Digitized by the Internet Archive

in 2012 with funding from

Royal Ontario Museum

http://archive.org/details/newspeciesofglauOOpete

®03

S/ty/

tier*

&

.s^-**

iSlifflhii. . ""'u MUSEUM LlbKAKIES

3 1761 05162347 8

ROM

Life Sciences
Occasional Papers

Royal Ontario Museum
August 17, 1973

No. 22

A New Species of Glauconycteris
(Vespertilionidae, Chiroptera)

by R. L. Peterson1 and Donald A. Smith2

In a systematic study of the African genus
Glauconycteris Dobson, we have found a
series of white-winged specimens that repre-
sent a new species that has not been pre-
viously described. The genera (or sub-
genera) Chalinolobus Peters of the Austral-
asian region and Glauconycteris from Africa
are closely related as discussed by Dobson
(1875, 1878), Miller (1907), Ryan
(1966), Hayman (1967), Hayman and
Hill (1971), and Koopman (1971). Argu-
ments have been advanced that all species
belong to one genus {Chalinolobus), that
the two should be considered subgenera, or
that generic distinction should be retained
for the two taxa, an option that has been
widely followed in recent years. We tenta-
tively recognize Glauconycteris as a genus
pending the results of our own analysis of
variation within each of the African species,
now in progress.

We have compared the new taxon with
all previously described species of Glau-
conycteris, but in our opinion, it needs
critical comparison only with G. argentata
(Dobson) and G. variegata (Tomes) as it
differs so distinctly either in size or in pelage
and wing coloration (or both) from all other

known species. We are pleased to name this
new species after Mr. Robert Glen, Nairobi,
Kenya, who collected the holotype, assisted
in the collection of the Uganda series of the
new species, and who has been a field com-
panion extraordinaire.

Materials and Methods — A total of 154
specimens of Glauconycteris from Africa,
representing eight different species, have
been examined and measured in this study
(see specimens examined). The specimens
examined were from the collections of the
following institutions: American Museum of
Natural History, British Museum (Natural
History), Carleton University Museum of
Biology, Carnegie Museum, Field Museum
of Natural History, Institut Royal des Sci-
ences Naturelles de Belgique, Los Angeles
County Museum, Museum of Comparative
Zoology - Harvard, National Museum of
Rhodesia, Royal Ontario Museum and
United States National Museum. All mea-
surements are given in millimetres and
weights in grams. The measurements used
follow those as defined by Peterson (1972).
Specimens were sorted by sex and age, and

1 Department of Mammalogy, Royal Ontario Museum, Toronto, Ontario.

2 Museum of Zoology, Department of Biology, Carleton University, Ottawa, Ontario.

«„««« « """^Z-

X'iW

Fig. 2 — Upper dentition of three species of Glauconycteris (scale bars = 1 mm). Left, G. argentata ROM
50501 $ , Kakamega Forest, Kenya (C-M3 4.3 mm). Centre, G.g/em Holotype rom 57125 9, Lomie.Cameroun
(C-M3 4.75 mm). Right, G. variegata rom 40141 $ , Budongo Forest, Uganda (C-M3 4.8 mm).

the measurements were analyzed using an
IBM/370 computer at the University of
Toronto Computer Centre. The means (X)
and standard deviations (SD) of all char-
acters were calculated for male and female
adults, and the differences in character
means between species were examined using
Student's /-test (P). Differences were con-
sidered significant at the 0.05 probability
level. Selected cranial specimens were photo-
graphed with a scanning electron microscope
using the technique described by Howden
and Ling (1973).

Glauconycteris gleni sp. n.

Holotype — ROM 57125, adult female, skin
and skull collected 16 June 1970 by Robert
Glen and V. Ngam in a tropical forest region
near Lomie, Cameroun, 3°10'N, 13°37'E.
Geographic distribution — Known only from
the type locality in Cameroun and from
Malabigambo Forest near Sango Bay, Ma-
saka District, southern Uganda, 1°05'N,
31°30'E.

Diagnosis — Wings and uropatagium white
with dark brown pigment outlining the bones
of the wings, hind limbs, and tail, the basal
portions of the calcars and veinations of the
interfemoral membrane, and to a lesser ex-
tent the proximal areas of the wings between
humeri and hind limbs (Fig. 1). The ears
are relatively large for the genus and have
broad tragi. The pelage of the dorsum is
quadricoloured with a pale clay-coloured ter-
minal band, a narrow and dark-brown sub-
terminal zone, a pale, buffy, sub-basal zone,
and a dark slate-gray basal band. There are
no distinct dorsolateral bands of pale pelage
as in G. argentata. Ventral hairs are bi-
coloured, with the base gray and the whitish
terminal band varying from pure white to a
wash of pale yellowish buff. The length of
the tibia relative to forearm length is short
compared with other species of the genus
(Tables 1 and 2).

The size and general shape of the skull
of G. gleni is similar to that of G. variegata
but differs from the latter by the combin-
ation of a more rounded braincase (equal in

Fig. 1 — Dorsal and ventral aspects of the holotype of Glauconycteris gleni ROM 57125 9
(total length 100, forearm length 41.4).

a

w

a

S>

Oj

^

J.

«

tn

a

B

nO

<u

■o

S

(/}

1"

<5

U

bo

—

o

(3

•a
c

A

/— *s

E
E

DioSatADa '£)

Tg \U3\8 -Q

u

a

-I

CO

u

<

-

H

o

(A

"3
E

"3
-o

CD

■—
O

Ml

Q

I

s
1

}U3\S •£) 1p

vwiuaSAD [)

X

|x

IX

ooooooooo

n O Q O m o o

ooo o o o o

ooooooooooooooooo
A A A VI VI VI A A A A VI VI VI VI VI VI A

oo r-l no — on — Ttr~Tj- — mmoooooomTl-

OOOOOOOOO

A VI A VI VI A VI VI VI

NONOONOOflOmoOON

•n \o — o t- r- no cn no

<N<N— i <N — <N — — —

ooooooo
VI VI VI VI VI VI VI

en t~~ oo m rn no m
m oo O pi — CN -t

mm OOOOOOOOOO — 00\0(N

ONOOOONOONOOONONOOONONONONONONNOO

OOOOOOOOO

vovoooor-ONt-^oooo

OOOOOOO
r~ oo oo oo oo oo c-

m on r- o

III I III I J I I III I J. 1
oo O t-~ — m r- r«-i — o I-- OO^IMOVO
00 rf ............ o

xOM»inOr--"0"iMvorn
— mm — <N m — < — m

m<N — r-Tj-mOTj-oo — or--r-~Or-- m

in oo r- so

• -NO ■N"1NON 00 — I <N <N • r- fi

•* m • o O • •

iih *a- ,-. o\ oo no no no -tMnvirtmm

I I I i I I T i I iiiiiii

mmtNOr-r-NOmin cj m m (^ r-~ m o

m oo ..(».....

• ■^■OooMnin'o tj- no • Tt on m m

mrirtoxos^imwi ^j- no ^j- Tt O m m

\OMrtfn-'(N^Doo^FHtH(Noornr^o>x
m — »nmr--OmTtmooooOmTtoo,nNO

<N — — (NCn) — — — <N

<N rj- m r- Tt Tt r-~

O — O NO i-l — l f-

fN| — _ _ —I — . O

(N — OOO — — ■OO — O O — O O m O

ONONONONONONONONONONONONONONONr^ON

m • ■ O in in •

Or-OTtt-- — (NNOmO — — no ■ • O <N
« Tt ~- — — Tt Tt — <N Tt — — m On no m — i
M I I IJllll I I I J J J. I
no Tt oo m — ■ OOTrOONOot^mTtoOm
ON TJ- — on •

^nn^Mn m

OOOOOOOOO

00r-~ON0000ONNOONON

in O — m
• • >n • I— <N O r- On

*t ■* • ©

Till
i tj- o >n i— i i~-

m<N <N
IT) . .

■ in in

ooooooo

ON ON ON ON ON ON on

in in in (N in
no Tt oo r-~ • C4 —

• • • • O ■ •

■* no ■* ■* — in m

I 111 III
m in m m on On on

ON — I

• • tj- ^r on ■* <n

m no

r-NOONONNOt--ONOOfNTtoomt— m ■* OTtNOinON<N<N«NO

^^DooooOOrt^-iri'tr^^^rnxm^ — r^(Noom©r--inTt

Tt <N <N rj- — i O Tt

mmr^\oo-io\

ONinoomNOO©in~-oo©©inooinooO TtmrtONooooininin

ON Tt _«TtTt — (Nm — — m ON— — —

<N

'tvO'ttO'niN

inmoNr-- — © — no — NOmt^OmoOfN
no — oo<Nr--mONOTtinmNO<NinmONin
fNTtmNONOO<Nt"~<NmminOinr--TtTt

c«-im — — © — — O — i—i OO — © © © —

in r- © — OnOnOOOn
oo in on

nOMSn*"t«r-(NVO • • fN ©

OOOOOOOOO OOOOOOO

— Tt ON — — Tt Tt — (Nt*— —

i i i i i _1 i J. J. J i _l

m m (N (N

i

(N ON

• • Tt in in on on r-- oo

mN

— — -*ONoor-ininin

O tj- rn in r^ on o

.'TV

— r- t— oo o '

in no t+ -^- on t* en

_L I 1 1 J. _L I

Tt oo o — i oo tt r-

fN^^r^ioooor^ininin

■* m -^t rf 00 t fS

ooin^rinfN — OooONOOTtr-NDTj-oo on

■TtON — Ol^l — (N — — ON00(N<N00 — OON

i in — in —

iininoininooo

>oooooooo

^IfNTtONoor-ininin

OO^^OinOinintn

OOOOOOOOO

T* NO TT TJ- 0O Tt (N

OOOOOOO
OOOOOOO

OOOOOOOOOOOOOOOOO
A A VI VI VI VI VI VI VI A A VI A VI VI VI VI

OOOOOOOOO
VI VI VI VI A VI A A A

OOOOOOO

VI VI VI VI VI VI VI

Q.— <N CL— <

-I

<

H u c o

" ■" •- C
• O « •-

■5.2

IS:

E °

u- ^-
'— 00
i- C

c

O.t!

r: .2P

J" ■

i—] rn

S 3 S

5 u J

-C — -C

a-2o-

00

Q

d ro

eu2

00

B

V5 fl)

Si sz
PhPl,

: _ _ S J2 '5

<oo«-5'>i; j? ra *

< ^ — ~ =2^-^

c
o

i i; B) J

00

c

o<-.s2 2.

u '

OOofl," r«T3« Q ■»- ^ TTa it P O g «S -

§•£ o

S ^ . ;-oa>;™!:<;-6bui i^i i
££ mOUcSNSDol^ouo. SuuSuu

2?U

|§-S§|S||

iM^^O'

vivSdtAva •£)

Tg IU9JS •£>

<3

to

6

IU3\8 •£) ts>
VWIU33AV •£)

a

CO

IX

Q
so

oi

Q

t/3

IX

ooooooooooooooooo
A A A VI VI VI A A A A VI A VI VI VI VI A

<N©^o<D©i--©t--r--*ON'*j-©pria\©vo

OOOOONOOON<NfNfN<N(N(N<N<NtN<NcN-<t

Or-r--©'nT)-aN©r<">m
00 m Tf

©0©©0©©©0
A A A VI VI A VI VI VI

o\<s^to\©fNa\-^-oo

fN<N-H<N — <N — i— <rt

SO m © V) m ©
©©©©©O

© ©'©'©©©©

VI VI A VI A VI VI

OHmt>inooH

©O©©©©©©© ©©©©©O©

I I

©rr ..
©Tt — I

J. I i

I I I

• rl

r-r-r~NOr-^r~t^r^

in 00 NO

■* m • ©

-H-HTj-rtON00NOmND

I I I I I I I I I

t^ O N m t^ o\ m io r>

• co

■ CO

r~ (-• i— r- r- r— no

m

m in

00 © <N © • 00 m

....©..
Tf r- in in ~^ in r*-j

I I I I I I I
m r-~ r- r~ o\ <N ©

■ in m

'tvO^-tOMnm

^J-m^Ooooo'sOm in -t * "ct * O ^ n

m©Tj-\o©^DOino\-^'rj-oo(Nr^(N'nin

tJ- in r^ « i^ -i (N t on
NO © ^)- ON © — (N-h©

© no oo rfr ■* "3- r-4

— -H©©-H ©_ |

fn(N©©-H©©©©-iOO«©©m(N

©©©©©©©O© ©0©©©O©

nonononononononononononononononono ininNOin<n"ninNONO no no no no no no no

© © no r- m — m r- on ©
r-- © © •

Nor-fNcNNOr^ — — — < no

t--inoo'^-in©(N(N-Hoomr--vooornONin

On -^ -h on •

u->©©in«r-©a\mr-NOC4oo

© © in — n m r- m © oo © <N r— t— oo © in
m©r-~o\'*™Tfvo©-H©©Tt(Nr-©(N

m *-i — i >n in

■ • ■* ■ oo r4 on r- no

t ■* •©

-■-h^- — oooominin
I I I I I I I I I

mm^ONoor^-ininin

in in oo no • m ©
. . . . © . .

II I I I I
(N in no ln on (N in

• © • • • • r-

■* • Tf ^1- on in •

no (N

fNNO^OOO-^-r-ON^O— <lnON©ONON<^lTt-
ONNO<NrN]r^rnON^HNOr--ON'^-NO(N(NNO'n
■r)-(NmrgNOON-^NOoo©-^-inONTj-inoor--

Tj-mTfONooooininin

ONNO»^r--r-©'^"r^r^i
OfNir-ON© — fNr^in

■* no tt tj- © m in

NO OO NO © © 00 ©
— T)- © -h _ I -h [-

rj- m — -h © © ^ © © i-i © © © © © on i-H ©©©©©©©©© ©OOO©©©

vOl^^tO-HOOm^O ©

-q- r'lf) ■

— Hm©TfONroinNOinm<N(Noo© • m fN

II I I ill I 111 III I J. I
flv Tt n\ .

ON ■*

• ■ ON ■

ND©ONm — ooo©^tooNO(Nr~

m oo

• • no t-> r- © © oo ~-
m <N

— ««0\MOO\0"1VO

I J I I I I I I I

r*-)©ONONr-inr*-)mm

in ■ • •

fN <N m oo r— • m m m

ON in rfr l~- >n On .

Tno ■* ■* on Tf m
III III
in in ~^ <N r-~ m \o

(N fN f on oo r- m in m Tt no ^t ■* On ^r <n

©©inininininin©

©©©©OOOO©

© m © © © © in

©©©OOO©

©0©©©©0©©0©©©©©©©

A A VI VI VI VI A V VI VI A A VI VI VI VI A

J

<

Z

oi

43

"3)

C

o

H

u

o

X

LU

"c5

_o

<*-

0

n

<HhK

CJ .3

<=<r

E °

£ oo

i- a

WhI

a— i

E§c
o3«J2

£2£

n

00

O

mQ

i— I rn

- - a-^
x b x

— S — .
5S u S
o-Seui

00

5

(N (X

, X X

; .c js

00

3

©o©©©©©©© ©©o©©o©

VI VI A VI A A A A VI VI VI VI VI VI VI A

00 * S 50

i— I an"3 J3 -C! *3 »h "3 > u C 2?

<cSa^*2> .2

O. U
D.3

3 •« JS
'■^ -^ . "-1 O "S >>« C<u'Eoo^'^. I_l I

width to that of G. variegata) and a nar-
rower mastoid width. The upper inner in-
cisor is usually bilobate, but the develop-
ment of the posterior lobe varies consider-
ably in individual specimens. The upper pre-
molar has a large hypocone with its lingual
margin lying in line with the lingual margins
of the canine and the first molar (M1) in
occlusal view (Fig. 2). The lower incisors
are typically trilobate (Fig. 3), but the
outer incisor (I3) has a well-developed cin-
gulum extending along its anterior surface
below the central lobe. The occlusal surfaces
of I3 and I2 are each characterized by the
development of a posterior projection of the
cingulum to form a heel-like lobe (Fig. 4).

Measurements (mm) of the holotype — -Ex-
ternal: Total length 100, tail vertebrae 46,
hind foot 8, ear 16, tibia 16.4, forearm 41.1,
third digit - metacarpal 40.2, first phalanx
15.2, second phalanx 21.3, fourth digit -
metacarpal 37.8, first phalanx 10.9, second
phalanx 9.7, fifth digit - metacarpal 34.9,
first phalanx 7.8, second phalanx 6.9, wing
span 305, weight 9.25 grams. Skull: Great-
est length 14.2, condylobasal length 13.6,
palatal length 4.1, zygomatic width 9.9,
mastoid width 8.4, breadth of braincase 8.2,
height of braincase 5.9, supraorbital process
width 5.3, interorbital width (from front)
5.2, postorbital constriction 4.5, width
across molars (M3-M3) 6.4, length of tooth-
row (C-M3) 4.75, width across upper ca-
nines (C-C) 4.6, length of mandible
(condylo-incisive) 9.9, length of lower
toothrow (C-M3) 5.2, width across lower
canines (C-C) 3.0.

Specimens examined — Glauconycteris gleni
sp. n. — The holotype from Cameroun
(rom); plus nine adult and two subadult
males, five adult and one subadult females

all from Malabigambo Forest near Sango
Bay, Masaka District, southern Uganda in
the collection of the Los Angeles County
Museum. G. argentata (Dobson) - Camer-
oun, 8; French Congo, 2; Uganda, 6; Kenya,
49; Tanzania, 5. G. variegata (Tomes) -
Cameroun, 2; Congo, 11; Dahomey, 2;
Ghana, 2; Nigeria, 2; Zambia, 2; Sudan, 1;
Uganda, 9; Malawi, 1; South Africa (Zulu-
land), 1. G. albogutattus J. A. Allen -
Cameroun, 1; Congo, 1. G. beatrix Thomas
(including G. humeralis J. A. Allen) -
French Congo, 1; Gabon, 1; Congo, 5;
Uganda, 9; Kenya, 6. G. egeria Thomas -
Cameroun, 1 ; Uganda, 4. G. machadoi Hay-
man - Angola, 1. G. superba Hayman -
Congo, 1 ; Ghana, 1 ; Ivory Coast, 1 .

Comparisons — Glauconycteris gleni appears
morphologically intermediate between G.
argentata and G. variegata. Externally, G.
gleni resembles a pale-coloured G. argentata
in the veination of the interfemoral mem-
brane, but markings are considerably darker
and the membrane much whiter. G. gleni
lacks the dorsolateral bands of pale-coloured
pelage on the back, characteristic of G.
argentata, and the white ventral pelage con-
trasts sharply with the dark-greyish or
brownish coloration of that species. The
ear of G. gleni is larger, the tragus is wider,
and the tibia is distinctly shorter than in G.
argentata. G. gleni is significantly larger in
11 of 16 external measurements in both
sexes (Tables 1 and 2). In cranial charac-
ters, G. gleni is significantly larger (P <
.001 ) without overlap in measurements of
greatest length, condylobasal length, length
of toothrow (C-M3), and length of man-
dible, while the postorbital constriction of
G. gleni is significantly narrower (P <
0.001) in both sexes compared with G.
argentata (Tables 1 and 2). Of the 16

Fig. 3 — Anterior aspect of the lower incisors and canines of the same three specimens shown in Fig. 2
(scale bars = 1 mm). Left, G. argentata (C-C 4.5 mm). Centre, G. gleni (C-C 4.6 mm)*.
Right, G. variegata (C-C 5.0 mm).

Fig. 4 — Occlusal aspect of lower jaws of the same specimens shown in Fig. 2 (scale bars = 1 mm).
Left, G. argentata (length 9.0, C-M3 4.65 mm). Centre, G. gleni (length 9.9, C-M3 5.2 mm).
Right, G. variegata (length 9.9, C-M3 5.3 mm).

cranial characters compared, G. gleni and
G. argentata males differed significantly in

12 and females in 10. In dental morphology,
specimens of G. gleni tended to differ from
those of G. argentata in having a larger
hypocone on the upper premolar and in the
development of the cingula on the anterior
surface of the crown of I3, as well as on the
posterior portion of both I2 and I3 (Figs.
2-4).

When compared with G. variegata, G.
gleni lacks the heavily pigmented markings
on the wings, and the markings on the inter-
femoral membrane are less complex (typic-
ally, lines are laterally oriented and antero-
posterior orientation of cross-veination is
limited). The coloration of the pelage is
strikingly different from any known example
of G. variegata. The dorsal pelage is dis-
tinctly quadricoloured instead of tricoloured
as in G. variegata. Although similar in
shape, the ears are longer and the tragi are
broader in the new species. The tibia of G.
gleni is distinctly shorter (17.0 or less) than
that of G. variegata (18.3 or more; see
Tables 1 and 2). Of the 16 external mea-
surements (Tables 1 and 2), G. gleni males
are significantly smaller in nine and females
in eight when compared with G. variegata.
Although the crania of the two species are
similar in size, differences are significant in

13 of the 16 measurements in males and 10
of the 16 in females. In dental morphology,
G. gleni also differs from G. variegata by
the shorter first commissure and longer third
commissure of M3 and by the greater devel-
opment of the hypocone of P4 (Fig. 2), as
well as by minor differences in the develop-
ment of the cingula of the lower incisors
(Figs. 3 and 4).

Each of the remaining known species of
Glauconycteris can be distinguished from
G. gleni as follows:

G. poensis (Gray) - wings pale gray in-
stead of white, general pelage coloration
darker, and greatest length of skull less than
13.0.

G. albogutattus Allen - pelage coloration
much darker, wing membranes dark instead

of white, tibia greater than 17.0, and great-
est length of skull shorter (13.1 and 13.2).

G. beatrix Thomas - pelage and wing
membranes dark, ear shorter (12.0 or less),
skull shorter (greatest length, 12.0 or less),
and length of mandible less than 9.0.

G. egeria Thomas - pelage and wing
membranes quite dark combined with strik-
ing whitish dorsolateral stripes on the back,
larger square-shaped ears, greatest length
of skull less than 13.5, and length of
mandible less than 9.5.

G. machadoi Hayman - darker pelage
coloration, as well as wing and interfemoral
membranes with sharply defined veinations;
otherwise differs in the same way as G.
variegata.

G. superba Hayman - pelage black and
white above, black below, forearm length
greater than 44.0, greatest length of skull
more than 16.0.

Acknowledgments — We acknowledge the in-
valuable assistance of many individuals and
institutions that have facilitated our studies.
We particularly thank Dr. Donald R. Patten
of the Los Angeles County Museum for per-
mission to study specimens in his care. Ms.
Judy L. Eger, Ms. Liliane Lortie, and Mr.
Geoffrey E. Turner provided statistical and
other assistance. Dr. H. F. Howden, Depart-
ment of Biology, Carleton University, per-
mitted our use of the scanning electron
microscope and Mr. Lewis Ling took the
cranial photographs (Figs 2-4). Messrs.
John Williams and Robert Glen collected
much of the comparative material aided by
the financial support and cooperation of
Mrs. Noreen Curry. Dr. J. R. Tamsitt critic-
ally appraised the manuscript and gave edi-
torial assistance.

This study results from a larger bat re-
search program that has had the financial
support of the Canadian National Sports-
men's Show and of the National Research
Council of Canada (Operating Grant
A-2385).

Literature Cited

DOBSON, G. E.

1875 On the genus Chalinolobus, with descriptions of new or little known species.

Proc. Zool. Soc. Lond., pt. 3, pp. 381-388.
1878 Catalogue of the Chiroptera in the collection of the British Museum. Printed by

Order of the Trustees of the British Museum, London, 567 pp.

HAYMAN, R. W.

1967 Chiroptera. In Meester, J. ed. Smithsonian preliminary identification manual
for African mammals, pt. 2. Washington, D.C., Smithsonian Institution. 155
pp. Mimeographed.

HAYMAN, R. W. AND J. E. HILL

1971 Chiroptera. In Meester, J. and H. W. Setzer, eds. The Mammals of Africa; an
identification manual, pt. 2. Washington, D.C., Smithsonian Institution Press. 73
pp.

HOWDEN, H. F. AND L. E. C. LING

1973 Scanning electron microscopy: Low-magnification pictures of uncoated zoolo-
gical specimens. Science, vol. 179, no. 4071, pp. 386-388.

KOOPMAN, K. F.

1971 Taxonomic notes on Chalinolobus and Glauconycteris (Chiroptera, Vespertilioni-
dae). Am. Mus. Novit., no. 2451, pp. 1-10.

MILLER, G. S.

1907 The families and genera of bats. Bull. U.S. Natl. Mus. no. 57, pp. 1-282.

PETERSON, R. L.

1972 Systematic status of the African molossid bats Tadarida congica, T. niangarae
and T. trevori. Life Sci. Contr. R. Cnt. Mus., no. 85, pp. 1-32.

RYAN, R. M.

1966 A new and some imperfectly known Australian Chalinolobus and the taxonomic
status of African Glauconycteris. J. Mammal., vol. 47, no. 1, pp. 86-91.

Suggested citation: Life Sci. Occ. Pap., R. Ont. Mus.

All manuscripts considered for publication are subject to the scrutiny and editorial policies of the Life
Sciences Editorial Board and to review by persons outside the Museum staff who are authorities in the
particular field involved.

price: 754

© The Royal Ontario Museum, 1973

100 Queen's Park, Toronto, Canada M5S 2C6

PRINTED AT THE UNIVERSITY OF TORONTO PRESS .

12

*n3S

thai

Ol

'*Kn

■o

*»v*an ^

... - •

*>