularis Kunth.] F Cyperus compressus L. F Cyp dentatus Torr. R ¢8 dVO YALVA AAVAV TAC AHL AO VYOTA UV TNOSVA FHL AO LSITIOAHD ANV MAIAWAAO OSF R SUF LS LPM RIF RB SOB C _ TS _ state status Cyperus erythrorhizos Muhl. a 2 F Cyperus esculentus L. V Cyperus lupulinus (Spreng.) Marcks [C. filiculmis es = Vahl.] Cyperus strigosus L. V Dulichium arundinaceum (L.) Britt. var. arundinaceum R VO Eleocharis acicularis (L.) R. & S. EF Eleocharis compressa Sullivant var. compressa R NJE, PE, $1 (NJ, PA) Eleocharis elliptica Kunth 11f@) Eleocharis obtusa (Willd.) Schultes var. obtusa R $2 (NJ) Eleocharis ovata (Roth) R. & S. R Eleocharis palustris (L.) R. & S. R Py ris q ing fl ei \¢,.l [Scirp R pauciflorus Lightf.] Eleocharis tenuis (Willd.) Schultes var. tenuis R Eriophorum virginicum L. R R Eriophon um UI di ] (E £ | ) Fern R R $3 (NJ), PT, $2 (PA) Rhynchospora capitellata Tort. R Sch i I pung (V. } l) Palla [S. americanus F (Pers.) Volk. ex Schinz & P. Kellen, Scirpus americanus Pers., Scirpus p. Vahl.] Cal ] Pe DOE F ay] \e¢ jal [S inf Gray] R £ \ a? $2 (NJ), PE, S1 (PA) oo oa VINOLUVa OSF R SUF LS LPM RIF RB SOB Cc TS State status Dioscoreaceae Eriocaulaceae Hemerocallidaceae Hyacinthaceae Hydrocharitaceae Hypoxidaceae Schoenoplectus tabernaemontani K. C. Gemel [Scirpus validus Vahl.] Scirpus atrovirens Willd. Scirpus cyperinus (L.) Kunth. Scirpus expansus Fern. Scirpus microcarpus J. & K. Pers. Scirpus polyphyllus Vahl. Scleria pauciflora Muhl. ex Willd. var. pauciflora R Trichophorum a Palla [Scirpus R verecundus Fern Dioscorea villosa L. VO Eriocaulon aquaticum (Hill) Druce [E. septangulare Withering] * Hemerocallis fulva (L.) L. vO VO * Hemerocallis lilioasphodelus L. fe) *Muscari botryoides (L.) Miller R *Ornithogalum umbellatum L. F Elodea canadensis Michx. Elodea nuttallii (Planch.) St. John Najas flexilis (Willd.) Rostk. & Schmidt Vallisneria americana Michx. var. americana Hypoxis hirsuta (L.) Cov. F *Crocus vernus (L.) Hill O I R F NJE, $1 (NJ) $1? (NJ), PT, S2 (PA) 28 dV9 WALVA AYVAV TAC AHL AO VYO TA UV TNOSVA FHL AO LSITHOAHS ANV AAIAYAAO OSF R SUF LS LPM RIF RB SOB C_ TS _ state status *Iris germanica L. R * Iris pseudacorus L. VO *Iris sibirica L. R Tris versicolor L. F Sisyrinchium angustifolinm P. Mill. Vv F/O Sisyrinchium montanum Greene var. crebrum Fern. R R NJE, $2 (NJ) Sisyrinchium mucronatum Michx. Vv R Juncaceae Juncus acuminatus Michx. FiO: ¥ Juncus brachycephalus (Engelm.) Buch. R R $3 (NJ), PT, S2 (PA) Juncus bufonius L. Vv Juncus canadensis J. Gray ex La Harpe i Juncus dichotomus Ell. R R PE, $1 (PA) Juncus dudleyi Weig. (J. tenuis var. dudleyi (Wieg.) F. R $3 (NJ) J. Herm.] Juncus effusus L. var. solutus HO: #4. F/0 Juncus marginatus Rostk. F/O F/O Juncus nodosus Coville VO VO $3 (NJ) Juncus subcaudatus (Englem.) Corville & Blake F F Juncus tenuis Willd. var. tenuis FO OV Luzula multiflora (Ehrh.) Lej. VO VO F/O F Liliaceae Clintonia borealis (Ait.) Raf. Vv $3 (NJ) Vv Erythronium americanum Ker-Gawl. VINOLYVA “oF KR SUF LS ECPM RIP RB SOB C TS state status Melanthiaceae Orchidaceae Lilium canadense L. ssp. canadense * Lilium lancifolium Thunb. Lilium philadelphicum L. Medeola virginiana L. Streptopus amplexifolius (L.) DC. Streptopus roseus var. perspectus Fassett Chamaelirium luteum (L.) Gray Trillium cernuum L. var. cernunm Trillium erectum L. var. erectum Trillium undulatum Willd. Mi eine Deethn fan ¥ Alolcath cis bcheiduim t a Walt., Melanthium L Desr.} Veratrum vi 2(L.) Ait. f. [Melanthium v. L.) Veratrum viride Ait. Coeloglossum viride (L.) Hartman [Habenaria viridis (L.) R. Br. Ex Ait. F.] Corallorhiza maculata (Raf.) Raf. Corallorhiza odontorhiza (Willd.) Nutt. Corallorhiza trifida Chatelein Cypripedium acaule Ait. Cypripedium parviflorum var. pubescens Willd. [C. calceolus var. pubescens (Willd.) Correll, C. pubescens Willd. V/O 1/O F/O S2 (NJ) NJE, PT, S1 (NJ, PA) NJE, $1 (NJ) $3 (NJ) $2 (PA) $2 (NJ) NJE, S1 (NJ, PA) $2 (NJ), SH (PA) $2 (NJ) PE, $1 (PA) 68 dV9O WALVA FUVAV TAC AHL AO VYOTA UV TNOSVA AHL AO LSITIOAHO GNV AAIAYTAO State status 3 Cor K @e US Tee RF RS S08 C SS *Epipactis helleborine (L.) Crantz F R Galearis spectabilis (L.) Raf. [Orchis s. L.] Vo Goodyera pubescens (Willd.) R. Br. F Goodyera repens (L.) R. Br. ex Ait. f. R R $2 (PA) Isotria verticillata Raf. VO Liparis liliifolia (L.) L.C. Rich. ex Ker-Gawl. R NJE, SX.1 (NJ), Liparis loeselii (L.) L. C. Rich. R R sais Listera cordata (L.) R. Br. ex Ait. f. R R NJE, PE, $1 (NJ, PA) Malaxis unifolia Michx. V/O SH (NJ) Platanthera flava var. herbiola (R. Br. ex Ait. f.) Luer fe) $2 (NJ) PH aoah f. (L.) R. Br., Plantathera f. (L.) Lindl. var. Hostekcn lacera (Michx.) D, Don Vo fe) R Platanthera psycodes (L.) Lindl. R R R $2 (NJ) Spiranthes cernua (L.) L. C. Rich. F F/O Spiranthes lacera var. gracilis (Bigelow) Luer [S. R R gracilis (Bigelow) Beck] Spiranthes lucida (H. Eaton) Ames R R $2 (NJ), $3 (PA) Spiranthes ochroleuca (Rydb.) Rydb. VO $3 (NJ) Poaceae Agrostis hyemalis (Walt.) B. S. P. Vv F/O Agrostis perennans (Walt.) Tuckerm. F * Agrostis stolonifera L. var. palustris ar F/O VINOLYV4 OSF -R SUF. LS LPM RIF RB -SOB..C...TS .. statestatus Alopecurus aequalis Sobol. R R $2 (NJ), S3 (PA) si Andropogon gerardii Vitman vO VO < *Anthoxanthum odoratum L. F F/O < Aristida dichotoma Chapman var. dichotoma F Z * Arrbenatherum elatius (L.) J. & C. Presl. F F/O F/O . Avena sativa L. R VO 2 a CNN itcsce, ec FOR 2 Bromus altissimus Pursh [B. latiglumis (Shear) A. S. F/O & Hitche.] * Bromus inermis Leyss oo Sy < *Bromus japonicus Thunb, ex Murr. + ae Bromus kalmii A. Gray R $2 (NJ), $3 (PA) Bromus pubescens Muhl. ex Willd. F/O a2 * Bromus tectorum L. F 2 Calamagrostis canadensis (Michx.) Beauv. var. F/O FIO F SI canadensis Calamagrostis cinnoides (Torr.) Eat. (?) [C. coarctata : (Torr.) Eat.] tH Cinna arundinacea L. F/O F/O O z *Dactylis glomerata L. Vv oO bs Danthonia compressa Austin fe) Vo S Danthonia spicata (L.) Beauv. ex R. & S. F F/O s Deschampsia cespitosa (L.) Beauv. F F $3 (NJ, PA) 5 Deschampsia flexuosa (L.) Trin. vo = OSF R SUF LS LPM RIF RB SOB C__ TS _ statestatus Dichanthelium acuminatum (Sw.) Gould & C. A. Clark Dichanthelium boreale (Nash) Freckmann R NJE, S1 (NJ), SH (PA) Dichanthelium boscii Gould & C. A. Clark VO I O Dichanthelium clandestinum (L.) Gould F F/O Dichanthelium depauperatum (Muhl.) Gould. R Dichanthelium dichotomum (L.) Gould [Panicum d. L.] 1l/O Dichanthelium cas i al ex Nash)Gould [MO R_ I/O [Panicum I. Scribn. ex Nash] * Digitaria sanguinalis (L.) Scop. F V * Echinochloa crusgalli (L.) Gaertn. var. crusgalli om *Eleusine indica (L.) Gaertn. V Elymu densis L. var. canadensis F/O F/O Elymus hystrix L. [Hystrix patula Moench] * Elymus repens (L.) Desv. ex B. D. Jackson F F Elymus villosus Muhl. ex Willd. R R R Elymus virginicus L. F/O F/O F/O *Elytrigia repens (L.) Desv. ex B. D. Jackson F F [Agropyron r.] Eragrostis pelt ot (L.) Nees F F/O * Eragrostis cilianensis (All.) Janchen VO VO Eragrostis hypnoides (Lam.) BSP. R R Eragrostis spectabilis (Pursh) Steud. » dineae 74 8 | SO i) VINOLYVa CSF RK. SUF LS. LEM: RIP RB. SOB. G FS. ‘state status Festuca obtusa Biehler [F. subverticillata (Pers.) R exeev *Festuca ovina L. V F/O * Festuca ovina L. var. capillata ~ DD 3 Glyceria acutiflora Torr. F/O F/O Glyceria canadensis (Michx.) Trin. F/O F/O Glyceria melicaria (Michx.) C. E. Hubbard F/O F/O F/O F/O $3 (NJ) Glyceria striata (Lam.) A. S. Hitche. F/O. F/O- F/O *Holcus lanatus L. F F F/O Leersia oryzoides L. Sw. F r Leersia virginica Willd. HO. F *Lolium perenne L. F . Microstegium pg (Trin.) A. Camus [Eulalia V V viminea (Trin e] Muh Sci pits (Poir.) Fern. F/O F/O F/O Muhlenbergia sobolifera (Muhl.) Trin. Vo R Oryzopsis asperifolia Michx. R NJE, $1 (NJ) Panicum capillare L. F Panicum rigidulum Nees R R Panicum virgatum L. F F/O Paspalum setaceum var. setaceum VO VO Phalaris arundinacea L. x *Phleum pratense L. Vo F/O €6 dVO YALVA FTUVAV TAC AH.L AO VYOTA UV TNOSVA AHL AO .LSITIOAHO AUNV AAIAYAAO OSF RR SUP LS LPM RIF RB SOB C TS © state status “Phragmites australis (Cav.) Trin. ex Steud. ssp. F stralis *Poa annua L. VOSS *Poa compressa L. r F oO Poa languida A. Hitche. R $2 (NJ, PA) *Poa nemoralis L. R R Poa paludigena Fern. & Wieg. R PT, S3 (PA) Poa palustris L. VO V/O * Poa pratensis L. Ee F/O 2 Poa saltuensis Fern. & Wieg. R NJE, SH (NJ) Poa sylvestris A. Gray R SH (NJ) Schedonorus pratensis (Festuca p. Hudson, Lolium oo arundinaceum (Schreb.) S.J. Darbyshire] Schizachyrium scoparium (Michx.) Nash var. V F scoparium [Andropogon scoparius Michx.] *Secale cereale L. VO Rk *Setaria faberi Herrm. F *Setaria italica (L.) Beauv. F Setaria parviflora (Poir.) Kerguelen Setaria pumila (Poir.) Roemer & J.A. Schultes [S. F glauca (L.) Beauv.] Setaria viridis (L.) Beauv. V Sorghastrum nutans (L.) Nash V/O I I O Spartina pectinata Link +6 VINOLYVE KR SUP LS LaM Rie RB SOR Ut TS _ state status Sphenopholis obtusata var. major (Torr.) K.S. Erdman [S. intermedia (Rydb.) Rydb.] Sphenopholis obtusata (Michx.) Scribn. var. obtusata Sphenopholis pensylvanica (L.) A. §. Hitche. Sporobolus vaginiflorus (Torr.) Wood Tridens flavus (L.) Smyth [Triodia flava (L.) A. S. Hitchce.] * Triticum aestivum (L.) Vulpia octoflora (Walt.) Rydb. var. glauca [Festuca o. Walt.] *Zea mays L. Zizania aquatica L. Pontederiaceae Heteranthera dubia (Jacq.) Mac. M. [Zosterella d. acq.) Small] Pontederia cordata L. Potamogetonaceae Potamogeton alpinus Balbi *Potamogeton crispus L. Potamogeton diversifolius Raf. Potamogeton epibydrus Raf. Potamogeton foliosus Raf. Potamogeton illinoensis Morong Potamogeton natans L. Potamogeton nodosus Poir. Potamogeton perfoliatus L. F/O = /O I VO O F/O F/O SU (NJ) R PR, $3 (PA) R F R NJE, $1 (NJ) F ie) F F R NJE, $1 (NJ) R F R S6 dV5 WALVA AUVAV TAC AHL AO VUOTA UV INOSVA AHL AO LSITIOAHO GNV MALATTAO OSF R SUF tS ‘TPM RIF RB SOB (Accessed December, 2008). SMALL, J. K. 1933. Manual of the southeastern flora. Hafner Publishing Co., New York. 1554 pp. [Reprinted in two vols. in 2004 by The Blackburn Press, Caldwell, NJ.] SNYDER, D. B. 1996.The genus Rhexia in New Jersey. Bartonia 59: 55-70. SWALLEN, J. R. 1961. A new species of Panicum from New Jersey. Rhodora 63: 235-236. STONE, W. 1911. The plants of southern New Jersey with especial reference to the flora of the Pine Barrens and the geographic distribution of the species. Annual Report of the New Jersey State Museum for 1910 (pt. 2): 21-828. [Reprinted in 1973 by Quarterman Publications, Inc., Boston]. TAYLOR, P. G. 1989. The genus Utricularia: a taxonomic monograph. Kew Bulletin Additional Series XI WHeERRY, E. T. 1959. Goose Pond saved. Bartonia 29: 2. Bartonia No. 65: 115-116, 2011 William R. Overlease (1925-2007) When Bill passed away on November 30, 2007, we lost more than a botanist. We lost an ornithologist, an ecologist, a historian, a mentor, and a friend. Bill was born October 2, 1925, in Elkhart, Indiana, where he spent his early years. Then like so many of his generation he answered the call of his nation, spending four years from 1943 to 1946 with the US Navy in the Pacific. This tour of duty awoke in him a desire to travel, and in later years he managed to visit all seven continents. But Bill was always happy to come home, not only to Indiana, where he spent a number of years after the war working as a naturalist in the state park system, but to Benzie County, Michigan, where he spent many summers exploring the woods, swamps, and dunes of that part of the lower peninsula, collecting plants, studying birds, and conducting ecological studies. Bill was tireless in his quest to learn why plants and birds they grew where they grew and lived where they did. He was fascinated by plant distribution and succession and forest ecology. After graduating from Michigan State University in 1950 with a bachelor of science degree in forestry, followed up by a master of science in conservation (1952) and a doctorate in botany and plant pathology (1964) he moved to West Chester State College (later University), where he taught ecology and botany from 1963 to 1986. Once there, among other projects, he set up a plant succession study in an abandoned cornfield at the southwest corner of Tigue Road and New Street, which he monitored for 34 years (see pp. 97-104 of this issue). One of his greatest contributions while at West Chester, and one he was 115 116 BARTONIA very proud of, was helping to persuade the Board of Trustees in 1973 to formally establish the Robert B Gordon Natural Area at the South Campus as a permanent natural laboratory to study plants and animals in a suburban setting. Before and after its establishment as a preserve he spent much time there studying the structure and composition of the woody vegetation. With his teaching load, field studies, and other scholarly duties, it was often a wonder that Bill was able to get out in the field as much as he did. But when he did, he was an exhaustive and careful note taker. And when he got back, he spent innumerable hours writing them up, posting to his catalog of index cards and then hammering the data into shape for publication. Bill published innumerable articles on plants, birds, and other subjects, never letting weather, illness, or anything else interfere with his work. Botany was not his only love. Bill spent probably as many hours studying birds, both in Indiana, where he participated in many summer surveys both before and after his sojourn at West Chester, Michigan, and in Chester County, where he was a regular participant in the annual Audubon Christmas Count, as he did plants. He was an amateur historian, gathering, writing down, and publishing the tales and oral traditions of the frontier days of the Midwest. Not surprisingly, as befitted a man who spent years in Bruce Catton country, he was also a Civil War buff, collecting lyrics of marching songs, photographing old cemeteries, and gathering and publishing information on many of the unsung and forgotten contributors to the Northern war effort. It has been said that behind every great man is a woman. This was so true for Bill, for without his devoted wife, Edith, he would not have accomplished nearly as much he did. They first met at Turkey Run State Park in Indiana, in the 1950s, and were married in a log church there in 1955. Edith and Bill were inseparable, traveling, working, and writing together. Their last papers, published jointly, are in this issue (pp. 97-111). Bill was an incurable optimist, always able to see the silver lining in every dark cloud, to find the rare plant lurking amidst a mass of invasive aliens. As a result he was able to inspire generations of students to emulate him, not only in the field of botany, but in other pursuits. His memorial service and dedication ceremony on April 18, 2009 at West Chester was attended by a horde of former pupils ranging from college presidents to landscapers. This member of the greatest generation will be missed. JACK HOLT NOVEMBER 2010 Bartonia No. 65: 117-118, 2011 Ann Elizabeth Wolff Newbold (1922-2008) Ann Elizabeth Newbold, a fine, self-taught botanist who has made significant contributions to our knowledge of the flora of Pennsylvania and a past president of the Philadelphia Botanical Club (January 1977-May 1978), died of congestive heart failure on July 14, 2008 at Lima Estates, an assisted living facility in Media, Pennsylvania. After suffering several strokes, Ann was compelled to live life in a wheelchair during most of her last five years. For a woman accustomed to traveling, botanizing, playing tennis, skating, and skiing up to age 80, such confinement often proved difficult to accept. Throughout her life, Ann Newbold was a champion of racial equality and nuclear disarmament. She staunchly supported the Freedom Riders that headed to the Deep South. During the Vietnam War, she co-founded the Pottstown Peace Committee and served as a draft counselor, proud that she was able to persuade many young men to stay out of the war. In 1965, while standing in a silent Quaker peace vigil, Ann and her daughter suffered the indignity of being bombarded by eggs and tomatoes. Ann’s love of wildflowers was nurtured during childhood on the Wolff’s family apple orchard in Lima, Pennsylvania. At their home overlooking a pond in Bechtelsville, Montgomery County, she and her late husband Albert G. Newbold (died 1982) transformed a ten-acre parcel of meadow and woodland into a native wildflower garden, the Glenn Flora Preserve. This remarkable preserve was frequently a destination of field trips, especially to observe spring ephemerals or late-season asters and goldenrods. It also served as a “classroom” for students enrolled in Ann’s plant identification classes. Ann Newbold was a frequent participant on field trips sponsored by the Philadelphia Botanical Club and the leader or co-leader of 21 excursions conducted from 1976 through 1997, each of them documented in various issues of Bartonia. Her deliberate and intensive approach in the field stemmed from a desire to put a precise Latin name on every species enco is g dees to break out their plant manuals on the spot. Rarely did the group advance far beyond the parking lot in the first hour of botanizing. Ann once told the writer that “keying out” plants was one of her favorite activities. On extended joint field meetings of the Northeast Section of the Botanical Society of America, the Torrey Botanical Society, and the Philadelphia Botanical Club, she would often provide dissecting scopes and make arrangements for keying difficult species, especially members of the Cyperaceae. These valuable sessions often lasted late into the night. Gerry Moore fondly remembered: “My lasting recollection of Ann was her rainbow colored umbrella hat. She would wear it on days when rain was in the forecast, opening the umbrella when the rains came.” Teaching courses in botany and plant identification at the Barnes Foundation and the Morris Arboretum of the University of Pennsylvania provided Ann with great satisfaction. Equally rewarding was her work as a botanical consultant. During 1980-1982 she conducted a vegetation survey in an extensive area around a Pennsylvania nuclear power plant near Berwick. Another major assignment included her pl ities and vegetation diversity inventory of Valley Forge National Historical Park in the 1990s. Ann also provided major input for “A Natural Areas Inventory of Montgomery County, Pennsylvania,” a document Produced in the 1990s by the Pennsylvania Science Office of the Nature Conservancy for the Montgomery County Planning Commission. Throughout the 1980s, Ann Newbold served 117 118 BARTONIA as a steadfast member of a survey team (along with director Ann Rhoads, Richard Mellon, Roger Latham, and Jack Holt) of the Eastern Pennsylvania Rare Plant Survey, a project of the Morris Arboretum in cooperation with the Pennsylvania Natural Diversity Inventory. The persistent efforts of this team have resulted in the discovery of numerous county and state records. In the 1990s, Ann was engaged by the Delaware Heritage Program to serve on a similar survey team seeking rare plants especially in New Castle County, Delaware. A significant chapter in Ann Newbold’s life began in 1987 when she met Dr. Heinrich Zoller, professor emeritus at the University of Basel in Switzerland, and leader of a weeklong field excursion in the Swiss Alps that she attended during an international botanical congress. Their deep love of wildflowers and nature was an immediate bond that kindled an abiding relationship. A number of Philadelphia Botanical Club members will recall celebrating their “Affirmation of Love” reception at Bechtelsville on August 26, 1989. Ann’s daughter Monica recalls: “For the next 15 years, their story was a wonder. He would come to Pennsylvania or she would go to Switzerland for a month or two. Then the two of them would spend some time apart, pursuing their independent lives. Then again they would switch to visit the other person’s country. Together they went on botanical trips to Madagascar, Jamaica, the Swiss Alps, the U.S. West, Alaska, and more. When Mom moved into Lima Estates, the traveling stopped, but the love did not. She and Heinrich spoke to each other often by phone.” During the final two and a half to three years of Ann’s life, Heinrich’s own failing health made him too weak to endure an overseas flight to visit Ann. Ann Newbold was survived by Heinrich Zoller (since d d) andi ived by son J. Denis Newbold of Cochranville, Pennsylvania, son Edward Newbold of Seattle, daughter Monica Shay of New York, brother Kenneth Wolff of Lima, Pennsylvania, and three grandchildren. PUBLICATIONS BY ANN NEWBOLD IN BARTONIA No. 44: 32-36, 1976 — The Flora of Sunrise Mill Park, Montgomery County, Pennsylvania No. 45: 5, 1978 Additions to the Check-List of the Flora of Montgomery County No. 45: 15-17, 1978 Additions to the Flora of Sunrise Mill Park, Montgomery County, Pennsylvania No. 46: 49-50, 1979 Additions to the Check-List of the Flora of Montgomery County, II No. 47: 36, 1980 Additions to the Check-List of the Flora of Montgomery County, III (with Mariane McCabe) No. 51: 77, 1985 Montia chamissoi Rediscovered along the Delaware River Wayne County, Pennsylvania (with Ann F. Rhoads, Richard H. Mellon, and Roger E. Latham) No. 51: 115, 1985 Montgomery County Flower (Tulipa sylvestris) No. 52: 78, 1986 Additions to the Check-List of the Flora of Montgomery County, IV (with Jack Holt) No. 53: 41-43, 1987 Vascular Plants in Lord Petre’s Herbarium Collection by John Bartram (with Alfred E. Schuyler) No. 53: 47, 1987 Obituary: Leonard Sweetman No. 56: 65-67, 1990 Delaware Rare Plant Survey—Finding and Relocating Rare Plants of the Piedmont of Delaware (with Janet Ebert, Jack Holt, and Alfred E. Schuyler) TED GORDON JuLy 2010 Bartonia No. 65: 119-120, 2011 Naomi Dicker (1926-2009) I (Anderson) believe I first met Naomi on a joint Torrey Botanical Club-Philadelphia Botanical Club field trip to New Jersey Coastal Ponds, led by Dr. Alfred E. Schuyler on September 6, 1981. She had waist-long black hair, which she wore in a single braid. And she spoke softly, but with great precision. Her botanical expertise was evident, but she never was aggressive in her identifications. Only rarely would she correct somebody — just kind of hint that perhaps another identification was possible. Generally her possibilities turned out to be actualities. She was very ready to share her expertise. Her specialty was grasses, and she introduced many aspiring field botanists into the world of the glume and the lemma. Naomi had a wonderful command of language, enjoyed playing with words, and in addition had a great knowledge of word origins. She had studied linguistics; and was one of the few people around who could read (and understand) works such as Beowulf in their original Old English language. Not a terribly marketable skill, but one that made for some interesting conversation. As one might expect, she was also interested in art and history. She was enamored of New York City, and enjoyed showing out-of-towners the historic sites, architectural gems, and hidden public artworks of Manhattan. And she not infrequently housed visiting botanists in her apartment on west 93rd Street. 119 120 BARTONIA In the 1980s, the Torrey Botanical Club had a field trip committee, chaired by Richard Stalter. Naomi was one of several members, but was certainly the most active. In 1988, she promoted a meeting to discuss Matters of Interest to New York City Area Field Botanists. There were about 90 participants; the meeting was held at The New York Botanical Garden on 8 December 1988. Among other results, this led to the formation of a Metropolitan Flora Committee. In time, this provided the impetus for Brooklyn Botanic Garden’s Metropolitan Flora project. Naomi was an active Torrey Botanical Club trip leader. Between 1984 and 1997 she typically led one trip per season. Her trips usually took place in the vicinity of her beloved New York City within the five boroughs, or just across the Hudson in New Jersey. Especially noteworthy were the urban botany trips she led, often jointly with the New York City Audubon Society. Starting somewhere in midtown Manhattan’s west side, she would walk her group north for several miles, examining the flora of sidewalk cracks, vacant lots, and street tree plots. It was not unusual for passers-by to join in, at least for a while. She would routinely find 80 or more plant species on these trips. Shortly after I (Moore) began working at Brooklyn Botanic Garden in 2000, Naomi began volunteering with the New York Metropolitan Flora project. As expected, she focused primarily on grass identifications, often spending hours working on a single specimen and writing detailed notes to accompany her identification. She kept her own supply of fine dissecting needles that would allow her to dissect the smallest spikelets. I will always remember how delighted she would be when I would bring her material of an unusual grass species, such as Amphicarpum amphicarpon, Gymnopogon ambiguus, Piptochaetium avenaceum, Setaria magna, or Tripsacum dactyloides. When working late, Naomi noticed that I would frequently listen to baseball games, and this led to numerous discussions about the sport, such as keeping score, something that both of us did when attending games. Naomi was a New York Giants fan who loved going to games at the Polo Grounds. She would often discuss the Giants 1951 season when the Giants came from 13 games behind in August to eventually defeat the Brooklyn Dodgers in a three game playoff thanks to Bobby Thompson’s three run homerun (The Shot Heard ’Round the World) in the bottom of the ninth of the third game. On 14 August 2003, Naomi was at Brooklyn Botanic Garden when New York City (and other areas of the Northeast and Midwest) experienced a blackout. This prevented her from returning to her Manhattan apartment as the subways were not running, and she ended up staying at my Brooklyn apartment that night where we chatted about botany, baseball, and other subjects for much of the evening. Another subject that Naomi was devoted to was opera, and she frequently went to shows with other opera buffs, including Paul Harwood, the manager of Brooklyn Botanic Garden’s herbarium. Naomi was especially interested in Richard Wagner, and she had spent years working on a paper on Wagner's alliteration, stress and word choice in the libretti of the Ring Cycle (this ms remained unfinished at the time of her passing on July 14, 2009). She is acknowledged by John Louis DiGaetani in his book Wagner and Suicide published in 2003. One thing Naomi, Paul, and I had talked about but never did do was to go to Philadelphia and see a Phillies game and an opera on the same day. Naomi will be missed for her grass determinations and wonderful company. KARL ANDERSON GERRY MOORE SEPTEMBER 2009 Bartonia No. 65: 121-122, 2011 Franklin S. Hirst (1928-2009) On 3 August 2009, the botanical world lost one of its best friends when Frank Hirst, a field botany legend in the Middle Atlantic area, passed away from heart complications at the age of 81. As a young man, Frank frequented the Pine Barrens area of New Jersey. It was there that he forged a deep appreciation of all that was natural and right with the world. Frank and his brother Bob became interested in botany and scoured every nook and cranny of the Pine Barrens looking for rare and unusual plants. The list of species they found reads like a who’s who in New Jersey botany. In 1959 they found a species of grass that they had never seen before. This grass was later described as a new species, Panicum hirstii by Jason R. Swallen of the Smithsonian Institution (Rhodora 63: 235-236. 1961), the holotype for the name being a specimen collected by Frank Hirst in Atlantic County, New Jersey. Today there are only five known locations for this grass in the world, which is also referred to as Dichanthelium hirstii (Swallen) Kartesz by those who recognize the genus Dichanthelium. n 1971 Frank moved to the Eastern Shore of Maryland to get away from the ever encroaching development of coastal New Jersey. Being a plumber by trade, he decided to teach at the Worcester brig Vocational oo As a teacher, he took full advantage of his summers off by botanizing the he Delmarva Peninsula. In the early 1980s, he did td Welk for the then fledeling Maryland Natural Heritage Program. Many of the more promising areas he discovered were later purchased and turned into nature preserves by the Maryland/D.C. Chapter of the Nature Conservancy (TNC). He did similar work for the Delaware Natural Heritage Program as well and discovered some of their rarest plants. Besides his botanical contributions, Frank donated countless hours of volunteer work for TNC, especially at the Nassawango Creek Preserve. 121 122 BARTONIA Some of his more notable accomplishments were finding species that were new to the states with southern cutgrass (Leersia hexandra), a population of Harper’s fimbry (Finbristylis 1 (fm. ean ee | RF Ve aR perpusilla) in Maryland, sticky hedgehyssop (Gratio ifolia) in Delaware, Fernald’s ticktrefoil (Desmodium fernaldit) in Maryland, and Canby’s dropwort (Oxypolis canbyi) in Maryland. Th i i ti that went along ith the di y of Canby’s dr opwort. It was late in the afternoon of a day spent visiting many potential rare-plant habitats. Frank and his companions, George Fenwick and Dan Boone of the Maryland Natural Heritage Program, decided to visit one last area that appeared to be a coastal plain pond on the aerial photo. As they entered the large pond, Frank looked out across the mass of vegetation and yelled out excitedly, “There’s Oxypolis canbyi\” Considering that this plant had not been seen in 100 years on Delmarva and that Frank had never seen this species, this may have seemed like an outrageous statement, but not for Frank. He had done his h | d had pored the literati line d i gs, and phot f plants that were historically found on Delmarva. At this point George Fenwick replied, “Either you’re the greatest damn botanist that ever lived or you’re crazy.” I think the truth was closer to the first part of his statement, because later that night they verified that the plant was indeed Canby’s dropwort. Around 1989, I became obsessed with botany and took it upon myself to learn all of the plants I saw. The following year, Frank took me under his wing and greatly accelerated my climb up the learning curve. Later that year, we started what was to be 15 years of working together on botanical contracts with the Nature Conservancy of Maryland and Delaware _and the Natural Heritage Programs of Maryland and Delaware. The close friendship that developed between us was as precious as the great memories I now possess of those days. During our many years of botanizing together, I helped Frank carry on his legacy. I had figured that, by then, Frank had already found all of the rare plants there were to find, but together we added many new species to the state lists. Perhaps the greatest of our discoveries was in 1990, when we found the curly-grass fern (Schizaea pusilla) in Delaware. Before that day, the rather odd range of this fern was Newfoundland, New Brunswick, Nova Scotia, Long Island, New Jersey, and Peru. We were slogging through a wet Atlantic white cedar swamp, when we suddenly entered a clearing. Frank immediately commented, “This looks just like a cedar bog in New Jersey. I could picture almost anything in here!” About 30 seconds later, he yelled out, “I don’t believe it! Here’s curly-grass fern.” Since it was early in my botanical career, I didn’t fully realize the significance of what we had just found until I got home and looked it up. On many occasions, while botanizing with Frank, I would see a plant that seemed to be in a strange habitat and I would invariably ask him why is that plant growing there? Frank always had the same wry response, “Plants are where you find ’em.” I always knew that answer was coming, but I still asked. It was such a simplistic answer, but yet on another level it was profound. We really don’t understand all of the complex interrelationships that determine why certain plants occupy the niches where we find them. Frank had another stock answer to a question he was often asked, “What is your favorite wildflower?” His answer was always “It’s the one I have in my hand right now.” I think that summed up Frank’s philosophy very accurately. He appreciated everything that Nature had to offer and felt it was all worth saving. He will be missed. RONALD M.WILSON May 2010 Bartonia No. 65: 123-125, 2011 BOOK REVIEWS Uncorking the Past: The Quest for Wine, Beer, and Other Alcoholic Beverages, by Patrick E. McGovern. 2009. University of California Press, Berkeley. xv + 330 pages. $29.95. If you like botany, wine, and beer, this book is a must. Despite its omission from the title, there is a lot of botany in this book, and it’s good, drinkable botany. In the first chapter, McGovern introduces two hypotheses: The drunken monkey hypothesis proposes that alcoholism is rooted in the evolutionary history of primates, who had a powerful incentive to overindulge in fermented fruits and high-sugar resources that were available only in season. The Paleolithic hypothesis proposes that hunter-gatherers went beyond an unconscious craving for fermented fruit to a “more conscious, intentional production and consumption of a fermented beverage.” After presenting these hypotheses and commenting on the human benefits of alcohol, McGovern takes us on a world tour of fermented beverages. Some highlights follow. Starting with China, McGovern discusses his analysis of residue in ancient pottery of the world’s earliest known alcoholic beverage, made around 9,000 years ago. The ingredients were rice (Oryza sativa), grapes (possibly Vitis amurensis or V. quinquangularis), hawthorn fruits (Crataegus pinnatifida and/or C. cuneata), and honey. Neither beer nor wine, the beverage is described as a mixed fermented beverage, or grog. On to the Near East, we learn about the earliest known wine made with Eurasian grapes (V. vinifera), which also had resin of the terebinth tree (Pistacia atlantica). This information comes from McGovern’s analysis of residue in jars from Iran that were 7400-7000 years old. Also from Iran, a jug around 5,500 years old containing calcium oxalate (beerstone) provides chemical evidence of the earliest beer made from barley (Hordeum vulgare), the most widely consumed beverage in Mesopotamia. McGovern identifies Scotland as the site of the earliest known fermented beverage made from grain and honey in Europe. Vats dating from the mid-fourth millennium BCE contained pollen of cereal, meadowsweet (Filipendula ulmaria), and heather (Calluna vulgaris). McGovern suggests this may have been a “Nordic grog” with added herbs such as cloudberry (Rubus chamaemorus) and lingonberry (Vaccinium vitis-idaea)—two of my own favorite fruits from Newfoundland, where they are known as bakeapple and partridgeberry, respectively. We then get a fascinating survey of ancient fermented beverages in Europe and western Asia made with diverse ingredients including mugwort (Artemisia vulgaris), cranberries (Vaccinium oxycoccos), bog myrtle (Myrica gale), and saffron (Crocus sativus). Moving to South America, McGovern discusses the domestication of maize (Zea mays) in Mexico around 6,000 years ago. Humans’ first taste of it may have been wine made from sugary juice squeezed from its stalks. Chicha, beer made by fermenting the kernels, apparently came later. Similarly, the juicy, fermentable, sugary pulp in the fruit of the cacao, or chocolate, tree (Theobroma cacao) probably led to its domestication in Mesoamerica. There also is archaeological evidence of large amounts of wine made from drupes of the Peruvian pepper tree (Schinus moille) in southern Peru around 600 CE. In Africa south of the Sahara, the king of beers is made from fermented grains of Sorghum bicolor. Domesticated about 8,000 years ago, sorghum in the form of beer currently provides much of the caloric intake for hundreds of millions of Africans. In western sub-Saharan 123 124 BARTONIA Africa, beer is made from malted sorghum grains and clarified with bark of the raisin tree (Grewia flavescens) and okra (Abelmoschus esculentus). McGovern concludes that the available evidence suggests the initial domestication of grains was motivated by a desire to increase alcoholic-beverage production. Similarly, beverages made from the sweet fruits of grapes, figs, dates, and cacao probably prompted the domestication of these fruits. With his detailed account of the human consumption of fermented beverages over the past 9,000 years in different parts of the world, McGovern gives a strong argument for considering alcohol as a driving force for plant domestication, and thus civilization. ALFRED E. SCHUYLER Curator Emeritus of Botany Academy of Natural Sciences Benjamin Smith Barton: Naturalist and Physician in Jeffersonian America by Joseph Ewan and Nesta Dunn Ewan (edited by Victoria C. Hollowell, Eileen P. Duggan, and Marshall R. Crosby). 2007. Monographs in Systematic Botany 100. Missouri Botanical Garden Press, St. Louis. xxvi + 1134 pages. $55.00. The husband/wife team of Joseph and Nesta Ewan has long been familiar to Philadelphia naturalists and bibliophiles. They have spent many fruitful years in Philadelphia museums and libraries where they have plucked many “plums” that have enhanced our appreciation of colonial and early 19th century American natural history. Now we finally have their lengthy biography of Benjamin Smith Barton, Professor of Botany and Natural History in the Medical School of the University of Pennsylvania from 1790-1815. The book is not a light read. To me it is an archive crammed full of unpublished documents with a wealth of information to be distilled by curious readers. There are 95 pages of illustrations, which include some outstanding botanical drawings and paintings in the archives of the American Philosophical Society. Of particular appeal to me are nine paintings by the talented French artist Pierre Jean Francois Turpin. These are among “at least fifty ‘indigenous plants’ Turpin drew for Barton.” We can also appreciate the art of William Bartram and Frederick Pursh among these illustrations. Barton had diverse interests, but his legacy is botanical. The Academy houses what remains of much of his herbarium with plant collections made by him and other well- known botanists of the day. He authored the first American textbook of botany in 1803, which Catherine the Great ordered translated into French for her gratification. We continue to appreciate the accomplishments of his protégés: Frederick Pursh, Meriwether Lewis, and Thomas Nuttall. Without Barton’s support, Frederick Pursh would not have produced his classic Flora Americae Septentrionalis where he described and illustrated plants collected by Meriwether Lewis. Without his “crash course” in botany from Barton, we assume that Lewis would have been ill prepared for collecting and preserving botanical specimens. Today these specimens in the herbarium of the Academy of Natural Sciences are th ignifi | history artifacts of the Lewis and Clark Expedition. BOOK REVIEWS 125 And finally, Thomas Nuttall, who received his initial funding from Barton, went on to a distinguished career as America’s foremost botanical traveler and collector. His work remains significant today because, like Frederick Pursh, he was an author who named and described many new American plant species. There is much more in the Ewans’ book. People interested in the history of medicine at the University of Pennsylvania will be rewarded as well as those with an interest in late 18th and early 19th century intellectual activities in Philadelphia. It fosters appreciation of the rich intellectual heritage of Philadelphia when it was the “Athens” of America. ALFRED E, SCHUYLER Curator Emeritus of Botany Academy of Natural Sciences Bartonia No. 65: 126-147, 2011 2007-2008 FIELD TRIPS Reports reviewed, formatted, and edited by TED GORDON. 2007 Field Trips 18 March-13 October: Various Plant Communities in the Pinelands National Reserve, New Jersey. The Pinelands Preservation Alliance (PPA) sponsored fourteen all-day Saturday lecture- workshop sessions in association with a Special Pinelands Plant Course at their PPA headquarters in Vincentown. Focusing on rare plants with legal protection in the Pinelands, each session was followed by a field trip, many of these with special guest speakers and field trip leaders. Some of the highlights included the following: 31 March—leader Russell Juelg, dwarf forest ecology and Corema conradit. 28 April—leader Russell Juelg, New Jersey Conservation Foundation’s Four Mile Spring Preserve, characteristic Pine Barrens uplands, Pitch Pine lowlands, and a mixed-community wetland complex with Listera australis. 19 May—leader Renee Scagnelli, Tuckahoe area, a diverse wetland community, and a severely disturbed site with Ophioglossum vulgatum. 7 July—leader Ted Gordon, an intermittent pond (“Hampton-Central Big Pond”) near Hampton Furnace, Scleria minor, Calamovilfa brevipilis, Carex barrattit, Muhlenbergia torreyana, Nymphoides cordata, and Panicum hirstu. 4 August—leader Ernie Schuyler, Scleria nitida in a diverse, semi-dry, disturbed area near the Lebanon Fire Tower, Schoenoplectus novae-angliae in tidal freshwater where Route 542 crosses the Merrygold Branch, and Rhynchospora knieskernii in an intermittent pond in the Stafford Forge Wildlife Management Area. 18 August—leader Bill Olson, a recently burned forest near Atsion. In unburned, diverse, wetland areas adjacent to the burn, we found Lobelia canbyi, Rhynchospora cephalantha, and Panicum scabriusculum. 8 September—leader Wayne Ferren, Eleocharis olivacea var. reductiseta in tidal wetlands of the Wading River, and Eryngium aquaticum and Eriocaulon parkeri in tidal wetlands of the Mullica River. In addition to the guest lecturers and trip leaders mentioned above, we also had trip/instruction sessions with Gerry Moore, Ken Cameron, Joe Arsenault, and Karl Anderson. Registrants: 20. Report by Russell Juelg. 15 April: Biodiversity Blitz at the John A. Phillips Preserve, Old Bridge, Middlesex County, New Jersey. Trip cancelled because of stormy weather. On 14 April, however, Bill Olson visited the site and observed the following species: b dg: grass,A d. pog gi: icus; hai bittercress, Cardamine hi =P yl i sedge, Carex pensylvanica; shaved sedge, Carex tonsa; ground-pine, Dendrolycopodium obscurum; buttonweed, Diodia teres; spring whitlow-grass, Draba verna; threeway sedge, Dulichium arundinaceum; weeping lovegrass, Eragrostis curvula; love grass, Eragrostis spectabilis; common rush, Juncus effusus; scirpus-like rush, Juncus scirpoides; dewberry, Rubus hispidus; glaucous greenbrier, Smilax glauca; round-leaved greenbrier, Smilax rotundifolia; atrichum moss, Atrichum angustatum; little broom moss, Dicrane 126 2007-2008 FIELD TRIPS 127 heteromalla; hypnum moss, Hypnum imponens; pin cushion moss, Leucobryum glaucum; platygyrium moss, Platygyrium repens; common haircap moss, Polytrichum commune; juniper haircap moss, Polytrichum juniperinum; Ohio polytrichum moss, Polytrichum ohioense; a peat moss, Sphagnum sp.; lingzhi, Ganoderma lucidum; candleflame lichen, Candelaria concolor; mealy pixie-cup lichen, Cladonia chloropheae; common powderhorn lichen, Cladonia coniocraea ; British soldier lichen, Cladonia cristatella; common g ield lichen, Flavoparmelia caperata; bottlebrush shield lichen, Parmelia squarrosa; parmotrema lichen, Parmotrema hypotropum; wreath lichen, Phaeophyscia pusilloides; placynthiella lichen, Placynthiella uliginosa; rough-speckled shield lichen, Punctelia subrudecta; and oak apple gall wasp, Amphibolips confluenta. Richard Lear (richard.lear@co.middlesex.nj.us) of Middlesex County, Department of Parks and Recreation is maintaining a comprehensive species list for this site. Report by coordinators: Richard Lear and Lena Struwe. 28 April: Duke Island Park, Bridgewater, Somerset County, New Jersey. Joint trip with the Torrey Botanical Society. Our plan for the day was to search the floodplain of the Raritan River for spring ephemerals, but torrential rains had caused the river to flood the area on the day previous. Overnight the water had receded, leaving a coating of mud on much of the low vegetation. Despite that, a few hardy souls braved the mud and debris in the east section of the park, finding a fine display of robust Mertensia virginica, which was in full bloom and only partially mud-coated. Several of the plants had all pink or all white flowers, rather than blue. With diligent searching, we were able to find other species that were identifiable and in good shape. In flower we found Caulophyllum thalictroides, Erythronium americanum, Floerkea proserpinacoides, Viola pubescens, V. sororia, Cardamine pensylvanica, and Claytonia virginica. Hydrophyllum virginianum was inbud. Beds of Allium tricoccum were frequent, but we found leaves only. The invasive Ranunculus ficaria, also in bloom, covered large patches of ground. Of course, the huge old-growth trees were awesome: Platanus occidentalis, Acer saccharinum, Nyssa sylvatica, Fraxinus americana, Carya cordiformis, and Liriodendron tulipifera. Tiring of the mud, we decided after lunch to travel to nearby Sourland Mountain, where the vegetation was refreshingly green. Some species we found in flower there were Arisaema triphyllum, Chrysosplenium americanum, Claytonia virginica, Ranunculus abortivus, Thalictrum thalictroides, Viola pubescens, Hepatica americana, Luzula multiflora, and Obolaria virginica. Abundant Orchis (Galearis) spectabilis was in bud, as were Podophyllum peltatum, Polygonatum pubescens, and Geranium maculatum. A nice selection of ferns was also seen, including Adiantum pedatum, Athyrium thelypterioides (syn. Deparia acrostichoides), A. filix-femina, Botrychium virginianum, Dryopteris marginalis, Osmunda claytoniana, Polypodium virginianum, Polystichum acrostichotdes, Phegopteris hexagonoptera, and several others. Thanks to Janet Novak for keeping a plant list for the day. Attendance: 7. Report by leader: Linda Kelly. 12 May: Crosswicks Creek, Cream Ridge, Monmouth County, New Jersey. Leader: Linda Rohleder. No report received. 10 June: Biodiversity Blitz at John A. Phillips Preserve, Old Bridge, Middlesex County, New Jersey. Joint trip arranged by Chrysler Herbarium, Rutgers University 128 BARTONIA (Department of Ecology, Evolution and Natural Resources) and Middlesex County, Department of Parks and Recreation. During this day, 92 species were cataloged from data sheets that were sent in to Middlesex County Parks and Recreation Office. Species that were seen included eastern kingbird, Tyrannus tyrannus; cedar waxwing, Bombycilla cedrorum; pine warbler, Dendroica pinus; ovenbird, Seirus aurocapilla; scarlet tanager, Piranga olivacea; red-tailed hawk, Buteo jamaicensis; Carolina chickadee, Poecile carolinensis; wood thrush, Hylocichla mustelina; indigo bunting, Passerina cyanea; common yellowthroat, Geothlypis trichas; black-and- white warbler, Mniotilta varia; red-eyed vireo, Vireo olivaceus; wild turkey, Meleagris gallopavo; red-bellied gnatcatcher, Polioptila caerulea; veery, Turdus migratorius; yellow- billed cuckoo, Coccyzus americanus; great crested flycatcher, Myiarchus crinitus; red admiral, Vanessa atalanta; wolf’s milk slime, Lycogala epidendrum; Indian pipe, Monotropa uniflora; skunk cabbage, Symplocarpus foetidus; lone star tick, Amblyomma americanum; orchard spider, Leucauge venusta; partridgeberry, Mitchella repens; polyphemus, Antheraea polyphemus; royal fern, Osmunda regalis; Carolina locust, Dissosteira carolina; turkey tail, Trametes versicolor; greater bladder sedge, Carex intumescens; bracker fungi, Postia caesia; alsike clover, Trifolium hybridum; black medic, Medicago lupulina; white clover, Trifolium repens; hairy vetch, Vicia villosa; wild peppergrass, Lepidium densif] 3p ial rye Lolium perenne; willow herb, Epilobium ciliatum; lowbush blueberry, Vaccinium pallidum; sheep laurel, Kalmia angustifolia; staggerbush, Lyonia mariana; yellow wood sorrel, Oxalis stricta; dangleberry, Gaylussacia frondosa; rabbit-foot clover, Trifolium arvense; red fox, Vulpes vulpes; comma butterfly, Polygonia c-album; mourning cloak butterfly, Nymphalis antiopa; southern ringneck snake, Diadophis punctatus punctatus; American kestrel, Falco sparverius; spicebush swallowtail, Papilio troilus; wood satyr, Megisto cymela; netted chain fern, Woodardia areolata; Virginia chain fern, Woodardia virginica; wolf’s milk slime, Lycogala epidendrum; spring field cricket, Grylus veletis; scarlet tanager, Piranga olivacea; pink lady’s slipper, Cypripedium acaule; box turtle, Terrapene carolina; spring azure, Celestrina ladon. (See also trip report for 15 April.) Attendance: 16. Report by coordinators: Richard Lear and Lena Struwe. Ws LAID, 17-21 June: Elkins, West Virginia. Joint Annual Field Meeting of the Torrey Botanical Society, the Philadelphia Botanical Club, and the Northeastern Section of the Botanical Society of America. The meeting took place at Davis and Elkins College in Elkins, West Virginia, a gateway to the high Allegheny Mountains. The first day of field trips visited sites on Cheat Mountain in the area of Gaudineer Knob, including upland limestone forest, balsam fir swamp, young and old-growth red spruce forest, and riverscour prairie. The second day was at Dolly Sods, including several types of mountain laurel shrubland (with Kalmia latifolia in peak bloom), cranberry-beakrush peatland (with Vaccinium oxycoccos in peak bloom), and the picturesque red spruce-heath rocky woodland. The third day was spent at Canaan Valley National Wildlife Refuge and State Park to observe limestone meadow, beaver pond, oxbow fen, and several types of balsam fir swamp. The field trip leaders were Jim Vanderhorst, Elizabeth Byers, and Brian Streets from West Virginia Natural Heritage Program; Dr. Katherine Gregg of West Virginia Wesleyan College; and Leah Ceperley from Canaan Valley National Wildlife Refuge. Evening lectures were given by Elizabeth Byers, Katherine Gregg, Bill Roody (West Virginia DNR), and Rodney Bartgis (The Nature Conservancy). 2007-2008 FIELD TRIPS 129 Attendance: 47, 42 full-time and 5 part-time participants representing 11 northeastern states plus the District of Columbia and Florida. Thanks to Karl Anderson for serving as the treasurer. Report by Conference Chair: Larry Klotz. Assistant Chair: Marcia Minichiello. June 30: Tucquan Preserve, Erbs Mill, Lancaster County, Pennsylvania. Joint trip with the Delaware Nature Society. The group met at the entrance to the Tucquan Preserve in west-central Lancaster County north of the Holtwood dam. From the bucolic parking lot they went down the wooded trail along the south side of Tucquan Creek. The first portion of the walk proceeded through relatively level, rich oak-tulip forest, where Carex torta grew on gravelly bars. Interesting woody species encountered here included Asimina triloba, Cercis canadensis, Chionanthus virginicus, Hydrangea arborescens, and local colonies of the orchid Goodyera pubescens. Further west, as the slopes came down to the water’s edge, the terrain grew increasingly rocky and difficult to traverse, as oaks, hemlocks, and great laurel dominated the landscape. Blueberries, partridge-berry, and striped pipsissewa were frequently encountered here. Less steep, richer ravines and slopes yielded Corylus cornuta, Adiantum pedatum, Mitella diphylla, Trillium erectum £. album, Ostrya virginiana, and Caulophyllum thalictroides. Occasional areas of glade-like, level ground along the stream, including where we ate lunch, were lush with a mixture of weedy native and non-native herbs, ferns, and shrubs. Notable plants growing in this habitat included Lilium canadense, Salvia lyrata, Monarda clinopodia, Veratrum viride, and a particularly large, fruiting specimen of Fraxinus nigra. Further downstream enormous outcrops of schistose, metamorphic rocks started jutting out from the hillsides. Here moister, shaded cracks were frequently lined with Asplenium montanum and occasional sprays of Aquilegia canadensis. Close to the Susquehanna the gorge opened up, with sycamore and silver maple taking over the canopy, shading flood- tolerant herbs including Hydrophyllum virginianum. Notable were large boulders in the stream bed now half-covered with sand, mud, and silt accumulated since the damming of the river. Dominating the arid, steep banks and cinders along a railroad were a mixture of drought- tolerant grasses and herbs, mainly Andropogon virginicus and Schizachyrium scoparium, Eupatorium rotundifolium v. rotundifolium, Hypericum gentianoides, and Linaria canadensis. The moister west bank of the railroad was home to a wide variety of disturbance- loving species, including Desmodium spp., while Lythrum salicaria was the most noticeable herb on the bars and mudflats lining the Susquehanna. The wooded north slope of Tucquan Creek, although weedy, had its share of less common plants, including Tradescantia virginiana, Silene stellata, Solidago ulmifolia, and Bromus pubescens, growing on the rich, loamy soil. Further east atop the ridge grew a deer-browsed oak-heath woodland, which gave way to a rich oak-tulip woodland. The most notable species seen here was a thin population of Carex jamesii, growing along an overgrown woodland lane. Downslope near the stream the botanists clambered through yet more rocky woodland, which yielded a fine population of Conopholis americana in fruit. Passiflora lutea was observed by a few stragglers. The following is a list of species compiled by Jack Holt during the field trip: Acalypha rhomboidea, Acer negundo, A. rubrum, A. saccharinum, A. saccharum, Adiantum pedatum, Aegopodium podagraria, Ageratina altissima (Eupatorium rugosum), Agrimonia pubescens, Agrostis perennans, Ailanthus altissima, Ajuga reptans, Alliaria petiolata, Allium 130 BARTONIA canadense, Amaranthus spinosus, Ambrosia artemisiifolia, A. trifida, Amelanchier arborea, Amphicarpaea bracteata, Andropogon virginicus, Anemone quinquefolia, Anthoxanthum odoratum, Apios americana, panos cannabinum, Aquilegia canadensis, Arabis laevigata, Are ifolia, Arisaema triphyllum, Artemisia vulgaris, Asarum canadense, Asimina ike Asplenium montanum, Athyrium filix-femina, Berberis thunbergii, Betula lenta, B. nigra, Bidens frondosa, Bidens sp., Boehmeria cylindrica, Botrychium dissectum, B. virginianum, Brachyelytrum erectum, Bromus pubescens, B. tectorum, Callitriche sp., Calystegia sepium, Cardamine hirsuta, C. pensylvanica, Carex amphibola, C. blanda, C. communis, C. debilis, C. digitalis, C. gracillima, C. jamesi, C. laxiculmis, C. laxiflora, C. pensylvanica, C. radiata, C. sparganioides, C. swanit, C. torta, C. tribuloides, C. virescens, Carpinus caroliniana, Carya alba, C. cordiformis, C. glabra, Castanea dentata, Caulophyllum thalictroides, Celastrus orbiculatus, Celtis occidentalis, Cercis canadensis, Chamaecrista nictitans, Chamaesyce maculata, C. nutans, Chelone glabra, Chenopodium album, Chimaphila maculata, Chionanthus virginicus, Chrysosplenium americanum, Cimicifuga racemosa, Cinna arundinacea, Circaea lutetiana, Coincya monensis, Collinsonia canadensis, Commelina communis, Conopholis americana, Conyza canadensis, Cornus alternifolia, Corylus cornuta, Cryptotaenia canadensis, Danthonia compressa, D. spicata, Dennstaedtia punctilobula, Deparia acrostichoides, Desmodium glutinosum, D. nudiflorum, D. paniculatum, D. perplexum, Dichanthelium boscii, D. clandestinum, D. dichotomum, D. latifolium, Dioscorea quaternata, D. villosa, Diphasiastrum digitatum, Dryopteris intermedia, D. marginalis, Duchesnea indica, Elymus riparius, Eragrostis hypnoides, Erigeron annuus, E. philadelphicus, Eupatorium perfoliatum, E. rotundifolium v. rotundifolium, E. fistulosum (Eutrochium f.), E. purpureum, Eurybia (Aster) divaricata, Fagus grandifolia, Fallopia japonicum (Polygonum cuspidatum), F. scandens, Festuca elatior, F. obtusa, Fraxinus americana, Fraxinus nigra, Galinsoga ciliata, Galium aparine, G. circaezans, G. triflorum, Geranium carolinianum, G. maculatum, Geum canadense, G. verum, Glechoma hederacea, Glyceria striata, Goodyera pubescens, Hamamelis virginiana, Hedera helix, Helianthus decapetalus, Hemerocallis fulva, Hepatica americana, Hieractum venosum, Hosta ventricosa, Huperzia lucidula, Hydrangea arborescens, Hydrocotyle americana, Hydrophyllum virgimianum, Hypericum gentianoides, H. mutilum, Ilex verticillata, Impatiens pallida, Juglans nigra, Juncus tenuis, Juniperus virginiana, Kalmia latifolia, Lactuca biennis, Lechea sp., Leersia virginica, Lepidium virginianum, Ligustrum obtusifolium, Lilium canadense, Linaria canadensis, Lindera benzoin, Lindernia dubia, Liriodendron tulipifera, Lobelia inflata, Lonicera japonica, L. morrowil, Luzula echinata, Lycopodium obscurum, Lycopus virginicus, Lysimachia ciliata, L. quadrifolia, Lythrum salicaria, Maianthemum canadense, M. (Smilacina) racemosum, Malus sp., Matteuccia struthiopteris, Medeola virginiana, Melilotus officinalis, Microstegium vimineum, Mitchella repens, Mitella diphylla, Monarda clinopodia, Monotropa uniflora, Muhlenbergia schreberi, Myosoton aquaticum, Nyssa sylvatica, Oecnothera — Onoclea sensibilis, Osmorhiza claytoni, O. longistylis, Osmunda cinnamomea, O. claytoniana, Ostrya virginiana, Oxalis dillenit, Parthenocissus quinquefolia, Passiflora lutea, ee tomentosa, Penthorum sedoides, Perilla frutescens, Persicaria (Polygonum) arifolia, longiseta, P. perfoliata, P. virginiana, Phalaris arundinacea, Physalis subglabrata, eaters americana, Pilea pumila, Plantago rugelit, Platanus occidentalis, Poa compressa, P. sylvestris, P. trivialis, Podophyllum peltatum, Polygonatum biflorum, P. pubescens, Polygonum aviculare, Polypodium virginianum, Polystichum miottiheidies Potentilla canadensis, P. norvegica, P. simplex, Prenanthes altissima, Prunella vulgaris, Prunus avium, P. serotina, 2007-2008 FIELD TRIPS 131 Quercus coccinea, Q. montana (Q. prinus), Q. rubra, Q. velutina, Ranunculus abortivus, Ranunculus sceleratus, Rhododendron maximum, R. periclymenoides, Ribes sp., Robinia pseudoacacia, Rosa multiflora, Rubus allegheniensis, R. flagellaris, R. phoenicolasius, Rudbeckia laciniata, Rumex crispus, R. obtusifolius, Salix nigra, Salvia lyrata, Sambucus canadensis, Sanguinaria canadensis, Sanicula canadensis, S. trifoliata, Saponaria officinalis, Sassafras albidum, Schizachyrium scoparium, Scirpus georgianus, Scrophularia marilandica, Scutellaria elliptica, Senecio aureus, S. vulgaris, Sicyos angulatus, Silene antirrhina, S. stellata, Smilax glauca, S. herbacea, S. pulverulenta, S. rotundifolia, Solanum nigrum, Solidago bicolor, S. caesia, S. canadensis, S. flexicaulis, S. gigantea, S. ulmifolia, Sphenopholis intermedia, Stellaria longifolia, a (Aster) cordifolium, S. lanceolatum, S. lateriflorum, S. puniceum, S pus foetidus, Teucrium canadense, Thalictrum pubescens, Thalictrum sioketesiaies Thelypteris noveboracensis, Tilia americana, Toxicodendron radicans, Tradescantia virginiana, Trillium erectum v. album, Triodanis perfoliata, Tsuga canadensis, Ulmus rubra, Urtica dioica, Uvularia perfoliata, U. sessilifolia, Vaccinium corymbosum, V. pallidum, V. stamineum, Veratrum viride, Verbascum thapsus, Verbena urticifolia, Verbesina alternifolia, Veronica officinalis, Viburnum acerifolium, V. dentatum, Vinca minor, Viola arvensis, V. blanda, V. conspersa, V. eriocarpon, V. ON api V. sororia, V. striata, Vitis aestivalis, V. labrusca, V. vulpina, Xanthium strumarium Attendance: 10. Leader: William Ryan. Report by Jack Holt. 21 July: Peaslee Wildlife Management Area, Cumberland County, New Jersey. Joint meeting with the Torrey Botanical Society and the Gloucester County Nature Club. The morning of this trip was spent near Cumberland Pond. Drosera intermedia was in bloom along the shore, Pontederia cordata was in bloom in the shallows, and the flowers of Nymphaea odorata decorated the open waters. Also present were Peltandra virginica, Lysimachia terrestris, Decodon verticillatus, Triadenum virginicum, Chamaedaphne calyculata, Rhynchospora alba, and Xyris smalliana. An adjacent field, which at first glance seemed an l exploration, yielded Asclepias tuberosa, Hypericum stragulum, H. gentianoides, Solidago odora, Jasione montana, Opuntia humifusa, Diodia teres, Eupatorium pilosum, E. album, E. hyssopifolium, and E. rotundifolium, to name just a few species, all in bloom. A very good find was the rare Desmodium strictum, also in bloom. Asclepias amplexicaulis was found with fruit. A damp depression in this field had Rhexia virginica, R. mariana, Apios americana, and Ludwigia alternifolia in bloom. A brief foray to a forested wetland edge just downstream from the lake found one flowering plant of Platanthera lacera and numerous plants of P. clavellata. Clumps of iris- like leaves near the site of the Cumberland Furnace, showing no sign that they had ever flowered, were identified by Janet Novak as Belamcanda chinensis. Trees and shrubs, here and near the lake, included Quercus stellata, Q. alba, Q. falcata, Q. prinoides, Leucothoe racemosa, Vaccinium corymbosum, and Rhododendron viscosum. After lunch, a visit was made to a railroad crossing just north of Port Elizabeth. Here, the showy Ipomopsis rubra, a western species, has been established for over a century—it is mentioned in Witmer Stone’s Plants of Southern New Jersey (1911) but has never spread from the site. It was found in bloom. Also seen at this location were numerous non- flowering plants of Desmodium strictum, along with Monarda punctata in good bloom. Carya pallida was found here - an unfamiliar tree species to some trip participants. The next stops of the day were a utility line cut north of Halberton, and an intermittent pond nearby. The cut had Polygala nuttallii, Lobelia nuttalli, Rhynchospora capitellata, 132 BARTONIA Rhexia mariana, Carex complanata, and a large population of the uncommon and inconspicuous Croton willdenowii. Oldenlandia uniflora was found growing in the middle of a sand road. The pond, a circular opening about 100 meters across, rimmed with Vaccinium corymbosum and Smilax rotundifolia, was water-filled to an average depth of about 50 centimeters. Several large clones of Panicum hemitomon were seen here with their seldom-seen flowers and fruit. In many areas, the pond was covered with Nymphaea odorata, but there were patches of Brasenia schreberi, some with flowers. The dominant vegetation was Schoenoplectus subterminalis (Scirpus s.). Eleocharis microcarpa, Xyris smalliana, Myriophyllum sp., and Utricularia sp. were the only other visible plants. After the “official” trip, several participants followed leader Linda Kelly to a tidal marsh south of Mays Landing, where hundreds of Lilium superbum were found in bloom. Also here were Gratiola aurea, Scutellaria integrifolia, Sium suave, and Mikania scandens. Many thanks to Janet Novak for compiling a plant list for the day and for helping with some difficult identifications. Attendance: 11. Report by leaders: Karl Anderson and Linda Kelly. 29 July: East Plains - Stafford Forge Wildlife Management Area (SFWMA), Ocean County & Warren Grove Gunnery Range (WGR), Burlington County, New Jersey. Botanizing commenced on the SFWMA ca. 0.25 mile east of the gated WGR entrance. In Watering Place Pond were several flowering specimens of Utricularia purpurea, purple bladderwort; Sphagnum cuspidatum and S. trinitense, two submersed peat mosses; and a single specimen of Chrysemys scripta elegans, red-eared turtle (introduced). Here we failed to re-locate an occurrence of Sphagnum macrophyllum, but near the pond outlet were a couple of plants of Muhlenbergia torreyana, Torrey’s muhly, and about a dozen fruiting culms of Rhynchospora knieskernii, Knieskern’s beaked rush, both elements discovered a number of years ago. In the adjacent plains or dwarf pine-scrub oak community ona slope east of the pond, we observed eight scattered, well-established, 2- to 3-year-old seedlings of Corema conradii, broom crowberry, that survived the severe fire of May 2007. Occurring on a 1-acre patch, these seedlings were spared because they were surrounded by a substantial buffer of nearly bare mineral soil. In contrast, a nearby patch of several mature cushions of broom crowberry growing among a contiguous fuel layer of dense pine sprouts and shrubs was totally expunged by an earlier fire. Clearly, wide sandy buffers (actually created by fire) devoid of a fuel-load enhance the survival chances of populations of this state endangered shrub. Along an abandoned airstrip within the WGR Target Zone, we visited highly productive rare species habitats severely impacted by such anthropogenic disturbances as bulldozing, scraping, detonation of munitions, mowing, and prescribed burning (the latter two still ongoing). These disturbances have helped to maintain early successional conditions that benefit the pioneer species that thrive here. An estimated 6000 fruiting tufts of Knieskern’s beaked rush were observed in a periodically prescribed-burned seepage strip 100 m long x 3 m wide that parallels the central segment of the runway on the east side. An even larger population of this federally-threatened sedge occurred to the southwest of the runway near its southern end in a regenerating area marred by parallel “sight lines” or bulldozed strips that once d igati aids. There are at least eight other significant occurrences of Knieskern’s beaked rush within the borders of the WGR. Together these occurrences rival those that thrive in the shallow, abandoned, clay pits and powerline right-of-way at Reega 2007-2008 FIELD TRIPS 133 in Atlantic County. For a list of associated species of the above two WGR sites, see Bartonia No. 63: 8 September, 57-58, 2006. Three of co-leader Bien’s Drexel University students showed us their Gentiana autumnalis study plots equipped with monitoring wells. Here the impacts of groundwater fluctuation and prescribed burning were being monitored and evaluated. Situated on the west side of the northern half of the runway, this parcel has fairly consistently produced thousands of flowering gentians annually. Associated species here were Scleria minor, slender nutrush; Rhynchospora knieskernii; Calamovilfa brevipilis, pine barren reedgrass; and Panicum wrightianum, Wright's panic grass. All of these species have a “wetland affinity” or are deemed to be moisture dependent. Thus it was perplexing to see them growing side-by-side with such upland species as Quercus ilicifolia, bear oak; Q. marilandica, blackjack oak; Hudsonia ericoides, pine barren heather; Scleria pauciflora var. caroliniana, Carolina nutrush; and Eurybia compacta, slender aster. Soil borings of this habitat, exhibiting pit and mound relief, have verified the presence of a semi-permeable clay fragipan that allows rain water to perch near the surface for extended periods. The leaders have designated this parcel a “complex hydroxeric habitat.”(See Bartonia No. 64: 9-10, 2009 for the location and further discussion of this habitat.) Plans to devote the remainder of the afternoon to conducting de novo surveys of lesser- known areas along the Oswego River, the northern border of WGR, came to an abrupt halt when a severe cloudburst produced torrential rain. We were fortunate to prevent our vehicles from getting stuck in deep puddles and ruts made slick by underlying clay. Attendance: 18. Leaders: Ted Gordon and Walt Bien. Report by T. Gordon. 4 August: Intermittent Hampton-Central Ponds and Vicinity, Wharton State Forest, Burlington County, New Jersey. Entering the Wharton State Forest at Hampton Gate via Carranza Road, we were pleased to learn that a small, though well established, occurrence of Pityopsis falcata was still flourishing along the east road shoulder just beyond the intersection of Moores Meadow Road. The rare sickle-leaved golden aster, associated with Plantago aristata and Diodia teres, was growing in a 12 meter-long strip of sparsely vegetated sand. From the site of nearby Hampton Furnace, we traveled east on High Crossing Road, a sand trail, to reach our primary destination, the “Hampton-Central Ponds.” En route, we stopped briefly at the edge of a former bog-ore excavation to record the following typical plants growing in wet sand: Panicum verrucosum, Juncus canadensis, J. pelocarpus, J. effusus var. pylaei, Carex barrattii, Cyperus dentatus, Eleocharis tuberculosa, Rhynchospora capitellata, Triadenum virginicum, Hypericum canadense,* Lobelia nuttallii,* Polygala lutea,* Rhexia virginica,* Xyris difformis var. difformis,* and Xyris torta.* To reach the ponds, we had to traverse on foot a parcel of pitch pine lowland that was devastated by an extremely hot fire on July 18, 1999. Our passage was complicated by uneven terrain marred by depressions, haphazardly scattered dead tree trunks, thickets of regenerating shrubs and pine sprouts, and humid weather with a temperature approaching 95°. A thorough survey of this diverse, recovering landscape yielded the following 39 species: trees & shrubs - seedlings and saplings of both Acer rubrum forma trilobum and Ilex opaca, tall sprouts of Pinus rigida, Chamaedaphne calyculata, Clethra alnifolia,* Gaultheria procumbens, Gaylussacia baccata, G. dumosa, G. frondosa, Hudsonia ericoides, Ilex glabra, Kalmia angustifolia, Leucothoe racemosa, Lyonia mariana, Morella pensylvanica, Pyxidanthera barbulata, Vaccinium corymbosum vat. corymbosum, V. corymbosum var. caesariense, V. pallidum; herbs—Hypericum denticulatum var. 134 BARTONIA denticulatum,* Lechea racemulosa, Lobelia nuttallit,* Minuartia caroliniana, Platanthera cristata* (4 in a moist depression), Sabatia difformis,* Xerophyll. phodeloides; gra Andropogon glomeratus, A. virginicus, Calamovilfa brevipilis, Panicum mattamuskeetense, P. virgatum, Schizachyrium scoparium var. scoparium; sedges - Rhynchospora capitellata, Scleria triglomerata; ferns - Pteridium aquilinum, var. latiusculum, Woodwardia virginica; mosses - Sphagnum tenerum, S. magellanicum, and S. fallax (?). Two sedges, Rhynchospora torreyana and Scleria minor, the herb Bartonia virginica, and the fern Schizaea pusilla were seen here in July, 2004, but not during this exploration. We next entered “Hampton-Central Big Pond,” where on July 17, 2004, T. Gordon and R. Juelg di d f the globally imperiled grass, Panicum hirstii (= Dichanthelium hirsti). It was our intention to determine the extent and vigor of this population. However, since only about a dozen of these extremely narrow, wand-like culms still retained their fruit, it proved too difficult to see the delicate, bare inflorescences jutting out from the shallow water. A general survey of the pond resulted in finding the following 46 species, excluding woody plants: herbs—Brasenia schreberi,* Decodon verticillatus, Drosera intermedia,* Eriocaulon aquaticum, E. compressum* (rosettes only), E. decangulare,* Hypericum denticulatum var. denticulatum,* Lobelia canbyi,* L. nuttallii,* Lophiola aurea, Nymphaea odorata,* Nymphoides cordata,* Polygala cruciata,* Proserpinaca pectinata,* Rhexia virginica,* Sabatia difformis,* Triadenum virginicum, Utricularia striata,* Xyris smalliana;** rushes—Juncus biflorus,* J. canadensis,’ J. pelocarpus;* sedges—Carex barrattit, C. bullata, C. livida, C. striata, Cladium mariscoides,*” Dulichium arundinaceum, Eleocharis olivacea, E. robbinsi,\ E. tuberculosa, Rhynchospora alba,*" R. capitellata, R. cephalantha, R. gracilenta, R. fusca, Scleria minor; grasses— Andropogon glomeratus, Calamovilfa brevipilis, Mublenbergia torreyana, Panicum hirstii, P. spretum, P. virgatum, Saccharum giganteum; ferns—Woodwardia virginica; mosses— Sphagnum cuspidatum. Species seen here in 2004 but not during this survey were Cyperus dentatus, Sphagnum cyclophyllum, S. macrophyllum, and Platanthera blephariglottis. However, the current survey added 9 new species to the 2004 survey list. (See also Bartonia No. 63: 17 July, 62-63, 2006.) To reach adjacent “South Pond,” much smaller than Big Pond and totally devoid of water, we followed a short, slightly elevated causeway. A casual survey resulted in finding the following species: herbs—Eriocaulon aquaticum, E. compressum, Hypericum denticulatum var. denticulatum,* Lachnanthes caroliniana, Lophiola aurea, Nymphaea odorata, Proserpinaca pectinata,’ Sabatia difformis,* Utricularia striata,* Xyris smalliana;* rushes—Juncus canadensis, J. pelocarpus;\ sedges—Cladium mariscoides,*\ Eleocharis olivacea,* E. robbinsi,® E. tuberculosa, Rhynchospora alba,’ R. cephalantha,® R. fusca; grasses—Calamovilfa brevipilis, Mublenbergia torreyana, Panicum spretum, Saccharum giganteum; ferns—Woodwardia virginica. Species seen here in 2004 (pond flooded) but not in 2007 were Scleria minor, Rhynchospora pallida, and Utricularia juncea. The smallest pond of the cluster, “Southwest Pond,” also devoid of water, yielded the following species: herbs—Lobelia canbyi,* Proserpinaca pectinata,\ Xyris smalliana;*" rushes—Juncus pelocarpus;\ sedges—Cladium mariscoides,*\ Eleocharis robbinsii, Rhynchospora alba,** R. cephalantha;\ grasses - Muhlenbergia torreyana;\ ferns— Woodwardia virginica.’ Seen in 2004, Eriocaulon compressum’ and Lachnanthes caroliniana were either overlooked or not present on this occasion. The flora of individual intermittent ponds in close proximity can vary dramatically. The three ponds surveyed had in common only those ten species marked with a caret (). It is anticipated that this number will increase with additional, timely field work. VIS — 2007-2008 FIELD TRIPS 135 We returned to Carranza Road to see a small occurrence of Stylisma pickeringii var. pickeringi, Pickering’ s morning glory, in bloom near the Carranza Memorial. A few trailing vines were growing in extremely dry sand along the railroad tracks in association with Pinus echinata, P. rigida, Cyperus grayi, Diodia teres, Hudsonia ericoides, and H. tomentosa. Further to the south, we searched unsuccessfully for an old Gcrarrente of Stylisma at the “lost” village of Sandy Ridge. This fire-impacted site contained Jonactis (Aster) linariifolius, Solidago odora, M. elampyrum lineare var. pectinatum, Diodia teres, pe Liatris graminifolia. Our final stop, a moist road shoulder and shrub thicket along Route 563 below the cranberry village of Hog Wallow, harbored a spectacular floral display of 372 specimens of Platanthera blephariglottis, white fringed orchid. [* flowering plant; * indicates plant present in all three “Hampton-Central Ponds.” Thanks go to Bill Standaert for maintaining a list of species observed. Attendance: 8. Report by leader: Ted Gordon. 25 August: Daleville, Chester County, Pennsylvania. In the Elk Creek watershed, we visited the extensive parcel of land owned by Anna Francis, a participant in the Conservation Reserve Enhancement Program (CREP), which rewards landowners for instating conservation practices on their land. Serving as our trip leader, the CREP facilitator gave a brief talk regarding the property, which has been extensively planted with trees. The tract, at the upper end of the headwaters of Elk Creek, was a dairy farm prior to current ownership; horses now graze the upland portions. Over 900 trees were planted along the riparian corridor during the initial planting in 1997, with an additional 2200 trees in 2003 along with 300 shrubs. The most common species planted included Fraxinus pennsylvanica, Platanus occidentalis, and Betula nigra, with some Liquidambar styracaflua, Quercus palustris, Q. bicolor, Liriodendron tulipifera, Acer saccharinum, and one or two Fagus grandifolia. Shrubs included Alnus serrulata, Cephalanthus occidentalis, Cornus amomum, Ilex verticillata, Sambucus canadensis, and Viburnum dentatum. The leader further discussed the evolution of the planting program, including how some local non-natives such as Liguidambar and Betula nigra have been dropped from her area planting list; the survival and growth rates of various species; the problems associated with mislabeled species, notably Alnus glutinosa; and methods of protecting the saplings from both deer and invasive species, especially Phalaris arundinacea, Agrostis gigantea, and Microstegium vimineum at this site; and protecting animals that use the tree tubes for shelter and food. Of special interest to the participants were the reason why netting is fitted atop the tubes (to reduce bluebird mortality), and the — in the north of the state with black bears destroying trees to get at wasp nests inside the tubes. We then went into the field both upstream and downstream, venturing into the extensive open and shrubby wetlands along the edges of the watercourse, where in the course of sloshing through the often thick herbage we encountered some of the unusual species known on the property. These included Calamagrostis canadensis, Sagittaria australis, Rhexia virginica in bloom, a nice flowering population of Oxypolis rigidior, and a sedge resembling Carex tetanica and, to some degree, C. meadii. This last Carex has been examined by Tony Reznicek and other sedge specialists. It apparently has not yet been determined whether it is an anomalous form of an existing Carex or a new taxon. We finished the trip by examining an adjoining Brandywine Conservancy property, a low woodland dominated by oaks and red maple, and walking along the edge of a large, rectangular, abandoned farm pond. Attendance: 6. Leader: Jessie Benjamin. Report by Jack Holt. 136 BARTONIA 8 September: Welsh Mountain, Chester County, Pennsylvania. On a brief walk along the edge of Struble Lake near the fishermen’s parking lot, we saw A galinis tenuifolia in bloom both in the tall grass at water’s edge and in the rough-mown lawn. We th yed by car to the wooded upp h slope of Welsh Mountain near the junction of Berks, Chester, and Lancaster Counties. This tract was recently put under a conservation easement. We slowly walked up a gas line cut that ran straight up the mountain, observing numerous acid-tolerant edge and meadow plants. Frequent cutting had favored numerous low-growing and mat-forming species. These included four clubmosses, Dendrolycopodium obscurum, D. hickeyi, Lycopodium clavatum, and Diphasiastrum digitatum, Epigaea repens, Lespedeza repens, Viola primulifolia, V. sagittata, V. lanceolata, and Spiranthes cernua (in bud). The Lycopodium, Epigaea, and Viola lanceolata frequently formed large colonies, the first-named frequently found draped over the high edges of the cut. Other notable species included Eupatorium pilosum low down, and a single sprig of Spiraea tomentosa, atop the cut. Large colonies of Kalmuia latifolia, including cut-over mats, lined the north edge of the cut. We then ventured into the woods on either side of the cut. The slopes of Welsh Mountain possess an increasingly rare habitat in southeastern Pennsylvania, a mature oak-heath upland forest that has not yet been heavily impacted by deer. Chestnut oak was the most common oak, mixed with red, black, and white, with black gum, cherry birch, and a few red maples mixed in. Members of the Ericaceae dominated the shrub and ground flora, with Kalmia latifolia practically the only tall shrub, frequently forming near-impenetrable thickets. Lower down were the huckleberries and blueberries, consisting mainly of Gaylussacia baccata, G. frondosa, and Vaccinium pallidum, along with an occasional V. stamineum and V. corymbosum. Except for Medeola virginiana, few herbs grew with the heaths, but notable species included a local colony of Trientalis borealis, a nice colony of Goodyera pubescens, and the occasional Cypripedium acaule. After lunch some of the botanists walked along the lower edge of the woods above an expanding housing development, where we looked at a population of Jsotria verticillata and a patch of Viburnum nudum var. cassinoides. Of concern to the leaders was the high mortality observed among the Kalmia. The previous year, most of the mountain laurel south of the utility cut was dead or dying, while most to the north was healthy and thriving. This year the southern population was re-sprouting from the bases while the northern thickets were dying off. Attendance: 11. Report by leaders: Janet Ebert and Jack Holt. 15 September: Sandy Hook, Gateway National Recreation Area, Monmouth County, New Jersey. Joint trip with Torrey Botanical Society. The beach was our first stop, to avoid potential crowds later in the day. Our primary objectives were two rare species, Amaranthus pumilus and Polygonum glaucum. We found both. We also saw large populations of Artemisia campestris around the parking lot and along the path to the beach and Helianthus petiolaris among the dunes along the road. Our next stop was an J/ex opaca forest. Sandy Hook offers one of the best examples of this habitat in the region. Many of the trees are over one hundred years old, impressively large, and quite beautiful. The forest is on the bay side. We encountered salt marsh along the boardwalk on the way in and upper-marsh habitat near the entrance to the forest. Species we saw of note were Bassia hyssopifolia, Limonium carolinianum, Maianthemum stellatum, Salicornia europaea, Salicornia virginica, and Salsola kali, We then proceeded to a large parking lot in the center of Sandy Hook where a “mystery plant” had been reported before the trip. Since Sandy Hook is home to a number of species 2007-2008 FIELD TRIPS 137 common in other regions, but rare in the Northeast, we had reason to be hopeful. Once again Sand Hook’s reputation did not disappoint. The plant was Eriogonum annuum, a member of the Polygonaceae common on the Great Plains but very rare in the East. After a stop for lunch, we set out to look for another relatively-recent arrival from a distant region, Carex macrocephala. This species is native to the Pacific Northwest, Alaska, and northeastern Asia, but is only found at three sites in the East. It somewhat resembles Carex kobomugi in its growth habit. Not far from the parking lot we were surprised by a small population of Celastrus scandens. Then, along the way we passed scattered populations of Panicum amarum. Far into the dunes we found a large swale filled with Carex macrocephala among more P. amarum. Our last stop was Plum Island, just north of the park entrance. Here we observed Atriplex cristata (A. arenaria), Other plants observed during the course of the trip were Baptisia tinctoria, Cakile edentula, Lathyrus japonicus, Lechea maritima, Plantago psyllium, Berberis vulgaris, and Hudsonia tomentosa. Thanks to Linda Kelly for keeping a species list. Attendance: 16. Report by leader: David Austin 16 September: Silver Lake/Delhaas Woods and 5-Mile Woods, Bucks County, Pennsylvania On this trip we examined the geology and how it influenced the plant communities. At Silver Lake Nature Center east of Bath Road and Delhaas Woods, west of Bath Road near Bristol, we looked at relatively undisturbed habitats underlain by pre-Wisconsin glacial deposits. These habitats exhibited very acidic conditions (pH ~ 3.8—4.5) that harbored acidophiles like Andropogon glomeratus, Carex cf. bullata, Clethra alnifolia, Lilium superbum, Liquidambar styraciflua, Lyonia ligustrina, Maianthemum canadense, Quercus phellos, Q. falcata, apparent Q. phellos X falcata, Rhexia mariana, Rhododendron cf. periclymenoides, Rhynchospora capitellata, Spiraea latifolia, Symphyotrichum (Aster) novi-belgii, and Viola lanceolata. A small, previously unknown colony of Corallorhiza odontorhiza was discovered in a disturbed area. In the marshes of Mill Creek, east of Bath Road in the Park, the soils were much richer, having washed down from the agriculturally enriched Pennsauken and Bridgeton formations and Wissahickon schist, which tends to be richer. Species observed here included Cephalanthus occidentalis, Chelone glabra, Clematis virginiana, Convallaria majalis, Cornus amomum, Eupatorium dubium, Hibiscus palustris, Lythrum salicaria, Mikania scandens, Nuphar sp., Leersia oryzoides, Sambucus canadensis, Saururus cernuus, and many other common species. At Five-Mile Woods Preserve in Lower Makefield Township, Bucks County, we looked at the influence of undisturbed Pennsauken, Bridgeton, and Chickies Quartzite formations, all of which tend to be very acidic. These conditions provided habitat for less-common to rare species in Bucks County like Bartonia paniculata, Botrychium dissectum, Chimaphila maculata, Clethra alnifolia, Lyonia ligustrina, Medeola virginiana, Monotropa uniflora, Tipularia discolor, Vaccinium corymbosum, and Viola primulifolia. In areas of former agricultural use and around old foundations on the same three geologic formations, species associated with richer soil were found. These included Actaea pachypoda, Botrychium virginianum, Celtis occidentalis, Lindera benzoin, Lonicera japonica, Microstegium vimineum, and Polystichum acrostichoides. Thanks go to Janet Novak for maintaining a species list. Attendance: 6. Report by leader: Rick Mellon. 138 BARTONIA 2008 Field Trips 15 March: In Search of Mistletoe, Winslow Township, Camden County and Monroe Township, Gloucester County, New Jersey. Joint trip with the Flora of New Jersey Project. The trip started in Folsom, Atlantic County, where a brief history of the region and the biology of Phoradendron leucarpum were discussed. Our goal was to see American mistletoe at multiple stations along the Great Egg Harbor River. Winter is the best time to find American mistletoe. Without the deciduous canopy foliage, colonies of this unusual parasite are readily visible. The parasite’s affinity in New Jersey with Nyssa sylvatica, black gum, was discussed. From Folsom, we drove a short distance to the Piney Hollow Road entrance into Winslow Fish and Wildlife Manag Area. This state-owned tract supports many Pinelands habitats as well as a substantial population of American mistletoe. We entered the forest following the management area’s dirt roads. Our first stop was Inskip’s, the site of a historic inn and the famed Blue Hole. We searched the river banks for a population observed by the trip leader a few weeks prior. After a fruitless search it was discovered the tree in which a fine colony had existed was toppled during a winter storm. The large black gum was found sprawled across the river and its colony of mistletoe lost in the fast-moving current of the Great Egg Harbor River. The group laughed at the coincidence and continued to survey for additional colonies. Approximately 100 meters downstream, a small population was found at the top of a gnarled black gum. This was the only occurrence we observed near the historic site. We left Inskip’s and followed the old Blue Anchor Fireline north toward Winslow Road. We stopped approximately ‘4 mile before the paved county highway at the edge of a large, circular, wet depression. Here we discussed the regional geology and landscape formation while searching the tree line for mistletoe. We discovered one small colony of mistletoe in a young black gum sapling on the edge of the dirt road. We drove north from this depression, crossed Winslow Road, and followed the south side of the Atlantic City Expressway to a manmade excavation using a dirt service road. The group searched the tree line surrounding the pond, identifying squirrel’s nests and witch’s brooms but no additional mistletoe populations. The pond edge was inspected and the following species were identified: Rhynchospora macrostachya, R. alba, R. capitellata, Lycopodiella alopecuroides, Triadenum virginicum, Andropogon glomeratus, and Juncus canadensis. The fourth stop was on the Winslow Road bridge spanning the Great Egg Harbor River. From this vantage point, we were able to identify many colonies of American mistletoe. Dozens of large and small colonies occupied the lowest part of the river’s floodplain, which apparently is providing ideal conditions to sustain this large occurrence. Each colony occupied the upper branches of the floodplain’s large-stature black gums. After a brief lunch, the group drove north on Morgan Road to the intersection with Malaga Road in the historic village of New Brooklyn. We parked in a Winslow Fish and Wildlife Management Area parking lot and immediately spied dozens of large and small mistletoe colonies. The group was able to walk among the black gum trees supporting mistletoe and found parts of the plants on the forest floor. Fruiting specimens were examined and the anatomy of the parasite discussed. At this station one silver maple tree in an adjacent residential landscape was observed with a small colony of American mistletoe. This may be the only existing New Jersey documentation of mistletoe in Acer saccharinum, silver maple. Report by leader: Joe Arsenault. 2007-2008 FIELD TRIPS 139 5 April: Field Botany Party, Pinelands Preservation Alliance Headquarters, Vincentown, New Jersey. Short trip to Smithville, Burlington County. No report received. 3 May: Bowman’s Hill Wildflower Preserve (BHWP) “Phyto-Bio Blitz,” New Hope, Pennsylvania. Joint trip with BHWP staff and volunteers. BHWP features over 700 species of Pennsylvania plants, both native and originally non- native, growing along trails in a natural setting of woodland, meadow, and pond. The total acreage of the preserve is 134 acres, of which approximately 100 acres are surrounded bya deer exclusion fence, erected in 1994. The phytobioblitz was just the beginning of an ongoing comprehensive survey with multiple goals. It is important to know what is growing on the preserve. It is also desirable to verify presence and current condition of some of the plants that were planted in previous years, going back into the 1950s. Finally, by using the Plant Stewardship Index it is possible to compare the “nativeness” of vegetation in selected areas inside and outside the deer fence. For a full explanation of the technique and rationale visit the BHWP website: www.bhwp.org/db. For the phytoblitz, the preserve was divided into sixteen areas of about ten acres each. Groups of volunteers spent the morning and afternoon sampling the vegetation. Some of the species found included Caltha palustris, Dicentra eximia, Mertensia virginica, Caulophyllum thalictroides, Polemonium repens, Trillium grandiflorum, T. cernuum, T. luteum and Viola labradorica. For a complete lists of species found, see the preserve’s website. A delicious lunch in the Moore Pavilion and a catered dinner were served by BHWP. Attendance; 16. Report by Coordinator: Jeannine Vannais. 2-4 May: Field Workshop - Spring Wildflowers of the Delaware National Recreation Area, Pocono Environmental Education Center. Instructor: Bill Olson. Event was cancelled. 10 May: Flora of Ker-Feal (country estate of Albert Barnes), West Pikeland Township, Chester County, Pennsylvania. Joint trip with students of the Barnes Foundation. We first examined cultivated plants in the yard near the house and then walked though fallow fields and early successional forest to a relatively late successional forest at the west end of the property. We saw most of the cultivated trees and shrubs around the house, outbuildings, and along the driveway that were reported on a previous trip in 2002 (Bartonia 62: 126). We noticed that two plants not reported on the previous trip, Primula Japonica and Photinia villosa, were spreading to other parts of the property. Large stands of the invasive Elaeagnus umbellata were abundant in and along the margins of fallow fields. An early successional forest along the horse trail west of the driveway had numerous non native and invasive plants including Acer platanoides, Phellodendron amurense, Euonymus alatus, Ligustrum obtusifolium, Rhodotypos scandens, and Celastrus orbiculatus. The forest at the west end of the property was dominated by Liriodendron tulipifera, but in steeper portions there were numerous trees of Quercus rubra, Quercus velutina, and Fagus grandifolia. Lindera benzoin and Viburnum acerifolium were dominant understory shrubs. Spring wildflowers included Arisaema triphyllum, Claytonia virginica, Galearis spectabilis, Maianthemum racemosum, Obolaria virginica, Podophyllum peltatum, Sanguinaria canadensis, Uvularia perfoliata, Viola cucullata, and Viola pubescens. Attendance: ca 15. Report by leader: Alfred E. Schuyler. 140 BARTONIA 1-5 June: The Pinelands National Reserve, Sections of Ocean and Burlington Counties, New Jersey. Joint Field Meeting of the Northeastern Section of the Botanical Society of America, the Philadelphia Botanical Club, and the Torrey Botanical Society. Participants were housed at the Lighthouse Center for Natural Resource Education on Barnegat Bay in Waretown, Ocean County. Five experts on the regional flora served as trip leaders: Joseph Arsenault of Arsenault Environmental Consulting; Dr. Walter Bien of Drexel University; Ted Gordon, Director of Pine Barrens Inventories; Dr. Gerry Moore, Director of Science, Brooklyn Botanic Garden; and William Olson of Maser Consulting. Travel to a diverse tapestry of wetland and upland plant communities was in 4 vans and 4 private vehicles. In preparation for Monday’s excursions, Walter Bien’s Sunday evening PowerPoint program, “Fire Effects on the Pitch Pine Plains,” provided a comprehensive overview of the globally imperiled dwarf pitch pine (Pinus rigida) communities created and maintained by frequent intense fires. On Monday, following in the footsteps of botanists of the 19th and 20th centuries (J.H. Redfield, Constantine Rafi , John Torrey, Witmer Stone) in search of Corema conradii (broom berry) at it limit of range, we paused briefly at historic Cedar Bridge Tavern that had welcomed these travelers who then went on to seek the crowberry on the West Plains. Our next stop at a nearby plantation of towering white pines set out as seedlings in 1960 by forester Silas Little convinced us that soil impoverishment was not the cause of stunting in the plains. At the Le Clare homestead on the Little Plains near Warren Grove we again saw planted, non-native trees that were protected from wildfire towering over the endemic stunted pines and scrub oaks in adjacent fire ravaged areas. Also observed were many members of the heath family, including large areas of Letophyllum buxifolium (sand myrtle), some still in bloom. Extensive carpets of broom crowberry and patches of Pyxidanthera barbulata (pyxie) were also seen. After lunch we viewed the Lower Plains from an observation tower at the Warren Grove Air National Guard Gunnery Range. Here we visited anthropogenically disturbed sites harboring many rare pioneer species recently documented by Bien and Gordon. Of special note were Rhynchospora knieskernii, (Knieskern’s beaked rush), a large population of Gentiana autumnalis (pine barren gentian), and Muhlenbergia torreyana (Torrey’s muhly). Bien also discussed regeneration plots on severely disturbed sites maintained by his graduate students. The evening slide-illustrated program, “Cedar Swamps, Savannahs, and Quaking Bogs of the New Jersey Pine Barrens,” was presented by Ted Gordon. It provided an in-depth overview of a number of wetland communities visited the following day. On Tuesday morning near Wells Mills County Park we walked through an upland pitch pine (Pinus rigida)-blackjack oak (Quercus marilandica) forest with some Q. ilicifolia (bear oak), Q. stellata (post oak), and Sassafras albidum (sassafras). No additional tree species can tolerate the severe fires that shaped this community. The understory was dominated by Gaylussacia frondosa (dangleberry), G. baccata (black huckleberry), and Vaccinium pallidum (lowbush blueberry). We also visited an adjacent fire-scorched pitch pine lowland dominated by Calamovilfa brevipilis (pine barren reedgrass) and Gaylussacia dumosa (dwarf huckleberry). Both of these communities produced large populations of Xerophyllum asphodeloides (turkeybeard) whose flowers were severely browsed by deer. Next we entered an adjoining cedar swamp along Cold Brook to see Schizaea pusilla (curly grass fern) and, in anthesis, Arethusa bulbosa (dragon mouth). At the Oswego Cranberry/Blueberry Research Center near Jenkins, Dr. Amy Howell presented a PowerPoint program titled “The Health Benefits of Cranberries and Blueberries.” In a canal at the center we noted the rare Utricularia inflata (swollen bladderwort), the rare U. purpurea (purple bladderwort), and the common U. striata 2007-2008 FIELD TRIPS 141 (striped bladderwort). After lunch we focused on the riverine savannahs, quaking bogs, and cedar swamps along the Oswego River above Martha Furnace. Highlights here were colorful sphagnum carpets, an abundance of flowering Sarracenia purpurea (pitcher plant), Utricularia cornuta (horned bladderwort), Narthecium americanum (bog asphodel), and Eriocaulon compressum (early pipewort). At the furnace site Liparis loeselii (Loesel’s twayblade) and a small glade harboring a dozen fern species, including Ophioglossum pusillum (northern adder’s tongue), attracted attention. The evening PowerPoint program by Gerry Moore was titled “Rare and Endangered Species of the New Jersey Pine Barrens.” This presentation provided the participants an Opportunity to appreciate in flower the many special plants that thrive throughout the growing season. On Wednesday at Webbs Mills, the group traversed a boardwalk over a bog created and favorably impacted by turf removal. Prolonged flooding by beaver had delayed flowering of many species. Carex exilis (coastal sedge) and a few dragon mouth were in bloom; bog asphodel and Lophiola aurea (gold crest) were in bud. The four leaders made a strong effort to teach the group to identify bog plants in their non-flowering state. At Whitesbog, birthplace of the cultivated blueberry, we saw several banners of Calopogon tuberosus (grasspink), Polygala lutea (orange milkwort), Minuartia caroliniana (pine barren sandwort), Cephalanthus occidentalis (buttonbush), Nymphaea odorata (fragrant white water lily; inc. forma rosea), Itea virginica (Virginia willow), and two highlights, Uvularia puberula var. nitida (pine barren bellwort), and New Jersey’s only native occurrence of Schwalbea americana (chaffseed). After botanizing the edge of Pakim Pond in Brendan Byrne (Lebanon) State Forest, we examined an advanced growth Chamaecyparis thyoides (Atlantic white cedar) swamp along Shinn Branch. Here we noted J/ex laevigata (smooth winter holly), Trientalis borealis (star flower), Smilax walteri (coral greenbrier), Thelypteris simulata (bog fern), and Woodwardia areolata (netted chainfern). In the evening a fine banquet was held at the nearby Captain’s Inn in Forked River. Potential sites for next year’s meeting were discussed, leading to the possibility of the Pocono Mountains. Mark Demitroff, University of Delaware Permafrost Group, presented a PowerPoint program titled “Pine Barrens Wetlands: Geographic Reflections of South Jersey’s Glacial Past.” Mark made a convincing case that permafrost played an important role in shaping regional wetlands in southern New Jersey. Thanks go to the treasurer Karl Anderson; the assistant chair Walter Bien; German Georgieff, Chief Naturalist of Wells Mills County Park, his staff and associates for providing transportation; Dr. Nicholi Vorsa, Director of the Marucci Blueberry and Cranberry Research Center, for use of their facilities; and Patricia Gordon for assistance with all phases of this event. Attendance: 58. Report by conference chairperson: Ted Gordon. 22 June: Haycock Mountain, Bucks County, Pennsylvania. We climbed partway up the east side of Haycock Mountain on a trail that took us through rocky woods and past boulder fields. Actaea racemosa (Cimicifuga racemosa; black cohosh) was abundant. In the rockier areas, we saw Sambucus racemosa (red elderberry), a very large cluster of Botrychium virginianum (rattlesnake fern), and a single plant of Adlumia fungosa (Allegheny vine). In general the woods had few invasive plants, perhaps because the deer population is kept in check by hunting (Haycock Mountain is within one of the state 142 BARTONIA game lands). Other plants included Amphicarpaea bracteata, Asarum canadensis, Asclepias quadrifolia, Castanea dentata, Collinsonia canadensis, Conopholis americana, Cornus alternifolia, Desmodium nudiflorum, Galearis spectabilis, Galium triflorum, Geranium maculatum, Hepatica nobilis var. americana, Mitchella repens, Phegopteris hexagonoptera, Podophyllum peltatum, Rhododendron periclymenoides (?), Rubus odoratus, Sanguinaria canadensis, Scutellaria elliptica, Thalictrum thalictroides, Uvularia perfoliata, U. sessilifolia, Viburnum prunifolium, Viola palmata, V. pubescens, and a past-bloom orchid that we tentatively identified as Liparis liliifolia. A second stop was State Game Lands 56. The main attraction was a small moist meadow east of Lake Warren. This meadow appears to be kept open by mowing. We noted numerous plants of Liatris spicata (blazing-star), Krigia biflora (two-flowered cynthia), and Pedicularis canadensis (wood betony). In wet ditches we noted two small annuals, Juncus bufonius (toad rush) and Gratiola neglecta (clammy hedge-hyssop). On somewhat higher ground we found lots of Asclepias quadrifolia (fourleaf milkweed) and Aristolochia serpentarta (Virginia snakeroot). Other plants included Asclepias purpurascens, Cynoglossum virginianum, Hypericum perforatum, Hypoxis hirsuta, Lysimachia ciliata, Oenothera perennis, O. fruticosa, Penstemon digitalis, Polygala verticillata, Rosa caroliniana, Sisyrinchium angustifolium, and Triodanis perfoliata. Finally, we made a quick stop at Nockamixon Cliffs, which is a high wall of hornfels (a metamorphic rock) rising above the Delaware River. The north-facing orientation of the cliffs ensures that the site stays cool, and the cliffs have long been known as a site of northern plants, notably Rhodiola rosea (Sedum rosea; roseroot). We think we were able to spot this plant high on the cliffs, but, because we had neglected to bring a spotting scope, we could not be certain. Other highlights were Adiantum pedatum, Aquilegia canadensis, Arabis lyrata, Asarum canadensis, Heuchera americana, Hydrangea arborescens, Penstemon hirsutus, and Saxifraga virginiensis. Attendance: 6. Report by leader: Janet Novak. 12 July: Ramapo Mountains, Bergen, Passaic Counties, New Jersey. Joint trip with the Torrey Botanical Society. Steve Glenn, filled in for the listed leader, Gerry Moore, who was out of the country and unable to attend. The group began the trip botanzing areas adjacent to the New Jersey State Botanical Garden. Of note here were escaped specimens of Gymmocladus dioicus and a cut-leaved cultivar of elderberry, most likely Sambucus canadensis ‘Maxima’. The non-native orchid Epipactus helleborine was also noted here in flower. The group also visited Swan Pond where they observed recently planted meadows. Here the orchid Platanthera lacera was also noted in bloom. The group then walked along the Halifax Trail enroute to Glasmere Ponds. Along this trail birches and hickories were studied with the following species being identified: Betula alleghaniensis, Betula lenta, Betula populifolia (at Swan Pond B. nigra was also observed as a possible cultivated remnant), Carya cordiformis, C. glabra, and C. tomentosa. Viburnums were also studied with the following species being identified: V. acerifolium, Vv. dilatatum, v. prunifolium, V. rafinesquianum, and V. recognitum. Herbs that i ded Carex and Pycnanthemum, with the following species being recorded: C. annectans, G lurida, C. Sts - rosea, C. Pe Cc stricta, C. swanit, P. incanum, P tenuifolium, and Pvt ire altata and in bloom pe Glasmere Ponds the highlight was seeing the rare gbladdécveont species, Utricularia radiata in bloom, growing with U. macrorhiza (also in bloom), Brasenia schreberi, 2007-2008 FIELD TRIPS 143 Ceratophyllum demersum, Myriophyllum spicatum, Nuphar lutea ssp. advena, Nymphaea odorata, Potamogeton robbinsii, and Sparganium americanum. Other species of note at Glasmere Ponds included: Aureolaria flava, Cryptotaenia canadensis, Frangula alnus, Hydrocotyle anericana, Lyonia ligustrina, Salix eriocephala, and Sanicula canadensis, Thanks to Bill Standaert for maintaining a species list. Attendance 11. Report by leader Steven Glenn. 19 July: Burden Hill Forest, Salem County, New Jersey. Our first stop was a Natural Lands Trust (NLT) property of <100 acres located east of Telegraph Road (Route 540) about 1 mile SSE of the village of Alloway. Situated on the exposed, unconsolidated marine deposit of the Kirkwood Formation, this Outer Coastal Plain parcel of silt, loamy sands, and sandy clays provides a mesic to wet environment suitable for plants that are more at home on the Inner Coastal Plain of the Delaware Valley. We explored both upland and wetland forests adjacent to open farmland. The upland was dominated by a forest canopy of large and mature Fagus grandifolia, associated with similarly large specimens of Quercus rubra, Q. phellos, Q. falcata, Q. alba, Carya tomentosa, C. glabra, Liquidambar styraciflua, Liriodendron tulipifera, Ilex opaca, Prunus serotina, and Sassafras albidum. Relatively devoid of saplings and shrubs, the understory contained only scattered individuals or clusters of species such as Viburnum dentatum, Vaccinium corymbosum, V. pallidum, Kalmia latifolia, Leucothoe racemosa, Hamamelis virginiana, Clethra alnifolia, Smilax rotundifolia, S. glauca, Rubus hispidus, Euonymus americanus, and Fagus seedlings. Herbs observed included Medeola virginiana, Uvularia sessilifolia, Maianthemum canadense, M. racemosum ssp. racemosum, Dioscorea villosa, Goodyera pubescens, Tipularia discolor,* Mitchella repens, Chimaphila maculata, Monotropa uniflora, Epifagus virginiana,* Onoclea sensibilis, Osmunda cinnamomea, Thelypteris noveboracenstis, Dryopteris intermedia,* Dennstaedtia punctilobula, Viola affinis,* Dichanthelinm (Panicum) clandestinum, D. dichotomum, Leersia virginica, Carex blanda, C. swanii, and Lycopodium obscurum. This suite of species reflected a mesic, nutrient-rich environment not typically associated with Outer Coastal Plain landscapes. It appeared that this mature forest was never totally clearcut, but trees were selectively harvested over many generations. _ Immediately to the south along a tributary of Deep Run and in areas surrounding the mixed hardwood stand, we examined a wetland complex with steep stream banks and deeply incised, clayey flats. Here the canopy was primarily occupied by Nyssa sylvatica, and Acer rubrum. Occasionally flooded peri-glacial depressions, also ringed by these canopy species, were widely distributed within this forest as well as the upland forest. Scattered on the moist flats and in depressions were the following species: Vaccinium corymbosum, Rhododendron periclymenoides, R. viscosum, Woodwardia areolata, W. virginica, Carex albolutescens, C. crinita var. crinita, C. intumescens, C. lupulina,* C. lurida, C. seorsa,* C. stricta, C. tribuloides,* Carex sp.,* Juncus effusus, Viola xprimulifolia, Lilium superbum, and Dichanthelium (Panicum) microcarpon. It should be noted that several trees, shrubs, and herbs listed for the upland forest were also seen in the wetland. Both of the above communities were bisected by several canals, whose date of construction and function were not known to us. (We were surprised by the apparent absence of Carpinus caroliniana, ironwood or blue beech.) On leaving the forest, we documented the following species along a path bordering the adjacent farm field: Fraxinus pennsylvanica,* Cornus florida, Juniperus virginiana, Diospyros virginiana, Parthenocissus quinquefolia, Toxicodendron radicans, R ltiflora, Rhexia mariana, Spiraea tomentosa, Chamaecrista nictitans, Sisyrinchium atlanticum,* 144 BARTONIA Microstegium vimineum, Dichantheli imatum var. lindheimeri,* and Mublenbergia schreberi. Thanks go to Marilyn Patterson, former owner of the property, for allowing us to eat lunch in her garden and providing refreshing drinks on this hot day. About 1 mile to the south we briefly visited another NLT property, the “Telegraph Road Site,” on the west side of Route 540. Here we learned that recent timber harvesting had no immediate negative impact on small occurrences of Helonias bullata, Listera australis, and Amianthium muscitoxicum concealed in a hardwood swamp of Acer, Nyssa, and Magnolia virginiana. Among the other associated species we observed were Symplocarpus foetidus, Lilium superbum, Platanthera blephariglottis (in fruit), Sphagnum magellanicum,* Sphagnum sp. (section Sphagnum*), Carex folliculata, C. stricta, C. trisperma, C, collinsii, Eleocharis tuberculosa,* Woodwardia areolata, W. virginica, Osmunda cinnamomea, Mitchella repens, Gaylussacia frondosa, Rhododendron viscosum, Vaccinium corymbosum, and Clethra alnifolia. Approximately 4.6 miles to the southwest and 1.2 miles due west of Jericho, we examined a portion of an oak-pine stand that roughly fell within an area that the late Silas Little, federal forester, referred to as the “Salem Barrens” (Bulletin of the Torrey Botanical Club Vol. 100, No. 2: 100-101, 1973). South of Hell Neck Road and north of the Buckhorn Gun Club, this upland forest, situated on the western-most expression of the Cohansey Formation, was underlain by very sandy, well drained soils of low fertility. During Little’s investigation, there existed here a 38-year-old stand (plantation) of Pinus echinata being invaded by various oaks and other hardwoods. Our survey of this general vicinity yielded an extensive list of canopy species: Quercus alba, Q, coccinea, Q. falcata, Q. marilandica, Q. prinus, Q. stellata, Q. velutina, Sassafras albidum, Carya pallida, Nyssa sylvatica, Prunus serotina, Juniperus virginiana, Diospyros virginiana, Ilex opaca, Acer rubrum, Cornus florida, Fagus grandifolia, and only scattered specimens of Pinus virginiana, P. rigida, and P. echinata. The understory was dominated by Gaylussacia baccata, G. frondosa, and Vaccinium pallidum. Among the minor components were Castanea dentata (sprouts only), Vaccinium corymbosum, Smilax glauca, Lyonia mariana, Toxicodendron radicans, Vitis aestivalis, and Amelanchier stolonifera (=A. spicata).* Likely because of thick leaf litter only a few herbs were found: Carex pensylvanica, C. lucorum var. lucorum,* Danthonia sericea,* Nuttallanthus canadensis, Melampyrum lineare, Chimaphila maculata, and Epigaea repens. Because of a scarcity of Pinus echinata, we suspected that we may have failed to relocate the precise spot of Little’s stand. It was clear, however, that in the absence of fire and harvesting, the forest of this immediate vicinity (as Little had predicted) has become dominated by oaks, with only a minor component of pine. We added 17 new species to our survey list for the Burden Hill Forest (marked with asterisks), bringing the total count to 408. Attendance: 11. Report by leaders: Ted Gordon and Joe Arsenault. 26 July: Intermittent Ponds, Westecunk Creek Watershed, Stafford Forge Wildlife Management Area, Ocean County, New Jersey. Joint trip with participants of the Pinelands Preservation Alliance’s Special Plant Course. About 0.35 miles south of the fire house in Warren Grove, we veered southeast onto Cervetto Road on a 2.0 mile drive to our first stop. Recent heavy rains had filled deep, clay-lined potholes and treacherous ruts that brought our car caravan to a sudden halt. It was necessary to hike the remaining 1.5 miles through transitional pitch pine-shrub oaks plains, recovering from a severe May 2007 fire. En route we observed several fine patches of Epigaea repens and Pyxidanthera barbulata and the following herbs: Hypericum 2007-2008 FIELD TRIPS 145 gentianoides, Tephrosia virginica, Lechea racemulosa, Helianthemum canadense, Solidago odora var. odora, Eurybia compacta (Aster gracilis), Melampyrum lineare var. pectinatum, Apocynum androsaemifolium, and Euphorbia ipecacuanhae. At the head of Three-Foot Branch of Westecunk Creek, directly north of the junction of another sand road, we turned into “3-Foot Headwaters Pond,” partially concealed by a band of ericaceous shrubs and saplings. Underlain by Mullica sandy loam and devoid of standing water, the moist, open pond bottom of 100 yards x 21 yards contained a few thousand immature tufts of Rhynchospora knieskernii, a globally rare beaked rush. Associated rare species included emerging seedling of Scleria reticularis (LP—Pinelands listed), several rosettes of fruiting Panicum wrightianum (= Dichanthelium w.; an endangered grass), and some culms of the grass Muhlenbergia torreyana. Other species noted were Amphicarpum purshu, Panicum longifolium, P. virgatum, Cuscuta gronovil, Juncus scirpoides, Drosera filiformis, D. intermedia, Hypericum canadense, Lobelia nuttallii, Rhexia virginica, Triadenum virginicum, Xyris difformis var. difformis, Eleocharis tricostata, Rhynchospora capitellata, R. chalarocephala, R. fusca, and R. torreyana. We next explored nearby “Round Pond,” a peculiar roadedge, kettle-hole spung about 200 yards below the road split along the south-bearing road to the west. Bordered by a pitch pine lowland of ericaceous shrubs, [lex glabra, and saplings of Acer rubrum forma trilobum and Nyssa sylvatica, this 33-yard-wide pond, underlain by Mullica sandy loam, was also devoid of standing water. We observed several hundred young tufts of Rhynchospora knieskernii, associated with 6 other rare elements: Scleria reticularis and Calamovilfa brevipilis (both Pinelands-listed), Panicum wrightianum (= Dichanthelium w.), Muhlenbergia torreyana, Lobelia canbyi (one flower only), and a few leaves of Nymphoides cordata, a new find. Other companion species were Eurybia compacta, Rubus hispidus, Vaccinium macrocarpon, Gratiola aurea, Iris prismatica, Polygala cruciata var. aquilonia, Symphyotrichum (Aster) dumosum var. dumosum, Juncus effusus, J. pelocarpus, Andropogon glomeratus var. glomeratus, and all species of Rhynchospora (except R. torreyana) seen at the first pond. Since most known Knieskern’s beaked rush populations occupy sites of anthropogenic disturbance, the presence of these two natural occurrences in intermittent ponds is of special significance. All rare species discovered at these sites were already documented by the leader in Gordon, T. 1996. “De Novo Field Survey of Rhynchospora knieskernit in New Jersey.” Office of Natural Lands Management. Trenton, NJ. During the 2008 survey, it was found that these elements were still well-established. It was also learned that “Round Pond” had recently been marred by off-road vehicles. It appeared, however, that the site was able to recover from this impact. Rising temperature and humidity made the “march” back to our vehicles unpleasant. Refreshed after eating a picnic lunch, we continued to “Pond 74” on the west side of Route 539 a few hundred yards north of Governors Branch. Behaving like an intermittent pond, this shallow, abandoned, gravel excavation was almost completely dried up. In addition to Eriocaulon aquaticum, Cyperus dentatus, Rhynchospora alba, Scirpus cyperinus, and species of Lycopodiella, this pit harbored almost all the species that were documented for the two natural ponds, including a small occurrence of Rhynchospora knieskernii. On occasions, several thousand specimens of the latter sedge have been seen here. On the shore and wet, sandy flats bordering the northeast segment of the upper reservoir at Stafford Forge, we added the following species: Dulichium arundinaceum var. arundinaceum, Eleocharis robbinsii, Lachnanthes caroliniana, Nuphar lutea ssp. lutea, Nymphaea odorata ssp. odorata, Sabatia difformis, Sparganium americanum, and 146 BARTONIA Xyris smalliana. We entered the adjacent cedar swamp/quaking bog expressly to see Xyris fimbriata, the state-endangered fringed yellow-eyed grass, growing on hummocks. Among the many species seen here were Carex exilis, Eriophorum virginicum, Rhynchospora alba, R. chalarocephala, R. gracilenta, Eriocaulon compressum (decomposing), Juncus canadensis, Pogonia ophioglossoides (many leaves), Calopogon tuberosus (fruiting capsules), Oclemena (Aster) nemoralis, Rhexia mariana, Polygala lutea, Sarracenia purpurea, Lycopodiella caroliniana var. caroliniana, Woodwardia virginica, Sphagnum magellanicum, S. papillosum, S. pulchrum, and S. tenerum. At the Stafford Business Park adjacent to the landfill in Manahawkin, we visited the leader’s Rhynchospora knieskernii and Spiranthes tuberosa transplant plots established in November 2006. This controversial transplanting of rare species had the approval of the Pinelands Commission. While neither species was visible at this time, the initial response in 2007 was positive. It was clear, however, that the beaked rush was under severe stress from competing plants and the absence of suitable soil. In another section of the project’s buffer, we saw the following rare plants: Scleria pauciflora var. caroliniana, Pycnanthemum setosum, and Eupatorium hyssopifolium var. laciniatum. The trip concluded at Warren Grove where a small contingent paused to admire Bill Smith’s recently constructed bog featuring a spectacular display of pitcher plant species and a variety of other carnivorous plants. Thanks go to Bill Standaert for maintaining a list of species observed. Attendance: 22. Report by leader: Ted Gordon. 9 August: Bowman’s Hill Wildflower Preserve (BHWP), Bucks County, Pennsylvania. Joint trip with the Torrey Botanical Society. The 134-acre BHWP features over 700 species of plants, along trails in a naturalistic setting. All are species that are native or naturalized in P lvania (though not necessarily in Bucks County). It is a good place in which to observe some plants species that, though more or less local, are rather uncommonly seen in the wild. The morning of this trip was spent in a three-acre meadow at the entrance of the preserve, where plantings of such species as Helenium autumnale, Rudbeckia laciniata, Hibiscus moscheutos, Hypericum pyramidatum, Monarda fistulosa, Coreopsis tripteris, Pycnanthemum incanum, Physostegia virginiana, Senna hebecarpa, and Lobelia cardinalis made a fine show and attracted numerous butterflies and photographers. A list of species was compiled to assist the preserve in its ongoing efforts to survey their collections. After lunch in a picnic pavilion that had been reserved for our use, the field trip group moved into the central area of the preserve and walked the Marsh Marigold Trail, the Violet Trail, and the Audubon Trail. Again, species lists were compiled to complement lists made during a one-day bioblitz earlier in the year. Some of the species seen in bloom or in fruit were Elephantopus carolinianus, Aralia racemosa, Helianthus decapetalus, Rudbeckia triloba, Agastache scrophularifolia, Heliopsis helianthoides, Silphium perfoliatum, and Monarda clinopodia. Many thanks go to the Plant Stewardship Index coordinator, Jeannine Vannais, for her hospitality to the group. Attendance: 10. Leaders: Karl Anderson and Linda Kelly. Report by K. Anderson. 27 September: Delaware Bay Shore, Cumberland County, New Jersey. Joint trip with the Torrey Botanical Society. The trip began in downtown Dividing Creek. Here along 2007-2008 FIELD TRIPS 147 the roadsides the group observed large patches of the non-native grass Chloris verticillata, which is currently spreading rapidly on New Jersey’s Coastal Plain. The group then made a brief stop to Dividing Creek Crossing to see a small stand of Eupatorium resinosum in late bloom in an open boggy area adjacent to the railroad. The group’s next stop was along Hansey Creek Road. Here along the roadsides the group saw Lobelia puberula and Solidago latissimifolia (=S. elliottii) in bloom and the rare oak species Quercus michauxii and Q. nigra in fruit. The group’s last stop of the day was a recently disturbed sandy tidal area along Berrytown Road. Here the group identified numerous species of the chenopods, including: Atriplex patula, Bassia hirsuta, Chenopodium album, Chenpodium ambrosioides, Cycloloma atriplicifolium, Salicornia depressa, Salsola kali, and Suaeda maritima. Two rare species, Leptochloa fusca ssp. fascicularis (=Diplachne maritima) and Setaria magna, were also observed here. Attendance 10. Report by Leaders: Renee Brecht and Gerry Moore. Bartonia No. 65: 148-149, 2011 Program of Meetings September 2007—May 2011 Date Subject Speaker 2007 27Sept | Members’ Reports on Summer Botanizing 25 Oct Benjamin Smith Barton: Naturalist and Physician in SORPOPSOOIA AMON on lie so eh ace ev ec ed onaks wes Alfred E. Schuyler 15 Nov ___Lichens of the Limestone Barrens of the Great Northern Peninsula. ot Newloundland fais 554s + ace ew acl James C. Lendemer 20Dec ___ Botanical Collections in the Tepui Highlands of Guyana Go Siee al Gx Si eces Pe RE ce el wearer eee Erin A. Tripp 2008 24 Jan Revealing Camera Studies of Plants in the Florida Eivetades Aes Bera ek ss es ce ca eee Stevens Heckscher 28Feb — The Diversity and Evolution of Rainforest Tree Communities in the New World Tropics ............. Benjamin Torke 27Mar The Ascomycete Genus Neolecta in New England......... David Hewitt 24: Ape +. dhighlights of the Andean Flora: ccc homes. dane i Alina Freire-Fierro 22 May Rare Plant Conservation in the New si Pinelands... :3::s.: G. Russell Juelg 25 Sep Members’ Reports on Summer Botanizin 23 Oct Two Centuries of Philadelphia Botany in Névfoundian d Hus pF Sle ee Ann F. Rhoads and Alfred E. Schuyler 20Nov To Boldly Botanize: Thomas Nuttall’s Explorations in the Early Nineteenth-Century North America... ..........5.000. Steve Grund 18Dec The Latest on the Systematics of North American Atipheeco8re. Se eb an BRE Elizabeth H. Zacharias 2009 22 Jan Ecosystems and Plants of Cambodia and Thailand... .. . Tatyana Livshultz 26Feb — Notes on the Natural History and Evolution of the Cape Flows, Southern Affieg foc os ae Chris Hardy 26Mar _—_— Pennsylvania’s “Little Newfoundland”: Glacial Till Barrens ontie Pocono Plaga’. 6.0252 FG 2, BSE. es ee Roger Latham 2> Ape Vasa Perns of North America. 2... . os 2 Robbin Moran 25 May * The Piivate Lite of Inillgim rs et ee en John Gyer 148 24 Sep 22 Oct 19 Nov 17 Dec 2010 28 Jan 25 Feb 25 Mar 22 Apr 27 May 23 Sep 28 Oct 18 Nov 16 Dec 2011 27 Jan 24 Feb 24 Mar 28 Apr 26 May PROGRAM OF MEETINGS 149 Members’ Reports on Summer Botanizin Darwin and Botany in a Changing World: 150 Years after The Origin of Species Rare Plant Conservation in the Pinelands of New Jersey: umn’ Came tines ee eee Ce oe re ee Emile DeVito Biology and Restoration of Schwalbea americana (American ‘Chatiseed) in New Jersey. 6 oS ee Jay Kelley Mui: Wares 68 is ie ee a Steve Eisenhauer Meeting cancelled Ongin and Evolution of Beer. i. 65 i495 6 ie oe eh Ernie Schuyler Thomas Horsfield: Philadelphia to Java to London HeviIGtd as niles. ate bu eth Barbara Ceiga and Ernie Schuyler Vignettes from the: Coastal Plain) 20 oe oenon eee. Stevens Heckscher Members’ Reports on Summer Botanizing Climate Change Research in Northern Mongolia: the Peclopwcr warner Pastures. 0. ee ee Brenda Casper Diatoms: Philadelphia’s Most Common Algae are Beautiful and Excellent Ecological Indicators... 0.0. 6. soe. Donald Charles Floral Scents, Color and Architecture: A Look at the Floral Phenotypes of Pawpaws (Asimina: Annonaceae)......... Kate Goodrich Meeting cancelled Advancing Our Understanding of the Flora of the Northeastern U.S.A. and Adjacent Canada.............. Robert Naczi Putting Milkweeds in Context: Reconstructing The Biogeography and Ecology of the Evolution of Floral Novelty Pelton fon te Sei Gates ce cle oi on ee ne Tatyana Livshultz Mycorrhizal Fungi and Their Interactions with Plants in Rasy EVO CRG 6 ee con eh a Se + a Amy Karpati Lewis David von Schweinitz: Mycologist, Botanist, Illustrator David Hewitt eR Re ce we cee le a eo a ee Ne ee ee em OR eS ae a Om aes ee Bartonia No. 65: 150-157, 2011 2009-2011 Membership List Honorary Members FARLEY, ELIZABETH B.—600 Cathedral Rd., Apt A308, Philadelphia PA 19128, 215-984-8721, EBFAR@comcast.net SCHUYLER, ALFRED E.—The Academy of Natural Sciences, 1900 Benj. Franklin Pwky., Philadelphia, PA 19103, 215-299-1193, schuyler@ansp.org Life Members ARSENAULT, JOE—961 Clark Ave., Franklinville, NJ 08322, njplants@aol.com BARNARD, EDWARD—8038 Crittenden Street, Philadelphia, PA 19118, nedbarnard@gmail.com Bien, WALTER—144 Summit Ave, Langhorne PA 19047, 215-752-3762, wbien205@comcast.net EIsENMAN, SASHA—Temple University Dept. of Landscape Architecture and Horticulture, 201 Dixon Hall, 580 Meetinghouse Road, Ambler, PA 19002, 267-468-8168, eisenman@temple.edu FRANK, KEN—2508 Pine St., Philadelphia, PA 19103, 215-732-9280, kenfrank@aol.com FRANKLIN, CAROL—532 W. Springfield Ave., Philadelphia, PA 19118, 215-248-2783 GREENLAND, CHRISTINE MANVILLE—790 E. Street Rd., Warminster, PA 18974, 215-322-4105 Gross, MICHAEL F.—c/o Georgian Court University Biology, 900 Lakewood Ave, Lakewood, NJ 08701, 732-987-2373, gross@georgian.edu Hewitt, David A.—Department of Botany, Academy of Natural Sciences, 1900 Benjamin Franklin Parkway, Philadelphia, PA 19103, 617-495-2368, davidhewitt@post.harvard.edu Hott, Rosert J.—1611 Smith Bridge Road, Chadds Ford, PA 19317, 610-584-5578, jack@ mobilelifts.com Hurwitz, ALISHEBA—215 Harrogate Road, Wynnewood, PA 19096, 610-322-1217, ali.hurwitz@ gmail.com IRETON, Mary Lou—213 4th Ave., Haddon Heights, NJ 08035, 856-547-1118 LauER, DaviD—49 Cornell Ave., Churchville, PA 18966, 215-357-2646, DML1000@comcast.net MACIEJEWSKI, STEPHEN—2030 Fitzwater St., Philadelphia, PA 19146, teciu@verizon.net McLean, ELIZABETH—139 Cherry Ln., Wynnewood, PA 19096-1208, 610-642-4196, Epmclean@ comcast.net McLEan, WILLIAM—139 Cherry Ln., Wynnewood, PA 19096-1208, 610-642-4196, Epmclean@ comcast.net Moore, GERRY—East National Technology Support Center, Natural Resources Conservation Service, USDA, 2901 East Lee St., Suite 2100, Greensboro, NC 27401, 336-370-3337, Gerry. Moore@gnb.usda.gov Novak, JANET—19 West Willow Grove Ave, Philadelphia, PA 19118, 215-248-2642, janet@indri. org O’ HERRON, JOHN—PO Box 126, 129 Juniper St., Cornwall, PA 17016-0126, 717-277-0587, joherronII@msn.com OLSON, WILLIAM—1005 Lakewood-Farmingdale Rd., Howell, NJ 07727, 732-938-3187, Wolson3@ optonline.com ALMER, MATTHEW—1518 Millstone River Rd., Hillsborogh, NJ 08844, 908-359-2073, mp2434@ columbia.edu PaTRICK, RUTH—750 Thomas Rd, Philadelphia, PA 19118 PETROW, CAROL—60 McLeod Pond Rd., Glenmoore, PA 19343 150 MEMBERSHIP LIST 151 QUIGLEY, PATRICIA ANN—1080 Quarry Hall Road, Norristown, PA 19403, 610-584-1829, paqinc@aol.com RHOADS, ANN—3 Blythewood Rd., Doylestown, PA 18901, 215-348-8139, rhoadsaf@exchange. upenn.edu ROBERTS, WILLIAM H.—1806 Delancey St, Philadelphia, PA 19103, 215-569-5632, roberts@ blankrome.com STAILEY, HELEN M.—6555 Tabor Ave. Alpha 1709, Philadelphia, PA 19111-5385, 215-697-8037 STALTER, RICHARD—Dept. of Biology, St. John’s University, Jamaica, NY 11439, 718-990-6269, stalterr@stjohns.edu STARTUP, MICHAEL—718 Price St., West Chester, PA 19382, 610-692-5943, mstartup@haverford. d edu STEVENS, CHARLES E.—615 Preston Pl., Charlottesville, VA 22903, 804-293-8658 TEESE, PAUL—1183 Apple Road, Quakertown, PA 18951, 215-862-2924, teese@bhwp.org THOMPSON, SUE—129 E. Sycamore St., Pittsburg, PA 15211-1719, 412-622-3295, sthomp@ andrew.cmu.edu TREADWAY, SUSAN—1509 Monk Rd., Gladwyne, PA 19035, 610-642-8050, sptreadway@aol.com Sponsoring Members CLARKE, TERENCE—15 East 6th Rd., Broad Channel, NY 11693, chelseaclarke@att.net FERREN, WAYNE JR.—67 Main St., Vincentown, NJ 08088, 609-859-8682, wrfjrl@comcast.net HENRY FOUNDATION FOR BOTANICAL RESEARCH—P.O. Box 7, Gladwyne, PA 19035-0007, 610- 525-2037 JUELG, RUssELL—79 Grassy Lake Rd., Shamong, NJ 08088, 609-654-6455, russell@pinelandsalliance.or: McBribe, DoNNA—4 Gentian Street, Browns Mills, NJ 08015, 609-893-3732, donnamcbride57@ gmail.com McComse, Ropert—445 E. Main Street, Moorestown, NJ 08057, 609-313-5000, robmccombe@ verizon.net O’BRIEN, JOAN—PO Box 277, Maple Shade, NJ 08052, 610-329-1029 SOBEL, MARILYN—329 Haverford Place, Swarthmore, PA 19081, 610-604-0604, m_sobel@msn.com WHITFORD, STEVE—41 Main Street, Port Republic, NJ 08241, whitfordsp@gmail.com Regular and Student Members ANDERSON, KARL—46 N. Childs St., Woodbury, NJ 08096-1535, 856-845-7075, karlanderson6@ aol.com ANGuUs-ANDERSON, BLAKE—306 Hunter Street, Gloucester City, NJ 08030, 856-456-6132, blakeaaa@comcast.net ARMSTRONG, REBEKAH—1357 E. Hewson St., Philadelphia, PA 19125, 215-713-7611, rebekah. armstrong@gmail.com AUSTIN, JASON—419 Hillside Ave., Pine Beach, NJ 08741, 732-966-7541, diamonddobbs@yahoo. BAKER, MIKE—6 Hill Lane, Cream Ridge, NJ 08514, 609-577-9004, mike@mikebaker.com BarRINGER, KERRY—126 Hutton Street, Jersey City, PA 07307, 201-659-1158, kerrybarringer@ .com BAuCHSPIES, JAMES T.—4320 Chetwin Terrace, Easton, PA 18045-4908, 610-253-8925 BAUMGARTEN, JONI—4405 Osage Ave, Apt. 3, Philadelphia, PA 19104, 215-870-5654, joni. baumgarten@gmail.com BENJAMIN, JESSIE L.—1062 Glen Hall Rd., Kennett Square, PA 19348, 484-887-8612, jessie@ taprootnativedesign.com BrssELMAN, THOMAS H.—147 Canna St, Browns Mills, NJ 08015-2040, oji1160@verizon.net Best, JAMES—1262 Ashbourne Rd., Elkins Park, PA 19027, jvb262@comcast.net 152 BARTONIA BIDDLE, DORRELL—701 Washington Ave., Palmyra, NJ 08065, 856-499-2343, dell2733@aol.com BINCKLEY, SARAH—8101 Gladstone Road, Wyndmoor, PA 19038, 828-686-7358 BLAUSTEIN, MAXIMILIAN—6102 Ardleigh St., Philadelphia, PA 19138, maximilianjb@gmail.com BLock, TIMOTHY—203 Juniper Court, Zieglerville, PA 19492, 610-754-1885, Block@exchange. penn.edu BowELL, MICHAEL W.—2148 Bodine Rd., Malvern, PA 19355, 610-827-1268, mwb@createascene. com BoyLe, MARY—1080 Plowshare Road, Blue Bell, PA 19422, 484-231-1408, mboyle299@yahoo.com BRAXTON, J.W.—4712 Windsor Ave., Philadelphia, PA 19143, 215-724-1571, morbraxton@aol.com BRECHT, DAWN—2244 East Main St, Millville, NJ 08332, pbfiddler@yahoo.com BRENNER, HUGH—202 Edgemont Ave, Ardmore, PA 19003, hbrenner54@gmail.com Buck, WILLIAM R.—Institute of Systematic Botany, New York Botanical Garden, Bronx, NY 10458-5126, 718-817-8624, bbuck@nybg.org CANTRELL, GREGORY—48 Mariners Cove, Freehold, NJ 07728, gregory_cantrell@yahoo.com aaee: sensi E. Highland Ave., Philadelphia, PA 19118, 215-242-0734, carrwechsler@ CHAMIERLA AMY—570 State Route 2, Shelburne NH 03581, 207-443-4718, chamberlainedit@ aol.co ce REBECCA—4200 Pine St. Unit 204, Philadelphia, PA 19104, rebeccachance@gmail.com CHERRY, LYNN—927 Black Rock Road, Gladwyne, PA 19035, rudy927@yahoo.com CoLe, MARGARET—106 Woodridge Place, Laurel, MD 20724-1800, 202-230-6616, m.e.cole@ me.com CONNOR, JACK—35 Clarks Landing Rd., Port Republic, NJ 08241, jconnor@stockton.edu CONNOR, JESSE—35 Clarks Landing Rd., Port Republic, NJ 08241 Coomss, ALEXANDRA—1086 King Road, Apt. FIR-114, Malvern, PA 19355, naturechic14@ hotmail.com CouRTNEY, JOHN—439 Gladstone Ave., Haddonfield, NJ 08033, 856-429-4987, courtneyjohn9@ gmail.com CRAWFORD, ALLEN—508 Emma St, Mount Holly, NJ 08060, email@planktonart.com Crooks, CARLA—403 Watson Ave, Horsham, PA 19044, carlitacru 00.com UNNINGHAM, RoBERT—9 W. Whites Bog Road, Brown Mills, NJ 08015, beltane00@aol.com Davis, pe A.—1510 Bellona Ave., Lutherville, MD 21093, 410-252-4154, charliedavis1@ sigm: me” tite M. —14 Cemetery Lane, Schwenksville, PA 19473, 610-316-0036, entoman@comcast. DE Coad. LinDA—1100 Concord Dr., Bridgewater, NJ 08807, lindadecastro@optonline.net DeLL, RACHEL—PO Box 7399, PMB 450, Breckenridge, CO 80424, rachelrdell@gmail.com DENNE-HINNOV, Bo£L—32 Morton Court, Lawrenceville, NJ 08648, 609-921-1272, hinnov2001@ yahoo.com DENNY, sant Mount Gilead Rd., Fredericktown, OH 43019-9513, 740-694-6087, guydenny@ecr. DIEDRICH myers Te Highland Terrace, Pitman, NJ 08071, 856-589-8455 DIETRICH, "JuLE—2914 W. Flora St., Philadelphia, PA 19121, 215-898-1708, julie2124@gmail.com Dyo ts, IsSABELLE—6626 Morris Park Road, Philadelphia, PA 19151, isabelledijols@yahoo.com aise JiLL—56 Tumble Falls Rd., Stockton, NJ 08559-1309, jsdodds@biostarassociates.com RN, RUTH—1014 Dolphin Ave., Croydon, PA 19021, 215-499-0104, rdorn@aesop.rutgers Dear , THOMAS J.—2022 Montrose Street, Philadelphia, PA 19146, sini SO SEs net EBERT, JANET—1611 Smith Bridge Road, Chadds Ford, PA 19317-9765, 610-459-0585, jack@ mobilelifts.com EDINGER, GREG—442 ae Rd., , Greenwich, NY 12834, 518-692-1725 EsTEN, JOHN—2401 Per Ave. 5C-52, Philadelphia, PA 19130, 215-235-1521 MEMBERSHIP LIST 153 FERGESEN, SUSAN—209 Laurel Court, Morgantown, PA 19543, 610-913-0542, sfergesen@yahoo. FIELD, STEPHEN—5 Evelyn Ave., Vineland, NJ ewe FIELD, THERESA—5 Evelyn Ave., Vineland, NJ 0836 FINE, NORMAN—73 Nature Lane, Sewell, NJ 08080, a mama _— BEVERLY—634 Revere Road, Merion Station, PA 19066, 610-667- sss, beverlyfitts@verizon. .. Tont ANN—662 West Johnson Street, Philadelphia, PA 19144, 215-858-1692, toniann@ philadelphiagardens.com FoLin, GRANT—319 Titan Street, Philadelphia, PA 19147, gfolin@verizon.net ForD, JOANNE B.—729 Westview St., Philadelphia, PA 19119, 215-844-8054, Joanne.B.Ford@ gmail.com Fox, ANN—5741 Cranberry Ct., Egg Harbor City, NJ 08215, atthebog812@verizon.net FREIRE-FIERRO, ALINA—1147 S Mole St, Philadelphia, PA 19146-3102, alina_ff@yahoo.com FULLERTON, WILLIAM—1104 Blythe Ave., Drexel Hill, PA 19026, 610-446-0968, sharnbill@msn. com GarBACK, Mary E.—3839 Janice St., Philadelphia, PA 19114-2826, 215-332-7105 Gorr, ELINOR 1—275 Bryn Mawr Ave., Apt. K17, Bryn Mawr, PA 19010-4246 GoopricH, KATE—Dept. of Biology, Widener University, 1 University Place, Chester, PA 19013, 610-499-1086, kgoodrich@mail.widener.edu sie PaTRICIA—31 Burrs Mill Road, Southampton, NJ 08088, 609-859-3566, Gordontrish@ bt TED—31 Burrs Mill Road, Southampton, NJ 08088, 609-859-3566, pbitgordon@aol.com Graciz, CAROL—19 North Lake Circle, South Salem, NY 10590-1014, carol.gracie@gmail.com Gray, DENNIS—668 Princeton Ave., Collingswood, NJ 08108, 856-858-6642, gray@marine.rutgers. edu GREMBOWICZ, DUANE—2210 Tiiu Court, Toms River, NJ 08755, 732-349-0364, dgrembowicz@ ocean.edu RIM, DEBRA—710 Valentine Street Apt 2, Bellefonte, PA 16823, 814-234-2344 ext 223, dgrim@ inligzsearch com ND, STEVEN—1609 Cumberland St, Pittsburgh, PA 15205, SGrund@paconserve.org ons Joun—P.O. Box 185, Clarksboro, NJ 08020, 856-423-3889, fernhill@voicenet.com HACKER, SUSAN—1501 Darmstadt Ave., Egg Harbor City, NJ 08215, 609-965-3297, susanspero1@ verizon.net HAkGELE, ELIZABETH—235 Mather Road, Jenkintown, PA 19046, haegele@sas.upenn.edu HALLIWELL, THOMAS—19 Kings Rd., Netcong, NJ 07857, 973-347-6071, widget a a net Harpesty, GAIL—488 Big Oak Road, Morrisville, PA 19067, evergreem@aol.co HARLAN, — Quaker Hill Rd., Warren, PA 16365, 814-406-4239, maggie811@ atlanticbb.n Harris, an M. —4401 W St NW, Washington, DC 20007-1133, 202-338-9083, jes405@hotmail. m co Harris, Louis C.—1002 Abington Rd., Cherry Hill, NJ 08034, “eee sie net Heavers, NATHAN—171 Longview Drive, Princeton, NJ 08540, heavers@gm mail.co HECKSCHER, STEVENS—1030 E. Lancaster Ave. Apt. 314, Bryn Mawr, PA 19010, 610-527- 6607, sheckscher103@comcast.net HELD, MICHAEL E.—Department of Biology, St. Peter’s College, Jersey City, NJ 07306, 201-761- 6430, mheld@spc.edu HELM, SARA—303 W. Miner St., Apt. 2 Rear, West Chester, PA 19382, sara. helm.wallace@gmail.com g gunewiabos Larry—420 Walvebiook Drive, Norristown, PA 19403, 484-744-7035, allbethere@ comcast.n Hokss, ae Capano Dr. Apt. C5, Newark, DE 19702, choess@stwing.org 154 BARTONIA Homes, MARION—438 South 44th St., Philadelphia, PA 19104, a.triphyllum@gmail.com Hrena, ALYCE—639 Lafayette Ave., Trenton, NJ 08610, 609-888-2672, alycehreha@optonline.net Hucues, COLIN—P.O. Box 707, Exton, PA 19341, 610-524-0424, km2442@comcast.net Hucues, STUART—721 E. Butler Pike, Maple Glen, PA 19002, stuhu2@comcast.net HUMBERT, KAREN—2401 Pennsylvania Ave., # 7A6, Philadelphia, PA 19130, kchumbert@hotmail. o HuTCHEON, DaviD—3229 Iris Ave., Trevose, PA 19053, 267-994-5103, davidhutcheon@comcast. net INSKEEP, rien it N. Wayne Ave. Apt. 204, Wayne, PA 19087, 610-989-9358, mike@ gentlehelpers.co oS a Bainbridge, Apt 3, Philadelphia, PA 19147, jakubowski_anna@yahoo. TANGA: JEFFREY—1801 Buttonwood St. Suite 1610, Philadelphia, PA 19130, janoski@sas.upenn. d edu JeNEY, RopyN—6 Chestnut Ave., Haddon Township, NJ 08108, 856-287-3727, robyn_jeney@ mail.com Jess, RoBIN—55 Lahiere Ave., Edison, NJ 08817, 732-572-5928, rjess5928@aol.com JOHNSON, ALAN—63 Central Avenue, Audubon, NJ 08106-1903, aajohnson1@gmail.com JOHNSON, KATHERINE—312 E. Gorgas Ln., Philadelphia, PA 19119, 267-614-0587, bluejaykj- PBC@bacteriophages JOHNSON, KirsTEN—550 W University Pky, Baltimore, MD 21210, kirsten2924@gmail.com KarGER, MERMIE—P.O. Box 707, Exton, PA 19341, 610-524-0424, km2442@comcast.net Ke.ty, LinpA—60 Forest Drive, Lakewood, NJ 08701, 732-363-1266, kelly LC01@aol.com KHAN, NANCY—4323 Osage Ave., Philadelphia, PA 19104, nancyrkhan@yahoo.com KIMELMAN, GAY—2212 St. James St., Philadelphia, PA 19103, 215-563-0285, gkimelman@yahoo. om KLIGMAN, DouGLAS—1198 Rossiter Lane, Radnor, PA 19087, 484-432-0794, kligahm@earthlink. et Kiotz, LARRY—27 Colonial Ct, Shippensburg, PA 17257-9588, 717-532-8978 (h) 717-477-1402 (w), larry_klotz@hotmail.com Kos_er, EvVELYN—118 South 21st Street #106, Philadelphia, PA 19103, 215-988-1774, koblerev@ verizon.net Kuna, JOsEPH—3523 Hamilton St., Philadelphia, PA 19104, joseph.kuna@yahoo.com LaripiGc, Kim J.—712 Wood Lane, Haddonfield, NJ 08033, 856-354-3296, kim.laidig@gmail.com LAMONT, Eric—717 Sound Shore Rd., Riverhead, NY 11901, 631-722-5542, elamont@optonline.net LATHAM, ROGER—P.O. Box Rose Valley, PA 19086-0057, 610-565-3405, rel@ continentalconservation.u trex; CHARLES—105 Kendall Rd., Kendall Park, NJ 08824, 732-821-8310, oes edu Leck, MARY—105 Kendall Road, Kendall Park, NJ 08824, 732-821-8310, leck@rider.edu = ras East 6th Road, Broad Channel, NY 11693, 718-945-5009, Sicksedhadtie@iaichiicl’ Herp WILLIAM M.—Duke University, 420 Anderson St., Box 90341, Durham, NC 27708-0341, bill. lefevre@duke.edu LENDEMER, JAMES—1245 Park Ave. 16J, New York, NY 10128, 215-801-6948, jlendemer@nybg.org Levin, MicHAEL H.—414 Mill Rd., Havertown, PA 19083, 610-449-7400, eraincmhl@aol.com LEWANDOWSKI, RICK J.—Mt. Cuba $88 P.O.Box 3570, Greenville, DE 19807-0570, 302-239- 4244, rlewandowski@mtcubacenter.or saad TATYANA—2122 Mt. Vernon’ St., #2, Philadelphia, PA 19130, 617-710-7090, livshultz@ ansp.O' Eervici: CaroL—Biology Department, Dickinson College, P.O.Box 1773, Carlisle, PA 17103- 2896, 717-245-1360, loeffler@dickinson.edu MEMBERSHIP LIST 155 LoFurno, MicHakEL—2028 Fitzwater St., Philadelphia, PA 19146, 215-732-0849, mlfcomp@aol. com LONGWOOD GaRDENS LisraRy—P.O. Box 501, Kennett Square, PA 19348-0501, 610-388-5440, library@longwoodgardens.org LYNCH, JEAN—PO Box 127, Cape May, NJ 08204, jean.lynch.1@gmail.co Maher, CAROLE—774 W. Church Road, Elkins Park, PA 19027, S hclecaes asin et Mater, AGNES LEE—1815 JFK Blvd. Apt. 2209, Philadelphia, PA 19103, 267-773-7784, maier. agnes@gmail.com MALONE, SARA—1275 County Road 519, Frenchtown, NJ 08825, 908-996-3402, malonesa@sas. upenn.edu Martin, Harris W.—5100 Bay Rd., Bensalem, PA 19020, 215-208-5264, harrismartin3@yahoo. co MartTAN, STEVE—1475 Mt. Holly Road, B5 Cooper Valley Village, Edgewater Park, NJ 08010, 609- 877-4292, stevemattan@comcast.net McAvoy, WILLIAM A.—1291 Fords Corner Rd., Hartly, DE 19953, 302-492-3541, william. mcavoy@state.de.us McConneLL, ScoTr—PO Box 572, Haddonfield, NJ 08033, scitchyrooroo@yahoo.co McCrea, EsTHEr J. —404 Cheswick Place, Apt. 408, Bryn Mawr, PA 19010, 610-527- 1089 McGanney, JULIANA—2307 Lombard Street, Philadelphia, PA 19146, 917-309-6222, juliemcganney@gmail.com McKay, NAN—Box 370, 214 P lvania Ave, Mt. Gretna, PA 17064, nanmckay@gmail.com —— RICHARD—200 South Flint Court, Yardley, PA 19067, 215-493-0697, Se adilinsinieicinet on Mary BeTH—40 Jennings Rd., Medford, NJ 08055, 609-790-6044, mltn_mrybth@yahoo. com MickLE, ANN M.—Department of Biology, LaSalle University, 1900 W. Olney Ave., Philadelphia, PA 19141, 215-951-1254, mickle@lasalle.edu MILNER, Norma M.—305 Westtown Circle, West Chester, PA 19382-7653, 610-344-0860, normamilner@comcast.net MITCHELL, JOHN—25 Central Park West Apt. 9P, New York, NY 10023, 917-842-6940, katherinemitc@gmail.com MITCHELL, nine aya Central Park West Apt. 9P, New York, NY 10023, 917-842-6940, kashevinentive@eriis il.co sags geil Fue Racloay III, Salem Blvd, Berwick, PA 18603, 570-542-2191, jimm37@ verizon.n Moroz, regener Park Ave. 16J, New York, NY 10128, 215-801-6948, jlendemer@nybg.org Morris, WAYNE—45-12 Pearson Street, Long Island City, NY 11101, 718-361-7627, treeman16@ .com Morse, LARRY—PO Box 34237, Washington, DC 20043, 202-347-6822 MUELLER, Betry Lois—118 Race St., Apt. 11, Kennett Square, PA 19348, 610-444-9977, seagull9977@verizon.net MurpHy, Mary—2042 Maplewood Ave., Abington, PA 19001, 215-694-4222, bam842@cs.com Naczi, RoBERT—New York Botanical Garden, 2900 Southern Blvd., Bronx, NY 10458, 718-817- 8087, rnaczi@nybg.org NESWALD, BARBARA—2034 Arch St., Philadelphia, PA 19103, 215-563-5412, b.neswald@comcast. net NEWSTEAD, EDWIN—270 Roseland Ave., Essex Fells, NJ 07021, 973-226-7651, enew001@aol. com NICKERSON, Gary W. JR.—159 Holly Ave, Penndel, PA 19047, gwn099@ O’BrIEN, TERRY—3 Mystery Rose Lane, West Grove, PA 19390, 484-667-8418, obrien@rowan.edu Pace, MarTIN—9 Old Covered Bridge Rd, Newtown Square, PA 19073-1211, Martin_Page@msn. com 156 BARTONIA — a Christina Mill Dr., Newark, DE 19711, 302-353-9052, zoepanchen@ sympat Peants ets —234 Broughton Lane, Villanova, PA 19085, mikiperez@comcast.net PEREZ, NykiA—2125 E. Norris St., Philadelphia, PA 19125, 215-284-6383, nykia.perez@gmail.com PLyLerR, DoROTHY—418 Cadadeud Dr., Kennett Square, PA 19348-2050 PUNNETT, HopeE—6635 Wissahickon Ave, Philadelphia, PA 19119, hpunnett@verizon.net Rapis, RICHARD—69 Ogden Ave., Rockaway, NJ 07866, 973-586-0845, isotria@verizon.net RocHE, LESLIE—73 Hull Ave., Srsekold, NJ 07728 ROSENBERG, ANN—5 Westview Road, Bryn Mawr, PA 19010, 610-525-8683, rosiebig@voicenet. com ROSENBERG, D1CcK—5 Westview Road, Bryn Mawr, PA 19010, 610-525-8683, rosiebig@voicenet. co RucH, PAMELA—228 Cherokee St., Emmaus, PA 18044, 484-225-6679, pamruch@hotmail.com RupyJ, EricH $.—1021 Irwins Choice, Bel Air, MD 21014, 410-420-8973, erudyj@aol.com Rue, MARGARET—209 Arden Rd., Gulph Mills, PA 19428 SCHMIDT, GERRIE—733 Bradford Alley, Philadelphia, PA 19147, 267-324-3081, gerrichope@gmail. co SCHMITT, WILLIAM—1470 Burgoyne Road, Downingtown, PA 19335, 610-363-7693, schmitt@ temple.edu SCHNEIDER, GEORGE—345 Nursery Rd., Wellsville, PA 17365, 717-292-4035, gwsam@att.net SCHNEIDER, ew 340 Sugartown Road, Apt. C93, Devon, PA 19333, 610-964-0210, schndrw@aol.co SCHUBERT, panien rome Road, Apt. D, Springfield, PA 19064, 610-328-2759, paulschubert88@yahoo ssgusw JOHN—55 Hertzog Shoat Road, Mertztown, PA 19539, 610-682-2809, johnscott302@aol. Serve, KENNETH T.—219 Maple St., Jersey Shore, PA 17740, 570-398-2546, ksettlem1@ com Pe peel Burnham Road, Philadelphia, PA 19119, beezsveet@gmail.co SLATER, co fovepsgpiose New Holland Road, Mohnton, PA 19540, 610-775- ~— mslater@ voicenet.c SMEJKAL, Bi--97 Old Well Road, Easton, PA 18042, isotria@verizon.net SMITH, A. cgnnermaaniete Route 539, Warren Grove, Barnegat, NJ 08005-3010, 609-709-0769, bogmanbill@gmail.co SMYTH, BRAD—840 S. aed Street, Philadelphia, PA 19147, smyth555@yahoo.com SOLOMON, SEAN—6626 Morris Park Road, Philadelphia, PA 19151, isabelledijols@yahoo.com Spitz, LAUREN—370 David Drive, Langhorne, PA 19047, 267-391-6286, lauren.spitz@comcast.net STAHL, STANLEY—104 Harvard Ave., Lancaster, PA 17603, 717-293-0292, sweepersuzi@yahoo.com STANDAERT, WILLIAM F.—45 Maltbie Ave., Apt. 19B, Midland Park, NJ 07432, 201-612-9069, wiszzz@aol.com STANTON, MICHAEL—15 Higbee Ave., Somers Point, NJ 08244, Michael0448@comcast.net SWEETMAN, HAROLD—631 Berwyn Baptist Rd., Devon, PA 17333 SZUTARSKI, MARK—2301N Lakeshore Drive, Browns Mills, NJ 08015-2909, 609-735-1382, mszutal1046@aol.com TayLor, SARAH—3622 Calumet St., Philadelphia, PA 19129, 610-357-0564, sarahlatifa@hotmail. com TERRAMIN, NANCy—66 Llangollen Lane, Newtown Square, PA 19073, nterramin@gmail.com TIPPING, Curis—1080 Plowshare Road, Blue Bell, PA 19422, 484-231-1408, mboyle299@yahoo. com TREHER, AMANDA—426 Burnside Street, Apt 2, Annapolis, MD 21403, amanda.treher@gmail.com TREXLER, NANCyY—7501 Germantown Ave, Philadelphia PA 19119-1606, 215-242-1309, ntrexler@ MEMBERSHIP LIST 104 VERNICK, SANDRA—44 Seven Oaks Circle, Holmdel, NJ 07733, sandra.barenbaum@gmail.com WECHSLER, DouG—314 E. 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